Plant

Palm / Cycad
Evergreen
S.W. Mexico, Zone 9-12
Leaf scars or bases

Bark Type

Leaf scars or bases

Persistent leaf bases attached to the trunk or leaf scars that remain on the trunk.
Standard

Growth Habit

Standard

A plant with a free standing erect stem and terminal foliage.
Slow
8 - 18 m (27 - 60 ft )
7 m (23 ft)
14
Yes
Medium

Plant Overview

This fan palm has a single trunk with persistent split leaf bases and forms a dense rounded crown. The grey-green fronds are deeply divided and the creamy cup-shaped flowers appear in a large cluster amongst the foliage during spring, followed by black fruit.

 

Sabal uresana Trel. is naturally found in the state of Sabal in north-west Mexico and extends to south-western Chihuahua and northern Sinaloa growing along the Sierra Madre Occidental in valleys and foothills in oak forests to an altitude of 1,200 m (3,937 ft). It is also found in lowland dry tropical forests, commonly along river banks and drainage lines or desert scrub and coastal dunes. It prefers a very well drained poor to moderately fertile moist calcareous based sandy soil that is slightly acidic to alkaline, with a pH range from 6.0 to 8.0. It grows in an open to exposed sunny position and is drought, heat and frost-cold (-9º C) tolerant.

 

The Sonoran Palmetto is grown for its foliage colour and its upright growth habit. It is planted in parks or large gardens singulary or in groups along avenues. It is also planted for a tropical effect around a large water feature or used along river banks. It tolerates warm temperate regions (zone 9) and is suitable for coastal or inland positions. It is slow growing establishes in 4 to 6 years and is long lived. This palm is also grown in tubs and planter boxes when young and in cold regions it is placed in a conservatory. Once established it has a medium water requirement, (Scale: 2-drops from 3) preferring organic rich moist to dry sandy soil and will tolerate prolonged dry periods.

I.D. 667

UK hardiness zone H2

Climate zones H1, H2, 13 - 17, 19 - 24

USDA Zone 9-1

 

Sabal (SAY-bal) uresana (yoo-RES-ana)

 

Etymology

Genus:  Sabal - presumably named after Italian botanist, Liberato Sabbati (b. 1714)

Species:  uresana - from Ures, Mexico, where the plant is quite common

 

Arecaceae (ar-ek-AY-see-ee)        

(Palmae)

Palm Family

This family consists predominantly of tropical monocotyledons that form climbing and tree like habits with divided pinnate leaves and axillary inflorescence that produce ample indehiscent fruit.

 

Distribution

These plants are mainly distributed in tropical and warm temperate areas of the world from Central and South America to the Malay Peninsula and Africa in environments with poor drainage or permanent water such as mangrove swamps and low land rainforest.  A small number of species are found in temperate regions.

 

Diagnostic Features

Palms are long lived and form solitary or multiple trunks that are rarely branched and can form a clump of, rarely with underground stems. They have a single apical bud and if it is killed by frost or mechanical damage the stem dies.

 

The leaves (fronds) of adult plants appear in regular numbers per stem at the apex with their tubular leaf bases clasping around the stem. These forms a pseudo-stem (crown-shaft) as in Arecoid palms or the dead leaf bases form a parchment-like weaved mat and may split below the petiole. The distinctive petiole is armed or unarmed with spines or teeth and leaf scars or leaf bases may be persistent on the trunk. They vary in size from 150 mm (6 in) to 25 m (82 ft) in length.

 

The leaf blade is plicate from bud and is normally simple and adult fronds are palmately, pinnate and occasionally bipinnate with pinnate ribs. Caryota spp. is palmately lobed simple with ribs and appears compound with the petiole acting as the rachis (Costapalmate). All have ligula terminating the petiole and have leaflets that fold upwards then down wards to the petiole or rachis and have one to several parallel nerved veins joined by transverse veinlets.

 

The inflorescence is simple or branched into a huge panicle and normally arises from the leaf bases and forms with a protective overlapping woody bracts that remains until the flowers developed.

 

The numerous small flowers occur individually or 2 - 3 together in a cluster and are monoecious or dioecious with the perianth segments in 2 whorls. The female flowers are generally 3 merious with 3 somewhat imbricate sepals and petals and have 3 stamens. Male flowers sometimes have 4 petals and sepals. The flowers of pollinated by wind, ants, bees, beetles and flies.

 

There are three or more stamens and sometimes are adnate to the perianth or have the filaments joined into a tube or disk.

 

The gynoecium normally three-carpelled and may be fused or unfused. The ovary is superior with 3 carpels that have a single ovule and in many species only one ovule per flower that matures into one seed. The styles are rarely elongated and normally the arrangement is a three lobed fleshy stigma that is sessile on the ovary.

 

The fruit normally contains a single seed and it is a drupe or berry with a woody or fibrous surface that is smooth, prickly or hairy mesocarp is fleshy or dry and rarely produced 2 - 10 seeds.

 

The indehiscent seeds are commonly brightly coloured and have endosperm that is oily or fatty and a hard coated.

 

Note:

The palms have many uses from making spears to the fruit and leaf bases being eaten. There are of great horticulture importance and are used in many gardens.

 

This plant tolerates between USDA zones 9a to 12a and grows to 18 m (55 ft)

Fahrenheit         20º to 55º F

These temperatures represent the lowest average.

Celsius             -6.6º to 12.7º C

 

Attention

All photographs and data are covered by copyright. Apart from any fair dealing for the purpose of private study, research, reference or review, as permitted under the Copyright Act, no part may be reproduced by any means with out written permission. All inquiries should be addressed to www.plantfile.com attention Peter Kirkland.

Leaf

Compound

Compound

The leaf that is divided into separate units (leaflets).
Costapalmate

Leaf Shape

Costapalmate

When a palmate leaf is almost circular with the petiole extending into the blade as a mid rib and is divided to about half with segments that are forked at the apex. As in some palm species.
Spiral

Leaf Arrangement

Spiral

Leaves arrangment is twisted like a screw.
Entire

Leaf Margin

Entire

A leaf margin with no irregularities (smooth).
Silvery blue
1800 - 2500 mm ( 70.9 - 98.4 in )

Additional Information

The arching silvery-blue to grey green costapalmate fronds are up to 2.5 m (8 ft) wide and have stiff spreading segments that tend to droop at the tips. They are accompanied with a few cotton-like threads. The petiole is short and spreading at the base and persists on the trunk.

 

Note:

Inland specimen’s foliage is distinctly silvery-blue and the coastal plants tend to be glaucous-green.

Flower

Cyathiform

Botanic Flower Description

Cyathiform

A cup like shape.
Odorless
Panicle

Flower Inflorescence

Panicle

Branched with large loose clusters.
Creamy white
4 - 7 mm ( 0.2 - 0.3 in )

Flowering Season

(Southern Hemisphere)

Jan Feb Mar Apr May Jun
Jul Aug Sep Oct Nov Dec

Additional Information

The small 7 mm (¼ in) wide creamy-white cyathiform flowers are bisexual with 3 fused sepals and 3 fused petals. They are arranged in a large panicle that appears in the leaf bases during early summer.

Fruit

Drupe

Fruit Type

Drupe

A succulent fruit composed of an outer fleshy layer "
Black
No
12 - 18 mm ( 0.5 - 0.7 in )

Fruiting Season

(Southern Hemisphere)

Jan Feb Mar Apr May Jun
Jul Aug Sep Oct Nov Dec

Additional Information

The 18 mm (7/8 in) diameter spherical drupe has a thin dry mesocarp and contains a thin black seed.

Environment

Well drained, poor-fertile, moist-dry sandy soil, acidic-alkaline pH 6.0-8.0
Tubs, planter boxes, under glass in cold regions
Full sun, open to exposed position, drought-heat and light frost tolerant
Sub-tropical
Palm leaf-scab, leaf blight, fungal leaf spots, fusarium wilt, red spider mite

Cultural Uses

The Sonoran Palmetto is grown for its foliage colour and its upright growth habit. It is planted in parks or large gardens singulary or in groups as a specimen or used along avenues. Itis also planted for a tropical effect around a large water feature or used along river banks. It tolerates warm temperate regions (zone 9) and is suitable for coastal or inland regions. It is slow growing establishes in 4 to 6 years and is long lived. This palm is also grown in tubs and planter boxes when young and in cold regions it is placed in a conservatory.

Cultivation

Not normally required, remove spent foliage
Mulch the soil to keep root zone cool and moist during summer, apply complete fertiliser in spring

Propagation

Sow ripe seeds in a well-drained media and maintain a temperature of 19º to 27º C (66º to 81º F). Germination takes up to 3 months.

 

Palm Seed (General)

Palm seed should be sown as soon as possible after collection as the viability period is short, ripe mature fruit is essential. Do not allow the seed to dry out and if placed in a container of water any seed that floats should be discarded. After selecting seeds dust with a fungicide, as they are prone to fungal attack.

 

A soil mix composed of half peat moss and half perlite is a reliable media and then placed the containers over bottom heat of 28º C (80º F) and don't allow it dry out. It may take up to 1 or 2 years for germination of some species this may be accelerated by scarification or soaking the seeds in gibberellic at 1000 ppm for two days.

 

Propagation by Seed (General)

 

Germination

In order for a seed to germinate it must fulfil three conditions.

 

1. The embryo must be alive (a viable seed).

 

2. The seed must have no dormancy-inducing physiological, physical or chemical barrier to germination; also the seed must be nondormant.

 

3. The seed must have the appropriate environmental requirements, water, temperature and oxygen.

The interaction between these requirements and dormancy is complex and may lead to different environmental requirements that avoid the dormancy of a seed.

 

Sowing Seeds in Containers

There are two general methods for germinating seeds.

Seeds in a flat or germinating bed, through which seedlings are pricked-out then, transplanted into another flat with wider spacing or directly to an individual pot.

 

2. Sowing seeds by placing them in to flats with the appropriate spacing or into individual pots.

This method is normally carried out with medium to large seeds such as woody plants and plants that are difficult to transplant.  

Seedling production normally occurs in a greenhouse / glasshouse, cold frames and on hot beds.

 

Method of Seed Sowing

Fine seed is sown in pots or flats that are no deeper than 70 to 80 mm. using a sterilised well-drained media (soil). Fill the container to 20 mm from the top and sprinkle sieved peat to 3 mm depth.

Press the media down level and firm with a piece of timber and then thoroughly moisten.

 

Mix the fine seed with washed sand and then sow thinly on the surface. These may be lightly covered with sand.

Larger seeds may be covered with media or a hole is dibbled and the seed is placed in the media.

 

Watering Methods

For watering you may either mist the containers from above or place the container in tepid water and allow the water to raise through the pot to the surface of the media, then drain away and do not fill to the top of the container.

 

Place a piece of glass over the pot and store in a protected warm environment (glasshouse).

Seeds germinate best in darkness so shade the containers if in direct sunlight.

 

After the seedlings have sprouted remove the glass and ease the seedlings into direct light.

When the seedlings are large enough prick them out and transplant into larger containers then place them in a shade house to harden off.

Many seeds have different methods of seed preparation for germination such as nicking or cutting the seed coat to allow water penetration, also placing seeds in hot water and allowing it to cool off.

This is particularly important as it is softening the seed coat.

Pests

83
Two-spotted Mite, Red Spider Mite
Tetranychus urticae
Acarina
Tetranychidae

PEST

   NAME

     Two-spotted Mite, Red Spider Mite

     Tetranychus urticae

   ORDER

     Acarina

   FAMILY

     Tetranychidae


Description of the Pest

Also known as the red spider mite. Females are pale green or yellowish, depending on the host plant, and have two dark lateral markings; the mite becomes red in winter, retaining their dark markings. Nymphs are six-legged, with another pair of legs appearing as the mite matures. Males are smaller and narrower. Fully-grown adults are just visible to the naked eye. Two-spotted mites spread by crawling between nearby plants or movement of dead leaves.


Appearance and Distribution of the Pest

Found world-wide; an introduced pest in Australia. They congregate in protected places, such as under bark and at the base of trees, during winter. During spring, they become green in colour, and migrate back into the leaves. During heavy infestations, the leaves may be covered in visible webs, which they spin as they feed. Leaves may eventually wither and fall. Mites can spread via the movement of dead leaves, or in webs that have become attached to birds or large insects. They initially appear on the undersides of leaves.



Spruce Spider Mite (Oligonychus ununguis) is a tiny greenish black adult which lays eggs on twigs where they overwinter. The pale green young spiders suck the sap turning the leaves yellow to brown. Heavy infestations form webbing and the pest is found on Abies and Juniperus species.


Banana spider mite (Tetranychus lambi) is a major widespread pest of bananas.  It differs from two spotted mite by not producing copious amount of webbing.  It is highly active during the dry spring to summer period and with the onset of the wet season mite numbers are reduced.  The warm dry conditions that are created under plastic bunch covers is ideal for building up banana spider mite numbers.

Damage is normally confined to the underside of leaves appearing as rusty patches that coalesce along the leaf veins eventually turning the whole leaf brown-grey before it collapses.  Fruit is damaged, close to the bunch stalk causing the area to become dull red purple-black, which in turn becomes dry then cracks.


  Damage fruit


Control methods include careful water management during dry periods, and the reduction of dust from roadways.  Regular desuckering and leaf trimming of plants will assist with a good coverage when spraying miticides.


Life Cycle

Mites have a gradual metamorphosis, with several nymphal stages. Each female lays up to 100 eggs that hatch in 7-14 days, with several generations appearing throughout the year. Females may become inactive during cold weather.


Webbing         


Period of Activity

The Two-spotted mite is most active in hot dry conditions. Under optimum conditions, the population can double every four days. It produces large quantities of webbing for over-wintering nests. Many plants are only susceptible to this insect when cultivated under glass.


Damage Caused

Adults and nymphs lacerated the undersides of the leaves with there rasping mouth parts, although infestations on both surfaces are not uncommon. Infestations cause leaf mottling leaf fall; premature leaf loss causes loss of vigour and reduces the quality and quantity of future crops. Repeated infestations, year after year, may weaken root growth and kill herbaceous plants.



Susceptible Plants

A wide range of plants are attacked by the Red Spider Mite including annuals, fruit trees and vegetables, ornamental shrubs and trees.

Note

Many plant species are more susceptible to Red Spider Mite when they are cultivated under glass.


Other species of mite that are mentioned below have simular characteristics.


Calluna, Rose, Tropaeolum and Viola and species are infested with the Red Spider Mite (Tetranychus telarius) commonly in greenhouse situations.


Musa species are attacked by two spotted mite and banana spider mite damaging foliage and fruit.


Juglans species can be infested with up to four types of mites including red spider.


Cultural Control

Heavy rain or irrigation can reduce numbers; some plants may benefit from replanting in cooler locations. Generally, however, infested material should be completely removed and destroyed.

Preventative measures such as removing weeds or mulching around trees or shrubs or scrubbing the loose bark of susceptible trees during winter helps reduce numbers. During spring sticky bands can be wrapped around trunks close to the ground to trap the mites.


Biological Control

Natural predators include lacewings, ladybirds and thrips help keep the numbers down. Insecticide-resistant predatory mites (Typhlodromus occidenyalis) are also available commercially to control the Two-spotted Mite only on a large scale, as they require ample mites to survive.


Chemical Control

Spraying should be carried out as a last resort as many predators are killed during the operation and spraying can have the opposite effect by increasing numbers in the long term. Dimethoate will reduce numbers; however, Two-spotted mites are resistant to insecticides in some areas. Dusting with wettable sulphur may also prove effective.

Note

Always read the label for registration details and direction of use prior to application of any chemicals.


87
Scale Insect
Various Scale Species
Hemiptera

PEST

 

   NAME

     Scale Insect

     Various Scale Species

 

   ORDER

     Hemiptera

 

Description of the Pest

Generally scales are soft bodied insects that have a hard (armoured) or soft covering to hide under. They have piercing and sucking mouth parts that are attached to the host, feed off sap and soft scales commonly producing sweet honeydew, which in turn attracts sooty mould and ants.

The adult female has a circular or oval covering depending on the species and is up to 8mm across. The first stage (crawlers) hatch and wander around the leaf surface until finding a suitable place to suck sap, normally in colonies and the smaller male is relatively inconspicuous.

 

Hard Scale                   Soft Scale, attending Ants

 

Cactus Scale (Diaspis echinocacti) has a circular greyish female and a narrow white male scale and is commonly found on house plants.

 

Chain Scales (Pulvinaria species) adult females are obvious with large group of eggs that are white or cottony-like, and the tiny young light green scales are flat and oval-shaped up to 2mm long. The legged nymphs are normally arranged from head to tail along the mid rib of the leaf, and may move to a new position to feed. They excrete honeydew and attract sooty mould and are found on Acacia and Acronychia species.

 

Chinese Wax Scale

 

Chinese Wax Scale (Ceroplastes sinensis) is a domed wax scale that has dark spots around its margin and immature scales form waxy material around there margins.

 

Fern Scale on Aspidistra elatior

 

Fern Scale or Coconut Scale (Pinnaspis aspidistrae) appears as flecks up to 0.15mm long with a white covering over the male congregating on the underside of the fronds on the axils and among the sporangia causing them to turn yellow. Many species of fern are susceptible to infestation.

 

                  Flat Brown Scale

 

Flat Brown Scale (Eucalymnatus tessellates) are light brown up to 0.5mm long, flat and closely attached both sides of the leaf and causing yellowing of the foliage.

 

Juniper Scale (Diaspis carueli) is tiny and circular, white maturing to grey-black and as it feeds the needles turn yellow and die.

 

Oleander Scale (Aspidiotus hederae) is a pale yellow circular scale up to 3mm across and is found in dense colonies on the stem or leaves.

 

Tea-tree Scale (Eriococcus orariensis) are a creamy blue colour normally packed along the branches and are plump and rounded to 4mm across.

 

                  Wattle Tick Scale

 

Tick or Wattle Scale (Cryptes baccatus) adult is domed, blue-slate colour with a leathery covering up to 10mm long. All stages of growth are found in groups of over forty, packed along the stems and normally tended by ants as they produce large amounts of honeydew. A serious pest of Acacia species found inland or coastal from temperate to sub tropical climates and commonly accompanied by Sooty Mould.

 

Toxic Scale (Hemiberlesia lataniae) is a tiny flat rounded scale up to 0.15mm long and is white to pale pink. It is normally found in colonies on the small branches and twigs of shrubs. It injects a toxic substance into the host as it sucks sap causing the death of the branch.

 

Wattle Scale (Pseudococcus albizziae) is soft, plump and secrets cotton-like threads. It is not a true scale insect and is simular to mealy bugs. It is reddish-brown up to 0.4mm long and secrets large amounts of honeydew as it sucks sap in colonies along the branches.

 

Life Cycle

These insects have a Hemimetabolous life cycle, ie. When the immature nymphs resemble the adults.

 

Appearance of the Pest

All parts of the plant above the soil may be attacked, but normally the stems and leaves and scale tends to favour well-lit positions.

 

Period of Activity

The nymphs and females are active for most of the year, in warm climates. Once they selected a position they attach and don't move. Normally the winged or wingless males are mobile and only soft scales produce honeydew.

 

Susceptible Plants

There is a wide range of susceptible plants including citrus, willows, holly, and many ornamentals, such as roses or Paeonia species. It also attacks indoor or glasshouse plants and Australian native plants such as wattles, hakeas, grevilleas and eucalyptus.

 

Acacia species are attacked by the Tick or Wattle Scale, which infest twigs and small branches and heavy infestations will kill the host plant.

 

Acer species are attacked by the Cotton Maple Scale (Pulvinaria innumerabilia) which prefers Acer saccharinum. Nymphs first attack the leaves and the brown adult scale is covered in a woolly mass up to 14mm across, normally found on the underside of the stems and twigs.

 

Acmena smithii, Melaleuca, Syzygium and Pittosporum species are attacked by the Chinese Wax Scale.

 

Aesculus species are attacked by several scale insects including the Walnut Scale (Aspidiotus juglans-regiae) which is saucer-shaped and attacks the main trunks.

 

Agave species are susceptible to several types of scale including (Aspidiotus nerii), (Aonidiella aurantii) and (Pinnaspis strachani), but generally do not require control.

 

Asplenium australasicum

 

Asplenium australasicum is susceptible to Coconut Scale or Fern Scale (Pinnaspis aspidistrae). It is normally found on the under side of the fronds. Small infestations cause little damage.

 

Bougainvillea species may be attacked by the soft scale (Coccus hesperidum) outdoors or under glass.

 

Calluna and Vaccinium species are attacked by the Oyster Shell Scale (Lepidosaphes ulmi).

 

Camellia species may be attacked by the Florida Red Scale (Chrysomphalus aonidum), which is small, circular and black and is found firmly attached to the underside of the leaf along the veins. On inspection after removing the scale the insect has a pale yellow body. Camellias are also attacked by a large variety of scale insects including Tea Scale and Camellia Scale.

 

Carpinus species may be attacked by the scale (Phenacoccus acericola). It is found on the underside of the leaves forming a white cotton-like clump along the veins.

 

Casuarina and Allocasuarina species may be attacked by the Casuarina Scale (Frenchia casuarinae), a black hard scale that is upright to 4mm with a pinkish body. During attachment the surrounding tissue swells up and in time can, form galls. This weakens the wood and in severe infestations may kill the tree.

 

Cotoneaster species are attacked by up to four species of scale including the Oyster Shell Scale (Lepidosaphes ulmi).

 

Cupressus species are attacked by Bark Scale (Ehrhornia cupressi) is pink and covered in white wax. Heavy infestations cause the leaves to turn yellow or reddish.

 

Flat Brown Scale on Cycas revoluta

 

Cycads, palms and some species of Callistemon are attacked by the Flat Brown Scale.

 

Erica species are attacked by several species of scale including, Greedy, Oleander and Oystershell scale.

 

Jasminum species can be infested with up to twelve types of scale.

 

Juniperus x media and other conifer species are attacked by the Juniper Scale.

 

Leptospermum species are attacked by the Tea-tree Scale which produces ample honey dew that promotes sooty mould.

 

Palm and Fern species are susceptible to attack by the Coconut Scale or Fern Scale (Pinnaspis aspidistrae) which infests the underside of the leaves. They are also hosts for many other scale species such as red, cottony cushion and tea scale.

 

Pinus species are attacked by several species of scale including the Pine Tortoise Scale (Toumeyella numismaticum) and the Red Pine Scale (Matsucoccus resinosae).

 

Polygonum odoratum is attacked by a small brown scale.

 

 

Sorbus aucuparia is attacked by a five species of scale insect, including Black Cottony Maple, San Jose and Scurfy. Generally they suck on the sap of the new growth and leaves.

 

Strelitzia species are attacked by the Greedy Scale (Aspidiotus camelliae).

 

Damage Caused

Leaves become yellow and are shed prematurely and there may be twig or stem die-back. When the infestation occurs on fruit, the fruit is small and its skin becomes pitted and cracked. Small trees and saplings that are heavily infested may be seriously damaged or die. Sooty mould can cover fruit or leaves causing a secondary problem.

 

Cactus Scale can completely cover the host cactus sucking sap and causing it to die.

 

Cultural Control

Dead or damaged parts of the plant should be removed and destroyed including fallen fruit. Small infestations may be removed by hand or squashed on the stems. Healthy plants are less susceptible to attack, so maintain vigour of the plant and avoid using high-nitrogen fertiliser that produces excessive soft young growth.

When pruning susceptible plants paint the cuts with antifungal sealant paint as scale insects are attracted to the sweet smell of the sap. This will reduce the infection rate of the plant.

 

Biological Control

Natural predators such as parasitic wasps may reduce numbers of active nymphs; parasitic wasps are bred commercially in some areas for this purpose. It should be noted, however, that wasps would avoid dusty conditions.

Other predators that assist in control are assassin bugs, ladybirds, lacewings, hover flies and scale eating caterpillars. A variety of birds also attack scales.

The control of ants that transport aphid from one host to another also reduces infestation and can be carried out by applying at least three greased bandages 5mm apart around the stem or trunk of the plant.

 

Chemical Control

Spray the entire plant with dilute white oil solution; a follow-up spray may be required after four weeks, for heavy infestations. Spraying of chemicals will also kill of natural predators and in some cases the secondary scale infestation is more prolific especially when using copper based chemicals.

Some chemical controls, such as methidathion, are available - please seek advice from your local nursery as to the suitable product for your area.

Note

Always read the label for registration details and direction of use prior to application of any chemicals.


Diseases

26
Leaf Spot (General)
Various Leaf Spot Species

DISEASE

   NAME

     Leaf Spot (General)

     Various Leaf Spot Species



Description

There is a wide variety of fungal leaf spots that infect perennials, shrub and trees. Some are specific to the host while others can affect a range of plants.


Symptoms

Generally light brown to purplish or blackish spots appear on the leaf and form concentric rings of fruiting bodies. The spots may leave holes, perforating the leaf or expand with pale green to yellowish margins and when the holes merge the leaf normally dies. There are many different types of leaf spot, some are discussed below.


Alocasia species     


Alternaria Leaf Spot (Alternaria nelumbii) forms a small reddish brown spots that are boarded in light green, and as they develop in size the leaf curls and dies from the margin inwards. Normally occurs on Nelumbo species (water lilies).


Helminthosporium Disease


Helminthosporium Disease (Bipolris species), (Drechslera species) and (Exserophilum species) are responsible for several leaf spots that occur on all Turf Grass species. Generally they form black or white spots that may be faded and produce masses of spores in the thatch during late summer, under humid conditions. The life cycle is short and when conditions are favourable spores are splashed onto the foliage from the thatch, causing wide spread infection. Cynodon dactylon (common couch) is most susceptible and found in bowling or golf greens where it is a serious problem.


Banana Leaf Spot     


Banana Leaf Spot (Mycosphaerella musicola) is found on many species of banana causing pale yellow streaks on the young leaves to turn brown with dark spots. The leaf then becomes dried, brown and dead commencing from the margins, eventually the leaf dies. Control requires removal of infected foliage or the spraying of a fungicide and fungicides should not be used during the fruiting period.


                  Lophostemon confertus (Brush Box)


Leaf Spot on Brush Box (Elsinoe species). This is a casual fungus that attacks the epidermal layer of the leaf, forming circular spots that are up to 25mm across and are often restricted by the main vein.  These spots are a dull yellowish brown but can also have purplish patterns.  A leaf may have more than one spot develop on its surface and normally appears on scattered leaves throughout the tree.  This doesn't affect the vigour of Lophostemon confertus.


Palm Leaf-scab (Graphiola phoeicis) appears as yellow spots and develop into scabs or warts that are outwards hard and dark but with a soft centre with powdery yellowish brown spores. The infected leaves eventually die.


Palm Leaf Spot, Chamaedorea elegans


Palm Leaf Spot (Pestaloptiopsis species) appears as a small spot with a dark centre on the leaves and affects palms that are growing in shaded humid positions and normally control is not required, though infected fronds should be removed.


Source and Dispersal

Infection source is other contaminated plants and the spores are spread by wind or by splashing water.  The fruiting bodies are black spots that appear on the damaged tissue releasing spores.


Favoured Conditions

This fungus prefers a warm humid environment and leafy plants with soft new growth, particularly if they are crowded.

        

Affected Plants

There are many ornamental and native plants that are hosts to a wide range of fungal leaf spots. Some specific ones are listed below. Plants such as Cornus or Paeonia species are infected by a large variety of leaf spots, while other plants attract a specific leaf spot.

Generally a healthy plant can tolerate fungal leaf spot attack, though it may make the plant look unsightly. In trees and shrubs it is difficult to control and generally not necessary, but in perennials and annuals control may be necessary in order to save the plant.


Acalypha and Arctotis species are infected by up to three leaf spots including (Cercospora acalyphae) and (Ramularia acalyphae) that rarely require control.


Acer species are infected by Purple Eye (Phyllosticta minima) which forms spots with brownish centres and purplish margins causing the death of the leaves.


Acer species are also infected by Tar Spot (Rhytisma acerinum) which forms round black spots that have yellow margins. Not normally seen on cultivated trees, but seen in forests.


Adiantum, Asplenium, Blechnum, Cyathea, Davallia, Nephrolepis, Platycerium, Polypodium and Pteris species are infected by the leaf spot (Pseudocercopora species) which forms circular brown spots on the fronds and heavy infection can defoliate a plant.


Aesculus species are occasionally infected with the leaf spot (Septoria hippocastani) which forms small brown spots.


Agave species are susceptible to the leaf spot (Coniothyrium concentricum), which appear as greyish spots up to 20mm (1in) across with concentric rings and black fruiting bodies. Affected leaves are destroyed as the infection spreads.


Albizia julibrissin is susceptible to the fungal leaf spot (gloeosporium aletridis), which does not normally require control.


Amelanchler, Chaenomeles, Crataegus and Rhaphiolepis species Mespilus germanica are infected by the leaf spot (Fabraea maculata) which may cause considerable damage during wet periods.


Aquilegia species can be infected by three types of Leaf Spot including (Ascochyta aquilegiae), (Cercospora aquilegiae) and (Septoria aquilegiae), normally appearing during humid conditions forming spots on the leaves.


Arbutus species are infected by two leaf spots (Septoria Unedonis) which produces small brown spots on the leaves and (Elsinoe mattirolianum).


Arctostaphylos manzanita is infected by the leave spot (Cryptostictis arbuti) which damages leaves but is not normally detrimental to the shrub.


Aspidistra species are infected by the leaf spot (Colletotrichum omnivorum) causing whitish spots on the leaves and petiole.


Aster species are infected by many leaf spots including (Alternaria species), (Cercosporella cana), ( Ovularia asteris) and (Septoria asteris).


Aucuba species are infected by several leaf spots, usually as a secondary infection after aphid attack. These include (Phyllosticta aucubae) and (Phyllostica aucubae).


                     Azalea


Azalea (Rhododendron species) are susceptible to Leaf Scorch (Septoria azalea). This fungal disease forms reddish- brown spots which expand and engulf the leaf, with fruiting bodies appearing in the centre. Infected leaves die, then fall and the branchlets wilt. This problem is more serious during wet periods and may require control using a fungicide.


Banksia robur


Banksia species are infected by several leaf spots causing chlorotic areas that have brown centres and is not normally a major problem for the plant.


Betula species may be infected by the Leaf Spots (Gloeosporium betularum) that forms brown spots with darker margins and (Cylindrosporium betulae) that also forms brown spots with faded indefinite margins.


Bougainvillea species are infected by the leaf spot (Cercosporidium bougainvilleae) which forms rounded spots with dark margins that yellowish ting. Infected leaves die and fall from the plant.


Calendula species are infected by the Leaf Spot (Cercospora calendulae) which rapidly infects the plant spotting the leaves and killing the plant.


Callicarpa species may be infected by the leaf spot (Atractilina callicarpae) forming irregular brownish spot or (Cercospora callicarpae) which can defoliate the plant in subtropical climates.


Campsis species may be infected by several fungal leaf spots including (Phyllosticta tecomae), (Septoria tecomae) and (Cercospora duplicata).


Carpinus species are infected by the leaf spots (Gloeosporium robergei), (Gnomoniella fimbriata) and (Septoria carpinea), all are minor infections not normally requiring control.


Carya species are infected by several leaf spots including (Gnomonia caryae) that infects leaves with irregular reddish spots on the upper surface with corresponding brown spore producing spots on the underside. It also has a secondary spore release that occurs on the dead leaves where it over winters. Other leaf spots include (monochaetia desmazierii) and (Marssonina juglandis).


Ceanothus species are susceptible to the leaf spot (Cercospora ceanothi) and (Phyllosticta ceanothi) both are of minor importance not requiring control.


Celtis species are infected by many leaf spots including (Cercosporella celtidis), (Cylindrosporium celtidis), (Phleospora celtidis) and (Septogloeum celtidis).


Chrysanthemums species are infected by the leaf spot (Septoria species) which forms yellow spots appear toward the edge of the leaves; these become enlarged brownish patches with yellow margins.  Damaged areas may converge and in severe attacks and the leaves may fall prematurely or flower production is reduced.


Clematis species are infected by the fungal disease (Ascochyta clematidina) which may cause stem rot or leaf spots that are water soaked areas with reddish margins. The infection spreads from the leaves to the stem causing wilting and eventually girdling the stem killing the plant. There are many fungal leaf spots that infect this plant including (Cercospora rubigo) and (Septoria clematidis)


Dracaena deremensis


Cordyline and Dracaena species may be infected by the leaf spot (Phyllosticta maculicola) which forms small brownish spots that have yellowish margins and has black fruiting bodies that forms coils of spores. These plants are also susceptible to other leaf spots such as (Glomerella cincta) and (Phyllosticta dracaaaenae). Keep foliage dry to avoid infection.


Cynodon dactylon,  Pennisetum clandestinum and many other Turf Grasses are susceptible to Helminthosporium Disease.


Daphne species are infected by the leaf spot (Gloeosporium mezerei) and (Marssonina daphnes) both of which form thickish brown spots that are seen on both sides of the leaves. Infected leaves turn yellowish before dieing.


Dendranthema species are infected by many leaf spots such as (Septoria chrysanthemi) which first forms yellowish spots up to 25mm (1in) across that become black. Infected leaves die prematurely and persist on the plant.


Dianthus species may be infected by the leaf spot (Septoria dianthi). It forms light brown rounded spots that have a purplish border. The scattered spots on the lower leaves can also be found on the stems and the spores are dispersed by water from the tiny black fruiting bodies.


Dieffenbachia species are infected by several leaf spot fungi including (Cephalosporium species) and (Myrothecium species).


Eucalyptus species


Eucalyptus species are infected by many fungal leaf spots such as (Mycosphaeralla species), (Hendersonia species) and (Monocheatia monochaeta). Generally leaf spots appear on the juvenile or new leaves causing brownish spots that enlarge and may have a purplish halo around the margin.  Mature adult leaves are not normally infected and the trees rarely require control measures.


Fern species are infected by the leaf spot, (Alternaria polypodii). This fungus appears as brown circular or oblong spots that congregate along the margins of the pinnae causing the fronds to turn brown and die.  It is spread by wind currents from plant to plant and control methods include removing infected fronds and maintaining a drier atmosphere.


Ficus species are infected by various fungal leaf spot including (Pseudocercospora species). Generally the fungal attack forms circular or irregular dark coloured spots on the leaves eventually causing them to fall prematurely.


Ficus elastica is susceptible to many fungal leaf spots including (Alternaria species), (Leptostromella elastica) and (Phyllosticta roberti).


                    Strawberry


Fragaria x ananassa (Strawberry) is infected by the fungal leaf spot (Mycospharella fragariae). The mature leaf is initially infected with well defined brown spots that that turn light grey with red-purplish margins. As the spots merge they form large brown blotches and the leaf turns yellow then dies. This fungal attack normally occurs on plants in poor health and can be a serious problem early in the season seriously damaging stock.


Fraxinus species


Fraxinus species are infected by the leaf spot (Gloeosporium aridum) giving the leaf a scorched appearance as large blotches appear from the margin or apex and turn brown with a papery texture. It is more prevalent during rainy periods and infected leaves fall prematurely. Collect and depose of fallen leaves otherwise control is not normally required.


Fuchsia species may be infected by the leaf spot (Septoria species) or ( Cercospora species), both form spots with dead centres and dark margins.


Gladiolus species are infected by Hard Rot or Leaf Spot (Septoria gladioli). On the corms reddish brown circular water soaked spots become large and sunken. These areas dry out and form obvious margins. The leaves may also have these symptoms but is not commonly seen.


Hemerocallis species are infected by several leaf spots including (Cercospora hemerocallis) and (Heterosporium iridis). These may be in the form of black spots or brownish spots that converge killing the leaf. Infected leaves should be removed and burnt.


Hibiscus species


Hibiscus rosa-sinensis, Hibiscus syriacus and Hibiscus tiliaceus are susceptible to several fungal leaf spots including (Ascochyta abelmoschi), (Cerospora kellermanii) and (Phyllosticta hibiscina). All cause spotting or blotching of the leaf surface; remove and destroy infected parts.


Hydrangea species are infected by four fungal species including (Ascochyta hydrangeae), (Phyllosticta hydrangeae) and (Septoria hydrangeae).


Iris species are infected by several fungal leaf spots including (Alternaria iridicola) and (Macosphaerella species).

Iris species are also infected by the leaf spot (Didymellina macrospore) that forms greyish spots with brown water soaked borders and coalesce on the upper part of the leaf. This casual organism commonly occurs after flowering killing the leaves but will not infect the bulbs. The bulbs become weak over several seasons due to the decreased foliage.

There is also a Bacterial Leaf Spot (Bacterium tardicrescens) that is commonly mistaken as a fungal problem causing translucent spots that coalesce and involve the entire leaf. Normally found on Iris species.


Laburnum anagyroides is infected by the Leaf Spot (Phyllosticta cytisii). The leaf forms light grey spots with no definite margin and mature to brown. The black fruiting bodies appear as dots in the centre of the spot.


Leucanthemum species are infected by the leaf spot (Cerocspora chrysanthemi) and (Septoria leucanthemi).


Magnolia species are susceptible to many species including (Alternaria tenuis), (Mycosphaerella milleri) and (Phyllosticta species). Leaves generally turn brown from the apex or margins turning brown or spots appear on the leaf surface and leaves become yellow before withering and dieing. Normally the make the tree look poorly but have little effect on its growth. Control is not normally required.


Nerium oleander is susceptible to several fungal leaf spots including (Cercospora nerella), (Cercospora repens), (Gloesporium species) and (Phyllosticta nerii). Infected leaves should be removed but generally control is not required.


Nyssa sylvatica is infected by the leaf spot (Mycosphaerella nyssaecola) forming irregular purplish blotches.


Orchids such as Cattleya, Cymbidium, Cypripedium, Dendrobium, Epidendrum, Paphiopedilum, Phalaenopsis and Zygopetalum species are infected by several leaf spots including (Cerospora, Colletotrichum and Phyllosticta species). Normally forming dark or dead, circular or irregular areas on the leaves.


Palms such as Syagrus, Howea, Phoenix, Roystonea and Washingtonia species are infected by Leaf-scab (Graphiola phoeicis).


Palms such as Archontophoenix, Caryota, Chamaedorea, Cocos, Dypsis, Howea, Liculia, Linospadix, Livistona, Phoenix, Ptychosperma, Rhapis, Roystonea, Syagrus, Washingtonia and Wodyetia species are susceptible to several fungal leaf spots including;

(Bipolaris spp.), (Cylindrocladium spp.), (Colletotrichum spp.) and (Pestalotiopsis spp.).


Generally the circular leaf spots are brown and may have a yellow halo such as Palm Ring Spot (Bipolaris incurvata). They vary in size from small to large depending on the species. When a plant is healthy it recovers from attack, but heavy infections can defoliate, causing the collapse of the plant.


Palms are also infected by the Brachybasidium Leaf Spot (Brachybasidium pinangae). This fungus forms angular leaf lesions that produce fruiting bodies on the underside and is commonly found on Archontophoenix species.


                  Archontophoenix cunninghamiana


Passiflora species are infected with many types of leaf spot such as (Alternaria passiflorae).


Phoenix species are susceptible to False Smut (Graphiola phoenicis). This fungus forms yellow leaf spots that become hard with a raised with a blackish scab, which produces masses of powdery spores that are thread-like.


                  Pittosporum species


Pittosporum species are susceptible to the leaf spots (Alternaria tenuissima), (Phyllostica species) and (Cercospora pittospori). Circular or angular dark spots appear on the leaves and are surrounded by necrotic areas that are yellowish. Generally removal of infected leaves is adequate control.


Poa species and other cool season grasses are infected by Winter Fusarium Leaf Disease (Fusarium species), which causes small pale spots that are water soaked to appear on the leaves that turn red-brown. Infected leaves become bleached then wither and die, but the infection will not affect the crown or roots of the plant. It can be identified by pink, cotton-like mycelium and the plant prefers cold wet weather.


Populus species are infected by several fungal leaf spots including (Ciborinia bifrons, Ciborinia confundens), and (Mycosphaerella populicola).


Prunus species are infected by several leaf spots including (Cercospora circumscissa and Septoria ravenelii).


Pseudotsuga menziesii Douglas Fir is infected by the Leaf Cast (Rhabdocline pseudotsugae) Symptoms include the needles becoming yellowish at the apex and extending down the needle and spreading to others during moist spring weather turning them brown. Brownish scorched areas are noticeable on the tree from a distance. Control; is not normally required for mature trees but nursery stock may require spraying with a copper based fungicide.  


Psidium guajava (Guava) is infected by (Glomerella cingulate). This fungus courses spots to appear on leaves and mummifies and blackens immature fruit or rots mature fruit.  This fungus can devastate a guava crop.  


Quercus species are infected by several types of leaf spot including (Cylindrosporium microspilum) and (Marssonina martini). These attacks tend top take place later in the season and normally not detrimental to the tree.


Rhododendron species are infected by a large variety of fungal leaf spots including (Cercospora rhododendri) and (lophodermium melaleucum)


Salix species are infected by several fungal leaf spots including (Ascochyta salicis) and (Septogloeum salicinum).


Senecio species are infected by the fungal leaf spot (Alternaria cinerariae) and (Cercospora species), forming dark rounded or angular spots.


Spiraea species are attacked by the fungal leaf spot (Cylindrosporium filipendulae).


Stenotaphrum secundatum (Buffalo) turf grass is susceptible to Grey Leaf Spot (Pyricularia grisea) in domestic and commercial situations devastating lawns. This fungal disease infects the stems and leaves with small brown lesions that enlarge rapidly forming grey-brown spots that have darker borders or surrounded by yellow chlorotic areas. This infection is commonly found on newly laid turf but will also infect established lawns. It is most prevalent during warm humid periods in soil with a high nitrogen level.


Syringa species are attacked by up to six species of leaf spot including (Cercospora lilacis) and (Phyllostica species).


Syzygium species


Syzygium species are infected by fungal leaf spots but normally control is not required.


Tagetes species are infected by the leaf spot (Septoria tageticola), which starts at the base and moves progressively up through the plant, covering the leaves in grey to black spots.


Trillium species are host to several leaf spots, including (Colletotrichum peckii) (Gloeosporium Trillii) (Heterosporium trillii).


Ulmus species are infected by many fungal leaf spots including (Gnomonia ulmea) and (Cercospora sphaeriaeformis).


Veronica species are infected by the leaf spot (Septoria veronicae). The symptoms include small violet to brown spots appear on the upper surface of the leaf and correspondingly yellowish brown on the underside. The spots converge forming a scorched shot-hole appearance and eventually death of the leaf.


Vaccinium ovatum


Vaccinium ovatum is infected by the leaf spot (Rhytisma vaccinii) and (Dothichiza caroliniana).


Vicia species are infected by the leaf spot (Erostrotheca multiformis), which forms greyish spots that enlarge and may defoliate the plant.


Wisteria species are infected by three fungal leaf spots (Phyllostica wisteriae), (Septoria wisteriae) and (Phomatospora wisteriae).


Non-chemical Control

Remove and destroy infected plant material and avoid overhead watering.  When planting select infection resistant varieties. Practice crop rotation and add pot ash to the soil to decrease the plants venerability to the disease. Many species of fungus overwinter in fallen leaves, remove and destroy any litter under the plant.  

Winter Fusarium Leaf Disease in Turf Grasses can be minimised by aerating the soil, reducing thatch and avoid excessive nitrogen in the soil.


Chemical Control

Protective fungicides such as zineb or copper oxychloride should be sprayed at the first sign of infection and cuttings should be sprayed as they start to grow.

Note

Always read the label for registration details and direction of use prior to application of any chemicals.


72
Leaf Blight (General)
Various Leaf Blight Species

DISEASE

   NAME

     Leaf Blight (General)

     Various Leaf Blight Species



Description

A fungal problem that attacks the leaves and may spread to other parts.


Symptoms

Generally light to dark brown patches or spots appear on the leaves. These join together causing the leaf to become dried and withered. Infected leaves fall prematurely and dieback occurs in the stems. The overall effect is a sparsely foliaged tree.

On Carrots the fungus (Alternaria dauci) forms small spots that turn the affected area yellow. This can spread over the entire plant resulting in the death of the plant.


Leptosphaerulina Leaf Blight (Leptosphaerulina species) infects Turf Grass. Generally the leaves turn light brown and wilt and from a distance, appear as irregular yellow brown patches. The leaves initially die back from the tip forming water soaked lesions that extend to the sheath and give them a bleached appearance. The fruiting bodies develop on the dead material and the spores are distributed by wind. This fungus appears on many grasses but is only a significant problem for Agrostis palustris (Bent) particularly during hot humid conditions.


Palm Leaf Blight (Pestalotia palmarum) forms grey brown spots on the leaf blades and penetrates into the leaflet axil turning the leaflets brown and killing the frond. Spores appear in black masses on the upper leaf surface.  Another Palm Blight (Pestalotia palmicola) infects leaflet tips and progresses down throughout the frond turning it brown.


                 


Strawberry leaf blight (Phomopsis obscurans) infects the leaves primarily with a triangular shaped patch that is reddish turning to brown. It is also found on the petioles and stolons and the tiny black fruiting bodies occur on damaged areas.


The fungal Leaf Blight (Didymascella thujina) forms brownish raised spots on small leaves tuning them brown during late spring and giving them a scorched, dead appearance, eventually falling from the plant during autumn. Commonly found on large Thuja species, but may also attack smaller varieties.


Umbrella leaf blight (Alternaria panax) forms dark brown to black patches that enlarge and distort immature leaves.


Source and Dispersal

The spores originate from other infected trees and are dispersed by wind or infected nursery stock. Once a plant is infected it can moves rapidly throughout. Certain species of tree can tolerate infection.


Favoured Conditions

It prefers humid still conditions with warm temperatures and is particularly a problem in tropical areas.        


Affected Plants

A wide range of plants including vegetables, shrubs and trees may be affected.


Hibiscus species leaves are infected by Pellicularia koleroga a soil born fungus.


Leaf Blight (Pestalotia palmarum) infects several palms including Syagrus romanzoffianum and leaf blight (Pestalotia palmicola) attacks Phoenix, Syagrus and Washingtonia species.


Primula species are infected by the Leaf Blight (Phyllosticta prinulicola).


Syringa species are infected with the leaf blights (Cladosporium herbarum) and (Heterosporium syringae) which form large brown spots on the leaves that become dry and fall out.


Tilia species particularly young trees are in infected by the casual fungus (Cercospora microsora), which forms brown spots with darker borders, resulting in the death of the leaf.


Tsuga species are infected by the leaf blight (didymascella tsugae), which turns the leaves brown causing them to fall prematurely.


Non-chemical Control

On small trees remove and destroy the infected parts or in a nursery remove and destroy infected stock. Perennials and annuals should be removed and destroyed.  Remove any fallen leaves from around the base of the plant as they contain fruiting bodies.


Chemical Control

Plants may be sprayed at five day intervals with copper oxychloride if possible. Turf infected with Leptosphaerulina Leaf Blight may be sprayed with a broad spectrum fungicide.

Note

Always read the label for registration details and direction of use prior to application of any chemicals.


76
Fungi (General)
Various Fungal species

DISEASE

   NAME

     Fungi (General)

     Various Fungal species


Description

A fungus is a plant that lacks chlorophyll and conductive tissue. Generally they are made up of branched threads called 'hyphae' and collectively form a vegetative body called 'mycelium'. The fungus is small but the fruiting bodies can become very large up to 600mm across such as bracket fungi or mushrooms. Common fungi are mould and mildews. problem that attacks the roots causing them to rot.

Fungus can reproduce many ways but primarily it is asexually, simular to cuttings of a plant and often occurs with minute portions of the mycelium (spores) separating. The spores can be arranged in a structure such as a sporangia or pycnidia or develop without an enclosed structure called a "conidia". Either way the fungus propagates very rapidly.  Sexually reproduction occurs when two nuclei unite and form sexual fruiting bodies (zygospore).


Strelitzia reginae flower


Symptoms

Fungus attacks all the above or below ground level parts of the plant living within the tissue of the plant and are very small and not normally detected until the fruiting body appears. However parasitic types such as powdery mildew or rust are visible on the outer surface of the plant.

Fungi hyphae may be divided by cross walls and known as "septate" while others with no cross walls are known as "nonseptate". These are the fungi responsible for cell leakage as in rot.


Back Mold (Chalariopsis thielavioides) affects understocks of grafted Rosa species by inhibiting the development of callus. It is whitish-grey maturing to black and can be found in the pith of the rose stem.


Black Root Rot (Chalara elegans).This recently introduced fungal disease in Australia (1993) affect plants by blackening the root systems and turning leaves yellow or purple. It is difficult to identify specifically as other pathogenic root diseases and nutritional deficiencies have simular characteristics.  

The asexual spores are dispersed by wind or water. It is also transmitted on insects and in contaminated growing media or plants preferring humid moist conditions.

This fungus affects a wide range of ornamental plants including; annuals, perennials and shrubs. Examples are Begonia, Boronia, Camellia, Cyclamen, Fuchsia, Gerbera, Grevillea, Impatiens, Pansy, Petunia, Rosa species and Snapdragon.


Black Stem Rot (Pythium splendens) normally is a rot that occurs in cuttings turning the stem progressively black and shrunken. The leaves fall and the plant becomes stunted, eventually dieing.


Bleeding Necrosis (Botyosphaeria ribis) attacks and kills the inner wood causing the bark to split open and bleed sap giving it an oily appearance.


Blight (Endothia parasitica) is a serious pest of Castanea species, entering the twigs and small branches, and then progressively travelling throughout the tree killing it. It may form cankers on the base of the trunk or in the dead branches above with the amber coloured fruiting bodies pushing there way through the bark.


Copper Web ((Rhizoctonia crocorum). This fungal disease appears in defined patches causing the corms in the centre to become a black powdery mass. Corms on the outer ring of the patch that are partially infected forming a felty mass of violet threads on the corm scales. These threads extend into the soil and large sclerotia forms in the soil and on the corms. Healthy corms become infected from contaminated soil that contains mycelium and sclerotia.


Dry Rot (Phyllosticta concave) forms small circular spots that increase to a diameter of 30mm, and then becomes sunken as the cells collapse. The infected area develops minute black fruiting bodies.


Dutch Elm Disease (Ceratocystis ulmi) is a serious fungal problem of Ulmus species that initially causes yellowing then wilting of the leaves that turn brown and die. This may be seen on certain branches of the tree and on inspection under the bark the sapwood reveals brown streaks. A cross section of the affected branch displays round spots that are dark brown. This infection normally spreads quickly throughout, killing the tree in one to two seasons.


Dieback in Camellia (Glomerella cingulate) is a pathogenic fungus that infecting existing wounds such as leaf scars or mechanical damage, forming a sunken area (canker) that spreads around the stem causing die back. The affected plant has new shoots that are brown-black and the tips curl, forming a 'Shepard's Crook' appearance. The leaves also die but are persistent on the plant and the spores are found in soil or on other infected plants.


Curvularia Leaf Spot (Curvularia species) in Turf Grass. This is normally a secondary weak fungal infection that forms spots on the leaves that lengthens turning the leaves greyish. The leaf shrivels then dies and infected areas appear as weak patches in the turf. Preventive measures include minimising leaf wetness and excessive use of nitrogen fertiliser.


Fairy Rings Blue Couch     


Fairy Rings are a fungal problem in Turf Grass and is caused by several species including (Lycoperdon species), (Marasmius species) and (Tricholoma species). Rings appear in the turf as fruiting bodies or dead grass and as lush green foliage. The mycelia expand radially in the turf feeding on soil nutrients and organic matter with water present.


Under severs conditions the mycelia consume all available nutrients resulting in the death of the turf. Lush turf can result from a less developed infection, where the decomposing hyphal releases nitrogen. This available nitrogen may be beneficial to the turf but some forms of nitrogen are detrimental.


Leaf Blister (Taphrina coerulescens) appears as yellowish circular raised areas on the upper side and depressions on the underside of leaves, up to 15mm across. As the fungus spreads the leaf dies but remains attached to the tree and this infection is commonly found on Quercus species..


Leaf Blotch (Guignardia aesculi) forms small or large water soaked spots that are reddish with a bright yellow margin and form black fruiting bodies in the centre. The affected leaf and petiole have a scorched appearance before falling, found on Aesculus species


Grevillea robusta     Leaf Scorch


Leaf Scorch (Verrucispora proteacearum) is a fungal disease that infects leaves causing large parts of the leaf to turn grey-brown, giving the appearance that it has been singed by fire. Black fruiting bodies appear on the affected areas and the leaf soon withers then dies. New, mature leaves are affected during very wet periods towards the end of the branches and Grevillea and Hakea species are susceptible.


Melting Out (Helminthosporium vegans) forms bluish black spots with straw coloured centres on the leaves and may be found on the sheath, encircling it causing Foot Rot. It infects grasses particularly Poa pratensis. There is another fungus that is simular Helminthosporium Blight (Helminthosporium dictyoides) that infects Poa, Festuca and Agrostis species.


Pad decay (Aspergilus alliaceus) infects Cereus and Opuntia species and occurs at during periods of high temperature.  The yellow spores at the epidermal layer through wounds and germinate on mass causing the area to become soft and spongy. An anthracnose called Shot Hole is a similar forming brownish spots the turn grey, and then black destroying pads. Control methods include physically removing damaged pads and allowing the Sun to heal wounds.


Potato Gangrene (Phoma foveate) is a soil borne fungus that infects the roots during harvest primarly through wounds and develops during storage. The potatoes rot from the inside forming rounded depressions on the surface and have a strong odour of rotten fish.


Root Rot Fungi (Phymatotrichum omnivorum) and (Pellicularia filamentosa) cause the roots to rot and the plant suddenly wilts then dies.


Root Rot (Pythium debaryanum) forms water soaked dark brown streaks that affect all parts of the plant causing wilting then dieing. It infects Ranunculus species, it also infects cactus species by forming brown spotting and wilting that appears at the base of the plant then extends towards the top. It quickly spreads from plant to plant in collections and is controlled by avoiding over watering, excessive humidity and are using a sterilised soil when potting up.

This fungus also is responsible for damping off of seedlings in a glasshouse environment.


Spring Dead Spot     


Spring Dead Spot (Leptosphaeri species) is a fungal disease that infects Couch Grass. It first appears during autumn as pale bleaches areas up to 500mm (20in) wide and persists throughout winter. In spring the affected areas do not recover or recover slowly and on inspection the roots or rhizomes are rotted. Runners from the surrounding healthy turf will help with recovery and all signs of the problem disappear by mid summer.


Cactus species      Pachypodium species


Stem Rot (Helminthosporium cactivorum) forms well defined yellow lesions that mature into soft dark brown rot. It commonly infects Cactus species entering through the stomates or wounds. Heavily infected plants collapse and die.


Stem Rot or Basal Rot (Pellicularia rolfsii) is a soil borne fungus that infects the stem root junction and extends into the leaves. In orchids the leaves become discoloured, dry and detach from the base which is covered in a fungal growth that produces sclerotia. The sclerotia is whitish to yellow then becoming dark brown and can be viable for up to four years.


White Mold  (Ramularia desta f. odorati) occurs on both sides of the leaf and looks simular to powdery mildew but forms faint dull, reddish brown elongated spots on the leaf that may be depressed or along the margin where they have a watery appearance. Tufts of hyphae develop in the stomates.


Wilt (Ceratocystis fagacearum) causes leaves to curl then turn brown and the sap wood may also turn brown or black. Heavy infection may kill a tree within two seasons and is found on Quercus species and other ornamental trees.


Witches Broom may be a fungal problem that causes a proliferation of small axillary shoots to appear at the end of the branches. Little is known about this problem, though it affects a wide range of plants including Eucalyptus, Leptospermum and Pinus species.


Source and Dispersal

Fungus is found in the soil or on other infected plants and after releasing the spores, they are dispersed by wind or are transmitted in infected stock, insects and with splashing water.


Wilt is transmitted by infected root stocks, several species of insect and contaminated tools.


Dutch Elm Disease is transmitted by bark beetles such as (Scolytus multistriatus) and (Hylurgopinus rufipes). These beetles deposit eggs in the sapwood where the lava tunnel and pupate. The emerging beetles tunnel the bark and carry the fungus to fresh feeding sites on the tree. Infected beetles may also be transported to fresh sites in waist material.


Favoured Conditions

Prefers cool moist conditions with temperatures from 10º to 25ºC and is more common from autumn to spring when it is wet.    

        

Affected Plants

A wide range of plants and all parts can be infected by various fungal diseases. Bleeding Necrosis is found in Liquidambar species and Stem Rot or Dry Rot infects Cactus species such as Opuntia and Pelargonium.


Abies species are infected by several fungi that cause Leaf Cast which turn the needles yellow to brown then fall prematurely.


Abutilon species are infected by the Stem Rot (Macrophomina phaseolin) affecting the lower stems and is not commonly seen.


Achillea, Cuphea, Leucanthemum, Euphorbia species are infected by the Stem Rot (Pellicularia filamentosa) which enters through the roots and rots the base of the stem.


Alternanthera species are infected by the Leaf Blight (Phyllosticta amaranthi) which forms small brown spots on the leaves causing them to curl and die.


Aloe, Astrophytum, Copiapoa, Echinocactus, Espostoa, Ferocactus, Gymnocalycium, Kalanchoe and Schlumbergerera species are infected by Bipolaris Stem Rot (Bipolaris cactivora). This infection affects many cacti species causing rot in the stems with a blackish appearance.


Amelanchler is affected by the Witches Broom (Apiosporina collinsii).


Antirrhinum species are infected by the Blight (Phyllosticta antirrhini) that forms light brown spots on the upper-side of the leaf and on the stem. As the spots enlarge they turn greyish with black fruiting bodies in the centre, then become brown and killing the affected areas.


Begonia species are infected by the Stem Rot (Pythium ultimum) turning stems black then becoming soft and causing the plant to collapse. This is the same fungus that causes Damping-off.


Betula species are affected by the Leaf Blister (Taphrina bacteriosperma) which curls the leaves and forms reddish blisters.


Chamaedorea and other cain-like species are infected with Gliocladium Stem Rot (Gliocladium vermoseni) which forms a dark basil stem rot generally on damaged plants and produces orange-pink spores.  The mature leaves are first affected and eventually the stems or cains rot and die.


Crocus and Gladiolus species are infected by the Dry Rot (Stromatinia gladioli), which causes lesions on the corms and rots the leaf sheath.


Crocus, Iris, Tulipa, and Narcissus species are infected Copper Web ((Rhizoctonia crocorum).


Dianthus species are infected by Phialophora Wilt (Phialophora cinerescens) that causes the leaves to fade and plants to wilt. There is obvious vascular discoloration which is very dark. It is not found in Australia.


Erythrina x sykesii may be infected by the Root Rot Fungi (Phymatotrichum omnivorum).


Fern species are infected by Tip Blight (Phyllosticta pteridis). This blight produces ash-grey spots with purple brown margins and the fruiting bodies appear as black pimple like spots.  It is transmitted by air or moisture and in infected fronds become brown and die.  Control methods include sprang fungicide on leaves or reducing humidity and avoid wetting the fronds.


Forsythia species are infected by Stem Gall (Phomopsis species). It forms rounded growths along the stems causing them to die and look unsightly.


Gladiolus species are infected by Penicillium Rot of Corms (Penicillium gladioli). This disease forms deeply sunken reddish brown areas that become corky and produce a greenish fungal growth.


Hedera species are susceptible to several Fungal Leaf Spots including (Glomerella cingulate), (Phyllosticta concentrica) and (Ramularia hedericola). All of which cause yellowish spots that develop into dry brown blotches that kill the leaf.


Larix species are susceptible to Leaf Cast (Hypodermella laricis). This fungus attacks the needles and spur shoots turning them yellow at first then brown after which small black fruiting bodies appear on the leaves during winter.

There are several other fungi including (Cladosporium species) and (Lophodermium laricis) cause leaf blight or leaf casts.


Orchids such as Cattleya, Cymbidium, Cypripedium, Dendrobium, Epidendrum, Oncidium, Paphiopedilum, Phalaenopsis and Zygopetalum species are infected by Phomopsis Rot (Phomopsis species). This fungal problem forms a firm brown rot that appears on the leaves, pseudobulbs and rhizomes. The affected areas have yellow margins and the centre is covered in tiny black specks (fruiting bodies). Cattleya species are particularly susceptible. These plants are also infected by Psudobulb Rot (Mycolleptodiscus coloratus implicated). Dark spots appear on the pseudobulbs eventually causing extensive rot and killing the bulb.


Palms are infected by the fungus Butt Rot (Ganoderma sulcatum). The fungus entered the lower trunk normally as a result of mechanical damage (lawn mower). Symptoms include stunting of new growth and yellowing of the lower leaves. Fruiting bodies become evident at the base of the trunk. There is no effective control method and replanting in infected soil should be avoided.



Kikuyu Yellows     


Pennisetum clandestinum (Kikuyu) is susceptible to Kikuyu Yellows (Verrucalvus flavofaciens), thisis a water mould that infects the roots and causes them to rot. The infection extends up the stem and onto the leaves with yellow discolouration and can be limited to a small or large area up to 1m (3ft) wide.

It is found in warm temperate to sub tropical regions and dispersed in infected soil or plant material. There is no chemical control, nitrogen fertiliser masks the symptoms and complete fertiliser encourages stronger roots to fight the disease.


Pittosporum, Antirrhinum, Aquilegia, Echinops and Orchid species are infected by the Stem Rot or Basal Rot (Pellicularia rolfsii) commonly in the northern hemisphere and preferring humid glasshouse conditions.


Solidago species are infected by the fungal Scab (Elsinoe solidaginis) which covers the leaves and stunts the growth of the plant. Young plants may be killed.


Trillium species are infected by the stem rot (Pellicularia rolfsii) and ( Ciborinia trillii). This normally occurs in wet soils and is detrimental to the plants life.


Tsuga species are infected by Sapwood Rot or Butt Rot (Ganoderma lucidum) and (Coniophora puteana), which attacks the sapwood close to the bark, towards the base of the tree. Commonly killing the host.


Tulipa species are affected Blue Mold (Penicillium species) and the fungus (Rhizopus stolonifer) causing rot in the bulbs.


Vinca species are infected by the soil born Root Rot (Pellicularia filamentosa) which rots the stems and roots.


Viola species may be infected with the Scab (Sphaceloma violae) which attacks all parts of the plant including the seed capsule forming yellowish spots that turn brown and in leaves fall out. Stems and petioles can be girdled killing the upper part.

Viola species are also infected with the Stem Rot (Myrothecium roridum) which attacks the stems at ground level causing them to become dry and brittle. The leaves show symptoms by turning purplish-black and this fungus also infects Alcea and Antirrhinum species.


Non-chemical Control

Generally remove and destroy any infected plants or plant parts, when replanting, avoid using susceptible species for 3 years.  When growing crops space the plants to reduce the humidity and airflow and cultivate the soil to increase the drainage.  Remove weed growth from around the susceptible plants.

Avoid over watering the surrounding soil which encourages fungal development. In the case of trees remove any infected branches and heavily infected trees should be cut down and removed. This infected material should be disposed or burnt. Damaged trees should have the wounds dressed and sealed as a preventative measure particularly for Dieback in Camellia.


Deter Potato Gangrene by planting clean stock and be careful not to damage the crop when weeding. When harvesting the tubers choose a dryer period and be careful not to damage them.


     Fairy Rings


Fairy Rings in Turf are difficult to control and may appear or disappear sporadically. Cultural practice such as minimal thatch build-up, regular aeration and a reduction of organic matter spread on the turf will reduce infection.


Chemical Control

No suitable fungicides available, though drenching or spraying the soil with the fungicide dichloran helps control soil born fungi.

Note

Always read the label for registration details and direction of use prior to application of any chemicals.


74
Fusarium Wilt (General)
Various Wilt Species

DISEASE

   NAME

     Fusarium Wilt (General)

     Various Wilt Species


Description

Fusarium wilt is caused by specialised strains of the common soil fungus, Fusarium oxysporum.This fungus is microscopic, and that can live in the soil for many years and primarily attacks plants by entering through the roots. This affects corms, stems and leaves by moving through the water conductive tissue in the plant.

The specialised strains are known as formae speciales (f. sp. or special forms) and each f. sp. has a relatively narrow host range. For example Fusarium oxysporum f. sp. cubense causes Panama wilt of banana, F. oxysporum f.sp. vasinfectum infects cotton and F. oxysporum f. sp. lycopersici attacks tomatoes. Each strain is specific to that narrow host range and will not affect other hosts; this is important as it makes control options involving the planting of other plant species possible.  


Symptoms

Generally the plant wilts quickly during hot weather or lacks growth and vigour. Typically the symptoms are often indistinguishable from drought stress. Leaves yellow and wither or stems split open near the base. Individual branches may collapse and die or entire plants perish. One of the distinctive symptoms is a one-sided yellowing and death of a leaf or branch. Corms become rotted with open wounds and affected stems, when split open are yellowish. There are variations in the symptoms depending on the type of plant but generally plants wilt, turn brown and die.



Basal Rot (Fusarium oxysporum) infects Tulipa and Crocus species by turning leaves reddish, which wilt and normally die. It also affects the bulbs by forming few roots and rotting the base, turning them dull white.

This disease also infects cactus such as Cereus species and Zygocactus truncatus causing soft black rot that appears at the tips of the plant then spread rapidly downwards.  It is particularly virulent in hot beds, under glass. The infected plants cannot be cured and should be disposed off site.


Fusarium Patch, Bent<      Fusarium Patch, Blue Grass


Other Fusarium (or related species) diseases

Fusarium Patch (Microdochium nivale formerly known as Fusarium nivale) is also known as Crown or Root Rot and infects warm or cool season Turf Grasses. The fungus infects the roots or rhizome turning them black, sometimes with a pink ting. During hot periods light green to brown patches appear in the turf as the damaged roots are unable to supply water to the leaves. This diseases is not a significant disease in Australia

Minimise irrigation of infected areas to reduce spread of infection as the spores are distributed in water.


Root Rot (Fusarium verticillioides) is a seed born fungus that infects Strelitzia reginae. It also forms a Leaf Spot in Dracaena species where rounded or irregular yellowish to reddish spots with a pail green border form. These are found on immature leaves and heavy infections may rot the crown.


Source and Dispersal

The fungal spores are dispersed by wind from infected plants or parts of plants. It is also dispersed by splashing water or contaminated stock or the movement of soil. With Fusarium wilt of canary island date palms the transfer of infected plant material on chain saws has been shown to be an important way in which the fungus can be transmitted from one tree to another. It is essential in these cases to sterilise the chain and chain bar between trimming the fronds on each tree.


Fusarium oxysporum produces three types of species, macroconidia, microconidia and chlamydospores. The latter two spore types play very important roles in the disease. The microconidia move in the vascular system of the plant and collect at the sieve plates in the xylem of plant. Here they germinate and grow and eventually block the sieve plate thus stopping the flow of water in the plant. The chlamydospores are specialised survival spores that persist in roots and in the soil for very long periods of time, thus allowing the fungus to survive periods when there are no susceptible plants present.


Favoured Conditions

The fungus prefers warm humid climates and is commonly seen in coastal regions and may continue living in soil for many years.


Affected Plants

Many species are infected including tomatoes, carnation, cucurbits and freesia. Die Back (Fusarium solani var. martii) causes damage to the branches of Ilex species. Fusarium also attacks palm species such as Phoenix causing wilt.

Albizia and Alternantera species are infected by the wilt (Fusarium oxysporum var. periciosum ) that causes the leaves to wilt, shrivel and die. This infection also extends to the branches and eventually may kill the tree.


Callistephus species are attacked by the wilt (Fusarium oxysporum f. sp. callistephi) which infects the vascular bundles turning brown, particularly on one side of the stem causing the plant to become stunted, eventually dieing.


Dendranthema and Thymus species are susceptible to the Stem Rot (Fusarium species) and (Pellicularia filamentosa) which rots the base of the stem.


Dianthus species are infected by Bud Drop (Fusarium poae). This fungus rots flower buds prior to opening and infected flowers open with distorted, unattractive petals. The control of thrips is essential. Dianthus species are also infected by Wilt (Fusarium oxysporum) that causes yellowing and wilting of the lower leaves, normally on one side. The infection causes stems to rot with vascular discolouration.


                  Fusarium Wilt Image by B. Sonsie

                  Image by Dr Brett Summerell


The only palms susceptible to Fusarium wilt are Phoenix species, especially Phoenix canariensis and juvenile Washingtonia filifera. Initially the leaflets turn yellowish then brown, on mature fronds causing them to die. Eventually only tufts of new growth remain the plant. As the infection continues the plant collapses and dies.


Fusarium Patch


Turf grasses are infected by Fusarium Patch (Microdochium nivale). Plants include; Digitaria didactyla (Blue Couch), Cynodon species (Couch), Eremochioa ophiuroides (Centipede Grass), Festuca species, Paspalum vaginatum (Salt Water Couch), Pennisetum clandestinum (Kikuyu), Poa species (Bluegrass), Stenotaphrum secundatum (Buffalo) and Zoysia species. Generally the fungus produces small dead circular patches in the lawn.


Non-chemical Control

Fusarium diseases are extremely difficult diseases to control primarily because of a lack of a chemical control option and because the fungus is able to persist in soil for long periods of time. Crop rotation to non susceptible plants is advised and consequently it will be important to have the disease accurately diagnosed to ensure the identity of the pathogen and the possible rotation options.

Remove and destroy any infected plants. In the case of large trees or palms, it will be necessary to remove the infected plant, this not only means cutting down the above ground parts, but also removing the roots of the plant. These will then need to be removed and destroyed.

Fusarium Patch can be minimised by reducing thatch and aerating the soil regularly or improving the drainage.


Chemical Control

There are no effective chemical control options currently available for plant diseases caused by Fusarium species.

Note

Always read the label for registration details and direction of use prior to application of any chemicals.


Amendments by

Dr Brett Summerell
Director Science and Public Programs
Royal Botanic Gardens Trust, Sydney


Inflorescence
Leaf to 2.5m (7 ft) long
Rounded crown
Bark and leaf bases

Plant Photo Gallery - Click thumbnails to enlarge

Climate zone

This Plant tolerates zones 9-12

Average Lowest Temperature : 10º C 50º F

USDA : 9, 10, 11, 12

This USDA (United States Department of Agriculture) hardiness zone chart can be used to indicate a plant’s ability to withstand average minimum temperatures. However, other factors such as soil type, pH, and moisture, drainage, humidity and exposure to sun and wind will also have a direct effect on your plant’s survival. Use this chart only as a guide, always keep the other factors in mind when deciding where, when and what to plant.

A plant's individual USDA zone can be found in the Plant Overview.

Climate Description

Sub-tropical
This zone has ample rain with coastal breezes during summer and periodic high humid summer temperatures with mild winters.
Frosts and droughts rarely occur.

Plant growth

Tropical and some warm temperature native and exotic plants grow well.

Glossary

Dictionary Growth Habit
Leaf Type Botanic Flower Description
Leaf Shape Flower Inflorescence
Leaf Arrangement Fruit Type
Leaf Margin Bark Type
Leaf Apex And Bases Flower Description