The flora of Hatila Valley National Park and its close environs (Artvin)

Ozgur Eminagaoglu

This study was aimed to determine floristic content of Yukarımaden and Aşağımaden villages and their surroundings in Artvin between June and September 2015. It is located in the northeast of the Eastern Black Sea Region and is taken place in Colchic province of Euro-Siberian floristic area of Holarctic Region and lies between 400 and 2830 m. As a result of this study, total 566 plant taxa belong to 85 families and 358 genera were determined.7 of them belong to Pteridophyta, the remaining 559 taxa were Magnoliophyta which included 7 taxa from Pinidae, 1 taxa from Gnetidae and 551 taxa from Magnoliidae. The largest families recorded were as follows: Poaceae 38 taxa, Lamiaceae 38 taxa, Rosaceae 38 taxa, Asteraceae 36 taxa, Fabaceae 32 taxa, Brassicaceae 26 taxa, Caryophyllaceae 22 taxa, Boraginaceae 22 taxa, Ranunculaceae 20 taxa and Apiaceae 16 taxa. The percentage of phytogeographical origins of 227 taxa (40.1%) was determined. The distribution of the taxa according to the phytogeographical regions was as follows: 130 taxa (22.9%) Euro-Siberian, 83 taxa (14%). Irano-Turanian, 14 taxa (2.4%) Mediterranean, 339 taxa (59.9%) cosmopolit. In the study area, 57 rare plant taxa were identified, 18 of which were endemic. Endemism is 3.2% and included 18 endemic taxa. The endemic and rare non-endemic plants in the study area have been indicated to IUCN threat catagories. Also, one species has been identified under the BERN and CITES contracts in the field.

This study has been carried out for determination and contribution to the flora of Kamilet Valley where has quite rich floristic composition during 2011-2016. The research area under review is taken place in Colchic province of Euro-Siberian floristic area of Holarctic and is found in the A8 square according to grid system of Davis. On the other hand, the study area lies between 300 and 3264 m. elevation above the sea level. In this study, total 651 plant taxa belong to 107 families and 361 genera were determined. 41 of them belong to Pteridophyta, the remaining 610 taxa were Spermatophyta which included 5 taxa from Gymnospermae and 605 taxa from Angiospermae. Angiospermae also included 504 taxa from Dicotylodonae and 101 taxa from Monocotyledonae. The largest families recorded were as follows: Asteraceae 79 taxa, Poaceae 44 taxa, Rosaceae 38 taxa, Fabaceae 30 taxa, Brassicaceae 28 taxa, Lamiaceae 28 taxa, Caryophyllaceae 21 taxa, Ranunculaceae 20 taxa, Boraginaceae 18 taxa and Apiaceae 17 taxa. The percentage of phytogeographical origins of 378 taxa (%58.07) was determined. The distribution of the taxa according to the phytogeographical regions was as follows: 326 taxa (86.24%) Euro-Siberian, 19 taxa (5.03%) Irano-Turanian, 5 taxa (1.32%) Mediterranean, 28 taxa (7.41%) cosmopolit. The life form spectrum of the taxon was as follows: Hemicryptophytes 253 taxa, Cryptophytes 196 taxa, Phanerophytes 46 taxa, Chamaephytes 68 taxa, Therophytes 86 taxa and Vascular Parasites 2 taxa. Endemism is 2.92% and included 19 endemic taxa. The endemic and rare non-endemic plants in the study area have been indicated to IUCN threat catagories.

This paper describes a floristic study on Bağlıca Campus area of Başkent University in Ankara province. 779 vascular plant specimens were collected between 2008 March and 2009 August in 26 field trips. 373 species, 382 taxa of natural plants belonging to 216 genera and 51 families were determined, which of 63 taxa are endemic to Turkey (16.5%). Except for these, 45 taxa are detected as cultivated plants. The distribution of the phytogeographical elements is as follows: Irano – Turanian 119 taxa (31.2%), Mediterranean 20 taxa (5.2%), Euro – Siberian 15 taxa (3.9%), pluriregional or phytogeographically unknown 228 taxa (59.7%). As a result of present paper, the three largest families are Asteraceae with 52 taxa (13.6%), Fabaceae with 44 taxa (11.5%) and Lamiaceae with 33 taxa (8.6%). The largest three genera are Astragalus with 19 taxa (5.0%), Salvia with 11 taxa (2.9%) and Centaurea with 10 taxa (2.6%). The results of our research are compared with the studied neighbouring areas by m...

Turk J Bot 27 (2003) 1-27 © TÜB‹TAK Research Article The Flora of Hatila Valley National Park and its Close Environs (Artvin) Özgür EM‹NA⁄AO⁄LU Kafkas University, Faculty of Forestry, Department of Forest Botany, 08000, Artvin - TURKEY eminagaoglu@hotmail.com Rahim ANfi‹N Black Sea Technical University, Faculty of Forestry, Department of Forest Botany, 61080, Trabzon - TURKEY rahimansin@hotmail.com Received: 11.10.2000 Accepted: 11.07.2002 Abstract: This investigation was carried out between 1994 and 1997 in order to contribute to knowledge on Artvin’s flora. The Artvin area reflects the eastern part of the Black Sea region’s rich floristic composition. The research area under review is in the Colchic province of the Euro-Siberian floristic area in the Holarctic region and is found in the A8 square according to grid system of Davis. In the studied area, 769 taxa belonging to 374 genera and 95 families were determined. Of the 769 taxa, 26 Pteridophytes and 743 Spermatophytes were detected. Spermatophytes also contained 10 Gymnospermae and 733 Angiospermae taxa. The largest five families are Compositae with 73 taxa (9.5%), Leguminosae with 67 taxa (8.7%), Rosaceae with 50 taxa (6.5%), Cruciferae with 49 taxa (6.4%) and Labiatae with 42 taxa (5.5%). The largest genus is Trifolium L. (14 taxa) and the second is Astragalus L. (13 taxa). The distribution of the taxa according to the phytogeographical regions was as follows: 35.6% EuroSiberian, 6.9% Irano-Turanian, 2.2% Mediterranean and 55.3% unknown. Endemism is 7.4% and included 57 endemic taxa. Key Words: Flora, National Park, Hatila Valley, Artvin. Hatila Vadisi Milli Park› ve Yak›n Çevresinin Floras› (Artvin) Özet: Bu çal›flma, 1994-1997 y›llar› aras›nda, Do¤u Karadeniz Bölgesinin kuzeyinde, oldukça zengin floristik içeri¤e sahip Artvin’in floras›na katk›da bulunmak amac›yla gerçeklefltirilmifltir. Araflt›rma alan›, Davis taraf›ndan uygulanan grid sistemine göre A8 karesinde yer almakta olup, Holarktik fitoco¤rafik bölgenin, Avrupa-Sibirya flora alan›n›n Kolflik kesimde yer almaktad›r. Araflt›rma sonuçlar›na göre; 95 familya ve 374 cinse iliflkin toplam 769 takson kaydedilmifltir. Bu taksonlar›n 26’s› Pteridophyta, 743’ü Spermatophyta bölümüne iliflkindir. Spermatophyta bölümünde, 10 takson Gymnospermae ve 733 takson Angiospermae’dir. Araflt›rma alan›nda en fazla türe sahip befl familya s›ras›yla Compositae 73 takson (% 9.5), Leguminosae 67 takson (% 8.7), Rosaceae 50 takson (% 6.5), Cruciferae 49 takson (% 6.4) ve Labiatae 42 takson (% 5.5). En büyük cins Trifolium L. (14 takson), ikincisi ise Astragalus L. (13 takson)’tur. Taksonlar›n, fitoco¤rafik bölgelere göre da¤›l›mlar› ise flöyledir: Avrupa-Sibirya % 35.6, ‹ran-Turan % 6.9, Akdeniz % 2.2 ve orijini bilinmeyenler % 55.3’tür. Taksonlar›n 57 adeti (% 7.4) endemiktir. Anahtar Sözcükler: Flora, Milli Park, Hatila Vadisi, Artvin. Introduction The aim of this study was to contribute to knowledge about Artvin’s flora, as this area reflects the eastern part of the Black Sea region’s rich floristic composition. In nearby areas, some floristic and vegetation studies were carried out by Anflin (1979, 1983, 1984), Düzenli (1988), Vural (1996), and Küçük (1998). The study area is in the A8 square according to Davis (1971)’s grid system and within the Colchic province of the Euro-Siberian floristic area in the Holarctic region (Zohary, 1973). The altitude of the area is between 170 and 3224 m. The area under review has a length of 24.4 km and extends from the Çoruh River (170 m) to Mount Kurt (3224 m) and has a width of 14.3 km from Alacada¤ (2844 m) to Mount Keçi (2047 m). The study area includes a sharp valley water catchment area that has about 3000 m elevation differences. The area is situated between lat 41°3'-41°13'30'' N and long 41°31'- 41°48' E and is some 25 000 ha in area (Figure 1). 1 The Flora of Hatila Valley National Park and its Close Environs (Artvin) Çoruh River F›st›kl› ARTV‹N Hatipo¤lu Figure 1. E N Topographical map of the study area. S W Gökçeyurt Bakacak Arganet Gölgeli Meydanlar Cerat Hill (1812 m) Demirci Alacada¤ Mount Keçi (2844 m) Köklü (2047 m) Kanac›k Davütet Mount Otluca Direkli (2887 m) Perikaya Hill (2296 m) K›l›çl› Seçinge Hill (1938 m) Mount Kurt (3224 m) Ar›p›nar 1 A 2 3 4 5 6 7 8 9 fiaflk›rat B C Tuzlu Hill (3060 m) Climatological data were obtained from Artvin Meteorological station. The annual average rainfall is 661.03 mm and the average warmth is 12.3 °C in the area (Meteoroloji Bülteni, 1995). The climate type of the area is semi-humid and the main vegetation type of the area is humid forest (Çepel, 1966). The rainfall regime of the study area is “winter-spring-autumn-summer” (Central Mediterranean) of Mediterranean origin. The region is in line with the rainfall-heat formula (Q = 89.4) and a low precipitation/cold bioclimate type is dominant (Akman, 1999). Upper Cretaceous aged volcanic blocks (basalt, gabbro and peritoneum) form the main geologic structure. There are seven large soil groups in the study area, namely, brown forest soils, noncalcareous brown forest soils, chestnut soils, alluvial soils, colluvial soils, red-yellow podzolic soils and high mountain meadow soils. The most widespread is the brown forest soils type (Topraksu Genel Müdürlü¤ü, 1975). 2 Materials and Methods The materials of this study included some 2000 plant samples collected from the research area between 1994 and 1997. Efforts were made to collect both flowering and fruiting specimens. The plant samples were prepared according to established herbarium techniques. The Flora of Turkey (Davis, 1965-85; Davis et al., 1988) and other sources (Komarov, 1934-60; Grosheim, 1939-1967) were utilised in the identification of the plant samples. This identification was carried out in the Herbarium of the Forestry Faculty, Black Sea Technical University, Trabzon (KATO) and ISTO (‹stanbul) herbarium. All plant samples were kept in KATO. The list of plants is presented according to the sequence given in the Flora of Turkey (Davis, 1965-85). The taxa we were unable to collect during this study but were cited in the Flora of Turkey were added to the floristic list. Since the research area was wholly situated within the A8 grid square, this information was not Ö. EM‹NA⁄AO⁄LU, R. ANfi‹N, repeated after the taxa names. After the name of every species in the floristic list, the geographical location, habitat of plant, altitude, collection date, collector name and number, endemism, IUCN threat categories, and phytogeographical region are given. Author abbreviations of plant names are given according to Brummitt and Powell (1992). Since 19 taxa defined as new records for the A8 square in this manuscript were published before (Anflin et al., 1997), only the names of these 19 taxa are given in this study. The endemic and rare non-endemic plants in the study area have been classified according to IUCN threat categories (Ekim et al., 2000). Abbreviations The abbreviations used in the text and in the floristic list are as follows: Cosm: Cosmopolitan, DD: Data Deficient, E.: East, EN: Endangered, Euro-Sib: EuroSiberian element, Eux.: Euxine element, Hyr-Eux: Hyrcano-Euxine element, Ir-Tur: Irano-Turanian element, LR: Lower Risk, (lc: least concern, cd: conservation dependent, nt: near threatened), m: metre, Medit.: Mediterranean element, mt: mountain, ÖE: Özgür Emina¤ao¤lu, and VU: Vulnerable. Results and Discussion In this study, a total of 769 taxa from 87 families and 324 genera were identified. Twenty-six of them belonged to Pteridophyta, and the remaining 743 taxa were Spermatophyta, which included 10 taxa from the Table 1. Gymnospermae and 733 taxa from the Angiospermae. The Angiospermae also included 644 taxa from the Magnoliopsida and 89 taxa from the Liliopsida. The dispersion of the plant taxa that were defined in the study area according to the large taxonomical groups is shown in Table 1. After the identification of all taxa, the phytogeographical distributions of 344 taxa were determined. The percentage of phytogeographical origins of 344 taxa was 274 (35.6%) Euro-Siberian, 53 (6.9%) Irano-Turanian, and 17 (2.2%) Mediterranean. Twentynine taxa are cosmopolitan (3.8%). The rest of the 396 taxa (51.5%) are either multi-area elements or single elements which have not yet been accepted as members of the phytogeographical area (Table 2). Euro-Siberian elements were higher in ratio when the phytogeographical distribution of taxa was compared with the other elements in the study area. The abundance of Euro-Siberian elements (35.6%) and widespread presence of taxa such as Picea orientalis (L.) Link, Abies nordmanniana (Stev.) Spach. subsp. nordmanniana, Pinus sylvestris L. (Pinaceae), and Alnus glutinosa (L.) Gaertn subsp. barbata (C.A.Mey.) Yalt. (Betulaceae) can be attributed to the high influence of the Colchic province in the study area. However, the contribution of the IranoTuranian (6.9%) and Mediterranean (2.2%) elements to the flora of the area cannot be overlooked. As seen in Table 2, the results of our study area were in agreement with those of neighboring areas because it is in the same phytogeographical region. The percentage of The dispersion of taxa according to the large taxonomical groups. Number of Families Number of Genera Number of Taxa Pteridophyta Gymnospermae Dicotyledones Monocotyledones 12 4 71 8 15 6 305 48 26 10 644 89 Total 95 374 769 Table 2. Distribution of species in phytogeographical regions and comparison with studies carried out in nearby areas. Number of taxa (%) Euro–Sib. Ir –Tur. Medit. Cosm. Pluriregional Present study 35.6 6.9 2.2 3.8 51.5 Küçük (1998) 34.4 7.9 1.1 1.6 55.0 Düzenli (1988) 50.5 7.5 5.0 1.0 36.0 3 The Flora of Hatila Valley National Park and its Close Environs (Artvin) Mediterranean elements in our study was higher than that stated by Küçük (1998). A possible explanation is that our study area is under the influence of the Mediterranean climate and a local Mediterranean climate occurs in some areas close to the Black Sea region. The flora of the study area mainly resembles Caucasian flora, but it also shares similarities with Japanese and North American flora. A comparison of families in terms of the largest number of species found in this study and to previous studies carried out in nearby regions is given in Table 3. As seen in Table 3, the results of this article were also in agreement with the other similar studies. Differences in several taxa might be derived from the results of dissimilarities in climate and habitat. The largest families in terms of number of genera are Compositae (39), Cruciferae (26), Gramineae (23), Rosaceae (22), Leguminosae (19), Labiatae (19), Boraginaceae (15), Ranunculaceae (14), Umbelliferae (12), Liliaceae (11), Scrophulariaceae (11), and Caryophyllaceae (11). The richest families in terms of number of taxa are Compositae (73), Leguminosae (67), Rosaceae (50), Cruciferae (49), Labiatae (42), Gramineae (38), Scrophulariaceae (32), Ranunculaceae (32), Caryophyllaceae (28), Boraginaceae (25), and Liliaceae (21). The richest genera in terms of the number of taxa are Trifolium L. (14), Astragalus L. (13), Ranunculus L. (10), Veronica L. (10), Campanula L. (10), Geranium L. (8) and Vicia L. (8). Table 3. Endemism was 7.4% and included 57 endemic taxa. The endemism ratio of the study area is very low compared with the average endemism ratio (34.5%) of the Flora of Turkey in volume 11 (Güner et al., 2000). The reasons for the low endemism ratio of the study area may lie in the similarities between Caucasian flora and the studied area. Pseudomacchie, forest, subalpine, alpine and stream are the main vegetation types in the area (Anflin et al., 2000). Forest vegetation is the most common formation type in this area. In conclusion, the study area is mainly under the influence of a semi-humid climate, which is suitable for growing Euro-Siberian, Euxine, HyrcanoEuxine and Colchic elements. Pseudomacchie vegetation was distributed throughout the area, especially at altitudes of 200-600 m. There are a few Pinus pinea L. (Pinaceae) stands distributed sporadically at the junction of the Hatila valley and the Çoruh River named F›st›kl› village (formerly Naflviye). On the other hand, it has been questioned whether Pinus pinea L. stands here are natural distribution or plantation. In the stands of Pinus pinea L. in F›st›kl› village, many herbaceous and woody plants specifically growing in the Mediterranean region are located in the transition zone between the Euro-Siberian and Mediterranean regions. Anflin et al. (1997) believed that the Pinus pinea L. stands had distributed indigenously because of the many other herbaceous and woody Mediterranean taxa. They are as follows: Trachomitum venetum (L.) Woodson subsp. sarmatiense (Woodson) Avet. (Apocynaceae), Ostrya carpinifolia Scop. (Betulaceae), Cistus creticus L., C. salviifolius L. Comparison of the 10 families containing the most species in studies conducted in nearby regions. Number of taxa (%) Families Compositae Leguminosae Rosaceae Cruciferae Labiatae Gramineae Scrophulariaceae Caryophyllaceae Boraginaceae Liliaceae Others 4 Present study Anflin (1979) Düzenli (1988) Küçük (1998) 9.5 8.7 8.9 8.1 14.0 8.1 10.2 4.5 6.5 6.4 5.5 4.9 4.2 3.7 3.3 2.7 44.6 5.6 3.6 4.7 4.0 3.9 2.9 1.9 2.4 54.1 6.0 3.8 4.7 4.9 3.4 5.2 2.5 2.0 45.4 6.8 2.7 6.0 3.9 3.7 2.7 2.1 2.4 55.0 Ö. EM‹NA⁄AO⁄LU, R. ANfi‹N, (Cistaceae), Olea europaea L. var. sylvestris (Mill.) Lehr, Jasminum fruticans L. (Oleaceae), Cotinus coggygria Scop., Rhus coriaria L. (Anacardiaceae), Silene italica (L.) Pers. (Caryophyllaceae), Trifolium campestre Schreb (Legüminosae), Euphorbia paralias L., E. peplis L. (Euphorbiaceae), Erodium malacoides (L.) L'Hérit. (Geraniaceae), Sideritis montana L. subsp. montana (Labiatae). Botanists have been interested in F›st›kl› village because of its rich flora. This richness depends on the climatic variety and geographical structure of the area. Many botanists have studied the flora, vegetation and plant-geography of the surrounding area including, Mirov (1967) and Zohary (1973). Although the economy of the area depends on agriculture and forest products, there is no visible vegetation destruction in the area. The exception to this is the roadsides. As a result, the Hatila Valley National Park has a very rich flora mainly because of its different climates and habitats. Unfortunately, COMINCO runs a gold mine in this area and has been destroying the flora. Therefore, it is important that this area be protected as soon as possible. Being a detailed floristic study, this report may be a significant basis to measure possible variations in the future due to economic, industrial and pollution effects, and the most evident contribution is to the flora of Turkey. References Akman Y (1999). ‹klim ve Biyoiklim (Biyoiklim Metodlar› ve Türkiye ‹klimleri). Ankara: Kariyer Matbaac›l›k. Anflin R (1979). Trabzon-Meryemana Araflt›rma Orman› Floras› ve Saf Ladin Meflcerelerinde Floristik Araflt›rmalar. Trabzon: Karadeniz Gazetecilik ve Matbaac›l›k. Anflin R (1983). Türkiye'nin flora bölgeleri ve bu bölgelerde yay›lan asal vejetasyon tipleri (The Floristic Regions and the Major Vegetation Types of Turkey). Karadeniz Teknik Üniversitesi Orman Fakültesi Dergisi 6 (2): 318-339. Anflin R (1984). Artvin-Atila (Hatila) vadisinin bitki örtüsü ve bu örtünün filogenetik özellikleri (Vegetation of the Atila Valley in Artvin Area and Its Filogenetical Features). Karadeniz Teknik Üniversitesi Orman Fakültesi Dergisi 7(1): 84-89. Anflin R, Emina¤ao¤lu Ö, & Yüksek T (1997). Research on the Flora and Soil Properties of the Pinus pinea L. Stands Near F›st›kl› Village in Artvin. XI. Dünya Ormanc›l›k Kongresi, Antalya 13-22 Ekim 1997, XI. Dünya Ormanc›l›k Kongresi Tebli¤leri, Cilt 5, Antalya. Anflin R, Özkan ZC, Abay G, & Emina¤ao¤lu Ö (1997). New floristic record from A8 (Artvin) OT Sistematik Botanik Dergisi 4(1): 95-98. Anflin R, Özkan ZC, Emina¤ao¤lu Ö (2000). Artvin-Atila (Hatila) Vadisi Milli Park›n›n Vejetasyon Yap›s›na Genel Bir Bak›fl. Artvin Orman Fakültesi Dergisi (Journal of Artvin Forestry Faculty of Kafkas University) 1 (1): 59-71. Brummitt RK & Powell CE (eds) (1992). Authors of Plant Names. Kew: Royal Botanic Gardens, Kew. Çepel N (1966). Orman Yetiflme Muhiti Tan›m›n›n Pratik Esaslar› ve Orman Yetiflme Muhiti Haritac›l›¤›. ‹stanbul: Kutulmufl Matbaas›. Davis PH, Harper PC, & Hedge IC (1971). Plant Life of South West Asia. Edinburgh: The Botanical Society of Edinburgh. Davis PH (1965-85). Flora of Turkey and the East Aegean Islands, Vols. I-IX, Edinburgh: Edinburgh University Press. Davis PH, Mill RR & Tan K (1988). Flora of Turkey and the East Aegean Islands, Vol. X, Supplement. Edinburgh: Edinburgh University Press. Düzenli A (1988). Neuve aux Sintaxons Phytosociologiques pour la Vegetation de Turquie. Ecology Mediterranean, 14, 1(2): 143148. Ekim T, Koyuncu M, Vural M, Duman H, Aytaç Z, & Ad›güzel N (2000). Türkiye Bitkileri K›rm›z› Kitab›, E¤relti ve Tohumlu Bitkiler (Red Data Book of Turkish Plants, Pteridophyta and Spermatophyta). Ankara: Bar›flcan Ofset. Grossheim AA (1939-1967). Flora Kavkaza. Vol. 1-7, Baku and Leningrad. Güner A, Özhatay N, Ekim T, & Bafler KHC (2000). Flora of Turkey and the East Aegean Islands (Supplement-II). Vol. XI. Edinburgh: Edinburgh University Press. Komarow, VL (ed.) (1934-78). Flora of the USSR. Leningrad. (English translation by IPST, Vol. I-XXIV, Jerusalem: Israel). Küçük M (1998). Kürtün (Gümüflhane)-Örümcek Ormanlar›n›n Floras› ve Saf Meflcere Tiplerinin Floristik Kompozisyonu. T.C. Orman Bakanl›¤› Do¤u Karadeniz Ormanc›l›k Araflt›rma Enstitüsü, DKOA Yay›n No :5, Teknik Bülten No: 5, Trabzon: ‹nk›lap Matbaac›l›k Ltd. fiti. Meteoroloji Bülteni (1995). T.C. Baflbakanl›k Devlet Meteoroloji ‹flleri Genel Müdürlü¤ü Artvin Meteoroloji ‹l Müdürlü¤ü 1948 - 1980 Verileri. Artvin. Mirov NT (1967). The Genus Pinus. New York: The Ronald Press Company. Topraksu Genel Müdürlü¤ü (1975). Artvin ‹li Toprak Kayna¤› Envanter Raporu. Ankara. Vural M (1996). Rize’nin Yüksek Da¤ Vejetasyonu. Turk J Bot 20: 83102. Zohary M (1973). Geobotanical Foundations of the Middle East. Vols I-II, Amsterdam: Gustav Fischer Verlag. 5 The Flora of Hatila Valley National Park and its Close Environs (Artvin) APPENDIX The sequence of families, genera and species in the following list follows the system adopted in the Flora of Turkey. PTERIDOPHYTA BLECHNACEAE Pteridium aquilinum (L.) Kuhn Blechnum spicant (L.) Roth Saçinge hill, open forest, roadside, 1950 m, 13.05.1995, ÖE 1103. Tafll›ca village, Demirci, shady bank, 1250 m, 12.07.1997, ÖE 1842. EQUISETACEAE Cosm. POLYPODIACEAE Equisetum hyemale L. THELYPTERIDACEAE Polypodium vulgare L. subsp. vulgare Karada¤ hill, streamside, 1555 m, 14.04.1995, ÖE 1100. E. ramossimum Desf. Tafll›ca village, K›l›çl›, moist meadow, 1750 m, 10.06.1997, ÖE 1831. E. fluviatile L. Tafll›ca village, Köklü, streamside, 1800 m, 04.06,1997, ÖE 1832. E. palustre L. Tafll›ca village, Köklü, roadside, moist soil, 1400 m, 13.09.1995, ÖE 1101. E. arvense L. Tafll›ca village, Kanac›k, waste place, 1750 m, 21.05.1997, ÖE 1833. E. telmateia Ehrh. Tafll›ca village, Köklü, streamside, 1250 m, 10.06.1997, ÖE 1834. LYCOPODIACEAE Lycopodium selago L. Tafll›ca village, Bakacak, rocky place, 1200 m, 21.05.1997, ÖE 1835. L. clavatum L. Tafll›ca village, Ar›p›nar, shady bank, 1900 m, 02.06.1997, ÖE 1836. L. alpinum L. Tafll›ca village, fiaflk›rat plateau, alpine slope, 2200 m, 10.06.1997, ÖE 1837. OSMUNDACEAE Osmunda regalis L. Cerat hill, open Picea Dietr. forest, moist soil, 1250 m, 21.05.1995, ÖE 1102. ADIANTHACEAE Adiantum capillus-veneris L. Tafll›ca village, Direkli, stony slope, 1650 m, 12.06.1996, ÖE 1838. PTERIDACEAE Pteris cretica L. Tafll›ca village, Arganet, stony slope, 1200 m, 11.07.1996, ÖE 1839. 6 HYPOLEPIDACEAE Thelypteris limbosperma (All.) Fuchs Tafll›ca village, Meydanlar, in the Picea Dietr. forest, 1400 m, 06.06.1995, ÖE 1104. ASPLENIACEAE Asplenium trichomanes L. Tafll›ca village, Gökçeyurt, rocky and stony slope, 970 m, 13.07.1995, ÖE 1106. A. adiantum-nigrum L. Hatipo¤lu rocky and stony slope, 550 m, 13.07.1995, ÖE 1105. A. septentrionale (L.) H.Hoffm. F›st›kl› village, rocky and stony slope, 1930 m, 13.07.1995, ÖE 1107. Ceterach officinarum DC. F›st›kl› village, pathside, rocky and stony slope, 300 m, 06.04.1995, ÖE 1108. Phyllitis scolopendrium (L.) Newman Tafll›ca village, Gölgeli, open area, stony and moist soil, shady area, 1300 m, 15.07.1995, ÖE 1109. Tafll›ca village, Arganet, roadside, rocky area, 1450 m, 16.05.1995, ÖE 1113. Cosm. SPERMATOPHYTA GYMNOSPERMAE PINACEAE Abies nordmanniana (Stev.) Spach subsp. nordmanniana Saçinge hill, mixed forest, 1500 m, 13.07.1995, ÖE 1114, Mount Kurt, mixed forest, 1950 m, 10.06.1996, ÖE 1676, Eux. Picea orientalis (L.) Link Tafll›ca village, Gökçeyurt, mixed forest, 650 m, 16.05.1995, ÖE 1677, Tafll›ca village, Meydanlar, as pure stand or mixed forest with Pinus sylvestris L., Fagus orientalis Lipsky, Abies nordmanniana (Stev.) Spach, 1300 m. 13.07.1995, ÖE 1115, fiaflk›rat plateau, mixed forest, 2250 m, 10.06.1996, ÖE 1678, ATHYRIACEAE Eux. Athrium filix-foemina (L.) Roth Pinus sylvestris L. Tafll›ca village, Bakacak, along the water channel side, mixed forest, moist soil, 1150 m, 17.05.1995, ÖE 1110. Cosm. ASPIDIACEAE Polystichum lonchitis (L.) Roth Tafll›ca village, Demirci, roadside, rocky and stony slope, 1700 m, 16.08.1995, ÖE 1112. Dryopteris abbreviata (DC.) Newman Tafll›ca village, Ar›p›nar, rocky slope, 1800 m, 10.07.1997, ÖE 1840. D. dilatata (Hoffm.) Gray Saçinge hill, rocky slope, 1750 m, 08.07.1997, ÖE 1841. D. liliana Golitsin Tafll›ca village, Direkli, open area, wet soil, 1600 m, 16.08.1995, ÖE 1111. VU. Tafll›ca village, Meydanlar, as pure stand or mixed forest with Picea orientalis (L.) Link, Fagus orientalis Lipsky, Abies nordmanniana (Stev.) Spach, 850 m, 13.07.1995, ÖE 1116, Mount Kurt, mixed forest, 2200 m, 10.06.1996, ÖE 1679, Euro-Sib. P. pinea L. F›st›kl› village, together with shrub species as follow: Cistus creticus L., C. salviifolius L., Cotinus coggygria Scop., Carpinus orientalis Miller, Buxus sempervirens L., 550 m, 13.07.1995, ÖE 1117, Medit. TAXACEAE Taxus baccata L. Saçinge hill, solitary in mixed forest, 1000 m, 13.07.1995, ÖE 1118. Ö. EM‹NA⁄AO⁄LU, R. ANfi‹N, Juniperus communis hemisphaerica (Presl) Nyman Ranunculus cappadocicus Willd A. nasutum Fisch. ex Rchb. CUPRESSACEAE L. subsp. Tafll›ca village, Alacada¤, stony slope, 1800 m, 13.07.1995, ÖE 1119. J. communis L. subsp. alpina (Sm.) Celak. Tafll›ca village, Alacada¤, subalpine slope, 2300 m, 16.06.1996, ÖE 1522. J. oxycedrus L. subsp. oxycedrus F›st›kl› village, rocky and stony slope, 1450 m, macchie, 13.07.1995, ÖE 1120. J. foetidissima Willd. Tafll›ca village, Gökçeyurt, rocky and stony slope, together with Quercus L. and Carpinus L., 1200 m, 13.07.1995, ÖE 1121. Saçinge hill, streamside, 1800 m, 10.06.1997, ÖE 1846. Eux. Eux. Delphinium flexuosum M.Bieb. R. buhsei Boiss. Tafll›ca village, Ar›p›nar, slope, 1750 m, 13.07.1995, ÖE 1131. Hyr-Eux (mt). Consolida hohenackeri (Boiss.) Grossh. R. oreophilus M.Bieb. F›st›kl› village, dry slope, 400 m, 16.07.1997, ÖE 1847 Ir-Tur. Perikaya hill, rocky slope, 1900 m, 18.08.1997, ÖE 1848. Ephedra major Host C. regalis Gray subsp. regalis narcissiflora Euro-Sib. RANUNCULACEAE A. blanda Schott & Kotschy Helleborus orientalis Lam. F›st›kl› village, open forest, 470 m, 04.04.1995, ÖE 1132, Eux. Tafll›ca village, Meydanlar, meadow, 1250 m, 12.07.1997, ÖE 1843. Ir-Tur. 670 m, Tafll›ca village, Ar›p›nar, streamside, 1850 m, 17.06.1996, ÖE 1844. Tafll›ca village, Konac›k, moist rocky and stony slope, 1900 m, 12.04.1995, ÖE 1125. A. caucasica Willd. ex Rupr. VU, Eux. Ir-Tur. Saçinge hill, Picea Dietr. forest, 1400 m, 13.07.1995, ÖE 1124. 2800 m, Aconitum orientale Mill. Tafll›ca village, Ar›p›nar, meadow, 1700 m, 21.06.1997, ÖE 1845. Tafll›ca village, Ar›p›nar, water meadow, 1750 m, 25.05.2997, ÖE 1856. Tafll›ca village, Meydanlar, meadow, 1150 m, 21.05.1995, ÖE 1135. R. muricatus L. F›st›kl› village, muddy roadside, 700 m, 21.05.1997, ÖE 1857. R. arvensis L. R. pinardii (Stev.) Boiss. Tafll›ca village, Meydanlar, Picea Dietr. forest, 1200 m, 21.05.1995, ÖE 1592. Tafll›ca village, fiaflk›rat plateau, streamside, open forest, 2100 m, 03.06.1995, ÖE 1127. Thalictrum minus L. var. minus 400 m, Adonis aestivalis L. subsp. aestivalis Caltha polypetala Hochst. ex Lorent Tafll›ca village, Meydanlar, moist meadows, 1450 m, 03.06.1995, ÖE 1590. Aquilegia olympica Boiss. Tafll›ca village, Meydanlar, roadside, as climbing form on Carpinus L., 1350 m, 13.07.1995, ÖE 1129. F›st›kl› village, roadside, 13.07.1995, ÖE 1130. m, Tafll›ca village, Konac›k, waste place, 1650 m, 13.07.1995, ÖE 1591. A. albana Stev. C. orientalis L. Actaea spicata L. 2300 R. constantinopolitanus (DC.) d' Urv. Clematis vitalba L. Trollius ranunculinus (Smith) Stearn Alacada¤, rocky slope, 10.07.1997, ÖE 1855. R. grandiflorus L. Mount Otluca, alpine slope, 2650 m, 10.06.1997, ÖE 1850. N. segetalis M.Bieb. Tuzlu hill, streamside, 07.04.1996, ÖE 1128. subsp. Mount Keçi, roadside, moist soil, 1980 m, 12.04.1996, ÖE 1126. Nigella latisecta P.H.Davis F›st›kl› village, roadside, 13.06.1995, ÖE 1133. L. Mount Kurt, subalpine slope, 2400 m, 17.07.1997, ÖE 1849. DICOTYLEDONAE R. dissectus M.Bieb. subsp. napellifolius (DC.) P.H.Davis R. repens L. Çoruh: above Artvin, 800 m, D. 29695. Anemone narcissiflora Mount Otluca, stony slope, 2400 m, 06.08.1997, ÖE 1854. Euro-Sib. C. orientalis (Gay) Schröd. Ir-Tur. ANGIOSPERMAE Mount Keçi, rocky slope, 1900 m, 10.07.1997, ÖE 1853. Eux. EPHEDRACEAE Çoruh riverside, rocky slope, open area in the mixed forest, together with Acer L., Rhus L., Ficus L., Paliurus Mill., 400 m, 13.07.1995, ÖE 1123. Tafll›ca village, Meydanlar, Picea Dietr. forest, 1200 m, 21.05.1995, ÖE 1134. Tafll›ca village, Direkli, stony slope, 1450 m, 10.07.1997, ÖE 1851. A. flammea Jacq. Tafll›ca village, Ar›p›nar, stony place, 1700 m, 16.06.1997, ÖE 1852. Tafll›ca village, Ar›p›nar, damp meadow, 1850 m, 10.07.1997, ÖE 1858. BERBERIDACEAE Berberis vulgaris L. Hatipo¤lu, open forest, 12.10.1995, ÖE 1136. 1970 m, Eux. Eux. 7 The Flora of Hatila Valley National Park and its Close Environs (Artvin) Rapistrum rugosum (L.) All. PAPAVERACEAE Chelidonium majus L. Hatipo¤lu, roadside, 27.05.1995, ÖE 1137. 435 m, Conringia perfoliata (C.A.Mey.) Busch Euro-Sib. F›st›kl› village, roadside, 12.05.1997, ÖE 1864. Glaucium leiocarpum Boiss. Tafll›ca village, Bakacak, open forest, 950 m, 13.07.1995, ÖE 1139. Papaver orientale L. var. parviflora Busch Mount Otluca, stony slope, 2100 m, 15.08.1997, ÖE 1859. Ir-Tur. P. fugax Poiret var. fugax Perikaya hill, dry slope, 2000 m, 10.06.1997, ÖE 1860. P. dubium L. F›st›kl› village, roadside, 12.04.1996, ÖE 1140. 670 m, F›st›kl› village, stony slope, 550 m, 10.06.1997, ÖE 1861. Corydalis conorhiza Ledeb. Tuzlu hill, alpine vegetation, 2700 m, 10.07.1997, ÖE 2008. m, Lepidium campestre (L.) R.Br. Tafll›ca village, Meydanlar, waste place, 1250 m, 10.06.1997, ÖE 1865. L. perfoliatum L. Tafll›ca village, Gökçeyurt, stony slope, 1150 m, 05.06.1997, ÖE 1866. Cardaria draba (L.) Desv. subsp. chalepensis (L.) O.E.Schulz Tafll›ca village, Demirci, roadside, 1750 m, 14.04.1996, ÖE 1219. Thlaspi arvense L. Cosm. C. caucasica DC. Tafll›ca village, Direkli, moist soil, 1500 m, 28.02.1996, ÖE 1138. Tafll›ca village, Ar›p›nar, waste place, 1700 m, 10.06.1997, ÖE 1867. macrophyllum Tafll›ca village, K›l›çl›, roadside, 1850 m, 12.04.1996, ÖE 1228 Eux. Eux. Capsella bursa-pastoris (L.) Medik. Fumaria officinalis L. F›st›kl› village, open area, cultivated ground, 350 m, 1142, 13.04.1996, ÖE 1673. F. microcarpa Boiss. ex Hausskn. 1750 Tafll›ca village, Davütet, roadside, open forest, 1450 m, 13.07.1995, ÖE 1213. Cosm. Neslia paniculata (L.) Desv. m, F. asepala Boiss. Tafll›ca village, Gökçeyurt, roadside, 1050 m, 10.07.1997, ÖE 2009. CRUCIFERAE (BRASSICACEAE) Brassica elongata Ehrh. Tafll›ca village, Meydanlar, cultivated field, 1350 m, 10.06.1997, ÖE 1862 Ir-Tur. Hirschfeldia incana (L.) Lag.-Foss. Çoruh: beside the river Çoruh, Koch. Raphanus raphanistrum L. F›st›kl› village, open forest, 350 m, 15.04.1995, ÖE 1229. desertorum Stapf F›st›kl› village, fieldside, 18.06.1995, ÖE 1211. var. 360 m, A. xanthocarpum Boiss. Cerat hill, rocky slope, 1700 m, 10.06.1997, ÖE 1869. A. pseudo-mouradicum Hausskn. & Bornm. ex Baumg. Tafll›ca village, K›l›çl›, open area, 2100 m, 18.06.1995, ÖE 1210. Endemic, LR(lc). A. peltaroides Boiss. subsp. peltaroides Tafll›ca village, Ar›p›nar, roadside, 2200 m, 27.06.1995, ÖE 1209. Endemic, LR(lc), Ir-Tur. A. paphlagonicum (Hausskn.) Dudley Tafll›ca village, Konac›k, open forest, 1450 m, 27.06.1995, ÖE 1208. Endemic, LR(lc), Ir-Tur. A. murale Waldst. & Kit. var. murale T. huetii Boiss. Pachyphragma (Hoffm.) Busch Eux. 8 650 Tafll›ca village, Bakacak, slope, moist soil, 1000 m, 09.05.1995, ÖE 1232. P. arenarium M.Bieb. Mount Keçi, roadside, 12.07.1995, ÖE 1141. F›st›kl› village, cultivated place, 450 m, 10.06.1997, ÖE 1863. Alyssum Desertorum Tafll›ca village, Ar›p›nar, rocky slope, 1800 m, 21.07.1997, ÖE 1868. Alyssoides utriculata (L.) Munoz Med. Tafll›ca village, Gökçeyurt, roadside, 1250 m, 21.04.1995, ÖE 1207. Fibigia clypeata (L.) Medik. F›st›kl› village, rocky slope, 600 m, 21.04.1995, ÖE 1225. F. eriocarpa (DC.) Boiss. F›st›kl› village, roadside, rocky area, 470 m, 12.04.1996, ÖE 1223. F. suffruticosa (Vent.) Sweet Saçinge hill, roadside, stony slope, 1850 m, 21.04.1995, ÖE 1224. Tuzlu hill, subalpine slope, rocky place, 2200 m, 10.06.1997, ÖE 1870. Draba bruniifolia armeniaca Coode & Cullen Stev. Perikaya hill, slope, 16.06.1995, ÖE 1220. subsp. 1800 m, Endemic, LR(lc). D. hispida Willd. Mount Keçi, alpine slope, 2000 m, 13.04.1996, ÖE 1221. Eux (mt). D. siliquosa M.Bieb. fiaflk›rat plateau, alpine slope, 2200 m, 26.07.1997, ÖE 1871. Hyr-Eux (mt). D. muralis L. Tafll›ca village, K›l›çl›, fieldside, 1550 m, 09.04.1996, ÖE 1222. D. nemorosa L. Tuzlu hill, alpine slope, 2400 m, 10.06.1997, ÖE 1872. Erophila verna (L.) Chevall. subsp. verna Tafll›ca village, Ar›p›nar, slope, 1800 m, 20.06.1997, ÖE 1873. Arabis brachycarpa Rupr. Tuzlu hill, alpine pasture, 2800 m, 10.06.1997, ÖE 1874. Eux (mt). Ö. EM‹NA⁄AO⁄LU, R. ANfi‹N, A. caucasica Willd. subsp. caucasica F›st›kl› village, roadside, stony slope, 540 m, 05.04.1996, ÖE 1212. A. nova Vill. Perikaya hill, stony slope, 2100 m, 20.06.1997, ÖE 1875. Turritis glabra L. Mount Otluca, slope, 12.07.1997, ÖE 1876. 2200 m, Nasturtium officinale R.Br. Tafll›ca village, Bakacak, open forest, streamside, wet soil, 950 m, 16.06.1995, ÖE 1127. Rorippa austriaca (Crantz) Besser Tafll›ca village, Davütet, wet place, 1350 m, 10.06.1997, ÖE 1877. R. sylvestre (L.) Bess. Tafll›ca village, Meydanlar, streamside, 1250 m, 12.09.1995, ÖE 1230. Barbarea plantaginea DC. Tafll›ca village, Ar›p›nar, streamside, 1750 m, 10.07.1997, ÖE 1878. Cardamine bulbifera (L.) Crantz Tafll›ca village, Direkli, Picea Dietr. forest, moist soil, 1600 m, 12.04.1996, ÖE 1214. H. matronalis L. subsp. adzharica (Tzvelev) Cullen Tafll›ca village, Meydanlar, meadow, 1300 m, 23.06.1995, ÖE 1226. H. transcaucasica Tzvelev Çoruh: Artvin, 2100 m, Stn. & Hend. 5931. Erysimum cuspidatum (M.Bieb) DC. Tafll›ca village, Direkli, stony slope, 1500 m, 10.06.1997, ÖE 1880. Alliaria petiolata (M.Bieb.) Cavara & Grande Tafll›ca village, Meydanlar, open forest, 1150 m, 04.04.1995, ÖE 1206. Sisymbrium officinale (L.) Scop. Hatipo¤lu, roadside, wet soil, 650 m, 11.03.1996, ÖE 1231. S. loeselii L. Tafll›ca village, Meydanlar, roadside, 1250 m, 10.06.1997, ÖE 1881. CAPPARACEAE Capparis ovata Desf. var. herbacea (Willd.) Zohary Tafll›ca village, Çoruh riverside, roadside, 250 m, 16.05.1996, ÖE 1205. Euro-Sib. CISTACEAE C. quinquefolia (M.Bieb.) Schmalh. Cistus creticus L. Tafll›ca village, K›l›çl›, roadside, open forest, 1680 m, 12.04.1996, ÖE 1216. Euro-Sib. C. raphanifolia Pourr. subsp. acris (Gris) O.E.Schulz Cerat hill, open area, streamside, 1850 m, 20.06.1995, ÖE 1217. Euro-Sib. C. tenera Gmel. Tafll›ca village, Direkli, streamside, 1540 m, 16.06.1995, ÖE 1218. Hyr-Eux. C. impatiens L.var. pectinata (Palla) Trautv. Tafll›ca village, Demirci, mixed forest, wet soil, shady area, 1550 m, 12.04.1996, ÖE 1215. Euro-Sib. Hesperis bicuspidata (Willd.) Poiret F›st›kl› village, 350 m, 15.04.1996, ÖE 1195. Omni-Medit. C. salviifolius L. F›st›kl› village, Pinus pinea L. forest, 370 m, 15.04.1996, ÖE 1196. Medit. Helianthemum nummullarium (L.) Mill. subsp. tomentosum (Scop.) Schinz & Thell. F›st›kl› village, stony and rocky slope, 700 m, 03.05.1995, ÖE 1197. H. nummullarium (L.) Mill. subsp. ovatum Viv. Tafll›ca village, Meydanlar, stony and rocky slope, 1300 m, 21.08.1995, ÖE 1198. Fumana arabica (L.) Spach var. arabica Tafll›ca village, Direkli, mixed forest, 1450 m, 20.07.1997, ÖE 1882. VIOLACEAE Viola odorata L. Tafll›ca village, Meydanlar, moist soil, shady area, 1250 m, 15.04.1995, ÖE 1191. Cosm. V. suavis M.Bieb. F›st›kl› village, open forest, 740 m, 15.04.1996, ÖE 1194. V. alba Besser Tafll›ca village, Demirci, open forest, 1540 m, 13.04.1996, ÖE 1187. V. sieheana Becker Tafll›ca village, Demirci, stony slope, 1350 m, 12.05.1995, ÖE 1193. V. canina L. F›st›kl› village, roadside, 15.04.1996, ÖE 1192. 470 m, V. altaica Ker Gawl. subsp. oreades (M.Bieb.) Becker Tuzlu hill, alpine slope, 2900 m, 14.05.1996, ÖE 1189. V. kitaibeliana Roem. & Schult. F›st›kl› village, fieldside, 14.05.1996, ÖE 1190. 550 m, V. arvensis Murray Tafll›ca village, Demirci, meadow, 1400 m, 06.06.1997, ÖE 1883. POLYGALACEAE Polygala supina Schreb. Saçinge hill, open area, stony slope, 1850 m, 17.05.1995, ÖE 1396. P. pruinosa Boiss. subsp. pruinosa F›st›kl› village, open area, slope, 570 m, 07.05.1995, ÖE 1395. P. major Jacq. Tafll›ca village, Meydanlar, stony slope, 900-1350 m, 07.05.1995, ÖE 1394. Euro-Sib. P. anatolica Boiss. & Heldr. F›st›kl› village, slope, 17.05.1995, ÖE 1393. 450 m, P. vulgaris L. Tafll›ca village, Meydanlar, open forest, 1300 m, 07.05.1995, ÖE 1397. Euro-Sib. Tafll›ca village, Ar›p›nar, rocky slope, 1750 m, 20.07.1997, ÖE 1879. 9 The Flora of Hatila Valley National Park and its Close Environs (Artvin) CARYOPHYLLACEAE Arenaria rotundifolia M.Bieb. subsp. rotundifolia Tafll›ca village, Bakacak, meadow, 1350 m, 12.08.1995, ÖE 1593. A. pseudacantholimon Bornm. Tuzlu hill, alpine slope, 2600 m, 06.06.1997, ÖE 1884. Endemic, LR (lc), Ir-Tur. A. armeniaca Boiss. Tafll›ca village, Ar›p›nar, dry rocky slope, 1700 m, 16.06.1997, ÖE 1885. Endemic, LR (lc), Ir-Tur. Minuartia imbricata (M.Bieb) Woronow Tuzlu hill, rocky-stony slope, 2750 m, 10.06.1997, ÖE 1886. Eux (mt). M. aizoides (Boiss.) Bornm. Tafll›ca village, Gökçeyurt, roadside, damp places, 1200 m, 10.06.1994, ÖE 1674. M. circassica (Albow) Woron. Perikaya hill, alpine slope, 2050 m, 06.06.1997, ÖE 1887. Eux (mt). Stellaria media (L.) Vill. subsp. media F›st›kl› village, Hatila streamside, 470 m, 14.04.1996, ÖE 1175. S. holostea L. Tafll›ca village, Ar›p›nar, slope, 1850 m, 12.07.1997, ÖE 1888. Euro-Sib. Myosoton aquaticum (L.) Moench F›st›kl› village, streamside, moist soil, 400 m, 15.05.1996, ÖE 1167. Euro-Sib. Cerastium cerastioides (L.) Britt. Tafll›ca village, Bakacak, meadow, 1250 m, 12.08.1995, ÖE 1595. C. armeniacum Gren. Saçinge hill, slope, 1750 m, 10.06.1997, ÖE 1889. Endemic, LR (lc). C. chlorifolium Fisch. & Mey. Tafll›ca village, Ar›p›nar, meadow, 1550 m, 10.07.1997, ÖE 1890. 10 C. gnaphalodes Fenzl S. compacta Fisch. Tuzlu hill, slope, 2700 m, 24.08.1997, ÖE 1891. Endemic, LR (lc). S. alba (Mill.) Krause subsp. divaricata (Rchb.) Walters C. glomeratum Thuill. F›st›kl› village, slope, 28.04.1997, ÖE 1892. Tafll›ca village, Ar›p›nar, open forest, roadside, 1450 m, 17.07.1995, ÖE 1173. 600 m, Cosm. Sagina procumbens L. Mount Otluca, waste place, 2200 m, 20.07.1997, ÖE 1893. Dianthus floribundus Boiss. Tafll›ca village, Direkli, roadside, 1600 m, 16.08.1995, ÖE 1166. Cerat hill, roadside, 1400 m, 13.07.1995, ÖE 1170, Ar›p›nar, roadside 2200 m, 21.07.1995, ÖE 1171. S. alba (Mill.) Krause subsp. eriocalycina (Boiss.) Walters Tafll›ca village, Gökçeyurt, roadside, 950 m, 21.07.1995, ÖE 1172. ILLECEBRACEAE Scleranthus uncinatus Schur Ir-Tur. Çoruh: mountain above Artvin, D 29805. D. crinitus Sm. var. crinitus POLYGONACEAE Tafll›ca village, Bakacak, rocky and dry soil, 800 m, 13.07.1995, ÖE 1165. D. artwinensis Schischk. Tafll›ca village, Meydanlar, roadside, 950 m, 03.08.1995, ÖE 1163. Endemic, LR(lc). D. calocephalus Boiss. Tafll›ca village, K›l›çl›, roadside, rocky and stony slope, open forest, 1450 m, 21.07.1995, ÖE 1164. Polygonum alpinum All. Tuzlu hill, alpine slope, 2850 m, 12.08.1997, ÖE 1895. Euro-Sib. P. bistorta L. subsp. carneum (Koch) Coode & Cullen Saçinge hill, moist slope, 1850 m, 13.07.1995, ÖE 1176. Eux (mt). Rumex acetocella L. Petrorhagia saxifraga (L.) Link Tafll›ca village, Meydanlar, dry soil, 1100 m, 03.08.1995, ÖE 1168. Tafll›ca village, Ar›p›nar, streamside, open forest, 1500 m, 15.07.1995, ÖE 1177. Euro-Sib. Cosm. Saponaria orientalis L. R. scutatus L. Tafll›ca village, Bakacak, rocky and dry soil, 800 m, 13.07.1995, ÖE 1596. Gypsophila tenuifolia M.Bieb. Tafll›ca village, Meydanlar, meadow, 1300 m, 06.08.1995, ÖE 1597. Eux (mt). R. tuberosus L. subsp. horizontalis (C.Koch) Rech. Tafll›ca village, fiaflk›rat plateau, subalpine slope, 2100 m, 11.06.1995, ÖE 1178. R. caucasicus Rech Silene italica (L.) Pers. Hatipo¤lu, open forest, 13.07.1995, ÖE 1169. Tafll›ca village, Ar›p›nar, meadow, 1700 m, 10.06.1997, ÖE 1896. 745 m, Medit. S. spergulifolia (Desf.) M.Bieb. Tafll›ca village, Ar›p›nar, stony slope, 1700 m, 10.06.1997, ÖE 1894. Ir-Tur. S. vulgaris (Moench) Garcke var. vulgaris Saçinge hill, roadside, open forest, 1800 m, 21.07.1995, ÖE 1174. Mount Keçi, slope, 12.08.1997, ÖE 1897. 1900 m, R. crispus L. Tafll›ca village, Köklü, meadow, 1450 m, 10.06.1997, ÖE 1898. CHENOPODIACEAE Chenopodium foliosum (Moench) Asch. F›st›kl› village, roadside, 10.06.1997, ÖE 1899. 450 m, Ö. EM‹NA⁄AO⁄LU, R. ANfi‹N, Phytolacca americana L. F›st›kl› village, roadside, 13.07.1995, ÖE 1162. G. pyrenaicum Burm. A. calvertii (Boiss.) Boiss. PHYTOLACCACEAE 530 m, Cosm. TAMARICACEAE Tamarix tetrandra Pall. ex M.Bieb. Çoruh riverside, 350 m, 07.10.1995, ÖE 1199. T. smyrnensis Bunge Çoruh riverside, 170 m, 10.06.1997, ÖE 1900. GUTTIFERAE Hypericum androsaemum L. Tafll›ca village, Meydanlar, roadside, streamside, 1300 m, 13.07.1995, ÖE 1179, F›st›kl› village, streamside, 370 m, 15.06.1996, ÖE 1180. Euro-Sib. H. bupleuroides Gris. Tafll›ca village, Davütet, open area in mixed forest, moist soil, 1300 m, 11.08.1995, ÖE 1182. Eux. H. montanum L. Tuzlu hill, stony slope, 2250 m, 20.07.1997, ÖE 1901. Tafll›ca village, Ar›p›nar, rocky slope, 1600 m, 20.07.1997, ÖE 1902. Endemic, LR (lc), Ir-Tur. Tafll›ca village, Bakacak, roadside, 1450 m, 17.06.1995, ÖE 1599. Althaea cannabina L. TILIACEAE Saçinge hill, roadside, mixed forest, 1320 m, 12.08.1995, 10549, ÖE 1185. Eux. Eux. H. orientale L. Tafll›ca village, fiaflk›rat plateau, subalpine slope, 2200 m, 17.06.1995, ÖE 1183, Ar›p›nar, roadside, 1650 m, 11.05.1996, ÖE 1675. H. perforatum L. Saçinge hill, open forest, wet soil, 1900 m, 03.08.1995, ÖE 1184. Tafll›ca village, Meydanlar, stony slope, 1300 m, 07.05.1995, ÖE 1390. MALVACEAE Malva sylvestris L. Tafll›ca village, Bakacak, roadside, 1400 m, 17.06.1995, ÖE 1598. Alcea hohenackeri (Boiss. & Huet) Boiss. Tafll›ca village, Direkli, roadside, open forest, 1450 m, 21.07.1995, ÖE 1186. 550 m, Medit. Endemic, LR(lc). Oxalis acetosella L. Tafll›ca village, Meydanlar, Picea Dietr. forest, moist soil, 1070 m, 15.07.1995, ÖE 1378. Medit. L. catharticum L. Tafll›ca village, Ar›p›nar, stony slope, 1750 m, 16.07.1995, ÖE 1392. Euro-Sib. O. corniculata L. Tafll›ca village, Bakacak, mixed forest, 1500 m, 10.07.1996, ÖE 1602. Cosm. BALSAMINACEAE Hatipo¤lu, Picea Dietr. forest side, 450 m, 14.03.1996, ÖE 1381. G. purpureum Vill. Tafll›ca village, Gölgeli, rocky and stony slope, 1000 m, 18.07.1995, ÖE 1386. G. robertianum L. Impatiens noli-tangere L. Tafll›ca village, Demirci, streamside, 1150 m, 19.07.1995, ÖE 1389. Euro-Sib. ACERACEAE Acer trautvetteri Medw. Tafll›ca village, Bakacak, open forest, 1025 m, 21.06.1995, ÖE 1387. G. molle L. subsp. molle Tafll›ca village, fiaflk›rat plateau, mixed forest, 2550 m, 14.05.1996, ÖE 1375. Eux. 500 m, G. psilostemon Ledeb. Tafll›ca village, Konac›k, Picea Dietr. forest, 1950 m, 21.06.1995, ÖE 1385. Eux. G. sylvaticum L. Tafll›ca village, Ar›p›nar, mixed forest, 1950 m, 18.07.1995, ÖE 1388. Euro-Sib. F›st›kl› village, open forest, 300 m, 14.03.1996, ÖE 1380. OXALIDACEAE L. bienne Mill. F›st›kl› village, fieldside, 13.03.1995, ÖE 1382. E. malacoides (L.) L' Hér. Tafll›ca village, Direkli, rocky area, 1450 m, 11.04.1996, ÖE 1379. Linum aroanium Boiss. & Orph. Geranium lucidum L. Tafll›ca village, Gölgeli, rocky slope, 850 m, 16.07.1996, ÖE 1601. E. amanum Boiss. & Kotschy LINACEAE H. bithynicum Boiss. Hyr-Eux (mt). Ir-Tur. Tilia rubra DC. subsp. caucasica (Rupr.) V.Engl. GERANIACEAE Mount Keçi, mixed forest, 2000 m, 18.07.1995, ÖE 1383. Erodium oxyrrhynchum M.Bieb. Tafll›ca village, Meydanlar, meadow, 1300 m, 16.07.1996, ÖE 1600. Euro-Sib. Tafll›ca village, Gölgeli, moist soil, 750 m, 21.08.1995, ÖE 1181. G. platypetalum Fisch. & Mey. A. excubita Iljin F›st›kl› village, fieldside, 07.05.1995, ÖE 1391. Tafll›ca village, Meydanlar, open forest, 970 m, 18.07.1995, ÖE 1384. A. cappadocicum cappadocicum Gled. var. Tafll›ca village, Direkli, mixed forest, 8001850 m, 17.08.1995, ÖE 1370. Hyr-Eux. A. cappadocicum stenocarpum Yalt. Gled. var. Tafll›ca village, Konac›k, mixed forest, 1370 m, 17.08.1995, ÖE 1371. Endemic, VU, Eux. 11 The Flora of Hatila Valley National Park and its Close Environs (Artvin) A. platanoides L. F›st›kl› village, roadside, 1300 m, 13.06.1995, ÖE 1374. Euro-Sib. A. divergens Pax var. divergens Çoruh riverside, 350 m, 17.07.1995, ÖE 1372. Endemic, VU. A. campestre L. subsp. campestre Tafll›ca village, Arganet, mixed forest, 1300 m, 24.08.1995, ÖE 1368. A. campestre L. subsp. leiocarpum (Opiz) Pax Hatipo¤lu, rocky and stony slope, 850 m, 24.08.1995, ÖE 1369. Euro-Sib. A. hyrcanum Fisch. & Mey. subsp. hyrcanum Tafll›ca village, Meydanlar, roadside, stony slope, 1400 m, 13.06.1995, ÖE 1373. Euro-Sib. RHAMNACEAE Paliurus spina - christi Mill. F›st›kl› village, roadside, stony slope, dry soil, 470 m, 12.09.1995, ÖE 1365. Frangula alnus Mill. subsp. alnus Tafll›ca village, Gölgeli, streamside, 1700 m, 12.09.1995, ÖE 1366. Euro-Sib. Rhamnus imeretinus Booth Tafll›ca village, Demirci, mixed forest, 1500 m, 12.09.1995, ÖE 1367. Eux. R. microcarpus Boiss. Tafll›ca village, Meydanlar, open forest, 1200 m, 16.06.1996, ÖE 1603. Eux (mt). AQUIFOLIACEAE Ilex colchica Pojark. Tafll›ca village, Davütet, mixed forest, moist soil, 970 m, 13.07.1995. ÖE 1359. Eux. ANACARDIACEAE Cotinus coggygria Scop. Çoruh riverside, roadside, 500 m, 21.06.1995, ÖE 1376. Medit. 12 Rhus coriaria L. Çoruh riverside, roadside, stony slope, 750 m, 18.08.1995, ÖE 1377. Medit. Pistacia terebinthus palaestina (Boiss.) Engler L. subsp. Euonymus latifolius (L.) Mill. subsp. latifolius Tafll›ca village, Köklü, mixed forest, 1300 m, 15.07.1995, ÖE 1358. Euro-Sib. Tafll›ca village, Direkli, mixed forest, moist soil, 1450 m, 15.07.1995, ÖE 1357. Endemic, LR(nt). LEGUMINOSAE (FABACEAE) A. aureus Willd. Tafll›ca village, Meydanlar, stony slope, 1350 m, 10.07.1997, ÖE 1904. Ir-Tur. Cerat hill, open forest, 1600 m, 12.06.1996, ÖE 1606. A. woronowii Bornm. Tafll›ca village, fiaflk›rat plateau, alpine vegetation, 2500 m, 12.06.1996, ÖE 1607. A. falcatus Lam. Chamaecytisus hirsutus (L.) Link 650 m, Mount Keçi, Pinus L. forest, 1700 m, 10.06.1997, ÖE 1905. Eux (mt). Genista tinctoria L. A. adzharicus Popov 460 m, Euro-Sib. Cerat hill, waste place, 1650 m, 16.07.1996, ÖE 1608. A. sanguinolentus M.Bieb. G. albida Willd. Tafll›ca village, Gölgeli, rocky and stony slope, 950 m, 24.04.1996, ÖE 1310. Argyrolobium biebersteinii Ball F›st›kl› village, rocky place, 540 m, 15.05.1995, ÖE 1298. Galega officinalis L. Tafll›ca village, Arganet, dry soil, 1350 m, 20.07.1997, ÖE 1906. A. frickii Bunge Perikaya hill, mixed forest, 2100 m, 17.05.1995, ÖE 1302. Eux (mt). A. taochius Woron. Tafll›ca village, Meydanlar, open forest, 1200 m, 13.07.1995, ÖE 1308. Colutae armena Boiss. & Huet 750 F›st›kl› village, roadside, 10.06.1997, ÖE 1907. 600 m, Psoralea acaulis Stev. m, Astragalus oreades Mey. Tafll›ca village, Meydanlar, waste place, 1200 m, 10.06.1996, ÖE 1604. A. flaccidus M.Bieb. Tafll›ca village, Bakacak, roadside, 1400 m, 22.08.1996, ÖE 1605. A. glycyphyllos L. subsp. glycyphyllos Tafll›ca village, Bakacak, forestside, 1300 m, 17.05.1995, ÖE 1299. Euro-Sib. Tafll›ca village, Ar›p›nar, mixed forest, 1850 m, 11.07.1995, ÖE 1301. A. sommieri Freyn E. latifolius (L.) Mill. subsp. cauconis Coode & Cullen F›st›kl› village, roadside, 18.08.1995, ÖE 1304. Euro-Sib. Ir-Tur. CELASTRACEAE F›st›kl› village, roadside, 19.04.1995, ÖE 1309. Tafll›ca village, Arganet, open forest, 1350 m, 11.07.1995, ÖE 1300. A. fraxinifolius DC. Tafll›ca village, Gölgeli, mixed forest, 1100 m, 20.07.1997, ÖE 1903. F›st›kl› village, roadside, 12.05.1996, ÖE 1303. A. glycyphyllos L. subsp. glycyphylloides (DC.) Matthews Tafll›ca village, Direkli, slope, 1560 m, 06.06.1997, ÖE 1908. Eux. Vicia crocea (Desf.) B.Fedtsch. Tafll›ca village, Demirci, Picea orientalis (L.) Link forest, 1320 m, 17.09.1995, ÖE 1342. Hyr-Eux. V. dadianorum Sommer & Lév. Tafll›ca village, Meydanlar, slope, 1300 m, 21.07.1997, ÖE 1908. Eux. Ö. EM‹NA⁄AO⁄LU, R. ANfi‹N, V. cassubica L. L. cicera L. Tafll›ca village, Meydanlar, roadside, 1200 m, 22.05.1995, ÖE 1340. Euro-Sib. Tafll›ca village, Gölgeli, mixed forest, 1100 m, 22.05.1995, ÖE 1341. Euro-Sib. V. villosa Roth subsp. villosa Tafll›ca village, Davütet, roadside, stony slope, 1350 m, 12.07.1995, ÖE 1346. V. sepium L. Tafll›ca village, K›l›çl›, mixed forest, wet soil, 1600 m, 25.06.1995, ÖE 1345. Pisum sativum L. subsp. elatius (M.Bieb.) Asch. & Graebn. var. elatius Saçinge hill, open slope, 1700 m, 16.04.1996, ÖE 1325. Ononis arvensis L. V. grandiflora Scop. var. grandiflora 600 m, Euro-Sib. V. sativa L. subsp. nigra (L.) Ehrh. var. nigra Tafll›ca village, Meydanlar, roadside, rocky slope, 1350 m, 28.06.1995, ÖE 1344. Lathyrus aureus (Stev.) Brândză F›st›kl› village, open forest, 450 m, 12.04.1995, ÖE 1311. Euro-Sib. L. vernus (L.) Bernh. 560 m, Euro-Sib. L. laxiflorus (Desf.) Kuntze subsp. laxiflorus Tafll›ca village, Demirci, Picea Dietr. forest, 1320 m, 13.07.1995, ÖE 1313. L. roseus Stev. Tafll›ca village, Kanac›k, Pinus L. forest, 1500 m, 10.06.1997, ÖE 1908. Hyr-Eux. L. tuberosus L. Saçinge hill, open forest, moist soil, 1500 m, 16.07.1995, ÖE 1315. Euro-Sib. L. rotundifolius Willd. subsp. miniatus (M.Bieb. ex Stev.) P.H. Davis Tafll›ca village, Meydanlar, meadow pasture, 1350 m, 16.07.1995, ÖE 1316. Hyr-Eux. T. trichocephalum M.Bieb. Tafll›ca village, Ar›p›nar, open forest, 1680 m, 12.07.1995, ÖE 1338. Tafll›ca village, Ar›p›nar, open forest, 1850 m, 12.07.1995, ÖE 1328. T. arvense L. var arvense 1900 m, Cosm. T. montanum L. subsp. humboldtianum (A.Br. & Asch.) Hossain Mount Keçi, open slope, 1850 m, 22.08.1995, ÖE 1334. T. ambiguum M.Bieb. Saçinge hill, open area, 1900 m, 12.07.1995, ÖE 1327. Euro-Sib. T. hybridum L. var. hybridum Eux. Tafll›ca village, Arganet, roadside, 1400 m, 12.07.1995, ÖE 1329. T. alpestre L. Tafll›ca village, Meydanlar, moist and dry slope, 1400 m, 21.07.1997, ÖE 1909. Mount Kurt, roadside, 23.05.1996, ÖE 1337. Tafll›ca village, Demirci, roadside, 1370 m, 12.07.1995, ÖE 1333. T. canescens Willd. F›st›kl› village, roadside, fieldside, 340 m, 25.04.1995, ÖE 1314. Trifolium repens L. var repens Euro-Sib. F›st›kl› village, roadside, 16.07.1995, ÖE 1317. Tafll›ca village, Meydanlar, Pinus L. forest, 1350 m, 17.03.1996, ÖE 1312. L. nissolia L. V. cracca L subsp. cracca F›st›kl› village, roadside, 13.05.1995, ÖE 1343. T. ochroleucum Huds. Tafll›ca village, Demirci, roadside, 1400 m, 21.08.1995, ÖE 1331. Tafll›ca village, Meydanlar, meadow, 1250 m, 16.06.1995, ÖE 1607. Melilotus officinalis (L.) Desr. Tafll›ca village, Gölgeli, open area, 750 m, 13.07.1995, ÖE 1323. Cosm. Medicago lupulina L. Tafll›ca village, Bakacak, open forest, wet soil, 1100 m, 07.03.1996, ÖE 1321. M. sativa L. subsp. sativa Mount Keçi, stony slope, 1850 m, 19.06.1995, ÖE 1322. M. arabica (L.) Huds. F›st›kl› village, open forest, 450 m, 07.03.1996, ÖE 1320. Cosm. Cosm. T. campestre Schreb. Dorycnium graecum (L.) Ser. F›st›kl› village, open forest, 400 m, 22.08.1995, ÖE 1330. T. tumens Stev. ex M.Bieb. Tafll›ca village, Arganet, roadside, 900 m, 23.05.1996, ÖE 1339. Hyr-Eux. T. pratense L. var. pratense Tafll›ca village, Bakacak, open forest, 1100 m, 22.08.1995, ÖE 1335. Cosm. T. pratense L. var. sativum Schreb. Tafll›ca village, Arganet, open area, 1300 m, 23.05.1996, ÖE 1336. T. medium L. var. medium Tafll›ca village, Meydanlar, open forest, 1050 m, 22.08.1995, ÖE 1332. Cosm. F›st›kl› village, open slopes, 370 m, 15.07.1995, ÖE 1307. Eux. D. pentaphyllum herbaceum (Vill.) Rouy Scop. subsp. Tafll›ca village, Meydanlar, roadside, 1400 m, 20.07.1997, ÖE 1909. Lotus corniculatus L. var. corniculatus Hatipo¤lu, roadside, 13.07.1995, ÖE 1318. 690 m, L. corniculatus L. var. tenuifolius L. Tafll›ca village, K›l›çl›, streamside, 1500 m, 13.07.1995, ÖE 1319. Anthllis vulneraria L. subsp. polyphylla (DC.) Nyman Tafll›ca village, Meydanlar, open forest, 1350 m, 15.05.1995, ÖE 1297. Euro-Sib. 13 The Flora of Hatila Valley National Park and its Close Environs (Artvin) Coronilla coronata L. R. caucasicus Focke Alacada¤, alpine vegetation, 2750 m, 16.06.1996, ÖE 1608. C. orientalis Mill. var. orientalis Mount Otluca, mixed forest, 1800 m, 10.06.1997, ÖE 1913. Eux. Tafll›ca village, Direkli, roadside, 1470 m, 13.06.1995, ÖE 1305. C. varia L. subsp. varia Tafll›ca village, Bakacak, open forest, 750 m, 13.06.1995, ÖE 1306. Onobryhis armena Boiss. & Huet Tafll›ca village, Ar›p›nar, meadow, 1650 m, 10.06.1997, ÖE 1910. O. altissima Grossh. Tafll›ca village, Meydanlar, open forest, 1230 m, 19.06.1995, ÖE 1324. O. huetiana Boiss. Tafll›ca village, Direkli, meadow, 1700 m, 21.07.1997, ÖE 1911. Cerat hill, roadside, 23.07.1995, ÖE 1287. 1540 m, Aruncus vulgaris Raf. Tafll›ca village, Ar›p›nar, mixed forest, wet soil, 1575 m, 13.07.1995, ÖE 1270. Tafll›ca village, Köklü, open forest, 1300 m, 23.07.1995, ÖE 1282. P. crantzii (Crantzow) Beck ex Fritsch var. crantzii Tafll›ca village, Ar›p›nar, open area, 1970 m, 23.07.1995, ÖE 1284. 480 m, Eux. Prunus spinosa L. subsp. dasyphylla (Schur) Domin Tafll›ca village, Meydanlar, open forest, wet soil, 1200 m, 12.05.1995, ÖE 1279. Tafll›ca village, Davütet, mixed forest, 1650 m, 12.06.1996, ÖE 1610. Rubus saxatilis L. Cerat hill, open 30.06.1997, ÖE 1912. place, 1750 m, R. idaeus L. Tafll›ca village, Köklü, roadside, stony slope, 1370 m, 21.08.1995, ÖE 1290. Cosm. R. caesius L. Tafll›ca village, Bakacak, open forest, 1570 m, 22.07.1996, ÖE 1611. R. platyphyllos C.Koch. Tafll›ca village, Meydanlar, slope, 1270 m, 21.08.1995, ÖE 1291. Eux. 14 Fragaria vesca L. parviflora Willd. var. Alacada¤, alpine vegetation, 2600 m, 10.06.1995, ÖE 1612. Geum coccineum Sm. Tuzlu hill, marshy slope, 2500 m, 06.06.1997, ÖE 1914. Endemic, LR(nt), Eux. A. retinervis Buser Tuzlu hill, alpine pasture, 3000 m, 10.06.1997, ÖE 1916. Tafll›ca village, Bakacak, open forest, 1550 m, 20.07.1996, ÖE 1616. R. iberica Stev. Tafll›ca village, Kanac›k, open area, 1450 m, 10.06.1997, ÖE 1918. R. montana Chaix subsp. woronowii (Lonacz.) O.Nilsson Mount Keçi, open forest, 1800 m, 28.06.1997, ÖE 1919. R. canina L. F›st›kl› village, open forest, 450 m, 08.06.1995, ÖE 1289 Cosm. Tafll›ca village, Demirci, open forest, 1550 m, 18.07.1995, ÖE 1281. G. urbanum L. Saçinge hill, in forest, 12.07.1996, ÖE 1613. Tafll›ca village, K›l›çl›, roadside, 1600 m, 28.06.1997, ÖE 1917. Mespilus germanica L. Euro-Sib. 1600 m, Agrimonia eupatoria L. Tafll›ca village, Direkli, open forest, wet soil, 1480 m, 12.09.1995, ÖE 1267. Aremonia agrimonoides (L.) DC. Tafll›ca village, Meydanlar, meadow, 1350 m, 06.06.1997, ÖE 1915. Euro-Sib. Tafll›ca village, fiaflk›rat plateau, alpine slope, 1800 m, 05.05.1995, ÖE 1268. R. elymaitica Boiss. & Hausskn. Tafll›ca village, Gökçeyurt, open forest, 970 m, 23.07.1995, ÖE 1285. Cerasus avium (L.) Moench Eux (mt). Rosa pimpinellifolia L. F›st›kl› village, fieldside, 08.08.1995, ÖE 1286. Sibbaldia parviflora Tafll›ca village, fiaflk›rat plateau, alpine vegetation, 2800 m, 20.07.1996, ÖE 1615. Eux (mt). Tafll›ca village, Davütet, Hatila streamside, mixed forest, 1350 m, 16.05.1996, ÖE 1280. Tafll›ca village, Meydanlar, mixed forest, 1300 m, 16.06.1996, ÖE 1609. Tafll›ca village, fiaflk›rat plateau, alpine vegetation, 2600 m, 20.07.1996, ÖE 1614. A. sintenisii Rothm. Tafll›ca village, Demirci, open area, 1350 m, 23.07.1995, ÖE 1283. P. micrantha Ramond ex DC. Laurocerasus officinalis Roem. Tafll›ca village, Bakacak, roadside, dry soil, 1550 m, 10.06.1996, ÖE 1613. A. caucasica Buser P. argentea L. P. reptans L. ROSACEAE subsp. Eux (mt). Euro-Sib. Euro-Sib. Endemic, LR (nt), Ir-Tur. Scop. Alchemilla sericea Willd. Potentilla rupestris L. P. recta L. Endemic, LR (lc). Sanguisorba minor muricata (Spach) Briq. Hyr-Eux. Cotoneaster nummularia Fisch. & Mey. Tafll›ca village, Meydanlar, open forest, rocky and stony slope, 1200 m, 18.08.1995, ÖE 1271. C. transcaucasicus Pojark. Tafll›ca village, Demirci, stony slope, 1500 m, 12.09.1995, ÖE 1272. VU. Ö. EM‹NA⁄AO⁄LU, R. ANfi‹N, C. morulus Pojark. F›st›kl› village, rocky and stony slope, 750 m, 12.09.1995, ÖE 1273. Pyracantha coccinea Roem. Çoruh riverside, 550 m, 15.05.1996, ÖE 1288. Crataegus pentagyna Waldst., Kit. ex Willd. Tafll›ca village, Meydanlar, stony slope, 1250 m, 20.07.1997, ÖE 1920. Euro-Sib. C. orientalis Pall. ex M.Bieb. var. orientalis Saçinge hill, open forest, 1870 m, 11.08.1995. C. curvicephala Lindm. Tafll›ca village, Ar›p›nar, open forest, 1870 m, 23.09.1995, ÖE 1274. C. monogyna Jacq. subsp. monogyna Tafll›ca village, Demirci, open forest, 1370 m, 11.08.1995, ÖE 1277. C. monogyna Jacq. subsp. azarella (Gris.) Franco Tafll›ca village, Gökçeyurt, open forest, 720 m, 27.08.1995, ÖE 1276. C. microphylla C.Koch Tafll›ca village, Demirci, open forest, 1325 m, 23.09.1995, ÖE 1275. Hyr-Eux. Sorbus aucuparia L. Tafll›ca village, fiaflk›rat plateau, subalpine slope, 2400 m, 11.08.1995, ÖE 1293. Euro-Sib. S. umbellata (Desf.) Fritsch var. cretica (Lindl.) Schneid. F›st›kl› village, Quercus scrub, rocky area, 2250 m, 17.08.1995, ÖE 1294. S. umbellata (Desf.) Fritsch var. orbiculata (Kárpáti) Gabr. Tafll›ca village, Meydanlar, mixed forest, 1450 m, 17.08.1995, ÖE 1295. Pyrus communis L. subsp. communis Tafll›ca village, Bakacak, open forest, 1400 m, 10.06.1996, ÖE 1618. P. elaeagnifolia elaeagnifolia Pall. subsp. (Lam.) F›st›kl› village, open forest, 700 m, 10.08.1995, ÖE 1269. PUNICACEAE Punica granatum L. Çoruh riverside, 260 m, 16.05.1996, ÖE 1349. LYTHRACEAE Lytrum salicaria L. F›st›kl› village, open area, streamside, 450 m, 08.08.1995, ÖE 1347. Euro-Sib. ONAGRACEAE Circeae lutetiana L. Tafll›ca village, Gölgeli, mixed forest, 750 m, 13.07.1995, ÖE 1352. C. alpina L. Tafll›ca village, K›l›çl›, mixed forest, 1400 m, 20.07.1997, ÖE 1921. Epilobium angustifolium L. Saçinge hill, roadside, open forest, 1600 m, 13.07.1995, ÖE 1350. E. montanum L. Mount Keçi, open forest, 1850 m, 13.07.1995, ÖE 1351. Euro-Sib. DATISCACEAE Datisca cannabina L. Tafll›ca village, Gölgeli, streamside, 1000 m, 10.06.1997, ÖE 1922. CRASSULACEAE S. torminalis (L.) Crantz var. torminalis Sedum telephium L. subsp. maximum (L.) Krock. Tafll›ca village, Meydanlar, mixed forest, 1250 m, 11.07.1995, ÖE 1296. Tafll›ca village, Bakacak, roadside, rocky slope, 1070 m, 16.08.1995, ÖE 1263. Euro-Sib. Malus sylvestris Mill. subsp. orientalis (Uglitzk.) Browicz var. orientalis Tafll›ca village, Meydanlar, mixed forest, 1300 m, 10.06.1996, ÖE 1617. Saçinge hill, Picea Dietr. forest, 8501850 m, 21.08.1995, ÖE 1262. Hyr-Eux (mt). S. alpestre Vill. Tafll›ca village, Demirci, open forest, 1570 m, 11.09.1995, ÖE 1292. Amelanchier rotundifolia Dum.Cours. subsp. rotundifolia S. stoloniferum Gmel. Tafll›ca village, Davütet, roadside, rocky place, 1400 m, 06.06.1997, ÖE 1923. S. album L. Cerat hill, roadside, stony slope, 1500 m, 13.05.1996, ÖE 1257. S. gracile C.A.Mey. Tafll›ca village, Ar›p›nar, rocky area, 2400 m, 13.05.1996, ÖE 1258. Hyr.-Eux. S. pallidum M.Bieb. var. pallidum Tafll›ca village, K›l›çl›, roadside, rocky slope, 1350 m, 21.08.1995, ÖE 1260. S. pallidum M.Bieb. var. bithynicum (Boiss.) Chamb. Tafll›ca village, Demirci, roadside, Picea Dietr. forest, 1570 m, 18.07.1995, ÖE 1259. Eux. Sempervivum glabrifolium Boriss. F›st›kl› village, rocky and stony slope, 750 m, 16.08.1995, ÖE 1264. Endemic, LR (cd). S. davisii Muirhead Tafll›ca village, Gökçeyurt, stony slope, 1050 m, 06.06.1997, ÖE 1926. SAXIFRAGACEAE Saxifraga paniculata Mill. cartilaginea (Willd.) D.A.Webb Tafll›ca village, Demirci, open forest, rocky area, 1470 m, 18.05.1995, ÖE 1265. Hyr-Eux. S. artvinensis Matthews Perikaya hill, stony slope, 1800 m, 20.07.1997, ÖE 1927. Endemic, VU, Eux (mt). S. rotundifolia L. Tafll›ca village, Meydanlar, stony slope, 1170 m, 12.06.1995, ÖE 1266. Euro-Sib. Euro-Sib. S. spurium M.Bieb. S. cymbalaria L. var. cymbalaria Tafll›ca village, K›l›çl›, rocky area, 1800 m, 25.07.1995, ÖE 1261. subsp. Tafll›ca village, Köklü, streamside, 1500 m, 06.06.1997, ÖE 1928. Hyr.-Eux. 15 The Flora of Hatila Valley National Park and its Close Environs (Artvin) Daucus carota L. GROSSULARIACEAE Ribes biebersteinii Berl. ex DC. Tafll›ca village, fiaflk›rat plateau, subalpine slope, 2250 m, 18.07.1995, ÖE 1256. R. alpinum L. Tuzlu hill, subalpine slope, 2300 m, 20.07.1997, ÖE 1929. UMBELLIFERAE (APIACEAE) Sanicula europaea L. Tafll›ca village, Ar›p›nar, Picea Dietr. forest, 1750 m, 29.06.1995, ÖE 1402. Euro-Sib. Tafll›ca village, Arganet, on the ground mixed forest, 1370 m, 13.07.1995, ÖE 1398. Tafll›ca village, Meydanlar, meadow, 1130 m, 29.06.1995, ÖE 1400 Eux. Saçinge hill, open forest, 1500 m, 26.08.1996, ÖE 1619. Chaerophyllum bulbosum L. Tafll›ca village, Meydanlar, open area, 1350 m, 16.06.1996, ÖE 1620. Carum carvi L. Tafll›ca village, Bakacak, open area, 1400 m, 22.06.1996, ÖE 1621. C. caucasicum (M.Bieb.) Boiss. Pimpinella affinis Ledeb. Tafll›ca village, Arganet, open forest, 1350 m, 29.06.1995, ÖE 1401. Foeniculum vulgare Mill. Tafll›ca village, Meydanlar, open forest, 1300 m, 10.08.1996, ÖE 1622. Ferulago latiloba Schischkin slope, Tafll›ca village, Gölgeli, roadside, mixed forest, 960 m, 23.05.1995, ÖE 1354. C. sanguinea (Wangerin) Chamb. L. subsp. cilicica F›st›kl› village, mixed forest, 560 m, 23.05.1995, ÖE 1355. E.Medit. m, Heracleum apiifolium Boiss. Tafll›ca village, Arganet, streamside, 1800 m, 08.09.1996, ÖE 1623. Torilis japonica (Houtt.) DC. Tafll›ca village, Meydanlar, open forest, shady area, 1240 m, 19.07.1995, ÖE 1403. Caucalis platycarpos L. Tafll›ca village, Meydanlar, meadow, 1200 m, 10.08.1996, ÖE 1624. Hatipo¤lu, mixed 23.05.1995, ÖE 1353. 1870 m, Centranthus longiflorus Stev. subsp. longiflorus Hatipo¤lu, roadside, 10.07.1996, ÖE 1626. 600 m, Ir-Tur. Valerianella lasiocarpa (Stev.) Betcke Tafll›ca village, Meydanlar, meadow, 1300 m, 06.06.1997, ÖE 1932. DIPSACACEAE forest, 760 m, Euro-Sib. Sambucus ebulus L. Tafll›ca village, Davütet, roadside, 1200 m, 06.07.1995, ÖE 1500. Euro-Sib. S. nigra L. Tafll›ca village, Ar›p›nar, roadside, 1900 m, 06.07.1995, ÖE 1501. Euro-Sib. Mount Keçi, mixed forest, 1950 m, 09.06.1995, ÖE 1502. Euro-Sib. V. orientale Pall. Saçinge hill, open forest, 1750 m, 12.07.1995, ÖE 1504. Eux. V. opulus L. Tafll›ca village, Meydanlar, roadside, 1400 m, 09.06.1995, ÖE 1503. Euro-Sib. Saçinge hill, roadside, 23.06.1995, ÖE 1505. Ir-Tur. C. mas L. Viburnum lantana L. 800 Valeriana alliariifolia Adams Cosm. Cornus sanguinea L. subsp. australis (C.A.Mey.) Jáv. CAPRIFOLIACEAE Mount Kurt, alpine slope, 2400 m, 20.07.1997, ÖE 1930. Mount Kurt, open area, alpine slope, 2600 m, 10.9.1995, ÖE 1499. VALERIANACEAE Eux. subsp. L. caucasica Pall. subsp. orientalis (Lam.) Chamb. & Long Endemic, LR (lc). Tafll›ca village, Meydanlar, mixed forest, 1400 m, ÖE 1399. Pall. Tafll›ca village, Ar›p›nar, stony slope, 1950 m, 18.07.1995, ÖE 1498. H. colchica (C.Koch) C.Koch Euro-Sib. Eryngium caeruleum M.Bieb. caucasica Cosm. CORNACEAE Astrantia maxima Pall. subsp. maxima 16 ARALIACEAE Hedera helix L. Eux. Hatipo¤lu, open 06.06.1997, ÖE 1931. Tafll›ca village, Köklü, open area, 1500 m, 16.06.1996, ÖE 1625. Lonicera caucasica Dipsacus pilosus L. Tafll›ca village, Davütet, roadside, 1450 m, 23.06.1995, ÖE 1507. Cephalaria aristata C.Koch F›st›kl› village, fieldside, 23.06.1995, ÖE 1506. Scabiosa columbaria columbaria var. columbaria 770 L. m, subsp. Mount Keçi, stony slope, 1950 m, 23.06.1995, ÖE 1508. S. caucasica M.Bieb. Tafll›ca village, Ar›p›nar, stony slope, 2300 m, 17.08.1995, ÖE 1509. COMPOSITAE (ASTERACEAE) Xanthium strumarium strumarium L. subsp. Tafll›ca village, Gökçeyurt, open place, 1100 m, 10.06.1996, ÖE 1627. Telekia speciosa (Schreb.) Baumg. Tafll›ca village, Bakacak, Picea Dietr. forest, moist soil, 1170 m, 15.07.1995, ÖE 1551. Euro-Sib. Ö. EM‹NA⁄AO⁄LU, R. ANfi‹N, Pallenis spinosa (L.) Cass. F›st›kl› village, fieldside, 08.04.1995, ÖE 1539. Tussilago farfara L. 250 m, Cosm. Inula helenium L. subsp. orgyalis (Boiss.) Grierson Mount Kurt, streamside, open area, 2650 m, 07.09.1995, ÖE 1534. Endemic, LR(nt), Eux. Helichrysum artvinense P.H.Davis & Kupicha Tafll›ca village, fiaflk›rat plateau, open area, dry soil, 2650 m, 27.07.1995, ÖE 1531. Endemic, EN. H. orientale (L.) DC. Tafll›ca village, Ar›p›nar, open area, 2350 m, 27.07.1995, ÖE 1532. Solidago virgaurea L. subsp. virgaurea Tafll›ca village, Köklü, streamside, 1500 m, 23.09.1995, ÖE 1547. Aster alpinus L. Tuzlu hill, alpine pasture, 2800 m, 10.06.1997, ÖE 1933. A. caucasicus Willd. Tafll›ca village, Ar›p›nar, open area, 1900 m, 24.08.1995, ÖE 1514. Eux. Bellis perennis L. Tafll›ca village, Bakacak, open area, 750 m, 12.03.1996, ÖE 1515. Euro-Sib. Doronicum orientale Hoffm. F›st›kl› village, 370 m, 12.03.1996, ÖE 1530. Senecio pandurifolius C.Koch F›st›kl› village, stony slope, 350 m, 14.04.1996, ÖE 1544. Eux. S. taraxacifolius (M.Bieb.) DC. var. taraxacifolius Tuzlu hill, rocky area, 2980 m, 09.09.1995, ÖE 1545. S. vernalis Waldst. & Kit. Tafll›ca village, Gölgeli, moist soil, shady area, 800 m, 04.06.1995, ÖE 1546. S. viscosus L. Tafll›ca village, Direkli, rocky slope, 1600 m, 20.07.1997, ÖE 1934. T. parthenium (L.) Sch.Bip. Tafll›ca village, Gölgeli, roadside, moist soil, 1300 m, 02.03.1996, ÖE 1556. Euro-Sib. Tripleurospermum oreades (Boiss.) Rech. var. oreades Petasites hybridus (L.) Gaertn. Tafll›ca village, Gökçeyurt, streamside, 1300 m, 12.06.1996, ÖE 1628. P. albus (L.) Gaertn. Tafll›ca village, Gölgeli, streamside, open forest, 1250 m, 04.04.1995, ÖE 1540. Euro-Sib. Tafll›ca village, Gökçeyurt, open area, wet soil, 1000 m, 06.08.1995, ÖE 1529. Euro-Sib. Anthemis calcarea Sosn. var. calcarea 2400 m, A. marschalliana Willd. pectinata (Boiss.) Grierson subsp. Tafll›ca village, fiaflk›rat plateau, alpine vegetation, 2800 m, 22.06.1996, ÖE 1629. Eux. F›st›kl› village, rocky slope, 650 m, 10.06.1996, ÖE 1630. Cousinia woronowii Bornm. Alacada¤, rocky slope, 20.07.1997, ÖE 1937. 1900 m, Endemic, LR (nt), Ir-Tur. Tafll›ca village, Mount Keçi, open area, 1950 m, 12.07.1995, ÖE 1513. Euro-Sib. Onopordum acanthium L. Tafll›ca village, Ar›p›nar, slope, 1700 m, 10.06.1997, ÖE 1938. A. cotula L. Tuzlu hill, alpine pasture, 2750 m, 20.07.1997, ÖE 1936. A. tinctoria L. var. tinctoria Tafll›ca village, Bakacak, roadside, 1100 m, 05.07.1995, ÖE 1511. A. tinctoria L. var. pallida DC. 1700 m, Achillea biserrata M.Bieb. Circium caucasicum (Adams) Petr. Mount Otluca, alpine vegetation, 2400 m, 04.08.1996, ÖE 1631. Eux (mt). C. pseudopersonata Boiss. & Bal. subsp. pseudopersonata Mount Keçi, open stony slope, 1800 m, 20.06.1997, ÖE 1939. Endemic, LR (lc), Eux. Tafll›ca village, Ar›p›nar, wet place, 1800 m, 11.07.1995, ÖE 1510. Leucanthemum vulgare Lam. 760 C. obvalatum (M.Bieb.) Fisch. Perikaya hill, roadside, 13.08.1995, ÖE 1524. Eux. m, Euro-Sib. Tanacetum macrophyllum (Waldst. & Kit.) Schultz Tafll›ca village, Demirci, roadside, 1450 m, 11.08.1995, ÖE 1548. Euro-Sib. Endemic, LR(cd). Arctium minus (hill) Bernh. subsp. pubens (Bab.) Arènes Endemic, CR. Hatipo¤lu, roadside, 19.07.1995, ÖE 1537. F›st›kl› village, roadside, rocky slope, 380 m, 14.04.1996, ÖE 1555. Artemisia vulgaris L. Eupotarium cannabinum L. Saçinge hill, roadside, 05.07.1995, ÖE 1512. Tafll›ca village, Köklü, open forest, 1200 m, 12.03.1996, ÖE 1554. T. fissurale (Sosn.) E.Hossain Eur-Sib. Mount Kurt, slope, 10.06.1997, ÖE 1935. Tafll›ca village, Konac›k, roadside, waste place, 1560 m, 11.08.1995, ÖE 1549. 2250 m, VU, Hyr-Eux. C. arvense (L.) Scop. subsp. arvense Tafll›ca village, Kanac›k, roadside, streamside, 1230 m, 14.10.1995, ÖE 1525. Carduus hamulosus hamulosus Ehrh. subsp. Tafll›ca village, Demirci, roadside, 1450 m, 02.06.1995, ÖE 1516. 17 The Flora of Hatila Valley National Park and its Close Environs (Artvin) Centaurea salicifolia M.Bieb. ex Willd. subsp. abbreviata C.Koch Tafll›ca village, Davütet, rocky and stony slope, 1370 m, 07.08.1995, ÖE 1519. Eux. Tafll›ca village, K›l›çl›, open forest, 1860 m, 13.08.1995, ÖE 1520. Eux. 650 m, C. hedgei Wagenitz Tuzlu hill, alpine slope, 2850 m, 06.06.1997, ÖE 1940. Endemic, VU. C. nigrifimbra (C.Koch) Sosn. Mount Otluca, subalpine slope, 2060 m, 21.08.1995, ÖE 1517. Eux (mt). Endemic, LR (lc), Ir-Tur. Tragopogon reticulatus Boiss. & Huet T. aureus Boiss. Endemic, LR(lc). Tafll›ca village, Meydanlar, stony slope, 1100 m, 14.04.1996, ÖE 1521. Endemic, LR(cd). F›st›kl› village, open area, dry soil, 300 m, 17.07.1995, ÖE 1557. Ir-Tur. Alacada¤, stony slope, dry soil, 1600 m, 10.06.1997, ÖE 1941. Echinops galaticus Freyn Tafll›ca village, Demirci, roadside, 1450 m, 09.08.1995, ÖE 1528. Eux. 750 m, L. crispus Vill. subsp. asper (Waldst. & Kit.) Röhl. var. asper Sonchus asper (L.) glaucescens (Jord.) Ball. subsp. Endemic, DD, Eux. H. subhastulatum (Zahn) Juxip Tuzlu hill, alpine pasture, 2800 m, 20.07.1997, ÖE 1944. Endemic, DD, Eux. Artvin:inter Selalet et Chinzart ad margines sylvae, Woronow. Endemic, EN, Eux. Cichorium intybus L. H. bornmuelleri Freyn Cosm. Scorzonera inaequiscapa Boiss. F›st›kl› village, dry soil, open area, 340 m, 07.06.1995, ÖE 1543. Endemic, LR(nt). hill Hieracium variegatisguamum (Zahn) Juxip Ir-Tur. Hatipo¤lu, cultivated fields, moist soil, 250 m, 16.04.1996, ÖE 1523. Pilosella x ruprechtii (Boiss.) Sell & West Tafll›ca village, Gölgeli, rocky slope, 1600 m, 12.08.1996, ÖE 1635. P. piloselloides (Vill.) Soják subsp. megalomastix (NP.) Sell & West Tafll›ca village, Demirci, open area, slope, 1560 m, 13.07.1995, ÖE 1541. Tafll›ca village, Direkli, mixed forest, 1850 m, 17.09.1995, ÖE 1542. H. diaphanoidiceps (Woronow & Zahn) Juxip E. pungens Trautv. var. pungens Endemic, DD, Eux. F›st›kl› village, grassy slope, 560 m, 17.07.1995, ÖE 1536. Tafll›ca village, Demirci, mixed forest, 1600 m, 28.06.1997, ÖE 1943. X. cylindraceum Sm. Artvin, prope stationem Gurdshani, 26.07.1911, Woronow. Prenanthes abietina (Boiss. & Ball.) Kirp. Cerat hill, open woods, 1600 m, 17.04.1996, ÖE 1634. Xeranthemum annuum L. H. subartvinense Juxip Leontodon asperimus (Willd.) Ball. Tafll›ca village, Gökçeyurt, rocky slope, 1200 m, 10.08.1996, ÖE 1633. C. woronowi Bornm. Artvin, in pineto supra Ghurdzan, Woronow. Endemic, DD, Eux. S. tomentosa L. Tafll›ca village, Ar›p›nar, open area, subalpine slope, 2100 m, 17.06.1995, ÖE 1553 Endemic, VU, Eux. 18 Saçinge hill, rocky slope, 1750 m, 09.08.1995, ÖE 1632. F›st›kl› village, open area, stony slope, 560 m, 15.06.1995, ÖE 1552. C. pecho Albov Hatipo¤lu, open forest, 09.08.1995, ÖE 1527. H. artvinense (Woronow & Zahn) Juxip Mount Otluca, rocky slope, 2050 m, 18.07.1997, ÖE 1942. C. simplicicaulis Boiss. & Huet F›st›kl› village, roadside, 21.07.1995, ÖE 1518. S. seidlitzii Boiss. Mount Otluca, rocky place, 2000 m, 10.08.1997, ÖE 1945. Endemic, LR (lc). H. subrosulatum Freyn & Sint. Tafll›ca village, Meydanlar, roadside, 1230 m, 08.08.1995, ÖE 1533. Endemic, DD. Eux. P. cacaliifolia (M.Bieb.) Beauverd Cerat hill, mixed forest, 1780 m, 10.10.1997, ÖE 1946. Eux. Mycelis muralis (L.) Dum. Tafll›ca village, Bakacak, open forest, 1230 m, 13.07.1995, ÖE 1538. Euro-Sib. Lapsana communis intermedia (M.Bieb.) Hayek L. subsp. Tafll›ca village, Meydanlar, mixed forest, 1370 m, 17.07.1995, ÖE 1535. Taraxacum buttleri Soest Tafll›ca village, Meydanlar, rocky and stony slope, 1100 m, 12.06.1995, ÖE 1550. Crepis paludosa (L.) Moench Tafll›ca village, K›l›çl›, open forest, 1700 m, 05.07.1995, ÖE 1526. Euro-Sib. CAMPANULACEAE Campanula latifolia L. Perikaya hill, slope, 10.06.1997, ÖE 1947. 1800 m, Euro-Sib. C. rapunculoides rapunculoides L. subsp. Tafll›ca village, Gökçeyurt, open area, wet soil, 850 m, 14.06.1995, ÖE 1488. Euro-Sib. Ö. EM‹NA⁄AO⁄LU, R. ANfi‹N, C. rapunculoides L. subsp. cordifolia (C.Koch) Damboldt Tafll›ca village, Bakacak, Picea Dietr. forest, stony slope, 920 m, 14.06.1996, ÖE 1489. C. alliariifolia Willd. Tafll›ca village, Direkli, roadside, 1830 m, 11.07.1995, ÖE 1484. Eux. C. collina Sims Mount Keçi, open slope, 1650 m, 10.06.1997, ÖE 1948. Eux. C. aucheri A.DC. Mount Kurt, alpine vegetation, 2700 m, 12.06.1996, ÖE 1636. Eux (mt). C. lactiflora M.Bieb. Tafll›ca village, fiaflk›rat plateau, mixed forest, 2400 m, 10.08.1995, ÖE 1485. Eux. C. stevenii M.Bieb. subsp. stevenii Tafll›ca village, fiaflk›rat plateau, open area, stony slope, 2200 m, 14.06.1996, ÖE 1490. R. caucasicum Pall. Tafll›ca village, fiaflk›rat plateau, subalpine slope, 2200 m, 16.05.1996, ÖE 1236. Eux. R. ponticum L. subsp. ponticum var. ponticum Tafll›ca village, Meydanlar, mixed forest, 1700 m, ÖE 1238. Eux. Tafll›ca village, Meydanlar, open forest, 1400 m, 12.03.1996, ÖE1241. Epigaea gaultherioides (Boiss. & Ball.) Takht. Tafll›ca village, fiaflk›rat plateau, subalpine slope, moist soil, 2230 m, 17.05.1996, ÖE 1234. Eux. P. elatior (L.) hill subsp. pallasii (Lehm.) W.W.Sm. & Forrest Tafll›ca village, fiaflk›rat plateau, subalpine slope, 2370 m, 12.04.1996, ÖE 1252. P. elatior (L.) hill subsp. pseudoelatior (Kusn.) W.W.Sm. & Forrest Mount Kurt, subalpine slope, 2400 m, 14.05.1996, ÖE 1253. VU, Eux (mt). P. auriculata Lam. Tuzlu hill, streamside, alpine slope, 2700 m, 14.05.1996, ÖE 1251. VU, Eux. Ir-Tur. Arbutus andrachne L. P. algida Adams Çoruh riverside, 14.05.1996, ÖE 1233. slope, 500 m, Medit. Vaccinium myrtillus L. Tafll›ca village, fiak›rat plateau, subalpine slope, 2170 m, 16.08.1996, ÖE 1243. Euro-Sib. C. pontica Albov V. arctostaphylos L. Tafll›ca village, Meydanlar, mixed forest, 1350 m, 19.07.1995, ÖE 1242. Eux. Eux. C. olympica Boiss. Pyrola media Sw. Perikaya hill, subalpine slope, 1950 m, 19.07.1995, ÖE 1486. Tafll›ca village, Meydanlar, roadside, 1300 m, 13.03.1996, ÖE 1255. Euro-Sib. R. x sochadzeae Kharadze & Davlian. Hyr-Eux. Tafll›ca village, Ar›p›nar, subalpine slope, 1970 m, 14.04.1996, ÖE 1487. P. vulgaris Huds. subsp. sibthorpii (Hoffmanns.) W.W.Sm. & Forrest Cerat hill, Picea Dietr. forest, 1570 m, 13.07.1995, ÖE 1235. Tuzlu hill, alpine vegetation, wet pasture, 2700 m, 10.07.1996, ÖE 1637. Cyclamen coum Mill. var. coum F›st›kl› village, Pinus pinea L. forest, 480 m, 13.03.1996, ÖE 1248. C. coum Miller (C.Koch) Medik. Lysimachia verticillaris Spreng. Tafll›ca village, Ar›p›nar, moist soil, 1750 m, 19.07.1995, ÖE 1250. Hyr-Eux. Anagallis arvensis L. var. arvensis Çoruh riverside, 350 m, 12.07.1995, ÖE 1247. Euro-Sib. EBENACEAE ERICACEAE Orthilia secunda (L.) House Diospyros lotus L. Tafll›ca village, Meydanlar, mixed forest, 1200 m, 16.05.1996, ÖE 1237. Eux. R. ungernii Trautv. Tafll›ca village, fiaflk›rat plateau, subalpine slope, 2550 m, 21.06.1995, ÖE 1240. VU, Eux. R. smirnovii Trautv. Tafll›ca village, fiaflk›rat plateau, subalpine slope, 2400 m, 16.05.1996, ÖE 1239. Endemic, EN, Eux. Tafll›ca village, Meydanlar, rocky area, acid soil, 1450 m, 17.07.1995, ÖE 1244. Monotropa hypopithys L. Cerat hill, mixed forest, 1370 m, 13.07.1995, ÖE 1245. PRIMULACEAE Primula vulgaris Huds. subsp. vulgaris Tafll›ca village, Meydanlar, wet soil, Picea Dietr. forest, 1400 m, 13.07.1995, ÖE 1254. Euro-Sib. caucasicum F›st›kl› village, rocky and stony slope, 370 m, 13.03.1996, ÖE 1249. Eux. Rhododendron luteum Sweet var. Hatipo¤lu, roadside, mixed forest, wet soil, 580 m, 17.08.1995, ÖE 1246. OLEACEAE Jasminium fruticans L. F›st›kl› village, open area, 350 m, 16.05.1996, ÖE 1462. Medit. Fraxinus angustifolia Vahl subsp. oxycarpa (M.Bieb.ex Willd.) Franco & Rocha Afonso Tafll›ca village, Gökçeyurt, roadside, 900 m, 19.09.1995, ÖE 1461. Euro-Sib. 19 The Flora of Hatila Valley National Park and its Close Environs (Artvin) Olea europaea L. var. sylvestris (Mill.) Lehr F›st›kl› village, open area, 350 m, 06.06.1996, ÖE 1638. Medit. Osmanthus decorus (Boiss. & Ball.) Kasapl›gil Tafll›ca village, Davütet, streamside, mixed forest, 1230 m, 16.03.1996, ÖE 1463. VU, Eux. APOCYNACEAE Trachomitum venetum (L.) Woodson subsp. sarmatiense (Woodson) Avet. Çoruh riverside, 260 m, 17.08.1996, ÖE 1408. E. Medit. Vinca herbacea Waldst. & Kit. F›st›kl› village, sunny stony slope, 470 m, 12.03.1996, ÖE 1409. GENTIANACEAE Centaurium erythraea Rafn subsp. erythraea Tafll›ca village, Meydanlar, stony slope, dry soil, 900 m, 27.07.1995, ÖE 1404. Euro-Sib. Gentiana asclepiadea L. Tafll›ca village, Demirci, roadside, mixed forest, 1500 m, 14.09.1995, ÖE 1405. Euro-Sib. G. pyrenaica L. Saçinge hill, open forest, 1650 m, 18.07.1997, ÖE 1949. Euro-Sib. G. septenfida Pall. Tafll›ca village, Ar›p›nar, open area in the Abies Mill. forest, 2150 m, 11.08.1995, ÖE 1406. Hyr-Eux (mt). G. verna L. subsp. pontica (Soltok) Hayek Tuzlu hill, alpine slope, 3000 m, 20.06.1997, ÖE 1950. Hyr-Eux (mt). Gentianella ciliata (L.) Borkh. subsp. blepharophora (Bordz.) Pritch. Saçinge hill, open area, 1450 m, 12.09.1995, ÖE 1407. Hyr-Eux (mt). 20 G. caucasea (G.Lodd. ex Sims) Holub C. montanum L. Tuzlu hill, subalpine pasture, 2300 m, 10.06.1997, ÖE 1951. Perikaya hill, alpine slope, 2200 m, 18.06.1997, ÖE 1952. Eux (mt). Euro-Sib. Swertia iberica Fisch. ex C.A.Mey. Lithospermum officinale L. Tuzlu hill, alpine vegetation, wet place, 2750 m, 10.07.1996, ÖE 1639. Tafll›ca village, fiaflk›rat plateau, slope, 2200 m, 10.07.1997, ÖE 1954. CONVOLVULACEAE Euro-Sib. Convolvulus cantabrica L. L. purpurocaeruleum L. F›st›kl› village, Pinus pinea L. stand, 350 m, 12.04.1995, ÖE 1414. C. arvensis L. Euro-Sib. F›st›kl› village, roadside, dry soil, 500 m, 06.06.1996, ÖE 1640. Cosm. Calystegia silvatica (Kit.) Griseb. Buglossoides arvensis (L.) Johnston Mount Kurt, slope, 18.07.1997, ÖE 1953. 2700 m, Echium vulgare L. Tafll›ca village, Davütet, roadside, 1450 m, 14.07.1995, ÖE 1413. F›st›kl› village, roadside, 13.06.1995, ÖE 1419. 750 m, Euro-Sib. CUSCUTACEAE Cuscuta epithymum epithymum Tafll›ca village, Gökçeyurt, roadside, 1100 m, 12.03.1996, ÖE 1421. (L.) L. var. Tafll›ca village, Gölgeli, roadside, on the Sambucus ebulus L., 1450 m, 14.07.1995, ÖE 1415. BORAGINACEAE Cerat hill, open forest, 1700 m, 18.07.1997, ÖE 1955. Ir-Tur. O. circinatum Riedl F›st›kl› village, roadside, 17.06.1995, ÖE 1426. Heliotropium ellipticum Ledeb. Tafll›ca village, Bakacak, roadside, dry soil, 950 m, 13.06.1995, ÖE 1420. Ir-Tur. 900 m, Endemic, EN, Eux. O. microcarpum Steven ex DC. F›st›kl› village, open forest, 470 m, 13.04.1996, ÖE 1427. Myosotis lazica Popov Tafll›ca village, Köklü, roadside, moist soil, 1100 m, 17.07.1995, ÖE 1423. VU, Eux. M. sylvatica Ehrh. ex Hoffm. subsp. cyanea Vestergr. Perikaya hill, open area, dry soil, 2200 m, 17.07.1995, ÖE 1424. M. laxa Lehm. subsp. caespitosa (Schultz) Hyl. ex Nordh. Tafll›ca village, Ar›p›nar, roadside, wet soil, 1800 m, 13.06.1995, ÖE 1422. Omphalodes cappadocica (Willd.) DC. F›st›kl› village, roadside, 12.03.1996, ÖE 1425. Onosma sericeum Willd. 750 m, Eux. Cynoglossum creticum Mill. Tafll›ca village, Ar›p›nar, subalpine vegetation, 2300 m, 26.07.1996, ÖE 1641. Ir-Tur. Cerinthe minor L. subsp. minor Tafll›ca village, Konac›k, Picea Dietr. forest, 1450 m, 15.07.1995, ÖE 1418. Euro-Sib. C. glabra Mill. Tafll›ca village, K›l›çl›, open area, 1900 m, 17.07.1995, ÖE 1417. Euro-Sib. Symphytum asperum Lepech. Tafll›ca village, Ar›p›nar, slope, 1550 m, 10.07.1997, ÖE 1956. Hyr-Eux. S. armeniacum Buckn. Saçinge hill, stony slope, 1800 m, 15.07.1997, ÖE 1956. Endemic, DD, Eux. Ö. EM‹NA⁄AO⁄LU, R. ANfi‹N, S. tauricum Willd. V. thapsus L. Tafll›ca village, Kanac›k, mixed forest, 1450 m, 12.05.1995, ÖE 1428. Euro-Sib. slope, 750 m, Eux. Brunnera macrophylla (Adams) Johnst. Tafll›ca village, Meydanlar, open area in the Picea Dietr. forest, 1350 m, 12.03.1996, ÖE 1416. VU, Eux. Anchusa azurea Mill. var. azurea Tafll›ca village, Ar›p›nar, waste place, 2200 m, 28.06.1996, ÖE 1642. Nonea versicolor (Steven) Sweet Saçinge hill, stony slope, 1750 m, 06.06.1996, ÖE 1643. N. intermedia Ledb. Mount Keçi, alpine slope, 1950 m, 10.06.1997, ÖE 1957. Alkanna cordifolia C.Koch Çoruh: Artvin, 500 m, Stainton 8184. Endemic, LR (lc), Ir-Tur. SOLANACEAE Solanum dulcamara L. Tafll›ca village, Ar›p›nar, streamside, 1550 m, 20.07.1997, ÖE 1958. Physalis alkekengi L. Tafll›ca village, Bakacak, roadside, wet soil, 1450 m, 16.10.1995, ÖE 1412. Atropa belladonna L. Tafll›ca village, K›l›çl›, roadside, 1700 m, 13.07.1995, ÖE 1410. Euro-Sib. Datura stramonium L. F›st›kl› village, open area, 600 m, 06.06.1996, ÖE 1644. Cosm. Hyoscyamus niger L. Hatipo¤lu, roadside, open area, 850 m, 13.04.1996, ÖE 1411. SCROPHULARIACEAE Verbascum gracilescens Hub.-Mor. Tafll›ca village, Meydanlar, roadside, 1050 m, 17.07.1995, ÖE 1475. Endemic, EN, Ir-Tur. Tafll›ca village, Ar›p›nar, stony slope, dry soil, 12.07.1999, ÖE 1645. V. saccatum C.Koch Trachystemon orientalis (L.) G.Don Hatipo¤lu, open 07.03.1996, ÖE 1429. V. anagallis-aquatica L. Cosm. Tafll›ca village, Demirci, roadside, 1300 m, 01.07.1995, ÖE 1476. Ir-Tur. V. artvinense Wulf Endemic, DD, Eux. Scrophularia chrysantha Jaub. & Spach Tafll›ca village, Davütet, on rocks, 1600 m, 06.04.1996, ÖE 1473. Eux. S. kotschyana Benth. S. umbrosa Dum. V. liwanensis C.Koch Çoruh riverside, on the rocks, 250 m, 03.03.1995, ÖE 1479. VU, Eux. V. orientalis Mill. subsp. orientalis Tafll›ca village, Ar›p›nar, roadside, 1660 m, 06.06.1995, ÖE 1481. Tafll›ca village, fiaflk›rat plateau, alpine slope, 2450 m, 01.04.1996, ÖE 1482. Eux. Tafll›ca village, Meydanlar, streamside, 1400 m, 20.07.1997, ÖE 1959. Linaria genistifolia (L.) Mill. subsp. artvinensis P.H.Davis Çoruh: Artvin, 400 m, Stainton 8290. V. officinalis L. Tafll›ca village, Demirci, open forest, 1350 m, 06.06.1995, ÖE 1480. Melampyrum arvense L. var. arvense Tafll›ca village, Direkli, open forest, 1400 m, 16.05.1996, ÖE 1467. Endemic, VU, Eux. L. vulgaris Mill. Tafll›ca village, Meydanlar, meadow, 1300 m, 20.08.1997, ÖE 1960. L. subsp. Tafll›ca village, Meydanlar, open forest, 1150 m, 21.06.1995, ÖE 1464. Euro-Sib. Veronica gentianoides Vahl. Tuzlu hill, alpine pasture, 2850 m, 10.06.1997, ÖE 1961. Hyr-Eux (mt). V. verna L. Tuzlu hill, alpine pasture, 2900 m, 10.06.1997, ÖE 1962. Euro-Sib. V. persica Poir. F›st›kl› village, open area, 450 m, 12.03.1996, ÖE 1483. V. ceratocarpa C.A.Mey. Tafll›ca village, Meydanlar, meadow, 1300 m, 08.06.1995, ÖE 1477. VU, Hyr-Eux. F›st›kl› village, open forest, 350 m, 06.04.1996, ÖE 1478. V. peduncularis M.Bieb. Tafll›ca village, Gölgeli, open area, 800 m, 06.04.1996, ÖE 1474. ferruginea V. jacquinii Baumg. Euro-Sib. Artvin, Lomaflen, Machljuse, 09.06.1909, Nesterow (holo. LE) Digitalis ferruginea Tafll›ca village, Ar›p›nar, streamside, 1500 m, 18.07.1997, ÖE 1963. Euro-Sib. M. arvense L. var elatius Boiss. Tafll›ca village, Ar›p›nar, open area, 1750 m, 08.07.1995, ÖE 1468. Endemic, LR(nt). Euphrasia hirtella Jordan ex Reuter Alacada¤, slope, 1900 m, 12.07.1997, ÖE 1964. Euro-Sib. E. pectinata Ten. Tafll›ca village, fiaflk›rat plateau, subalpine slope, 2100 m, 13.05.1996, ÖE 1465. Euro-Sib. Pedicularis caucasica M.Bieb. Mount Kurt, alpine slope, 2700 m, 10.06.1997, ÖE 1965. Hyr-Eux (mt) P. atropurpurea Nordm. Tafll›ca village, fiaflk›rat plateau, subalpine slope, 2300 m, 16.08.1995, ÖE 1469. VU, Eux (mt). 21 The Flora of Hatila Valley National Park and its Close Environs (Artvin) P. condensata M.Bieb. Tafll›ca village, Demirci, slope, 1700 m, 08.07.1995, ÖE 1470. Eux. P. nordmanniana Bunge Perikaya hill, sublpine slope, 1900 m, 20.07.1997, ÖE 1966. Eux (mt). Rhinanthus angustifolius C.C.Gmel. subsp. grandiflorus (Wallr.) D.A.Webb Tafll›ca village, Meydanlar, meadow, wet soil, 1250 m, 05.06.1995, ÖE 1471. subsp. Galeobdolon luteum Hudson subsp. montanum (Pers.) R.Mill Tafll›ca village, Demirci, rocky slope, 1350 m, 04.06.1995, ÖE 1455. Tafll›ca village, Köklü, meadow, 1500 m, 28.06.1997, ÖE 1972. Teucrium chamaedrys trapenzuntinus Rech. L. Eux. Galeopsis bifida Boem. T. orientale L. Tafll›ca village, Gökçeyurt, roadside, 1050 m, 07.09.1995, ÖE 1456. Ir-Tur. Tafll›ca village, Gölgeli, stony slope, rocky area, 850 m, 20.09.1995, ÖE 1457. Rhynchocorys orientalis (L.) Benth. Tafll›ca village, fiaflk›rat plateau, meadow, 2000 m, 08.07.1997, ÖE 1967. F›st›kl› village, stony slope, 600 m, 18.06.1997, ÖE 1971. R. stricta (C.Koch) Albov Tafll›ca village, Gökçeyurt, open area, meadow, 950 m, 05.06.1995, ÖE 1472. Eux. Lathraea squamaria L. Tafll›ca village, Köklü, stony slope, 1230 m, 14.04.1996, ÖE 1466. OROBANCHACEAE Orobanche bungeana Beck Tafll›ca village, Ar›p›nar, roadside, 1550 m, 10.06.1997, ÖE 1968. Hyr-Eux. VERBENACEAE Verbena officinalis L. Tafll›ca village, Ar›p›nar, roadside, 1800 m, 19.08.1995, ÖE 1430. Cosm. LABIATAE (LAMIACEAE) Ajuga orientalis L. Tuzlu hill, alpine slope, 2900 m, 16.06.1996, ÖE 1431. A. genevensis L. Mount Kurt, alpine slope, 2700 m, 10.06.1997, ÖE 1969. Euro-Sib. A. reptans L. Tafll›ca village, Meydanlar, meadow, 1350 m, 20.06.1997, ÖE 1970. Euro-Sib. S. orientalis L. subsp. orientalis Tuzlu hill, alpine slope, 2780 m, 12.08.1995, ÖE 1449. Ir-Tur. purpureum var. Tafll›ca village, Direkli, mixed forest, moist soil, 1480 m, 14.06.1995, ÖE 1437. Euro-Sib. L. album L. Euro-Sib. L. crinitum Montbret & Aucher ex Benth. Mount Otluca, Picea Dietr. forest, 1850 m, 12.09.1995, ÖE 1435. Eux. L. tomentosum tomentosum Ballota nigra L. subsp. nigra Tafll›ca village, Köklü, open forest, 28.06.1997, ÖE 1974. Sideritis armeniaca Bornm. Endemic, LR(cd), Ir-Tur. S. montana L. subsp. montana F›st›kl› village, open area, 330 m, 12.05.1995, ÖE 1451. Medit. Stachys sylvatica L. Tafll›ca village, Meydanlar, mixed forest, 1350 m, 07.09.1995, ÖE 1454. Euro-Sib. Tafll›ca village, Ar›p›nar, rocky slope, 1670 m, 12.04.1996, ÖE 1434. Willd. var. F›st›kl› village, stony slope, 450 m, 24.08.1995, ÖE 1439. Ir-Tur. L. sulfureum Hausskn. & Sint ex R.B.Mill. Mount Kurt, in crevices on rocks, 2880 m, 16.07.1995, ÖE 1438. Endemic, DD, Ir-Tur. L. galactophyllum Boiss. & Reut. Tafll›ca village, Ar›p›nar, roadside, 1670 m, 18.07.1996, ÖE 1436. Endemic, LR(lc), Ir-Tur. 22 L. Euro-Sib. Tafll›ca village, Demirci, stony slope, 1350 m, 03.09.1995, ÖE 1450. Eux. Lamium purpureum Leonurus quinquelobatus Gilib. Tafll›ca village, Ar›p›nar, meadow, 1600 m, 10.06.1997, ÖE 1973. T. polium L. Scutellaria albida L. subsp. colchica (Rech.) Edm. Eux. Tafll›ca village, Kanac›k, slope, 1500 m, 20.06.1997, ÖE 1973. S. iberica M.Bieb. subsp. iberica var. iberica Hatipo¤lu, open forest, 12.05.1995, ÖE 1452. 550 m, Ir-Tur. S. macrantha (C.Koch) Stearn Mount Kurt, mixed forest, 2350 m, 07.09.1995, ÖE 1453. Eux. Prunella vulgaris L. Tafll›ca village, Ar›p›nar, fieldside, 1920 m, 12.08.1995, ÖE 1445. Euro-Sib. P. laciniata (L.) L. Tafll›ca village, Arganet, Pinus L. forest, 1200 m, 06.06.1997, ÖE 1975. Origanum rotundifolium Boiss. Tafll›ca village, Meydanlar, roadside, 1350 m, 14.06.1995, ÖE 1442. Eux. Ö. EM‹NA⁄AO⁄LU, R. ANfi‹N, O. vulgare L. subsp. vulgare Tafll›ca village, Demirci, slope, dry soil, 1280 m, 14.06.1995, ÖE 1444. Euro-Sib. O. vulgare L. subsp. viride (Boiss.) Hayek Tafll›ca village, Meydanlar, roadside, 1270 m, 14.06.1995, ÖE 1443. Satureja spicigera (C.Koch) Boiss. Tafll›ca village, Gölgeli, moist soil, 1250 m, 03.09.1995, ÖE 1448. Eux. Calamintha grandiflora (L.) Moench Tafll›ca village, Demirci, roadside, moist soil, 1350 m, 15.07.1995, ÖE 1432. Euro-Sib. Clinopodium vulgare L. subsp. vulgare Tafll›ca village, Arganet, stony slope, 1760 m, 13.06.1995, ÖE 1433. Euro-Sib. Micromeria elliptica C.Koch Çoruh riverside, 350 m, 18.07.1997, ÖE 1976. Thymus fallax Fisch. & Mey. Tafll›ca village, Köklü, stony slope, dry soil, 1450 m, 20.09.1995, ÖE 1458. Ir-Tur. T. praecox Opiz subsp. grossheimii (Ronniger) Jalas var. grossheimii Tafll›ca village, Ar›p›nar, roadside, open area, 1900 m, 17.08.1995, ÖE 1460. T. longicaulis C.Presl subsp. longicaulis var. subisophyllus (Borbás) Jalas Tafll›ca village, Demirci, roadside, open area, 1350 m, 17.08.1995, ÖE 1459. Mentha pulegium L. Tafll›ca village, Meydanlar, streamside, wet soil, 1030 m, 14.06.1995, ÖE 1441. M. longifolia (L.) Huds. subsp. longifolia Tafll›ca village, Demirci, streamside, 1470 m, 14.06.1995, ÖE 1440. Alacada¤, stony slope, 20.06.1997, ÖE 1977. m, Perikaya hill, mixed forest, dry soil, 1950 m, 12.07.1995, ÖE 1361. Euro-Sib. S. pachystachys Trautv. BUXACEAE Tafll›ca village, fiaflk›rat plateau, subalpine vegetation, 2000 m, 06.07.1997, ÖE 1978. Ir-Tur. Saçinge hill, roadside, 06.06.1997, ÖE 1979. Buxus sempervirens L. F›st›kl› village, mixed forest, stony slope, 350 m, 12.03.1996, ÖE 1360. Euro-Sib. S. sclarea L. 1350 m. S. verticillata L. subsp. verticillata Tafll›ca village, Demirci, mixed forest, 1450 m, 14.07.1995, ÖE 1447. URTICACEAE Urtica dioica L. Tafll›ca village, Gölgeli, roadside, open forest, 1050 m, 13.07.1995, ÖE 1149. Euro-Sib. PLANTAGINACEAE MORACEAE Plantago major L. subsp. major Morus alba L. Tafll›ca village, Ar›p›nar, open area, waste place, 1800 m, 28.06.1996, ÖE 1646. P. lanceolata L. Tafll›ca village, Gökçeyurt, roadside, 890 m, 13.05.1995, ÖE 1460. THYMELAEACEAE Daphne glomerata Lam. Tuzlu hill, subalpine slope, 2000 m, 13.07.1995, ÖE 1348. LORANTHACEAE Viscum album L. subsp. album Tafll›ca village, Meydanlar, Pinus L. forest, 1300 m, 13.05.1996, ÖE 1356. EUPHORBIACEAE Mercurialis perennis L. Tafll›ca village, Meydanlar, meadow, 1200 m, 10.07.1996, ÖE 1647. Euro-Sib. Euphorbia peplis L. Tafll›ca village, Gölgeli, stony slope, 900 m, 16.08.1995, ÖE 1363. Medit. E. stricta L. Tafll›ca village, Direkli, Picea Dietr. forest, 1650 m, 12.07.1995, ÖE 1364. Euro-Sib. Salvia rosifolia Sm. E. paralias L. Endemic, (LR (lc), Ir-Tur. 1700 Endemic, LR (cd), Ir-Tur. Eux. Tafll›ca village, Meydanlar, mixed forest, 1300 m, 12.07.1995, ÖE 1446. E. amygdaloides L. var amygdaloides S. huberi Hedge F›st›kl› village, open area, dry soil, 350 m, 16.08.1995, ÖE 1362. Medit. Tafll›ca village, Köklü, fieldside, 1360 m, 14.07.1995, ÖE 1148. Ficus carica L. subsp. carica F›st›kl› village, open area in the Pinus pinea L. stand, 570 m, 13.07.1995, ÖE 1146, Çoruh riverside, 250 m, 16.05.1996, ÖE 1147. ULMACEAE Ulmus glabra Huds. Tafll›ca village, Meydanlar, mixed forest, moist soil, 1250 m, 13.07.1995, ÖE 1144. Euro-Sib. U. minor Mill. subsp. minor Hatipo¤lu, roadside, 15.07.1995, ÖE 1145. 800 m, JUGLANDACEAE Juglans regia L. Tafll›ca village, Meydanlar, mixed forest, streamside, 1370 m, 13.07.1995, ÖE 1150. PLATANACEAE Platanus orientalis L. Çoruh riverside, 300 m, 16.07.1996, ÖE 1143. FAGACEAE Fagus orientalis Lipsky Tafll›ca village, fiaflk›rat plateau, subalpine slope, moist soil, 2250 m, 13.07.1995, ÖE 1152. Euro-Sib. Castanea sativa Mill. Tafll›ca village, Ar›p›nar, mixed forest, 1650 m, 05.05.1996, ÖE 1151. Euro-Sib. 23 The Flora of Hatila Valley National Park and its Close Environs (Artvin) Quercus hartwissiana Steven S. caucasica Andersson Tafll›ca village, Meydanlar, mixed forest, 1550 m, 13.07.1995, ÖE 1491. Eux. Eux. Q. petraea (Matt.) Liebl. subsp. iberica (Steven ex M.Bieb.) Krassiln. F›st›kl› village, roadside, mixed forest, 350 m, 13.07.1995, ÖE 1153. BETULACEAE forest, 850 m, Euro-Sib. C. orientalis Mill. F›st›kl› village, mixed forest, 590 m, 13.07.1995, ÖE 1159. Ostrya carpinifolia Scop. Tafll›ca village, Gökçeyurt, mixed forest, roadside, 1300 m, 16.06.1995, ÖE 1161. Medit. Corylus avellana L. var. avellana Tafll›ca village, Bakacak, roadside, 350 m, 16.08.1995, ÖE 1160. Betula pendula Roth Tafll›ca village, fiaflk›rat plateau, roadside, 1950 m, 14.05.1995, ÖE 1156. B. litwinowii Doluch. Tafll›ca village, fiaflk›rat plateau, with shrubs in the mixed forest as follows: Rhododendron caucasicum Pallas, Sorbus aucuparia L., 2200 m, 14.04.1996, ÖE 1155. Eux. B. recurvata (I.V.Vassil) I. V.Vassil Tafll›ca village, fiaflk›rat plateau, with shrubs in the mixed forest as follows: Rhododendron L., Lonicera L., Sorbus aucuparia L., 1350 m, 21.05.1996, ÖE 1157. Eux. Alnus glutinosa (L.) Gaertn. subsp. barbata (C.A.Mey.) Yalt. Tafll›ca village, Gölgeli, moist soil, along the Hatila streamside, 1100 m, 14.04.1996, ÖE 1154. Eux. Saçinge hill, open forest, 1800 m, 12.04.1996, ÖE 1203. Euro-Sib. Tafll›ca village, Ar›p›nar, roadside, streamside, 1890 m, 21.04.1996, ÖE 1204. Eux. Populus tremula L. Tafll›ca village, fiaflk›rat plateau, subalpine slope, 2450 m, 12.06.1995, ÖE 1200. RUBIACEAE Crucianella gilanica Trin. subsp. pontica (Ehrend.) Ehrend. Tafll›ca village, Bakacak, stony slope, 1075 m, 23.06.1995, ÖE 1492. Eux. Asperula orientalis Boiss. & Hohen. Tafll›ca village, Gölgeli, roadside, 900 m, 13.05.1995, ÖE 1491. Ir-Tur. Galium rotundifolium L. Tafll›ca village, Meydanlar, open forest, 1270 m, 18.07.1995, ÖE 1494. Euro-Sib. G. odoratum (L.) Scop. Tafll›ca village, Bakacak, mixed forest, 950 m, 16.05.1995, ÖE 1493. Euro-Sib. G. uliginosum L. F›st›kl› village, moist soil, 760 m, 25.06.1995, ÖE 1495. VU, Euro-Sib. G. verum L. subsp. verum Tafll›ca village, Köklü, riverside, 1430 m, 07.07.1995, ÖE 1496. Euro-Sib. Cruciata taurica (Pall. ex Willd.) Ehrend. Tafll›ca village, Ar›p›nar, stony slope, 1500 m, 20.06.1997, ÖE 1980. SALICACEAE Ir-Tur. Salix alba L. Rubia tinctorum L. Tafll›ca village, Gölgeli, streamside, 850 m, 12.10.1995, ÖE 1201. Euro-Sib. 24 S. caprea L. S. pseudomedemii E.Wolf Carpinus betulus L. Hatipo¤lu, mixed 13.07.1995, ÖE 1158. Tafll›ca village, Ar›p›nar, roadside, 1600 m, 12.05.1996, ÖE 1202. Tafll›ca village, Bakacak, roadside, 970 m, 16.05.1996, ÖE 1497. Ir-Tur. MONOCOTYLEDONAE ALISMATACEAE Alisma plantago-aquatica L. Hatila streamside, 1500 m, 06.06.1996, ÖE 1648. Euro-Sib. LILIACEAE Smilax excelsa L. Hatipo¤lu, mixed 08.05.1995, ÖE 1586. forest, 750 m, Eux. Ruscus aculeatus L. var. angustifolius Boiss. F›st›kl› village, open area, 250 m, 03.03.1996, ÖE 1581. Polygonatum verticillatum (L.) All. Tafll›ca village, Meydanlar, mixed forest, wet soil, 1300 m, 08.06.1996, ÖE 1649. Euro-Sib. Allium gramineum C.Koch Tafll›ca village, Ar›p›nar, subalpine slope, stony slope, 2200 m, 10.07.1996, ÖE 1650. A. rollovii Grossh. Perikaya hill, stony slope, 2000 m, 20.06.1997, ÖE 1981. Scilla monanthos C.Koch Tafll›ca village, fiaflk›rat plateau, subalpine slope, 1900 m, 03.03.1996, ÖE 1583. Eux. S. siberica Haw. (Grossh.) Mordak subsp. armena Tuzlu hill, alpine slope, 2500 m, 07.03.1996, ÖE 1584. Ir-Tur. S. leepi Speta F›st›kl› village, rocky slope, 650 m, 03.03.1996, ÖE 1582. Endemic, LR(nt), Ir-Tur. Ornithogalum oligophyllum E.D.Clarke Tuzlu hill, alpine pasture, 3000 m, 20.06.1997, ÖE 1982. O. platyphylum Boiss. Tafll›ca village, Köklü, open area, 1500 m, 09.05.1996, ÖE 1580. Ir-Tur. O. umbellatum L. F›st›kl› village, dry 14.04.1995, ÖE 1579. soil, 350 m, Ö. EM‹NA⁄AO⁄LU, R. ANfi‹N, Muscari coeleste Fomin Gladiolus atroviolaceus Boiss. Tafll›ca village, fiaflk›rat plateau, open area, 2200 m, 12.03.1996, ÖE 1577. Endemic, LR(lc), Ir-Tur. slope, 650 m, M. neglectum Guss. Tafll›ca village, Gölgeli, roadside, 450 m, 12.03.1996, ÖE 1578. Lilium carniolicum Bernh. ex W.Koch subsp. ponticum (C.Koch) P.H.Davis & Hend. var. ponticum Tafll›ca village, Ar›p›nar, open Picea Dietr. forest, 1750 m, 15.05.1995, ÖE 1574. VU. L. carniolicum Bernh. ex W.Koch subsp. ponticum (C.Koch) P.H.Davis & Hend. var. artvinense (Miscz.) P.H.Davis & Hend. Tafll›ca village, Ar›p›nar, open Picea Dietr. forest, 2000 m, 14.07.1995, ÖE 1575. Endemic, EN, Eux. Gagea reticulata (Pall.) Schult. F›st›kl› village, open area, 750 m, 12.03.1996, ÖE 1572 Ir-Tur. G. bulbifera (Pall.) Schult. Saçinge hill, open area, 1870 m, 04.04.1996, ÖE 1571 Euro-Sib. G. tenuissima Miscz. F›st›kl› village, stony slope, 370 m, 12.03.1996, ÖE 1573. Endemic, DD, Eux. Veratrum album L. Tafll›ca village, fiaflk›rat plateau, alpine vegetation, 2700 m, 10.07.1996, ÖE 1651. Euro-Sib. Colchicum umbrosum Steven Tafll›ca village, fiaflk›rat plateau, subalpine slope, 2270 m, 12.10.1995, ÖE 1570. Eux. IRIDACEAE Crocus biflorus Mill. subsp. artvinensis (J.Phil.) Mathew Tuzlu hill, alpine vegetation, 3000 m, 10.04.1996, ÖE 1652. Endemic, EN, Euro-Sib. Tafll›ca village, Meydanlar, open Picea Dietr. forest, 1350 m, 02.05.1996, ÖE 1585. Ir-Tur. M. armeniacum Leichtlin ex Baker Hatipo¤lu, rocky 12.03.1996, ÖE 1576. DIOSCOREACEAE Juncus inflexus L. Cephalanthera rubra (L.) Rich. Tafll›ca village, Ar›p›nar, mixed forest, 1650 m, 10.06.1997, ÖE 1983. C. longifolia (L.) Fritsch Tafll›ca village, Ar›p›nar, open forest, 1700 m, 10.06.1996, ÖE 1653. Euro-Sib. Çoruh: Tütüncüler, 35 km from ArtvinÇoruh River junction, 1700 m, Kit-Tan 1290. J. effusus L. Tafll›ca village, Ar›p›nar, streamside, wet soil, 1950 m, 13.07.1995, ÖE 1558. Cosm. J. filiformis L. Epipactis helleborine (L.) Crantz Tafll›ca village, Gökçeyurt, open forest, 900 m, 01.07.1997, ÖE 1984. E. microphylla (Ehrh.) Sw. Tafll›ca village, Direkli, open area, wet soil, 1470 m, 18.07.1995, ÖE 1588. Limodorum abortivum (L.) Sw. Tafll›ca village, Ar›p›nar, mixed forest, 1600 m, 01.07.1997, ÖE 1985. Goodyera repens (L.) R.Br. Saçinge hill, mixed forest, 1750 m, 10.06.1997, ÖE 1986. Platanthera bifolia (L.) Rich. Çoruh: above Artvin, 2000 m, D 29705. Tafll›ca village, Bakacak, wet grassland, 1500 m, 22.08.1996, ÖE 1656. J. bufonius L. Tafll›ca village, fiaflk›rat plateau, wet place, 2200 m, 01.07.1997, ÖE 1988. Cosm. Luzula forsteri (Sm.) DC. Tafll›ca village, Meydanlar, open area, wet place, 1250 m, 1150 m, 20.08.1996, ÖE 1657. Euro-Sib. L. sylvatica (Huds.) Gaudin Tafll›ca village, Ar›p›nar, damp slope, 1600 m, 01.07.1997, ÖE 1989. Euro-Sib. Euro-Sib. P. chlorantha (Custer) Rchb. Tafll›ca village, Meydanlar, open Picea Dietr. forest, 1200 m, 17.07.1995, ÖE 1589. Orchis purpurea Huds. L. pseudosudetica V.I.Krecz. (V.I.Krecz.) Tuzlu hill, alpine pasture, 3000 m, 06.06.1997, ÖE 1990. Eux (mt). Tafll›ca village, Gölgeli, open forest, 1560 m, 10.06.1996, ÖE 1654. Euro-Sib. (Steud.) Tafll›ca village, Demirci, slope, 1450 m, 12.03.1995, ÖE 1587. Eux. D. euxina (Nevski) H.Baumann & Künkele var. euxina Tafll›ca village, Ar›p›nar, wet meadow, 1600 m, 01.07.1997, ÖE 1987. Eux. Tafll›ca village, Meydanlar, mixed forest, 1200 m, 28.06.1996, ÖE 1655. JUNCACEAE ORCHIDACEAE Dactilorhiza urvilleana Baumann & Künkele Tamus communis L. subsp. communis L. campestris (L.) DC. Tafll›ca village, Ar›p›nar, roadside, 1800 m, 03.05.1996, ÖE 1559. Euro-Sib. CYPERACEAE Carex spicata Huds. Saçinge hill, roadside, 01.07.1997, ÖE 1991. 1500 m, Euro-Sib. C. echinata Murray Çoruh: Artvin to Atila, 1600-2000 m, Kit-Tan 1068. Euro-Sib. 25 The Flora of Hatila Valley National Park and its Close Environs (Artvin) C. rostrata Stokes Tafll›ca village, fiaflk›rat plateau, 2000 m, marshy place, 10.08.1997, ÖE 1992. Euro-Sib. B. tectorum L. F. drymeja Mert. & Koch Tafll›ca village, Meydanlar, meadow, 1270 m, 10.06.1996, ÖE 1661. Euro-Sib. B. erectus Hudson C. sylvatica Huds. subsp. latifrons (V.I.Krecz) O.Nilsson Mount Keçi, dry 20.07.1997, ÖE 1996. Tafll›ca village, Ar›p›nar, open forest, moist soil, 1750 m, 03.05.1996, ÖE 1560. Avena sterilis L. subsp. ludoviciana (Durieu) Gillet Eux. C. digitata L. Tafll›ca village, fiaflk›rat plateau, grassland, 2200 m, 01.07.1997, ÖE 1993. Eux. C. atrata L. subsp. atrata Saçinge hill, meadow, wet place, 1700 m, 02.08.1996, ÖE 1658. Euro-Sib (mt). C. nigra (L.) Reichard subsp. nigra Perikaya hill, waste place, 1900 m, 02.08.1996, ÖE 1659. GRAMINEAE (POACEAE) Brachypodium pinnatum (L.) P.Beauv. Tafll›ca village, Meydanlar, meadow, 1200 m, 22.07.1996, ÖE 1659. Euro-Sib. Elymus nodosus (Nevski) Melderis subsp. sinuatus (Nevski) Melderis Tafll›ca village, Ar›p›nar, stony slope, 1600 m, 18.07.1997, ÖE 1994. Ir-Tur. E. repens (L.) Gould subsp. repens Tafll›ca village, Bakacak, meadow, 1500 m, 02.08.1996, ÖE 1660. Triticum monococcum L. Tafll›ca village, Arganet, open forest, 970 m, 14.04.1996, ÖE 1569. Hordeum geniculatum All. Tafll›ca village, Demirci, meadow, wet soil, 1400 m, 13.07.1995, ÖE 1565. Euro-Sib. Bromus racemosus L. Tafll›ca village, Meydanlar, wet meadow, 1250 m, 28.07.1997, ÖE 1995. Euro-Sib. B. japonicus Thunb. subsp. japonicus Tafll›ca village, fiaflk›rat plateau, meadow, 2300 m, 15.05.1996, ÖE 1561. 26 Tafll›ca village, Bakacak, mixed forest, 1000 m, 15.07.1995, ÖE 1564. soil, 1700 m, Tafll›ca village, Gökçeyurt, meadow, 1150 m, 22.08.1996, ÖE 1662. Trisetum rigidum (M.Bieb.) Roem. & Schult. Mount Keçi, slope, 16.06.1996, ÖE 1568. 1750 m, F. djimilensis Boiss. & Bal. Tuzlu hill, wet alpine meadow, 2400 m, 10.07.1997, ÖE 2003. Euro-Sib (mt). Lolium rigidum Gaudin var. rigidum Tafll›ca village, Gökçeyurt, meadow, wet soil, 800 m, 19.07.1995, ÖE 1566. Poa annua L. Tafll›ca village, Meydanlar, meadow, 1200 m, 10.07.1996, ÖE 1666. Ir-Tur. Cosm. T. flavescens (L.) P.Beauv. P. trivialis L. Mount Kurt, alpine pasture, 2700 m, 01.07.1997, ÖE 1997. Euro-Sib. Deschampsia caespitosa (L.) P.Beauv. Saçinge hill, subalpine vegetation, 1950 m, 10.06.1996, ÖE 1663. Calamagrostis arundinaceae (L.) Roth Tafll›ca village, Ar›p›nar, slope, 1800 m, 22.08.1996, ÖE 1664. Euro-Sib. Agrostis planifolia C.Koch Tuzlu hill, alpine pasture, 2750 m, 10.08.1997, ÖE 1998. Ir-Tur. A. stolonifera L. Tafll›ca village, Meydanlar, wet place, 1300 m, 01.07.1997, ÖE 1999. Alopecurus arundinaceus Poir. Perikaya hill, wet meadow, 2100 m, 10.06.1997, ÖE 2000. Phleum alpinum L. Tuzlu hill, wet alpine meadow, 2700 m, 20.07.1997, ÖE 2001. Euro-Sib. P. pratense L. Tafll›ca village, Bakacak, open area, wet place, 1500 m, 07.07.1995, ÖE 1665. Euro-Sib. Festuca gigantea (L.) Vill. Tafll›ca village, Meydanlar, wet meadow, 1200 m, 01.07.1997, ÖE 2002. Euro-Sib. Tafll›ca village, Meydanlar, meadow, 1300 m, 01.07.1997, ÖE 2004. P. pratensis L. Tafll›ca village, Meydanlar, meadow, 1230 m, 10.07.1996, ÖE 1667. P. nemoralis L. Tafll›ca village, Meydanlar, meadow, 1230 m, 10.07.1996, ÖE 1668. P. sterilis M.Bieb. Tafll›ca village, Ar›p›nar, meadow, 1500 m, 20.08.1997, ÖE 2005. P. bulbosa L. Tafll›ca village, Ar›p›nar, open area, dry soil, 1700 m, 15.05.1996, ÖE 1567. Dactylis glomerata L. subsp. hispanica (Roth) Nyman Tafll›ca village, Meydanlar, meadow, 1230 m, 10.07.1996, ÖE 1669. Cynosurus cristatus L. Tafll›ca village, Köklü, open Picea Dietr. forest, moist soil, 1250 m, 13.07.1995, ÖE 1562. Euro-Sib. C. echinatus L. Tafll›ca village, Ar›p›nar, meadow, 1850 m, 17.07.1995, ÖE 1563. Briza media L. Tafll›ca village, Meydanlar, meadow, 1230 m, 10.07.1996, ÖE 1669. Melica uniflora Retz. Saçinge hill, open forest, 1600 m, 18.08.1997, ÖE 2006. Euro-Sib. Ö. EM‹NA⁄AO⁄LU, R. ANfi‹N, Nardus stricta L. Tuzlu hill, alpine pasture, 2900 m, 22.08.1996, ÖE 1670. Stipa arabica Trin. & Rupr. Cynodon dactylon (L.) Pers. var. dactylon Saçinge hill, slope, 1400 m, 22.08.1996, ÖE 1672. Zea mays L. Tafll›ca village, Meydanlar, cultivated place, 1200 m, 10.08.1997, ÖE 2007. Tafll›ca village, Ar›p›nar, meadow, 1850 m, 10.06.1996, ÖE 1671. 27

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The flora of Hatila Valley National Park and its close environs (Artvin)

The flora of Hatila Valley National Park and its close environs (Artvin)

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