Mycosphere
Towards incorporating anamorphic fungi in a natural classification – checklist
and notes for 2010
Hyde KD1,2*, McKenzie EHC3 and KoKo TW1
1
School of Mae Fah Luang University, 333 M. 1. T. Tasud Muang District, Chiang Rai 57100, Thailand.
Botany and Microbiology Department, College of Science, King Saud University, Riyadh, Saudi Arabia.
3
Manaaki Whenua Landcare Research, Private Bag 92170, Auckland, New Zealand.
2
Hyde KD, McKenzie EHC, KoKo TW. (2011). Towards incorporating anamorphic fungi in a
natural classification – checklist and notes for 2010. Mycosphere 2(1), 1–88.
A complilation of anamorphic names for both Ascomycota and Basidiomycota is provided which
compises 2873 genera. The genera are listed against a backbone of teleomorphic relationships
where known. The study reveals that 699 genera and 94 anamorph-like genera are linked to
teleomorphic genera names, 447 genera (one anamorph-like genus) are linked to teleomorph
families, orders or classes, while for more than 1728 (60.15%) genera no teleomorph link is known.
Key words –Asexual fungi – life cycle – sexual fungi – taxonomy
Article Information
Received 4 February 2011
Accepted 7 February 2011
Published online 10 March 2011
*Corresponding author: Hyde KD – e-mail – kdhyde3@gmail.com
Introduction
The purpose of the present paper is to
collate the knowledge of anamorphic fungi in
relationship to their teleomorphic states. It
attempts to classify the anamorphic genera in a
natural biological framework for the Ascomycota based on the Outline of Ascomycota 2009
(Lumbsch & Huhndorf 2010) and for the
Basidiomycota based on Kirk et al. (2008). The
present paper has used data from the Dictionary of the Fungi (Kirk et al. 2008), Index
Fungorum (http://www.indexfungorum.org/na
mes/Names.asp), Species Fungorum (http://
www.speciesfungorum.org/Names/Names.asp)
and Index of Fungi (Anon. 2009–2010) as well
as literature published in 2009 and 2010.
There is now debate in the systematics
community to place one name on a fungus
species as opposed to the present system
(especially in Ascomycota) of naming both
sexual and asexual morphs (Samuels et al.
2009). Also, as more and more strains of
anamorphic genera are sequenced (eg. Shenoy
et al. 2010), these genera will be classified in
families, orders and classes of Ascomycota and
Basidiomycota. This will result in more and
more anamorphic states being described and
illustrated without being given formal names
and therefore it is important to have a forum
where anamorphic data can be brought to the
attention of researchers. It is hoped that this
compilation will facilitate the move towards
one fungus one name as advocated by many
mycologists (Shenoy et al. 2007, 2010). By
placing all anamorphic genera against the
backbone classification of the sexual genera it
will be possible to establish 1) what are the
anamorphs of various species, genera, families
or orders, 2) which genera have no information,
3) which are the older or more commonly used
names for each genus, and 4) illustrate the
redundancy of the dual classification system.
This compilation also provides a single
place where anamorph connections can be
accessed and critically scrutinized so that a
stable and reliable system can be developed
1
based on published data. We do not claim that
this compilation is complete or definitive; however, it provides a forum for displaying data
and can be added to or changed as more data
becomes available or errors are discovered. In
future issues we will explore various connections and provide more data.
In the long term it will be desirable if
Myconet (http://www.fieldmuseum.org/myco
net) is merged with this document, however,
this should occur at some time in the future,
when more data is available.
Material and methods
The overall scheme for the classification
of the anamorphic fungi follows the Outline of
Ascomycota 2009 (Lumbsch & Huhndorf
2010), with additional data on anamorphic
Basidiomycota from Kirk et al. (2008). This
scheme is chosen as the schemes in the
Dictionary of the Fungi, Index Fungorum, and
MycoBank differ to a greater or lesser extent
and it was felt wise to follow what the authors
consider is an admirable attempt to keep
abreast of the current literarure as is routinely
carried out in the Outline of Ascomycota. This
compilation also annotates changes made in the
form of notes on anamorphic fungi (marked
with an asterisk after the entry, with notes in
the later “Notes” section) so that changes can
easily be followed; this is not always true of
the available databases.
Anamorphic genera or ‘-like’ genera are
listed in bold under the appropriate fungal
classification and the link to a teleomorph
genus given where found. Dubious genera are
excluded but can be added if they are later
found to be good genera. Fossil fungi are also
excluded as it would be impossible with
present techniques available to provide a natural classification for them. A question mark
before the entry means that its placement in
that family or order is not fully established and
generally follows Lumbsch & Huhndorf (2010).
A question mark before anamorph means it is
not clear that this is the anamorph of this genus,
although there are many cases without a question mark where the link has not equivocally
been proven. An asterisk (*) following the
names indicates an entry in the “Notes” section.
In many genus records in Index Fungorum the species is linked to more than one
2
anamorphic genus. Such records can be located
by searching the genus throughout the list for
repeat entries. The synonyms of genera are not
listed as they can be easily found in Index
Fungorum.
We encourage colleagues to inform us of
any missing entries, new data or misunderstandings, which will be updated in future
compilations. We would also invite experts of
specific groups to contribute to future compilations on their specific group(s) of interest and
encourage reviews or illustrated accounts of
families, orders or other groups relating to
teleomorph and anamorph links. Reviews of
anamorphic genera with sections or series, such
as Aspergillus, Penicillium and Verticillium,
the powdery mildews, many of the yeasts and
yeast-like genera and the anamorphic smuts
and rusts are particularly needed. Similarly the
lichenized ascomycete anamorphic states are
poorly documented.
As there is a move towards using one
name for both the sexual and asexual states of a
fungus we also encourage authors to submit
articles or opinions concerning the use of one
name. This might be in the form of a general
critic, arguments for or against using one name,
or an article that specifically addresses an
individual group or genus with arguments for
adopting the anamorph, teleomorph or both
names.
Index to Ascomycota
Phylum ASCOMYCOTA Caval-Sm.
Subphylum TAPHRINOMYCOTINA O.E.
Erikss. & Winka
Class Taphrinomycetes O.E. Erikss. & Winka
Taphrinales Gäum. & C.W. Dodge
Protomycetaceae Gray
Saitoella Goto, Sugiy., Hamam. & Komag.
1987
Taphrinaceae Gäum.
Lalaria R.T. Moore 1990, anamorphic
Taphrina Fr. 1815
Subphylum SACHAROMYCOTINA
Erikss. & Winka
O.E.
Class Saccharomycetes O.E. Erikss. & Winka
Saccharomycetales Kudrjanzev
Mycosphere
Dipodascaceae Engl. & E. Gilg
?Candida Berkhout 1923, anamorphic
Sporopachydermia Rodr. Mir. 1978
Galactomyces Redhead & Malloch 1977,
anamorphic Dipodascus Lagerh. 1892
Geotrichum Link 1809, anamorphic Dipodascus Lagerh. 1892
Geotrichum-like, anamorphic Basidioascus Matsush. 2003
Saprochaete Coker & Shanor 1939, anamorphic Magnusiomyces Zender 1926
Endomycetaceae J. Schröt.
Fusidium Link 1809, possibly anamorphic
Ascocephalophora K. Matsush. &
Matsush. 1995
Lipomycetaceae E.K. Novák & Zsolt
Myxozyma Van der Walt, Weijman & Arx
1981
Metschnikowiaceae T. Kamienski
Candida Berkhout 1923, anamorphic
Clavispora Rodr. Mir. 1979 and
Metschnikowia T. Kamienski 1899
Pichiaceae Zender
Brettanomyces N.H. Claussen ex Custers
1940, anamorphic Dekkera Van der
Walt 1964
Eeniella M.T. Sm., Bat. Vegte & Scheffers
1981, anamorphic Dekkera Van der
Walt
Enantiothamnus Pinoy 1911
Hyphopichia Arx & Van der Walt 1976
Trichomonascaceae Kurtzman & Robnett
Blastobotrys Klopotek 1967, anamorphic
Trichomonascus H.S. Jacks. 1948
Candida Berkhout 1923, ?anamorphic
Zygoascus M.T. Sm. 1986
Saccharomycetales, genera incertae sedis
Aciculoconidium D.S. King & S.C. Jong
1976
Botryozyma Shann & M.T. Sm. 1992,
anamorphic Ascobotryozyma J. Kerrigan, M.T. Sm. & J.D. Rogers 2001
Candida Berkhout 1923, anamorphic
Starmerella C.A. Rosa & Lachance
1998
Cicadomyces Šulc 1911
Macrorhabdus Tomasz., Logan, Snowden,
Kurtzman & Phalen 2003
Oosporidium Stautz 1931
Pseudomycoderma H. Will 1916
Schizoblastosporion Cif. 1930
Trigonopsis Schachner 1929
Saccharomycetes, genera incertae sedis
Selenotila Lagerh. 1892
Selenozyma Yarrow 1977
Subphylum PEZIZOMYCOTINA O.E. Erikss.
& Winka
Class Arthoniomycetes O.E. Erikss. & Winka
Arthoniales Henssen ex D. Hawksw. & O.E.
Erikss.
Arthoniaceae Reichenb. ex Reichenb.
Helicobolomyces Matzer 1995, anamorphic Arthonia Ach. 1806
Septocyta Petr. 1927, ?anamorphic Arthonia Ach. 1806
Subhysteropycnis Wedin & Hafellner 1998,
anamorphic Arthonia Ach. 1806
Saccharomycetaceae G. Winter
Candida Berkhout 1923, ?anamorphic
Ogataea Y. Yamada, K. Maeda &
Mikata 1994, Lodderomyces Van der
Walt 1966 and Spathaspora N.H.
Nguyen, S.O. Suh & M. Blackw. 2006
Roccellaceae Chevall.
Sporhaplus H.B.P. Upadhyay 1964, anamorphic Mazosia A. Massal 1854
Saccharomycodaceae Kudrjanzev
Candida Berkhout 1923, anamorphic
Lodderomyces Van der Walt 1966 and
Spathaspora N.H. Nguyen, S.O. Suh &
M. Blackw. 2006
?Kloeckera Janke 1923, anamorphic Hanseniaspora Zikes 1911
Class Dothideomycetes sensu O.E. Erikss. &
Winka
Subclass Dothideomycetidae P.M. Kirk, P.F.
Cannon, J.C. David & J.A. Stalpers ex C.L.
Schoch, Spatafora, Crous & Shoemaker
Capnodiales Woron.
Antennulariellaceae Woron.
3
Antennariella Bat. & Cif. 1963*, anamorphic Antennulariella Woron. 1915*
Capnodendron S. Hughes 1976, anamorphic Antennulariella Woron. 1915
Capnofrasera S. Hughes 2003
Heteroconium Petr. 1949, anamorphic
Antennulariella Woron. 1915
Hoornsmania Crous 2007
Rachicladosporium Crous, U. Braun &
C.F. Hill 2007*
Toxicocladosporium Crous & U. Braun
2007*
Verrucocladosporium K. Schub., Aptroot
& Crous 2007*
Capnodiaceae (Sacc.) Höhn. ex Theiss.
Acanthorus Bat. & Cavalc. 1967
Antennariella Bat. & Cif. 1963*
Apiosporium Kunze 1817
?Ciferrioxyphium Bat. & H. Maia 1963,
anamorphic Aithaloderma P. Syd.
Conidiocarpus Woron. 1927, ?anamorphic
Phragmocapnias Theiss. & Syd. 1918
and Scorias Fr. 1825
Conidioxyphium Bat. & Cif. 1963*
Fumagospora G. Arnaud 1911*, anamorphic Capnodium Mont. 1848
Fumiglobus D.R. Reynolds & G.S. Gilbert
2006
?Leptoxyphium Speg. 1918*, anamorphic
Aithaloderma P. Syd. 1913
Microxiphium (Harv. ex Berk. & Desm.)
Thüm. 1879 (as Microxyphium)*
?Mycogelidium W.Y. Zhuang 2007*
Phaeoxyphiella Bat. & Cif. 1963, anamorphic Capnodium Mont. 1848
Polychaetella Speg. 1918, anamorphic
Capnodium Mont. 1848
Scolecoxyphium Cif. & Bat. 1956,
anamorphic Capnodium Mont. 1848
and Scorias Fr. 1825
Tripospermum Speg. 1918, anamorphic
Phragmocapnias Theiss. & Syd. 1918
and Trichomerium Speg. 1918
Dissoconiaceae Crous & de Hoog*
Dissoconium de Hoog, Oorschot &
Hijwegen 1983*
Ramichloridium Stahel ex de Hoog 1977*
Coccodiniaceae Höhn. ex O.E. Erikss.
Bisbyopeltis Bat. & A.F. Vital 1957
Microxiphium (Harv. ex Berk. & Desm.)
Thüm. 1879*, anamorphic Dennisiella
Bat. & Cif. 1962
Davidiellaceae C.L. Schoch, Spatafora, Crous
& Shoemaker
Cladosporium Link 1816, anamorphic
Davidiella Crous & U. Braun 2003*
Dichocladosporium K. Schub., U. Braun
& Crous 2007
Graphiopsis Trail 1889*, anamorphic
Davidiella Crous & U. Braun 2003
4
Metacapnodiaceae S. Hughes & Corlett
Capnobotrys S. Hughes 1970, anamorphic
Metacapnodium Speg. 1918
Capnocybe S. Hughes 1966
Capnophialophora S. Hughes 1966, anamorphic Metacapnodium Speg. 1918
Capnosporium S. Hughes 1976, anamorphic Metacapnodium Speg. 1918
Hormiokrypsis Bat. & Nascim. 1957, anamorphic Metacapnodium Speg. 1918
Mycosphaerellaceae Lindau
Anguillosporella U. Braun 1995
Asperisporium Maubl. 1913
Asteromella-like, anamorphic Gillotia
Sacc. & Trotter 1913
Cercospora Fresen. 1863*
Cercosporella Sacc. 1880*
Clypeispora A.W. Ramaley 1991, ?anamorphic Mycosphaerella Johanson
1884
Coniothryium-like, anamorphic Bruneosphaerella Crous 2009*
Deightoniella S. Hughes 1952
Didymochora Höhn. 1918, anamorphic
Euryachora Fuckel 1870
Distocercospora N. Pons & B. Sutton
1988
Dothistroma Hulbary 1941*
Elletevera Deighton 1969
Floricola Kohlm. & Volkm.-Kohlm. 2000
Fusicladiella Höhn. 1919, anamorphic
Mycosphaerella Johanson 1884
Laocoön J.C. David 1997
Lecanosticta Syd. 1922*
Microcyclospora Jana Frank, Schroers &
Crous 2010*
Microcyclosporella Jana Frank, Schroers
& Crous 2010*
Mycosphere
Miuraea Hara 1948*, anamorphic Mycosphaerella Johanson 1884
Parastenella J.C. David 1991*
Passalora Fr. 1849*, anamorphic Mycosphaerella Johanson 1884
Penidiella Crous & U. Braun 2007*
Periconiella Sacc. 1885*, ?anamorphic
Allosoma Syd. 1926
Phacellium Bonord. 1860, anamorphic
Mycosphaerella Johanson 1884
Phaeophleospora Rangel 1916*
Phloeospora Wallr. 1833*, anamorphic
Mycosphaerella Johanson 1884
Placosphaeria-like, anamorphic Euryachora Fuckel 1870
Polythrincium Kunze 1817*, anamorphic
Cymadothea F.A. Wolf 1935
Pseudocercospora Speg. 1910*
Pseudocercosporella Deighton 1973*,
anamorphic Mycosphaerella Johanson
1884
Pseudocercosporidium Deighton 1973
Quasiphloeospora B. Sutton, Crous &
Shamoun 1996
Ramichloridium Stahel ex de Hoog 1977*
Ramularia Unger 1833*, anamorphic
Mycosphaerella Johanson 1884*
Ramularia-like, anamorphic Melanodothis
R.H. Arnold 1972
Ramulariopsis Speg. 1910, anamorphic
Mycosphaerella Johanson 1884
Ramulispora Miura 1920*
Rhabdospora (Durieu & Mont. ex Sacc.)
Sacc. 1884
Rhexocercosporidium U. Braun 1994
Semipseudocercospora J.M. Yen 1983
Septoria Sacc. 1884*
Sirosporium Bubák & Serebrian. 1912
Sonderhenia H.J. Swart & J. Walker
1988*
Stenella Syd. 1930, anamorphic Mycosphaerella Johanson 1884*
Stenellopsis B. Huguenin 1966
Thedgonia-like*
Trochophora R.T. Moore 1955*
Verrucisporota D.E. Shaw & Alcorn
1993*
Zasmidium Fr. 1849*
Capnodiales, genera incertae sedis
Capnocheirides J.L. Crane & S. Hughes
1982
Elasticomyces Zucconi & Selbmann 2008
Heptaster Cif., Bat. & Nascim. 1956
Hormonema-like, anamorphic Xenomeris
Syd. 1924
Houjia G.Y. Sun & Crous 2010*
Micropustulomyces R.W. Barreto 1995
Phaeotheca Sigler, Tsuneda & J.W.
Carmich. 1981
Pseudovirgaria H.D. Shin, U. Braun,
Arzanlou & Crous 2007
Racodium Pers. 1794*
Recurvomyces Selbmann & de Hoog 2008
Scolecostigmina U. Braun 1999
Sporidesmajora Batzer & Crous 2010*
Dothideales Lindau
Dothideaceae Chevall.
Lecanosticta Syd. 1922, anamorphic Scirrhia Nitschke ex Fuckel 1870
Podoplaconema Petr. 1921, anamorphic
Omphalospora Theiss. & Syd. 1915
Dothioraceae Theiss. & H. Syd.
Aureobasidium Viala & G. Boyer 1891*,
anamorphic Columnosphaeria Munk
1953 and Saccothecium Fr. 1836
Dothichiza Lib. ex Roum. 1880, anamorphic Dothiora Fr. 1849 and Sydowia Bres. 1895
Hormonema Lagerb. & Melin 1927,
anamorphic Dothiora Fr. 1849 and
Sydowia Bres. 1895*
Japonia Höhn. 1909, anamorphic Yoshinagaia Henn. 1904
Kabatina R. Schneid. & Arx 1966
?Rhizosphaera L. Mangin & Har. 1907,
anamorphic Phaeocryptopus Naumov
1915 and Dimerina Theiss. 1912
Sclerophoma Höhn. 1909, anamorphic
Sydowia Bres. 1895
Teratosphaeriaceae Crous & U. Braun
Acidomyces B.J. Baker, M.A. Lutz, S.C.
Dawson, P.L. Bond & Banfield ex
Selbmann, de Hoog & De Leo 2008
Batcheloromyces Marasas, P.S. van Wyk
& Knox-Dav. 1975*
Baudoinia J.A. Scott & Unter. 2007*
Catenulostroma Crous & U. Braun 2007*
Capnobotryella Sugiy. 1987*
Cibiessia Crous 2007*
5
Colletogloeopsis Crous & M.J. Wingf.
1997*
Cystocoleus Thwaites 1849*
Davisoniella H.J. Swart 1988*
Devriesia Seifert & N.L. Nick. 2004*
Friedmanniomyces Onofri 1999*
Hortaea Nishim. & Miyaji 1984*
Kirramyces J. Walker, B. Sutton & Pascoe
1992, anamorphic Teratosphaeria Syd.
& P. Syd. 1912*
Nothostrasseria Nag Raj 1983
Passalora Fr. 1849*
Penidiella Crous & U. Braun 2007*
Phacellium Bonord. 1860*
Phaeothecoidea Crous 2007*
Pseudocercosporella Deighton 1973*
Pseudoramichloridium Cheewangkoon &
Crous 2010*
Pseudotaeniolina J.L. Crane & Schokn.
1986*
Ramichloridium Stahel ex de Hoog 1977*
Readeriella Syd. & P. Syd. 1908*
Sporidesmium Link 1809*
Staninwardia B. Sutton 1971*
Stenella Syd. 1930*
Trimmatostroma-like, anamorphic Teratosphaeria Syd. & P. Syd. 1912
Tripospermum Speg. 1918*
Myriangiales Starbäck
Elsinoaceae Höhn. ex Sacc. & Trotter
Endosporium Tsuneda 2008
Sphaceloma de Bary 1874 anamorphic
Elsinoë Racib. 1900
Xenodiella Syd. 1935, anamorphic Xenodium Syd. 1935
Myriangiales, genera incertae sedis
Phaeosclera Sigler, Tsuneda & J.W.
Carmich. 1981*
Sarcinomyces Lindner 1898*
Subclass Pleosporomycetidae C.L. Schoch,
Spatafora, Crous & Shoemaker
Pleosporales Luttrell ex M.E. Barr
Amniculicolaceae Yin. Zhang, C.L. Schoch, J.
Fourn., Crous & K.D. Hyde
Anguillospora Ingold 1942*
Cucurbitariaceae G. Winter
Camarosporium Schulzer 1870, anamorphic Cucurbitaria Gray 1821
6
Coniothyrium-like, anamorphic Curreya
Sacc. 1883
Diplodia-like, anamorphic Cucurbitaria
Gray 1821
?Phaeoseptoria Speg. 1908, anamorphic
Rhytidiella Zalasky 1968
Phialophorophoma Linder 1944*
Phoma Sacc. 1880*
Pleurophoma Höhn. 1914*
Pleurostromella Petr. 1922, anamorphic
Cucurbitaria Gray 1821 and Gibberidea Fuckel 1870
Pyrenochaeta De Not. 1849, anamorphic
Cucurbitaria Gray 1821*
Pyrenochaetopsis Gruyter, Aveskamp, &
Verkley, 2010*
?Megaloseptoria Naumov 1925, anamorphic Cucurbitaria Gray 1821
Syntholus A.W. Ramaley & M.E. Barr
1997, anamorphic Syncarpella Theiss.
& Syd. 1915
Diademaceae Shoemaker & C.E. Babc.
Alternaria-like anamorphic Comoclathris
Clem. 1909
Didymellaceae Gruyter, Aveskamp & Verkley
Ascochyta Lib. 1830, anamorphic Didymella Sacc. ex D. Sacc. 1880*
Ampelomyces Ces. ex Schltdl. 1852*
Boeremia Aveskamp, Gruyter & Verkley
2010*
Chaetasbolisia Speg. 1918*
Dactuliochaeta G.L. Hartm. & J.B.
Sinclair 1988, anamorphic Didymella
Sacc. 1880
Epicoccum Link 1815*
Microsphaeropsis Höhn. 1917*
Peyronellaea Gold. ex Togliani 1952*
Phoma Sacc. 1880, anamorphic Didymella
Sacc. ex D. Sacc. 1880*
Piggotia Berk. & Broome 1851, possibly
anamorphic Platychora Petr. 1925
Pithoascus Arx 1973, ?anamorphic
Leptosphaerulina McAlpine 1902
Pithomyces Berk. & Broome 1875, anamorphic Leptosphaerulina McAlpine
1902
Stagonosporopsis Died. 1912*, anamorphic Didymella Sacc. ex D. Sacc. 1880
Mycosphere
Didymosphaeriaceae Munk
Cytoplea Bizz. & Sacc. 1885, anamorphic
Roussoëlla Sacc. 1888
Dothidotthiaceae Crous & A.J.L. Phillips
Dothiorella Sacc. 1880, ?anamorphic
Dothidotthia Höhn. 1918*
Thyrostroma Höhn. 1911, anamorphic
Dothidotthia Höhn. 1918
Hypsostromataceae Huhndorf
Pleurophomopsis–like, anamorphic Hypsostroma Huhndorf
Lentitheciaceae Yin. Zhang, C.L. Schoch, J.
Fourn., Crous & K.D. Hyde
Stagonospora (Sacc.) Sacc. 1884*
Leptosphaeriaceae M.E. Barr
Coniothyrium Corda 1840, anamorphic
Leptosphaeria Ces. & De Not. 1863*
Coniothyrium-like, anamorphic Neophaeosphaeria M.P.S. Câmara, M.E. Palm &
A.W. Ramaley 2003
Phoma Sacc. 1880, anamorphic Leptosphaeria Ces. & De Not. 1863*
Plenodomus Preuss 1851
Pyrenochaeta De Not. 1849, possibly
anamorphic Leptosphaeria Ces. & De
Not. 1863*
Lindgomycetaceae K. Hiray., Kaz. Tanaka &
Shearer
Taeniolella S. Hughes 1958*
Lophiostomataceae Sacc.
Ascochyta Lib. 1830, anamorphic Trichometasphaeria Munk 1953
Pleuorphomopsis-like, anamorphic Lophiostoma Ces. & De Not. 1863
Massariaceae Nitschke
?Aplosporella-like, anamorphic Dubitatio
Speg. 1882
Myxocyclus Riess 1852, anamorphic
Massaria De Not. 1844
Torula Pers. 1794*
Massarinaceae Munk
Aquaticheirospora Kodsueb & W.H. Ho
2007*
Ceratophoma Höhn. 1917, anamorphic
Massarina Sacc. 1883
Cheirosporium L. Cai & K.D. Hyde 2008*
Helminthosporium Link 1809*
Neottiosporina Subram. 1961*
Pseudodictyosporium Matsush. 1971
Melanommataceae G. Winter
Aposphaeria Sacc. 1880*
Beverwykella Tubaki 1975*
Nigrolentilocus R.F. Castañeda & Heredia
2001, anamorphic Melanomma Nitschke ex Fuckel 1870
Monodictys S. Hughes 1958, ?anamorphic
Ohleria Fuckel 1868
Monotosporella S. Hughes 1958*
Pyrenochaeta De Not. 1849, anamorphic
Byssosphaeria Cooke 1879
Sporidesmiella P.M. Kirk 1982
Montagnulaceae M.E. Barr
Aschersonia Mont. 1848, anamorphic
Montagnula Berl. 1896
Microdiplodia Allesch. 1901, possibly
anamorphic Karstenula Speg. 1879
Microsphaeropsis Höhn. 1917, anamorphic Paraphaeosphaeria O.E. Erikss.
1967*
Paraconiothyrium Verkley 2004*, possibly anamorphic Paraphaeosphaeria
O.E. Erikss. 1967
Phaeosphaeriaceae M.E. Barr
Amarenographium O.E. Erikss. 1982,
anamorphic Amarenomyces O.E. Erikss.
1981 (recently synonymised with Phaeosphaeria I. Miyake 1909)
Ampelomyces Ces. ex Schltdl. 1852*
Chaetosphaeronema Moesz 1915*
Coniothyrium Corda 1840, anamorphic
Ophiobolus Riess 1854
Hendersonia-like, anamorphic Austropleospora R.G. Shivas & L. Morin 2010 and
Phaeosphaeria I. Miyake 1909*
Neosetophoma Gruyter, Aveskamp &
Verkley 2010*
?Parahendersonia A.W. Ramaley 1995,
anamorphic Chaetoplea (Sacc.) Clem.
1931
Paraphoma Morgan-Jones & J.F. White
1983*
7
Phaeoseptoria Speg. 1908, anamorphic
Phaeosphaeria I. Miyake 1909
Phaeostagonospora A.W. Ramaley 1997,
anamorphic Phaeosphaeriopsis M.P.S.
Câmara, M.E. Palm & A.W. Ramaley
2003
Phoma Sacc. 1880, anamorphic Ophiobolus Riess 1854
Rhabdospora (Durieu & Mont. ex Sacc.)
Sacc. 1884, anamorphic Ophiobolus
Riess 1854
Scolecosporiella Petr. 1921, anamorphic
Ophiosphaerella Speg. and Phaeosphaeria I. Miyake 1909
Setophoma Gruyter, Aveskamp & Verkley
2010*
?Sphaerellopsis Cooke 1883, anamorphic
Eudarluca Speg. 1908
Stagonospora (Sacc.) Sacc. 1884, anamorphic Phaeosphaeria I. Miyake 1909*
Tiarospora Sacc. & Marchal 1885
Pleomassariaceae M.E. Barr
Ceuthodiplospora Died. 1912, anamorphic
Splanchnonema Corda
?Prosthemium Kunze 1817, anamorphic
Pleomassaria Speg. 1880*
Myxocyclus Riess 1852, anamorphic
Splanchnonema Corda 1829
Scolicosporium Lib. ex Roum. 1880,
anamorphic Asteromassaria Höhn.
1917
Shearia Petr. 1924, anamorphic Pleomassaria Speg. 1880
Stegonsporium Corda 1827, anamorphic
Splanchnonema Corda 1829
Pleosporaceae Nitschke
Alternaria Nees 1816, anamorphic Lewia
M.E. Barr & E.G. Simmons 1986
Alternariaster E.G. Simmons 2007
Brachycladium Corda 1838, anamorphic
Crivellia Shoemaker & Inderbitzin
2006
Bipolaris Shoemaker 1959, anamorphic
Cochliobolus Drechsler 1934
Chalastospora E.G. Simmons 2007*
Curvularia Boedijn 1933, anamorphic
Cochliobolus Drechsler 1934
Dactuliophora C.L. Leakey 1964
Dendryphiella Bubák & Ranoj. 1914*
Dendryphion Wallr. 1833
8
Dictyosporium Corda 1836*
Digitodesmium P.M. Kirk 1981
Drechslera S. Ito 1930, anamorphic Pyrenophora Fr. 1849
Edenia M.C. González, Anaya, Glenn,
Saucedo & Hanlin 2007*
Embellisia E.G. Simmons 1971, anamorphic Lewia M.E. Barr & E.G. Simmons
Exserohilum K.J. Leonard & Suggs 1974,
anamorphic Setosphaeria K.J. Leonard
& Suggs 1974
Gibbago E.G. Simmons 1986
Lemonniera De Wild. 1894*
Marielliottia Shoemaker 1999, anamorphic
Pyrenophora Fr. 1849
Nimbya E.G. Simmons 1989, anamorphic
Macrospora Fuckel 1870
Prathoda Subram. 1956
Stemphylium Wallr. 1833, anamorphic
Pleospora Rabenh. ex Ces. & De Not.
1863*
Teretispora E.G. Simmons 2007
Ulocladium Preuss 1851*
Sporormiaceae Munk
?Amorosia Mantle & D. Hawksw. 2006
Phoma Sacc. 1880*
Phoma-like, anamorphic Westerdykella
Stolk 1955*
Teichosporaceae M.E. Barr
Coniothyrium-like, anmorphic Immothia
M.E. Barr
Tetraplosphaeriaceae Kaz. Tanaka & K. Hiray
Pseudotetraploa Kaz. Tanaka & K.
Hirayama 2009*
Quadricrura Kaz. Tanaka, K. Hirayama &
Sat. Hatak. 2009*
Tetraploa Berk. & Broome 1850,
anamorphic Tetraplosphaeria Kaz.
Tanaka & K. Hirayama 2009*
Tetraploa-like, anamorphic Triplosphaeria
Kaz. Tanaka & K. Hirayama 2009 and
Polyplosphaeria Kaz. Tanaka & K.
Hirayama 2009*
Venturiaceae E. Müll. & Arx ex M.E. Barr
Anungitea B. Sutton 1973
Anungitopsis R.F. Castañeda & W.B.
Kendr. 1990
Mycosphere
Cylindrosympodium W.B. Kendr. & R.F.
Castañeda 1990
Dictyodochium Sivan. 1984, anamorphic
Gibbera Fr. 1825
Fusicladium Bonord. 1851, anamorphic
Acantharia Theiss. & Syd. 1918, Dibotryon Thesis. & Syd. 1915 and ?Venturia Sacc. 1882*
Fusicladium-like, anamorphic Antennularia Rchb. 1828
Piggotia Berk. & Broome 1851
Sclerophoma Höhn. 1909, ?anamorphic
Xenomeris Syd. 1924
Spilodochium Syd. 1927, anamorphic
Pseudoparodiella F. Stevens 1927
Stigmina-like, anamorphic Gibbera Fr.
1825 and Phragmogibbera Samuels &
Rogerson 1990
Ulocladium-like, anamorphic Lasiobotrys
Kunze 1823
Veronaeopsis Arzanlou & Crous 2007
Zeloasperisporium R.F. Castañeda 1996
Pleosporales, genera incertae sedis
Berkleasmium Zobel 1854
Briansuttonia R.F. Castañeda, Minter &
Saikawa 2004
Cheiromoniliophora Tzean & J.L. Chen
1990
Clavariopsis De Wild. 1895
Diplococcium Grove 1885*
Ellisembia Subram. 1992
Fusculina Crous & Summerell 2006
Mycocentrospora Deighton 1972
Ochrocladosporium Crous & U. Braun
2007
Phoma Sacc. 1880*
Periconia Tode 1791
Pleurophomopsis Petr. 1924, anamorphic
Astrosphaeriella Syd. & P. Syd. 1913
Polyschema H.P. Upadhyay 1966*
Pseudochaetosphaeronema Punith. 1979
Repetophragma Subram. 1992*
Sclerostagonospora Höhn. 1917*
Sirodesmium De Not. 1849*
Spadicoides S. Hughes 1958*
Sporidesmium Link 1809*
Versicolorisporium Sat. Hatak., Kaz.
Tanaka & Y. Harada 2008
Zalerion-like, anamorphic Trematosphaeria Fuckel 1870
Pleosporomycetidae, genera incertae sedis
Acrogenospora M.B. Ellis 1971, anamorphic Farlowiella Sacc. 1891
Dothideomycetes, orders incertae sedis
Acrospermales Minter, Peredo & A.T. Watson
Acrospermaceae Fuckel
Gonatophragmium Deighton 1969, anamorphic Acrospermum Tode 1790
Botryosphaeriales C.L. Schoch, Crous &
Shoemaker
Botryosphaeriaceae Theiss. & H. Syd.*
Aplosporella Speg. 1880*
Bahusutrabeeja Subram. & Bhat 1977*
Barriopsis A.J.L. Phillips, A. Alves &
Crous 2008*
Dichomera Cooke 1878
Diplodia Fr. 1834*
Diplodia-like, anamorphic Neodeightonia
C. Booth 1970 and Saccharata Denman
& Crous 2004
Dothichiza-like, anamorphic Leptoguignardia E. Müll. 1955
Dothiorella Sacc. 1880, anamorphic Spencermartinsia A.J.L. Phillips, A. Alves
& Crous 2008
Fusicoccum Corda 1829, anamorphic
Botryosphaeria Ces. & De Not. 1863
Fusicoccum-like, anamorphic Melanops
Nitschke ex Fuckel 1870* and Saccharata Denman & Crous 2004
Leptodothiorella Höhn. 1918, anamorphic
Guignardia Viala & Ravaz 1892
Lasiodiplodia Ellis & Everh. 1896*
Macrophomina Petr. 1923*
Microdiplodia Allesch. 1901
Neofusicoccum Crous, Slippers & A.J.L.
Phillips 2006, with Botryosphaeria-like
teleomorphs*
Neoscytalidium Crous & Slippers 2006
Phaeobotryon Theiss. & Syd. 1915*
Phyllosticta Pers. 1818, anamorphic Guignardia Viala & Ravaz 1892
Pseudofusicoccum Mohali, Slippers & M.J.
Wingf. 2006
Sphaeropsis Sacc. 1880
Botryosphaeriales, genera incertae sedis
Camarosporium Schulzer 1870
Dichomera Cooke 1878
Hendersonula Speg. 1880
9
Hysteriales Lindau
Hysteriaceae Chevall.
Aposphaeria-like, anamorphic Gloniopsis
De Not. 1847* and Hysterobrevium E.
Boehm & C.L. Schoch 2010*
Plenodomus Preuss 1851, anamorphic
Psiloglonium Höhn. 1918*
Septonema Corda 1837, anamorphic Oedohysterium E. Boehm & C.L. Schoch
2009*
Sporidesmium Link 1809, anamorphic
Psiloglonium Höhn. 1918*
Jahnulales Pang, Abdel-Wahab, El-Sharouney,
E.B.G. Jones & Sivichai
Aliquandostipitaceae Inderbitzin
Brachiosphaera Nawawi 1976*
Xylomyces Goos, R.D. Brooks & Lamore
1977*
Mytilinidiales Boehm, C.L. Schoch &
Spatafora
Gloniaceae (Corda) Boehm, C.L. Schoch &
Spatafora
Cleistonium Speer 1986, anamorphic
Glonium Mühl. 1813
Mytilinidiaceae Kirschst.
Camaroglobulus Speer 1986, anamorphic
Mytilinidion Duby 1862
Chalara-like, anamorphic Quasiconcha
M.E. Barr & M. Blackw. 1981
Papulospora-like, anamorphic Lophium Fr.
1818
Septonema-like anamorphic Mytilinidion
Duby 1862
Patellariales D. Hawksw. & O.E. Erikss.
Patellariaceae Corda
Aposphaeria-like, anamorphic Rhytidhysteron Speg. 1881
Diplodia-like, anamorphic Rhytidhysteron
Speg. 1881
Dothideomycetes, families incertae sedis
Ascoporiaceae Kutorga & D. Hawksw.
Plectophomella-like, anamorphic Pseudosolidum Lloyd 1923
Asterinaceae Hansf.
?Acarella Syd. 1927, anamorphic Morenoina Theiss. 1913
10
Asterostomella Speg. 1886, anamorphic
Asterina Lév.
Asterostromina Bat. & A.F. Vital 1957,
anamorphic Asterodothis Theiss.
Asterostomula Theiss. 1916, anamorphic
Echidnodes Theiss. & Syd. 1918
?Clasterosporium
Schwein.
1832,
anamorphic Eupelte Syd. 1924
Clasterosporium-like, anamorphic Asterina Lév. 1845
Elachopeltis Syd. 1927, ?anamorphic
Aphanopeltis Syd. 1927
Leprieurina G. Arnaud 1918, anamorphic
Prillieuxina G. Arnaud 1918
Peltasterella Bat. & H. Maia 1959, ?anamorphic Yamamotoa Bat. 1960
Pirozynskia Subram. 1972, anamorphic
Eupelte Syd. 1924
Mahanteshamomyces Hosag. & C.K. Biju
2004
Septoidium-like, anamorphic Eupelte Syd.
1924
Septothyrella Höhn. 1911, anamorphic
Uleothyrium Petr. 1929
Sirothyriella Höhn. 1910, anamorphic
Morenoina Thesis. 1913
?Thyrinula Petr. & Syd. 1924, anamorphic
Aulographina Arx & E. Müll.
Shivomyces Hosag. 2004
Triposporium Corda 1837, anamorphic
Batistinula Arx 1960
Corynesporascaceae Sivan.
Corynespora Güssow 1906, anamorphic
Corynesporasca Sivan.
Dacampiaceae Körb.
Cyclothyrium Petr. 1923, anamorphic
Polycoccum Saut. ex Körb. 1865
Lecythophora-like, anamorphic Munkovalsaria Aptroot 1995
Englerulaceae Henn.
Capnodiastrum Speg. 1886, anamorphic
Englerula Henn. 1904 and Rhytidenglerula Höhn. 1918
Questieriella G. Arnaud ex S. Hughes
1983, anamorphic Schiffnerula Höhn.
1909
Mitteriella Syd. 1933, anamorphic Schiffnerula Höhn. 1909
Mycosphere
Sarcinella Sacc. 1880, anamorphic Schiffnerula Höhn. 1909
Eremomcetaceae Malloch & Cain
Arthrographis G. Cochet ex Sigler & J.W.
Carmich. 1976, anamorphic Eremomyces Malloch & Cain
Trichosporiella-like, anamorphic Rhexothecium Samson & Mouch. 1975
Euantennariaceae Hughes & Corlett
Antennatula Fr. ex F. Strauss 1850,
anamorphic Euantennaria Speg. and
Strigopodia Bat. 1957
Capnokyma S. Hughes 1975
Capnophialophora S. Hughes 1966,
anamorphic Strigopodia Bat. 1957
Hormisciomyces Bat. & Nascim. 1957,
anamorphic Euantennaria Speg.
Hyphosoma Syd. 1924, anamorphic Strigopodia Bat. 1957
Plokamidomyces Bat., C.A.A. Costa & Cif.
1957, anamorphic Trichopeltheca Bat.,
C.A.A. Costa & Cif. 1958
Racodium Fr. 1829, anamorphic Strigopodia Bat. 1957
Trichothallus F. Stevens 1925, anamorphic
Trichopeltheca Bat., C.A.A. Costa &
Cif. 1958
Fenestellaceae M.E. Barr
Pleurostromella Petr. 1922, anamorphic
Fenestella Tul. & C. Tul. 1863
Leptopeltidaceae Höhn. ex Trotter
Idriella-like, anamorphic Dothiopeltis E.
Mull. 1957
Leptothyrium-like, anamorphic Leptopeltis
Höhn. 1917
Meliolinaceae S. Hughes
Briania D.R. Reynolds 1989, anamorphic
Meliolina Syd. & P. Syd. 1914
Micropeltidaceae Clem. & Shear
Cyclopeltella Petr. 1953, anamorphic
Cyclopeltis Petr. 1953
Sirothyriella Höhn. 1910, anamorphic
Stomiopeltis Theiss. 1914
Sporidesmium-like anamorphic Stomiopeltis Theiss. 1914 (as Akaropeltella
M.L. Farr, 1972)
Microthyriaceae Sacc.
Asterostomula Theiss. 1916, anamorphic
Asterinella Theiss. 1912
Asteromella-like, anamorphic Asterinella
Theiss. 1912
Eriothyrium Speg. 1888 (possible link
to Asterinema Bat. & Gayão 1953)
Hansfordiella S. Hughes 1951, anamorphic Trichothyrium Speg. 1889
Isthmospora F. Stevens 1918, anamorphic
Trichothyrium Speg. 1889
Leptothyrium Kunze 1823, anamorphic
Microthyrium Desm. 1841
Xenogliocladiopsis Crous & W.B. Kendr.
1994, anamorphic Arnaudiella Petr.
1927
Parmulariaceae E. Müll. & Arx ex M.E. Barr
?Melanoplaca Syd. & P. Syd. 1917,
anamorphic Dothidasteroma Höhn.
1909
Placomelan Cif. 1962, anamorphic
Dothidasteroma Höhn. 1909
Parodiellaceae Theiss. & H. Syd. ex M.E. Barr
?Ascochytopsis Henn. 1905, anamorphic
Parodiella Speg. 1880
Parodiopsidaceae Toro
Chuppia-like, anamorphic Neoparodia
Petr. & Cif. 1932
Cicinnobella Henn. 1904, anamorphic
Perisporiopsis Henn. 1904 (as Phaeodimeriella Speg. 1908) and Dimerium
(Sacc. & P. Syd.) Sacc. & D. Sacc.
1905
Exosporinella Bender 1932, possibly
anamorphic Parodiellina Henn. ex G.
Arnaud 1918
Ophiotrichum Kunze 1849 ?anamorphic
Leptomeliola Höhn. 1919
Sarcinella-like, anamorphic Neoparodia
Petr. & Cif. 1932
Septoidium G. Arnaud 1921, anamorphic
Alina Racib. 1909, possibly Perisporiopsis Henn. 1904* and Pilgeriella
Henn. 1900
Tretospora M.B. Ellis 1976, anamorphic
Balladyna Racib. and Balladynopsis
Theiss. & Syd.
11
Planistromellaceae M.E. Barr
Aposphaeria-like, anamorphic Microcyclus
Sacc., Syd. & P. Syd. 1904
Fusicladium Bonord. 1851, anamorphic
Microcyclus Sacc., Syd. & P. Syd. 1904
Hyphospora A.W. Ramaley 1996, anamorphic Comminutispora A.W. Ramaley 1996
Kellermania Ellis & Everh. 1885,
anamorphic Planistroma A.W. Ramaley
1991
Lecanosticta Syd. 1922, anamorphic
Eruptio M.E. Barr 1996
Pazschkeella Syd. & P. Syd. 1901,
anamorphic Microcyclus Sacc., Syd. &
P. Syd. 1904
Piptarthron Mont. ex Höhn. 1918,
anamorphic Planistroma A.W. Ramaley
1991
Polystomellaceae Theiss. & H. Syd.
Lasmenia Speg. 1886, anamorphic Munkiella Speg. 1885
Asteromella Pass. & Thüm. 1880 (as
Stictochorella Höhn. 1917), anamorphic
Dothidella Speg. 1880
Pseudoperisporiaceae Toro
Chaetosticta Petr. & Syd. 1925, anamorphic Lasiostemma Theiss., Syd. & P.
Syd. 1917 and ?Nematostoma Syd. & P.
Syd. 1914
Schizothyriaceae Höhn. ex Trotter, Sacc., D.
Sacc. & Traverso
Plenotrichaius Bat. & Valle 1961, anamorphic Mycerema Bat., J.L. Bezerra &
Cavalc 1963
Zygophiala E.W. Mason 1945, anamorphic
Schizothyrium Desm. 1849*
Tubeufiaceae M.E. Barr
Annellospermosporella P.R. Johnst. 1999,
anamorphic Malacaria Syd. 1930
Aquaphila Goh, K.D. Hyde & W.H. Ho
1998, anamorphic Tubeufia Penz. &
Sacc. 1898
Araneomyces Höhn. 1909, anamorphic
Paranectriella (P. Henn. & Sacc.) Höhn.
Guelichia Speg. 1886, anamorphic Puttemansia Henn. 1902
12
Helicoma Corda 1837, anamorphic
Tubeufia Penz. & Sacc. 1898
Helicoön Morgan 1892
Helicomyces Link 1809, anamorphic
Acanthostigma De Not. (1863)
Helicosporium Nees 1816, anamorphic
Acanthostigma De Not. (1863), Thaxteriellopsis Sivan., Panwar & S.J. Kaur
1977, Tubeufia Penz. & Sacc. 1898
Kamalomyces R.K. Verma, N. Sharma &
Soni 2008
Mirandina G. Arnaud ex Matsush. 1975,
anamorphic Taphrophila Scheuer 1988
Monodictys-like, anamorphic Tubeufia
Penz. & Sacc. 1898
Pendulispora M.B. Ellis 1961, anamorphic
Tubeufia Penz. & Sacc. 1898
Peziotrichum (Sacc.) Lindau 1900, anamorphic Podonectria Petch 1921
Tetracrium Henn. 1902, anamorphic Puttemansia Henn. 1902
Titaea Sacc. 1876, anamorphic Paranectriella (Henn. ex Sacc. & D. Sacc.)
Höhn. 1910
Xenosporium Penz. & Sacc. 1901, anamorphic Acanthostigmella Höhn. 1905 and
Chaetosphaerulina I. Hino 1938 (as
Thaxteriellopsis Sivan., Panwar & S.J.
Kaur 1977).
Vizellaceae H.J. Swart
Chrysogloeum Petr. 1959, anamorphic
Blasdalea Sacc. & P. Syd. 1902
Manginula G. Arnaud 1918, anamorphic
Vizella Sacc. 1883
Dothideomycetes, genera incertae sedis
Acrocalymma Alcorn & J.A.G. Irwin
(1987), anamorphic Massarina, however Massarina has been shown to be
polyphyletic and its placement is therefore presently unclear.
Ascochyta-like, anamorphic Gilletiella
Sacc. & P. Syd. 1899
Asteromella Pass. & Thüm. 1880
Bactrodesmium Cooke 1883, anamorphic
Stuartella Fabre 1879
Bahusakala Subram. 1958
Blastostroma C.Z. Wei, Y. Harada &
Katum. 1998, anamorphic Mycodidymella C.Z. Wei, Y. Harada & Katum.
1998
Mycosphere
Brachyconidiella R.F. Castañeda & W.B.
Kendr. 1990
Camarosporula Petr. 1954, anamorphic
Anthracostroma Petr. 1954
Cenococcum Moug. & Fr. 1829
Chaetasbolisia Speg. 1918, anamorphic
Pseudomorfea Punith. 1981*
Cladoriella Crous 2006
Coniosporium Link 1809*
Coronospora M.B. Ellis 1971, anamorphic
Ascocoronospora Matsush. 2003
Cryomyces Selbmann, de Hoog, Mazzaglia,
Friedmann & Onofri 2005*
?Cyclothyrium Petr. 1923, anamorphic
Thyridaria Sacc. 1875
Dendryphiopsis S. Hughes 1953, anamorphic Kirschsteiniothelia D. Hawksw.
1985
Diederichia D. Hawksw. 2003
Dilophospora Desm. 1840, anamorphic
Lidophia J. Walker & B. Sutton 1974
Diplodia-like, anamorphic Otthia Nitschke
ex Fuckel 1870
Disculina Höhn. 1916
Hansfordiellopsis Deighton 1960, anamorphic Koordersiella Höhn. 1909
Helicodendron Peyronel 1918, anamorphic
Tyrannosorus Unter. & Malloch 1995
Hiospira R.T. Moore 1962, anamorphic
Brooksia Hansf. 1956
Iledon Samuels & J.D. Rogers 1986,
anamorphic Botryohypoxylon Samuels
& J.D. Rogers 1986
Megaloseptoria Naumov 1925
Monodictys S. Hughes 1958
Peltaster Syd. & P. Syd. 1917
Phaeothecoidiella Batzer & Crous 2010*
Phanerococculus Cif. 1954, anamorphic
Koordersiella Höhn. 1909
Phialophora-like, anamorphic Tirisporella
E.B.G. Jones, K.D. Hyde & Alias 1996
Phoma-like, anamorphic Tremateia
Kohlm., Volkm.-Kohlm. & O.E. Erikss.
1995
Placosphaeria (De Not.) Sacc. 1880
Plectopycnis Bat. & A.F. Vital 1959
Pleurostromella Petr. 1922, anamorphic
Gibberidea Fuckel 1870
Pseudorobillarda M. Morelet 1968*
Pycnopleiospora C.Z. Wei, Y. Harada &
Katum. 1997, anamorphic Pseudodidy-
mella C.Z. Wei, Y. Harada & Katum.
1997
Ramalia Bat. 1957
Rhizopycnis D.F. Farr 1998
Scleroconidioma Tsuneda, Currah & Thormann 2000
Septoriella Oudem. 1889
Sympodiella-like, anamorphic Sympoventuria Crous & Seifert 2007
Troposporella P. Karst. 1892*
Xenostigmina Crous 1998*
Class Eurotiomycetes Tehler ex O.E. Eriksson
& K. Winka
Subclass Chaetothyriomycetidae Doweld
Chaetothyriales M.E. Barr
Chaetothyriaceae Hansf. ex M.E. Barr
Cyphellophora G.A. de Vries 1962*
Merismella Syd. 1927, anamorphic
Chaetothyrium Speg.
Stanhughesia Constant. 1989, anamorphic
Ceramothyrium Bat. & H. Maia 1956
Vonarxia Bat. 1960*
Herpotrichiellaceae Munk
Brycekendrickomyces Crous & M.J. Wingf.
2009*
Cladophialophora Borelli 1980, anamorphic Capronia Sacc. 1883*
Exophiala J.W. Carmich. 1966, anamorphic Capronia Sacc. 1883
Fonsecaea Negroni 1936, anamorphic
Capronia Sacc. 1883
Metulocladosporiella Crous, Schroers, J.Z.
Groenew., U. Braun & K. Schub. 2006
Nadsoniella Issatsch. 1914
Phaeococcomyces de Hoog 1979
Phialophora Medlar 1915
Racodium Pers. 1794, anamorphic Capronia Sacc. 1883
Rhinocladiella Nannf. 1934, anamorphic
Capronia Sacc. 1883
Sorocybe Fr. 1849
Thysanorea Arzanlou, W. Gams & Crous
2007
Chaetothyriales M.E. Barr, genera incertae
sedis
Coniosporium Link 1809, the type species
is considered as Sirodesmium De Not.
1849*
13
Phaeoannellomyces McGinnis & Schell
1985
Staninwardia B. Sutton 1971*
Strelitziana Arzanlou & Crous 2006*
Pyrenulales Fink ex D. Hawksw. & O.E.
Erikss.
Pyrenulales, genera incertae sedis
Lyromma Bat. & H. Maia 1965
Chaetothyriomycetidae, families incertae sedis
Rhynchostomataceae Winka & O. E. Erikss.
Arthropycnis Constant. 1992, anamorphic
Rhynchostoma P. Karst.
Strigulaceae Zahlbr.
Discosiella Syd. & P. Syd. 1912, anamorphic Strigula Fr. 1823
Chaetothyriomycetidae, genera incertae sedis
Peyronelia Cif. & Gonz. Frag. 1927,
anamorphic Glyphium Nitschke ex F.
Lehm. 1886
Phaeomoniella Crous & W. Gams 2000*
Rhabdospora-like, anamorphic Dolabra C.
Booth & W.P. Ting 1964*
Taeniolella S. Hughes 1958*, anamorphic
Glyphium Nitschke ex F. Lehm. 1886
Subclass Eurotiomycetidae Geiser & Lutzoni
Coryneliales Seaver & Chardon
Coryneliaceae Sacc. ex Berl. & Voglino
Exophiala J.W. Carmich. 1966, anamorphic Bicornispora Checa, Barrasa, M.N.
Blanco & A.T. Martínez 1996
Eurotiales G.W. Martin ex Benny & Kimbr.
Monascaceae J. Schröt.
Basipetospora G.T. Cole & W.B. Kendr.
1968, anamorphic Monascus Tiegh.
?Fraseriella Cif. & A.M. Corte 1957,
anamorphic Xeromyces Fraser
Trichocomaceae E. Fisch.
Aspergillus P. Micheli ex Link 1809,
anamorphic Dichlaena Durieu & Mont.
1849, Emericella Berk. & Broome 1857,
Eurotium Link 1809, Fennellia B.J.
Wiley & E.G. Simmons 1973, Hemicarpenteles A.K. Sarbhoy & Elphick 1968,
Neopetromyces Frisvad & Samson 2000,
Neosartorya Malloch & Cain 1973*,
14
Petromyces Malloch & Cain 1973* and
Warcupiella Subram. 1972
Eladia G. Sm. 1961*
Geosmithia-like anamorphic Chromocleista Yaguchi & Udagawa 1993 and
Talaromyces C.R. Benj. 1955
Isaria Pers. 1794, anamorphic Byssochlamys Westling
Merimbla Pitt 1979, anamorphic Talaromyces C.R. Benj. 1955*
Paecilomyces Bainier 1907, anamorphic
Byssochlamys Westling 1909*
Paecilomyces-like, anamorphic Coonemeria Mouch. 1997
Penicillium Link 1809, anamorphic
Eupenicillium F. Ludw. 1892*, Talaromyces C.R. Benj. 1955* and Trichocoma Jungh. 1838
Polypaecilum G. Sm. 1961, anamorphic
Dactylomyces Sopp 1912, Dichotomomyces Saito ex D.B.Scott 1970 and
Thermoascus Miehe 1907
Pseudocordyceps Hauman 1936, anamorphic Penicilliopsis Solms 1887
Sagenomella W. Gams 1978, anamorphic
Sagenoma Stolk & G.F. Orr 1974
Sarophorum Syd. & P. Syd. 1916,
anamorphic Penicilliopsis Solms 1887
Stilbodendron Syd. & P. Syd. 1916,
anamorphic Penicilliopsis Solms 1887
Stilbothamnium Henn. 1896
Thysanophora W.B. Kendr. 1961
Torulomyces Delitsch 1943
Eurotiales, genera incertae sedis
Thermomyces Tsikl. 1899
Eurotiomycetidae genera incertae sedis
Penicillium Link 1809, anamorphic Paratalaromyces Matsush. 2003
Onygenales Cif. ex Benny & Kimbr.
Ajellomycetaceae Untereiner, J.A. Scott &
Sigler
Emmonsia Cif. & Montemart. 1959,
anamorphic Ajellomyces McDonough &
A.L. Lewis 1968
Histoplasma Darling 1906, anamorphic
Ajellomyces McDonough & A.L. Lewis
1968
Lacazia Taborda, V.A. Taborda &
McGinnis 1999*
Mycosphere
Loboa Cif., P.C. Azevedo, Campos &
Carneiro 1956
Paracoccidioides F.P. Almeida 1930
Chrysosporium-like, anamorphic Ascosphaera L.S. Olive & Spiltoir 1955 and
Bettsia Skou 1972
Arachnomycetaceae Gibas, Sigler & Currah
Onychocola Sigler 1990, anamorphic
Arachnomyces Massee & E.S. Salmon
1902
Onygenales, genera incertae sedis
Arthropsis Sigler, M.T. Dunn & J.W.
Carmich. 1982
Lacazia Taborda, V.A. Taborda &
McGinnis 1999
Gymnoascaceae Baran.
Chrysosporium-like, anamorphic Orromyces B. Sur & G.R. Ghosh 1987
Malbranchea Sacc. 1882, anamorphic
Byssoonygena Guarro, Punsola & Cano
1987 and Gymnoascus Baran. 1872
Oncocladium Wallr. 1833, anamorphic
Gymnoascus Baran. 1872
Onygenaceae Berk.
Castanedomyces Cano, L.B. Pitarch &
Guarro 2002
Chrysosporium Corda 1833 anamorphic
Amauroascus J. Schröt., Aphanoascus
Zukal 1890*, Apinisia La Touche 1968,
Arachnotheca Arx 1971, Nannizziopsis
Currah 1985, ?Polytolypa J.A. Scott &
Malloch 1993*, Pectinotrichum Varsavsky & G.F. Orr 1971, ?Renispora Sigler
& J.W. Carmich. 1979*
Coccidioides G.W. Stiles 1896
Malbranchea-like, anamorphic Auxarthron
G.F. Orr & Kuehn 1963, Byssoonygena
Guarro, Punsola & Cano 1987 and
Uncinocarpus Sigler & G.F. Orr 1976
Arthrodermataceae Currah
Chrysosporium Corda 1833, anamorphic
Shanorella R.K. Benj. 1956
Chrysosporium-like, anamorphic Ctenomyces Eidam 1880
Epidermophyton Sabour. 1907, anamorphic Arthroderma Curr. 1860
Microsporum Gruby 1843, anamorphic
Arthroderma Curr. 1860
Myceliophthora Costantin 1892,
anamorphic Arthroderma Curr. 1860
Trichophyton Malmsten 1848, anamorphic
Arthroderma Curr. 1860
Ascosphaeraceae L.S. Olive & Spiltoir
Subclass Mycocaliciomycetidae Tibell
Mycocaliciales Tibell & Wedin
Mycocaliciaceae A.F.W. Schmid
Asterophoma D. Hawksw. 1981, anamorphic Chaenothecopsis Vain. 1927
Catenomycopsis Tibell & Constant. 1991,
anamorphic Chaenothecopsis Vain.
1927
Phialophora-like, anamorphic Chaenothecopsis Vain. 1927 and Mycocalicium
Vain. 1890
Class Laboulbeniomycetes Engler
Laboulbeniales Engler
Pyxidiophorales P.F. Cannon
Pyxidiophoraceae Arnold
Chalara-like, anamorphic Pyxidiophora
Bref. & Tavel 1891
Gliocephalis Matr. 1899
Pleurocatena G. Arnaud ex Aramb.,
Gamundí, W. Gams & G.R.W. Arnold
2007, anamorphic Pyxidiophora Bref.
& Tavel 1891
Thaxteriola Speg. 1918, anamorphic
Pyxidiophora Bref. & Tavel 1891
Class Lecanoromycetes O.E. Erikss. & Winka
Subclass Ostropomycetidae Reeb, Lutzoni &
Cl. Roux
Agyriales Clem. & Shear
Trapeliaceae M. Choisy ex Hertel
Epithyrium
(Sacc.)
Trotter
1931,
anamorphic Sarea Fr. 1825
Pycnidiella Höhn. 1915, anamorphic Sarea
Fr. 1825
Ostropales Nannf.
Odontotremataceae D. Hawksw. & Sherwood
Pycnidial anamorph associated with
Claviradulomyces P.R. Johnst., D.C.
Park, H.C. Evans, R.W. Barreto & D.J.
Soares 2010
15
Stictidaceae Fr.
?Coleophoma-like, anamorphic Stictophacidium Rehm 1888
?Ebollia Minter & Caine 1980, anamorphic Stictophacidium Rehm 1888
Phacidiella-like anamorphic Acarosporina
Sherwood 1977
Stictospora Cif. 1957, anamorphic Stictis
Pers. 1800
Subclass Lecanoromycetidae P.M. Kirk, P.F.
Cannon, J.C. David & Stalpers ex Miadl.,
Lutzoni & Lumbsch
Lecanorales Nannf.
Pilocarpaceae Zahlbr.
Pyriomyces Bat. & H. Maia 1965,
anamorphic Byssoloma Trevis. 1853
Szczawinskia A. Funk 1984
Stereocaulaceae Chevall.
Lepraria Ach. 1803
Lecanorales, genera incertae sedis
Leprocaulon Nyl. 1878
Vouauxiomyces Dyko & D. Hawks. 1979,
anamorphic Abrothallus De Not. 1845
Class Leotiomycetes O.E. Erikss. & Winka
Cyttariales Luttr. ex Gamundı´
Cyttariaceae Speg.
Cytariella Palm 1932, anamorphic Cyttaria Berk. 1842
Erysiphales Gwynne-Vaughan
Erysiphaceae Tul. & C. Tul.
Oidiopsis Scalia 1902, anamorphic Leveillula G. Arnaud 1921
Oidium Link 1824, anamorphic Arthrocladiella Vassilkov 1960, Brasiliomyces
Viégas 1944, Cystotheca Berk. & M.A.
Curtis 1860, Erysiphe R. Hedw. ex DC.
1805 and Golovinomyces (U. Braun)
V.P. Gelyuta 1988, Neoerysiphe U.
Braun 1999*, Podosphaera Kunze 1823
and Sawadaea Miyabe 1914
Ovulariopsis Pat. & Har. 1900, anamorphic Phyllactinia Lév. 1851
Streptopodium R.Y. Zheng & G.Q. Chen
1978, anamorphic Pleochaeta Sacc. &
Speg. 1881
Helotiales Nannf.
16
Bulgariaceae Fr.
Crinula Fr. 1821, anamorphic Holwaya
Sacc. 1889
Endomelaconium Petr. 1940, anamorphic
Bulgaria Fr. 1822
Dermateaceae Fr.
Cadophora-like, anamorphic Phaeomollisia T.N. Sieber & Grünig 2009
Casaresia Gonz. Frag. 1920, anamorphic
Mollisia (Fr.) P. Karst. 1871
Chondropodium-like, anamorphic Waltonia Saho 1970
Corniculariella P. Karst. 1884, anamorphic Dermea Fr. 1825 and Durandiella
Seaver 1932
Cryptocline Petr. 1924, anamorphic
Trochila Fr. 1849
Cryptosporiopsis Bubák & Kabát 1912*,
anamorphic Neofabraea H.S. Jacks.
1913, Pezicula Tul. & C. Tul. 1865
and ?Ocellaria (Tul. & C. Tul.) P. Karst.
1871
Cryptosympodula Verkley 1999, anamorphic Scleropezicula Verkley 1999
Cylindrocolla Bonord. 1851, anamorphic
Calloria Fr. 1836
Cylindrosporium Grev. 1822, anamorphic
Pyrenopeziza Fuckel 1870
Cystodendron Bubák 1914, anamorphic
Mollisia (Fr.) P. Karst. 1871 and
Niptera Fr. 1849
Diplosporonema Höhn. 1917, anamorphic
Pyrenopeziza Fuckel 1917*
Discogloeum Petr. 1923, anamorphic
Spilopodia Boud. 1885
Entomosporium Lév. 1856, anamorphic
Diplocarpon F.A. Wolf 1912
Eriosporella Höhn. 1916, possibly
anamorphic Laetinaevia Nannf. 1932
Foveostroma DiCosmo 1978, anamorphic
Dermea Fr. 1825
Gelatinosporium Peck 1873, anamorphic
Dermea Fr. 1825 and Durandiella
Seaver 1932
Gloeosporidiella Petr. 1921, anamorphic
Drepanopeziza (Kleb.) Höhn. 1917 and
Pseudopeziza Fuckel 1870
Hainesia Ellis & Sacc. 1884, anamorphic
Discohainesia Nannf. 1932
Helicodendron Peyronel 1918, anamorphic
Mollisia (Fr.) P. Karst. 1871
Mycosphere
Laetinaevia Nannf. 1932, anamorphic
Naeviopsis B. Hein 1976
Marssonina Magnus 1906, anamorphic
Diplocarpon F.A. Wolf 1912 &
Drepanopeziza (Kleb.) Höhn. 1917
Melanodiscus Höhn. 1918, anamorphic
Spilopodia Boud. 1885
Microgloeum Petr. 1922, anamorphic
Blumeriella Arx 1961
Monostichella Höhn. 1916, anamorphic
Drepanopeziza (Kleb.) Höhn. 1917
Phacidiella P. Karst. 1884, anamorphic
Pyrenopeziza Fuckel 1870
Phialocephala W.B. Kendr. 1961*
Phialocephala-like, anamorphic Phaeomollisia T.N. Sieber & Grünig 2009
Phialophora-like, anamorphic Dibeloniella Nannf. 1932
Phloeosporella Höhn. 1924, anamorphic
Blumeriella Arx 1961
Phlyctema Desm. 1847, anamorphic Neofabraea H.S. Jacks. 1913
Pilidium Kunze 1823, anamorphic Discohainesia Nannf. 1932
Sporonema Desm. 1847, ?anamorphic
Leptotrochila P. Karst. 1871
Trichosporiella Kamyschko 1960, anamorphic Laetinaevia Nannf. 1932
Helotiaceae Rehm
Articulospora Ingold 1942, anamorphic
Hymenoscyphus Gray 1821
Ascoconidium Seaver 1942, anamorphic
Sageria A. Funk 1975
Bloxamia Berk. & Broome 1854, anamorphic Bisporella Sacc. 1884
Bothrodiscus Shear 1907, anamorphic
Ascocalyx Naumov 1926
Brunchorstia Erikss. 1891, anamorphic
Gremmeniella M. Morelet 1969
Bryophytomyces Cif. 1953, anamorphic
Hymenoscyphus Gray 1821
Coryne Nees 1816, anamorphic Ascocoryne J.W. Groves & D.E. Wilson
1967
Cystotricha Berk. & Broome 1850, ?anamorphic Xylogramma Wallr. 1833
Deltosperma W.Y. Zhuang 1988, anamorphic Unguiculariopsis Rehm 1909
Dendrostilbella Höhn. 1905, anamorphic
Claussenomyces Kirschst. 1923
Dichaenopsella Petr. 1952, type is
anamorphic Godroniopsis Diehl & E.K.
Cash 1929
Digitosporium Gremmen 1953 anamorphic
Crumenulopsis J.W. Groves 1969
Dimorphospora Tubaki 1958, anamorphic
Hymenoscyphus Gray 1821
Dothiorina Höhn. 1911, anamorphic Chlorociboria Seaver ex Ramamurthi, Korf
& L.R. Batra 1958
Endomelaconium Petr. 1940, anamorphic
Austrocenangium Gamundí 1997
Fuckelia Bonord. 1864, anamorphic
Godronia Moug. & Lév. 1846
Geniculospora Sv. Nilsson ex Marvanová
& Sv. Nilsson 1971, anamorphic Hymenoscyphus Gray 1821
Gliomastix-like anamorphic Ascoclavulina
Y. Otani 1974
Heteropatella Fuckel 1874, anamorphic
Heterosphaeria Grev. 1824
Hormonema-like, anamorphic Pragmopora A. Massal 1855
Idriella P.E. Nelson & S. Wilh. 1956,
anamorphic Hymenoscyphus Gray 1821
Periperidium Darker 1963, anamorphic
Micraspis Darker 1963
Phialophora-like, anamorphic Pseudopezicula Korf 1975
Pragmopycnis B. Sutton & A. Funk 1975,
anamorphic Pragmopora A. Massal
1855
Pseudospiropes M.B. Ellis 1971, anamorphic Strossmayeria Schulzer 1881
Rhinocladiella-like, anamorphic Bioscypha Syd. 1927
Rhizothyrium Naumov 1915, anamorphic
Rhizocalyx Petr. 1928
Scytalidium-like, anamorphic Hymenoscyphus Gray 1821
Sirodothis Clem. 1909, anamorphic Tympanis Tode 1790
Sporonema Desm. 1847, anamorphic
Godronia Moug. & Lév. 1846
Symphyosirella Seifert 2009, anamorphic
Symphyosirinia E.A. Ellis 1956*
Titaeospora Bubák 1916, anamorphic
Stamnaria Fuckel 1870
Topospora Fr. 1836, anamorphic Godronia
Moug. & Lév. 1846
17
Tricladium Ingold 1942, anamorphic
Cudoniella Sacc. 1889 and Hymenoscyphus Gray 1821*
Varicosporium W. Kegel 1906, anamorphic Hymenoscyphus Gray 1821*
Hemiphacidiaceae Korf
Meria Vuill. 1896, anamorphic Rhabdocline Syd. 1922
Rhabdogloeum Syd. 1922, anamorphic
Rhabdocline Syd. 1922
Rhabdogloeopsis Petr. 1925, anamorphic
Sarcotrochila Höhn. 1917
Hyaloscyphaceae Nannf.
Acleistia Bayl. Ell. (1917) anamorphic
Calycina Nees ex Gray 1821
Brefeldochium Verkley 2005, anamorphic
Polydesmia Boud. 1885
Chalara-like, anamorphic Calycellina
Höhn., Phaeoscypha Spooner 1984 and
Tapesina Lambotte 1887
Cheiromycella Höhn. 1910, anamorphic
Hyaloscypha Boud. 1885
Clathrosphaerina Beverw. 1951, anamorphic Hyaloscypha Boud. 1885
Haplographium Berk. & Broome 1859,
anamorphic Dematioscypha Svrček
1977
Mycoarthris Marvanová & P.J. Fisher
2002
Naemospora Roth ex Kuntze 1898,
possibly anamorphic Lachnellula P.
Karst. 1884
Phialophora-like, anamorphic Cistella
Quél. and Hyalopeziza Fuckel 1870
Pseudaegerita J.L. Crane & Schokn. 1981,
anamorphic Hyaloscypha Boud. 1885
Septonema-like anamorphic Ciliolarina
Svrcek 1977
Tricladium Ingold 1942, anamorphic
Hydrocina Scheuer 1991
Leotiaceae Corda
Alatospora Ingold 1942
Aureohyphozyma Hosoya & Y. Otani 1995,
anamorphic Gelatinipulvinella Hosoya
& Y. Otani 1995
Halenospora E.B.G. Jones 2009*
Myrioconium-like, anamorphic Neobulgaria Petr. 1921
18
Phialophora-like, anamorphic Neobulgaria Petr. 1921*
Loramycetaceae Dennis ex Digby & Goos
Anguillospora-like, anamorphic Loramyces W. Weston 1929
Phacidiaceae Fr.
Allantophomopsis Petr. 1925, anamorphic
Phacidium Fr. 1815
Apostrasseria Nag Raj 1983, anamorphic
Lophophacidium Lagerb. 1949 and
Phacidium Fr. 1815
Blennoria Moug. & Fr. 1825, anamorphic
Phacidium Fr. 1815
Ceuthospora Grev. 1826, anamorphic
Phacidium Fr. 1815
?Coma Nag Raj & W.B. Kendr. 1972,
anamorphic Ascocoma H.J. Swart
Rutstroemiaceae Holst-Jensen, L.M. Kohn &
T. Schumacher
Helicodendron Peyronel 1918, anamorphic
Lambertella Höhn. 1918
Myrioconium Syd. & P. Syd. 1912, ?anamorphic Lanzia Sacc. 1884, Rutstroemia P. Karst. 1871 and Scleromitrula S.
Imai 1941
Phialophora-like, anamorphic Rutstroemia
P. Karst. 1871
Sclerotiniaceae Whetzel ex Whetzel
Acarosporium Bubák & Vleugel ex Bubák
1911, anamorphic Pycnopeziza W.L.
White & Whetzel 1938
Amerosporium Speg. 1882, anamorphic
Zoellneria Velen. 1934
Amphobotrys Hennebert 1973, anamorphic
Botryotinia Whetzel 1945
Botrytis P. Micheli ex Pers. 1794,
anamorphic Botryotinia Whetzel 1945
Cristulariella Höhn. 1916, anamorphic
Nervostroma Narumi & Y. Harada 2006
Haradamyces Masuya, Kusunoki, Kosaka
& Aikawa 2009*
Hinomyces Narumi & Y. Harada 2006,
anamorphic Grovesinia M.N. Cline, J.L.
Crane & S.D. Cline 1983
Monilia Bonord. 1851, anamorphic Monilinia Honey 1928 and Phaeosclerotinia
Hori 1916
Mycosphere
Mycopappus Redhead & G.P. White 1985,
anamorphic Redheadia Y. Suto &
Suyama 2005
Myrioconium Syd. & P. Syd. 1912,
anamorphic Ciborinia Whetzel 1945,
Encoelia (Fr.) P. Karst. 1871, Martininia Dumont & Korf 1970, Myriosclerotinia N.F. Buchw. 1947 and
Sclerotinia Fuckel 1870
Ovulitis N.F. Buchw. 1970, anamorphic
Ovulinia F.A. Weiss 1940
Sclerotium Tode 1790, anamorphic Myriosclerotinia N.F. Buchw. 1947, Sclerotinia Fuckel 1870* and Stromatinia
(Boud.) Boud. 1907*
Septotis N.F. Buchw. ex Arx 1970,
anamorphic Septotinia Whetzel ex J.W.
Groves & M.E. Elliott 1961
Streptobotrys Hennebert 1973, anamorphic
Streptotinia Whetzel 1945
Valdensia Peyronel 1923, anamorphic
Valdensinia Peyronel 1953
Verrucobotrys Hennebert 1973, anamorphic Seaverinia Whetzel 1945
Vibrisseaceae Korf
Acephala Grünig & T.N. Sieber 2005*
Anavirga B. Sutton 1975, anamorphic
Vibrissea Fr. 1822
Phialocephala W.B. Kendr. 1961, some
possibly anamorphic Vibrissea Fr.
1822*
Phialophora-like anamorphic Chlorovibrissea L.M. Kohn 1989
Helotiales, genera incertae sedis
Cadophora Lagerb. & Melin 1927*
Catenulifera Hosoya 2002, anamorphic
Hyphodiscus Kirschst. 1906*
Chaetochalara B. Sutton & Piroz. 1965
Chalara (Corda) Rabenh. 1844*
Clathrosporium Nawawi & Kuthub. 1987
Crucellisporium M.L. Farr 1968*
Cystotricha Berk. & Broome 1850
Dactylaria Sacc. 1880
Diplococcium Grove 1885*
Endoconidium Prill. & Delacr. 1891,
anamorphic Gloeotinia M. Wilson,
Noble & E.G. Gray 1954
Fontanospora Dyko 1978*
Glarea Bills & Paláez 1999
Helgardia Crous & W. Gams 2003, anamorphic Oculimacula Crous & W.
Gams 2003*
Hyalodendriella Crous 2007
Hyphodiscus Kirschst. 1906
Hysteropezizella Höhn. 1917
Leptodontidium de Hoog 1979*
Libartania Nag Raj 1979
Margaritispora Ingold 1942
Mirandina G. Arnaud ex Matsush. 1975
Monochaetiellopsis B. Sutton & DiCosmo
1977, anamorphic Hypnotheca Tommerup 1970
Mycochaetophora Hara & Ogawa 1931*
Neozythia Petr. 1958
Rhexocercosporidium U. Braun 1994*
Rhizocladosporium Crous & U. Braun
2007
Rhodesia Grove 1937
Rhynchosporium Heinsen ex A.B. Frank
1897*
Scytalidium Pesante 1957
Sphaerographium Sacc. 1884
Spirosphaera Beverw. 1953
Tetracladium De Wild. 1893
Thedgonia B. Sutton 1973*
Trimmatostroma Corda 1837
Variocladium Descals & Marvanová
1999*
Rhytismatales M.E. Barr ex Minter
Ascodichaenaceae D. Hawksw. & Sherwood
Dichaenopsella Petr. 1952, anamorphic
Ascodichaena Butin 1977
Macroallantina Speer 1987, anamorphic
Delpinoina Kuntze, 1891
Polymorphum Chevall. 1822, anamorphic
Ascodichaena Butin 1977
Cryptomycetaceae Höhn.
?Phacidiopycnis Potebnia 1912, anamorphic Potebniamyces Smerlis 1962
Rhytismataceae Chevall.
Crandallia Ellis & Sacc.
morphic Bifusella Höhn.
Duplicaria Fuckel 1870
Conostroma Moesz 1921,
Colpoma Wallr. 1833
Cryocaligula Minter 1986,
Ploioderma Darker. 1967
1897, ana1917 and
anamorphic
anamorphic
19
Hysterodiscula Petr. 1942, anamorphic
Duplicaria Fuckel 1870
Leptostroma Fr. 1815, anamorphic Lophodermium Chevall. 1826
Melasmia Lév. 1846, anamorphic Rhytisma Fr. 1818*
Tryblidiopycnis Höhn. 1918, anamorphic
Tryblidiopsis P. Karst. 1871
Rhytismatales, genera incertae sedis
Fuligomyces Morgan-Jones & Kamal 1984
Uyucamyces H.C. Evans & Minter 1985,
anamorphic Ocotomyces H.C. Evans &
Minter 1985
Leotiomycetes, families incertae sedis
Myxotrichaceae Currah
Geomyces Traaen 1914, anamorphic
Pseudogymnoascus Raillo 1929
Gymnostellatospora Udagawa, Uchiy. &
Kamiya 1993, ?anamorphic Pseudogymnoascus Raillo 1929
Oidiodendron Robak 1932, anamorphic
Byssoascus Arx 1971 and Myxotrichum
Kunze 1823
Malbranchea Sacc. 1882, ?anamorphic
Myxotrichum Kunze 1823
Leotiomycetes, genera incertae sedis
Chaetomella Fuckel 1870
Collophora Damm & Crous 2010*
Eleutheromyces Fuckel 1870,
Geniculospora Sv. Nilsson ex Marvanová
& Sv. Nilsson 1971
Hyphozyma de Hoog & M.T. Sm. 1981
Leohumicola N.L. Nick. 2005
Meliniomyces Hambl. & Sigler 2005
Scytalidium Pesante 1957, anamorphic
Xylogone Arx & T. Nilsson 1969*
Tiarosporella Höhn. 1924, anamorphic
Darkera H.S. Whitney, J. Reid & Piroz.
1975
Class Orbiliomycetes O.E. Erikss. & Baral
Orbiliales Baral, O.E. Erikss., G. Marson & E.
Weber
Orbiliaceae Nannf.
Arthrobotrys Corda 1939, anamorphic
Orbilia Fr. 1836*
Brachyphoris Juan Chen, L.L. Xu, B. Liu
& Xing Z. Liu 2007, anamorphic
Hyalorbilia Baral & G. Marson 2000
20
Dactylella Grove 1884, anamorphic Orbilia Fr. 1836
Dicranidion Harkn. 1885, anamorphic Orbilia Fr. 1836
Drechslerella Subram. 1964, anamorphic
Orbilia Fr. 1836*
Dwayaangam Subram. 1978, anamorphic
Orbilia Fr. 1836
Gamsylella M. Scholler, Hagedorn & A.
Rubner 1999, anamorphic Orbilia Fr.
1836
Monacrosporium Oudem. 1885, anamorphic Orbilia Fr. 1836
Trinacrium Riess 1852, anamorphic Orbilia Fr. 1836
Class Pezizomycetes sensu O.E. Erikss. &
Winka
Pezizales J. Schröt.
Ascobolaceae Boud. ex Sacc.
Rhizostilbella Wolk 1914, anamorphic
Ascobolus Pers. 1792
Caloscyphaceae Harmaja
Geniculodendron G.A. Salt 1974, anamorphic Caloscypha Boud. 1885
Chorioactidaceae Pfister
Kumanasamuha P.Rag. Rao & D. Rao
1964, anamorphic Chorioactis Kupfer
ex Eckblad 1968*
Verticicladium Preuss 1851, anamorphic
Desmazierella Lib. 1829
Morchellaceae Reichenb.
Costantinella Matr. 1892, anamorphic
Morchella Dill. ex Pers. 1794
Pezizaceae Dumort.
Chromelosporium Corda 1833, anamorphic Muciturbo P.H.B. Talbot 1989,
Peziza Dill. ex Fr. 1822 and Plicaria
Fuckel 1870
Glischroderma Fuckel 1870
Oedocephalum Preuss 1851, anamorphic
Iodophanus Korf 1967 and Peziza Dill.
ex Fr. 1822
Ostracoderma Fr. 1825, anamorphic Peziza Dill. ex Fr. 1822
Pyronemataceae Corda
Actinospora Ingold 1952,
Mycosphere
Actinosporella Descals, Marvanová & J.
Webster 1999, anamorphic Miladina
Svrček 1972
Ascorhizoctonia Chin S. Yang & Korf
1985, anamorphic Tricharina Eckblad
1968
Complexipes C. Walker 1979, anamorphic
Tricharina Eckblad 1968 and Wilcoxina
Chin S. Yang & Korf 1985
Dichobotrys Hennebert 1973, ?anamorphic
Pyropyxis Egger 1984 and Trichophaea
Boud. 1885
Micronematobotrys Xiang Sun & L.D.
Guo 2010
Scytalidium-like, anamorph of Anthracobia Boud. 1885
Rhizinaceae Bonord.
Phymatotrichopsis Hennebert 1973
Sarcoscyphaceae LeGal ex Eckblad
Molliardiomyces Paden 1984, anamorphic
Nanoscypha Denison 1972, Phillipsia
Berk. 1881, Pithya Fuckel 1870 and
Sarcoscypha (Fr.) Boud. 1885
Sarcosomataceae Kobayasi
Conoplea Pers. 1797 and ?anamorphic
Plectania Fuckel 1870
Strumella Fr. 1849, anamorphic Urnula Fr.
1849
Verticicladium Preuss 1851, anamorphic
Sarcosoma Casp. 1891
Pezizales, genera incertae sedis
Cephaliophora Thaxt. 1903
Class Sordariomycetes sensu O.E. Erikss. &
Winka
Subclass Hypocreomycetidae O.E. Erikss. &
Winka
Coronophorales Nannf.
Chaetosphaerellaceae Huhndorf, A.N. Mill. &
F.A. Fernández
Oedemium Link 1824, anamorphic
Chaetosphaerella E. Müll. & C. Booth
1972
Veramycina Subram. 1995, anamorphic
Chaetosphaerella E. Müll. & C. Booth
1972
Nitschkiaceae (Fitzp.) Nannf.
Acremonium-like anamorphic
nitschkea Speg.
Acantho-
Hypocreales Lindau
Bionectriaceae Samuels & Rossman
Acremonium-like, anamorphic Dimerosporiella Speg. 1908, Ijuhya Starbäck
1899*, Hydropisphaera Dumont 1822*,
Lasionectria (Sacc.) Cooke 1884, Mycoarachis Malloch & Cain 1970*,
Mycocitrus Moller 1901, Nectriella
Nitschke ex Fuckel 1870, Nectriopsis
Maire 1911*, Nigrosabulum Malloch &
Cain 1970*, Protocreopsis Yoshim. Doi
1977*, Selinia P. Karst. 1876*, and
Stilbocrea Pat. 1900 and Verrucostoma
Hirooka, Tak. Kobay. & Chaverri
2010*
Albosynnema E.F. Morris 1967
Clonostachys Corda 1839, anamorphic
Bionectria Speg. 1919
Dendrodochium-like, anamorphic Nectriella Nitschke ex Fuckel 1870
Didymostilbe Henn. 1902, anamorphic
Peethambara Subram. & Bhat 1978
Gliocladium-like, anamorphic Roumegueriella Speg. 1880
Gliomastix Guég. 1905, anamorphic Hydropisphaera Dumort. 1822*
Gracilistilbella Seifert 2000, anamorphic
Stilbocrea Pat. 1900
Rhopalocladium Schroers, Samuels & W.
Gams 1999, anamorphic Nectriopsis
Maire 1911
Stanjemonium W. Gams, O'Donnell,
Schroers & M. Chr. 1999
Stilbella-like, anamorphic Stilbocrea Pat.
1900
Tubercularia-like, anamorphic Selinia P.
Karst. 1876
Verticillium-like, anamorphic Nectriopsis
Maire 1911*
Vesicladiella Crous & M.J. Wingf. 1994
Clavicipitaceae (Lindau) Earle ex Rogerson
Acremonium-like, anamorphic Neobarya
Lowen 1986
Albomyces I. Miyake 1908, anamorphic
Aciculosporium I. Miyake
Aschersonia Mont. 1848, anamorphic
Hypocrella Sacc. and Moelleriella Bres.
1897.*
21
Blistum B. Sutton 1973, anamorphic
Berkelella (Sacc.) Sacc. 1891
Corallocytostroma Y.N. Yu & Z.Y. Zhang
1980
Drechmeria W. Gams & H.-B. Jansson
1985
Ephelis Fr. 1849, anamorphic Atkinsonella
Diehl, Balansia Speg., 1885, Epichloë
(Fr.) Tul. & C. Tul., 1865 and Linearistroma Höhn. 1910
Harposporium Lohde 1874, anamorphic
Podocrella Seaver 1928
Hirsutella-like, anamorphic Orbiocrella D.
Johnson, G.H. Sung, Hywel-Jones &
Spatafora 2009 and Moelleriella Bres.
1897*
Metarhizium Sorokīn 1879, anamorphic
Metacordyceps G.H. Sung, J.M. Sung,
Hywel-Jones & Spatafora 2007*
Neotyphodium Glenn, C.W. Bacon &
Hanlin 1996, anamorphic Epichloë (Fr.)
Tul. & C. Tul. 1865
Nomuraea Maubl. 1903
Polycephalomyces Kobayasi 1941, anamorphic Berkelella (Sacc.) Sacc. 1891
Pseudomeria G.L. Barron 1980
Sphacelia Lév. 1827, anamorphic Atkinsonella Diehl 1950 and Claviceps Tul.
1853
Cordycipitaceae Kreisel ex G.M. Sung, J.M.
Sung, Hywel-Jones & Spatafora
Akanthomyces Lebert 1858, anamorphic
Cordyceps Fr. 1824 and Torrubiella
Boud. 1885*
Beauveria Vuill. 1912, anamorphic Cordyceps Fr. 1824
Engyodontium de Hoog 1978
Gibellula Cavara 1894, anamorphic Torrubiella Boud. 1885*
Granulomanus de Hoog & Samson 1978,
anamorphic Torrubiella Boud. 1885
Isaria Pers. 1794
Lecanicillium W. Gams & Zare 2001,
anamorphic Cordyceps Fr. 1824 and
Torrubiella Boud. 1885*
Microhilum H.Y. Yip & A.C. Rath 1989
?Microides H.Y.Yip & A.C. Rath 1989
Pseudogibellula Samson & H.C. Evans
1973
Rotiferophthora G.L. Barron 1991
22
Simplicillium W. Gams & Zare 2001,
possibly anamorphic Torrubiella Boud.
1885*
Hypocreaceae De Not.
Acremonium-like, anamorphic Protocrea
Petch 1937 and Pseudohypocrea
Yoshim. Doi 1972
Acrostalagmus Corda 1838
Arachnocrea Z. Moravec 1956, Protocrea
Petch 1937 and Sarawakus Lloyd 1924
Cladobotryum Nees 1816, anamorphic
Hypomyces (Fr.) Tul. & C. Tul. 1860
Gliocladium Corda 1840, anamorphic
Hypocrea Fr. 1825* and Sphaerostilbella (P. Henn.) Sacc. & D. Sacc. 1905
Gliocladium-like, anamorphic Sarawakus
Lloyd 1924
Mycogone Link 1809, anamorphic Hypomyces (Fr.) Tul. & C. Tul. 1860
Sepedonium Link 1809, anamorphic Hypomyces (Fr.) Tul. & C. Tul. 1860
Sibirina G.R.W. Arnold 1970, anamorphic
Hypomyces (Fr.) Tul. & C. Tul. 1860
Stephanoma Wallr. 1833, anamorphic
Hypomyces (Fr.) Tul. & C. Tul. 1860
Stromatocrea W.B. Cooke 1952, anamorphic Hypocreopsis P. Karst. 1873
Trichoderma Pers. 1794, anamorphic
Hypocrea Fr. 1825 and Podostroma P.
Karst. 1892
Trichoderma-like, anamorphic Sarawakus
Lloyd 1924
Verticillium-like anamorphic Aphysiostroma Barrasa, A.T. Martínez & G.
Moreno 1986*
Nectriaceae Tul. & C. Tul.
Acremonium-like, anamorphic Cosmospora Rabenh. 1862*, Nectria (Fr.) Fr.
1849 and Neocosmospora E.F. Sm.
1899
Actinostilbe Petch 1925, anamorphic Lanatonectria Samuels & Rossman 1999
Antipodium Piroz. 1974, anamorphic
Ophionectria Sacc. 1878*
Aphanocladium W. Gams 1971
Calostilbella Höhn. 1919, anamorphic
Calostilbe Sacc. & Syd. 1902
Chaetopsina Rambelli 1956, anamorphic
Chaetopsinectria J. Luo & W.Y.
Mycosphere
Zhuang* and Cosmospora Rabenh.
1862
Ciliciopodium violaceum Corda 1831,
anamorphic Nectria (Fr.) Fr. 1849
Curvicladiella Decock & Crous 2006
Curvicladium Decock & Crous 1998
Cyanochyta Höhn. 1915, anamorph of
Gibberella Sacc. 1877
Cyanophomella Höhn. 1918, anamorph of
Gibberella Sacc. 1877
Cylindrocarpon Wollenw. 1913, anamorphic Neonectria Wollenw. 1917*
Cylindrocladiella Boesew. 1982, anamorphic Cosmospora Rabenh. 1862 and
Nectricladiella Crous & C.L. Schoch
2000
Cylindrocladium Morgan 1892, anamorphic Calonectria De Not. 1867*
Dacryoma Samuels 1988, anamorphic
Nectria (Fr.) Fr. 1849
Dematiocladium Allegr., Aramb., Cazau &
Crous 2005
Dichomera Cooke 1878, type is anamorph
of Gibberella Sacc. 1877
Flagellospora Ingold 1942, anamorphic
Nectria (Fr.) Fr. 1849
Fusarium Link 1809, anamorphic ?Albonectria Rossman & Samuels 1999, Cyanonectria Samuels & Chaverri 2009*,
Gibberella Sacc. 1877 and ?Haematonectria Samuels & Nirenberg 1999*
and unknown teleomorphs*
Fusarium-like, anamorphic Corallomycetella Henn. 1904 and Cosmospora
Rabenh. 1862
Fusidium Link 1809, type is anamorphic
Neonectria Wollenw. 1917
Gliocephalotrichum J.J. Ellis & Hesselt.
1962, anamorphic Leuconectria Rossman, Samuels & Lowen 1993*
Gliocladiopsis S.B. Saksena 1954, anamorphic Glionectria Crous & S.L.
Schoch 2000
Gyrostroma Naumov 1914, anamorphic
Nectria (Fr.) Fr. 1849*
Heliscus Sacc. 1880, anamorphic Nectria
(Fr.) Fr. 1849
Mariannaea G. Arnaud ex Samson 1974*
?Mirandina G. Arnaud ex Matsush. 1975,
anamorphic Chaetonectrioides Matsush.
Nalanthamala Subram. 1956, anamorphic
Rubrinectria Rossman & Samuels 1999
Penicillifer Emden 1968, anamorphic Viridispora Samuels & Rossman 1999*
Pleurocolla Petr. 1924
Rhizostilbella-like, anamorphic Corallomycetella Henn. 1904
Septofusidium W. Gams 1971
Stilbella Lindau 1900, anamorphic Cosmospora Rabenh. 1862
Trichothecium-like, anamorphic Rodentomyces Doveri, Pecchia, Sarrocco &
Vannacci 2010*
Tubercularia Tode 1790, anamorphic
Nectria (Fr.) Fr. 1849
Verticillium-like, anamorphic Cosmospora
Rabenh. 1862*
Volutella Fr. 1832, anamorphic Cosmospora Rabenh. 1862 and Pseudonectria
Seaver 1909*
Xenocylindrocladium Decock, Hennebert
& Crous 1997, anamorphic Xenocalonectria Crous & C.L. Schoch 2000
Zythiostroma Höhn. ex Falck 1923,
anamorphic Nectria (Fr.) Fr. 1849*
Niessliaceae Kirschst.
Acremonium-like, anamorphic Trichosphaerella E. Bommer, M. Rousseau &
Sacc. 1891 and Valetoniellopsis Samuels & M.E. Barr 1998
Custingophora-like, anamorphic Melanopsamma Niessl 1876
Monocillium S.B. Saksena 1955, anamorphic Hyaloseta A.W. Ramaley 2001
Monocillium S.B. Saksena 1955, anamorphic Niesslia Auersw. 1869
Ophiocordycipitaceae G.H. Sung, J.M. Sung,
Hywel-Jones & Spatafora
?Chaunopycnis W. Gams 1979
Haptocillium W. Gams & Zare 2001
Hirsutella Pat. 1892, anamorphic Ophiocordyceps Petch 1931*
Hymenostilbe Petch 1931, anamorphic
Ophiocordyceps Petch 1931
Paraisaria Samson & B.L. Brady 1983,
anamorphic Ophiocordyceps Petch
1931*
Syngliocladium Petch 1932, anamorphic
Ophiocordyceps Petch 1931
Tolypocladium W. Gams 1971, anamorphic Elaphocordyceps G.H. Sung &
Spatafora 2007
23
Verticillium-like, anamorphic Elaphocordyceps G.H. Sung & Spatafora 2007
Hypocreales, genera incertae sedis
Acremonium Link 1809
Acremonium-like, anamorphic Emericellopsis J.F.H. Beyma 1940 and
Leucosphaerina Arx 1987
Ascochytopsis-like, anamorphic Eucasphaeria Crous 2007
Calcarisporium Preuss 1851
Cancellidium Tubaki 1975
Cephalosporiopsis Peyronel 1915
Didymostilbe Henn. 1902, anamorphic
Peethambara Subram. & Bhat 1978
Escovopsis J.J. Muchovej & Della Lucia
1990
Geosmithia Pitt 1979*
Hapsidospora Malloch & Cain 1970
Haptospora G.L. Barron 1991
Illosporiopsis D. Hawksw. 2001
Illosporium Mart. 1817
Imicles Shoemaker & Hambl. 2001
Memnoniella Höhn. 1923
Munkia Speg. 1886
Myrothecium Tode 1790
Neomunkia Petr. 1947
Pseudomicrodochium B. Sutton 1975
Sarocladium W. Gams & D. Hawksw.
1976
Septomyrothecium Matsush. 1971
Sorosporella Sorokīn 1888
Spicellum Nicot & Roquebert 1976
Sporothrix Hektoen & C.F. Perkins 1901,
anamorphic Leucosphaerina Arx 1987
Stachybotrys Corda 1837
Stilbella Lindau 1900
Tilachlidium Preuss 1851
Trichothecium Link 1809
Ustilaginoidea Bref. 1895
Melanosporales N. Zhang & M. Blackw.
Ceratostomataceae G. Winter
Gonatobotrys Corda 1839, anamorphic
Melanospora Corda 1837
Harzia Costantin 1888, anamorphic Melanospora Corda 1837
Papulaspora Preuss 1851, anamorphic ?Melanospora Corda 1837
Proteophiala Cif. 1958, anamorphic Melanospora Corda 1837
24
[Sphaerodes Clem. 1909 is a teleomorph
which was reported to produce ampulliform phialides on irregularly branched
conidiophores diectrly on ascomata or
on surrounding hyphae (Vujanovic &
Goh, 2009)]
Microascales Luttr. ex Benny & Kimbr.
Chadefaudiellaceae Faurel & Schotter ex
Benny & Kimbr.
Arthrographis–like, anamorphic Faurelina
Locq.-Lin. 1975
Halosphaeriaceae E. Müll. & Arx ex Kohlm.
Cirrenalia Meyers & R.T. Moore 1960*
Clavariopsis-like, anamorphic Corollospora Werderm. 1922
Clavatospora Sv. Nilsson ex Marvanová &
Sv. Nilsson 1971, anamorphic Corollospora Werderm. 1922
Culcitaina Meyers & R.T. Moore 1960
Halosigmoidea Nakagiri, K.L. Pang &
E.B.G. Jones 2009, anamorphic Corollospora Werderm. 1922*
Humicola-like, anamorphic Pseudolignincola Chatmala & E.B.G. Jones 2006,
Monodictys-like, anamorphic Nereiospora
E.B.G. Jones, R.G. Johnson & S.T.
Moss 1983
Periconia-like, anamorphic Halosphaeria
Linder 1944
Sigmoidea J.L. Crane 1968 anamorphic
Corollospora Werderm. 1922
Trichocladium Harz 1871, anamorphic
Halosphaeria Linder 1944
Varicosporina Meyers & Kohlm. 1965,
anamorphic Corollospora Werderm.
1922*
Microascaceae Luttr. ex Malloch
Atrhrographis-like, anamorphic Pithoascus Arx 1973
Brachyconidiellopsis Decock, R.F. Castañeda & Adhikari 2004
Cephalotrichum Link 1809, anamorphic
Microascus Zukal 1885
Echinobotryum Corda 1831
Graphium Corda 1837, anamorphic Microascus Zukal 1885, Parascedosporium
Gilgado, Gené, Cano & Guarro 2007,
Petriella Curzi 1930 and Pseudallescheria Negr. & I. Fisch. 1944*
Mycosphere
Humicola-like, anamorphic Lophotrichus
R.K. Benj. 1949
Parascedosporium Gilgado, Gené, Cano &
Guarro 2007
Scedosporium Sacc. ex Castell. & Chalm.
1919, anamorphic Pseudallescheria
Negr. & I. Fisch. 1944* and possibly
Petriella Curzi 1930
Scopulariopsis Bainier 1907, anamorphic
Microascus Zukal 1885 and ?Petriella
Curzi 1930 and Pithoascus Arx 1973
Trichurus Clem. 1896
Wardomyces F.T. Brooks & Hansf. 1923,
anamorphic Microascus Zukal 1885
Wardomycopsis Udagawa & Furuya 1978,
anamorphic Microascus Zukal 1885
Microascales, genera incertae sedis
Bisporostilbella Brandsb. & E.F. Morris
1971
Chalara-like anamorphic Ceratocystis
Ellis & Halst. 1890 and Cornuvesica
C.D. Viljoen, M.J. Wingf. & K. Jacobs
2000
?Custingophora Stolk, Hennebert &
Klopotek 1968, anamorphic Gondwanamyces G.J. Marais & M.J. Wingf.
1998*
Gabarnaudia Samson & W. Gams 1974,
anamorphic Sphaeronaemella P. Karst.
1884*
Sporendocladia G. Arnaud ex Nag Raj &
W.B. Kendr. 1975
Thielaviopsis Went 1893, anamorphic
Ceratocystis Ellis & Halst. 1890
Hypocreomycetidae, families incertae sedis
Glomerellaceae Locq. ex Seifert & W. Gams
Colletotrichum Corda 1831, anamorphic
Glomerella Spauld. & Schrenk 1903*
Plectosphaerellaceae W. Gams, Summerbell &
Zare
Acremonium Link 1809*
Gibellulopsis Bat. & H. Maia 1959
Musicillium Zare & W. Gams 2007
Plectosporium M.E. Palm, W. Gams &
Nirenberg 1995, anamorphic Plectosphaerella Kleb. 1929
Verticillium Nees 1816
Hypocreomycetidae, genera incertae sedis
Canalisporium Nawawi & Kuthub. 1989,
anamorphic Ascothailandia Sri-indrasutdhi, Boonyuen, Sivichai & E.B.G.
Jones 2010*
Dictyochaeta-like, anamorphic Ascocodinaea Samuels, Candoussau & Magni
1997
Glomerulispora Abdel-Wahab, AbdelAziz & Nagahama 2010, ?anamorphic
Torpedospora Meyers 1957*
Moheitospora Abdel-Wahab, Abdel-Aziz
& Nagahama 2010, anamorphic Juncigena Kohlm., Volkm.-Kohlm & O.E.
Erikss. 1997*
Sporoschismopsis Hol.-Jech. & Hennebert
1972, anamorphic Porosphaerellopsis
Samuels & E. Müll. 1982
Subclass Sordariomycetidae O.E. Erikss. &
Winka
Calosphaeriales M.E. Barr*
Calosphaeriaceae Munk
Calosphaeriophora Réblová, L. Mostert,
W. Gams & Crous 2004, anamorphic
Calosphaeria Tul. & C. Tul.
Phaeocrella Réblová, L. Mostert, W.
Gams & Crous 2004, anamorphic Togniniella Réblová, L. Mostert, W. Gams
& Crous 2004
Phialophora-like, anamorphic Jattaea Berl.
1900
Stachybotrys-like, anamorphic Jattaea
Berl. 1900
Pleurostomataceae Réblová, L. Mostert, W.
Gams & Crous
Pleurostomophora Vijaykr., L. Mostert,
Jeewon, W. Gams, K.D. Hyde & Crous
2004, anamorphic Pleurostoma Tul. &
C. Tul. 1863
Chaetosphaeriales Huhndorf, A.N. Mill. &
F.A. Fernández
Chaetosphaeriaceae Réblová, M.E. Barr &
Samuels
Cacumisporium Preuss 1851, anamorphic
Chaetosphaeria Tul. & C. Tul. 1863
Catenularia Grove 1886, anamorphic
Chaetosphaeria Tul. & C. Tul. 1863
Chalara-like, anamorphic Ascochalara
Réblová 1999 and Chaetosphaeria Tul.
& C. Tul. 1863
25
Chloridium Link 1809, anamorphic Chaetosphaeria Tul. & C. Tul. 1863 and
Melanopsammella Höhn. 1920
Codinaea Maire 1937, anamorphic Chaetosphaeria Tul. & C. Tul. 1863
Codinaeopsis Morgan-Jones 1976
Cordana Preuss 1851, ?anamorphic Porosphaerella E. Müll. & Samuels 1982
Craspedodidymum Hol.-Jech. 1972, anamorphic Chaetosphaeria Tul. & C. Tul.
1863
Cryptophiale Piroz. 1968, anamorphic
Chaetosphaeria Tul. & C. Tul. 1863
Cylindrotrichum Bonord. 1851, anamorphic Chaetosphaeria Tul. & C. Tul.
1863
Dictyochaeta Speg. 1923, anamorphic
Chaetosphaeria Tul. & C. Tul. 1863
and Striatosphaeria Samuels & E. Müll.
1979
Dictyochaetopsis Aramb. & Cabello 1990
Dischloridium B. Sutton 1977, anamorphic
Australiasca Sivan. & Alcorn 2002
Exserticlava S. Hughes 1978, anamorphic
Chaetosphaeria Tul. & C. Tul. 1863
Fusichalara S. Hughes & Nag Raj 1973,
anamorphic Chaetosphaeria Tul. & C.
Tul. 1863
Gonytrichum Nees & T. Nees 1818,
anamorphic Chaetosphaeria Tul. & C.
Tul. 1863 and ?Melanopsammella Höhn.
1920
Hemicorynespora M.B. Ellis 1972, anamorphic Chaetosphaeria Tul. & C. Tul.
1863
Kylindria DiCosmo, S.M. Berch & W.B.
Kendr. 1983
Lecythothecium Réblová & Winka 2001
Menispora Pers. 1822, anamorphic Chaetosphaeria Tul. & C. Tul. 1863
Nawawia Marvanová 1980*
Paliphora Sivan. & B. Sutton 1985*
Phaeostalagmus W. Gams 1976, anamorphic Chaetosphaeria Tul. & C. Tul.
1863
Phialogeniculata Matsush. 1971
Phialophora-like, anamorphic Chaetosphaeria Tul. & C. Tul. 1863
Pseudobotrytis Krzemien. & Badura 1954,
anamorphic Porosphaerella E. Müll. &
Samuels 1982
26
Sporidesmium-like, anamorphic Umbrinosphaeria Réblová 1999
Sporoschisma Berk. & Broome 1847,
anamorphic Melanochaeta E. Müll.,
Harr & Sulmont 1969
Thozetella Kuntze 1891
Zanclospora S. Hughes & W.B. Kendr.
1965, anamorphic Chaetosphaeria Tul.
& C. Tul. 1863
Chaetosphaeriales, genera incertae sedis
Rattania Prabhugaonkar & Bhat 2009*
Coniochaetales Huhndorf, A.N. Mill. & F.A.
Fernández
Coniochaetaceae Malloch & Cain
Lecythophora Nannf. 1934, anamorphic
Coniochaeta (Sacc.) Cooke 1887*
Phialophora-like anamorphic Barrina
A.W. Ramaley 1997
Diaporthales Nannf.
Cryphonectriaceae Gryzenh. & M.J. Wingf.
Aurapex Gryzenh. & M.J. Wingf. 2006
Chrysoporthella Gryzenh. & M.J. Wingf.
2004, anamorphic Chrysoporthe Gryzenh. & M.J. Wingf. 2004*
Endothiella Sacc. 1906, anamorphic Cryphonectria (Sacc.) Sacc. & D. Sacc.
1905*
Foliocryphia Cheewangkoon & Crous
2010*
Prosopidicola Crous & C.L. Lennox 2004
Ursicollum Gryzenh. & M.J. Wingf. 2006
Diaporthaceae Höhn. ex Wehm.
Mazzantiella Höhn. 1925, anamorphic
Mazzantia Mont. 1855
Phomopsis (Sacc.) Bubák 1905, anamorphic Diaporthe Nitschke 1870
Gnomoniaceae G. Winter
Asteroma DC. 1815, anamorphic Gnomoniella Sacc. 1881 and Plagiostoma
Fuckel 1870 and Pleuroceras Riess.
The type of the genus is considered to
be Montagnellina stellaris (Pers.)
Theiss. & Syd. 1915
Cylindrosporella Höhn. 1916, type is
anamorphic Gnomonia Ces. & De Not.
1863, others Gnomoniella Sacc. 1881
and Pleuroceras Riess 1854
Mycosphere
Depazea Fr. 1818, ?anamorphic Pleuroceras Riess 1854
Diplodina Westend. 1857, anamorphic
Plagiostoma Fuckel 1870
Discosporium Sacc. & P. Syd. 1902 anamorphic Cryptodiaporthe Petr. 1921 C.
Tul.]
Discula Sacc. 1884, anamorphic Apiognomonia Höhn. 1917, Gnomonia Ces. &
De Not. 1863 and Amphiporthe Petr.
1971
Gloeosporidina Petr. 1921, anamorphic
Apiognomonia Höhn. 1917
Mazzantiella Höhn. 1925, anamorphic
Mamianiella Höhn. 1917
Millerburtonia Cif. 1951, anamorphic Plagiostoma Fuckel 1870
Phaeocytostroma Petr. 1921, anamorphic
Clypeoporthe Höhn 1919
Septogloeum Sacc. 1880, ?anamorphic
Pleuroceras Riess 1854
Uniseta Ciccar. 1948, anamorphic Plagiostoma Fuckel 1870
?Zythia Fr. 1849
Melanconidaceae G. Winter
Dumortieria Westend. 1857 (as Cytosporina Sacc. 1884), anamorphic Melanamphora Lafl. 1976 and Melogramma
Fr. 1849
Harknessia Cooke 1881, anamorphic Wuestneia Auersw. ex Fuckel 1863
Hendersonula Speg. 1880, anamorphic
Pseudovalsella Höhn. 1918 and Dictyoporthe Petr. 1955
Mastigosporella Höhn. 1914, anamorphic
Wuestneiopsis J. Reid & Dowsett 1990
Melanconiopsis Ellis & Everh. 1900,
anamorphic Massariovalsa Sacc. 1882
Melanconium Link 1809, anamorphic
Melanconis Tul. & C. Tul. 1863
Stilbospora Pers. 1794, ?anamorphic Prosthecium Fresen. 1852
Tubakia B. Sutton 1973, anamorphic
Dicarpella Syd. & P. Syd. 1921
Pseudoplagiostomaceae Cheewangkoon, M.J.
Wingf. & Crous
Cryptosporiopsis-like, anamorphic Pseudoplagiostoma Cheewangkoon, M.J.
Wingf. & Crous 2010*
Pseudovalsaceae M.E. Barr
Coryneum Nees 1816, anamorphic Pseudovalsa Ces. & De Not. 1863
Schizoparmeaceae Rossman
Coniella Höhn. 1918*
Pilidiella Petr. & Syd. 1927, anamorphic
Schizoparme Shear 1923*
Sydowiellaceae Lar. N. Vassiljeva
Cylindrosporella Höhn. 1916, anamorphic
Stegophora Syd. & P. Syd. 1916
Phoma-like, anamorphic Hapalocystis
Auersw. ex Fuckel 1863
Togniniaceae Réblová, L. Mostert, W. Gams
& Crous
Phaeoacremonium W. Gams, Crous & M.J.
Wingf. 1996, anamorphic Togninia Berl.
1900
Valsaceae Tul. & C. Tul.
Amphicytostroma Petr. 1921, anamorphic
Amphiporthe Petr. 1971
Cytospora Ehrenb. 1818, anamorphic
Leucostoma (Nitschke) Höhn. 1917*,
Valsa Fr. 1849*, Valsella Fuckel 1870
and Valseutypella Höhn. 1919
Harpostroma Höhn. 1928, anamorphic
Leptosillia Höhn. 1928
Lasmenia Speg. 1886, ?anamorphic
Chadefaudiomyces Kamat, Rao, Patil &
Ullasa 1974
Phomopsis (Sacc.) Bubák 1905, anamorphic Clypeoporthella Petr. 1924
Diaporthales, genera incertae sedis
?Greeneria Scribn. & Viala 1887
Apoharknessia Crous & S.J. Lee 2004
Apomelasmia Grove 1937
Asterosporium Kunze 1819*
Craspedodidymum Hol.-Jech. 1972, anamorphic Thailandiomyces Pinruan, Sakay., K.D. Hyde & E.B.G. Jones 2008
Cytospora-like, anamorphic Pachytrype
Berl. ex M.E. Barr, J.D. Rogers & Y.M.
Ju 1993
Dwiroopa Subram. & Muthumary
Hypodermina Höhn. 1916
Macrohilum H.J. Swart 1988
Natarajania Pratibha & Bhat 2006*
27
Phomopsis-like, anamorphic Caudospora
Starbäck 1889
Rabenhorstia Fr. 1849, anamorphic Hercospora Fr. 1825
Sirococcus Preuss 1855
Stenocarpella Syd. & P. Syd. 1917
Synchaetomella Decock & Seifert 2005
Magnaporthales Thongk., Vijakr. & K.D.
Hyde
Magnaporthaceae P.F. Cannon
Clasterosporium Schwein. 1832, anamorphic Clasterosphaeria Sivan. 1984
Harpophora W. Gams 2000, anamorphic
Gaeumannomyces Arx & D.L. Olivier
1952
Mycoleptodiscus Ostaz. 1968, anamorphic
Omnidemptus P.F. Cannon & Alcorn
1994
Pyricularia Sacc. 1880, anamorphic Magnaporthe R.A. Krause & R.K. Webster
1972
Ophiostomatales Benny & Kimbr.
Kathistaceae Malloch & M. Blackw.
Mattirolella S. Colla 1929
Termitaria Thaxt. 1920
Termitariopsis M. Blackw., Samson &
Kimbr. 1980
Ophiostomataceae Nannf.
Acremonium-like, anamorphic Fragosphaeria Shear 1923
Ambrosiella Brader ex Arx & Hennebert
1965, anamorphic Grosmannia Goid.
1936 and Ophiostoma Syd. & P. Syd.
1919*
Dryadomyces Gebhardt 2005, anamorphic
Grosmannia Goid. 1936*
Graphilbum H.P. Upadhyay & W.B.
Kendr. 1975, anamorphic Ophiostoma
Syd. & P. Syd. 1919
Graphium Corda 1837, anamorphic Ophiostoma Syd. & P. Syd. 1919*
Hyalobelemnospora Matsush. 1993, anamorphic Subbaromyces Hesselt. 1953
Hyalorhinocladiella H.P. Upadhyay &
W.B. Kendr. 1975, anamorphic Ceratocystiopsis H.P. Upadhyay & W.B.
Kendr. 1975, and Ophiostoma Syd. & P.
Syd. 1919*
28
Hyalorhinocladiella-like, anamorphic Klasterskya Petr. 1940
Leptographium Lagerb. & Melin 1927,
anamorphic Grosmannia Gold. 1936
and Ophiostoma Syd. & P. Syd. 1919*
Pesotum J.L. Crane & Schokn. 1973,
anamorphic Ophiostoma Syd. & P. Syd.
1919*
Pesotum-like, anamorphic Grosmannia
Gold. 1936
Phialographium H.P. Upadhyay & W.B.
Kendr. 1974, anamorphic Ophiostoma
Syd. & P. Syd. 1919
Raffaelea Arx & Hennebert 1965, anamorphic Grosmannia Goid. 1936*
Sporothrix Hektoen & C.F. Perkins 1901,
anamorphic Ophiostoma Syd. & P. Syd.
1919*
Sordariales Chad. ex D. Hawksw. & O.E.
Erikss.
Chaetomiaceae G. Winter
Botryotrichum Sacc. & Marchal 1885,
anamorphic Chaetomium Kunze 1817
Humicola Traaen 1914
Papulaspora Preuss 1851, anamorphic
Chaetomium Kunze 1817
Phialophora-like, anamorph of Chaetomidium (Zopf) Sacc. 1882
Taifanglania Z.Q. Liang, Y.F. Han, H.L.
Chu & R.T.V. Fox 2009
Trichocladium Harz 1871
Lasiosphaeriaceae Nannf.
Angulimaya Subram. & Lodha 1964,
anamorphic Bombardioidea C. Moreau
ex N. Lundqv. 1972
?Bagadiella Cheewangkoon & Crous
2010*
Cladorrhinum Sacc. & Marchal 1885,
anamorphic Apiosordaria Arx & W.
Gams 1967, Cercophora Fuckel 1870
and Podospora Ces. 1856*
Mammaria Ces. ex Rabenh. 1854,
anamorphic Pseudocercophora Subram.
& Sekar 1986
Phialophora-like, anamorphic Cercophora
Fuckel 1870, Eosphaeria Höhn. 1917,
Lasiosphaeria Ces. & De Not. 1863 and
Podospora Ces. 1856
Mycosphere
Sordariaceae G. Winter
Chrysonilia Arx 1981, anamorphic Neurospora Shear & B.O. Dodge 1927
?Virgariella-like, anamorphic Effetia Bartoli, Maggi & Persiani 1984
Sordariales, genera incertae sedis
Madurella Brumpt 1905
Ramophialophora M. Calduch, Stchigel,
Gené & Guarro 2004*
?Trichocladium-like, anamorphic Ascolacicola Ranghoo & K.D. Hyde 1998
Ypsilonia Lév. 1846, anamorphic Acanthotheciella Höhn. 1911
Sordariomycetidae, families incertae sedis
Amplistromataceae Huhndorf, A.N. Mill., M.
Greif & Samuels
Acrodontium-like, anamorphic Amplistroma Huhndorf, A.N. Mill., M. Greif &
Samuels 2009*
Pseudogliomastix W. Gams 1985, anamorphic Wallrothiella Sacc. 1882
Annulatascaceae S.W. Wong, K.D. Hyde &
E.B.G. Jones
Phaeoisaria-like, anamorphic Rhamphoria
Niessl 1876
Phialophora-like, anamorphic Teracosphaeria Réblová & Seifert 2007
Cephalothecaceae Höhn.
Acremonium-like, anamorphic Albertiniella Kirschst. 1936
Chalara-like, anamorphic Cryptendoxyla
Malloch & Cain 1970
Phialemonium W. Gams & McGinnis
1983
Helminthosphaeriaceae Samuels, Candoussau
& Magní
Diplococcium Grove 1885, anamorphic
Helminthosphaeria Fuckel 1870
Endophragmiella B. Sutton 1973, anamorphic Echinosphaeria A.N. Mill. &
Huhndorf 2004
Selenosporella-like, anamorphic Ruzenia
O. Hilber 2002
Spadicoides S. Hughes 1958, anamorphic
Tengiomyces Réblová 1999
Vermiculariopsiella Bender 1932, anamorphic Echinosphaeria A.N. Mill. &
Huhndorf 2004*
Sordariomycetidae, genera incertae sedis
Brachysporiella Bat. 1952, anamorphic
Ascotaiwania Sivan. & H.S. Chang
1992
Conioscypha Höhn. 1904, anamorphic
Conioscyphascus Réblová & Seifert
2004
Endophragmiella-like, anamorphic Phaeotrichosphaeria Sivan. 1983
Menisporopsis S. Hughes 1952, anamorphic Menisporopascus Matsush. 2003
Phialophora-like, anamorphic Linocarpon
Syd. & P. Syd. 1917
Pleurothecium Höhn. 1919, anamorphic
Carpoligna F.A. Fernández & Huhndorf 1999
Ramichloridium-like, anamorphic Barbatosphaeria Reblova 2008
Rhodoveronaea Arzanlou, W. Gams &
Crous 2007*
Spadicoides S. Hughes 1958*
Sporothrix-like, anamorphic Barbatosphaeria Reblova 2008
Subclass Xylariomycetidae sensu O.E. Erikss.
& Winka
Xylariales Nannf.
Amphisphaeriaceae G. Winter
Bartalinia Tassi 1900
?Beltraniella Subram. 1952, ?anamorphic
Leiosphaerella Höhn. 1919
Bleptosporium Steyaert 1961, anamorphic
Amphisphaeria Ces. & De Not. 1863
Ceratosporium Schwein. 1832, anamorphic Iodosphaeria Samuels, E. Müll. &
O. Petrini 1987
?Chaetoconis Clem. 1909, anamorphic
Ceriospora Niessl 1876
Discosia Lib. 1837
Doliomyces Steyaert 1961
?Hyalotiopsis Punith. 1970, anamorphic
Ellurema Nag Raj & W.B. Kendr. 1986
Labridella Brenckle 1929, anamorphic
Griphosphaerioma Höhn. 1918
Labridium Vestergr. 1897, anamorphic
Discostroma Clem. 1909
29
Microdochium Syd. 1924, anamorphic
Monographella Petr. 1924
Monochaetia (Sacc.) Allesch. 1902
Monochaetinula Muthumary, Abbas & B.
Sutton 1986
Monochaetiopsis L. Cai, Jeewon & K.D.
Hyde 2003, anamorphic Dyrithiopsis L.
Cai, R. Jeewon & K.D. Hyde 2003
Morinia Berl. & Bres. 1889
?Pestalotia De Not. 1841, anamorphic
Broomella Sacc. 1883
Pestalotiopsis Steyaert 1949, anamorphic
Neobroomella Petr. 1947 and Pestalosphaeria M.E. Barr 1975
Phlogicylindrium Crous, Summerb. &
Summerell 2006
Sarcostroma Cooke 1871, anamorphic
Griphosphaerioma Höhn. 1918
Seimatosporium Corda 1833, anamorphic
anamorphic Discostroma Clem. 1909
Seiridium Nees 1816, anamorphic Blogiascospora Shoemaker, E. Müll. &
Morgan-Jones 1966 and Lepteutypa
Petr. 1923
?Selenosporella G. Arnaud ex MacGarvie
1969, anamorphic Oxydothis Penz. &
Sacc. 1898
Selenosporella-like, anamorphic Iodosphaeria Samuels, E. Müll. & O. Petrini
1987
Sporocadus Corda 1839
Synnemapestaloides T. Handa & Y.
Harada 2004
?Truncatella Steyaert 1949 anamorphic
Broomella Sacc. 1883
Zetiasplozna Nag Raj 1993
Diatrypaceae Nitschke
Dumortieria Westend. 1857 (some as
Cytosporina Sacc. 1884), anamorphic
Cryptosphaeria Ces. & De Not. 1863,
Diatrype Fr. 1849, Eutypa Tul. & C.
Tul. 1863 and Eutypella (Nitschke)
Sacc. 1875
Libertella Desm. 1830, anamorphic
Diatrype Fr. 1849, Diatrypella (Ces. &
De. Not.) De. Not. 1863, Eutypa Tul. &
C. Tul. 1863 and Eutypella (Nitschke)
Sacc. 1875
Hyponectriaceae Petr.
Aureobasidium Viala & G. Boyer 1891,
Discosphaerina Höhn. 1917
30
Beltraniella Subram. 1952, anamorphic
Pseudomassaria Jacz. 1894
Hormonema-like, anamorphic Discosphaerina Hohn 1917
Kabatia Bubák 1904, anamorphic Discosphaerina Höhn. 1917
Kabatiella Bubák 1907, anamorphic Discosphaerina Höhn. 1917
Sarcophoma Höhn. 1916, anamorphic Discosphaerina Hohn 1917
Selenophoma Maire 1907, anamorphic
Discosphaerina Hohn 1917
Graphostromataceae M.E. Barr, J.D. Rogers &
Y.M. Ju
Nodulisporium-like, anamorphic Graphostroma Piroz. 1974
Xylariaceae Tul. & C. Tul.
Acanthodochium Samuels, J.D. Rogers &
Nagas., anamorphic Astrocystis Berk. &
Broome and Collodiscula I. Hino &
Katum. 1955
Arthroxylaria Seifert & W. Gams 2002
Dematophora R. Hartig 1883, anamorphic
Rosellinia De Not. 1844*
?Dicyma Boulanger 1897, anamorphic
Ascotricha Berk. 1838
Geniculisynnema Okane & Nakagiri 2007,
anamorphic Nemania Gray 1821
Geniculosporium Chesters & Greenh.
1964, anamorphic Entoleuca Syd. 1922,
Leprieuria Laessøe, J.D. Rogers &
Whalley 1989, Nemania Gray 1821,
Phylacia Lév. 1845, Podosordaria Ellis
& Holw. 1897, Rosellinia De Not. 1844
and Xylaria Hill ex Schrank 1789
Geniculosporium-like, anamorphic Halorosellinia Whalley, E.B.G. Jones, K.D.
Hyde & Læssøe 2000, Jumillera J.D.
Rogers, Y.M. Ju & San Martín 1997
and Kretzschmaria Fr. 1849
Hypocreodendron Henn. 1897, anamorphic Discoxylaria J.C. Lindq. & J.E.
Wright 1964
Libertella Desm. 1830
Libertella-like, anamorphic Barrmaelia
Rappaz 1995, Creosphaeria Theiss.
1910, Jumillera J.D. Rogers, Y.M. Ju &
San Martín 1997 and Lopadostoma
(Nitschke) Traverso 1906
Mycosphere
Lindquistia Subram. & Chandrash. 1977,
anamorphic Poronia Willd. 1787
Masoniomyces J.L. Crane & Dumont 1975,
anamorphic Camillea Fr. 1849
Mirandina-like, anamorphic Kretzschmariella Viégas 1944
Moelleroclavus Henn. 1902, anamorphic
Xylaria Hill ex Schrank 1789
Muscodor Worapong, Strobel & W.M.
Hess 2001
Nodulisporium Preuss 1849, anamorphic
Annulohypoxylon Y.M. Ju, J.D. Rogers
& H.M. Hsieh 2005, Biscogniauxia
Kuntze 1891, Daldinia Ces. & De Not.
1863, Entonaema Möller 1901, Hypoxylon Bull. 1791*, Induratia Samuels, E.
Müll. & O. Petrini, Pyrenomyxa Morgan 1895 (as Pulveria Malloch &
Rogerson 1977), Thamnomyces Ehrenb.
1820, Theissenia Maubl. 1914, Thuemenella Penz. & Sacc. 1898 and
Xylaria Hill ex Schrank
Nodulosporium-like, anamorphic Rhopalostroma D. Hawksw. 1977 and Ruwenzoria J. Fornier, M. Stadler, Læssøe &
C. Decock 2010*
Padixonia Subram. 1972, anamorphic
Xylaria Hill ex Schrank
Rhinocladiella-like, anamorphic Obolarina Pouzar 1986
Sporothrix-like, anamorphic Rostrohypoxylon J. Fourn. & M. Stadler 2010* and
Ruwenzoria J. Fornier, M. Stadler,
Læssøe & C. Decock 2010*
Triplicaria P. Karst. 1889, anamorphic
Hypoxylon Bull. 1791
Virgaria Nees 1816, anamorphic Ascovirgaria J.D. Rogers & Y.M. Ju 2002
Virgariella-like, anamorphic Hypoxylon
Bull. 1791*
Vivantia J.D. Rogers, Y.M. Ju & Cand.
1996
Xylocladium P. Syd. ex Lindau 1900,
anamorphic Camillea Fr. 1849
Xylocoremium J.D. Rogers 1984, anamorphic Xylaria Hill ex Schrank 1789
Xylariales, genera incertae sedis
Basifimbria Subram. & Lodha 1968
Chaetochalara B. Sutton & Piroz. 1965
Chalara (Corda) Rabenh. 1844*
Hadrotrichum Fuckel 1865
Humicola-like, anamorphic Ascotrichella
Valldos. & Guarro 1988
Melanographium Sacc. 1913, possibly
anamorphic Lasiobertia Sivan. 1978
Nodulisporium-like, anamorphic Coniolariella D. Garcıá, Stchigel & Guarro
Rhinocladiella-like, anamorphic Coniolariella D. Garcıá, Stchigel & Guarro
Selenosporella-like, anamorphic Diatrypasimilis J. Zhou, & Kohlm. 2010*
Sordariomycetes, orders incertae sedis
Lulworthiales Kohlm., Spatafora & Volkm.Kohlm.
Lulworthiaceae Kohlm., Spatafora & Volkm.Kohlm.
Anguillospora Ingold 1942, anamorphic
Lindra I.M. Wilson 1956
Cumulospora I. Schmidt 1985, anamorphic
Lulwoana Kohlm., Volkm.-Kohlm., J.
Campb., Spatafora & Gräfenhan 2005
and Lulworthia G.K. Sutherl. 1916*
Halazoon Abdel-Aziz, Abdel-Wahab &
Nagahama 2010, and Lulworthia G.K.
Sutherl. 1916*
Hydea K.L. Pang & E.B.G. Jones 2010*
Matsusporium E.B.G. Jones & K.L. Pang
2010*
Moleospora Abdel-Wahab, Abdel-Aziz &
Nagahama 2010*
Moromyces Abdel-Wahab, K.L. Pang,
Nagahama, Abdel-Aziz & E.B.G. Jones
2010
Zalerion R.T. Moore & Meyers 1962,
anamorphic Lulwoana Kohlm., Volkm.Kohlm., J. Campb., Spatafora &
Gräfenhan 2005*
Phyllachorales M.E. Barr
Phyllachoraceae Theiss. & H. Syd.
Acerviclypeatus Hanlin 1990, anamorphic
Ophiodothella Boyd 1934
Baeumleria Petr. & Syd. 1927, anamorphic
Phyllachora Nitschke ex Fuckel 1870
(as Trabutia Sacc. & Roum. 1881)
Colletogloeum-like, anamorphic Coccodiella Hara 1909
Cyclodomus Höhn. 1909, anamorphic
Maculatifrons K.D. Hyde 1996
Diachorella Höhn. 1918, anamorphic
Diachora Müll. 1893
31
Hysterodiscula Petr. 1942, anamorphic
Orphnodactylis Malloch & A. Mallik
1998
Linochora Höhn. 1910, anamorphic Phyllachora Nitschke ex Fuckel 1870
Mycohypallage B. Sutton 1963, anamorphic Deshpandiella Kamat & Ullasa
1973
Oswaldina Rangel 1921, anamorphic
Apiosphaeria Höhn. 1909
Polystigmina Sacc. 1884, anamorphic
Polystigma DC. 1815
Pseudothiopsella Petr. 1928, anamorphic
Pseudothiella Petr. 1928
Rhodosticta Woron. 1911, anamorphic
Polystigma DC. 1815 and Stigmatula
(Sacc.) Syd. & P. Syd. 1901
Trichosphaeriales M.E. Barr 1983
Trichosphaeriaceae G. Winter 1885
Brachysporium Sacc. 1886, anamorphic
Cryptadelphia Réblová & Seifert 2004
Koorchaloma Subram. 1953, anamorphic
Kananascus Nag Raj 1984
Stanjehughesia Subram. 1992, anamorphic
Miyoshiella Kawam. 1929 and Umbrinosphaeria Réblová 1999
Stromatographium Höhn. 1907, anamorphic Fluviostroma Samuels & E. Müll.
1980
Sordariomycetes, family incertae sedis
Apiosporaceae K.D. Hyde, J. Fröhlich, Joanne
E. Taylor & M.E. Barr
Arthrinium Kunze 1817, anamorphic Apiospora Sacc. 1875
Cordella Speg. 1886, anamorphic Apiospora Sacc. 1875
Pteroconium Sacc. ex Grove 1914, anamorphic Apiospora Sacc. 1875
Scyphospora L.A. Kantsch. 1928, anamorphic Apiospora Sacc. 1875
Didymostilbe-like, anamorphic Ornatispora K.D. Hyde, Goh, Joanne E. Taylor, J.
Fröhl. 1999
Dinemasporium Lév. 1846, anamorphic
Phomatospora Sacc. 1875
Ellisembia Subram. 1992
Myrmecridium Arzanlou, W. Gams &
Crous 2007
Nigrospora Zimm. 1902, anamorphic
Khuskia H.J. Huds. 1963
Penicillifer Emden 1968, anamorphic
Stellosetifera Matsush.1996
Phomatosporella Tak. Kobay. & K. Sasaki
1982, anamorphic Phomatospora Sacc.
1875
Ramichloridium-like, anamorphic Tectonidula Réblová 2009*
Selenosporella G. Arnaud ex MacGarvie
1969
Sporothrix-like, anamorphic Phomatospora Sacc. 1875 and Tectonidula
Réblová 2009*
ASCOMYCOTA, families incertae sedis
Families and genera that cannot be
placed in any of the classes and orders accepted
in the present classification with a high degree
of probability are listed below.
Amorphothecaceae Parbery
Hormoconis Arx & G.A. de Vries 1973,
anamorphic Amorphotheca Parbery
1969
Batistiaceae Samuels & K.F. Rodrigues
Acrostroma Seifert 1987, anamorphic
Batistia Cif. 1958
Pseudeurotiaceae Malloch & Cain
Teberdinia Sogonov, W. Gams, Summerb.
& Schroers 2005, anamorphic Pseudeurotium J.F.H. Beyma 1937
Thyridiaceae O.E. Erikss. & J.Z. Yue
Pleurocytospora Petr. 1923, anamorphic
Thyridium Nitschke 1867
Seuratiaceae Vuill. ex M.E. Barr
Atichia Flot. 1850, anamorphic Seuratia
Pat. 1904
Sordariomycetes, genera incertae sedis
Cryptomycella Höhn. 1925, anamorphic
Cryptomycina Höhn. 1977
?Custingophora Stolk, Hennebert & Klopotek 1968*
ASCOMYCOTA, genera incertae sedis
Abgliophragma R.Y. Roy & Gujarati 1966
Abropelta B. Sutton 1986
Acanthoderma Syd. & P. Syd. 1917
Acarellina Bat. & H. Maia 1960
32
Mycosphere
Acaroconium Kocourk. & D. Hawksw.
2008
Acarocybellina Subram. 1992
Acarocybiopsis J. Mena, A. Hern. Gut. &
Mercado 1999
Acaropeltis Petr. 1937
Achoropeltis Syd. 1929
Aciculariella G. Arnaud 1954
Acinula Fr. 1822
Acontium Morgan 1902
Acremoniula G. Arnaud 1954
Acrocybe Syd. 1937
Acroconidiellina M.B. Ellis 1971
Acrocylindrium Bonord 1851
Acrodictyella W.A. Baker & Partr. 2001
Acrodictyopsis P.M. Kirk 1983
Acrodictys M.B. Ellis 1961
Acrodontiella U. Braun & Scheuer 1995
Acrodontium de Hoog 1972
Acrophialophora Edward 1961
Acrophragmis Kiffer & Reisinger 1970
Acrospeira Berk. & Broome 1857
Acrospira Mont. 1857
Acrosporium Bonord. 1851
Acrostaurus Deighton & Piroz. 1972
Acrotheciella Koord. 1907
Acrothecium (Corda) Preuss 1851
Actinochaete Ferro 1907
Actinocladium Ehrenb. 1819
Actinodochium Syd. 1927
Actinonema Pers. 1822
Actinotexis Arx 1960
Actinothecium Ces. 1854
Actinothyrium Kunze 1823
Acumispora Matsush. 1980
Aegeritella Bałazy & J. Wiśn. 1974
Aenigmatospora R.F. Castañeda, Saikawa,
Guarro & Calduch 1999
Agaricodochium X.J. Liu 1981
Agarwalia D.P. Tiwari & P.D. Agrawal
1974
Agarwalomyces R.K. Verma & Kamal
1987
Agrabeeja Subram. 1995
Agyriella Sacc. 1884
Agyriellopsis Höhn. 1903
Ahmadia Syd. 1939
Ajrekarella Kamat & Kalani 1964
Alatosessilispora K. Ando & Tubaki 1984
Albophoma Tak. Kobay., Masuma, Omura
& Kyoto Watan. 1994
Alciphila Harmaja 2002
Aleurodomyces Buchner 1912
Algonquinia R.F. Castañeda & W.B.
Kendr. 1991
Allonema Syd. 1934
Alloneottiosporina Nag Raj 1993
Allothyriella Bat., Cif. & Nascim. 1959
Allothyrina Bat. & J.L. Bezerra 1964
Allothyriopsis Bat., Cif. & H. Maia 1959
Alpakesa Subram. & K. Ramakr. 1954
Alpakesiopsis Abbas, B. Sutton, Ghaffar &
A. Abbas 2003
Alveophoma Alcalde 1952
Alysidiella Crous 2006
Alysidiopsis B. Sutton 1973
Amallospora Penz. 1897
Amblyosporium Fresen. 1863
Amerodiscosiella M.L. Farr 1961
Amerodiscosiellina Bat. & Cavalc. 1966
Amerosporiopsis Petr. 1941
Amerosympodula Matsush. 1996
Amoenodochium Peláez & R.F. Castañeda
1996
Amoenomyces R.F. Castañeda, Saikawa &
Hennebert 1996
Amphichaetella Höhn. 1916
Amphophialis R.F. Castañeda, W.B. Kendr.
& Guarro 1998
Amphoropycnium Bat. 1963
Ampullicephala R.F. Castañeda, Minter &
M. Stadler 2009*
Ampullifera Deighton 1960
Ampulliferina B. Sutton 1969
Ampulliferinites Kalgutkar & Sigler 1995,
fossil fungi
Anaphysmene Bubák 1906
Anarhyma M.H. Pei & Z.W. Yuan 1986
Anaselenosporella Heredia, R.F. Castañeda & R.M. Arias 2010*
Ancoraspora Mig. Rodr. 1982
Ancorasporella J. Mena, Mercado &
Heredia 1998
Angiopomopsis Höhn. 1912
Angulospora Sv. Nilsson 1962
Annellodentimyces Matsush. 1985
Annellodochium Deighton 1969
Annellolacinia B. Sutton 1964
Annellophora S. Hughes 1952
Annellophorella Subram. 1962
Annellophragmia Subram. 1963
Annellosympodia McTaggart, R.G. Shivas
& U. Braun 2007
Antennopsis R. Heim 1952
33
Anthina Fr. 1832
Anthopsis Fil. March., A. Fontana & Luppi
Mosca 1977
Anthracoderma Speg. 1888
Antromyces Fresen. 1850
Anulohypha Cif. 1962
Aoria Cif. 1962
Aphanofalx B. Sutton 1986
Apiocarpella Syd. & P. Syd. 1919
Apogloeum Petr. 1954
Aporellula B. Sutton 1986
Aposporella Thaxt. 1920
Arachnophora Hennebert 1963
Arachnospora R.F. Castañeda, Minter &
Camino 2003
Arborillus Munt.-Cvetk. & Gómez-Bolea
1998
Arborispora K. Ando 1986
Arbusculina Marvanová & Descals 1987
Arcuadendron Sigler & J.W. Carmich.
1976
Argopericonia B. Sutton & Pascoe 1987
Aristastoma Tehon 1933
Arnaudina Trotter 1931
Arthrobotryum Ces. 1854
Arthrocladium Papendorf 1969
Arthrocristula Sigler, M.T. Dunn & J.W.
Carmich. 1982
Arthrosporium Sacc. 1880
Arthrowallemia R.F. Castañeda, Dania
García & Guarro 1998
Articularia Höhn. 1909
Articulophora C.J.K. Wang & B. Sutton
1982
Artocarpomyces Subram. 1996
Arxiella Papendorf 1967
Asbolisiomyces Bat. & H. Maia 1961
Ascochytopsis Henn. 1905
Ascochytulina Petr. 1922
Ascospora Mont. 1849
Ashtaangam Subram. 1995
Aspilaima Bat. & H. Maia 1961
Astelechia Cif. 1962
Asterinothyriella Bat. & Cif. 1959
Asterinothyrium Bat., Cif. & H. Maia 1959
Asteroconium Syd. & P. Syd. 1903
Asterostomopora Bat. & H. Maia 1960
Asterostomopsis Bat., Cif. & H. Maia 1959
Asterostomulina Bat., J.L. Bezerra & H.
Maia 1964
Astrodochium Ellis & Everh. 1897
Astronatelia Bat. & H. Maia 1962
34
Atractilina Dearn. & Barthol. 1924
Atractium Link 1809
Atrosetaphiale Matsush. 1995
Aurantiosacculus Dyko & B. Sutton 1979
Aurosphaeria Sun J. Lee, Strobel, Eisenman, Geary, Vargas & S.A. Strobel
2009
Avesicladiella W.P. Wu, B. Sutton &
Gange 1997
Avettaea Petr. & Syd. 1927
Bachmanniomyces D. Hawksw. 1981
Bacillopeltis Bat. 1957
Bactridium Kunze 1817
Bactrodesmiella M.B. Ellis 1959
Badarisama Kunwar, J.B. Manandhar &
J.B. Sinclair 1986
Bahuchashaka Subram. 1978
Bahukalasa Subram. & Chandrash. 1979
Bahusandhika Subram. 1956
Balaniopsis P.M. Kirk 1985
Balanium Wallr. 1833
Barbarosporina Ķirulis 1942
Barnettella D. Rao & P.Rag. Rao 1964
Basauxia Subram. 1995
Basididyma Cif. 1962
Basramyces Abdullah, Abdulk. & Goos
1989
Batistina Peres 1961
Beauveriphora Matsush 1975
Beccopycnidium F. Stevens 1930
Beejadwaya Subram. 1978
Belemnospora P.M. Kirk 1981
Bellulicauda B. Sutton 1967
Beltrania Penz. 1882
Beltraniopsis Bat. & J.L. Bezerra 1960
Beniowskia Racib. 1900
Benjaminia S. Ahmad 1967
Benjpalia Subram. & Bhat 1989
Bharatheeya D'Souza & Bhat 2002
Bhargavaella Sarj. Singh & K.S. Srivast.
1980
Bhatia W.A. Baker & Morgan-Jones
2009*
Bibanasiella R.F. Castañeda & W.B.
Kendr. 1991
Biflagellospora Matsush. 1975
Biflagellosporella Matsush. 1993
Bilboque Viégas 1960
Bilgramia Panwar, Purohit & Chouhan
1974
Bimeris Petr. 1949
Bioconiosporium Bat. & J.L. Bezerra 1964
Mycosphere
Biophomopsis Petr. 1931
Bispora Corda 1837
Bisseomyces R.F. Castañeda 1985
Bizzozeriella Speg. 1888
Blarneya D. Hawksw., Coppins & P.
James 1980
Blastacervulus H.J. Swart 1988
Blastocatena Subram. & Bhat 1989
Blastoconium Cif. 1931
Blastodictys M.B. Ellis 1976
Blastofusarioides Matsush. 1996
Blastophorella Boedijn 1937
Blastophorum Matsush. 1971
Blastophragma Subram. 1995
Blennoriopsis Petr. 1919
Blodgettia Harv. 1858
Bomplandiella Speg. 1886
Bostrichonema Ces. 1867
Botryodeorsum T.P. Devi, N. Mathur,
Chowdhry, Jasvir Singh & O. Prakash
2006 Botryoderma Papendorf & H.P.
Upadhyay 1969
Botryodiplodina Dias & Sousa da Câmara
1954
Botryomonilia Goos & Piroz. 1975
Botryomyces Greco 1916
Botryosporium Corda 1831
Brachydesmiella G. Arnaud ex S. Hughes
1961
Brachyhelicoon G. Arnaud 1952
Brachysporiellina Subram. & Bhat 1989
Brachysporiopsis Yanna, W.H. Ho & K.D.
Hyde 2004
Brefeldiopycnis Petr. & Cif. 1932
Brevicatenospora R.F. Castañeda, Minter
& Saikawa 2006
Briosia Cavara 1888
Broomeola Kuntze 1891
Brycekendrickia Nag Raj 1973
Bulbilopycnis Matsush. 1996
Bulbocatenospora R.F. Castañeda & Iturr.
2000
Bullaserpens Bat., J.L. Bezerra & Cavalc.
1965
Cacumisporium Preuss 1851
Calcarispora Marvanová & Marvan 1963
Calcarispora Marvanová & Marvan 1963
Calcarisporiella de Hoog 1974
Callistospora Petr. 1955
Calocline Syd. 1939
Camarographium Bubák 1916
Camaropycnis E.K. Cash 1945
Camarosporellum Tassi 1902
Camarosporiopsis Abbas, B. Sutton &
Ghaffar 2000
Camarosporulum Tassi 1902
Camposporidium Nawawi & Kuthub. 1988
Camposporium Harkn. 1884
Camptomeris Syd. 1927
Campylospora Ranzoni 1953
Candelabrum Beverw. 1951
Candelosynnema K.D. Hyde & Seifert
1992
Capillaria Pers. 1822
Capitorostrum Bat. 1957
Capsicumyces Gamundí, Aramb. &
Giaiotti 1979
Carmichaelia N.D. Sharma 1980
Carrismyces R.F. Castañeda & Heredia
2000
Castanedaea W.A. Baker & Partr. 2001
Catenella Bat. & Peres 1963
Catenocuneiphora Matsush. 2003
Catenophora Luttr. 1940
Catenophoropsis Nag Raj & W.B. Kendr.
1988
Catenospegazzinia Subram. 1991
Catenosubulispora Matsush. 1971
Catenosynnema Kodsueb, K.D. Hyde &
W.H. Ho 2007
Catenulaster Bat. & C.A.A. Costa 1959
Catinopeltis Bat. & C.A.A. Costa 1957
Catinula Lév. 1848
Ceeveesubramaniomyces Pratibha, K.D.
Hyde & Bhat 2005
Cephalodochium Bonord. 1851
Ceracea Cragin 1885
Ceratocladium Corda 1839
Ceratophorum Sacc. 1880
Ceratopycnis Höhn. 1915
Ceratosporella Höhn. 1923
Cercosperma G. Arnaud ex B. Sutton &
Hodges 1983
Ceuthosira Petr. 1924
Ceuthosporella Petr. & Syd. 1923
Chaetantromycopsis H.P.
Upadhyay,
Cavalc. & A.A. Silva 1986
Chaetendophragmia Matsush. 1971
Chaetendophragmiopsis B. Sutton &
Hodges 1978
Chaetoblastophorum Morgan-Jones 1977
Chaetoconidium Zukal 1887
Chaetocytostroma Petr. 1920
Chaetodiplis Clem. 1931
35
Chaetodiplodia P. Karst. 1884
Chaetodiplodina Speg. 1910
Chaetopeltaster Katum. 1975
Chaetophiophoma Speg. 1910
Chaetophoma Cooke 1878
Chaetopsis Grev. 1825
Chaetopyrena Pass. 1881
Chaetoseptoria Tehon 1937
Chaetostromella P. Karst. 1895
Chalarodendron C.J.K. Wang & B. Sutton
1984
Chalarodes McKenzie 1991
Chantransiopsis Thaxt. 1914
Characonidia Bat. & Cavalc. 1965
Chardonia Cif. 1930
Charomyces Seifert 1987
Chasakopama Manohar., Bagyan., N.K.
Rao & Kunwar 2009*
Cheilaria Lib. 1830
Cheiroidea W.A. Baker & Morgan-Jones
2009*
Cheiromyceopsis Mercado & J. Mena
1988
Cheiromyces Berk. & M.A. Curtis 1857
Cheiromycina B. Sutton 1986
Cheiropolyschema Matsush. 1980
Cheirospora Moug. & Fr. 1825
Chiastospora Riess 1852
Chikaneea B. Sutton 1973
Chionomyces Deighton & Piroz. 1972
Chithramia Nag Raj 1988
Chlamydomyces Bainier 1907
Chlamydopsis Hol.-Jech. & R.F. Castañeda 1986
Chlamydorubra K.B. Deshp. & K.S.
Deshp. 1966
Choanatiara DiCosmo 1984
Choreospora Constant. & R. Sant. 1987
Chromosporium Corda 1829
Chrysachne Cif. 1938
Chrysalidopsis Steyaert 1961
Chuppia Deighton 1965
Cicadocola Brain 1923
Ciferria Gonz. Frag. 1925
Ciferriella Petr. 1930
Ciferrina Petr. 1932
Ciferriopeltis Bat. & H. Maia 1965
Ciliochora Höhn. 1919
Ciliochorella Syd. 1935
Ciliophora Petr. 1929
Ciliophorella Petr. 1940
Ciliosporella Petr. 1927
36
Circinoconis Boedijn 1942
Circinotrichum Nees 1816
Cirrosporium S. Hughes 1980
Cissococcomyces Brain 1923
Civisubramaniania Vittal & Dorai 1986
Cladochasiella Marvanová 1997
Cladoconidium Bandoni & Tubaki 1985
Cladographium Peyronel 1918
Cladoniicola Diederich, van den Boom &
Aptroot 2001
Cladosporiella Deighton 1965
Clasteropycnis Bat. & Cavalc. 1963
Clathroconium Samson & H.C. Evans
1982
Clauzadeomyces Diederich 1994
Clavariana Nawawi 1976
Cleistocystis Sousa da Câmara 1931
Cleistophoma Petr. & Syd. 1927
Clithramia Nag Raj 1988
Clohesyomyces K.D. Hyde 1993
Clypeochorella Petr. 1923
Clypeopatella Petr. 1942
Clypeophialophora Bat. & Peres 1962
Clypeopycnis Petr. 1925
Clypeoseptoria F. Stevens & P.A. Young
1925
Clypeostagonospora Punith. 1981
Coccogloeum Petr. 1955
Codonmyces Calat. & Etayo 1999
Coelographium (Sacc.) Gäum. 1920
Coelosporium Link 1824
Colemaniella Agnihothr. 1974
Coleodictyospora Charles 1929
Coleophoma Höhn. 1907
Coleoseptoria Petr. 1940
Colispora Marvanová 1988
Collecephalus J.A. Spencer 1972
Collembolispora Marvanová & Pascoal
2003
Colletoconis de Hoog & Aa 1978
Colletosporium Link 1824
Collodochium Höhn. 1902
Collostroma Petr. 1947
Columnodomus Petr. 1941
Columnophora Bubák & Vleugel 1916
Columnothyrium Bubák 1916
Comatospora Piroz. & Shoemaker 1971
Comocephalum Syd. 1939
Condylospora Nawawi 1976
Coniambigua Etayo & Diederich 1995
Conicomyces R.C. Sinclair, Eicker &
Morgan-Jones 1983
Mycosphere
Conidiocarpus Woron. 1927
Conioscyphopsis Goh & K.D. Hyde 1998
Coniothyrina Syd. 1912
Coniozyma Crous 2008
Conjunctospora Udagawa & Uchiy. 1999
Conostoma Bat. & J.L. Bezerra 1965
Consetiella Hol.-Jech. & Mercado 1982
Cooksonomyces H.J. Swart & D.A.
Griffiths 1974
Corallinopsis Lagarde 1917
Corallomyces Fr. 1849
Cordana Preuss 1851
Coremiella Bubák & K. Krieg. 1912
Corethropsis Corda 1839
Cornucopiella Höhn. 1915
Cornutispora Piroz. 1973
Cornutostilbe Seifert 1990
Corynecercospora V.K. Pal, Akhtar, N.
Ahmad, Kamal & D.K. Agarwal 2006
Corynesporella Munjal & H.S. Gill 1961
Corynesporina Subram. 2009*
Corynesporopsis P.M. Kirk 1981
Costanetoa Bat. & J.L. Bezerra 1963
Craneomyces Morgan-Jones, R.C. Sinclair
& Eicker 1987
Cremasteria Meyers & R.T. Moore 1960
Creodiplodina Petr. 1957
Creonecte Petr. 1949
Creoseptoria Petr. 1937
Creothyriella Bat. & C.A.A. Costa 1957
Cribropeltis Tehon 1933
Crinitospora B. Sutton & Alcorn 1985
Cristula Chenant. 1920
Crucellisporiopsis Nag Raj 1983
Crustodiplodina Punith. 1988
Cryptoceuthospora Petr. 1921
Cryptocoryneopsis B. Sutton 1980
Cryptocoryneum Fuckel 1870
Cryptophiale Piroz. 1968
Cryptophialoidea Kuthub. & Nawawi
1987
Cryptosporium Kunze 1817
Cryptostroma P.H. Greg. & S. Waller
1952
Cryptumbellata Udagawa & Uchiy. 1999
Ctenosporium R. Kirschner 2006
Cubasina R.F. Castañeda 1986
Culicidospora R.H. Petersen 1960
Culicinomyces Couch, Romney & B. Rao
1974
Curculiospora G. Arnaud 1954
Curucispora Matsush. 1981
Curvidigitus Sawada 1943
Curvulariopsis M.B. Ellis 1961
Cyanopatella Petr. 1949
Cyclomarsonina Petr. 1965
Cylindrocarpostylus R. Kirschner &
Oberw. 1999
Cylindrogloeum Petr. 1941
Cylindromyces Manohar., D.K. Agarwal &
N.K. Rao 2004
Cylindrothyrium Maire 1907
Cylindroxyphium Bat. & Cif. 1963
Cymbothyrium Petr. 1947
Cyrenella Goch. 1981
Cystidiella Malan 1943
Cystotrichiopsis B.A. Abbas, B. Sutton &
Ghaffar 2001
Cytodiscula Petr. 1931
Cytogloeum Petr. 1925
Cytonaema Höhn. 1914
Cytoplacosphaeria Petr. 1920
Cytopleastrum Abbas, B. Sutton, Ghaffar
& A. Abbas 2004
Cytosphaera Died. 1916
Cytosporella Sacc. 1884
Cytostagonospora Bubák 1916
Dacrymycella Bizz. 1885
Dactylifera Alcorn 1987
Dactylosporium Harz 1871
Dasysticta Speg. 1912
Davisiella Petr. 1924
Dearnessia Bubák 1916
Deccanodia Singhai 1974
Deichmannia Alstrup & D. Hawksw. 1990
Delortia Pat. & Gaillard 1888
Dendroclathra Voglmayr & G. Delgado
2001
Dendrodomus Bubák 1915
Dendrographiella Agnihothr. 1972
Dendrographium Massee 1892
Dendroseptoria Alcalde 1948
Dendrospora Ingold 1943
Dendrosporium Plakidas & Edgerton ex
J.L. Crane 1972
Dendryphiosphaera Lunghini & Rambelli
1978
Dennisographium Rifai 1977
Denticularia Deighton 1972
Dentocircinomyces R.F. Castañeda & W.B.
Kendr. 1990
Derexia Naras. 1970
Descalsia A. Roldán & Honrubia 1989
Desertella Mouch. 1979
37
Desmidiospora Thaxt. 1891
Dexhowardia J.J. Taylor 1970
Diademospora B.E. Söderstr. & Bååth
1979
Dialaceniopsis Bat. 1959
Diaphanium Fr. 1836
Diarimella B. Sutton 1980
Dicellispora Sawada 1944
Dichelostroma Bat. & Peres 1963
Dicholobodigitus G.P. White & Illman
1988
Dichotomophthora Mehrl. & Fitzp. ex P.N.
Rao 1966
Dichotomophthoropsis M.B. Ellis 1971
Dictyoarthrinium S. Hughes 1952
Dictyoarthrinopsis Bat. & Cif. 1958
Dictyocatenulata Finley & E.F. Morris
1967
Dictyodesmium S. Hughes 1951
Dictyophrynella Bat. & Cavalc. 1964
Dictyopolyschema M.B. Ellis 1976
Dictyorostrella U. Braun 1999
Dictyospiropes M.B. Ellis 1976
Didymobotryum Sacc. 1886
Didymochaeta Sacc. & Ellis 1898
Didymochaetina Bat. & J.L. Bezerra 1965
Didymopsis Sacc. & Marchal 1885
Didymosporina Höhn. 1916
Didymothozetia Rangel 1915
Didymotrichella G. Arnaud 1954
Diedickea Syd. & P. Syd. 1913
Digitodochium Tubaki & Kubono 1989
Digitomyces Mercado, M. Calduch &
Gené 2003
Digitopodium U. Braun, Heuchert & K.
Schub. 2005
Digitoramispora R.F. Castañeda & W.B.
Kendr. 1990
Digitosarcinella S. Hughes 1984
Dimastigosporium Faurel & Schotter 1965
Diplocladiella G. Arnaud ex M.B. Ellis
1976
Diplodinis Clem. 1931
Diplodinula Tassi 1902
Diplolaeviopsis Giralt & D. Hawksw.
1991
Diploöspora Grove 1916
Diploplenodomus Died. 1912
Diplozythiella Died. 1916
Discocolla Prill. & Delacr. 1894
Discogloeum Petr. 1923
Discomycetoidea Matsush. 1993
38
Discosiellina Subram. & K.R.C. Reddy
1972
Discosporina Höhn. 1927
Discotheciella Syd. & P. Syd 1917
Discozythia Petr. 1922
Dissoacremoniella Kiril. 1970
Ditangifibula G.C. Adams 1996
Divinia Cif. 1955
Domingoella Petr. & Cif. 1932
Dothideodiplodia Murashk. 1927
Dothioropsis Riedl 1974
Drechsleromyces Subram. 1978
Drepanospora Berk. & M.A. Curtis 1875
Drudeola Kuntze 1891
Drumopama Subram. 1957
Dryosphaera Jørg. Koch & E.B.G. Jones
1989
Dualomyces Matsush. 1987
Dwayabeeja Subram. 1958
Dwayaloma Subram. 1957
Dwayalomella Brisson, Piroz. & Pauzé
1975
Dwibahubeeja N. Srivast., A.K. Srivast. &
Kamal 1995
Dwibeeja Subram. 1995
Dwiroopella Subram. & Muthumary 1986
Echinocatena R. Campb. & B. Sutton
1977
Echinochondrium Samson & Aa 1975
Echinoconidiophorum Pereira-Carv. &
Dianese 2009*
Elachopeltella Bat. & Cavalc. 1964
Elattopycnis Bat. & Cavalc. 1964
Elegantimyces Goh, K.M. Tsui & K.D.
Hyde 1998
Eleutheromycella Höhn. 1908
Ellisiella Bat. 1956
Elotespora R.F. Castañeda & Heredia
2010*
Enantioptera Descals 1983
Endobotrya Berk. & M.A. Curtis 1874
Endobotryella Höhn. 1909
Endocalyx Berk. & Broome 1876
Endoconospora Gjaerum 1971
Endocoryneum Petr. 1922
Endogenospora R.F. Castañeda, O.
Morillo & Minter 2010*
Endophragmiopsis M.B. Ellis 1966
Endoplacodium Petr. 1949
Endoramularia Petr. 1923
Endosporoideus W.H. Ho, Yanna, K.D.
Hyde & Goh 2005
Mycosphere
Endostilbum Malençon 1964
Endozythia Petr. 1959
Enerthidium Syd. 1939
Engelhardtiella A. Funk 1973
Enridescalsia R.F. Castañeda & Guarro
1998
Enthallopycnidium F. Stevens 1925
Entoderma Hanula, Andreadis & M.
Blackw. 1991
Epaphroconidia Calat. & V. Atienza 1995
Ephelidium C.W. Dodge & E.D. Rudolph
1955
Epicladonia D. Hawksw. 1981
Epiclinium Fr. 1849
Epicoccospora Budathoki & S.K. Singh
1995
Epidermidophyton E. Lang 1879
Epidochiopsis P. Karst. 1892
Episporogoniella U. Braun 1994
Epistigme Syd. 1924
Erysiphopsis Speg. 1910
Erythrogloeum Petr. 1953
Esdipatilia Phadke 1981
Esteya J.Y. Liou, J.Y. Shih & Tzean 1999
Eurotiopsis P. Karst. 1889
Evanidomus Caball. 1941
Everhartia Sacc. & Ellis 1882
Everniicola D. Hawksw. 1982
Eversia J.L. Crane & Schokn. 1977
Evlachovaea Borisov & Tarasov 1999
Excipularia Sacc. 1884
Excipulariopsis P.M. Kirk & Spooner
1982
Exochalara W. Gams & Hol.-Jech. 1976
Exophoma Weedon 1926
Exosporella Höhn. 1912
Exosporiella P. Karst. 1892
Exosporium Link 1809
Exserticlava S. Hughes 1978
Favostroma B. Sutton & E.M. Davison
1983
Fecundostilbum Prameela & Chowdhry
2009*
Feltgeniomyces Dieder. 1990
Fenestroconidia Calat. & Etayo 1999
Filosporella Nawawi 1976
Fissuricella Pore, D'Amatao & Ajello
1977
Flabellocladia Nawawi 1985
Flabellospora Alas. 1968
Flagellospora Ingold 1942
Flahaultiella Seifert 2009*
Flosculomyces B. Sutton 1978
Frigidispora K.D. Hyde & Goh 1999
Fujimyces Minter & Caine 1980
Fuligomyces Morgan-Jones & Kamal 1984
Funicularius K.K. Baker & Zaim 1979
Furcaspora Bonar 1965
Fusamen (Sacc.) P. Karst. 1890
Fusariella Sacc. 1884
Fusariopsis Horta 1919
Fuscophialis B. Sutton 1977
Fusella Sacc. 1886
Fusicatena K. Matsush. & Matsush. 1996
Fusichalara S. Hughes & Nag Raj 1973
Fusicolla Bonord. 1851
Fusisporella Speg. 1910
Fusticeps J. Webster & R.A. Davey 1980
Gamosporella Speg. 1888
Gampsonema Nag Raj 1975
Gangliophora Subram. 1992
Gangliostilbe Subram. & Vittal 1976
Garnaudia Borowska 1977
Gaubaea Petr. 1942
Geastrumia Bat. 1960
Gelatinocrinis Matsush. 1995
Gelatinopycnis Dyko & B. Sutton 1979
Geminoarcus K. Ando 1993
Gemmulina Descals & Marvanová 1999
Geniopila Marvanová & Descals 1985
Geosmithia Pitt 1979*
Gerulajacta Preuss 1855
Gilmaniella G.L. Barron 1964
Giulia Tassi 1904
Glaphyriopsis B. Sutton & Pascoe 1987
Glenosporopsis O.M. Fonseca 1943
Glioannellodochium Matsush. 1989
Glioblastocladium Matsush. 1989
Gliophragma Subram. & Lodha 1964
Globoconidiopsis G.F. Sepúlveda, PereiraCarv. & Dianese 2009*
Globoconidium G.F. Sepúlveda, PereiraCarv. & Dianese 2009*
Globuliroseum Sullia & K.R. Khan 1984
Gloeodes Colby 1920
Gloeosporiella Cavara 1892
Gloiosphaera Höhn. 1902
Glutinium Fr. 1849
Glycyphila Mont. 1851
Goidanichiella G.L. Barron ex W. Gams
2009*
Gonatobotryum Sacc. 1880
Gonatophragmiella Rak. Kumar, A.N. Rai
& Kamal 1994
39
Gonatorrhodum Corda 1839
Goniopila Marvanová & Descals 1985
Gonyella Syd. & P. Syd. 1919
Goosiella Morgan-Jones, Kamal & R.K.
Verma 1986
Goosiomyces N.K. Rao & Manohar. 1989
Gorgomyces M. Gönczöl & Révay 1985
Grallomyces F. Stevens 1918
Granularia Sacc. & Ellis ex Sacc. 1882
Graphiothecium Fuckel 1870
Groveolopsis Boedijn 1951
Gymnoxyphium Cif., Bat. & I.J. Araujo
1963
Gyoerffyella Kol 1928
Gyrothrix (Corda) Corda 1842
Hadronema Syd. & P. Syd. 1909
Hadrosporium Syd. 1938
Halysiomyces E.G. Simmons 1981
Halysium Corda 1837
Hansfordia S. Hughes 1951
Hansfordiopeltis Bat. & C.A.A. Costa
1956
Hansfordiopeltopsis M.L. Farr 1986
Hapalosphaeria Syd. 1908
Haplariella Syd. & P. Syd. 1908
Haplariopsis Oudem. 1903
Haplobasidion Erikss. 1889
Haplolepis Syd. 1925
Haplosporidium Speg. 1912
Haptocara Drechsler 1975
Harmoniella V.N. Boriss. 1981
Harpagomyces Wilcz. 1911
Harpographium Sacc. 1880
Hawksworthia Manohar., N.K. Rao, D.K.
Agarwal & Kunwar 2004
Hawksworthiana U. Braun 1988
Heimiodora Nicot 1960
Helhonia B. Sutton 1980
Helicoceras Linder 1931
Helicofilia Matsush. 1983
Helicogoosia Hol.-Jech. 1991
Helicominopsis Deighton 1960
Helicorhoidion S. Hughes 1958
Helicosingula P.S. van Wyk, Marasas,
Baard & Knox-Dav. 1985
Helicosporina G. Arnaud ex Rambelli
1960
Helicostilbe Höhn. 1902
Helicothyrium I. Hino & Katum. 1961
Helicoubisia Lunghini & Rambelli 1979
Heliocephala V.G. Rao, K.A. Reddy & de
Hoog 1984
40
Heliscella Marvanová 1980
Heliscina Marvanová 1980
Helminthophora Bonord. 1851
Helminthosporiomyces G.F. Sepúlveda,
Pereira-Carv. & Dianese 2009*
Hemibeltrania Piroz. 1963
Hemicorynesporella Subram. 1992
Hemidothis Syd. & P. Syd. 1916
Hemisphaeropsis Petr. 1947
Hemisynnema Subram. 1995
Hendersoniella Tassi 1900
Hendersonina E.J. Butler 1913
Hendersoniopsis Höhn. 1918
Hendersonulina Petr. 1951
Henicospora P.M. Kirk & B. Sutton 1980
Hennebertia M. Morelet 1969
Hermatomyces Speg. 1910
Herposira Syd. 1938
Herreromyces R.F. Castañeda & W.B.
Kendr. 1991
Heterocephalum Thaxt. 1903
Heteroconidium Sawada 1944
Heterosporiopsis Petr. 1950
Hexacladium D.L. Olivier 1983
Heydenia Fresen. 1852
Himantia Pers. 1801
Hinoa Hara & I. Hino 1961
Hirudinaria Ces. 1856
Hobsoniopsis D. Hawksw. 2001
Hoehneliella Bres. & Sacc. 1902
Holubovaea Mercado 1983
Holubovaniella R.F. Castañeda 1985
Homalopeltis Bat. & Valle 1961
Hormiactella Sacc. 1886
Hormiactis Preuss 1851
Hormiscioideus M. Blackw. & Kimbr.
1979
Hormisciopsis Sumst. 1914
Hormocephalum Syd. 1939
Hormocladium Höhn. 1923
Hormococcus Robak 1956
Hormographis Guarro, Punsola & Arx
1986
Hughesinia J.C. Lindq. & Gamundí 1970
Humicolopsis Cabral & S. Marchand 1976
Humicolopsis Verona 1977
Hyalocamposporium Révay & J. Gönczöl
2007
Hyalocladium Mustafa 1977
Hyalodictyum Woron. 1916
Hyalohelicomina T. Yokoy. 1974
Mycosphere
Hyalopleiochaeta R.F. Castañeda, Guarro
& Cano 1996
Hyalosynnema Matsush. 1975
Hyalothyridium Tassi 1900
Hyalotiastrum Abbas, B. Sutton, Ghaffar
& Abbas 2003
Hyalotiella Papendorf 1967
Hydrometrospora J. Gönczöl & Révay
1985
Hymenella Fr. 1822
Hymeniopeltis Bat. 1959
Hymenobactron (Sacc.) Höhn. 1916
Hymenopsis Sacc. 1886
Hyphochlaena Cif. 1962
Hyphodiscosia Lodha & K.R.C. Reddy
1974
Hyphodiscosioides Matsush. 1993
Hyphopolynema Nag Raj 1977
Hyphostereum Pat. 1892
Hyphothyrium B. Sutton & Pascoe 1989
Hypocline Syd. 1939
Hypogloeum Petr. 1923
Hysteridium P. Karst. 1905
Hysteropycnis Hilitzer 1929
Ialomitzia Gruia 1964
Icerymyces Brain 1923
Idiocercus B. Sutton 1967
Impudentia Vujanović 2003
Inesiosporium R.F. Castañeda & W. Gams
1997
Infrafungus Cif. 1951
Infundibulomyces Plaingam, Somrith. &
E.B.G. Jones 2003
Inifatiella R.F. Castañeda 1985
Intercalarispora J.L. Crane & Schokn.
1983
Ionophragmium Peres 1961
Irpicomyces Deighton 1969
Isariella Henn. 1908
Ischnostroma Syd. & P. Syd. 1914
Isthmolongispora Matsush. 1971
Isthmophragmospora Kuthub. & Nawawi
1992
Isthmotricladia Matsush. 1971
Ityorhoptrum P.M. Kirk 1986
Iyengarina Subram. 1958
Jahniella Petr. 1921
Jainesia Gonz. Frag. & Cif. 1925
Janetia M.B. Ellis 1976
Javonarxia Subram. 1995
Jerainum Nawawi & Kuthub. 1992
Jubispora B. Sutton & H.J. Swart 1986
Junctospora Minter & Hol.-Jech. 1981
Junewangia W.A. Baker & Morgan-Jones
2002
Kalchbrenneriella Diederich & M.S.
Christ. 2002
Kaleidosporium Van Warmelo & B.
Sutton 1981
Kamatella Anahosur 1969
Kameshwaromyces Kamal, R.K. Verma &
Morgan-Jones 1986
Karsteniomyces D. Hawksw. 1980
Kionocephala P.M. Kirk 1986
Kionochaeta P.M. Kirk & B. Sutton 1986
Kmetia Bres. & Sacc. 1902
Kmetiopsis Bat. & Peres 1960
Knemiothyrium Bat. & J.L. Bezerra 1960
Knufia L.J. Hutchison & Unter. 1996
Kodonospora K. Ando 1993
Kolletes Kohlm. & Volkm.-Kohlm. 2005
Kontospora A. Roldán, Honrubia &
Marvanová 1990
Koorchalomella Chona, Munjal & J.N.
Kapoor 1958
Korunomyces Hodges & F.A. Ferreira
1981
Kostermansinda Rifai 1968
Kostermansindiopsis R.F. Castañeda 1986
Kramabeeja G.V. Rao & K.A. Reddy 1981
Kramasamuha Subram. & Vittal 1973
Kreiseliella Braun 1991
Krishnamyces Hosag. 2003
Kyphophora B. Sutton 1991
Lacellina Sacc. 1913
Lacellinopsis Subram. 1953
Lachnodochium Marchal 1895
Laciniocladium Petri 1917
Lactydina Subram. 1978
Laeviomyces D. Hawksw. 1981
Lagenomyces Cavalc. & A.A. Silva 1972
Lambdasporium Matsush. 1971
Lambinonia Sérus. & Diederich 2005
Lamproconium (Grove) Grove 1937
Lappodochium Matsush. 1975
Lasiodiplodiella Zambett. 1955
Lasiophoma Speg. 1918
Lasiothyrium Syd. & P. Syd. 1913
Lasmenia Speg. 1886
Lasmeniella Petr. & Syd. 1927
Latericonis G.V. Rao, K.A. Reddy & de
Hoog 1984
Lateriramulosa Matsush. 1971
41
Laterispora Uecker, W.A. Ayers & P.B.
Adams 1982
Lauriomyces R.F. Castañeda 1990
Lawalreea Dieder. 1990
Leandria Rangel 1915
Lecaniocola Brain 1923
Lecanostictopsis B. Sutton & Crous 1997
Lecythispora Chowdhry 1985
Lecythophora-like, anamorphic Igneocumulus A.W. Ramaley 2003
Leeina Petr. 1923
Leightoniomyces D. Hawksw. & B. Sutton
1977
Lembuncula Cif. 1954
Lemkea Morgan-Jones & R.C. Sinclair
1983
Lepisticola W. Gams 2009*
Leprieurina G. Arnaud 1918, also see
Prillieuxina
Leprieurinella Bat. & H. Maia 1961
Leptochlamys Died. 1921
Leptodermella Höhn. 1915
Leptodiscella Papendorf 1969
Leptomelanconium Petr. 1923
Leptophyllosticta I.E. Brezhnev 1939
Leptostromella (Sacc.) Sacc. 1884
Leptothyrella Sacc. 1885
Leptothyrina Höhn. 1915
Leptothyrium Kunze 1823
Leucodochium Syd. & P. Syd. 1917
Leucopenicillifer G.R.W. Arnold 1971
Leuliisinea Matsush. 1985
Libertiella Speg. & Roum. 1880
Lichenobactridium Diederich & Etayo
1995
Lichenoconium Petr. & Syd. 1927
Lichenodiplis Dyko & D. Hawksw. 1979
Lichenodiplisiella S.Y. Kondr. & Kudratov 2002
Lichenohendersonia Calat. & Etayo 2001
Lichenopuccinia D. Hawksw. & Hafellner
1984
Lichenostella Calat. & Etayo 1999
Lichenosticta Zopf 1898
Linkosia A. Hern. Gut. & B. Sutton 1997
Linochorella Syd. & P. Syd. 1912
LinodochiumHöhn. 1909
Listeromyces Penz. & Sacc. 1901
Lobatopedis P.M. Kirk 1979
Loliomyces Maire 1937
Lomaantha Subram. 1954
Lomachashaka Subram. 1956
42
Lunulospora Ingold 1942
Luzfridiella R.F. Castañeda & W.B. Kendr.
1991
Lylea Morgan-Jones 1975
Lysotheca Cif. 1962
Mackenziella Yanna & K.D. Hyde 2009*
Macrodiplodia Sacc. 1884
Macrodiplodiopsis Petr. 1922
Macrotrichum Grev. 1825
Magdalaenaea G. Arnaud 1952
Mahabalella B. Sutton & S.D. Patil 1966
Mammariopsis L.J. Hutchison & J. Reid
1988
Manginella Bat. & H. Maia 1961
Manoharachariella Bagyan., N.K. Rao &
Kunwar 2009*
Manoharachariomyces N.K. Rao, D.K.
Agarwal & Kunwar 2005
Mapletonia B. Sutton 1991
Margarinomyces Laxa 1930
Martinellisia V.G. Rao & Varghese 1977
Massalongina Bubák 1916
Massariothea Syd. 1939
Mastigomyces Imshen. & Kriss 1933
Mastigosporium Riess 1852
Matsushimaea Subram. 1978
Matsushimiella R.F. Castañeda & Heredia
2001
Matsushimomyces V.G. Rao & Varghese
1979
Mauginiella Cavara 1925
Medusamyces G.L. Barron & Szijarto 1990
Megacapitula J.L. Chen & Tzean 1993
Megalodochium Deighton 1960
Melanocephala S. Hughes 1979
Melanodiscus Höhn. 1918
Melanographium Sacc. 1913
Melanophoma Papendorf & J.W. du Toit
1967
Melanosella Örösi-Pál 1936
Melophia Sacc. 1884
Menoidea L. Mangin & Har. 1907
Mercadomyces J. Mena 1988
Metadiplodia Syd. 1937
Metarhiziopsis D.W. Li, R.S. Cowles &
Vossbrinck 2008
Metazythia Petr. 1950
Metazythiopsis M. Morelet 1988
Microblastosporon Cif. 1930
Microclava F. Stevens 1917
Microdiscula Höhn. 1915
Mycosphere
Microdothiorella C.A.A. Costa & Sousa
da Câmara 1955
Microhendersonula Dias & Sousa da
Câmara 1952
Microperella Höhn. 1909
Microtyle Speg. 1919
Microxyphiella Speg. 1918
Microxyphiopsis Bat. 1963
Milospium D. Hawksw. 1975
Mindoa Petr. 1949
Miniancora Marvanová & Bärl. 1989
Minimelanolocus R.F. Castañeda & Heredia 2001
Minimidochium B. Sutton 1970
Minutoexcipula V. Atienza & D. Hawksw.
1994
Minutophoma D. Hawksw. 1981
Mirimyces Nag Raj 1993
Mixtoconidium Etayo 1995
Mohgaonidium Singhai 1974
Monilochaetes Halst. ex Harter 1916
Monochaetiella E. Castell. 1943
Monochaetopsis Pat. 1931
Monodia Breton & Faurel 1970
Monodidymaria U. Braun 1994
Monodisma Alcorn 1975
Monosporiella Speg. 1918
Monosporium Bonord. 1851, nom. illegit.
Monotrichum Gäum. 1922
Moorella P.Rag. Rao & D. Rao 1964
Moralesia Urries 1956
Morosporium Renault & Roche 1898
Morrisographium M. Morelet 1968
Mucobasispora Mustafa & Abdul-Wahid
1990
Mucosetospora M. Morelet 1972
Muiaria Thaxt. 1914
Muiogone Thaxt. 1914
Muirella R. Sprague 1959
Multicladium K.B. Deshp. & K.S. Deshp.
1966
Multipatina Sawada 1928
Mycelephas R.F. Castañeda* 2009
Myceloderma Ducomet 1907
Mycobacillaria Naumov 1915
Mycocentrodochium K. Matsush. & Matsush. 1996
Mycochlamys S. Marchand & Cabral 1976
Mycoenterolobium Goos 1970
Mycofalcella Marvanová, Om-Kalth. & J.
Webster 1993
Mycomyces Wyss-Chod. 1928
Mycopara Bat. & J.L. Bezerra 1960
Mycospraguea U. Braun & Rogerson 1993
Mycosticta Höhn. 1918
Mycosylva M.C. Tulloch 1973
Mycotribulus Nag Raj & W.B. Kendr.
1970
Mycousteria M.L. Farr 1986
Myiocoprula Petr. 1955
Myriellina Höhn. 1915
Myriodontium Samson & Polon. 1978
Myrmecomyces Jouvenaz & Kimbr. 1991
Myropyxis Ces. ex Rabenh. 1851
Myrotheciastrum Abbas & B. Sutton 1988
Mystrosporiella Munjal & Kulshr. 1969
Myxoparaphysella Caball. 1941
Myxosporella Sacc. 1881
Myxosporidiella Negru 1960
Myxostomellina Syd. 1931
Myxothyriopsis Bat. & A.F. Vital 1956
Myxothyrium Bubák & Kabát 1915
Naemaspora Pers. 1796
Naemosphaera P. Karst. 1888
Naemosphaerella Höhn. 1923
Nagrajia R.F. Castañeda & W.B. Kendr.
1991
Nagrajomyces Melnik 1984
Nakatopsis Whitton, McKenzie & K.D.
Hyde 2001
Nanoschema B. Sutton 1980
Naothyrsium Bat. 1960
Naranus Ts. Watan. 1995
Navaneethospora V.G. Rao 1994
Necraphidium Cif. 1951
Negeriella Henn. 1897
Nematogonum Desm. 1834
Nematographium Goid. 1935
Nemozythiella Höhn. 1925
Neoalpakesa Punith. 1981
Neoarbuscula B. Sutton 1983
Neobarclaya Sacc. 1899
Neodiplodina Petr. 1954
Neofuckelia Zeller & Goodd. 1935
Neohendersonia Petr. 1921
Neoheteroceras Nag Raj 1993
Neojohnstonia B. Sutton 1983
Neoligniella Naumov 1951
Neomarssoniella U. Braun 1991
Neomelanconium Petr. 1940
Neoovularia U. Braun 1992
Neopeltis Syd. 1937
Neopericonia Kamal, A.N. Rai & MorganJones 1983
43
Neophoma Petr. & Syd. 1927
Neoplaconema B. Sutton 1977
Neoramularia U. Braun 1991
Neospegazzinia Petr. & Syd. 1936
Neosporidesmium Mercado & J. Mena
1988
Neottiospora Desm. 1843
Neta Shearer & J.L. Crane 1971
Nidulispora Nawawi & Kuthub. 1990
Nigromacula Etayo 2002
Nigropuncta D. Hawksw. 1981
Nosophloea Fr. 1849
NothosporaPeyronel 1913
NummosporaE. Müll. & Shoemaker 1964
Nusia Subram. 1995
Nyctalospora E.F. Morris 1972
Obeliospora Nawawi & Kuthub. 1990
Obstipipilus B. Sutton 1968
Obstipispora R.C. Sinclair & MorganJones 1979
Ochroconis de Hoog & Arx 1974
Octopodotus Kohlm. & Volkm.-Kohlm.
2003
Odontodictyospora Mercado 1984
Oedocephalum Preuss 1851
Oedothea Syd. 1930
Ojibwaya B. Sutton 1973
Olpitrichum G.F. Atk. 1894
Omega B. Sutton & Minter 1988
Ommatosporella Bat., J.L. Bezerra &
Poroca 1967
Oncopodiella G. Arnaud ex Rifai 1965
Oncopodium Sacc. 1904
Oncospora Kalchbr. 1880
Oncosporella P. Karst. 1887
Oncostroma Bat. & Marasas 1966
Onychophora W. Gams, P.J. Fisher & J.
Webster 1984
Oothyrium Syd. 1939
Ophiosira Petr. 1955
Orbimyces Linder 1944
Ordus K. Ando & Tubaki 1983
Ostracodermidium Mukerji 1973
Ovadendron Sigler & J.W. Carmich. 1976
Paathramaya Subram. 1956
Pachycladina Marvanová 1987
Pactilia Fr. 1837
Palawaniopsis Bat., Cif. & Nascim. 1959
Panchanania Subram. & N.G. Nair 1966
Papilionospora V.G. Rao & B. Sutton
1976
Pappimyces B. Sutton & Hodges 1975
44
Paraaoria R.K. Verma & Kamal 1987
Paraarthrocladium Matsush. 1993
Paraceratocladium R.F. Castañeda 1987
Parachionomyces Thaung 1979
Paracostantinella Subram. & Sudha 1989
Paracryptophiale Kuthub. & Nawawi
1994
Paracytospora Petr. 1925
Paradactylaria Subram. & Sudha 1989
Paradactylella Matsush. 1993
Paradendryphiopsis M.B. Ellis 1976
Paradictyoarthrinium Matsush. 1996
Paradidymobotryum C.J.K. Wang & B.
Sutton 1984
Paradiplodia Speg. ex Trotter 1931
Paradischloridium Bhat & B. Sutton 1985
Paradiscula Petr. 1941
Paraepicoccum Matsush. 1993
Parafulvia Kamal, A.N. Rai & MorganJones 1983
Paraharknessia Matsush. 2003
Parahyalotiopsis Nag Raj 1976
Paramassariothea Subram. & Muthumary
1979
Paramenisporopsis Matsush. 2003
Parapericonia M.B. Ellis 1976
Parapericoniella U. Braun, Heuchert & K.
Schub. 2005
Paraphaeoisaria de Hoog & MorganJones 1978
Paraphialocephala Budathoki 1997
Parapithomyces Thaung 1976
Parapleurotheciopsis P.M. Kirk 1982
Parapyricularia M.B. Ellis 1972
Pararobillarda Matsush. 1996
Parasarcopodium Melnik, S.J. Lee &
Crous 2004
Parasphaeropsis Petr. 1953
Paraspora Grove 1884
Parastigmatellina Bat. & C.A.A. Costa
1959
Parasympodiella Ponnappa 1975
Paratetraploa M.K.M. Wong & K.D.
Hyde 2002
Paratomenticola M.B. Ellis 1976
Paratrichoconis Deighton & Piroz. 1972
Paraulocladium R.F. Castañeda 1986
Parvosympodium A.W. Ramaley 2002
Paspalomyces Linder 1933
Patouillardiella Speg. 1889
Patriciomyces D. Hawksw. 2001
Pellionella (Sacc.) Sacc. 1902
Mycosphere
Peltasterella Bat. & H. Maia 1959
Peltasterinostroma Punith. 1975
Peltasteropsis Bat. & H. Maia 1959
Peltistroma Henn. 1904
Peltistromella Höhn. 1907
Peltosoma Syd. 1925
Peltostromellina Bat. & A.F. Vital 1959
Peltostromopsis Bat. & A.F. Vital 1959
Penzigomyces Subram. 1992
Perelegamyces R.F. Castañeda & W.B.
Kendr. 1990
Periola Fr. 1822
Perizomella Syd. 1927
Pestalozziella Sacc. & Ellis ex Sacc. 1882
Petrakia Syd. & P. Syd. 1913
Petrakiopsis Subram. & K.R.C. Reddy
1968
Phacostroma Petr. 1955
Phacostromella Petr. 1955
Phaeoblastophora Partr. & Morgan-Jones
2002
Phaeobotrys M. Calduch, Gené & Guarro
2002
Phaeocandelabrum R.F. Castañeda, Heredia, Saikawa 2009*
Phaeocytostroma Petr. 1921
Phaeodactylella Udaiyan 1992
Phaeodactylium Agnihothr. 1968
Phaeodiscula Cub. 1891
Phaeodomus Höhn. 1909
Phaeohiratsukaea Udagawa & Iwatsu
1990
Phaeohymenula Petr. 1954
Phaeoidiomyces Dorn.-Silva & Dianese
2004
Phaeoisaria Höhn. 1909
Phaeolabrella Speg. 1912
Phaeomonilia R.F. Castañeda, Heredia &
R.M. Arias 2007
Phaeomonostichella Keissl. ex Petr. 1941
Phaeophloeosporella Crous & B. Sutton
1997
Phaeophomopsis Höhn. 1917
Phaeosphaera Bat. & Cif. 1963
Phaeosporobolus D. Hawksw. & Hafellner
1986
Phaeothyrium Petr. 1947
Phaeotrichoconis Subram. 1956
Phalangispora Nawawi & J. Webster 1982
Phialoarthrobotryum Matsush. 1975
Phialogangliospora Udaiyan & V.S.
Hosag. 1992
Phialomyces P.C. Misra & P.H.B. Talbot
1964
Phialophaeoisaria Matsush. 1995
Phialoselanospora Udaiyan 1992
Phialosporostilbe Mercado & J. Mena
1985
Phialostele Deighton 1969
Phialotubus R.Y. Roy & Leelav. 1966
Phloeoconis Fr. 1849
Phloeosporina Höhn. 1924
Phlyctaeniella Petr. 1922
Phomachora Petr. & Syd. 1925
Phomachorella Petr. 1947
Phomyces Clem. 1931
Phragmocephala E.W. Mason & S.
Hughes 1951
Phragmoconidium G.F. Sepúlveda, Pereira-Carv. & Dianese 2009*
Phragmodochium Höhn. 1924
Phragmogloeum Petr. 1954
Phragmopeltis Henn. 1904
Phragmospathula Subram. & N.G. Nair
1966
Phragmospathulella J. Mena & Mercado
1986
Phragmotrichum Kunze 1823
Phylloedium Fr. 1825
Phyllohendersonia Tassi 1902
Physalidiella Rulamort 1990
Physalidiopsis R.F. Castañeda & W.B.
Kendr. 1990
Pilulina G. Arnaud 1954
Pinatubo J.B. Manandhar & Mew 1996
Piperivora Siboe, P.M. Kirk & P.F.
Cannon 1999
Piricauda Bubák 1914
Piricaudilium Hol.-Jech. 1988
Piricaudiopsis J. Mena & Mercado 1987
Pirispora Faurel & Schotter 1966
Pirostomella Sacc. 1914
Pirozynskiella S. Hughes 2007
Pithosira Petr. 1949
Pittostroma Kowalski & T.N. Sieber 1992
Placella Syd. 1938
Placodiplodia Bubák 1916
Placonema (Sacc.) Petr. 1921
Placonemina Petr. 1921
Plagiostigmella Petr. 1949
Plectonaemella Höhn. 1915
Plectopeltis Syd. 1927
Plectophomella Moesz 1922
Plectophomopsis Petr. 1922
45
Plectosira Petr. 1929
Plectronidiopsis Nag Raj 1979
Plectronidium Nag Raj 1977
Plectrothrix Shear 1902
Pleiochaeta (Sacc.) S. Hughes 1951
Plenocatenulis Bat. & Cif. 1959
Plenophysa Syd. & P. Syd. 1920
Plenotrichopsis Bat. 1961
Plenotrichum Syd. 1927
Plenozythia Syd. & P. Syd. 1916
Pleocouturea G. Arnaud 1911
Plesiospora Drechsler 1971
Pleurodesmospora Samson, W. Gams &
H.C. Evans 1979
Pleurodiscula Höhn. 1926
Pleurodiscus Petr. 1931
Pleurodomus Petr. 1934
Pleuropedium Marvanová & S.H. Iqbal
1973
Pleurophragmium Costantin 1888
Pleuroplaconema Petr. 1923
Pleuroplacosphaeria Syd. 1928
Pleurotheciopsis B. Sutton 1973
Pleurothyriella Petr. & Syd. 1925
Pleurothyrium Bubák 1916
Pleurothyrium Bubák 1916
Pocillopycnis Dyko & B. Sutton 1979
Podosporiella Ellis & Everh. 1894
Podosporium Schwein. 1832
Poikilosperma Bat. & J.L. Bezerra 1961
Polybulbophiale Goh & K.D. Hyde 1998
Polycladium Ingold 1959
Polydesmus Mont. 1845
Polyetron Bat. & Peres 1963
Polylobatispora Matsush. 1996
Polyscytalina G. Arnaud 1954
Polyscytalum Riess 1853
Polystomellomyces Bat. 1959
Polystratorictus Matsush. 1993
Polysynnema Constant. & Seifert 1988
Polythrinciella Bat. & H. Maia 1960
Polythrinciopsis J. Walker 1966
Polytretophora Mercado 1983
Porobeltraniella Gusmão 2004
Porocladium Descals 1976
Poroisariopsis M. Morelet 1971
Poropeltis Henn. 1904
Porophilomyces U. Braun 2000
Porosubramaniania Hol.-Jech. 1985
Porrectotheca Matsush. 1996
Powellia Bat. & Peres 1964
Prathigada Subram. 1956
46
Prismaria Preuss 1851
Proboscispora Punith. 1984
Prophytroma Sorokīn 1877
Prosthemiella Sacc. 1881
Protostegia Cooke 1880
Protostegiomyces Bat. & A.F. Vital 1955
Protostroma Bat. 1957
Psammina Sacc. & M. Rousseau ex E.
Bommer & M. Rousseau 1891
Pseudoacrodictys W.A. Baker & MorganJones 2003
Pseudoanguillospora S.H. Iqbal 1974
Pseudoaristastoma Suj. Singh 1979
Pseudoasperisporium U. Braun 2000
Pseudobasidiospora Dyko & B. Sutton
1978
Pseudobeltrania Henn. 1902
Pseudocanalisporium R.F. Castañeda &
W.B. Kendr. 1991
Pseudocenangium P. Karst. 1886
Pseudochuppia Kamal, A.N. Rai &
Morgan-Jones 1984
Pseudoclathrosphaerina Voglmayr 1997
Pseudoconium Petr. 1969
Pseudocytoplacosphaeria Punith. & Spooner 2002
Pseudocytospora Petr. 1923
Pseudodichomera Höhn. 1918
Pseudodidymaria U. Braun 1993
Pseudodiplodia (P. Karst.) Sacc. 1884
Pseudodiscula Laubert 1911
Pseudoepicoccum M.B. Ellis 1971
Pseudofuscophialis Sivan. & H.S. Chang
1995
Pseudogaster Höhn. 1907
Pseudogliophragma Phadke & V.G. Rao
1980
Pseudographiella E.F. Morris 1966
Pseudohansfordia G.R.W. Arnold 1970
Pseudohelicomyces Garnica & E. Valenz.
2000
Pseudohendersonia Crous & M.E. Palm
1999
Pseudolachnea Ranoj. 1910
Pseudoneottiospora Faurel & Schotter
1965
Pseudopapulaspora N.D. Sharma 1977
Pseudopatellina Höhn. 1908
Pseudopeltistroma Katum. 1975
Pseudopetrakia M.B. Ellis 1971
Pseudophloeosporella U. Braun 1993
Mycosphere
Pseudophragmotrichum W.P. Wu, B.
Sutton & Gange 1998
Pseudopolystigmina Murashk. 1928
Pseudoramularia Matsush. 1983
Pseudorhizopogon Kobayasi 1983
Pseudoschizothyra Punith. 1980
Pseudoseptoria Speg. 1910
Pseudostegia Bubák 1906
Pseudostilbella Munt.-Cvetk. & GómezBolea 1995
Pseudostracoderma A.E. Martinez &
Godeas 1997
Pseudothyrium Höhn. 1927
Pseudotorula Subram. 1958
Pseudotracylla B. Sutton & Hodges 1976
Pseudotrichoconis W.A. Baker & MorganJones 2001
Pseudozythia Höhn. 1902
Pterulopsis Wakef. & Hansf. 1943
Pterygosporopsis P.M. Kirk 1983
Pucciniospora Speg. 1886
Pulchromyces Hennebert 1973
Pullospora Faurel & Schotter 1965
Pulvinella A.W. Ramaley 2001
Pulvinotrichum Gamundí, Aramb. &
Giaiotti 1981
Punctillina Toro 1934
Pycnidioarxiella Punith. & N.D. Sharma
1980
Pycnidiopeltis Bat. & C.A.A. Costa 1959
Pycnis Bref. 1881
Pycnodactylus Bat., A.A. Silva & Cavalc.
1967
Pycnodallia Kohlm. & Volkm.-Kohlm.
2001
Pycnoharknessia Matsush. 1996
Pycnomma Syd. 1924
Pycnomoreletia Rulamort 1990
Pycnopleiospora C.Z. Wei, Y. Harada &
Katum. 1997
Pycnoseynesia Kuntze 1898
Pycnothera N.D. Sharma & G.P. Agarwal
1974
Pycnothyriella Bat. 1952
Pycnothyrium Died. 1913
Pyramidospora Sv. Nilsson 1962
Pyrenyllium Clem. 1909
Pyrgostroma Petr. 1951
Pyriculariopsis M.B. Ellis 1971
Pyripnomyces Cavalc. 1972
Quadracaea Lunghini, Pinzari & Zucconi
1996
Quadricladium Nawawi & Kuthub. 1989
Quasidiscus B. Sutton 1991
Queenslandia Bat. & H. Maia 1959
Quezelia Faurel & Schotter 1965
Radiatispora Matsush. 1995
Radiciseta Sawada & Katsuki 1959
Radulidium Arzanlou, W. Gams & Crous
2007
Raizadenia S.L. Srivast. 1981
Ramakrishnanella Kamat & Ullasa ex
Ullasa 1970
Ramaraomyces N.K. Rao, Manohar. &
Goos 1989
Ramicephala Voglmayr & G. Delgado
2003
Ramoacrodictys G.Z. Zhao 2009*
Ramoconidiifera B. Sutton, Carmarán &
A.I. Romero 1996
Ramulaspera Lindr. 1902
Ranojevicia Bubák 1910
Redbia Deighton & Piroz. 1972
Refractohilum D. Hawksw. 1977
Reichlingia Diederich & Scheid. 1996
Remersonia Samson & Seifert 1997
Repetoblastiella R.F. Castañeda, Minter &
M. Stadler 2009*
Retroconis de Hoog & Bat. Vegte 1989
Rhabdoclema Syd. 1939
Rhabdomyces Balbiani 1889
Rhabdostromella Höhn. 1915
Rhexoacrodictys W.A. Baker & MorganJones 2002
Rhexoampullifera P.M. Kirk 1982
Rhexodenticula W.A. Baker & MorganJones 2001
Rhexoprolifer Matsush. 1996
Rhinocladium Sacc. & Marchal 1885
Rhinotrichella G. Arnaud ex de Hoog
1977
Rhipidocephalum Trail 1888
Rhizohypha Chodat & Sigr. 1911
Rhizosphaerina B. Sutton 1986
Rhodesiopsis B. Sutton & R. Campb. 1979
Rhodothallus Bat. & Cif. 1959
Rhombostilbella Zimm. 1902
Rhynchodiplodia Briosi & Farneti 1906
Rhynchomyces Willk. 1866
Rhynchoseptoria Unamuno 1940
Rhynchosporina Arx 1957
Riclaretia Peyronel 1915
Rileya A. Funk 1979
Robakia Petr. 1952
47
Robillarda Sacc. 1882
Rogergoosiella A. Hern. Gut. & J. Mena
1996
Roigiella R.F. Castañeda 1984
Roscoepoundia Kuntze 1898
Rosulomyces S. Marchand & Cabral 1976
Rota Bat., Cif. & Nascim. 1959
Rotaea Ces. ex Schltdl. 1851
Rubikia H.C. Evans & Minter 1985
Ruggieria Cif. & Montemart. 1958
Rutola J.L. Crane & Schokn. 1978
Sadasivania Subram. 1957
Sagrahamala Subram. 1972
Saliastrum myrtilli (Allesch.) Kujala 1946
Sanjuanomyces R.F. Castañeda & W.B.
Kendr. 1991
Santapauinda Subram. 1995
Saprophragma K.B. Deshp. & K.S. Deshp.
1966
Saprotaphrina Verona & Rambelli 1962
Sarbhoyomyces Saikia 1981
Sarcinodochium Höhn. 1905
Sarcinosporon D.S. King & S.C. Jong
1975
Sarcopodium Ehrenb. 1818
Satchmopsis B. Sutton & Hodges 1975
Scaphidium Clem. 1901
Scenomyces F. Stevens 1927
Sceptrifera Deighton 1965
Schizoderma Kunze 1825
Schizothyra Bat. & C.A.A. Costa 1957
Schizothyrella Thüm. 1880
Schizothyropsis Bat. & A.F. Vital 1960
Schizotrichum McAlpine 1903
Schroeteria G. Winter 1881
Schwarzmannia Pisareva 1968
Scirrhophoma Petr. 1941
Sclerococcum Fr. 1825
Scleroconium Syd. 1935
Sclerodiscus Pat. 1890
Sclerodothiorella Died. 1912
Sclerographiopsis Deighton 1973
Sclerographium Berk. 1854
Scleromeris Syd. 1926
Scleropycnis Syd. & P. Syd. 1911
Sclerotiella A.K. Sarbhoy & A. Sarbhoy
1975
Sclerozythia Petch 1937
Scolecobasidiella M.B. Ellis 1971
Scolecobasidium E.V. Abbott 1927
Scolecodochium K. Matsush. & Matsush.
1996
48
Scolecotheca Søchting & B. Sutton 1997
Scolecozythia Curzi 1927
Scoliotidium Bat. & Cavalc. 1963
Scopaphoma Dearn. & House 1925
Scopulariella Gjaerum 1971
Scorpiosporium S.H. Iqbal 1974
Scothelius Bat., J.L. Bezerra & Cavalc.
1965
Scutisporus K. Ando & Tubaki 1985
Scutopeltis Bat. & H. Maia 1957
Scutopycnis Bat. 1957
Seifertia Partr. & Morgan-Jones 2002
Seimatosporiopsis B. Sutton, Ghaffer &
Abbas 1972
Selenodriella R.F. Castañeda & W.B.
Kendr. 1990
Selenosira Petr. 1957
Selenosporopsis R.F. Castañeda & W.B.
Kendr. 1991
Septocyta Petr. 1927
Septocytella Syd. 1929
Septodochium Matsush. 1971
Septogloeum Sacc. 1880
Septomyxella (Höhn.) Höhn. 1923
Septopatella Petr. 1925
Septosporiopsis W.A. Baker & MorganJones 2009*
Septosporium Corda 1831
Septothyrella Höhn. 1911
Septotrullula Höhn. 1902
Sessiliospora D. Hawksw. 1979
Setolibertella Punith. & Spooner 1999
Setophiale Matsush. 1995
Setosporella Mustafa & Abdul-Wahid
1989
Setosynnema D.E. Shaw & B. Sutton 1985
Seychellomyces Matsush. 1981
Seynesiopsis Henn. 1904
Shawiella Hansf. 1957
Sheariella Petr. 1952
Shrungabeeja V.G. Rao & K.A. Reddy
1981
Siamia V. Robert, Decock & R.F. Castañeda 2000
Sigmatomyces Sacc. & P. Syd. 1913
Sirexcipula Bubák 1907
Sirocyphis Clem. 1909
Sirodochiella Höhn. 1925
Sirogloea Petr. 1923
Siroligniella Naumov 1926
Sirophoma Höhn. 1917
Siroplacodium Petr. 1940
Mycosphere
Siropleura Petr. 1934
Siroscyphellina Petr. 1923
Sirosperma Syd. & P. Syd. 1916
Sirosphaera Syd. & P. Syd. 1913
Sirosporonaemella Naumov 1951
Sirothecium P. Karst. 1887
Sirothyriella Höhn. 1910
Sirothyrium Syd. & P. Syd. 1916
Sirozythia Höhn. 1904
Sirozythiella Höhn. 1909
Sitochora H.B.P. Upadhyay 1964
Slimacomyces Minter 1986
Solheimia E.F. Morris 1967
Solicorynespora R.F. Castañeda & W.B.
Kendr. 1990
Soloacrospora W.B. Kendr. & R.F. Castañeda 1991
Solosympodiella Matsush. 1971
Soloterminospora Matsush. 1996
Spegazzinia Sacc. 1879
Speiropsis Tubaki 1958
Spermatoloncha Speg. 1908
Spermochaetella Cif. 1954
Spermospora R. Sprague 1948
Spermosporella Deighton 1969
Spermosporella Deighton 1969
Sphaeridium Fresen. 1852
Sphaeriostromella Bubák 1916
Sphaeriothyrium Bubák 1916
Sphaerocolla P. Karst. 1892
Sphaerocybe Magrou & Marneffe 1946
Sphaeromma H.B.P. Upadhyay 1964
Sphaeronema Fr. 1815
Sphaerophoma Petr. 1924
Sphaerosporium Schwein. 1832
Sphaerulomyces Marvanová 1977
Sphondylocephalum Stalpers 1974
Spicularia Pers. 1822
Spinulospora Deighton 1973
Spiralum J.L. Mulder 1975
Spiropes Cif. 1955
Spondylocladiella Linder 1934
Spondylocladiopsis M.B. Ellis 1963
Sporendonema Desm. 1827
Sporocystis Morgan 1902
Sporoglena Sacc. 1894
Sporophiala P.Rag. Rao 1970
Sporophora Luteraan 1952
Stachybotryella Ellis & Barthol. 1902
Stachybotryna Tubaki & T. Yokoy. 1971
Stachycoremium Seifert 1986
Stachylidium Link 1809
Stagonopatella Petr. 1927
Stagonopsis Sacc. 1884
Stagonosporina Tassi 1902
Stagonostromella Petr. & Syd. 1927
Staheliella Emden 1974
Stalagmochaetia Cif. & Bat. 1963
Staphylotrichum J. Mey. & Nicot 1957
Stauriella Sivichai & E.B.G. Jones 2004
Stauronema (Sacc.) Syd., P. Syd. & E.J.
Butler 1916
Stauronematopsis Abbas, B. Sutton &
Ghaffar 2002
Staurophoma Höhn. 1907
Stegolerium Strobel, W.M. Hess & E.J.
Ford 2001
Stegonsporiopsis Van Warmelo & B.
Sutton 1981
Stellomyces Morgan-Jones, R.C. Sinclair
& Eicker 1987
Stellopeltis Bat. & A.F. Vital 1959
Stellospora Alcorn & B. Sutton 1984
Stellothyriella Bat. & Cif. 1959
Stenocephalopsis Chamuris & C.J.K.
Wang 1998
Stenocephalum Chamuris & C.J.K. Wang
1990
Stenocladiella Marvanová & Descals 1987
Stenospora Deighton 1969
Stephanosporium Dal Vesco 1961
Stephembruneria R.F. Castañeda 1988
Sterigmatobotrys Oudem. 1886
Stevensomyces E.F. Morris & Finley 1965
Stevensonula Petr. 1952
Stichospora Petr. 1927
Stictopatella Höhn. 1918
Stictosepta Petr. 1964
Stigmatellina Bat. & H. Maia 1960
Stigmella Lév. 1842
Stigmopeltis Syd. 1927
Stilbellula Boedijn 1951
Stilbophoma Petr. 1942
Stilbospora Pers. 1794, also see Prosthecium
Strasseria Bres. & Sacc. 1902
Strasseriopsis B. Sutton & Tak. Kobay.
1970
Stratiphoromyces Goh & K.D. Hyde 1998
Striosphaeropsis Verkley & Aa 1997
Stromatopogon Zahlbr. 1897
Stromatopycnis A.F. Vital 1956
Stromatostysanus Höhn. 1919
Strongylothallus Bat. & Cif. 1959
49
Strumellopsis Höhn. 1909
Stygiomyces Coppins & S.Y. Kondr. 1995
Stylaspergillus B. Sutton, Alcorn & P.J.
Fisher 1982
Subicularium M.L. Farr & Goos 1989
Subramania D. Rao & P.Rag. Rao 1964
Subramanianospora Narayanan, J.K.
Sharma & Minter 2003
Subramaniomyces Varghese & V.G. Rao
1980
Subulispora Tubaki 1971
Surculiseries Okane, Nakagiri & Tad. Ito
2001
Sutravarana Subram. & Chandrash. 1977
Suttoniella S. Ahmad 1961
Suttonina H.C. Evans 1984
Syamithabeeja Subram. & Natarajan 1976
Sylviacollaea Cif. 1963
Symbiotaphrina Kühlw. & Jurzitza ex W.
Gams & Arx 1980
Symphyosira Preuss 1853*
Symphysos Bat. & Cavalc. 1967
Sympodiella W.B. Kendr. 1958
Sympodiocladium Descals 1982
Sympodioclathra Voglmayr 1997
Sympodioplanus R.C. Sinclair & Boshoff
1997
Synchronoblastia Uecker & F.L. Caruso
1988
Syncladium Rabenh. 1859
Synnemacrodictys W.A. Baker & MorganJones 2009*
Synnemaseimatoides K. Matsush. &
Matsush. 1996
Synnematomyces Kobayasi 1981
Synnemellisia N.K. Rao, Manohar. &
Goos 1989
Synnmukerjiomyces Aneja & R. Kumar
1999
Synostomina Petr. 1949
Systremmopsis Petr. 1923
Taeniolina M.B. Ellis 1976
Talekpea Lunghini & Rambelli 1979
Tandonia M.D. Mehrotra 1991
Tarsodisporus Bat. & A.A. Silva 1965
Tassia Syd. & P. Syd. 1919
Tawdiella K.B. Deshp. & K.S. Deshp.
1966
Taxomyces Strobel, A. Stierle, D. Stierle &
W.M. Hess 1993
Tectacervulus A.W. Ramaley 1992
Telligia Hendr. 1948
50
Temerariomyces B. Sutton 1993
Teratosperma Syd. & P. Syd. 1909
Tetrabrachium Nawawi & Kuthub. 1987
Tetrabrunneospora Dyko 1978
Tetrachaetum Ingold 1942
Tetracoccosporium Szabó 1905
Tetrameronycha Speg. ex W. Rossi & M.
Blackw. 1990
Tetranacriella Kohlm. & Volkm.-Kohlm.
2001
Tetranacrium H.J. Huds. & B. Sutton 1964
Tetraposporium S. Hughes 1951
Textotheca Matsush. 1996
Thallospora L.S. Olive 1948
Thaptospora B. Sutton & Pascoe 1987
Tharoopama Subram. 1956
Thirumalacharia Rathaiah 1981
Tholomyces Matsush. 2003
Thoracella Oudem. 1900
Thrinacospora Petr. 1948
Thyriostromella Bat. & C.A.A. Costa 1959
Thyrostromella Höhn. 1919
Thyrsidiella Höhn. ex Höhn. 1909
Thyrsidina Höhn. 1905
Tiarosporellivora Punith. 1981
Ticogloea G. Weber, Spaaij & W. Gams
1994
Tilakiopsis V.G. Rao 1994
Titaeopsis B. Sutton & Deighton 1984
Titaeospora Bubák 1916, also see
Stamnaria
Tomenticola Deighton 1969
Tompetchia Subram. 1985
Torulopsiella Bender 1932
Toxosporiella B. Sutton 1986
Toxosporiopsis B. Sutton & Sellar 1966
Toxosporium Vuill. 1896
Tracylla (Sacc.) Tassi 1904
Trematophoma Petr. 1924
Tremellidium Petr. 1927
Tretocephala Subram. 1995
Tretophragmia Subram. & Natarajan 1974
Tretospeira Piroz. 1972
Tretovularia Deighton 1984
Triacutus G.L. Barron & Tzean 1981
Triadelphia Shearer & J.L. Crane 1971
Tribolospora D.A. Reid 1966
Tricellula Beverw. 1954
Trichaegum Corda 1837
Trichobolbus Bat. 1964
Trichobotrys Penz. & Sacc. 1901
Trichoconis Clem. 1909
Mycosphere
Trichodiscula Vouaux 1910
Trichodochium Syd. 1927
Trichomatoclava G.F. Sepúlveda, PereiraCarv. & Dianese 2009*
Trichomatosphaera Pereira-Carv., G.F.
Sepúlveda & Dianese 2009*
Trichopeltulum Speg. 1889
Trichoseptoria Cavara 1892
Trichosporodochium Dorn.-Silva & Dianese 2004
Tricladiella K. Ando & Tubaki 1984
Tricladiopsis Descals 1982
Tricladiospora Nawawi & Kuthub. 1988
Tridentaria Preuss 1852
Trifurcospora K. Ando & Tubaki 1988
Triglyphium Fresen. 1852
Trigonosporium Tassi 1900
Tripoconidium Subram. 1978
Triposporina Höhn. 1912
Triramulispora Matsush. 1975
Triscelophorus Ingold 1944
Triscelosporium Nawawi & Kuthub. 1987
Trisulcosporium H.J. Huds. & B. Sutton
1964
Tritirachium Limber 1940
Troposporium Harkn. 1884
Troposporopsis Whitton, McKenzie &
K.D. Hyde 1999
Trullula Ces. 1852
Tryssglobulus B. Sutton & Pascoe 1987
Tuberculariopsis Höhn. 1909
Tuberculispora Deighton & Piroz. 1972
Tulipispora Révay & Gönczöl 2009*
Tumularia Descals & Marvanová 1987
Tunicago B. Sutton & Pollack 1977
Turturconchata J.L. Chen, T.L. Huang &
Tzean 1999
Tylomyces Cortini 1921
Tympanosporium W. Gams 1974
Uberispora Piroz. & Hodges 1973
Ubrizsya Negru 1965
Ulocoryphus Michaelides, L. Hunter &
W.B. Kendr. 1982
Umbellidion B. Sutton & Hodges 1975
Uncispora R.C. Sinclair & Morgan-Jones
1979
Urohendersonia Speg. 1902
Urohendersoniella Petr. 1955
Uvarispora Goos & Piroz. 1975
Vagnia D. Hawksw. & Miądl. 1997
Vamsapriya Gawas & Bhat 2006
Vanakripa Bhat, W.B. Kendr. & Nag Raj
1993
Vanbeverwijkia Agnihothr. 1961
Vanderystiella Henn. 1908
Vanibandha Manohar., N.K. Rao, Kunwar
& D.K. Agarwal 2006
Vanterpoolia A. Funk 1982
Vargamyces Tóth 1980
Variocladium Descals & Marvanová 1999
Vasculomyces S.F. Ashby 1913
Vasudevella Chona, Munjal & Bajaj 1957
Velutipila D. Hawksw. 1987
Ventrographium H.P. Upadhyay, Cavalc.
& A.A. Silva 1986
Venustisporium R.F. Castañeda & Iturr.
1999
Venustocephala Matsush. 1995
Venustosynnema R.F. Castañeda & W.B.
Kendr. 1990
Veracruzomyces Mercado, Guarro, Heredia & J. Mena 2002
Veramycella G. Delgado 2009*
Veramyces Matsush. 1993
Verdipulvinus A.W. Ramaley 1999
Vermispora Deighton & Piroz. 1972
Vermisporium H.J. Swart & M.A. Will.
1983
Veronaea Cif. & Montemart. 1957
Veronaella Subram. & K.R.C. Reddy 1975
Veronidia Negru 1964
Verrucariella S. Ahmad 1967
Verrucophragmia Crous, M.J. Wingf. &
W.B. Kendr. 1994
Verticicladus Matsush. 1993
Verticimonosporium Matsush. 1971
Vesiculohyphomyces Armando, PereiraCarv. & Dianese 2009*
Vestigium Piroz. & Shoemaker 1972
Virgariella S. Hughes 1953
Virgatospora Finley 1967
Viscomacula R. Sprague 1951
Volucrispora Haskins 1958
Volutellis Clem. & Shear 1931
Volutellopsis Speg. 1910
Vouauxiella Petr. & Syd. 1927
Waihonghopes Yanna & K.D. Hyde 2002
Wardinella Bat. & Peres 1960
Waydora B. Sutton 1976
Websteromyces W.A. Baker & Partr. 2000
Weissia Bat. & M.P. Herrera 1964
Weufia Bhat & B. Sutton 1985
Wiesneriomyces Koord. 1907
51
Wojnowicia Sacc. 1899
Xanthoriicola D. Hawksw. 1973
Xenidiocercus Nag Raj 1993
Xenochalara M.J. Wingf. & Crous 2000
Xenodomus Petr. 1922
Xenoheteroconium Bhat, W.B. Kendr. &
Nag Raj 1993
Xenokylindria DiCosmo, S.M. Berch &
W.B. Kendr. 1983
Xenopeltis Syd. & P. Syd. 1919
Xenoplaca Petr. 1949
Xenostroma Höhn. 1915
Xepicula Nag Raj 1993
Xepiculopsis Nag Raj 1993
Xeroconium D. Hawksw. 1981
Xiambola Minter & Hol.-Jech. 1981
Xiphomyces Syd. & P. Syd. 1916
Xylochia B. Sutton 1983
Xyloglyphis Clem. 1909
Xylohypha (Fr.) E.W. Mason 1960
Xylohyphopsis W.A. Baker & Partr. 2000
Yalomyces Nag Raj 1993
Ybotromyces Rulamort 1986
Yinmingella Goh, K.M. Tsui & K.D. Hyde
1999
Ypsilina J. Webster, Descals & Marvanová
1999
Yuccamyces Gour, Dyko & B. Sutton 1979
Yunnania H.Z. Kong 1998
Zakatoshia B. Sutton 1973
Zanclospora S. Hughes & W.B. Kendr.
1965
Zebrospora McKenzie 1991
Zelandiocoela Nag Raj 1993
Zelopelta B. Sutton & R.D. Gaur 1984
Zelosatchmopsis Nag Raj 1991
Zelotriadelphia R.F. Castañeda, Saikawa,
M. Stadler & Iturr. 2005
Zetesimomyces Nag Raj 1988
Zevadia J.C. David & D. Hawksw. 1995
Zilingia Petr. 1934
Zinzipegasa Nag Raj 1993
Zopheromyces B. Sutton & Hodges 1977
Zunura Nag Raj 1993
Zygosporium Mont. 1842
Zythia Fr. 1849
Zyxiphora B. Sutton 1981
Phylum BASIDIOMYCOTA R.T. Moore
Subphylum AGARICOMYCOTINA Doweld
Class Agaricomycetes Doweld
Agaricales Underw.
52
Agaricaceae Chevall.
Attamyces Kreisel 1972
Coccobotrys Boud. & Pat. 1900, anamorphic Lepiota (Pers.) Gray 1821
Fistulinaceae Lotsy
Confistulina Stalpers 1983, anamorphic
Fistulina Bull. 1791
Lyophyllaceae Jülich
Termitosphaera Cif. 1935, anamorphic
Termitomyces R. Heim 1942
Ugola Adans. 1763, anamorphic Asterophora Ditmar 1809
Mycenaceae Overeem
Decapitatus Redhead & Seifert 2000,
anamorphic Mycena (Pers.) Roussel
1806
Niaceae Jülich
Peyronelina P.J. Fisher, J. Webster & D.F.
Kane 1976*
Psathyrellaceae Vilgalys, Moncalvo &
Redhead
Hormographiella Guarro & Gené 1992,
Coprinellus P. Karst. 1879
Rhacophyllus Berk. & Broome 1871,
anamorphic Coprinopsis P. Karst. 1881
(or Coprinus?)
Pleurotaceae Kühner
Antromycopsis Pat. & Trab. 1897,
anamorphic Pleurotus (Fr.) P. Kumm.
Nematoctonus Drechsler 1941, anamorphic
Hohenbuehelia Schulzer 1866
Tricholomataceae R. Heim ex Pouzar
Nothoclavulina Singer 1970, anamorphic
Arthrosporella Singer 1970
Tilachlidiopsis Keissl. 1924, anamorphic
Dendrocollybia R.H. Petersen &
Redhead 2001
Typhulaceae Jülich
Sclerotium Tode 1790, anamorphic
Typhula (Pers.) Fr. 1818
Agaricales, genera incertae sedis
Disporotrichum Stalpers 1984
Fibulochlamys A.I. Romero & Cabral
1989*
Mycosphere
Atheliales Jülich
Atheliaceae Jülich
Fibulomyces Jülich 1972, anamorphic
Taeniospora Marvanová 1977
Fibularhizoctonia G.C. Adams & Kropp
1996, anamorphic Athelia Pers. 1822
Sclerotium Tode 1790, anamorphic Athelia
Pers. 1822*
Taeniospora Marvanová 1977, anamorphic
Fibulomyces Jülich 1972
Auriculariales, genera incertae sedis
Oliveorhiza P. Roberts 1998, anamorphic
Oliveonia Donk 1958
Cantharellales Gäum
Botryobasidiaceae Jülich
Allescheriella Henn. 1897, anamorphic
Botryobasidium Donk 1931
Alysidium Kunze 1817, anamorphic
Botryobasidium Donk 1931
Haplotrichum Link 1824, anamorphic
Botryobasidium Donk 1931
Ceratobasidiaceae G.W. Martin
Acanthellorhiza P. Roberts, anamorphic
Heteroacanthella 1990
Ceratorhiza R.T. Moore 1987, anamorphic
Ceratobasidium D.P. Rogers 1935*
Rhizoctonia DC. 1805 (and as Thanatophytum Nees 1816), anamorphic Helicobasidium Pat. 1885 and Thanatephorus Donk 1956
Hydnaceae Chevall.
Burgoa Goid. 1938, anamorphic Sistotrema Fr. 1821
Ingoldiella D.E. Shaw 1972, anamorphic
Sistotrema Fr. 1821
Osteomorpha G. Arnaud ex Watling &
W.B. Kendr. 1979, anamorphic Sistotrema Fr. 1821
Tulasnellaceae Juel
Epulorhiza R.T. Moore 1987, anamorphic
Tulasnella J. Schröt. 1888
Cantharellales, genera incertae sedis
Minimedusa Weresub & P.M. LeClair
1971*
Corticiales K.H. Larss.
Corticiaceae Herter
Chrysorhiza T.F. Andersen & Stalpers
1996, anamorphic Waitea Warcup &
P.H.B. Talbot 1962
Erythricium J. Erikss. & Hjortstam 1970
Marchandiomyces Dieder. & D. Hawksw.
1990, anamorphic Marchandiobasidium
Diederich & Schultheis 2003
Michenera Berk. & M.A. Curtis 1868,
anamorphic Licrostroma P.A. Lemke
1964
Tretopileus B.O. Dodge 1946
Hymenochaetales Oberw.
Schizoporaceae Jülich
Echinodia Pat. 1918, anamorphic Echinoporia Ryvarden 1980
Hymenochaetales, genera incertae sedis
Caeruleomyces Stalpers 2000
Polyporales Gäum.
Fomitopsidaceae Jülich
Ptychogaster Corda 1838, anamorphic
Oligoporus Bref. 1888
Sporotrichum Link 1809, anamorphic
Laetiporus Murrill 1904 and Pycnoporellus Murrill 1905
Ganodermataceae Donk
Thermophymatospora Udagawa, Awao &
Abdullah 1986, anamorphic Ganoderma P. Karst. 1881
Meruliaceae P. Karst.
Aegerita Pers. 1801, anamorphic Bulbillomyces Jülich 1974
Bornetina L. Mangin & Viala 1903,
anamorphic Diacanthodes Singer 1945
Sporotrichopsis Staplers (2000), anamorphic Abortiporus Murrill
Phanerochaetaceae Jülich
Erythricium J. Erikss. & Hjortstam 1970,
anamorph of Phanerochaete P. Karst.
1889
Sporotrichum Link 1809, anamorphic
Phanerochaete P. Karst. 1889
Polyporaceae Fr. ex Corda
Digitellus Paulet 1791, anamorphic Lentinus Fr. 1825
53
Mycelithe Gasp. 1841, anamorphic Polyporus P. Micheli ex Adans. 1763
Pachyma Fr. 1822, anamorphic Lentinus
Fr. 1825, Macrohyporia I. Johans. &
Ryvarden 1979, Polyporus P. Micheli
ex Adans. 1763 and Wolfiporia Ryvarden & Gilb. 1984
Russulales Kreisel ex P.M. Kirk, P.F. Cannon
& J.C. David
Bondarzewiaceae Kotl. & Pouzar
Spiniger Stalpers 1974, anamorphic Heterobasidion Bref. 1888
Echinodontiaceae Donk
Spiniger Stalpers 1974, anamorphic Laurilia Pouzar 1959
Lachnocladiaceae D.A. Reid
Spiniger Stalpers 1974,
Dichostereum Pilát 1926,
anamorphic
Stereaceae Pilát
Aleurocystis Lloyd ex G. Cunn. 1956,
Acaromyces Boekhout, Scorzetti, Gerson
& Sztejnb. 2003
Aleurocystis Lloyd ex G. Cunn. 1956,
anamorphic Licrostroma P.A. Lemke
1964
Matula Massee 1888, anamorphic Aleurocystis Lloyd ex G. Cunn.1956
Sebacinales M. Weiss, Selosse, Rexer, A. Urb.
& Oberw.
Sebacinaceae K. Wells & Oberw.
Craterocolla Bref. 1888
Ditangium P. Karst. 1867, anamorphic
Craterocolla Bref. 1888
Chaetospermum Sacc. 1892, ?anamorphic
Efibulobasidium K. Wells 1975*
Opadorhiza T.F. Andersen & R.T. Moore
1996, anamorphic Sebacina Tul. & C.
Tul. 1871
Thelephorales Corner ex Oberw.
Thelephoraceae Chevall.
Parahaplotrichum W.A. Baker & Partr.
2001, anamorphic Botryobasidium Rick
1959
Trechisporales K.H. Larss.
Hydnodontaceae Jülich
54
Aegeritina Jülich 1984, anamorphic
Subulicystidium Parmasto 1968
Agaricomycetes, genera incertae sedis
Akenomyces G. Arnaud ex D. Hornby
1984
Arthrodochium R.F. Castañeda & W.B.
Kendr. 1990
Arualis Katz 1980
Cenangiomyces Dyko & B. Sutton 1979
Corticomyces A.I. Romero & S.E. López
1989
Cruciger R. Kirschner & Oberw. 1999
Dendrosporomyces Nawawi, J. Webster &
R.A. Davey 1977
Ellula Nag Raj 1980
Fibulocoela Nag Raj 1978
Fibulotaeniella Marvanová & Bärl. 1988
Geotrichopsis Tzean & Estey 1991
Gloeosynnema Seifert & G. Okada 1988
Glomerulomyces A.I. Romero & S.E.
López 1989
Glutinoagger Sivan. & Watling 1980
Myriococcum Fr. 1823
Nyctalina G. Arnaud 1952
Pagidospora Drechsler 1960
Pycnovellomyces R.F. Castañeda 1987
Riessia Fresen. 1852
Riessiella Jülich 1985
Titaeella G. Arnaud ex K. Ando & Tubaki
1985
Tricladiomyces Nawawi 1985
Class Dacrymycetes Doweld
Dacrymycetales Henn.
Dacrymycetaceae J. Schröt.
Cerinosterus R.T. Moore 1987
Dacryoscyphus R. Kirschner & Zhu L.
Yang 2005
Class Tremellomycetes Hibbett, Matheny, &
Manfr. Binder
Cystofilobasidiales
Fell,
Roeijmans
&
Boekhout
Cystofilobasidiaceae K. Wells & Bandoni
Guehomyces Fell & Scorzetti 2004
Itersonilia Derx 1948
Mrakiella Margesin & Fell 2008
Phaffia M.W. Mill., Yoney. & Soneda
1976, anamorphic Xanthophyllomyces
Golubev 1995
Mycosphere
Rhodozyma Phaff, M.W. Mill., Yoney. &
Soneda 1972, anamorphic Xanthophyllomyces Golubev 1995
Tausonia Babeva 1998
Udeniomyces Nakase & Takem. 1992
Filobasidiales Jülich
Filobasidiaceae L.S. Olive
Cryptococcus Vuill. 1901, anamorphic
Filobasidium L.S. Olive 1968
Tremellales Fr.
Cuniculitremaceae J.P. Samp., R. Kirschner &
M. Weiss
Fellomyces Y. Yamada & I. Banno 1984
Kockovaella Nakase, I. Banno & Y.
Yamada 1991
Sterigmatosporidium G. Kraep. & U.
Schulze 1983, anamorphic Cuniculitrema J.P. Samp. & R. Kirschner 2001
Hyaloriaceae Lindau
Helicomyxa R. Kirschner & Chee J. Chen
2004
Tremellaceae Fr.
Bullera Derx 1930
Bulleribasidium J.P. Samp., M. Weiss & R.
Bauer 2002
Bulleromyces Boekhout & Á. Fonseca
1991
Cryptococcus Vuill. 1901, anamorphic
Filobasidiella Kwon-Chung 1976
Hormomyces Bonord. 1851, anamorphic
Tremella Pers. 1794
Hormomyces-like anamorphic Biatoropsis
Räsänen 1934
Tsuchiyaea Y. Yamada, H. Kawas., Itoh, I.
Banno & Nakase 1988
Trichosporonaceae Nann.
Asterotremella Prillinger, Lopandic &
Sugita 2007
Trichosporon Behrend 1890
Tritirachium-like, anamorphic Neorehmia
Höhn. 1902
Tremellales genera incertae cedis
Derxomyces F.Y. Bai & Q.M. Wang 2008
Hannaella F.Y. Bai & Q.M. Wang 2008
Tremellina Bandoni 1986
Tremellomycetes, genera incertae sedis
Heteromycophaga P. Roberts 1997
Moniliella Stolk & Dakin 1966
Trichosporonoides Haskins & J.F.T.
Spencer 1967
AGARICOMYCOTINA, genera incertae sedis
Microstella K. Ando & Tubaki 1984
Subphylum
PUCCINIOMYCOTINA
R.
Bauer, Begerow, J.P. Samp., M. Weiss &
Oberw
Class Agaricostilbomycetes R. Bauer, Begerow, J.P. Samp., M. Weiss & Oberw.
Agaricostilbales Oberw. & R. Bauer
Agaricostilbaceae Oberw. & R. Bauer
Bensingtonia Ingold 1986
Sterigmatomyces Fell 1966
Chionosphaeraceae Oberw. & Bandoni
Kurtzmanomyces Y. Yamada, Itoh, H.
Kawas., I. Banno & Nakase 1989
Mycogloea L.S.Olive 1950
Class Atractiellomycetes R. Bauer, Begerow,
J.P. Samp., M. Weiss & Oberw.
Atractiellales Oberw. & Bandoni
Phleogenaceae Gäum.
Basidiopycnides J. Reid, Eyjólfsd. &
Georg Hausner 2008, anamorphic Basidiopycnis Oberw., R. Kirschner, R.
Bauer, Begerow & Arenal 2006*
Saccoblastiaceae Jülich
Infundibura Nag Raj & W.B. Kendr. 1981
Atractiellales, genera incertae sedis
Hobsonia Berk. ex Massee 1891
Leucogloea R. Kirschner 2004
Class Classiculomycetes R. Bauer, Begerow,
J.P. Samp., M. Weiss & Oberw.
Classiculales R. Bauer, Begerow, Oberw. &
Marvanová
Classiculaceae Bauer, Begerow, Oberw. &
Marvanová
Jaculispora H.J. Huds. & Ingold 1960
Naiadella Marvanová & Bandoni 1987,
anamorphic Classicula R. Bauer, Begerow, Oberw. & Marvanová 2003
55
Class Cryptomycocolacomycetes R. Bauer,
Begerow, J.P. Samp., M. Weiss & Oberw.
Cryptomycocolacales Oberw. & R. Bauer
Cryptomycocolacaceae Oberw. & R. Bauer
Colacosiphon R. Kirschner, R. Bauer &
Oberw. 2001
Class Cystobasidiomycetes R. Bauer, Begerow,
J.P. Samp., M. Weiss & Oberw.
Erythrobasidiales, genera incertae sedis
Bannoa Hamam. 2002
Erythrobasidium Hamam., Sugiy. & Komag. 1988
Class Microbotryomycetes R. Bauer, Begerow,
J.P. Samp., M. Weiss & Oberw.
Heterogastridiales Oberw. & R. Bauer
Heterogastridiaceae Oberw. & R. Bauer
Hyalopycnis Höhn. 1918, anamorphic
Heterogastridium Oberw. & R. Bauer
1990
Leucosporidiales J.P. Samp., M. Weiss & R.
Bauer
Leucosporidiaceae Jülich
Leucosporidiella Samp. 2003
Microbotryales R. Bauer & Oberw., genera
incertae sedis
Reniforma Pore & Sorenson 1990
Sporidiobolales Doweld
Sporidiobolaceae R.T. Moore
Blastoderma B. Fisch. & Brebeck 1894,
anamorphic Sporidiobolus Nyland 1950
Rhodomyces Wettst. 1885, anamorphic
Sporidiobolus Nyland 1950
Sporobolomyces Kluyver & C.B. Niel
1924, anamorphic Sporidiobolus Nyland 1950
Sporidiobolales, genera incertae cedis
Ballistosporomyces Nakase, G. Okada &
Sugiy., in Nakase, Okada, Sugiyama, Itoh &
Suzuki 1989
Erythrobasidium Hamam., Sugiy. &
Komag. 1988, anamorphic Rhodosporidium Banno 1967
Rhodotorula F.C. Harrison 1927, anamorphic Rhodosporidium Banno 1967
Microbotryomycetes, genera incertae sedis
56
Crucella Marvanová & Suberkr. 1990,
anamorphic Camptobasidium Marvanová & Suberkr. 1990
Class Pucciniomycetes R. Bauer, Begerow, J.P.
Samp., M. Weiss & Oberw.
Helicobasidiales R. Bauer, Begerow, J.P.
Samp., M. Weiss & Oberw.
Helicobasidiaceae P.M. Kirk
Thanatophytum Nees 1816, anamorphic
Helicobasidium Pat. 1885
Tuberculina Tode ex Sacc. 1880, anamorphic Helicobasidium Pat. 1885
Platygloeales R.T. Moore
Eocronartiaceae Jülich
Glomopsis D.M. Hend. 1961, anamorphic
Herpobasidium Lind 1908
Uncolaceae Buriticá
Calidion Syd. & P. Syd. 1919
Pucciniales Clem. & Shear
Coleosporiaceae Dietel
Chrysomyxa Unger 1840, ?anamorphic
Peridermium (Link) J.C. Schmidt &
Kunze 1817
Cronartiaceae Dietel
Cronartium Fr. 1815, ?anamorphic
Peridermium (Link) J.C. Schmidt &
Kunze 1817
Phakopsoraceae Cummins & Hirats. f.
Aeciure Buriticá & J.F. Hennen 1994,
anamorphic Arthuria H.S. Jacks
Milesia F.B. White 1878, anamorphic
Phakopsora Dietel 1895 but see
Milesina Magnus 1909*
Macabuna Buriticá & J.F. Hennen 1994
Malupa Y. Ono, Buriticá & J.F. Hennen
1992
Physopella Arthur 1906, anamorphic Cerotelium Arthur 1906 and Phakopsora
Dietel 1895*
Uredendo Buriticá & J.F. Hennen 1994,
anamorphic Phakopsora Dietel 1895
Uredostilbe Buriticá & J.F. Hennen 1994
Phragmidiaceae Corda
Gerwasia Racib. 1909, anamorphic Campanulospora Salazar-Yepes, Pardo-
Mycosphere
Card. & Buriticá 2007, Morispora
Salazar-Yepes, Pardo-Card. & Buriticá
2007 and Scutelliformis Salazar-Yepes,
Pardo-Card. & Buriticá 2007
Physonema Lév. 1847, anamorphic Phragmidium Link 1815
Puccineaceae Chevall.
Caeoma Link 1809
Roestelia Rebent. 1804, anamorphic Gymnosporangium R. Hedw. ex DC. 1805
Pucciniastraceae Gäum. ex Leppik
Milesia F.B. White 1878, anamorphic
Milesina Magnus 1909, but see Phakopsora Dietel 1895*
Peridiopsora Kamat & Sathe 1969,
anamorphic Milesina Magnus 1909
Pomatomyces Oerst. 1864, anamorphic
Thekopsora Magnus 1875
Pucciniastrum G.H. Otth 1861, ?anamorphic Peridermium (Link) J.C.
Schmidt & Kunze 1817
Uropyxidaceae Cummins & Y. Hirats.
Canasta A.A. Carvalho & J.F. Hennen
2010, anamorphic Prospodium Arthur
1907*
Pucciniales, genera incertae sedis
Elateraecium Thirum., F. Kern & B.V.
Patil 1966, anamorphic Hiratsukamyces
Thirum., F. Kern & B.V. Patil 1975
Intrapes J.F. Hennen & Figueiredo 1979
Uraecium Arthur 1933
Uredo Pers. 1801
Septobasidiales Couch ex Donk
Septobasidiaceae Racib.
Johncouchia S. Hughes & Cavalc. 1983,
anamorphic Septobasidium Pat. 1892
Subphylum USTILAGINOMYCOTINA R.
Bauer, Begerow, J.P. Samp., M. Weiss &
Oberw.
Class Exobasidiomycetes Begerow, M. Stoll,
R. Bauer
Exobasidiomycetes, orders incertae sedis
Doassansiales R. Bauer & Oberw.
Doassansiaceae R.T. Moore ex P.M. Kirk, P.F.
Cannon & J.C. David
Savulescuella Cif. 1959,
Doassansia Cornu 1883
anamorphic
Entylomatales R. Bauer & Oberw.
Entylomataceae R. Bauer & Oberw.
Entylomella Höhn. 1924, anamorphic
Entyloma de Bary 1874
Tilletiopsis Derx 1948, anamorphic Entyloma de Bary 1874 and Melanotaenium
de Bary 1874
Microstromatales R. Bauer & Oberw.
Microstromataceae Jülich
Sympodiomycopsis Sugiy., Tokuoka &
Komag. 1991
Class Ustilaginomycetes R. Bauer, Oberw. &
Vánky
Urocystidales R. Bauer & Oberw.
Glomosporiaceae Cif.
Rhombiella Liro 1939, anamorphic Thecaphora Fingerh. 1836
Thecaphorella H. Scholz & I. Scholz 1988,
anamorphic Thecaphora Fingerh. 1836
Ustilaginales G. Winter
Anthracoideaceae Denchev
Crotalia Liro 1938, anamorphic Anthracoidea Bref. 1895
Cintractiellaceae Vánky
Naiadella Marvanová & Bandoni 1987,
anamorphic Classicula R. Bauer,
Begerow, Oberw. & Marvanová 2003
Ustilaginaceae Tul. & C. Tul.
Pseudozyma Bandoni 1985
Ustilaginomycetes, orders incertae sedis
Urocystidiales R. Bauer & Oberw.
Urocystidaceae Begerow, R. Bauer, & Oberw.
Paepalopsis J.G. Kühn 1882, anamorphic
Urocystis Rabenh. ex Fuckel 1870
USTILAGINOMYCOTINA,
sedis
Malasseziales R.T. Moore
Malassezia Baill. 1889
order
incertae
BASIDIOMYCOTA, class incertae sedis
Wallemiomycetes Zalar, de Hoog & Schroers
Wallemiales Zalar, de Hoog & Schroers
57
Wallemiaceae R.T. Moore
Wallemia Johan-Olsen 1887
BASIDIOMYCOTA, genera incertae sedis
Anastomyces W.P. Wu, B. Sutton & Gange
1997
Anguillomyces Marvanová & Bärl. 2000
Arcispora Marvanová & Bärl. 1998
Discussion
The study reveals that there are approximately 2873 anamorphic genera names of
which 699 genera and 94 anamorph-like genera
are linked to teleomorphic genera names, 447
are linked to teleomorph families, orders or
classes, while for about 1728 (60.25%) genera
no teleomorph link is known.
The data used in this compilation are
mostly from Index Fungorum, Species Fungorum and Kirk et al. (2008), plus publications
during 2009 and 2010 that were available
before the manuscript was sent to press. Thus it
is not inclusive and it is planned to improve the
compilation in future issues. Take the case of
Hysteriaceae in which four anamorphic genera
are listed (Acrogenospora, Aposphaeria, Plenodomus, Septonema). Boehm et al. (2009a)
however state that the coelomycetous pycnidial
states (e.g. Aposphaeria, Hysteropycnis) and
dematiaceous hyphomycetous anamorphs (e.g.
Coniosporium, Septonema, Sirodesmium, Sphaeronaema and Sporidesmium) have been
described for this family. Four anamorphs are
listed for Mytilinidiaceae in this compilation
(Camaroglobulus, Chalara-like, Papulasporalike, Septonema-like), however Boehm et al.
(2009a) state that anamorphic states in the
family are primarily coelomycetous (e.g. Aposphaeria, Camaroglobulus, Dothiorella, Pyrenochaeta, Sclerochaeta) and less frequently
hyphomycetous (e.g. Chalara-like, Papulaspora, Peyronelia, Septonema). It is obvious
that similar linkages will be missing from other
entries. However, many anamorph/teleomorph
conections were recorded based on appearance
of the two states in close proximity on a
substate and the connection has not inequiviqually been proven. It was also difficult
to locate all new genera, especially data on
anamorphic placement between the publication
of Kirk et al. (2008) and literature published
before 2009. We hope to incorporate any
58
missing data in future issues. In future issues
we will also examine the literature from which
connections are extracted to establish their
basis.
Although we have used data from peer
reviewed published data we have no way to
confirm the validity of the species names used
in various phylogenetic analyses. Where generic types were sequenced we have stated this,
but again it is not clear whether correct names
have been applied to these taxa or whether they
are strains of type material (in most cases they
are not). Because of their wide ranging occurrence as anamorphic ascomycota, we have only
dealt with Phoma species when publications
specifically resolve this genus.
The naming of fungi has long been
problematic because of the dual nomenclature
system. The system was introduced because it
was previously often impossible to link a
sexual form with the asexual form and therefore there was a need to be able to name both
forms (Shenoy et al. 2007). There are extensive
discussions on this topic, which will not be
repeated here, but they generally agree that the
dual nomenclature system is becoming redundant and should be replaced with a modern
system using one name for a biological species
(Shenoy et al. 2007, 2010). With the advent of
the molecular era it is often possible to link the
anamorph with the teleomorph, and if there is
no obvious teleomorph it is becoming increasing feasible to establish the relationship of the
anamorphic genus within the teleomorph taxonomic framework.
There are presently at least four schools
of though on how anamorphic genera and species names should be dealt with in the future
and the above compilation helps to reveal some
of the problems associated with any approach.
The first school believes that only one name
should be used for a biological species and that
this name should follow the sexual state since
this state has more biological significance.
Although this argument is compelling the mycological community must deal with important
asexual genera that are very well known to the
wider scientific community (e.g. Aspergillus,
Penicillium, Trichoderma), which would disappear from usage, and perhaps make a ridicule
of fungal taxonomic approaches. This could be
overcome by conserving some of the important
Mycosphere
anamorph or teleomorph names. A second
school believes that the earliest introduced
name should be used. Again this idea has merit
but may mean that some well known genera
would disappear from usage; however conservation of certain names could also be applied to
solve this problem. The third school favour
status quo, and again there is merit in keeping a
system that does work, but is confusing to most
within and outside the fungal taxonomic community. A fourth school has started to adopt
these approaches. Several authors have described new teleomorphic genera and do not
give names to the asexual states but describe
them as part of the teleomorph species. The
problem with this approach is that it is hard for
researchers to link a characteristic anamorph
(i.e. those with a specific set of features) with a
teleomorph genus and such data does not get
entered in databases; thus we have tried to
include these in this compilation. Other authors
have introduced new anamorphic species and
described the teleomorph state. This approach
has generally been adopted where the anamorphic name is better known to the scientific
community or where the teleomorph name is
polyphyletic. Whichever of the above approaches is most appropriate requires considerable
debate and is not elaborated on here as it is not
the purpose of the compilation. It will be
several years before a consensus is reached;
however, there are moves ahead to promote
various approaches and the next few years will
be interesting and perhaps controversial
There has been a long tradition of not
naming the anamorphic stages in the Basidiomycota (Kirschner & Oberwinkler 2009),
however many have been named and are listed
in the compilation.
Notes (1-307)
Notes are provided on recent published
data on anamorphic taxa. Notes are provided if
new data is published in relation to the phylogeny, if changes in anamorph or teleomorph
linkages have been revealed and if new genera
are introduced. If well known connections are
confirmed we may provide data from one or
more publications, however, in many cases we
did not cite these data as the known connections are unchanged.
Acremonium Link (1)
In a survey of mycotic skin and nail
lesions from humans in Chiriquí, Western
Panama, a darkly pigmented Acremonium-like
taxon was isolated. Morphological investigation and phylogenetic analysis based on LSU
rDNA and ITS sequences revealed that this
species represents an undescribed member of
the Plectosphaerellaceae. The species is described as Acremonium collariferum because of
its similarities to other Acremonium species
and its distinct collarette (Weisenborn et al.
2010).
Acremonium-like (2)
Plishka et al. (2009) reported an Acremonium anamorph for Nigrosabulum globosum
Malloch & Cain of Bionectriaceae, while
Lechat & Courtecuisse (2010) described a new
species of Ijuhya with an Acremonium-like
anamorph.
Acephala Grünig & T.N. Sieber (3)
Grünig et al. (2009) found that two
species clustered near Vibrissea (Vibrisseaceae)
while one unnamed species clustered near
Loramyces showing the polyphyletic nature of
this genus.
Akanthomyces Lebert (4)
These spider pathogens formed a well
supported subclade but were not linked to
Torrubiella (Johnson et al. 2009).
Ambrosiella Brader ex Arx & Hennebert (5)
Multigene phylogenetic analysis showed
this genus to have Ophiostoma and Grosmannia anamorphs (Massoumi Alamouti et al.
2009). Five species of Ambrosiella clustered in
a novel clade within Ophiostoma, while two
species clustered with Raffaelea and Dryadomyces and were related to Grosmannia.
Ampelomyces Ces. ex Schltdl. (6)
One species of this genus clustered in
Didymellaceae following multigene phylogenetic analysis (Averkamp et al. 2010).
Amplistroma Huhndorf, A.N. Mill., M. Greif
& Samuels (7)
59
Huhndorf et al. (2009) described this
teleomorphic genus in Amplistromataceae with
Acrodontium-like anamorphs.
Aposphaeria Sacc. (14)
Aveskamp et al. (2010) confirmed one
species belonged in Melanommataceae following multigene analysis.
Ampullicephala R.F. Castañeda, Minter & M.
Stadler (8)
Castañeda Ruiz et al. (2009a) described
this new genus to accommodate Pleurotheciopsis setiformis R.F. Castañeda. No teleomorph relationship is known.
Aposphaeria-like (15)
Boehm et al. (2009a) described Gloniopsis and the new teleomorph genus Hysterobrevium as having Aposphaeria-like anamorphs.
Anaselenosporella Heredia, R.F. Castañeda &
R.M. Arias (9)
Castañeda Ruiz et al. (2010a) introduced
this new monotypic hyphomycete genus from
Mexico. No teleomorph relationship is known.
Aquaticheirospora Kodsueb & W.H. Ho (16)
Shearer et al. (2009) showed the generic
type to be anamorphic Massarinaceae based on
phylogenetic analyses of SSU and/or LSU
genes.
Anguillospora Ingold (10)
Shearer et al. (2009) showed the generic
type to be anamorphic Amniculicolaceae based
on phylogenetic analyses of SSU and/or LSU
genes.
Arthrobotrys Corda (17)
Li et al. (2009) described a new member
of this genus as anamorphic Orbilia.
Antennariella Bat. & Cif. (11)
Crous et al. (2009a) showed one species
in this genus to be a member of Capnodiaceae
in a phylogeny derived from partial LSU genes
indicating that the anamorphic genus may be
polyphyletic as it has already been linked with
Antennulariella in Antennulariellaceae. Cheewangkoon et al. (2009) introduced a new
species in the genus in Capnodiales based on
LSU gene analysis.
Anthonectria Döbbeler (12)
Döbbeler (2010) introduced a new
teleomorphic genus from the developing sporophytes of Drepanolejeunea sp. which had a
pulvinate anamorphic state with ramifying
cylindrical conidiogenous cells and small
globose colourless conidia.
Aplosporella Speg. (13)
Taylor et al. (2009) described a new
species from Australia. The teleomorph is
unknown, however some species clustered
basal to Botryosphaeriaceae. This genus has
over 300 species and appears to be heterogenous; therefore not all species are likely to
belong in Botryosphaeriaceae.
60
Aschersonia Mont. (18)
Confirmed as anamorphic Moelleriella
(Tadych et al. 2009) and anamorphic Hypocrella (Mongkolsamrit et al. 2009).
Ascochyta Lib. (19)
Ascochyta pisi Lib. was linked to the
teleomorph Didymella pisi Chilvers, J.D.
Rogers & Peever (Chilvers et al. 2009).
Aveskamp et al. (2010) noted the polyphyletic
nature of this genus.
Aspergillus P. Micheli ex Link (20)
Two species of teleomorphic Neosartorya with anamorphs in Aspergillus section
Fumigati were described by Yaguchi et al.
(2010).
Aureobasidium Viala & G. Boyer (21)
Suetrong et al. (2009) showed one
species to be anamorphic Dothideaceae based
on phylogenetic analyses of four genes.
Asterosporium Kunze (22)
Tanaka et al. (2010) transferred one
species in this genus to Prosthemium and found
that another species lay in Diaporthales. The
type species was not used in this study.
Mycosphere
Aurosphaeria Sun J. Lee, Strobel, Eisenman,
Geary, Vargas & S.A. Strobel (23)
Lee et al. (2009) described this new
endophytic pycnidial coelomycetous genus
from the Bolivian Amazon. Molecular data
provided no clues for the placement of this
taxon other than in the Ascomycota.
Bagadiella Cheewangkoon & Crous (24)
Cheewangkoon et al. (2009) introduced
this genus for a single species from Eucalyptus.
The phylogenetic placement is not clear
although it was related to Plectosphaera in
Phyllachoraceae. The authors state that Bagadiella is similar to Cladorrhinum and is related
to Apiosordaria in Lasiosphaeriaceae and we
have followed the latter classification here.
Bahusutrabeeja Subram. & Bhat (25)
Shenoy et al. (2010) found the type
species is phylogenetically related to Neodeightonia (Botryosphaeriaceae).
Barriopsis A.J.L. Phillips, A. Alves & Crous
(26)
Abdollahzadeh et al. (2009) describe a
second species in the genus and show this and
the generic type to be anamorphic Botryosphaeriaceae based on molecular sequence
analysis.
Basidiopycnides J. Reid, Eyjólfsd. & Georg
Hausner (27)
Kirschner & Oberwinkler (2009) considered this monotypic genus to be identical with
the hyphomycete anamorph of Basidiopycnis,
which is also monotypic.
Batcheloromyces Marasas, P.S. van Wyk &
Knox-Dav. (28)
Crous et al. (2009b) showed the generic
type and one species clustered in Clade 2 of
Teratosphaeriaceae, while one putative species
clustered in Clade 4. It has a Catenulostroma
synanamorph (Crous 2009).
Baudoinia J.A. Scott & Unter. (29)
This is anamorphic Teratosphaeriaceae
(Crous et al. 2009a,d) based on molecular
sequence analysis.
Beverwykella Tubaki (30)
Zhang et al. (2009) showed the generic
type to be anamorphic Melanommataceae
based on phylogenetic analyses of five genes.
Bhatia W.A. Baker & Morgan-Jones (31)
Gams et al. (2009) introduced this new
hyphomycete genus to accommodate a species
of Acrodictys which differs from other genera
in the Acrodictys complex. No teleomorph
relationship is known.
Boeremia Aveskamp, Gruyter& Verkley (32)
This genus was introduced by Aveskamp
et al. (2010) to accommodate ten species and
ten varieties mostly based on combination of
Phoma species and varieties and is accommodated in Didymellaceae.
Brachiosphaera Nawawi (33)
Shearer et al. (2009) conducted molecular sequence-based phylogenetic analyses using
nuclear ribosomal sequences (SSU and/or LSU)
and showed this to be anamorphic Aliquandostipitaceae, which comprise only freshwater
species.
Bruneosphaerella Crous (35)
This teleomorphic genus was introduced
by Crous et al. (2009a) and is a member of
Mycosphaerellaceae in a phylogeny derived
from partial LSU gene analysis; the anamorph
was stated to be Coniothyrium-like and possibly Phaeophloeospora.
Brycekendrickomyces Crous & M.J. Wingf.
(35)
Introduced by Crous et al. (2009d) to
accommodate a new hyphomycete genus from
Indonesia which clustered in Herpotrichiellaceae based on LSU sequence analysis.
Cadophora Lagerb. & Melin (36)
Nekoduka et al. (2010) found that three
species of this genus clustered in Helotiales
following ITS gene sequence analysis.
Cadophora-like (37)
Grünig et al. (2009) found this to be the
anamorph of Phaeomollisia. However, they
could not distinguish clearly between Cadophora and Phialocephala reporting intermediate forms.
61
Canalisporium Nawawi & Kuthub. (38)
Sri-indrasutdhi et al. (2010) described the
new teleomorph genus Ascothailandia which
produced a Canalisporium anamorph. Phylogenetic analysis of 18S and 28S rDNA sequences showed Ascothailandia, the generic
type and nine other Canalisporium species to
belong in Hypocreomycetidae.
Canasta A.A. Carvalho & J.F. Hennen (39)
This anamorphic genus (Carvalho &
Hennen 2010) was described to accommodate
species closely related to Prospodium (Uropyxidaceae)
Capnobotryella Sugiy. (40)
Crous et al. (2009a,d) confirmed the
generic type to be anamorphic Teratosphaeriaceae based on molecular sequence analysis.
Catenulifera Hosoya (41)
Bogale et al. (2010) confirmed this genus
to be anamorphic Hyphodiscus with all species
analysed clustering in a monophyletic clade
with morphologically similar species of Phialophora, with the exception of P. hyalina. Two
species of Phialophora were transferred to
Catenulifera and one new species was described. The phylogenetic positions of Catenulifera and Hyphodiscus within the Helotiales
could not be resolved.
Catenulostroma Crous & U. Braun (42)
Crous et al. (2009a,b) confirmed the
generic type and five other species to be
anamorphic Teratosphaeriaceae based on
partial LSU gene analysis.
Cercospora Fresen. (43)
Crous et al. (2009a,d) showed seven
species in this genus to be members of Mycosphaerellaceae in a phylogeny derived from
SSU, 5.8S rDNA and LSU analyses.
Cercosporella Sacc. (44)
Kirschner (2009) showed species in this
genus to cluster in Mycosphaerellaceae with
one species close to Ramularia and others
close to Mycosphaerella and Pseudocercospora. Crous et al. (2009a,d) showed one
species in this genus to be a member of
62
Mycosphaerellaceae in a phylogeny derived
from SSU, 5.8S rDNA and LSU analyses.
Chaetopsina Rambelli (45)
Luo & Zhang (2010) introduced a new
teleomorphic genus Chaetopsinectria, which is
distinct from Cosmospora and has Chaetopsina
anamorphs.
Chaetosphaeronema Moesz (46)
Zhang et al. (2009) showed one species
to be anamorphic Phaeosphaeriaceae based on
phylogenetic analyses of five genes.
Cheirosporium L. Cai & K.D. Hyde (47)
Shearer et al. (2009) showed the generic
type to be anamorphic Massarinaceae based on
phylogenetic analyses of SSU and LSU genes.
Chrysosporium Corda (48)
Nováková & Kolařik (2010) described a
new species of this genus in Onygenales from
bat guano in caves. They showed by rDNA
sequence analysis that the species was possibly
a relative of Renispora, while C. chiropterorum
Beguin & Larcher was linked to Polytolypa.
Cibiessia Crous (49)
Crous et al. (2009b) showed that two
strains of the generic type and two other species clustered with Readeriella clade of
Teratosphaeriaceae.
Cladophialophora Borelli (50)
Koukol (2010) isolated a strain of this
genus from pine litter which was treated as a
new species, C. matsushimae Koukol. A 28S
rDNA data matrix showed this strain to be
basal in Cladophialophora confirming the
polyphyly of the genus.
Chaetospermum Sacc. (51)
Kirschner & Oberwinkler (2009) found
evidence to support the relationship between
this genus and the basidiomycete Efibulobasidium.
Chalara (Corda) Rabenh. (52)
Phyogenetic relationships of 26 strains of
Chalara and allied genera were investigated by
analysis of rDNA (Cai et al. 2009a). Most
Mycosphere
Chalara strains clustered in a strongly supported monophyletic lineage across a wide
range of Helotiales, while a few strains seemed
close to Xylariales, however, the generic type
strain was not used in this study. They also
suggest that the phenotypes of Chalara-like
species have multiple evolutionary origins.
Chaetasbolisia Speg. (53)
Aveskamp et al. (2010) found the generic
type to cluster in Didymellaceae in a multigene
phylogenetic analysis.
Chaetochalara B. Sutton & Piroz. (54)
Cai et al. (2009a) showed this genus to be
phylogenetically inseparable from Chalara
with strains clustering in Helotiales and Xylariales, however, they did not use the generic
type strain in their study.
Chaetosphaeronema Moesz (55)
Gruyter et al. (2010) proposed a new
combination to this genus, which along with C.
hispidulum (Corda) Moesz, is anamorphic
Phaeosphaeriaceae.
Chalastospora E.G. Simmons (56)
Crous et al. (2009e) showed the generic
type and four other taxa in the genus to be
anamorphic Pleosporaceae based on analysis
of LSU sequence data.
Chasakopama Manohar., Bagyan., N.K. Rao
& Kunwar (57)
Introduced by Manoharachary et al.
(2009) for a new hyphomycete genus characterized by discrete, polyblastic, denticulate
conidiogenous cells and single campanulate
condia with two dark bands. No teleomorph
relationship is known.
Cheiroidea W.A. Baker & Morgan-Jones (58)
Gams et al. (2009) introduced this new
hyphomycete genus to accommodate a species
of Acrodictys primarily based on its peculiar
cheiroid conida. No teleomorph relationship is
known.
Chrysoporthe Gryzenh. & M.J. Wingf. (59)
Chungu et al. (2010) described two
anamorphic stem canker pathogens under the
teleomorph name Chrysoporthe even though
the teleomorph stage was not formed and is
unknown. They argued that it is unnessecary to
introduce these new species under the anamorphic name (Chrysoporthella) as this has
confused understanding of species in the genus.
To further ensure a stable and less confusing
taxonomy for Chrysoporthe, the authors combine the only anamorphic species (Chrysoporthella hodgesiana Gryzenh. & M.J. Wingf.)
linked to the genus as Chrysoporthe hodgesiana (Gryzenh. & M.J. Wingf.) Chungu,
Gryzenh. & M.J. Wingf. This encourages the
use of a one name system for the genus.
Chrysosporium Corda (60)
Liang et al. (2009a) described a new
species in the genus. ITS sequence data
confirmed that this and other species used in
the phylogenetic analysis have a close phylogenetic relationship with the teleomorph genus
Aphanoascus.
Cirrenalia Meyers & R.T. Moore (61)
The type species Cirrenalia macrocephala (Kohlm.) Meyers & R.T. Moore was
well placed in the Halosphaeriaceae while
other taxa were assigned to other genera
(Abdel-Wahab et al. 2010).
Cladosporium Link (62)
The generic type and two species in this
genus were shown to be members of Davidiellaceae in a phylogeny derived from five
genes (Schoch et al. 2009). Crous et al. (2009a)
also demonstrated that four species of Cladosporium were members of Davidiellaceae in a
phylogeny derived from three genes.
Claviradulomyces P.R. Johnst., D.C. Park, H.C.
Evans, R.W. Barreto & D.J. Soares (63)
Evans et al. (2010a) introduced this teleomorph new genus in Odontotremataceae for a
discomycete from Western Africa. A putative
pycnidial anamorph was consistently associated with the gall-like eruptions, often together
with the teleomorph. The conidia were morphologically similar to the ascospores but a
link was not confirmed as cultures were not
derived from single ascospores.
Collophora Damm & Crous (64)
Damm et al. (2010) introduced this new
coelomycetous genus with five new species
63
which they placed in Leotiomycetes based on
sequence analysis. The isolates were from
Prunus trees and resembled Lecythophora but
were distinguishable morphologically and phylogenetically.
Gams et al. (2009) validated this new
hyphomycete genus to accommodate a species
of Acrodictys primarily based on its peculiar
cheiroid conida. No teleomorph relationship is
known.
Colletogloeopsis Crous & M.J. Wingf. (65)
Crous et al. (2009b) showed by analysis
of LSU sequence data that six species clustered
in Clade 4 of Teratosphaeriaceae with Teratosphaeria sensu stricto.
Crucellisporium M.L. Farr (72)
Marincowitz et al. (2010) showed this to
be anamorphic Helotiales.
Colletotrichum Corda (66)
Cai et al. (2009b), Damm et al. (2009)
and Hyde et al. (2009) extensively reviewed
this genus with links to the Glomerella teleomorphs supported by molecular phylogenetic
analysis, while Prihastuti et al. (2009) and
Yang et al. (2009) described the Glomerella
teleomorph state for some species.
Conidioxyphium Bat. & Cif. (67)
Schoch et al. (2009) and Crous et al.
(2009a) showed one species in this genus to be
a member of Capnodiaceae in a phylogeny
derived from combined analysis of five genes
or from partial LSU genes.
Coniella Höhn. (68)
Cheewangkoon et al. (2010) confirmed
one species to be a member of Schizoparmaceae.
Coniosporium Link (69)
Sert & Sterflinger (2010) described a
new species of Coniosporium from historical
marble monuments which judged by SSU
phylogeny data belonged to Chaetothyriales
and was related to Mycocalicium Vain. ex
Reinke. Ruibal et al. (2009) found that two
species inhabiting rocks were anamorphic
Dothideomycetes indicating the genus is polyphyletic.
Coniothyrium Corda (70)
Aveskamp et al. (2010) used four species
in this genus in a multigene phylogeny of
Phoma species which clustered in four separate
clades indicating the polyphyletic nature of this
genus. The generic type clustered in the
Leptosphaeriaceae/Pleosporaceae clade.
Corynesporina Subram. (71)
64
Cryomyces Selbmann, de Hoog, Mazzaglia,
Friedmann & Onofri (73)
Ruibal et al. (2009) confirmed that two
species inhabiting rocks were anamorphic
Dothideomycetes.
Cryptosporiopsis Bubák & Kabát (74)
Cheewangkoon et al. (2010) described
two new species in the genus that they
considered belonging to Dermateaceae.
Cumulospora I. Schmidt (75)
Both species of this genus clustered in
Lulworthiales, Lulworthiaceae with Cumulospora marina I. Schmidt linked to an unknown
Lulworthia species and Cumulospora varia
Chatmala & Somrith. 2004 (as Moromyces
varius) grouping with Lulwoana uniseptata
(Nakagiri) Kohlm., Volkm.-Kohlm., J. Campb.,
Spatafora & Gräfenhan 2005 (Abdel-Wahab et
al. 2010).
Custingophora Stolk, Hennebert & Klopotek
(76)
Kolařik & Hulcr (2009) accepted a
broader concept for Custingophora when they
combined Knoxdaviesia. They described a new
species of Gondwanamyces and listed the
anamorph as Custingophora-like.
Cylindrocladium Morgan (77)
Lombard et al. (2010) provided a monograph of Calonectria and its Cylindrocladium
anamorph.
Cyphellophora G.A. de Vries (78)
Crous et al. (2009e) confirm the placement of this genus in Chaetothyriaceae based
on LSU gene sequence analysis.
Cystocoleus Thwaites (79)
Ruibal et al. (2009) showed the generic
type is anamorphic Teratosphaeriaceae.
Mycosphere
Cytospora Ehrenb. (80)
Fotouhifar et al. (2010) examined the
associated teleomorphs of this genus in Iran,
which clustered with Leucostoma and Valsa
species.
Diplodia Fr. (88)
Begoude (2010) studied Botryosphaeriaceae associated with Terminalia catappa
and phylogenetic analysis confirmed placement
of Diplodia.
Davisoniella H.J. Swart (81)
Crous (2009) list this as anamorphic
Teratosphaeriaceae.
Dissoconiaceae Crous & de Hoog (89)
This family was introduced in Capnodiales for taxa with Mycosphaerella-like teleomorphs and Dissoconium anamorphs based on
two-gene phylogenetic analyses using nucSSU
and nucLSU rDNA (Crous et al. 2009a).
Dematophora-like (82)
Takemoto et al. (2009) introduced a new
species of Rosellinia with a Dematophora-like
anamorph with synnemata and conspicuously
cicatrized and geniculate conidiophores.
Dendryphiella Bubák & Ranoj. (83)
Shearer et al. (2009), Suetrong et al.
(2009) and Zhang et al. (2009a) showed
species in this genus to be anamorphic Pleosporaceae based on phylogenetic analyses of
various genes.
Devriesia Seifert & N.L. Nick. (84)
Crous et al. (2009a,d) confirmed four
species to be anamorphic Teratosphaeriaceae
based on partial LSU gene analysis. Koukol
(2010) found a strain from pine litter that
clustered with D. americana Crous & Dugan in
Teratosphaeriaceae. Frank et al. (2010) described a new species in the genus which is
shown to be paraphyletic, and to represent
several lineages of which only Devriesia s.str.
is thermotolerant. They suggested that further
collections are required, before the latter generic complex can be resolved.
Dissoconium de Hoog, Oorschot & Hijwegen
(90)
Crous et al. (2009a,d) confirmed the
generic type and four species to be anamorphic
Dissoconiaceae based on partial LSU gene
analysis.
Dothiorella Sacc. (91)
Two new species are described in the
genus in a paper on Botryosphaeriaceae from
Australia. The teleomorph is stated as assumed
to be Dothidotthia based on phylogenetic
analysis (Taylor et al. 2009) and if this was the
case the genus should belong in Dothidothiaceae? Zhang et al. (2009b) described a new
species from Populus in China with a
Spencermartinsia teleomorph.
Dothistroma Hulbary (92)
Crous et al. (2009a,d) showed two
species in this genus to be members of
Mycosphaerellaceae in a phylogeny derived
from SSU, 5.8S rDNA and LSU analyses.
Diatrypasimilis J. Zhou & Kohlm. (85)
This new teleomorphic genus was described with a Selenosporella-like anamorph by
Chalkley et al. (2010).
Diplosporonema Höhn. (93)
This genus was described as new to
Turkey and it was noted to be the anamorph of
Pyrenopezia (Erdoǧdu & Hüseyin 2009)
Dictyosporium Corda (86)
Crous et al. (2009e) showed one species
in Pleosporaceae incertae sedis based on LSU
gene sequence analysis.
Drechslerella Subram. (94)
Yu et al. (2009) described a new species
in this genus as anamorphic Orbilia.
Diplococcium Grove (87)
Shenoy et al. (2010) referred the generic
type to Helotiales and a second species to
Pleosporales showing that the genus does not
represent a natural grouping.
Dryadomyces Gebhardt (95)
Massoumi Alamouti et al. (2009) showed
this is anamorphic Grosmannia and possibly
forms a monophyletic clade with some Ambrosiella species and Raffaelea.
65
Echinoconidiophorum
Pereira-Carv.
&
Dianese (96)
This new genus was introduced by
Pereira-Carvalho et al. (2009) to accommodate
a Brazilian hyphomycetous species growing on
living leaves of Gomidesia. No taxonomic
placement was given.
Edenia M.C. González, Anaya, Glenn,
Saucedo & Hanlin (97)
Crous et al. (2009d) showed the generic
type to be anamorphic Pleosporaceae based on
LSU gene data analysis.
Eladia G. Sm. (98)
Although this genus is regarded as a
synonym of Penicillium (Kirk et al. 2008),
Jiang & Zhang (2009) described two new
species in the genus.
Elotespora R.F. Castañeda & Heredia (99)
This remarkable new genus was introduced by Castañeda Ruiz et al. (2010b). It has
minute cupulate conidiomata in which a single
muriform conidium is formed. It was not linked
to any teleomorph.
Endogenospora R.F. Castañeda, O. Morillo &
Minter (100)
Castañeda Ruiz et al. (2010c) introduced
this new monotypic hyphomycete genus from
Venuzuala. No teleomorph relationship is
known.
Endothiella Sacc. 1906 (101)
Cheewangkoon et al. (2010) confirmed
one species in this genus to be Schizoparmaceae.
Epicoccum Link (102)
Aveskamp et al. (2010) reinstated this
genus to accommodate the generic type and
two Phoma species, which they formerly transferred. Multigene phylogenetic analysis showed the genus to lie in Didymellaceae.
Fecundostilbum Prameela & Chowdhry (103)
Devi (2009) introduced this new anamorphic genus with no known taxonomic placement.
Fibulochlamys A.I. Romero & Cabral (104)
66
Madrid et al. (2010a) described a new
species in this genus that is related to members
of gilled Agaricales.
Flahaultiella Seifert (105)
Gams et al. (2009) introduced this genus
for Flahaultia G. Arnaud 1951, which is
preoccupied by a red alga. They suspected this
is a mycoparasite and not the anamorph of
Sebacina sp., as had sometimes been thought,
as they were unable to culture it. No taxonomic
placement was given.
Foliocryphia Cheewangkoon & Crous (106)
Cheewangkoon et al. (2010) introduced
this monotypic genus that clustered within
Cryphonectriaceae based on phylogenetic analysis.
Fontanospora Dyko (107)
Two strains were included in molecular
phylogenetic analysis and clustered in Fontanospora clade 1 in Vasicosporium clade 1 in
Helotiales (Campbell et al. 2009).
Friedmanniomyces Onofri (108)
Schoch et al. (2009) showed that the two
known species in this genus are anamorphic
Teratosphaeriaceae in a phylogeny derived
from five genes.
Fumagospora G. Arnaud (109)
Crous et al. (2009a) showed one species
in this genus to be a member of Capnodiaceae
in a phylogeny derived from partial LSU gene
analysis.
Fusarium Link (110)
A new genus Cyanonectria was introduced to accommodate Nectria cyanostoma
Sacc. & Flageolet which has a Fusarium
anamorph (Samuels et al. 2009). The authors
consider that Fusarium morphology in not
monophyletic and state that “whether Fusarium
is dismantled into newly revived and described
genera, or whether they are referred to the
corresponding teleomorph name, the genus
name Fusarium in its current broad sense has
limited phylogenetic significance”. Schroers et
al. (2009) discovered a new lineage of Fusarium comprising the F. dimerum group that
Mycosphere
was not linked to any teleomorph but clustered
in Nectriaceae.
trichomes of living leaves of Eriosema No
taxonomic placement was given.
Fusicladium Bonord. (111)
Seyran et al. (2010) showed that the
teleomorph of F. effusum G. Winter was Venturia inaequalis (Cooke) G. Winter using
analysis of the mitochondrial cytochrome b
gene. Koukol (2010) also introduced three new
species of this genus in the Venturiaceae.
Gabarnaudia Samson & W. Gams (112)
The relationship with Sphaeronaemella
was confirmed by Jung et al. (2010).
Glomerulispora Abdel-Wahab, Abdel-Aziz &
Nagahama (118)
Abdel-Wahab et al. (2010) introduced
this genus for a marine hyphomycete from
Egypt with helicospores, which differs from
Moheitospora in smaller sized and larger
number of conidial cells. The taxa clustered
close to Torpedospora which may be the
teleomorph in the clade named TBM and was
included in the Hypocreales incertae sedis
along with this genus.
Geosmithia Pitt (113)
The characters of this genus appear in
several different genera of ambrosia fungi in
different lineages in Ophiostomatales (Raffaellea, Dryadomycyes), Micorascales (Ambrosiella) and Saccharomycetales and appears to
represent an optimal phenotype for this habit
(Kolařík & Kirkendall 2010). The new species
described in this paper represented new
lineages in Ascomycota.
Gnomoniopsis Berl. (119)
This status of this teleomorphic genus
was reconfirmed and pycnidial anamorphs
were confirmed in culture for some species
(Walker et al. 2010). The pycnidia range from
hyaline to cream, yellow or light brown, back
and even pink and conidia ranged from oval to
fusiform, globose, femur-shaped and lacked or
had varying numbers of guttules. Unfortunately
no illustrations were provided.
Gibellula Cavara (114)
Gibellula was linked to Torrubiella and
was shown to be specific to spiders in a
monophyletic clade (Johnson et al. 2009)
Goidanichiella G.L. Barron ex W. Gams (120)
Gams et al. (2009) validated this genus
which had been wrongly typified. The taxonomic placement is unknown.
Gliomastix Guég. (115)
Lechat et al. (2010) described a new
species of Hydropisphaera with a Gliomastix
anamorphic state. No teleomorph had previously been linked to Gliomastix and the genus can now be accomodated in Bionectriaceae.
Graphiopsis Trail (121)
Crous et al. (2009a) showed one species
in this genus to be a member of Davidiellaceae
in a phylogeny derived partial LSU genes.
Globoconidiopsis G.F. Sepúlveda, PereiraCarv. & Dianese (116)
This new genus was introduced by
Pereira-Carvalho et al. (2009) to accommodate
a Brazilian hyphomycetous species growing on
trichomes of living leaves of Brysonima. No
taxonomic placement was given.
Globoconidium G.F. Sepúlveda, Pereira-Carv.
& Dianese (117)
This new genus was introduced by
Pereira-Carvalho et al. (2009) to accommodate
a Brazilian hyphomycetous species growing on
Graphium Corda (122)
Grobbelaar et al. (2009) confirmed this to
be the anamorph of Ophiostoma, while Kolařík
& Hulcr (2009) gave details of a Graphium sp.,
but did not formally describe it.
Halazoon Abdel-Aziz, Abdel-Wahab &
Nagahama (123)
This genus was introduced for a marine
hyphomycete genus from Egypt with brown
helicoid conidia. Phylogenetic analysis showed
it to form a well-supported monophyletic group
along with Cirrenalia fusca I. Schmidt with
two Lulworthia species in the Lulworthiales,
Lulworthiaceae (Abdel-Wahab et al. 2010).
67
Halenospora E.B.G. Jones (124)
Jones et al. (2009a) introduced this new
marine hyphomycete genus to accommodate a
Zalerion species which grouped with Glarea
lozoyensis Bills & Peláez in Leotiaceae.
Halosigmoidea Nakagiri, K.L. Pang & E.B.G.
Jones (125)
Introduced by Jones et al. (2009b) to
accommodate species of Sigmoidea with Corollospora teleomorphs.
Haradamyces Masuya, Kusunoki, Kosaka &
Aikawa (126)
A new anamorphic genus was introduced
for a Myrioconium-like taxon causing zonate
leaf blight disease of Cornus florida in Japan
(Masuya et al. 2009). Molecular analysis
placed it in the Sclerotiniaceae, Leotiomycetes.
Helgardia Crous & W. Gams (127)
Nekoduka et al. (2010) found that two
species of this genus clustered in Helotiales
following ITS gene sequence analysis along
with the teleomorph Oculimacula.
Helminthosporiomyces
G.F.
Sepúlveda,
Pereira-Carv. & Dianese (128)
This new genus was introduced by
Pereira-Carvalho et al. (2009) to accommodate
a Brazilian hyphomycetous species growing on
trichomes of living leaves of Davilla. No
taxonomic placement was given.
Helminthosporium Link (129)
Suetrong et al. (2009) showed the generic
type and one other species to be anamorphic
Massarinaceae based on phylogenetic analyses
of four genes.
Hendersonia Berk. (130)
Although Hendersonia is regarded as a
synonym of Stagonospora it contains many
species that are still accommodated in the
genus and thus needs a thorough revision.
Morin et al. (2010) described a new teleomorph
genus Austropleospora with a Hendersonia
anamorph which is Camarosporium-like and
clustered in Pleosporales in molecular phylogenetic analysis.
68
Hirsutella Pat. (131)
Phylogenetic analysis confirmed this
genus to be anamorphic Ophiocordyceps
(Johnson et al. 2009). Johnson et al. (2009)
also introduced a new ascomycetous genus
Orbiocrella in Clavicipitaceae with a Hirsutella-like anamorph. The anamorph however
is atypical of Hirsutella and did not cluster
with other Hirsutella species. They concluded
that the genus should have a narrow definition
that is phylogenetically informative. The Hirsutella-like anamorph of Moelleriella was also
confirmed (Tadych et al. 2009). Evans et al.
(2010b) described two new species in this
genus from the red ant and Zou et al. (2010)
describe a single species and all were anamorphic Ophiocordyceps in Ophiocordycipitaceae.
Hormonema Lagerb. & Melin (132)
Talgø et al. (2010) reported that Hormonema dematioides Lagerb. & Melin is anamorphic Sydowia polyspora (Bref. & Tavel) E.
Müll. (Dothioraceae) and is important in
current season needle necrosis of Abies spp.
Hortaea Nishim. & Miyaji (133)
Crous et al. (2009a) confirmed one
species to be anamorphic Teratosphaeriaceae
based on partial LSU gene analysis.
Houjia G.Y. Sun & Crous (134)
Yang et al. (2010) describe two new
species in this new hyphomycetous genus
which they placed in Capnodiales based on
phylogenetic analysis.
Hyalorhinocladiella H.P. Upadhyay & W.B.
Kendr. (135)
Jacobs et al. (2010) confirmed this to be
the anamorph of Ophiostoma.
Hydea K.L. Pang & E.B.G. Jones (136)
This genus was introduced to accommodate the hyphomycetous Cirrenalia pygmea
Kohlm., which has brown helicoid conidia
distinguished by its relatively large apical cell.
The genus did not group with any other
Lulworthiales, Lulworthiaceae forming a
monophyletic clade (Abdel-Wahab et al. 2010).
Mycosphere
Kirramyces J. Walker, B. Sutton & Pascoe
(137)
Crous et al. (2009b) showed by analysis
of LSU sequence data that four species
clustered in Clade 4 of Teratosphaeriaceae
with Teratosphaeria sensu stricto.
Kumanasamuha P.Rag. Rao & D. Rao (138)
A new anamorphic species was described
for this genus by Nagao et al. (2009) and found
to be an anamorph of the type of Chorioactis in
Chorioactidaceae.
Lacazia Taborda, V.A. Taborda & McGinnis
(139)
Vilela et al. (2009) showed that Lacazia
is an anamorphic taxon that is distinct from
Paracoccidioides in the Ajellomycetaceae.
Lasiodiplodia Ellis & Everh. (140)
Begoude (2010) studied Botryosphaeriaceae associated with Terminalia catappa
and phylogenetic analysis confirmed placement
of Lasiodiplodia in this family.
Lecanicillium W. Gams & Zare (141)
Johnson et al. (2009) confirmed by
phylogenetic analysis that this is anamorphic
Cordyceps. Sukarno et al. (2009) described
three new species in the genus from Indonesia.
They adopted this genus, even though it is
paraphyletic, because it is a convenient genus
for characterizing Verticillium-like species
whose teleomorphs have not yet been discovered.
Lecanosticta Syd. (142)
Crous et al. (2009a,d) showed one
species in this genus to be a member of Mycosphaerellaceae in a phylogeny derived from
SSU, 5.8S rDNA and LSU analyses.
Lemonniera De Wild. (143)
Shearer et al. (2009) showed one species
in this genus to be anamorphic Pleosporaceae
based on phylogenetic analyses of SSU and/or
LSU genes.
Lecythophora Nannf. (144)
Damm et al. (2010) described two new
species of the teleomorph genus Coniochaeta
(Coniochaetaceae, Coniochaetales) with Lecythophora anamorphs from Prunus wood.
Lepisticola W. Gams (145)
This genus was introduced as a new
name for the illegitimate name Harziella
Costantin & Matr. 1899 by Gams et al. (2009).
The taxonomic placement is unclear.
Leptodontidium de Hoog (146)
Nekoduka et al. (2010) found that one
species of this genus clustered in Helotiales
following ITS gene sequence analysis along
with the teleomorph Oculimacula.
Leptographium Lagerb. & Melin (147)
Jacobs et al. (2010) confirmed this to be
the anamorph of Ophiostoma. Harrington et al.
(2010) provided a discussion of the merit of
Ophiostoma species with Leptographium anamorphs being recognized as a separate genus,
Grosmannia, concluding that the current phylogenetic analyses are ambiguous.
Leptoxyphium Speg. (148)
Schoch et al. (2009) and Crous et al.
(2009a) showed one species in this genus to be
a member of Capnodiaceae in a phylogeny
derived from combined five gene or LSU gene
analysis. Cheewangkoon et al. (2010) introduced a new species that clustered in Capnodiales in LSU gene analysis.
Mackenziella Yanna & K.D. Hyde (149)
This nom. nov. was introduced in Gams
et al. (2009) to accommodate Mackenziea
Yanna & K.D. Hyde 2002, which is a homonym of Mackenziea Nees 1847. The taxonomic
placement is unclear.
Macrophomina Petr. (150)
Suetrong et al. (2009) showed the generic
type to be anamorphic Botryosphaeriaceae
based on phylogenetic analyses of four genes.
Manoharachariella Bagyan., N.K. Rao &
Kunwar (151)
This new genus producing solitary,
doliform, dictyospetate conidia was introduced
from India by Bagyanarayana et al. (2009). The
taxonomic placement is unclear although listed
as Dothideomycetes in Index Fungorum.
69
Mariannaea G. Arnaud ex Samson (152)
Cai et al. (2010) described a new species
of this genus confirming its affinity with
Nectriaceae.
position was determined by means of 18S and
28S rDNA sequence analysis and suggested
that Micronematobotrys is a member of Pyronemataceae.
Matsusporium E.B.G. Jones & K.L. Pang (153)
This genus was introduced to accommodate the hyphomycetous Cirrenalia tropicalis Kohlm., which has brown helicoid conidia. The genus clustered adjacent to Lulworthia
grandispora Meyers in the Lulworthiales,
Lulworthiaceae (Abdel-Wahab et al. 2010).
Microsphaeropsis Höhn. (160)
Aveskamp et al. (2010) showed the
generic type to be anamorphic Didymellaceae
following multigene phylogenetic analysis.
Melanops Nitschke ex Fuckel (154)
Phillips & Alves (2009) epitypified this
teleomorph genus which has Fusicoccum-like
anamorphs and clusters in Botryosphaeriaceae
in phylogenetic analysis.
Merimbla Pitt (155)
Peterson et al. (2010) described several
species in Hamigera with Merimbla anamorphic states.
Metarhizium Sorokīn (156)
A new species of Metacordyceps (teleomorphic genus) with a Metarhizium anamorph
was introduced by Li et al. (2010).
Microcyclospora Jana Frank, Schroers &
Crous (157)
This new genus was introduced with
three new species causing sooty blotch on fruit.
Molecular phylogenetics analysis showed the
genus to cluster in Mycosphaerellaceae (Frank
et al. 2010).
Microcyclosporella Jana Frank, Schroers &
Crous 2010* (158)
Frank et al. (2010) described this new
monotypic Pseudocercosporella-like genus for
species causing sooty blotch on apple which is
accommodated in Mycosphaerellaceae according to molecular phylogenetic analysis.
Micronematobotrys Xiang Sun & L.D. Guo
(159
Sun & Guo (2010) introduced this
hyphomycete genus for an endophyte isolated
from Quercus and Ulmus west of Beijing. This
genus differed from other Botrytis-like genera
in having micronematous conidiophores and
coloured verrucose conidia. The phylogenetic
70
Microxiphium (Harv. ex Berk. & Desm.)
Thüm. (161)
Schoch et al. (2009), Crous et al. (2009a)
and Ruibal et al. (2009) showed species in this
genus to be members of Capnodiaceae in a
phylogeny derived from molecular analysis
indicating that this genus is polyphyletic as it
has already been linked to Dennisiella in
Coccodiniaceae.
Milesia F.B. White (162)
Salazar-Yepes (2009) described a new
species of Phakopsora with a Milesia
anamorph.
Minimedusa Weresub & P.M. LeClair (163)
Cheewangkoon et al. (2009) introduced a
new combination and phylogenetic analysis of
the LSU gene showed it to cluster in Cantharellales.
Miuraea Hara 1948 (164)
Crous et al. (2009a) showed one species
in this genus to be a member of Mycosphaerellaceae in a phylogeny derived from SSU,
5.8S rDNA and LSU analyses.
Moheitospora Abdel-Wahab, Abdel-Aziz &
Nagahama (165)
Abdel-Wahab et al. (2010) introduced
this genus for a marine hyphomycete from
Egypt with helicospores, which differed from
Cirrenalia species in smaller conidial cells.
The taxa clustered close to Juncigena in the
clade named TBM and was included in the
Hypocreales incertae sedis along with this
genus. They also transferred Cirrenalia adarca
Kohlm., Volkm.-Kohlm & O.E. Erikss. to Moheitospora establishing the link to Juncigena
adarca Kohlm., Volkm.-Kohlm & O.E. Erikss.
Moleospora Abdel-Wahab, Abdel-Aziz &
Nagahama (166)
Mycosphere
This genus was introduced for a marine
hyphomycete genus on Phragmites from Egypt
with young brown helicoid conidia which
eventually become a contorted mass of cells.
Phylogenetic analysis showed it to form a wellsupported monophyletic group in the Lulworthiales, Lulworthiaceae (Abdel-Wahab et
al. 2010).
Monotosporella S. Hughes (167)
Zhang et al. (2009a) showed one species
to be anamorphic Melanommataceae based on
phylogenetic analyses of five genes.
Moromyces Abdel-Wahab, K.L. Pang,
Nagahama, Abdel-Aziz & E.B.G. Jones (168)
Abdel-Wahab et al. (2010) introduced
this genus to accommodate two Cumulospora
species based on phylogenetic data. This
helicosporous hyphomyceteus genus clustered
basal to Lulwoana uniseptata and Zalerion
maritima (Linder) Anastasiou in the Lulworthiaceae, Lulworthiales.
Mycelephas R.F. Castañeda (169)
Gams et al. (2009) introduced this
hyphomycete genus as a nom. nov. for Arnoldiella R.F. Castañeda 1984. No teleomorph
relationship is known.
Mycochaetophora Hara & Ogawa (170)
Nekoduka et al. (2010) recollected M.
gentianae Tak. Kobay., Kasuyama & Nasu of
this genus and found it to cluster in Helotiales.
Mycogelidium W.Y. Zhuang (171)
Kirschner & Oberwinkler (2009) showed
this genus, which had been described as the
sole member of a new family Mycogelidiaceae,
Atractiellales in Basidiomycota, to be anamorphic Capnodiales, possibly Capnodiaceae.
Mycosphaerellaceae Lindau (172)
There are many more anamorphic genera
listed in Kirk et al. (2008) than in Crous (2009)
and this group needs revision.
Nawawia Marvanová (173)
Crous et al. (2009f) described a new
species which clustered with Chaetosphaeria
species in molecular phylogenetic analysis.
Shenoy et al. (2010) found this genus to
be related to Diaporthales.
Neofusicoccum Crous, Slippers & A.J.L.
Phillips (175)
Begoude (2010) studied Botryosphaeriaceae associated with Terminalia catappa
and phylogenetic analysis confirmed placement
of Neofusicoccum in this family. They have
Botryosphaeria-like teleomorphs (Taylor et al.
2009).
Neosetophoma Gruyter, Aveskamp & Verkley
(176)
Gruyter et al. (2010) described this new
genus which is close to Phoma section, but
differ in morphology and molecular phylogeny.
This is anamorphic Phaeosphaeriaceae. It differs in having yellowish conidia that are
attenuate at one end and a Stagnosporopsis
synanamorph.
Neottiosporina Subram. (177)
Zhang et al. (2009a) showed one species
to be anamorphic Massarinaceae based on
phylogenetic analyses of five genes.
Nodulisporium Preuss (178)
Platas et al. (2009) confirm that several
species in this genus are anamorphic Hypoxylon.
Nodulisporium-like (179)
Zare et al. (2010) described a new
species of Coniolariella with a Nodulisporiumlike anamorph.
Oidium Link (180)
Takamatsu et al. (2009) conifrmed this as
anamorphic Golovinomyces and Neoerysiphe.
Paecilomyces Bainier (181)
Samson et al. (2009) confirmed the link
between Paecilomyces and Byssochlamys
based on analysis of three genes, Byssochlamys
comprised nine species with five forming the
teleomorph, while four are asexual
Paliphora Sivan. & B. Sutton (182)
Shenoy et al. (2010) found this to be
anamorphic Chaetosphaeriaceae.
Natarajania Pratibha & Bhat (174)
71
Paraconiothyrium Verkley (183)
Schoch et al. (2010) confirmed this as
anamorphic Montagnulaceae following phylogenetic analysis.
Paraisaria Samson & B.L. Brady (184)
Evans et al. (2010b) described a new
species in this genus from the red ant that were
anamorphic Ophiocordyceps in Ophiocordycipitaceae.
Paraphoma Morgan-Jones & J.F. White (185)
Phoma section Paraphoma, Pyrenochaeta and Pleurophoma were studied by
Gruyter et al. (2010) using SSU and LSU
sequence data. Phoma section Paraphoma was
highly polyphyletic and distantly related to
Phoma section Phoma. Phoma radicina (McAlpine) Boerema, the type of Phoma section
Paraphoma is anamorphic Phaeosphaeriaceae.
Parastenella J.C. David (186)
Kirschner & Chen (2010) describe a new
species in this genus which clusters in
Mycosphaerellaceae in phylogenetic analysis.
Passalora Fr. 1849 (187)
Crous et al. (2009a) showed twelve
species to be members of Mycosphaerellaceae
in a phylogeny derived from SSU, 5.8S rDNA
and LSU analyses and one unidentified species
to be anamorphic Teratosphaeriaceae based on
partial LSU gene analysis indicating the polyphyletic nature of this anamorphic form which
is also anamorphic Mycosphaerella (Mycosphaerellaceae).
Penicillium Link (188)
Peterson & Horn (2009) introduced two
new Penicillium species, which cluster
amongst Eupenicillium species, indicating relationships of the anamorphs and teleomorphs of
these genera. Peterson et al. (2010) combined a
Hamigera with a Talaromyces species, listing a
Penicillium anamorph.
Penidiella Crous & U. Braun (189)
Crous et al. (2009a,c,d) confirmed the
generic type and five other species to be
anamorphic Teratosphaeriaceae and one species to be a member of Mycosphaerellaceae
based on partial LSU gene analysis indicating
72
the polyphyletic nature of this anamorphic
form.
Periconiella Sacc. (190)
Crous et al. (2009a) showed two species
in this genus to be a member of Mycosphaerellaceae in a phylogeny derived from
partial LSU gene analysis.
Pesotum J.L. Crane & Schokn. (191)
Grobbelaar et al. (2009) confirmed this to
be the anamorph of Ophiostoma.
Petromyces Malloch & Cain (192)
Horn et al. (2009a) described the sexual
state of Aspergillus parasiticus Speare as
Petromyces parasiticus B.W. Horn, I. Carbone
& Ramirez-Prado, and Horn et al. (2009b)
described the sexual state of A. flavus Link. as
Petromyces flavus B.W. Horn, I. Carbone &
G.G. Moore.
Peyronellaea Gold. ex Togliani (193)
The section Peyronellaea was studied by
Aveskamp et al. (2009) and phylogenetic
results were unable to support the section as a
taxonomic entity. The section is typified by the
production of dictyochlamydospores but this
feature has developed or was lost many times
during the evolution of Phoma. Likely teleomorphs were Didymella (Didymellaceae), Leptosphaeria (Leptosphaeriaceae), while another
group had no associated teleomorphs in Pleosporales. Aveskamp et al. (2010) however,
reinstated the genus when they used a multigene phylogenetic analysis to test the relationship of a large number of Phoma taxa.
Peyronelina P.J. Fisher, J. Webster & D.F.
Kane (194)
A species of Peyronelina grew in
association with a Flagelloscypha-like basidiomycete on decayed wood near a stream (Yamaguchi et al. 2009). Phylogenetic analysis
based on partial LSU sequences showed it to
cluster in the Flagelloscypha clade, nested
within the Nia clade of Hymenomycetes.
Phacellium Bonord. (195)
Crous et al. (2009a) confirmed one
species to be anamorphic Teratosphaeriaceae
based on partial LSU gene analysis indicating
Mycosphere
the polyphyletic nature of this anamorphic
form, which is also thought to be anamorphic
Mycosphaerella (Mycosphaerellaceae).
Phaeobotryon Theiss. & Syd. (196)
This genus was reinstated by Phillips et
al. (2008) and comprises four species. Abdollahzadeh et al. (2009) described a further
species for the genus and showed two species
to be anamorphic Botryosphaeriaceae based on
molecular sequence analysis.
Phaeocandelabrum R.F. Castañeda, Heredia,
Saikawa (197)
This new genus with complex brown
conidia was introduced by Castañeda Ruiz et al.
(2009b) with three species. No teleomorph
relationship is known.
Phaeomoniella Crous & W. Gams (198)
Damm et al. (2010) describe four new
species in this genus. Rossman et al. (2010)
showed the generic type belongs in a novel
lineage of Chaetothyriomycetidae, Eurotiomycetes using combined phylogenetic analysis of
four genes.
Phaeophleospora Rangel (199)
Crous et al. (2009a,d) showed four species in this genus to be members of Mycosphaerellaceae in a phylogeny derived from
partial LSU gene analysis and Crous et al.
(2009b) also showed one Phaeomoniella–like
species to cluster in Clade of Teratosphaeriaceae.
Phaeosclera Sigler, Tsuneda & J.W. Carmich.
(200)
Ruibal et al. (2009) showed the generic
type inhabiting rocks is anamorphic Myriangiales.
Phloeospora Wallr. (201)
Crous et al. (2009a) showed one species
in this genus to be a member of Mycosphaerellaceae in a phylogeny derived from
SSU, 5.8S rDNA and LSU analyses.
Phaeothecoidea Crous (202)
Crous et al. (2009a,d) confirmed the
generic type and two species to be anamorphic
Teratosphaeriaceae based on partial LSU gene
analysis. It is not clear if some species are
anamorphic Mycosphaerella (Mycosphaerellaceae) as previously recorded.
Phaeothecoidiella Batzer & Crous (203)
Yang et al. (2010) described two new
species in this new anamorphic genus which
they placed in Dothideomycetes following
phylogenetic analysis.
Phialocephala W.B. Kendr. (204)
Grünig et al. (2009) analyzed sequence
data from several species in this genus and
found them to be scattered in the Helotiales.
One example was the anamorph of their newly
introduced Phaeomollisia, while others clustered near Loramyces, Mollisia and Vibrissea
species indicating the polyphyletic nature of
this genus. Wang et al. (2009) also described a
new species of this genus in the Helotiales.
Phialophora-like, anamorphic Neobulgaria
Petr. 1921 (205)
Johnston et al. (2010) described a new
species of Neobulgaria which stains vascular
tissues of kiwifruit and has a Phialophora-like
anamorph.
Phialophorophoma Linder 1944 (206)
Aveskamp et al. (2010) showed one
species (P. littoralis) to be related to Cucurbitariaceae following phylogenetic analysis.
Phoma Sacc. (207)
Aveskamp et al. (2010) used a multigene
phylogenetic analysis to test the relationship of
a large number of Phoma taxa. Phoma species
clustered in six distinct clades in Pleosporales
and was highly polyphyletic. Most species
including the generic type clustered in Didymellaceae. Two species of Phoma clustered in
Sporormiaceae (Aveskamp et al. 2010) and
their status as Phoma species should be reconsidered; similarly two species clustered in
Cucurbitariaceae. In the same study, six
species of Phoma were retrieved in Phaeosphaeriaceae, eleven in the Leptosphaeriaceae/Pleosporaceae clade and seven in the
Didymellaceae clade, the latter incorporating
the generic type of Phoma.
73
Phragmoconidium G.F. Sepúlveda, PereiraCarv. & Dianese (208)
This new genus was introduced by
Pereira-Carvalho et al. (2009) to accommodate
a Brazilian hyphomycetous species growing on
trichomes of Emotum. No taxonomic placement was given.
Physopella Arthur (209)
Salazar-Yepes & Carvalho (2009) described a new species of Cerotelium with a
Physopella anamorph.
Pilidiella Petr. & Syd. (210)
Cheewangkoon et al. (2010) confirmed
two species in this genus to be Schizoparmaceae.
Plenodomus Preuss (211)
Boehm et al. (2009b) mentioned the
connection of this genus with the teleomorph
genus Psiloglonium.
Pleurophoma Höhn. (212)
Aveskamp et al. (2010) showed one
species to be related to Cucurbitariaceae
following phylogenetic analysis.
Polyschema H.P. Upadhyay (213)
Shenoy et al. (2010) found the generic
type and three other taxa species to represent a
monophyletic lineage in Pleosporales.
Polythrincium Kunze (214)
Crous et al. (2009a,d) showed one
species in this genus to be a member of
Mycosphaerellaceae in a phylogeny derived
from partial LSU gene analysis, while Simon et
al. (2009) confirmed P. trifollii to be the
anamorph of Cymadothea.
Prosthemium Kunze (215)
The phylogenetic placement of several
species in this genus was investigated and
found to be anamorphic Pleomassaria lying
basal, but within Melanommataceae (Tanaka et
al. 2010).
Pseudocercospora Speg. (216)
Crous et al. (2009a,d) showed 27 taxa in
this genus to be members of Mycosphae-
74
rellaceae in a phylogeny derived from SSU,
5.8S rDNA and LSU analyses.
Pseudocercosporella Deighton (217)
Crous et al. (2009a) showed three taxa in
this genus to be members of Mycosphaerellaceae and one species to be anamorphic
Teratosphaeriaceae in a phylogeny derived
from SSU, 5.8S rDNA and LSU analyses
indicating the possible polyphyletic nature of
this anamorphic form which is also anamorphic
Mycosphaerella (Mycosphaerellaceae). Frank
et al. (2010) introduced an epitype for the
genus.
Pseudoplagiostoma Cheewangkoon, M.J.
Wingf. & Crous (218)
Cheewangkoon et al. (2010) introduced
this teleomorph genus to accommodate the
generic type Cryptosporiopsis eucalypti Sankaran & B. Sutton and two other species, which
was accommodated in the new family Pseudoplagiostomaceae (Diaporthales). The teleomorph was only known in the generic type.
Pseudoramichloridium Cheewangkoon &
Crous (219)
Cheewangkoon et al. (2009) introduced
this new genus with two new species. The
genus resembles Ramichloridium and is placed
in Teratosphaeriaceae following phylogenetic
analysis.
Pseudorobillarda M. Morelet (220)
Suetrong et al. (2009) showed the generic
type and two other species to be anamorphic
Dothideomycetes based on phylogenetic analyses of four genes.
Pseudotaeniolina J.L. Crane & Schokn. (221)
Crous et al. (2009a) confirmed the one
species to be anamorphic Teratosphaeriaceae
in a phylogeny derived from SSU, 5.8S rDNA
and LSU analyses.
Pseudotetraploa Kaz. Tanaka & K. Hirayama
(222)
Tanaka et al. (2009) introduced this new
anamorphic genus in Tetraplosphaeriaceae
based on SSU and LSU combined sequence
analyses with three new species. No teleomorph is known.
Mycosphere
Pyrenochaeta De Not. 1849 (223)
Gruyter et al. (2010) confirmed that
several species in this genus are anamorphic
Cucurbitariaceae. Herpotrichia parasitica,
which is known to have a Pyrenochaeta anamorph clustered basally in Cucurbitariaceae.
Pyrenochaetopsis Gruyter, Aveskamp &
Verkley (224)
Gruyter et al. (2010) described this new
genus to accommodate several species formerly accommodated in Pyrenochaeta and Phoma.
The genus is close to Pyrenochaeta and belongs in Cucurbitariaceae. It is distinguished
by setose pycnidia as in Pyrenochaeta but has
Phoma-like conidiogenous cells.
Quadricrura Kaz. Tanaka, K. Hirayama & Sat.
Hatak. (225)
Tanaka et al. (2009) introduced this new
anamorphic genus in Tetraplosphaeriaceae
based on SSU and LSU combined sequence
analyses with three new species. No teleomorph is known.
Rachicladosporium Crous, U. Braun & C.F.
Hill (226)
Crous et al. (2009a) described a new
species in this genus and showed two species to
be members of Davidiellaceae in a phylogeny
derived from partial LSU genes.
Racodium Pers. (227)
Ruibal et al. (2009) showed the one
species is anamorphic Capnodiales.
Raffaelea Arx & Hennebert (228)
Kolařik & Hulcr (2009) described a new
species in the genus, which lies within the
Ophiostomataceae and is closest to an Ambrosiella sp., while Massoumi Alamouti et al.
(2009) showed this is anamorphic Grosmannia.
Harrington et al. (2010) regarded Raffaelea as
the proper asexual genus for cycloheximidetolerant symbionts of ambrosia beetles with
Ophiostoma teleomorphs.
Ramichloridium Stahel ex de Hoog (229)
Crous et al. (2009a,d) confirmed five
taxa in this genus to be members of Mycosphaerellaceae, four species to be anamorphic
Dissoconiaceae and one species to be anamor-
phic Teratosphaeriaceae based on partial LSU
gene analysis indicating the polyphyletic nature
of this anamorphic form.
Ramichloridium-like (230)
Réblová & Štěpánek (2009) introduced a
new genus Tectonidula (Sordariomycetes,
genera incertae sedis) with Ramichloridiumlike and Sporothrix-like anamorphs.
Ramoacrodictys G.Z. Zhao (231)
This new genus was introduced to
accommodate a species previously described in
Acrodictys (Zhao et al. 2009). No familial
placement was given.
Ramophialophora M. Calduch, Stchigel, Gené
& Guarro (232)
Madrid et al. (2010a) described a new
species in this genus, which occurs in Sordariales and possibly Lasiosphaeriaceae. They
also discussed the placement of Cladorrhinum.
Ramularia Unger (233)
Kirschner (2009) showed species in this
genus to cluster in Mycosphaerellaceae. Crous
et al. (2009a) showed six taxa in this genus to
be members of Mycosphaerellaceae in a
phylogeny derived from partial LSU gene
analysis.
Ramulispora Miura (234)
Crous et al. (2009a,d) showed one
species in this genus to be a member of
Mycosphaerellaceae in a phylogeny derived
from SSU, 5.8S rDNA and LSU analyses.
Rattania Prabhugaonkar & Bhat (235)
This new sporodochial hyphomycete was
described to accommodate an endophytic hyphomycete from Calamus in India (Prabhugaonkar & Bhat 2009). Shenoy et al. (2010)
found this to be anamorphic Chaetosphaeriales.
Readeriella Syd. & P. Syd. (236)
Crous et al. (2009a,d) confirmed 16
species to be anamorphic Teratosphaeriaceae
based on partial LSU gene analysis, while
Crous et al. (2009b) showed three strains of the
generic type and one other species to sit in
Clade 1 of Teratosphaeriaceae.
75
Repetoblastiella R.F. Castañeda, Minter & M.
Stadler (237)
Castañeda Ruiz et al. (2010d) introduced
this new monotypic hyphomycete genus from
Mexico. No teleomorph relationship is known.
Repetophragma Subram. (238)
Suetrong et al. (2009) showed one
species to be anamorphic Pleosporales based
on phylogenetic analyses of four genes.
Rhabdospora-like (239)
Rossman et al. (2010) redescribe a
species in the teleomorph genus Dolabra which
had been given the invalid anamorph name,
Rhabdospora
nepheliae.
Combined
phylogenetic analysis of four genes showed it
to belong in a novel lineage of Eurotiomycetes.
Rhexocercosporidium U. Braun (240)
Nekoduka et al. (2010) found that the
generic type and one species of this genus
clustered in Helotiales following ITS gene
sequence analysis.
Rhodoveronaea Arzanlou, W. Gams & Crous
(241)
Réblová (2009) found the teleomorph of
this genus and phylogenetic analysis showed in
to be in Sordariomycetidae and not Annulatascaceae.
Rhynchosporium Heinsen ex A.B. Frank (242)
Nekoduka et al. (2010) found that two
species of this genus clustered in Helotiales
following ITS gene sequence analysis.
Rhytisma Fr. (243)
Three new species of this teleomorph
genus were described by Suto (2009) and all
species produced minute conidia in conidiomata. These, however, never germinated in
repeated experiments and were regarded as
spermogonia and spermatia.
Rostrohypoxylon J. Fourn. & M. Stadler (244)
The genus of Xylariaceae was described
with Virgariella-like and Sporothrix-like anamorphs (Fournier et al. 2010a).
Ruwenzoria J. Fornier, M. Stadler, Læssøe &
C. Decock (245)
76
The genus of Xylariaceae was described
with Nodulosporium-like and Sporothrix-like
anamorphs (Stadler et al. 2010).
Sarcinomyces Lindner (246)
Ruibal et al. (2009) showed the generic
type inhabiting rocks is anamorphic Myriangiales.
Scedosporium Sacc. ex Castell. & Chalm. (247)
The link with Pseudallescheria was
confirmed by Rainer & Kaltseis (2010) when
describing three new species.
Sclerostagonospora Höhn. (248)
Possibly anamorphic Didymosphaerella
Cooke (Montagnulaceae) but link shown by
occurrence with the teleomorph and not proven
(Chlebicki 2009).
Sclerotium Tode (249)
Xu et al. (2010) assessed the phylogenetic placement of eight Sclerotium taxa. Two
species previously thought to be Basidiomycota
clustered with Ascomycota and had affinities
with the teleomorph genera Sclerotinia and
Stromatinia along with a third Sclerotium
species already known to be Ascomycota. The
other Sclerotium species were placed in Basidiomycota and clustered with Athelia (3 taxa)
or Ceratobasidium (2 species); the latter two
species were transferred to Ceratorhiza which
is the known anamorph of Ceratobasidium
species.
Scytalidium Pesante (250)
Kang et al. (2010) described two species
in this genus as anamorphic Xylogone, which
clustered with the generic type in a distinct
clade in Leotiomycetes.
Setophoma Gruyter, Aveskamp & Verkley
(251)
Gruyter et al. (2010) described this new
genus, which is close to Phoma section Paraphoma, but differs in morphology and molecular phylogeny. The genus lies in Phaeosphaeriaceae.
Septoria Sacc. (252)
A new species of the teleomorphic genus
Mycosphaerella was described by Lima et al.
(2009) with a Septoria anamorph. Crous et al.
Mycosphere
(2009a,d) showed seven taxa in this genus to
be members of Mycosphaerellaceae in a
phylogeny derived from SSU, 5.8S rDNA and
LSU analyses.
Septoidium G. Arnaud (253)
Chaverri & Gazis (2010) described a new
species of Perisporiopsis with Septoidium
macro- and microconidial anamorphs.
Septonema Corda (254)
Boehm et al. (2009a) described a new
teleomorphic genus Oedohysterium with Septonema anamorphs.
Septosporiopsis W.A. Baker & Morgan-Jones
(255)
Gams et al. (2009) introduced this new
hyphomycete genus to accommodate a species
of Acrodictys which differs from other genera
in the Acrodictys complex. No teleomorph
relationship is known.
Simplicillium W. Gams & Zare (256)
Two species from this genus were the
first diverging lineages of Corycipitaceae
(Johnson et al. 2009) but their occurrence as
hyperparasites of Torrubiella and not anamorphs cannot be dismissed.
Sirodesmium De Not. (257)
Ruibal et al. (2009) showed one species
inhabiting rocks is anamorphic Pleosporales.
Sonderhenia H.J. Swart & J. Walker (258)
Crous et al. (2009a,d) showed two taxa in
this genus to be members of Mycosphaerellaceae in a phylogeny derived from partial
LSU gene analysis.
Spadicoides S. Hughes (259)
Shenoy et al. (2010) found the generic
type to be associated with Porosphaerella
species as sister taxa to the Coniochaetales and
not in Helminthosphaeriaceae. Two other
species also clustered in Sordariomycetes
incertae sedis, while a fourth species clustered
in Pleosporales indicating the polyphyletic
nature of the genus.
Sphaerodes Clem. (260)
This teleomorphic genus was reported to
produce ampulliform phialides on irregularly
branched conidiophores directly on ascomata
or on surrounding hyphae (Vujanovic & Goh
2009)
Sporidesmajora Batzer & Crous (261)
Yang et al. (2010) described this monotypic new hyphomycete genus which they
place in Capnodiales based on phylogenetic
analysis.
Sporidesmium Link (262)
Crous et al. (2009a) confirmed one
species to be anamorphic Teratosphaeriaceae
based on partial LSU gene analysis indicating
the polyphyletic nature of this anamorphic
form which is also anamorphic Mycosphaerella
(Mycosphaerellaceae). Boehm et al. (2009a)
mentioned the link with the teleomorph Psiloglonium Höhn. 1918
Sporothrix Hektoen & C.F. Perkins (263)
Grobbelaar et al. (2009) confirmed this to
be the anamorph of Ophiostoma. Zanzot et al.
(2010) and Madrid et al. (2010b) also described
a new species of Ophiostoma with Sporothrix
anamorphs.
Sporothrix-like (264)
Réblová & Štěpánek (2009) introduced a
new genus Tectonidula (Sordariomycetes,
genera incertae sedis) with Ramichloridiumlike and Sporothrix-like anamorphs.
Stagonospora (Sacc.) Sacc. (265)
Zhang et al. (2009a) showed one species
to be anamorphic Lentitheciaceae based on
phylogenetic analyses of five genes, indicating
the polyphyletic nature, while Aveskamp et al.
(2010) showed one species to be anamorphic
Phaeosphaeriaceae.
Stagonosporopsis Died. (266)
Aveskamp et al. (2010) emended this
genus with 19 species and two varieties and
showed it to belong in Didymellaceae following multigene phylogenetic analysis.
Staninwardia B. Sutton (267)
Crous et al. (2009a,d) confirmed one
species to be anamorphic Teratosphaeriaceae
77
based on partial LSU gene analysis indicating
the probable polyphyletic nature of this anamorphic form which is also anamorphic Chaetothyriales.
Stemphylium Wallr. (268)
The teleomorphs of this genus where
known are anamorphic Pleospora and this was
confirmed by Inderbitzin et al. (2009).
Stenella Syd. (269)
Crous et al. (2009a,d) confirmed one
species to be anamorphic Teratosphaeriaceae
in a phylogeny derived from SSU, 5.8S rDNA
and LSU analyses indicating the polyphyletic
nature of this anamorphic form which is also
anamorphic Mycosphaerella (Mycosphaerellaceae).
Stigmina Sacc. (270)
Crous et al. (2009a) showed a Stigmina
synanamoprh to be a member of Mycosphaerellaceae in a phylogeny derived from SSU,
5.8S rDNA and LSU analyses.
Strelitziana Arzanlou & Crous (271)
Zhang et al. (2009c) introduced a new
species in this genus as anamorphic Chaetothyriales with no teleomorph connection.
Symphyosira Preuss (272)
See entry under Symphyosirella Seifert.
Symphyosirella Seifert (273)
This genus was proposed by Gams et al.
(2009) to solve the problem of the missing type
of Symphyosira Preuss 1853 and was introduced to accommodate two relatively well
understood Symphyosira species, which are
obligate parasites of seeds and have Symphyosirina teleomorphs, but not the type S. lutea
Preuss that is excluded.
Synnemacrodictys W.A. Baker & MorganJones (274)
Gams et al. (2009) introduced this new
hyphomycete genus to accommodate a species
of Acrodictys which differed from other genera
in the Acrodictys complex in having synnematous conidiomata and lack of percurrent
proliferation of the conidiogenous cells. No
teleomorph relationship is known.
78
Taeniolella S. Hughes (275)
Shearer et al. (2009) showed one species
in this genus to be anamorphic Lindgomycetaceae based on phylogenetic analyses of SSU
and/or LSU genes.
Taifanglania Z.Q. Liang, Y.F. Han, H.L. Chu
& R.T.V. Fox (276)
This genus was introduced by Liang et al.
(2009b) for Paecilomyces–like species in
Chaetomiaceae based in ITS and SSU gene
sequence data analysis.
Teratosphaeria Syd. & P. Syd. (277)
Crous et al. (2009b) introduced several
species in this teleomorphic genus where the
sexual characters are unknown and the species
is only known by its anamorphic characters but
were linked to Teratosphaeria by molecular
data. They synonymised taxa previously accommodated in the anamorphic genera Colletogloeopsis, Coniothyrium, Kirramyces and
Readeriella.
Tetraploa Berk. & Broome (278)
Tanaka et al. (2009) introduced the new
teleomorph genus Tetraplosphaeria (Tetraplosphaeriaceae) to accommodate this genus
which has four setose appendages, including
the generic type, which was supported by SSU
and LSU combined sequence analyses. A
similar anamorph with only three setose appendages was found in Triplosphaeria, while in
Polyplosphaeria the Tetraploa-like conidia had
three to eight setose appendages.
Thedgonia B. Sutton (279)
Crous et al. (2009e) and Nekoduka et al.
(2010) found that the generic type of this genus
clustered in Helotiales following LSU and ITS
gene sequence analysis.
Thedgonia-like (280)
Crous et al. (2009a) showed this taxon to
be a member of Mycosphaerellaceae in a
phylogeny derived from SSU, 5.8S rDNA and
LSU analyses.
Torula Pers. (281)
Zhang et al. (2009a) showed one species
to be anamorphic Massariaceae based on
phylogenetic analyses of five genes.
Mycosphere
Toxicocladosporium Crous & U. Braun (282)
Crous et al. (2009a,c) showed the generic
type plus five species in this genus to be
members of Davidiellaceae in a phylogeny
derived from partial LSU genes.
Tricladium Ingold 1942 (283)
Campbell et al. (2009) studied 16 species
of this genus and molecular phylogenetic
analysis showed it to be polyphyletic with
species clustering in distinct five clades with
the generic type, Tricladium splendens Ingold,
clustering in clade 1 of the Helotiales. Other
species clustered in clades 2-4 of Helotiales
while T. caudatum Kuzuha clustered in Rhytismatales.
Trichothecium-like (284)
Doveri et al. (2010) described the teleomorph Rodentomyces with a Trichotheciumlike anamorph.
Trichomatoclava G.F. Sepúlveda, Pereira-Carv.
& Dianese (285)
This new genus was introduced by
Pereira-Carvalho et al. (2009) to accommodate
a Brazilian hyphomycetous species growing on
trichomes of Parinari. No taxonomic placement was given.
The genus is reinstated with two species
and found to be occur in a basal clade of
Dothideomycetes (Tsui & Berbee 2010).
Tulipispora Révay & Gönczöl (290)
Révay et al. (2009) described this genus
from submerged, decaying wood. Conidia
superficially resemble those of Triscelophorus
and Triramulispora species, but the taxon
differs somewhat in the conidial morphology
and especially in the ontogeny.
Ulocladium Preuss 1851 (291)
Runa et al. (2009) found that ten Ulocladium species clustered in a core Ulocladium
clade with all taxa possessing the key diagnostic feature of obovoid conidia. A. cheiranthi
(Lib.) P.C. Bolle and E. indefessa E.G.
Simmons, which did not have this feature, also
clustered in this group. The status of the genus
as an independent lineage basal to the core
Alternaria clade was strongly supported. Two
Ulocladium species clustered in a sister clade
to the core Ulocladium indicating some members of the genus are polyphyletic. No teleomorph genera were used in their analysis but
the taxa can be placed in the Pleosporaceae
with Alternaria.
Trichomatosphaera Pereira-Carv., G.F. Sepúlveda & Dianese (286)
This new genus was introduced by
Pereira-Carvalho et al. (2009) to accommodate
a Brazilian hyphomycetous species. No taxonomic placement was given.
Varicosporina Meyers & Kohlm. (292)
A new teleomorph species Corollospora
anglusa Abdel-Wahab & Nagahama was
described by Abdel-Wahab et al. (2009) and
was linked to the anamorph state Varicosporina which was also described as a new species
V. anglusa Abdel-Wahab & Nagahama.
Tripospermum Speg. (287)
Crous et al. (2009a) confirmed one
species to be anamorphic Teratosphaeriaceae
based on partial LSU gene analysis indicating
the probable polyphyletic nature of this anamorphic form which is also thought to be
anamorphic Capnodiaceae.
Varicosporium W. Kegel (293)
Phylogenetic analysis showed this genus
was polyphyletic and clustered in three distinct
clades in Helotiales (Campbell et al. 2009).
The generic type, V. elodeae W. Kegel,
clustered in Varicosporium clade 1 which
included one Hymenoscyphus species.
Trochophora R.T. Moore (288)
Crous et al. (2009e) positioned this genus
in Mycosphaerellaceae following analysis of
LSU gene sequences.
Variocladium Descals & Marvanová (294)
Phylogenetic analysis showed this genus
clustered in Helotiales (Campbell et al. 2009).
Troposporella P. Karst. (289)
Veramycella G. Delgado (295)
This new genus was introduced for a
taxon from dead palm leaves possessing a
79
unique combination of features (Delgado 2009).
The conidia are distinctive in having 3distoseptate, subhyaline to pale olivaceous,
polymorphic conidia formed by polyblastic,
sympodial conidiogenesis. The placement of
the genus is unknown.
Verrucisporota D.E. Shaw & Alcorn (296)
Crous et al. (2009a,d) showed a possible
strain of the generic type and one other taxon
in this genus to be members of Mycosphaerellaceae in a phylogeny derived from partial
LSU gene analysis.
Virgariella-like (297)
Fournier et al. (2010b) described a new
Hypoxylon species from Northern Ireland with
a Virgariella-like anamorph.
Vermiculariopsiella Bender (298)
Dhargalkar & Bhat (2009) described a
new species of the teleomorphic genus Echinosphaeria with a Vermiculariopsiella anamorph.
Verrucocladosporium K. Schub., Aptroot &
Crous (299)
Crous et al. (2009a) showed one species
in this genus to be a member of Davidiellaceae
in a phylogeny derived from partial LSU genes.
Verrucostoma Hirooka, Tak. Kobay. &
Chaverri (300)
This new teleomorphic genus was introduced by Hirooka et al. (2010) and had an
Acremonium-like anamorph. They also discussed several anamorphic genera in the Bionectriaceae and Nectriceae.
Vesiculohyphomyces Armando, Pereira-Carv.
& Dianese (301)
This new genus was introduced by
Pereira-Carvalho et al. (2009) to accommodate
a Brazilian hyphomycetous species growing on
trichomes of Caryocar. No taxonomic placement was given.
Vonarxia Bat. (302)
Crous et al. (2009e) showed one species
in this genus to be anamorphic Chaetothyriaceae based on LSU gene data analysis.
Xenostigmina Crous (303)
80
Crous et al. (2009e) showed this genus to
belong in Dothideomycetes incertae sedis and
not Mycosphaerellaceae in LSU gene sequence
analysis.
Xylomyces Goos, R.D. Brooks & Lamore (304)
Shearer et al. (2009) conducted molecular sequence-based phylogenetic analyses using
nuclear ribosomal sequences (SSU and/or LSU)
and showed this to be anamorphic Aliquandostipitaceae, which comprise only freshwater
species.
Zalerion R.T. Moore & Meyers (305)
Zalerion maritima (Linder) Anastasiou
1963 clustered in Lulworthiales, Lulworthiaceae with relatively weak support (AbdelWahab et al. 2010).
Zasmidium Fr. (306)
Crous et al. (2009a) showed five taxa in
this genus to be members of Mycosphaerellaceae in a phylogeny derived from partial
LSU gene analysis.
Zygophiala E.W. Mason (307)
Crous et al. (2009a,d) confirmed two
species to be anamorphic Schizothyrium (Schizothyriaceae) based on partial LSU gene
analysis. Ma et al. (2010) also confirmed this
genus to be anamorphic Schizothyrium.
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