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Biodiversity Assessment of Michigan Technical Report Prepared by: John J. Paskus, A. L. Derosier, E. H. Schools, H. D. Enander, B. S. Slaughter, M. A. Kost, and R. L. Rogers Michigan Natural Features Inventory P.O. Box 30444 Lansing, MI 48909-7944 For: Michigan Department of Natural Resources Wildlife Division March 31, 2008 Report Number 2007-11 Suggested citation: Paskus J. J., A. L. Derosier, E. H. Schools, H. D. Enander, B. S. Slaughter, M. A. Kost, and R. L. Rogers. 2007. Biodiversity Assessment of Michigan Technical Report. Report to the Wildlife Division, Michigan Department of Natural Resources. Report number MNFI 2007-11. Michigan Natural Features Inventory, Lansing, MI. Copyright 2007 Michigan State Board of Trustees. Michigan State University Extension programs and materials are open to all without regards to race, color, national origin, gender, religion, age disability, political beliefs, sexual orientation, marital status, or family status. Cover photos: top left, limestone bedrock lakeshore (Patrick J. Comer); top middle, dwarf lake iris (Thomas Arter); top right, northern blue (David Cuthrell); top right, piping plover (MDNR staff); bottom left, American lotus (MNFI staff); bottom middle, wood turtle (Jim H. Harding); bottom middle, pugnose shiner (Konrad Schmidt); bottom right, Tahquamenon Falls State Park aerial (David Kenyon). Acknowledgements We would like to thank the Michigan Department of Natural Resources (MDNR) Wildlife Division for funding this effort to assess the biodiversity of Michigan’s terrestrial and aquatic ecosystems. The Nature Conservancy also provided important funding to upgrade our computer technology for handling the massive amounts of data needed to complete this project. We especially would like to thank Ray Rustem, MDNR Natural Heritage Coordinator, for having the vision to see the importance of this project, and ensuring it was adequately funded over the past few years. More so than any other MNFI project, this effort relied very heavily on data collected by many present and former Michigan Natural Features Inventory (MNFI) field scientists and contributors. This includes all staff that collected data, entered data in the database, provided quality control and assurance of the data, and/or secured funds that supported these activities. In particular, we are very grateful for the work of: Michael Penskar and Ryan O-Connor for meticulously reviewing all existing plant records in the MNFI database, as well as processing the majority of the botany data backlog from outside sources; Dave Cuthrell, Michael Sanders, and Bradford Yocum for processing the biggest backlog of data (animals), compared to all other programs, that primarily came from outside sources such as the U. S. Forest Service, and the MDNR; and Shari Gregory for transcribing the natural community data that was unprocessed from previous project work. We would like to thank the following people for reviewing and providing feedback on earlier versions of the aquatic methodology: Patricia Soranno and Mary Bremigan of Michigan State University, Lizhu Wang of the MDNR Institute of Fisheries Research (IFR), and Michelle DePhillip and Matthew Herbert of The Nature Conservancy (TNC). We would also like to thank Paul Seelbach (IFR), Arthur Cooper (IFR), Christine Geddes (IFR), and Michelle DePhillip (TNC) for help with and discussions about available aquatics GIS data.We would also like to thank Michael Donovan, Paul Seelbach, Peter Badra, Doug Pearsall, Matthew Herbert, and Patrick Doran for reviewing the report and providing valuable insight. Much appreciated administrative support was provided by Sue Ridge, Lyn Scrimger, Nancy Toben, Connie Brinson, and Patrick Brown. Table of Contents Introduction .................................................................................................................................................. 1 Brief Summary of Michigan’s biological diversity ...................................................................................... 4 Approach ...................................................................................................................................................... 8 Terrestrial Biodiversity Assessment Methodology ..................................................................................... 17 Aquatic Biodiversity Assessment Methodology ........................................................................................ 61 Looking for Patterns: Integrating the Data Layers Together .................................................................... 127 Next Steps ................................................................................................................................................. 135 References Cited ....................................................................................................................................... 137 List of Figures Figure 1. Percentage of species tracked by major taxon ............................................................................. 4 Figure 2. Percentage of element occurrences by major taxon ..................................................................... 5 Figure 3. Percentage of natural communities by global rank ........................................................................ 6 Figure 4. Percentage of natural communities by state rank ......................................................................... 6 Figure 5. Regional landscapes of Michigan. ................................................................................................12 Figure 6. Ecological Drainage Units (EDUs) of the Great Lakes. ..............................................................14 Figure 7. IFMAP landcover classification, 2000. .........................................................................................19 Figure 8. Circa 1800 vegetation map. ..........................................................................................................22 Figure 9. All forest patches with boundaries defined by all roads, with a 0, 90, 210, and 300 m buffer ......28 Figure 10. Natural vegetation core areas defined by the no road, major road, and all road data layers ......32 Figure 11. Natural vegetation core areas defined by all roads with a 0, 90, 210, and 300 m buffer . ..........33 Figure 12. Potentially unchanged vegetation core areas ..............................................................................37 Figure 13. Large functional landscape patches defined by all roads with a 0, 90, 210, and 300 m buffer . 40 Figure 14. Large functional landscape patches defined by no roads, major roads, and all roads ................41 Figure 15. Frequency of rare terrestrial species using all last observed dates. ...........................................44 Figure 16. Likelihood of a known rare terrestrial species occurrence still occurring ..................................47 Figure 17. Bio-rarity scores for all element occurrences using all last observed dates - top 10%. .............50 Figure 18: Bio-rarity scores for rare terrestrial species with last observed dates > 1985 - top 10%. .........51 Figure 19. Best two terrestrial element occurrences by sub-subsection or subsection. ..............................53 Figure 20. High quality natural communities with an EO rank of >B/C. .....................................................56 Figure 21. Three best occurrences of each natural community type at the statewide scale. ......................60 Figure 22. Map of size and temperature river classification framework used in analysis. ..........................65 Figure 23. Map of gradient classification framework used in analysis. .......................................................66 Figure 24. Map of connectivity and shoreline complexity for lake classification framework . ....................68 Figure 25. Map of proximate geology lake classification framework used in analysis. ...............................69 Figure 26. Unique river ecosystems in Michigan using the 5% rule. ...........................................................74 Figure 27. Unique river ecosystems in Michigan using the 1% rule. ...........................................................75 Figure 28. Unique river ecosystems in Michigan by EDU using the 5% rule. ............................................76 Figure 29. Unique river ecosystems in Michigan by EDU using the 1% rule. ............................................77 Figure 30. High quality river ecosystems in Michigan by EDU. ..................................................................79 Figure 31. Rivers in Michigan with unimpeded access to the Great Lakes. ................................................81 Figure 32. Intact watersheds of headwater streams in Michigan. ...............................................................83 Figure 33. Percent natural land cover in watersheds of headwater streams in Michigan. ..........................84 Figure 34. Unique lake ecosystems in Michigan using the 5% rule. ............................................................87 Figure 35. Unique lake ecosystems in Michigan using the 1% rule. ............................................................88 Figure 36. Unique lake ecosystems in Michigan by EDU using the 5% rule. .............................................90 Figure 37. Unique lake ecosystems in Michigan by EDU using the 1% rule. .............................................91 Figure 38. High quality lakes by EDU. ........................................................................................................93 Figure 39. Percent natural land cover by sub-watershed. ...........................................................................95 Figure 40. Percent natural land cover in riparian areas by subwatershed. ..................................................96 Figure 41. Land cover analysis by subwatershed. .......................................................................................97 Figure 42. Number of dams per river mile by sub-watershed. .................................................................. 100 Figure 43. Number of road crossings per river mile by sub-watershed. .................................................... 101 Figure 44. Fragmentation analysis by sub-watershed. ............................................................................... 102 Figure 45. Number of DEQ non-point source pollution permits per river mile by sub-watershed. ........... 103 Figure 46. Percent impervious surface by sub-watershed. ....................................................................... 104 Figure 47. Number of active mines per river mile by sub-watershed. ....................................................... 105 Figure 48. Pollution analysis by sub-watershed. ........................................................................................ 106 Figure 49. Sub-watersheds in Michigan scored from least-modified to most-modified. ............................ 108 Figure 50. Frequency counts of aquatic element occurrences by PLSS. .................................................. 112 Figure 51. Frequency counts of aquatic element occurrences without loon EOs by PLSS. ..................... 113 Figure 52. Element occurrence likelihood by PLSS. .................................................................................. 116 Figure 53. Element occurrence biological rarity by PLSS. ........................................................................ 117 Figure 54. Aquatic rare species richness per river mile by sub-watershed. .............................................. 120 Figure 55. Aquatic species of greatest conservation need richness.. ........................................................ 124 Figure 56. Locations of the best occurrences for each element by watershed. ........................................ 125 Figure 57. Prioritized terrestrial biodiversity areas . .................................................................................. 128 Figure 58. Prioritized aquatic biodiversity areas ........................................................................................ 130 Figure 59. High priority great lakes shoreline sites. ................................................................................... 132 List of Tables Table 1. Aquatic and terrestrial species summary ........................................................................................ 4 Table 2. Summary of Natural Communities tracked by MNFI .................................................................... 5 Table 3. IFMAP land cover classification. ...................................................................................................18 Table 4. Summary of Circa 1800 Vegetation Classification. ...................................................................... 21 Table 5. Modified IFMAP land cover classes. ........................................................................................... 24 Table 6. Natural vegetation communities organized by patch type and minimum size. ............................ 26 Table 7. All forest patches with different road and buffer combinations applied. ..................................... 27 Table 8. Summary of natural vegetation core areas in the SLP ecoregional section. ................................. 30 Table 9. Summary of natural vegetation core areas in the NLP ecoregional section. ................................ 30 Table 10. Summary of natural vegetation core areas in the UP ecoregional section. ................................ 31 Table 11. Summary of potentially unchanged vegetation core areas statewide. ........................................ 36 Table 12. Summary of large functional landscape patches statewide. ....................................................... 39 Table 13. Total number of best two terrestrial element occurrences by sub-subsection or subsection. ..... 52 Table 14. Summary of high quality natural communities with an EO rank of > BC. ................................ 55 Table 15. Summary of classification of river valley segments and statewide uniqueness analysis. .......... 71 Table 16. Number of statewide unique VSECs in each EDU using the 5% and 1% rule. ......................... 71 Table 17. Names of rivers within EDUs that have unique VSECs using the 1% rule statewide. ...............72 Table 18. Summary of general river and VSEC statistics within EDUs. ....................................................72 Table 19. Summary of unique river ecosystems by EDUs based on the 5% rule. ......................................72 Table 20. Summary of unique river ecosystems by EDUs based on the 1% rule. ......................................73 Table 21. Names of additional rivers within EDUs that have unique VSECs using the 1% rule . ..............73 Table 22. Landscape variables used to determine quality ............................................................................78 Table 23. Summary of the number of river reaches classified as common ecosystems. ............................78 Table 24. Number of 100% natural headwater watersheds in each EDU. .................................................82 Table 25. Summary of classification of lakes and uniqueness analysis. ......................................................85 Table 26. Number of statewide unique lakes in each EDU using the 5% and 1% rule. .............................85 Table 27. Summary of general lake statistics within EDUs. ........................................................................86 Table 28. Summary of unique lake ecosystems by EDU based on the 5% rule. ........................................86 Table 29. Summary of unique lake ecosystems by EDU based on the 1% rule. ........................................89 Table 30. Summary of the number of high quality lakes by size class in each EDU. ..................................92 Table 31. Percent of sub-watersheds in each EDU in each score category of the functional analysis. . . 109 Table 32. Frequency of element occurrences and number of species occurring in EDUs. ....................... 111 Table 33. Summary statistics on river miles per sub-watershed. ............................................................. 119 Table 34. Summary statistics of 19 sub-watersheds that had <0.1 mi of river. ........................................ 119 Table 35. Species richness per river mile by EDU. .................................................................................. 119 Table 36. Average species of greatest conservation need (SGCN) richness per river mile by EDU. ...... 122 Table 37. Important terrestrial biodiversity area data layers. ................................................................... 126 Table 38. Prioritized terrestrial biodiversity area descriptions. ................................................................ 127 Table 39. Summary of prioritized terrestrial biodiversity area scores. .................................................... 127 List of Appendices Appendix A - Rare terrestrial plant list ................................................................................................... A - 1 Appendix B - Rare terrestrial animal list .............................................................................................. A - 10 Appendix C - Rare aquatic plant list .................................................................................................... A - 14 Appendix D - Rare aquatic animal list ................................................................................................. A - 15 Appendix E - Global and State rank descriptions ................................................................................ A - 17 Appendix F - MNFI Natural Community List ...................................................................................... A - 18 Appendix G - Description of Ecological Drainage Units ..................................................................... A - 20 Appendix H - Natural Vegetation Type datalayers and descriptions .................................................... A - 22 Appendix I - Natural vegetation core area datalayers and descriptions ............................................... A - 54 Appendix J - Potentially unchanged natural vegetation core area datalayers ...................................... A - 72 Appendix K - Matrix vegetation datalayers and descriptions ............................................................... A - 77 Appendix L - EO based datalayers and descriptions ............................................................................ A - 81 Appendix M - Aquatic datalayers and descriptions .............................................................................. A - 83 Introduction Michigan is approximately 37 million acres in size, and contains over 43,000 miles of rivers and streams, nearly 11,000 inland lakes, as well as over 4,500 miles of Great Lakes shoreline. Its diverse glaciated terrain contains a variety of forest, wetland, and grassland communities that provide habitat to over 15,000 native species of insects, 1,815 native species of vascular plants, and 691 native species of animals (Evers 1994). Several of these species, such as Michigan monkey flower and dwarf lake iris are only found in the Great Lakes region. Michigan’s landscape, however, has undergone major changes over the last century and the pace of this change is rapidly increasing. Between 1982 and 1997, acreage of developed land in Michigan grew by over 30 percent. If current trends continue, projections indicate that the built areas of Michigan will increase by 178% between 1980 and 2040 (Public Sector Consultants 2001). In addition to direct habitat destruction, sprawling development patterns are continuing to fragment Michigan’s remaining forests, grasslands, and wetlands, as well as alter hydrologic routing and increase levels of stormwater. As a result of these and other changes to the landscape that have occurred since the early 1800’s, 665 species of the state’s plants, birds, mammals, reptiles, amphibians, fish, insects, and mollusks are listed as threatened, endangered, and special concern. In addition, 46 plants and 10 animals are currently extinct or extirpated in Michigan (Michigan Natural Features Inventory 2006). The major factor contributing to this loss of biological diversity or biodiversity is loss of habitat. Since the mid 1800’s, Michigan has lost over 99 percent of its prairies, oak savannas, and oak and oak-pine barrens. What remains of these communities tend to be small, isolated patches. Michigan has also lost approximately 35 percent of its wetlands through conversion to urban and agricultural land uses, with most of these losses occurring in the southern portion of the Lower Peninsula. In some counties, over 75 percent of the wetlands have been lost. In addition, Michigan has lost approximately 50% of its forest cover, with the majority of that loss occurring in the southern Lower Peninsula. One of the first steps towards conserving Michigan’s natural heritage is knowing what is left on the landscape. With limited resources it is especially important to be able to identify and prioritize the best places to conserve biodiversity. Before too many resources have been allocated, and before too much of our precious natural heritage is lost, a focused effort to assess Michigan’s biodiversity needs to be conducted. This technical report was born out of the MDNR’s Wildlife Action Plan (WAP), which was officially approved by the Natural Resource Council in November, 2005 (Eagle et al. 2005). Of the fourteen threats identified in the WAP, fragmentation was listed as one of the two most important threats to the future of Michigan’s wildlife and landscape features. Conservation needs identified in the WAP to address fragmentation include: 1) identifying large tracts and systems to target for protection, 2) identifying areas of biological significance, 3) identifying lands that serve as important linkages between isolated patches of priority landscape features, and 4) completing an analysis of biodiversity elements to identify areas of high biodiversity regardless of ownership type. In addition, another key issue identified in the plan focuses on ecosystem representation and networks. Conservation needs to address this issue include: 1) establishing a cooperative system that captures the full variety of landscape features and associated wildlife and 2) identifying and protecting additional important lands in representative networks (Eagle et al. 2005). To address the conservation needs stated in the WAP the following key questions need to be answered: 1. How do we go about conducting a biodiversity assessment for the state of Michigan? 1 2. What type of framework should be used to organize the landscape and conduct a statewide analysis? 3. What parts of biodiversity should we focus on for conservation? 4. Where are the best places to conserve these elements of biodiversity? Goals Ultimately, Michigan’s biodiversity needs to be protected by maintaining and restoring all natural community and aquatic ecosystem types, as well as viable populations of all native species in natural patterns of abundance and distribution. The primary goal of this initial effort was to gather, develop, and assess a series of data layers for both terrestrial and aquatic natural features that could be used for future conservation planning efforts at multiple scales. Ultimately, we hope this project provides a foundation for end users to target potentially important terrestrial and aquatic biodiversity areas across the state for biological surveys, and eventually strategic conservation at a variety of scales. This work will inform one of the most important conservation strategies identified in the WAP; the development of a cooperative, voluntary based, statewide, conservation network by providing information and data layers focused on: 1) large, intact natural landscapes, 2) rare species hotspots, 3) representative natural areas and high quality natural communities, 4) functional watersheds, and 5) rare and high quality stream segments and lakes. One of the key decisions made early on in this project was to provide end users with a series of data layers, that can be mixed and matched depending on the end users needs and preferences, to construct a conservation network or help set priorities for inventory. One of the shortcomings with providing a statewide conservation network is that watershed councils, township planning commissions, and park managers all have different conservation values, as well as different needs to help them assess important conservation areas. Providing access to multiple data layers allows the end user to determine their own methods of analysis for identifying important conservation areas for whatever jurisdiction or region that may be of interest. Likewise, it was also decided that at least one possible conservation network alternative would be provided from a scientific point of view. This gives end users the option of utilizing an existing, defensible product, or at the very least an alternative that can be modified to best suit their needs. Major Steps The four major steps of this project were to: 1) review other state biodiversity projects; 2) enhance the natural heritage database; 3) develop an approach and methodology for a GIS biodiversity assessment; and 4) conduct the GIS analysis and develop a technical product. Review Other State Biodiversity Projects Before initiating a biodiversity project in Michigan, we explored and summarized other state level biodiversity conservation efforts from around the country. We expected that only a few such projects existed. In fact, we found that 24 states were involved in some sort of statewide terrestrial biodiversity project since the early 1990’s, however, only three state projects involved aquatic biodiversity (Florida, Massachusetts, and Missouri). Only a few of these projects were completed as of December 2002, most were a work in progress, and some were just getting under way. In total, 35 different projects from 24 states were reviewed. We used these previous efforts to inform the approach we developed to ensure we captured the best components. Enhance the Natural Heritage Database The natural heritage database is a critical component of the biodiversity assessment. In fact it is probably the heart of the assessment. When this project was initiated, the Michigan Natural Features 2 Inventory (MNFI) had a high volume of data backlog to review in each of the field science disciplines, particularly zoology and botany. In addition, many of our existing Element Occurrences’ (EOs) were missing relevant pieces of information such as EO rank, or required a comprehensive review due to revised EO specifications or other issues. During the course of this project, over 2,500 element occurrence records were added to the MNFI database. In addition, all plant, animal, and natural community records were quality checked for element occurrence rank and spatial location accuracy. For the terrestrial natural communities, a new procedure was developed to improve the standardization of natural community ranks, provide consistent identification of natural community observations, and provide consistent specifications for each type of natural community. Develop an Approach and Methodology As stated previously, Michigan’s biodiversity comes from both terrestrial and aquatic ecosystems. These ecosystems and associated species have evolved, function, and are classified very differently. Terrestrial ecosystems are primarily influenced by climate, landforms, soils, and vegetation and are described in terms of biomes, ecoregions, landscapes, and vegetation types. Whereas aquatic ecosystems are also influenced by these variables, they are further defined by how water flows over the landscape and are described in terms of basins, watersheds, and water body types. As a result, species distributions and migrations for most aquatic species tend to be restricted to watersheds. Terrestrial species on the other hand tend to be much less restricted, and many terrestrial animal species use both terrestrial and aquatic habitats to complete their life cycle or to exploit resources. Another complication is the discrepancy in available classification frameworks. A solid framework and classification system had already existed and has been tracked in the MNFI database for terrestrial systems in Michigan for several decades. On the other hand, frameworks and classifications for aquatic systems have only recently been described, are still under development and have not been tracked by MNFI. In Michigan, it is critical that both terrestrial and aquatic elements are taken into account, in order to sufficiently address biodiversity. As a result of the functional and practical differences, the terrestrial and aquatic analyses were generally conducted separately. By having both, it also makes it possible to combine data layers and identify places that are important for both terrestrial and aquatic biodiversity. We outlined a methodology and conducted preliminary analyses for conducting a statewide assessment for terrestrial and aquatic biodiversity. We used a fine and coarse filter approach with prioritization to represent biodiversity at a variety of levels; this approach has been frequently used and advocated for (Angermeir and Schlosser 1995, Grossman et al. 1998, Abel et al. 2000, Noss 2004). More information will be given on the fine and coarse filter approach in the approach section of this document. This methodology brings together existing and newly created data to begin assessing Michigan’s biodiversity statewide and regionally as well as identifying weaknesses or information gaps needed to create a more robust assessment. Develop Technical Product The primary purpose of this effort was to produce a technical report detailing a methodology for conducting a statewide assessment for biodiversity in Michigan. This report brings together existing GIS data layers and produces new data layers with associated metadata relevant to the assessment. This information can be used to begin answering key conservation questions, and to address some of the most important conservation needs outlined in Michigan’s WAP. As a follow up to this project, it is our intention to utilize the information resulting from this project along with additional input, information, and analyses to create a user-friendly publication, such as Massachusetts’ BioMap and Living Waters or Oregon’s Living Landscape, for the state of Michigan. 3 Brief Summary of Michigan’s Biological Diversity What exactly are we trying to conserve? Most conservation references today focus on the conservation of an area’s biological diversity or biodiversity. Biodiversity is most simply defined as the variety of life on earth and its processes. More specifically, it is the variety of living organisms, the genetic differences among them, the communities and ecosystems in which they occur, and the ecological and evolutionary processes that keep them functioning, yet ever changing and adapting (Noss and Cooperrider 1994). It is typically measured at several levels of organization: genes, species, natural communities, and landscape ecosystems. The principles of biological protection and restoration are based on several assumptions: 1) biodiversity depends on functioning ecosystems, 2) biodiversity, at all levels, is integral to ecosystem function, 3) priority should be given to keystone species, 4) ecological redundancy is important to the long-term persistence of ecosystems, and 5) natural processes and disturbances are critical to the health and evolutionary pathways of native ecosystems and their associated biota (Armstrong, 1993). In addition, it is important to realize that native ecosystems are complex systems that we still do not fully understand. The MNFI database tracks a total of 665 different plant and animal species (Table 1, Appendix A, B, C, and D). The majority, 417 (62%) are plants, and the next largest category is insects with 94 (14%) (figure 1). The five species that have gone extinct include: one bird (passenger pigeon), three fish (deepwater cisco, blackfin cisco, and bluepike), and one snail (acorn ramshorn). Of the 665 species tracked, 94 or 14% have a global rank of G1-G3 as assigned by NatureServe (Figure 2). G1 refers to species that are considered critically imperiled on a global scale; G2 means that a species is considered globally imperiled, and G3 means that a species is either very rare throughout its range or found locally in a restricted range. Although plants have the highest number of G1-G3 species, 58% of the mussels tracked by MNFI have a global rank of G1- G3. This represents approximately 20% of all native mussels found in Michigan. In addition, 40% of the reptiles and 32% of the insects tracked in the MNFI database have a global rank of G1 – G3. For more information about global and state ranks, please refer to Appendix E. Table 1. Aquatic and terrestrial species summary (MNFI 2006) Major Taxon plants mammals breeding birds reptiles amphibians fish insects snails mussels Totals Total Total Native Extinct State X State E State T State SC Tracked # of EOs *G1-G3 % G1-G3 1,815 46 52 210 109 417 5,923 32 8% 68 4 2 4 10 79 1 10% 238 1 1 8 13 21 43 3,056 3 7% 28 2 2 6 10 1,211 4 40% 23 1 1 2 4 149 0 0% 136 3 6 8 7 11 35 761 7 20% 15-20,000 8 11 75 94 1,061 30 32% 180 1 2 2 29 33 207 7 21% 46 8 2 8 19 700 11 58% 5 53 94 250 265 665 13,147 95 14% 4 Figure 1. Percent of species tracked by major taxon insects 14% snails mussels 5% 3% fish amphibians 5% 1% reptiles 2% Plants 62% breeding birds 6% mammals 2% Figure 2. Percent of element occurrences by major taxon fish 6% insects 8% snails 2% mussels 5% amphibians 1% plants 45% reptiles 9% breeding birds 23% mammals 1% There were a total of 1,371 natural community element occurrences in the MNFI database with the most recent last observed date of September 28, 2006 (Table 2). This represented about 9 % of the total MNFI database (plants, animals, and natural communities). Of the 1,371 natural community element occurrences in the MNFI database, 68 % (932) of these occurrences had an element occurrence rank of BC or higher (A, AB, B, BC). These ranks were interpreted to mean that these occurrences are high quality and viable over a long period of time. The spatial extent of natural communities with a BC rank or higher totaled 390,919 acres; approximately 1 % of the landscape. Of the 74 different types of natural communities tracked by MNFI, 56 % are considered to be critically imperiled or imperiled in Michigan (SX - S2) (Figure 3). Incredibly, 90 % of Michigan’s natural communities are considered to be at least rare or uncommon in Michigan (SX - S3), and 64 % are considered to be at least very rare or local throughout their range (G1-G3) (Figure 4). All prairies and savannahs (grassland dominated systems) in Michigan have a state rank of SX – S3 and a global rank of G1 - G3. Bur oak plains, a type of savannah historically found in the interlobate region of the 5 Table 2. Summary of Natural Communities tracked by MNFI Major Natural Community Groupings Upland Forest Lowland Forest Non-forested wetlands Prairie Savanna/barrens Other (mostly Great Lakes shoreline) # of # of MNFI Acres SX S1 S2 S3 S4 S5 EOs types 7 255 105,277 7 7 184 42,890 6 1 24 598 135,643 5 5 8 4 2 5 47 884 3 2 8 75 13,209 1 4 3 23 74 1,371 431,964 Totals 4 15 212 134,061 1 16 25 25 Figure 3. Percentage of natural communities by state rank S5 3% SX 1% S4 7% S1 22% S3 34% S2 33% Figure 4. Percentage of natural communities by global rank G5 0% GNR 3% GU 11% G1 8% 4 G2 14% G4 22% G3 42% 6 5 2 G1 G2 G3 G4 G5 GNR GU 3 1 2 2 3 6 3 3 4 4 3 1 15 5 6 1 2 1 1 5 1 6 10 32 16 0 1 1 2 8 southern Lower Peninsula, is the only natural community considered extirpated from Michigan. In terms of the 6 major categories of natural communities identified in table 2, the non-forested wetlands category contains the most natural community types tracked by MNFI with 24 (32 %). The non-forested wetlands category also has the highest number of element occurrences at 598, which represents approximately 44 % of all natural community EO’s in Michigan (For a list natural communities tracked by MNFI, please see Appendix F). 7 Approach Different Types of Approaches To help the inform assessment of Michigan’s biological diversity, we reviewed and summarized other state level biodiversity conservation efforts from around the country. We expected only a few such projects existed. In fact, we found that 24 states were involved in some sort of statewide terrestrial biodiversity project since the early 1990’s, and only three states had conducted aquatic biodiversity projects. Few of these projects were completed (as of 2002), the majority were still a work in progress, and some were just getting under way. In total, there were 35 different projects to review (several states had multiple projects), and only 16 projects developed a repeatable methodology. The approaches taken by these 16 projects were categorized into four different types: status, fine filter, coarse filter, and prioritization. Status refers to the current status and trends of biodiversity in the state without identifying conservation priorities or specific sites on the landscape. Fine filter focuses on species that slip through the cracks such as rare, focal, or restricted species. Coarse filter focuses on natural communities, ecological hubs, core areas, connecting corridors, enduring features (e.g., land type associations), and large blocks of undeveloped land. The main idea behind the coarse filter is that these larger features, such as natural communities, capture the majority of common species associated with that feature. Prioritization involves ranking the final set of sites based on some sort of value system. For a species or natural community, this could be based on its global or state rarity rank, and/or element occurrence rank, i.e. its viability. Of the 16 projects that developed assessment methodology, states either conducted: 1) a status assessment, or employed: 2) a coarse filter approach, 3) a combination of a fine and coarse filter approach, 4) a combination of a coarse filter and prioritization approach, or 5) a combination of a fine filter, coarse filter, and prioritization approach. Maine was the only state to conduct strictly a status assessment. Five projects (Florida: ecological network project, Illinois, Indiana, Missouri: aquatic integrity areas, and Wisconsin) employed the coarse filter approach, while six projects (Delaware, Florida: closing the gaps, Massachusetts: biomap and living waters, Oregon, and Vermont) used a combination of fine filter and coarse filter. Maryland was the only state to use a coarse filter-prioritization approach, while New Jersey, Florida (Florida Forever conservation needs assessment), and Missouri (GAP) were the only projects to use a combination of fine filter, coarse filter, and prioritization. In our opinion, the best assessment methodologies were developed by the states of Florida, Missouri, and New Jersey. These states used a fine and coarse filter approach with prioritization. It should be noted that Florida was motivated by legislation to acquire land based on a scientific approach, and Missouri has been working on their assessment since 1997. Other commendable assessments were developed by Massachusetts (and used by Delaware), Oregon, Maryland, and Vermont. All four of these states used both a fine and coarse filter approach but decided not to prioritize the final selection of sites based on ecological significance. Below is a brief summary of the Florida, New Jersey, and Missouri assessments. Florida Forever Conservation Needs Assessment The Florida Forever Conservation Needs Assessment was prepared by the Florida Natural Areas Inventory in 2000. It was funded by the Florida Department of Environmental Protection, Division of State Lands and was initiated by the Florida Forever Act, a 10 year, $3-billion land and water conservation program. The act specifically states that acquisition should be based on a 8 comprehensive assessment of Florida’s natural resources and planned so as to protect the integrity of ecosystems. The goal of the project was to develop and compile statewide resource data to evaluate the protection status of these resources and guide decisions about future conservation efforts. Three overlay models were developed for the report: 1) a biodiversity model, 2) a water resources model, and 3) an integrated conservation priorities model. The biodiversity model overlayed the Strategic Habitat Conservation Areas (SHCA), Florida Natural Areas Inventory (FNAI) element occurrence records, Habitat Conservation Priorities (HCP), ecological greenways and underrepresented natural community data layers. Overlap was addressed by halving the weighting factor for individual species habitats in the FNAI data layer that were common to both FNAI and SHCA. Areas of the natural community data layer that overlapped with SHCAs were removed from the natural community data layer. The water resources model combined the floodplain, surface water, wetlands, and aquifer recharge data layers. The floodplain data layer was scored significantly less to reduce double counting. The integrated model combined the biodiversity model, water resources model, and two additional layers – coastal resources and recreation. Scores for each model were lumped into five priority classes. A GAP Analysis For Riverine Ecosystems Of Missouri The GAP Analysis for Riverine Ecosystems of Missouri (Sowa et al. 2005, 2007), prepared by the Missouri Resource Assessment Partnership (MoRAP), was started in 1997 and completed in 2005. It was funded by the U.S. Geological Survey’s National Water Quality Assessment Program, the U.S. Department of Defense-Legacy Program, and the Missouri Department of Conservation. This project is a bit different from the other state efforts in that it is a GAP analysis. The GAP project set out to identify riverine ecosystems, habitats, and species that are not adequately represented within existing conservation lands. To accomplish this they created a hierarchical riverine ecosystem classification using GIS. This classification scheme incorporated and nested ecological drainage units, aquatic ecological system types, and valley segment types. They also predicted species distributions based on available data and the create classification. By using this data in conjunction with public ownership and stewardship lands, and a human-threat index, a conservation plan for Missouri was developed. New Jersey Landscape Project The New Jersey Landscape Project (2001) was prepared by the Endangered and Nongame Species Program, New Jersey Division of Fish, and Wildlife and Rutgers University. The goal of the project was to protect New Jersey’s biological diversity by maintaining and enhancing rare wildlife populations within healthy, functioning ecosystems. To achieve this goal, the project set out to identify and map areas of critical habitat for rare species within each of the five major landscape regions. Continuous patches for each habitat type are delineated and then intersected with endangered and threatened species location data. Patches were classified based on conservation status of species present (i.e., patches with federally listed species were given a higher ranking than patches with state listed species). Only seconds precision records with a last observation date of 1970 or greater were used. The project also identified critical area maps for species dependent on forests, forested wetlands, emergent wetlands, grasslands, and dunes. Highest rank was assigned to patches with federally listed species, followed by state endangered, state threatened, non-listed state priority species, and finally patches that met the minimum size requirement (different for each habitat type). In addition, each patch was coded with the number of listed species present as well as the total number of species records within the patch. 9 Our Approach There are essentially five basic concepts that form the foundation of the Michigan statewide biodiversity assessment: 1) representation, 2) regionalization, 3) quality (viability), 4) core ecological areas, and 5) supporting natural landscapes. Each of these concepts can be applied to both the terrestrial and aquatic analysis. Representation To truly conserve biodiversity, The Nature Conservancy (TNC) recommends that there be a sufficient number, distribution, and quality of each native species and ecosystem to ensure their long term persistence within an ecoregion (1996). Capturing multiple examples is necessary to capture variability and to ensure persistence in the face of natural and human disturbances. However, it is an impossible task to track all native species of biota. The native biota of an area includes innumerable species unknown or at best poorly known to science, embedded in numerous ecosystems whose webs of biotic and abiotic interactions are only poorly understood (Parrish et al. 2003). Ideally, conservation decisions would be based on definitive knowledge of the distribution and viability of native species within an ecosystem. However, it is impossible to track all native species and their biotic and abiotic interactions. Coarse Filter - Fine Filter Approach: One solution to this problem is to identify conservation targets. TNC defines conservation targets as a limited number of species, natural communities or ecosystems chosen to represent the biodiversity of a given area. Due to the limitations of using individual species as filters for other species, it is recommended to initially select ecological communities or ecosystems as coarse filter targets (Noss et al. 1994). Ecological communities or ecosystems are often defined as the sum of the assemblages of populations of plants, animal, bacteria, and fungi and their environment (Groves 2003). If ecological communities are to work as coarse filters for all associated plants and animals they must (Anderson et al. 1999): 1) be conserved as often as possible at a size and scale that they naturally occurred prior to major human impacts; 2) be conserved as part of dynamic, intact, landscape mosaics; 3) maintain some level of connectivity between communities; and 4) contain a full complement of their associated flora and fauna in so far as it is known. In addition, TNC also recommends that smaller and rarer natural community types (lakeplain prairie, prairie fen, coastal plain marsh, bog) should be represented at a higher number in the landscape than larger and more common community types such as mesic southern forest. The coarse filter approach should then be followed by the selection of species with unique ecological requirements that cannot be met through the conservation of natural communities or ecosystems. Wide ranging, rare, extremely localized or keystone species are all likely to need fine filter strategies (Abell et al. 2002). Furthermore, the spatial scale at which organisms use the environment differs tremendously among species and depends on body size, food habits, mobility, and other factors. Hence, no coarse filter will be a complete assessment of biodiversity protection status and needs. However, species that are not addressed using the coarse filter, such as narrow endemics and wideranging mammals or fish, can be captured by the safety net of the fine filter. Community-level (coarse-filter) protection is a complement to, not a substitute for, protection of individual rare species (Donovan et al. 2004). One approach is to identify a set of species typical of or restricted to a particular community in the ecoregion and then use available information on their space, resource, and breeding habitat needs to 10 determine minimum area requirements for the community type (Anderson et. al. 1999). Building on this concept, Lambeck (1997) recommends the use of a suite of focal species to define different spatial and compositional attributes that must be present in a landscape and their appropriate management regimes. All species considered at risk are grouped according to the processes that threaten their persistence. Within each group, the species most sensitive to the threat is used to define the minimum acceptable level at which that threat can occur. Species are categorized as either arealimited, resource-limited, dispersal-limited, and/or process-limited (Lambeck 1997). Combined, this has commonly been referred to as the coarse filter-fine filter approach to biological conservation. Representative Outliers: High quality and/or rare occurrences of species may not be located in high biodiversity value areas or large functional sites. As mentioned earlier, to truly conserve biodiversity, there needs to be a sufficient number, distribution, and quality of each native species to ensure their long term persistence within an ecoregion. Since it is impossible to track all species and their occurrences in Michigan, only species tracked by the MNFI database were considered (endangered, threatened, and special concern). It is important to ensure that a sufficient number of occurrences for each rare and declining species in Michigan are identified for protection regardless of landscape context and integrity and possibly even viability. These outlier occurrences may actually be more important than populations that are located in more contiguous settings because they may contain unique genomes. Regionalization To adequately ensure representation, species and ecosystems need to be distributed across their range. A critical step to ensuring representation is determining a regionalization framework. For the terrestrial analysis, we used Albert’s (1995) regional landscape ecosystems of Michigan. Albert’s approach to classifying landscapes in the upper Midwest can be characterized as multifactor and multilevel in orientation. Landscape units are delineated based on multiple abiotic factors (bedrock geology, glacial landforms, soils, hydrology, and climate). This approach provides a basis for understanding patterns of species distribution, natural disturbance regimes, and natural processes. The classification is also hierarchical; the landscape is viewed as a series of various sized ecosystems nested within one another. The three hierarchical levels are section, subsection, and sub-subsection. There are 4 sections, 22 subsections, and 38 sub-subsections in Michigan (Albert 1995). Section boundaries were used for the coarse scale terrestrial analysis. Due to the relative intactness of the vegetation in the Upper Peninsula, the western and eastern Upper Peninsula sections were combined for this study. Related to this, the boundary between the northern and southern Lower Peninsula was modified slightly in order to minimize false fragmentation of vegetation patches that fell along the section boundary. Subsection and sub-subsection boundaries were used as a surrogate to capture potential genetic diversity for the species representation analysis (Figure 5). All levels were used to identify high quality natural communities. For the aquatic assessment, we used Ecological Drainage Units (EDU’s) of the Great Lakes as the regionalization framework (TNC 2001, Higgins et al. 2005). EDU’s are aggregates of watersheds based on hydrologic units that share similar ecological characteristics such as climate, hydrologic regime, physiography, and zoogeographic history. EDU’s and ecoregions do share similar characteristics but EDU’s are based on watersheds which provide a more effective framework for aquatic ecosystems and species distributions. EDU’s have been shown to be effective in landscapebased classification efforts for both riverine and lake ecosystems (Higgins et al. 2005, Cheruveilil in prep) and have been used in other biodiversity planning efforts (Sowa et al. 2005, 2007). This regionalization will allow us to break the state up into meaningful units to ensure representation of aquatic ecosystems and populations. There are a total of nine Ecological Drainage Units in Michigan 11 Figure 5. Regional landscapes of Michigan. 12 (Figure 6). We combined the Bayfield Peninsula and Uplands (12) EDU with the Western Upper Pennisula and Keweenaw Penninsula (6) EDU and the Western Lake Erie (2) EDU with the Southeast Michigan Interlobate and Lake Plain (16) EDU together for a total of 7 EDUs for this analysis. For a detailed description of each EDU see Appendix G. Quality (viability) TNC defines ecological integrity as the ability of an ecological system to support and maintain a community of organisms that has species composition, diversity, and functional organization comparable to those of natural habitats within a region (reference sites). An ecosystem or species has integrity or is viable when its dominant ecological characteristics - composition, structure, function, and processes - occur within their natural ranges of variation and can withstand and recover from most disturbances. In other words, ecosystems and populations of plants and animals should be self-sustaining. Integrity expresses itself in the characteristics of resistance and resilience. TNC recommends using three criteria to assess integrity: 1) size, 2) current condition, and 3) landscape context. Size: Stability and resilience of a terrestrial natural community tend to increase with the size of the patch. For a natural community occurrence to persist over long time frames, it much be large enough to sustain, absorb, and buffer disturbances. For rivers, size is thought of more as longitudinal intactness, although there is little research to suggest optimal stream lengths for the preservation of natural processes. However, research is working towards identifying minimal units for river conservation (Fausch et al. 2002, Allen 2004). On the other hand, the persistence of small patch natural communities however, such as depressional wetlands or lakes, is largely dependent on the surrounding landscape context rather than size. Evidence also suggests that species loss is strongly correlated with the size and landscape context of the area (Newmark 1987). For species, size is a quantitative measure of the area and/or abundance of an occurrence. Components of this factor are: a) area of occupancy; b) population abundance; c) population density; and d) population fluctuation (NatureServe 2003) Current Condition: Current condition refers to the viability of the occurrence. For a natural community or ecosystem, condition refers to native species diversity, threats, presence of exotic species, and is affected by: 1) anthropogenic impacts (exp. fragmentation, pollution) and 2) biological legacies. TNC defines biological legacies as critical features that take hundreds to thousands of years to develop. In forests these might include: presence of fallen logs and rotting wood, a well developed moss and herbaceous understory, structural complexity in the canopy and understory layers, a reservoir of soil organic matter for nutrient storage, seed banks, and evidence of intact nutrient cycles. For rivers these might include: channel sinuosity, riffle – pool – run composition, and available substrates. For species, condition refers to demographics, reproductive success, degree of threats, and extent and quality of critical habitat. For many animals, condition is very difficult to determine due to the intensity and duration of sampling required to get scientifically defensible data. As a result, the majority of animal occurrences in the MNFI database (64%) are given an element occurrence rank of E for 13 Figure 6. Ecological Drainage Units (EDUs) of the Great Lakes. 14 extant. In other words, if an individual or several individuals are found within a given area, their presence alone does not allow scientists to comment on the long-term viability of the population. Landscape Context: Landscape context for terrestrial ecosystems refers to the size of the surrounding natural vegetation patch or block, proximity and extent of incompatible land uses, and the potential for ecological processes to occur at natural rates and scales. Surrounding landscape functionality (context) is an issue for all communities, but particularly for patch types that depend on easily disrupted processes occurring at scales larger than those of the individual community. Examples of key threats to consider in the surrounding landscape include: fire suppression, diversion of groundwater, coastal revetments, impervious surface, and agricultural runoff. For aquatic ecosystems, landscape context must be viewed at different spatial scales. Ecological processes (i.e. hydrologic, geomorphic) function at the catchment (or watershed) scale for each reach of a stream or the catchment of a lake. Additionally, adjacent and upstream riparian areas can have a strong influence on the functionality of a stream reach and the availability of habitat. Landscape context in aquatic ecosystems refers to the proximity and extent of incompatible land uses, the potential for ecological processes to occur at natural rates and scales, and the amount of natural land cover within the catchment and the riparian area. Examples of key threats to consider in the surrounding landscape include: dams, impervious surface, erosion, diversion of groundwater, agricultural runoff, and road crossings. Core Ecological Areas Large Functional Landscapes: These are the best areas to conserve terrestrial biodiversity over the long term, maintain essential ecological processes and services and provide habitat for common species. These areas also provide the best opportunity for supporting viable populations of rare species and high quality natural communities. Landscape integrity is critical to maintaining the long-term viability of species and natural communities. Landscape integrity addresses the health of the larger ecosystem, as well as large scale stresses impacting individual components across the landscape. Without landscape integrity, maintaining fragmented patches of habitat and isolated populations of flora and fauna becomes akin to keeping a patient alive on a respirator in the hopes that a cure will be discovered in the future. Fragmentation is one of the greatest threats to biodiversity. Large functional landscapes provide the best chances for mitigating the effects of roads, invasive species, pollution, development, and other threats to biodiversity, and allow natural processes to occur at more natural rates and scales. Natural disturbances such as flooding, wildfires, tornadoes, ice storms, insect outbreaks, and disease alter the landscape and ultimately help create the variety of ecosystems needed to provide habitat for Michigan’s native species. Functional Watersheds: Aquatic species conservation is a very difficult task given the interconnected nature of rivers and the high vulnerability of both lake and river ecosystems to human disturbance. Parallel to the large functional terrestrial landscapes, functional watersheds provide the best opportunity to conserve biodiversity over the long-term, maintain essential ecological processes and services and provide habitat for common species. By identifying watersheds that have a relatively high degree of integrity, we can focus conservation efforts on those watersheds that can have the greatest long-term impact on aquatic conservation, including rare species. Functional watersheds are areas that can be characterized as having: 1) high percentage of natural land cover, 2) low imperviousness, 3) intact riparian buffers, and 4) minimal road/stream crossings, dams, point source pollution sites, and nearby mining operations. 15 Biological Rarity Hotspots (biological rarity score): This concept has often been referred to as biological hotspots. The general idea is to prioritize spatially defined areas on the landscape that contain a large number of rare or declining species and natural communities. These areas may not coincide with the conditions of the other concepts (representation, quality, functional watersheds, and large functional landscapes), but in general areas that contain concentrations of globally imperiled species and/or occurrences of rare species or natural communities with high viability should receive higher priority over areas with concentrations of element occurrences with low viability and/or more secure species. Supporting Natural Landscape This concept was borrowed from the Massachusetts BioMap project (2000), however our interpretation of the supporting natural landscape is much narrower. We define the supporting natural landscape as natural lands not included as part of the large functional landscapes described in the previous section. Although these lands do not contain known occurrences of rare species or natural communities, or are not part of a large, high quality, undeveloped roadless area, these lands provide potential ecological services or functions. They provide the potential for connectivity between important wildlife habitat areas, buffering large intact patches from incompatible land uses, and allowing natural processes such as flooding to occur at more natural rates and scales. These lands may be smaller fragments, degraded due to human activity, or intensively managed for natural resources such as timber, game species, or other recreational pursuits. The important point here is that these areas have potential natural resource value that should be evaluated by the local or regional community or land manager. Primary evaluation should be based on the ecological value these lands could provide to nearby or adjacent terrestrial and aquatic core ecological areas and representative plant, animal, and natural community occurrences. Products One of the things we noticed from the other state biodiversity projects was the tendency to develop only one solution. However, we realized up front that different end users have different needs and values. The very concept of conservation is inherently based on values. A group or an individual conserves things in the natural environment based on what they think is important. Potential criteria for conservation may include: function, aesthetics, services, goods, recreation, jobs, and/or the needs of future generations. Another very important point to consider is that there will always be uncertainty in the data used in the analysis, and new information will have the tendency to change outcomes, sometimes significantly. Some of the data sets available for this statewide analysis are outdated, incomplete, and/ or have a level of accuracy that may be appropriate at the statewide or regional scale but may not be appropriate to use at a smaller scale. A local unit of government for example, may have a more recent and/or more accurate land cover data set than the statewide IFMAP land cover data set used in this analysis. By providing only one composite product, we eliminate the opportunity for end users to incorporate better data sets. To address these challenges, we decided to focus on flexibility instead of the creation of one solution that somehow fits every end user’s needs. The primary goal of this initial effort was to gather, develop, and assess a series of data layers for both terrestrial and aquatic natural features that could be used for future conservation planning efforts. We addressed this by creating a wide-ranging series of data layers with associated documentation, as well as creating several composite maps to show end users different ways the various data layers can be integrated to develop various conservation network designs at multiple scales. 16 Terrestrial Biodiversity Assessment Methodology Introduction The analysis used in the assessment of Michigan’s terrestrial biodiversity was based on two major categories of data: land cover and element occurrences of natural features. The two land cover datasets used were developed from two different projects: the Michigan GAP Analysis project developed by the Michigan Department of Natural Resources (MDNR), and the circa 1800 vegetation of Michigan project developed by the Michigan Natural Features Inventory (MNFI). The element occurrence dataset is a continuously updated database developed and maintained by the Michigan Natural Features Inventory (MNFI). From the land cover datasets we developed a large number of new data layers that can be used to identify and prioritize core vegetation areas, potentially unchanged vegetation, large functional landscapes, important patches of different vegetation cover types, and large supporting landscapes. The MNFI element occurrence database identifies places on the land that contain unique elements of biodiversity – rare species and high quality natural communities, which MNFI refers to as element occurrences. The database, which is updated periodically throughout the year, contains a wealth of detailed information that was used to identify and prioritize areas based on frequency, likelihood of persistence, viability, and/or rarity of EOs. Both land cover and EOs of natural features are discussed in more detail below. Categories of land cover based datasets developed by this project: 1. Natural vegetation core areas - by ecoregional section 2. Potentially unchanged natural vegetation core areas - by ecoregional section 3. Natural vegetation types - statewide 4. Large functional landscapes – statewide Categories of MNFI EO based datasets developed by this project: 1. EO frequency count 2. EO likelihood 3. Bio-rarity score 4. Best two occurrences of each terrestrial species by sub-subsection 5. High quality natural communities (> B/C rank) 6. Best three occurrences of each natural community (statewide, section, subsection, and subsubsection) A table summarizing the EO based datasets can be found in Appendix L. Coarse Filter: Land Cover Data The following paragraph was primarily borrowed from the Michigan GAP Analysis Project Final Report (Donovan et al. 2004).Vegetation patterns are an integrated reflection of the physical and chemical factors that shape the environment of a given land area (Whittaker 1965). They also are determinants for overall biodiversity patterns (Franklin 1993, Levin 1981, Noss 1990), and they can be used as a currency for habitat types in conservation evaluations (Specht 1975, Austin 1991). The central concept is that the physiognomic and floristic characteristics of vegetation (and, in the absence of vegetation, other physical structures) across the land surface can be used to define biologically meaningful biogeographic patterns. 17 IFMAP land cover Description: A major component of the Michigan GAP Analysis Project was the development of a statewide digital land coverage called the Integrated Forest Monitoring Assessment and Prescription (IFMAP) land cover (Figure 7). IFMAP was developed to assess the distribution and protection status of terrestrial vertebrate species in Michigan, and to assist with forest inventory on state lands (Donovan et al. 2004). We decided to use the IFMAP land coverage for this assessment because it is currently the most up-to-date, statewide, digital land coverage for Michigan. The IFMAP land coverage was derived from classification of Landsat Thematic Mapper (TM) imagery taken between 1999 and 2001. The data is stored in a raster format with a cell resolution of 30 meters. Both supervised and unsupervised classication techniques were used in conjunction with multiple ancillary data sources to produce 32 categories of land cover ranging from high density residential to lowland deciduous forest (Table 3). Table 3. IFMAP land cover classification. Class Name Low Intensity Urban High Intensity Urban Airports Roads / Paved Non-vegetated Farmland Row Crops Forage Crops / Non-tilled herbaceous Orchards / Vineyards / Nursery Herbaceous Openland Upland Shrub / Low-density trees Parks / Golf Courses Northern Hardwood Association Oak Association Aspen Association Other Upland Deciduous Mixed Upland Deciduous Pines Other Upland Conifers Mixed Upland Conifers Upland Mixed Forest Water Lowland Deciduous Forest Lowland Coniferous Forest Lowland Mixed Forest Floating Aquatic Lowland Shrub Emergent Wetland Mixed Non-Forest Wetland Sand / Soil Exposed Rock Mud Flats Other Bare / Sparsely Vegetated Value 1 2 3 4 5 6 7 9 10 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 35 18 Figure 7. IFMAP landcover classification, 2000. 19 Limitations: IFMAP data products and assessments represent a snapshot in time generally representing the date of the satellite imagery (1999-2001). Users of the data must be aware of the static nature of the products. IFMAP data are derived from remote sensing and modeling. Any decisions based on the data must be supported by ground-verification and more detailed analyses. An accuracy assessment of the final land cover layer determined it to be 87 percent accurate at level 2 in the hierarchical classication scheme. At the next level of classification detail (level 3), class accuracies range from 36 to 87 percent. Overall accuracy was 80.7 percent for the non-forested types and 67.9 percent for forested types (Donovan et al. 2004). Please see the Space Imaging Report “Review of Remote Sensing Technologies used in the IFMAP Project” (Space Imaging 2004) for a complete discussion of the accuracy assessment and associated tables. MNFI Circa 1800 Vegetation of Michigan Description: Between 1816 and 1855 Government Land Office Surveyors mapped a one-mile grid across the entire surface of Michigan, starting in the southeast near Lake Erie and finishing along the Wisconsin border along Lake Superior. The Land Office Surveyors were not only creating a grid for land sales, they were also recording information about the land and its vegetation, describing the fertility of the soil, mapping bedrock exposures, and recording the size and species of the trees. As they measured out the boundaries of townships and sections, surveyors made notes on the topography, soils, and vegetation they encountered along each one mile section line. Surveyors were instructed to note the exact location of wetlands, lakes and streams, comment on the agricultural potential of soils, and note the quantity and quality of timber resources as they were encountered along each section line (White 1984, Caldwell 1990). With this information plotted over topography maps, ecologists interpreted cover type boundaries primarily using the locations of dominant tree species and associated landforms. Wetland boundaries were interpolated between section lines by using associated elevation lines as they were depicted on the topographic maps. Ecologists consulted surface geology maps, soils maps, and earlier vegetation maps throughout the process of interpretation. Once cover type boundaries were interpreted and assigned codes, the maps were proofed and then digitized (Figure 8, Table 4) (Comer et al. 1995). Limitations: Given that these surveys were not undertaken as a scientific sample of vegetation, they should not be considered as such. It is important to place the circa 1800 vegetation map within the context of the times when the surveys were conducted. Aspects of long-term climatic cycles, Native American activities, and the European fur trade, all had the potential to influence natural patterns on the landscape traversed by surveyors in the nineteenth century. The interpolated boundary line between each section line should be considered an approximation that could differ on the ground depending on local variation not apparent on topographic maps. Upland and wetland boundaries in interior sections should be most accurate where topography is abrupt. Given the scale of survey data, much of the small-scale variation one normally encounters in natural environments was not well represented. One should assume that wetlands which naturally occur as relatively small, complex shapes, totaling less than 50 acres in area, are highly under-represented in this data layer (Comer et al. 1995). 20 Table 4. Summary of Circa 1800 Vegetation Classification. Cover Type Aspen-birch forest Beech-sugar maple forest Beech-sugar maple-hemlock forest black ash swamp Black oak barren Cedar swamp Exposed bedrock Grassland Hemlock-white pine forest Hemlock-yellow birch forest Jack pine-red pine forest Lake/river Mixed conifer swamp Mixed hardwood swamp Mixed oak forest Mixed oak savannah Mixed pine-oak forest Muskeg/bog Oak-hickory forest Oak/pine barrens Pine barrens Sand dune Shrub swamp/emergent marsh Spruce-fir-cedar forest Sugar maple-basswood forest Sugar maple-hemlock forest Sugar maple-yellow birch forest Wet prairie White pine-mixed hardwood forest White pine-red pine forest White pine-white oak forest Total Acres 292,266 5,845,677 6,346,662 280,705 719,043 1,254,093 9,209 73,088 1,962,192 295,314 1,112,655 799,203 4,290,553 1,421,462 418,363 1,061,564 106,331 287,610 1,888,010 112,051 270,330 18,365 608,044 954,169 213,036 2,321,507 948,608 382,029 1,185,681 1,272,127 437,231 37,187,178 Coarse Filter: Land Cover Analysis Introduction The primary purpose of the land cover analysis was to identify the most important natural vegetation areas in the state. Ideally, condition would be one of the primary variables to prioritize or rank one patch over another. Other relevant variables include size, core area, shape, proximity, connectivity, and landscape context. Due to the large area of analysis, high degree of variation from one part of the state to another, and the high number of pixels that needed to be processed, we decided to minimize the number of variables, and focus primarily on: 1) total size, 2) core area, and 3) condition. Using these three variables, five different types of land cover analyses were conducted for the whole state. 21 Figure 8. Circa 1800 vegetation map. 22 1. 2. 3. 4. Land cover analyses Natural vegetation core areas - by ecoregional section Potentially unchanged natural vegetation core areas - by ecoregional section Natural vegetation types - statewide Large functional landscapes – statewide All four analyses are based on the 2001 IFMAP land coverage, and one includes the MNFI circa 1800 vegetation data layer. Two analyses provide information on an ecoregional section basis, and the other two analyze natural vegetation patches from a statewide perspective. As stated earlier, the boundaries of Albert’s (1995) four ecoregional sections were modified to minimize problems associated with artificially fragmenting natural vegetation patches that fell along the section boundaries. The western and eastern Upper Peninsula were combined, and the boundary between the Northern Lower Peninsula and Southern Lower Peninsula were slightly modified to follow existing breaks in the vegetation. A brief discussion of how these data layers can be used in combination with other data layers is provided at the end of this chapter, as well as the chapter entitled: Looking for Patterns: Bringing the Data Layers Together. IFMAP Reclassification A modified version of the IFMAP land cover classes was created to help minimize inaccuracies and to simplify a land cover analysis of the whole state. For example the aspen, oak, and maple layers were combined together to form an upland deciduous forest type layer, rather than treating each forest type individually. In total, eight different natural land cover types were identified for this project: 1) upland deciduous forest, 2) upland mixed forest, 3) upland conifer forest, 4) lowland deciduous forest, 5) lowland mixed forest, 6) lowland conifer forest, 7) grassland, and 8) non-forested wetlands (table 5). Roads Three different road data layers were used in the analysis to distinguish between patches of vegetation. The first data layer did not include any roads, the second data layer used only major roads to differentiate between patches, while the third data layer used all roads to identify vegetation patches. The road data layer used in the analysis is the Michigan Geographic Framework Statewide All Roads Layer Version 5a. All road arcs identified in the Framework were converted to a 30 meter raster dataset. Roads were used to differentiate and define vegetation patches due to their widespread yet uneven distribution across the landscape, combined with their potential impact on wildlife and ecological processes. According to Diamondback (1990), the construction and maintenance of roads is among the most widespread form of modification in the United States during the past century. Road construction kills sessile and slow moving organisms in the path of or areas influenced by the road. Existing roads: 1) cause mortality of both vertebrates and invertebrates from collision with vehicles, 2) modify animal behavior (such as altered home range, altered movements, altered reproductive success and altered escape patterns), and 3) increase the spread of exotic species (Trombulak and Frissell 2000). Species prone to road kill include moose, white-tailed deer, raccoon, opossum, wolf, barn owl, eastern screech owl, American kestrel, frogs, turtles, amphibians, and flying invertebrates such as butterflies. Research has shown that many different types of animal species are impacted by roads. Black bear in North Carolina shift their home range away from areas with high road densities (Brody and Pelton 1989), and several species of rodents, such as white footed mice and prairie voles, will not cross 23 Table 5. Modified IFMAP land cover classes. IFMAP Class Name Low Intensity Urban High Intensity Urban Airports Roads / Paved Non-vegetated Farmland Row Crops Forage Crops / Non-tilled herbaceous Orchards / Vineyards / Nursery Herbaceous Openland Upland Shrub / Low-density trees Parks / Golf Courses Northern Hardwood Association Oak Association Aspen Association Other Upland Deciduous Mixed Upland Deciduous Pines Other Upland Conifers Mixed Upland Conifers Upland Mixed Forest Water Lowland Deciduous Forest Lowland Coniferous Forest Lowland Mixed Forest Floating Aquatic Lowland Shrub Emergent Wetland Mixed Non-Forest Wetland Sand / Soil Exposed Rock Mud Flats Other Bare / Sparsely Vegetated Value 1 2 3 4 5 6 7 9 10 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 35 IFMAP_Code 110 123 121 122 2111 2112 2113 222 310 320 350 411 412 413 414 419 421 423 429 431 500 611 612 613 621 622 623 629 710 720 730 790 Natural Vegetation New Class Name X X Filtered grassland Filtered grassland X X X X X X X X X X X X X X X X X Upland decidious forest Upland decidious forest Upland decidious forest Upland decidious forest Upland mixed forest Upland conifer forest Upland conifer forest Upland conifer forest Upland mixed forest Water Lowland decidious forest Lowland conifer forest Lowland mixed forest Non-forested wetland Non-forested wetland Non-forested wetland Non-forested wetland roads as narrow as 3 meters (Swihart and Slade, 1984). Productivity of bald eagles in Oregon and Illinois declined with proximity to roads (Anthony and Issacs 1989, Paruk 1987), and they preferentially nested away from roads. Sandhill cranes also avoid nesting near paved and gravel roads (Norling et al. 1992). In Ontario, it was discovered that the local abundance of toads and frogs was inversely related to traffic density on adjacent roads. Despite the lower populations adjacent to highly trafficked roads, roadkill relative to abundance was higher on highly traveled roads (Fahrig et al. 1995). More recently, a study conducted in upstate New York found that turtle populations in high road density areas had a much higher proportion of males than populations found in low road density areas. The study suggests that more female turtles are killed on roads presumably during nesting migration (Gibbs and Steen 2005). Grassland and forest interior birds also appear to be affected by roads. The population density of the most sensitive forest interior species (cuckoo) in a recent study was significantly reduced within a distance of 650 meters from the road (Forman and Deblinger 1999). Similarly, in a Netherlands study, 24 the most sensitive grassland species (black tailed godwit) was significantly reduced in density within a distance of 930 meters from the road (Reijen et al. 1996). Buffers In addition to roads, three different buffers were applied to roads and non-natural landcover classes to represent the potential impact of incompatible edges on wildlife and natural processes: 90 meters, 210 meters, and 300 meters. Initially we intended to use 100 meter increments; however, the IFMAP raster land cover data layer consists of 30 meter pixels. As a result we chose to substitute 90 meters for 100 meters, and 210 meters for 200 meters. These distances were chosen based on a literature review of buffers. Rodgers et al. (1997) found that flushing distances of waterbirds extended to 100 meters, and a 100 meter buffer around forests was found to be sufficient for a relatively sensitive guild of bird species (Sandilands and Hounsell 1994). Bolger et al. (1997) found that the abundance of interior habitat bird species was reduced within 200 meters of an edge, and Sandilands and Hounsell (1994) found that a 200 meter buffer around a forest was sufficient for a second more sensitive guild of bird species. Lastly, Brittingham and Temple (1983) found that nest parasitism by brown headed cowbirds decreased with distance away from forest edge, but extended greater than 300 meters into the forest, and Environment Canada (2004) recommended that natural lands should be buffered up to 300 meters to avoid the negative effects of edges on wildlife. Natural vegetation types - statewide Description: This analysis focused on the different natural vegetation communities found in Michigan. Each patch of natural vegetation was buffered from roads and non-natural land cover using several different buffer widths, and then either selected or removed based on the large patch, small patch, matrix size criteria developed by The Nature Conservancy (Anderson et al. 1999). A matrix community is defined as a large, regional sized cover type that ranges in size from 2,000 to 100,000 hectares. They typically encompass a variety of large and small patch communities. Examples of matrix communities include: northern hardwood forests, deserts, mangrove swamps, tallgrass prairies, and tundra. For Michigan, only upland deciduous forests were categorized as a matrix community type. Large patch communities are communities that are relatively easy to define spatially, and range in size from 20 to 2,000 hectares. In Michigan, large patch communities include: forested wetlands, coastal wetlands, barrens and savannas, and upland conifer forests. Small patch communities were defined as communities with a very limited, highly defined spatial extent that are typically embedded with larger community types. Sizes typically range from .1 hectares to 20 hectares. Examples of small patch communities in Michigan include: fen, coastal plain marsh, emergent marsh, dry sand prairie, and bog. The 11 different vegetation categories used in the analysis were: 1) forest, 2) upland forest, 3) upland deciduous forest, 4) upland mixed forest, 5) upland coniferous forest, 6) lowland forest, 7) lowland deciduous forest, 8) lowland mixed forest, 9) lowland coniferous forest, 10) filtered grassland, and 11) non-forested wetland (Table 6). Due to the large amount of anthropogenic grasslands in Michigan, a process was used to identify existing grasslands that were also historically grasslands; these patches are referred to as filtered grassland. Twelve different data layers were developed for each of the 11 vegetation type categories mentioned above, for a total of 132 data layers. The 12 data layers are: 1) no roads – no buffer, 2) no roads - 90 m buffer, 3) no roads – 210 m buffer, 4) no roads – 300 m buffer, 5) major roads – no buffer, 6) major roads – 90 m buffer, 7) major roads – 210 m buffer, 8) major roads – 300 m buffer, 9) all roads – no buffer, 10) all roads – 90 m buffer, 11) all roads – 210 m buffer, and 12) all roads – 300 m buffer. 25 Table 6. Natural vegetation communities organized by patch type and minimum size. Natural Vegetation Types forest upland forest upland decidous forest upland mixed forest upland coniferous forest lowland forest lowland deciduous forest lowland mixed forest lowland coniferous forest filtered grassland non-forested wetland Patch Type matrix matrix matrix matrix large large large large large large small Minimum size (hectares) 2,000 2,000 2,000 2,000 20 20 20 20 20 20 0.1 Please refer to appendix H for metadata. Use: The analysis can be used to identify the largest most intact patches for each of the 11 vegetation type categories. These data layers can also be used to analyze patch statistics for each of the 11 types such as mean and median patch size, range, total acreage, etc. Limitations: As mentioned earlier, IFMAP land coverage is limited in accuracy. In addition, the IFMAP land cover was documented from satellite imagery taken between 1999 and 2001. Some areas of land have been altered since that time period rendering the land cover outdated for those areas. File names: nva2 (grid) nva2_buffered (grid) Data source: Michigan Geographic Framework statewide all roads layer version 5a Lu2001v2_g – IFMAP circa 2000 land use data for entire state of Michigan in grid format. Results: Due to the large number of data layers associated with this analysis, the results section only focused on the all forest category. There were a total of 24,617 patches of forestland in the state totaling 17,860,005 acres (Table 7). When a minimum patch size of 5,000 acres was applied, total acres of all forest dropped to 15,024,720 acres (only a 16% decrease). However, when all roads are used to define patch boundaries, and a 300 meter buffer is applied to each road and non-natural landcover, forest area dropped to 628,640 acres (a 96.5 % decrease). This demonstrates that although forest (both upland and lowland combined) is the dominant land cover in the state, roads have a tremendous impact on Michigan’s forest ecosystems (Figure 9). 26 Table 7. All forest patches with different road and buffer combinations applied. Vegetation Type All Forest All Forest All Forest All Forest All Forest All Forest All Forest All Forest All Forest All Forest All Forest All Forest road layer none none none none major major major major all all all all Total acres # of minimum acres with % acres buffer of natural patches size road layer that with road size in vegetation patches meet layer that meters type (acres) minimum size meet minimum size 17,860,005 17,860,005 17,860,005 17,860,005 17,818,578 17,818,578 17,818,578 17,818,578 16,834,320 16,834,320 16,834,320 16,834,320 24,617 24,617 24,617 24,617 25,496 25,496 25,496 25,496 48,097 48,097 48,097 48,097 5,000 5,000 5,000 5,000 5,000 5,000 5,000 5,000 5,000 5,000 5,000 5,000 15,024,720 15,024,720 15,024,720 15,024,720 14,674,106 14,674,106 14,674,106 14,674,106 6,820,601 6,820,601 6,820,601 6,820,601 84% 84% 84% 84% 82% 82% 82% 82% 41% 41% 41% 41% acres with road layer and buffer 0 15,024,720 90 5,889,270 210 2,185,384 300 1,147,199 0 14,674,106 90 5,779,619 210 2,177,455 300 1,145,941 0 6,820,601 90 2,900,842 210 1,206,356 300 628,640 % acres with road layer and buffer that meet minimum size 100% 39% 15% 8% 100% 39% 15% 8% 100% 43% 18% 9% Natural vegetation core areas - by ecoregional section Description: All natural vegetation types identified by the IFMAP land coverage were combined together to form a new natural vegetation core area data layer. All natural vegetation patches greater than a threshold size, with the threshold dependent on the modified ecoregional section were selected. The Upper Peninsula (UP) threshold was set at 5000 acres, the Northern Lower Peninsula (NLP) threshold was set at 2,500 acres, and the Southern Lower Peninsula (SLP) was set at 500 acres. These select patches were then buffered inward with a series of three buffer sizes (90 meter, 210 meter, and 300 meter). For each buffer, all natural vegetation patches greater than a threshold size, with the threshold dependent on the ecoregional section, were extracted. Water, which includes lakes, ponds, and large river segments, was originally included as part of the natural vegetation data layer. Once the buffers were applied, water bodies with a surface area greater than 10 acres were subtracted out of the data layer and the remaining patches were regrouped and extracted based on the ecoregional thresholds mentioned above. Threshold sizes were set for each of the three ecoregions based on the percentage of natural lands remaining, degree of fragmentation, and mean patch size using all roads with no buffer to define patches. The Nature Conservancy suggests using a 5,000 acre minimal size for matrix patches, however, due to the wide variation in patch sizes between the UP and SLP, thresholds had to be customized to each ecoregional section. We decided to keep the 5,000 acre threshold for the UP due to its high percentage of natural lands (86 %) and large mean patch size (1,299 acres). A threshold of 2,500 acres was set for the NLP due to its moderate amount of natural lands (53%) and mean patch size (341 acres). Additionally, 2,500 acres is within the range needed for to support a female black bear and cubs (Roger and Allen 1987) and 75-80 % of all highly sensitive bird species (Herkert et al. 1993). The threshold for the SLP was set at only 500 acres due to its relatively low percentage of natural lands (25%) and small mean patch size (108 acres). However, 500 acres was found to be sufficient for supporting 80 % of all expected bird species (Tate 1998). 27 Figure 9. All forest patches with boundaries defined by all roads, with 0, 90, 210, and 300 meter buffers applied to roads and non-natural vegetative landcover. 28 Twenty-four different data layers were developed for natural vegetation core areas based on roads and buffers. Twelve data layers included water in the analysis, and the remaining 12 did not include water in the analysis. The 12 data layers are: 1) no roads – no buffer, 2) no roads - 90 m buffer, 3) no roads – 210 m buffer, 4) no roads – 300 m buffer, 5) major roads – no buffer, 6) major roads – 90 m buffer, 7) major roads – 210 m buffer, 8) major roads – 300 m buffer, 9) all roads – no buffer, 10) all roads – 90 m buffer, 11) all roads – 210 m buffer, and 12) all roads – 300 m buffer. The remaining 12 data layers are the same except that water was removed from the analysis. Please refer to appendix I for metadata. Use: The natural vegetation core areas can be used to identify the largest patches of natural vegetation within each ecoregion. Limitations: As mentioned earlier, IFMAP land coverage is limited in accuracy. In addition, the IFMAP land cover was documented from satellite imagery taken between 1999 and 2001. Some areas of land have been altered since that time period rendering the land cover outdated for those areas. File names: natveg2 (grid) _natveg2 (grid - water removed) Data source: Lu2001v2_g – IFMAP circa 2000 land use data for entire state of Michigan in grid format. Michigan Geographic Framework statewide all roads layer version 5a Results: Total area of natural vegetation in the SLP equals 4,266,953 acres (20 % of the statewide total) (Table 8). This represents 27 % of the SLP region. Using 500 acres as a minimum patch size, the total area of natural vegetation in the SLP dropped to 3,065,733 acres (a 38 % decrease). Mean patch size was 4,194 acres. When all roads were used to define patch boundaries, and a 300 meter buffer was applied to each road and non-natural landcover, total area of natural vegetation in the SLP decreased to 14,373 acres. This represents only 0.3 % of the original area. These numbers indicate that natural vegetation in the SLP primarily consists of small, isolated, highly fragmented patches that are heavily impacted by roads (Figures 10 and 11). Total area of natural vegetation in the NLP equals 7,325,525 acres (35 % of the statewide total) (Table 9). This represents 67 % of the NLP region. Using 2,500 acres as a minimum patch size, the total area of natural vegetation in the NLP dropped to 6,845,366 acres (a 7 % decrease). Mean patch size was 60,047 acres. When all roads are used to define patch boundaries, and a 300 meter buffer was applied to each road and non-natural landcover, total area of natural vegetation in the NLP decreased to 409,586 acres. This represents only 5.6 % of the original area. These numbers indicate that natural vegetation in the NLP primarily consists of moderately sized, somewhat fragmented patches that are impacted primarily by minor roads (Figures 10 and 11). 29 Table 8. Summary of natural vegetation core areas in the SLP ecoregional section. Ecoregions (modified) water removed SLP SLP SLP SLP SLP SLP SLP SLP SLP SLP SLP SLP road layer none none none none major major major major all all all all Total acres # of min. acres with of natural patches size road layer vegetation (acres) that meet minimum size 4,266,953 4,266,953 4,266,953 4,266,953 4,251,419 4,251,419 4,251,419 4,251,419 3,829,234 3,829,234 3,829,234 3,829,234 19,859 19,859 19,859 19,859 20,414 20,414 20,414 20,414 35,192 35,192 35,192 35,192 500 500 500 500 500 500 500 500 500 500 500 500 3,065,733 3,065,733 3,065,733 3,065,733 3,000,175 3,000,175 3,000,175 3,000,175 1,135,828 1,135,828 1,135,828 1,135,828 % acres buffer acres with % acres with road size in road layer with road layer that meters and buffer layer and meet buffer that minimum meet size minimum size 72% 0 3,065,733 100% 72% 90 328,973 11% 72% 210 126,979 4% 72% 300 51,526 2% 71% 0 3,000,175 100% 71% 90 328,973 11% 71% 210 126,979 4% 71% 300 51,526 2% 30% 0 1,135,828 100% 30% 90 144,204 13% 30% 210 39,883 4% 30% 300 14,373 1% Mean patch size 4,194 1,574 1,549 1,145 3,375 1,574 1,549 1,145 917 936 928 898 Table 9. Summary of natural vegetation core areas in the NLP ecoregional section. Ecoregions road Total acres # of min. (modified) - layer of natural patches size water vegetation (acres) removed acres with road layer that meet minimum size NLP NLP NLP NLP NLP NLP NLP NLP NLP NLP NLP NLP 6,845,366 6,845,366 6,845,366 6,845,366 6,845,366 6,845,366 6,845,366 6,845,366 2,730,501 2,730,501 2,730,501 2,730,501 none none none none major major major major all all all all 7,325,535 7,325,535 7,325,535 7,325,535 7,305,038 7,305,038 7,305,038 7,305,038 6,859,681 6,859,681 6,859,681 6,859,681 4,712 4,712 4,712 4,712 5,229 5,229 5,229 5,229 20,080 20,080 20,080 20,080 2,500 2,500 2,500 2,500 2,500 2,500 2,500 2,500 2,500 2,500 2,500 2,500 30 % acres buffer with road size in layer that meters meet minimum size 93% 93% 93% 93% 94% 94% 94% 94% 40% 40% 40% 40% 0 90 210 300 0 90 210 300 0 90 210 300 acres with road layer and buffer 6,845,366 2,872,189 1,638,581 835,520 6,845,366 2,872,189 1,638,581 835,520 2,730,501 1,971,399 905,964 409,586 % acres Mean with road patch size layer and buffer that meet minimum size 100% 60,047 42% 16,699 24% 13,655 12% 9,495 100% 60,047 42% 16,699 24% 13,655 12% 9,495 100% 5,450 72% 5,357 33% 5,148 15% 5,319 Total area of natural vegetation in the UP was 9,502,487 acres (45 % of the statewide total) (table 10). This represented 88 % of the UP region. Using 5,000 acres as a minimum patch size, the total area of natural vegetation in the UP dropped to 9,354,185 acres (only a 2 % decrease). The mean patch size was 1,169,273 acres. When all roads were used to define patch boundaries, and a 300 meter buffer was applied to each road and non-natural landcover, total area of natural vegetation in the UP decreased to 2,659,822 acres. This represents 28 % of the original area. These numbers indicate that natural vegetation in the UP primarily consists of large, highly connected patches that are somewhat impacted by minor roads (Figures 10 and 11). Table 10. Summary of natural vegetation core areas in the UP ecoregional section. Ecoregion( modified) UP UP UP UP UP UP UP UP UP UP UP UP road Total acres # of min. layer of natural patches size vegetation (acres) none none none none major major major major all all all all 9,502,487 9,502,487 9,502,487 9,502,487 9,484,693 9,484,693 9,484,693 9,484,693 9,247,664 9,247,664 9,247,664 9,247,664 1,944 1,944 1,944 1,944 2,296 2,296 2,296 2,296 7,117 7,117 7,117 7,117 5,000 5,000 5,000 5,000 5,000 5,000 5,000 5,000 5,000 5,000 5,000 5,000 acres with % acres buffer road layer with road size in that meet layer that meters minimum meet size minimum size acres with road layer and buffer 9,354,185 9,354,185 9,354,185 9,354,185 9,291,547 9,291,547 9,291,547 9,291,547 7,019,981 7,019,981 7,019,981 7,019,981 9,354,185 6,859,935 5,395,673 4,242,582 9,291,547 6,824,489 5,372,619 4,224,697 7,019,981 4,997,897 3,645,053 2,659,822 31 98% 98% 98% 98% 98% 98% 98% 98% 76% 76% 76% 76% 0 90 210 300 0 90 210 300 0 90 210 300 Mean % acres with road patch size (acres) layer and buffer that meet minimum size 100% 1,169,273 73% 107,186 58% 78,198 45% 44,659 100% 154,859 73% 80,288 58% 62,472 45% 40,235 100% 18,093 71% 16,333 52% 16,129 38% 16,834 Figure 10. Natural vegetation core areas defined by the no road, major road, and all road data layers, and a 210 m buffer along roads and non-natural vegetation landcover. 32 Figure 11. Natural vegetation core areas defined by all roads with a 0, 90, 210, 300 m buffer applied to roads and non-natural vegetation landcover. 33 Potentially unchanged natural vegetation core areas - by ecoregional section Description: The potentially unchanged natural vegetation core areas analysis identifies patches with pixels that appear to contain the same vegetation that was recorded in circa 1800. In order to accomplish this, MNFI staff created a table that crosswalks each of the circa 1800 vegetation types (31) to each of the 32 IFMAP level 2 class types. Each of the IFMAP level 2 classes were crosswalked to one of the eleven modified IFMAP vegetation classes created by MNFI, and then lumped together to form an unchanged vegetation data layer. All unchanged natural vegetation patches greater than a threshold size (see below) were selected. These select patches were then buffered inward by 90 meters. Again, patches greater than a threshold size were reselected. Water, which includes lakes, ponds, and large river segments, was originally included as part of the natural vegetation data layer. Once the 90 meter buffer was applied, water bodies greater than 10 acres were subtracted out of the data layer and the remaining patches were regrouped and extracted based on the ecoregional section thresholds. Threshold sizes were set for each of the three ecoregional sections based on the percentage of potentially unchanged natural lands remaining, degree of fragmentation, and mean patch size. Due to the relatively small size of potentially unchanged natural vegetation patches across the state, it was determined that minimum thresholds would be set at 10% of the natural land patch minimum threshold sizes by ecoregional section. Therefore, the UP was set at 500 acres, the NLP was set at 250 acres, and the SLP was set at 50 acres. Six different data layers of potentially unchanged natural vegetation core areas were developed for each ecoregional section based on roads and buffers (for a total of 18 data layers). The 6 data layers were: 1) no roads – no buffer, 2) no roads - 90 m buffer, 3) major roads – no buffer, 4) major roads – 90 m buffer, 50 all roads – no buffer, and 6) all roads – 90 m buffer. Please refer to appendix J for metadata. Use: The potentially unchanged vegetation analysis can be used to identify what appears to be the least modified or altered patches of natural vegetation by ecoregional section. Unchanged vegetation was used to identify those areas that appear to be unchanged between circa 1800 and circa 2000. It can be assumed that these patches have a higher probability of being in high quality condition compared to patches that appear to be changed. This data layer can also be used to analyze unchanged natural vegetation patch statistics such as mean and median patch size, maximum size, and total acreage either at the statewide scale or by ecoregional section. Limitations: As mentioned earlier, IFMAP land cover is limited in accuracy. In addition, vegetation coverage was documented from satellite imagery taken between 1999 and 2001, and some areas have been altered since that time period. The circa 1800 vegetation data layer is based on general land office survey notes taken along section lines in the early to mid 1800’s. This limited information from surveyor notes had to be extrapolated out to the remainder of the section (1 square mile), which means the majority of area within each section is based on scientific interpretation rather than empirical data. File name: unchanged (grid) 34 Data source: Michigan Geographic Framework statewide all roads layer version 5a Lu1800_g – circa 1800 vegetation for entire state of Michigan in grid format Lu2001v2_g – IFMAP circa 2000 land use data for entire state of Michigan in grid format Results: Total area of potentially unchanged natural vegetation in the SLP was 663,803 acres (10.5 % of the statewide total) (Table 11). This represented only 4.3 % of the SLP region. Using 50 acres as a minimum patch size, the total area of potentially unchanged natural vegetation in the SLP dropped to 395,140 acres (a 40% decrease). The mean patch size was 126 acres. When all roads are used to define patch boundaries, and a 300 meter buffer is applied to each road and non-natural landcover, total area of potentially unchanged natural vegetation in the SLP decreased to 6,241 acres. This represents .9 % of the original area of potentially unchanged natural vegetation. These numbers indicate that potentially unchanged natural vegetation in the SLP consists of very small, isolated, and highly fragmented patches that are heavily impacted by both major and minor roads (Figure 12). Total area of potentially unchanged natural vegetation in the NLP was 1,652,985 acres (26 % of the statewide total) (Table 11). This represented only 15 % of the NLP region. Using 250 acres as a minimum patch size, the total area of potentially unchanged natural vegetation in the NLP dropped to 1,071,634 acres (a 35 % decrease). The mean patch size was 1,142 acres. When all roads were used to define patch boundaries, and a 300 meter buffer was applied to each road and non-natural landcover, total area of potentially unchanged natural vegetation in the NLP decreased to 110,485 acres. This represents 6.7 % of the original area of potentially unchanged natural vegetation. These numbers indicate that potentially unchanged natural vegetation in the NLP consists of moderately sized, somewhat fragmented patches that are impacted by minor roads (Figure 12). Total area of potentially unchanged natural vegetation in the UP was 4,032,176 acres (63.5 % of the statewide total) (Table 11). This represented 37.5 % of the UP region. Using 500 acres as a minimum patch size, the total area of potentially unchanged natural vegetation in the UP dropped to 3,273,235 acres (a 29 % decrease). The mean patch size was 4,696 acres. When all roads are used to define patch boundaries, and a 300 meter buffer was applied to each road and non-natural landcover, total area of potentially unchanged natural vegetation in the UP decreased to 668,238 acres. This represents only 16.6 % of the original area of potentially unchanged natural vegetation. These numbers indicate that potentially unchanged natural vegetation in the UP consists of very large, connected patches that are impacted by minor roads. The largest patches of potentially unchanged vegetation in the state are concentrated in the northern half of the eastern UP (Figure 12). 35 Table 11. Summary of potentially unchanged vegetation core areas statewide. Ecoregion road (Modified) layer Total acres of potentially unchanged natural vegetation # of patches UP UP UP UP UP UP NLP NLP NLP NLP NLP NLP SLP SLP SLP SLP SLP SLP 4,032,176 4,032,176 4,025,336 4,025,336 3,894,087 3,894,087 1,648,104 1,648,104 1,652,985 1,652,985 1,510,390 1,510,390 663,803 663,803 660,273 660,273 549,943 549,943 9,386 9,386 9,553 9,553 12,529 12,529 11,661 11,661 11,538 11,538 14,266 14,266 13,690 13,690 13,631 13,631 12,870 12,870 none none major major all all none none major major all all none none major major all all min. patch size (acres) 500 500 500 500 500 500 250 250 250 250 250 250 50 50 50 50 50 50 acres with road layer that meet minimum size % acres with road layer that meet minimum size buffer size in meters 3,273,235 3,273,235 3,253,778 3,253,778 2,820,870 2,820,870 1,071,634 1,071,634 1,059,367 1,059,367 750,674 750,674 395,140 395,140 393,055 393,055 294,670 294,670 81% 81% 81% 81% 72% 72% 65% 65% 64% 64% 50% 50% 60% 60% 60% 60% 54% 54% 0 90 0 90 0 90 0 90 0 90 0 90 0 90 0 90 0 90 36 acres with % acres with Mean road layer road layer patch and buffer and buffer size that meet (acres) minimum size 3,273,235 941,821 3,253,778 845,117 2,820,870 668,238 1,071,634 213,692 1,059,367 210,306 750,674 110,485 395,140 10,963 393,055 10,787 294,670 6,241 100% 29% 100% 26% 100% 24% 100% 20% 100% 20% 100% 15% 100% 3% 100% 3% 100% 2% 4,696 1,351 4,253 3,422 2,345 1,877 1,142 1,068 1,101 922 590 511 126 169 125 166 103 104 Figure 12. Potentially unchanged vegetation core areas defined by no road, major road, and all road data layers with a 0 m buffer. 37 Large Functional Landscapes Description: The large functional landscape analysis is a statewide look at natural vegetation core areas, without differentiating by ecoregion. In that sense, these patches are identical to the patches created for the natural vegetation core areas analysis. The difference is that these natural vegetation core areas were selected based on the matrix community criterion of 5,000 acres or greater as defined by The Nature Conservancy (Anderson et al. 1999). Patches greater than the minimum threshold are buffered inward using three different buffer sizes (90 m, 210 m, 300 m). After buffering, patches greater then the 5,000 acre criterion were reselected and retained. Water, which includes lakes, ponds, and large river segments, was originally included as part of the natural vegetation data layer. After each buffer was applied, water was subtracted out of the landscape, and the remaining patches regrouped, and reselected based on the minimum size threshold of 5,000 acres. Twenty-four different data layers were developed for large functional landscape patches based on different road and buffer combinations. Twelve data layers included water in the analysis, and the remaining 12 did not include water in the analysis. The 12 data layers are: 1) no roads – no buffer, 2) no roads - 90 m buffer, 3) no roads – 210 m buffer, 4) no roads – 300 m buffer, 5) major roads – no buffer, 6) major roads – 90 m buffer, 7) major roads – 210 m buffer, 8) major roads – 300 m buffer, 9) all roads – no buffer, 10) all roads – 90 m buffer, 11) all roads – 210 m buffer, and 12) all roads – 300 m buffer. The remaining 12 data layers are the same as above except that water was removed from the analysis. Please refer to appendix K for metadata. Use: The purpose of this analysis was to identify the largest most intact areas of natural vegetation in the state – sites that have the potential to function as matrix communities now or in the future. All natural vegetation types were combined to create one natural vegetation data layer. The reason for combining them together is that matrix communities typically contain numerous large and small patch natural community types. Limitations: As mentioned earlier, IFMAP land cover is limited in accuracy. In addition, vegetation coverage was documented from satellite imagery taken between 1999 and 2001. Some areas have been altered since that time period. File names: natveg2_matrix (grid) natveg2_matrix\_water_out (grid water removed) Data source: Michigan Geographic Framework statewide all roads layer version 5a Lu2001v2_g – IFMAP circa 2000 land use data for entire state of Michigan in grid format. 38 Results: Total area of natural vegetation in the state of Michigan (including water) was 22,084,814 acres (Table 12). Using a minimum patch size of 5,000 acres to define large functional landscapes, and removing water from these patches, the area of natural vegetation decreased to 18,749,300 (a 15 % decrease). When all roads were used to define patch boundaries, and a 300 m buffer was applied to each road and non-natural landcover, large functional landscapes decreased to 2,825,288 acres, or 12.8% of the original area of natural vegetation in the state. This demonstrates that 85% of the vegetation in Michigan is considered to be part of a large functional landscape patch, and that minor roads have a high impact on large functional landscapes in the state. The vast majority of large functional landscape patches are located in the UP (Figures 13 and 14). Table 12. Summary of large functional landscape patches statewide. Matrix Vegetation water removed Matrix Matrix Matrix Matrix Matrix Matrix Matrix Matrix Matrix Matrix Matrix Matrix road Total acres of # of layer natural patches vegetation none none none none major major major major all all all all 22,084,814 22,084,814 22,084,814 22,084,814 22,029,008 22,029,008 22,029,008 22,029,008 20,909,709 20,909,709 20,909,709 20,909,709 18,634 18,634 18,634 18,634 19,657 19,657 19,657 19,657 45,280 45,280 45,280 45,280 min. size (acres) 5,000 5,000 5,000 5,000 5,000 5,000 5,000 5,000 5,000 5,000 5,000 5,000 acres with % acres buffer road layer with road size in that meet layer that meters minimum meet min. size size acres with road layer and buffer 18,749,300 18,749,300 18,749,300 18,749,300 18,239,182 18,239,182 18,239,182 18,239,182 8,934,329 8,934,329 8,934,329 8,934,329 18,749,300 11,437,341 7,025,589 4,994,044 18,239,182 11,270,153 6,954,357 4,957,123 8,934,329 6,123,369 3,927,944 2,825,288 39 85% 85% 85% 85% 83% 83% 83% 83% 43% 43% 43% 43% 0 90 210 300 0 90 210 300 0 90 210 300 % acres with road layer and buffer that meet minimum size 100% 61% 37% 27% 100% 62% 38% 27% 100% 69% 44% 32% Figure 13. Large functional landscape patches defined by all roads with 0, 90, 210, and 300 m buffers applied. 40 Figure 14. Large functional landscape patches defined by no road, major road, and all road data layers with a 210 m buffer applied. 41 Fine Filter - Element Occurrence Data Description: The Michigan Natural Features Inventory has been inventorying and tracking Michigan’s threatened, endangered, and special concern species and high quality natural communities since 1979. As of September, 2006, MNFI tracked 417 plant species, 248 animal species, and 74 natural community types. In addition to species and natural communities, MNFI also tracks other natural features such as colonial bird nesting colonies and significant geological features. The tracked species include those with Federal and State legal protection and special concern species, which have no legal protection. Like the special concern species, natural communities also have no legal protection status. As of September, 2006, The MNFI database contained approximately 14,532 records of these natural features (plants, animals, and natural communities). Data sources include museum and herbarium collections, published reports, MNFI field surveys, and information from cooperators. Database records span a range from historic information to very current information from the latest field season. The MNFI database is continually being updated and is the most complete record of Michigan’s sensitive species and natural features. The MNFI database is a Natural Heritage database and utilizes Natural Heritage methodology and data standards originally designed by The Nature Conservancy and now maintained by Natureserve (www.natureserve.org). The MNFI database is more than a presence/absence database. Among other information, it contains dates of sightings, global and state imperilment rankings for species, and a quality (or viability) ranking for individual occurrences. Definitions of the global and state (or subnational) rankings can be found in appendix A. The quality ranking is an A – D scale with A being the highest quality. Other codes such as E for extant, H for historic, and X for extirpated are also used. Extant is used when not enough information is available to assess population viability. The standards for applying a quality rank to an occurrence vary by species and community, but generally fall into three main categories: size, condition, and context. See the chapter entitled approach for more information. Limitations: The primary limitations to MNFI’s element occurrence database are: 1) it contains static information – each element occurrence is updated infrequently 2) a lack of a statewide systematic survey, and 3) the presence of very old and/or general (non location specific) records. Biological information from the field is collected annually from MNFI staff and other reliable contributors. Once this information is entered into the database, it may be decades before it gets updated. For example, approximately 36 % of the records in the database are over 20 years old. More significantly, there has never been a systematic survey of element occurrences in the state. This means that something can be said about the biological significance of an area containing element occurrence records, however nothing can be said definitively about the biological significance of areas with no known element occurrence records. This is where the quote “absence of evidence is not evidence of absence” comes into play. Related to this, is that there have been small areas of the state that have been systematically surveyed; however they are predominantly owned by public agencies or non-governmental organizations such as The Nature Conservancy. 42 Fine Filter - Element Occurrence Data Analysis EO Frequency Count Description: The EO frequency count is a count of all element occurrences that fall within a given public land survey system (PLSS) section. The model utilizes a statewide GIS data layer (Environmental Systems Research Institution (ESRI) shapefile) of the PLSS sections. A numeric count field is added to the section shapefile theme table. Each section shape is selected in turn and intersected with the MNFI GIS database. The number of occurrences intersecting each section shape is counted and that value is calculated into the count field in the section shapefile theme table. A cutoff date of September 1, 2006 was used to create the EO frequency datasets. All records added to the Michigan Natural Features database after this date are not included in this analysis. A total of 6 data layers were developed for the terrestrial EO frequency count. They are differentiated by element categories and last observed dates. The 9 data layers are: 1) all species (no natural communities) – all dates, 2) all species (no natural communities) – only dates > 1985, 3) only terrestrial species – all dates, 4) only terrestrial species – only dates > 1985, 5) all element occurrences – all dates, and 6) all element occurrences – only dates > 1985. Use: The EO frequency count is a relatively simple representation of the MNFI data. It is designed to show users where there are concentrations of known species or natural community occurrences in the MNFI database. While the EO frequency count provides limited information, it does fulfill its intended purpose. Users can see if there are known occurrences in the vicinity of a proposed project or delineate those areas where there are concentrations of occurrences. All species information is removed so locations of particularly sensitive species cannot be determined from the model. Limitations: The primary disadvantage is that it provides very limited information. The user only knows that the known boundary of an occurrence overlaps the boundary of the area of interest. No allowance is made for the age of the record, relative importance of the species, or the extent of potential habitat within the occurrence boundary. File names: Ter_EO_trs_0906.shp freq_ter_trs_v9-06.shp Data source: Biot_p – Biotics polygon database created directly from Biotics from version created September 1, 2006. Results: Values vary depending on which dates and natural feature elements are utilized in the analysis. Using only terrestrial species and all last observed dates in the database, frequency values for PLSS sections range from a low of 0 to a high of 65 (Figure 15). Using the Jenk’s optimization classification method to define groupings, < 1 % (51) of all PLSS sections fell into the highest category (scores 34 65). Geographic areas that fell into the highest category included: northern half of Isle Royale, southwest corner of the Lower Peninsula, and eastern Washtenaw County (figure 15). 43 Figure 15. Frequency of rare terrestrial species using all last observed dates. 44 EO Likelihood Description: The overall modeling process of EO likelihood consists of grouping species into habitat guilds, creating a habitat layer for each guild, using the habitat layer to redefine the spatial extent of the appropriate occurrences, intersecting the spatially redefined occurrences with political boundaries (PLSS unit), and then assigning each political unit a likelihood value. The process starts by grouping species into habitat classes and assigning a habitat identifier code to each species occurrence. Features in the MNFI database other than species and natural communities, such as geological formations, are removed from the analysis. Next a habitat layer is created for each habitat class. The habitat layers are then used to redefine the spatial extent of the occurrences. This is accomplished by selecting all the occurrences with a given habitat code then clipping the selected occurrences using the appropriate habitat layer as the clipping overlay theme. The result of this operation produces a new theme for each habitat group. In each new theme the spatial extent of each occurrence is replaced by the spatial extent of the habitat within the original boundary of the occurrence. The new theme retains all the database attributes of the original occurrence database. Where fragmented habitat patches occur within an occurrence boundary, the occurrence will be converted from a single shape to multiple shapes. The clipping operation was not performed on natural community occurrences because the communities have a defined spatial extent. The natural communities are selected out of the occurrence database and converted to a separate layer. The themes for each habitat group and the natural community themes are then all merged together. After merging the themes for each habitat type into a single theme, the merged theme is dissolved on the unique code number assigned to each individual occurrence. This operation consolidates all the separate shapes for each occurrence into a single shape. Each occurrence is then assigned a value based on the age of the record. This value is used to represent the likelihood of the occurrence still existing. Occurrences with a last observed date of no later than 1982 are assigned a value of one, occurrences between 1970 and 1982 are assigned a value of 0.5, and occurrences prior to 1972 are assigned a value of 0.25. All natural community records are assigned a value of one. To create the EO likelihood value for the PLSS data set, all records in the PLSS data set are selected and assigned a “No Status” value. Next the records in the species database with the lowest likelihood of still existing (value = 0.25) are selected. The PLSS data set is intersected with the species database and the selected PLSS records are assigned a value of “Low.” Next those records with a moderate likelihood of still existing are selected (value = 0.5). The PLSS data set is intersected with the species database and the selected PLSS records are assigned a value of “Moderate.” Finally the records in the species database with the highest likelihood of still existing (value = 1) are selected. The PLSS data set is intersected with the species database and the selected PLSS records are assigned a value of “High.” Performing the selections and intersections in this order insures that a higher likelihood value in any PLSS feature will override a lower likelihood value. A cutoff date of September 1, 2006 was used to create EO likelihood datasets. All records added to the Michigan Natural Features database after this date are not included in this analysis. A total of six data layers were developed for the terrestrial EO likelihood count. They are differentiated by element categories and last observed dates. The six data layers are: 1) all species (no natural communities) – all dates, 2) all species (no natural communities) – only dates > 1985, 3) only terrestrial species – all dates, 4) only terrestrial species – only dates > 1985, 5) all element occurrences – all dates, and 6) all element occurrences – only dates > 1985. 45 Use: The EO likelihood model is designed to help protect biodiversity and minimize potential regulatory problems by directing development away from those areas with a high likelihood of encountering a sensitive species. Because no specific species information is presented, the model reduces the sensitivity of the underlying MNFI data. A high probability indicates that the area of interest contains the spatial extent of an occurrence, there is potential habitat within the area, and the occurrence has been observed in the recent past. A low probability indicates that the area contains the spatial extent of an historic species occurrence and there is potential habitat within the area. While the low likelihood indicates that the underlying occurrences are historic, there is still a possibility that the species persists in appropriate habitat. In the recent past, MNFI botanists have reconfirmed three 100 year old plant records. A moderate likelihood indicates, by default, something between the other two values. The EO likelihood model provides users with a higher level of information than the simple EO frequency count. Unlike the EO frequency count, which only implies that the extent of an occurrence lies within an area of interest, the EO likelihood model delineates those areas where there is a higher likelihood of encountering a known occurrence of a sensitive species or natural community. Also, by utilizing potential habitat within the known extent of the occurrences, areas without potential habitat are eliminated from consideration. The EO likelihood model can be used in the context of both land use planning efforts and conservation planning efforts. By delineating areas with a high likelihood of encountering a sensitive species or natural community, the model can be used to direct development away from those areas, or to identify areas worthy of conservation efforts. Limitations: One shortcoming of the EO likelihood model is that all high likelihood areas are treated the same. Whether there is one recent occurrence in the area or thirty recent occurrences, the same high likelihood value is assigned to the area. There is also no allowance for the relative imperilment of the species found in any unit of interest, and there is no numeric value assigned to any of the units of interest that allow them to be compared to each other. File names: Ter_EO_trs_0906.shp likelihood_ter_trs_v9-06.shp Data source: Biot_p – Biotics polygon database created directly from Biotics from version created September 1, 2006. Results: The number of PLSS sections that fell into any one category varied depending on the last observed dates of the natural feature elements used in the analysis. Using only rare terrestrial species and all last observed dates in the MNFI database, 17 % of all PLSS sections in the state fell into the high probability category (Figure 16). 46 Figure 16. Likelihood of a known rare terrestrial species occurrence still occurring in its last observed location using all last observed dates. 47 Bio-rarity Score Description: In addition to the EO likelihood value described above, each element occurrence is also assigned three other values based on: 1) the species global status, 2) the species state status, and 3) on the occurrence’s viability rank. The greater the threat of imperilment to the species, the higher the value assigned to the occurrence. In a similar manner, the higher the quality or viability of each occurrence, the higher the value assigned to it. The biodiversity value of each occurrence is then calculated by adding the values for the global status, state status, and the quality ranking, then multiplying the sum by the EO likelihood value described above. To calculate the biodiversity value of a given PLSS feature, each feature in the PLSS theme is selected in sequence. Next, all the species occurrences intersecting the PLSS feature are selected. The biodiversity values of the selected species occurrences are summed and assigned to the PLSS feature. The result is a value for each PLSS unit that is the sum of the biodiversity values of all occurrences falling within the PLSS unit. A cutoff date of September 1, 2006 was used to create the bio-rarity datasets. All records added to the Michigan Natural Features database after this date are not included in this analysis. A total of six data layers were developed for the terrestrial bio-rarity score. They are differentiated by element categories and last observed dates. The six data layers are: 1) all species (no natural communities) – all dates, 2) all species (no natural communities) – only dates > 1985, 3) only terrestrial species – all dates, 4) only terrestrial species – only dates > 1985, 5) all element occurrences – all dates, and 6) all element occurrences – only dates > 1985. Use: Unlike the EO likelihood model, the bio-rarity score allows similar areas to be compared to each other to determine their relative contributions to biodiversity. Because resources for conservation are generally limited, the bio-rarity score can help direct limited resources to those areas where the resources will have the greatest conservation impact. Limitations: As with other element occurrence based information, this data layer is limited by: 1) static information, which is updated infrequently, 2) incomplete data, and 3) old and/or general (non location specific) records. File names: Ter_EO_trs_0906.shp br_ter_trs_v9-06.shp br_ter85_trs_v9-06.shp Data source: Biot_p – Biotics polygon database created directly from Biotics from version created September 1, 2006. Results: Values vary depending on which dates and natural feature elements are utilized in the analysis. Using only rare terrestrial species and all last observed dates in the MNFI database, bio-rarity values for PLSS sections range from a low of 0 to a high of 357.88. Using quantiles to statistically define groupings, PLSS sections with bio-rarity scores > 23.13 fell into the top 10 % of scores. A few spatially distinct areas that fell into the highest category included: northern half of Isle Royale, Allegan 48 State Game Area, Fort Custer Recreation Area, southeast Newaygo County, southern Oceana and northern Muskegon Counties, northern Lake Michigan and Lake Huron shorelines, and the central high plains of the northern Lower Peninsula (Figure 17 and 18). These results may be due to survey bias and/or the naturally high concentrations of natural features in these areas. Best two occurrences of each terrestrial species by sub-subsection Description: The two highest ranking occurrences of each rare terrestrial plant and animal tracked by MNFI were identified for each sub-subsection (as described by Albert et. al., 1995). There are a total of 398 terrestrial plants (appendix A) and 174 animals (appendix B) currently tracked by MNFI. There are a total of 38 sub-subsections, plus 7 sub-sections that do not contain any sub-subsections, in Michigan (for a total of 45 units used in this analysis). A cutoff date of September 1, 2006 was used to create this dataset. All records added to the MNFI database after this date were not included in the analysis. Use: In some cases, important element occurrences may be located outside areas deemed significant due to other natural assets such as size, intactness, connectivity, and quality. Identifying areas with high quality element occurrences regardless of natural vegetation quality or landscape context can be important for ensuring adequate biological representation, and in turn protecting potential genetic variability. How many occurrences of each element are enough for sufficient representation is a difficult question to answer. Two was chosen simply because it is more than one. However, given 45 units and the wide geographic range of some of these species and communities, 2 element occurrences per unit could theoretically add up to 90 occurrences of each element statewide. Limitations: As with the other element occurrence based information, this data layer is limited by: 1) static information, which is updated infrequently, 2) incomplete data, and 3) old and/or general (non location specific) records. File names: best2_ter_subsubsection_trs_0906.shp best2_ter_subsub_summed_trs_0906.shp Data source: Biot_p – Biotics polygon database created directly from Biotics from version created September 1, 2006. Results: As a result of this analysis, 3,768 occurrences (out of 9,985 total terrestrial element occurrences) were identified as one of the best two occurrences of each terrestrial species by sub-subsection. This represents approximately 38% of all terrestrial species element occurrences.The three subsubsections with the highest number of best two terrestrial element occurrences are: 1) Battle Creek Outwash Plain (340 EO’s), 2) Maumee Lake Plain (236 EO’s), and 3) Southern Lake Michigan Lake Plain (231 EO’s). All three are located in the southern Lower Peninsula (Table 13, Figure 19). 49 Fig. 17. Bio-rarity scores for all element occurrences using all last observed dates - top 10%. 50 Figure 18: Bio-rarity scores for rare terrestrial species with last observed dates > 1985 - top 10%. 51 Table 13. Total number of best two terrestrial element occurrences by sub-subsection or subsection. Subsubsection or subsection Name of sub-subsection or subsection 0 611 612 613 621 622 631 632 633 641 642 651 652 660 711 712 721 722 723 730 740 751 752 761 762 763 811 812 813 821 822 831 832 833 910 920 931 932 950 961 962 963 971 972 973 980 Total Maumee Lake Plain Ann Arbor Moraines Jackson Interlobate Battle Creek Outwash Plain Cassopolis Ice-Contact Ridges Berrien Springs Southern Lake Michigan Lake Plain Jamestown Lansing Greenville Sandusky Lake Plain Lum Interlobate Saginaw Bay Lake Plain Standish Wiggins Lake Cadillac Grayling Outwash Plain Vanderbilt Moraines newaygo Outwash Plain Manistee Williamsburg Traverse City Onaway Stutsmanville Cheboygan St. Ignace Rudyard Escanaba/Door Peninsula Seney Sand Lake Plain Grand Marais Sandy End Moraine and Outwash Northern Lake Michigan Till Plain Gwinn Deerton Spead Eagle-Dunbar Barrens Michigame Highland Brule and Paint Rivers Winegar Moraine Lac Veaux Desert Outwash Plain Gogebic-Penokee Iron Range Ewen Baraga Gay Calumet Isle Royale Lake Superior Lake Plain 52 Total # of best 2 terrestrial EO's 31 236 199 185 340 172 150 231 22 197 61 101 23 83 44 5 42 92 57 108 137 7 41 73 14 84 159 31 90 72 100 53 14 11 39 67 13 46 3 45 24 18 13 101 113 21 3,768 Figure 19. Best two terrestrial element occurrences by sub-subsection or subsection. 53 High quality natural communities Description: The MNFI database contains records of high quality and/or rare natural communities. Currently, MNFI tracks 74 different natural community types (Appendix B). As of September 28, 2006, the database contained 1,371 natural community records which represent approximately 9% of the total records for plants, animals, and natural communities. High quality natural communities were defined as those communities with a B/C element occurrence rank or higher. A “C” ranked community, which was not included in the high quality category, means that the natural community is moderately degraded and long-term viability is estimated to be fair. A cutoff date of September 28, 2006 was used to create this dataset. All records added to the MNFI database after this date were not included in this analysis. Use: High quality natural communities represent the best, most viable known occurrences of the 74 different natural community types found in Michigan (as recognized by MNFI). Natural communities are important because they provide the environment necessary for plants and animals to persist and evolve over the long-term. High quality natural communities provide the genetic material needed for changing environmental conditions and restoration projects. They also are a good benchmark for guiding the planning, implementation, and monitoring of natural community restoration and management projects. Limitation: As with the other element occurrence based information, this data layer is limited by: 1) static information, which is updated infrequently, 2) incomplete data, and 3) old and/or general (non location specific) records. In addition, EO ranks for natural communities have a certain degree of inconsistency due to human judgment, changes in EO rank specifications over time, and an emphasis on qualitative criteria. In addition, approximately 64 natural community occurrences were missing acreage information. File names: community_w_best_attributes.shp natcomm_bcrank.shp Data sources: Biot_p – Biotics polygon database created directly from Biotics from version with a last observed date of 09/28/2006. Biot_x – Biotics point database created directly from Biotics from version with a last observed date of 09/28/2006. Results: Of the 1,371 natural community element occurrences in the MNFI database, 68 % (932) of these occurrences had an element occurrence rank of BC or higher (A, AB, B, BC) (Table 14). These ranks were interpreted to mean that these occurrences are high quality and viable over a long period of time. The spatial extent of natural communities with a BC rank or higher totaled 390,919 acres. This represents approximately 1 % of the landscape in Michigan (Figure 20). 54 Table 14. Summary of high quality natural communities with an EO rank of > BC. Total # of natural community EO's 1,371 Total acres 431,964 Total # > B/C rank 932 % of Total acres total > B/C rank 68% 390,919 % of Total acres 90% Best occurrences of each natural community type Description: The highest quality occurrences of Michigan’s 74 natural community types (Appendix B) were identified at four scales: statewide, Ecological section, Ecological subsection, and Ecological subsubsection (Albert 1995). At each scale, the three highest-quality examples of each community type were identified. The rankings were nested so that the highest quality occurrence of a natural community at the broad scale (statewide or section) was also the highest quality occurrence at the appropriate local scale (subsection and sub-subsection). It is important to note that the MNFI natural community classification was revised after this analysis was completed. Rankings were primarily based on existing EO data in the MNFI database. All occurrences of each natural community type are ranked according to condition/quality, size, and landscape context. Rankings for these factors are combined to calculate an overall EO Rank on an A-D scale, with Aranked occurrences representing the highest quality sites, C-ranked occurrences meeting the minimal standards for a community to be included in the MNFI database as an element occurrence, and Dranked communities representing occurrences for rare communities that are not represented by any A-C ranked (high quality) examples. For the purposes of determining the highest quality EOs for each community type, C-ranked occurrences were generally omitted from consideration, unless no Aor B-ranked occurrences were documented at a particular scale. Due to the fact that element occurrences have been documented and ranked by different surveyors over the course of approximately 25 years, and given that the tools and methods for assessing community quality have evolved over that time, A-ranked occurrences were not necessarily assumed to be of higher quality than AB- or B-ranked occurrences. For each community type, field notes and the most recent aerial photographs (1998) were consulted to identify the highest quality occurrences at each scale. Digital maps for each occurrence were checked against hand-drawn maps for accuracy, and, in many cases, occurrences were remapped. Two primary reasons for remapping were inaccurate digitization of the original maps and post-survey changes in spatial extent of occurrences due to anthropogenic disturbance or development. In some instances, significant changes in acreage associated with remapping warranted lowering or raising the overall EO Rank. In some cases, especially for natural communities with a high number of occurrences (e.g., prairie fen, bog), there are many occurrences of equal rank at one or multiple scales. For example, in subsection 7.3, there are five B-ranked bog occurrences. In sub-subsection 6.1.3, there are three Aranked prairie fen occurrences. In these instances, occurrences were ranked relative to each other based on the best available information regarding condition/quality, size, and landscape context, in the following manner. 55 Figure 20. High quality natural communities with an EO rank of >B/C. 56 1) Condition Condition ranking was based primarily on field notes and interpretation of aerial photos. Community intactness, structure, anthropogenic impacts, presence/abundance of invasive species, vascular plant species diversity, and presence/representation of typical, indicator, or rare vascular plant species were assessed. Community intactness and anthropogenic impacts were assessed from field notes in addition to inspection of 1998 aerial photographs. Information on invasive species, structure, diversity, and presence of rare species relied on existing field notes, although some aspects of structure could be confirmed through inspection of aerial photographs. One caveat particular to species-level data is that some community occurrences were more thoroughly surveyed (either spatially or temporally) than other sites. Therefore, apparent differences in diversity between one site and another of similar rank may be an artifact of sample effort rather than an actual biological difference between the sites. In addition, the manner in which a community occurrence is mapped often affects its condition rank, which in turn affects the overall EO Rank. 2) Landscape context Historically, high quality occurrences of natural communities in the MNFI database were ranked primarily or entirely based on condition/quality ranks, as long as minimum size criteria were met. However, the field of conservation has moved towards landscape-level approaches, and landscape context is vitally important to viability of natural community occurrences and the conservation of biodiversity over the short and especially long term. A 30-acre old-growth mesic southern forest bordered by residential development on all sides is not as viable as a 30-acre old-growth mesic southern forest surrounded by 150 acres of second- and third-growth forest in an agricultural setting. For sites of similar EO ranks, landscape context was used to determine the highest quality occurrences. Landscape context consists of two levels: buffer of associated natural communities and overall landscape condition. These levels were broken down using the following criteria, ranked from best condition to worst condition. • Landscape buffer condition o Buffered by associated natural communities o Buffered, but not by associated natural communities o Agricultural buffer o Developed buffer • Overall landscape condition o Natural - Landscape is largely natural cover. o Partially agriculture - EO in partially agricultural landscape. o Agriculture - EO in predominantly agricultural landscape. o Urban - EO surrounded in part or wholly by urban/suburban development, regardless of remaining buffer type (natural, agriculture). Buffers were visually inspected using 1998 aerial photographs for high quality EOs in order to determine the highest quality community occurrence of a similar or identical EO Rank. In some instances, landscape context was poor enough to warrant lowering the overall EO rank for particular occurrences. This was especially true in fast-developing regions of the state where large amounts of land were converted from natural cover or agriculture to suburban and exurban development in the time between the original date the community occurrence was surveyed and 1998. 57 3) Size Sites of small size are more vulnerable to successional changes, dominance by exotic species, and “island” effects than sites of large size. Large sites are more likely to support higher-level ecosystem functions and are less vulnerable to local extirpations and elimination via natural or non-natural successional processes. Size was used as a tiebreaker if condition and landscape context were of similar rank, or if an overwhelming difference in size balanced out slightly lower condition and landscape context ranks. Due to differences in element occurrence mapping strategy used by different surveyors, size was checked against condition to ensure apparent “size” based on EO acreage accurately reflected the size of the high quality community occurrence. One consideration outside of the traditional EO specifications was also used to determine which occurrences were highlighted. Certain communities (e.g., prairie fen) are characterized by subtypes. If these subtypes are unique (e.g., lakebed marl fens dominated by calciphiles vs. streamside prairie fens dominated by prairie forbs and grasses), representation of the variation on the landscape was addressed in situations where high quality occurrences of more than one subtype existed. Based on the above criteria and considerations, the three highest quality element occurrences for each of the 74 natural community types currently known from Michigan were identified at each of the four scales (statewide, Section, Subsection, Sub-subsection), with the understanding that the accuracy of this assessment is limited by the amount of biological information available for each occurrence. Use: High quality natural communities represent the best, most viable known occurrences of the 74 different natural community types found in Michigan (as recognized by MNFI). Natural communities are important because they provide the environment necessary for plants and animals to persist and evolve over the long-term. High quality natural communities provide the genetic material needed for changing environmental conditions and restoration projects. They also are a good benchmark for guiding the planning, implementation, and monitoring of natural community restoration and management projects. Limitations: Determination of the highest-quality examples of each community type relies on existing information, some of which dates to the early 1980s. Selection of approximately three occurrences for each community at each scale should compensate for the potential uncertainty relating to accuracy. Quality (especially with regards to vascular plant diversity) relies heavily on sampling method and effort. High diversity in particular community occurrences will be incorporated into the overall ranking system, but apparent low diversity (e.g., short plant species lists) often reflects sampling effort rather than complete biological inventory. File names: natcomm_combined.shp natcomm_state.shp natcomm_section.shp natcomm_subsections.shp natcomm_subsubsection.shp 58 Data source: Biot_x – Biotics point database created directly from Biotics from version with a last observed date of 09/28/2006. Results: Further analysis needs to be completed by identify any significant trends regarding the three best occurrences of each natural community type at the statewide scale (Figure 21). Areas of relatively high concentrations of high quality natural communities included: 1) Pinckney-Waterloo Recreation Areas, 2) southwestern Huron County (along the Saginaw Bay shoreline), 3) northern Marquette County, and 4) the tip of the Keweenaw Peninsula. 59 Figure 21. Three best occurrences of each natural community type at the statewide scale. 60 Aquatic Biodiversity Assessment Methodology Introduction The analysis used in the assessment of Michigan’s aquatic biodiversity was based on two major categories of data: Landscape-based classifications for ecosystems and element occurrences of natural features. Since MNFI does not currently track aquatic natural communities, the aquatic assessment had to rely heavily on previously developed classifications and data by other entities. The two landscape-based ecosystem classifications were developed from multiple projects. The river classification framework used was first proposed by Seelbach et al. (1997) and was then revised by Brenden et al. (2008). This latest version was modeled using expert opinion as the final review. The lake classification framework used was developed by Higgins et al. (1998). Both of these frameworks are based on landscape-level data. The element occurrence dataset is a continuously updated database developed and maintained by the Michigan Natural Features Inventory (MNFI). Using the ecosystem classification frameworks, we developed new data layers that can be used to identify and prioritize potentially unique aquatic ecosystems, important areas for Great Lakes migrating species, intact headwater watersheds, and functional sub-watersheds. The MNFI element occurrence database identifies places on the land that contain unique elements of biodiversity – rare species and high quality natural communities, which MNFI refers to as element occurrences (EOs). The database, which is updated periodically throughout the year, contains a wealth of detailed information that was used to identify and prioritize areas based on frequency, likelihood of persistence, viability, and/or rarity of EOs. Both aquatic ecosystems and EOs of natural features are discussed in more detail below. Categories of ecosystem level datasets developed by this project: 1. Unique river and lake ecosystems – by EDU and statewide 2. High quality rivers and lakes – by EDU and statewide 3. Rivers with access to the Great Lakes 4. Level of intactness of headwater watersheds – statewide 5. Functional sub-watersheds and watersheds – statewide Categories of MNFI EO based datasets developed by this project: 1. EO frequency count 2. EO likelihood 3. Bio-rarity score 4. Rare species richness by sub-watershed 5. Species of greatest conservation need richness by sub-watershed 6. Best two occurrences of each rare aquatic species by watershed For a list aquatic datalayers and descriptions see Appendix M. The list of EO based aquatic datalayers can be found in appendix L. Defining uniqueness Defining what is rare or unique is often subjective and can be difficult to quantify. Rare species are often determined using geographic distribution, habitat specificity, and population size (Rabinowitz 1981, Rabinowitz et al. 1986). However, community rarity or uniqueness has received much less attention (Izco 1998). We do not know how many ecosystems are needed to ensure continued persistence but we expect that frequency of occurrence and geographical range are important components. Uniqueness is affected by the number of individual ecosystems, the classification framework used, and how uniqueness is defined. 61 We define ecosystem uniqueness using geographic range and frequency of occurrence. We considered those ecosystem types occurring in only one watershed statewide as having a restricted geographic range and hence unique. Additionally, we defined uniqueness as those lakes or rivers that have the fewest occurrences and that make up 5% and 1% of the total number of lakes or rivers in Michigan or within an EDU. We defined uniqueness using the 5% and 1% to provide options. We felt that this scheme captured what we intuitively felt was unique or rare, and that it was easily applied to different classifications. If new classifications are introduced in Michigan this analysis could be easily reassessed. Determining representation There is little guidance for abundance and distributional goals for the preservation of ecosystems. Although most literature agrees that smaller and rarer ecosystems should be represented in higher quantities across the landscape than larger and more common ecosystems, specific numbers are not agreed upon. One school of thought suggests using a percentage of the historic distribution, but these percentages vary greatly from 10% to 40% (Tear et al. 2005). Even following the lowest percentage could require large numbers of sites to be protected which could be impractical and unmanageable. Yet others suggest targeting a specific number of ecosystems, but again these numbers vary and can seem too limited (Smith et al. 2001). The question of how much is enough to protect species and ecosystems is unknown. We chose to represent all unique ecosystems, as well as high quality common ecosystems as follows: 10 small rivers, 5 medium rivers, 1 large river, 10 unconnected ponds or small lakes, 5 connected ponds or small lakes, 5 medium lakes, and 1 large lake within each EDU. These were minimum quotas to ensure representation. However, when there were ties in the scores for quality, all occurrences with that score were selected. Determining quality It should be noted that all of the quality analysis conducted on aquatic ecosystems in this report rely on the surrounding terrestrial landscape and not field data. Aquatic ecosystems are so tied and intricately linked to the surrounding lands and watershed that it is difficult to separate the aquatic ecosystem from the terrestrial landscape. The coarse filter approach is generally based on identifying areas of land that have intact natural processes. For terrestrial ecosystems it is relatively easier to determine the size needed to allow for natural processes to occur in different types of ecosystems or natural communities. Aquatic ecosystems are inherently more difficult to assess because the surrounding landscape has such a direct influence. For example, it is easy to draw a boundary around a lake. The natural processes that function within that lake are sediment and nutrient dynamics, internal water movements, water retention, turbidity, water temperature, and oxygen concentration, to name a few. Since most of these processes rely on external inputs from the landscape or water bodies within the watershed of the lake, the lake can not function without these external inputs. Because these inputs are difficult and time-consuming to gather information on, we had to rely on landscape or terrestrial surrogates to determine if natural ecosystem processes are occurring in the aquatic systems. Coarse-Filter: Aquatic Ecosystem Data Watershed and sub-watershed defined Description: Throughout this document we use the terminology watershed and sub-watershed. Watershed is defined as the 8-digit hydrologic units or HUC’s, and sub-watershed is defined here as the 12-digit HUC’s, sometimes called sub-basins. There are 57 watersheds and 2,319 sub-watersheds in Michigan. 62 Limitations: Hydrologic units (or HUC’s) were initially delineated to break the state up into similarly sized units based on hydrology. These units are often termed sub-watersheds. However, they are not hydrologically accurate. A true watershed is defined by all waters draining from an area to a particular point. HUC’s often break up true watersheds such that a point in a HUC can actually get all of its water from a completely different HUC. We used HUC’s as a way to summarize the data with full knowledge that the use of these units does not provide a full picture of the area needed to protect or manage for important species or ecosystems. File name: mi_subwatersheds.shp Data source: The 8 digit HUCs are from the National Hydrography Dataset (NHD). The 12 digit HUCs are from the DEQ, but they did not cover all of Michigan. Parts of Ohio and Indiana’s 12 digit Watershed boundary units (WBUs) were used to fill in the missing area, and the final layer was clipped to the Michigan state boundary. River classification Description: Riverine ecosystems were delineated using river valley segments (VSECs) as defined by the DNR Fisheries Division as of August 2007 (Seelbach et al. 1997, Brenden et al. 2008). VSECs are relatively large stretches of river that have similar hydrology, limnology, channel morphology, and riparian dynamics. VSECs often change at stream junctions or landform boundaries. VSECs use catchment size, hydrology, water chemistry, water temperature, valley character, and channel character as the basis for delineation. VSECs are made up of reaches, which are segments with similar hydrologic characteristics, such as a stretch of stream between two confluences or a lake. A reach is the smallest unit in the hydrology data layer. VSECs defined the boundaries of river ecosystems in this analysis. The classification we used to determine different types of river ecosystems is based on size, water temperature, and gradient. Physical, chemical, and biological changes occur on a longitudinal gradient from the headwaters to the very large rivers (Vannote et al. 1980). Headwaters and small tributaries tend to be shaded and rely on energy inputs from riparian vegetation; their macroinvertebrate communities tend to be dominated by shredders. Medium rivers tend to be less shaded and rely on energy inputs from primary production; their macroinvertebrate communities tend to be dominated by grazers. Large rivers tend to rely on energy inputs from upstream and their macroinvertebrate communities tend to be dominated by collectors. Fish, mussel, and aquatic plant communities all vary as well. Rivers do vary from this general model (the river continuum concept), however it provides insight into how size is an important factor in determining and defining river communities. Water temperature is also important because species have optimum temperature preferences. Gradient provides a measure of channel morphology which correlates to valley shape, sinuosity, water velocity, and substrate size. All three factors are important in determining species compositions in rivers. Four size classes were defined using drainage areas of VSECs, following the Wildlife Action Plan (Eagle et al. 2005): headwaters and small tributaries are less than 40 mi2, medium rivers are between 40 and 179 mi2, large rivers are between 180 and 620 mi2, and very large rivers are greater than 620 mi2. Four classes of temperature were defined for each VSEC, generally defined as: cold (<19°C), 63 cool (19-21°C), and warm (>21°C). Three classes of gradient were defined: low (an average gradient less than 0.001), moderate (between 0.001 and 0.006), and high (greater than 0.006). Gradient classes were defined using the 25th and 75th percentiles of all stream reach gradients. See figures 23 and 23 for map of classification framework used. Limitations: Classification requires discrete boundaries however riverine ecosystems are essentially a continuum. As a result, river classification is inherently difficult. The main limitation to using VSECs is that the current VSEC framework is still under construction. We used the most current version (August, 2007), yet the MDNR Fisheries Division is continuing to refine and evaluate the framework. They are working on finalizing version 3. Since there has been significant work already towards evaluation of VSECs, we decided to proceed with our analysis using this version, which had change quite a bit from version 1. We do not expect major changes in the boundaries of the current VSECs, the reach identifier (pugap_code) is provided in the analysis. One limitation with our classification is that the gradient classes are not necessarily ecologically based. However, we were unable to find literature backing specific gradient breaks. To build a stronger classification, future research is needed to determine or document gradient classes that are ecologically meaningful. File name: vsec_size_temp.shp, vsec_gradient.shp Data source: Institute for Fisheries Research, Michigan Department of Natural Resources, version as of August 2007. groundwater_vsec_statewide_6_29_07.shp Lake classification Description: Lake ecosystems were classified using Higgins et al. (1998), which was based on available GIS data. Most of the data used in this classification were queried from or calculated using queried information from available data layers. Lakes were classified based on size, connectivity, shoreline complexity, and proximate geology. These particular variables were used based on available data, literature, and expert review. Size provides a measure of the availability and types of habitat in a lake (Eagle et al. 2005). Most small lakes are shallow, unstratified, have relatively high nutrient concentrations, and are somewhat likely to have low oxygen levels in winter. Additionally, they can either be turbid due to wind resuspension with no rooted plants or dominated by rooted plants with clear water. Succession is also a factor with these ecosystems because over time they fill in with sediments and become marsh. Small lakes can range from not stratified to fully stratified throughout the summer, and low winter oxygen levels can lead to winter kills. In lakes that stratify, a true pelagic or open-water zone develops and is distinct from the shallow littoral (or nearshore) zone. In medium lakes stratification and winter oxygen levels are also variable. They tend to have more complexity in their shoreline (lakes with many bays) and basin (lakes with more than one deep hole). Large lakes tend to be more homogenous in their chemical and biological makeup, but more diverse in their habitats than smaller lakes. They also are dominated by the pelagic zone. Connectivity refers to whether or not there are stream connections to the lake. Streams can influence a lake through the input or removal of water and nutrients as well as an exchange of species. Shoreline complexity becomes more important as lake size increases, increasing habitat variation. We used proximate geology as a surrogate for lake hydrology. All of these factors can influence species composition and communities. Typically ponds only have one community of fish, however as size increases, the pelagic habitat becomes more abundant and a pelagic fish community will be also present. 64 Figure 22. Map of size and temperature river classification framework used in analysis. 65 Figure 23. Map of gradient classification framework used in analysis. 66 We modified the size classes that Higgins et al. (1998) used as follows: ponds are >2 and <= 10 acres, small lakes are >10 and <100 acres, medium lakes are >= 100 and < 1000 acres, and large lakes are >1000. These size classes generally follow the Wildlife Action Plan (ponds <5 acres, small lakes 5-99 acres, medium lakes 100-999 acres, and large >1000 acres), however we increased the size range of ponds because water bodies less than 10 acres are often treated differently than larger lakes. For example, they are not typically surveyed or monitored. See Figures 24 and 25 for map of classification framework used. It should be noted that the Institute of Fisheries Research (MNDR) and Michigan State University are currently working on a lake classification for Michigan. For this effort we used Higgins et al. (1998) because it was both available and statewide in coverage. As more detailed and accurate classifications for Michigan become available, they should be evaluated for use in a statewide biodiversity assessment. Limitations: This classification is based on coarse scale data. To date there has been no ground-truthing and little analysis to determine accuracy and precision of assigned lake types in this classification. There are also many “single occurrence” lake types in this classification that may not be ecologically meaningful but artifacts of the classification process, which needs to be recognized in the unique lakes analysis. Although there are some critical issues with using this classification, it is currently the only lake classification for Michigan that is statewide and available in GIS format. Lake ecosystems undergo succession and begin to fill in with sediment; this process is important to keep in mind when setting conservation priorities, especially for ponds. MNFI typically distinguishes ponds from marshes if they have an open water area. Those “ponds” that have macrophytes across the entire water surface were identified as marsh for our work. Sampling for ponds can be difficult because they can be difficult to find, and during dry years could be designated as a marsh. We hope that by representing a variety of different types (Abell et al. 2002) of ponds that we will account for this process at least partially. File name: milakes_conn_shoreline.shp, milakes_proxgeol.shp Data source: The Nature Conservancy – Great Lakes Program, Higgins et al. 1998: milakes_w_attributes.shp Great Lakes Classification of areas within the Great Lakes is still largely in its infancy. The MNFI database contains point and polygon data for rare species found within the Great Lakes, however this data may or may not show important or critical areas for these species. Because most Great Lakes species can have large scale movements, single date location data does not provide adequate information when determining important areas for management and conservation. In addition, there have been other efforts focused on modeling important habitats for fish that we will currently defer to (Koonce et al. 1999). Due to lake of good information, habitats within the Great Lakes will not be considered in this analysis. Great Lakes nearshore areas are addressed in the terrestrial portion of this assessment. However, in the future this analysis should be revisited with both terrestrial and aquatic functions and processes in mind. The current analysis may miss out on important processes and functions of nearshore areas for fully aquatic species since this analysis was mainly based on coastal wetlands and did not include 67 Figure 24. Map of connectivity and shoreline complexity for lake classification framework used in analysis. 68 Figure 25. Map of proximate geology lake classification framework used in analysis. 69 other types of shoreline / nearshore types. The Institute of Fisheries Research and the University of Michigan are currently working on Great Lakes classification in Michigan. The US Geological Survey is also undertaking efforts to classify habitats in the Great Lake region through their Aquatic GAP program. As these efforts become available they should be examined for their use in expanding the statewide biodiversity assessment. Coarse-Filter: Aquatic Ecosystem Analysis Unique River Ecosystems statewide and by EDU Description: River ecosystems or VSECs were classified as unique using a 5% and 1% rule at two scales: EDU and statewide. See previous section on defining uniqueness for more detail. Use: By highlighting unique VSECs, we hope to capture potentially unique and important ecosystems that contribute to the diversity regionally and statewide. These layers will provide a relatively simple representation of where unique ecosystems are located within an EDU and statewide, and will help direct future survey efforts to determine true rarity, importance, and condition of these ecosystems. Limitations: Unique VSECs identified may be an artifact of the classification process and the accuracy of available digital data. As a result, true rarity is uncertain. But it does provide a basis that will help direct future survey efforts and analysis. In addition, we do not include a landscape context analysis with this layer because we are looking for rarity and not necessarily the best of the unique ecosystems. See river classification section for limitations associated with data used in this analysis. File names: vsec_unique_statewide_5pct.shp, vsec_unique_statewide_1pct.shp vsec_unique_edu_5pct.shp, vsec_unique_edu_1pct.shp Data source: Institute for Fisheries Research, Michigan Department of Natural Resources, version as of August 2007: groundwater_vsec_statewide_6_29_07.shp Results - statewide: There were 29,037 river reaches used in our analysis, which were aggregated into VSECs. Seventysix VSECs were removed from the analysis because they were not fully classified, leaving a total population of 9,961 VSECs statewide. These VSECs were categorized into one of 45 river types (there were a possible total of 48). Overall, river types were well represented statewide (Table 15). The number of VSECs within a type for headwaters and small tributaries ranged from 97 to 1,793, medium rivers ranged from 3 to 155, large rivers ranged from 1 to 78, and very large rivers ranged from 2 to 92. No headwaters and small tributaries were designated as unique statewide; large rivers and very large rivers dominated unique river ecosystems statewide (Table 15). This may be an artifact of the classification framework we used. Using the 5% rule, a total of 498 VSECs were targeted to be designated as unique (9,961*.05=498). Due to the number of VSECs within a type, a total of 524 VSECs were selected as unique statewide (Figure 26). The types of rivers selected as unique were very large rivers (except warm, low gradient types), all large rivers, all high gradient medium rivers, and cold low gradient medium rivers. The number of VSECs designated as unique increases from the southeast part of the state to the 70 northwest part of the state. The Southeast Michigan Interlobate and Lake Plain (16+2) EDU had the fewest VSECs selected as unique statewide with 26, whereas the Central Upper Peninsula (8) EDU had the most VSECS designated with 105 (Table 16). Using the 1% rule, a total of 109 VSECs were selected as unique statewide (Table 15, 16). Again, very large rivers were selected as well as mainly high gradient large rivers, and high gradient medium rivers (Figure 27). The Southeast Lake Michigan (3) EDU had the fewest designated as unique statewide with 2 VSECs and the Northern Lake Michigan, Lake Huron, and Straits of Mackinac (5) EDU has the most with 46 VSECs when using the 1% rule (Table 17). Results – in EDU: The number of VSECs in an EDU ranged from 722 to 2,049 and the number of river types ranged from 22 to 40 (Table 18); 48 was the maximum potential. The minimum number of VSECs in river types for all EDU’s was one and the maximum number ranged from 225 to 760. Although no headwaters and small tributaries were designated as unique statewide, they were designated as unique within EDUs. Using the 5% rule, a total of 36 to102 VSECs were targeted as unique dependent upon EDU (Figure 28). In the end, a total of 566 VSECs were selected as unique across EDUs (Table 19). The number of river types and VSECs designated as unique ranged from 11 to 22 and 37 to 124, respectively. The Northern Lake Michigan, Lake Huron, and Straits Of Mackinac (5) EDU had the most VSECs selected, where as the Eastern Upper Peninsula (7) EDU had the least. All river sizes were represented in the selected unique ecosystems. Using the 1% a total of 129 VSECs were selected as unique in EDUs (Table 20, 21, Figure 29). The number of river types and VSECs selected as unique across EDUs ranged from 6 to 10 and 12 to 26, respectively. In this analysis, the Southeast Lake Michigan (3) EDU had the most VSECs selected and the Eastern Upper Peninsula (7) EDU and the Western Upper Peninsula and Keweenaw Peninsula (6+12) EDU had the fewest. Not all river sizes were represented across EDUs. Table 15. Summary of classification of river valley segments (VSECs) and statewide uniqueness analysis. Number of VSECs Number of river types Minimum number of VSECs in a river type Maximum number of VSECs in a river type Number of river types in only one watershed Headwaters / Small Tributaries 8513 12 97 155 0 Medium Rivers 904 12 3 155 0 Large Rivers 346 12 1 78 1 Very Large Rivers 198 9 2 92 0 51 0 0 0 0 40 72 5 29 3 27 346 12 35 6 22 106 8 45 6 Maximum number of watersheds a river type occurred Number of unique VSECs (5%) Number of unique river types (5%) Number of unique VSECs (1%) Number of unique river types (1%) Table 16. Number of statewide unique VSECs in each EDU using the 5% and 1% rule. Number of unique VSECs statewide (5%) Number of unique VSECs statewide (1%) 2+16 26 3 3 80 2 71 4 81 5 5 145 46 7 31 8 8 105 29 6+12 56 16 Table 17. Names of rivers within EDUs that have unique VSECs using the 1% rule statewide. EDU Southeast Michigan Interlobate and Lake Plain (16+2) Southeast Lake Michigan (3) Saginaw Bay (4) Rivers with unique VSECs Huron, Saline, and unnamed Coldwater and Portage unnamed, Hemmingway and Whittle Drain, North Branch of the Flint River, Pine River, Sugar River, and Tittabawassee River Ausable River (mainstem and north branch), Baker Creek, Black River, Boardman Northern Lake Michigan, Lake Huron, River, Crumley Creek, Flinton Creek, Hudson Creek, Little Manistee River, Manistee and Straits Of Mackinac River, Manton Creek, Muskegon River, Pere Marquette River, Pine River, South Branch of the White River, Sturgeon River, Thunder Bay River, and West Branch of (5) Big Creek. Eastern Upper Tahquamenon River and Two Hearted River Peninsula (7) Central Upper unnamed, Daults Creek, Dead River, Huron River, Menominnee River, Michigamee Peninsula (8) River, Silver Creek, Silver River, Six-mile Creek, Sturgeon River, West Branch Huron River, West Branch Sturgeon River, Yellow Dog Creek Western Upper Black River, Ontonagon River (main, east, middle, and west branches), Jackson Creek, Peninsula and Montreal River, Pelton River, Portage River, Presque Isle River, Slate River, Sparkling Keweenaw Peninsula Creek, and Sturgeon River (6+12) Table 18. Summary of general river and VSEC statistics within EDUs. Number of river miles Total number of VSECs Number of actual river types Minimum number of VSECs in river types Maximum number of VSECs in river types Number of headwater / small tributary VSECs Number of medium VSECs Number of large VSECs Number of very large VSECs Ecological Drainage Unit 4 5 7 13,091 7,416 3,034 1,913 1,888 722 31 40 32 2+16 4,648 1,024 22 3 9,127 2,043 29 8 4,559 1,414 37 6+12 3,463 957 36 1 1 1 1 1 1 1 283 580 760 623 225 383 306 898 94 18 14 1,768 170 66 39 1,648 164 69 32 1,546 192 87 63 623 71 17 11 1,191 143 50 30 839 70 39 9 Table 19. Summary of unique river ecosystems by EDUs based on the 5% rule. Number of river types Number of VSECs Number of headwaters/ small VSECs Number of medium VSECs Number of large VSECs Number of very large VSECS Common Number of river types Number of VSECs Unique 2+16 11 57 3 13 119 4 16 101 5 22 124 7 16 37 8 18 74 6+12 20 54 11 14 18 14 12 967 20 46 40 13 18 1924 11 37 42 11 17 1812 10 16 56 42 19 1764 6 9 11 11 16 685 9 22 22 21 21 1314 5 21 19 9 19 903 72 Table 20. Summary of unique river ecosystems by EDUs based on the 1% rule. Unique Common Number of river types Number of VSECs Number of headwaters/ small tributary VSECs Number of medium VSECs Number of large VSECs Number of very large VSECS Number of river types Number of VSECs 2+16 6 16 3 7 26 4 10 25 5 9 20 7 9 12 8 8 18 6+12 9 12 3 7 0 6 17 1,008 0 7 19 0 24 2,017 3 8 13 1 23 1,888 0 5 10 5 32 1,868 3 0 4 5 23 710 0 5 4 9 31 1,370 2 4 4 2 30 945 Table 21. Names of additional rivers within EDUs that have unique VSECs using the 1% rule in each EDU. EDU Southeast Michigan Interlobate and Lake Plain (16+2) Southeast Lake Michigan (3) Saginaw Bay (4) Northern Lake Michigan, Lake Huron, and Straits Of Mackinac (5) Eastern Upper Peninsula (7) Central Upper Peninsula (8) Western Upper Peninsula and Keweenaw Peninsula (6+12) Rivers with unique VSECs Clinton River, and the St. Joseph River (main stem, east fork west branch, west branch) 15 new rivers Au Gres River, Cedar River, North Branch Chippewa River, Gamble Creek, Silver Creek, and West Branch Rifle River Carp Lake River Manistique River and Munuscony Rivers Black Creek, Escanaba River, Otter River, Walton River, and the West Branch of the Cedar River Flintsteel River, Little Gratiot River, Salmon Trout River, Tenmile Creek, and Tobacco River High-Quality Common River Ecosystems within EDUs Description: River ecosystems or VSECs were classified as common in an EDU using a greater than 5% rule; see previous section on defining uniqueness for more detail. Quality of common VSECs were assessed using Wang et al.’s (2006) analysis of landscape-level GIS data (Table 22). Quality will be relative within each EDU. Use: This analysis provides a relatively simple representation of where potential high-quality river ecosystems are located in each EDU and will help direct survey efforts to determine true condition and importance. Limitations: One main limitation of this data layer is that it does not include representation of all common river ecosystems. In addition, no field survey data was used to determine true condition and integrity of the ecosystems, so the individual VSECs highlighted may not be the best representatives available. Local factors that are not captured in this analysis could drive the quality of ecosystems. However, it does provide a basis to start from that will help direct future survey efforts. See river classification section for limitations associated with data used in this analysis. File name: vsec_HQ_edu.shp 73 Figure 26. Unique river ecosystems in Michigan using the 5% rule. 74 Figure 27. Unique river ecosystems in Michigan using the 1% rule. 75 Figure 28. Unique river ecosystems in Michigan by EDU for the 5% rule. 76 Figure 29. Unique river ecosystems in Michigan by EDU for the 1% rule. 77 Data source: Institute for Fisheries Research, Michigan Department of Natural Resources, version as of August 2007: groundwater_vsec_statewide_6_29_07.shp Institute for Fisheries Research, Michigan Department of Natural Resources: mi_epastar_nhd_stresref.shp Results: Of the 9,935 VSECs, 9,369 were classified as common in an EDU using the greater than 5% rule. Using the disturbance classification created by Wang et al. (2006), we selected the highest quality of the common river ecosystems (Figure 30). However, the Wang et al. (2006) analysis was conducted at the reach level. VSECs are made up of multiple reaches, and consequently, VSECs were not consistently classified in their disturbance classification. Reaches within a single VSEC could have different associated quality. For example, if a VSEC was made up of 4 reaches, each reach could have a different disturbance class (e.g. reference, no impact, degraded, reference). Therefore, only those reaches classified as reference within common VSECs were selected in our analysis (Table 23). Future work should review the entire VSEC and identify those common VSECs with an overall high quality. The most common type of headwater and small tributary streams were cool or warm with moderate gradient. The most common type of medium rivers was warm moderate gradient. The most common type of large river and very large river types were warm, low gradient. Table 22. Landscape variables used to determine quality (from Wang et al. 2006). Network watershed encompasses all areas upstream from the stream reach. Variables for all streams Active mining (#/10000 km2) Network watershed agricultural land use (%) Network watershed urban land use (%) MDEQ’s permitted point source facilities (#/100 km 2) MDEQ’s permitted point source facilities having direct connection with stream (#/100 km2) USEPA’s toxic release inventory sites (#/10000 km2) Population density (#/km2) Road crossing (#/km2) Road density (km/km2) Total nitrogen plus (phosphorus*10) loading (kg/l/yr) Watershed area treated with manure from barn yards (m/km) Additional variables for coldwater streams Total nitrogen plus (phosphorus*10) yield (kg/l/year) Additional variables for warmwater streams Dam density (#/100 km2) USEPA’s toxic release inventory sites discharging into surface water (#/10000 km2) Table 23. Summary of the number of river reaches classified as common ecosystems. River size headwaters/small tributaries medium rivers large rivers very large rivers Count 26,100 4,361 686 511 78 Figure 30. High quality river ecosystems in Michigan by EDU. 79 Rivers with Unimpeded Access To The Great Lakes Description: This shapefile shows river stretches still accessible to the Great Lakes. These data were obtained from Institute of Fisheries Research, Michigan Department of Natural Resources. Use: This layer identifies rivers that may have important habitats for migrating fish species, such as suckers, redhorse, salmon, and sturgeon, and ecosystem function in terms of connectivity. Limitations: This layer provides limited information since it is not coupled with migrating or exotic species data. File name: mi_epastar_nhd_damseg.shp Data source: mi_epastar_nhd_damseg.shp. Produced and supplied by Great Lakes GIS project of the University of Michigan and Michigan Department of Natural Resources, Institute for Fisheries Research, Ann Arbor, Michigan, USA, (May, 2007). Results: Many of the rivers highlighted in Figure 31 in the Saginaw Bay (4) EDU area are ditched streams that may be seasonal and may not be important to migrating fish species. Future work should compare accessible rivers with known species data to help determine priority areas for migrating fish. Intact Headwaters in Michigan Description: A land cover analysis was conducted to identify intactness of headwater (stream order 1) watersheds. Headwater watersheds with 100% natural cover were identified. Additionally, percent naturalness for all headwater watersheds was provided. Use: Headwaters are critical ecosystems that can serve as refuge areas, sources of organic material, and stream cooling. They are important areas for fish, macroinvertebrates, amphibians, and reptiles. These ecosystems are also very sensitive to disturbance and any negative impacts to them can cause negative impacts downstream. Limitations: Land coverage data is limited in accuracy and is static. IFMAP land coverage is limited in accuracy. In addition, the IFMAP land cover was documented from satellite imagery taken between 1999 and 2001. Some areas of land have been altered since that time period rendering the land cover outdated for those areas. File name: headwaters100Natural.shp headwatersPctnatural.shp 80 Figure 31. Rivers in Michigan with unimpeded access to the Great Lakes. 81 Data sources: Michigan Department of Natural Resources. 2003. Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Lower Peninsula and Upper Peninsula) GIS data layer, version 1 (2003). lu2000_f. reach_watersheds.shp. Date unknown. Produced and supplied by Great Lakes GIS project of the University of Michigan and Michigan Department of Natural Resources, Institute for Fisheries Research, Ann Arbor, Michigan, USA, (May, 2007). mi_nhd_gap.shp. Date unknown. Produced and supplied by the University of Michigan and Michigan Department of Natural Resources, Institute for Fisheries Research, Ann Arbor, Michigan, USA, (September, 2005). Results: There are 43,288 miles of river in Michigan according to the data layer we used, and more than half of them, 25,227 miles, are first order streams. There are 19,426 first order reach watersheds out of 35,858 reach watersheds in Michigan. Headwater (first order) watersheds account for 22,802,925 acres in Michigan. There are 1,116 headwater watersheds with 100% natural land cover and they make up about 670,274 acres in Michigan. Most of the 100% natural headwater watersheds occur in the Upper Peninsula, however there are also some located in the Lower Peninsula (Figure 32). The majority of the natural headwaters were found in the Central Upper Peninsula (7) EDU, and the fewest were found in the Southeast Michigan Interlobate and Lake Plain (16+2) EDU (Table 24). By decreasing the threshold to 87% naturalness, more headwater watersheds in the southern Lower Peninsula were included (Figure 33). Table 24. Number of 100% natural headwater watersheds in each EDU. EDU 3 4 5 6+12 7 8 16+2 Count 9 32 93 322 201 457 2 Unique Lake Ecosystems in by statewide and by EDU Description: Lake ecosystems were classified as unique using a 5% and 1% rule; see previous section on defining uniqueness for more detail. Use: By highlighting unique lakes, we hope to capture potentially unique and important ecosystems that contribute to the diversity of Michigan and the Great Lakes Region. This analysis will provide a relatively simple representation of where in Michigan unique ecosystems are located and will help direct future survey efforts to determine true rarity, importance, and condition of these ecosystems. 82 Figure 32. Intact watersheds of headwater streams in Michigan. 83 Figure 33. Percent natural land cover in watersheds of headwater streams in Michigan. 84 Limitations: Unique lake types identified may be an artifact of the classification process and the accuracy of available digital data. Although true rarity is uncertain, this analysis provides a basis that will help direct future survey efforts. In addition, a landscape context analysis is not included with this layer because we are looking for rarity and not necessarily the best of the unique. See lake classification section for limitations associated with data used in this analysis. File name: lake_unique_statewide_5pct.shp, lake_unique_statewide_1pct.shp lake_unique_edu_5pct.shp, lake_unique_edu_1pct Data source: The Nature Conservancy – Great Lakes Program, Higgins et al. 1998 milakes_w_attributes.shp Results - statewide: There were 10,772 lakes used in our analysis. Originally, the dataset we used had a universe of 11,172 lakes but 372 were removed due to small size (<=2 acres), lack of proximate geology value, or lack of EDU assignment. The current EDU layer does not cover most islands and the boundary lines are at a coarser scale than the state boundary. Statewide there are 157 lake types. Twenty-three lake types occurred in only one watershed. The number of lakes within a type for ponds ranged from 1 to 1,226, for small lakes from 1 to 1,128, for medium lakes from 1 to 116, and for large lakes from 1 to 18 (Table 25). There were 61 lake types with five or fewer lakes. Lakes were identified as unique within each of the four size classes. Table 25. Summary of classification of lakes and uniqueness analysis. Ponds 5,136 5,101 38 1 1,226 7 44 14 53 12 23 Total number of lakes* Number of lakes in analysis Number of lake types Minimum number of lakes in a lake type Maximum number of lakes in a lake type Number of lake types in only one watershed Maximum number of watersheds a lake type occurred Number of unique lake types (5%) Number of unique lakes (5%) Number of unique lake types (1%) Number of unique lakes (1%) Small Lakes 4,837 4,805 52 1 1,128 3 45 24 165 13 35 Medium Lakes 873 792 50 1 116 8 28 40 281 17 38 Large Lakes 86 74 17 1 18 5 12 17 74 14 32 Using the 5% rule, a total of 539 lakes were targeted as unique, the actual number selected was 573 assigned among 95 lake types (Figure 34). Lakes selected as unique were scattered across the state and no pattern was apparent. The Eastern Upper Peninsula (7) EDU had the fewest lakes identified, while the Northern Lake Michigan, Lake Huron, and Straits of Machinac (5) EDU had the most (Table 26) lakes identified. Table 26. Number of statewide unique lakes in each EDU using the 5% and 1% rule. Number of unique lakes statewide (5%) Number of unique lakes statewide (1%) 16+2 46 11 3 98 16 85 4 104 21 5 127 32 7 28 8 8 88 21 6+12 82 19 Using the 1% rule, a total of 108 lakes were selected as unique statewide (Table 25). The Eastern Upper Peninsula (7) EDU again had the fewest lakes selected, whereas the Northern Lake Michigan, Lake Huron, and Straits of Mackinac (5) EDU had the most (Figure 35) lakes identified. Results – by EDU: The number of lakes in an EDU ranged from 550 to 2,547 and the number of lake types ranged from 56 to 99 (Table 27). The minimum number of lakes in a lake type for all EDUs was one and the maximum ranged from 92 to 379. Table 27. Summary of general lake statistics within EDUs. Number of lakes Number of ponds Number of small lakes Number of medium lakes Number of large lakes Number of possible lake types Number of actual lakes types Minimum number of lakes in a type Maximum number of lakes in a type 16+2 1,123 522 511 89 1 176 79 1 169 3 2,547 1,238 1,126 177 6 176 88 1 367 4 1,446 769 589 82 6 208 99 1 92 5 2,304 970 1,089 207 38 192 94 1 379 7 1,362 605 647 100 10 160 78 1 198 8 1,413 710 594 100 9 208 85 1 135 6+12 550 287 222 37 4 176 56 1 134 Using the 5% rule, a total of 33 to 131 lakes were targeted as unique dependent upon EDU (Figure 36). A total of 577 lakes were selected as unique across EDUs (Table 28). The Western Upper Peninsula and Keweenaw Peninsula (6+12) EDU had the fewest lakes identified and the Southeast Lake Michigan (3) EDU had the most lakes identified. The Northern Lake Michigan, Lake Huron, and Straits of Mackinac (5) EDU had the highest number of unique lake types. Overall, unique lakes selected were typically spread out throughout an EDU and were distributed across size classes. In general, small lakes and medium lakes were represented more than ponds and large lakes. Table 28. Summary of unique lake ecosystems by EDU based on the 5% rule. Unique Common Number of lake types Number of lakes Number of ponds Number of small lakes Number of medium lakes Number of large lakes Number of lake types Number of lakes 16+2 38 65 12 21 31 1 41 1058 3 46 131 31 29 65 6 42 2416 86 4 44 74 8 22 38 6 55 1372 5 48 123 15 37 46 25 46 2181 7 37 74 15 29 20 10 41 1315 8 47 77 19 27 26 5 38 1336 6+12 28 33 6 11 12 4 28 517 Figure 34. Unique lake ecosystems in Michigan using the 5% rule. 87 Figure 35. Unique lake ecosystems in Michigan using the 1% rule. 88 Using the 1% rule, a total of 185 lakes were selected as unique across EDUs (Table29). The number of lakes and lake types designated as unique ranged from 17 to 41 and 17 to 26, respectively. The Eastern Upper Peninsula (7) EDU had the fewest lakes selected, whereas the Southeast Lake Michigan (3) EDU had the most (Figure 37). All lakes size classes were represented, except in the Southeast Lake Michigan (3) EDU where no large lakes were designated as unique. In general, ponds and large lakes were less represented than small and medium lakes. Table 29. Summary of unique lake ecosystems by EDU based on the 1% rule. Unique Common Number of lake classes Number of lakes Number of ponds Number of small lakes Number of medium lakes Number of large lakes Number of lake classes Number of lakes 16+2 19 19 2 7 9 1 60 1104 3 26 41 5 14 22 0 62 2506 4 23 23 2 8 9 4 76 1423 5 25 37 8 7 15 7 69 2267 7 17 17 3 6 5 3 61 1372 8 25 25 9 5 8 3 60 1388 6+12 23 23 4 7 8 4 33 527 High-Quality Common Lake Ecosystems within EDUs Description: Lake ecosystems were classified as common in an EDU using a greater than 5% rule; see previous section on defining uniqueness for more details. Quality of common lakes was assessed by calculating percent natural land use and road density in a 500 m buffer around each lake (Soranno et al. in prep). Values of the landscape variables were put into classes and lakes were ranked according to lowest road density and highest percent natural land use. Land use (Allen 2004) is known to affect the quality of aquatic ecosystems and species. Road density was included as part of the landscape context analysis because we felt true land use may be masked in the IFMAP data. Natural vegetation buffers often surround lakes, even if housing density is high. Quality was relative within each EDU. For this analysis we targeted 10 unconnected ponds or small lakes, 5 connected ponds or small lakes, 5 medium lakes, and 1 large or very large lake ecosystem in each EDU with the best landscape context. No threshold values for quality were used, just target numbers of lakes. The best qualilty lakes were seleceted until we got our target number. However, more lakes than the target number could be selected if many lakes had the same quality value. Use: This analysis provides a relatively simple representation of where potential high-quality lake ecosystems are located in each EDU, and helps direct survey efforts to determine true condition and importance. Limitations: One main limitation of this data layer is that it does not include representation of all common lake ecosystems. In addition, no field survey data was used to determine true condition and integrity of the ecosystems. Individual lakes highlighted may not be the best representatives available, because local factors that are not captured in this analysis could drive the quality of an ecosystems. However this analysis does provide a basis to help direct future survey efforts. See lake classification section for limitations associated with data used in this analysis. 89 Figure 36. Unique lake ecosystems in Michigan by EDU using the 5% rule. 90 Figure 37. Unique lake ecosystems in Michigan by EDU using the 1% rule. 91 File name: lake_HQ_edu.shp Data sources: The Nature Conservancy – Great Lakes Program, Higgins et al. 1998 milakes_w_attributes.shp Michigan Department of Natural Resources. 2003. Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Lower Peninsula and Upper Peninsula) GIS data layer, version 1 (2003). lu2000_f. Forest, Minerals and Fire Management Division, Michigan Department of Natural Resources (MDNR), Lansing, Michigan. Michigan Center for Geographic Information. 2006. Michigan Geographic Framework v6b. roads_only_6b.shp. Results: As to be expected, more high quality ponds were selected than larger lakes (Table 30). The Eastern Upper Peninsula (7) EDU had the most high quality lakes selected in each of the four size classes, and the Saginaw Bay (4) EDU had the fewest (Figure 38), likely due to the paucity of lakes in that EDU. Table 30. Summary of the number of high quality lakes by size class in each EDU. Ponds Small lakes Medium lakes Large lakes Total: 2 +16 26 20 6 0 52 3 55 40 7 1 103 4 32 8 4 1 45 5 52 13 43 0 108 7 147 91 66 4 308 8 60 26 50 0 136 6 + 12 29 24 21 1 75 Total: 401 222 197 7 827 Functional (or least modified) sub-watersheds Description: This analysis integrated land cover, fragmentation, and pollution analyses into a shapefile that highlights functional sub-watersheds (huc-12). Three different analyses (land cover, fragmentation, and pollution) were conducted and scored between 1 and 5 using quantiles, 1 being the least disturbed and 5 being the most disturbed. A single metric was pooled to determine the 2 least disturbed subwatersheds within each watershed and the least disturbed sub-watersheds statewide. Use: The quality of an aquatic ecosystem is largely dependent upon its landscape context, which include those areas upstream. To truly protect or manage a river or lake its contributing watershed must be taken into account. This analysis provides a method for assessing the quality of sub-watersheds based on available data. This information can be used to direct future surveys or target conservation efforts. Limitations: We call this analysis “functional sub-watersheds,” however true functionality is unknown. This layer is essentially our “best-guess” based on available data. Functionality and disturbance are complicated processes, and in this analysis we are only targeting a few potential indicators. 92 Figure 38. High quality lakes by EDU. 93 Land cover analysis Description: The land cover analysis was based on a combination of natural land cover for the entire catchment and within the riparian zones. All natural vegetation types identified by the IFMAP land coverage were combined together to form a new all natural vegetation data layer. Natural vegetation included grassland/herbaceous, shrubland, forest, and wetland. The percent of sub-watershed with natural land cover was determined and placed in one of 5 classes based on quartiles. Additionally, all rivers and lakes were buffered outward by 60 m to create the riparian zone for analysis. The percent of natural land cover within riparian zones was determined and placed in one of 5 classes based on quartiles. These two analyses were added together and divided by 2 to determine the overall class (1-5) for each sub-watershed. Use: This analysis was used to rank sub-watersheds in terms of natural cover, and was one component of the functional watershed analysis. Limitations: IFMAP land coverage is limited in accuracy. In addition, the IFMAP land cover was documented from satellite imagery taken between 1999 and 2001. Some areas of land have been altered since that time period rendering the land cover outdated for those areas. File names: pctNat_subwatershed.shp pctNat_Riparian_subwatershed.shp Data source: Michigan Department of Natural Resources. 2003. Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Lower Peninsula and Upper Peninsula) GIS data layer, version 1 (2003). lu2000_f. Forest, Minerals and Fire Management Division, Michigan Department of Natural Resources (MDNR), Lansing, Michigan. Results: The percent natural land cover in sub-watersheds showed the expected; sub-watersheds in the northern Lower Peninsula and Upper Peninsula have more natural land cover (Figure 40). However, when only riparian land cover was considered, more sub-watersheds in the southern Lower Peninsula have relatively good natural riparian buffers (Figure 41), and more sub-watersheds in the northern Lower Peninsula and Upper Peninsula have poor riparian cover relative to the overall sub-watershed land cover. Many of the sub-watersheds with poor riparian cover are located in more urban environments (exp. Alpena, Escanaba, Sault St. Marie). When both natural land cover in the entire sub-watershed and within the riparian buffer were combined, the map shows something in between the two analyses (Figure 42). 94 Figure 39. Percent natural land cover by sub-watershed. 95 Figure 40. Percent natural land cover in riparian areas by subwatershed. 96 Figure 41. Land cover analysis by subwatershed. 97 Fragmentation analysis Description: This analysis provides information on the level of fragmentation of the rivers in each subwatershed. There are two major fragmentation factors for rivers that can be easily gleaned from GIS data: dams and road crossings. Both can alter hydrologic flows, sediment exchange, and disrupt fish and mussel movements and population exchanges. Not all road and stream crossings fragment aquatic habitats, but if improperly installed and maintained they can. Because the quality of road crossing cannot be determined using available data, we treated all crossings as a level of fragmentation. In this analysis, the number of dams per river mile and the number of road and stream crossings per river mile in each sub-watershed were calculated, ranked within each watershed, and placed in one of 5 classes based on quartiles. These two analyses were added together and then divided by 2 to determine the overall class (1-5) for each sub-watershed. Use: This analysis provides information on which sub-watersheds have the least fragmentation, and was used to calculate the overall functional score of each sub-watershed. Limitations: Sub-watersheds with <0.1 miles of river were eliminated from the analysis. The data used in this analysis are static and hence may be outdated for some areas. Additionally, these are not the only factors that create fragmentation in aquatic ecosystems, however they are the easiest to determine given the available data. Even if sub-watersheds have no dams and few road crossings, they still can be substantially impacted by fragmentation upstream or downstream from the sub-watershed boundaries. File names: damCount_subwatershed.shp rdxStrCount_subwatershed.shp fragementation_subwatershed.shp Data sources: dams.shp from MDEQ Michigan Center for Geographic Information. 2006. Michigan Geographic Framework v6b. roads_only_6b.shp. Institute for Fisheries Research, Michigan Department of Natural Resources. mi_nhd_gap.shp. Date unknown. Produced and supplied by the University of Michigan and Michigan Department of Natural Resources, Institute for Fisheries Research, Ann Arbor, Michigan, USA, (September, 2005). Results: Some sub-watersheds, according to the available data, have very small sections of river, which can create high numbers of dams and stream crossings per river mile. For example, the sub-watershed with the highest number of dams (9) per river mile. This sub-watershed actually had 5 dams in 0.54 miles of river. The sub-watershed with the highest number of road crossings (22), resulted from a sub-watershed with 0.18 miles of river and 4 stream crossings. 98 Road crossings are a larger fragmentation issue in the southern Lower Peninsula, whereas dams are a bigger issue in the northern Lower Peninsula and the Upper Peninsula (Figure 42, 43). Figure 44 suggests that fragmentation is a major issue for aquatic ecosystems in Michigan across the state; there are few areas where fragmentation is not an issue. Pollution analysis Description: This analysis provides an overall pollution score to each sub-watershed. This metric includes a variety of variables to target both point and non-point source pollution. The number of DEQ permitted point source facilities and active mining operations was calculated. In addition, the percent impervious surface for each sub-watershed was calculated. As in the previous analyses, each was placed in one of 5 classes (1-5) within a sub-watershed based on quartiles. The overall pollution metric was calculated by adding together each individual rank and divided by 3. This resultant metric ranged from 1 to 5, with 1 being the least polluted and 5 being the most polluted. Use: This analysis provides a broad look at both point and non-point source pollution within subwatersheds. Those sub-watersheds with the least pollution threats were identified. The overall metric was used to calculate the overall functional score for each sub-watershed. Limitations: The point source and toxic release site data used in this analysis is static and may be outdated for some areas. IFMAP land coverage is limited in accuracy. In addition, the IFMAP land cover was documented from satellite imagery taken between 1999 and 2001. Some areas of land have been altered since that time period rendering the land cover outdated for those areas. File names: imperv_subwatershed.shp npdesCount_subwatershed.shp mineCount_subwatershed.shp pollution_subwatershed.shp Data sources: U.S. Geological Survey (USGS) Active Mines and Mineral Processing Plants in the United States in 2003. http://mrdata.usgs.gov. mineplant_mi_georef.shp. Published 2005. USGS, Reston, VA. Michigan Department of Environmental Quality (DEQ), Non-point source data, npdes_gw_permits1_georef.shp. Michigan Department of Natural Resources. 2003. Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Lower Peninsula and Upper Peninsula) GIS data layer, version 1 (2003). G:\gis\landu\lu2000_f. Forest, Minerals and Fire Management Division, Michigan Department of Natural Resources (MDNR), Lansing, Michigan. 99 Figure 42. Number of dams per river mile in sub-watersheds. 100 Figure 43. Number of road crossings per river mile in sub-watersheds. 101 Figure 44. Fragmentation analysis by sub-watersheds. 102 Figure 45. Number of DEQ non-point source pollution permits per river mile in sub-watersheds. 103 Figure 46. Percent impervious surface in sub-watersheds. 104 Figure 47. Number of active mines per river mile in sub-watersheds. 105 Figure 48. Pollution analysis by sub-watersheds. 106 Results: The non-point source pollution (Figure 45) indicator shows a similar pattern as the impervious surface pollution indicator (Figure 46); they closely follow locations of cities or major towns. Active mines (Figure 47) are more limited in area as a pollution threat. The overall pollution analysis (Figure 48) shows that most of the Lower Peninsula of Michigan has moderate to high pollution threats. Many studies have shown that watersheds with as little as 10 to 20% impervious surfaces are heavily degraded (Paul and Meyer 2001). Yet, for much of Michigan impervious surfaces range between 6 and 10% suggesting that most of Michigan’s streams still have the potential for healthy natural processes to exist. Overall Functional Sub-Watershed Results: File name: functional_subwatersheds.shp As expected, the Upper Peninsula had the most sub-watersheds that were classified as least modified, and the southern Lower Peninsula had the most sub-watersheds classified as most modified (Figure 49). The sub-watersheds with the lowest score within each EDU occurred along the coast or at the border between Michigan and Indiana. This is likely an artifact of the small size of these subwatersheds and lack of rivers in these areas. Overall, the Lower Peninsula had very few sub-watersheds that scored a 1 (least-modified). In the Upper Peninsula between 13 and 26% of sub-watersheds scored a 1. When the top two leastmodified scores (1 and 2) are combined, greater than 70% of sub-watersheds are in good functioning condition (Table 31). The Northern Lake Michigan, Lake Huron, and Straits of Mackinac (5) EDU had the next group of most functional sub-watersheds. The Saginaw Bay (4) and Southeast Lake Michigan (3) EDUs were ranked fairly similarly with most sub-watersheds having moderate amounts of modification. The Southeast Michigan Interlobate and Lake Plain (16+2) EDU, as expected, contained the most-modified (or least functional) sub-watersheds in the State. There were 145 sub-watersheds that were classified as highly functional with a score of 1. In the Northern Lake Michigan, Lake Huron, and Straits of Mackinac (5) EDU these sub-watersheds occurred in Black, Cheboygan, Lone Lake – Ocqueoc, Manistee, and Pere Marquete – White watersheds. In the Eastern Upper Peninsula (7) EDU they occurred in the Betsy – Chocolay, Brevoort – Millecoquin, Carp – Pine, Fishdam – Sturgeon, Manistique, St. Marys, Tahquamenon, and Waiska watersheds. In the Central Upper Peninsula (8) EDU they occurred in the Brule, Cedar – Ford, Dead – Kelsey, Escanaba, Menominee, Ontonagon, and Sturgeon watersheds. And in the Western Upper Peninsula and Keweenaw Peninsula (6+12) EDU they occurred in the Black – Presque Isle, Dead – Kelsey, Keweenaw Peninsula, and Ontonagon watersheds. The most functional sub-watersheds, with a score of 2, for the other EDUs are following: In the Saginaw Bay (4) EDU occurred in the Au Gres – Riffle, Flint, Kawkawlin – Pine, Pine, Shiawassee, Tittabawassee, and Upper Grand watersheds. In the Southeast Lake Michigan (3) EDU they occurred in the Black – Macatawa, Kalamazoo, Muskegon, Pine, Shiawassee, St. Joseph, and Upper Grand watersheds. And in the Southeast Michigan Interlobate and Lake Plain (16+2) EDU they occurred in the Flint, Huron, and St. Joseph watersheds. 107 Figure 49. Sub-watersheds in Michigan scored from least-modified to most-modified. 108 Table 31. Percent of sub-watersheds in each EDU in each score category of the functional analysis. Scores of 1 are the least modified sub-watersheds and scores of 5 are the most modified subwatersheds. Score 1 2 3 4 5 16+2 0 1 32 58 10 3 0 3 45 46 7 4 0 5 55 37 3 5 3 37 44 14 2 7 23 49 25 3 0 8 13 61 22 4 1 6+12 26 58 14 2 0 Fine-Filter: Element Occurrence Data Description: The Michigan Natural Features Inventory has been inventorying and tracking Michigan’s threatened, endangered, and special concern species and high quality natural communities since 1979. As of September, 2006, MNFI tracked 417 plant species, 248 animal species, and 74 natural community types. In addition to species and natural communities, MNFI also tracks other natural features such as colonial bird nesting colonies and significant geological features. The tracked species include those with Federal and State legal protection and special concern species which have no legal protection. Like the special concern species, natural communities also have no legal protection status. As of September, 2006, The MNFI database contained approximately 14,532 records of these natural features (plants, animals, and natural communities). Data sources include museum and herbarium collections, published reports, MNFI field surveys, and information from cooperators. Database records span a range from historic information to very current information from the latest field season. The data in the MNFI database are based on ground-truthed observations by reliable experts and are continually updated. The MNFI database is the most complete record of Michigan’s sensitive species and natural features. The MNFI database is a Natural Heritage database and utilizes Natural Heritage methodology and data standards originally designed by The Nature Conservancy and now maintained by Natureserve (www.natureserve.org). The MNFI database is more than a presence/absence database. Among other information, it contains dates of sightings, global and state imperilment rankings for species, and a quality (or viability) ranking for individual occurrences. Definitions of the global and state (or subnational) rankings can be found in appendix A. The quality ranking is an A – D scale with A being the highest quality. Other codes such as E for extant, H for historic, and X for extirpated are also used. The standards for applying a quality rank to an occurrence vary by species and community, but generally fall into three main categories: size, condition, and context. Approximately 50% of the mussels tracked by MNFI are considered globally critically imperiled (G1), imperiled (G2), or vulnerable (G3). This represents approximately 20% of all native mussels found in Michigan. In addition, 40% of the reptiles and 32% of the insects tracked in the MNFI database have a global rank of G1 – G3 some of which rely on aquatic ecosystems. For a list of aquatic species used in these analyses see Appendix D. Limitations: The primary limitations to MNFI’s element occurrence database are 1) it contains static information – each element occurrence is updated infrequently, 2) lack of a statewide systematic survey, and 3) in some cases, very old and/or general (non location specific) records. Biological information from the 109 field is collected annually from MNFI staff and other reliable contributors. Once this information is entered into the database, it may be decades before it gets updated. For example, approximately 36 % of the records in the database are over 20 years old. More significantly, there has never been a systematic survey of element occurrences in the state. This means that something can be said about the biological significance of an area containing element occurrence records, however nothing can be said definitively about the biological significance of areas with no known element occurrence records. This is where the quote “absence of evidence is not evidence of absence” comes into play. Related to this, is that there have been small areas of the state that have been systematically surveyed; however they are predominantly owned by public agencies or non-governmental organizations such as The Nature Conservancy. Fine-Filter: Element Occurrence Data Analysis EO Frequency Count Description: The EO frequency count is a count of all element occurrences that fall within a given public land survey system (PLSS) section. The model utilizes a statewide GIS data layer (Environmental Systems Research Institution (ESRI) shapefile) of the PLSS sections. A numeric count field is added to the section shapefile theme table. Each section shape is selected in turn and intersected with the MNFI GIS database. The number of aquatic occurrences intersecting each section shape is counted and that value is calculated into the count field in the section shapefile theme table. A cutoff date of September 1, 2006 was used to create the EO frequency datasets. All records added to the Michigan Natural Features database after this date are not included in this analysis. This analysis is based on terrestrial boundaries (1 mile blocks) to allow for this analysis to be easily combined or overlaid with the terrestrial analysis. Use: The EO frequency count is a relatively simple representation of the MNFI data. It is designed to show users where there are concentrations of known species or natural community occurrences in the MNFI database. While the EO frequency count provides limited information, it does fulfill its intended purpose. Users can see if there are known occurrences in the vicinity of a proposed project or delineate those areas where there are concentrations of occurrences. All species information is removed so locations of particularly sensitive species cannot be determined from the model. Limitations: The primary disadvantage is that it provides very limited information. The user only knows that the known boundary of an occurrence overlaps the boundary of the area of interest. No allowance is made for the age of the record, relative importance of the species, or the extent of potential habitat within the occurrence boundary. File name: Aq_EO_trs_0906.shp Data source: Biot_p – Biotics polygon database created directly from Biotics from version created September 1, 2006. 110 Results: As the map shows (Figure 50), there are limited concentrations of rare aquatic species across the state. In the Upper Peninsula and the northern Lower Peninsula the high frequency counts are driven by common loon occurrences (Table 32), whereas, in the southern portion of the Lower Peninsula the high concentrations are driven by fully aquatic species, fish and mussels (Figure 51). The areas with the highest frequency counts of aquatic rare species are found in the Lower Grand River watershed, the St. Joseph River of the Maumee, the River Raisin, and the Black River in the St. Clair river watershed. This is not surprising since the Lake Erie basin and portions of the Southeast Lake Michigan (3) EDU have the most diverse aquatic species assemblages due to species range distributions. Table 32. Frequency of element occurrences (with and without loons) and number of species occurring in EDUs. 3 4 5 6+12 7 8 2+16 Frequency (all aquatic sp) 384 273 438 157 172 181 524 Frequency (no Loons) 376 222 196 38 53 53 523 Species Count 30 26 25 13 10 16 35 EO Likelihood Description: The likelihood modeling process consists of grouping species into habitat guilds, creating a habitat layer for each guild, using the habitat layer to redefine the spatial extent of the corresponding occurrences, and intersecting the spatially redefined occurrences with political boundaries such as Public Land Survey System (PLSS) units. Each political unit is then assigned the “highest” likelihood value for all occurrences that fall within it’s boundary. Aquatic species’ habitat layers were created from either stream lines, the water class in the current land cover layer, or a combination of the two. The habitat layers are then used to redefine the spatial extent of the occurrences. The spatial extent of each occurrence is replaced by the spatial extent of the habitat within. After the overlay process, each occurrence still retains all database attribute values, including the date of the last observation. A value is assigned based on this field and is used to represent the likelihood that the occurrence still exists. Occurrences with a last observed date of no later than 1982 are assigned a value of one, occurrences between 1970 and 1982 are assigned a value of 0.5, and occurrences prior to 1972 are assigned a value of 0.25. These likelihood values are then aggregated up to a PLSS data set. First all records in the PLSS data set are selected and assigned a No Status value. Next the records in the occurrence layer with the lowest likelihood of still existing (value = 0.25) are selected. The PLSS data set is intersected with the occurrence layer and the selected PLSS records are assigned a value of “Low”. Next those records with a moderate likelihood of still existing are selected (value = 0.5). The PLSS data set is intersected with the occurrence layer and the selected PLSS records are assigned a value of “Moderate”. Finally 111 Figure 50. Frequency counts of aquatic element occurrences by PLSS. 112 Figure 51. Frequency counts of aquatic element occurrences without loon EOs by PLSS. 113 the occurrences with the highest likelihood of still existing (value = 1) are selected. The PLSS data set is intersected with the selected occurrence features and the selected PLSS records are assigned a value of “High”. Performing the selections and intersections in this order insures that a higher likelihood value in any PLSS feature will override a lower likelihood value. The element occurrence database for this model was accessed September 1, 2006. Any records added to the Michigan Natural Features database after this date are not included in this analysis. This analysis is aggregated to terrestrial boundaries (1 mile blocks) to allow for merging or overlay with the terrestrial analysis. This analysis is based on terrestrial boundaries (1 mile blocks) to allow for this analysis to be easily combined or overlaid with the terrestrial analysis. Use: The EO likelihood model is designed to help protect biodiversity and minimize potential regulatory problems by directing development away from those areas with a high likelihood of encountering a sensitive species. Because no specific species information is presented, the model reduces the sensitivity of the underlying MNFI data. A high probability indicates that the area of interest contains the spatial extent of an occurrence, there is potential habitat within the area, and the occurrence has been observed in the recent past. A low probability indicates that the area contains the spatial extent of an historic species occurrence and there is potential habitat within the area. While the low likelihood indicates that the underlying occurrences are historic, there is still a possibility that the species persists in appropriate habitat. In the recent past, MNFI botanists have reconfirmed three 100 year old plant records. A moderate likelihood indicates, by default, something between the other two values. The EO likelihood model provides users with a higher level of information than the simple EO frequency count. Unlike the EO frequency count, which only implies that the extent of an occurrence lies within an area of interest, the EO likelihood model delineates those areas where there is a higher likelihood of encountering a sensitive species or natural community. Also, by utilizing potential habitat within the known extent of the occurrences, areas without potential habitat are eliminated from consideration. The EO likelihood model can be used in the context of both land use planning efforts and conservation planning efforts. By delineating areas with high likelihood of encountering sensitive species or natural communities, the model can be used to direct development away from those areas, or to identify areas worthy of conservation efforts. Limitations: One shortcoming of the EO likelihood model is that all high likelihood areas are treated the same. Whether there is one recent occurrence in the area or thirty recent occurrences, the same high likelihood value is assigned to the area. There is also no allowance for the relative imperilment of the species found in any unit of interest, and there is no numeric value assigned to any of the units of interest that allow them to be compared to each other. File name: Aq_EO_trs_0906.shp 114 Data source: Biot_p – Biotics polygon database created directly from Biotics from version created September 1, 2006. Results: The results of this analysis did not provide significant additional information than the EO frequency count for the aquatic species (Figure 52). This is due to two main issues. The first is the coarseness of the available aquatic habitat data used. The habitat information was taken from IFMAP, and there is only one category for aquatic habitats (water body), whereas the terrestrial habitat was able to be broken up into more categories and hence provide more information. Second, many of the aquatic EOs are relatively old records. Little work has been conducted over the last 10 years on rare fish, macroinvertebrates, and macrophytes. Bio-rarity Score Description: In addition to the EO likelihood value described above, each element occurrence is also assigned three other values, one based on the species global status, one based on the species state status, and one based on the occurrence viability rank. The greater the threat of imperilment to the species, the higher the value assigned to the occurrence. In a similar manner, the higher the quality or viability of each occurrence, the higher the value assigned to it. The biodiversity value of each occurrence is then calculated by adding the values for the global status, state status, and the quality ranking, then multiplying the sum by the EO likelihood value described above. To calculate the biodiversity value of a given PLSS feature, each feature in the PLSS theme is selected in sequence. Next, all the species occurrences intersecting the PLSS feature are selected. Then the biodiversity values of the selected species occurrences are summed and assigned to the PLSS feature. The result is a value for each PLSS unit that is the sum of the biodiversity values of all occurrences falling within the PLSS unit. A cutoff date of September 1, 2006 was used to create the bio-rarity datasets. All records added to the Michigan Natural Features database after this date are not included in this analysis. This analysis is based on terrestrial boundaries (1 mile blocks) to allow for this analysis to be easily combined or overlaid with the terrestrial analysis. Use: Unlike the EO likelihood model, the bio-rarity score allows similar areas to be compared to each other to determine their relative contributions to biodiversity. Because resources for conservation are generally limited, the bio-rarity score can help direct limited resources to those areas where the resources will have the greatest conservation impact. Limitations: As with other element occurrence based information, this data layer is limited by: 1) static information, which is updated infrequently, 2) incomplete data, and 3) old and/or general (non location specific) records. File name: Aq_EO_trs_0906.shp 115 Figure 52. Aquatic element occurrence likelihood map by PLSS. 116 Figure 53. Aquatic element occurrence biological rarity by PLSS. 117 Data source: Biot_p – Biotics polygon database created directly from Biotics from version created September 1, 2006. Results: This analysis did provide some additional information to the EO frequency analysis (Figure 53), but it is likely less informative than the terrestrial analysis. More individual or small clumps of PLSS units are highlighted as important likely due to the status of the EO (S and G rank). This analysis is not as informative because of the limitations with the EO likelihood analysis and the EO rank. For most aquatic species the EO rank is simply extant. We do not have enough information for most aquatic EOs to determine viability of occurrences. Rare Species Richness Description: This analysis counts the number of rare (state-listed and special concern) aquatic plant and animal species that fall within a given sub-watershed. The model utilizes a statewide GIS data layer (Environmental Systems Research Institution (ESRI) shapefile) of sub-watersheds and normalizes the data by river miles (Table 33). River miles are used because the majority of aquatic rare species use riverine habitats. One hundred twenty-five sub-watersheds did not contain a river, based on the NHD hydrology layer. An additional 19 sub-watersheds contained less than 0.1 miles of river, probably due to the inherent geometric inaccuracy of the spatial data layers (Table 34). These 146 sub-watersheds were removed from the analysis. A numeric count field was added to the sub-watershed shapefile theme table and the total number of species based on the MNFI Biotics database was determined. The count was then divided by sub-watershed river miles and then placed in categories based on quartiles. Use: Species richness is another relatively simple representation of the MNFI data. It is designed to show users where there are known rare species rich areas. While the species richness analysis provides limited information, it does fulfill its intended purpose. Limitations: As with the other element occurrence based information, this data layer is limited by: 1) static information, which is updated infrequently, 2) incomplete data, and 3) old and/or general (non location specific) records. File name: aq_EO_richness_subwatershed.shp Data sources: Institute for Fisheries Research, Michigan Department of Natural Resources. mi_epastar_nhd_stresref.shp Biot_p – Biotics polygon database created directly from Biotics from version created September 1, 2006. Results: The results of this analysis generally follow the EO frequency count analysis (Figure 54). Listedspecies richness ranged from 0 to 13 species per sub-watershed. Only four sub-watersheds, three within the River Raisin watershed and one draining directly to Lake St. Clair, had 10 or more listedspecies. Thirty-four sub-watersheds had greater than 5 listed-species; these occurred in the Dead (1 118 sub-watershed), Muskegon (2), Lower Grand (2), St. Joseph (2), St. Joseph of the Maumee (2), Tiffin (1), Raisin (10), Huron (8), Lake St. Clair (4), and Clinton (2) watersheds. When the data was standardize the results are slightly altered. The number of listed-species per river mile ranged from 0 to 6.5. Thirteen sub-watersheds had greater than 2 listed-species per river mile, including: 2 subwatersheds in the Huron watershed with 6.5 and 2.9 listed-species per river mile, four sub-watersheds in the St. Joseph watershed with between 4.4 and 5.6 listed-species per river mile, two subwatersheds in the Upper Grand watershed with 4.07 listed-species per river mile in each, and one subwatershed in the Huron, Lower Grand, Betsie-Platte, Black, Ottawa-Stony, and Manistique watersheds with 2.9, 2.7, 2.5, 2.4, 5.7, and 2.1 listed-species per river mile, respectively. The Southeast Michigan Interlobate and Lake Plain (16+2) EDU had the greatest aquatic species richness and the Saginaw Bay (4) EDU had the least (Table 35). Table 33. Summary statistics on river miles per sub-watershed. Count Minimum Maximum Mean Std Deviation 2,319 0 331.55 19.93 19.05 Table 34. Summary statistics of 19 sub-watersheds that had <0.1 mi of river. Minimum Maximum Mean Median Standard Deviation 0 6.5 0.08 0 0.33 Table 35. Species richness per river mile by EDU. EDU 3 4 5 6+12 7 8 16+2 Richness/river mi (x1000) 3.287 1.986 3.371 3.754 3.296 3.51 7.53 Species of Greatest Conservation Need Richness Description: This analysis counts the number of aquatic animal species of greatest conservation need (SGCN), as listed in Michigan’s Wildlife Action Plan (Eagle et al. 2005), that fall within a given sub-watershed. The model utilizes a statewide GIS data layer (Environmental Systems Research Institution (ESRI) shapefile) of sub-watersheds and normalizes the data by river miles. River miles are used because the majority of aquatic rare species use riverine habitats. A numeric count field is added to the subwatershed shapefile theme table. Each sub-watershed shape is selected in turn and intersected with the available SGCN GIS data. Species richness intersecting each sub-watershed shape is counted and that value is calculated into the count field in the sub-watershed shapefile theme table and then placed in categories based on quantiles. 119 Figure 54. Aquatic rare species richness per river mile in sub-watersheds. Categories are based on quantiles. 120 In addition to the state-listed aquatic animal species, the following SGCN have available point location data: Mussels: pimpleback (Quadrula pustulosa), cylindrical papershell (Anodontoides ferussacianus), creek heelsplitter (Lasmignona compressa), black sandshell (Ligumia recta), threehorn wartyback (Obliquaria reflexa), and kidneyshell (Ptychobranchus fasciolaris). Amphibians (fully aquatic only): mudpuppy (Necturus maculosus maculosus) and western lesser siren (Siren intermedia nettingi). Fish (all SGCN fish have available point location data): brassy minnow (Hybognathus hankinsoni), striped shiner (Luxilus chrysocephalus), silver chub (Macrhybopsis storeriana), river chub (Nocomis micropogon), finescale dace (Phoxinus neogaeus), lake chubsucker (Erimyzon sucetta), spotted sucker (Minytrema melanops), black redhorse (Moxostoma duquesnei), golden redhorse (Moxostoma erythrurum), brown bullhead (Ameiurus nebulosus), stonecat (Noturus flavus), tadpole madtom (Noturus gyrinus), grass pickerel (Esox americanus), pirate perch (Aphredoderus sayanus), slimy sculpin (Cottus cognatus), fantail darter (Etheostoma flabellare), and least darter (Etheostoma microperca). Use: The species of greatest conservation need richness count is another relatively simple representation of known areas important to species biodiversity. While the species richness count provides limited information, it does fulfill its intended purpose. Limitations: As with the other species based information, this data layer is limited by: 1) static information, which is updated infrequently and 2) incomplete data because field sampling is limited, especially for particular species. File name: aq_SGCN_richness_subwatershed.shp Data sources: Institute for Fisheries Research, Michigan Department of Natural Resources mi_epastar_nhd_stresref.shp The Nature Conservancy – Great Lakes Program, Higgins et al. 1998 milakes_w_attributes.shp Digital Water Atlas v1, Fish Atlas 03, Institute for Fisheries Research, MI DNR Fisheries Division Mussel data from University of Michigan Museum of Zoology, created September 27, 2007 Biot_p – Biotics polygon database created directly from Biotics from version created September 1, 2006. 121 Results: This analysis highlighted different sub-watersheds (Figure 55) than the previous analysis. Thirteen sub-watersheds had greater than 15 SGCN located within their borders: 6 sub-watersheds were in the River Raisin watershed, 4 were in the Huron watershed, 2 in the Muskegon watershed, and 1 in the Clinton watershed. Fifty-three sub-watersheds had greater than 10 SGCN within their borders and 1,137 sub-watersheds had no aquatic SGCN reported. However, once the data was standardized by river miles the location of “hot spots” changed. Six sub-watersheds had greater than 5 SGCN per river mile including: 1 sub-watershed in the St. Joseph watershed (Lake Michigan Basin) with 26.6 SGCN per river mile, 2 sub-watersheds in the Huron watershed with 8.7 and 7 SGCN per river mile, one sub-watershed in the Upper Grand watershed with 5.4 SGCN per river mile, and one subwatershed in the Ottawa-Stony and the Betsie-Platte watershed, with 17 and 9.8 SGCN per river mile, respectively. There were 2,617 sub-watersheds with less than 1 SGCN per river mile. The sub-watersheds with high SGCN richness did not always coincide with high listed-species richness due to the plant species that were included in the listed-species list but not in the SGCN list. The Southeast Lake Michigan (3) and the Southeast Michigan Interlobate and Lake Plain (16+2) were the richest EDUs for SGCN. Whereas, the Saginaw Bay (4) and the Western Upper Peninsula and Keweenaw Peninsula (6+12) were the least rich EDUs (Table 36). Table 36. Average species of greatest conservation need (SGCN) richness per river mile by EDU. EDU 3 4 5 6+12 7 8 16+2 Average SGCN richness per river mile 0.29 0.06 0.18 0.06 0.14 0.08 0.44 Best Two Occurrences of Each Element by Watershed Description: The two highest ranking occurrences of each rare aquatic species tracked by MNFI were identified for each watershed and, when possible, at least 10 occurrences across the state were represented. There are a total of 19 aquatic plants (appendix C) and 74 animals (appendix D) currently tracked by MNFI. For this analysis, aquatic plants were strictly defined as plants that are floating or submerged. The ranking of occurrences used viability ranking in EO data, year EO was last observed, and landscape context. Again, there is often little available data to provide an accurate viability ranking, since most animal EOs received an extant ranking. Thus, the other two ranking factors were more important. The most recent EOs are ranked higher. Landscape context for river EOs was accessed using the analysis conducted by Wang et al. (2006), which classifies river reaches across a disturbance gradient (reference to disturbed). Landscape context for lakes was determined by analyzing land use and road density within a 500m buffer around the lakes. Land use is known to affect the quality of aquatic ecosystems and species (Allen 2004). We added road density as part of our landscape context analysis because we felt true land use may be masked in the IFMAP data because often natural vegetation buffers surround lakes, even if housing density is high since many roads are not at a scale that is detectable on Landsat satellite imagery. For those cases where EO viability, last observed date, and landscape quality was a tie, all occurrences were included. 122 Use: In some cases, important element occurrences may be located outside areas deemed significant due to other natural assets such as size, intactness, connectivity, and quality. Identifying areas with high quality EOs regardless of landscape context can be important for ensuring adequate biological representation, and in turn protecting potential genetic variability. Limitations: As with the other element occurrence based information, this data layer is limited by: 1) static information, which is updated infrequently, 2) incomplete data, and 3) old and/or general (non location specific) records. File name: best2_aq_watershed_0906.shp Data sources: Biot_p – Biotics polygon database created directly from Biotics from version created September 1, 2006. Institute for Fisheries Research, Michigan Department of Natural Resources mi_epastar_nhd_stresref.shp The Nature Conservancy – Great Lakes Program, Higgins et al. 1998 milakes_w_attributes.shp Results: A total of 977 EOs were selected to represent the best 2 aquatic EOs within each watershed. The majority of EOs selected for riverine plants occurred in the Erie Basin and the northern tip of the Lower Peninsula, while the majority of EOs selected for lake plants occurred in the western Lower Peninsula and the Upper Peninsula (Figure 56). For riverine fish, the majority of EOs selected were located in the southern lower peninsula, and for lake fish the majority of EOs selected occurred throughout the state. Invertebrate EOs selected, including mussels, were mainly located in the Lower Peninsula. The Raisin, St. Joseph (Lake Michigan Basin), and Huron watersheds had the most EOs selected, partly due to species distributions and sampling effort. Discussion The methodology outlined here provides a key first step in assessing Michigan’s aquatic biodiversity statewide. However due to the nature of the data used in this assessment, we can only point to areas with potential importance to Michigan’s biodiversity. There has been no comprehensive statewide systematic survey to identify locations or habitat types for rare species in Michigan. Currently, we can only provide information based on available known data which has been inconsistently collected. This is not sufficient for understanding what these species need and how best to manage and protect them. As classification frameworks for aquatic habitats become available and finalized in Michigan, we will be able to design systematic surveys to search for rare aquatic species as well as unique ecosystems. This next step will allow us to begin truly quantifying Michigan’s aquatic biodiversity. As stated previously, the methodology developed for this project is a good first step. However, due to the nature of the project and available funding we were unable to conduct a detailed field-expert review. We view this as a critical next step to a robust statewide assessment. Due to the coarseness 123 of some of the data and the emphasis on modeling, we need to begin scrutinizing the results to ensure that we are targeting what is important to Michigan’s biodiversity (species and ecosystems). We also need to ensure that we do not miss key components of Michigan’s biodiversity. In the future we hope to bring together a variety of experts to begin reviewing the results of this project. Additionally, we want to tie this work with other aquatic efforts in the state and continue to develop a more robust statewide assessment of biodiversity. As aquatic habitat classifications become more refined in Michigan, we would like to update our analyses to ensure they provide the most current state of knowledge. We also want to look at this work in the context of The Nature Conservancy’s conservation priority areas and the Wildlife Divisions (DNR) protected lands. The Great Lakes GAP analysis, when completed, will provide more detailed information on important habitats to the diversity of fish in Michigan and will provide information about important Great Lakes’ habitats. Additionally, there are some datasets that we were unable to incorporate but would like to in the future, such as riparian ecosystems of the Lower Peninsula of Michigan (Baker) and Great Lakes ecosystems (in progress – Rutherford and Geddes, Aquatic GAP). By assessing this work in the context of how it fits in with other efforts in the state and a field-based expert review, we will be able to develop a more accurate assessment of Michigan’s aquatic biodiversity. 124 Figure 55. Aquatic species of greatest conservation need richness. Categories are based on quantiles. 125 Figure 56. Locations of the best occurrences for each element by watershed. 126 Looking for Patterns: Integrating the Data Layers Together Introduction As stated earlier in the report, it was decided that the best way to address the various needs of potential end users was to develop a series of data layers that could be used individually or in combination with each other. The previous two chapters addressed the different data layers that were developed for both terrestrial and aquatic biodiversity; however, we haven’t addressed how these data layers may be combined to identify important biodiversity areas based on several variables. From the authors’ perspective, there are two major methods to combining the data layers; merging and prioritizing. Merging is when several data layers containing different datasets are combined together to form an aggregate, and all areas identified are given the same priority. Areas where there is overlap between two or more data layers are not given a higher priority over an area with just one data layer. Data layers that seemed important to incorporate into the identification of core terrestrial biodiversity areas included: 1) bio-rarity hotspots, 2) natural vegetation core areas, 3) high quality natural communities, and 4) potentially unchanged natural vegetation core areas (Table 37). Table 37. Important terrestrial biodiversity area data layers. Data Layer Bio-rarity Hotspots Description Only terrestrial species tracked in MNFI database; only top 10% of scores Natural Vegetation Core Areas All natural vegetation patches that meet a minimum size threshold determined by ecoregion, split by major roads and buffered 210 meters from roads and non-natural land cover High Quality Natural Communities All natural communities with an EO rank of A-B/C Potentially unchanged natural vegetation All potentially unchanged natural vegetation patches that meet a core areas - by ecoregion minimum size threshold determined by ecoregion; split by major roads; no buffer. Prioritizing involves the same steps as merging. The difference is that areas which overlap are given a higher priority. The assumption is that areas containing several components of biodiversity have a higher value than areas that only contain one, and therefore are more valuable. Another way to view this is from an economic perspective. If two areas of approximately the same size contain different values, it makes sense to apply limited resources to the area with more value. Data layers incorporated into the identification of prioritized core terrestrial biodiversity areas included: 1) biorarity hotspots, 2) natural vegetation core areas, 3) high quality natural communities, and 4) potentially unchanged natural vegetation core areas. The resulting data layer is displayed as pixels with a score ranging from 0 (no data layers) to 4 (all four data layers) (Table 38). Note that a score of 3 or 4 requires the occurrence of a high quality natural community and/or high biorarity score. Both of these data layers are based on field observations that are biased towards certain species and natural communties, as well as certain areas of the state. One way to interpret this analysis is that: 1) all areas receiving a score of one or greater are important, 2) areas recieving a score of three or four may be the best places to focus on initially, and 3) a score of zero does not mean an area is unimportant to biodiversity conservation (could be due to lack of survey effort). 127 Table 38. Prioritized terrestrial biodiversity area descriptions. Data Layers Score Bio-rarity Hotspots Existing Data type Grid 1 2 Converted to: 30m2 pixel Natural Vegetation Core Areas 1 30m pixel 30m2 pixel High Quality Natural Communities 1 polygon 30m2 pixel Potentially Unchanged Natural Vegetation Core Areas – by ecoregion 1 30m2 pixel 30m2 pixel In addition to the prioritized biodiversity areas, since many of these areas are small and/or isolated, it seemed important to incorporate those lands that may support these core biodiversity areas. This is called the supporting natural landscape, a term borrowed from the Massachusetts BioMap project. The supporting natural landscape was defined as all natural vegetation patches with no roads and no buffer that intersected with a core biodiversity area. We also provided one example of prioritized core aquatic biodiversity areas in the state. We incorporated the two best classes of the functional sub-watersheds with the best two classes of the SGCN richness data layer. The resulting data layer displays sub-watersheds where the two data layers overlap (Figure 58). Terrestrial Results: A total of 12,609,097 acres fell into one of four categories of prioritized terrestrial biodiversity areas in the state. Using the criteria described above, these areas combined to represent approximately 35% of the total area of the state (not including inland water) (Table 39). Although the majority of these areas were located in the UP and NLP, the highest priority areas with scores of 3 and 4 were distributed across the state (Figure 57). High priority areas in the UP included: 1) Seney National Wildlife Refuge, 2) Grand Island National Recreation Area, 3) area just north of St. Ignace, 4) Lake Michigan shoreline in western Mackinac County and eastern Schoolcraft County, 5) Tahquamenon State Park, 6) Porcupine Mountains Wilderness State Park, 7) northern Marquette County, and 8) the north portion of the Keweenaw Peninsula. High priority areas in the NLP included: 1) Wilderness State Park, 2) Thompson’s Harbor State Park, 3) eastern portion of Thunder Bay – east of Alpena, 4) large portions of the Au Sable watershed, 5) southeast Newaygo County, and 6) the Blue Lakes region of Oceana and Muskegon Counties. High priority areas in the SLP included: 1) Allegan State Game Area, 2) Fort Custer Recreation Area, 3) Pinckney-Waterloo Recreation Areas, and 4) St. Clair Flats. Table 39. Summary of prioritized terrestrial biodiversity area scores. Score Total area in acres 1 2 3 4 Total 9,045,789 3,371,944 184,995 6,369 12,609,097 % of State (not including water) 24.90% 9.28% 0.51% 0.02% 34.71% Aquatic Results: A total of 78 sub-watersheds were selected as relatively functional and important to species of greatest conservation need. The selected sub-watersheds occurred in all EDUs but were most 128 Figure 57. Prioritized terrestrial biodiversity areas displayed at a 1 mile2 resolution. 129 prevalent in the Northern Lake Michigan, Lake Huron, and Straits of Mackinac and Easter Upper Peninsula EDUs, with 23 and 26 sub-watersheds selected, respectively. Five of the sub-watersheds occurred in the Southeast Michigan Interlobate and Lake Plain EDU (Bean Creek, the West Fork of the West Branch of the St. Joseph River, the River Raisin, and two are along the lake shore); the Southeast Lake Michigan (3) EDU had 6 sub-watersheds highlighted (Grand River, Looking Glass River, Battle Creek, South Branch of the Kalamazoo River, St. Joseph River, and Pigeon River); Saginaw Bay (4) EDU had 5 (South Branch of the Flint River, Shiawassee River, Molasses River, Black River, and one along the lake shore), Central Upper Peninsula (8) EDU had 9, and the Western Uppper Peninsula and Keweenaw Peninsula (6+12) EDU had 4 sub-watersheds (Tenderfoot Creek, West Branch of the Presque Isle River, and two watersheds that drain directly into Lake Superior) highlighted. Additional ways to bring the data layers together Aside from the example provided above to identify and prioritize potentially important biodiversity areas, there are many additional ways to analyze or overlay the different data layers described in this report to identify important natural resource areas in the state. The first example provided below focuses on identifying and prioritizing sites along the Great lakes shoreline. This analysis is important to conduct as a separate product due to the global and regional significance of the Great Lakes shoreline in Michigan. In addition, three other major categories of analysis that could be further explored include (but not limited to): 1) bio-rarity hotspots, 2) high quality natural communities, and 3) natural land cover types. Examples of a few analyses that could be conducted are listed under each of the headings. Lastly, it is important to identify gaps in protection by overlaying the data layer or layers you think are most important with the latest conservation lands or public lands data layer. Great Lakes shoreline An analysis was conducted to identify and prioritize sites along the Great Lakes shoreline which support concentrations of threatened and endangered species. The first step of the analysis involved selecting all natural community element occurrences, and all plant and animal occurrences from the MNFI database within a distance of 0.5 miles of the Michigan portion of the Great Lakes shoreline. Plant and animal occurrences greater than 20 years old were discarded. The shoreline layer was derived from the Michigan County layer, at 1:24,000 scale, and consists of a line delineating the entire Great Lakes shoreline of Michigan. The resulting features were buffered by 0.5 kilometers, and the boundaries between overlapping buffers were dissolved to create a new layer of shoreline sites. The newly created sites were then scored using specific criteria outlined in the biological rarity score. The biological rarity model is generated by assigning each element occurrence a value based on the age of the record. This value is used to represent the probability that the occurrence still exist. Each element occurrence is also assigned three other values, one based on the species global status, one based on the species State status, and one based on the element occurrence quality rank. The greater the threat of imperilment to the species and the higher the quality of each occurrence, the higher the value assigned to the occurrence. Sites were then ranked based on the summed biological rarity scores. File name: GL shoreline sites\sites.shp Data source: Biot_p – Biotics polygon database created directly from Biotics from version created September 1, 2006. 130 Figure 58. Prioritized aquatic biodiversity areas based on species of greatest conservation concern and functional sub-watersheds. 131 Results: A total of 1,960 element occurrences (all natural communities and only plant and animal occurrences observed within the last 20 years) were located within .5 miles of one of the Great Lakes. This represents 13% of the database. Once these occurrences were buffered by 0.5 kilometers and merged together, a total of 461 distinct sites were identified along the Great Lakes shoreline. Biorarity scores ranged from a low of 4 to a high of 1,957. The five sites with the highest scores were: 1) north half of Isle Royale, 2) Schoolcraft County shoreline, 3) Wilderness State Park, 4) Seiner’s Point to Big Knob Campground, and 5) Drummond Island-Maxton Plains (Figure 59). Bio-rarity hotspots High terrestrial species bio-rarity score. Purpose is to identify areas with high unique natural features value regardless of patch or landscape integrity. [High terrestrial species bio-rarity score] intersected with [all natural vegetation – all roads – 210m buffer]. Purpose is to identify areas with high unique natural features value located within landscapes of high ecological integrity. These sites are important because they contain a concentration of high quality natural features that have the best opportunity for long-term viability. [High terrestrial species bio-rarity score] intersected with [matrix – all roads – 210m buffer]. Purpose is to identify areas with high unique natural features value located within landscapes of high ecological integrity. These sites are important because they contain a concentration of high quality natural features that have the best opportunity for long-term viability. High quality natural communities [High quality natural communities] intersected with [all natural vegetation – all roads - 210m buffer] – Identify high quality natural communities located within landscapes of high integrity. [High quality natural communities] intersected with [matrix – all roads – 210m buffer] - Identify high quality natural communities located within landscapes of high integrity. Natural landcover types [natural vegetation types – all roads – 210m buffer]. Purpose is to identify areas with high patch integrity regardless of landscape integrity. [natural vegetation types – all roads – 210m buffer] intersected with [all natural vegetation – all roads - 210m buffer]. Purpose is to identify areas with high patch and landscape integrity that have the potential to harbor a high diversity of plants and animals and/or rare species. [natural vegetation types – all roads – 210m buffer] intersected with [matrix – all roads – 210m buffer] - Identify areas with high patch and landscape integrity that have the potential to harbor a high diversity of plants and animals and/or rare species. Unique Aquatic ecosystems [Unique river ecosystems] and [Unique lake ecosystems] intersected with [functional subwatersheds] – Identify where there are higher concentrations of unique ecosystems within functional subwatersheds. This analysis could also help prioritize areas to survey for aquatic elements. Unique river ecosystems] and [Unique lake ecosystems] intersected with [SGCN richness] – Identity sub-watersheds that may be key areas for overall aquatic biodiversity. 132 Figure 59. High priority great lakes shoreline sites. 133 Ownership Patterns Lastly, it is important to also highlight important biodiversity lands that are under the highest degree of threat. The simplest way to accomplish this is to overlay the various data layers mentioned earlier with a public lands data layer. When using land ownership this way, we are assuming that the public lands shown on this data layer are at least somewhat protected from development or habitat destruction. From this perspective, the resulting maps will highlight private lands that fall within important biodiversity areas. Based on the most recent Conservation and Recreation Lands (CARL) database, developed by Ducks Unlimited and The Nature Conservancy, approximately 21% of the land in Michigan is under public ownership. However, these ownership patterns are not evenly distributed. The Eastern Upper Peninsula leads the state with 47% of the land in public ownership. This is followed by the western Upper Peninsula with 35%, the northern Lower Peninsula with 25%, and the southern Lower Peninsula with only 5%. 134 Next Steps Assessing the state of Michigan’s biodiversity and identifying important areas for conservation is far from complete. The primary focus of this initial effort was to gather, develop, and assess a series of data layers for both terrestrial and aquatic natural features that could be used for future conservation planning efforts at multiple scales. Given this basis of information, there are five categories of next steps: 1) gathering needed data, 2) conducting field-expert reviews, 3) examining this work in light of other efforts in the state (TNC’s conservation priority areas, and the terrestrial and aquatic Michigan GAP projects), 4) updating models, and 5) setting conservation priorities. First, this project helped crystallize that more data is needed to aid in more effective models and analyses. One element that is still missing and that plays a key role in conducting a critical assessment of Michigan’s biodiversity is a comprehensive, systematic biological survey. Very few places in Michigan have had a systematic survey of its natural features. The vast majority of areas where surveys have been conducted are publicly owned, and our knowledge of the places that have been surveyed is incomplete. We need comprehensive statewide data, as well as more data on species viability. The majority of rare animal element occurrences in the MNFI database have an EO rank of E for extant. As a result, all of the data layers that utilize rare animal occurrence data, specifically the EO rank, are not as robust as they could be. Predictive models for species can help identify areas with potentially high species diversity or areas important for particular guilds of species, such as wading birds. As part of the Michigan GAP Project, predictive models were developed for 327 vertebrate terrestrial species. The Michigan Aquatic GAP analysis is still in progress. However, in order to obtain the level of confidence needed to effectively model where important natural features occur across the state of Michigan, we need field data that is more comprehensive, accurate, and complete. Other data needs include: • Biotic and abiotic surveys of significant sites identified through GIS models to determine if those sites truly are significant and/or unique. • A scientifically defensible lakes classification system in Michigan. • Defined riverine natural communities with associated species • Improved methodology for identifying high quality natural land cover Second, since this project relied on broad GIS data and modeling to conduct our analyses, a detailed field-expert review is needed to determine the accuracy and validity of our methods. Three key reasons to include regional experts in the review of our work are to: 1) gather data to fine tune the models; 2) set priorities for field surveys, and 3) expand ownership of the assessment. Although we were unable to include a large stakeholder or user group in the development of this project, we understand the importance of stakeholder input. We believe an expert review is an important next step. Third, we need to tie our work to other statewide efforts in Michigan. Future aquatic classifications (lakes, Great Lakes) should be examined for their utility in an updated aquatic assessment. We would also like to examine how our results fit in with TNC’s conservation priority areas for Michigan and the Great Lakes, as well as the terrestrial and aquatic Michigan GAP projects. Examining the variety of conservation and natural resource efforts in Michigan allows us to more accurately identify where there are gaps in knowledge. Fourth, to begin setting conservation priorities for Michigan’s biodiversity, we need to determine 135 important areas for both terrestrial and aquatic biodiversity, and identify and design an interconnected network of conservation areas, including connecting corridors. Although the initial efforts for the aquatic and terrestrial assessment needed to be completed separately, we now need to determine the best way to bring these two different components together. By connecting these components in a scientifically defensible, efficient, and meaningful way, we can begin prioritizing areas across the state that are potentially important for both aquatic and terrestrial biodiversity. More than likely important areas will compliment each other, and GIS tools that evaluate adjacency and proximity will help identify where these areas of terrestrial and aquatic features converge on the landscape. Fifth, this report provides examples of how the data could be used to spatially identify important areas on the landscape; however designing an interconnected network of conservation areas is a bit more complicated. One key element that still needs to be addressed is connecting corridors or linkages between important areas and sites. Corridors can be difficult to identify because: 1) their location and design are dependent on the specific requirements of the biotic and/or abiotic target(s), and 2) obstacles such as roads, development, dams, large scale intensive agricultural operations, railroads, and other non-natural land cover types fragment the landscape, restrict opportunities and lead to numerous design challenges. One way to address an interconnected network of conservation areas is by developing green infrastructure plans at multiple scales. Green infrastructure plans essentially consist of three design elements: hubs, sites, and linkages. Hubs are large areas of natural land that act as anchors for a variety of natural processes, and provide an origin or destination for many species of wildlife. Hubs tend to have a wide diversity of habitats, and are resilient to natural disturbances such as fire, flooding, and wind throw. At the next finer scale, it is important to identify sites. Sites are smaller landscape areas that incorporate smaller-scale ecologically important features. They tend to be well defined, isolated places on the landscape, such as an isolated wetland, a sink hole, or a great blue heron rookery. Once these are identified, it is important to identify a suite a species that would benefit from a corridor or linkage between two or more hubs or sites. Again, multiple scales need to be considered. Wide ranging terrestrial species such as black bear, moose, elk, martin, or bobcat, or migrating fish might be good candidates for the design and incorporation of linkages at the 102 to 103 m scale. On a finer scale, smaller ranging species that require multiple habitat types for survival, such as many of Michigan’s snakes and turtles, or species whose populations are characterized as meta-populations, might be good candidates for the design of very site specific travel corridors at the 101 m scale. Determining effective and meaningful conservation areas is a difficult and complicated endeavor, but through this report we now have data to help Michigan’s resource agencies, conservation organizations, and concerned citizens begin the process. This report provides the basis for the next steps in completing a comprehensive and robust assessment of Michigan’s biodiversity. Despite the several areas of improvement mentioned above, the data layers provided in this report reflect the best information currently available on Michigan’s biodiversity at the state and regional scales. Data layers that are particularly weak due to lack of empirical data are the unique lakes and streams analyses. However, the rest of the data layers provided in this report, particularly the terrestrial layers, should meet the majority of end user needs. The information and data on terrestrial and aquatic biodiversity can be used by: 1) government agencies to help develop conservation plans at multiple scales, 2) local units of government that are interested in creating green infrastructure plans or updating their parks and recreation plans, 3) watershed councils for watershed planning and protection, and 4) land conservancies for prioritizing lands for permanent protection. All of these efforts should include field visits to verify the modeling results. 136 References Cited Allan, J.D. 2004. Landscapes and riverscapes: the influence of land use on river ecosystems. Annual Reviews of Ecology, Evolution and Systematics 35:257-284. Abell, R., D.M. Olson, E. Dinerstein, P.T. Hurley, et al. 2000. Freshwater ecoregions of North America: a conservation assessment. World Wildlife Fund – United States. Island Press, Washington, D.C., USA. Abell, R., M. Thieme, E. Dinerstein, and D. Olson. 2002. 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Science 147:250-259. 141 Appendix A - Rare terrestrial plant list Species Adlumia fungosa Agalinis gattingeri Agalinis skinneriana Agoseris glauca Agrimonia rostellata Agropyron spicatum Allium schoenoprasum Amerorchis rotundifolia Amorpha canescens Androsace occidentalis Angelica venenosa Antennaria parvifolia Antennaria rosea Arabis missouriensis var. deamii Arabis perstellata sensu lato Arenaria macrophylla Aristida dichotoma Aristida longespica Aristida tuberculosa Aristolochia serpentaria Arnica cordifolia Artemisia ludoviciana Asclepias hirtella Asclepias ovalifolia Asclepias purpurascens Asclepias sullivantii Asplenium montanum Asplenium rhizophyllum Asplenium ruta-muraria Asplenium scolopendrium var. americanum Asplenium trichomanes-ramosum Aster furcatus Aster modestus Aster praealtus Aster sericeus Astragalus canadensis Astragalus neglectus Baptisia lactea Baptisia leucophaea Bartonia paniculata Beckmannia syzigachne Berula erecta Besseya bullii Betula murrayana Botrychium acuminatum Botrychium campestre Botrychium hesperium Botrychium mormo Botrychium pallidum Common name Climbing fumitory Gattinger's gerardia Skinner's gerardia Prairie or pale agoseris Beaked agrimony Bluebunch wheatgrass Wild chives Round-leaved orchis Leadplant Rock-jasmine Hairy angelica Pussy-toes Rosy pussytoes Missouri rock-cress Rock cress Big-leaf sandwort Shinner's three-awned grass Three-awned grass Beach three-awned grass Virginia snakeroot Heart-leaved arnica Western mugwort Tall green milkweed Dwarf milkweed Purple milkweed Sullivant's milkweed Mountain spleenwort Walking fern Wall-rue Hart's-tongue fern Green spleenwort Forked aster Great northern aster Willow aster Western silvery aster Canadian milk-vetch Cooper's milk-vetch White or prairie false indigo Cream wild indigo Panicled screw-stem Slough grass Cut-leaved water-parsnip Kitten-tails Murray birch Acute-leaved moonwort Prairie moonwort Western moonwort Goblin moonwort pale moonwort Fed State SC E E T SC X T E SC E SC SC T SC T T X T T T E T T E SC T X T E LT E T T T SC T T SC SC E T T T T SC E T T T SC Grank G4 G4 G3 G5 G5 G5 G5 G5 G5 G5 G5 G5 G5 G4G5QT3?Q G5 G4 G5 G5 G5 G4 G5 G5 G5 G5? G5? G5 G5 G5 G5 G4T3 G4 G3 G5 G5 G5 G5 G4 G4Q G4G5T4T5 G5 G5 G4G5 G3 G1Q G1 G3G4 G3G4 G3 G2G3 A-1 Srank S3 S1 S1 S2 S1S2 SX S2 S1 S3 SH S3 S1 SH S2 S1 S1 SX S2 S1 S2 S1 S1 S2 S1 S3 S2 SH S2S3 S1 S1 S2S3 S1 S1 S3 S2 S1S2 S3 S3 S1 S2 S2 S2 S1S2 S1 S1 S2 S2 S2 S3 Appendix A - Rare terrestrial plant list - continued Species Bouteloua curtipendula Braya humilis Bromus pumpellianus Buchnera americana Cacalia plantaginea Calamagrostis lacustris Calamagrostis stricta Calypso bulbosa Camassia scilloides Carex albolutescens Carex assiniboinensis Carex atratiformis Carex concinna Carex conjuncta Carex crus-corvi Carex davisii Carex decomposita Carex festucacea Carex frankii Carex gravida Carex haydenii Carex heleonastes Carex lupuliformis Carex media Carex nigra Carex novae-angliae Carex oligocarpa Carex platyphylla Carex richardsonii Carex rossii Carex scirpoidea Carex seorsa Carex squarrosa Carex straminea Carex tincta Carex trichocarpa Carex typhina Carex wiegandii Castanea dentata Castilleja septentrionalis Ceanothus sanguineus Celtis tenuifolia Chamaerhodos nuttallii var. keweenawensis Chasmanthium latifolium Chelone obliqua Cirsium hillii Cirsium pitcheri Clematis occidentalis Collinsia parviflora A-2 Common name Side-oats grama grass Low northern rock-cress Pumpelly's brome grass Blue-hearts Prairie indian-plantain Northern reedgrass Narrow-leaved reedgrass Calypso or fairy-slipper Wild-hyacinth Greenish-white sedge Assiniboia sedge Sedge Beauty sedge Sedge Raven's-foot sedge Davis's sedge Log sedge Fescue sedge Frank's sedge Sedge Hayden's sedge Hudson bay sedge False hop sedge Sedge Black sedge New england sedge Eastern few-fruited sedge Broad-leaved sedge Richardson's sedge Ross's sedge Bulrush sedge Sedge Sedge Straw sedge Sedge Hairy-fruited sedge Cat-tail sedge Wiegand's sedge American chestnut Pale indian paintbrush Redstem ceanothus Dwarf hackberry Keweenaw rock-rose Wild-oats Purple turtlehead Hill's thistle Pitcher's thistle Purple clematis Small blue-eyed mary Fed State T T T X SC T T T T T T T SC T T SC X SC SC X X E T T E T T T SC T T T SC E SC SC T T E T T SC E T E SC LT T SC T Grank G5 G5 G5T4 G5? G4G5 G3Q G5 G5 G4G5 G5 G4G5 G5 G4G5 G4G5 G5 G4 G3 G5 G5 G5 G5 G4 G4 G5T5? G5 G5 G4 G5 G4 G5 G5 G4 G4G5 G5 G4G5 G4 G5 G3 G4 G5 G4G5 G5 G5T1Q G5 G4 G3 G3 G5 G5 Srank S1S2 S1 S2 SX S3 S1 S1 S2 S2 S2 S2 S2 S3 S1 SH S3 SX S1 S2S3 SX SX S1 S2 S2S3 S1 S1 S2 S1 S3S4 S2 S2 S2 S1 SH SNR S2 S1 S2 S1S2 S2S3 S2 S3 S1 S1 S1 S3 S3 S3 S2 Appendix A - Rare terrestrial plant list - continued Species Commelina erecta Coreopsis palmata Corydalis flavula Crataegus douglasii Cryptogramma acrostichoides Cryptogramma stelleri Cuscuta campestris Cuscuta glomerata Cuscuta indecora Cuscuta pentagona Cuscuta polygonorum Cyperus acuminatus Cyperus flavescens Cypripedium arietinum Cypripedium candidum Cystopteris laurentiana Dalea purpurea Dalibarda repens Danthonia compressa Danthonia intermedia Dasistoma macrophylla Dennstaedtia punctilobula Dentaria maxima Diarrhena americana Digitaria filiformis Disporum hookeri Disporum maculatum Disporum trachycarpum Dodecatheon meadia Draba arabisans Draba cana Draba glabella Draba incana Draba nemorosa Draba reptans Drosera anglica Dryopteris celsa Dryopteris filix-mas Dryopteris fragrans Echinacea purpurea Echinodorus tenellus Eleocharis atropurpurea Eleocharis caribaea Eleocharis compressa Eleocharis engelmannii Eleocharis equisetoides Eleocharis melanocarpa Eleocharis microcarpa Eleocharis nitida Common name Slender day-flower Prairie coreopsis Yellow fumewort Douglas's hawthorn American rock-brake Slender cliff-brake Field dodder Rope dodder Dodder Dodder Knotweed dodder Nut-grass Yellow nut-grass Ram's head lady's-slipper White lady-slipper Laurentian fragile fern Purple prairie-clover False-violet Flat oat grass Wild oat-grass Mullein foxglove Hay-scented fern Large toothwort Beak grass Slender finger-grass Fairy bells Nodding mandarin northern fairy bells Shooting-star Rock whitlow-grass Ashy whitlow-grass Smooth whitlow-grass Twisted whitlow-grass Whitlow-grass Creeping whitlow-grass English sundew Log fern Male fern Fragrant cliff woodfern Purple coneflower Dwarf burhead Purple spike-rush Spike-rush Flattened spike-rush Engelmann's spike-rush Horsetail spike-rush Black-fruited spike-rush Small-fruited spike-rush Slender spike-rush Fed State X T T SC E SC SC SC SC SC SC X SC SC T SC X T SC SC T X T T X E X T E SC T E T X T SC T SC SC X E E T T SC SC SC E E Grank G5 G5 G5 G5 G5 G5 G5T5 G5 G5 G5 G5 G5 G5 G3 G4 G3 G5 G5 G5 G5 G4 G5 G5Q G4? G5 G5 G3G4 G5 G5 G4 G5 G4G5 G5 G5 G5 G5 G4 G5 G5 G4 G5? G4G5 G4G5 G4 G4G5Q G4 G4 G5 G3G4 A-3 Srank SX S2 S2 S3S4 S2 S3S4 SH SH SH SH S2 SX S2S3 S3 S2 S1S2 SX S1S2 S1 S1S2 S1S2 SNR S1S2 S2 SX S1 SX S1 S1 S3 S1 S1 S1 SX S1 S3 S2 S3 S3 SX S1 S1 S1 S2 S2S3 S3 S3 S1 S1 Appendix A - Rare terrestrial plant list - continued Species Eleocharis parvula Eleocharis radicans Eleocharis tricostata Elymus glaucus Elymus mollis Empetrum nigrum Equisetum telmateia Eragrostis capillaris Eragrostis pilosa Erigeron acris Erigeron hyssopifolius Eryngium yuccifolium Euonymus atropurpurea Eupatorium fistulosum Eupatorium sessilifolium Euphorbia commutata Euphrasia hudsoniana Euphrasia nemorosa Festuca scabrella Filipendula rubra Fimbristylis puberula Fraxinus profunda Fuirena squarrosa Galearis spectabilis Galium kamtschaticum Gentiana flavida Gentiana linearis Gentiana puberulenta Gentiana saponaria Gentianella quinquefolia Geum triflorum Geum virginianum Gillenia trifoliata Glyceria acutiflora Gnaphalium sylvaticum Gratiola aurea Gratiola virginiana Gymnocarpium jessoense Gymnocarpium robertianum Gymnocladus dioicus Hedyotis nigricans Hedysarum alpinum Helianthus hirsutus Helianthus microcephalus Helianthus mollis Hemicarpha micrantha Hibiscus laevis Hibiscus moscheutos Hieracium paniculatum A-4 Common name Dwarf spike-rush Spike-rush Three-ribbed spike-rush Blue wild-rye American dune wild-rye Black crowberry Giant horsetail Love grass Small love grass fleabane Hyssop-leaved fleabane Rattlesnake-master Wahoo Hollow-stemmed joe-pye-weed Upland boneset Tinted spurge Eyebright Common eyebright Rough fescue Queen-of-the-prairie Chestnut sedge Pumpkin ash Umbrella-grass Showy orchis Bedstraw White gentian Narrow-leaved gentian Downy gentian Soapwort gentian Stiff gentian Prairie-smoke Pale avens Bowman's root Manna grass Cudweed Hedge-hyssop Round-fruited hedge hyssop Northern oak fern Limestone oak fern Kentucky coffee-tree Hedyotis Alpine sainfoin Whiskered sunflower Small wood sunflower Downy sunflower Dwarf-bulrush Smooth rose-mallow Swamp rose-mallow Panicled hawkweed Fed State T X T SC SC T X SC SC SC T T SC T T T T T T T X T T T T E T E X T T SC T X T T T E T SC X E SC X T SC SC SC SC Grank G5 G5 G4 G5 G5 G5 G5 G5 G4 G5 G5 G5 G5 G5? G5 G5 G5? G5 G5 G4G5 G5 G4 G4G5 G5 G5 G4 G4G5 G4G5 G5 G5 G5 G5 G4G5 G5 G3G4 G5 G5 G5 G5 G5 G5 G5 G5 G5 G4G5 G5 G5 G5 G5 Srank S1 SX S2 S3 S3 S2 SX SH SH SR S1 S2 S3 S1 S1 S1 SNR S1 S2S3 S2 SX S2 S2 S2 S1 S1 S2 S1 SX S2 S2S3 S1S2 S1 SX S1 S1S2 S1 S1 S2 S3S4 SX S1 S3 SX S2 S3 SH S3S4 S2 Appendix A - Rare terrestrial plant list - continued Species Houstonia caerulea Huperzia appalachiana Huperzia selago Hybanthus concolor Hydrastis canadensis Hymenoxys herbacea Hypericum gentianoides Hypericum sphaerocarpum Ipomoea pandurata Iris lacustris Isoetes engelmannii Isotria medeoloides Isotria verticillata Jeffersonia diphylla Juncus brachycarpus Juncus militaris Juncus scirpoides Juncus stygius Juncus vaseyi Justicia americana Kuhnia eupatorioides Lactuca floridana Lactuca pulchella Lechea minor Lechea pulchella Lechea stricta Lespedeza procumbens Leucospora multifida Liatris punctata Liatris squarrosa Linum sulcatum Linum virginianum Liparis liliifolia Listera auriculata Lithospermum incisum Lithospermum latifolium Littorella uniflora Lonicera involucrata Ludwigia alternifolia Ludwigia sphaerocarpa Luzula parviflora Lycopodiella margueriteae Lycopodiella subappressa Lycopus virginicus Lygodium palmatum Lysimachia hybrida Mertensia virginica Mikania scandens Mimulus alatus Common name bluets mountain fir-moss Fir clubmoss Green violet Goldenseal Lakeside daisy Gentian-leaved st. john's-wort Round-fruited st. john's-wort Wild potato-vine Dwarf lake iris Appalachian quillwort Smaller whorled pogonia Whorled pogonia Twinleaf Short-fruited rush Bayonet rush Scirpus-like rush Moor rush Vasey's rush Water-willow False boneset Woodland lettuce Blue lettuce Least pinweed Leggett's pinweed Erect pinweed Trailing bush-clover conobea Dotted blazing-star Blazing-star Furrowed flax Virginia flax Purple twayblade Auricled twayblade Narrow-leaved puccoon Broad-leaved puccoon American shore-grass Black twinberry Seedbox Globe-fruited seedbox Small-flowered woodrush northern prostrate clubmoss Northern appressed clubmoss Virginia water-horehound Climbing fern Swamp candles Virginia bluebells Mikania Wing-stemmed monkey-flower Fed State SC E SC SC T LT E SC T T LT T E LT E T SC T T T T T T SC T T SC T SC X SC X X SC T SC SC X SC SC T SC T T T SC T E SC T X X Grank G5 G4G5 G5 G5 G4 G2 G5 G5 G5 G3 G4 G2 G5 G5 G4G5 G4 G5 G5 G5? G5 G5 G5 G5T5 G5 G5 G4? G5 G5 G5 G5 G5 G4G5 G5 G3 G5 G4 G5 G4G5 G5 G5 G5 G2 G2 G5 G4 G5 G5 G5 G5 Srank SNR S? S3 S3 S2 S1 S3 S1 S2 S3 S1 S1 S2 S3 S1S2 S1 S2 S1S2 S1S2 S2 S2 S2 SH SH S1S2 S1 SX SNR SX SX S2S3 S2 S3 S2S3 SX S2 S2S3 S2 S3 S1 S1 S2 S2 S2 S1 S2 S2 SX SX A-5 Appendix A - Rare terrestrial plant list - continued Species Mimulus glabratus var. michiganensis Mimulus guttatus Monarda didyma Morus rubra Muhlenbergia cuspidata Muhlenbergia richardsonis Onosmodium molle Ophioglossum vulgatum Oplopanax horridus Opuntia fragilis Orobanche fasciculata Oryzopsis canadensis Osmorhiza depauperata Oxalis violacea Panax quinquefolius Panicum leibergii Panicum longifolium Panicum microcarpon Panicum polyanthes Panicum verrucosum Parnassia palustris Paronychia fastigiata Pellaea atropurpurea Penstemon calycosus Penstemon gracilis Penstemon pallidus Petasites sagittatus Phacelia franklinii Phaseolus polystachios Phleum alpinum Phlox bifida Phlox maculata Pinguicula vulgaris Piperia unalascensis Plantago cordata Platanthera ciliaris Platanthera leucophaea Poa alpina Poa canbyi Poa paludigena Polemonium reptans Polygala cruciata Polygala incarnata Polygonatum biflorum var. melleum Polygonum careyi Polygonum viviparum Polymnia uvedalia Polytaenia nuttallii Populus heterophylla A-6 Common name Michigan monkey-flower Western monkey-flower Oswego tea Red mulberry Plains muhly Mat muhly Marbleweed Southeastern adder's tongue Devil's-club Fragile prickly-pear Fascicled broom-rape Canada rice-grass Sweet cicely Violet wood-sorrel Ginseng Leiberg's panic-grass Long-leaved panic-grass Small-fruited panic-grass Round-seed panic grass Warty panic-grass Marsh grass-of-parnassus Low-forked chickweed Purple cliff-brake Smooth beard tongue Slender beard-tongue Pale beard tongue Sweet coltsfoot Franklin's phacelia Wild bean Mountain timothy Cleft phlox Spotted phlox Butterwort Alaska orchid Heart-leaved plantain Orange or yellow fringed orchid Prairie fringed orchid Alpine bluegrass Canby's bluegrass Bog bluegrass Jacob's ladder or greek-valerian Cross-leaved milkwort Pink milkwort Honey-flowered solomon-seal Carey's smartweed Alpine bistort Large-flowered leafcup Prairie-parsley Swamp or black cottonwood Fed State LE E SC X T X T X T T E T T T T T T T SC E T T SC T T E SC T T SC X T T SC SC E T LT E T E T T SC X X T T T X E Grank G5T1 G5 G5 G5 G4 G5 G4G5 G5 G4 G4G5 G4 G5 G5 G5 G3G4 G5 G4 G5T5 G5T5 G4 G5 G5 G5 G5 G5 G5 G5 G5 G4 G5 G5? G5 G5 G5 G4 G5 G2 G5 G4G5 G3 G5 G5 G5 G5TH G4 G5 G4G5 G5 G5 Srank S1 S1 SX S2 SX S2 SX S1 S2 S1 S2 S2 S2 S1 S2S3 S2 S2 S2 S1 S1 S2 SH S2 S2 S1 S3 S1S2 S1 SH SX S1 S1 S3 S2S3 S1 S2 S1 S1S2 S1 S2 S2 S3 SX SX S1S2 S1S2 S1 SX S1 Appendix A - Rare terrestrial plant list - continued Species Potentilla paradoxa Potentilla pensylvanica Proserpinaca pectinata Prunus alleghaniensis var. davisii Psilocarya scirpoides Pterospora andromedea Pycnanthemum muticum Pycnanthemum pilosum Pycnanthemum verticillatum Quercus shumardii Ranunculus ambigens Ranunculus cymbalaria Ranunculus lapponicus Ranunculus macounii Ranunculus rhomboideus Rhexia mariana var. mariana Rhexia virginica Rhynchospora globularis Rhynchospora macrostachya Ribes oxyacanthoides Rotala ramosior Rubus acaulis Rudbeckia subtomentosa Ruellia humilis Ruellia strepens Rumex occidentalis Sabatia angularis Sagina nodosa Sagittaria montevidensis Salix pellita Salix planifolia Sanguisorba canadensis Sarracenia purpurea ssp. heterophylla Saxifraga paniculata Saxifraga tricuspidata Scirpus clintonii Scirpus hallii Scirpus olneyi Scirpus torreyi Scleria pauciflora Scleria reticularis Scleria triglomerata Scutellaria elliptica Scutellaria incana Scutellaria nervosa Scutellaria ovata Scutellaria parvula Senecio congestus Senecio indecorus Common name Sand cinquefoil Prairie cinquefoil Mermaid-weed Alleghany or sloe plum Bald-rush Pine-drops Mountain-mint Hairy mountain-mint Whorled mountain-mint Shumard's oak Spearwort Seaside crowfoot Lapland buttercup Macoun's buttercup Prairie buttercup Maryland meadow-beauty Meadow-beauty Globe beak-rush Tall beak-rush Northern gooseberry Tooth-cup Dwarf raspberry Sweet coneflower Hairy ruellia Smooth ruellia Western dock Rose-pink Pearlwort Arrowhead Satiny willow Tea-leaved willow Canadian burnet Yellow pitcher-plant Encrusted saxifrage Prickly saxifrage Clinton's bulrush Hall's bulrush Olney's bulrush Torrey's bulrush Few-flowered nut-rush Netted nut-rush Tall nut-rush Hairy skullcap Downy skullcap Skullcap Heart-leaved skullcap Small skullcap Marsh-fleabane Rayless mountain ragwort Fed State T T E SC T T T T SC SC T T T T T T SC E SC SC SC E X T T E T T T SC T T T T T SC T T SC E T SC SC X T X T X T Grank G5 G5 G5 G4T3Q G4 G5 G5 G5T5 G5 G5 G4 G5 G5 G5 G5 G5T5 G5 G5 G4 G5 G5 G5 G5 G5 G4G5 G5 G5 G5 G4G5 G5 G5 G5 G5T1T2Q G5 G4G5 G4 G2 G4Q G5? G5 G4 G5 G5 G5 G5 G5 G4 G5 G5 A-7 Srank SU S1 S1 S3 S2 S2 S1 S2 S2 S2 SH S1 S1S2 S1 S2 S1S2 S3 S1 S3S4 S3 S3 S1 S? S1 S1 SNR S2 S2 S1S2 S2 SH S1 S1 S1 S2 S3 S2 S1 S2S3 S1 S2 S3 S3 SX S1 SX S2 SX S1 Appendix A - Rare terrestrial plant list - continued Species Silene stellata Silene virginica Silphium integrifolium Silphium laciniatum Silphium perfoliatum Sisyrinchium atlanticum Sisyrinchium farwellii Sisyrinchium hastile Sisyrinchium strictum Smilax herbacea Solidago bicolor Solidago houghtonii Solidago missouriensis Spiranthes ochroleuca Spiranthes ovalis Sporobolus clandestinus Sporobolus heterolepis Stellaria crassifolia Stellaria longipes Strophostyles helvula Tanacetum huronense Thalictrum venulosum var. confine Tipularia discolor Tofieldia pusilla Tomanthera auriculata Tradescantia bracteata Tradescantia virginiana Trichostema brachiatum Trichostema dichotomum Trillium nivale Trillium recurvatum Trillium sessile Trillium undulatum Trillium viride Triphora trianthophora Triplasis purpurea Trisetum spicatum Vaccinium cespitosum Vaccinium uliginosum Vaccinium vitis-idaea Valeriana edulis var. ciliata Valerianella chenopodiifolia Valerianella umbilicata Viburnum edule Viburnum prunifolium Viola epipsila Viola novae-angliae Viola pedatifida Vitis vulpina A-8 Common name Starry campion Fire pink Rosinweed Compass-plant Cup-plant Atlantic blue-eyed-grass Farwell's blue-eyed-grass Blue-eyed-grass Blue-eyed-grass Smooth carrion-flower White goldenrod Houghton's goldenrod Missouri goldenrod Yellow ladies'-tresses Lesser ladies'-tresses Dropseed Prairie dropseed Fleshy stitchwort Stitchwort Trailing wild bean Lake huron tansy Veiny meadow-rue Cranefly orchid False asphodel Eared false foxglove Long-bracted spiderwort Virginia spiderwort False pennyroyal Bastard pennyroyal Snow trillium Prairie trillium Toadshade Painted trillium Green trillium Three-birds orchid Sand grass Downy oat-grass Dwarf bilberry Alpine blueberry Mountain-cranberry Edible valerian Goosefoot corn-salad Corn-salad Squashberry or mooseberry Black haw Northern marsh violet New england violet Prairie birdfoot violet Frost grape Fed State T T T T T T X X SC SC SC LT T T SC T SC SC T SC SC T SC T T X X SC T T T T T E X T SC SC T T E T T T T SC T T T T Grank G5 G5 G5 G5 G5 G5 GHQ GUGHQ G2Q G5 G5 G3 G5 G4 G5? G5 G5 G5 G5 G5 G5T4T5 G5T4?Q G4G5 G5 G3 G5 G5 G5 G5 G4 G5 G4G5 G5 G4G5 G3G4 G4G5 G5 G5 G5 G5 G5T3 G5 G3G5 G5 G5 G4 G4Q G5 G5 Srank S2 S1 S2 S1S2 S2 S2 SX SX S2 S3 S3 S3 SNR S3 S1 S1 S3 S1S2 S2 S3 S3 S3 S1 S2 SX SX S2 S1 S2 S2 S2S3 S2S3 S1S2 SX S1 S2 S2S3 S1S2 S2 S1 S2 S1 S2 S2S3 S3 SH S2 S1 S1S2 Appendix A - Rare terrestrial plant list - continued Species Wisteria frutescens Woodsia alpina Woodsia obtusa Woodwardia areolata Zizania aquatica var. aquatica Zizia aptera Common name Wisteria Northern woodsia Blunt-lobed woodsia Netted chain-fern Wild-rice Prairie golden alexanders Fed State T T T X T T Grank G5 G4 G5 G5 G5T5 G5 Srank S1 S1 S1S2 SX S2S3 S1S2 A-9 Appendix B - Rare terrestrial animal list Scientic Name AMPHIBIANS Acris crepitans blanchardi Ambystoma opacum Ambystoma texanum Pseudacris triseriata maculata BIRDS Protonotaria citrea Rallus elegans Seiurus motacilla Spiza americana Sterna caspia Sterna forsteri Sterna hirundo Sturnella neglecta Pandion haliaetus Phalaropus tricolor Picoides arcticus Tympanuchus phasianellus Tyto alba Wilsonia citrina Xanthocephalus xanthocephalus Ammodramus henslowii Ammodramus savannarum Asio flammeus Accipiter cooperii Accipiter gentilis Botaurus lentiginosus Buteo lineatus Asio otus Chondestes grammacus Circus cyaneus Cistothorus palustris Charadrius melodus Chlidonias niger Cygnus buccinator Dendroica cerulea Dendroica discolor Dendroica dominica Dendroica kirtlandii Coturnicops noveboracensis Falcipennis canadensis Falco columbarius Falco peregrinus Gallinula chloropus Haliaeetus leucocephalus Ixobrychus exilis Lanius ludovicianus migrans A - 10 Common Name State Federal State Global rank rank status status Blanchard's Cricket Frog Marbled Salamander Smallmouth Salamander Boreal Chorus Frog SC T E SC G5T5 G5 G5 G5T5 S2S3 S1 S1 S1 Prothonotary Warbler King Rail Louisiana Waterthrush Dickcissel Caspian Tern Forster's Tern Common Tern Western Meadowlark Osprey Wilson's Phalarope Black-backed Woodpecker Sharp-tailed Grouse Barn Owl Hooded Warbler Yellow-headed Blackbird Henslow's Sparrow Grasshopper Sparrow Short-eared Owl Cooper's Hawk Northern Goshawk American Bittern Red-shouldered Hawk Long-eared Owl Lark Sparrow Northern Harrier Marsh Wren Piping Plover Black Tern Trumpeter Swan Cerulean Warbler Prairie Warbler Yellow-throated Warbler Kirtland's Warbler Yellow Rail Spruce Grouse Merlin Peregrine Falcon Common Moorhen Bald Eagle Least Bittern Migrant Loggerhead Shrike SC E SC SC T SC T SC T SC SC SC E SC SC T SC E SC SC SC T T X SC SC E SC T SC E T E T SC T E SC T T E G5 G4 G5 G5 G5 G5 G5 G5 G5 G5 G5 G4 G5 G5 G5 G4 G5 G5 G5 G5 G4 G5 G5 G5 G5 G5 G3 G4 G4 G4 G5 G5 G1 G4 G5 G5 G4 G5 G4 G5 G4T3Q S3 S1 S2S3 S3 S2 S2 S2 S4 S4 S2 S2 S3S4 S1 S3 S2 S2S3 S3S4 S1 S3S4 S3 S3S4 S3S4 S2 SX S3 S3S4 S1 S3 S3 S3 S1 S1 S1 S1S2 S2S3 S1S2 S1 S3 S4 S2 S1 LE LE Appendix B - Rare terrestrial animal list - continued Scientic Name Common Name Nycticorax nycticorax Beetles Nicrophorus americanus Dryobius sexnotatus Liodessus cantralli Lordithon niger Somatochlora hineana Somatochlora incurvata Butterflies and Moths Lycaeides idas nabokovi Lycaeides melissa samuelis Merolonche dolli Meropleon ambifusca Oarisma poweshiek Neonympha mitchellii mitchellii Erebia discoidalis Euphyes dukesi Euxoa aurulenta Fixsenia favonius ontario Euchloe ausonides Incisalia henrici Incisalia irus Hesperia ottoe Heterocampa subrotata Heteropacha rileyana Hemileuca maia Eacles imperialis pini Erora laeta Erynnis baptisiae Erynnis persius persius Chlosyne gorgone carlota Brachionycha borealis Atrytonopsis hianna Basilodes pepita Battus philenor Boloria freija Boloria frigga Calephelis mutica Catocala amestris Catocala dulciola Catocala illecta Catocala robinsoni Acronicta falcula Pachypolia atricornis Phyciodes batesii Oeneis macounii Oncocnemis piffardi Papaipema aweme Black-crowned Night-heron American Burying Beetle Six-banded Longhorn Beetle Cantrall's Bog Beetle Black Lordithon Rove Beetle Hine's Emerald Incurvate Emerald Northern Blue Karner Blue Doll's Merolonche Newman's Brocade Poweshiek Skipperling Mitchell's Satyr Red-disked Alpine Dukes' Skipper Dune Cutworm Northern Hairstreak Large Marble Henry's Elfin Frosted Elfin Ottoe Skipper Small Heterocampa Riley's Lappet Moth Barrens Buckmoth Pine Imperial Moth Early Hairstreak Wild Indigo Duskywing Persius Duskywing Gorgone Checkerspot Boreal Brachionyncha Dusted Skipper Gold Moth Pipevine Swallowtail Freija Fritillary Frigga Fritillary Swamp Metalmark Three-staff Underwing Quiet Underwing Magdalen Underwing Robinson's Underwing Corylus Dagger Moth Three-horned Moth Tawny Crescent Macoun's Arctic 3-striped Oncocnemis Aweme Borer State Federal State Global rank rank status status SC G5 S2S3 LE LE LE LE E SC SC SC E SC G2G3 GNR GNR GU G2G3 G4 SH SH S1S3 SH S1 S1S2 T T SC SC T E SC T SC SC SC SC T T SC SC SC SC SC SC T SC SC T SC SC SC SC SC E SC SC SC SC SC SC SC SC SC G5TU G5T2 G3G4 G3G4 G2G3 G1G2T1T2 G5 G3 G5 G4T4 G5 G5 G3 G3G4 G4G5 G4 G5 G5T3T4 G3G4 G5 G5T1T3 G5T5 G4 G4G5 G4 G5 G5 G5 G3 G4 G3 G5 G4 G2G4 G3G4 G4 G5 G4 GH S2 S2 S1S2 S1S2 S1S2 S1 S2S3 S1 S1S2 S1 S1S2 S2S3 S2S3 S1S2 S1S2 S1S2 S2S3 S2S3 S2S3 S2S3 S3 S2S3 S1S2 S2S3 S1S2 S1S2 S3S4 S3S4 S1S2 S1 S1S2 S2S3 S2S3 S2S3 S1S2 S4 S2S3 S1S2 SH A - 11 Appendix B - Rare terrestrial animal list - continued Scientic Name Common Name Papaipema beeriana Papaipema cerina Papaipema maritima Papaipema sciata Papaipema silphii Papaipema speciosissima Pyrgus wyandot Spartiniphaga inops Speyeria idalia Polygonia gracilis Proserpinus flavofasciata Schinia indiana Schinia lucens Pygarctia spraguei Cicadas and Hoppers Prosapia ignipectus Philaenarcys killa Dorydiella kansana Flexamia delongi Flexamia huroni Flexamia reflexus Lepyronia angulifera Lepyronia gibbosa Damselflies and Dragonflies Tachopteryx thoreyi Williamsonia fletcheri Grasshoppers and Crickets Trimerotropis huroniana Psinidia fenestralis Scudderia fasciata Orchelimum concinnum Orchelimum delicatum Orphulella pelidna Paroxya hoosieri Atlanticus davisi Appalachia arcana Melanoplus flavidus Oecanthus laricis Oecanthus pini Neoconocephalus lyristes Neoconocephalus retusus Mammals Myotis sodalis Microtus ochrogaster Microtus pinetorum Felis concolor Felis lynx Alces alces Blazing Star Borer Golden Borer Maritime Sunflower Borer Culvers Root Borer Silphium Borer Moth Regal Fern Borer Grizzled Skipper Spartina Moth Regal Fritillary Hoary Comma Yellow-banded Day-sphinx Phlox Moth Leadplant Flower Moth Sprague's Pygarctia A - 12 Federal State Global rank status status SC G3 SC G4 SC G3 SC G3G4 T G3G4 SC G4 SC G1G2Q SC G2G4 E G3 SC G5 SC G4 E G2G4 E G4 SC G5 State rank S1S2 S1S2 S1S2 S2S3 S1S2 S2S3 S1S2 S1S2 SH S3 S2S3 S1S2 S1 S2S3 Red-legged Spittlebug Spittlebug Leafhopper Leafhopper Huron River Leafhopper Leafhopper Angular Spittlebug Great Plains Spittlebug SC SC SC SC SC SC SC T G4 GNR GNR GNR GNR GNR G3 G3G4 S2S3 S1S2 S1S2 S1S2 S1 S1 S1S2 S1S2 Grey Petaltail Ebony Boghaunter SC SC G4 G3G4 S1S3 S1S2 Lake Huron Locust Atlantic-coast Locust Pine Katydid Red-faced Meadow Katydid Delicate Meadow Katydid Green Desert Grasshopper Hoosier Locust Davis's Shield-bearer Secretive Locust Blue-legged Locust Tamarack Tree Cricket Pinetree Cricket Bog Conehead Conehead Grasshopper T SC SC SC SC SC SC SC SC SC SC SC SC SC G2G3 G5 GNR GNR GNR G5 G5 GNR G2G3 G4 G1G2 GNR GNR GNR S2S3 S1S3 S1S3 S2S3 S1S3 S1S3 S2S3 S2S3 S2S3 S1S3 S1S2 S1S2 S1S3 S1 E E SC E E SC G2 G5 G5 G5 G5 G5 S1 S1 S3S4 SH S1 S4 Indiana Bat or Indiana Myotis Prairie Vole Woodland Vole Cougar Lynx Moose LE PS LT Appendix B - Rare terrestrial animal list - continued Scientic Name Common Name Canis lupus Cryptotis parva Pipistrellus subflavus Sorex fumeus Reptiles Nerodia erythrogaster neglecta Glyptemys insculpta Emydoidea blandingii Clonophis kirtlandii Cnemidophorus sexlineatus Clemmys guttata Pantherophis gloydi Pantherophis spiloides Sistrurus catenatus catenatus Terrapene carolina carolina Snails Pupilla muscorum Philomycus carolinianus Planogyra asteriscus Vertigo bollesiana Vertigo cristata Vertigo elatior Vertigo hubrichti Vertigo modesta Vertigo modesta parietalis Vertigo morsei Vertigo nylanderi Vertigo paradoxa Vertigo pygmaea Vallonia gracilicosta albula Xolotrema denotata Discus patulus Appalachina sayanus Anguispira kochi Catinella exile Gastrocopta holzingeri Guppya sterkii Hendersonia occulta Euconulus alderi Mesodon elevatus Mesomphix cupreus Gray Wolf Least Shrew Eastern Pipistrelle Smoky Shrew Copperbelly Watersnake Wood Turtle Blanding's Turtle Kirtland's Snake Six-lined Racerunner Spotted Turtle Eastern Fox Snake Black Rat Snake Eastern Massasauga Eastern Box Turtle Widespread Column Carolina Mantleslug Eastern Flat-whorl Delicate Vertigo Land Snail Tapered Vertigo Hubricht's Vertigo Cross Vertigo Land Snail Six-whorl Vertigo Deep-throat Vertigo Land Snail Crested Vertigo Land Snail Velvet Wedge Domed Disc Spike-lip Crater Banded Globe Land Snail Lambda Snaggletooth Snail Land Snail Cherrystone Drop Land Snail Proud Globe Copper Button Federal State Global rank status status LT T G4 T G5 SC G5 SC G5 State rank S3 S1S2 S2 S1 LT C E SC SC E SC T T SC SC SC G5T2T3 G4 G4 G2 G5 G5 G5T3 G5T5 G3G4T3T4 G5T5 S1 S2S3 S3 S1 SU S2 S2 S3 S3S4 S2S3 SC SC SC SC SC SC SC SC SC SC SC SC SC SC SC SC SC SC SC SC SC T SC SC SC G5 G5 G3G4 G3 G4 G5 G3 G5 G5T1 G2G3 G2 G3G4Q G5 G4Q G5 G5 G5 G5 G2 G5 G5Q G5 G3Q G5 G5 SU SU S3 S2 S3 S3 S2 S1 S1 S2 S1 S3 SU S1 SU SU SU SU SU S1 S1 S1 S2 SU SU A - 13 Appendix C - Rare aquatic plant list Species Armoracia lacustris Callitriche hermaphroditica Callitriche heterophylla Lemna valdiviana Myriophyllum alterniflorum Myriophyllum farwellii Nelumbo lutea Nuphar pumila Nymphaea tetragona Potamogeton bicupulatus Potamogeton confervoides Potamogeton hillii Potamogeton pulcher Potamogeton vaseyi Ruppia maritima Subularia aquatica Utricularia inflata Utricularia subulata Wolffia papulifera A - 14 Common name Lake cress Autumnal water-starwort Large water-starwort Pale duckweed Alternate-leaved water-milfoil Farwell's water-milfoil American lotus Small yellow pond-lily pygmy water-lily Waterthread pondweed Alga pondweed Hill's pondweed Spotted pondweed Vasey's pondweed Widgeon-grass Awlwort Floating bladderwort Zigzag bladderwort Water-meal Fed State T SC T X SC T T E E T SC T T T T E E T T Grank G4? G5 G5 G5 G5 G5 G4 G5T4T5 G5 G4 G4 G3 G5 G4 G5 G5 G5 G5 G4 Srank S2 S2 S1 SX S2S3 S2 S2 S1S2 S1 S2 S3 S2 S2 SH S1 S1 S1 S1 S1 Appendix D - Rare aquatic animal list Species BIRDS Gavia immer INSECTS Brychius hungerfordi Cordulegaster erronea Gomphus lineatifrons Gomphus quadricolor Ophiogomphus anomalus Ophiogomphus howei Stenelmis douglasensis Stylurus amnicola Stylurus laurae Stylurus notatus Stylurus plagiatus FISH Acipenser fulvescens Ammocrypta pellucida Clinostomus elongatus Coregonus artedi Coregonus hubbsi Coregonus johannae Coregonus kiyi Coregonus nigripinnis Coregonus reighardi Coregonus zenithicus Coregonus zenithicus bartletti Cottus ricei Erimyzon oblongus Etheostoma zonale Fundulus dispar Hiodon tergisus Hybopsis amblops Ictiobus niger Lepisosteus oculatus Macrhybopsis storeriana Moxostoma carinatum Notropis anogenus Notropis chalybaeus Notropis photogenis Notropis texanus Noturus miurus Noturus stigmosus Opsopoeodus emiliae Percina copelandi Percina shumardi Phoxinus erythrogaster Polyodon spathula Common Name Federal State Global State status Status Rank Rank Common loon Hungerford's crawling water beetle Tiger spiketail Splendid clubtail Rapids clubtail Extra-striped snaketail Pygmy snaketail Douglas stenelmis riffle beetle Riverine snaketail Laura's snaketail Elusive snaketail Russet-tipped clubtail Lake Sturgeon Eastern Sand Darter Redside Dace Cisco or Lake Herring Ives Lake Cisco Deepwater Cisco Kiyi Blackfin Cisco Shortnose Cisco Shortjaw Cisco Siskiwit Lake Cisco Spoonhead Sculpin Creek Chubsucker Banded Darter Starhead Topminnow Mooneye Bigeye Chub Black Buffalo Spotted Gar Silver Chub River Redhorse Pugnose Shiner Ironcolor Shiner Silver Shiner Weed Shiner Brindled Madtom Northern Madtom Pugnose Minnow Channel Darter River Darter Southern Redbelly Dace Paddlefish LE T G5 S3S4 E SC SC SC SC SC SC SC SC SC SC G1 G4 G4 G3G4 G3 G3 G1G3 G4 G4 G3 G5 S1 S1S2 S2S3 S2S3 S1 S1 S1S2 S1S2 S1S2 S1S2 S1S2 T T E T SC X SC X X T SC SC E SC SC T X SC SC SC T SC X E X SC E E E E E X G3G4 G3 G4 G5 G1Q GX G3 GXQ GH G3 GHQ G5 G5 G5 G4 G5 G5 G5 G5 G5 G4 G3 G4 G5 G5 G5 G3 G5 G4 G5 G5 G4 S2 S1S2 S1S2 S3 S1 SX S3 SX SH S2 S1 S3 S1S2 S1 S2 S2 SH S3 S2S3 S2S3 S1 S3 S1 S1 S1 S2S3 S1 S1 S1S2 S1 S1 SX A - 15 Appendix D - Rare aquatic animal list continued Species Common Name Sander canadensis Stizostedion vitreum glaucum Thymallus arcticus MUSSELS Alasmidonta marginata Alasmidonta viridis Anodonta subgibbosa Cyclonaias tuberculata Dysnomia sulcata Epioblasma obliquata perobliqua Epioblasma torulosa rangiana Epioblasma triquetra Lampsilis fasciola Leptodea leptodon Obovaria olivaria Obovaria subrotunda Pleurobema clava Pleurobema coccineum Simpsonaias ambigua Toxolasma lividus Venustaconcha ellipsiformis Villosa fabalis Villosa iris SNAILS Acella haldemani Fontigens nickliniana Planorbella multivolvis Planorbella smithi Pomatiopsis cincinnatiensis Pyrgulopsis letsoni Stagnicola contracta Stagnicola petoskeyensis Sauger Bluepike Arctic Grayling A - 16 Elktoe Slippershell Mussel Lake Floater Purple Wartyback Catspaw White Catspaw Northern Riffleshell Snuffbox Wavy-rayed Lampmussel Scaleshell Hickorynut Round Hickorynut Clubshell Round Pigtoe Salamander Mussel Purple Lilliput Ellipse Rayed Bean Rainbow Spindle Lymnaea Watercress Snail Acorn Ramshorn Aquatic Snail Brown Walker Gravel Pyrg Deepwater Pondsnail Petoskey Pondsnail Federal State status Status T X X LE LE LE LE LE C Global Rank G5 G5TX G5 State Rank S1 SX SX SC SC T SC E E E E T SC SC E E SC E E SC E SC S2S3 S2S3 S1 S2S3 SH SH S1 S1 S2 SU S2S3 S1 S1 S2S3 S1 S1 S2S3 S1 S2S3 G4 G4G5 G1Q G5 G1 G1T1 G2T2 G3 G4 G1 G4 G4 G2 G4 G3 G2 G3G4 G1G2 G5 SC SC E SC SC SC T E S3 SU SX S2 SU SU S1 SH G3 G5 GX G2 G4 G5 G1 GH Appendix E - Global and State rank descriptions GLOBAL RANKS G1 = critically imperiled globally because of extreme rarity (5 or fewer occurrences rangewide or very few remaining individuals or acres) or because of some factor(s) making it especially vulnerable to extinction. G2 = imperiled globally because of rarity (6 to 20 occurrences or few remaining individuals or acres) or because of some factor(s) making it very vulnerable to extinction throughout its range. G3 = either very rare and local throughout its range or found locally (even abundantly at some of its locations) in a restricted range (e.g. a single western state, a physiographic region in the East) or because of other factor(s) making it vulnerable to extinction throughout its range; in terms of occurrences, in the range of 21 to 100. G4 = apparently secure globally, though it may be quite rare in parts of its range, especially at the periphery. G5 = demonstrably secure globally, though it may be quite rare in parts of its range, especially at the periphery. GH = of historical occurrence throughout its range, i.e. formerly part of the established biota, with the expectation that it may be rediscovered (e.g. Bachman’s Warbler). GU = possibly in peril range-wide, but status uncertain; need more information. GX = believed to be extinct throughout its range (e.g. Passenger Pigeon) with virtually no likelihood that it will be rediscovered. STATE RANKS S1 = critically imperiled in the state because of extreme rarity (5 or fewer occurrences or very few remaining individuals or acres) or because of some factor(s) making it especially vulnerable to extirpation in the state. S2 = imperiled in state because of rarity (6 to 20 occurrences or few remaining individuals or acres) or because of some factor(s) making it very vulnerable to extirpation from the state. S3 = rare or uncommon in state (on the order of 21 to 100 occurrences). S4 = apparently secure in state, with many occurrences. S5 = demonstrably secure in state and essentially ineradicable under present conditions. SA = accidental in state, including species (usually birds or butterflies) recorded once or twice or only at very great intervals, hundreds or even thousands of miles outside their usual range. SE = an exotic established in the state; may be native elsewhere in North America (e.g. house finch or catalpa in eastern states). SH = of historical occurrence in state and suspected to be still extant. SN = regularly occurring, usually migratory and typically nonbreeding species. SR = reported from state, but without persuasive documentation which would provide a basis for either accepting or rejecting the report. SRF = reported falsely (in error) from state but this error persisting in the literature. SU = possibly in peril in state, but status uncertain; need more information. SX = apparently extirpated from state. A - 17 Appendix F - MNFI Natural Community List (Names in italics represent categories that are not currently tracked as separate natural communities) Community Name Alvar [Alvar grassland] Bedrock glade Basalt bedrock glade Igneous bedrock glade Limestone bedrock glade [Alvar glade] Sandstone bedrock glade Volcanic conglomerate bedrock glade Bedrock lakeshore Basalt bedrock lakeshore Igneous bedrock lakeshore Limestone pavement lakeshore [Alvar pavement] Volcanic conglomerate bedrock lakeshore Bog Boreal forest Bur oak plains Cave Cliff Dry acid cliff Dry non-acid cliff Moist acid cliff Moist non-acid cliff Coastal plain marsh Cobble beach [Cobble shore] Dry northern forest [Pine forest] Dry sand prairie Dry southern forest [Oak forest] Dry-mesic northern forest [Pine-hardwood forest] Dry-mesic southern forest [Oak-hardwood forest] Emergent marsh Great Lakes barrens Great Lakes marsh Hardwood-conifer swamp Hillside prairie Inland salt marsh Interdunal wetland Intermittent wetland [Boggy seepage wetland] Inundated shrub swamp Lakeplain mesic sand prairie Lakeplain oak openings Lakeplain wet prairie Lakeplain wet-mesic prairie Lakeshore cliff Basalt lakeshore cliff Sandstone lakeshore cliff Volcanic conglomerate lakeshore cliff Mesic northern forest [Northern hardwood forest; Hemlock-hardwood forest] Mesic prairie Mesic sand prairie Mesic southern forest [Southern hardwood forest] Muskeg A - 18 State Rank Global Rank S1 G2 S2 S2 S2 S2? S2 G3 G3G4 G2? G3G4 G3 S2 S2 S2 S2 S4 S3 SX S1 G4G5 G? G? G4G5 G5 G4G5 G1 G4? S2? S2 S2? S2 S2 S3 S3 S2 S3 S3 S3 S4 S2 S3 S3 S1 S1 S2 S3 S3 S1 S1 S2 S2 G4G5 G4G5 G4G5 G4G5 G2? G4G5 G4 G2G3 G4? G4? G4? G5 G2 G4 G3G4 G3 G1 G3? G3 G4 G1 G1 G2G3 G2 S1 S2 S1 S3 S1 S1 S3 S3 G3? G3? G3? G4 G1G2 G2 G3G4 G4G5 Appendix F - MNFI Natural Community List - Continued Northern bald [Krummholz ridgetop] Northern fen Northern shrub thicket Northern swamp Northern wet meadow Northern wet-mesic prairie Oak barrens Oak openings Oak-pine barrens Open dunes Patterned fen Pine barrens Poor conifer swamp Poor fen Prairie fen Relict conifer swamp Rich conifer swamp Sand/gravel beach Sinkhole Southern floodplain forest Southern shrub-carr Southern swamp Southern wet meadow Submergent marsh Wet prairie Wet-mesic prairie Wooded dune and swale complex Woodland prairie S1 S3 S5 S3? S4 S1 S2 S1 S2 S3 S2 S2 S4 S3 S3 S3 S3 S3 S2 S3 S5 S3 S3 S4 S2 S2 S3 S2 G3G4 G4G5 G5? G4? G4G5 G? G3 G1 G2? G3G5 G3G4 G2 G5 G3G4 G3G4 G3 G4 G3? G3G5 G3G5 G5 G4? G4? G5 G3? G2G3 G3 G3 A - 19 Appendix G - Description of Ecological Drainage Units There are nine Ecological Drainage Units in Michigan, we combined them into 7 EDUs. The following paragraphs briefly describe each one in terms of climate, within ecoregion sections and subsections, major landforms, water features, and zoogeography. (16) Southeast Michigan Interlobate and Lake Plain (SEMILP) contains most of the Lake Erie drainage in Michigan. Mean annual temperature is 48.6ˆF (sd 1.1) and has a mean annual precipitation of 30.5 inches (sd 4.8). This EDU contains many kettle lakes, ponds, and wetland complexes in the interlobate headwaters region. In the lake and till plains, there are few lakes but many low gradient streams. Historically, all streams flow to the Ohio River via the Teays River but today they all flow into western Lake Erie and Lake St. Clair. (2) Only a small portion of the Western Lake Erie (WLE) EDU is in Michigan, most of the EDU is in Ohio. The mean annual temperature in this EDU is 48.6-50.1ˆF (sd 1.0-1.2) and the mean annual precipitation is between 30.5-34.3 (sd 4.6-4.8) inches. This EDU mainly has low gradient, surface water-fed streams except in the interlobate area (along the glacial boundary) where moderate gradient streams occur. Historically, all streams drained to the Ohio River via Teays River but today they all flow into western Lake Erie. Because only a small area of this EDU is in Michigan, it will be combined with the SEMILP EDU for this analysis. (4) The Saginaw Bay (SB) EDU if found in the lower half of the Huron River Basin. The mean annual temperature is 48.5 to 43.3 (sd 1.08) ˆF and the mean annual precipitation is 29.2 (sd 3.8) to 31.7 (sd 4.56) inches from south to north respectively. Many of the streams in this EDU are intermittent. Those that are perennial are part of the Saginaw River system and are generally low gradient streams. Historically, all streams drained west out to the Grand River into Lake Chicago but today they drain to Saginaw Bay and Lake St. Clair. (3) The Southeast Lake Michigan (SELM) EDU is the southern portion of the Lake Michigan basin. Mean annual air temperatures range from 48.6 (sd 1.15) to 47.4 (sd 1.11) ˆF and mean annual precipitation is 35.1 (sd 4.9) to 31.7 (sd 4.56) inches with the rain shadow from west to east. This EDU has three major river systems (Grand, Kalamazoo, and St. Joseph) which flow east to west. There are many kettle lakes in the interlobate region to the east, which forms the headwaters of all three river systems. Historically, all waters in this region drained west out the Grand River into Lake Chicago, today all rivers flow west to southern Lake Michigan. (5) The Northern Lake Michigan, Lake Huron, and Straits of Mackinac (NLMLHSM) EDU encompasses the northern half of the lower peninsula of Michigan. Mean annual air temperatures range from 46.1 (sd 1.16) to 43.3 (sd 1.08) ˆF from west to east and mean annual precipitation ranges from 33.1 (sd 4.38) to 29.5 (sd 3.29) inches from west to each with a rain shadow from southwest to northwest. There are kettle lakes in the outwash plains areas. In the lake plain area there are some large lakes, lakes of many geneses, and intermittent streams. Groundwater streams can be found in the outwash surrounded by coarse moraines and ice contact. Historically, this area likely drained to the St. Lawrence River via the Ottowa River and Champlain Sea but today, rivers drain west to Lake Michigan, east to Lake Huron, and north to the straits. The Lake Michigan and Lake Huron drainage divide roughly bisects this EDU. A - 20 Appendix G - Description of Ecological Drainage Units - Continued (7) In the Eastern Upper Peninsula (EUP) EDU the mean annual temperature is 41.1 (sd 1.06) ˆF and the mean annual precipitation is 32.5 (sd 4.07) inches. This EDU has many small and medium sized low-gradient streams which are underlain by deep sandy outwash deposits or sedimentary rock. They are also often connected to wetlands. Historically, the streams in this area likely drained to the St. Lawrence River via the Ottowa River and Champlain Sea, but today waters drain to the north to Lake Superior and to the south to Lakes Michigan and Huron and to the St. Mary’s River. (8) In the Central Upper Peninsula (CUP) EDU the mean annual temperature is 40.4 (sd 1.22) ˆF and the mean annual precipitation is 32.5 (sd 4.39) inches. Half of this EDU is within the Menominee River drainage. There are many lakes, spring ponds, springs, wetlands, and streams in this EDU. Kettle lakes are common. Streams tend to be low in density and have dendritic drainages and high spring and fall water flows with relatively low flows in the summer. These low gradient streams are underlain by sandy outwash, limestone, or shale. Historically, the waters in this EDU drained south to the Mississippi River via a connection through Green Bay (Wolf/Fox Rivers), but today it drains north to Lake Superior and south to northern Lake Michigan / Green Bay. (6) The Western Upper Peninsula and Keweenaw Peninsula (WUPKP) EDU has mean annual air temperatures of 40.42 (sd 1.22) ˆF and a mean annual precipitation of 32.5 (sd 4.39) inches. This EDU has many kettle lakes in the outwash plains. Historically, the waters in this EDU drained to the upper Mississippi River via St. Croix River drainage of glacial Lake Duluth with a possible connection to Hudson Bay and Lake Agassiz. Today the waters drain to the southwest into Lake Superior. (12) A very small portion of Michigan is in the Bayfield Peninsula and Uplands (BPU) EDU. The mean annual temperature in this EDU is 41.41 (sd 1.16) ˆF and the mean annual precipitation is 31.29 (sd 5.39) inches, this precipitation. There are few lakes in this EDU and the streams are low gradient and flow from west to east into Lake Michigan. Historically, this EDU drained to the Mississippi River via the Fox River, but today it drains to western Lake Michigan. Only a very small portion of this EDU is in Michigan, hence we will combine it with the WUPKP EDU during our analysis because it is in the same ecoregion. A - 21 Appendix H - Natural Vegetation Type datalayers and descriptions File Name allforgps allfor_min allfor_90 allfor_210 allfor_300 allforjgps allforj_min allforj_90 allforj_210 allforj_300 allformgps allform_min allform_90 allform_210 allform_300 upfgps upf_min upf_90 upf_210 upf_300 upfjgps upfj_min upfj_90 upfj_210 upfj_300 upfmgps upfm_min upfm_90 upfm_210 upfm_300 updecgps updec_min updec_90 updec_210 updec_300 updecjgps updecj_min updecj_90 updecj_210 updecj_300 updecmgps updecm_min updecm_90 n/a n/a upmgps A - 22 Vegetation Type All Forest All Forest All Forest All Forest All Forest All Forest All Forest All Forest All Forest All Forest All Forest All Forest All Forest All Forest All Forest Upland Forest Upland Forest Upland Forest Upland Forest Upland Forest Upland Forest Upland Forest Upland Forest Upland Forest Upland Forest Upland Forest Upland Forest Upland Forest Upland Forest Upland Forest Upland decidious forest Upland decidious forest Upland decidious forest Upland decidious forest Upland decidious forest Upland decidious forest Upland decidious forest Upland decidious forest Upland decidious forest Upland decidious forest Upland decidious forest Upland decidious forest Upland decidious forest Upland decidious forest Upland decidious forest Upland mixed forest road layer none none none none none major major major major major all all all all all none none none none none major major major major major all all all all all none none none none none major major major major major all all all all all none minimum size buffer patches size in (acres) meters 20 0 5000 0 5000 90 5000 210 5000 300 20 0 5000 0 5000 90 5000 210 5000 300 20 0 5000 0 5000 90 5000 210 5000 300 20 0 5000 0 5000 90 5000 210 5000 300 20 0 5000 0 5000 90 5000 210 5000 300 20 0 5000 0 5000 90 5000 210 5000 300 20 0 5000 0 5000 90 5000 210 5000 300 20 0 5000 0 5000 90 5000 210 5000 300 20 0 5000 0 5000 90 5000 210 5000 300 20 0 Appendix H - Natural Vegetation Type datalayers and descriptions - continued File Name upmix_min n/a n/a n/a upmjgps upmixj_min n/a n/a n/a upmmgps upmixm_min n/a n/a n/a upcongps upcon_min upcon_90 upcon_210 upcon_300 upconjgps upconj_min upconj_90 upconj_210 upconj_300 upconmgps upconm_min upconm_90 upconm_210 upconm_300 wetforgps wetfor_min wetfor_90 wetfor_210 wetfor_300 wetforjgps wetforj_min wetforj_90 wetforj_210 wetforj_300 wetformgps wetform_min wetform_90 wetform_210 wetform_300 lowdecgps lowdec_min Vegetation Type Upland mixed forest Upland mixed forest Upland mixed forest Upland mixed forest Upland mixed forest Upland mixed forest Upland mixed forest Upland mixed forest Upland mixed forest Upland mixed forest Upland mixed forest Upland mixed forest Upland mixed forest Upland mixed forest Upland coniferous forest Upland coniferous forest Upland coniferous forest Upland coniferous forest Upland coniferous forest Upland coniferous forest Upland coniferous forest Upland coniferous forest Upland coniferous forest Upland coniferous forest Upland coniferous forest Upland coniferous forest Upland coniferous forest Upland coniferous forest Upland coniferous forest Lowland forest Lowland forest Lowland forest Lowland forest Lowland forest Lowland forest Lowland forest Lowland forest Lowland forest Lowland forest Lowland forest Lowland forest Lowland forest Lowland forest Lowland forest Lowland deciduous forest Lowland deciduous forest road layer none none none none major major major major major all all all all all none none none none none major major major major major major all all all all none none none none none major major major major major all all all all all none none minimum size buffer patches size in (acres) meters 5000 0 5000 90 5000 210 5000 300 20 0 5000 0 5000 90 5000 210 5000 300 20 0 5000 0 5000 90 5000 210 5000 300 20 0 50 0 50 90 50 210 50 300 20 0 50 0 50 90 50 210 50 300 20 0 50 0 50 90 50 210 50 300 20 0 50 0 50 90 50 210 50 300 20 0 50 0 50 90 50 210 50 300 20 0 50 0 50 90 50 210 50 300 20 0 50 0 A - 23 Appendix H - Natural Vegetation Type datalayers and descriptions - continued File Name lowdec_90 lowdec_210 lowdec_300 lowdecgps lowdecj_min lowdecj_90 lowdecj_210 lowdecj_300 lowdecmgps lowdecm_min lowdecm_90 lowdecm_210 lowdecm_300 lowmixgps lowmix_min lowmix_90 n/a n/a lowmixjgps lowmixj_min lowmixj_90 n/a n/a lowmixmgps lowmixm_min lowmixm_90 n/a n/a lowcongps lowcon_min lowcon_90 lowcon_210 lowcon_300 lowconjgps lowconj_min lowconj_90 lowconj_210 lowconj_300 lowconmgps lowconm_min lowconm_90 lowconm_210 lowconm_300 grassgps grass_min grass_90 A - 24 Vegetation Type Lowland deciduous forest Lowland deciduous forest Lowland deciduous forest Lowland deciduous forest Lowland deciduous forest Lowland deciduous forest Lowland deciduous forest Lowland deciduous forest Lowland deciduous forest Lowland deciduous forest Lowland deciduous forest Lowland deciduous forest Lowland deciduous forest Lowland mixed forest Lowland mixed forest Lowland mixed forest Lowland mixed forest Lowland mixed forest Lowland mixed forest Lowland mixed forest Lowland mixed forest Lowland mixed forest Lowland mixed forest Lowland mixed forest Lowland mixed forest Lowland mixed forest Lowland mixed forest Lowland mixed forest Lowland coniferous forest Lowland coniferous forest Lowland coniferous forest Lowland coniferous forest Lowland coniferous forest Lowland coniferous forest Lowland coniferous forest Lowland coniferous forest Lowland coniferous forest Lowland coniferous forest Lowland coniferous forest Lowland coniferous forest Lowland coniferous forest Lowland coniferous forest Lowland coniferous forest Filtered grassland Filtered grassland Filtered grassland road layer none none none major major major major major all all all all all none none none none none major major major major major all all all all all none none none none none none major major major major all all all all all none none none minimum size buffer patches size in (acres) meters 50 90 50 210 50 300 20 0 50 0 50 90 50 210 50 300 20 0 50 0 50 90 50 210 50 300 20 0 50 0 50 90 50 210 50 300 20 0 50 0 50 90 50 210 50 300 20 0 50 0 50 90 50 210 50 300 20 0 50 0 50 90 50 210 50 300 20 0 50 0 50 90 50 210 50 300 20 0 50 0 50 90 50 210 50 300 20 0 50 0 50 90 Appendix H - Natural Vegetation Type datalayers and descriptions - continued File Name grass_210 grass_300 grassjgps grassj_min grassj_90 grassj_210 grassj_300 grassmgps grassm_min grassm_90 grassm_210 grassm_300 nonforgps nonfor_90 nonfor_210 nonfor_300 nonforjgps nonforj_90 nonforj_210 nonforj_300 nonformgps nonform_90 nonform_210 nonform_300 Vegetation Type Filtered grassland Filtered grassland Filtered grassland Filtered grassland Filtered grassland Filtered grassland Filtered grassland Filtered grassland Filtered grassland Filtered grassland Filtered grassland Filtered grassland Non-forested wetland Non-forested wetland Non-forested wetland Non-forested wetland Non-forested wetland Non-forested wetland Non-forested wetland Non-forested wetland Non-forested wetland Non-forested wetland Non-forested wetland Non-forested wetland road layer none none major major major major major all all all all all none none none none major major major major all all all all minimum size buffer patches size in (acres) meters 50 210 50 300 20 0 50 0 50 90 50 210 50 300 20 0 50 0 50 90 50 210 50 300 0.1 0 0.1 90 0.1 210 0.1 300 0.1 0 0.1 90 0.1 210 0.1 300 0.1 0 0.1 90 0.1 210 0.1 300 A - 25 Appendix H - Natural Vegetation Type datalayers and descriptions - continued All forests Allforgps Patches of all forest types. All forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted and grouped together into patches. Allforjgps Patches of all forest types, cut by major roads. All forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. Major roads are converted to raster and the road raster removed from the forest land cover types. The remaining forest cover type cells are then grouped together into patches. Allformgps Patches of all forest types, cut by all roads. All forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. All roads are converted to raster and the road raster removed from the forest land cover types. The remaining forest cover type cells are then grouped together into patches. Allfor_min Patches of all forest types, greater than 2000 hectares. All forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted and grouped together into patches. Patches greater than or equal to 2000 hectares are extracted. Allforj_min Patches of all forest types, cut by major roads, greater than 2000 hectares. All forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. Major roads are converted to raster and the road raster removed from the forest land cover types. The remaining forest cover type cells are then grouped together into patches. Patches greater than or equal to 2000 hectares are extracted. Allform_min Patches of all forest types, cut by all roads, greater than 2000 hectares. All forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. All roads are converted to raster and the road raster removed from the forest land cover types. The remaining forest cover type A - 26 Appendix H - Natural Vegetation Type datalayers and descriptions - continued cells are then grouped together into patches. Patches greater than or equal to 2000 hectares are extracted. Allfor_90 Patches of all forest types, greater than 2000 hectares, after buffering inward 90 meters. All forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted and grouped together into patches. Patches greater than or equal to 2000 hectares are extracted. Patches are shrunk by 3 cells (90 meters), regrouped into patches, and patches greater than or equal to 2000 hectares extracted. Allfor_210 Patches of all forest types, greater than 2000 hectares, after buffering inward 210 meters. All forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted and grouped together into patches. Patches greater than or equal to 2000 hectares are extracted. Patches are shrunk by 7 cells (210 meters), regrouped into patches, and patches greater than or equal to 2000 hectares extracted. Allfor_300 Patches of all forest types, greater than 2000 hectares, after buffering inward 300 meters. All forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted and grouped together into patches. Patches greater than or equal to 2000 hectares are extracted. Patches are shrunk by 10 cells (300 meters), regrouped into patches, and patches greater than or equal to 2000 hectares extracted. Allforj_90 Patches of all forest types, cut by major roads, greater than 2000 hectares, after buffering inward 90 meters. All forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. Major roads are converted to raster and the road raster removed from the forest land cover types. The remaining forest cover type cells are then grouped together into patches. Patches greater than or equal to 2000 hectares are extracted. Patches are shrunk by 3 cells (90 meters), regrouped into patches, and patches greater than or equal to 2000 hectares extracted. Allforj_210 Patches of all forest types, cut by major roads, greater than 2000 hectares, after buffering inward 210 meters. All forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. Major roads are converted to raster and the road raster removed from the forest land cover types. The remaining forest cover type cells are then grouped together into patches. Patches greater than or equal to 2000 A - 27 Appendix H - Natural Vegetation Type datalayers and descriptions - continued hectares are extracted. Patches are shrunk by 7 cells (210 meters), regrouped into patches, and patches greater than or equal to 2000 hectares extracted. Allforj_300 Patches of all forest types, cut by major roads, greater than 2000 hectares, after buffering inward 300 meters. All forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. Major roads are converted to raster and the road raster removed from the forest land cover types. The remaining forest cover type cells are then grouped together into patches. Patches greater than or equal to 2000 hectares are extracted. Patches are shrunk by 10 cells (300 meters), regrouped into patches, and patches greater than or equal to 2000 hectares extracted. Allform_90 Patches of all forest types, cut by all roads, greater than 2000 hectares, after buffering inward 90 meters. All forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. All roads are converted to raster and the road raster removed from the forest land cover types. The remaining forest cover type cells are then grouped together into patches. Patches greater than or equal to 2000 hectares are extracted. Patches are shrunk by 3 cells (90 meters), regrouped into patches, and patches greater than or equal to 2000 hectares extracted. Allform_210 Patches of all forest types, cut by all roads, greater than 2000 hectares, after buffering inward 210 meters. All forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. All roads are converted to raster and the road raster removed from the forest land cover types. The remaining forest cover type cells are then grouped together into patches. Patches greater than or equal to 2000 hectares are extracted. Patches are shrunk by 7 cells (210 meters), regrouped into patches, and patches greater than or equal to 2000 hectares extracted. Allform_300 Patches of all forest types, cut by all roads, greater than 2000 hectares, after buffering inward 300 meters. All forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. All roads are converted to raster and the road raster removed from the forest land cover types. The remaining forest cover type cells are then grouped together into patches. Patches greater than or equal to 2000 hectares are extracted. Patches are shrunk by 10 cells (300 meters), regrouped into patches, and patches greater than or equal to 2000 hectares extracted. A - 28 Appendix H - Natural Vegetation Type datalayers and descriptions - continued Lowland coniferous forests Lcgps Patches of lowland coniferous forest types. Lowland coniferous forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted and grouped together into patches. Lcjgps Patches of lowland coniferous forest types, cut by major roads. Lowland coniferous forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. Major roads are converted to raster and the road raster removed from the forest land cover types. The remaining lowland coniferous forest cover type cells are then grouped together into patches. Lcmgps Patches of lowland coniferous forest types, cut all roads. Lowland coniferous forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. All roads are converted to raster and the road raster removed from the forest land cover types. The remaining lowland coniferous forest cover type cells are then grouped together into patches. Lowcon_min Patches of lowland coniferous forest types, greater than 50 hectares. Lowland coniferous forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted and grouped together into patches. Patches greater than or equal to 50 hectares are extracted. Lowconj _min Patches of lowland coniferous forest types, cut by major roads, greater than 50 hectares. Lowland coniferous forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. Major roads are converted to raster and the road raster removed from the lowland coniferous forest land cover types. The remaining lowland coniferous forest cover type cells are then grouped together into patches. Patches greater than or equal to 50 hectares are extracted. Lowconm _min Patches of lowland coniferous forest types, cut by all roads, greater than 50 hectares. Lowland coniferous forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. All roads are A - 29 Appendix H - Natural Vegetation Type datalayers and descriptions - continued converted to raster and the road raster removed from the lowland coniferous forest land cover types. The remaining lowland coniferous forest cover type cells are then grouped together into patches. Patches greater than or equal to 50 hectares are extracted. Lowcon _90 Patches of lowland coniferous forest types, greater than 50 hectares, after buffering inward 90 meters. Lowland coniferous forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted and grouped together into patches. Patches greater than or equal to 50 hectares are extracted. Patches are shrunk by 3 cells (90 meters), regrouped into patches, and patches greater than or equal to 50 hectares extracted. Lowcon _210 Patches of lowland coniferous forest types, greater than 50 hectares, after buffering inward 210 meters. Lowland coniferous forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted and grouped together into patches. Patches greater than or equal to 50 hectares are extracted. Patches are shrunk by 7 cells (210 meters), regrouped into patches, and patches greater than or equal to 50 hectares extracted. Lowcon _300 Patches of lowland coniferous forest types, greater than 50 hectares, after buffering inward 300 meters. Lowland coniferous forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted and grouped together into patches. Patches greater than or equal to 50 hectares are extracted. Patches are shrunk by 10 cells (300 meters), regrouped into patches, and patches greater than or equal to 50 hectares extracted. Lowconj_90 Patches of lowland coniferous forest types, cut by major roads, greater than 50 hectares, after buffering inward 90 meters. Lowland coniferous forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. Major roads are converted to raster and the road raster removed from the lowland coniferous forest land cover types. The remaining lowland coniferous forest cover type cells are then grouped together into patches. Patches greater than or equal to 50 hectares are extracted. Patches are shrunk by 3 cells (90 meters), regrouped into patches, and patches greater than or equal to 50 hectares extracted. A - 30 Appendix H - Natural Vegetation Type datalayers and descriptions - continued Lowconj_210 Patches of lowland coniferous forest types, cut by major roads, greater than 50 hectares, after buffering inward 210 meters. Lowland coniferous forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. Major roads are converted to raster and the road raster removed from the lowland coniferous forest land cover types. The remaining lowland coniferous forest cover type cells are then grouped together into patches. Patches greater than or equal to 50 hectares are extracted. Patches are shrunk by 7 cells (210 meters), regrouped into patches, and patches greater than or equal to 50 hectares extracted. Lowconj_300 Patches of lowland coniferous forest types, cut by major roads, greater than 50 hectares, after buffering inward 300 meters. Lowland coniferous forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. Major roads are converted to raster and the road raster removed from the lowland coniferous forest land cover types. The remaining lowland coniferous forest cover type cells are then grouped together into patches. Patches greater than or equal to 50 hectares are extracted. Patches are shrunk by 10 cells (300 meters), regrouped into patches, and patches greater than or equal to 50 hectares extracted. Lowconm_90 Patches of lowland coniferous forest types, cut by all roads, greater than 50 hectares, after buffering inward 90 meters. Lowland coniferous forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. All roads are converted to raster and the road raster removed from the lowland coniferous forest land cover types. The remaining lowland coniferous forest cover type cells are then grouped together into patches. Patches greater than or equal to 50 hectares are extracted. Patches are shrunk by 3 cells (90 meters), regrouped into patches, and patches greater than or equal to 50 hectares extracted. Lowconm_210 Patches of lowland coniferous forest types, cut by all roads, greater than 50 hectares, after buffering inward 210 meters. Lowland coniferous forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. All roads are converted to raster and the road raster removed from the lowland coniferous forest land cover types. The remaining lowland coniferous forest cover type cells are then grouped together into patches. Patches greater than or equal to 50 hectares are extracted. Patches are shrunk by 7 cells (210 meters), regrouped into patches, and patches greater than or equal to 50 hectares extracted. Lowconm_300 Patches of lowland coniferous forest types, cut by all roads, greater than 50 hectares, after buffering inward 300 meters. A - 31 Appendix H - Natural Vegetation Type datalayers and descriptions - continued Lowland coniferous forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. All roads are converted to raster and the road raster removed from the lowland coniferous forest land cover types. The remaining lowland coniferous forest cover type cells are then grouped together into patches. Patches greater than or equal to 50 hectares are extracted. Patches are shrunk by 10 cells (300 meters), regrouped into patches, and patches greater than or equal to 50 hectares extracted. Lowland deciduous Ldgps Patches of lowland deciduous forest types. Lowland deciduous forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted and grouped together into patches. Ldjgps Patches of lowland deciduous forest types, cut by major roads. Lowland deciduous forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. Major roads are converted to raster and the road raster removed from the lowland forest land cover types. The remaining lowland coniferous forest cover type cells are then grouped together into patches. Ldmgps Patches of lowland deciduous forest types, cut by all roads. Lowland deciduous forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. All roads are converted to raster and the road raster removed from the lowland forest land cover types. The remaining lowland deciduous forest cover type cells are then grouped together into patches. Lowdec_min Patches of lowland deciduous forest types, greater than 50 hectares. Lowland deciduous forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted and grouped together into patches. Patches greater than or equal to 50 hectares are extracted. Lowdecj _min Patches of lowland deciduous forest types, cut by major roads, greater than 50 hectares. Lowland deciduous forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. Major roads are converted to raster and the road raster removed from the lowland deciduous forest land cover types. The remaining lowland deciduous forest cover type cells are then grouped together into patches. Patches greater than or equal to 50 hectares are extracted. A - 32 Appendix H - Natural Vegetation Type datalayers and descriptions - continued Lowdecm _min Patches of lowland deciduous forest types, cut by all roads, greater than 50 hectares. Lowland deciduous forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. All roads are converted to raster and the road raster removed from the lowland deciduous forest land cover types. The remaining lowland deciduous forest cover type cells are then grouped together into patches. Patches greater than or equal to 50 hectares are extracted. Lowdec _90 Patches of lowland deciduous forest types, greater than 50 hectares, after buffering inward 90 meters. Lowland deciduous forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted and grouped together into patches. Patches greater than or equal to 2000 hectares are extracted. Patches are shrunk by 3 cells (90 meters), regrouped into patches, and patches greater than or equal to 50 hectares extracted. Lowdecj_90 Patches of lowland deciduous forest types, cut by major roads, greater than 50 hectares, after buffering inward 90 meters. Lowland deciduous forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. Major roads are converted to raster and the road raster removed from the lowland deciduous forest land cover types. The remaining lowland deciduous forest cover type cells are then grouped together into patches. Patches greater than or equal to 50 hectares are extracted. Patches are shrunk by 3 cells (90 meters), regrouped into patches, and patches greater than or equal to 50 hectares extracted. Lowdecm_90 Patches of lowland deciduous forest types, cut by all roads, greater than 50 hectares, after buffering inward 90 meters. Lowland coniferous forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. All roads are converted to raster and the road raster removed from the lowland deciduous forest land cover types. The remaining lowland deciduous forest cover type cells are then grouped together into patches. Patches greater than or equal to 50 hectares are extracted. Patches are shrunk by 3 cells (90 meters), regrouped into patches, and patches greater than or equal to 50 hectares extracted. Lowland mixed Lmgps Patches of lowland mixed forest types. A - 33 Appendix H - Natural Vegetation Type datalayers and descriptions - continued Lowland mixed forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted and grouped together into patches. Lmjgps Patches of lowland mixed forest types, cut by major roads. Lowland mixed forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. Major roads are converted to raster and the road raster removed from the forest land cover types. The remaining lowland mixed forest cover type cells are then grouped together into patches. Lmmgps Patches of lowland mixed forest types, cut by all roads. Lowland mixed forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. All roads are converted to raster and the road raster removed from the forest land cover types. The remaining lowland mixed forest cover type cells are then grouped together into patches. Lowmix_min Patches of lowland mixed forest types, greater than 50 hectares. Lowland mixed forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted and grouped together into patches. Patches greater than or equal to 50 hectares are extracted. Lowmixj _min Patches of lowland mixed forest types, cut by major roads, greater than 50 hectares. Lowland mixed forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. Major roads are converted to raster and the road raster removed from the lowland mixed forest land cover types. The remaining lowland mixed forest cover type cells are then grouped together into patches. Patches greater than or equal to 50 hectares are extracted. Lowmixm _min Patches of lowland mixed forest types, cut by all roads, greater than 50 hectares. Lowland mixed forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. All roads are converted to raster and the road raster removed from the lowland mixed forest land cover types. The remaining lowland mixed forest cover type cells are then grouped together into patches. Patches greater than or equal to 50 hectares are extracted. A - 34 Appendix H - Natural Vegetation Type datalayers and descriptions - continued Lowmix_90 Patches of lowland deciduous forest types, greater than 50 hectares, after buffering inward 90 meters. Lowland mixed forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted and grouped together into patches. Patches greater than or equal to 2000 hectares are extracted. Patches are shrunk by 3 cells (90 meters), regrouped into patches, and patches greater than or equal to 50 hectares extracted. Lowmixj_90 Patches of lowland mixed forest types, cut by major roads, greater than 50 hectares, after buffering inward 90 meters. Lowland mixed forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. Major roads are converted to raster and the road raster removed from the lowland mixed forest land cover types. The remaining lowland mixed forest cover type cells are then grouped together into patches. Patches greater than or equal to 50 hectares are extracted. Patches are shrunk by 3 cells (90 meters), regrouped into patches, and patches greater than or equal to 50 hectares extracted. Lowmixm_90 Patches of lowland mixed forest types, cut by all roads, greater than 50 hectares, after buffering inward 90 meters. Lowland mixed forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. All roads are converted to raster and the road raster removed from the lowland mixed forest land cover types. The remaining lowland mixed forest cover type cells are then grouped together into patches. Patches greater than or equal to 50 hectares are extracted. Patches are shrunk by 3 cells (90 meters), regrouped into patches, and patches greater than or equal to 50 hectares extracted. Upland coniferous Upcgps Patches of upland coniferous forest types. Upland coniferous forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted and grouped together into patches. Upcjgps Patches of upland coniferous forest types, cut by major roads. Upland coniferous forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. Major roads are converted to raster and the road raster removed from the upland coniferous forest land cover types. The remaining upland coniferous forest cover type cells are then grouped together into patches. A - 35 Appendix H - Natural Vegetation Type datalayers and descriptions - continued Upcmgps Patches of upland coniferous forest types, cut by all roads. Upland coniferous forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. All roads are converted to raster and the road raster removed from the upland forest coniferous land cover types. The remaining upland coniferous forest cover type cells are then grouped together into patches. Upcon_min Patches of upland coniferous forest types, greater than 50 hectares. Upland coniferous forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted and grouped together into patches. Patches greater than or equal to 50 hectares are extracted. Upconj _min Patches of upland coniferous forest types, cut by major roads, greater than 50 hectares. Upland coniferous forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. Major roads are converted to raster and the road raster removed from the upland coniferous forest land cover types. The remaining upland coniferous forest cover type cells are then grouped together into patches. Patches greater than or equal to 50 hectares are extracted. Upconm _min Patches of upland coniferous forest types, cut by all roads, greater than 50 hectares. Upland coniferous forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. All roads are converted to raster and the road raster removed from the upland coniferous forest land cover types. The remaining upland coniferous forest cover type cells are then grouped together into patches. Patches greater than or equal to 50 hectares are extracted. Upcon _90 Patches of upland coniferous forest types, greater than 50 hectares, after buffering inward 90 meters. Upland coniferous forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted and grouped together into patches. Patches greater than or equal to 2000 hectares are extracted. Patches are shrunk by 3 cells (90 meters), regrouped into patches, and patches greater than or equal to 50 hectares extracted. Upcon _210 Patches of upland coniferous forest types, greater than 50 hectares, after buffering inward 210 meters. A - 36 Appendix H - Natural Vegetation Type datalayers and descriptions - continued Upland coniferous forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted and grouped together into patches. Patches greater than or equal to 2000 hectares are extracted. Patches are shrunk by 7 cells (210 meters), regrouped into patches, and patches greater than or equal to 50 hectares extracted. Upcon _300 Patches of upland coniferous forest types, greater than 50 hectares, after buffering inward 300 meters. Upland coniferous forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted and grouped together into patches. Patches greater than or equal to 2000 hectares are extracted. Patches are shrunk by 10 cells (300 meters), regrouped into patches, and patches greater than or equal to 50 hectares extracted. Upconj_90 Patches of upland coniferous forest types, cut by major roads, greater than 50 hectares, after buffering inward 90 meters. Upland coniferous forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. Major roads are converted to raster and the road raster removed from the upland coniferous forest land cover types. The remaining upland coniferous forest cover type cells are then grouped together into patches. Patches greater than or equal to 50 hectares are extracted. Patches are shrunk by 3 cells (90 meters), regrouped into patches, and patches greater than or equal to 50 hectares extracted. Upconj_210 Patches of upland coniferous forest types, cut by major roads, greater than 50 hectares, after buffering inward 210 meters. Upland coniferous forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. Major roads are converted to raster and the road raster removed from the upland coniferous forest land cover types. The remaining upland coniferous forest cover type cells are then grouped together into patches. Patches greater than or equal to 50 hectares are extracted. Patches are shrunk by 7 cells (210 meters), regrouped into patches, and patches greater than or equal to 50 hectares extracted. Upconj_300 Patches of upland coniferous forest types, cut by major roads, greater than 50 hectares, after buffering inward 300 meters. Upland coniferous forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. Major roads are converted to raster and the road raster removed from the upland coniferous forest land cover types. The remaining upland coniferous forest cover type cells are then grouped together into A - 37 Appendix H - Natural Vegetation Type datalayers and descriptions - continued patches. Patches greater than or equal to 50 hectares are extracted. Patches are shrunk by 10 cells (300 meters), regrouped into patches, and patches greater than or equal to 50 hectares extracted. Upconm_90 Patches of upland coniferous forest types, cut by all roads, greater than 50 hectares, after buffering inward 90 meters. Upland coniferous forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. All roads are converted to raster and the road raster removed from the upland coniferous forest land cover types. The remaining upland coniferous forest cover type cells are then grouped together into patches. Patches greater than or equal to 50 hectares are extracted. Patches are shrunk by 3 cells (90 meters), regrouped into patches, and patches greater than or equal to 50 hectares extracted. Upconm_210 Patches of upland coniferous forest types, cut by all roads, greater than 50 hectares, after buffering inward 210 meters. Upland coniferous forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. All roads are converted to raster and the road raster removed from the upland coniferous forest land cover types. The remaining upland coniferous forest cover type cells are then grouped together into patches. Patches greater than or equal to 50 hectares are extracted. Patches are shrunk by 7 cells (210 meters), regrouped into patches, and patches greater than or equal to 50 hectares extracted. Upconm_300 Patches of upland coniferous forest types, cut by all roads, greater than 50 hectares, after buffering inward 300 meters. Upland coniferous forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. All roads are converted to raster and the road raster removed from the upland coniferous forest land cover types. The remaining upland coniferous forest cover type cells are then grouped together into patches. Patches greater than or equal to 50 hectares are extracted. Patches are shrunk by 10 cells (300 meters), regrouped into patches, and patches greater than or equal to 50 hectares extracted. Upland deciduous Updgps Patches of upland deciduous forest types. Upland deciduous forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted and grouped together into patches. Updjgps Patches of upland deciduous forest types, cut by major roads. A - 38 Appendix H - Natural Vegetation Type datalayers and descriptions - continued Upland deciduous forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. Major roads are converted to raster and the road raster removed from the upland deciduous forest land cover types. The remaining upland deciduous forest cover type cells are then grouped together into patches. Updmgps Patches of upland deciduous forest types, cut by all roads. Upland deciduous forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. All roads are converted to raster and the road raster removed from the upland deciduous forest land cover types. The remaining upland deciduous forest cover type cells are then grouped together into patches. Updec_min Patches of deciduous forest types, greater than 2000 hectares. Upland deciduous forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted and grouped together into patches. Patches greater than or equal to 2000 hectares are extracted. Updecj_min Patches of deciduous forest types, cut by major roads, greater than 2000 hectares. Upland deciduous forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. Major roads are converted to raster and the road raster removed from the upland deciduous forest land cover types. The remaining upland deciduous forest cover type cells are then grouped together into patches. Patches greater than or equal to 2000 hectares are extracted. Updecm_min Patches of deciduous forest types, cut by all roads, greater than 2000 hectares. Upland deciduous forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. All roads are converted to raster and the road raster removed from the upland deciduous forest land cover types. The remaining upland deciduous forest cover type cells are then grouped together into patches. Patches greater than or equal to 2000 hectares are extracted. Updec_90 Patches of deciduous forest types, greater than 2000 hectares, after buffering inward 90 meters. Upland deciduous forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted and grouped together into patches. Patches greater than or equal to 2000 hectares are extracted. Patches are shrunk by 3 cells (90 meters), regrouped into patches, and patches greater than or equal to 2000 hectares extracted. A - 39 Appendix H - Natural Vegetation Type datalayers and descriptions - continued Updec_210 Patches of deciduous forest types, greater than 2000 hectares, after buffering inward 210 meters. Upland deciduous forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted and grouped together into patches. Patches greater than or equal to 2000 hectares are extracted. Patches are shrunk by 7 cells (210 meters), regrouped into patches, and patches greater than or equal to 2000 hectares extracted. Updec_300 Patches of deciduous forest types, greater than 2000 hectares, after buffering inward 300 meters. Upland deciduous forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted and grouped together into patches. Patches greater than or equal to 2000 hectares are extracted. Patches are shrunk by 10 cells (300 meters), regrouped into patches, and patches greater than or equal to 2000 hectares extracted. Updecj_90 Patches of upland deciduous forest types, cut by major roads, after buffering inward 90 meters. Upland deciduous forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. Major roads are converted to raster and the road raster removed from the upland deciduous forest land cover types. The remaining upland deciduous forest cover type cells are then grouped together into patches. Patches greater than or equal to 50 hectares are extracted. Patches are shrunk by 3 cells (90 meters), regrouped into patches, and patches greater than or equal to 2000 hectares extracted. Updecj_210 Patches of upland deciduous forest types, cut by major roads, after buffering inward 210 meters. Upland deciduous forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. Major roads are converted to raster and the road raster removed from the upland deciduous forest land cover types. The remaining upland deciduous forest cover type cells are then grouped together into patches. Patches greater than or equal to 50 hectares are extracted. Patches are shrunk by 7 cells (210 meters), regrouped into patches, and patches greater than or equal to 2000 hectares extracted. Updecj_300 Patches of upland deciduous forest types, cut by major roads, after buffering inward 300 meters. Upland deciduous forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. Major roads are converted to raster and the road raster removed from the upland deciduous forest land cover types. The remaining upland deciduous forest cover type cells are then grouped together into patches. Patches greater than or equal to 50 hectares are extracted. Patches are shrunk by 10 cells (300 meters), regrouped into patches, and patches greater than or equal to 2000 hectares extracted. A - 40 Appendix H - Natural Vegetation Type datalayers and descriptions - continued Updecm_90 Patches of upland deciduous forest types, cut by all roads, after buffering inward 90 meters. Upland deciduous forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. All roads are converted to raster and the road raster removed from the upland deciduous forest land cover types. The remaining upland deciduous forest cover type cells are then grouped together into patches. Patches greater than or equal to 50 hectares are extracted. Patches are shrunk by 3 cells (90 meters), regrouped into patches, and patches greater than or equal to 2000 hectares extracted. Upland forests Upfgps Patches of all upland forest types. All upland forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted and grouped together into patches. Upfjgps Patches of all upland forest types, cut by major roads. All upland forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. Major roads are converted to raster and the road raster removed from the upland forest land cover types. The remaining upland forest cover type cells are then grouped together into patches. Upfmgps Patches of all upland forest types, cut by all roads. All upland forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. All roads are converted to raster and the road raster removed from the upland forest land cover types. The remaining upland forest cover type cells are then grouped together into patches. Upf_min Patches of all upland forest types, greater than 2000 hectares. All upland forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted and grouped together into patches. Patches greater than or equal to 2000 hectares are extracted. Upfj_min Patches of all upland forest types, cut by major roads, greater than 2000 hectares. All upland forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. Major roads are converted to raster and the road raster removed from the upland forest land cover types. The A - 41 Appendix H - Natural Vegetation Type datalayers and descriptions - continued remaining upland forest cover type cells are then grouped together into patches. Patches greater than or equal to 2000 hectares are extracted. Upfm_min Patches of all upland forest types, cut by all roads, greater than 2000 hectares. All upland forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. All roads are converted to raster and the road raster removed from the upland forest land cover types. The remaining upland forest cover type cells are then grouped together into patches. Patches greater than or equal to 2000 hectares are extracted. Upf_90 Patches of all upland forest types, greater than 2000 hectares, after buffering inward 90 meters. All upland forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted and grouped together into patches. Patches greater than or equal to 2000 hectares are extracted. Patches are shrunk by 3 cells (90 meters), regrouped into patches, and patches greater than or equal to 2000 hectares extracted. Upfor_210 Patches of all upland forest types, greater than 2000 hectares, after buffering inward 210 meters. All upland forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted and grouped together into patches. Patches greater than or equal to 2000 hectares are extracted. Patches are shrunk by 7 cells (210 meters), regrouped into patches, and patches greater than or equal to 2000 hectares extracted. Upf_300 Patches of all upland forest types, greater than 2000 hectares, after buffering inward 300 meters. All upland forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted and grouped together into patches. Patches greater than or equal to 2000 hectares are extracted. Patches are shrunk by 10 cells (300 meters), regrouped into patches, and patches greater than or equal to 2000 hectares extracted. Upfj_90 Patches of all upland forest types, cut by major roads, greater than 2000 hectares, after buffering inward 90 meters. All upland forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. Major roads are converted to raster and the road raster removed from the upland forest land cover types. The remaining upland forest cover type cells are then grouped together into patches. Patches greater than or equal to 2000 hectares are extracted. Patches are shrunk by 3 cells (90 meters), regrouped into patches, and patches greater than or equal to 2000 hectares extracted. A - 42 Appendix H - Natural Vegetation Type datalayers and descriptions - continued Upfj_210 Patches of all upland forest types, cut by major roads, greater than 2000 hectares, after buffering inward 210 meters. All upland forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. Major roads are converted to raster and the road raster removed from the upland forest land cover types. The remaining upland forest cover type cells are then grouped together into patches. Patches greater than or equal to 2000 hectares are extracted. Patches are shrunk by 7 cells (210 meters), regrouped into patches, and patches greater than or equal to 2000 hectares extracted. Upfj_300 Patches of all upland forest types, cut by major roads, greater than 2000 hectares, after buffering inward 300 meters. All upland forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. Major roads are converted to raster and the road raster removed from the upland forest land cover types. The remaining upland forest cover type cells are then grouped together into patches. Patches greater than or equal to 2000 hectares are extracted. Patches are shrunk by 10 cells (300 meters), regrouped into patches, and patches greater than or equal to 2000 hectares extracted. Upfm_90 Patches of all upland forest types, cut by all roads, greater than 2000 hectares, after buffering inward 90 meters. All upland forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. All roads are converted to raster and the road raster removed from the upland forest land cover types. The remaining upland forest cover type cells are then grouped together into patches. Patches greater than or equal to 2000 hectares are extracted. Patches are shrunk by 3 cells (90 meters), regrouped into patches, and patches greater than or equal to 2000 hectares extracted. Upfm_210 Patches of all upland forest types, cut by all roads, greater than 2000 hectares, after buffering inward 210 meters. All upland forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. All roads are converted to raster and the road raster removed from the upland forest land cover types. The remaining upland forest cover type cells are then grouped together into patches. Patches greater than or equal to 2000 hectares are extracted. Patches are shrunk by 7 cells (210 meters), regrouped into patches, and patches greater than or equal to 2000 hectares extracted. A - 43 Appendix H - Natural Vegetation Type datalayers and descriptions - continued Upfm_300 Patches of all upland forest types, cut by all roads, greater than 2000 hectares, after buffering inward 300 meters. All upland forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. All roads are converted to raster and the road raster removed from the upland forest land cover types. The remaining upland forest cover type cells are then grouped together into patches. Patches greater than or equal to 2000 hectares are extracted. Patches are shrunk by 10 cells (300 meters), regrouped into patches, and patches greater than or equal to 2000 hectares extracted. Wetland forests Wforgps Patches of all wetland forest types. All wetland forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted and grouped together into patches. Wforjgps Patches of all wetland forest types, cut by major roads. All wetland forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. Major roads are converted to raster and the road raster removed from the wetland forest land cover types. The remaining wetland forest cover type cells are then grouped together into patches. Wformgps Patches of all wetland forest types, cut by all roads. All wetland forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. All roads are converted to raster and the road raster removed from the wetland forest land cover types. The remaining wetland forest cover type cells are then grouped together into patches. Wetfor_min Patches of all wetland forest types, greater than 50 hectares. All wetland forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted and grouped together into patches. Patches greater than or equal to 50 hectares are extracted. Wetforj_min Patches of all wetland forest types, cut by major roads, greater than 50 hectares. A - 44 Appendix H - Natural Vegetation Type datalayers and descriptions - continued All wetland forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. Major roads are converted to raster and the road raster removed from the wetland forest land cover types. The remaining wetland forest cover type cells are then grouped together into patches. Patches greater than or equal to 50 hectares are extracted. Wetform_min Patches of all wetland forest types, cut by all roads, greater than 50 hectares. All wetland forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. All roads are converted to raster and the road raster removed from the wetland forest land cover types. The remaining wetland forest cover type cells are then grouped together into patches. Patches greater than or equal to 50 hectares are extracted. Wetfor_90 Patches of all wetland forest types, greater than 50 hectares, after buffering inward 90 meters. All wetland forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted and grouped together into patches. Patches greater than or equal to 50 hectares are extracted. Patches are shrunk by 3 cells (90 meters), regrouped into patches, and patches greater than or equal to 50 hectares extracted. Wetfor_210 Patches of all wetland forest types, greater than 50 hectares, after buffering inward 210 meters. All wetland forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted and grouped together into patches. Patches greater than or equal to 50 hectares are extracted. Patches are shrunk by 7 cells (210 meters), regrouped into patches, and patches greater than or equal to 50 hectares extracted. Wetfor_300 Patches of all wetland forest types, greater than 50 hectares after buffering inward 300 meters. All wetland forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted and grouped together into patches. Patches greater than or equal to 50 hectares are extracted. Patches are shrunk by 10 cells (300 meters), regrouped into patches, and patches greater than or equal to 50 hectares extracted. Wetforj_90 Patches of all wetland forest types, cut by major roads, greater than 50 hectares after buffering inward 90 meters. All wetland forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. Major roads are converted to raster and the road raster removed from the wetland forest land cover types. The A - 45 Appendix H - Natural Vegetation Type datalayers and descriptions - continued remaining wetland forest cover type cells are then grouped together into patches. Patches greater than or equal to 50 hectares are extracted. Patches are shrunk by 3 cells (90 meters), regrouped into patches, and patches greater than or equal to 50 hectares extracted. Wetforj_210 Patches of all wetland forest types, cut by major roads, greater than 50 hectares after buffering inward 210 meters. All wetland forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. Major roads are converted to raster and the road raster removed from the wetland forest land cover types. The remaining wetland forest cover type cells are then grouped together into patches. Patches greater than or equal to 50 hectares are extracted. Patches are shrunk by 7 cells (210 meters), regrouped into patches, and patches greater than or equal to 50 hectares extracted. Wetforj_300 Patches of all wetland forest types, cut by major roads, greater than 50 hectares after buffering inward 300 meters. All wetland forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. Major roads are converted to raster and the road raster removed from the wetland forest land cover types. The remaining wetland forest cover type cells are then grouped together into patches. Patches greater than or equal to 50 hectares are extracted. Patches are shrunk by 10 cells (300 meters), regrouped into patches, and patches greater than or equal to 50 hectares extracted. Wetform_90 Patches of all wetland forest types, cut by all roads, greater than 50 hectares after buffering inward 90 meters. All wetland forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. All roads are converted to raster and the road raster removed from the wetland forest land cover types. The remaining wetland forest cover type cells are then grouped together into patches. Patches greater than or equal to 50 hectares are extracted. Patches are shrunk by 3 cells (90 meters), regrouped into patches, and patches greater than or equal to 50 hectares extracted. Wetform_210 Patches of all wetland forest types, cut by all roads, greater than 50 hectares after buffering inward 210 meters. All wetland forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. All roads are converted to raster and the road raster removed from the wetland forest land cover types. The remaining wetland forest cover type cells are then grouped together into patches. Patches greater than or equal to 50 hectares are extracted. Patches are shrunk by 7 cells (210 meters), regrouped into patches, and patches greater than or equal to 50 hectares extracted. A - 46 Appendix H - Natural Vegetation Type datalayers and descriptions - continued Wetform_300 Patches of all wetland forest types, cut by all roads, greater than 50 hectares after buffering inward 300 meters. All wetland forest land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. All roads are converted to raster and the road raster removed from the wetland forest land cover types. The remaining wetland forest cover type cells are then grouped together into patches. Patches greater than or equal to 50 hectares are extracted. Patches are shrunk by 10 cells (300 meters), regrouped into patches, and patches greater than or equal to 50 hectares extracted. Grasslands Grassgps Patches of current grasslands in areas know to have historic grasslands. All grassland cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) that coincide with grassland cover types in the Circa 1800 vegetation layer (BLACK OAK BARREN, EXPOSED BEDROCK, GRASSLAND, JACK PINE-RED PINE FOREST, MIXED OAK FOREST, MIXED OAK SAVANNA, MIXED PINE-OAK FOREST, OAK-HICKORY FOREST, OAK/PINE BARRENS, PINE BARRENS, SAND DUNE, WHITE PINE-RED PINE FOREST, WHITE PINE-WHITE OAK FOREST, WET PRAIRIE) are extracted and grouped together into patches. Grassjgps Patches of current grasslands in areas know to have historic grasslands, cut by major roads. All grassland cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) that coincide with grassland cover types in the Circa 1800 vegetation layer (BLACK OAK BARREN, EXPOSED BEDROCK, GRASSLAND, JACK PINE-RED PINE FOREST, MIXED OAK FOREST, MIXED OAK SAVANNA, MIXED PINE-OAK FOREST, OAK-HICKORY FOREST, OAK/PINE BARRENS, PINE BARRENS, SAND DUNE, WHITE PINE-RED PINE FOREST, WHITE PINE-WHITE OAK FOREST, WET PRAIRIE) are extracted and grouped together into patches. Major roads are converted to raster and the road raster removed from the grassland cover types. The remaining grassland cover type cells are then grouped together into patches. Grassmgps Patches of current grasslands in areas know to have historic grasslands, cut by all roads. All grassland cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) that coincide with grassland cover types in the Circa 1800 vegetation layer (BLACK OAK BARREN, EXPOSED BEDROCK, GRASSLAND, JACK PINE-RED PINE FOREST, MIXED OAK FOREST, MIXED OAK SAVANNA, MIXED PINE-OAK FOREST, OAK-HICKORY FOREST, OAK/PINE BARRENS, PINE BARRENS, SAND DUNE, WHITE PINE-RED PINE FOREST, WHITE PINE-WHITE OAK FOREST, WET PRAIRIE) are extracted and grouped together into patches. All roads are converted to raster and the road raster removed from grassland cover types. The remaining grassland cover type cells are then grouped together into patches. A - 47 Appendix H - Natural Vegetation Type datalayers and descriptions - continued Grass_min Patches of current grasslands in areas know to have historic grasslands, greater than 50 hectares. All grassland cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) that coincide with grassland cover types in the Circa 1800 vegetation layer (BLACK OAK BARREN, EXPOSED BEDROCK, GRASSLAND, JACK PINE-RED PINE FOREST, MIXED OAK FOREST, MIXED OAK SAVANNA, MIXED PINE-OAK FOREST, OAK-HICKORY FOREST, OAK/PINE BARRENS, PINE BARRENS, SAND DUNE, WHITE PINE-RED PINE FOREST, WHITE PINE-WHITE OAK FOREST, WET PRAIRIE) are extracted and grouped together into patches and patches greater than or equal to 50 hectares are extracted. Grassj_min Patches of current grasslands in areas know to have historic grasslands, cut by major roads, greater than 50 hectares. All grassland cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) that coincide with grassland cover types in the Circa 1800 vegetation layer (BLACK OAK BARREN, EXPOSED BEDROCK, GRASSLAND, JACK PINE-RED PINE FOREST, MIXED OAK FOREST, MIXED OAK SAVANNA, MIXED PINE-OAK FOREST, OAK-HICKORY FOREST, OAK/PINE BARRENS, PINE BARRENS, SAND DUNE, WHITE PINE-RED PINE FOREST, WHITE PINE-WHITE OAK FOREST, WET PRAIRIE) are extracted and grouped together into patches. Major roads are converted to raster and the road raster removed from the grassland cover types. The remaining grassland cover type cells are then grouped together into patches and patches greater than or equal to 50 hectares are extracted. Grassm_min Patches of current grasslands in areas know to have historic grasslands, cut by all roads, greater than 50 hectares. All grassland cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) that coincide with grassland cover types in the Circa 1800 vegetation layer (BLACK OAK BARREN, EXPOSED BEDROCK, GRASSLAND, JACK PINE-RED PINE FOREST, MIXED OAK FOREST, MIXED OAK SAVANNA, MIXED PINE-OAK FOREST, OAK-HICKORY FOREST, OAK/PINE BARRENS, PINE BARRENS, SAND DUNE, WHITE PINE-RED PINE FOREST, WHITE PINE-WHITE OAK FOREST, WET PRAIRIE) are extracted and grouped together into patches. Major roads are converted to raster and the road raster removed from the grassland cover types. The remaining grassland cover type cells are then grouped together into patches and patches greater than or equal to 50 hectares are extracted. Grass_90 Patches of current grasslands in areas known to have historic grasslands, greater than 50 hectares after buffering inward 90 meters. All grassland cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) that coincide with grassland cover types in the A - 48 Appendix H - Natural Vegetation Type datalayers and descriptions - continued Circa 1800 vegetation layer (BLACK OAK BARREN, EXPOSED BEDROCK, GRASSLAND, JACK PINE-RED PINE FOREST, MIXED OAK FOREST, MIXED OAK SAVANNA, MIXED PINE-OAK FOREST, OAK-HICKORY FOREST, OAK/PINE BARRENS, PINE BARRENS, SAND DUNE, WHITE PINE-RED PINE FOREST, WHITE PINE-WHITE OAK FOREST, WET PRAIRIE) are extracted and grouped together into patches and patches greater than or equal to 50 hectares are extracted. These patches are then shrunk by 3 cells (90 meters), regrouped into patches, and patches greater than or equal to 50 hectares extracted. Grass_210 Patches of current grasslands in areas known to have historic grasslands, greater than 50 hectares after buffering inward 210 meters. All grassland cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) that coincide with grassland cover types in the Circa 1800 vegetation layer (BLACK OAK BARREN, EXPOSED BEDROCK, GRASSLAND, JACK PINE-RED PINE FOREST, MIXED OAK FOREST, MIXED OAK SAVANNA, MIXED PINE-OAK FOREST, OAK-HICKORY FOREST, OAK/PINE BARRENS, PINE BARRENS, SAND DUNE, WHITE PINE-RED PINE FOREST, WHITE PINE-WHITE OAK FOREST, WET PRAIRIE) are extracted and grouped together into patches and patches greater than or equal to 50 hectares are extracted. These patches are then shrunk by 7 cells (210 meters), regrouped into patches, and patches greater than or equal to 50 hectares extracted. Grass_300 Patches of current grasslands in areas known to have historic grasslands, greater than 50 hectares after buffering inward 300 meters. All grassland cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) that coincide with grassland cover types in the Circa 1800 vegetation layer (BLACK OAK BARREN, EXPOSED BEDROCK, GRASSLAND, JACK PINE-RED PINE FOREST, MIXED OAK FOREST, MIXED OAK SAVANNA, MIXED PINE-OAK FOREST, OAK-HICKORY FOREST, OAK/PINE BARRENS, PINE BARRENS, SAND DUNE, WHITE PINE-RED PINE FOREST, WHITE PINE-WHITE OAK FOREST, WET PRAIRIE) are extracted and grouped together into patches and patches greater than or equal to 50 hectares are extracted. These patches are then shrunk by 10 cells (300 meters), regrouped into patches, and patches greater than or equal to 50 hectares extracted. Grassj_90 Patches of current grasslands in areas known to have historic grasslands, cut by major roads, greater than 50 hectares after buffering inward 90 meters. All grassland cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) that coincide with grassland cover types in the Circa 1800 vegetation layer (BLACK OAK BARREN, EXPOSED BEDROCK, GRASSLAND, JACK PINE-RED PINE FOREST, MIXED OAK FOREST, MIXED OAK SAVANNA, MIXED PINE-OAK FOREST, OAK-HICKORY FOREST, OAK/PINE BARRENS, PINE BARRENS, SAND DUNE, WHITE PINE-RED PINE FOREST, WHITE PINE-WHITE OAK FOREST, WET PRAIRIE) A - 49 Appendix H - Natural Vegetation Type datalayers and descriptions - continued are extracted and grouped together into patches. Major roads are converted to raster and the road raster removed from the grassland cover types. The remaining grassland cover type cells are then grouped together into patches and patches greater than or equal to 50 hectares are extracted. These patches are then shrunk by 3 cells (90 meters), regrouped into patches, and patches greater than or equal to 50 hectares extracted. Grassj_210 Patches of current grasslands in areas known to have historic grasslands, cut by major roads, greater than 50 hectares after buffering inward 210 meters. All grassland cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) that coincide with grassland cover types in the Circa 1800 vegetation layer (BLACK OAK BARREN, EXPOSED BEDROCK, GRASSLAND, JACK PINE-RED PINE FOREST, MIXED OAK FOREST, MIXED OAK SAVANNA, MIXED PINE-OAK FOREST, OAK-HICKORY FOREST, OAK/PINE BARRENS, PINE BARRENS, SAND DUNE, WHITE PINE-RED PINE FOREST, WHITE PINE-WHITE OAK FOREST, WET PRAIRIE) are extracted and grouped together into patches. Major roads are converted to raster and the road raster removed from the grassland cover types. The remaining grassland cover type cells are then grouped together into patches and patches greater than or equal to 50 hectares are extracted. These patches are then shrunk by 7 cells (210 meters), regrouped into patches, and patches greater than or equal to 50 hectares extracted. Grassj_300 Patches of current grasslands in areas known to have historic grasslands, cut by major roads, greater than 50 hectares after buffering inward 300 meters. All grassland cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) that coincide with grassland cover types in the Circa 1800 vegetation layer (BLACK OAK BARREN, EXPOSED BEDROCK, GRASSLAND, JACK PINE-RED PINE FOREST, MIXED OAK FOREST, MIXED OAK SAVANNA, MIXED PINE-OAK FOREST, OAK-HICKORY FOREST, OAK/PINE BARRENS, PINE BARRENS, SAND DUNE, WHITE PINE-RED PINE FOREST, WHITE PINE-WHITE OAK FOREST, WET PRAIRIE) are extracted and grouped together into patches. Major roads are converted to raster and the road raster removed from the grassland cover types. The remaining grassland cover type cells are then grouped together into patches and patches greater than or equal to 50 hectares are extracted. These patches are then shrunk by 10 cells (300 meters), regrouped into patches, and patches greater than or equal to 50 hectares extracted. Grassm_90 Patches of current grasslands in areas known to have historic grasslands, cut by all roads, greater than 50 hectares after buffering inward 90 meters. All grassland cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) that coincide with grassland cover types in the Circa 1800 vegetation layer (BLACK OAK BARREN, EXPOSED BEDROCK, GRASSLAND, JACK PINE-RED PINE FOREST, MIXED OAK FOREST, MIXED OAK SAVANNA, MIXED PINE-OAK FOREST, OAK-HICKORY FOREST, OAK/PINE BARRENS, PINE BARRENS, SAND A - 50 Appendix H - Natural Vegetation Type datalayers and descriptions - continued DUNE, WHITE PINE-RED PINE FOREST, WHITE PINE-WHITE OAK FOREST, WET PRAIRIE) are extracted and grouped together into patches. Major roads are converted to raster and the road raster removed from the grassland cover types. The remaining grassland cover type cells are then grouped together into patches and patches greater than or equal to 50 hectares are extracted. These patches are then shrunk by 3 cells (90 meters), regrouped into patches, and patches greater than or equal to 50 hectares extracted. Grassm_210 Patches of current grasslands in areas known to have historic grasslands, cut by all roads, greater than 50 hectares after buffering inward 210 meters. All grassland cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) that coincide with grassland cover types in the Circa 1800 vegetation layer (BLACK OAK BARREN, EXPOSED BEDROCK, GRASSLAND, JACK PINE-RED PINE FOREST, MIXED OAK FOREST, MIXED OAK SAVANNA, MIXED PINE-OAK FOREST, OAK-HICKORY FOREST, OAK/PINE BARRENS, PINE BARRENS, SAND DUNE, WHITE PINE-RED PINE FOREST, WHITE PINE-WHITE OAK FOREST, WET PRAIRIE) are extracted and grouped together into patches. Major roads are converted to raster and the road raster removed from the grassland cover types. The remaining grassland cover type cells are then grouped together into patches and patches greater than or equal to 50 hectares are extracted. These patches are then shrunk by 7 cells (210 meters), regrouped into patches, and patches greater than or equal to 50 hectares extracted. Grassm_300 Patches of current grasslands in areas known to have historic grasslands, cut by all roads, greater than 50 hectares after buffering inward 300 meters. All grassland cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) that coincide with grassland cover types in the Circa 1800 vegetation layer (BLACK OAK BARREN, EXPOSED BEDROCK, GRASSLAND, JACK PINE-RED PINE FOREST, MIXED OAK FOREST, MIXED OAK SAVANNA, MIXED PINE-OAK FOREST, OAK-HICKORY FOREST, OAK/PINE BARRENS, PINE BARRENS, SAND DUNE, WHITE PINE-RED PINE FOREST, WHITE PINE-WHITE OAK FOREST, WET PRAIRIE) are extracted and grouped together into patches. Major roads are converted to raster and the road raster removed from the grassland cover types. The remaining grassland cover type cells are then grouped together into patches and patches greater than or equal to 50 hectares are extracted. These patches are then shrunk by 10 cells (300 meters), regrouped into patches, and patches greater than or equal to 50 hectares extracted. Non-forested wetlands Nforgps Patches of all non-forested wetland types. All non-forested wetland land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted and grouped together into patches. A - 51 Appendix H - Natural Vegetation Type datalayers and descriptions - continued Nforjgps Patches of all non-forested wetland types, cut by major roads. All non-forested wetland land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. Major roads are converted to raster and the road raster removed from the non-forested wetland land cover types. The remaining non-forested wetland cover type cells are then grouped together into patches. Nformgps Patches of all non-forested wetland types, cut by major roads. All non-forested wetland land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. All roads are converted to raster and the road raster removed from the non-forested wetland land cover types. The remaining non-forested wetland cover type cells are then grouped together into patches. Nonfor_90 Patches of all non-forested wetland types, after buffering inward 90 meters. All non-forested wetland land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted and grouped together into patches. Patches are shrunk by 3 cells (90 meters) and then regrouped into patches. Nonfor_210 Patches of all non-forested wetland types, after buffering inward 210 meters. All non-forested wetland land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted and grouped together into patches. Patches are shrunk by 7 cells (210 meters) and regrouped into patches. Nonfor_300 Patches of all non-forested wetland types, after buffering inward 210 meters. All non-forested wetland land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted and grouped together into patches. Patches are shrunk by 10 cells (300 meters) and regrouped into patches. Nonforj_90 Patches of all non-forested wetland types, cut by major roads, after buffering inward 90 meters. All non-forested wetland land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. Major roads are converted to raster and the road raster removed from the non-forested wetland land cover types. The remaining non-forested wetland cover type cells are then grouped together into patches. Patches are shrunk by 3 cells (90 meters) and then regrouped into patches. A - 52 Appendix H - Natural Vegetation Type datalayers and descriptions - continued Nonforj_210 Patches of all non-forested wetland types, cut by major roads, after buffering inward 210 meters. All non-forested wetland land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. Major roads are converted to raster and the road raster removed from the non-forested wetland land cover types. The remaining non-forested wetland cover type cells are then grouped together into patches. Patches are shrunk by 7 cells (210 meters) and then regrouped into patches. Nonforj_300 Patches of all non-forested wetland types, cut by major roads, after buffering inward 300 meters. All non-forested wetland land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. Major roads are converted to raster and the road raster removed from the non-forested wetland land cover types. The remaining non-forested wetland cover type cells are then grouped together into patches. Patches are shrunk by 10 cells (300 meters) and then regrouped into patches. Nonform_90 Patches of all non-forested wetland types, cut by major roads, after buffering inward 90 meters. All non-forested wetland land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. All roads are converted to raster and the road raster removed from the non-forested wetland land cover types. The remaining non-forested wetland cover type cells are then grouped together into patches. Patches are shrunk by 3 cells (90 meters) and then regrouped into patches. Nonform _210 Patches of all non-forested wetland types, cut by major roads, after buffering inward 210 meters. All non-forested wetland land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. All roads are converted to raster and the road raster removed from the non-forested wetland land cover types. The remaining non-forested wetland cover type cells are then grouped together into patches. Patches are shrunk by 7 cells (210 meters) and then regrouped into patches. Nonform_300 Patches of all non-forested wetland types, cut by major roads, after buffering inward 300 meters. All non-forested wetland land cover types in the Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover (Michigan DNR, 2003) are extracted. All roads are converted to raster and the road raster removed from the non-forested wetland land cover types. The remaining non-forested wetland cover type cells are then grouped together into patches. Patches are shrunk by 10 cells (300 meters) and then regrouped into patches. A - 53 Appendix I - natural vegetation core area datalayers and descriptions File Name nat2up nat2up_c1 nat2up_c2 nat2up_c3 nat2mup nat2mup_c1 nat2mup_c2 nat2mup_c3 nat2jup nat2jup_c1 nat2jup_c2 nat2jup_c3 nat2nlp nat2nlp_c1 nat2nlp_c2 nat2nlp_c3 nat2jnlp nat2jnlp_c1 nat2jnlp_c2 nat2jnlp_c3 nat2mnlp nat2mnlp_c1 nat2mnlp_c2 nat2mnlp_c3 nat2slp nat2slp_c1 nat2slp_c2 nat2slp_c3 nat2jslp nat2jslp_c1 nat2jslp_c2 nat2jslp_c3 nat2mslp nat2mslp_c1 nat2mslp_c2 nat2mslp_c3 A - 54 Description all natural vegetation all natural vegetation all natural vegetation all natural vegetation all natural vegetation all natural vegetation all natural vegetation all natural vegetation all natural vegetation all natural vegetation all natural vegetation all natural vegetation all natural vegetation all natural vegetation all natural vegetation all natural vegetation all natural vegetation all natural vegetation all natural vegetation all natural vegetation all natural vegetation all natural vegetation all natural vegetation all natural vegetation all natural vegetation all natural vegetation all natural vegetation all natural vegetation all natural vegetation all natural vegetation all natural vegetation all natural vegetation all natural vegetation all natural vegetation all natural vegetation all natural vegetation road Ecoregion layer UP none UP none UP none UP none UP major UP major UP major UP major UP all UP all UP all UP all NLP none NLP none NLP none NLP none NLP major NLP major NLP major NLP major NLP all NLP all NLP all NLP all SLP none SLP none SLP none SLP none SLP major SLP major SLP major SLP major SLP all SLP all SLP all SLP all minimum buffer size size in (acres) meters 5,000 0 5,000 90 5,000 210 5,000 300 5,000 0 5,000 90 5,000 210 5,000 300 5,000 0 5,000 90 5,000 210 5,000 300 2,500 0 2,500 90 2,500 210 2,500 300 2,500 0 2,500 90 2,500 210 2,500 300 2,500 0 2,500 90 2,500 210 2,500 300 500 0 500 90 500 210 500 300 500 0 500 90 500 210 500 300 500 0 500 90 500 210 500 300 Appendix I - natural vegetation core area datalayers and descriptions - continued minimum patch sizes dependent on ecoregion Nat2up Michigan Upper Peninsula natural vegetation classes, patches greater than 5000 acres. All natural vegetation classes, including water, are grouped together. Patches greater than or equal to 5000 acres are extracted. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than 5000 acres extracted. Nat2nlp Michigan Northern Lower Peninsula natural vegetation classes, patches greater than 2500 acres. All natural vegetation classes, including water, are grouped together. Patches greater than or equal to 2500 acres are extracted. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than 2500 acres extracted. Nat2slp Michigan Southern Lower Peninsula natural vegetation classes, patches greater than 500 acres. All natural vegetation classes, including water, are grouped together. Patches greater than or equal to 500 acres are extracted. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than 500 acres extracted. Nat2jup Michigan Upper Peninsula natural vegetation classes, patches greater than 5000 acres after cutting by major roads. All natural vegetation classes, including water, are grouped together. Major roads are converted to raster and then removed from the natural vegetation classes. Patches are re-grouped and those greater than or equal to 5000 acres are extracted. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than 5000 acres extracted. Nat2jnlp Michigan Northern Lower Peninsula natural vegetation classes, patches greater than 2500 acres after cutting by major roads. All natural vegetation classes, including water, are grouped together. Major roads are converted to raster and then removed from the natural vegetation classes. Patches are re-grouped and those greater than or equal to 2500 acres are extracted. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than 2500 acres extracted. Nat2jslp Michigan Southern Lower Peninsula natural vegetation classes, patches greater than 500 acres after cutting by major roads. All natural vegetation classes, including water, are grouped together. Major roads are converted to raster and then removed from the natural vegetation classes. Patches are re-grouped and those A - 55 Appendix I - natural vegetation core area datalayers and descriptions - continued greater than or equal to 500 acres are extracted. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than 500 acres extracted. Nat2mup Michigan Upper Peninsula natural vegetation classes, patches greater than 5000 acres after cutting by all roads. All natural vegetation classes, including water, are grouped together. All roads are converted to raster and then removed from the natural vegetation classes. Patches are re-grouped and those greater than or equal to 5000 acres are extracted. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than 5000 acres extracted. Nat2mnlp Michigan Northern Lower Peninsula natural vegetation classes, patches greater than 2500 acres after cutting by all roads. All natural vegetation classes, including water, are grouped together. All roads are converted to raster and then removed from the natural vegetation classes. Patches are re-grouped and those greater than or equal to 2500 acres are extracted. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than 2500 acres extracted. Nat2mslp Michigan Southern Lower Peninsula natural vegetation classes, patches greater than 500 acres after cutting by all roads. All natural vegetation classes, including water, are grouped together. All roads are converted to raster and then removed from the natural vegetation classes. Patches are re-grouped and those greater than or equal to 500 acres are extracted. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than 500 acres extracted. Nat2up_c Michigan Upper Peninsula natural vegetation classes, patches greater than 5000 acres after buffering inward 90 meters. All natural vegetation classes, including water, are grouped together. Patches greater than or equal to 5000 acres are extracted. These patches are then shrunk by 3 cells (90 meters), regrouped, and those patches greater than or equal to 5000 acres are extracted. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than 5000 acres extracted. Nat2nlp_c Michigan Northern Lower Peninsula natural vegetation classes, patches greater than 2500 acres after buffering inward 90 meters. All natural vegetation classes, including water, are grouped together. Patches greater than or equal to 2500 acres are extracted. These patches are then shrunk by 3 cells (90 meters), regrouped, and those patches greater than or equal to 2500 acres are extracted. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than 2500 acres extracted. A - 56 Appendix I - natural vegetation core area datalayers and descriptions - continued Nat2slp_c Michigan Southern Lower Peninsula natural vegetation classes, patches greater than 500 acres after buffering inward 90 meters. All natural vegetation classes, including water, are grouped together. Patches greater than or equal to 500 acres are extracted. These patches are then shrunk by 3 cells (90 meters), regrouped, and those patches greater than or equal to 500 acres are extracted. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than 500 acres extracted. Nat2jup_c Michigan Upper Peninsula natural vegetation classes, patches greater than 5000 acres after cutting by major roads and buffering inward 90 meters. All natural vegetation classes, including water, are grouped together. Major roads are converted to raster and then removed from the natural vegetation classes. Patches are re-grouped and those greater than or equal to 5000 acres are extracted. These patches are then shrunk by 3 cells (90 meters), regrouped, and those patches greater than or equal to 5000 acres are extracted. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than 5000 acres extracted. Nat2jnlp_c Michigan Northern Lower Peninsula natural vegetation classes, patches greater than 2500 acres after cutting by major roads and buffering inward 90 meters. All natural vegetation classes, including water, are grouped together. Major roads are converted to raster and then removed from the natural vegetation classes. Patches are re-grouped and those greater than or equal to 2500 acres are extracted. These patches are then shrunk by 3 cells (90 meters), regrouped, and those patches greater than or equal to 2500 acres are extracted. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than 2500 acres extracted. Nat2jslp_c Michigan Southern Lower Peninsula natural vegetation classes, patches greater than 500 acres after cutting by major roads and buffering inward 90 meters. All natural vegetation classes, including water, are grouped together. Major roads are converted to raster and then removed from the natural vegetation classes. Patches are re-grouped and those greater than or equal to 500 acres are extracted. These patches are then shrunk by 3 cells (90 meters), regrouped, and those patches greater than or equal to 500 acres are extracted. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than 500 acres extracted. Nat2mup_c Michigan Upper Peninsula natural vegetation classes, patches greater than 5000 acres after cutting by all roads and buffering inward 90 meters. A - 57 Appendix I - natural vegetation core area datalayers and descriptions - continued All natural vegetation classes, including water, are grouped together. All roads are converted to raster and then removed from the natural vegetation classes. Patches are re-grouped and those greater than or equal to 5000 acres are extracted. These patches are then shrunk by 3 cells (90 meters), regrouped, and those patches greater than or equal to 5000 acres are extracted. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than 5000 acres extracted. Nat2mnlp_c Michigan Northern Lower Peninsula natural vegetation classes, patches greater than 2500 acres after cutting by all roads and buffering inward 90 meters. All natural vegetation classes, including water, are grouped together. All roads are converted to raster and then removed from the natural vegetation classes. Patches are re-grouped and those greater than or equal to 2500 acres are extracted. These patches are then shrunk by 3 cells (90 meters), regrouped, and those patches greater than or equal to 2500 acres are extracted. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than 2500 acres extracted. Nat2mslp_c Michigan Southern Lower Peninsula natural vegetation classes, patches greater than 500 acres after cutting by all roads and buffering inward 90 meters. All natural vegetation classes, including water, are grouped together. All roads are converted to raster and then removed from the natural vegetation classes. Patches are re-grouped and those greater than or equal to 500 acres are extracted. These patches are then shrunk by 3 cells (90 meters), regrouped, and those patches greater than or equal to 500 acres are extracted. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than 500 acres extracted. Nat2up_c1 Michigan Upper Peninsula natural vegetation classes, patches greater than 5000 acres after buffering inward 90 meters. All natural vegetation classes, including water, are grouped together. Patches greater than or equal to 5000 acres are extracted. These patches are then shrunk by 3 cells (90 meters), regrouped, and those patches greater than or equal to 5000 acres are extracted. To eliminate small connectors, the patches are further shrunk by 3 cells (90 meters) grouped, and patches greater than or equal to 5000 acres are extracted, then 3 cells (90 meters) added back to the patches. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than 5000 acres extracted. Nat2nlp_c1 Michigan Northern Lower Peninsula natural vegetation classes, patches greater than 2500 acres after buffering inward 90 meters. All natural vegetation classes, including water, are grouped together. Patches greater than or equal to 2500 acres are extracted. These patches are then shrunk by 3 cells (90 meters), regrouped, and those A - 58 Appendix I - natural vegetation core area datalayers and descriptions - continued patches greater than or equal to 2500 acres are extracted. To eliminate small connectors, the patches are further shrunk by 3 cells (90 meters) grouped, and patches greater than or equal to 2500 acres are extracted, then 3 cells (90 meters) added back to the patches. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than 2500 acres extracted. Nat2slp_c1 Michigan Southern Lower Peninsula natural vegetation classes, patches greater than 500 acres after buffering inward 90 meters. All natural vegetation classes, including water, are grouped together. Patches greater than or equal to 500 acres are extracted. These patches are then shrunk by 3 cells (90 meters), regrouped, and those patches greater than or equal to 500 acres are extracted. To eliminate small connectors, the patches are further shrunk by 3 cells (90 meters) grouped, and patches greater than or equal to 500 acres are extracted, then 3 cells (90 meters) added back to the patches. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than 500 acres extracted. Nat2jup_c1 Michigan Upper Peninsula natural vegetation classes, patches greater than 5000 acres after cutting by major roads and buffering inward 90 meters. All natural vegetation classes, including water, are grouped together. Major roads are converted to raster and then removed from the natural vegetation classes. Patches are re-grouped and those greater than or equal to 5000 acres are extracted. These patches are then shrunk by 3 cells (90 meters), regrouped, and those patches greater than or equal to 5000 acres are extracted. To eliminate small connectors, the patches are further shrunk by 3 cells (90 meters) grouped, and patches greater than or equal to 5000 acres are extracted, then 3 cells (90 meters) added back to the patches. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than 5000 acres extracted. Nat2jnlp_c1 Michigan Northern Lower Peninsula natural vegetation classes, patches greater than 2500 acres after cutting by major roads and buffering inward 90 meters. All natural vegetation classes, including water, are grouped together. Major roads are converted to raster and then removed from the natural vegetation classes. Patches are re-grouped and those greater than or equal to 2500 acres are extracted. These patches are then shrunk by 3 cells (90 meters), regrouped, and those patches greater than or equal to 2500 acres are extracted. To eliminate small connectors, the patches are further shrunk by 3 cells (90 meters) grouped, and patches greater than or equal to 2500 acres are extracted, then 3 cells (90 meters) added back to the patches. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than 2500 acres extracted. Nat2jslp_c1 Michigan Southern Lower Peninsula natural vegetation classes, patches greater than 500 acres after cutting by major roads and buffering inward 90 meters. A - 59 Appendix I - natural vegetation core area datalayers and descriptions - continued All natural vegetation classes, including water, are grouped together. Major roads are converted to raster and then removed from the natural vegetation classes. Patches are re-grouped and those greater than or equal to 500 acres are extracted. These patches are then shrunk by 3 cells (90 meters), regrouped, and those patches greater than or equal to 500 acres are extracted. To eliminate small connectors, the patches are further shrunk by 3 cells (90 meters) grouped, and patches greater than or equal to 500 acres are extracted, then 3 cells (90 meters) added back to the patches. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than 500 acres extracted. Nat2mup_c1 Michigan Upper Peninsula natural vegetation classes, patches greater than 5000 acres after cutting by all roads and buffering inward 90 meters. All natural vegetation classes, including water, are grouped together. All roads are converted to raster and then removed from the natural vegetation classes. Patches are re-grouped and those greater than or equal to 5000 acres are extracted. These patches are then shrunk by 3 cells (90 meters), regrouped, and those patches greater than or equal to 5000 acres are extracted. To eliminate small connectors, the patches are further shrunk by 3 cells (90 meters) grouped, and patches greater than or equal to 5000 acres are extracted, then 3 cells (90 meters) added back to the patches. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than 5000 acres extracted. Nat2mnlp_c1 Michigan Northern Lower Peninsula natural vegetation classes, patches greater than 2500 acres after cutting by all roads and buffering inward 90 meters. All natural vegetation classes, including water, are grouped together. All roads are converted to raster and then removed from the natural vegetation classes. Patches are re-grouped and those greater than or equal to 2500 acres are extracted. These patches are then shrunk by 3 cells (90 meters), regrouped, and those patches greater than or equal to 2500 acres are extracted. To eliminate small connectors, the patches are further shrunk by 3 cells (90 meters) grouped, and patches greater than or equal to 2500 acres are extracted, then 3 cells (90 meters) added back to the patches. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than 2500 acres extracted. Nat2mslp_c1 Michigan Southern Lower Peninsula natural vegetation classes, patches greater than 500 acres after cutting by all roads and buffering inward 90 meters. All natural vegetation classes, including water, are grouped together. All roads are converted to raster and then removed from the natural vegetation classes. Patches are re-grouped and those greater than or equal to 500 acres are extracted. These patches are then shrunk by 3 cells (90 meters), regrouped, and those patches greater than or equal to 500 acres are extracted. To eliminate small connectors, the patches are further shrunk by 3 cells (90 meters) grouped, and patches greater than or equal to 500 acres are extracted, then 3 cells (90 meters) added back to the patches. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than 500 acres extracted. A - 60 Appendix I - natural vegetation core area datalayers and descriptions - continued Nat2up_c2 Michigan Upper Peninsula natural vegetation classes, patches greater than 5000 acres after buffering inward 210 meters. All natural vegetation classes, including water, are grouped together. Patches greater than or equal to 5000 acres are extracted. These patches are then shrunk by 3 cells (90 meters), regrouped, and those patches greater than or equal to 5000 acres are extracted. To eliminate small connectors, the patches are further shrunk by 3 cells (90 meters) grouped, and patches greater than or equal to 5000 acres are extracted, then 3 cells (90 meters) added back to the patches. These patches are shrunk a further four cells (120 meters, 210 meters total) and patches greater than or equal to 5000 acres are extracted. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than 5000 acres extracted. Nat2nlp_c2 Michigan Northern Lower Peninsula natural vegetation classes, patches greater than 2500 acres after buffering inward 210 meters. All natural vegetation classes, including water, are grouped together. Patches greater than or equal to 2500 acres are extracted. These patches are then shrunk by 3 cells (90 meters), regrouped, and those patches greater than or equal to 2500 acres are extracted. To eliminate small connectors, the patches are further shrunk by 3 cells (90 meters) grouped, and patches greater than or equal to 2500 acres are extracted, then 3 cells (90 meters) added back to the patches. These patches are shrunk a further four cells (120 meters, 210 meters total) and patches greater than or equal to 2500 acres are extracted. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than 2500 acres extracted. Nat2slp_c2 Michigan Southern Lower Peninsula natural vegetation classes, patches greater than 500 acres after buffering inward 210 meters. All natural vegetation classes, including water, are grouped together. Patches greater than or equal to 500 acres are extracted. These patches are then shrunk by 3 cells (90 meters), regrouped, and those patches greater than or equal to 500 acres are extracted. To eliminate small connectors, the patches are further shrunk by 3 cells (90 meters) grouped, and patches greater than or equal to 500 acres are extracted, then 3 cells (90 meters) added back to the patches. These patches are shrunk a further four cells (120 meters, 210 meters total) and patches greater than or equal to 500 acres are extracted. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than 500 acres extracted. Nat2jup_c2 Michigan Upper Peninsula natural vegetation classes, patches greater than 5000 acres after cutting by major roads and buffering inward 210 meters. All natural vegetation classes, including water, are grouped together. Major roads are converted to raster and then removed from the natural vegetation classes. Patches are re-grouped and those greater than or equal to 5000 acres are extracted. These patches are then shrunk by 3 cells (90 meters), A - 61 Appendix I - natural vegetation core area datalayers and descriptions - continued regrouped, and those patches greater than or equal to 5000 acres are extracted. To eliminate small connectors, the patches are further shrunk by 3 cells (90 meters) grouped, and patches greater than or equal to 5000 acres are extracted, then 3 cells (90 meters) added back to the patches. These patches are shrunk a further four cells (120 meters, 210 meters total) and patches greater than or equal to 5000 acres are extracted. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than 5000 acres extracted. Nat2jnlp_c2 Michigan Northern Lower Peninsula natural vegetation classes, patches greater than 2500 acres after cutting by major roads and buffering inward 210 meters. All natural vegetation classes, including water, are grouped together. Major roads are converted to raster and then removed from the natural vegetation classes. Patches are re-grouped and those greater than or equal to 2500 acres are extracted. These patches are then shrunk by 3 cells (90 meters), regrouped, and those patches greater than or equal to 2500 acres are extracted. To eliminate small connectors, the patches are further shrunk by 3 cells (90 meters) grouped, and patches greater than or equal to 2500 acres are extracted, then 3 cells (90 meters) added back to the patches. These patches are shrunk a further four cells (120 meters, 210 meters total) and patches greater than or equal to 2500 acres are extracted. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than 2500 acres extracted. Nat2jslp_c2 Michigan Southern Lower Peninsula natural vegetation classes, patches greater than 500 acres after cutting by major roads and buffering inward 210 meters. All natural vegetation classes, including water, are grouped together. Major roads are converted to raster and then removed from the natural vegetation classes. Patches are re-grouped and those greater than or equal to 500 acres are extracted. These patches are then shrunk by 3 cells (90 meters), regrouped, and those patches greater than or equal to 500 acres are extracted. To eliminate small connectors, the patches are further shrunk by 3 cells (90 meters) grouped, and patches greater than or equal to 500 acres are extracted, then 3 cells (90 meters) added back to the patches. These patches are shrunk a further four cells (120 meters, 210 meters total) and patches greater than or equal to 500 acres are extracted. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than 500 acres extracted. Nat2mup_c2 Michigan Upper Peninsula natural vegetation classes, patches greater than 5000 acres after cutting by all roads and buffering inward 210 meters. All natural vegetation classes, including water, are grouped together. All roads are converted to raster and then removed from the natural vegetation classes. Patches are re-grouped and those greater than or equal to 5000 acres are extracted. These patches are then shrunk by 3 cells (90 meters), regrouped, and those patches greater than or equal to 5000 acres are extracted. To eliminate small connectors, the patches are further shrunk by 3 cells (90 meters) grouped, and patches greater than or equal to 5000 acres are extracted, then 3 cells (90 meters) added back to the patches. These patches are shrunk a further four cells (120 meters, 210 meters total) and patches greater than or equal to 5000 A - 62 Appendix I - natural vegetation core area datalayers and descriptions - continued acres are extracted. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than 5000 acres extracted. Nat2mnlp_c2 Michigan Northern Lower Peninsula natural vegetation classes, patches greater than 2500 acres after cutting by all roads and buffering inward 210 meters. All natural vegetation classes, including water, are grouped together. All roads are converted to raster and then removed from the natural vegetation classes. Patches are re-grouped and those greater than or equal to 2500 acres are extracted. These patches are then shrunk by 3 cells (90 meters), regrouped, and those patches greater than or equal to 2500 acres are extracted. To eliminate small connectors, the patches are further shrunk by 3 cells (90 meters) grouped, and patches greater than or equal to 2500 acres are extracted, then 3 cells (90 meters) added back to the patches. These patches are shrunk a further four cells (120 meters, 210 meters total) and patches greater than or equal to 2500 acres are extracted. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than 2500 acres extracted. Nat2mslp_c2 Michigan Southern Lower Peninsula natural vegetation classes, patches greater than 500 acres after cutting by all roads and buffering inward 210 meters. All natural vegetation classes, including water, are grouped together. All roads are converted to raster and then removed from the natural vegetation classes. Patches are re-grouped and those greater than or equal to 500 acres are extracted. These patches are then shrunk by 3 cells (90 meters), regrouped, and those patches greater than or equal to 500 acres are extracted. To eliminate small connectors, the patches are further shrunk by 3 cells (90 meters) grouped, and patches greater than or equal to 500 acres are extracted, then 3 cells (90 meters) added back to the patches. These patches are shrunk a further four cells (120 meters, 210 meters total) and patches greater than or equal to 500 acres are extracted. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than 500 acres extracted. Nat2up_c3 Michigan Upper Peninsula natural vegetation classes, patches greater than 5000 acres after buffering inward 300 meters. All natural vegetation classes, including water, are grouped together. Patches greater than or equal to 5000 acres are extracted. These patches are then shrunk by 3 cells (90 meters), regrouped, and those patches greater than or equal to 5000 acres are extracted. To eliminate small connectors, the patches are further shrunk by 3 cells (90 meters) grouped, and patches greater than or equal to 5000 acres are extracted, then 3 cells (90 meters) added back to the patches. These patches are shrunk a further four cells (120 meters, 210 meters total) and patches greater than or equal to 5000 acres are extracted. The remaining patches are then shrunk a further 3 cells (90 meters, 300 meters total) and all patches greater than or equal to 5000 acres are extracted. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than 500 acres extracted. A - 63 Appendix I - natural vegetation core area datalayers and descriptions - continued Nat2nlp_c3 Michigan Northern Lower Peninsula natural vegetation classes, patches greater than 2500 acres after buffering inward 300 meters. All natural vegetation classes, including water, are grouped together. Patches greater than or equal to 2500 acres are extracted. These patches are then shrunk by 3 cells (90 meters), regrouped, and those patches greater than or equal to 2500 acres are extracted. To eliminate small connectors, the patches are further shrunk by 3 cells (90 meters) grouped, and patches greater than or equal to 2500 acres are extracted, then 3 cells (90 meters) added back to the patches. These patches are shrunk a further four cells (120 meters, 210 meters total) and patches greater than or equal to 2500 acres are extracted. The remaining patches are then shrunk a further 3 cells (90 meters, 300 meters total) and all patches greater than or equal to 2500 acres are extracted. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than 2500 acres extracted. Nat2slp_c3 Michigan Southern Lower Peninsula natural vegetation classes, patches greater than 500 acres after buffering inward 300 meters. All natural vegetation classes, including water, are grouped together. Patches greater than or equal to 500 acres are extracted. These patches are then shrunk by 3 cells (90 meters), regrouped, and those patches greater than or equal to 500 acres are extracted. To eliminate small connectors, the patches are further shrunk by 3 cells (90 meters) grouped, and patches greater than or equal to 500 acres are extracted, then 3 cells (90 meters) added back to the patches. These patches are shrunk a further four cells (120 meters, 210 meters total) and patches greater than or equal to 500 acres are extracted. The remaining patches are then shrunk a further 3 cells (90 meters, 300 meters total) and all patches greater than or equal to 500 acres are extracted. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than 500 acres extracted. Nat2jup_c3 Michigan Upper Peninsula natural vegetation classes, patches greater than 5000 acres after cutting by major roads and buffering inward 300 meters. All natural vegetation classes, including water, are grouped together. Major roads are converted to raster and then removed from the natural vegetation classes. Patches are re-grouped and those greater than or equal to 5000 acres are extracted. These patches are then shrunk by 3 cells (90 meters), regrouped, and those patches greater than or equal to 5000 acres are extracted. To eliminate small connectors, the patches are further shrunk by 3 cells (90 meters) grouped, and patches greater than or equal to 5000 acres are extracted, then 3 cells (90 meters) added back to the patches. These patches are shrunk a further four cells (120 meters, 210 meters total) and patches greater than or equal to 5000 acres are extracted. The remaining patches are then shrunk a further 3 cells (90 meters, 300 meters total) and all patches greater than or equal to 5000 acres are extracted. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than 5000 acres extracted. Nat2jnlp_c3 Michigan Northern Lower Peninsula natural vegetation classes, patches greater than 2500 acres after cutting by major roads and buffering inward 300 meters. A - 64 Appendix I - natural vegetation core area datalayers and descriptions - continued All natural vegetation classes, including water, are grouped together. Major roads are converted to raster and then removed from the natural vegetation classes. Patches are re-grouped and those greater than or equal to 2500 acres are extracted. These patches are then shrunk by 3 cells (90 meters), regrouped, and those patches greater than or equal to 2500 acres are extracted. To eliminate small connectors, the patches are further shrunk by 3 cells (90 meters) grouped, and patches greater than or equal to 2500 acres are extracted, then 3 cells (90 meters) added back to the patches. These patches are shrunk a further four cells (120 meters, 210 meters total) and patches greater than or equal to 2500 acres are extracted. The remaining patches are then shrunk a further 3 cells (90 meters, 300 meters total) and all patches greater than or equal to 2500 acres are extracted. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than 2500 acres extracted. Nat2jslp_c3 Michigan Southern Lower Peninsula natural vegetation classes, patches greater than 500 acres after cutting by major roads and buffering inward 300 meters. All natural vegetation classes, including water, are grouped together. Major roads are converted to raster and then removed from the natural vegetation classes. Patches are re-grouped and those greater than or equal to 500 acres are extracted. These patches are then shrunk by 3 cells (90 meters), regrouped, and those patches greater than or equal to 500 acres are extracted. To eliminate small connectors, the patches are further shrunk by 3 cells (90 meters) grouped, and patches greater than or equal to 500 acres are extracted, then 3 cells (90 meters) added back to the patches. These patches are shrunk a further four cells (120 meters, 210 meters total) and patches greater than or equal to 500 acres are extracted. The remaining patches are then shrunk a further 3 cells (90 meters, 300 meters total) and all patches greater than or equal to 500 acres are extracted. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than 500 acres extracted. Nat2mup_c3 Michigan Upper Peninsula natural vegetation classes, patches greater than 5000 acres after cutting by all roads and buffering inward 300 meters. All natural vegetation classes, including water, are grouped together. All roads are converted to raster and then removed from the natural vegetation classes. Patches are re-grouped and those greater than or equal to 5000 acres are extracted. These patches are then shrunk by 3 cells (90 meters), regrouped, and those patches greater than or equal to 5000 acres are extracted. To eliminate small connectors, the patches are further shrunk by 3 cells (90 meters) grouped, and patches greater than or equal to 5000 acres are extracted, then 3 cells (90 meters) added back to the patches. These patches are shrunk a further four cells (120 meters, 210 meters total) and patches greater than or equal to 5000 acres are extracted. The remaining patches are then shrunk a further 3 cells (90 meters, 300 meters total) and all patches greater than or equal to 5000 acres are extracted. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than 5000 acres extracted. Nat2mnlp_c3 Michigan Northern Lower Peninsula natural vegetation classes, patches greater than 2500 acres after cutting by all roads and buffering inward 300 meters. A - 65 Appendix I - natural vegetation core area datalayers and descriptions - continued All natural vegetation classes, including water, are grouped together. All roads are converted to raster and then removed from the natural vegetation classes. Patches are re-grouped and those greater than or equal to 2500 acres are extracted. These patches are then shrunk by 3 cells (90 meters), regrouped, and those patches greater than or equal to 2500 acres are extracted. To eliminate small connectors, the patches are further shrunk by 3 cells (90 meters) grouped, and patches greater than or equal to 2500 acres are extracted, then 3 cells (90 meters) added back to the patches. These patches are shrunk a further four cells (120 meters, 210 meters total) and patches greater than or equal to 2500 acres are extracted. The remaining patches are then shrunk a further 3 cells (90 meters, 300 meters total) and all patches greater than or equal to 2500 acres are extracted. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than 2500 acres extracted. Nat2mslp_c3 Michigan Southern Lower Peninsula natural vegetation classes, patches greater than 500 acres after cutting by all roads and buffering inward 300 meters. All natural vegetation classes, including water, are grouped together. All roads are converted to raster and then removed from the natural vegetation classes. Patches are re-grouped and those greater than or equal to 500 acres are extracted. These patches are then shrunk by 3 cells (90 meters), regrouped, and those patches greater than or equal to 500 acres are extracted. To eliminate small connectors, the patches are further shrunk by 3 cells (90 meters) grouped, and patches greater than or equal to 500 acres are extracted, then 3 cells (90 meters) added back to the patches. These patches are shrunk a further four cells (120 meters, 210 meters total) and patches greater than or equal to 500 acres are extracted. The remaining patches are then shrunk a further 3 cells (90 meters, 300 meters total) and all patches greater than or equal to 500 acres are extracted. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than 500 acres extracted. Merged core areas Nat2 Natural vegetation patches with the patch size dependent on ecoregion (Michigan Upper Peninsula, Northern Lower Peninsula, or Southern Lower Peninsula). All natural vegetation classes, including water, are grouped together. Three raster datasets are created, one for the Upper Peninsula, one for the Northern Lower Peninsula, and one for the Southern Lower Peninsula. For each of the three raster datasets, patches greater than or equal to a threshold (UP 5000 acres, NLP 2500 acres, SLP 500 acres) are extracted. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than the criteria are extracted. The three raster datasets are then merged into one statewide raster. Nat2j Natural vegetation patches with the patch size dependent on ecoregion (Michigan Upper Peninsula, Northern Lower Peninsula, or Southern Lower Peninsula) after the patches are cut by major roads. All natural vegetation classes, including water, are grouped together. Major roads are converted to raster and then removed from the natural vegetation classes. Three raster datasets are created, one for the Upper Peninsula, one for the Northern Lower Peninsula, and one for the Southern Lower A - 66 Appendix I - natural vegetation core area datalayers and descriptions - continued Peninsula. For each of the three raster datasets, patches greater than or equal to a threshold (UP 5000 acres, NLP 2500 acres, SLP 500 acres) are extracted. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than the criteria are extracted. The three raster datasets are then merged into one statewide raster. Nat2m Natural vegetation patches with the patch size dependent on ecoregion (Michigan Upper Peninsula, Northern Lower Peninsula, or Southern Lower Peninsula) after the patches are cut by all roads. All natural vegetation classes, including water, are grouped together. All roads are converted to raster and then removed from the natural vegetation classes. Three raster datasets are created, one for the Upper Peninsula, one for the Northern Lower Peninsula, and one for the Southern Lower Peninsula. For each of the three raster datasets, patches greater than or equal to a threshold (UP 5000 acres, NLP 2500 acres, SLP 500 acres) are extracted. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than the criteria are extracted. The three raster datasets are then merged into one statewide raster. Nat2_c Natural vegetation patches with the patch size dependent on ecoregion (Michigan Upper Peninsula, Northern Lower Peninsula, or Southern Lower Peninsula), after buffering inward 90 meters. All natural vegetation classes, including water, are grouped together. Three raster datasets are created, one for the Upper Peninsula, one for the Northern Lower Peninsula, and one for the Southern Lower Peninsula. For each of the three raster datasets, patches greater than or equal to a threshold (UP 5000 acres, NLP 2500 acres, SLP 500 acres) are extracted. These patches are then shrunk by 3 cells (90 meters), regrouped, and those patches greater than or equal to the ecoregional thresholds are extracted. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those greater than or equal to the ecoregional thresholds are extracted. The three raster datasets are then merged into one statewide raster. Nat2j_c Natural vegetation patches with the patch size dependent on ecoregion (Michigan Upper Peninsula, Northern Lower Peninsula, or Southern Lower Peninsula), after cutting by major roads and buffering inward 90 meters. All natural vegetation classes, including water, are grouped together. Major roads are converted to raster and then removed from the natural vegetation classes. Three raster datasets are created, one for the Upper Peninsula, one for the Northern Lower Peninsula, and one for the Southern Lower Peninsula. For each of the three raster datasets, patches greater than or equal to a threshold (UP 5000 acres, NLP 2500 acres, SLP 500 acres) are extracted. These patches are then shrunk by 3 cells (90 meters), regrouped, and those patches greater than or equal to the ecoregional thresholds are extracted. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those greater than or equal to the ecoregional thresholds are extracted. The three raster datasets are then merged into one statewide raster. A - 67 Appendix I - natural vegetation core area datalayers and descriptions - continued Nat2m_c Natural vegetation patches with the patch size dependent on ecoregion (Michigan Upper Peninsula, Northern Lower Peninsula, or Southern Lower Peninsula), after cutting by all roads and buffering inward 90 meters. All natural vegetation classes, including water, are grouped together. All roads are converted to raster and then removed from the natural vegetation classes. Three raster datasets are created, one for the Upper Peninsula, one for the Northern Lower Peninsula, and one for the Southern Lower Peninsula. For each of the three raster datasets, patches greater than or equal to a threshold (UP 5000 acres, NLP 2500 acres, SLP 500 acres) are extracted. These patches are then shrunk by 3 cells (90 meters), regrouped, and those patches greater than or equal to the ecoregional thresholds are extracted. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those greater than or equal to the ecoregional thresholds are extracted. The three raster datasets are then merged into one statewide raster. Nat2_c1 Natural vegetation patches with the patch size dependent on ecoregion (Michigan Upper Peninsula, Northern Lower Peninsula, or Southern Lower Peninsula), after buffering inward 90 meters. All natural vegetation classes, including water, are grouped together. Three raster datasets are created, one for the Upper Peninsula, one for the Northern Lower Peninsula, and one for the Southern Lower Peninsula. For each of the three raster datasets patches greater than or equal to a threshold (UP 5000 acres, NLP 2500 acres, SLP 500 acres) are extracted. These patches are then shrunk by 3 cells (90 meters), regrouped, and those patches greater than or equal to the ecoregional threshold are extracted. To eliminate small connectors, the patches are further shrunk by 3 cells (90 meters) grouped, and patches greater than or equal to the ecoregional threshold are extracted, then 3 cells (90 meters) added back to the patches. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than the ecoregional threshold extracted. The three raster datasets are then merged into one statewide raster. Nat2j_c1 Natural vegetation patches with the patch size dependent on ecoregion (Michigan Upper Peninsula, Northern Lower Peninsula, or Southern Lower Peninsula), after cutting by major roads and buffering inward 90 meters. All natural vegetation classes, including water, are grouped together. Major roads are converted to raster and then removed from the natural vegetation classes. Three raster datasets are created, one for the Upper Peninsula, one for the Northern Lower Peninsula, and one for the Southern Lower Peninsula. For each of the three raster datasets patches greater than or equal to a threshold (UP 5000 acres, NLP 2500 acres, SLP 500 acres) are extracted. These patches are then shrunk by 3 cells (90 meters), regrouped, and those patches greater than or equal to the ecoregional threshold are extracted. To eliminate small connectors, the patches are further shrunk by 3 cells (90 meters) grouped, and patches greater than or equal to the ecoregional threshold are extracted, then 3 cells (90 meters) added back to the patches. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than the ecoregional threshold extracted. The three raster datasets are then merged into one statewide raster. A - 68 Appendix I - natural vegetation core area datalayers and descriptions - continued Nat2m_c1 Natural vegetation patches with the patch size dependent on ecoregion (Michigan Upper Peninsula, Northern Lower Peninsula, or Southern Lower Peninsula), after cutting by all roads and buffering inward 90 meters. All natural vegetation classes, including water, are grouped together. All roads are converted to raster and then removed from the natural vegetation classes. Three raster datasets are created, one for the Upper Peninsula, one for the Northern Lower Peninsula, and one for the Southern Lower Peninsula. For each of the three raster datasets patches greater than or equal to a threshold (UP 5000 acres, NLP 2500 acres, SLP 500 acres) are extracted. These patches are then shrunk by 3 cells (90 meters), regrouped, and those patches greater than or equal to the ecoregional threshold are extracted. To eliminate small connectors, the patches are further shrunk by 3 cells (90 meters) grouped, and patches greater than or equal to the ecoregional threshold are extracted, then 3 cells (90 meters) added back to the patches. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than the ecoregional threshold extracted. The three raster datasets are then merged into one statewide raster. Nat2_c2 Natural vegetation patches with the patch size dependent on ecoregion (Michigan Upper Peninsula, Northern Lower Peninsula, or Southern Lower Peninsula), after buffering inward 210 meters. All natural vegetation classes, including water, are grouped together. Three raster datasets are created, one for the Upper Peninsula, one for the Northern Lower Peninsula, and one for the Southern Lower Peninsula. For each of the three raster datasets patches greater than or equal to a threshold (UP 5000 acres, NLP 2500 acres, SLP 500 acres) are extracted. These patches are then shrunk by 3 cells (90 meters), regrouped, and those patches greater than or equal to the ecoregional threshold are extracted. To eliminate small connectors, the patches are further shrunk by 3 cells (90 meters) grouped, and patches greater than or equal to the ecoregional threshold are extracted, then 3 cells (90 meters) added back to the patches. These patches are shrunk a further four cells (120 meters, 210 meters total) and patches greater than or equal to the ecoregional threshold are extracted. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than the ecoregional threshold extracted. The three raster datasets are then merged into one statewide raster. Nat2j_c2 Natural vegetation patches with the patch size dependent on ecoregion (Michigan Upper Peninsula, Northern Lower Peninsula, or Southern Lower Peninsula), after cutting by major roads and buffering inward 210 meters. All natural vegetation classes, including water, are grouped together. Major roads are converted to raster and then removed from the natural vegetation classes. Three raster datasets are created, one for the Upper Peninsula, one for the Northern Lower Peninsula, and one for the Southern Lower Peninsula. For each of the three raster datasets patches greater than or equal to a threshold (UP 5000 acres, NLP 2500 acres, SLP 500 acres) are extracted. These patches are then shrunk by 3 cells (90 meters), regrouped, and those patches greater than or equal to the ecoregional threshold are extracted. To eliminate small connectors, the patches are further shrunk by 3 cells (90 meters) grouped, and patches greater than or equal to the ecoregional threshold are extracted, then 3 cells (90 meters) added back to the patches. These patches are shrunk a further four cells (120 meters, 210 A - 69 Appendix I - natural vegetation core area datalayers and descriptions - continued meters total) and patches greater than or equal to the ecoregional threshold are extracted. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than the ecoregional threshold extracted. The three raster datasets are then merged into one statewide raster. Nat2m_c2 Natural vegetation patches with the patch size dependent on ecoregion (Michigan Upper Peninsula, Northern Lower Peninsula, or Southern Lower Peninsula), after cutting by all roads and buffering inward 210 meters. All natural vegetation classes, including water, are grouped together. All roads are converted to raster and then removed from the natural vegetation classes. Three raster datasets are created, one for the Upper Peninsula, one for the Northern Lower Peninsula, and one for the Southern Lower Peninsula. For each of the three raster datasets patches greater than or equal to a threshold (UP 5000 acres, NLP 2500 acres, SLP 500 acres) are extracted. These patches are then shrunk by 3 cells (90 meters), regrouped, and those patches greater than or equal to the ecoregional threshold are extracted. To eliminate small connectors, the patches are further shrunk by 3 cells (90 meters) grouped, and patches greater than or equal to the ecoregional threshold are extracted, then 3 cells (90 meters) added back to the patches. These patches are shrunk a further four cells (120 meters, 210 meters total) and patches greater than or equal to the ecoregional threshold are extracted. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than the ecoregional threshold extracted. The three raster datasets are then merged into one statewide raster. Nat2_c3 Natural vegetation patches with the patch size dependent on ecoregion (Michigan Upper Peninsula, Northern Lower Peninsula, or Southern Lower Peninsula), after buffering inward 300 meters. All natural vegetation classes, including water, are grouped together. Three raster datasets are created, one for the Upper Peninsula, one for the Northern Lower Peninsula, and one for the Southern Lower Peninsula. For each of the three raster datasets patches greater than or equal to a threshold (UP 5000 acres, NLP 2500 acres, SLP 500 acres) are extracted. These patches are then shrunk by 3 cells (90 meters), regrouped, and those patches greater than or equal to the ecoregional threshold are extracted. To eliminate small connectors, the patches are further shrunk by 3 cells (90 meters) grouped, and patches greater than or equal to the ecoregional threshold are extracted, then 3 cells (90 meters) added back to the patches. These patches are shrunk a further four cells (120 meters, 210 meters total) and patches greater than or equal to the ecoregional threshold are extracted. The remaining patches are then shrunk a further 3 cells (90 meters, 300 meters total) and all patches greater than or equal to the ecoregional threshold are extracted. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than the ecoregional threshold extracted. The three raster datasets are then merged into one statewide raster. Nat2j_c3 Natural vegetation patches with the patch size dependent on ecoregion (Michigan Upper Peninsula, Northern Lower Peninsula, or Southern Lower Peninsula), after cutting by major roads and buffering inward 300 meters. A - 70 Appendix I - natural vegetation core area datalayers and descriptions - continued All natural vegetation classes, including water, are grouped together. Major roads are converted to raster and then removed from the natural vegetation classes. Three raster datasets are created, one for the Upper Peninsula, one for the Northern Lower Peninsula, and one for the Southern Lower Peninsula. For each of the three raster datasets patches greater than or equal to a threshold (UP 5000 acres, NLP 2500 acres, SLP 500 acres) are extracted. These patches are then shrunk by 3 cells (90 meters), regrouped, and those patches greater than or equal to the ecoregional threshold are extracted. To eliminate small connectors, the patches are further shrunk by 3 cells (90 meters) grouped, and patches greater than or equal to the ecoregional threshold are extracted, then 3 cells (90 meters) added back to the patches. These patches are shrunk a further four cells (120 meters, 210 meters total) and patches greater than or equal to the ecoregional threshold are extracted. The remaining patches are then shrunk a further 3 cells (90 meters, 300 meters total) and all patches greater than or equal to the ecoregional threshold are extracted. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than the ecoregional threshold extracted. The three raster datasets are then merged into one statewide raster. Nat2m_c3 Natural vegetation patches with the patch size dependent on ecoregion (Michigan Upper Peninsula, Northern Lower Peninsula, or Southern Lower Peninsula), after cutting by all roads and buffering inward 300 meters. All natural vegetation classes, including water, are grouped together. All roads are converted to raster and then removed from the natural vegetation classes. Three raster datasets are created, one for the Upper Peninsula, one for the Northern Lower Peninsula, and one for the Southern Lower Peninsula. For each of the three raster datasets patches greater than or equal to a threshold (UP 5000 acres, NLP 2500 acres, SLP 500 acres) are extracted. These patches are then shrunk by 3 cells (90 meters), regrouped, and those patches greater than or equal to the ecoregional threshold are extracted. To eliminate small connectors, the patches are further shrunk by 3 cells (90 meters) grouped, and patches greater than or equal to the ecoregional threshold are extracted, then 3 cells (90 meters) added back to the patches. These patches are shrunk a further four cells (120 meters, 210 meters total) and patches greater than or equal to the ecoregional threshold are extracted. The remaining patches are then shrunk a further 3 cells (90 meters, 300 meters total) and all patches greater than or equal to the ecoregional threshold are extracted. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than the ecoregional threshold extracted. The three raster datasets are then merged into one statewide raster. A - 71 Appendix J - Potentially unchanged natural vegetation core area datalayers and descriptions File Name up_un_core up_un_500 up_un_500_c1 up_unj_500 up_unj_500_c1 up_unm_500 up_unm_500_c1 nlp_un_core nlp_un_250 nlp_un_250_c1 nlp_unj_250 nlp_unj_250_c1 nlp_unm_250 nlp_unm_250_c1 slp_un_core slp_un_50 slp_un_50_c1 slp_unj_50 slp_unj_50_c1 slp_unm_50 slp_unm_50_c1 A - 72 Ecoregion UP UP UP UP UP UP UP NLP NLP NLP NLP NLP NLP NLP SLP SLP SLP SLP SLP SLP SLP road layer none none none major major all all none none none major major all all none none none major major all all minimum buffer size in size (acres) meters 0 0 500 0 500 90 500 0 500 90 500 0 500 90 0 0 250 0 250 90 250 0 250 90 250 0 250 90 0 0 50 0 50 90 50 0 50 90 50 0 50 90 Appendix J - Potentially unchanged natural vegetation core area datalayers and descriptions continued un_core Potentially unchanged vegetation communities. The Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover dataset (Michigan DNR, 2003) and the Circa 1800 Vegetation dataset (Michigan Natural Features Inventory, 1995) are combined. Vegetation communities in the two datasets that are similar to each other are extracted and grouped into patches. un_core_90 Potentially unchanged vegetation communities after buffering inward 90 meters. The Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover dataset (Michigan DNR, 2003) and the Circa 1800 Vegetation dataset (Michigan Natural Features Inventory, 1995) are combined. Vegetation communities in the two datasets that are similar to each other are extracted and grouped into patches. These patches are then shrunk inward by 90 meters (3 cells) and regrouped into patches. unj_core Potentially unchanged vegetation communities after removing major roads. The Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover dataset (Michigan DNR, 2003) and the Circa 1800 Vegetation dataset (Michigan Natural Features Inventory, 1995) are combined. Vegetation communities in the two datasets that are similar to each other are extracted. Major roads from the Michigan Framework are converted to a raster and subtracted from the potentially unchanged vegetation communities. The remaining unchanged vegetation cells are then grouped into patches. unj_core_90 Potentially unchanged vegetation communities after removing major roads and buffering inward 90 meters. The Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover dataset (Michigan DNR, 2003) and the Circa 1800 Vegetation dataset (Michigan Natural Features Inventory, 1995) are combined. Vegetation communities in the two datasets that are similar to each other are extracted. Major roads from the Michigan Framework are converted to a raster and subtracted from the potentially unchanged vegetation communities. The remaining unchanged vegetation cells are then grouped into patches and these patches shrunk by 90 meters (3 cells) and regrouped. unm_core Potentially unchanged vegetation communities after removing major roads. The Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover dataset (Michigan DNR, 2003) and the Circa 1800 Vegetation dataset (Michigan Natural Features Inventory, 1995) are combined. Vegetation communities in the two datasets that are similar to each other are extracted. All roads from the Michigan Framework are converted to a raster and subtracted A - 73 Appendix J - Potentially unchanged natural vegetation core area datalayers and descriptions from the potentially unchanged vegetation communities. The remaining unchanged vegetation cells are then grouped into patches. unm_core_90 Potentially unchanged vegetation communities after removing major roads and buffering inward 90 meters. The Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover dataset (Michigan DNR, 2003) and the Circa 1800 Vegetation dataset (Michigan Natural Features Inventory, 1995) are combined. Vegetation communities in the two datasets that are similar to each other are extracted. All roads from the Michigan Framework are converted to a raster and subtracted from the potentially unchanged vegetation communities. The remaining unchanged vegetation cells are then grouped into patches and these patches shrunk by 90 meters (3 cells) and regrouped. Nlp_un_core Potentially unchanged vegetation communities in Michigan’s Northern Lower Peninsula. The Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover dataset (Michigan DNR, 2003) and the Circa 1800 Vegetation dataset (Michigan Natural Features Inventory, 1995) are combined. Vegetation communities in the two datasets that are similar to each other are extracted and grouped into patches. Nlp_unj_core Potentially unchanged vegetation communities in Michigan’s Northern Lower Peninsula after removing major roads. The Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover dataset (Michigan DNR, 2003) and the Circa 1800 Vegetation dataset (Michigan Natural Features Inventory, 1995) are combined. Vegetation communities in the two datasets that are similar to each other are extracted. Major roads from the Michigan Framework are converted to a raster and subtracted from the potentially unchanged vegetation communities. The remaining unchanged vegetation cells are then grouped into patches. Nlp_unm_core Potentially unchanged vegetation communities in Michigan’s Northern Lower Peninsula after removing all roads. The Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover dataset (Michigan DNR, 2003) and the Circa 1800 Vegetation dataset (Michigan Natural Features Inventory, 1995) are combined. Vegetation communities in the two datasets that are similar to each other are extracted. All roads from the Michigan Framework are converted to a raster and subtracted from the potentially unchanged vegetation communities. The remaining unchanged vegetation cells are then grouped into patches. Slp_un_core Potentially unchanged vegetation communities in Michigan’s Southern Lower Peninsula. A - 74 Appendix J - Potentially unchanged natural vegetation core area datalayers and descriptions The Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover dataset (Michigan DNR, 2003) and the Circa 1800 Vegetation dataset (Michigan Natural Features Inventory, 1995) are combined. Vegetation communities in the two datasets that are similar to each other are extracted and grouped into patches. Slp_unj_core Potentially unchanged vegetation communities in Michigan’s Southern Lower Peninsula after removing major roads. The Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover dataset (Michigan DNR, 2003) and the Circa 1800 Vegetation dataset (Michigan Natural Features Inventory, 1995) are combined. Vegetation communities in the two datasets that are similar to each other are extracted. Major roads from the Michigan Framework are converted to a raster and subtracted from the potentially unchanged vegetation communities. The remaining unchanged vegetation cells are then grouped into patches. Slp_unm_core Potentially unchanged vegetation communities in Michigan’s Southern Lower Peninsula after removing all roads. The Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover dataset (Michigan DNR, 2003) and the Circa 1800 Vegetation dataset (Michigan Natural Features Inventory, 1995) are combined. Vegetation communities in the two datasets that are similar to each other are extracted. All roads from the Michigan Framework are converted to a raster and subtracted from the potentially unchanged vegetation communities. The remaining unchanged vegetation cells are then grouped into patches. Up_un_core Potentially unchanged vegetation communities in Michigan’s Upper Peninsula. The Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover dataset (Michigan DNR, 2003) and the Circa 1800 Vegetation dataset (Michigan Natural Features Inventory, 1995) are combined. Vegetation communities in the two datasets that are similar to each other are extracted and grouped into patches. Up_unj_core Potentially unchanged vegetation communities in Michigan’s Upper Peninsula after removing major roads. The Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover dataset (Michigan DNR, 2003) and the Circa 1800 Vegetation dataset (Michigan Natural Features Inventory, 1995) are combined. Vegetation communities in the two datasets that are similar to each other are extracted. Major roads from the Michigan Framework are converted to a raster and subtracted from the potentially unchanged vegetation communities. The remaining unchanged vegetation cells are then grouped into patches. A - 75 Appendix J - Potentially unchanged natural vegetation core area datalayers and descriptions Up_unm_core Potentially unchanged vegetation communities in Michigan’s Upper Peninsula after removing all roads. The Integrated Forest Monitoring Assessment and Prescription (IFMAP) / GAP Landuse/Landcover dataset (Michigan DNR, 2003) and the Circa 1800 Vegetation dataset (Michigan Natural Features Inventory, 1995) are combined. Vegetation communities in the two datasets that are similar to each other are extracted. All roads from the Michigan Framework are converted to a raster and subtracted from the potentially unchanged vegetation communities. The remaining unchanged vegetation cells are then grouped into patches. A - 76 Appendix K - Matrix vegetation datalayers and descriptions File Name nat2_min nat2_90 nat2_210 nat2_300 nat2j_min nat2j_90 nat2j_210 nat2j_300 nat2m_min nat2m_90 nat2m_210 nat2m_300 Description Matrix vegetation patches statewide water removed Matrix vegetation patches statewide water removed Matrix vegetation patches statewide water removed Matrix vegetation patches statewide water removed Matrix vegetation patches statewide water removed Matrix vegetation patches statewide water removed Matrix vegetation patches statewide water removed Matrix vegetation patches statewide water removed Matrix vegetation patches statewide water removed Matrix vegetation patches statewide water removed Matrix vegetation patches statewide water removed Matrix vegetation patches statewide water removed road layer minimum size (acres) none 5000 none 5000 none 5000 none 5000 major 5000 major 5000 major 5000 major 5000 all 5000 all 5000 all 5000 all 5000 A - 77 Appendix K - Matrix vegetation datalayers and descriptions Nat2gps All natural vegetation classes, including water, are grouped together. Nat2jgps All natural vegetation classes, including water, are grouped together. Major roads are converted to raster and then removed from the natural vegetation classes. Nat2mgps All natural vegetation classes, including water, are grouped together. All roads are converted to raster and then removed from the natural vegetation classes. Matrix vegetation, patch sizes not differentiated by ecoregion. Nat2_min Michigan statewide natural vegetation classes, patches greater than 2000 hectares. All natural vegetation classes, including water, are grouped together. Patches greater than or equal to 2000 hectares are extracted. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than 2000 hectares extracted. Nat2j_min Michigan statewide natural vegetation classes, patches greater than 2000 hectares after cutting by major roads. All natural vegetation classes, including water, are grouped together. Major roads are converted to raster and then removed from the natural vegetation classes. Patches are re-grouped and those greater than or equal to 2000 hectares are extracted. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than 2000 hectares extracted. Nat2m_min Michigan statewide natural vegetation classes, patches greater than 2000 hectares after cutting by major roads. All natural vegetation classes, including water, are grouped together. All roads are converted to raster and then removed from the natural vegetation classes. Patches are re-grouped and those greater than or equal to 2000 hectares are extracted. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than 2000 hectares extracted. Nat2_90 Michigan statewide natural vegetation classes, patches greater than 2000 hectares, after buffering inward 90 meters. All natural vegetation classes, including water, are grouped together. Patches greater than or equal to 2000 hectares are extracted. These patches are shrunk by 3 cells (90 meters) and patches greater than or equal to 2000 hectares extracted. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than 2000 hectares extracted A - 78 Appendix K - Matrix vegetation datalayers and descriptions Nat2j_90 Michigan statewide natural vegetation classes, patches greater than 2000 hectares after cutting by major roads and buffering inward 90 meters. All natural vegetation classes, including water, are grouped together. Major roads are converted to raster and then removed from the natural vegetation classes. Patches are re-grouped and those greater than or equal to 2000 hectares are extracted. These patches are shrunk by 3 cells (90 meters) and patches greater than or equal to 2000 hectares extracted. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than 2000 hectares extracted. Nat2m_90 Michigan statewide natural vegetation classes, patches greater than 2000 hectares after cutting by all roads and buffering inward 90 meters. All natural vegetation classes, including water, are grouped together. All roads are converted to raster and then removed from the natural vegetation classes. Patches are re-grouped and those greater than or equal to 2000 hectares are extracted. These patches are shrunk by 3 cells (90 meters) and patches greater than or equal to 2000 hectares extracted. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than 2000 hectares extracted. Nat2_210 Michigan statewide natural vegetation classes, patches greater than 2000 hectares, after buffering inward 210 meters. All natural vegetation classes, including water, are grouped together. Patches greater than or equal to 2000 hectares are extracted. These patches are shrunk by 7 cells (210 meters) and patches greater than or equal to 2000 hectares extracted. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than 2000 hectares extracted. Nat2j_210 Michigan statewide natural vegetation classes, patches greater than 2000 hectares after cutting by major roads and buffering inward 210 meters. All natural vegetation classes, including water, are grouped together. Major roads are converted to raster and then removed from the natural vegetation classes. Patches are re-grouped and those greater than or equal to 2000 hectares are extracted. These patches are shrunk by 7 cells (210 meters) and patches greater than or equal to 2000 hectares extracted. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than 2000 hectares extracted. Nat2m_210 Michigan statewide natural vegetation classes, patches greater than 2000 hectares after cutting by all roads and buffering inward 210 meters. All natural vegetation classes, including water, are grouped together. All roads are converted to raster and then removed from the natural vegetation classes. Patches are re-grouped and those greater than or equal to 2000 hectares are extracted. These patches are shrunk by 7 cells (210 meters) and patches A - 79 Appendix K - Matrix vegetation datalayers and descriptions greater than or equal to 2000 hectares extracted. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than 2000 hectares extracted. Nat2_300 Michigan statewide natural vegetation classes, patches greater than 2000 hectares, after buffering inward 3000 meters. All natural vegetation classes, including water, are grouped together. Patches greater than or equal to 2000 hectares are extracted. These patches are shrunk by 10 cells (300 meters) and patches greater than or equal to 2000 hectares extracted. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than 2000 hectares extracted. Nat2j_300 Michigan statewide natural vegetation classes, patches greater than 2000 hectares after cutting by major roads and buffering inward 300 meters. All natural vegetation classes, including water, are grouped together. Major roads are converted to raster and then removed from the natural vegetation classes. Patches are re-grouped and those greater than or equal to 2000 hectares are extracted. These patches are shrunk by 10 cells (300 meters) and patches greater than or equal to 2000 hectares extracted. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than 2000 hectares extracted. Nat2m_300 Michigan statewide natural vegetation classes, patches greater than 2000 hectares after cutting by all roads and buffering inward 300 meters. All natural vegetation classes, including water, are grouped together. All roads are converted to raster and then removed from the natural vegetation classes. Patches are re-grouped and those greater than or equal to 2000 hectares are extracted. These patches are shrunk by 10 cells (300 meters) and patches greater than or equal to 2000 hectares extracted. Water bodies greater than ten acres are then removed from the patches, the patches regrouped, and those larger than 2000 hectares extracted. A - 80 Appendix L - EO based datalayers and descriptions File name EO Frequency Ter_EO_trs_0906.shp Ter_EO_trs_0906.shp Ter_EO_trs_0906.shp Ter_EO_trs_0906.shp Ter_EO_trs_0906.shp Ter_EO_trs_0906.shp Aq_EO_trs_0906.shp Aq_EO_trs_0906.shp Aq_EO_trs_0906.shp Aq_EO_trs_0906.shp F_noc F_noc_85 F_ter F_ter_85 F_all F_all_85 F_aq F_aq_85 F_aq_nl Faq85_nl all species - no communities - all dates all species - no communities - only dates > 1985 only terrestrial species - all dates only terrestrial species - only dates > 1985 all element occurrences - all dates all element occurrences - only dates > 1985 only aquatic species - all dates only aquatic species - only dates > 1985 only aquatic species - all dates - no loons only aquatic species - only dates > 1985 - no loons EO Likelihood Ter_EO_trs_0906.shp Ter_EO_trs_0906.shp Ter_EO_trs_0906.shp Ter_EO_trs_0906.shp Ter_EO_trs_0906.shp Ter_EO_trs_0906.shp Aq_EO_trs_0906.shp Aq_EO_trs_0906.shp Aq_EO_trs_0906.shp Aq_EO_trs_0906.shp L_noc L_noc_85 L_ter L_ter_85 L_all L_all_85 L_aq L_aq_85 L_aq_nl Laq85_nl all species - no communities - all dates all species - no communities - only dates > 1985 only terrestrial species - all dates only terrestrial species - only dates > 1985 all element occurrences - all dates all element occurrences - only dates > 1985 only aquatic species - all dates only aquatic species - only dates > 1985 only aquatic species - all dates - no loons only aquatic species - only dates > 1985 - no loons Bio-rarity Ter_EO_trs_0906.shp Ter_EO_trs_0906.shp Ter_EO_trs_0906.shp Ter_EO_trs_0906.shp Ter_EO_trs_0906.shp Ter_EO_trs_0906.shp Aq_EO_trs_0906.shp Aq_EO_trs_0906.shp Aq_EO_trs_0906.shp Aq_EO_trs_0906.shp B_noc B_noc_85 B_ter B_ter_85 B_all B_all_85 B_aq B_aq_85 B_aq_nl Baq85_nl all species - no communities - all dates all species - no communities - only dates > 1985 only terrestrial species - all dates only terrestrial species - only dates > 1985 all element occurrences - all dates all element occurrences - only dates > 1985 only aquatic species - all dates only aquatic species - only dates > 1985 only aquatic species - all dates - no loons only aquatic species - only dates > 1985 - no loons Best 2 occurrences of each species best2_ter_subsubsection_trs_0906.shp best2_ter_subsub_summed_trs_0906.shp best2_aq_watershed_0906.shp Aquatic species richness aq_EO_richness_subwatershed.shp aq_SGCN_richness_subwatershed.shp Field Description best 2 occurrences of each terrestrial species for each sub-subsection the sum of the best 2 occurrences of each terrestrial species by sub-subsection best 2 occurrences of each aquatic species by watershed aquatic rare species richness per river mile by subwatershed aquatic species of greatest conservaton need per river mile by sub-watershed A - 81 Appendix L - EO based datalayers and descriptions - continued File name High quality natural communities natcomm_bcrank.shp natcomm_combined.shp natcomm_state.shp natcomm_section.shp natcomm_subsections.shp natcomm_subsubsection.shp A - 82 Field Description all natural communities with an EO rank > B/C the best 3 occurrences of each natural community type in the state and by section, subsection, and subsubsection the best 3 occurrences of each natural community type in the state the best 3 occurrences of each natural community type for each section (4) the best 3 occurrences of each natural community type for each subsection (22) the best 3 occurrences of each natural community type for each sub-subsection (38) Appendix M - Aquatic datalayers and descriptions File name Rivers mi_subwatersheds.shp vsec_size_temp.shp vsec_gradient.shp vsec_unique_statewide_5pct.shp vsec_unique_statewide_1pct.shp vsec_unique_edu_5pct.shp vsec_unique_edu_1pct.shp vsec_HQ_edu.shp Lakes milakes_conn_shoreline.shp Description michigan subwatersheds River classification framework - one of 2 shapefiles - this one shows the size and water temperature classes used in this report River classification framework - one of 2 shapefiles - this one shows the gradient classes used in this report Potentially unique vsecs statewide using 5% rule Potentially unique vsecs statewide using 1% rule Potentially unique vsecs within an EDU using 5% rule Potentially unique vsecs within an EDU using 1% rule High quality common vsecs by EDU lake_unique_statewide_5pct.shp lake_unique_statewide_1pct.shp lake_unique_EDU_5pct.shp lake_unique_EDU_1pct.shp lake_HQ_edu.shp Lake classification framework - one of 2 shapefiles - this one shows the connectivity and shoreline classes used in this report Lake classification framework - one of 2 shapefiles - this one shows the proximate geology classes used in this report Potentially unique lakes statewide using 5% rule Potentially unique lakes statewide using 1% rule Potentially unique lakes within an EDU using 5% rule Potentially unique lakes within an EDU using 1% rule High quality common lakes by EDU - based on landscape context Subwatersheds pctNat_subwatershed.shp pctNat_Riparian_subwatershed.shp rdXStrCount_subwatershed.shp Percent natural landcover in sub-watershed Percent natural landcover in riparian zones of sub-watershed Number of road and stream crossings per river mile in sub-watershed milakes_proxgeol.shp damCount_subwatershed.shp fragmentation_subwatershed.shp imperv_subwatershed.shp npdesCount_subwatershed.shp mineCounts_subwatershed.shp pollution_subwatershed.shp functional_subwatershed.shp headwaters100natural.shp headwatersPctnatural.shp Number of dams per river mile in sub-watershed Overall fragmentation analysis metric by sub-watersheds Percent impervious suface in sub-watersheds Number of DEQ non-point source pollution permits per river mile in sub-watersheds Number of active mines per river mile in sub-watersheds Overall pollution analysis metric by sub-watersheds Overall functional analysis metric by sub-watersheds 100 percent natural landcover in catchments of headwater (order 1) streams Percent natural landcover in catchments of headwater (order 1) streams A - 83