51 • November 2002: 771–785 Paz-Bermúdez & al. • Typification of lichen names Typification of names of lichen taxa described by G. Sampaio and some others, deposited in Porto Herbarium (PO)* Graciela Paz-Bermúdez 1, Helena Aguiar-Branco & Elisa Folhadela Herbário do Instituto de Botânica “Dr. Gonçalo Sampaio”, Universidade de Porto, Rua do Campo Alegre 1191, 4150-181 Porto, Portugal 1 Permanent address: Departamento de Botánica, Facultade de Bioloxía, Universidade de Santiago, 15706 Santiago de Compostela, Spain. E-mail: bvgraci@usc.es Gonçalo Sampaio authored 103 new nam es of lichens, comprising one new genus, 60 new species, 14 new infraspecific taxa, 25 new com binations and two nomina nova. In addition, he proposed 42 “names” that were not validly published, 18 of these being simple nomina nuda and 21 apparently representing invalid attempts to publish new combinations. The type m aterial is deposited in the Herbarium of University of Porto (PO), except in three cases. Apart from a few specimens from Spain, the material derives from Portugal. For the names of all new taxa the type collections are listed, either a holotype or a lectotype (usually here designated). In addition, a few lichen names authored by M aurice Bouly de Lesdain or William Nylander and based on Portuguese m aterial preserved at PO are listed with indications of type. KEYWORDS: herbarium, lichens, Portugal, Sampaio, Spain, typification. INTRODUCTION The Herbarium of the University of Porto (PO), located in the Institute of Botany “Dr. Gonçalo Sampaio”, is one of the most important on the Iberian Peninsula with about 122,000 specimens belonging to all plant groups. The lichen herbarium includes about 6000 specimens and comprises a general herbarium with specimens from Europe, a Portuguese herbarium with local specimens, and several special collections, the most important being Gonçalo Sampaio’s collection with c. 4000 specimens, mostly from Portugal. Gonçalo António da Silva Ferreira Sampaio (1865–1937), Professor of Botany in the University of Porto, was a renowned investigator mainly through his studies on vascular plants, but he also studied non-vascular groups, mainly lichens (Pires de Lima, 1938). In fact, for over two decades Sampaio collected lichens all over Portugal, mainly in the provinces of Minho, Trásos-Montes e Alto Douro and Douro Litoral. According to our interpretation, Sampaio authored the name of one lichen genus, 87 names of lichen species (60 described as new species), and 14 names of infraspecific taxa. However, it is sometimes difficult to distinguish between new names and new nomenclatural combinations in Sampaio’s papers. This is due to his usual custom of citing only one author for a name even in nomenclatural combinations, viz., the author of the combination. In some cases (especially in Sampaio & Crespí, 1927) the epithet used and the taxonomic relationships expressed allow one to presume that Sampaio intended to make new combinations, so that it is difficult to interpret whether the names should be considered nomina nuda or combinations without basionym author citation. In the simple case in which the presumed new combination appears alone, these have been treated as technically nomina nuda, not being validly published under the provisions of Art. 33.2 of the ICBN (Greuter & al., 2000), even where Sampaio’s intent to publish a new combination seems clear. In a number of cases, however, the situation is complicated by the fact that the presumed new combination has a synonym associated with it. If not treated as a new combination, the name would be validly published as an illegitimate replacement name for the cited synonym, which was clearly not in the least Sampaio’s intention. Art. 33.6, dealing with incorrect citation of a basionym, treats as validly published combinations that would otherwise be validly published either as nomina nova or as names of new taxa, whereas Art. 33.2 specifies that, where no basionym is cited at all, a presumed new combination is validly published if it would otherwise be validly published as the name of a *In memoriam Aurea C. Carvalho (†), our colleague and friend who started this research with us. 771 Paz-Bermúdez & al. • Typification of lichen names new taxon, rather surprisingly making no mention of nomina nova. On the advice of the Nomenclature Editor, and solely for the purposes of this list, we have treated these names, not as heterotypic nomina nova which could be very disturbing nomenclaturally, but as though they were new combinations, albeit not validly published under Art. 33.2. For completeness, and to avoid possible confusion, all names published by Sampaio are listed and an indication given as to the actual authorship of the name, if it is not, in fact, that claimed by Sampaio. Sampaio sometimes used the abbreviation “r.” in naming infraspecific taxa. This is probably an abbreviation for “raça” (race); these infraspecific taxa are technically unranked and are so indicated in the list. Three new species and one variety were described by Sampaio together with the French lichenologist Marcelo Bouly de Lesdain. In addition, some type specimens of lichen names published by other authors were found by us in Sampaio’s herbarium and they are also listed. They comprise two new species described by Marcelo Bouly de Lesdain alone, who received the Portuguese material from G. Sampaio, and nine new species described by William Nylander from specimens sent by Isaac Newton (1840–1906), who was an important Porto Herbarium collector. MATERIAL AND METHODS We have analysed the herbarium specimens used by G. Sampaio, W. Nylander and Bouly de Lesdain comparing them with the original descriptions in order to recognize the holotypes, and when there are more than one original specimen we have designated lectotypes or indicated syntypes. It is important to note the presence of type material in the exsiccata edited and distributed by G. Sampaio: Lichenes de Portugal, Fasc. I-III (nos. 1-300) March 1923; this set of exsiccata is in PO and (according to Stafleu & Cowan, 1985) also in COLO, FH, LD, M and UPS, but Sayre (1969) indicates that 15 sets in all were prepared. The exsiccata have printed labels but no separate catalogue was published. RESULTS In the following lists of names of taxa, we have cited the paper where the original description was published, the Porto herbarium number of the type specimen, and the type category, together with data on the label; we have added the country and the province in brackets. If a taxon received more than one name from Sampaio, that is indicated too. Most of the specimens were collected 772 51 • November 2002: 771–785 and identified by G. Sampaio, who was very careful with his notes. We cite both the paper where the original description was made, and also the page reference for the original description in the compilation of Sampaio´s lichen papers (Sampaio, 1970). Lists of the names of taxa described by G. Sampaio together with B. de Lesdain, by M. Bouly de Lesdain, alone, and by W. Nylander, are presented after the main list. We have used the recent Iberian check-list of the lichens and lichenicolous fungi (Llimona & Hladun, 2001) to indicate the present taxonomic status of the Sampaio taxa. The species names marked with an asterisk (*) are under “Nomina Inquirenda” in that work. Buellia jorgei, Lecania badiella, Lecanora lisbonensis, Lemmopsis affinis, Ramalina portuensis and Rinodina lesdainii are accepted species in Llimona & Hladun (2001). I . List of a ll t a x a n a m ed by S a m pa io, wit h t ypifica tion of th e n a m es of h is n ew ta xa The entries are distinguished typographically as between names for which types were sought (bold italic), validly published names for which selection of a type is not appropriate, e.g., new combinations (italic), and those binary designations (“names”) that were used but not validly published by Sampaio (“italic in double quotes”). For only two names of new taxa described by Sampaio could types not be located, those of Pertusaria lutescens var. conimbricensis and Physma hispanicum. Acarospora alberti Samp., Líqu. inéd.: 5. 15 Dec 1920, as “Alberti”; Sampaio, 1970: 112. – Lectotype (designated here): [Portugal: Minho:] Ponte de Lima: Sá; nas paredes (muito rara), 28 Sep 1920, G. Sampaio (PO 2465L). Syntype: [Portugal: Minho:] in Sampaio: Lich. Portugal nº 115, Ponte de Lima: Sá (paredes), Nov 1920, G. Sampaio (PO 5643L), as “Acarospora lesdaini Harm. var. alberti Samp”. Remarks: = Acarospora smaragdula (Wahlenb.) A. Massal. var. alberti (Samp.) H. Magn. (Llimona & Hladun, 2001). Acarospora flavorubens Bagl. & Car. var. angulosa Samp., Brotéria, Sér. Bot. 15(1): 21. 1 Apr 1917 Sampaio, 1970: 37. – Lectotype (designated here): [Portugal: Douro Litoral:] Foz do Douro, carreiros ao Castelo do Queijo, 24 Feb 1917, G. Sampaio (PO 1316L, isolectotype PO 1316aL). Syntype: [Portugal: Minho:] Ponte de Lima: Sá, nas paredes, 10 Sep 1916, G. Sampaio (PO 1182L). Remarks: (1) = Acarospora lesdainii Harm. ex A. L. Sm. (Sampaio, 1924, 1970: 174). (2) = Acarospora smaragdula (Wahlenb.) A. Massal. subsp. lesdainii 51 • November 2002: 771–785 (Harm. ex A.L. Sm.) Clauzade & Roux (Llimona & Hladun, 2001). *Acarospora granatensis Samp., Líqu. esp. Willkomm: 142. Post May 1917; Sampaio, 1970: 76. – Syntype: [España: Andalucía:] Granada: Arcos do aqueducto da Alhambra, 8 Oct 1844, Willkomm [det. G. Sampaio] (PO 4713L). “Acarospora lesdainii Samp.”, in Sampaio & Crespí, Bol. Real Soc. Esp. Hist. Nat. 27(3): 143. Mar 1927; Sampaio, 1970: 191, nom. nud. = Acarospora lesdainii Harm. ex A. L. Sm. (1918). Remarks: (1) Apparently an error for A. lesdainii Harm. ex A. L. Sm., cited by Sampaio, 1924; 1970: 174. (2) = Acarospora smaragdula (Wahlenb.) A. Massal. subsp. lesdainii (Harm. ex A. L. Sm.) Clauzade & Roux (Llimona & Hladun, 2001). Acarospora lesdainii Harm. ex A. L. Sm. “var. alberti (Samp.) Samp.” in Sampaio, in Sampaio & Crespí, Bol. Real Soc. Esp. Hist. Nat. 27(3): 143. Mar 1927; Sampaio, 1970: 191,, as “Acarospora lesdaini Harm. var. Alberti Samp.” : A. alberti Samp. Remarks: Clearly intended as a new combination based on his earlier A. alberti (Sampaio, 1920); Sampaio had also earlier used the varietal name on Lich. Portugal nº 115 (1920), the type of A. alberti, but as a nomen nudum. Paz-Bermúdez & al. • Typification of lichen names [Portugal: Estremadura:] Lisboa: Arieiro, 6 Dec 1918, G. Sampaio (PO 2449L). Remarks: = Acarospora nodulosa (Dufour) Hue var. reagens (Zahlbr.) Clauzade & Roux (Llimona & Hladun, 2001). Alectoria dichotoma (Hoffm.) Samp., Brotéria, Sér. Bot. 15(3): 128. 1 Dec 1917, as “Alectoria dichotoma (Usnea dichotoma Hoff.; Alectoria sarmentosa Ach.) var. variegata nob.”; Sampaio, 1970: 48: Usnea dichotoma Hoffm. Alectoria dichotoma (Hoffm.) Samp. var. variegata Samp., Brotéria, Sér. Bot. 15(3): 128. 1 Dec 1917, as “Alectoria dichotoma (Usnea dichotoma Hoff.; Alectoria sarmentosa Ach.) var. variegata nob.”; Sampaio, 1970: 48. – Holotype: [Portugal: Beira Alta:] Serra da Estrela (“Estrêla” in protologue), nos penedos, perto do Sanatório de Manteigas, 6 Aug 1916, G. Sampaio (PO 1009L). Remarks: = Alectoria ochroleuca (Hoffm.) A. Massal. (Llimona & Hladun, 2001). Amphiloma lobulatum (Flörke) Samp., in Sampaio & Crespí, Bol. Real Soc. Esp. Hist. Nat. 27(3): 143. Mar 1927, as “Amphiloma lobulatum Samp. (Caloplaca lobulata Oliv.)”; Sampaio, 1970: 192 : Lecanora lobulata Flörke. Remarks: = Caloplaca lobulata (Flörke) Hellb. (Llimona & Hladun, 2001). Acarospora magnussonii Samp., Bol. Soc. Brot., ser. 2, 2: 173. Post Mar 1924, as “Magnussoni”; Sampaio, 1970: 175. – Holotype: [Portugal: Minho:] Sá: Vale de Pereiras, nos pegões do aqueducto, Apr 1923, G. Sampaio (PO 5597L). Remarks: = Acarospora veronensis A. Massal. (Llimona & Hladun, 2001). Arthonia algarbica Samp., Ann. Sci. Acad. Polytechn. Porto 13(1): 28 Jul 1918; Sampaio, 1970: 94. – Holotype: [Portugal: Algarve:] Portimão, nas alfarrobeiras, 7 Apr 1917, G. Sampaio (PO 1361bL). Remarks: = Arthonia pruinata (Torss.) A. L. Sm. (Llimona & Hladun, 2001). Acarospora varzinensis Samp., Líqu. inéd.: 4. 15 Dec 1920; Sampaio, 1970: 112. – Syntypes: [Portugal: Douro Litoral:] Póvoa de Varzim, paredes do litoral, 17 Jan 1920, G. Sampaio (PO 2233L); [Portugal: Douro Litoral:] in Sampaio: Lich. Portugal nº 113, Póvoa de Varzim (paredes da beira-mar), Mar 1919, G. Sampaio (PO 5602L). Remarks: (1) = Acarospora subrufula (Nyl.) H. Olivier (Llimona & Hladun, 2001). (2) The syntype from Foz do Douro was not located. Arthopyrenia cinereopruinosa (Schaer.) A. Massal. var. olivetorum Samp., in Sampaio & Crespí, Bol. Real Soc. Esp. Hist. Nat. 27(3): 138. Mar 1927, as “Arthopyrenia cinereopruinosa Koerb. var. olivetorum Samp.”; Sampaio, 1970: 185. – Lectotype (designated here): [Portugal: Minho:] Ponte de Lima: Sá, nas oliveiras, 10 Mar 1919, G. Sampaio (PO 2141L; isolectotype PO 2141aL). Remarks: = Arthopyrenia cinereopruinosa (Schaer.) A. Massal. (Llimona & Hladun, 2001). Acarospora zahlbruckneri Samp., Brotéria, Sér. Bot. 19(1): 31. Mar 1921, as “Zahlbruckneri”; Sampaio, 1970: 135. – Lectotype (designated here): [Portugal: Ribatejo:] Santarém, nos muros do convento de Santa Clara, 2 Dec 1918, G. Sampaio (PO 2448L). Syntype: *Aspicilia transmontana Samp., Bol. Soc. Brot., ser. 2, 2: 169. Post Mar 1924; Sampaio, 1970: 170. – Holotype: [Portugal: Trás-os-Montes e Alto Douro:] Moncorvo, Mar 1922, J. Santos Júnior [det. G. Sampaio], (PO 2562L). 773 Paz-Bermúdez & al. • Typification of lichen names “Bacidia milliaria Samp.”, Lich. Portugal nº 147. Mar 1923, nom. nud. 51 • November 2002: 771–785 *Bacidia sulphurella Samp., Bol. Soc. Brot., ser. 2, 2: 174. Post Mar 1924; Sampaio, 1970: 176. – Holotype: [Portugal: Minho:] S. Gens (“Póvoa de Lanhoso: S. Gens” in protologue), nas oliveiras (rara), 3 Jul 1923, G. Sampaio (PO 5595L). “Buellia subcinerascens Samp.”, Brotéria, Sér. Bot. 15(3): 139, nom. inval., as “Buellia subcinerascens nob. in herb.” Sampaio, 1970: 58. Remarks: (1) Apparently intended to be a new combination based on Lecidea subcinerascens Nyl., but without any reference to a basionym, so not validly published; (2) = Buellia subcinerascens (Nyl.) Zahlbr. (Llimona & Hladun, 2001). *Blastenia subarenaria Samp., Ann. Sci. Acad. Polytechn. Porto 12(1): 48. Post 12 Jun 1917(c); Sampaio, 1970: 83. – Holotype: [Portugal: Algarve:] Portimão: Praia da Rocha, 7 Apr 1917, G. Sampaio (PO 1451L). Remarks: º Lecanora subarenaria (Samp.) Samp. (Sampaio, 1917d: 136; 1970: 55). Calicium brunneolum Ach. var. stemonoides Samp., Brotéria, Sér. Bot. 15(1): 12. 1 Apr 1917; Sampaio, 1970: 27. – Holotype: [Portugal: Estremadura:] Sintra: no parque, (“Cintra: na mata, sôbre as árvores” in protologue), 23 Aug 1916, G. Sampaio (PO 1356L). Remarks: = Chaenotheca brunneola (Ach.) Müll. Arg. (Llimona & Hladun, 2001). Buellia duartei Samp., Líqu. inéd.: 1. 15 Dec 1920, as “Duartei”; Sampaio, 1970: 109. – Lectotype (designated here): [Portugal: Minho:] Póvoa de Lanhoso: S. Gens (Nasce, na Lage Longa), 29 Sep 1919, G. Sampaio (PO 1993L; isolectotype PO 2054L). Remarks: = Buellia dispersa A. Massal. (Llimona & Hladun, 2001). Caloplaca herminica Samp., Ann. Sci. Acad. Polytechn. Porto 12(1): 47. Post 12 Jun 1917; Sampaio, 1970: 82. – Holotype: [Portugal: Beira Alta:] Serra da Estrela, nos penedos musgosos, 6 Aug 1916, Dr. Miguel Machado [det. G. Sampaio] (PO 1038L). Remarks: (1) º Lecanora herminica (Samp.) Samp. (Sampaio, 1917d: 135; 1970: 53). (2) = Caloplaca congrediens (Nyl.) Zahlbr. (Llimona & Hladun, 2001). Buellia jorgei Samp., Bol. Soc. Brot., ser. 2, 2: 177. Post Mar 1924, as “Jorgei”; Sampaio, 1970: 180. – Lectotype (designated here): [Portugal: Minho:] Ponte de Lima: Sá, nas macieiras, 7 Mar 1919, G. Sampaio (PO 2150L). Syntype: [Portugal: Minho:] Paredes de Coura, nos carvalhos, 28 Sep 1915, G. Sampaio (PO 2580L). Remarks: Sampaio (1924) also cited a syntype from Melgaço: Peso which was not located. Buellia myriocarpa Mudd. “var. aequata Samp.”, Lich. Portugal nº 179. Mar 1923, nom. nud. Buellia myriocarpa Mudd. var. lusitanica Samp., Bol. Soc. Brot., ser. 2, 2: 177. Post Mar 1924; Sampaio, 1970: 180. – Lectotype (designated here): [Portugal: Estremadura:] Mafra, castanheiros, Jan 1922, Dr. A. R. Jorge [det. G. Sampaio], in Sampaio: Lich. Portugal nº 178 (PO 5603L; isolectotypes in other herbaria). Remarks: = Buellia sardiniensis J. Steiner (Llimona & Hladun, 2001). *Buellia pseudosaxatilis Samp., Ann. Sci. Acad. Polytechn. Porto 12(1): 49. Post 12 Jun 1917; Sampaio, 1970: 84. – Holotype: [Portugal: Estremadura:] Sintra: Castelo dos Mouros, 23 Aug 1916, G. Sampaio (PO 1132L). Remarks: º Lecidea pseudosaxatilis (Samp.) Samp. (Sampaio, 1917d: 139; 1970: 59). “Caloplaca lallavei (Clemente) Samp.”, Ann. Sci. Acad. Polytechn. Porto 13(1): 34, 28 Jul 1918, as “Coloplaca Lallavei Samp. (comb. n.); Lecidea Lallavei Clem.; Lecanora Lallavei Nyl.”; Sampaio, 1970: 101. = Caloplaca lallavei (Clemente) Flagev (1886). Remarks: = Caloplaca erythrocarpa (Pers.) Zwackh (Llimona & Hladun, 2001). *Caloplaca lesdainii Samp., Rev. Estud. Univ. Porto 1(2): 37. Mar 1916, as “Lesdaini”; Sampaio, 1970: 1. – Holotype: [Portugal: Beira Litoral:] Buçaco (“Bussaco” in protologue), 9 Jan 1916, G. Sampaio (PO 786L). Remarks: º Lecanora lesdainii (Samp.) Samp. (Sampaio, 1916b:73; 1970: 13). Caloplaca peregrina Samp., Ann. Sci. Acad. Polytechn. Porto 12(1): 47. Post 12 Jun 1917; Sampaio, 1970: 82. – Lectotype (designated here): [Portugal: Trásos-Montes e Alto Douro:] Foz-Tua: margem do Douro, 11 Oct 1916, G. Sampaio (PO 1305L). Syntype: PO1304L. [Portugal: Beira Alta:] Barca de Alva (“Barca d’Alva” in protologue): margem do Águeda, penedo, 12 Oct 1916, G. Sampaio (PO 1304L; isosyntype PO 1304aL). Remarks: º Lecanora peregrina (Samp.) Samp. (Sampaio, 1917d: 133; 1970: 52). Carlosia Samp., A Aguia, sér. 3, 2(22): 192. Jun 774 51 • November 2002: 771–785 Paz-Bermúdez & al. • Typification of lichen names 1923; Sampaio, 1970: 159. Type: Carlosia lusitanica Samp. Remarks: (1) Sampaio (1923) included only one species, C. lusitanica, in his new genus Carlosia and so it necessarily provides the type. (2) = Thelomma A. Massal. (Tibell, 1976) 162; 1970: 158). (2) This specimen was discussed by Burgaz & al. (1999), where it is indicated that it belongs to Cladonia subscariosa Nyl. s. lat. Carlosia lusitanica Samp., A Aguia, sér. 3, 2(22): 192. Jun 1923; Sampaio, 1970: 160. Type: [Portugal: Trás-os-Montes e Alto Douro:] Tabuaço, 1923, Joaquim Sampaio [det. G. Sampaio], (LISU 1685), Remarks: = Thelomma siliceum (Fée) Tibell (Tibell, 1976: 248; Llimona & Hladun, 2001). Cladonia verticillata Hoff. “[unranked] cervicornis Samp.”, in Sampaio & Crespí, Bol. Real Soc. Esp. Hist. Nat. 27(3): 151. Mar 1927, nom. nud.; Sampaio, 1970: 201. Remarks: (1) Technically a nomen nudum because without references, but clearly refers to Lichen cervicornis Ach. (2) = Cladonia cervicornis (Ach.) Flot. (Llimona & Hladun, 2001). “Catillaria melastigma (A. Zahlbr.) Samp.”, in Sampaio & Crespí, Bol. Real Soc. Esp. Hist. Nat. 27(3): 144. Mar 1927, as “Catillaria melastigma Samp. (Lecidea chalybaea Borr.)”: Catillaria chalybeia f. melastigma A. Zahlbr. Remarks: (1) Although he did not cite the basionym, Sampaio evidently intended to publish a new combination based on Catillaria chalybeia f. melastigma A. Zahlbr. As his intended new combination would otherwise be validly published (as an illegitimate nomen novum for Lecidea chalybaea Borrer), on analogy with Art. 33.6(c) of the ICBN we are listing it as though a new combination even although not validly published under Art. 33.2. (2) = Catillaria chalybeia (Borrer) A. Massal. (Llimona & Hladun, 2001). *Chiodecton fragosoi Samp., Bol. Soc. Brot., ser. 2, 2: 167. Post Mar 1924, as “Fragosoi”; Sampaio, 1970: 168. – Syntype: [Portugal: Minho:] Ponte de Lima: muro do Vale das Pereiras, 21 Mar 1923, G. Sampaio (PO 5551L; isosyntypes PO 5551aL, PO 5551bL, PO 5551cL, PO 5551dL, PO 5551eL). Remarks: Sampaio (1924) also cited a syntype from Póvoa de Lanhoso which was not located. Cladonia foliacea Will. “[unranked] convoluta Samp.”, in Sampaio & Crespí, Bol. Real Soc. Esp. Hist. Nat. 27(3): 151. Mar 1927, nom. nud.; Sampaio, 1970: 201. Remarks: Technically a nomen nudum because without references, although clearly refers to Lichen convolutus Lam. = Cladonia convoluta (Lam.) Anders. Cladonia subturgida Samp., Ann. Sci. Acad. Polytechn. Porto 13(1): 38. Jul 1918; Sampaio, 1970: 106. – Holotype: [Portugal: Beira Alta:] Barca de Alva (“Barca d’Alva” in protologue), na terra, 13 Dec 1916, G. Sampaio (PO 1245L). Remarks: (1) = C. turgida Hoffm. (Sampaio, 1922: Cladonia uncialis “var. gracillima Samp.”, Lich. Portugal nº 291. Mar 1923, nom. nud. Collema anemoides Samp., Ann. Sci. Acad. Polytechn. Porto 13(1): 25. Jul 1918; Sampaio, 1970: 90. – Holotype: [Portugal: Beira Litoral:] Coimbra: monte de Santa Clara, 16 Feb 1918, G. Sampaio (PO 1634L). Remarks: = Collema crispum (Huds.) F. H. Wigg. (Llimona & Hladun, 2001). Collema harmandii Samp., Ann. Sci. Acad. Polytechn. Porto 13(1): 25. Jul 1918, as “Collema Harmandi Samp. (n. n.) Collema pulposulum Harm. (1905) non Nyl. (1864)”, nom. nov. : Collema pulposulum Harm.; Sampaio, 1970: 91. Remarks: = Collema coccophorum Tuck. (Llimona & Hladun, 2001). *Collemopsidium stenosporum Samp., Ann. Sci. Acad. Polytechn. Porto 13(1): 25. Jul 1918; Sampaio, 1970: 90. – Holotype: [Portugal: Algarve:] Portimão (“Vila Nova de Portimão” in protologue), 6 Apr 1917, G. Sampaio (PO 1692L). Coniocybe brunneola (Ach.) Samp., Brotéria, Sér. Bot. 20(3): 153. 10 Oct 1922, as “Coniocybe bruneola Samp. (Chaenotheca brunneola Mull-Arg)”; Sampaio, 1970: 147 : Calicium brunneolum Ach. Remarks: = Chaenotheca brunneola (Ach.) Müll. Arg. (Llimona & Hladun, 2001). Coniocybe chrysocephala (Ach.) Samp., Brotéria, Sér. Bot. 20(3): 153. 10 Oct 1922, as “Coniocybe chrysocephala Samp.; Chaenotheca chrysocephala Fr. fil.”: Calicium chrysocephalum Ach.; Sampaio, 1970: 147. Remarks: = Chaenotheca chrysocephala (Ach.) Th. Fr. (Llimona & Hladun, 2001). “Cornicularia intricata Samp.”, Lich. Portugal nº 268. Mar 1923, nom. nud. 775 Paz-Bermúdez & al. • Typification of lichen names 51 • November 2002: 771–785 Cyphelium zahlbruckneri Samp., Bol. Soc. Brot., ser. 2, 2: 161. Post Mar 1924, as “Zahlbruckneri”; Sampaio, 1970: 161. – Holotype: [Portugal: Trás-osMontes e Alto Douro:] Tabuaço (“Taboaço” in protologue): alto da mata do hospital, nos penedos, 10 Mar 1923, Joaquim Sampaio, [det. G. Sampaio] (PO 2632L). Remarks: = Cyphelium lecideinum (Nyl.) Trevis. (Tibell, 1971; Llimona & Hladun, 2001). Gyalecta deminuta Samp., Bol. Soc. Brot., ser 2, 2: 171. Post Mar 1924; Sampaio, 1970: 169. – Holotype: [Portugal: Minho:] Ponte de Lima: Carcaveira, nas ribas do caminho sobre Marsupella emarginata (Ehrh.) Dum., Apr 1923, G. Sampaio (PO 2636L; isotype in W, Vìzda, 1966). Remarks: = Bryophagus gloecapsa Nitschke (Llimona & Hladun, 2001). “Dimelaena castanoplaca Samp.”, Lich. Portugal nº 200. Mar 1923, nom. nud. *Gyalecta limica Samp., Rev. Estud. Univ. Porto 1(2): 38. Mar 1916, as “Gyaleta límica”; Sampaio, 1970: 2. – Holotype: [Portugal: Minho:] Ponte de Lima: Sá, nas cascas apodrecidas dos carvalhos velhos, 10 Aug 1915, G. Sampaio (PO 795L). Remarks: (1) º Lecidea limica (Samp.) Samp. (Sampaio, 1916b: 77; 1970: 18). (2) = Pachyphiale carneola Ar. (Sampaio, 1922: 154; 1970: 148), as “Pachyphiale carneola Ar.; Gyalecta cornea A. L. Sm.; Pachyphiale limica Samp.”. (3) Holotype identified in 1964 by A. Vìzda as Pachyphiale cornea (With.) Poetsch. “Diploschistes cinereocaesius (Sw.) Samp.”, in Sampaio & Crespí, Bol. Real Soc. Esp. Hist. Nat. 27(3): 137. Mar 1927 : Lichen cinereocaesius Sw.; Sampaio, 1970: 194 = Diploschistes cinereocaesius (Sw.) Vain. (1921). Diploschistes cinereocaesius (Sw.) Vain. “var. violarius Samp.”, in Sampaio & Crespí, Bol. Real Soc. Esp. Hist. Nat. 27(3): 137. Mar 1927, nom. nud., as “Diploschistes cinereocaesius Samp. var. violarius Samp.”; Sampaio, 1970: 194. Remarks: (1) Clearly intended to be a new combination based on Pertusaria violaria Nyl. but no reference is provided to the basionym, so technically a nomen nudum. (2) = Diploschistes scruposus (Schreb.) Norman (Llimona & Hladun, 2001). “Girardia cantabrica” Samp., in Sampaio & Crespí, Bol. Real Soc. Esp. Hist. Nat. 27(3): 137. Mar 1927, nom. nud.; Sampaio, 1970: 183. Remarks: (1) Apparently intended to be a new combination based on Ephebeia cantabrica Nyl., but no basionym cited. (2) See Paz-Bermúdez (2001). “Girardia lanata” Samp., Lich. Portugal nº 36. Mar 1923, nom. nud. Gyalecta decipiens Samp., Ann. Sci. Acad. Polytechn. Porto 13(1): 31. Jul 1918; Sampaio, 1970: 98. – Lectotype (designated here): [Portugal: Beira Litoral:] Coimbra: Choupal, 15 Feb 1918, G. Sampaio (PO 1699L). Syntype: [Portugal: Beira Litoral:] Coimbra: Choupal, 6 Jan 1916, G. Sampaio (PO 1624L). Remarks: (1) Sampaio (1924: 169; 1970: 170) referred G. decipiens to Gyalecta croatica Schul. & A. Zahlbr. following a re-identification of Portuguese specimens by G. Lettrau in 1922. (2) In 1964, A. Vìzda (in herb.) identified the two original syntypes as Gyalecta truncigena (Ach.) Hepp var. derivata (Nyl.) Boistel. (3) = Gyalecta derivata (Nyl.) H. Olivier (Llimona & Hladun, 2001). 776 “Hypogymnia tubulosa Samp.”, Lich. Portugal nº 238. Mar 1923, nom. nud. Koerberia lusitanica Samp., Bol. Soc. Brot., ser. 2, 2: 167. Post Mar 1924, as “Kœrberia”; Sampaio, 1970: 167. – Lectotype (designated here): [Portugal: Minho:] Póvoa de Lanhoso, nas rochas do castelo, 28 Jun 1920, G. Sampaio (PO 5490L, isolectotypes in Sampaio: Lich Portugal nº 55, PO 5604L, and other herbaria). Remarks: (1) Henssen (1963) reported that the holotype of this name was: “Koerberia lusitanica Samp. 1924, p. 167. Orig. coll.: Portugal, Sampaio (holotype: PO)”, but this is not an effective lectotypification under Art. 9.8 of the ICBN (Greuter & al., 2000) because he did not distinguish between the duplicates at PO. (2) = Koerberia sonomensis (Tuck.) Henssen (Llimona & Hladun, 2001). Lecania badiella Samp., Líqu. inéd.: 3. 15 Dec 1920; Sampaio, 1970: 110. – Holotype: [Portugal: Baixo Alentejo:] Beja, Apr 1917, G. Sampaio (PO 1874L). *Lecania citrinella Samp., Ann. Sci. Acad. Polytechn. Porto 12(1): 49. Post 12 Jun 1917; Sampaio, 1970: 83. – Holotype: [Portugal: Douro Litoral:] Leça da Palmeira, nas rochas marítimas, 17 Jan 1917, G. Sampaio (PO 1252L; topotype in LISU, M. Mayrhofer 1988). Remarks: (1) º Lecanora citrinella (Samp.) Samp. (Sampaio, 1917d: 130; 1970: 49). (2) The holotype specimen was identified as Lecanora actophila Wedd. by D. M. Rawson in 1969. 51 • November 2002: 771–785 “Lecania lesdainii (Samp.) Samp.”, Brotéria, Sér. Bot. 15(3): 144. 1 Dec 1917 (as “Lecanora Lesdaini Samp. (Lecania Lesdaini nob. in herb.)”; Sampaio, 1970: 64, nom. inval. pro syn. See Caloplaca lesdainii Samp. Lecanora calvosina Samp., Bol. Soc. Brot., ser. 2, 2: 170. Post Mar 1924; Sampaio, 1970: 171. – Holotype: [Portugal: Minho:] Póvoa de Lanhoso, S. Gens (“S. Gens de Calvos” in protologue), 9 Jul 1923, G. Sampaio (PO 2638L). Remarks: = Schaereria fuscocinerea (Nyl.) Clauzade & Roux (Llimona & Hladun, 2001). *Lecanora celestinoi Samp., Líqu. inéd.: 6. 15 Dec 1920, as “Celestini”; Sampaio, 1970: 113. – Lectotype (designated here): [Portugal: Minho:] Ponte de Lima: Sá; nas figueiras, macieiras, oliveiras, japoneiras, 10 Mar 1919, G. Sampaio (PO 2096L). Syntypes: [Portugal: Douro Litoral:] Póvoa de Varzim, junto ao cemitério, Feb 1920, (PO 2285L); [Portugal: Minho:] Braga, Falperra, 19 Jun 1920, G. Sampaio (PO 2378L). Lecanora circumrubens Samp., Ann. Sci. Acad. Polytechn. Porto 12(1): 47. Post 12 Jun 1917; Sampaio, 1970: 81. – Holotype: [Portugal: Algarve:] Portimão: Algarve; nas figueiras, 6 Apr 1917, G. Sampaio (PO 1401L). Remarks: (1) Redescribed by Sampaio, 1917d: 132; 1970: 51. (2) = Lecanora rubicunda Bagl. (Llimona & Hladun, 2001). Lecanora citrinella (Samp.) Samp., Brotéria, Sér. Bot. 15(3): 130. 1 Dec 1917; Sampaio, 1970: 49. : Lecania citrinella Samp. Sampaio (1917d) described Lecanora citrinella in Latin, but because he indicates that he earlier described the species under Lecania from the same material, it must be treated as a new combination, not a new species. Lecanora conimbricensis (Samp.) Samp., Brotéria, Sér. Bot. 14(2): 75. 1 Jul 1916; Sampaio, 1970: 15. : Rinodina conimbricensis Samp. Sampaio (1916b) described Lecanora conimbricensis in Latin, but as he described it from the same material under Rinodina (Sampaio, 1916a), it must be treated as a new combination, not a new species. Lecanora conimbricensis (Samp.) Samp. var. tumidula (Samp.) Samp., Brotéria, Sér. Bot. 14(2): 75. 1 Jul 1916; Sampaio, 1970: 16. (“tumídula”) : Rinodina conimbricensis Samp. var. tumidula Samp. Sampaio (1916b) described Lecanora conimbricensis var. tumidula in Latin, but as he had already described it from the same material as Rinodina conimbricensis Paz-Bermúdez & al. • Typification of lichen names var. tumidula (Sampaio, 1916a), it must be treated as a new combination, not a new variety. Lecanora gerezina Samp., nom. nov., Brotéria, Sér. Bot. 19(1): 17. Mar 1921; Sampaio, 1970: 136. : Lecanora tristis Samp. (1917) non Merezhk. (1913). Remarks: = Aspicilia epiglypta (Norrl.) Arnold (Llimona & Hladun, 2001). Lecanora herminica (Samp.) Samp., Brotéria, Sér. Bot. 15(3): 133. 1 Dec 1917; Sampaio, 1970: 53. : Caloplaca herminica Samp. Sampaio (1917d) described Lecanora herminica in Latin, but because he indicates that he earlier described the species under Caloplaca from the same material, it must be treated as a new combination, not a new species. Lecanora lesdainii (Samp.) Samp., Brotéria, Sér. Bot. 14(2): 73. 1 Jul 1916; Sampaio, 1970: 13. (“Lesdaini”) : Caloplaca lesdainii Samp. Sampaio (1916b) described Lecanora lesdainii in Latin, but because he indicated that he earlier described the species under Caloplaca from the same material, it must be treated as a new combination, not a new species. Lecanora lisbonensis Samp., Brotéria, Sér. Bot. 19(1): 33. Mar 1921; Sampaio, 1970: 137. – Syntypes: [Portugal: Algarve:] Praia das Maçãs, 23 Sep 1918, G. Sampaio (PO 991L); [Portugal: Beira Alta:] Viseu (“Vizeu” in protologue): nos sucalcos, 16 Apr 1916, G. Sampaio (PO 991aL); [Portugal: Beira Alta:] Viseu (“Vizeu” in protologue): penedos, 16 Apr 1916, G. Sampaio (PO 991bL); [Portugal: Algarve:] Portimão: Praia da Rocha, 1916, G. Sampaio (PO 1563L); [Portugal: Estremadura:] S. Martinho do Porto: rochedos marítimos, 12 Oct 1917, A. R. Jorge [det. G. Sampaio] (PO 1567L); [Portugal: Beira Litoral:] Buarcos: nas muralhas velhas do jardim, 13 Feb 1918, G. Sampaio (PO 1817L); [Portugal: Douro Litoral:] Póvoa de Varzim: estrada de Vila do Conde, muros, 4 Mar 1920, G. Sampaio (PO 2317L)); [Portugal: Estremadura:] in Sampaio: Lich. Portugal nº 85, Lisboa, Jan 1922, R. Jorge [det. G. Sampaio] (PO5607L), as “Psoroma lisbonense Samp.” Remarks: The syntypes from Sintra, Cascais and Almada were probably distributed under nº 85 of the exsiccata Lichenes de Portugal. Lecanora pachycarpa Samp., Brotéria, Sér. Bot. 15(1): 15. 1 Apr 1917; Sampaio, 1970: 31. – Holotype: [Portugal: Trás-os-Montes e Alto Douro:] Moncorvo (“Moncôrvo” in protologue): Carviçais; numa parede, 14 Oct 1916, G. Sampaio (PO 1184L; isotype PO 1184aL). Remarks: = Lecanora swartzii (Ach.) Ach. (Llimona 777 Paz-Bermúdez & al. • Typification of lichen names & Hladun, 2001). Lecanora peregrina (Samp.) Samp., Brotéria, Sér. Bot. 15(3): 133. 1 Dec 1917; Sampaio, 1970: 52. : Caloplaca peregrina Samp. Sampaio (1917d) described Lecanora peregrina in Latin, but because he indicates that he earlier described the species under Caloplaca from the same material, it must be treated as a new combination, not a new species. Lecanora pruinella (Bagl.) Samp. var. cintrana Samp., Brotéria, Sér. Bot. 14(2): 74. 1 Jul 1916, as “Lecanora pruinella (Rinodina pruinella Bagl.) var. cintrana Samp.”; Sampaio, 1970: 15. – Holotype: [Portugal: Estremadura:] Sintra, 27 Apr 1916, G. Sampaio (PO 919L). Remarks: (1) º Rinodina cintrana (Samp.) Samp. (Sampaio, 1922: 161; 1970: 155). (2) = Rinodina pruinella Bagl. (Llimona & Hladun, 2001). *Lecanora rustica Samp., Bol. Soc. Brot., ser. 2, 2: 170. Post Mar 1924; Sampaio, 1970: 171. – Holotype: [Portugal: Trás-os-Montes e Alto Douro:] Tabuaço: caminho velho de Távora, Joaquim Sampaio [det. G. Sampaio], 8 Mar 1923, (PO 2634L). Lecanora subarenaria (Samp.) Samp., Brotéria, Sér. Bot., 15(3): 136. 1 Dec 1917; Sampaio, 1970: 55. : Blastenia subarenaria Samp. Sampaio (1917d) described Lecanora subarenaria in Latin, but because he indicates that he earlier described the species under Blastenia from the same material, it must be treated as a new combination, not a new species. Lecanora tristis Samp., Brotéria, Sér. Bot. 15(1): 17. 1 Apr 1917, nom. illeg., non Lecanora tristis Merezhk. 1913; Sampaio, 1970: 32. – Holotype: [Portugal: Trásos-Montes e Alto Douro:] Serra do Gerês: Borrageira, 22 Sep 1916, G. Sampaio (PO 1119L). Remarks: = Aspicilia epiglypta (Norrl.) Arnold (Llimona & Hladun, 2001). Lecidea chodatii Samp., Líqu. inéd.: 6. 15 Dec 1920, as “Chodati”; Sampaio, 1970: 111. – Lectotype (Knoph, 1990: 88): [Portugal: Douro Litoral:] Póvoa de Varzim, nas paredes, Mar 1920, G. Sampaio, in Sampaio: Lich. Portugal nº 103 (PO 5644aL; isolectotype PO 5644L and in other herbaria). Remarks: (1) This specimen was identified as Lecidella chodati (Samp.) Knoph & Leuckert by J. G. Knoph in 1989. (2) = Lecidella granulosula (Nyl.) Knoph & Leuckert (Llimona & Hladun, 2001). *Lecidea flavigrana Samp., Brotéria, Sér. Bot. 778 51 • November 2002: 771–785 20(3): 156. 10 Oct 1922; Sampaio, 1970: 151. – Holotype: [Portugal: Trás-os-Montes e Alto Douro:] Bragança: estrada de Macedo; num choupo (rara), 15 Oct 1921, G. Sampaio (PO 2524L). Lecidea limica (Samp.) Samp., Brotéria, Sér. Bot. 14(2): 77. 1 Jul 1916. Sampaio, 1970: 18. : Gyalecta limica Samp. Sampaio (1916b) described Lecidea limica in Latin, but because he indicates that he earlier described the species under Gyalecta from the same material, it must be treated as a new combination, not a new species. *Lecidea machadoi Samp., Bol. Soc. Brot., ser. 2, 2: 171. Post Mar 1924, as “Machadoi”; Sampaio, 1970: 173. – Holotype: [Portugal: Minho:] Póvoa de Lanhoso: S. Gens; no lenho velho de oliveira, Mar 1923, Joaquim Sampaio [det. G. Sampaio] (PO 5598L). Lecidea macrocarpoides Samp., Brotéria, Sér. Bot. 15(1): 25. 1 Apr 1917; Sampaio, 1970: 41. – Lectotype (designated here): [Portugal: Minho:] Ponte de Lima: Sá, 14 Aug 1915, G. Sampaio (PO 421L). Syntype: [Portugal: Minho:] Ponte de Lima: Sá, 9 Sep 1916, G. Sampaio (PO 1306L). Remarks: (1) = Lecidea platycarpoides Bagl. (Sampaio, 1922: 156; 1970: 151). (2) = Porpidia platycarpoides (Bagl.) Hertel (Llimona & Hladun, 2001). (3) Sampaio (1917) also cited three syntypes from Porto which were not located. Lecidea nigrescens (Anzi) Samp., Brotéria, Sér. Bot. 14(2): 79. 1 Jul 1916, as “Lecidea nigrescens Samp. (Toninia nigrescens Anzi)”; Sampaio, 1970: 20 : Toninia nigrescens Anzi. Remarks: = Toninia cinereovirens (Schaer.) A. Massal (Llimona & Hladun, 2001). *Lecidea populicola Samp., Ann. Sci. Acad. Polytechn. Porto 13(1): 36. Jul 1918; Sampaio, 1970: 104. – Holotype: [Portugal: Beira Litoral:] Coimbra: Choupal, 15 Feb 1918, G. Sampaio (PO 1639L). Lecidea pseudosaxatilis (Samp.) Samp., Brotéria, Sér. Bot., 15(3): 139. 1 Dec 1917; Sampaio, 1970: 59: Buellia pseudosaxatilis Samp. Sampaio (1917d) described Lecidea pseudosaxatilis in Latin, but because he indicates that he earlier described the species under Buellia from the same material, it must be treated as a new combination, not a new species. *Lecidea rigata Samp., Bol. Soc. Brot., ser. 2, 2: 172. Post Mar 1924; Sampaio, 1970: 173. – Holotype: 51 • November 2002: 771–785 [Portugal: Minho:] Póvoa de Lanhoso: Porto de Bois; nos penedos do rio, Sep 1923, G. Sampaio (PO 2639L). *Leciographa fragosoi Samp., Bol. Soc. Brot., ser. 2, 2: 178. Post Mar 1924, as “Fragosoi”; Sampaio, 1970: 181. – Lectotype (designated here, first suggested by Rolf Santesson but not published by him): [Portugal: Douro Litoral:] Vila do Conde, nun muro, junto ao Castelo (litoral), 4 Mar 1920, G. Sampaio (PO 2298L). Syntype: [Portugal: Douro Litoral:] Porto: Castelo do Queijo, nos penedos do mar, 1 Jan 1921,G. Sampaio, (PO 2454L). “Leiophloea biformis (Borrer) Samp.”, in Sampaio & Crespí, Bol. Real Soc. Esp. Hist. Nat. 27(3): 137. Mar 1927, as “Lejophlea biformis Samp. (Acrocordia biformis Oliv.)”; Sampaio, 1970: 185 : Verrucaria biformis Borrer = Leiophloea biformis (Borrer) Trevis (1860). Remarks: = Anisomeridium biforme (Borrer) R. C. Harris (Llimona & Hladun, 2001). Leiophloea sphaeroides (Wallr.) Samp., Bol. Soc. Brot., ser. 2, 2: 162. Post Mar 1924, as “Leiophloea sphaeroides Samp. (comb. n.); Verrucaria sphaeoroides Wallr. (1831); Acrocordia sphaeoroides Arn. (1885); Arthopyrenia sphaeoroides A. Zahlbr. (1903)”; Sampaio, 1970: 162 : Verrucaria sphaeoroides Wallr. Remarks: = Acrocordia gemmata (Ach.) Massal. (Llimona & Hladun, 2001). Lemmopsis affinis Samp., Ann. Sci. Acad. Polytechn. Porto 13(1): 26. Jul 1918, as “affine”; Sampaio, 1970: 91. – Holotype: [Portugal: Trás-osMontes e Alto Douro:] Bragança: perto do rio Sabor, 10 Aug 1915, G. Sampaio (PO 1694L). “Lepidoma deceptorium Samp.”, Lich. Portugal nº 109. Mar 1923, nom. nud. “Lepidoma tabacinum Samp.”, Lich. Portugal nº 107. Mar 1923, nom. nud. “Lepraria candelaris (L.) Samp.”, in Sampaio & Crespí, Bol. Real Soc. Esp. Hist. Nat. 27: 151. Mar 1927; Sampaio, 1970: 201 : Byssus candelaris L. = Lepraria candelaris (L.) Fr. (1824). Remarks: = Chrysothrix candelaris (L.) J. R. Laundon (Llimona & Hladun, 2001). *Leptorhaphis epidermidis (Ach.) Th Fr. var. olivetorum Samp., Bol. Soc. Brot., ser. 2, 2: 162. Post Mar 1924; Sampaio, 1970: 163. – Lectotype (designated here): [Portugal: Beira Litoral:] Coimbra: Penedo da Paz-Bermúdez & al. • Typification of lichen names Saudade, numa oliveira, 7 Jan 1922, G. Sampaio (PO 2514L) – Syntype: [Portugal: Estremadura:] Óbidos, nas oliveiras, Jun 1918, A. Ricardo Jorge [det. G. Sampaio] (PO 2471L). Remarks: According to Aguirre-Hudson (1991:116) = Leptorhaphis oleae (A. Massal.) Körb. (?), identification made on the basis of description since the author could not see the type material. Leptorhaphis pinicola Samp., Bol. Soc. Brot., ser. 2, 2: 163. Post Mar 1924; Sampaio, 1970: 163. – Holotype: [Portugal: Estremadura:] Sintra: Estefãnia; Quinta dos Plátanos, nos pinheiros, Jan 1922, A. Ricardo Jorge [det. G. Sampaio] (PO 2530L). Remarks: (1) Aguirre-Hudson (1991:153) designated a neotype (in LISU) for this name because Sampaio’s type material was not found at PO at that time. Now that the holotype has been detected, the neotypification by Aguirre-Hudson is superseded. (2) = Cresporhaphis pinicola (Samp.) M. B. Aguirre (Llimona & Hladun, 2001). “Lethagrium aggregatum Samp.”, Lich. Portugal nº 46. Mar 1923, nom. nud. Lobaria mollissima Samp., Líqu. inéd.: 7. 15 Dec 1920; Sampaio, 1970: 114. – Isolectotype: [Portugal: Minho:] in Sampaio: Lich. Portugal nº 226, Castelo de Lanhoso, penedos musgosos, Jun 1920, Joaquim Sampaio [det. G. Sampaio] (PO 5599L, PO 2388L; lectotype in UPS, Jørgensen 2000: 672). Syntype: [Portugal: Minho:] Ponte de Lima: entre Sá e Santa Comba, 18 Sep 1920, G. Sampaio (PO 2426L). Remarks: = Erioderma mollisimum (Samp.) Du Rietz (Llimona & Hladun, 2001). Lopadium athalloides (Nyl.) Samp., Brotéria, Sér. Bot. 19(1): 19. Mar 1921; as “Lopadium athalloides Samp. (comb. n.); Lecidea athalloides Nyl.; Lopadium Newtoni Samp. in “Liq. ined.” pag. 2 (1920)”; Sampaio, 1970: 122: Lecidea athalloides Nyl. Remarks: = Gyalideopsis athalloides (Nyl.) Vezda (Llimona & Hladun, 2001). Lopadium newtonii Samp., Líqu. inéd.: 2. 15 Dec 1920, as “Newtoni”; Sampaio, 1970: 110. – Holotype: [Portugal: Douro Litoral:] Foz do Douro, Dec 1887, Isaac Newton [det. G. Sampaio] (PO 1649L). Remarks: (1) = Lopadium athalloides (Nyl.) Samp. (Sampaio, 1921: 19; 1970: 122). (2) = Gyalideopsis athalloides (Nyl.) Vìzda (Llimona & Hladun, 2001). *Microglaena cordeiroi (Harm.) Samp., Brotéria, Sér. Bot. 20(3): 151. 10 Oct 1922, as “Microglaena Cordeiri Samp.; Verrucaria Cordeiri Harm. (1909); 779 Paz-Bermúdez & al. • Typification of lichen names Microglaena modesta P. Cot. non Jat.”; Sampaio, 1970: 145 : Verrucaria Cordeiri Harm. “Ochrolechia subtartarea Samp.”, Lich. Portugal nº 89. Mar 1923, nom. nud. Omphalaria granitica Samp., Brotéria, Sér. Bot. 14(2): 66. 1 Jul 1916; Sampaio, 1970: 6. – Holotype: [Portugal: Minho:] Melgaço: Peso, rochedos graníticos na margem do rio Minho, 21 Aug 1915, G. Sampaio (PO 470L). Remarks: = Lichinella cribellifera (Nyl.) Moreno & Egea (Llimona & Hladun, 2001). Identified by P. Moreno & J. M. Egea under this name. Opegrapha vulgata Ach. “var. devulgata Samp.”, Lich. Portugal nº 13. Mar 1923, nom. nud. Pachyphiale carneolutea (Turner) Samp., Brotéria, Sér. Bot. 19(1): 22. Mar 1921, as “Pachyphiale carneolutea Samp. (comb. n.); Gyalecta carneolutea Oliv.; Pachyphiale lecanorina Stein. (?)”; Sampaio, 1970: 126 : Parmelia carneolutea Turner. Remarks: = Cryptolechia carneolutea (Turner) A. Massal. (Llimona & Hladun, 2001). “Pachyphiale limica (Samp.) Samp.”, Brotéria, Sér. Bot. 20(3): 153. 10 Oct 1922; Sampaio, 1970: 148, nom. inval. pro syn. See Gyalecta limica Samp. Parmelia conspersa Ach. “[unranked] lusitana Samp.”, in Sampaio & Crespí, Bol. Real Soc. Esp. Hist. Nat. 27: 148. Mar 1927; Sampaio, 1970: 197, nom. nud. Remarks: = Xanthoparmelia verrucigera (Nyl.) Hale; Parmelia verrucigera Nyl. (Llimona & Hladun, 2001). Parmelia stygia Ach. var. herminica Samp., Brotéria, Sér. Bot. 15(1): 13. 1 Apr 1917; Sampaio, 1970: 28. – Lectotype (designated here): [Portugal: Beira Alta:] Serra da Estrela: Charcos, nos penedos, 8 Aug 1916, G. Sampaio (PO 1004L). Remarks: (1) The syntypes from Serra do Gerez were not located. (2) Parmelia stygia. var. herminica Samp. is not included in Llimona & Hladun (2001) Pertusaria aspicilioides Samp., Brotéria, Sér. Bot. 15(1): 22. 1 Apr 1917; Sampaio, 1970: 38. – Holotype: [Portugal: Minho:] Ponte de Lima: Sá, Sep 1916, G. Sampaio (PO 5452L). Remarks: Pertusaria aspicilioides Samp. is not included in Llimona & Hladun (2001) *Pertusaria lutescens Lamy var. conimbricensis 780 51 • November 2002: 771–785 Samp., Líqu. esp. Willkomm: 143. Post May 1917; Sampaio, 1970: 76. Potential type material: [España: Andalucía:] Lomas de Alcornoque, 22 Feb 1845, leg. Willkomm. “Pertusaria pseudocorallina (Lilj.) Samp.”, in Sampaio & Crespí, Bol. Real Soc. Esp. Hist. Nat. 27(3): 140. Mar 1927, as “Pertusaria pseudocorallina Samp. (P. Westringii Leight.)”; Sampaio, 1970: 188, nom. inval. = Pertusaria pseudocorallina (Lilj.) Arnold (1887). Remarks: Although he did not cite the basionym, Sampaio evidently intended to publish a new combination based on Lichen pseudocorallinus Lilj. As his intended new combination would otherwise be validly published (as an illegitimate nomen novum for Pertusaria Westringii Leight.), on analogy with Art. 33.6(c) of the ICBN we are listing it as a new combination that is superfluous having already been made by Arnold in 1887. Pertusaria pseudocorallina Samp. “var. concreta Samp.”, in Sampaio & Crespí, Bol. Real Soc. Esp. Hist. Nat. 27(3): 140. Mar 1927; Sampaio, 1970: 188, nom. nud. Remarks: = Pertusaria pseudocorallina (Lilj.) Arnold (Llimona & Hladun, 2001). Physcia caesia Nyl. var. perrugosa Samp., Brotéria, Sér. Bot. 20(3): 162. 10 Oct 1922; Sampaio, 1970: 175. – Lectotype (designated here): [Portugal: Minho:] Serra de Arga: Santa Justa, 13 Sep 1916, G. Sampaio (PO 1081L). Remarks: (1) = Physcia caesia (Hoffm.) Fürnr. (Llimona & Hladun, 2001). (2) The syntypes from Póvoa de Lanhoso and Braga were not located. Physcia hispida Tuck. “[unranked] leptalea Samp.”, in Sampaio & Crespí, Bol. Real Soc. Esp. Hist. Nat. 27(3): 146. Mar 1927, as “Physcia hispida Tuck. [Ph. tenella Ach. (1798), DC. (1805)] r. leptalea Samp.”; Sampaio, 1970: 196. Remarks: (1) Apparently intended to be a new combination based on Lichen leptaleus Ach., but no basionym is cited and thus the name is not validly published. (2) = Physcia leptalea (Ach.) DC. (Llimona & Hladun, 2001). Physma hispanicum Samp., Líqu. esp. Willkomm: 136. Post May 1917; Sampaio, 1970: 69. Potential type material: [España: Andalucía:] circa Malacam, Willkomm 1844 [det. Flotow as Collema aggregatum Nyl.]. Remarks: = Lempholemma hispanicum (Samp.) Zahlbr. (Llimona & Hladun, 2001). 51 • November 2002: 771–785 Pilocarpon leucoblepharum (Nyl.) Vain. var. chloroticum Samp., Bol. Soc. Brot., ser. 2, 2: 171. Post Mar 1924; Sampaio, 1970: 196. – Lectotype (designated here, first suggested by Rolf Santesson but not published by him): [Portugal: Estremadura:] Sintra, na tapada num Abies, 23 Aug 1916, G. Sampaio (PO 1134L). Syntype: [Portugal: Beira Litoral:] Buçaco, 24 Sep 1918, A. Ricardo Jorge [det. G. Sampaio] (PO 1836L). Remarks: = Byssoloma leucoblepharum (Nyl.) Vain. (Llimona & Hladun, 2001). *Polyblastia exigua Samp., Brotéria, Sér. Bot. 20(3): 152. 10 Oct 1922; Sampaio, 1970: 146. Holotype: [Portugal: Trás-os-Montes e Alto Douro:] Bragança: Panacal, 19 Oct 1921, G. Sampaio (PO 2553L). “Psoroma lisbonense Samp.”, Lich. Portugal nº 85. Mar 1923, nom. nud.: Lecanora lisbonensis Samp. Remarks: Clearly intended as a new combination based on his earlier Lecanora lisbonensis (Sampaio, 1921). “Psoroma murale (Schreb.) Samp.”, in Sampaio & Crespí, Bol. Real Soc. Esp. Hist. Nat. 27(3): 142. Mar 1927, as “Psoroma murale Samp. (Lecanora saxicola Ach.)”: Lichen murale Schreb.; Sampaio, 1970: 190, Remarks: (1) Although he did not cite the basionym, Sampaio evidently intended to publish a new combination based on Lichen murale Schreb. As his intended new combination would otherwise be validly published (as an illegitimate nomen novum for Lecanora saxicola Ach.), on analogy with Art. 33.6(c) of the ICBN we are listing it as though a new combination even although not validly published under Art. 33.2. (2) = Lecanora muralis (Schreb.) Rabenh. (Llimona & Hladun, 2001). *Psorotichia henriquesii Samp., O Instituto 63(5): 245–246. May 1918, as “Henriquesi”; Sampaio, 1970: 87. – Holotype: [Portugal: Beira Litoral:] Coimbra: Santa Clara, 16 Feb 1918, G. Sampaio (PO 1674L). *Psorotichia macrospora Samp., Bol. Soc. Brot., ser. 2, 2: 166. Post Mar 1924; Sampaio, 1970: 166. – Holotype: [Portugal: Douro Litoral:] Porto: Quinta Amarela; nos muros, Mar 1923, G. Sampaio (PO 2627L). Pyrenopsis anemoides Samp., Bol. Soc. Brot., ser. 2, 2: 165. Post Mar 1924; Sampaio, 1970: 166. – Lectotype (designated here): [Portugal: Minho:] Póvoa de Lanhoso: S. Gens, Barreiros; nas paredes, 1 Sep 1919, G. Sampaio (PO 2191L). Syntypes: [Portugal: Minho:] Póvoa de Lanhoso: Frades em S. Roque, numa parede, 29 Aug 1919, G. Sampaio (PO 2192L); [Portugal: Minho:] Póvoa de Lanhoso: entre o Terrão e Porto de Bois, numa parede, Paz-Bermúdez & al. • Typification of lichen names 3 Sep 1919, G. Sampaio (PO 2193L; isosyntype PO 2193aL). Remarks: = Collema crispum (Huds.) F. H. Wigg. (Llimona & Hladun, 2001). *Pyrenopsis calvosina Samp., Bol. Soc. Brot., ser. 2, 2: 165. Post Mar 1924; Sampaio, 1970: 165. – Lectotype (designated here): [Portugal: Minho:] Póvoa de Lanhoso: Alto de Calvos, nas paredes, G. Sampaio (PO 2521L). Syntype: [Portugal: Minho:] Ponte de Lima: Sá, 12 Nov 1920, G. Sampaio (PO 2522L, isosyntype PO 2522aL). Ramalina portuensis Samp., Bol. Soc. Brot., ser. 2, 2: 179. Post Mar 1924; Sampaio, 1970: 182. – Lectotype (designated by Krog & James 1977: 38, as “holotype”): [Portugal: Douro Litoral:] in Sampaio: Lich. Portugal nº 276, Leça de Palmeira: Boa Nova (pinheiros), Oct 1922, G. Sampaio (LISU; isolectotypes PO 5600L and in other herbaria). *Rhizocarpon discrepans Samp., Ann. Sci. Acad. Polytechn. Porto 12(1): 50. Post 12 Jun 1917; Sampaio, 1970: 85. – Holotype: [Portugal: Trás-os-Montes e Alto Douro:] Gerês: Borrageira, 22 Sep 1916, G. Sampaio (PO 1234L). Rhizocarpon geographicum DC. var. lavatum Samp., Bol. Soc. Brot., ser. 2, 2: 178. Post Mar 1924; Sampaio, 1970: 181. Lectotype (designated by Runemark, 1956: 113): [Portugal: Minho:] Póvoa de Lanhoso, G. Sampaio, (UPS) Remarks: (1) Runemark (1956: 113): “Portugal, P. de Lanhoso, Sampaio (Magn. Lectotype)”. Herb. Magnusson is now in UPS. (2) = R. sublucidum var. lavatum (Samp.) Räsänen (Räsänen, 1949: 85). (3) There is no suggestion that Sampaio was basing his variety on Lichen lavatum Fr. and the lectotype specimen is not, in fact, Rhizocarpon lavatum (Fr.) Hazsl., as stated in Llimona & Hladun (2001), but Rhizocarpon lindsayanum subsp. kittilense Räsänen (Runemark 1956) (= Rhizocarpon geographicum (L.) DC. in Llimona & Hladun, 2001). Rinodina atrocinerella (Nyl.) Boist. var. macrospora Samp., Ann. Sci. Acad. Polytechn. Porto 13(1): 30. Jul 1918; Sampaio, 1970: 96. – Holotype: [Portugal: Trásos-Montes e Alto Douro:] Foz-Tua, 11 Oct 1916, G. Sampaio (PO 1223L). Remarks: = Buellia atrocinerella (Nyl.) Scheid. (Llimona & Hladun, 2001). Rinodina cintrana (Samp.) Samp., Brotéria, Sér. Bot. 20(3): 161. 10 Oct 1922; Sampaio, 1970: 155. : 781 Paz-Bermúdez & al. • Typification of lichen names Lecanora pruinella Bagl. var. cintrana Samp. Rinodina confinis Samp., Bol. Soc. Brot., ser. 2, 2: 177. Post Mar 1924; Sampaio, 1970: 179. – Lectotype (designated here): [Portugal: Beira Litoral:] Coimbra: Santo Antonio dos Olivais; nas oliveiras, 6 Jan 1916, G. Sampaio (PO 710L) – Syntypes: [Portugal: Minho:] Ponte de Lima: Sá, troncos velhos duma oliveira, 15 Sep 1915, G. Sampaio (PO 661L); [Portugal: Minho:] Póvoa de Lanhoso: S. Gens, nas oliveiras, 10 Sep 1919, G. Sampaio (PO 1915L); [Portugal: Minho:] Póvoa de Lanhoso: S. Gens, nos carvalhos, 2 Sep 1919, G. Sampaio (PO 1917L). [Portugal: Minho:] Póvoa do Lanhoso, S. Gens, 8-1919, G. Sampaio (M, UPS) in Giralt & Mayrhofer (1995). Remarks: (1) Sampaio (1924) also cited a syntype from Póvoa de Varzim: Laundes which was not located. (2) The nº 198 of the exsiccata Lichenes de Portugal, was reported by Giralt & Mayrhofer (1995: 142) as isotype; but that is not correct because its locality (Póvoa de Varzim: Anjos) was not listed in the protologue by G. Sampaio. Rinodina confragosa (Ach.) Körb. var. olivieri Samp., Bol. Soc. Brot., ser. 2, 2: 178. Post Mar 1924, as “Olivieri”; Sampaio, 1970: 178. Lectotype (designated here): [Portugal: Minho:] Póvoa de Lanhoso, nos rochedos do castelo, 7 Nov 1921, G. Sampaio, in Sampaio: Lich. Portugal nº 194 (PO 5601L; isotypes PO 2490L and other herbaria). Remarks: (1) The name Rinodina confragosa var. olivieri was apparently first published in Sampaio, Lich. Portugal nº 194, but as a nomen nudum because the label has not a description. Mayrhofer & al. (1993: 301) cite incorrectly the exsiccata´s specimens as type material. (2) = Rinodina santorinensis Steiner var. olivieri (Samp.) H. Mayrhofer & Sattler (Llimona & Hladun, 2001). Rinodina conimbricensis Samp., Rev. Estud. Univ. Porto 1(2): 37. Mar 1916; Sampaio, 1970: 1. – Lectotype (designated here): [Portugal: Beira Litoral:] Coimbra: Santo António (“Antônio” in protologue) dos Olivais, 6 Jan 1916, G. Sampaio (PO 741L) – Syntype: [Portugal: Beira Alta:] Viseu, paredes sobre os musgos, 18 Apr 1916, G. Sampaio (PO 894L). Remarks: (1) º Lecanora conimbricensis (Samp.) Samp. (Sampaio, 1916b: 75; 1970: 15). (2) = Rinodina intermedia Bagl. (Llimona & Hladun, 2001). Rinodina conimbricensis Samp. var. tumidula Samp., Rev. Estud. Univ. Porto 1(2): 38. Mar 1916, as “tumídula”; Sampaio, 1970: 2. – Holotype: [Portugal: Beira Litoral:] Coimbra: Santo António dos Olivais, terra e musgos, 6 Jan 1916, G. Sampaio (PO 712L). 782 51 • November 2002: 771–785 Remarks: (1) º Lecanora conimbricensis (Samp.) Samp. var. tumidula (Samp.) Samp. (Sampaio, 1916b: 75; 1970: 16. (2) = Rinodina intermedia Bagl. (Llimona & Hladun, 2001). Rinodina lesdainii Samp., Bol. Soc. Brot., ser. 2, 2: 175. Post Mar 1924, as “Lesdaini”; Sampaio, 1970: 178. – Lectotype (designated here): [Portugal: Minho:] Póvoa de Lanhoso: S. Gens; nas lages de paredes, 28 Mar 1923, G. Sampaio (PO 2635L); isolectotypes: PO 5550L; PO 5550aL; PO 5550bL; PO 5550cL; PO 5550dL; PO 5550eL; PO 5550fL and in W! (Mayrhofer & Poelt, 1979). Schismatomma diplotommoides (Bagl.) Samp., Brotéria, Sér. Bot. 15(3): 140. 1 Dec 1917, as “Schismatomma diplotommoides nob. (Lecania diplotommoides Bagl.)”; Sampaio, 1970: 60 : Lecania diplotommoides Bagl. Remarks: = Schismatomma dirinellum (Nyl.) Zahlbr. (Llimona & Hladun, 2001). “Schismatomma graphidioides (Leight.) Samp.”, Brotéria, Sér. Bot. 15(3): 140. 1 Dec 1917, as “Schismatomma graphidioides nob. (Platygrapha rimata Nyl.)”; Sampaio, 1970: 60 : Chiodecton graphidioides Leight. Remarks: (1) Although he did not cite the basionym, Sampaio evidently intended to publish a new combination based on Chiodecton graphidioides Leight. As his intended new combination would otherwise be validly published (as an illegitimate nomen novum for Platygrapha rimata Nyl.), on analogy with Art. 33.6(c) of the ICBN, we are listing it as though a new combination even although not validly published under Art. 33.2. The new combination was made validly by Zahlbruckner in 1919. (2) = Schismatomma graphidioides (Leight.) Zahlbr. (Llimona & Hladun, 2001). Setaria jubata Ach. “var. chalybeioformis Samp.”, Lich. Portugal nº 269. Mar 1923, nom. nud. Solenopsora holophaea (Mont.) Samp., Brotéria, Sér. Bot. 19(1): 26. Mar. 1921, as “Solenopsora holophaea; Parmelia holophaea Mont.”; Sampaio, 1970: 130 : Parmelia holophaea Mont. Solenopsora olivacea (Fr.) Samp. “var. spadicea Samp.”, in Sampaio & Crespí, Bol. Real Soc. Esp. Hist. Nat. 27(3): 144. Mar 1927, as “Solenopsora olivacea Samp. var. spadicea Samp”; Sampaio, 1970: 192, nom. nud. Remarks: (1) Apparently intended to be a new combination based on Lecanora spadicea Flot. but no other 51 • November 2002: 771–785 citation given and hence technically a nomen nudum. (2) = Lecania spadicea (Flot.) Zahlbr. (Llimona & Hladun, 2001). “Solenopsora subdisparata Samp.”, Lich. Portugal nº 140. Mar 1923, nom. nud. “Toninia albilabra (Dufour) Samp.”, Ann. Sci. Acad. Polytechn. Porto 13(1): 32. Jul 1918, as “Toninia albilabra Samp. (comb. n.); Biatora albilabra Duf. ex Fr.; Lecidea albilabra Nyl.; Thalloidima albilabrum Flag.”; Sampaio, 1970: 99 = Toninia albilabra (Dufour) H. Olivier (1911). “Toninia lurida (Arnold) Samp.”, Ann. Sci. Acad. Polytechn. Porto 13(1): 32. Jul 1918, as “Toninia lurida Samp. (comb. n.); Thalloidima luridum Bagl.”; Sampaio, 1970: 99 = Toninia lurida (Arnold) H. Olivier (1911). Remarks: = Toninia physaroides (Opiz) Zahlbr. (Llimona & Hladun, 2001). Toninia sabulosa (Massal.) Samp., Ann. Sci. Acad. Polytechn. Porto 13(1): 32. Jul 1918, as “Toninia sabulosa Samp. (comb. n.); Bilimbia sabulosa Mass.; Lecidea sabuletorum ß syncomista Floerk.; Toninia syncomista Fr. fil.”; Sampaio, 1970: 99 : Bilimbia sabulosa Massal. Remarks: = Mycobilimbia lobulata (Sommerf.) Hafellner (Llimona & Hladun, 2001). *Verrucaria carrissoi Samp., Brotéria, Sér. Bot. 20(3): 150. 10 Oct 1922, as “Carrisoi”; Sampaio, 1970: 144. – Holotype: [Portugal: Beira Litoral:] Coimbra: Monte de Santa Clara, nas rochas calcáreas, 11 Jan 1922, G. Sampaio (PO 2536L). *Verrucaria limae Samp., Bol. Soc. Brot., ser. 2, 2: 163. Post Mar 1924, as “Limae”; Sampaio, 1970: 164. – Lectotype (designated here): [Portugal: Minho:] Ponte de Lima: Sá, paredes nos montes, Feb 1923, G. Sampaio (PO 2631L). Remarks: The syntype from Póvoa de Lanhoso was not located. Xanthocarpia aurantiaca (Lightf.) Samp., in Sampaio & Crespí, Bol. Real Soc. Esp. Hist. Nat. 27: 143. Mar 1927, as “Xanthocarpia aurantiaca Samp. (Placodium aurantiacum Hepp in Fl. Europ. núm. 399)”; Sampaio, 1970: 192. : Lichen aurantiacus Lightf. Remarks: = Caloplaca flavorubescens (Huds.) J. R. Laundon (Llimona & Hladun, 2001). “Xanthocarpia irrubescens Samp.”, Lich. Portugal nº 124. Mar 1923, nom. nud. Paz-Bermúdez & al. • Typification of lichen names “Xanthocarpia phlogina Samp.”, Lich. Portugal nº 123. Mar 1923, nom. nud. Xanthocarpia vitellinula (Nyl.) Samp., in Sampaio & Crespí, Bol. Real Soc. Esp. Hist. Nat. 27: 143, as “Xanthocarpia vitellinula Samp. (Lecanora vitellinula Nyl.)”; Sampaio, 1970: 192 : Lecanora vitellinula Nyl. Remarks: = Caloplaca vitellinula (Nyl.) H. Olivier (Llimona & Hladun, 2001). II. New t a xa describ ed by ot h er a u t hors In the following lists, those names for which types have been located are in bold italic, whereas those for which types could not be located are in italic. M. B ou ly de Le sda in & G. S a m pa io Acarospora duriana de Lesd. & Sampaio, Bull. Soc. Bot. France 68: 205. 1921. Potential type material: [Portugal: Trás-os-Montes e Alto Douro:] Foz-Tua: margem do Douro, G. Sampaio [det. M. Bouly de Lesdain]. *Lecanora bracarensis de Lesd. & Sampaio, Bull. Soc. Bot. France 68: 204. 1921. Potential type material: [Portugal: Minho:] Braga, Falperra, G. Sampaio [det. M. Bouly de Lesdain]. *Lecanora limica de Lesd. & Samp., Bull. Soc. Bot. France 68: 203. 1921, as “Limica”. Holotype: [Portugal: Minho:] Ponte de Lima: Sá (“sur les pierres granitiques d’un mur” in protologue), Sep 1916, G. Sampaio [det. M. Bouly de Lesdain] (PO 2416L). *Lecanora piniperda Körber var. lusitanica de Lesd. & Sampaio, Bull. Soc. Bot. France 68: 204. 1921. Lectotype (designated here): [Portugal: Minho:] Ponte de Lima: Sá, numa austrália, Sep 1920, G. Sampaio [det. M. Bouly de Lesdain] (PO 2417L). Syntype: [Portugal: Minho:] Póvoa de Lanhoso, numa pereira velha, 26 Oct 1920, G. Sampaio [det. M. Bouly de Lesdain] (PO 2418L). M. B ou ly de Le sda in Lecania sampaiana de Lesd., Bull. Soc. Bot. France 68: 204. 1921. Lectotype (designated here): [Portugal: Douro Litoral:] Póvoa de Varzim, penedo da beira mar, 17 May 1920, G. Sampaio [det. M. Bouly de Lesdain] (PO 2351L). Syntype: [Portugal: Douro Litoral:] Vila do Conde, 783 Paz-Bermúdez & al. • Typification of lichen names 51 • November 2002: 771–785 Mar 1920, G. Sampaio [det. M. Bouly de Lesdain] (PO 2349L). Remarks: = *Candelariella subdisparata (Nyl.) Zahlbr. (Llimona & Hladun, 2001). Microglaena sampaiana de Lesd., Bull. Soc. Bot. France 68: 206. 1921, as “Microglœna Sampaiana”. Holotype (designated by H. Mayrhofer): [Portugal: Minho:] Póvoa de Lanhoso, entre Calvos e as Vinhas, 29 Sep 1919, G. Sampaio [det. M. Bouly de Lesdain] (PO 2027L). Remarks: = Thelenella inductula (Nyl.) H. Mayrhofer (Llimona & Hladun, 2001). *Lecidea portuensis Nyl., Bol. Soc. Brot., 6: 241. 1888, as “Portuensis” = Lecidea Oportensis Nyl., Flora 63: 12. 1880. Syntype: [Portugal: Douro Litoral:] Valongo: Lugar do Seixo, Jun 1879, Isaac Newton [det. W. Nylander] (PO 2910L). W. Nylander (taxa described from Portugal) *Arthonia baeastroidea Nyl., Flora 64: 6. 1881. Syntype: [Portugal: Douro Litoral:] Porto: Ramal (Seixo), Sep 1880, Isaac Newton [det. W. Nylander] (PO 2872L). Chiodecton spilocarpum Nyl., Flora 63: 391. 1880. Holotype: H-NYL. 4483. [Portugal: Douro Litoral:] Porto. in Torrente & Egea (1989). Syntype: [Portugal: Douro Litoral:] Vila Nova de Gaia: Cabedelo, 27 May 1880, Isaac Newton [det. W. Nylander] (PO 2804L). Remarks: (1) Identified by Torrente & Egea (1989) as Lecanactis grumulosa (Dufour) Fries (2) = Lecanographa grumulosa (Dufour) Egea & Torrente (Llimona & Hladun, 2001). Lecanora glaucolutescens Nyl., Flora 63: 389. 1880, as “glauco-lutescens”. Syntype: [Portugal: Douro Litoral:] Requesende, May 1879, Isaac Newton [det. W. Nylander] (PO 2843L). Lecanora plumbella Nyl., Bol. Soc. Brot. 6: 227. 1888. Potential type material: [Portugal: Douro Litoral:] Porto, Isaac Newton [W. Nylander]. Remarks: = Rinodina atrocinerea (Hook.) Körb. (Llimona & Hladun, 2001). Lecanora quartzina Nyl., Flora 64: 3. 1881. Syntype: [Portugal: Douro Litoral:] Castelo do Queijo: Foz, Aug 1880, Isaac Newton [det. W. Nylander] (PO 2842L). Remarks: = Lecanora actophila Wedd. (Llimona & Hladun, 2001). Lecanora subdisparata Nyl., Flora 63: 10. 1880. Syntype: [Portugal: Douro Litoral:] Lordelo: Estrada da Foz, 11 Jul 1880, Isaac Newton, [det. W. Nylander] (PO 5406L). 784 Lecidea vexabilis Nyl., Flora 63: 390. 1880. Syntype: [Portugal: Douro Litoral:] Isaac Newton [det. W. Nylander] (PO 2820L). Remarks: = Mycobilimbia sabuletorum (Schreber) Hafellner (Llimona & Hladun, 2001). Pyrenopsis triptococca Nyl., Flora 64: 2. 1881. Potential type material: [Portugal: Douro Litoral:] Porto, Isaac Newton [det. W. Nylander]. Ramalina digitellata Nyl., Flora 63: 10. 1880. Syntype: [Portugal: Douro Litoral:] Porto: Arrábida, May 1879, Isaac Newton [det. W. Nylander] (PO 5527L). Remarks: = Ramalina capitata (Ach.) Nyl. var. digitellata (Nyl.) Nimis (Llimona & Hladun, 2001). Thelotrema leiospodium Nyl., Flora 63: 11. 1880 Syntypes: [Portugal: Douro Litoral:] Serra do Pilar: Estrada marginal, Jun 1879, Isaac Newton [det. W. Nylander] (PO 5429L); [Portugal: Douro Litoral:] Leça da Palmeira: S. Gens, Sep 1880, Isaac Newton [det. W. Nylander] (PO 5430L). Remarks: = Thelotrema wightii (Taylor) Nyl. (Llimona & Hladun, 2001). ACKNOWLEDGEMENTS This study was financially supported by a FCT grant (Ministério da Ciência e da Tecnologia de Portugal, reference number SFRH/BPD/3559/2000) to Graciela Paz-Bermúdez. We are very grateful to an anonymous reviewer for critical advice and to John McNeill for helpful suggestions and support. LITERATURE CITED Aguirre-Hudson, B. 1991. A taxonomic study of the species referred to the ascomycete genus Leptorhaphis. Bull. Brit. 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