World checklist of hornworts and liverworts

Alfons Schäfer-Verwimp; Barbara Crandall-stotler; Christine Cargill; Matt Von Konrat; Tamás Pócs; Konstantinova, N.A.; Jiří Váňa

A working checklist of accepted taxa worldwide is vital in achieving the goal of developing an online flora of all known plants by 2020 as part of the Global Strategy for Plant Conservation. We here present the first-ever worldwide checklist for liverworts (Marchantiophyta) and hornworts (Anthocerotophyta) that includes 7486 species in 398 genera representing 92 families from the two phyla. The checklist has far reaching implications and applications, including providing a valuable tool for taxonomists and systematists, analyzing phytogeographic and diversity patterns, aiding in the assessment of floristic and taxonomic knowledge, and identifying geographical gaps in our understanding of the global liverwort and hornwort flora. The checklist is derived from a working data set centralizing nomenclature, taxonomy and geography on a global scale. Prior to this effort a lack of centralization has been a major impediment for the study and analysis of species richness, conservation and systematic research at both regional and global scales. The success of this checklist, initiated in 2008, has been underpinned by its community approach involving taxonomic specialists working towards a consensus on taxonomy, nomenclature and distribution.

PhytoKeys 59: 1–828 (2016) doi: 10.3897/phytokeys.59.6261 http://phytokeys.pensoft.net World checklist of hornworts and liverworts A peer-reviewed open-access journal 1 CHECKLIST Launched to accelerate biodiversity research World checklist of hornworts and liverworts Lars Söderström1,*, Anders Hagborg2,*, Matt von Konrat2,*, Sharon Bartholomew-Began3, David Bell4, Laura Briscoe2, Elizabeth Brown5, †, D. Christine Cargill6, Denise P. Costa7, Barbara J. Crandall-Stotler8, Endymion D. Cooper9, Gregorio Dauphin10, John J. Engel2, Kathrin Feldberg11, David Glenny12, S. Robbert Gradstein13, Xiaolan He14, Jochen Heinrichs11, Jörn Hentschel15, Anna Luiza Ilkiu-Borges16, Tomoyuki Katagiri17, Nadezhda A. Konstantinova18, Juan Larraín2, David G. Long19, Martin Nebel20, Tamás Pócs21, Felisa Puche22, Elena Reiner-Drehwald23, Matt A.M. Renner5, Andrea Sass-Gyarmati21, Alfons Schäfer-Verwimp24, José Gabriel Segarra Moragues25, Raymond E. Stotler8, †, Phiangphak Sukkharak26, Barbara M. hiers27, Jaime Uribe28, Jiří Váňa29, Juan Carlos Villarreal30, Martin Wigginton31, Li Zhang32, Rui-Liang Zhu33 1 Department of Biology, Norwegian University of Science and Technology, N-7491 Trondheim, Norway 2 Department of Science and Education, Field Museum, 1400 South Lake Shore Drive, Chicago, IL 60605– 2496, United States of America 3 Department of Biology, West Chester University, West Chester, PA 19383, United States of America 4 Department of Botany, University of British Columbia, 6270 University Boulevard, Vancouver, BC, V6T 1Z4, Canada 5 Royal Botanic Gardens and Domain Trust, Mrs Macquaries Road, Sydney NSW2000, Australia 6 Centre for Australian National Biodiversity Research, Australian National Herbarium, GPO Box 1600, Canberra, ACT 2601, Australia 7 Instituto de Pesquisas Jardim Botânico do Rio de Janeiro, Rua Pacheco Leão 915, 22460-030, Rio de Janeiro, RJ, Brazil 8 Department of Plant Biology, Southern Illinois University, Carbondale, Illinois 62901-6509, United States of America 9 CMNS-Cell Biology and Molecular Genetics, 2107 Bioscience Research Building, University of Maryland, College Park, MD 20742-4451, United States of America 10 Apartado 5-1500, Acosta, Costa Rica 11 Systematic Botany and Mycology, Ludwig Maximilian University of Munich, Menzinger Str. 67, 80638 Munich, Germany 12 Allan Herbarium, Landcare Research, P O Box 69-040, Lincoln 7608, New Zealand 13 Muséum National d’Histoire Naturelle, Department Systématique et Evolution, C.P. 39, 57 Rue Cuvier, 75231 Paris 05, France 14 Botany Unit, Finnish Museum of Natural History, University of Helsinki, P.O. Box 7, Helsinki FI-00014, Finland 15 Department of Systematic Botany with Herbarium Haussknecht and Botanical Garden, Friedrich Schiller University, Fürstengraben 1, 07737 Jena, Germany 16 Museu Paraense Emilio Goeldi, Coordenaçao de Botanica, Av. Magalhaes Barata 376, 66040-1 70 Belem, Para, Brazil 17 Department of Biological Science, Graduate School of Science, Hiroshima University, Kagamiyama 1–3–1, Higashihiroshimashi, Hiroshima 739–8526, Japan 18 N.A. Avrorin Polar-Alpine Botanical Garden–Institute of Kola SC RAS, 184236 Kirovsk-6, Russia 19 Royal Botanic Garden, Edinburgh EH3 5LR, United Kingdom 20 Staatliches Copyright Lars Söderström et al. This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited. 2 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Museum für Naturkunde Stuttgart, Rosenstein 1, 70191 Stuttgart, Germany 21 Botany Department, Institute of Biology, Eszterházy Károly College, Eger, Pf. 43, H-3301, Hungary 22 Departamento de Botánica, Facultad de Ciencias Biológicas, Universitat de València. C/ Dr. Moliner 50, E-46100, Burjassot (Valencia), Spain 23 Albrecht-von-Haller-Institut für Planzenwissenschaften, Department of Systematics, Biodiversity and Evolution of Plants, Untere Karspüle 2, 37073 Göttingen, Germany 24 Mittlere Letten 11, 88634 Herdwangen-Schönach, Germany 25 Centro de Investigaciones sobre Desertiicación (CIDE-CSIC-UV-GV), C/ Carretera de Moncada-Náquera Km. 4.5, E-46113, Moncada (Valencia), Spain 26 Department of Biology, Faculty of Science, Burapha University, Mueang, 20131 Chonburi, hailand 27 William and Lynda Steere Herbarium, he New York, Botanical Garden, Bronx, New York 10458-5126, United States of America 28 Instituto de Ciencias Naturales. Universidad Nacional de Colombia. Apartado 7495, Bogotá D.C., Colombia 29 Department of Botany, Charles University, Benátská 2, CZ-128 01 Praha 2, Czech Republic 30 Department of Biology, Ludwig-Maximilians-Universität, Menzinger Str. 67, D-80638, München, Germany 31 36, Big Green, Warmington, Peterborough PE8 6TU, United Kingdom 32 Shenzhen Key Laboratory of Southern Subtropical Plant Diversity, Fairylake Botanical Garden, 160 Xianhu Rd., Liantang, Shenzhen 518004, Guangdong, China 33 Department of Biology, School of Life Sciences, East China Normal University, 3663 Zhong Shan North Road, Shanghai 200062, China * Editors Corresponding author: Lars Söderström (lars.soderstrom@ntnu.no) Academic editor: Lyubomir Penev | Received 20 June 2015 | Accepted 25 September 2015 | Published 29 January 2016 Citation: Söderström L, Hagborg A, von Konrat M, Bartholomew-Began S, Bell D, Briscoe L, Brown E, Cargill DC, Costa DP, Crandall-Stotler BJ, Cooper ED, Dauphin G, Engel JJ, Feldberg K, Glenny D, Gradstein SR, He X, Heinrichs J, Hentschel J, Ilkiu-Borges AL, Katagiri T, Konstantinova NA, Larraín J, Long DG, Nebel M, Pócs T, Felisa Puche F, Reiner-Drehwald E, Renner MAM, Sass-Gyarmati A, Schäfer-Verwimp A, Moragues JGS, Stotler RE, Sukkharak P, hiers BM, Uribe J, Váňa J, Villarreal JC, Wigginton M, Zhang L, Zhu R-L (2015) World checklist of hornworts and liverworts. PhytoKeys 59: 1–828. doi: 10.3897/phytokeys.59.6261 his checklist is dedicated to Elizabeth Brown and Ray Stotler, who sadly left us during our work with it. “……Charles Darwin wrote of his desire to provide inancial support ‘for the formation of a perfect M.S. catalogue of all known plants’ (Darwin 1881, letter 13570). It is a personal embarrassment to me, and should be chastening to us all, that more than 120 years later we still have not delivered on that commitment.” (Crane 2004). Abstract A working checklist of accepted taxa worldwide is vital in achieving the goal of developing an online lora of all known plants by 2020 as part of the Global Strategy for Plant Conservation. We here present the irst-ever worldwide checklist for liverworts (Marchantiophyta) and hornworts (Anthocerotophyta) that includes 7486 species in 398 genera representing 92 families from the two phyla. he checklist has far reaching implications and applications, including providing a valuable tool for taxonomists and systematists, analyzing phytogeographic and diversity patterns, aiding in the assessment of loristic and World checklist of hornworts and liverworts 3 taxonomic knowledge, and identifying geographical gaps in our understanding of the global liverwort and hornwort lora. he checklist is derived from a working data set centralizing nomenclature, taxonomy and geography on a global scale. Prior to this efort a lack of centralization has been a major impediment for the study and analysis of species richness, conservation and systematic research at both regional and global scales. he success of this checklist, initiated in 2008, has been underpinned by its community approach involving taxonomic specialists working towards a consensus on taxonomy, nomenclature and distribution. Keywords Marchantiophyta, Anthocerophyta, nomenclature, taxonomy Introduction he natural world is changing fast (Balmford et al. 2005). he Global Strategy for Plant Conservation (GSPC) was adopted by the Conference of Parties (COP) of the Convention on Biological Diversity (CBD) in April 2002 (http://www.cbd.int/decision/cop/?id=7183; accessed 2014.06.02). he Strategy set out 16 outcome-oriented targets that were to be achieved by 2010. he GSPC was designed as a framework for action to halt the loss of plant diversity. Target 1 of the Strategy was to complete “a widely accessible working list of all known plant species, as a step towards a complete world Flora” (Lughadha 2004). As early as 1881 Charles Darwin expressed a wish to have a catalogue of all known plants (Crane 2004). However, over 125 years later, this wish is not yet fulilled. Without a working checklist, many of the other objectives in the GSPC cannot be met and botanists around the world cannot communicate about plants on a global basis (Crane 2004). A working list of known plant species is critical for (I) its underpinning role in the efective implementation of the other targets through provision of baseline information; (II) increasing the accessibility and use of accurate botanical name information for research, conservation and sustainable use; and (III) real-world politics and how taxonomists respond to the decisions of policy makers (Paton et al. 2008). Version 1.0 of he Plant List (http://www.theplantlist.org/) was released in December 2010 aimed to be comprehensive for species of vascular plants (lowering plants, conifers, ferns and their allies) and of bryophytes (mosses, liverworts and hornworts), as a response to the 2010 Target 1 of the GSPC and to a clear global need for such data. he Plant List is a broad collaboration, coordinated by the Royal Botanic Gardens, Kew, and the Missouri Botanical Garden (MO), involving diverse partnerships. Paton (2013) noted he Plant List represents a work in progress with future versions planned to improve the quality of names and decrease the number of unresolved names. Target 1 has since been revised and agreed at the 10th COP of the CBD in Nagoya, Japan, with the goal of developing ‘‘an online lora of all known plants’’ by 2020 (http://www.cbd.int/decision/cop/default.shtml?id=12283; accessed 2014.06.02). Paton (2013) identiied several factors that might assist in the implementation of the 4 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) revised target, stressing that achieving a working list for plant taxa worldwide was a vital step to an online lora. Earlier, Paton et al. (2008) assessed the progress made at that time and discussed prospects for the completion of Target 1. Paton noted that good progress had been made in bryophytes (mosses, liverworts and hornworts), ferns and gymnosperms with widely accessible working lists either complete or almost so for those groups. For bryophytes alone, he tabulated 13,370 accepted species noting that the data was largely derived from Tropicos. However, although Tropicos is an indispensable reference for anyone dealing with bryophyte names and the database is very strong for mosses, the nomenclatural and auxiliary data for liverworts is less complete, especially for larger genera (von Konrat et al. 2008a). he lack of a central source providing a synthesis of nomenclatural data and global distributional data was the impetus toward developing the current checklist of liverworts and hornworts (von Konrat et al. 2008a, 2010d, Söderström et al. 2012e). Checklists are powerful and important tools that can integrate the almost overwhelmingly scattered information concerning taxonomy, systematics, nomenclature, distribution, and even frequency (Söderström et al. 2008). Ecological and biological significance of liverworts and hornworts Liverworts and hornworts (Figure 1) are of critical biological, ecological, and phylogenetic signiicance (e.g. Asakawa 1998, Longton 1992, Hallingbäck and Hodgetts 2000, Gradstein et al. 2001b, Wellman et al. 2003, Qiu et al. 2007). Liverworts are found on soil, rocks, and trees throughout the world, from coastal Antarctica to the tundra of the Northern Hemisphere, and from semi-arid areas of Australia to the Amazon rainforests (Hallingbäck and Hodgetts 2000). Although there are xero-tolerant taxa, the majority of liverworts are found in relatively humid and shaded terrestrial ecosystems (Gradstein and Costa 2003). Liverworts and hornworts are an important component of the vegetation in many regions of the world, constituting a major part of the biodiversity in moist forest, wetlands, mountain, and tundra ecosystems (Hallingbäck and Hodgetts 2000). Liverworts, hornworts and mosses ofer microhabitats that are critical to the survival of a tremendous diversity of organisms such as single-celled eukaryotes, protozoa, and numerous groups of invertebrates (Gerson 1982). heir structural contribution to levels of diversity might be as signiicant as that of vascular plants, albeit at a smaller scale (von Konrat et al. 2008a). hese plants are also important environmental and ecological indicators (Rao 1982, Pitcairn et al. 1995, Gradstein et al. 2001b, Giordano et al. 2004). Liverworts, in concert with mosses and hornworts, play a signiicant role in the global carbon budget (O’Neill 2000) and CO2 exchange (De Lucia et al. 2003), plant succession (Cremer and Mount 1965), net production and phytomass (Frahm 1990), nutrient cycling (Coxson 1991), and water retention (Pócs 1980a, Gradstein et al. 2001b). hese groups of land plants also have been used as indicators of past climate change, to validate climate models, and as potential indicators of global warming (Gignac 2001, Tuba et al. 2011). World checklist of hornworts and liverworts 5 Figure 1. Habit images of selected lineages of Anthocerotophyta (hornworts) and Marchantiophyta (liverworts). A Phaeomegaceros squamuliger, photo by J. Hollinger B Haplomitrium mnioides, photo by L. Zhang C Conocephalum japonicum, photo by D. G. Long D Colura vitiensis, photo by T. Pócs E Pleurozia gigantea, photo by L. Söderström F Riccardia spegazziniana, photo by Juan Larraín. Species numbers here are estimated to exist only 215 hornwort species. By comparison estimates of liverwort species richness have varied considerably, by as much as 50%, with estimates ranging from 4,500 to 9,000 (e.g. Pearson 1995, Forrest et al. 2006). In the last decade or so estimates in the range of 5,000 to 6,000 liverwort species have been widely accepted (e.g. Gradstein and Costa 2003, Gradstein and Ilkiu-Borges 2009, Heinrichs et al. 2007). von Konrat et al. (2010a) provided a mean estimate of 7,500 6 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) for the number of liverwort species based on estimating rates of synonymy in a sample of recently monographed and revised taxa. A standardized global worldwide liverwort checklist with strong community participation, as presented here, coupled with the critical need for ongoing monographs and revisions, will aid in arriving at clearer estimates of liverwort and hornwort diversity. Signiicantly, a list of standardized names reviewed by taxonomic experts will enable more meaningful geographical and biogeographical species comparative studies. he global checklist of accepted liverwort and hornwort taxa has vast potential, not only in aiding our understanding of liverwort diversity, patterns, and processes, but also to the broader biological community (von Konrat et al. 2008a). Below we provide context to the worldwide checklist, including a brief historical account and current informational resources, structure and layout, systematic and classiication concepts, and methodology, including a brief overview of the underlying data set and nomenclatural elements. Major historical works Schuster (1966d) has provided a detailed account on the history of hepaticology, up to the irst half of the 20th century. Earlier, Verdoorn (1934c) also outlined a valuable historical review of important igures in hepaticology. In the early 19th century, possibly the two most inluential persons who provided an early framework for taxonomic concepts were W.J. Hooker and G. Raddi. During the middle of the 19th century C.G. Nees von Esenbeck, C.M. Gottsche and J.B.W. Lindenberg provided vastly signiicant contributions towards developing a classiication framework. Frahm and Eggers (2001) provided a brief biography of these workers. his was later followed by R. Spruce and the Austrian-born H. Leitgeb in the second half 19th century. Leitgeb in particular was instrumental in liverwort plant morphology and anatomy including a six part treatise, “Untersuchungen ueber die Lebermoose”, from 1874 to 1881. Towards the end of 19th century and early 20th century V. Schifner, F. Stephani, and A. Evans were prominent hepaticologists. In the irst half of the 20th century A. Evans, H. Buch, F. Verdoorn, and K. Müller (Frib.) were leading inluential hepaticologists. In the second half of the 20th century, R. Grolle and R.M. Schuster were of extraordinary inluence in working towards contemporary classiication schemes. Many of these groundbreaking concepts still serve the foundation from which our current classiication scheme is derived. R.M. Schuster’s major contribution to botany and hepaticology lies in the astounding new diversity of liverworts he added to our knowledge. He described a staggering 463 species, 83 genera and 15 families new to science. It is diicult to name another contemporary botanist who discovered this much new diversity of a major clade of land plants (Qiu et al. 2013). R.M. Schuster’s contribution to botany went beyond hepatics. He was one of the irst botanists who recognized the importance of Wallace’s Line in plant biogeography, separating Australia of Gondwanaland from Southeast Asia of Laurasia (Cronquist 1988). World checklist of hornworts and liverworts 7 Figure 2. Number of novel liverwort species, excluding new combinations, which have been described over the last 250 years, with an inset of the number described from 2001–2012. Figure 2 depicts the number of novel liverwort species, excluding new combinations that have been described over the last 250 years. he irst major peak corresponds to the works of several early 19th century botanists, including Synopsis Hepaticarum by Gottsche, Lindenberg, and Nees (1844–1847). he three decades leading into, but prior to the highest peak, between 1860 and 1890, correspond to publications by a number of prominent bryologists including W. Mitten, J.D. Hooker, T. Taylor, and V. Schifner. he second and highest peak of almost 1,200 names, in the early 1900s corresponds largely to the plethora of taxa described by F. Stephani (1898–1924) in his monumental work Species Hepaticarum. he periods of highest rates of new species described in the 1830s and around 1900 are the same for seed plants (Mutke and Barthlott 2005). he third peak over the four decades between 1950 and 1980 can be attributed mainly to the works of R.M. Schuster, H. Inoue, and S. Hattori. H. Inoue and S. Hattori were inluential on large species-rich genera such as Plagiochila, Frullania, and Porella. he decline in newly described species since 1970 does not necessarily translate to the conclusion that taxonomists are closer to discovering all known species. he almost 200 novel liverwort species that have been described in the past six years alone still represent a signiicant number, considering the relatively few liverwort taxonomists and monographers. Moreover, bryological exploration has been very uneven in many parts of the world. For example, many areas of the Neotropics still remain without a single bryophyte record (Gradstein et al. 2001a). Paradoxically, scores of new species are still being discovered and described in relatively well-studied 8 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) areas such as New Zealand, e.g. over 50 new taxa since 2000 alone (e.g. Engel and Schuster 2001, Engel and Glenny 2008a, Renner and de Lange 2011, von Konrat et al. 2012b). Recent attention to cryptic speciation in bryophytes is also revealing novel liverwort species (e.g. Szweykowski et al. 2005). he combination of collecting in yetto-be explored areas, the continued discovery of species in well-studied regions, and an increased understanding of the biology of liverworts (including cryptic speciation), will lead to a signiicant number of newly discovered species in the foreseeable future. he corollary of this, coupled with increased monographic and revisional work, will be the increased discovery and the unravelling of synonymy. Contemporary resources Data availability and information needs associated with liverworts and hornworts have been reviewed extensively by von Konrat et al. (2008a, 2010a). A fundamental problem common to all nomenclatural indexing projects is the dispersed nature of the biological literature, some of which may date back over 250 years (Lughadha 2004). Our own data records, from 1990 to the present alone, indicate there are over 190 periodical titles and non-serials in which liverwort nomenclatural novelties were published. he most useful and successful web-accessible bryophyte nomenclatural database is Tropicos (http://www.tropicos.org), which ofers name data with references and type information as well as links to specimen data of its holdings. Tropicos has therefore become an indispensable reference for anyone dealing with bryophyte names. However, as stated above, the database is particularly strong for mosses and far less comprehensive for liverworts. Moreover, Tropicos does not prescribe taxonomic disposition of names, i.e., there is no attempt to adopt a single consistent view on the status of any particular name (Lughadha 2004, von Konrat et al. 2008a). A major nomenclatural work is the Index Hepaticarum (http://www.ville-ge.ch/ musinfo/bd/cjb/hepatic/index.php), which includes all efectively published liverwort epithets spanning 12 volumes with the closing date of 1973. he indices were prepared as a purely nomenclatural resource and did not claim to express any particular taxonomic concept. Recently, Crosby and Engel (2006) provided an equally valuable nomenclatural resource and catalog of names at all ranks for liverworts and hornworts published during 1974 to 2000. Subsequently, indices of the citations for names published for bryophytes have been compiled for the years 2001-2004 (Crosby and Magill 2005) and 2005 (Crosby and Magill 2006). he Early Land Plants Today initiative has continued this series (von Konrat et al. 2010c, Söderström et al. 2012b, 2014). he application of molecular phylogenetics continues to generate new insights into the evolutionary history of liverworts and hornworts. In recent years, inferences made from these phylogenies have especially revolutionized liverwort classiication (CrandallStotler et al. 2009). Yet, the reconstruction of the phylogenetic history of the Marchantiophyta and Anthocerotophyta remains an ongoing efort and the classiication is luid (von Konrat et al. 2010a). he current checklist largely follows the comprehensive World checklist of hornworts and liverworts 9 phylogenetic liverwort classiication scheme provided by Crandall- Stotler et al. (2009) and the hornwort classiication scheme of Stotler and Crandall-Stotler (2005). hus the checklist largely adheres to their higher classiication namely from class to suborder, and the majority of family and genera arrangements provided therein. However, there are some notable exceptions, especially in the systematic treatment of liverworts, which largely either relect publications subsequent to 2009, or are a slight departure in taxonomic opinion based on earlier works, e.g. the concepts of Saccogynaceae and Stephaniellaceae depart from Crandall-Stotler et al. (2009). Since 2010 there have been a number of realignments and novel hypotheses proposed that were generated from conclusions based on molecular evidence which have inluenced the systematic treatment presented here. hese include, for example, Adelanthaceae (e.g. Konstantinova and Vilnet 2009, Feldberg et al. 2010b, 2010a, Söderström et al. 2010b, Vilnet et al. 2010), Anastrophyllaceae (e.g. Söderström et. al 2010b), Gymnomitriaceae (e.g. Vilnet et al 2007a, 2010, Váňa et al. 2010b, Shaw et al. 2015), Lepidoziaceae (Cooper 2013), Lophocoleaceae (Söderström et al. 2013b), and Scapaniaceae (e.g. Vilnet et al. 2010, Heinrichs et al. 2012a). In all cases, any such departures, new families, or realignments are noted and very briely discussed throughout the checklist. As noted by Crandall-Stotler et al. (2009) many small families still remain to be investigated using molecular data, and many of the large families and genera that have been sampled appear to be either polyphyletic or paraphyletic. he systematic treatment as outlined here also includes the addition of subfamily and infrageneric classiication; ranks not presented by Crandall-Stotler et al. (2008b, 2009). Although there are arguably a number of ambiguous placements, especially at the infrageneric rank, and there remain many areas of contention, particularly at the familial and generic level, the systematic treatment itself serves as an excellent synopsis of contemporary taxonomic opinion, including infrageneric ranks. he infrageneric treatments will also aid future eforts in providing a taxonomic framework that will be particularly useful for extremely large genera. In large part, the infrageneric treatments only deal with those taxa that represent phylogenetic clades with strong support. Care has been taken to reduce ambiguity and those taxa where infrageneric placement is weak or without evidence are simply listed under the genus and not placed systematically. Structure and format of the checklist he checklist is presented in a taxonomic framework as outlined above with four main sections. he irst is arranged systematically with taxa ordered alphabetically within the nearest higher rank. A brief discussion for selected families is provided in the main body of the text under the respective families. his does not appear uniformly for all families, but typically for larger more complicated families. Rather, a brief description is provided mainly where concepts depart from Crandall-Stotler et al. (2009) or if new information has come to light since that publication. Subfamilies and infrageneric ranks have been incorporated where appropriate. However, if the subgeneric place- 10 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ment is unclear the species are listed under “Incertae sedis”. From the rank of genus and below, all entries include a three star conidence ranking (described below), accepted taxon name, authority, abbreviated publication title, page number and date, which is cross- referenced to the full citation in the reference list, as well as the basionym if appropriate. A typical entry appears as follows: *** *** Oleolophozia L.Söderstr., De Roo et Hedd., Phytotaxa 3: 50, 2010 (Söderström et al. 2010b). Oleolophozia perssonii (H.Buch et S.W.Arnell) L.Söderstr., De Roo et Hedd., Phytotaxa 3: 51, 2010 (Söderström et al. 2010b). Bas.: Lophozia perssonii H.Buch et S.W.Arnell, Bot. Not. 97: 382, 1944 (Buch 1944). here are comprehensive footnotes throughout the systematic section of the checklist. hese are applied to selected names representing the rank of genus and below only. Footnotes without any literature citation represent observations from the authors of the respective family. Footnotes are wide ranging in their content. Generally, footnotes can be categorized into the following: i) general statements relating to taxonomic opinion, ii) comments on the type specimen, iii) comments about possible conspeciicity, including citation of conlicting opinions, and iv) comment about species complexes. his section is followed by a list of taxa in genera that we do not recognize. hose names represent taxa that have not been studied recently and have not been recognized in any recent treatment. However, many of those names are old and may gain priority over some recognized taxa once their identity is determined. he next section is an alphabetical list of taxa of the ranks of species and below. he alphabetical list includes the conidence level, the taxon name and authority as well as a reference to the page of the corresponding entry in the systematic section. hus, the alphabetical list provides a rapid gateway requiring no prior knowledge of higher taxonomy, but also serves as an index to the corresponding name in the systematic section that includes more detailed information. he last section is a reference list where full bibliographic citations are given to all references for taxa included in the checklist. Titles and references have all been checked and veriied, except in a few cases where we could not get hold of the publications. Taxon confidence levels Signiicantly, each accepted taxon is qualiied using a three level ranking system that summarizes our knowledge about a taxon. he coding convention we are adopting largely follows that described by von Konrat et al. (2010d) using one to three stars, which has been applied to recent regional checklists produced by the Early Land Plants Today initiative, e.g. Söderström et al. (2010a, 2011a, 2011b, 2012g, 2013e). he conventions are briely outlined below coupled with samples illustrating how these conventions are applied in practice. he application of a conidence level to a taxon’s World checklist of hornworts and liverworts 11 status and whether it represents a genuine “species” that is reached through community consensus may go towards reining species estimates using an evidence-based approach (von Konrat et al. 2008a, 2008b). he conidence levels coupled with the detailed annotations in the form of footnotes may also provide a rapid assessment of taxa and help aid and drive future research into speciic taxonomic or nomenclatural problems and issues. he coding convention: * Serious doubts. here are doubts about the value of the taxon. It can be that there are conlicting views without any substantial evidence in any direction, conlicting views with substantiating evidence supporting one or both positions, or evidence pointing towards synonymization but it is premature to do it. We have adhered to the principle that it is better to keep a taxon with one star (and preferably a note) than to synonymize it too quickly. Example: Bazzania asymmetrica is conspeciic with Bazzania macgregorii in Grolle (1968a), but Kitagawa (1979a) kept them separate. Example: Nardia kamtschatica may be conspeciic with Nardia assamica (Váňa 1976c), but the type specimen could not be studied. ** Knowledge problem. he taxon is not well known by the person evaluating it. It may be a newly described species or a species originally not well described and not restudied recently. Example: Jungermannia erectii Ajit P.Singh et V.Nath was recently described and has not been independently studied by someone with a global overview. It is therefore diicult to evaluate. *** Accepted. A good taxon as currently understood based on personal experience or on taxonomic revisions that have been convincingly performed. Nomenclature and/or taxonomic position may, however, be questioned. Elements may be excluded from the taxon, but the taxon with the current type will still be accepted. Example: H. Bischler-Causse revised the genus Marchantia worldwide in a series of publications. Although she had a broad species concept, she also recognized infraspeciic taxa. he taxa that she recognized without doubt should be accepted unless new evidence against it exists. Adopting a narrower species concept so that many of her subspecies are elevated to species does not change her view of what a good taxon is. Methodology he foundation of the checklist is in the underlying data set from which it is derived. It was briely described by von Konrat et al. (2008a, 2010a). he working data sets now includes a bibliography of 25,000 publications; approximately 39,000 published liverwort names (including “accepted” taxa, synonyms, invalid and illegitimate names). he data quality and standards were outlined by von Konrat et al. (2010d). In summary, for authorities and for the citation abbreviations, we follow the standards set by the on-line version of Authors of Plant Names at the Royal Bota- 12 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) nical Gardens, Kew Website (http://www.ipni.org), with whom we collaborate closely and provide with updated data records. Publications and journal abbreviations follow Taxonomic Literature: A selective guide to botanical publications and collections with dates, commentaries and types (Staleu and Cowan 1976–2009) (http://www. sil.si.edu/digitalcollections/tl- 2/search.cfm) and Botanico Periodicum Huntianum Lawrence et al. (1968) (http://huntbot.andrew.cmu.edu/HIBD/Departments/Databases.shtml). here has been an intense systematic efort focusing on data quality. In all but a few cases nomenclatural data has been veriied against original publications. Söderström and Hagborg have checked and conirmed almost every original publication (three publications have not been available) for correct author, title and journal/ book citation, date of publication as far as possible, page number for the protologue, and if the name is validly and legitimately published according to the International Code of Nomenclature for algae, fungi and plants (Melbourne Code; McNeill et al. 2012). Most signiicantly, the checklist has been community driven and collaborative. Broader participation by taxonomic specialists and regional experts has lead to the checklist containing high quality data (cf. Söderström et al. 2008). Development of the checklist has included three international meetings (the irst one in 2009) and generated 74 published notes on taxonomy and nomenclature under the auspices of the Early Land Plants Today initiative from 2012 to 2015. he series of meetings were instrumental in providing a framework for direct interaction with taxonomic experts, workshops reviewing names, and helping identify potential participants who were taxonomic experts in speciic taxonomic groups, whether these were individual genera or entire families. he notes, published in the journal Phytotaxa, have included 38 authors from 13 countries. hese are the results of several years of revisions of taxonomic groups by taxonomic experts. Following these revisions and working with participants identifying nomenclatural and taxonomic problems that would impact the worldwide checklist, Söderström, Hagborg and von Konrat led editorial eforts in compiling manuscripts with baseline information and nomenclatural or taxonomic issues that required resolving. he work was then coordinated with the taxonomic experts who drove and led the process and often identiied further issues. he series of notes on taxonomy and nomenclature of liverworts and hornworts provided updated nomenclature, corrected invalid and illegitimate names and described new taxa based on studies (mainly molecular) that did not draw nomenclatural conclusions. hose notes are all open access and any changes were efective immediately as the Melbourne Code of Botanical Nomenclature allows publishing on internet from 1 Jan. 2012 (Knapp et al. 2011). his checklist builds on all published taxonomic and nomenclatural papers, that have come to our attention until June 30, 2015. World checklist of hornworts and liverworts 13 Summary statistics We here present the irst-ever worldwide checklist for liverworts (Marchantiophyta) and hornworts (Anthocerotophyta) that includes 8,078 taxa (species and below) in 7,486 species representing 398 genera, 92 families, 20 orders and 7 classes from the two phyla. he list includes 3,533 species with three stars, 2,988 species with two stars and 915 species with one star. he checklist also has extra utility in that it contains 3,106 references in the bibliography that serve as a powerful bibliographic resource for liverwort and hornwort systematic and taxonomic research. Concluding remarks he marked-up publication form of the current checklist by PhytoKeys provides a virtual instrument with a linked environment both internally (e.g. within an article) and externally (GBIF, IPNI, Tropicos, Wikispecies, etc.) that will undoubtedly help accelerate taxonomic research. he published checklist was the irst phase in providing a worldwide list of accepted names. he next phase is to establish a generally recognized online repository to augment the huge underlying informational auxiliary data of over 25,000 publications, almost 39,000 published names, and the over 700,000 geographical observations. Several features of Web-based technology make it an essential tool, including 1) a vehicle to facilitate data access, 2) to unify the vastly scattered data on distributional information and nomenclature, 3) ofering dynamic rather than static data in a searchable forum. he broader accessibility to the wealth of auxiliary data will help augment monographic and revisional work for many taxonomic groups, aid in identifying the need for increased loristic and survey work in many regions throughout the world, and have broad implications and applications beyond taxonomic research such as conservation science. he current Early Land Plants Today model has strong participation from taxonomic experts and an online resource will provide an opportunity to expand stakeholders to include ecologists, conservationists, scientists from other disciplines and general interest groups. However, such an efort can only be successful if it comes with sustained funding and infrastructure rather than depending on an ad hoc commitment by a few individuals, however dedicated. he project will help augment monographic and revisional work for many taxonomic groups and aid in identifying the need for increased loristic and survey work in many regions throughout the world. Although there are many challenges ahead to obtain high quality data, quantifying global liverwort diversity is a tractable, multifaceted and scientiically important goal, and everyone stands to gain by fostering this endeavour. he success of the project will lie on strong collaboration between institutions and the bryological community in general. 14 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Taxonomic synopsis Anthocerotophyta ................................................................................................. 27 Anthocerotopsida de Bary ex Jancz. .................................................................... 27 Anthocerotidae Rosenv. ................................................................................. 27 Anthocerotales Limpr. ............................................................................... 27 Anthocerotaceae Dumort. ...................................................................... 27 Anthoceros L. ................................................................................ 27 Folioceros D.C.Bharadwaj ........................................................... 31 Dendrocerotidae R.J.Duf .............................................................................. 32 Dendrocerotales Hässel .............................................................................. 32 Dendrocerotaceae J.Haseg. .................................................................... 32 Dendroceros Nees ......................................................................... 32 Megaceros Campb. ....................................................................... 27 Nothoceros (R.M.Schust.) J.Haseg............................................... 35 Phaeomegaceros R.J.Duf............................................................. 35 Phymatocerotales R.J.Duf ......................................................................... 36 Phymatocerotaceae R.J.Duf .................................................................. 36 Phymatoceros Stotler.................................................................... 36 Notothylatidae R.J.Duf................................................................................. 36 Notothyladales Hyvönen et Piippo ............................................................ 36 Notothyladaceae Müll.Frib. ex Prosk. .................................................... 36 Notothyladoideae Grolle ................................................................... 37 Notothylas Sull. ex A.Gray ........................................................... 37 Phaeocerotoideae Hässel .................................................................... 38 Mesoceros Piippo.......................................................................... 38 Paraphymatoceros Hässel ............................................................ 38 Phaeoceros Prosk. ........................................................................ 38 Leiosporocerotopsida Stotler et Crand.-Stotl. ................................................. 41 Leiosporocerotales Hässel ........................................................................... 41 Leiosporocerotaceae Hässel ex Ochyra ................................................... 41 Leiosporoceros Hässel................................................................... 41 Marchantiophyta ........................................................................................... 41 Haplomitriopsida Stotler et Crand.-Stotl. ...................................................... 41 Haplomitriidae Stotler et Crand.-Stotl. ...................................................... 41 Calobryales Hamlin ............................................................................... 41 Haplomitriaceae Dědeček .................................................................. 41 Haplomitrium Nees ..................................................................... 41 Treubiidae Stotler et Crand.-Stotl. ............................................................. 42 Treubiales Schljakov .............................................................................. 42 Treubiaceae Verd............................................................................... 42 Apotreubia S.Hatt. et Mizut. ....................................................... 42 Treubia K.I.Goebel...................................................................... 43 World checklist of hornworts and liverworts 15 Jungermanniopsida Stotler et Crand.-Stotl. ........................................................ 43 Jungermanniidae Engl.................................................................................... 43 Jungermanniales H.Klinggr........................................................................ 43 Cephaloziineae Schljakov ....................................................................... 43 Adelanthaceae Grolle ......................................................................... 43 Adelanthoideae K.Feldberg ............................................................ 43 Adelanthus Mitt. ...................................................................... 44 Pseudomarsupidium Herzog ................................................... 44 Wettsteinia Schifn. ................................................................. 45 Jamesonielloideae Inoue ................................................................ 45 Cuspidatula Steph. .................................................................. 45 Denotarisia Grolle ................................................................... 46 Nothostrepta R.M.Schust. ....................................................... 46 Pisanoa Hässel ......................................................................... 46 Protosyzygiella (Inoue) R.M.Schust. ....................................... 46 Syzygiella Spruce...................................................................... 46 Vanaea (Inoue et Gradst.) Inoue et Gradst. ........................... 49 Anastrophyllaceae L.Söderstr. ............................................................ 49 Anastrepta (Lindb.) Schifn. .................................................... 50 Anastrophyllum (Spruce) Steph............................................... 50 Barbilophozia Loeske .............................................................. 51 Biantheridion (Grolle) Konstant. et Vilnet ............................ 52 Chandonanthus Mitt. .............................................................. 52 Crossocalyx Meyl. ..................................................................... 52 Gymnocolea (Dumort.) Dumort. ............................................ 52 Hamatostrepta Váňa et D.G.Long .......................................... 52 Hattoria R.M.Schust. .............................................................. 53 Isopaches H.Buch ..................................................................... 53 Neoorthocaulis L.Söderstr. ...................................................... 53 Orthocaulis H.Buch ................................................................. 53 Plicanthus R.M.Schust. ........................................................... 54 Schizophyllopsis Váňa et L.Söderstr. ....................................... 54 Schljakovia Konstant. et Vilnet ............................................... 54 Schljakovianthus Konstant. et Vilnet ...................................... 55 Sphenolobopsis R.M.Schust. et N.Kitag. ................................. 55 Sphenolobus (Lindb.) Berggr................................................... 55 Tetralophozia (R.M.Schust.) Schljakov .................................. 55 Zantenia (S.Hatt.) Váňa et J.J.Engel....................................... 56 Cephaloziaceae Mig. .......................................................................... 56 Alobielloideae R.M.Schust............................................................. 56 Alobiella (Spruce) Schifn. ....................................................... 56 Alobiellopsis R.M.Schust. ........................................................ 56 Cephalozioideae Müll.Frib. ........................................................... 57 16 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Cephalozia (Dumort.) Dumort. .............................................. 57 Fuscocephaloziopsis Fulford .................................................... 59 Nowellia Mitt. .......................................................................... 61 Odontoschismatoideae H.Buch ex Grolle ...................................... 62 Odontoschisma (Dumort.) Dumort. ....................................... 62 Schifnerioideae R.M.Schust. ......................................................... 64 Schifneria Steph...................................................................... 64 Trabacelluloideae R.M.Schust. ...................................................... 64 Haesselia Grolle et Gradst. ...................................................... 64 Trabacellula Fulford ............................................................... 64 Cephaloziellaceae Douin.................................................................... 64 Allisoniella E.A.Hodgs. ............................................................ 65 Amphicephalozia R.M.Schust.................................................. 65 Anastrophyllopsis (R.M.Schust.) Váňa et L.Söderstr. ............. 65 Cephalojonesia Grolle.............................................................. 66 Cephalomitrion R.M.Schust. ................................................... 66 Cephaloziella (Spruce) Schifn. ............................................... 66 Cephaloziopsis (Spruce) Schifn. ............................................. 72 Chaetophyllopsis R.M.Schust. ................................................. 72 Cylindrocolea R.M.Schust. ...................................................... 72 Gottschelia Grolle .................................................................... 74 Gymnocoleopsis (R.M.Schust.) R.M.Schust. ........................... 74 Herzogobryum Grolle .............................................................. 74 Kymatocalyx Herzog ................................................................ 74 Lophonardia R.M.Schust. ....................................................... 75 Nothogymnomitrion R.M.Schust............................................. 75 Obtusifolium S.W.Arnell ......................................................... 75 Oleolophozia L.Söderstr. ......................................................... 75 Phycolepidozia R.M.Schust. .................................................... 75 Protolophozia (R.M.Schust.) Schljakov .................................. 75 Lophoziaceae Cavers .......................................................................... 77 Andrewsianthus R.M.Schust. .................................................. 77 Gerhildiella Grolle................................................................... 78 Heterogemma (Jørg.) Konstant. et Vilnet ............................... 78 Lophozia (Dumort.) Dumort. ................................................. 79 Lophoziopsis Konstant. et Vilnet ............................................. 80 Pseudocephaloziella R.M.Schust. ............................................ 81 Trilophozia (R.M.Schust.) Bakalin ......................................... 81 Tritomaria Schifn. ex Loeske ................................................. 82 Scapaniaceae Mig............................................................................... 82 Diplophyllum (Dumort.) Dumort........................................... 82 Douinia (C.E.O.Jensen) H.Buch ............................................ 84 Pseudotritomaria Konstant. et Vilnet ..................................... 84 World checklist of hornworts and liverworts 17 Saccobasis H.Buch ................................................................... 85 Scapania (Dumort.) Dumort. ................................................. 85 Schistochilopsis (N.Kitag.) Konstant. ...................................... 92 Jungermanniineae R.M.Schust. ex Stotler et Crand.-Stotl...................... 93 Acrobolbaceae E.A.Hodgs.................................................................. 93 Enigmella G.A.M.Scott et K.G.Beckm. ................................... 93 Acrobolboideae R.M.Schust. ex Briscoe ......................................... 93 Acrobolbus Nees ....................................................................... 93 Austrolophozioideae R.M.Schust. ex Crand.-Stotl. ........................ 96 Austrolophozia R.M.Schust. .................................................... 96 Goebelobryum Grolle............................................................... 97 Lethocoleoideae Grolle .................................................................. 97 Lethocolea Mitt. ....................................................................... 97 Saccogynidioideae Crand.-Stotl. .................................................... 97 Saccogynidium Grolle .............................................................. 97 Antheliaceae R.M.Schust. .................................................................. 98 Anthelia (Dumort.) Dumort. .................................................. 98 Arnelliaceae Nakai ............................................................................. 99 Arnellia Lindb. ........................................................................ 99 Balantiopsidaceae H.Buch ................................................................. 99 Acroscyphella N.Kitag. et Grolle.............................................. 99 Pseudoisotachis Váňa ............................................................... 99 Balantiopsidoideae J.J.Engel et Váňa ............................................. 99 Balantiopsis Mitt. .................................................................... 99 Isotachidoideae Grolle ................................................................. 101 Isotachis Mitt. ........................................................................ 101 Neesioscyphus Grolle .............................................................. 102 Ruizanthoideae R.M.Schust. ex J.J.Engel et G.L.Merr. ................ 103 Ruizanthus R.M.Schust. ........................................................ 103 Blepharidophyllaceae R.M.Schust. ex J.J.Engel ................................ 103 Blepharidophyllum Ångstr. ................................................... 103 Clandarium (Grolle) R.M.Schust. ........................................ 103 Calypogeiaceae Arnell ...................................................................... 103 Calypogeia Raddi ................................................................... 103 Eocalypogeia (R.M.Schust.) R.M.Schust............................... 107 Metacalypogeia (S.Hatt.) Inoue............................................. 107 Mizutania Furuki et Z.Iwats. ................................................ 107 Mnioloma Herzog .................................................................. 107 Endogemmataceae Konstant. ........................................................... 109 Endogemma Konstant............................................................ 109 Geocalycaceae H.Klinggr. ................................................................ 109 Geocalyx Nees ........................................................................ 109 Gymnomitriaceae H.Klinggr. .......................................................... 109 18 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Acrolophozia R.M.Schust. ..................................................... 110 Nanomarsupella R.M.Schust. ex A.Hagborg ........................ 110 Paramomitrion R.M.Schust. ................................................. 110 Gymnomitrioideae T.Jensen........................................................ 110 Cryptocoleopsis Amakawa ...................................................... 110 Gymnomitrion Corda ............................................................ 110 Marsupella Dumort. .............................................................. 112 Poeltia Grolle ......................................................................... 114 Prasanthus Lindb. ................................................................. 114 Nardioideae Váňa ........................................................................ 114 Nardia Gray ........................................................................... 114 Gyrothyraceae R.M.Schust. ............................................................. 115 Gyrothyra M.Howe................................................................ 115 Harpanthaceae Arnell ...................................................................... 116 Harpanthus Nees ................................................................... 116 Hygrobiellaceae Konstant. et Vilnet................................................. 116 Hygrobiella Spruce ................................................................ 116 Jackiellaceae R.M.Schust. ................................................................ 116 Jackiella Schifn. .................................................................... 116 Jungermanniaceae Rchb. ................................................................. 117 Delavayelloideae Grolle ............................................................... 117 Delavayella Steph. ................................................................. 117 Liochlaena Nees ..................................................................... 117 Jungermannioideae Dumort. ....................................................... 118 Eremonotus Lindb. et Kaal. ex Pearson ................................ 118 Jungermannia L. .................................................................... 118 Mesoptychioideae R.M.Schust. ................................................... 120 Mesoptychia (Lindb.) A.Evans ............................................... 120 Rivulariella D.H.Wagner ...................................................... 122 Notoscyphaceae Crand.-Stotl. ......................................................... 122 Notoscyphus Mitt. .................................................................. 122 Saccogynaceae Heeg ........................................................................ 122 Saccogyna Dumort. ................................................................ 122 Solenostomataceae Stotler et Crand.-Stotl. ...................................... 123 Aponardia (R.M.Schust.) Váňa ............................................. 123 Arctoscyphus Hässel ............................................................... 123 Cryptocolea R.M.Schust. ....................................................... 123 Diplocolea Amakawa ............................................................. 123 Solenostoma Mitt. .................................................................. 123 Southbyaceae Váňa .......................................................................... 135 Gongylanthus Nees ................................................................ 135 Southbya Spruce .................................................................... 135 Stephaniellaceae R.M.Schust. .......................................................... 136 World checklist of hornworts and liverworts 19 Stephaniella J.B.Jack ............................................................. 136 Stephaniellidium S.Winkl. ex Grolle .................................... 136 Trichotemnomataceae R.M.Schust. ................................................. 136 Trichotemnoma R.M.Schust.................................................. 136 Lophocoleineae Schljakov .................................................................... 137 Blepharostomataceae W.Frey et M.Stech ......................................... 137 Blepharostoma (Dumort.) Dumort. ...................................... 137 Brevianthaceae J.J.Engel et R.M.Schust. .......................................... 137 Brevianthus J.J.Engel et R.M.Schust. ................................... 137 Tetracymbaliella Grolle ......................................................... 137 Chonecoleaceae R.M.Schust. ex Grolle ............................................ 138 Chonecolea Grolle .................................................................. 138 Grolleaceae Solari ex R.M.Schust..................................................... 138 Grollea R.M.Schust. .............................................................. 138 Herbertaceae Müll.Frib. ex Fulford et Hatcher ................................ 138 Herbertus Gray ...................................................................... 138 Triandrophyllum Fulford et Hatcher .................................... 171 Lepicoleaceae R.M.Schust................................................................ 141 Lepicolea Dumort. ................................................................. 142 Vetaforma Fulford et J.Taylor............................................... 142 Lepidoziaceae Limpr. ....................................................................... 142 Meinungeria Frank Müll. ...................................................... 143 Bazzanioideae Rodway................................................................. 143 Acromastigum A.Evans .......................................................... 143 Bazzania Gray ....................................................................... 146 Mastigopelma Mitt................................................................. 163 Drucelloideae R.M.Schust. .......................................................... 163 Drucella E.A.Hodgs. .............................................................. 163 Lembidioideae R.M.Schust.......................................................... 163 Dendrolembidium Herzog..................................................... 163 Hygrolembidium R.M.Schust. ............................................... 164 Isolembidium R.M.Schust. .................................................... 164 Kurzia G.Martens .................................................................. 164 Lembidium Mitt. ................................................................... 168 Megalembidium R.M.Schust. ................................................ 168 Pseudocephalozia R.M.Schust. .............................................. 168 Lepidozioideae Müll.Frib. ........................................................... 169 Ceramanus E.D.Cooper ........................................................ 169 Lepidozia (Dumort.) Dumort. .............................................. 169 Neolepidozia Fulford et J.Taylor........................................... 176 Tricholepidozia (R.M.Schust.) E.D.Cooper ......................... 179 Micropterygioideae Grolle ........................................................... 181 Micropterygium Gottsche ...................................................... 181 20 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Mytilopsis Spruce ................................................................... 182 Protocephalozioideae R.M.Schust................................................ 182 Protocephalozia (Spruce) K.I.Goebel.................................... 182 Zoopsidoideae R.M.Schust. ......................................................... 183 Amazoopsis J.J.Engel et G.L.Merr. ........................................ 183 Hyalolepidozia S.W.Arnell ex Grolle .................................... 183 Monodactylopsis (R.M.Schust.) R.M.Schust. ........................ 183 Neogrollea E.A.Hodgs............................................................ 183 Odontoseries Fulford ............................................................. 183 Paracromastigum Fulford et J.Taylor ................................... 183 Psiloclada Mitt....................................................................... 185 Pteropsiella Spruce ................................................................ 185 Telaranea Spruce ex Schifn. ................................................. 185 Zoopsidella R.M.Schust. ........................................................ 187 Zoopsis Hook.f. ex Gottsche .................................................. 188 Lophocoleaceae Vanden Berghen..................................................... 189 Bragginsella R.M.Schust........................................................ 189 Chiloscyphus Corda................................................................ 189 Clasmatocolea Spruce ............................................................ 191 Conoscyphus Mitt. .................................................................. 193 Cryptolophocolea L.Söderstr.................................................. 193 Deceptifrons J.J.Engel et Váňa............................................... 196 Evansianthus R.M.Schust. et J.J.Engel ................................. 196 Hepatostolonophora J.J.Engel et R.M.Schust. ...................... 196 Heteroscyphus Schifn. ........................................................... 197 Lamellocolea J.J.Engel ........................................................... 204 Leptophyllopsis R.M.Schust. .................................................. 204 Leptoscyphopsis R.M.Schust. ................................................. 204 Leptoscyphus Mitt. ................................................................. 204 Lophocolea (Dumort.) Dumort. ............................................ 208 Otoscyphus J.J.Engel, Bardat et houvenot ........................... 215 Pachyglossa Herzog et Grolle ................................................ 216 Perdusenia Hässel .................................................................. 216 Pigafettoa C.Massal. .............................................................. 217 Platycaulis R.M.Schust.......................................................... 217 Stolonivector J.J.Engel ........................................................... 217 Xenocephalozia R.M.Schust. ................................................. 217 Mastigophoraceae R.M.Schust......................................................... 217 Dendromastigophora R.M.Schust. ........................................ 217 Mastigophora Nees ................................................................ 217 Plagiochilaceae Müll.Frib................................................................. 218 Acrochila R.M.Schust. ........................................................... 218 Chiastocaulon Carl ................................................................ 219 World checklist of hornworts and liverworts 21 Dinckleria Trevis. .................................................................. 219 Pedinophyllopsis R.M.Schust. et Inoue ................................. 219 Pedinophyllum Lindb. ex Nordst. ......................................... 219 Plagiochila (Dumort.) Dumort. ............................................ 219 Plagiochilidium Herzog......................................................... 252 Plagiochilion S.Hatt............................................................... 252 Pseudolophocolea R.M.Schust. et J.J.Engel ........................... 253 Xenochila R.M.Schust. .......................................................... 253 Pseudolepicoleaceae Fulford et J.Taylor ........................................... 253 Archeophylla R.M.Schust....................................................... 253 Castanoclobos J.J.Engel et Glenny ........................................ 254 Chaetocolea Spruce ................................................................ 254 Herzogiaria Fulford ex Hässel............................................... 254 Isophyllaria E.A.Hodgs. et Allison ........................................ 254 Pseudolepicolea Fulford et J.Taylor ...................................... 254 Temnoma Mitt. ...................................................................... 255 Trichocoleaceae Nakai ..................................................................... 256 Eotrichocolea R.M.Schust...................................................... 256 Leiomitra Lindb. ................................................................... 256 Trichocolea Dumort. ............................................................. 257 Myliineae J.J.Engel et Braggins ex Crand.-Stotl. .................................. 258 Myliaceae Schljakov ......................................................................... 258 Mylia Gray ............................................................................. 258 Perssoniellineae R.M.Schust. ............................................................... 259 Schistochilaceae H.Buch .................................................................. 259 Schistochila Dumort. ............................................................. 259 Porellales Schljakov .................................................................................. 264 Jubulineae Müll.Frib. .......................................................................... 264 Frullaniaceae Lorch.......................................................................... 264 Frullania Raddi ..................................................................... 264 Jubulaceae H.Klinggr....................................................................... 295 Jubula Dumort. ..................................................................... 295 Neohattoria Kamim. .............................................................. 296 Nipponolejeunea S.Hatt. ....................................................... 296 Lejeuneaceae Cavers ........................................................................ 297 Lejeuneoideae ............................................................................. 297 trib. Brachiolejeuneeae ................................................................ 297 subtrib. Brachiolejeuneinae Gradst. ............................................. 297 Acanthocoleus R.M.Schust. .................................................... 297 Blepharolejeunea S.W.Arnell ................................................ 298 Brachiolejeunea (Spruce) Schifn. ......................................... 298 Dicranolejeunea (Spruce) Schifn. ........................................ 299 Lindigianthus Kruijt et Gradst. ............................................. 299 22 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Odontolejeunea (Spruce) Schifn. ......................................... 299 subtrib. Stictolejeuneinae Gradst. ................................................ 299 Neurolejeunea (Spruce) Schifn............................................. 299 Stictolejeunea (Spruce) Schifn. ............................................ 300 trib. Lejeuneeae Dumort. ............................................................ 300 Dactylophorella R.M.Schust.................................................. 300 Metalejeunea Grolle .............................................................. 300 Pictolejeunea Grolle .............................................................. 300 subtrib. Ceratolejeuneinae Gradst. ............................................... 301 Ceratolejeunea (Spruce) J.B.Jack et Steph. ........................... 301 Luteolejeunea Piippo ............................................................. 304 Otigoniolejeunea (Spruce) Schifn. ....................................... 304 subtrib. Cheilolejeuneinae Gradst. ............................................... 304 Aureolejeunea R.M.Schust. ................................................... 304 Cheilolejeunea (Spruce) Steph. ............................................. 305 Cyrtolejeunea A.Evans ........................................................... 316 Cystolejeunea A.Evans............................................................ 316 Omphalanthus Lindenb. et Nees ........................................... 316 subtrib. Cololejeuneinae Gradst. ................................................. 317 Aphanotropis Herzog ............................................................. 317 Calatholejeunea K.I.Goebel .................................................. 317 Cololejeunea (Spruce) Steph. ................................................ 317 Colura (Dumort.) Dumort. ................................................... 340 Diplasiolejeunea (Spruce) Schifn. ........................................ 344 Haplolejeunea Grolle ............................................................. 348 Macrocolura R.M.Schust. ...................................................... 348 Myriocoleopsis Schifn. .......................................................... 348 Nephelolejeunea Grolle .......................................................... 349 Schusterolejeunea Grolle ....................................................... 349 Siphonolejeunea Herzog ........................................................ 350 Tuyamaella S.Hatt................................................................. 350 subtrib. Cyclolejeuneinae Gradst. ................................................ 350 Bromeliophila R.M.Schust. ................................................... 350 Cyclolejeunea A.Evans ........................................................... 351 Prionolejeunea (Spruce) Schifn............................................ 351 subtrib. Drepanolejeuneinae Gradst. ........................................... 353 Capillolejeunea S.W.Arnell ................................................... 353 Drepanolejeunea (Spruce) Steph........................................... 353 Vitalianthus R.M.Schust. et Giancotti .................................. 361 subtrib. Echinolejeuneinae Gradst. .............................................. 361 Anoplolejeunea (Spruce) Schifn. .......................................... 361 Echinolejeunea R.M.Schust. .................................................. 362 Kymatolejeunea Grolle .......................................................... 362 World checklist of hornworts and liverworts 23 subtrib. Leiolejeuneinae Schäf.-Verw. et Heinrichs ...................... 362 Leiolejeunea A.Evans ............................................................. 362 subtrib. Lejeuneinae Gradst. ........................................................ 362 Harpalejeunea (Spruce) Schifn. ........................................... 362 Hattoriolejeunea Mizut. ........................................................ 364 Lejeunea Lib. ......................................................................... 364 Microlejeunea (Spruce) Steph. .............................................. 385 Taxilejeunea (Spruce) Steph. ................................................ 388 subtrib. Lepidolejeuneinae Gradst. .............................................. 391 Lepidolejeunea R.M.Schust. .................................................. 391 Otolejeunea Grolle et Tixier .................................................. 392 Rectolejeunea A.Evans ........................................................... 393 subtrib. Leptolejeuneinae Heinrichs et Schäf.-Verw. .................... 394 Leptolejeunea (Spruce) Steph. ............................................... 394 subtrib. Pycnolejeuneinae Heinrichs et Schäf.-Verw. .................... 396 Pycnolejeunea (Spruce) Schiffn. ........................................... 396 subtrib. Xylolejeuneinae Henrichs et Schäf.-Verw. ....................... 398 Xylolejeunea Xiao L.He et Grolle .......................................... 398 trib. Symbiezidieae Gradst. .......................................................... 398 Symbiezidium Trevis. ............................................................ 398 Ptychanthoideae Mizut. ............................................................... 399 Acrolejeunea (Spruce) Schifn. .............................................. 399 Archilejeunea (Spruce) Steph. ............................................... 401 Bryopteris (Nees) Lindenb..................................................... 403 Caudalejeunea Schifn........................................................... 403 Cephalantholejeunea R.M.Schust. ........................................ 405 Cephalolejeunea Mizut. ......................................................... 405 Frullanoides Raddi ................................................................ 405 Fulfordianthus Gradst. .......................................................... 405 Lopholejeunea (Spruce) Steph. .............................................. 406 Marchesinia Gray .................................................................. 408 Mastigolejeunea (Spruce) Steph. ........................................... 409 Phaeolejeunea Mizut. ............................................................ 410 Ptychanthus Nees ................................................................... 411 Schifneriolejeunea Verd........................................................ 411 Spruceanthus Verd. ................................................................ 412 hysananthus Lindenb. ......................................................... 413 Tuzibeanthus S.Hatt.............................................................. 414 Verdoornianthus Gradst. ....................................................... 414 Porellineae R.M.Schust. ....................................................................... 415 Goebeliellaceae Verd........................................................................ 415 Goebeliella Steph. .................................................................. 415 Lepidolaenaceae Nakai .................................................................... 415 24 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Gackstroemia Trevis. ............................................................. 415 Lepidogyna R.M.Schust. ........................................................ 416 Lepidolaena Dumort. ............................................................ 416 Porellaceae Cavers nom. conserv........................................................ 417 Ascidiota C.Massal. ................................................................ 417 Porella L................................................................................. 417 Radulineae R.M.Schust. ...................................................................... 424 Radulaceae Müll.Frib. ..................................................................... 424 Radula Dumort. .................................................................... 425 Ptilidiales Schljakov ................................................................................. 434 Herzogianthaceae Stotler et Crand.-Stotl. ........................................ 434 Herzogianthus R.M.Schust. ................................................... 434 Neotrichocoleaceae Inoue ................................................................ 434 Neotrichocolea S.Hatt. ........................................................... 434 Trichocoleopsis S.Okamura ................................................... 434 Ptilidiaceae H.Klinggr. .................................................................... 434 Ptilidium Nees ....................................................................... 434 Metzgeriidae Barthol.-Began ........................................................................ 435 Metzgeriales Chalaud ............................................................................... 435 Aneuraceae H.Klinggr. .................................................................... 435 Aneura Dumort. .................................................................... 435 Lobatiriccardia (Mizut. et S.Hatt.) Furuki........................... 437 Riccardia Gray ....................................................................... 438 Verdoornia R.M.Schust. ........................................................ 456 Metzgeriaceae H.Klinggr. ................................................................ 456 Metzgeria Raddi..................................................................... 457 Steereella Kuwah. .................................................................. 461 Vandiemenia Hewson ........................................................... 461 Pleuroziales Schljakov .............................................................................. 462 Pleuroziaceae Müll.Frib. .................................................................. 462 Pleurozia Dumort. ................................................................ 462 Pelliidae He-Nygrén .................................................................................... 463 Fossombroniales Schljakov ....................................................................... 463 Calyculariineae He-Nygrén.................................................................. 463 Calyculariaceae He-Nygrén ............................................................. 463 Calycularia Mitt. ................................................................... 463 Fossombroniineae R.M.Schust. ex Stotler et Crand.-Stotl. ................... 463 Allisoniaceae Schljakov .................................................................... 463 Allisonia Herzog .................................................................... 463 Fossombroniaceae Hazsl. nom. conserv. ............................................ 463 Fossombronia Raddi .............................................................. 463 Petalophyllaceae Stotler et Crand.-Stotl. .......................................... 468 Petalophyllum Nees et Gottsche ............................................ 468 World checklist of hornworts and liverworts 25 Sewardiella Kashyap ............................................................. 468 Makinoiineae He-Nygrén .................................................................... 468 Makinoaceae Nakai ......................................................................... 468 Makinoa Miyake.................................................................... 468 Pallaviciniales W.Frey et M.Stech ............................................................ 468 Pallaviciniineae R.M.Schust. ................................................................ 468 Hymenophytaceae R.M.Schust. ....................................................... 468 Hymenophyton Dumort......................................................... 468 Moerckiaceae K.I.Goebel ex Stotler et Crand.-Stotl. ........................ 469 Hattorianthus R.M.Schust. et Inoue..................................... 469 Moerckia Gottsche ................................................................. 469 Pallaviciniaceae Mig......................................................................... 469 Pallavicinioideae Mig. ex Grolle .................................................. 469 Jensenia Lindb. ...................................................................... 469 Pallavicinia Gray ................................................................... 470 Podomitrium Mitt. ................................................................ 471 Symphyogynoideae R.M.Schust. ex Grolle .................................. 471 Greeneothallus Hässel............................................................ 471 Seppeltia Grolle ..................................................................... 471 Symphyogyna Nees et Mont. .................................................. 471 Symphyogynopsis Grolle......................................................... 473 Xenothallus R.M.Schust. ....................................................... 473 Sandeothallaceae R.M.Schust. ......................................................... 474 Sandeothallus R.M.Schust..................................................... 474 Phyllothalliineae R.M.Schust. .............................................................. 474 Phyllothalliaceae E.A.Hodgs. ex T.Katag. ........................................ 474 Phyllothallia E.A.Hodgs. ....................................................... 474 Pelliales He-Nygrén ................................................................................. 474 Noterocladaceae W.Frey et M.Stech ................................................ 474 Noteroclada Taylor ex Hook.f. et Wilson ............................. 474 Pelliaceae H.Klinggr. ....................................................................... 475 Pellia Raddi ........................................................................... 475 Marchantiopsida Cronquist.............................................................................. 476 Blasiidae He-Nygrén .................................................................................... 476 Blasiales Stotler et Crand.-Stotl. ............................................................... 476 Blasiaceae H.Klinggr........................................................................ 476 Blasia L. ................................................................................. 476 Cavicularia Steph. ................................................................. 476 Marchantiidae Engl...................................................................................... 476 Lunulariales H.Klinggr. ........................................................................... 476 Lunulariaceae H.Klinggr.................................................................. 476 Lunularia Adans. ................................................................... 476 Marchantiales Limpr. ............................................................................... 476 26 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Aytoniaceae Cavers .......................................................................... 476 Asterella P.Beauv. .................................................................. 477 Cryptomitrium Austin ex Underw. ....................................... 481 Mannia Corda ....................................................................... 481 Plagiochasma Lehm. .............................................................. 482 Reboulia Raddi ...................................................................... 483 Cleveaceae Cavers ............................................................................ 484 Athalamia Falc. ...................................................................... 484 Clevea Lindb. ......................................................................... 484 Peltolepis Lindb. .................................................................... 484 Sauteria Nees ......................................................................... 485 Conocephalaceae Müll.Frib. ex Grolle ............................................. 485 Conocephalum Hill ................................................................ 485 Corsiniaceae Engl. ........................................................................... 486 Corsinioideae Schifn....................................................................... 486 Corsinia Raddi ....................................................................... 486 Cronisioideae R.M.Schust. .............................................................. 486 Cronisia Berk. ........................................................................ 486 Cyathodiaceae Stotler et Crand.-Stotl. ............................................. 486 Cyathodium Kunze ................................................................ 486 Dumortieraceae D.G.Long .............................................................. 487 Dumortiera Nees ................................................................... 487 Exormothecaceae Müll.Frib. ex Grolle ............................................. 487 Aitchisoniella Kashyap........................................................... 487 Exormotheca Mitt. ................................................................. 488 Stephensoniella Kashyap ....................................................... 488 Marchantiaceae Lindl. ..................................................................... 488 Bucegioideae R.M.Schust. ........................................................... 488 Bucegia Radian ...................................................................... 488 Marchantioideae Schifn. ............................................................. 489 Marchantia L. ........................................................................ 489 Preissia Corda ........................................................................ 492 Monocleaceae A.B.Frank ................................................................. 492 Monoclea Hook. .................................................................... 492 Monosoleniaceae Inoue ................................................................... 492 Monosolenium Grif. .............................................................. 492 Oxymitraceae Müll.Frib. ex Grolle .................................................. 493 Oxymitra Bisch. ex Lindenb.................................................. 493 Ricciaceae Rchb. .............................................................................. 493 Riccia L. ................................................................................. 493 Ricciocarpos Corda ................................................................ 503 Targioniaceae Dumort..................................................................... 503 World checklist of hornworts and liverworts 27 Targionia L. ........................................................................... 503 Wiesnerellaceae Inoue...................................................................... 504 Wiesnerella Schifn. ............................................................... 504 Neohodgsoniales D.G.Long ..................................................................... 504 Neohodgsoniaceae D.G.Long .......................................................... 504 Neohodgsonia Perss................................................................ 504 Sphaerocarpales Cavers ............................................................................ 504 Monocarpaceae D.J.Carr ex Schelpe ................................................ 504 Monocarpus D.J.Carr ............................................................ 504 Riellaceae Engl................................................................................. 504 Austroriella Cargill et J.Milne ............................................... 504 Riella Mont. ........................................................................... 505 Sphaerocarpaceae Heeg.................................................................... 506 Geothallus Campb. ................................................................ 506 Sphaerocarpos Boehm. .......................................................... 506 Taxonomic list ANTHOCEROTOPHYTA ANTHOCEROTOPSIDA de Bary ex Jancz. Anthocerotidae Rosenv. Anthocerotales Limpr. *** Anthocerotaceae Dumort. by J.C. Villarreal and D.C. Cargill Notes on nomenclature and taxonomy can also be found in Cargill et al. (2013b) and Villarreal et al. (2015) *** Anthoceros L., Sp. Pl. 1: 1139, 1753 (Linnaeus 1753). 1 *** Anthoceros adscendens Lehm. et Lindenb., Nov. Stirp. Pug. 4: 24, 1832 (Lehmann 1832). *** Anthoceros agrestis Paton, J. Bryol. 10 (3): 257, 1979 (Paton 1979a), nom. conserv. 2 *** Anthoceros alpinus Steph., Sp. Hepat. (Stephani) 6: 425, 1923 (Stephani 1923). 1 2 Anthoceros includes Aspiromitus, but some taxa have neither been transferred nor synonymized. hey are listed in the “Names in genera not currently accepted” section below. Anthoceros agrestis is conserved against the earlier Anthoceros nagasakiensis Steph. 1916. Anthoceros cristatus Steph. 1916 is also an earlier synonym and a proposal to conserve Anthoceros agrestis against it is in progress. 28 * *** *** ** *** *** ** * ** ** ** *** ** * *** *** *** ** ** ** * ** 3 4 5 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Anthoceros angustifolius Gottsche, Lindenb. et Nees, Syn. Hepat. 4: 582, 1846 (Gottsche et al. 1846). 3 Anthoceros angustus Steph., Sp. Hepat. (Stephani) 5: 1001, 1916 (Stephani 1916b). Anthoceros bharadwajii Udar et A.K.Asthana, Proc. Indian Natl. Sci. Acad., B 51 (4): 484, 1985 (Udar and Asthana 1985b). Anthoceros buettneri Steph., Sp. Hepat. (Stephani) 5: 997, 1916 (Stephani 1916b). Anthoceros capricornii Cargill et G.A.M.Scott, J. Hattori Bot. Lab. 82: 55, 1997 (Cargill and Scott 1997). Anthoceros caucasicus Steph., Izv. Kavkazsk. Muz. 8: 87, 1914 (Voronov’ 1914). Anthoceros cavernosus Steph., Sp. Hepat. (Stephani) 5: 998, 1916 (Stephani 1916b). Anthoceros chambensis Kashyap, J. Bombay Nat. Hist. Soc. 25 (2): 281, 1917 (Kashyap 1917). Anthoceros chungii Khanna, J. Indian Bot. Soc. 17 (5/6): 316, 1938 (Khanna 1938). Anthoceros crispatus Grif., Not. pl. asiat. 2: 349, 1849 (Griith 1849). Anthoceros dimorphus Sim, Trans. Roy. Soc. South Africa 15 (1): 114, 1926 (Sim 1926). Anthoceros erectus Kashyap, New Phytol. 14 (1): 9, 1915 (Kashyap 1915). Anthoceros expansus (Steph.) J.C.Villarreal et Cargill, Phytotaxa 208 (1): 92, 2015 (Villarreal et al. 2015). Bas.: Aspiromitus expansus Steph., Sp. Hepat. (Stephani) 5: 961, 1916 (Stephani 1916b). Anthoceros ferdinandi-muelleri Steph., Sp. Hepat. (Stephani) 5: 1007, 1916 (Stephani 1916b). 4 Anthoceros fragilis Steph., Sp. Hepat. (Stephani) 5: 1006, 1916 (Stephani 1916b). Anthoceros fusiformis Austin, Bull. Torrey Bot. Club 6 (4): 28, 1875 [1876] (Austin 1875a). Anthoceros fusiformis var. taiwanensis J.Haseg., Acta Phytotax. Geobot. 44 (2): 100, 1993 (Hasegawa 1993b). Anthoceros gasongorii Gola, Mem. Reale Accad. Sci. Torino (ser. 2) 65 (1): 10, 1916 (Gola 1916). Anthoceros granulatus Gottsche, Mexik. Leverm.: 275, 1863 (Gottsche 1863). Anthoceros harrisanus (Steph.) Parihar, Univ. Allahabad Stud., Bot. 1961-2: 31, 1962 (Parihar 1962). Bas.: Aspiromitus harrisanus Steph., Sp. Hepat. (Stephani) 5: 965, 1916 (Stephani 1916b). Anthoceros helmsii Steph., Hedwigia 32 (3): 142, 1893 (Stephani 1893b). 5 Anthoceros jamesonii Taylor ex Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 171, 1855 (Mitten 1855). Anthoceros angustifolius (type from Europe) has neither been recognized in any recent treatment nor synonymized. he name may have priority once its identity is determined. Anthoceros ferdinandi-muelleri is similar to Anthoceros punctatus, but Anthoceros still needs to be revised for Australia. Anthoceros helmsii was treated as conspeciic with Anthoceros muscoides by Campbell (1982), but it is in need of a revision. World checklist of hornworts and liverworts * 29 *** Anthoceros javanicoides H.A.Mill., Phytologia 47 (4): 319, 1981 (Miller 1981). Nom. nov. pro Anthoceros javanicus Steph., Sp. Hepat. (Stephani) 5: 988, 1916 (Stephani 1916b), nom. illeg. 6 Anthoceros jungermannioides Schwein., Spec. Fl. Amer. Crypt.: 25, 1821 (Schweinitz 1821). Anthoceros kajumas (K.I.Goebel) Prosk., Bull. Torrey Bot. Club 78 (4): 347, 1951 (Proskauer 1951a). Bas.: Aspiromitus kajumas K.I.Goebel, Ann. Jard. Bot. Buitenzorg 39: 69, 1928 (Goebel 1928). Anthoceros koshii Khanna, J. Indian Bot. Soc. 15 (4): 237, 1936 (Khanna 1936). Anthoceros lamellatus Steph., Sp. Hepat. (Stephani) 5: 1000, 1916 (Stephani 1916b). Anthoceros laminifer Steph., J. Linn. Soc., Bot. 29 (201): 266, 1892 (Stephani 1892b). Anthoceros macounii M.Howe, Bull. Torrey Bot. Club 25 (1): 19, 1898 (Howe 1898a). Anthoceros macrosporus Steph., Sp. Hepat. (Stephani) 5: 1005, 1916 (Stephani 1916b). Anthoceros maritimus Steph., Sp. Hepat. (Stephani) 5: 984, 1916 (Stephani 1916b). Anthoceros megasporus Meijer, J. Hattori Bot. Lab. 18: 6, 1957 (Meijer 1957). Anthoceros muscoides Colenso, Trans. & Proc. New Zealand Inst. 16: 361, 1884 (Colenso 1884). Anthoceros myriandroecius Steph., Wiss. Ergebn. Deut. Zentr.-Afr. Exped. (190708), Bot. 2: 134, 1911 (Stephani 1911a). Anthoceros natalensis Steph., Akad. Wiss. Wien, Math.-Naturwiss. Kl., Denkschr. 88: 732, 1913 (Stephani 1913b). Anthoceros neesii Prosk., Leberm. Eur. 2 (9): 1312, 1958 (Proskauer 1958). Nom. nov. pro Anthoceros punctatus α* monocarpus Nees, Naturgesch. Eur. Leberm. 4: 339, 1838 (Nees 1838a). Anthoceros niger Steph., Sp. Hepat. (Stephani) 5: 1005, 1916 (Stephani 1916b). Anthoceros orizabensis (Steph.) Hässel, Candollea 45 (1): 211, 1990 (Hässel 1990b). Bas.: Aspiromitus orizabensis Steph., Sp. Hepat. (Stephani) 5: 965, 1916 (Stephani 1916b). Anthoceros pandei Udar et A.K.Asthana, J. Indian Bot. Soc. 64: 305, 1985 (Udar and Asthana 1985a). Anthoceros patagonicus Hässel, Candollea 45 (1): 207, 1990 (Hässel 1990b). Anthoceros patagonicus subsp. gremmenii J.C.Villarreal, J.J.Engel et Váňa, Mem. New York Bot. Gard. 105: 32, 2013 (Váňa and Engel 2013). Anthoceros peruvianus Steph., Sp. Hepat. (Stephani) 5: 999, 1916 (Stephani 1916b). Anthoceros pinnatus Steph., Bol. Soc. Brot. 4: 154 [182], 1885 [1886] (Stephani 1885c). Anthoceros punctatus L., Sp. Pl. 1: 1139, 1753 (Linnaeus 1753). 7 6 7 Anthoceros javanicoides belongs to the Anthoceros punctatus species complex (Söderström et al. 2010a). Anthoceros punctatus is a species complex in need of a revision. ** * * *** *** *** *** ** * ** *** ** *** ** ** *** *** *** ** ** 30 * *** *** *** ** ** ** ** *** *** *** *** ** * * * * * * * 8 9 10 11 12 13 14 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Anthoceros pusillus Colenso, Trans. & Proc. New Zealand Inst. 18: 255, 1886 (Colenso 1886b). Anthoceros rosulans J.Haseg., J. Hattori Bot. Lab. 60: 379, 1986 (Hasegawa 1986b). Anthoceros sambesianus Steph., Sp. Hepat. (Stephani) 5: 996, 1916 (Stephani 1916b). Anthoceros scariosus Austin, Proc. Acad. Nat. Sci. Philadelphia 21: 230, 1869 (Austin 1869). Anthoceros schroederi Steph., 52 (5): 306, 1912 (Stephani 1912a). Anthoceros serratus Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 90, 1911 (Stephani 1911b). Anthoceros simulans M.Howe, Proc. Calif. Acad. Sci. (ser. 4) 21 (17): 204, 1934 (Howe 1934). Anthoceros spongiosus Steph., Sp. Hepat. (Stephani) 5: 1003, 1916 (Stephani 1916b). Anthoceros subtilis Steph., Sp. Hepat. (Stephani) 5: 1003, 1916 (Stephani 1916b). Anthoceros telaganus Steph., Sp. Hepat. (Stephani) 5: 1005, 1916 (Stephani 1916b). Anthoceros tristanianus J.C.Villarreal, J.J.Engel et Váňa, Mem. New York Bot. Gard. 105: 33, 2013 (Váňa and Engel 2013). Anthoceros tuberculatus Lehm. et Lindenb., Nov. Stirp. Pug. 4: 25, 1832 (Lehmann 1832). Anthoceros venosus Lindenb. et Gottsche, Syn. Hepat. 4: 584, 1846 (Gottsche et al. 1846). Excluded from the genus Anthoceros aethyopicus Gola, Ann. Bot. (Rome) 13 (1): 73, 1914 (Gola 1914a). 8 Anthoceros brunnthaleri Steph., Akad. Wiss. Wien, Math.-Naturwiss. Kl., Denkschr. 88: 732, 1913 (Stephani 1913b). 9 Anthoceros loribundus Steph., Sp. Hepat. (Stephani) 5: 977, 1916 (Stephani 1916b). 10 Anthoceros mildbraedii Steph., Sp. Hepat. (Stephani) 6: 428, 1923 (Stephani 1923). 11 Anthoceros parvifrons Steph., 52 (5): 307, 1912 (Stephani 1912a). 12 Anthoceros pseudocostus Steph., Sp. Hepat. (Stephani) 5: 977, 1916 (Stephani 1916b). 13 Anthoceros rossoi Gola, Mem. Reale Accad. Sci. Torino (ser. 2) 65 (1): 10, 1916 (Gola 1916). 14 Anthoceros aethyopicus is probably a Phaeoceros species (Wigginton 2009). Anthoceros brunnthaleri was annotated by Hasegawa in 1993 as probably conspeciic with Phymatoceros bulbiculosus but the synonymy seems not to have been published. Anthoceros loribundus type specimen in G was annotated by Hasegawa in 1993 as conspeciic with Anthoceros parvifrons which is a Phaeoceros species. Anthoceros mildbraedii is probably conspeciic with Phaeoceros carolinianus (Wigginton and Grolle 1996). Anthoceros parvifrons is annotated as a Phaeoceros species by Hasegawa in 1993 but the new combination has never been published. Anthoceros pseudocostus is a Phaeoceros species of uncertain status (Grolle 1995). Anthoceros rossoi is a Phaeoceros species of uncertain status (Wigginton 2009). World checklist of hornworts and liverworts 31 *** Folioceros D.C.Bharadwaj, Geophytology 1 (1): 9, 1971 (Bharadwaj 1971). *** Folioceros amboinensis (Schifn.) Piippo, Acta Bot. Fenn. 148: 36, 1993 (Piippo 1993b). Bas.: Anthoceros amboinensis Schifn., Leberm., Forschungsr. Gazelle 4 (4): 45, 1890 (Schifner 1890). ** Folioceros apiahynus (Steph.) Hässel, Candollea 45 (1): 215, 1990 (Hässel 1990b). Bas.: Anthoceros apiahynus Steph., Sp. Hepat. (Stephani) 5: 999, 1916 (Stephani 1916b). ** Folioceros argillaceus (Steph.) J.C.Villarreal et Cargill, Phytotaxa 208 (1): 93, 2015 (Villarreal et al. 2015). Bas.: Aspiromitus argillaceus Steph., Sp. Hepat. (Stephani) 5: 970, 1916 (Stephani 1916b). *** Folioceros assamicus D.C.Bharadwaj, Geophytology 1 (1): 10, 1971 (Bharadwaj 1971). ** Folioceros dilatatus (Steph.) J.C.Villarreal et Cargill, Phytotaxa 208 (1): 93, 2015 (Villarreal et al. 2015). Bas.: Anthoceros dilatatus Steph., Bot. Jahrb. Syst. 8 (2): 95, 1886 (Stephani 1886d). *** Folioceros dixitianus (Mahab.) D.C.Bharadwaj, Geophytology 5 (2): 227, 1975 (Bharadwaj 1975). Bas.: Aspiromitus dixitianus Mahab., Curr. Sci. 10 (12): 532, 1941 (Mahabale 1941). *** Folioceros fuciformis (Mont.) D.C.Bharadwaj, Geophytology 5 (2): 227, 1975 (Bharadwaj 1975). Bas.: Anthoceros fuciformis Mont., Ann. Sci. Nat. Bot. (sér. 2) 20: 296, 1844 (Montagne 1844b). *** Folioceros glandulosus (Lehm. et Lindenb.) D.C.Bharadwaj, Geophytology 5 (2): 227, 1975 (Bharadwaj 1975). Bas.: Anthoceros glandulosus Lehm. et Lindenb., Nov. Stirp. Pug. 4: 26, 1832 (Lehmann 1832). *** Folioceros incurvus (Steph.) D.C.Bharadwaj, Geophytology 5 (2): 227, 1975 (Bharadwaj 1975). Bas.: Anthoceros incurvus Steph., Hedwigia 32 (3): 143, 1893 (Stephani 1893b). *** Folioceros indicus D.C.Bharadwaj, Geophytology 8 (1): 114, 1978 (Bharadwaj 1978). *** Folioceros kashyapii S.C.Srivast. et A.K.Asthana, Bryologist 92 (2): 219, 1989 (Srivastava and Asthana 1989). *** Folioceros mangaloreus (Steph.) D.C.Bharadwaj, Geophytology 5 (2): 227, 1975 (Bharadwaj 1975). Bas.: Aspiromitus mangaloreus Steph., Sp. Hepat. (Stephani) 5: 967, 1916 (Stephani 1916b). *** Folioceros paliformis D.K.Singh, Bull. Bot. Surv. India 29: 176, 1987 [1989] (Singh 1987b). *** Folioceros physocladus D.C.Bharadwaj, Geophytology 8 (1): 115, 1978 (Bharadwaj 1978). Based on: Anthoceros physocladus Schifn. ex Pandé, Proc. Indian Sci. Congr. Assoc. 47 (2): 98, 1960 (Pandé 1960), nom. inval. *** Folioceros pinnilobus (Steph.) D.C.Bharadwaj, Geophytology 5 (2): 227, 1975 (Bharadwaj 1975). Bas.: Anthoceros pinnilobus Steph., Akad. Wiss. Wien, Math.Naturwiss. Kl., Denkschr. 81: 299, 1907 (Stephani 1907a). *** Folioceros satpurensis D.C.Bharadwaj et K.P.Srivast., Geophytology 8 (1): 112, 1978 (Bharadwaj 1978). Based on: Anthoceros satpurensis K.P.Srivast., Proc. Indian Sci. Congr. Assoc. 47 (3): 337, 1960 (Srivastava 1960), nom. inval. 32 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Folioceros udarii A.K.Asthana et S.C.Srivast., Cryptog. Bryol. Lichénol. 7 (2): 151, 1986 (Asthana and Srivastava 1986). *** Folioceros verruculosus (J.Haseg.) R.L.Zhu et M.J.Lai, Ann. Bot. Fenn. 48 (5): 383, 2011 (Wang et al. 2011). Bas.: Anthoceros verruculosus J.Haseg., Acta Phytotax. Geobot. 44 (2): 103, 1993 (Hasegawa 1993b). Dendrocerotidae R.J.Duff, J.C.Villarreal, Cargill et Renzaglia Dendrocerotales Hässel *** Dendrocerotaceae J.Haseg. by J.C. Villarreal and D.C. Cargill *** Dendroceros Nees, Syn. Hepat. 4: 579, 1846 (Gottsche et al. 1846). ** subg. Apoceros R.M.Schust., Phytologia 63 (3): 200, 1987 (Schuster 1987b). *** Dendroceros cavernosus J.Haseg., J. Hattori Bot. Lab. 47: 306, 1980 (Hasegawa 1980). *** Dendroceros diicilis Steph., Sp. Hepat. (Stephani) 5: 1009, 1917 (Stephani 1917b). *** Dendroceros muelleri Steph., Hedwigia 28 (2): 133, 1889 (Stephani 1889a). *** Dendroceros ogeramnangus Piippo, Acta Bot. Fenn. 148: 40, 1993 (Piippo 1993b). *** Dendroceros pedunculatus Steph., Sitzungsber. Naturf. Ges. Leipzig 36: 14, 1909 (Stephani 1909c). ** Dendroceros subdiicilis S.Hatt., Bot. Mag. (Tokyo) 64 (755/756): 119, 1951 (Hattori 1951c). ** subg. Dendroceros *** Dendroceros acutilobus Steph., Sitzungsber. Naturf. Ges. Leipzig 36: 18, 1909 (Stephani 1909c). *** Dendroceros borbonicus Steph., Bull. Soc. Roy. Bot. Belgique, Compt. Rend. 32 (2): 31, 1893 [1894] (Stephani 1893e). *** Dendroceros crispus (Sw.) Nees, Syn. Hepat. 4: 581, 1846 (Gottsche et al. 1846). Bas.: Anthoceros crispus Sw., Prodr. (Swartz): 146, 1788 (Swartz 1788). *** Dendroceros foliicola J.Haseg., J. Hattori Bot. Lab. 47: 296, 1980 (Hasegawa 1980). *** Dendroceros japonicus Steph., Sitzungsber. Naturf. Ges. Leipzig 36: 15, 1909 (Stephani 1909c). *** Dendroceros javanicus (Nees) Nees, Syn. Hepat. 4: 582, 1846 (Gottsche et al. 1846). Bas.: Anthoceros javanicus Nees, Enum. Pl. Crypt. Javae: 1, 1830 (Nees 1830). *** Dendroceros subplanus Steph., Sitzungsber. Naturf. Ges. Leipzig 36: 20, 1909 (Stephani 1909c). *** Dendroceros tubercularis S.Hatt., Bot. Mag. (Tokyo) 58 (685): 6, 1944 (Hattori 1944b). World checklist of hornworts and liverworts 33 *** Dendroceros validus Steph., Sp. Hepat. (Stephani) 5: 1016, 1917 (Stephani 1917b). ** Dendroceros vesconianus Gottsche, J. Bot. (Morot) 12: 150, 1898 (Bescherelle 1898). ** Dendroceros wattsianus Steph., Sitzungsber. Naturf. Ges. Leipzig 36: 17, 1909 (Stephani 1909c). * *** ** ** ** ** *** * ** *** ** ** * * *** ** ** ** Incertae sedis Dendroceros adglutinatus (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 4: 580, 1846 (Gottsche et al. 1846). Bas.: Monoclea adglutinata Hook.f. et Taylor, London J. Bot. 4: 96, 1845 (Hooker and Taylor 1845). 15 Dendroceros africanus Steph., Sitzungsber. Naturf. Ges. Leipzig 36: 18, 1909 (Stephani 1909c). Dendroceros allionii Steph., Sp. Hepat. (Stephani) 5: 1014, 1917 (Stephani 1917b). Dendroceros australis Steph., Sitzungsber. Naturf. Ges. Leipzig 36: 17, 1909 (Stephani 1909c). Dendroceros breutelii Nees, Syn. Hepat. 4: 581, 1846 (Gottsche et al. 1846). Dendroceros breutelii var. surinamensis Lindenb. et Gottsche, Linnaea 24 (6): 639, 1851 [1852] (Lindenberg and Gottsche 1851a). Dendroceros cichoraceus (Mont.) Gottsche, Bot. Zeitung (Berlin) Beil. 16: 16, 1858 (Gottsche 1858). Bas.: Anthoceros cichoraceus Mont., Ann. Sci. Nat. Bot. (sér. 3) 4: 355, 1845 (Montagne 1845b). Dendroceros crassicostatus Steph., Sp. Hepat. (Stephani) 5: 1015, 1917 (Stephani 1917b). Dendroceros crassinervis (Nees) Gottsche, Bot. Zeitung (Berlin) Beil. 16: 18, 1858 (Gottsche 1858). Bas.: Anthoceros crassinervis Nees, Syn. Hepat. 4: 589, 1846 (Gottsche et al. 1846). Dendroceros crispatus (Hook.) Nees, Syn. Hepat. 4: 579, 1846 (Gottsche et al. 1846). Bas.: Monoclea crispata Hook., Bot. Misc. 1: 117, 1830 (Hooker 1830). Dendroceros crispatus var. simplicior Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 574, 1885 (Spruce 1885). Dendroceros cucullatus Steph., Sp. Hepat. (Stephani) 6: 429, 1923 (Stephani 1923). 16 Dendroceros exalatus Steph., Sitzungsber. Naturf. Ges. Leipzig 36: 14, 1909 (Stephani 1909c). 17 Dendroceros gracilis Steph., Sp. Hepat. (Stephani) 5: 1015, 1917 (Stephani 1917b). Dendroceros granulatus Mitt., Fl. vit.: 419, 1871 [1873] (Mitten 1871). Dendroceros herasii M.Infante, J. Bryol. 32 (4): 285, 2010 (Infante 2010). Dendroceros humboldtensis Hürl., Bauhinia 1 (3): 254, 1960 (Hürlimann 1960). Dendroceros paivae C.A.Garcia, Sérgio et J.C.Villarreal, Cryptog. Bryol. 33 (1): 5, 2012 (Garcia et al. 2012). 15 Dendroceros adglutinatus is conspeciic with Dendroceros crispus in Proskauer (1960), but later authors accept it. 16 Dendroceros cucullatus was treated as conspeciic with Dendroceros diicilis by Hasegawa (1980), but re-instated by Chantanaorrapint et al. (2014). 17 Dendroceros exalatus was reported from the Moluccas in Stephani (1909c), but Hasegawa (1980) states that this is a Brazilian species. Dendroceros has not been recently revised for Brazil. 34 * * ** *** ** ** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Dendroceros rarus Steph., Sp. Hepat. (Stephani) 5: 1014, 1917 (Stephani 1917b). Dendroceros reticulus Herzog, Memoranda Soc. Fauna Fl. Fennica 26: 37, 1950 [1951] (Herzog 1950b). 18 Dendroceros rigidus Steph., Sp. Hepat. (Stephani) 5: 1017, 1917 (Stephani 1917b). Dendroceros seramensis J.Haseg., Acta Phytotax. Geobot. 37 (1/3): 10, 1986 (Hasegawa 1986a). Dendroceros subtropicus C.J.Wild, Trans. Nat. Hist. Soc. Queensland 1: 49, 1893 (Wild 1893). Dendroceros tahitensis Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 30 (5): 138, 1873 (Ångström 1873). *** Megaceros Campb., Ann. Bot. (Oxford) 21 (4): 484, 1907 (Campbell 1907). ** Megaceros aneuriformis Steph., Sp. Hepat. (Stephani) 5: 949, 1916 (Stephani 1916b). *** Megaceros austronesophilus Cargill et Seppelt, Austral. Syst. Bot. 26 (5): 372, 2013 (Cargill et al. 2013a). ** Megaceros ciliatus K.I.Goebel, Ann. Jard. Bot. Buitenzorg 39: 74, 1928 (Goebel 1928). *** Megaceros denticulatus (Lehm.) Steph., Sp. Hepat. (Stephani) 5: 956, 1916 (Stephani 1916b). Bas.: Anthoceros denticulatus Lehm., Nov. Stirp. Pug. 10: 25, 1857 (Lehmann 1857). *** Megaceros lagellaris (Mitt.) Steph., Sp. Hepat. (Stephani) 5: 951, 1916 (Stephani 1916b). Bas.: Anthoceros lagellaris Mitt., Fl. vit.: 419, 1871 [1873] (Mitten 1871). *** Megaceros gracilis (Reichardt) Steph., Sp. Hepat. (Stephani) 5: 955, 1916 (Stephani 1916b). Bas.: Anthoceros gracilis Reichardt, Verh. K.K. Zool.-Bot. Ges. Wien 16: 957, 1866 (Reichardt 1866). ** Megaceros leptohymenius (Hook.f. et Taylor) Steph., Sp. Hepat. (Stephani) 5: 955, 1916 (Stephani 1916b). Bas.: Monoclea leptohymenia Hook.f. et Taylor, London J. Bot. 3: 575, 1844 (Hooker and Taylor 1844d). *** Megaceros pellucidus (Colenso) E.A.Hodgs., J. Roy. Soc. New Zealand 2 (1): 115, 1972 (Hodgson 1972). Bas.: Anthoceros pellucidus Colenso, Trans. & Proc. New Zealand Inst. 17: 263, 1885 (Colenso 1885). ** Megaceros tjibodensis Campb., Ann. Bot. (Oxford) 21 (4): 484, 1907 (Campbell 1907). * * Excluded from the genus 19 Megaceros lavens (Spruce) Campb., Ann. Bot. (Oxford) 21 (4): 483, 1907 (Campbell 1907). Bas.: Anthoceros lavens Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 575, 1885 (Spruce 1885). Megaceros jamesonii (Taylor) Steph., Biblioth. Bot. 87 (2): 268, 1916 (Stephani 1916a). Bas.: Dendroceros jamesonii Taylor, London J. Bot. 7: 285, 1848 (Taylor 1848b). 18 Dendroceros reticulus was mentioned by Hasegawa (1980). He did not synonymise it with Dendroceros javanicus, he just stated that Herzog thought it similar to Dendroceros elegans which is conspeciic with Dendroceros javanicus. 19 Megaceros lavens and Megaceros jamesonii are probably conspeciic with Nothoceros vincentianus (Villarreal et al. 2010a). World checklist of hornworts and liverworts 35 *** Nothoceros (R.M.Schust.) J.Haseg., J. Hattori Bot. Lab. 76: 32, 1994 (Hasegawa 1994b). Bas.: Megaceros subg. Nothoceros R.M.Schust., Phytologia 63 (3): 200, 1987 (Schuster 1987b). *** Nothoceros aenigmaticus J.C.Villarreal et K.D.McFarland, Bryologist 113 (1): 109, 2010 (Villarreal et al. 2010b). Based on: Megaceros aenigmaticus R.M.Schust., Hepat. Anthocerotae N. Amer. 6: 830, 1992 (Schuster 1992d), nom. inval. *** Nothoceros canaliculatus (Pagán) J.C.Villarreal, Hässel et N.Salazar, Bryologist 110 (2): 283, 2007 (Villarreal et al. 2007). Bas.: Dendroceros canaliculatus Pagán, Bryologist 45 (4): 111, 1942 (Pagán 1942a). *** Nothoceros endiviifolius (Mont.) J.Haseg. ex J.C.Villarreal, Hässel et N.Salazar, Bryologist 110 (2): 283, 2007 (Villarreal et al. 2007). Bas.: Anthoceros endiviifolius Mont., Voy. Pole Sud, Bot. 1: 211, 1845 (Montagne 1845c). ** Nothoceros fuegiensis (Steph.) J.C.Villarreal, Bryologist 113 (1): 109, 2010 (Villarreal et al. 2010b). Bas.: Megaceros fuegiensis Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 91, 1911 (Stephani 1911b). *** Nothoceros giganteus (Lehm. et Lindenb.) J.Haseg. ex J.C.Villarreal, Hässel et N.Salazar, Bryologist 110 (2): 283, 2007 (Villarreal et al. 2007). Bas.: Anthoceros giganteus Lehm. et Lindenb., Nov. Stirp. Pug. 4: 25, 1832 (Lehmann 1832). *** Nothoceros minarum (Nees) J.C.Villarreal, Molec. Phylogen. Evol. 78: 34, 2014 (Villarreal and Renner 2014). Bas.: Anthoceros minarum Nees, Naturgesch. Eur. Leberm. 4: 340, 1838 (Nees 1838a). ** Nothoceros renzagliensis J.C.Villarreal, L.V.Campos et Uribe, Syst. Bot. 37 (1): 32, 2012 (Villarreal et al. 2012). *** Nothoceros schizophyllus (Steph.) J.C.Villarreal, Molec. Phylogen. Evol. 78: 34, 2014 (Villarreal and Renner 2014). Bas.: Megaceros schizophyllus Steph., Sp. Hepat. (Stephani) 5: 949, 1916 (Stephani 1916b). *** Nothoceros superbus J.C.Villarreal, Hässel et N.Salazar, Bryologist 110 (2): 280, 2007 (Villarreal et al. 2007). *** Nothoceros vincentianus (Lehm. et Lindenb.) J.C.Villarreal, Bryologist 113 (1): 111, 2010 (Villarreal et al. 2010b). Bas.: Anthoceros vincentianus Lehm. et Lindenb., Nov. Stirp. Pug. 6: 16, 1834 (Lehmann 1834). ** Phaeomegaceros R.J.Duf, J.C.Villarreal, Cargill et Renzaglia, Bryologist 110 (2): 241, 2007 (Duf et al. 2007). *** Phaeomegaceros coriaceus (Steph.) R.J.Duf, J.C.Villarreal, Cargill et Renzaglia, Bryologist 110 (2): 241, 2007 (Duf et al. 2007). Bas.: Anthoceros coriaceus Steph., Sp. Hepat. (Stephani) 5: 991, 1916 (Stephani 1916b). *** Phaeomegaceros imbriatus (Gottsche) R.J.Duf, J.C.Villarreal, Cargill et Renzaglia, Bryologist 110 (2): 241, 2007 (Duf et al. 2007). Bas.: Anthoceros imbriatus Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 187, 1864 (Gottsche 1864). *** Phaeomegaceros foveatus (J.Haseg.) J.C.Villarreal, Nova Hedwigia 91 (3/4): 352, 2010 (Villarreal et al. 2010a). Bas.: Phaeoceros foveatus J.Haseg., Bryol. Res. 7 (12): 374, 2001 (Hasegawa 2001). 36 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Phaeomegaceros hirticalyx (Steph.) R.J.Duf, J.C.Villarreal, Cargill et Renzaglia, Bryologist 110 (2): 241, 2007 (Duf et al. 2007). Bas.: Aspiromitus hirticalyx Steph., Sp. Hepat. (Stephani) 5: 966, 1916 (Stephani 1916b). *** Phaeomegaceros plicatus (Mitt.) J.C.Villarreal, J.J.Engel et Váňa, Mem. New York Bot. Gard. 105: 85, 2013 (Váňa and Engel 2013). Bas.: Anthoceros plicatus Mitt., Rep. Challenger, Bot. 1 (2): 178, 1884 (Mitten 1884a). *** Phaeomegaceros skottsbergii (Steph.) R.J.Duf, J.C.Villarreal, Cargill et Renzaglia, Bryologist 110 (2): 241, 2007 (Duf et al. 2007). Bas.: Anthoceros skottsbergii Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 90, 1911 (Stephani 1911b). *** Phaeomegaceros squamuliger (Spruce) J.C.Villarreal, Nova Hedwigia 91 (3/4): 351, 2010 (Villarreal et al. 2010a). Bas.: Anthoceros squamuliger Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 576, 1885 (Spruce 1885). ** Phaeomegaceros squamuliger subsp. hasselii J.C.Villarreal, Cargill et Goinet, Nova Hedwigia 91 (3/4): 352, 2010 (Villarreal et al. 2010a). Phymatocerotales R.J.Duff, J.C.Villarreal, Cargill et Renzaglia *** Phymatocerotaceae R.J.Duff, J.C.Villarreal, Cargill et Renzaglia by J.C. Villarreal and D.C. Cargill *** Phymatoceros Stotler, W.T.Doyle et Crand.-Stotl., Phytologia 87 (2): 113, 2005 (Stotler et al. 2005). *** Phymatoceros bulbiculosus (Brot.) Stotler, W.T.Doyle et Crand.-Stotl., Phytologia 87 (2): 114, 2005 (Stotler et al. 2005). Bas.: Anthoceros bulbiculosus Brot., Fl. lusit. 2: 430, 1804 [1805] (Brotero 1804). *** Phymatoceros phymatodes (M.Howe) R.J.Duf, J.C.Villarreal, Cargill et Renzaglia, Bryologist 110 (2): 240, 2007 (Duf et al. 2007). Bas.: Anthoceros phymatodes M.Howe, Bull. Torrey Bot. Club 25 (1): 12, 1898 (Howe 1898a). Notothylatidae R.J.Duff, J.C.Villarreal, Cargill et Renzaglia Notothyladales Hyvönen et Piippo *** Notothyladaceae Müll.Frib. ex Prosk. by J.C. Villarreal and D.C. Cargill World checklist of hornworts and liverworts ** 37 Notothyladoideae Grolle *** Notothylas Sull. ex A.Gray, Amer. J. Sci. Arts (ser. 2) 1 (1): 74, 1846 (Gray 1846). ** subg. Notothylas *** Notothylas anaporata Udar et D.K.Singh, Rev. Bryol. Lichénol. 45 (2): 202, 1979 (Udar and Singh 1979b). *** Notothylas breutelii (Gottsche) Gottsche, Bot. Zeitung (Berlin) Beil. 16: 21, 1858 (Gottsche 1858). Bas.: Anthoceros breutelii Gottsche, Syn. Hepat. 4: 583, 1846 (Gottsche et al. 1846). *** Notothylas depressispora J.Haseg., Acta Phytotax. Geobot. 30 (1/3): 26, 1979 (Hasegawa 1979). *** Notothylas dissecta Steph., Sp. Hepat. (Stephani) 5: 1020, 1917 (Stephani 1917b). *** Notothylas himalayensis Udar et D.K.Singh, J. Bryol. 11 (3): 451, 1981 (Udar and Singh 1981a). *** Notothylas indica Kashyap, Proc. Lahore Philos. Soc. 4: 54, 1925 (Kashyap and Dutt 1925). *** Notothylas javanica (Sande Lac.) Gottsche, Bot. Zeitung (Berlin) Beil. 16: 20, 1858 (Gottsche 1858). Bas.: Blasia javanica Sande Lac., Syn. hepat. jav.: 94, 1856 [1857] (Sande Lacoste 1856b). *** Notothylas orbicularis (Schwein.) Sull., Amer. J. Sci. Arts (ser. 2) 1 (1): 75, 1846 (Gray 1846). Bas.: Targionia orbicularis Schwein., Spec. Fl. Amer. Crypt.: 23, 1821 (Schweinitz 1821). *** Notothylas pandei Udar et V.Chandra, Geophytology 7 (2): 142, 1977 (Udar and Chandra 1977). ** subg. Notothyloides A.K.Asthana et S.C.Srivast., Bryophyt. Biblioth. 42: 106, 1991 (Asthana and Srivastava 1991). *** Notothylas khasiana Udar et D.K.Singh, J. Indian Bot. Soc. 60: 112, 1981 (Udar and Singh 1981b). *** Notothylas pleidereri Udar et D.K.Singh, Lindbergia 5 (1): 28, 1979 (Udar and Singh 1979a). Incertae sedis *** Notothylas decurva (Mitt.) Steph., Cat. Afr. Pl. (Hiern) 2 (2): 320, 1901 (Stephani 1901d). Bas.: Anthoceros decurvus Mitt., Trans. Linn. Soc. London 23 (1): 58, 1860 (Mitten 1860a). *** Notothylas labellata Steph., Cat. Afr. Pl. (Hiern) 2 (2): 320, 1901 (Stephani 1901d). *** Notothylas galapagensis M.Howe, Proc. Calif. Acad. Sci. (ser. 4) 21 (17): 203, 1934 (Howe 1934). ** Notothylas irregularis Chantanaorr., Acta Bot. Hung. 56 (3/4): 270, 2014 (Chantanaorrapint 2014). 38 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Notothylas kashyapii D.K.Singh, Indian J. Forest. 23 (4): 386, 2000 (Singh and Semwal 2000). *** Notothylas nepalensis D.K.Singh, J. Bombay Nat. Hist. Soc. 84 (3): 650, 1987 [1988] (Singh 1987a). *** Notothylas temperata J.Haseg., Acta Phytotax. Geobot. 30 (1/3): 20, 1979 (Hasegawa 1979). *** Notothylas udarii D.K.Singh et Semwal, Phytotaxonomy 1: 35, 2001 (Singh and Semwal 2001). * Notothylas verdoornii Khanna, Rev. Bryol. Lichénol. 6: 118, 1933 (Khanna 1933). *** Notothylas vitalii Udar et D.K.Singh, Misc. Bryol. Lichenol. 8 (9): 173, 1980 (Udar and Singh 1980). ** Notothylas yunnanensis T.Peng et R.L.Zhu, Phytotaxa 156 (3): 157, 2014 (Peng and Zhu 2014). ** Phaeocerotoideae Hässel * * * Mesoceros Piippo, Acta Bot. Fenn. 148: 30, 1993 (Piippo 1993b). 20 Mesoceros mesophoros Piippo, Acta Bot. Fenn. 148: 30, 1993 (Piippo 1993b). Mesoceros porcatus Piippo, Haussknechtia, Beih. 9: 279, 1999 (Piippo 1999). ** Paraphymatoceros Hässel, Phytologia 88 (2): 208, 2006 (Hässel 2006a). *** Paraphymatoceros diadematus Hässel, Phytologia 88 (2): 209, 2006 (Hässel 2006a). *** Paraphymatoceros hallii (Austin) Hässel, Phytologia 88 (2): 209, 2006 (Hässel 2006a). Bas.: Anthoceros hallii Austin, Bull. Torrey Bot. Club 6 (4): 26, 1875 [1876] (Austin 1875a). *** Paraphymatoceros pearsonii (M.Howe) J.C.Villarreal et Cargill, Phytotaxa 208 (1): 94, 2015 (Villarreal et al. 2015). Bas.: Anthoceros pearsonii M.Howe, Bull. Torrey Bot. Club 25 (1): 8, 1898 (Howe 1898a). *** Paraphymatoceros proskaueri (Stotler, Crand.-Stotl. et W.T.Doyle) J.C.Villarreal et Cargill, Phytotaxa 208 (1): 94, 2015 (Villarreal et al. 2015). Bas.: Phaeoceros proskaueri Stotler, Crand.-Stotl. et W.T.Doyle, Fieldiana, Bot. (n.ser.) 47: 232, 2008 (Crandall-Stotler et al. 2008a). *** Phaeoceros Prosk., Bull. Torrey Bot. Club 78 (4): 346, 1951 (Proskauer 1951a). ** Phaeoceros austroandinus Hässel, Candollea 44 (2): 721, 1989 (Hässel 1989a). ** Phaeoceros bolusii (Sim) S.W.Arnell, Hepat. South Africa: 401, 1963 (Arnell 1963b). Bas.: Anthoceros bolusii Sim, Trans. Roy. Soc. South Africa 15 (1): 114, 1926 (Sim 1926). 20 Mesoceros was described from a collection that contains a mixture of both Anthoceros and Phaeoceros material. World checklist of hornworts and liverworts 39 ** Phaeoceros brevicapsulus (Steph.) Hässel, Candollea 44 (2): 725, 1989 (Hässel 1989a). Bas.: Anthoceros brevicapsulus Steph., Sp. Hepat. (Stephani) 5: 981, 1916 (Stephani 1916b). *** Phaeoceros carolinianus (Michx.) Prosk., Bull. Torrey Bot. Club 78 (4): 347, 1951 (Proskauer 1951a). Bas.: Anthoceros carolinianus Michx., Fl. bor.-amer. (Michaux) 2: 280, 1803 (Michaux 1803). *** Phaeoceros delicatus E.O.Campb. et Outred, New Zealand J. Bot. 33 (3): 285, 1995 (Campbell and Outred 1995). *** Phaeoceros dendroceroides (Steph.) Hässel, Beih. Nova Hedwigia 134: 487, 2009 (Hässel and Rubies 2009). Bas.: Anthoceros dendroceroides Steph., Sp. Hepat. (Stephani) 5: 984, 1916 (Stephani 1916b). 21 *** Phaeoceros engelii Cargill et Fuhrer, Fieldiana, Bot. (n.ser.) 47: 248, 2008 (Cargill and Fuhrer 2008). ** Phaeoceros erectus Udar et D.K.Singh, Geophytology 11 (2): 257, 1981 (Udar and Singh 1981c). Nom. nov. pro Anthoceros erectus Steph., Sp. Hepat. (Stephani) 5: 984, 1916 (Stephani 1916b), nom. illeg. *** Phaeoceros evanidus (Steph.) Cargill et Fuhrer, Fieldiana, Bot. (n.ser.) 47: 245, 2008 (Cargill and Fuhrer 2008). Bas.: Anthoceros evanidus Steph., Sp. Hepat. (Stephani) 5: 990, 1916 (Stephani 1916b). *** Phaeoceros exiguus (Steph.) J.Haseg., J. Hattori Bot. Lab. 60: 387, 1986 (Hasegawa 1986b). Bas.: Anthoceros exiguus Steph., Sp. Hepat. (Stephani) 5: 988, 1916 (Stephani 1916b). *** Phaeoceros lexivalvis (Gottsche et Nees) Hässel, Candollea 44 (2): 728, 1989 (Hässel 1989a). Bas.: Anthoceros lexivalvis Gottsche et Nees, Syn. Hepat. 4: 586, 1846 (Gottsche et al. 1846). *** Phaeoceros fulvisporus (Steph.) J.Haseg., Trop. Bryol. 8: 52, 1993 (Hasegawa 1993a). Bas.: Anthoceros fulvisporus Steph., 52 (5): 306, 1912 (Stephani 1912a). *** Phaeoceros gemmifer (Horik.) J.Haseg., J. Hattori Bot. Lab. 57: 252, 1984 (Hasegawa 1984). Bas.: Anthoceros gemmifer Horik., Sci. Rep. Tôhoku Imp. Univ., Ser. 4, Biol. 4 (2): 426, 1929 (Horikawa 1929a). ** Phaeoceros gualaquizanus (Steph.) Gradst., J. Hattori Bot. Lab. 45: 123, 1979 (Gradstein and Hekking 1979). Bas.: Anthoceros gualaquizanus Steph., Sp. Hepat. (Stephani) 5: 979, 1916 (Stephani 1916b). *** Phaeoceros himalayensis (Kashyap) Prosk. ex Bapna et G.G.Vyas, J. Hattori Bot. Lab. 25: 88, 1962 (Bapna and Vyas 1962). Bas.: Anthoceros himalayensis Kashyap, New Phytol. 14 (1): 8, 1915 (Kashyap 1915). *** Phaeoceros huebschmannii Hässel, Veröf. Geobot. Inst. ETH Stiftung Rübel Zürich 91: 301, 1986 (Hässel 1986b). *** Phaeoceros inlatus (Steph.) Cargill et Fuhrer, Fieldiana, Bot. (n.ser.) 47: 246, 2008 (Cargill and Fuhrer 2008). Bas.: Anthoceros inlatus Steph., Sp. Hepat. (Stephani) 5: 990, 1916 (Stephani 1916b). 21 Phaeoceros dendroceroides may belong to Phaeomegaceros 40 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Phaeoceros kashyapii A.K.Asthana et S.C.Srivast., Bryophyt. Biblioth. 42: 129, 1991 (Asthana and Srivastava 1991). *** Phaeoceros laevis (L.) Prosk., Bull. Torrey Bot. Club 78 (4): 347, 1951 (Proskauer 1951a). Bas.: Anthoceros laevis L., Sp. Pl. 1: 1139, 1753 (Linnaeus 1753). ** Phaeoceros maranguensis (Steph.) Bapna, Univ. Udaipur Res. Stud. 3: 138, 1966 (Bapna 1966). Bas.: Aspiromitus maranguensis Steph., Sp. Hepat. (Stephani) 5: 960, 1916 (Stephani 1916b). * Phaeoceros microsporus (Steph.) Hässel, Candollea 44 (2): 721, 1989 (Hässel 1989a). Bas.: Aspiromitus microsporus Steph., Sp. Hepat. (Stephani) 5: 963, 1916 (Stephani 1916b). 22 *** Phaeoceros minutus (Mitt.) S.W.Arnell, Hepat. South Africa: 403, 1963 (Arnell 1963b). Bas.: Anthoceros minutus Mitt., J. Linn. Soc., Bot. 16 (91): 195, 1877 (Mitten 1877). *** Phaeoceros mohrii (Austin) Hässel, Candollea 44 (2): 721, 1989 (Hässel 1989a). Bas.: Anthoceros mohrii Austin, Bull. Torrey Bot. Club 6 (52): 304, 1879 (Austin 1879). *** Phaeoceros oreganus (Austin) Hässel, Candollea 44 (2): 718, 1989 (Hässel 1989a). Bas.: Anthoceros oreganus Austin, Bull. Torrey Bot. Club 6 (4): 26, 1875 [1876] (Austin 1875a). *** Phaeoceros parvulus (Schifn.) J.Haseg., J. Hattori Bot. Lab. 57: 248, 1984 (Hasegawa 1984). Bas.: Anthoceros parvulus Schifn., Österr. Bot. Z. 49 (11): 391, 1899 (Schifner 1899c). *** Phaeoceros perpusillus Chantanaorr., Acta Bot. Hung. 51 (1/2): 30, 2009 (Chantanaorrapint 2009). ** Phaeoceros pichinchensis (Spruce) Hässel, Candollea 44 (2): 725, 1989 (Hässel 1989a). Bas.: Anthoceros pichinchensis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 577, 1885 (Spruce 1885). ** Phaeoceros propagulifer (Steph.) Prosk., Bull. Torrey Bot. Club 78 (4): 347, 1951 (Proskauer 1951a). Bas.: Anthoceros propagulifer Steph., Sp. Hepat. (Stephani) 5: 1001, 1916 (Stephani 1916b). * Phaeoceros striatisporus J.Haseg., J. Hattori Bot. Lab. 75, 268 (Hasegawa 1994a).23 *** Phaeoceros tenuis (Spruce) Hässel, Veröf. Geobot. Inst. ETH Stiftung Rübel Zürich 91: 303, 1986 (Hässel 1986b). Bas.: Anthoceros tenuis Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: cxcvi, 1889 [1890] (Spruce 1889). ** Phaeoceros tigrinus (Gola) J.C.Villarreal, Phytotaxa 208 (1): 95, 2015 (Villarreal et al. 2015). Bas.: Anthoceros tigrinus Gola, Ann. Bot. (Rome) 13 (1): 72, 1914 (Gola 1914a). *** Phaeoceros tuberosus (Taylor) Prosk., J. Indian Bot. Soc. 42A: 185, 1964 (Proskauer 1964). Bas.: Anthoceros tuberosus Taylor, London J. Bot. 5: 412, 1846 (Taylor 1846b). *** Phaeoceros udarii A.K.Asthana et V.Nath, Proc. Natl. Acad. Sci. India, B 63 (4): 461, 1993 (Asthana and Nath 1993). ** Phaeoceros wrightii (Steph.) Hässel, Candollea 44 (2): 722, 1989 (Hässel 1989a). Bas.: Anthoceros wrightii Steph., Sp. Hepat. (Stephani) 5: 999, 1916 (Stephani 1916b). 22 Phaeoceros microsporus may be conspeciic with Phaeoceros mohrii in Schuster (1992d), but he did not study any material. 23 Phaeoceros striatisporus was nested within Phaeoceros in Li et al. (2011). Morphologically it was supposed to difer by its unique spores which, however, turns out to be fungal infections (Villarreal & Renner 2012). World checklist of hornworts and liverworts 41 LEIOSPOROCEROTOPSIDA Stotler et Crand.-Stotl. Leiosporocerotales Hässel *** Leiosporocerotaceae Hässel ex Ochyra by J.C. Villarreal and D.C. Cargill *** Leiosporoceros Hässel, J. Bryol. 14 (2): 255, 1986 (Hässel 1986a). *** Leiosporoceros dussii (Steph.) Hässel, J. Bryol. 14 (2): 255, 1986 (Hässel 1986a). Bas.: Anthoceros dussii Steph., Hedwigia 32 (3): 142, 1893 (Stephani 1893b). MARCHANTIOPHYTA HAPLOMITRIOPSIDA Stotler et Crand.-Stotl. Haplomitriidae Stotler et Crand.-Stotl. Calobryales Hamlin *** Haplomitriaceae Dědeček by S. Bartholomew-Began he treatment of Haplomitriaceae follows Bartholomew-Began (1991). Morphogenetic and molecular studies support the infrageneric ranks of Haplomitrium (CrandallStotler et al. 2009, Forrest et al. 2006). *** Haplomitrium Nees, Naturgesch. Eur. Leberm. 1: 109, 1833 (Nees 1833c) nom. conserv. *** subg. Calobryum (Nees) R.M.Schust., Nova Hedwigia 13 (1/2): 40, 1967 (Schuster 1967c). Bas.: Calobryum Nees, Syn. Hepat. 4: 507, 1846 (Gottsche et al. 1846). *** Haplomitrium blumei (Nees) R.M.Schust., J. Hattori Bot. Lab. 26: 225, 1963 (Schuster 1963b). Bas.: Monoclea blumei Nees, Enum. Pl. Crypt. Javae: 2, 1830 (Nees 1830). *** Haplomitrium mnioides (Lindb.) R.M.Schust., J. Hattori Bot. Lab. 26: 225, 1963 (Schuster 1963b). Bas.: Rhopalanthus mnioides Lindb., Not. Sällsk. Fauna Fl. Fenn. Förh. 13: 391, 1874 (Lindberg 1874a). *** subg. Haplomitrium ** sect. Archibryum (R.M.Schust.) J.J.Engel, Ann. Missouri Bot. Gard. 68 (4): 675, 1981 (Engel 1981). Bas.: Haplomitrium subg. Archibryum R.M.Schust., Nova Hedwigia 13 (1/2): 28, 1967 (Schuster 1967c). 42 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Haplomitrium gibbsiae (Steph.) R.M.Schust., J. Hattori Bot. Lab. 26: 225, 1963 (Schuster 1963b). Bas.: Calobryum gibbsiae Steph., Sp. Hepat. (Stephani) 6: 76, 1917 (Stephani 1917a). *** Haplomitrium intermedium Berrie, Proc. Linn. Soc. New South Wales (ser. 2) 87 (399): 191, 1963 (Berrie 1963). ** sect. Haplomitrium *** Haplomitrium hookeri (Lyell ex Sm.) Nees, Naturgesch. Eur. Leberm. 1: 111, 1833 (Nees 1833c). Bas.: Jungermannia hookeri Lyell ex Sm., Engl. Bot. 35: tab. 2555, 1814 (Smith and Sowerby 1814). *** Haplomitrium hookeri var. minutum (E.O.Campb.) Barthol.-Began, Bryophyt. Biblioth. 41: 230, 1991 (Bartholomew-Began 1991). Bas.: Steereomitrium minutum E.O.Campb., Mem. New York Bot. Gard. 45: 569, 1987 (Campbell 1987). *** Haplomitrium monoicum J.J.Engel, Ann. Missouri Bot. Gard. 68 (4): 668, 1981 [1982] (Engel 1981). *** Haplomitrium ovalifolium R.M.Schust., Bryologist 74 (2): 136, 1971 (Schuster 1971d). Treubiidae Stotler et Crand.-Stotl. Treubiales Schljakov *** Treubiaceae Verd. by R. Stotler and B.J. Crandall-Stotler Stech et al. (2002) showed a clear molecular distinction between Treubia and Apotreubia, which is in accordance with their morphological diferences and supports their recognition as separate genera. *** Apotreubia S.Hatt. et Mizut., Bryologist 69 (4): 491, 1966 [1967] (Hattori et al. 1966). *** Apotreubia hortoniae Konstant., Phytotaxa 76 (3): 33, 2013 (Konstantinova et al. 2013b). Based on: Apotreubia hortoniae R.M.Schust. et Konstant., J. Hattori Bot. Lab. 78: 55, 1995 (Schuster and Konstantinova 1995), nom. inval. *** Apotreubia nana (S.Hatt. et Inoue) S.Hatt. et Mizut., Bryologist 69 (4): 492, 1966 [1967] (Hattori et al. 1966). Bas.: Treubia nana S.Hatt. et Inoue, J. Hattori Bot. Lab. 11: 99, 1954 (Hattori and Inoue 1954). ** Apotreubia pusilla (R.M.Schust.) Grolle, Acta Bot. Fenn. 125: 63, 1984 (Grolle and Piippo 1984). Bas.: Treubia pusilla R.M.Schust., Nova Hedwigia 15: 515, 1968 (Schuster 1968b). World checklist of hornworts and liverworts 43 *** Apotreubia yunnanensis Higuchi, Cryptog. Bryol. Lichénol. 19 (4): 321, 1998 (Higuchi 1998). *** Treubia K.I.Goebel, Ann. Jard. Bot. Buitenzorg 9 (1): 1, 1890 [1891] (Goebel 1890) nom. conserv. *** Treubia insignis K.I.Goebel, Ann. Jard. Bot. Buitenzorg 9 (1): 1, 1890 [1891] (Goebel 1890). ** Treubia insignis subsp. bracteata (Steph.) R.M.Schust. et G.A.M.Scott, J. Hattori Bot. Lab. 32: 241, 1969 (Schuster and Scott 1969). Bas.: Treubia bracteata Steph., Bot. Jahrb. Syst. 23 (1/2, 3): 302, 1896 (Stephani 1896a). ** Treubia insignis subsp. caledonica R.M.Schust. et G.A.M.Scott, J. Hattori Bot. Lab. 32: 243, 1969 (Schuster and Scott 1969). ** Treubia insignis subsp. vitiensis R.M.Schust. et G.A.M.Scott, J. Hattori Bot. Lab. 32: 242, 1969 (Schuster and Scott 1969). *** Treubia lacunosa (Colenso) Prosk., Bryologist 58 (3): 199, 1955 (Proskauer 1955). Bas.: Noteroclada lacunosa Colenso, Trans. & Proc. New Zealand Inst. 18: 248, 1886 (Colenso 1886b). ** Treubia lacunosoides T.Pfeif., W.Frey et M.Stech, Nova Hedwigia 75 (1/2): 249, 2002 (Pfeifer et al. 2002). *** Treubia pygmaea R.M.Schust., Phytologia 56 (7): 460, 1985 (Schuster 1985c). *** Treubia scapanioides R.M.Schust., J. Hattori Bot. Lab. 32: 246, 1969 (Schuster and Scott 1969). *** Treubia tahitensis (Nadeaud) Besch., J. Bot. (Morot) 12: 147, 1898 (Bescherelle 1898). Bas.: Gottschea tahitensis Nadeaud, Énum. Pl. Tahiti: 7, 1873 (Nadeaud 1873). *** Treubia tasmanica R.M.Schust. et G.A.M.Scott, J. Hattori Bot. Lab. 32: 248, 1969 (Schuster and Scott 1969). JUNGERMANNIOPSIDA Stotler et Crand.-Stotl. Jungermanniidae Engl. Jungermanniales H.Klinggr. Cephaloziineae Schljakov *** Adelanthaceae Grolle by K. Feldberg, J. Váňa and J. Heinrichs he subfamily Jamesonielloideae was excluded from Lophoziaceae/Scapaniaceae by De Roo et al. (2007) and this status was conirmed by e.g. Vilnet et al. (2010). he family was described and deined by Feldberg et al. (2010a). Some taxonomic and nomenclatural notes can also be found in Feldberg et al. (2010b, 2011), Váňa et al. (2014d). 44 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Adelanthoideae K.Feldberg, Heinrichs et Váňa *** Adelanthus Mitt., J. Proc. Linn. Soc., Bot. 7 (28): 243, 1864 (Mitten 1864b) nom. conserv. ** sect. Adelanthus *** Adelanthus falcatus (Hook.) Mitt., J. Proc. Linn. Soc., Bot. 7 (28): 243, 1864 (Mitten 1864b). Bas.: Jungermannia falcata Hook., Musci Exot. 1: tab. 89, 1818 (Hooker 1818). *** Adelanthus occlusus (Hook.f. et Taylor) Carrington, Trans. Bot. Soc. Edinburgh 10: 381, 1870 (Carrington 1870). Bas.: Jungermannia occlusa Hook.f. et Taylor, London J. Bot. 3: 369, 1844 (Hooker and Taylor 1844a). ** sect. Calyptrocolea (R.M.Schust.) Grolle, J. Hattori Bot. Lab. 35: 331, 1972 (Grolle 1972c). Bas.: Calyptrocolea R.M.Schust., Rev. Bryol. Lichénol. 34 (3/4): 685, 1966 (Schuster 1966a). *** Adelanthus aureomarginatus R.M.Schust., Phytologia 39 (4): 250, 1978 (Schuster 1978a). *** Adelanthus gemmiparus (R.M.Schust.) E.A.Hodgs., Trans. Roy. Soc. New Zealand, Biol. Sci. 11 (18): 241, 1970 (Hodgson 1970). Bas.: Calyptrocolea gemmipara R.M.Schust., Rev. Bryol. Lichénol. 34 (3/4): 695, 1966 [1967] (Schuster 1966a). ** Adelanthus lingulatus J.J.Engel et Váňa, Mem. New York Bot. Gard. 105: 26, 2013 (Váňa and Engel 2013). *** Adelanthus tenuis J.J.Engel et Grolle, J. Hattori Bot. Lab. 35: 333, 1972 (Grolle 1972c). ** sect. Lindenbergiani Grolle, J. Hattori Bot. Lab. 35: 331, 1972 (Grolle 1972c). *** Adelanthus carabayensis (Mont.) Grolle, J. Hattori Bot. Lab. 35: 348, 1972 (Grolle 1972c). Bas.: Plagiochila carabayensis Mont., Ann. Sci. Nat. Bot. (sér. 4) 5: 348, 1856 (Montagne 1856c). *** Adelanthus integerrimus Grolle, J. Hattori Bot. Lab. 35: 340, 1972 (Grolle 1972c). *** Adelanthus lindenbergianus (Lehm.) Mitt., J. Proc. Linn. Soc., Bot. 7 (28): 244, 1864 (Mitten 1864b). Bas.: Jungermannia lindenbergiana Lehm., Linnaea 4: 367, 1829 (Lehmann 1829). ** sect. Pittieri Grolle, J. Hattori Bot. Lab. 35: 331, 1972 (Grolle 1972c). *** Adelanthus pittieri (Steph.) Grolle, J. Hattori Bot. Lab. 35: 337, 1972 (Grolle 1972c). Bas.: Tylimanthus pittieri Steph., Sp. Hepat. (Stephani) 6: 250, 1922 (Stephani 1922). *** Adelanthus squarrosus Grolle, J. Hattori Bot. Lab. 67: 243, 1989 (Grolle 1989d). World checklist of hornworts and liverworts 45 *** Pseudomarsupidium Herzog, Svensk Bot. Tidskr. 47 (1): 42, 1953 (Herzog 1953b). *** Pseudomarsupidium aureocinctum (R.M.Schust.) J.J.Engel, Novon 17 (3): 312, 2007 (Engel 2007). Bas.: Adelanthus decipiens subsp. aureocinctus R.M.Schust., Phytologia 39 (4): 250, 1978 (Schuster 1978a). *** Pseudomarsupidium borneensis (Grolle) Váňa, L.Söderstr., A.Hagborg et von Konrat, Phytotaxa 65: 60, 2012 (Váňa et al. 2012f). Bas.: Adelanthus borneensis Grolle, J. Hattori Bot. Lab. 35: 362, 1972 (Grolle 1972c). *** Pseudomarsupidium decipiens (Hook.) Grolle, Trans. Brit. Bryol. Soc. 4 (3): 443, 1963 (Grolle 1963a). Bas.: Jungermannia decipiens Hook., Brit. Jungermann.: tab. 50, 1813 (Hooker 1813). *** Pseudomarsupidium piliferum (Steph.) Herzog ex Grolle, Trans. Brit. Bryol. Soc. 4 (3): 443, 1963 (Grolle 1963a). Bas.: Marsupidium piliferum Steph., Bull. Herb. Boissier (sér. 2) 8 (8): 602 (386), 1908 (Stephani 1908e). *** Wettsteinia Schifn., Ann. Jard. Bot. Buitenzorg, suppl. 2: 44, 1898 (Schifner 1898c). *** Wettsteinia densiretis (Herzog) Grolle, J. Hattori Bot. Lab. 28: 99, 1965 (Grolle 1965f). Bas.: Tylimanthus densiretis Herzog, Nat. Hist. Juan Fernandez (Botany) 2 (5): 712, 1942 (Herzog 1942a). *** Wettsteinia inversa (Sande Lac.) Schifn., Ann. Jard. Bot. Buitenzorg, suppl. 2: 45, 1898 (Schifner 1898c). Bas.: Plagiochila inversa Sande Lac., Ann. Mus. Bot. Lugduno-Batavi 1: 289, 1864 (Sande Lacoste 1864). *** Wettsteinia rotundifolia (Horik.) Grolle, J. Hattori Bot. Lab. 28: 100, 1965 (Grolle 1965f). Bas.: Adelanthus rotundifolius Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 181, 1934 (Horikawa 1934). *** Wettsteinia schusteriana Grolle, J. Hattori Bot. Lab. 28: 99, 1965 (Grolle 1965f). ** Jamesonielloideae Inoue *** Cuspidatula Steph., Bull. Herb. Boissier (sér. 2) 1 (10): 1141 (124), 1901 (Stephani 1901c). *** Cuspidatula contracta (Reinw., Blume et Nees) Steph., Bull. Herb. Boissier (sér. 2) 1 (11): 1141 (124), 1901 (Stephani 1901c). Bas.: Jungermannia contracta Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 233, 1824 [1825] (Reinwardt et al. 1824a). *** Cuspidatula laccida (Steph.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 142, 2010 (Feldberg et al. 2010b). Bas.: Anastrophyllum laccidum Steph., Sp. Hepat. (Stephani) 6: 105, 1917 (Stephani 1917a). *** Cuspidatula lexicaulis (Nees) Váňa et L.Söderstr., Phytotaxa 76 (3): 35, 2013 (Váňa et al. 2013h). Bas.: Jungermannia lexicaulis Nees, Linnaea 6 (4): 604, 1831 (Nees 1831). 46 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Cuspidatula kirkii (Steph.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 142, 2010 (Feldberg et al. 2010b). Bas.: Jamesoniella kirkii Steph., Hedwigia 34 (2): 47, 1895 (Stephani 1895c). *** Cuspidatula monodon (Taylor) Steph., Bull. Herb. Boissier (sér. 2) 1 (11): 1143 (126), 1901 (Stephani 1901c). Bas.: Jungermannia monodon Taylor, Nov. Stirp. Pug. 8: 7, 1844 (Lehmann 1844). *** Cuspidatula orbicularis (Grolle) Váňa et L.Söderstr., Phytotaxa 76 (3): 36, 2013 (Váňa et al. 2013h). Bas.: Jamesoniella orbicularis Grolle, Feddes Repert. 82 (1): 42, 1971 (Grolle 1971b). *** Cuspidatula robusta (Austin) Váňa et L.Söderstr., Phytotaxa 76 (3): 36, 2013 (Váňa et al. 2013h). Bas.: Jungermannia robusta Austin, Proc. Acad. Nat. Sci. Philadelphia 21: 219, 1869 (Austin 1869). ** Denotarisia Grolle, Feddes Repert. 82 (1): 6, 1971 (Grolle 1971b). *** Denotarisia linguifolia (De Not.) Grolle, Feddes Repert. 82 (1): 6, 1971 (Grolle 1971b). Bas.: Plagiochila linguifolia De Not., Epat. Borneo: 13, 1874 (De Notaris 1874). ** Nothostrepta R.M.Schust., Phytologia 45 (5): 420, 1980 (Schuster 1980b). *** Nothostrepta biida (Steph.) R.M.Schust., Phytologia 45 (5): 420, 1980 (Schuster 1980b). Bas.: Plagiochila biida Steph., Annuario Reale Ist. Bot. Roma 2: 86, 1885 [1886] (Stephani 1885d). *** Nothostrepta longissima (Steph.) R.M.Schust., Phytologia 45 (5): 420, 1980 (Schuster 1980b). Bas.: Anastrophyllum longissimum Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 17): 13, 1901 (Stephani 1901b). ** Pisanoa Hässel, Lindbergia 14 (3): 179, 1988 [1989] (Hässel 1988). *** Pisanoa chilensis Hässel, Lindbergia 14 (3): 179, 1988 [1989] (Hässel 1988). ** Protosyzygiella (Inoue) R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 334, 1980 (Schuster 1980c). Bas.: Syzygiella subg. Protosyzygiella Inoue, J. Hattori Bot. Lab. 29: 180, 1966 (Inoue 1966c). *** Protosyzygiella pseudoconnexa (Herzog) R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 334, 1980 (Schuster 1980c). Bas.: Plagiochila pseudoconnexa Herzog, Rev. Bryol. Lichénol. 21 (3/4): 259, 1952 [1953] (Herzog 1952d). *** Syzygiella Spruce, J. Bot. 14: 234, 1876 (Spruce 1876a). 24 *** subg. Anomalae (Inoue) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 143, 2010 (Feldberg et al. 2010b). Bas.: Syzygiella sect. Anomalae Inoue, J. Hattori Bot. Lab. 29: 183, 1966 (Inoue 1966c). 24 Syzygiella includes Jamesoniella (cf. Feldberg et al. 2010a), but one taxon has neither been transferred nor synonymized. It is listed in the “Names in genera not currently accepted” section below. World checklist of hornworts and liverworts 47 *** Syzygiella anomala (Lindenb. et Gottsche) Steph., Bull. Herb. Boissier (sér. 2) 2 (5): 471 (190), 1902 (Stephani 1902f). Bas.: Plagiochila anomala Lindenb. et Gottsche, Syn. Hepat. 5: 646, 1847 (Gottsche et al. 1847). *** Syzygiella bilobata Inoue, J. Hattori Bot. Lab. 29: 186, 1966 (Inoue 1966c). ** Syzygiella ciliata Gradst. et A.R.Benitez, Nova Hedwigia 99 (1/2): 115, 2014 (Gradstein and Benitez 2014). *** Syzygiella concreta (Gottsche) Spruce, J. Bot. 14: 234, 1876 (Spruce 1876a). Bas.: Jungermannia concreta Gottsche, Mexik. Leverm.: 82, 1863 (Gottsche 1863). *** Syzygiella manca (Mont.) Steph., Hedwigia 31 (1): 14, 1892 (Jack and Stephani 1892). Bas.: Chiloscyphus mancus Mont., Syll. Gen. Sp. Crypt.: 63, 1856 (Montagne 1856b). *** Syzygiella pectiniformis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 501, 1885 (Spruce 1885). *** Syzygiella tonduzana Steph., Sp. Hepat. (Stephani) 6: 118, 1917 (Stephani 1917a). *** Syzygiella trigonifolia (Steph.) Herzog, Hedwigia 74 (2): 87, 1934 (Herzog 1934a). Bas.: Jamesoniella trigonifolia Steph., Biblioth. Bot. 87 (2): 185, 1916 (Stephani 1916a). *** subg. Cryptochila (R.M.Schust.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 143, 2010 (Feldberg et al. 2010b). Bas.: Cryptochila R.M.Schust., J. Hattori Bot. Lab. 26: 284, 1963 (Schuster 1963b). *** Syzygiella acinacifolia (Hook.f. et Taylor) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 143, 2010 (Feldberg et al. 2010b). Bas.: Jungermannia acinacifolia Hook.f. et Taylor, London J. Bot. 3: 367, 1844 (Hooker and Taylor 1844a). *** Syzygiella nigrescens (Steph.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 143, 2010 (Feldberg et al. 2010b). Bas.: Jamesoniella nigrescens Steph., Hedwigia 34 (2): 48, 1895 (Stephani 1895c). *** Syzygiella paludosa (Steph.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 143, 2010 (Feldberg et al. 2010b). Bas.: Jamesoniella paludosa Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 17): 11, 1901 (Stephani 1901b). *** Syzygiella pseudocclusa (E.A.Hodgs.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 143, 2010 (Feldberg et al. 2010b). Bas.: Jamesoniella pseudocclusa E.A.Hodgs., Trans. Roy. Soc. New Zealand 85 (4): 583, 1958 (Hodgson 1958). *** Syzygiella sonderi (Gottsche) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 143, 2010 (Feldberg et al. 2010b). Bas.: Jungermannia sonderi Gottsche, Linnaea 28 (5): 550, 1856 [1857] (Gottsche 1856). *** Syzygiella spegazziniana (Spruce ex C.Massal.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 144, 2010 (Feldberg et al. 2010b). Bas.: Jungermannia spegazziniana Spruce ex C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 216, 1885 (Massalongo 1885). *** subg. Pseudoplagiochila Inoue, J. Hattori Bot. Lab. 29: 182, 1966 (Inoue 1966c). *** Syzygiella ovalifolia Inoue, J. Hattori Bot. Lab. 29: 191, 1966 (Inoue 1966c). 48 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) * Syzygiella securifolia (Nees) Inoue, J. Hattori Bot. Lab. 46: 232, 1979 (Inoue 1979a). Bas.: Plagiochila securifolia Nees, Sp. Hepat. (Lindenberg) 2-4: 58, 1840 (Lindenberg 1840). 25 *** Syzygiella subintegerrima (Reinw., Blume et Nees) Spruce, J. Linn. Soc., Bot. 30 (210): 362, 1895 (Gepp 1895b). Bas.: Jungermannia subintegerrima Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 238, 1824 [1825] (Reinwardt et al. 1824a). *** Syzygiella tasmanica (Hook.f. et Taylor) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 144, 2010 (Feldberg et al. 2010b). Bas.: Jungermannia tasmanica Hook.f. et Taylor, London J. Bot. 5: 274, 1846 (Taylor 1846a). *** subg. Roivainenia (Perss.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 144, 2010 (Feldberg et al. 2010b). Bas.: Roivainenia Perss., Nova Hedwigia 3 (1): 43, 1961 (Persson and Grolle 1961). *** Syzygiella jacquinotii (Mont.) Hentschel, K.Feldberg, Váňa et Heinrichs, Cryptog. Bryol. 31 (2): 144, 2010 (Feldberg et al. 2010b). Bas.: Jungermannia jacquinotii Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 250, 1843 (Montagne 1843). *** subg. Syzygiella *** Syzygiella autumnalis (DC.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 144, 2010 (Feldberg et al. 2010b). Bas.: Jungermannia autumnalis DC., Fl. Franç. (DC. & Lamarck), 5 (6): 202, 1815 (De Candolle and Lamarck 1815). *** Syzygiella campanulata Herzog, Rev. Bryol. Lichénol. 11 (1): 9, 1938 [1939] (Herzog 1938a). *** Syzygiella colorata (Lehm.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 144, 2010 (Feldberg et al. 2010b). Bas.: Jungermannia colorata Lehm., Linnaea 4: 366, 1829 (Lehmann 1829). ** Syzygiella colorata var. collenchymata J.J.Engel et Váňa, Mem. New York Bot. Gard. 105: 102, 2013 (Váňa and Engel 2013). *** Syzygiella contigua Steph., Bull. Herb. Boissier (sér. 2) 2 (5): 470 (189), 1902 (Stephani 1902f). Nom. nov. pro Jungermannia contigua Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 118, 1864 (Gottsche 1864), nom. illeg. *** Syzygiella elongella (Taylor) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 144, 2010 (Feldberg et al. 2010b). Bas.: Jungermannia elongella Taylor, London J. Bot. 5: 274, 1846 (Taylor 1846a). *** Syzygiella macrocalyx (Mont.) Spruce, J. Bot. 14: 234, 1876 (Spruce 1876a). Bas.: Jungermannia macrocalyx Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 248, 1843 (Montagne 1843). *** Syzygiella nipponica (S.Hatt.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 145, 2010 (Feldberg et al. 2010b). Bas.: Jamesoniella nipponica S.Hatt., J. Jap. Bot. 19 (11): 350, 1943 (Hattori 1943b). 25 Syzygiella securifolia is a sister taxon to Syzygiella subintegerrima (Feldberg et al. 2010b) and it is debatable if they are diferent enough to warrant separation on the species level. Morphologically they are very similar (Söderström et al. 2010a). World checklist of hornworts and liverworts 49 ** Syzygiella oenops (Lindenb. et Gottsche) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 145, 2010 (Feldberg et al. 2010b). Bas.: Jungermannia oenops Lindenb. et Gottsche, Syn. Hepat. 5: 673, 1847 (Gottsche et al. 1847). *** Syzygiella perfoliata (Sw.) Spruce, J. Bot. 14: 234, 1876 (Spruce 1876a). Bas.: Jungermannia perfoliata Sw., Prodr. (Swartz): 143, 1788 (Swartz 1788). *** Syzygiella purpurascens (Steph.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 145, 2010 (Feldberg et al. 2010b). Bas.: Jamesoniella purpurascens Steph., Bull. Soc. Roy. Bot. Belgique, Compt. Rend. 30 (2): 200, 1891 [1892] (Stephani 1891b). *** Syzygiella rubricaulis (Nees) Steph., Bull. Herb. Boissier (sér. 2) 2 (5): 468 (187), 1902 (Stephani 1902f). Bas.: Jungermannia rubricaulis Nees, Fl. Bras. (Martius) 1 (1): 344, 1833 (Nees 1833a). *** Syzygiella setulosa Steph., Bull. Herb. Boissier (sér. 2) 2 (5): 469 (188), 1902 (Stephani 1902f). *** Syzygiella teres (Carrington et Pearson) Váňa, Phytotaxa 76 (3): 35, 2013 (Váňa et al. 2013h). Bas.: Jungermannia teres Carrington et Pearson, Pap. & Proc. Roy. Soc. Tasmania 1887: 9, 1888 (Carrington and Pearson 1888b). *** Syzygiella undata (Mont.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 145, 2010 (Feldberg et al. 2010b). Bas.: Jungermannia undata Mont., Ann. Sci. Nat. Bot. (sér. 4) 14: 183, 1860 (Montagne 1860). Incertae sedis *** Syzygiella eatonii (Austin) Inoue, J. Jap. Bot. 37 (12): 359, 1962 (Inoue 1962a). Bas.: Plagiochila eatonii Austin, Trans. Connecticut Acad. Arts 8 (15): 257, 1891 (Evans 1891). *** Syzygiella uleana Steph., Hedwigia 44 (4): 224, 1905 (Stephani 1905a). ** Vanaea (Inoue et Gradst.) Inoue et Gradst., Trop. Bryol. 1: 33, 1989 (Gradstein and Florschütz-de Waard 1989). Bas.: Anastrophyllum subg. Vanaea Inoue et Gradst., Bull. Natl. Sci. Mus. Tokyo, B 14 (3): 88, 1988 (Inoue and Gradstein 1988). *** Vanaea plagiochiloides (Inoue et Gradst.) Inoue et Gradst., Trop. Bryol. 1: 33, 1989 (Gradstein and Florschütz-de Waard 1989). Bas.: Anastrophyllum plagiochiloides Inoue et Gradst., Bull. Natl. Sci. Mus. Tokyo, B 14 (3): 88, 1988 (Inoue and Gradstein 1988). *** Anastrophyllaceae L.Söderstr., De Roo et Hedd. by J. Váňa and L. Söderström Anastrophyllaceae was described by Söderström et. al (2010b) from elements usually included in Lophoziaceae. Further taxonomic and nomenclatural notes can be found in Váňa et al. (2013k, 2013a). he complex of Chandonanthus/Plicanthus/Tetralophozia should be checked using molecular methods before generic and speciic status of most 50 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) of the species can be conirmed. he placement of Hattoria and Zantenia in the family is still provisional. he placement of Isopaches is also unclear, de Roo et al. (2007) placed it in the family, but the study of Vilnet et al. (2010) showed that it can not be placed there. *** Anastrepta (Lindb.) Schifn., Hepat. (Engl.-Prantl): 85, 1893 (Schifner 1893b). Bas.: Jungermannia sect. Anastrepta Lindb., Kongl. Svenska Vetensk.-Akad. Handl. (n.ser.) 23 (5): 40, 1889 (Lindberg and Arnell 1889). *** Anastrepta orcadensis (Hook.) Schifn., Hepat. (Engl.-Prantl): 85, 1893 (Schifner 1893b). Bas.: Jungermannia orcadensis Hook., Brit. Jungermann.: tab. 71, 1815 (Hooker 1815). *** Anastrophyllum (Spruce) Steph., Hedwigia 32 (3): 139, 1893 (Stephani 1893b). Bas.: Jungermannia subg. Anastrophyllum Spruce, J. Bot. 14: 235, 1876 (Spruce 1876a). *** Anastrophyllum alpinum Steph., Sp. Hepat. (Stephani) 6: 103, 1917 (Stephani 1917a). *** Anastrophyllum assimile (Mitt.) Steph., Hedwigia 32 (3): 140, 1893 (Stephani 1893b). Bas.: Jungermannia assimilis Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 93, 1860 [1861] (Mitten 1860c). *** Anastrophyllum auritum (Lehm.) Steph., Bull. Herb. Boissier (sér. 2) 1 (11): 1137 (120), 1901 (Stephani 1901c). Bas.: Jungermannia aurita Lehm., Linnaea 4: 368, 1829 (Lehmann 1829). *** Anastrophyllum ciliatum Steph., Hedwigia 32 (3): 139, 1893 (Stephani 1893b). *** Anastrophyllum donnianum (Hook.) Steph., Hedwigia 32 (3): 140, 1893 (Stephani 1893b). Bas.: Jungermannia donniana Hook., Brit. Jungermann.: tab. 39, 1813 (Hooker 1813). *** Anastrophyllum ellipticum Inoue, Bull. Natl. Sci. Mus. Tokyo, B 4 (1): 13, 1978 (Inoue 1978b). 26 ** Anastrophyllum esenbeckii (Mont.) Steph., Hedwigia 32 (3): 140, 1893 (Stephani 1893b). Bas.: Jungermannia esenbeckii Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 247, 1843 (Montagne 1843). *** Anastrophyllum issum Steph., Bull. Herb. Boissier 5 (10): 845, 1897 (Stephani 1897c). *** Anastrophyllum joergensenii Schifn., Hedwigia 49 (4): 396, 1910 (Schifner 1910b). * Anastrophyllum lignicola D.B.Schill et D.G.Long, Ann. Bot. Fenn. 39 (2): 130, 2002 (Schill and Long 2002). 27 *** Anastrophyllum michauxii (F.Weber) H.Buch, Memoranda Soc. Fauna Fl. Fennica 8: 289, 1932 [1933] (Buch 1932). Bas.: Jungermannia michauxii F.Weber, Hist. Musc. Hepat. Prodr.: 76, 1815 (Weber 1815). 26 Anastrophyllum ellipticum is only tentatively placed in the genus following Mamontov & Vilnet (2013). 27 Anastrophyllum lignicola is possibly conspeciic with Anastrophyllum ellipticum and its generic placement is uncertain. World checklist of hornworts and liverworts 51 *** Anastrophyllum nigrescens (Mitt.) Steph., Hedwigia 32 (3): 140, 1893 (Stephani 1893b). Bas.: Jungermannia nigrescens Mitt., Hooker’s J. Bot. Kew Gard. Misc. 3: 358, 1851 (Mitten 1851). *** Anastrophyllum obtusum Herzog, Trans. Brit. Bryol. Soc. 1 (4): 285, 1950 (Herzog 1950a). *** Anastrophyllum piligerum (Nees) Steph., Hedwigia 32 (3): 140, 1893 (Stephani 1893b). Bas.: Jungermannia piligera Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 414, 1824 [1825] (Reinwardt et al. 1824b). *** Anastrophyllum squarrosum Herzog, Ann. Bryol. 5: 72, 1932 (Herzog 1932b). *** Anastrophyllum stellatum R.M.Schust., Phytologia 39 (4): 243, 1978 (Schuster 1978a). *** Anastrophyllum tubulosum (Nees) Grolle, J. Hattori Bot. Lab. 28: 101, 1965 (Grolle 1965e). Bas.: Jungermannia tubulosa Nees, Enum. Pl. Crypt. Javae: 32, 1830 (Nees 1830). *** Barbilophozia Loeske, Verh. Bot. Vereins Prov. Brandenburg 49 (1): 37, 1908 (Loeske 1908). ** subg. Barbilophozia *** Barbilophozia barbata (Schmidel ex Schreb.) Loeske, Verh. Bot. Vereins Prov. Brandenburg 49 (1): 37, 1908 (Loeske 1908). Bas.: Jungermannia barbata Schmidel ex Schreb., Spic. Fl. Lips.: 107, 1771 (Schreber 1771). *** Barbilophozia hatcheri (A.Evans) Loeske, Verh. Bot. Vereins Prov. Brandenburg 49 (1): 37, 1908 (Loeske 1908). Bas.: Jungermannia hatcheri A.Evans, Bull. Torrey Bot. Club 25 (8): 417, 1898 (Evans 1898). *** Barbilophozia lycopodioides (Wallr.) Loeske, Verh. Bot. Vereins Prov. Brandenburg 49 (1): 37, 1908 (Loeske 1908). Bas.: Jungermannia lycopodioides Wallr., Comp. l. Germ. 2 (III): 76, 1831 (Bluf and Fingerhuth 1831). *** Barbilophozia rubescens (R.M.Schust. et Damsh.) Kartt. et L.Söderstr., Ann. Bot. Fenn. 29 (2): 120, 1992 (Söderström et al. 1992). Bas.: Lophozia rubescens R.M.Schust. et Damsh., Phytologia 63 (5): 325, 1987 (Schuster and Damsholt 1987). ** subg. Sudeticae (Schljakov) L.Söderstr., De Roo et Hedd., Phytotaxa 3: 50, 2010 (Söderström et al. 2010b). Bas.: Lophozia sect. Sudeticae Schljakov, Pečen. Mchi Sev. SSSR 3: 113, 1980 (Shliakov 1980a). *** Barbilophozia sudetica (Nees ex Huebener) L.Söderstr., De Roo et Hedd., Phytotaxa 3: 50, 2010 (Söderström et al. 2010b). Bas.: Jungermannia sudetica Nees ex Huebener, Hepaticol. germ.: 142, 1834 (Hübener 1834). 28 28 Barbilophozia sudetica was transferred from Lophozia by Söderström et al. (2010a), but transferred to the new genus Pseudolophozia Konstant. and Vilnet (sister to Barbilophozia) by Konstantinova and Vilnet (2009). 52 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Biantheridion (Grolle) Konstant. et Vilnet, Arctoa 18: 67, 2009 [2010] (Konstantinova and Vilnet 2009). Bas.: Jamesoniella sect. Biantheridion Grolle, Trans. Brit. Bryol. Soc. 4 (4): 662, 1964 (Grolle 1964i). *** Biantheridion undulifolium (Nees) Konstant. et Vilnet, Arctoa 18: 67, 2009 [2010] (Konstantinova and Vilnet 2009). Bas.: Jungermannia schraderi β undulifolia Nees, Naturgesch. Eur. Leberm. 1: 306, 1833 (Nees 1833c). *** Chandonanthus Mitt., Handb. N. Zeal. l. 2: 753, 1867 (Hooker 1867). *** Chandonanthus squarrosus (Menzies) Mitt., Handb. N. Zeal. l. 2: 753, 1867 (Hooker 1867). Bas.: Jungermannia squarrosa Menzies, Musci Exot. 1: tab. 78, 1818 (Hooker 1818). *** Crossocalyx Meyl., Bull. Soc. Vaud. Sci. Nat. 60 (249): 266, 1939 (Meylan 1939). *** Crossocalyx hellerianus (Nees ex Lindenb.) Meyl., Bull. Soc. Vaud. Sci. Nat. 60 (249): 266, 1939 (Meylan 1939). Bas.: Jungermannia helleriana Nees ex Lindenb., Syn. hepat. eur: 64, 1829 (Lindenberg 1829). *** Crossocalyx tenuis (Harry Williams) Schljakov, Novosti Sist. Nizš. Rast. 15: 246, 1978 (Shliakov 1978). Bas.: Anastrophyllum tenue Harry Williams, Bryologist 71 (1): 34, 1968 (Williams 1968). *** Gymnocolea (Dumort.) Dumort., Recueil Observ. Jungerm.: 17, 1835 (Dumortier 1835). Bas.: Jungermannia sect. Gymnocolea Dumort., Syll. Jungerm. Europ.: 52, 1831 (Dumortier 1831). *** Gymnocolea borealis (Frisvoll et Moen) R.M.Schust., Lindbergia 12 (1): 7, 1986 (Schuster 1986). Bas.: Lophozia borealis Frisvoll et Moen, Lindbergia 6 (2): 138, 1980 [1981] (Frisvoll and Moen 1980). ** Gymnocolea fascinifera Potemkin, Arctoa 2: 76, 1993 (Potemkin 1993). *** Gymnocolea inlata (Huds.) Dumort., Recueil Observ. Jungerm.: 17, 1835 (Dumortier 1835). Bas.: Jungermannia inlata Huds., Fl. Angl. (Hudson), ed. 2: 511, 1778 (Hudson 1778). * Gymnocolea inlata subsp. acutiloba (Schifn.) R.M.Schust. et Damsh. ex L.Söderstr. et Váňa, Lindbergia 27 (1): 43, 2002 (Söderström et al. 2002). Bas.: Lophozia acutiloba Schifn., Hedwigia 48 (3): 187, 1909 (Schifner 1909a). 29 *** Hamatostrepta Váňa et D.G.Long, Fieldiana, Bot. (n.ser.) 47: 134, 2008 (Váňa and Long 2008). *** Hamatostrepta concinna Váňa et D.G.Long, Fieldiana, Bot. (n.ser.) 47: 134, 2008 (Váňa and Long 2008). 29 Gymnocolea inlata subsp. acutiloba is a problematic taxon that sometimes has been treated as conspeciic with Gymnocolea inlata, sometimes as a separate species. World checklist of hornworts and liverworts 53 ** Hattoria R.M.Schust., Rev. Bryol. Lichénol. 30 (1/2): 69, 1961 (Schuster 1961a). *** Hattoria yakushimensis (Horik.) R.M.Schust., Rev. Bryol. Lichénol. 30 (1/2): 70, 1961 (Schuster 1961a). Bas.: Anastrophyllum yakushimense Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 149, 1934 (Horikawa 1934). *** Isopaches H.Buch, Memoranda Soc. Fauna Fl. Fennica 8: 287, 1932 [1933] (Buch 1932). *** Isopaches alboviridis (R.M.Schust.) Schljakov, Novosti Sist. Nizš. Rast. 16: 205, 1979 (Shliakov 1979). Bas.: Lophozia alboviridis R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 487, 1969 (Schuster 1969b). *** Isopaches bicrenatus (Schmidel ex Hofm.) H.Buch, Memoranda Soc. Fauna Fl. Fennica 8: 288, 1932 [1933] (Buch 1932). Bas.: Jungermannia bicrenata Schmidel ex Hofm., Deutschl. Fl., heil 2 (Hofm.): 11 (addenda), 1795 [1796] (Hofmann 1795). *** Isopaches decolorans (Limpr.) H.Buch, Memoranda Soc. Fauna Fl. Fennica 8: 288, 1932 [1933] (Buch 1932). Bas.: Jungermannia decolorans Limpr., Jahresber. Schles. Ges. Vaterl. Cult. 57: 316, 1879 [1880] (Limpricht 1879). *** Isopaches pumicicola (Berggr.) Bakalin, Arctoa 17: 162, 2008 [2009] (Bakalin 2008b). Bas.: Lophozia pumicicola Berggr., New Zealand Hepat.: 21, 1898 (Berggren 1898). *** Neoorthocaulis L.Söderstr., De Roo et Hedd., Phytotaxa 3: 49, 2010 (Söderström et al. 2010b). *** Neoorthocaulis attenuatus (Mart.) L.Söderstr., De Roo et Hedd., Phytotaxa 3: 49, 2010 (Söderström et al. 2010b). Bas.: Jungermannia quinquedentata δ attenuata Mart., Fl. crypt. erlang.: 177, 1817 (Martius 1817). *** Neoorthocaulis binsteadii (Kaal.) L.Söderstr., De Roo et Hedd., Phytotaxa 3: 49, 2010 (Söderström et al. 2010b). Bas.: Jungermannia binsteadii Kaal., Skr. Vidensk.Selsk. Christiana, Math.-Naturvidensk. Kl. 1898 (9): 9, 1898 (Kaalaas 1898). *** Neoorthocaulis loerkei (F.Weber et D.Mohr) L.Söderstr., De Roo et Hedd., Phytotaxa 3: 50, 2010 (Söderström et al. 2010b). Bas.: Jungermannia loerkei F.Weber et D.Mohr, Bot. Taschenb. (Weber): 410, 1807 (Weber and Mohr 1807). ** Neoorthocaulis hyperboreus (R.M.Schust.) L.Söderstr., De Roo et Hedd., Phytotaxa 3: 50, 2010 (Söderström et al. 2010b). Bas.: Lophozia loerkei var. hyperborea R.M.Schust., Bull. Natl. Mus. Canada 164: 21, 1959 (Schuster et al. 1959). *** Orthocaulis H.Buch, Memoranda Soc. Fauna Fl. Fennica 8: 293, 1932 [1933] (Buch 1932). *** Orthocaulis atlanticus (Kaal.) H.Buch, Memoranda Soc. Fauna Fl. Fennica 8: 294, 1932 [1933] (Buch 1932). Bas.: Jungermannia atlantica Kaal., Skr. Vidensk.-Selsk. Christiana, Math.-Naturvidensk. Kl. 1898 (9): 11, 1898 (Kaalaas 1898). * Orthocaulis cavifolius H.Buch et S.W.Arnell, Memoranda Soc. Fauna Fl. Fennica 26: 71, 1951 (Buch 1951). 54 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Plicanthus R.M.Schust., Nova Hedwigia 74 (3/4): 484, 2002 (Schuster 2002a). *** Plicanthus birmensis (Steph.) R.M.Schust., Beih. Nova Hedwigia 119: 223, 2002 (Schuster 2002b). Bas.: Chandonanthus birmensis Steph., Bull. Soc. Roy. Bot. Belgique 38 (1): 43, 1899 (Stephani 1899h). 30 ** Plicanthus diicilis (Steph.) L.Söderstr. et Váňa, Phytotaxa 81 (1): 30, 2013 (Váňa et al. 2013a). Bas.: Chandonanthus diicilis Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 101, 1914 (Stephani and Watts 1914). ** Plicanthus giganteus (Steph.) R.M.Schust., Nova Hedwigia 74 (3/4): 485, 2002 (Schuster 2002a). Bas.: Chandonanthus giganteus Steph., Wiss. Ergebn. Deut. Zentr.-Afr. Exped. (1907-08), Bot. 2: 124, 1911 (Stephani 1911a). 31 *** Plicanthus hirtellus (F.Weber) R.M.Schust., Nova Hedwigia 74 (3/4): 492, 2002 (Schuster 2002a). Bas.: Jungermannia hirtella F.Weber, Hist. Musc. Hepat. Prodr.: 50, 1815 (Weber 1815). *** Schizophyllopsis Váňa et L.Söderstr., Phytotaxa 152 (1): 48, 2013 (Váňa et al. 2013f). Nom. nov. pro Anastrophyllum subg. Schizophyllum R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 739, 1969 (Schuster 1969b). 32 ** Schizophyllopsis aristata (Herzog ex N.Kitag.) Váňa et L.Söderstr., Phytotaxa 152 (1): 48, 2013 (Váňa et al. 2013f). Bas.: Anastrophyllum bidens var. aristatum Herzog ex N.Kitag., J. Hattori Bot. Lab. 33: 216, 1970 (Kitagawa 1970). *** Schizophyllopsis bidens (Reinw., Blume et Nees) Váňa et L.Söderstr., Phytotaxa 152 (1): 48, 2013 (Váňa et al. 2013f). Bas.: Jungermannia bidens Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 208, 1824 [1825] (Reinwardt et al. 1824a). ** Schizophyllopsis lanciloba (Steph.) Váňa et L.Söderstr., Phytotaxa 152 (1): 48, 2013 (Váňa et al. 2013f). Bas.: Anastrophyllum lancilobum Steph., Sp. Hepat. (Stephani) 6: 107, 1917 (Stephani 1917a). *** Schizophyllopsis papillosa (J.J.Engel et Braggins) Váňa et L.Söderstr., Phytotaxa 152 (1): 48, 2013 (Váňa et al. 2013f). Bas.: Anastrophyllum papillosum J.J.Engel et Braggins, J. Bryol. 20 (2): 381, 1998 (Engel and Braggins 1998). *** Schizophyllopsis sphenoloboides (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 152 (1): 49, 2013 (Váňa et al. 2013f). Bas.: Anastrophyllum sphenoloboides R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 741, 1969 (Schuster 1969b). *** Schljakovia Konstant. et Vilnet, Arctoa 18: 66, 2009 [2010] (Konstantinova and Vilnet 2009). 30 Plicanthus birmensis seems to be closely related to Tetralophozia (Vilnet et al. 2010), but Grolle (1995) considered it a possibly depauperate form of Plicanthus hirtellus. However, it may be a good species, possibly of the genus Tetralophozia. 31 Plicanthus giganteus is often treated as a subspecies of Plicanthus hirtellus (e.g. Pócs and Lye 1999). 32 Schizophyllopsis was originally published as Schizophyllum (R.M.Schust.) Váňa et L.Söderstr. (Váňa et al. 2013k), but that is a later homonym of Schizophyllum Fr. (1815; Fungus) and Nutt. (1841; Asteraceae). World checklist of hornworts and liverworts 55 *** Schljakovia kunzeana (Huebener) Konstant. et Vilnet, Arctoa 18: 66, 2009 [2010] (Konstantinova and Vilnet 2009). Bas.: Jungermannia kunzeana Huebener, Hepaticol. germ.: 115, 1834 (Hübener 1834). *** Schljakovianthus Konstant. et Vilnet, Arctoa 18: 66, 2009 [2010] (Konstantinova and Vilnet 2009). *** Schljakovianthus quadrilobus (Lindb.) Konstant. et Vilnet, Arctoa 18: 66, 2009 [2010] (Konstantinova and Vilnet 2009). Bas.: Jungermannia quadriloba Lindb., Kongl. Svenska Vetensk.-Akad. Handl. (n.ser.) 23 (5): 55, 1889 (Lindberg and Arnell 1889). *** Sphenolobopsis R.M.Schust. et N.Kitag., Nova Hedwigia 22: 152, 1971 [1972] (Schuster 1971b). *** Sphenolobopsis pearsonii (Spruce) R.M.Schust., Nova Hedwigia 22: 153, 1971 [1972] (Schuster 1971b). Bas.: Jungermannia pearsonii Spruce, J. Bot. 19: 33, 1881 (Spruce 1881b). *** Sphenolobus (Lindb.) Berggr., New Zealand Hepat.: 22, 1898 (Berggren 1898). Bas.: Jungermannia sect. Sphenolobus Lindb., Not. Sällsk. Fauna Fl. Fenn. Förh. 13: 369, 1874 (Lindberg 1874a). *** Sphenolobus austroamericanus (Váňa) Váňa, Phytotaxa 81 (1): 30, 2013 (Váňa et al. 2013a). Bas.: Anastrophyllum austroamericanum Váňa, J. Hattori Bot. Lab. 48: 225, 1980 (Váňa 1980). *** Sphenolobus minutus (Schreb. ex D.Crantz) Berggr., New Zealand Hepat.: 22, 1898 (Berggren 1898). Bas.: Jungermannia minuta Schreb. ex D.Crantz, Forts. Hist. Grönland: 285, 1770 (Crantz 1770; non vidi). *** Sphenolobus saxicola (Schrad.) Steph., Bull. Herb. Boissier (sér. 2) 2 (2): 168 (160), 1902 (Stephani 1902d). Bas.: Jungermannia saxicola Schrad., Syst. Samml. Crypt. Gew. 2: 4, 1797 (Schrader 1797). ** Tetralophozia (R.M.Schust.) Schljakov, Novosti Sist. Nizš. Rast. 13: 227, 1976 (Shliakov 1976). Bas.: Chandonanthus subg. Tetralophozia R.M.Schust., J. Hattori Bot. Lab. 23: 206, 1960 [1961] (Schuster 1960a). *** Tetralophozia cavallii (Gola) Váňa, Trop. Bryol. 8: 102, 1993 (Váňa 1993). Bas.: Blepharostomum cavallii Gola, Ann. Bot. (Rome) 6 (2): 274, 1907 (Gola 1907). *** Tetralophozia iliformis (Steph.) Urmi, J. Bryol. 12 (3): 394, 1983 (Urmi 1983). Bas.: Chandonanthus iliformis Steph., Sp. Hepat. (Stephani) 3: 644, 1909 (Stephani 1909a). ** Tetralophozia pilifera (Steph.) R.M.Schust., Nova Hedwigia 74 (3/4): 482, 2002 (Schuster 2002a). Bas.: Chandonanthus pilifer Steph., Sp. Hepat. (Stephani) 3: 644, 1909 (Stephani 1909a). *** Tetralophozia setiformis (Ehrh.) Schljakov, Novosti Sist. Nizš. Rast. 13: 228, 1976 (Shliakov 1976). Bas.: Jungermannia setiformis Ehrh., Hannover. Mag. 22 (8): 142, 1784 (Ehrhart 1784). 56 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Zantenia (S.Hatt.) Váňa et J.J.Engel, Mem. New York Bot. Gard. 105: 29, 2013 (Váňa and Engel 2013). Bas.: Anastrophyllum subg. Zantenia S.Hatt., Bot. Mag. (Tokyo) 79 (937): 342, 1966 (Hattori 1966a). *** Zantenia borneensis (Herzog) Váňa et J.J.Engel, Mem. New York Bot. Gard. 105: 29, 2013 (Váňa and Engel 2013). Bas.: Anastrophyllum borneense Herzog, Trans. Brit. Bryol. Soc. 1 (4): 282, 1950 (Herzog 1950a). *** Zantenia denticulata (Grolle) Váňa et J.J.Engel, Mem. New York Bot. Gard. 105: 30, 2013 (Váňa and Engel 2013). Bas.: Anastrophyllum denticulatum Grolle, Nova Hedwigia 16: 148, 1968 (Grolle 1968d). *** Zantenia karstenii (Schifn.) Váňa et J.J.Engel, Mem. New York Bot. Gard. 105: 30, 2013 (Váňa and Engel 2013). Bas.: Anastrophyllum karstenii Schifn., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 60 (2): 268, 1893 (Schifner 1893a). *** Zantenia prionophylla (S.Hatt.) Váňa et J.J.Engel, Mem. New York Bot. Gard. 105: 30, 2013 (Váňa and Engel 2013). Bas.: Anastrophyllum prionophyllum S.Hatt., Bot. Mag. (Tokyo) 79 (937): 342, 1966 (Hattori 1966a). *** Cephaloziaceae Mig. by J. Váňa with contributions by S.R. Gradstein (Odontoschisma) Cephaloziaceae was recently studied by Vilnet et al. (2012) and Feldberg et al. (2013). Some taxonomic and nomenclatural notes were published by Váňa et al. (2013g, 2013i) and Gradstein et al. (2014b). ** Alobielloideae R.M.Schust. ** Alobiella (Spruce) Schifn., Hepat. (Engl.-Prantl): 98, 1893 (Schifner 1893b). Bas.: Cephalozia subg. Alobiella Spruce, Cephalozia: 28, 1882 (Spruce 1882). *** Alobiella husnotii (Spruce) Schifn., Hepat. (Engl.-Prantl): 98, 1893 (Schifner 1893b). Bas.: Cephalozia husnotii Spruce, Cephalozia: 30, 1882 (Spruce 1882). ** Alobiellopsis R.M.Schust., Nova Hedwigia 10 (1/2): 25, 1965 (Schuster 1965b). *** Alobiellopsis acroscypha (Spruce) R.M.Schust., Nova Hedwigia 10 (1/2): 25, 1965 (Schuster 1965b). Bas.: Cephalozia acroscypha Spruce, Cephalozia: 30, 1882 (Spruce 1882). *** Alobiellopsis dominicensis (Spruce) Fulford, Mem. New York Bot. Gard. 11 (3): 350, 1968 (Fulford 1968). Bas.: Alobiella dominicensis Spruce, J. Linn. Soc., Bot. 30 (210): 355, 1895 (Gepp 1895b). *** Alobiellopsis heteromorpha (Lehm.) R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 12 (3): 682, 1969 (Schuster 1969a). Bas.: Jungermannia heteromorpha Lehm., Linnaea 4: 362, 1829 (Lehmann 1829). World checklist of hornworts and liverworts 57 *** Alobiellopsis parvifolia (Steph.) R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 12 (3): 679, 1969 (Schuster 1969a). Bas.: Alobiella parvifolia Steph., Bull. Herb. Boissier (sér. 2) 8 (8): 568 (352), 1908 (Stephani 1908e). *** Alobiellopsis pillansii (Sim) R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 12 (3): 683, 1969 (Schuster 1969a). Bas.: Cephalozia pillansii Sim, Trans. Roy. Soc. South Africa 15 (1): 87, 1926 (Sim 1926). ** Cephalozioideae Müll.Frib. *** Cephalozia (Dumort.) Dumort., Recueil Observ. Jungerm.: 18, 1835 (Dumortier 1835). Bas.: Jungermannia sect. Cephalozia Dumort., Syll. Jungerm. Europ.: 60, 1831 (Dumortier 1831). *** Cephalozia acuminata (Herzog) Váňa, Phytotaxa 112 (1): 11, 2013 (Váňa et al. 2013g). Bas.: Hygrobiella acuminata Herzog, Trans. Brit. Bryol. Soc. 1 (4): 292, 1950 (Herzog 1950a). ** Cephalozia acutiloba (Inoue) Váňa, Phytotaxa 112 (1): 11, 2013 (Váňa et al. 2013g). Bas.: Metahygrobiella acutiloba Inoue, Bull. Natl. Sci. Mus. Tokyo (n.ser.) 10 (2): 155, 1967 (Inoue 1967c). *** Cephalozia albula Steph., Hedwigia 32 (5): 318, 1893 (Stephani 1893d). Nom. nov. pro Jungermannia albula Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 93, 1860 [1861] (Mitten 1860c), nom. illeg. *** Cephalozia ambigua C.Massal., Malpighia 21 (7/8): 310, 1907 (Massalongo 1907). *** Cephalozia austrigena R.M.Schust. ex J.J.Engel, Novon 17 (3): 312, 2007 (Engel 2007). Based on: Cephalozia bicuspidata subsp. austrigena R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 712, 1974 (Schuster 1974), nom. inval. *** Cephalozia badia (Gottsche) Steph., Bull. Herb. Boissier (sér. 2) 8 (7): 483 (313), 1908 (Stephani 1908f). Bas.: Jungermannia badia Gottsche, Int. Polarforsch., Deutsch. Exped. 2: 452, 1890 (Gottsche 1890). *** Cephalozia bicuspidata (L.) Dumort., Recueil Observ. Jungerm.: 18, 1835 (Dumortier 1835). Bas.: Jungermannia bicuspidata L., Sp. Pl. 1: 1132, 1753 (Linnaeus 1753). * Cephalozia bicuspidata subsp. lammersiana (Huebener) R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 730, 1974 (Schuster 1974). Bas.: Jungermannia lammersiana Huebener, Flora 15 (20): 306, 1832 (Hübener 1832). *** Cephalozia chilensis (J.J.Engel et R.M.Schust.) R.M.Schust., Beih. Nova Hedwigia 119: 29, 2002 (Schuster 2002b). Bas.: Metahygrobiella chilensis J.J.Engel et R.M.Schust., Brittonia 40 (2): 203, 1988 (Engel and Schuster 1988). *** Cephalozia conchata (Grolle et Váňa) Váňa, Syst. Bot. 40 (1): 38, 2015 (Shaw et al. 2015). Bas.: Jungermannia conchata Grolle et Váňa, Fragm. Florist. Geobot. 37 (1): 3, 1992 (Grolle and Váňa 1992). *** Cephalozia crossii Spruce, Cephalozia: 46, 1882 (Spruce 1882). *** Cephalozia darjeelingensis Udar et D.Kumar, Geophytology 6 (1): 36, 1976 (Udar and Kumar 1976). 58 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Cephalozia drucei (R.M.Schust.) Váňa, Phytotaxa 112 (1): 11, 2013 (Váňa et al. 2013g). Bas.: Metahygrobiella drucei R.M.Schust., J. Hattori Bot. Lab. 26: 273, 1963 (Schuster 1963b). *** Cephalozia fuegiensis Váňa, Phytotaxa 112 (1): 12, 2013 (Váňa et al. 2013g). Nom. nov. pro Hygrobiella dusenii Steph., Sp. Hepat. (Stephani) 6: 444, 1924 (Stephani 1924). *** Cephalozia hamatiloba Steph., Bull. Herb. Boissier (sér. 2) 8 (6): 427 (303), 1908 (Stephani 1908g). *** Cephalozia hamatiloba subsp. siamensis (N.Kitag.) Váňa, Acta Bot. Fenn. 177: 16, 2004 (Koponen et al. 2004). Bas.: Cephalozia siamensis N.Kitag., J. Hattori Bot. Lab. 32: 293, 1969 (Kitagawa 1969c). *** Cephalozia lacinulata (J.B.Jack ex Gottsche et Rabenh.) Spruce, Cephalozia: 45, 1882 (Spruce 1882). Bas.: Jungermannia lacinulata J.B.Jack ex Gottsche et Rabenh., Hepat. Eur., Leberm. 62-64: no. 624, 1877 (Gottsche and Rabenhorst 1877). *** Cephalozia lucens (A.Evans) Steph., Bull. Herb. Boissier (sér. 2) 8 (7): 496 (326), 1908 (Stephani 1908f). Bas.: Jungermannia lucens A.Evans, Trans. Connecticut Acad. Arts 8 (15): 258, 1891 (Evans 1891). *** Cephalozia macgregorii (Steph.) Váňa, Phytotaxa 112 (1): 12, 2013 (Váňa et al. 2013g). Bas.: Hygrobiella macgregorii Steph., Hedwigia 34 (2): 45, 1895 (Stephani 1895c). *** Cephalozia macounii (Austin) Austin, Hepat. bor.-amer.: 14, 1873 (Austin 1873). Bas.: Jungermannia macounii Austin, Proc. Acad. Nat. Sci. Philadelphia 21: 222, 1869 (Austin 1869). *** Cephalozia maxima Steph., Sp. Hepat. (Stephani) 6: 441, 1924 (Stephani 1924). *** Cephalozia mollusca (De Not.) Váňa, Phytotaxa 112 (1): 12, 2013 (Váňa et al. 2013g). Bas.: Jungermannia mollusca De Not., Epat. Borneo: 16, 1874 (De Notaris 1874). * Cephalozia neesiana Steph., Bull. Herb. Boissier (sér. 2) 8 (6): 429 (305), 1908 (Stephani 1908g). 33 *** Cephalozia nishimurae (N.Kitag.) Váňa, Phytotaxa 112 (1): 12, 2013 (Váňa et al. 2013g). Bas.: Hygrobiella nishimurae N.Kitag., Misc. Bryol. Lichenol. 9 (4): 69, 1982 (Kitagawa 1982). ** Cephalozia pachygyna R.M.Schust. ex J.J.Engel, Novon 17 (3): 313, 2007 (Engel 2007). ** Cephalozia physocaula (Hook.f. et Taylor) Steph., Bull. Herb. Boissier (sér. 2) 8 (7): 485 (315), 1908 (Stephani 1908f). Bas.: Jungermannia physocaula Hook.f. et Taylor, London J. Bot. 3: 455, 1844 (Hooker and Taylor 1844b). ** Cephalozia schusteriana J.J.Engel, Novon 17 (3): 313, 2007 (Engel 2007). *** Cephalozia stolonacea (Herzog) Váňa, Phytotaxa 112 (1): 12, 2013 (Váňa et al. 2013g). Bas.: Hygrobiella stolonacea Herzog, Trans. Brit. Bryol. Soc. 1 (4): 293, 1950 (Herzog 1950a). *** Cephalozia tubulata (Hook.f. et Taylor) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 417, 1877 (Trevisan 1877). Bas.: Jungermannia tubulata Hook.f. et Taylor, London J. Bot. 3: 463, 1844 (Hooker and Taylor 1844b). 33 Cephalozia neesiana may be conspeciic with Cephalozia hamatiloba (Váňa 1988). World checklist of hornworts and liverworts * * * * * 59 Excluded from the genus Cephalozia hians Steph., Sp. Hepat. (Stephani) 6: 441, 1924 (Stephani 1924). 34 Cephalozia indica Udar et D.Kumar, Geophytology 6 (1): 42, 1976 (Udar and Kumar 1976). 35 Cephalozia kodaikanalensis G.Asthana et Saumya Srivast., Geophytology 43 (1): 63, 2013 (Asthana and Srivastava 2013). 36 Cephalozia parvifolia Arnell, Rev. Bryol. 25 (1): 1, 1898 (Arnell 1898). 37 Cephalozia tricuspidata (Nees) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 417, 1877 (Trevisan 1877). Bas.: Jungermannia tricuspidata Nees, Enum. Pl. Crypt. Javae: 31, 1830 (Nees 1830). 38 *** Fuscocephaloziopsis Fulford, Mem. New York Bot. Gard. 11 (3): 353, 1968 (Fulford 1968). ** Fuscocephaloziopsis ainis (Lindb. ex Steph.) Váňa et L.Söderstr., Phytotaxa 112 (1): 9, 2013 (Váňa et al. 2013g). Bas.: Cephalozia ainis Lindb. ex Steph., Bull. Herb. Boissier (sér. 2) 8 (4): 277 (291), 1908 (Stephani 1908j). *** Fuscocephaloziopsis africana (Váňa) Váňa et L.Söderstr., Phytotaxa 112 (1): 9, 2013 (Váňa et al. 2013g). Bas.: Cephalozia africana Váňa, Beih. Nova Hedwigia 90: 187, 1988 (Váňa 1988). *** Fuscocephaloziopsis albescens (Hook.) Váňa et L.Söderstr., Phytotaxa 112 (1): 9, 2013 (Váňa et al. 2013g). Bas.: Jungermannia albescens Hook., Brit. Jungermann.: tab. 72, 1815 (Hooker 1815). * Fuscocephaloziopsis albescens var. islandica (Nees) Váňa et L.Söderstr., Phytotaxa 112 (1): 9, 2013 (Váňa et al. 2013g). Bas.: Jungermannia islandica Nees, Naturgesch. Eur. Leberm. 2: 29, 1836 (Nees 1836). *** Fuscocephaloziopsis baldwinii (C.M.Cooke) Váňa et L.Söderstr., Phytotaxa 112 (1): 9, 2013 (Váňa et al. 2013g). Bas.: Cephalozia baldwinii C.M.Cooke, Trans. Connecticut Acad. Arts 12 (1): 35, 1904 (Cooke 1904). ** Fuscocephaloziopsis biloba (Herzog) Fulford, Mem. New York Bot. Gard. 11 (3): 355, 1968 (Fulford 1968). Bas.: Alobiella biloba Herzog, Feddes Repert. Spec. Nov. Regni Veg. 57 (1/2): 166, 1955 (Herzog 1955). *** Fuscocephaloziopsis catenulata (Huebener) Váňa et L.Söderstr., Phytotaxa 112 (1): 9, 2013 (Váňa et al. 2013g). Bas.: Jungermannia catenulata Huebener, Hepaticol. germ.: 169, 1834 (Hübener 1834). *** Fuscocephaloziopsis catenulata subsp. nipponica (S.Hatt.) Váňa et L.Söderstr., Phytotaxa 112 (1): 9, 2013 (Váňa et al. 2013g). Bas.: Cephalozia nipponica S.Hatt., Bull. Tokyo Sci. Mus. 11: 74, 1944 (Hattori 1944d). 34 35 36 37 38 Cephalozia hians is probably an Isotachis or Lophocolea species. Cephalozia indica is a Lophocolea species and probably conspeciic with “Cephalozia” kodaikanalensis. Cephalozia kodaikanalensis is a Lophocolea species, possibly conspeciic with “Cephalozia” indica. Cephalozia parvifolia is a Cephaloziella species although the type specimen has not been found. Cephalozia tricuspidata is a Lepidozia species (Schifner 1898b). 60 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Fuscocephaloziopsis connivens (Dicks.) Váňa et L.Söderstr., Phytotaxa 112 (1): 9, 2013 (Váňa et al. 2013g). Bas.: Jungermannia connivens Dicks., Fasc. Pl. Crypt. Brit. 4: 19, 1801 (Dickson 1801). *** Fuscocephaloziopsis connivens subsp. issa (Steph.) Váňa et L.Söderstr., Phytotaxa 112 (1): 9, 2013 (Váňa et al. 2013g). Bas.: Cephalozia issa Steph., Hedwigia 30 (5): 204, 1891 (Stephani 1891a). *** Fuscocephaloziopsis connivens subsp. sandwicensis (Mont.) Váňa et L.Söderstr., Phytotaxa 112 (1): 9, 2013 (Váňa et al. 2013g). Bas.: Jungermannia sandwicensis Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 249, 1843 (Montagne 1843). *** Fuscocephaloziopsis crassifolia (Lindenb. et Gottsche) Váňa et L.Söderstr., Phytotaxa 112 (1): 10, 2013 (Váňa et al. 2013g). Bas.: Jungermannia crassifolia Lindenb. et Gottsche, Syn. Hepat. 5: 685, 1847 (Gottsche et al. 1847). *** Fuscocephaloziopsis gollanii (Steph.) Váňa et L.Söderstr., Phytotaxa 112 (1): 10, 2013 (Váňa et al. 2013g). Bas.: Cephalozia gollanii Steph., Bull. Herb. Boissier (sér. 2) 8 (6): 428 (304), 1908 (Stephani 1908g). *** Fuscocephaloziopsis leucantha (Spruce) Váňa et L.Söderstr., Phytotaxa 112 (1): 10, 2013 (Váňa et al. 2013g). Bas.: Cephalozia leucantha Spruce, Cephalozia: 68, 1882 (Spruce 1882). *** Fuscocephaloziopsis loitlesbergeri (Schifn.) Váňa et L.Söderstr., Phytotaxa 112 (1): 10, 2013 (Váňa et al. 2013g). Bas.: Cephalozia loitlesbergeri Schifn., Österr. Bot. Z. 62 (1): 10, 1912 (Schifner 1912b). *** Fuscocephaloziopsis lunulifolia (Dumort.) Váňa et L.Söderstr., Phytotaxa 112 (1): 10, 2013 (Váňa et al. 2013g). Bas.: Jungermannia lunulifolia Dumort., Syll. Jungerm. Europ.: 61, 1831 (Dumortier 1831). 39 *** Fuscocephaloziopsis macrostachya (Kaal.) Váňa et L.Söderstr., Phytotaxa 112 (1): 10, 2013 (Váňa et al. 2013g). Bas.: Cephalozia macrostachya Kaal., Rev. Bryol. 29 (1): 8, 1902 (Kaalaas 1902). *** Fuscocephaloziopsis macrostachya subsp. australis (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 112 (1): 10, 2013 (Váňa et al. 2013g). Bas.: Cephalozia macrostachya subsp. australis R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 754, 1974 (Schuster 1974). *** Fuscocephaloziopsis macrostachya subsp. macrostachya var. spinilora (Schifn.) Váňa et L.Söderstr., Phytotaxa 112 (1): 10, 2013 (Váňa et al. 2013g). Bas.: Cephalozia spinilora Schifn., Hedwigia 54 (6): 323, 1914 (Schifner 1914a). *** Fuscocephaloziopsis monticola (J.D.Godfrey) Váňa et L.Söderstr., Phytotaxa 112 (1): 11, 2013 (Váňa et al. 2013g). Bas.: Schoieldia monticola J.D.Godfrey, Bryologist 79 (3): 315, 1976 (Godfrey 1976). *** Fuscocephaloziopsis pachycaulis (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 112 (1): 11, 2013 (Váňa et al. 2013g). Bas.: Cephalozia pachycaulis R.M.Schust., Bryologist 96 (4): 623, 1993 (Schuster 1993b). 39 Fuscocephaloziopsis lunulifolia is a species complex also including Fuscocephaloziopsis ainis and Fuscocephaloziopsis schusteri. World checklist of hornworts and liverworts 61 *** Fuscocephaloziopsis pleniceps (Austin) Váňa et L.Söderstr., Phytotaxa 112 (1): 11, 2013 (Váňa et al. 2013g). Bas.: Jungermannia pleniceps Austin, Proc. Acad. Nat. Sci. Philadelphia 21: 222, 1869 (Austin 1869). ** Fuscocephaloziopsis pleniceps var. caroliniana (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 112 (1): 11, 2013 (Váňa et al. 2013g). Bas.: Cephalozia pleniceps var. caroliniana R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 780, 1974 (Schuster 1974). *** Fuscocephaloziopsis pulvinata (Steph.) Fulford, Mem. New York Bot. Gard. 11 (3): 355, 1968 (Fulford 1968). Bas.: Alobiella pulvinata Steph., Bull. Herb. Boissier (sér. 2) 8 (8): 572 (356), 1908 (Stephani 1908e). * Fuscocephaloziopsis schusteri (Sushil K.Singh et D.K.Singh) Váňa, Phytotaxa 183 (4): 291, 2014 (Váňa et al. 2014a). Bas.: Cephalozia schusteri Sushil K.Singh et D.K.Singh, Lindbergia 32 (1): 1, 2007 (Singh and Singh 2007a). *** Fuscocephaloziopsis subintegra Gradst. et Váňa, Cryptog. Bryol. 25 (3): 274, 2004 (Parolly et al. 2004). *** Fuscocephaloziopsis zoopsioides (Horik.) Váňa et L.Söderstr., Phytotaxa 112 (1): 11, 2013 (Váňa et al. 2013g). Bas.: Cephalozia zoopsioides Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 178, 1934 (Horikawa 1934). *** Nowellia Mitt., Nat. hist. Azores: 321, 1870 (Mitten 1870). ** sect. Acronowellia R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 816, 1974 (Schuster 1974). *** Nowellia reedii H.Rob., Bryologist 73 (1): 150, 1970 (Robinson 1970). *** Nowellia yunckeri Fulford, Mem. New York Bot. Gard. 11 (3): 329, 1968 (Fulford 1968). ** sect. Metanowellia (Grolle) R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 816, 1974 (Schuster 1974). Bas.: Nowellia subg. Metanowellia Grolle, J. Hattori Bot. Lab. 31: 33, 1968 (Grolle 1968b). *** Nowellia borneensis (De Not.) Schifn., Hepat. (Engl.-Prantl): 98, 1893 (Schifner 1893b). Bas.: Jungermannia curvifolia var. borneensis De Not., Epat. Borneo: 19, 1874 (De Notaris 1874). *** Nowellia dominicensis Steph., Sp. Hepat. (Stephani) 6: 443, 1924 (Stephani 1924). *** Nowellia evansii Grolle, J. Hattori Bot. Lab. 31: 33, 1968 (Grolle 1968b). *** Nowellia langii Pearson, J. Linn. Soc., Bot. 46 (305): 25, 1922 (Pearson 1922b). *** Nowellia pusilla Grolle, J. Hattori Bot. Lab. 31: 37, 1968 (Grolle 1968b). *** Nowellia wrightii (Gottsche ex Spruce) Steph. ex Duss, Enum. musc. Antilles franç., Hép.: 21, 1903 (Duss 1903). Bas.: Cephalozia wrightii Gottsche ex Spruce, J. Linn. Soc., Bot. 30 (210): 354, 1895 (Gepp 1895b). 62 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** sect. Nowellia *** Nowellia aciliata (P.C.Chen et P.C.Wu) Mizut., Hikobia 11: 469, 1994 (Mizutani 1994). Bas.: Nowellia curvifolia var. aciliata P.C.Chen et P.C.Wu, Obs. l. Hwangs.: 6, 1965 (Chen and Wu 1965). *** Nowellia curvifolia (Dicks.) Mitt., Nat. hist. Azores: 321, 1870 (Mitten 1870). Bas.: Jungermannia curvifolia Dicks., Fasc. Pl. Crypt. Brit. 2: 15, 1790 (Dickson 1790). ** Odontoschismatoideae H.Buch ex Grolle *** Odontoschisma (Dumort.) Dumort., Recueil Observ. Jungerm.: 19, 1835 (Dumortier 1835). Bas.: Pleuroschisma sect. Odontoschisma Dumort., Syll. Jungerm. Europ.: 68, 1831 (Dumortier 1831). ** sect. Cladopodiella (H.Buch) Gradst., S.C.Aranda et Vanderp., Taxon 63 (5): 1017, 2014 (Aranda et al. 2014). Bas.: Cladopodiella H.Buch, Memoranda Soc. Fauna Fl. Fennica 1: 89, 1927 (Buch 1927). *** Odontoschisma francisci (Hook.) L.Söderstr. et Váňa, Phytotaxa 112 (1): 12, 2013 (Váňa et al. 2013g). Bas.: Jungermannia francisci Hook., Brit. Jungermann.: tab. 49, 1813 (Hooker 1813). ** sect. Denudata R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 833, 1974 (Schuster 1974). *** Odontoschisma brasiliense Steph., Bull. Herb. Boissier (sér. 2) 8 (8): 585 (369), 1908 (Stephani 1908e). ** Odontoschisma cleeii Gradst., S.C.Aranda et Vanderp., Taxon 63 (5): 1017, 2014 (Aranda et al. 2014). *** Odontoschisma denudatum (Mart.) Dumort., Recueil Observ. Jungerm.: 19, 1835 (Dumortier 1835). Bas.: Jungermannia scalaris var. denudata Mart., Fl. crypt. erlang.: 183, 1817 (Martius 1817). ** Odontoschisma denudatum subsp. naviculare (Steph.) Gradst., S.C.Aranda et Vanderp., Taxon 63 (5): 1019, 2014 (Aranda et al. 2014). Bas.: Jamesoniella navicularis Steph., Sp. Hepat. (Stephani) 6: 101, 1917 (Stephani 1917a). ** Odontoschisma denudatum subsp. sandvicense (Ångstr.) Gradst., S.C.Aranda et Vanderp., Taxon 63 (5): 1019, 2014 (Aranda et al. 2014). Bas.: Sphagnoecetis sandvicensis Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 29 (4): 22, 1872 (Ångström 1872). *** Odontoschisma elongatum (Lindb.) A.Evans, Rhodora 14 (157): 13, 1912 (Evans 1912b). Bas.: Odontoschisma denudatum f. elongatum Lindb., Helsingf. Dagbl. 1874 (45, 16 Feb): 2, 1874 (Lindberg 1874b). *** Odontoschisma engelii Gradst. et Burghardt, Fieldiana, Bot. (n.ser.) 47: 194, 2008 (Gradstein and Burghardt 2008). World checklist of hornworts and liverworts 63 *** Odontoschisma longilorum (Taylor) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 419, 1877 (Trevisan 1877). Bas.: Sphagnoecetis longilora Taylor, London J. Bot. 5: 281, 1846 (Taylor 1846a). *** Odontoschisma macounii (Austin) Underw., Bull. Illinois State Lab. Nat. Hist. 2 (1): 92, 1884 (Underwood 1884). Bas.: Sphagnoecetis macounii Austin, Bull. Torrey Bot. Club 3 (3): 13, 1872 (Austin 1872). *** Odontoschisma portoricense (Hampe et Gottsche) Steph., Hedwigia 27 (11/12): 296, 1888 (Stephani 1888c). Bas.: Sphagnoecetis portoricensis Hampe et Gottsche, Linnaea 25 (3): 343, 1852 [1853] (Hampe and Gottsche 1852). ** Odontoschisma pseudogrosseverrucosum Gradst., S.C.Aranda et Vanderp., Taxon 63 (5): 1019, 2014 (Aranda et al. 2014). *** Odontoschisma purpuratum Herzog, Trans. Brit. Bryol. Soc. 1 (4): 297, 1950 (Herzog 1950a). *** Odontoschisma soratamum Fulford, Mem. New York Bot. Gard. 11 (3): 338, 1968 (Fulford 1968). *** Odontoschisma variabile (Lindenb. et Gottsche) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 419, 1877 (Trevisan 1877). Bas.: Sphagnoecetis variabilis Lindenb. et Gottsche, Syn. Hepat. 5: 688, 1847 (Gottsche et al. 1847). ** Odontoschisma zhui Gradst., S.C.Aranda et Vanderp., Taxon 63 (5): 1020, 2014 (Aranda et al. 2014). ** sect. Iwatsukia (N.Kitag.) Gradst., S.C.Aranda et Vanderp., Phytotaxa 162 (4): 232, 2014 (Gradstein et al. 2014b). Bas.: Iwatsukia N.Kitag., J. Hattori Bot. Lab. 27: 178, 1964 (Kitagawa 1964). *** Odontoschisma biidum (Fulford) Gradst., S.C.Aranda et Vanderp., Phytotaxa 162 (4): 232, 2014 (Gradstein et al. 2014b). Bas.: Cladomastigum biidum Fulford, Acta Bot. Venez. 2 (5/8): 80, 1967 (Fulford 1967). *** Odontoschisma jishibae (Steph.) L.Söderstr. et Váňa, Phytotaxa 112 (1): 13, 2013 (Váňa et al. 2013g). Bas.: Cephalozia jishibae Steph., Sp. Hepat. (Stephani) 6: 437, 1924 (Stephani 1924). *** Odontoschisma spinosum (Fulford) Gradst., S.C.Aranda et Vanderp., Phytotaxa 162 (4): 232, 2014 (Gradstein et al. 2014b). Bas.: Cladomastigum spinosum Fulford, Mem. New York Bot. Gard. 23: 840, 1972 (Fulford 1972). ** sect. Neesia Gradst., S.C.Aranda et Vanderp., Taxon 63 (5): 1017, 2014 (Aranda et al. 2014). *** Odontoschisma luitans (Nees) L.Söderstr. et Váňa, Phytotaxa 112 (1): 12, 2013 (Váňa et al. 2013g). Bas.: Jungermannia luitans Nees, Flora 6 (2): 30, 1823 (Link 1823). ** sect. Odontoschisma *** Odontoschisma grosseverrucosum Steph., Bull. Herb. Boissier (sér. 2) 8 (8): 593 (377), 1908 (Stephani 1908e). 64 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Odontoschisma sphagni (Dicks.) Dumort., Recueil Observ. Jungerm.: 19, 1835 (Dumortier 1835). Bas.: Jungermannia sphagni Dicks., Fasc. Pl. Crypt. Brit. 1: 6, 1785 (Dickson 1785). ** Odontoschisma steyermarkii Gradst. et Ilk.-Borg., Nova Hedwigia 100 (1/2): 38, 2015 [2014 online] (Gradstein and Ilkiu-Borges 2015). * ** Incertae sedis Odontoschisma obcordatum (Spruce) Steph., Bull. Herb. Boissier (sér. 2) 8 (8): 586 (370), 1908 (Stephani 1908e). Bas.: Cephalozia obcordata Spruce, Cephalozia: 61, 1882 (Spruce 1882). 40 Schiffnerioideae R.M.Schust. ** Schifneria Steph., Österr. Bot. Z. 44 (1): 1, 1894 (Stephani 1894c). *** Schifneria hyalina Steph., Österr. Bot. Z. 44 (1): 1, 1894 (Stephani 1894c). ** Trabacelluloideae R.M.Schust. ** Haesselia Grolle et Gradst., J. Hattori Bot. Lab. 64: 327, 1988 (Grolle and Gradstein 1988). *** Haesselia acuminata Gradst., Trop. Bryol. 1: 30, 1989 (Gradstein and Florschützde Waard 1989). *** Haesselia roraimensis Grolle et Gradst., J. Hattori Bot. Lab. 64: 327, 1988 (Grolle and Gradstein 1988). ** Trabacellula Fulford, Acta Bot. Venez. 2 (5/8): 86, 1967 (Fulford 1967). *** Trabacellula tumidula Fulford, Acta Bot. Venez. 2 (5/8): 86, 1967 (Fulford 1967). *** Cephaloziellaceae Douin by J. Váňa he treatment of Cephaloziellaceae follows what was outlined in Váňa et al. (2013e). Some nomenclatural and taxonomic notes can also be found in Söderström et al. (2013a) and Váňa et al. (2013b, 2013l). he placement of Phycolepidozia follows Gradstein et al. (2014a). 40 Odontoschisma obcordatum is to be excluded from Odontoschisma but its identity remains unclear as the type is not found. Based on the original description this species might belong to Alobiellopsis (Gradstein & Ilkiu-Borges 2014: 81). World checklist of hornworts and liverworts 65 *** Allisoniella E.A.Hodgs., Trans. Roy. Soc. New Zealand, Bot. 3 (4): 80, 1965 (Hodgson 1965). ** sect. Allisoniella *** Allisoniella nigra (Rodway) R.M.Schust., Nova Hedwigia 22: 137, 1971 [1972] (Schuster 1971b). Bas.: Sphenolobus niger Rodway, Tasm. Bryoph.: 33, 1917 (Rodway 1917b). ** Allisoniella nigra var. acutiloba J.J.Engel, Novon 17 (3): 313, 2007 (Engel 2007). ** Allisoniella nigra subsp. novaezelandiae R.M.Schust., Nova Hedwigia 22: 143, 1971 [1972] (Schuster 1971b). *** Allisoniella recurva R.M.Schust., Nova Hedwigia 22: 146, 1971 [1972] (Schuster 1971b). *** Allisoniella subbipartita (C.Massal.) R.M.Schust. et J.J.Engel, Nova Hedwigia 22: 147, 1971 [1972] (Schuster 1971b). Bas.: Cephalozia subbipartita C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 235, 1885 (Massalongo 1885). *** Allisoniella tasmanica R.M.Schust., Nova Hedwigia 22: 145, 1971 [1972] (Schuster 1971b). ** sect. Protomarsupella (R.M.Schust.) R.M.Schust., Nova Hedwigia 22: 150, 1971 (Schuster 1971b). Bas.: Protomarsupella R.M.Schust., Rev. Bryol. Lichénol. 34 (1/2): 264, 1966 (Schuster 1966b). *** Allisoniella scottii (R.M.Schust.) R.M.Schust., Nova Hedwigia 22: 151, 1971 [1972] (Schuster 1971b). Bas.: Protomarsupella scottii R.M.Schust., Rev. Bryol. Lichénol. 34 (1/2): 267, 1966 (Schuster 1966b). *** Amphicephalozia R.M.Schust., Nova Hedwigia 22: 131, 1971 [1972] (Schuster 1971b). *** Amphicephalozia africana Váňa et M.Wigginton, J. Bryol. 30 (1): 55, 2008 (Váňa and Wigginton 2008). *** Amphicephalozia amplexicaulis R.M.Schust., Nova Hedwigia 22: 133, 1971 [1972] (Schuster 1971b). *** Amphicephalozia geisslerae Pócs et Váňa, Polish Bot. J. 46 (2): 145, 2001 (Pócs and Váňa 2001). *** Anastrophyllopsis (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 81 (1): 15, 2013 (Váňa et al. 2013k). Bas.: Anastrophyllum sect. Anastrophyllopsis R.M.Schust., Beih. Nova Hedwigia 119: 310, 2002 (Schuster 2002b). *** Anastrophyllopsis involutifolia (Mont. ex Gottsche, Lindenb. et Nees) Váňa et L.Söderstr., Phytotaxa 81 (1): 15, 2013 (Váňa et al. 2013k). Bas.: Jungermannia involutifolia Mont. ex Gottsche, Lindenb. et Nees, Syn. Hepat. 1: 81, 1844 (Gottsche et al. 1844). *** Anastrophyllopsis revoluta (Steph.) Váňa et L.Söderstr., Phytotaxa 81 (1): 16, 2013 (Váňa et al. 2013k). Bas.: Anastrophyllum revolutum Steph., Hedwigia 32 (3): 139, 1893 (Stephani 1893b). 66 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Anastrophyllopsis subcomplicata (Lehm. et Lindenb.) Váňa et L.Söderstr., Phytotaxa 81 (1): 16, 2013 (Váňa et al. 2013k). Bas.: Jungermannia subcomplicata Lehm. et Lindenb., Nov. Stirp. Pug. 7: 4, 1838 (Lehmann 1838). ** Cephalojonesia Grolle, Rev. Bryol. Lichénol. 37 (4): 763, 1970 [1971] (Grolle and Vanden Berghen 1970). *** Cephalojonesia incuba Grolle et Vanden Berghen, Rev. Bryol. Lichénol. 37 (4): 764, 1970 [1971] (Grolle and Vanden Berghen 1970). *** Cephalojonesia incuba subsp. mexicana Burghardt, Gradst. et Váňa, J. Hattori Bot. Lab. 100: 35, 2006 (Burghardt et al. 2006). ** Cephalomitrion R.M.Schust., Nova Hedwigia 61 (3/4): 550, 1995 (Schuster 1995b). *** Cephalomitrion aterrimum (Steph.) R.M.Schust., Nova Hedwigia 61 (3/4): 554, 1995 (Schuster 1995b). Bas.: Cephalozia aterrima Steph., Bull. Herb. Boissier (sér. 2) 8 (7): 501 (331), 1908 (Stephani 1908f). *** Cephaloziella (Spruce) Schifn., Hepat. (Engl.-Prantl): 98, 1893 (Schifner 1893b) nom. conserv. Bas.: Cephalozia subg. Cephaloziella Spruce, Cephalozia: 62, 1882 (Spruce 1882). 41 ** subg. Cephaloziella *** Cephaloziella aenigmatica R.M.Schust., Nova Hedwigia 63 (1/2): 20, 1996 (Schuster 1996d). *** Cephaloziella anthelioides S.W.Arnell, Bot. Not. 105: 322, 1952 (Arnell 1952a). *** Cephaloziella arctogena (R.M.Schust.) Konstant., Arctoa 3: 126, 1994 (Konstantinova and Vasil’ev 1994). Bas.: Cephaloziella rubella var. arctogena R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 125, 1980 (Schuster 1980c). ** Cephaloziella arenaria (Steph.) R.M.Schust., J. Hattori Bot. Lab. 26: 280, 1963 (Schuster 1963b). Bas.: Cephalozia arenaria Steph., Bull. Herb. Boissier (sér. 2) 8 (7): 504 (334), 1908 (Stephani 1908f). *** Cephaloziella aspericaulis Jørg., Bergens Mus. Skr. (n.ser.) 16: 197, 1934 (Jørgensen 1934). *** Cephaloziella baumgartneri Schifn., Verh. K.K. Zool.-Bot. Ges. Wien 56 (3): 273, 1906 (Schifner 1906c). ** Cephaloziella breviperianthia C.Gao, Fl. Hepat. Chin. Boreali-Orient.: 208, 1981 (Gao and Chang 1981). ** Cephaloziella brinkmannii Douin, Mém. Soc. Bot. France 29: 75, 1920 (Douin 1920). *** Cephaloziella capensis (Sim) S.W.Arnell, Bot. Not. 105: 326, 1952 (Arnell 1952a). Bas.: Cephalozia capensis Sim, Trans. Roy. Soc. South Africa 15 (1): 87, 1926 (Sim 1926). 41 Cephaloziella is a large genus and the subgeneric division is unclear. he traditional subdivisions used here do not necessary agree with recent phylogenetic studies. World checklist of hornworts and liverworts 67 *** Cephaloziella capillaris (Steph.) Douin, Mém. Soc. Bot. France 29: 59, 1920 (Douin 1920). Bas.: Cephalozia capillaris Steph., Bull. Herb. Boissier (sér. 2) 8 (7): 496 (326), 1908 (Stephani 1908f). *** Cephaloziella crassigyna (R.M.Schust.) R.M.Schust., Nova Hedwigia 61 (3/4): 556, 1995 (Schuster 1995b). Bas.: Cephaloziella aterrima var. crassigyna R.M.Schust., Nova Hedwigia 22: 211, 1971 [1972] (Schuster 1971b). *** Cephaloziella crispata N.Kitag., J. Hattori Bot. Lab. 32: 301, 1969 (Kitagawa 1969c). *** Cephaloziella densifolia R.M.Schust., Nova Hedwigia 22: 199, 1971 [1972] (Schuster 1971b). ** Cephaloziella densifolia var. dubia R.M.Schust., Nova Hedwigia 22: 202, 1971 [1972] (Schuster 1971b). *** Cephaloziella divaricata (Sm.) Schifn., Hepat. (Engl.-Prantl): 99, 1893 (Schifner 1893b). Bas.: Jungermannia divaricata Sm., Engl. Bot. 10: tab. 719, 1800 (Smith and Sowerby 1800). ** Cephaloziella divaricata var. scabra (M.Howe) Haynes, Bryologist 12 (4): 68, 1909 (Haynes 1909). Bas.: Cephalozia divaricata var. scabra M.Howe, Mem. Torrey Bot. Club 7: 129, 1899 (Howe 1899). 42 ** Cephaloziella dusenii Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 6): 49, 1900 (Stephani 1900b). *** Cephaloziella elachista (J.B.Jack ex Gottsche et Rabenh.) Schifn., Sitzungsber. deutsch. naturwiss.-med. Vereins Böhmen “Lotos” Prag 48: 336, 1900 (Schifner 1900d). Bas.: Jungermannia elachista J.B.Jack ex Gottsche et Rabenh., Hepat. Eur., Leberm. 58-59: no. 574, 1873 (Gottsche and Rabenhorst 1873a). ** Cephaloziella elachista var. spinophylla (C.Gao) C.Gao, Fl. Bryoph. Sin. 9: 181, 2003 (Gao 2003). Bas.: Cephaloziella spinophylla C.Gao, Fl. Hepat. Chin. BorealiOrient.: 208, 1981 (Gao and Chang 1981). ** Cephaloziella elegans (Heeg) Schifn., Sitzungsber. deutsch. naturwiss.-med. Vereins Böhmen “Lotos” Prag 48: 336, 1900 (Schifner 1900d). Bas.: Cephalozia elegans Heeg, Rev. Bryol. 20 (5): 82, 1893 (Heeg 1893). *** Cephaloziella exigua R.M.Schust., Nova Hedwigia 63 (1/2): 49, 1996 (Schuster 1996d). *** Cephaloziella exililora (Taylor) Douin, Mém. Soc. Bot. France 29: 72, 1920 (Douin 1920). Bas.: Jungermannia exililora Taylor, London J. Bot. 5: 279, 1846 (Taylor 1846a). ** Cephaloziella fragillima (Spruce) Fulford, Mem. New York Bot. Gard. 11 (4): 409, 1976 (Fulford 1976). Bas.: Cephalozia fragillima Spruce, Mem. Torrey Bot. Club 1 (3): 131, 1890 (Spruce 1890). *** Cephaloziella garsidei S.W.Arnell, Rev. Bryol. Lichénol. 23 (1/2): 173, 1954 (Arnell 1954a). *** Cephaloziella grandiretis (R.M.Schust.) R.M.Schust., Nova Hedwigia 63 (1/2): 35, 1996 (Schuster 1996d). Bas.: Cephaloziella byssacea subsp. grandiretis R.M.Schust., Nova Hedwigia 22: 195, 1971 [1972] (Schuster 1971b). 42 he correct name for the variety was discussed by Söderström et al. (2012d). 68 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Cephaloziella grimsulana (J.B.Jack ex Gottsche et Rabenh.) Lacout., Hépat. France: 52, 1905 (Lacouture 1905). Bas.: Jungermannia grimsulana J.B.Jack ex Gottsche et Rabenh., Hepat. Eur., Leberm. 53–55: no. 526, 1872 (Gottsche and Rabenhorst 1872). *** Cephaloziella hampeana (Nees) Schifn. ex Loeske, Moosl. Harz.: 92, 1903 (Loeske 1903). Bas.: Jungermannia hampeana Nees, Naturgesch. Eur. Leberm. 3: 560, 1838 (Nees 1838b). ** Cephaloziella hebridensis Steph., Hedwigia 32 (5): 316, 1893 (Stephani 1893d). *** Cephaloziella herzogiana (Pandé et K.P.Srivast.) Udar et D.Kumar, Geophytology 6 (1): 45, 1976 (Udar and Kumar 1976). Bas.: Cephalozia herzogiana Pandé et K.P.Srivast., Feddes Repert. Spec. Nov. Regni Veg. 58: 75, 1955 (Pandé and Srivastava 1955). ** Cephaloziella heteroica (C.M.Cooke) Douin, Mém. Soc. Bot. France 29: 76, 1920 (Douin 1920). Bas.: Cephalozia heteroica C.M.Cooke, Trans. Connecticut Acad. Arts 12 (1): 38, 1904 (Cooke 1904). ** Cephaloziella hyalina Douin, Mém. Soc. Bot. France 29: 77, 1920 (Douin 1920). * Cephaloziella hyalina var. rappii (Douin) R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 56, 1980 (Schuster 1980c). Bas.: Cephaloziella rappii Douin, Mém. Soc. Bot. France 29: 77, 1920 (Douin 1920). *** Cephaloziella inaequalis R.M.Schust., Nova Hedwigia 22: 186, 1971 [1972] (Schuster 1971b). *** Cephaloziella invisa R.M.Schust., Nova Hedwigia 63 (1/2): 30, 1996 (Schuster 1996d). ** Cephaloziella kilohanensis (C.M.Cooke) Douin, Mém. Soc. Bot. France 29: 85, 1920 (Douin 1920). Bas.: Cephalozia kilohanensis C.M.Cooke, Trans. Connecticut Acad. Arts 12 (1): 37, 1904 (Cooke 1904). *** Cephaloziella longii Váňa, Folia Geobot. Phytotax. 27 (2): 193, 1992 (Váňa 1992). *** Cephaloziella lycopodioides (Sim) S.W.Arnell, Bot. Not. 105: 321, 1952 (Arnell 1952a). Bas.: Cephalozia lycopodioides Sim, Trans. Roy. Soc. South Africa 15 (1): 85, 1926 (Sim 1926). ** Cephaloziella mammillifera R.M.Schust. et Damsh., Phytologia 63 (5): 327, 1987 (Schuster and Damsholt 1987). *** Cephaloziella massalongi (Spruce) Müll.Frib., Lebermoose 2 (17): 191, 1913 (Müller 1913a). Bas.: Cephalozia massalongi Spruce, Cephalozia: 71, 1882 (Spruce 1882). *** Cephaloziella muelleriana R.M.Schust., Nova Hedwigia 63 (1/2): 24, 1996 (Schuster 1996d). *** Cephaloziella natalensis (Sim) S.W.Arnell, Hepat. South Africa: 338, 1963 (Arnell 1963b). Bas.: Cephalozia natalensis Sim, Trans. Roy. Soc. South Africa 15 (1): 86, 1926 (Sim 1926). ** Cephaloziella nicholsonii Douin, Rev. Bryol. 40 (6): 81, 1913 (Douin 1913b). *** Cephaloziella nothogena R.M.Schust., Nova Hedwigia 63 (1/2): 37, 1996 (Schuster 1996d). ** Cephaloziella obtusilobula R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 108, 1980 (Schuster 1980c). World checklist of hornworts and liverworts 69 ** Cephaloziella patulifolia (Steph.) Douin, Mém. Soc. Bot. France 29: 70, 1920 (Douin 1920). Bas.: Cephalozia patulifolia Steph., Bull. Herb. Boissier (sér. 2) 8 (7): 509 (339), 1908 (Stephani 1908f). *** Cephaloziella phyllacantha (C.Massal. et Carestia) Müll.Frib., Lebermoose 2 (17): 194, 1913 (Müller 1913a). Bas.: Anthelia phyllacantha C.Massal. et Carestia, Nuovo Giorn. Bot. Ital. 12 (4): 340, 1880 (Massalongo and Carestia 1880). *** Cephaloziella polystratosa (R.M.Schust. et Damsh.) Konstant., Bot. Zhurn. (Moscow & Leningrad) 85 (10): 127, 2000 (Konstantinova 2000). Bas.: Cephaloziella byssacea var. polystratosa R.M.Schust. et Damsh., Phytologia 63 (5): 327, 1987 (Schuster and Damsholt 1987). *** Cephaloziella pseudocrassigyna R.M.Schust. ex J.J.Engel, Novon 17 (3): 313, 2007 (Engel 2007). *** Cephaloziella pulcherrima R.M.Schust., Nova Hedwigia 22: 203, 1971 [1972] (Schuster 1971b). ** Cephaloziella pulcherrima subsp. sphagnicola R.M.Schust., Nova Hedwigia 22: 205, 1971 [1972] (Schuster 1971b). ** Cephaloziella pungens Steph. ex Fulford, Mem. New York Bot. Gard. 11 (4): 410, 1976 (Fulford 1976). *** Cephaloziella rubella (Nees) Warnst., Krypt.-Fl. Brandenburg, Leber- & Torfmoose: 231, 1902 (Warnstorf 1902). Bas.: Jungermannia rubella Nees, Naturgesch. Eur. Leberm. 2: 236, 1836 (Nees 1836). *** Cephaloziella schelpei S.W.Arnell, Bot. Not. 110 (1): 18, 1957 (Arnell 1957a). *** Cephaloziella spegazziniana (C.Massal.) Douin, Mém. Soc. Bot. France 29: 69, 1920 (Douin 1920). Bas.: Cephalozia spegazziniana C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 234, 1885 (Massalongo 1885). *** Cephaloziella spinicaulis Douin, Mém. Soc. Bot. France 29: 62, 1920 (Douin 1920). *** Cephaloziella spinigera (Lindb.) Jørg., Bergens Mus. Skr. (n.ser.) 16: 189, 1934 (Jørgensen 1934). Bas.: Cephalozia spinigera Lindb., Musci Scand.: 4, 1879 (Lindberg 1879). *** Cephaloziella stellulifera (Taylor ex Carrington et Pearson) Croz., Rev. Bryol. 30 (2): 31, 1903 (Crozals 1903a). Bas.: Jungermannia stellulifera Taylor ex Carrington et Pearson, Hepat. Brit. Exsicc. Fasc. I: no. 32, 1878 (Carrington and Pearson 1878). *** Cephaloziella stephanii Schifn. ex Douin, Mém. Soc. Bot. France 29: 85, 1920 (Douin 1920). ** Cephaloziella stolonifera R.M.Schust., Phytologia 39 (6): 426, 1978 (Schuster 1978b). ** Cephaloziella subtilis (Lindenb. et Gottsche) Steph., Hedwigia 32 (5): 318, 1893 (Stephani 1893d). Bas.: Jungermannia subtilis Lindenb. et Gottsche, Syn. Hepat. 5: 685, 1847 (Gottsche et al. 1847). *** Cephaloziella sumatrana Schifn. ex Douin, Mém. Soc. Bot. France 29: 59, 1920 (Douin 1920). *** Cephaloziella tabularis S.W.Arnell, Bot. Not. 105: 318, 1952 (Arnell 1952a). *** Cephaloziella transvaalensis S.W.Arnell, Bot. Not. 110 (1): 19, 1957 (Arnell 1957a). *** Cephaloziella triplicata S.W.Arnell, Bot. Not. 115: 203, 1962 (Arnell 1962a). 70 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Cephaloziella umtaliensis S.W.Arnell, Bot. Not. 110 (1): 20, 1957 (Arnell 1957a). *** Cephaloziella uncinata R.M.Schust., Meddel. Grønland 199 (1): 316, 1974 (Schuster and Damsholt 1974). ** Cephaloziella uncinata var. brevigyna R.M.Schust. et Damsh., Phytologia 63 (5): 327, 1987 (Schuster and Damsholt 1987). ** Cephaloziella uncinata var. sphagnicola R.M.Schust., Meddel. Grønland 199 (1): 323, 1974 (Schuster and Damsholt 1974). *** Cephaloziella vaginans Steph., Wiss. Ergebn. Deut. Zentr.-Afr. Exped. (1907-08), Bot. 2: 119, 1911 (Stephani 1911a). *** Cephaloziella varians (Gottsche) Steph., Wiss. Ergebn. Schwed. Südpolar-Exped. [1901–1903] 4 (1): 4, 1905 (Stephani 1905e). Bas.: Jungermannia varians Gottsche, Int. Polarforsch., Deutsch. Exped. 2: 452, 1890 (Gottsche 1890). *** Cephaloziella verrucosa Steph., Hedwigia 32 (5): 318, 1893 (Stephani 1893d). * Cephaloziella villaumei (Steph.) Váňa, Phytotaxa 112 (1): 3, 2013 (Váňa et al. 2013e). Bas.: Cephalozia villaumei Steph., Sp. Hepat. (Stephani) 6: 437, 1924 (Stephani 1924). 43 ** Cephaloziella violacea Schljakov, Novosti Sist. Nizš. Rast. 15: 242, 1978 (Shliakov 1978). ** Cephaloziella welwitschii (Steph.) Douin, Mém. Soc. Bot. France 29: 58, 1920 (Douin 1920). Bas.: Cephalozia welwitschii Steph., Bull. Herb. Boissier (sér. 2) 8 (6): 432 (308), 1908 (Stephani 1908g). ** subg. Dichiton (Mont.) Müll.Frib., Lebermoose 2 (27): 787, 1916 (Müller 1916). Bas.: Dichiton Mont., Syll. Gen. Sp. Crypt.: 52, 1856 (Montagne 1856b). *** Cephaloziella calyculata (Durieu et Mont.) Müll.Frib., Lebermoose 2 (27): 787, 1916 (Müller 1916). Bas.: Jungermannia calyculata Durieu et Mont., Ann. Sci. Nat. Bot. (sér. 3) 11: 34, 1849 (Montagne 1849). *** Cephaloziella integerrima (Lindb.) Warnst., Krypt.-Fl. Brandenburg, Leber- & Torfmoose: 232, 1902 (Warnstorf 1902). Bas.: Cephalozia integerrima Lindb., Acta Soc. Sci. Fenn. 10: 502, 1875 (Lindberg 1875). ** subg. Evansia (Douin et Schifn.) Müll.Frib., Lebermoose 2 (27): 787, 1916 (Müller 1916). Bas.: Evansia Douin et Schifn., Rev. Bryol. 40 (5): 66, 1913 (Douin 1913a). *** Cephaloziella antillana (Besch. et Spruce) Fulford, Mem. New York Bot. Gard. 11 (4): 405, 1976 (Fulford 1976). Bas.: Cephalozia antillana Besch. et Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: clxxxiii, 1889 [1890] (Bescherelle and Spruce 1889). *** Cephaloziella dentata (Raddi) Steph., Bull. Herb. Boissier 5 (2): 78, 1897 (Stephani 1897b). Bas.: Jungermannia dentata Raddi, Jungermanniogr. Etrusca: 21, 1818 (Raddi 1818a). *** Cephaloziella hirta (Steph.) R.M.Schust., J. Hattori Bot. Lab. 26: 280, 1963 (Schuster 1963b). Bas.: Cephalozia hirta Steph., Bull. Herb. Boissier (sér. 2) 8 (8): 561 (345), 1908 (Stephani 1908e). 43 Cephaloziella villaumei is possibly conspeciic with Cephaloziella anthelioides. World checklist of hornworts and liverworts 71 ** Cephaloziella squarrosula (Trevis.) Steph., Hedwigia 32 (5): 318, 1893 (Stephani 1893d). Bas.: Cephalozia squarrosula Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 417, 1877 (Trevisan 1877). *** Cephaloziella subspinosa R.M.Schust., Nova Hedwigia 22: 210, 1971 [1972] (Schuster 1971b). ** subg. Prionolobus (Spruce) Müll.Frib., Lebermoose 2 (16): 110, 1912 (Müller 1912). Bas.: Cephalozia subg. Prionolobus Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 508, 1885 (Spruce 1885). *** Cephaloziella acanthophora (S.Hatt.) Horik., Hikobia 1 (2): 79, 1951 (Horikawa 1951c). Bas.: Prionolobus acanthophorus S.Hatt., Bull. Tokyo Sci. Mus. 11: 29, 1944 (Hattori 1944d). *** Cephaloziella biokoensis Váňa et Frank Müll., Trop. Bryol. 24: 1, 2003 (Váňa and Müller 2003). *** Cephaloziella granatensis (J.B.Jack ex Steph.) Fulford, Mem. New York Bot. Gard. 11 (4): 411, 1976 (Fulford 1976). Bas.: Cephalozia granatensis J.B.Jack ex Steph., Bull. Herb. Boissier (sér. 2) 8 (7): 500 (330), 1908 (Stephani 1908f). *** Cephaloziella grisea R.M.Schust., Phytologia 39 (6): 425, 1978 (Schuster 1978b). * Cephaloziella meghalayensis Udar et Ad.Kumar, Lindbergia 8 (1): 34, 1982 (Udar and Kumar 1982d). 44 *** Cephaloziella microphylla (Steph.) Douin, Mém. Soc. Bot. France 29: 59, 1920 (Douin 1920). Bas.: Cephalozia microphylla Steph., Bull. Herb. Boissier (sér. 2) 8 (7): 513 (343), 1908 (Stephani 1908f). *** Cephaloziella tenuissima (Lehm.) Steph., Hedwigia 32 (5): 318, 1893 (Stephani 1893d). Bas.: Jungermannia tenuissima Lehm., Linnaea 4: 367, 1829 (Lehmann 1829). *** Cephaloziella turneri (Hook.) Müll.Frib., Lebermoose 2 (17): 202, 1913 (Müller 1913a). Bas.: Jungermannia turneri Hook., Brit. Jungermann.: tab. 29, 1812 (Hooker 1812). Incertae sedis ** Cephaloziella dentifolia Udar et Ad.Kumar, Lindbergia 8 (1): 30, 1982 (Udar and Kumar 1982d). ** Cephaloziella ilum (Trevis.) Steph., Hedwigia 32 (5): 318, 1893 (Stephani 1893d). Bas.: Cephalozia ilum Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 417, 1877 (Trevisan 1877). * Cephaloziella lexuosa C.Gao et K.C.Chang, Bull. Bot. Res., Harbin 4 (3): 88, 1984 (Chang and Gao 1984). 45 ** Cephaloziella intricata Schifn. ex Douin, Mém. Soc. Bot. France 29: 59, 1920 (Douin 1920). 44 Cephaloziella meghalayensis is probably conspeciic with Cephaloziella acanthophora or Cephaloziella microphylla. 45 Cephaloziella lexuosa may be a form of Plicanthus birmensis or a juvenile form of Tetralophozia iliformis. 72 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Cephaloziella levieri Schifn. ex Douin, Mém. Soc. Bot. France 29: 80, 1920 (Douin 1920). *** Cephaloziella pellucida R.M.Schust., Nova Hedwigia 63 (1/2): 57, 1996 (Schuster 1996d). 46 * Cephaloziella pygmaea (Spruce) Váňa, Phytotaxa 183 (4): 290, 2014 (Váňa et al. 2014a). Bas.: Cephalozia pygmaea Spruce, Cephalozia: 69, 1882 (Spruce 1882). 47 * Cephaloziella secundifolia Pearson, Ann. Bryol. 4: 106, 1931 (Pearson 1931b). 48 ** Cephaloziella sinensis Douin, Rev. Bryol. 41 (1): 8, 1914 (Douin 1914). ** Cephaloziopsis (Spruce) Schifn., Hepat. (Engl.-Prantl): 85, 1893 (Schifner 1893b). Bas.: Jungermannia sect. Cephaloziopsis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 511, 1885 (Spruce 1885). *** Cephaloziopsis intertexta (Gottsche) R.M.Schust., Nova Hedwigia 22: 183, 1971 [1972] (Schuster 1971b). Bas.: Jungermannia intertexta Gottsche, Syn. Hepat. 1: 107, 1844 (Gottsche et al. 1844). ** Chaetophyllopsis R.M.Schust., J. Hattori Bot. Lab. 23: 69, 1960 [1961] (Schuster 1960b). *** Chaetophyllopsis whiteleggei (Carrington et Pearson) R.M.Schust. ex Hamlin, Rec. Domin. Mus. 7: 255, 1972 (Hamlin 1972). Bas.: Jungermannia whiteleggei Carrington et Pearson, Proc. Linn. Soc. New South Wales (ser. 2) 2 (4): 1051, 1888 (Carrington and Pearson 1888a). ** Cylindrocolea R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 12 (3): 666, 1969 (Schuster 1969a). ** subg. Cylindrocolea *** Cylindrocolea kiaeri (Austin) Váňa, Phytotaxa 112 (1): 2, 2013 (Váňa et al. 2013e). Bas.: Jungermannia kiaeri Austin, Bull. Torrey Bot. Club 6 (3): 18, 1875 (Austin 1875b). *** Cylindrocolea sanctae-helenae M.Wigginton, Polish Bot. J. 58 (1): 107, 2013 (Wigginton 2013). ** sect. Cylindrocolea *** Cylindrocolea andersonii R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 33, 1980 (Schuster 1980c). *** Cylindrocolea brasiliensis D.P.Costa, N.D.Santos et Váňa, Bryologist 111 (4): 667, 2008 (Costa et al. 2008). *** Cylindrocolea chevalieri (Steph.) R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 12 (3): 666, 1969 (Schuster 1969a). Bas.: Alobiella chevalieri Steph., Bull. Herb. Boissier (sér. 2) 8 (8): 567 (351), 1908 (Stephani 1908e). 46 Cephaloziella pellucida is probably not a Cephaloziella, but the type specimen has not been found. 47 Cephaloziella pygmaea is possibly conspeciic with Cephaloziella granatensis (Váňa et al. 2014a). 48 Cephaloziella secundifolia could not be studied by Fulford (1976) and she did not know what it is. World checklist of hornworts and liverworts 73 *** Cylindrocolea gittinsii (E.W.Jones) R.M.Schust., Nova Hedwigia 22: 172, 1971 [1972] (Schuster 1971b). Bas.: Cephaloziella gittinsii E.W.Jones, Trans. Brit. Bryol. Soc. 3 (3): 437, 1958 (Jones 1958). *** Cylindrocolea madagascariensis (Steph.) R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 12 (3): 666, 1969 (Schuster 1969a). Bas.: Lophozia madagascariensis Steph., Sp. Hepat. (Stephani) 6: 112, 1917 (Stephani 1917a). *** Cylindrocolea nigerica (E.W.Jones) R.M.Schust., Nova Hedwigia 22: 172, 1971 [1972] (Schuster 1971b). Bas.: Cephaloziella nigerica E.W.Jones, Trans. Brit. Bryol. Soc. 3 (3): 435, 1958 (Jones 1958). *** Cylindrocolea novae-caledoniae (Grolle) R.M.Schust., Nova Hedwigia 22: 161, 1971 [1972] (Schuster 1971b). Bas.: Cephaloziella novae-caledoniae Grolle, Rev. Bryol. Lichénol. 29 (3/4): 208, 1960 [1961] (Grolle 1960a). *** Cylindrocolea planifolia (Steph.) R.M.Schust., Nova Hedwigia 22: 164, 1971 [1972] (Schuster 1971b). Bas.: Cephaloziella planifolia Steph., Hedwigia 32 (5): 317, 1893 (Stephani 1893d). *** Cylindrocolea sprucei R.M.Schust., Nova Hedwigia 22: 163, 1971 [1972] (Schuster 1971b). *** Cylindrocolea ugandica (E.W.Jones) R.M.Schust., Nova Hedwigia 22: 171, 1971 [1972] (Schuster 1971b). Bas.: Cephaloziella ugandica E.W.Jones, Trans. Brit. Bryol. Soc. 3 (3): 433, 1958 (Jones 1958). ** sect. Platycaulis R.M.Schust., Nova Hedwigia 22: 173, 1971 (Schuster 1971b). *** Cylindrocolea recurvifolia (Steph.) Inoue, J. Jap. Bot. 47 (11): 348, 1972 (Inoue 1972a). Bas.: Cephalozia recurvifolia Steph., Bull. Herb. Boissier (sér. 2) 8 (7): 497 (327), 1908 (Stephani 1908f). *** Cylindrocolea tagawae (N.Kitag.) R.M.Schust., Nova Hedwigia 22: 174, 1971 [1972] (Schuster 1971b). Bas.: Cephaloziella tagawae N.Kitag., J. Hattori Bot. Lab. 32: 303, 1969 (Kitagawa 1969c). ** subg. Cylindroscyphus (Douin) R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 20, 1980 (Schuster 1980c). Bas.: Cephaloziella subg. Cylindroscyphus Douin, Mém. Soc. Bot. France 29: 56, 1920 (Douin 1920). *** Cylindrocolea abyssinica (Gola) Váňa, Phytotaxa 112 (1): 2, 2013 (Váňa et al. 2013e). Bas.: Cephaloziella abyssinica Gola, Ann. Bot. (Rome) 13 (1): 68, 1914 (Gola 1914a). ** Cylindrocolea obtusifolia Fulford, Mem. New York Bot. Gard. 11 (4): 401, 1976 (Fulford 1976). ** Cylindrocolea reticulata Udar et Ad.Kumar, Lindbergia 8 (3): 181, 1982 [1983] (Udar and Kumar 1982c). *** Cylindrocolea rhizantha (Mont.) R.M.Schust., Nova Hedwigia 22: 175, 1971 [1972] (Schuster 1971b). Bas.: Jungermannia rhizantha Mont., Hist. Phys. Cuba, Bot., Pl. Cell.: 454, 1842 (Montagne 1842a). 74 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Gottschelia Grolle, J. Hattori Bot. Lab. 31: 13, 1968 (Grolle 1968c). *** Gottschelia maxima (Steph.) Grolle, J. Bryol. 25 (1): 6, 2003 (Grolle et al. 2003). Bas.: Tylimanthus maximus Steph., Sp. Hepat. (Stephani) 6: 249, 1922 (Stephani 1922). *** Gottschelia schizopleura (Spruce) Grolle, J. Hattori Bot. Lab. 31: 16, 1968 (Grolle 1968c). Bas.: Jungermannia schizopleura Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 517, 1885 (Spruce 1885). ** Gymnocoleopsis (R.M.Schust.) R.M.Schust., Phytologia 39 (4): 243, 1978 (Schuster 1978a). Bas.: Gymnocolea subg. Gymnocoleopsis R.M.Schust., Bryologist 70 (1): 111, 1967 (Schuster 1967b). *** Gymnocoleopsis capensis (S.W.Arnell) R.M.Schust., J. Hattori Bot. Lab. 78: 123, 1995 (Schuster 1995c). Bas.: Lophozia capensis S.W.Arnell, Svensk Bot. Tidskr. 47 (1): 112, 1953 (Arnell 1953a). *** Gymnocoleopsis cylindriformis (Mitt.) R.M.Schust., J. Hattori Bot. Lab. 78: 126, 1995 (Schuster 1995c). Bas.: Jungermannia cylindriformis Mitt., J. Linn. Soc., Bot. 15 (84): 196, 1876 (Mitten 1876b). *** Herzogobryum Grolle, Rev. Bryol. Lichénol. 32 (3/4): 160, 1963 [1964] (Grolle 1963d). Nom. nov. pro Chondrophyllum Herzog, Rev. Bryol. Lichénol. 21 (1/2): 46, 1952 (Herzog 1952f). *** Herzogobryum atrocapillum (Hook.f. et Taylor) Grolle, Österr. Bot. Z. 113 (2): 228, 1966 (Grolle 1966j). Bas.: Gymnomitrion atrocapillum Hook.f. et Taylor, London J. Bot. 5: 258, 1846 (Taylor 1846a). *** Herzogobryum iliforme R.M.Schust., Phytologia 45 (5): 422, 1980 (Schuster 1980b). *** Herzogobryum molle Grolle, Österr. Bot. Z. 113 (2): 226, 1966 (Grolle 1966j). *** Herzogobryum vermiculare (Schifn.) Grolle, J. Hattori Bot. Lab. 28: 103, 1965 (Grolle 1965e). Bas.: Gymnomitrion vermiculare Schifn., Leberm., Forschungsr. Gazelle 4 (4): 2, 1890 (Schifner 1890). ** Kymatocalyx Herzog, Memoranda Soc. Fauna Fl. Fennica 25: 56, 1950 (Herzog 1950c). *** Kymatocalyx africanus Váňa et M.Wigginton, Haussknechtia, Beih. 9: 158, 1999 (Gradstein and Váňa 1999). *** Kymatocalyx dominicensis (Spruce) Váňa, Österr. Bot. Z. 118 (5): 575, 1970 (Váňa 1970b). Bas.: Jungermannia dominicensis Spruce, J. Linn. Soc., Bot. 30 (210): 363, 1895 (Gepp 1895b). *** Kymatocalyx madagascariensis (Steph.) Gradst. et Váňa, Haussknechtia, Beih. 9: 164, 1999 (Gradstein and Váňa 1999). Bas.: Acrobolbus madagascariensis Steph., Bull. Herb. Boissier (sér. 2) 2 (5): 460 (179), 1902 (Stephani 1902f). *** Kymatocalyx rhizomaticus (Herzog) Gradst. et Váňa, Haussknechtia, Beih. 9: 166, 1999 (Gradstein and Váňa 1999). Bas.: Stenorrhipis rhizomatica Herzog, Trans. Brit. Bryol. Soc. 1 (4): 290, 1950 (Herzog 1950a). World checklist of hornworts and liverworts 75 *** Lophonardia R.M.Schust., Phytologia 39 (4): 244, 1978 (Schuster 1978a). *** Lophonardia jamesonii (Mont.) L.Söderstr. et Váňa, Phytotaxa 81 (1): 19, 2013 (Söderström et al. 2013a). Bas.: Jungermannia jamesonii Mont., Syll. Gen. Sp. Crypt.: 60, 1856 (Montagne 1856b). *** Lophonardia laxifolia (Mont.) L.Söderstr. et Váňa, Phytotaxa 81 (1): 20, 2013 (Söderström et al. 2013a). Bas.: Sarcocyphos laxifolius Mont., Ann. Sci. Nat. Bot. (sér. 3) 4: 346, 1845 (Montagne 1845b). *** Lophonardia tristaniana (S.W.Arnell) L.Söderstr. et Váňa, Phytotaxa 81 (1): 20, 2013 (Söderström et al. 2013a). Bas.: Jungermannia tristaniana S.W.Arnell, Results Norweg. Sci. Exped. Tristan da Cunha 42: 12, 1958 (Arnell 1958b). *** Nothogymnomitrion R.M.Schust., J. Hattori Bot. Lab. 80: 43, 1996 (Schuster 1996a). *** Nothogymnomitrion erosum (Carrington et Pearson) R.M.Schust., J. Hattori Bot. Lab. 80: 44, 1996 (Schuster 1996a). Bas.: Cesius erosus Carrington et Pearson, Pap. & Proc. Roy. Soc. Tasmania 1887: 8, 1888 (Carrington and Pearson 1888b). *** Obtusifolium S.W.Arnell, Ill. Moss Fl. Fennosc. Hep.: 309, 1956 (Arnell 1956b). *** Obtusifolium obtusum (Lindb.) S.W.Arnell, Ill. Moss Fl. Fennosc. Hep.: 133, 1956 (Arnell 1956b). Bas.: Jungermannia obtusa Lindb., Musci Scand.: 7, 1879 (Lindberg 1879). *** Oleolophozia L.Söderstr., De Roo et Hedd., Phytotaxa 3: 50, 2010 (Söderström et al. 2010b). *** Oleolophozia perssonii (H.Buch et S.W.Arnell) L.Söderstr., De Roo et Hedd., Phytotaxa 3: 51, 2010 (Söderström et al. 2010b). Bas.: Lophozia perssonii H.Buch et S.W.Arnell, Bot. Not. 97: 382, 1944 (Buch 1944). ** Phycolepidozia R.M.Schust., Bull. Torrey Bot. Club 93 (6): 438, 1966 [1967] (Schuster 1966f). ** subg. Metaphycolepidozia Gradst., J.-P.Frahm et U.Schwarz, Taxon 63 (3): 506, 2014 (Gradstein et al. 2014a). ** Phycolepidozia indica Gradst., J.-P.Frahm et U.Schwarz, Taxon 63 (3): 499, 2014 (Gradstein et al. 2014a). ** subg. Phycolepidozia *** Phycolepidozia exigua R.M.Schust., Bull. Torrey Bot. Club 93 (6): 440, 1966 [1967] (Schuster 1966f). ** Protolophozia (R.M.Schust.) Schljakov, Novosti Sist. Nizš. Rast. 16: 204, 1979 (Shliakov 1979). Bas.: Lophozia subg. Protolophozia R.M.Schust., Nova Hedwigia 15: 472, 1968 (Schuster 1968b). 76 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Protolophozia androgyna R.M.Schust. ex Váňa et L.Söderstr., Phytotaxa 76 (3): 51, 2013 (Váňa et al. 2013l). Based on: Lophozia androgyna R.M.Schust., Beih. Nova Hedwigia 119: 266, 2002 (Schuster 2002b), nom. inval. ** Protolophozia autoica (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 76 (3): 51, 2013 (Váňa et al. 2013l). Bas.: Lophozia autoica R.M.Schust., Nova Hedwigia 15: 479, 1968 (Schuster 1968b). ** Protolophozia crispata (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 76 (3): 51, 2013 (Váňa et al. 2013l). Bas.: Lophozia crispata R.M.Schust., Nova Hedwigia 15: 474, 1968 (Schuster 1968b). *** Protolophozia druceae (Grolle et E.A.Hodgs.) Váňa et L.Söderstr., Phytotaxa 76 (3): 51, 2013 (Váňa et al. 2013l). Bas.: Lophozia druceae Grolle et E.A.Hodgs., J. Roy. Soc. New Zealand 2 (1): 112, 1972 (Hodgson 1972). *** Protolophozia elongata (Steph.) Schljakov, Novosti Sist. Nizš. Rast. 16: 204, 1979 (Shliakov 1979). Bas.: Lophozia elongata Steph., Bull. Herb. Boissier (sér. 2) 2 (1): 41 (141), 1902 (Stephani 1902c). *** Protolophozia herzogiana (E.A.Hodgs. et Grolle) Váňa et L.Söderstr., Phytotaxa 76 (3): 51, 2013 (Váňa et al. 2013l). Bas.: Lophozia herzogiana E.A.Hodgs. et Grolle, Rev. Bryol. Lichénol. 31 (3/4): 152, 1962 [1963] (Grolle 1962c). *** Protolophozia lancistipa (Grolle) Váňa et L.Söderstr., Phytotaxa 76 (3): 51, 2013 (Váňa et al. 2013l). Bas.: Andrewsianthus lancistipus Grolle, Marion Prince Edw. Is: 230, 1971 (Grolle 1971d). *** Protolophozia leucorhiza (Mitt.) Váňa et L.Söderstr., Phytotaxa 76 (3): 51, 2013 (Váňa et al. 2013l). Bas.: Jungermannia leucorhiza Mitt., J. Linn. Soc., Bot. 15 (82): 68, 1876 (Mitten 1876a). *** Protolophozia longilora (Herzog) L.Söderstr. et Váňa, Phytotaxa 76 (3): 51, 2013 (Váňa et al. 2013l). Bas.: Orthocaulis longilorus Herzog, Rev. Bryol. Lichénol. 23 (1/2): 32, 1954 (Herzog 1954). ** Protolophozia monoica (E.A.Hodgs.) Váňa et L.Söderstr., Phytotaxa 76 (3): 52, 2013 (Váňa et al. 2013l). Bas.: Metahygrobiella monoica E.A.Hodgs., Trans. Roy. Soc. New Zealand, Bot. 3 (4): 76, 1965 (Hodgson 1965). *** Protolophozia multicuspidata (Hook.f. et Taylor) Váňa et L.Söderstr., Phytotaxa 76 (3): 52, 2013 (Váňa et al. 2013l). Bas.: Jungermannia multicuspidata Hook.f. et Taylor, London J. Bot. 3: 375, 1844 (Hooker and Taylor 1844a). ** Protolophozia nivicola (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 76 (3): 52, 2013 (Váňa et al. 2013l). Bas.: Lophozia nivicola R.M.Schust., Nova Hedwigia 15: 477, 1968 (Schuster 1968b). *** Protolophozia perssoniana (H.A.Mill.) Váňa et L.Söderstr., Phytotaxa 76 (3): 52, 2013 (Váňa et al. 2013l). Bas.: Lophozia perssoniana H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 508, 1963 (Miller 1963). ** Protolophozia subalpina (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 76 (3): 52, 2013 (Váňa et al. 2013l). Bas.: Lophozia autoica var. subalpina R.M.Schust., Nova Hedwigia 15: 482, 1968 (Schuster 1968b). World checklist of hornworts and liverworts 77 *** Protolophozia tasmanica (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 76 (3): 52, 2013 (Váňa et al. 2013l). Bas.: Lophozia tasmanica R.M.Schust., Nova Hedwigia 15: 484, 1968 (Schuster 1968b). *** Protolophozia verruculosa (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 76 (3): 52, 2013 (Váňa et al. 2013l). Bas.: Lophozia verruculosa R.M.Schust., Phytologia 39 (4): 242, 1978 (Schuster 1978a). *** Lophoziaceae Cavers by L. Söderström and J. Váňa Lophoziaceae was broadly deined by de Roo et al. (2007) excluding Anastrophyllaceae and Jamesonielloideae and also moving several traditionally included genera to other families. Vilnet et al. (2010) further reined the family. Nomenclatural and taxonomic notes can also be found in Vilnet et al. (2007b, 2008), Söderström et al. (2013c) and Váňa et al. (2013m). he placement of Gerhildiella and Pseudocephaloziella in the family is provisional. *** Andrewsianthus R.M.Schust., Rev. Bryol. Lichénol. 30 (1/2): 66, 1961 (Schuster 1961a). ** subg. Andrewsianthus *** Andrewsianthus aberrans (Nees et Mont.) Grolle, Trans. Brit. Bryol. Soc. 4 (3): 440, 1963 (Grolle 1963a). Bas.: Jungermannia aberrans Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 250, 1843 (Montagne 1843). *** Andrewsianthus bidens (Mitt. ex Steph.) R.M.Schust., Nova Hedwigia 15: 492, 1968 (Schuster 1968b). Bas.: Lophozia bidens Mitt. ex Steph., Bull. Herb. Boissier (sér. 2) 2 (1): 41 (141), 1902 (Stephani 1902c). *** Andrewsianthus bilobus (Mitt.) Grolle, Trans. Brit. Bryol. Soc. 4 (3): 437, 1963 (Grolle 1963a). Bas.: Gymnanthe biloba Mitt., J. Proc. Linn. Soc., Bot. 7 (27): 166, 1863 (Mitten 1863). *** Andrewsianthus carinatus Grolle, Marion Prince Edw. Is: 229, 1971 (Grolle 1971d). *** Andrewsianthus cavifolius Grolle et Váňa, J. Hattori Bot. Lab. 38: 640, 1974 (Váňa 1974b). *** Andrewsianthus chimbuensis R.M.Schust., Nova Hedwigia 15: 491, 1968 (Schuster 1968b). *** Andrewsianthus kinabaluensis N.Kitag., J. Hattori Bot. Lab. 33: 205, 1970 (Kitagawa 1970). *** Andrewsianthus koponenii Váňa et Piippo, Ann. Bot. Fenn. 26 (3): 284, 1989 (Váňa and Piippo 1989). *** Andrewsianthus mizutanii N.Kitag., J. Hattori Bot. Lab. 32: 307, 1969 (Kitagawa 1969a). 78 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Andrewsianthus papillosus N.Kitag., J. Hattori Bot. Lab. 33: 207, 1970 (Kitagawa 1970). *** Andrewsianthus perigonialis (Hook.f. et Taylor) R.M.Schust., Beih. Nova Hedwigia 119: 336, 2002 (Schuster 2002b). Bas.: Jungermannia perigonialis Hook.f. et Taylor, London J. Bot. 3: 368, 1844 (Hooker and Taylor 1844a). *** Andrewsianthus puniceus (Nees) R.M.Schust. ex Grolle, Trans. Brit. Bryol. Soc. 4 (3): 439, 1963 (Grolle 1963a). Bas.: Jungermannia punicea Nees, Enum. Pl. Crypt. Javae: 32, 1830 (Nees 1830). *** Andrewsianthus recurvifolius (Nees) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 710, 1969 (Schuster 1969b). Bas.: Jungermannia recurvifolia Nees, Enum. Pl. Crypt. Javae: 32, 1830 (Nees 1830). *** Andrewsianthus sundaicus (Schifn.) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 710, 1969 (Schuster 1969b). Bas.: Anastrophyllum sundaicum Schifn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 202, 1898 (Schifner 1898a). *** Andrewsianthus zantenii Váňa, J. Hattori Bot. Lab. 38: 645, 1974 (Váňa 1974b). ** subg. Cephalolobus (R.M.Schust.) R.M.Schust., Beih. Nova Hedwigia 119: 326, 2002 (Schuster 2002b). Bas.: Cephalolobus R.M.Schust., Rev. Bryol. Lichénol. 34 (1/2): 244, 1966 (Schuster 1966b). *** Andrewsianthus hodgsoniae (R.M.Schust.) R.M.Schust. ex J.J.Engel, Novon 17 (3): 311, 2007 (Engel 2007). Bas.: Cephalolobus hodgsoniae R.M.Schust., Rev. Bryol. Lichénol. 34 (1/2): 254, 1966 (Schuster 1966b). *** Andrewsianthus marionensis (S.W.Arnell) Grolle, Marion Prince Edw. Is: 232, 1971 (Grolle 1971d). Bas.: Lophozia marionensis S.W.Arnell, Svensk Bot. Tidskr. 47 (3): 421, 1953 (Arnell 1953c). *** Andrewsianthus scabrellus (C.Massal.) R.M.Schust. ex J.J.Engel, Novon 17 (3): 311, 2007 (Engel 2007). Bas.: Cephalozia scabrella C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 233, 1885 (Massalongo 1885). ** Andrewsianthus sphenoloboides (R.M.Schust.) R.M.Schust. ex J.J.Engel, Novon 17 (3): 311, 2007 (Engel 2007). Bas.: Cephalolobus sphenoloboides R.M.Schust., Rev. Bryol. Lichénol. 34 (1/2): 251, 1966 (Schuster 1966b). ** Gerhildiella Grolle, Rev. Bryol. Lichénol. 34 (1/2): 187, 1966 (Grolle 1966a). *** Gerhildiella rossneriana Grolle, Rev. Bryol. Lichénol. 34 (1/2): 187, 1966 (Grolle 1966a). *** Heterogemma (Jørg.) Konstant. et Vilnet, Arctoa 18: 67, 2009 [2010] (Konstantinova and Vilnet 2009). Bas.: Lophozia sect. Heterogemma Jørg., Bergens Mus. Skr. (n.ser.) 16: 146, 1934 (Jørgensen 1934). *** Heterogemma capitata (Hook.) Konstant. et Vilnet, Arctoa 18: 67, 2009 [2010] (Konstantinova and Vilnet 2009). Bas.: Jungermannia capitata Hook., Brit. Jungermann.: tab. 80, 1816 (Hooker 1816a). World checklist of hornworts and liverworts 79 *** Heterogemma laxa (Lindb.) Konstant. et Vilnet, Arctoa 18: 67, 2009 [2010] (Konstantinova and Vilnet 2009). Bas.: Jungermannia laxa Lindb., Acta Soc. Sci. Fenn. 10: 529, 1875 (Lindberg 1875). *** Heterogemma patagonica (Herzog et Grolle) L.Söderstr. et Váňa, Phytotaxa 97 (2): 29, 2013 (Söderström et al. 2013c). Bas.: Lophozia patagonica Herzog et Grolle, Rev. Bryol. Lichénol. 28 (3/4): 343, 1959 [1960] (Grolle 1959a). *** Lophozia (Dumort.) Dumort., Recueil Observ. Jungerm.: 17, 1835 (Dumortier 1835). Bas.: Jungermannia sect. Lophozia Dumort., Syll. Jungerm. Europ.: 53, 1831 (Dumortier 1831). 49 *** Lophozia ascendens (Warnst.) R.M.Schust., Bryologist 55 (3): 180, 1952 (Schuster 1952). Bas.: Sphenolobus ascendens Warnst., Hedwigia 57 (1/2): 63, 1916 (Warnstorf 1916). * Lophozia austrosibirica Bakalin, Ann. Bot. Fenn. 40 (1): 49, 2003 (Bakalin 2003). ** Lophozia ciliata Damsh., L.Söderstr. et H.Weibull, Lindbergia 25 (1): 3, 2000 (Söderström et al. 2000). *** Lophozia guttulata (Lindb. et Arnell) A.Evans, Proc. Wash. Acad. Sci. 2: 302, 1900 (Evans 1900c). Bas.: Jungermannia guttulata Lindb. et Arnell, Kongl. Svenska Vetensk.-Akad. Handl. (n.ser.) 23 (5): 51, 1889 (Lindberg and Arnell 1889). * Lophozia jamaicensis (Nees) Steph., Bull. Herb. Boissier (sér. 2) 1 (2): 128 (155), 1901 (Stephani 1901h). Bas.: Jungermannia jamaicensis Nees, Syn. Hepat. 1: 105, 1844 (Gottsche et al. 1844). 50 ** Lophozia lacerata N.Kitag., Hikobia 3 (3): 172, 1963 (Kitagawa 1963a). ** Lophozia lantratoviae Bakalin, Ann. Bot. Fenn. 40 (1): 47, 2003 (Bakalin 2003). ** Lophozia murmanica Kaal., Rep. Second Norweg. Arctic Exped. 11: 34, 1906 (Bryhn 1906). ** Lophozia paciica Bakalin, Bryologist 114 (2): 302, 2011 (Bakalin 2011). * Lophozia pallida (Steph.) Grolle, J. Jap. Bot. 39 (6): 174, 1964 (Grolle 1964a). Bas.: Anastrophyllum pallidum Steph., Bull. Herb. Boissier (sér. 2) 1 (10): 1131 (114), 1901 (Stephani 1901c). 51 ** Lophozia savicziae Schljakov, Novosti Sist. Nizš. Rast. 10: 299, 1973 (Shliakov 1973). * Lophozia schusterana Schljakov, Novosti Sist. Nizš. Rast. 12: 320, 1975 (Shliakov 1975). *** Lophozia silvicola H.Buch, Beret. 18 Skand. Naturforskarmøde: 228, 1929 (Buch 1929). *** Lophozia silvicoloides N.Kitag., J. Hattori Bot. Lab. 28: 276, 1965 (Kitagawa 1965). 49 Lophozia has many nomenclatural and taxonomic problems. his treatment follows Söderström et al. (2013c). 50 Lophozia jamaicensis is probably not a Lophozia, but the type specimen is missing in Nees’s herbarium (Söderström et al. 2010a). 51 Lophozia pallida may be conspeciic with Lophozia guttulata (Bakalin 2003). 80 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) * Lophozia subapiculata R.M.Schust. et Damsh., Meddel. Grønland 199 (1): 104, 1974 (Schuster and Damsholt 1974). 52 ** Lophozia udarii S.Srivast., S.C.Srivast. et K.K.Rawat, Nelumbo 55: 130, 2013 (Srivastava et al. 2013). 53 *** Lophozia ventricosa (Dicks.) Dumort., Recueil Observ. Jungerm.: 17, 1835 (Dumortier 1835). Bas.: Jungermannia ventricosa Dicks., Fasc. Pl. Crypt. Brit. 2: 14, 1790 (Dickson 1790). 54 * Lophozia wenzelii (Nees) Steph., Bull. Herb. Boissier (sér. 2) 2 (1): 35 (135), 1902 (Stephani 1902c). Bas.: Jungermannia wenzelii Nees, Naturgesch. Eur. Leberm. 2: 58, 1836 (Nees 1836). 55 * Excluded from the genus Lophozia serpens (Dumort.) Dumort., Recueil Observ. Jungerm.: 17, 1835 (Dumortier 1835). Bas.: Jungermannia serpens Dumort., Syll. Jungerm. Europ.: 56, 1831 (Dumortier 1831). 56 ** Lophoziopsis Konstant. et Vilnet, Arctoa 18: 66, 2009 [2010] (Konstantinova and Vilnet 2009). *** Lophoziopsis excisa (Dicks.) Konstant. et Vilnet, Arctoa 18: 66, 2009 [2010] (Konstantinova and Vilnet 2009). Bas.: Jungermannia excisa Dicks., Fasc. Pl. Crypt. Brit. 3: 11, 1793 (Dickson 1793). ** Lophoziopsis excisa var. elegans (R.M.Schust.) Konstant. et Vilnet, Arctoa 18: 66, 2009 [2010] (Konstantinova and Vilnet 2009). Bas.: Lophozia excisa var. elegans R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 522, 1969 (Schuster 1969b). ** Lophoziopsis excisa var. infuscata (R.M.Schust. et Damsh.) Konstant. et Vilnet, Arctoa 18: 66, 2009 [2010] (Konstantinova and Vilnet 2009). Bas.: Lophozia excisa var. infuscata R.M.Schust. et Damsh., Meddel. Grønland 199 (1): 94, 1974 (Schuster and Damsholt 1974). ** Lophoziopsis excisa var. succulenta (R.M.Schust. et Damsh.) Konstant. et Vilnet, Arctoa 18: 66, 2009 [2010] (Konstantinova and Vilnet 2009). Bas.: Lophozia excisa var. 52 Lophozia subapiculata may be a form of Lophozia guttulata, but Bakalin (2005) treated it as conspeciic with Lophozia ventricosa. 53 Lophozia udarii has recently been described using the generic concept of Lophozia in the wide sense (as e.g. Schuster 1969b). We do not know where to refer the species. It is possibly a Schistochilopsis species close to Schistochilopsis incisa. 54 Lophozia ventricosa is very variable and many segregates at various subspeciic levels have been described. he species complex also includes at least Lophozia austrosibirica, Lophozia schusteriana and Lophozia wenzelii. 55 Lophozia wenzelii is closely related to Lophozia ventricosa according to De Roo et al. (2007) and Vilnet et al. (2010) based on molecular and morphological evidence. It is variable and many varieties are described. he variability and relationships to related species needs further study. 56 Lophozia serpens is probably conspeciic with Schistochilopsis incisa. World checklist of hornworts and liverworts *** ** *** ** *** ** * ** 81 succulenta R.M.Schust. et Damsh., Meddel. Grønland 199 (1): 96, 1974 (Schuster and Damsholt 1974). Lophoziopsis longidens (Lindb.) Konstant. et Vilnet, Arctoa 18: 66, 2009 [2010] (Konstantinova and Vilnet 2009). Bas.: Jungermannia longidens Lindb., Helsingf. Dagbl. 1876 (323, 26 Nov.): 2, 1876 (Lindberg 1876a). Lophoziopsis longidens subsp. arctica (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 97 (2): 28, 2013 (Söderström et al. 2013c). Bas.: Lophozia longidens subsp. arctica R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 539, 1969 (Schuster 1969b). 57 Lophoziopsis pellucida (R.M.Schust.) Konstant. et Vilnet, Arctoa 18: 67, 2009 [2010] (Konstantinova and Vilnet 2009). Bas.: Lophozia pellucida R.M.Schust., Canad. J. Bot. 39 (4): 978, 1961 (Schuster 1961b). Lophoziopsis pellucida var. minor (R.M.Schust.) L.Söderstr. et Váňa, Phytotaxa 97 (2): 28, 2013 (Söderström et al. 2013c). Bas.: Lophozia pellucida var. minor R.M.Schust., Canad. J. Bot. 39 (4): 984, 1961 (Schuster 1961b). Lophoziopsis polaris (R.M.Schust.) Konstant. et Vilnet, Arctoa 18: 67, 2009 [2010] (Konstantinova and Vilnet 2009). Bas.: Lophozia alpestris subsp. polaris R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 614, 1969 (Schuster 1969b). Lophoziopsis polaris var. sphagnorum (R.M.Schust.) Konstant. et Vilnet, Arctoa 18: 67, 2009 [2010] (Konstantinova and Vilnet 2009). Bas.: Lophozia alpestris f. sphagnorum R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 619, 1969 (Schuster 1969b). Lophoziopsis propagulifera (Gottsche) Konstant. et Vilnet, Arctoa 18: 67, 2009 [2010] (Konstantinova and Vilnet 2009). Bas.: Jungermannia propagulifera Gottsche, Int. Polarforsch., Deutsch. Exped. 2: 451, 1890 (Gottsche 1890). 58 Lophoziopsis rubrigemma (R.M.Schust.) Konstant. et Vilnet, Arctoa 18: 67, 2009 [2010] (Konstantinova and Vilnet 2009). Bas.: Lophozia rubrigemma R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 621, 1969 (Schuster 1969b). ** Pseudocephaloziella R.M.Schust., Phytologia 39 (4): 243, 1978 (Schuster 1978a). *** Pseudocephaloziella epiphytica R.M.Schust., Phytologia 39 (4): 243, 1978 (Schuster 1978a). *** Trilophozia (R.M.Schust.) Bakalin, Monogr. Lophozia: 34, 2005 (Bakalin 2005). Bas.: Tritomaria subg. Trilophozia R.M.Schust., Amer. Midl. Naturalist 49 (2): 382, 1953 (Schuster 1953). *** Trilophozia quinquedentata (Huds.) Bakalin, Monogr. Lophozia: 34, 2005 (Bakalin 2005). Bas.: Jungermannia quinquedentata Huds., Fl. Angl. (Hudson): 433, 1762 (Hudson 1762). 57 Lophoziopsis longidens var. arctica was treated as a form of Lophoziopsis pellucida by Bakalin (2005). 58 Lophoziopsis propagulifera belongs to the Lophoziopsis excisa species complex. It is probably only a modiication (cf. discussion by Váňa and Engel 2013). 82 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Trilophozia quinquedentata var. asymmetrica (Horik.) L.Söderstr. et Váňa, Phytotaxa 97 (2): 32, 2013 (Söderström et al. 2013c). Bas.: Lophozia asymmetrica Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 153, 1934 (Horikawa 1934). *** Tritomaria Schifn. ex Loeske, Hedwigia 49 (1/2): 13, 1909 (Loeske 1909). Based on: Tritomaria Schifn., Ber. Naturwiss.-Med. Vereins Innsbruck 31 [Beilage]: 12, 1908 (Schifner 1908a). *** Tritomaria exsecta (Schmidel) Schifn. ex Loeske, Hedwigia 49 (1/2): 13, 1909 (Loeske 1909). Bas.: Jungermannia exsecta Schmidel, Syst. Samml. Crypt. Gew. 2: 5, 1797 (Schrader 1797), nom. conserv. ** Tritomaria exsecta subsp. novaezelandiae J.J.Engel, Bryologist 109 (1): 61, 2006 (Engel 2006b). *** Tritomaria exsectiformis (Breidl.) Schifn. ex Loeske, Hedwigia 49 (1/2): 13, 1909 (Loeske 1909). Bas.: Jungermannia exsectiformis Breidl., Mitt. Naturwiss. Vereins Steiermark 30: 321, 1894 (Breidler 1894). ** Tritomaria exsectiformis subsp. arctica R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 661, 1969 (Schuster 1969b). ** Tritomaria exsectiformis subsp. camerunensis S.W.Arnell ex Váňa, Phytotaxa 167 (2): 216, 2014 (Váňa et al. 2014b). Based on: Tritomaria camerunensis S.W.Arnell, Svensk Bot. Tidskr. 52 (1): 64, 1958 (Arnell 1958a), nom. inval. *** Tritomaria ferruginea (Grolle) Váňa, Phytotaxa 81 (1): 24, 2013 (Váňa et al. 2013m). Bas.: Andrewsianthus ferrugineus Grolle, Khumbu Himal 1 (4): 275, 1966 (Grolle 1966k). * Tritomaria koreana Bakalin, S.S.Choi et B.Y.Sun, Arctoa 18: 163, 2009 [2010] (Bakalin et al. 2009a). * Tritomaria mexicana Bakalin, Arctoa 17: 162, 2008 [2009] (Bakalin 2008b). *** Tritomaria scitula (Taylor) Jørg., Bergens Mus. Aarbok 1919/20 (7): 3, 1922 (Jørgensen 1922). Bas.: Jungermannia scitula Taylor, London J. Bot. 5: 274, 1846 (Taylor 1846a). *** Scapaniaceae Mig. by J. Váňa Scapaniaceae, in its classical concept, was recognized as monophyletic and nested within Lophoziaceae s. lat. (cf. Davis 2004, Schill et al. 2004, Yatsentyuk et al. 2004, He-Nygrén et al. 2006). More recent molecular studies classiied Scapaniaceae as a sister clade to Lophoziaceae, and the genus Schistochilopsis, a segregate of the genus Lophozia s. lat., more closely related to Scapaniaceae than to Lophoziaceae (cf. de Roo et al. 2007, Vilnet et al. 2007). he genus Scapania was recently studied by morphological methods by Potemkin (2002), by molecular methods by Vilnet et al. (2007) and especially Heinrichs et al. (2012a). Some taxonomic and nomenclatural notes World checklist of hornworts and liverworts 83 can be found in Váňa et al. (2012a, 2013j, 2015) and Konstantinova et al. (2013a). he placement of Pseudotritomaria, Saccobasis and Schistochilopsis in the family follows Vilnet et al. (2010). *** Diplophyllum (Dumort.) Dumort., Recueil Observ. Jungerm.: 15, 1835 (Dumortier 1835) nom. conserv. Bas.: Jungermannia sect. Diplophyllum Dumort., Syll. Jungerm. Europ.: 44, 1831 (Dumortier 1831). *** subg. Austrodiplophyllum R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 11 (1): 18, 1968 (Schuster 1968a). ** Diplophyllum recurvifolium C.Massal., Atti Reale Ist. Veneto Sci. Lett. Arti 87 (2): 221, 1928 (Massalongo 1928). *** Diplophyllum squarrosum Steph., Sp. Hepat. (Stephani) 4: 116, 1910 (Stephani 1910b). *** Diplophyllum verrucosum R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 11 (1): 19, 1968 (Schuster 1968a). *** subg. Diplophyllum *** sect. Diplophyllum *** Diplophyllum albicans (L.) Dumort., Recueil Observ. Jungerm.: 16, 1835 (Dumortier 1835). Bas.: Jungermannia albicans L., Sp. Pl. 1: 1133, 1753 (Linnaeus 1753). *** sect. Protodiplophyllum (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 76 (3): 29, 2013 (Váňa et al. 2013j). Bas.: Diplophyllum subg. Protodiplophyllum R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 192, 1974 (Schuster 1974). *** Diplophyllum africanum S.W.Arnell, Ark. Bot. (n.ser.) 3 (16): 527, 1956 (Arnell 1956e). *** Diplophyllum andicola R.M.Schust., Phytologia 39 (4): 248, 1978 (Schuster 1978a). *** Diplophyllum andrewsii A.Evans, Bryologist 25 (2): 28, 1922 (Evans 1922). * Diplophyllum androgynum J.J.Engel et G.L.Merr., J. Hattori Bot. Lab. 84: 277, 1998 (Engel and Smith Merrill 1998). * Diplophyllum angustifolium J.J.Engel et G.L.Merr., J. Hattori Bot. Lab. 84: 262, 1998 (Engel and Smith Merrill 1998). *** Diplophyllum apiculatum (A.Evans) Steph., Sp. Hepat. (Stephani) 4: 110, 1910 (Stephani 1910b). Bas.: Diplophylleia apiculata A.Evans, Bot. Gaz. 34 (5): 372, 1902 (Evans 1902b). ** Diplophyllum dioicum R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 11 (1): 20, 1968 (Schuster 1968a). ** Diplophyllum exiguum Steph., Sp. Hepat. (Stephani) 6: 500, 1924 (Stephani 1924). * Diplophyllum gemmiparum J.J.Engel et G.L.Merr., J. Hattori Bot. Lab. 84: 255, 1998 (Engel and Smith Merrill 1998). 84 * *** * *** *** ** *** *** ** *** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Diplophyllum incrassatum J.J.Engel et G.L.Merr., J. Hattori Bot. Lab. 84: 265, 1998 (Engel and Smith Merrill 1998). Diplophyllum nanum Herzog, Memoranda Soc. Fauna Fl. Fennica 26: 48, 1950 [1951] (Herzog 1950b). Diplophyllum novum J.J.Engel et G.L.Merr., J. Hattori Bot. Lab. 84: 274, 1998 (Engel and Smith Merrill 1998). Diplophyllum obtusatum (R.M.Schust.) R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 215, 1974 (Schuster 1974). Bas.: Diplophyllum apiculatum var. obtusatum R.M.Schust., Amer. Midl. Naturalist 49 (2): 432, 1953 (Schuster 1953). Diplophyllum obtusifolium (Hook.) Dumort., Recueil Observ. Jungerm.: 16, 1835 (Dumortier 1835). Bas.: Jungermannia obtusifolia Hook., Brit. Jungermann.: tab. 26, 1812 (Hooker 1812). 59 Diplophyllum obtusifolium subsp. domesticum (Gottsche) Váňa, Phytotaxa 76 (3): 29, 2013 (Váňa et al. 2013j). Bas.: Jungermannia domestica Gottsche, Linnaea 28 (5): 548, 1856 [1857] (Gottsche 1856). Diplophyllum serrulatum (Müll.Frib.) Steph., Sp. Hepat. (Stephani) 4: 112, 1910 (Stephani 1910b). Bas.: Diplophylleia serrulata Müll.Frib., Bull. Herb. Boissier (sér. 2) 3 (1): 34, 1903 (Müller 1903). Diplophyllum taxifolium (Wahlenb.) Dumort., Recueil Observ. Jungerm.: 16, 1835 (Dumortier 1835). Bas.: Jungermannia taxifolia Wahlenb., Fl. Lapp. (Wahlenberg): 389, 1812 (Wahlenberg 1812). Diplophyllum taxifolium var. mucronatum R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 203, 1974 (Schuster 1974). Diplophyllum trollii Grolle, Khumbu Himal 1 (4): 273, 1966 (Grolle 1966k). *** Douinia (C.E.O.Jensen) H.Buch, Scapan. N.-Eur. Sib.: 13, 1928 (Buch 1928). Bas.: Diplophylleia subg. Douinia C.E.O.Jensen, Danmarks mosser: 145, 1915 (Jensen 1915). *** Douinia imbricata (M.Howe) Konstant. et Vilnet, Phytotaxa 76 (3): 31, 2013 (Konstantinova et al. 2013a). Bas.: Scapania imbricata M.Howe, Bull. New York Bot. Gard. 2 (6): 104, 1901 (Howe 1901b). *** Douinia ovata (Dicks.) H.Buch, Scapan. N.-Eur. Sib.: 14, 1928 (Buch 1928). Bas.: Jungermannia ovata Dicks., Fasc. Pl. Crypt. Brit. 3: 11, 1793 (Dickson 1793). *** Douinia plicata (Lindb.) Konstant. et Vilnet, Phytotaxa 76 (3): 31, 2013 (Konstantinova et al. 2013a). Bas.: Diplophyllum plicatum Lindb., Acta Soc. Sci. Fenn. 10: 235, 1872 [1873] (Lindberg 1872b). ** Pseudotritomaria Konstant. et Vilnet, Arctoa 18: 66, 2009 [2010] (Konstantinova and Vilnet 2009). 59 Diplophyllum obtusifolium is a species complex also including Diplophyllum androgynum, Diplophyllum angustifolium, Diplophyllum gemmiparum, Diplophyllum incrassatum and Diplophyllum novum. World checklist of hornworts and liverworts 85 *** Pseudotritomaria heterophylla (R.M.Schust.) Konstant. et Vilnet, Arctoa 18: 66, 2009 [2010] (Konstantinova and Vilnet 2009). Bas.: Tritomaria heterophylla R.M.Schust., Canad. J. Bot. 36 (2): 272, 1958 (Schuster 1958b). ** Saccobasis H.Buch, Memoranda Soc. Fauna Fl. Fennica 8: 291, 1932 [1933] (Buch 1932). *** Saccobasis polita (Nees) H.Buch, Memoranda Soc. Fauna Fl. Fennica 8: 292, 1932 [1933] (Buch 1932). Bas.: Jungermannia polita Nees, Naturgesch. Eur. Leberm. 2: 145, 1836 (Nees 1836). ** Saccobasis polymorpha (R.M.Schust.) Schljakov, Novosti Sist. Nizš. Rast. 16: 205, 1979 (Shliakov 1979). Bas.: Tritomaria polita subsp. polymorpha R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 700, 1969 (Schuster 1969b). *** Scapania (Dumort.) Dumort., Recueil Observ. Jungerm.: 14, 1835 (Dumortier 1835) nom. conserv. Bas.: Radula sect. Scapania Dumort., Syll. Jungerm. Europ.: 38, 1831 (Dumortier 1831). ** subg. Ascapania Grolle, Khumbu Himal 1 (4): 268, 1966 (Grolle 1966k). *** Scapania contorta Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 101, 1860 [1861] (Mitten 1860c). ** subg. Gracilidae (H.Buch) Váňa, Hentschel, Joch.Müll. et Heinrichs, PhytoKeys 10: 15, 2012 (Váňa et al. 2012a). Bas.: Scapania sect. Gracilidae H.Buch, Scapan. N.-Eur. Sib.: 106, 1928 (Buch 1928). *** Scapania ampliata Steph., Bull. Herb. Boissier 5 (2): 106, 1897 (Stephani 1897b). ** Scapania ampliata subsp. queenslandica M.L.Hicks, J. Hattori Bot. Lab. 69: 130, 1991 (Hicks 1991). *** Scapania bolanderi Austin, Proc. Acad. Nat. Sci. Philadelphia 21: 218, 1869 (Austin 1869). *** Scapania gracilis Lindb., Morgonbladet (Helsinki) 1873 (286, 9 Dec): 2, 1873 (Lindberg 1873a). ** Scapania macroparaphyllia T.Cao, C.Gao et J.Sun, Acta Phytotax. Sin. 42 (2): 180, 2004 (Cao et al. 2004). *** Scapania maxima Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 223, 1934 (Horikawa 1934). *** Scapania nipponica (Amakawa et S.Hatt.) Amakawa, J. Hattori Bot. Lab. 30: 319, 1967 (Amakawa 1967a). Bas.: Scapania bolanderi var. nipponica Amakawa et S.Hatt., J. Hattori Bot. Lab. 14: 83, 1955 (Amakawa and Hattori 1955). ** Scapania paraphyllia T.Cao, C.Gao, J.Sun et B.R.Zuo, Acta Phytotax. Sin. 45 (3): 311, 2007 (Zuo et al. 2007b). *** Scapania subnimbosa Steph., Sp. Hepat. (Stephani) 4: 150, 1910 (Stephani 1910b). 86 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** subg. Macroscapania R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 248, 1974 (Schuster 1974). * Scapania geppii Steph., Hedwigia 44 (1): 14, 1904 (Stephani 1904i). *** Scapania portoricensis Hampe et Gottsche, Linnaea 25 (3): 342, 1852 [1853] (Hampe and Gottsche 1852). ** Scapania portoricensis var. boliviensis (Steph.) Herzog, Ann. Bryol. 1: 110, 1928 (Herzog 1928). Bas.: Scapania boliviensis Steph., Biblioth. Bot. 87 (2): 231, 1916 (Stephani 1916a). ** Scapania portoricensis var. organensis (Herzog) Herzog, Ann. Bryol. 1: 110, 1928 (Herzog 1928). Bas.: Scapania organensis Herzog, Repert. Spec. Nov. Regni Veg. 21 (1/7): 27, 1925 (Herzog 1925a). ** Scapania portoricensis var. roraimensis Warnst., Hedwigia 63 (2): 109, 1921 (Warnstorf 1921). ** subg. Plicaticalyx Müll.Frib., Bull. Herb. Boissier (sér. 2) 3 (1): 36, 1903 (Müller 1903). ** sect. Planifoliae (Müll.Frib.) Potemkin, J. Hattori Bot. Lab. 85: 56, 1998 (Potemkin 1998). Bas.: Scapania Gruppe Planifoliae Müll.Frib., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 83: 286, 1905 (Müller 1905). *** Scapania davidii Potemkin, Ann. Bot. Fenn. 38 (2): 83, 2001 (Potemkin 2001). ** Scapania ferruginaeoides T.Cao, C.Gao et J.Sun, Guihaia 24 (1): 23, 2004 (Sun et al. 2004). ** Scapania gaochii X.Fu ex T.Cao, Phytotaxa 97 (2): 26, 2013 (Cao et al. 2013). Based on: Scapania gaochii X.Fu ex T.Cao, Acta Bot. Yunnan. 25 (5): 541, 2003 (Cao et al. 2003), nom. inval. *** Scapania harae Amakawa, J. Hattori Bot. Lab. 27: 5, 1964 (Amakawa 1964a). *** Scapania nimbosa Taylor, Nov. Stirp. Pug. 8: 6, 1844 (Lehmann 1844). *** Scapania ornithopodioides (With.) Waddell, Moss Exch. Club Cat. Hepat.: 4, 1897 (Waddell 1897). Bas.: Jungermannia ornithopodioides With., Bot. arr. veg. Gr. Brit. 2: 695, 1776 (Withering 1776). *** Scapania rotundifolia W.E.Nicholson, Symb. Sin. 5: 31, 1930 (Nicholson et al. 1930). *** Scapania secunda Steph., Mém. Soc. Nat. Sci. Nat. Math. Cherbourg 29: 226, 1894 (Stephani 1894b). *** Scapania zhukovae Potemkin, Arctoa 9: 121, 2000 (Potemkin 2000b). ** sect. Plicaticalyx (Müll.Frib.) Potemkin, Ann. Bot. Fenn. 39 (4): 326, 2002 (Potemkin 2002). Bas.: Scapania subg. Plicaticalyx Müll.Frib., Bull. Herb. Boissier (sér. 2) 3 (1): 36, 1903 (Müller 1903). *** Scapania ciliatospinosa Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 222, 1934 (Horikawa 1934). World checklist of hornworts and liverworts 87 *** Scapania ferruginea (Lehm. et Lindenb.) Lehm. et Lindenb., Syn. Hepat. 1: 72, 1844 (Gottsche et al. 1844). Bas.: Jungermannia ferruginea Lehm. et Lindenb., Nov. Stirp. Pug. 4: 20, 1832 (Lehmann 1832). *** Scapania hians Steph. ex Müll.Frib., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 83: 223, 1905 (Müller 1905). ** Scapania hians subsp. salishensis J.D.Godfrey et G.Godfrey, Bryologist 81 (3): 362, 1978 (Godfrey and Godfrey 1978). *** Scapania orientalis Steph. ex Müll.Frib., Bull. Herb. Boissier (sér. 2) 1 (6): 606, 1901 (Müller 1901a). *** Scapania pseudocontorta Potemkin, Arctoa 9: 115, 2000 (Potemkin 2000b). *** Scapania sinikkae Potemkin, Ann. Bot. Fenn. 38 (2): 85, 2001 (Potemkin 2001). *** Scapania spiniloba Potemkin, Arctoa 9: 117, 2000 (Potemkin 2000b). ** subg. Pseudomacrodiplophyllum Váňa, Hentschel, Joch.Müll. et Heinrichs, PhytoKeys 10: 15, 2012 (Váňa et al. 2012a). *** Scapania microdonta (Mitt.) Müll.Frib., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 83: 262, 1905 (Müller 1905). Bas.: Martinellius microdontus Mitt., Trans. Linn. Soc. London, Bot. 3 (3): 196, 1891 (Mitten 1891). *** subg. Scapania ** sect. Aequilobae (Müll.Frib.) H.Buch, Scapan. N.-Eur. Sib.: 110, 1928 (Buch 1928). Bas.: Scapania Gruppe Aequilobae Müll.Frib., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 83: 219, 1905 (Müller 1905). *** Scapania aequiloba (Schwägr.) Dumort., Recueil Observ. Jungerm.: 14, 1835 (Dumortier 1835). Bas.: Jungermannia aequiloba Schwägr., Hist. Musc. Hepat. Prodr.: 24, 1814 (Schwägrichen 1814). *** Scapania aspera M.Bernet et Bernet, Cat. hép. Suisse: 42, 1888 (Bernet 1888). ** sect. Americanae Váňa, Hentschel, Joch.Müll. et Heinrichs, PhytoKeys 10: 15, 2012 (Váňa et al. 2012a). *** Scapania americana Müll.Frib., Bull. Herb. Boissier (sér. 2) 3 (1): 44, 1903 (Müller 1903). ** sect. Apiculatae H.Buch, Scapan. N.-Eur. Sib.: 53, 1928 (Buch 1928). *** Scapania apiculata Spruce, Hep. Pyr. Exsic.: no. 15, 1847 (Spruce 1847). *** Scapania carinthiaca J.B.Jack ex Lindb., Rev. Bryol. 7 (4): 77, 1880 (Lindberg 1880a). *** Scapania carinthiaca var. massalongi Müll.Frib., Bull. Herb. Boissier (sér. 2) 1 (6): 598, 1901 (Müller 1901a). *** Scapania umbrosa (Schrad.) Dumort., Recueil Observ. Jungerm.: 14, 1835 (Dumortier 1835). Bas.: Jungermannia umbrosa Schrad., Syst. Samml. Crypt. Gew. 2: 5, 1797 (Schrader 1797). 88 ** *** *** *** *** *** *** *** *** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) sect. Ciliatae Grolle, Khumbu Himal 1 (4): 272, 1966 (Grolle 1966k). Scapania bhutanensis Amakawa, Fl. E. Himalaya 2: 230, 1971 (Hattori 1971a). Scapania ciliata Sande Lac., Prolus. l. jap.: 209, 1867 (Sande Lacoste 1867). Scapania ciliata subsp. hawaiica (Müll.Frib.) Potemkin, Ann. Bot. Fenn. 39 (4): 321, 2002 (Potemkin 2002). Bas.: Scapania hawaiica Müll.Frib., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 83: 160, 1905 (Müller 1905). Scapania hirosakiensis Steph. ex Müll.Frib., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 83: 120, 1905 (Müller 1905). Scapania hollandiae W.S.Hong, Bryologist 83 (1): 56, 1980 (Hong 1980). Scapania koponenii Potemkin, Ann. Bot. Fenn. 37 (1): 41, 2000 (Potemkin 2000c). Scapania lepida Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 101, 1860 [1861] (Mitten 1860c). Scapania sandei Schifn. ex Müll.Frib., Bull. Herb. Boissier (sér. 2) 1 (6): 612, 1901 (Müller 1901a). ** sect. Compactae (Müll.Frib.) H.Buch, Scapan. N.-Eur. Sib.: 101, 1928 (Buch 1928). Bas.: Scapania Gruppe Compactae Müll.Frib., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 83: 53, 1905 (Müller 1905). *** Scapania compacta (Roth) Dumort., Recueil Observ. Jungerm.: 14, 1835 (Dumortier 1835). Bas.: Jungermannia compacta Roth, Tent. Fl. Germ. 3: 375, 1800 (Roth 1800). *** Scapania kaurinii Ryan, Bot. Not. 42: 210, 1889 (Ryan 1889). *** Scapania spitsbergensis (Lindb.) Müll.Frib., Bull. Herb. Boissier (sér. 2) 1 (6): 607, 1901 (Müller 1901a). Bas.: Martinellius spitsbergensis Lindb., Kongl. Svenska Vetensk.-Akad. Handl. (n.ser.) 23 (5): 31, 1889 (Lindberg and Arnell 1889). ** sect. Curtae (Müll.Frib.) H.Buch, Scapan. N.-Eur. Sib.: 55, 1928 (Buch 1928). Bas.: Scapania Gruppe Curtae Müll.Frib., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 83: 245, 1905 (Müller 1905). *** Scapania curta (Mart.) Dumort., Recueil Observ. Jungerm.: 14, 1835 (Dumortier 1835). Bas.: Jungermannia curta Mart., Fl. crypt. erlang.: 148, 1817 (Martius 1817). ** Scapania curta var. grandiretis R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 393, 1974 (Schuster 1974). ** Scapania curta var. isoloba R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 396, 1974 (Schuster 1974). *** Scapania diplophylloides Amakawa et S.Hatt., J. Hattori Bot. Lab. 9: 59, 1953 (Amakawa and Hattori 1953). *** Scapania esterhuyseniae S.W.Arnell, Bot. Not. 110 (1): 26, 1957 (Arnell 1957a). *** Scapania fulfordiae W.S.Hong, Bryologist 83 (1): 46, 1980 (Hong 1980). *** Scapania gamundiae R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 11 (1): 14, 1968 (Schuster 1968a). World checklist of hornworts and liverworts 89 *** Scapania helvetica Gottsche, Hepat. Eur., Leberm. 42-44: no. 426, 1868 (Gottsche and Rabenhorst 1868). *** Scapania irrigua (Nees) Nees, Syn. Hepat. 1: 67, 1844 (Gottsche et al. 1844). Bas.: Jungermannia irrigua Nees, Naturgesch. Eur. Leberm. 1: 193, 1833 (Nees 1833c). ** Scapania irrigua subsp. rufescens (Loeske) R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 471, 1974 (Schuster 1974). Bas.: Scapania irrigua f. rufescens Loeske, Moosl. Harz.: 71, 1903 (Loeske 1903). *** Scapania lingulata H.Buch, Meddel. Soc. Fauna Fl. Fenn. 42: 92, 1916 (Buch 1916). ** Scapania lingulata var. microphylla (Warnst.) R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 415, 1974 (Schuster 1974). Bas.: Scapania microphylla Warnst., Hedwigia 63 (2): 75, 1921 (Warnstorf 1921). ** Scapania magadanica S.S.Choi, Bakalin et B.Y.Sun, Bot. Paciica 1: 46, 2012 (Choi et al. 2012). *** Scapania mucronata H.Buch, Meddel. Soc. Fauna Fl. Fenn. 42: 91, 1916 (Buch 1916). *** Scapania obcordata (Berggr.) S.W.Arnell, Ark. Bot. (n.ser.) 4 (6): 117, 1959 (Arnell and Mårtensson 1959). Bas.: Sarcocyphos obcordatus Berggr., Kongl. Svenska Vetensk.-Akad. Handl. (n.ser.) 13 (7): 96, 1875 (Berggren 1875). ** Scapania parvifolia Warnst., Hedwigia 63 (2): 78, 1921 (Warnstorf 1921). *** Scapania praetervisa Meyl., Jahresber. Naturf. Ges. Graubündens (n.f.) 64: 364, 1926 (Meylan 1926). *** Scapania scandica (Arnell et H.Buch) Macvicar, Stud. handb. Brit. hepat. (ed. 2): 394, 1926 (Macvicar 1926). Bas.: Martinellius scandicus Arnell et H.Buch, Bot. Not. 74: 1, 1921 (Arnell and Buch 1921). ** Scapania scandica var. argutedentata H.Buch, Scapan. N.-Eur. Sib.: 75, 1928 (Buch 1928). ** Scapania scandica var. dimorpha R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 453, 1974 (Schuster 1974). ** Scapania scandica var. grandiretis (Schljakov) Schljakov, Pečen. Mchi Sev. SSSR 4: 152, 1981 (Shliakov 1981). Bas.: Scapania parvifolia var. grandiretis Schljakov, Novosti Sist. Nizš. Rast. 8: 332, 1971 (Shliakov 1971). *** Scapania uliginosa (Lindenb.) Dumort., Recueil Observ. Jungerm.: 14, 1835 (Dumortier 1835). Bas.: Jungermannia undulata var. uliginosa Lindenb., Syn. hepat. eur: 58, 1829 (Lindenberg 1829). *** Scapania valdonii Váňa, Bedn.-Ochyra et Cykowska, Nova Hedwigia 89 (1/2): 126, 2009 (Váňa et al. 2009). *** Scapania zemliae S.W.Arnell, Svensk Bot. Tidskr. 41: 215, 1947 (Arnell 1947). ** sect. Cuspiduligerae H.Buch, Scapan. N.-Eur. Sib.: 125, 1928 (Buch 1928). *** Scapania cuspiduligera (Nees) Müll.Frib., Lebermoose 2 (22): 472, 1915 (Müller 1915a). Bas.: Jungermannia cuspiduligera Nees, Naturgesch. Eur. Leberm. 1: 180, 1833 (Nees 1833c). ** Scapania cuspiduligera var. diplophyllopsis R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 361, 1974 (Schuster 1974). 90 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** sect. Grolleoscapania Potemkin, Ann. Bot. Fenn. 39 (4): 329, 2002 (Potemkin 2002). *** Scapania karl-muelleri Grolle, Khumbu Himal 1 (4): 270, 1966 (Grolle 1966k). ** sect. Hyperboreae Váňa, Hentschel, Joch.Müll. et Heinrichs, PhytoKeys 10: 16, 2012 (Váňa et al. 2012a). *** Scapania hyperborea Jørg., Forh. Vidensk.-Selsk. Kristiania 1894 (8): 56, 1894 (Jørgensen 1894). *** Scapania paludicola Loeske et Müll.Frib., Lebermoose 2 (21): 425, 1915 (Müller 1915b). ** Scapania paludicola var. viridigemma R.M.Schust., Bull. Natl. Mus. Canada 122: 20, 1950 [1951] (Schuster 1950). *** Scapania tundrae (Arnell) H.Buch, Scapan. N.-Eur. Sib.: 99, 1928 (Buch 1928). Bas.: Martinellius tundrae Arnell, Bot. Not. 74: 289, 1921 (Arnell 1921). ** sect. Kaalaasia (H.Buch) Jørg., Bergens Mus. Skr. (n.ser.) 16: 210, 1934 (Jørgensen 1934). Bas.: Scapania subg. Kaalaasia H.Buch, Scapan. N.-Eur. Sib.: 47, 1928 (Buch 1928). *** Scapania calcicola (Arnell et J.Perss.) Ingham, Naturalist (Hull) 564: 11, 1904 (Ingham 1904). Bas.: Martinellius calcicola Arnell et J.Perss., Rev. Bryol. 30 (6): 97, 1903 (Arnell 1903). *** Scapania gymnostomophila Kaal., Bot. Not. 49: 21, 1896 (Kaalaas 1896). *** Scapania ligulifolia R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 306, 1974 (Schuster 1974). *** Scapania pseudocalcicola R.M.Schust., Phytologia 63 (5): 327, 1987 (Schuster and Damsholt 1987). Scapania sect. Muelleria Potemkin, Ann. Bot. Fenn. 39 (4): 320, 2002 (Potemkin 2002). *** Scapania himalayica Müll.Frib. ex Herzog, Ann. Bryol. 12: 81, 1939 (Herzog 1939b). *** Scapania schljakovii Potemkin, Ann. Bot. Fenn. 38 (2): 87, 2001 (Potemkin 2001). * ** sect. Nemorosae (Müll.Frib.) H.Buch, Scapan. N.-Eur. Sib.: 152, 1928 (Buch 1928). Bas.: Scapania Gruppe Nemorosae Müll.Frib., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 83: 155, 1905 (Müller 1905). *** Scapania brevicaulis Taylor, London J. Bot. 5: 272, 1846 (Taylor 1846a). *** Scapania crassiretis Bryhn, Rev. Bryol. 19 (1): 7, 1892 (Bryhn 1892). * Scapania degenii Schifn. ex Müll.Frib., Lebermoose 2 (22): 497, 1915 (Müller 1915a). 60 * Scapania glaucoviridis Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 221, 1934 (Horikawa 1934). 61 60 Scapania degenii is conspeciic with Scapania brevicaulis in Potemkin (1998), but recognized by Konstantinova et al. (2009). 61 Scapania glaucoviridis was treated as conspeciic with Scapania parvitexta by Potemkin (2002), but accepted by Zuo et al. (2007a). World checklist of hornworts and liverworts 91 ** Scapania grossidens Steph. ex Müll.Frib., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 83: 146, 1905 (Müller 1905). *** Scapania hedbergii S.W.Arnell, Ark. Bot. (n.ser.) 3 (16): 556, 1956 (Arnell 1956e). *** Scapania integerrima Steph., Sp. Hepat. (Stephani) 4: 148, 1910 (Stephani 1910b). *** Scapania matveyevae Potemkin, Arctoa 9: 97, 2000 (Potemkin 2000a). *** Scapania nemorea (L.) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 160, 1963 [1964] (Grolle 1963d). Bas.: Jungermannia nemorea L., Syst. Nat., ed. 10., 2: 1337, 1759 (Linnaeus 1759). ** Scapania parvidens Steph., Hedwigia 44 (1): 15, 1904 (Stephani 1904i). 62 *** Scapania parvitexta Steph., Bull. Herb. Boissier 5 (2): 107, 1897 (Stephani 1897b). *** Scapania rigida Nees, Syn. Hepat. 1: 69, 1844 (Gottsche et al. 1844). ** sect. Scapania *** Scapania gigantea Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 1: 15, 1931 (Horikawa 1931b). ** Scapania grandiloba Steph., Sp. Hepat. (Stephani) 6: 502, 1924 (Stephani 1924). *** Scapania komagadakensis Amakawa, J. Hattori Bot. Lab. 31: 96, 1968 (Amakawa 1968a). *** Scapania obscura (Arnell et C.E.O.Jensen) Schifn., Österr. Bot. Z. 58 (10): 377, 1908 (Schifner 1908b). Bas.: Martinellius obscurus Arnell et C.E.O.Jensen, Moose Sarekgeb.: 91, 1907 (Arnell and Jensen 1907). ** Scapania paludosa (Müll.Frib.) Müll.Frib., Mitt. Bad. Bot. Vereins 4 (182/183): 287, 1902 (Müller 1902). Bas.: Scapania undulata var. paludosa Müll.Frib., Beih. Bot. Centralbl. 10 (4/5): 220, 1901 (Müller 1901b). *** Scapania ruidula Warnst., Hedwigia 63 (2): 94, 1921 (Warnstorf 1921). *** Scapania serrulata R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 539, 1974 (Schuster 1974). *** Scapania subalpina (Nees ex Lindenb.) Dumort., Recueil Observ. Jungerm.: 14, 1835 (Dumortier 1835). Bas.: Jungermannia subalpina Nees ex Lindenb., Syn. hepat. eur: 55, 1829 (Lindenberg 1829). ** Scapania subalpina var. haynesiae Frye et L.Clark, Univ. Wash. Publ. Biol. 6 (4): 638, 1946 (Frye and Clark 1946). ** Scapania subalpina var. muddiae C.D.Bird et W.S.Hong, Bryologist 83 (1): 51, 1980 (Hong 1980). *** Scapania undulata (L.) Dumort., Recueil Observ. Jungerm.: 14, 1835 (Dumortier 1835). Bas.: Jungermannia undulata L., Sp. Pl. 1: 1132, 1753 (Linnaeus 1753). ** sect. Scapaniella (H.Buch) Potemkin, J. Hattori Bot. Lab. 85: 43, 1998 (Potemkin 1998). Bas.: Scapaniella H.Buch, Scapan. N.-Eur. Sib.: 33, 1928 (Buch 1928). 62 Scapania parvidens was treated as conspeciic with Scapania parvitexta by Potemkin (2002), but accepted by Zuo et al. (2007a) and Choi et al. (2012). 92 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Scapania glaucocephala (Taylor) Austin, Bull. Torrey Bot. Club 6 (16): 85, 1876 (Austin 1876c). Bas.: Jungermannia glaucocephala Taylor, London J. Bot. 5: 277, 1846 (Taylor 1846a). *** Scapania glaucocephala var. saxicola (R.M.Schust.) Potemkin, Bryologist 102 (1): 36, 1999 (Potemkin 1999). Bas.: Scapania saxicola R.M.Schust., Amer. Midl. Naturalist 49 (2): 448, 1953 (Schuster 1953). * Scapania scapanioides (C.Massal.) Grolle, Feddes Repert. 87 (3/4): 235, 1976 (Grolle 1976a). Bas.: Jungermannia scapanioides C.Massal., Hepaticol. ven.: 64, 1879 (Massalongo 1879). 63 ** sect. Simmonsiae (R.M.Schust.) Váňa, Hentschel, Joch.Müll. et Heinrichs, PhytoKeys 10: 16, 2012 (Váňa et al. 2012a). Bas.: Scapania subsect. Simmonsiae R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 612, 1974 (Schuster 1974). *** Scapania simmonsii Bryhn et Kaal., Rep. Second Norweg. Arctic Exped. 11: 51, 1906 (Bryhn 1906). ** sect. Sphaeriferae Konstant. et Potemkin, Ann. Bot. Fenn. 31 (2): 125, 1994 (Konstantinova and Potemkin 1994). *** Scapania sphaerifera H.Buch et Tuom., Memoranda Soc. Fauna Fl. Fennica 11: 227, 1936 (Buch and Tuomikoski 1936). ** sect. Stephaniae Potemkin, J. Hattori Bot. Lab. 85: 57, 1998 (Potemkin 1998). Based on: Scapania sect. Stephaniae Amakawa et S.Hatt., J. Hattori Bot. Lab. 12: 94, 1954 (Amakawa and Hattori 1954). nom. inval. *** Scapania griithii Schifn., Österr. Bot. Z. 49 (6): 204, 1899 (Schifner 1899a). *** Scapania javanica Gottsche, Natuurk. Tijdschr. Ned.-Indië 4: 575, 1853 (Gottsche 1853). * Scapania javanica var. scabra Schifn., Arch. Hydrobiol., suppl. 21 (3/4): 397, 1955 (Schifner 1955). *** Scapania ligulata Steph., Hedwigia 44 (1): 14, 1904 (Stephani 1904i). ** Scapania ligulata subsp. stephanii (Müll.Frib.) Potemkin, Piippo et T.J.Kop., Ann. Bot. Fenn. 41 (6): 423, 2004 (Potemkin et al. 2004). Bas.: Scapania stephanii Müll.Frib., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 83: 273, 1905 (Müller 1905). ** sect. Verrucosae Potemkin, J. Hattori Bot. Lab. 85: 54, 1998 (Potemkin 1998). *** Scapania udarii S.C.Srivast. et A.Srivast., J. Indian Bot. Soc. 72: 237, 1993 (Srivastava and Srivastava 1993). *** Scapania verrucosa Heeg, Rev. Bryol. 20 (5): 81, 1893 (Heeg 1893). *** Schistochilopsis (N.Kitag.) Konstant., Arctoa 3: 125, 1994 (Konstantinova and Vasil’ev 1994). Bas.: Lophozia subg. Schistochilopsis N.Kitag., J. Hattori Bot. Lab. 28: 289, 1965 (Kitagawa 1965). 63 Scapania scapanioides is possibly conspeciic with Scapania glaucocephala (Potemkin 2002). World checklist of hornworts and liverworts 93 *** Schistochilopsis cornuta (Steph.) Konstant., Arctoa 3: 125, 1994 (Konstantinova and Vasil’ev 1994). Bas.: Schistochila cornuta Steph., Sp. Hepat. (Stephani) 4: 84, 1909 (Stephani 1909d). *** Schistochilopsis grandiretis (Lindb. ex Kaal.) Konstant., Arctoa 3: 125, 1994 (Konstantinova and Vasil’ev 1994). Bas.: Jungermannia grandiretis Lindb. ex Kaal., Nyt Mag. Naturvidensk. 33 (4/5): 322, 1893 (Kaalaas 1893b). ** Schistochilopsis hyperarctica Konstant. et L.Söderstr., Phytotaxa 162 (4): 240, 2014 (Konstantinova et al. 2014b). Based on: Lophozia hyperarctica R.M.Schust., Canad. J. Bot. 39 (4): 967, 1961 (Schuster 1961b), nom. inval. *** Schistochilopsis incisa (Schrad.) Konstant., Arctoa 3: 125, 1994 (Konstantinova and Vasil’ev 1994). Bas.: Jungermannia incisa Schrad., Syst. Samml. Crypt. Gew. 2: 5, 1797 (Schrader 1797). * Schistochilopsis nakanishii (Inoue) Konstant., Arctoa 3: 125, 1994 (Konstantinova and Vasil’ev 1994). Bas.: Lophozia nakanishii Inoue, Bull. Natl. Sci. Mus. Tokyo (n.ser.) 9 (1): 37, 1966 (Inoue 1966a). 64 ** Schistochilopsis opacifolia (Culm. ex Meyl.) Konstant., Arctoa 3: 125, 1994 (Konstantinova and Vasil’ev 1994). Bas.: Lophozia opacifolia Culm. ex Meyl., Beitr. Kryptogamenl. Schweiz 6 (4): 174, 1924 (Meylan 1924). 65 *** Schistochilopsis setosa (Mitt.) Konstant., Arctoa 3: 125, 1994 (Konstantinova and Vasil’ev 1994). Bas.: Jungermannia setosa Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 92, 1860 [1861] (Mitten 1860c). Jungermanniineae R.M.Schust. ex Stotler et Crand.-Stotl. *** Acrobolbaceae E.A.Hodgs. by L. Briscoe and J.J. Engel ** Enigmella G.A.M.Scott et K.G.Beckm., J. Bryol. 17 (2): 297, 1992 (Beckmann and Scott 1992). ** Enigmella thallina G.A.M.Scott et K.G.Beckm., J. Bryol. 17 (2): 297, 1992 (Beckmann and Scott 1992). ** Acrobolboideae R.M.Schust. ex Briscoe *** Acrobolbus Nees, Syn. Hepat. 1: 5, 1844 (Gottsche et al. 1844). ** Acrobolbus africanus (Pearson) Briscoe, Phytotaxa 202 (1): 59, 2015 (Briscoe et al. 2015). Bas.: Tylimanthus africanus Pearson, Forh. Vidensk.-Selsk. Kristiania 1887 (9): 14, 1887 (Pearson 1887b). 64 Schistochilopsis nakanishii is possibly conspeciic with Schistochilopsis incisa. 65 Schistochilopsis opacifolia is sometimes treated as a subspecies of Schistochilopsis incisa (e.g. Bisang 1991). 94 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Acrobolbus anisodontus (Hook.f. et Taylor) Briscoe, Phytotaxa 202 (1): 59, 2015 (Briscoe et al. 2015). Bas.: Jungermannia anisodonta Hook.f. et Taylor, London J. Bot. 4: 79, 1845 (Hooker and Taylor 1845). ** Acrobolbus antillanus R.M.Schust., J. Hattori Bot. Lab. 90: 143, 2001 (Schuster 2001a). ** Acrobolbus azoricus (Grolle et Perss.) Briscoe, Phytotaxa 202 (1): 59, 2015 (Briscoe et al. 2015). Bas.: Tylimanthus azoricus Grolle et Perss., Svensk Bot. Tidskr. 60 (1): 169, 1966 (Grolle and Persson 1966). *** Acrobolbus caducifolius R.M.Schust., J. Hattori Bot. Lab. 90: 154, 2001 (Schuster 2001a). *** Acrobolbus ciliatus (Mitt.) Schifn., Hepat. (Engl.-Prantl): 86, 1893 (Schifner 1893b). Bas.: Gymnanthe ciliata Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 100, 1860 [1861] (Mitten 1860c). *** Acrobolbus cinerascens (Lehm. et Lindenb.) Bastow, Pap. & Proc. Roy. Soc. Tasmania 1887: 242, 1888 (Bastow 1888). Bas.: Jungermannia cinerascens Lehm. et Lindenb., Nov. Stirp. Pug. 4: 46, 1832 (Lehmann 1832). *** Acrobolbus concinnus (Mitt.) Steph., Trans. & Proc. New Zealand Inst. 24: 399, 1892 (Colenso 1892). Bas.: Gymnanthe concinna Mitt., Bot. antarct. voy. III (Fl. Tasman. 2): 230, 1860 (Mitten 1860b). ** Acrobolbus cuneifolius (Steph.) Briscoe, Phytotaxa 202 (1): 59, 2015 (Briscoe et al. 2015). Bas.: Tylimanthus cuneifolius Steph., Bull. Herb. Boissier (sér. 2) 5 (12): 1138 (10), 1905 (Stephani 1905b). *** Acrobolbus diversifolius R.M.Schust., J. Hattori Bot. Lab. 90: 150, 2001 (Schuster 2001a). *** Acrobolbus epiphytus (Colenso) Briscoe, Phytotaxa 202 (1): 59, 2015 (Briscoe et al. 2015). Bas.: Marsupidium epiphytum Colenso, Trans. & Proc. New Zealand Inst. 21: 64, 1889 (Colenso 1889). *** Acrobolbus lavicans (J.J.Engel et Grolle) Briscoe et J.J.Engel, Phytotaxa 202 (1): 59, 2015 (Briscoe et al. 2015). Bas.: Marsupidium lavicans J.J.Engel et Grolle, J. Hattori Bot. Lab. 34: 438, 1971 (Engel and Grolle 1971). *** Acrobolbus gradsteinii (Grolle) Briscoe, Phytotaxa 202 (1): 59, 2015 (Briscoe et al. 2015). Bas.: Marsupidium gradsteinii Grolle, J. Hattori Bot. Lab. 66: 337, 1989 (Grolle 1989a). ** Acrobolbus integrifolius (A.Evans) Briscoe, Phytotaxa 202 (1): 59, 2015 (Briscoe et al. 2015). Bas.: Tylimanthus integrifolius A.Evans, Trans. Connecticut Acad. Arts 8 (15): 259, 1891 (Evans 1891). *** Acrobolbus knightii (Mitt.) Briscoe, Phytotaxa 202 (1): 59, 2015 (Briscoe et al. 2015). Bas.: Marsupidium knightii Mitt., Handb. N. Zeal. l. 2: 753, 1867 (Hooker 1867). ** Acrobolbus kunkelii (Hässel et Solari) Briscoe et J.J.Engel, Phytotaxa 202 (1): 60, 2015 (Briscoe et al. 2015). Bas.: Tylimanthus kunkelii Hässel et Solari, Darwiniana 17: 574, 1972 (Hässel and Solari 1972). *** Acrobolbus laxus (Lehm. et Lindenb.) Briscoe, Phytotaxa 202 (1): 60, 2015 (Briscoe et al. 2015). Bas.: Plagiochila laxa Lehm. et Lindenb., Sp. Hepat. (Lindenberg) 2-4: 68, 1840 (Lindenberg 1840). World checklist of hornworts and liverworts 95 *** Acrobolbus limbatus (Steph.) Briscoe et J.J.Engel, Phytotaxa 202 (1): 60, 2015 (Briscoe et al. 2015). Bas.: Tylimanthus limbatus Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 25, 1911 (Stephani 1911b). *** Acrobolbus lophocoleoides (Mitt.) Mitt., Handb. N. Zeal. l. 2: 753, 1867 (Hooker 1867). Bas.: Gymnanthe lophocoleoides Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 144, 1854 (Mitten 1854). ** Acrobolbus madeirensis (Grolle et Perss.) Briscoe, Phytotaxa 202 (1): 60, 2015 (Briscoe et al. 2015). Bas.: Tylimanthus madeirensis Grolle et Perss., Svensk Bot. Tidskr. 60 (1): 166, 1966 (Grolle and Persson 1966). ** Acrobolbus mittenii Steph., Bull. Herb. Boissier (sér. 2) 2 (5): 460 (179), 1902 (Stephani 1902f). *** Acrobolbus ochrophyllus (Hook.f. et Taylor) R.M.Schust., Rev. Bryol. Lichénol. 30 (1/2): 64, 1961 (Schuster 1961a). Bas.: Jungermannia ochrophylla Hook.f. et Taylor, London J. Bot. 3: 368, 1844 (Hooker and Taylor 1844a). *** Acrobolbus papillosus (J.J.Engel et Glenny) Briscoe, Phytotaxa 202 (1): 60, 2015 (Briscoe et al. 2015). Bas.: Marsupidium papillosum J.J.Engel et Glenny, Nova Hedwigia 87 (3/4): 289, 2008 (Engel and Glenny 2008c). *** Acrobolbus perpusillus (Colenso) Briscoe, Phytotaxa 202 (1): 60, 2015 (Briscoe et al. 2015). Bas.: Tylimanthus perpusillus Colenso, Trans. & Proc. New Zealand Inst. 19: 286, 1887 (Colenso 1887). *** Acrobolbus perpusillus var. denticulatus (J.J.Engel et Glenny) Briscoe, Phytotaxa 202 (1): 60, 2015 (Briscoe et al. 2015). Bas.: Marsupidium perpusillum var. denticulatum J.J.Engel et Glenny, Nova Hedwigia 87 (3/4): 284, 2008 (Engel and Glenny 2008c). *** Acrobolbus plagiochiloides (J.J.Engel et Glenny) Briscoe, Phytotaxa 202 (1): 60, 2015 (Briscoe et al. 2015). Bas.: Marsupidium plagiochiloides J.J.Engel et Glenny, Nova Hedwigia 87 (3/4): 284, 2008 (Engel and Glenny 2008c). *** Acrobolbus pseudosaccatus (Grolle) Briscoe, Phytotaxa 202 (1): 60, 2015 (Briscoe et al. 2015). Bas.: Tylimanthus pseudosaccatus Grolle, Nova Hedwigia 6 (3/4): 391, 1963 (Grolle 1963c). *** Acrobolbus renifolius (Hässel et Solari) Briscoe et J.J.Engel, Phytotaxa 202 (1): 60, 2015 (Briscoe et al. 2015). Bas.: Tylimanthus renifolius Hässel et Solari, Darwiniana 17: 583, 1972 (Hässel and Solari 1972). ** Acrobolbus ruwenzorensis (S.W.Arnell) Briscoe, Phytotaxa 202 (1): 60, 2015 (Briscoe et al. 2015). Bas.: Tylimanthus ruwenzorensis S.W.Arnell, Ark. Bot. (n.ser.) 3 (16): 560, 1956 (Arnell 1956e). *** Acrobolbus saccatus (Hook.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 423, 1877 (Trevisan 1877). Bas.: Jungermannia saccata Hook., Musci Exot. 1: tab. 16, 1818 (Hooker 1818). *** Acrobolbus setulosus (Mitt.) Briscoe, Phytotaxa 202 (1): 60, 2015 (Briscoe et al. 2015). Bas.: Gymnanthe setulosa Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 144, 1854 (Mitten 1854). *** Acrobolbus spinifolius R.M.Schust., J. Hattori Bot. Lab. 90: 137, 2001 (Schuster 2001a). 96 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Acrobolbus sumatranus (Schifn.) Briscoe, Phytotaxa 202 (1): 61, 2015 (Briscoe et al. 2015). Bas.: Lophozia sumatrana Schifn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 203, 1898 (Schifner 1898a). *** Acrobolbus surculosus (Nees) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 423, 1877 (Trevisan 1877). Bas.: Scapania surculosa Nees, Syn. Hepat. 1: 62, 1844 (Gottsche et al. 1844). *** Acrobolbus tenellus (Taylor) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 423, 1877 (Trevisan 1877). Bas.: Gymnanthe tenella Taylor, Nov. Stirp. Pug. 8: 1, 1844 (Lehmann 1844). *** Acrobolbus tenellus var. diversifolius (E.A.Hodgs.) Briscoe, Phytotaxa 202 (1): 61, 2015 (Briscoe et al. 2015). Bas.: Tylimanthus diversifolius E.A.Hodgs., Trans. Roy. Soc. New Zealand 85 (4): 575, 1958 (Hodgson 1958). *** Acrobolbus urvilleanus (Mont.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 423, 1877 (Trevisan 1877). Bas.: Plagiochila urvilleana Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 247, 1843 (Montagne 1843). *** Acrobolbus viridis (Mitt.) Briscoe et J.J.Engel, Phytotaxa 202 (1): 61, 2015 (Briscoe et al. 2015). Bas.: Tylimanthus viridis Mitt., J. Linn. Soc., Bot. 15 (84): 197, 1876 (Mitten 1876b). *** Acrobolbus wilsonii Nees, Syn. Hepat. 1: 5, 1844 (Gottsche et al. 1844). ** Acrobolbus wilsonii var. andinus Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 522, 1885 (Spruce 1885). * ** Excluded from the genus Acrobolbus bispinosus (J.B.Jack et Steph.) Steph., Bull. Herb. Boissier (sér. 2) 2 (5): 459 (178), 1902 (Stephani 1902f). Bas.: Tylimanthus bispinosus J.B.Jack et Steph., Hedwigia 31 (1): 26, 1892 (Jack and Stephani 1892). 66 Austrolophozioideae R.M.Schust. ex Crand.-Stotl., Váňa et Stotler ** Austrolophozia R.M.Schust., J. Hattori Bot. Lab. 26: 282, 1963 (Schuster 1963b). ** Austrolophozia andina R.M.Schust., Nova Hedwigia 15: 495, 1968 (Schuster 1968b). *** Austrolophozia camensis (Steph.) Grolle ex Hässel et Solari, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 3 (6): 240, 1970 (Hässel and Solari 1970). Bas.: Tylimanthus camensis Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 24, 1911 (Stephani 1911b). *** Austrolophozia paradoxa R.M.Schust., J. Hattori Bot. Lab. 26: 282, 1963 (Schuster 1963b). 66 Acrobolbus bispinosus is a Plagiochila species (Burghardt and Gradstein 2008). World checklist of hornworts and liverworts 97 *** Goebelobryum Grolle, J. Hattori Bot. Lab. 25: 135, 1962 (Grolle 1962b). *** Goebelobryum grossitextum (Steph.) Grolle, J. Hattori Bot. Lab. 25: 137, 1962 (Grolle 1962b). Bas.: Marsupidium grossitextum Steph., Sp. Hepat. (Stephani) 6: 446, 1924 (Stephani 1924). *** Goebelobryum unguiculatum (Hook.f. et Taylor) Grolle, J. Hattori Bot. Lab. 25: 137, 1962 (Grolle 1962b). Bas.: Jungermannia unguiculata Hook.f. et Taylor, London J. Bot. 5: 279, 1846 (Taylor 1846a). *** Goebelobryum vermiculare J.J.Engel et Glenny, Nova Hedwigia 95 (3/4): 320, 2012 (Engel and Glenny 2012). ** Lethocoleoideae Grolle *** Lethocolea Mitt., Handb. N. Zeal. l. 2: 751, 1867 (Hooker 1867) nom. conserv. ** Lethocolea congesta (Lehm.) S.W.Arnell, Bot. Not. 108: 311, 1955 (Arnell 1955b). Bas.: Jungermannia congesta Lehm., Linnaea 4: 365, 1829 (Lehmann 1829). *** Lethocolea glossophylla (Spruce) Grolle, Bot. Mag. (Tokyo) 78 (921): 83, 1965 (Grolle 1965c). Bas.: Symphyomitra glossophylla Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 503, 1885 (Spruce 1885). ** Lethocolea indica G.Asthana et Maurya, Natl. Acad. Sci. Lett. 37 (6): 535, 2014 (Asthana and Maurya 2014). *** Lethocolea javanica (Schifn.) Grolle, Bot. Mag. (Tokyo) 78 (921): 83, 1965 (Grolle 1965c). Bas.: Symphyomitra javanica Schifn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 193, 1898 (Schifner 1898a). ** Lethocolea naruto-toganensis Furuki, Bryologist 104 (2): 306, 2001 (Furuki 2001). *** Lethocolea pansa (Taylor) G.A.M.Scott et K.G.Beckm., Symp. Biol. Hung. 35: 212, 1987 (Scott and Beckmann 1987). Bas.: Jungermannia pansa Taylor, London J. Bot. 5: 275, 1846 (Taylor 1846a). *** Lethocolea radicosa (Lehm. et Lindenb.) Grolle, Bot. Mag. (Tokyo) 78 (921): 83, 1965 (Grolle 1965c). Bas.: Jungermannia radicosa Lehm. et Lindenb., Nov. Stirp. Pug. 6: 35, 1834 (Lehmann 1834). * Lethocolea repens S.Winkl., Mitt. Inst. Colombo-Alemán Invest. Ci. 3: 67, 1969 (Winkler 1969). ** Saccogynidioideae Crand.-Stotl., Váňa et Stotler *** Saccogynidium Grolle, J. Hattori Bot. Lab. 23: 43, 1960 [1961] (Grolle 1960d). ** sect. Decurvum Grolle, J. Hattori Bot. Lab. 23: 59, 1960 (Grolle 1960d). *** Saccogynidium decurvum (Mitt.) Grolle, J. Hattori Bot. Lab. 23: 59, 1960 [1961] (Grolle 1960d). Bas.: Lophocolea decurva Mitt., Bot. antarct. voy. III (Fl. Tasman. 2): 227, 1860 (Mitten 1860b). 98 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** sect. Jugata Grolle, J. Hattori Bot. Lab. 23: 55, 1960 (Grolle 1960d). ** Saccogynidium rigidulum (Nees) Grolle, J. Hattori Bot. Lab. 23: 52, 1960 [1961] (Grolle 1960d). Bas.: Jungermannia rigidula Nees, Enum. Pl. Crypt. Javae: 25, 1830 (Nees 1830). ** sect. Saccogynidium *** Saccogynidium australe (Mitt.) Grolle, J. Hattori Bot. Lab. 23: 49, 1960 [1961] (Grolle 1960d). Bas.: Saccogyna australis Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 145, 1854 (Mitten 1854). ** Saccogynidium caldense (Ångstr.) Grolle, J. Hattori Bot. Lab. 23: 44, 1960 [1961] (Grolle 1960d). Bas.: Chiloscyphus caldensis Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 33 (7): 80, 1876 [1877] (Ångström 1876). ** Saccogynidium goebelii (Herzog) Grolle, J. Hattori Bot. Lab. 23: 52, 1960 [1961] (Grolle 1960d). Bas.: Leioscyphus goebelii Herzog, Ann. Bryol. 5: 89, 1932 (Herzog 1932a). ** Saccogynidium muricellum (De Not.) Grolle, J. Hattori Bot. Lab. 36: 80, 1972 [1973] (Grolle and Schultze-Motel 1972). Bas.: Chiloscyphus muricellus De Not., Epat. Borneo: 24, 1874 (De Notaris 1874). *** Saccogynidium vasculosum (Hook.f. et Taylor) Grolle, J. Hattori Bot. Lab. 23: 46, 1960 [1961] (Grolle 1960d). Bas.: Jungermannia vasculosa Hook.f. et Taylor, London J. Bot. 3: 461, 1844 (Hooker and Taylor 1844b). Incertae sedis ** Saccogynidium chiloscyphoides R.M.Schust., J. Hattori Bot. Lab. 26: 272, 1963 (Schuster 1963b). ** Saccogynidium irregularospinum C.Gao, T.Cao et M.J.Lai, Bryologist 104 (1): 129, 2001 (Gao et al. 2001). *** Antheliaceae R.M.Schust. *** Anthelia (Dumort.) Dumort., Recueil Observ. Jungerm.: 18, 1835 (Dumortier 1835). Bas.: Jungermannia sect. Anthelia Dumort., Syll. Jungerm. Europ.: 63, 1831 (Dumortier 1831). *** Anthelia julacea (L.) Dumort., Recueil Observ. Jungerm.: 18, 1835 (Dumortier 1835). Bas.: Jungermannia julacea L., Sp. Pl., ed. 2: 1601, 1763 (Linnaeus 1763). ** Anthelia juratzkana (Limpr.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 416, 1877 (Trevisan 1877). Bas.: Jungermannia juratzkana Limpr., Hedwigia 15 (2): 18, 1876 (Limpricht 1876). World checklist of hornworts and liverworts *** 99 Arnelliaceae Nakai Crandall-Stotler et al. (2009) placed Stephaniella and Stephaniellidium in Arnelliaceae following de Roo et al. (2007). However, Váňa et al. (2012e) argued based on further molecular evidence that Arnelliaceae should be retained as a monotypic family with a single species. *** Arnellia Lindb., Kongl. Svenska Vetensk.-Akad. Handl. (n.ser.) 23 (5): 35, 1889 (Lindberg and Arnell 1889). *** Arnellia fennica (Gottsche) Lindb., Kongl. Svenska Vetensk.-Akad. Handl. (n.ser.) 23 (5): 35, 1889 (Lindberg and Arnell 1889). Bas.: Jungermannia fennica Gottsche, Hepat. Eur., Leberm. 42-44: no. 418, 1868 (Gottsche and Rabenhorst 1868). *** Balantiopsidaceae H.Buch by J.J. Engel with contribution by J. Váňa (Neesioscyphus) he placement of Pseudoisotachis in Balantiopsidaceae is only preliminary (cf. Váňa 2013). *** Acroscyphella N.Kitag. et Grolle, Acta Phytotax. Geobot. 36 (1/3): 58, 1985 (Kitagawa and Grolle 1985). Nom. nov. pro Acroscyphus N.Kitag., Acta Phytotax. Geobot. 35 (1/3): 1, 1984 (Kitagawa 1984). *** Acroscyphella iwatsukii (N.Kitag.) N.Kitag. et Grolle, Acta Phytotax. Geobot. 36 (1/3): 58, 1985 (Kitagawa and Grolle 1985). Bas.: Acroscyphus iwatsukii N.Kitag., Acta Phytotax. Geobot. 35 (1/3): 3, 1984 (Kitagawa 1984). *** Acroscyphella phoenicorhiza (Grolle) N.Kitag. et Grolle, Acta Phytotax. Geobot. 36 (1/3): 58, 1985 (Kitagawa and Grolle 1985). Bas.: Neesioscyphus phoenicorhizus Grolle, Österr. Bot. Z. 111 (1): 27, 1964 (Grolle 1964e). *** Acroscyphella tjiwideiensis (Sande Lac.) N.Kitag. et Grolle, Acta Phytotax. Geobot. 36 (1/3): 58, 1985 (Kitagawa and Grolle 1985). Bas.: Chiloscyphus tjiwideiensis Sande Lac., Ned. Kruidk. Arch. 3: 418, 1854 [1855] (Sande Lacoste 1854). ** Pseudoisotachis Váňa, Polish Bot. J. 58 (1): 55, 2013 (Váňa 2013). ** Pseudoisotachis pocsii Váňa, Polish Bot. J. 58 (1): 55, 2013 (Váňa 2013). *** Balantiopsidoideae J.J.Engel et Váňa *** Balantiopsis Mitt., Handb. N. Zeal. l. 2: 751, 1867 (Hooker 1867). *** Balantiopsis asymmetrica (Herzog) J.J.Engel, Nova Hedwigia 16: 93, 1968 (Engel 1968). Bas.: Balantiopsis latifolia var. asymmetrica Herzog, Rev. Bryol. Lichénol. 23 (1/2): 53, 1954 (Herzog 1954). 100 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Balantiopsis bisbiida (Steph.) Steph., Sp. Hepat. (Stephani) 4: 101, 1910 (Stephani 1910b). Bas.: Isotachis bisbiida Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 17): 24, 1901 (Stephani 1901b). *** Balantiopsis brasiliensis Steph., Sp. Hepat. (Stephani) 4: 104, 1910 (Stephani 1910b). *** Balantiopsis cancellata (Nees) Steph., Sp. Hepat. (Stephani) 4: 103, 1910 (Stephani 1910b). Bas.: Ptilidium cancellatum Nees, Syn. Hepat. 2: 251, 1845 (Gottsche et al. 1845a). *** Balantiopsis ciliaris S.Hatt., J. Jap. Bot. 41 (5): 129, 1966 (Hattori 1966b). ** Balantiopsis ciliaris subsp. novoguineensis S.Hatt., J. Jap. Bot. 41 (5): 131, 1966 (Hattori 1966b). *** Balantiopsis convexiuscula Berggr., New Zealand Hepat.: 44, 1898 (Berggren 1898). *** Balantiopsis crocea Herzog, Beih. Bot. Centralbl. 60B (1/2): 12, 1939 (Herzog 1939c). *** Balantiopsis diplophylla (Hook.f. et Taylor) Mitt., Handb. N. Zeal. l. 2: 753, 1867 (Hooker 1867). Bas.: Jungermannia diplophylla Hook.f. et Taylor, London J. Bot. 3: 377, 1844 (Hooker and Taylor 1844a). ** Balantiopsis diplophylla var. hockenii (Berggr.) J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 37: 11, 1997 (Engel and Smith Merrill 1997). Bas.: Balantiopsis hockenii Berggr., New Zealand Hepat.: 46, 1898 (Berggren 1898). *** Balantiopsis erinacea (Hook.f. et Taylor) Mitt., Handb. N. Zeal. l. 2: 753, 1867 (Hooker 1867). Bas.: Jungermannia erinacea Hook.f. et Taylor, London J. Bot. 3: 462, 1844 (Hooker and Taylor 1844b). *** Balantiopsis lingulata R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 11 (1): 26, 1968 (Schuster 1968a). *** Balantiopsis montana (Colenso) J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 37: 12, 1997 (Engel and Smith Merrill 1997). Bas.: Chiloscyphus montanus Colenso, Trans. & Proc. New Zealand Inst. 21: 62, 1889 (Colenso 1889). ** Balantiopsis neocaledonica Pearson, J. Linn. Soc., Bot. 46 (305): 28, 1922 (Pearson 1922b). ** Balantiopsis paucidens Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 17): 29, 1901 (Stephani 1901b). *** Balantiopsis purpurata Mitt., Rep. Challenger, Bot. 1 (3, 1): 86, 1884 (Mitten 1884b). *** Balantiopsis rosea Berggr., New Zealand Hepat.: 43, 1898 (Berggren 1898). *** Balantiopsis splendens (Steph.) J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 37: 55, 1997 (Engel and Smith Merrill 1997). Bas.: Isotachis splendens Steph., Hedwigia 34 (2): 49, 1895 (Stephani 1895c). *** Balantiopsis tumida Berggr., New Zealand Hepat.: 45, 1898 (Berggren 1898). *** Balantiopsis verrucosa J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 37: 16, 1997 (Engel and Smith Merrill 1997). World checklist of hornworts and liverworts *** 101 Isotachidoideae Grolle *** Isotachis Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 148, 1854 (Mitten 1854). *** Isotachis armata (Nees) Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 121, 1864 (Gottsche 1864). Bas.: Jungermannia armata Nees, Syn. Hepat. 1: 129, 1844 (Gottsche et al. 1844). *** Isotachis aubertii (Schwägr.) Mitt., J. Linn. Soc., Bot. 22 (146): 322, 1886 (Mitten 1886b). Bas.: Jungermannia aubertii Schwägr., Hist. Musc. Hepat. Prodr.: 19, 1814 (Schwägrichen 1814). * Isotachis boliviensis Gottsche, Sp. Hepat. (Stephani) 3: 670, 1909 (Stephani 1909a). ** Isotachis chinensis C.Gao, T.Cao et J.Sun, Bryologist 105 (4): 694, 2002 [2003] (Gao et al. 2002). ** Isotachis erythrorhiza (Lehm. et Lindenb.) Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 338, 1885 (Spruce 1885). Bas.: Jungermannia erythrorhiza Lehm. et Lindenb., Nov. Stirp. Pug. 4: 44, 1832 (Lehmann 1832). *** Isotachis fragilis Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 67, 1911 (Stephani 1911b). ** Isotachis grandis Carrington et Pearson, Proc. Linn. Soc. New South Wales (ser. 2) 2 (4): 1041, 1888 (Carrington and Pearson 1888a). *** Isotachis grossidens Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 69, 1911 (Stephani 1911b). ** Isotachis hastatistipula (Steph.) J.J.Engel, Phytotaxa 183 (4): 299, 2014 (Engel et al. 2014). Bas.: Balantiopsis hastatistipula Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 98, 1914 (Stephani and Watts 1914). * Isotachis hians Steph., Sp. Hepat. (Stephani) 3: 665, 1909 (Stephani 1909a). *** Isotachis humectata (Hook.f. et Taylor) Steph., Sp. Hepat. (Stephani) 3: 654, 1909 (Stephani 1909a). Bas.: Jungermannia humectata Hook.f. et Taylor, London J. Bot. 3: 462, 1844 (Hooker and Taylor 1844b). ** Isotachis inlata Steph., Arch. Mus. Nac. Rio de Janeiro 13: 113, 1905 (Stephani 1905c). *** Isotachis intortifolia (Hook.f. et Taylor) Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 121, 1864 (Gottsche 1864). Bas.: Jungermannia intortifolia Hook.f. et Taylor, London J. Bot. 3: 374, 1844 (Hooker and Taylor 1844a). ** Isotachis japonica Steph., Sp. Hepat. (Stephani) 3: 652, 1909 (Stephani 1909a). * Isotachis lacustris Herzog, Hedwigia 74 (2): 94, 1934 (Herzog 1934a). *** Isotachis lopezii (R.M.Schust.) Gradst., Mem. New York Bot. Gard. 84: 66, 1999 (Gradstein 1999). Bas.: Ruizanthus lopezii R.M.Schust., Phytologia 39 (4): 241, 1978 (Schuster 1978a). *** Isotachis lyallii Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 149, 1854 (Mitten 1854). *** Isotachis minima Pearson, Univ. Calif. Publ. Bot. 10 (4): 322, 1923 (Pearson 1923). *** Isotachis montana Colenso, Trans. & Proc. New Zealand Inst. 21: 68, 1889 (Colenso 1889). 102 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Isotachis multiceps (Lindenb. et Gottsche) Gottsche, Mexik. Leverm.: 105, 1863 (Gottsche 1863). Bas.: Jungermannia multiceps Lindenb. et Gottsche, Syn. Hepat. 5: 687, 1847 (Gottsche et al. 1847). 67 ** Isotachis multiceps var. fendleri Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 125, 1864 (Gottsche 1864). ** Isotachis nigella Herzog, Memoranda Soc. Fauna Fl. Fennica 27: 107, 1952 (Herzog 1952c). *** Isotachis obtusa Steph., Sp. Hepat. (Stephani) 6: 354, 1922 (Stephani 1922). ** Isotachis olivacea R.M.Schust., J. Hattori Bot. Lab. 83: 207, 1997 (Schuster and Engel 1997). *** Isotachis plicata J.J.Engel, J. Hattori Bot. Lab. 83: 210, 1997 (Schuster and Engel 1997). ** Isotachis pusilla Steph., Sp. Hepat. (Stephani) 3: 655, 1909 (Stephani 1909a). ** Isotachis riparia Rodway, Tasm. Bryoph.: 63, 1917 (Rodway 1917b). *** Isotachis serrulata (Sw.) Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 121, 1864 (Gottsche 1864). Bas.: Jungermannia serrulata Sw., Prodr. (Swartz): 143, 1788 (Swartz 1788). *** Isotachis spegazziniana C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 220, 1885 (Massalongo 1885). * Isotachis sprucei Beauverd, Sp. Hepat. (Stephani) 6: 572, 1924 (Stephani 1924). Nom. nov. pro Isotachis triida Steph., Sp. Hepat. (Stephani) 6: 356, 1922 (Stephani 1922), nom. illeg. *** Isotachis stephanii E.S.Salmon, Rev. Bryol. 28 (4): 75, 1901 (Salmon 1901). * Isotachis vexans Steph., Sp. Hepat. (Stephani) 3: 662, 1909 (Stephani 1909a). *** Isotachis westlandica (E.A.Hodgs.) R.M.Schust., Nova Hedwigia 15: 455, 1968 (Schuster 1968b). Bas.: Rhizocaulia westlandica E.A.Hodgs., Trans. Roy. Soc. New Zealand, Bot. 3 (4): 78, 1965 (Hodgson 1965). *** Neesioscyphus Grolle, Österr. Bot. Z. 111 (1): 19, 1964 (Grolle 1964e). *** Neesioscyphus allionii (Steph.) Grolle, Rev. Bryol. Lichénol. 34 (1/2): 185, 1966 (Grolle 1966c). Bas.: Isotachis allionii Steph., Sp. Hepat. (Stephani) 6: 350, 1922 (Stephani 1922). *** Neesioscyphus argillaceus (Nees) Grolle, Österr. Bot. Z. 111 (1): 24, 1964 (Grolle 1964e). Bas.: Jungermannia argillacea Nees, Fl. Bras. (Martius) 1 (1): 338, 1833 (Nees 1833a). *** Neesioscyphus bicuspidatus (Steph.) Grolle, Rev. Bryol. Lichénol. 34 (1/2): 182, 1966 (Grolle 1966c). Bas.: Isotachis bicuspidata Steph., Symb. Antill. 2: 471, 1901 (Stephani 1901f). *** Neesioscyphus carneus (Nees) Grolle, Österr. Bot. Z. 111 (1): 20, 1964 (Grolle 1964e). Bas.: Jungermannia carnea Nees, Fl. Bras. (Martius) 1 (1): 337, 1833 (Nees 1833a). *** Neesioscyphus homophyllus (Nees) Grolle, Österr. Bot. Z. 111 (2/3): 188, 1964 (Grolle 1964f). Bas.: Jungermannia homophylla Nees, Fl. Bras. (Martius) 1 (1): 336, 1833 (Nees 1833a). 67 Isotachis multiceps is sometimes treated as a Hypoisotachis, but recent molecular studies (Forrest et al. 2006, Cooper et al. 2012b) point towards inclusion in Isotachis. World checklist of hornworts and liverworts ** 103 Ruizanthoideae R.M.Schust. ex J.J.Engel et G.L.Merr. *** Ruizanthus R.M.Schust., Phytologia 39 (4): 240, 1978 (Schuster 1978a). *** Ruizanthus venezuelanus R.M.Schust., Phytologia 39 (4): 240, 1978 (Schuster 1978a). *** Blepharidophyllaceae R.M.Schust. ex J.J.Engel by J.J. Engel *** Blepharidophyllum Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 30 (5): 151, 1873 (Ångström 1873). *** Blepharidophyllum densifolium (Hook.) Ångstr. ex C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 208, 1885 (Massalongo 1885). Bas.: Jungermannia densifolia Hook., Musci Exot. 1: tab. 36, 1818 (Hooker 1818). *** Blepharidophyllum vertebrale (Gottsche) Ångstr. ex C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 208, 1885 (Massalongo 1885). Bas.: Scapania vertebralis Gottsche, Syn. Hepat. 1: 72, 1844 (Gottsche et al. 1844). *** Clandarium (Grolle) R.M.Schust., New Manual Bryol. 1: 541, 1983 [1984] (Schuster 1983a). Bas.: Blepharidophyllum subg. Clandarium Grolle, J. Hattori Bot. Lab. 28: 65, 1965 (Grolle 1965a). *** Clandarium clandestinum (Mont.) R.M.Schust., New Manual Bryol. 1: 541, 1983 [1984] (Schuster 1983a). Bas.: Plagiochila clandestina Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 247, 1843 (Montagne 1843). *** Clandarium gottscheanum (Grolle) R.M.Schust., New Manual Bryol. 1: 541, 1983 [1984] (Schuster 1983a). Bas.: Blepharidophyllum gottscheanum Grolle, J. Hattori Bot. Lab. 28: 69, 1965 (Grolle 1965a). *** Clandarium xiphophyllum (Grolle) R.M.Schust., Phytologia 56 (2): 68, 1984 (Schuster 1984). Bas.: Blepharidophyllum xiphophyllum Grolle, J. Hattori Bot. Lab. 28: 65, 1965 (Grolle 1965a). *** Calypogeiaceae Arnell by M.A.M. Renner Calypogeiaceae is shown to be monophyletic (Masuzaki et al. 2010b) and the classiication follows Schuster (2000a) and Masuzaki et al. (2010b). *** Calypogeia Raddi, Jungermanniogr. Etrusca: 31, 1818 (Raddi 1818a) nom. conserv. 104 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** subg. Asperifoliae (Warnst.) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 115, 1969 (Schuster 1969b). Bas.: Calypogeia [unranked] Asperifoliae Warnst., Bryol. Z. 1 (7): 111, 1917 (Warnstorf 1917). *** Calypogeia arguta Nees et Mont., Naturgesch. Eur. Leberm. 3: 24, 1838 (Nees 1838b). ** Calypogeia sullivantii Austin, Hepat. bor.-amer.: 19, 1873 (Austin 1873). ** *** ** ** *** *** ** *** ** ** ** *** *** *** *** ** ** ** subg. Calypogeia Calypogeia andicola Bischl., Candollea 18: 79, 1962 (Bischler 1962a). Calypogeia annabonensis Steph., Sp. Hepat. (Stephani) 6: 447, 1924 (Stephani 1924). Calypogeia azorica Bischl., Rev. Bryol. Lichénol. 37 (1): 116, 1970 (Bischler 1970). Calypogeia azurea Stotler et Crotz, Taxon 32 (1): 74, 1983 (Stotler and Crotz 1983). Calypogeia bidentula (F.Weber) Nees, Syn. Hepat. 2: 199, 1845 (Gottsche et al. 1845a). Bas.: Jungermannia bidentula F.Weber, Hist. Musc. Hepat. Prodr.: 38, 1815 (Weber 1815). Calypogeia falcata Bischl., Candollea 18: 112, 1962 (Bischler 1962c). Calypogeia issa (L.) Raddi, Jungermanniogr. Etrusca: 33, 1818 (Raddi 1818a). Bas.: Mnium issum L., Sp. Pl. 1: 1114, 1753 (Linnaeus 1753), nom. conserv. 68 Calypogeia issa subsp. neogaea R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 169, 1969 (Schuster 1969b). Calypogeia goebelii (Schifn.) Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 677 (409), 1908 (Stephani 1908d). Bas.: Kantius goebelii Schifn., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 60 (2): 260, 1893 (Schifner 1893a). Calypogeia goebelii var. siamensis N.Kitag., Beih. Nova Hedwigia 90: 165, 1988 (Kitagawa 1988). Calypogeia grandistipula (Steph.) Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 669 (401), 1908 (Stephani 1908d). Bas.: Kantius grandistipulus Steph., Hedwigia 34 (2): 52, 1895 (Stephani 1895c). Calypogeia integristipula Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 662 (394), 1908 (Stephani 1908d). Calypogeia laxa Gottsche et Lindenb., Syn. Hepat. 5: 713, 1847 (Gottsche et al. 1847). Calypogeia lechleri (Steph.) Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 680 (412), 1908 (Stephani 1908d). Bas.: Kantius lechleri Steph., Hedwigia 34 (2): 53, 1895 (Stephani 1895c). 69 Calypogeia lechleri var. densifolia (Steph.) Bischl., Candollea 18: 101, 1962 (Bischler 1962c). Bas.: Kantius densifolius Steph., Hedwigia 34 (2): 52, 1895 (Stephani 1895c). Calypogeia longifolia Steph., Sp. Hepat. (Stephani) 6: 449, 1924 (Stephani 1924). Calypogeia lophocoleoides Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 677 (409), 1908 (Stephani 1908d). 68 Calypogeia issa contains two well separated taxa (Buczkowska et al. 2011), but a formal taxonomic treatment is not yet available. 69 Calypogeia lechleri was treated as conspeciic with Calypogeia leptoloma by Bischler (1962c) with hesitation (type specimen not found). World checklist of hornworts and liverworts 105 ** Calypogeia mascarenensis Bischl., Rev. Bryol. Lichénol. 37 (1): 89, 1970 (Bischler 1970). ** Calypogeia microstipula (Steph.) Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 670 (402), 1908 (Stephani 1908d). Bas.: Kantius microstipulus Steph., Hedwigia 34 (2): 53, 1895 (Stephani 1895c). *** Calypogeia miquelii Mont. ex Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 200, 1845 (Gottsche et al. 1845a). *** Calypogeia muelleriana (Schifn.) Müll.Frib., Beih. Bot. Centralbl. 10 (4/5): 217, 1901 (Müller 1901b). Bas.: Kantius muellerianus Schifn., Sitzungsber. deutsch. naturwiss.-med. Vereins Böhmen “Lotos” Prag 48: 342, 1900 (Schifner 1900d). 70 ** Calypogeia muelleriana subsp. blomquistii R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 187, 1969 (Schuster 1969b). *** Calypogeia neesiana (C.Massal. et Carestia) Müll.Frib., Verh. Bot. Vereins Prov. Brandenburg 47: 320, 1905 (Loeske 1905). Bas.: Kantius trichomanis var. neesianus C.Massal. et Carestia, Nuovo Giorn. Bot. Ital. 12 (4): 351, 1880 (Massalongo and Carestia 1880). ** Calypogeia neesiana subsp. subalpina (Inoue) Inoue, Mem. Natl. Sci. Mus. (Tokyo) 4: 58, 1971 (Inoue 1971a). Bas.: Calypogeia subalpina Inoue, J. Jap. Bot. 37 (4): 103, 1962 (Inoue 1962b). *** Calypogeia oblata Herzog, Svensk Bot. Tidskr. 51 (1): 189, 1957 (Herzog 1957a). *** Calypogeia peruviana Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 9: 47, 1838 (Montagne 1838). *** Calypogeia rhombifolia (Spruce) Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 667 (399), 1908 (Stephani 1908d). Bas.: Kantius rhombifolius Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 413, 1885 (Spruce 1885). ** Calypogeia rhombifolia var. colombiana Bischl., Candollea 18: 104, 1962 (Bischler 1962c). *** Calypogeia sphagnicola (Arnell et J.Perss.) Warnst. et Loeske, Verh. Bot. Vereins Prov. Brandenburg 47: 320, 1905 (Loeske 1905). Bas.: Kantius sphagnicola Arnell et J.Perss., Rev. Bryol. 29 (2): 26, 1902 (Arnell 1902). 71 *** Calypogeia subintegra (Gottsche, Lindenb. et Nees) Bischl., Candollea 18: 75, 1962 (Bischler 1962a). Bas.: Calypogeia peruviana β subintegra Gottsche, Lindenb. et Nees, Syn. Hepat. 5: 712, 1847 (Gottsche et al. 1847). ** Calypogeia subintegra var. dussiana (Steph.) Bischl., Candollea 18: 77, 1962 (Bischler 1962a). Bas.: Calypogeia dussiana Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 672 (404), 1908 (Stephani 1908d). 70 Calypogeia muelleriana contains two well separated taxa (Buczkowska 2010, Buczkowska and Bączkiewicz 2011). One corresponds well to the type of the species while the name of the other is unknown. 71 Calypogeia sphagnicola is a species complex including at least three genetically and morphologically well separated taxa (Buczkowska et al. 2012b, 2012a). hey all need to be compared with types of many existing names in the genus before the correct name can be assigned. 106 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Calypogeia suecica (Arnell et J.Perss.) Müll.Frib., Beih. Bot. Centralbl. 17 (2): 224, 1904 (Müller 1904). Bas.: Kantius suecicus Arnell et J.Perss., Rev. Bryol. 29 (2): 29, 1902 (Arnell 1902). *** Calypogeia tenax (Spruce) Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 664 (396), 1908 (Stephani 1908d). Bas.: Kantius tenax Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 416, 1885 (Spruce 1885). ** Calypogeia uncinulatula Herzog, Hedwigia 67 (6): 250, 1927 (Herzog 1927). *** * ** ** ** ** ** ** * ** ** ** ** ** ** Incertae sedis Calypogeia aeruginosa Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 107, 1860 [1861] (Mitten 1860c). Calypogeia amazonica (Spruce) Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 680 (412), 1908 (Stephani 1908d). Bas.: Kantius amazonicus Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 415, 1885 (Spruce 1885). 72 Calypogeia angusta Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 663 (395), 1908 (Stephani 1908d). Calypogeia apiculata (Steph.) Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 668 (400), 1908 (Stephani 1908d). Bas.: Kantius apiculatus Steph., Hedwigia 34 (2): 51, 1895 (Stephani 1895c). Calypogeia asakawana S.Hatt. ex Inoue, J. Jap. Bot. 39 (4): 107, 1964 (Inoue 1964a). Calypogeia ceylanica S.Hatt. et Mizut., Candollea 23: 288, 1968 (Hattori 1968). Calypogeia contracta Inoue, Bull. Natl. Sci. Mus. Tokyo, B 1 (4): 139, 1975 (Inoue 1975b). Calypogeia cuspidata (Steph.) Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 669 (401), 1908 (Stephani 1908d). Bas.: Kantius cuspidatus Steph., Bull. Herb. Boissier 5 (10): 846, 1897 (Stephani 1897c). Calypogeia decurrens (Steph.) Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 675 (407), 1908 (Stephani 1908d). Bas.: Kantius decurrens Steph., Hedwigia 34 (2): 52, 1895 (Stephani 1895c). Calypogeia formosana Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 186, 1934 (Horikawa 1934). Calypogeia fujisana Inoue, Bull. Natl. Sci. Mus. Tokyo, B 1 (4): 135, 1975 (Inoue 1975b). Calypogeia granulata Inoue, J. Jap. Bot. 43 (10/11): 468, 1968 (Inoue 1968b). Calypogeia japonica Steph., Sp. Hepat. (Stephani) 6: 448, 1924 (Stephani 1924). Calypogeia khasiana Ajit P.Singh et V.Nath, Taiwania 52 (4): 320, 2007 (Singh and Nath 2007a). Calypogeia latissima Steph., Sp. Hepat. (Stephani) 6: 449, 1924 (Stephani 1924). 72 Calypogeia amazonica was treated as conspeciic with Calypogeia miquelii in Gradstein et al. (1994), but it has been accepted by later authors (e.g. Churchill et al. 2008, Söderström et al. 2013e). World checklist of hornworts and liverworts 107 *** Calypogeia lunata Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 107, 1860 [1861] (Mitten 1860c). ** Calypogeia marginella Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 106, 1860 [1861] (Mitten 1860c). ** Calypogeia obovata R.M.Schust., Phytologia 39 (4): 242, 1978 (Schuster 1978a). * Calypogeia steyermarkii Fulford, Mem. New York Bot. Gard. 11 (3): 305, 1968 (Fulford 1968). ** Calypogeia tosana (Steph.) Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 678 (410), 1908 (Stephani 1908d). Bas.: Kantius tosanus Steph., Hedwigia 34 (2): 54, 1895 (Stephani 1895c). ** Calypogeia udarii Sudipa Das et D.K.Singh, Nelumbo 53: 194, 2011 (Das and Singh 2011). ** Eocalypogeia (R.M.Schust.) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 861, 1995 (Schuster 1995a). Bas.: Metacalypogeia subg. Eocalypogeia R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 107, 1969 (Schuster 1969b). ** Eocalypogeia quelpaertensis (S.Hatt. et Inoue) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 861, 1995 (Schuster 1995a). Bas.: Metacalypogeia quelpaertensis S.Hatt. et Inoue, J. Hattori Bot. Lab. 25: 129, 1962 (Hattori et al. 1962). ** Eocalypogeia schusterana (S.Hatt. et Mizut.) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 861, 1995 (Schuster 1995a). Bas.: Metacalypogeia schusterana S.Hatt. et Mizut., Misc. Bryol. Lichenol. 4 (8): 121, 1967 (Hattori and Mizutani 1967). ** Metacalypogeia (S.Hatt.) Inoue, J. Hattori Bot. Lab. 21: 231, 1959 (Inoue 1959b). Bas.: Calypogeia subg. Metacalypogeia S.Hatt., J. Hattori Bot. Lab. 18: 83, 1957 (Hattori 1957b). *** Metacalypogeia alternifolia (Nees) Grolle, Österr. Bot. Z. 111 (2/3): 185, 1964 (Grolle 1964f). Bas.: Mastigobryum alternifolium Nees, Syn. Hepat. 2: 216, 1845 (Gottsche et al. 1845a). ** Metacalypogeia cordifolia (Steph.) Inoue, J. Hattori Bot. Lab. 21: 233, 1959 (Inoue 1959b). Bas.: Calypogeia cordifolia Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 661 (393), 1908 (Stephani 1908d). ** Mizutania Furuki et Z.Iwats., J. Hattori Bot. Lab. 67: 291, 1989 (Furuki and Iwatsuki 1989). *** Mizutania riccardioides Furuki et Z.Iwats., J. Hattori Bot. Lab. 67: 291, 1989 (Furuki and Iwatsuki 1989). *** Mnioloma Herzog, Ann. Bryol. 3: 115, 1930 (Herzog 1930a). ** subg. Caracoma (Bischl.) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 833, 1995 (Schuster 1995a). Bas.: Calypogeia subg. Caracoma Bischl., Candollea 18: 26, 1962 (Bischler 1962b). 108 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Mnioloma bolivianum (Fulford) R.M.Schust., Beih. Nova Hedwigia 118: 509, 2000 (Schuster 2000a). Bas.: Calypogeia boliviana Fulford, Mem. New York Bot. Gard. 11 (3): 291, 1968 (Fulford 1968). *** Mnioloma caespitosum (Spruce) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 839, 1995 (Schuster 1995a). Bas.: Kantius caespitosus Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 412, 1885 (Spruce 1885). *** Mnioloma cellulosum (Spreng.) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 836, 1995 (Schuster 1995a). Bas.: Jungermannia cellulosa Spreng. Syst. Veg. (ed. 16) [Sprengel] 4 (1): 232, 1827 (Sprengel 1827a). *** Mnioloma crenulatum (Bischl.) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 839, 1995 (Schuster 1995a). Bas.: Calypogeia crenulata Bischl., Candollea 18: 35, 1962 (Bischler 1962b). *** Mnioloma cyclostipum (Spruce) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 843, 1995 (Schuster 1995a). Bas.: Kantius cyclostipus Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 411, 1885 (Spruce 1885). ** Mnioloma elliottii (Steph.) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 841, 1995 (Schuster 1995a). Bas.: Calypogeia elliottii Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 663 (395), 1908 (Stephani 1908d). ** Mnioloma issistipulum (Bischl.) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 847, 1995 (Schuster 1995a). Bas.: Calypogeia issistipula Bischl., Candollea 18: 47, 1962 (Bischler 1962b). *** Mnioloma fuscum (Lehm.) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 848, 1995 (Schuster 1995a). Bas.: Jungermannia fusca Lehm., Linnaea 4: 360, 1829 (Lehmann 1829). *** Mnioloma nephrostipum (Spruce) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 847, 1995 (Schuster 1995a). Bas.: Kantius nephrostipus Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 412, 1885 (Spruce 1885). ** Mnioloma novaezelandiae J.J.Engel, Cryptog. Bryol. 27 (1): 111, 2006 (Engel 2006a). *** Mnioloma parallelogramum (Spruce) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 847, 1995 (Schuster 1995a). Bas.: Kantius parallelogramus Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 413, 1885 (Spruce 1885). ** Mnioloma retusum (Bischl.) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 839, 1995 (Schuster 1995a). Bas.: Calypogeia retusa Bischl., Candollea 18: 33, 1962 (Bischler 1962b). ** Mnioloma stamatotonum M.A.M.Renner et E.A.Br., Fieldiana, Bot. (n.ser.) 47: 173, 2008 (Renner and Brown 2008). *** Mnioloma venezuelanum (Fulford) R.M.Schust., Beih. Nova Hedwigia 118: 509, 2000 (Schuster 2000a). Bas.: Calypogeia venezuelana Fulford, Mem. New York Bot. Gard. 11 (3): 287, 1968 (Fulford 1968). ** subg. Mnioloma, Fragm. Florist. Geobot. 40 (2): 833, 1995 (Schuster 1995a). *** Mnioloma rhynchophyllum Herzog, Ann. Bryol. 3: 120, 1930 (Herzog 1930a). World checklist of hornworts and liverworts *** 109 Endogemmataceae Konstant., Vilnet et A.V.Troitsky by N.A. Konstantinova Vilnet et al. (2011) described the monotypic family Endogemmataceae based on molecular evidence after a re-evaluation of Solenostomataceae. *** Endogemma Konstant., Vilnet et A.V.Troitsky, Folia Cryptog. Estonica 48: 132, 2011 (Vilnet et al. 2011). *** Endogemma caespiticia (Lindenb.) Konstant., Vilnet et A.V.Troitsky, Folia Cryptog. Estonica 48: 132, 2011 (Vilnet et al. 2011). Bas.: Jungermannia caespiticia Lindenb., Syn. hepat. eur: 67, 1829 (Lindenberg 1829). *** Geocalycaceae H.Klinggr. Placement of Geocalycaceae in Jungermanniinae follows Shaw et al. (2015). *** Geocalyx Nees, Naturgesch. Eur. Leberm. 1: 97, 1833 (Nees 1833c). *** Geocalyx caledonicus Steph., Bull. Herb. Boissier (sér. 2) 8 (3): 205 (265), 1908 (Stephani 1908h). *** Geocalyx graveolens (Schrad.) Nees, Naturgesch. Eur. Leberm. 2: 397, 1836 (Nees 1836). Bas.: Jungermannia graveolens Schrad., Syst. Samml. Crypt. Gew. 2: 6, 1797 (Schrader 1797). ** Geocalyx lancistipulus (Steph.) S.Hatt., J. Jap. Bot. 28 (8): 234, 1953 (Hattori 1953a). Bas.: Lophocolea lancistipula Steph., Sp. Hepat. (Stephani) 6: 281, 1922 (Stephani 1922). ** Geocalyx orientalis Besch. et Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: clxxxix, 1889 [1890] (Bescherelle and Spruce 1889). *** Gymnomitriaceae H.Klinggr. by J. Váňa he treatment of the family follows Váňa et al. (2010b) with some modiications. Nardioideae is included following Vilnet et al. (2010) and Váňa et al. (2014c). Some re-arrangements in Apomarsupella, Gymnomitrion and Marsupella were done by Vilnet et al (2010) and Shaw et al. (2015). Further notes on nomenclature and taxonomy can be found in Váňa et al. (2013d). Herzogobryum and Nothogymnomitrion were removed from the family by Váňa et al. (2013e). he placement of Acrolophozia, Nanomarsupella and Paramomitrion is provisional. Inclusion of Cryptocoleopsis follows Shaw et al (2015). 110 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Acrolophozia R.M.Schust., Rev. Bryol. Lichénol. 34 (1/2): 259, 1966 (Schuster 1966b). *** Acrolophozia fuegiana R.M.Schust., Nova Hedwigia 15: 499, 1968 (Schuster 1968b). *** Acrolophozia pectinata R.M.Schust., Rev. Bryol. Lichénol. 34 (1/2): 261, 1966 (Schuster 1966b). *** Acrolophozia sulcata Hässel, J. Bryol. 11 (1): 108, 1980 (Hässel 1980). ** Nanomarsupella R.M.Schust. ex A.Hagborg, L.Söderstr. et von Konrat, Phytotaxa 112 (1): 16, 2013 (Hagborg et al. 2013). Based on: Marsupella subg. Nanomarsupella R.M.Schust., Phytologia 39 (4): 248, 1978 (Schuster 1978a). *** Nanomarsupella xenophylla (R.M.Schust.) R.M.Schust. ex A.Hagborg, L.Söderstr. et von Konrat, Phytotaxa 112 (1): 16, 2013 (Hagborg et al. 2013). Bas.: Marsupella xenophylla R.M.Schust., Phytologia 39 (4): 248, 1978 (Schuster 1978a). ** Paramomitrion R.M.Schust., J. Hattori Bot. Lab. 80: 134, 1996 (Schuster 1996a). *** Paramomitrion paradoxum R.M.Schust., J. Hattori Bot. Lab. 80: 135, 1996 (Schuster 1996a). *** Gymnomitrioideae T.Jensen ** Cryptocoleopsis Amakawa, J. Hattori Bot. Lab. 21: 274, 1959 (Amakawa 1959a). *** Cryptocoleopsis imbricata Amakawa, J. Hattori Bot. Lab. 21: 274, 1959 (Amakawa 1959a). *** Gymnomitrion Corda, Gen. hepat.: 651, 1829 (Corda 1829) nom. conserv. *** Gymnomitrion adustum Nees, Naturgesch. Eur. Leberm. 1: 120, 1833 (Nees 1833c). *** Gymnomitrion africanum (Steph.) Horik., Acta Phytotax. Geobot. 13: 212, 1943 (Horikawa 1943). Bas.: Acolea africana Steph., Sp. Hepat. (Stephani) 6: 77, 1917 (Stephani 1917a). *** Gymnomitrion alpinum (Gottsche ex Husn.) Schifn., Österr. Bot. Z. 53 (7): 280, 1903 (Schifner 1903a). Bas.: Sarcocyphos alpinus Gottsche ex Husn., Hepaticol. gall. 1: 13, 1875 (Husnot 1875). *** Gymnomitrion asperulatum R.M.Schust., Acta Acad. Ped. Agr., Sect. Biol. 24: 114, 2003 (Váňa 2003). *** Gymnomitrion atroilum Váňa, J. Hattori Bot. Lab. 41: 411, 1976 (Váňa 1976b). *** Gymnomitrion bolivianum (Steph.) Váňa, Novon 20 (2): 225, 2010 (Váňa et al. 2010c). Bas.: Anastrophyllum bolivianum Steph., Biblioth. Bot. 87 (2): 186, 1916 (Stephani 1916a). *** Gymnomitrion brevissimum (Dumort.) Warnst., Hedwigia 53 (3): 196, 1913 (Warnstorf 1913). Bas.: Acolea brevissima Dumort., Syll. Jungerm. Europ.: 76, 1831 (Dumortier 1831). World checklist of hornworts and liverworts 111 *** Gymnomitrion commutatum (Limpr.) Schifn., Magyar Bot. Lapok 13: 304, 1914 [1915] (Schifner 1914b). Bas.: Sarcocyphos commutatus Limpr., Jahresber. Schles. Ges. Vaterl. Cult. 57: 314, 1879 [1880] (Limpricht 1879). *** Gymnomitrion concinnatum (Lightf.) Corda, Gen. hepat.: 651, 1829 (Corda 1829). Bas.: Jungermannia concinnata Lightf., Fl. Scot. 2: 786, 1777 (Lightfoot 1777), nom. conserv. *** Gymnomitrion corallioides Nees, Naturgesch. Eur. Leberm. 1: 118, 1833 (Nees 1833c). *** Gymnomitrion crenatilobum Grolle, Khumbu Himal 1 (4): 278, 1966 (Grolle 1966k). *** Gymnomitrion crenulatum Gottsche ex Carrington, Trans. Bot. Soc. Edinburgh 7 (3): 444, 1863 (Carrington 1863). *** Gymnomitrion crystallocaulon (Grolle) Váňa, Crand.-Stotl. et Stotler, Syst. Bot. 40 (1): 39, 2015 (Shaw et al. 2015). Bas.: Marsupella crystallocaulon Grolle, Khumbu Himal 1 (4): 281, 1966 (Grolle 1966k). ** Gymnomitrion incompletum (Gottsche) R.M.Schust. ex Váňa, J. Hattori Bot. Lab. 40: 186, 1976 (Váňa 1976a). Bas.: Jungermannia incompleta Gottsche, Linnaea 28 (5): 551, 1856 [1857] (Gottsche 1856). *** Gymnomitrion laceratum (Steph.) Horik., Acta Phytotax. Geobot. 13: 212, 1943 (Horikawa 1943). Bas.: Acolea lacerata Steph., Sp. Hepat. (Stephani) 6: 78, 1917 (Stephani 1917a). *** Gymnomitrion miniatum Lindenb. et Gottsche, Syn. Hepat. 4: 617, 1846 (Gottsche et al. 1846). *** Gymnomitrion minutulum (Hässel) Váňa, Novon 20 (2): 225, 2010 (Váňa et al. 2010c). Bas.: Marsupella minutula Hässel, J. Bryol. 11 (1): 123, 1980 (Hässel 1980). ** Gymnomitrion moralesae Váňa, J. Hattori Bot. Lab. 48: 230, 1980 (Váňa 1980). *** Gymnomitrion mucronulatum (N.Kitag.) N.Kitag., Acta Phytotax. Geobot. 19 (2/3): 53, 1962 (Kitagawa 1962a). Bas.: Gymnomitrion concinnatum var. mucronulatum N.Kitag., Acta Phytotax. Geobot. 18 (2/3): 38, 1959 (Kitagawa 1959). ** Gymnomitrion mucrophorum R.M.Schust., Bryologist 98 (2): 243, 1995 (Schuster 1995d). *** Gymnomitrion nigrum (Grolle et Váňa) Váňa, Novon 20 (2): 225, 2010 (Váňa et al. 2010c). Bas.: Marsupella nigra Grolle et Váňa, J. Hattori Bot. Lab. 40: 186, 1976 (Váňa 1976a). *** Gymnomitrion noguchianum S.Hatt., J. Jap. Bot. 27 (2): 55, 1952 (Hattori 1952b). *** Gymnomitrion obtusilobum N.Kitag., Bull. Univ. Mus. Univ. Tokyo 8: 229, 1975 (Hattori 1975e). *** Gymnomitrion obtusum Lindb., Morgonbladet (Helsinki) 1877 (30, 6 Feb): 2, 1877 (Lindberg 1877a). *** Gymnomitrion paciicum Grolle, Trans. Brit. Bryol. Soc. 5 (1): 92, 1966 (Grolle 1966f). *** Gymnomitrion revolutum (Nees) H.Philib., Rev. Bryol. 17 (3): 34, 1890 (Philibert 1890). Bas.: Sarcocyphos revolutus Nees, Naturgesch. Eur. Leberm. 2: 419, 1836 (Nees 1836). 112 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Gymnomitrion revolutum subsp. novoguineanensis (R.M.Schust.) Váňa, Crand.-Stotl. et Stotler, Syst. Bot. 40 (1): 39, 2015 (Shaw et al. 2015). Bas.: Apomarsupella revoluta subsp. novoguineanensis R.M.Schust., J. Hattori Bot. Lab. 80: 90, 1996 (Schuster 1996a). *** Gymnomitrion rubidum (Mitt.) Váňa, Crand.-Stotl. et Stotler, Syst. Bot. 40 (1): 39, 2015 (Shaw et al. 2015). Bas.: Jungermannia rubida Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 90, 1860 [1861] (Mitten 1860c). *** Gymnomitrion setaceum Grolle et Váňa, J. Hattori Bot. Lab. 41: 411, 1976 (Váňa 1976b). *** Gymnomitrion sinense Müll.Frib., Rev. Bryol. Lichénol. 20 (1/2): 176, 1951 (Müller 1951b). ** Gymnomitrion strictum (Berggr.) R.M.Schust., J. Hattori Bot. Lab. 26: 280, 1963 (Schuster 1963b). Bas.: Cesius strictus Berggr., New Zealand Hepat.: 2, 1898 (Berggren 1898). ** Gymnomitrion strictum var. inaequale R.M.Schust., J. Hattori Bot. Lab. 80: 118, 1996 (Schuster 1996a). *** Gymnomitrion subintegrum (S.W.Arnell) Váňa, Novon 20 (2): 225, 2010 (Váňa et al. 2010c). Bas.: Marsupella subintegra S.W.Arnell, Ark. Bot. (n.ser.) 3 (16): 545, 1956 (Arnell 1956e). *** Gymnomitrion truncatoapiculatum Herzog, Hedwigia 74 (2): 81, 1934 (Herzog 1934a). *** Gymnomitrion verrucosum W.E.Nicholson, Symb. Sin. 5: 10, 1930 (Nicholson et al. 1930). *** Marsupella Dumort., Commentat. Bot. (Dumortier): 114, 1822 (Dumortier 1822). *** Marsupella alata S.Hatt. et N.Kitag., Mem. Coll. Sci. Kyoto Imp. Univ., Ser. B, Biol. 27: 79, 1960 (Kitagawa 1960b). *** Marsupella andreaeoides (Lindb.) Müll.Frib., Feddes Repert. Spec. Nov. Regni Veg. 54 (2/3): 214, 1951 (Müller 1951a). Bas.: Cesius andreaeoides Lindb., Meddel. Soc. Fauna Fl. Fenn. 14: 68, 1887 (Lindberg 1887a). *** Marsupella apiculata Schifn., Österr. Bot. Z. 53 (6): 249, 1903 (Schifner 1903b). *** Marsupella aquatica (Lindenb.) Schifn., Sitzungsber. deutsch. naturwiss.-med. Vereins Böhmen “Lotos” Prag 44 (8): 267, 1896 [1897] (Schifner 1896a). Bas.: Jungermannia emarginata var. aquatica Lindenb., Syn. hepat. eur: 75, 1829 (Lindenberg 1829). *** Marsupella arctica (Berggr.) Bryhn et Kaal., Rep. Second Norweg. Arctic Exped. 11: 26, 1906 (Bryhn 1906). Bas.: Sarcocyphos emarginatus var. arcticus Berggr., Kongl. Svenska Vetensk.-Akad. Handl. (n.ser.) 13 (7): 96, 1875 (Berggren 1875). *** Marsupella boeckii (Austin) Lindb. ex Kaal., Nyt Mag. Naturvidensk. 33 (4/5): 409, 1893 (Kaalaas 1893b). Bas.: Sarcocyphos boeckii Austin, Bull. Torrey Bot. Club 3 (3): 9, 1872 (Austin 1872). *** Marsupella bolanderi (Austin) Underw., Zoe 1 (12): 365, 1891 (Underwood 1891). Bas.: Sarcocyphos bolanderi Austin, Bull. Torrey Bot. Club 3 (3): 9, 1872 (Austin 1872). World checklist of hornworts and liverworts 113 *** Marsupella condensata (Ångstr. ex C.Hartm.) Lindb. ex Kaal., Nyt Mag. Naturvidensk. 33 (4/5): 420, 1893 (Kaalaas 1893b). Bas.: Gymnomitrion condensatum Ångstr. ex C.Hartm., Handb. Skand. l. (ed. 10): 128, 1871 (Hartman 1871). *** Marsupella disticha Steph., Bull. Herb. Boissier (sér. 2) 1 (2): 164 (25), 1901 (Stephani 1901h). *** Marsupella emarginata (Ehrh.) Dumort., Recueil Observ. Jungerm.: 24, 1835 (Dumortier 1835). Bas.: Jungermannia emarginata Ehrh., Hannover. Mag. 22 (8): 141, 1784 (Ehrhart 1784). ** Marsupella emarginata subsp. tubulosa (Steph.) N.Kitag., Mem. Coll. Sci. Kyoto Imp. Univ., Ser. B, Biol. 27: 76, 1960 (Kitagawa 1960b). Bas.: Marsupella tubulosa Steph., Bull. Herb. Boissier 5 (2): 99, 1897 (Stephani 1897b). ** Marsupella emarginata subsp. tubulosa var. apertifolia (Steph.) N.Kitag., J. Hattori Bot. Lab. 26: 89, 1963 (Kitagawa 1963b). Bas.: Marsupella apertifolia Steph., Bull. Herb. Boissier (sér. 2) 1 (2): 162 (23), 1901 (Stephani 1901h). ** Marsupella emarginata subsp. tubulosa var. patens N.Kitag., Mem. Coll. Sci. Kyoto Imp. Univ., Ser. B, Biol. 27: 77, 1960 (Kitagawa 1960b). ** Marsupella emarginata subsp. tubulosa var. tubulosa (Steph.) N.Kitag. ex Váňa et L.Söderstr., Phytotaxa 183 (4): 288, 2014 (Váňa et al. 2014c). Bas.: Marsupella tubulosa Steph., Bull. Herb. Boissier 5 (2): 99, 1897 (Stephani 1897b). *** Marsupella funckii (F.Weber et D.Mohr) Dumort., Recueil Observ. Jungerm.: 24, 1835 (Dumortier 1835). Bas.: Jungermannia funckii F.Weber et D.Mohr, Bot. Taschenb. (Weber): 422, 1807 (Weber and Mohr 1807). *** Marsupella microphylla R.M.Schust., Phytologia 39 (4): 249, 1978 (Schuster 1978a). ** Marsupella minutissima N.Kitag., Mem. Coll. Sci. Kyoto Imp. Univ., Ser. B, Biol. 27: 81, 1960 (Kitagawa 1960b). *** Marsupella neesii Sande Lac. ex Schifn., Consp. Hepat. Arch. Ind.: 70, 1898 (Schifner 1898b). Based on: Sarcocyphos neesii Nees ex Sande Lac., Ann. Mus. Bot. Lugduno-Batavi 1: 288, 1864 (Sande Lacoste 1864), nom. inval. *** Marsupella paroica R.M.Schust., Bryologist 60 (2): 145, 1957 (Schuster 1957b). *** Marsupella profunda Lindb., Rev. Bryol. 14 (2): 19, 1887 (Lindberg 1887b). *** Marsupella pseudofunckii S.Hatt., J. Hattori Bot. Lab. 4: 63, 1950 (Hattori 1950). *** Marsupella sparsifolia (Lindb.) Dumort., Bull. Soc. Roy. Bot. Belgique 13: 128, 1874 (Dumortier 1874). Bas.: Sarcocyphos sparsifolius Lindb., Not. Sällsk. Fauna Fl. Fenn. Förh. 9: 280, 1868 (Lindberg 1868b). ** Marsupella sparsifolia subsp. childii R.M.Schust., Phytotaxa 183 (4): 288, 2014 (Váňa et al. 2014c). Based on: Marsupella sparsifolia subsp. childii R.M.Schust., J. Hattori Bot. Lab. 80: 61, 1996 (Schuster 1996a), nom. inval. *** Marsupella sphacelata (Giesecke ex Lindenb.) Dumort., Recueil Observ. Jungerm.: 24, 1835 (Dumortier 1835). Bas.: Jungermannia sphacelata Giesecke ex Lindenb., Syn. hepat. eur: 76, 1829 (Lindenberg 1829). *** Marsupella spiniloba R.M.Schust. et Damsh., Phytologia 63 (5): 326, 1987 (Schuster and Damsholt 1987). 114 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Marsupella sprucei (Limpr.) Bernet, Cat. hép. Suisse: 33, 1888 (Bernet 1888). Bas.: Sarcocyphos sprucei Limpr., Flora 64 (5): 72, 1881 (Limpricht 1881). ** Marsupella stableri Spruce, Rev. Bryol. 8 (6): 96, 1881 (Spruce 1881a). *** Marsupella stoloniformis N.Kitag., J. Hattori Bot. Lab. 30: 201, 1967 (Kitagawa 1967b). ** Marsupella stoloniformis subsp. vermiformis R.M.Schust., J. Hattori Bot. Lab. 80: 72, 1996 (Schuster 1996a). *** Marsupella yakushimensis (Horik.) S.Hatt., Bull. Tokyo Sci. Mus. 11: 80, 1944 (Hattori 1944d). Bas.: Sphenolobus yakushimensis Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 156, 1934 (Horikawa 1934). *** Poeltia Grolle, Khumbu Himal 1 (4): 280, 1966 (Grolle 1966k). *** Poeltia campylata Grolle, Khumbu Himal 1 (4): 280, 1966 (Grolle 1966k). *** Prasanthus Lindb., Kongl. Svenska Vetensk.-Akad. Handl. (n.ser.) 23 (5): 62, 1889 (Lindberg and Arnell 1889). ** Prasanthus jamalicus Potemkin, Ann. Bot. Fenn. 29 (4): 319, 1992 (Potemkin 1992). *** Prasanthus suecicus (Gottsche) Lindb., Kongl. Svenska Vetensk.-Akad. Handl. (n. ser.) 23 (5): 62, 1889 (Lindberg and Arnell 1889). Bas.: Gymnomitrion suecicum Gottsche, Fl. Danica 16 (48): 20, 1871 (Lange 1871). *** Nardioideae Váňa *** Nardia Gray, Nat. Arr. Brit. Pl. 1: 694, 1821 (Gray 1821) nom. conserv. *** Nardia arnelliana Grolle, Bot. Mag. (Tokyo) 77 (914): 297, 1964 (Grolle 1964b). *** Nardia assamica (Mitt.) Amakawa, J. Hattori Bot. Lab. 26: 23, 1963 (Amakawa 1963). Bas.: Jungermannia assamica Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 91, 1860 [1861] (Mitten 1860c). *** Nardia breidleri (Limpr.) Lindb., Helsingf. Dagbl. 1880 (311, 15 Nov.): 2, 1880 (Lindberg 1880b). Bas.: Alicularia breidleri Limpr., Jahresber. Schles. Ges. Vaterl. Cult. 57: 311, 1879 [1880] (Limpricht 1879). *** Nardia compressa (Hook.) Gray, Nat. Arr. Brit. Pl. 1: 694, 1821 (Gray 1821). Bas.: Jungermannia compressa Hook., Brit. Jungermann.: tab. 58, 1813 (Hooker 1813). *** Nardia lagelliformis Inoue, J. Jap. Bot. 46 (1): 1, 1971 (Inoue 1971b). *** Nardia geoscyphus (De Not.) Lindb., Not. Sällsk. Fauna Fl. Fenn. Förh. 13: 371, 1874 (Lindberg 1874a). Bas.: Alicularia geoscyphus De Not., Mem. Reale Accad. Sci. Torino (ser. 2) 18: 486, 1859 (De Notaris 1859). * Nardia geoscyphus var. dioica Bakalin, Arctoa 18: 87, 2009 [2010] (Bakalin et al. 2009b). * Nardia geoscyphus var. suberecta (Lindb. ex Kaal.) Váňa, Phytotaxa 76 (3): 37, 2013 (Váňa et al. 2013d). Bas.: Nardia haematosticta var. suberecta Kaal., Nyt Mag. Naturvidensk. 33 (4/5): 395, 1893 (Kaalaas 1893b). World checklist of hornworts and liverworts 115 *** Nardia grollei Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 491, 2009 (Váňa and Long 2009). *** Nardia insecta Lindb., Helsingf. Dagbl. 1878 (315, 18 Nov.): 2, 1878 (Lindberg 1878). *** Nardia japonica Steph., Bull. Herb. Boissier 5 (2): 101, 1897 (Stephani 1897b). * Nardia kamtschatica Arnell et C.E.O.Jensen, Hedwigia 67 (1/2): 111, 1927 (Arnell 1927). 73 * Nardia leptocaulis C.Gao, Fl. Hepat. Chin. Boreali-Orient.: 205, 1981 (Gao and Chang 1981). *** Nardia lescurii (Austin) Underw., Bull. Illinois State Lab. Nat. Hist. 2 (1): 115, 1884 (Underwood 1884). Bas.: Alicularia lescurii Austin, Hepat. bor.-amer.: 4, 1873 (Austin 1873). ** Nardia minutifolia Furuki, Bryol. Res. 9 (3): 73, 2006 (Furuki 2006b). ** Nardia nuda (Lindenb. et Gottsche) Váňa, Folia Geobot. Phytotax. 8 (2): 193, 1973 (Váňa 1973b). Bas.: Jungermannia nuda Lindenb. et Gottsche, Syn. Hepat. 5: 668, 1847 (Gottsche et al. 1847). *** Nardia poeltii Váňa, J. Hattori Bot. Lab. 36: 73, 1972 [1973] (Váňa 1972b). *** Nardia scalaris Gray, Nat. Arr. Brit. Pl. 1: 694, 1821 (Gray 1821). * Nardia scalaris var. botryoidea (R.M.Schust.) Váňa, Phytotaxa 76 (3): 38, 2013 (Váňa et al. 2013d). Bas.: Nardia scalaris subsp. botryoidea R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 862, 1969 (Schuster 1969b). * Nardia scalaris var. harae (Amakawa) Váňa, Phytotaxa 76 (3): 38, 2013 (Váňa et al. 2013d). Bas.: Nardia harae Amakawa, J. Jap. Bot. 32 (2): 38, 1957 (Amakawa 1957c). *** Nardia subclavata (Steph.) Amakawa, J. Jap. Bot. 32 (2): 40, 1957 (Amakawa 1957c). Bas.: Jungermannia subclavata Steph., Sp. Hepat. (Stephani) 6: 93, 1917 (Stephani 1917a). *** Nardia succulenta (A.Rich.) Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 519, 1885 (Spruce 1885). Bas.: Jungermannia succulenta A.Rich., Nov. Stirp. Pug. 4: 43, 1832 (Lehmann 1832). *** Nardia unispiralis Amakawa, J. Jap. Bot. 32 (6): 167, 1957 (Amakawa 1957a). ** Gyrothyraceae R.M.Schust. Placement of Gyrothyraceae in Jungermanniinae follows Shaw et al. (2015). ** Gyrothyra M.Howe, Bull. Torrey Bot. Club 24 (4): 201, 1897 (Howe 1897a). *** Gyrothyra underwoodiana M.Howe, Bull. Torrey Bot. Club 24 (4): 202, 1897 (Howe 1897a). 73 Nardia kamtschatica may be conspeciic with Nardia assamica (Váňa 1976c), but the type specimen could not be studied. 116 *** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Harpanthaceae Arnell Davis (2004) and Hentschel et al. (2006) resolved Harpanthus as an independent lineage which is recognized here as a monogeneric family as originally construed by Arnell (1928). *** Harpanthus Nees, Naturgesch. Eur. Leberm. 2: 351, 1836 (Nees 1836). *** Harpanthus drummondii (Taylor) Grolle, Österr. Bot. Z. 112 (3): 274, 1965 (Grolle 1965g). Bas.: Chiloscyphus drummondii Taylor, London J. Bot. 5: 283, 1846 (Taylor 1846a). *** Harpanthus lotovianus (Nees) Nees, Naturgesch. Eur. Leberm. 2: 353, 1836 (Nees 1836). Bas.: Jungermannia lotoviana Nees, Flora 16 (26): 408, 1833 (Nees 1833b). *** Harpanthus scutatus (F.Weber et D.Mohr) Spruce, Trans. Bot. Soc. Edinburgh 3 (1/4): 209, 1850 (Spruce 1850). Bas.: Jungermannia scutata F.Weber et D.Mohr, Bot. Taschenb. (Weber): 408, 1807 (Weber and Mohr 1807). ** Hygrobiellaceae Konstant. et Vilnet by N. Konstantinova he family Hygrobiellaceae was validated by Konstantinova et al. (2014a). *** Hygrobiella Spruce, Cephalozia: 73, 1882 (Spruce 1882). ** Hygrobiella intermedia Bakalin et Vilnet, Pl. Syst. Evol. 300 (10): 2286, 2014 (Bakalin and Vilnet 2014). *** Hygrobiella laxifolia (Hook.) Spruce, Cephalozia: 74, 1882 (Spruce 1882). Bas.: Jungermannia laxifolia Hook., Brit. Jungermann.: tab. 59, 1813 (Hooker 1813). ** Hygrobiella squamosa Bakalin et Vilnet, Pl. Syst. Evol. 300 (10): 2286, 2014 (Bakalin and Vilnet 2014). *** Jackiellaceae R.M.Schust. by J. Váňa he monogeneric status of Jackiella is supported by a molecular study based on three loci by Hendry et al. (2007). *** Jackiella Schifn., Hep. Fl. Buitenzorg: 211, 1900 (Schifner 1900a). ** Jackiella angustifolia Herzog, Trans. Brit. Bryol. Soc. 1 (4): 296, 1950 (Herzog 1950a). ** Jackiella ceylanica Schifn. ex Steph., Bull. Herb. Boissier (sér. 2) 8 (3): 212 (272), 1908 (Stephani 1908h). World checklist of hornworts and liverworts 117 *** Jackiella curvata E.A.Hodgs. et Allison, Trans. Roy. Soc. New Zealand 85 (4): 571, 1958 (Hodgson 1958). *** Jackiella javanica Schifn., Hep. Fl. Buitenzorg: 212, 1900 (Schifner 1900a). ** Jackiella javanica var. cavifolia Schifn., Hep. Fl. Buitenzorg: 213, 1900 (Schifner 1900a). ** Jackiella javanica var. cordifolia Schifn., Hep. Fl. Buitenzorg: 213, 1900 (Schifner 1900a). ** Jackiella renifolia Schifn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 70: 218, 1900 [1901] (Schifner 1900c). ** Jackiella sinensis (W.E.Nicholson) Grolle, Österr. Bot. Z. 111 (2/3): 186, 1964 (Grolle 1964f). Bas.: Aplozia sinensis W.E.Nicholson, Symb. Sin. 5: 12, 1930 (Nicholson et al. 1930). ** Jackiella singapurensis Schifn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 70: 218, 1900 [1901] (Schifner 1900c). ** Jackiella singapurensis var. philippinensis N.Kitag., Misc. Bryol. Lichenol. 9 (1): 9, 1981 (Kitagawa 1981b). Excluded from the genus * Jackiella unica Steph., Sp. Hepat. (Stephani) 6: 318, 1922 (Stephani 1922). 74 *** Jungermanniaceae Rchb. by J. Váňa he treatment of Jungermanniaceae follows Shaw et al. (2015). Many old names in Jungermannia are still neither synonymized nor transferred and we do not know their value. Some of them may prove to be older names of currently accepted taxa. We list those doubtful names in a separate section below. ** Delavayelloideae Grolle ** Delavayella Steph., Hedwigia 33 (1): 4, 1894 (Stephani 1894a). ** Delavayella serrata Steph., Mém. Soc. Nat. Sci. Nat. Math. Cherbourg 29: 211, 1894 (Stephani 1894b). ** Delavayella serrata var. purpurea P.C.Chen, Feddes Repert. Spec. Nov. Regni Veg. 58: 38, 1955 (Chen 1955). ** Liochlaena Nees, Syn. Hepat. 2: 150, 1845 (Gottsche et al. 1845a). *** Liochlaena lanceolata Nees, Syn. Hepat. 2: 150, 1845 (Gottsche et al. 1845a). 74 Jackiella unica was identiied as a Colura species before the B herbarium burned (Grolle 1966d). 118 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Liochlaena subulata (A.Evans) Schljakov, Pečen. Mchi Sev. SSSR 4: 71, 1981 (Shliakov 1981). Bas.: Jungermannia subulata A.Evans, Trans. Connecticut Acad. Arts 8 (15): 258, 1891 (Evans 1891). ** Jungermannioideae Dumort. ** Eremonotus Lindb. et Kaal. ex Pearson, Hepat. Br. Isl. 1 (6-15): 200, 1900 (Pearson 1900). *** Eremonotus myriocarpus (Carrington) Lindb. et Kaal. ex Pearson, Hepat. Br. Isl. 1 (6-15): 201, 1900 (Pearson 1900). Bas.: Jungermannia myriocarpa Carrington, Hepat. Brit. Exsicc. Fasc. II: no. 96, 1879 (Carrington and Pearson 1879). *** Jungermannia L., Sp. Pl. 1: 1131, 1753 (Linnaeus 1753). 75 *** Jungermannia atrovirens Dumort., Syll. Jungerm. Europ.: 51, 1831 (Dumortier 1831). *** Jungermannia borealis Damsh. et Váňa, Lindbergia 4 (1/2): 5, 1977 (Damsholt and Váňa 1977). * Jungermannia erectii Ajit P.Singh et V.Nath, Hepat. Khasi Jaintia Hills: E. Himal.: 117, 2007 (Singh and Nath 2007b). 76 *** Jungermannia exsertifolia Steph., Sp. Hepat. (Stephani) 6: 86, 1917 (Stephani 1917a). ** Jungermannia exsertifolia subsp. cordifolia (Dumort.) Váňa, Folia Geobot. Phytotax. 8 (3): 268, 1973 (Váňa 1973c). Bas.: Aplozia cordifolia Dumort., Bull. Soc. Roy. Bot. Belgique 13: 59, 1874 (Dumortier 1874). *** Jungermannia gollanii Steph., Sp. Hepat. (Stephani) 6: 86, 1917 (Stephani 1917a). ** Jungermannia konstantinovae Bakalin et Vilnet, Arctoa 18: 161, 2009 [2010] (Bakalin and Vilnet 2009). *** Jungermannia ovatotrigona (Steph.) Grolle, Feddes Repert. 82 (1): 90, 1971 (Grolle 1971b). Bas.: Jamesoniella ovatotrigona Steph., Biblioth. Bot. 87 (2): 184, 1916 (Stephani 1916a). *** Jungermannia polaris Lindb., Öfvers. Kongl. Vetensk.-Akad. Förh. 23 (10): 560, 1866 [1867] (Lindberg 1866). *** Jungermannia pumila With., Arr. Brit. Pl., ed. 3, 3: 883, 1796 (Withering 1796). 75 Jungermannia was used for almost all leafy liverworts from Linnaeus (1753) and well into the 19th century. Some taxa have neither been transferred nor synonymized. As the names often are very old, they may gain priority over existing names if proven to be a synonym of something. hey are listed in the “Names in genera not currently accepted” section below. 76 Jungermannia erectii was described on the basis of sterile material and it is not certain if it belongs in Jungermannia or Solenostoma (Váňa and Long 2009). World checklist of hornworts and liverworts * * * * * * * * * * * * * * * * * * * * 119 Taxa doubtfully belonging to the genus 77 Jungermannia amentacea Bertol., Mem. Reale Accad. Sci. Ist. Bologna (ser. 2) 1: 19, 1862 (Bertoloni 1862). Jungermannia brasiliensis Raddi, Critt. Brasil.: 15, 1822 (Raddi 1822). 78 Jungermannia chinensis Osbeck, Dagb. Ostind. Resa: 221, 1757 (Osbeck 1757). 79 Jungermannia cordata Vill., Hist. Pl. Dauphiné (Villars) 3: 923, 1789 (Villars 1789). Jungermannia crenulata Schmidel, Jungerm. Char.: 20, 1760 (Schmidel 1760). Jungermannia creutzeri Kremer, Monogr. hépat. Moselle: 26, 1837 (Krémer 1837). Jungermannia digitata C.F.W.Meissn. ex Spreng. Syst. Veg. (ed. 16) [Sprengel] 4 (2): 326, 1827 (Sprengel 1827b). Jungermannia dubioides H.A.Mill., Phytologia 47 (4): 322, 1981 (Miller 1981). Nom. nov. pro Jungermannia dubia Nees, Prodr. Fl. Norfolk.: 5, 1833 (Endlicher 1833), nom. illeg. Jungermannia fernandeziana Mitt., Rep. Challenger, Bot. 1 (3, 1): 85, 1884 (Mitten 1884b). Jungermannia hexagona Schwägr., Hist. Musc. Hepat. Prodr.: 18, 1814 (Schwägrichen 1814). Jungermannia holandriana Kremer, Monogr. hépat. Moselle: 25, 1837 (Krémer 1837). Jungermannia incerta Gottsche, Abh. Naturwiss. Vereins Bremen 7: 344, 1882 (Gottsche 1882). 80 Jungermannia laterilora Hampe ex Gottsche, Mexik. Leverm.: 82, 1863 (Gottsche 1863). Jungermannia lescuriana Austin, Rep. (Annual) Regents Univ. State New York State Cab. Nat. Hist. 19: 67, 1866 (Peck 1866). 81 Jungermannia longiretis Besch. et Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: clxxxv, 1889 [1890] (Bescherelle and Spruce 1889). 82 Jungermannia mastigophora Spreng. Neue Entdeck. Planzenk. 2: 99, 1821 (Sprengel 1821). 83 Jungermannia michelii Mérat, Nouv. l. env. Paris (ed. 2) 1: 219, 1821 (Mérat 1821). 84 Jungermannia minima Scop., Fl. Carniol. (ed. 2) 2: 350, 1772 (Scopoli 1772). Jungermannia odorata With., Bot. arr. veg. Gr. Brit. 2: 693, 1776 (Withering 1776). Jungermannia peltata Schmidel, Jungerm. Char.: 14, 1760 (Schmidel 1760). 77 Jungermannia was used for almost all leafy hepatics from Linnaeus (1753) and well into the 19th century. he following names have not been studied recently and are neither accepted nor synonymized. Many of them may have priority when their identity is determined. 78 Jungermannia brasiliensis is a taxon of uncertain status. Costa (2009) could not ind any authentic material and did not know what it is. 79 Jungermannia chinensis may be conspeciic with Cephalozia connivens or Cephalozia bicuspidata. 80 Jungermannia incerta is a doubtful taxon (Grolle 1995). he type specimen was burned in B. 81 Jungermannia lescuriana is a Cephaloziella species. 82 Jungermannia longiretis is an Isotachis species. 83 Jungermannia mastigophora is probably a Mastigophora species. 84 Jungermannia michelii is a Fossombronia species. 120 * * * * * * * * * * ** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Jungermannia quadridigitata Grif., Not. pl. asiat. 2: 314, 1849 (Griith 1849). 85 Jungermannia sauteri De Not. ex Rabenh., Hedwigia 1 (20): 121, 1857 (Rabenhorst 1857). Jungermannia secunda Hampe ex Gottsche, Mexik. Leverm.: 82, 1863 (Gottsche 1863). Jungermannia stereocaulis Bory, Voy. Uranie, Bot. 4: 130, 1827 (Gaudichaud 1827). 86 Jungermannia submersa Kremer, Monogr. hépat. Moselle: 36, 1837 (Krémer 1837). Jungermannia sullivantiana Austin, Rep. (Annual) Regents Univ. State New York State Cab. Nat. Hist. 19: 66, 1866 (Peck 1866). 87 Jungermannia supina Hofm., Deutschl. Fl., heil 2 (Hofm.): 86, 1795 [1796] (Hofmann 1795). Jungermannia tenuis Ehrh., Beitr. Naturk. (Ehrhart) 4: 45, 1789 (Ehrhart 1789). 88 Jungermannia uncifolia Steph., Hedwigia 34 (2): 51, 1895 (Stephani 1895c). Jungermannia vernicosa Cass. ex Mérat, Nouv. l. env. Paris (ed. 2) 1: 221, 1821 (Mérat 1821). Mesoptychioideae R.M.Schust. *** Mesoptychia (Lindb.) A.Evans, Ottawa Naturalist 17: 15, 1903 (Evans 1903a). Bas.: Jungermannia sect. Mesoptychia Lindb., Kongl. Svenska Vetensk.-Akad. Handl. (n.ser.) 23 (5): 39, 1889 (Lindberg and Arnell 1889). *** Mesoptychia badensis (Gottsche ex Rabenh.) L.Söderstr. et Váňa, Phytotaxa 65: 52, 2012 (Váňa et al. 2012b). Bas.: Jungermannia badensis Gottsche ex Rabenh., Hepat. Eur., Leberm. 9-10: no. 95, 1859 (Rabenhorst 1859). *** Mesoptychia bantriensis (Hook.) L.Söderstr. et Váňa, Phytotaxa 65: 52, 2012 (Váňa et al. 2012b). Bas.: Jungermannia bantriensis Hook., Brit. Jungermann.: tab. 41, 1813 (Hooker 1813). ** Mesoptychia bantriensis subsp. wallischii (Ştefănuţ) L.Söderstr. et Váňa, Phytotaxa 65: 52, 2012 (Váňa et al. 2012b). Bas.: Leiocolea bantriensis subsp. wallischii Ştefănuţ, Hornwort Liverwort Romania: 21, 2008 (Ştefănuţ 2008). * Mesoptychia chichibuensis (Inoue) L.Söderstr. et Váňa, Phytotaxa 65: 53, 2012 (Váňa et al. 2012b). Bas.: Lophozia chichibuensis Inoue, J. Jap. Bot. 36 (2): 41, 1961 (Inoue 1961b). * Mesoptychia collaris (Nees) L.Söderstr. et Váňa, Phytotaxa 65: 53, 2012 (Váňa et al. 2012b). Bas.: Jungermannia collaris Nees, Fl. crypt. erlang.: xv, 1817 (Martius 1817). 85 Jungermannia quadridigitata is conspeciic with Kurzia paucilora in Schuster (1969b), but that species does not occur in India (Sharma and Srivastava 1993). It is probably some other Kurzia species. 86 Jungermannia stereocaulis is a doubtful taxon. Engel (1990b) did not ind any type specimen and did not know what it is. 87 Jungermannia sullivantiana is a Cephaloziella species. 88 Jungermannia tenuis is a Lejeunea species. World checklist of hornworts and liverworts * *** *** ** ** *** * *** *** * *** ** *** *** 121 Mesoptychia itzgeraldiae (Paton et A.R.Perry) L.Söderstr. et Váňa, Phytotaxa 65: 53, 2012 (Váňa et al. 2012b). Bas.: Leiocolea itzgeraldiae Paton et A.R.Perry, J. Bryol. 18 (3): 470, 1995 (Paton and Perry 1995). Mesoptychia gillmanii (Austin) L.Söderstr. et Váňa, Phytotaxa 65: 53, 2012 (Váňa et al. 2012b). Bas.: Jungermannia gillmanii Austin, Bull. Torrey Bot. Club 3 (3): 12, 1872 (Austin 1872). Mesoptychia heterocolpos (hed. ex Hartm.) L.Söderstr. et Váňa, Phytotaxa 65: 53, 2012 (Váňa et al. 2012b). Bas.: Jungermannia heterocolpos hed. ex Hartm., Handb. Skand. l. (ed. 3): 328, 1838 (Hartman 1838). Mesoptychia heterocolpos var. arctica (S.W.Arnell) L.Söderstr. et Váňa, Phytotaxa 65: 53, 2012 (Váňa et al. 2012b). Bas.: Leiocolea arctica S.W.Arnell, Svensk Bot. Tidskr. 44 (2): 374, 1950 (Arnell 1950). Mesoptychia heterocolpos var. harpanthoides (Bryhn et Kaal.) L.Söderstr. et Váňa, Phytotaxa 65: 53, 2012 (Váňa et al. 2012b). Bas.: Lophozia harpanthoides Bryhn et Kaal., Rep. Second Norweg. Arctic Exped. 11: 31, 1906 (Bryhn 1906). Mesoptychia igiana (S.Hatt.) L.Söderstr. et Váňa, Phytotaxa 65: 54, 2012 (Váňa et al. 2012b). Bas.: Lophozia igiana S.Hatt., J. Jap. Bot. 31 (7): 201, 1956 (Hattori 1956a). Mesoptychia mamatkulovii (Duda) L.Söderstr. et Váňa, Phytotaxa 65: 54, 2012 (Váňa et al. 2012b). Bas.: Lophozia mamatkulovii Duda, Trans. Brit. Bryol. Soc. 6 (1): 82, 1970 (Duda 1970). Mesoptychia mayebarae (S.Hatt.) L.Söderstr. et Váňa, Phytotaxa 65: 54, 2012 (Váňa et al. 2012b). Bas.: Cephalozia mayebarae S.Hatt., J. Hattori Bot. Lab. 3: 37, 1948 [1950] (Hattori 1948a). Mesoptychia morrisoncola (Horik.) L.Söderstr. et Váňa, Phytotaxa 65: 54, 2012 (Váňa et al. 2012b). Bas.: Lophozia morrisoncola Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 150, 1934 (Horikawa 1934). Mesoptychia polymorpha Stotler, Crand.-Stotl. et Bakalin, Polish Bot. J. 58 (1): 82, 2013 (Crandall-Stotler et al. 2013). Mesoptychia rutheana (Limpr.) L.Söderstr. et Váňa, Phytotaxa 65: 54, 2012 (Váňa et al. 2012b). Bas.: Jungermannia rutheana Limpr., Jahresber. Schles. Ges. Vaterl. Cult. 61: 207, 1884 (Limpricht 1884). Mesoptychia rutheana var. laxa (Schifn. ex Burrell) L.Söderstr. et Váňa, Phytotaxa 65: 54, 2012 (Váňa et al. 2012b). Bas.: Lophozia schultzii var. laxa Schifn. ex Burrell, J. Bot. 49: 217, 1911 (Burrell 1911). Mesoptychia sahlbergii (Lindb. et Arnell) A.Evans, Ottawa Naturalist 17: 15, 1903 (Evans 1903a). Bas.: Jungermannia sahlbergii Lindb. et Arnell, Kongl. Svenska Vetensk.-Akad. Handl. (n.ser.) 23 (5): 40, 1889 (Lindberg and Arnell 1889). Mesoptychia subcrispa (Herzog) L.Söderstr. et Váňa, Phytotaxa 65: 54, 2012 (Váňa et al. 2012b). Bas.: Lophozia subcrispa Herzog, Ann. Naturhist. Mus. Wien 53 (1): 362, 1942 [1943] (Herzog 1942b). 122 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Mesoptychia turbinata (Raddi) L.Söderstr. et Váňa, Phytotaxa 65: 55, 2012 (Váňa et al. 2012b). Bas.: Jungermannia turbinata Raddi, Jungermanniogr. Etrusca: 18, 1818 (Raddi 1818a). * Mesoptychia ussuriensis (Bakalin) L.Söderstr. et Váňa, Phytotaxa 65: 55, 2012 (Váňa et al. 2012b). Bas.: Leiocolea ussuriensis Bakalin, Arctoa 17: 103, 2008 [2009] (Bakalin 2008c). ** Rivulariella D.H.Wagner, Phytoneuron 2013 (10): 2, 2013 (Wagner 2013). ** Rivulariella gemmipara (A.Evans) D.H.Wagner, Phytoneuron 2013 (10): 2, 2013 (Wagner 2013). Bas.: Chiloscyphus gemmiparus A.Evans, Bryologist 41 (3): 50, 1938 (Evans 1938b). ** Notoscyphaceae Crand.-Stotl., Váňa et Stotler Shaw et al. (2015) circumscribed Notoscyphaceae as monogeneric based on molecular and morphological evidence. *** Notoscyphus Mitt., Fl. vit.: 407, 1871 [1873] (Mitten 1871). *** Notoscyphus lutescens (Lehm. et Lindenb.) Mitt., Fl. vit.: 407, 1871 [1873] (Mitten 1871). Bas.: Jungermannia lutescens Lehm. et Lindenb., Nov. Stirp. Pug. 4: 16, 1832 (Lehmann 1832). ** Saccogynaceae Heeg Shaw et al. (2015) recognized Saccogynaceae as being monogeneric. *** Saccogyna Dumort., Commentat. Bot. (Dumortier): 113, 1822 (Dumortier 1822) nom. conserv. * Saccogyna ligulata Steph., Bull. Herb. Boissier (sér. 2) 8 (3): 207 (267), 1908 (Stephani 1908h). ** Saccogyna subacuta Steph., Sp. Hepat. (Stephani) 6: 316, 1922 (Stephani 1922). *** Saccogyna viticulosa (L.) Dumort., Syll. Jungerm. Europ.: 74, 1831 (Dumortier 1831). Bas.: Jungermannia viticulosa L., Sp. Pl. 1: 1131, 1753 (Linnaeus 1753). * Excluded from the genus Saccogyna tridens Steph., Sp. Hepat. (Stephani) 6: 317, 1922 (Stephani 1922). 89 89 Saccogyna tridens is a Bazzania species, but the ainity is unclear and the type specimen was burned in B (Grolle and Piippo 1984). World checklist of hornworts and liverworts *** 123 Solenostomataceae Stotler et Crand.-Stotl. by J. Váňa Solenostomataceae was irst deined by Crandall-Stotler et al. (2009) and reined to the current circumscription in Shaw et al (2015). he placement of Aponardia is provisional following Váňa et al. (2012c) and the placement of Arctoscyphus, Cryptocolea and Diplocolea in the family is also provisional. Taxonomic and nomenclatural notes can also be found in Váňa and Long (2009) and Váňa et al. (2012d, 2013c). ** Aponardia (R.M.Schust.) Váňa, Phytotaxa 65: 46, 2012 (Váňa et al. 2012c). Bas.: Nardia subg. Aponardia R.M.Schust., Beih. Nova Hedwigia 119: 360, 2002 (Schuster 2002b). *** Aponardia huerlimannii (Váňa et Grolle) Váňa, Phytotaxa 65: 46, 2012 (Váňa et al. 2012c). Bas.: Nardia huerlimannii Váňa et Grolle, Österr. Bot. Z. 118 (3): 233, 1970 (Váňa 1970c). ** Arctoscyphus Hässel, Lindbergia 16 (4): 133, 1990 [1992] (Hässel 1990a). ** Arctoscyphus fuegiensis (C.Massal.) Hässel, Cryptog. Bryol. Lichénol. 17 (3): 164, 1996 (Hässel 1996). Bas.: Leioscyphus repens var. fuegiensis C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 212, 1885 (Massalongo 1885). ** Arctoscyphus ronsmithii Hässel, Lindbergia 16 (4): 133, 1990 [1992] (Hässel 1990a). ** Cryptocolea R.M.Schust., Amer. Midl. Naturalist 49 (2): 414, 1953 (Schuster 1953). *** Cryptocolea imbricata R.M.Schust., Amer. Midl. Naturalist 49 (2): 417, 1953 (Schuster 1953). ** Diplocolea Amakawa, J. Jap. Bot. 37 (9): 274, 1962 (Amakawa 1962). *** Diplocolea sikkimensis Amakawa, J. Jap. Bot. 37 (9): 274, 1962 (Amakawa 1962). *** Solenostoma Mitt., J. Proc. Linn. Soc., Bot. 8 (29): 51, 1864 [1865] (Mitten 1864c) nom. conserv. 90 ** subg. Eucalyx (Lindb.) Váňa, Crand.-Stotl. et Stotler, Syst. Bot. 40 (1): 38, 2015 (Shaw et al. 2015). Bas.: Nardia sect. Eucalyx Lindb., Acta Soc. Sci. Fenn. 10: 525, 1875 (Lindberg 1875). *** Solenostoma bilobum (S.Hatt. ex Amakawa) Potemkin et Nyushko, Liverworts and Hornworts of Russia 1: 286, 2009 (Potemkin and Sofronova 2009). Bas.: Plectocolea biloba S.Hatt. ex Amakawa, J. Jap. Bot. 32 (7): 216, 1957 (Amakawa 1957d). 90 Solenostoma subdivisions are following Shaw et al. (2015), but the subgeneric placement of several species is preliminary and only based on morphological characteristics. Plectocolea is synonymous with Solenostoma, but one taxon has neither been transferred nor synonymized. It is listed in the “Names in genera not currently accepted” section below. 124 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Solenostoma emarginatum (Amakawa) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 136, 2010 (Váňa et al. 2010a). Bas.: Plectocolea emarginata Amakawa, J. Jap. Bot. 33 (11): 340, 1958 (Amakawa 1958b). *** Solenostoma lagellatum (S.Hatt.) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 501, 2009 (Váňa and Long 2009). Bas.: Plectocolea lagellata S.Hatt., J. Hattori Bot. Lab. 3: 13, 1948 [1950] (Hattori 1948b). *** Solenostoma hokkaidense (Váňa) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 137, 2010 (Váňa et al. 2010a). Bas.: Jungermannia hokkaidensis Váňa, J. Hattori Bot. Lab. 35: 314, 1972 (Váňa 1972a). *** Solenostoma obovatum (Nees) C.Massal., Epat. erb. critt. ital.: 17, 1903 (Massalongo 1903). Bas.: Jungermannia obovata Nees, Naturgesch. Eur. Leberm. 1: 332, 1833 (Nees 1833c). *** Solenostoma obscurum (A.Evans) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 1013, 1969 (Schuster 1969b). Bas.: Nardia obscura A.Evans, Rhodora 21 (249): 159, 1919 (Evans 1919a). *** Solenostoma schusteranum (J.D.Godfrey et G.Godfrey) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 137, 2010 (Váňa et al. 2010a). Bas.: Jungermannia schusterana J.D.Godfrey et G.Godfrey, J. Hattori Bot. Lab. 46: 109, 1979 (Godfrey and Godfrey 1979). *** Solenostoma subtilissimum (Schifn.) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 1027, 1969 (Schuster 1969b). Bas.: Nardia subtilissima Schifn., Ann. K. K. Naturhist. Hofmus. 23: 136, 1909 (Schifner 1909b). ** subg. Metasolenostoma Váňa, Crand.-Stotl. et Stotler, Syst. Bot. 40 (1): 38, 2015 (Shaw et al. 2015). *** Solenostoma fusiforme (Steph.) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 944, 1969 (Schuster 1969b). Bas.: Nardia fusiformis Steph., Bull. Herb. Boissier 5 (2): 99, 1897 (Stephani 1897b). *** Solenostoma gracillimum (Sm.) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 972, 1969 (Schuster 1969b). Bas.: Jungermannia gracillima Sm., Engl. Bot. 32: tab. 2238, 1811 (Smith and Sowerby 1811). *** Solenostoma handelii (Schifn.) Müll.Frib., Beitr. Kryptogamenl. Schweiz 10 (2): 38, 1947 (Müller 1947). Bas.: Nardia handelii Schifn., Ann. K. K. Naturhist. Hofmus. 23: 135, 1909 (Schifner 1909b). ** Solenostoma lignicola (Schifn.) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 137, 2010 (Váňa et al. 2010a). Bas.: Nardia lignicola Schifn., Ann. K. K. Naturhist. Hofmus. 23: 137, 1909 (Schifner 1909b). *** Solenostoma limbatifolium (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 504, 2009 (Váňa and Long 2009). Bas.: Jungermannia limbatifolia Amakawa, J. Hattori Bot. Lab. 31: 112, 1968 (Amakawa 1968b). ** Solenostoma philippinense Váňa, Phytotaxa 152 (1): 43, 2013 (Váňa et al. 2013c). Based on: Solenostoma gracillimum subsp. camiguinense Bakalin, Polish Bot. J. 58 (1): 134, 2013 (Bakalin 2013). World checklist of hornworts and liverworts 125 *** Solenostoma rubrum (Gottsche) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 975, 1969 (Schuster 1969b). Bas.: Jungermannia rubra Gottsche, Bot. Gaz. 13 (5): 113, 1888 (Underwood 1888). *** Solenostoma suborbiculatum (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 508, 2009 (Váňa and Long 2009). Bas.: Jungermannia suborbiculata Amakawa, J. Hattori Bot. Lab. 31: 112, 1968 (Amakawa 1968b). ** subg. Plectocolea Mitt., J. Linn. Soc., Bot. 8 (31): 156, 1864 (Mitten 1864a). *** Solenostoma balfourii (Váňa) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 136, 2010 (Váňa et al. 2010a). Bas.: Jungermannia balfourii Váňa, Folia Geobot. Phytotax. 9 (3): 279, 1974 (Váňa 1974a). *** Solenostoma borneense (Amakawa) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 136, 2010 (Váňa et al. 2010a). Bas.: Jungermannia borneensis Amakawa, J. Hattori Bot. Lab. 33: 160, 1970 (Amakawa 1970). *** Solenostoma callithrix (Lindenb. et Gottsche) Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 486 (48), 1901 (Stephani 1901a). Bas.: Jungermannia callithrix Lindenb. et Gottsche, Syn. Hepat. 5: 673, 1847 (Gottsche et al. 1847). *** Solenostoma champawatense (S.N.Srivast. et Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 496, 2009 (Váňa and Long 2009). Bas.: Jungermannia champawatensis S.N.Srivast. et Amakawa, Proc. Natl. Acad. Sci. India, B 61 (2): 205, 1991 (Srivastava and Amakawa 1991). *** Solenostoma comatum (Nees) C.Gao, Fl. Hepat. Chin. Boreali-Orient.: 73, 1981 (Gao and Chang 1981). Bas.: Jungermannia comata Nees, Enum. Pl. Crypt. Javae: 78, 1830 (Nees 1830). *** Solenostoma comatum var. novae-guineae (Váňa) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 136, 2010 (Váňa et al. 2010a). Bas.: Jungermannia comata var. novae-guineae Váňa, J. Hattori Bot. Lab. 37: 187, 1973 (Váňa 1973a). *** Solenostoma crenuliforme (Austin) Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 494 (56), 1901 (Stephani 1901a). Bas.: Jungermannia crenuliformis Austin, Bull. Torrey Bot. Club 3 (3): 10, 1872 (Austin 1872). *** Solenostoma decolor (Schifn.) R.M.Schust. ex Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 136, 2010 (Váňa et al. 2010a). Bas.: Jungermannia decolor Schifn., Leberm., Forschungsr. Gazelle 4 (4): 10, 1890 (Schifner 1890). *** Solenostoma dusenii (Steph.) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 136, 2010 (Váňa et al. 2010a). Bas.: Nardia dusenii Steph., Hedwigia 30 (5): 209, 1891 (Stephani 1891a). *** Solenostoma erectum (Amakawa) C.Gao, Fl. Hepat. Chin. Boreali-Orient.: 66, 1981 (Gao and Chang 1981). Bas.: Plectocolea erecta Amakawa, J. Jap. Bot. 32 (10): 307, 1957 (Amakawa 1957b). * Solenostoma lagellalioides C.Gao, Fl. Hepat. Chin. Boreali-Orient.: 205, 1981 (Gao and Chang 1981). 126 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Solenostoma lavialbicans (Amakawa et Grolle) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 501, 2009 (Váňa and Long 2009). Bas.: Jungermannia lavialbicans Amakawa et Grolle, J. Hattori Bot. Lab. 31: 108, 1968 (Amakawa 1968b). *** Solenostoma fossombronioides (Austin) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 1027, 1969 (Schuster 1969b). Bas.: Jungermannia fossombronioides Austin, Proc. Acad. Nat. Sci. Philadelphia 21: 220, 1869 (Austin 1869). *** Solenostoma glaucum (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 502, 2009 (Váňa and Long 2009). Bas.: Jungermannia glauca Amakawa, Fl. E. Himalaya: 511, 1966 (Hattori 1966c). * Solenostoma gongshanense (C.Gao et J.Sun) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 502, 2009 (Váňa and Long 2009). Bas.: Jungermannia gongshanensis C.Gao et J.Sun, Bull. Bot. Res., Harbin 27 (2): 140, 2007 (Sun and Duan 2007). *** Solenostoma haskarlianum (Nees) R.M.Schust. ex Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 502, 2009 (Váňa and Long 2009). Bas.: Alicularia haskarliana Nees, Syn. Hepat. 1: 12, 1844 (Gottsche et al. 1844). *** Solenostoma hattorianum (Amakawa) Potemkin et Nyushko, Liverworts and Hornworts of Russia 1: 287, 2009 (Potemkin and Sofronova 2009). Bas.: Plectocolea hattoriana Amakawa, J. Jap. Bot. 33 (11): 341, 1958 (Amakawa 1958b). *** Solenostoma hirticalyx (Steph.) R.M.Schust. ex Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 137, 2010 (Váňa et al. 2010a). Bas.: Jungermannia hirticalyx Steph., Sp. Hepat. (Stephani) 6: 87, 1917 (Stephani 1917a). *** Solenostoma horikawanum (Amakawa) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 137, 2010 (Váňa et al. 2010a). Bas.: Plectocolea horikawana Amakawa, J. Jap. Bot. 32 (7): 219, 1957 (Amakawa 1957d). *** Solenostoma hyalinum (Lyell) Mitt., Nat. hist. Azores: 319, 1870 (Mitten 1870). Bas.: Jungermannia hyalina Lyell, Brit. Jungermann.: tab. 63, 1814 (Hooker 1814). *** Solenostoma infuscum (Mitt.) Hentschel, Pl. Syst. Evol. 268 (1/4): 152, 2007 (Hentschel et al. 2007a). Bas.: Plectocolea infusca Mitt., Trans. Linn. Soc. London, Bot. 3 (3): 196, 1891 (Mitten 1891). *** Solenostoma infuscum var. ovicalyx (Steph.) Potemkin et Sofronova, Liverworts and Hornworts of Russia 1: 288, 2009 (Potemkin and Sofronova 2009). Bas.: Solenostoma ovicalyx Steph., Sp. Hepat. (Stephani) 6: 82, 1917 (Stephani 1917a). ** Solenostoma kurilense (Bakalin) Váňa, Phytotaxa 152 (1): 40, 2013 (Váňa et al. 2013c). Bas.: Plectocolea lagellata var. kurilensis Bakalin, Arctoa 18: 90, 2009 [2010] (Bakalin et al. 2009b). * Solenostoma lixingjiangii (C.Gao et X.L.Bai) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 504, 2009 (Váňa and Long 2009). Bas.: Jungermannia lixingjiangii C.Gao et X.L.Bai, Philipp. Scientist 38: 128, 2001 (Gao and Bai 2001). *** Solenostoma marginatum (S.Hatt.) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 983, 1969 (Schuster 1969b). Bas.: Plectocolea marginata S.Hatt., J. Hattori Bot. Lab. 3: 40, 1948 [1950] (Hattori 1948a). World checklist of hornworts and liverworts 127 *** Solenostoma micranthum (Mitt.) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 137, 2010 (Váňa et al. 2010a). Bas.: Plectocolea micrantha Mitt., Fl. vit.: 405, 1871 [1873] (Mitten 1871). ** Solenostoma montanum (Steph.) Váňa, Phytotaxa 65: 44, 2012 (Váňa et al. 2012d). Bas.: Nardia montana Steph., Hedwigia 28 (3): 164, 1889 (Stephani 1889d). * Solenostoma multicarpum (C.Gao et J.Sun) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 505, 2009 (Váňa and Long 2009). Bas.: Jungermannia multicarpa C.Gao et J.Sun, Bull. Bot. Res., Harbin 27 (2): 139, 2007 (Sun and Duan 2007). ** Solenostoma nilgiriense (A.Alam, Ad.Kumar et S.C.Srivast.) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 505, 2009 (Váňa and Long 2009). Bas.: Jungermannia nilgiriensis A.Alam, Ad.Kumar et S.C.Srivast., Bull. Bot. Surv. India 49 (1/4): 220, 2007 (Alam et al. 2007). *** Solenostoma obliquifolium (Schifn.) R.M.Schust. ex Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 137, 2010 (Váňa et al. 2010a). Bas.: Nardia obliquifolia Schifn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 191, 1898 (Schifner 1898a). *** Solenostoma onraedtii (Váňa) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 137, 2010 (Váňa et al. 2010a). Bas.: Jungermannia onraedtii Váňa, Folia Geobot. Phytotax. 9 (3): 282, 1974 (Váňa 1974a). * Solenostoma orbicularifolium (Piippo ex C.Gao et Bai) Váňa, Phytotaxa 222 (2): 199, 2015 (Söderström et al. 2015d). Bas.: Jungermannia orbicularifolia Piippo ex C.Gao et Bai, Philipp. Scientist 38: 129, 2001 (Gao and Bai 2001). *** Solenostoma otianum (S.Hatt.) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 984, 1969 (Schuster 1969b). Bas.: Plectocolea otiana S.Hatt., J. Jap. Bot. 28 (6): 183, 1953 (Hattori 1953b). *** Solenostoma ovalifolium (Amakawa) Váňa, Phytotaxa 152 (1): 40, 2013 (Váňa et al. 2013c). Bas.: Plectocolea infusca var. ovalifolia Amakawa, J. Jap. Bot. 34 (4): 115, 1959 (Amakawa 1959b). *** Solenostoma paroicum (Schifn.) R.M.Schust., Amer. Midl. Naturalist 49 (2): 402, 1953 (Schuster 1953). Bas.: Nardia paroica Schifn., Lotos 58: 320, 1910 (Schifner 1910d). *** Solenostoma plagiochilaceum (Grolle) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 505, 2009 (Váňa and Long 2009). Bas.: Jungermannia plagiochilacea Grolle, J. Hattori Bot. Lab. 58: 197, 1985 (Grolle 1985b). *** Solenostoma polyrhizoides (Grolle ex Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 505, 2009 (Váňa and Long 2009). Bas.: Jungermannia polyrhizoides Grolle ex Amakawa, J. Hattori Bot. Lab. 29: 262, 1966 (Amakawa 1966). *** Solenostoma radicellosum Mitt., J. Linn. Soc., Bot. 8 (31): 156, 1864 [1865] (Mitten 1864a). *** Solenostoma renauldii (Steph.) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 137, 2010 (Váňa et al. 2010a). Bas.: Jungermannia renauldii Steph., Bull. Soc. Roy. Bot. Belgique, Compt. Rend. 30 (2): 201, 1891 [1892] (Stephani 1891b). 128 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Solenostoma rigidulum (S.Hatt.) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 983, 1969 (Schuster 1969b). Bas.: Plectocolea rigidula S.Hatt., J. Jap. Bot. 27 (2): 53, 1952 (Hattori 1952b). *** Solenostoma rosulans (Steph.) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 507, 2009 (Váňa and Long 2009). Bas.: Nardia rosulans Steph., Bull. Herb. Boissier 5 (2): 101, 1897 (Stephani 1897b). *** Solenostoma rotundatum Amakawa, J. Jap. Bot. 31 (2): 50, 1956 (Amakawa 1956). *** Solenostoma rubripunctatum (S.Hatt.) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 898, 1969 (Schuster 1969b). Bas.: Plectocolea rubripunctata S.Hatt., J. Hattori Bot. Lab. 3: 41, 1948 [1950] (Hattori 1948a). * Solenostoma rupicola (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 507, 2009 (Váňa and Long 2009). Bas.: Jungermannia rupicola Amakawa, J. Hattori Bot. Lab. 22: 23, 1960 (Amakawa 1960b). 91 *** Solenostoma sikkimense (Schifn. ex Steph.) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 508, 2009 (Váňa and Long 2009). Bas.: Jungermannia sikkimensis Schifn. ex Steph., Sp. Hepat. (Stephani) 6: 92, 1917 (Stephani 1917a). *** Solenostoma tetragonum (Lindenb.) R.M.Schust. ex Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 509, 2009 (Váňa and Long 2009). Bas.: Jungermannia tetragona Lindenb., Bot. Zeitung (Berlin) 6 (25): 462, 1848 (Meissner 1848). * Solenostoma tetragonum var. kodaikanalense A.Alam, D.Sharma et So.Yadav, Phytotaxonomy 12: 70, 2012 (Alam et al. 2012). *** Solenostoma torticalyx (Steph.) C.Gao, Fl. Hepat. Chin. Boreali-Orient.: 69, 1981 (Gao and Chang 1981). Bas.: Jungermannia torticalyx Steph., Sp. Hepat. (Stephani) 6: 94, 1917 (Stephani 1917a). *** Solenostoma truncatum (Nees) R.M.Schust. ex Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 509, 2009 (Váňa and Long 2009). Bas.: Jungermannia truncata Nees, Enum. Pl. Crypt. Javae: 29, 1830 (Nees 1830). ** Solenostoma truncatum var. setulosum (Herzog) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 510, 2009 (Váňa and Long 2009). Bas.: Plectocolea setulosa Herzog, J. Hattori Bot. Lab. 14: 33, 1955 (Herzog and Noguchi 1955). *** Solenostoma tuberculiferum (Herzog) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 138, 2010 (Váňa et al. 2010a). Bas.: Aplozia tuberculifera Herzog, Ann. Bryol. 5: 84, 1932 (Herzog 1932a). *** Solenostoma unispire (Amakawa) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 138, 2010 (Váňa et al. 2010a). Bas.: Plectocolea unispiris Amakawa, J. Jap. Bot. 29 (6): 178, 1954 (Amakawa 1954). *** Solenostoma virgatum (Mitt.) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 510, 2009 (Váňa and Long 2009). Bas.: Plectocolea virgata Mitt., Trans. Linn. Soc. London, Bot. 3 (3): 197, 1891 (Mitten 1891). 91 Solenostoma rupicola is conspeciic with Solenostoma rosulans in Váňa (1975a), but accepted by Váňa and Long (2009). World checklist of hornworts and liverworts 129 *** Solenostoma vulcanicola (Schifn.) Nyushko ex Potemkin et Sofronova, Liverworts and Hornworts of Russia 1: 289, 2009 (Potemkin and Sofronova 2009). Bas.: Nardia vulcanicola Schifn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 191, 1898 (Schifner 1898a). * Solenostoma zangmuii (C.Gao et X.L.Bai) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 510, 2009 (Váňa and Long 2009). Bas.: Jungermannia zangmuii C.Gao et X.L.Bai, Philipp. Scientist 38: 135, 2001 (Gao and Bai 2001). ** subg. Solenostoma *** Solenostoma amoenum (Lindenb. et Gottsche) R.M.Schust. ex Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 136, 2010 (Váňa et al. 2010a). Bas.: Jungermannia amoena Lindenb. et Gottsche, Syn. Hepat. 5: 674, 1847 (Gottsche et al. 1847). *** Solenostoma amplexifolium (Hampe) Váňa et Schäf.-Verw., Acta Bot. Hung. 51 (3/4): 408, 2009 (Schäfer-Verwimp and Pócs 2009). Bas.: Plagiochila amplexifolia Hampe, Nov. Stirp. Pug. 7: 6, 1838 (Lehmann 1838). ** Solenostoma appalachianum R.M.Schust. ex Bakalin, Arctoa 23: 127, 2014 (Bakalin 2014). Based on: Solenostoma appalachianum R.M.Schust., Rhodora 60 (717): 246, 1958 (Schuster 1958a), nom. inval. 92 *** Solenostoma appressifolium (Mitt.) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 494, 2009 (Váňa and Long 2009). Bas.: Jungermannia appressifolia Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 91, 1860 [1861] (Mitten 1860c). 93 *** Solenostoma appressifolium var. minor (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 495, 2009 (Váňa and Long 2009). Bas.: Jungermannia clavellata var. minor Amakawa, J. Hattori Bot. Lab. 26: 25, 1963 (Amakawa 1963). *** Solenostoma appressifolium var. nigricans (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 495, 2009 (Váňa and Long 2009). Bas.: Jungermannia decolyana var. nigricans Amakawa, J. Hattori Bot. Lab. 30: 185, 1967 (Amakawa 1967b). *** Solenostoma ariadne (Taylor) R.M.Schust. ex Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 495, 2009 (Váňa and Long 2009). Bas.: Jungermannia ariadne Taylor, Nov. Stirp. Pug. 8: 9, 1844 (Lehmann 1844). *** Solenostoma atrobrunneum (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 495, 2009 (Váňa and Long 2009). Bas.: Jungermannia atrobrunnea Amakawa, J. Hattori Bot. Lab. 30: 192, 1967 (Amakawa 1967b). *** Solenostoma atrorevolutum (Grolle ex Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 496, 2009 (Váňa and Long 2009). Bas.: Jungermannia atrorevoluta Grolle ex Amakawa, J. Hattori Bot. Lab. 29: 255, 1966 (Amakawa 1966). *** Solenostoma atrovirens Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 493 (55), 1901 (Stephani 1901a). 92 Solenostoma appalachianum is placed in synonymy with Solenostoma pyrilorum by Schuster (1969b) but Shaw et al. (2015) shows that the North American specimens difers from the East Asiatic specimens so named. 93 Solenostoma appressifolium seems to be a species complex (Shaw et al. 2015). 130 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Solenostoma baueri (Schifn.) Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 495 (57), 1901 (Stephani 1901a). Bas.: Aplozia baueri Schifn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 195, 1898 (Schifner 1898a). *** Solenostoma bengalense (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 496, 2009 (Váňa and Long 2009). Bas.: Jungermannia bengalensis Amakawa, J. Hattori Bot. Lab. 31: 112, 1968 (Amakawa 1968b). ** Solenostoma brevilorum Kashyap et R.S.Chopra, Liverworts W. Himal. 2: 85, 1932 (Kashyap and Chopra 1932). *** Solenostoma caeleste (Inoue et Váňa) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 136, 2010 (Váňa et al. 2010a). Bas.: Jungermannia caelestis Inoue et Váňa, Stud. Cryptog. Papua N. Guinea: 16, 1979 (Inoue 1979b). * Solenostoma caoi (C.Gao et X.L.Bai) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 496, 2009 (Váňa and Long 2009). Bas.: Jungermannia caoi C.Gao et X.L.Bai, Philipp. Scientist 38: 152, 2001 (Gao and Bai 2001). *** Solenostoma caucasicum (Váňa) Konstant., Arctoa 1: 123, 1992 (Konstantinova et al. 1992). Bas.: Jungermannia caucasica Váňa, Preslia 42: 96, 1970 (Váňa 1970a). *** Solenostoma chenianum (C.Gao, Y.H.Wu et Grolle) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 496, 2009 (Váňa and Long 2009). Bas.: Jungermannia cheniana C.Gao, Y.H.Wu et Grolle, Nova Hedwigia 77 (1/2): 190, 2003 (Gao et al. 2003). *** Solenostoma clavellatum Mitt. ex Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 491 (53), 1901 (Stephani 1901a). *** Solenostoma confertissimum (Nees) Schljakov, Novosti Sist. Nizš. Rast. 17: 239, 1980 (Shliakov 1980b). Bas.: Jungermannia confertissima Nees, Naturgesch. Eur. Leberm. 1: 291, 1833 (Nees 1833c). *** Solenostoma conilorum (Schifn.) Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 497 (59), 1901 (Stephani 1901a). Bas.: Jungermannia conilora Schifn., Leberm., Forschungsr. Gazelle 4 (4): 10, 1890 (Schifner 1890). *** Solenostoma crassulum (Nees et Mont.) Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 497 (59), 1901 (Stephani 1901a). Bas.: Jungermannia crassula Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 5: 54, 1836 (Nees and Montagne 1836). ** Solenostoma cryptogynum R.M.Schust. ex J.J.Engel, Novon 17 (3): 311, 2007 (Engel 2007). Based on: Solenostoma cryptogynum R.M.Schust., Beih. Nova Hedwigia 119: 380, 2002 (Schuster 2002b), nom. inval. *** Solenostoma cyclops (S.Hatt.) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 945, 1969 (Schuster 1969b). Bas.: Jungermannia cyclops S.Hatt., J. Hattori Bot. Lab. 3: 5, 1948 [1950] (Hattori 1948b). *** Solenostoma diversiclavellatum (Amakawa et Grolle) R.M.Schust. ex Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 501, 2009 (Váňa and Long 2009). Bas.: Jungermannia diversiclavellata Amakawa et Grolle, J. Hattori Bot. Lab. 31: 107, 1968 (Amakawa 1968b). *** Solenostoma dulongense Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 497, 2009 (Váňa and Long 2009). World checklist of hornworts and liverworts 131 *** Solenostoma exsertum (A.Evans) Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 490 (52), 1901 (Stephani 1901a). Bas.: Nardia exserta A.Evans, Trans. Connecticut Acad. Arts 8 (15): 259, 1891 (Evans 1891). *** Solenostoma faurieanum (Beauverd) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 945, 1969 (Schuster 1969b). Bas.: Jungermannia faurieana Beauverd, Sp. Hepat. (Stephani) 6: 571, 1924 (Stephani 1924). *** Solenostoma lagellare (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 501, 2009 (Váňa and Long 2009). Bas.: Jungermannia lagellaris Amakawa, J. Hattori Bot. Lab. 29: 258, 1966 (Amakawa 1966). *** Solenostoma lavorevolutum (Váňa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 501, 2009 (Váňa and Long 2009). Bas.: Jungermannia lavorevoluta Váňa, J. Hattori Bot. Lab. 36: 63, 1972 [1973] (Váňa 1972b). *** Solenostoma grollei (D.G.Long et Váňa) K.Feldberg, Hentschel, Bombosch, D.G.Long, Váňa et Heinrichs, Pl. Syst. Evol. 280 (3/4): 244, 2009 (Feldberg et al. 2009). Bas.: Gottschelia grollei D.G.Long et Váňa, J. Bryol. 29 (3): 167, 2007 (Long and Váňa 2007). *** Solenostoma grosseverrucosum (Amakawa et S.Hatt.) Váňa, Crand.-Stotl. et Stotler, Syst. Bot. 40 (1): 39, 2015 (Shaw et al. 2015). Bas.: Horikawaella grosseverrucosa Amakawa et S.Hatt., Bull. Univ. Mus. Univ. Tokyo 8: 216, 1975 (Hattori 1975e). *** Solenostoma heterolimbatum (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 503, 2009 (Váňa and Long 2009). Bas.: Jungermannia heterolimbata Amakawa, J. Hattori Bot. Lab. 30: 183, 1967 (Amakawa 1967b). *** Solenostoma hewsoniae (Amakawa et Grolle) R.M.Schust. ex Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 136, 2010 (Váňa et al. 2010a). Bas.: Jungermannia hewsoniae Amakawa et Grolle, J. Hattori Bot. Lab. 31: 108, 1968 (Amakawa 1968b). *** Solenostoma hiugaense Amakawa, J. Jap. Bot. 31 (2): 47, 1956 (Amakawa 1956). *** Solenostoma indrodayanum (Sushil K.Singh et D.K.Singh) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 503, 2009 (Váňa and Long 2009). Bas.: Jungermannia indrodayana Sushil K.Singh et D.K.Singh, Cryptog. Bryol. 28 (2): 103, 2007 (Singh and Singh 2007b). *** Solenostoma inundatum (Hook.f. et Taylor) Mitt. ex Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 490 (52), 1901 (Stephani 1901a). Bas.: Jungermannia inundata Hook.f. et Taylor, London J. Bot. 3: 559, 1844 (Hooker and Taylor 1844d). *** Solenostoma javanicum (Schifn.) Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 494 (56), 1901 (Stephani 1901a). Bas.: Aplozia javanica Schifn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 193, 1898 (Schifner 1898a). ** Solenostoma kanaii (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 503, 2009 (Váňa and Long 2009). Bas.: Jungermannia kanaii Amakawa, J. Hattori Bot. Lab. 30: 194, 1967 (Amakawa 1967b). *** Solenostoma kashyapii (S.C.Srivast., S.Srivast. et D.Sharma) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 503, 2009 (Váňa and Long 2009). Bas.: Jungermannia 132 *** *** *** * * *** *** * *** *** *** *** *** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) kashyapii S.C.Srivast., S.Srivast. et D.Sharma, Lindbergia 28 (3): 131, 2003 (Srivastava et al. 2003). Solenostoma lanigerum (Mitt.) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 503, 2009 (Váňa and Long 2009). Bas.: Jungermannia lanigera Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 91, 1860 [1861] (Mitten 1860c). Solenostoma macrocarpum (Schifn. ex Steph.) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 504, 2009 (Váňa and Long 2009). Bas.: Jungermannia macrocarpa Schifn. ex Steph., Sp. Hepat. (Stephani) 6: 87, 1917 (Stephani 1917a). Solenostoma mamatkulovii (Váňa et Zerov) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 137, 2010 (Váňa et al. 2010a). Bas.: Jungermannia mamatkulovii Váňa et Zerov, Preslia 49: 181, 1977 (Váňa 1977). Solenostoma microphyllum C.Gao, Fl. Hepat. Chin. Boreali-Orient.: 206, 1981 (Gao and Chang 1981). Solenostoma microrevolutum (C.Gao et X.L.Bai) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 505, 2009 (Váňa and Long 2009). Bas.: Jungermannia microrevoluta C.Gao et X.L.Bai, Philipp. Scientist 38: 146, 2001 (Gao and Bai 2001). Solenostoma mildbraedii (Steph.) R.M.Schust., Beih. Nova Hedwigia 119: 387, 2002 (Schuster 2002b). Bas.: Jungermannia mildbraedii Steph., Wiss. Ergebn. Deut. Zentr.-Afr. Exped. (1907-08), Bot. 2: 113, 1911 (Stephani 1911a). Solenostoma niveum (Grolle) R.M.Schust. ex Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 137, 2010 (Váňa et al. 2010a). Bas.: Jungermannia nivea Grolle, Misc. Bryol. Lichenol. 6 (1): 1, 1971 (Grolle 1971c). Solenostoma novazelandiae R.M.Schust., Beih. Nova Hedwigia 119: 380, 2002 (Schuster 2002b). 94 Solenostoma ohbae (Amakawa) C.Gao, Bryol. Xizang: 495, 1985 (Li 1985). Bas.: Jungermannia ohbae Amakawa, Bull. Univ. Mus. Univ. Tokyo 8: 218, 1975 (Hattori 1975e). Solenostoma orbiculatum (Colenso) R.M.Schust., Beih. Nova Hedwigia 119: 380, 2002 (Schuster 2002b). Bas.: Gymnomitrion orbiculatum Colenso, Trans. & Proc. New Zealand Inst. 18: 236, 1886 (Colenso 1886b). Solenostoma parvitextum (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 505, 2009 (Váňa and Long 2009). Bas.: Jungermannia parvitexta Amakawa, J. Hattori Bot. Lab. 30: 187, 1967 (Amakawa 1967b). Solenostoma patoniae (Grolle, D.B.Schill et D.G.Long) K.Feldberg, Hentschel, Bombosch, D.G.Long, Váňa et Heinrichs, Pl. Syst. Evol. 280 (3/4): 244, 2009 (Feldberg et al. 2009). Bas.: Gottschelia patoniae Grolle, D.B.Schill et D.G.Long, J. Bryol. 25 (1): 3, 2003 (Grolle et al. 2003). Solenostoma pocsii (Váňa) Bakalin, Arctoa 18: 160, 2009 [2010] (Bakalin and Vilnet 2009). Bas.: Jungermannia pocsii Váňa, Folia Geobot. Phytotax. 10 (4): 365, 1975 (Váňa 1975b). 94 Solenostoma novaezelandiae is possibly conspeciic with Solenostoma orbiculatum (Engel and Glenny 2008a). World checklist of hornworts and liverworts 133 *** Solenostoma poeltii (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 505, 2009 (Váňa and Long 2009). Bas.: Jungermannia poeltii Amakawa, J. Hattori Bot. Lab. 29: 258, 1966 (Amakawa 1966). *** Solenostoma pseudocyclops (Inoue) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 506, 2009 (Váňa and Long 2009). Bas.: Jungermannia pseudocyclops Inoue, Bull. Natl. Sci. Mus. Tokyo (n.ser.) 9 (1): 37, 1966 (Inoue 1966a). ** Solenostoma pseudopyrilorum Bakalin et Vilnet, Arctoa 18: 159, 2009 [2010] (Bakalin and Vilnet 2009). *** Solenostoma purpuratum (Mitt.) Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 489 (51), 1901 (Stephani 1901a). Bas.: Jungermannia purpurata Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 91, 1860 [1861] (Mitten 1860c). *** Solenostoma pyrilorum Steph., Sp. Hepat. (Stephani) 6: 83, 1917 (Stephani 1917a). 95 *** Solenostoma pyrilorum var. gracillimum (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 506, 2009 (Váňa and Long 2009). Bas.: Jungermannia pyrilora var. gracillima Amakawa, Fl. E. Himalaya 2: 228, 1971 (Hattori 1971a). ** Solenostoma pyrilorum var. major (S.Hatt.) Bakalin, Arctoa 17: 230, 2008 [2009] (Bakalin 2008a). Bas.: Jungermannia monticola f. major S.Hatt., J. Hattori Bot. Lab. 3: 8, 1948 [1950] (Hattori 1948b). *** Solenostoma pyrilorum var. minutissimum (Amakawa) Bakalin, Arctoa 16: 208, 2007 [2008] (Bakalin 2007). Bas.: Jungermannia pyrilora var. minutissima Amakawa, J. Hattori Bot. Lab. 22: 61, 1960 (Amakawa 1960b). 96 *** Solenostoma raujeanum (Grolle ex Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 507, 2009 (Váňa and Long 2009). Bas.: Jungermannia raujeana Grolle ex Amakawa, J. Hattori Bot. Lab. 29: 262, 1966 (Amakawa 1966). *** Solenostoma ricleii Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 507, 2009 (Váňa and Long 2009). Nom. nov. pro Jungermannia grollei Amakawa, J. Hattori Bot. Lab. 29: 260, 1966 (Amakawa 1966). *** Solenostoma sanguinolentum (Grif.) Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 489 (51), 1901 (Stephani 1901a). Bas.: Jungermannia sanguinolenta Grif., Not. pl. asiat. 2: 302, 1849 (Griith 1849). *** Solenostoma schaulianum (Steph.) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 508, 2009 (Váňa and Long 2009). Bas.: Jungermannia schauliana Steph., Sp. Hepat. (Stephani) 6: 90, 1917 (Stephani 1917a). *** Solenostoma shimizuanum (S.Hatt. ex Váňa) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 138, 2010 (Váňa et al. 2010a). Bas.: Jungermannia shimizuana S.Hatt. ex Váňa, J. Hattori Bot. Lab. 35: 315, 1972 (Váňa 1972a). *** Solenostoma speciosum (Horik.) Hentschel, K.Feldberg, Bombosch, D.G.Long, Váňa et Heinrichs, Pl. Syst. Evol. 280 (3/4): 244, 2009 (Feldberg et al. 2009). Bas.: Anastrophyllum speciosum Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 147, 1934 (Horikawa 1934). 95 Solenostoma pyrilorum is a species complex (Shaw et al. 2015). 96 Solenostoma pyrilorum var. minutissimum was treated as a separate species, Solenostoma rishiriense Amakawa by Bakalin and Vilnet (2009). 134 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Solenostoma speciosum subsp. villosum (R.M.Schust.) Hentschel, K.Feldberg, Bombosch, D.G.Long, Váňa et Heinrichs, Pl. Syst. Evol. 280 (3/4): 246, 2009 (Feldberg et al. 2009). Bas.: Scaphophyllum speciosum subsp. villosum R.M.Schust., Bryologist 101 (3): 434, 1998 (Schuster 1998b). *** Solenostoma sphaerocarpum (Hook.) Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 499 (61), 1901 (Stephani 1901a). Bas.: Jungermannia sphaerocarpa Hook., Brit. Jungermann.: tab. 74, 1815 (Hooker 1815). *** Solenostoma stephanii (Schifn.) Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 496 (58), 1901 (Stephani 1901a). Bas.: Aplozia stephanii Schifn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 195, 1898 (Schifner 1898a). *** Solenostoma stoloniferum (Steph.) S.W.Arnell, Hepat. South Africa: 316, 1963 (Arnell 1963b). Bas.: Nardia stolonifera Steph., Hedwigia 31 (3): 128, 1892 (Stephani 1892d). *** Solenostoma strictum (Schifn.) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 138, 2010 (Váňa et al. 2010a). Bas.: Aplozia stricta Schifn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 194, 1898 (Schifner 1898a). *** Solenostoma subacutum (Herzog) Váňa, Crand.-Stotl. et Stotler, Syst. Bot. 40 (1): 39, 2015 (Shaw et al. 2015). Bas.: Anastrophyllum subacutum Herzog, Ann. Bryol. 12: 75, 1939 (Herzog 1939b). *** Solenostoma subrubrum (Schifn. ex Steph.) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 508, 2009 (Váňa and Long 2009). Bas.: Jungermannia subrubra Schifn. ex Steph., Sp. Hepat. (Stephani) 6: 93, 1917 (Stephani 1917a). * Solenostoma sunii Bakalin et Vilnet, Bot. Paciica 3 (2): 15, 2014 (Bakalin et al. 2014). * Solenostoma totopapillosum (E.A.Hodgs.) R.M.Schust., Bryologist 100 (3): 366, 1997 (Schuster 1997a). Bas.: Jungermannia totopapillosa E.A.Hodgs., J. Roy. Soc. New Zealand 2 (1): 111, 1972 (Hodgson 1972). 97 *** Solenostoma udarii (S.C.Srivast. et P.Singh) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 510, 2009 (Váňa and Long 2009). Bas.: Jungermannia udarii S.C.Srivast. et P.Singh, Recent Stud. Indian Bryoph.: 152, 1995 (Srivastava and Singh 1995). * Solenostoma ventroversum (Grolle) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 510, 2009 (Váňa and Long 2009). Bas.: Jungermannia ventroversa Grolle, Khumbu Himal 1 (4): 284, 1966 (Grolle 1966k). 98 *** Solenostoma zantenii (Amakawa) R.M.Schust. ex Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 510, 2009 (Váňa and Long 2009). Bas.: Jungermannia zantenii Amakawa, J. Hattori Bot. Lab. 31: 110, 1968 (Amakawa 1968b). * Solenostoma zengii (C.Gao et X.L.Bai) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 510, 2009 (Váňa and Long 2009). Bas.: Jungermannia zengii C.Gao et X.L.Bai, Philipp. Scientist 38: 151, 2001 (Gao and Bai 2001). 97 Solenostoma totopapillosum is possibly conspeciic with Solenostoma inundatum (Shaw et al. 2015). 98 Solenostoma ventroversum is possibly conspeciic with Solenostoma appressifolium. World checklist of hornworts and liverworts *** 135 Southbyaceae Váňa, Crand.-Stotl., Stotler et D.G.Long by J. Váňa Southbyaceae has traditionally been placed in Arnelliaceae (e.g. Crandall-Stotler et al. 2009) or kept as a separate family (e.g. Vanden Berghen 1957) that has never been validated. However, it was re-established, rearranged and validated in the study by Váňa et al. (2012e). *** Gongylanthus Nees, Naturgesch. Eur. Leberm. 2: 405, 1836 (Nees 1836). *** Gongylanthus dusenii Steph., Bull. Herb. Boissier (sér. 2) 6 (5): 385 (41), 1906 (Stephani 1906a). *** Gongylanthus ericetorum (Raddi) Nees, Naturgesch. Eur. Leberm. 2: 407, 1836 (Nees 1836). Bas.: Calypogeia ericetorum Raddi, Jungermanniogr. Etrusca: 31, 1818 (Raddi 1818a). *** Gongylanthus granatensis (Gottsche) Steph., Bull. Herb. Boissier (sér. 2) 6 (5): 385 (41), 1906 (Stephani 1906a). Bas.: Lindigina granatensis Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 138, 1864 (Gottsche 1864). *** Gongylanthus himalayensis Grolle, Khumbu Himal 1 (4): 285, 1966 (Grolle 1966k). *** Gongylanthus javanicus Grolle, J. Jap. Bot. 40 (7): 206, 1965 (Grolle 1965d). *** Gongylanthus liebmanianus (Lindenb. et Gottsche) Steph., Bull. Herb. Boissier (sér. 2) 6 (5): 388 (44), 1906 (Stephani 1906a). Bas.: Gymnanthe liebmaniana Lindenb. et Gottsche, Syn. Hepat. 5: 712, 1847 (Gottsche et al. 1847). *** Gongylanthus limbatus (Herzog) Grolle et Váňa, Folia Geobot. Phytotax. 9 (2): 198, 1974 (Váňa 1974c). Bas.: Aplozia limbata Herzog, Beih. Bot. Centralbl. 61B (3): 561, 1942 (Herzog 1942d). *** Gongylanthus muelleri (Gottsche) Steph., Bull. Herb. Boissier (sér. 2) 6 (5): 388 (44), 1906 (Stephani 1906a). Bas.: Lindigia muelleri Gottsche, Mexik. Leverm.: 121, 1863 (Gottsche 1863). ** Gongylanthus oniscoides (Spruce) Steph., Bull. Herb. Boissier (sér. 2) 6 (5): 386 (42), 1906 (Stephani 1906a). Bas.: Calypogeia oniscoides Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 448, 1885 (Spruce 1885). *** Gongylanthus richardsii E.W.Jones, Trans. Brit. Bryol. Soc. 4 (4): 650, 1964 (Jones 1964). *** Southbya Spruce, Ann. Mag. Nat. Hist. (ser. 2) 3 (18): 501, 1849 (Spruce 1849). *** Southbya gollanii Steph., Bull. Herb. Boissier (sér. 2) 6 (5): 381 (37), 1906 (Stephani 1906a). *** Southbya nigrella (De Not.) Henriq., Bol. Soc. Brot. 4: 244, 1886 [1887] (Henriques 1886). Bas.: Jungermannia nigrella De Not., Mem. Reale Accad. Sci. Torino (ser. 2) 1: 315, 1838 (De Notaris 1838). 136 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Southbya organensis Herzog, Memoranda Soc. Fauna Fl. Fennica 25: 53, 1950 (Herzog 1950c). *** Southbya tophacea (Spruce) Spruce, Ann. Mag. Nat. Hist. (ser. 2) 3 (18): 501, 1849 (Spruce 1849). Bas.: Jungermannia tophacea Spruce, Hep. Pyr. Exsic.: no. 23, 1847 (Spruce 1847). ** Stephaniellaceae R.M.Schust. by J. Váňa In the absence of molecular data, the phylogenetic ainities of Stephaniella and Stephaniellidium remain equivocal, but there is no evidence to support their placement in either the Arnelliaceae or Southbyaceae where they have been placed recently. hus the family as construed by Schuster (2002b) is retained. *** Stephaniella J.B.Jack, Hedwigia 33 (1): 11, 1894 (Jack 1894). * Stephaniella boliviensis Steph., Biblioth. Bot. 87 (2): 182, 1916 (Stephani 1916a). *** Stephaniella hamata Steph., Bull. Herb. Boissier (sér. 2) 1 (10): 1024 (87), 1901 (Stephani 1901c). *** Stephaniella paraphyllina J.B.Jack, Hedwigia 33 (1): 11, 1894 (Jack 1894). *** Stephaniella rostrata U.Schmitt, Österr. Bot. Z. 115 (2): 124, 1968 (Schmitt and Winkler 1968). *** Stephaniella uncifolia S.Winkl., Österr. Bot. Z. 115 (2): 124, 1968 (Schmitt and Winkler 1968). ** Stephaniellidium S.Winkl. ex Grolle, Acta Bot. Fenn. 121: 38, 1983 (Grolle 1983b). Based on: Stephaniellidium S.Winkl., Mitt. Inst. Colombo-Alemán Invest. Ci. 3: 60, 1969 (Winkler 1969). *** Stephaniellidium sleumeri (Müll.Frib.) S.Winkl. ex Grolle, Acta Bot. Fenn. 121: 38, 1983 (Grolle 1983b). Bas.: Stephaniella sleumeri Müll.Frib., Rev. Bryol. Lichénol. 20 (1/2): 177, 1951 (Müller 1951b). *** Trichotemnomataceae R.M.Schust. *** Trichotemnoma R.M.Schust., Nova Hedwigia 15: 440, 1968 (Schuster 1968b). *** Trichotemnoma corrugatum (Steph.) R.M.Schust., Nova Hedwigia 15: 440, 1968 (Schuster 1968b). Bas.: Blepharostoma corrugatum Steph., Hedwigia 32 (5): 315, 1893 (Stephani 1893d). World checklist of hornworts and liverworts 137 Lophocoleineae Schljakov *** Blepharostomataceae W.Frey et M.Stech he recognition of Blepharostomataceae follows Frey and Stech (2008) following reconsideration of molecular and morphological data available at that time. *** Blepharostoma (Dumort.) Dumort., Recueil Observ. Jungerm.: 18, 1835 (Dumortier 1835). Bas.: Jungermannia sect. Blepharostoma Dumort., Syll. Jungerm. Europ.: 65, 1831 (Dumortier 1831). ** Blepharostoma arachnoideum M.Howe, Mem. Torrey Bot. Club 7: 140, 1899 (Howe 1899). ** Blepharostoma indicum G.Asthana, M.Saxena et Maurya, J. Bryol. 35 (4): 267, 2013 (Asthana et al. 2013). ** Blepharostoma minor Horik., Hikobia 1 (2): 104, 1951 [1952] (Horikawa 1951b). *** Blepharostoma trichophyllum (L.) Dumort., Recueil Observ. Jungerm.: 18, 1835 (Dumortier 1835). Bas.: Jungermannia trichophylla L., Sp. Pl. 1: 1135, 1753 (Linnaeus 1753). ** Blepharostoma trichophyllum subsp. brevirete (Bryhn et Kaal.) R.M.Schust., Bull. Natl. Mus. Canada 164: 16, 1959 (Schuster et al. 1959). Bas.: Blepharostoma trichophyllum var. brevirete Bryhn et Kaal., Rep. Second Norweg. Arctic Exped. 11: 46, 1906 (Bryhn 1906). *** Brevianthaceae J.J.Engel et R.M.Schust. by L. Söderström, J. Váňa, R. Stotler, B.J. Crandall-Stotler and J.J. Engel he treatment of Brevianthaceae follows Söderström et al. (2013b). *** Brevianthus J.J.Engel et R.M.Schust., Phytologia 47 (4): 317, 1981 (Engel and Schuster 1981). *** Brevianthus lavus (Grolle) J.J.Engel et R.M.Schust., Phytologia 47 (4): 318, 1981 (Engel and Schuster 1981). Bas.: Jackiella lava Grolle, J. Hattori Bot. Lab. 33: 222, 1970 (Grolle 1970b). ** Brevianthus lavus subsp. crenulatus J.J.Engel, Nova Hedwigia 93 (3/4): 406, 2011 (Engel 2011). * Brevianthus hypocanthidium M.A.M.Renner et J.J.Engel, PhytoKeys 50: 46, 2015 (Renner et al. 2015). *** Tetracymbaliella Grolle, Nova Hedwigia 3 (1): 48, 1961 (Grolle 1961b). ** Tetracymbaliella comptonii (Pearson) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 164, 1963 [1964] (Grolle 1963d). Bas.: Chiloscyphus comptonii Pearson, J. Linn. Soc., Bot. 46 (305): 23, 1922 (Pearson 1922b). *** Tetracymbaliella cymbalifera (Hook.f. et Taylor) Grolle, Nova Hedwigia 3 (1): 50, 1961 (Grolle 1961b). Bas.: Jungermannia cymbalifera Hook.f. et Taylor, Bot. Antarct. Voy. I (Fl. Antarct. 1): 151, 1845 (Taylor and Hooker 1845). 138 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Tetracymbaliella decipiens (Gottsche) Grolle, Nova Hedwigia 3 (1): 49, 1961 (Grolle 1961b). Bas.: Chiloscyphus decipiens Gottsche, Syn. Hepat. 2: 176, 1845 (Gottsche et al. 1845a). *** Tetracymbaliella subsimplex (Austin) J.J.Engel, Phytotaxa 207 (2): 185, 2015 (Engel 2015b). Bas.: Polyotus subsimplex Austin, Bull. Torrey Bot. Club 6 (7): 46, 1875 (Austin 1875c). ** Chonecoleaceae R.M.Schust. ex Grolle ** Chonecolea Grolle, Rev. Bryol. Lichénol. 25 (3/4): 294, 1956 [1957] (Grolle 1956). ** Chonecolea acutiloba (Schifn.) R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 321, 1980 (Schuster 1980c). Bas.: Clasmatocolea acutiloba Schifn., Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 63, 1964 (Schifner and Arnell 1964). ** Chonecolea andina Grolle et Váňa, J. Hattori Bot. Lab. 48: 229, 1980 (Váňa 1980). ** Chonecolea doellingeri (Nees) Grolle, Rev. Bryol. Lichénol. 25 (3/4): 295, 1956 [1957] (Grolle 1956). Bas.: Jungermannia doellingeri Nees, Syn. Hepat. 1: 104, 1844 (Gottsche et al. 1844). ** Chonecolea ruwenzorensis E.W.Jones, J. Bryol. 13 (4): 498, 1986 (Jones 1986). ** Chonecolea schusteri Udar et Ad.Kumar, Bryologist 85 (3): 315, 1982 (Udar and Kumar 1982a). ** Chonecolea verae Potemkin, Proc. int. meeting 90th anniv. Abramova: 165, 2005 (Potemkin 2005). ** Grolleaceae Solari ex R.M.Schust. ** Grollea R.M.Schust., Nova Hedwigia 8 (3/4): 288, 1964 (Schuster 1964b). ** Grollea antheliopsis R.M.Schust., Nova Hedwigia 8 (3/4): 288, 1964 (Schuster 1964b). *** Herbertaceae Müll.Frib. ex Fulford et Hatcher by D. Bell *** Herbertus Gray, Nat. Arr. Brit. Pl. 1: 705, 1821 (Gray 1821). 99 *** Herbertus aduncus (Dicks.) Gray, Nat. Arr. Brit. Pl. 1: 705, 1821 (Gray 1821). Bas.: Jungermannia adunca Dicks., Fasc. Pl. Crypt. Brit. 3: 12, 1793 (Dickson 1793). 99 Herbertus includes Schisma, but a few taxa have neither been transferred nor synonymized. hey are listed in the “Names in genera not currently accepted” section below. World checklist of hornworts and liverworts 139 ** Herbertus arcticus (Inoue et Steere) Schljakov, Novosti Sist. Nizš. Rast. 19: 209, 1982 (Shliakov 1982). Bas.: Herbertus sakuraii subsp. arcticus Inoue et Steere, J. Hattori Bot. Lab. 44: 266, 1978 (Steere and Inoue 1978). ** Herbertus armitanus (Steph.) H.A.Mill., J. Hattori Bot. Lab. 28: 324, 1965 (Miller 1965). Bas.: Schisma armitanum Steph., Sp. Hepat. (Stephani) 4: 28, 1909 (Stephani 1909d). ** Herbertus asparus Tixier, Bryophyt. Biblioth. 18: 64, 1979 (Tixier 1979a). *** Herbertus bivittatus Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 343, 1885 (Spruce 1885). ** Herbertus borealis Crundw., Trans. Brit. Bryol. Soc. 6 (1): 41, 1970 (Crundwell 1970). ** Herbertus buchii Juslén, Ann. Bot. Fenn. 43 (6): 416, 2006 (Juslén 2006a). ** Herbertus ceylanicus (Steph.) Abeyw., Ceylon J. Sci., Biol. Sci. 2 (1): 43, 1959 (Abeywickrama 1959). Bas.: Schisma ceylanicum Steph., Sp. Hepat. (Stephani) 4: 22, 1909 (Stephani 1909d). ** Herbertus circinatus (Steph.) H.A.Mill., J. Hattori Bot. Lab. 28: 318, 1965 (Miller 1965). Bas.: Schisma circinatum Steph., Sp. Hepat. (Stephani) 4: 25, 1909 (Stephani 1909d). *** Herbertus delavayi (Steph.) Steph., Hedwigia 34 (2): 43, 1895 (Stephani 1895c). Bas.: Schisma delavayi Steph., Mém. Soc. Nat. Sci. Nat. Math. Cherbourg 29: 228, 1894 (Stephani 1894b). 100 *** Herbertus dicranus (Gottsche, Lindenb. et Nees) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 397, 1877 (Trevisan 1877). Bas.: Sendtnera dicrana Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 239, 1845 (Gottsche et al. 1845a). ** Herbertus durandii (Steph.) Herzog, Rev. Bryol. Lichénol. 11 (1): 25, 1938 [1939] (Herzog 1938a). Bas.: Schisma durandii Steph., Trans. Linn. Soc. London, Bot. 6 (1): 99, 1901 (Stephani 1901e). * Herbertus evittatus (Steph.) H.A.Mill., J. Hattori Bot. Lab. 28: 327, 1965 (Miller 1965). Bas.: Schisma evittatum Steph., Sp. Hepat. (Stephani) 6: 357, 1922 (Stephani 1922). ** Herbertus gaochienii X.Fu, Fl. Bryoph. Sin. 9: 38, 2003 (Gao 2003). ** Herbertus gracilis (Mont.) Steph., Bull. Herb. Boissier 5 (10): 842, 1897 (Stephani 1897c). Bas.: Mastigophora gracilis Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 254, 1843 (Montagne 1843). ** Herbertus guangdongii P.J.Lin ex Piippo, Bryobrothera 1: 206, 1992 (Lin et al. 1992). ** Herbertus hawaiiensis H.A.Mill., J. Hattori Bot. Lab. 28: 317, 1965 (Miller 1965). ** Herbertus helleri (Steph.) W.E.Nicholson, Rev. Bryol. Lichénol. 13: 143, 1942 (Nicholson 1942). Bas.: Schisma helleri Steph., Sp. Hepat. (Stephani) 4: 29, 1909 (Stephani 1909d). ** Herbertus herpocladioides E.B.Scott et H.A.Mill., Bryologist 62 (2): 116, 1959 (Scott and Miller 1959). 100 Herbertus delavayi is a species complex that requires further studies (Bell et al. 2012). It also includes Herbertus borealis and may include several, partly cryptic species. 140 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Herbertus hutchinsiae (Gottsche et Rabenh.) A.Evans, Bull. Torrey Bot. Club 44 (4): 214, 1917 (Evans 1917c). Bas.: Sendtnera adunca var. hutchinsiae Gottsche et Rabenh., Hepat. Eur., Leberm. 21-22: no. 210, 1862 (Rabenhorst 1862). *** Herbertus juniperoideus (Sw.) Grolle, Rev. Bryol. Lichénol. 30 (1/2): 80, 1961 (Grolle 1961a). Bas.: Jungermannia juniperoidea Sw., Prodr. (Swartz): 144, 1788 (Swartz 1788). 101 *** Herbertus juniperoideus subsp. acanthelius (Spruce) K.Feldberg et Heinrichs, Bot. J. Linn. Soc. 151: 326, 2006 (Feldberg and Heinrichs 2006). Bas.: Herbertus acanthelius Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 341, 1885 (Spruce 1885). *** Herbertus juniperoideus subsp. pensilis (Taylor) K.Feldberg et Heinrichs, Bot. J. Linn. Soc. 151: 329, 2006 (Feldberg and Heinrichs 2006). Bas.: Sendtnera pensilis Taylor, London J. Bot. 5: 372, 1846 (Taylor 1846b). ** Herbertus kurzii (Steph.) R.S.Chopra, J. Indian Bot. Soc. 22: 247, 1943 (Chopra 1943). Bas.: Schisma kurzii Steph., Sp. Hepat. (Stephani) 4: 24, 1909 (Stephani 1909d). ** Herbertus leratii (Steph.) H.A.Mill., J. Hattori Bot. Lab. 28: 327, 1965 (Miller 1965). Bas.: Schisma leratii Steph., Sp. Hepat. (Stephani) 6: 360, 1922 (Stephani 1922). ** Herbertus lonchobasis H.A.Mill., J. Hattori Bot. Lab. 28: 306, 1965 (Miller 1965). ** Herbertus longiissus Steph., Hedwigia 34 (2): 44, 1895 (Stephani 1895c). ** Herbertus longispinus J.B.Jack et Steph., Hedwigia 31 (1): 15, 1892 (Jack and Stephani 1892). ** Herbertus mauritianus N.G.Hodgetts, J. Bryol. 30 (4): 247, 2008 (Hodgetts 2008). *** Herbertus norenus D.G.Long, D.Bell et H.H.Blom, Molec. Ecol. Res. 12: 44, 2012 (Bell et al. 2012). ** Herbertus oldieldianus (Steph.) Rodway, Tasm. Bryoph.: 72, 1917 (Rodway 1917b). Bas.: Schisma oldieldianum Steph., Sp. Hepat. (Stephani) 4: 20, 1909 (Stephani 1909d). ** Herbertus pilifer Schifn., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 60 (2): 252, 1893 (Schifner 1893a). ** Herbertus pocsii N.G.Hodgetts, J. Bryol. 30 (4): 249, 2008 (Hodgetts 2008). ** Herbertus pumilus Steph., Hedwigia 34 (2): 44, 1895 (Stephani 1895c). ** Herbertus ramosus (Steph.) H.A.Mill., J. Hattori Bot. Lab. 28: 314, 1965 (Miller 1965). Bas.: Schisma ramosum Steph., Sp. Hepat. (Stephani) 4: 23, 1909 (Stephani 1909d). ** Herbertus runcinatus (Taylor) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 397, 1877 (Trevisan 1877). Bas.: Sendtnera runcinata Taylor, London J. Bot. 5: 372, 1846 (Taylor 1846b). *** Herbertus sendtneri (Nees) Lindb., Hepat. Scand. Exsicc.: no. 4, 1874 (Lindberg and Lackström 1874). Bas.: Schisma sendtneri Nees, Naturgesch. Eur. Leberm. 3: 575, 1838 (Nees 1838b). ** Herbertus spicatus N.G.Hodgetts, J. Bryol. 30 (4): 244, 2008 (Hodgetts 2008). 101 Herbertus juniperoideus is here treated with two subspecies, but they may deserve species rank instead. World checklist of hornworts and liverworts 141 *** Herbertus stramineus (Dumort.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 397, 1877 (Trevisan 1877). Bas.: Schisma stramineum Dumort., Syll. Jungerm. Europ.: 77, 1831 (Dumortier 1831). ** Herbertus streimannii M.L.So, Syst. Bot. 28 (1): 13, 2003 (So 2003). *** Herbertus tenuis A.Evans, Bull. Torrey Bot. Club 44 (4): 219, 1917 (Evans 1917c). ** Herbertus udarii D.Kumar et N.Manocha, Geophytology 29 (1/2): 105, 1999 [2000] (Kumar and Manocha 1999). * Excluded from the genus Herbertus subrotundatus X.Fu et Y.J.Yi, Acta Phytotax. Sin. 39 (1): 89, 2001 (Yi et al. 2001). 102 *** Triandrophyllum Fulford et Hatcher, Bryologist 64 (4): 349, 1961 [1962] (Fulford and Hatcher 1961). Based on: Triandrophyllum Fulford et Hatcher, Bryologist 61 (4): 277, 1958 [1959] (Fulford and Hatcher 1958). *** Triandrophyllum eophyllum (R.M.Schust.) Gradst., Mem. New York Bot. Gard. 86: 104, 2001 (Gradstein et al. 2001a). Bas.: Olgantha eophylla R.M.Schust., Nova Hedwigia 63 (3/4): 535, 1996 (Schuster 1996b). ** Triandrophyllum fernandeziense (S.W.Arnell) Grolle ex Fulford et Hatcher, Bryologist 64 (4): 351, 1961 [1962] (Fulford and Hatcher 1961). Bas.: Acromastigum fernandeziense S.W.Arnell, Ark. Bot. (n.ser.) 4 (1): 10, 1957 (Arnell 1957b). ** Triandrophyllum heterophyllum (Steph.) Grolle, J. Jap. Bot. 39 (8): 238, 1964 (Grolle 1964g). Bas.: Mastigophora heterophylla Steph., Sp. Hepat. (Stephani) 6: 367, 1922 (Stephani 1922). *** Triandrophyllum subtriidum (Hook.f. et Taylor) Fulford et Hatcher, Bryologist 64 (4): 350, 1961 [1962] (Fulford and Hatcher 1961). Bas.: Jungermannia subtriida Hook.f. et Taylor, London J. Bot. 3: 579, 1844 (Hooker and Taylor 1844c). ** Triandrophyllum subtriidum var. triidum (Gottsche) Solari, Bol. Soc. Argent. Bot. 15 (2/3): 201, 1973 (Solari 1973). Bas.: Sendtnera triida Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 142, 1864 (Gottsche 1864). ** Triandrophyllum symmetricum J.J.Engel, Haussknechtia, Beih. 9: 115, 1999 (Engel 1999b). *** Lepicoleaceae R.M.Schust. by M. von Konrat Lepicoleaceae has conventionally been considered a monogeneric family only containing Lepicolea (e.g. Crandall-Stotler et al. 2009). However, we expand Lepicoleaceae here to include the monogeneric family Vetaformaceae. A number of molecular analy102 Herbertus subrotundus does not belong in Herbertus (Juslén 2006a), but it is unclear where it belongs. 142 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ses have shown that Vetaforma dusenii is sister to Lepicolea (e.g. He-Nygrén et al. 2006, Juslén 2006b) together forming a robust clade. *** Lepicolea Dumort., Recueil Observ. Jungerm.: 20, 1835 (Dumortier 1835). *** Lepicolea attenuata (Mitt.) Steph., J. Linn. Soc., Bot. 29 (201): 276, 1892 (Stephani 1892b). Bas.: Sendtnera attenuata Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 153, 1854 (Mitten 1854). *** Lepicolea magellanica (Gola) Solari, Lindbergia 9 (2): 86, 1983 (Solari 1983b). Bas.: Lepicolea scolopendra var. magellanica Gola, Nuovo Giorn. Bot. Ital. (n.ser.) 29 (1/4): 170, 1922 [1923] (Gola 1922). *** Lepicolea norrisii Piippo, Ann. Bot. Fenn. 25 (1): 55, 1988 (Piippo 1988). *** Lepicolea ochroleuca (Spreng.) Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 345, 1885 (Spruce 1885). Bas.: Jungermannia ochroleuca Spreng. Syst. Veg. (ed. 16) [Sprengel] 4 (2): 325, 1827 (Sprengel 1827b). *** Lepicolea pruinosa (Taylor) Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 345, 1885 (Spruce 1885). Bas.: Sendtnera pruinosa Taylor, London J. Bot. 5: 373, 1846 (Taylor 1846b). *** Lepicolea ramentiissa Herzog, Biblioth. Bot. 88: 30, 1920 [1921] (Herzog 1920). *** Lepicolea rara (Steph.) Grolle, Nova Hedwigia 16: 152, 1968 (Grolle 1968d). Bas.: Lepidozia rara Steph., Sp. Hepat. (Stephani) 3: 618, 1909 (Stephani 1909a). 103 *** Lepicolea rigida (De Not.) E.B.Scott, Nova Hedwigia 2: 148, 1960 (Scott 1960). Bas.: Sendtnera rigida De Not., Mem. Reale Accad. Sci. Torino (ser. 2) 16: 229, 1857 (De Notaris 1857). *** Lepicolea scolopendra (Hook.) Dumort. ex Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 398, 1877 (Trevisan 1877). Bas.: Jungermannia scolopendra Hook., Musci Exot. 1: tab. 40, 1818 (Hooker 1818). *** Lepicolea yakusimensis (S.Hatt.) S.Hatt., J. Hattori Bot. Lab. 10: 42, 1953 (Hattori 1953c). Bas.: Lepicolea scolopendra var. yakusimensis S.Hatt., J. Hattori Bot. Lab. 2: 9, 1947 [1948] (Hattori 1947b). *** Vetaforma Fulford et J.Taylor, Nova Hedwigia 4 (1/2): 81, 1962 (Fulford 1962b). *** Vetaforma dusenii (Steph.) Fulford et J.Taylor, Nova Hedwigia 4 (1/2): 82, 1962 (Fulford 1962b). Bas.: Lepidozia dusenii Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 6): 52, 1900 (Stephani 1900b). *** Lepidoziaceae Limpr. by E.D. Cooper he classiication within Lepidoziaceae is in a state of lux. he treatment provided here follows the interim classiication proposed by Cooper (2013) based on recent molecular phylogenetic studies (Heslewood and Brown 2007, Cooper et al. 2011, 2012a, 2012b) 103 Lepicolea rara is possibly a species complex and warrants further investigation (Schuster 2000a). World checklist of hornworts and liverworts 143 incorporating nomenclatural changes from Cooper et al. (2013). Several higher taxa are unlikely to represent monophyletic units, but the phylogenetic data currently available are insuicient to re-circumscribe the doubtful subfamilies and genera (Cooper 2013). Infrageneric taxa have been retained where phylogenetic data are absent or inconclusive. ** Meinungeria Frank Müll., Bryologist 110 (3): 494, 2007 (Müller 2007). ** Meinungeria mouensis Frank Müll., Bryologist 110 (3): 494, 2007 (Müller 2007). *** Bazzanioideae Rodway *** Acromastigum A.Evans, Bull. Torrey Bot. Club 27 (3): 103, 1900 (Evans 1900b). 104 ** subg. Acromastigum *** Acromastigum caledonicum (Steph.) Grolle, Österr. Bot. Z. 111 (2/3): 243, 1964 (Grolle 1964h). Bas.: Acolea caledonica Steph., Nova Caledonia, Bot. 1: 19, 1914 (Stephani 1914c). *** Acromastigum cavifolium R.M.Schust., J. Hattori Bot. Lab. 26: 257, 1963 (Schuster 1963b). ** Acromastigum herzogii Grolle, Österr. Bot. Z. 111 (2/3): 250, 1964 (Grolle 1964h). *** Acromastigum homodictyon (Herzog) Grolle, Österr. Bot. Z. 111 (2/3): 245, 1964 (Grolle 1964h). Bas.: Acromastigum integrifolium var. homodictyon Herzog, Ark. Bot. (n.ser.) 3 (3): 44, 1953 (Herzog 1953a). ** Acromastigum integrifolium (Austin) A.Evans, Bull. Torrey Bot. Club 27 (3): 103, 1900 (Evans 1900b). Bas.: Mastigobryum integrifolium Austin, Bot. Bull. (Hanover) 1 (7): 32, 1876 (Austin 1876b). *** Acromastigum stellare N.Kitag., Acta Phytotax. Geobot. 36 (4/6): 109, 1985 (Kitagawa 1985). ** Acromastigum stenophyllum R.M.Schust., Nova Hedwigia 15: 465, 1968 (Schuster 1968b). *** Acromastigum verticale (Steph.) E.A.Hodgs., Trans. Roy. Soc. New Zealand 82 (1): 18, 1954 (Hodgson 1954). Bas.: Bazzania verticalis Steph., Hedwigia 32 (4): 214, 1893 (Stephani 1893c). ** subg. Inaequilatera (Schifn.) Grolle, J. Hattori Bot. Lab. 44: 2, 1978 (Grolle 1978b). Bas.: Bazzania sect. Inaequilaterae Schifn., Hepat. (Engl.-Prantl): 101, 1893 (Schifner 1893b). *** Acromastigum adaptatum Hürl., Bauhinia 7 (4): 263, 1983 (Hürlimann 1983). *** Acromastigum anisostomum (Lehm. et Lindenb.) A.Evans, Ann. Bryol., Suppl. 3: 48, 1934 (Evans 1934). Bas.: Jungermannia anisostoma Lehm. et Lindenb., Nov. Stirp. Pug. 6: 57, 1834 (Lehmann 1834). 104 Acromastigum is here divided into two subgenera which are supported by recent molecular studies (Heslewood and Brown 2007, Cooper et al. 2011). 144 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Acromastigum anisostomum var. minutum E.A.Hodgs., Trans. Roy. Soc. New Zealand, Bot. 3 (4): 71, 1965 (Hodgson 1965). ** Acromastigum aurescens A.Evans, Ann. Bryol., Suppl. 3: 45, 1934 (Evans 1934). ** Acromastigum bancanum (Sande Lac.) A.Evans, Ann. Bryol., Suppl. 3: 20, 1934 (Evans 1934). Bas.: Mastigobryum bancanum Sande Lac., Ann. Mus. Bot. Lugduno-Batavi 1: 301, 1864 (Sande Lacoste 1864). ** Acromastigum brotheri (Steph.) A.Evans, Ann. Bryol., Suppl. 3: 70, 1934 (Evans 1934). Bas.: Mastigobryum brotheri Steph., Sp. Hepat. (Stephani) 3: 536, 1909 (Stephani 1909a). ** Acromastigum capillare (Steph.) A.Evans, Ann. Bryol., Suppl. 3: 37, 1934 (Evans 1934). Bas.: Mastigobryum capillare Steph., Sp. Hepat. (Stephani) 6: 457, 1924 (Stephani 1924). *** Acromastigum colensoanum (Mitt.) A.Evans ex Reimers, Hedwigia 73 (3/4): 142, 1933 (Reimers 1933). Bas.: Mastigobryum colensoanum Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 147, 1854 (Mitten 1854). *** Acromastigum cunninghamii (Steph.) A.Evans, Ann. Bryol., Suppl. 3: 106, 1934 (Evans 1934). Bas.: Bazzania cunninghamii Steph., Hedwigia 32 (4): 205, 1893 (Stephani 1893c). ** Acromastigum curtilobum A.Evans, Ann. Bryol., Suppl. 3: 97, 1934 (Evans 1934). *** Acromastigum divaricatum (Nees) A.Evans ex Reimers, Hedwigia 73 (3/4): 142, 1933 (Reimers 1933). Bas.: Mastigobryum divaricatum Nees, Syn. Hepat. 2: 219, 1845 (Gottsche et al. 1845a). ** Acromastigum echinatiforme (De Not.) A.Evans, Ann. Bryol., Suppl. 3: 64, 1934 (Evans 1934). Bas.: Mastigobryum echinatiforme De Not., Epat. Borneo: 38, 1874 (De Notaris 1874). * Acromastigum echinatum (Gottsche) A.Evans, Ann. Bryol., Suppl. 3: 147, 1934 (Evans 1934). Bas.: Mastigobryum echinatum Gottsche, Syn. Hepat. 2: 218, 1845 (Gottsche et al. 1845a). 105 *** Acromastigum exiguum (Steph.) A.Evans, Ann. Bryol., Suppl. 3: 75, 1934 (Evans 1934). Bas.: Mastigobryum exiguum Steph., Hedwigia 25 (1): 6, 1886 (Stephani 1886e). ** Acromastigum exile (Lindenb.) A.Evans, Ann. Bryol., Suppl. 3: 24, 1934 (Evans 1934). Bas.: Mastigobryum exile Lindenb., Syn. Hepat. 2: 217, 1845 (Gottsche et al. 1845a). *** Acromastigum ilum (Steph.) A.Evans, Ann. Bryol., Suppl. 3: 31, 1934 (Evans 1934). Bas.: Bazzania ilum Steph., Hedwigia 32 (4): 206, 1893 (Stephani 1893c). *** Acromastigum ilum var. papillosum N.Kitag., Acta Phytotax. Geobot. 36 (4/6): 112, 1985 (Kitagawa 1985). ** Acromastigum imbriatum (Steph.) A.Evans, Ann. Bryol., Suppl. 3: 153, 1934 (Evans 1934). Bas.: Mastigobryum imbriatum Steph., Sp. Hepat. (Stephani) 3: 538, 1909 (Stephani 1909a). 105 Acromastigum echinatum may be conspeciic with Acromastigum inaequilaterum since all specimens identiied by Piippo et al. (2002) belong there, but she did not see the type specimen. World checklist of hornworts and liverworts 145 ** Acromastigum furcatifolium (Steph.) E.A.Br., Phytotaxa 65: 58, 2012 (Brown et al. 2012). Bas.: Lepidozia furcatifolia Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 112, 1914 (Stephani and Watts 1914). *** Acromastigum inaequilaterum (Lehm. et Lindenb.) A.Evans, Ann. Bryol., Suppl. 3: 129, 1934 (Evans 1934). Bas.: Jungermannia inaequilatera Lehm. et Lindenb., Nov. Stirp. Pug. 6: 56, 1834 (Lehmann 1834). ** Acromastigum interstisiale E.A.Br. et M.A.M.Renner, Telopea 17: 268, 2014 (Brown and Renner 2014). ** Acromastigum laetevirens (Sande Lac. ex Steph.) A.Evans, Ann. Bryol., Suppl. 3: 94, 1934 (Evans 1934). Bas.: Mastigobryum laetevirens Sande Lac. ex Steph., Hedwigia 25 (4): 133, 1886 (Stephani 1886b). ** Acromastigum laevigatum A.Evans, Ann. Bryol., Suppl. 3: 101, 1934 (Evans 1934). ** Acromastigum linganum (De Not.) A.Evans, Ann. Bryol., Suppl. 3: 118, 1934 (Evans 1934). Bas.: Mastigobryum linganum De Not., Epat. Borneo: 37, 1874 (De Notaris 1874). ** Acromastigum lobuliferum A.Evans, Ann. Bryol., Suppl. 3: 157, 1934 (Evans 1934). ** Acromastigum longirete Grolle, J. Hattori Bot. Lab. 44: 11, 1978 (Grolle 1978b). *** Acromastigum mooreanum (Steph.) E.A.Hodgs., Trans. Roy. Soc. New Zealand 82 (1): 19, 1954 (Hodgson 1954). Bas.: Bazzania mooreana Steph., Hedwigia 33 (1): 1, 1894 (Stephani 1894a). *** Acromastigum moratii N.Kitag., Acta Phytotax. Geobot. 36 (4/6): 119, 1985 (Kitagawa 1985). ** Acromastigum obliquatum (Mitt.) A.Evans, Ann. Bryol., Suppl. 3: 110, 1934 (Evans 1934). Bas.: Bazzania obliquata Mitt., Hedwigia 32 (4): 211, 1893 (Stephani 1893c). ** Acromastigum tenax (Steph.) A.Evans, Ann. Bryol., Suppl. 3: 41, 1934 (Evans 1934). Bas.: Mastigobryum tenax Steph., Sp. Hepat. (Stephani) 6: 483, 1924 (Stephani 1924). ** ** *** ** ** ** ** ** Incertae sedis Acromastigum fumosum E.A.Br. et M.A.M.Renner, Telopea 17: 274, 2014 (Brown and Renner 2014). Acromastigum leptophyllum Herzog, Trans. Brit. Bryol. Soc. 1 (4): 309, 1950 (Herzog 1950a). Acromastigum marginatum E.A.Hodgs., Trans. Roy. Soc. New Zealand 82 (1): 22, 1954 (Hodgson 1954). Acromastigum microstictum A.Evans, Ann. Bryol., Suppl. 3: 115, 1934 (Evans 1934). Acromastigum prismaticale E.A.Br. et M.A.M.Renner, Telopea 17: 281, 2014 (Brown and Renner 2014). Acromastigum pusillum N.Kitag., Acta Phytotax. Geobot. 36 (4/6): 116, 1985 (Kitagawa 1985). Acromastigum rigidum R.M.Schust., Nova Hedwigia 64 (3/4): 617, 1997 (Schuster 1997b). Acromastigum subechinatiforme Hürl., Bauhinia 7 (4): 266, 1983 (Hürlimann 1983). 146 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Bazzania Gray, Nat. Arr. Brit. Pl. 1: 704, 1821 (Gray 1821) nom. conserv. 106 *** Bazzania acanthostipa Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 381, 1885 (Spruce 1885). *** Bazzania accreta (Lehm. et Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (Trevisan 1877). Bas.: Mastigobryum accretum Lehm. et Lindenb., Syn. Hepat. 2: 222, 1845 (Gottsche et al. 1845a). ** Bazzania acinaciformis Steph., Akad. Wiss. Wien, Math.-Naturwiss. Kl., Denkschr. 81: 290, 1907 (Stephani 1907a). ** Bazzania acuminata (Lindenb. et Gottsche) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (Trevisan 1877). Bas.: Mastigobryum acuminatum Lindenb. et Gottsche, Syn. Hepat. 5: 719, 1847 (Gottsche et al. 1847). ** Bazzania acutifolia (Steph.) Schifn., Consp. Hepat. Arch. Ind.: 147, 1898 (Schifner 1898b). Bas.: Mastigobryum acutifolium Steph., Hedwigia 24 (5): 214, 1885 (Stephani 1885a). *** Bazzania adnexa (Lehm. et Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (Trevisan 1877). Bas.: Jungermannia adnexa Lehm. et Lindenb., Nov. Stirp. Pug. 4: 58, 1832 (Lehmann 1832). ** Bazzania adnexa var. aucklandica (Lindenb. et Gottsche) J.J.Engel et G.L.Merr., Bryologist 97 (3): 319, 1994 (Engel and Smith Merrill 1994). Bas.: Mastigobryum novae-hollandiae f. aucklandicum Lindenb. et Gottsche, Sp. Hepat. (Lindenberg) 8-11: 33, 1851 (Lindenberg and Gottsche 1851b). *** Bazzania ainis (Lindenb. et Gottsche) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 415, 1877 (Trevisan 1877). Bas.: Mastigobryum aine Lindenb. et Gottsche, Syn. Hepat. 5: 720, 1847 (Gottsche et al. 1847). ** Bazzania albifolia Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 195, 1934 (Horikawa 1934). ** Bazzania ambigua (Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (Trevisan 1877). Bas.: Mastigobryum ambiguum Lindenb., Syn. Hepat. 2: 217, 1845 (Gottsche et al. 1845a). *** Bazzania amblyphylla Meagher, Nova Hedwigia 92 (3/4): 487, 2011 (Meagher 2011). ** Bazzania aneityensis (Steph.) Tixier, Bull. Mus. Natl. Hist. Nat. (Sér. 3), Bot. 10 (190): 80, 1973 (Tixier 1973d). Bas.: Mastigobryum aneityense Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 121, 1914 (Stephani and Watts 1914). ** Bazzania angusta (Steph.) Herzog, Trans. Brit. Bryol. Soc. 1 (4): 307, 1950 (Herzog 1950a). Bas.: Mastigobryum angustum Steph., Sp. Hepat. (Stephani) 6: 453, 1924 (Stephani 1924). ** Bazzania angustifalcata Herzog, Feddes Repert. Spec. Nov. Regni Veg. 57 (1/2): 166, 1955 (Herzog 1955). 106 Bazzania can not be subdivided based on recent molecular studies, so infrageneric taxa are not used here. he genus includes Mastigobryum, but several taxa have neither been transferred nor synonymized. hey are listed in the “Names in genera not currently accepted” section below. World checklist of hornworts and liverworts 147 ** Bazzania angustifolia Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 198, 1934 (Horikawa 1934). ** Bazzania angustisedens (Steph.) N.Kitag., J. Hattori Bot. Lab. 36: 445, 1972 [1973] (Kitagawa 1972). Bas.: Mastigobryum angustisedens Steph., Bull. Herb. Boissier (sér. 2) 8 (10): 745 (429), 1908 (Stephani 1908c). ** Bazzania angustistipula N.Kitag., J. Hattori Bot. Lab. 30: 268, 1967 (Kitagawa 1967a). *** Bazzania appendiculata (Mitt.) S.Hatt., Fl. E. Himalaya: 505, 1966 (Hattori 1966c). Bas.: Mastigobryum appendiculatum Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 105, 1860 [1861] (Mitten 1860c). ** Bazzania approximata Onr., Bull. Jard. Bot. Natl. Belg. 47 (1/2): 139, 1977 (Onraedt 1977). ** Bazzania arcuata (Lindenb. et Gottsche) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (Trevisan 1877). Bas.: Mastigobryum arcuatum Lindenb. et Gottsche, Syn. Hepat. 5: 718, 1847 (Gottsche et al. 1847). ** Bazzania arcuata var. mamillosa Gradst. et A.R.Benitez, Nova Hedwigia 99 (1/2): 113, 2014 (Gradstein and Benitez 2014). ** Bazzania armatistipula (Steph.) Fulford, Ann. Cryptog. Phytopathol. 3: 114, 1946 (Fulford 1946). Bas.: Mastigobryum armatistipulum Steph., Bull. Herb. Boissier (sér. 2) 8 (11): 866 (490), 1908 (Stephani 1908a). ** Bazzania asperrima Steph., Rev. Bryol. 34 (3): 48, 1907 (Paris 1907). * Bazzania asymmetrica (Steph.) N.Kitag., Bull. Nara Univ. Educ., B 28 (2): 77, 1979 (Kitagawa 1979a). Bas.: Mastigobryum asymmetricum Steph., Sp. Hepat. (Stephani) 6: 454, 1924 (Stephani 1924). 107 ** Bazzania aterrima (Steph.) N.Kitag., Bull. Nara Univ. Educ., B 26 (2): 77, 1977 (Kitagawa 1977). Bas.: Mastigobryum aterrimum Steph., Sp. Hepat. (Stephani) 6: 454, 1924 (Stephani 1924). *** Bazzania aurescens Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 374, 1885 (Spruce 1885). ** Bazzania avia Meagher, Nova Hedwigia 97 (3/4): 529, 2013 (Meagher 2013). ** Bazzania azorica H.Buch et Perss., Bryophyt. Azoren Madeira: 3, 1941 (Buch and Persson 1941). ** Bazzania baldwinii Austin, Trans. Connecticut Acad. Arts 8 (15): 255, 1891 (Evans 1891). ** Bazzania bernieri (Steph.) Inoue et H.A.Mill., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 8 (2): 142, 1965 (Inoue and Miller 1965). Bas.: Mastigobryum bernieri Steph., Bull. Herb. Boissier (sér. 2) 8 (11): 852 (476), 1908 (Stephani 1908a). *** Bazzania bescherellei Steph., Hedwigia 32 (4): 204, 1893 (Stephani 1893c). ** Bazzania bhutanica N.Kitag. et Grolle, J. Hattori Bot. Lab. 61: 269, 1986 [1987] (Kitagawa and Grolle 1986). ** Bazzania bicrenata N.Kitag., J. Hattori Bot. Lab. 47: 127, 1980 (Kitagawa 1980). 107 Bazzania asymmetrica is conspeciic with Bazzania macgregorii in Grolle (1968a), but Kitagawa (1979a) kept them separate. 148 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Bazzania bidens (Gottsche et Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 415, 1877 (Trevisan 1877). Bas.: Mastigobryum bidens Gottsche et Lindenb., Syn. Hepat. 2: 228, 1845 (Gottsche et al. 1845a). ** Bazzania bidens var. heterodonta Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 372, 1885 (Spruce 1885). ** Bazzania bidentula (Steph.) Yasuda, Shokubutsugaku Kakuron: 711, 1911 (Yasuda 1911). Bas.: Pleuroschisma bidentulum Steph., Mém. Soc. Nat. Sci. Nat. Math. Cherbourg 29: 222, 1894 (Stephani 1894b). *** Bazzania bilobata N.Kitag., J. Hattori Bot. Lab. 30: 257, 1967 (Kitagawa 1967a). ** Bazzania borneensis N.Kitag., J. Hattori Bot. Lab. 37: 263, 1973 (Kitagawa 1973). Nom. nov. pro Mastigobryum borneense Steph., Bull. Herb. Boissier (sér. 2) 8 (11): 839 (463), 1908 (Stephani 1908a), nom. illeg. ** Bazzania brasiliensis (Gottsche et Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 415, 1877 (Trevisan 1877). Bas.: Mastigobryum brasiliense Gottsche et Lindenb., Syn. Hepat. 2: 227, 1845 (Gottsche et al. 1845a). ** Bazzania brighamii (Austin) A.Evans, Trans. Connecticut Acad. Arts 8 (15): 255, 1891 (Evans 1891). Bas.: Mastigobryum brighamii Austin, Bull. Torrey Bot. Club 5 (3): 16, 1874 (Austin 1874). ** Bazzania cadens N.Kitag., J. Hattori Bot. Lab. 47: 129, 1980 (Kitagawa 1980). *** Bazzania calcarata (Sande Lac.) Schifn., Consp. Hepat. Arch. Ind.: 149, 1898 (Schifner 1898b). Bas.: Mastigobryum calcaratum Sande Lac., Ann. Mus. Bot. Lugduno-Batavi 1: 304, 1864 (Sande Lacoste 1864). ** Bazzania callida (Sande Lac. ex Steph.) Abeyw., Ceylon J. Sci., Biol. Sci. 2 (1): 45, 1959 (Abeywickrama 1959). Bas.: Mastigobryum callidum Sande Lac. ex Steph., Hedwigia 24 (6): 246, 1885 (Stephani 1885b). *** Bazzania canelensis (Steph.) Fulford, Ann. Cryptog. Phytopathol. 3: 152, 1946 (Fulford 1946). Bas.: Mastigobryum canelense Steph., Sp. Hepat. (Stephani) 3: 518, 1909 (Stephani 1909a). ** Bazzania caudata (Steph.) Herzog, Trans. Brit. Bryol. Soc. 1 (4): 301, 1950 (Herzog 1950a). Bas.: Mastigobryum caudatum Steph., Bull. Herb. Boissier (sér. 2) 8 (12): 947 (497), 1908 (Stephani 1908b). *** Bazzania caudistipula (Steph.) Inoue et H.A.Mill., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 8 (2): 141, 1965 (Inoue and Miller 1965). Bas.: Mastigobryum caudistipulum Steph., Bull. Herb. Boissier (sér. 2) 8 (12): 945 (495), 1908 (Stephani 1908b). ** Bazzania ceylanica (Mitt.) Steph., Bot. Jahrb. Syst. 23 (1/2, 3): 306, 1896 (Stephani 1896a). Bas.: Mastigobryum ceylanicum Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 105, 1860 [1861] (Mitten 1860c). ** Bazzania chilensis (Steph.) Fulford, Ann. Cryptog. Phytopathol. 3: 51, 1946 (Fulford 1946). Bas.: Mastigobryum chilense Steph., Hedwigia 24 (6): 247, 1885 (Stephani 1885b). *** Bazzania chimantensis Fulford, Bryologist 63 (2): 89, 1960 (Fulford 1960). World checklist of hornworts and liverworts 149 ** Bazzania cincinnata (De Not.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (Trevisan 1877). Bas.: Mastigobryum cincinnatum De Not., Epat. Borneo: 34, 1874 (De Notaris 1874). *** Bazzania citharodes Meagher, Nova Hedwigia 86 (3/4): 481, 2008 (Meagher 2008). ** Bazzania combinata (J.B.Jack et Steph.) Steph., Bot. Jahrb. Syst. 23 (1/2, 3): 306, 1896 (Stephani 1896a). Bas.: Mastigobryum combinatum J.B.Jack et Steph., Bot. Centralbl. 60 (4): 102, 1894 (Jack and Stephani 1894). ** Bazzania commutata (Lindenb. et Gottsche) Schifn., Consp. Hepat. Arch. Ind.: 149, 1898 (Schifner 1898b). Bas.: Mastigobryum commutatum Lindenb. et Gottsche, Sp. Hepat. (Lindenberg) 8-11: 97, 1851 (Lindenberg and Gottsche 1851b). * Bazzania comorensis Steph., Bull. Soc. Roy. Bot. Belgique, Compt. Rend. 30 (2): 197, 1891 [1892] (Stephani 1891b). ** Bazzania confertifolia (Steph.) Herzog, Trans. Brit. Bryol. Soc. 1 (4): 298, 1950 (Herzog 1950a). Bas.: Mastigobryum confertifolium Steph., Bull. Herb. Boissier (sér. 2) 8 (11): 858 (482), 1908 (Stephani 1908a). ** Bazzania conistipula (Steph.) H.A.Mill., Phytologia 47 (4): 320, 1981 (Miller 1981). Bas.: Mastigobryum conistipulum Steph., Sp. Hepat. (Stephani) 6: 458, 1924 (Stephani 1924). ** Bazzania conophylla (Sande Lac.) Schifn., Consp. Hepat. Arch. Ind.: 150, 1898 (Schifner 1898b). Bas.: Mastigobryum conophyllum Sande Lac., Ann. Mus. Bot. Lugduno-Batavi 1: 304, 1864 (Sande Lacoste 1864). ** Bazzania consanguinea (Hampe et Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (Trevisan 1877). Bas.: Mastigobryum consanguineum Hampe et Lindenb., Linnaea 20 (3): 327, 1847 (Hampe 1847). ** Bazzania consociata (Steph.) H.A.Mill., Phytologia 47 (4): 320, 1981 (Miller 1981). Bas.: Mastigobryum consociatum Steph., Sp. Hepat. (Stephani) 6: 458, 1924 (Stephani 1924). * Bazzania corbieri (Steph.) Meagher, Nova Hedwigia 86 (3/4): 483, 2008 (Meagher 2008). Bas.: Mastigobryum corbieri Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 122, 1914 (Stephani and Watts 1914). 108 ** Bazzania crassidentata Fulford, Bull. Torrey Bot. Club 86 (5): 338, 1959 (Fulford 1959b). ** Bazzania crassitexta Steph., Hedwigia 32 (4): 205, 1893 (Stephani 1893c). * Bazzania crenata (Trevis.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 415, 1877 (Trevisan 1877). Bas.: Pleuroschisma crenatum Trevis., Herb. Crypt. Trev. 2: 30, 1853 (Trevisan 1853; non vidi). 109 ** Bazzania cubensis (Gottsche ex Steph.) Pagán, Bryologist 42 (2): 38, 1939 (Pagán 1939b). Bas.: Mastigobryum cubense Gottsche ex Steph., Hedwigia 24 (6): 248, 1885 (Stephani 1885b). 108 Bazzania corbieri may be conspeciic with Bazzania crenata (Meagher 2008). 109 Bazzania crenata from Europe has neither been recognized in any recent treatment nor synonymized. he name may have priority once the identity is established. 150 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Bazzania cucullata Onr., Bull. Jard. Bot. Natl. Belg. 47 (1/2): 142, 1977 (Onraedt 1977). *** Bazzania cuneistipula (Gottsche et Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (Trevisan 1877). Bas.: Mastigobryum cuneistipulum Gottsche et Lindenb., Syn. Hepat. 2: 225, 1845 (Gottsche et al. 1845a). * Bazzania curvidens Steph., Bull. Soc. Roy. Bot. Belgique, Compt. Rend. 30 (2): 197, 1891 [1892] (Stephani 1891b). ** Bazzania debilis N.Kitag., J. Hattori Bot. Lab. 30: 256, 1967 (Kitagawa 1967a). ** Bazzania deciduifolia Onr., Bull. Jard. Bot. Natl. Belg. 47 (1/2): 144, 1977 (Onraedt 1977). *** Bazzania decrescens (Lehm. et Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (Trevisan 1877). Bas.: Jungermannia decrescens Lehm. et Lindenb., Nov. Stirp. Pug. 4: 57, 1832 (Lehmann 1832). 110 ** Bazzania decrescens var. dentistipula Kiaer et Pearson, Forh. Vidensk.-Selsk. Kristiania 1892 (14): 6, 1893 (Pearson 1893). ** Bazzania decrescens subsp. molleri (Steph.) E.W.Jones, J. Bryol. 8 (3): 303, 1975 (Jones 1975). Bas.: Mastigobryum molleri Steph., Bot. Jahrb. Syst. 8 (2): 84, 1886 (Stephani 1886d). ** Bazzania decrescens subsp. pumila (Mitt.) Pócs, Trop. Bryol. 9: 129, 1994 (Pócs 1994c). Bas.: Bazzania pumila Mitt., J. Linn. Soc., Bot. 22 (146): 322, 1886 (Mitten 1886b). ** Bazzania densa (Sande Lac.) Schifn., Consp. Hepat. Arch. Ind.: 151, 1898 (Schifner 1898b). Bas.: Mastigobryum densum Sande Lac., Ned. Kruidk. Arch. 3: 418, 1854 [1855] (Sande Lacoste 1854). ** Bazzania densa var. connata (Sande Lac.) Schifn., Consp. Hepat. Arch. Ind.: 151, 1898 (Schifner 1898b). Bas.: Mastigobryum densum β connatum Sande Lac., Ann. Mus. Bot. Lugduno-Batavi 1: 302, 1864 (Sande Lacoste 1864). *** Bazzania denticulata (Lindenb. et Gottsche) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (Trevisan 1877). Bas.: Mastigobryum denticulatum Lindenb. et Gottsche, Syn. Hepat. 5: 718, 1847 (Gottsche et al. 1847). ** Bazzania denticulifera Mägd., Nova Hedwigia 38: 53, 1983 (Mägdefrau 1983). ** Bazzania denudata (Lindenb. et Gottsche) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (Trevisan 1877). Bas.: Mastigobryum denudatum Lindenb. et Gottsche, Syn. Hepat. 2: 216, 1845 (Gottsche et al. 1845a). *** Bazzania deplanchei (Gottsche) Jovet-Ast, Rev. Bryol. Lichénol. 18 (1/2): 83, 1949 (Jovet-Ast 1949b). Bas.: Mastigobryum deplanchei Gottsche, Bull. Herb. Boissier (sér. 2) 8 (12): 955 (505), 1908 (Stephani 1908b). ** Bazzania deplanchei var. ilamentosa Tixier, Cryptog. Bryol. Lichénol. 6 (2): 179, 1985 (Tixier 1985b). ** Bazzania desciscens (Steph.) Meijer, Blumea 10 (2): 382, 1960 (Meijer 1960). Bas.: Mastigobryum desciscens Steph., Bull. Herb. Boissier (sér. 2) 8 (11): 862 (486), 1908 (Stephani 1908a). 110 Bazzania decrescens is a species complex also including also Bazzania comorensis, Bazzania curvidens and Bazzania mascarena (Grolle 1995). World checklist of hornworts and liverworts 151 ** Bazzania didericiana (Gottsche ex Steph.) Steph., Bull. Herb. Boissier 5 (10): 841, 1897 (Stephani 1897c). Bas.: Mastigobryum didericianum Gottsche ex Steph., Hedwigia 24 (6): 249, 1885 (Stephani 1885b). ** Bazzania diminuta Herzog, Trans. Brit. Bryol. Soc. 1 (4): 305, 1950 (Herzog 1950a). * Bazzania distans (Nees) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (Trevisan 1877). Bas.: Mastigobryum distans Nees, Syn. Hepat. 2: 216, 1845 (Gottsche et al. 1845a). 111 *** Bazzania diversicuspis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 373, 1885 (Spruce 1885). ** Bazzania drepanophylla Herzog, Trans. Brit. Bryol. Soc. 1 (4): 301, 1950 (Herzog 1950a). ** Bazzania dulitensis Herzog, Trans. Brit. Bryol. Soc. 1 (4): 299, 1950 (Herzog 1950a). ** Bazzania dulongensis L.P.Zhou et Li Zhang, J. Bryol. 34 (1): 22, 2012 (Zhou et al. 2012). ** Bazzania eggersiana (Steph.) Pagán, Bryologist 42 (2): 39, 1939 (Pagán 1939b). Bas.: Mastigobryum eggersianum Steph., Bull. Herb. Boissier (sér. 2) 8 (11): 844 (468), 1908 (Stephani 1908a). ** Bazzania elmeri (Steph.) N.Kitag., J. Hattori Bot. Lab. 36: 446, 1972 [1973] (Kitagawa 1972). Bas.: Mastigobryum elmeri Steph., Leal. Philipp. Bot. 6: 2289, 1914 (Stephani 1914a). ** Bazzania elongata Fulford, Bull. Torrey Bot. Club 86 (5): 337, 1959 (Fulford 1959b). ** Bazzania emarginata (Steph.) C.M.Cooke, Trans. Connecticut Acad. Arts 12 (1): 17, 1904 (Cooke 1904). Bas.: Mastigobryum didericianum var. emarginatum Steph., Hedwigia 24 (6): 249, 1885 (Stephani 1885b). ** Bazzania engelii Glenny, Fieldiana, Bot. (n.ser.) 47: 176, 2007 (Glenny and Bartlett 2007). *** Bazzania erosa (Reinw., Blume et Nees) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 415, 1877 (Trevisan 1877). Bas.: Jungermannia erosa Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 230, 1824 [1825] (Reinwardt et al. 1824a). ** Bazzania erosa var. pulopenangensis (Lindenb. et Gottsche) Schifn., Consp. Hepat. Arch. Ind.: 157, 1898 (Schifner 1898b). Bas.: Mastigobryum erosum δ pulopenangense Lindenb. et Gottsche, Sp. Hepat. (Lindenberg) 8-11: 99, 1851 (Lindenberg and Gottsche 1851b). *** Bazzania exempta J.J.Engel, J. Hattori Bot. Lab. 99: 197, 2006 (Engel 2006c). *** Bazzania falcata (Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 415, 1877 (Trevisan 1877). Bas.: Mastigobryum falcatum Lindenb., Syn. Hepat. 2: 231, 1845 (Gottsche et al. 1845a). ** Bazzania falcifolia (Steph.) H.A.Mill., Phytologia 47 (4): 320, 1981 (Miller 1981). Bas.: Mastigobryum falcifolium Steph., Akad. Wiss. Wien, Math.-Naturwiss. Kl., Denkschr. 88: 33, 1911 (Stephani 1911d). 111 Bazzania distans is probably conspeciic with Bazzania loricata (Meijer 1960). 152 * *** ** ** *** ** ** * ** *** ** *** *** *** ** ** ** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Bazzania fallax (Sande Lac.) Schifn., Consp. Hepat. Arch. Ind.: 158, 1898 (Schifner 1898b). Bas.: Mastigobryum fallax Sande Lac., Ann. Mus. Bot. Lugduno-Batavi 1: 304, 1864 (Sande Lacoste 1864). Bazzania fasciculata (Steph.) Meagher, Australas. Bryol. Newslett. 46: 6, 2002 (Meagher 2002). Bas.: Mastigobryum fasciculatum Steph., Bull. Herb. Boissier (sér. 2) 8 (10): 748 (432), 1908 (Stephani 1908c). Bazzania fauriana (Steph.) S.Hatt., Bot. Mag. (Tokyo) 59 (693/694): 27, 1946 (Hattori 1946). Bas.: Mastigobryum faurianum Steph., Bull. Herb. Boissier (sér. 2) 8 (11): 843 (467), 1908 (Stephani 1908a). Bazzania iliformis Steph., Hedwigia 28 (2): 131, 1889 (Stephani 1889a). Bazzania laccida (Dumort.) Grolle, Lindbergia 1 (3/4): 200, 1972 [1973] (Grolle 1972b). Bas.: Pleuroschisma laccidum Dumort., Syll. Jungerm. Europ.: 71, 1831 (Dumortier 1831). Bazzania lavescens (Sande Lac. ex Steph.) Schifn., Consp. Hepat. Arch. Ind.: 158, 1898 (Schifner 1898b). Bas.: Mastigobryum lavescens Sande Lac. ex Steph., Hedwigia 25 (1): 6, 1886 (Stephani 1886e). Bazzania leischeri (Steph.) Abeyw., Ceylon J. Sci., Biol. Sci. 2 (1): 45, 1959 (Abeywickrama 1959). Bas.: Mastigobryum leischeri Steph., Bull. Herb. Boissier (sér. 2) 8 (10): 773 (457), 1908 (Stephani 1908c). Bazzania francana (Steph.) N.Kitag., J. Hattori Bot. Lab. 36: 446, 1972 [1973] (Kitagawa 1972). Bas.: Mastigobryum francanum Steph., Sp. Hepat. (Stephani) 6: 463, 1924 (Stephani 1924). Bazzania friabilis N.Kitag. et T.Kodama, J. Hattori Bot. Lab. 39: 67, 1975 (Kitagawa and Kodama 1975b). Bazzania fuhreri Meagher, Nova Hedwigia 92 (3/4): 488, 2011 (Meagher 2011). Bazzania fuscescens A.Evans, Pap. Michigan Acad. Sci. 17: 85, 1932 [1933] (Evans 1932b). Bazzania gamscottii Meagher, Nova Hedwigia 92 (3/4): 492, 2011 (Meagher 2011). Bazzania gedeana (Steph.) Meijer, Blumea 10 (2): 378, 1960 (Meijer 1960). Bas.: Mastigobryum gedeanum Steph., Sp. Hepat. (Stephani) 3: 540, 1909 (Stephani 1909a). Bazzania gracilis (Hampe et Gottsche) Steph., Hedwigia 27 (11/12): 279, 1888 (Stephani 1888c). Bas.: Mastigobryum gracile Hampe et Gottsche, Linnaea 25 (3): 346, 1852 [1853] (Hampe and Gottsche 1852). Bazzania grandiretis (Steph.) Herzog, Trans. Brit. Bryol. Soc. 1 (4): 298, 1950 (Herzog 1950a). Bas.: Mastigobryum grandirete Steph., Bull. Herb. Boissier (sér. 2) 8 (10): 747 (431), 1908 (Stephani 1908c). Bazzania griithiana (Steph.) Mizut., J. Hattori Bot. Lab. 30: 82, 1967 (Mizutani 1967). Bas.: Mastigobryum griithianum Steph., Bull. Herb. Boissier (sér. 2) 8 (12): 959 (509), 1908 (Stephani 1908b). Bazzania gunniana (Steph.) H.A.Mill., Phytologia 47 (4): 320, 1981 (Miller 1981). Bas.: Mastigobryum gunnianum Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 123, 1914 (Stephani and Watts 1914). World checklist of hornworts and liverworts 153 ** Bazzania hainanensis L.P.Zhou et Li Zhang, J. Bryol. 34 (1): 25, 2012 (Zhou et al. 2012). ** Bazzania halconiensis (Steph.) N.Kitag., J. Hattori Bot. Lab. 36: 447, 1972 [1973] (Kitagawa 1972). Bas.: Mastigobryum halconiense Steph., Bull. Herb. Boissier (sér. 2) 8 (10): 759 (443), 1908 (Stephani 1908c). ** Bazzania hamatifolia (Steph.) H.A.Mill., Phytologia 47 (4): 320, 1981 (Miller 1981). Bas.: Mastigobryum hamatifolium Steph., Bull. Herb. Boissier (sér. 2) 8 (11): 862 (486), 1908 (Stephani 1908a). ** Bazzania harpago (De Not.) Schifn., Consp. Hepat. Arch. Ind.: 159, 1898 (Schifner 1898b). Bas.: Mastigobryum harpago De Not., Epat. Borneo: 29, 1874 (De Notaris 1874). ** Bazzania hebridensis (Steph.) H.A.Mill., Phytologia 47 (4): 320, 1981 (Miller 1981). Bas.: Mastigobryum hebridense Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 124, 1914 (Stephani and Watts 1914). ** Bazzania herminieri (Gottsche ex Steph.) Pagán, Bryologist 45 (4): 90, 1942 (Pagán 1942b). Bas.: Mastigobryum herminieri Gottsche ex Steph., Hedwigia 25 (1): 8, 1886 (Stephani 1886e). ** Bazzania herzogiana Meijer, Blumea 10 (2): 371, 1960 (Meijer 1960). Nom. nov. pro Bazzania remotifolia Herzog, Trans. Brit. Bryol. Soc. 1 (4): 304, 1950 (Herzog 1950a), nom. illeg. ** Bazzania heterostipa (Steph.) Fulford, Bull. Torrey Bot. Club 86 (6): 410, 1959 (Fulford 1959a). Bas.: Mastigobryum heterostipum Steph., Sp. Hepat. (Stephani) 3: 532, 1909 (Stephani 1909a). *** Bazzania himalayana (Mitt.) Schifn., Österr. Bot. Z. 49 (4): 132, 1899 (Schifner 1899b). Bas.: Mastigobryum himalayanum Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 105, 1860 [1861] (Mitten 1860c). *** Bazzania hochstetteri (Reichardt) E.A.Hodgs., Trans. Roy. Soc. New Zealand 82 (1): 11, 1954 (Hodgson 1954). Bas.: Mastigobryum hochstetteri Reichardt, Verh. K.K. Zool.-Bot. Ges. Wien 16: 959, 1866 (Reichardt 1866). *** Bazzania hookeri (Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (Trevisan 1877). Bas.: Mastigobryum hookeri Lindenb., Syn. Hepat. 2: 226, 1845 (Gottsche et al. 1845a). ** Bazzania horridula Schifn., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 60 (2): 258, 1893 (Schifner 1893a). ** Bazzania inaequabilis Steph., Trans. Connecticut Acad. Arts 12 (1): 21, 1904 (Cooke 1904). ** Bazzania inaequitexta Steph., Hedwigia 32 (4): 208, 1893 (Stephani 1893c). ** Bazzania incrassata (Steph.) N.Kitag., J. Hattori Bot. Lab. 36: 448, 1972 [1973] (Kitagawa 1972). Bas.: Mastigobryum incrassatum Steph., Sp. Hepat. (Stephani) 6: 469, 1924 (Stephani 1924). ** Bazzania indica (Gottsche et Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 415, 1877 (Trevisan 1877). Bas.: Mastigobryum indicum Gottsche et Lindenb., Syn. Hepat. 2: 230, 1845 (Gottsche et al. 1845a). 154 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Bazzania indigenarum (Steph.) N.Kitag., Bull. Nara Univ. Educ., B 26 (2): 82, 1977 (Kitagawa 1977). Bas.: Mastigobryum indigenarum Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 124, 1914 (Stephani and Watts 1914). ** Bazzania insignis (De Not.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (Trevisan 1877). Bas.: Mastigobryum insigne De Not., Epat. Borneo: 26, 1874 (De Notaris 1874). * Bazzania intermedia (Gottsche et Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 415, 1877 (Trevisan 1877). Bas.: Mastigobryum intermedium Gottsche et Lindenb., Sp. Hepat. (Lindenberg) 8-11: 82, 1851 (Lindenberg and Gottsche 1851b). ** Bazzania intermedia var. sarawakiana (De Not.) Schifn., Consp. Hepat. Arch. Ind.: 162, 1898 (Schifner 1898b). Bas.: Mastigobryum intermedium var. sarawakianum De Not., Epat. Borneo: 32, 1874 (De Notaris 1874). ** Bazzania involuta (Mont.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (Trevisan 1877). Bas.: Herpetium involutum Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 253, 1843 (Montagne 1843). ** Bazzania involuta var. submutica (Lindenb. et Gottsche) J.J.Engel et G.L.Merr., Bryologist 97 (3): 314, 1994 (Engel and Smith Merrill 1994). Bas.: Mastigobryum novae-hollandiae γ3 submuticum Lindenb. et Gottsche, Sp. Hepat. (Lindenberg) 8-11: 33, 1851 (Lindenberg and Gottsche 1851b). ** Bazzania involutiformis (De Not.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (Trevisan 1877). Bas.: Mastigobryum involutiforme De Not., Epat. Borneo: 28, 1874 (De Notaris 1874). ** Bazzania irregularis (Steph.) Schifn., Consp. Hepat. Arch. Ind.: 163, 1898 (Schifner 1898b). Bas.: Mastigobryum irregulare Steph., Hedwigia 25 (4): 133, 1886 (Stephani 1886b). *** Bazzania jamaicensis (Lehm. et Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (Trevisan 1877). Bas.: Herpetium jamaicense Lehm. et Lindenb., Nov. Stirp. Pug. 7: 7, 1838 (Lehmann 1838). ** Bazzania japonica (Sande Lac.) Lindb., Acta Soc. Sci. Fenn. 10: 224, 1872 [1873] (Lindberg 1872b). Bas.: Mastigobryum japonicum Sande Lac., Ann. Mus. Bot. Lugduno-Batavi 1: 303, 1864 (Sande Lacoste 1864). ** Bazzania japonica var. sumatrana Herzog, Ann. Naturhist. Mus. Wien 53 (1): 366, 1942 [1943] (Herzog 1942b). ** Bazzania javanica (Sande Lac.) Schifn., Consp. Hepat. Arch. Ind.: 163, 1898 (Schifner 1898b). Bas.: Mastigobryum javanicum Sande Lac., Ned. Kruidk. Arch. 3: 418, 1854 [1855] (Sande Lacoste 1854). ** Bazzania kernii Steph., Hedwigia 32 (4): 208, 1893 (Stephani 1893c). ** Bazzania kokawana N.Kitag. et T.Kodama, J. Jap. Bot. 50 (1): 11, 1975 (Kitagawa and Kodama 1975a). ** Bazzania latifolia Steph., Hedwigia 32 (4): 209, 1893 (Stephani 1893c). World checklist of hornworts and liverworts * ** ** ** ** ** ** *** ** *** ** *** *** ** 155 Bazzania lehmanniana (Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (Trevisan 1877). Bas.: Mastigobryum lehmannianum Lindenb., Syn. Hepat. 2: 223, 1845 (Gottsche et al. 1845a). 112 Bazzania leratii (Beauverd) H.A.Mill., Phytologia 47 (4): 320, 1981 (Miller 1981). Bas.: Mastigobryum leratii Beauverd, Sp. Hepat. (Stephani) 6: 477, 1924 (Stephani 1924). Bazzania lessonii (Steph.) H.A.Mill., Phytologia 47 (4): 321, 1981 (Miller 1981). Bas.: Mastigobryum lessonii Steph., Sp. Hepat. (Stephani) 3: 531, 1909 (Stephani 1909a). Bazzania levieri (Steph.) N.Kitag., J. Hattori Bot. Lab. 36: 448, 1972 [1973] (Kitagawa 1972). Bas.: Mastigobryum levieri Steph., Bull. Herb. Boissier (sér. 2) 8 (12): 944 (494), 1908 (Stephani 1908b). Bazzania liebmaniana (Lindenb. et Gottsche) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (Trevisan 1877). Bas.: Mastigobryum liebmanianum Lindenb. et Gottsche, Syn. Hepat. 5: 719, 1847 (Gottsche et al. 1847). Bazzania linearis Herzog, Trans. Brit. Bryol. Soc. 1 (4): 303, 1950 (Herzog 1950a). Bazzania linguiformis (Sande Lac.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (Trevisan 1877). Bas.: Mastigobryum linguiforme Sande Lac., Plagiochila Sandei: 8, 1856 (Sande Lacoste 1856c). Bazzania longa (Nees) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 415, 1877 (Trevisan 1877). Bas.: Jungermannia longa Nees, Linnaea 6 (4): 623, 1831 (Nees 1831). Bazzania longa var. papillata (Steph.) Fulford, Ann. Cryptog. Phytopathol. 3: 94, 1946 (Fulford 1946). Bas.: Mastigobryum papillatum Steph., Sp. Hepat. (Stephani) 3: 526, 1909 (Stephani 1909a). Bazzania longicaulis (Sande Lac.) Schifn., Consp. Hepat. Arch. Ind.: 165, 1898 (Schifner 1898b). Bas.: Mastigobryum longicaule Sande Lac., Ann. Mus. Bot. Lugduno-Batavi 1: 303, 1864 (Sande Lacoste 1864). Bazzania longicaulis var. latiareata Herzog, Memoranda Soc. Fauna Fl. Fennica 26: 46, 1950 [1951] (Herzog 1950b). Bazzania longistipula (Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 415, 1877 (Trevisan 1877). Bas.: Mastigobryum longistipulum Lindenb., Syn. Hepat. 2: 228, 1845 (Gottsche et al. 1845a). Bazzania loricata (Reinw., Blume et Nees) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (Trevisan 1877). Bas.: Jungermannia loricata Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 233, 1824 [1825] (Reinwardt et al. 1824a). Bazzania lowii (Sande Lac. ex Steph.) Schifn., Consp. Hepat. Arch. Ind.: 166, 1898 (Schifner 1898b). Bas.: Mastigobryum lowii Sande Lac. ex Steph., Hedwigia 25 (5): 204, 1886 (Stephani 1886f). 112 Bazzania lehmanniana is possibly conspeciic with Bazzania arcuata or Bazzania longa (Meagher 2012). 156 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Bazzania luzonensis (Steph.) Del Ros., Philipp. J. Sci. 100 (3/4): 231, 1971 (Del Rosario 1971). Bas.: Mastigobryum luzonense Steph., Sp. Hepat. (Stephani) 6: 472, 1924 (Stephani 1924). ** Bazzania macgregorii Steph., Hedwigia 32 (4): 210, 1893 (Stephani 1893c). ** Bazzania macrostipula Fulford, Bull. Torrey Bot. Club 86 (6): 407, 1959 (Fulford 1959a). ** Bazzania magna Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 197, 1934 (Horikawa 1934). ** Bazzania magnistipula N.Kitag., J. Hattori Bot. Lab. 47: 132, 1980 (Kitagawa 1980). ** Bazzania malaccensis (Steph.) Tixier, Gard. Bull. Singapore 25 (3): 342, 1971 (Tixier 1971). Bas.: Mastigobryum malaccense Steph., Bull. Herb. Boissier (sér. 2) 8 (12): 944 (494), 1908 (Stephani 1908b). ** Bazzania manillana (Gottsche ex Steph.) S.Hatt., Bot. Mag. (Tokyo) 64 (755/756): 113, 1951 (Hattori 1951c). Bas.: Mastigobryum manillanum Gottsche ex Steph., Hedwigia 25 (5): 204, 1886 (Stephani 1886f). * Bazzania marginata (Steph.) N.Kitag., J. Hattori Bot. Lab. 36: 449, 1972 [1973] (Kitagawa 1972). Bas.: Mastigobryum marginatum Steph., Rev. Bryol. 35 (2): 31, 1908 (Stephani 1908l). ** Bazzania marginella (Herzog) N.Kitag. et T.Kodama, Bull. Osaka Mus. Nat. Hist. 27: 17, 1973 (Kitagawa and Kodama 1973). Bas.: Mastigobryum marginellum Herzog, Ann. Bryol. 5: 91, 1932 (Herzog 1932a). * Bazzania mascarena (Steph.) Herzog, Bot. Not. 100 (4): 334, 1947 (Herzog 1947). Bas.: Mastigobryum mascarenum Steph., Hedwigia 25 (5): 205, 1886 (Stephani 1886f). ** Bazzania mayebarae S.Hatt., J. Hattori Bot. Lab. 19: 91, 1958 (Hattori and Mizutani 1958). * Bazzania menzelii E.D.Cooper, Phytotaxa 97 (2): 52, 2013 (Cooper et al. 2013). Nom. nov. pro Acromastigum emarginatum Herzog, Trans. Brit. Bryol. Soc. 1 (4): 309, 1950 (Herzog 1950a). ** Bazzania merrillana (Steph.) Inoue ex Bonner, Index Hepat. 3: 359, 1963 (Bonner 1963). Bas.: Mastigobryum merrillanum Steph., Bull. Herb. Boissier (sér. 2) 8 (12): 944 (494), 1908 (Stephani 1908b). ** Bazzania minuta (Austin) A.Evans, Trans. Connecticut Acad. Arts 8 (15): 255, 1891 (Evans 1891). Bas.: Mastigobryum minutum Austin, Bull. Torrey Bot. Club 5 (3): 17, 1874 (Austin 1874). ** Bazzania minutidens (Steph.) Inoue et H.A.Mill., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 8 (2): 142, 1965 (Inoue and Miller 1965). Bas.: Mastigobryum minutidens Steph., Sp. Hepat. (Stephani) 6: 474, 1924 (Stephani 1924). ** Bazzania minutiserra (Steph.) N.Kitag., Bull. Nara Univ. Educ., B 26 (2): 77, 1977 (Kitagawa 1977). Bas.: Mastigobryum minutiserrum Steph., Sp. Hepat. (Stephani) 6: 474, 1924 (Stephani 1924). World checklist of hornworts and liverworts 157 ** Bazzania missionum (Herzog) Jovet-Ast, Rev. Bryol. Lichénol. 20 (1/2): 96, 1951 (Jovet-Ast 1951). Bas.: Mastigobryum missionum Herzog, Ann. Bryol. 5: 93, 1932 (Herzog 1932a). ** Bazzania mittenii (Steph.) Steph., Hedwigia 28 (2): 132, 1889 (Stephani 1889a). Bas.: Mastigobryum mittenii Steph., Hedwigia 25 (6): 245, 1886 (Stephani 1886c). *** Bazzania monilinervis (Lehm. et Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (Trevisan 1877). Bas.: Jungermannia monilinervis Lehm. et Lindenb., Nov. Stirp. Pug. 4: 56, 1832 (Lehmann 1832). ** Bazzania morokensis (Steph.) Grolle, J. Hattori Bot. Lab. 31: 1, 1968 (Grolle 1968a). Bas.: Mastigobryum morokense Steph., Bull. Herb. Boissier (sér. 2) 8 (10): 765 (449), 1908 (Stephani 1908c). *** Bazzania nitida (F.Weber) Grolle, Rev. Bryol. Lichénol. 29 (3/4): 210, 1960 [1961] (Grolle 1960a). Bas.: Jungermannia nitida F.Weber, Hist. Musc. Hepat. Prodr.: 43, 1815 (Weber 1815). *** Bazzania novae-zelandiae (Mitt.) Besch. et C.Massal., Miss. sci. Cape Horn, Bot. 5: 233, 1889 (Bescherelle and Massalongo 1889). Bas.: Mastigobryum novae-zelandiae Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 148, 1854 (Mitten 1854). ** Bazzania nudicaulis A.Evans, Bryologist 26 (6): 62, 1923 [1924] (Evans 1923b). ** Bazzania nuuanuensis C.M.Cooke, Trans. Connecticut Acad. Arts 12 (1): 15, 1904 (Cooke 1904). ** Bazzania obcuneata (Steph.) H.A.Mill., Phytologia 47 (4): 321, 1981 (Miller 1981). Bas.: Mastigobryum obcuneatum Steph., Bull. Herb. Boissier (sér. 2) 8 (11): 863 (487), 1908 (Stephani 1908a). ** Bazzania obtusata (Mitt.) Abeyw., Ceylon J. Sci., Biol. Sci. 2 (1): 45, 1959 (Abeywickrama 1959). Bas.: Mastigobryum obtusatum Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 106, 1860 [1861] (Mitten 1860c). ** Bazzania okaritana Meagher et Glenny, J. Bryol. 29 (1): 60, 2007 (Meagher and Glenny 2007). ** Bazzania orbanii Pócs, Acta Biol. Pl. Agr. 1: 16, 2010 [2011] (Pócs 2010c). ** Bazzania ovistipula (Steph.) Abeyw., Ceylon J. Sci., Biol. Sci. 2 (1): 45, 1959 (Abeywickrama 1959). Bas.: Mastigobryum ovistipulum Steph., Bull. Herb. Boissier (sér. 2) 8 (10): 760 (444), 1908 (Stephani 1908c). ** Bazzania pallidevirens (Steph.) Fulford, Ann. Cryptog. Phytopathol. 3: 42, 1946 (Fulford 1946). Bas.: Mastigobryum pallidevirens Steph., Bull. Herb. Boissier (sér. 2) 8 (11): 849 (473), 1908 (Stephani 1908a). ** Bazzania papillosa S.W.Arnell, Svensk Bot. Tidskr. 59 (1): 67, 1965 (Arnell 1965). ** Bazzania paradoxa (Sande Lac.) Steph., Bot. Jahrb. Syst. 23 (1/2, 3): 307, 1896 (Stephani 1896a). Bas.: Mastigobryum paradoxum Sande Lac., Ned. Kruidk. Arch. 3: 419, 1854 [1855] (Sande Lacoste 1854). *** Bazzania parisii (Steph.) N.Kitag., J. Hattori Bot. Lab. 47: 135, 1980 (Kitagawa 1980). Bas.: Mastigobryum parisii Steph., Bull. Herb. Boissier (sér. 2) 8 (10): 769 (453), 1908 (Stephani 1908c). 158 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Bazzania parvitexta Steph., Hedwigia 32 (4): 211, 1893 (Stephani 1893c). ** Bazzania patens (Mont.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (Trevisan 1877). Bas.: Herpetium patens Mont., Ann. Sci. Nat. Bot. (sér. 2) 20: 295, 1844 (Montagne 1844b). ** Bazzania patentistipa (Sande Lac.) Schifn., Consp. Hepat. Arch. Ind.: 168, 1898 (Schifner 1898b). Bas.: Mastigobryum patentistipum Sande Lac., Ann. Mus. Bot. Lugduno-Batavi 1: 302, 1864 (Sande Lacoste 1864). ** Bazzania paucidens (Steph.) H.A.Mill., Phytologia 47 (4): 321, 1981 (Miller 1981). Bas.: Mastigobryum paucidens Steph., Bull. Herb. Boissier (sér. 2) 8 (11): 860 (484), 1908 (Stephani 1908a). ** Bazzania pearsonii Steph., Hedwigia 32 (4): 212, 1893 (Stephani 1893c). *** Bazzania pectinata (Lindenb. et Gottsche) Schifn., Nova Acta Acad. Caes. Leop.Carol. German. Nat. Cur. 60 (2): 259, 1893 (Schifner 1893a). Bas.: Mastigobryum pectinatum Lindenb. et Gottsche, Sp. Hepat. (Lindenberg) 8-11: 84, 1851 (Lindenberg and Gottsche 1851b). ** Bazzania perfalcata N.Kitag., J. Hattori Bot. Lab. 47: 135, 1980 (Kitagawa 1980). ** Bazzania perrotana E.W.Jones, J. Bryol. 8 (3): 310, 1975 (Jones 1975). Nom. nov. pro Mastigobryum perrotanum Steph., Sp. Hepat. (Stephani) 6: 476, 1924 (Stephani 1924), nom. illeg. ** Bazzania peruviana (Nees) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (Trevisan 1877). Bas.: Mastigobryum peruvianum Nees, Syn. Hepat. 2: 220, 1845 (Gottsche et al. 1845a). *** Bazzania phyllobola Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 372, 1885 (Spruce 1885). *** Bazzania placophylla (Taylor) Grolle, Rev. Bryol. Lichénol. 27 (1/2): 54, 1958 (Grolle 1958). Bas.: Jungermannia placophylla Taylor, London J. Bot. 5: 276, 1846 (Taylor 1846a). ** Bazzania platycnema (Schwägr. ex Steph.) H.A.Mill., Bryologist 63 (2): 121, 1960 (Miller 1960). Bas.: Mastigobryum platycnemum Schwägr. ex Steph., Bull. Herb. Boissier (sér. 2) 8 (10): 776 (460), 1908 (Stephani 1908c). ** Bazzania pompeana (Sande Lac.) Mitt., Trans. Linn. Soc. London, Bot. 3 (3): 200, 1891 (Mitten 1891). Bas.: Mastigobryum pompeanum Sande Lac., Ann. Mus. Bot. Lugduno-Batavi 1: 304, 1864 (Sande Lacoste 1864). *** Bazzania praerupta (Reinw., Blume et Nees) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (Trevisan 1877). Bas.: Jungermannia praerupta Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 229, 1824 [1825] (Reinwardt et al. 1824a). * Bazzania praerupta var. obliquata (Nees) Schifn., Consp. Hepat. Arch. Ind.: 170, 1898 (Schifner 1898b). Bas.: Jungermannia obliquata Nees, Enum. Pl. Crypt. Javae: 62, 1830 (Nees 1830). ** Bazzania praerupta var. thermalis Schifn., Arch. Hydrobiol., suppl. 21 (3/4): 396, 1955 (Schifner 1955). World checklist of hornworts and liverworts 159 ** Bazzania pseudovittata N.Kitag. et T.Kodama, J. Hattori Bot. Lab. 39: 69, 1975 (Kitagawa and Kodama 1975b). ** Bazzania pusilla (Mitt.) Steph., Bot. Jahrb. Syst. 23 (1/2, 3): 307, 1896 (Stephani 1896a). Bas.: Mastigobryum pusillum Mitt., Fl. vit.: 406, 1871 [1873] (Mitten 1871). ** Bazzania pycnophylla (Taylor) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (Trevisan 1877). Bas.: Mastigobryum pycnophyllum Taylor, London J. Bot. 5: 371, 1846 (Taylor 1846b). ** Bazzania quadratistipula H.A.Mill., Phytologia 47 (4): 321, 1981 (Miller 1981). Nom. nov. pro Mastigobryum quadratum Steph., Sp. Hepat. (Stephani) 6: 477, 1924 (Stephani 1924), nom. illeg. ** Bazzania rabenhorstii (Steph.) Abeyw., Ceylon J. Sci., Biol. Sci. 2 (1): 45, 1959 (Abeywickrama 1959). Bas.: Mastigobryum rabenhorstii Steph., Bull. Herb. Boissier (sér. 2) 8 (10): 774 (458), 1908 (Stephani 1908c). ** Bazzania recurva (Mont.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (Trevisan 1877). Bas.: Herpetium recurvum Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 253, 1843 (Montagne 1843). * Bazzania recurva var. major (Sande Lac.) Schifn., Consp. Hepat. Arch. Ind.: 172, 1898 (Schifner 1898b). Bas.: Mastigobryum recurvum var. majus Sande Lac., Ann. Mus. Bot. Lugduno-Batavi 1: 304, 1864 (Sande Lacoste 1864). ** Bazzania relexa (Gottsche) Steph., Rev. Bryol. 18 (4): 56, 1891 (Renauld and Cardot 1891). Bas.: Mastigobryum relexum Gottsche, Abh. Naturwiss. Vereins Bremen 7: 347, 1882 (Gottsche 1882). ** Bazzania reinwardtii (Sande Lac.) Schifn., Consp. Hepat. Arch. Ind.: 172, 1898 (Schifner 1898b). Bas.: Mastigobryum reinwardtii Sande Lac., Pl. Ind. Batav. Orient.: 22, 1856 (Sande Lacoste 1856a). ** Bazzania renistipula Steph., Hedwigia 32 (4): 212, 1893 (Stephani 1893c). ** Bazzania revoluta (Steph.) N.Kitag., J. Hattori Bot. Lab. 36: 450, 1972 [1973] (Kitagawa 1972). Bas.: Mastigobryum revolutum Steph., Bull. Herb. Boissier (sér. 2) 8 (12): 961 (511), 1908 (Stephani 1908b). *** Bazzania rimosa Meagher, Nova Hedwigia 86 (3/4): 489, 2008 (Meagher 2008). ** Bazzania roccatii Gola, Ann. Bot. (Rome) 6 (2): 273, 1907 (Gola 1907). *** Bazzania roraimensis (Steph.) Fulford, Ann. Cryptog. Phytopathol. 3: 27, 1946 (Fulford 1946). Bas.: Mastigobryum roraimense Steph., Trans. Linn. Soc. London, Bot. 6 (1): 97, 1901 (Stephani 1901e). ** Bazzania sandvicensis (Gottsche ex Steph.) Steph., Bull. Herb. Boissier 5 (10): 841, 1897 (Stephani 1897c). Bas.: Mastigobryum sandvicense Gottsche ex Steph., Hedwigia 25 (5): 207, 1886 (Stephani 1886f). ** Bazzania sauropoda Meagher, Austrobaileya 7 (1): 129, 2005 (Meagher 2005a). *** Bazzania scalaris Meagher, Telopea 11 (3): 247, 2006 (Meagher 2006). *** Bazzania schlimiana (Gottsche) Fulford, Bull. Torrey Bot. Club 86 (6): 401, 1959 (Fulford 1959a). Bas.: Mastigobryum schlimianum Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 140, 1864 (Gottsche 1864). ** Bazzania schultze-motelii N.Kitag., J. Hattori Bot. Lab. 47: 138, 1980 (Kitagawa 1980). 160 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Bazzania schusterana N.Kitag., J. Hattori Bot. Lab. 47: 139, 1980 (Kitagawa 1980). *** Bazzania schwaneckiana (Hampe et Gottsche) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (Trevisan 1877). Bas.: Mastigobryum schwaneckianum Hampe et Gottsche, Linnaea 25 (3): 345, 1852 [1853] (Hampe and Gottsche 1852). * Bazzania scutigera (Nees et Mont.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (Trevisan 1877). Bas.: Herpetium scutigerum Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 9: 44, 1838 (Montagne 1838). ** Bazzania semicordata (Lindenb. et Gottsche) Kuntze, Revis. Gen. Pl. 2: 832, 1891 (Kuntze 1891). Bas.: Mastigobryum semicordatum Lindenb. et Gottsche, Syn. Hepat. 5: 720, 1847 (Gottsche et al. 1847). * Bazzania serpentina (Nees) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 415, 1877 (Trevisan 1877). Bas.: Jungermannia serpentina Nees, Enum. Pl. Crypt. Javae: 62, 1830 (Nees 1830). 113 ** Bazzania serrapiculata Inoue et H.A.Mill., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 8 (2): 141, 1965 (Inoue and Miller 1965). ** Bazzania serrata Fulford, Bull. Torrey Bot. Club 86 (5): 321, 1959 (Fulford 1959b). ** Bazzania serrulatoides Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 200, 1934 (Horikawa 1934). * Bazzania sikkimensis (Steph.) Herzog, Ann. Bryol. 12: 78, 1939 (Herzog 1939b). Bas.: Mastigobryum sikkimense Steph., Hedwigia 44 (2): 73, 1905 (Stephani 1905h). 114 ** Bazzania spinosa S.Okamura, J. Coll. Sci. Imp. Univ. Tokyo 38 (4): 2, 1916 (Okamura 1916). ** Bazzania spiralis (Reinw., Blume et Nees) Meijer, Blumea 10 (2): 381, 1960 (Meijer 1960). Bas.: Jungermannia spiralis Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 231, 1824 [1825] (Reinwardt et al. 1824a). ** Bazzania spruceana Steph., Hedwigia 32 (4): 213, 1893 (Stephani 1893c). ** Bazzania squarrosa (Steph.) H.A.Mill., Phytologia 47 (4): 321, 1981 (Miller 1981). Bas.: Mastigobryum squarrosum Steph., Bull. Herb. Boissier (sér. 2) 8 (10): 776 (460), 1908 (Stephani 1908c). *** Bazzania stolonifera (Sw.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 415, 1877 (Trevisan 1877). Bas.: Jungermannia stolonifera Sw., Prodr. (Swartz): 144, 1788 (Swartz 1788). * Bazzania stolonifera var. granatensis (Gottsche) Fulford, Ann. Cryptog. Phytopathol. 3: 51, 1946 (Fulford 1946). Bas.: Mastigobryum stoloniferum var. granatense Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 141, 1864 (Gottsche 1864). ** Bazzania stresemannii (Herzog) N.Kitag., Bull. Nara Univ. Educ., B 28 (2): 77, 1979 (Kitagawa 1979a). Bas.: Mastigobryum stresemannii Herzog, Beih. Bot. Centralbl. 38 (2): 324, 1921 (Herzog 1921). 113 Bazzania serpentina is possibly conspeciic with Bazzania bidens (Frahm et al. 1990). 114 Bazzania sikkimensis is possibly conspeciic with Bazzania sumbavensis (Long and Grolle 1990). World checklist of hornworts and liverworts 161 ** Bazzania subacuta (Mitt.) Steph., Bot. Jahrb. Syst. 23 (1/2, 3): 307, 1896 (Stephani 1896a). Bas.: Mastigobryum subacutum Mitt., Fl. vit.: 406, 1871 [1873] (Mitten 1871). ** Bazzania subaequitexta (Steph.) N.Kitag., Bull. Nara Univ. Educ., B 28 (2): 81, 1979 (Kitagawa 1979a). Bas.: Mastigobryum subaequitextum Steph., Bull. Herb. Boissier (sér. 2) 8 (11): 859 (483), 1908 (Stephani 1908a). ** Bazzania subintegra (Steph.) L.Söderstr. et A.Hagborg, Phytotaxa 202 (1): 69, 2015 (Söderström et al. 2015c). Bas.: Mastigobryum subintegrum Steph., Bull. Herb. Boissier (sér. 2) 8 (10): 775 (459), 1908 (Stephani 1908c). ** Bazzania sublonga Fulford, Bull. Torrey Bot. Club 86 (5): 334, 1959 (Fulford 1959b). ** Bazzania subserrifolia (Beauverd) H.A.Mill., Phytologia 47 (4): 321, 1981 (Miller 1981). Bas.: Mastigobryum subserrifolium Beauverd, Sp. Hepat. (Stephani) 6: 480, 1924 (Stephani 1924). ** Bazzania subserrulata A.Evans, Pap. Michigan Acad. Sci. 17: 97, 1932 [1933] (Evans 1932b). *** Bazzania subtilis (Sande Lac.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (Trevisan 1877). Bas.: Mastigobryum subtile Sande Lac., Ann. Mus. Bot. Lugduno-Batavi 1: 302, 1864 (Sande Lacoste 1864). ** Bazzania succulenta N.Kitag., J. Hattori Bot. Lab. 47: 141, 1980 (Kitagawa 1980). ** Bazzania sumatrana (Sande Lac. ex Steph.) Steph., Hedwigia 32 (4): 209, 1893 (Stephani 1893c). Bas.: Mastigobryum sumatranum Sande Lac. ex Steph., Hedwigia 25 (6): 234, 1886 (Stephani 1886c). * Bazzania sumbavensis (Gottsche ex Steph.) Steph., Hedwigia 32 (4): 204, 1893 (Stephani 1893c). Bas.: Mastigobryum sumbavense Gottsche ex Steph., Hedwigia 25 (6): 236, 1886 (Stephani 1886c). 115 *** Bazzania taleana (Gottsche) Fulford, Ann. Cryptog. Phytopathol. 3: 54, 1946 (Fulford 1946). Bas.: Mastigobryum taleanum Gottsche, Mexik. Leverm.: 131, 1863 (Gottsche 1863). *** Bazzania tayloriana (Mitt.) Kuntze, Revis. Gen. Pl. 2: 832, 1891 (Kuntze 1891). Bas.: Mastigobryum taylorianum Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 147, 1854 (Mitten 1854). ** Bazzania temariana (Steph.) H.A.Mill., Phytologia 47 (4): 321, 1981 (Miller 1981). Bas.: Mastigobryum temarianum Steph., Sp. Hepat. (Stephani) 3: 532, 1909 (Stephani 1909a). *** Bazzania tessellata Meagher, Nova Hedwigia 92 (3/4): 492, 2011 (Meagher 2011). ** Bazzania tiaoloensis Mizut. et K.C.Chang, J. Hattori Bot. Lab. 60: 432, 1986 (Mizutani and Chang 1986). *** Bazzania tricrenata (Wahlenb.) Lindb., Musci Fenn. Exsic., fasc. 2: [2 (adnot.)], 1872 (Brotherus 1872). Bas.: Jungermannia tricrenata Wahlenb., Fl. Carpat. Princ.: 364, 1814 (Wahlenberg 1814). ** Bazzania tricrenata var. fulfordiae W.S.Hong, Bryologist 91 (4): 331, 1988 (Hong 1988). 115 Bazzania sumbavensis is possibly conspeciic with Bazzania japonica (Söderström et al. 2010a). 162 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Bazzania tridens (Reinw., Blume et Nees) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 415, 1877 (Trevisan 1877). Bas.: Jungermannia tridens Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 228, 1824 [1825] (Reinwardt et al. 1824a). *** Bazzania tridens var. assamica (Steph.) Pócs, J. Hattori Bot. Lab. 32: 86, 1969 (Pócs 1969). Bas.: Mastigobryum assamicum Steph., Hedwigia 24 (5): 216, 1885 (Stephani 1885a). ** Bazzania tridens var. cornutistipula (Steph.) Pócs, J. Hattori Bot. Lab. 32: 83, 1969 (Pócs 1969). Bas.: Mastigobryum cornutistipulum Steph., Rev. Bryol. 35 (2): 35, 1908 (Stephani 1908l). *** Bazzania trilobata (L.) Gray, Nat. Arr. Brit. Pl. 1: 704, 1821 (Gray 1821). Bas.: Jungermannia trilobata L., Sp. Pl. 1: 1133, 1753 (Linnaeus 1753). ** Bazzania trilobata var. depauperata (Müll.Frib.) Grolle, Lindbergia 1 (3/4): 197, 1972 [1973] (Grolle 1972b). Bas.: Pleuroschisma trilobatum var. depauperatum Müll.Frib., Lebermoose 2 (18): 266, 1913 (Müller 1913b). *** Bazzania uncigera (Reinw., Blume et Nees) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 415, 1877 (Trevisan 1877). Bas.: Jungermannia uncigera Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 230, 1824 [1825] (Reinwardt et al. 1824a). ** Bazzania uncigera var. brevifolia Herzog, Trans. Brit. Bryol. Soc. 1 (4): 301, 1950 (Herzog 1950a). ** Bazzania uncigera var. gibba (Sande Lac.) Meijer, Blumea 10 (2): 378, 1960 (Meijer 1960). Bas.: Mastigobryum gibbum Sande Lac., Plagiochila Sandei: 8, 1856 (Sande Lacoste 1856c). ** Bazzania undulata Herzog, Trans. Brit. Bryol. Soc. 1 (4): 306, 1950 (Herzog 1950a). ** Bazzania vietnamica Pócs, J. Hattori Bot. Lab. 32: 90, 1969 (Pócs 1969). ** Bazzania vitiana Mitt., Hedwigia 32 (4): 214, 1893 (Stephani 1893c). *** Bazzania vittata (Gottsche) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (Trevisan 1877). Bas.: Mastigobryum vittatum Gottsche, Syn. Hepat. 2: 216, 1845 (Gottsche et al. 1845a). ** Bazzania vittata var. luxurians (De Not.) Schifn., Consp. Hepat. Arch. Ind.: 178, 1898 (Schifner 1898b). Bas.: Mastigobryum vittatum var. luxurians De Not., Epat. Borneo: 29, 1874 (De Notaris 1874). ** Bazzania wallichiana (Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 415, 1877 (Trevisan 1877). Bas.: Mastigobryum wallichianum Lindenb., Syn. Hepat. 2: 229, 1845 (Gottsche et al. 1845a). 116 ** Bazzania watanabei Inoue, J. Jap. Bot. 34 (9): 269, 1959 (Inoue 1959a). * Bazzania wattsiana (Steph.) Meagher, Australas. Bryol. Newslett. 50: 8, 2005 (Meagher 2005b). Bas.: Mastigobryum wattsianum Steph., Bull. Herb. Boissier (sér. 2) 8 (11): 850 (474), 1908 (Stephani 1908a). 117 116 Bazzania wallichiana seems to be a species complex also including Bazzania francana, Bazzania intermedia and Bazzania marginata (Meagher 2010). 117 Bazzania wattsiana is possibly conspeciic with Bazzania tridens (Meagher 2010). World checklist of hornworts and liverworts 163 ** Bazzania wiltensii (Sande Lac. ex Steph.) Schifn., Consp. Hepat. Arch. Ind.: 180, 1898 (Schifner 1898b). Bas.: Mastigobryum wiltensii Sande Lac. ex Steph., Hedwigia 25 (6): 237, 1886 (Stephani 1886c). ** Bazzania wooroonooran Meagher, Nova Hedwigia 100 (3/4): 549, 2015 (Meagher 2015). ** Bazzania wrightii (Gottsche ex Steph.) Steph., Hedwigia 27 (11/12): 279, 1888 (Stephani 1888c). Bas.: Mastigobryum wrightii Gottsche ex Steph., Hedwigia 25 (6): 237, 1886 (Stephani 1886c). ** Bazzania yoshinagana (Steph.) Yasuda, Shokubutsugaku Kakuron: 711, 1911 (Yasuda 1911). Bas.: Mastigobryum yoshinaganum Steph., Bull. Herb. Boissier (sér. 2) 8 (11): 866 (490), 1908 (Stephani 1908a). ** Bazzania zollingeri (Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (Trevisan 1877). Bas.: Mastigobryum zollingeri Lindenb., Bot. Zeitung (Berlin) 6 (25): 462, 1848 (Meissner 1848). *** Bazzania zonulata Meagher, Nova Hedwigia 86 (3/4): 491, 2008 (Meagher 2008). * Mastigopelma Mitt., Fl. vit.: 406, 1871 [1873] (Mitten 1871). 118 ** Mastigopelma fragile (Steph.) N.Kitag., J. Hattori Bot. Lab. 36: 454, 1972 [1973] (Kitagawa 1972). Bas.: Mastigobryum fragile Steph., Sp. Hepat. (Stephani) 6: 463, 1924 (Stephani 1924). ** Mastigopelma pulvinulatum (De Not.) Grolle, J. Hattori Bot. Lab. 33: 39, 1970 (Grolle 1970c). Bas.: Mastigobryum pulvinulatum De Not., Epat. Borneo: 40, 1874 (De Notaris 1874). ** Mastigopelma simplex Mitt., Fl. vit.: 406, 1871 [1873] (Mitten 1871). ** Mastigopelma subissum Grolle, J. Hattori Bot. Lab. 33: 36, 1970 (Grolle 1970c). *** Drucelloideae R.M.Schust. *** Drucella E.A.Hodgs., Trans. Roy. Soc. New Zealand, Bot. 2 (3): 45, 1962 (Hodgson 1962b). *** Drucella integristipula (Steph.) E.A.Hodgs., Trans. Roy. Soc. New Zealand, Bot. 2 (3): 45, 1962 (Hodgson 1962b). Bas.: Lepidozia integristipula Steph., Sp. Hepat. (Stephani) 6: 331, 1922 (Stephani 1922). *** Lembidioideae R.M.Schust. ** Dendrolembidium Herzog, Ark. Bot. (n.ser.) 1 (13): 497, 1951 (Herzog 1951b). *** Dendrolembidium dendroides (Carrington et Pearson) Herzog, Ark. Bot. (n.ser.) 1 (13): 500, 1951 (Herzog 1951b). Bas.: Lembidium dendroides Carrington et Pearson, 118 Mastigopelma may be better placed in Bazzania, but it is retained here until further studied. 164 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Proc. Linn. Soc. New South Wales (ser. 2) 2 (4): 1047, 1888 (Carrington and Pearson 1888a). *** Dendrolembidium tenax (Grev.) Herzog, Ark. Bot. (n.ser.) 1 (13): 499, 1951 (Herzog 1951b). Bas.: Jungermannia tenax Grev., Ann. Lyceum Nat. Hist. New York 1 (2): 277, 1825 (Greville 1825). ** Hygrolembidium R.M.Schust., J. Hattori Bot. Lab. 26: 277, 1963 (Schuster 1963b). *** Hygrolembidium acrocladum (Berggr.) R.M.Schust., J. Hattori Bot. Lab. 26: 277, 1963 (Schuster 1963b). Bas.: Aplozia acroclada Berggr., New Zealand Hepat.: 9, 1898 (Berggren 1898). ** Hygrolembidium andinum (Herzog) R.M.Schust., Nova Hedwigia 10 (1/2): 23, 1965 (Schuster 1965b). Bas.: Lembidium andinum Herzog, Rev. Bryol. Lichénol. 23 (1/2): 48, 1954 (Herzog 1954). *** Hygrolembidium australe (Steph.) Grolle, J. Jap. Bot. 41 (8): 229, 1966 (Grolle 1966d). Bas.: Hygrobiella australis Steph., Bull. Herb. Boissier (sér. 2) 8 (8): 574 (358), 1908 (Stephani 1908e). *** Hygrolembidium boschianum (Sande Lac.) R.M.Schust., J. Hattori Bot. Lab. 26: 277, 1963 (Schuster 1963b). Bas.: Jungermannia boschiana Sande Lac., Ned. Kruidk. Arch. 3: 521, 1855 (Sande Lacoste 1855). ** Hygrolembidium isophyllum R.M.Schust., Nova Hedwigia 15: 467, 1968 (Schuster 1968b). *** Hygrolembidium rigidum R.M.Schust. et J.J.Engel, Phytologia 62 (1): 9, 1987 (Schuster and Engel 1987a). *** Hygrolembidium triquetrum J.J.Engel et R.M.Schust., Phytologia 62 (1): 11, 1987 (Schuster and Engel 1987a). ** Hygrolembidium ventrosum (Mitt.) Grolle, Marion Prince Edw. Is: 233, 1971 (Grolle 1971d). Bas.: Lembidium ventrosum Mitt., J. Linn. Soc., Bot. 15 (82): 69, 1876 (Mitten 1876a). ** Isolembidium R.M.Schust., Nova Hedwigia 15: 466, 1968 (Schuster 1968b). *** Isolembidium anomalum (Rodway) Grolle, J. Bryol. 10 (3): 264, 1979 (Grolle 1979b). Bas.: Lembidium anomalum Rodway, Tasm. Bryoph.: 70, 1917 (Rodway 1917b). ** Isolembidium anomalum var. cucullatum (E.A.Hodgs.) J.J.Engel et R.M.Schust., J. Hattori Bot. Lab. 63: 268, 1987 (Schuster and Engel 1987b). Bas.: Lembidium cucullatum E.A.Hodgs., Rec. Domin. Mus. 4 (11): 110, 1962 (Hodgson 1962a). *** Kurzia G.Martens, Flora 53 (27): 417, 1870 (0). ** subg. Kurzia *** Kurzia capillaris (Sw.) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 173, 1963 [1964] (Grolle 1963b). Bas.: Jungermannia capillaris Sw., Prodr. (Swartz): 144, 1788 (Swartz 1788). World checklist of hornworts and liverworts 165 *** Kurzia capillaris subsp. capillaris var. verrucosa (Steph.) Pócs, Proc. hird Meeting Bryol. C. & E. Europe: 111, 1984 (Pócs 1984b). Bas.: Lepidozia verrucosa Steph., Hedwigia 24 (4): 167, 1885 (Stephani 1885f). *** Kurzia capillaris subsp. paramicola Pócs, Acta Biol. Pl. Agr. 2: 102, 2012 (Pócs 2012a). *** Kurzia capillaris subsp. stephanii (Renauld ex Steph.) Pócs, Proc. hird Meeting Bryol. C. & E. Europe: 111, 1984 (Pócs 1984b). Bas.: Lepidozia stephanii Renauld ex Steph., Bot. Gaz. 15 (11): 287, 1890 (Stephani 1890c). *** Kurzia gonyotricha (Sande Lac.) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 167, 1963 [1964] (Grolle 1963b). Bas.: Lepidozia gonyotricha Sande Lac., Ned. Kruidk. Arch. 3: 521, 1855 (Sande Lacoste 1855). ** Kurzia nemoides (Hook.f. et Taylor) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 173, 1963 [1964] (Grolle 1963b). Bas.: Jungermannia nemoides Hook.f. et Taylor, London J. Bot. 4: 84, 1845 (Hooker and Taylor 1845). ** subg. Micrisophylla (Fulford) J.J.Engel ex R.M.Schust., J. Hattori Bot. Lab. 48: 347, 1980 (Schuster 1980a). Bas.: Micrisophylla Fulford, Brittonia 14 (1): 124, 1962 (Fulford 1962a). ** Kurzia saddlensis (Besch. et C.Massal.) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 174, 1963 [1964] (Grolle 1963b). Bas.: Lepidozia saddlensis Besch. et C.Massal., Bull. Mens. Soc. Linn. Paris 1 (79): 637, 1886 (Bescherelle and Massalongo 1886). ** subg. Microlepidozia (Spruce) R.M.Schust., J. Hattori Bot. Lab. 48: 355, 1980 (Schuster 1980a). Bas.: Lepidozia subg. Microlepidozia Spruce, J. Bot. 14: 165, 1876 (Spruce 1876b). ** Kurzia calcarata (Steph.) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 178, 1963 [1964] (Grolle 1963b). Bas.: Lepidozia calcarata Steph., Sp. Hepat. (Stephani) 3: 592, 1909 (Stephani 1909a). ** Kurzia compacta (Steph.) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 178, 1963 [1964] (Grolle 1963b). Bas.: Lepidozia compacta Steph., Sp. Hepat. (Stephani) 3: 592, 1909 (Stephani 1909a). ** Kurzia helophila R.M.Schust., J. Hattori Bot. Lab. 48: 368, 1980 (Schuster 1980a). ** Kurzia helophila var. laccida R.M.Schust. ex J.J.Engel, Novon 17 (3): 310, 2007 (Engel 2007). *** Kurzia hippurioides (Hook.f. et Taylor) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 178, 1963 [1964] (Grolle 1963b). Bas.: Jungermannia hippurioides Hook.f. et Taylor, London J. Bot. 3: 287 [387], 1844 (Hooker and Taylor 1844a). ** Kurzia hippurioides var. ornata J.J.Engel et G.L.Merr., J. Hattori Bot. Lab. 80: 228, 1996 (Engel and Smith Merrill 1996). ** Kurzia irregularis (Steph.) Grolle, J. Hattori Bot. Lab. 36: 548, 1972 [1973] (Grolle 1972a). Bas.: Lepidozia irregularis Steph., Wiss. Ergebn. Deut. Zentr.-Afr. Exped. (1907-08), Bot. 2: 120, 1911 (Stephani 1911a). ** Kurzia moniliformis J.J.Engel, Cryptog. Bryol. 26 (1): 73, 2005 (Engel 2005). 166 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Kurzia paucilora (Dicks.) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 171, 1963 [1964] (Grolle 1963b). Bas.: Jungermannia paucilora Dicks., Fasc. Pl. Crypt. Brit. 2: 15, 1790 (Dickson 1790). Incertae sedis ** Kurzia abbreviata Mizut., J. Hattori Bot. Lab. 38: 379, 1974 (Mizutani 1974). ** Kurzia abietinella (Herzog) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 170, 1963 [1964] (Grolle 1963b). Bas.: Lepidozia abietinella Herzog, Trans. Brit. Bryol. Soc. 1 (4): 311, 1950 (Herzog 1950a). *** Kurzia bisetula (Steph.) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 170, 1963 [1964] (Grolle 1963b). Bas.: Lepidozia bisetula Steph., Sp. Hepat. (Stephani) 6: 323, 1922 (Stephani 1922). ** Kurzia borneensis Mizut., J. Hattori Bot. Lab. 38: 377, 1974 (Mizutani 1974). ** Kurzia brasiliensis (Steph.) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 174, 1963 [1964] (Grolle 1963b). Bas.: Psiloclada brasiliensis Steph., Sp. Hepat. (Stephani) 3: 550, 1909 (Stephani 1909a). ** Kurzia brevicalycina (Steph.) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 175, 1963 [1964] (Grolle 1963b). Bas.: Lepidozia brevicalycina Steph., Sp. Hepat. (Stephani) 3: 580, 1909 (Stephani 1909a). * Kurzia caduciloba R.M.Schust., Beih. Nova Hedwigia 118: 273, 2000 (Schuster 2000a). * Kurzia cucullifolia (Steph.) R.M.Schust., J. Hattori Bot. Lab. 48: 350, 1980 (Schuster 1980a). Bas.: Lepidozia cucullifolia Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 6): 51, 1900 (Stephani 1900b). 119 *** Kurzia lagellifera (Steph.) Grolle, J. Jap. Bot. 39 (3): 80, 1964 (Grolle 1964c). Bas.: Lepidozia lagellifera Steph., Sp. Hepat. (Stephani) 3: 571, 1909 (Stephani 1909a). *** Kurzia fragilifolia R.M.Schust., J. Hattori Bot. Lab. 48: 364, 1980 (Schuster 1980a). ** Kurzia fragillima (Herzog) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 174, 1963 [1964] (Grolle 1963b). Bas.: Lepidozia fragillima Herzog, Nat. Hist. Juan Fernandez (Botany) 2 (5): 726, 1942 (Herzog 1942a), nom. illeg. ** Kurzia geniculata Mizut., J. Hattori Bot. Lab. 38: 383, 1974 (Mizutani 1974). ** Kurzia hawaica (C.M.Cooke) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 170, 1963 [1964] (Grolle 1963b). Bas.: Lepidozia hawaica C.M.Cooke, Trans. Connecticut Acad. Arts 12 (1): 8, 1904 (Cooke 1904). ** Kurzia hispida (Steph.) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 170, 1963 [1964] (Grolle 1963b). Bas.: Lepidozia hispida Steph., Sp. Hepat. (Stephani) 3: 607, 1909 (Stephani 1909a). 119 Kurzia cucullifolia is of doubtful status (Engel 1978). World checklist of hornworts and liverworts 167 ** Kurzia lateconica (Steph.) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 175, 1963 [1964] (Grolle 1963b). Bas.: Lepidozia lateconica Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 113, 1914 (Stephani and Watts 1914). ** Kurzia lineariloba Mizut., J. Hattori Bot. Lab. 38: 382, 1974 (Mizutani 1974). *** Kurzia longicaulis Piippo, Acta Bot. Fenn. 131: 174, 1985 (Piippo 1985b). ** Kurzia makinoana (Steph.) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 171, 1963 [1964] (Grolle 1963b). Bas.: Lepidozia makinoana Steph., Bull. Herb. Boissier 5 (2): 94, 1897 (Stephani 1897b). ** Kurzia mauiensis (H.A.Mill.) H.A.Mill., J. Hattori Bot. Lab. 30: 274, 1967 (Miller 1967). Bas.: Microlepidozia mauiensis H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 496, 1963 (Miller 1963). ** Kurzia mollis (Steph.) J.J.Engel et R.M.Schust., Bryologist 79 (4): 514, 1976 [1977] (Engel 1976b). Bas.: Lepidozia mollis Steph., Sp. Hepat. (Stephani) 3: 601, 1909 (Stephani 1909a). *** Kurzia nivicola (R.M.Schust.) E.D.Cooper, Phytotaxa 97 (2): 52, 2013 (Cooper et al. 2013). Bas.: Telaranea nivicola R.M.Schust., Nova Hedwigia 15: 460, 1968 (Schuster 1968b). *** Kurzia pallescens Grolle, Rev. Bryol. Lichénol. 32 (3/4): 177, 1963 [1964] (Grolle 1963b). ** Kurzia pallida Piippo, Acta Bot. Fenn. 131: 178, 1985 (Piippo 1985b). *** Kurzia quinquespina J.J.Engel et G.L.Merr., J. Hattori Bot. Lab. 80: 217, 1996 (Engel and Smith Merrill 1996). ** Kurzia reversa (Carrington et Pearson) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 175, 1963 [1964] (Grolle 1963b). Bas.: Lepidozia reversa Carrington et Pearson, J. Bot. 27: 225, 1889 (Carrington and Pearson 1889). ** Kurzia setiformis (De Not.) J.J.Engel et R.M.Schust., Bryologist 79 (4): 514, 1976 [1977] (Engel 1976b). Bas.: Lepidozia setiformis De Not., Mem. Reale Accad. Sci. Torino (ser. 2) 16: 225, 1857 (De Notaris 1857). ** Kurzia sexida (Steph.) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 178, 1963 [1964] (Grolle 1963b). Bas.: Lepidozia sexida Steph., Sp. Hepat. (Stephani) 3: 582, 1909 (Stephani 1909a). ** Kurzia sinensis K.C.Chang, Bull. Bot. Res., Harbin 4 (3): 83, 1984 (Chang and Gao 1984). ** Kurzia sylvatica (A.Evans) Grolle, Herzogia 3: 77, 1973 (Grolle 1973a). Bas.: Lepidozia sylvatica A.Evans, Rhodora 6 (69): 186, 1904 (Evans 1904b). *** Kurzia tasmanica (Steph.) E.D.Cooper, Phytotaxa 97 (2): 52, 2013 (Cooper et al. 2013). Bas.: Lepidozia tasmanica Steph., Sp. Hepat. (Stephani) 3: 580, 1909 (Stephani 1909a). ** Kurzia tayloriana (H.A.Mill.) H.A.Mill., J. Hattori Bot. Lab. 30: 274, 1967 (Miller 1967). Bas.: Microlepidozia tayloriana H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 497, 1963 (Miller 1963). 168 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Kurzia tenerrima (Mitt.) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 171, 1963 [1964] (Grolle 1963b). Bas.: Lepidozia tenerrima Mitt., Sp. Hepat. (Stephani) 3: 607, 1909 (Stephani 1909a). ** Kurzia touwii N.Kitag., Acta Phytotax. Geobot. 29 (1/5): 56, 1978 (Kitagawa 1978). ** Kurzia trichoclados (Müll.Frib.) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 171, 1963 [1964] (Grolle 1963b). Bas.: Lepidozia trichoclados Müll.Frib., Hedwigia 38 (4): 197, 1899 (Müller 1899). *** Kurzia trilobata (R.M.Schust.) R.M.Schust., Beih. Nova Hedwigia 118: 270, 2000 (Schuster 2000a). Bas.: Kurzia quadriseta var. trilobata R.M.Schust., J. Hattori Bot. Lab. 48: 363, 1980 (Schuster 1980a). ** Kurzia verticellata (Carrington) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 178, 1963 [1964] (Grolle 1963b). Bas.: Lepidozia verticellata Carrington, Pap. & Proc. Roy. Soc. Tasmania 1887: 3, 1888 (Carrington and Pearson 1888b). ** Lembidium Mitt., Handb. N. Zeal. l. 2: 754, 1867 (Hooker 1867) nom. conserv. *** Lembidium berggrenii Herzog, Ark. Bot. (n.ser.) 1 (13): 485, 1951 (Herzog 1951b). *** Lembidium longifolium R.M.Schust., Phytologia 45 (5): 420, 1980 (Schuster 1980b). *** Lembidium nutans (Hook.f. et Taylor) Mitt., Handb. N. Zeal. Fl. 2: 754, 1867 (Mitten 1867). Bas.: Jungermannia nutans Hook.f. et Taylor, London J. Bot. 3: 289 [389], 1844 (Hooker and Taylor 1844a). ** Lembidium nutans var. lagelliferum E.A.Hodgs., Rec. Domin. Mus. 4 (11): 110, 1962 (Hodgson 1962a). ** Megalembidium R.M.Schust., J. Hattori Bot. Lab. 26: 258, 1963 (Schuster 1963b). *** Megalembidium insulanum (W.Martin et E.A.Hodgs.) R.M.Schust., J. Hattori Bot. Lab. 26: 258, 1963 (Schuster 1963b). Bas.: Lembidium insulanum W.Martin et E.A.Hodgs., Trans. & Proc. Roy. Soc. New Zealand 78 (4): 497, 1950 (Martin 1950). ** Pseudocephalozia R.M.Schust., Nova Hedwigia 10 (1/2): 21, 1965 (Schuster 1965b). ** sect. Lobulatae R.M.Schust., J. Hattori Bot. Lab. 36: 371, 1972 (Schuster 1972). ** Pseudocephalozia cucullata J.J.Engel et R.M.Schust., J. Hattori Bot. Lab. 38: 694, 1974 (Schuster and Engel 1974). *** Pseudocephalozia lobulata (Herzog) R.M.Schust., J. Hattori Bot. Lab. 36: 371, 1972 [1973] (Schuster 1972). Bas.: Lembidium lobulatum Herzog, Arch. Esc. Fárm. Fac. Ci. Méd. Córdoba 7: 24, 1938 (Herzog and Hosseus 1938). *** Pseudocephalozia quadriloba (Steph.) R.M.Schust., J. Hattori Bot. Lab. 36: 371, 1972 [1973] (Schuster 1972). Bas.: Isotachis quadriloba Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 6): 54, 1900 (Stephani 1900b). World checklist of hornworts and liverworts 169 ** sect. Pseudocephalozia ** Pseudocephalozia lepidozioides R.M.Schust., Nova Hedwigia 10 (1/2): 22, 1965 (Schuster 1965b). ** Pseudocephalozia leptodictyon R.M.Schust., J. Hattori Bot. Lab. 36: 369, 1972 [1973] (Schuster 1972). *** Pseudocephalozia paludicola R.M.Schust., Nova Hedwigia 10 (1/2): 21, 1965 (Schuster 1965b). *** Lepidozioideae Müll.Frib. *** Ceramanus E.D.Cooper, Phytotaxa 97 (2): 53, 2013 (Cooper et al. 2013). *** Ceramanus centipes (Lindenb. et Gottsche) E.D.Cooper, Phytotaxa 97 (2): 54, 2013 (Cooper et al. 2013). Bas.: Lepidozia centipes Lindenb. et Gottsche, Syn. Hepat. 2: 204, 1845 (Gottsche et al. 1845a). *** Ceramanus clatritexta (Steph.) E.D.Cooper, Phytotaxa 97 (2): 54, 2013 (Cooper et al. 2013). Bas.: Lepidozia clatritexta Steph., Sp. Hepat. (Stephani) 3: 583, 1909 (Stephani 1909a). *** Ceramanus elegans (Colenso) E.D.Cooper, Phytotaxa 97 (2): 54, 2013 (Cooper et al. 2013). Bas.: Lepidozia elegans Colenso, Trans. & Proc. New Zealand Inst. 21: 65, 1889 (Colenso 1889). *** Ceramanus grossiseta (Steph.) E.D.Cooper, Phytotaxa 97 (2): 54, 2013 (Cooper et al. 2013). Bas.: Lepidozia grossiseta Steph., Sp. Hepat. (Stephani) 3: 584, 1909 (Stephani 1909a). *** Ceramanus perfragilis (J.J.Engel et G.L.Merr.) E.D.Cooper, Phytotaxa 97 (2): 54, 2013 (Cooper et al. 2013). Bas.: Telaranea perfragilis J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 72, 2004 (Engel and Smith Merrill 2004). *** Ceramanus pruinosa (Herzog) E.D.Cooper, Phytotaxa 97 (2): 54, 2013 (Cooper et al. 2013). Bas.: Lepidozia pruinosa Herzog, Memoranda Soc. Fauna Fl. Fennica 27: 93, 1952 (Herzog 1952c). *** Ceramanus tuberifera (J.J.Engel et R.M.Schust.) E.D.Cooper, Phytotaxa 97 (2): 54, 2013 (Cooper et al. 2013). Bas.: Telaranea tuberifera J.J.Engel et R.M.Schust., Fieldiana, Bot. (n.ser.) 14: 2, 1983 (Engel and Schuster 1983). *** Lepidozia (Dumort.) Dumort., Recueil Observ. Jungerm.: 19, 1835 (Dumortier 1835) nom. conserv. Bas.: Pleuroschisma sect. Lepidozia Dumort., Syll. Jungerm. Europ.: 69, 1831 (Dumortier 1831). 120 ** Lepidozia acantha J.J.Engel, Fieldiana, Bot. (n.ser.) 42: 71, 2001 (Engel and Schuster 2001). 120 Lepidozia seems to be a monophyletic genus, but its infrataxonomy still has to be tested using molecular data. Grolle and Piippo (1984) could not study Lepidozia cordistipula and Lepidozia palmicola since the types were destroyed in B. 170 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Lepidozia aequiloba Steph., Sp. Hepat. (Stephani) 6: 319, 1922 (Stephani 1922). ** Lepidozia africana Steph., Sp. Hepat. (Stephani) 6: 320, 1922 (Stephani 1922). *** Lepidozia alstonii Fulford, Mem. New York Bot. Gard. 11 (2): 211, 1966 (Fulford 1966). ** Lepidozia ambigua De Not., Epat. Borneo: 25, 1874 (De Notaris 1874). ** Lepidozia andicola Beauverd, Sp. Hepat. (Stephani) 6: 572, 1924 (Stephani 1924). Nom. nov. pro Lepidozia appendiculata Steph., Biblioth. Bot. 87 (2): 225, 1916 (Stephani 1916a), nom. illeg. ** Lepidozia appressifolia Steph., Sp. Hepat. (Stephani) 3: 583, 1909 (Stephani 1909a). *** Lepidozia armata Steph., Sp. Hepat. (Stephani) 3: 567, 1909 (Stephani 1909a). ** Lepidozia asymmetrica Steph., Sp. Hepat. (Stephani) 3: 586, 1909 (Stephani 1909a). ** Lepidozia auriculata Mitt., Sp. Hepat. (Stephani) 3: 579, 1909 (Stephani 1909a). ** Lepidozia australis (Lehm. et Lindenb.) Mitt., Fl. vit.: 406, 1871 [1873] (Mitten 1871). Bas.: Jungermannia australis Lehm. et Lindenb., Nov. Stirp. Pug. 6: 28, 1834 (Lehmann 1834). ** Lepidozia bidens J.J.Engel, Fieldiana, Bot. (n.ser.) 42: 79, 2001 (Engel and Schuster 2001). ** Lepidozia biloba Herzog, Ann. Bryol. 4: 83, 1931 (Herzog 1931b). *** Lepidozia bisbiida Steph., Sp. Hepat. (Stephani) 3: 593, 1909 (Stephani 1909a). ** Lepidozia borneensis Steph., Sp. Hepat. (Stephani) 3: 625, 1909 (Stephani 1909a). ** Lepidozia bragginsiana E.D.Cooper et M.A.M.Renner, Phytotaxa 173 (2): 118, 2014 (Cooper and Renner 2014). * Lepidozia brasiliensis Steph., Sp. Hepat. (Stephani) 3: 571, 1909 (Stephani 1909a). 121 ** Lepidozia brevidentata Mitt., Fl. vit.: 406, 1871 [1873] (Mitten 1871). ** Lepidozia brevifolia Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 104, 1860 [1861] (Mitten 1860c). ** Lepidozia brevifolia var. planifolia Schifn., Ann. Bryol. 8: 155, 1935 (Verdoorn 1935). ** Lepidozia brotheri Steph., Sp. Hepat. (Stephani) 3: 623, 1909 (Stephani 1909a). ** Lepidozia bufalona Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 111, 1914 (Stephani and Watts 1914). ** Lepidozia bursifera S.Hatt. et Grolle, J. Hattori Bot. Lab. 30: 115, 1967 (Grolle 1967b). ** Lepidozia caespitosa Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 362, 1885 (Spruce 1885). ** Lepidozia caledonica Steph., Rev. Bryol. 35 (2): 31, 1908 (Stephani 1908l). ** Lepidozia caledonica var. tenuisecta Hürl., Bauhinia 8 (2): 109, 1985 (Hürlimann 1985). * Lepidozia ceramensis Herzog, Hedwigia 66 (6): 340, 1926 (Herzog 1926). ** Lepidozia cherydrion Hürl., Bauhinia 8 (2): 109, 1985 (Hürlimann 1985). ** Lepidozia chiloensis Steph., Sp. Hepat. (Stephani) 6: 322, 1922 (Stephani 1922). 121 Lepidozia brasiliensis is conspeciic with Kurzia brasiliensis in Pócs (1984b), possibly conspeciic with Lepidozia cupressina in Gradstein and Costa (2003), but it has later been accepted by many authors (e.g. Schäfer-Verwimp and Pócs 2009) World checklist of hornworts and liverworts 171 ** Lepidozia chordulifera Taylor, London J. Bot. 5: 371, 1846 (Taylor 1846b). *** Lepidozia cladorhiza (Reinw., Blume et Nees) Nees, Syn. Hepat. 2: 210, 1845 (Gottsche et al. 1845a). Bas.: Jungermannia cladorhiza Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 203, 1824 [1825] (Reinwardt et al. 1824a). *** Lepidozia coilophylla Taylor, London J. Bot. 5: 370, 1846 (Taylor 1846b). ** Lepidozia coilophylla var. apiculiloba (Steph.) Fulford, Mem. New York Bot. Gard. 11 (2): 194, 1966 (Fulford 1966). Bas.: Lepidozia apiculiloba Steph., Sp. Hepat. (Stephani) 6: 321, 1922 (Stephani 1922). ** Lepidozia communis Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 111, 1914 (Stephani and Watts 1914). *** Lepidozia concinna Colenso, Trans. & Proc. New Zealand Inst. 18: 244, 1886 (Colenso 1886b). ** Lepidozia cordata Lindenb., Syn. Hepat. 2: 207, 1845 (Gottsche et al. 1845a). * Lepidozia cordistipula Steph., Sp. Hepat. (Stephani) 6: 345, 1922 (Stephani 1922). ** Lepidozia crassitexta Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 111, 1914 (Stephani and Watts 1914). *** Lepidozia cupressina (Sw.) Lindenb., Syn. Hepat. 2: 207, 1845 (Gottsche et al. 1845a). Bas.: Jungermannia cupressina Sw., Prodr. (Swartz): 144, 1788 (Swartz 1788). ** Lepidozia cupressina subsp. natalensis (Steph.) Pócs, Proc. hird Meeting Bryol. C. & E. Europe: 109, 1984 (Pócs 1984b). Bas.: Lepidozia natalensis Steph., Sp. Hepat. (Stephani) 3: 562, 1909 (Stephani 1909a). * Lepidozia cupressina subsp. pinnata (Hook.) Pócs, Proc. hird Meeting Bryol. C. & E. Europe: 109, 1984 (Pócs 1984b). Bas.: Jungermannia reptans var. pinnata Hook., Brit. Jungermann.: tab. 75, 1815 (Hooker 1815). * Lepidozia cupressina subsp. quinqueida (Steph.) Pócs, Proc. hird Meeting Bryol. C. & E. Europe: 109, 1984 (Pócs 1984b). Bas.: Lepidozia quinqueida Steph., Wiss. Ergebn. Deut. Zentr.-Afr. Exped. (1907-08), Bot. 2: 123, 1911 (Stephani 1911a). ** Lepidozia decaisnei Steph., Sp. Hepat. (Stephani) 3: 588, 1909 (Stephani 1909a). ** Lepidozia dendritica Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 362, 1885 (Spruce 1885). ** Lepidozia densa Herzog, Repert. Spec. Nov. Regni Veg. 21 (1/7): 26, 1925 (Herzog 1925a). *** Lepidozia digitata Herzog, Trans. & Proc. Roy. Soc. New Zealand 68 (1): 45, 1938 (Herzog 1938c). *** Lepidozia eenii S.W.Arnell, Svensk Bot. Tidskr. 57 (2): 190, 1963 (Arnell 1963a). ** Lepidozia elobata R.M.Schust., Fieldiana, Bot. (n.ser.) 42: 74, 2001 (Engel and Schuster 2001). ** Lepidozia erosa Steph., Sp. Hepat. (Stephani) 3: 621, 1909 (Stephani 1909a). ** Lepidozia erronea Herzog, Nat. Hist. Juan Fernandez (Botany) 2 (5): 725, 1942 (Herzog 1942a). Nom. nov. pro Lepidozia fernandeziensis Steph., Sp. Hepat. (Stephani) 6: 326, 1922 (Stephani 1922), nom. illeg. 172 ** * ** *** ** ** ** ** ** * *** *** ** * ** ** ** *** ** *** *** ** ** *** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Lepidozia everettii Steph., Sp. Hepat. (Stephani) 3: 622, 1909 (Stephani 1909a). Lepidozia everettii var. javensis Herzog, Ann. Bryol. 5: 78, 1932 (Herzog 1932b). Lepidozia fauriana Steph., Sp. Hepat. (Stephani) 3: 631, 1909 (Stephani 1909a). Lepidozia ferdinandi-muelleri Steph., Sp. Hepat. (Stephani) 3: 614, 1909 (Stephani 1909a). Lepidozia ilamentosa (Lehm. et Lindenb.) Lehm. et Lindenb., Syn. Hepat. 2: 206, 1845 (Gottsche et al. 1845a). Bas.: Jungermannia ilamentosa Lehm. et Lindenb., Nov. Stirp. Pug. 6: 29, 1834 (Lehmann 1834). Lepidozia istulosa Mitt., Fl. vit.: 406, 1871 [1873] (Mitten 1871). Lepidozia lexuosa Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 103, 1860 [1861] (Mitten 1860c). Lepidozia fuegiensis Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 63, 1911 (Stephani 1911b). Lepidozia fugax J.J.Engel, Fieldiana, Bot. (n.ser.) 42: 63, 2001 (Engel and Schuster 2001). Lepidozia gedena Steph., Sp. Hepat. (Stephani) 6: 327, 1922 (Stephani 1922). Lepidozia glaucescens J.J.Engel, Fieldiana, Bot. (n.ser.) 42: 101, 2001 (Engel and Schuster 2001). Lepidozia glaucophylla (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 207, 1845 (Gottsche et al. 1845a). Bas.: Jungermannia glaucophylla Hook.f. et Taylor, London J. Bot. 3: 580, 1844 (Hooker and Taylor 1844c). Lepidozia grandifolia Steph., Sp. Hepat. (Stephani) 3: 625, 1909 (Stephani 1909a). Lepidozia griseola Herzog, Hedwigia 66 (6): 340, 1926 (Herzog 1926). Lepidozia groenlandica Lehm., Nov. Stirp. Pug. 10: 7, 1857 (Lehmann 1857). Lepidozia gwamii Piippo, Ann. Bot. Fenn. 21 (4): 311, 1984 (Piippo 1984b). Lepidozia hampeana Lindenb., Syn. Hepat. 2: 208, 1845 (Gottsche et al. 1845a). Lepidozia haskarliana (Gottsche, Lindenb. et Nees) Steph., Sp. Hepat. (Stephani) 3: 614, 1909 (Stephani 1909a). Bas.: Lepidozia supradecomposita β haskarliana Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 202, 1845 (Gottsche et al. 1845a). Lepidozia hastatistipula Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 113, 1914 (Stephani and Watts 1914). Lepidozia hirta Steph., Sp. Hepat. (Stephani) 3: 599, 1909 (Stephani 1909a). Lepidozia holorhiza (Reinw., Blume et Nees) Nees, Syn. Hepat. 2: 210, 1845 (Gottsche et al. 1845a). Bas.: Jungermannia holorhiza Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 204, 1824 [1825] (Reinwardt et al. 1824a). Lepidozia holorhiza var. laxa (Nees) Schifn., Consp. Hepat. Arch. Ind.: 186, 1898 (Schifner 1898b). Bas.: Jungermannia holorhiza β laxa Nees, Enum. Pl. Crypt. Javae: 14, 1830 (Nees 1830). Lepidozia inaequalis (Lehm. et Lindenb.) Lehm. et Lindenb., Syn. Hepat. 2: 209, 1845 (Gottsche et al. 1845a). Bas.: Jungermannia inaequalis Lehm. et Lindenb., Nov. Stirp. Pug. 5: 1, 1833 (Lehmann 1833). Lepidozia incurvata Lindenb., Syn. Hepat. 2: 203, 1845 (Gottsche et al. 1845a). World checklist of hornworts and liverworts ** ** *** ** ** *** ** *** ** ** ** ** ** ** *** * *** ** ** * ** ** * 173 Lepidozia infuscata Mitt., Fl. vit.: 406, 1871 [1873] (Mitten 1871). Lepidozia integrifolia Doei, J. Hattori Bot. Lab. 63: 421, 1987 (Doei 1987). Lepidozia jamaicensis Steph., Sp. Hepat. (Stephani) 3: 568, 1909 (Stephani 1909a). Lepidozia kashyapii D.Singh et D.K.Singh, Nova Hedwigia 94 (1/2): 222, 2012 (Singh and Singh 2012). Lepidozia kinabaluensis Mizut., J. Hattori Bot. Lab. 38: 372, 1974 (Mizutani 1974). Lepidozia kirkii Steph., Sp. Hepat. (Stephani) 3: 598, 1909 (Stephani 1909a). Lepidozia lacerifolia Steph., Sp. Hepat. (Stephani) 6: 332, 1922 (Stephani 1922). Lepidozia laevifolia (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 208, 1845 (Gottsche et al. 1845a). Bas.: Jungermannia laevifolia Hook.f. et Taylor, London J. Bot. 3: 285 [385], 1844 (Hooker and Taylor 1844a). Lepidozia laevifolia var. acutiloba J.J.Engel, Fieldiana, Bot. (n.ser.) 42: 61, 2001 (Engel and Schuster 2001). Lepidozia laevifolia var. alpina R.M.Schust. et J.J.Engel, Fieldiana, Bot. (n.ser.) 42: 62, 2001 (Engel and Schuster 2001). Lepidozia lindigiana Steph., Sp. Hepat. (Stephani) 3: 573, 1909 (Stephani 1909a). Lepidozia loheri Steph., Sp. Hepat. (Stephani) 3: 621, 1909 (Stephani 1909a). Lepidozia longifolia Steph., Sp. Hepat. (Stephani) 3: 606, 1909 (Stephani 1909a). Lepidozia loriana Steph., Sp. Hepat. (Stephani) 6: 333, 1922 (Stephani 1922). Lepidozia macrocolea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 363, 1885 (Spruce 1885). Lepidozia massartiana Schifn. ex Steph., Sp. Hepat. (Stephani) 3: 611, 1909 (Stephani 1909a). Based on: Lepidozia massartiana Schifn., Hedwigia 39 (4): 196, 1900 (Schifner 1900b), nom. inval. 122 Lepidozia microphylla (Hook.) Lindenb., Syn. Hepat. 2: 202, 1845 (Gottsche et al. 1845a). Bas.: Jungermannia microphylla Hook., Musci Exot. 1: tab. 80, 1818 (Hooker 1818). Lepidozia microstipula Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 114, 1914 (Stephani and Watts 1914). Lepidozia minima Steph., Sp. Hepat. (Stephani) 6: 335, 1922 (Stephani 1922). Lepidozia minor (Gottsche, Lindenb. et Nees) Solari, Boll. Mus. Civico Storia Nat. Verona 10: 203, 1983 [1985] (Solari and Hässel 1983). Bas.: Lepidozia truncatella β minor Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 209, 1845 (Gottsche et al. 1845a). 123 Lepidozia miqueliana Sande Lac., Ann. Mus. Bot. Lugduno-Batavi 1: 301, 1864 (Sande Lacoste 1864). Lepidozia montana Steph., Sp. Hepat. (Stephani) 3: 587, 1909 (Stephani 1909a). Lepidozia newtonii Steph., Sp. Hepat. (Stephani) 3: 623, 1909 (Stephani 1909a). 122 Lepidozia massartiana has not been studied recently. Stephani (1909a) confused it with Telaranea cuneifolia (Söderström et al. 2010a). 123 Lepidozia minor may be conspeciic with a known taxon. Lepidozia minor sensu Solari and Hässel (1983) is probably another taxon perhaps without name. 174 * *** ** ** *** *** ** *** * * ** *** * ** ** *** ** *** * ** *** * *** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Lepidozia nova Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 115, 1914 (Stephani and Watts 1914). Lepidozia novae-zelandiae Steph., Sp. Hepat. (Stephani) 3: 595, 1909 (Stephani 1909a). Lepidozia novae-zelandiae var. heterostipa R.M.Schust., Fieldiana, Bot. (n.ser.) 42: 70, 2001 (Engel and Schuster 2001). Lepidozia novae-zelandiae var. minima R.M.Schust., Fieldiana, Bot. (n.ser.) 42: 71, 2001 (Engel and Schuster 2001). Lepidozia obtusiloba Steph., Sp. Hepat. (Stephani) 3: 598, 1909 (Stephani 1909a). Lepidozia obtusiloba var. parvula J.J.Engel, Fieldiana, Bot. (n.ser.) 42: 48, 2001 (Engel and Schuster 2001). Lepidozia omeiensis P.C.Chen ex Mizut. et K.C.Chang, J. Hattori Bot. Lab. 60: 421, 1986 (Mizutani and Chang 1986). Lepidozia ornata J.J.Engel, Fieldiana, Bot. (n.ser.) 42: 49, 2001 (Engel and Schuster 2001). Lepidozia pallida Steph., Sp. Hepat. (Stephani) 3: 604, 1909 (Stephani 1909a). 124 Lepidozia palmicola Steph., Sp. Hepat. (Stephani) 6: 346, 1922 (Stephani 1922). Lepidozia paschalis Steph., Sp. Hepat. (Stephani) 6: 336, 1922 (Stephani 1922). Lepidozia patens Lindenb., Syn. Hepat. 2: 202, 1845 (Gottsche et al. 1845a). Lepidozia paucifolia Steph., Sp. Hepat. (Stephani) 3: 610, 1909 (Stephani 1909a). 125 Lepidozia paupercula Steph., Sp. Hepat. (Stephani) 6: 337, 1922 (Stephani 1922). Lepidozia pearsonii Spruce, J. Bot. 19: 34, 1881 (Spruce 1881b). Lepidozia pendulina (Hook.) Lindenb., Syn. Hepat. 2: 208, 1845 (Gottsche et al. 1845a). Bas.: Jungermannia pendulina Hook., Musci Exot. 1: tab. 60, 1818 (Hooker 1818). Lepidozia peruviensis Steph., Sp. Hepat. (Stephani) 3: 575, 1909 (Stephani 1909a). Lepidozia pinnaticruris Spruce ex Steph., Sp. Hepat. (Stephani) 3: 579, 1909 (Stephani 1909a). Lepidozia plumula Herzog, Beih. Bot. Centralbl. 38 (2): 331, 1921 (Herzog 1921). Lepidozia portoricensis Fulford, Mem. New York Bot. Gard. 11 (2): 187, 1966 (Fulford 1966). Lepidozia procera Mitt., Bot. antarct. voy. III (Fl. Tasman. 2): 231, 1860 (Mitten 1860b). Lepidozia pseudocupressina Schifn., Krit. Bemerk. Eur. Lebermoose 14: 9, 1919 (Schifner 1919). Lepidozia pumila J.J.Engel, Fieldiana, Bot. (n.ser.) 42: 76, 2001 (Engel and Schuster 2001). 124 Lepidozia pallida is possibly Lepidozia chordulifera since all specimens cited by Stephani (1911b) and examined by Engel (1978) belong to this species, except the supposed type specimen which does not contain any Lepidozia material. 125 Lepidozia paucifolia is conspeciic with Lepidozia fauriana in del Rosario (1977), but Tan and Engel (1986) kept them apart. World checklist of hornworts and liverworts 175 *** Lepidozia quadridens (Nees) Nees, Syn. Hepat. 2: 209, 1845 (Gottsche et al. 1845a). Bas.: Jungermannia quadridens Nees, Enum. Pl. Crypt. Javae: 18, 1830 (Nees 1830). ** Lepidozia quadriida Lindenb., Syn. Hepat. 2: 203, 1845 (Gottsche et al. 1845a). *** Lepidozia reptans (L.) Dumort., Recueil Observ. Jungerm.: 19, 1835 (Dumortier 1835). Bas.: Jungermannia reptans L., Sp. Pl. 1: 1133, 1753 (Linnaeus 1753). ** Lepidozia richardsii Herzog, Trans. Brit. Bryol. Soc. 1 (4): 313, 1950 (Herzog 1950a). ** Lepidozia rigida Steph., Sp. Hepat. (Stephani) 6: 340, 1922 (Stephani 1922). ** Lepidozia robusta Steph., Mém. Soc. Nat. Sci. Nat. Math. Cherbourg 29: 217, 1894 (Stephani 1894b). * Lepidozia rufescens Steph., Biblioth. Bot. 87 (2): 226, 1916 (Stephani 1916a). ** Lepidozia sandvicensis Lindenb., Syn. Hepat. 2: 201, 1845 (Gottsche et al. 1845a). ** Lepidozia schwabei Herzog, Rev. Bryol. Lichénol. 23 (1/2): 49, 1954 (Herzog 1954). ** Lepidozia sellingiana H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 494, 1963 (Miller 1963). *** Lepidozia septemida Steph., Sp. Hepat. (Stephani) 3: 588, 1909 (Stephani 1909a). ** Lepidozia serpens Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 364, 1885 (Spruce 1885). *** Lepidozia serrulata J.J.Engel, J. Hattori Bot. Lab. 96: 273, 2004 (Engel 2004a). *** Lepidozia setigera Steph., Sp. Hepat. (Stephani) 3: 599, 1909 (Stephani 1909a). ** Lepidozia sikkimensis Steph., Sp. Hepat. (Stephani) 6: 341, 1922 (Stephani 1922). *** Lepidozia spinosissima (Hook.f. et Taylor) Mitt., Bot. antarct. voy. II (Fl. Nov.Zel. 2): 146, 1854 (Mitten 1854). Bas.: Sendtnera spinosissima Hook.f. et Taylor, London J. Bot. 5: 373, 1846 (Taylor 1846b). * Lepidozia squamifolia Steph., Sp. Hepat. (Stephani) 6: 341, 1922 (Stephani 1922). *** Lepidozia squarrosa Steph., Sp. Hepat. (Stephani) 3: 573, 1909 (Stephani 1909a). ** Lepidozia stahlii Steph., Sp. Hepat. (Stephani) 3: 616, 1909 (Stephani 1909a). ** Lepidozia stuhlmannii Steph., Bot. Jahrb. Syst. 20 (3): 308, 1895 (Stephani 1895a). ** Lepidozia stuhlmannii var. carnosa (Steph.) Pócs, Lidia 4 (1): 22, 1997 (Lye and Pócs 1997). Bas.: Lepidozia carnosa Steph., Wiss. Ergebn. Deut. Zentr.-Afr. Exped. (1907-08), Bot. 2: 122, 1911 (Stephani 1911a). ** Lepidozia stuhlmannii subsp. pulvinata (Steph.) Pócs, Trop. Bryol. 9: 127, 1994 (Pócs 1994c). Bas.: Lepidozia pulvinata Steph., Wiss. Ergebn. Deut. Zentr.-Afr. Exped. (1907-08), Bot. 2: 121, 1911 (Stephani 1911a). ** Lepidozia subdichotoma Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 361, 1885 (Spruce 1885). ** Lepidozia subintegra Lindenb., Syn. Hepat. 2: 201, 1845 (Gottsche et al. 1845a). ** Lepidozia subtransversa Steph., Bull. Herb. Boissier 5 (2): 95, 1897 (Stephani 1897b). ** Lepidozia subtrichodes Steph., Sp. Hepat. (Stephani) 3: 615, 1909 (Stephani 1909a). ** Lepidozia succida Mitt., Trans. Linn. Soc. London 23 (1): 57, 1860 (Mitten 1860a). ** Lepidozia supradecomposita Lindenb., Syn. Hepat. 2: 202, 1845 (Gottsche et al. 1845a). * Lepidozia supradecomposita var. falcifolia Herzog, Mitt. Inst. Allg. Bot. Hamburg 7 (3): 191, 1931 (Herzog 1931a). 176 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Lepidozia suyungii C.Gao et X.L.Bai, J. Hattori Bot. Lab. 92: 192, 2002 (Gao and Bai 2002). ** Lepidozia terricola Steph., Sp. Hepat. (Stephani) 3: 585, 1909 (Stephani 1909a). ** Lepidozia triangulifolia Steph., Sp. Hepat. (Stephani) 6: 344, 1922 (Stephani 1922). *** Lepidozia trichodes (Reinw., Blume et Nees) Nees, Syn. Hepat. 2: 203, 1845 (Gottsche et al. 1845a). Bas.: Jungermannia trichodes Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 199, 1824 [1825] (Reinwardt et al. 1824a). * Lepidozia tricuspidata Steph., Sp. Hepat. (Stephani) 6: 344, 1922 (Stephani 1922). ** Lepidozia tunguraguae Steph., Sp. Hepat. (Stephani) 6: 345, 1922 (Stephani 1922). ** Lepidozia ubangiensis Steph., Sp. Hepat. (Stephani) 3: 561, 1909 (Stephani 1909a). *** Lepidozia udarii S.C.Srivast., D.Kumar et D.Sharma, Cryptog. Bryol. Lichénol. 9 (3): 237, 1988 (Srivastava et al. 1988). *** Lepidozia ulothrix (Schwägr.) Lindenb., Syn. Hepat. 2: 210, 1845 (Gottsche et al. 1845a). Bas.: Jungermannia ulothrix Schwägr., Hist. Musc. Hepat. Prodr.: 21, 1814 (Schwägrichen 1814). ** Lepidozia vitrea Steph., Bull. Herb. Boissier 5 (2): 96, 1897 (Stephani 1897b). ** Lepidozia wattsiana Steph., Sp. Hepat. (Stephani) 3: 586, 1909 (Stephani 1909a). ** Lepidozia weymouthiana Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 117, 1914 (Stephani and Watts 1914). * * Excluded from the genus Lepidozia hexiloba Pearson, Ann. Cryptog. Exot. 4 (2): 67, 1931 (Pearson 1931a). 126 Lepidozia parvistipa Taylor, London J. Bot. 5: 370, 1846 (Taylor 1846b). 127 *** Neolepidozia Fulford et J.Taylor, Brittonia 11 (2): 81, 1959 (Fulford and Taylor 1959a). *** Neolepidozia aubertii (Jovet-Ast) E.D.Cooper, Phytotaxa 97 (2): 55, 2013 (Cooper et al. 2013). Bas.: Lepidozia aubertii Jovet-Ast, Candollea 11: 35, 1947 (Jovet-Ast 1947a). *** Neolepidozia autoica (J.J.Engel et G.L.Merr.) E.D.Cooper, Phytotaxa 97 (2): 55, 2013 (Cooper et al. 2013). Bas.: Telaranea autoica J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 124, 2004 (Engel and Smith Merrill 2004). *** Neolepidozia capilligera (Schwägr.) Fulford et J.Taylor, Brittonia 11 (2): 84, 1959 (Fulford and Taylor 1959a). Bas.: Jungermannia capilligera Schwägr., Hist. Musc. Hepat. Prodr.: 21, 1814 (Schwägrichen 1814). *** Neolepidozia consobrina (J.J.Engel et G.L.Merr.) E.D.Cooper, Phytotaxa 97 (2): 55, 2013 (Cooper et al. 2013). Bas.: Telaranea consobrina J.J.Engel et G.L.Merr., Novon 9 (3): 339, 1999 (Engel and Smith Merrill 1999a). 126 Lepidozia hexiloba is possibly an Arachniopsis or Telaranea species (Wigginton and Grolle 1996). 127 Lepidozia parvistipa is possibly conspeciic with Ceramanus clatritexta. World checklist of hornworts and liverworts 177 *** Neolepidozia cuneifolia (Steph.) Fulford et J.Taylor, Brittonia 11 (2): 85, 1959 (Fulford and Taylor 1959a). Bas.: Lepidozia cuneifolia Steph., Sp. Hepat. (Stephani) 3: 618, 1909 (Stephani 1909a). *** Neolepidozia disparata (J.J.Engel et G.L.Merr.) E.D.Cooper, Phytotaxa 97 (2): 55, 2013 (Cooper et al. 2013). Bas.: Telaranea disparata J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 147, 2004 (Engel and Smith Merrill 2004). *** Neolepidozia disticha (Steph.) Fulford et J.Taylor, Brittonia 11 (2): 85, 1959 (Fulford and Taylor 1959a). Bas.: Lepidozia disticha Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 62, 1911 (Stephani 1911b). *** Neolepidozia gibbsiana (Steph.) E.D.Cooper, Phytotaxa 97 (2): 55, 2013 (Cooper et al. 2013). Bas.: Lepidozia gibbsiana Steph., Sp. Hepat. (Stephani) 6: 328, 1922 (Stephani 1922). *** Neolepidozia heterotexta (Steph.) E.D.Cooper, Phytotaxa 97 (2): 55, 2013 (Cooper et al. 2013). Bas.: Lepidozia heterotexta Steph., Sp. Hepat. (Stephani) 6: 329, 1922 (Stephani 1922). *** Neolepidozia hodgsoniae (J.J.Engel et G.L.Merr.) E.D.Cooper, Phytotaxa 97 (2): 55, 2013 (Cooper et al. 2013). Bas.: Telaranea hodgsoniae J.J.Engel et G.L.Merr., Phytologia 79 (3): 251, 1995 [1996] (Engel and Smith Merrill 1995). *** Neolepidozia longitudinalis (Herzog) E.D.Cooper, Phytotaxa 97 (2): 56, 2013 (Cooper et al. 2013). Bas.: Lepidozia longitudinalis Herzog, Trans. Brit. Bryol. Soc. 1 (4): 312, 1950 (Herzog 1950a). *** Neolepidozia mamillosa (Schifn.) E.D.Cooper, Phytotaxa 97 (2): 56, 2013 (Cooper et al. 2013). Bas.: Lepidozia mamillosa Schifn., Nova Acta Acad. Caes. Leop.Carol. German. Nat. Cur. 60 (2): 254, 1893 (Schifner 1893a). *** Neolepidozia meridiana (E.A.Hodgs.) E.D.Cooper, Phytotaxa 97 (2): 56, 2013 (Cooper et al. 2013). Bas.: Lepidozia meridiana E.A.Hodgs., Trans. Roy. Soc. New Zealand 83 (4): 611, 1956 (Hodgson 1956). *** Neolepidozia oligophylla (Lehm. et Lindenb.) Fulford et J.Taylor, Brittonia 11 (2): 84, 1959 (Fulford and Taylor 1959a). Bas.: Jungermannia oligophylla Lehm. et Lindenb., Nov. Stirp. Pug. 6: 26, 1834 (Lehmann 1834). *** Neolepidozia ophiria (Gottsche) E.D.Cooper, Phytotaxa 97 (2): 56, 2013 (Cooper et al. 2013). Bas.: Lepidozia ophiria Gottsche, Sp. Hepat. (Stephani) 3: 611, 1909 (Stephani 1909a). *** Neolepidozia palmata (J.J.Engel et G.L.Merr.) E.D.Cooper, Phytotaxa 97 (2): 56, 2013 (Cooper et al. 2013). Bas.: Telaranea palmata J.J.Engel et G.L.Merr., Novon 9 (3): 344, 1999 (Engel and Smith Merrill 1999a). *** Neolepidozia paludicola (E.A.Hodgs.) E.D.Cooper, Phytotaxa 97 (2): 56, 2013 (Cooper et al. 2013). Bas.: Lepidozia meridiana var. paludicola E.A.Hodgs., Trans. Roy. Soc. New Zealand 83 (4): 611, 1956 (Hodgson 1956). *** Neolepidozia papulosa (Steph.) Fulford et J.Taylor, Brittonia 11 (2): 85, 1959 (Fulford and Taylor 1959a). Bas.: Lepidozia papulosa Steph., Sp. Hepat. (Stephani) 3: 609, 1909 (Stephani 1909a). 178 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Neolepidozia parvifolia (Steph.) Fulford et J.Taylor, Brittonia 11 (2): 85, 1959 (Fulford and Taylor 1959a). Bas.: Lepidozia parvifolia Steph., Sp. Hepat. (Stephani) 6: 337, 1922 (Stephani 1922). *** Neolepidozia patentissima (Hook.f. et Taylor) E.D.Cooper, Phytotaxa 97 (2): 56, 2013 (Cooper et al. 2013). Bas.: Jungermannia patentissima Hook.f. et Taylor, London J. Bot. 3: 286 [386], 1844 (Hooker and Taylor 1844a). *** Neolepidozia patentissima var. ampliata (J.J.Engel et G.L.Merr.) E.D.Cooper, Phytotaxa 97 (2): 56, 2013 (Cooper et al. 2013). Bas.: Telaranea patentissima var. ampliata J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 48, 2004 (Engel and Smith Merrill 2004). *** Neolepidozia patentissima var. zebrina (J.J.Engel et G.L.Merr.) E.D.Cooper, Phytotaxa 97 (2): 57, 2013 (Cooper et al. 2013). Bas.: Telaranea patentissima var. zebrina J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 50, 2004 (Engel and Smith Merrill 2004). *** Neolepidozia pennata (J.J.Engel et G.L.Merr.) E.D.Cooper, Phytotaxa 97 (2): 57, 2013 (Cooper et al. 2013). Bas.: Telaranea pennata J.J.Engel et G.L.Merr., Phytologia 79 (3): 252, 1995 [1996] (Engel and Smith Merrill 1995). *** Neolepidozia planifolia (Steph.) E.D.Cooper, Phytotaxa 97 (2): 57, 2013 (Cooper et al. 2013). Bas.: Lepidozia planifolia Steph., Sp. Hepat. (Stephani) 3: 629, 1909 (Stephani 1909a). *** Neolepidozia praenitens (Lehm. et Lindenb.) E.D.Cooper, Phytotaxa 97 (2): 57, 2013 (Cooper et al. 2013). Bas.: Jungermannia praenitens Lehm. et Lindenb., Nov. Stirp. Pug. 6: 27, 1834 (Lehmann 1834). *** Neolepidozia praenitens var. dentifolia (J.J.Engel et G.L.Merr.) E.D.Cooper, Phytotaxa 97 (2): 57, 2013 (Cooper et al. 2013). Bas.: Telaranea praenitens var. dentifolia J.J.Engel et G.L.Merr., Phytologia 79 (3): 253, 1995 [1996] (Engel and Smith Merrill 1995). *** Neolepidozia quadristipula (Steph.) E.D.Cooper, Phytotaxa 97 (2): 57, 2013 (Cooper et al. 2013). Bas.: Lepidozia quadristipula Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 115, 1914 (Stephani and Watts 1914). *** Neolepidozia rectangularis (R.M.Schust.) E.D.Cooper, Phytotaxa 97 (2): 57, 2013 (Cooper et al. 2013). Bas.: Telaranea rectangularis R.M.Schust., Phytologia 39 (4): 241, 1978 (Schuster 1978a). *** Neolepidozia seriatitexta (Steph.) Fulford, Mem. New York Bot. Gard. 11 (2): 215, 1966 (Fulford 1966). Bas.: Lepidozia seriatitexta Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 6): 53, 1900 (Stephani 1900b). *** Neolepidozia tetrapila (Hook.f. et Taylor) E.D.Cooper, Phytotaxa 97 (2): 57, 2013 (Cooper et al. 2013). Bas.: Lepidozia tetrapila Hook.f. et Taylor, London J. Bot. 5: 370, 1846 (Taylor 1846b). *** Neolepidozia tetrapila var. cancellata (Colenso) E.D.Cooper, Phytotaxa 97 (2): 57, 2013 (Cooper et al. 2013). Bas.: Lepidozia cancellata Colenso, Trans. & Proc. New Zealand Inst. 18: 244, 1886 (Colenso 1886b). World checklist of hornworts and liverworts 179 *** Neolepidozia tetrapila var. roseana (Steph.) E.D.Cooper, Phytotaxa 97 (2): 58, 2013 (Cooper et al. 2013). Bas.: Lepidozia roseana Steph., Sp. Hepat. (Stephani) 3: 590, 1909 (Stephani 1909a). *** Neolepidozia tridactylis (Lehm. et Lindenb.) E.D.Cooper, Phytotaxa 97 (2): 58, 2013 (Cooper et al. 2013). Bas.: Jungermannia tridactylis Lehm. et Lindenb., Nov. Stirp. Pug. 4: 41, 1832 (Lehmann 1832). *** Neolepidozia triida (Steph.) E.D.Cooper, Phytotaxa 97 (2): 58, 2013 (Cooper et al. 2013). Bas.: Lepidozia triida Steph., Wiss. Ergebn. Deut. Zentr.-Afr. Exped. (1907-08), Bot. 2: 120, 1911 (Stephani 1911a). *** Neolepidozia verruculosa (J.J.Engel et G.L.Merr.) E.D.Cooper, Phytotaxa 97 (2): 58, 2013 (Cooper et al. 2013). Bas.: Telaranea verruculosa J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 197, 2004 (Engel and Smith Merrill 2004). *** Neolepidozia wallichiana (Gottsche) Fulford et J.Taylor, Brittonia 11 (2): 84, 1959 (Fulford and Taylor 1959a). Bas.: Lepidozia wallichiana Gottsche, Syn. Hepat. 2: 204, 1845 (Gottsche et al. 1845a). *** Tricholepidozia (R.M.Schust.) E.D.Cooper, Phytotaxa 97 (2): 58, 2013 (Cooper et al. 2013). Bas.: Telaranea subg. Tricholepidozia R.M.Schust., J. Hattori Bot. Lab. 26: 256, 1963 (Schuster 1963b). *** Tricholepidozia chaetocarpa (Pearson) E.D.Cooper, Phytotaxa 97 (2): 58, 2013 (Cooper et al. 2013). Bas.: Lepidozia chaetocarpa Pearson, J. Linn. Soc., Bot. 46 (305): 27, 1922 (Pearson 1922b). *** Tricholepidozia fernandeziensis (Steph.) E.D.Cooper, Phytotaxa 97 (2): 58, 2013 (Cooper et al. 2013). Bas.: Lepidozia fernandeziensis Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 63, 1911 (Stephani 1911b). *** Tricholepidozia ferruginea (J.J.Engel et G.L.Merr.) E.D.Cooper, Phytotaxa 97 (2): 58, 2013 (Cooper et al. 2013). Bas.: Telaranea ferruginea J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 159, 2004 (Engel and Smith Merrill 2004). *** Tricholepidozia issifolia (Steph.) E.D.Cooper, Phytotaxa 97 (2): 58, 2013 (Cooper et al. 2013). Bas.: Lepidozia issifolia Steph., Sp. Hepat. (Stephani) 3: 610, 1909 (Stephani 1909a). *** Tricholepidozia jowettiana (H.A.Mill.) E.D.Cooper, Phytotaxa 97 (2): 59, 2013 (Cooper et al. 2013). Bas.: Telaranea jowettiana H.A.Mill., J. Bryol. 14 (2): 235, 1986 (Miller 1986). *** Tricholepidozia kaindina (Grolle) E.D.Cooper, Phytotaxa 97 (2): 59, 2013 (Cooper et al. 2013). Bas.: Telaranea kaindina Grolle, J. Hattori Bot. Lab. 31: 9, 1968 (Grolle 1968a). *** Tricholepidozia kogiana (Steph.) E.D.Cooper, Phytotaxa 97 (2): 59, 2013 (Cooper et al. 2013). Bas.: Lepidozia kogiana Steph., Sp. Hepat. (Stephani) 6: 332, 1922 (Stephani 1922). *** Tricholepidozia lawesii (Steph.) E.D.Cooper, Phytotaxa 97 (2): 59, 2013 (Cooper et al. 2013). Bas.: Lepidozia lawesii Steph., Hedwigia 28 (4): 264, 1889 (Stephani 1889c). 180 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Tricholepidozia leratii (Steph.) E.D.Cooper, Phytotaxa 167 (2): 218, 2014 (Cooper et al. 2014). Bas.: Lepidozia leratii Steph., Sp. Hepat. (Stephani) 6: 333, 1922 (Stephani 1922). *** Tricholepidozia lindenbergii (Gottsche) E.D.Cooper, Phytotaxa 97 (2): 59, 2013 (Cooper et al. 2013). Bas.: Lepidozia lindenbergii Gottsche, Syn. Hepat. 2: 213, 1845 (Gottsche et al. 1845a). *** Tricholepidozia lindenbergii var. complanata (J.J.Engel et G.L.Merr.) E.D.Cooper, Phytotaxa 97 (2): 59, 2013 (Cooper et al. 2013). Bas.: Telaranea lindenbergii var. complanata J.J.Engel et G.L.Merr., Phytologia 79 (3): 252, 1995 [1996] (Engel and Smith Merrill 1995). *** Tricholepidozia lindenbergii var. mellea (J.J.Engel et G.L.Merr.) E.D.Cooper, Phytotaxa 97 (2): 59, 2013 (Cooper et al. 2013). Bas.: Telaranea lindenbergii var. mellea J.J.Engel et G.L.Merr., Phytologia 79 (3): 252, 1995 [1996] (Engel and Smith Merrill 1995). *** Tricholepidozia lindenbergii var. papillata (J.J.Engel et G.L.Merr.) E.D.Cooper, Phytotaxa 97 (2): 59, 2013 (Cooper et al. 2013). Bas.: Telaranea lindenbergii var. papillata J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 83, 2004 (Engel and Smith Merrill 2004). *** Tricholepidozia marginata (J.J.Engel et G.L.Merr.) E.D.Cooper, Phytotaxa 97 (2): 59, 2013 (Cooper et al. 2013). Bas.: Telaranea marginata J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 166, 2004 (Engel and Smith Merrill 2004). *** Tricholepidozia martinii (E.A.Hodgs.) E.D.Cooper, Phytotaxa 97 (2): 59, 2013 (Cooper et al. 2013). Bas.: Lepidozia martinii E.A.Hodgs., Trans. Roy. Soc. New Zealand 83 (4): 602, 1956 (Hodgson 1956). *** Tricholepidozia melanesica (H.A.Mill.) E.D.Cooper, Phytotaxa 97 (2): 60, 2013 (Cooper et al. 2013). Bas.: Telaranea melanesica H.A.Mill., J. Bryol. 14 (2): 237, 1986 (Miller 1986). *** Tricholepidozia neesii (Lindenb.) E.D.Cooper, Phytotaxa 97 (2): 60, 2013 (Cooper et al. 2013). Bas.: Lepidozia neesii Lindenb., Syn. Hepat. 2: 212, 1845 (Gottsche et al. 1845a). *** Tricholepidozia octoloba (Del Ros.) E.D.Cooper, Phytotaxa 97 (2): 60, 2013 (Cooper et al. 2013). Bas.: Telaranea octoloba Del Ros., Philipp. J. Sci. 100 (3/4): 239, 1971 (Del Rosario 1971). *** Tricholepidozia plumulosa (Lehm. et Lindenb.) E.D.Cooper, Phytotaxa 97 (2): 60, 2013 (Cooper et al. 2013). Bas.: Jungermannia plumulosa Lehm. et Lindenb., Nov. Stirp. Pug. 6: 30, 1834 (Lehmann 1834). *** Tricholepidozia pulcherrima (Steph.) E.D.Cooper, Phytotaxa 97 (2): 60, 2013 (Cooper et al. 2013). Bas.: Lepidozia pulcherrima Steph., Sp. Hepat. (Stephani) 3: 600, 1909 (Stephani 1909a). *** Tricholepidozia pulcherrima var. mooreana (Steph.) E.D.Cooper, Phytotaxa 97 (2): 60, 2013 (Cooper et al. 2013). Bas.: Lepidozia mooreana Steph., Sp. Hepat. (Stephani) 3: 585, 1909 (Stephani 1909a). World checklist of hornworts and liverworts 181 *** Tricholepidozia quadriseta (Steph.) E.D.Cooper, Phytotaxa 97 (2): 60, 2013 (Cooper et al. 2013). Bas.: Lepidozia quadriseta Steph., Sp. Hepat. (Stephani) 3: 582, 1909 (Stephani 1909a). *** Tricholepidozia remotifolia (E.A.Hodgs.) E.D.Cooper, Phytotaxa 97 (2): 60, 2013 (Cooper et al. 2013). Bas.: Telaranea remotifolia E.A.Hodgs., Rec. Domin. Mus. 4 (11): 107, 1962 (Hodgson 1962a). *** Tricholepidozia semperiana (Steph.) E.D.Cooper, Phytotaxa 97 (2): 60, 2013 (Cooper et al. 2013). Bas.: Lepidozia semperiana Steph., Sp. Hepat. (Stephani) 3: 612, 1909 (Stephani 1909a). *** Tricholepidozia tetradactyla (Hook.f. et Taylor) E.D.Cooper, Phytotaxa 97 (2): 61, 2013 (Cooper et al. 2013). Bas.: Jungermannia tetradactyla Hook.f. et Taylor, London J. Bot. 3: 286 [386], 1844 (Hooker and Taylor 1844a). *** Tricholepidozia trichocoleoides (Herzog) E.D.Cooper, Phytotaxa 97 (2): 61, 2013 (Cooper et al. 2013). Bas.: Lepidozia trichocoleoides Herzog, Trans. Brit. Bryol. Soc. 1 (4): 314, 1950 (Herzog 1950a). ** Micropterygioideae Grolle ** Micropterygium Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 233, 1845 (Gottsche et al. 1845a). ** Micropterygium angustistipulum Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 385, 1885 (Spruce 1885). *** Micropterygium bialatum Fulford, Mem. New York Bot. Gard. 11 (2): 268, 1966 (Fulford 1966). *** Micropterygium bolivarense Fulford, Mem. New York Bot. Gard. 11 (2): 263, 1966 (Fulford 1966). *** Micropterygium campanense Spruce, Hedwigia 73 (3/4): 157, 1933 (Reimers 1933). *** Micropterygium carinatum (Grev.) Reimers, Hedwigia 76 (3): 166, 1936 (Reimers 1936). Bas.: Jungermannia carinata Grev., Ann. Lyceum Nat. Hist. New York 1 (2): 276, 1825 (Greville 1825). *** Micropterygium conchifolium Reimers, Hedwigia 73 (3/4): 155, 1933 (Reimers 1933). *** Micropterygium duidae Reimers, Hedwigia 73 (3/4): 147, 1933 (Reimers 1933). ** Micropterygium exalatum Steph., Sp. Hepat. (Stephani) 3: 547, 1909 (Stephani 1909a). ** Micropterygium grandistipulum Steph., Trans. Linn. Soc. London, Bot. 6 (1): 98, 1901 (Stephani 1901e). ** Micropterygium laeve H.Rob., Bol. Soc. Venez. Ci. Nat. 32 (132/133): 252, 1976 (Robinson 1976b). ** Micropterygium lechleri Reimers, Hedwigia 73 (3/4): 184, 1933 (Reimers 1933). *** Micropterygium leiophyllum Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 386, 1885 (Spruce 1885). 182 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Micropterygium longicellulatum Uribe et E.L.Linares, Phytotaxa 213 (3): 297, 2015 (Uribe and Linares 2015). ** Micropterygium parvistipulum Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 383, 1885 (Spruce 1885). *** Micropterygium pterygophyllum (Nees) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 413, 1877 (Trevisan 1877). Bas.: Jungermannia pterygophylla Nees, Fl. Bras. (Martius) 1 (1): 377, 1833 (Nees 1833a). * Micropterygium pterygophyllum var. lancifolium Reimers, Hedwigia 73 (3/4): 176, 1933 (Reimers 1933). ** Micropterygium reimersianum Herzog, Hedwigia 81 (5/6): 226, 1943 (Herzog 1943b). *** Micropterygium steyermarkii Fulford, Mem. New York Bot. Gard. 11 (2): 270, 1966 (Fulford 1966). *** Micropterygium tatei Reimers, Hedwigia 73 (3/4): 150, 1933 (Reimers 1933). *** Micropterygium tenax (Steph.) Grolle, J. Bryol. 10 (3): 265, 1979 (Grolle 1979b). Bas.: Harpalejeunea tenax Steph., Trans. Linn. Soc. London, Bot. 6 (1): 100, 1901 (Stephani 1901e). *** Micropterygium trachyphyllum Reimers, Hedwigia 73 (3/4): 186, 1933 (Reimers 1933). 128 * Micropterygium trachyphyllum var. brasiliense Reimers, Hedwigia 73 (3/4): 195, 1933 (Reimers 1933). * Micropterygium trachyphyllum var. cubense Reimers, Hedwigia 73 (3/4): 188, 1933 (Reimers 1933). * Micropterygium trachyphyllum var. guadeloupense Reimers, Hedwigia 73 (3/4): 190, 1933 (Reimers 1933). * Micropterygium trachyphyllum var. jamaicense Reimers, Hedwigia 73 (3/4): 190, 1933 (Reimers 1933). *** Micropterygium tumidulum Fulford, Mem. New York Bot. Gard. 11 (2): 272, 1966 (Fulford 1966). ** Mytilopsis Spruce, Cephalozia: 90, 1882 (Spruce 1882). *** Mytilopsis albifrons Spruce, Cephalozia: 91, 1882 (Spruce 1882). ** Protocephalozioideae R.M.Schust. ** Protocephalozia (Spruce) K.I.Goebel, Flora 77 (2): 83, 1893 (Goebel 1893b). Bas.: Cephalozia subg. Protocephalozia Spruce, Cephalozia: 24, 1882 (Spruce 1882). *** Protocephalozia ephemeroides (Spruce) K.I.Goebel, Flora 77 (2): 83, 1893 (Goebel 1893b). Bas.: Cephalozia ephemeroides Spruce, Cephalozia: 24, 1882 (Spruce 1882). 128 Micropterygium trachyphyllum was split into several varieties by Reimers (1933), but they do not seem to have been accepted by any later author and Söderström et al. (2011a) were not sure about their status. World checklist of hornworts and liverworts *** 183 Zoopsidoideae R.M.Schust. ** Amazoopsis J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 242, 2004 (Engel and Smith Merrill 2004). *** Amazoopsis diplopoda (Pócs) J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 245, 2004 (Engel and Smith Merrill 2004). Bas.: Arachniopsis diplopoda Pócs, Proc. hird Meeting Bryol. C. & E. Europe: 114, 1984 (Pócs 1984b). *** Amazoopsis dissotricha (Spruce) J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 247, 2004 (Engel and Smith Merrill 2004). Bas.: Arachniopsis dissotricha Spruce, Cephalozia: 86, 1882 (Spruce 1882). *** Amazoopsis gracilis J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 246, 2004 (Engel and Smith Merrill 2004). Hyalolepidozia S.W.Arnell ex Grolle, Rev. Bryol. Lichénol. 32 (3/4): 179, 1963 [1964] (Grolle 1963b). Based on: Hyalolepidozia S.W.Arnell, Bot. Not. 115: 203, 1962 (Arnell 1962a). ** Hyalolepidozia bicuspidata (C.Massal.) S.W.Arnell ex Grolle, Rev. Bryol. Lichénol. 32 (3/4): 179, 1963 [1964] (Grolle 1963b). Bas.: Lepidozia bicuspidata C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 239, 1885 (Massalongo 1885). * Monodactylopsis (R.M.Schust.) R.M.Schust., Nova Hedwigia 56 (1/2): 45, 1993 (Schuster 1993a). Bas.: Arachniopsis subg. Monodactylopsis R.M.Schust., Nova Hedwigia 10 (1/2): 24, 1965 (Schuster 1965b). ** Monodactylopsis monodactyla (Spruce) R.M.Schust., Nova Hedwigia 69 (3/4): 523, 1999 (Schuster 1999a). Bas.: Cephalozia monodactyla Spruce, Cephalozia: 28, 1882 (Spruce 1882). * ** Neogrollea E.A.Hodgs., Trans. Roy. Soc. New Zealand, Bot. 3 (4): 70, 1965 (Hodgson 1965). *** Neogrollea notabilis E.A.Hodgs., Trans. Roy. Soc. New Zealand, Bot. 3 (4): 70, 1965 (Hodgson 1965). * * Odontoseries Fulford, Mem. New York Bot. Gard. 11 (3): 364, 1968 (Fulford 1968). Odontoseries chimantana Fulford, Mem. New York Bot. Gard. 11 (3): 366, 1968 (Fulford 1968). *** Paracromastigum Fulford et J.Taylor, Brittonia 13 (4): 336, 1961 (Fulford and Taylor 1961). *** Paracromastigum denticulatum (Steph.) E.D.Cooper, Phytotaxa 97 (2): 61, 2013 (Cooper et al. 2013). Bas.: Lembidium denticulatum Steph., Sp. Hepat. (Stephani) 6: 444, 1924 (Stephani 1924). 184 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Paracromastigum drucei (R.M.Schust.) R.M.Schust., J. Hattori Bot. Lab. 38: 700, 1974 (Schuster and Engel 1974). Bas.: Pseudocephalozia drucei R.M.Schust., Nova Hedwigia 10 (1/2): 22, 1965 (Schuster 1965b). *** Paracromastigum dusenii (Steph.) R.M.Schust., J. Hattori Bot. Lab. 38: 700, 1974 (Schuster and Engel 1974). Bas.: Alobiella dusenii Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 6): 48, 1900 (Stephani 1900b). ** Paracromastigum iordlandiae R.M.Schust. et J.J.Engel, Brittonia 48 (2): 167, 1996 (Schuster and Engel 1996). *** Paracromastigum furcifolium (Steph.) R.M.Schust., J. Hattori Bot. Lab. 26: 276, 1963 (Schuster 1963b). Bas.: Cephalozia furcifolia Steph., Bull. Herb. Boissier (sér. 2) 8 (7): 485 (315), 1908 (Stephani 1908f). ** Paracromastigum granatense (Gottsche) R.M.Schust., J. Hattori Bot. Lab. 48: 341, 1980 (Schuster 1980a). Bas.: Lepidozia granatensis Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 139, 1864 (Gottsche 1864). ** Paracromastigum longiscyphum (Taylor) R.M.Schust. et J.J.Engel, Brittonia 48 (2): 167, 1996 (Schuster and Engel 1996). Bas.: Jungermannia longiscypha Taylor, London J. Bot. 5: 280, 1846 (Taylor 1846a). ** Paracromastigum macrostipum (Steph.) R.M.Schust., J. Hattori Bot. Lab. 26: 276, 1963 (Schuster 1963b). Bas.: Cephalozia macrostipa Steph., Hedwigia 32 (5): 315, 1893 (Stephani 1893d). ** Paracromastigum micromera (Spruce) R.M.Schust., J. Hattori Bot. Lab. 26: 276, 1963 (Schuster 1963b). Bas.: Cephalozia micromera Spruce, Cephalozia: 32, 1882 (Spruce 1882). ** Paracromastigum microphyllum (R.M.Schust. ex J.J.Engel) E.D.Cooper, Phytotaxa 97 (2): 61, 2013 (Cooper et al. 2013). Bas.: Hyalolepidozia microphylla R.M.Schust. ex J.J.Engel, Novon 17 (3): 310, 2007 (Engel 2007). *** Paracromastigum pachyrhizum (Nees) Fulford, Mem. New York Bot. Gard. 11 (3): 390, 1968 (Fulford 1968). Bas.: Jungermannia pachyrhiza Nees, Fl. Bras. (Martius) 1 (1): 339, 1833 (Nees 1833a). ** Paracromastigum ryszardii Váňa, Bedn.-Ochyra et Cykowska, Nova Hedwigia 89 (1/2): 122, 2009 (Váňa et al. 2009). ** Paracromastigum stipulatum (Herzog) Fulford, Mem. New York Bot. Gard. 11 (3): 390, 1968 (Fulford 1968). Bas.: Cephalozia stipulata Herzog, Memoranda Soc. Fauna Fl. Fennica 27: 105, 1952 (Herzog 1952c). ** Paracromastigum subsimplex (Steph.) Fulford et J.Taylor, Brittonia 13 (4): 336, 1961 (Fulford and Taylor 1961). Bas.: Lepidozia subsimplex Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 66, 1911 (Stephani 1911b). ** Paracromastigum succulentum (Sim) J.J.Engel et G.L.Merr., Bryologist 104 (1): 151, 2001 (Engel and Smith Merrill 2001). Bas.: Lepidozia succulenta Sim, Trans. Roy. Soc. South Africa 15 (1): 90, 1926 (Sim 1926). ** Paracromastigum tristanianum (R.M.Schust.) J.J.Engel et R.M.Schust., J. Hattori Bot. Lab. 38: 700, 1974 (Schuster and Engel 1974). Bas.: Pseudocephalozia tristaniana R.M.Schust., Nova Hedwigia 10 (1/2): 23, 1965 (Schuster 1965b). World checklist of hornworts and liverworts 185 ** Paracromastigum vastilobum (Steph.) J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 255, 2004 (Engel and Smith Merrill 2004). Bas.: Lepidozia vastiloba Steph., Sp. Hepat. (Stephani) 3: 581, 1909 (Stephani 1909a). ** Psiloclada Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 143, 1854 (Mitten 1854). *** Psiloclada clandestina Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 143, 1854 (Mitten 1854). ** Psiloclada clandestina subsp. melanesica R.M.Schust., J. Hattori Bot. Lab. 48: 411, 1980 (Schuster 1980a). ** Psiloclada clandestina subsp. spinosa (S.W.Arnell) R.M.Schust., J. Hattori Bot. Lab. 48: 410, 1980 (Schuster 1980a). Bas.: Lepidozia spinosa S.W.Arnell, Bot. Not. 107: 427, 1954 (Arnell 1954c). ** Pteropsiella Spruce, J. Bot. 14: 161, 1876 (Spruce 1876b). *** Pteropsiella frondiformis Spruce, J. Bot. 14: 161, 1876 (Spruce 1876b). ** Pteropsiella metzgeriiformis R.M.Schust., Nova Hedwigia 10 (1/2): 25, 1965 (Schuster 1965b). ** Telaranea Spruce ex Schifn., Hepat. (Engl.-Prantl): 103, 1893 (Schifner 1893b) nom. conserv. Based on: Telaranea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 365, 1885 (Spruce 1885). 129 * sect. Telaranea *** Telaranea apiahyna (Steph.) Fulford, Brittonia 15 (1): 71, 1963 (Fulford 1963). Bas.: Lepidozia apiahyna Steph., Sp. Hepat. (Stephani) 3: 572, 1909 (Stephani 1909a). *** Telaranea bicruris (Steph.) M.Howe, Bull. Torrey Bot. Club 29 (5): 287, 1902 (Howe 1902). Bas.: Lepidozia bicruris Steph., Hedwigia 24 (4): 166, 1885 (Stephani 1885f). *** Telaranea blepharostoma (Steph.) Fulford, Brittonia 15 (1): 73, 1963 (Fulford 1963). Bas.: Lepidozia blepharostoma Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 17): 22, 1901 (Stephani 1901b). *** Telaranea breviseta (Herzog) J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 131, 2004 (Engel and Smith Merrill 2004). Bas.: Lepidozia sejuncta var. breviseta Herzog, Nat. Hist. Juan Fernandez (Botany) 2 (5): 723, 1942 (Herzog 1942a). *** Telaranea chaetophylla (Spruce) Schifn., Hepat. (Engl.-Prantl): 103, 1893 (Schifner 1893b). Bas.: Lepidozia chaetophylla Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 365, 1885 (Spruce 1885). *** Telaranea europaea J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 150, 2004 (Engel and Smith Merrill 2004). 129 Telaranea is a heterogeneous genus and several segregates are warranted. Additional studies are needed to clarify the relationships of the species retained here (Cooper 2013). 186 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Telaranea fragilis Mizut., J. Hattori Bot. Lab. 40: 449, 1976 (Mizutani 1976a). *** Telaranea granulata J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 103, 2004 (Engel and Smith Merrill 2004). *** Telaranea longifolia (M.Howe) J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 163, 2004 (Engel and Smith Merrill 2004). Bas.: Telaranea nematodes var. longifolia M.Howe, Bull. Torrey Bot. Club 29 (5): 286, 1902 (Howe 1902). *** Telaranea nematodes (Gottsche ex Austin) M.Howe, Bull. Torrey Bot. Club 29 (5): 284, 1902 (Howe 1902). Bas.: Cephalozia nematodes Gottsche ex Austin, Bull. Torrey Bot. Club 6 (52): 302, 1879 (Austin 1879). *** Telaranea panchoi Del Ros., Philipp. J. Sci. 100 (3/4): 238, 1971 (Del Rosario 1971). *** Telaranea pellucida J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 179, 2004 (Engel and Smith Merrill 2004). *** Telaranea pseudozoopsis (Herzog) Fulford, Brittonia 15 (1): 71, 1963 (Fulford 1963). Bas.: Lepidozia pseudozoopsis Herzog, Nat. Hist. Juan Fernandez (Botany) 2 (5): 723, 1942 (Herzog 1942a). *** Telaranea redacta (Steph.) J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 186, 2004 (Engel and Smith Merrill 2004). Bas.: Lepidozia redacta Steph., Wiss. Ergebn. Deut. Zentr.-Afr. Exped. (1907-08), Bot. 2: 119, 1911 (Stephani 1911a). *** Telaranea rosarioana H.A.Mill., J. Bryol. 14 (2): 240, 1986 (Miller 1986). *** Telaranea setosa J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 192, 2004 (Engel and Smith Merrill 2004). *** Telaranea trisetosa (Steph.) Grolle, J. Hattori Bot. Lab. 29: 280, 1966 (Grolle 1966g). Bas.: Lepidozia trisetosa Steph., Sp. Hepat. (Stephani) 3: 607, 1909 (Stephani 1909a). * *** ** *** *** *** *** sect. Tenuifoliae (R.M.Schust.) J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 112, 2004 (Engel and Smith Merrill 2004). Bas.: Arachniopsis sect. Tenuifoliae R.M.Schust., Beih. Nova Hedwigia 118: 461, 2000 (Schuster 2000a). Telaranea anomala R.M.Schust. ex J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 121, 2004 (Engel and Smith Merrill 2004). Telaranea bischleriana Pócs, Acta Bot. Hung. 48 (1/2): 120, 2006 (Pócs 2006c). Telaranea coactilis (Spruce) J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 140, 2004 (Engel and Smith Merrill 2004). Bas.: Arachniopsis coactilis Spruce, Cephalozia: 85, 1882 (Spruce 1882). Telaranea confervoides J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 143, 2004 (Engel and Smith Merrill 2004). Based on: Arachniopsis pecten var. confervoides R.M.Schust., Beih. Nova Hedwigia 118: 455, 2000 (Schuster 2000a), nom. inval. Telaranea diacantha (Mont.) J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 145, 2004 (Engel and Smith Merrill 2004). Bas.: Jungermannia diacantha Mont., Ann. Sci. Nat. Bot. (sér. 4) 5: 349, 1856 (Montagne 1856c). Telaranea herzogii (E.A.Hodgs.) E.A.Hodgs., Rec. Domin. Mus. 4 (11): 106, 1962 (Hodgson 1962a). Bas.: Lepidozia herzogii E.A.Hodgs., Trans. & Proc. Roy. Soc. New Zealand 78 (4): 500, 1950 (Martin 1950). World checklist of hornworts and liverworts 187 *** Telaranea inaequalis R.M.Schust. ex J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 117, 2004 (Engel and Smith Merrill 2004). ** Telaranea maorensis Pócs, Acta Bot. Hung. 48 (1/2): 124, 2006 (Pócs 2006c). *** Telaranea microstipulata R.M.Schust., Phytologia 39 (4): 241, 1978 (Schuster 1978a). *** Telaranea monocera (Mitt. ex R.M.Schust. et Grolle) J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 168, 2004 (Engel and Smith Merrill 2004). Bas.: Arachniopsis monocera Mitt. ex R.M.Schust. et Grolle, Nova Hedwigia 10 (1/2): 25, 1965 (Schuster 1965b). *** Telaranea pecten (Spruce) J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 178, 2004 (Engel and Smith Merrill 2004). Bas.: Arachniopsis pecten Spruce, Cephalozia: 85, 1882 (Spruce 1882). *** Telaranea sejuncta (Ångstr.) S.W.Arnell, Bot. Not. 110 (1): 18, 1957 (Arnell 1957a). Bas.: Blepharostoma sejunctum Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 33 (7): 78, 1876 [1877] (Ångström 1876). *** Telaranea tenuifolia J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 193, 2004 (Engel and Smith Merrill 2004). Based on: Arachniopsis tenuifolia R.M.Schust., Beih. Nova Hedwigia 118: 461, 2000 (Schuster 2000a), nom. inval. Incertae sedis ** Telaranea azorica (H.Buch et Perss.) Pócs in Schumacker et Váňa, Identif. keys liverw. hornw. Europe: 160, 2005 (Schumacker and Váňa 2005). Bas.: Lepidozia azorica H.Buch et Perss., Bryophyt. Azoren Madeira: 4, 1941 (Buch and Persson 1941). ** Telaranea indica (S.C.Srivast. et P.K.Verma) A.E.D.Daniels et P.Daniel, Bull. Bot. Surv. India 49 (1/4): 231, 2007 (Daniels and Daniel 2007). Bas.: Arachniopsis indica S.C.Srivast. et P.K.Verma, Natl. Acad. Sci. Lett. 27 (7/8): 270, 2004 [2006] (Srivastava and Verma 2004). ** Telaranea major (Herzog) J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 165, 2004 (Engel and Smith Merrill 2004). Bas.: Arachniopsis major Herzog, Trans. Brit. Bryol. Soc. 1 (4): 294, 1950 (Herzog 1950a). ** Zoopsidella R.M.Schust., Nova Hedwigia 10 (1/2): 24, 1965 (Schuster 1965b). ** Zoopsidella antillana (Steph.) H.Rob., Bol. Soc. Venez. Ci. Nat. 32 (132/133): 254, 1976 (Robinson 1976b). Bas.: Zoopsis antillana Steph., Bull. Herb. Boissier (sér. 2) 8 (4): 268 (282), 1908 (Stephani 1908j). ** Zoopsidella antillana subsp. jamaicensis R.M.Schust., Nova Hedwigia 69 (1/2): 132, 1999 (Schuster 1999b). *** Zoopsidella caledonica (Steph.) R.M.Schust., Taxon 18 (1): 57, 1969 (Schuster 1969c). Bas.: Zoopsis caledonica Steph., Sp. Hepat. (Stephani) 6: 318, 1922 (Stephani 1922). ** Zoopsidella cynosurandra (Steph.) R.M.Schust., Nova Hedwigia 10 (1/2): 24, 1965 (Schuster 1965b). Bas.: Zoopsis cynosurandra Steph., Bull. Herb. Boissier (sér. 2) 8 (4): 269 (283), 1908 (Stephani 1908j). 188 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Zoopsidella integrifolia (Spruce) R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 12 (3): 666, 1969 (Schuster 1969a). Bas.: Cephalozia integrifolia Spruce, Cephalozia: 29, 1882 (Spruce 1882). ** Zoopsidella macella (Spruce) R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 12 (3): 666, 1969 (Schuster 1969a). Bas.: Cephalozia macella Spruce, Cephalozia: 29, 1882 (Spruce 1882). *** Zoopsidella serra (Spruce) R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 12 (3): 666, 1969 (Schuster 1969a). Bas.: Cephalozia serra Spruce, Cephalozia: 32, 1882 (Spruce 1882). ** Zoopsis Hook.f. ex Gottsche, Lindenb. et Nees, Syn. Hepat. 4: 473, 1846 (Gottsche et al. 1846). ** subg. Eozoopsis R.M.Schust., J. Hattori Bot. Lab. 36: 373, 1972 (Schuster 1972). *** Zoopsis leitgebiana (Carrington et Pearson) Bastow, Pap. & Proc. Roy. Soc. Tasmania 1887: 269, 1888 (Bastow 1888). Bas.: Cephalozia leitgebiana Carrington et Pearson, Pap. & Proc. Roy. Soc. Tasmania 1887: 3, 1888 (Carrington and Pearson 1888b). *** Zoopsis macrophylla R.M.Schust., Nova Hedwigia 68 (1/2): 14, 1999 (Schuster 1999d). ** subg. Zoopsis *** Zoopsis argentea (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 4: 473, 1846 (Gottsche et al. 1846). Bas.: Jungermannia argentea Hook.f. et Taylor, London J. Bot. 3: 400, 1844 (Hooker and Taylor 1844a). ** Zoopsis argentea var. lagelliformis (Colenso) R.M.Schust., Nova Hedwigia 68 (1/2): 38, 1999 (Schuster 1999d). Bas.: Zoopsis lagelliformis Colenso, Trans. & Proc. New Zealand Inst. 18: 250, 1886 (Colenso 1886b). *** Zoopsis bicruris Glenny et E.A.Br., J. Bryol. 28 (4): 332, 2006 (Renner et al. 2006). ** Zoopsis liukiuensis Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 1: 65, 1931 (Horikawa 1931a). *** Zoopsis matawaia M.A.M.Renner, J. Bryol. 28 (4): 334, 2006 (Renner et al. 2006). *** Zoopsis nitida Glenny, Braggins et R.M.Schust., J. Bryol. 19 (4): 776, 1997 (Glenny et al. 1997). *** Zoopsis setulosa Leitg., Mitt. Naturwiss. Vereins Steiermark 13: 24, 1876 (Leitgeb 1876). Incertae sedis *** Zoopsis ceratophylla (Spruce) Hamlin, Rec. Domin. Mus. 7: 311, 1972 (Hamlin 1972). Bas.: Cephalozia ceratophylla Spruce, Cephalozia: 32, 1882 (Spruce 1882). World checklist of hornworts and liverworts 189 * Zoopsis ciliata Colenso, Trans. & Proc. New Zealand Inst. 20: 253, 1888 (Colenso 1888). 130 * Zoopsis martinicensis Steph., Bull. Herb. Boissier (sér. 2) 8 (4): 268 (282), 1908 (Stephani 1908j). ** Zoopsis setigera K.I.Goebel, Flora 77 (2): 93, 1893 (Goebel 1893b). ** Zoopsis uleana Steph., Hedwigia 44 (4): 225, 1905 (Stephani 1905a). *** Lophocoleaceae Vanden Berghen by B.J. Crandall-Stotler, R. Stotler, J. Váňa, J.J.Engel and L. Söderström Söderström et al. (2013b) outlined the current status of Lophocoleaceae noting that several taxonomic problems in delimitating genera remain. he placement of several species is also unclear. Nomenclatural and taxonomic notes can be found in Söderström et al. (2013b, 2013f ). ** Bragginsella R.M.Schust., Bryologist 100 (3): 363, 1997 (Schuster 1997a). *** Bragginsella anomala R.M.Schust., Bryologist 100 (3): 363, 1997 (Schuster 1997a). *** Chiloscyphus Corda, Gen. hepat.: 651, 1829 (Corda 1829) nom. conserv. ** Chiloscyphus kashyapii A.Srivast. et S.C.Srivast., Indian Geocalyc.: 34, 2002 (Srivastava and Srivastava 2002). *** Chiloscyphus pallescens (Ehrh.) Dumort., Syll. Jungerm. Europ.: 67, 1831 (Dumortier 1831). Bas.: Jungermannia pallescens Ehrh., Deutschl. Fl., heil 2 (Hofm.): 87, 1795 [1796] (Hofmann 1795). * Chiloscyphus pallescens var. fragilis (Roth) Müll.Frib., Ber. Deutsch. Bot. Ges. 59 (10): 429, 1942 (Müller 1942). Bas.: Jungermannia fragilis Roth, Tent. Fl. Germ. 3: 370, 1800 (Roth 1800). 131 *** Chiloscyphus polyanthos (L.) Corda, Gen. hepat.: 651, 1829 (Corda 1829). Bas.: Jungermannia polyanthos L., Sp. Pl. 1: 1131, 1753 (Linnaeus 1753). * Chiloscyphus polyanthos var. rivularis (Schrad.) Lindb. et Arnell, Kongl. Svenska Vetensk.-Akad. Handl. (n.ser.) 23 (5): 24, 1889 (Lindberg and Arnell 1889). Bas.: Jungermannia pallescens var. rivularis Schrad., Syst. Samml. Crypt. Gew. 2: 7, 1797 (Schrader 1797). 132 130 Zoopsis ciliata was not studied by Engel and Glenny (2008a) and they did not know what it may be. 131 Chiloscyphus pallescens var. fragilis is a problematic taxon sometimes treated as a species, sometimes as conspeciic with Chiloscyphus pallescens, sometimes as conspeciic with Chiloscyphus polyanthos. 132 Chiloscyphus polyanthos var. rivularis is a problematic taxon sometimes treated as a species, sometimes as conspeciic with Chiloscyphus polyanthos. 190 ** ** ** * ** * ** ** ** ** * * ** ** ** ** ** * ** ** ** ** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Incertae sedis 133 Chiloscyphus acutus Steph., Sp. Hepat. (Stephani) 6: 302, 1922 (Stephani 1922). Chiloscyphus alpicola J.J.Engel, Phytotaxa 207 (2): 181, 2015 (Engel 2015b). Chiloscyphus beesleyanus Pearson, J. Linn. Soc., Bot. 46 (305): 22, 1922 (Pearson 1922b). Chiloscyphus biidus Schifn., Hep. Fl. Buitenzorg: 200, 1900 (Schifner 1900a). 134 Chiloscyphus breviculus B.Y.Yang et W.C.Lee, Bot. Bull. Acad. Sin. (n.ser.) 5 (2): 190, 1964 (Yang and Lee 1964). Chiloscyphus brevistipulus Steph., Sp. Hepat. (Stephani) 6: 303, 1922 (Stephani 1922). Chiloscyphus chinnarensis Manju, K.P.Rajesh et Madhus., Acta Bot. Hung. 53 (1/2): 152, 2011 (Manju et al. 2011). Chiloscyphus confertifolius Steph., Sp. Hepat. (Stephani) 6: 304, 1922 (Stephani 1922). Chiloscyphus confertus Steph., Sp. Hepat. (Stephani) 6: 305, 1922 (Stephani 1922). Chiloscyphus cornutistipulus Steph., Sp. Hepat. (Stephani) 6: 303, 1922 (Stephani 1922). Chiloscyphus durus (Steph.) Hässel, Revista Mus. Argent. Ci. Nat. (n.ser.) 1 (2): 122, 1999 (Hässel 1999). Bas.: Lophocolea dura Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 43, 1911 (Stephani 1911b). 135 Chiloscyphus ernstianus Steph., Sp. Hepat. (Stephani) 6: 306, 1922 (Stephani 1922). Chiloscyphus etesseanus Steph., Bull. Herb. Boissier (sér. 2) 7 (10): 845 (217), 1907 (Stephani 1907b). Chiloscyphus francanus Steph., Sp. Hepat. (Stephani) 6: 306, 1922 (Stephani 1922). Chiloscyphus graefeanus Steph., Sp. Hepat. (Stephani) 6: 307, 1922 (Stephani 1922). Chiloscyphus greenwelliae (H.A.Mill.) H.A.Mill., J. Hattori Bot. Lab. 30: 275, 1967 (Miller 1967). Bas.: Lophocolea greenwelliae H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 504, 1963 (Miller 1963). Chiloscyphus hookeri J.J.Engel, J. Hattori Bot. Lab. 36: 150, 1972 [1973] (Engel 1972). Chiloscyphus hookeri var. constantifolius J.J.Engel, J. Hattori Bot. Lab. 36: 155, 1972 [1973] (Engel 1972). Chiloscyphus integerrimus Schifn., Hep. Fl. Buitenzorg: 197, 1900 (Schifner 1900a). Chiloscyphus kehdingianus (Steph.) N.Kitag., Hikobia, Suppl. 1: 68, 1981 (Kitagawa 1981a). Bas.: Lophocolea kehdingiana Steph., Sp. Hepat. (Stephani) 6: 278, 1922 (Stephani 1922). Chiloscyphus kilauensis Steph., Sp. Hepat. (Stephani) 6: 309, 1922 (Stephani 1922). Chiloscyphus koeppensis (Gottsche) Steph., Bull. Herb. Boissier (sér. 2) 8 (2): 139 (255), 1908 (Stephani 1908i). Bas.: Jungermannia koeppensis Gottsche, Int. Polarforsch., Deutsch. Exped. 2: 452, 1890 (Gottsche 1890). 133 Chiloscyphus is here treated in a very narrow sense (cf. Söderström et al. 2013b), but a large number of species remain to be assigned to other genera. It is unclear if the taxa included here belong to Cryptolophocolea, Lophocolea or some other genus in Lophocoleaceae. 134 Chiloscyphus biidus is possibly conspeciic with Heteroscyphus aselliformis (Schifner 1900a). 135 Chiloscyphus durus is conspeciic with Leptoscyphus expansus in Grolle (1962a), but it probably belongs to one of its segregates. It was accepted by Hässel (1999). World checklist of hornworts and liverworts ** ** ** ** ** ** ** ** *** ** ** ** ** ** ** ** ** * ** * 191 Chiloscyphus laceratus Steph., Sp. Hepat. (Stephani) 6: 310, 1922 (Stephani 1922). Chiloscyphus lambertonii H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 506, 1963 (Miller 1963). Chiloscyphus latistipus Steph., Sp. Hepat. (Stephani) 6: 309, 1922 (Stephani 1922). Chiloscyphus lepervanchei (Steph.) J.J.Engel et R.M.Schust., Nova Hedwigia 39: 418, 1984 [1985] (Engel and Schuster 1984). Bas.: Lophocolea lepervanchei Steph., Bull. Herb. Boissier (sér. 2) 7 (4): 310 (174), 1907 (Stephani 1907d). Chiloscyphus longiissus Steph., Sp. Hepat. (Stephani) 6: 310, 1922 (Stephani 1922). Chiloscyphus propagulifer Schifn., Hep. Fl. Buitenzorg: 208, 1900 (Schifner 1900a). Chiloscyphus purpureus Steph., Sp. Hepat. (Stephani) 6: 312, 1922 (Stephani 1922). Chiloscyphus quadricilius Steph., Sp. Hepat. (Stephani) 6: 312, 1922 (Stephani 1922). Chiloscyphus quadridentatus (Spruce) J.J.Engel et R.M.Schust., Nova Hedwigia 39: 422, 1984 [1985] (Engel and Schuster 1984). Bas.: Lophocolea quadridentata Spruce, Mem. Torrey Bot. Club 1 (3): 137, 1890 (Spruce 1890). Chiloscyphus rotundifolius Mitt., Rep. Challenger, Bot. 1 (3, 1): 85, 1884 (Mitten 1884b). Chiloscyphus rotundiphyllus H.A.Mill., Phytologia 47 (4): 321, 1981 (Miller 1981). Nom. nov. pro Chiloscyphus rotundifolius Steph., Sp. Hepat. (Stephani) 6: 313, 1922 (Stephani 1922), nom. illeg. Chiloscyphus scaberulus Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: cc, 1889 [1890] (Spruce 1889). Chiloscyphus septatus J.J.Engel, Fieldiana, Bot. (n.ser.) 48: 125, 2010 (Engel 2010). Chiloscyphus similis Steph., Rev. Bryol. 35 (2): 28, 1908 (Stephani 1908l). Chiloscyphus skottsbergianus H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 508, 1963 (Miller 1963). Chiloscyphus subacuminatus Herzog, Ark. Bot. (n.ser.) 3 (3): 49, 1953 (Herzog 1953a). Chiloscyphus subsimilis Steph., Sp. Hepat. (Stephani) 6: 314, 1922 (Stephani 1922). Chiloscyphus tridens Steph., Sp. Hepat. (Stephani) 6: 315, 1922 (Stephani 1922). Chiloscyphus trigonifolius Steph., Sp. Hepat. (Stephani) 6: 316, 1922 (Stephani 1922). Chiloscyphus venustulus Colenso, Trans. & Proc. New Zealand Inst. 21: 60, 1889 (Colenso 1889). ** Clasmatocolea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 440, 1885 (Spruce 1885). 136 *** Clasmatocolea bisexualis Glenny et J.J.Engel, New Zealand J. Bot. 51 (1): 23, 2013 (Glenny and Engel 2013). *** Clasmatocolea crassiretis (Herzog) Grolle, Nova Acta Leop. (n.ser.) 25 (161): 69, 1962 [1963] (Grolle 1962a). Bas.: Lophocolea crassiretis Herzog, Trans. & Proc. Roy. Soc. New Zealand 65 (3): 354, 1936 (Herzog 1936b). *** Clasmatocolea ctenophylla (Schifn.) Grolle, Rev. Bryol. Lichénol. 29 (1/2): 71, 1960 (Grolle 1960c). Bas.: Lophocolea ctenophylla Schifn., Leberm., Forschungsr. Gazelle 4 (4): 12, 1890 (Schifner 1890). 136 Clasmatocolea is here treated without subdivisions although Engel (1980a, 2015) did so. 192 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Clasmatocolea cucullistipula (Steph.) Grolle, Rev. Bryol. Lichénol. 29 (1/2): 71, 1960 (Grolle 1960c). Bas.: Lophocolea cucullistipula Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 6): 37, 1900 (Stephani 1900b). *** Clasmatocolea fasciculata (Nees) Grolle, Rev. Bryol. Lichénol. 29 (1/2): 72, 1960 (Grolle 1960c). Bas.: Jungermannia fasciculata Nees, Horae Phys. Berol.: 46, 1820 (Nees 1820). *** Clasmatocolea fulvella (Hook.f. et Taylor) Grolle, Rev. Bryol. Lichénol. 29 (1/2): 72, 1960 (Grolle 1960c). Bas.: Jungermannia fulvella Hook.f. et Taylor, London J. Bot. 3: 464, 1844 (Hooker and Taylor 1844b). *** Clasmatocolea gayana (Mont.) Grolle, Rev. Bryol. Lichénol. 29 (1/2): 72, 1960 (Grolle 1960c). Bas.: Jungermannia gayana Mont., Ann. Sci. Nat. Bot. (sér. 3) 4: 349, 1845 (Montagne 1845b). *** Clasmatocolea humilis (Hook.f. et Taylor) Grolle, Rev. Bryol. Lichénol. 29 (1/2): 72, 1960 (Grolle 1960c). Bas.: Jungermannia humilis Hook.f. et Taylor, London J. Bot. 3: 468, 1844 (Hooker and Taylor 1844b). *** Clasmatocolea humilis var. polymorpha J.J.Engel, Phytologia 41 (5): 309, 1979 (Engel 1979a). *** Clasmatocolea humilis var. suspecta (C.Massal.) J.J.Engel, Phytologia 41 (5): 309, 1979 (Engel 1979a). Bas.: Lophocolea puccioana β suspecta C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 228, 1885 (Massalongo 1885). *** Clasmatocolea inlexispina (Hook.f. et Taylor) J.J.Engel, Bryologist 94 (4): 436, 1991 (Engel 1991b). Bas.: Jungermannia inlexispina Hook.f. et Taylor, London J. Bot. 4: 82, 1845 (Hooker and Taylor 1845). *** Clasmatocolea marginata (Steph.) Grolle, Nova Acta Leop. (n.ser.) 25 (161): 73, 1962 [1963] (Grolle 1962a). Bas.: Leioscyphus marginatus Steph., Bull. Herb. Boissier (sér. 2) 6 (3): 223 (23), 1906 (Stephani 1906g). *** Clasmatocolea minutiretis J.J.Engel et Grolle, Phytologia 41 (5): 309, 1979 (Engel 1979a). *** Clasmatocolea moniliformis J.J.Engel, Phytologia 41 (5): 310, 1979 (Engel 1979a). *** Clasmatocolea navistipula (Steph.) Grolle, Feddes Repert. 82 (1): 88, 1971 (Grolle 1971b). Bas.: Lophocolea navistipula Steph., Bull. Herb. Boissier (sér. 2) 6 (7): 543 (57), 1906 (Stephani 1906f). *** Clasmatocolea navistipula var. parceramosa J.J.Engel, Phytologia 41 (5): 311, 1979 (Engel 1979a). *** Clasmatocolea notophylla (Hook.f. et Taylor) Grolle, J. Jap. Bot. 41 (8): 228, 1966 (Grolle 1966d). Bas.: Jungermannia notophylla Hook.f. et Taylor, London J. Bot. 3: 376, 1844 (Hooker and Taylor 1844a). *** Clasmatocolea obvoluta (Hook.f. et Taylor) Grolle, Rev. Bryol. Lichénol. 29 (1/2): 72, 1960 (Grolle 1960c). Bas.: Jungermannia obvoluta Hook.f. et Taylor, London J. Bot. 4: 80, 1845 (Hooker and Taylor 1845). *** Clasmatocolea obvoluta var. cookiana (C.Massal.) J.J.Engel, Phytologia 41 (5): 311, 1979 (Engel 1979a). Bas.: Lophocolea cookiana C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 224, 1885 (Massalongo 1885). World checklist of hornworts and liverworts 193 *** Clasmatocolea puccioana (De Not.) Grolle, Rev. Bryol. Lichénol. 29 (1/2): 72, 1960 (Grolle 1960c). Bas.: Jungermannia puccioana De Not., Mem. Reale Accad. Sci. Torino (ser. 2) 16: 221, 1857 (De Notaris 1857). *** Clasmatocolea rigens (Hook.f. et Taylor) J.J.Engel, J. Hattori Bot. Lab. 36: 156, 1972 [1973] (Engel 1972). Bas.: Jungermannia rigens Hook.f. et Taylor, London J. Bot. 3: 461, 1844 (Hooker and Taylor 1844b). *** Clasmatocolea strongylophylla (Hook.f. et Taylor) Grolle, Rev. Bryol. Lichénol. 29 (1/2): 73, 1960 (Grolle 1960c). Bas.: Jungermannia strongylophylla Hook.f. et Taylor, London J. Bot. 3: 370, 1844 (Hooker and Taylor 1844a). *** Clasmatocolea trachyopa (Hook.f. et Taylor) Grolle, Rev. Bryol. Lichénol. 29 (1/2): 73, 1960 (Grolle 1960c). Bas.: Jungermannia trachyopa Hook.f. et Taylor, London J. Bot. 3: 471, 1844 (Hooker and Taylor 1844b). *** Clasmatocolea vermicularis (Lehm.) Grolle, Rev. Bryol. Lichénol. 29 (1/2): 78, 1960 (Grolle 1960c). Bas.: Jungermannia vermicularis Lehm., Linnaea 4: 361, 1829 (Lehmann 1829). *** Clasmatocolea verrucosa J.J.Engel, Bryologist 83 (2): 220, 1980 (Engel 1980b). *** Conoscyphus Mitt., Fl. vit.: 404, 1871 [1873] (Mitten 1871). ** Conoscyphus koponenii Piippo, Mamontov et Potemkin, Acta Bryolichenol. Asiat. 5: 20, 2014 (Piippo et al. 2014). *** Conoscyphus trapezioides (Sande Lac.) Schifn., Consp. Hepat. Arch. Ind.: 125, 1898 (Schifner 1898b). Bas.: Chiloscyphus trapezioides Sande Lac., Ned. Kruidk. Arch. 3: 417, 1854 [1855] (Sande Lacoste 1854). *** Cryptolophocolea L.Söderstr., Crand.-Stotl., Stotler et Váňa, Phytotaxa 97 (2): 39, 2013 (Söderström et al. 2013b). Based on: Plagiochila sect. Connatae Lindenb., Monogr. hep. gen. Plagiochilae: xxix, 1844 [1843] (Lindenberg 1844). *** Cryptolophocolea aculeata (Mitt.) L.Söderstr., Phytotaxa 112 (1): 21, 2013 (Söderström et al. 2013f). Bas.: Chiloscyphus aculeatus Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 140, 1854 (Mitten 1854). ** Cryptolophocolea chiloscyphoidea (Lindenb.) L.Söderstr. et Crand.-Stotl., Phytotaxa 112 (1): 18, 2013 (Söderström et al. 2013f). Bas.: Plagiochila chiloscyphoidea Lindenb., Nov. Stirp. Pug. 8: 4, 1844 (Lehmann 1844). *** Cryptolophocolea ciliolata (Nees) L.Söderstr., Crand.-Stotl., Stotler et Váňa, Phytotaxa 97 (2): 39, 2013 (Söderström et al. 2013b). Bas.: Jungermannia ciliolata Nees, Enum. Pl. Crypt. Javae: 68, 1830 (Nees 1830). ** Cryptolophocolea compacta (Mitt.) L.Söderstr., Phytotaxa 112 (1): 19, 2013 (Söderström et al. 2013f). Bas.: Lophocolea compacta Mitt., Trans. Linn. Soc. London, Bot. 3 (3): 198, 1891 (Mitten 1891). *** Cryptolophocolea connata (Sw.) L.Söderstr. et Váňa, Phytotaxa 112 (1): 19, 2013 (Söderström et al. 2013f). Bas.: Jungermannia connata Sw., Prodr. (Swartz): 143, 1788 (Swartz 1788). 194 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Cryptolophocolea connatifolia (J.J.Engel) L.Söderstr., Phytotaxa 112 (1): 21, 2013 (Söderström et al. 2013f). Bas.: Chiloscyphus connatifolius J.J.Engel, Phytologia 83 (1): 42, 1997 [1998] (Engel 1997). *** Cryptolophocolea costata (Nees) L.Söderstr., Phytotaxa 112 (1): 19, 2013 (Söderström et al. 2013f). Bas.: Jungermannia costata Nees, Enum. Pl. Crypt. Javae: 69, 1830 (Nees 1830). *** Cryptolophocolea edentata (J.J.Engel) L.Söderstr., Phytotaxa 112 (1): 21, 2013 (Söderström et al. 2013f). Bas.: Chiloscyphus edentatus J.J.Engel, Phytologia 83 (1): 43, 1997 [1998] (Engel 1997). ** Cryptolophocolea explanata (Mitt.) Váňa et Crand.-Stotl., Phytotaxa 202 (1): 69, 2015 (Söderström et al. 2015c). Bas.: Lophocolea explanata Mitt., Fl. vit.: 404, 1871 [1873] (Mitten 1871). ** Cryptolophocolea leischeri (Steph.) L.Söderstr., Phytotaxa 112 (1): 21, 2013 (Söderström et al. 2013f). Bas.: Lophocolea leischeri Steph., Bull. Herb. Boissier (sér. 2) 6 (11): 952 (132), 1906 (Stephani 1906c). *** Cryptolophocolea guadalupensis (Steph.) L.Söderstr. et Váňa, Phytotaxa 112 (1): 19, 2013 (Söderström et al. 2013f). Bas.: Lophocolea guadalupensis Steph., Bull. Herb. Boissier (sér. 2) 7 (1): 65 (153), 1907 (Stephani 1907c). *** Cryptolophocolea helmsiana (Steph.) L.Söderstr., Phytotaxa 112 (1): 19, 2013 (Söderström et al. 2013f). Bas.: Lophocolea helmsiana Steph., Bull. Herb. Boissier (sér. 2) 6 (9): 794 (94), 1906 (Stephani 1906e). *** Cryptolophocolea leucophylla (Hook.f. et Taylor) L.Söderstr., Phytotaxa 112 (1): 19, 2013 (Söderström et al. 2013f). Bas.: Jungermannia leucophylla Hook.f. et Taylor, London J. Bot. 3: 384, 1844 (Hooker and Taylor 1844a). * Cryptolophocolea levieri (Schifn.) L.Söderstr., Phytotaxa 112 (1): 21, 2013 (Söderström et al. 2013f). Bas.: Lophocolea levieri Schifn., Hep. Fl. Buitenzorg: 182, 1900 (Schifner 1900a). 137 * Cryptolophocolea lilliena (Steph.) L.Söderstr., Phytotaxa 112 (1): 21, 2013 (Söderström et al. 2013f). Bas.: Lophocolea lilliena Steph., Sp. Hepat. (Stephani) 6: 282, 1922 (Stephani 1922). 138 *** Cryptolophocolea martiana (Nees) L.Söderstr., Crand.-Stotl. et Stotler, Phytotaxa 112 (1): 20, 2013 (Söderström et al. 2013f). Bas.: Lophocolea martiana Nees, Syn. Hepat. 2: 152, 1845 (Gottsche et al. 1845a). ** Cryptolophocolea martiana subsp. bidentula (Nees) L.Söderstr., Crand.-Stotl. et Stotler, Phytotaxa 112 (1): 20, 2013 (Söderström et al. 2013f). Bas.: Chiloscyphus bidentulus Nees, Syn. Hepat. 2: 181, 1845 (Gottsche et al. 1845a). ** Cryptolophocolea martiana var. perissodonta (Spruce) Gradst., Phytoneuron 2015 (22): 1, 2015 (Bernal et al. 2015). Bas.: Lophocolea martiana var. perissodonta Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 432, 1885 (Spruce 1885). 137 Cryptolophocolea levieri is possibly conspeciic with Cryptolophocolea ciliolata (Söderström et al. 2010a). 138 Cryptolophocolea lilliena is possibly conspeciic with Cryptolophocolea martiana (Wigginton and Grolle 1996). World checklist of hornworts and liverworts * *** *** *** *** ** * * *** *** * *** * 195 Cryptolophocolea massalongoana (Schifn.) L.Söderstr., Phytotaxa 112 (1): 20, 2013 (Söderström et al. 2013f). Bas.: Lophocolea massalongoana Schifn., Hep. Fl. Buitenzorg: 183, 1900 (Schifner 1900a). 139 Cryptolophocolea mitteniana (Colenso) L.Söderstr., Phytotaxa 112 (1): 21, 2013 (Söderström et al. 2013f). Bas.: Isotachis mitteniana Colenso, Trans. & Proc. New Zealand Inst. 21: 69, 1889 (Colenso 1889). Cryptolophocolea mitteniana var. obtusa (J.J.Engel) L.Söderstr., Phytotaxa 112 (1): 21, 2013 (Söderström et al. 2013f). Bas.: Chiloscyphus mittenianus var. obtusus J.J.Engel, Phytologia 83 (1): 44, 1997 [1998] (Engel 1997). Cryptolophocolea mitteniana var. symmetrica (J.J.Engel) L.Söderstr., Phytotaxa 112 (1): 22, 2013 (Söderström et al. 2013f). Bas.: Chiloscyphus mittenianus var. symmetricus J.J.Engel, Phytologia 83 (1): 44, 1997 [1998] (Engel 1997). Cryptolophocolea pallida (Mitt.) L.Söderstr., Phytotaxa 112 (1): 22, 2013 (Söderström et al. 2013f). Bas.: Lophocolea pallida Mitt., Bot. antarct. voy. II (Fl. Nov.Zel. 2): 135, 1854 (Mitten 1854). Cryptolophocolea pallidovirens (Hook.f. et Taylor) L.Söderstr., Phytotaxa 112 (1): 20, 2013 (Söderström et al. 2013f). Bas.: Jungermannia pallidovirens Hook.f. et Taylor, London J. Bot. 3: 473, 1844 (Hooker and Taylor 1844b). Cryptolophocolea proteus (Herzog) L.Söderstr., Phytotaxa 112 (1): 22, 2013 (Söderström et al. 2013f). Bas.: Lophocolea proteus Herzog, Feddes Repert. Spec. Nov. Regni Veg. 57 (1/2): 164, 1955 (Herzog 1955). Cryptolophocolea pycnophylla (Spruce) L.Söderstr., Phytotaxa 112 (1): 22, 2013 (Söderström et al. 2013f). Bas.: Lophocolea pycnophylla Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 434, 1885 (Spruce 1885). Cryptolophocolea regularis (Steph.) L.Söderstr., Phytotaxa 112 (1): 23, 2013 (Söderström et al. 2013f). Bas.: Chiloscyphus regularis Steph., Hedwigia 32 (5): 325, 1893 (Stephani 1893d). Cryptolophocolea spinifera (Hook.f. et Taylor) L.Söderstr., Phytotaxa 112 (1): 20, 2013 (Söderström et al. 2013f). Bas.: Jungermannia spinifera Hook.f. et Taylor, London J. Bot. 3: 381, 1844 (Hooker and Taylor 1844a). Cryptolophocolea stephanii (Schifn.) L.Söderstr., Phytotaxa 112 (1): 23, 2013 (Söderström et al. 2013f). Bas.: Lophocolea stephanii Schifn., Hep. Fl. Buitenzorg: 181, 1900 (Schifner 1900a). 140 Cryptolophocolea subopposita (J.J.Engel) L.Söderstr., Phytotaxa 112 (1): 23, 2013 (Söderström et al. 2013f). Bas.: Chiloscyphus suboppositus J.J.Engel, Phytologia 83 (1): 45, 1997 [1998] (Engel 1997). Cryptolophocolea thermarum (Schifn.) L.Söderstr., Phytotaxa 112 (1): 23, 2013 (Söderström et al. 2013f). Bas.: Lophocolea thermarum Schifn., Hep. Fl. Buitenzorg: 180, 1900 (Schifner 1900a). 141 139 Cryptolophocolea massalongoana is possibly conspeciic with Cryptolophocolea costata (Piippo 1985a). 140 Cryptolophocolea stephanii is doubtfully distinct from Cryptolophocolea ciliolata (Söderström et al. 2010a). 141 Cryptolophocolea thermarum is possibly conspeciic with Cryptolophocolea ciliolata (Söderström et al. 2010a). 196 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Cryptolophocolea trialata (Gottsche) L.Söderstr., Phytotaxa 112 (1): 23, 2013 (Söderström et al. 2013f). Bas.: Lophocolea trialata Gottsche, Linnaea 28 (5): 552, 1856 [1857] (Gottsche 1856). ** Cryptolophocolea tricorata (Hässel) Crand.-Stotl. et Stotler, Phytotaxa 112 (1): 23, 2013 (Söderström et al. 2013f). Bas.: Chiloscyphus tricoratus Hässel, Nova Hedwigia 70 (3/4): 456, 2000 (Hässel 2000). *** Cryptolophocolea tuberculata (J.J.Engel) L.Söderstr., Phytotaxa 112 (1): 23, 2013 (Söderström et al. 2013f). Bas.: Chiloscyphus tuberculatus J.J.Engel, Phytologia 83 (1): 45, 1997 [1998] (Engel 1997). * Cryptolophocolea whittieriana (Inoue et H.A.Mill.) L.Söderstr., Phytotaxa 112 (1): 24, 2013 (Söderström et al. 2013f). Bas.: Lophocolea whittieriana Inoue et H.A.Mill., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 8 (2): 143, 1965 (Inoue and Miller 1965). 142 *** Deceptifrons J.J.Engel et Váňa, Mem. New York Bot. Gard. 105: 54, 2013 (Váňa and Engel 2013). *** Deceptifrons plagiochiloides J.J.Engel et Váňa, Mem. New York Bot. Gard. 105: 54, 2013 (Váňa and Engel 2013). *** Evansianthus R.M.Schust. et J.J.Engel, Bryologist 76 (4): 516, 1973 (Schuster and Engel 1973). *** Evansianthus georgiensis (Gottsche) R.M.Schust. et J.J.Engel, Bryologist 76 (4): 518, 1973 (Schuster and Engel 1973). Bas.: Lophocolea georgiensis Gottsche, Int. Polarforsch., Deutsch. Exped. 2: 453, 1890 (Gottsche 1890). *** Hepatostolonophora J.J.Engel et R.M.Schust., J. Hattori Bot. Lab. 46: 91, 1979 (Engel 1979b). Nom. nov. pro Stolonophora J.J.Engel et R.M.Schust., Fieldiana, Bot. 36 (11): 111, 1975 (Engel and Schuster 1975). *** Hepatostolonophora abnormis (Besch. et C.Massal.) J.J.Engel et R.M.Schust., J. Hattori Bot. Lab. 46: 94, 1979 (Engel 1979b). Bas.: Leioscyphus abnormis Besch. et C.Massal., Bull. Mens. Soc. Linn. Paris 1 (79): 629, 1886 (Bescherelle and Massalongo 1886). ** Hepatostolonophora conica (Steph.) Hässel, Bol. Soc. Argent. Bot. 22 (1/4): 123, 1983 (Hässel 1983). Bas.: Plagiochila conica Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 28, 1911 (Stephani 1911b). *** Hepatostolonophora paucistipula (Rodway) J.J.Engel, J. Hattori Bot. Lab. 46: 103, 1979 (Engel 1979b). Bas.: Lophocolea paucistipula Rodway, Pap. & Proc. Roy. Soc. Tasmania 1916: 46, 1917 (Rodway 1917a). *** Hepatostolonophora rotata (Hook.f. et Taylor) J.J.Engel, J. Hattori Bot. Lab. 46: 98, 1979 (Engel 1979b). Bas.: Jungermannia rotata Hook.f. et Taylor, London J. Bot. 3: 560, 1844 (Hooker and Taylor 1844d). 142 Cryptolophocolea whittieriana is possibly conspeciic with Cryptolophocolea helmsiana (Hodgson 1967). World checklist of hornworts and liverworts 197 *** Hepatostolonophora rotata var. perssonii (R.M.Schust.) J.J.Engel, J. Hattori Bot. Lab. 46: 103, 1979 (Engel 1979b). Bas.: Calyptrocolea perssonii R.M.Schust., Rev. Bryol. Lichénol. 34 (3/4): 699, 1966 [1967] (Schuster 1966a). *** Heteroscyphus Schifn., Österr. Bot. Z. 60 (5): 171, 1910 (Schifner 1910a) nom. conserv. ** Heteroscyphus acutangulus (Schifn.) Schifn., Österr. Bot. Z. 60 (5): 172, 1910 (Schifner 1910a). Bas.: Chiloscyphus acutangulus Schifn., Hep. Fl. Buitenzorg: 200, 1900 (Schifner 1900a). ** Heteroscyphus allodontus (Hook.f. et Taylor) J.J.Engel et R.M.Schust., Nova Hedwigia 39: 399, 1984 [1985] (Engel and Schuster 1984). Bas.: Jungermannia allodonta Hook.f. et Taylor, London J. Bot. 3: 382, 1844 (Hooker and Taylor 1844a). ** Heteroscyphus amboinensis (Schifn.) Schifn., Österr. Bot. Z. 60 (5): 172, 1910 (Schifner 1910a). Bas.: Chiloscyphus endlicherianus var. amboinensis Schifn., Leberm., Forschungsr. Gazelle 4 (4): 15, 1890 (Schifner 1890). ** Heteroscyphus ammophilus (Colenso) R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 248, 1980 (Schuster 1980c). Bas.: Chiloscyphus ammophilus Colenso, Trans. & Proc. New Zealand Inst. 21: 59, 1889 (Colenso 1889). ** Heteroscyphus ammophilus var. obtusifolius J.J.Engel et G.L.Merr., Nova Hedwigia 99 (1/2): 158, 2014 (Engel 2014). *** Heteroscyphus argutus (Reinw., Blume et Nees) Schifn., Österr. Bot. Z. 60 (5): 172, 1910 (Schifner 1910a). Bas.: Jungermannia arguta Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 206, 1824 [1825] (Reinwardt et al. 1824a). ** Heteroscyphus argutus var. brevidens (Schifn.) Herzog et Nog., J. Hattori Bot. Lab. 14: 40, 1955 (Herzog and Noguchi 1955). Bas.: Chiloscyphus argutus var. brevidens Schifn., Hep. Fl. Buitenzorg: 195, 1900 (Schifner 1900a). *** Heteroscyphus aselliformis (Reinw., Blume et Nees) Schifn., Österr. Bot. Z. 60 (5): 172, 1910 (Schifner 1910a). Bas.: Jungermannia aselliformis Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 412, 1824 [1825] (Reinwardt et al. 1824b). ** Heteroscyphus assurgentifolius J.J.Engel, Nova Hedwigia 99 (1/2): 158, 2014 (Engel 2014). ** Heteroscyphus baduinus (Nees) Schifn., Österr. Bot. Z. 60 (5): 171, 1910 (Schifner 1910a). Bas.: Jungermannia baduina Nees, Enum. Pl. Crypt. Javae: 26, 1830 (Nees 1830). ** Heteroscyphus balnetii (Herzog) Grolle, Acta Bot. Fenn. 125: 68, 1984 (Grolle and Piippo 1984). Bas.: Chiloscyphus balnetii Herzog, Ann. Bryol. 5: 89, 1932 (Herzog 1932a). *** Heteroscyphus billardierei (Schwägr.) Schifn., Österr. Bot. Z. 60 (5): 172, 1910 (Schifner 1910a). Bas.: Jungermannia billardierei Schwägr., Hist. Musc. Hepat. Prodr.: 19, 1814 (Schwägrichen 1814). ** Heteroscyphus billardierei var. clasmatocoleoides (J.J.Engel et G.L.Merr.) J.J.Engel, Nova Hedwigia 100 (3/4): 565, 2015 (Engel 2015a). Bas.: Heteroscyphus circum- 198 ** * * ** ** *** ** ** *** *** *** ** ** *** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) dentatus var. clasmatocoleoides J.J.Engel et G.L.Merr., Polish Bot. J. 58 (1): 95, 2013 (Engel 2013b). Heteroscyphus brassii (Grolle) Grolle, Acta Bot. Fenn. 125: 68, 1984 (Grolle and Piippo 1984). Bas.: Chiloscyphus brassii Grolle, J. Hattori Bot. Lab. 31: 2, 1968 (Grolle 1968a). Heteroscyphus caesius (Schifn.) Schifn., Österr. Bot. Z. 60 (5): 172, 1910 (Schifner 1910a). Bas.: Chiloscyphus caesius Schifn., Hep. Fl. Buitenzorg: 207, 1900 (Schifner 1900a). 143 Heteroscyphus caledonicus (Steph.) Schifn., Österr. Bot. Z. 60 (5): 172, 1910 (Schifner 1910a). Bas.: Chiloscyphus caledonicus Steph., Bull. Herb. Boissier (sér. 2) 7 (10): 844 (216), 1907 (Stephani 1907b). 144 Heteroscyphus chlorophyllus (Hook.f. et Taylor) Schifn., Österr. Bot. Z. 60 (5): 172, 1910 (Schifner 1910a). Bas.: Jungermannia chlorophylla Hook.f. et Taylor, London J. Bot. 3: 562, 1844 (Hooker and Taylor 1844d). Heteroscyphus ciliatus (Steph.) Schifn., Österr. Bot. Z. 60 (5): 172, 1910 (Schifner 1910a). Bas.: Chiloscyphus ciliatus Steph., Hedwigia 32 (5): 320, 1893 (Stephani 1893d). Heteroscyphus coalitus (Hook.) Schifn., Österr. Bot. Z. 60 (5): 172, 1910 (Schifner 1910a). Bas.: Jungermannia coalita Hook., Musci Exot. 2: tab. 123, 1820 (Hooker 1820). Heteroscyphus coalitus var. simplicifolius J.J.Engel, Nova Hedwigia 100 (3/4): 572, 2015 (Engel 2015a). Heteroscyphus combinatus (Nees) Schifn., Österr. Bot. Z. 60 (5): 172, 1910 (Schifner 1910a). Bas.: Jungermannia combinata Nees, Enum. Pl. Crypt. Javae: 22, 1830 (Nees 1830). Heteroscyphus conjugatus (Mitt.) J.J.Engel et R.M.Schust., Nova Hedwigia 39: 399, 1984 [1985] (Engel and Schuster 1984). Bas.: Chiloscyphus conjugatus Mitt., Bot. antarct. voy. III (Fl. Tasman. 2): 227, 1860 (Mitten 1860b). Heteroscyphus contortuplicatus (Nees et Mont.) Grolle, J. Hattori Bot. Lab. 55: 504, 1984 (Grolle 1984a). Bas.: Jungermannia contortuplicata Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 5: 54, 1836 (Nees and Montagne 1836). Heteroscyphus cuneistipulus (Steph.) Schifn., Österr. Bot. Z. 60 (5): 172, 1910 (Schifner 1910a). Bas.: Chiloscyphus cuneistipulus Steph., Hedwigia 32 (5): 322, 1893 (Stephani 1893d). Heteroscyphus darjeelingensis A.Srivast. et S.C.Srivast., Indian Geocalyc.: 115, 2002 (Srivastava and Srivastava 2002). Heteroscyphus deceptifrons J.J.Engel, Nova Hedwigia 99 (1/2): 162, 2014 (Engel 2014). Heteroscyphus dentammophilus J.J.Engel et G.L.Merr., Polish Bot. J. 58 (1): 96, 2013 (Engel 2013b). 143 Heteroscyphus caesius is possibly conspeciic with Heteroscyphus splendens (Söderström et al. 2010a). 144 Heteroscyphus caledonicus is possibly conspeciic with Heteroscyphus amboinensis Miller et al. (1983) with a question mark. World checklist of hornworts and liverworts 199 ** Heteroscyphus denticulatus (Mitt.) Schifn., Beih. Bot. Centralbl. 29: 106, 1912 (Schifner 1912c). Bas.: Chiloscyphus denticulatus Mitt., Nat. hist. Azores: 320, 1870 (Mitten 1870). ** Heteroscyphus deplanchei (Steph.) Schifn., Österr. Bot. Z. 60 (5): 172, 1910 (Schifner 1910a). Bas.: Chiloscyphus deplanchei Steph., Bull. Herb. Boissier (sér. 2) 7 (8): 693 (203), 1907 (Stephani 1907e). ** Heteroscyphus diestianus (Sande Lac.) Piippo, Acta Bot. Fenn. 131: 165, 1985 (Piippo 1985a). Bas.: Chiloscyphus diestianus Sande Lac., Ann. Mus. Bot. Lugduno-Batavi 1: 296, 1864 (Sande Lacoste 1864). ** Heteroscyphus divergenticiliatus (Steph.) Fulford, Mem. New York Bot. Gard. 11 (4): 501, 1976 (Fulford 1976). Bas.: Lophocolea divergenticiliata Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 6): 38, 1900 (Stephani 1900b). ** Heteroscyphus dubius (Gottsche) Schifn., Österr. Bot. Z. 60 (5): 172, 1910 (Schifner 1910a). Bas.: Chiloscyphus dubius Gottsche, Abh. Naturwiss. Vereins Bremen 7: 346, 1882 (Gottsche 1882). *** Heteroscyphus echinellus (Lindenb. et Gottsche) J.J.Engel et Xiao L.He, Bryologist 113 (1): 155, 2010 (Engel and He 2010). Bas.: Lophocolea echinella Lindenb. et Gottsche, Syn. Hepat. 5: 703, 1847 (Gottsche et al. 1847). ** Heteroscyphus echinellus var. hyalinus J.J.Engel, Bryologist 113 (1): 161, 2010 (Engel and He 2010). ** Heteroscyphus elliottii (Steph.) Pagán, Bryologist 42 (1): 7, 1939 (Pagán 1939a). Bas.: Chiloscyphus elliottii Steph., Bull. Herb. Boissier (sér. 2) 8 (1): 55 (231), 1908 (Stephani 1908k). * Heteroscyphus elliottii var. portoricensis Fulford, Mem. New York Bot. Gard. 11 (4): 489, 1976 (Fulford 1976). * Heteroscyphus falcifolius (Steph.) Schifn., Österr. Bot. Z. 60 (5): 172, 1910 (Schifner 1910a). Bas.: Chiloscyphus falcifolius Steph., Bull. Herb. Boissier (sér. 2) 7 (8): 698 (208), 1907 (Stephani 1907e). *** Heteroscyphus issistipus (Hook.f. et Taylor) Schifn., Österr. Bot. Z. 60 (5): 172, 1910 (Schifner 1910a). Bas.: Jungermannia issistipa Hook.f. et Taylor, London J. Bot. 3: 384, 1844 (Hooker and Taylor 1844a). ** Heteroscyphus issistipus var. multispinus (E.A.Hodgs. et Allison) J.J.Engel, Nova Hedwigia 99 (1/2): 165, 2014 (Engel 2014). Bas.: Chiloscyphus multispinus E.A.Hodgs. et Allison, Trans. & Proc. Roy. Soc. New Zealand 73 (1): 37, 1943 (Hodgson 1943). ** Heteroscyphus issistipus var. repandus J.J.Engel, Nova Hedwigia 99 (1/2): 165, 2014 (Engel 2014). ** Heteroscyphus laccidus (Mitt.) A.Srivast. et S.C.Srivast., Indian Geocalyc.: 85, 2002 (Srivastava and Srivastava 2002). Bas.: Lophocolea laccida Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 99, 1860 [1861] (Mitten 1860c). ** Heteroscyphus leischeri (Steph.) D.G.Long et Rubas., Ceylon J. Sci., Biol. Sci. 43 (1): 26, 2014 (Long and Rubasinghe 2014). Bas.: Chiloscyphus leischeri Steph., Sp. Hepat. (Stephani) 6: 306, 1922 (Stephani 1922). 200 * ** ** ** ** ** ** ** ** ** ** *** ** ** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Heteroscyphus fragilicilius (Schifn.) Schifn., Österr. Bot. Z. 60 (5): 172, 1910 (Schifner 1910a). Bas.: Chiloscyphus fragilicilius Schifn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 70: 210, 1900 [1901] (Schifner 1900c). Heteroscyphus furcistipulus (E.A.Hodgs.) J.J.Engel et R.M.Schust., Nova Hedwigia 39: 400, 1984 [1985] (Engel and Schuster 1984). Bas.: Chiloscyphus furcistipulus E.A.Hodgs., Trans. Roy. Soc. New Zealand, Bot. 3 (4): 79, 1965 (Hodgson 1965). Heteroscyphus giganteus (Steph.) Hürl., Bauhinia 12 (1/2): 114, 1998 (Hürlimann 1998). Bas.: Chiloscyphus giganteus Steph., Sp. Hepat. (Stephani) 6: 307, 1922 (Stephani 1922). Heteroscyphus graefei (J.B.Jack et Steph.) Grolle, Wiss. Z. Friedrich-Schiller-Univ. Jena, Math.-Naturwiss. Reihe 29 (4): 645, 1980 (Grolle 1980d). Bas.: Lophocolea graefei J.B.Jack et Steph., Bot. Centralbl. 60 (4): 101, 1894 (Jack and Stephani 1894). Heteroscyphus grandilorus (Steph.) Hürl., Bauhinia 12 (1/2): 114, 1998 (Hürlimann 1998). Bas.: Chiloscyphus grandilorus Steph., Sp. Hepat. (Stephani) 6: 307, 1922 (Stephani 1922). Heteroscyphus grandistipus (Steph.) Schifn., Österr. Bot. Z. 60 (5): 172, 1910 (Schifner 1910a). Bas.: Chiloscyphus grandistipus Steph., Bot. Gaz. 15 (11): 283, 1890 (Stephani 1890c). Heteroscyphus gunnianus (Mitt.) J.J.Engel et R.M.Schust., Nova Hedwigia 39: 400, 1984 [1985] (Engel and Schuster 1984). Bas.: Chiloscyphus gunnianus Mitt., Bot. antarct. voy. III (Fl. Tasman. 2): 228, 1860 (Mitten 1860b). Heteroscyphus hastatus (E.A.Hodgs.) J.J.Engel et R.M.Schust., Nova Hedwigia 39: 400, 1984 [1985] (Engel and Schuster 1984). Bas.: Chiloscyphus hastatus E.A.Hodgs., Trans. Roy. Soc. New Zealand, Bot. 3 (11): 181, 1967 (Hodgson 1967). Heteroscyphus hebridensis (Steph.) Schifn., Österr. Bot. Z. 60 (5): 172, 1910 (Schifner 1910a). Bas.: Chiloscyphus hebridensis Steph., Hedwigia 32 (5): 323, 1893 (Stephani 1893d). Heteroscyphus heterophyllus (Steph.) J.J.Engel et R.M.Schust., Nova Hedwigia 39: 400, 1984 [1985] (Engel and Schuster 1984). Bas.: Chiloscyphus heterophyllus Steph., Sp. Hepat. (Stephani) 6: 308, 1922 (Stephani 1922). Heteroscyphus hyalinus (Steph.) A.Srivast. et S.C.Srivast., Indian Geocalyc.: 118, 2002 (Srivastava and Srivastava 2002). Bas.: Lophocolea hyalina Steph., Bull. Soc. Roy. Bot. Belgique 38 (1): 46, 1899 (Stephani 1899h). Heteroscyphus integrifolius (Lehm. et Lindenb.) Fulford, Mem. New York Bot. Gard. 11 (4): 495, 1976 (Fulford 1976). Bas.: Jungermannia integrifolia Lehm. et Lindenb., Nov. Stirp. Pug. 6: 32, 1834 (Lehmann 1834). Heteroscyphus iwatsukii (S.Hatt.) Piippo, Willdenowia 18 (2): 522, 1989 (Piippo 1989a). Bas.: Saccogynidium iwatsukii S.Hatt., J. Jap. Bot. 39 (7): 206, 1964 (Hattori 1964). Heteroscyphus jackii (Steph.) Schifn., Österr. Bot. Z. 60 (5): 172, 1910 (Schifner 1910a). Bas.: Chiloscyphus jackii Steph., Bot. Centralbl. 60 (4): 102, 1894 (Jack and Stephani 1894). World checklist of hornworts and liverworts 201 *** Heteroscyphus knightii (Steph.) Grolle, J. Hattori Bot. Lab. 61: 251, 1986 [1987] (Grolle 1986a). Bas.: Chiloscyphus knightii Steph., Bull. Herb. Boissier (sér. 2) 8 (2): 129 (245), 1908 (Stephani 1908i). ** Heteroscyphus levieri (Steph.) Schifn., Österr. Bot. Z. 60 (5): 172, 1910 (Schifner 1910a). Bas.: Chiloscyphus levieri Steph., Bull. Herb. Boissier (sér. 2) 8 (2): 132 (248), 1908 (Stephani 1908i). ** Heteroscyphus limosus (Carrington et Pearson) Schifn., Österr. Bot. Z. 60 (5): 172, 1910 (Schifner 1910a). Bas.: Chiloscyphus limosus Carrington et Pearson, Pap. & Proc. Roy. Soc. Tasmania 1887: 6, 1888 (Carrington and Pearson 1888b). ** Heteroscyphus lingulatus (Colenso) J.J.Engel et R.M.Schust., Nova Hedwigia 39: 400, 1984 [1985] (Engel and Schuster 1984). Bas.: Chiloscyphus lingulatus Colenso, Trans. & Proc. New Zealand Inst. 21: 61, 1889 (Colenso 1889). ** Heteroscyphus lophocoleoides S.Hatt., Bull. Tokyo Sci. Mus. 11: 45, 1944 (Hattori 1944d). *** Heteroscyphus lyallii (Mitt.) R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 248, 1980 (Schuster 1980c). Bas.: Chiloscyphus lyallii Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 140, 1854 (Mitten 1854). *** Heteroscyphus magellanicus (Steph.) J.J.Engel et R.M.Schust., Nova Hedwigia 39: 400, 1984 [1985] (Engel and Schuster 1984). Bas.: Chiloscyphus magellanicus Steph., Bull. Herb. Boissier (sér. 2) 8 (2): 140 (256), 1908 (Stephani 1908i). ** Heteroscyphus mamillatus Piippo, Nova Hedwigia 56 (3/4): 360, 1993 (Piippo 1993a). ** Heteroscyphus marginatus (Steph.) Fulford, Mem. New York Bot. Gard. 11 (4): 491, 1976 (Fulford 1976). Bas.: Lophocolea marginata Steph., Sp. Hepat. (Stephani) 6: 282, 1922 (Stephani 1922). *** Heteroscyphus menziesii (Mitt.) J.J.Engel, Polish Bot. J. 58 (1): 99, 2013 (Engel 2013b). Bas.: Chiloscyphus menziesii Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 139, 1854 (Mitten 1854). ** Heteroscyphus merapiensis (Steph.) Piippo, Ann. Bot. Fenn. 30 (3): 200, 1993 (Piippo 1993c). Bas.: Chiloscyphus merapiensis Steph., Sp. Hepat. (Stephani) 6: 311, 1922 (Stephani 1922). ** Heteroscyphus miradorensis (Steph.) Schifn., Österr. Bot. Z. 60 (5): 172, 1910 (Schifner 1910a). Bas.: Chiloscyphus miradorensis Steph., Bull. Herb. Boissier (sér. 2) 8 (1): 56 (232), 1908 (Stephani 1908k). *** Heteroscyphus mononuculus J.J.Engel, J. Hattori Bot. Lab. 90: 241, 2001 (Engel 2001). ** Heteroscyphus mononuculus var. ammophilopsis J.J.Engel, Nova Hedwigia 99 (1/2): 166, 2014 (Engel 2014). ** Heteroscyphus mononuculus var. bilobus J.J.Engel, Nova Hedwigia 99 (1/2): 167, 2014 (Engel 2014). ** Heteroscyphus montagnei (Steph.) Fulford, Mem. New York Bot. Gard. 11 (4): 497, 1976 (Fulford 1976). Bas.: Chiloscyphus montagnei Steph., Bull. Herb. Boissier (sér. 2) 8 (2): 141 (257), 1908 (Stephani 1908i). 202 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Heteroscyphus multiidus (Steph.) J.J.Engel et R.M.Schust., Nova Hedwigia 39: 400, 1984 [1985] (Engel and Schuster 1984). Bas.: Chiloscyphus multiidus Steph., Bull. Herb. Boissier (sér. 2) 8 (2): 132 (248), 1908 (Stephani 1908i). ** Heteroscyphus multiidus var. subintegerrimus J.J.Engel, Nova Hedwigia 100 (3/4): 562, 2015 (Engel 2015a). ** Heteroscyphus nadeaudii (Steph.) Schifn., Österr. Bot. Z. 60 (5): 172, 1910 (Schifner 1910a). Bas.: Chiloscyphus nadeaudii Steph., Bull. Herb. Boissier (sér. 2) 7 (10): 848 (220), 1907 (Stephani 1907b). ** Heteroscyphus orbiculatus A.Srivast. et S.C.Srivast., Indian Geocalyc.: 140, 2002 (Srivastava and Srivastava 2002). ** Heteroscyphus palniensis A.Srivast. et S.C.Srivast., Indian Geocalyc.: 130, 2002 (Srivastava and Srivastava 2002). ** Heteroscyphus pandei S.C.Srivast. et A.Srivast., Lindbergia 15 (6): 197, 1989 [1991] (Srivastava and Srivastava 1989b). *** Heteroscyphus parallelifolius J.J.Engel, Polish Bot. J. 58 (1): 99, 2013 (Engel 2013b). ** Heteroscyphus parvulus (Schifn.) Schifn., Österr. Bot. Z. 60 (5): 171, 1910 (Schifner 1910a). Bas.: Chiloscyphus parvulus Schifn., Hep. Fl. Buitenzorg: 206, 1900 (Schifner 1900a). ** Heteroscyphus parvus A.Srivast. et S.C.Srivast., Indian Geocalyc.: 112, 2002 (Srivastava and Srivastava 2002). ** Heteroscyphus perfoliatus (Mont.) Schifn., Österr. Bot. Z. 60 (5): 171, 1910 (Schifner 1910a). Bas.: Lophocolea perfoliata Mont., Ann. Sci. Nat. Bot. (sér. 2) 18: 12, 1842 (Montagne 1842b). ** Heteroscyphus pertusus (Lehm.) Fulford, Mem. New York Bot. Gard. 11 (4): 505, 1976 (Fulford 1976). Bas.: Chiloscyphus pertusus Lehm., Nov. Stirp. Pug. 10: 7, 1857 (Lehmann 1857). *** Heteroscyphus planiusculus (Hook.f. et Taylor) J.J.Engel, J. Hattori Bot. Lab. 68: 315, 1990 (Engel 1990a). Bas.: Jungermannia planiuscula Hook.f. et Taylor, London J. Bot. 3: 382, 1844 (Hooker and Taylor 1844a). ** Heteroscyphus planus (Mitt.) Schifn., Österr. Bot. Z. 60 (5): 171, 1910 (Schifner 1910a). Bas.: Chiloscyphus planus Mitt., J. Linn. Soc., Bot. 8 (31): 157, 1864 [1865] (Mitten 1864a). *** Heteroscyphus polyblepharis (Spruce) Schifn., Österr. Bot. Z. 60 (5): 172, 1910 (Schifner 1910a). Bas.: Chiloscyphus polyblepharis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 442, 1885 (Spruce 1885). *** Heteroscyphus polychaetus (Spruce) Hentschel et Heinrichs, Taxon 56 (4): 1139, 2007 (Hentschel et al. 2007c). Bas.: Lophocolea polychaeta Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 436, 1885 (Spruce 1885). ** Heteroscyphus polycladus (Hook.f. et Lév.) R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 248, 1980 (Schuster 1980c). Bas.: Jungermannia polyclada Hook.f. et Lév., Choix Pl. Nouv.-Zel.: 8, 1846 (Raoul 1846). World checklist of hornworts and liverworts 203 ** Heteroscyphus rectangulatus (Herzog) Piippo, Ann. Bot. Fenn. 30 (3): 200, 1993 (Piippo 1993c). Bas.: Chiloscyphus rectangulatus Herzog, Ann. Naturhist. Mus. Wien 53 (1): 364, 1942 [1943] (Herzog 1942b). ** Heteroscyphus saccogynoides Herzog, J. Hattori Bot. Lab. 14: 40, 1955 (Herzog and Noguchi 1955). ** Heteroscyphus sarawaketanus Piippo, Acta Bot. Fenn. 131: 143, 1985 (Piippo 1985a). *** Heteroscyphus sinuosus (Hook.) Schifn., Österr. Bot. Z. 60 (5): 172, 1910 (Schifner 1910a). Bas.: Jungermannia sinuosa Hook., Musci Exot. 2: tab. 113, 1820 (Hooker 1820). ** Heteroscyphus spectabilis (Steph.) Schifn., Österr. Bot. Z. 60 (5): 172, 1910 (Schifner 1910a). Bas.: Chiloscyphus spectabilis Steph., Hedwigia 30 (5): 205, 1891 (Stephani 1891a). ** Heteroscyphus spinifer C.Gao, T.Cao et Y.H.Wu, J. Bryol. 26 (2): 97, 2004 (Gao et al. 2004). *** Heteroscyphus splendens (Lehm. et Lindenb.) Grolle, Acta Bot. Fenn. 125: 68, 1984 (Grolle and Piippo 1984). Bas.: Jungermannia splendens Lehm. et Lindenb., Nov. Stirp. Pug. 4: 22, 1832 (Lehmann 1832). ** Heteroscyphus splendidus (E.A.Hodgs.) J.J.Engel et R.M.Schust., Nova Hedwigia 39: 400, 1984 [1985] (Engel and Schuster 1984). Bas.: Chiloscyphus splendidus E.A.Hodgs., Trans. Roy. Soc. New Zealand, Bot. 3 (11): 180, 1967 (Hodgson 1967). *** Heteroscyphus stolonifer J.J.Engel, Polish Bot. J. 58 (1): 102, 2013 (Engel 2013b). *** Heteroscyphus succulentus (Gottsche) Schifn., Österr. Bot. Z. 60 (5): 171, 1910 (Schifner 1910a). Bas.: Chiloscyphus succulentus Gottsche, Natuurk. Tijdschr. Ned.-Indië 4: 576, 1853 (Gottsche 1853). ** Heteroscyphus supinus (Hook.f. et Taylor) R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 248, 1980 (Schuster 1980c). Bas.: Chiloscyphus supinus Hook.f. et Taylor, London J. Bot. 5: 284, 1846 (Taylor 1846a). ** Heteroscyphus tener (Steph.) Schifn., Österr. Bot. Z. 60 (5): 172, 1910 (Schifner 1910a). Bas.: Chiloscyphus tener Steph., Bull. Herb. Boissier (sér. 2) 7 (8): 695 (205), 1907 (Stephani 1907e). *** Heteroscyphus thraustus (Spruce) Fulford, Mem. New York Bot. Gard. 11 (4): 495, 1976 (Fulford 1976). Bas.: Lophocolea thrausta Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 437, 1885 (Spruce 1885). ** Heteroscyphus timppae Piippo, Ann. Bot. Fenn. 29 (3): 246, 1992 (Piippo 1992). *** Heteroscyphus triacanthus (Hook.f. et Lév.) Schifn., Österr. Bot. Z. 60 (5): 172, 1910 (Schifner 1910a). Bas.: Jungermannia triacantha Hook.f. et Lév., Choix Pl. Nouv.-Zel.: 8, 1846 (Raoul 1846). ** Heteroscyphus triacanthus var. magnistipulatus J.J.Engel, Nova Hedwigia 99 (1/2): 167, 2014 (Engel 2014). ** Heteroscyphus tridentatus (Sande Lac.) Grolle, Acta Bot. Fenn. 125: 68, 1984 (Grolle and Piippo 1984). Bas.: Lophocolea tridentata Sande Lac., Ann. Mus. Bot. Lugduno-Batavi 1: 296, 1864 (Sande Lacoste 1864). 204 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Heteroscyphus turgidus (Schifn.) Schifn., Österr. Bot. Z. 60 (5): 171, 1910 (Schifner 1910a). Bas.: Chiloscyphus turgidus Schifn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 70: 212, 1900 [1901] (Schifner 1900c). *** Heteroscyphus valdiviensis (Mont.) Schifn., Österr. Bot. Z. 60 (5): 172, 1910 (Schifner 1910a). Bas.: Chiloscyphus valdiviensis Mont., Ann. Sci. Nat. Bot. (sér. 3) 4: 351, 1845 (Montagne 1845b). ** Heteroscyphus varians (Steph.) J.J.Engel, J. Hattori Bot. Lab. 68: 315, 1990 (Engel 1990a). Bas.: Lophocolea varians Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 119, 1914 (Stephani and Watts 1914). ** Heteroscyphus wettsteinii (Schifn.) Schifn., Österr. Bot. Z. 60 (5): 172, 1910 (Schifner 1910a). Bas.: Chiloscyphus wettsteinii Schifn., Hep. Fl. Buitenzorg: 202, 1900 (Schifner 1900a). ** Heteroscyphus zollingeri (Gottsche) Schifn., Österr. Bot. Z. 60 (5): 171, 1910 (Schifner 1910a). Bas.: Chiloscyphus zollingeri Gottsche, Natuurk. Tijdschr. Ned.-Indië 4: 576, 1853 (Gottsche 1853). *** Lamellocolea J.J.Engel, J. Hattori Bot. Lab. 70: 65, 1991 (Engel 1991c). *** Lamellocolea granditexta (Steph.) J.J.Engel, J. Hattori Bot. Lab. 70: 66, 1991 (Engel 1991c). Bas.: Lophocolea granditexta Steph., Bull. Herb. Boissier (sér. 2) 6 (10): 881 (106), 1906 (Stephani 1906d). *** Lamellocolea integrostia J.J.Engel et Glenny, Bryologist 114 (1): 23, 2011 (Engel and Glenny 2011). *** Leptophyllopsis R.M.Schust., J. Hattori Bot. Lab. 26: 269, 1963 (Schuster 1963b). *** Leptophyllopsis laxa (Mitt.) R.M.Schust. ex Hamlin, Rec. Domin. Mus. 7: 284, 1972 (Hamlin 1972). Bas.: Chiloscyphus laxus Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 142, 1854 (Mitten 1854). *** Leptoscyphopsis R.M.Schust., Phytologia 39 (4): 246, 1978 (Schuster 1978a). *** Leptoscyphopsis paradoxa R.M.Schust., Phytologia 39 (4): 246, 1978 (Schuster 1978a). *** Leptoscyphus Mitt., Hooker’s J. Bot. Kew Gard. Misc. 3: 358, 1851 (Mitten 1851). 145 ** subg. Anomylia (R.M.Schust.) R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 272, 1980 (Schuster 1980c). Bas.: Anomylia R.M.Schust., Amer. Midl. Naturalist 62 (1): 51, 1959 (Schuster 1959a). *** Leptoscyphus cuneifolius (Hook.) Mitt., Hooker’s J. Bot. Kew Gard. Misc. 3: 358, 1851 (Mitten 1851). Bas.: Jungermannia cuneifolia Hook., Brit. Jungermann.: tab. 64, 1814 (Hooker 1814). 145 he subdivision of Leptoscyphus follows Vanderpoorten et al. (2010). World checklist of hornworts and liverworts 205 ** Leptoscyphus cuneifolius subsp. fragilis (J.B.Jack et Steph.) Grolle, Nova Acta Leop. (n.ser.) 25 (161): 28, 1962 [1963] (Grolle 1962a). Bas.: Leioscyphus fragilis J.B.Jack et Steph., Hedwigia 31 (1): 20, 1892 (Jack and Stephani 1892). ** subg. Austroleptoscyphus Vanderp., Schäf.-Verw. et D.G.Long, Taxon 59 (1): 183, 2010 (Vanderpoorten et al. 2010). *** Leptoscyphus antarcticus (C.Massal.) Solari, Cryptog. Bryol. Lichénol. 7 (3): 219, 1986 (Solari 1986). Bas.: Leioscyphus antarcticus C.Massal., Atti Reale Ist. Veneto Sci. Lett. Arti 87 (2): 229, 1928 (Massalongo 1928). *** Leptoscyphus australis (Gottsche, Lindenb. et Nees) R.M.Schust., J. Hattori Bot. Lab. 26: 270, 1963 (Schuster 1963b). Bas.: Chiloscyphus australis Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 189, 1845 (Gottsche et al. 1845a). *** Leptoscyphus belmoranus (Steph.) J.J.Engel, J. Hattori Bot. Lab. 74: 33, 1993 (Engel 1993). Bas.: Lophocolea belmorana Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 117, 1914 (Stephani and Watts 1914). *** Leptoscyphus excipulatus (Steph.) J.J.Engel, J. Hattori Bot. Lab. 74: 33, 1993 (Engel 1993). Bas.: Lophocolea excipulata Steph., Bull. Herb. Boissier (sér. 2) 6 (9): 790 (90), 1906 (Stephani 1906e). *** Leptoscyphus innovatus (E.A.Hodgs.) J.J.Engel, J. Hattori Bot. Lab. 74: 33, 1993 (Engel 1993). Bas.: Lophocolea innovata E.A.Hodgs., Trans. Roy. Soc. New Zealand 80 (3/4): 347, 1952 [1953] (Hodgson 1952). *** Leptoscyphus longistipulus (Steph.) J.J.Engel, Bryologist 94 (4): 436, 1991 (Engel 1991b). Bas.: Lophocolea longistipula Steph., Bull. Herb. Boissier (sér. 2) 6 (10): 884 (109), 1906 (Stephani 1906d). ** subg. Leptoscyphus ** sect. Hexagonistipa Grolle, Nova Acta Leop. (n.ser.) 25 (161): 46, 1962 (Grolle 1962a). *** Leptoscyphus gibbosus (Taylor) Mitt., Hooker’s J. Bot. Kew Gard. Misc. 3: 358, 1851 (Mitten 1851). Bas.: Chiloscyphus gibbosus Taylor, London J. Bot. 5: 283, 1846 (Taylor 1846a). *** Leptoscyphus gradsteinii Vanderp., Schäf.-Verw. et D.G.Long, Taxon 59 (1): 182, 2010 (Vanderpoorten et al. 2010). *** Leptoscyphus hexagonus (Nees) Grolle, Nova Acta Leop. (n.ser.) 25 (161): 47, 1962 [1963] (Grolle 1962a). Bas.: Chiloscyphus hexagonus Nees, Syn. Hepat. 2: 177, 1845 (Gottsche et al. 1845a). *** Leptoscyphus jackii (Steph.) Grolle, Nova Acta Leop. (n.ser.) 25 (161): 48, 1962 [1963] (Grolle 1962a). Bas.: Leioscyphus jackii Steph., Hedwigia 31 (1): 21, 1892 (Jack and Stephani 1892). *** Leptoscyphus physocalyx (Hampe et Gottsche) Gottsche, Bot. Zeitung (Berlin) Beil. 16: 33, 1858 (Gottsche 1858). Bas.: Jungermannia physocalyx Hampe et Gottsche, Linnaea 20 (3): 326, 1847 (Hampe 1847). 206 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Leptoscyphus sotiauxii Vanderp., Schäf.-Verw. et D.G.Long, Taxon 59 (1): 183, 2010 (Vanderpoorten et al. 2010). ** sect. Leptoscyphus *** Leptoscyphus aequatus (Hook.f. et Taylor) Mitt., Hooker’s J. Bot. Kew Gard. Misc. 3: 358, 1851 (Mitten 1851). Bas.: Jungermannia aequata Hook.f. et Taylor, London J. Bot. 3: 465, 1844 (Hooker and Taylor 1844b). *** Leptoscyphus intermedius Grolle, Nova Acta Leop. (n.ser.) 25 (161): 32, 1962 [1963] (Grolle 1962a). *** Leptoscyphus lambinonii Vanderp., Schäf.-Verw. et D.G.Long, Taxon 59 (1): 179, 2010 (Vanderpoorten et al. 2010). *** Leptoscyphus obcordatus (Spruce) Grolle, Nova Acta Leop. (n.ser.) 25 (161): 33, 1962 [1963] (Grolle 1962a). Bas.: Leioscyphus obcordatus Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 446, 1885 (Spruce 1885). *** Leptoscyphus ovatus (Spruce) Grolle, Nova Acta Leop. (n.ser.) 25 (161): 45, 1962 [1963] (Grolle 1962a). Bas.: Leioscyphus ovatus Spruce, J. Linn. Soc., Bot. 30 (210): 357, 1895 (Gepp 1895b). *** Leptoscyphus porphyrius (Nees) Grolle, Österr. Bot. Z. 117 (1): 3, 1969 (Grolle 1969a). Bas.: Chiloscyphus porphyrius Nees, Syn. Hepat. 2: 185, 1845 (Gottsche et al. 1845a). *** Leptoscyphus porphyrius subsp. azoricus (H.Buch et Perss.) Vanderp. et Heinrichs, Taxon 59 (1): 181, 2010 (Vanderpoorten et al. 2010). Bas.: Mylia azorica H.Buch et Perss., Bryophyt. Azoren Madeira: 7, 1941 (Buch and Persson 1941). ** subg. Physoscyphus Grolle, Nova Acta Leop. (n.ser.) 25 (161): 51, 1962 (Grolle 1962a). ** sect. Homaloscyphus Grolle, Nova Acta Leop. (n.ser.) 25 (161): 58, 1962 (Grolle 1962a). *** Leptoscyphus chilensis (De Not.) Hässel, J. Hattori Bot. Lab. 91: 207, 2001 (Hässel 2001). Bas.: Lophocolea chilensis De Not., Mem. Reale Accad. Sci. Torino (ser. 2) 16: 222, 1857 (De Notaris 1857). * Leptoscyphus diicilis (Steph.) Fulford, Mem. New York Bot. Gard. 11 (4): 534, 1976 (Fulford 1976). Bas.: Chiloscyphus diicilis Steph., Biblioth. Bot. 87 (2): 222, 1916 (Stephani 1916a). 146 *** Leptoscyphus diversifolius (Gottsche) Grolle, Nova Acta Leop. (n.ser.) 25 (161): 58, 1962 [1963] (Grolle 1962a). Bas.: Lophocolea diversifolia Gottsche, Syn. Hepat. 2: 166, 1845 (Gottsche et al. 1845a). *** Leptoscyphus expansus (Lehm.) Grolle, Nova Acta Leop. (n.ser.) 25 (161): 60, 1962 [1963] (Grolle 1962a). Bas.: Jungermannia expansa Lehm., Linnaea 4: 361, 1829 (Lehmann 1829). 146 Leptoscyphus diicilis is conspeciic with Leptoscyphus expansus in Grolle (1962a), but it was accepted by Vanderpoorten et al. (2010). World checklist of hornworts and liverworts 207 *** Leptoscyphus hedbergii (S.W.Arnell) R.M.Schust., Amer. Midl. Naturalist 62 (1): 13, 1959 (Schuster 1959a). Bas.: Mylia hedbergii S.W.Arnell, Ark. Bot. (n.ser.) 3 (16): 547, 1956 (Arnell 1956e). * Leptoscyphus huidobroanus (Mont.) Gottsche, Bot. Zeitung (Berlin) Beil. 16: 33, 1858 (Gottsche 1858). Bas.: Chiloscyphus huidobroanus Mont., Ann. Sci. Nat. Bot. (sér. 3) 4: 352, 1845 (Montagne 1845b). 147 *** Leptoscyphus huonicus Piippo, Acta Bot. Fenn. 131: 152, 1985 (Piippo 1985a). *** Leptoscyphus magellanicus (Gola) Hässel, J. Hattori Bot. Lab. 91: 214, 2001 (Hässel 2001). Bas.: Lophozia magellanica Gola, Nuovo Giorn. Bot. Ital. (n.ser.) 29 (1/4): 165, 1922 [1923] (Gola 1922). ** sect. Physoscyphus Grolle, Nova Acta Leop. (n.ser.) 25 (161): 51, 1962 (Grolle 1962a). *** Leptoscyphus amphibolius (Nees) Grolle, Nova Acta Leop. (n.ser.) 25 (161): 54, 1962 [1963] (Grolle 1962a). Bas.: Jungermannia amphibolia Nees, Fl. Bras. (Martius) 1 (1): 334, 1833 (Nees 1833a). *** Leptoscyphus infuscatus (Mitt.) E.W.Jones ex Grolle, Nova Acta Leop. (n.ser.) 25 (161): 52, 1962 [1963] (Grolle 1962a). Bas.: Leioscyphus infuscatus Mitt., J. Linn. Soc., Bot. 22 (146): 321, 1886 (Mitten 1886b). ** subg. Spinoscyphus Vanderp., Schäf.-Verw. et D.G.Long, Taxon 59 (1): 183, 2010 (Vanderpoorten et al. 2010). *** Leptoscyphus cleeii Fulford, Mem. New York Bot. Gard. 11 (4): 534, 1976 (Fulford 1976). *** Leptoscyphus spectabilis (Steph.) Grolle, J. Bryol. 11 (2): 328, 1980 [1981] (Grolle 1980c). Bas.: Lophocolea spectabilis Steph., Bull. Herb. Boissier (sér. 2) 7 (4): 302 (166), 1907 (Stephani 1907d). Incertae sedis *** Leptoscyphus autoicus (J.J.Engel et Gradst.) Vanderp. et Gradst., J. Bryol. 34 (4): 252, 2012 (Vanderpoorten et al. 2012). Bas.: Physotheca autoica J.J.Engel et Gradst., Taxon 52 (4): 764, 2003 (Engel and Gradstein 2003). ** Leptoscyphus beckettianus (Steph.) R.M.Schust. ex J.J.Engel, Nova Hedwigia 93 (3/4): 402, 2011 (Engel 2011). Bas.: Chiloscyphus beckettianus Steph., Bull. Herb. Boissier (sér. 2) 8 (1): 59 (235), 1908 (Stephani 1908k). ** Leptoscyphus compactus (Colenso) J.J.Engel, Nova Hedwigia 100 (3/4): 579, 2015 (Engel 2015a). Bas.: Chiloscyphus compactus Colenso, Trans. & Proc. New Zealand Inst. 21: 63, 1889 (Colenso 1889). 147 Leptoscyphus huidobroanus is conspeciic with Leptoscyphus expansus in Váňa and Engel (2013) rejecting the conclusion by Hässel (2001) that it is an independent species, but it was accepted by Vanderpoorten et al. (2010). 208 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Leptoscyphus erraticus (W.Martin et E.A.Hodgs.) J.J.Engel, Nova Hedwigia 99 (1/2): 168, 2014 (Engel 2014). Bas.: Chiloscyphus erraticus W.Martin et E.A.Hodgs., Trans. & Proc. Roy. Soc. New Zealand 78 (4): 497, 1950 (Martin 1950). *** Leptoscyphus horizontalis (Hook.) Kühnem., Revista Centro Estud. Doct. Ci. Nat. 1: 176, 1937 (Kühnemann 1937). Bas.: Jungermannia horizontalis Hook., Musci Exot. 1: tab. 96, 1818 (Hooker 1818). ** Leptoscyphus normalis (Steph.) J.J.Engel, Nova Hedwigia 100 (3/4): 579, 2015 (Engel 2015a). Bas.: Lophocolea normalis Steph., Sp. Hepat. (Stephani) 6: 285, 1922 (Stephani 1922). ** Leptoscyphus physanthus (Hook.f. et Taylor) J.J.Engel, Nova Hedwigia 99 (1/2): 168, 2014 (Engel 2014). Bas.: Jungermannia physantha Hook.f. et Taylor, London J. Bot. 3: 561, 1844 (Hooker and Taylor 1844d). ** Leptoscyphus subemarginatus (Hook.f. et Taylor) J.J.Engel, Bryologist 94 (4): 436, 1991 (Engel 1991b). Bas.: Lophocolea subemarginata Hook.f. et Taylor, London J. Bot. 5: 367, 1846 (Taylor 1846b). *** Leptoscyphus trapezoïdes (Mont.) L.Söderstr., Phytotaxa 112 (1): 27, 2013 (Söderström et al. 2013f). Bas.: Lophocolea trapezoïdes Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 251, 1843 (Montagne 1843). *** Lophocolea (Dumort.) Dumort., Recueil Observ. Jungerm.: 17, 1835 (Dumortier 1835). Bas.: Jungermannia sect. Lophocolea Dumort., Syll. Jungerm. Europ.: 59, 1831 (Dumortier 1831). *** Lophocolea aberrans Lindenb. et Gottsche, Syn. Hepat. 5: 696, 1847 (Gottsche et al. 1847). *** Lophocolea aequifolia Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 5: 55, 1836 (Nees and Montagne 1836). * Lophocolea angustistipula Steph., Sp. Hepat. (Stephani) 6: 260, 1922 (Stephani 1922). *** Lophocolea anisoloba (J.J.Engel et Glenny) L.Söderstr., Phytotaxa 112 (1): 25, 2013 (Söderström et al. 2013f). Bas.: Chiloscyphus anisolobus J.J.Engel et Glenny, Bryologist 111 (1): 118, 2008 (Engel and Glenny 2008b). ** Lophocolea anomala Steph., Sp. Hepat. (Stephani) 6: 300, 1922 (Stephani 1922). ** Lophocolea anomoda (Mont.) Steph., Hedwigia 32 (5): 327, 1893 (Stephani 1893d). Bas.: Chiloscyphus anomodus Mont., Ann. Sci. Nat. Bot. (sér. 3) 4: 352, 1845 (Montagne 1845b). ** Lophocolea apalachicola R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 195, 1980 (Schuster 1980c). *** Lophocolea aperticaulis (J.J.Engel) L.Söderstr., Phytotaxa 112 (1): 25, 2013 (Söderström et al. 2013f). Bas.: Chiloscyphus aperticaulis J.J.Engel, J. Hattori Bot. Lab. 95: 229, 2004 (Engel 2004b). ** Lophocolea aphelophylla (Hässel) Váňa, Phytotaxa 112 (1): 25, 2013 (Söderström et al. 2013f). Bas.: Chiloscyphus aphelophyllus Hässel, J. Hattori Bot. Lab. 98: 123, 2005 (Hässel 2005). World checklist of hornworts and liverworts 209 ** Lophocolea apophylla (Hässel) Váňa, Phytotaxa 112 (1): 25, 2013 (Söderström et al. 2013f). Bas.: Chiloscyphus apophyllus Hässel, J. Hattori Bot. Lab. 98: 126, 2005 (Hässel 2005). *** Lophocolea appalachiana R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 208, 1980 (Schuster 1980c). ** Lophocolea ascensionis Steph., Sp. Hepat. (Stephani) 6: 261, 1922 (Stephani 1922). ** Lophocolea asperrima Steph., Bull. Herb. Boissier (sér. 2) 6 (11): 949 (129), 1906 (Stephani 1906c). ** Lophocolea atra Gola, Nuovo Giorn. Bot. Ital. (n.ser.) 29 (1/4): 167, 1922 [1923] (Gola 1922). *** Lophocolea attenuata Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 6): 34, 1900 (Stephani 1900b). *** Lophocolea australis Gottsche, Linnaea 28 (5): 553, 1856 [1857] (Gottsche 1856). ** Lophocolea autoica Steph., Sp. Hepat. (Stephani) 6: 262, 1922 (Stephani 1922). ** Lophocolea baldwinii Steph., Bull. Herb. Boissier (sér. 2) 6 (11): 950 (130), 1906 (Stephani 1906c). ** Lophocolea bartlettii H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 506, 1963 (Miller 1963). ** Lophocolea bewsii (Sim) Grolle, Trans. Brit. Bryol. Soc. 3 (4): 588, 1959 (Grolle 1959b). Bas.: Leptoscyphus bewsii Sim, Trans. Roy. Soc. South Africa 15 (1): 103, 1926 (Sim 1926). ** Lophocolea bicuspidata Steph., Sp. Hepat. (Stephani) 6: 263, 1922 (Stephani 1922). *** Lophocolea bidentata (L.) Dumort., Recueil Observ. Jungerm.: 17, 1835 (Dumortier 1835). Bas.: Jungermannia bidentata L., Sp. Pl. 1: 1132, 1753 (Linnaeus 1753). 148 ** Lophocolea biidistipula Steph., Sp. Hepat. (Stephani) 6: 264, 1922 (Stephani 1922). *** Lophocolea bispinosa (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 162, 1845 (Gottsche et al. 1845a). Bas.: Jungermannia bispinosa Hook.f. et Taylor, London J. Bot. 3: 378, 1844 (Hooker and Taylor 1844a). ** Lophocolea bootanensis Steph., Sp. Hepat. (Stephani) 6: 265, 1922 (Stephani 1922). ** Lophocolea boulyana Steph., Sp. Hepat. (Stephani) 6: 264, 1922 (Stephani 1922). ** Lophocolea bowiena Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 117, 1914 (Stephani and Watts 1914). *** Lophocolea brookwoodiana Paton et Sheahan, J. Bryol. 28 (3): 163, 2006 (Paton and Sheahan 2006). ** Lophocolea caespitans Steph., Bull. Herb. Boissier (sér. 2) 6 (11): 949 (129), 1906 (Stephani 1906c). *** Lophocolea calcarea Steph., Bull. Herb. Boissier (sér. 2) 6 (10): 884 (109), 1906 (Stephani 1906d). ** Lophocolea caledonica Steph., Sp. Hepat. (Stephani) 6: 267, 1922 (Stephani 1922). 148 Lophocolea bidentata is a species complex as discussed by Váňa and Engel (2013) and it has a complicated nomenclatural history causing many misunderstandings. At least Lophocolea coadunata and Lophocolea humifusa also belong to the complex. 210 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Lophocolea canaliculata (Gottsche, Lindenb. et Nees) Steph., Bull. Herb. Boissier (sér. 2) 6 (9): 786 (86), 1906 (Stephani 1906e). Bas.: Chiloscyphus canaliculatus Gottsche, Lindenb. et Nees, Syn. Hepat. 5: 710, 1847 (Gottsche et al. 1847). *** Lophocolea canaliculata var. concava (J.J.Engel) L.Söderstr., Phytotaxa 112 (1): 26, 2013 (Söderström et al. 2013f). Bas.: Chiloscyphus canaliculatus var. concavus J.J.Engel, Fieldiana, Bot. (n.ser.) 48: 107, 2010 (Engel 2010). ** Lophocolea cervicornis Steph., Biblioth. Bot. 87 (2): 219, 1916 (Stephani 1916a). ** Lophocolea ciliifera Steph., Bull. Herb. Boissier (sér. 2) 6 (8): 660 (76), 1906 (Stephani 1906h). * Lophocolea coadunata (Sw.) Mont., Voy. Amér. Mérid., Bot. 7 (1): 76, 1839 (Montagne 1839b). Bas.: Jungermannia coadunata Sw., Fl. Ind. Occid. 3: 1850, 1806 (Swartz 1806). 149 ** Lophocolea concreta Mont., Ann. Sci. Nat. Bot. (sér. 3) 4: 350, 1845 (Montagne 1845b). ** Lophocolea convexula Mitt., Fl. vit.: 405, 1871 [1873] (Mitten 1871). ** Lophocolea corticola Steph., Sp. Hepat. (Stephani) 6: 268, 1922 (Stephani 1922). *** Lophocolea decurrens Herzog, Trans. & Proc. Roy. Soc. New Zealand 65 (3): 352, 1936 (Herzog 1936b). * Lophocolea deningeri Herzog, Beih. Bot. Centralbl. 38 (2): 321, 1921 (Herzog 1921). ** Lophocolea dentilora Steph., Bull. Herb. Boissier (sér. 2) 6 (7): 550 (64), 1906 (Stephani 1906f). ** Lophocolea diformis Nees, Syn. Hepat. 2: 166, 1845 (Gottsche et al. 1845a). ** Lophocolea discedens (Lehm. et Lindenb.) Lehm. et Lindenb., Syn. Hepat. 2: 167, 1845 (Gottsche et al. 1845a). Bas.: Jungermannia discedens Lehm. et Lindenb., Nov. Stirp. Pug. 5: 3, 1833 (Lehmann 1833). * Lophocolea dusenii Steph., Cat. Afr. Pl. (Hiern) 2 (2): 314, 1901 (Stephani 1901d). *** Lophocolea erosa (J.J.Engel) L.Söderstr., Phytotaxa 112 (1): 26, 2013 (Söderström et al. 2013f). Bas.: Chiloscyphus erosus J.J.Engel, Phytologia 83 (1): 43, 1997 [1998] (Engel 1997). *** Lophocolea excisifolia Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 118, 1914 (Stephani and Watts 1914). *** Lophocolea fertilis (J.J.Engel) L.Söderstr., Phytotaxa 112 (1): 26, 2013 (Söderström et al. 2013f). Bas.: Chiloscyphus fertilis J.J.Engel, Phytologia 83 (1): 43, 1997 [1998] (Engel 1997). ** Lophocolea lavicans Steph., Sp. Hepat. (Stephani) 6: 300, 1922 (Stephani 1922). *** Lophocolea loribunda Steph., Bull. Herb. Boissier (sér. 2) 6 (10): 886 (111), 1906 (Stephani 1906d). ** Lophocolea foliicola Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 428, 1885 (Spruce 1885). ** Lophocolea fragillima Steph., Sp. Hepat. (Stephani) 6: 273, 1922 (Stephani 1922). 149 Lophocolea coadunata belongs to the Lophocolea bidentata species complex with complicated nomenclature (Váňa and Engel 2013). World checklist of hornworts and liverworts 211 *** Lophocolea fragmentissima R.M.Schust., Phytologia 39 (4): 245, 1978 (Schuster 1978a). *** Lophocolea fragrans (Moris et De Not.) Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 166, 1845 (Gottsche et al. 1845a). Bas.: Jungermannia fragrans Moris et De Not., Fl. Caprariae: 177, 1839 (Moris and De Notaris 1839). ** Lophocolea glaziovii Steph., Bull. Herb. Boissier (sér. 2) 6 (11): 961 (141), 1906 (Stephani 1906c). ** Lophocolea gollanii (Steph.) Váňa, Phytotaxa 112 (1): 26, 2013 (Söderström et al. 2013f). Bas.: Chiloscyphus gollanii Steph., Bull. Herb. Boissier (sér. 2) 7 (10): 837 (209), 1907 (Stephani 1907b). ** Lophocolea granatensis Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 126, 1864 (Gottsche 1864). ** Lophocolea griithiana Steph., Sp. Hepat. (Stephani) 6: 274, 1922 (Stephani 1922). ** Lophocolea hahnii Steph., Bull. Herb. Boissier (sér. 2) 6 (8): 660 (76), 1906 (Stephani 1906h). ** Lophocolea haskarliana Gottsche, Syn. Hepat. 2: 153, 1845 (Gottsche et al. 1845a). *** Lophocolea hattorii (J.J.Engel) L.Söderstr., Phytotaxa 112 (1): 26, 2013 (Söderström et al. 2013f). Bas.: Chiloscyphus hattorii J.J.Engel, J. Hattori Bot. Lab. 74: 29, 1993 (Engel 1993). ** Lophocolea hawaica Steph., Bull. Herb. Boissier (sér. 2) 6 (11): 945 (125), 1906 (Stephani 1906c). ** Lophocolea heterodonta Steph., Sp. Hepat. (Stephani) 6: 275, 1922 (Stephani 1922). ** Lophocolea heteromorpha Steph., Sp. Hepat. (Stephani) 6: 275, 1922 (Stephani 1922). *** Lophocolea heterophylla (Schrad.) Dumort., Recueil Observ. Jungerm.: 17, 1835 (Dumortier 1835). Bas.: Jungermannia heterophylla Schrad., J. Bot. (Schrader) 5: 66, 1802 [1803] (Schrader 1802). ** Lophocolea heterophylla subsp. cladogyna R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 223, 1980 (Schuster 1980c). ** Lophocolea horikawana S.Hatt., Bull. Tokyo Sci. Mus. 11: 50, 1944 (Hattori 1944d). ** Lophocolea howeana Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 118, 1914 (Stephani and Watts 1914). * Lophocolea humifusa (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 5: 695, 1847 (Gottsche et al. 1847). Bas.: Jungermannia humifusa Hook.f. et Taylor, London J. Bot. 3: 472, 1844 (Hooker and Taylor 1844b). ** Lophocolea humistrata (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 5: 701, 1847 (Gottsche et al. 1847). Bas.: Jungermannia humistrata Hook.f. et Taylor, London J. Bot. 4: 82, 1845 (Hooker and Taylor 1845). ** Lophocolea itoana Inoue, J. Jap. Bot. 31 (11): 340, 1956 (Inoue 1956). ** Lophocolea javanica Schifn., Hep. Fl. Buitenzorg: 178, 1900 (Schifner 1900a). ** Lophocolea koponenii (Piippo) Váňa, Phytotaxa 112 (1): 26, 2013 (Söderström et al. 2013f). Bas.: Chiloscyphus koponenii Piippo, Ann. Bot. Fenn. 35 (1): 55, 1998 (Piippo 1998). 212 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Lophocolea kurzii Sande Lac., Ann. Mus. Bot. Lugduno-Batavi 1: 296, 1864 (Sande Lacoste 1864). * Lophocolea kurzii var. siamensis N.Kitag., Acta Phytotax. Geobot. 30 (1/3): 33, 1979 (Kitagawa 1979b). ** Lophocolea laceristipula Steph., Sp. Hepat. (Stephani) 6: 281, 1922 (Stephani 1922). ** Lophocolea latistipula Steph., Sp. Hepat. (Stephani) 6: 281, 1922 (Stephani 1922). *** Lophocolea lauterbachii Steph., Bull. Herb. Boissier (sér. 2) 6 (11): 938 (118), 1906 (Stephani 1906c). * Lophocolea laxissima Herzog, Ann. Bryol. 5: 77, 1932 (Herzog 1932b). ** Lophocolea ledermannii Steph., Sp. Hepat. (Stephani) 6: 300, 1922 (Stephani 1922). *** Lophocolea lenta (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 162, 1845 (Gottsche et al. 1845a). Bas.: Jungermannia lenta Hook.f. et Taylor, London J. Bot. 3: 379, 1844 (Hooker and Taylor 1844a). *** Lophocolea leptantha (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 5: 694, 1847 (Gottsche et al. 1847). Bas.: Jungermannia leptantha Hook.f. et Taylor, London J. Bot. 3: 471, 1844 (Hooker and Taylor 1844b). *** Lophocolea liebmanniana Gottsche, Mexik. Leverm.: 113, 1863 (Gottsche 1863). ** Lophocolea lindmannii Steph., Bull. Herb. Boissier (sér. 2) 6 (11): 960 (140), 1906 (Stephani 1906c). *** Lophocolea longiciliata Herzog, Memoranda Soc. Fauna Fl. Fennica 27: 96, 1952 (Herzog 1952c). ** Lophocolea lucida (Spreng.) Mont., Voy. Amér. Mérid., Bot. 7 (2): 78, 1839 (Montagne 1839a). Bas.: Jungermannia lucida Spreng. Nov. Stirp. Pug. 5: 2, 1833 (Lehmann 1833). ** Lophocolea madagascariensis Gottsche, Abh. Naturwiss. Vereins Bremen 7: 344, 1882 (Gottsche 1882). ** Lophocolea magna (Udar et V.Nath) Váňa, Phytotaxa 183 (4): 291, 2014 (Váňa et al. 2014a). Bas.: Cephaloziella magna Udar et V.Nath, Geophytology 6 (1): 105, 1976 (Udar and Nath 1976). *** Lophocolea mediinfrons (J.J.Engel et Braggins) L.Söderstr., Phytotaxa 112 (1): 26, 2013 (Söderström et al. 2013f). Bas.: Chiloscyphus mediinfrons J.J.Engel et Braggins, Fieldiana, Bot. (n.ser.) 48: 119, 2010 (Engel 2010). ** Lophocolea micronesica Inoue et H.A.Mill., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 11 (1): 4, 1968 (Inoue and Miller 1968). * Lophocolea microstipula Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 6): 43, 1900 (Stephani 1900b). 150 *** Lophocolea minor Nees, Naturgesch. Eur. Leberm. 2: 330, 1836 (Nees 1836). ** Lophocolea minutistipula Steph., Sp. Hepat. (Stephani) 6: 283, 1922 (Stephani 1922). * Lophocolea mollis (Nees) Nees, Syn. Hepat. 2: 158, 1845 (Gottsche et al. 1845a). Bas.: Jungermannia mollis Nees, Enum. Pl. Crypt. Javae: 24, 1830 (Nees 1830). 151 150 Lophocolea microstipula is possibly conspeciic with Lophocolea sabuletorum (Engel 1978), but it was accepted by Hässel and Rubies (2009). 151 Lophocolea mollis is possibly conspeciic with Lophocolea diformis (Grolle 1995). World checklist of hornworts and liverworts 213 * Lophocolea morobeana Piippo, Acta Bot. Fenn. 131: 160, 1985 (Piippo 1985a). 152 ** Lophocolea muhavurensis (S.W.Arnell) S.W.Arnell ex Pócs, Acta Bot. Acad. Sci. Hung. 25 (3/4): 227, 1979 [1980] (Bizot and Pócs 1979). Bas.: Chiloscyphus muhavurensis S.W.Arnell, Ark. Bot. (n.ser.) 3 (16): 526, 1956 (Arnell 1956e). *** Lophocolea muricata (Lehm.) Nees, Syn. Hepat. 2: 169, 1845 (Gottsche et al. 1845a). Bas.: Jungermannia muricata Lehm., Linnaea 4: 363, 1829 (Lehmann 1829). * Lophocolea muricata var. major Pearson, Forh. Vidensk.-Selsk. Kristiania 1892 (14): 10, 1893 (Pearson 1893). ** Lophocolea nakajimae S.Hatt. et Inoue, J. Hattori Bot. Lab. 21: 221, 1959 (Inoue 1959c). *** Lophocolea novae-zeelandiae (Lehm. et Lindenb.) Nees, Syn. Hepat. 2: 168, 1845 (Gottsche et al. 1845a). Bas.: Jungermannia novae-zeelandiae Lehm. et Lindenb., Nov. Stirp. Pug. 6: 33, 1834 (Lehmann 1834). *** Lophocolea novae-zeelandiae var. meridionalis (Steph.) L.Söderstr., Phytotaxa 112 (1): 26, 2013 (Söderström et al. 2013f). Bas.: Lophocolea meridionalis Steph., Bull. Herb. Boissier (sér. 2) 6 (10): 888 (113), 1906 (Stephani 1906d). ** Lophocolea orbigniana Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 5: 55, 1836 (Nees and Montagne 1836). ** Lophocolea papulimarginata H.A.Mill., Phytologia 47 (4): 323, 1981 (Miller 1981). Nom. nov. pro Lophocolea papulosa Steph., Sp. Hepat. (Stephani) 6: 286, 1922 (Stephani 1922), nom. illeg. ** Lophocolea parca (Gottsche) Fulford et Sharp, Mem. New York Bot. Gard. 63: 19, 1990 (Fulford and Sharp 1990). Bas.: Jungermannia parca Gottsche, Mexik. Leverm.: 94, 1863 (Gottsche 1863). ** Lophocolea parva Steph., Sp. Hepat. (Stephani) 6: 287, 1922 (Stephani 1922). *** Lophocolea parvispinea (J.J.Engel) L.Söderstr., Phytotaxa 112 (1): 26, 2013 (Söderström et al. 2013f). Bas.: Chiloscyphus parvispineus J.J.Engel, Phytologia 83 (1): 44, 1997 [1998] (Engel 1997). ** Lophocolea parvistipula Steph., Sp. Hepat. (Stephani) 6: 287, 1922 (Stephani 1922). *** Lophocolea patulistipa Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 50, 1911 (Stephani 1911b). *** Lophocolea perpusilla (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 163, 1845 (Gottsche et al. 1845a). Bas.: Jungermannia perpusilla Hook.f. et Taylor, London J. Bot. 3: 380, 1844 (Hooker and Taylor 1844a). ** Lophocolea piacenzai (Gola) Váňa, Phytotaxa 112 (1): 26, 2013 (Söderström et al. 2013f). Bas.: Lophozia piacenzai Gola, Atti Reale Accad. Sci. Torino, Cl. Sci. Fis. Mat. Nat. 49: 759, 1914 (Gola 1914b). ** Lophocolea pilistipula Steph., Sp. Hepat. (Stephani) 6: 288, 1922 (Stephani 1922). * Lophocolea pinnatistipula Steph., Biblioth. Bot. 87 (2): 220, 1916 (Stephani 1916a). 152 Lophocolea morobeana is conspeciic with Chiloscyphus kurzii in Piippo (1987), but it was accepted as very close to Chiloscyphus kurzii by Ariyanti et al. (2009). 214 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Lophocolea platensis C.Massal., Atti Accad. Sci. Med. Nat. Ferrara 80 (3/4): 12, 1906 (Massalongo 1906a). ** Lophocolea purpurea Steph., Sp. Hepat. (Stephani) 6: 289, 1922 (Stephani 1922). ** Lophocolea pusilla Steph., Sp. Hepat. (Stephani) 6: 290, 1922 (Stephani 1922). ** Lophocolea randii S.W.Arnell, Svensk Bot. Tidskr. 47 (3): 420, 1953 (Arnell 1953c). ** Lophocolea rara Steph., Sp. Hepat. (Stephani) 6: 290, 1922 (Stephani 1922). ** Lophocolea rectangularis Herzog, Rev. Bryol. Lichénol. 23 (1/2): 43, 1954 (Herzog 1954). ** Lophocolea rectangulata Mitt., Fl. vit.: 404, 1871 [1873] (Mitten 1871). 153 *** Lophocolea rupicola Steph., Bull. Herb. Boissier (sér. 2) 6 (10): 874 (99), 1906 (Stephani 1906d). *** Lophocolea sabuletorum (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 5: 697, 1847 (Gottsche et al. 1847). Bas.: Jungermannia sabuletorum Hook.f. et Taylor, London J. Bot. 3: 469, 1844 (Hooker and Taylor 1844b). ** Lophocolea salacensis Steph., Bull. Herb. Boissier (sér. 2) 6 (11): 943 (123), 1906 (Stephani 1906c). ** Lophocolea savesiana Steph., Bull. Herb. Boissier (sér. 2) 6 (11): 942 (122), 1906 (Stephani 1906c). *** Lophocolea semiteres (Lehm.) Mitt., J. Linn. Soc., Bot. 16 (91): 188, 1877 (Mitten 1877). Bas.: Jungermannia semiteres Lehm., Linnaea 4: 363, 1829 (Lehmann 1829). 154 *** Lophocolea semiteres var. retusa (J.J.Engel) L.Söderstr., Phytotaxa 112 (1): 27, 2013 (Söderström et al. 2013f). Bas.: Chiloscyphus semiteres var. retusus J.J.Engel, Phytologia 83 (1): 44, 1997 [1998] (Engel 1997). ** Lophocolea serrata Mitt., St. Helena: 368, 1875 (Mitten 1875). ** Lophocolea siamensis Steph., Sp. Hepat. (Stephani) 6: 293, 1922 (Stephani 1922). ** Lophocolea sikkimensis (Steph.) Herzog et Grolle, Rev. Bryol. Lichénol. 27 (3/4): 164, 1958 [1959] (Herzog and Grolle 1958). Bas.: Herpocladium sikkimense Steph., Sp. Hepat. (Stephani) 6: 349, 1922 (Stephani 1922). * Lophocolea silvestris Gottsche, Abh. Naturwiss. Vereins Bremen 7: 345, 1882 (Gottsche 1882). ** Lophocolea steetziae De Not., Epat. Borneo: 20, 1874 (De Notaris 1874). *** Lophocolea striatella (C.Massal.) Schifn., Leberm., Forschungsr. Gazelle 4 (4): 13, 1890 (Schifner 1890). Bas.: Chiloscyphus striatellus C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 232, 1885 (Massalongo 1885). ** Lophocolea subbidentata Herzog, Rev. Bryol. Lichénol. 23 (1/2): 43, 1954 (Herzog 1954). ** Lophocolea subcostata Steph., Sp. Hepat. (Stephani) 6: 295, 1922 (Stephani 1922). *** Lophocolea subporosa Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 137, 1854 (Mitten 1854). 153 Lophocolea rectangulata may be a Cryptolophocolea species. 154 Lophocolea semiteres is a species complex also including Lophocolea platensis and Lophocolea undulata. World checklist of hornworts and liverworts 215 ** Lophocolea subporosa var. inlexifolia (Steph.) L.Söderstr., Phytotaxa 112 (1): 27, 2013 (Söderström et al. 2013f). Bas.: Lophocolea inlexifolia Steph., Sp. Hepat. (Stephani) 6: 278, 1922 (Stephani 1922). ** Lophocolea subviridis (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 5: 699, 1847 (Gottsche et al. 1847). Bas.: Jungermannia subviridis Hook.f. et Taylor, London J. Bot. 3: 473, 1844 (Hooker and Taylor 1844b). ** Lophocolea sumatrana Schifn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 70: 195, 1900 [1901] (Schifner 1900c). *** Lophocolea sylvatica Mitt., Rep. Challenger, Bot. 1 (3, 1): 84, 1884 (Mitten 1884b). ** Lophocolea tenera Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 33 (7): 79, 1876 [1877] (Ångström 1876). ** Lophocolea tenerrima Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 439, 1885 (Spruce 1885). ** Lophocolea teptepensis Piippo, Acta Bot. Fenn. 131: 163, 1985 (Piippo 1985a). *** Lophocolea textilis (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 5: 696, 1847 (Gottsche et al. 1847). Bas.: Jungermannia textilis Hook.f. et Taylor, London J. Bot. 3: 468, 1844 (Hooker and Taylor 1844b). ** Lophocolea textiloidea J.J.Engel, Phytologia 41 (5): 311, 1979 (Engel 1979a). Nom. nov. pro Chiloscyphus lucidus Mitt., J. Linn. Soc., Bot. 15 (82): 64, 1876 (Mitten 1876a), nom. illeg. *** Lophocolea trichocoleoides (Glenny, J.J.Engel et He-Nygrén) L.Söderstr., Phytotaxa 112 (1): 27, 2013 (Söderström et al. 2013f). Bas.: Chiloscyphus trichocoleoides Glenny, J.J.Engel et He-Nygrén, J. Bryol. 31 (2): 100, 2009 (Glenny et al. 2009). ** Lophocolea tricuspidata Herzog, Rev. Bryol. Lichénol. 11 (1): 17, 1938 [1939] (Herzog 1938a). *** Lophocolea tristaniana S.W.Arnell, Results Norweg. Sci. Exped. Tristan da Cunha 42: 19, 1958 (Arnell 1958b). * Lophocolea undulata Mont., Ann. Sci. Nat. Bot. (sér. 3) 4: 351, 1845 (Montagne 1845b). *** Lophocolea villosa Mitt., Sp. Hepat. (Stephani) 6: 299, 1922 (Stephani 1922). ** Lophocolea wacei (S.W.Arnell ex J.J.Engel et Váňa) Váňa et L.Söderstr., Phytotaxa 112 (1): 27, 2013 (Söderström et al. 2013f). Bas.: Chiloscyphus wacei S.W.Arnell ex J.J.Engel et Váňa, Mem. New York Bot. Gard. 105: 48, 2013 (Váňa and Engel 2013). *** Lophocolea wambana Piippo, Acta Bot. Fenn. 131: 163, 1985 (Piippo 1985a). ** Lophocolea werthii (J.J.Engel et R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 112 (1): 27, 2013 (Söderström et al. 2013f). Bas.: Chiloscyphus werthii J.J.Engel et R.M.Schust., Nova Hedwigia 39: 425, 1984 [1985] (Engel and Schuster 1984). ** Lophocolea widgrenii Steph., Bull. Herb. Boissier (sér. 2) 7 (1): 66 (154), 1907 (Stephani 1907c). Nom. nov. pro Lophocolea pallida Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 33 (7): 80, 1876 [1877] (Ångström 1876), nom. illeg. ** Otoscyphus J.J.Engel, Bardat et houvenot, Cryptog. Bryol. 33 (3): 280, 2012 (Engel et al. 2012). 216 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Otoscyphus crassicaulis (Steph.) J.J.Engel, Bardat et houvenot, Cryptog. Bryol. 33 (3): 280, 2012 (Engel et al. 2012). Bas.: Lophocolea crassicaulis Steph., Sp. Hepat. (Stephani) 6: 268, 1922 (Stephani 1922). *** Pachyglossa Herzog et Grolle, Rev. Bryol. Lichénol. 27 (3/4): 150, 1958 [1959] (Herzog and Grolle 1958). *** Pachyglossa austrigena (Hook.f. et Taylor) L.Söderstr., Phytotaxa 112 (1): 24, 2013 (Söderström et al. 2013f). Bas.: Jungermannia austrigena Hook.f. et Taylor, London J. Bot. 3: 466, 1844 (Hooker and Taylor 1844b). *** Pachyglossa austrigena subsp. okaritana (Steph.) L.Söderstr., Phytotaxa 112 (1): 24, 2013 (Söderström et al. 2013f). Bas.: Lophocolea okaritana Steph., Bull. Herb. Boissier (sér. 2) 6 (9): 785 (85), 1906 (Stephani 1906e). *** Pachyglossa boveana (C.Massal.) L.Söderstr., Phytotaxa 112 (1): 24, 2013 (Söderström et al. 2013f). Bas.: Lophocolea boveana C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 225, 1885 (Massalongo 1885). ** Pachyglossa dissitifolia Herzog et Grolle, Rev. Bryol. Lichénol. 27 (3/4): 155, 1958 [1959] (Herzog and Grolle 1958). * Pachyglossa exilis (Herzog et Grolle) Hässel et Solari, Transecta botánica de la Patagonia austral: 324, 1985 (Hässel and Solari 1985). Bas.: Pachyglossa spegazziniana var. exilis Herzog et Grolle, Rev. Bryol. Lichénol. 27 (3/4): 159, 1958 [1959] (Herzog and Grolle 1958). 155 ** Pachyglossa issa (Mitt.) Herzog et Grolle, Rev. Bryol. Lichénol. 28 (3/4): 346, 1959 [1960] (Grolle 1959c). Bas.: Herpocladium issum Mitt., J. Linn. Soc., Bot. 15 (82): 69, 1876 (Mitten 1876a). *** Pachyglossa gottscheoides (Besch. et C.Massal.) L.Söderstr., Phytotaxa 112 (1): 25, 2013 (Söderström et al. 2013f). Bas.: Lophocolea gottscheoides Besch. et C.Massal., Bull. Mens. Soc. Linn. Paris 1 (79): 631, 1886 (Bescherelle and Massalongo 1886). ** Pachyglossa grolleana Váňa, Cryptog. Bryol. 26 (1): 86, 2005 (Váňa and Gremmen 2005). ** Pachyglossa otiphylla (Hook.f. et Taylor) Váňa, Phytotaxa 112 (1): 25, 2013 (Söderström et al. 2013f). Bas.: Jungermannia otiphylla Hook.f. et Taylor, London J. Bot. 3: 466, 1844 (Hooker and Taylor 1844b). ** Pachyglossa spegazziniana (C.Massal.) Herzog et Grolle, Rev. Bryol. Lichénol. 27 (3/4): 159, 1958 [1959] (Herzog and Grolle 1958). Bas.: Lophocolea spegazziniana C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 225, 1885 (Massalongo 1885). ** Pachyglossa tenacifolia (Hook.f. et Taylor) Herzog et Grolle, Rev. Bryol. Lichénol. 27 (3/4): 153, 1958 [1959] (Herzog and Grolle 1958). Bas.: Jungermannia tenacifolia Hook.f. et Taylor, Bot. Antarct. Voy. I (Fl. Antarct. 1): 152, 1845 (Taylor and Hooker 1845). ** Perdusenia Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 7 (2): 11, 1989 (Hässel 1989b). 155 Pachyglossa exilis may be a Chiloscyphus species. World checklist of hornworts and liverworts 217 ** Perdusenia rheophila Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 7 (2): 11, 1989 (Hässel 1989b). ** Pigafettoa C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 237, 1885 (Massalongo 1885). ** Pigafettoa crenulata C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 237, 1885 (Massalongo 1885). ** Platycaulis R.M.Schust., Phytologia 39 (4): 245, 1978 (Schuster 1978a). *** Platycaulis renifolius R.M.Schust., Phytologia 39 (4): 245, 1978 (Schuster 1978a). *** Stolonivector J.J.Engel, J. Hattori Bot. Lab. 69: 80, 1991 (Engel 1991a). ** Stolonivector clasmatocoleoides J.J.Engel, Nova Hedwigia 88 (3/4): 339, 2009 (Engel 2009). *** Stolonivector iordlandiae (E.A.Hodgs.) J.J.Engel, J. Hattori Bot. Lab. 69: 82, 1991 (Engel 1991a). Bas.: Lophocolea iordlandiae E.A.Hodgs., Trans. Roy. Soc. New Zealand 80 (3/4): 340, 1952 [1953] (Hodgson 1952). ** Stolonivector iordlandiae var. nodulosus J.J.Engel, Nova Hedwigia 93 (3/4): 403, 2011 (Engel 2011). *** Stolonivector gremmenii (Váňa) Váňa, Phytotaxa 112 (1): 28, 2013 (Söderström et al. 2013f). Bas.: Chiloscyphus gremmenii Váňa, Cryptog. Bryol. 26 (1): 81, 2005 (Váňa and Gremmen 2005). ** Stolonivector obtusilobus J.J.Engel, Nova Hedwigia 88 (3/4): 337, 2009 (Engel 2009). ** Stolonivector waipouensis J.J.Engel, J. Hattori Bot. Lab. 93: 70, 2003 (Engel 2003). ** Xenocephalozia R.M.Schust., Nova Hedwigia 10 (1/2): 25, 1965 (Schuster 1965b). ** Xenocephalozia navicularis (Steph.) R.M.Schust., Nova Hedwigia 10 (1/2): 25, 1965 (Schuster 1965b). Bas.: Lophocolea navicularis Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 6): 43, 1900 (Stephani 1900b). *** Mastigophoraceae R.M.Schust. ** Dendromastigophora R.M.Schust., Mem. New York Bot. Gard. 45: 738, 1987 (Schuster 1987a). *** Dendromastigophora lagellifera (Hook.) R.M.Schust., Mem. New York Bot. Gard. 45: 738, 1987 (Schuster 1987a). Bas.: Jungermannia lagellifera Hook., Musci Exot. 1: tab. 59, 1818 (Hooker 1818). *** Mastigophora Nees, Naturgesch. Eur. Leberm. 3: 89, 1838 (Nees 1838b) nom. conserv. 156 156 Mastigophora includes (Mastigophora appendiculata, Mastigophora guineensis, Mastigophora pyramidana and Mastigophora valida) which Grolle and Piippo (1984) could not study since the types were destroyed in B. 218 * * ** *** ** ** * * * * ** ** * ** *** *** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Mastigophora appendiculata Steph., Sp. Hepat. (Stephani) 6: 368, 1922 (Stephani 1922). Mastigophora attenuata (Taylor) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 416, 1877 (Trevisan 1877). Bas.: Lepidozia attenuata Taylor, London J. Bot. 5: 369, 1846 (Taylor 1846b). Mastigophora caledonica Steph., Rev. Bryol. 35 (2): 31, 1908 (Stephani 1908l). Mastigophora diclados (Brid. ex F.Weber) Nees, Naturgesch. Eur. Leberm. 3: 18, 1838 (Nees 1838b). Bas.: Jungermannia diclados Brid. ex F.Weber, Hist. Musc. Hepat. Prodr.: 56, 1815 (Weber 1815). Mastigophora diclados var. borneensis (De Not.) Schifn., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 60 (2): 251, 1893 (Schifner 1893a). Bas.: Sendtnera diclados var. borneensis De Not., Epat. Borneo: 42, 1874 (De Notaris 1874). Mastigophora diclados var. ramentiissa Herzog, Trans. Brit. Bryol. Soc. 1 (4): 315, 1950 (Herzog 1950a). Mastigophora diclados var. villosa Herzog, Ann. Bryol. 5: 82, 1932 (Herzog 1932b). Mastigophora guineensis Steph., Sp. Hepat. (Stephani) 6: 369, 1923 (Stephani 1923). Mastigophora humillima (Taylor) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 416, 1877 (Trevisan 1877). Bas.: Lepidozia humillima Taylor, London J. Bot. 5: 369, 1846 (Taylor 1846b). Mastigophora pyramidana Steph., Sp. Hepat. (Stephani) 6: 369, 1923 (Stephani 1923). Mastigophora sepikiana Piippo, Ann. Bot. Fenn. 23 (1): 2, 1986 (Piippo 1986b). Mastigophora tuberculata D.H.Mill. et H.A.Mill., J. Hattori Bot. Lab. 75: 181, 1994 (Miller and Miller 1994). Mastigophora valida Steph., Sp. Hepat. (Stephani) 6: 369, 1923 (Stephani 1923). Mastigophora viridula (Nees) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 416, 1877 (Trevisan 1877). Bas.: Jungermannia viridula Nees, Flora 6 (2): 30, 1823 (Link 1823). Mastigophora woodsii (Hook.) Nees, Naturgesch. Eur. Leberm. 3: 95, 1838 (Nees 1838b). Bas.: Jungermannia woodsii Hook., Brit. Jungermann.: tab. 66, 1814 (Hooker 1814). Plagiochilaceae Müll.Frib. by L. Söderström he placement of Pedinophyllopsis in Plagiochilaceae follows He-Nygrén and Piippo (2003). he inclusion of Pseudolophocolea in the family follows Söderström et al. (2013b). he subgeneric division of Plagiochila follows the review by Söderström et al. (2015b). ** Acrochila R.M.Schust., J. Hattori Bot. Lab. 26: 285, 1963 (Schuster 1963b). World checklist of hornworts and liverworts 219 *** Acrochila biserialis (Lehm. et Lindenb.) Grolle, J. Jap. Bot. 39 (8): 236, 1964 (Grolle 1964g). Bas.: Plagiochila biserialis Lehm. et Lindenb., Sp. Hepat. (Lindenberg) 5: 126, 1843 (Lindenberg 1843). ** Acrochila caledonica (Steph.) Inoue, J. Jap. Bot. 42 (6): 182, 1967 (Inoue 1967d). Bas.: Plagiochila caledonica Steph., Rev. Bryol. 35 (2): 32, 1908 (Stephani 1908l). ** Chiastocaulon Carl, Flora 126: 58, 1931 (Carl 1931a). *** Chiastocaulon dendroides (Nees) Carl, Flora 126: 59, 1931 (Carl 1931a). Bas.: Jungermannia dendroides Nees, Enum. Pl. Crypt. Javae: 77, 1830 (Nees 1830). ** Dinckleria Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 421, 1877 (Trevisan 1877). *** Dinckleria fruticella (Hook.f. et Taylor) J.J.Engel et Heinrichs, Cryptog. Bryol. 29 (2): 194, 2008 (Engel and Heinrichs 2008). Bas.: Jungermannia fruticella Hook.f. et Taylor, London J. Bot. 3: 565, 1844 (Hooker and Taylor 1844d). *** Dinckleria pleurata (Hook.f. et Taylor) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 421, 1877 (Trevisan 1877). Bas.: Jungermannia pleurata Hook.f. et Taylor, London J. Bot. 3: 372, 1844 (Hooker and Taylor 1844a). ** Pedinophyllopsis R.M.Schust. et Inoue, Phytologia 47 (4): 311, 1981 (Schuster and Engel 1981). ** Pedinophyllopsis abdita (Sull.) R.M.Schust. et Inoue, Phytologia 47 (4): 311, 1981 (Schuster and Engel 1981). Bas.: Plagiochila abdita Sull., Hooker’s J. Bot. Kew Gard. Misc. 2: 317, 1850 (Sullivant 1850). ** Pedinophyllum Lindb. ex Nordst., Bot. Not. 1874: 156, 1874 (Nordstedt 1874). *** Pedinophyllum autoicum (Steph.) Inoue, Bull. Natl. Sci. Mus. Tokyo (n.ser.) 9 (4): 575, 1966 (Inoue 1966b). Bas.: Plagiochila autoica Steph., Sp. Hepat. (Stephani) 6: 126, 1917 (Stephani 1917a). *** Pedinophyllum interruptum (Nees) Kaal., Nyt Mag. Naturvidensk. 33 (1): 190, 1893 (Kaalaas 1893a). Bas.: Jungermannia interrupta Nees, Naturgesch. Eur. Leberm. 1: 165, 1833 (Nees 1833c). *** Pedinophyllum monoicum (Steph.) Grolle, Nova Hedwigia 2: 287, 1960 (Grolle 1960b). Bas.: Plagiochila monoica Steph., Bull. Herb. Boissier (sér. 2) 3 (4): 331 (315), 1903 (Stephani 1903c). ** Pedinophyllum truncatum (Steph.) Inoue, J. Hattori Bot. Lab. 23: 35, 1960 [1961] (Inoue 1960). Bas.: Clasmatocolea truncata Steph., Bull. Herb. Boissier 5 (2): 87, 1897 (Stephani 1897b). *** Plagiochila (Dumort.) Dumort., Recueil Observ. Jungerm.: 14, 1835 (Dumortier 1835) nom. conserv. Bas.: Radula sect. Plagiochila Dumort., Syll. Jungerm. Europ.: 42, 1831 (Dumortier 1831). 220 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Plagiochila heteromalla (Lehm. et Lindenb.) Lindenb., Sp. Hepat. (Lindenberg) 2-4: 83, 1840 (Lindenberg 1840). Bas.: Jungermannia heteromalla Lehm. et Lindenb., Nov. Stirp. Pug. 6: 62, 1834 (Lehmann 1834). *** sect. Adianthoideae Lindenb., Monogr. hep. gen. Plagiochilae: xx, 1844 (Lindenberg 1844). *** Plagiochila adianthoides (Sw.) Lindenb., Sp. Hepat. (Lindenberg) 2-4: 77, 1840 (Lindenberg 1840). Bas.: Jungermannia adianthoides Sw., Prodr. (Swartz): 142, 1788 (Swartz 1788). * Plagiochila adianthoides var. aspergillifera Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 474, 1885 (Spruce 1885). *** Plagiochila cristata (Sw.) Lindenb., Sp. Hepat. (Lindenberg) 1: 33, 1839 (Lindenberg 1839). Bas.: Jungermannia cristata Sw., Prodr. (Swartz): 143, 1788 (Swartz 1788). *** Plagiochila grandicrista Steph., Bull. Herb. Boissier (sér. 2) 5 (10): 931 (581), 1905 (Stephani 1905f). *** Plagiochila herminieri Steph., Bull. Herb. Boissier (sér. 2) 5 (8): 748 (547), 1905 (Stephani 1905i). *** sect. Africanae Heinrichs, Taxon 54 (2): 319, 2005 (Heinrichs et al. 2005). *** Plagiochila barteri Mitt., J. Linn. Soc., Bot. 22 (146): 320, 1886 (Mitten 1886b). ** Plagiochila barteri var. valida (Steph.) Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 51 (1/2): 73, 1981 (Vanden Berghen 1981). Bas.: Plagiochila valida Steph., Bull. Herb. Boissier (sér. 2) 4 (6): 587 (438), 1904 (Stephani 1904b). *** Plagiochila colorans Steph., Wiss. Ergebn. Deut. Zentr.-Afr. Exped. (1907-08), Bot. 2: 116, 1911 (Stephani 1911a). *** sect. Arrectae Carl, Ann. Bryol., Suppl. 2: 52, 1931 (Carl 1931b). * Plagiochila arnelliana Steph., Bull. Herb. Boissier (sér. 2) 2 (10): 861 (230), 1902 (Stephani 1902g). 157 *** Plagiochila badia Mitt., Rep. Challenger, Bot. 1 (3, 1): 84, 1884 (Mitten 1884b). *** Plagiochila bidens Gottsche, Ann. Sci. Nat. Bot. (sér. 4) 8: 322, 1857 (Gottsche 1857). *** Plagiochila bifaria (Sw.) Lindenb., Sp. Hepat. (Lindenberg) 5: 127, 1843 (Lindenberg 1843). Bas.: Jungermannia bifaria Sw., Prodr. (Swartz): 145, 1788 (Swartz 1788). ** Plagiochila bifaria var. rosea (R.M.Schust.) Heinrichs, Org. Divers. Evol. 4 (1/2): 112, 2004 (Heinrichs et al. 2004). Bas.: Rhodoplagiochila rosea R.M.Schust., Phytologia 39 (4): 247, 1978 (Schuster 1978a). *** Plagiochila chacabucensis Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 27, 1911 (Stephani 1911b). *** Plagiochila emeiensis Grolle et M.L.So, Bryologist 101 (2): 282, 1998 (Grolle and So 1998a). 157 Plagiochila arnelliana may be conspeciic with Plagiochila bifaria (Heinrichs et al. 1998). World checklist of hornworts and liverworts * *** *** * ** *** *** *** *** *** *** *** *** ** *** *** *** *** 221 Plagiochila fragilis Taylor, London J. Bot. 7: 198, 1848 (Taylor 1848a). 158 Plagiochila lunata S.W.Arnell, Bot. Not. 115: 204, 1962 (Arnell 1962a). Plagiochila pachyloma Taylor, London J. Bot. 5: 267, 1846 (Taylor 1846a). Plagiochila pachyloma var. elatior Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 480, 1885 (Spruce 1885). Plagiochila papillifolia Steph., Biblioth. Bot. 87 (2): 207, 1916 (Stephani 1916a). Plagiochila parviramifera Inoue, J. Hattori Bot. Lab. 46: 317, 1979 (Mizutani 1979a). Plagiochila pseudoattenuata S.W.Arnell, Bot. Not. 115: 206, 1962 (Arnell 1962a). Plagiochila punctata (Taylor) Taylor, London J. Bot. 5: 261, 1846 (Taylor 1846a). Bas.: Jungermannia punctata Taylor, Trans. Bot. Soc. Edinburgh 1: 179, 1844 (Taylor 1844b). Plagiochila renauldii Steph., Bull. Herb. Boissier (sér. 2) 4 (2): 156 (408), 1904 (Stephani 1904f). Plagiochila retrorsa Gottsche, Mexik. Leverm.: 67, 1863 (Gottsche 1863). Plagiochila rubescens (Lehm. et Lindenb.) Lindenb., Sp. Hepat. (Lindenberg) 2-4: 46, 1840 (Lindenberg 1840). Bas.: Jungermannia rubescens Lehm. et Lindenb., Nov. Stirp. Pug. 6: 63, 1834 (Lehmann 1834). Plagiochila sichuanensis Grolle et M.L.So, Bryologist 101 (2): 284, 1998 (Grolle and So 1998a). Plagiochila spinulosa (Dicks.) Dumort., Recueil Observ. Jungerm.: 15, 1835 (Dumortier 1835). Bas.: Jungermannia spinulosa Dicks., Fasc. Pl. Crypt. Brit. 2: 14, 1790 (Dickson 1790). Plagiochila sticticola Mont. et Gottsche, Ann. Sci. Nat. Bot. (sér. 4) 6: 198, 1856 (Montagne 1856a). Plagiochila stricta Lindenb., Sp. Hepat. (Lindenberg) 1: 20, 1839 (Lindenberg 1839). Plagiochila tronadoris Herzog, Darwiniana 11 (2): 214, 1957 (Herzog 1957b). Plagiochila uncialis (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 5: 628, 1847 (Gottsche et al. 1847). Bas.: Jungermannia uncialis Hook.f. et Taylor, London J. Bot. 3: 459, 1844 (Hooker and Taylor 1844b). Plagiochila wilmsiana Steph., Sp. Hepat. (Stephani) 6: 240, 1921 (Stephani 1921). sect. Caducifoliae J.J.Engel et G.L.Merr., Nova Hedwigia 96 (3/4): 407, 2013 (Engel and Smith Merrill 2013). ** Plagiochila caducifolia Inoue et R.M.Schust., J. Hattori Bot. Lab. 34: 71, 1971 (Inoue and Schuster 1971). * sect. Cardotiae Inoue, Bull. Natl. Sci. Mus. Tokyo (n.ser.) 8 (3): 386, 1965 (Inoue 1965a). *** Plagiochila cumingiana Steph., Bull. Herb. Boissier (sér. 2) 4 (1): 32 (404), 1904 (Stephani 1904g). * 158 Plagiochila fragilis may be conspeciic with Plagiochila bifaria (Heinrichs et al. 1998). 222 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Plagiochila denticulata Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 95, 1860 [1861] (Mitten 1860c). *** Plagiochila fragillima Steph., Bull. Herb. Boissier (sér. 2) 3 (6): 522 (326), 1903 (Stephani 1903d). *** Plagiochila pseudorenitens Schifn., Österr. Bot. Z. 49 (4): 132, 1899 (Schifner 1899b). *** Plagiochila stevensiana Steph., Bull. Herb. Boissier (sér. 2) 3 (2): 110 (290), 1903 (Stephani 1903b). * sect. Cobanae Carl, Ann. Bryol., Suppl. 2: 79, 1931 (Carl 1931b). ** Plagiochila cobana Steph., Sp. Hepat. (Stephani) 6: 138, 1918 (Stephani 1918). ** Plagiochila detecta M.L.So et Grolle, Nova Hedwigia 71 (3/4): 391, 2000 (So and Grolle 2000b). *** Plagiochila singularis Schifn., Hep. Fl. Buitenzorg: 158, 1900 (Schifner 1900a). *** Plagiochila tagawae Inoue, J. Hattori Bot. Lab. 38: 561, 1974 (Inoue 1974a). ** Plagiochila tixieri Inoue, Bull. Natl. Sci. Mus. Tokyo, B 1 (3): 87, 1975 (Inoue 1975a). *** Plagiochila zhuensis Grolle et M.L.So, Bryologist 102 (2): 200, 1999 (Grolle and So 1999c). *** sect. Cucullatae Schifn., Hep. Fl. Buitenzorg: 107, 1900 (Schifner 1900a). *** Plagiochila bantamensis (Reinw., Blume et Nees) Mont., Voy. Amér. Mérid., Bot. 7 (2): 82, 1839 (Montagne 1839a). Bas.: Jungermannia bantamensis Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 235, 1824 [1825] (Reinwardt et al. 1824a). *** Plagiochila blepharophora (Nees) Lindenb., Sp. Hepat. (Lindenberg) 2-4: 102, 1840 (Lindenberg 1840). Bas.: Jungermannia blepharophora Nees, Enum. Pl. Crypt. Javae: 71, 1830 (Nees 1830). ** Plagiochila chauviniana Mont., Ann. Sci. Nat. Bot. (sér. 3) 11: 34, 1849 (Montagne 1849). ** Plagiochila clavatosaccata Steph., Bull. Herb. Boissier (sér. 2) 4 (1): 25 (397), 1904 (Stephani 1904g). ** Plagiochila grossispina Steph., Sp. Hepat. (Stephani) 6: 162, 1918 (Stephani 1918). *** Plagiochila integerrima Steph., Bot. Jahrb. Syst. 8 (2): 83, 1886 (Stephani 1886d). * Plagiochila integrilobula Schifn., Hep. Fl. Buitenzorg: 170, 1900 (Schifner 1900a). 159 ** Plagiochila johannis-winkleri Herzog, Mitt. Inst. Allg. Bot. Hamburg 7 (3): 187, 1931 (Herzog 1931a). * Plagiochila kuhliana Sande Lac., Ann. Mus. Bot. Lugduno-Batavi 1: 292, 1864 (Sande Lacoste 1864). 160 159 Plagiochila integrilobula is possibly conspeciic with Plagiochila blepharophora (Söderström et al. 2010a). 160 Plagiochila kuhliana is possibly conspeciic with Plagiochila sciophila (Söderström et al. 2010a). World checklist of hornworts and liverworts 223 ** Plagiochila kurzii Steph., Bull. Herb. Boissier (sér. 2) 3 (2): 112 (292), 1903 (Stephani 1903b). ** Plagiochila reischeckiana Steph., Bull. Herb. Boissier (sér. 2) 3 (4): 331 (315), 1903 (Stephani 1903c). *** Plagiochila sandei Dozy ex Sande Lac., Plagiochila Sandei: 5, 1856 (Sande Lacoste 1856c). *** Plagiochila sciophila Nees, Sp. Hepat. (Lindenberg) 2-4: 100, 1840 (Lindenberg 1840). ** Plagiochila sciophila subsp. ciliigera (R.M.Schust.) L.Söderstr., Phytotaxa 208 (1): 84, 2015 (Söderström et al. 2015b). Bas.: Plagiochila japonica subsp. ciliigera R.M.Schust., Amer. Midl. Naturalist 62 (2): 354, 1959 (Schuster 1959b). * Plagiochila stephanii Schifn., Hep. Fl. Buitenzorg: 166, 1900 (Schifner 1900a). 161 ** Plagiochila subplana Lindenb., Sp. Hepat. (Lindenberg) 2-4: 73, 1840 (Lindenberg 1840). ** Plagiochila vitiensis Mitt., Bonplandia 9 (24): 367, 1861 (Mitten 1861). *** sect. Denticulatae Schifn., Hep. Fl. Buitenzorg: 106, 1900 (Schifner 1900a). *** Plagiochila alternans Lindenb. et Gottsche, Syn. Hepat. 5: 648, 1847 (Gottsche et al. 1847). ** Plagiochila ansata (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 5: 649, 1847 (Gottsche et al. 1847). Bas.: Jungermannia ansata Hook.f. et Taylor, London J. Bot. 3: 457, 1844 (Hooker and Taylor 1844b). *** Plagiochila banksiana Gottsche, Ann. Sci. Nat. Bot. (sér. 4) 8: 329, 1857 (Gottsche 1857). ** Plagiochila banksiana var. echinophora Inoue et R.M.Schust., J. Hattori Bot. Lab. 34: 62, 1971 (Inoue and Schuster 1971). ** Plagiochila chonotica Taylor, London J. Bot. 5: 260, 1846 (Taylor 1846a). ** Plagiochila equitans Gottsche, Ann. Sci. Nat. Bot. (sér. 4) 8: 331, 1857 (Gottsche 1857). * Plagiochila fragmentissima Inoue et R.M.Schust., J. Hattori Bot. Lab. 34: 155, 1971 (Inoue and Schuster 1971). 162 *** Plagiochila gigantea Lindenb., Sp. Hepat. (Lindenberg) 2-4: 115, 1840 (Lindenberg 1840). Nom. nov. pro Jungermannia gigantea Hook., Musci Exot. 1: tab. 93, 1818 (Hooker 1818), nom. illeg. ** Plagiochila gigantea var. inermis J.J.Engel et G.L.Merr., Nova Hedwigia 91 (3/4): 513, 2010 (Engel and Smith Merrill 2010). *** Plagiochila gregaria (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 5: 654, 1847 (Gottsche et al. 1847). Bas.: Jungermannia gregaria Hook.f. et Taylor, London J. Bot. 3: 564, 1844 (Hooker and Taylor 1844d). ** Plagiochila hookeriana Lindenb., Sp. Hepat. (Lindenberg) 2-4: 81, 1840 (Lindenberg 1840). 161 Plagiochila stephanii is possibly conspeciic with Plagiochila blepharophora (Söderström et al. 2010a). 162 Plagiochila fragmentissima may be conspeciic with Plagiochila gregaria (M.L. So and D. Glenny, pers. comm.). 224 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Plagiochila latifrons Gottsche et Hampe, Linnaea 27 (5): 553, 1854 (Hampe 1854). ** Plagiochila minutula (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 5: 652, 1847 (Gottsche et al. 1847). Bas.: Jungermannia minutula Hook.f. et Taylor, London J. Bot. 3: 459, 1844 (Hooker and Taylor 1844b). *** Plagiochila nobilis Gottsche, Bot. Zeitung (Berlin) Beil. 16: 37, 1858 (Gottsche 1858). ** Plagiochila obovata Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 33, 1911 (Stephani 1911b). *** Plagiochila ovata Lindenb. et Gottsche, Syn. Hepat. 5: 656, 1847 (Gottsche et al. 1847). *** Plagiochila retrospectans Lindenb., Sp. Hepat. (Lindenberg) 5: 123, 1843 (Lindenberg 1843). Nom. nov. pro Jungermannia retrospectans Nees, Linnaea 6 (4): 619, 1831 (Nees 1831), nom. illeg. ** Plagiochila riparia Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 34, 1911 (Stephani 1911b). ** Plagiochila rutlandii Steph., Bull. Herb. Boissier (sér. 2) 4 (8): 776 (454), 1904 (Stephani 1904c). ** Plagiochila subpectinata Besch. et C.Massal., Bull. Mens. Soc. Linn. Paris 1 (79): 628, 1886 (Bescherelle and Massalongo 1886). *** sect. Durae Carl, Ann. Bryol., Suppl. 2: 123, 1931 (Carl 1931b). ** Plagiochila acanthocaulis Sull., Hooker’s J. Bot. Kew Gard. Misc. 2: 317, 1850 (Sullivant 1850). ** Plagiochila angulata Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 6): 26, 1900 (Stephani 1900b). ** Plagiochila bicornuta Steph., Bot. Jahrb. Syst. 23 (1/2, 3): 305, 1896 (Stephani 1896a). ** Plagiochila crozetensis Kaal., Nyt Mag. Naturvidensk. 49 (2/3): 92, 1911 (Kaalaas 1911). *** Plagiochila deltoidea Lindenb., Sp. Hepat. (Lindenberg) 5: 132, 1843 (Lindenberg 1843). ** Plagiochila deltoidea var. densa J.J.Engel et G.L.Merr., Nova Hedwigia 91 (3/4): 506, 2010 (Engel and Smith Merrill 2010). ** Plagiochila dura De Not., Mem. Reale Accad. Sci. Torino (ser. 2) 16: 214, 1857 (De Notaris 1857). ** Plagiochila heterodonta (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 5: 638, 1847 (Gottsche et al. 1847). Bas.: Jungermannia heterodonta Hook.f. et Taylor, London J. Bot. 3: 460, 1844 (Hooker and Taylor 1844b). *** Plagiochila ramosissima (Hook.) Lindenb., Sp. Hepat. (Lindenberg) 2-4: 87, 1840 (Lindenberg 1840). Bas.: Jungermannia ramosissima Hook., Musci Exot. 1: tab. 92, 1818 (Hooker 1818). *** sect. Duseniae Carl, Ann. Bryol., Suppl. 2: 126, 1931 (Carl 1931b). ** Plagiochila dusenii Steph., Bull. Herb. Boissier (sér. 2) 4 (10): 979 (475), 1904 (Stephani 1904a). ** Plagiochila elata Taylor, London J. Bot. 5: 259, 1846 (Taylor 1846a). World checklist of hornworts and liverworts 225 ** Plagiochila lechleri Gottsche, Ann. Sci. Nat. Bot. (sér. 4) 8: 325, 1857 (Gottsche 1857). ** Plagiochila validissima Steph., Biblioth. Bot. 87 (2): 214, 1916 (Stephani 1916a). * sect. Flexicaules Carl, Ann. Bryol., Suppl. 2: 127, 1931 (Carl 1931b). ** Plagiochila lexicaulis Mont., Syn. Hepat. 5: 629, 1847 (Gottsche et al. 1847). *** sect. Fruticosae Inoue, Gen. Plagiochila SE Asia: 50, 1984 (Inoue 1984b). *** Plagiochila assamica Steph., Sp. Hepat. (Stephani) 6: 125, 1917 (Stephani 1917a). ** Plagiochila benitoi Inoue ex Piippo, J. Hattori Bot. Lab. 72: 122, 1992 (Piippo and Tan 1992). *** Plagiochila frondescens (Nees) Lindenb., Sp. Hepat. (Lindenberg) 2-4: 52, 1840 (Lindenberg 1840). Bas.: Jungermannia frondescens Nees, Linnaea 6 (4): 610, 1831 (Nees 1831). *** Plagiochila fruticosa Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 94, 1860 [1861] (Mitten 1860c). ** Plagiochila mauiensis Steph., Sp. Hepat. (Stephani) 6: 184, 1921 (Stephani 1921). *** Plagiochila pulcherrima Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 1: 63, 1931 (Horikawa 1931a). ** Plagiochila tamiensis Steph., Sp. Hepat. (Stephani) 6: 235, 1921 (Stephani 1921). *** *** *** *** ** ** *** *** *** *** *** sect. Fuscoluteae Carl, Ann. Bryol., Suppl. 2: 46, 1931 (Carl 1931b). Plagiochila aerea Taylor, London J. Bot. 5: 263, 1846 (Taylor 1846a). Plagiochila dependula Taylor, London J. Bot. 5: 265, 1846 (Taylor 1846a). Plagiochila fuscolutea Taylor, London J. Bot. 5: 263, 1846 (Taylor 1846a). Plagiochila heterophylla Lindenb., Nov. Stirp. Pug. 10: 2, 1857 (Lehmann 1857). Plagiochila heterophylla var. beauverdii (Steph.) Heinrichs, Bryophyt. Biblioth. 58: 148, 2002 (Heinrichs 2002). Bas.: Plagiochila beauverdii Steph., Biblioth. Bot. 87 (2): 191, 1916 (Stephani 1916a). Plagiochila longiramea Steph., Biblioth. Bot. 87 (2): 204, 1916 (Stephani 1916a). Plagiochila paraphyllina Herzog, Hedwigia 74 (2): 89, 1934 (Herzog 1934a). Plagiochila rudischusteri H.Rob., Beih. Nova Hedwigia 90: 199, 1988 (Robinson 1988). Plagiochila subbidentata Taylor, Ann. Mag. Nat. Hist. 20 (135): 381, 1847 (Taylor 1847a). Plagiochila tabinensis Steph., Bull. Herb. Boissier (sér. 2) 2 (10): 862 (231), 1902 (Stephani 1902g). *** sect. Glaucescentes Carl, Ann. Bryol., Suppl. 2: 70, 1931 (Carl 1931b). *** Plagiochila buchtiniana Steph., Sp. Hepat. (Stephani) 6: 135, 1918 (Stephani 1918). *** Plagiochila diversifolia Lindenb. et Gottsche, Syn. Hepat. 5: 640, 1847 (Gottsche et al. 1847). *** Plagiochila longispina Lindenb. et Gottsche, Syn. Hepat. 5: 642, 1847 (Gottsche et al. 1847). 226 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** sect. Hylacoetes Carl, Ann. Bryol., Suppl. 2: 50, 1931 (Carl 1931b). *** Plagiochila amicta Steph., Bull. Herb. Boissier (sér. 2) 5 (9): 895 (561), 1905 (Stephani 1905g). *** Plagiochila boryana Gottsche, Bull. Soc. Roy. Bot. Belgique 31: 118, 1892 (Stephani 1892c). *** Plagiochila breuteliana Lindenb., Sp. Hepat. (Lindenberg) 5: 150, 1843 (Lindenberg 1843). *** Plagiochila canelensis Steph., Bull. Herb. Boissier (sér. 2) 5 (10): 926 (576), 1905 (Stephani 1905f). *** Plagiochila cucullifolia J.B.Jack et Steph., Hedwigia 31 (1): 24, 1892 (Jack and Stephani 1892). *** Plagiochila cucullifolia var. anomala Heinrichs et Gradst., Pl. Syst. Evol. 242 (1/4): 208, 2003 (Heinrichs et al. 2003). *** Plagiochila dimorpha Lindenb. et Gottsche, Syn. Hepat. 5: 627, 1847 (Gottsche et al. 1847). *** Plagiochila dominicensis Taylor, London J. Bot. 5: 270, 1846 (Taylor 1846a). *** Plagiochila ecuadorica (Inoue) L.Söderstr., Phytotaxa 208 (1): 84, 2015 (Söderström et al. 2015b). Bas.: Steereochila ecuadorica Inoue, Mem. New York Bot. Gard. 45: 279, 1987 (Inoue 1987a). *** Plagiochila ensiformis Taylor, London J. Bot. 5: 265, 1846 (Taylor 1846a). *** Plagiochila labellilora Steph., Bull. Herb. Boissier (sér. 2) 2 (10): 880 (249), 1902 (Stephani 1902g). ** Plagiochila guevarii H.Rob., Bryologist 70 (1): 48, 1967 (Robinson 1967). ** Plagiochila husnotii Steph., Bull. Herb. Boissier (sér. 2) 5 (2): 178 (506), 1905 (Stephani 1905d). *** Plagiochila macrostachya Lindenb., Sp. Hepat. (Lindenberg) 2-4: 75, 1840 (Lindenberg 1840). *** Plagiochila patriciae Heinrichs et H.Anton, Bryophyt. Biblioth. 58: 107, 2002 (Heinrichs 2002). *** Plagiochila superba (Nees ex Spreng.) Mont. et Nees, Voy. Amér. Mérid., Bot. 7 (2): 81, 1839 (Montagne 1839a). Bas.: Jungermannia superba Nees ex Spreng. Syst. Veg. (ed. 16) [Sprengel] 4 (2): 326, 1827 (Sprengel 1827b). *** Plagiochila superba var. macrotricha (Spruce) Heinrichs, Bryophyt. Biblioth. 58: 114, 2002 (Heinrichs 2002). Bas.: Plagiochila macrotricha Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 476, 1885 (Spruce 1885). *** Plagiochila turgida Herzog, Hedwigia 72 (6): 196, 1932 (Herzog 1932c). *** Plagiochila vincentina Lindenb., Sp. Hepat. (Lindenberg) 2-4: 39, 1840 (Lindenberg 1840). * sect. Jacquinotiae Hässel, Nova Hedwigia 89 (1/2): 85, 2009 (Hässel 2009). ** Plagiochila jacquinotii Mont., Voy. Pole Sud, Bot. 1: 273, 1845 (Montagne 1845c). * sect. Kaalaasiae Carl, Ann. Bryol., Suppl. 2: 103, 1931 (Carl 1931b). ** Plagiochila amboynensis Taylor, London J. Bot. 5: 260, 1846 (Taylor 1846a). World checklist of hornworts and liverworts 227 * sect. Longilorae Carl, Ann. Bryol., Suppl. 2: 130, 1931 (Carl 1931b). ** Plagiochila longilora Mont., Syn. Hepat. 5: 651, 1847 (Gottsche et al. 1847). * sect. Oligodontes Carl, Ann. Bryol., Suppl. 2: 130, 1931 (Carl 1931b). ** Plagiochila lophocoleoides Mont., Ann. Sci. Nat. Bot. (sér. 3) 4: 348, 1845 (Montagne 1845b). *** sect. Peculiares Schifn., Hep. Fl. Buitenzorg: 107, 1900 (Schifner 1900a). ** Plagiochila aspericaulis Grolle et M.L.So, Syst. Bot. 24 (3): 307, 1999 (Grolle and So 1999b). ** Plagiochila caulimammillosa Grolle et M.L.So, J. Bryol. 20 (1): 42, 1998 (Grolle and So 1998c). *** Plagiochila devexa Steph., Bull. Herb. Boissier (sér. 2) 3 (4): 340 (324), 1903 (Stephani 1903c). Nom. nov. pro Plagiochila delexa Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 97, 1860 [1861] (Mitten 1860c), nom. illeg. *** Plagiochila durelii Schifn., Österr. Bot. Z. 49 (4): 131, 1899 (Schifner 1899b). ** Plagiochila durelii subsp. guizhouensis Grolle et M.L.So, Syst. Bot. 24 (3): 304, 1999 (Grolle and So 1999b). ** Plagiochila grollei Inoue, Bull. Natl. Sci. Mus. Tokyo (n.ser.) 8 (3): 384, 1965 (Inoue 1965a). ** Plagiochila hoei Inoue, J. Hattori Bot. Lab. 40: 421, 1976 (Inoue 1976b). ** Plagiochila hyalodermica Grolle et M.L.So, Bryologist 100 (4): 470, 1997 (Grolle and So 1997b). ** Plagiochila magna Inoue, J. Hattori Bot. Lab. 28: 216, 1965 (Inoue 1965c). ** Plagiochila paraphyllosa Grolle et M.L.So, Syst. Bot. 24 (3): 298, 1999 (Grolle and So 1999b). ** Plagiochila peculiaris Schifn., Hep. Fl. Buitenzorg: 157, 1900 (Schifner 1900a). *** Plagiochila perserrata Herzog, Symb. Sin. 5: 19, 1930 (Nicholson et al. 1930). ** Plagiochila philippinensis Steph., Bull. Herb. Boissier (sér. 2) 3 (6): 526 (330), 1903 (Stephani 1903d). *** Plagiochila pseudopoeltii Inoue, Bull. Natl. Sci. Mus. Tokyo (n.ser.) 8 (3): 382, 1965 (Inoue 1965a). *** Plagiochila renitens (Nees) Lindenb., Sp. Hepat. (Lindenberg) 2-4: 90, 1840 (Lindenberg 1840). Bas.: Jungermannia renitens Nees, Enum. Pl. Crypt. Javae: 76, 1830 (Nees 1830). ** Plagiochila semidecurrens (Lehm. et Lindenb.) Lindenb., Sp. Hepat. (Lindenberg) 5: 142, 1843 (Lindenberg 1843). Bas.: Jungermannia semidecurrens Lehm. et Lindenb., Nov. Stirp. Pug. 4: 21, 1832 (Lehmann 1832). ** Plagiochila semidecurrens var. alaskana (A.Evans) Inoue, J. Hattori Bot. Lab. 28: 216, 1965 (Inoue 1965c). Bas.: Plagiochila alaskana A.Evans, Bull. Torrey Bot. Club 41 (12): 590, 1914 [1915] (Evans 1914a). *** Plagiochila vexans Schifn. ex Steph., Sp. Hepat. (Stephani) 6: 237, 1921 (Stephani 1921). 228 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Plagiochila zangii Grolle et M.L.So, Bryologist 100 (4): 467, 1997 (Grolle and So 1997b). *** Plagiochila zonata Steph., Mém. Soc. Nat. Sci. Nat. Math. Cherbourg 29: 225, 1894 (Stephani 1894b). *** sect. Plagiochila ** Plagiochila arctica Bryhn et Kaal., Rep. Second Norweg. Arctic Exped. 11: 41, 1906 (Bryhn 1906). * Plagiochila arctica var. intermedia R.M.Schust., Amer. Midl. Naturalist 62 (1): 152, 1959 (Schuster 1959a). *** Plagiochila asplenioides (L.) Dumort., Recueil Observ. Jungerm.: 14, 1835 (Dumortier 1835). Bas.: Jungermannia asplenioides L., Sp. Pl. 1: 1131, 1753 (Linnaeus 1753). *** Plagiochila bischleriana Grolle et M.L.So, Cryptog. Bryol. Lichénol. 18 (3): 191, 1997 (Grolle and So 1997a). ** Plagiochila britannica Paton, J. Bryol. 10 (3): 245, 1979 (Paton 1979b). *** Plagiochila chinensis Steph., Mém. Soc. Nat. Sci. Nat. Math. Cherbourg 29: 223, 1894 (Stephani 1894b). *** Plagiochila circumdentata Steph., Bull. Herb. Boissier (sér. 2) 4 (8): 778 (456), 1904 (Stephani 1904c). ** Plagiochila circumdentata var. carinata J.J.Engel et G.L.Merr., Nova Hedwigia 91 (3/4): 511, 2010 (Engel and Smith Merrill 2010). ** Plagiochila circumserrata Inoue et Grolle, Bull. Natl. Sci. Mus. Tokyo, B 5 (1): 27, 1979 (Inoue 1979c). ** Plagiochila columbiana A.Evans, Bot. Gaz. 21 (4): 189, 1896 (Evans 1896). *** Plagiochila delavayi Steph., Mém. Soc. Nat. Sci. Nat. Math. Cherbourg 29: 224, 1894 (Stephani 1894b). *** Plagiochila elegans Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 97, 1860 [1861] (Mitten 1860c). *** Plagiochila hakkodensis Steph., Bull. Herb. Boissier 5 (2): 103, 1897 (Stephani 1897b). ** Plagiochila korthalsiana Molk., Ned. Kruidk. Arch. 3: 416, 1854 [1855] (Sande Lacoste 1854). ** Plagiochila loriana Steph., Bull. Herb. Boissier (sér. 2) 3 (7): 608 (354), 1903 (Stephani 1903e). ** Plagiochila microdentata M.L.So, New Zealand J. Bot. 38 (3): 426, 2000 (So 2000b). ** Plagiochila mundaliensis Steph., Bull. Herb. Boissier (sér. 2) 3 (6): 536 (340), 1903 (Stephani 1903d). ** Plagiochila orbicularis (S.Hatt.) S.Hatt., J. Hattori Bot. Lab. 3: 26, 1948 [1950] (Hattori 1948b). Bas.: Plagiochila ovalifolia var. orbicularis S.Hatt., Bull. Tokyo Sci. Mus. 11: 61, 1944 (Hattori 1944d). *** Plagiochila ovalifolia Mitt., Trans. Linn. Soc. London, Bot. 3 (3): 193, 1891 (Mitten 1891). *** Plagiochila porelloides (Torr. ex Nees) Lindenb., Sp. Hepat. (Lindenberg) 2-4: 61, 1840 (Lindenberg 1840). Bas.: Jungermannia porelloides Torr. ex Nees, Naturgesch. Eur. Leberm. 1: 170, 1833 (Nees 1833c). World checklist of hornworts and liverworts 229 ** Plagiochila porelloides var. norvegica (H.H.Blom et Holten) Schumacker et Váňa, Identif. keys liverw. hornw. Europe: 131, 2005 (Schumacker and Váňa 2005). Bas.: Plagiochila norvegica H.H.Blom et Holten, Lindbergia 14 (1): 8, 1988 (Blom and Holten 1988). ** Plagiochila porelloides var. subarctica (Jørg.) Lammes, Fl. Fenn. 6: 54, 1977 (Koponen et al. 1977). Bas.: Plagiochila asplenioides var. subarctica Jørg., Bergens Mus. Skr. (n.ser.) 16: 173, 1934 (Jørgensen 1934). ** Plagiochila schoieldiana Inoue, Bull. Natl. Sci. Mus. Tokyo (n.ser.) 15 (1): 183, 1972 (Inoue 1972c). *** Plagiochila secretifolia Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 98, 1860 [1861] (Mitten 1860c). ** Plagiochila spinosa M.L.So, New Zealand J. Bot. 38 (3): 428, 2000 (So 2000b). ** Plagiochila sumatrana Schifn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 70: 183, 1900 [1901] (Schifner 1900c). ** Plagiochila taiwanensis Inoue, Bull. Natl. Sci. Mus. Tokyo, B 8 (4): 136, 1982 (Inoue 1982). *** Plagiochila trapezoidea Lindenb., Sp. Hepat. (Lindenberg) 2-4: 112, 1840 (Lindenberg 1840). Nom. nov. pro Jungermannia asplenioides β australis Nees, Enum. Pl. Crypt. Javae: 73, 1830 (Nees 1830). ** Plagiochila uniformis Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 98, 1860 [1861] (Mitten 1860c). ** sect. Poeltiae Inoue, Bull. Natl. Sci. Mus. Tokyo (n.ser.) 8 (3): 394, 1965 (Inoue 1965a). *** Plagiochila biondiana C.Massal., Hepat. Shen-si: 15, 1897 (Massalongo 1897). *** Plagiochila carringtonii (Balf. ex Carrington) Grolle, Trans. Brit. Bryol. Soc. 4 (4): 656, 1964 (Grolle 1964i). Bas.: Adelanthus carringtonii Balf. ex Carrington, Trans. Bot. Soc. Edinburgh 10: 380, 1870 (Carrington 1870). ** Plagiochila carringtonii subsp. lobuchensis Grolle, Trans. Brit. Bryol. Soc. 4 (4): 660, 1964 (Grolle 1964i). *** Plagiochila duthiana Steph., Bull. Herb. Boissier (sér. 2) 3 (6): 527 (331), 1903 (Stephani 1903d). ** Plagiochila erlangensis M.L.So, Haussknechtia, Beih. 9: 350, 1999 (So 1999). *** Plagiochila poeltii Inoue et Grolle, Trans. Brit. Bryol. Soc. 4 (4): 656, 1964 (Grolle 1964i). *** Plagiochila recurvata (W.E.Nicholson) Grolle, Trans. Brit. Bryol. Soc. 4 (4): 654, 1964 (Grolle 1964i). Bas.: Jamesoniella carringtonii var. recurvata W.E.Nicholson, Symb. Sin. 5: 13, 1930 (Nicholson et al. 1930). *** Plagiochila retusa Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 96, 1860 [1861] (Mitten 1860c). *** Plagiochila wallichiana Steph., Bull. Herb. Boissier (sér. 2) 3 (6): 523 (327), 1903 (Stephani 1903d). ** Plagiochila wangii Inoue, J. Jap. Bot. 37 (6): 187, 1962 (Inoue 1962c). ** Plagiochila yulungensis Piippo, Ann. Bot. Fenn. 34 (4): 283, 1997 (Piippo 1997). 230 *** *** ** ** * *** *** *** *** ** *** *** *** ** ** ** ** *** *** *** *** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) sect. Rutilantes Carl, Ann. Bryol., Suppl. 2: 83, 1931 (Carl 1931b). Plagiochila bicuspidata Gottsche, Mexik. Leverm.: 43, 1863 (Gottsche 1863). Plagiochila caduciloba H.L.Blomq., Bryologist 42 (5): 114, 1939 (Blomquist 1939). Plagiochila chimborazensis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 469, 1885 (Spruce 1885). Plagiochila cuneata Lindenb. et Gottsche, Syn. Hepat. 5: 632, 1847 (Gottsche et al. 1847). 163 Plagiochila debilis Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 97, 1860 [1861] (Mitten 1860c). Plagiochila delexa Mont. et Gottsche, Ann. Sci. Nat. Bot. (sér. 4) 6: 192, 1856 (Montagne 1856a). Plagiochila defolians Grolle et M.L.So, Syst. Bot. 23 (4): 459, 1998 [1999] (Grolle and So 1998b). Plagiochila exigua (Taylor) Taylor, London J. Bot. 5: 264, 1846 (Taylor 1846a). Bas.: Jungermannia exigua Taylor, Trans. Bot. Soc. Edinburgh 1: 179, 1844 (Taylor 1844b). Plagiochila fracta Pócs, Phytotaxa 195 (2): 183, 2015 (Pócs 2015a). Plagiochila ghatiensis Steph., Sp. Hepat. (Stephani) 6: 159, 1918 (Stephani 1918). Plagiochila grossa Grolle et M.L.So, Syst. Bot. 23 (4): 461, 1998 [1999] (Grolle and So 1998b). Plagiochila gymnocalycina (Lehm. et Lindenb.) Mont. et Nees, Voy. Amér. Mérid., Bot. 7 (2): 81, 1839 (Montagne 1839a). Bas.: Jungermannia gymnocalycina Lehm. et Lindenb., Nov. Stirp. Pug. 5: 28, 1833 (Lehmann 1833). Plagiochila gymnocalycina var. surinamensis (Molk.) Heinrichs et D.S.Rycroft, J. Hattori Bot. Lab. 100: 139, 2006 (Heinrichs et al. 2006). Bas.: Plagiochila surinamensis Molk., Syn. hepat. jav.: 103, 1856 [1857] (Sande Lacoste 1856b). Plagiochila loriloba Herzog ex Carl, Ann. Bryol., Suppl. 2: 47, 1931 (Carl 1931b). Plagiochila maderensis Gottsche ex Steph., Bull. Herb. Boissier (sér. 2) 4 (4): 350 (426), 1904 (Stephani 1904e). Plagiochila oresitropha Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 467, 1885 (Spruce 1885). Plagiochila pectinata Lindenb., Sp. Hepat. (Lindenberg) 1: 14, 1839 (Lindenberg 1839). Plagiochila rutilans Lindenb., Sp. Hepat. (Lindenberg) 2-4: 47, 1840 (Lindenberg 1840). Plagiochila rutilans var. moritziana (Lindenb. et Gottsche) Heinrichs, Bryologist 105 (2): 197, 2002 (Heinrichs et al. 2002). Bas.: Plagiochila moritziana Lindenb. et Gottsche, Linnaea 20 (3): 323, 1847 (Hampe 1847). Plagiochila rutilans var. standleyi (Herzog ex Carl) Heinrichs et D.S.Rycroft, Bryologist 104 (3): 357, 2001 (Heinrichs et al. 2001). Bas.: Plagiochila standleyi Herzog ex Carl, Ann. Bryol., Suppl. 2: 80, 1931 (Carl 1931b). 163 Plagiochila cuneata is conspeciic with Plagiochila bursata in Inoue (1980), but accepted by Grolle and Heinrichs (1999). World checklist of hornworts and liverworts 231 * Plagiochila steyermarkii H.Rob., Bryologist 68 (1): 93, 1965 (Robinson 1965). 164 *** Plagiochila trichostoma Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 113, 1864 (Gottsche 1864). sect. Strombifoliae Inoue et R.M.Schust., J. Hattori Bot. Lab. 34: 130, 1971 (Inoue and Schuster 1971). ** Plagiochila strombifolia Taylor, Nov. Stirp. Pug. 8: 5, 1844 (Lehmann 1844). * *** sect. Tayloriae Carl, Ann. Bryol., Suppl. 2: 140, 1931 (Carl 1931b). *** Plagiochila annotina Lindenb., Sp. Hepat. (Lindenberg) 1: 34, 1839 (Lindenberg 1839). *** Plagiochila baylisii Inoue et R.M.Schust., J. Hattori Bot. Lab. 34: 150, 1971 (Inoue and Schuster 1971). *** Plagiochila bazzanioides J.J.Engel et G.L.Merr., Novon 9 (1): 29, 1999 (Engel and Smith Merrill 1999b). *** Plagiochila chenii Grolle et M.L.So, Syst. Bot. 25 (1): 6, 2000 (Grolle and So 2000). *** Plagiochila circinalis (Lehm. et Lindenb.) Lehm., Sp. Hepat. (Lindenberg) 5: 124, 1843 (Lindenberg 1843). Bas.: Jungermannia circinalis Lehm. et Lindenb., Nov. Stirp. Pug. 4: 64, 1832 (Lehmann 1832). ** Plagiochila circinalis var. hemicardia (Hook.f. et Taylor) J.J.Engel et G.L.Merr., Nova Hedwigia 91 (3/4): 512, 2010 (Engel and Smith Merrill 2010). Bas.: Jungermannia hemicardia Hook.f. et Taylor, London J. Bot. 3: 371, 1844 (Hooker and Taylor 1844a). *** Plagiochila colensoi Hook.f. et Taylor, London J. Bot. 5: 269, 1846 (Taylor 1846a). ** Plagiochila colensoi var. quinquespina (Steph.) J.J.Engel et G.L.Merr., Nova Hedwigia 91 (3/4): 504, 2010 (Engel and Smith Merrill 2010). Bas.: Plagiochila quinquespina Steph., Bull. Herb. Boissier (sér. 2) 3 (4): 328 (312), 1903 (Stephani 1903c). ** Plagiochila corticola Steph., Mém. Soc. Nat. Sci. Nat. Math. Cherbourg 29: 224, 1894 (Stephani 1894b). ** Plagiochila fasciculata Lindenb., Sp. Hepat. (Lindenberg) 1: 7, 1839 (Lindenberg 1839). *** Plagiochila fusca Sande Lac., Ned. Kruidk. Arch. 3: 417, 1854 [1855] (Sande Lacoste 1854). ** Plagiochila fuscella (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 5: 648, 1847 (Gottsche et al. 1847). Bas.: Jungermannia fuscella Hook.f. et Taylor, London J. Bot. 3: 373, 1844 (Hooker and Taylor 1844a). ** Plagiochila fuscella var. novae-zelandiae (E.A.Hodgs.) J.J.Engel et G.L.Merr., Nova Hedwigia 89 (3/4): 294, 2009 (Engel and Smith Merrill 2009). Bas.: Plagiochila retrospectans var. novae-zelandiae E.A.Hodgs., Trans. & Proc. Roy. Soc. New Zealand 73 (4): 293, 1944 (Hodgson 1944). 164 Plagiochila steyermarkii is conspeciic with Plagiochila aerea in Inoue in Gradstein and Hekking (1979), but this was rejected by Grolle and Heinrichs (1999). 232 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Plagiochila gracilis Lindenb. et Gottsche, Syn. Hepat. 5: 632, 1847 (Gottsche et al. 1847). *** Plagiochila gymnoclada Sande Lac., Plagiochila Sandei: 6, 1856 (Sande Lacoste 1856c). *** Plagiochila himalayana Schifn., Österr. Bot. Z. 49 (4): 131, 1899 (Schifner 1899b). ** Plagiochila incurvicolla (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 5: 651, 1847 (Gottsche et al. 1847). Bas.: Jungermannia incurvicolla Hook.f. et Taylor, London J. Bot. 3: 564, 1844 (Hooker and Taylor 1844d). ** Plagiochila incurvicolla var. lonchoscypha (Herzog) J.J.Engel et G.L.Merr., Nova Hedwigia 91 (3/4): 505, 2010 (Engel and Smith Merrill 2010). Bas.: Plagiochila lonchoscypha Herzog, Trans. & Proc. Roy. Soc. New Zealand 68 (1): 42, 1938 (Herzog 1938c). ** Plagiochila microdictyon Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 131, 1854 (Mitten 1854). ** Plagiochila monospiris Inoue et Grolle, J. Hattori Bot. Lab. 36: 489, 1972 [1973] (Inoue 1972b). *** Plagiochila nitens Inoue, Willdenowia 18 (2): 561, 1989 (Inoue 1989b). ** Plagiochila pseudocapillaris Inoue, Bull. Natl. Sci. Mus. Tokyo (n.ser.) 11 (3): 302, 1968 (Inoue 1968a). *** Plagiochila pseudoirma Herzog, Symb. Sin. 5: 17, 1930 (Nicholson et al. 1930). ** Plagiochila radiculosa Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 132, 1854 (Mitten 1854). ** Plagiochila spathulifolia Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 96, 1860 [1861] (Mitten 1860c). *** Plagiochila stephensoniana Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 133, 1854 (Mitten 1854). *** sect. Trabeculatae S.Hatt. ex Inoue, J. Hattori Bot. Lab. 20: 75, 1958 (Inoue 1958). ** Plagiochila austinii A.Evans, Rhodora 16 (184): 68, 1914 (Evans 1914b). *** Plagiochila lexuosa Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 94, 1860 [1861] (Mitten 1860c). ** Plagiochila sullivantii Gottsche, Bot. Gaz. 21 (4): 191, 1896 (Evans 1896). ** Plagiochila sullivantii var. spinigera R.M.Schust., Amer. Midl. Naturalist 62 (2): 323, 1959 (Schuster 1959b). *** Plagiochila trabeculata Steph., Bull. Herb. Boissier (sér. 2) 3 (2): 103 (283), 1903 (Stephani 1903b). *** sect. Vagae Lindenb., Monogr. hep. gen. Plagiochilae: xv, 1844 (Lindenberg 1844). *** Plagiochila abietina (Nees) Mont. et Nees, Voy. Amér. Mérid., Bot. 7 (2): 81, 1839 (Montagne 1839a). Bas.: Jungermannia abietina Nees, Enum. Pl. Crypt. Javae: 76, 1830 (Nees 1830). ** Plagiochila abrupta Lehm. et Lindenb., Sp. Hepat. (Lindenberg) 2-4: 106, 1840 (Lindenberg 1840). World checklist of hornworts and liverworts 233 ** Plagiochila aequatorialis Gottsche, Ann. Sci. Nat. Bot. (sér. 4) 8: 334, 1857 (Gottsche 1857). *** Plagiochila africana Steph., Bull. Herb. Boissier (sér. 2) 2 (12): 973 (263), 1902 (Stephani 1902h). *** Plagiochila akiyamae Inoue, Bull. Natl. Sci. Mus. Tokyo, B 12 (3): 73, 1986 (Inoue 1986). *** Plagiochila angusta Lindenb., Sp. Hepat. (Lindenberg) 5: 148, 1843 (Lindenberg 1843). *** Plagiochila angustitexta Steph., Bull. Herb. Boissier (sér. 2) 2 (12): 977 (267), 1902 (Stephani 1902h). *** Plagiochila arbuscula (Brid. ex Lehm. et Lindenb.) Lindenb., Sp. Hepat. (Lindenberg) 1: 23, 1839 (Lindenberg 1839). Bas.: Jungermannia arbuscula Brid. ex Lehm. et Lindenb., Nov. Stirp. Pug. 4: 63, 1832 (Lehmann 1832). ** Plagiochila arbuscula var. rekohuensis J.J.Engel et G.L.Merr., Nova Hedwigia 91 (3/4): 509, 2010 (Engel and Smith Merrill 2010). * Plagiochila aspera Steph., Sp. Hepat. (Stephani) 6: 125, 1917 (Stephani 1917a). 165 ** Plagiochila aspleniformis R.M.Schust., Amer. Midl. Naturalist 63 (1): 51, 1960 (Schuster 1960c). ** Plagiochila beddomei Steph., Bull. Herb. Boissier (sér. 2) 3 (10): 876 (361), 1903 (Stephani 1903f). *** Plagiochila boivinii Steph., Bull. Herb. Boissier (sér. 2) 2 (12): 987 (277), 1902 (Stephani 1902h). *** Plagiochila bryopteroides Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 499, 1885 (Spruce 1885). ** Plagiochila contigua Gottsche, Mexik. Leverm.: 30, 1863 (Gottsche 1863). *** Plagiochila corrugata (Nees) Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 5: 52, 1836 (Nees and Montagne 1836). Bas.: Jungermannia corrugata Nees, Fl. Bras. (Martius) 1 (1): 378, 1833 (Nees 1833a). *** Plagiochila cuspidata Steph., Sp. Hepat. (Stephani) 6: 144, 1918 (Stephani 1918). ** Plagiochila cymata Inoue et Grolle, Bull. Natl. Sci. Mus. Tokyo, B 1 (3): 95, 1975 (Inoue 1975a). ** Plagiochila delexirama Taylor, London J. Bot. 5: 262, 1846 (Taylor 1846a). *** Plagiochila dichotoma (P.Beauv.) Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 5: 53, 1836 (Nees and Montagne 1836). Bas.: Carpolepidum dichotomum P.Beauv., Fl. Oware 1 (3): 23, 1805 (Palisot de Beauvois 1805b). * Plagiochila dichotoma var. luitans Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 490, 1885 (Spruce 1885). *** Plagiochila dissecta Steph., Bull. Herb. Boissier (sér. 2) 3 (7): 600 (346), 1903 (Stephani 1903e). 165 Plagiochila aspera is conspeciic with Plagiochila trigona Steph. (=Plagiochila metcalii) in Inoue (1970b), but it was accepted by So (2000a, 2001a). 234 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Plagiochila disticha (Lehm. et Lindenb.) Lindenb., Sp. Hepat. (Lindenberg) 2-4: 107, 1840 (Lindenberg 1840). Bas.: Jungermannia disticha Lehm. et Lindenb., Nov. Stirp. Pug. 6: 64, 1834 (Lehmann 1834). ** Plagiochila distinctifolia Lindenb., Sp. Hepat. (Lindenberg) 1: 17, 1839 (Lindenberg 1839). *** Plagiochila divergens Steph., Hedwigia 30 (6): 268, 1891 (Stephani 1891c). *** Plagiochila drepanophylla Sande Lac., Syn. hepat. jav.: 103, 1856 [1857] (Sande Lacoste 1856b). * Plagiochila drepanophylla var. minor Gottsche, Abh. Naturwiss. Vereins Bremen 7: 341, 1882 (Gottsche 1882). *** Plagiochila efusa Steph., Bot. Jahrb. Syst. 20 (3): 310, 1895 (Stephani 1895a). * Plagiochila efusa var. decurrens Steph., Bot. Jahrb. Syst. 20 (3): 310, 1895 (Stephani 1895a). *** Plagiochila ericicola Steph., Bull. Herb. Boissier (sér. 2) 4 (6): 590 (441), 1904 (Stephani 1904b). ** Plagiochila exinnovata Steph., Bull. Herb. Boissier (sér. 2) 3 (7): 600 (346), 1903 (Stephani 1903e). ** Plagiochila fastigiata Lindenb. et Gottsche, Syn. Hepat. 5: 657, 1847 (Gottsche et al. 1847). *** Plagiochila labellata Steph., Bot. Jahrb. Syst. 8 (2): 82, 1886 (Stephani 1886d). ** Plagiochila loridana A.Evans, Bot. Gaz. 21 (4): 190, 1896 (Evans 1896). *** Plagiochila fordiana Steph., Bull. Herb. Boissier (sér. 2) 3 (2): 104 (284), 1903 (Stephani 1903b). ** Plagiochila francana Steph., Sp. Hepat. (Stephani) 6: 157, 1918 (Stephani 1918). *** Plagiochila furcifolia Mitt., Trans. Linn. Soc. London, Bot. 3 (3): 194, 1891 (Mitten 1891). *** Plagiochila fusifera Taylor, London J. Bot. 5: 268, 1846 (Taylor 1846a). *** Plagiochila hampeana Gottsche, Bot. Zeitung (Berlin) Beil. 16: 38, 1858 (Gottsche 1858). * Plagiochila heterospina Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 128, 1914 (Stephani and Watts 1914). * Plagiochila heterostipa Steph., Hedwigia 31 (3): 129, 1892 (Stephani 1892d). 166 ** Plagiochila incerta Gottsche, Ann. Sci. Nat. Bot. (sér. 4) 8: 324, 1857 (Gottsche 1857). *** Plagiochila indica Mitt. ex Steph., Bull. Herb. Boissier (sér. 2) 3 (6): 532 (336), 1903 (Stephani 1903d). ** Plagiochila invisa (R.M.Schust.) R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 513, 1980 (Schuster 1980c). Bas.: Plagiochila ludoviciana var. invisa R.M.Schust., Amer. Midl. Naturalist 63 (1): 101, 1960 (Schuster 1960c). *** Plagiochila javanica (Sw.) Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 5: 52, 1836 (Nees and Montagne 1836). Bas.: Jungermannia javanica Sw., Meth. Musc.: 35, 1781 (Swartz 1781). 166 Plagiochila heterostipa is possibly conspeciic with Plagiochila corymbulosa (Jones 1962). World checklist of hornworts and liverworts 235 *** Plagiochila junghuhniana Sande Lac., Ned. Kruidk. Arch. 3: 416, 1854 [1855] (Sande Lacoste 1854). *** Plagiochila khasiana Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 95, 1860 [1861] (Mitten 1860c). ** Plagiochila kiaeri Gottsche, Abh. Naturwiss. Vereins Bremen 7: 341, 1882 (Gottsche 1882). ** Plagiochila kiaeri var. capensis (Steph.) M.Wigginton et Grolle, Bryophyt. Biblioth. 50: 182, 1996 (Wigginton and Grolle 1996). Bas.: Plagiochila capensis Steph., Bull. Herb. Boissier (sér. 2) 4 (4): 350 (426), 1904 (Stephani 1904e). ** Plagiochila kiaeri var. myriocarpa (Pearson) Pócs, J. E. Afr. Nat. Hist. 96 (1): 38, 2007 (Pócs and Luke 2007). Bas.: Plagiochila myriocarpa Pearson, Ark. Bot. 19 (5): 5, 1924 (Pearson 1924b). *** Plagiochila kunmingensis Piippo, Ann. Bot. Fenn. 34 (4): 281, 1997 (Piippo 1997). ** Plagiochila kurokawae Inoue, J. Hattori Bot. Lab. 32: 104, 1969 (Inoue 1969). *** Plagiochila laetevirens Lindenb., Sp. Hepat. (Lindenberg) 2-4: 101, 1840 (Lindenberg 1840). ** Plagiochila lamellistipula Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 491, 1885 (Spruce 1885). *** Plagiochila lastii Mitt., J. Linn. Soc., Bot. 22 (146): 320, 1886 (Mitten 1886b). ** Plagiochila latifolia Steph., Bull. Herb. Boissier (sér. 2) 5 (8): 742 (541), 1905 (Stephani 1905i). * Plagiochila loloënsis Steph., Bull. Herb. Boissier (sér. 2) 4 (2): 166 (418), 1904 (Stephani 1904f). 167 ** Plagiochila manillana Mont. et Gottsche, Ann. Sci. Nat. Bot. (sér. 4) 6: 189, 1856 (Montagne 1856a). ** Plagiochila massalongoana Schifn., Hep. Fl. Buitenzorg: 136, 1900 (Schifner 1900a). ** Plagiochila mastigophoroides Inoue, J. Hattori Bot. Lab. 32: 99, 1969 (Inoue 1969). ** Plagiochila metcalii Steph., Bull. Herb. Boissier (sér. 2) 3 (6): 533 (337), 1903 (Stephani 1903d). ** Plagiochila micropteryx Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 107, 1864 (Gottsche 1864). ** Plagiochila miradorensis Gottsche, Mexik. Leverm.: 31, 1863 (Gottsche 1863). ** Plagiochila miradorensis var. convoluta R.M.Schust., Amer. Midl. Naturalist 63 (1): 113, 1960 (Schuster 1960c). * Plagiochila moenkemeyeri Steph., Bull. Herb. Boissier (sér. 2) 4 (2): 160 (412), 1904 (Stephani 1904f). 168 *** Plagiochila montagnei Nees, Ann. Sci. Nat. Bot. (sér. 2) 5: 53, 1836 (Nees and Montagne 1836). 167 Plagiochila loloënsis is probably an entire-leaved form of Plagiochila moenkemeyeri (Wigginton and Grolle 1996). 168 Plagiochila moenkemeyeri is possibly conspeciic with Plagiochila winteri (Jones 1962), but he expressed some doubts on the synonymy. 236 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Plagiochila morobensis Inoue et Piippo, Ann. Bot. Fenn. 26 (2): 203, 1989 (Piippo 1989b). ** Plagiochila multipinnula Herzog et S.Hatt., J. Hattori Bot. Lab. 14: 36, 1955 (Herzog and Noguchi 1955). *** Plagiochila neckeroidea Mitt., Trans. Linn. Soc. London 23 (1): 57, 1860 (Mitten 1860a). *** Plagiochila nepalensis Lindenb., Sp. Hepat. (Lindenberg) 2-4: 93, 1840 (Lindenberg 1840). ** Plagiochila norfolkiensis Steph., Bull. Herb. Boissier (sér. 2) 3 (10): 877 (362), 1903 (Stephani 1903f). *** Plagiochila obtusa Lindenb., Sp. Hepat. (Lindenberg) 2-4: 42, 1840 (Lindenberg 1840). * Plagiochila owaihiensis Nees et Lindenb., Sp. Hepat. (Lindenberg) 1: 30, 1839 (Lindenberg 1839). 169 ** Plagiochila paciica Mitt., Fl. vit.: 407, 1871 [1873] (Mitten 1871). ** Plagiochila parallela Steph., Bull. Herb. Boissier (sér. 2) 2 (8): 686 (223), 1902 (Stephani 1902a). *** Plagiochila parvifolia Lindenb., Sp. Hepat. (Lindenberg) 1: 28, 1839 (Lindenberg 1839). *** Plagiochila patentissima Lindenb., Sp. Hepat. (Lindenberg) 2-4: 64, 1840 (Lindenberg 1840). *** Plagiochila patula (Sw.) Lindenb., Sp. Hepat. (Lindenberg) 1: 21, 1839 (Lindenberg 1839). Bas.: Jungermannia patula Sw., Fl. Ind. Occid. 3: 1844, 1806 (Swartz 1806). ** Plagiochila patula var. brevifolia Gottsche, Mexik. Leverm.: 10, 1863 (Gottsche 1863). ** Plagiochila paucidens Steph., Bull. Herb. Boissier (sér. 2) 3 (2): 117 (297), 1903 (Stephani 1903b). ** Plagiochila pensilis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 497, 1885 (Spruce 1885). ** Plagiochila peradenyensis Schifn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 70: 172, 1900 [1901] (Schifner 1900c). ** Plagiochila perdentata M.L.So et Grolle, Syst. Bot. 26 (3): 460, 2001 (So and Grolle 2001). *** Plagiochila pinnilora Steph., Hedwigia 30 (5): 212, 1891 (Stephani 1891a). *** Plagiochila praemorsa Steph., Bot. Jahrb. Syst. 8 (2): 92, 1886 (Stephani 1886d). ** Plagiochila propinqua Sande Lac., Plagiochila Sandei: 5, 1856 (Sande Lacoste 1856c). ** Plagiochila purpurascens Steph., Sp. Hepat. (Stephani) 6: 197, 1921 (Stephani 1921). *** Plagiochila raddiana Lindenb., Sp. Hepat. (Lindenberg) 1: 9, 1839 (Lindenberg 1839). * Plagiochila ragazzii Gola, Ann. Bot. (Rome) 13 (1): 67, 1914 (Gola 1914a). *** Plagiochila repanda (Schwägr.) Lindenb., Sp. Hepat. (Lindenberg) 2-4: 62, 1840 (Lindenberg 1840). Bas.: Jungermannia repanda Schwägr., Hist. Musc. Hepat. Prodr.: 26, 1814 (Schwägrichen 1814). 169 Plagiochila owaihiensis was considered conspeciic with Plagiochila remyana by Inoue (1976c), but So (2000a) did not agree. World checklist of hornworts and liverworts 237 ** Plagiochila repanda var. perrotana (Steph.) Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 51 (1/2): 65, 1981 (Vanden Berghen 1981). Bas.: Plagiochila perrotana Steph., Bull. Herb. Boissier (sér. 2) 4 (6): 586 (437), 1904 (Stephani 1904b). *** Plagiochila rodriguezii Steph., Bot. Gaz. 15 (11): 290, 1890 (Stephani 1890c). *** Plagiochila rudolii Pócs, Beih. Nova Hedwigia 90: 223, 1988 (Pócs 1988). *** Plagiochila salacensis Gottsche, Natuurk. Tijdschr. Ned.-Indië 4: 576, 1853 (Gottsche 1853). *** Plagiochila salvadorica Steph., Hedwigia 30 (6): 272, 1891 (Stephani 1891c). ** Plagiochila serrialata Herzog, Hedwigia 72 (6): 212, 1932 (Herzog 1932c). *** Plagiochila shangaica Steph., Sp. Hepat. (Stephani) 6: 216, 1921 (Stephani 1921). *** Plagiochila simplex (Sw.) Lindenb., Sp. Hepat. (Lindenberg) 2-4: 54, 1840 (Lindenberg 1840). Bas.: Jungermannia simplex Sw., Prodr. (Swartz): 143, 1788 (Swartz 1788). *** Plagiochila squamulosa Mitt., J. Proc. Linn. Soc., Bot. 7 (27): 165, 1863 (Mitten 1863). ** Plagiochila squamulosa var. crispulocaudata (Gottsche) Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 51 (1/2): 74, 1981 (Vanden Berghen 1981). Bas.: Plagiochila crispulocaudata Gottsche, Abh. Naturwiss. Vereins Bremen 7: 340, 1882 (Gottsche 1882). ** Plagiochila squamulosa var. sinuosa (Mitt.) Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 51 (1/2): 75, 1981 (Vanden Berghen 1981). Bas.: Plagiochila sinuosa Mitt., J. Linn. Soc., Bot. 22 (146): 319, 1886 (Mitten 1886b). ** Plagiochila streimannii Inoue, J. Jap. Bot. 63 (11): 365, 1988 (Inoue 1988c). *** Plagiochila strictifolia Steph., Hedwigia 30 (5): 210, 1891 (Stephani 1891a). ** Plagiochila sublabellata Colenso, Trans. & Proc. New Zealand Inst. 21: 51, 1889 (Colenso 1889). ** Plagiochila subjavanica M.L.So, Austral. Syst. Bot. 13 (5): 804, 2000 (So 2000a). *** Plagiochila subtropica Steph., Bull. Soc. Roy. Bot. Belgique 38 (1): 46, 1899 (Stephani 1899h). ** Plagiochila tamariscina Steph., Bull. Herb. Boissier (sér. 2) 2 (8): 685 (222), 1902 (Stephani 1902a). ** Plagiochila tecta Inoue et Grolle, J. Hattori Bot. Lab. 33: 319, 1970 (Inoue 1970a). *** Plagiochila terebrans Nees et Mont., Sp. Hepat. (Lindenberg) 2-4: 98, 1840 (Lindenberg 1840). *** Plagiochila teysmannii Sande Lac., Plagiochila Sandei: 6, 1856 (Sande Lacoste 1856c). ** Plagiochila thyoides Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 498, 1885 (Spruce 1885). ** Plagiochila tocarema Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 106, 1864 (Gottsche 1864). ** Plagiochila ulata Inoue et Grolle, Bull. Natl. Sci. Mus. Tokyo, B 1 (3): 91, 1975 (Inoue 1975a). ** Plagiochila undata Sull., Amer. J. Sci. Arts (ser. 2) 1 (1): 73, 1846 (Gray 1846). ** Plagiochila undata subsp. crispata (Gottsche) R.M.Schust., Amer. Midl. Naturalist 63 (1): 122, 1960 (Schuster 1960c). Bas.: Plagiochila crispata Gottsche, Mexik. Leverm.: 71, 1863 (Gottsche 1863). 238 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Plagiochila ungarangana Sande Lac., Syn. hepat. jav.: 10, 1856 [1857] (Sande Lacoste 1856b). *** Plagiochila virginica A.Evans, Prelim. cat. l. W. Virginia: 497, 1892 (Evans 1892a). ** Plagiochila virginica var. caroliniana R.M.Schust., Amer. Midl. Naturalist 63 (1): 15, 1960 (Schuster 1960c). ** Plagiochila virginica var. euryphylla R.M.Schust., Amer. Midl. Naturalist 63 (1): 21, 1960 (Schuster 1960c). *** Plagiochila wightii Nees, Sp. Hepat. (Lindenberg) 2-4: 43, 1840 (Lindenberg 1840). ** Plagiochila wilhelmina Inoue, J. Hattori Bot. Lab. 33: 317, 1970 (Inoue 1970a). sect. Zanteniae (Inoue) Inoue, Gen. Plagiochila SE Asia: 45, 1984 (Inoue 1984b). Bas.: Plagiochila subsect. Zanteniae Inoue, J. Hattori Bot. Lab. 32: 109, 1969 (Inoue 1969). ** Plagiochila zantenii Inoue, J. Hattori Bot. Lab. 32: 107, 1969 (Inoue 1969). * * ** ** * * ** * * ** ** * * ** ** ** Incertae sedis Plagiochila abscendens Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 104, 1864 (Gottsche 1864). Plagiochila aculeata (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 5: 627, 1847 (Gottsche et al. 1847). Bas.: Jungermannia aculeata Hook.f. et Taylor, London J. Bot. 3: 578, 1844 (Hooker and Taylor 1844c). Plagiochila acuta Steph., Bull. Herb. Boissier (sér. 2) 3 (7): 607 (353), 1903 (Stephani 1903e). Plagiochila albertii Steph., Biblioth. Bot. 87 (2): 188, 1916 (Stephani 1916a). Plagiochila aliena Gottsche, Mexik. Leverm.: 22, 1863 (Gottsche 1863). 170 Plagiochila allionii Steph., Sp. Hepat. (Stephani) 6: 120, 1917 (Stephani 1917a). Plagiochila ambigua Lindenb. et Hampe, Linnaea 24 (6): 640, 1851 [1852] (Hampe 1851a). Plagiochila ampliata Steph., Biblioth. Bot. 87 (2): 189, 1916 (Stephani 1916a). Plagiochila andicola Mont. et Gottsche, Ann. Sci. Nat. Bot. (sér. 4) 6: 187, 1856 (Montagne 1856a). Plagiochila andina Steph., Sp. Hepat. (Stephani) 6: 121, 1917 (Stephani 1917a). Plagiochila angolensis Steph., Sp. Hepat. (Stephani) 6: 122, 1917 (Stephani 1917a). Plagiochila angusteoblonga Steph., Biblioth. Bot. 87 (2): 189, 1916 (Stephani 1916a). Plagiochila angustisedens Steph., Bull. Herb. Boissier (sér. 2) 5 (8): 743 (542), 1905 (Stephani 1905i). Plagiochila angustispina Steph., Bull. Herb. Boissier (sér. 2) 2 (8): 671 (208), 1902 (Stephani 1902a). Plagiochila apicalis Gottsche, Mexik. Leverm.: 29, 1863 (Gottsche 1863). 170 Plagiochila aliena is conspeciic with Plagiochila ludoviciana in Schuster (1960c), but others (e.g. Fulford and Sharp 1990, Heinrichs et al. 1998, Dauphin 2005) recognize it. World checklist of hornworts and liverworts 239 ** Plagiochila appalachiana Inoue, J. Hattori Bot. Lab. 40: 415, 1976 (Inoue 1976b). Nom. nov. pro Plagiochila yokogurensis subsp. fragilifolia R.M.Schust., J. Hattori Bot. Lab. 18: 18, 1957 (Schuster 1957c). * Plagiochila arcta S.Winkl., Rev. Bryol. Lichénol. 42 (3): 818, 1976 (Winkler 1976). ** Plagiochila arcuata Lindenb., Sp. Hepat. (Lindenberg) 2-4: 91, 1840 (Lindenberg 1840). ** Plagiochila arenacea Schifn., Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 53, 1964 (Schifner and Arnell 1964). * Plagiochila artsii Pócs, J. Hattori Bot. Lab. 100: 334, 2006 (Pócs 2006a). ** Plagiochila atrovirens Taylor, London J. Bot. 5: 266, 1846 (Taylor 1846a). ** Plagiochila balansae Gottsche, Contr. Étude Plagiochila: 149, 1928 (Dugas 1928). ** Plagiochila baldwinii Austin, Trans. Connecticut Acad. Arts 8 (15): 257, 1891 (Evans 1891). * Plagiochila bamingensis Steph., Bull. Mus. Natl. Hist. Nat. 18 (2): 117, 1912 (Corbière 1912). ** Plagiochila bancroftii Steph., Sp. Hepat. (Stephani) 6: 127, 1917 (Stephani 1917a). *** Plagiochila barbadensis Steph., Bull. Herb. Boissier (sér. 2) 5 (9): 897 (563), 1905 (Stephani 1905g). * Plagiochila barbata Steph., Biblioth. Bot. 87 (2): 190, 1916 (Stephani 1916a). * Plagiochila barbeyi Steph., Biblioth. Bot. 87 (2): 190, 1916 (Stephani 1916a). * Plagiochila batava Steph., Sp. Hepat. (Stephani) 6: 128, 1917 (Stephani 1917a). * Plagiochila berggrenii Steph., Biblioth. Bot. 87 (2): 191, 1916 (Stephani 1916a). ** Plagiochila beskeana Steph., Bull. Herb. Boissier (sér. 2) 2 (10): 863 (232), 1902 (Stephani 1902g). ** Plagiochila bialata Mitt., Fl. vit.: 407, 1871 [1873] (Mitten 1871). * Plagiochila biapiculata Steph., Bull. Herb. Boissier (sér. 2) 5 (9): 891 (557), 1905 (Stephani 1905g). * Plagiochila bicaudata Steph., Sp. Hepat. (Stephani) 6: 130, 1918 (Stephani 1918). ** Plagiochila biciliata Steph., Bull. Herb. Boissier (sér. 2) 3 (6): 529 (333), 1903 (Stephani 1903d). ** Plagiochila bicornis Hampe et Gottsche, Linnaea 25 (3): 338, 1852 [1853] (Hampe and Gottsche 1852). * Plagiochila bidentula Steph., Sp. Hepat. (Stephani) 6: 130, 1918 (Stephani 1918). ** Plagiochila binghamiae A.Evans, Trans. Connecticut Acad. Arts 18 (5): 304, 1914 (Evans 1914c). ** Plagiochila binominata Steph., Sp. Hepat. (Stephani) 6: 131, 1918 (Stephani 1918). ** Plagiochila bitexta Dugas, Contr. Étude Plagiochila: 58, 1928 (Dugas 1928). ** Plagiochila blepharobasis Herzog, Hedwigia 72 (6): 216, 1932 (Herzog 1932c). ** Plagiochila bogotensis Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 98, 1864 (Gottsche 1864). * Plagiochila boliviana Spruce, Mem. Torrey Bot. Club 1 (3): 137, 1890 (Spruce 1890). * Plagiochila borneensis Steph., Sp. Hepat. (Stephani) 6: 132, 1918 (Stephani 1918). 171 171 Plagiochila borneensis is a doubtful taxon. No specimen could be found by Inoue (1984b) or So and Grolle (2000a). 240 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Plagiochila brassii Inoue et Grolle, J. Hattori Bot. Lab. 36: 492, 1972 [1973] (Inoue 1972b). * Plagiochila brevicalycina Lindenb. et Gottsche, Linnaea 20 (3): 322, 1847 (Hampe 1847). ** Plagiochila brunneola Steph., Bull. Herb. Boissier (sér. 2) 4 (2): 164 (416), 1904 (Stephani 1904f). *** Plagiochila bryhnii Steph., Biblioth. Bot. 87 (2): 192, 1916 (Stephani 1916a). ** Plagiochila bunburii Taylor, London J. Bot. 5: 269, 1846 (Taylor 1846a). * Plagiochila byssacea Hampe, Linnaea 20 (3): 326, 1847 (Hampe 1847). ** Plagiochila caducidentata R.M.Schust., Phytologia 39 (4): 247, 1978 (Schuster 1978a). ** Plagiochila caldana Steph., Bull. Herb. Boissier (sér. 2) 2 (10): 879 (248), 1902 (Stephani 1902g). ** Plagiochila callaensis Steph., Sp. Hepat. (Stephani) 6: 136, 1918 (Stephani 1918). * Plagiochila camusii Steph., Biblioth. Bot. 87 (2): 193, 1916 (Stephani 1916a). * Plagiochila capillicaulis Steph., Biblioth. Bot. 87 (2): 193, 1916 (Stephani 1916a). ** Plagiochila caribbeania R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 412, 1980 (Schuster 1980c). * Plagiochila cava Steph., Bull. Herb. Boissier (sér. 2) 4 (12): 1213 (501), 1904 (Stephani 1904h). ** Plagiochila ceylanica Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 98, 1860 [1861] (Mitten 1860c). ** Plagiochila chacoënsis Herzog, Feddes Repert. Spec. Nov. Regni Veg. 55 (1): 7, 1952 (Herzog 1952h). ** Plagiochila chiloënsis Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 6): 27, 1900 (Stephani 1900b). ** Plagiochila chinantlana Gottsche, Mexik. Leverm.: 12, 1863 (Gottsche 1863). * Plagiochila chiovendae Gola, Ann. Bot. (Rome) 13 (1): 67, 1914 (Gola 1914a). * Plagiochila cinchonae Steph., Bull. Herb. Boissier (sér. 2) 5 (10): 920 (570), 1905 (Stephani 1905f). * Plagiochila circumvoluta Gerola, Lav. Bot. Ist. Bot. Univ. Padova 12: 475, 1947 (Gerola 1947). *** Plagiochila cleeii Inoue, Stud. Cryptog. S. Peru: 95, 1987 (Inoue 1987c). * Plagiochila concava Nees, Sp. Hepat. (Lindenberg) 2-4: 70, 1840 (Lindenberg 1840). ** Plagiochila conduplicata Steph., Sp. Hepat. (Stephani) 6: 139, 1918 (Stephani 1918). * Plagiochila conferta Steph., Sp. Hepat. (Stephani) 6: 139, 1918 (Stephani 1918). ** Plagiochila confertissima Steph., Bull. Herb. Boissier (sér. 2) 5 (2): 182 (510), 1905 (Stephani 1905d). ** Plagiochila connata Lindenb. et Gottsche, Syn. Hepat. 5: 645, 1847 (Gottsche et al. 1847). ** Plagiochila contorta Lindenb. et Hampe, Linnaea 24 (3): 301, 1851 [1852] (Hampe 1851b). ** Plagiochila convoluta Steph., Sp. Hepat. (Stephani) 6: 141, 1918 (Stephani 1918). ** Plagiochila convolutifolia Steph., Sp. Hepat. (Stephani) 6: 142, 1918 (Stephani 1918). World checklist of hornworts and liverworts 241 *** Plagiochila corymbulosa Pearson, Forh. Vidensk.-Selsk. Kristiania 1887 (9): 14, 1887 (Pearson 1887b). ** Plagiochila costariensis Horik., Acta Phytotax. Geobot. 13: 213, 1943 (Horikawa 1943). Nom. nov. pro Plagiochila pinnata Steph., Bull. Herb. Boissier (sér. 2) 5 (8): 749 (548), 1905 (Stephani 1905i), nom. illeg. ** Plagiochila crispabilis Lindenb., Sp. Hepat. (Lindenberg) 1: 15, 1839 (Lindenberg 1839). * Plagiochila crispabilis var. minima Lindenb., Sp. Hepat. (Lindenberg) 5: 155, 1843 (Lindenberg 1843). ** Plagiochila cristophylla Steph., Bull. Herb. Boissier (sér. 2) 3 (2): 117 (297), 1903 (Stephani 1903b). *** Plagiochila cuatrecasii H.Rob., Bryologist 70 (1): 47, 1967 (Robinson 1967). ** Plagiochila cubensis Steph., Sp. Hepat. (Stephani) 6: 143, 1918 (Stephani 1918). ** Plagiochila cucullata Lindenb. et Gottsche, Syn. Hepat. 5: 642, 1847 (Gottsche et al. 1847). * Plagiochila cuervina Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 96, 1864 (Gottsche 1864). ** Plagiochila delapsa Inoue, Beih. Nova Hedwigia 90: 171, 1988 (Inoue 1988a). * Plagiochila delognei Steph., Biblioth. Bot. 87 (2): 196, 1916 (Stephani 1916a). ** Plagiochila denigrata Inoue, Willdenowia 18 (2): 557, 1989 (Inoue 1989b). ** Plagiochila densa Herzog, Hedwigia 72 (6): 222, 1932 (Herzog 1932c). ** Plagiochila densilora Herzog, Hedwigia 72 (6): 226, 1932 (Herzog 1932c). ** Plagiochila densiramosa Steph., Biblioth. Bot. 87 (2): 196, 1916 (Stephani 1916a). ** Plagiochila deppeana Steph., Bull. Herb. Boissier (sér. 2) 5 (9): 886 (552), 1905 (Stephani 1905g). ** Plagiochila desciscens Steph., Bull. Herb. Boissier (sér. 2) 2 (10): 867 (236), 1902 (Stephani 1902g). * Plagiochila dilatata Steph., Bull. Herb. Boissier (sér. 2) 5 (9): 887 (553), 1905 (Stephani 1905g). * Plagiochila distans Colenso, Trans. & Proc. New Zealand Inst. 19: 283, 1887 (Colenso 1887). 172 ** Plagiochila divaricata Lindenb., Sp. Hepat. (Lindenberg) 5: 147, 1843 (Lindenberg 1843). * Plagiochila doerleri Steph., Biblioth. Bot. 87 (2): 196, 1916 (Stephani 1916a). ** Plagiochila dolichoblasta Herzog, Trans. Brit. Bryol. Soc. 1 (4): 286, 1950 (Herzog 1950a). ** Plagiochila dussiana Steph., Symb. Antill. (Urban) 3 (2): 277, 1902 (Stephani 1902e). ** Plagiochila echinata R.M.Schust., Amer. Midl. Naturalist 62 (2): 341, 1959 (Schuster 1959b). ** Plagiochila echinella Gottsche, Ann. Sci. Nat. Bot. (sér. 4) 8: 332, 1857 (Gottsche 1857). * Plagiochila ecuadorensis Steph., Sp. Hepat. (Stephani) 6: 149, 1918 (Stephani 1918). 172 Plagiochila distans is a doubtful taxon. he type specimen could not be found (Hodgson 1944, Hamlin 1972). 242 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) * Plagiochila efuseintricata Steph., Biblioth. Bot. 87 (2): 197, 1916 (Stephani 1916a). ** Plagiochila eggersii Inoue, Bull. Natl. Sci. Mus. Tokyo, B 14 (4): 138, 1988 (Inoue 1988b). ** Plagiochila ekmanii S.W.Arnell, Bryologist 59 (4): 274, 1956 (Arnell 1956c). * Plagiochila electa Steph., Bull. Herb. Boissier (sér. 2) 5 (8): 739 (538), 1905 (Stephani 1905i). * Plagiochila elegantula Herzog, Feddes Repert. Spec. Nov. Regni Veg. 57 (1/2): 161, 1955 (Herzog 1955). * Plagiochila emarginatobidentula Steph., Biblioth. Bot. 87 (2): 197, 1916 (Stephani 1916a). * Plagiochila erythraeae Herzog, Hedwigia 78 (3/4): 224, 1938 (Herzog 1938d). ** Plagiochila estrellensis Herzog, Repert. Spec. Nov. Regni Veg. 21 (1/7): 23, 1925 (Herzog 1925a). ** Plagiochila eurydictyon Herzog, Hedwigia 72 (6): 207, 1932 (Herzog 1932c). * Plagiochila excisa Steph., Sp. Hepat. (Stephani) 6: 153, 1918 (Stephani 1918). ** Plagiochila exesa Lindenb. et Gottsche, Syn. Hepat. 5: 629, 1847 (Gottsche et al. 1847). * Plagiochila exilis Colenso, Trans. & Proc. New Zealand Inst. 19: 282, 1887 (Colenso 1887). 173 ** Plagiochila expansa Gottsche, Mexik. Leverm.: 18, 1863 (Gottsche 1863). ** Plagiochila facallonia Steph., Bull. Herb. Boissier (sér. 2) 2 (10): 871 (240), 1902 (Stephani 1902g). * Plagiochila falcato-oblonga Steph., Biblioth. Bot. 87 (2): 198, 1916 (Stephani 1916a). ** Plagiochila fallax Lindenb. et Hampe, Linnaea 24 (3): 300, 1851 [1852] (Hampe 1851b). ** Plagiochila faxinensis Schifn., Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 60, 1964 (Schifner and Arnell 1964). ** Plagiochila fendleri Mont., Ann. Sci. Nat. Bot. (sér. 4) 6: 198, 1856 (Montagne 1856a). * Plagiochila ilicicola Steph., Bull. Herb. Boissier (sér. 2) 4 (4): 351 (427), 1904 (Stephani 1904e). ** Plagiochila issidentoides Taylor, London J. Bot. 5: 264, 1846 (Taylor 1846a). * Plagiochila labellifrons Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 488, 1885 (Spruce 1885). ** Plagiochila lava Steph., Sp. Hepat. (Stephani) 6: 156, 1918 (Stephani 1918). ** Plagiochila lavescens (Gottsche, Lindenb. et Nees) Gottsche, Mexik. Leverm.: 52, 1863 (Gottsche 1863). Bas.: Plagiochila guilleminiana β lavescens Gottsche, Lindenb. et Nees, Syn. Hepat. 5: 644, 1847 (Gottsche et al. 1847). * Plagiochila lavorufescens Steph., Biblioth. Bot. 87 (2): 199, 1916 (Stephani 1916a). ** Plagiochila footei A.Evans, Trans. Connecticut Acad. Arts 18 (5): 306, 1914 (Evans 1914c). * Plagiochila formosa Nees, Contr. Étude Plagiochila: 147, 1928 (Dugas 1928). 173 Plagiochila exilis is a doubtful taxon. he type specimen could not be found (Hodgson 1944, Hamlin 1972). World checklist of hornworts and liverworts 243 ** Plagiochila fragmentata R.M.Schust., Phytologia 45 (5): 421, 1980 (Schuster 1980b). * Plagiochila frausa Gottsche, Mexik. Leverm.: 66, 1863 (Gottsche 1863). * Plagiochila frausa var. boliviana Spruce, Mem. Torrey Bot. Club 1 (3): 134, 1890 (Spruce 1890). ** Plagiochila frayjorgensis Hässel, Bol. Soc. Argent. Bot. 22 (1/4): 103, 1983 (Hässel 1983). Nom. nov. pro Plagiochila modesta Herzog, Rev. Bryol. Lichénol. 23 (1/2): 34, 1954 (Herzog 1954), nom. illeg. * Plagiochila gafatensis Gottsche ex Schweinf., Beitr. Fl. Aethiop.: 227, 1866 [1867] (Schweinfurth 1866). ** Plagiochila gaudichaudii Mont. et Gottsche, Ann. Sci. Nat. Bot. (sér. 4) 6: 193, 1856 (Montagne 1856a). ** Plagiochila geniculata Lindenb., Sp. Hepat. (Lindenberg) 5: 131, 1843 (Lindenberg 1843). * Plagiochila geppii Steph., Biblioth. Bot. 87 (2): 199, 1916 (Stephani 1916a). ** Plagiochila germana Gottsche, Mexik. Leverm.: 34, 1863 (Gottsche 1863). ** Plagiochila germanii Steph., Bull. Herb. Boissier (sér. 2) 5 (10): 938 (588), 1905 (Stephani 1905f). * Plagiochila gibbilora Steph., Bull. Herb. Boissier (sér. 2) 4 (6): 590 (441), 1904 (Stephani 1904b). ** Plagiochila gittinsii Inoue, Bull. Natl. Sci. Mus. Tokyo, B 3 (4): 139, 1977 (Inoue 1977a). * Plagiochila glauca Carl, Ann. Bryol., Suppl. 2: 129, 1931 (Carl 1931b). 174 * Plagiochila gracilicaulis Spruce, Mem. Torrey Bot. Club 1 (3): 132, 1890 (Spruce 1890). ** Plagiochila gracillima Austin, Trans. Connecticut Acad. Arts 8 (15): 256, 1891 (Evans 1891). * Plagiochila granatensis Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 111, 1864 (Gottsche 1864). ** Plagiochila granditexta Steph., Bull. Herb. Boissier (sér. 2) 4 (2): 165 (417), 1904 (Stephani 1904f). ** Plagiochila grateloupii Mont., Ann. Sci. Nat. Bot. (sér. 4) 6: 188, 1856 (Montagne 1856a). ** Plagiochila guatemalensis Steph., Sp. Hepat. (Stephani) 6: 163, 1918 (Stephani 1918). ** Plagiochila guttisquama Inoue et Grolle, Bull. Natl. Sci. Mus. Tokyo, B 5 (1): 29, 1979 (Inoue 1979c). ** Plagiochila gymnocalyx Inoue, Trop. Bryol. 1: 34, 1989 (Gradstein and Florschützde Waard 1989). ** Plagiochila haeseliae Inoue, Stud. Cryptog. S. Chile: 97, 1984 (Inoue 1984a). * Plagiochila hans-meyeri Steph., Sp. Hepat. (Stephani) 6: 164, 1918 (Stephani 1918). ** Plagiochila harlingii S.W.Arnell, Svensk Bot. Tidskr. 56 (2): 346, 1962 (Arnell 1962b). 174 Plagiochila glauca is a doubtful taxon. Hässel and Rubies (2009) could not ind any type material. 244 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Plagiochila haumanii Herzog, Repert. Spec. Nov. Regni Veg. 41 (14/25): 285, 1937 (Herzog 1937). ** Plagiochila hawaica Steph., Bull. Herb. Boissier (sér. 2) 3 (7): 598 (344), 1903 (Stephani 1903e). ** Plagiochila heteracantha Steph., Sp. Hepat. (Stephani) 6: 166, 1918 (Stephani 1918). * Plagiochila heterofolia Steph., Biblioth. Bot. 87 (2): 200, 1916 (Stephani 1916a). * Plagiochila hieronymii Steph., Biblioth. Bot. 87 (2): 201, 1916 (Stephani 1916a). ** Plagiochila hiroshiana Pócs, J. Hattori Bot. Lab. 100: 335, 2006 (Pócs 2006a). ** Plagiochila hoehnei Herzog, Hedwigia 72 (6): 220, 1932 (Herzog 1932c). * Plagiochila holstii Steph., Hedwigia 45 (4): 214, 1906 (Stephani 1906b). Nom. nov. pro Plagiochila prostrata Steph., Bull. Herb. Boissier (sér. 2) 4 (2): 168 (420), 1904 (Stephani 1904f), nom. illeg. ** Plagiochila horrida Gottsche, Mexik. Leverm.: 74, 1863 (Gottsche 1863). * Plagiochila huatuscana Gottsche, Mexik. Leverm.: 24, 1863 (Gottsche 1863). ** Plagiochila huerlimannii Inoue, J. Hattori Bot. Lab. 33: 307, 1970 (Inoue 1970b). * Plagiochila humboldtiana Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 112, 1864 (Gottsche 1864). ** Plagiochila hyalina Lindenb., Syn. Hepat. 5: 640, 1847 (Gottsche et al. 1847). * Plagiochila incisa Dugas, Contr. Étude Plagiochila: 112, 1928 (Dugas 1928). ** Plagiochila inlata Steph., Bull. Herb. Boissier (sér. 2) 3 (11): 961 (376), 1903 (Stephani 1903a). * Plagiochila informifolia Steph., Biblioth. Bot. 87 (2): 202, 1916 (Stephani 1916a). ** Plagiochila infuscata Mitt., J. Linn. Soc., Bot. 15 (82): 63, 1876 (Mitten 1876a). * Plagiochila injasutiensis S.W.Arnell, Bot. Not. 110: 404, 1957 (Arnell 1957d). ** Plagiochila inouei Grolle, J. Bryol. 10 (3): 269, 1979 (Grolle 1979b). Nom. nov. pro Plagiochila nudiuscula Inoue, Bull. Natl. Sci. Mus. Tokyo, B 3 (2): 45, 1977 (Inoue 1977c), nom. illeg. ** Plagiochila insularia Mitt., St. Helena: 366, 1875 (Mitten 1875). ** Plagiochila intertexta Hook.f. et Taylor, London J. Bot. 5: 267, 1846 (Taylor 1846a). * Plagiochila inuensis Steph., Biblioth. Bot. 87 (2): 202, 1916 (Stephani 1916a). ** Plagiochila irregularis Gottsche, Mexik. Leverm.: 14, 1863 (Gottsche 1863). ** Plagiochila itatiajensis Steph., Bull. Herb. Boissier (sér. 2) 2 (10): 874 (243), 1902 (Stephani 1902g). ** Plagiochila jamaicensis Lindenb. et Hampe, Linnaea 24 (3): 302, 1851 [1852] (Hampe 1851b). ** Plagiochila jaramillii H.Rob., Bryologist 70 (1): 48, 1967 (Robinson 1967). * Plagiochila karstenii Steph., Hedwigia 45 (4): 214, 1906 (Stephani 1906b). Nom. nov. pro Plagiochila patentispina Steph., Bull. Herb. Boissier (sér. 2) 4 (1): 28 (400), 1904 (Stephani 1904g), nom. illeg. 175 * Plagiochila kaulfussiana Steph., Sp. Hepat. (Stephani) 6: 172, 1918 (Stephani 1918). ** Plagiochila keckiana Steph., Bull. Herb. Boissier (sér. 2) 5 (4): 358 (526), 1905 (Stephani 1905j). 175 Plagiochila karstenii is possibly conspeciic with Plagiochila bantamensis (So and Grolle 2000a). World checklist of hornworts and liverworts 245 ** Plagiochila kerneriana S.W.Arnell, Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 60, 1964 (Schifner and Arnell 1964). ** Plagiochila koponenii Inoue et Piippo, Ann. Bot. Fenn. 26 (2): 216, 1989 (Piippo 1989b). * Plagiochila lacerifolia Steph., Biblioth. Bot. 87 (2): 202, 1916 (Stephani 1916a). * Plagiochila lachenaudii Steph., Biblioth. Bot. 87 (2): 202, 1916 (Stephani 1916a). * Plagiochila laciniosa Dugas, Contr. Étude Plagiochila: 68, 1928 (Dugas 1928). * Plagiochila laevifolia Gola, Ann. Bot. (Rome) 6 (2): 273, 1907 (Gola 1907). ** Plagiochila lansbergii Gottsche, Mexik. Leverm.: 13, 1863 (Gottsche 1863). ** Plagiochila latitrigona Schifn., Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 48, 1964 (Schifner and Arnell 1964). * Plagiochila laxiramea Steph., Biblioth. Bot. 87 (2): 203, 1916 (Stephani 1916a). * Plagiochila lecontei Steph., Bull. Herb. Boissier (sér. 2) 4 (2): 165 (417), 1904 (Stephani 1904f). 176 * Plagiochila ledermanniana Beauverd, Sp. Hepat. (Stephani) 6: 572, 1924 (Stephani 1924). Nom. nov. pro Plagiochila cucullifolia Steph., Sp. Hepat. (Stephani) 6: 243, 1922 (Stephani 1922), nom. illeg. 177 * Plagiochila ledieui Steph., Bull. Herb. Boissier (sér. 2) 4 (2): 161 (413), 1904 (Stephani 1904f). 178 ** Plagiochila lehmanniana S.W.Arnell, Svensk Bot. Tidskr. 55 (1): 205, 1961 (Arnell 1961). ** Plagiochila liebmanniana Lehm. et Lindenb., Sp. Hepat. (Lindenberg) 2-4: 97, 1840 (Lindenberg 1840). * Plagiochila lignicola Spruce, Mem. Torrey Bot. Club 1 (3): 135, 1890 (Spruce 1890). * Plagiochila lindaui Steph., Biblioth. Bot. 87 (2): 204, 1916 (Stephani 1916a). * Plagiochila lindigiana Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 105, 1864 (Gottsche 1864). ** Plagiochila lingua Steph., Bull. Herb. Boissier (sér. 2) 2 (8): 677 (214), 1902 (Stephani 1902a). * Plagiochila longiissa Steph., Bull. Herb. Boissier (sér. 2) 5 (9): 891 (557), 1905 (Stephani 1905g). * Plagiochila lotsyana Steph., Biblioth. Bot. 87 (2): 205, 1916 (Stephani 1916a). * Plagiochila luteola Steph., Bull. Herb. Boissier (sér. 2) 5 (2): 175 (503), 1905 (Stephani 1905d). ** Plagiochila lutescens Steph., Sp. Hepat. (Stephani) 6: 179, 1921 (Stephani 1921). ** Plagiochila luzonensis Grolle et M.L.So, Bryologist 102 (1): 69, 1999 (Grolle and So 1999a). 176 Plagiochila lecontei is allied to and perhaps conspeciic with Plagiochila ledieui and Plagiochila multilora (Jones 1962). 177 Plagiochila ledermanniana is a doubtful taxon. he type specimen was burned in B (Piippo 1989b). 178 Plagiochila ledieui is allied to and perhaps conspeciic with Plagiochila lecontei and Plagiochila multilora (Jones 1962). 246 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) * Plagiochila macra Taylor, London J. Bot. 7: 198, 1848 (Taylor 1848a). ** Plagiochila macrifolia Taylor, London J. Bot. 5: 270, 1846 (Taylor 1846a). * Plagiochila macrifolia var. angustifolia Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 470, 1885 (Spruce 1885). *** Plagiochila martiana (Nees) Lindenb., Sp. Hepat. (Lindenberg) 1: 12, 1839 (Lindenberg 1839). Bas.: Jungermannia martiana Nees, Linnaea 6 (4): 617, 1831 (Nees 1831). * Plagiochila matanga Steph., Sp. Hepat. (Stephani) 6: 183, 1921 (Stephani 1921). ** Plagiochila maunakeana Steph., Sp. Hepat. (Stephani) 6: 186, 1921 (Stephani 1921). ** Plagiochila maximiliana Gottsche, Mexik. Leverm.: 18, 1863 (Gottsche 1863). ** Plagiochila meghalayensis K.K.Rawat et S.C.Srivast., Geophytology 35 (1/2): 49, 2005 (Rawat and Srivastava 2005). ** Plagiochila meridana Gottsche, Ann. Sci. Nat. Bot. (sér. 4) 8: 328, 1857 (Gottsche 1857). ** Plagiochila microdonta Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 97, 1860 [1861] (Mitten 1860c). * Plagiochila mildbreadiana Beauverd, Sp. Hepat. (Stephani) 6: 571, 1924 (Stephani 1924). Nom. nov. pro Plagiochila andongensis Steph., Sp. Hepat. (Stephani) 6: 122, 1917 (Stephani 1917a), nom. illeg. ** Plagiochila minarum Herzog, Hedwigia 72 (6): 207, 1932 (Herzog 1932c). ** Plagiochila minutiretis Reimers, Repert. Spec. Nov. Regni Veg. 21 (8/20): 264, 1925 (Reimers 1925). ** Plagiochila miqueliana Lehm. et Lindenb., Sp. Hepat. (Lindenberg) 2-4: 95, 1840 (Lindenberg 1840). ** Plagiochila molliuscula Inoue, Ann. Bot. Fenn. 13 (3): 134, 1976 (Engel 1976a). ** Plagiochila mollusca Lehm., Nov. Stirp. Pug. 10: 4, 1857 (Lehmann 1857). ** Plagiochila moniliformis R.M.Schust., Phytologia 39 (4): 247, 1978 (Schuster 1978a). ** Plagiochila muelleriana Gottsche, Mexik. Leverm.: 38, 1863 (Gottsche 1863). * Plagiochila multilora Steph., Planzenw. Ost-Afrikas C: 64, 1895 (Stephani 1895d). 179 ** Plagiochila mutila Gottsche, Ann. Sci. Nat. Bot. (sér. 4) 8: 327, 1857 (Gottsche 1857). * Plagiochila naranjoënsis Steph., Bull. Herb. Boissier (sér. 2) 2 (8): 687 (224), 1902 (Stephani 1902a). ** Plagiochila neesiana Lindenb., Sp. Hepat. (Lindenberg) 2-4: 71, 1840 (Lindenberg 1840). * Plagiochila neglecta Steph., Bull. Herb. Boissier (sér. 2) 5 (4): 351 (519), 1905 (Stephani 1905j). * Plagiochila negrensis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 466, 1885 (Spruce 1885). * Plagiochila negrii Gola, Ann. Bot. (Rome) 13 (1): 66, 1914 (Gola 1914a). 179 Plagiochila multilora is allied to and perhaps conspeciic with Plagiochila lecontei and Plagiochila ledieui (Jones 1962). World checklist of hornworts and liverworts * * * ** ** * ** ** ** * * * ** ** ** * ** * * ** ** ** ** * ** 247 Plagiochila nigricaulis Steph., Biblioth. Bot. 87 (2): 206, 1916 (Stephani 1916a). Plagiochila notha Steph., Bull. Herb. Boissier (sér. 2) 5 (8): 737 (536), 1905 (Stephani 1905i). Plagiochila nova Steph., Sp. Hepat. (Stephani) 6: 189, 1921 (Stephani 1921). Plagiochila nudifolia (Steph.) Grolle, Feddes Repert. 82 (1): 89, 1971 (Grolle 1971b). Bas.: Jamesoniella nudifolia Steph., Biblioth. Bot. 87 (2): 184, 1916 (Stephani 1916a). Plagiochila nutans Steph., Bull. Herb. Boissier (sér. 2) 3 (11): 960 (375), 1903 (Stephani 1903a). Plagiochila obliquetruncata Steph., Biblioth. Bot. 87 (2): 207, 1916 (Stephani 1916a). Plagiochila oblonga Inoue, Bull. Natl. Sci. Mus. Tokyo (n.ser.) 8 (3): 398, 1965 (Inoue 1965a). Plagiochila oerstediana Lindenb. et Hampe, Linnaea 24 (3): 301, 1851 [1852] (Hampe 1851b). Plagiochila olivacea Steph., Bull. Herb. Boissier (sér. 2) 5 (2): 190 (518), 1905 (Stephani 1905d). Plagiochila ornata Wilson ex Lindenb., Sp. Hepat. (Lindenberg) 5: 164, 1843 (Lindenberg 1843). Plagiochila ovato-obconica Steph., Sp. Hepat. (Stephani) 6: 191, 1921 (Stephani 1921). Plagiochila ovifolia Steph., Sp. Hepat. (Stephani) 6: 191, 1921 (Stephani 1921). Plagiochila palangiensis S.C.Srivast., K.K.Rawat et P.K.Verma, Natl. Acad. Sci. Lett. 29 (7/8): 267, 2006 (Srivastava et al. 2006). Plagiochila panamensis Inoue, Bull. Natl. Sci. Mus. Tokyo, B 15 (3): 94, 1989 (Inoue 1989a). Plagiochila paramicola Herzog, Beih. Bot. Centralbl. 61B (3): 563, 1942 (Herzog 1942d). Plagiochila parvitexta Steph., Bull. Herb. Boissier (sér. 2) 2 (8): 674 (211), 1902 (Stephani 1902a). Plagiochila parvivittata Inoue, Bull. Natl. Sci. Mus. Tokyo, B 13 (2): 50, 1987 (Inoue 1987b). Plagiochila parvula Steph., Akad. Wiss. Wien, Math.-Naturwiss. Kl., Denkschr. 88: 726, 1913 (Stephani 1913b). Plagiochila pastasensis Steph., Sp. Hepat. (Stephani) 6: 197, 1921 (Stephani 1921). Plagiochila patentispina Steph., Bull. Herb. Boissier (sér. 2) 2 (10): 877 (246), 1902 (Stephani 1902g). Plagiochila patuloides Schifn., Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 58, 1964 (Schifner and Arnell 1964). Plagiochila paucidentata Mont. et Gottsche, Ann. Sci. Nat. Bot. (sér. 4) 6: 197, 1856 (Montagne 1856a). Plagiochila paupercula Gottsche, Mexik. Leverm.: 22, 1863 (Gottsche 1863). Plagiochila pearceana Steph., Bull. Herb. Boissier (sér. 2) 5 (8): 742 (541), 1905 (Stephani 1905i). Plagiochila pellucida Lindenb. et Gottsche, Linnaea 20 (3): 321, 1847 (Hampe 1847). 248 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Plagiochila perrottetiana Mont. et Gottsche, Ann. Sci. Nat. Bot. (sér. 4) 6: 195, 1856 (Montagne 1856a). ** Plagiochila perrottetiana var. minor Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 101, 1864 (Gottsche 1864). ** Plagiochila pilifera Steph., Sp. Hepat. (Stephani) 6: 198, 1921 (Stephani 1921). ** Plagiochila pinnatidens Steph., Sp. Hepat. (Stephani) 6: 199, 1921 (Stephani 1921). ** Plagiochila pittieri Steph., Bull. Herb. Boissier (sér. 2) 2 (8): 673 (210), 1902 (Stephani 1902a). ** Plagiochila platyphylla Herzog, Hedwigia 72 (6): 223, 1932 (Herzog 1932c). ** Plagiochila polopolensis Herzog, Hedwigia 67 (6): 261, 1927 (Herzog 1927). * Plagiochila prostrata Steph., Bull. Herb. Boissier (sér. 2) 2 (8): 668 (205), 1902 (Stephani 1902a). * Plagiochila pseudopatula Herzog, Hedwigia 72 (6): 227, 1932 (Herzog 1932c). ** Plagiochila pseudoradicans Herzog, Repert. Spec. Nov. Regni Veg. 21 (1/7): 24, 1925 (Herzog 1925a). ** Plagiochila ptychanthoidea Steph., Bull. Herb. Boissier (sér. 2) 3 (2): 121 (301), 1903 (Stephani 1903b). ** Plagiochila pulchella Steph., Bull. Herb. Boissier (sér. 2) 2 (10): 871 (240), 1902 (Stephani 1902g). ** Plagiochila pulvinata Steph., Bull. Herb. Boissier (sér. 2) 3 (6): 526 (330), 1903 (Stephani 1903d). ** Plagiochila ratkowskiana Inoue, Brunonia 3 (1): 141, 1980 (Inoue 1980). ** Plagiochila regeliana Steph., Bull. Herb. Boissier (sér. 2) 2 (8): 675 (212), 1902 (Stephani 1902a). ** Plagiochila remyana Steph., Bull. Herb. Boissier (sér. 2) 3 (11): 963 (378), 1903 (Stephani 1903a). *** Plagiochila revolvens Mitt., Hooker’s J. Bot. Kew Gard. Misc. 3: 358, 1851 (Mitten 1851). ** Plagiochila rigidula Lindenb. et Gottsche, Linnaea 20 (3): 323, 1847 (Hampe 1847). ** Plagiochila rosana Steph., Bull. Herb. Boissier (sér. 2) 5 (9): 892 (558), 1905 (Stephani 1905g). * Plagiochila rubricaulis Steph., Bot. Jahrb. Syst. 20 (3): 311, 1895 (Stephani 1895a). * Plagiochila ruifolia Steph., Biblioth. Bot. 87 (2): 209, 1916 (Stephani 1916a). ** Plagiochila rufoviridis Spruce, Mem. Torrey Bot. Club 1 (3): 136, 1890 (Spruce 1890). * Plagiochila rusbyi Spruce, Mem. Torrey Bot. Club 1 (3): 135, 1890 (Spruce 1890). * Plagiochila sabensis Steph., Sp. Hepat. (Stephani) 6: 214, 1921 (Stephani 1921). ** Plagiochila sachapatensis Steph., Bull. Herb. Boissier (sér. 2) 2 (8): 679 (216), 1902 (Stephani 1902a). ** Plagiochila salazariae Inoue, Bull. Natl. Sci. Mus. Tokyo, B 15 (3): 91, 1989 (Inoue 1989a). ** Plagiochila saltuensis Spruce ex Steph., Bull. Herb. Boissier (sér. 2) 5 (10): 927 (577), 1905 (Stephani 1905f). World checklist of hornworts and liverworts 249 ** Plagiochila saltuensis var. spinosissima Herzog, Feddes Repert. Spec. Nov. Regni Veg. 57 (1/2): 163, 1955 (Herzog 1955). * Plagiochila samoana Steph., Akad. Wiss. Wien, Math.-Naturwiss. Kl., Denkschr. 85: 197, 1910 (Stephani 1910a). ** Plagiochila sarmentosa (Lehm. et Lindenb.) Lindenb., Sp. Hepat. (Lindenberg) 2-4: 86, 1840 (Lindenberg 1840). Bas.: Jungermannia sarmentosa Lehm. et Lindenb., Linnaea 9 (4): 427, 1835 (Lehmann 1835). * Plagiochila schifneri Steph., Biblioth. Bot. 87 (2): 209, 1916 (Stephani 1916a). * Plagiochila schinzei Steph., Biblioth. Bot. 87 (2): 209, 1916 (Stephani 1916a). * Plagiochila schmidtii Steph., Biblioth. Bot. 87 (2): 209, 1916 (Stephani 1916a). * Plagiochila schraderbergii Steph., Sp. Hepat. (Stephani) 6: 244, 1922 (Stephani 1922). 180 ** Plagiochila schubertiana Steph., Sp. Hepat. (Stephani) 6: 208, 1921 (Stephani 1921). ** Plagiochila schusteri Inoue et Grolle, J. Hattori Bot. Lab. 33: 326, 1970 (Inoue 1970a). ** Plagiochila scissifolia Steph., Bull. Herb. Boissier (sér. 2) 2 (10): 865 (234), 1902 (Stephani 1902g). * Plagiochila scotica Macvicar ex Steph., Sp. Hepat. (Stephani) 6: 209, 1921 (Stephani 1921). 181 ** Plagiochila semiamplexicaulis Steph., Bull. Herb. Boissier (sér. 2) 5 (10): 936 (586), 1905 (Stephani 1905f). ** Plagiochila semiermis Dugas, Contr. Étude Plagiochila: 66, 1928 (Dugas 1928). ** Plagiochila silvatica Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 108, 1864 (Gottsche 1864). * Plagiochila similis Steph., Biblioth. Bot. 87 (2): 210, 1916 (Stephani 1916a). ** Plagiochila sisparensis Steph., Sp. Hepat. (Stephani) 6: 207, 1921 (Stephani 1921). * Plagiochila slateri Steph., Biblioth. Bot. 87 (2): 211, 1916 (Stephani 1916a). ** Plagiochila solitaria Gottsche, Sp. Hepat. (Stephani) 6: 225, 1921 (Stephani 1921). *** Plagiochila spinifera Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 30 (5): 113, 1873 (Ångström 1873). * Plagiochila sprucei Steph., Bull. Herb. Boissier (sér. 2) 2 (10): 860 (229), 1902 (Stephani 1902g). * Plagiochila staudtiana Steph., Bull. Herb. Boissier (sér. 2) 4 (4): 352 (428), 1904 (Stephani 1904e). ** Plagiochila stipata Steph., Sp. Hepat. (Stephani) 6: 209, 1921 (Stephani 1921). * Plagiochila stolzii Steph., Sp. Hepat. (Stephani) 6: 244, 1922 (Stephani 1922). ** Plagiochila subcontigua Herzog, Hedwigia 72 (6): 230, 1932 (Herzog 1932c). * Plagiochila subconvoluta Gottsche, Mexik. Leverm.: 24, 1863 (Gottsche 1863). * Plagiochila subdenudata Steph., Bull. Herb. Boissier (sér. 2) 2 (8): 673 (210), 1902 (Stephani 1902a). 180 Plagiochila schraderbergii is a doubtful taxon. he type specimen was burned in B (Piippo 1989b). 181 Plagiochila scotica was described from Europe, but it has neither been recognized in any recent European study nor synonymized. 250 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Plagiochila subedentata Steph., Bull. Herb. Boissier (sér. 2) 2 (8): 679 (216), 1902 (Stephani 1902a). ** Plagiochila subfragilis Inoue, Stud. Cryptog. S. Peru: 98, 1987 (Inoue 1987c). * Plagiochila subhyalina Steph., Biblioth. Bot. 87 (2): 212, 1916 (Stephani 1916a). ** Plagiochila subligulata Steph., Sp. Hepat. (Stephani) 6: 224, 1921 (Stephani 1921). ** Plagiochila sublyallii M.L.So, New Zealand J. Bot. 39 (1): 109, 2001 (So 2001b). * Plagiochila subrara Herzog, Hedwigia 74 (2): 87, 1934 (Herzog 1934a). * Plagiochila subrotundifolia Steph., Bull. Herb. Boissier (sér. 2) 5 (10): 935 (585), 1905 (Stephani 1905f). ** Plagiochila subundulata Lindenb., Sp. Hepat. (Lindenberg) 5: 137, 1843 (Lindenberg 1843). * Plagiochila suringarii Steph., Sp. Hepat. (Stephani) 6: 208, 1921 (Stephani 1921). ** Plagiochila sylvicultrix Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 486, 1885 (Spruce 1885). * Plagiochila symmetrica Steph., Bull. Herb. Boissier (sér. 2) 2 (10): 876 (245), 1902 (Stephani 1902g). ** Plagiochila tabinana Gottsche, Mexik. Leverm.: 46, 1863 (Gottsche 1863). ** Plagiochila talinayi S.W.Arnell, Svensk Bot. Tidskr. 50 (2): 312, 1956 (Arnell 1956d). ** Plagiochila tambillensis Loitl., Diagn. pl. nov.: 22, 1894 (Szyszylowicz 1894). ** Plagiochila tarapotensis Steph., Bull. Herb. Boissier (sér. 2) 2 (10): 863 (232), 1902 (Stephani 1902g). * Plagiochila tenera Steph., Sp. Hepat. (Stephani) 6: 231, 1921 (Stephani 1921). ** Plagiochila tenuis Lindenb., Sp. Hepat. (Lindenberg) 2-4: 50, 1840 (Lindenberg 1840). * Plagiochila tenuispica Steph., Sp. Hepat. (Stephani) 6: 234, 1921 (Stephani 1921). ** Plagiochila thamniopsis Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: cc, 1889 [1890] (Spruce 1889). * Plagiochila thollonii Steph., Bull. Herb. Boissier (sér. 2) 2 (12): 980 (270), 1902 (Stephani 1902h). ** Plagiochila thrausta Inoue et Grolle, Bull. Natl. Sci. Mus. Tokyo, B 5 (1): 34, 1979 (Inoue 1979c). * Plagiochila tonduzana Steph., Sp. Hepat. (Stephani) 6: 227, 1921 (Stephani 1921). ** Plagiochila tortuosa Lindenb. et Gottsche, Mexik. Leverm.: 70, 1863 (Gottsche 1863). * Plagiochila tovarina Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 102, 1864 (Gottsche 1864). * Plagiochila trabeculatospinosa Herzog, Hedwigia 72 (6): 213, 1932 (Herzog 1932c). * Plagiochila trabutii Steph., Biblioth. Bot. 87 (2): 213, 1916 (Stephani 1916a). ** Plagiochila trianae Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 114, 1864 (Gottsche 1864). ** Plagiochila trichomanes Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: cc, 1889 [1890] (Spruce 1889). * Plagiochila tricuspis Steph., Sp. Hepat. (Stephani) 6: 229, 1921 (Stephani 1921). World checklist of hornworts and liverworts 251 ** Plagiochila tristaniana Váňa et J.J.Engel, Mem. New York Bot. Gard. 105: 89, 2013 (Váňa and Engel 2013). Nom. nov. pro Plagiochila fragilifolia S.W.Arnell, Results Norweg. Sci. Exped. Tristan da Cunha 42: 25, 1958 (Arnell 1958b), nom. illeg. ** Plagiochila tristis Steph., Bull. Herb. Boissier (sér. 2) 2 (10): 880 (249), 1902 (Stephani 1902g). ** Plagiochila trollii Herzog, Hedwigia 74 (2): 90, 1934 (Herzog 1934a). ** Plagiochila truncata Gottsche, Mexik. Leverm.: 25, 1863 (Gottsche 1863). ** Plagiochila uleana Steph., Bull. Herb. Boissier (sér. 2) 2 (10): 868 (237), 1902 (Stephani 1902g). ** Plagiochila umbrosioides L.Söderstr., Phytotaxa 208 (1): 85, 2015 (Söderström et al. 2015b). Nom. nov. pro Plagiochila umbrosa Steph., Sp. Hepat. (Stephani) 6: 235, 1921 (Stephani 1921), nom. illeg. * Plagiochila unduavensis Steph., Bull. Herb. Boissier (sér. 2) 2 (10): 882 (251), 1902 (Stephani 1902g). * Plagiochila usambarana Steph., Bull. Herb. Boissier (sér. 2) 2 (12): 980 (270), 1902 (Stephani 1902h). ** Plagiochila variedentata Steph., Bull. Herb. Boissier (sér. 2) 2 (10): 887 (256), 1902 (Stephani 1902g). ** Plagiochila vastifolia Steph., Bull. Herb. Boissier (sér. 2) 2 (8): 680 (217), 1902 (Stephani 1902a). ** Plagiochila velata Inoue et Piippo, Ann. Bot. Fenn. 26 (2): 207, 1989 (Piippo 1989b). * Plagiochila venezuelana Steph., Bull. Herb. Boissier (sér. 2) 5 (8): 749 (548), 1905 (Stephani 1905i). * Plagiochila ventricosotrigona Steph., Biblioth. Bot. 87 (2): 214, 1916 (Stephani 1916a). * Plagiochila verrucosa Steph., Bull. Herb. Boissier (sér. 2) 5 (9): 885 (551), 1905 (Stephani 1905g). * Plagiochila vetustisilva Steph., Sp. Hepat. (Stephani) 6: 245, 1922 (Stephani 1922). 182 * Plagiochila viminea Spruce, Mem. Torrey Bot. Club 1 (3): 134, 1890 (Spruce 1890). * Plagiochila viridis Steph., Sp. Hepat. (Stephani) 6: 239, 1921 (Stephani 1921). ** Plagiochila viridonigra (E.A.Hodgs.) Inoue, Bryologist 68 (2): 218, 1965 (Inoue 1965b). Bas.: Syzygiella viridonigra E.A.Hodgs., Rec. Domin. Mus. 4 (11): 120, 1962 (Hodgson 1962a). ** Plagiochila vittiana Inoue, Beih. Nova Hedwigia 90: 171, 1988 (Inoue 1988a). ** Plagiochila vittifolia Steph., Sp. Hepat. (Stephani) 6: 239, 1921 (Stephani 1921). ** Plagiochila vulcanica Steph., Bull. Herb. Boissier (sér. 2) 2 (8): 671 (208), 1902 (Stephani 1902a). ** Plagiochila wacei S.W.Arnell ex Váňa et J.J.Engel, Mem. New York Bot. Gard. 105: 89, 2013 (Váňa and Engel 2013). * Plagiochila wallisiana Steph., Bull. Herb. Boissier (sér. 2) 5 (10): 936 (586), 1905 (Stephani 1905f). 182 Plagiochila vetustisilva is a doubtful taxon. he type specimen was burned in B (Piippo 1989b). 252 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Plagiochila wattsiana J.J.Engel et G.L.Merr., Nova Hedwigia 91 (3/4): 511, 2010 (Engel and Smith Merrill 2010). Nom. nov. pro Plagiochila wattsii Steph., Sp. Hepat. (Stephani) 6: 240, 1921 (Stephani 1921), nom. illeg. ** Plagiochila wettsteiniana S.W.Arnell, Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 53, 1964 (Schifner and Arnell 1964). * Plagiochila weymouthiana Steph., Biblioth. Bot. 87 (2): 215, 1916 (Stephani 1916a). ** Plagiochila wiemanniana S.W.Arnell, Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 54, 1964 (Schifner and Arnell 1964). *** Plagiochila winteri Steph., Bull. Herb. Boissier (sér. 2) 2 (12): 981 (271), 1902 (Stephani 1902h). ** Plagiochila wrightii Steph., Bull. Herb. Boissier (sér. 2) 2 (8): 681 (218), 1902 (Stephani 1902a). ** Plagiochila xalapensis Gottsche, Mexik. Leverm.: 21, 1863 (Gottsche 1863). * Plagiochila xanthochroma Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 489, 1885 (Spruce 1885). * Plagiochila yoshinagana Steph., Sp. Hepat. (Stephani) 6: 242, 1922 (Stephani 1922). ** Plagiochila zacuapana Gottsche, Mexik. Leverm.: 20, 1863 (Gottsche 1863). ** Plagiochilidium Herzog, Mitt. Inst. Allg. Bot. Hamburg 7 (3): 186, 1931 (Herzog 1931a). ** Plagiochilidium bidentulum (Steph.) Grolle, J. Hattori Bot. Lab. 65: 408, 1988 (Grolle 1988c). Bas.: Tylimanthus bidentulus Steph., Bull. Herb. Boissier (sér. 2) 5 (12): 1134 (6), 1905 (Stephani 1905b). ** Plagiochilion S.Hatt., Biosphaera 1 (1): 7, 1947 (Hattori 1947a). *** Plagiochilion braunianum (Nees) S.Hatt., Biosphaera 1 (1): 7, 1947 (Hattori 1947a). Bas.: Jungermannia brauniana Nees, Enum. Pl. Crypt. Javae: 80, 1830 (Nees 1830). ** Plagiochilion combinatum (Mitt.) Inoue, J. Hattori Bot. Lab. 27: 55, 1964 (Inoue 1964b). Bas.: Plagiochila combinata Mitt., Fl. vit.: 408, 1871 [1873] (Mitten 1871). *** Plagiochilion conjugatum (Hook.) R.M.Schust., J. Hattori Bot. Lab. 26: 285, 1963 (Schuster 1963b). Bas.: Jungermannia conjugata Hook., Musci Exot. 1: tab. 91, 1818 (Hooker 1818). ** Plagiochilion imbriatum (Mitt.) Inoue, J. Hattori Bot. Lab. 27: 57, 1964 (Inoue 1964b). Bas.: Plagiochila imbriata Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 97, 1860 [1861] (Mitten 1860c). ** Plagiochilion giulianettii (Steph.) Inoue, J. Hattori Bot. Lab. 27: 57, 1964 (Inoue 1964b). Bas.: Plagiochila giulianettii Steph., Bull. Herb. Boissier (sér. 2) 4 (1): 30 (402), 1904 (Stephani 1904g). ** Plagiochilion herzogii Inoue, Bull. Natl. Sci. Mus. Tokyo (n.ser.) 14 (2): 270, 1971 (Inoue 1971c). ** Plagiochilion intermedium R.M.Schust., Phytologia 45 (5): 421, 1980 (Schuster 1980b). World checklist of hornworts and liverworts 253 ** Plagiochilion mayebarae S.Hatt., J. Hattori Bot. Lab. 3: 39, 1948 [1950] (Hattori 1948a). *** Plagiochilion oppositum (Reinw., Blume et Nees) S.Hatt., Biosphaera 1 (1): 7, 1947 (Hattori 1947a). Bas.: Jungermannia opposita Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 236, 1824 [1825] (Reinwardt et al. 1824a). ** Plagiochilion pachycephalum (De Not.) Inoue, J. Hattori Bot. Lab. 27: 55, 1964 (Inoue 1964b). Bas.: Plagiochila pachycephala De Not., Epat. Borneo: 14, 1874 (De Notaris 1874). *** Plagiochilion proliferum (Mitt.) R.M.Schust., J. Hattori Bot. Lab. 26: 285, 1963 (Schuster 1963b). Bas.: Plagiochila prolifera Mitt., Bot. antarct. voy. II (Fl. Nov.Zel. 2): 130, 1854 (Mitten 1854). ** Plagiochilion theriotanum (Steph.) Inoue, J. Hattori Bot. Lab. 27: 59, 1964 (Inoue 1964b). Bas.: Plagiochila theriotana Steph., Sp. Hepat. (Stephani) 6: 228, 1921 (Stephani 1921). ** Pseudolophocolea R.M.Schust. et J.J.Engel, Lindbergia 8 (2): 71, 1982 (Schuster and Engel 1982). Based on: Pseudolophocolea R.M.Schust. et J.J.Engel, Phytologia 47 (4): 310, 1981 (Schuster and Engel 1981). ** Pseudolophocolea denticulata R.M.Schust. et J.J.Engel, Lindbergia 8 (2): 73, 1982 (Schuster and Engel 1982). Based on: Pseudolophocolea denticulata R.M.Schust. et J.J.Engel, Phytologia 47 (4): 311, 1981 (Schuster and Engel 1981), nom. inval. ** Xenochila R.M.Schust., Amer. Midl. Naturalist 62 (1): 15, 1959 (Schuster 1959a). ** Xenochila integrifolia (Mitt.) Inoue, Bull. Natl. Sci. Mus. Tokyo (n.ser.) 6 (4): 373, 1963 (Inoue 1963). Bas.: Plagiochila integrifolia Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 96, 1860 [1861] (Mitten 1860c). *** Pseudolepicoleaceae Fulford et J.Taylor by M. von Konrat Blepharostoma was recognized as an element within Pseudolepicoleaceae by CrandallStotler et al. (2009), but we follow the concept of Frey and Stech (2008) with Blepharostoma as the single genus in Blepharostomataceae. *** Archeophylla R.M.Schust., J. Hattori Bot. Lab. 26: 263, 1963 (Schuster 1963b). ** Archeophylla paradoxa R.M.Schust., Trans. Brit. Bryol. Soc. 4 (5): 810, 1965 (Schuster 1965c). ** Archeophylla pungens (Herzog) R.M.Schust., Candollea 21 (1): 86, 1966 (Schuster 1966e). Bas.: Blepharostoma pungens Herzog, Rev. Bryol. Lichénol. 29 (3/4): 189, 1960 [1961] (Herzog 1960). 254 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Archeophylla schusteri (E.A.Hodgs. et Allison) R.M.Schust., J. Hattori Bot. Lab. 26: 263, 1963 (Schuster 1963b). Bas.: Temnoma schusteri E.A.Hodgs. et Allison, Trans. Roy. Soc. New Zealand, Bot. 1 (12): 147, 1962 (Hodgson and Allison 1962). *** Castanoclobos J.J.Engel et Glenny, Novon 17 (4): 424, 2007 (Engel and Glenny 2007). *** Castanoclobos julaceus (Hatcher ex J.J.Engel) J.J.Engel et Glenny, Novon 17 (4): 425, 2007 (Engel and Glenny 2007). Bas.: Leiomitra julacea Hatcher ex J.J.Engel, Novon 9 (1): 26, 1999 (Engel 1999a). ** Chaetocolea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 346, 1885 (Spruce 1885). *** Chaetocolea palmata Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 346, 1885 (Spruce 1885). *** Herzogiaria Fulford ex Hässel, Lindbergia 7 (1): 23, 1981 (Hässel 1981). *** Herzogiaria teres (Steph.) Fulford ex Hässel, Lindbergia 7 (1): 24, 1981 (Hässel 1981). Bas.: Lepicolea teres Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 17): 26, 1901 (Stephani 1901b). ** Isophyllaria E.A.Hodgs. et Allison, Trans. Roy. Soc. New Zealand, Bot. 3 (4): 68, 1965 (Hodgson 1965). *** Isophyllaria attenuata (Rodway) E.A.Hodgs., J. Roy. Soc. New Zealand 2 (1): 111, 1972 (Hodgson 1972). Bas.: Isotachis attenuata Rodway, Pap. & Proc. Roy. Soc. Tasmania 1916: 47, 1917 (Rodway 1917a). ** Isophyllaria fuegiana (Hässel) R.M.Schust., Beih. Nova Hedwigia 118: 141, 2000 (Schuster 2000a). Bas.: Fulfordiella fuegiana Hässel, Comun. Mus. Argent. Ci. Nat. “Bernardino Rivadavia,” Ci. Bot. 2 (9): 48, 1974 (Hässel 1974). *** Pseudolepicolea Fulford et J.Taylor, Nova Hedwigia 1 (3/4): 412, 1959 [1960] (Fulford and Taylor 1959b). ** Pseudolepicolea andoi (R.M.Schust.) Inoue, Bull. Natl. Sci. Mus. Tokyo, B 4 (3): 94, 1978 (Inoue 1978a). Bas.: Lophochaete andoi R.M.Schust., J. Hattori Bot. Lab. 26: 261, 1963 (Schuster 1963b). ** Pseudolepicolea fryei (Perss.) Grolle et Ando, Hikobia 3 (3): 180, 1963 (Ando 1963). Bas.: Lepicolea fryei Perss., Bryologist 49 (2): 47, 1946 (Persson 1946). *** Pseudolepicolea grolleana (R.M.Schust.) Grolle, Ann. Bot. Fenn. 21 (1): 30, 1984 (Piippo 1984a). Bas.: Archeochaete grolleana R.M.Schust., Nova Hedwigia 15: 441, 1968 (Schuster 1968b). *** Pseudolepicolea kuehnemannii (R.M.Schust.) Hässel, Fl. Criptog. Tierra del Fuego 15: 125, 1975 (Hässel and Solari 1975). Bas.: Archeochaete kuehnemannii R.M.Schust., J. Hattori Bot. Lab. 26: 262, 1963 (Schuster 1963b). World checklist of hornworts and liverworts 255 *** Pseudolepicolea quadrilaciniata (Sull.) Fulford et J.Taylor, Nova Hedwigia 1 (3/4): 413, 1959 [1960] (Fulford and Taylor 1959b). Bas.: Sendtnera quadrilaciniata Sull., Hooker’s J. Bot. Kew Gard. Misc. 2: 317, 1850 (Sullivant 1850). *** Pseudolepicolea temnomoides (R.M.Schust.) Váňa et J.J.Engel, Mem. New York Bot. Gard. 105: 92, 2013 (Váňa and Engel 2013). Bas.: Archeochaete temnomoides R.M.Schust., Candollea 21 (1): 129, 1966 (Schuster 1966e). ** Pseudolepicolea trollii (Herzog) Grolle et Ando, Hikobia 3 (3): 177, 1963 (Ando 1963). Bas.: Blepharostoma trollii Herzog, Ann. Bryol. 12: 80, 1939 (Herzog 1939b). * Pseudolepicolea trollii var. darjeelingensis S.Hatt. et Mizut., J. Hattori Bot. Lab. 31: 252, 1968 (Hattori and Mizutani 1968). *** Temnoma Mitt., Handb. N. Zeal. l. 2: 750, 1867 (Hooker 1867). *** Temnoma angustifolium R.M.Schust., Candollea 21 (2): 279, 1966 [1967] (Schuster 1966c). *** Temnoma chaetophyllum R.M.Schust., Phytologia 39 (4): 239, 1978 (Schuster 1978a). ** Temnoma palmatum (Lindb. ex Pearson) R.M.Schust., Bryologist 62 (4): 240, 1959 [1960] (Schuster 1959c). Bas.: Blepharostoma palmatum Lindb. ex Pearson, J. Bot. 25: 193, 1887 (Pearson 1887a). ** Temnoma palmatum var. cuneatum R.M.Schust., Candollea 21 (2): 347, 1966 [1967] (Schuster 1966c). ** Temnoma palmatum var. laxifolium R.M.Schust., Candollea 21 (2): 345, 1966 [1967] (Schuster 1966c). ** Temnoma palmatum var. pseudospiniferum R.M.Schust., Candollea 21 (2): 348, 1966 [1967] (Schuster 1966c). ** Temnoma patagonicum R.M.Schust., Candollea 21 (2): 313, 1966 [1967] (Schuster 1966c). *** Temnoma paucisetigerum R.M.Schust., Candollea 21 (2): 266, 1966 [1967] (Schuster 1966c). *** Temnoma pilosum (A.Evans) R.M.Schust., Bryologist 62 (4): 240, 1959 [1960] (Schuster 1959c). Bas.: Blepharostoma pilosum A.Evans, Bull. Torrey Bot. Club 25 (8): 413, 1898 (Evans 1898). *** Temnoma pulchellum (Hook.) Mitt., Handb. N. Zeal. l. 2: 753, 1867 (Hooker 1867). Bas.: Jungermannia pulchella Hook., Musci Exot. 1: tab. 94, 1818 (Hooker 1818). *** Temnoma quadriidum (Mitt.) Mitt. ex E.A.Hodgs. et Allison, Trans. Roy. Soc. New Zealand, Bot. 1 (12): 142, 1962 (Hodgson and Allison 1962). Bas.: Jungermannia quadriida Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 128, 1854 (Mitten 1854). *** Temnoma quadripartitum (Hook.) Mitt., J. Linn. Soc., Bot. 15 (82): 68, 1876 (Mitten 1876a). Bas.: Jungermannia quadripartita Hook., Musci Exot. 2: tab. 117, 1820 (Hooker 1820). ** Temnoma quadripartitum var. pseudopungens R.M.Schust., Candollea 21 (2): 312, 1966 [1967] (Schuster 1966c). 256 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Temnoma quadripartitum var. randii (S.W.Arnell) R.M.Schust., Candollea 21 (2): 307, 1966 [1967] (Schuster 1966c). Bas.: Lepidozia randii S.W.Arnell, Svensk Bot. Tidskr. 47 (3): 417, 1953 (Arnell 1953c). *** Temnoma setigerum (Lindenb.) R.M.Schust., Nova Hedwigia 5: 35, 1963 (Schuster 1963c). Bas.: Jungermannia setigera Lindenb., Syn. Hepat. 1: 131, 1844 (Gottsche et al. 1844). ** Temnoma setigerum var. hawaiicum Inoue, Bull. Natl. Sci. Mus. Tokyo (n.ser.) 17 (3): 228, 1974 (Inoue 1974b). ** Temnoma townrowii R.M.Schust., Candollea 21 (2): 351, 1966 [1967] (Schuster 1966c). *** Trichocoleaceae Nakai by T. Katagiri he treatment of Trichocoleaceae follows what was outlined in Katagiri and Deguchi (2012) and Katagiri et al. (2012, 2013). Recent nomenclatural and taxonomic notes can also be found in Katagiri (2013). *** Eotrichocolea R.M.Schust., J. Hattori Bot. Lab. 26: 252, 1963 (Schuster 1963b). *** Eotrichocolea furukii T.Katag., Bryologist 115 (4): 519, 2012 (Katagiri et al. 2012). *** Eotrichocolea polyacantha (Hook.f. et Taylor) R.M.Schust., J. Hattori Bot. Lab. 26: 264, 1963 (Schuster 1963b). Bas.: Jungermannia polyacantha Hook.f. et Taylor, London J. Bot. 3: 290 [390], 1844 (Hooker and Taylor 1844a). *** Leiomitra Lindb., Acta Soc. Sci. Fenn. 10: 515, 1875 (Lindberg 1875). ** subg. Brachygyna R.M.Schust., Nova Hedwigia 73 (3/4): 480, 2001 (Schuster 2001b). ** Leiomitra mastigophoroides R.M.Schust., Phytologia 45 (5): 416, 1980 (Schuster 1980b). ** subg. Leiomitra, Nova Hedwigia 73 (3/4): 480, 2001 (Schuster 2001b). *** Leiomitra breviseta (Steph.) R.M.Schust., Beih. Nova Hedwigia 118: 152, 2000 (Schuster 2000a). Bas.: Trichocolea breviseta Steph., Sp. Hepat. (Stephani) 4: 60, 1909 (Stephani 1909d). *** Leiomitra capillata Lindb., Acta Soc. Sci. Fenn. 10: 515, 1875 (Lindberg 1875). *** Leiomitra elegans (Lehm.) Hässel, Novon 12 (4): 465, 2002 (Hässel 2002). Bas.: Trichocolea elegans Lehm., Nov. Stirp. Pug. 10: 8, 1857 (Lehmann 1857). ** Leiomitra elliottii (Steph.) R.M.Schust., Nova Hedwigia 73 (3/4): 469, 2001 (Schuster 2001b). Bas.: Trichocolea elliottii Steph., Sp. Hepat. (Stephani) 4: 55, 1909 (Stephani 1909d). World checklist of hornworts and liverworts 257 *** Leiomitra laccida Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 349, 1885 (Spruce 1885). ** Leiomitra hirticaulis R.M.Schust., Nova Hedwigia 73 (3/4): 472, 2001 (Schuster 2001b). *** Leiomitra lanata (Hook.) R.M.Schust., Phytologia 45 (5): 417, 1980 (Schuster 1980b). Bas.: Jungermannia lanata Hook., Musci Exot. 2: tab. 116, 1820 (Hooker 1820). *** Leiomitra merrillana (Steph.) T.Katag., Bryologist 115 (4): 488, 2012 (Katagiri and Deguchi 2012). Bas.: Trichocolea merrillana Steph., Sp. Hepat. (Stephani) 6: 374, 1923 (Stephani 1923). *** Leiomitra paraphyllina Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 350, 1885 (Spruce 1885). ** Leiomitra robusta (Steph.) R.M.Schust., Nova Hedwigia 73 (3/4): 469, 2001 (Schuster 2001b). Bas.: Trichocolea robusta Steph., Sp. Hepat. (Stephani) 4: 58, 1909 (Stephani 1909d). ** Leiomitra smaragdina Hässel, Novon 12 (4): 467, 2002 (Hässel 2002). *** Leiomitra tomentosa (Sw.) Lindb., Acta Soc. Sci. Fenn. 10: 515, 1875 (Lindberg 1875). Bas.: Jungermannia tomentosa Sw., Prodr. (Swartz): 145, 1788 (Swartz 1788). *** Trichocolea Dumort., Commentat. Bot. (Dumortier): 113, 1822 (Dumortier 1822) nom. conserv. ** Trichocolea argentea Herzog, Arch. Bot. São Paulo 1 (2): 40, 1925 (Herzog 1925b). *** Trichocolea breviissa Steph., Sp. Hepat. (Stephani) 4: 54, 1909 (Stephani 1909d). ** Trichocolea comptonii Pearson, J. Linn. Soc., Bot. 46 (305): 27, 1922 (Pearson 1922b). *** Trichocolea ilicaulis Steph., Sp. Hepat. (Stephani) 4: 59, 1909 (Stephani 1909d). * Trichocolea loccosa Herzog et Hatcher, Lloydia 20 (3): 148, 1957 [1958] (Hatcher 1957). 183 ** Trichocolea geniculata Pearson, J. Linn. Soc., Bot. 46 (305): 28, 1922 (Pearson 1922b). ** Trichocolea gracillima Austin, Bot. Gaz. 3 (1): 6, 1878 (Austin 1878). *** Trichocolea hatcheri E.A.Hodgs., Trans. Roy. Soc. New Zealand, Bot. 3 (4): 69, 1965 (Hodgson 1965). *** Trichocolea iriomotensis T.Katag., Hattoria 4: 6, 2013 (Katagiri et al. 2013). *** Trichocolea japonica T.Katag., Bryologist 114 (4): 744, 2011 (Katagiri et al. 2011). *** Trichocolea magna T.Katag., Hattoria 4: 7, 2013 (Katagiri et al. 2013). ** Trichocolea minutifolia Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 134, 1914 (Stephani and Watts 1914). *** Trichocolea mollissima (Hook.f. et Taylor) Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 132, 1864 (Gottsche 1864). Bas.: Jungermannia mollissima Hook.f. et Taylor, London J. Bot. 3: 290 [390], 1844 (Hooker and Taylor 1844a). 183 Trichocolea loccosa is treated as a variety of Trichocolea laccida (=Leiomitra laccida by Schultze-Motel & Menzel (1987). 258 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Trichocolea pluma (Reinw., Blume et Nees) Mont., Voy. Bonite, Bot. 1: 238, 1846 (Montagne 1846). Bas.: Jungermannia pluma Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 209, 1824 [1825] (Reinwardt et al. 1824a). *** Trichocolea rigida R.M.Schust., Nova Hedwigia 15: 447, 1968 (Schuster 1968b). *** Trichocolea rudimentaris Steph., Sp. Hepat. (Stephani) 6: 376, 1923 (Stephani 1923). * Trichocolea sprucei Steph., Sp. Hepat. (Stephani) 4: 59, 1909 (Stephani 1909d). Nom. nov. pro Trichocolea gracillima Spruce, J. Linn. Soc., Bot. 30 (210): 353, 1895 (Gepp 1895b), nom. illeg. 184 *** Trichocolea tomentella (Ehrh.) Dumort., Syll. Jungerm. Europ.: 67, 1831 (Dumortier 1831). Bas.: Jungermannia tomentella Ehrh., Hannover. Mag. 21 (18): 277, 1783 (Ehrhart 1783). ** Trichocolea udarii D.K.Singh, Bull. Bot. Surv. India 25: 177, 1983 [1985] (Singh 1983a). ** Trichocolea wattsiana Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 135, 1914 (Stephani and Watts 1914). Myliineae J.J.Engel et Braggins ex Crand.-Stotl.,Váňa, Stotler et J.J.Engel *** Myliaceae Schljakov by L. Söderström Söderström et al. (2015c) outlines the controversy as to the placement of Mylia anomala where they advocate recognition at the subgeneric level. It has previously been recognized as a segregate genus, Leiomylia J.J.Engel and Braggins (cf. also Shaw et al. 2015). *** Mylia Gray, Nat. Arr. Brit. Pl. 1: 693, 1821 (Gray 1821) nom. conserv. ** subg. Anomalae (R.M.Schust. ex Potemkin) L.Söderstr., Phytotaxa 202 (1): 70, 2015 (Söderström et al. 2015c). Bas.: Mylia sect. Anomalae R.M.Schust. ex Potemkin, Arctoa 2: 1, 1993 (Potemkin and Kazanovsky 1993). *** Mylia anomala (Hook.) Gray, Nat. Arr. Brit. Pl. 1: 693, 1821 (Gray 1821). Bas.: Jungermannia anomala Hook., Brit. Jungermann.: tab. 34, 1813 (Hooker 1813). ** subg. Mylia *** Mylia taylorii (Hook.) Gray, Nat. Arr. Brit. Pl. 1: 693, 1821 (Gray 1821). Bas.: Jungermannia taylorii Hook., Brit. Jungermann.: tab. 57, 1813 (Hooker 1813). *** Mylia verrucosa Lindb., Acta Soc. Sci. Fenn. 10: 236, 1872 [1873] (Lindberg 1872b). 184 Trichocolea sprucei is suggested to be a variety of Trichocolea laccida (=Leiomitra laccida) in SchultzeMotel & Menzel (1987) World checklist of hornworts and liverworts 259 *** Mylia verrucosa subsp. nuda (Inoue et B.Y.Yang) Potemkin et Kazan., Arctoa 2: 5, 1993 (Potemkin and Kazanovsky 1993). Bas.: Mylia nuda Inoue et B.Y.Yang, Taiwania 12 (1): 35, 1966 (Inoue and Yang 1966). Perssoniellineae R.M.Schust. *** Schistochilaceae H.Buch by X. He and D. Glenny He et al. (2014b) provided an historical account of Schistochilaceae summarizing studies that have showed that the phylogenetic structure of the family does not match units that have resulted from morphologically-based investigations. Further studies are needed until a natural division of the family can be proposed. hus, we follow the broad concept of He and Glenny (2010) and He and Sun (2013) here, treating Schistochilaceae as comprising a single genus as discussed by He et al. (2014b). *** Schistochila Dumort., Recueil Observ. Jungerm.: 15, 1835 (Dumortier 1835). ** Schistochila acuminata Steph., Sp. Hepat. (Stephani) 4: 81, 1909 (Stephani 1909d). *** Schistochila aequiloba Steph., Sp. Hepat. (Stephani) 4: 80, 1909 (Stephani 1909d). *** Schistochila alata (Lehm.) Schifn., Hepat. (Engl.-Prantl): 111, 1893 (Schifner 1893b). Bas.: Jungermannia alata Lehm., Linnaea 4: 359, 1829 (Lehmann 1829). *** Schistochila aligera (Nees et Blume) J.B.Jack et Steph., Hedwigia 31 (1): 12, 1892 (Jack and Stephani 1892). Bas.: Jungermannia aligera Nees et Blume, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 11: 135, 1823 (Blume and Nees 1823). * Schistochila aligera var. laxa (Nees) Schifn., Consp. Hepat. Arch. Ind.: 212, 1898 (Schifner 1898b). Bas.: Jungermannia aligera γ laxa Nees, Enum. Pl. Crypt. Javae: 68, 1830 (Nees 1830). *** Schistochila altissima E.A.Hodgs., Trans. Roy. Soc. New Zealand, Bot. 3 (4): 85, 1965 (Hodgson 1965). ** Schistochila altissima subsp. polystratosa R.M.Schust. et J.J.Engel, Phytologia 30 (4): 241, 1975 (Schuster and Engel 1975). *** Schistochila antara Grolle, J. Hattori Bot. Lab. 29: 249, 1966 (Grolle 1966h). *** Schistochila appendiculata (Hook.) Dumort. ex Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 392, 1877 (Trevisan 1877). Bas.: Jungermannia appendiculata Hook., Musci Exot. 1: tab. 15, 1818 (Hooker 1818). * Schistochila baileyana Steph., Sp. Hepat. (Stephani) 4: 85, 1909 (Stephani 1909d). 185 *** Schistochila balfouriana (Hook.f. et Taylor) Steph., Sp. Hepat. (Stephani) 4: 91, 1909 (Stephani 1909d). Bas.: Jungermannia balfouriana Hook.f. et Taylor, London J. Bot. 3: 556, 1844 (Hooker and Taylor 1844d). 185 Schistochila baileyana is possibly conspeciic with Schistochila beccariana (D. Meagher, pers. comm.). 260 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Schistochila beccariana (De Not.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 392, 1877 (Trevisan 1877). Bas.: Gottschea beccariana De Not., Epat. Borneo: 9, 1874 (De Notaris 1874). *** Schistochila berggrenii (J.J.Engel et R.M.Schust.) Xiao L.He et Glenny, Austral. Syst. Bot. 23 (4): 237, 2010 (He and Glenny 2010). Bas.: Pachyschistochila berggrenii J.J.Engel et R.M.Schust., J. Hattori Bot. Lab. 58: 476, 1985 (Schuster and Engel 1985). *** Schistochila berteroana (Hook.) Steph., Sp. Hepat. (Stephani) 4: 96, 1909 (Stephani 1909d). Bas.: Jungermannia berteroana Hook., Bot. Misc. 2: 148, 1831 (Hooker 1831). *** Schistochila blumei (Nees) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 392, 1877 (Trevisan 1877). Bas.: Jungermannia blumei Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 11: 136, 1823 (Blume and Nees 1823). ** Schistochila brassii Grolle, J. Hattori Bot. Lab. 31: 5, 1968 (Grolle 1968a). ** Schistochila caledonica Steph., Sp. Hepat. (Stephani) 4: 77, 1909 (Stephani 1909d). *** Schistochila carnosa (Mitt.) Steph., Sp. Hepat. (Stephani) 4: 93, 1909 (Stephani 1909d). Bas.: Gottschea carnosa Mitt., J. Linn. Soc., Bot. 15 (82): 72, 1876 (Mitten 1876a). *** Schistochila caudata R.M.Schust. et J.J.Engel, Phytologia 30 (4): 242, 1975 (Schuster and Engel 1975). *** Schistochila childii (R.M.Schust. et J.J.Engel) Xiao L.He et Glenny, Austral. Syst. Bot. 23 (4): 237, 2010 (He and Glenny 2010). Bas.: Pachyschistochila childii R.M.Schust. et J.J.Engel, J. Hattori Bot. Lab. 58: 471, 1985 (Schuster and Engel 1985). *** Schistochila chlorophylla (Colenso) E.A.Hodgs., Trans. & Proc. Roy. Soc. New Zealand 71 (3): 192, 1941 (Hodgson 1941). Bas.: Gottschea chlorophylla Colenso, Trans. & Proc. New Zealand Inst. 18: 240, 1886 (Colenso 1886b). *** Schistochila ciliata (Mitt.) Steph., Sp. Hepat. (Stephani) 4: 87, 1909 (Stephani 1909d). Bas.: Gottschea ciliata Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 151, 1854 (Mitten 1854). *** Schistochila compacta (Colenso) E.A.Hodgs., Trans. & Proc. Roy. Soc. New Zealand 71 (3): 192, 1941 (Hodgson 1941). Bas.: Gottschea compacta Colenso, Trans. & Proc. New Zealand Inst. 16: 349, 1884 (Colenso 1884). *** Schistochila conchophylla Herzog ex E.A.Hodgs. et Allison, Trans. & Proc. Roy. Soc. New Zealand 71 (3): 191, 1941 (Hodgson 1941). ** Schistochila conchophylla var. multidentata (J.J.Engel) Xiao L.He et Glenny, Phytotaxa 173 (1): 92, 2014 (He et al. 2014b). Bas.: Gottschea conchophylla var. multidentata J.J.Engel, Nova Hedwigia 93 (3/4): 407, 2011 (Engel 2011). *** Schistochila congoana Steph., Sp. Hepat. (Stephani) 4: 71, 1909 (Stephani 1909d). ** Schistochila cookei (H.A.Mill.) R.M.Schust., J. Hattori Bot. Lab. 42: 274, 1977 (Schuster and Engel 1977). Bas.: Fulfordistria cookei H.A.Mill., Phytologia 20 (5): 320, 1970 (Miller 1970). ** Schistochila crinita Grolle, J. Hattori Bot. Lab. 31: 7, 1968 (Grolle 1968a). World checklist of hornworts and liverworts 261 *** Schistochila cristata Steph., Hedwigia 28 (4): 274, 1889 (Stephani 1889c). *** Schistochila cunninghamii Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 17): 27, 1901 (Stephani 1901b). ** Schistochila doriae (De Not.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 392, 1877 (Trevisan 1877). Bas.: Gottschea doriae De Not., Epat. Borneo: 10, 1874 (De Notaris 1874). ** Schistochila engleriana Steph., Sp. Hepat. (Stephani) 4: 69, 1909 (Stephani 1909d). *** Schistochila exalata Herzog, Rev. Bryol. Lichénol. 29 (3/4): 191, 1960 [1961] (Herzog 1960). ** Schistochila ijiensis H.Buch et Herzog, Memoranda Soc. Fauna Fl. Fennica 27: 94, 1952 (Herzog 1952c). *** Schistochila glaucescens (Hook.) A.Evans, Rev. Bryol. 32 (4): 60, 1905 (Evans 1905c). Bas.: Jungermannia glaucescens Hook., Musci Exot. 1: tab. 39, 1818 (Hooker 1818). ** Schistochila hattorii Grolle, J. Hattori Bot. Lab. 29: 247, 1966 (Grolle 1966h). ** Schistochila integerrima Steph., Sp. Hepat. (Stephani) 6: 492, 1924 (Stephani 1924). ** Schistochila isotachyphylla (J.J.Engel et R.M.Schust.) Xiao L.He et Glenny, Austral. Syst. Bot. 23 (4): 237, 2010 (He and Glenny 2010). Bas.: Paraschistochila isotachyphylla J.J.Engel et R.M.Schust., J. Hattori Bot. Lab. 58: 429, 1985 (Schuster and Engel 1985). *** Schistochila kirkiana Steph., Sp. Hepat. (Stephani) 4: 86, 1909 (Stephani 1909d). *** Schistochila kunkelii S.W.Arnell, Ark. Bot. (n.ser.) 4 (1): 12, 1957 (Arnell 1957b). ** Schistochila lacerata Steph., Sp. Hepat. (Stephani) 6: 492, 1924 (Stephani 1924). *** Schistochila lamellata (Hook.) Dumort. ex A.Evans, Contr. U.S. Natl. Herb. 1 (5): 141, 1892 (Evans 1892b). Bas.: Jungermannia lamellata Hook., Musci Exot. 1: tab. 49, 1818 (Hooker 1818). *** Schistochila laminigera (Hook.f. et Taylor) A.Evans, Contr. U.S. Natl. Herb. 1 (5): 141, 1892 (Evans 1892b). Bas.: Jungermannia laminigera Hook.f. et Taylor, London J. Bot. 3: 456, 1844 (Hooker and Taylor 1844b). *** Schistochila latiloba (R.M.Schust. et J.J.Engel) Xiao L.He et Glenny, Austral. Syst. Bot. 23 (4): 237, 2010 (He and Glenny 2010). Bas.: Pachyschistochila latiloba R.M.Schust. et J.J.Engel, J. Hattori Bot. Lab. 58: 493, 1985 (Schuster and Engel 1985). *** Schistochila lehmanniana (Lindenb.) Steph., Sp. Hepat. (Stephani) 4: 86, 1909 (Stephani 1909d). Bas.: Jungermannia lehmanniana Lindenb., Nov. Stirp. Pug. 4: 60, 1832 (Lehmann 1832). *** Schistochila leucophylla (Lehm. ex Gottsche, Lindenb. et Nees) Steph., Sp. Hepat. (Stephani) 4: 98, 1910 (Stephani 1910b). Bas.: Gottschea leucophylla Lehm. ex Gottsche, Lindenb. et Nees, Syn. Hepat. 1: 17, 1844 (Gottsche et al. 1844). ** Schistochila macrodonta W.E.Nicholson, Symb. Sin. 5: 29, 1930 (Nicholson et al. 1930). ** Schistochila minor C.Gao et Y.H.Wu, J. Hattori Bot. Lab. 95: 267, 2004 (Gao and Wu 2004). 262 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Schistochila monticola R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 14 (4): 623, 1971 (Schuster 1971c). *** Schistochila muricata E.A.Hodgs. et Allison, Trans. & Proc. Roy. Soc. New Zealand 71 (3): 186, 1941 (Hodgson 1941). *** Schistochila nadeaudiana Steph., Sp. Hepat. (Stephani) 4: 76, 1909 (Stephani 1909d). ** Schistochila neesii (Mont.) Lindb., J. Linn. Soc., Bot. 13 (67): 194, 1872 [1873] (Lindberg 1872a). Bas.: Gottschea neesii Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 244, 1843 (Montagne 1843). *** Schistochila nitidissima R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 11 (1): 27, 1968 (Schuster 1968a). *** Schistochila nivicola (R.M.Schust. et J.J.Engel) Xiao L.He et Glenny, Austral. Syst. Bot. 23 (4): 237, 2010 (He and Glenny 2010). Bas.: Pachyschistochila nivicola R.M.Schust. et J.J.Engel, J. Hattori Bot. Lab. 58: 512, 1985 (Schuster and Engel 1985). *** Schistochila nobilis (Hook.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 392, 1877 (Trevisan 1877). Bas.: Jungermannia nobilis Hook., Musci Exot. 1: tab. 11, 1818 (Hooker 1818). *** Schistochila nuda Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 215, 1934 (Horikawa 1934). *** Schistochila pachyphylla (Lehm.) Steph., Sp. Hepat. (Stephani) 4: 99, 1910 (Stephani 1910b). Bas.: Jungermannia pachyphylla Lehm., Nov. Stirp. Pug. 6: 61, 1834 (Lehmann 1834). *** Schistochila papillifera R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 11 (1): 27, 1968 (Schuster 1968a). *** Schistochila parvistipula Rodway, Pap. & Proc. Roy. Soc. Tasmania 1916: 47, 1917 (Rodway 1917a). *** Schistochila pellucida R.M.Schust. et J.J.Engel, J. Hattori Bot. Lab. 58: 286, 1985 (Schuster and Engel 1985). *** Schistochila piligera Steph., Bot. Gaz. 15 (11): 291, 1890 (Stephani 1890c). *** Schistochila pinnatifolia (Hook.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 392, 1877 (Trevisan 1877). Bas.: Jungermannia pinnatifolia Hook., Musci Exot. 2: tab. 114, 1820 (Hooker 1820). *** Schistochila pluriciliata R.M.Schust. et J.J.Engel, J. Hattori Bot. Lab. 58: 395, 1985 (Schuster and Engel 1985). *** Schistochila pseudociliata R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 14 (4): 623, 1971 (Schuster 1971c). *** Schistochila quadriida A.Evans, Contr. U.S. Natl. Herb. 1 (5): 141, 1892 (Evans 1892b). ** Schistochila ramentacea Steph., Sp. Hepat. (Stephani) 6: 494, 1924 (Stephani 1924). *** Schistochila relexa (Mont.) Steph., Sp. Hepat. (Stephani) 4: 97, 1910 (Stephani 1910b). Bas.: Gottschea relexa Mont., Ann. Sci. Nat. Bot. (sér. 3) 4: 347, 1845 (Montagne 1845b). World checklist of hornworts and liverworts 263 *** Schistochila relexistipula J.J.Engel et R.M.Schust., Phytologia 30 (4): 245, 1975 (Schuster and Engel 1975). *** Schistochila reinwardtii (Nees) Schifn., Consp. Hepat. Arch. Ind.: 218, 1898 (Schifner 1898b). Bas.: Jungermannia reinwardtii Nees, Enum. Pl. Crypt. Javae: 66, 1830 (Nees 1830). *** Schistochila repleta (Hook.f. et Taylor) Steph., Sp. Hepat. (Stephani) 4: 90, 1909 (Stephani 1909d). Bas.: Jungermannia repleta Hook.f. et Taylor, London J. Bot. 3: 557, 1844 (Hooker and Taylor 1844d). ** Schistochila rubriseta Steph., Sp. Hepat. (Stephani) 4: 78, 1909 (Stephani 1909d). ** Schistochila schultzei Steph., Sp. Hepat. (Stephani) 6: 494, 1924 (Stephani 1924). *** Schistochila sciophila R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 14 (4): 632, 1971 (Schuster 1971c). *** Schistochila sciurea (Nees) Schifn., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 60 (2): 251, 1893 (Schifner 1893a). Bas.: Jungermannia sciurea Nees, Enum. Pl. Crypt. Javae: 34, 1830 (Nees 1830). *** Schistochila simulans (C.Massal.) Xiao L.He et Yu Sun, Polish Bot. J. 58 (2): 473, 2013 (He and Sun 2013). Bas.: Cephalozia simulans C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 236, 1885 (Massalongo 1885). *** Schistochila spegazziniana (C.Massal.) Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 6): 60, 1900 (Stephani 1900b). Bas.: Gottschea spegazziniana C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 206, 1885 (Massalongo 1885). *** Schistochila sphagnoides (Schwägr.) Lindb. ex Steph., Sp. Hepat. (Stephani) 4: 70, 1909 (Stephani 1909d). Bas.: Jungermannia sphagnoides Schwägr., Hist. Musc. Hepat. Prodr.: 23, 1814 (Schwägrichen 1814). *** Schistochila splachnophylla (Hook.f. et Taylor) Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 17): 28, 1901 (Stephani 1901b). Bas.: Jungermannia splachnophylla Hook.f. et Taylor, London J. Bot. 3: 455, 1844 (Hooker and Taylor 1844b). ** Schistochila stratosa (Mont.) A.Evans, Contr. U.S. Natl. Herb. 1 (5): 141, 1892 (Evans 1892b). Bas.: Gottschea stratosa Mont., Ann. Sci. Nat. Bot. (sér. 3) 4: 346, 1845 (Montagne 1845b). *** Schistochila subhyalina R.M.Schust., Phytologia 30 (4): 246, 1975 (Schuster and Engel 1975). ** Schistochila subhyalina var. grandidentata (J.J.Engel et R.M.Schust.) Xiao L.He et Glenny, Austral. Syst. Bot. 23 (4): 237, 2010 (He and Glenny 2010). Bas.: Pachyschistochila subhyalina var. grandidentata J.J.Engel et R.M.Schust., J. Hattori Bot. Lab. 58: 507, 1985 (Schuster and Engel 1985). *** Schistochila subimmersa J.J.Engel et R.M.Schust., Phytologia 30 (4): 247, 1975 (Schuster and Engel 1975). *** Schistochila succulenta (J.J.Engel et R.M.Schust.) Xiao L.He et Glenny, Austral. Syst. Bot. 23 (4): 237, 2010 (He and Glenny 2010). Bas.: Pachyschistochila succulenta J.J.Engel et R.M.Schust., J. Hattori Bot. Lab. 58: 517, 1985 (Schuster and Engel 1985). 264 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Schistochila tasmanica Steph., Sp. Hepat. (Stephani) 4: 86, 1909 (Stephani 1909d). *** Schistochila trispiralis R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 11 (1): 28, 1968 (Schuster 1968a). *** Schistochila tuloides (Hook.f. et Taylor) Steph., Sp. Hepat. (Stephani) 4: 89, 1909 (Stephani 1909d). Bas.: Jungermannia tuloides Hook.f. et Taylor, London J. Bot. 3: 558, 1844 (Hooker and Taylor 1844d). *** Schistochila undulatifolia Piippo, Ann. Bot. Fenn. 23 (1): 8, 1986 (Piippo 1986b). *** Schistochila virescens R.M.Schust., Phytologia 30 (4): 248, 1975 (Schuster and Engel 1975). *** Schistochila vitreocincta (Herzog) Xiao L.He et Glenny, Austral. Syst. Bot. 23 (4): 237, 2010 (He and Glenny 2010). Bas.: Perssoniella vitreocincta Herzog, Ark. Bot. (n.ser.) 2 (4): 265, 1952 (Herzog 1952g). ** Schistochila volans Grolle, J. Hattori Bot. Lab. 29: 244, 1966 (Grolle 1966h). *** Schistochila yakushimensis N.Ohnishi et Deguchi, Bryologist 106 (3): 451, 2003 (Ohnishi and Deguchi 2003). ** Schistochila zantenii Grolle, J. Hattori Bot. Lab. 29: 243, 1966 (Grolle 1966h). Porellales Schljakov Jubulineae Müll.Frib. *** Frullaniaceae Lorch by M. von Konrat, J. Hentschel, J. Uribe, P. Sukkharak, J. Heinrichs, J. Larraín, R. Stotler and L. Zhang he subgeneric treatment of Frullaniaceae follows Hentschel et al. (2015), which includes a slight departure from formal infrageneric ranks where they apply clades referred to as Diastaloba I, II, III, and IV that appear to represent distinct subgenera. *** Frullania Raddi, Jungermanniogr. Etrusca: 9, 1818 (Raddi 1818a). *** subg. Chonanthelia Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 8, 1884 (Spruce 1884). * Frullania lammea Taylor, Trans. & Proc. Bot. Soc. Edinburgh 15: 29, 1884 (Spruce 1884). *** Frullania lindmanii Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 23 (III, 2): 19, 1897 (Stephani 1897a). ** Frullania paranensis Steph., Sp. Hepat. (Stephani) 4: 607, 1911 (Stephani 1911e). ** Frullania spegazzinii M.E.Reiner, Bol. Soc. Argent. Bot. 25 (3/4): 310, 1988 (Reiner 1988). *** sect. Chonanthelia (Spruce) Yuzawa ex Hentschel et von Konrat, Phytotaxa 220 (2): 129, 2015 (Hentschel et al. 2015). Bas.: Frullania subg. Chonanthelia Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 8, 1884 (Spruce 1884). World checklist of hornworts and liverworts 265 *** Frullania gibbosa Nees, Syn. Hepat. 3: 411, 1845 (Gottsche et al. 1845b). *** sect. Cladocarpicae Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 43, 1884 (Spruce 1884). *** Frullania beauverdii Steph., Biblioth. Bot. 87 (2): 241, 1916 (Stephani 1916a). *** Frullania blepharozia Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 18, 1884 (Spruce 1884). *** Frullania bogotensis Steph., Sp. Hepat. (Stephani) 4: 327, 1910 (Stephani 1910b). * Frullania brachycarpa Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: cciv, 1889 [1890] (Spruce 1889). *** Frullania confertiloba Steph., Sp. Hepat. (Stephani) 4: 326, 1910 (Stephani 1910b). *** Frullania decidua Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 30, 1884 (Spruce 1884). *** Frullania ecklonii (Spreng.) Gottsche, Lindenb. et Nees, Syn. Hepat. 3: 413, 1845 (Gottsche et al. 1845b). Bas.: Jungermannia ecklonii Spreng. Syst. Veg. (ed. 16) [Sprengel] 4 (2): 324, 1827 (Sprengel 1827b). * Frullania ecklonii var. robustior (Gottsche, Lindenb. et Nees) Sim, Trans. Roy. Soc. South Africa 15 (1): 39, 1926 (Sim 1926). Bas.: Frullania ecklonii a robustior Gottsche, Lindenb. et Nees, Syn. Hepat. 5: 771, 1847 (Gottsche et al. 1847). * Frullania ecklonii var. rufescens Gottsche, Lindenb. et Nees, Syn. Hepat. 5: 771, 1847 (Gottsche et al. 1847). * Frullania ecklonii var. tenerior (Gottsche, Lindenb. et Nees) Sim, Trans. Roy. Soc. South Africa 15 (1): 39, 1926 (Sim 1926). Bas.: Frullania ecklonii b tenerior Gottsche, Lindenb. et Nees, Syn. Hepat. 5: 771, 1847 (Gottsche et al. 1847). *** Frullania holostipula S.Hatt. et D.G.Griin, Misc. Bryol. Lichenol. 8 (3): 47, 1978 (Hattori and Griin 1978). *** Frullania megalostipa Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 15, 1884 (Spruce 1884). *** Frullania obscura (Sw.) Mont., Ann. Sci. Nat. Bot. (sér. 2) 14: 333, 1840 (Montagne 1840a). Bas.: Jungermannia obscura Sw., Fl. Ind. Occid. 3: 1869, 1806 (Swartz 1806). *** Frullania obscura var. spiniloba (Steph.) Hentschel et von Konrat, Phytotaxa 220 (2): 136, 2015 (Hentschel et al. 2015). Bas.: Frullania spiniloba Steph., Sp. Hepat. (Stephani) 4: 336, 1910 (Stephani 1910b). *** Frullania ringens Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 17, 1884 (Spruce 1884). *** Frullania rio-janeirensis (Raddi) Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 33 (7): 88, 1876 [1877] (Ångström 1876). Bas.: Frullanoides rio-janeirensis Raddi, Critt. Brasil.: 13, 1822 (Raddi 1822). *** Frullania sphaerocephala Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 17, 1884 (Spruce 1884). ** Frullania tunguraguana L.Clark et Frye, Bryologist 55 (2): 133, 1952 (Clark and Frye 1952). Nom. nov. pro Frullania brachyclada Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 15, 1884 (Spruce 1884), nom. illeg. 266 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** sect. Pluricarinatae (Yuzawa, Mues et S.Hatt.) Hentschel et von Konrat, Phytotaxa 220 (2): 129, 2015 (Hentschel et al. 2015). Bas.: Frullania ser. Pluricarinatae Yuzawa, Mues et S.Hatt., J. Hattori Bot. Lab. 63: 428, 1987 (Yuzawa et al. 1987). *** Frullania albertii Steph., Biblioth. Bot. 87 (2): 237, 1916 (Stephani 1916a). *** Frullania arsenii Steph., Sp. Hepat. (Stephani) 6: 530, 1924 (Stephani 1924). ** Frullania bonariensis M.E.Reiner, Bol. Soc. Argent. Bot. 25 (3/4): 313, 1988 (Reiner 1988). *** Frullania cuencensis Taylor, London J. Bot. 5: 406, 1846 (Taylor 1846b). *** Frullania depressa Mitt., J. Proc. Linn. Soc., Bot. 7 (27): 168, 1863 (Mitten 1863). *** Frullania dusenii Steph., Arch. Mus. Nac. Rio de Janeiro 13: 115 (9), 1905 (Stephani 1905c). *** Frullania gradsteinii Yuzawa, Mues et S.Hatt., J. Hattori Bot. Lab. 63: 429, 1987 (Yuzawa et al. 1987). *** Frullania haematocysta Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 54, 1884 (Spruce 1884). *** Frullania jelskii Loitl., Diagn. pl. nov.: 18, 1894 (Loitlesberger 1894). *** Frullania laxilora Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 26, 1884 (Spruce 1884). * Frullania laxilora var. crossii Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 27, 1884 (Spruce 1884). *** Frullania neurota Taylor, London J. Bot. 5: 400, 1846 (Taylor 1846b). *** Frullania planifolia Steph., Sp. Hepat. (Stephani) 4: 337, 1910 (Stephani 1910b). *** Frullania pluricarinata Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 168, 1864 (Gottsche 1864). ** Frullania sandvicensis Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 29 (4): 28, 1872 (Ångström 1872). *** Frullania standaertii Steph., Sp. Hepat. (Stephani) 4: 342, 1910 (Stephani 1910b). *** Frullania stenostipa Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 29, 1884 (Spruce 1884). *** Frullania tetraptera Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 9: 47, 1838 (Montagne 1838). ** Frullania trinervis (Lehm.) Drège, Flora, Beig. 26: 186, 1843 (Drège 1843). Bas.: Jungermannia trinervis Lehm., Linnaea 9 (4): 426, 1835 (Lehmann 1835). 186 *** Frullania winteri Steph., Sp. Hepat. (Stephani) 4: 338, 1910 (Stephani 1910b). *** Frullania winteri var. vanderhammenii (Haarbrink) Yuzawa, J. Hattori Bot. Lab. 70: 233, 1991 (Yuzawa 1991). Bas.: Frullania vanderhammenii Haarbrink, Lindbergia 7 (1): 56, 1981 (Haarbrink 1981). *** subg. Diastaloba Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 55, 1884 (Spruce 1884). ** Frullania antaresensis S.Hatt., J. Hattori Bot. Lab. 47: 92, 1980 (Hattori 1980d). 186 Frullania trinervis is possibly a species complex (Vanden Berghen 1976b). World checklist of hornworts and liverworts * *** ** ** ** ** ** ** * * *** *** *** ** ** ** 267 Frullania armatifolia Verd., Bull. Jard. Bot. Buitenzorg (sér. 3) 12 (1): 61, 1932 (Verdoorn 1932a). Frullania curvilobula Schäf.-Verw., D.F.Peralta et S.M.Siqueira, Phytotaxa 57 (4): 27, 2012 (Schäfer-Verwimp et al. 2012). Frullania gracilicaulis S.Hatt., J. Hattori Bot. Lab. 43: 421, 1977 [1978] (Hattori 1977b). Frullania humbertii Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 46 (1/2): 24, 1976 (Vanden Berghen 1976b). Frullania hypoleucula S.Hatt., J. Hattori Bot. Lab. 57: 413, 1984 (Hattori 1984a). Frullania incurva S.Hatt., J. Hattori Bot. Lab. 65: 431, 1988 (Hattori 1988a). Frullania klotzschii Nees, Sp. Hepat. (Stephani) 4: 558, 1911 (Stephani 1911e). Frullania letestui Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 46 (1/2): 38, 1976 (Vanden Berghen 1976b). Frullania miradorensis Lindenb. et Gottsche, Syn. Hepat. 5: 781, 1847 (Gottsche et al. 1847). Frullania odontostipa Spruce, Mem. Torrey Bot. Club 1 (3): 120, 1890 (Spruce 1890). Frullania pilibracteola S.Hatt., J. Hattori Bot. Lab. 43: 428, 1977 [1978] (Hattori 1977b). Frullania pilistipula Steph., Sp. Hepat. (Stephani) 4: 648, 1911 (Stephani 1911e). Frullania ramuligera (Nees) Mont., Ann. Sci. Nat. Bot. (sér. 2) 18: 14, 1842 (Montagne 1842b). Bas.: Jungermannia ramuligera Nees, Enum. Pl. Crypt. Javae: 52, 1830 (Nees 1830). Frullania subpilibracteola S.Hatt., J. Hattori Bot. Lab. 43: 434, 1977 [1978] (Hattori 1977b). Frullania subtilissima (Nees ex Mont.) Lindenb., Syn. Hepat. 3: 443, 1845 (Gottsche et al. 1845b). Bas.: Frullania atrata β subtilissima Nees ex Mont., Ann. Sci. Nat. Bot. (sér. 2) 14: 333, 1840 (Montagne 1840a). Frullania vandenberghenii Pócs, Acta Bot. Acad. Sci. Hung. 25 (3/4): 229, 1979 [1980] (Bizot and Pócs 1979). Nom. nov. pro Frullania epiphylla Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 46 (1/2): 30, 1976 (Vanden Berghen 1976b), nom. illeg. grp. Diastaloba I ** Frullania akiyamae S.Hatt., J. Hattori Bot. Lab. 60: 240, 1986 (Hattori 1986b). ** Frullania apiculata (Reinw., Blume et Nees) Nees, Syn. Hepat. 3: 452, 1845 (Gottsche et al. 1845b). Bas.: Jungermannia apiculata Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 222, 1824 [1825] (Reinwardt et al. 1824a). ** Frullania apiculata var. goebelii Schifn., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 60 (2): 222, 1893 (Schifner 1893a). *** Frullania armitiana Steph., Sp. Hepat. (Stephani) 4: 538, 1911 (Stephani 1911e). 268 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Frullania armitiana var. inlexula S.Hatt., J. Hattori Bot. Lab. 65: 415, 1988 (Hattori 1988a). ** Frullania aterrima (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 3: 450, 1845 (Gottsche et al. 1845b). Bas.: Jungermannia aterrima Hook.f. et Taylor, London J. Bot. 3: 395, 1844 (Hooker and Taylor 1844a). *** Frullania attenuata Steph., Sp. Hepat. (Stephani) 4: 538, 1911 (Stephani 1911e). *** Frullania bella Steph., Sp. Hepat. (Stephani) 4: 643, 1911 (Stephani 1911e). ** Frullania changii S.Hatt. et C.Gao, J. Jap. Bot. 60 (1): 1, 1985 (Hattori and Gao 1985). * Frullania claviloba Steph., Sp. Hepat. (Stephani) 4: 651, 1911 (Stephani 1911e). 187 *** Frullania colliculosa von Konrat, Braggins, Hentschel et Heinrichs, Nova Hedwigia 91 (3/4): 494, 2010 (von Konrat et al. 2010b). *** Frullania cordistipula (Reinw., Blume et Nees) Nees, Voy. Amér. Mérid. 7 (2): 68, 1839 (Montagne 1839a). Bas.: Jungermannia cordistipula Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 220, 1824 [1825] (Reinwardt et al. 1824a). 188 ** Frullania cordistipula var. dentistipula S.Hatt., J. Hattori Bot. Lab. 60: 242, 1986 (Hattori 1986b). * Frullania cordistipula var. mutica (Gottsche, Lindenb. et Nees) Schifn., Consp. Hepat. Arch. Ind.: 323, 1898 (Schifner 1898b). Bas.: Frullania cordistipula β mutica Gottsche, Lindenb. et Nees, Syn. Hepat. 3: 454, 1845 (Gottsche et al. 1845b). * Frullania crenatiloba Steph., Sp. Hepat. (Stephani) 6: 551, 1924 (Stephani 1924). 189 *** Frullania curvistipula Steph., Sp. Hepat. (Stephani) 4: 548, 1911 (Stephani 1911e). ** Frullania curvistipula var. falcatidentata S.Hatt., Misc. Bryol. Lichenol. 9 (6): 124, 1982 (Hattori 1982a). ** Frullania curvistipula var. lamii Verd., Ann. Bryol., Suppl. 1: 91, 1930 (Verdoorn 1930c). ** Frullania cuspidifolia Steph., Sp. Hepat. (Stephani) 4: 543, 1911 (Stephani 1911e). * Frullania degelii S.W.Arnell, Svensk Bot. Tidskr. 53 (4): 503, 1959 (Arnell 1959). ** Frullania dentifera S.Hatt. et Streimann, J. Hattori Bot. Lab. 59: 102, 1985 (Hattori and Streimann 1985). *** Frullania dentiloba S.Hatt., J. Jap. Bot. 50 (6): 161, 1975 (Hattori 1975a). ** Frullania durifolia Steph., Hedwigia 33 (3): 162, 1894 (Stephani 1894d). ** Frullania exilis Taylor, London J. Bot. 5: 405, 1846 (Taylor 1846b). ** Frullania gabonensis Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 46 (1/2): 81, 1976 (Vanden Berghen 1976b). *** Frullania gracilis (Reinw., Blume et Nees) Nees, Syn. Hepat. 3: 452, 1845 (Gottsche et al. 1845b). Bas.: Jungermannia gracilis Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 221, 1824 [1825] (Reinwardt et al. 1824a). 187 Frullania claviloba is possibly conspeciic with Frullania gracilis (Söderström et al. 2010a). 188 Frullania cordistipula is a species complex also including Frullania serrata. 189 Frullania crenatiloba is conspeciic with Frullania apiculata in Verdoorn (1930c), but it was accepted by So and Wang (2006). World checklist of hornworts and liverworts * ** ** *** ** ** *** *** ** *** ** *** ** ** ** ** ** * ** ** ** ** ** ** *** 269 Frullania gracilis var. brevior (Gottsche, Lindenb. et Nees) Schifn., Consp. Hepat. Arch. Ind.: 327, 1898 (Schifner 1898b). Bas.: Frullania gracilis β brevior Gottsche, Lindenb. et Nees, Syn. Hepat. 3: 453, 1845 (Gottsche et al. 1845b). Frullania gracilis var. vittata S.Hatt., J. Hattori Bot. Lab. 60: 243, 1986 (Hattori 1986b). Frullania gracilis subsp. zennoskei S.Hatt. et haithong, J. Hattori Bot. Lab. 44: 183, 1978 (Hattori and haithong 1978b). Frullania hasskarliana Lindenb., Syn. Hepat. 3: 453, 1845 (Gottsche et al. 1845b). Frullania hasskarliana var. gracilis S.Hatt., J. Hattori Bot. Lab. 60: 243, 1986 (Hattori 1986b). Frullania hasskarliana var. parvidentata S.Hatt., J. Hattori Bot. Lab. 60: 245, 1986 (Hattori 1986b). Frullania hattorii von Konrat et Braggins, New Zealand J. Bot. 41 (1): 56, 2003 (von Konrat and Braggins 2003). Frullania hodgsoniae von Konrat, Braggins, Hentschel et Heinrichs, Nova Hedwigia 91 (3/4): 492, 2010 (von Konrat et al. 2010b). Nom. nov. pro Frullania aterrima var. lepida E.A.Hodgs., Trans. & Proc. Roy. Soc. New Zealand 77 (3): 386, 1949 (Hodgson 1949). Frullania hottana S.Hatt., J. Hattori Bot. Lab. 40: 479, 1976 (Hattori 1976d). Frullania inconstans Verd., Ann. Bryol., Suppl. 1: 83, 1930 (Verdoorn 1930c). Frullania inconstans var. grossedentata Kamim. et S.Hatt., J. Hattori Bot. Lab. 37: 530, 1973 (Hattori and Kamimura 1973). Frullania johnsonii Steph., Hedwigia 33 (3): 163, 1894 (Stephani 1894d). Frullania macgregorii Steph., Hedwigia 33 (3): 154, 1894 (Stephani 1894d). Frullania macgregorii var. rostellula (S.Hatt.) S.Hatt., J. Hattori Bot. Lab. 51: 220, 1982 (Hattori 1982d). Bas.: Frullania reimersii var. rostellula S.Hatt., J. Hattori Bot. Lab. 38: 258, 1974 (Hattori 1974c). Frullania madens Steph., Sp. Hepat. (Stephani) 6: 553, 1924 (Stephani 1924). Frullania mehrana S.Hatt., Recent Adv. Bot.: 66, 1976 (Hattori 1976e). Frullania motoyana Steph., Sp. Hepat. (Stephani) 4: 646, 1911 (Stephani 1911e). Frullania multilacera Steph., Sp. Hepat. (Stephani) 4: 650, 1911 (Stephani 1911e). 190 Frullania multilacera subsp. gracilior S.Hatt., Mem. New York Bot. Gard. 45: 547, 1987 (Hattori 1987a). Frullania multilacera var. lacerissima S.Hatt., J. Hattori Bot. Lab. 39: 294, 1975 (Hattori 1975d). Frullania multilaceroides S.Hatt., Mem. New York Bot. Gard. 45: 549, 1987 (Hattori 1987a). Frullania neosheana S.Hatt., J. Hattori Bot. Lab. 45: 350, 1979 (Hattori 1979b). Frullania papillata Steph., Sp. Hepat. (Stephani) 4: 615, 1911 (Stephani 1911e). Frullania pulogensis Steph., Sp. Hepat. (Stephani) 4: 545, 1911 (Stephani 1911e). Frullania purpurea Steph., Sp. Hepat. (Stephani) 4: 626, 1911 (Stephani 1911e). 190 Frullania multilacera is possibly conspeciic with Frullania vaga. 270 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Frullania reimersii Verd., Ann. Bryol., Suppl. 1: 84, 1930 (Verdoorn 1930c). ** Frullania saepisdentata S.Hatt. et Streimann, J. Hattori Bot. Lab. 59: 116, 1985 (Hattori and Streimann 1985). ** Frullania schifneri Verd., Ann. Bryol. 2: 150, 1929 (Verdoorn 1929a). ** Frullania schusterana S.Hatt., J. Hattori Bot. Lab. 36: 411, 1972 [1973] (Hattori 1972b). ** Frullania seriatifolia Steph., Hedwigia 33 (3): 167, 1894 (Stephani 1894d). ** Frullania serrata Gottsche, Syn. Hepat. 3: 453, 1845 (Gottsche et al. 1845b). ** Frullania serrata var. ceramensis S.Hatt., J. Hattori Bot. Lab. 60: 247, 1986 (Hattori 1986b). ** Frullania serrata subsp. grolleana (S.Hatt.) S.Hatt., J. Hattori Bot. Lab. 51: 225, 1982 (Hattori 1982d). Bas.: Frullania grolleana S.Hatt., J. Hattori Bot. Lab. 36: 416, 1972 [1973] (Hattori 1972b). ** Frullania serrata var. hamatispina (S.Hatt.) S.Hatt., J. Hattori Bot. Lab. 51: 225, 1982 (Hattori 1982d). Bas.: Frullania serrata subsp. hamatispina S.Hatt., J. Hattori Bot. Lab. 38: 262, 1974 (Hattori 1974c). ** Frullania serrata var. pertenuis (Nees) Schifn., Consp. Hepat. Arch. Ind.: 342, 1898 (Schifner 1898b). Bas.: Jungermannia cordistipula γ pertenuis Nees, Enum. Pl. Crypt. Javae: 49, 1830 (Nees 1830). ** Frullania serrata subsp. spinistipula S.Hatt., J. Hattori Bot. Lab. 51: 225, 1982 (Hattori 1982d). Nom. nov. pro Frullania spinistipula S.Hatt., J. Hattori Bot. Lab. 36: 413, 1972 [1973] (Hattori 1972b), nom. illeg. ** Frullania setacea S.Hatt., J. Hattori Bot. Lab. 65: 447, 1988 (Hattori 1988a). ** Frullania sheana S.Hatt., J. Hattori Bot. Lab. 45: 356, 1979 (Hattori 1979b). ** Frullania simmondsii Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 109, 1914 (Stephani and Watts 1914). *** Frullania sinuata Sande Lac., Ned. Kruidk. Arch. 3: 424, 1854 [1855] (Sande Lacoste 1854). ** Frullania steereana S.Hatt., Mem. New York Bot. Gard. 45: 553, 1987 (Hattori 1987a). ** Frullania stipatiloba Steph., Hedwigia 33 (3): 168, 1894 (Stephani 1894d). ** Frullania subdentata Steph., Sp. Hepat. (Stephani) 4: 545, 1911 (Stephani 1911e). ** Frullania subdentata var. latistipula (S.Hatt.) S.Hatt., J. Hattori Bot. Lab. 51: 228, 1982 (Hattori 1982d). Bas.: Frullania curvistipula var. latistipula S.Hatt., J. Hattori Bot. Lab. 44: 530, 1978 (Hattori 1978b). ** Frullania submultilacera S.Hatt. et Koike, J. Hattori Bot. Lab. 75: 190, 1994 (Koike 1994). ** Frullania subocellata S.Hatt., J. Hattori Bot. Lab. 60: 248, 1986 (Hattori 1986b). *** Frullania taxodiocola R.M.Schust., Phytologia 53 (5): 364, 1983 (Schuster 1983b). *** Frullania ternatensis Gottsche, Syn. Hepat. 4: 465, 1846 (Gottsche et al. 1846). ** Frullania ternatensis var. non-appendiculata S.Hatt., J. Hattori Bot. Lab. 38: 174, 1974 (Hattori 1974a). World checklist of hornworts and liverworts 271 *** Frullania trichodes Mitt., Bonplandia 10 (2): 19, 1862 (Mitten 1862). ** Frullania vaga Mitt., Fl. vit.: 418, 1871 [1873] (Mitten 1871). *** Frullania vaginata (Sw.) Nees, Syn. Hepat. 4: 465, 1846 (Gottsche et al. 1846). Bas.: Jungermannia vaginata Sw., Meth. Musc.: 35, 1781 (Swartz 1781). * Frullania vaginata var. nigricans (Gottsche, Lindenb. et Nees) Schifn., Consp. Hepat. Arch. Ind.: 348, 1898 (Schifner 1898b). Bas.: Frullania vaginata β nigricans Gottsche, Lindenb. et Nees, Syn. Hepat. 4: 465, 1846 (Gottsche et al. 1846). * Frullania van-zantenii Kamim. et S.Hatt., J. Hattori Bot. Lab. 37: 528, 1973 (Hattori and Kamimura 1973). 191 ** Frullania venusta S.Hatt., J. Hattori Bot. Lab. 38: 217, 1974 (Hattori 1974d). ** Frullania verdoorniana S.Hatt., J. Hattori Bot. Lab. 37: 122, 1973 (Hattori 1973b). *** Frullania vitalii Yuzawa et S.Hatt., J. Jap. Bot. 63 (1): 30, 1988 (Yuzawa and Hattori 1988a). ** Frullania vittata S.Hatt., J. Hattori Bot. Lab. 38: 270, 1974 (Hattori 1974c). ** Frullania vivipara Pócs, Fieldiana, Bot. (n.ser.) 47: 151, 2008 (Pócs 2008a). ** Frullania wairua von Konrat et Braggins, New Zealand J. Bot. 43 (4): 886, 2005 (von Konrat and Braggins 2005). ** Frullania warnckeana S.Hatt., J. Hattori Bot. Lab. 38: 213, 1974 (Hattori 1974d). ** Frullania warnckeana var. dentosa S.Hatt., Misc. Bryol. Lichenol. 7 (8): 162, 1977 (Hattori 1977a). *** *** ** *** *** ** ** grp. Diastaloba II Frullania baumannii S.Hatt., J. Hattori Bot. Lab. 43: 410, 1977 [1978] (Hattori 1977b). Frullania congesta Gottsche, Lindenb. et Nees, Syn. Hepat. 3: 451, 1845 (Gottsche et al. 1845b). Nom. nov. pro Jungermannia congesta Hook.f. et Taylor, London J. Bot. 3: 396, 1844 (Hooker and Taylor 1844a), nom. illeg. Frullania ocellata S.Hatt. et Kamim., J. Hattori Bot. Lab. 37: 531, 1973 (Hattori and Kamimura 1973). Frullania ptychantha Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 257, 1843 (Montagne 1843). Frullania repandistipula Sande Lac., Ned. Kruidk. Arch. 3: 422, 1854 [1855] (Sande Lacoste 1854). Frullania repandistipula subsp. queenslandica S.Hatt., Mem. New York Bot. Gard. 45: 550, 1987 (Hattori 1987a). Frullania repandistipula subsp. spinibractea S.Hatt., Bull. Natl. Sci. Mus. Tokyo, B 1 (4): 150, 1975 (Hattori 1975f). grp. Diastaloba III ** Frullania grossiclava Steph., Sp. Hepat. (Stephani) 4: 384, 1910 (Stephani 1910b). 191 Frullania van-zantenii is possibly conspeciic with Frullania serrata. 272 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Frullania loricata Pearson, Forh. Vidensk.-Selsk. Kristiania 1890 (2): 6, 1891 (Pearson 1891). * Frullania loricata var. laxa Pearson, Forh. Vidensk.-Selsk. Kristiania 1890 (2): 8, 1891 (Pearson 1891). *** Frullania usambarana Schifn., Hedwigia 33 (3): 160, 1894 (Stephani 1894d). ** Frullania usambarana var. reducta Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 46 (1/2): 91, 1976 (Vanden Berghen 1976b). * *** *** *** *** * ** * grp. Diastaloba IV Frullania brunea (Spreng.) Drège, Flora, Beig. 26: 186, 1843 (Drège 1843). Bas.: Jungermannia brunea Spreng. Syst. Veg. (ed. 16) [Sprengel] 4 (2): 325, 1827 (Sprengel 1827b). 192 Frullania caulisequa (Nees) Mont., Ann. Sci. Nat. Bot. (sér. 2) 12: 51, 1839 (Montagne 1839c). Bas.: Jungermannia caulisequa Nees, Fl. Bras. (Martius) 1 (1): 373, 1833 (Nees 1833a). Frullania grossifolia Steph., Sp. Hepat. (Stephani) 4: 633, 1911 (Stephani 1911e). Frullania hypoleuca Nees, Observ. bot.: 470, 1843 (Gottsche et al. 1843). Frullania lindenbergii Lehm., Nov. Stirp. Pug. 8: 17, 1844 (Lehmann 1844). Frullania lindenbergii var. fusca Gottsche, Lindenb. et Nees, Syn. Hepat. 5: 780, 1847 (Gottsche et al. 1847). Frullania ponapensis S.Hatt. et Koike, J. Hattori Bot. Lab. 75: 186, 1994 (Koike 1994). Frullania tricarinata Sande Lac., Plagiochila Sandei: 10, 1856 (Sande Lacoste 1856c). 193 subg. Diversitextae (Kamim.) S.Hatt., J. Hattori Bot. Lab. 59: 154, 1985 (Hattori and Lin 1985a). Bas.: Frullania subsect. Diversitextae Kamim., J. Hattori Bot. Lab. 24: 80, 1961 (Kamimura 1961). *** Frullania diversitexta Steph., Bull. Herb. Boissier 5 (2): 89, 1897 (Stephani 1897b). * *** subg. Frullania * Frullania amamiensis Kamim., Bull. Kochi Gakuen Jun. Coll. 1: 51, 1970 (Kamimura 1970). ** Frullania amplicrania Steph., Sp. Hepat. (Stephani) 4: 404, 1910 (Stephani 1910b). *** Frullania ampullifera J.B.Jack et Steph., Hedwigia 33 (3): 139, 1894 (Stephani 1894d). *** Frullania anderssonii Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 30 (5): 144, 1873 (Ångström 1873). ** Frullania angustistipa Steph., Akad. Wiss. Wien, Math.-Naturwiss. Kl., Denkschr. 81: 297, 1907 (Stephani 1907a). 192 Frullania brunea belongs to the Frullania lindenbergii complex (Vanden Berghen 1976b). 193 Frullania tricarinata is possibly conspeciic with Frullania hypoleuca (Söderström et al. 2010a). World checklist of hornworts and liverworts 273 ** Frullania aposinensis S.Hatt. et P.J.Lin, J. Hattori Bot. Lab. 59: 131, 1985 (Hattori and Lin 1985a). Nom. nov. pro Frullania chinensis Steph., Sp. Hepat. (Stephani) 4: 469, 1911 (Stephani 1911e), nom. illeg. ** Frullania appendistipula S.Hatt., J. Hattori Bot. Lab. 36: 424, 1972 [1973] (Hattori 1972b). ** Frullania appendistipula var. spinifera S.Hatt., J. Hattori Bot. Lab. 38: 226, 1974 (Hattori 1974c). ** Frullania auriculata S.Hatt., Bull. Natl. Sci. Mus. Tokyo, B 11 (1): 11, 1985 (Hattori 1985). ** Frullania benjaminiana Inoue, Bull. Natl. Sci. Mus. Tokyo, B 1 (3): 109, 1975 (Hattori 1975b). ** Frullania bergmanii S.Hatt., J. Hattori Bot. Lab. 38: 192, 1974 (Hattori 1974d). *** Frullania berthoumieui Steph., Hedwigia 33 (3): 140, 1894 (Stephani 1894d). ** Frullania bhutanensis S.Hatt., Fl. E. Himalaya 2: 232, 1971 (Hattori 1971a). ** Frullania blastopetala S.Hatt., J. Hattori Bot. Lab. 57: 407, 1984 (Hattori 1984a). *** Frullania bonincola S.Hatt., J. Hattori Bot. Lab. 44: 551, 1978 (Hattori 1978b). Nom. nov. pro Frullania viridis Horik., Sci. Rep. Tôhoku Imp. Univ., Ser. 4, Biol. 5 (4): 646, 1929 [1930] (Horikawa 1929c), nom. illeg. * Frullania brevicalycina Steph., Hedwigia 33 (3): 141, 1894 (Stephani 1894d). ** Frullania brittoniae A.Evans, Trans. Connecticut Acad. Arts 10 (1): 15, 1899 (Evans 1899). *** Frullania bullata Steph., Sp. Hepat. (Stephani) 4: 371, 1910 (Stephani 1910b). *** Frullania cafraria Steph., Hedwigia 33 (3): 141, 1894 (Stephani 1894d). *** Frullania calcarata Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 30 (5): 137, 1873 (Ångström 1873). ** Frullania carrii Kamim. et S.Hatt., J. Hattori Bot. Lab. 37: 520, 1973 (Hattori and Kamimura 1973). ** Frullania chenii S.Hatt. et P.J.Lin, J. Jap. Bot. 60 (4): 106, 1985 (Hattori and Lin 1985b). ** Frullania chilensis Steph., Hedwigia 33 (3): 142, 1894 (Stephani 1894d). *** Frullania chodatii Beauverd, Sp. Hepat. (Stephani) 6: 533, 1924 (Stephani 1924). Based on: Frullania grandistipula Steph., Sp. Hepat. (Stephani) 6: 533, 1924 (Stephani 1924), nom. inval. *** Frullania cobrensis Gottsche, Hedwigia 33 (3): 142, 1894 (Stephani 1894d). ** Frullania consociata Steph., Sp. Hepat. (Stephani) 4: 461, 1911 (Stephani 1911e). * Frullania contracta Steph., Sp. Hepat. (Stephani) 4: 469, 1911 (Stephani 1911e). 194 * Frullania cornuta Steph., Sp. Hepat. (Stephani) 4: 467, 1911 (Stephani 1911e). 195 *** Frullania crassitexta Steph., Sp. Hepat. (Stephani) 4: 423, 1910 (Stephani 1910b). *** Frullania crispiplicata Yuzawa et S.Hatt., J. Jap. Bot. 58 (2): 43, 1983 (Yuzawa and Hattori 1983). 194 Frullania contracta is conspeciic with Frullania squarrosa in Verdoorn (1930c), but it was accepted by Hattori (1986e). 195 Frullania cornuta is closely related to the very variable Frullania ornithocephala. 274 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Frullania cristata S.Hatt., J. Hattori Bot. Lab. 49: 165, 1981 (Hattori 1981a). * Frullania cuneiloba Nees, Syn. Hepat. 3: 427, 1845 (Gottsche et al. 1845b). ** Frullania cyparioides (Schwägr.) Nees, Naturgesch. Eur. Leberm. 3: 210, 1838 (Nees 1838b). Bas.: Jungermannia cyparioides Schwägr., Hist. Musc. Hepat. Prodr.: 14, 1814 (Schwägrichen 1814). *** Frullania davurica Hampe ex Gottsche, Lindenb. et Nees, Syn. Hepat. 3: 422, 1845 (Gottsche et al. 1845b). ** Frullania debilis Steph. ex S.Hatt., J. Hattori Bot. Lab. 38: 190, 1974 (Hattori 1974d). ** Frullania deppii Lehm., Nov. Stirp. Pug. 8: 15, 1844 (Lehmann 1844). ** Frullania diptera (Lehm.) Drège, Flora, Beig. 26: 186, 1843 (Drège 1843). Bas.: Jungermannia diptera Lehm., Linnaea 9 (4): 425, 1835 (Lehmann 1835). *** Frullania duthiana Steph., Sp. Hepat. (Stephani) 4: 351, 1910 (Stephani 1910b). ** Frullania duthiana var. appendiculata S.Hatt., Fl. E. Himalaya 2: 234, 1971 (Hattori 1971a). ** Frullania duthiana var. laevis S.Hatt., Bull. Univ. Mus. Univ. Tokyo 8: 233, 1975 (Hattori 1975e). ** Frullania duthiana var. szechuanensis S.Hatt. et C.Gao, J. Jap. Bot. 60 (1): 2, 1985 (Hattori and Gao 1985). ** Frullania echinantha S.Hatt., J. Hattori Bot. Lab. 38: 233, 1974 (Hattori 1974c). ** Frullania echinatella S.Hatt., J. Hattori Bot. Lab. 65: 423, 1988 (Hattori 1988a). * Frullania elegans Lehm., Nov. Stirp. Pug. 10: 16, 1857 (Lehmann 1857). *** Frullania elephantum S.Hatt., J. Hattori Bot. Lab. 43: 416, 1977 [1978] (Hattori 1977b). ** Frullania epiphylla S.Hatt., J. Hattori Bot. Lab. 38: 235, 1974 (Hattori 1974c). * Frullania epiphylla subsp. ijiensis S.Hatt., Bull. Natl. Sci. Mus. Tokyo, B 11 (1): 14, 1985 (Hattori 1985). ** Frullania erostrata S.Hatt., J. Hattori Bot. Lab. 38: 237, 1974 (Hattori 1974c). * Frullania esenbeckiana Beauverd, Sp. Hepat. (Stephani) 6: 570, 1924 (Stephani 1924). Nom. nov. pro Frullania grossiloba Steph., Sp. Hepat. (Stephani) 4: 531, 1911 (Stephani 1911e), nom. illeg. ** Frullania evelynae S.Hatt. et haithong, J. Hattori Bot. Lab. 44: 179, 1978 (Hattori and haithong 1978b). * Frullania evelynae var. devendrae Sushil K.Singh et Barbhuiya, Taiwania 57 (2): 109, 2012 (Singh and Barbhuiya 2012). * Frullania evelynae var. srivastavae Sushil K.Singh et Barbhuiya, Taiwania 57 (2): 112, 2012 (Singh and Barbhuiya 2012). *** Frullania eymae S.Hatt., J. Hattori Bot. Lab. 39: 284, 1975 (Hattori 1975d). ** Frullania eymae var. crispidentata S.Hatt. et Streimann, J. Hattori Bot. Lab. 59: 105, 1985 (Hattori and Streimann 1985). *** Frullania falciloba Lehm., Nov. Stirp. Pug. 8: 20, 1844 (Lehmann 1844). ** Frullania falsicornuta S.Hatt., J. Hattori Bot. Lab. 60: 212, 1986 (Hattori 1986e). ** Frullania fauriana Steph., Hedwigia 33 (3): 144, 1894 (Stephani 1894d). World checklist of hornworts and liverworts 275 ** Frullania fengyangshanensis R.L.Zhu et M.L.So, Bryologist 100 (3): 356, 1997 (Zhu and So 1997b). ** Frullania ferdinandi-muelleri Steph., Sp. Hepat. (Stephani) 4: 417, 1910 (Stephani 1910b). * Frullania lexuosa S.Hatt., J. Hattori Bot. Lab. 54: 146, 1983 (Hattori 1983). 196 ** Frullania fuscovirens Steph., Sp. Hepat. (Stephani) 4: 401, 1910 (Stephani 1910b). ** Frullania fuscovirens var. gemmipara (R.M.Schust. et S.Hatt.) S.Hatt. et P.J.Lin, J. Hattori Bot. Lab. 59: 135, 1985 (Hattori and Lin 1985a). Bas.: Frullania gemmipara R.M.Schust. et S.Hatt., J. Hattori Bot. Lab. 44: 547, 1978 (Hattori 1978b). ** Frullania gaoligongensis X.L.Bai et C.Gao, Hikobia 13 (1): 87, 1999 (Bai and Gao 1999). ** Frullania gemmulosa S.Hatt. et haithong, J. Hattori Bot. Lab. 43: 449, 1977 [1978] (Hattori et al. 1977). *** Frullania gigantea Steph., Sp. Hepat. (Stephani) 4: 467, 1911 (Stephani 1911e). ** Frullania giraldiana C.Massal., Hepat. Shen-si: 41, 1897 (Massalongo 1897). 197 ** Frullania giraldiana var. handelii (Verd.) S.Hatt., J. Hattori Bot. Lab. 36: 123, 1972 [1973] (Hattori 1972c). Bas.: Frullania nepalensis var. handelii Verd., Symb. Sin. 5: 41, 1930 (Nicholson et al. 1930). ** Frullania globosa S.Hatt. et Streimann, J. Hattori Bot. Lab. 59: 107, 1985 (Hattori and Streimann 1985). ** Frullania grandilobula S.Hatt. et Piippo, Acta Bot. Fenn. 133: 45, 1986 (Hattori and Piippo 1986). *** Frullania grandistipula Lindenb., Syn. Hepat. 3: 430, 1845 (Gottsche et al. 1845b). ** Frullania hainanensis S.Hatt. et P.J.Lin, J. Jap. Bot. 61 (10): 307, 1986 (Hattori and Lin 1986). ** Frullania hamatiloba Steph., Sp. Hepat. (Stephani) 4: 400, 1910 (Stephani 1910b). ** Frullania handelii Verd., Symb. Sin. 5: 36, 1930 (Nicholson et al. 1930). ** Frullania handel-mazzettii S.Hatt., J. Hattori Bot. Lab. 49: 150, 1981 (Hattori 1981a). ** Frullania hattoriantha Udar et V.Nath, Misc. Bryol. Lichenol. 9 (2): 44, 1981 (Udar and Nath 1981). * Frullania hebridensis Steph., Sp. Hepat. (Stephani) 4: 469, 1911 (Stephani 1911e). 198 ** Frullania hicksiae S.Hatt., Cryptog. Bryol. Lichénol. 5 (1/2): 182, 1984 (Hattori 1984c). ** Frullania higuchii Yuzawa, Koike et S.Hatt., J. Hattori Bot. Lab. 75: 194, 1994 (Yuzawa and Koike 1994). ** Frullania hiroshii S.Hatt., Bull. Natl. Sci. Mus. Tokyo, B 6 (1): 34, 1980 (Hattori 1980b). * Frullania hirtilora Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 35, 1884 (Spruce 1884). 196 Frullania lexuosa is possibly conspeciic with Frullania rubella. 197 Frullania giraldiana belongs to the Frullania nepalensis complex (Hattori 1973b). 198 Frullania hebridensis belongs to the Frullania ericoides species complex (Verdoorn 1930c, 1930a). 276 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Frullania howeana Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 107, 1914 (Stephani and Watts 1914). ** Frullania ignatovii Sofronova, Mamontov et Potemkin, Novosti Sist. Nizš. Rast. 47: 335, 2013 (Sofronova et al. 2013). ** Frullania incisoduthiana S.Hatt., J. Hattori Bot. Lab. 46: 391, 1979 (Mizutani 1979c). * Frullania incisoduthiana var. parva S.Hatt., J. Hattori Bot. Lab. 46: 391, 1979 (Mizutani 1979c). * Frullania incisostipula Steph., Sp. Hepat. (Stephani) 6: 541, 1924 (Stephani 1924). ** Frullania inlexiloba S.Hatt., J. Hattori Bot. Lab. 57: 415, 1984 (Hattori 1984a). ** Frullania inouei S.Hatt., Bull. Natl. Sci. Mus. Tokyo, B 6 (1): 36, 1980 (Hattori 1980b). *** Frullania irregularis S.Hatt. et Piippo, Acta Bot. Fenn. 133: 46, 1986 (Hattori and Piippo 1986). *** Frullania jackii Gottsche, Hepat. Eur., Leberm. 29-30: no. 294, 1863 (Gottsche and Rabenhorst 1863). ** Frullania jacobsii S.Hatt., Bull. Natl. Sci. Mus. Tokyo, B 12 (4): 129, 1986 (Hattori 1986c). * Frullania jacquinotii Gottsche, J. Bot. (Morot) 12: 138, 1898 (Bescherelle 1898). ** Frullania jovetiana von Konrat et Hentschel, Phytotaxa 220 (2): 135, 2015 (Hentschel et al. 2015). Based on: Frullania pseudericoides S.Hatt., Bull. Natl. Sci. Mus. Tokyo, B 12 (4): 132, 1986 (Hattori 1986c), nom. illeg. ** Frullania kagoshimensis Steph., Sp. Hepat. (Stephani) 4: 353, 1910 (Stephani 1910b). ** Frullania kagoshimensis subsp. hunanensis (S.Hatt.) S.Hatt. et P.J.Lin, J. Hattori Bot. Lab. 59: 137, 1985 (Hattori and Lin 1985a). Bas.: Frullania hunanensis S.Hatt., J. Hattori Bot. Lab. 49: 152, 1981 (Hattori 1981a). ** Frullania kalimantanensis S.Hatt., Bull. Natl. Sci. Mus. Tokyo, B 12 (4): 127, 1986 (Hattori 1986c). ** Frullania kashyapii Verd., Ann. Bryol. 5: 162, 1932 (Dixon et al. 1932). ** Frullania kitagawana S.Hatt., J. Hattori Bot. Lab. 57: 417, 1984 (Hattori 1984a). ** Frullania laeviperiantha X.L.Bai et C.Gao, Phytotaxa 220 (2): 136, 2015 (Hentschel et al. 2015). Based on: Frullania laeviperiantha X.L.Bai et C.Gao, Nova Hedwigia 70 (1/2): 135, 2000 (Bai and Gao 2000), nom. inval. * Frullania lancistyla Steph., Sp. Hepat. (Stephani) 4: 389, 1910 (Stephani 1910b). * Frullania latilora Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 34, 1884 (Spruce 1884). ** Frullania latogaleata Herzog, Trans. Brit. Bryol. Soc. 1 (3): 189, 1949 (Herzog 1949b). ** Frullania lepida S.Hatt. et Piippo, Acta Bot. Fenn. 133: 48, 1986 (Hattori and Piippo 1986). * Frullania levieri Steph., Sp. Hepat. (Stephani) 4: 388, 1910 (Stephani 1910b). ** Frullania linii S.Hatt., J. Hattori Bot. Lab. 49: 155, 1981 (Hattori 1981a). * Frullania longistyla Yuzawa et S.Hatt., J. Jap. Bot. 63 (11): 361, 1988 (Yuzawa and Hattori 1988b). 199 199 Frullania longistyla is closely related to Frullania brevicalycina (Yuzawa and Hattori 1988b). World checklist of hornworts and liverworts 277 * Frullania ludoviciae Steph., Rev. Bryol. 35 (2): 29, 1908 (Stephani 1908l). 200 ** Frullania lushanensis S.Hatt. et P.J.Lin, J. Hattori Bot. Lab. 59: 137, 1985 (Hattori and Lin 1985a). * Frullania macularis Taylor, London J. Bot. 5: 403, 1846 (Taylor 1846b). ** Frullania maymyoensis Svihla, Bryologist 61 (4): 376, 1958 [1959] (Svihla 1958). ** Frullania microauriculata Verd., Ann. Bryol. 2: 126, 1929 (Verdoorn 1929a). ** Frullania microauriculata var. rotundior Verd., Ann. Jard. Bot. Buitenzorg 40: 141, 1929 (Verdoorn 1929b). ** Frullania microrhyncha L.Clark et Svihla, Bryologist 53 (1): 63, 1950 (Clark and Svihla 1950). ** Frullania mizoramensis Sushil K.Singh et Barbhuiya, Taiwania 57 (2): 106, 2012 (Singh and Barbhuiya 2012). * Frullania montana Steph., Sp. Hepat. (Stephani) 4: 455, 1911 (Stephani 1911e). 201 ** Frullania multituberculata Hentschel et von Konrat, Phytotaxa 220 (2): 136, 2015 (Hentschel et al. 2015). Nom. nov. pro Frullania kalimantanensis Piippo et S.Hatt., J. Hattori Bot. Lab. 72: 117, 1992 (Piippo and Tan 1992), nom. illeg. ** Frullania mutilata Steph., Sp. Hepat. (Stephani) 4: 673, 1911 (Stephani 1911e). ** Frullania nadeaudii Steph., Sp. Hepat. (Stephani) 4: 465, 1911 (Stephani 1911e). *** Frullania nepalensis (Spreng.) Lehm. et Lindenb., Syn. Hepat. 3: 422, 1845 (Gottsche et al. 1845b). Bas.: Jungermannia nepalensis Spreng. Syst. Veg. (ed. 16) [Sprengel] 4 (2): 324, 1827 (Sprengel 1827b). 202 *** Frullania nicholsonii E.A.Hodgs., Trans. & Proc. Roy. Soc. New Zealand 77 (3): 368, 1949 (Hodgson 1949). Nom. nov. pro Frullania berggrenii W.E.Nicholson, Bryologist 28 (2): 17, 1925 (Nicholson 1925), nom. illeg. ** Frullania nigricaulis (Reinw., Blume et Nees) Nees, Syn. Hepat. 3: 457, 1845 (Gottsche et al. 1845b). Bas.: Jungermannia nigricaulis Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 225, 1824 [1825] (Reinwardt et al. 1824a). * Frullania nigricaulis var. elongata Verd., Ann. Bryol., Suppl. 1: 155, 1930 (Verdoorn 1930c). ** Frullania nivimontana S.Hatt., Bull. Natl. Sci. Mus. Tokyo, B 8 (3): 95, 1982 (Hattori 1982c). *** Frullania nobilis Steph., Hedwigia 33 (3): 154, 1894 (Stephani 1894d). ** Frullania nobilis var. cochleata (Steph.) S.Hatt., J. Hattori Bot. Lab. 59: 111, 1985 (Hattori and Streimann 1985). Bas.: Frullania cochleata Steph., Sp. Hepat. (Stephani) 4: 681, 1911 (Stephani 1911e). ** Frullania novocurvirostris S.Hatt., J. Hattori Bot. Lab. 49: 370, 1981 (Hattori 1981b). Nom. nov. pro Frullania curvirostris J.B.Jack et Steph., Hedwigia 33 (3): 143, 1894 (Stephani 1894d), nom. illeg. 200 Frullania ludoviciae seems to be close to Frullania pusilla (Hattori 1986e). 201 Frullania montana is possibly conspeciic with Frullania relexistipula. 202 Frullania nepalensis is a species complex (Hattori 1973b). 278 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Frullania oahuensis Hampe ex Gottsche, Lindenb. et Nees, Observ. bot.: 471, 1843 (Gottsche et al. 1843). ** Frullania obovata S.Hatt., Bull. Natl. Sci. Mus. Tokyo, B 8 (3): 97, 1982 (Hattori 1982c). ** Frullania obtusangula Hentschel et von Konrat, Phytotaxa 220 (2): 137, 2015 (Hentschel et al. 2015). ** Frullania okinawensis Kamim., Misc. Bryol. Lichenol. 9 (4): 90, 1982 (Kamimura 1982). ** Frullania orbicularis Austin, Proc. Acad. Nat. Sci. Philadelphia 21: 227, 1869 (Austin 1869). *** Frullania orientalis Sande Lac., Plagiochila Sandei: 10, 1856 (Sande Lacoste 1856c). ** Frullania orinocensis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 30, 1884 (Spruce 1884). *** Frullania ornithocephala (Reinw., Blume et Nees) Nees, Syn. Hepat. 3: 425, 1845 (Gottsche et al. 1845b). Bas.: Jungermannia ornithocephala Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 216, 1824 [1825] (Reinwardt et al. 1824a). * Frullania ornithocephala var. major (Nees) Schifn., Consp. Hepat. Arch. Ind.: 336, 1898 (Schifner 1898b). Bas.: Jungermannia ornithocephala β major Nees, Enum. Pl. Crypt. Javae: 47, 1830 (Nees 1830). ** Frullania ornithocephala var. pilosa Verd., Ann. Bryol. 2: 129, 1929 (Verdoorn 1929a). ** Frullania ornithocephala var. tuberculosa S.Hatt., J. Hattori Bot. Lab. 38: 179, 1974 (Hattori 1974a). ** Frullania pachyderma S.Hatt., J. Hattori Bot. Lab. 44: 525, 1978 (Hattori 1978b). ** Frullania pallidevirens Steph., Sp. Hepat. (Stephani) 4: 454, 1911 (Stephani 1911e). ** Frullania pariharii S.Hatt. et haithong, J. Jap. Bot. 53 (5): 130, 1978 (Hattori and haithong 1978a). * Frullania parvifolia Steph., Sp. Hepat. (Stephani) 4: 354, 1910 (Stephani 1910b). ** Frullania pauciramea Steph., Sp. Hepat. (Stephani) 4: 458, 1911 (Stephani 1911e). ** Frullania pauciramea var. pauciramella S.Hatt. et Piippo, Acta Bot. Fenn. 133: 50, 1986 (Hattori and Piippo 1986). ** Frullania paucirameoides S.Hatt. et Piippo, Acta Bot. Fenn. 133: 51, 1986 (Hattori and Piippo 1986). ** Frullania pedicellata Steph., Bull. Herb. Boissier 5 (2): 90, 1897 (Stephani 1897b). *** Frullania physantha Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 121, 1860 [1861] (Mitten 1860c). ** Frullania piptophylla S.Hatt., J. Hattori Bot. Lab. 47: 87, 1980 (Hattori 1980d). ** Frullania piptophylla var. minor S.Hatt., J. Hattori Bot. Lab. 60: 247, 1986 (Hattori 1986b). ** Frullania piptophylloides S.Hatt., J. Hattori Bot. Lab. 47: 90, 1980 (Hattori 1980d). * Frullania plicata Hentschel et von Konrat, Phytotaxa 220 (2): 137, 2015 (Hentschel et al. 2015). Nom. nov. pro Frullania acutiloba Gerola, Lav. Bot. Ist. Bot. Univ. Padova 12: 477, 1947 (Gerola 1947), nom. illeg. World checklist of hornworts and liverworts 279 ** Frullania pocsantha haithong et S.Hatt., Bull. Natl. Sci. Mus. Tokyo, B 3 (4): 149, 1977 (haithong and Hattori 1977). ** Frullania pran-nathii M.Dey et D.K.Singh, J. Jap. Bot. 83 (5): 281, 2008 (Dey and Singh 2008). ** Frullania pringlei Fulford et Sharp, Mem. New York Bot. Gard. 63: 45, 1990 (Fulford and Sharp 1990). Nom. nov. pro Frullania spicata Steph., Sp. Hepat. (Stephani) 4: 392, 1910 (Stephani 1910b), nom. illeg. ** Frullania prominula S.Hatt. et Streimann, J. Hattori Bot. Lab. 59: 112, 1985 (Hattori and Streimann 1985). ** Frullania propaginea S.Hatt. et Streimann, J. Hattori Bot. Lab. 59: 114, 1985 (Hattori and Streimann 1985). * Frullania pseudericoides S.Hatt., J. Hattori Bot. Lab. 51: 256, 1982 (Hattori 1982d). Nom. nov. pro Frullania sharpii subsp. subrostrata S.Hatt., J. Hattori Bot. Lab. 38: 265, 1974 (Hattori 1974c). ** Frullania pseudoschensiana S.Hatt., J. Hattori Bot. Lab. 47: 101, 1980 (Hattori 1980d). ** Frullania pseudoschensiana var. darjeelingensis S.Hatt., J. Hattori Bot. Lab. 49: 149, 1981 (Hattori 1981a). ** Frullania pullei Verd., Nova Guinea 14: 542, 1930 (Verdoorn 1930b). ** Frullania pusilla Mitt., Fl. vit.: 417, 1871 [1873] (Mitten 1871). *** Frullania pycnantha (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 3: 411, 1845 (Gottsche et al. 1845b). Bas.: Jungermannia pycnantha Hook.f. et Taylor, London J. Bot. 3: 566, 1844 (Hooker and Taylor 1844d). ** Frullania queenslandica Steph., Sp. Hepat. (Stephani) 4: 424, 1910 (Stephani 1910b). ** Frullania recurvistipula S.Hatt., Bull. Natl. Sci. Mus. Tokyo, B 1 (3): 114, 1975 (Hattori 1975b). *** Frullania relexistipula Sande Lac., Ned. Kruidk. Arch. 3: 422, 1854 [1855] (Sande Lacoste 1854). ** Frullania relexistipula var. squarrosa S.Hatt. et Piippo, Acta Bot. Fenn. 133: 54, 1986 (Hattori and Piippo 1986). ** Frullania remotidens S.Hatt., J. Hattori Bot. Lab. 51: 257, 1982 (Hattori 1982d). * Frullania reptans Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 161, 1855 (Mitten 1855). *** Frullania retusa Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 119, 1860 [1861] (Mitten 1860c). ** Frullania retusa var. gymnantha S.Hatt. et haithong, J. Jap. Bot. 53 (5): 131, 1978 (Hattori and haithong 1978a). ** Frullania retusa var. hirsuta S.Hatt. et haithong, J. Hattori Bot. Lab. 44: 191, 1978 (Hattori and haithong 1978b). *** Frullania rhystocolea Herzog, Symb. Sin. 5: 39, 1930 (Nicholson et al. 1930). ** Frullania rhytidantha S.Hatt., J. Hattori Bot. Lab. 47: 97, 1980 (Hattori 1980d). ** Frullania rigida Steph., Sp. Hepat. (Stephani) 4: 371, 1910 (Stephani 1910b). *** Frullania riparia Hampe, Nov. Stirp. Pug. 7: 14, 1838 (Lehmann 1838). 280 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Frullania rizalii Piippo et S.Hatt., J. Hattori Bot. Lab. 72: 119, 1992 (Piippo and Tan 1992). *** Frullania rostellata Mitt., Handb. N. Zeal. l. 2: 755, 1867 (Hooker 1867). ** Frullania rubella Gottsche, Hedwigia 28 (3): 159, 1889 (Stephani 1889d). ** Frullania rubella var. elongata (Steph.) S.Hatt., J. Hattori Bot. Lab. 54: 166, 1983 (Hattori 1983). Bas.: Frullania elongata Steph., Sp. Hepat. (Stephani) 4: 423, 1910 (Stephani 1910b). ** Frullania rupicola Steph., Sp. Hepat. (Stephani) 6: 534, 1924 (Stephani 1924). ** Frullania saipanensis S.Hatt. et Koike, J. Hattori Bot. Lab. 75: 188, 1994 (Koike 1994). *** Frullania schensiana C.Massal., Hepat. Shen-si: 40, 1897 (Massalongo 1897). ** Frullania schusteri S.Hatt., Beih. Nova Hedwigia 90: 154, 1988 (Hattori 1988b). ** Frullania scottiana S.Hatt., Mem. New York Bot. Gard. 45: 551, 1987 (Hattori 1987a). ** Frullania setchellii Pearson, Univ. Calif. Publ. Bot. 10 (4): 326, 1923 (Pearson 1923). ** Frullania shanensis Svihla, Bryologist 60 (4): 359, 1957 (Svihla 1957). ** Frullania sharpantha Udar et Ad.Kumar, Misc. Bryol. Lichenol. 9 (9): 192, 1983 (Udar and Kumar 1983a). ** Frullania sharpii S.Hatt., J. Hattori Bot. Lab. 38: 180, 1974 (Hattori 1974a). ** Frullania sinensis Steph., Nuovo Giorn. Bot. Ital. (n.ser.) 13 (4): 349, 1906 (Levier 1906). ** Frullania sinosphaerantha S.Hatt. et P.J.Lin, J. Hattori Bot. Lab. 59: 144, 1985 (Hattori and Lin 1985a). ** Frullania sphaerantha S.Hatt., J. Hattori Bot. Lab. 47: 99, 1980 (Hattori 1980d). ** Frullania sphaerolobulata S.H.Lin, Tunghai Journal 38: 104, 1997 (Lin and Chen 1997). *** Frullania spinifera Taylor, London J. Bot. 5: 407, 1846 (Taylor 1846b). ** Frullania spinigastria S.Hatt., J. Hattori Bot. Lab. 45: 358, 1979 (Hattori 1979b). ** Frullania spiniplica S.Hatt., J. Hattori Bot. Lab. 36: 428, 1972 [1973] (Hattori 1972b). *** Frullania spongiosa Steph., Hedwigia 33 (3): 147, 1894 (Stephani 1894d). ** Frullania squamuligera Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 33, 1884 (Spruce 1884). *** Frullania squarrosula (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 3: 412, 1845 (Gottsche et al. 1845b). Bas.: Jungermannia squarrosula Hook.f. et Taylor, London J. Bot. 4: 88, 1845 (Hooker and Taylor 1845). ** Frullania subcaduca S.Hatt., J. Hattori Bot. Lab. 38: 267, 1974 (Hattori 1974c). ** Frullania subclavata Steph., Sp. Hepat. (Stephani) 4: 354, 1910 (Stephani 1910b). ** Frullania subnigricaulis S.Hatt., J. Hattori Bot. Lab. 37: 89, 1973 (Hattori 1973c). ** Frullania subnigricaulis var. subtruncata S.Hatt., J. Hattori Bot. Lab. 39: 308, 1975 (Hattori 1975d). * Frullania subpedicellata S.Hatt., J. Hattori Bot. Lab. 47: 93, 1980 (Hattori 1980d). 203 ** Frullania subsquarrosa S.Hatt., J. Hattori Bot. Lab. 36: 429, 1972 [1973] (Hattori 1972b). 203 Frullania subpedicellata is possibly conspeciic with Frullania pedicellata. World checklist of hornworts and liverworts 281 ** Frullania subvalida S.Hatt. et haithong, J. Jap. Bot. 53 (6): 173, 1978 (Hattori and haithong 1978c). *** Frullania svihlana S.Hatt., J. Hattori Bot. Lab. 54: 180, 1983 (Hattori 1983). * Frullania taiheizana Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 241, 1934 (Horikawa 1934). ** Frullania tamsuina Steph., Sp. Hepat. (Stephani) 4: 444, 1910 (Stephani 1910b). *** Frullania taradakensis Steph., Sp. Hepat. (Stephani) 4: 352, 1910 (Stephani 1910b). ** Frullania tenuirostris Steph., Sp. Hepat. (Stephani) 4: 462, 1911 (Stephani 1911e). ** Frullania togashiana S.Hatt., Bull. Natl. Sci. Mus. Tokyo, B 1 (3): 118, 1975 (Hattori 1975b). ** Frullania tubercularis S.Hatt. et P.J.Lin, J. Jap. Bot. 60 (4): 107, 1985 (Hattori and Lin 1985b). *** Frullania usamiensis Steph., Bull. Herb. Boissier 5 (2): 91, 1897 (Stephani 1897b). ** Frullania valdiviensis J.B.Jack et Steph., Hedwigia 33 (3): 149, 1894 (Stephani 1894d). ** Frullania valida Steph., Sp. Hepat. (Stephani) 4: 402, 1910 (Stephani 1910b). ** Frullania variegata Steph., Hedwigia 33 (3): 149, 1894 (Stephani 1894d). * Frullania victoriensis Steph., Sp. Hepat. (Stephani) 4: 418, 1910 (Stephani 1910b). *** Frullania vittiana S.Hatt., Bryologist 90 (4): 368, 1987 [1988] (Hattori 1987b). ** Frullania wangii S.Hatt. et P.J.Lin, J. Hattori Bot. Lab. 59: 146, 1985 (Hattori and Lin 1985a). *** Frullania yuennanensis Steph., Hedwigia 33 (3): 161, 1894 (Stephani 1894d). ** Frullania yuennanensis var. siamensis (N.Kitag., haithong et S.Hatt.) S.Hatt. et P.J.Lin, J. Hattori Bot. Lab. 59: 148, 1985 (Hattori and Lin 1985a). Bas.: Frullania siamensis N.Kitag., haithong et S.Hatt., J. Hattori Bot. Lab. 43: 452, 1977 [1978] (Hattori et al. 1977). ** Frullania yuzawana S.Hatt., J. Hattori Bot. Lab. 49: 157, 1981 (Hattori 1981a). ** Frullania zangii S.Hatt. et P.J.Lin, J. Hattori Bot. Lab. 59: 149, 1985 (Hattori and Lin 1985a). ** Frullania zennoskeana S.Hatt., J. Jap. Bot. 59 (10): 308, 1984 (Hattori 1984b). ** sect. Acutilobae Verd., Ann. Bryol., Suppl. 1: 44, 1930 (Verdoorn 1930c). ** Frullania allanii E.A.Hodgs., Trans. & Proc. Roy. Soc. New Zealand 77 (3): 371, 1949 (Hodgson 1949). *** Frullania clavata (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 3: 428, 1845 (Gottsche et al. 1845b). Bas.: Jungermannia clavata Hook.f. et Taylor, London J. Bot. 4: 88, 1845 (Hooker and Taylor 1845). ** Frullania hamaticoma Steph., Hedwigia 28 (3): 158, 1889 (Stephani 1889d). *** Frullania monocera (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 3: 418, 1845 (Gottsche et al. 1845b). Bas.: Jungermannia monocera Hook.f. et Taylor, London J. Bot. 4: 89, 1845 (Hooker and Taylor 1845). ** Frullania monocera var. acutiloba (Mitt.) Hentschel et von Konrat, Phytotaxa 220 (2): 136, 2015 (Hentschel et al. 2015). Bas.: Frullania acutiloba Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 120, 1860 [1861] (Mitten 1860c). 282 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Frullania monocera var. depauperata S.Hatt., J. Hattori Bot. Lab. 57: 419, 1984 (Hattori 1984a). ** Frullania monocera var. schifneri (Verd.) S.Hatt., J. Hattori Bot. Lab. 46: 120, 1979 (Hattori 1979a). Bas.: Frullania acutiloba var. schifneri Verd., Ann. Bryol. 2: 123, 1929 (Verdoorn 1929a). *** Frullania monocera var. subhampeana (E.A.Hodgs.) Hentschel et von Konrat, Phytotaxa 220 (2): 136, 2015 (Hentschel et al. 2015). Bas.: Frullania subhampeana E.A.Hodgs., Trans. & Proc. Roy. Soc. New Zealand 77 (3): 370, 1949 (Hodgson 1949). *** Frullania monocera var. undulata (Kamim.) Hentschel et von Konrat, Phytotaxa 220 (2): 136, 2015 (Hentschel et al. 2015). Bas.: Frullania undulata Kamim., J. Hattori Bot. Lab. 24: 50, 1961 (Kamimura 1961). ** Frullania osumiensis (S.Hatt.) S.Hatt., J. Hattori Bot. Lab. 16: 87, 1956 (Iwatsuki and Hattori 1956). Bas.: Frullania hampeana var. osumiensis S.Hatt., Bull. Tokyo Sci. Mus. 11: 144, 1944 (Hattori 1944d). * Frullania pallidula S.Hatt., Beih. Nova Hedwigia 90: 152, 1988 (Hattori 1988b). ** Frullania pseudomonocera S.Hatt., J. Hattori Bot. Lab. 60: 216, 1986 (Hattori 1986e). ** Frullania seriata Gottsche, Hedwigia 28 (3): 160, 1889 (Stephani 1889d). ** Frullania spinistipula Steph., Sp. Hepat. (Stephani) 4: 463, 1911 (Stephani 1911e). ** Frullania streimannii S.Hatt., J. Hattori Bot. Lab. 54: 176, 1983 (Hattori 1983). *** sect. Australes Verd., Ann. Bryol., Suppl. 1: 58, 1930 (Verdoorn 1930c). *** Frullania anomala E.A.Hodgs., Trans. & Proc. Roy. Soc. New Zealand 77 (3): 374, 1949 (Hodgson 1949). * Frullania baileyana Steph., Sp. Hepat. (Stephani) 4: 417, 1910 (Stephani 1910b). *** Frullania baladina Gottsche, Hedwigia 33 (3): 140, 1894 (Stephani 1894d). 204 * Frullania belmorensis Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 106, 1914 (Stephani and Watts 1914). *** Frullania campanulata Sande Lac., Ned. Kruidk. Arch. 3: 422, 1854 [1855] (Sande Lacoste 1854). ** Frullania campanulata var. caduca Verd., Ann. Bryol., Suppl. 1: 41, 1930 (Verdoorn 1930c). ** Frullania campanulata var. malesiaca (Verd.) S.Hatt., Bull. Natl. Sci. Mus. Tokyo, B 1 (4): 158, 1975 (Hattori 1975f). Bas.: Frullania malesiaca Verd., Ann. Bryol., Suppl. 1: 59, 1930 (Verdoorn 1930c). * Frullania cataractarum Steph., Sp. Hepat. (Stephani) 4: 657, 1911 (Stephani 1911e). * Frullania crawfordii Steph., Hedwigia 33 (3): 143, 1894 (Stephani 1894d). *** Frullania dentata S.Hatt., J. Hattori Bot. Lab. 38: 231, 1974 (Hattori 1974c). ** Frullania dentata var. secernens S.Hatt., J. Hattori Bot. Lab. 65: 422, 1988 (Hattori 1988a). *** Frullania errans Verd., Ann. Bryol., Suppl. 1: 59, 1930 (Verdoorn 1930c). 204 Frullania baladina is a species complex also including Frullania fulfordiae. World checklist of hornworts and liverworts 283 ** Frullania errans var. angulistipula S.Hatt., J. Hattori Bot. Lab. 36: 431, 1972 [1973] (Hattori 1972b). *** Frullania fugax (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 3: 445, 1845 (Gottsche et al. 1845b). Bas.: Jungermannia fugax Hook.f. et Taylor, London J. Bot. 4: 87, 1845 (Hooker and Taylor 1845). 205 ** Frullania fulfordiae S.Hatt., Bryologist 90 (4): 365, 1987 [1988] (Hattori 1987b). *** Frullania glomerata (Lehm. et Lindenb.) Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 9: 46, 1838 (Montagne 1838). Bas.: Jungermannia glomerata Lehm. et Lindenb., Nov. Stirp. Pug. 5: 21, 1833 (Lehmann 1833). *** Frullania incumbens Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 162, 1855 (Mitten 1855). *** Frullania inlexa Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 120, 1860 [1861] (Mitten 1860c). ** Frullania media (E.A.Hodgs.) S.Hatt., J. Hattori Bot. Lab. 54: 153, 1983 (Hattori 1983). Bas.: Frullania fugax var. media E.A.Hodgs., Trans. & Proc. Roy. Soc. New Zealand 77 (3): 375, 1949 (Hodgson 1949). ** Frullania mizutanii Kamim. et S.Hatt., J. Hattori Bot. Lab. 37: 524, 1973 (Hattori and Kamimura 1973). *** Frullania obscurifolia Mitt., Philos. Trans. 168: 400, 1879 (Mitten 1879). *** Frullania patagonica Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 88, 1911 (Stephani 1911b). ** Frullania pentapleura Taylor, London J. Bot. 5: 402, 1846 (Taylor 1846b). ** Frullania polyptera Taylor, London J. Bot. 5: 401, 1846 (Taylor 1846b). ** Frullania polyptera var. angustata (Mitt.) S.Hatt., Bull. Natl. Sci. Mus. Tokyo, B 1 (2): 74, 1975 (Hattori 1975g). Bas.: Frullania angustata Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 122, 1860 [1861] (Mitten 1860c). ** Frullania probosciphora Taylor, London J. Bot. 5: 402, 1846 (Taylor 1846b). ** Frullania pulchella Herzog, Rev. Bryol. Lichénol. 23 (1/2): 60, 1954 (Herzog 1954). *** Frullania sinskeana J.J.Engel et B.C.Tan, J. Hattori Bot. Lab. 60: 335, 1986 (Tan and Engel 1986). Nom. nov. pro Frullania spathulistipa Steph., Sp. Hepat. (Stephani) 4: 415, 1910 (Stephani 1910b), nom. illeg. ** Frullania socotrana Mitt., Trans. Roy Soc. Edinburgh 31: 335, 1888 (Mitten 1888). * Frullania solanderiana Colenso, Trans. & Proc. New Zealand Inst. 21: 75, 1889 (Colenso 1889). 206 ** Frullania subincumbens S.Hatt., Bryologist 90 (4): 367, 1987 [1988] (Hattori 1987b). * Frullania subtropica Steph., Sp. Hepat. (Stephani) 4: 416, 1910 (Stephani 1910b). * Frullania tjibodensis S.Hatt. et haithong, J. Jap. Bot. 52 (10): 289, 1977 (Hattori and haithong 1977). 207 205 Frullania fugax is a species complex possibly including Frullania baileyana, Frullania belmorensis, Frullania cataractarum, Frullania media and Frullania subtropica (Hattori 1979a). 206 Frullania solanderiana is morphologically similar to Frullania pentapleura and possibly conspeciic. 207 Frullania tjibodensis is possibly conspeciic with Frullania campanulata (Söderström et al. 2010a). 284 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Frullania tuyamae S.Hatt. et haithong, J. Jap. Bot. 53 (6): 175, 1978 (Hattori and haithong 1978c). *** sect. Frullania *** Frullania appalachiana R.M.Schust., Phytologia 53 (5): 366, 1983 (Schuster 1983b). *** Frullania azorica Sim-Sim, Sérgio, Mues et Kraut, Cryptog. Bryol. Lichénol. 16 (2): 112, 1995 (Sim-Sim et al. 1995). *** Frullania catalinae A.Evans, Trans. Connecticut Acad. Arts 10 (1): 11, 1899 (Evans 1899). *** Frullania dilatata (L.) Dumort., Recueil Observ. Jungerm.: 13, 1835 (Dumortier 1835). Bas.: Jungermannia dilatata L., Sp. Pl. 1: 1133, 1753 (Linnaeus 1753). ** Frullania dilatata subsp. asiatica S.Hatt., J. Jap. Bot. 57 (9): 258, 1982 (Hattori 1982b). *** Frullania eboracensis Lehm., Nov. Stirp. Pug. 8: 14, 1844 (Lehmann 1844). 208 *** Frullania ericoides (Nees) Mont., Ann. Sci. Nat. Bot. (sér. 2) 12: 51, 1839 (Montagne 1839c). Bas.: Jungermannia ericoides Nees, Fl. Bras. (Martius) 1 (1): 346, 1833 (Nees 1833a). 209 * Frullania ericoides var. laxa (Gottsche, Lindenb. et Nees) Schifn., Consp. Hepat. Arch. Ind.: 324, 1898 (Schifner 1898b). Bas.: Frullania squarrosa γ laxa Gottsche, Lindenb. et Nees, Syn. Hepat. 5: 772, 1847 (Gottsche et al. 1847). * Frullania ericoides var. minor Kamim., Bull. Kochi Gakuen Jun. Coll. 2: 22, 1971 (Kamimura 1971). ** Frullania ericoides var. verrucosa (Kamim.) Hentschel et von Konrat, Phytotaxa 220 (2): 134, 2015 (Hentschel et al. 2015). Bas.: Frullania squarrosa var. verrucosa Kamim., J. Hattori Bot. Lab. 24: 19, 1961 (Kamimura 1961). *** Frullania fragilifolia (Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 3: 437, 1845 (Gottsche et al. 1845b). Bas.: Jungermannia fragilifolia Taylor, Ann. Mag. Nat. Hist. 12 (76): 172, 1843 (Taylor 1843). ** Frullania fuegiana Steph., Sp. Hepat. (Stephani) 4: 428, 1910 (Stephani 1910b). ** Frullania hattoriana J.D.Godfrey et G.Godfrey, J. Hattori Bot. Lab. 48: 321, 1980 (Godfrey and Godfrey 1980). ** Frullania koponenii S.Hatt., Ann. Bot. Fenn. 15 (2): 111, 1978 (Koponen et al. 1978). *** Frullania muscicola Steph., Hedwigia 33 (3): 146, 1894 (Stephani 1894d). *** Frullania oakesiana Austin, Proc. Acad. Nat. Sci. Philadelphia 21: 225, 1869 (Austin 1869). ** Frullania oakesiana subsp. takayuensis (Steph.) R.M.Schust., Hepat. Anthocerotae N. Amer. 5: 195, 1992 (Schuster 1992b). Bas.: Frullania takayuensis Steph., Sp. Hepat. (Stephani) 4: 399, 1910 (Stephani 1910b). 208 Frullania eboracensis is a species complex also including Frullania appalachiana, Frullania parvistipula and Frullania virginica. 209 Frullania ericoides is enormously variable (Schuster 1992b) and is a species complex with at least two lineages (Hentschel et al. 2009). World checklist of hornworts and liverworts 285 *** Frullania parvistipula Steph., Sp. Hepat. (Stephani) 4: 397, 1910 (Stephani 1910b). ** Frullania sabaliana R.M.Schust., Phytologia 53 (5): 365, 1983 (Schuster 1983b). ** Frullania semivillosa Lindenb. et Gottsche, Syn. Hepat. 5: 774, 1847 (Gottsche et al. 1847). ** Frullania stylifera (R.M.Schust.) R.M.Schust., Hepat. Anthocerotae N. Amer. 5: 210, 1992 (Schuster 1992b). Bas.: Frullania inlata var. stylifera R.M.Schust., Phytologia 53 (5): 366, 1983 (Schuster 1983b). * Frullania subdilatata C.Massal., Nuovo Giorn. Bot. Ital. (n.ser.) 13 (4): 349, 1906 (Levier 1906). *** Frullania virginica Lehm., Nov. Stirp. Pug. 8: 19, 1844 (Lehmann 1844). ** sect. Irregulares E.A.Hodgs. ex S.Hatt., J. Hattori Bot. Lab. 54: 143, 1983 (Hattori 1983). * Frullania astrolabea Steph., Sp. Hepat. (Stephani) 4: 460, 1910 (Stephani 1910b). 210 *** Frullania deplanata Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 161, 1855 (Mitten 1855). *** Frullania morobensis S.Hatt. et Streimann, J. Hattori Bot. Lab. 59: 109, 1985 (Hattori and Streimann 1985). *** Frullania patula Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 159, 1854 (Mitten 1854). *** Frullania scandens Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 258, 1843 (Montagne 1843). ** sect. Planae R.M.Schust., Phytologia 57 (5): 372, 1985 (Schuster 1985a). *** Frullania plana Sull., Mem. Amer. Acad. Arts (n.ser.) 4: 175, 1849 (Sullivant 1849). *** subg. Homotropantha Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 35, 1884 (Spruce 1884). *** Frullania delexa Mitt., Bonplandia 10 (2): 19, 1862 (Mitten 1862). *** Frullania integristipula (Nees) Nees, Syn. Hepat. 3: 431, 1845 (Gottsche et al. 1845b). Bas.: Jungermannia integristipula Nees, Enum. Pl. Crypt. Javae: 54, 1830 (Nees 1830). ** Frullania integristipula var. emarginata Verd., Ann. Bryol. 2: 153, 1929 (Verdoorn 1929a). ** Frullania macrophylla S.Hatt., J. Hattori Bot. Lab. 47: 220, 1980 (Hattori 1980a). ** Frullania sabahana S.Hatt., J. Hattori Bot. Lab. 40: 493, 1976 (Hattori 1976d). ** Frullania sackawana Steph., Bull. Herb. Boissier 5 (2): 91, 1897 (Stephani 1897b). ** Frullania sarawakensis S.Hatt., J. Hattori Bot. Lab. 40: 496, 1976 (Hattori 1976d). ** Frullania umbonata Mitt., Sp. Hepat. (Stephani) 4: 579, 1911 (Stephani 1911e). *** Frullania utriculata Steph., Hedwigia 33 (3): 152, 1894 (Stephani 1894d). *** sect. Fallaces Verd., Rev. Bryol. Lichénol. 1: 112, 1928 (Verdoorn 1928a). * Frullania fallax Gottsche, Syn. Hepat. 3: 432, 1845 (Gottsche et al. 1845b). 210 Frullania astrolabea is possibly conspeciic with Frullania scandens (Hattori 1981b). 286 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Frullania intermedia (Reinw., Blume et Nees) Nees, Syn. Hepat. 3: 434, 1845 (Gottsche et al. 1845b). Bas.: Jungermannia intermedia Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 218, 1824 [1825] (Reinwardt et al. 1824a). 211 ** Frullania intermedia subsp. morokensis (Steph.) S.Hatt., J. Hattori Bot. Lab. 47: 194, 1980 (Hattori 1980a). Bas.: Frullania morokensis Steph., Sp. Hepat. (Stephani) 4: 578, 1911 (Stephani 1911e). * Frullania intermedia var. non-apiculata S.Hatt., J. Hattori Bot. Lab. 39: 291, 1975 (Hattori 1975d). 212 ** Frullania novoguineensis Schifn., Leberm., Forschungsr. Gazelle 4 (4): 37, 1890 (Schifner 1890). ** Frullania regularis Schifn., Leberm., Forschungsr. Gazelle 4 (4): 38, 1890 (Schifner 1890). *** * ** ** *** sect. Nodulosae Verd., Rev. Bryol. Lichénol. 1: 116, 1928 (Verdoorn 1928a). Frullania brotheri Steph., Hedwigia 33 (3): 150, 1894 (Stephani 1894d). 213 Frullania hamata Steph., Sp. Hepat. (Stephani) 4: 582, 1911 (Stephani 1911e). Frullania leeuwenii Verd., Nova Guinea 14: 545, 1930 (Verdoorn 1930b). Frullania nodulosa (Reinw., Blume et Nees) Nees, Syn. Hepat. 3: 433, 1845 (Gottsche et al. 1845b). Bas.: Jungermannia nodulosa Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 217, 1824 [1825] (Reinwardt et al. 1824a). ** sect. Remotilobae Verd., Rev. Bryol. Lichénol. 1: 119, 1928 (Verdoorn 1928a). *** Frullania heteromorpha Schifn., Leberm., Forschungsr. Gazelle 4 (4): 38, 1890 (Schifner 1890). ** Frullania remotiloba Steph., Hedwigia 33 (3): 152, 1894 (Stephani 1894d). * subg. Mammillosae S.Hatt., J. Hattori Bot. Lab. 60: 226, 1986 (Hattori 1986e). ** Frullania huerlimannii S.Hatt., Bull. Natl. Sci. Mus. Tokyo, B 2 (3): 84, 1976 (Hattori 1976b). * Frullania huerlimannii var. dioica S.Hatt., J. Hattori Bot. Lab. 57: 413, 1984 (Hattori 1984a). ** Frullania involvens S.Hatt. et Kamim., J. Hattori Bot. Lab. 37: 526, 1973 (Hattori and Kamimura 1973). ** Frullania iriomotensis S.Hatt., J. Jap. Bot. 55 (5): 133, 1980 (Hattori 1980c). ** Frullania mammillosa S.Hatt., J. Hattori Bot. Lab. 43: 424, 1977 [1978] (Hattori 1977b). 211 Frullania intermedia is a species complex (Hattori 1980a). 212 Frullania intermedia var. non-apiculata was reduced to a synonym of subsp. intermedia by Hattori (1980a), but it was re-instated by Hattori (1985). 213 Frullania brotheri is probably just a high-elevation form of Frullania nodulosa (Söderström et al. 2010a). World checklist of hornworts and liverworts 287 *** Frullania meijeri S.Hatt., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 17 (4): 307, 1974 (Hattori 1974b). ** Frullania notarisii Steph., Sp. Hepat. (Stephani) 4: 651, 1911 (Stephani 1911e). ** Frullania papulosa Steph., Sp. Hepat. (Stephani) 4: 654, 1911 (Stephani 1911e). ** Frullania rudoliana S.Hatt., J. Hattori Bot. Lab. 36: 437, 1972 [1973] (Hattori 1972b). *** Frullania thiersiae S.Hatt., Beih. Nova Hedwigia 90: 156, 1988 (Hattori 1988b). ** Frullania tixieri S.Hatt., J. Jap. Bot. 51 (7): 193, 1976 (Hattori 1976a). *** subg. Meteoriopsis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 37, 1884 (Spruce 1884). * Frullania caldensis Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 33 (7): 89, 1876 [1877] (Ångström 1876). ** Frullania evoluta Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 122, 1860 [1861] (Mitten 1860c). *** Frullania tagawana (S.Hatt. et haithong) S.Hatt., J. Hattori Bot. Lab. 59: 160, 1985 (Hattori and Lin 1985a). Bas.: Frullania evoluta var. tagawana S.Hatt. et haithong, J. Hattori Bot. Lab. 43: 441, 1977 [1978] (Hattori et al. 1977). *** *** ** *** *** *** *** ** * * * sect. Intumescentes R.M.Schust., Phytologia 57 (5): 370, 1985 (Schuster 1985a). Frullania aculeata Taylor, London J. Bot. 5: 407, 1846 (Taylor 1846b). Frullania ambronnii Steph., Biblioth. Bot. 87 (2): 242, 1916 (Stephani 1916a). Frullania atrata (Sw.) Nees ex Mont., Ann. Sci. Nat. Bot. (sér. 2) 12: 51, 1839 (Montagne 1839c). Bas.: Jungermannia atrata Sw., Prodr. (Swartz): 144, 1788 (Swartz 1788). Frullania beyrichiana (Lehm. et Lindenb.) Lehm. et Lindenb., Syn. Hepat. 3: 460, 1845 (Gottsche et al. 1845b). Bas.: Jungermannia beyrichiana Lehm. et Lindenb., Nov. Stirp. Pug. 5: 25, 1833 (Lehmann 1833). Frullania bicornistipula Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 46, 1884 (Spruce 1884). Frullania brasiliensis Raddi, Critt. Brasil.: 12, 1822 (Raddi 1822). 214 Frullania brasiliensis var. elegantula Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 50, 1884 (Spruce 1884). Frullania breuteliana Gottsche, Syn. Hepat. 3: 461, 1845 (Gottsche et al. 1845b). 215 Frullania compacta Gottsche, Sp. Hepat. (Stephani) 4: 493, 1911 (Stephani 1911e). 216 Frullania crenulifolia J.B.Jack et Steph., Hedwigia 31 (1): 14, 1892 (Jack and Stephani 1892). 214 Frullania brasiliensis is a species complex with controversial synonymy (Schuster 1992b, Hentschel et al. 2009). 215 Frullania breuteliana is possibly conspeciic with Frullania beyrichiana. 216 Frullania compacta is conspeciic with Frullania brasiliensis in Stotler (1969), but Schuster (1992b) doubts the correctness of this. 288 ** ** *** ** * *** * * ** * *** * *** * *** ** ** * *** ** ** *** ** *** ** *** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Frullania crispiloba Steph., Hedwigia 33 (3): 156, 1894 (Stephani 1894d). Frullania curviramea Steph., Sp. Hepat. (Stephani) 4: 684, 1911 (Stephani 1911e). Frullania ecuadorensis Steph., Sp. Hepat. (Stephani) 4: 526, 1911 (Stephani 1911e). Frullania formosa Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 46, 1884 (Spruce 1884). Frullania granatensis Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 173 (79), 1864 (Gottsche 1864). 217 Frullania griithsiana Gottsche, Syn. Hepat. 4: 466, 1846 (Gottsche et al. 1846). Frullania guadalupensis Gottsche, Sp. Hepat. (Stephani) 4: 496, 1911 (Stephani 1911e). Frullania gualaquizana Steph., Sp. Hepat. (Stephani) 4: 531, 1911 (Stephani 1911e). Frullania hamilora Herzog et L.Clark, Bryologist 56 (3): 180, 1953 (Clark and Schultz 1953). Frullania humilis Spruce, Mem. Torrey Bot. Club 1 (3): 119, 1890 (Spruce 1890). Frullania intumescens (Lehm. et Lindenb.) Lehm. et Lindenb., Syn. Hepat. 3: 460, 1845 (Gottsche et al. 1845b). Bas.: Jungermannia intumescens Lehm. et Lindenb., Nov. Stirp. Pug. 6: 52, 1834 (Lehmann 1834). Frullania laticaulis Spruce, Mem. Torrey Bot. Club 1 (3): 120, 1890 (Spruce 1890). Frullania lobatohastata Steph., Sp. Hepat. (Stephani) 4: 499, 1911 (Stephani 1911e). Frullania longistipula var. apiculata Demaret et Vanden Berghen, Bull. Jard. Bot. État Bruxelles 20 (1): 4, 1950 (Demaret and Vanden Berghen 1950). Frullania macrocephala (Lehm. et Lindenb.) Lehm. et Lindenb., Syn. Hepat. 3: 460, 1845 (Gottsche et al. 1845b). Bas.: Jungermannia macrocephala Lehm. et Lindenb., Nov. Stirp. Pug. 5: 20, 1833 (Lehmann 1833). Frullania madothecoides Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 47, 1884 (Spruce 1884). Frullania meridana Steph., Sp. Hepat. (Stephani) 4: 500, 1911 (Stephani 1911e). Frullania microcephala Gottsche, Mexik. Leverm.: 251, 1863 (Gottsche 1863). Frullania montagnei Gottsche, Syn. Hepat. 3: 456, 1845 (Gottsche et al. 1845b). Frullania moritziana Lindenb. et Gottsche, Syn. Hepat. 5: 782, 1847 (Gottsche et al. 1847). Frullania osculatiana De Not., Mem. Reale Accad. Sci. Torino (ser. 2) 16: 236, 1857 (De Notaris 1857). Frullania paradoxa Lehm. et Lindenb., Syn. Hepat. 3: 462, 1845 (Gottsche et al. 1845b). Frullania pearceana Steph., Sp. Hepat. (Stephani) 4: 515, 1911 (Stephani 1911e). Frullania pittieri Steph., Primit. l. costar.: 113, 1892 [1893] (Stephani 1892e). Frullania rigescens Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 52, 1884 (Spruce 1884). Frullania setigera Steph., Hedwigia 33 (3): 159, 1894 (Stephani 1894d). 217 Frullania granatensis is possibly conspeciic with Frullania beyrichiana, but the type specimen was lost in B (Stotler 1969). World checklist of hornworts and liverworts 289 ** Frullania speciosa Herzog, Memoranda Soc. Fauna Fl. Fennica 25: 59, 1950 (Herzog 1950c). ** Frullania supradecomposita (Lehm. et Lindenb.) Lehm. et Lindenb., Syn. Hepat. 3: 431, 1845 (Gottsche et al. 1845b). Bas.: Jungermannia supradecomposita Lehm. et Lindenb., Nov. Stirp. Pug. 5: 23, 1833 (Lehmann 1833). * Frullania trianae Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 173, 1864 (Gottsche 1864). ** Frullania triquetra Lindenb. et Gottsche, Syn. Hepat. 5: 780, 1847 (Gottsche et al. 1847). * Frullania trollii Herzog, Beih. Bot. Centralbl. 61B (3): 575, 1942 (Herzog 1942d). *** Frullania uleana Steph., Hedwigia 33 (3): 155, 1894 (Stephani 1894d). ** sect. Meteoriopsis Uribe, von Konrat et Hentschel, Phytotaxa 220 (2): 132, 2015 (Hentschel et al. 2015). *** Frullania convoluta Lindenb. et Hampe, Linnaea 24 (3): 303, 1851 [1852] (Hampe 1851b). * Frullania convoluta var. ampliata Herzog, Rev. Bryol. Lichénol. 20 (1/2): 129, 1951 [1952] (Herzog 1951a). *** Frullania darwinii Gradst. et Uribe, Cryptog. Bryol. 25 (4): 296, 2004 (Uribe 2004). *** Frullania dulimensis Uribe, Cryptog. Bryol. 27 (3): 309, 2006 (Uribe 2006). *** Frullania grandifolia Steph., Sp. Hepat. (Stephani) 4: 684, 1911 (Stephani 1911e). *** Frullania peruviana Gottsche, Syn. Hepat. 4: 465, 1846 (Gottsche et al. 1846). *** Frullania phalangilora Steph., Biblioth. Bot. 87 (2): 247, 1916 (Stephani 1916a). *** Frullania weberbaueri Steph., Sp. Hepat. (Stephani) 4: 510, 1911 (Stephani 1911e). *** sect. Obtusilobae Verd., Ann. Bryol., Suppl. 1: 81, 1930 (Verdoorn 1930c). ** Frullania angulata Mitt., J. Proc. Linn. Soc., Bot. 7 (27): 169, 1863 (Mitten 1863). ** Frullania angulata var. laciniata Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 46 (1/2): 72, 1976 (Vanden Berghen 1976b). ** Frullania apicalis Mitt., Philos. Trans. 168: 401, 1879 (Mitten 1879). ** Frullania apicalis var. camerunensis Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 46 (1/2): 53, 1976 (Vanden Berghen 1976b). * Frullania borbonica Lindenb., Syn. Hepat. 3: 455, 1845 (Gottsche et al. 1845b). 218 *** Frullania capensis Gottsche, Syn. Hepat. 3: 449, 1845 (Gottsche et al. 1845b). ** Frullania donnellii Austin, Bull. Torrey Bot. Club 6 (52): 301, 1879 (Austin 1879). ** Frullania eplicata Steph., Sp. Hepat. (Stephani) 4: 679, 1911 (Stephani 1911e). ** Frullania imerinensis Steph., Sp. Hepat. (Stephani) 4: 484, 1911 (Stephani 1911e). *** Frullania kunzei (Lehm. et Lindenb.) Lehm. et Lindenb., Syn. Hepat. 3: 449, 1845 (Gottsche et al. 1845b). Bas.: Jungermannia kunzei Lehm. et Lindenb., Nov. Stirp. Pug. 6: 50, 1834 (Lehmann 1834). 218 Frullania borbonica is possibly conspeciic with Frullania apicalis (Vanden Berghen 1976b). 290 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Frullania kunzei var. maritima R.M.Schust., Phytotaxa 220 (2): 135, 2015 (Hentschel et al. 2015). Based on: Frullania kunzei var. maritima R.M.Schust., J. Hattori Bot. Lab. 70: 145, 1991 (Schuster 1991b), nom. inval. ** Frullania longistipula Steph., Bull. Soc. Roy. Bot. Belgique, Compt. Rend. 30 (2): 199, 1891 [1892] (Stephani 1891b). *** Frullania meyeniana Lindenb., Syn. Hepat. 3: 455, 1845 (Gottsche et al. 1845b). ** Frullania meyeniana var. dioica S.Hatt., J. Hattori Bot. Lab. 43: 426, 1977 [1978] (Hattori 1977b). ** Frullania onraedtii Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 46 (1/2): 60, 1976 (Vanden Berghen 1976b). ** Frullania papuana Verd., Ann. Bryol., Suppl. 1: 82, 1930 (Verdoorn 1930c). *** Frullania schimperi Nees, Syn. Hepat. 3: 454, 1845 (Gottsche et al. 1845b). 219 ** Frullania schimperi var. laciniata Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 46 (1/2): 59, 1976 (Vanden Berghen 1976b). *** subg. Microfrullania (R.M.Schust.) R.M.Schust., Hepat. Anthocerotae N. Amer. 5: 34, 1992 (Schuster 1992b). Bas.: Neohattoria subg. Microfrullania R.M.Schust., J. Hattori Bot. Lab. 33: 280, 1970 (Schuster 1970a). *** Frullania fertilis De Not., Mem. Reale Accad. Sci. Torino (ser. 2) 16: 235, 1857 (De Notaris 1857). *** Frullania knightbridgei von Konrat et de Lange, PhytoKeys 8: 28, 2012 (von Konrat et al. 2012b). *** Frullania magellanica F.Weber et Nees, Syn. Hepat. 3: 446, 1845 (Gottsche et al. 1845b). Nom. nov. pro Jungermannia magellanica Spreng. Ann. Wetterauischen Ges. Gesammte Naturk. 1: 25, 1809 (Sprengel 1809), nom. illeg. ** Frullania magellanica subsp. tristaniana (S.W.Arnell) Váňa et J.J.Engel, Mem. New York Bot. Gard. 105: 59, 2013 (Váňa and Engel 2013). Bas.: Frullania tristaniana S.W.Arnell, Results Norweg. Sci. Exped. Tristan da Cunha 42: 9, 1958 (Arnell 1958b). * Frullania matafaoica H.A.Mill., Phytologia 47 (4): 322, 1981 (Miller 1981). Nom. nov. pro Frullania minutissima Pearson, Amer. Samoa: 140, 1924 (Pearson 1924a), nom. illeg. *** Frullania toropuku von Konrat, de Lange et Larraín, Polish Bot. J. 58 (2): 439, 2013 (von Konrat et al. 2013). *** sect. Amphijubula (R.M.Schust.) von Konrat, Hentschel, Heinrichs et Braggins, Bryologist 114 (1): 53, 2011 (von Konrat et al. 2011). Bas.: Amphijubula R.M.Schust., J. Hattori Bot. Lab. 33: 298, 1970 (Schuster 1970a). *** Frullania lobulata (Hook.) Hook. et Nees, Syn. Hepat. 3: 445, 1845 (Gottsche et al. 1845b). Bas.: Jungermannia lobulata Hook., Musci Exot. 2: tab. 119, 1820 (Hooker 1820). 219 Frullania schimperi belongs to a species complex also including Frullania onraedtii, Frullania capensis, Frullania apicalis, Frullania imerinensis and Frullania meyeniana (Vanden Berghen 1976b). World checklist of hornworts and liverworts 291 *** Frullania microcaulis Gola, Nuovo Giorn. Bot. Ital. (n.ser.) 29 (1/4): 172, 1922 [1923] (Gola 1922). *** Frullania truncatistyla von Konrat, Hentschel, Heinrichs et Braggins, Bryologist 114 (1): 63, 2011 (von Konrat et al. 2011). *** sect. Microfrullania (R.M.Schust.) von Konrat et Hentschel, Phytotaxa 220 (2): 133, 2015 (Hentschel et al. 2015). Bas.: Neohattoria sect. Microfrullania R.M.Schust., J. Hattori Bot. Lab. 33: 288, 1970 (Schuster 1970a). *** Frullania chevalieri (R.M.Schust.) R.M.Schust., Hepat. Anthocerotae N. Amer. 5: 34, 1992 (Schuster 1992b). Bas.: Neohattoria chevalieri R.M.Schust., J. Hattori Bot. Lab. 33: 289, 1970 (Schuster 1970a). 220 *** Frullania microscopica Pearson, J. Linn. Soc., Bot. 46 (305): 33, 1922 (Pearson 1922b). ** Frullania neocaledonica J.J.Engel, Novon 9 (3): 344, 1999 (Engel and Smith Merrill 1999a). Nom. nov. pro Neohattoria caledonica R.M.Schust., J. Hattori Bot. Lab. 33: 291, 1970 (Schuster 1970a). *** Frullania parhamii (R.M.Schust.) R.M.Schust. ex von Konrat, L.Söderstr. et A.Hagborg, Telopea 13 (3): 407, 2011 (Söderström et al. 2011a). Bas.: Neohattoria parhamii R.M.Schust., J. Hattori Bot. Lab. 26: 243, 1963 (Schuster 1963b). *** sect. Regulares Verd., Ann. Bryol., Suppl. 1: 133, 1930 (Verdoorn 1930c). *** Frullania junghuhniana Gottsche, Syn. Hepat. 3: 444, 1845 (Gottsche et al. 1845b). 221 ** Frullania junghuhniana var. bisexualis S.Hatt., J. Hattori Bot. Lab. 40: 485, 1976 (Hattori 1976d). ** Frullania junghuhniana var. tenella (Sande Lac.) Grolle et S.Hatt., Misc. Bryol. Lichenol. 9 (6): 123, 1982 (Hattori 1982a). Bas.: Frullania tenella Sande Lac., Ned. Kruidk. Arch. 3: 423, 1854 [1855] (Sande Lacoste 1854). ** Frullania mcveanii S.Hatt., J. Hattori Bot. Lab. 37: 55, 1973 (Hattori 1973a). ** Frullania pseudomeyeniana S.Hatt., J. Hattori Bot. Lab. 60: 231, 1986 (Hattori 1986e). *** Frullania rostrata (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 3: 445, 1845 (Gottsche et al. 1845b). Bas.: Jungermannia rostrata Hook.f. et Taylor, London J. Bot. 4: 87, 1845 (Hooker and Taylor 1845). 222 ** Frullania scalaris S.Hatt., J. Hattori Bot. Lab. 43: 432, 1977 [1978] (Hattori 1977b). ** subg. Saccophora Verd., Ann. Bryol. 2: 121, 1929 (Verdoorn 1929a). *** Frullania gaudichaudii (Nees et Mont.) Nees et Mont., Syn. Hepat. 3: 435, 1845 (Gottsche et al. 1845b). Bas.: Jubula gaudichaudii Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 5: 64, 1836 (Nees and Montagne 1836). 220 Frullania chevalieri is a species complex (Hattori 1984a). 221 Frullania junghuhniana is a species complex (Hattori 1976b). 222 Frullania rostrata is a species complex with several lineages, some of which have been recently described (e.g. von Konrat et al. 2013). 292 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Frullania gaudichaudii var. ceylanica (Nees) S.Hatt., J. Hattori Bot. Lab. 47: 104, 1980 (Hattori 1980d). Bas.: Frullania ceylanica Nees, Syn. Hepat. 3: 436, 1845 (Gottsche et al. 1845b). *** Frullania hedrantha S.Hatt. et Kamim., J. Hattori Bot. Lab. 37: 519, 1973 (Hattori and Kamimura 1973). ** Frullania immersa Steph., Bot. Jahrb. Syst. 23 (1/2, 3): 315, 1896 (Stephani 1896a). ** Frullania pancheri Gottsche, Hedwigia 33 (3): 159, 1894 (Stephani 1894d). ** Frullania papillilobula S.Hatt., Bull. Natl. Sci. Mus. Tokyo, B 1 (4): 141, 1975 (Hattori 1975f). ** Frullania sublignosa Steph., Hedwigia 33 (3): 148, 1894 (Stephani 1894d). subg. Steerea (S.Hatt. et Kamim.) R.M.Schust., Hepat. Anthocerotae N. Amer. 5: 32, 1992 (Schuster 1992b). Bas.: Steerea S.Hatt. et Kamim., J. Hattori Bot. Lab. 34: 429, 1971 (Hattori and Kamimura 1971). *** Frullania clemensiana Verd., Ned. Kruidk. Arch. (ser. 3) 42 (2): 493, 1932 (Verdoorn 1932b). * *** subg. hyopsiella Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 41, 1884 (Spruce 1884). *** Frullania acicularis Hentschel et von Konrat, Phytotaxa 220 (2): 134, 2015 (Hentschel et al. 2015). Nom. nov. pro Frullania tamarisci var. azorica J.-P.Frahm, Trop. Bryol. 27: 102, 2006 (Frahm 2006). 223 ** Frullania alstonii Verd., Ann. Bryol., Suppl. 1: 76, 1930 (Verdoorn 1930c). ** Frullania aoshimensis Horik., Sci. Rep. Tôhoku Imp. Univ., Ser. 4, Biol. 4 (1): 64, 1929 (Horikawa 1929b). *** Frullania appendiculata Steph., Bull. Herb. Boissier 5 (2): 88, 1897 (Stephani 1897b). 224 *** Frullania asagrayana Mont., Ann. Sci. Nat. Bot. (sér. 2) 18: 14, 1842 (Montagne 1842b). 225 *** Frullania calcarifera Steph., Bol. Soc. Brot. 4: 241, 1886 [1887] (Henriques 1886). 226 *** Frullania californica (M.Howe) A.Evans, Trans. Connecticut Acad. Arts 10 (1): 25, 1899 (Evans 1899). Bas.: Frullania asagrayana var. californica M.Howe, Erythea 2 (6): 98, 1894 (Howe 1894). *** Frullania densiloba Steph. ex A.Evans, Proc. Wash. Acad. Sci. 8: 157, 1906 (Evans 1906b). *** Frullania franciscana M.Howe, Erythea 2 (6): 99, 1894 (Howe 1894). 223 Frullania acicularis was shown to be a species separate from Frullania tamarisci (Heinrichs et al. 2010, Vilnet et al. (2014). It may, however, not be the earliest available name. 224 Frullania appendiculata was shown by Vilnet et al. (2014) to be a species separate from Frullania tamarisci. It may, however, not be the earliest available name. 225 Frullania asagrayana consists of two geographically separated clades (Ramaiya et al. 2010). 226 Frullania calcarifera was shown to be a species separate from Frullania tamarisci by Heinrichs et al. (2010) and Vilnet et al. (2014). It may, however, not be the earliest available name. World checklist of hornworts and liverworts 293 ** Frullania iwatsukii S.Hatt., J. Hattori Bot. Lab. 35: 240, 1972 (Hattori 1972a). *** Frullania microphylla (Gottsche) Pearson, J. Bot. 32: 328, 1894 (Pearson 1894). Bas.: Frullania tamarisci var. microphylla Gottsche, Hepat. Eur., Leberm. 21-22: no. 109 [209], 1862 (Rabenhorst 1862). *** Frullania moniliata (Reinw., Blume et Nees) Mont., Ann. Sci. Nat. Bot. (sér. 2) 18: 13, 1842 (Montagne 1842b). Bas.: Jungermannia moniliata Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 224, 1824 [1825] (Reinwardt et al. 1824a). 227 *** Frullania nisquallensis Sull., Mem. Amer. Acad. Arts (n.ser.) 4: 175, 1849 (Sullivant 1849). *** Frullania polysticta Lindenb., Syn. Hepat. 3: 440, 1845 (Gottsche et al. 1845b). ** Frullania pseudoalstonii Tsudo et J.Haseg., Bryol. Res. 9 (3): 44, 2006 (Tsudo and Hasegawa 2006). ** Frullania punctata Reimers, Hedwigia 71 (1/2): 36, 1931 (Reimers 1931). ** Frullania schaefer-verwimpii Yuzawa et S.Hatt., J. Jap. Bot. 64 (2): 37, 1989 (Yuzawa and Hattori 1989). ** Frullania selwyniana Pearson, List. Canad. Hepat.: 1, 1890 (Pearson 1890). *** Frullania sergiae Sim-Sim, Fontinha, Mues et Lion, Nova Hedwigia 71 (1/2): 186, 2000 (Sim-Sim et al. 2000). *** Frullania subarctica Vilnet, Borovich. et Bakalin, Phytotaxa 173 (1): 67, 2014 (Vilnet et al. 2014). 228 *** Frullania tamarisci (L.) Dumort., Recueil Observ. Jungerm.: 13, 1835 (Dumortier 1835). Bas.: Jungermannia tamarisci L., Sp. Pl. 1: 1134, 1753 (Linnaeus 1753). *** Frullania tenerifae (F.Weber) Nees, Naturgesch. Eur. Leberm. 3: 239, 1838 (Nees 1838b). Bas.: Jungermannia tenerifae F.Weber, Hist. Musc. Hepat. Prodr.: 23, 1815 (Weber 1815). ** Frullania trigona L.Clark, Jovet-Ast et Frye, Bryologist 50 (1): 52, 1947 (Clark et al. 1947). Incertae sedis Frullania ainis Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 257, 1843 (Montagne 1843). 229 * Frullania allionii Steph., Sp. Hepat. (Stephani) 4: 394, 1910 (Stephani 1910b). * Frullania alpina Steph., Sp. Hepat. (Stephani) 4: 533, 1911 (Stephani 1911e). ** Frullania alternans Nees, Syn. Hepat. 3: 430, 1845 (Gottsche et al. 1845b). * Frullania apertilobula Gerola, Lav. Bot. Ist. Bot. Univ. Padova 12: 478, 1947 (Gerola 1947). ** Frullania armata Herzog et L.Clark, Bryologist 57 (1): 36, 1954 (Clark 1954). * 227 Frullania moniliata is a species complex (Vilnet et al. 2014). 228 Frullania subarctica was shown to be a species separate from the Frullania tamarisci complex (Vilnet et al. 2014), but it is maybe not the oldest name. 229 Frullania ainis is possibly conspeciic with Frullania ecklonii (Arnell 1963b). 294 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Frullania bolanderi Austin, Proc. Acad. Nat. Sci. Philadelphia 21: 226, 1869 (Austin 1869). *** Frullania boveana C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 244, 1885 (Massalongo 1885). * Frullania caespitans Beauverd, Sp. Hepat. (Stephani) 6: 537, 1924 (Stephani 1924). Nom. nov. pro Frullania campanulata Steph., Biblioth. Bot. 87 (2): 242, 1916 (Stephani 1916a), nom. illeg. * Frullania canaliculata Gottsche, Sp. Hepat. (Stephani) 4: 391, 1910 (Stephani 1910b). ** Frullania capillaris Steph., Sp. Hepat. (Stephani) 4: 616, 1911 (Stephani 1911e). * Frullania cavallii Gola, Ann. Bot. (Rome) 6 (2): 275, 1907 (Gola 1907). * Frullania chiapasana Steph., Sp. Hepat. (Stephani) 4: 392, 1910 (Stephani 1910b). * Frullania chilcootiensis Steph., Bot. Jahrb. Syst. 8 (2): 98, 1886 (Stephani 1886a). * Frullania chiovendae Gola, Ann. Bot. (Rome) 13 (1): 71, 1914 (Gola 1914a). ** Frullania ciliata Lindenb. et Gottsche, Syn. Hepat. 5: 775, 1847 (Gottsche et al. 1847). ** Frullania cinchonae Gottsche, Syn. Hepat. 3: 455, 1845 (Gottsche et al. 1845b). * Frullania complicata Steph., Sp. Hepat. (Stephani) 4: 671, 1911 (Stephani 1911e). * Frullania cordaeana Lindenb., Syn. Hepat. 3: 463, 1845 (Gottsche et al. 1845b). ** Frullania cuneatistipula Steph., Sp. Hepat. (Stephani) 6: 538, 1924 (Stephani 1924). * Frullania cuspiloba Steph., Sp. Hepat. (Stephani) 4: 394, 1910 (Stephani 1910b). * Frullania dispar Nees, Syn. Hepat. 3: 429, 1845 (Gottsche et al. 1845b). * Frullania guatemalensis Steph., Sp. Hepat. (Stephani) 4: 497, 1911 (Stephani 1911e). *** Frullania inlata Gottsche, Syn. Hepat. 3: 424, 1845 (Gottsche et al. 1845b). ** Frullania inlata var. communis R.M.Schust., Phytologia 57 (5): 372, 1985 (Schuster 1985a). ** Frullania inlata var. dioica S.Hatt. et haithong, J. Hattori Bot. Lab. 44: 185, 1978 (Hattori and haithong 1978b). ** Frullania inlata var. mayebarae (S.Hatt.) K.Yamada, Misc. Bryol. Lichenol. 6 (9): 163, 1974 (Yamada 1974). Bas.: Frullania mayebarae S.Hatt., Bot. Mag. (Tokyo) 65 (763/764): 13, 1952 (Hattori 1952a). * Frullania laetevirens Hampe ex Gottsche, Lindenb. et Nees, Syn. Hepat. 3: 420, 1845 (Gottsche et al. 1845b). * Frullania larjiana Sushil K.Singh et D.K.Singh, J. Bryol. 27 (2): 105, 2005 (Singh and Singh 2005). * Frullania larjiana var. didyhatii S.N.Srivast. et M.Rai, Geophytology 41 (1/2): 109, 2011 (Srivastava and Rai 2011). * Frullania leana Austin, Proc. Acad. Nat. Sci. Philadelphia 21: 227, 1869 (Austin 1869). * Frullania madagascariensis Gottsche, Abh. Naturwiss. Vereins Bremen 7: 364, 1882 (Gottsche 1882). * Frullania mauritiana Austin, Proc. Acad. Nat. Sci. Philadelphia 21: 227, 1869 (Austin 1869). *** Frullania mirabilis J.B.Jack et Steph., Hedwigia 31 (1): 15, 1892 (Jack and Stephani 1892). World checklist of hornworts and liverworts 295 ** Frullania monoica Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 6): 67, 1900 (Stephani 1900b). * Frullania ocanniensis Steph., Sp. Hepat. (Stephani) 6: 542, 1924 (Stephani 1924). *** Frullania platycalyx Herzog, Feddes Repert. Spec. Nov. Regni Veg. 55 (1): 10, 1952 (Herzog 1952h). * Frullania pyricalycina Steph., Sp. Hepat. (Stephani) 4: 394, 1910 (Stephani 1910b). ** Frullania quillotensis (Nees et Mont.) Nees et Mont., Syn. Hepat. 3: 427, 1845 (Gottsche et al. 1845b). Bas.: Jubula quillotensis Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 5: 64, 1836 (Nees and Montagne 1836). ** Frullania reicheana Steph., Sp. Hepat. (Stephani) 4: 427, 1910 (Stephani 1910b). * Frullania sabanetica Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 170, 1864 (Gottsche 1864). * Frullania semienana Gola, Ann. Bot. (Rome) 13 (1): 71, 1914 (Gola 1914a). ** Frullania serrifolia Steph., Sp. Hepat. (Stephani) 4: 526, 1911 (Stephani 1911e). ** Frullania subpyricalycina Herzog, Rev. Bryol. Lichénol. 23 (1/2): 57, 1954 (Herzog 1954). ** Frullania subtruncata Steph., Sp. Hepat. (Stephani) 4: 389, 1910 (Stephani 1910b). ** Frullania turfosa Lindenb. et Gottsche, Syn. Hepat. 5: 779, 1847 (Gottsche et al. 1847). ** Frullania udarii V.Nath et Ajit P.Singh, Curr. Sci. 91 (6): 744, 2006 (Nath and Singh 2006). ** Frullania valparaisiana Lehm., Nov. Stirp. Pug. 10: 17, 1857 (Lehmann 1857). ** Frullania wagneri Steph., Sp. Hepat. (Stephani) 4: 392, 1910 (Stephani 1910b). *** Jubulaceae H.Klinggr. by J. Hentschel and M. von Konrat he systematic placement of Neohattoria herzogii has been contentious since its description six decades ago. It has been interpreted as either a member of the genus Frullania or segregated into its own genus, Neohattoria, due to morphological similarities with both Frullania and Jubula. We follow Larrain et al. (2015) that provided molecular evidence supporting the recognition of the genus Neohattoria and its inclusion within the Jubulaceae, together with Jubula and Nipponolejeunea. *** Jubula Dumort., Commentat. Bot. (Dumortier): 112, 1822 (Dumortier 1822) nom. conserv. *** Jubula blepharophylla Grolle, J. Hattori Bot. Lab. 28: 47, 1965 (Grolle 1965b). ** Jubula hattorii Udar et V.Nath, Misc. Bryol. Lichenol. 8 (3): 49, 1978 (Udar and Nath 1978). ** Jubula hattorii var. muthukuzhiana A.E.D.Daniels et P.Daniel, Bryol. Southernm. W Ghats: 181, 2013 (Daniels and Daniel 2013). 296 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Jubula himalayensis S.C.Srivast. et D.Sharma, Proc. Indian Acad. Sci. Pl. Sci. 100 (2): 85, 1990 (Srivastava and Sharma 1990). *** Jubula hutchinsiae (Hook.) Dumort., Syll. Jungerm. Europ.: 36, 1831 (Dumortier 1831). Bas.: Jungermannia hutchinsiae Hook., Brit. Jungermann.: tab. 1, 1812 (Hooker 1812). 230 ** Jubula hutchinsiae subsp. australiae Pócs et A.Cairns, Nova Hedwigia 86 (1/2): 232, 2008 (Pócs and Cairns 2008). ** Jubula hutchinsiae subsp. bogotensis (Steph.) Verd., Ann. Cryptog. Exot. 1 (2): 215, 1928 (Verdoorn 1928b). Bas.: Jubula bogotensis Steph., Sp. Hepat. (Stephani) 4: 687, 1911 (Stephani 1911e). 231 *** Jubula hutchinsiae subsp. caucasica Konstant. et Vilnet, Arctoa 20: 234, 2011 (Konstantinova and Vilnet 2011). *** Jubula hutchinsiae subsp. japonica (Steph.) Horik. et Ando, J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 6: 300, 1954 (Horikawa and Ando 1954). Bas.: Jubula japonica Steph., Bull. Herb. Boissier 5 (2): 92, 1897 (Stephani 1897b). *** Jubula hutchinsiae subsp. javanica (Steph.) Verd., Ann. Cryptog. Exot. 1 (2): 216, 1928 (Verdoorn 1928b). Bas.: Jubula javanica Steph., Sp. Hepat. (Stephani) 4: 688, 1911 (Stephani 1911e). 232 *** Jubula hutchinsiae subsp. pennsylvanica (Steph.) Verd., Ann. Cryptog. Exot. 1 (2): 215, 1928 (Verdoorn 1928b). Bas.: Frullania pennsylvanica Steph., Hedwigia 22 (10): 147, 1883 (Stephani 1883). ** Jubula kwangsiensis C.Gao et K.C.Chang, Bull. Bot. Res., Harbin 4 (3): 89, 1984 (Chang and Gao 1984). *** Neohattoria Kamim., J. Jap. Bot. 37 (7): 218, 1962 (Kamimura 1962). Nom. nov. pro Hattoria Kamim., J. Hattori Bot. Lab. 24: 93, 1961 (Kamimura 1961). *** Neohattoria herzogii (S.Hatt.) Kamim., J. Jap. Bot. 37 (7): 218, 1962 (Kamimura 1962). Bas.: Frullania herzogii S.Hatt., Feddes Repert. Spec. Nov. Regni Veg. 58: 53, 1955 (Hattori 1955). *** Nipponolejeunea S.Hatt., Bull. Tokyo Sci. Mus. 11: 124, 1944 (Hattori 1944d). *** Nipponolejeunea pilifera (Steph.) S.Hatt., Bull. Tokyo Sci. Mus. 11: 125, 1944 (Hattori 1944d). Bas.: Pycnolejeunea pilifera Steph., Sp. Hepat. (Stephani) 5: 624, 1914 (Stephani 1914b). *** Nipponolejeunea subalpina (Horik.) S.Hatt., Bull. Tokyo Sci. Mus. 11: 125, 1944 (Hattori 1944d). Bas.: Pycnolejeunea subalpina Horik., J. Jap. Bot. 15: 360, 1939 (Horikawa 1939). 230 Jubula hutchinsiae has subspecies that may be better recognized at the species level. 231 Jubula hutchinsiae subsp. bogotensis is closely related to subsp. pennsylvanica (Pätsch et al. 2010). 232 Jubula hutchinsiae subsp. javanica includes a cryptic taxon from China (Pätsch et al. 2010). World checklist of hornworts and liverworts *** 297 Lejeuneaceae Cavers by S.R. Gradstein, T. Pócs and R.-L. Zhu with contributions by G. Dauphin (Ceratolejeunea), X. He (Pycnolejeunea), A.-L. Ilkiu-Borges (Prionolejeunea), E. ReinerDrehwald (Rectolejeunea, Lejeunea), A. Sass-Gyarmati (Lopholejeunea), A. Schäfer-Verwimp (Diplasiolejeunea) and P. Sukkharak (hysananthus) he subdivision of Lejeuneaceae follows Gradstein (2013c) with updates from Heinrichs et al. (2013, 2014) and Schäfer-Verwimp et al. (2014). Some nomenclatural and taxonomic notes can also be found in Gradstein (2013b), Shu and Zhu (2014), Pócs et al. (2015a, 2015c), Qui et al. (2014) and Söderström et al. (2015a). *** *** *** Lejeuneoideae trib. Brachiolejeuneeae subtrib. Brachiolejeuneinae Gradst. ** Acanthocoleus R.M.Schust., Bull. Torrey Bot. Club 97 (6): 339, 1970 [1971] (Schuster 1970b). *** Acanthocoleus aberrans (Lindenb. et Gottsche) Kruijt, Bryophyt. Biblioth. 36: 62, 1988 (Kruijt 1988). Bas.: Lejeunea aberrans Lindenb. et Gottsche, Syn. Hepat. 5: 751, 1847 (Gottsche et al. 1847). *** Acanthocoleus aberrans var. laevis Gradst., Fl. Neotrop. Monogr. 62: 193, 1994 (Gradstein 1994). *** Acanthocoleus chrysophyllus (Lehm.) Kruijt, Bryophyt. Biblioth. 36: 72, 1988 (Kruijt 1988). Bas.: Jungermannia chrysophylla Lehm., Linnaea 9 (4): 423, 1835 (Lehmann 1835). ** Acanthocoleus elgonensis Gyarmati et Pócs, Cryptog. Bryol. 35 (2): 120, 2014 (SassGyarmati and Pócs 2014). *** Acanthocoleus gilvus (Gottsche) Kruijt, Bryophyt. Biblioth. 36: 79, 1988 (Kruijt 1988). Bas.: Lejeunea gilva Gottsche, Abh. Naturwiss. Vereins Bremen 7: 353, 1882 (Gottsche 1882). *** Acanthocoleus javanicus (Steph.) Kruijt, Bryophyt. Biblioth. 36: 85, 1988 (Kruijt 1988). Bas.: Dicranolejeunea javanica Steph., Sp. Hepat. (Stephani) 5: 169, 1912 (Stephani 1912c). *** Acanthocoleus juddii Kruijt, Bryophyt. Biblioth. 36: 93, 1988 (Kruijt 1988). ** Acanthocoleus madagascariensis (Steph.) Kruijt, Bryophyt. Biblioth. 36: 98, 1988 (Kruijt 1988). Bas.: Dicranolejeunea madagascariensis Steph., Sp. Hepat. (Stephani) 5: 158, 1912 (Stephani 1912c). *** Acanthocoleus trigonus (Nees et Mont.) Gradst., Contr. Univ. Michigan Herb. 18: 101, 1992 (Gradstein 1992b). Bas.: Lejeunea trigona Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 5: 61, 1836 (Nees and Montagne 1836). 298 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Acanthocoleus yoshinaganus (S.Hatt.) Kruijt, Bryophyt. Biblioth. 36: 105, 1988 (Kruijt 1988). Bas.: Lopholejeunea subfusca var. yoshinagana S.Hatt., Bot. Mag. (Tokyo) 58 (686): 38, 1944 (Hattori 1944a). *** Blepharolejeunea S.W.Arnell, Svensk Bot. Tidskr. 56 (2): 335, 1962 (Arnell 1962b). *** Blepharolejeunea chimantaensis van Slageren et Kruijt, Beih. Nova Hedwigia 80: 126, 1985 (van Slageren and Kruijt 1985). *** Blepharolejeunea fuegiana (Besch. et C.Massal.) Gradst., Beih. Nova Hedwigia 80: 130, 1985 (van Slageren and Kruijt 1985). Bas.: Lejeunea fuegiana Besch. et C. Massal., Bull. Mens. Soc. Linn. Paris 1 (79): 638, 1886 (Bescherelle and Massalongo 1886). *** Blepharolejeunea incongrua (Lindenb. et Gottsche) van Slageren et Kruijt, Beih. Nova Hedwigia 80: 133, 1985 (van Slageren and Kruijt 1985). Bas.: Lejeunea incongrua Lindenb. et Gottsche, Syn. Hepat. 5: 750, 1847 (Gottsche et al. 1847). *** Blepharolejeunea saccata (Steph.) van Slageren et Kruijt, Beih. Nova Hedwigia 80: 138, 1985 (van Slageren and Kruijt 1985). Bas.: Dicranolejeunea saccata Steph., Hedwigia 35 (3): 78, 1896 (Stephani 1896b). *** Blepharolejeunea securifolia (Steph.) R.M.Schust., Phytologia 45 (5): 424, 1980 (Schuster 1980b). Bas.: Brachiolejeunea securifolia Steph., Sp. Hepat. (Stephani) 5: 128, 1912 (Stephani 1912c). *** Brachiolejeunea (Spruce) Schifn., Hepat. (Engl.-Prantl): 128, 1893 (Schifner 1893b). Bas.: Lejeunea subg. Brachiolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 129, 1884 (Spruce 1884). *** Brachiolejeunea conduplicata (Steph.) Gradst., Fl. Neotrop. Monogr. 62: 175, 1994 (Gradstein 1994). Bas.: Archilejeunea conduplicata Steph., Sp. Hepat. (Stephani) 4: 712, 1911 (Stephani 1911e). *** Brachiolejeunea fernandeziana S.W.Arnell, Ark. Bot. (n.ser.) 4 (1): 18, 1957 (Arnell 1957b). *** Brachiolejeunea laxifolia (Taylor) Schifn., Hepat. (Engl.-Prantl): 128, 1893 (Schifner 1893b). Bas.: Phragmicoma laxifolia Taylor, London J. Bot. 6: 341, 1847 (Taylor 1847b). *** Brachiolejeunea leiboldiana (Gottsche et Lindenb.) Schifn., Hedwigia 33 (4): 182, 1894 (Schifner 1894). Bas.: Phragmicoma leiboldiana Gottsche et Lindenb., Syn. Hepat. 2: 296, 1845 (Gottsche et al. 1845a). *** Brachiolejeunea phyllorhiza (Nees) Kruijt et Gradst., Nova Hedwigia 43 (3/4): 299, 1986 (Kruijt and Gradstein 1986). Bas.: Jungermannia phyllorhiza Nees, Fl. Bras. (Martius) 1 (1): 348, 1833 (Nees 1833a). *** Brachiolejeunea spruceana (C.Massal.) Schifn., Hepat. (Engl.-Prantl): 128, 1893 (Schifner 1893b). Bas.: Lejeunea spruceana C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 246, 1885 (Massalongo 1885). World checklist of hornworts and liverworts 299 *** Dicranolejeunea (Spruce) Schifn., Hepat. (Engl.-Prantl): 128, 1893 (Schifner 1893b). Bas.: Lejeunea subg. Dicranolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 138, 1884 (Spruce 1884). *** Dicranolejeunea axillaris (Nees et Mont.) Schifn., Hepat. (Engl.-Prantl): 128, 1893 (Schifner 1893b). Bas.: Lejeunea axillaris Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 5: 59, 1836 (Nees and Montagne 1836). * Dicranolejeunea bovonei Gola, Nuovo Giorn. Bot. Ital. (n.ser.) 27 (2/4): 248, 1920 (Gola 1920). *** Lindigianthus Kruijt et Gradst., Beih. Nova Hedwigia 80: 165, 1985 (Kruijt and Gradstein 1985). *** Lindigianthus cipaconeus (Gottsche) Kruijt et Gradst., Beih. Nova Hedwigia 80: 166, 1985 (Kruijt and Gradstein 1985). Bas.: Lejeunea cipaconea Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 150, 1864 (Gottsche 1864). *** Odontolejeunea (Spruce) Schifn., Hepat. (Engl.-Prantl): 127, 1893 (Schifner 1893b). Bas.: Lejeunea subg. Odontolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 142, 1884 (Spruce 1884). *** Odontolejeunea decemdentata (Spruce) Steph., Sp. Hepat. (Stephani) 5: 175, 1912 (Stephani 1912c). Bas.: Lejeunea decemdentata Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 148, 1884 (Spruce 1884). *** Odontolejeunea lunulata (F.Weber) Schifn., Hepat. (Engl.-Prantl): 128, 1893 (Schifner 1893b). Bas.: Jungermannia lunulata F.Weber, Hist. Musc. Hepat. Prodr.: 33, 1815 (Weber 1815). *** Odontolejeunea rhomalea (Spruce) Steph., Sp. Hepat. (Stephani) 5: 176, 1912 (Stephani 1912c). Bas.: Lejeunea rhomalea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 147, 1884 (Spruce 1884). *** subtrib. Stictolejeuneinae Gradst. *** Neurolejeunea (Spruce) Schifn., Hepat. (Engl.-Prantl): 131, 1893 (Schifner 1893b). Bas.: Lejeunea subg. Neurolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 84, 1884 (Spruce 1884). *** Neurolejeunea breutelii (Gottsche) A.Evans, Bull. Torrey Bot. Club 34 (1): 13, 1907 (Evans 1907b). Bas.: Lejeunea breutelii Gottsche, Syn. Hepat. 3: 324, 1845 (Gottsche et al. 1845b). ** Neurolejeunea breutelii var. africana Pócs, Herzogia 28 (1): 63, 2015 (Pócs et al. 2015b). *** Neurolejeunea catenulata (Nees) Schifn., Hepat. (Engl.-Prantl): 131, 1893 (Schifner 1893b). Bas.: Lejeunea catenulata Nees, Syn. Hepat. 3: 323, 1845 (Gottsche et al. 1845b). *** Neurolejeunea sastreana Gradst., Bryologist 92 (3): 345, 1989 (Gradstein 1989). 300 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Neurolejeunea seminervis (Spruce) Schifn., Hepat. (Engl.-Prantl): 131, 1893 (Schifner 1893b). Bas.: Lejeunea seminervis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 84, 1884 (Spruce 1884). *** Stictolejeunea (Spruce) Schifn., Hepat. (Engl.-Prantl): 131, 1893 (Schifner 1893b). Bas.: Lejeunea subg. Stictolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 81, 1884 (Spruce 1884). *** subg. Leptostictolejeunea R.M.Schust., Phytologia 56 (2): 70, 1984 (Schuster 1984). *** Stictolejeunea balfourii (Mitt.) E.W.Jones, J. Bryol. 9 (1): 50, 1976 (Jones 1976). Bas.: Lejeunea balfourii Mitt., Philos. Trans. 168: 398, 1879 (Mitten 1879). *** Stictolejeunea balfourii var. bekkeri Gradst., Beih. Nova Hedwigia 80: 214, 1985 (Gradstein 1985b). *** Stictolejeunea iwatsukii Mizut., J. Hattori Bot. Lab. 44: 134, 1978 (Mizutani 1978). *** subg. Stictolejeunea *** Stictolejeunea squamata (Willd.) Schifn., Hepat. (Engl.-Prantl): 131, 1893 (Schifner 1893b). Bas.: Jungermannia squamata Willd. ex F.Weber, Hist. Musc. Hepat. Prodr.: 33, 1815 (Weber 1815). *** trib. Lejeuneeae Dumort. ** Dactylophorella R.M.Schust., Phytologia 45 (5): 427, 1980 (Schuster 1980b). *** Dactylophorella muricata (Gottsche) R.M.Schust., Phytologia 45 (5): 427, 1980 (Schuster 1980b). Bas.: Lejeunea muricata Gottsche, Syn. Hepat. 3: 348, 1845 (Gottsche et al. 1845b). *** Metalejeunea Grolle, Bryophyt. Biblioth. 48: 17, 1995 (Grolle 1995). ** Metalejeunea crassitexta (J.B.Jack et Steph.) Pócs, Telopea 13 (3): 456, 2011 (Pócs et al. 2011). Bas.: Microlejeunea crassitexta J.B.Jack et Steph., Bot. Centralbl. 60 (4): 106, 1894 (Jack and Stephani 1894). *** Metalejeunea cucullata (Reinw., Blume et Nees) Grolle, Bryophyt. Biblioth. 48: 100, 1995 (Grolle 1995). Bas.: Jungermannia cucullata Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 227, 1824 [1825] (Reinwardt et al. 1824a). *** Metalejeunea winkleri R.L.Zhu et Grolle, Nova Hedwigia 74 (3/4): 498, 2002 (Zhu and Grolle 2002). *** Pictolejeunea Grolle, Feddes Repert. 88 (4): 248, 1977 (Grolle 1977b). *** Pictolejeunea levis Grolle et M.E.Reiner, J. Bryol. 27 (3): 281, 2005 (Grolle and Reiner-Drehwald 2005). World checklist of hornworts and liverworts 301 *** Pictolejeunea mizutanii Grolle, Feddes Repert. 88 (4): 255, 1977 (Grolle 1977b). Nom. nov. pro Cheilolejeunea picta Mizut., J. Hattori Bot. Lab. 33: 226, 1970 (Mizutani 1970). ** Pictolejeunea piconii Pócs, Acta Bot. Hung. 49 (1/2): 110, 2007 (Pócs 2007). *** Pictolejeunea picta (Steph.) Grolle, Feddes Repert. 88 (4): 252, 1977 (Grolle 1977b). Bas.: Prionolejeunea picta Steph., Sp. Hepat. (Stephani) 5: 223, 1913 (Stephani 1913a). *** Pictolejeunea reginae Ilk.-Borg., Brittonia 54 (4): 318, 2002 [2003] (Ilkiu-Borges 2002). *** Pictolejeunea sprucei Grolle, Feddes Repert. 88 (4): 249, 1977 (Grolle 1977b). ** subtrib. Ceratolejeuneinae Gradst. *** Ceratolejeunea (Spruce) J.B.Jack et Steph., Hedwigia 31 (1): 16, 1892 (Jack and Stephani 1892). Bas.: Lejeunea subg. Ceratolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 198, 1884 (Spruce 1884). 233 *** subg. Ceratolejeunea *** Ceratolejeunea andringitrae Pócs, Polish Bot. J. 56 (2): 144, 2011 (Pócs 2011c). ** Ceratolejeunea atlantica Alvarenga et Ilk.-Borg., Nova Hedwigia 86 (1/2): 238, 2008 (Ilkiu-Borges and Alvarenga 2008). ** Ceratolejeunea belangeriana (Gottsche) Steph., Sp. Hepat. (Stephani) 5: 396, 1913 (Stephani 1913a). Bas.: Lejeunea belangeriana Gottsche, Syn. Hepat. 3: 398, 1845 (Gottsche et al. 1845b). ** Ceratolejeunea beninensis E.W.Jones et Vanden Berghen, Bull. Jard. Bot. État Bruxelles 21 (1/2): 63, 1951 (Vanden Berghen 1951b). *** Ceratolejeunea brevinervis (Spruce) A.Evans, Bull. Torrey Bot. Club 32 (6): 282, 1905 (Evans 1905a). Bas.: Lejeunea brevinervis Spruce, J. Linn. Soc., Bot. 30 (210): 342, 1895 (Gepp 1895b). *** Ceratolejeunea ceratantha (Nees et Mont.) Schifn., Bot. Jahrb. Syst. 23 (5): 582, 1897 (Schifner 1897). Bas.: Lejeunea ceratantha Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 14: 335, 1840 (Montagne 1840a). ** Ceratolejeunea coalita (Ångstr.) Steph., Sp. Hepat. (Stephani) 5: 402, 1913 (Stephani 1913a). Bas.: Lejeunea coalita Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 30 (5): 135, 1873 (Ångström 1873). *** Ceratolejeunea coarina (Gottsche) Schifn., Hepat. (Engl.-Prantl): 125, 1893 (Schifner 1893b). Bas.: Lejeunea coarina Gottsche, Syn. Hepat. 3: 395, 1845 (Gottsche et al. 1845b). 233 he treatment of Ceratolejeunea follows Dauphin (2003) except that his subg. Caduciloba becomes subg. Ceratolejeunea and his subg. Ceratolejeunea becomes subg. Ceratophora as a consequence of the introduction of ICN Art. 41.4 (cf. Söderström et al. 2015a). 302 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Ceratolejeunea confusa R.M.Schust., J. Elisha Mitchell Sci. Soc. 72 (2): 313, 1956 (Schuster 1956a). *** Ceratolejeunea cornuta (Lindenb.) Steph., Planzenw. Ost-Afrikas C: 65, 1895 (Stephani 1895d). Bas.: Jungermannia cornuta Lindenb., Syn. hepat. eur: 23, 1829 (Lindenberg 1829). *** Ceratolejeunea cubensis (Mont.) Schifn., Hepat. (Engl.-Prantl): 125, 1893 (Schifner 1893b). Bas.: Lejeunea cubensis Mont., Hist. Phys. Cuba, Bot., Pl. Cell.: 481, 1842 (Montagne 1842a). *** Ceratolejeunea dentistipula Steph., Sp. Hepat. (Stephani) 5: 407, 1913 (Stephani 1913a). *** Ceratolejeunea fallax (Lehm. et Lindenb.) Bonner, Candollea 14: 189, 1953 (Bonner 1953a). Bas.: Jungermannia fallax Lehm. et Lindenb., Nov. Stirp. Pug. 5: 17, 1833 (Lehmann 1833). *** Ceratolejeunea ilaria (Taylor) Steph., Sp. Hepat. (Stephani) 5: 412, 1913 (Stephani 1913a). Bas.: Lejeunea ilaria Taylor, Nov. Stirp. Pug. 8: 28, 1844 (Lehmann 1844). ** Ceratolejeunea loribunda Steph., Sp. Hepat. (Stephani) 5: 412, 1913 (Stephani 1913a). *** Ceratolejeunea guianensis (Nees et Mont.) Steph., Sp. Hepat. (Stephani) 5: 416, 1913 (Stephani 1913a). Bas.: Lejeunea guianensis Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 14: 335, 1840 (Montagne 1840a). * Ceratolejeunea karstenii Steph., Sp. Hepat. (Stephani) 5: 420, 1913 (Stephani 1913a). * Ceratolejeunea kuerschneri Eb.Fisch. et Vanderp., Beih. Nova Hedwigia 138: 87, 2010 (Fischer and Vanderpoorten 2010). 234 *** Ceratolejeunea laetefusca (Austin) R.M.Schust., J. Elisha Mitchell Sci. Soc. 72 (2): 306, 1956 (Schuster 1956a). Bas.: Lejeunea laetefusca Austin, Bot. Bull. (Hanover) 1 (8): 36, 1876 (Austin 1876a). * Ceratolejeunea ledermannii Steph., Sp. Hepat. (Stephani) 6: 399, 1923 (Stephani 1923). *** Ceratolejeunea malleigera (Spruce) Steph., Sp. Hepat. (Stephani) 5: 422, 1913 (Stephani 1913a). Bas.: Lejeunea malleigera Spruce, Mem. Torrey Bot. Club 1 (3): 123, 1890 (Spruce 1890). ** Ceratolejeunea maranhensis Silva Brito et Ilk.-Borg., Nova Hedwigia 95 (3/4): 424, 2012 (Brito and Ilkiu-Borges 2012). ** Ceratolejeunea minor Mizut., J. Hattori Bot. Lab. 49: 311, 1981 (Mizutani 1981). *** Ceratolejeunea minuta G.Dauphin, Fl. Neotrop. Monogr. 90: 66, 2003 (Dauphin 2003). ** Ceratolejeunea moniliata Herzog, Mitt. Inst. Allg. Bot. Hamburg 7 (3): 205, 1931 (Herzog 1931a). ** Ceratolejeunea oculata (Gottsche) Steph., Bull. Herb. Boissier 5 (10): 842, 1897 (Stephani 1897c). Bas.: Lejeunea oculata Gottsche, Syn. Hepat. 3: 357, 1845 (Gottsche et al. 1845b). *** Ceratolejeunea oxygonia Steph., Sp. Hepat. (Stephani) 5: 429, 1913 (Stephani 1913a). 234 Ceratolejeunea kuerschneri is very similar to Ceratolejeunea papulilora and may be conspeciic with it. World checklist of hornworts and liverworts 303 *** Ceratolejeunea papulilora Steph., Sp. Hepat. (Stephani) 5: 430, 1913 (Stephani 1913a). *** Ceratolejeunea patentissima (Hampe et Gottsche) A.Evans, Bull. Torrey Bot. Club 32 (6): 286, 1905 (Evans 1905a). Bas.: Lejeunea patentissima Hampe et Gottsche, Linnaea 25 (3): 355, 1852 [1853] (Hampe and Gottsche 1852). *** Ceratolejeunea pungens Steph., Sp. Hepat. (Stephani) 5: 434, 1913 (Stephani 1913a). *** Ceratolejeunea rubiginosa Steph., Hedwigia 34 (5): 237, 1895 (Stephani 1895b). *** Ceratolejeunea saroltae Pócs, Polish Bot. J. 56 (2): 150, 2011 (Pócs 2011c). * Ceratolejeunea sinensis P.C.Chen et P.C.Wu, Acta Phytotax. Sin. 9 (3): 232, 1964 (Chen and Wu 1964). 235 ** Ceratolejeunea singapurensis (Lindenb.) Schifn., Consp. Hepat. Arch. Ind.: 273, 1898 (Schifner 1898b). Bas.: Lejeunea singapurensis Lindenb., Syn. Hepat. 3: 397, 1845 (Gottsche et al. 1845b). *** Ceratolejeunea spinosa (Gottsche) Steph., Hedwigia 34 (5): 238, 1895 (Stephani 1895b). Bas.: Lejeunea spinosa Gottsche, Syn. Hepat. 3: 402, 1845 (Gottsche et al. 1845b). *** Ceratolejeunea temnantha (Spruce) M.E.Reiner, Cryptog. Bryol. 32 (2): 95, 2011 (Reiner-Drehwald 2011). Bas.: Lejeunea temnantha Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 250, 1884 (Spruce 1884). *** Ceratolejeunea umbonata Steph., Sp. Hepat. (Stephani) 5: 446, 1913 (Stephani 1913a). ** Ceratolejeunea vitiensis Steph., Sp. Hepat. (Stephani) 5: 448, 1913 (Stephani 1913a). ** Ceratolejeunea zenkeri Steph., Sp. Hepat. (Stephani) 5: 449, 1914 (Stephani 1914b). *** subg. Ceratophora R.M.Schust., J. Elisha Mitchell Sci. Soc. 72 (2): 294, 1956 (Schuster 1956a). *** Ceratolejeunea desciscens (Sande Lac.) Schifn., Hepat. (Engl.-Prantl): 126, 1893 (Schifner 1893b). Bas.: Lejeunea desciscens Sande Lac., Syn. hepat. jav.: 107, 1856 [1857] (Sande Lacoste 1856b). *** Ceratolejeunea globulifera Herzog, Rev. Bryol. Lichénol. 13: 23, 1942 (Herzog 1942e). *** Ceratolejeunea grandiloba J.B.Jack et Steph., Hedwigia 31 (1): 16, 1892 (Jack and Stephani 1892). ** Ceratolejeunea grandiloba subsp. inlata (Mizut.) Gradst., Phytotaxa 81 (1): 5, 2013 (Gradstein 2013d). Bas.: Ceratolejeunea inlata Mizut., J. Hattori Bot. Lab. 49: 313, 1981 (Mizutani 1981). *** Ceratolejeunea szyszylowiczii (Loitl.) Steph., Sp. Hepat. (Stephani) 5: 443, 1913 (Stephani 1913a). Bas.: Lejeunea szyszylowiczii Loitl., Diagn. pl. nov.: 19, 1894 (Loitlesberger 1894). Incertae sedis ** Ceratolejeunea aliena Herzog, Trans. Brit. Bryol. Soc. 2 (1): 71, 1952 (Herzog 1952a). 235 Ceratolejeunea sinensis may be conspeciic with Drepanolejeunea erecta (Zhu et al. 2005). 304 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Luteolejeunea Piippo, Acta Bot. Fenn. 132: 56, 1986 (Piippo 1986a). *** Luteolejeunea herzogii (Buchloh) Piippo, Acta Bot. Fenn. 132: 57, 1986 (Piippo 1986a). Bas.: Stictolejeunea herzogii Buchloh, Nova Hedwigia 3 (4): 515, 1961 (Buchloh 1961). *** Otigoniolejeunea (Spruce) Schifn., Hepat. (Engl.-Prantl): 125, 1893 (Schifner 1893b). Bas.: Lejeunea subg. Otigoniolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 226, 1884 (Spruce 1884). * Otigoniolejeunea crenulata Steph., Sp. Hepat. (Stephani) 6: 408, 1923 (Stephani 1923). 236 *** Otigoniolejeunea huctumalcensis (Lindenb. et Gottsche) Y.M.Wei, R.L.Zhu et Gradst., Phytotaxa 162 (4): 237, 2014 (Wei et al. 2014). Bas.: Lejeunea huctumalcensis Lindenb. et Gottsche, Syn. Hepat. 5: 762, 1847 (Gottsche et al. 1847). * Otigoniolejeunea ledermannii Steph., Sp. Hepat. (Stephani) 6: 409, 1923 (Stephani 1923). 237 *** Otigoniolejeunea portoricensis (Hampe et Gottsche) Y.M.Wei, R.L.Zhu et Gradst., Phytotaxa 162 (4): 237, 2014 (Wei et al. 2014). Bas.: Lejeunea portoricensis Hampe et Gottsche, Linnaea 25 (3): 352, 1852 [1853] (Hampe and Gottsche 1852). *** subtrib. Cheilolejeuneinae Gradst. * Aureolejeunea R.M.Schust., Phytologia 39 (6): 428, 1978 (Schuster 1978b). 238 *** Aureolejeunea aurifera R.M.Schust., Phytologia 39 (6): 429, 1978 (Schuster 1978b). *** Aureolejeunea lumae (Herzog) van Slageren, Meded. Bot. Mus. Herb. Rijks Univ. Utrecht 544: 121, 1985 (van Slageren 1985). Bas.: Brachiolejeunea lumae Herzog, Beih. Bot. Centralbl. 60B (1/2): 15, 1939 (Herzog 1939c). *** Aureolejeunea paramicola (Herzog) R.M.Schust., Phytologia 61 (7): 446, 1987 (Schuster 1987c). Bas.: Brachiolejeunea paramicola Herzog, Hedwigia 74 (2): 95, 1934 (Herzog 1934a). *** Aureolejeunea quinquecarinata R.M.Schust., Phytologia 39 (6): 429, 1978 (Schuster 1978b). *** Aureolejeunea tonduzana (Steph.) Gradst., Phytotaxa 76 (3): 46, 2013 (Gradstein 2013a). Bas.: Archilejeunea tonduzana Steph., Sp. Hepat. (Stephani) 4: 721, 1911 (Stephani 1911e). 236 Otigoniolejeunea crenulata is a doubtful taxon. he type specimen was burned in B and it is unclear where the taxon belongs (Grolle and Piippo (1984). 237 Otigoniolejeunea ledermannii is a doubtful taxon. he type specimen was burned in B and it is unclear where the taxon belongs (Grolle and Piippo (1984). 238 Aureolejeunea is nested in Cheilolejeunea (Ye et al. 2011, Gradstein 2013a, Ye et al. 2015). A formal transfer of the genus and its species to Cheilolejeunea is found in Ye et al. (2015). World checklist of hornworts and liverworts 305 *** Cheilolejeunea (Spruce) Steph., Bot. Gaz. 15 (11): 284, 1890 (Stephani 1890c). Bas.: Lejeunea subg. Cheilolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 251, 1884 (Spruce 1884). 239 ** Cheilolejeunea sandvicensis (Prantl) Steph., Bull. Herb. Boissier 5 (10): 842, 1897 (Stephani 1897c). Bas.: Lejeunea sandvicensis Prantl, Hedwigia 29: xvii, 1890 (Prantl 1890). ** subg. Cheilolejeunea *** Cheilolejeunea acanthina (Spruce) Gradst. et Ilk.-Borg., Mem. New York Bot. Gard. 76 (4): 62, 2009 (Gradstein and Ilkiu-Borges 2009). Bas.: Lejeunea acanthina Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 182, 1884 (Spruce 1884). *** Cheilolejeunea adnata (Kunze ex Lehm.) Grolle, J. Bryol. 9 (4): 529, 1977 [1978] (Grolle 1977a). Bas.: Jungermannia adnata Kunze ex Lehm., Nov. Stirp. Pug. 6: 46, 1834 (Lehmann 1834). ** Cheilolejeunea adnata var. autoica Gradst. et Ilk.-Borg., Mem. New York Bot. Gard. 76 (4): 64, 2009 (Gradstein and Ilkiu-Borges 2009). ** Cheilolejeunea albovirens (Hook.f. et Taylor) E.A.Hodgs., Rec. Domin. Mus. 4 (11): 127, 1962 (Hodgson 1962a). Bas.: Jungermannia albovirens Hook.f. et Taylor, London J. Bot. 3: 397, 1844 (Hooker and Taylor 1844a). *** Cheilolejeunea aneogyna (Spruce) A.Evans, Trans. Connecticut Acad. Arts 10 (8): 440, 1900 (Evans 1900a). Bas.: Lejeunea aneogyna Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 254, 1884 (Spruce 1884). ** Cheilolejeunea ascensionis (Hook.f. et Taylor) Grolle, Haussknechtia 4: 45, 1988 (Grolle 1988a). Bas.: Jungermannia ascensionis Hook.f. et Taylor, London J. Bot. 4: 91, 1845 (Hooker and Taylor 1845). *** Cheilolejeunea asperilora (Spruce) Gradst. et Ilk.-Borg., Mem. New York Bot. Gard. 76 (4): 62, 2009 (Gradstein and Ilkiu-Borges 2009). Bas.: Lejeunea asperilora Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 183, 1884 (Spruce 1884). ** Cheilolejeunea australis Solari, Comun. Mus. Argent. Ci. Nat. “Bernardino Rivadavia,” Ci. Bot. 2 (11): 70, 1981 (Solari 1981). ** Cheilolejeunea baumannii Hürl., Bauhinia 11 (3): 160, 1995 (Hürlimann 1995). *** Cheilolejeunea beyrichii (Lindenb.) M.E.Reiner, Nova Hedwigia 83 (3/4): 474, 2006 (Reiner-Drehwald 2006). Bas.: Lejeunea beyrichii Lindenb., Syn. Hepat. 3: 371, 1845 (Gottsche et al. 1845b). ** Cheilolejeunea boninensis Mizut., J. Hattori Bot. Lab. 51: 153, 1982 (Mizutani 1982). 239 Cheilolejeunea is here organized into subgenera that are maintained for practical purposes. hey may not be natural lineages. An improved infrageneric classiication based on molecular analysis is in preparation (Ye et al. 2015). he genus includes Euosmolejeunea, Strepsilejeunea and Trachylejeunea, but several taxa have neither been transferred nor synonymized. hey are listed under “Names in genera not currently accepted” below. 306 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Cheilolejeunea caducifolia (Gradst. et Schäf.-Verw.) W.Ye et R.L.Zhu, J. Bryol. 32 (4): 280, 2010 (Ye and Zhu 2010). Bas.: Leucolejeunea caducifolia Gradst. et Schäf.-Verw., J. Hattori Bot. Lab. 74: 64, 1993 (Gradstein et al. 1993). ** Cheilolejeunea camerunensis S.W.Arnell, Svensk Bot. Tidskr. 52 (1): 63, 1958 (Arnell 1958a). ** Cheilolejeunea celebensis (Steph.) Mizut., J. Hattori Bot. Lab. 36: 157, 1972 [1973] (Mizutani 1972a). Bas.: Trachylejeunea celebensis Steph., Sp. Hepat. (Stephani) 5: 312, 1913 (Stephani 1913a). ** Cheilolejeunea chenii R.L.Zhu et M.L.So, Taxon 48 (4): 663, 1999 (Zhu et al. 1999). Nom. nov. pro Neurolejeunea fukiensis P.C.Chen et P.C.Wu, Acta Phytotax. Sin. 9 (3): 227, 1964 (Chen and Wu 1964). *** Cheilolejeunea clypeata (Schwein.) W.Ye et R.L.Zhu, J. Bryol. 32 (4): 280, 2010 (Ye and Zhu 2010). Bas.: Jungermannia clypeata Schwein., Spec. Fl. Amer. Crypt.: 12, 1821 (Schweinitz 1821). ** Cheilolejeunea compressa (Herzog) Grolle, Bryophyt. Biblioth. 48: 38, 1995 (Grolle 1995). Bas.: Strepsilejeunea compressa Herzog, Bot. Not. 100 (4): 325, 1947 (Herzog 1947). *** Cheilolejeunea conchifolia (A.Evans) W.Ye et R.L.Zhu, J. Bryol. 32 (4): 280, 2010 (Ye and Zhu 2010). Bas.: Archilejeunea conchifolia A.Evans, Mem. Torrey Bot. Club 8 (2): 128, 1902 (Evans 1902a). *** Cheilolejeunea cordigera (Steph.) Grolle, J. Bryol. 9 (4): 530, 1977 [1978] (Grolle 1977a). Bas.: Hygrolejeunea cordigera Steph., Hedwigia 35 (3): 100, 1896 (Stephani 1896b). ** Cheilolejeunea coronalis (Gottsche) R.M.Schust., Phytologia 45 (5): 431, 1980 (Schuster 1980b). Bas.: Lejeunea coronalis Gottsche, Syn. Hepat. 3: 361, 1845 (Gottsche et al. 1845b). * Cheilolejeunea curvatilobula (Herzog) Grolle, J. Bryol. 21 (1): 41, 1999 (Grolle and Reiner-Drehwald 1999). Bas.: Harpalejeunea curvatilobula Herzog, Feddes Repert. Spec. Nov. Regni Veg. 57 (1/2): 184, 1955 (Herzog 1955). *** Cheilolejeunea decursiva (Sande Lac.) R.M.Schust., Beih. Nova Hedwigia 9: 112, 1963 (Schuster 1963a). Bas.: Lejeunea decursiva Sande Lac., Ned. Kruidk. Arch. 3: 522, 1855 (Sande Lacoste 1855). *** Cheilolejeunea discoidea (Lehm. et Lindenb.) R.M.Schust. et Kachroo, J. Linn. Soc., Bot. 56 (368): 509, 1961 (Kachroo and Schuster 1961). Bas.: Jungermannia discoidea Lehm. et Lindenb., Nov. Stirp. Pug. 6: 47, 1834 (Lehmann 1834). ** Cheilolejeunea diversifolia Augier, Ann. Fac. Sci. Univ. Féd. Cameroun 11: 66, 1972 (Augier 1972). ** Cheilolejeunea ecarinata Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 54 (1/2): 11, 1984 (Vanden Berghen 1984b). ** Cheilolejeunea erostrata R.M.Schust., Phytologia 39 (6): 427, 1978 (Schuster 1978b). *** Cheilolejeunea exinnovata E.W.Jones, J. Bryol. 12 (1): 37, 1982 (Jones 1982). *** Cheilolejeunea fragrantissima (Spruce) R.M.Schust., Phytologia 45 (5): 431, 1980 (Schuster 1980b). Bas.: Lejeunea fragrantissima Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 243, 1884 (Spruce 1884). World checklist of hornworts and liverworts * * ** ** *** ** ** ** * ** *** ** ** ** ** ** ** *** 307 Cheilolejeunea fukiensis (P.C.Chen et P.C.Wu) Piippo, J. Hattori Bot. Lab. 68: 133, 1990 (Piippo 1990). Bas.: Euosmolejeunea fukiensis P.C.Chen et P.C.Wu, Acta Phytotax. Sin. 9 (3): 232, 1964 (Chen and Wu 1964). 240 Cheilolejeunea galliotii Steph., Sp. Hepat. (Stephani) 5: 656, 1914 (Stephani 1914b). Cheilolejeunea germanii (Besch. et Spruce) Grolle, Acta Bot. Fenn. 125: 64, 1984 (Grolle and Piippo 1984). Bas.: Lejeunea germanii Besch. et Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: clxxxvii, 1889 [1890] (Bescherelle and Spruce 1889). Cheilolejeunea ghatensis G.Asthana, S.C.Srivast. et A.K.Asthana, Lindbergia 20 (2/3): 132, 1995 [1996] (Asthana et al. 1995). Cheilolejeunea gradsteinii (Grolle et Piippo) W.Ye et R.L.Zhu, J. Bryol. 32 (4): 280, 2010 (Ye and Zhu 2010). Bas.: Leucolejeunea gradsteinii Grolle et Piippo, Ann. Bot. Fenn. 27 (2): 122, 1990 (Grolle and Piippo 1990). Cheilolejeunea grandibracteata Steph., Sp. Hepat. (Stephani) 5: 657, 1914 (Stephani 1914b). Cheilolejeunea hamlinii Grolle, Wiss. Z. Friedrich-Schiller-Univ. Jena, Math.-Naturwiss. Reihe 31 (2): 212, 1982 (Grolle 1982). Nom. nov. pro Strepsilejeunea curnowii Steph., Hedwigia 35 (3): 129, 1896 (Stephani 1896b). Cheilolejeunea hawaica Steph., Bull. Herb. Boissier 5 (10): 847, 1897 (Stephani 1897c). Cheilolejeunea herzogiana Steph., Biblioth. Bot. 87 (2): 267, 1916 (Stephani 1916a). Cheilolejeunea implexicaulis (Hook.f. et Taylor) R.M.Schust., J. Hattori Bot. Lab. 26: 245, 1963 (Schuster 1963b). Bas.: Jungermannia implexicaulis Hook.f. et Taylor, London J. Bot. 3: 397, 1844 (Hooker and Taylor 1844a). Cheilolejeunea insecta Grolle et Gradst., Taxon 50 (4): 1071, 2001 [2002] (Grolle et al. 2001). Cheilolejeunea insignis Jovet-Ast et Tixier, Rev. Bryol. Lichénol. 31 (1/2): 25, 1962 (Jovet-Ast and Tixier 1962). Cheilolejeunea intricata (Steph.) J.J.Engel, Bryologist 79 (4): 514, 1976 [1977] (Engel 1976b). Bas.: Harpalejeunea intricata Steph., Sp. Hepat. (Stephani) 5: 269, 1913 (Stephani 1913a). Cheilolejeunea invaginata R.M.Schust., Phytologia 39 (6): 427, 1978 (Schuster 1978b). Cheilolejeunea jamaicensis Steph., Hedwigia 34 (5): 241, 1895 (Stephani 1895b). Cheilolejeunea japonica (Horik.) W.Ye et R.L.Zhu, J. Bryol. 32 (4): 280, 2010 (Ye and Zhu 2010). Bas.: Archilejeunea japonica Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 1: 84, 1932 (Horikawa 1932a). Cheilolejeunea kitagawae W.Ye et R.L.Zhu, J. Bryol. 32 (4): 280, 2010 (Ye and Zhu 2010). Nom. nov. pro Leucolejeunea paroica N.Kitag., Acta Phytotax. Geobot. 18 (7): 190, 1960 (Kitagawa 1960a). Cheilolejeunea lacerata C.J.Bastos et Gradst., J. Bryol. 28 (2): 133, 2006 (Bastos and Gradstein 2006). 240 Cheilolejeunea fukiensis may be conspeciic with Lejeunea lava (Zhu and So 2001), but they did not see the type specimen. 308 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Cheilolejeunea larsenii Mizut., Dansk Bot. Ark. 27 (1): 95, 1969 (Hattori and Mizutani 1969). * Cheilolejeunea laurentii Steph., Sp. Hepat. (Stephani) 5: 647, 1914 (Stephani 1914b). 241 ** Cheilolejeunea leptophylla (Ångstr.) Steph., Sp. Hepat. (Stephani) 5: 657, 1914 (Stephani 1914b). Bas.: Lejeunea leptophylla Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 33 (7): 86, 1876 [1877] (Ångström 1876). * Cheilolejeunea longilora (Taylor) R.M.Schust., Phytologia 45 (5): 431, 1980 (Schuster 1980b). Bas.: Lejeunea longilora Taylor, London J. Bot. 5: 396, 1846 (Taylor 1846b). 242 ** Cheilolejeunea longispina (Herzog) R.M.Schust., Beih. Nova Hedwigia 9: 111, 1963 (Schuster 1963a). Bas.: Harpalejeunea longispina Herzog, Rev. Bryol. Lichénol. 23 (1/2): 61, 1954 (Herzog 1954). ** Cheilolejeunea loriana (Steph.) W.Ye et R.L.Zhu, J. Bryol. 32 (4): 280, 2010 (Ye and Zhu 2010). Bas.: Symbiezidium lorianum Steph., Sp. Hepat. (Stephani) 5: 106, 1912 (Stephani 1912c). ** Cheilolejeunea ludoviciae Steph., Sp. Hepat. (Stephani) 5: 668, 1914 (Stephani 1914b). ** Cheilolejeunea lurida (Lindenb.) Steph., Sp. Hepat. (Stephani) 5: 658, 1914 (Stephani 1914b). Bas.: Lejeunea lurida Lindenb., Syn. Hepat. 3: 379, 1845 (Gottsche et al. 1845b). ** Cheilolejeunea macroloba (Herzog) Grolle, Wiss. Z. Friedrich-Schiller-Univ. Jena, Math.-Naturwiss. Reihe 31 (2): 212, 1982 (Grolle 1982). Bas.: Strepsilejeunea macroloba Herzog, Nat. Hist. Juan Fernandez (Botany) 2 (5): 742, 1942 (Herzog 1942a). *** Cheilolejeunea malaccensis (G.Hofm.) Xiao L.He, Ann. Bot. Fenn. 33 (1): 59, 1996 (He 1996b). Bas.: Pycnolejeunea malaccensis G.Hofm., Ann. Bryol. 8: 118, 1935 (Hofman 1935). ** Cheilolejeunea mammifera R.M.Schust., Phytologia 45 (5): 429, 1980 (Schuster 1980b). *** Cheilolejeunea mariana (Gottsche) B.M.hiers et Gradst., Mem. New York Bot. Gard. 52: 75, 1989 (hiers and Gradstein 1989). Bas.: Lejeunea mariana Gottsche, Syn. Hepat. 3: 337, 1845 (Gottsche et al. 1845b). ** Cheilolejeunea mexicana Steph., Sp. Hepat. (Stephani) 6: 417, 1923 (Stephani 1923). * Cheilolejeunea micholitzii (Steph.) R.M.Schust. et Kachroo, J. Linn. Soc., Bot. 56 (368): 509, 1961 (Kachroo and Schuster 1961). Bas.: Pycnolejeunea micholitzii Steph., Sp. Hepat. (Stephani) 5: 627, 1914 (Stephani 1914b). 243 241 Cheilolejeunea laurentii is not a Cheilolejeunea species (Jones 1954b, Wigginton and Grolle 1996). We do not know where to refer it. 242 Cheilolejeunea longilora is possibly conspeciic with Cheilolejeunea trifaria (Gradstein and Costa 2003). 243 Cheilolejeunea micholitzii is conspeciic with Cheilolejeunea longidens in hiers (1992b), but accepted by Hürlimann (1995). World checklist of hornworts and liverworts 309 ** Cheilolejeunea microscypha (Hook.f. et Taylor) M.Wigginton, J. Bryol. 34 (4): 269, 2012 (Wigginton 2012). Bas.: Jungermannia microscypha Hook.f. et Taylor, London J. Bot. 4: 90, 1845 (Hooker and Taylor 1845). *** Cheilolejeunea mizutanii W.Ye et R.L.Zhu, J. Bryol. 32 (4): 281, 2010 (Ye and Zhu 2010). Nom. nov. pro Hygrolejeunea decurrens Steph., Hedwigia 35 (3): 101, 1896 (Stephani 1896b). *** Cheilolejeunea neblinensis Ilk.-Borg. et Gradst., Nova Hedwigia 87 (3/4): 522, 2008 (Ilkiu-Borges and Gradstein 2008). ** Cheilolejeunea nietneri (Steph.) Mizut., J. Hattori Bot. Lab. 37: 191, 1973 (Mizutani 1973). Bas.: Harpalejeunea nietneri Steph., Sp. Hepat. (Stephani) 6: 392, 1923 (Stephani 1923). *** Cheilolejeunea norisiae G.Dauphin et Gradst., J. Bryol. 25 (4): 259, 2003 (Dauphin and Gradstein 2003). ** Cheilolejeunea novaezelandiae R.M.Schust., Phytologia 56 (7): 459, 1985 (Schuster 1985c). *** Cheilolejeunea obcordata Herzog, Mitt. Inst. Allg. Bot. Hamburg 7 (3): 209, 1931 (Herzog 1931a). ** Cheilolejeunea obtruncata (Mont.) Solari, J. Hattori Bot. Lab. 54: 539, 1983 (Solari 1983a). Bas.: Lejeunea obtruncata Mont., Ann. Sci. Nat. Bot. (sér. 3) 4: 354, 1845 (Montagne 1845b). ** Cheilolejeunea obtusa (Herzog) Solari, J. Hattori Bot. Lab. 54: 537, 1983 (Solari 1983a). Bas.: Harpalejeunea obtusa Herzog, Rev. Bryol. Lichénol. 29 (3/4): 193, 1960 [1961] (Herzog 1960). ** Cheilolejeunea obtusifolia (Steph.) S.Hatt., J. Hattori Bot. Lab. 18: 116, 1957 (Hattori 1957a). Bas.: Harpalejeunea obtusifolia Steph., Sp. Hepat. (Stephani) 5: 265, 1913 (Stephani 1913a). ** Cheilolejeunea occlusa (Herzog) T.Kodama et N.Kitag., Bull. Osaka Mus. Nat. Hist. 28: 40, 1974 (Kitagawa and Kodama 1974). Bas.: Strepsilejeunea occlusa Herzog, Trans. Brit. Bryol. Soc. 1 (4): 320, 1950 (Herzog 1950a). ** Cheilolejeunea orientalis (Gottsche) Mizut., J. Hattori Bot. Lab. 35: 399, 1972 (Mizutani 1972b). Bas.: Lejeunea orientalis Gottsche, Syn. Hepat. 3: 371, 1845 (Gottsche et al. 1845b). ** Cheilolejeunea ovistipula Steph., Hedwigia 34 (5): 244, 1895 (Stephani 1895b). ** Cheilolejeunea panurensis (Spruce) Steph., Sp. Hepat. (Stephani) 5: 660, 1914 (Stephani 1914b). Bas.: Lejeunea panurensis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 255, 1884 (Spruce 1884). ** Cheilolejeunea papillata Solari, Comun. Mus. Argent. Ci. Nat. “Bernardino Rivadavia,” Ci. Bot. 2 (11): 72, 1981 (Solari 1981). ** Cheilolejeunea paroica Mizut., J. Hattori Bot. Lab. 46: 364, 1979 (Mizutani 1979b). ** Cheilolejeunea pirilora Schifn., Bot. Jahrb. Syst. 23 (5): 592, 1897 (Schifner 1897). Based on: Lejeunea pirilora Gottsche ex Pol., J. Bot. 15: 227, 1877 (Polakowski 1877), nom. inval. 310 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Cheilolejeunea polystachya (Spruce) Gradst. et Ilk.-Borg., Mem. New York Bot. Gard. 76 (4): 62, 2009 (Gradstein and Ilkiu-Borges 2009). Bas.: Lejeunea polystachya Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 250, 1884 (Spruce 1884). ** Cheilolejeunea renneri (G.Hofm.) Xiao L.He, Ann. Bot. Fenn. 33 (1): 62, 1996 (He 1996b). Bas.: Pycnolejeunea renneri G.Hofm., Ann. Bryol. 8: 117, 1935 (Hofman 1935). *** Cheilolejeunea rotundistipula (Lindenb. ex Lehm.) Malombe, Acta Bot. Hung. 51 (3/4): 322, 2009 (Malombe 2009). Bas.: Jungermannia rotundistipula Lindenb. ex Lehm., Linnaea 4: 360, 1829 (Lehmann 1829). ** Cheilolejeunea rufescens (Lindenb.) Grolle, Wiss. Z. Friedrich-Schiller-Univ. Jena, Math.-Naturwiss. Reihe 31 (2): 212, 1982 (Grolle 1982). Bas.: Lejeunea rufescens Lindenb., Syn. Hepat. 3: 366, 1845 (Gottsche et al. 1845b). ** Cheilolejeunea stenoschiza (Ångstr.) A.Evans, Trans. Connecticut Acad. Arts 10 (8): 436, 1900 (Evans 1900a). Bas.: Lejeunea stenoschiza Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 29 (4): 26, 1872 (Ångström 1872). ** Cheilolejeunea subcrenulata (Spruce) R.M.Schust., Phytologia 45 (5): 431, 1980 (Schuster 1980b). Bas.: Lejeunea subcrenulata Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 245, 1884 (Spruce 1884). * Cheilolejeunea suborbicularis (Herzog) H.A.Mill., Bonner et Bischl., Nova Hedwigia 4: 559, 1962 [1963] (Miller et al. 1962). Bas.: Microlejeunea suborbicularis Herzog, Memoranda Soc. Fauna Fl. Fennica 26: 65, 1950 [1951] (Herzog 1950b). *** Cheilolejeunea suprema (Grolle et Piippo) W.Ye et R.L.Zhu, J. Bryol. 32 (4): 281, 2010 (Ye and Zhu 2010). Bas.: Leucolejeunea suprema Grolle et Piippo, Ann. Bot. Fenn. 27 (2): 123, 1990 (Grolle and Piippo 1990). ** Cheilolejeunea surrepens (Mitt.) E.W.Jones, J. Bryol. 9 (1): 49, 1976 (Jones 1976). Bas.: Lejeunea surrepens Mitt., Philos. Trans. 168: 399, 1879 (Mitten 1879). *** Cheilolejeunea turgida (Mitt.) W.Ye et R.L.Zhu, J. Bryol. 32 (4): 281, 2010 (Ye and Zhu 2010). Bas.: Lejeunea turgida Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 110, 1860 [1861] (Mitten 1860c). * Cheilolejeunea ulugurica Malombe, Eb.Fisch. et Pócs, Acta Biol. Pl. Agr. 1: 24, 2010 [2011] (Malombe et al. 2010). 244 *** Cheilolejeunea unciloba (Lindenb.) Malombe, Acta Bot. Hung. 51 (3/4): 325, 2009 (Malombe 2009). Bas.: Lejeunea unciloba Lindenb., Syn. Hepat. 3: 331, 1845 (Gottsche et al. 1845b). ** Cheilolejeunea upoluensis S.W.Arnell, Svensk Bot. Tidskr. 50 (3): 516, 1956 (Arnell 1956a). ** Cheilolejeunea valenciae (Steph.) Xiao L.He, Ann. Bot. Fenn. 33 (1): 55, 1996 (He 1996a). Bas.: Pycnolejeunea valenciae Steph., Sp. Hepat. (Stephani) 5: 605, 1914 (Stephani 1914b). 244 Cheilolejeunea ulugurica is possibly conspeciic with Cheilolejeunea chenii (Ye et al. 2013a). World checklist of hornworts and liverworts 311 ** Cheilolejeunea virescens (Lehm. et Lindenb.) Steph., Sp. Hepat. (Stephani) 5: 662, 1914 (Stephani 1914b). Bas.: Lejeunea virescens Lehm. et Lindenb., Nov. Stirp. Pug. 7: 21, 1838 (Lehmann 1838). * Cheilolejeunea viridis Steph., Sp. Hepat. (Stephani) 5: 673, 1914 (Stephani 1914b). ** Cheilolejeunea warnstorii (Steph.) Solari, Darwiniana 20 (3/4): 387, 1976 (Solari 1976). Bas.: Strepsilejeunea warnstorii Steph., Hedwigia 35 (3): 131, 1896 (Stephani 1896b). * Cheilolejeunea wrightii Steph., Sp. Hepat. (Stephani) 5: 662, 1914 (Stephani 1914b). *** Cheilolejeunea xanthocarpa (Lehm. et Lindenb.) Malombe, Acta Bot. Hung. 51 (3/4): 326, 2009 (Malombe 2009). Bas.: Jungermannia xanthocarpa Lehm. et Lindenb., Nov. Stirp. Pug. 5: 8, 1833 (Lehmann 1833). ** Cheilolejeunea xanthophylla (Lindenb.) Steph., Sp. Hepat. (Stephani) 5: 663, 1914 (Stephani 1914b). Bas.: Lejeunea xanthophylla Lindenb., Syn. Hepat. 3: 370, 1845 (Gottsche et al. 1845b). ** subg. Euosmolejeunea (Spruce) Kachroo, Ceylon J. Sci., Biol. Sci. 8 (1): 6, 1968 (Kachroo 1968). Bas.: Lejeunea subg. Euosmolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 241, 1884 (Spruce 1884). ** Cheilolejeunea aciculifera R.M.Schust., Phytologia 45 (5): 428, 1980 (Schuster 1980b). ** Cheilolejeunea acutangula (Nees) Grolle, J. Hattori Bot. Lab. 45: 173, 1979 (Grolle 1979c). Bas.: Jungermannia acutangula Nees, Fl. Bras. (Martius) 1 (1): 357, 1833 (Nees 1833a). *** Cheilolejeunea asperrima (Steph.) Grolle, J. Hattori Bot. Lab. 58: 197, 1985 (Grolle 1985b). Bas.: Taxilejeunea asperrima Steph., Biblioth. Bot. 87 (2): 259, 1916 (Stephani 1916a). ** Cheilolejeunea birmensis (Steph.) Mizut., J. Hattori Bot. Lab. 27: 139, 1964 (Mizutani 1964a). Bas.: Strepsilejeunea birmensis Steph., Sp. Hepat. (Stephani) 5: 286, 1913 (Stephani 1913a). ** Cheilolejeunea campbelliensis (Steph.) R.M.Schust., J. Hattori Bot. Lab. 26: 245, 1963 (Schuster 1963b). Bas.: Strepsilejeunea campbelliensis Steph., Hedwigia 35 (3): 128, 1896 (Stephani 1896b). ** Cheilolejeunea cedercreutzii (H.Buch et Perss.) Grolle, Feddes Repert. 87 (3/4): 188, 1976 (Grolle 1976a). Bas.: Euosmolejeunea cedercreutzii H.Buch et Perss., Bryophyt. Azoren Madeira: 9, 1941 (Buch and Persson 1941). ** Cheilolejeunea choachina (Gottsche) Gradst., Mem. New York Bot. Gard. 84: 69, 1999 (Gradstein 1999). Bas.: Lejeunea choachina Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 156, 1864 (Gottsche 1864). *** Cheilolejeunea clausa (Nees et Mont.) R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 863, 1980 (Schuster 1980c). Bas.: Lejeunea clausa Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 14: 337, 1840 (Montagne 1840a). ** Cheilolejeunea clavata Mizut., J. Hattori Bot. Lab. 46: 361, 1979 (Mizutani 1979b). 312 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Cheilolejeunea comans (Spruce) R.M.Schust., Phytologia 45 (5): 431, 1980 (Schuster 1980b). Bas.: Lejeunea comans Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 246, 1884 (Spruce 1884). ** Cheilolejeunea convexa (S.W.Arnell) S.W.Arnell, Hepat. South Africa: 209, 1963 (Arnell 1963b). Bas.: Lejeunea convexa S.W.Arnell, Bot. Not. 106: 272, 1953 (Arnell 1953b). ** Cheilolejeunea cookiensis (Steph.) R.M.Schust. et Kachroo, J. Linn. Soc., Bot. 56 (368): 509, 1961 (Kachroo and Schuster 1961). Bas.: Pycnolejeunea cookiensis Steph., Sp. Hepat. (Stephani) 5: 617, 1914 (Stephani 1914b). ** Cheilolejeunea cordistipula (Steph.) Grolle ex E.W.Jones, J. Bryol. 13 (3): 395, 1985 (Jones 1985). Bas.: Strepsilejeunea cordistipula Steph., Sp. Hepat. (Stephani) 5: 276, 1913 (Stephani 1913a). ** Cheilolejeunea ischeri Malombe, Acta Bot. Hung. 51 (3/4): 319, 2009 (Malombe 2009). ** Cheilolejeunea gaoi R.L.Zhu, M.L.So et Grolle, Bryologist 103 (3): 499, 2000 (Zhu et al. 2000). ** Cheilolejeunea hallingii B.M.hiers, Brittonia 44 (2): 160, 1992 (hiers 1992a). *** Cheilolejeunea inlexa (Hampe) Grolle, J. Hattori Bot. Lab. 45: 174, 1979 (Grolle 1979c). Bas.: Lejeunea inlexa Hampe, Nov. Stirp. Pug. 7: 22, 1838 (Lehmann 1838). *** Cheilolejeunea intertexta (Lindenb.) Steph., Bull. Herb. Boissier 5 (2): 79, 1897 (Stephani 1897b). Bas.: Lejeunea intertexta Lindenb., Syn. Hepat. 3: 379, 1845 (Gottsche et al. 1845b). *** Cheilolejeunea krakakammae (Lindenb.) R.M.Schust., Beih. Nova Hedwigia 9: 112, 1963 (Schuster 1963a). Bas.: Lejeunea krakakammae Lindenb., Syn. Hepat. 3: 353, 1845 (Gottsche et al. 1845b). *** Cheilolejeunea laciniata D.F.Peralta et M.E.Reiner, Bryologist 116 (1): 54, 2013 (Peralta and Reiner-Drehwald 2013). *** Cheilolejeunea laevicalyx (J.B.Jack et Steph.) Grolle, J. Hattori Bot. Lab. 65: 403, 1988 (Grolle 1988c). Bas.: Strepsilejeunea laevicalyx J.B.Jack et Steph., Sp. Hepat. (Stephani) 5: 283, 1913 (Stephani 1913a). ** Cheilolejeunea laeviuscula (Mitt.) Steph., Sp. Hepat. (Stephani) 5: 668, 1914 (Stephani 1914b). Bas.: Lejeunea laeviuscula Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 114, 1860 [1861] (Mitten 1860c). *** Cheilolejeunea lindenbergii (Gottsche) Mizut., J. Hattori Bot. Lab. 33: 226, 1970 (Mizutani 1970). Bas.: Lejeunea lindenbergii Gottsche, Syn. Hepat. 3: 336, 1845 (Gottsche et al. 1845b). ** Cheilolejeunea mimosa (Hook.f. et Taylor) R.M.Schust., J. Hattori Bot. Lab. 26: 245, 1963 (Schuster 1963b). Bas.: Jungermannia mimosa Hook.f. et Taylor, London J. Bot. 3: 398, 1844 (Hooker and Taylor 1844a). ** Cheilolejeunea nana R.M.Schust., Phytologia 39 (6): 426, 1978 (Schuster 1978b). ** Cheilolejeunea ngongensis Malombe et Pócs, Acta Bot. Hung. 51 (3/4): 317, 2009 (Malombe 2009). ** Cheilolejeunea nipponica (S.Hatt.) S.Hatt., Misc. Bryol. Lichenol. 1 (14): 1, 1957 (Hattori 1957c). Bas.: Strepsilejeunea nipponica S.Hatt., Bull. Tokyo Sci. Mus. 11: 134, 1944 (Hattori 1944d). World checklist of hornworts and liverworts 313 *** Cheilolejeunea omphalogastria Pócs, Trop. Bryol. 9: 131, 1994 (Pócs 1994b). *** Cheilolejeunea oncophylla (Ångstr.) Grolle et M.E.Reiner, J. Bryol. 19 (4): 781, 1997 (Grolle and Reiner-Drehwald 1997). Bas.: Lejeunea oncophylla Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 33 (7): 86, 1876 [1877] (Ångström 1876). ** Cheilolejeunea ornata C.J.Bastos, J. Bryol. 33 (1): 86, 2011 (Bastos 2011). ** Cheilolejeunea osumiensis (S.Hatt.) Mizut., Misc. Bryol. Lichenol. 8 (7): 148, 1980 (Mizutani 1980). Bas.: Euosmolejeunea osumiensis S.Hatt., Bull. Tokyo Sci. Mus. 11: 105, 1944 (Hattori 1944d). ** Cheilolejeunea pluriplicata (Pearson) R.M.Schust., Phytologia 45 (5): 430, 1980 (Schuster 1980b). Bas.: Lejeunea pluriplicata Pearson, Forh. Vidensk.-Selsk. Kristiania 1887 (9): 5, 1887 (Pearson 1887b). ** Cheilolejeunea pocsii E.W.Jones, J. Bryol. 15 (1): 156, 1988 (Jones 1988). ** Cheilolejeunea polyantha A.Evans, Mem. Torrey Bot. Club 8 (2): 141, 1902 (Evans 1902a). ** Cheilolejeunea polyantha var. caduciloba R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 887, 1980 (Schuster 1980c). *** Cheilolejeunea revoluta (Herzog) Gradst. et Grolle, J. Hattori Bot. Lab. 74: 59, 1993 (Gradstein et al. 1993). Bas.: Pycnolejeunea revoluta Herzog, Feddes Repert. Spec. Nov. Regni Veg. 57 (1/2): 193, 1955 (Herzog 1955). *** Cheilolejeunea rigidula (Nees ex Mont.) R.M.Schust., Castanea 36 (2): 102, 1971 (Schuster 1971a). Bas.: Lejeunea rigidula Nees ex Mont., Ann. Sci. Nat. Bot. (sér. 2) 14: 336, 1840 (Montagne 1840a). ** Cheilolejeunea ruwenzorensis (S.W.Arnell) R.M.Schust., Beih. Nova Hedwigia 9: 112, 1963 (Schuster 1963a). Bas.: Euosmolejeunea ruwenzorensis S.W.Arnell, Ark. Bot. (n.ser.) 3 (16): 529, 1956 (Arnell 1956e). ** Cheilolejeunea ryukyuensis Mizut., J. Hattori Bot. Lab. 51: 162, 1982 (Mizutani 1982). ** Cheilolejeunea subopaca (Mitt.) Mizut., J. Hattori Bot. Lab. 26: 183, 1963 (Mizutani 1963). Bas.: Lejeunea subopaca Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 116, 1860 [1861] (Mitten 1860c). *** Cheilolejeunea trifaria (Reinw., Blume et Nees) Mizut., J. Hattori Bot. Lab. 27: 132, 1964 (Mizutani 1964b). Bas.: Jungermannia trifaria Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 226, 1824 [1825] (Reinwardt et al. 1824a). * Cheilolejeunea udarii G.Asthana, S.C.Srivast. et A.K.Asthana, Lindbergia 20 (2/3): 142, 1995 [1996] (Asthana et al. 1995). 245 ** Cheilolejeunea usambarana (Steph.) Grolle, J. Hattori Bot. Lab. 46: 344, 1979 (Grolle 1979d). Bas.: Strepsilejeunea usambarana Steph., Akad. Wiss. Wien, Math.-Naturwiss. Kl., Denkschr. 88: 731, 1913 (Stephani 1913b). ** Cheilolejeunea verrucosa Steph., Sp. Hepat. (Stephani) 5: 673, 1914 (Stephani 1914b). 245 Cheilolejeunea udarii is possibly conspeciic with Cheilolejeunea krakakammae (Zhu 2006a). 314 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** subg. Renilejeunea R.M.Schust., Beih. Nova Hedwigia 9: 112, 1963 (Schuster 1963a). *** Cheilolejeunea montagnei (Gottsche ex Mont.) R.M.Schust., Beih. Nova Hedwigia 9: 112, 1963 (Schuster 1963a). Bas.: Lejeunea montagnei Gottsche ex Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 261, 1843 (Montagne 1843). ** subg. Xenolejeunea Kachroo et R.M.Schust., J. Linn. Soc., Bot. 56 (368): 496, 1961 (Kachroo and Schuster 1961). ** Cheilolejeunea ceylanica (Gottsche) R.M.Schust. et Kachroo, J. Linn. Soc., Bot. 56 (368): 509, 1961 (Kachroo and Schuster 1961). Bas.: Lejeunea ceylanica Gottsche, Syn. Hepat. 3: 359, 1845 (Gottsche et al. 1845b). ** Cheilolejeunea eximia (Jovet-Ast et Tixier) R.L.Zhu et M.L.So, Beih. Nova Hedwigia 121: 114, 2001 (Zhu and So 2001). Bas.: Pycnolejeunea eximia Jovet-Ast et Tixier, Rev. Bryol. Lichénol. 31 (1/2): 31, 1962 (Jovet-Ast and Tixier 1962). *** Cheilolejeunea falsinervis (Sande Lac.) R.M.Schust. et Kachroo, J. Linn. Soc., Bot. 56 (368): 509, 1961 (Kachroo and Schuster 1961). Bas.: Lejeunea falsinervis Sande Lac., Ned. Kruidk. Arch. 3: 421, 1854 [1855] (Sande Lacoste 1854). ** Cheilolejeunea gardneri (Mitt.) Mizut., J. Hattori Bot. Lab. 26: 181, 1963 (Mizutani 1963). Bas.: Lejeunea gardneri Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 115, 1860 [1861] (Mitten 1860c). ** Cheilolejeunea gigantea (Steph.) R.M.Schust. et Kachroo, J. Linn. Soc., Bot. 56 (368): 509, 1961 (Kachroo and Schuster 1961). Bas.: Pycnolejeunea gigantea Steph., Hedwigia 35 (3): 125, 1896 (Stephani 1896b). ** Cheilolejeunea huerlimannii Tixier, Misc. Bryol. Lichenol. 9 (9): 184, 1983 (Tixier 1983b). ** Cheilolejeunea hyalomarginata R.L.Zhu et Frank Müll., Bryologist 115 (2): 218, 2012 (Zhu and Müller 2012). *** Cheilolejeunea incisa (Gottsche) R.M.Schust. et Kachroo, J. Linn. Soc., Bot. 56 (368): 509, 1961 (Kachroo and Schuster 1961). Bas.: Lejeunea incisa Gottsche, Syn. Hepat. 3: 360, 1845 (Gottsche et al. 1845b). ** Cheilolejeunea incisa var. teretilora B.M.hiers, J. Hattori Bot. Lab. 82: 321, 1997 (hiers 1997a). ** Cheilolejeunea longidens (Steph.) R.M.Schust. et Kachroo, J. Linn. Soc., Bot. 56 (368): 509, 1961 (Kachroo and Schuster 1961). Bas.: Pycnolejeunea longidens Steph., Sp. Hepat. (Stephani) 5: 634, 1914 (Stephani 1914b). ** Cheilolejeunea meyeniana (Nees, Lindenb. et Gottsche) R.M.Schust. et Kachroo, J. Linn. Soc., Bot. 56 (368): 509, 1961 (Kachroo and Schuster 1961). Bas.: Lejeunea meyeniana Nees, Lindenb. et Gottsche, Observ. bot.: 472, 1843 (Gottsche et al. 1843). ** Cheilolejeunea obtusilobula (S.Hatt.) S.Hatt., Misc. Bryol. Lichenol. 1 (14): 2, 1957 (Hattori 1957c). Bas.: Pycnolejeunea obtusilobula S.Hatt., J. Hattori Bot. Lab. 3: 44, 1948 [1950] (Hattori 1948a). ** Cheilolejeunea parvidens B.M.hiers, J. Hattori Bot. Lab. 82: 323, 1997 (hiers 1997a). World checklist of hornworts and liverworts 315 ** Cheilolejeunea streimannii Pócs et Ninh, Acta Bot. Hung. 47 (1/2): 162, 2005 (Pócs and Ninh 2005). *** Cheilolejeunea trapezia (Nees) Kachroo et R.M.Schust., J. Linn. Soc., Bot. 56 (368): 509, 1961 (Kachroo and Schuster 1961). Bas.: Jungermannia trapezia Nees, Enum. Pl. Crypt. Javae: 41, 1830 (Nees 1830). ** Cheilolejeunea ventricosa (Schifn. ex P.Syd.) Xiao L.He, Acta Bot. Fenn. 163: 60, 1999 (He 1999). Bas.: Pycnolejeunea ventricosa Schifn. ex P.Syd., Just’s Bot. Jahresber. 19: 246, 1894 (Sydow 1894). *** Cheilolejeunea vittata (Steph. ex G.Hofm.) R.M.Schust. et Kachroo, J. Linn. Soc., Bot. 56 (368): 509, 1961 (Kachroo and Schuster 1961). Bas.: Pycnolejeunea vittata Steph. ex G.Hofm., Ann. Bryol. 8: 115, 1935 (Hofman 1935). Incertae sedis ** Cheilolejeunea gottscheana C.J.Bastos, J. Bryol. 34 (4): 316, 2012 (Bastos 2012b). Nom. nov. pro Strepsilejeunea lindenbergii Steph., Hedwigia 35 (3): 130, 1896 (Stephani 1896b). * Cheilolejeunea heteroclada (Spruce) Schifn., Hepat. (Engl.-Prantl): 124, 1893 (Schifner 1893b). Bas.: Lejeunea heteroclada Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 256, 1884 (Spruce 1884). 246 ** Cheilolejeunea minutilobula Amakawa, J. Jap. Bot. 35 (12): 365, 1960 (Amakawa 1960a). ** Cheilolejeunea norrisii (Grolle) M.A.M.Renner, Bryologist 115 (4): 550, 2012 (Renner 2012). Bas.: Lejeunea norrisii Grolle, Wiss. Z. Friedrich-Schiller-Univ. Jena, Math.-Naturwiss. Reihe 31 (2): 215, 1982 (Grolle 1982). ** Cheilolejeunea oscilla M.A.M.Renner, Bryologist 115 (4): 551, 2012 (Renner 2012). ** Cheilolejeunea papulosa Schifn., Hepat. (Engl.-Prantl): 124, 1893 (Schifner 1893b). Nom. nov. pro Lejeunea papulosa Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 258, 1884 (Spruce 1884), nom. illeg. *** Cheilolejeunea riparia (Steph.) M.E.Reiner, Nova Hedwigia 95 (3/4): 467, 2012 (Reiner-Drehwald and Grolle 2012). Bas.: Potamolejeunea riparia Steph., Sp. Hepat. (Stephani) 5: 639, 1914 (Stephani 1914b). *** Cheilolejeunea rotalis (Hook.f. et Taylor) M.Wigginton, J. Bryol. 34 (4): 270, 2012 (Wigginton 2012). Bas.: Jungermannia rotalis Hook.f. et Taylor, London J. Bot. 4: 89, 1845 (Hooker and Taylor 1845). ** Cheilolejeunea tenerrima (Steph.) C.J.Bastos, J. Bryol. 34 (4): 317, 2012 (Bastos 2012b). Bas.: Strepsilejeunea tenerrima Steph., Sp. Hepat. (Stephani) 5: 286, 1913 (Stephani 1913a). 246 Cheilolejeunea heteroclada was considered conspeciic with Cheilolejeunea aneogyna by X.-L. He (in Gradstein and Costa 2003), but it was tentatively accepted by Reiner-Drehwald and Grolle (2012). Lejeunea heteroclada var. subandina Spruce may be a diferent taxon (cf. Reiner-Drehwald & Grolle 2012). 316 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Cheilolejeunea urubuensis (Zartman et I.L.Ackerman) R.L.Zhu et Y.M.Wei, Phytotaxa 152 (1): 50, 2013 (Wei et al. 2013). Bas.: Vitalianthus urubuensis Zartman et I.L.Ackerman, Bryologist 105 (2): 267, 2002 (Zartman and Ackerman 2002). * Cyrtolejeunea A.Evans, Bull. Torrey Bot. Club 30 (10): 553, 1903 (Evans 1903c). *** Cyrtolejeunea holostipa (Spruce) A.Evans, Bull. Torrey Bot. Club. 30 (10): 553, 1903 (Evans 1903c). Bas.: Lejeunea holostipa Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 171 (Spruce 1884). ** Cystolejeunea A.Evans, Bull. Torrey Bot. Club 33 (1): 16, 1906) (Evans 1906a) *** Cystolejeunea lineata (Lehm. et Lindenb.) A.Evans, Bull. Torrey Bot. Club. 33 (1): 17, 1906 (Evans 1906a). Bas.: Jungermannia lineata Lehm. et Lindenb., Nov. Stirp. Pug. 4: 53, 1832 (Lehmann 1832). * ** *** ** ** ** *** *** *** Omphalanthus Lindenb. et Nees, Syn. Hepat. 2: 303, 1845 (Gottsche et al. 1845a). 247 Omphalanthus baracoensis Mustelier, M.E.Reiner et Gradst., J. Bryol. 29 (2): 95, 2007 (Reiner-Drehwald et al. 2007). Omphalanthus iliformis (Sw.) Nees, Syn. Hepat. 2: 304, 1845 (Gottsche et al. 1845a). Bas.: Jungermannia iliformis Sw., Prodr. (Swartz): 144, 1788 (Swartz 1788). Omphalanthus iliformis var. platycoleus (Herzog) Gradst., Phytotaxa 76 (3): 46, 2013 (Gradstein 2013a). Bas.: Omphalanthus platycoleus Herzog, Feddes Repert. Spec. Nov. Regni Veg. 57 (1/2): 171, 1955 (Herzog 1955). Omphalanthus iliformis var. wallisii (Prantl) Gradst., Phytotaxa 76 (3): 46, 2013 (Gradstein 2013a). Bas.: Lejeunea wallisii Prantl, Hedwigia 31: xvii, 1892 (Prantl 1892). Omphalanthus huanucensis (Gottsche) Gradst., Beih. Nova Hedwigia 80: 109, 1985 (Gradstein and Buskes 1985). Bas.: Lejeunea huanucensis Gottsche, Syn. Hepat. 3: 335, 1845 (Gottsche et al. 1845b). Omphalanthus jackii (Prantl) Gradst., Proc. Kon. Ned. Akad. Wetensch. C 80: 410, 1977 (Gradstein et al. 1977). Bas.: Lejeunea jackii Prantl, Hedwigia 31: xvii, 1892 (Prantl 1892). Omphalanthus ovalis (Lindenb. et Gottsche) Gradst., Proc. Kon. Ned. Akad. Wetensch. C 80: 411, 1977 (Gradstein et al. 1977). Bas.: Lejeunea ovalis Lindenb. et Gottsche, Syn. Hepat. 5: 754, 1847 (Gottsche et al. 1847). Omphalanthus roccatii (Gola) R.M.Schust., Beih. Nova Hedwigia 9: 96, 1963 (Schuster 1963a). Bas.: Acrolejeunea roccatii Gola, Ann. Bot. (Rome) 6 (2): 275, 1907 (Gola 1907). 247 Omphalanthus is nested in Cheilolejeunea (Wilson et al. 2007, Ye et al. 2011, Gradstein 2013a). A formal transfer of the genus and its species to Cheilolejeunea is found in Ye et al. (2015). World checklist of hornworts and liverworts *** 317 subtrib. Cololejeuneinae Gradst. ** Aphanotropis Herzog, Trans. Brit. Bryol. Soc. 2 (1): 63, 1952 (Herzog 1952b). *** Aphanotropis saxicola Herzog, Trans. Brit. Bryol. Soc. 2 (1): 63, 1952 (Herzog 1952b). ** Calatholejeunea K.I.Goebel, Ann. Jard. Bot. Buitenzorg 39: 8, 1928 (Goebel 1928). *** Calatholejeunea lamii Mizut., J. Hattori Bot. Lab. 56: 334, 1984 (Mizutani 1984a). *** Calatholejeunea paradoxa (Schifn.) K.I.Goebel, Ann. Jard. Bot. Buitenzorg 39: 8, 1928 (Goebel 1928). Bas.: Lejeunea paradoxa Schifn., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 60 (2): 243, 1893 (Schifner 1893a). *** Cololejeunea (Spruce) Steph., Hedwigia 30 (5): 208, 1891 (Stephani 1891a). Bas.: Lejeunea subg. Cololejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 291, 1884 (Spruce 1884). 248 ** Cololejeunea micrandroecia (Spruce) M.Menzel, Willdenowia 14: 492, 1984 [1985] (Menzel 1984). Bas.: Lejeunea micrandroecia Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 298, 1884 (Spruce 1884). * *** *** *** *** *** subg. Aphanolejeunea (A.Evans) Pócs, Phytotaxa 202 (1): 64, 2015 (Pócs et al. 2015c). Bas.: Aphanolejeunea A.Evans, Bull. Torrey Bot. Club 38 (6): 272, 1911 (Evans 1911). Cololejeunea berneckerae Pócs, Polish Bot. J. 54 (1): 4, 2009 (Pócs and Bernecker 2009). Nom. nov. pro Aphanolejeunea pocsii Bern.-Lück., Nova Hedwigia 66 (1/2): 168, 1998 (Bernecker-Lücking 1998). Cololejeunea cingens (Herzog) Bernecker et Pócs, Polish Bot. J. 54 (1): 4, 2009 (Pócs and Bernecker 2009). Bas.: Aphanolejeunea cingens Herzog, Svensk Bot. Tidskr. 46 (1): 104, 1952 (Herzog 1952e). Cololejeunea cornutissima (R.M.Schust.) Stotler et Crand.-Stotl., Bryologist 80 (3): 411, 1977 (Stotler and Crandall-Stotler 1977). Bas.: Aphanolejeunea cornutissima R.M.Schust., Bryologist 59 (3): 217, 1956 (Schuster 1956c). Cololejeunea costaricensis (Bern.-Lück.) Bernecker et Pócs, Polish Bot. J. 54 (1): 5, 2009 (Pócs and Bernecker 2009). Bas.: Aphanolejeunea costaricensis Bern.-Lück., Nova Hedwigia 66 (1/2): 164, 1998 (Bernecker-Lücking 1998). Cololejeunea cubensis Pócs, Polish Bot. J. 54 (1): 5, 2009 (Pócs and Bernecker 2009). Nom. nov. pro Aphanolejeunea evansii Herzog, Beih. Bot. Centralbl. 61B (3): 583, 1942 (Herzog 1942d). 248 Cololejeunea is here organized into subgenera that are not supported by the molecular study of Yu et al. (2013). hey are referred to following the traditional use (e.g. as in Pócs et al. 2014). Leptocolea and Physocolea also belong here, but a few taxa have neither been transferred nor synonymized. hey are listed in the “Names in genera not currently accepted” section below. 318 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Cololejeunea gracilis (Jovet-Ast) Pócs, Cryptog. Bryol. 29 (3): 233, 2008 (Dauphin et al. 2008). Bas.: Aphanolejeunea gracilis Jovet-Ast, Rev. Bryol. Lichénol. 16 (1/2): 21, 1947 [1948] (Jovet-Ast 1947b). ** Cololejeunea gracilis var. linearifolia (R.M.Schust.) Pócs, Acta Bot. Hung. 56 (1/2): 189, 2014 (Pócs et al. 2014). Bas.: Aphanolejeunea gracilis var. linearifolia R.M.Schust., Phytologia 45 (5): 434, 1980 (Schuster 1980b). *** Cololejeunea grossepapillosa (Horik.) N.Kitag., Hikobia, Suppl. 1: 68, 1981 (Kitagawa 1981a). Bas.: Aphanolejeunea grossepapillosa Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 1: 92, 1932 (Horikawa 1932a). ** Cololejeunea iwatsukiana (Pócs) Pócs, Polish Bot. J. 54 (1): 6, 2009 (Pócs and Bernecker 2009). Bas.: Aphanolejeunea iwatsukiana Pócs, Hikobia 11: 457, 1994 (Pócs 1994d). *** Cololejeunea jovetastiana (Pócs) Pócs, Polish Bot. J. 54 (1): 6, 2009 (Pócs and Bernecker 2009). Bas.: Aphanolejeunea jovetastiana Pócs, Cryptog. Bryol. Lichénol. 5 (3): 251, 1984 (Pócs 1984a). ** Cololejeunea lisowskii (Pócs) Pócs, Polish Bot. J. 54 (1): 6, 2009 (Pócs and Bernecker 2009). Bas.: Aphanolejeunea lisowskii Pócs, Cryptog. Bryol. Lichénol. 5 (3): 259, 1984 (Pócs 1984a). *** Cololejeunea madeirensis Schifn., Hedwigia 41 (5): 279, 1902 (Schifner 1902). *** Cololejeunea microscopica (Taylor) Schifn., Hepat. (Engl.-Prantl): 122, 1893 (Schifner 1893b). Bas.: Jungermannia microscopica Taylor, Mackay, Fl. Hibern. 2: 59, 1836 (Taylor 1836a). *** Cololejeunea microscopica var. africana (Pócs) Pócs et Bernecker, Cryptog. Bryol. 29 (3): 234, 2008 (Dauphin et al. 2008). Bas.: Aphanolejeunea exigua var. africana Pócs, Cryptog. Bryol. Lichénol. 5 (3): 247, 1984 (Pócs 1984a). *** Cololejeunea microscopica var. exigua (A.Evans) Pócs, Mem. New York Bot. Gard. 76 (4): 73, 2009 (Gradstein and Ilkiu-Borges 2009). Bas.: Aphanolejeunea exigua A.Evans, Bull. Torrey Bot. Club 38 (6): 273, 1911 (Evans 1911). *** Cololejeunea minuscula Pócs, Polish Bot. J. 54 (1): 7, 2009 (Pócs and Bernecker 2009). Nom. nov. pro Aphanolejeunea minuta R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 1310, 1980 (Schuster 1980c). ** Cololejeunea moramangae Tixier, Bull. Acad. Malgache (n.ser) 55 (1/2): 241, 1977 [1979] (Tixier 1977a). *** Cololejeunea norrisii (Pócs) Pócs, Polish Bot. J. 54 (1): 7, 2009 (Pócs and Bernecker 2009). Bas.: Aphanolejeunea norrisii Pócs, Acta Bot. Fenn. 165: 95, 1999 (Pócs and Piippo 1999). *** Cololejeunea papillosa (K.I.Goebel) Mizut., J. Hattori Bot. Lab. 29: 156, 1966 (Mizutani 1966). Bas.: Physocolea papillosa K.I.Goebel, Ann. Jard. Bot. Buitenzorg 39: 41, 1928 (Goebel 1928). *** Cololejeunea sicifolia (Gottsche ex A.Evans) Pócs et Bernecker, Polish Bot. J. 54 (1): 8, 2009 (Pócs and Bernecker 2009). Bas.: Aphanolejeunea sicifolia Gottsche ex A.Evans, Bull. Torrey Bot. Club 38 (6): 277, 1911 (Evans 1911). World checklist of hornworts and liverworts 319 *** Cololejeunea sicifolia subsp. jamaicensis (R.M.Schust.) Bernecker et Pócs, Polish Bot. J. 54 (1): 8, 2009 (Pócs and Bernecker 2009). Bas.: Aphanolejeunea jamaicensis R.M.Schust., Phytologia 45 (5): 434, 1980 (Schuster 1980b). *** Cololejeunea sintenisii (Steph.) Pócs, Cryptog. Bryol. 29 (3): 235, 2008 (Dauphin et al. 2008). Bas.: Aphanolejeunea sintenisii Steph., Sp. Hepat. (Stephani) 5: 861, 1916 (Stephani 1916b). *** Cololejeunea subsphaeroidea (R.M.Schust.) Pócs, Cryptog. Bryol. 29 (3): 235, 2008 (Dauphin et al. 2008). Bas.: Aphanolejeunea subsphaeroidea R.M.Schust., Phytologia 39 (6): 431, 1978 (Schuster 1978b). ** Cololejeunea thiersiae (Pócs) Pócs, Polish Bot. J. 54 (1): 9, 2009 (Pócs and Bernecker 2009). Bas.: Aphanolejeunea thiersiae Pócs, Hikobia 11: 459, 1994 (Pócs 1994d). ** Cololejeunea veillonii Tixier, Nova Hedwigia 31: 757, 1979 (Tixier 1979b). *** Cololejeunea winkleri (M.I.Morales et A.Lücking) Pócs, Mem. New York Bot. Gard. 76 (4): 78, 2009 (Gradstein and Ilkiu-Borges 2009). Bas.: Aphanolejeunea winkleri M.I.Morales et A.Lücking, Nova Hedwigia 60 (1/2): 120, 1995 (Morales and Lücking 1995). * ** ** ** ** subg. Austrocololejeunea Tixier, Nova Hedwigia 31: 776, 1979 (Tixier 1979b). Cololejeunea australis Tixier, Nova Hedwigia 31: 781, 1979 (Tixier 1979b). Cololejeunea caledonica Gottsche, Hedwigia 34 (5): 246, 1895 (Stephani 1895b). Cololejeunea sophiana Tixier, Bot. Not. 128: 428, 1975 [1976] (Tixier 1975a). Cololejeunea virotana Tixier, Nova Hedwigia 31: 777, 1979 (Tixier 1979b). * subg. Chlorocolea R.M.Schust., Beih. Nova Hedwigia 9: 178, 1963 (Schuster 1963a). Cololejeunea ceratilobula (P.C.Chen) R.M.Schust., Beih. Nova Hedwigia 9: 179, 1963 (Schuster 1963a). Bas.: Leptocolea ceratilobula P.C.Chen, Feddes Repert. Spec. Nov. Regni Veg. 58: 49, 1955 (Chen 1955). Cololejeunea desciscens Steph., Hedwigia 34 (5): 248, 1895 (Stephani 1895b). Cololejeunea linopteroides H.Rob., Bryologist 67 (4): 457, 1964 (Robinson 1964). Cololejeunea rotundilobula (P.C.Wu et P.J.Lin) Piippo, J. Hattori Bot. Lab. 68: 134, 1990 (Piippo 1990). Bas.: Pedinolejeunea rotundilobula P.C.Wu et P.J.Lin, Acta Phytotax. Sin. 16 (2): 69, 1978 (Wu and Lin 1978). 249 Cololejeunea sigmoidea Jovet-Ast et Tixier, Rev. Bryol. Lichénol. 31 (1/2): 27, 1962 (Jovet-Ast and Tixier 1962). Cololejeunea sigmoidea var. dubia Tixier, Bryophyt. Biblioth. 27: 114, 1985 (Tixier 1985a). Cololejeunea standleyi Herzog, Rev. Bryol. Lichénol. 20 (1/2): 172, 1951 [1952] (Herzog 1951a). *** *** *** * *** ** *** 249 Cololejeunea rotundilobula is possibly conspeciic with Cololejeunea sigmoidea (Pócs et al. 2013). 320 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Cololejeunea stylilobula Tixier, Phytotaxa 202 (1): 65, 2015 (Pócs et al. 2015c). Based on: Cololejeunea stylilobula Tixier, Bryophyt. Biblioth. 27: 119, 1985 (Tixier 1985a), nom. inval. *** Cololejeunea zangii R.L.Zhu et M.L.So, Syst. Bot. 24 (4): 501, 1999 [2000] (Zhu and So 1999b). *** subg. Chlorolejeunea Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 81, 1953 (Benedix 1953). 250 ** Cololejeunea lacinulata Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 82, 1953 (Benedix 1953). *** Cololejeunea madothecoides (Steph.) Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 81, 1953 (Benedix 1953). Bas.: Physocolea madothecoides Steph., Sp. Hepat. (Stephani) 5: 898, 1916 (Stephani 1916b). ** Cololejeunea ombrophila Tixier, Nat. Hist. Bull. Siam Soc. 23 (4): 550, 1970 (Tixier 1970a). *** Cololejeunea stotleriana Gradst., Ilk.-Borg. et Vanderp., Bryologist 114 (1): 13, 2011 (Gradstein et al. 2011). *** subg. Chondriolejeunea Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 75, 1953 (Benedix 1953). 251 *** Cololejeunea chinii Tixier, Nat. Hist. Bull. Siam Soc. 24 (3/4): 445, 1973 (Tixier 1973b). *** Cololejeunea pseudostipulata Schifn. ex P.Syd., Just’s Bot. Jahresber. 19: 246, 1894 (Sydow 1894). Based on: Cololejeunea pseudostipulata Schifn., Leberm., Forschungsr. Gazelle 4 (4): 33, 1890 (Schifner 1890), nom. inval. *** Cololejeunea shimizui N.Kitag., Acta Phytotax. Geobot. 23 (5/6): 185, 1969 (Kitagawa 1969b). *** Cololejeunea shimizui var. phangngana N.Kitag., Acta Phytotax. Geobot. 23 (5/6): 187, 1969 (Kitagawa 1969b). *** subg. Cololejeunea, 252 ** Cololejeunea albodentata P.C.Chen et P.C.Wu, Acta Phytotax. Sin. 9 (3): 252, 1964 (Chen and Wu 1964). ** Cololejeunea armata Tixier, Gard. Bull. Singapore 26 (1): 149, 1972 (Tixier 1972b). *** Cololejeunea bhutanica Grolle et Mizut., J. Bryol. 15 (2): 281, 1989 (Grolle 1989e). 250 Cololejeunea subg. Chlorolejeunea forms a separate clade in Yu et al. (2013) with three sequenced species (Cololejeunea ceratilobula, Cololejeunea madothecoides (the type) and Cololejeunea raduliloba). 251 Cololejeunea subg. Chondriolejeunea is supported by Yu et al. (2013) who included Cololejeunea chinii, Cololejeunea pseudostipulata (the type, not sequenced) and Cololejeunea shimizui. 252 Subg. Cololejeunea includes subg. Aphanolejeunea in the phylogeny by Yu et al. (2013). World checklist of hornworts and liverworts 321 ** Cololejeunea biddlecomiae (Austin) A.Evans, Mem. Torrey Bot. Club 8 (2): 168, 1902 (Evans 1902a). Bas.: Lejeunea biddlecomiae Austin, List. Canad. Hepat.: 5, 1890 (Pearson 1890). * Cololejeunea caihuaella But et P.C.Wu, Hepat. Fl. Hong Kong: 132, 2009 (Wu and But 2009). *** Cololejeunea calcarea (Lib.) Steph., Bot. Gaz. 17 (6): 171, 1892 (Stephani 1892f). Bas.: Lejeunea calcarea Lib., Ann. Gen. Sci. Phys. 6: 373, 1820 (Libert 1820). ** Cololejeunea capuronii Tixier, Bull. Acad. Malgache (n.ser) 55 (1/2): 241, 1977 [1979] (Tixier 1977a). ** Cololejeunea dinghuiana R.L.Zhu et Y.F.Wang, J. E. China Norm. Univ., Nat. Sci. Ed. 2: 91, 1992 (Zhu and Wang 1992). ** Cololejeunea dolichodonta Tixier, Bryophyt. Biblioth. 27: 215, 1985 (Tixier 1985a). *** Cololejeunea dozyana (Sande Lac.) Schifn., Hedwigia 39 (4): 199, 1900 (Schifner 1900b). Bas.: Lejeunea dozyana Sande Lac., Ned. Kruidk. Arch. 3: 522, 1855 (Sande Lacoste 1855). *** Cololejeunea elegans Steph., Hedwigia 30 (5): 208, 1891 (Stephani 1891a). ** Cololejeunea falcidentata R.M.Schust., Nova Hedwigia 15: 507, 1968 (Schuster 1968b). ** Cololejeunea ilidens Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 49, 1953 (Benedix 1953). * Cololejeunea frahmii Tixier, Trop. Bryol. 11: 63, 1995 (Tixier 1995b). 253 *** Cololejeunea grolleana Pócs, J. Hattori Bot. Lab. 48: 312, 1980 (Pócs 1980b). *** Cololejeunea haskarliana (Lehm.) Schifn., Consp. Hepat. Arch. Ind.: 244, 1898 (Schifner 1898b). Bas.: Lejeunea haskarliana Lehm., Nov. Stirp. Pug. 8: 26, 1844 (Lehmann 1844). *** Cololejeunea hyalina G.Asthana et S.C.Srivast., Bryophyt. Biblioth. 60: 25, 2003 (Asthana and Srivastava 2003). * Cololejeunea kahuziensis Tixier, Trop. Bryol. 11: 63, 1995 (Tixier 1995b). 254 *** Cololejeunea karnatakensis G.Asthana et S.C.Srivast., Bryophyt. Biblioth. 60: 26, 2003 (Asthana and Srivastava 2003). ** Cololejeunea kodamae Kamim., Feddes Repert. Spec. Nov. Regni Veg. 58: 55, 1955 (Kamimura 1955). *** Cololejeunea kolombangarae Pócs, Acta Bryolichenol. Asiat. 4: 67, 2011 (Pócs and Piippo 2011). ** Cololejeunea kolombangarae subsp. sepikensis Pócs, Acta Bryolichenol. Asiat. 4: 68, 2011 (Pócs and Piippo 2011). *** Cololejeunea konratii Pócs, Acta Bot. Hung. 54 (1/2): 156, 2012 (Pócs 2012b). *** Cololejeunea kuciana Pócs et Schäf.-Verw., Polish Bot. J. 57 (1): 51, 2012 (Pócs and Schäfer-Verwimp 2012). *** Cololejeunea longiana Grolle et Mizut., J. Bryol. 15 (2): 284, 1989 (Grolle 1989e). 253 Cololejeunea frahmii is probably only a depauperate form of Cololejeunea elegans or Cololejeunea zenkeri. 254 Cololejeunea kahuziensis is probably only a depauperate form of Cololejeunea elegans or Cololejeunea zenkeri. 322 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Cololejeunea macounii (Spruce) A.Evans, Mem. Torrey Bot. Club 8 (2): 171, 1902 (Evans 1902a). Bas.: Lejeunea macounii Spruce, Bull. Torrey Bot. Club 17 (10): 259, 1890 (Underwood 1890). ** Cololejeunea magillii Pócs, J. Hattori Bot. Lab. 74: 49, 1993 (Pócs 1993). *** Cololejeunea malanjae Steph., Bot. Jahrb. Syst. 30 (2): 261, 1901 (Stephani 1901g). ** Cololejeunea mamillata (Ångstr.) E.A.Hodgs., Trans. Roy. Soc. New Zealand, Bot. 3 (11): 184, 1967 (Hodgson 1967). Bas.: Lejeunea mamillata Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 29 (4): 14, 1872 (Ångström 1872). ** Cololejeunea mizutaniana Udar et G.Srivast., Misc. Bryol. Lichenol. 9 (7): 138, 1983 (Udar and Srivastava 1983). *** Cololejeunea mocambiquensis S.W.Arnell, Mitt. hüring. Bot. Ges. 1 (1): 7, 1955 [1956] (Arnell 1955a). ** Cololejeunea nanhutashanensis J.D.Yang et S.H.Lin, Phytotaxa 177 (1): 56, 2014 (Yang and Lin 2014). *** Cololejeunea nilgiriensis G.Asthana et S.C.Srivast., Bryophyt. Biblioth. 60: 27, 2003 (Asthana and Srivastava 2003). *** Cololejeunea ornata A.Evans, Bryologist 41 (4): 73, 1938 (Evans 1938c). ** Cololejeunea parva Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 47 (1/2): 239, 1977 (Vanden Berghen 1977). *** Cololejeunea planilora Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 60, 1953 (Benedix 1953). *** Cololejeunea platyneura (Spruce) A.Evans, Mem. Torrey Bot. Club 8 (2): 172, 1902 (Evans 1902a). Bas.: Lejeunea platyneura Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 299, 1884 (Spruce 1884). *** Cololejeunea pluridentata P.C.Wu et J.S.Lou, Acta Phytotax. Sin. 16 (4): 105, 1978 (Wu and Lou 1978). ** Cololejeunea pretiosa Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 62, 1953 (Benedix 1953). *** Cololejeunea pseudocristallina P.C.Chen et P.C.Wu, Acta Phytotax. Sin. 9 (3): 257, 1964 (Chen and Wu 1964). ** Cololejeunea pseudoplagiophylla P.C.Wu et J.S.Lou, Acta Phytotax. Sin. 16 (4): 106, 1978 (Wu and Lou 1978). ** Cololejeunea pseudoschmidtii Tixier, Gard. Bull. Singapore 26 (1): 145, 1972 (Tixier 1972b). ** Cololejeunea ramromensis Pócs, Biodiv., biogeogr., nat. conserv. Wallacea N. Guinea II: 113, 2014 (Chantanaorrapint and Pócs 2014). *** Cololejeunea rosellata Mizut., J. Hattori Bot. Lab. 29: 159, 1966 (Mizutani 1966). *** Cololejeunea rossettiana (C.Massal.) Schifn., Hepat. (Engl.-Prantl): 122, 1893 (Schifner 1893b). Bas.: Lejeunea rossettiana C.Massal., Nuovo Giorn. Bot. Ital. 21 (3): 487, 1889 (Massalongo 1889). *** Cololejeunea runssorensis (Steph.) Pócs, Acta Bot. Acad. Sci. Hung. 21 (3/4): 371, 1975 (Pócs 1975). Bas.: Aphanolejeunea runssorensis Steph., Sp. Hepat. (Stephani) 5: 858, 1916 (Stephani 1916b). World checklist of hornworts and liverworts 323 ** Cololejeunea schaeferi Grolle, J. Bryol. 13 (4): 488, 1986 (Grolle 1986b). *** Cololejeunea schmidtii Steph., Bot. Tidsskr. 24 (3): 278, 1902 (Stephani 1902b). ** Cololejeunea schmidtii var. acutepapillosa Pócs, Acta Bot. Hung. 54 (1/2): 165, 2012 (Pócs 2012b). ** Cololejeunea selaginellicola Tixier, Gard. Bull. Singapore 26 (1): 149, 1972 (Tixier 1972b). *** Cololejeunea serrata (Steph.) Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 53, 1953 (Benedix 1953). Bas.: Physocolea serrata Steph., Sp. Hepat. (Stephani) 5: 905, 1916 (Stephani 1916b). ** Cololejeunea shikokiana (Horik.) S.Hatt., Bull. Tokyo Sci. Mus. 11: 101, 1944 (Hattori 1944d). Bas.: Physocolea shikokiana Horik., Bot. Mag. (Tokyo) 46 (544): 182, 1932 (Horikawa 1932b). ** Cololejeunea spinosa (Horik.) Pandé et R.N.Misra, J. Indian Bot. Soc. 22 (2/4): 166, 1943 (Pandé and Misra 1943). Bas.: Physocolea spinosa Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 1: 70, 1931 (Horikawa 1931a). *** Cololejeunea stellaris Pócs, Acta Bryolichenol. Asiat. 4: 70, 2011 (Pócs and Piippo 2011). ** Cololejeunea subkodamae Mizut., J. Hattori Bot. Lab. 60: 448, 1986 (Mizutani 1986a). ** Cololejeunea tamasii Schäf.-Verw., Phytotaxa 60: 9, 2012 (Schäfer-Verwimp 2012). ** Cololejeunea tanneri Pócs, Acta Bot. Hung. 31 (1/4): 126, 1985 (Pócs 1985). *** Cololejeunea tenella Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 55, 1953 (Benedix 1953). ** Cololejeunea tenella var. dentiloba Onr., Bull. Jard. Bot. Natl. Belg. 59 (3/4): 436, 1989 (Onraedt 1989). ** Cololejeunea thiersiana Tixier, Cryptog. Bryol. Lichénol. 16 (3): 229, 1995 (Tixier 1995a). Based on: Cololejeunea thiersiana Tixier, Candollea 46 (2): 291, 1991 (Tixier 1991), nom. inval. ** Cololejeunea tuiwawana Pócs, Acta Bot. Hung. 54 (1/2): 165, 2012 (Pócs 2012b). *** Cololejeunea verrucosa Steph., Hedwigia 34 (5): 253, 1895 (Stephani 1895b). ** Cololejeunea verrucosa var. rectispina (Herzog) Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 55, 1953 (Benedix 1953). Bas.: Physocolea verrucosa var. rectispina Herzog, Mitt. Inst. Allg. Bot. Hamburg 7 (3): 214, 1931 (Herzog 1931a). *** Cololejeunea zenkeri (Steph.) E.W.Jones, Trans. Brit. Bryol. Soc. 2 (3): 420, 1954 (Jones 1954c). Bas.: Aphanolejeunea zenkeri Steph., Sp. Hepat. (Stephani) 5: 858, 1916 (Stephani 1916b). subg. Cryptolejeunea Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 77, 1953 (Benedix 1953). *** Cololejeunea angustilora (Steph.) Mizut., J. Hattori Bot. Lab. 28: 113, 1965 (Mizutani 1965). Bas.: Leptocolea angustilora Steph., Sp. Hepat. (Stephani) 5: 848, 1916 (Stephani 1916b). * 324 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Cololejeunea drepanolejeuneoides (Horik.) R.M.Schust., Beih. Nova Hedwigia 9: 174, 1963 (Schuster 1963a). Bas.: Boninoleptocolea drepanolejeuneoides Horik., Bot. Mag. (Tokyo) 50 (598): 558, 1936 (Horikawa 1936). 255 ** Cololejeunea hattoriana Mizut. et Pócs, Ann. Bot. Fenn. 31 (3): 188, 1994 (Pócs et al. 1994). Nom. nov. pro Campylolejeunea pusilla Mizut., J. Hattori Bot. Lab. 29: 154, 1966 (Mizutani 1966). *** Cololejeunea inlectens (Mitt.) Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 79, 1953 (Benedix 1953). Bas.: Lejeunea inlectens Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 117, 1860 [1861] (Mitten 1860c). ** Cololejeunea mouensis (Tixier) H.A.Mill., Phytologia 47 (4): 322, 1981 (Miller 1981). Bas.: Campylolejeunea mouensis Tixier, Nova Hedwigia 31: 727, 1979 (Tixier 1979b). *** Cololejeunea vesicaria (Sande Lac.) Schifn., Consp. Hepat. Arch. Ind.: 247, 1898 (Schifner 1898b). Bas.: Lejeunea vesicaria Sande Lac., Syn. hepat. jav.: 74, 1856 [1857] (Sande Lacoste 1856b). * *** ** * *** *** *** *** ** *** *** subg. Diaphanae R.M.Schust., J. Elisha Mitchell Sci. Soc. 72 (1): 103, 1956 (Schuster 1956b). Cololejeunea amphibola B.M.hiers, Beih. Nova Hedwigia 90: 130, 1988 (hiers 1988). Cololejeunea antillana Pócs, Polish Bot. J. 54 (1): 3, 2009 (Pócs and Bernecker 2009). Nom. nov. pro Aphanolejeunea longifolia Jovet-Ast, Rev. Bryol. Lichénol. 16 (1/2): 23, 1947 [1948] (Jovet-Ast 1947b). Cololejeunea augieri Tixier, Trop. Bryol. 11: 48, 1995 (Tixier 1995b). Cololejeunea azorica V.Allorge et Jovet-Ast, Mitt. hüring. Bot. Ges. 1 (2/3): 17, 1955 (Allorge and Jovet-Ast 1955). Cololejeunea camillii (Lehm.) A.Evans, Bryologist 15 (4): 59, 1912 (Evans 1912a). Bas.: Lejeunea camillii Lehm., Nov. Stirp. Pug. 10: 15, 1857 (Lehmann 1857). Cololejeunea contractiloba A.Evans, Amer. J. Bot. 5 (3): 131, 1918 (Evans 1918). Cololejeunea cornuta E.W.Jones, Trans. Brit. Bryol. Soc. 2 (3): 436, 1954 (Jones 1954c). Cololejeunea crenata (A.Evans) Pócs, Polish Bot. J. 54 (1): 5, 2009 (Pócs and Bernecker 2009). Bas.: Aphanolejeunea crenata A.Evans, Bull. Torrey Bot. Club 38 (6): 276, 1911 (Evans 1911). Cololejeunea diaphana A.Evans, Bull. Torrey Bot. Club 32 (4): 184, 1905 (Evans 1905b). Cololejeunea erostrata (Herzog) Bernecker et Pócs, Polish Bot. J. 54 (1): 5, 2009 (Pócs and Bernecker 2009). Bas.: Physocolea erostrata Herzog, Beih. Bot. Centralbl. 61B (3): 582, 1942 (Herzog 1942d). 255 Cololejeunea drepanolejeuneoides may based on its description be close to Cololejeunea angustilora and especially to Cololejeunea inlectens, forming a natural group. World checklist of hornworts and liverworts 325 ** Cololejeunea gradsteinii M.J.Lai et R.L.Zhu, Ann. Bot. Fenn. 45 (5): 334, 2008 (Lai et al. 2008). Nom. nov. pro Cololejeunea pusilla Tixier, Nat. Hist. Bull. Siam Soc. 23 (4): 552, 1970 (Tixier 1970a), nom. illeg. ** Cololejeunea koponenii (Pócs) Pócs, Polish Bot. J. 54 (1): 6, 2009 (Pócs and Bernecker 2009). Bas.: Aphanolejeunea koponenii Pócs, Acta Bot. Fenn. 165: 91, 1999 (Pócs and Piippo 1999). *** Cololejeunea lanceolata E.W.Jones, Trans. Brit. Bryol. Soc. 2 (3): 428, 1954 (Jones 1954c). *** Cololejeunea moralesiae (Bern.-Lück.) Bernecker et Pócs, Polish Bot. J. 54 (1): 7, 2009 (Pócs and Bernecker 2009). Bas.: Aphanolejeunea moralesiae Bern.-Lück., Nova Hedwigia 66 (1/2): 166, 1998 (Bernecker-Lücking 1998). *** Cololejeunea morobensis (Pócs) Pócs, Polish Bot. J. 54 (1): 7, 2009 (Pócs and Bernecker 2009). Bas.: Aphanolejeunea morobensis Pócs, Ann. Bot. Fenn. 31 (3): 180, 1994 (Pócs et al. 1994). *** Cololejeunea obtusifolia (E.W.Jones) Tixier, Trop. Bryol. 11: 42, 1995 (Tixier 1995b). Bas.: Cololejeunea pusilla var. obtusifolia E.W.Jones, Trans. Brit. Bryol. Soc. 2 (3): 427, 1954 (Jones 1954c). * Cololejeunea obtusifolia var. madecassa (Tixier) Pócs, Phytotaxa 202 (1): 64, 2015 (Pócs et al. 2015c). Bas.: Cololejeunea androphylla var. madecassa Tixier, Bull. Acad. Malgache (n.ser) 55 (1/2): 216, 1977 [1979] (Tixier 1977a). *** Cololejeunea papilliloba (Steph.) Steph., Hedwigia 34 (5): 250, 1895 (Stephani 1895b). Bas.: Lejeunea papilliloba Steph., Hedwigia 29 (2): 73, 1890 (Stephani 1890b). ** Cololejeunea papulosa R.M.Schust., Phytologia 45 (5): 433, 1980 (Schuster 1980b). *** Cololejeunea paucifolia (Spruce) Bernecker et Pócs, Polish Bot. J. 54 (1): 8, 2009 (Pócs and Bernecker 2009). Bas.: Lejeunea paucifolia Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: cxciv, 1889 [1890] (Spruce 1889). *** Cololejeunea peponiformis Mizut., J. Hattori Bot. Lab. 33: 258, 1970 (Mizutani 1970). ** Cololejeunea pseudocuspidata Tixier, Bauhinia 8 (4): 228, 1987 (Hürlimann 1987). *** Cololejeunea pterocolea Herzog, Svensk Bot. Tidskr. 46 (1): 103, 1952 (Herzog 1952e). *** Cololejeunea pusilla Steph., Hedwigia 34 (5): 251, 1895 (Stephani 1895b). ** Cololejeunea spathulata Jovet-Ast, Rev. Bryol. Lichénol. 29 (1/2): 36, 1960 (Jovet-Ast 1960). ** Cololejeunea taurifolia Inoue et H.A.Mill., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 8 (2): 152, 1965 (Inoue and Miller 1965). *** Cololejeunea wightii Steph., Hedwigia 34 (5): 253, 1895 (Stephani 1895b). *** Cololejeunea yelitzae Pócs et Bernecker, Acta Bot. Hung. 55 (3/4): 386, 2013 (Pócs and Bernecker 2013). subg. Leptocolea (Spruce) Schifn., Hepat. (Engl.-Prantl): 122, 1893 (Schifner 1893b). Bas.: Lejeunea sect. Leptocolea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 294, 1884 (Spruce 1884). ** Cololejeunea acuminata Mizut., J. Hattori Bot. Lab. 33: 261, 1970 (Mizutani 1970). * 326 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Cololejeunea aequabilis (Sande Lac.) Schifn., Consp. Hepat. Arch. Ind.: 242, 1898 (Schifner 1898b). Bas.: Lejeunea aequabilis Sande Lac., Ann. Mus. Bot. LugdunoBatavi 1: 310, 1864 (Sande Lacoste 1864). ** Cololejeunea altimontana Pócs, Acta Bryolichenol. Asiat. 4: 78, 2011 (Pócs and Piippo 2011). *** Cololejeunea amaniensis Pócs, Acta Bot. Hung. 31 (1/4): 120, 1985 (Pócs 1985). *** Cololejeunea angustibracteata Schifn. ex P.Syd., Just’s Bot. Jahresber. 19: 246, 1894 (Sydow 1894). Based on: Cololejeunea angustibracteata Schifn., Leberm., Forschungsr. Gazelle 4 (4): 34, 1890 (Schifner 1890), nom. inval. *** Cololejeunea apiculata (E.W.Jones) R.M.Schust., Beih. Nova Hedwigia 9: 174, 1963 (Schuster 1963a). Bas.: Leptocolea apiculata E.W.Jones, Trans. Brit. Bryol. Soc. 2 (3): 415, 1954 (Jones 1954c). * Cololejeunea arrectifolia (Mitt.) Steph., Bot. Jahrb. Syst. 23 (1/2, 3): 309, 1896 (Stephani 1896a). Bas.: Lejeunea arrectifolia Mitt., Fl. vit.: 415, 1871 [1873] (Mitten 1871). ** Cololejeunea aurantia (Tixier) houvenot, Cryptog. Bryol. 32 (4): 290, 2011 (houvenot et al. 2011). Bas.: Jovetastella aurantia Tixier, Cryptog. Bryol. Lichénol. 3 (1): 29, 1982 (Tixier 1982). ** Cololejeunea bandamiae Tixier, Bull. Acad. Malgache (n.ser) 55 (1/2): 220, 1977 [1979] (Tixier 1977a). ** Cololejeunea bebourensis Tixier, Bull. Acad. Malgache (n.ser) 55 (1/2): 229, 1977 [1979] (Tixier 1977a). ** Cololejeunea bergmansiana Tixier, Bull. Jard. Bot. Natl. Belg. 59 (3/4): 440, 1989 (Tixier 1989). ** Cololejeunea bidentula (Steph.) E.W.Jones, Trans. Brit. Bryol. Soc. 2 (3): 423, 1954 (Jones 1954c). Bas.: Physocolea bidentula Steph., Sp. Hepat. (Stephani) 5: 868, 1916 (Stephani 1916b). ** Cololejeunea bifalcata Pócs, Acta Bot. Hung. 54 (1/2): 146, 2012 (Pócs 2012b). *** Cololejeunea blepharophylla Pócs, Acta Bot. Hung. 54 (1/2): 149, 2012 (Pócs 2012b). ** Cololejeunea borhidiana Pócs, J. Hattori Bot. Lab. 48: 305, 1980 (Pócs 1980b). ** Cololejeunea bosseriana Tixier, Bull. Acad. Malgache (n.ser) 55 (1/2): 235, 1977 [1979] (Tixier 1977a). ** Cololejeunea calcarata E.W.Jones, Bull. Brit. Mus. (Nat. Hist.), Bot. 11 (3): 235, 1983 (Jones and Harrington 1983). * Cololejeunea camusii Tixier, Bull. Acad. Malgache (n.ser) 55 (1/2): 227, 1977 [1979] (Tixier 1977a). *** Cololejeunea ceatocarpa (Ångstr.) Steph., Bull. Herb. Boissier 5 (10): 842, 1897 (Stephani 1897c). Bas.: Lejeunea ceatocarpa Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 29 (4): 27, 1872 (Ångström 1872). *** Cololejeunea ceylanica Onr., Acta Bot. Acad. Sci. Hung. 25 (1/2): 107, 1979 (Onraedt 1979). World checklist of hornworts and liverworts * *** *** ** ** ** ** * ** ** * ** ** ** ** * *** 327 Cololejeunea chamlongiana Tixier, Ann. Fac. Sci. Univ. Phnom Penh 3: 181, 1970 (Tixier 1970b). 256 Cololejeunea chenii Tixier, Bryophyt. Biblioth. 27: 219, 1985 (Tixier 1985a). Nom. nov. pro Cololejeunea plagiophylla var. grossipapillosa P.C.Chen et P.C.Wu, Acta Phytotax. Sin. 9 (3): 254, 1964 (Chen and Wu 1964). Cololejeunea ciliata Pócs, Ann. Bot. Fenn. 31 (3): 182, 1994 (Pócs et al. 1994). Cololejeunea comptonii (Pearson) H.A.Mill., Phytologia 47 (4): 321, 1981 (Miller 1981). Bas.: Leptocolea comptonii Pearson, J. Linn. Soc., Bot. 46 (305): 40, 1922 (Pearson 1922b). Cololejeunea cookei A.Evans, Trans. Connecticut Acad. Arts 10 (8): 447, 1900 (Evans 1900a). Cololejeunea cordilora Steph., Hedwigia 34 (5): 246, 1895 (Stephani 1895b). Cololejeunea crateris Pócs, Acta Bot. Hung. 54 (1/2): 153, 2012 (Pócs 2012b). Cololejeunea crenulata (Pearson) H.A.Mill., Phytologia 47 (4): 321, 1981 (Miller 1981). Bas.: Leptocolea crenulata Pearson, J. Linn. Soc., Bot. 46 (305): 41, 1922 (Pearson 1922b). 257 Cololejeunea cucullifolia (Herzog) E.A.Hodgs., Rec. Domin. Mus. 4 (11): 127, 1962 (Hodgson 1962a). Bas.: Physocolea cucullifolia Herzog, Trans. & Proc. Roy. Soc. New Zealand 68 (1): 45, 1938 (Herzog 1938c). Cololejeunea cuneifolia Steph., Hedwigia 31 (4): 166, 1892 (Stephani 1892g). Cololejeunea dadeuniana Tixier, Ann. Fac. Sci. Univ. Phnom Penh 3: 186, 1970 (Tixier 1970b). Cololejeunea decemplicata (Steph.) Tixier, Bull. Acad. Malgache (n.ser) 55 (1/2): 213, 1977 [1979] (Tixier 1977a). Bas.: Physocolea decemplicata Steph., Sp. Hepat. (Stephani) 5: 869, 1916 (Stephani 1916b). Cololejeunea decliviloba Steph., Hedwigia 34 (5): 247, 1895 (Stephani 1895b). Cololejeunea dentata (E.W.Jones) R.M.Schust., Beih. Nova Hedwigia 9: 175, 1963 (Schuster 1963a). Bas.: Leptocolea dentata E.W.Jones, Trans. Brit. Bryol. Soc. 2 (2): 161, 1953 (Jones 1953a). Cololejeunea denticulata (Horik.) S.Hatt., Bull. Tokyo Sci. Mus. 11: 99, 1944 (Hattori 1944d). Bas.: Physocolea denticulata Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 287, 1934 (Horikawa 1934). Cololejeunea dentilobula (Steph.) R.M.Schust., J. Hattori Bot. Lab. 26: 241, 1963 (Schuster 1963b). Bas.: Physocolea dentilobula Steph., Sp. Hepat. (Stephani) 5: 892, 1916 (Stephani 1916b). 258 Cololejeunea dianae M.Wigginton, J. Bryol. 28 (4): 364, 2006 (Wigginton 2006). 256 Cololejeunea chamlongiana is possibly conspeciic with Cololejeunea papulilora. 257 Cololejeunea crenulata is possibly conspeciic with Cololejeunea angustilora judging from the protologue. 258 Cololejeunea dentilobula is possibly conspeciic with Cololejeunea decliviloba judging from Stephani’s Icones (unpublished). 328 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Cololejeunea dilatata (Steph.) Mizut., J. Hattori Bot. Lab. 28: 113, 1965 (Mizutani 1965). Bas.: Leptocolea dilatata Steph., Sp. Hepat. (Stephani) 5: 850, 1916 (Stephani 1916b). *** Cololejeunea diplasiolejeunoides Tixier, Bryologist 82 (4): 602, 1979 (Tixier 1979d). *** Cololejeunea distalopapillata (E.W.Jones) R.M.Schust., Beih. Nova Hedwigia 9: 173, 1963 (Schuster 1963a). Bas.: Leptocolea distalopapillata E.W.Jones, Trans. Brit. Bryol. Soc. 3 (2): 202, 1957 (Jones 1957b). * Cololejeunea efusa (Mitt.) Steph., Bot. Jahrb. Syst. 23 (1/2, 3): 309, 1896 (Stephani 1896a). Bas.: Lejeunea efusa Mitt., Fl. vit.: 415, 1871 [1873] (Mitten 1871). 259 *** Cololejeunea elephantorum Tixier, Ann. Fac. Sci. Univ. Phnom Penh 3: 185, 1970 (Tixier 1970b). ** Cololejeunea ellipsoidea R.M.Schust., Nova Hedwigia 15: 508, 1968 (Schuster 1968b). *** Cololejeunea equialbi Tixier, Ann. Fac. Sci. Univ. Phnom Penh 3: 178, 1970 (Tixier 1970b). *** Cololejeunea ilicis (Herzog) Piippo, J. Hattori Bot. Lab. 68: 133, 1990 (Piippo 1990). Bas.: Leptocolea ilicis Herzog, Symb. Sin. 5: 52, 1930 (Nicholson et al. 1930). * Cololejeunea ischeri Tixier, Trop. Bryol. 11: 59, 1995 (Tixier 1995b). 260 ** Cololejeunea lavicans (Steph.) Mizut., J. Hattori Bot. Lab. 28: 115, 1965 (Mizutani 1965). Bas.: Physocolea lavicans Steph., Sp. Hepat. (Stephani) 5: 893, 1916 (Stephani 1916b). * Cololejeunea fredericii Onr., Rev. Bryol. Lichénol. 44 (1): 80, 1978 (Onraedt 1978). 261 ** Cololejeunea fusca (Steph.) Mizut., J. Hattori Bot. Lab. 28: 120, 1965 (Mizutani 1965). Bas.: Physocolea fusca Steph., Sp. Hepat. (Stephani) 5: 893, 1916 (Stephani 1916b). ** Cololejeunea gottschei (Steph.) Pandé, K.P.Srivast. et Ahmad, J. Indian Bot. Soc. 36 (3): 345, 1957 (Pandé et al. 1957). Bas.: Physocolea gottschei Steph., Sp. Hepat. (Stephani) 5: 894, 1916 (Stephani 1916b). *** Cololejeunea grossestyla M.Wigginton, J. Bryol. 28 (4): 369, 2006 (Wigginton 2006). *** Cololejeunea grushvitzkiana Pócs, Bot. Zhurn. (Moscow & Leningrad) 56 (5): 674, 1971 (Pócs 1971). *** Cololejeunea hainanensis R.L.Zhu, J. Hattori Bot. Lab. 78: 87, 1995 (Zhu 1995). *** Cololejeunea harrisii Pócs, Acta Bot. Acad. Sci. Hung. 21 (3/4): 357, 1975 (Pócs 1975). *** Cololejeunea hildebrandii (Austin) Steph., Bull. Herb. Boissier 5 (10): 842, 1897 (Stephani 1897c). Bas.: Lejeunea hildebrandii Austin, Bot. Bull. (Hanover) 1 (8): 35, 1876 (Austin 1876a). *** Cololejeunea hirta Steph., Bull. Misc. Inform. Kew 1899 (151/152): 125, 1899 (MacGregor 1899). ** Cololejeunea hodgsoniae (Herzog) E.A.Hodgs., Trans. Roy. Soc. New Zealand, Bot. 3 (11): 184, 1967 (Hodgson 1967). Bas.: Physocolea hodgsoniae Herzog, Trans. & Proc. Roy. Soc. New Zealand 68 (1): 46, 1938 (Herzog 1938c). 259 Cololejeunea efusa may be conspeciic with Cololejeunea trichomanis. 260 Cololejeunea leischeri is near to or conspeciic with Cololejeunea bidentula. 261 Cololejeunea fredericii is possibly conspeciic with Cololejeunea polyantha. World checklist of hornworts and liverworts 329 *** Cololejeunea horikawana (S.Hatt.) Mizut., J. Hattori Bot. Lab. 24: 254, 1961 (Mizutani 1961). Bas.: Leptocolea horikawana S.Hatt., J. Jap. Bot. 18 (11): 653, 1942 (Hattori 1942). *** Cololejeunea huerlimannii Tixier, Nova Hedwigia 31: 773, 1979 (Tixier 1979b). ** Cololejeunea inlexifolia R.M.Schust., Phytologia 56 (7): 458, 1985 (Schuster 1985c). *** Cololejeunea iradieri M.Infante et Heras, Trop. Bryol. 17: 14, 1999 (Infante and Heras 1999). ** Cololejeunea irianensis Tixier, Bull. Jard. Bot. Natl. Belg. 59 (3/4): 440, 1989 (Tixier 1989). *** Cololejeunea johannis-winkleri (Herzog) R.L.Zhu, Nova Hedwigia 79 (3/4): 528, 2004 (Zhu et al. 2004). Bas.: Leptocolea johannis-winkleri Herzog, Mitt. Inst. Allg. Bot. Hamburg 7 (3): 214, 1931 (Herzog 1931a). ** Cololejeunea kegelii Steph., Hedwigia 34 (5): 249, 1895 (Stephani 1895b). ** Cololejeunea khanii Tixier, Dacca Univ. Stud., B 15: 9, 1967 (Tixier 1967). * Cololejeunea kohkongensis Tixier, Bryophyt. Biblioth. 27: 302, 1985 (Tixier 1985a). 262 *** Cololejeunea lichenyae R.D.Porley, N.G.Hodgetts et M.Wigginton, J. Bryol. 29 (1): 7, 2007 (Wigginton et al. 2007). * Cololejeunea lobulilineata Tixier, Trop. Bryol. 11: 42, 1995 (Tixier 1995b). 263 *** Cololejeunea longifolia (Mitt.) Benedix ex Mizut., J. Hattori Bot. Lab. 26: 184, 1963 (Mizutani 1963). Bas.: Lejeunea longifolia Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 117, 1860 [1861] (Mitten 1860c). *** Cololejeunea magna (Tixier) M.Infante et Heras, Trop. Bryol. 17: 17, 1999 (Infante and Heras 1999). Bas.: Cololejeunea harrisii var. magna Tixier, Trop. Bryol. 11: 56, 1995 (Tixier 1995b). ** Cololejeunea magniica Pócs, Acta Bryolichenol. Asiat. 4: 88, 2011 (Pócs and Piippo 2011). ** Cololejeunea magnilobula (Horik.) S.Hatt., Bull. Tokyo Sci. Mus. 11: 99, 1944 (Hattori 1944d). Bas.: Physocolea magnilobula Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 288, 1934 (Horikawa 1934). * Cololejeunea micronesica H.A.Mill. et Bonner, Beih. Nova Hedwigia 11: 63, 1963 (Miller et al. 1963). 264 ** Cololejeunea mooreaensis Tixier, Bauhinia 8 (4): 226, 1987 (Hürlimann 1987). * Cololejeunea ninguana Tixier, Nova Hedwigia 31: 776, 1979 (Tixier 1979b). 265 ** Cololejeunea obcordata (Austin) A.Evans, Trans. Connecticut Acad. Arts 10 (8): 448, 1900 (Evans 1900a). Bas.: Lejeunea obcordata Austin, Bot. Bull. (Hanover) 1 (8): 36, 1876 (Austin 1876a). 262 Cololejeunea kohkongensis may be conspeciic with Cololejeunea plagiophylla. 263 Cololejeunea lobulilineata seems to be a depauperate form of some other species, maybe Cololejeunea obtusifolia. 264 Cololejeunea micronesica is possibly conspeciic with Cololejeunea cookei ide Schuster (1980c, 1983a). 265 Cololejeunea ninguana is possibly conspeciic with Cololejeunea decliviloba (Yu et al. 2013). 330 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Cololejeunea obliqua (Nees et Mont.) Schifn., Bot. Jahrb. Syst. 23 (5): 586, 1897 (Schifner 1897). Bas.: Lejeunea obliqua Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 264, 1843 (Montagne 1843). * Cololejeunea oblongiperianthia (P.C.Wu et J.S.Lou) Piippo, J. Hattori Bot. Lab. 68: 134, 1990 (Piippo 1990). Bas.: Leptocolea oblongiperianthia P.C.Wu et J.S.Lou, Acta Phytotax. Sin. 16 (4): 109, 1978 (Wu and Lou 1978). 266 ** Cololejeunea oleana Sim, Trans. Roy. Soc. South Africa 15 (1): 49, 1926 (Sim 1926). ** Cololejeunea ovalifolia A.Evans, Trans. Connecticut Acad. Arts 10 (8): 450, 1900 (Evans 1900a). ** Cololejeunea panchoana Tixier, Ann. Fac. Sci. Univ. Phnom Penh 3: 183, 1970 (Tixier 1970b). *** Cololejeunea paniensis (Tixier) Grolle, J. Bryol. 8 (4): 485, 1975 (Grolle 1975d). Bas.: Jovetastella paniensis Tixier, Rev. Bryol. Lichénol. 39 (4): 662, 1973 [1974] (Tixier 1973c). ** Cololejeunea papulilora Steph., Akad. Wiss. Wien, Math.-Naturwiss. Kl., Denkschr. 85: 199, 1910 (Stephani 1910a). ** Cololejeunea pentagona (Mitt.) Steph., Bot. Jahrb. Syst. 23 (1/2, 3): 309, 1896 (Stephani 1896a). Bas.: Lejeunea pentagona Mitt., Fl. vit.: 416, 1871 [1873] (Mitten 1871). *** Cololejeunea plagiochiliana Tixier, Bot. Not. 128: 428, 1975 [1976] (Tixier 1975a). ** Cololejeunea planiuscula Tixier, Cryptog. Bryol. Lichénol. 16 (3): 229, 1995 (Tixier 1995a). Based on: Cololejeunea planiuscula Tixier, Candollea 46 (2): 289, 1991 (Tixier 1991), nom. inval. *** Cololejeunea pseudoserrata Tixier, Nova Hedwigia 31: 770, 1979 (Tixier 1979b). * Cololejeunea pteroporum Tixier, Bryophyt. Biblioth. 27: 271, 1985 (Tixier 1985a). 267 ** Cololejeunea pulchella (Mitt.) R.M.Schust., J. Hattori Bot. Lab. 26: 241, 1963 (Schuster 1963b). Bas.: Lejeunea pulchella Mitt., Bot. antarct. voy. II (Fl. Nov.Zel. 2): 157, 1854 (Mitten 1854). ** Cololejeunea pulchella var. stylifera R.M.Schust., Phytologia 56 (7): 458, 1985 (Schuster 1985c). *** Cololejeunea quadridentata (S.Hatt.) Grolle, Acta Bot. Fenn. 125: 65, 1984 (Grolle and Piippo 1984). Bas.: Leptocolea quadridentata S.Hatt., Bot. Mag. (Tokyo) 64 (755/756): 117, 1951 (Hattori 1951c). ** Cololejeunea retusula (Mitt.) H.A.Mill., Phytologia 47 (4): 322, 1981 (Miller 1981). Bas.: Lejeunea retusula Mitt., Fl. vit.: 416, 1871 [1873] (Mitten 1871). ** Cololejeunea salgadoi Onr., Bull. Jard. Bot. Natl. Belg. 59 (3/4): 433, 1989 (Onraedt 1989). *** Cololejeunea sanctae-helenae M.Wigginton, J. Bryol. 28 (4): 366, 2006 (Wigginton 2006). ** Cololejeunea serrulata Steph., Hedwigia 34 (5): 252, 1895 (Stephani 1895b). 266 Cololejeunea oblongiperianthia is possibly conspeciic with Cololejeunea trichomanis (Zhu and So 2001). 267 Cololejeunea pteroporum is possibly conspeciic with Cololejeunea spathulifolia. World checklist of hornworts and liverworts 331 *** Cololejeunea setiloba A.Evans, Bryologist 16 (4): 51, 1913 (Evans 1913). *** Cololejeunea siangensis G.Asthana et S.C.Srivast., Bryophyt. Biblioth. 60: 57, 2003 (Asthana and Srivastava 2003). *** Cololejeunea skottsbergii Herzog, Nat. Hist. Juan Fernandez (Botany) 2 (5): 749, 1942 (Herzog 1942a). *** Cololejeunea societatis Tixier, Bauhinia 8 (4): 230, 1987 (Hürlimann 1987). * Cololejeunea spathulifolia (Steph.) H.A.Mill., Phytologia 47 (4): 322, 1981 (Miller 1981). Bas.: Leptocolea spathulifolia Steph., Sp. Hepat. (Stephani) 5: 855, 1916 (Stephani 1916b). 268 ** Cololejeunea spruceana Tixier, Cryptog. Bryol. Lichénol. 16 (3): 229, 1995 (Tixier 1995a). Based on: Cololejeunea spruceana Tixier, Candollea 46 (2): 286, 1991 (Tixier 1991), nom. inval. ** Cololejeunea stenophylla Herzog, Bot. Not. 100 (4): 330, 1947 (Herzog 1947). *** Cololejeunea streimannii Pócs, Acta Bryolichenol. Asiat. 4: 96, 2011 (Pócs and Piippo 2011). ** Cololejeunea streimannii subsp. solomonensis Pócs, Acta Bryolichenol. Asiat. 4: 96, 2011 (Pócs and Piippo 2011). *** Cololejeunea subalpina Pócs, Acta Bryolichenol. Asiat. 4: 98, 2011 (Pócs and Piippo 2011). ** Cololejeunea subcristata A.Evans, Bryologist 20 (2): 24, 1917 (Evans 1917b). * Cololejeunea takamakae Tixier, Bryophyt. Biblioth. 27: 319, 1985 (Tixier 1985a). 269 *** Cololejeunea tanzaniae Pócs, J. Hattori Bot. Lab. 48: 312, 1980 (Pócs 1980b). ** Cololejeunea tenuiparietata Tixier, Trop. Bryol. 11: 56, 1995 (Tixier 1995b). ** Cololejeunea teurnoumensis Tixier, Ann. Fac. Sci. Univ. Phnom Penh 3: 183, 1970 (Tixier 1970b). *** Cololejeunea timoi Pócs, Acta Bryolichenol. Asiat. 4: 98, 2011 (Pócs and Piippo 2011). ** Cololejeunea touwii Pócs, Acta Bryolichenol. Asiat. 4: 101, 2011 (Pócs and Piippo 2011). *** Cololejeunea tranninhiana Tixier, Ann. Hist.-Nat. Mus. Natl. Hung. 66: 97, 1974 (Tixier 1974). *** Cololejeunea trichomanis (Gottsche) Besch., Rev. Bryol. 19 (1): 14, 1892 (Bescherelle 1892). Bas.: Lejeunea trichomanis Gottsche, Abh. Naturwiss. Vereins Bremen 7: 362, 1882 (Gottsche 1882). * Cololejeunea tuksapiana Tixier, Ann. Fac. Sci. Univ. Phnom Penh 3: 186, 1970 (Tixier 1970b). 270 ** Cololejeunea vulcania Tixier, Ann. Fac. Sci. Univ. Phnom Penh 3: 181, 1970 (Tixier 1970b). *** Cololejeunea yoshinagana (S.Hatt.) Mizut., J. Hattori Bot. Lab. 24: 250, 1961 (Mizutani 1961). Bas.: Leptocolea yoshinagana S.Hatt., Bull. Tokyo Sci. Mus. 11: 115, 1944 (Hattori 1944d). 268 Cololejeunea spathulifolia is possibly conspeciic with Cololejeunea obliqua (Pócs and Piippo 2011). 269 Cololejeunea takamakae is possibly conspeciic with Cololejeunea angustilora. 270 Cololejeunea tuksapiana is possibly conspeciic with Cololejeunea aequalis. 332 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) subg. Metzgeriopsis (K.I.Goebel) Pócs, Acta Bryolichenol. Asiat. 4: 106, 2011 (Pócs and Piippo 2011). Bas.: Metzgeriopsis K.I.Goebel, Ann. Jard. Bot. Buitenzorg 7 (1): 54, 1888 (Goebel 1888). *** Cololejeunea metzgeriopsis (K.I.Goebel) Gradst., R.Wilson, Ilk.-Borg. et Heinrichs, Bot. J. Linn. Soc. 151 (3): 306, 2006 (Gradstein et al. 2006). Bas.: Lejeunea metzgeriopsis K.I.Goebel, Flora 72 (1): 2, 1889 (Goebel 1889). * *** subg. Pedinolejeunea Benedix ex Mizut., J. Hattori Bot. Lab. 24: 240, 1961 (Mizutani 1961). ** Cololejeunea abnormis Mizut., J. Hattori Bot. Lab. 33: 260, 1970 (Mizutani 1970). *** Cololejeunea adhaesiva (Mitt.) R.M.Schust., Beih. Nova Hedwigia 9: 177, 1963 (Schuster 1963a). Bas.: Lejeunea adhaesiva Mitt., J. Linn. Soc., Bot. 22 (146): 325, 1886 (Mitten 1886b). ** Cololejeunea adnata Tixier, Bull. Acad. Malgache (n.ser) 55 (1/2): 194, 1977 [1979] (Tixier 1977a). *** Cololejeunea africana (Steph.) R.M.Schust., Beih. Nova Hedwigia 9: 173, 1963 (Schuster 1963a). Bas.: Physocolea africana Steph., Sp. Hepat. (Stephani) 5: 867, 1916 (Stephani 1916b). ** Cololejeunea ambeliensis Tixier, Bryophyt. Biblioth. 27: 142, 1985 (Tixier 1985a). ** Cololejeunea amieuensis Tixier, Nova Hedwigia 31: 748, 1979 (Tixier 1979b). ** Cololejeunea andapania Tixier, Bull. Acad. Malgache (n.ser) 55 (1/2): 178, 1977 [1979] (Tixier 1977a). *** Cololejeunea angulata (Steph.) Mizut., J. Hattori Bot. Lab. 28: 108, 1965 (Mizutani 1965). Bas.: Leptocolea angulata Steph., Sp. Hepat. (Stephani) 5: 847, 1916 (Stephani 1916b). *** Cololejeunea ankaiana Tixier, Bryophyt. Biblioth. 27: 62, 1985 (Tixier 1985a). ** Cololejeunea attilana Pócs, Magyar Bot. Kut. Ezredf. Tanul. Borhidi: 186, 2002 (Pócs 2002a). *** Cololejeunea auriculata (E.W.Jones) R.M.Schust., Beih. Nova Hedwigia 9: 177, 1963 (Schuster 1963a). Bas.: Leptocolea auriculata E.W.Jones, Trans. Brit. Bryol. Soc. 2 (2): 152, 1953 (Jones 1953b). ** Cololejeunea autoica (Steph.) Grolle, Bryophyt. Biblioth. 48: 43, 1995 (Grolle 1995). Bas.: Physocolea autoica Steph., Sp. Hepat. (Stephani) 5: 867, 1916 (Stephani 1916b). *** Cololejeunea bekkeri Tixier, Cryptog. Bryol. Lichénol. 16 (3): 229, 1995 (Tixier 1995a). Based on: Cololejeunea bekkeri Tixier, Candollea 46 (2): 269, 1991 (Tixier 1991), nom. inval. *** Cololejeunea bischleriana Tixier, Bradea 3 (6): 36, 1980 (Tixier 1980a). ** Cololejeunea bolovenensis Tixier, Nat. Hist. Bull. Siam Soc. 24 (3/4): 442, 1973 (Tixier 1973b). *** Cololejeunea borbonica Tixier, Bull. Acad. Malgache (n.ser) 55 (1/2): 188, 1977 [1979] (Tixier 1977a). World checklist of hornworts and liverworts 333 * Cololejeunea brunelii Tixier, Dacca Univ. Stud., B 15: 10, 1967 (Tixier 1967). 271 *** Cololejeunea cardiocarpa (Mont.) A.Evans, Mem. Torrey Bot. Club 8 (2): 172, 1902 (Evans 1902a). Bas.: Lejeunea cardiocarpa Mont., Hist. Phys. Cuba, Bot., Pl. Cell.: 476, 1842 (Montagne 1842a). * Cololejeunea chittagongensis Tixier, Bryophyt. Biblioth. 27: 97, 1985 (Tixier 1985a). 272 *** Cololejeunea cocoscola Tixier, Cryptog. Bryol. Lichénol. 14 (3): 353, 1993 (Tixier 1993). ** Cololejeunea cremersii Tixier, Cryptog. Bryol. Lichénol. 16 (3): 229, 1995 (Tixier 1995a). Based on: Cololejeunea cremersii Tixier, Candollea 46 (2): 271, 1991 (Tixier 1991), nom. inval. ** Cololejeunea cristata (Steph.) R.M.Schust., Beih. Nova Hedwigia 9: 173, 1963 (Schuster 1963a). Bas.: Physocolea cristata Steph., Sp. Hepat. (Stephani) 5: 869, 1916 (Stephani 1916b). ** Cololejeunea cuneata (Lehm. et Lindenb.) Herzog, Bot. Not. 100 (4): 320, 1947 (Herzog 1947). Bas.: Jungermannia cuneata Lehm. et Lindenb., Nov. Stirp. Pug. 4: 56, 1832 (Lehmann 1832). * Cololejeunea deroinii Tixier, Cryptog. Bryol. Lichénol. 14 (3): 355, 1993 (Tixier 1993). * Cololejeunea deslooveri Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 47 (1/2): 227, 1977 (Vanden Berghen 1977). 273 * Cololejeunea dzumacensis Tixier, Nova Hedwigia 31: 754, 1979 (Tixier 1979b). 274 *** Cololejeunea ecuadoriensis Pócs, Acta Bot. Hung. 44 (3/4): 372, 2002 (Pócs 2002b). * Cololejeunea epiphylla G.Asthana et A.Shukla, Cryptog. Bryol. 31 (3): 218, 2010 (Asthana and Shukla 2010). 275 ** Cololejeunea issilobula Herzog, Trans. Brit. Bryol. Soc. 1 (4): 323, 1950 (Herzog 1950a). ** Cololejeunea lorencei Tixier, Cryptog. Bryol. Lichénol. 14 (3): 355, 1993 (Tixier 1993). ** Cololejeunea foliicola S.C.Srivast. et G.Srivast., Proc. Indian Acad. Sci. Pl. Sci. 99 (2): 86, 1989 (Srivastava and Srivastava 1989a). ** Cololejeunea fructumarginata Tixier, Bryophyt. Biblioth. 27: 58, 1985 (Tixier 1985a). *** Cololejeunea furcilobulata (Berrie et E.W.Jones) R.M.Schust., Beih. Nova Hedwigia 9: 178, 1963 (Schuster 1963a). Bas.: Leptocolea furcilobulata Berrie et E.W.Jones, Trans. Brit. Bryol. Soc. 2 (3): 417, 1954 (Jones 1954c). *** Cololejeunea geissleriana Tixier, Bradea 3 (6): 37, 1980 (Tixier 1980a). 271 Cololejeunea brunelii is conspeciic with Cololejeunea madothecoides in Tixier (1985a), but it belongs to the Cololejeunea raduliloba, Cololejeunea furcilobulata, Cololejeunea paucimarginata species complex. 272 Cololejeunea chittagongensis is probably conspeciic with Cololejeunea schwabei. 273 Cololejeunea deslooveri is possibly conspeciic with Cololejeunea cristata. 274 Cololejeunea dzumacensis is possibly conspeciic with Cololejeunea lanciloba, but the type specimen has not been localized. 275 Cololejeunea epiphylla is closely related to Cololejeunea chittagongensis and Cololejeunea schwabei. 334 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Cololejeunea georgiana Tixier, Bryophyt. Biblioth. 27: 145, 1985 (Tixier 1985a). *** Cololejeunea guadelupensis Tixier, Cryptog. Bryol. Lichénol. 16 (3): 229, 1995 (Tixier 1995a). Based on: Cololejeunea guadelupensis Tixier, Candollea 46 (2): 276, 1991 (Tixier 1991), nom. inval. *** Cololejeunea hebridensis Tixier, Bot. Not. 128: 425, 1975 [1976] (Tixier 1975a). *** Cololejeunea hinidumae Onr., Acta Bot. Acad. Sci. Hung. 25 (1/2): 109, 1979 (Onraedt 1979). * Cololejeunea hoabinhiana Tixier, Ann. Hist.-Nat. Mus. Natl. Hung. 66: 91, 1974 (Tixier 1974). 276 ** Cololejeunea hoeana Tixier, Bryophyt. Biblioth. 27: 56, 1985 (Tixier 1985a). ** Cololejeunea hungii Tixier, Ann. Hist.-Nat. Mus. Natl. Hung. 66: 91, 1974 (Tixier 1974). *** Cololejeunea indosinica Tixier, Bryophyt. Biblioth. 27: 63, 1985 (Tixier 1985a). ** Cololejeunea inoueana Mizut., J. Hattori Bot. Lab. 57: 440, 1984 (Mizutani 1984c). ** Cololejeunea japonica (Schifn.) Mizut., J. Hattori Bot. Lab. 24: 241, 1961 (Mizutani 1961). Bas.: Leptocolea japonica Schifn., Ann. Bryol. 2: 92, 1929 (Schifner 1929). *** Cololejeunea jonesii Pócs, Acta Bot. Acad. Sci. Hung. 21 (3/4): 361, 1975 (Pócs 1975). *** Cololejeunea kapingaensis H.A.Mill., Bryologist 59 (3): 170, 1956 (Miller 1956). ** Cololejeunea kiriromensis Tixier, Bryophyt. Biblioth. 27: 147, 1985 (Tixier 1985a). *** Cololejeunea kulenensis Tixier, Bryophyt. Biblioth. 27: 71, 1985 (Tixier 1985a). Nom. nov. pro Leptocolea verdoornii Herzog, Ann. Bryol. 5: 97, 1932 (Herzog 1932a). ** Cololejeunea laevigata (Mitt.) R.M.Schust., J. Hattori Bot. Lab. 26: 241, 1963 (Schuster 1963b). Bas.: Lejeunea laevigata Mitt., Bot. antarct. voy. II (Fl. Nov.Zel. 2): 157, 1854 (Mitten 1854). *** Cololejeunea lanciloba Steph., Hedwigia 34 (5): 250, 1895 (Stephani 1895b). *** Cololejeunea latilobula (Herzog) Tixier, Bryophyt. Biblioth. 27: 156, 1985 (Tixier 1985a). Bas.: Leptocolea latilobula Herzog, Symb. Sin. 5: 54, 1930 (Nicholson et al. 1930). *** Cololejeunea latistyla R.L.Zhu, Hikobia 11: 544, 1994 (Zhu et al. 1994). *** Cololejeunea leloutrei (E.W.Jones) R.M.Schust., Beih. Nova Hedwigia 9: 173, 1963 (Schuster 1963a). Bas.: Leptocolea leloutrei E.W.Jones, Trans. Brit. Bryol. Soc. 2 (2): 146, 1953 (Jones 1953b). ** Cololejeunea leloutrei var. microlobulata Tixier, Bryophyt. Biblioth. 27: 78, 1985 (Tixier 1985a). ** Cololejeunea leloutrei var. ulugurica Pócs ex Tixier, Bryophyt. Biblioth. 27: 74, 1985 (Tixier 1985a). *** Cololejeunea lemuriana Tixier, Bull. Acad. Malgache (n.ser) 55 (1/2): 191, 1977 [1979] (Tixier 1977a). ** Cololejeunea littoralis Tixier, Bryophyt. Biblioth. 27: 163, 1985 (Tixier 1985a). 276 Cololejeunea hoabinhiana is possibly conspeciic with Cololejeunea lanciloba. World checklist of hornworts and liverworts 335 ** Cololejeunea longistylis A.Evans, Trans. Connecticut Acad. Arts 10 (8): 453, 1900 (Evans 1900a). *** Cololejeunea magnistyla (Horik.) Mizut., J. Hattori Bot. Lab. 24: 243, 1961 (Mizutani 1961). Bas.: Leptocolea magnistyla Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 1: 131, 1932 (Horikawa 1932c). * Cololejeunea malaccensis Tixier, Bryophyt. Biblioth. 27: 42, 1985 (Tixier 1985a). 277 ** Cololejeunea malayana Tixier, Bryophyt. Biblioth. 27: 154, 1985 (Tixier 1985a). *** Cololejeunea marginata (Lehm. et Lindenb.) Pearson, Forh. Vidensk.-Selsk. Kristiania 1892 (8): 9, 1892 (Pearson 1892). Bas.: Jungermannia marginata Lehm. et Lindenb., Nov. Stirp. Pug. 5: 11, 1833 (Lehmann 1833). * Cololejeunea maritima Tixier, Nova Hedwigia 31: 752, 1979 (Tixier 1979b). *** Cololejeunea minutilobula Herzog, Rev. Bryol. Lichénol. 20 (1/2): 171, 1951 [1952] (Herzog 1951a). *** Cololejeunea nigerica (E.W.Jones) R.M.Schust., Beih. Nova Hedwigia 9: 177, 1963 (Schuster 1963a). Bas.: Leptocolea nigerica E.W.Jones, Trans. Brit. Bryol. Soc. 2 (2): 150, 1953 (Jones 1953b). ** Cololejeunea occidentalis (E.W.Jones) Vanden Berghen, Rev. Bryol. Lichénol. 44 (4): 449, 1978 (Vanden Berghen 1978). Bas.: Leptocolea cristata var. occidentalis E.W.Jones, Trans. Brit. Bryol. Soc. 2 (2): 149, 1953 (Jones 1953b). ** Cololejeunea onraedtii Tixier, Bull. Acad. Malgache (n.ser) 55 (1/2): 202, 1977 [1979] (Tixier 1977a). *** Cololejeunea paciica Pócs, Acta Bot. Hung. 54 (1/2): 158, 2012 (Pócs 2012b). ** Cololejeunea panamensis G.Dauphin et Pócs, Trop. Bryol. 27: 76, 2006 (Dauphin et al. 2006). *** Cololejeunea paucimarginata Tixier, Bryophyt. Biblioth. 27: 100, 1985 (Tixier 1985a). ** Cololejeunea perakensis Tixier, Bryophyt. Biblioth. 27: 95, 1985 (Tixier 1985a). *** Cololejeunea plagiophylla Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 73, 1953 (Benedix 1953). *** Cololejeunea planissima (Mitt.) Abeyw., Ceylon J. Sci., Biol. Sci. 2 (1): 73, 1959 (Abeywickrama 1959). Bas.: Lejeunea planissima Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 117, 1860 [1861] (Mitten 1860c). ** Cololejeunea planissima var. chagosensis Pócs, J. Bryol. 28 (1): 14, 2006 (Seaward et al. 2006). ** Cololejeunea praeruptorum Tixier, Bull. Acad. Malgache (n.ser) 55 (1/2): 205, 1977 [1979] (Tixier 1977a). ** Cololejeunea producta (Mitt.) S.Hatt., Fl. E. Himalaya: 533, 1966 (Hattori 1966c). Bas.: Lejeunea producta Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 117, 1860 [1861] (Mitten 1860c). 277 Cololejeunea malaccensis may be conspeciic with Cololejeunea stylosa. 336 * *** * ** ** *** *** ** *** *** *** ** *** *** ** *** ** ** ** *** *** ** ** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Cololejeunea punctata (Gottsche) Pearson, Forh. Vidensk.-Selsk. Kristiania 1892 (14): 9, 1893 (Pearson 1893). Bas.: Lejeunea punctata Gottsche, Abh. Naturwiss. Vereins Bremen 7: 361, 1882 (Gottsche 1882). 278 Cololejeunea raduliloba Steph., Hedwigia 34 (5): 251, 1895 (Stephani 1895b). Cololejeunea reineckeana Steph., Bot. Jahrb. Syst. 23 (1/2, 3): 309, 1896 (Stephani 1896a). Cololejeunea saltuum Tixier, Bull. Acad. Malgache (n.ser) 55 (1/2): 188, 1977 [1979] (Tixier 1977a). Cololejeunea sambiroana Tixier, Bryologist 82 (4): 608, 1979 (Tixier 1979d). Cololejeunea saroltae Pócs, Acta Bot. Hung. 54 (1/2): 160, 2012 (Pócs 2012b). Cololejeunea schusteri Pócs, Acta Bot. Hung. 44 (3/4): 376, 2002 (Pócs 2002b). Cololejeunea schwabei Herzog, J. Hattori Bot. Lab. 14: 54, 1955 (Herzog and Noguchi 1955). Cololejeunea selangorensis Tixier, Bryophyt. Biblioth. 27: 166, 1985 (Tixier 1985a). Cololejeunea shibiensis Mizut., J. Hattori Bot. Lab. 57: 437, 1984 (Mizutani 1984c). Cololejeunea smitinandii Tixier, Bryophyt. Biblioth. 27: 131, 1985 (Tixier 1985a). Based on: Cololejeunea smitinandii Tixier, Nat. Hist. Bull. Siam Soc. 24 (3/4): 439, 1973 (Tixier 1973b), nom. inval. Cololejeunea stoniana Tixier, Bot. Not. 128: 429, 1975 [1976] (Tixier 1975a). Cololejeunea stylosa Steph., Trans. Connecticut Acad. Arts 10 (8): 454, 1900 (Evans 1900a). Based on: Cololejeunea stylosa Steph., Hedwigia 27 (11/12): 289, 1888 (Stephani 1888c), nom. inval. Cololejeunea subcardiocarpa Tixier, Bradea 3 (6): 39, 1980 (Tixier 1980a). Cololejeunea subinlata Tixier, Bull. Acad. Malgache (n.ser) 55 (1/2): 182, 1977 [1979] (Tixier 1977a). Cololejeunea submarginata Tixier, Bradea 3 (6): 40, 1980 (Tixier 1980a). Cololejeunea subminutilobula Mizut., J. Hattori Bot. Lab. 24: 282, 1961 (Mizutani 1961). Nom. nov. pro Leptocolea minutilobula Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 1: 16, 1931 (Horikawa 1931b). Cololejeunea subscariosa (Spruce) Pócs, Acta Bot. Hung. 56 (1/2): 197, 2014 (Pócs et al. 2014). Bas.: Lejeunea subscariosa Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 300, 1884 (Spruce 1884). Cololejeunea subtriapiculata Tixier, Nova Hedwigia 31: 744, 1979 (Tixier 1979b). Cololejeunea succinea Tixier, Bull. Acad. Malgache (n.ser) 55 (1/2): 194, 1977 [1979] (Tixier 1977a). Cololejeunea surinamensis Tixier, Bradea 3 (6): 42, 1980 (Tixier 1980a). Cololejeunea tahitensis Tixier, Cryptog. Bryol. Lichénol. 14 (3): 359, 1993 (Tixier 1993). Cololejeunea tamatavensis Tixier, Bull. Acad. Malgache (n.ser) 55 (1/2): 191, 1977 [1979] (Tixier 1977a). 278 Cololejeunea punctata is a doubtful taxon. he type specimen has been destroyed and the identity is unclear (Grolle 1995). World checklist of hornworts and liverworts * * ** * * * *** *** ** ** *** 337 Cololejeunea taprobanea Tixier, Bryophyt. Biblioth. 27: 158, 1985 (Tixier 1985a). Cololejeunea thailandensis Tixier, Nat. Hist. Bull. Siam Soc. 24 (3/4): 441, 1973 (Tixier 1973b). Cololejeunea triapiculata (Herzog) Tixier, Gard. Bull. Singapore 25 (3): 344, 1971 (Tixier 1971). Bas.: Leptocolea triapiculata Herzog, Ann. Bryol. 5: 95, 1932 (Herzog 1932a). Cololejeunea tribracteata Tixier, Trop. Bryol. 11: 46, 1995 (Tixier 1995b). 279 Cololejeunea tridentata Tixier, Bryophyt. Biblioth. 27: 83, 1985 (Tixier 1985a). Cololejeunea uchimae Amakawa, J. Jap. Bot. 33 (5): 142, 1958 (Amakawa 1958a). 280 Cololejeunea verwimpii Tixier, Cryptog. Bryol. Lichénol. 16 (3): 230, 1995 (Tixier 1995a). Cololejeunea vidaliana Tixier, Nat. Hist. Bull. Siam Soc. 24 (3/4): 444, 1973 (Tixier 1973b). Cololejeunea vietnamensis Tixier, Bryophyt. Biblioth. 27: 127, 1985 (Tixier 1985a). Cololejeunea vitaliana Tixier, Cryptog. Bryol. Lichénol. 16 (3): 230, 1995 (Tixier 1995a). Cololejeunea yakusimensis (S.Hatt.) Mizut., J. Hattori Bot. Lab. 57: 430, 1984 (Mizutani 1984c). Bas.: Leptocolea lanciloba var. yakusimensis S.Hatt., J. Jap. Bot. 18 (11): 655, 1942 (Hattori 1942). subg. Protocolea R.M.Schust., Beih. Nova Hedwigia 9: 171, 1963 (Schuster 1963a). ** Cololejeunea chuahiana Pócs, Polish Bot. J. 47 (1): 11, 2002 (Pócs 2002c). 281 ** Cololejeunea dauphinii R.L.Zhu, J. Bryol. 28 (3): 277, 2006 (Zhu 2006b). Nom. nov. pro Cololejeunea tixieri M.I.Morales et G.Dauphin, Trop. Bryol. 14: 133, 1998 (Morales and Dauphin 1998), nom. illeg. 282 ** Cololejeunea discilora Tixier, Bryologist 82 (4): 604, 1979 (Tixier 1979d). 283 * *** subg. Taeniolejeunea (Zwickel) Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 21, 1953 (Benedix 1953). Bas.: Taeniolejeunea Zwickel, Ann. Bryol. 6: 106, 1933 (Zwickel 1933). * Cololejeunea amoena Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 25, 1953 (Benedix 1953). 284 *** Cololejeunea appressa (A.Evans) Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 31, 1953 (Benedix 1953). Bas.: Leptocolea appressa A.Evans, Bull. Torrey Bot. Club 39 (12): 606, 1912 [1913] (Evans 1912d). 279 Cololejeunea tribracteata is possibly conspeciic with Cololejeunea africana. 280 Cololejeunea uchimae is conspeciic with Cololejeunea raduliloba in Mizutani (1961), but it is accepted in recent Japanese checklists. 281 Cololejeunea chuahiana is probably a Myriocoleopsis species (Yu et al. 2014). 282 Cololejeunea dauphinii is probably a Myriocoleopsis species (Yu et al. 2014). 283 Cololejeunea discilora is probably a Myriocoleopsis species (Yu et al. 2014). 284 Cololejeunea amoena is possibly conspeciic with Cololejeunea loccosa (Söderström et al. 2010a). 338 * * * ** *** ** * ** *** * ** ** ** * ** ** ** ** *** ** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Cololejeunea bachmaensis Tixier, Rev. Bryol. Lichénol. 36 (3/4): 560, 1969 [1970] (Tixier 1969). Cololejeunea bontocensis Tixier, Rev. Bryol. Lichénol. 36 (3/4): 583, 1969 [1970] (Tixier 1969). 285 Cololejeunea crassipapillata Tixier, Rev. Bryol. Lichénol. 36 (3/4): 565, 1969 [1970] (Tixier 1969). Cololejeunea eustacei Pócs, J. Bryol. 29 (2): 83, 2007 (Müller and Pócs 2007). Cololejeunea falcata (Horik.) Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 29, 1953 (Benedix 1953). Bas.: Physocolea falcata Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 1: 22, 1931 (Horikawa 1931b). Cololejeunea falcata var. madecassa Tixier, Bryologist 82 (4): 606, 1979 (Tixier 1979d). Cololejeunea lavida P.C.Wu et J.S.Lou, Acta Phytotax. Sin. 16 (4): 103, 1978 (Wu and Lou 1978). 286 Cololejeunea lavovittata Pócs, Acta Bryolichenol. Asiat. 4: 120, 2011 (Pócs and Piippo 2011). Cololejeunea loccosa (Lehm. et Lindenb.) Schifn., Consp. Hepat. Arch. Ind.: 243, 1898 (Schifner 1898b). Bas.: Jungermannia loccosa Lehm. et Lindenb., Nov. Stirp. Pug. 5: 26, 1833 (Lehmann 1833). Cololejeunea loccosa var. amoenoides Tixier, Cryptog. Bryol. Lichénol. 2 (1): 59, 1981 (Tixier 1981). Cololejeunea loccosa var. angustibracteata Tixier, Cryptog. Bryol. Lichénol. 2 (1): 65, 1981 (Tixier 1981). Cololejeunea loccosa var. aurita Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 24, 1953 (Benedix 1953). Cololejeunea loccosa var. conivens Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 24, 1953 (Benedix 1953). Cololejeunea loccosa var. ocellata Tixier, Cryptog. Bryol. Lichénol. 2 (1): 69, 1981 (Tixier 1981). Cololejeunea loccosa var. plicata Tixier, Cryptog. Bryol. Lichénol. 2 (1): 65, 1981 (Tixier 1981). Cololejeunea loccosa var. trivittata Tixier, Cryptog. Bryol. Lichénol. 2 (1): 62, 1981 (Tixier 1981). Cololejeunea gresicola Tixier, Rev. Bryol. Lichénol. 36 (3/4): 569, 1969 [1970] (Tixier 1969). Cololejeunea gynophthalma Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 32, 1953 (Benedix 1953). Cololejeunea inlata Steph., Hedwigia 34 (5): 249, 1895 (Stephani 1895b). Cololejeunea khiavensis Tixier, Rev. Bryol. Lichénol. 36 (3/4): 557, 1969 [1970] (Tixier 1969). 285 Cololejeunea bontocensis is possibly conspeciic with Cololejeunea pseudostephanii. 286 Cololejeunea lavida may be conspeciic with Cololejeunea perainis (Zhu and So 2001). World checklist of hornworts and liverworts 339 ** Cololejeunea koratensis Tixier, Rev. Bryol. Lichénol. 36 (3/4): 585, 1969 [1970] (Tixier 1969). * Cololejeunea manlinensis Tixier, Rev. Bryol. Lichénol. 36 (3/4): 563, 1969 [1970] (Tixier 1969). ** Cololejeunea maquilingensis Tixier, Rev. Bryol. Lichénol. 36 (3/4): 579, 1969 [1970] (Tixier 1969). ** Cololejeunea mutabilis Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 27, 1953 (Benedix 1953). ** Cololejeunea nakajimae S.Hatt., J. Hattori Bot. Lab. 10: 57, 1953 (Hattori and Kodama 1953). *** Cololejeunea ocellata (Horik.) Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 38, 1953 (Benedix 1953). Bas.: Leptocolea ocellata Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 1: 86, 1932 (Horikawa 1932a). *** Cololejeunea ocelloides (Horik.) Mizut., J. Hattori Bot. Lab. 24: 277, 1961 (Mizutani 1961). Bas.: Leptocolea ocelloides Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 280, 1934 (Horikawa 1934). *** Cololejeunea perainis (Schifn.) Schifn., Consp. Hepat. Arch. Ind.: 245, 1898 (Schifner 1898b). Bas.: Lejeunea perainis Schifn., Nova Acta Acad. Caes. Leop.Carol. German. Nat. Cur. 60 (2): 242, 1893 (Schifner 1893a). * Cololejeunea perainis var. ciconiae Tixier, Bull. Jard. Bot. Natl. Belg. 59 (3/4): 444, 1989 (Tixier 1989). ** Cololejeunea perainis var. elegans Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 35, 1953 (Benedix 1953). ** Cololejeunea perainis var. serrulata Schifn. ex Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 35, 1953 (Benedix 1953). * Cololejeunea polisiana Tixier, Rev. Bryol. Lichénol. 36 (3/4): 550, 1969 [1970] (Tixier 1969). 287 *** Cololejeunea pseudoloccosa (Horik.) Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 36, 1953 (Benedix 1953). Bas.: Leptocolea pseudoloccosa Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 1: 87, 1932 (Horikawa 1932a). *** Cololejeunea pseudostephanii Tixier, Rev. Bryol. Lichénol. 36 (3/4): 571, 1969 [1970] (Tixier 1969). ** Cololejeunea setosa Mizut., J. Hattori Bot. Lab. 29: 163, 1966 (Mizutani 1966). *** Cololejeunea sharpii Mizut., J. Hattori Bot. Lab. 39: 258, 1975 (Mizutani 1975). *** Cololejeunea siamensis Steph., Bot. Tidsskr. 24 (3): 279, 1902 (Stephani 1902b). *** Cololejeunea sphaerodonta Mizut., J. Hattori Bot. Lab. 29: 165, 1966 (Mizutani 1966). *** Cololejeunea stephanii Schifn. ex Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 40, 1953 (Benedix 1953). ** Cololejeunea subloccosa Mizut., J. Hattori Bot. Lab. 57: 168, 1984 (Mizutani 1984b). ** Cololejeunea subocelloides Mizut., J. Hattori Bot. Lab. 57: 163, 1984 (Mizutani 1984b). 287 Cololejeunea polisiana may be conspeciic with Cololejeunea perainis. 340 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) * Cololejeunea tamdaoensis Tixier, Rev. Bryol. Lichénol. 36 (3/4): 567, 1969 [1970] (Tixier 1969). ** Cololejeunea verdoornii (S.Hatt.) S.Hatt., J. Hattori Bot. Lab. 17: 75, 1956 [1957] (Hattori 1956b). Bas.: Taeniolejeunea verdoornii S.Hatt., J. Jap. Bot. 17: 459, 1941 (Hattori 1941). ** Cololejeunea yipii R.L.Zhu, Beih. Nova Hedwigia 121: 346, 2001 (Zhu and So 2001). ** Cololejeunea zantenorum Pócs, Acta Bryolichenol. Asiat. 4: 127, 2011 (Pócs and Piippo 2011). Incertae sedis ** Cololejeunea conchifolia (Gottsche) Gradst., J. Hattori Bot. Lab. 45: 109, 1979 (Gradstein and Hekking 1979). Bas.: Lejeunea conchifolia Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 163, 1864 (Gottsche 1864). 288 ** Cololejeunea dankiaensis Tixier, Phytotaxa 220 (2): 199, 2015 (Söderström et al. 2015d). Based on: Cololejeunea dankiaensis Tixier, Rev. Bryol. Lichénol. 36 (3/4): 581, 1969 [1970] (Tixier 1969), nom. inval. ** Cololejeunea ensifera Tixier, Phytotaxa 220 (2): 199, 2015 (Söderström et al. 2015d). Based on: Cololejeunea ensifera Tixier, Rev. Bryol. Lichénol. 36 (3/4): 562, 1969 [1970] (Tixier 1969), nom. inval. ** Cololejeunea hamata Steph., Hedwigia 34 (5): 248, 1895 (Stephani 1895b). ** Cololejeunea herzogii K.I.Goebel, Biblioth. Bot. 87 (2): 269, 1916 (Stephani 1916a). ** Cololejeunea jamesii (Austin) M.E.Reiner et Pócs, Phytotaxa 208 (1): 98, 2015 (Pócs et al. 2015a). Bas.: Lejeunea jamesii Austin, Bull. Torrey Bot. Club 6 (30): 158, 1877 (Austin 1877). ** Cololejeunea sublatistyla Jian Wang bis et R.L.Zhu, Phytotaxa 161 (2): 165, 2014 (Wang et al. 2014a). ** Cololejeunea tixieri Onr., Bull. Jard. Bot. Natl. Belg. 59 (3/4): 436, 1989 (Onraedt 1989). * Cololejeunea variifolia (Mitt.) Steph., Bot. Jahrb. Syst. 23 (1/2, 3): 309, 1896 (Stephani 1896a). Bas.: Lejeunea variifolia Mitt., Fl. vit.: 415, 1871 [1873] (Mitten 1871). *** Colura (Dumort.) Dumort., Recueil Observ. Jungerm.: 12, 1835 (Dumortier 1835). Bas.: Lejeunea sect. Colura Dumort., Syll. Jungerm. Europ.: 32, 1831 (Dumortier 1831). ** subg. Colura ** sect. Colura *** Colura berghenii Jovet-Ast, Rev. Bryol. Lichénol. 22 (2/3): 245, 1953 [1954] (Jovet-Ast 1953). 288 Cololejeunea conchifolia is probably a Myriocoleopsis species (Yu et al. 2014). World checklist of hornworts and liverworts 341 *** Colura calyptrifolia (Hook.) Dumort., Recueil Observ. Jungerm.: 12, 1835 (Dumortier 1835). Bas.: Jungermannia calyptrifolia Hook., Brit. Jungermann.: tab. 43, 1813 (Hooker 1813). *** Colura hedbergiana Pócs, J. Bryol. 14 (3): 499, 1987 (Jones and Pócs 1987). *** Colura humbertii Jovet-Ast, Rev. Bryol. Lichénol. 22 (2/3): 251, 1953 [1954] (Jovet-Ast 1953). *** Colura irrorata (Spruce) Heinrichs, Y.Yu, Schäf.-Verw. et Pócs, Phytotaxa 66: 58, 2012 (Heinrichs et al. 2012c). Bas.: Myriocolea irrorata Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 305, 1884 (Spruce 1884). ** Colura junghuhniana (Prantl) Steph., Sp. Hepat. (Stephani) 5: 938, 1916 (Stephani 1916b). Bas.: Lejeunea junghuhniana Prantl, Hedwigia 29: xvi, 1890 (Prantl 1890). ** Colura medusa J.Eggers et Pócs, Chenia 11: 22, 2013 (Pócs 2013). ** Colura mizutanii Pócs, Chenia 11: 22, 2013 (Pócs 2013). *** Colura rhynchophora Jovet-Ast, Rev. Bryol. Lichénol. 17 (1/4): 27, 1948 [1949] (Jovet-Ast 1948). *** Colura tenuicornis (A.Evans) Steph., Sp. Hepat. (Stephani) 5: 942, 1916 (Stephani 1916b), nom. conserv. Bas.: Colurolejeunea tenuicornis A.Evans, Trans. Connecticut Acad. Arts 10 (8): 455, 1900 (Evans 1900a). ** sect. Gamolepis Jovet-Ast, Cryptog. Bryol. Lichénol. 4 (3): 207, 1983 (Jovet-Ast 1983). *** Colura cristata Jovet-Ast, Rev. Bryol. Lichénol. 22 (2/3): 291, 1953 [1954] (Jovet-Ast 1953). *** Colura greig-smithii Jovet-Ast, Rev. Bryol. Lichénol. 22 (2/3): 293, 1953 [1954] (Jovet-Ast 1953). *** Colura inlata K.I.Goebel, Ann. Jard. Bot. Buitenzorg 39: 11, 1928 (Goebel 1928). *** Colura jovet-astiae Grolle, J. Hattori Bot. Lab. 28: 44, 1965 (Grolle 1965b). Nom. nov. pro Colura undulata Jovet-Ast, Rev. Bryol. Lichénol. 30 (1/2): 7, 1961 (Jovet-Ast 1961), nom. illeg. *** Colura meijeri Jovet-Ast, Rev. Bryol. Lichénol. 22 (2/3): 290, 1953 [1954] (Jovet-Ast 1953). *** Colura obvoluta Jovet-Ast, Cryptog. Bryol. Lichénol. 4 (3): 207, 1983 (Jovet-Ast 1983). *** Colura ornata K.I.Goebel, Ann. Jard. Bot. Buitenzorg 9 (1): 26, 1890 [1891] (Goebel 1890). *** Colura palawanensis Jovet-Ast, Rev. Bryol. Lichénol. 22 (2/3): 305, 1953 [1954] (Jovet-Ast 1953). *** Colura valida Jovet-Ast, Rev. Bryol. Lichénol. 30 (1/2): 6, 1961 (Jovet-Ast 1961). *** Colura verdoornii Herzog et Jovet-Ast, Rev. Bryol. Lichénol. 22 (2/3): 288, 1953 [1954] (Jovet-Ast 1953). ** sect. Harmophyllum Grolle, J. Hattori Bot. Lab. 28: 44, 1965 (Grolle 1965b). *** Colura andoi Gradst. et Jovet-Ast, Hikobia 9: 355, 1986 (Gradstein 1986). 342 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Colura ari (Steph.) Steph., Sp. Hepat. (Stephani) 5: 936, 1916 (Stephani 1916b). Bas.: Colurolejeunea ari Steph., Hedwigia 35 (3): 73, 1896 (Stephani 1896b). *** Colura australiensis Jovet-Ast, Rev. Bryol. Lichénol. 22 (2/3): 260, 1953 [1954] (Jovet-Ast 1953). *** Colura brevistyla Herzog, Beih. Bot. Centralbl. 38 (2): 331, 1921 (Herzog 1921). *** Colura calderae Pócs, J. Bryol. 29 (2): 84, 2007 (Müller and Pócs 2007). *** Colura clementis Grolle, J. Hattori Bot. Lab. 28: 45, 1965 (Grolle 1965b). *** Colura conica (Sande Lac.) K.I.Goebel, Ann. Jard. Bot. Buitenzorg 39: 3, 1928 (Goebel 1928). Bas.: Lejeunea conica Sande Lac., Ann. Mus. Bot. Lugduno-Batavi 1: 311, 1864 (Sande Lacoste 1864). *** Colura corynophora (Nees, Lindenb. et Gottsche) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 402, 1877 (Trevisan 1877). Bas.: Lejeunea corynophora Nees, Lindenb. et Gottsche, Observ. bot.: 474, 1843 (Gottsche et al. 1843). *** Colura crenulata Grolle, J. Hattori Bot. Lab. 28: 46, 1965 (Grolle 1965b). *** Colura crispiloba Jovet-Ast, Cryptog. Bryol. Lichénol. 4 (3): 205, 1983 (Jovet-Ast 1983). ** Colura cylindrica Herzog, Svensk Bot. Tidskr. 46 (1): 106, 1952 (Herzog 1952e). *** Colura cymbalifera Herzog et Jovet-Ast, Rev. Bryol. Lichénol. 22 (2/3): 268, 1953 [1954] (Jovet-Ast 1953). *** Colura digitalis (Mitt.) Steph., Sp. Hepat. (Stephani) 5: 931, 1916 (Stephani 1916b). Bas.: Lejeunea digitalis Mitt., J. Linn. Soc., Bot. 22 (146): 325, 1886 (Mitten 1886b). ** Colura digitalis var. mucronata Pócs, Fragm. Florist. Geobot. 40 (1): 263, 1995 (Pócs 1995). *** Colura dusenii Steph., Sp. Hepat. (Stephani) 5: 931, 1916 (Stephani 1916b). Based on: Colurolejeunea dusenii Steph., Hedwigia 31 (4): 168, 1892 (Stephani 1892g). *** Colura fastigiata Jovet-Ast, Rev. Bryol. Lichénol. 27 (1/2): 28, 1958 (Jovet-Ast 1958). *** Colura istulosa Jovet-Ast, Cryptog. Bryol. Lichénol. 4 (3): 211, 1983 (Jovet-Ast 1983). *** Colura hattoriana Pócs, J. Hattori Bot. Lab. 74: 47, 1993 (Pócs 1993). *** Colura heimii Jovet-Ast, Rev. Bryol. Lichénol. 22 (2/3): 275, 1953 [1954] (Jovet-Ast 1953). *** Colura hemisphaerica Jovet-Ast, Rev. Bryol. Lichénol. 22 (2/3): 267, 1953 [1954] (Jovet-Ast 1953). *** Colura herzogii Jovet-Ast, Rev. Bryol. Lichénol. 22 (2/3): 261, 1953 [1954] (JovetAst 1953). *** Colura inuii Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 1: 68, 1931 (Horikawa 1931a). ** Colura koponenii Pócs, Chenia 11: 29, 2013 (Pócs 2013). ** Colura mauritiana Pócs, Cryptog. Bryol. Lichénol. 18 (3): 196, 1997 (Pócs 1997a). *** Colura maxima Jovet-Ast, Rev. Bryol. Lichénol. 22 (2/3): 284, 1953 [1954] (Jovet-Ast 1953). *** Colura mosenii Steph., Sp. Hepat. (Stephani) 5: 940, 1916 (Stephani 1916b). ** Colura norrisii Pócs, Chenia 11: 29, 2013 (Pócs 2013). World checklist of hornworts and liverworts 343 *** Colura obesa Jovet-Ast, Rev. Bryol. Lichénol. 22 (2/3): 273, 1953 [1954] (JovetAst 1953). *** Colura pallida Steph., Sp. Hepat. (Stephani) 5: 941, 1916 (Stephani 1916b). *** Colura pluridentata Jovet-Ast, Rev. Bryol. Lichénol. 22 (2/3): 265, 1953 [1954] (Jovet-Ast 1953). *** Colura speciosa Jovet-Ast, Rev. Bryol. Lichénol. 22 (2/3): 307, 1953 [1954] (Jovet-Ast 1953). ** Colura streimannii Pócs, Polish Bot. J. 60 (1): 7, 2015 (Pócs 2015b). *** Colura superba (Mont.) Steph., Sp. Hepat. (Stephani) 5: 941, 1916 (Stephani 1916b). Bas.: Lejeunea superba Mont., Ann. Sci. Nat. Bot. (sér. 3) 10: 115, 1848 (Montagne 1848). *** Colura thomeensis Pócs, Bryologist 114 (2): 363, 2011 (Pócs 2011b). *** Colura tortifolia (Nees et Mont.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 402, 1877 (Trevisan 1877). Bas.: Lejeunea tortifolia Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 265, 1843 (Montagne 1843). *** Colura tutuilana (Pearson) H.A.Mill., Phytologia 47 (4): 322, 1981 (Miller 1981). Bas.: Colurolejeunea tutuilana Pearson, Amer. Samoa: 151, 1924 (Pearson 1924a). *** Colura ulei Jovet-Ast, Rev. Bryol. Lichénol. 22 (2/3): 270, 1953 [1954] (Jovet-Ast 1953). *** Colura vitiensis Pócs et J.Eggers, Polish Bot. J. 52 (2): 88, 2007 (Pócs and Eggers 2007). ** sect. Oidocorys Jovet-Ast ex Grolle, J. Hattori Bot. Lab. 32: 140, 1969 (Grolle 1969c). *** Colura itatyana Steph., Sp. Hepat. (Stephani) 5: 932, 1916 (Stephani 1916b). *** Colura naumannii (Schifn.) Steph., Sp. Hepat. (Stephani) 5: 935, 1916 (Stephani 1916b). Bas.: Colurolejeunea naumannii Schifn., Hepat. (Engl.-Prantl): 121, 1893 (Schifner 1893b). *** Colura ornithocephala Herzog, Svensk Bot. Tidskr. 46 (1): 107, 1952 (Herzog 1952e). *** Colura pulcherrima Jovet-Ast, Rev. Bryol. Lichénol. 22 (2/3): 235, 1953 [1954] (Jovet-Ast 1953). ** Colura pulcherrima var. bartlettii Jovet-Ast, Cryptog. Bryol. Lichénol. 1 (3): 278, 1980 (Jovet-Ast 1980). *** Colura saccophylla E.A.Hodgs. et Herzog, Trans. & Proc. Roy. Soc. New Zealand 77 (2): 253, 1949 (Herzog 1949a). *** Colura schusteri Grolle, J. Hattori Bot. Lab. 32: 140, 1969 (Grolle 1969c). ** subg. Glotta Grolle et R.L.Zhu, J. Hattori Bot. Lab. 92: 187, 2002 (Grolle and Zhu 2002). ** sect. Glotta Grolle et R.L.Zhu, J. Hattori Bot. Lab. 92: 187, 2002 (Grolle and Zhu 2002). *** Colura bicornis Jovet-Ast, Rev. Bryol. Lichénol. 25 (3/4): 272, 1956 [1957] (Jovet-Ast 1956). *** Colura bisvoluta Herzog et Jovet-Ast, Rev. Bryol. Lichénol. 22 (2/3): 228, 1953 [1954] (Jovet-Ast 1953). 344 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Colura inornata Jovet-Ast, Rev. Bryol. Lichénol. 25 (3/4): 275, 1956 [1957] (Jovet-Ast 1956). *** Colura karstenii K.I.Goebel, Planzenbiol. Schilderungen 2 (1): 153, 1891 (Goebel 1891). *** Colura queenslandica B.M.hiers, Brittonia 39 (2): 175, 1987 (hiers 1987b). *** Colura saroltae Pócs, J. Bryol. 14 (3): 497, 1987 (Jones and Pócs 1987). *** Colura strophiolata Jovet-Ast, Rev. Bryol. Lichénol. 42 (4): 917, 1976 (Jovet-Ast 1976). *** Colura usambarica E.W.Jones, J. Bryol. 14 (3): 495, 1987 (Jones and Pócs 1987). ** sect. Heterophyllum Jovet-Ast, Cryptog. Bryol. Lichénol. 4 (3): 213, 1983 (JovetAst 1983). *** Colura acroloba (Prantl) Jovet-Ast, Rev. Bryol. Lichénol. 22 (2/3): 297, 1953 [1954] (Jovet-Ast 1953). Bas.: Lejeunea acroloba Prantl, Hedwigia 29: xiv, 1890 (Prantl 1890). *** Colura corniantha Grolle, J. Hattori Bot. Lab. 28: 44, 1965 (Grolle 1965b). Nom. nov. pro Colura cornuta Jovet-Ast, Rev. Bryol. Lichénol. 30 (1/2): 11, 1961 (Jovet-Ast 1961), nom. illeg. ** Colura denticulata Jovet-Ast, Rev. Bryol. Lichénol. 23 (1/2): 2, 1954 (Jovet-Ast 1954). *** Colura galeata Jovet-Ast, Rev. Bryol. Lichénol. 35 (1/4): 146, 1967 [1968] (JovetAst 1967a). ** Colura imperfecta Steph., Sp. Hepat. (Stephani) 5: 938, 1916 (Stephani 1916b). *** Colura siamensis Jovet-Ast, Rev. Bryol. Lichénol. 35 (1/4): 139, 1967 [1968] (Jovet-Ast 1967b). *** Colura tixieri Jovet-Ast, Rev. Bryol. Lichénol. 30 (1/2): 9, 1961 (Jovet-Ast 1961). *** Colura vietnamensis Jovet-Ast et Tixier, Rev. Bryol. Lichénol. 27 (3/4): 205, 1958 [1959] (Jovet-Ast and Tixier 1958). Incertae sedis ** Colura amboinensis Steph., Sp. Hepat. (Stephani) 5: 935, 1916 (Stephani 1916b). *** Colura hirta Steph., Sp. Hepat. (Stephani) 5: 932, 1916 (Stephani 1916b). *** Diplasiolejeunea (Spruce) Schifn., Hepat. (Engl.-Prantl): 121, 1893 (Schifner 1893b). Bas.: Lejeunea subg. Diplasiolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 301, 1884 (Spruce 1884). 289 *** Diplasiolejeunea utriculata Steph., Sp. Hepat. (Stephani) 5: 920, 1916 (Stephani 1916b). ** subg. Austrolejeuneopsis R.M.Schust., Bull. Torrey Bot. Club 97 (6): 349, 1970 (Schuster 1970b). *** Diplasiolejeunea alata Jovet-Ast, Rev. Bryol. Lichénol. 17 (1/4): 31, 1948 [1949] (Jovet-Ast 1948). 289 Diplasiolejeunea is here only provisionally accepted with subgenera following Schuster (1970b), but they are not supported by the molecular phylogeny by Dong et al. (2012) where the type species of subg. Austrolejeuneopsis and Physolejeunea are nested in subg. Diplasiolejeunea. World checklist of hornworts and liverworts 345 *** Diplasiolejeunea eggersii Pócs, Bryologist 109 (3): 408, 2006 (Pócs 2006b). *** Diplasiolejeunea erostrata Schäf.-Verw., Cryptog. Bryol. 25 (1): 3, 2004 (SchäferVerwimp 2004). * Diplasiolejeunea guadalupensis Steph., Sp. Hepat. (Stephani) 5: 924, 1916 (Stephani 1916b). 290 * Diplasiolejeunea heimii Jovet-Ast, Rev. Bryol. Lichénol. 29 (1/2): 39, 1960 (Jovet-Ast 1960). 291 ** Diplasiolejeunea involuta S.Winkl., Rev. Bryol. Lichénol. 35 (1/4): 320, 1967 [1968] (Winkler 1967). ** Diplasiolejeunea involuta subsp. andicola Pócs, Cryptog. Bryol. Lichénol. 19 (1): 12, 1998 (León et al. 1998). *** Diplasiolejeunea johnsonii A.Evans, Bull. Torrey Bot. Club 39 (12): 603, 1912 [1913] (Evans 1912d). ** Diplasiolejeunea johnsonii var. mexicana Jovet-Ast, Rev. Bryol. Lichénol. 29 (1/2): 39, 1960 (Jovet-Ast 1960). * Diplasiolejeunea montecristensis S.Winkl., Rev. Bryol. Lichénol. 35 (1/4): 321, 1967 [1968] (Winkler 1967). 292 *** Diplasiolejeunea papilionacea R.M.Schust., Phytologia 39 (6): 431, 1978 (Schuster 1978b). *** Diplasiolejeunea pauckertii (Nees) Steph., Sp. Hepat. (Stephani) 5: 924, 1916 (Stephani 1916b). Bas.: Lejeunea pauckertii Nees, Syn. Hepat. 3: 392, 1845 (Gottsche et al. 1845b). *** Diplasiolejeunea pluridentata Schäf.-Verw., Haussknechtia 8: 71, 2001 (SchäferVerwimp 2001b). *** Diplasiolejeunea replicata (Spruce) Steph., Sp. Hepat. (Stephani) 5: 926, 1916 (Stephani 1916b). Bas.: Lejeunea replicata Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 302, 1884 (Spruce 1884). ** Diplasiolejeunea rudolphiana Steph., Hedwigia 35 (3): 79, 1896 (Stephani 1896b). 293 *** Diplasiolejeunea runssorensis Steph., Bot. Jahrb. Syst. 20 (3): 318, 1895 (Stephani 1895a). *** Diplasiolejeunea unidentata (Lehm. et Lindenb.) Schifn., Bot. Jahrb. Syst. 23 (5): 583, 1897 (Schifner 1897). Bas.: Jungermannia unidentata Lehm. et Lindenb., Nov. Stirp. Pug. 6: 48, 1834 (Lehmann 1834). ** subg. Diplasiolejeunea *** Diplasiolejeunea armatiloba Steph., Hedwigia 35 (3): 80, 1896 (Stephani 1896b). 290 Diplasiolejeunea guadalupensis is a poorly known taxon with some similarities to the Diplasiolejeunea replicata species complex (Schäfer-Verwimp and Reiner-Drehwald 2009). 291 Diplasiolejeunea heimii is possibly conspeciic with Diplasiolejeunea replicata (Schäfer-Verwimp and Pócs 2009). 292 Diplasiolejeunea montecristensis is possibly conspeciic with Diplasiolejeunea replicata. 293 Diplasiolejeunea rudolphiana and Diplasiolejeunea unidentata are genetically very close and may be conspeciic (Dong et al. 2012). 346 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Diplasiolejeunea borhidiana Reyes Montoya, Acta Bot. Acad. Sci. Hung. 28 (1/2): 177, 1982 [1983] (Reyes 1982). *** Diplasiolejeunea brunnea Steph., Sp. Hepat. (Stephani) 5: 922, 1916 (Stephani 1916b). ** Diplasiolejeunea caribea Tixier, Bryophyt. Biblioth. 27: 377, 1985 (Tixier 1985a). *** Diplasiolejeunea cavifolia Steph., Sp. Hepat. (Stephani) 5: 918, 1916 (Stephani 1916b). Nom. nov. pro Lejeunea cavifolia Steph., Bot. Jahrb. Syst. 8 (2): 89, 1886 (Stephani 1886d), nom. illeg. *** Diplasiolejeunea cobrensis Steph., Sp. Hepat. (Stephani) 5: 923, 1916 (Stephani 1916b). ** Diplasiolejeunea cobrensis subsp. antsirananae Pócs, Candollea 56 (1): 72, 2001 (Pócs 2001). ** Diplasiolejeunea cogoensis M.Infante, Heras et Pócs, Trop. Bryol. 17: 9, 1999 (Infante et al. 1999). ** Diplasiolejeunea cubensis Tixier, Bryophyt. Biblioth. 27: 389, 1985 (Tixier 1985a). * Diplasiolejeunea glaziovii Tixier, Bryophyt. Biblioth. 27: 397, 1985 (Tixier 1985a). 294 *** Diplasiolejeunea ingekarolae Schäf.-Verw., Herzogia 19: 239, 2006 (Schäfer-Verwimp 2006). ** Diplasiolejeunea jonesii Tixier, Acta Bot. Hung. 30 (1/2): 21, 1984 (Tixier 1984). *** Diplasiolejeunea leiocarpa Jovet-Ast, Rev. Bryol. Lichénol. 16 (1/2): 34, 1947 [1948] (Jovet-Ast 1947b). *** Diplasiolejeunea malleiformis (A.Evans) Tixier, Bryophyt. Biblioth. 27: 351, 1985 (Tixier 1985a). Bas.: Diplasiolejeunea pellucida var. malleiformis A.Evans, Bull. Torrey Bot. Club 39 (5): 215, 1912 (Evans 1912c). ** Diplasiolejeunea palustrium Tixier, Acta Bot. Hung. 30 (1/2): 19, 1984 (Tixier 1984). *** Diplasiolejeunea pellucida (C.F.W.Meissn. ex Spreng.) Schifn., Hepat. (Engl.Prantl): 121, 1893 (Schifner 1893b). Bas.: Jungermannia pellucida C.F.W.Meissn. ex Spreng. Syst. Veg. (ed. 16) [Sprengel] 4 (2): 325, 1827 (Sprengel 1827b). ** Diplasiolejeunea phyllarthronii Tixier, Acta Bot. Hung. 30 (1/2): 19, 1984 (Tixier 1984). *** Diplasiolejeunea pocsii Reyes Montoya, Acta Bot. Acad. Sci. Hung. 28 (1/2): 173, 1982 [1983] (Reyes 1982). ** subg. Physolejeunea R.M.Schust., Bull. Torrey Bot. Club 97 (6): 348, 1970 (Schuster 1970b). *** Diplasiolejeunea andringitrae Schäf.-Verw., Cryptog. Bryol. 27 (4): 447, 2006 (Pócs and Schäfer-Verwimp 2006). *** Diplasiolejeunea cornuta Steph., Sp. Hepat. (Stephani) 5: 918, 1916 (Stephani 1916b). *** Diplasiolejeunea jovet-astiae Grolle, Feddes Repert. 73 (2): 84, 1966 (Grolle 1966b). *** Diplasiolejeunea kraussiana (Lindenb.) Steph., Sp. Hepat. (Stephani) 5: 919, 1916 (Stephani 1916b). Bas.: Lejeunea kraussiana Lindenb., Syn. Hepat. 3: 393, 1845 (Gottsche et al. 1845b). *** Diplasiolejeunea ornata Pócs et Schäf.-Verw., Cryptog. Bryol. 27 (4): 440, 2006 (Pócs and Schäfer-Verwimp 2006). 294 Diplasiolejeunea glaziovii may be conspeciic with Diplasiolejeunea cavifolia (Dong et al. 2012). World checklist of hornworts and liverworts 347 *** Diplasiolejeunea patelligera Herzog, Svensk Bot. Tidskr. 42 (3): 240, 1948 (Herzog 1948). *** Diplasiolejeunea plicatiloba (Hook.f. et Taylor) Grolle, J. Hattori Bot. Lab. 45: 175, 1979 (Grolle 1979c). Bas.: Jungermannia plicatiloba Hook.f. et Taylor, London J. Bot. 4: 92, 1845 (Hooker and Taylor 1845). *** Diplasiolejeunea pusilla Grolle, Feddes Repert. 86 (1/2): 79, 1975 (Grolle 1975a). *** Diplasiolejeunea ranomafanae Pócs, Cryptog. Bryol. 27 (4): 444, 2006 (Pócs and Schäfer-Verwimp 2006). *** Diplasiolejeunea villaumei Steph., Sp. Hepat. (Stephani) 5: 921, 1916 (Stephani 1916b). *** Diplasiolejeunea zakiae Tixier, Lindbergia 4 (1/2): 123, 1977 (Tixier 1977b). *** *** ** *** ** * ** *** ** * ** *** *** ** ** * *** Incertae sedis Diplasiolejeunea albifolia (Taylor) E.W.Jones, Trans. Brit. Bryol. Soc. 2 (3): 393, 1954 (Jones 1954a). Bas.: Lejeunea albifolia Taylor, London J. Bot. 5: 399, 1846 (Taylor 1846b). Diplasiolejeunea aulae E.W.Jones, J. Bryol. 7 (4): 552, 1973 [1974] (Jones 1973). Diplasiolejeunea auriculata Tixier, Rev. Bryol. Lichénol. 45 (2): 210, 1979 (Tixier 1979c). Diplasiolejeunea buckii Grolle, Beitr. Phytotax. 15: 105, 1992 (Grolle 1992a). Diplasiolejeunea columbica Tixier, Cryptog. Bryol. Lichénol. 4 (3): 233, 1983 (Tixier 1983a). Diplasiolejeunea comorensis Tixier, Rev. Bryol. Lichénol. 45 (2): 213, 1979 (Tixier 1979c). Diplasiolejeunea cyanguguensis Tixier, Trop. Bryol. 11: 67, 1995 (Tixier 1995b). Diplasiolejeunea deslooveri Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 47 (1/2): 217, 1977 (Vanden Berghen 1977). Diplasiolejeunea ensifera Tixier, Rev. Bryol. Lichénol. 45 (2): 215, 1979 (Tixier 1979c). Diplasiolejeunea evansii Tixier, Bryophyt. Biblioth. 27: 360, 1985 (Tixier 1985a). 295 Diplasiolejeunea gradsteinii Tixier, Trop. Bryol. 11: 67, 1995 (Tixier 1995b). Diplasiolejeunea grandirostrata Schäf.-Verw., Cryptog. Bryol. 25 (1): 7, 2004 (Schäfer-Verwimp 2004). Diplasiolejeunea grolleana Reyes Montoya, Acta Bot. Acad. Sci. Hung. 28 (1/2): 175, 1982 [1983] (Reyes 1982). Diplasiolejeunea hamata Tixier, Rev. Bryol. Lichénol. 45 (2): 217, 1979 (Tixier 1979c). Diplasiolejeunea insignis Tixier, Lindbergia 4 (1/2): 120, 1977 (Tixier 1977b). Diplasiolejeunea integerrima Tixier, Rev. Bryol. Lichénol. 45 (2): 219, 1979 (Tixier 1979c). 296 Diplasiolejeunea lanceolata Grolle, Beitr. Phytotax. 15: 107, 1992 (Grolle 1992a). 295 Diplasiolejeunea evansii is a doubtful taxon and the type specimen could not be found in PC (SchäferVerwimp 2004). 296 Diplasiolejeunea integerrima is a juvenile Acrolejeunea species, but the type is too poor to allow identiication (Grolle 1995). 348 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Diplasiolejeunea latipuensis Tixier, Cryptog. Bryol. Lichénol. 16 (3): 230, 1995 (Tixier 1995a). Based on: Diplasiolejeunea latipuensis Tixier, Candollea 46 (2): 294, 1991 (Tixier 1991), nom. inval. ** Diplasiolejeunea lemuriana Tixier, Lindbergia 4 (1/2): 120, 1977 (Tixier 1977b). ** Diplasiolejeunea longilobula Herzog, Trans. Brit. Bryol. Soc. 1 (4): 325, 1950 (Herzog 1950a). ** Diplasiolejeunea magnistipula Tixier, Ann. Fac. Sci. Yaoundé 20: 7, 1975 (Tixier 1975b). *** Diplasiolejeunea mayaykuensis Schäf.-Verw. et Heinrichs, Polish Bot. J. 58 (1): 143, 2013 (Schäfer-Verwimp et al. 2013). *** Diplasiolejeunea onraedtii Grolle, Feddes Repert. 89 (5/6): 301, 1978 (Grolle 1978a). ** Diplasiolejeunea ramicola Tixier, Rev. Bryol. Lichénol. 45 (2): 222, 1979 (Tixier 1979c). ** Diplasiolejeunea riclefgrollei Schäf.-Verw., Cryptog. Bryol. 26 (1): 37, 2005 (Schäfer-Verwimp 2005). ** Diplasiolejeunea subcornuta Tixier, Lindbergia 4 (1/2): 122, 1977 (Tixier 1977b). *** Diplasiolejeunea symoensii Vanden Berghen, Bull. Soc. Roy. Bot. Belgique 92: 126, 1960 (Vanden Berghen 1960b). ** Haplolejeunea Grolle, J. Hattori Bot. Lab. 39: 205, 1975 (Grolle 1975b). *** Haplolejeunea cucullata (Steph.) Grolle, J. Hattori Bot. Lab. 45: 176, 1979 (Grolle 1979c). Bas.: Cheilolejeunea cucullata Steph., Sp. Hepat. (Stephani) 5: 644, 1914 (Stephani 1914b). ** Haplolejeunea sticta Grolle, J. Hattori Bot. Lab. 39: 205, 1975 (Grolle 1975b). ** Macrocolura R.M.Schust., J. Hattori Bot. Lab. 75: 233, 1994 (Schuster 1994). *** Macrocolura sagittistipula (Spruce) R.M.Schust., J. Hattori Bot. Lab. 75: 233, 1994 (Schuster 1994). Bas.: Lejeunea sagittistipula Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 304, 1884 (Spruce 1884). *** Myriocoleopsis Schifn., Hedwigia 81 (5/6): 234, 1944 (Schifner 1944). *** Myriocoleopsis luviatilis (Steph.) M.E.Reiner et Gradst., J. Bryol. 19 (3): 639, 1997 (Reiner-Drehwald and Gradstein 1997). Bas.: Cololejeunea luviatilis Steph., Hedwigia 34 (5): 248, 1895 (Stephani 1895b). *** Myriocoleopsis gymnocolea (Spruce) M.E.Reiner et Gradst., J. Bryol. 19 (3): 640, 1997 (Reiner-Drehwald and Gradstein 1997). Bas.: Lejeunea gymnocolea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 296, 1884 (Spruce 1884). *** Myriocoleopsis minutissima (Sm.) R.L.Zhu, Y.Yu et Pócs, Phytotaxa 183 (4): 293, 2014 (Yu et al. 2014). Bas.: Jungermannia minutissima Sm., Engl. Bot. 23: tab. 1633, 1806 (Smith and Sowerby 1806). *** Myriocoleopsis minutissima subsp. myriocarpa (Nees et Mont.) R.L.Zhu, Y.Yu et Pócs, Phytotaxa 183 (4): 294, 2014 (Yu et al. 2014). Bas.: Lejeunea myriocarpa Nees et Mont., Hist. Phys. Cuba, Bot., Pl. Cell.: 473, 1842 (Montagne 1842a). World checklist of hornworts and liverworts 349 ** Myriocoleopsis vuquangensis (Pócs et Ninh) Pócs, Trop. Bryol. 31: 124, 2010 (Pócs 2010a). Bas.: Cololejeunea vuquangensis Pócs et Ninh, Acta Bot. Hung. 47 (1/2): 156, 2005 (Pócs and Ninh 2005). ** Nephelolejeunea Grolle, J. Hattori Bot. Lab. 37: 252, 1973 (Grolle 1973b). ** Nephelolejeunea bidentata B.M.hiers ex L.Söderstr. et A.Hagborg, Phytotaxa 202 (1): 65, 2015 (Pócs et al. 2015c). Based on: Austrolejeunea bidentata B.M.hiers, Bryologist 88 (4): 350, 1985 [1986] (hiers 1985), nom. inval. ** Nephelolejeunea carcharias M.A.M.Renner, Syst. Bot. 34 (4): 621, 2009 (Renner et al. 2009). *** Nephelolejeunea conchophylla Grolle, J. Hattori Bot. Lab. 48: 164, 1980 (Grolle 1980a). ** Nephelolejeunea fragilis (R.M.Schust.) L.Söderstr. et A.Hagborg, Phytotaxa 202 (1): 65, 2015 (Pócs et al. 2015c). Bas.: Cololejeunea fragilis R.M.Schust., Phytologia 56 (7): 458, 1985 (Schuster 1985c). *** Nephelolejeunea hamata Grolle, J. Hattori Bot. Lab. 48: 167, 1980 (Grolle 1980a). *** Nephelolejeunea hispida R.M.Schust. ex L.Söderstr. et A.Hagborg, Phytotaxa 202 (1): 66, 2015 (Pócs et al. 2015c). Based on: Austrolejeunea hispida R.M.Schust., Phytologia 47 (4): 305, 1981 (Schuster 1981a), nom. inval. *** Nephelolejeunea jarmaniana Grolle ex L.Söderstr. et A.Hagborg, Phytotaxa 202 (1): 66, 2015 (Pócs et al. 2015c). Based on: Austrolejeunea jarmaniana Grolle, Nova Hedwigia 55 (1/2): 112, 1992 (Grolle 1992b), nom. inval. *** Nephelolejeunea nudipes (Hook.f. et Taylor) L.Söderstr. et A.Hagborg, Phytotaxa 202 (1): 66, 2015 (Pócs et al. 2015c). Bas.: Jungermannia nudipes Hook.f. et Taylor, London J. Bot. 3: 568, 1844 (Hooker and Taylor 1844d). ** Nephelolejeunea occidentalis Pócs ex L.Söderstr. et A.Hagborg, Phytotaxa 202 (1): 66, 2015 (Pócs et al. 2015c). Based on: Austrolejeunea occidentalis Pócs, J. Hattori Bot. Lab. 99: 187, 2006 (Pócs 2006d), nom. inval. *** Nephelolejeunea papillosa Glenny, New Zealand J. Bot. 34 (2): 195, 1996 (Glenny 1996). *** Nephelolejeunea radulifolia (C.Massal.) L.Söderstr. et A.Hagborg, Phytotaxa 202 (1): 66, 2015 (Pócs et al. 2015c). Bas.: Lejeunea radulifolia C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 248, 1885 (Massalongo 1885). ** Nephelolejeunea secunda M.A.M.Renner ex L.Söderstr. et A.Hagborg, Phytotaxa 202 (1): 66, 2015 (Pócs et al. 2015c). Based on: Austrolejeunea secunda M.A.M.Renner, Bryologist 113 (4): 782, 2010 (Renner 2010), nom. inval. *** Nephelolejeunea talinayi (S.W.Arnell) Grolle, J. Hattori Bot. Lab. 37: 253, 1973 (Grolle 1973b). Bas.: Harpalejeunea talinayi S.W.Arnell, Svensk Bot. Tidskr. 50 (2): 309, 1956 (Arnell 1956d). ** Schusterolejeunea Grolle, J. Bryol. 11 (1): 105, 1980 (Grolle 1980b). Nom. nov. pro Cladocolea R.M.Schust., Beih. Nova Hedwigia 9: 155, 1963 (Schuster 1963a). 350 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Schusterolejeunea inundata (Spruce) Grolle, J. Bryol. 11 (1): 105, 1980 (Grolle 1980b). Bas.: Lejeunea inundata Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 278, 1884 (Spruce 1884). ** Siphonolejeunea Herzog, Nat. Hist. Juan Fernandez (Botany) 2 (5): 744, 1942 (Herzog 1942a). ** Siphonolejeunea elegantissima (Steph.) Grolle, J. Hattori Bot. Lab. 41: 405, 1976 (Grolle 1976b). Bas.: Trachylejeunea elegantissima Steph., Hedwigia 28 (4): 262, 1889 (Stephani 1889c). ** Siphonolejeunea neesii (Mont.) Bischl., Nova Hedwigia 17: 338, 1969 (Bischler 1969). Bas.: Lejeunea neesii Mont., Ann. Sci. Nat. Bot. (sér. 2) 5: 62, 1836 (Nees and Montagne 1836). *** Siphonolejeunea schifneri (Schifn.) Herzog, Svensk Bot. Tidskr. 42 (3): 230, 1948 (Herzog 1948). Bas.: Pycnolejeunea schifneri Schifn., Consp. Hepat. Arch. Ind.: 260, 1898 (Schifner 1898b). ** Tuyamaella S.Hatt., J. Hattori Bot. Lab. 5: 60, 1951 (Hattori 1951d). ** Tuyamaella angulistipa (Steph.) R.M.Schust. et Kachroo, J. Linn. Soc., Bot. 56 (368): 508, 1961 (Kachroo and Schuster 1961). Bas.: Pycnolejeunea angulistipa Steph., Hedwigia 35 (3): 123, 1896 (Stephani 1896b). ** Tuyamaella borneensis Tixier, Rev. Bryol. Lichénol. 39 (2): 241, 1973 (Tixier 1973a). ** Tuyamaella hattorii Tixier, Rev. Bryol. Lichénol. 31 (3/4): 188, 1962 [1963] (Tixier 1962). ** Tuyamaella jackii (Steph.) Tixier, Rev. Bryol. Lichénol. 39 (2): 235, 1973 (Tixier 1973a). Bas.: Pycnolejeunea jackii Steph., Sp. Hepat. (Stephani) 5: 611, 1914 (Stephani 1914b). *** Tuyamaella molischii (Schifn.) S.Hatt., J. Hattori Bot. Lab. 5: 62, 1951 (Hattori 1951d). Bas.: Pycnolejeunea molischii Schifn., Ann. Bryol. 2: 97, 1929 (Schifner 1929). ** Tuyamaella molischii var. brevistipa P.C.Wu et P.J.Lin, Acta Phytotax. Sin. 16 (2): 65, 1978 (Wu and Lin 1978). ** Tuyamaella molischii var. taiwanensis R.L.Zhu et M.L.So, Nova Hedwigia 70 (1/2): 190, 2000 (Zhu and So 2000). ** Tuyamaella serratistipa S.Hatt., Bot. Mag. (Tokyo) 64 (755/756): 118, 1951 (Hattori 1951c). ** subtrib. Cyclolejeuneinae Gradst. *** Bromeliophila R.M.Schust., J. Hattori Bot. Lab. 75: 226, 1994 (Schuster 1994). *** Bromeliophila helenae Gradst., Cryptog. Bryol. Lichénol. 18 (3): 218, 1997 (Gradstein 1997). World checklist of hornworts and liverworts 351 *** Bromeliophila natans (Steph.) R.M.Schust., J. Hattori Bot. Lab. 75: 226, 1994 (Schuster 1994). Bas.: Peltolejeunea natans Steph., Hedwigia 44 (4): 228, 1905 (Stephani 1905a). *** Cyclolejeunea A.Evans, Bull. Torrey Bot. Club 31 (4): 193, 1904 (Evans 1904a). *** Cyclolejeunea accedens (Gottsche) A.Evans, Bull. Torrey Bot. Club 31 (4): 201, 1904 (Evans 1904a). Bas.: Lejeunea accedens Gottsche, Syn. Hepat. 3: 339, 1845 (Gottsche et al. 1845b). *** Cyclolejeunea chitonia (Taylor) A.Evans, Bull. Torrey Bot. Club 31 (4): 194, 1904 (Evans 1904a). Bas.: Lejeunea chitonia Taylor, Nov. Stirp. Pug. 8: 27, 1844 (Lehmann 1844). *** Cyclolejeunea convexistipa (Lehm. et Lindenb.) A.Evans, Bull. Torrey Bot. Club 31 (4): 198, 1904 (Evans 1904a). Bas.: Jungermannia convexistipa Lehm. et Lindenb., Nov. Stirp. Pug. 6: 43, 1834 (Lehmann 1834). * Cyclolejeunea ecuadorensis Steph., Sp. Hepat. (Stephani) 5: 194, 1913 (Stephani 1913a). *** Cyclolejeunea foliorum (Nees) Grolle, J. Hattori Bot. Lab. 65: 403, 1988 (Grolle 1988c). Bas.: Lejeunea foliorum Nees, Syn. Hepat. 3: 326, 1845 (Gottsche et al. 1845b). * Cyclolejeunea integerrima (Steph.) Steph., Sp. Hepat. (Stephani) 5: 195, 1913 (Stephani 1913a). Bas.: Odontolejeunea integerrima Steph., Hedwigia 44 (4): 227, 1905 (Stephani 1905a). *** Cyclolejeunea luteola (Spruce) Grolle, Wiss. Z. Friedrich-Schiller-Univ. Jena, Math.-Naturwiss. Reihe 33: 761, 1984 (Grolle 1984b). Bas.: Lejeunea luteola Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 205, 1884 (Spruce 1884). *** Cyclolejeunea peruviana (Lehm. et Lindenb.) A.Evans, Bull. Torrey Bot. Club 31 (4): 196, 1904 (Evans 1904a). Bas.: Jungermannia peruviana Lehm. et Lindenb., Nov. Stirp. Pug. 5: 18, 1833 (Lehmann 1833). * Cyclolejeunea spectabilis Steph., Sp. Hepat. (Stephani) 5: 193, 1913 (Stephani 1913a). *** Prionolejeunea (Spruce) Schifn., Hepat. (Engl.-Prantl): 127, 1893 (Schifner 1893b). Bas.: Lejeunea subg. Prionolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 152, 1884 (Spruce 1884). *** Prionolejeunea aemula (Gottsche) A.Evans, Bull. Torrey Bot. Club 31 (4): 219, 1904 (Evans 1904a). Bas.: Lejeunea aemula Gottsche, Syn. Hepat. 3: 338, 1845 (Gottsche et al. 1845b). *** Prionolejeunea ampliretis Herzog, Feddes Repert. Spec. Nov. Regni Veg. 57 (1/2): 178, 1955 (Herzog 1955). *** Prionolejeunea arguta (Nees) Steph., Sp. Hepat. (Stephani) 5: 204, 1913 (Stephani 1913a). Bas.: Lejeunea arguta Nees, Syn. Hepat. 3: 338, 1845 (Gottsche et al. 1845b). *** Prionolejeunea decora (Taylor) Steph., Sp. Hepat. (Stephani) 5: 207, 1913 (Stephani 1913a). Bas.: Lejeunea decora Taylor, London J. Bot. 5: 393, 1846 (Taylor 1846b). *** Prionolejeunea denticulata (F.Weber) Schifn., Hepat. (Engl.-Prantl): 127, 1893 (Schifner 1893b). Bas.: Jungermannia denticulata F.Weber, Hist. Musc. Hepat. Prodr.: 30, 1815 (Weber 1815). 352 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Prionolejeunea diversitexta (Hampe et Gottsche) Steph., Sp. Hepat. (Stephani) 5: 209, 1913 (Stephani 1913a). Bas.: Lejeunea diversitexta Hampe et Gottsche, Linnaea 25 (3): 357, 1852 [1853] (Hampe and Gottsche 1852). *** Prionolejeunea exauriculata A.Evans, Bull. Torrey Bot. Club 31 (4): 223, 1904 (Evans 1904a). *** Prionolejeunea galliotii Steph., Sp. Hepat. (Stephani) 6: 388, 1923 (Stephani 1923). *** Prionolejeunea grata (Gottsche) Schifn., Hepat. (Engl.-Prantl): 127, 1893 (Schifner 1893b). Bas.: Lejeunea grata Gottsche, Abh. Naturwiss. Vereins Bremen 7: 359, 1882 (Gottsche 1882). *** Prionolejeunea grollei Ilk.-Borg. et Schäf.-Verw., Cryptog. Bryol. 26 (1): 29, 2005 (Ilkiu-Borges and Schäfer-Verwimp 2005). *** Prionolejeunea guadalupensis (Lindenb.) Steph., Sp. Hepat. (Stephani) 5: 211, 1913 (Stephani 1913a). Bas.: Lejeunea guadalupensis Lindenb., Syn. Hepat. 3: 340, 1845 (Gottsche et al. 1845b). *** Prionolejeunea limpida Herzog, Hedwigia 67 (6): 252, 1927 (Herzog 1927). *** Prionolejeunea magnistipula Herzog, Feddes Repert. Spec. Nov. Regni Veg. 57 (1/2): 176, 1955 (Herzog 1955). *** Prionolejeunea meissneri (Gottsche) Steph., Sp. Hepat. (Stephani) 5: 212, 1913 (Stephani 1913a). Bas.: Lejeunea meissneri Gottsche, Syn. Hepat. 3: 340, 1845 (Gottsche et al. 1845b). *** Prionolejeunea mucronata (Sande Lac.) Steph., Hedwigia 35 (3): 119, 1896 (Stephani 1896b). Bas.: Lejeunea mucronata Sande Lac., Syn. hepat. jav.: 106, 1856 [1857] (Sande Lacoste 1856b). *** Prionolejeunea muricatoserrulata (Spruce) Steph., Sp. Hepat. (Stephani) 5: 223, 1913 (Stephani 1913a). Bas.: Lejeunea muricatoserrulata Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 155, 1884 (Spruce 1884). *** Prionolejeunea principensis Vanden Berghen, Rev. Bryol. Lichénol. 29 (1/2): 65, 1960 (Vanden Berghen 1960a). *** Prionolejeunea recurvula (Spruce) Steph., Sp. Hepat. (Stephani) 5: 224, 1913 (Stephani 1913a). Bas.: Lejeunea recurvula Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 155, 1884 (Spruce 1884). *** Prionolejeunea scaberula (Spruce) Steph., Sp. Hepat. (Stephani) 5: 214, 1913 (Stephani 1913a). Bas.: Lejeunea scaberula Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 159, 1884 (Spruce 1884). *** Prionolejeunea schlimiana (Gottsche) Steph., Sp. Hepat. (Stephani) 5: 214, 1913 (Stephani 1913a). Bas.: Lejeunea schlimiana Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 154, 1864 (Gottsche 1864). *** Prionolejeunea trachyodes (Spruce) Steph., Sp. Hepat. (Stephani) 5: 215, 1913 (Stephani 1913a). Bas.: Lejeunea trachyodes Spruce, J. Linn. Soc., Bot. 30 (210): 338, 1895 (Gepp 1895b). World checklist of hornworts and liverworts * * ** 353 Excluded from the genus Prionolejeunea corbisieri Pearson, Natuurw. Tijdschr. 4 (5/6): 128, 1922 (Pearson 1922a). 297 Prionolejeunea maculata Herzog, Feddes Repert. Spec. Nov. Regni Veg. 57 (1/2): 180, 1955 (Herzog 1955). 298 subtrib. Drepanolejeuneinae Gradst. ** Capillolejeunea S.W.Arnell, Svensk Bot. Tidskr. 59 (1): 69, 1965 (Arnell 1965). ** Capillolejeunea geisslerae (Pócs) R.L.Zhu, Qiong He, Y.M.Wei et Pócs, Phytotaxa 175 (3): 167, 2014 (He et al. 2014a). Bas.: Drepanolejeunea geisslerae Pócs, Candollea 56 (1): 70, 2001 (Pócs 2001). ** Capillolejeunea mascarena S.W.Arnell, Svensk Bot. Tidskr. 59 (1): 69, 1965 (Arnell 1965). *** Drepanolejeunea (Spruce) Steph., Hedwigia 30 (5): 209, 1891 (Stephani 1891a). Bas.: Lejeunea subg. Drepanolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 186, 1884 (Spruce 1884). ** subg. Acantholejeunea R.M.Schust., Beih. Nova Hedwigia 9: 115, 1963 (Schuster 1963a). ** Drepanolejeunea dentistipula Steph., Sp. Hepat. (Stephani) 5: 343, 1913 (Stephani 1913a). ** Drepanolejeunea spinistipula Herzog, Svensk Bot. Tidskr. 42 (3): 238, 1948 (Herzog 1948). *** subg. Drepanolejeunea *** Drepanolejeunea aculeata Bischl., Rev. Bryol. Lichénol. 33 (1/2): 68, 1964 (Bischler 1964). ** Drepanolejeunea anderssonii (Ångstr.) A.Evans, Trans. Connecticut Acad. Arts 10 (8): 429, 1900 (Evans 1900a). Bas.: Lejeunea anderssonii Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 29 (4): 24, 1872 (Ångström 1872). *** Drepanolejeunea andina Herzog, Svensk Bot. Tidskr. 51 (1): 196, 1957 (Herzog 1957a). *** Drepanolejeunea angustifolia (Mitt.) Grolle, J. Jap. Bot. 40 (7): 206, 1965 (Grolle 1965d). Bas.: Lejeunea angustifolia Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 116, 1860 [1861] (Mitten 1860c). *** Drepanolejeunea ankasica E.W.Jones, Bull. Brit. Mus. (Nat. Hist.), Bot. 11 (3): 245, 1983 (Jones and Harrington 1983). 297 Prionolejeunea corbisieri is a Cheilolejeunea species (Ilkiu-Borges 2006). 298 Prionolejeunea maculata is a Cyclolejeunea species (Ilkiu-Borges 2006). 354 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Drepanolejeunea anoplantha (Spruce) Steph., Sp. Hepat. (Stephani) 5: 325, 1913 (Stephani 1913a). Bas.: Lejeunea anoplantha Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 189, 1884 (Spruce 1884). *** Drepanolejeunea appalachiana R.M.Schust., J. Elisha Mitchell Sci. Soc. 83 (4): 219, 1967 (Schuster 1967a). *** Drepanolejeunea araucariae Steph., Hedwigia 35 (3): 80, 1896 (Stephani 1896b). ** Drepanolejeunea araucariae var. chilensis Herzog, Acta Horti Gotob. 15: 161, 1943 (Herzog 1943a). ** Drepanolejeunea aucklandica Steph., Sp. Hepat. (Stephani) 5: 358, 1913 (Stephani 1913a). *** Drepanolejeunea aurita Bischl., Rev. Bryol. Lichénol. 33 (1/2): 146, 1964 (Bischler 1964). ** Drepanolejeunea bakeri Herzog, Ann. Bryol. 3: 148, 1930 (Herzog 1930b). *** Drepanolejeunea bidens (Prantl) A.Evans, Bull. Torrey Bot. Club 30 (1): 29, 1903 (Evans 1903b). Bas.: Lejeunea bidens Prantl, Hedwigia 29: xiv, 1890 (Prantl 1890). *** Drepanolejeunea biocellata A.Evans, Bull. Torrey Bot. Club 30 (1): 22, 1903 (Evans 1903b). ** Drepanolejeunea blumei Steph., Hedwigia 35 (3): 81, 1896 (Stephani 1896b). ** Drepanolejeunea brunnea Mizut., J. Hattori Bot. Lab. 33: 231, 1970 (Mizutani 1970). ** Drepanolejeunea caledonica Steph., Sp. Hepat. (Stephani) 5: 342, 1913 (Stephani 1913a). *** Drepanolejeunea campanulata (Spruce) Steph., Sp. Hepat. (Stephani) 5: 328, 1913 (Stephani 1913a). Bas.: Lejeunea campanulata Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 192, 1884 (Spruce 1884). ** Drepanolejeunea canceroides H.A.Mill. et Bonner, Beih. Nova Hedwigia 11: 53, 1963 (Miller et al. 1963). ** Drepanolejeunea capulata (Taylor) J.B.Jack et Steph., Hedwigia 31 (1): 13, 1892 (Jack and Stephani 1892). Bas.: Lejeunea capulata Taylor, London J. Bot. 5: 394, 1846 (Taylor 1846b). ** Drepanolejeunea capulata var. lagellifera S.W.Arnell, Österr. Akad. Wiss., Math.Naturwiss. Kl., Denkschr. 111: 100, 1964 (Schifner and Arnell 1964). ** Drepanolejeunea ciliata Mizut., J. Hattori Bot. Lab. 33: 232, 1970 (Mizutani 1970). *** Drepanolejeunea crassiretis A.Evans, Bull. Torrey Bot. Club 30 (1): 25, 1903 (Evans 1903b). *** Drepanolejeunea crucianella (Taylor) A.Evans, Bull. Torrey Bot. Club 30 (1): 33, 1903 (Evans 1903b). Bas.: Lejeunea crucianella Taylor, London J. Bot. 5: 393, 1846 (Taylor 1846b). *** Drepanolejeunea cultrella (Mitt.) Steph., Sp. Hepat. (Stephani) 5: 324, 1913 (Stephani 1913a). Bas.: Lejeunea cultrella Mitt., J. Proc. Linn. Soc., Bot. 7 (27): 168, 1863 (Mitten 1863). ** Drepanolejeunea cutervoensis (Loitl.) Grolle, J. Hattori Bot. Lab. 69: 186, 1991 (Grolle 1991). Bas.: Lejeunea cutervoensis Loitl., Diagn. pl. nov.: 21, 1894 (Loitlesberger 1894). World checklist of hornworts and liverworts 355 ** Drepanolejeunea decurviloba Steph., Sp. Hepat. (Stephani) 6: 397, 1923 (Stephani 1923). * Drepanolejeunea deslooveri Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 47 (1/2): 210, 1977 (Vanden Berghen 1977). 299 * Drepanolejeunea dissitifolia A.Evans, Bull. Torrey Bot. Club 30 (1): 28, 1903 (Evans 1903b). ** Drepanolejeunea elegans Herzog, Ann. Bryol. 9: 128, 1936 [1937] (Herzog 1936a). ** Drepanolejeunea erecta (Steph.) Mizut., J. Hattori Bot. Lab. 40: 442, 1976 (Mizutani 1976b). Bas.: Leptolejeunea erecta Steph., Bull. Soc. Roy. Bot. Belgique 38 (1): 44, 1899 (Stephani 1899h). ** Drepanolejeunea evansii Bischl. ex L.Söderstr., A.Hagborg et von Konrat, Phytotaxa 65: 47, 2012 (Söderström et al. 2012c). Based on: Drepanolejeunea evansii Bischl., Rev. Bryol. Lichénol. 33 (1/2): 75, 1964 (Bischler 1964), nom. inval. *** Drepanolejeunea fragilis Bischl. ex L.Söderstr., A.Hagborg et von Konrat, Phytotaxa 65: 47, 2012 (Söderström et al. 2012c). Based on: Drepanolejeunea fragilis Bischl., Rev. Bryol. Lichénol. 33 (1/2): 123, 1964 (Bischler 1964), nom. inval. ** Drepanolejeunea fulfordiae L.Söderstr., Phytotaxa 208 (1): 98, 2015 (Pócs et al. 2015a). Nom. nov. pro Drepanolejeunea papillosa Fulford, Mem. New York Bot. Gard. 23: 843, 1972 (Fulford 1972), nom. illeg. *** Drepanolejeunea granatensis (Prantl) Bischl., Rev. Bryol. Lichénol. 33 (1/2): 150, 1964 (Bischler 1964). Bas.: Lejeunea granatensis Prantl, Hedwigia 31: xvi, 1892 (Prantl 1892). ** Drepanolejeunea grandis Herzog, Ann. Bryol. 12: 113, 1939 (Herzog 1939a). *** Drepanolejeunea grollei M.E.Reiner et Schäf.-Verw., Candollea 51 (2): 475, 1996 (Reiner-Drehwald and Schäfer-Verwimp 1996). ** Drepanolejeunea grossidens Steph., Sp. Hepat. (Stephani) 5: 358, 1913 (Stephani 1913a). Based on: Drepanolejeunea grossidens Steph., Hedwigia 28 (3): 168, 1889 (Stephani 1889d), nom. inval. *** Drepanolejeunea hamatifolia (Hook.) Schifn., Hepat. (Engl.-Prantl): 126, 1893 (Schifner 1893b). Bas.: Jungermannia hamatifolia Hook., Brit. Jungermann.: tab. 51, 1813 (Hooker 1813). ** Drepanolejeunea hamulata Steph., Sp. Hepat. (Stephani) 5: 331, 1913 (Stephani 1913a). *** Drepanolejeunea helenae Pócs, Cryptog. Bryol. Lichénol. 18 (3): 198, 1997 (Pócs 1997a). ** Drepanolejeunea herzogii R.L.Zhu et M.L.So, Beih. Nova Hedwigia 121: 181, 2001 (Zhu and So 2001). Nom. nov. pro Strepsilejeunea ocellata Herzog, Memoranda Soc. Fauna Fl. Fennica 26: 57, 1950 [1951] (Herzog 1950b). 299 Drepanolejeunea deslooveri was treated as conspeciic with Drepanolejeunea hamatifolia by Tixier (1995b) and accepted uncritically as such by Wigginton and Grolle (1996) and Wigginton (2002). However, no published supporting evidence for the synonymy has been found, and it was therefore reinstated pending further studies (Wigginton 2009). 356 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Drepanolejeunea inchoata (C.F.W.Meissn.) Steph., Primit. l. costar.: 115, 1892 [1893] (Stephani 1892e). Bas.: Jungermannia inchoata C.F.W.Meissn., Nov. Stirp. Pug. 5: 19, 1833 (Lehmann 1833). ** Drepanolejeunea inchoata var. palmicola Pócs, Acta Bot. Hung. 51 (3/4): 381, 2009 (Schäfer-Verwimp and Pócs 2009). ** Drepanolejeunea inchoata var. roraimae (Steph. ex Zwickel) Bischl., Rev. Bryol. Lichénol. 33 (1/2): 48, 1964 (Bischler 1964). Bas.: Drepanolejeunea roraimae Steph. ex Zwickel, Ann. Bryol. 6: 121, 1933 (Zwickel 1933). *** Drepanolejeunea infundibulata (Spruce) A.Evans, Bull. Torrey Bot. Club 30 (1): 35, 1903 (Evans 1903b). Bas.: Lejeunea infundibulata Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 191, 1884 (Spruce 1884). *** Drepanolejeunea integribracteata Bischl., Rev. Bryol. Lichénol. 33 (1/2): 142, 1964 (Bischler 1964). ** Drepanolejeunea laevis (Mitt.) Steph., Sp. Hepat. (Stephani) 5: 357, 1913 (Stephani 1913a). Bas.: Lejeunea laevis Mitt., Fl. vit.: 416, 1871 [1873] (Mitten 1871). * Drepanolejeunea lancifolia (Gottsche) J.B.Jack et Steph., Hedwigia 31 (1): 13, 1892 (Jack and Stephani 1892). Bas.: Lejeunea lancifolia Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 155, 1864 (Gottsche 1864). 300 *** Drepanolejeunea lichenicola (Spruce) Steph., Sp. Hepat. (Stephani) 5: 335, 1913 (Stephani 1913a). Bas.: Lejeunea lichenicola Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 191, 1884 (Spruce 1884). ** Drepanolejeunea longii Grolle et R.L.Zhu, Ann. Bot. Fenn. 36 (2): 115, 1999 (Grolle and Zhu 1999). ** Drepanolejeunea macrodonta (Mitt.) Steph., Sp. Hepat. (Stephani) 5: 357, 1913 (Stephani 1913a). Bas.: Lejeunea macrodonta Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 116, 1860 [1861] (Mitten 1860c). ** Drepanolejeunea mawtmiana Ajit P.Singh et V.Nath, Hepat. Khasi Jaintia Hills: E. Himal.: 251, 2007 (Singh and Nath 2007b). ** Drepanolejeunea microcarpa Pearson, J. Linn. Soc., Bot. 46 (305): 36, 1922 (Pearson 1922b). ** Drepanolejeunea moluccensis Herzog, Ann. Bryol. 7: 88, 1934 (Herzog 1934b). *** Drepanolejeunea mosenii (Steph.) Bischl., Rev. Bryol. Lichénol. 35 (1/4): 118, 1967 [1968] (Bischler 1967). Bas.: Leptolejeunea mosenii Steph., Sp. Hepat. (Stephani) 5: 372, 1913 (Stephani 1913a). * Drepanolejeunea obliqua Steph., Hedwigia 35 (3): 82, 1896 (Stephani 1896b). 301 ** Drepanolejeunea obtriangulata T.Kodama, J. Hattori Bot. Lab. 41: 381, 1976 (Kodama 1976). 300 Drepanolejeunea lancifolia resembles Drepanolejeunea bidens and Drepanolejeunea araucariae, but the type material has not been found (Bischler 1964). 301 Drepanolejeunea obliqua is possibly conspeciic with Drepanolejeunea ternatensis (Söderström et al. 2010a). World checklist of hornworts and liverworts 357 ** Drepanolejeunea obtusifolia T.Yamag., J. Jap. Bot. 59 (11): 332, 1984 (Yamaguchi 1984). *** Drepanolejeunea orthophylla (Nees et Mont.) Bischl., Rev. Bryol. Lichénol. 35 (1/4): 102, 1967 [1968] (Bischler 1967). Bas.: Lejeunea orthophylla Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 265, 1843 (Montagne 1843). *** Drepanolejeunea palmifolia (Nees) Schifn., Hepat. (Engl.-Prantl): 126, 1893 (Schifner 1893b). Bas.: Jungermannia palmifolia Nees, Fl. Bras. (Martius) 1 (1): 366, 1833 (Nees 1833a). *** Drepanolejeunea pentadactyla (Mont.) Steph., Sp. Hepat. (Stephani) 5: 357, 1913 (Stephani 1913a). Bas.: Lejeunea pentadactyla Mont., Ann. Sci. Nat. Bot. (sér. 3) 10: 113, 1848 (Montagne 1848). ** Drepanolejeunea perissodonta (Spruce) Bischl., Rev. Bryol. Lichénol. 33 (1/2): 73, 1964 (Bischler 1964). Bas.: Lejeunea inchoata var. perissodonta Spruce, J. Linn. Soc., Bot. 30 (210): 340, 1895 (Gepp 1895b). *** Drepanolejeunea physifolia (Gottsche) Pearson, Forh. Vidensk.-Selsk. Kristiania 1892 (8): 8, 1892 (Pearson 1892). Bas.: Lejeunea physifolia Gottsche, Abh. Naturwiss. Vereins Bremen 7: 357, 1882 (Gottsche 1882). ** Drepanolejeunea pinnatiloba Schifn., Bot. Jahrb. Syst. 23 (5): 591, 1897 (Schifner 1897). ** Drepanolejeunea pleiodictya Herzog, Ann. Bryol. 7: 89, 1934 (Herzog 1934b). ** Drepanolejeunea propagulifera Herzog, Mitt. Inst. Allg. Bot. Hamburg 7 (3): 204, 1931 (Herzog 1931a). ** Drepanolejeunea pseudoneura (A.Evans) Grolle, J. Hattori Bot. Lab. 65: 405, 1988 (Grolle 1988c). Bas.: Harpalejeunea pseudoneura A.Evans, Trans. Connecticut Acad. Arts 10 (8): 427, 1900 (Evans 1900a). ** Drepanolejeunea pterocalyx (Herzog) Bischl., Rev. Bryol. Lichénol. 35 (1/4): 114, 1967 [1968] (Bischler 1967). Bas.: Leptolejeunea pterocalyx Herzog, Feddes Repert. Spec. Nov. Regni Veg. 57 (1/2): 187, 1955 (Herzog 1955). ** Drepanolejeunea pungens Bischl., Rev. Bryol. Lichénol. 33 (1/2): 104, 1964 (Bischler 1964). ** Drepanolejeunea ramentilora Steph., Sp. Hepat. (Stephani) 5: 338, 1913 (Stephani 1913a). * Drepanolejeunea ruandensis Vanden Berghen, Bull. Soc. Roy. Bot. Belgique 93: 63, 1961 (Vanden Berghen 1961). 302 *** Drepanolejeunea senticosa Bischl., Rev. Bryol. Lichénol. 33 (1/2): 96, 1964 (Bischler 1964). ** Drepanolejeunea sikkimensis (Udar et U.S.Awasthi) Grolle, J. Hattori Bot. Lab. 55: 503, 1984 (Grolle 1984a). Bas.: Leptolejeunea sikkimensis Udar et U.S.Awasthi, Misc. Bryol. Lichenol. 8 (6): 115, 1979 (Udar and Awasthi 1979). 302 Drepanolejeunea ruandensis is conspeciic with Drepanolejeunea cultrella in Vanden Berghen (1972a), but it was accepted by Tixier (1995b) and Wigginton and Grolle (1996). 358 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Drepanolejeunea spinosa Herzog, Feddes Repert. Spec. Nov. Regni Veg. 57 (1/2): 185, 1955 (Herzog 1955). ** Drepanolejeunea spinosocornuta Steph., Sp. Hepat. (Stephani) 5: 351, 1913 (Stephani 1913a). * Drepanolejeunea subdissitifolia Herzog, Memoranda Soc. Fauna Fl. Fennica 25: 65, 1950 (Herzog 1950c). ** Drepanolejeunea submuricata R.M.Schust., Phytotaxa 208 (1): 98, 2015 (Pócs et al. 2015a). Based on: Drepanolejeunea submuricata R.M.Schust., Nova Hedwigia 62 (1/2): 34, 1996 (Schuster 1996c), nom. inval. ** Drepanolejeunea subquadrata (Mitt.) Steph., Bot. Jahrb. Syst. 23 (1/2, 3): 311, 1896 (Stephani 1896a). Bas.: Lejeunea subquadrata Mitt., Fl. vit.: 416, 1871 [1873] (Mitten 1871). ** Drepanolejeunea subvittata (Herzog) Grolle, J. Hattori Bot. Lab. 69: 186, 1991 (Grolle 1991). Bas.: Harpalejeunea subvittata Herzog, Svensk Bot. Tidskr. 51 (1): 195, 1957 (Herzog 1957a). * Drepanolejeunea tenax K.I.Goebel, Ann. Jard. Bot. Buitenzorg 39: 15, 1928 (Goebel 1928). ** Drepanolejeunea tenera K.I.Goebel, Ann. Jard. Bot. Buitenzorg 39: 20, 1928 (Goebel 1928). 303 * Drepanolejeunea tenera var. litoceras Herzog, Ann. Bryol. 7: 85, 1934 (Herzog 1934b). *** Drepanolejeunea ternatensis (Gottsche) Schifn., Hepat. (Engl.-Prantl): 126, 1893 (Schifner 1893b). Bas.: Lejeunea ternatensis Gottsche, Syn. Hepat. 3: 346, 1845 (Gottsche et al. 1845b). ** Drepanolejeunea teysmannii (Gottsche) Steph., Hedwigia 35 (3): 84, 1896 (Stephani 1896b). Bas.: Lejeunea teysmannii Gottsche, Abh. Naturwiss. Vereins Bremen 7: 360, 1882 (Gottsche 1882). ** Drepanolejeunea tridactyla (Gottsche) Steph., Sp. Hepat. (Stephani) 5: 354, 1913 (Stephani 1913a). Bas.: Lejeunea tridactyla Gottsche, Syn. Hepat. 3: 347, 1845 (Gottsche et al. 1845b). *** Drepanolejeunea trigonophylla Steph., Hedwigia 35 (3): 85, 1896 (Stephani 1896b). ** Drepanolejeunea tristaniana S.W.Arnell, Results Norweg. Sci. Exped. Tristan da Cunha 42: 7, 1958 (Arnell 1958b). ** Drepanolejeunea tuyamae S.Hatt., Bot. Mag. (Tokyo) 64 (755/756): 116, 1951 (Hattori 1951c). ** Drepanolejeunea ualanensis Inoue et H.A.Mill., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 8 (2): 148, 1965 (Inoue and Miller 1965). ** Drepanolejeunea ungulata (Steph.) Grolle, J. Hattori Bot. Lab. 69: 187, 1991 (Grolle 1991). Bas.: Harpalejeunea ungulata Steph., Sp. Hepat. (Stephani) 5: 264, 1913 (Stephani 1913a). 303 Drepanolejeunea tenera is possibly conspeciic with Drepanolejeunea pentadactyla (Söderström et al. 2010a). World checklist of hornworts and liverworts 359 ** Drepanolejeunea urceolata R.M.Schust., Phytologia 39 (6): 427, 1978 (Schuster 1978b). ** Drepanolejeunea valiae Jovet-Ast, Rev. Bryol. Lichénol. 18 (1/2): 38, 1949 (JovetAst 1949a). ** Drepanolejeunea vandenberghenii Buchb. et Eb.Fisch., J. Bryol. 26 (4): 273, 2004 (Buchbender and Fischer 2004). *** Drepanolejeunea vesiculosa (Mitt.) Steph., Sp. Hepat. (Stephani) 5: 356, 1913 (Stephani 1913a). Bas.: Lejeunea vesiculosa Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 116, 1860 [1861] (Mitten 1860c). ** Drepanolejeunea yulensis Steph., Sp. Hepat. (Stephani) 5: 356, 1913 (Stephani 1913a). *** *** *** *** *** ** ** *** subg. Kolpolejeunea Grolle, J. Hattori Bot. Lab. 40: 193, 1976 (Grolle 1976c). Drepanolejeunea intermedia Zwickel, Ann. Bryol. 6: 119, 1933 (Zwickel 1933). Drepanolejeunea lyrata Grolle, J. Hattori Bot. Lab. 40: 199, 1976 (Grolle 1976c). Drepanolejeunea madagascariensis (Steph.) Grolle, Lindbergia 2 (3/4): 232, 1974 (Grolle and Onraedt 1974). Bas.: Leptolejeunea madagascariensis Steph., Sp. Hepat. (Stephani) 5: 363, 1913 (Stephani 1913a). Drepanolejeunea pocsii Grolle, J. Hattori Bot. Lab. 40: 209, 1976 (Grolle 1976c). Drepanolejeunea symoensii Vanden Berghen et Grolle, J. Hattori Bot. Lab. 49: 86, 1981 (Grolle 1981). Based on: Leptolejeunea symoensii Vanden Berghen, Bull. Soc. Roy. Bot. Belgique 93: 58, 1961 (Vanden Berghen 1961), nom. inval. Drepanolejeunea symoensii var. minor Tixier, Trop. Bryol. 11: 25, 1995 (Tixier 1995b). Drepanolejeunea trematodes (Nees) Bischl., Rev. Bryol. Lichénol. 35 (1/4): 125, 1967 [1968] (Bischler 1967). Bas.: Lejeunea trematodes Nees, Ann. Sci. Nat. Bot. (sér. 2) 5: 63, 1836 (Nees and Montagne 1836). ** subg. Pristolejeunea Grolle, J. Hattori Bot. Lab. 40: 193, 1976 (Grolle 1976c). ** Drepanolejeunea actinogyna Inuthai, R.L.Zhu et Chantanaorr., Bryologist 117 (2): 165, 2014 (Inuthai et al. 2014). *** Drepanolejeunea issicornua Steph., Sp. Hepat. (Stephani) 5: 344, 1913 (Stephani 1913a). * Drepanolejeunea hampeana Steph., Sp. Hepat. (Stephani) 5: 345, 1913 (Stephani 1913a). Nom. nov. pro Drepanolejeunea hampeana Steph., Hedwigia 29 (2): 70, 1890 (Stephani 1890b), nom. inval. ** Drepanolejeunea laciniata Qiong He et R.L.Zhu, Cryptog. Bryol. 33 (3): 292, 2012 (He et al. 2012a). *** Drepanolejeunea levicornua Steph., Sp. Hepat. (Stephani) 5: 347, 1913 (Stephani 1913a). * Drepanolejeunea longicornua (Herzog) Mizut., J. Hattori Bot. Lab. 68: 368, 1990 (Mizutani 1990). Bas.: Drepanolejeunea levicornua var. longicornua Herzog, Ann. Bryol. 3: 142, 1930 (Herzog 1930b). * Drepanolejeunea nymanii Steph., Sp. Hepat. (Stephani) 5: 348, 1913 (Stephani 1913a). 360 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Drepanolejeunea pulla (Mitt.) Grolle, J. Hattori Bot. Lab. 46: 349, 1979 (Grolle 1979d). Bas.: Lejeunea pulla Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 116, 1860 [1861] (Mitten 1860c). * Drepanolejeunea serricalyx Herzog, Ann. Bryol. 9: 126, 1936 [1937] (Herzog 1936a). *** Drepanolejeunea thwaitesiana (Mitt.) Steph., Sp. Hepat. (Stephani) 5: 350, 1913 (Stephani 1913a). Bas.: Lejeunea thwaitesiana Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 117, 1860 [1861] (Mitten 1860c). 304 ** Drepanolejeunea thwaitesiana var. zhengii R.L.Zhu, Beih. Nova Hedwigia 121: 197, 2001 (Zhu and So 2001). *** Drepanolejeunea tricornua Herzog, Ann. Bryol. 9: 124, 1936 [1937] (Herzog 1936a). *** subg. Rhaphidolejeunea (Herzog) Grolle et R.L.Zhu, Nova Hedwigia 70 (3/4): 376, 2000 (Grolle and Zhu 2000). Bas.: Rhaphidolejeunea Herzog, Mitth. hüring. Bot. Vereins 50: 104, 1943 (Herzog 1943c). ** Drepanolejeunea bidoupensis Pócs, Cryptog. Bryol. 34 (3): 293, 2013 (Pócs et al. 2013). *** Drepanolejeunea bischlerae (Grolle) Grolle et R.L.Zhu, Nova Hedwigia 70 (3/4): 391, 2000 (Grolle and Zhu 2000). Bas.: Rhaphidolejeunea bischlerae Grolle, J. Hattori Bot. Lab. 38: 653, 1974 (Grolle 1974b). *** Drepanolejeunea commutata Grolle et R.L.Zhu, Nova Hedwigia 70 (3/4): 377, 2000 (Grolle and Zhu 2000). *** Drepanolejeunea cyclops (Sande Lac.) Grolle et R.L.Zhu, Nova Hedwigia 70 (3/4): 391, 2000 (Grolle and Zhu 2000). Bas.: Lejeunea cyclops Sande Lac., Ann. Mus. Bot. Lugduno-Batavi 1: 310, 1864 (Sande Lacoste 1864). *** Drepanolejeunea leischeri (Steph.) Grolle et R.L.Zhu, Nova Hedwigia 70 (3/4): 379, 2000 (Grolle and Zhu 2000). Bas.: Leptolejeunea leischeri Steph., Sp. Hepat. (Stephani) 5: 382, 1913 (Stephani 1913a). ** Drepanolejeunea foliicola Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 1: 85, 1932 (Horikawa 1932a). ** Drepanolejeunea longicruris (Steph.) Grolle et R.L.Zhu, Nova Hedwigia 70 (3/4): 392, 2000 (Grolle and Zhu 2000). Bas.: Leptolejeunea longicruris Steph., Hedwigia 35 (3): 106, 1896 (Stephani 1896b). *** Drepanolejeunea polyrhiza (Nees) Grolle et R.L.Zhu, Nova Hedwigia 70 (3/4): 392, 2000 (Grolle and Zhu 2000). Bas.: Lejeunea polyrhiza Nees, Syn. Hepat. 3: 403, 1845 (Gottsche et al. 1845b). *** Drepanolejeunea siamensis (Bischl.) Grolle et R.L.Zhu, Nova Hedwigia 70 (3/4): 393, 2000 (Grolle and Zhu 2000). Bas.: Rhaphidolejeunea siamensis Bischl., Rev. Bryol. Lichénol. 36 (1/2): 86, 1968 [1969] (Bischler 1968). 304 Drepanolejeunea thwaitesiana is a species complex also including Drepanolejeunea hampeana, Drepanolejeunea longicornua, Drepanolejeunea nymannii and Drepanolejeunea serricalyx. World checklist of hornworts and liverworts 361 *** Drepanolejeunea spicata (Steph.) Grolle et R.L.Zhu, Nova Hedwigia 70 (3/4): 384, 2000 (Grolle and Zhu 2000). Bas.: Leptolejeunea spicata Steph., Hedwigia 35 (3): 108, 1896 (Stephani 1896b). *** Drepanolejeunea tibetana (P.C.Wu et J.S.Lou) Grolle et R.L.Zhu, Nova Hedwigia 70 (3/4): 386, 2000 (Grolle and Zhu 2000). Bas.: Rhaphidolejeunea tibetana P.C.Wu et J.S.Lou, Acta Phytotax. Sin. 16 (4): 102, 1978 (Wu and Lou 1978). *** Drepanolejeunea yunnanensis (P.C.Chen) Grolle et R.L.Zhu, Nova Hedwigia 70 (3/4): 388, 2000 (Grolle and Zhu 2000). Bas.: Rhaphidolejeunea yunnanensis P.C.Chen, Feddes Repert. Spec. Nov. Regni Veg. 58: 44, 1955 (Chen 1955). Incertae sedis *** Drepanolejeunea dactylophora (Nees, Lindenb. et Gottsche) J.B.Jack et Steph., Hedwigia 31 (1): 12, 1892 (Jack and Stephani 1892). Bas.: Lejeunea dactylophora Nees, Lindenb. et Gottsche, Observ. bot.: 473, 1843 (Gottsche et al. 1843). *** Drepanolejeunea dactylophora var. submuricata Herzog, Ann. Bryol. 4: 92, 1931 (Herzog 1931b). * Drepanolejeunea devendrae Sushil K.Singh et M.Dey, Nelumbo 54: 20, 2012 (Singh and Dey 2012). * Drepanolejeunea integerrima Herzog, Rev. Bryol. Lichénol. 20 (1/2): 146, 1951 [1952] (Herzog 1951a). 305 ** Vitalianthus R.M.Schust. et Giancotti, Nova Hedwigia 57 (3/4): 447, 1993 (Schuster and Giancotti 1993). *** Vitalianthus bischlerianus (K.C.Pôrto et Grolle) R.M.Schust. et Giancotti, Nova Hedwigia 57 (3/4): 448, 1993 (Schuster and Giancotti 1993). Bas.: Drepanolejeunea bischleriana K.C.Pôrto et Grolle, Cryptog. Bryol. Lichénol. 8 (4): 301, 1987 (Pôrto and Grolle 1987). ** Vitalianthus guangxianus R.L.Zhu, Qiong He et Y.M.Wei, J. Bryol. 34 (1): 32, 2012 (He et al. 2012b). ** subtrib. Echinolejeuneinae Gradst. *** Anoplolejeunea (Spruce) Schifn., Hepat. (Engl.-Prantl): 131, 1893 (Schifner 1893b). Bas.: Lejeunea subg. Anoplolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 129, 1884 (Spruce 1884). *** Anoplolejeunea conferta (C.F.W.Meissn. ex Spreng.) A.Evans, Bull. Torrey Bot. Club 35 (4): 175, 1908 (Evans 1908a). Bas.: Jungermannia conferta C.F.W.Meissn. ex Spreng. Syst. Veg. (ed. 16) [Sprengel] 4 (2): 325, 1827 (Sprengel 1827b). 305 Drepanolejeunea integerrima is a Leptolejeunea species (Bischler 1964), but it is probably a synonym since she did not transfer it. 362 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Echinolejeunea R.M.Schust., Beih. Nova Hedwigia 9: 187, 1963 (Schuster 1963a). *** Echinolejeunea papillata (Mitt.) R.M.Schust. ex Hamlin, Rec. Domin. Mus. 7: 260, 1972 (Hamlin 1972). Bas.: Lejeunea papillata Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 158, 1854 (Mitten 1854). ** Kymatolejeunea Grolle, Wiss. Z. Friedrich-Schiller-Univ. Jena, Math.-Naturwiss. Reihe 32 (6): 1005, 1984 (Grolle 1984c). *** Kymatolejeunea bartlettii Grolle, Wiss. Z. Friedrich-Schiller-Univ. Jena, Math.Naturwiss. Reihe 32 (6): 1005, 1984 (Grolle 1984c). ** subtrib. Leiolejeuneinae Schäf.-Verw. et Heinrichs ** Leiolejeunea A.Evans, Bull. Torrey Bot. Club 35 (8): 377, 1908 (Evans 1908b). *** Leiolejeunea grandilora A.Evans, Bull. Torrey Bot. Club 35 (8): 378, 1908 (Evans 1908b). *** subtrib. Lejeuneinae Gradst. *** Harpalejeunea (Spruce) Schifn., Hepat. (Engl.-Prantl): 126, 1893 (Schifner 1893b). Bas.: Lejeunea subg. Harpalejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 164, 1884 (Spruce 1884). ** subg. Cleeiolejeunea Grolle et M.E.Reiner, J. Bryol. 21 (1): 33, 1999 (Grolle and Reiner-Drehwald 1999). *** Harpalejeunea grandis Grolle et M.E.Reiner, J. Bryol. 21 (1): 32, 1999 (Grolle and Reiner-Drehwald 1999). ** subg. Harpalejeunea * Harpalejeunea acuta S.Winkl., Rev. Bryol. Lichénol. 42 (3): 812, 1976 (Winkler 1976). 306 *** Harpalejeunea ancistrodes (Spruce) Schifn., Hepat. (Engl.-Prantl): 127, 1893 (Schifner 1893b). Bas.: Lejeunea ancistrodes Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 169, 1884 (Spruce 1884). ** Harpalejeunea buenaventurae Herzog, Feddes Repert. Spec. Nov. Regni Veg. 57 (1/2): 183, 1955 (Herzog 1955). *** Harpalejeunea cinchonae (Nees) Schifn., Hepat. (Engl.-Prantl): 127, 1893 (Schifner 1893b). Bas.: Lejeunea cinchonae Nees, Syn. Hepat. 3: 342, 1845 (Gottsche et al. 1845b). 306 Harpalejeunea acuta is possibly conspeciic with Harpalejeunea cinchonae (Grolle and ReinerDrehwald 1999). World checklist of hornworts and liverworts 363 ** Harpalejeunea cinchonae var. strigulosa Herzog, Svensk Bot. Tidskr. 51 (1): 192, 1957 (Herzog 1957a). ** Harpalejeunea decurvicuspis (Besch. et C.Massal.) P.Syd., Just’s Bot. Jahresber. 19: 246, 1894 (Sydow 1894). Bas.: Lejeunea decurvicuspis Besch. et C.Massal., Bull. Mens. Soc. Linn. Paris 1 (79): 639, 1886 (Bescherelle and Massalongo 1886). ** Harpalejeunea emarginata Jovet-Ast, Rev. Bryol. Lichénol. 16 (1/2): 38, 1947 [1948] (Jovet-Ast 1947b). ** Harpalejeunea exocellata Herzog, Rev. Bryol. Lichénol. 20 (1/2): 144, 1951 [1952] (Herzog 1951a). ** Harpalejeunea grandistipula R.M.Schust., J. Hattori Bot. Lab. 87: 290, 1999 (Schuster 1999c). *** Harpalejeunea harpaphylla (Herzog) Bischl., Rev. Bryol. Lichénol. 33 (1/2): 164, 1964 (Bischler 1964). Bas.: Drepanolejeunea harpaphylla Herzog, Svensk Bot. Tidskr. 46 (1): 93, 1952 (Herzog 1952e). ** Harpalejeunea herzogii Jovet-Ast, Feddes Repert. Spec. Nov. Regni Veg. 58: 19, 1955 (Jovet-Ast 1955). ** Harpalejeunea longibracteata (Spruce) Steph., Sp. Hepat. (Stephani) 5: 254, 1913 (Stephani 1913a). Bas.: Lejeunea longibracteata Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: cciii, 1889 [1890] (Spruce 1889). ** Harpalejeunea marginalis (Hook.f. et Taylor) Steph., Sp. Hepat. (Stephani) 5: 271, 1913 (Stephani 1913a). Bas.: Jungermannia marginalis Hook.f. et Taylor, London J. Bot. 4: 91, 1845 (Hooker and Taylor 1845). *** Harpalejeunea molleri (Steph.) Grolle, Taxon 38 (1): 89, 1989 (Grolle 1989c). Bas.: Lejeunea molleri Steph., Hedwigia 26 (1): 3, 1887 (Stephani 1887). ** Harpalejeunea molleri subsp. integra (R.M.Schust.) Damsh., Ill. Fl. Nord. Liverw. Hornw.: 615, 2002 (Damsholt 2002). Bas.: Harpalejeunea ovata subsp. integra R.M.Schust., J. Elisha Mitchell Sci. Soc. 83 (4): 199, 1967 (Schuster 1967a). *** Harpalejeunea oxyphylla (Nees et Mont.) Steph., Sp. Hepat. (Stephani) 5: 255, 1913 (Stephani 1913a). Bas.: Lejeunea oxyphylla Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 264, 1843 (Montagne 1843). *** Harpalejeunea parasitica (Hook.f. et Taylor) Steph., Sp. Hepat. (Stephani) 5: 268, 1913 (Stephani 1913a). Bas.: Jungermannia parasitica Hook.f. et Taylor, London J. Bot. 3: 477, 1844 (Hooker and Taylor 1844b). ** Harpalejeunea pinaundensis Grolle, J. Hattori Bot. Lab. 46: 44, 1979 (Grolle 1979a). ** Harpalejeunea relexula A.Evans, Bull. Torrey Bot. Club 35 (8): 375, 1908 (Evans 1908b). ** Harpalejeunea scabra Gradst. et Schäf.-Verw., Cryptog. Bryol. 32 (2): 102, 2011 (Gradstein and Schäfer-Verwimp 2011). *** Harpalejeunea schifneri S.W.Arnell, Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 102, 1964 (Schifner and Arnell 1964). * Harpalejeunea solitaria (Gottsche) Steph., Sp. Hepat. (Stephani) 5: 246, 1913 (Stephani 1913a). Bas.: Lejeunea solitaria Gottsche, Abh. Naturwiss. Vereins Bremen 7: 356, 1882 (Gottsche 1882). 364 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Harpalejeunea stricta (Lindenb. et Gottsche) Steph., Sp. Hepat. (Stephani) 5: 259, 1913 (Stephani 1913a). Bas.: Lejeunea stricta Lindenb. et Gottsche, Syn. Hepat. 5: 756, 1847 (Gottsche et al. 1847). ** Harpalejeunea subacuta A.Evans, Bull. Torrey Bot. Club 30 (10): 547, 1903 (Evans 1903c). * Harpalejeunea tenuicuspis (Spruce) Schifn., Hepat. (Engl.-Prantl): 127, 1893 (Schifner 1893b). Bas.: Lejeunea tenuicuspis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 170, 1884 (Spruce 1884). 307 *** Harpalejeunea tridens (Besch. et Spruce) Steph., Sp. Hepat. (Stephani) 5: 263, 1913 (Stephani 1913a). Bas.: Lejeunea tridens Besch. et Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: clxxxi, 1889 [1890] (Bescherelle and Spruce 1889). ** Harpalejeunea uncinata Steph., Hedwigia 35 (3): 97, 1896 (Stephani 1896b). * Harpalejeunea uncinata var. setulosa Herzog, Feddes Repert. Spec. Nov. Regni Veg. 57 (1/2): 183, 1955 (Herzog 1955). * * * * Incertae sedis Harpalejeunea grossearmata Steph., Biblioth. Bot. 87 (2): 256, 1916 (Stephani 1916a). 308 Harpalejeunea renneri Herzog, Memoranda Soc. Fauna Fl. Fennica 26: 54, 1950 [1951] (Herzog 1950b). 309 Harpalejeunea spruceana Steph., Biblioth. Bot. 87 (2): 257, 1916 (Stephani 1916a). 310 Harpalejeunea vitrea Herzog, Memoranda Soc. Fauna Fl. Fennica 27: 94, 1952 (Herzog 1952c). 311 ** Hattoriolejeunea Mizut., J. Hattori Bot. Lab. 61: 303, 1986 [1987] (Mizutani 1986b). ** Hattoriolejeunea akiyamae Mizut., J. Hattori Bot. Lab. 61: 303, 1986 [1987] (Mizutani 1986b). *** Lejeunea Lib., Ann. Gen. Sci. Phys. 6: 373, 1820 (Libert 1820) nom. conserv. 312 307 Harpalejeunea tenuicuspis is possibly conspeciic with Harpalejeunea oxyphylla (Gradstein and Costa 2003, Grolle and Reiner-Drehwald 1999). 308 Harpalejeunea grossearmata is a Lejeunea species (Grolle and Reiner-Drehwald 1999). It was treated as a doubtful taxon by Gradstein et al. (2003). 309 Harpalejeunea renneri is a Lejeunea species and most likely conspeciic with something (Grolle and Reiner-Drehwald 1999). 310 Harpalejeunea spruceana is a Lejeunea species (non Lejeunea spruceana C.Massal. 1885) (Grolle and Reiner-Drehwald 1999). It was treated as a doubtful taxon by Gradstein et al. (2003). 311 Harpalejeunea vitrea is a Lejeunea species (non Lejeunea vitrea Nees, Lindenb. et Gottsche 1843) (Grolle and Reiner-Drehwald 1999). 312 Lejeunea is a large genus and its infrageneric classiication, which has not been studied in a rigorous manner, needs investigation although Heinrichs et al. (2013) showed two main clades on molecular grounds, provisionally named Lejeunea subg. Lejeunea and Lejeunea subg. Crossotolejeunea. However, their morphology is not yet understood. he subgenera listed here are recently accepted ones. Crossotolejeunea, Eulejeunea, Hygrolejeunea and Taxilejeunea also belong here, but several taxa have neither World checklist of hornworts and liverworts 365 ** Lejeunea tunquiniensis M.E.Reiner et Drehwald, Nova Hedwigia 100 (3/4): 584, 2015 (Reiner-Drehwald 2015). ** subg. Lejeunea *** Lejeunea abyssinica (Gola) Cufod., Phyton (Horn) 4: 75, 1952 (Cufodontis 1952). Bas.: Eulejeunea abyssinica Gola, Ann. Bot. (Rome) 13 (1): 70, 1914 (Gola 1914a). *** Lejeunea acanthogona Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 177, 1884 (Spruce 1884). ** Lejeunea acuminata (Lehm. et Lindenb.) Lehm., Nov. Stirp. Pug. 7: 22, 1838 (Lehmann 1838). Bas.: Jungermannia acuminata Lehm. et Lindenb., Nov. Stirp. Pug. 6: 49, 1834 (Lehmann 1834). ** Lejeunea acuta Mitt., J. Proc. Linn. Soc., Bot. 7 (27): 167, 1863 (Mitten 1863). ** Lejeunea acutata (Steph.) Solari, J. Hattori Bot. Lab. 54: 541, 1983 (Solari 1983a). Bas.: Strepsilejeunea acutata Steph., Hedwigia 35 (3): 127, 1896 (Stephani 1896b). *** Lejeunea adpressa Nees, Repert. Pharm. 76: 45, 1842 (von Flotow et al. 1842). 313 *** Lejeunea aethiopica E.W.Jones, J. Bryol. 13 (3): 387, 1985 (Jones 1985). ** Lejeunea alaskana (R.M.Schust. et Steere) Inoue et Steere, J. Hattori Bot. Lab. 44: 330, 1978 (Steere and Inoue 1978). Bas.: Hygrolejeunea alaskana R.M.Schust. et Steere, Bull. Torrey Bot. Club 85 (3): 190, 1958 (Schuster and Steere 1958). *** Lejeunea alata Gottsche, Syn. Hepat. 3: 406, 1845 (Gottsche et al. 1845b). ** Lejeunea alata var. patriciae Pócs, Candollea 56 (1): 72, 2001 (Pócs 2001). *** Lejeunea albescens (Steph.) Mizut., J. Hattori Bot. Lab. 33: 245, 1970 (Mizutani 1970). Bas.: Taxilejeunea albescens Steph., Hedwigia 35 (3): 132, 1896 (Stephani 1896b). * Lejeunea albilora Colenso, Trans. & Proc. New Zealand Inst. 21: 72, 1889 (Colenso 1889). ** Lejeunea aloba Sande Lac., Plagiochila Sandei: 10, 1856 (Sande Lacoste 1856c). ** Lejeunea alobifolia H.A.Mill., Phytologia 47 (4): 323, 1981 (Miller 1981). Nom. nov. pro Lejeunea aloba Steph., Sp. Hepat. (Stephani) 5: 767, 1915 (Stephani 1915b), nom. illeg. ** Lejeunea amaniensis E.W.Jones, J. Bryol. 13 (3): 392, 1985 (Jones 1985). ** Lejeunea ambigua Lindenb. et Gottsche, Syn. Hepat. 5: 764, 1847 (Gottsche et al. 1847). * Lejeunea amentulifera Steph., Sp. Hepat. (Stephani) 5: 707, 1915 (Stephani 1915b). ** Lejeunea androgyna R.M.Schust., Phytologia 45 (5): 432, 1980 (Schuster 1980b). * Lejeunea angulifolia Mitt., Philos. Trans. 168: 400, 1879 (Mitten 1879). 314 *** Lejeunea angusta (Lehm. et Lindenb.) Mont., Hist. Phys. Cuba, Bot., Pl. Cell.: 469, 1842 (Montagne 1842a). Bas.: Jungermannia angusta Lehm. et Lindenb., Nov. Stirp. Pug. 4: 52, 1832 (Lehmann 1832). been transferred nor synonymized. hey are listed in the “Names in genera not currently accepted” section below. 313 Lejeunea adpressa from South America is very similar to the palaeotropical Lejeunea anisophylla. he two taxa probably represent a single pantropical species (possibly with two subspecies). 314 Lejeunea angulifolia may be conspeciic with Lejeunea cocoes (Pócs 2011a). 366 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Lejeunea anisophylla Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 263, 1843 (Montagne 1843). *** Lejeunea anomala Lindenb. et Gottsche, Linnaea 24 (6): 636, 1851 [1852] (Lindenberg and Gottsche 1851a). * Lejeunea antillana Steph., Hedwigia 27 (11/12): 281, 1888 (Stephani 1888c). ** Lejeunea aphanes Spruce, J. Bot. 19: 36, 1881 (Spruce 1881b). *** Lejeunea apiculata Sande Lac., Ned. Kruidk. Arch. 3: 421, 1854 [1855] (Sande Lacoste 1854). ** Lejeunea aquatica Horik., Sci. Rep. Tôhoku Imp. Univ., Ser. 4, Biol. 5 (4): 643, 1929 [1930] (Horikawa 1929c). ** Lejeunea aquatica var. apiculata S.Hatt., Bull. Tokyo Sci. Mus. 11: 108, 1944 (Hattori 1944d). ** Lejeunea armitii (Steph.) Steph., Sp. Hepat. (Stephani) 5: 768, 1915 (Stephani 1915b). Bas.: Eulejeunea armitii Steph., Hedwigia 28 (3): 169, 1889 (Stephani 1889d). * Lejeunea asperifolia Steph., Sp. Hepat. (Stephani) 5: 708, 1915 (Stephani 1915b). *** Lejeunea asperrima Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 160, 1884 (Spruce 1884). ** Lejeunea asperula (Steph.) Mizut., J. Hattori Bot. Lab. 33: 236, 1970 (Mizutani 1970). Bas.: Taxilejeunea asperula Steph., Sp. Hepat. (Stephani) 5: 499, 1914 (Stephani 1914b). ** Lejeunea asprella Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 175, 1884 (Spruce 1884). * Lejeunea atheatostipa Spruce, J. Bot. 33: 83, 1895 (Gepp 1895a). ** Lejeunea barbata (Herzog) R.L.Zhu et M.J.Lai, Ann. Bot. Fenn. 48 (5): 376, 2011 (Wang et al. 2011). Bas.: Rectolejeunea barbata Herzog, J. Hattori Bot. Lab. 14: 49, 1955 (Herzog and Noguchi 1955). *** Lejeunea bermudiana (A.Evans) R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 1105, 1980 (Schuster 1980c). Bas.: Crossotolejeunea bermudiana A.Evans, Bull. Torrey Bot. Club 33 (3): 132, 1906 (Evans 1906c). ** Lejeunea bidentula Herzog, Symb. Sin. 5: 51, 1930 (Nicholson et al. 1930). ** Lejeunea biformis Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 162 (68), 1864 (Gottsche 1864). ** Lejeunea blepharogona Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 178, 1884 (Spruce 1884). ** Lejeunea blomquistii R.M.Schust., J. Elisha Mitchell Sci. Soc. 78 (1): 64, 1962 (Schuster 1962b). *** Lejeunea boliviensis (Steph.) R.L.Zhu et M.E.Reiner, Bryologist 107 (2): 237, 2004 (Zhu and Reiner-Drehwald 2004). Bas.: Strepsilejeunea boliviensis Steph., Biblioth. Bot. 87 (2): 257, 1916 (Stephani 1916a). ** Lejeunea boryana Mont., Ann. Sci. Nat. Bot. (sér. 2) 9: 47, 1838 (Montagne 1838). ** Lejeunea brenanii E.W.Jones, J. Bryol. 10 (4): 391, 1979 (Jones 1979). *** Lejeunea calcicola R.M.Schust., J. Elisha Mitchell Sci. Soc. 73 (2): 404, 1957 (Schuster 1957d). World checklist of hornworts and liverworts 367 ** Lejeunea calcicola var. mexicana R.M.Schust., J. Elisha Mitchell Sci. Soc. 73 (2): 408, 1957 (Schuster 1957d). ** Lejeunea canariensis (Steph.) Steph., Sp. Hepat. (Stephani) 5: 802, 1915 (Stephani 1915b). Bas.: Eulejeunea canariensis Steph., Mém. Soc. Bot. France 7: 42, 1907 (Pitard and Corbière 1907). *** Lejeunea cancellata Nees et Mont., Hist. Phys. Cuba, Bot., Pl. Cell.: 472, 1842 (Montagne 1842a). ** Lejeunea cantabrigiensis E.W.Jones, J. Bryol. 15 (4): 669, 1989 (Jones 1989). *** Lejeunea capensis Gottsche, Syn. Hepat. 3: 374, 1845 (Gottsche et al. 1845b). * Lejeunea caroliniana Austin, Bot. Bull. (Hanover) 1 (8): 36, 1876 (Austin 1876a). 315 *** Lejeunea catinulifera Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 233, 1884 (Spruce 1884). *** Lejeunea caulicalyx (Steph.) M.E.Reiner et Goda, J. Hattori Bot. Lab. 89: 13, 2000 (Reiner-Drehwald and Goda 2000). Bas.: Crossotolejeunea caulicalyx Steph., Sp. Hepat. (Stephani) 5: 237, 1913 (Stephani 1913a). *** Lejeunea cavifolia (Ehrh.) Lindb., Revis. crit. icon.: 43, 1871 (Lindberg 1871). Bas.: Jungermannia cavifolia Ehrh., Beitr. Naturk. (Ehrhart) 4: 45, 1789 (Ehrhart 1789). ** Lejeunea caviloba (Steph.) Besch., J. Bot. (Morot) 12: 140, 1898 (Bescherelle 1898). Bas.: Eulejeunea caviloba Steph., Hedwigia 35 (3): 86, 1896 (Stephani 1896b). *** Lejeunea cerina (Lehm. et Lindenb.) Lehm. et Lindenb., Syn. Hepat. 3: 391, 1845 (Gottsche et al. 1845b). Bas.: Jungermannia cerina Lehm. et Lindenb., Nov. Stirp. Pug. 5: 16, 1833 (Lehmann 1833). ** Lejeunea chaishanensis S.H.Lin, Yushania 9: 7, 1992 (Lin and Yang 1992). ** Lejeunea cladogyna A.Evans, Amer. J. Bot. 5 (3): 134, 1918 (Evans 1918). ** Lejeunea clavata Lindenb., Syn. Hepat. 3: 379, 1845 (Gottsche et al. 1845b). ** Lejeunea claviformis Lindenb. ex Steph., Sp. Hepat. (Stephani) 5: 727, 1915 (Stephani 1915b). ** Lejeunea cochleata Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 273, 1884 (Spruce 1884). ** Lejeunea cocoes Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 114, 1860 [1861] (Mitten 1860c). *** Lejeunea colensoana (Steph.) M.A.M.Renner, Austral. Syst. Bot. 23 (6): 455, 2010 (Renner et al. 2010b). Bas.: Taxilejeunea colensoana Steph., Hedwigia 35 (3): 132, 1896 (Stephani 1896b). ** Lejeunea compacta (Steph.) Steph., Sp. Hepat. (Stephani) 5: 771, 1915 (Stephani 1915b). Bas.: Eulejeunea compacta Steph., Bull. Herb. Boissier 5 (2): 93, 1897 (Stephani 1897b). ** Lejeunea concinnula Spruce et Steph., J. Bot. 25: 39, 1887 (Spruce 1887a). ** Lejeunea connatistipula (Steph.) Steph., Sp. Hepat. (Stephani) 5: 772, 1915 (Stephani 1915b). Bas.: Eulejeunea connatistipula Steph., Hedwigia 35 (3): 87, 1896 (Stephani 1896b). 315 Lejeunea caroliniana was poorly described and the type specimen so poor that Evans (1902a) could not identify it. 368 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Lejeunea contracta Mizut., J. Hattori Bot. Lab. 33: 248, 1970 (Mizutani 1970). ** Lejeunea controversa Gottsche, Hepat. Eur., Leberm. 56-57: no 556, 1873 (Gottsche and Rabenhorst 1873b). ** Lejeunea convexiloba M.L.So et R.L.Zhu, Bryologist 101 (1): 137, 1998 (So and Zhu 1998). ** Lejeunea corcovadae (Steph.) Bischl., Nova Hedwigia 5 (1/2): 406, 1963 (Bischler et al. 1963). Bas.: Microlejeunea corcovadae Steph., Sp. Hepat. (Stephani) 5: 820, 1915 (Stephani 1915b). ** Lejeunea cordilora Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 283, 1884 (Spruce 1884). ** Lejeunea corralensis A.Evans, Ann. Bryol. 3: 86, 1930 (Evans 1930b). *** Lejeunea corynantha Spruce, J. Linn. Soc., Bot. 30 (210): 344, 1895 (Gepp 1895b). ** Lejeunea crassiretis Mitt., Fl. vit.: 414, 1871 [1873] (Mitten 1871). *** Lejeunea cristulata (Steph.) M.E.Reiner et Goda, J. Hattori Bot. Lab. 89: 21, 2000 (Reiner-Drehwald and Goda 2000). Bas.: Crossotolejeunea cristulata Steph., Hedwigia 35 (3): 75, 1896 (Stephani 1896b). *** Lejeunea cristulilora (Steph.) M.E.Reiner et Goda, J. Hattori Bot. Lab. 89: 19, 2000 (Reiner-Drehwald and Goda 2000). Bas.: Crossotolejeunea cristulilora Steph., Sp. Hepat. (Stephani) 5: 231, 1913 (Stephani 1913a). ** Lejeunea cuspidistipula (Steph.) Steph. ex Watts, Proc. Linn. Soc. New South Wales (ser. 2) 27 (108): 493, 1903 (Watts 1903). Bas.: Eulejeunea cuspidistipula Steph., Hedwigia 35 (3): 88, 1896 (Stephani 1896b). ** Lejeunea cyanomontana R.M.Schust., Phytologia 45 (5): 432, 1980 (Schuster 1980b). ** Lejeunea cyanophora R.M.Schust., J. Hattori Bot. Lab. 26: 246, 1963 (Schuster 1963b). ** Lejeunea cyathearum E.W.Jones, J. Bryol. 8 (1): 86, 1974 (Jones 1974). *** Lejeunea cyathophora Mitt., Hooker’s J. Bot. Kew Gard. Misc. 3: 359, 1851 (Mitten 1851). ** Lejeunea denticalyx (Steph.) Steph., Sp. Hepat. (Stephani) 5: 794, 1915 (Stephani 1915b). Bas.: Eulejeunea denticalyx Steph., Hedwigia 28 (3): 169, 1889 (Stephani 1889d). ** Lejeunea denticuspis (Steph.) Mizut., J. Hattori Bot. Lab. 36: 160, 1972 [1973] (Mizutani 1972a). Bas.: Strepsilejeunea denticuspis Steph., Hedwigia 35 (3): 129, 1896 (Stephani 1896b). ** Lejeunea denudata (Pearson) J.J.Engel, Bryologist 78 (3): 361, 1975 (Engel 1975). Bas.: Eulejeunea denudata Pearson, J. Linn. Soc., Bot. 46 (305): 39, 1922 (Pearson 1922b). *** Lejeunea deplanata Nees, Syn. Hepat. 3: 368, 1845 (Gottsche et al. 1845b). ** Lejeunea deplanata var. cuspidata (Steph.) M.E.Reiner, Nova Hedwigia 91 (3/4): 529, 2010 (Reiner-Drehwald 2010). Bas.: Pycnolejeunea cuspidata Steph., Sp. Hepat. (Stephani) 5: 605, 1914 (Stephani 1914b). ** Lejeunea diaphana Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 168, 1884 (Spruce 1884). World checklist of hornworts and liverworts ** ** *** * ** ** ** ** ** * *** *** ** ** ** ** ** *** ** ** 369 Lejeunea dimorpha T.Kodama, J. Hattori Bot. Lab. 41: 384, 1976 (Kodama 1976). Lejeunea dipterocarpa E.W.Jones, J. Bryol. 7 (1): 44, 1972 (Jones 1972). Lejeunea discreta Lindenb., Syn. Hepat. 3: 361, 1845 (Gottsche et al. 1845b). Lejeunea disjecta Spruce, J. Linn. Soc., Bot. 30 (210): 347, 1895 (Gepp 1895b). Lejeunea diversicuspis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 176, 1884 (Spruce 1884). Lejeunea drummondii Taylor, London J. Bot. 5: 400, 1846 (Taylor 1846b). Lejeunea ecarinata (Steph.) J.M.Coult., Barnes et Arthur, Bot. Gaz. 15 (12): 349, 1890 (Coulter et al. 1890). Bas.: Eulejeunea ecarinata Steph., Bot. Gaz. 15 (11): 283, 1890 (Stephani 1890c). Lejeunea eckloniana Lindenb., Syn. Hepat. 3: 381, 1845 (Gottsche et al. 1845b). Lejeunea eifrigii Mizut., J. Hattori Bot. Lab. 33: 244, 1970 (Mizutani 1970). Nom. nov. pro Taxilejeunea acutiloba Eifrig, Monogr. Stud. Indomal. Art. Taxilejeunea: 94, 1937 (Eifrig 1937). Lejeunea elongella Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 161, 1864 (Gottsche 1864). Lejeunea erostrata M.E.Reiner et Goda, J. Hattori Bot. Lab. 89: 25, 2000 (Reiner-Drehwald and Goda 2000). Nom. nov. pro Crossotolejeunea parva Steph., Sp. Hepat. (Stephani) 5: 241, 1913 (Stephani 1913a). Lejeunea exilis (Reinw., Blume et Nees) Grolle, J. Hattori Bot. Lab. 46: 353, 1979 (Grolle 1979d). Bas.: Jungermannia exilis Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 227, 1824 [1825] (Reinwardt et al. 1824a). Lejeunea exilis var. abnormis (Herzog) G.E.Lee, Polish Bot. J. 58 (1): 61, 2013 (Lee and Gradstein 2013). Bas.: Byssolejeunea abnormis Herzog, Hedwigia 80 (1/2): 84, 1941 (Herzog 1941b). Lejeunea fernandeziana S.W.Arnell, Ark. Bot. (n.ser.) 4 (1): 16, 1957 (Arnell 1957b). Lejeunea irma Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 112, 1860 [1861] (Mitten 1860c). Lejeunea issistipula (Steph.) Steph., Sp. Hepat. (Stephani) 5: 775, 1915 (Stephani 1915b). Bas.: Eulejeunea issistipula Steph., Hedwigia 35 (3): 88, 1896 (Stephani 1896b). Lejeunea lagellaris Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 273, 1884 (Spruce 1884). Lejeunea lava (Sw.) Nees, Naturgesch. Eur. Leberm. 3: 277, 1838 (Nees 1838b). Bas.: Jungermannia lava Sw., Prodr. (Swartz): 144, 1788 (Swartz 1788). 316 Lejeunea lava subsp. moorei (Lindb.) R.M.Schust., J. Elisha Mitchell Sci. Soc. 73 (1): 161, 1957 (Schuster 1957a). Bas.: Lejeunea moorei Lindb., Acta Soc. Sci. Fenn. 10: 487, 1875 (Lindberg 1875). Lejeunea lava subsp. orientalis R.M.Schust., J. Elisha Mitchell Sci. Soc. 73 (1): 161, 1957 (Schuster 1957a). 316 Lejeunea lava is a species complex also including Lejeunea grossecristata. 370 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Lejeunea lava var. pellucida Lindenb. et Gottsche, Linnaea 24 (6): 634, 1851 [1852] (Lindenberg and Gottsche 1851a). ** Lejeunea lava subsp. tabularis (Spreng.) S.W.Arnell, Hepat. South Africa: 199, 1963 (Arnell 1963b). Bas.: Jungermannia tabularis Spreng. Syst. Veg. (ed. 16) [Sprengel] 4 (2): 325, 1827 (Sprengel 1827b). *** Lejeunea lavovirens Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 30 (5): 144, 1873 (Ångström 1873). ** Lejeunea leischeri (Steph.) Mizut., J. Hattori Bot. Lab. 33: 238, 1970 (Mizutani 1970). Bas.: Hygrolejeunea leischeri Steph., Sp. Hepat. (Stephani) 5: 560, 1914 (Stephani 1914b). ** Lejeunea loridana A.Evans, Bull. Torrey Bot. Club 32 (4): 185, 1905 (Evans 1905b). *** Lejeunea fulfordiae (Jovet-Ast) R.L.Zhu, Syst. Bot. 33 (4): 617, 2008 (Zhu and Cheng 2008). Bas.: Amblyolejeunea fulfordiae Jovet-Ast, Rev. Bryol. Lichénol. 17 (1/4): 25, 1948 [1949] (Jovet-Ast 1948). ** Lejeunea fusagasugana Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 158, 1864 (Gottsche 1864). ** Lejeunea galeata Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 172, 1884 (Spruce 1884). ** Lejeunea gayana Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 157, 1864 (Gottsche 1864). ** Lejeunea gibbiloba (Steph.) H.A.Mill., Phytologia 47 (4): 323, 1981 (Miller 1981). Bas.: Eulejeunea gibbiloba Steph., Sp. Hepat. (Stephani) 6: 418, 1923 (Stephani 1923). *** Lejeunea glaucescens Gottsche, Syn. Hepat. 3: 378, 1845 (Gottsche et al. 1845b). ** Lejeunea glaucescens var. acrogyna R.M.Schust., J. Elisha Mitchell Sci. Soc. 73 (2): 400, 1957 (Schuster 1957d). * Lejeunea glaucescens var. obsoleta R.M.Schust., J. Elisha Mitchell Sci. Soc. 73 (2): 395, 1957 (Schuster 1957d). ** Lejeunea globosilora (Steph.) Steph., Sp. Hepat. (Stephani) 5: 795, 1915 (Stephani 1915b). Bas.: Eulejeunea globosilora Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 6): 65, 1900 (Stephani 1900b). ** Lejeunea gomphocalyx Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 174, 1884 (Spruce 1884). ** Lejeunea gracilicaulis Spruce, Mem. Torrey Bot. Club 1 (3): 126, 1890 (Spruce 1890). ** Lejeunea gracilipes (Taylor) Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 213, 1884 (Spruce 1884). Bas.: Omphalanthus gracilipes Taylor, London J. Bot. 5: 385, 1846 (Taylor 1846b). ** Lejeunea gracilis Steph., Sp. Hepat. (Stephani) 5: 777, 1915 (Stephani 1915b). ** Lejeunea gradsteiniana Pócs, Acta Biol. Pl. Agr. 1: 55, 2010 [2011] (Pócs 2010d). Nom. nov. pro Ceratolejeunea aberrans Steph., Sp. Hepat. (Stephani) 6: 399, 1923 (Stephani 1923). ** Lejeunea gradsteinii G.E.Lee, Damanhuri et Latif, Acta Biol. Pl. Agr. 1: 29, 2010 [2011] (Lee et al. 2010). * Lejeunea grossecristata (Steph.) E.W.Jones, Trans. Brit. Bryol. Soc. 5 (3): 556, 1968 (Jones 1968). Bas.: Hygrolejeunea grossecristata Steph., Hedwigia 35 (3): 102, 1896 (Stephani 1896b). World checklist of hornworts and liverworts 371 *** Lejeunea grossiretis (Steph.) M.E.Reiner et Goda, J. Hattori Bot. Lab. 89: 27, 2000 (Reiner-Drehwald and Goda 2000). Bas.: Crossotolejeunea grossiretis Steph., Hedwigia 35 (3): 75, 1896 (Stephani 1896b). ** Lejeunea grossistipula Steph., Sp. Hepat. (Stephani) 5: 739, 1915 (Stephani 1915b). *** Lejeunea grossitexta (Steph.) M.E.Reiner et Goda, J. Hattori Bot. Lab. 89: 29, 2000 (Reiner-Drehwald and Goda 2000). Bas.: Crossotolejeunea grossitexta Steph., Sp. Hepat. (Stephani) 5: 240, 1913 (Stephani 1913a). * Lejeunea grossiuscula Gottsche ex Steph., Sp. Hepat. (Stephani) 5: 739, 1915 (Stephani 1915b). ** Lejeunea hahnii Solari, J. Hattori Bot. Lab. 54: 543, 1983 (Solari 1983a). Nom. nov. pro Microlejeunea grandistipula Steph., Hedwigia 35 (3): 114, 1896 (Stephani 1896b). * Lejeunea haitica Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 263, 1843 (Montagne 1843). ** Lejeunea hawaikiana M.A.M.Renner et de Lange, New Zealand J. Bot. 49 (3): 431, 2011 (Renner and de Lange 2011). Nom. nov. pro Stenolejeunea acuminata R.M.Schust., J. Hattori Bot. Lab. 89: 156, 2000 (Schuster 2000b). ** Lejeunea helenae Pearson, Forh. Vidensk.-Selsk. Kristiania 1886 (3): 6, 1886 (Pearson 1886). ** Lejeunea helmsiana (Steph.) Steph., Sp. Hepat. (Stephani) 5: 796, 1915 (Stephani 1915b). Bas.: Eulejeunea helmsiana Steph., Hedwigia 35 (3): 89, 1896 (Stephani 1896b). *** Lejeunea hepaticola (Steph.) Steph., Sp. Hepat. (Stephani) 5: 714, 1915 (Stephani 1915b). Bas.: Eulejeunea hepaticola Steph., Hedwigia 27 (2): 60, 1888 (Stephani 1888a). ** Lejeunea hibernica Bischl., H.A.Mill. et Bonner ex Grolle, Lindbergia 3 (1/2): 48, 1975 [1976] (Grolle 1975c). Based on: Lejeunea hibernica Bischl., H.A.Mill. et Bonner, Nova Hedwigia 3 (4): 455, 1961 [1962] (Bischler et al. 1961), nom. inval. ** Lejeunea hodgsoniana Grolle ex R.J.Lewington, Bever. et M.A.M.Renner, PhytoKeys 29: 2, 2013 (Lewington et al. 2013). ** Lejeunea howeana Grolle, Wiss. Z. Friedrich-Schiller-Univ. Jena, Math.-Naturwiss. Reihe 31 (2): 219, 1982 (Grolle 1982). Nom. nov. pro Cheilolejeunea wattsiana Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 102, 1914 (Stephani and Watts 1914). ** Lejeunea hui R.L.Zhu, Beih. Nova Hedwigia 121: 134, 2001 (Zhu and So 2001). ** Lejeunea humefacta Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 275, 1884 (Spruce 1884). ** Lejeunea hyalina (Steph.) L.Söderstr. et A.Hagborg, Phytotaxa 220 (2): 188, 2015 (Söderström et al. 2015a). Bas.: Pycnolejeunea hyalina Steph., Sp. Hepat. (Stephani) 5: 614, 1914 (Stephani 1914b). ** Lejeunea ibadana A.J.Harr. et E.W.Jones, J. Bryol. 12 (1): 40, 1982 (Jones 1982). ** Lejeunea immersa Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 186, 1884 (Spruce 1884). ** Lejeunea increscens Spruce, Mem. Torrey Bot. Club 1 (3): 124, 1890 (Spruce 1890). * Lejeunea inlatiloba (Steph.) H.A.Mill., Bonner et Bischl., Nova Hedwigia 14 (1): 67, 1967 (Miller et al. 1967). Bas.: Microlejeunea inlatiloba Steph., Sp. Hepat. (Stephani) 6: 422, 1923 (Stephani 1923). 372 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Lejeunea inlexiloba Prantl, Hedwigia 31: xvi, 1892 (Prantl 1892). Based on: Crossotolejeunea inlexiloba J.B.Jack et Steph., Hedwigia 31 (1): 16, 1892 (Jack and Stephani 1892), nom. inval. *** Lejeunea intricata Prantl, Hedwigia 31: xvi, 1892 (Prantl 1892). Based on: Crossotolejeunea intricata J.B.Jack et Steph., Hedwigia 31 (1): 17, 1892 (Jack and Stephani 1892), nom. inval. *** Lejeunea isophylla E.W.Jones, Trans. Brit. Bryol. Soc. 5 (3): 559, 1968 (Jones 1968). ** Lejeunea japonica Mitt., Trans. Linn. Soc. London, Bot. 3 (3): 203, 1891 (Mitten 1891). ** Lejeunea jardinii Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: ccv, 1889 [1890] (Spruce 1889). ** Lejeunea julacea Steph., Sp. Hepat. (Stephani) 5: 715, 1915 (Stephani 1915b). *** Lejeunea jungneri (Steph.) Steph., Cat. Afr. Pl. (Hiern) 2 (2): 318, 1901 (Stephani 1901d). Bas.: Eulejeunea jungneri Steph., Hedwigia 35 (3): 90, 1896 (Stephani 1896b). ** Lejeunea kashyapii M.Dey, D.K.Singh et D.Singh, J. Bryol. 30 (2): 126, 2008 (Dey et al. 2008). * Lejeunea kilimandjarensis Steph., Sp. Hepat. (Stephani) 5: 716, 1915 (Stephani 1915b). ** Lejeunea kinabalensis Mizut., J. Hattori Bot. Lab. 33: 246, 1970 (Mizutani 1970). ** Lejeunea kodamae Ikegami et Inoue, J. Jap. Bot. 36 (1): 7, 1961 (Inoue 1961a). ** Lejeunea konosensis Mizut., J. Hattori Bot. Lab. 71: 127, 1992 (Mizutani 1992). ** Lejeunea kuerschneriana Pócs, Beih. Nova Hedwigia 138: 100, 2010 (Pócs 2010b). *** Lejeunea laeta (Lehm. et Lindenb.) Lehm. et Lindenb., Syn. Hepat. 3: 380, 1845 (Gottsche et al. 1845b). Bas.: Jungermannia laeta Lehm. et Lindenb., Nov. Stirp. Pug. 6: 45, 1834 (Lehmann 1834). ** Lejeunea laii R.L.Zhu, J. Bryol. 30 (2): 173, 2008 (Wang and Zhu 2008). Nom. nov. pro Microlejeunea ramulosa Herzog, J. Hattori Bot. Lab. 14: 51, 1955 (Herzog and Noguchi 1955). *** Lejeunea lamacerina (Steph.) Schifn., Hedwigia 41 (5): 278, 1902 (Schifner 1902). Bas.: Eulejeunea lamacerina Steph., Hedwigia 35 (3): 91, 1896 (Stephani 1896b). 317 ** Lejeunea lamacerina subsp. gemminata R.M.Schust., J. Elisha Mitchell Sci. Soc. 73 (1): 168, 1957 (Schuster 1957a). ** Lejeunea latilobula (Herzog) R.L.Zhu et M.L.So, J. Bryol. 24 (2): 168, 2002 (Zhu and So 2002). Bas.: Taxilejeunea latilobula Herzog, Symb. Sin. 5: 50, 1930 (Nicholson et al. 1930). ** Lejeunea leptalea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 272, 1884 (Spruce 1884). * Lejeunea leratii (Steph.) Mizut., J. Hattori Bot. Lab. 33: 243, 1970 (Mizutani 1970). Bas.: Hygrolejeunea leratii Steph., Sp. Hepat. (Stephani) 5: 562, 1914 (Stephani 1914b). 318 317 Lejeunea lamacerina forms a North American and a Macaronesian/European clade and the separation of two subspecies are justiied if not treating them at species level (Heinrichs et al. 2013). 318 Lejeunea leratii is probably distinct from Lejeunea mimula (Lee 2013). World checklist of hornworts and liverworts 373 ** Lejeunea leucosis Besch. et Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: clxxxviii, 1889 [1890] (Bescherelle and Spruce 1889). * Lejeunea litoralis Steph., Sp. Hepat. (Stephani) 5: 778, 1915 (Stephani 1915b). ** Lejeunea lomana E.W.Jones, Bull. Brit. Mus. (Nat. Hist.), Bot. 11 (3): 257, 1983 (Jones and Harrington 1983). ** Lejeunea longicollis S.W.Arnell, Results Norweg. Sci. Exped. Tristan da Cunha 42: 12, 1958 (Arnell 1958b). ** Lejeunea longilobula Pócs, Beih. Nova Hedwigia 138: 112, 2010 (Pócs 2010b). Nom. nov. pro Lejeunea halei subsp. africana Pócs, J. Bryol. 29 (2): 89, 2007 (Müller and Pócs 2007). ** Lejeunea lowriana Steph., Sp. Hepat. (Stephani) 5: 779, 1915 (Stephani 1915b). *** Lejeunea lumbricoides (Nees) Nees, Syn. Hepat. 3: 342, 1845 (Gottsche et al. 1845b). Bas.: Jungermannia lumbricoides Nees, Enum. Pl. Crypt. Javae: 40, 1830 (Nees 1830). ** Lejeunea lunatigastria Tixier, Ann. Hist.-Nat. Mus. Natl. Hung. 66: 98, 1974 (Tixier 1974). ** Lejeunea lyratilora Prantl, Hedwigia 31: xvi, 1892 (Prantl 1892). Based on: Hygrolejeunea lyratilora Steph., Hedwigia 31 (4): 169, 1892 (Stephani 1892g), nom. inval. ** Lejeunea magohukui Mizut., Misc. Bryol. Lichenol. 7 (7): 133, 1977 (Mizutani 1977). ** Lejeunea mandonii (Steph.) Müll.Frib., Leberm. Eur. 2 (9): 1281, 1958 (Müller 1958). Bas.: Microlejeunea mandonii Steph., Hedwigia 35 (3): 115, 1896 (Stephani 1896b). *** Lejeunea masoalae Pócs, Beih. Nova Hedwigia 138: 103, 2010 (Pócs 2010b). ** Lejeunea massalongoana (Schifn. ex P.Syd.) Solari, J. Hattori Bot. Lab. 54: 542, 1983 (Solari 1983a).Bas.: Harpalejeunea massalongoana Schifn. ex P.Syd., Just’s Bot. Jahresber. 19: 246, 1894 (Sydow 1894). ** Lejeunea megalantha Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 172, 1884 (Spruce 1884). ** Lejeunea mehrana M.Dey, D.K.Singh et D.Singh, J. Bryol. 30 (2): 128, 2008 (Dey et al. 2008). *** Lejeunea meridensis Ilk.-Borg., Nova Hedwigia 80 (1/2): 59, 2005 (Ilkiu-Borges 2005). ** Lejeunea micholitzii Mizut., J. Hattori Bot. Lab. 33: 236, 1970 (Mizutani 1970). Nom. nov. pro Hygrolejeunea parvisaccata Steph., Sp. Hepat. (Stephani) 5: 567, 1914 (Stephani 1914b). ** Lejeunea microloba Taylor, London J. Bot. 5: 399, 1846 (Taylor 1846b). *** Lejeunea mimula Hürl., Bauhinia 11 (1): 12, 1993 (Hürlimann 1993). Nom. nov. pro Hygrolejeunea luteola Steph., Sp. Hepat. (Stephani) 5: 553, 1914 (Stephani 1914b). *** Lejeunea minutiloba A.Evans, Bull. Torrey Bot. Club 44 (11): 525, 1917 (Evans 1917d). ** Lejeunea minutiloba var. heterogyna R.M.Schust., J. Elisha Mitchell Sci. Soc. 73 (2): 425, 1957 (Schuster 1957d). ** Lejeunea mizutanii Grolle, J. Hattori Bot. Lab. 45: 178, 1979 (Grolle 1979c). Nom. nov. pro Cheilolejeunea zollingeri Steph., Hedwigia 34 (5): 245, 1895 (Stephani 1895b). ** Lejeunea molkenboeriana Sande Lac., Ned. Kruidk. Arch. 3: 421, 1854 [1855] (Sande Lacoste 1854). 374 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Lejeunea monimiae (Steph.) Steph., Sp. Hepat. (Stephani) 5: 747, 1915 (Stephani 1915b). Bas.: Eulejeunea monimiae Steph., Hedwigia 35 (3): 91, 1896 (Stephani 1896b). *** Lejeunea multidentata M.E.Reiner et Mustelier, J. Bryol. 26 (2): 103, 2004 (Reiner-Drehwald and Mustelier 2004). * Lejeunea musae (Spreng.) Gottsche, Lindenb. et Nees, Syn. Hepat. 3: 407, 1845 (Gottsche et al. 1845b). Bas.: Jungermannia musae Spreng. Ann. Wetterauischen Ges. Gesammte Naturk. 1: 25, 1809 (Sprengel 1809). ** Lejeunea muscicola Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 281, 1884 (Spruce 1884). * Lejeunea muscicola var. palmicola Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 282, 1884 (Spruce 1884). ** Lejeunea neelgherriana Gottsche, Syn. Hepat. 3: 354, 1845 (Gottsche et al. 1845b). ** Lejeunea nemoralis Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 159 (65), 1864 (Gottsche 1864). ** Lejeunea nepalensis Steph., Sp. Hepat. (Stephani) 5: 780, 1915 (Stephani 1915b). * Lejeunea nesiotica Besch. et Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: clxxxviii, 1889 [1890] (Bescherelle and Spruce 1889). ** Lejeunea neumanniana Nees, Repert. Pharm. 76: 44, 1842 (von Flotow et al. 1842). ** Lejeunea nietneri (Steph.) Steph. ex Watts, Proc. Linn. Soc. New South Wales (ser. 2) 26 (104): 633, 1902 (Watts 1902). Bas.: Eulejeunea nietneri Steph., Hedwigia 35 (3): 91, 1896 (Stephani 1896b). ** Lejeunea notata Gottsche, Abh. Naturwiss. Vereins Bremen 7: 361, 1882 (Gottsche 1882). ** Lejeunea nymannii Steph., Sp. Hepat. (Stephani) 5: 781, 1915 (Stephani 1915b). ** Lejeunea obfusca Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 114, 1860 [1861] (Mitten 1860c). ** Lejeunea obidensis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 277, 1884 (Spruce 1884). * Lejeunea obscura Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 112, 1860 [1861] (Mitten 1860c). 319 ** Lejeunea obtusata Gottsche, Abh. Naturwiss. Vereins Bremen 7: 354, 1882 (Gottsche 1882). ** Lejeunea okomuensis E.W.Jones, Trans. Brit. Bryol. Soc. 5 (4): 787, 1969 (Jones 1969). *** Lejeunea oligoclada Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: cxcix, 1889 [1890] (Spruce 1889). *** Lejeunea oracola M.A.M.Renner, Austral. Syst. Bot. 23 (6): 448, 2010 (Renner et al. 2010b). ** Lejeunea osculatiana De Not., Mem. Reale Accad. Sci. Torino (ser. 2) 16: 233, 1857 (De Notaris 1857). ** Lejeunea otiana S.Hatt., Bot. Mag. (Tokyo) 65 (763/764): 15, 1952 (Hattori 1952a). 319 Lejeunea obscura is possibly conspeciic with Lejeunea aloba (Söderström et al. 2010a). World checklist of hornworts and liverworts ** ** ** ** * *** * ** ** *** *** *** *** ** *** * *** * *** ** ** 375 Lejeunea ovalifolia Steph., Sp. Hepat. (Stephani) 5: 751, 1915 (Stephani 1915b). Lejeunea paciica Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 262, 1843 (Montagne 1843). Lejeunea pallescens Mitt., Hooker’s J. Bot. Kew Gard. Misc. 3: 360, 1851 (Mitten 1851). Lejeunea pallida Lindenb. et Gottsche, Syn. Hepat. 5: 762, 1847 (Gottsche et al. 1847). Lejeunea pallidissima Gola, Nuovo Giorn. Bot. Ital. (n.ser.) 27 (2/4): 247, 1920 (Gola 1920). 320 Lejeunea papilionacea Prantl, Hedwigia 31: xvii, 1892 (Prantl 1892). Based on: Hygrolejeunea papilionacea Steph., Hedwigia 31 (4): 169, 1892 (Stephani 1892g), nom. inval. Lejeunea paratropa Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 176, 1884 (Spruce 1884). 321 Lejeunea patagonica (Steph.) Steph., Sp. Hepat. (Stephani) 5: 797, 1915 (Stephani 1915b). Bas.: Eulejeunea patagonica Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 6): 66, 1900 (Stephani 1900b). Lejeunea patens Lindb., Acta Soc. Sci. Fenn. 10: 482, 1875 (Lindberg 1875). Lejeunea patersonii (Steph.) Steph., Sp. Hepat. (Stephani) 5: 784, 1915 (Stephani 1915b). Bas.: Eulejeunea patersonii Steph., Hedwigia 35 (3): 92, 1896 (Stephani 1896b). Lejeunea patriciae Schäf.-Verw., Candollea 56 (1): 64, 2001 (Schäfer-Verwimp 2001a). Nom. nov. pro Lejeunea pilifera Tixier, Gard. Bull. Singapore 25 (3): 351, 1971 (Tixier 1971), nom. illeg. Lejeunea paucidentata (Steph.) Grolle, J. Hattori Bot. Lab. 69: 191, 1991 (Grolle 1991). Bas.: Crossotolejeunea paucidentata Steph., Hedwigia 35 (3): 76, 1896 (Stephani 1896b). Lejeunea pectinella Mizut., J. Hattori Bot. Lab. 33: 239, 1970 (Mizutani 1970). Lejeunea perigonialis Gottsche, Mexik. Leverm.: 223, 1863 (Gottsche 1863). Lejeunea perpapillosa M.E.Reiner et K.C.Pôrto, Nova Hedwigia 85 (3/4): 542, 2007 (Reiner-Drehwald and Pôrto 2007). Lejeunea pertusa (Corda ex Nees et Mont.) Gottsche, Lindenb. et Nees, Syn. Hepat. 3: 407, 1845 (Gottsche et al. 1845b). Bas.: Jungermannia pertusa Corda ex Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 5: 63, 1836 (Nees and Montagne 1836). Lejeunea phyllobola Nees et Mont., Hist. Phys. Cuba, Bot., Pl. Cell.: 471, 1842 (Montagne 1842a). Lejeunea phyllobola var. turgidula Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 259, 1884 (Spruce 1884). Lejeunea planiloba A.Evans, Proc. Wash. Acad. Sci. 8: 147, 1906 (Evans 1906b). Lejeunea polilloensis Steph., Sp. Hepat. (Stephani) 5: 786, 1915 (Stephani 1915b). Lejeunea praetervisa Steph., Sp. Hepat. (Stephani) 5: 752, 1915 (Stephani 1915b). 320 Lejeunea pallidissima is possibly conspeciic with Lejeunea lavovirens (Vanden Berghen 1972b). 321 Lejeunea paratropa is closely related to or perhaps conspeciic with Lejeunea raddiana Lindenb. (Grolle and Reiner-Drehwald 1999). 376 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Lejeunea primordialis (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 3: 375, 1845 (Gottsche et al. 1845b). Bas.: Jungermannia primordialis Hook.f. et Taylor, London J. Bot. 4: 92, 1845 (Hooker and Taylor 1845). ** Lejeunea princeps (Steph.) Mizut., J. Hattori Bot. Lab. 34: 454, 1971 (Mizutani 1971a). Bas.: Hygrolejeunea princeps Steph., Sp. Hepat. (Stephani) 5: 568, 1914 (Stephani 1914b). ** Lejeunea procumbens Mitt., Fl. vit.: 413, 1871 [1873] (Mitten 1871). ** Lejeunea propagulifera Gradst., Phytotaxa 9: 54, 2010 (Söderström et al. 2010a). Nom. nov. pro Trachylejeunea schifneri Herzog, Svensk Bot. Tidskr. 42 (3): 239, 1948 (Herzog 1948). ** Lejeunea pteridis Besch. et Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: clxxxvii, 1889 [1890] (Bescherelle and Spruce 1889). *** Lejeunea ptosimophylla C.Massal., Nuovo Giorn. Bot. Ital. 13 (2): 123, 1881 (Massalongo 1881). *** Lejeunea puiggariana Steph., Sp. Hepat. (Stephani) 5: 754, 1915 (Stephani 1915b). *** Lejeunea pulverulenta (Steph.) M.E.Reiner, Cryptog. Bryol. 26 (1): 60, 2005 (Reiner-Drehwald 2005b). Bas.: Taxilejeunea pulverulenta Steph., Sp. Hepat. (Stephani) 5: 477, 1914 (Stephani 1914b). *** Lejeunea pulvinata Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 5: 61, 1836 (Nees and Montagne 1836). Nom. nov. pro Jungermannia pulvinata Lehm. et Lindenb., Nov. Stirp. Pug. 5: 15, 1833 (Lehmann 1833), nom. illeg. *** Lejeunea raddiana Lindenb., Syn. Hepat. 3: 342, 1845 (Gottsche et al. 1845b). ** Lejeunea radicans Lindenb. et Gottsche, Syn. Hepat. 5: 766, 1847 (Gottsche et al. 1847). ** Lejeunea ramosissima Steph., Bot. Jahrb. Syst. 8 (2): 88, 1886 (Stephani 1886d). ** Lejeunea rara Steph., Sp. Hepat. (Stephani) 5: 798, 1915 (Stephani 1915b). 322 * Lejeunea ravenelii Austin, Bot. Bull. (Hanover) 1 (8): 35, 1876 (Austin 1876a). 323 ** Lejeunea recurva M.E.Reiner, Polish Bot. J. 58 (2): 423, 2013 (Reiner-Drehwald et al. 2013). Nom. nov. pro Hygrolejeunea herzogii Steph., Biblioth. Bot. 87 (2): 265, 1916 (Stephani 1916a), nom. illeg. *** Lejeunea relexistipula (Lehm. et Lindenb.) Lehm. et Lindenb., Syn. Hepat. 3: 335, 1845 (Gottsche et al. 1845b). Bas.: Jungermannia relexistipula Lehm. et Lindenb., Nov. Stirp. Pug. 5: 10, 1833 (Lehmann 1833). ** Lejeunea relexistipula var. costaricensis (Steph.) M.E.Reiner, Nova Hedwigia 81 (3/4): 408, 2005 (Reiner-Drehwald 2005a). Bas.: Hygrolejeunea costaricensis Steph., Hedwigia 35 (3): 100, 1896 (Stephani 1896b). *** Lejeunea reinerae Ilk.-Borg., Nova Hedwigia 80 (1/2): 61, 2005 (Ilkiu-Borges 2005). Nom. nov. pro Echinocolea herzogii Mizut. et Grolle, Bot. Mag. (Tokyo) 77 (915): 333, 1964 (Grolle 1964d). 322 Lejeunea rara is the valid name for Lejeunea sinclairii Spruce (1884) hom. illeg. (non Mitten 1862 = hysananthus fruticosus), but Scott and Bradshaw (1985) confused the taxa and considered them conspeciic. Grolle (1982) accepted Spruce’s taxon without commenting on any homonym problem. 323 Lejeunea ravenelii was poorly described and the type specimen so poor that Evans (1902a) could not identify it. World checklist of hornworts and liverworts 377 ** Lejeunea resupinata (Steph.) Steph., Sp. Hepat. (Stephani) 5: 757, 1915 (Stephani 1915b). Bas.: Eulejeunea resupinata Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 23 (III, 2): 22, 1897 (Stephani 1897a). ** Lejeunea reticulata Herzog, Nat. Hist. Juan Fernandez (Botany) 2 (5): 747, 1942 (Herzog 1942a). *** Lejeunea rhigophila M.A.M.Renner, Austral. Syst. Bot. 23 (6): 453, 2010 (Renner et al. 2010b). ** Lejeunea rhodesiae (Sim) R.M.Schust., J. Hattori Bot. Lab. 25: 71, 1962 (Schuster 1962a). Bas.: Stylolejeunea rhodesiae Sim, Trans. Roy. Soc. South Africa 15 (1): 68, 1926 (Sim 1926). *** Lejeunea rionegrensis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 579, 1885 (Spruce 1885). Nom. nov. pro Lejeunea implexa Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 240, 1884 (Spruce 1884), nom. inval. ** Lejeunea riparia Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 113, 1860 [1861] (Mitten 1860c). * Lejeunea rothii (Schwägr.) Gottsche, Lindenb. et Nees, Syn. Hepat. 3: 407, 1845 (Gottsche et al. 1845b). Bas.: Jungermannia rothii Schwägr., Hist. Musc. Hepat. Prodr.: 17, 1814 (Schwägrichen 1814). *** Lejeunea rotundifolia Mitt., Hooker’s J. Bot. Kew Gard. Misc. 3: 359, 1851 (Mitten 1851). ** Lejeunea sanctae-helenae M.Wigginton, J. Bryol. 29 (1): 12, 2007 (Wigginton 2007). ** Lejeunea schusteri Grolle, Haussknechtia 8: 60, 2001 (Grolle 2001). Nom. nov. pro Rectolejeunea denudata R.M.Schust., J. Hattori Bot. Lab. 89: 143, 2000 (Schuster 2000c). ** Lejeunea semiscabrida Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 154 (60), 1864 (Gottsche 1864). ** Lejeunea semperi Steph., Sp. Hepat. (Stephani) 5: 788, 1915 (Stephani 1915b). ** Lejeunea seriata Lindenb. et Gottsche, Syn. Hepat. 5: 762, 1847 (Gottsche et al. 1847). ** Lejeunea sessililora (Steph.) Grolle, Wiss. Z. Friedrich-Schiller-Univ. Jena, Math.Naturwiss. Reihe 37: 171, 1988 (Grolle 1988b). Bas.: Macrolejeunea sessililora Steph., Sp. Hepat. (Stephani) 5: 512, 1914 (Stephani 1914b). ** Lejeunea setacea (Steph.) Steph., Sp. Hepat. (Stephani) 5: 719, 1914 (Stephani 1914b). Bas.: Eulejeunea setacea Steph., Bull. Mus. Natl. Hist. Nat. 18 (2): 120, 1912 (Corbière 1912). ** Lejeunea setiloba Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 281, 1884 (Spruce 1884). ** Lejeunea sharpii (R.M.Schust.) R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 991, 1980 (Schuster 1980c). Bas.: Taxilejeunea sharpii R.M.Schust., J. Elisha Mitchell Sci. Soc. 81 (1): 41, 1965 (Schuster 1965a). ** Lejeunea siccata Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 284, 1884 (Spruce 1884). ** Lejeunea silvatica Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 159 (65), 1864 (Gottsche 1864). 378 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Lejeunea smaragdina Besch. et Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: clxxxii, 1889 [1890] (Bescherelle and Spruce 1889). *** Lejeunea soae R.L.Zhu, Y.M.Wei, L.Söderstr., A.Hagborg et von Konrat, Phytotaxa 81 (1): 1, 2013 (Zhu et al. 2013). Nom. nov. pro Trachylejeunea chinensis Herzog, Symb. Sin. 5: 49, 1930 (Nicholson et al. 1930). ** Lejeunea solanicola Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 280, 1884 (Spruce 1884). *** Lejeunea sordida (Nees) Nees, Naturgesch. Eur. Leberm. 3: 278, 1838 (Nees 1838b). Bas.: Jungermannia sordida Nees, Enum. Pl. Crypt. Javae: 41, 1830 (Nees 1830). *** Lejeunea spiniloba Lindenb. et Gottsche, Syn. Hepat. 5: 770, 1847 (Gottsche et al. 1847). ** Lejeunea spinulilora Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 177, 1884 (Spruce 1884). ** Lejeunea sporadica Besch. et Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: clxxx, 1889 [1890] (Bescherelle and Spruce 1889). * Lejeunea squarrosa (Steph.) Steph., Sp. Hepat. (Stephani) 5: 719, 1915 (Stephani 1915b). Bas.: Eulejeunea squarrosa Steph., Bot. Jahrb. Syst. 20 (3): 317, 1895 (Stephani 1895a). ** Lejeunea squarrosula (Herzog) Solari, J. Hattori Bot. Lab. 54: 543, 1983 (Solari 1983a). Bas.: Strepsilejeunea squarrosula Herzog, Nat. Hist. Juan Fernandez (Botany) 2 (5): 741, 1942 (Herzog 1942a). * Lejeunea stephaniana Mizut., J. Hattori Bot. Lab. 27: 143, 1964 (Mizutani 1964a). Nom. nov. pro Strepsilejeunea heterophylla Steph., Sp. Hepat. (Stephani) 6: 395, 1923 (Stephani 1923). 324 ** Lejeunea stevensiana (Steph.) Mizut., J. Hattori Bot. Lab. 34: 452, 1971 (Mizutani 1971a). Bas.: Taxilejeunea stevensiana Steph., Hedwigia 35 (3): 136, 1896 (Stephani 1896b). ** Lejeunea subacuta Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 113, 1860 [1861] (Mitten 1860c). * Lejeunea subaquatica Schifn., Arch. Hydrobiol., suppl. 21 (3/4): 397, 1955 (Schifner 1955). * Lejeunea subbiida Steph. ex Duss, Enum. musc. Antilles franç., Hép.: 10, 1903 (Duss 1903). Nom. nov. pro Odontolejeunea subbiida Steph., Symb. Antill. (Urban) 3 (2): 277, 1902 (Stephani 1902e), nom. illeg. ** Lejeunea subigiensis (Steph.) Steph., Hedwigia 35 (3): 94, 1896 (Stephani 1896b). Bas.: Eulejeunea subigiensis Steph., Hedwigia 35 (3): 94, 1896 (Stephani 1896b). ** Lejeunea subrufula Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 289, 1884 (Spruce 1884). ** Lejeunea subsessilis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 282, 1884 (Spruce 1884). 324 Lejeunea stephaniana is doubtfully distinct from Lejeunea discreta (Söderström et al. 2010a). World checklist of hornworts and liverworts 379 *** Lejeunea subspathulata Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 173, 1884 (Spruce 1884). ** Lejeunea succulenta Herzog, Rev. Bryol. Lichénol. 20 (1/2): 171, 1951 [1952] (Herzog 1951a). ** Lejeunea sufruticola Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 279, 1884 (Spruce 1884). ** Lejeunea syoshii Inoue, Bull. Natl. Sci. Mus. Tokyo, B 3 (4): 143, 1977 (Inoue 1977b). *** Lejeunea talamancensis M.E.Reiner et Schäf.-Verw., Nova Hedwigia 87 (3/4): 414, 2008 (Reiner-Drehwald and Schäfer-Verwimp 2008). *** Lejeunea tamasii M.E.Reiner, N.Salazar et C.Chung, Polish Bot. J. 58 (2): 420, 2013 (Reiner-Drehwald et al. 2013). ** Lejeunea tamaspocsii G.E.Lee, Polish Bot. J. 58 (1): 66, 2013 (Lee and Gradstein 2013). *** Lejeunea tapajosensis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 223, 1884 (Spruce 1884). ** Lejeunea tarapotensis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 282, 1884 (Spruce 1884). ** Lejeunea tenella Taylor, London J. Bot. 5: 398, 1846 (Taylor 1846b). ** Lejeunea thallophora (Eifrig) Gradst., Phytotaxa 9: 54, 2010 (Söderström et al. 2010a). Bas.: Taxilejeunea thallophora Eifrig, Monogr. Stud. Indomal. Art. Taxilejeunea: 99, 1937 (Eifrig 1937). *** Lejeunea tonduzana (Steph.) M.E.Reiner, Polish Bot. J. 58 (2): 421, 2013 (ReinerDrehwald et al. 2013). Bas.: Hygrolejeunea tonduzana Steph., Hedwigia 35 (3): 105, 1896 (Stephani 1896b). ** Lejeunea trachygona Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 175, 1884 (Spruce 1884). *** Lejeunea trinitensis Lindenb., Syn. Hepat. 3: 381, 1845 (Gottsche et al. 1845b). ** Lejeunea trukensis H.A.Mill. et Bonner, Beih. Nova Hedwigia 11: 56, 1963 (Miller et al. 1963). ** Lejeunea tuberculosa Steph., Sp. Hepat. (Stephani) 5: 790, 1915 (Stephani 1915b). ** Lejeunea tumida Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 157, 1854 (Mitten 1854). ** Lejeunea uleana (Steph.) Steph., Sp. Hepat. (Stephani) 5: 765, 1915 (Stephani 1915b). Bas.: Eulejeunea uleana Steph., Hedwigia 35 (3): 94, 1896 (Stephani 1896b). *** Lejeunea umbilicata (Nees) Nees, Observ. bot.: 472, 1843 (Gottsche et al. 1843). Bas.: Jungermannia umbilicata Nees, Enum. Pl. Crypt. Javae: 42, 1830 (Nees 1830). ** Lejeunea utriculata (Steph.) Mizut., J. Hattori Bot. Lab. 33: 242, 1970 (Mizutani 1970). Bas.: Pycnolejeunea utriculata Steph., Hedwigia 35 (3): 126, 1896 (Stephani 1896b). ** Lejeunea vesicata Mitt., Fl. vit.: 415, 1871 [1873] (Mitten 1871). *** Lejeunea villaumei (Steph.) Grolle, J. Bryol. 9 (4): 536, 1977 [1978] (Grolle 1977a). Bas.: Otigoniolejeunea villaumei Steph., Sp. Hepat. (Stephani) 5: 516, 1914 (Stephani 1914b). ** Lejeunea vojtkoi Pócs, Beih. Nova Hedwigia 138: 107, 2010 (Pócs 2010b). ** Lejeunea vulgariformis Gottsche, Abh. Naturwiss. Vereins Bremen 7: 355, 1882 (Gottsche 1882). 380 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Lejeunea wattsiana (Steph.) H.A.Mill., Bonner et Bischl., Nova Hedwigia 14 (1): 66, 1967 (Miller et al. 1967). Bas.: Microlejeunea wattsiana Steph., Sp. Hepat. (Stephani) 5: 834, 1916 (Stephani 1916b). * Lejeunea wichurae (Steph.) Steph., Sp. Hepat. (Stephani) 5: 792, 1915 (Stephani 1915b). Bas.: Eulejeunea wichurae Steph., Hedwigia 35 (3): 94, 1896 (Stephani 1896b). ** Lejeunea wightii Lindenb., Syn. Hepat. 3: 379, 1845 (Gottsche et al. 1845b). *** Lejeunea xiphophylla (Herzog) M.E.Reiner, Nova Hedwigia 87 (3/4): 409, 2008 (Reiner-Drehwald and Schäfer-Verwimp 2008). Bas.: Taxilejeunea xiphophylla Herzog, Svensk Bot. Tidskr. 46 (1): 99, 1952 (Herzog 1952e). ** subg. Nanolejeunea R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 1092, 1980 (Schuster 1980c). ** Lejeunea curviloba Steph., Sp. Hepat. (Stephani) 5: 774, 1915 (Stephani 1915b). *** Lejeunea laetevirens Nees et Mont., Hist. Phys. Cuba, Bot., Pl. Cell.: 469, 1842 (Montagne 1842a). ** Lejeunea pallidevirens S.Hatt., J. Hattori Bot. Lab. 12: 80, 1954 (Hattori 1954). Nom. nov. pro Microlejeunea rotundistipula var. pallida S.Hatt., J. Hattori Bot. Lab. 5: 53, 1951 (Hattori 1951d). ** Lejeunea parva (S.Hatt.) Mizut., Misc. Bryol. Lichenol. 5 (10/12): 178, 1971 (Mizutani 1971b). Bas.: Microlejeunea rotundistipula f. parva S.Hatt., Bull. Tokyo Sci. Mus. 11: 123, 1944 (Hattori 1944d). *** Lejeunea prionoides Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 238, 1884 (Spruce 1884). *** Lejeunea ramulosa Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 274, 1884 (Spruce 1884). *** subg. Neopotamolejeunea (M.E.Reiner) Gradst. et M.E.Reiner, Syst. Bot. 32 (3): 487, 2007 (Gradstein and Reiner-Drehwald 2007). Bas.: Neopotamolejeunea M.E.Reiner, Nova Hedwigia 71 (3/4): 449, 2000 (Reiner-Drehwald 2000). ** Lejeunea juruana Gradst. et M.E.Reiner, Syst. Bot. 32 (3): 488, 2007 (Gradstein and Reiner-Drehwald 2007). Nom. nov. pro Potamolejeunea uleana Steph., Sp. Hepat. (Stephani) 5: 641, 1914 (Stephani 1914b). *** Lejeunea polyantha Mont., Ann. Sci. Nat. Bot. (sér. 4) 5: 350, 1856 (Montagne 1856c). ** Lejeunea tenera (Sw.) Gottsche, Lindenb. et Nees, Syn. Hepat. 3: 406, 1845 (Gottsche et al. 1845b). Bas.: Jungermannia tenera Sw., Prodr. (Swartz): 143, 1788 (Swartz 1788). *** Lejeunea topoensis Gradst. et M.E.Reiner, Syst. Bot. 32 (3): 488, 2007 (Gradstein and Reiner-Drehwald 2007). ** subg. Papillolejeunea (Pócs) R.M.Schust., J. Hattori Bot. Lab. 85: 84, 1998 (Schuster 1998a). Bas.: Papillolejeunea Pócs, Trop. Bryol. 13: 2, 1997 (Pócs 1997b). ** Lejeunea balazsii (Pócs) R.M.Schust., J. Hattori Bot. Lab. 85: 84, 1998 (Schuster 1998a). Bas.: Papillolejeunea balazsii Pócs, Trop. Bryol. 13: 3, 1997 (Pócs 1997b). World checklist of hornworts and liverworts 381 ** Lejeunea candida (Pócs) R.M.Schust., J. Hattori Bot. Lab. 85: 84, 1998 (Schuster 1998a). Bas.: Papillolejeunea candida Pócs, Trop. Bryol. 13: 8, 1997 (Pócs 1997b). ** Lejeunea falcata (Pócs et J.Eggers) Pócs, Phytotaxa 208 (1): 99, 2015 (Pócs et al. 2015a). Bas.: Papillolejeunea falcata Pócs et J.Eggers, Bryobrothera 5: 163, 1999 (Pócs and Eggers 1999). ** Lejeunea koponenii (Pócs et J.Eggers) Pócs, Phytotaxa 208 (1): 99, 2015 (Pócs et al. 2015a). Bas.: Papillolejeunea koponenii Pócs et J.Eggers, Bryobrothera 5: 159, 1999 (Pócs and Eggers 1999). ** Lejeunea papuana (Pócs) R.M.Schust., J. Hattori Bot. Lab. 85: 84, 1998 (Schuster 1998a). Bas.: Papillolejeunea papuana Pócs, Trop. Bryol. 13: 14, 1997 (Pócs 1997b). ** Lejeunea touwii (Pócs) R.M.Schust., J. Hattori Bot. Lab. 85: 84, 1998 (Schuster 1998a). Bas.: Papillolejeunea touwii Pócs, Trop. Bryol. 13: 11, 1997 (Pócs 1997b). * * ** * ** ** ** ** ** ** *** ** *** ** ** *** Incertae sedis Lejeunea apiahyna (Steph.) Sushil K.Singh, Phytotaxa 96 (1): 63, 2013 (Singh 2013). Bas.: Otigoniolejeunea apiahyna Steph., Sp. Hepat. (Stephani) 5: 514, 1914 (Stephani 1914b). Lejeunea aptycta Gottsche, Syn. Hepat. 3: 369, 1845 (Gottsche et al. 1845b). Lejeunea asthenica Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 222, 1884 (Spruce 1884). Lejeunea bethanica Gottsche, Syn. Hepat. 3: 381, 1845 (Gottsche et al. 1945b) Lejeunea bombonasensis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 222, 1884 (Spruce 1884). Lejeunea bornmuelleri (Steph.) M.E.Reiner, Nova Hedwigia 95 (3/4): 471, 2012 (Reiner-Drehwald and Grolle 2012). Bas.: Rectolejeunea bornmuelleri Steph., Sp. Hepat. (Stephani) 5: 682, 1914 (Stephani 1914b). Lejeunea caracensis Lindenb., Syn. Hepat. 3: 355, 1845 (Gottsche et al. 1845b). Lejeunea caripensis Lindenb. et Gottsche, Syn. Hepat. 5: 758, 1847 (Gottsche et al. 1847). Lejeunea chamissonis Lindenb., Syn. Hepat. 3: 378, 1845 (Gottsche et al. 1845b). Lejeunea chimborazensis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 215, 1884 (Spruce 1884). Lejeunea combuensis O.S.Moura, Ilk.-Borg. et M.E.Reiner, Nova Hedwigia 95 (1/2): 198, 2012 (Moura et al. 2012). Lejeunea concava Lindenb. et Gottsche, Syn. Hepat. 5: 759, 1847 (Gottsche et al. 1847). Lejeunea conformis Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 260, 1843 (Montagne 1843). Lejeunea cordistipula Lindenb. et Gottsche, Syn. Hepat. 5: 758, 1847 (Gottsche et al. 1847). Lejeunea cyrtotis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 229, 1884 (Spruce 1884). Lejeunea debilis (Lehm. et Lindenb.) Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 5: 60, 1836 (Nees and Montagne 1836). Bas.: Jungermannia debilis Lehm. et Lindenb., Nov. Stirp. Pug. 4: 51, 1832 (Lehmann 1832). 382 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Lejeunea devendrae (Sushil K.Singh) P.K.Verma et K.K.Rawat, J. Bryol. 36 (2): 161, 2014 (Verma and Rawat 2014). Bas.: Rectolejeunea devendrae Sushil K.Singh, Indian J. Forest. 34 (3): 341, 2011 (Singh 2011). ** Lejeunea dictyocalyx Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 218, 1884 (Spruce 1884). ** Lejeunea dipterota (Eifrig) G.E.Lee, Polish Bot. J. 58 (1): 61, 2013 (Lee and Gradstein 2013). Bas.: Taxilejeunea dipterota Eifrig, Monogr. Stud. Indomal. Art. Taxilejeunea: 96, 1937 (Eifrig 1937). *** Lejeunea drehwaldii Heinrichs et Schäf.-Verw., Phytotaxa 69: 14, 2012 (Heinrichs et al. 2012b). Nom. nov. pro Sphaerolejeunea umbilicata Herzog, Ann. Bryol. 11: 88, 1938 (Herzog 1938b). ** Lejeunea duncaniae (Sim) M.E.Reiner, Phytotaxa 208 (1): 98, 2015 (Pócs et al. 2015a). Bas.: Stylolejeunea duncaniae Sim, Trans. Roy. Soc. South Africa 15 (1): 68, 1926 (Sim 1926). ** Lejeunea edentata L.Söderstr., Phytotaxa 208 (1): 98, 2015 (Pócs et al. 2015a). Nom. nov. pro Cyclolejeunea marginata R.M.Schust., Phytologia 39 (6): 430, 1978 (Schuster 1978b). * Lejeunea emarginulilora Gottsche ex Steph., Sp. Hepat. (Stephani) 5: 734, 1915 (Stephani 1915b). 325 * Lejeunea epibrya Taylor, London J. Bot. 7: 199, 1848 (Taylor 1848a). ** Lejeunea estrellamontana M.A.M.Renner et Pócs, Phytotaxa 81 (1): 9, 2013 (Renner et al. 2013d). Nom. nov. pro Stenolejeunea issistipula R.M.Schust., J. Hattori Bot. Lab. 89: 167, 2000 (Schuster 2000b). * Lejeunea fawcettiae D.J.Carr, Proc. Roy. Soc. Victoria 117 (2): 325, 2005 (Carr 2005). Based on: Lejeunea fawcettiae D.J.Carr, Proc. Roy. Soc. Victoria 116 (2): 229, 2004 (Carr 2004), nom. inval. ** Lejeunea laccida Lindenb. et Gottsche, Syn. Hepat. 5: 758, 1847 (Gottsche et al. 1847). ** Lejeunea lavida Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 113, 1860 [1861] (Mitten 1860c). ** Lejeunea lorida Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 221, 1884 (Spruce 1884). ** Lejeunea fulva Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 237, 1884 (Spruce 1884). ** Lejeunea gottscheana Lindenb., Syn. Hepat. 3: 382, 1845 (Gottsche et al. 1845b). ** Lejeunea graminicolor Spruce, J. Linn. Soc., Bot. 30 (210): 343, 1895 (Gepp 1895b). *** Lejeunea grolleana (Bernecker) R.L.Zhu et W.Ye, J. Syst. Evol. 51 (4): 472, 2013 (Ye et al. 2013b). Bas.: Oryzolejeunea grolleana Bernecker, Haussknechtia, Beih. 9: 37, 1999 (Bernecker-Lücking 1999). *** Lejeunea herminieri (Steph.) R.L.Zhu, Phytotaxa 208 (1): 99, 2015 (Pócs et al. 2015a). Bas.: Archilejeunea herminieri Steph., Sp. Hepat. (Stephani) 4: 714, 1911 (Stephani 1911e). 325 Lejeunea emarginulilora is a Lejeunea (subg. Heterolejeunea) species (Reiner-Drehwald and Grolle 2012). World checklist of hornworts and liverworts 383 ** Lejeunea heterocheila Taylor, London J. Bot. 5: 394, 1846 (Taylor 1846b). * Lejeunea hygrophila Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 157, 1864 (Gottsche 1864). ** Lejeunea laevicalyx Gottsche, Mexik. Leverm.: 221, 1863 (Gottsche 1863). * Lejeunea laxa (Nees) Lindenb., Syn. Hepat. 3: 378, 1845 (Gottsche et al. 1845b). Bas.: Jungermannia thymifolia δ laxa Nees, Enum. Pl. Crypt. Javae: 43, 1830 (Nees 1830). ** Lejeunea leiantha Spruce, J. Linn. Soc., Bot. 30 (210): 345, 1895 (Gepp 1895b). ** Lejeunea leptoscypha Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: ccv, 1889 [1890] (Spruce 1889). *** Lejeunea lusoria (Lindenb. et Gottsche) Steph., Hedwigia 29 (3): 141, 1890 (Stephani 1890d). Bas.: Omphalanthus lusorius Lindenb. et Gottsche, Syn. Hepat. 5: 747, 1847 (Gottsche et al. 1847). ** Lejeunea luzonensis (Steph.) R.L.Zhu et M.J.Lai, Ann. Bot. Fenn. 48 (5): 376, 2011 (Wang et al. 2011). Bas.: Taxilejeunea luzonensis Steph., Hedwigia 35 (3): 134, 1896 (Stephani 1896b). ** Lejeunea macrorhyncha Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 220, 1884 (Spruce 1884). ** Lejeunea malangensis (Herzog) R.L.Zhu et Y.M.Wei, J. Bryol. 34 (4): 319, 2012 (Zhu and Wei 2012). Bas.: Trachylejeunea malangensis Herzog, Mitt. Inst. Allg. Bot. Hamburg 7 (3): 203, 1931 (Herzog 1931a). ** Lejeunea marasmodes Spruce, Mem. Torrey Bot. Club 1 (3): 125, 1890 (Spruce 1890). ** Lejeunea morobensis (Grolle) M.A.M.Renner et Pócs, Phytotaxa 81 (1): 9, 2013 (Renner et al. 2013d). Bas.: Stenolejeunea morobensis Grolle, J. Hattori Bot. Lab. 29: 76, 1966 (Grolle 1966e). ** Lejeunea novoguineensis Schifn., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 60 (2): 238, 1893 (Schifner 1893a). *** Lejeunea obtusangula Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 221, 1884 (Spruce 1884). * Lejeunea oerstediana Lindenb. et Hampe, Linnaea 24 (6): 641, 1851 [1852] (Hampe 1851a). 326 ** Lejeunea paraensis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 224, 1884 (Spruce 1884). ** Lejeunea parviloba Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 33 (7): 87, 1876 [1877] (Ångström 1876). * Lejeunea pleidereri Sushil K.Singh, Phytotaxa 96 (1): 63, 2013 (Singh 2013). Nom. nov. pro Otigoniolejeunea indica Steph., Sp. Hepat. (Stephani) 6: 408, 1923 (Stephani 1923). * Lejeunea prominula Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 157, 1864 (Gottsche 1864). 326 Lejeunea oerstediana is a poorly known taxon often placed in Euosmolejeunea (=Cheilolejeunea). We can not tell where it belongs until the type specimen is re-studied (Söderström et al. 2011a). 384 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Lejeunea pterigonia (Lehm. et Lindenb.) Mont., Ann. Sci. Nat. Bot. (sér. 2) 14: 337, 1840 (Montagne 1840a). Bas.: Jungermannia pterigonia Lehm. et Lindenb., Nov. Stirp. Pug. 6: 44, 1834 (Lehmann 1834). ** Lejeunea quinqueumbonata Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 230, 1884 (Spruce 1884). ** Lejeunea quinqueumbonata var. rotundata (Herzog) Sushil K.Singh, Phytotaxa 96 (1): 64, 2013 (Singh 2013). Bas.: Otigoniolejeunea quinqueumbonata var. rotundata Herzog, Rev. Bryol. Lichénol. 20 (1/2): 154, 1951 [1952] (Herzog 1951a). ** Lejeunea remotifolia Hampe ex Steph., Sp. Hepat. (Stephani) 5: 756, 1915 (Stephani 1915b). *** Lejeunea ruthii (A.Evans) R.M.Schust., J. Hattori Bot. Lab. 25: 23, 1962 (Schuster 1962a). Bas.: Microlejeunea ruthii A.Evans, Mem. Torrey Bot. Club 8 (2): 161, 1902 (Evans 1902a). ** Lejeunea ruthii var. alata R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 1063, 1980 (Schuster 1980c). ** Lejeunea sikorae (Steph.) Steph., Bull. Soc. Roy. Bot. Belgique, Mém. 32: 120, 1893 [1894] (Renauld and Cardot 1893). Bas.: Taxilejeunea sikorae Steph., Bull. Soc. Roy. Bot. Belgique, Compt. Rend. 32 (2): 34, 1893 [1894] (Stephani 1893e). ** Lejeunea srivastavae P.K.Verma et K.K.Rawat, Taiwania 58 (1): 8, 2013 (Verma and Rawat 2013). ** Lejeunea stenodentata M.A.M.Renner et Pócs, Phytotaxa 81 (1): 8, 2013 (Renner et al. 2013d). Nom. nov. pro Drepanolejeunea dentata Steph., Hedwigia 35 (3): 82, 1896 (Stephani 1896b). *** Lejeunea subelobata Carrington et Pearson, Proc. Linn. Soc. New South Wales (ser. 2) 2 (4): 1039, 1888 (Carrington and Pearson 1888a). ** Lejeunea subolivacea Mizut., J. Hattori Bot. Lab. 28: 121, 1965 (Mizutani 1965). Nom. nov. pro Eulejeunea olivacea Steph., Hedwigia 29 (2): 85, 1890 (Stephani 1890b). ** Lejeunea subplana (Steph.) C.J.Bastos, J. Bryol. 36 (3): 249, 2014 (Bastos 2014). Bas.: Trachylejeunea subplana Steph., Sp. Hepat. (Stephani) 5: 310, 1913 (Stephani 1913a). *** Lejeunea sulphurea (Lehm. et Lindenb.) Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 217, 1884 (Spruce 1884). Bas.: Jungermannia sulphurea Lehm. et Lindenb., Nov. Stirp. Pug. 5: 14, 1833 (Lehmann 1833). ** Lejeunea terricola Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: cxci, 1889 [1890] (Spruce 1889). ** Lejeunea urbanii (Steph.) Steph., Sp. Hepat. (Stephani) 5: 766, 1915 (Stephani 1915b). Bas.: Eulejeunea urbanii Steph., Hedwigia 27 (11/12): 301, 1888 (Stephani 1888b). ** Lejeunea venezuelana (R.M.Schust.) R.L.Zhu et W.Ye, J. Syst. Evol. 51 (4): 473, 2013 (Ye et al. 2013b). Bas.: Cyrtolejeunea venezuelana R.M.Schust., Phytologia 39 (6): 426, 1978 (Schuster 1978b). ** Lejeunea viridis R.M.Schust. ex L.Söderstr. et A.Hagborg, Phytotaxa 208 (1): 99, 2015 (Pócs et al. 2015a). Based on: Prionocolea viridissima R.M.Schust., J. Hattori Bot. Lab. 75: 215, 1994 (Schuster 1994), nom. inval. World checklist of hornworts and liverworts * * * * * * 385 Lejeunea zacuapana (Steph.) Prantl, Hedwigia 29: xviii, 1890 (Prantl 1890). Bas.: Eulejeunea zacuapana Steph., Hedwigia 29 (2): 87, 1890 (Stephani 1890b). Excluded from the genus Lejeunea elegans Gottsche, Syn. Hepat. 3: 364, 1845 (Gottsche et al. 1845b). 327 Lejeunea hieronymii Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: cciii, 1889 [1890] (Spruce 1889). 328 Lejeunea proboscidea Gottsche, Mexik. Leverm.: 225, 1863 (Gottsche 1863). 329 Lejeunea scabrilora Loitl., Diagn. pl. nov.: 22, 1894 (Szyszylowicz 1894). 330 Lejeunea trochantha Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: cxcii, 1889 [1890] (Spruce 1889). 331 ** Microlejeunea (Spruce) Steph., Hedwigia 27 (2): 61, 1888 (Stephani 1888a). Bas.: Lejeunea subg. Microlejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 286, 1884 (Spruce 1884). *** Microlejeunea acutifolia Steph., Hedwigia 35 (3): 113, 1896 (Stephani 1896b). ** Microlejeunea africana Steph., Hedwigia 27 (2): 61, 1888 (Stephani 1888a). ** Microlejeunea aligera (Mitt.) Steph., Sp. Hepat. (Stephani) 5: 827, 1915 (Stephani 1915b). Bas.: Lejeunea aligera Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 113, 1860 [1861] (Mitten 1860c). ** Microlejeunea ankasica E.W.Jones, J. Bryol. 10 (4): 394, 1979 (Jones 1979). ** Microlejeunea aphanella (Spruce) Steph., Sp. Hepat. (Stephani) 5: 816, 1915 (Stephani 1915b). Bas.: Lejeunea aphanella Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 290, 1884 (Spruce 1884). ** Microlejeunea atsuana Steph., Hedwigia 35 (3): 113, 1896 (Stephani 1896b). ** Microlejeunea bischlerae (B.M.hiers) B.M.hiers, Phytotaxa 65: 59, 2012 (hiers et al. 2012). Bas.: Lejeunea bischlerae B.M.hiers, Cryptog. Bryol. Lichénol. 18 (3): 223, 1997 (hiers 1997b). *** Microlejeunea bullata (Taylor) Steph., Hedwigia 29 (2): 90, 1890 (Stephani 1890b). Bas.: Lejeunea bullata Taylor, London J. Bot. 5: 398, 1846 (Taylor 1846b). * Microlejeunea byssoides (Gottsche) Pearson, Forh. Vidensk.-Selsk. Kristiania 1892 (8): 9, 1892 (Pearson 1892). Bas.: Lejeunea byssoides Gottsche, Abh. Naturwiss. Vereins Bremen 7: 358, 1882 (Gottsche 1882). ** Microlejeunea capillaris (Gottsche) Steph., Sp. Hepat. (Stephani) 5: 819, 1915 (Stephani 1915b). Bas.: Lejeunea capillaris Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 163, 1864 (Gottsche 1864). 327 328 329 330 Lejeunea elegans is a Cheilolejeunea species (Reiner-Drehwald 2006). Lejeunea hieronymii is a Cheilolejeunea species of uncertain status. Lejeunea proboscidea is a Drepanolejeunea species, but Bischler (1964) did not place it to any species. Lejeunea scabrilora was rejected from Dicranolejeunea by Kruijt (1988), but he did not refer it to any other genus. 331 Lejeunea trochantha is a Cheilolejeunea species (Reiner-Drehwald 2006). 386 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Microlejeunea cochlearifolia Steph., Hedwigia 27 (3/4): 113, 1888 (Stephani 1888d). 332 ** Microlejeunea colombiana Bischl., Nova Hedwigia 5 (1/2): 373, 1963 (Bischler et al. 1963). ** Microlejeunea constricta (Grolle) Grolle, J. Bryol. 21 (1): 41, 1999 (Grolle and Reiner-Drehwald 1999). Bas.: Harpalejeunea constricta Grolle, Nova Hedwigia 16: 150, 1968 (Grolle 1968d). ** Microlejeunea crenulifolia (Gottsche) Steph., Hedwigia 35 (3): 114, 1896 (Stephani 1896b). Bas.: Lejeunea crenulifolia Gottsche, Mexik. Leverm.: 227, 1863 (Gottsche 1863). *** Microlejeunea cystifera Herzog, Memoranda Soc. Fauna Fl. Fennica 25: 68, 1950 (Herzog 1950c). ** Microlejeunea dispar Jovet-Ast, Rev. Bryol. Lichénol. 27 (3/4): 191, 1959 (Jovet-Ast 1959). *** Microlejeunea epiphylla Bischl., Nova Hedwigia 5 (1/2): 378, 1963 (Bischler et al. 1963). *** Microlejeunea ilicuspis (Steph.) Heinrichs, Schäf.-Verw., Pócs et S.Dong, Phytotaxa 85 (2): 52, 2013 (Dong et al. 2013). Bas.: Drepanolejeunea ilicuspis Steph., Sp. Hepat. (Stephani) 5: 344, 1913 (Stephani 1913a). *** Microlejeunea ischeri (Tixier) Heinrichs, Schäf.-Verw., Pócs et S.Dong, Phytotaxa 85 (2): 52, 2013 (Dong et al. 2013). Bas.: Harpalejeunea ischeri Tixier, Trop. Bryol. 11: 29, 1995 (Tixier 1995b). ** Microlejeunea issistipula Steph., Sp. Hepat. (Stephani) 5: 810, 1915 (Stephani 1915b). *** Microlejeunea globosa (Spruce) Steph., Sp. Hepat. (Stephani) 5: 821, 1915 (Stephani 1915b). Bas.: Lejeunea globosa Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: cxciii, 1889 [1890] (Spruce 1889). ** Microlejeunea herzogiana Steph., Biblioth. Bot. 87 (2): 266, 1916 (Stephani 1916a). ** Microlejeunea indica (Udar et U.S.Awasthi) Y.M.Wei et R.L.Zhu, Phytotaxa 97 (2): 63, 2013 (Wei and Zhu 2013b). Bas.: Lejeunea indica Udar et U.S.Awasthi, Cryptog. Bryol. Lichénol. 2 (3): 345, 1981 (Udar and Awasthi 1981). ** Microlejeunea inlata Steph., Sp. Hepat. (Stephani) 5: 811, 1915 (Stephani 1915b). ** Microlejeunea kinabaluensis (Mizut.) Grolle, J. Bryol. 21 (1): 42, 1999 (Grolle and Reiner-Drehwald 1999). Bas.: Harpalejeunea kinabaluensis Mizut., J. Hattori Bot. Lab. 37: 200, 1973 (Mizutani 1973). *** Microlejeunea latitans (Hook.f. et Taylor) Heinrichs, Schäf.-Verw., Pócs et S.Dong, Phytotaxa 85 (2): 52, 2013 (Dong et al. 2013). Bas.: Jungermannia latitans Hook.f. et Taylor, London J. Bot. 3: 399, 1844 (Hooker and Taylor 1844a). ** Microlejeunea lunulatiloba Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 1: 27, 1931 (Horikawa 1931b). 333 * Microlejeunea magnilobula Gola, Ann. Bot. (Rome) 6 (2): 274, 1907 (Gola 1907). 332 Microlejeunea cochlearifolia is conspeciic with Microlejeunea kamerunensis in Jones (1969), but he used the younger name Microlejeunea kamerunensis Steph. 1915. 333 Microlejeunea lunulatiloba is conspeciic with Microlejeunea ulicina in Schuster (1962a), but that is not followed by many recent authors. World checklist of hornworts and liverworts 387 ** Microlejeunea mammillosa (Mizut.) Grolle, J. Bryol. 21 (1): 42, 1999 (Grolle and Reiner-Drehwald 1999). Bas.: Harpalejeunea mammillosa Mizut., J. Hattori Bot. Lab. 37: 192, 1973 (Mizutani 1973). ** Microlejeunea minutissima (Mizut.) Grolle, J. Bryol. 21 (1): 42, 1999 (Grolle and Reiner-Drehwald 1999). Bas.: Harpalejeunea minutissima Mizut., J. Hattori Bot. Lab. 37: 202, 1973 (Mizutani 1973). * Microlejeunea minutistipula Steph., Wiss. Ergebn. Deut. Zentr.-Afr. Exped. (1907–08), Bot. 2: 131, 1911 (Stephani 1911a). 334 *** Microlejeunea moniliata (Mizut.) R.L.Zhu et Y.M.Wei, Cryptog. Bryol. 34 (3): 308, 2013 (Wei and Zhu 2013a). Bas.: Lejeunea moniliata Mizut., J. Hattori Bot. Lab. 46: 357, 1979 (Mizutani 1979b). * Microlejeunea nepalensis Steph., Sp. Hepat. (Stephani) 5: 832, 1915 (Stephani 1915b). ** Microlejeunea nyandaruensis Pócs, Polish Bot. J. 47 (1): 14, 2002 (Pócs 2002c). ** Microlejeunea oblongistipula (Gottsche) Pearson, Forh. Vidensk.-Selsk. Kristiania 1892 (8): 9, 1892 (Pearson 1892). Bas.: Lejeunea oblongistipula Gottsche, Abh. Naturwiss. Vereins Bremen 7: 357, 1882 (Gottsche 1882). ** Microlejeunea ocellata (Herzog) Grolle, Haussknechtia 8: 60, 2001 (Grolle 2001). Bas.: Rectolejeunea ocellata Herzog, Trans. & Proc. Roy. Soc. New Zealand 77 (2): 255, 1949 (Herzog 1949a). * Microlejeunea ovistipula Steph., Wiss. Ergebn. Deut. Zentr.-Afr. Exped. (190708), Bot. 2: 131, 1911 (Stephani 1911a). 335 ** Microlejeunea papulosa (Gottsche) Pearson, Forh. Vidensk.-Selsk. Kristiania 1892 (8): 9, 1892 (Pearson 1892). Bas.: Lejeunea papulosa Gottsche, Abh. Naturwiss. Vereins Bremen 7: 358, 1882 (Gottsche 1882). ** Microlejeunea perpusilla (Spruce) Steph., Hedwigia 35 (3): 113, 1896 (Stephani 1896b). Bas.: Lejeunea perpusilla Spruce, J. Bot. 19: 36, 1881 (Spruce 1881b). ** Microlejeunea punctiformis (Taylor) Steph., Hedwigia 29 (2): 90, 1890 (Stephani 1890b). Bas.: Lejeunea punctiformis Taylor, London J. Bot. 5: 398, 1846 (Taylor 1846b). 336 ** Microlejeunea pyriformis (Lindenb. et Gottsche) Steph., Sp. Hepat. (Stephani) 5: 824, 1915 (Stephani 1915b). Bas.: Lejeunea pyriformis Lindenb. et Gottsche, Syn. Hepat. 5: 767, 1847 (Gottsche et al. 1847). ** Microlejeunea spinosa (Mizut.) Grolle, J. Bryol. 21 (1): 43, 1999 (Grolle and Reiner-Drehwald 1999). Bas.: Harpalejeunea spinosa Mizut., J. Hattori Bot. Lab. 37: 196, 1973 (Mizutani 1973). *** Microlejeunea squarrosa (Steph.) Heinrichs, Schäf.-Verw., Pócs et S.Dong, Phytotaxa 85 (2): 51, 2013 (Dong et al. 2013). Bas.: Strepsilejeunea squarrosa Steph., Hedwigia 35 (3): 130, 1896 (Stephani 1896b). 334 Microlejeunea minutistipula is probably either Microlejeunea cochlearifolia or Microlejeunea africana (Jones 1969). 335 Microlejeunea ovistipula is possibly conspeciic with Lejeunea hepaticola (Jones (1985, Wigginton and Grolle 1996). 336 Microlejeunea punctiformis was treated as conspeciic with Microlejeunea ulicina by Long and Grolle (1990), Fang et al. (1998) and Piippo (1990), but Zhu and So (2001) showed that it merits distinction. 388 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Microlejeunea strasbergii Bardat et Ah-Peng, Bryologist 114 (4): 669, 2011 (AhPeng and Bardat 2011). *** Microlejeunea subulistipa Steph., Hedwigia 35 (3): 115, 1896 (Stephani 1896b). * Microlejeunea szechuanensis P.C.Chen, Feddes Repert. Spec. Nov. Regni Veg. 58: 46, 1955 (Chen 1955). 337 ** Microlejeunea udarii P.K.Verma et S.C.Srivast., J. Bombay Nat. Hist. Soc. 108 (2): 122, 2011 [2012] (Verma and Srivastava 2011). *** Microlejeunea ulicina (Taylor) Steph., Hedwigia 29 (2): 88, 1890 (Stephani 1890b). Bas.: Jungermannia ulicina Taylor, Trans. Bot. Soc. Edinburgh 1 (1/4): 115, 1844 (Taylor 1844a). * Microlejeunea usambarensis Steph., Akad. Wiss. Wien, Math.-Naturwiss. Kl., Denkschr. 88: 730, 1913 (Stephani 1913b). ** Microlejeunea valenciana Steph., Sp. Hepat. (Stephani) 5: 815, 1915 (Stephani 1915b). * Microlejeunea victoriensis D.J.Carr, Proc. Roy. Soc. Victoria 117 (2): 322, 2005 (Carr 2005). *** Microlejeunea wallichiana (Lehm.) R.L.Zhu et Y.M.Wei, Cryptog. Bryol. 34 (3): 308, 2013 (Wei and Zhu 2013a). Bas.: Jungermannia wallichiana Lehm., Nov. Stirp. Pug. 3: 5, 1831 (Lehmann 1831). * ** ** ** ** ** ** ** ** ** * ** ** Taxilejeunea (Spruce) Steph., Hedwigia 28 (4): 262, 1889 (Stephani 1889c) nom. rejic. Bas.: Lejeunea subg. Taxilejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 212, 1884 (Spruce 1884). Taxilejeunea acutifolia (Steph.) Steph., Sp. Hepat. (Stephani) 5: 481, 1914 (Stephani 1914b). Bas.: Dicranolejeunea acutifolia Steph., Hedwigia 35 (3): 76, 1896 (Stephani 1896b). Taxilejeunea antillana Steph., Sp. Hepat. (Stephani) 5: 482, 1914 (Stephani 1914b). Taxilejeunea apiculata (Gottsche) J.B.Jack et Steph., Hedwigia 31 (1): 13, 1892 (Jack and Stephani 1892). Bas.: Omphalanthus apiculatus Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 149, 1864 (Gottsche 1864). Taxilejeunea argentina Steph., Sp. Hepat. (Stephani) 5: 482, 1914 (Stephani 1914b). Taxilejeunea arsenii Steph., Sp. Hepat. (Stephani) 6: 400, 1923 (Stephani 1923). Taxilejeunea auriculata Steph., Sp. Hepat. (Stephani) 5: 459, 1914 (Stephani 1914b). Taxilejeunea berteroana Steph., Sp. Hepat. (Stephani) 5: 483, 1914 (Stephani 1914b). Taxilejeunea beyrichiana Steph., Sp. Hepat. (Stephani) 5: 460, 1914 (Stephani 1914b). Taxilejeunea biapiculata Steph., Sp. Hepat. (Stephani) 5: 460, 1914 (Stephani 1914b). Taxilejeunea boliviana Steph., Biblioth. Bot. 87 (2): 260, 1916 (Stephani 1916a). Taxilejeunea brasiliensis Steph., Hedwigia 35 (3): 132, 1896 (Stephani 1896b). Taxilejeunea coilantha Herzog, Rev. Bryol. Lichénol. 20 (1/2): 159, 1951 [1952] (Herzog 1951a). 337 Microlejeunea szechuanensis may be conspeciic with Microlejeunea punctiformis. World checklist of hornworts and liverworts * * * * * ** ** ** ** * ** * ** ** ** ** ** * * ** * * * ** * ** ** ** ** 389 Taxilejeunea compressiuscula Steph., Sp. Hepat. (Stephani) 5: 501, 1914 (Stephani 1914b). Taxilejeunea convoluta Herzog, Biblioth. Bot. 88: 31, 1920 [1921] (Herzog 1920). Taxilejeunea cuervi (Gottsche) Steph., Sp. Hepat. (Stephani) 5: 486, 1914 (Stephani 1914b). Bas.: Omphalanthus cuervi Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 147, 1864 (Gottsche 1864). Taxilejeunea cuneistipula Steph., Sp. Hepat. (Stephani) 6: 401, 1923 (Stephani 1923). Taxilejeunea cuspidata Steph., Biblioth. Bot. 87 (2): 260, 1916 (Stephani 1916a). Taxilejeunea decurrens Steph., Sp. Hepat. (Stephani) 5: 487, 1914 (Stephani 1914b). Taxilejeunea delexa Steph., Sp. Hepat. (Stephani) 5: 502, 1914 (Stephani 1914b). Taxilejeunea densilora A.Evans, Bull. Torrey Bot. Club 48 (4): 121, 1921 (Evans 1921b). Taxilejeunea diaphana (Lehm.) Steph., Sp. Hepat. (Stephani) 5: 463, 1914 (Stephani 1914b). Bas.: Omphalanthus diaphanus Lehm., Nov. Stirp. Pug. 10: 12, 1857 (Lehmann 1857). Taxilejeunea dissitifolia Steph., Symb. Antill. 2: 472, 1901 (Stephani 1901f). Taxilejeunea eggersiana Schifn., Bot. Jahrb. Syst. 23 (5): 579, 1897 (Schifner 1897). Based on: Taxilejeunea eggersiana Steph., Hedwigia 27 (11/12): 285, 1888 (Stephani 1888c), nom. inval. Taxilejeunea elobulata Sim, Trans. Roy. Soc. South Africa 15 (1): 66, 1926 (Sim 1926). Taxilejeunea issistipula Steph., Sp. Hepat. (Stephani) 5: 488, 1914 (Stephani 1914b). Taxilejeunea foliicola Steph., Sp. Hepat. (Stephani) 5: 466, 1914 (Stephani 1914b). Taxilejeunea furcicornuta Grolle, J. Bryol. 8 (1): 93, 1974 (Grolle 1974a). Taxilejeunea fuscorufa Steph., Hedwigia 35 (3): 133, 1896 (Stephani 1896b). Taxilejeunea galapagensis Onr., Misc. Bryol. Lichenol. 9 (6): 117, 1982 (Onraedt 1982). Taxilejeunea ghatensis P.K.Verma et S.C.Srivast., Proc. Natl. Acad. Sci. India, B 77 (2): 211, 2007 (Verma and Srivastava 2007). Taxilejeunea giulianettii Steph., Sp. Hepat. (Stephani) 5: 502, 1914 (Stephani 1914b). Taxilejeunea gomphocalyx Herzog, Beih. Bot. Centralbl. 61B (3): 580, 1942 (Herzog 1942d). Taxilejeunea grandifolia Steph., Biblioth. Bot. 87 (2): 261, 1916 (Stephani 1916a). Taxilejeunea grandistipula Steph., Sp. Hepat. (Stephani) 5: 504, 1914 (Stephani 1914b). Taxilejeunea hamatifolia Steph., Biblioth. Bot. 87 (2): 261, 1916 (Stephani 1916a). Taxilejeunea himalayensis Herzog, Ann. Bryol. 12: 86, 1939 (Herzog 1939b). Taxilejeunea immersa Eifrig, Monogr. Stud. Indomal. Art. Taxilejeunea: 96, 1937 (Eifrig 1937). Taxilejeunea irregularis Steph., Sp. Hepat. (Stephani) 5: 490, 1914 (Stephani 1914b). Taxilejeunea jamaicensis A.Evans, Bull. Torrey Bot. Club 48 (4): 117, 1921 (Evans 1921b). Taxilejeunea jeringii Steph., Hedwigia 35 (3): 134, 1896 (Stephani 1896b). Taxilejeunea killipii Herzog, Feddes Repert. Spec. Nov. Regni Veg. 57 (1/2): 190, 1955 (Herzog 1955). 390 * * ** ** * ** * * ** * * ** ** ** ** ** * * * ** ** * ** * *** ** ** * ** ** ** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Taxilejeunea laevis (Gottsche) Steph., Hedwigia 44 (4): 229, 1905 (Stephani 1905a). Bas.: Omphalanthus laevis Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 148, 1864 (Gottsche 1864). Taxilejeunea langiana Pearson, Ann. Bryol. 4: 102, 1931 (Pearson 1931b). Taxilejeunea lindbergiana Steph., Sp. Hepat. (Stephani) 5: 491, 1914 (Stephani 1914b). Taxilejeunea linguifolia Steph., Sp. Hepat. (Stephani) 5: 471, 1914 (Stephani 1914b). Taxilejeunea maxima Steph., Sp. Hepat. (Stephani) 5: 473, 1914 (Stephani 1914b). Taxilejeunea mexicana Steph., Sp. Hepat. (Stephani) 6: 402, 1923 (Stephani 1923). Taxilejeunea microstipula Steph., Sp. Hepat. (Stephani) 5: 493, 1914 (Stephani 1914b). Taxilejeunea mucronata Steph., Sp. Hepat. (Stephani) 5: 473, 1914 (Stephani 1914b). Taxilejeunea multilora Steph., Hedwigia 35 (3): 135, 1896 (Stephani 1896b). Taxilejeunea muscicola Steph., Biblioth. Bot. 87 (2): 263, 1916 (Stephani 1916a). Taxilejeunea nilgiriensis P.K.Verma et S.C.Srivast., Proc. Natl. Acad. Sci. India, B 77 (2): 207, 2007 (Verma and Srivastava 2007). Taxilejeunea nymannii Steph., Sp. Hepat. (Stephani) 5: 507, 1914 (Stephani 1914b). Taxilejeunea obtusifolia Steph., Sp. Hepat. (Stephani) 5: 474, 1914 (Stephani 1914b). Taxilejeunea papulilora Steph., Sp. Hepat. (Stephani) 5: 494, 1914 (Stephani 1914b). Taxilejeunea parvibracteata Steph., Hedwigia 35 (3): 136, 1896 (Stephani 1896b). Taxilejeunea parvistipula Steph., Sp. Hepat. (Stephani) 6: 403, 1923 (Stephani 1923). Taxilejeunea paucidens Steph., Biblioth. Bot. 87 (2): 263, 1916 (Stephani 1916a). Taxilejeunea pendula Steph., Biblioth. Bot. 87 (2): 263, 1916 (Stephani 1916a). Taxilejeunea peruviana Steph., Sp. Hepat. (Stephani) 5: 475, 1914 (Stephani 1914b). Taxilejeunea planilobula Herzog, Rev. Bryol. Lichénol. 20 (1/2): 160, 1951 [1952] (Herzog 1951a). Taxilejeunea pulchrilora Pearson, Ark. Bot. 19 (5): 15, 1924 (Pearson 1924b). Taxilejeunea pusilla Steph., Biblioth. Bot. 87 (2): 264, 1916 (Stephani 1916a). Taxilejeunea renistipula (Lindenb.) Steph., Hedwigia 29 (3): 142, 1890 (Stephani 1890d). Bas.: Omphalanthus renistipulus Lindenb., Syn. Hepat. 3: 308, 1845 (Gottsche et al. 1845b). Taxilejeunea rufescens Steph., Biblioth. Bot. 87 (2): 264, 1916 (Stephani 1916a). Taxilejeunea serpillifolioides (Raddi) D.P.Costa, J. Bryol. 31 (4): 230, 2009 (Costa 2009). Bas.: Jungermannia serpillifolioides Raddi, Critt. Brasil.: 17, 1822 (Raddi 1822). Taxilejeunea setchellii Pearson, Amer. Samoa: 146, 1924 (Pearson 1924a). Taxilejeunea speciosa Herzog, Feddes Repert. Spec. Nov. Regni Veg. 57 (1/2): 188, 1955 (Herzog 1955). Taxilejeunea splendida Eifrig, Monogr. Stud. Indomal. Art. Taxilejeunea: 88, 1937 (Eifrig 1937). Taxilejeunea stephanii Eifrig, Monogr. Stud. Indomal. Art. Taxilejeunea: 90, 1937 (Eifrig 1937). Nom. nov. pro Hygrolejeunea nymanii Steph., Sp. Hepat. (Stephani) 5: 564, 1914 (Stephani 1914b). Taxilejeunea steyermarkii H.Rob., Phytologia 34 (1): 67, 1976 (Robinson 1976a). Taxilejeunea surinamensis (Lindenb. et Gottsche) Steph., Hedwigia 29 (3): 142, 1890 (Stephani 1890d). Bas.: Omphalanthus surinamensis Lindenb. et Gottsche, Linnaea 24 (6): 628, 1851 [1852] (Lindenberg and Gottsche 1851a). World checklist of hornworts and liverworts 391 * ** ** * Taxilejeunea suringarii Steph., Sp. Hepat. (Stephani) 5: 479, 1914 (Stephani 1914b). Taxilejeunea tenerrima Steph., Sp. Hepat. (Stephani) 6: 406, 1923 (Stephani 1923). Taxilejeunea tenuiplica Steph., Sp. Hepat. (Stephani) 5: 480, 1914 (Stephani 1914b). Taxilejeunea tjibodensis (Steph.) Eifrig, Monogr. Stud. Indomal. Art. Taxilejeunea: 91, 1937 (Eifrig 1937). Bas.: Hygrolejeunea tjibodensis Steph., Sp. Hepat. (Stephani) 5: 571, 1914 (Stephani 1914b). ** Taxilejeunea tonduzana Steph., Sp. Hepat. (Stephani) 5: 498, 1914 (Stephani 1914b). * Taxilejeunea uleana Steph., Hedwigia 35 (3): 136, 1896 (Stephani 1896b). * Taxilejeunea umbonata Steph., Sp. Hepat. (Stephani) 5: 481, 1914 (Stephani 1914b). * Taxilejeunea urbanii Steph., Symb. Antill. (Urban) 3 (2): 278, 1902 (Stephani 1902e). ** Taxilejeunea vallis-gratiae Steph., Hedwigia 35 (3): 137, 1896 (Stephani 1896b). ** subtrib. Lepidolejeuneinae Gradst. *** Lepidolejeunea R.M.Schust., Beih. Nova Hedwigia 9: 139, 1963 (Schuster 1963a). 338 ** subg. Kingiolejeunea (H.Rob.) R.M.Schust., Phytologia 45 (5): 425, 1980 (Schuster 1980b). Bas.: Kingiolejeunea H.Rob., Bryologist 70 (1): 53, 1967 (Robinson 1967). ** Lepidolejeunea auriculata Schäf.-Verw. et Heinrichs, Taxon 64 (2): 224, 2015 (Heinrichs et al. 2015). *** Lepidolejeunea cordiissa (Taylor) M.E.Reiner, Nova Hedwigia 83 (3/4): 478, 2006 (Reiner-Drehwald 2006). Bas.: Lejeunea cordiissa Taylor, London J. Bot. 5: 395, 1846 (Taylor 1846b). *** Lepidolejeunea grossepapulosa (Steph.) Piippo, Acta Bot. Fenn. 132: 49, 1986 (Piippo 1986a). Bas.: Prionolejeunea grossepapulosa Steph., Sp. Hepat. (Stephani) 5: 220, 1913 (Stephani 1913a). *** Lepidolejeunea involuta (Gottsche) Grolle, J. Hattori Bot. Lab. 55: 504, 1984 (Grolle 1984a). Bas.: Lejeunea involuta Gottsche, Syn. Hepat. 3: 350, 1845 (Gottsche et al. 1845b). ** subg. Lepidolejeunea *** Lepidolejeunea bidentula (Steph.) R.M.Schust., Phytologia 45 (5): 425, 1980 (Schuster 1980b). Bas.: Pycnolejeunea bidentula Steph., Bot. Centralbl. 60 (4): 107, 1894 (Jack and Stephani 1894). *** Lepidolejeunea bidentula var. novae-caledoniae Piippo, Acta Bot. Fenn. 132: 26, 1986 (Piippo 1986a). *** Lepidolejeunea falcata (Herzog) R.M.Schust., Beih. Nova Hedwigia 9: 139, 1963 (Schuster 1963a). Bas.: Hygrolejeunea falcata Herzog, Ark. Bot. (n.ser.) 3 (3): 57, 1953 (Herzog 1953a). 338 Lepidolejeunea includes Kingiolejeunea, but one taxon has neither been transferred nor synonymized. It is listed in the “Names in genera not currently accepted” section below. 392 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Lepidolejeunea graefei (J.B.Jack et Steph.) R.M.Schust., Phytologia 45 (5): 425, 1980 (Schuster 1980b). Bas.: Archilejeunea graefei J.B.Jack et Steph., Bot. Centralbl. 60 (4): 104, 1894 (Jack and Stephani 1894). *** Lepidolejeunea integristipula (J.B.Jack et Steph.) R.M.Schust., Phytologia 45 (5): 425, 1980 (Schuster 1980b). Bas.: Pycnolejeunea integristipula J.B.Jack et Steph., Bot. Centralbl. 60 (4): 107, 1894 (Jack and Stephani 1894). ** subg. Perilejeunea (Kachroo et R.M.Schust.) R.M.Schust., Phytologia 45 (5): 424, 1980 (Schuster 1980b). Bas.: Pycnolejeunea subg. Perilejeunea Kachroo et R.M.Schust., J. Linn. Soc., Bot. 56 (368): 493, 1961 (Kachroo and Schuster 1961). ** Lepidolejeunea cuspidata (Gottsche) Heinrichs et Schäf.-Verw., Taxon 64 (2): 224, 2015 (Heinrichs et al. 2015). Bas.: Lejeunea cuspidata Gottsche, Syn. Hepat. 3: 351, 1845 (Gottsche et al. 1845b). *** Lepidolejeunea delessertii (Nees et Mont.) Grolle, J. Hattori Bot. Lab. 55: 505, 1984 (Grolle 1984a). Bas.: Lejeunea delessertii Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 260, 1843 (Montagne 1843). *** Lepidolejeunea eluta (Nees) R.M.Schust., Beih. Nova Hedwigia 9: 139, 1963 (Schuster 1963a). Bas.: Jungermannia eluta Nees, Fl. Bras. (Martius) 1 (1): 362, 1833 (Nees 1833a). *** Lepidolejeunea serrulata (Steph.) Grolle, J. Hattori Bot. Lab. 55: 505, 1984 (Grolle 1984a). Bas.: Trachylejeunea serrulata Steph., Sp. Hepat. (Stephani) 5: 300, 1913 (Stephani 1913a). *** Lepidolejeunea sullivantii (Gottsche) M.E.Reiner, Nova Hedwigia 83 (3/4): 479, 2006 (Reiner-Drehwald 2006). Bas.: Lejeunea sullivantii Gottsche, Mexik. Leverm.: 196, 1863 (Gottsche 1863). Incertae sedis *** Lepidolejeunea borneensis (Steph.) R.M.Schust., Phytologia 45 (5): 425, 1980 (Schuster 1980b). Bas.: Hygrolejeunea borneensis Steph., Sp. Hepat. (Stephani) 5: 557, 1914 (Stephani 1914b). *** Lepidolejeunea longilobula (Amakawa) Xiao L.He, Acta Bot. Fenn. 163: 59, 1999 (He 1999). Bas.: Pycnolejeunea longilobula Amakawa, J. Jap. Bot. 40 (10): 307, 1965 (Amakawa 1965). *** Otolejeunea Grolle et Tixier, Nova Hedwigia 32: 609, 1980 (Tixier 1980b). *** subg. Allorgella (Tixier) Grolle, Haussknechtia 2: 53, 1985 (Grolle 1985a). Bas.: Allorgella Tixier, Nova Hedwigia 32: 612, 1980 (Tixier 1980b). *** Otolejeunea australiensis B.M.hiers, Brittonia 44 (2): 162, 1992 (hiers 1992a). *** Otolejeunea hoana (Tixier) Grolle, Haussknechtia 2: 54, 1985 [1986] (Grolle 1985a). Bas.: Allorgella hoana Tixier, Nova Hedwigia 32: 615, 1980 (Tixier 1980b). *** Otolejeunea rabenorii Tixier, Nova Hedwigia 46 (3/4): 376, 1988 (Tixier 1988). *** Otolejeunea schmidii (Tixier) Grolle, Haussknechtia 2: 54, 1985 [1986] (Grolle 1985a). Bas.: Allorgella schmidii Tixier, Nova Hedwigia 32: 613, 1980 (Tixier 1980b). World checklist of hornworts and liverworts 393 *** Otolejeunea schnellii (Tixier) R.L.Zhu et M.L.So, Ann. Bot. Fenn. 34 (4): 287, 1997 (Zhu and So 1997a). Bas.: Allorgella schnellii Tixier, Cryptog. Bryol. Lichenol. 16 (3): 230, 1995 (Tixier 1995a). *** Otolejeunea semperiana (Steph.) Grolle, Haussknechtia 2: 53, 1985 [1986] (Grolle 1985a). Bas.: Prionolejeunea semperiana Steph., Sp. Hepat. (Stephani) 5: 227, 1913 (Stephani 1913a). ** Otolejeunea subana Pocs, Acta Acad. Ped. Agr., Sect. Biol. 25: 50, 2004 (Pocs 2004). *** Otolejeunea zantenii Grolle, Haussknechtia 2: 54, 1985 [1986] (Grolle 1985a). *** subg. Otolejeunea *** Otolejeunea moniliata Grolle, Nova Hedwigia 32: 609, 1980 (Tixier 1980b). World checklist of hornworts and liverworts 353 *** subg. Phoxolejeunea Grolle, Haussknechtia 2: 49, 1985 (Grolle 1985a). ** Otolejeunea philippinensis R.L.Zhu et M.L.So, Syst. Bot. 23 (2): 231, 1998 (Zhu and So 1998). *** Otolejeunea streimannii Grolle, Haussknechtia 2: 49, 1985 [1986] (Grolle 1985a). *** Rectolejeunea A.Evans, Bull. Torrey Bot. Club 33 (1): 8, 1906 (Evans 1906a). * Rectolejeunea colombiana R.M.Schust., J. Hattori Bot. Lab. 89: 146, 2000 (Schuster 2000c). *** Rectolejeunea emarginulilora (Schifn.) A.Evans, Bull. Torrey Bot. Club 33 (1): 14, 1906 (Evans 1906a). Bas.: Cheilolejeunea emarginulilora Schifn., Bot. Jahrb. Syst. 23 (5): 585, 1897 (Schifner 1897). *** Rectolejeunea lagelliformis A.Evans, Bull. Torrey Bot. Club 33 (1): 9, 1906 (Evans 1906a). * Rectolejeunea lagelliformis subsp. hamata R.M.Schust., J. Hattori Bot. Lab. 89: 137, 2000 (Schuster 2000c). *** Rectolejeunea queenslandica (B.M.hiers) Xiao L.He, Ann. Bot. Fenn. 34 (2): 129, 1997 (He 1997). Bas.: Lepidolejeunea queenslandica B.M.hiers, Mem. New York Bot. Gard. 45: 556, 1987 (hiers 1987a). ** Rectolejeunea truncatilobula C.J.Bastos, J. Bryol. 34 (2): 144, 2012 (Bastos 2012a). *** Rectolejeunea versifolia (Schifn.) L.Söderstr. et A.Hagborg, Phytotaxa 220 (2): 188, 2015 (Söderström et al. 2015a). Bas.: Cheilolejeunea versifolia Schifn., Bot. Jahrb. Syst. 23 (5): 597, 1897 (Schifner 1897). * Excluded from the genus Rectolejeunea lindenbergii Steph., Sp. Hepat. (Stephani) 5: 689, 1914 (Stephani 1914b). 339 339 Rectolejeunea lindenbergii is a Lejeunea species (Reiner-Drehwald and Grolle 2012; non Lejeunea lindenbergii Gottsche). However, it may be conspeciic with something else. 394 * * ** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Rectolejeunea lindigiana Steph., Sp. Hepat. (Stephani) 5: 690, 1914 (Stephani 1914b). 340 Rectolejeunea pachyderma R.M.Schust., J. Hattori Bot. Lab. 89: 148, 2000 (Schuster 2000c). 341 subtrib. Leptolejeuneinae Heinrichs et Schäf.-Verw. *** Leptolejeunea (Spruce) Steph., Hedwigia 30 (6): 270, 1891 (Stephani 1891c). Bas.: Lejeunea subg. Leptolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 193, 1884 (Spruce 1884). *** Leptolejeunea amphiophthalma Zwickel, Ann. Bryol. 6: 117, 1933 (Zwickel 1933). ** Leptolejeunea apiculata (Horik.) S.Hatt., J. Hattori Bot. Lab. 5: 46, 1951 (Hattori 1951d). Bas.: Drepanolejeunea apiculata Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 266, 1934 (Horikawa 1934). ** Leptolejeunea arunachalensis Sudipa Das et D.K.Singh, J. Jap. Bot. 83 (6): 343, 2008 (Das and Singh 2008). *** Leptolejeunea astroidea (Mitt.) Steph., Sp. Hepat. (Stephani) 5: 363, 1913 (Stephani 1913a). Bas.: Lejeunea astroidea Mitt., Trans. Linn. Soc. London 23 (1): 58, 1860 (Mitten 1860a). ** Leptolejeunea australis Steph., Sp. Hepat. (Stephani) 5: 389, 1913 (Stephani 1913a). Based on: Leptolejeunea australis Steph., Hedwigia 28 (3): 173, 1889 (Stephani 1889d), nom. inval. ** Leptolejeunea balansae Steph., Hedwigia 35 (3): 105, 1896 (Stephani 1896b). * Leptolejeunea borneensis Herzog, Flora 135: 394, 1942 (Herzog 1942c). 342 ** Leptolejeunea brasiliensis Bischl., Nova Hedwigia 17: 301, 1969 (Bischler 1969). * Leptolejeunea convexistipa Bischl., Nova Hedwigia 17: 325, 1969 (Bischler 1969). * Leptolejeunea curvatifolia Steph., Sp. Hepat. (Stephani) 6: 398, 1923 (Stephani 1923). 343 ** Leptolejeunea denticulata Steph., Sp. Hepat. (Stephani) 5: 389, 1913 (Stephani 1913a). Based on: Leptolejeunea denticulata Steph., Hedwigia 28 (3): 174, 1889 (Stephani 1889d), nom. inval. ** Leptolejeunea dentistipula Steph., Sp. Hepat. (Stephani) 5: 379, 1913 (Stephani 1913a). ** Leptolejeunea diversilobulata Bischl., Nova Hedwigia 17: 313, 1969 (Bischler 1969). ** Leptolejeunea dolabriformis Pearson, J. Linn. Soc., Bot. 46 (305): 37, 1922 (Pearson 1922b). 340 341 342 343 Rectolejeunea lindigiana is a Lejeunea species of doubtful status (Reiner-Drehwald and Grolle 2012). Rectolejeunea pachyderma is a Lejeunea species of doubtful status (Reiner-Drehwald and Grolle 2012). Leptolejeunea borneensis may be conspeciic with Leptolejeunea vitrea. Leptolejeunea curvatifolia is possibly conspeciic with Drepanolejeunea thwaitesiana, but the type specimen was burned in B and the identity can not be ascertained (Grolle and Piippo 1984). World checklist of hornworts and liverworts 395 *** Leptolejeunea elliptica (Lehm. et Lindenb.) Besch., Rev. Bryol. 19 (1): 14, 1892 (Bescherelle 1892). Bas.: Jungermannia elliptica Lehm. et Lindenb., Nov. Stirp. Pug. 5: 13, 1833 (Lehmann 1833). * Leptolejeunea emarginata (Horik.) S.Hatt., J. Hattori Bot. Lab. 5: 46, 1951 (Hattori 1951d). Bas.: Drepanolejeunea emarginata Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 267, 1934 (Horikawa 1934). 344 *** Leptolejeunea epiphylla (Mitt.) Steph., Sp. Hepat. (Stephani) 5: 380, 1913 (Stephani 1913a). Bas.: Lejeunea epiphylla Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 118, 1860 [1861] (Mitten 1860c). *** Leptolejeunea exocellata (Spruce) A.Evans, Bull. Torrey Bot. Club 29 (8): 498, 1902 (Evans 1902c). Bas.: Lejeunea exocellata Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 195, 1884 (Spruce 1884). ** Leptolejeunea foliicola Steph., Hedwigia 35 (3): 106, 1896 (Stephani 1896b). ** Leptolejeunea integristipula Steph., Sp. Hepat. (Stephani) 6: 398, 1923 (Stephani 1923). * Leptolejeunea jamaicensis R.M.Schust., J. Elisha Mitchell Sci. Soc. 83 (4): 229, 1967 (Schuster 1967a). ** Leptolejeunea lancifolia Steph., Sp. Hepat. (Stephani) 5: 382, 1913 (Stephani 1913a). Nom. nov. pro Lejeunea lancifolia Mitt., Fl. vit.: 415, 1871 [1873] (Mitten 1871), nom. illeg. ** Leptolejeunea latifolia Herzog, Memoranda Soc. Fauna Fl. Fennica 26: 58, 1950 [1951] (Herzog 1950b). ** Leptolejeunea lepinii Steph., Sp. Hepat. (Stephani) 5: 383, 1913 (Stephani 1913a). ** Leptolejeunea ligulata Herzog, Flora 135: 429, 1942 (Herzog 1942c). *** Leptolejeunea maculata (Mitt.) Schifn., Consp. Hepat. Arch. Ind.: 275, 1898 (Schifner 1898b). Bas.: Lejeunea maculata Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 118, 1860 [1861] (Mitten 1860c). * Leptolejeunea massartiana Schifn. ex Herzog, Flora 135: 421, 1942 (Herzog 1942c). 345 ** Leptolejeunea micronesica Inoue et H.A.Mill., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 8 (2): 149, 1965 (Inoue and Miller 1965). ** Leptolejeunea minima Herzog, Memoranda Soc. Fauna Fl. Fennica 26: 60, 1950 [1951] (Herzog 1950b). ** Leptolejeunea mirikana M.Dey et D.K.Singh, Taiwania 55 (4): 355, 2010 (Dey and Singh 2010). *** Leptolejeunea moniliata Steph., Sp. Hepat. (Stephani) 5: 371, 1913 (Stephani 1913a). *** Leptolejeunea obfuscata (Spruce) Steph., Sp. Hepat. (Stephani) 5: 373, 1913 (Stephani 1913a). Bas.: Lejeunea obfuscata Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 579, 1885 (Spruce 1885). * Leptolejeunea punctata Herzog, Flora 135: 432, 1942 (Herzog 1942c). 344 Leptolejeunea emarginata is possibly conspeciic with Leptolejeunea apiculata (Zhu and So 2001). 345 Leptolejeunea massartiana is very possibly conspeciic with Leptolejeunea elliptica judging from the original description and illustration (Söderström et al. 2010a). 396 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Leptolejeunea radicosa (Nees ex Mont.) Grolle, J. Hattori Bot. Lab. 45: 178, 1979 (Grolle 1979c). Bas.: Lejeunea radicosa Nees ex Mont., Hist. Phys. Cuba, Bot., Pl. Cell.: 475, 1842 (Montagne 1842a). * Leptolejeunea renneri Herzog, Flora 135: 422, 1942 (Herzog 1942c). * Leptolejeunea revoluta P.C.Chen, Feddes Repert. Spec. Nov. Regni Veg. 58: 44, 1955 (Chen 1955). 346 ** Leptolejeunea rosulans Steph., Sp. Hepat. (Stephani) 5: 390, 1913 (Stephani 1913a). Based on: Leptolejeunea rosulans Steph., Hedwigia 28 (3): 174, 1889 (Stephani 1889d), nom. inval. ** Leptolejeunea serratifolia Schifn., Bot. Jahrb. Syst. 23 (5): 594, 1897 (Schifner 1897). ** Leptolejeunea serrulata Herzog, Flora 135: 426, 1942 (Herzog 1942c). ** Leptolejeunea spinistipula (Mizut.) Xiao L.He, Ann. Bot. Fenn. 34 (2): 127, 1997 (He 1997). Bas.: Pycnolejeunea spinistipula Mizut., J. Hattori Bot. Lab. 33: 255, 1970 (Mizutani 1970). ** Leptolejeunea subdentata Schifn. ex Herzog, Flora 135: 403, 1942 (Herzog 1942c). *** Leptolejeunea subrotundifolia Herzog, Flora 135: 424, 1942 (Herzog 1942c). *** Leptolejeunea tridentata Bischl., Nova Hedwigia 17: 335, 1969 (Bischler 1969). * Leptolejeunea trigonostipa (Spruce) Steph., Sp. Hepat. (Stephani) 5: 376, 1913 (Stephani 1913a). Bas.: Lejeunea trigonostipa Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 197, 1884 (Spruce 1884). ** Leptolejeunea tripuncta (Mitt.) Steph., Sp. Hepat. (Stephani) 5: 388, 1913 (Stephani 1913a). Bas.: Lejeunea tripuncta Mitt., Fl. vit.: 415, 1871 [1873] (Mitten 1871). ** Leptolejeunea truncatifolia Steph., Sp. Hepat. (Stephani) 5: 388, 1913 (Stephani 1913a). * Leptolejeunea udarii M.Dey et D.K.Singh, Taiwania 55 (4): 359, 2010 (Dey and Singh 2010). 347 *** Leptolejeunea vitrea (Nees) Schifn., Hepat. (Engl.-Prantl): 126, 1893 (Schifner 1893b). Bas.: Jungermannia vitrea Nees, Enum. Pl. Crypt. Javae: 56, 1830 (Nees 1830). ** subtrib. Pycnolejeuneinae Heinrichs et Schäf.Verw. ** Pycnolejeunea (Spruce) Schifn., Hepat. (Engl.-Prantl): 124, 1893 (Schifner 1893b). Bas.: Lejeunea subg. Pycnolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 246, 1884 (Spruce 1884). * Pycnolejeunea anotomensis Tixier, Bull. Soc. Hist. Nat. Afrique N. 63: 10, 1972 (Tixier 1972a). 348 ** Pycnolejeunea borneensis Steph., Sp. Hepat. (Stephani) 5: 632, 1914 (Stephani 1914b). 346 Leptolejeunea revoluta may be conspeciic with Leptolejeunea elliptica (Zhu and So 2001). 347 Leptolejeunea udarii is possibly conspeciic with Leptolejeunea latifolia (He et al. 2013). 348 Pycnolejeunea anotomensis is a Cheilolejeunea species. World checklist of hornworts and liverworts * * *** * *** *** ** ** * *** *** ** * *** ** ** * 397 Pycnolejeunea cavistipula (Steph.) Mizut., J. Hattori Bot. Lab. 36: 161, 1972 [1973] (Mizutani 1972a). Bas.: Strepsilejeunea cavistipula Steph., Hedwigia 35 (3): 128, 1896 (Stephani 1896b). 349 Pycnolejeunea connivens Schifn. ex P.Syd., Just’s Bot. Jahresber. 19: 246, 1894 (Sydow 1894). Based on: Pycnolejeunea connivens Gottsche ex Schifn., Leberm., Forschungsr. Gazelle 4 (4): 32, 1890 (Schifner 1890), nom. inval. Pycnolejeunea contigua (Nees) Grolle, J. Hattori Bot. Lab. 45: 179, 1979 (Grolle 1979c). Bas.: Jungermannia contigua Nees, Fl. Bras. (Martius) 1 (1): 360, 1833 (Nees 1833a). Pycnolejeunea convexifolia (Mitt.) Steph., Sp. Hepat. (Stephani) 5: 636, 1914 (Stephani 1914b). Bas.: Lejeunea convexifolia Mitt., Fl. vit.: 416, 1871 [1873] (Mitten 1871). 350 Pycnolejeunea decurviloba Steph., Hedwigia 35 (3): 125, 1896 (Stephani 1896b). Pycnolejeunea densistipula (Lehm. et Lindenb.) Steph., Sp. Hepat. (Stephani) 5: 602, 1914 (Stephani 1914b). Bas.: Lejeunea densistipula Lehm. et Lindenb., Nov. Stirp. Pug. 7: 20, 1838 (Lehmann 1838). Pycnolejeunea gradsteinii Ilk.-Borg., Bol. Inst. Bot. (São Paulo) 21 (1): 1, 2011 (Ilkiu-Borges 2011). Pycnolejeunea grandiocellata Steph., Bot. Tidsskr. 24 (3): 279, 1902 (Stephani 1902b). Pycnolejeunea grossiloba Steph., Sp. Hepat. (Stephani) 5: 629, 1914 (Stephani 1914b). 351 Pycnolejeunea macroloba (Nees et Mont.) Schifn., Hepat. (Engl.-Prantl): 124, 1893 (Schifner 1893b). Bas.: Lejeunea macroloba Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 260, 1843 (Montagne 1843). Pycnolejeunea monophthalma (R.M.Schust.) Xiao L.He, Acta Bot. Fenn. 163: 52, 1999 (He 1999). Bas.: Trachylejeunea monophthalma R.M.Schust., Bull. Torrey Bot. Club 97 (6): 345, 1970 [1971] (Schuster 1970b). Pycnolejeunea novae-caledoniae (Steph.) Horik., Acta Phytotax. Geobot. 13: 214, 1943 (Horikawa 1943). Bas.: Archilejeunea novae-caledoniae Steph., Sp. Hepat. (Stephani) 4: 729, 1911 (Stephani 1911e). Pycnolejeunea palmicola Steph., Sp. Hepat. (Stephani) 6: 413, 1923 (Stephani 1923). 352 Pycnolejeunea papillosa Xiao L.He, Acta Bot. Fenn. 163: 55, 1999 (He 1999). Pycnolejeunea porrectilobula C.J.Bastos et O.Yano, Nova Hedwigia 74 (3/4): 440, 2002 (Bastos and Yano 2002). Pycnolejeunea retusa R.M.Schust., J. Hattori Bot. Lab. 100: 402, 2006 (Schuster 2006). Pycnolejeunea schlimiana Steph., Sp. Hepat. (Stephani) 5: 615, 1914 (Stephani 1914b). 353 349 Pycnolejeunea cavistipula is possibly a Cheilolejeunea species. 350 Pycnolejeunea convexifolia is possibly conspeciic with Cheilolejeunea imbricata. 351 Pycnolejeunea grossiloba is a Cheilolejeunea species, but its taxonomic status is unclear (Grolle and Piippo 1984). 352 Pycnolejeunea palmicola is not a Pycnolejeunea species. 353 Pycnolejeunea schlimiana is not a Pycnolejeunea species. 398 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Pycnolejeunea schwaneckei (Steph.) Schifn. ex P.Syd., Just’s Bot. Jahresber. 19: 246, 1894 (Sydow 1894). Bas.: Lejeunea schwaneckei Steph., Hedwigia 27 (11/12): 290, 1888 (Stephani 1888c). ** Pycnolejeunea sphaeroides (Sande Lac.) J.B.Jack et Steph., Bot. Centralbl. 60 (4): 107, 1894 (Jack and Stephani 1894). Bas.: Lejeunea sphaeroides Sande Lac., Ann. Mus. Bot. Lugduno-Batavi 1: 309, 1864 (Sande Lacoste 1864). ** subtrib. Xylolejeuneinae Heinrichs et Schäf.Verw. *** Xylolejeunea Xiao L.He et Grolle, Ann. Bot. Fenn. 38 (1): 27, 2001 (He and Grolle 2001). *** Xylolejeunea aquarius (Spruce) Xiao L.He et Grolle, Ann. Bot. Fenn. 38 (1): 29, 2001 (He and Grolle 2001). Bas.: Lejeunea aquarius Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 185, 1884 (Spruce 1884). *** Xylolejeunea crenata (Nees et Mont.) Xiao L.He et Grolle, Ann. Bot. Fenn. 38 (1): 36, 2001 (He and Grolle 2001). Bas.: Lejeunea crenata Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 9: 48, 1838 (Montagne 1838). *** Xylolejeunea grolleana (Pócs) Xiao L.He et Grolle, Ann. Bot. Fenn. 38 (1): 32, 2001 (He and Grolle 2001). Bas.: Trachylejeunea grolleana Pócs, Haussknechtia, Beih. 9: 285, 1999 (Pócs 1999). *** Xylolejeunea muricella Xiao L.He et Grolle, Ann. Bot. Fenn. 38 (1): 34, 2001 (He and Grolle 2001). ** trib. Symbiezidieae Gradst. *** Symbiezidium Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 402, 1877 (Trevisan 1877). *** Symbiezidium barbilorum (Lindenb. et Gottsche) A.Evans, Bull. Torrey Bot. Club 34 (11): 540, 1907 [1908] (Evans 1907a). Bas.: Lejeunea barbilora Lindenb. et Gottsche, Linnaea 24 (6): 630, 1851 [1852] (Lindenberg and Gottsche 1851a). *** Symbiezidium dentatum Herzog, Feddes Repert. Spec. Nov. Regni Veg. 57 (1/2): 175, 1955 (Herzog 1955). *** Symbiezidium madagascariense Steph., Sp. Hepat. (Stephani) 5: 99, 1912 (Stephani 1912c). *** Symbiezidium transversale (Sw.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 403, 1877 (Trevisan 1877). Bas.: Jungermannia transversalis Sw., Prodr. (Swartz): 144, 1788 (Swartz 1788). *** Symbiezidium transversale var. hookerianum (Gottsche, Lindenb. et Nees) Gradst. et J.Beek, Beih. Nova Hedwigia 80: 237, 1985 (Gradstein and van Beek 1985). Bas.: Lejeunea transversalis β hookeriana Gottsche, Lindenb. et Nees, Syn. Hepat. 3: 311, 1845 (Gottsche et al. 1845b). World checklist of hornworts and liverworts *** 399 Ptychanthoideae Mizut. *** Acrolejeunea (Spruce) Schifn., Hepat. (Engl.-Prantl): 128, 1893 (Schifner 1893b) nom. conserv. Bas.: Lejeunea subg. Acrolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 115, 1884 (Spruce 1884). *** Acrolejeunea sandvicensis (Gottsche) Steph., Bot. Jahrb. Syst. 23 (1/2, 3): 312, 1896 (Stephani 1896a). Bas.: Phragmicoma sandvicensis Gottsche, Ann. Sci. Nat. Bot. (sér. 4) 8: 344, 1857 (Gottsche 1857). ** sect. Acrolejeunea *** Acrolejeunea emergens (Mitt.) Steph., Planzenw. Ost-Afrikas C: 65, 1895 (Stephani 1895d). Bas.: Phragmicoma emergens Mitt., Philos. Trans. 168: 397, 1879 (Mitten 1879). *** Acrolejeunea emergens var. confertissima (Steph.) Gradst., Bryophyt. Biblioth. 4: 76, 1975 (Gradstein 1975). Bas.: Acrolejeunea confertissima Steph., Hedwigia 31 (4): 165, 1892 (Stephani 1892g). *** Acrolejeunea heterophylla (A.Evans) Grolle et Gradst., J. Hattori Bot. Lab. 38: 332, 1974 (Gradstein 1974a). Bas.: Ptychocoleus heterophyllus A.Evans, Amer. J. Bot. 5 (3): 144, 1918 (Evans 1918). *** Acrolejeunea torulosa (Lehm. et Lindenb.) Schifn., Hepat. (Engl.-Prantl): 128, 1893 (Schifner 1893b). Bas.: Jungermannia torulosa Lehm. et Lindenb., Nov. Stirp. Pug. 6: 41, 1834 (Lehmann 1834). ** sect. Minores (Verd.) L.Söderstr. et A.Hagborg, Bryophyte Diversity Evol. 36 (1): 41, 2014 (Wang et al. 2014b). Bas.: Ptychocoleus sect. Minores Verd., Ann. Bryol., Suppl. 4: 132, 1934 (Verdoorn 1934a). *** Acrolejeunea arcuata (Nees) Grolle et Gradst., J. Hattori Bot. Lab. 38: 332, 1974 (Gradstein 1974a). Bas.: Jungermannia arcuata Nees, Enum. Pl. Crypt. Javae: 38, 1830 (Nees 1830). ** Acrolejeunea arcuata subsp. gradsteinii M.A.M.Renner, Phytotaxa 83 (1): 42, 2013 (Renner 2013). *** Acrolejeunea fertilis (Reinw., Blume et Nees) Schifn., Hepat. (Engl.-Prantl): 128, 1893 (Schifner 1893b). Bas.: Jungermannia fertilis Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 211, 1824 [1825] (Reinwardt et al. 1824a). *** Acrolejeunea parvula (Mizut.) Gradst., Bryophyt. Biblioth. 4: 115, 1975 (Gradstein 1975). Bas.: Ptychocoleus parvulus Mizut., Dansk Bot. Ark. 27 (1): 97, 1969 (Hattori and Mizutani 1969). *** Acrolejeunea pycnoclada (Taylor) Schifn., Hepat. (Engl.-Prantl): 128, 1893 (Schifner 1893b). Bas.: Ptychanthus pycnocladus Taylor, London J. Bot. 5: 385, 1846 (Taylor 1846b). *** Acrolejeunea pycnoclada subsp. latistipula Gradst., Bryophyt. Biblioth. 4: 113, 1975 (Gradstein 1975). 400 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Acrolejeunea tjibodensis (Verd.) Grolle et Gradst., J. Hattori Bot. Lab. 38: 333, 1974 (Gradstein 1974a). Bas.: Ptychocoleus tjibodensis Verd., Recueil Trav. Bot. Neerl. 30: 227, 1933 (Verdoorn 1933b). ** sect. Pusillae Gradst., Bryophyt. Biblioth. 4: 59, 1975 (Gradstein 1975). *** Acrolejeunea pusilla (Steph.) Grolle et Gradst., J. Hattori Bot. Lab. 38: 332, 1974 (Gradstein 1974a). Bas.: Archilejeunea pusilla Steph., Sp. Hepat. (Stephani) 4: 731, 1911 (Stephani 1911e). *** Acrolejeunea sikkimensis (Mizut.) Gradst., Bryophyt. Biblioth. 4: 83, 1975 (Gradstein 1975). Bas.: Ptychocoleus sikkimensis Mizut., Fl. E. Himalaya: 532, 1966 (Hattori 1966c). ** sect. Recurvatae Jian Wang bis et Gradst., Bryophyte Diversity Evol. 36 (1): 39, 2014 (Wang et al. 2014b). *** Acrolejeunea recurvata Gradst., Bryophyt. Biblioth. 4: 79, 1975 (Gradstein 1975). ** sect. Regulares (Verd.) Gradst., Bryophyt. Biblioth. 4: 63, 1975 (Gradstein 1975). Bas.: Ptychocoleus sect. Regulares Verd., Ann. Bryol., Suppl. 4: 143, 1934 (Verdoorn 1934a). *** Acrolejeunea allisonii Gradst., Bryophyt. Biblioth. 4: 103, 1975 (Gradstein 1975). *** Acrolejeunea aulacophora (Mont.) Steph., Bot. Jahrb. Syst. 20 (3): 317, 1895 (Stephani 1895a). Bas.: Phragmicoma aulacophora Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 259, 1843 (Montagne 1843). *** Acrolejeunea mollis (Hook.f. et Taylor) Schifn., Hedwigia 33 (4): 178, 1894 (Schifner 1894). Bas.: Ptychanthus mollis Hook.f. et Taylor, London J. Bot. 5: 384, 1846 (Taylor 1846b). *** Acrolejeunea securifolia (Nees) Steph., Hedwigia 34 (2): 59, 1895 (Stephani 1895c). Bas.: Jungermannia securifolia Nees, Prodr. Fl. Norfolk.: 5, 1833 (Endlicher 1833). *** Acrolejeunea securifolia subsp. caledonica (Steph.) Gradst., Bryophyt. Biblioth. 4: 100, 1975 (Gradstein 1975). Bas.: Ptychocoleus caledonicus Steph., Sp. Hepat. (Stephani) 5: 39, 1912 (Stephani 1912c). *** Acrolejeunea securifolia subsp. hartmannii (Steph.) Gradst., Bryophyt. Biblioth. 4: 99, 1975 (Gradstein 1975). Bas.: Ptychocoleus hartmannii Steph., Sp. Hepat. (Stephani) 5: 44, 1912 (Stephani 1912c). *** Acrolejeunea securifolia subsp. pallida (Ångstr.) Gradst., Bryophyt. Biblioth. 4: 101, 1975 (Gradstein 1975). Bas.: Phragmicoma pallida Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 30 (5): 132, 1873 (Ångström 1873). ** sect. Trocholejeunea (Schifn.) Jian Wang bis et Gradst., Bryophyte Diversity Evol. 36 (1): 38, 2014 (Wang et al. 2014b). Bas.: Trocholejeunea Schifn., Ann. Bryol. 5: 160, 1932 (Dixon et al. 1932). World checklist of hornworts and liverworts 401 *** Acrolejeunea crassicaulis (Steph.) Jian Wang bis et Gradst., Bryophyte Diversity Evol. 36 (1): 38, 2014 (Wang et al. 2014b). Bas.: Hygrolejeunea crassicaulis Steph., Sp. Hepat. (Stephani) 5: 550, 1914 (Stephani 1914b). *** Acrolejeunea infuscata (Mitt.) Jian Wang bis et Gradst., Bryophyte Diversity Evol. 36 (1): 38, 2014 (Wang et al. 2014b). Bas.: Lejeunea infuscata Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 111, 1860 [1861] (Mitten 1860c). ** Acrolejeunea meghalayensis (Ajit P.Singh et V.Nath) Jian Wang bis et Gradst., Bryophyte Diversity Evol. 36 (1): 39, 2014 (Wang et al. 2014b). Bas.: Trocholejeunea meghalayensis Ajit P.Singh et V.Nath, J. Jap. Bot. 83 (1): 2, 2008 (Singh and Nath 2008). ** Acrolejeunea sinensis (Jian Wang bis, R.L.Zhu et Gradst.) Jian Wang bis et Gradst., Bryophyte Diversity Evol. 36 (1): 39, 2014 (Wang et al. 2014b). Bas.: Trocholejeunea sinensis Jian Wang bis, R.L.Zhu et Gradst., Phytotaxa 174 (5): 296, 2014 (Wang et al. 2014c). * * * Excluded from the genus Acrolejeunea abnormis (Gottsche) Pearson, Forh. Vidensk.-Selsk. Kristiania 1892 (8): 4, 1892 (Pearson 1892). Bas.: Phragmicoma abnormis Gottsche, Abh. Naturwiss. Vereins Bremen 7: 352, 1882 (Gottsche 1882). 354 Acrolejeunea comptonii Pearson, J. Linn. Soc., Bot. 46 (305): 33, 1922 (Pearson 1922b). 355 Acrolejeunea inlexa (Gottsche) Pearson, Forh. Vidensk.-Selsk. Kristiania 1892 (8): 4, 1892 (Pearson 1892). Bas.: Phragmicoma inlexa Gottsche, Abh. Naturwiss. Vereins Bremen 7: 351, 1882 (Gottsche 1882). 356 *** Archilejeunea (Spruce) Steph., Hedwigia 27 (3/4): 113, 1888 (Stephani 1888d). Bas.: Lejeunea subg. Archilejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 88, 1884 (Spruce 1884). 357 *** Archilejeunea abbreviata (Mitt.) Vanden Berghen, Rev. Bryol. Lichénol. 20 (1/2): 117, 1951 (Vanden Berghen 1951c). Bas.: Lejeunea abbreviata Mitt., J. Proc. Linn. Soc., Bot. 7 (27): 167, 1863 (Mitten 1863). ** Archilejeunea africana Steph., Sp. Hepat. (Stephani) 4: 705, 1911 (Stephani 1911e). ** Archilejeunea alata Steph., Bull. Soc. Roy. Bot. Belgique, Compt. Rend. 32 (2): 33, 1893 [1894] (Stephani 1893e). 354 Acrolejeunea abnormis was excluded from Acrolejeunea by Gradstein (1975), but the type specimen was not localized. 355 Acrolejeunea comptonii is a Mastigolejeunea species (Gradstein 1975). he type specimen has not been localized. 356 Acrolejeunea inlexa was excluded from Acrolejeunea by Gradstein (1975), but the type specimen was not localized. 357 Archilejeunea is polyphyletic and several nomenclatural and taxonomic will be proposed (Shi et al. 2015a). 402 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Archilejeunea amakawana Inoue, J. Jap. Bot. 41 (1): 16, 1966 (Inoue 1966d). Nom. nov. pro Archilejeunea falcata Amakawa, J. Jap. Bot. 39 (5): 137, 1964 (Amakawa 1964b), nom. illeg. *** Archilejeunea auberiana (Mont.) Steph., Hedwigia 29 (3): 134, 1890 (Stephani 1890d). Bas.: Lejeunea auberiana Mont., Hist. Phys. Cuba, Bot., Pl. Cell.: 483, 1842 (Montagne 1842a). *** Archilejeunea autoica Vanden Berghen, Rev. Bryol. Lichénol. 20 (1/2): 119, 1951 (Vanden Bergen 1951b). *** Archilejeunea badia (Spruce) Steph., Sp. Hepat. (Stephani) 4: 711, 1911 (Stephani 1911e). Bas.: Lejeunea badia Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 92, 1884 (Spruce 1884). * Archilejeunea bilabiata (Mitt.) Steph., Sp. Hepat. (Stephani) 4: 723, 1911 (Stephani 1911e). Bas.: Phragmicoma bilabiata Mitt., Fl. vit.: 412, 1871 [1873] (Mitten 1871). 358 *** Archilejeunea bischleriana Gradst., Fl. Neotrop. Monogr. 62: 62, 1994 (Gradstein 1994). ** Archilejeunea bongardii Steph., Hedwigia 29 (1): 20, 1890 (Stephani 1890a). ** Archilejeunea brachyantha J.B.Jack et Steph., Bot. Centralbl. 60 (4): 104, 1894 (Jack and Stephani 1894). ** Archilejeunea brevilobula Steph., Sp. Hepat. (Stephani) 4: 706, 1911 (Stephani 1911e). *** Archilejeunea crispistipula (Spruce) Steph., Sp. Hepat. (Stephani) 4: 712, 1911 (Stephani 1911e). Bas.: Lejeunea crispistipula Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 93, 1884 (Spruce 1884). * Archilejeunea eberhardtii Steph., Sp. Hepat. (Stephani) 4: 725, 1911 (Stephani 1911e). ** Archilejeunea elobulata Steph., Sp. Hepat. (Stephani) 4: 707, 1911 (Stephani 1911e). ** Archilejeunea gradsteinii X.Q.Shi et R.L.Zhu, Nova Hedwigia 100 (3/4): 592, 2015 (Shi and Zhu 2015). ** Archilejeunea incrassata Steph., Rev. Bryol. 35 (2): 30, 1908 (Stephani 1908l). ** Archilejeunea jonesii Vanden Berghen, Rev. Bryol. Lichénol. 20 (1/2): 116, 1951 (Vanden Berghen 1951c). *** Archilejeunea juliformis (Nees) Gradst., Bryophyt. Biblioth. 4: 127, 1975 (Gradstein 1975). Bas.: Jungermannia juliformis Nees, Fl. Bras. (Martius) 1 (1): 351, 1833 (Nees 1833a). *** Archilejeunea kiushiana (Horik.) Verd., Ann. Bryol., Suppl. 4: 46, 1934 (Verdoorn 1934a). Bas.: Lopholejeunea kiushiana Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 1: 129, 1932 (Horikawa 1932c). *** Archilejeunea ludoviciana (De Not.) P.Geissler et Gradst., J. Hattori Bot. Lab. 75: 202, 1994 (Geissler and Gradstein 1994). Bas.: Phragmicoma ludoviciana De Not., Nov. Stirp. Pug. 10: 11, 1857 (Lehmann 1857). *** Archilejeunea ludoviciana subsp. porelloides (Spruce) Gradst., Fl. Neotrop. Monogr. 62: 58, 1994 (Gradstein 1994). Bas.: Lejeunea porelloides Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 90, 1884 (Spruce 1884). 358 Archilejeunea bilabiata is a doubtful taxon, the type specimen could not be found (Geissler and Gradstein 1994). World checklist of hornworts and liverworts 403 ** Archilejeunea nebeliana Gradst. et Schäf.-Verw., Cryptog. Bryol. 33 (2): 108, 2012 (Gradstein and Schäfer-Verwimp 2012). *** Archilejeunea olivacea (Hook.f. et Taylor) Steph., Hedwigia 29 (3): 134, 1890 (Stephani 1890d). Bas.: Jungermannia olivacea Hook.f. et Taylor, London J. Bot. 3: 568, 1844 (Hooker and Taylor 1844d). *** Archilejeunea parvilora (Nees) Steph., Hedwigia 29 (3): 134, 1890 (Stephani 1890d). Bas.: Jungermannia parvilora Nees, Fl. Bras. (Martius) 1 (1): 353, 1833 (Nees 1833a). ** Archilejeunea planifolia (Horik.) Mizut., J. Hattori Bot. Lab. 73: 175, 1993 (Mizutani 1993). Bas.: Leucolejeunea planifolia Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 1: 199, 1933 (Horikawa 1933). 359 *** Archilejeunea planiuscula (Mitt.) Steph., Sp. Hepat. (Stephani) 4: 731, 1911 (Stephani 1911e). Bas.: Lejeunea planiuscula Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 111, 1860 [1861] (Mitten 1860c). * * Excluded from the genus Archilejeunea negrensis Steph., Sp. Hepat. (Stephani) 4: 716, 1911 (Stephani 1911e). 360 Archilejeunea ovata Herzog, Rev. Bryol. Lichénol. 20 (1/2): 130, 1951 [1952] (Herzog 1951a). 361 *** Bryopteris (Nees) Lindenb., Syn. Hepat. 2: 284, 1845 (Gottsche et al. 1845a). Bas.: Frullania subg. Bryopteris Nees, Naturgesch. Eur. Leberm. 3: 211, 1838 (Nees 1838b). *** Bryopteris difusa (Sw.) Nees, Syn. Hepat. 2: 286, 1845 (Gottsche et al. 1845a). Bas.: Jungermannia difusa Sw., Prodr. (Swartz): 144, 1788 (Swartz 1788). *** Bryopteris ilicina (Sw.) Nees, Syn. Hepat. 2: 284, 1845 (Gottsche et al. 1845a). Bas.: Jungermannia ilicina Sw., Prodr. (Swartz): 145, 1788 (Swartz 1788). * Bryopteris issiloba Steph., Sp. Hepat. (Stephani) 6: 568, 1924 (Stephani 1924). 362 *** Bryopteris gaudichaudii Gottsche, Ann. Sci. Nat. Bot. (sér. 4) 8: 340, 1857 (Gottsche 1857). *** Caudalejeunea Schifn., Hepat. (Engl.-Prantl): 129, 1893 (Schifner 1893b). ** subg. Acaudalejeunea R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 779, 1980 (Schuster 1980c). *** Caudalejeunea grolleana Gradst., Acta Bot. Neerl. 23 (3): 334, 1974 (Gradstein 1974b). 359 Archilejeunea planifolia is conspeciic with Archilejeunea kiushiana in Hattori (1952c) and Gradstein and Geissler (1997), but it is a separate species in Mizutani (1993). 360 Archilejeunea negrensis is probably not an Archilejeunea species, but the type specimen is too poor to permit identiication (Gradstein and Buskes 1985). 361 Archilejeunea ovata is conspeciic with Lopholejeunea nigricans or Lopholejeunea subfusca (Gradstein 1994). 362 Bryopteris issiloba is probably conspeciic with Bryopteris difusa (Gradstein and Costa 2003). It was not accepted by Hartmann et al. (2006). 404 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** subg. Caudalejeunea ** Caudalejeunea africana (Steph.) Schifn., Hepat. (Engl.-Prantl): 129, 1893 (Schifner 1893b). Bas.: hysananthus africanus Steph., Bot. Jahrb. Syst. 8 (2): 93, 1886 (Stephani 1886d). *** Caudalejeunea hanningtonii (Mitt.) Schifn., Hepat. (Engl.-Prantl): 129, 1893 (Schifner 1893b). Bas.: Lejeunea hanningtonii Mitt., J. Linn. Soc., Bot. 22 (146): 324, 1886 (Mitten 1886b). ** Caudalejeunea katangensis Vanden Berghen, Explor. Hydrobiol. Lac Bangweolo Luapula: 94, 1972 (Vanden Berghen 1972b). *** Caudalejeunea lehmanniana (Gottsche) A.Evans, Bull. Torrey Bot. Club 34 (11): 554, 1907 [1908] (Evans 1907a). Bas.: Lejeunea lehmanniana Gottsche, Syn. Hepat. 3: 325, 1845 (Gottsche et al. 1845b). ** Caudalejeunea lewallei Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 42 (4): 434, 1972 (Vanden Berghen 1972a). ** subg. Vermilejeunea R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 778, 1980 (Schuster 1980c). *** Caudalejeunea cristiloba (Steph.) Gradst., Acta Bot. Neerl. 23 (3): 340, 1974 (Gradstein 1974b). Bas.: Acrolejeunea cristiloba Steph., Hedwigia 34 (2): 56, 1895 (Stephani 1895c). *** Caudalejeunea cristiloba subsp. samoana (Steph.) Gradst., Acta Bot. Neerl. 23 (3): 343, 1974 (Gradstein 1974b). Bas.: Caudalejeunea samoana Steph., Akad. Wiss. Wien, Math.-Naturwiss. Kl., Denkschr. 81: 296, 1907 (Stephani 1907a). ** Caudalejeunea dusenii Steph., Sp. Hepat. (Stephani) 5: 11, 1912 (Stephani 1912c). *** Caudalejeunea reniloba (Gottsche) Steph., Sp. Hepat. (Stephani) 5: 16, 1912 (Stephani 1912c). Bas.: Phragmicoma reniloba Gottsche, Syn. Hepat. 2: 301, 1845 (Gottsche et al. 1845a). ** Caudalejeunea yangambiensis (Vanden Berghen) E.W.Jones, Trans. Brit. Bryol. Soc. 3 (2): 192, 1957 (Jones 1957b). Bas.: Ptychocoleus yangambiensis Vanden Berghen, Bull. Soc. Roy. Bot. Belgique 84: 61, 1951 (Vanden Berghen 1951a). Incertae sedis ** Caudalejeunea acutifolia Gerola, Lav. Bot. Ist. Bot. Univ. Padova 12: 478, 1947 (Gerola 1947). * Caudalejeunea mauritiana Horik., Acta Phytotax. Geobot. 13: 214, 1943 (Horikawa 1943). Nom. nov. pro Dicranolejeunea africana Steph., Sp. Hepat. (Stephani) 5: 158, 1912 (Stephani 1912c). ** Caudalejeunea streimannii Gyarmati, Trop. Bryol. 22: 129, 2002 (Sass-Gyarmati 2002). ** Caudalejeunea tridentata R.L.Zhu, Y.M.Wei et Qiong He, Bryologist 114 (3): 469, 2011 (Zhu et al. 2011). World checklist of hornworts and liverworts 405 ** Cephalantholejeunea R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 798, 1980 (Schuster 1980c). *** Cephalantholejeunea temnanthoides (R.M.Schust.) R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 807, 1980 (Schuster 1980c). Bas.: Potamolejeunea temnanthoides R.M.Schust., Beih. Nova Hedwigia 9: 123, 1963 (Schuster 1963a). ** Cephalolejeunea Mizut., J. Hattori Bot. Lab. 46: 359, 1979 (Mizutani 1979b). *** Cephalolejeunea parvilobula Mizut., J. Hattori Bot. Lab. 46: 359, 1979 (Mizutani 1979b). *** Frullanoides Raddi, Critt. Brasil.: 13, 1822 (Raddi 1822). *** Frullanoides bahamensis (A.Evans) van Slageren, Meded. Bot. Mus. Herb. Rijks Univ. Utrecht 544: 81, 1985 (van Slageren 1985). Bas.: Brachiolejeunea bahamensis A.Evans, Bull. Torrey Bot. Club 35 (8): 383, 1908 (Evans 1908b). *** Frullanoides corticalis (Lehm. et Lindenb.) van Slageren, Meded. Bot. Mus. Herb. Rijks Univ. Utrecht 544: 84, 1985 (van Slageren 1985). Bas.: Jungermannia corticalis Lehm. et Lindenb., Nov. Stirp. Pug. 4: 50, 1832 (Lehmann 1832). *** Frullanoides densifolia Raddi, Critt. Brasil.: 14, 1822 (Raddi 1822). *** Frullanoides densifolia subsp. grandidentata (L.Clark) van Slageren, Meded. Bot. Mus. Herb. Rijks Univ. Utrecht 544: 95, 1985 (van Slageren 1985). Bas.: Brachiolejeunea grandidentata L.Clark, Proc. Calif. Acad. Sci. (ser. 4) 27 (18): 595, 1953 (Clark 1953). *** Frullanoides laciniatilora (Loitl.) van Slageren, Meded. Bot. Mus. Herb. Rijks Univ. Utrecht 544: 100, 1985 (van Slageren 1985). Bas.: Lejeunea laciniatilora Loitl., Diagn. pl. nov.: 19, 1894 (Loitlesberger 1894). *** Frullanoides liebmaniana (Lindenb. et Gottsche) van Slageren, Meded. Bot. Mus. Herb. Rijks Univ. Utrecht 544: 102, 1985 (van Slageren 1985). Bas.: Phragmicoma liebmaniana Lindenb. et Gottsche, Syn. Hepat. 5: 744, 1847 (Gottsche et al. 1847). *** Frullanoides mexicana van Slageren, Meded. Bot. Mus. Herb. Rijks Univ. Utrecht 544: 106, 1985 (van Slageren 1985). *** Frullanoides tristis (Steph.) van Slageren, Meded. Bot. Mus. Herb. Rijks Univ. Utrecht 544: 110, 1985 (van Slageren 1985). Bas.: Brachiolejeunea tristis Steph., Sp. Hepat. (Stephani) 5: 112, 1912 (Stephani 1912c). *** Fulfordianthus Gradst., Bryologist 95 (1): 44, 1992 (Gradstein 1992a). *** Fulfordianthus evansii (Fulford) Gradst., Bryologist 95 (1): 46, 1992 (Gradstein 1992a). Bas.: hysananthus evansii Fulford, Bull. Torrey Bot. Club 68 (1): 34, 1941 (Fulford 1941). *** Fulfordianthus pterobryoides (Spruce) Gradst., Bryologist 95 (1): 44, 1992 (Gradstein 1992a). Bas.: Lejeunea pterobryoides Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 109, 1884 (Spruce 1884). 406 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Lopholejeunea (Spruce) Steph., Bot. Gaz. 15 (11): 285, 1890 (Stephani 1890c) nom. rejic. Bas.: Lejeunea subg. Lopholejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 119, 1884 (Spruce 1884). ** subg. Lopholejeunea ** Lopholejeunea borbonica Steph., Hedwigia 35 (3): 109, 1896 (Stephani 1896b). ** Lopholejeunea jonesii Vanden Berghen, Bull. Jard. Bot. État Bruxelles 20 (2): 178, 1950 (Vanden Berghen 1950). ** Lopholejeunea laciniata E.W.Jones, Trans. Brit. Bryol. Soc. 3 (2): 194, 1957 (Jones 1957b). ** Lopholejeunea minima Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 54 (3/4): 437, 1984 (Vanden Berghen 1984a). * Lopholejeunea obtusilacera Herzog, Bull. Jard. Bot. État Bruxelles 20 (2): 172, 1950 (Vanden Berghen 1950). * Lopholejeunea paramultilacera Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 54 (3/4): 435, 1984 (Vanden Berghen 1984a). * Lopholejeunea quinquecarinata Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 54 (3/4): 408, 1984 (Vanden Berghen 1984a). ** Lopholejeunea renistipula (Mitt.) Steph., Sp. Hepat. (Stephani) 5: 94, 1912 (Stephani 1912c). Bas.: Phragmicoma renistipula Mitt., Fl. vit.: 413, 1871 [1873] (Mitten 1871). ** Lopholejeunea revoluta E.W.Jones, Trans. Brit. Bryol. Soc. 3 (2): 207, 1957 (Jones 1957b). ** sect. Eulophae Verd., Ann. Bryol., Suppl. 4: 87, 1934 (Verdoorn 1934a). *** Lopholejeunea applanata (Reinw., Blume et Nees) Schifn., Hepat. (Engl.-Prantl): 129, 1893 (Schifner 1893b). Bas.: Jungermannia applanata Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 210, 1824 [1825] (Reinwardt et al. 1824a). *** Lopholejeunea borneensis (Steph.) Verd., Ann. Bryol., Suppl. 4: 83, 1934 (Verdoorn 1934a). Bas.: Mastigolejeunea borneensis Steph., Sp. Hepat. (Stephani) 4: 777, 1912 (Stephani 1912b). *** Lopholejeunea erugata B.M.hiers, Brittonia 36 (2): 174, 1984 (hiers 1984). Nom. nov. pro Ptychocoleus inermis Steph., Sp. Hepat. (Stephani) 5: 27, 1912 (Stephani 1912c). *** Lopholejeunea eulopha (Taylor) Schifn., Hepat. (Engl.-Prantl): 129, 1893 (Schifner 1893b). Bas.: Lejeunea eulopha Taylor, London J. Bot. 5: 391, 1846 (Taylor 1846b). *** Lopholejeunea evansiana Verd., Nova Guinea 18: 4, 1934 (Verdoorn 1934d). *** Lopholejeunea grollei R.L.Zhu et Gradst., Monogr. Syst. Bot. Missouri Bot. Gard. 74: 36, 2005 (Zhu and Gradstein 2005). *** Lopholejeunea herzogiana Verd., Recueil Trav. Bot. Neerl. 30: 217, 1933 (Verdoorn 1933b). World checklist of hornworts and liverworts 407 *** Lopholejeunea hispidissima Steph., Sp. Hepat. (Stephani) 5: 80, 1912 (Stephani 1912c). *** Lopholejeunea loheri Steph., Sp. Hepat. (Stephani) 5: 77, 1912 (Stephani 1912c). *** Lopholejeunea minuta R.L.Zhu et Gradst., Nova Hedwigia 78 (3/4): 436, 2004 (Zhu and Gradstein 2004). *** Lopholejeunea nigricans (Lindenb.) Schifn., Consp. Hepat. Arch. Ind.: 293, 1898 (Schifner 1898b). Bas.: Lejeunea nigricans Lindenb., Syn. Hepat. 3: 316, 1845 (Gottsche et al. 1845b). *** Lopholejeunea plicatiscypha (Hook.f. et Taylor) Steph., Sp. Hepat. (Stephani) 5: 96, 1912 (Stephani 1912c). Bas.: Phragmicoma plicatiscypha Hook.f. et Taylor, London J. Bot. 5: 386, 1846 (Taylor 1846b). * Lopholejeunea proxima Steph., Sp. Hepat. (Stephani) 5: 89, 1912 (Stephani 1912c). 363 *** Lopholejeunea streimannii B.M.hiers et Gradst., Mem. New York Bot. Gard. 52: 37, 1989 (hiers and Gradstein 1989). ** Lopholejeunea vojtkoana Gyarmati, Nova Hedwigia 87 (3/4): 480, 2008 (SassGyarmati 2008). ** *** *** *** *** *** *** *** ** *** * *** sect. Lopholejeunea Lopholejeunea ceylanica Steph., Sp. Hepat. (Stephani) 5: 86, 1912 (Stephani 1912c). Lopholejeunea horticola Schifn., Ann. Bryol. 6: 133, 1933 (Herzog et al. 1933). Lopholejeunea latilobula Verd., Nova Guinea 18: 4, 1934 (Verdoorn 1934d). Lopholejeunea magna Mizut., J. Hattori Bot. Lab. 32: 131, 1969 (Mizutani 1969). Lopholejeunea recurvata Mizut., J. Hattori Bot. Lab. 46: 369, 1979 (Mizutani 1979b). Lopholejeunea soae R.L.Zhu et Gradst., Monogr. Syst. Bot. Missouri Bot. Gard. 74: 69, 2005 (Zhu and Gradstein 2005). Lopholejeunea subfusca (Nees) Schifn., Bot. Jahrb. Syst. 23 (5): 593, 1897 (Schifner 1897). Bas.: Jungermannia subfusca Nees, Enum. Pl. Crypt. Javae: 36, 1830 (Nees 1830). Lopholejeunea subfusca var. elongata Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 54 (3/4): 445, 1984 (Vanden Berghen 1984a). Lopholejeunea wiltensii Steph., Hedwigia 35 (3): 112, 1896 (Stephani 1896b). Lopholejeunea yapensis Steph., Sp. Hepat. (Stephani) 5: 81, 1912 (Stephani 1912c). 364 Lopholejeunea zollingeri (Steph.) Schifn., Consp. Hepat. Arch. Ind.: 296, 1898 (Schifner 1898b). Bas.: Lejeunea zollingeri Steph., Hedwigia 29 (1): 14, 1890 (Stephani 1890a). ** subg. Pholianthus B.M.hiers et Gradst., Mem. New York Bot. Gard. 52: 25, 1989 (hiers and Gradstein 1989). *** Lopholejeunea colensoi Steph., Sp. Hepat. (Stephani) 5: 97, 1912 (Stephani 1912c). *** Lopholejeunea pocsii Gyarmati, Cryptog. Bryol. 26 (4): 404, 2005 (Sass-Gyarmati 2005). 363 Lopholejeunea proxima is possibly conspeciic with Lopholejeunea nigricans (Zhu and Gradstein 2005). 364 Lopholejeunea yapensis is possibly conspeciic with Lopholejeunea subfusca (Mizutani 1985). 408 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** subg. Pteryganthus B.M.hiers, Brittonia 35 (1): 85, 1983 (hiers 1983). ** Lopholejeunea grandicrista Steph., Bull. Soc. Roy. Bot. Belgique, Compt. Rend. 32 (2): 34, 1893 [1894] (Stephani 1893e). *** Lopholejeunea leioptera Gyarmati, Candollea 56 (1): 80, 2001 (Sass-Gyarmati 2001). ** Lopholejeunea onraedtii Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 54 (3/4): 452, 1984 (Vanden Berghen 1984a). ** Lopholejeunea sphaerophora (Lehm. et Lindenb.) Steph., Sp. Hepat. (Stephani) 5: 68, 1912 (Stephani 1912c). Bas.: Jungermannia sphaerophora Lehm. et Lindenb., Nov. Stirp. Pug. 5: 9, 1833 (Lehmann 1833). * Lopholejeunea tixieriana Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 54 (3/4): 454, 1984 (Vanden Berghen 1984a). ** Lopholejeunea utriculata Steph., Sp. Hepat. (Stephani) 5: 69, 1912 (Stephani 1912c). Incertae sedis ** Lopholejeunea lepidoscypha Kiaer et Pearson, Forh. Vidensk.-Selsk. Kristiania 1892 (8): 5, 1892 (Pearson 1892). * Lopholejeunea multilacera Steph., Bot. Gaz. 15 (11): 285, 1890 (Stephani 1890c). ** Lopholejeunea udarii M.Dey et D.K.Singh, Nelumbo 53: 197, 2011 (Dey and Singh 2011). * * Excluded from the genus Lopholejeunea aberrantia Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 256, 1934 (Horikawa 1934). 365 Lopholejeunea vietnamica Tixier, Ann. Hist.-Nat. Mus. Natl. Hung. 66: 99, 1974 (Tixier 1974). 366 *** Marchesinia Gray, Nat. Arr. Brit. Pl. 1: 689, 1821 (Gray 1821) nom. conserv. *** subg. Marchesinia *** Marchesinia mackaii (Hook.) Gray, Nat. Arr. Brit. Pl. 1: 689, 1821 (Gray 1821). Bas.: Jungermannia mackaii Hook., Brit. Jungermann.: tab. 53, 1813 (Hooker 1813). *** subg. Marchesiniopsis R.M.Schust., J. Hattori Bot. Lab. 72: 358, 1992 (Schuster 1992a). *** Marchesinia bongardiana (Lehm. et Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 405, 1877 (Trevisan 1877). Bas.: Lejeunea bongardiana Lehm. et Lindenb., Nov. Stirp. Pug. 7: 18, 1838 (Lehmann 1838). 365 Lopholejeunea aberrantia is probably an Archilejeunea species, but the type specimen in HIRO is destroyed (Zhu and Gradstein 2005). 366 Lopholejeunea vietnamica is probably an Archilejeunea species (Zhu and Gradstein 2005). he type specimen was not found in PC. World checklist of hornworts and liverworts 409 *** Marchesinia brachiata (Sw.) Schifn., Hepat. (Engl.-Prantl): 128, 1893 (Schifner 1893b). Bas.: Jungermannia brachiata Sw., Prodr. (Swartz): 144, 1788 (Swartz 1788). *** Marchesinia deslooveri Vanden Berghen, Rev. Bryol. Lichénol. 42 (4): 926, 1976 (Vanden Berghen 1976a). *** Marchesinia excavata (Mitt.) Schifn., Hepat. (Engl.-Prantl): 128, 1893 (Schifner 1893b). Bas.: Lejeunea excavata Mitt., Trans. Linn. Soc. London 23 (1): 58, 1860 (Mitten 1860a). *** Marchesinia languida (Nees et Mont.) Steph., Sp. Hepat. (Stephani) 5: 149, 1912 (Stephani 1912c). Bas.: Lejeunea languida Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 5: 59, 1836 (Nees and Montagne 1836). *** Marchesinia nobilis (Gottsche) X.Q.Shi, R.L.Zhu et Gradst., Phytotaxa 195 (3): 249, 2015 (Shi et al. 2015b). Bas.: Lejeunea nobilis Gottsche, Abh. Naturwiss. Vereins Bremen 7: 353, 1882 (Gottsche 1882). *** Marchesinia robusta (Mitt.) Schifn., Hepat. (Engl.-Prantl): 128, 1893 (Schifner 1893b). Bas.: Lejeunea robusta Mitt., Hooker’s J. Bot. Kew Gard. Misc. 3: 359, 1851 (Mitten 1851). ** Mastigolejeunea (Spruce) Steph., Hedwigia 30 (5): 206, 1891 (Stephani 1891a). Bas.: Lejeunea subg. Mastigolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 100, 1884 (Spruce 1884). *** Mastigolejeunea auriculata (Wilson et Hook.) Steph., Bot. Gaz. 17 (6): 171, 1892 (Stephani 1892f). Bas.: Jungermannia auriculata Wilson et Hook., Musci Amer., S. States: no. 170, 1841 (Wilson 1841; non vidi). ** Mastigolejeunea auriculata var. rhodesica (Vanden Berghen) Sukkharak et Gradst., Nova Hedwigia 99 (3/4): 297, 2014 (Sukkharak and Gradstein 2014). Bas.: Brachiolejeunea rhodesica Vanden Berghen, Bull. Jard. Bot. État Bruxelles 21 (1/2): 94, 1951 (Vanden Berghen 1951d). *** Mastigolejeunea calcarata (Steph.) Verd., Blumea 1 (1): 230, 1934 (Verdoorn 1934b). Bas.: Archilejeunea calcarata Steph., Sp. Hepat. (Stephani) 4: 724, 1911 (Stephani 1911e). *** Mastigolejeunea lorea (Mitt.) Paris, Rev. Bryol. 33 (3): 42, 1906 (Paris 1906b). Bas.: Phragmicoma lorea Mitt., J. Linn. Soc., Bot. 22 (146): 323, 1886 (Mitten 1886b). *** Mastigolejeunea frauenfeldii (Reichardt) Verd., Blumea 1 (1): 230, 1934 (Verdoorn 1934b). Bas.: hysananthus frauenfeldii Reichardt, Verh. K.K. Zool.-Bot. Ges. Wien 16: 958, 1866 (Reichardt 1866). ** Mastigolejeunea gradsteinii Sukkharak, J. Bryol. 36 (1): 56, 2014 (Sukkharak 2014). 410 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Mastigolejeunea humilis (Gottsche) Schifn., Hepat. (Engl.-Prantl): 129, 1893 (Schifner 1893b). Bas.: Phragmicoma humilis Gottsche, Syn. Hepat. 2: 299, 1845 (Gottsche et al. 1845a). 367 *** Mastigolejeunea indica Steph., Sp. Hepat. (Stephani) 4: 776, 1912 (Stephani 1912b). *** Mastigolejeunea innovans (Spruce) Steph., Sp. Hepat. (Stephani) 4: 765, 1912 (Stephani 1912b). Bas.: Lejeunea innovans Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 103, 1884 (Spruce 1884). *** Mastigolejeunea ligulata (Lehm. et Lindenb.) Schifn., Hepat. (Engl.-Prantl): 129, 1893 (Schifner 1893b). Bas.: Jungermannia ligulata Lehm. et Lindenb., Nov. Stirp. Pug. 6: 39, 1834 (Lehmann 1834). *** Mastigolejeunea nigra Steph., Hedwigia 30 (5): 206, 1891 (Stephani 1891a). *** Mastigolejeunea plicatilora (Spruce) Steph., Sp. Hepat. (Stephani) 4: 766, 1912 (Stephani 1912b). Bas.: Lejeunea plicatilora Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 104, 1884 (Spruce 1884). *** Mastigolejeunea recondita (Steph.) Mizut., J. Hattori Bot. Lab. 32: 134, 1969 (Mizutani 1969). Bas.: Ptycholejeunea recondita Steph., Hedwigia 35 (3): 122, 1896 (Stephani 1896b). *** Mastigolejeunea repleta (Taylor) A.Evans, Mem. Torrey Bot. Club 8 (2): 131, 1902 (Evans 1902a). Bas.: Lejeunea repleta Taylor, London J. Bot. 5: 392, 1846 (Taylor 1846b). *** Mastigolejeunea truncata Mizut., J. Hattori Bot. Lab. 61: 292, 1986 [1987] (Mizutani 1986c). ** Mastigolejeunea turgida Steph., Hedwigia 31 (4): 170, 1892 (Stephani 1892g). *** Mastigolejeunea virens (Ångstr.) Steph., Sp. Hepat. (Stephani) 4: 776, 1912 (Stephani 1912b), nom. conserv. Bas.: hysananthus virens Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 30 (5): 131, 1873 (Ångström 1873). ** Phaeolejeunea Mizut., J. Hattori Bot. Lab. 31: 130, 1968 (Mizutani 1968). *** Phaeolejeunea amicorum (Hürl.) Pócs, Fieldiana, Bot. (n.ser.) 47: 140, 2008 (Pócs 2008b). Bas.: Phaeolejeunea etesseana subsp. amicorum Hürl., Bauhinia 9 (4): 263, 1991 (Hürlimann 1991). ** Phaeolejeunea etesseana (Steph.) Mizut., J. Hattori Bot. Lab. 31: 133, 1968 (Mizutani 1968). Bas.: Brachiolejeunea etesseana Steph., Sp. Hepat. (Stephani) 5: 133, 1912 (Stephani 1912c). ** Phaeolejeunea inermis (Steph.) Mizut., J. Hattori Bot. Lab. 31: 134, 1968 (Mizutani 1968). Bas.: Lopholejeunea inermis Steph., Sp. Hepat. (Stephani) 5: 92, 1912 (Stephani 1912c). *** Phaeolejeunea latistipula (Schifn. ex P.Syd.) Mizut., J. Hattori Bot. Lab. 31: 131, 1968 (Mizutani 1968). Bas.: Hygrolejeunea latistipula Schifn. ex P.Syd., Just’s Bot. Jahresber. 19: 246, 1894 (Sydow 1894). 367 Mastigolejeunea humilis has been treated as conspeciic with Mastigolejeunea auriculata by many recent authors, but it merits recognition based on molecular and morphological evidence (Sukkharak et al. 2011). World checklist of hornworts and liverworts 411 *** Ptychanthus Nees, Naturgesch. Eur. Leberm. 3: 211, 1838 (Nees 1838b). ** Ptychanthus africanus Steph., Wiss. Ergebn. Deut. Zentr.-Afr. Exped. (1907-08), Bot. 2: 131, 1911 (Stephani 1911a). * Ptychanthus stephensonianus (Mitt.) Steph., Sp. Hepat. (Stephani) 4: 754, 1912 (Stephani 1912b). Bas.: Lejeunea stephensoniana Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 155, 1854 (Mitten 1854). 368 *** Ptychanthus striatus (Lehm. et Lindenb.) Nees, Naturgesch. Eur. Leberm. 3: 212, 1838 (Nees 1838b). Bas.: Jungermannia striata Lehm. et Lindenb., Nov. Stirp. Pug. 4: 16, 1832 (Lehmann 1832). ** Ptychanthus striatus var. intermedius (Gottsche) Verd., Ann. Bryol., Suppl. 4: 122, 1934 (Verdoorn 1934a). Bas.: Ptychanthus intermedius Gottsche, Natuurk. Tijdschr. Ned.-Indië 4: 576, 1853 (Gottsche 1853). *** Schifneriolejeunea Verd., Ann. Bryol. 6: 89, 1933 (Verdoorn 1933a). ** sect. Pappeanae R.M.Schust. ex Gradst. et Vanden Berghen, Beih. Nova Hedwigia 80: 174, 1985 (Gradstein and Vanden Berghen 1985). Based on: Phragmilejeunea R.M.Schust., J. Hattori Bot. Lab. 11: 27, 1954 (Schuster and Hattori 1954). *** Schifneriolejeunea fragilis Gradst. et E.W.Jones, J. Bryol. 12 (1): 45, 1982 (Jones 1982). *** Schifneriolejeunea madagascariensis (Steph.) Gradst., J. Hattori Bot. Lab. 38: 333, 1974 (Gradstein 1974a). Bas.: Ptychocoleus madagascariensis Steph., Sp. Hepat. (Stephani) 5: 27, 1912 (Stephani 1912c). *** Schifneriolejeunea pappeana var. bidentata Gradst. et Vanden Berghen, Beih. Nova Hedwigia 80: 182, 1985 (Gradstein and Vanden Berghen 1985). *** Schifneriolejeunea pappeana var. integra Gradst. et Vanden Berghen, Beih. Nova Hedwigia 80: 182, 1985 (Gradstein and Vanden Berghen 1985). *** Schifneriolejeunea parviloba (Steph.) Gradst., J. Hattori Bot. Lab. 38: 335, 1974 (Gradstein 1974a). Bas.: Acrolejeunea parviloba Steph., Bot. Gaz. 15 (11): 286, 1890 (Stephani 1890c). *** Schifneriolejeunea pappeana (Nees) Gradst., J. Hattori Bot. Lab. 38: 335, 1974 (Gradstein 1974a). Bas.: Phragmicoma pappeana Nees, Syn. Hepat. 2: 296, 1845 (Gottsche et al. 1845a). ** sect. Schifneriolejeunea, Occas. Pap. Farlow Herb. Cryptog. Bot. 16: 72, 1981 (Gradstein and Terken 1981). *** Schifneriolejeunea altimontana Vanden Berghen, Rev. Bryol. Lichénol. 42 (4): 923, 1976 (Vanden Berghen 1976a). *** Schifneriolejeunea amazonica Gradst., Beih. Nova Hedwigia 80: 25, 1985 (Gradstein 1985a). 368 Ptychanthus stephensonianus was tentatively treated as a separate species by hiers and Gradstein (1989). Further work on the infraspeciic variation of Ptychanthus striatus is needed to determine the correct status. 412 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Schifneriolejeunea cumingiana (Mont.) Gradst., J. Hattori Bot. Lab. 38: 333, 1974 (Gradstein 1974a). Bas.: Phragmicoma cumingiana Mont., London J. Bot. 4: 7, 1845 (Montagne 1845a). *** Schifneriolejeunea ferruginea (Steph.) Gradst., J. Hattori Bot. Lab. 38: 333, 1974 (Gradstein 1974a). Bas.: Acrolejeunea ferruginea Steph., Hedwigia 34 (2): 57, 1895 (Stephani 1895c). *** Schifneriolejeunea occulta (Steph.) Gradst., J. Hattori Bot. Lab. 38: 333, 1974 (Gradstein 1974a). Bas.: Ptychocoleus occultus Steph., Sp. Hepat. (Stephani) 5: 25, 1912 (Stephani 1912c). *** Schifneriolejeunea omphalanthoides Verd., Ann. Bryol. 6: 91, 1933 (Verdoorn 1933c). *** Schifneriolejeunea tumida (Nees) Gradst., J. Hattori Bot. Lab. 38: 335, 1974 (Gradstein 1974a). Bas.: Ptychanthus tumidus Nees, Naturgesch. Eur. Leberm. 3: 213, 1838 (Nees 1838b). *** Schifneriolejeunea tumida var. haskarliana (Gottsche) Gradst. et Terken, Occas. Pap. Farlow Herb. Cryptog. Bot. 16: 77, 1981 (Gradstein and Terken 1981). Bas.: Phragmicoma haskarliana Gottsche, Syn. Hepat. 2: 299, 1845 (Gottsche et al. 1845a). *** Schifneriolejeunea nymannii (Steph.) Gradst. et Terken, Occas. Pap. Farlow Herb. Cryptog. Bot. 16: 79, 1981 (Gradstein and Terken 1981). Bas.: Archilejeunea nymannii Steph., Sp. Hepat. (Stephani) 4: 730, 1911 (Stephani 1911e). *** Schifneriolejeunea polycarpa (Nees) Gradst., J. Hattori Bot. Lab. 38: 335, 1974 (Gradstein 1974a). Bas.: Jungermannia polycarpa Nees, Fl. Bras. (Martius) 1 (1): 350, 1833 (Nees 1833a). *** Schifneriolejeunea pulopenangensis (Gottsche) Gradst., J. Hattori Bot. Lab. 38: 335, 1974 (Gradstein 1974a). Bas.: Phragmicoma pulopenangensis Gottsche, Syn. Hepat. 2: 299, 1845 (Gottsche et al. 1845a). ** Spruceanthus Verd., Ann. Bryol., Suppl. 4: 151, 1934 (Verdoorn 1934a). *** Spruceanthus macrostipulus (Steph.) Gradst., Trop. Bryol. 4: 13, 1991 (Gradstein 1991). Bas.: Mastigolejeunea macrostipula Steph., Sp. Hepat. (Stephani) 4: 767, 1912 (Stephani 1912b). ** Spruceanthus mamillilobulus (Herzog) Verd., Hepat. Select. Crit. 9: no. 447, 1936 (Verdoorn 1936; non vidi). Bas.: Ptychanthus mamillilobulus Herzog, Symb. Sin. 5: 44, 1930 (Nicholson et al. 1930). *** Spruceanthus pluriplicatus (Steph.) Gradst., Schriftenreihe Mensch, Kultur Umwelt z. Bergl. W Neug. 7: 14, 1981 (Hiepko and Schultze-Motel 1981). Bas.: Brachiolejeunea pluriplicata Steph., Sp. Hepat. (Stephani) 5: 135, 1912 (Stephani 1912c). *** Spruceanthus polymorphus (Sande Lac.) Verd., Ann. Bryol., Suppl. 4: 155, 1934 (Verdoorn 1934a). Bas.: Phragmicoma polymorpha Sande Lac., Ned. Kruidk. Arch. 3: 420, 1854 [1855] (Sande Lacoste 1854). *** Spruceanthus semirepandus (Nees) Verd., Ann. Bryol., Suppl. 4: 153, 1934 (Verdoorn 1934a). Bas.: Jungermannia semirepanda Nees, Enum. Pl. Crypt. Javae: 39, 1830 (Nees 1830). World checklist of hornworts and liverworts 413 *** Spruceanthus sulcatus (Nees) Gradst., Beih. Nova Hedwigia 80: 26, 1985 (Gradstein 1985a). Bas.: Jungermannia sulcata Nees, Enum. Pl. Crypt. Javae: 36, 1830 (Nees 1830). *** Spruceanthus theobromae (Spruce) Gradst., Beih. Nova Hedwigia 80: 26, 1985 (Gradstein 1985a). Bas.: Lejeunea theobromae Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 99, 1884 (Spruce 1884). *** Spruceanthus thozetianus (Gottsche et F.Muell.) B.M.hiers et Gradst., Mem. New York Bot. Gard. 52: 62, 1989 (hiers and Gradstein 1989). Bas.: Phragmicoma thozetiana Gottsche et F.Muell., Fragm. (Mueller): 63, 1880 (Gottsche 1880). *** hysananthus Lindenb., Nov. Stirp. Pug. 8: 24, 1844 (Lehmann 1844). ** sect. hysananthus ** subsect. Anguiformes Sukkharak, Phytotaxa 193 (1): 37, 2015 (Sukkharak 2015). *** hysananthus anguiformis (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 289, 1845 (Gottsche et al. 1845a). Bas.: Jungermannia anguiformis Hook.f. et Taylor, London J. Bot. 3: 567, 1844 (Hooker and Taylor 1844d). *** hysananthus pancheri (Steph.) Hürl., Bauhinia 9 (2): 167, 1989 (Hürlimann 1989). Bas.: Mastigolejeunea pancheri Steph., Sp. Hepat. (Stephani) 4: 771, 1912 (Stephani 1912b). ** subsect. hysananthus *** hysananthus aculeatus Herzog, Ann. Bryol. 4: 89, 1931 (Herzog 1931b). *** hysananthus amazonicus (Spruce) Schifn., Hepat. (Engl.-Prantl): 130, 1893 (Schifner 1893b). Bas.: Lejeunea amazonica Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 106, 1884 (Spruce 1884). *** hysananthus appendiculatus Steph., Sp. Hepat. (Stephani) 4: 794, 1912 (Stephani 1912b). ** hysananthus ciliaris (Sande Lac.) Sukkharak, Nova Hedwigia 99 (3/4): 339, 2014 (Sukkharak and Gradstein 2014). Bas.: Phragmicoma ciliaris Sande Lac., Ann. Mus. Bot. Lugduno-Batavi 1: 307, 1864 (Sande Lacoste 1864). *** hysananthus comosus Lindenb., Nov. Stirp. Pug. 8: 25, 1844 (Lehmann 1844). *** hysananthus convolutus Lindenb., Syn. Hepat. 2: 288, 1845 (Gottsche et al. 1845a). ** hysananthus convolutus var. laceratus (Steph.) Sukkharak, Phytotaxa 193 (1): 30, 2015 (Sukkharak 2015). Bas.: hysananthus laceratus Steph., Sp. Hepat. (Stephani) 4: 796, 1912 (Stephani 1912b). *** hysananthus discretus Sukkharak et Gradst., Cryptog. Bryol. 31 (2): 113, 2010 (Sukkharak and Gradstein 2010). *** hysananthus gottschei (J.B.Jack et Steph.) Steph., Sp. Hepat. (Stephani) 4: 787, 1912 (Stephani 1912b). Bas.: hysanolejeunea gottschei J.B.Jack et Steph., Hedwigia 31 (1): 20, 1892 (Jack and Stephani 1892). ** hysananthus gottschei var. continuus Sukkharak, Phytotaxa 193 (1): 33, 2015 (Sukkharak 2015). 414 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** hysananthus spathulistipus (Reinw., Blume et Nees) Lindenb., Syn. Hepat. 2: 287, 1845 (Gottsche et al. 1845a). Bas.: Jungermannia spathulistipa Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 212, 1824 [1825] (Reinwardt et al. 1824a). ** sect. Vittatae Verd., Ann. Bryol., Suppl. 4: 182, 1934 (Verdoorn 1934a). ** subsect. Sandeanthus (B.M.hiers et Gradst.) Sukkharak, Phytotaxa 193 (1): 43, 2015 (Sukkharak 2015). Bas.: hysananthus subg. Sandeanthus B.M.hiers et Gradst., Mem. New York Bot. Gard. 52: 66, 1989 (hiers and Gradstein 1989). *** hysananthus mollis Steph., Sp. Hepat. (Stephani) 4: 798, 1912 (Stephani 1912b). *** hysananthus montanus Gradst., Xiao L.He et Piippo, Acta Bot. Fenn. 174: 77, 2002 (Gradstein et al. 2002). *** hysananthus retusus (Reinw., Blume et Nees) B.M.hiers et Gradst., Mem. New York Bot. Gard. 52: 67, 1989 (hiers and Gradstein 1989). Bas.: Jungermannia retusa Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 214, 1824 [1825] (Reinwardt et al. 1824a). ** hysananthus retusus subsp. sellingii (Herzog) Sukkharak, Phytotaxa 193 (1): 47, 2015 (Sukkharak 2015). Bas.: Mastigolejeunea sellingii Herzog, Ark. Bot. (n.ser.) 3 (3): 60, 1953 (Herzog 1953a). ** subsect. Vittatae (Verd.) Sukkharak, Phytotaxa 193 (1): 40, 2015 (Sukkharak 2015). Bas.: hysananthus sect. Vittatae Verd., Ann. Bryol., Suppl. 4: 182, 1934 (Verdoorn 1934a). *** hysananthus fruticosus (Lindenb. et Gottsche) Schifn., Hepat. (Engl.-Prantl): 130, 1893 (Schifner 1893b). Bas.: Bryopteris fruticosa Lindenb. et Gottsche, Syn. Hepat. 5: 737, 1847 (Gottsche et al. 1847). ** Tuzibeanthus S.Hatt., Biosphaera 1 (1): 7, 1947 (Hattori 1947a). *** Tuzibeanthus chinensis (Steph.) Mizut., J. Hattori Bot. Lab. 24: 151, 1961 (Mizutani 1961). Bas.: Ptychanthus chinensis Steph., Sp. Hepat. (Stephani) 4: 744, 1911 (Stephani 1911e). ** Verdoornianthus Gradst., Bryologist 80 (4): 607, 1977 [1978] (Gradstein 1977). *** Verdoornianthus griinii Gradst., Bryologist 80 (4): 609, 1977 [1978] (Gradstein 1977). *** Verdoornianthus marsupiifolius (Spruce) Gradst., Bryologist 80 (4): 609, 1977 [1978] (Gradstein 1977). Bas.: Lejeunea marsupiifolia Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 118, 1884 (Spruce 1884). World checklist of hornworts and liverworts 415 Porellineae R.M.Schust. *** Goebeliellaceae Verd. by M. von Konrat and M.A.M. Renner *** Goebeliella Steph., Hedwigia 51 (1): 61, 1911 (Stephani 1911c). *** Goebeliella cornigera (Mitt.) Steph., Hedwigia 51 (1): 62, 1911 (Stephani 1911c). Bas.: Frullania cornigera Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 163, 1855 (Mitten 1855). *** Lepidolaenaceae Nakai by M. von Konrat he generic composition of Lepidolaenaceae follows Crandall-Stotler et al. (2009), except that the genus Jubulopsis was reduced to a synonym of Lepidolaena by von Konrat et al. (2012a). *** Gackstroemia Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 397, 1877 (Trevisan 1877). ** subg. Gackstroemia ** Gackstroemia ljungneri (Herzog) Grolle, J. Hattori Bot. Lab. 30: 17, 1967 (Grolle 1967a). Bas.: Lepidolaena ljungneri Herzog, Ark. Bot. 29A (21): 6, 1940 (Herzog 1940). *** Gackstroemia magellanica (Lam.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 397, 1877 (Trevisan 1877). Bas.: Jungermannia magellanica Lam., Encycl. (Lamarck) 3 (1): 284, 1789 (Lamarck 1789). *** Gackstroemia novae-zelandiae R.M.Schust. et J.J.Engel, Phytotaxa 118 (1): 10, 2013 (Engel 2013a). *** Gackstroemia weindorferi (Herzog) Grolle, J. Hattori Bot. Lab. 30: 20, 1967 (Grolle 1967a). Bas.: Lepidolaena weindorferi Herzog, Ann. Bryol. 6: 103, 1933 (Verdoorn 1933a). ** subg. Hariotiella (Besch. et C.Massal. ex Schifn.) Grolle, J. Hattori Bot. Lab. 30: 12, 1967 (Grolle 1967a). Bas.: Lepidolaena subg. Hariotiella Besch. et C.Massal. ex Schifn., Hepat. (Engl.-Prantl): 110, 1893 (Schifner 1893b). *** Gackstroemia hariotiana (Besch. et C.Massal.) Grolle, J. Hattori Bot. Lab. 30: 12, 1967 (Grolle 1967a). Bas.: Polyotus hariotianus Besch. et C.Massal., Bull. Mens. Soc. Linn. Paris 1 (79): 639, 1886 (Bescherelle and Massalongo 1886). 416 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Gackstroemia patagonica (Steph.) Grolle, J. Hattori Bot. Lab. 30: 14, 1967 (Grolle 1967a). Bas.: Lepidolaena patagonica Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 76, 1911 (Stephani 1911b). ** Gackstroemia schwabei (Herzog) Grolle, J. Hattori Bot. Lab. 30: 16, 1967 (Grolle 1967a). Bas.: Lepidolaena schwabei Herzog, Rev. Bryol. Lichénol. 29 (3/4): 191, 1960 [1961] (Herzog 1960). Incertae sedis *** Gackstroemia alpina R.M.Schust., J. Hattori Bot. Lab. 36: 349, 1972 [1973] (Schuster 1972). *** Lepidogyna R.M.Schust., Phytologia 45 (5): 419, 1980 (Schuster 1980b). *** Lepidogyna hodgsoniae (Grolle) R.M.Schust., Phytologia 45 (5): 419, 1980 (Schuster 1980b). Bas.: Lepidolaena hodgsoniae Grolle, J. Hattori Bot. Lab. 30: 29, 1967 (Grolle 1967a). *** Lepidogyna menziesii (Hook.) R.M.Schust., Phytologia 45 (5): 419, 1980 (Schuster 1980b). Bas.: Jungermannia menziesii Hook., Musci Exot. 2: tab. 118, 1820 (Hooker 1820). *** Lepidolaena Dumort., Recueil Observ. Jungerm.: 13, 1835 (Dumortier 1835). 369 *** Lepidolaena berggrenii E.A.Hodgs., Trans. Roy. Soc. New Zealand 87 (3/4): 205, 1959 (Hodgson 1959). ** Lepidolaena brachyclada (Lehm.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 393, 1877 (Trevisan 1877). Bas.: Frullania brachyclada Lehm., Nov. Stirp. Pug. 8: 21, 1844 (Lehmann 1844). *** Lepidolaena clavigera (Hook.) Dumort. ex Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 393, 1877 (Trevisan 1877). Bas.: Jungermannia clavigera Hook., Musci Exot. 1: tab. 70, 1818 (Hooker 1818). *** Lepidolaena novae-zelandiae (E.A.Hodgs. et S.W.Arnell) von Konrat, L.Söderstr. et A.Hagborg, Phytotaxa 65: 51, 2012 (von Konrat et al. 2012a). Bas.: Jubula novae-zelandiae E.A.Hodgs. et S.W.Arnell, Trans. Roy. Soc. New Zealand, Bot. 3 (4): 90, 1965 (Hodgson 1965). *** Lepidolaena palpebrifolia (Hook.) Dumort. ex Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 393, 1877 (Trevisan 1877). Bas.: Jungermannia palpebrifolia Hook., Musci Exot. 1: tab. 71, 1818 (Hooker 1818). *** Lepidolaena reticulata (Hook.f. et Taylor) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 393, 1877 (Trevisan 1877). Bas.: Jungermannia reticulata Hook.f. et Taylor, London J. Bot. 3: 395, 1844 (Hooker and Taylor 1844a). 369 Lepidolaena includes Polyotus, but a few taxa have neither been transferred nor synonymized. hey are listed in the “Names in genera not currently accepted” section below. World checklist of hornworts and liverworts 417 *** Lepidolaena taylorii (Gottsche) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 393, 1877 (Trevisan 1877). Bas.: Polyotus taylorii Gottsche, Syn. Hepat. 2: 246, 1845 (Gottsche et al. 1845a). *** Porellaceae Cavers nom. conserv. *** Ascidiota C.Massal., Nuovo Giorn. Bot. Ital. (n.ser.) 5 (2): 256, 1898 (Massalongo 1898). *** Ascidiota blepharophylla C.Massal., Nuovo Giorn. Bot. Ital. (n.ser.) 5 (2): 257, 1898 (Massalongo 1898). ** Ascidiota blepharophylla subsp. alaskana Steere et R.M.Schust., Bull. Torrey Bot. Club 87 (3): 213, 1960 (Steere and Schuster 1960). *** Porella L., Sp. Pl. 1: 1106, 1753 (Linnaeus 1753). ** Porella abyssinica (Nees) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 407, 1877 (Trevisan 1877). Bas.: Madotheca abyssinica Nees, Syn. Hepat. 2: 281, 1845 (Gottsche et al. 1845a). ** Porella abyssinica var. hoehnelii (Steph.) Pócs, Fragm. Florist. Geobot. 39 (1): 229, 1994 (Pócs 1994a). Bas.: Porella hoehnelii Steph., Hedwigia 30 (6): 266, 1891 (Stephani 1891c). ** Porella acutifolia (Lehm. et Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 408, 1877 (Trevisan 1877). Bas.: Madotheca acutifolia Lehm. et Lindenb., Nov. Stirp. Pug. 7: 8, 1838 (Lehmann 1838). ** Porella acutifolia var. hattoriana (Pócs) S.Hatt., Misc. Bryol. Lichenol. 8 (4): 79, 1979 (Hattori 1979c). Bas.: Porella plumosa var. hattoriana Pócs, J. Hattori Bot. Lab. 31: 82, 1968 (Pócs 1968). * Porella acutifolia var. linguifolia (Steph.) M.L.So, Syst. Bot. 27 (1): 5, 2002 (So 2002a). Bas.: Madotheca linguifolia Steph., Sp. Hepat. (Stephani) 4: 291, 1910 (Stephani 1910b). 370 ** Porella acutifolia subsp. tosana (Steph.) S.Hatt., J. Hattori Bot. Lab. 44: 100, 1978 (Hattori 1978a). Bas.: Madotheca tosana Steph., Bull. Herb. Boissier 5 (2): 97, 1897 (Stephani 1897b). * Porella andica (Gottsche) Hässel, Beih. Nova Hedwigia 134: 452, 2009 (Hässel and Rubies 2009). Bas.: Madotheca andica Gottsche, Ann. Sci. Nat. Bot. (sér. 4) 8: 339, 1857 (Gottsche 1857). 371 370 Porella acutifolia var. linguifolia was accepted by So (2002a), but it is conspeciic with Plagiochila viridissima in Hattori (1976f, 1986d). his synonymy is doubtful and further study is necessary to clarify the status. 371 Madotheca andica was transferred to Porella by Hässel (in Hässel and Rubies 2009), but she did not study the type specimen. It may be conspeciic with Plagiochila subsquarrosa. 418 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Porella arboris-vitae (With.) Grolle, Trans. Brit. Bryol. Soc. 5 (4): 770, 1969 (Grolle 1969b). Bas.: Jungermannia arboris-vitae With., Bot. arr. veg. Gr. Brit. 2: 697, 1776 (Withering 1776). ** Porella arboris-vitae subsp. nitidula (C.Massal.) S.Hatt., J. Hattori Bot. Lab. 40: 123, 1976 (Hattori 1976f). Bas.: Madotheca nitidula C.Massal., Bull. Soc. Bot. Ital. 1906: 141, 1906 (Massalongo 1906b). ** Porella baueri (Schifn.) C.E.O.Jensen, Danmarks mosser: 240, 1915 (Jensen 1915). Bas.: Madotheca baueri Schifn., Sitzungsber. deutsch. naturwiss.-med. Vereins Böhmen “Lotos” Prag 48: 346, 1900 (Schifner 1900d). ** Porella bolanderi (Austin) Pearson, List. Canad. Hepat.: 7, 1890 (Pearson 1890). Bas.: Madotheca bolanderi Austin, Bull. Torrey Bot. Club 3 (3): 14, 1872 (Austin 1872). ** Porella borellii (Gola) Parihar, Univ. Allahabad Stud., Bot. 1961-2: 15, 1962 (Parihar 1962). Bas.: Madotheca borellii Gola, Atti Reale Accad. Sci. Torino, Cl. Sci. Fis. Mat. Nat. 49: 760, 1914 (Gola 1914b). *** Porella brachiata (Taylor) Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 334, 1885 (Spruce 1885). Bas.: Madotheca brachiata Taylor, London J. Bot. 6: 341, 1847 (Taylor 1847b). *** Porella brasiliensis (Raddi) Schifn., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 60 (2): 246, 1893 (Schifner 1893a). Bas.: Schulthesia brasiliensis Raddi, Critt. Brasil.: 10, 1822 (Raddi 1822). * Porella brasiliensis var. ciliata (Gottsche, Lindenb. et Nees) Schifn., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 60 (2): 246, 1893 (Schifner 1893a). Bas.: Madotheca brasiliensis β ciliata Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 271, 1845 (Gottsche et al. 1845a). * Porella brasiliensis var. laevior (Gottsche, Lindenb. et Nees) Schifn., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 60 (2): 246, 1893 (Schifner 1893a). Bas.: Madotheca brasiliensis α laevior Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 271, 1845 (Gottsche et al. 1845a). ** Porella caespitans (Steph.) S.Hatt., J. Hattori Bot. Lab. 33: 50, 1970 (Hattori 1970). Bas.: Madotheca caespitans Steph., Mém. Soc. Nat. Sci. Nat. Math. Cherbourg 29: 218, 1894 (Stephani 1894b). 372 ** Porella caespitans var. cordifolia (Steph.) S.Hatt. ex T.Katag. et T.Yamag., Bryol. Res. 10 (5): 133, 2011 (Katagiri and Yamaguchi 2011). Bas.: Madotheca cordifolia Steph., Sp. Hepat. (Stephani) 4: 315, 1910 (Stephani 1910b). ** Porella caespitans subsp. latior (S.Hatt.) S.Hatt., J. Hattori Bot. Lab. 40: 127, 1976 (Hattori 1976f). Bas.: Porella acutifolia subsp. latior S.Hatt., J. Hattori Bot. Lab. 32: 325, 1969 (Hattori 1969). ** Porella caespitans var. nipponica S.Hatt., J. Hattori Bot. Lab. 33: 57, 1970 (Hattori 1970). 372 Porella caespitans was resolved as polyphyletic in the phylogeny of Hentschel et al. (2007b), suggesting that the species is a complex deserving further study. World checklist of hornworts and liverworts 419 ** Porella caespitans var. relexigastria (Pócs) S.Hatt., J. Hattori Bot. Lab. 40: 127, 1976 (Hattori 1976f). Bas.: Porella relexigastria Pócs, J. Hattori Bot. Lab. 31: 71, 1968 (Pócs 1968). ** Porella campylophylla (Lehm. et Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 408, 1877 (Trevisan 1877). Bas.: Jungermannia campylophylla Lehm. et Lindenb., Nov. Stirp. Pug. 6: 40, 1834 (Lehmann 1834). ** Porella campylophylla subsp. lancistipula (Steph.) S.Hatt., J. Hattori Bot. Lab. 44: 102, 1978 (Hattori 1978a). Bas.: Madotheca lancistipula Steph., Sp. Hepat. (Stephani) 6: 524, 1924 (Stephani 1924). ** Porella campylophylla var. ligulifera (Taylor) S.Hatt., J. Hattori Bot. Lab. 32: 333, 1969 (Hattori 1969). Bas.: Madotheca ligulifera Taylor, Nov. Stirp. Pug. 8: 10, 1844 (Lehmann 1844). ** Porella campylophylla var. tixieri (Pócs) S.Hatt., J. Hattori Bot. Lab. 40: 128, 1976 (Hattori 1976f). Bas.: Porella plumosa var. tixieri Pócs, J. Hattori Bot. Lab. 31: 82, 1968 (Pócs 1968). ** Porella canariensis (F.Weber) Underw., Rep. (Annual) Missouri Bot. Gard. 8: 186, 1897 (Trelease 1897). Bas.: Jungermannia platyphylla var. canariensis F.Weber, Hist. Musc. Hepat. Prodr.: 16, 1815 (Weber 1815). * Porella capehorniensis Swails, Nova Hedwigia 19: 244, 1970 (Swails 1970). 373 ** Porella capensis (Gottsche) Mitt., J. Linn. Soc., Bot. 22 (146): 323, 1886 (Mitten 1886b). Bas.: Madotheca capensis Gottsche, Syn. Hepat. 2: 270, 1845 (Gottsche et al. 1845a). * Porella caucasica Steph., Bot. Centralbl. 50 (3): 71, 1892 (Stephani 1892a). ** Porella chenii S.Hatt., J. Hattori Bot. Lab. 30: 129, 1967 (Hattori 1967). *** Porella chilensis (Lehm. et Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 407, 1877 (Trevisan 1877). Bas.: Jungermannia chilensis Lehm. et Lindenb., Nov. Stirp. Pug. 6: 36, 1834 (Lehmann 1834). * Porella chilensis var. antucensis (Gottsche) Hässel, Beih. Nova Hedwigia 134: 452, 2009 (Hässel and Rubies 2009). Bas.: Madotheca chilensis β antucensis Gottsche, Ann. Sci. Nat. Bot. (sér. 4) 8: 340, 1857 (Gottsche 1857). ** Porella chilensis var. fernandeziensis (Herzog) Swails, Nova Hedwigia 19: 236, 1970 (Swails 1970). Bas.: Madotheca chilensis f. fernandeziensis Herzog, Nat. Hist. Juan Fernandez (Botany) 2 (5): 736, 1942 (Herzog 1942a). ** Porella chilensis var. microloba (Herzog) Swails, Nova Hedwigia 19: 236, 1970 (Swails 1970). Bas.: Madotheca chilensis f. microloba Herzog, Nat. Hist. Juan Fernandez (Botany) 2 (5): 736, 1942 (Herzog 1942a). ** Porella chinensis (Steph.) S.Hatt., J. Hattori Bot. Lab. 30: 131, 1967 (Hattori 1967). Bas.: Madotheca chinensis Steph., Mém. Soc. Nat. Sci. Nat. Math. Cherbourg 29: 218, 1894 (Stephani 1894b). 373 Porella capehorniensis may be conspeciic with Plagiochila setigera and the type specimen is probably not from South America (Hässel and Rubies 2009). 420 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Porella chinensis var. crispata Udar et Shaheen, Misc. Bryol. Lichenol. 9 (4): 74, 1982 (Udar and Shaheen 1982). ** Porella chinensis var. decurrens (Steph.) S.Hatt., J. Hattori Bot. Lab. 44: 102, 1978 (Hattori 1978a). Bas.: Madotheca decurrens Steph., Sp. Hepat. (Stephani) 4: 289, 1910 (Stephani 1910b). ** Porella chinensis var. hattorii Udar et Shaheen, Misc. Bryol. Lichenol. 9 (7): 146, 1983 (Udar and Shaheen 1983a). ** Porella chinensis var. irregularis (Steph.) S.Hatt., J. Hattori Bot. Lab. 39: 270, 1975 (Hattori 1975c). Bas.: Madotheca irregularis Steph., Sp. Hepat. (Stephani) 4: 304, 1910 (Stephani 1910b). ** Porella circinnata Lindb., Not. Sällsk. Fauna Fl. Fenn. Förh. 13: 355, 1874 (Lindberg 1874a). *** Porella cordaeana (Huebener) Moore, Proc. Roy. Irish Acad. (ser. 2) 2: 618, 1877 (Moore 1877). Bas.: Jungermannia cordaeana Huebener, Hepaticol. germ.: 291, 1834 (Hübener 1834). ** Porella cranfordii Steph., Hedwigia 28 (4): 270, 1889 (Stephani 1889c). *** Porella crispata (Hook.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 407, 1877 (Trevisan 1877). Bas.: Jungermannia crispata Hook., Pl. crypt. (Hooker): tab. 4b, 1816 (Hooker 1816b). ** Porella cucullistipula Steph., Bull. Soc. Roy. Bot. Belgique, Compt. Rend. 32 (2): 38, 1893 [1894] (Stephani 1893e). ** Porella densifolia (Steph.) S.Hatt., J. Jap. Bot. 20: 109, 1944 (Hattori 1944c). Bas.: Madotheca densifolia Steph., Mém. Soc. Nat. Sci. Nat. Math. Cherbourg 29: 219, 1894 (Stephani 1894b). 374 ** Porella densifolia subsp. andamana S.Hatt., J. Hattori Bot. Lab. 32: 346, 1969 (Hattori 1969). ** Porella densifolia subsp. appendiculata (Steph.) S.Hatt., J. Hattori Bot. Lab. 32: 343, 1969 (Hattori 1969). Bas.: Madotheca appendiculata Steph., Sp. Hepat. (Stephani) 4: 301, 1910 (Stephani 1910b). ** Porella densifolia var. oviloba (Steph.) N.Kitag., Acta Phytotax. Geobot. 19 (2/3): 64, 1962 (Kitagawa 1962b). Bas.: Madotheca oviloba Steph., Sp. Hepat. (Stephani) 4: 312, 1910 (Stephani 1910b). ** Porella densifolia var. paraphyllina (P.C.Chen) Pócs, J. Hattori Bot. Lab. 31: 84, 1968 (Pócs 1968). Bas.: Madotheca paraphyllina P.C.Chen, Feddes Repert. Spec. Nov. Regni Veg. 58: 42, 1955 (Chen 1955). ** Porella densifolia var. pilosa S.Hatt. et K.C.Chang, Bull. Bot. Res., Harbin 8 (2): 44, 1988 (Chang 1988). ** Porella densifolia var. robusta (Steph.) S.Hatt., J. Hattori Bot. Lab. 32: 343, 1969 (Hattori 1969). Bas.: Madotheca robusta Steph., Sp. Hepat. (Stephani) 4: 313, 1910 (Stephani 1910b). 374 Porella densifolia was resolved as paraphyletic in the phylogeny of Hentschel et al. (2007b), suggesting that the species is a complex deserving further study. World checklist of hornworts and liverworts 421 *** Porella elegantula (Mont.) E.A.Hodgs., Svensk Bot. Tidskr. 42 (3): 277, 1948 (Hodgson and Sainsbury 1948). Bas.: Madotheca elegantula Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 255, 1843 (Montagne 1843). ** Porella faurieri (Steph.) S.Hatt., J. Jap. Bot. 20: 109, 1944 (Hattori 1944c). Bas.: Madotheca faurieri Steph., Sp. Hepat. (Stephani) 4: 315, 1910 (Stephani 1910b). ** Porella fengii P.C.Chen et S.Hatt., J. Hattori Bot. Lab. 30: 133, 1967 (Hattori 1967). ** Porella geheebii (Steph.) S.Hatt., Bot. Mag. (Tokyo) 64 (755/756): 114, 1951 (Hattori 1951c). Bas.: Madotheca geheebii Steph., Sp. Hepat. (Stephani) 4: 290, 1910 (Stephani 1910b). ** Porella gracillima Mitt., Trans. Linn. Soc. London, Bot. 3 (3): 202, 1891 (Mitten 1891). ** Porella grandifolia (Steph.) S.Hatt., J. Hattori Bot. Lab. 30: 136, 1967 (Hattori 1967). Bas.: Madotheca grandifolia Steph., Sp. Hepat. (Stephani) 4: 289, 1910 (Stephani 1910b). ** Porella grandiloba Lindb., Acta Soc. Sci. Fenn. 10: 234, 1872 [1873] (Lindberg 1872b). ** Porella grollei S.Hatt., J. Hattori Bot. Lab. 34: 411, 1971 (Hattori 1971b). ** Porella handelii S.Hatt., J. Hattori Bot. Lab. 33: 65, 1970 (Hattori 1970). ** Porella hattorii Udar et Shaheen, Lindbergia 9 (1): 70, 1983 (Udar and Shaheen 1983b). ** Porella hoeana S.Hatt., Misc. Bryol. Lichenol. 7 (5): 86, 1976 (Hattori 1976c). * Porella imbricata Lour., Fl. Cochinch. 2: 683, 1790 (Loureiro 1790). ** Porella inaequalis (Gottsche) Perss., Arch. Soc. Zool. Bot. Fenn. “Vanamo”, suppl. 9: 225, 1955 (Persson 1955). Bas.: Madotheca inaequalis Gottsche, Sp. Hepat. (Stephani) 4: 251, 1910 (Stephani 1910b). ** Porella japonica (Sande Lac.) Mitt., Trans. Linn. Soc. London, Bot. 3 (3): 202, 1891 (Mitten 1891). Bas.: Madotheca japonica Sande Lac., Syn. hepat. jav.: 105, 1856 [1857] (Sande Lacoste 1856b). ** Porella japonica subsp. appalachiana R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 682, 1980 (Schuster 1980c). ** Porella japonica var. densespinosa S.Hatt. et M.X.Zhang, J. Jap. Bot. 60 (11): 324, 1985 (Hattori and Zhang 1985). ** Porella javanica (Gottsche) Inoue, J. Hattori Bot. Lab. 30: 60, 1967 (Inoue 1967a). Bas.: Madotheca javanica Gottsche, Sp. Hepat. (Stephani) 4: 290, 1910 (Stephani 1910b). ** Porella latifolia J.S.Lou et Q.Li, Acta Phytotax. Sin. 25 (6): 482, 1987 (Lou 1987). *** Porella leiboldii (Lehm.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 407, 1877 (Trevisan 1877). Bas.: Madotheca leiboldii Lehm., Nov. Stirp. Pug. 8: 11, 1844 (Lehmann 1844). ** Porella longifolia (Steph.) S.Hatt., J. Hattori Bot. Lab. 32: 351, 1969 (Hattori 1969). Bas.: Madotheca longifolia Steph., Sp. Hepat. (Stephani) 4: 305, 1910 (Stephani 1910b). ** Porella madagascariensis (Nees et Mont.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 407, 1877 (Trevisan 1877). Bas.: Lejeunea madagascariensis Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 5: 57, 1836 (Nees and Montagne 1836). 422 * *** ** *** ** ** *** ** ** ** ** ** ** *** ** *** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Porella maxima (Steph.) M.L.So, Syst. Bot. 27 (1): 11, 2002 (So 2002a). Bas.: Madotheca maxima Steph., Sp. Hepat. (Stephani) 4: 291, 1910 (Stephani 1910b). 375 Porella mexicana (Hampe ex Gottsche, Lindenb. et Nees) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 407, 1877 (Trevisan 1877). Bas.: Madotheca mexicana Hampe ex Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 273, 1845 (Gottsche et al. 1845a). Porella montantii (Steph.) E.W.Jones, Trans. Brit. Bryol. Soc. 4 (3): 460, 1963 (Jones 1963). Bas.: Madotheca montantii Steph., Sp. Hepat. (Stephani) 4: 259, 1910 (Stephani 1910b). Porella navicularis (Lehm. et Lindenb.) Pfeif., Fl. Niederhessen 2: 234, 1855 (Pfeifer 1855). Bas.: Jungermannia navicularis Lehm. et Lindenb., Nov. Stirp. Pug. 6: 38, 1834 (Lehmann 1834). Porella nitens (Steph.) S.Hatt., Fl. E. Himalaya: 525, 1966 (Hattori 1966c). Bas.: Madotheca nitens Steph., Mém. Soc. Nat. Sci. Nat. Math. Cherbourg 29: 220, 1894 (Stephani 1894b). Porella oblongifolia S.Hatt., J. Jap. Bot. 19 (7): 200, 1943 (Hattori 1943c). 376 Porella obtusata (Taylor) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 407, 1877 (Trevisan 1877). Bas.: Madotheca obtusata Taylor, London J. Bot. 5: 380, 1846 (Taylor 1846b). Porella obtusata var. macroloba (Steph.) S.Hatt. et M.X.Zhang, J. Jap. Bot. 60 (11): 325, 1985 (Hattori and Zhang 1985). Bas.: Madotheca macroloba Steph., Sp. Hepat. (Stephani) 4: 292, 1910 (Stephani 1910b). Porella obtusiloba S.Hatt., J. Hattori Bot. Lab. 33: 69, 1970 (Hattori 1970). Porella perrottetiana (Mont.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 408, 1877 (Trevisan 1877). Bas.: Madotheca perrottetiana Mont., Ann. Sci. Nat. Bot. (sér. 2) 18: 15, 1842 (Montagne 1842b). Porella perrottetiana var. angustifolia Pócs, J. Hattori Bot. Lab. 31: 75, 1968 (Pócs 1968). Porella perrottetiana var. ciliatodentata (P.C.Chen et P.C.Wu) S.Hatt., J. Hattori Bot. Lab. 30: 144, 1967 (Hattori 1967). Bas.: Porella ciliatodentata P.C.Chen et P.C.Wu, Obs. l. Hwangs.: 8, 1965 (Chen and Wu 1965). Porella perrottetiana var. triciliata (Steph.) Pócs, J. Hattori Bot. Lab. 31: 75, 1968 (Pócs 1968). Bas.: Madotheca triciliata Steph., Sp. Hepat. (Stephani) 4: 308, 1910 (Stephani 1910b). Porella pinnata L., Sp. Pl. 1: 1106, 1753 (Linnaeus 1753). Porella planifolia J.S.Lou, Coll. Pap. Quing-Zang Huang-Den 1: 277, 1983 (Lou and Wang 1983). Porella platyphylla (L.) Pfeif., Fl. Niederhessen 2: 234, 1855 (Pfeifer 1855). Bas.: Jungermannia platyphylla L., Sp. Pl. 1: 1134, 1753 (Linnaeus 1753). 375 Porella maxima was accepted by So (2002a), but it was treated as conspeciic with Porella viridissima in Hattori (1976f, 1986d). his synonymy is doubtful and further study is necessary to clarify the status. 376 Porella oblongifolia is closely related to Porella densifolia (Hentschel et al. 2007b). World checklist of hornworts and liverworts 423 *** Porella platyphylloidea (Schwein.) Lindb., Morgonbladet (Helsinki) 1876 (287, 10 Dec.): 1, 1876 (Lindberg 1876c). Bas.: Jungermannia platyphylloidea Schwein., Spec. Fl. Amer. Crypt.: 9, 1821 (Schweinitz 1821). ** Porella plicata J.S.Lou, Acta Phytotax. Sin. 18 (1): 119, 1980 (Lou and Wu 1980). ** Porella plumosa (Mitt.) Parihar, Univ. Allahabad Stud., Bot. 1961-2: 17, 1962 (Parihar 1962). Bas.: Madotheca plumosa Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 108, 1860 [1861] (Mitten 1860c). ** Porella prolixa (Gottsche) E.W.Jones, Trans. Brit. Bryol. Soc. 4 (3): 460, 1963 (Jones 1963). Bas.: Madotheca prolixa Gottsche, Sp. Hepat. (Stephani) 4: 260, 1910 (Stephani 1910b). ** Porella pulcherrima Herzog et S.Hatt., Bull. Natl. Sci. Mus. Tokyo, B 12 (1): 34, 1986 (Hattori 1986a). *** Porella relexa (Lehm. et Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 408, 1877 (Trevisan 1877). Bas.: Jungermannia relexa Lehm. et Lindenb., Nov. Stirp. Pug. 5: 5, 1833 (Lehmann 1833). ** Porella revoluta (Lehm. et Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 407, 1877 (Trevisan 1877). Bas.: Jungermannia revoluta Lehm. et Lindenb., Nov. Stirp. Pug. 4: 18, 1832 (Lehmann 1832). ** Porella revoluta var. propinqua (C.Massal.) S.Hatt., J. Hattori Bot. Lab. 30: 148, 1967 (Hattori 1967). Bas.: Madotheca propinqua C.Massal., Hepat. Shen-si: 27, 1897 (Massalongo 1897). ** Porella roellii Steph., Bot. Centralbl. 45: 203, 1891 (Röll 1891). *** Porella saccata M.L.So, New Zealand J. Bot. 43 (1): 302, 2005 (So 2005b). ** Porella sichuanensis S.Hatt. et K.C.Chang, Bull. Bot. Res., Harbin 8 (2): 43, 1988 (Chang 1988). ** Porella spinulosa (Steph.) S.Hatt., J. Hattori Bot. Lab. 33: 74, 1970 (Hattori 1970). Bas.: Madotheca spinulosa Steph., Sp. Hepat. (Stephani) 6: 529, 1924 (Stephani 1924). *** Porella squamulifera (Taylor) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 407, 1877 (Trevisan 1877). Bas.: Madotheca squamulifera Taylor, London J. Bot. 5: 378, 1846 (Taylor 1846b). ** Porella stephaniana (C.Massal.) S.Hatt., J. Hattori Bot. Lab. 5: 81, 1951 (Hattori 1951b). Bas.: Madotheca stephaniana C.Massal., Hepat. Shen-si: 23, 1897 (Massalongo 1897). 377 ** Porella subdentata (Mitt.) Steph., Hedwigia 30 (5): 203, 1891 (Stephani 1891a). Bas.: Madotheca subdentata Mitt., J. Proc. Linn. Soc., Bot. 7 (27): 167, 1863 (Mitten 1863). ** Porella subdentata var. camerunensis E.W.Jones, Trans. Brit. Bryol. Soc. 4 (3): 456, 1963 (Jones 1963). ** Porella subobtusa (Steph.) S.Hatt., J. Jap. Bot. 20: 111, 1944 (Hattori 1944c). Bas.: Madotheca subobtusa Steph., Sp. Hepat. (Stephani) 4: 311, 1910 (Stephani 1910b). ** Porella subparaphyllina J.S.Lou, Acta Phytotax. Sin. 25 (6): 483, 1987 (Lou 1987). 377 Porella stephaniana is closely related to Porella densifolia (Hentschel et al. 2007b). 424 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Porella subsquarrosa (Nees et Mont.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 407, 1877 (Trevisan 1877). Bas.: Lejeunea subsquarrosa Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 5: 57, 1836 (Nees and Montagne 1836). *** Porella swailsii Grolle, J. Bryol. 10 (3): 270, 1979 (Grolle 1979b). Nom. nov. pro Madotheca apiculata Herzog, Feddes Repert. Spec. Nov. Regni Veg. 57 (1/2): 197, 1955 (Herzog 1955). *** Porella swartziana (F.Weber) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 407, 1877 (Trevisan 1877). Bas.: Jungermannia swartziana F.Weber, Hist. Musc. Hepat. Prodr.: 18, 1815 (Weber 1815). ** Porella triquetra (Steph.) E.W.Jones, Trans. Brit. Bryol. Soc. 4 (3): 454, 1963 (Jones 1963). Bas.: Madotheca triquetra Steph., Bot. Jahrb. Syst. 20 (3): 321, 1895 (Stephani 1895a). ** Porella truncata J.S.Lou, Acta Phytotax. Sin. 18 (1): 119, 1980 (Lou and Wu 1980). ** Porella ulophylla (Steph.) S.Hatt., Bull. Tokyo Sci. Mus. 11: 92, 1944 (Hattori 1944d). Bas.: Madotheca ulophylla Steph., Bull. Herb. Boissier 5 (2): 97, 1897 (Stephani 1897b). ** Porella undatorevoluta J.S.Lou, Acta Phytotax. Sin. 25 (6): 485, 1987 (Lou 1987). ** Porella urceolata S.Hatt., J. Hattori Bot. Lab. 33: 66, 1970 (Hattori 1970). ** Porella urogea (C.Massal.) S.Hatt., J. Hattori Bot. Lab. 32: 349, 1969 (Hattori 1969). Bas.: Madotheca urogea C.Massal., Hepat. Shen-si: 28, 1897 (Massalongo 1897). ** Porella vallis-gratiae (Gottsche) E.W.Jones, Trans. Brit. Bryol. Soc. 4 (3): 450, 1963 (Jones 1963). Bas.: Madotheca vallis-gratiae Gottsche, Sp. Hepat. (Stephani) 4: 261, 1910 (Stephani 1910b). ** Porella variabilis (Kashyap et R.S.Chopra) Parihar, Univ. Allahabad Stud., Bot. 1961-2: 17, 1962 (Parihar 1962). Bas.: Madotheca variabilis Kashyap et R.S.Chopra, Liverworts W. Himal. 2: 33, 1932 (Kashyap and Chopra 1932). ** Porella vernicosa Lindb., Acta Soc. Sci. Fenn. 10: 223, 1872 [1873] (Lindberg 1872b). ** Porella viridissima (Mitt.) Grolle, J. Hattori Bot. Lab. 36: 83, 1972 [1973] (Grolle and Schultze-Motel 1972). Bas.: Madotheca viridissima Mitt., Fl. vit.: 411, 1871 [1873] (Mitten 1871). ** Porella wataugensis (Sull.) Underw. ex M.Howe, Bull. Torrey Bot. Club 24 (11): 519, 1897 (Howe 1897b). Bas.: Madotheca wataugensis Sull., Musc. Hepat. U.S.: 700, 1856 (Sullivant 1856). Radulineae R.M.Schust. *** Radulaceae Müll.Frib. by M.A.M. Renner he treatment of Radulaceae follows Devos et al. (2011). Taxonomic and nomenclatural notes can also be found in Renner et al. (2013b, 2014). World checklist of hornworts and liverworts 425 *** Radula Dumort., Commentat. Bot. (Dumortier): 112, 1822 (Dumortier 1822) nom. conserv. *** subg. Amentuloradula Devos, M.A.M.Renner, Gradst., A.J.Shaw et Vanderp., Taxon 60 (6): 1630, 2011 (Devos et al. 2011). *** Radula amentulosa Mitt., Bonplandia 9 (24): 367, 1861 (Mitten 1861). *** Radula aneurismalis (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 262, 1845 (Gottsche et al. 1845a). Bas.: Jungermannia aneurismalis Hook.f. et Taylor, London J. Bot. 4: 86, 1845 (Hooker and Taylor 1845). ** Radula ceylanica K.Yamada, J. Jap. Bot. 50 (12): 373, 1975 (Yamada 1975b). *** Radula issifolia Steph., Sp. Hepat. (Stephani) 6: 507, 1924 (Stephani 1924). *** Radula formosa (C.F.W.Meissn. ex Spreng.) Nees, Syn. Hepat. 2: 258, 1845 (Gottsche et al. 1845a). Bas.: Jungermannia formosa C.F.W.Meissn. ex Spreng. Syst. Veg. (ed. 16) [Sprengel] 4 (2): 325, 1827 (Sprengel 1827b). *** Radula helix (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 260, 1845 (Gottsche et al. 1845a). Bas.: Jungermannia helix Hook.f. et Taylor, London J. Bot. 3: 475, 1844 (Hooker and Taylor 1844b). *** Radula hicksiae K.Yamada, Cryptog. Bryol. Lichénol. 5 (1/2): 191, 1984 (Yamada 1984a). *** Radula iwatsukii K.Yamada, J. Hattori Bot. Lab. 45: 275, 1979 (Yamada 1979b). ** Radula morobeana K.Yamada et Piippo, Ann. Bot. Fenn. 26 (4): 358, 1989 (Yamada and Piippo 1989). ** Radula multiamentula E.A.Hodgs., Rec. Domin. Mus. 4 (11): 122, 1962 (Hodgson 1962a). *** Radula ornata E.A.Br. et Pócs, Telopea 9 (3): 436, 2001 (Brown and Pócs 2001). *** Radula physoloba Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 255, 1843 (Montagne 1843). *** Radula pseudoscripta M.A.M.Renner, New Zealand J. Bot. 44 (3): 340, 2006 (Renner 2006). ** Radula queenslandica K.Yamada, J. Hattori Bot. Lab. 62: 192, 1987 (Yamada 1987). *** Radula scariosa Mitt., Bonplandia 9 (24): 367, 1861 (Mitten 1861). *** Radula splendida M.A.M.Renner et Devos, Nova Hedwigia 90 (1/2): 113, 2010 (Renner et al. 2010a). * Radula squarrosa K.Yamada, J. Jap. Bot. 65 (1): 1, 1990 (Yamada 1990). 378 *** Radula thiersiae K.Yamada, J. Hattori Bot. Lab. 62: 198, 1987 (Yamada 1987). *** Radula uvifera (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 258, 1845 (Gottsche et al. 1845a). Bas.: Jungermannia uvifera Hook.f. et Taylor, London J. Bot. 3: 292 [392], 1844 (Hooker and Taylor 1844a). * Radula vagans Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 85, 1911 (Stephani 1911b). ** Radula verrucosa K.Yamada, J. Hattori Bot. Lab. 45: 277, 1979 (Yamada 1979b). 378 Radula squarrosa is probably conspeciic with Radula morobeana. 426 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** subg. Cladoradula Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 315, 1885 (Spruce 1885). ** Radula auriculata Steph., Bull. Herb. Boissier 5 (2): 105, 1897 (Stephani 1897b). * Radula bipinnata Mitt., J. Proc. Linn. Soc., Bot. 7 (27): 166, 1863 (Mitten 1863). 379 *** Radula boryana (F.Weber) Nees ex Mont., Ann. Sci. Nat. Bot. (sér. 2) 18: 13, 1842 (Montagne 1842b). Bas.: Jungermannia boryana F.Weber, Hist. Musc. Hepat. Prodr.: 58, 1815 (Weber 1815). *** Radula campanigera Mont., London J. Bot. 3: 634, 1844 (Montagne 1844a). *** Radula campanigera subsp. obiensis (S.Hatt.) K.Yamada, J. Hattori Bot. Lab. 45: 309, 1979 (Yamada 1979b). Bas.: Radula obiensis S.Hatt., Bull. Tokyo Sci. Mus. 11: 83, 1944 (Hattori 1944d). ** Radula chinensis Steph., Nuovo Giorn. Bot. Ital. (n.ser.) 13 (4): 355, 1906 (Levier 1906). *** Radula gottscheana Taylor, London J. Bot. 5: 374, 1846 (Taylor 1846b). *** Radula perrottetii Gottsche, Hedwigia 23 (10): 154, 1884 (Stephani 1884a). ** Radula tenax Lindb., Acta Soc. Sci. Fenn. 10: 492, 1875 (Lindberg 1875). *** subg. Dactyloradula Devos, M.A.M.Renner, Gradst., A.J.Shaw et Vanderp., Taxon 60 (6): 1630, 2011 (Devos et al. 2011). *** Radula brunnea Steph., Sp. Hepat. (Stephani) 4: 232, 1910 (Stephani 1910b). *** subg. Metaradula R.M.Schust., Phytologia 56 (2): 69, 1984 (Schuster 1984). ** Radula acuminata Steph., Sp. Hepat. (Stephani) 4: 230, 1910 (Stephani 1910b). 380 ** Radula aguirrei R.M.Schust., Phytotaxa 202 (1): 70, 2015 (Söderström et al. 2015c). Based on: Radula aguirrei R.M.Schust., J. Hattori Bot. Lab. 70: 56, 1991 (Schuster 1991a), nom. inval. ** Radula anisotoma M.A.M.Renner, PhytoKeys 27: 30, 2013 (Renner et al. 2013a). *** Radula assamica Steph., Hedwigia 23 (10): 151, 1884 (Stephani 1884a). *** Radula australiana K.Yamada, J. Hattori Bot. Lab. 51: 323, 1982 (Yamada 1982a). *** Radula buccinifera (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 261, 1845 (Gottsche et al. 1845a). Bas.: Jungermannia buccinifera Hook.f. et Taylor, London J. Bot. 3: 580, 1844 (Hooker and Taylor 1844c). *** Radula demissa M.A.M.Renner, PhytoKeys 27: 53, 2013 (Renner et al. 2013a). ** Radula evansii Castle, Ann. Bryol. 11: 37, 1938 (Castle 1938). ** Radula laccida Lindenb. et Gottsche, Syn. Hepat. 5: 726, 1847 (Gottsche et al. 1847). ** Radula laccida var. brachycalyx Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 321, 1885 (Spruce 1885). *** Radula foricata M.A.M.Renner, Austral. Syst. Bot. 26 (4): 307, 2013 (Renner et al. 2013c). ** Radula grevilleana Taylor, Ann. Mag. Nat. Hist. 20 (135): 380, 1847 (Taylor 1847a). 379 Radula bipinnata is distinct from Radula boryana according to molecular evidence (Devos et al. 2011). 380 Radula acuminata may be conspeciic with Radula tjibodensis. World checklist of hornworts and liverworts 427 *** Radula imposita M.A.M.Renner, PhytoKeys 27: 65, 2013 (Renner et al. 2013a). ** Radula jovetiana K.Yamada, Cryptog. Bryol. Lichénol. 5 (1/2): 193, 1984 (Yamada 1984a). ** Radula kilgourii M.A.M.Renner, Austral. Syst. Bot. 26 (4): 313, 2013 (Renner et al. 2013c). ** Radula loriana Castle, J. Hattori Bot. Lab. 21: 6, 1959 (Castle 1959). *** Radula mammosa Spruce, Mem. Torrey Bot. Club 1 (3): 127, 1890 (Spruce 1890). *** Radula mittenii Steph., Hedwigia 23 (10): 148, 1884 (Stephani 1884a). *** Radula myriopoda M.A.M.Renner, Austral. Syst. Bot. 26 (4): 323, 2013 (Renner et al. 2013c). *** Radula notabilis M.A.M.Renner, PhytoKeys 27: 77, 2013 (Renner et al. 2013a). ** Radula nymannii Steph., Sp. Hepat. (Stephani) 4: 229, 1910 (Stephani 1910b). ** Radula protensa Lindenb., Bot. Zeitung (Berlin) 6 (25): 462, 1848 (Meissner 1848). ** Radula protensa var. erectilobula Schifn., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 60 (2): 247, 1893 (Schifner 1893a). ** Radula pseudolaccida E.W.Jones, J. Bryol. 9 (4): 501, 1977 [1978] (Jones 1977). ** Radula psychosis M.A.M.Renner, Austral. Syst. Bot. 26 (4): 328, 2013 (Renner et al. 2013c). *** Radula ratkowskiana K.Yamada, J. Jap. Bot. 59 (3): 94, 1984 (Yamada 1984b). *** Radula robinsonii Steph., Sp. Hepat. (Stephani) 4: 214, 1910 (Stephani 1910b). ** Radula stenocalyx Mont., Ann. Sci. Nat. Bot. (sér. 4) 3 (5): 315, 1855 (Montagne 1855). *** Radula strangulata Hook.f. et Taylor, London J. Bot. 5: 377, 1846 (Taylor 1846b). * Radula tjibodensis K.I.Goebel, Ann. Jard. Bot. Buitenzorg 7 (1): 53, 1888 (Goebel 1888). 381 *** Radula ventricosa Steph., Sp. Hepat. (Stephani) 4: 187, 1910 (Stephani 1910b). *** Radula yanoella R.M.Schust., Phytologia 56 (2): 72, 1984 (Schuster 1984). subg. Odontoradula K.Yamada, J. Hattori Bot. Lab. 45: 209, 1979 (Yamada 1979b). Radula acuta Mitt., Fl. vit.: 410, 1871 [1873] (Mitten 1871). Radula allisonii Castle, Rev. Bryol. Lichénol. 31 (3/4): 148, 1962 [1963] (Castle 1962). Radula amoena Herzog, Mitt. Inst. Allg. Bot. Hamburg 7 (3): 192, 1931 (Herzog 1931a). ** Radula anceps Sande Lac., Ned. Kruidk. Arch. 3: 419, 1854 [1855] (Sande Lacoste 1854). * Radula crenulata Schifn., Leberm., Forschungsr. Gazelle 4 (4): 21, 1890 (Schifner 1890). 382 *** Radula cuspidata Steph., Sp. Hepat. (Stephani) 4: 156, 1910 (Stephani 1910b). *** Radula decora Gottsche, Hedwigia 23 (10): 145, 1884 (Stephani 1884a). *** ** ** ** 381 Radula tjibodensis is a doubtful taxon. he type specimen has not been found. 382 Radula crenulata is conspeciic with Radula acuta in So (2006) and possibly conspeciic with Radula apiculata in Yamada and Piippo (1989), but difers morphologically from both and needs to be further studied. 428 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Radula emarginata K.Yamada et Piippo, Ann. Bot. Fenn. 26 (4): 352, 1989 (Yamada and Piippo 1989). ** Radula kojana Steph., Bull. Herb. Boissier 5 (2): 105, 1897 (Stephani 1897b). *** Radula lacerata Steph., Rev. Bryol. 35 (2): 33, 1908 (Stephani 1908l). *** Radula novae-hollandiae Hampe, Nov. Stirp. Pug. 7: 24, 1838 (Lehmann 1838). *** Radula ocellata K.Yamada, J. Hattori Bot. Lab. 45: 209, 1979 (Yamada 1979b). *** Radula plicata Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 154, 1854 (Mitten 1854). ** Radula pugioniformis M.A.M.Renner, PhytoKeys 27: 84, 2013 (Renner et al. 2013a). *** Radula pulchella Mitt., Hedwigia 23 (10): 149, 1884 (Stephani 1884a). *** Radula retrolexa Taylor, London J. Bot. 5: 378, 1846 (Taylor 1846b). *** Radula tasmanica Steph., Sp. Hepat. (Stephani) 4: 212, 1910 (Stephani 1910b). *** Radula weymouthiana Steph., Sp. Hepat. (Stephani) 4: 190, 1910 (Stephani 1910b). *** *** ** *** ** ** ** *** *** ** ** ** *** ** ** *** *** ** ** subg. Radula Radula acutiloba Steph., Hedwigia 28 (4): 271, 1889 (Stephani 1889c). 383 Radula appressa Mitt., Philos. Trans. 168: 397, 1879 (Mitten 1879). Radula aquilegia (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 260, 1845 (Gottsche et al. 1845a). Bas.: Jungermannia aquilegia Hook.f. et Taylor, London J. Bot. 3: 291 [391], 1844 (Hooker and Taylor 1844a). Radula australis Austin, Bot. Bull. (Hanover) 1 (7): 32, 1876 (Austin 1876b). Radula borneensis Steph., Sp. Hepat. (Stephani) 4: 209, 1910 (Stephani 1910b). Radula caduca K.Yamada, J. Hattori Bot. Lab. 45: 225, 1979 (Yamada 1979b). Radula carringtonii J.B.Jack, Flora 64 (25): 385, 1881 (Jack 1881). Radula complanata (L.) Dumort., Syll. Jungerm. Europ.: 38, 1831 (Dumortier 1831). Bas.: Jungermannia complanata L., Sp. Pl. 1: 1133, 1753 (Linnaeus 1753). Radula constricta Steph., Sp. Hepat. (Stephani) 6: 506, 1924 (Stephani 1924). Radula evelynae K.Yamada, J. Jap. Bot. 50 (4): 115, 1975 (Yamada 1975a). Radula fendleri Gottsche, Hedwigia 23 (10): 146, 1884 (Stephani 1884a). 384 Radula grandis Steph., J. Linn. Soc., Bot. 29 (201): 271, 1892 (Stephani 1892b). Radula japonica Gottsche, Hedwigia 23 (10): 152, 1884 (Stephani 1884a). Radula javanica Gottsche, Syn. Hepat. 2: 257, 1845 (Gottsche et al. 1845a). Radula jonesii Bouman, Dirkse et K.Yamada, J. Bryol. 15 (1): 161, 1988 (Bouman et al. 1988). Radula lindenbergiana Gottsche ex C.Hartm., Handb. Skand. l. (ed.9): 98, 1864 (Hartman 1864). Radula madagascariensis Gottsche, Abh. Naturwiss. Vereins Bremen 7: 349, 1882 (Gottsche 1882). Radula marojezica E.W.Jones, J. Bryol. 17 (2): 307, 1992 (Jones 1992). 383 Radula acutiloba is tentatively placed in subg. Radula. he specimen labelled Radula acutiloba in Devos et al. (2011) was misidentiied and should be Radula australiana. 384 Radula fendleri may be conspeciic with Radula madagascariensis. World checklist of hornworts and liverworts * ** ** ** ** * ** ** *** *** *** ** ** ** *** ** *** 429 Radula multilora Gottsche ex Schifn., Leberm., Forschungsr. Gazelle 4 (4): 20, 1890 (Schifner 1890). 385 Radula novoguineensis K.Yamada et Piippo, Ann. Bot. Fenn. 26 (4): 360, 1989 (Yamada and Piippo 1989). Radula obconica Sull., Manual (Gray): 688, 1848 (Gray 1848). Radula obtusiloba Steph., Bull. Herb. Boissier 5 (2): 105, 1897 (Stephani 1897b). Radula obtusiloba subsp. polyclada (A.Evans) S.Hatt., J. Hattori Bot. Lab. 29: 275, 1966 (Hattori 1966d). Bas.: Radula polyclada A.Evans, Bull. Torrey Bot. Club 41 (12): 607, 1914 [1915] (Evans 1914a). Radula oceania Castle, Rev. Bryol. Lichénol. 33 (3/4): 390, 1965 (Castle 1965). 386 Radula oreopsis M.A.M.Renner, Telopea 17: 123, 2014 (Renner 2014). Radula portoricensis Steph., Hedwigia 27 (11/12): 298, 1888 (Stephani 1888c). Radula prolifera Arnell, Ark. Bot. 13 (2): 12, 1913 (Arnell 1913). Radula quadrata Gottsche, Syn. Hepat. 2: 255, 1845 (Gottsche et al. 1845a). Radula relexa Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 255, 1843 (Montagne 1843). Radula sharpii K.Yamada, J. Jap. Bot. 60 (9): 260, 1985 (Yamada 1985a). Radula sumatrana Steph., Sp. Hepat. (Stephani) 4: 204, 1910 (Stephani 1910b). Radula tokiensis Steph., Hedwigia 23 (10): 150, 1884 (Stephani 1884a). Radula van-zantenii K.Yamada, J. Hattori Bot. Lab. 45: 260, 1979 (Yamada 1979b). Radula varilobula Castle, J. Hattori Bot. Lab. 21: 19, 1959 (Castle 1959). Radula wichurae Steph., Sp. Hepat. (Stephani) 4: 168, 1910 (Stephani 1910b). *** subg. Volutoradula Devos, M.A.M.Renner, Gradst., A.J.Shaw et Vanderp., Taxon 60 (6): 1629, 2011 (Devos et al. 2011). ** Radula ankeinensis Gottsche, Hedwigia 23 (10): 152, 1884 (Stephani 1884a). ** Radula antilleana Castle, J. Hattori Bot. Lab. 21: 48, 1959 (Castle 1959). ** Radula comorensis Steph., Hedwigia 23 (9): 132, 1884 (Stephani 1884c). ** Radula cubensis K.Yamada, J. Hattori Bot. Lab. 54: 241, 1983 (Yamada 1983). ** Radula diversifolia Steph., Sp. Hepat. (Stephani) 4: 212, 1910 (Stephani 1910b). *** Radula eggersii K.Yamada, J. Hattori Bot. Lab. 82: 339, 1997 (Yamada 1997). ** Radula episcia Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 318, 1885 (Spruce 1885). ** Radula loridana Castle, Rev. Bryol. Lichénol. 36 (1/2): 1, 1968 [1969] (Castle 1968). ** Radula fulvifolia (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 261, 1845 (Gottsche et al. 1845a). Bas.: Jungermannia fulvifolia Hook.f. et Taylor, London J. Bot. 4: 85, 1845 (Hooker and Taylor 1845). *** Radula hastata Steph., Sp. Hepat. (Stephani) 4: 163, 1910 (Stephani 1910b). * Radula holstiana Steph., Bot. Jahrb. Syst. 20 (3): 320, 1895 (Stephani 1895a). 385 Radula multilora is similar to, and may be conspeciic with Radula relexa, but it was accepted by Renner and de Lange (2009). 386 Radula oceania was considered conspeciic with Radula javanica by So (2006), but molecular data does not support it (Renner 2014). 430 *** ** ** ** ** ** ** ** ** *** * ** ** ** ** ** ** ** ** ** ** *** *** ** ** *** ** ** ** ** ** *** ** ** *** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Radula holtii Spruce, J. Bot. 25: 209, 1887 (Spruce 1887b). Radula husnotii Castle, J. Hattori Bot. Lab. 21: 45, 1959 (Castle 1959). Radula inlexa Gottsche, Hedwigia 23 (10): 148, 1884 (Stephani 1884a). Radula kegelii Gottsche ex Steph., Hedwigia 23 (10): 152, 1884 (Stephani 1884a). Radula macroloba Steph., Bull. Soc. Roy. Bot. Belgique 31: 121, 1892 (Stephani 1892c). Radula mazarunensis K.Yamada, Trop. Bryol. 1: 38, 1989 (Gradstein and Florschütz-de Waard 1989). Radula mexicana Lindenb. et Gottsche, Mexik. Leverm.: 150, 1863 (Gottsche 1863). Radula microloba Gottsche, Syn. Hepat. 2: 259, 1845 (Gottsche et al. 1845a). Radula neotropica Castle, J. Hattori Bot. Lab. 21: 31, 1959 (Castle 1959). Radula nudicaulis Steph., Sp. Hepat. (Stephani) 4: 174, 1910 (Stephani 1910b). Radula nudicaulis var. delicatula P.Allorge et V.Allorge, Rev. Bryol. Lichénol. 19 (1/2): 106, 1950 (Allorge and Allorge 1950). Radula pocsii K.Yamada, J. Hattori Bot. Lab. 54: 245, 1983 (Yamada 1983). Radula recubans Taylor, London J. Bot. 5: 376, 1846 (Taylor 1846b). Radula saccatiloba Steph., Hedwigia 23 (8): 129, 1884 (Stephani 1884b). Radula schaefer-verwimpii K.Yamada, J. Jap. Bot. 65 (1): 3, 1990 (Yamada 1990). Radula schoieldiana K.Yamada, J. Hattori Bot. Lab. 82: 337, 1997 (Yamada 1997). Radula stipatilora Steph., Sp. Hepat. (Stephani) 4: 159, 1910 (Stephani 1910b). Radula striata Mitt., Hedwigia 23 (10): 155, 1884 (Stephani 1884a). Radula subinlata Lindenb. et Gottsche, Syn. Hepat. 5: 724, 1847 (Gottsche et al. 1847). Radula sullivantii Austin, Hepat. bor.-amer.: 22, 1873 (Austin 1873). Radula tenera Mitt., Hedwigia 23 (10): 149, 1884 (Stephani 1884a). Radula voluta Taylor, Syn. Hepat. 2: 255, 1845 (Gottsche et al. 1845a). Incertae sedis Radula acutangula Steph., Bull. Herb. Boissier 5 (10): 848, 1897 (Stephani 1897c). Radula angulata Steph., Hedwigia 23 (8): 114, 1884 (Stephani 1884b). Radula bogotensis Steph., Hedwigia 23 (8): 115, 1884 (Stephani 1884b). Radula bolanderi Gottsche, Hedwigia 23 (10): 145, 1884 (Stephani 1884a). Radula boninensis Furuki et K.Yamada, J. Jap. Bot. 61 (10): 312, 1986 (Furuki and Yamada 1986). Radula brasilica K.Yamada, J. Hattori Bot. Lab. 74: 35, 1993 (Yamada 1993). Radula caespitosa Steph., Hedwigia 27 (3/4): 107, 1888 (Stephani 1888d). Radula campanulata Lindenb. et Gottsche, Syn. Hepat. 2: 256, 1845 (Gottsche et al. 1845a). Radula castlei Grolle, Bryologist 73 (4): 662, 1970 (Grolle 1970a). Radula cavifolia Hampe ex Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 259, 1845 (Gottsche et al. 1845a). Radula cochabambaensis K.Yamada, J. Hattori Bot. Lab. 74: 37, 1993 (Yamada 1993). Radula conferta Lindenb. et Gottsche, Syn. Hepat. 5: 729, 1847 (Gottsche et al. 1847). Radula cordata Mitt., Fl. vit.: 410, 1871 [1873] (Mitten 1871). World checklist of hornworts and liverworts ** *** * ** ** ** ** ** ** ** ** ** ** ** ** ** ** ** ** ** *** * *** *** *** *** ** ** *** ** ** ** 431 Radula costaricensis Gottsche, J. Bot. 15: 226, 1877 (Polakowski 1877). Radula curvilobula M.L.So, J. Hattori Bot. Lab. 98: 176, 2005 (So 2005a). Radula decurrens Mitt., Fl. vit.: 410, 1871 [1873] (Mitten 1871). 387 Radula densifolia Castle, Rev. Bryol. Lichénol. 33 (3/4): 385, 1965 (Castle 1965). Radula diaphana K.I.Goebel, Organogr. Pl., ed. 2, 2 (1): 677, 1915 (Goebel 1915). Radula dolabrata K.Yamada, J. Jap. Bot. 60 (9): 257, 1985 (Yamada 1985a). Radula elliottii Castle, J. Hattori Bot. Lab. 21: 12, 1959 (Castle 1959). Radula falcata Steph., Hedwigia 23 (8): 115, 1884 (Stephani 1884b). Radula fauriana Steph., Sp. Hepat. (Stephani) 4: 207, 1910 (Stephani 1910b). Radula fernandezana Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 84, 1911 (Stephani 1911b). Radula lavifolia (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 259, 1845 (Gottsche et al. 1845a). Bas.: Jungermannia lavifolia Hook.f. et Taylor, London J. Bot. 3: 476, 1844 (Hooker and Taylor 1844b). Radula fujitae Furuki, Bryol. Res. 9 (5): 143, 2007 (Furuki 2007). Radula galapagona Steph., Sp. Hepat. (Stephani) 4: 176, 1910 (Stephani 1910b). Radula gedena Gottsche, Hedwigia 23 (10): 146, 1884 (Stephani 1884a). Radula gracilis Mitt., Hedwigia 23 (10): 147, 1884 (Stephani 1884a). Radula gradsteinii K.Yamada, Trop. Bryol. 1: 37, 1989 (Gradstein and Florschützde Waard 1989). Radula grandifolia Steph., Sp. Hepat. (Stephani) 4: 184, 1910 (Stephani 1910b). Radula grollei K.Yamada et Piippo, Ann. Bot. Fenn. 26 (4): 379, 1989 (Yamada and Piippo 1989). Radula guyanensis K.Yamada, Trop. Bryol. 1: 38, 1989 (Gradstein and Florschützde Waard 1989). Radula hattorii K.Yamada, J. Jap. Bot. 60 (9): 259, 1985 (Yamada 1985a). Radula hawaiica M.L.So, J. Hattori Bot. Lab. 98: 177, 2005 (So 2005a). Radula hedingeri K.I.Goebel, Ann. Jard. Bot. Buitenzorg 7 (1): 51, 1888 (Goebel 1888). Radula inouei K.Yamada, J. Hattori Bot. Lab. 45: 262, 1979 (Yamada 1979b). Radula involvens Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 325, 1885 (Spruce 1885). Radula iwatsukiana K.Yamada, J. Hattori Bot. Lab. 58: 114, 1985 (Yamada 1985b). Radula jamaicensis Pearson, Ann. Bryol. 4: 103, 1931 (Pearson 1931b). Radula jamesonii Taylor, London J. Bot. 5: 375, 1846 (Taylor 1846b). Radula kinabaluensis K.Yamada, Misc. Bryol. Lichenol. 6 (6): 97, 1973 (Yamada 1973b). Radula kitagawae K.Yamada, J. Hattori Bot. Lab. 58: 116, 1985 (Yamada 1985b). Radula koponenii K.Yamada et Piippo, Ann. Bot. Fenn. 26 (4): 364, 1989 (Yamada and Piippo 1989). Radula kurzii Steph., Hedwigia 23 (10): 153, 1884 (Stephani 1884a). Radula laxiramea Steph., Sp. Hepat. (Stephani) 4: 178, 1910 (Stephani 1910b). 387 Radula decurrens may be conspeciic with Radula relexa. 432 ** ** *** *** ** * *** *** ** ** ** *** ** ** ** *** *** ** ** ** ** ** ** ** * *** *** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Radula leiboldii Steph., Hedwigia 23 (8): 116, 1884 (Stephani 1884b). Radula lewisii K.Yamada, J. Hattori Bot. Lab. 74: 39, 1993 (Yamada 1993). Radula ligula Steph., Sp. Hepat. (Stephani) 4: 228, 1910 (Stephani 1910b). Radula lingulata Gottsche, Syn. Hepat. 2: 260, 1845 (Gottsche et al. 1845a). Radula longiloba K.Yamada, J. Hattori Bot. Lab. 54: 243, 1983 (Yamada 1983). Radula longispica Steph., Sp. Hepat. (Stephani) 4: 183, 1910 (Stephani 1910b). 388 Radula marginata Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 261, 1845 (Gottsche et al. 1845a). Nom. nov. pro Jungermannia marginata Hook.f. et Taylor, London J. Bot. 3: 566, 1844 (Hooker and Taylor 1844a), nom. illeg. Radula mauiensis M.L.So, J. Hattori Bot. Lab. 98: 178, 2005 (So 2005a). Radula microlobula Castle, J. Hattori Bot. Lab. 21: 35, 1959 (Castle 1959). Radula minutilobula K.Yamada et Piippo, Ann. Bot. Fenn. 26 (4): 377, 1989 (Yamada and Piippo 1989). Radula mizutanii K.Yamada, J. Jap. Bot. 48 (5): 134, 1973 (Yamada 1973a). Radula nigra Pearson, J. Linn. Soc., Bot. 46 (305): 31, 1922 (Pearson 1922b). Radula nilgiriensis Udar et D.Kumar, J. Indian Bot. Soc. 61: 177, 1982 (Udar and Kumar 1982b). Radula norrisii K.Yamada et Piippo, Ann. Bot. Fenn. 26 (4): 374, 1989 (Yamada and Piippo 1989). Radula novivrieseana K.Yamada, J. Hattori Bot. Lab. 51: 326, 1982 (Yamada 1982a). Radula novocaledonica Hürl. et K.Yamada, J. Jap. Bot. 54 (8): 238, 1979 (Hürlimann and Yamada 1979). Radula novocaledoniensis K.Yamada, J. Hattori Bot. Lab. 58: 120, 1985 (Yamada 1985b). Radula obovata Castle, J. Hattori Bot. Lab. 21: 16, 1959 (Castle 1959). Radula obscura Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 107, 1860 [1861] (Mitten 1860c). Radula okamurana Steph., Sp. Hepat. (Stephani) 4: 209, 1910 (Stephani 1910b). Radula onraedtii K.Yamada, Misc. Bryol. Lichenol. 8 (6): 113, 1979 (Yamada 1979a). Radula opaciuscula (Spruce) Castle, J. Hattori Bot. Lab. 21: 22, 1959 (Castle 1959). Bas.: Radula episcia var. opaciuscula Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 319, 1885 (Spruce 1885). Radula ovalilobula K.Yamada, J. Hattori Bot. Lab. 45: 257, 1979 (Yamada 1979b). Radula oyamensis Steph., Hedwigia 23 (10): 149, 1884 (Stephani 1884a). Radula paganii Castle, J. Hattori Bot. Lab. 21: 33, 1959 (Castle 1959). 389 Radula pallens (Sw.) Nees ex Mont., Voy. Amér. Mérid. 7 (2): 71, 1839 (Montagne 1839a). Bas.: Jungermannia pallens Sw., Prodr. (Swartz): 143, 1788 (Swartz 1788). Radula pandei Udar et Dh.Kumar, Lindbergia 9 (2): 133, 1983 (Udar and Kumar 1983b). 388 Radula longispica may be conspeciic with Radula javanica. 389 Radula paganii is doubtfully distinct from Radula neotropica. World checklist of hornworts and liverworts 433 ** Radula patens K.Yamada, Cryptog. Bryol. Lichénol. 5 (1/2): 197, 1984 (Yamada 1984a). ** Radula peruviana K.Yamada, Beih. Nova Hedwigia 88: 79, 1987 (Schultze-Motel and Menzel 1987). ** Radula philippinensis K.Yamada, J. Hattori Bot. Lab. 45: 299, 1979 (Yamada 1979b). * Radula pinnulata Mitt., Fl. vit.: 410, 1871 [1873] (Mitten 1871). 390 ** Radula pseudostachya Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 319, 1885 (Spruce 1885). ** Radula punctata Steph., Hedwigia 23 (8): 135, 1884 (Stephani 1884b). ** Radula pusilla Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 320, 1885 (Spruce 1885). *** Radula rhombiloba Steph., Sp. Hepat. (Stephani) 4: 204, 1910 (Stephani 1910b). * Radula rupicola K.Yamada, J. Hattori Bot. Lab. 58: 124, 1985 (Yamada 1985b). 391 ** Radula santacruziana K.Yamada et Gradst., Trop. Bryol. 4: 67, 1991 (Yamada and Gradstein 1991). ** Radula silvestris Gottsche, Abh. Naturwiss. Vereins Bremen 7: 349, 1882 (Gottsche 1882). ** Radula sinskeana K.Yamada, J. Hattori Bot. Lab. 74: 41, 1993 (Yamada 1993). ** Radula sinuata Gottsche ex Steph., Sp. Hepat. (Stephani) 4: 161, 1910 (Stephani 1910b). * Radula socorana Gerola, Lav. Bot. Ist. Bot. Univ. Padova 12: 475, 1947 (Gerola 1947). ** Radula sonsonensis Steph., Sp. Hepat. (Stephani) 4: 201, 1910 (Stephani 1910b). ** Radula stellatogemmipara C.Gao et Y.H.Wu, Nova Hedwigia 80 (1/2): 239, 2005 (Gao and Wu 2005). ** Radula subsimplex Steph., Hedwigia 23 (8): 130, 1884 (Stephani 1884b). ** Radula subsquarrosa S.W.Arnell, Ark. Bot. (n.ser.) 4 (1): 15, 1957 (Arnell 1957b). ** Radula tabularis Steph., Hedwigia 23 (9): 131, 1884 (Stephani 1884c). ** Radula taylorii Steph., Hedwigia 23 (9): 133, 1884 (Stephani 1884c). ** Radula tectiloba Steph., Hedwigia 27 (11/12): 298, 1888 (Stephani 1888c). ** Radula tenuis K.Yamada, J. Hattori Bot. Lab. 54: 247, 1983 (Yamada 1983). ** Radula underwoodii Castle, J. Hattori Bot. Lab. 21: 37, 1959 (Castle 1959). ** Radula venezuelensis K.Yamada, Misc. Bryol. Lichenol. 9 (6): 122, 1982 (Yamada 1982b). *** Radula vieillardii Gottsche, Hedwigia 23 (10): 150, 1884 (Stephani 1884a). ** Radula visianica C.Massal., Ann. Bot. (Rome) 1 (4): 298, 1904 (Massalongo 1904). ** Radula vrieseana Sande Lac., Ann. Mus. Bot. Lugduno-Batavi 1: 305, 1864 (Sande Lacoste 1864). ** Radula wrightii Castle, J. Hattori Bot. Lab. 21: 15, 1959 (Castle 1959). 390 Radula pinnulata was accepted by Yamada (1979b) and, indirectly through comparison with Radula norrisii, by Yamada and Piippo (1989). It is conspeciic with Radula javanica according to So (2006). 391 Radula rupicola may be conspeciic with Radula vieillardii (So 2006). 434 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Radula xalapensis Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 5: 56, 1836 (Nees and Montagne 1836). Ptilidiales Schljakov ** Herzogianthaceae Stotler et Crand.-Stotl. ** Herzogianthus R.M.Schust., J. Hattori Bot. Lab. 23: 71, 1960 [1961] (Schuster 1960b). * Herzogianthus sanguineus R.M.Schust., Phytologia 56 (7): 457, 1985 (Schuster 1985c). 392 *** Herzogianthus vaginatus (Herzog) R.M.Schust., J. Hattori Bot. Lab. 23: 71, 1960 [1961] (Schuster 1960b). Bas.: Blepharostoma vaginatum Herzog, Trans. & Proc. Roy. Soc. New Zealand 65 (3): 355, 1936 (Herzog 1936b). ** Neotrichocoleaceae Inoue ** Neotrichocolea S.Hatt., J. Hattori Bot. Lab. 2: 9, 1947 [1948] (Hattori 1947b). ** Neotrichocolea bissetii (Mitt.) S.Hatt., J. Hattori Bot. Lab. 2: 10, 1947 [1948] (Hattori 1947b). Bas.: Mastigophora bissetii Mitt., Trans. Linn. Soc. London, Bot. 3 (3): 200, 1891 (Mitten 1891). ** Trichocoleopsis S.Okamura, Bot. Mag. (Tokyo) 25 (293): 159, 1911 (Okamura 1911). ** Trichocoleopsis sacculata (Mitt.) S.Okamura, Bot. Mag. (Tokyo) 25 (293): 159, 1911 (Okamura 1911). Bas.: Blepharozia sacculata Mitt., Trans. Linn. Soc. London, Bot. 3 (3): 200, 1891 (Mitten 1891). *** Ptilidiaceae H.Klinggr. by L. Söderström Ptilidiaceae was recently studied molecularly by Kreier et al. (2010) showing the occurrence of a possibly undescribed cryptic species from the Himalayas. *** Ptilidium Nees, Naturgesch. Eur. Leberm. 1: 95, 1833 (Nees 1833c). *** Ptilidium californicum (Austin) Pearson, List. Canad. Hepat.: 7, 1890 (Pearson 1890). Bas.: Lepidozia californica Austin, Bull. Torrey Bot. Club 6 (3): 19, 1875 (Austin 1875b). 392 Herzogianthus sanguineus is possibly conspeciic with Herzogianthus vaginatus. World checklist of hornworts and liverworts 435 *** Ptilidium ciliare (L.) Hampe, Prod. l. hercyn.: 76, 1836 (Hampe 1836). Bas.: Jungermannia ciliaris L., Sp. Pl. 1: 1134, 1753 (Linnaeus 1753). *** Ptilidium pulcherrimum (Weber) Vain., Meddel. Soc. Fauna Fl. Fenn. 3: 88, 1878 (Vainio 1878). Bas.: Jungermannia pulcherrima Weber, Spic. Fl. Goett.: 150, 1778 (Weber 1778). Metzgeriidae Barthol.-Began Metzgeriales Chalaud *** Aneuraceae H.Klinggr. by M. Nebel A molecular phylogeny of Aneuraceae was recently published by Preussing et al. (2010). heir study revealed Verdoorniaceae to be nested within Aneuraceae and Cryptothallus within Aneura (see also Wickett and Goinet 2008). Nomenclatural and taxonomic notes can also be found in Söderström et al. (2010a), Söderström et al. (2012a) and Nebel et al. (2013). *** Aneura Dumort., Commentat. Bot. (Dumortier): 115, 1822 (Dumortier 1822). 393 * Aneura amboinensis Steph., Bull. Herb. Boissier 7 (9): 678 (219), 1899 (Stephani 1899e). * Aneura augustae Steph., Sp. Hepat. (Stephani) 6: 430, 1923 (Stephani 1923). * Aneura bilora Colenso, Trans. & Proc. New Zealand Inst. 17: 262, 1885 (Colenso 1885). *** Aneura blasioides (Horik.) Furuki, J. Hattori Bot. Lab. 70: 311, 1991 (Furuki 1991). Bas.: Riccardia blasioides Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 1: 197, 1933 (Horikawa 1933). ** Aneura brasiliensis (Ångstr.) Steph., Hedwigia 32 (3): 137, 1893 (Stephani 1893b). Bas.: Pseudoneura brasiliensis Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 33 (7): 91, 1876 [1877] (Ångström 1876). * Aneura brevissima Steph., Sp. Hepat. (Stephani) 6: 21, 1917 (Stephani 1917a). ** Aneura cerebrata Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 94 (2): 185, 1970 (Hewson 1970b). *** Aneura crateriformis Furuki et D.G.Long, J. Bryol. 18 (2): 281, 1994 (Furuki and Long 1994). 393 Aneura includes also Acrostolia and Pseudoneura, but a few taxa have never been transferred nor synonymized. hey are listed in the section “Names in genera not currently accepted” below. he types of Aneura augustae, Aneura hunsteinii, Aneura latemultiida, Aneura ledermannii, Aneura subledermannii and Aneura subtenerrima were destroyed in B and their identity is doubtful (Grolle and Piippo 1984). 436 * ** * * ** ** ** ** * ** ** * *** * ** ** ** * ** * ** *** *** *** ** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Aneura crinita C.Massal., Bull. Soc. Bot. Ital. 1917 (8/9): 81, 1917 (Massalongo 1917). 394 Aneura crumii L.Söderstr., A.Hagborg et von Konrat, Phytotaxa 65: 43, 2012 (Söderström et al. 2012a). Nom. nov. pro Cryptothallus hirsutus H.A.Crum, Bryologist 99 (4): 438, 1996 (Crum and Bruce 1996). Aneura densa Steph., Sp. Hepat. (Stephani) 6: 24, 1917 (Stephani 1917a). Aneura denticulata Mitt. ex hurn, Timehri 5: 222, 1886 (hurn 1886). 395 Aneura eachamensis Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 94 (2): 184, 1970 (Hewson 1970b). Aneura erronea Steph., Sp. Hepat. (Stephani) 6: 20, 1917 (Stephani 1917a). Aneura eskuchei Hässel, Veröf. Geobot. Inst. ETH Stiftung Rübel Zürich 91: 294, 1986 (Hässel 1986b). Aneura gemmifera Furuki, J. Hattori Bot. Lab. 70: 309, 1991 (Furuki 1991). Aneura giangena Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 94 (2): 190, 1970 (Hewson 1970b). Aneura gibbsiana Steph., Sp. Hepat. (Stephani) 6: 28, 1917 (Stephani 1917a). Aneura glaucescens Steph., Sp. Hepat. (Stephani) 6: 28, 1917 (Stephani 1917a). Aneura goebeliana Steph., Sp. Hepat. (Stephani) 6: 28, 1917 (Stephani 1917a). Aneura hirsuta Furuki, J. Hattori Bot. Lab. 70: 317, 1991 (Furuki 1991). Aneura hunsteinii Steph., Sp. Hepat. (Stephani) 6: 431, 1923 (Stephani 1923). Aneura imbricata Colenso, Trans. & Proc. New Zealand Inst. 16: 359, 1884 (Colenso 1884). Aneura kaguaensis Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 94 (2): 190, 1970 (Hewson 1970b). Aneura keniae Gola, Mem. Reale Accad. Sci. Torino (ser. 2) 65 (1): 3, 1916 (Gola 1916). Aneura latemultiida Steph., Sp. Hepat. (Stephani) 6: 431, 1923 (Stephani 1923). Aneura latissima Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 544, 1885 (Spruce 1885). Aneura ledermannii Steph., Sp. Hepat. (Stephani) 6: 431, 1923 (Stephani 1923). Aneura macrostachya Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 550, 1885 (Spruce 1885). Aneura marianensis Furuki, Bryologist 97 (1): 87, 1994 (Furuki 1994b). Aneura maxima (Schifn.) Steph., Bull. Herb. Boissier 7 (10): 760 (270), 1899 (Stephani 1899f). Bas.: Riccardia maxima Schifn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 177, 1898 (Schifner 1898a). Aneura mirabilis (Malmb.) Wickett et Goinet, Bot. J. Linn. Soc. 156 (1): 11, 2008 (Wickett and Goinet 2008). Bas.: Cryptothallus mirabilis Malmb., Ann. Bryol. 6: 122, 1933 (von Malmborg 1933). Aneura novaecaledoniae R.M.Schust., Phytologia 56 (7): 451, 1985 (Schuster 1985c). 394 Aneura crinita (type from Europe) has neither been recognized in any recent European treatment nor synonymized. 395 Aneura denticulata is possibly conspeciic with Riccardia ciliolata (Hässel 2006b). World checklist of hornworts and liverworts 437 *** Aneura novaguineensis Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 94 (2): 189, 1970 (Hewson 1970b). * Aneura nymannii Steph., Sp. Hepat. (Stephani) 6: 35, 1917 (Stephani 1917a). ** Aneura pellucida Colenso, Trans. & Proc. New Zealand Inst. 18: 252, 1886 (Colenso 1886b). *** Aneura pinguis (L.) Dumort., Syll. Jungerm. Europ.: 86, 1831 (Dumortier 1831). Bas.: Jungermannia pinguis L., Sp. Pl. 1: 1136, 1753 (Linnaeus 1753). ** Aneura polyantha Colenso, Trans. & Proc. New Zealand Inst. 17: 262, 1885 (Colenso 1885). ** Aneura punctata Colenso, Trans. & Proc. New Zealand Inst. 18: 254, 1886 (Colenso 1886b). ** Aneura rodwayi Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 94 (2): 188, 1970 (Hewson 1970b). * Aneura roraimensis Steph., Trans. Linn. Soc. London, Bot. 6 (1): 94, 1901 (Stephani 1901e). ** Aneura rotangicola Steph., Sp. Hepat. (Stephani) 6: 432, 1923 (Stephani 1923). * Aneura serrulata Gottsche ex Steph., Sp. Hepat. (Stephani) 6: 42, 1917 (Stephani 1917a). ** Aneura sharpii Inoue et N.G.Mill., Bull. Natl. Sci. Mus. Tokyo, B 11 (3): 96, 1985 (Inoue and Miller 1985). 396 * Aneura singalangana (Schifn.) Steph., Bull. Herb. Boissier 7 (10): 751 (261), 1899 (Stephani 1899f). Bas.: Riccardia singalangana Schifn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 174, 1898 (Schifner 1898a). 397 ** Aneura subcanaliculata R.M.Schust., J. Hattori Bot. Lab. 67: 60, 1989 (Schuster 1989). * Aneura subledermannii Steph., Sp. Hepat. (Stephani) 6: 432, 1923 (Stephani 1923). * Aneura subtenerrima Steph., Sp. Hepat. (Stephani) 6: 432, 1923 (Stephani 1923). * Aneura vincentina Steph., Sp. Hepat. (Stephani) 6: 45, 1917 (Stephani 1917a). ** Lobatiriccardia (Mizut. et S.Hatt.) Furuki, J. Hattori Bot. Lab. 70: 319, 1991 (Furuki 1991). Bas.: Riccardia subg. Lobatiriccardia Mizut. et S.Hatt., J. Hattori Bot. Lab. 18: 38, 1957 (Mizutani and Hattori 1957). *** Lobatiriccardia alterniloba (Hook.f. et Taylor) Furuki, J. Hattori Bot. Lab. 70: 319, 1991 (Furuki 1991). Bas.: Jungermannia alterniloba Hook.f. et Taylor, London J. Bot. 3: 572, 1844 (Hooker and Taylor 1844d). ** Lobatiriccardia alterniloba var. gigantea (Steph.) Nebel, Phytotaxa 81 (1): 10, 2013 (Nebel et al. 2013). Bas.: Aneura gigantea Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 95, 1914 (Stephani and Watts 1914). ** Lobatiriccardia alterniloba var. robusta (Rodway) Nebel, Phytotaxa 81 (1): 10, 2013 (Nebel et al. 2013). Bas.: Aneura alterniloba f. robusta Rodway, Tasm. Bryoph.: 12, 1917 (Rodway 1917b). 396 Aneura sharpii is conspeciic with Aneura maxima in Schuster (1992b), but the taxa are not the same (D.G. Long, pers. comm.). 397 Aneura singalangana may be a form of Riccardia subexalata (Stephani 1899f ). 438 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Lobatiriccardia athertonensis (Hewson) Furuki, J. Hattori Bot. Lab. 70: 319, 1991 (Furuki 1991). Bas.: Aneura athertonensis Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 94 (2): 188, 1970 (Hewson 1970b). *** Lobatiriccardia coronopus (De Not.) Furuki, J. Hattori Bot. Lab. 100: 90, 2006 (Furuki 2006a). Bas.: Aneura coronopus De Not., Hedwigia 32 (1): 19, 1893 (Stephani 1893a). ** Lobatiriccardia coronopus subsp. australis (R.M.Schust.) Nebel, Preussing, Schäf.Verw. et D.Quandt, Taxon 59 (5): 1434, 2010 (Preussing et al. 2010). Bas.: Aneura lobata subsp. australis R.M.Schust., Phytologia 56 (7): 451, 1985 (Schuster 1985c). ** Lobatiriccardia oberwinkleri Nebel, Preussing, Schäf.-Verw. et D.Quandt, Taxon 59 (5): 1435, 2010 (Preussing et al. 2010). ** Lobatiriccardia subaquatica (R.M.Schust.) Nebel, Phytotaxa 81 (1): 10, 2013 (Nebel et al. 2013). Bas.: Aneura subaquatica R.M.Schust., Phytologia 56 (7): 450, 1985 (Schuster 1985c). ** Lobatiriccardia verdoornioides Nebel, Preussing, Schäf.-Verw. et D.Quandt, Taxon 59 (5): 1437, 2010 (Preussing et al. 2010). *** Lobatiriccardia yakusimensis (S.Hatt.) Furuki, J. Hattori Bot. Lab. 70: 321, 1991 (Furuki 1991). Bas.: Riccardia lobata var. yakusimensis S.Hatt., J. Hattori Bot. Lab. 6: 10, 1951 [1952] (Hattori 1951a). *** Lobatiriccardia yunnanensis Furuki et D.G.Long, J. Bryol. 29 (3): 161, 2007 (Furuki and Long 2007). *** Riccardia Gray, Nat. Arr. Brit. Pl. 1: 679, 1821 (Gray 1821) nom. conserv. 398 ** Riccardia pectinata var. fasciculata (Steph.) Hürl., Bauhinia 5 (4): 208, 1976 (Hürlimann 1976). Bas.: Aneura fasciculata Steph., Sp. Hepat. (Stephani) 6: 25, 1917 (Stephani 1917a). ** subg. Arceoneura Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 204, 1972 (Hässel 1972a). ** Riccardia marionensis R.M.Schust., J. Hattori Bot. Lab. 67: 65, 1989 (Schuster 1989). *** Riccardia prehensilis (Hook.f. et Taylor) C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 255, 1885 (Massalongo 1885). Bas.: Jungermannia prehensilis Hook.f. et Taylor, London J. Bot. 3: 480, 1844 (Hooker and Taylor 1844b). ** subg. Corioneura Furuki, J. Hattori Bot. Lab. 70: 394, 1991 (Furuki 1991). ** Riccardia argentolimbata Hewson et Grolle, J. Hattori Bot. Lab. 29: 70, 1966 (Grolle 1966i). *** Riccardia hattorii Furuki, J. Hattori Bot. Lab. 75: 257, 1994 (Furuki 1994a). ** subg. Hyaloneura R.M.Schust., Phytologia 56 (7): 452, 1985 (Schuster 1985c). 398 Riccardia includes Acrostolia, but a few taxa have neither been transferred nor synonymized. hey are listed under “Names in genera not currently accepted” below. World checklist of hornworts and liverworts 439 * Riccardia albomarginata (Steph.) Schifn., Consp. Hepat. Arch. Ind.: 53, 1898 (Schifner 1898b). Bas.: Aneura albomarginata Steph., Hedwigia 32 (1): 18, 1893 (Stephani 1893a). 399 ** Riccardia canaliculata (Nees) Kuntze, Revis. Gen. Pl. 2: 838, 1891 (Kuntze 1891). Bas.: Jungermannia canaliculata Nees, Enum. Pl. Crypt. Javae: 10, 1830 (Nees 1830). ** Riccardia pindensis Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 87, 1970 (Hewson 1970a). ** subg. Lophoneura Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 218, 1972 (Hässel 1972a). ** Riccardia fuegiensis C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 255, 1885 (Massalongo 1885). ** subg. Neoneura Furuki, J. Hattori Bot. Lab. 70: 385, 1991 (Furuki 1991). *** Riccardia spongiosa Furuki, J. Hattori Bot. Lab. 70: 385, 1991 (Furuki 1991). ** subg. Phycaneura R.M.Schust., J. Hattori Bot. Lab. 26: 294, 1963 (Schuster 1963b). ** Riccardia aequicellularis (Steph.) Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 79, 1970 (Hewson 1970a). Bas.: Aneura aequicellularis Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 95, 1914 (Stephani and Watts 1914). ** Riccardia asperulata R.M.Schust., J. Hattori Bot. Lab. 27: 209, 1964 (Schuster 1964a). ** *** *** *** *** *** *** *** *** ** subg. Riccardia Riccardia aeruginosa Furuki, J. Hattori Bot. Lab. 70: 345, 1991 (Furuki 1991). Riccardia arcuata Furuki, J. Hattori Bot. Lab. 70: 361, 1991 (Furuki 1991). Riccardia chamedryfolia (With.) Grolle, Trans. Brit. Bryol. Soc. 5 (4): 772, 1969 (Grolle 1969b). Bas.: Jungermannia chamedryfolia With., Bot. arr. veg. Gr. Brit. 2: 699, 1776 (Withering 1776). Riccardia cochleata (Hook.f. et Taylor) Kuntze, Revis. Gen. Pl. 2: 838, 1891 (Kuntze 1891). Bas.: Riccia cochleata Hook.f. et Taylor, London J. Bot. 4: 96, 1845 (Hooker and Taylor 1845). Riccardia eriocaula (Hook.) C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 256, 1885 (Massalongo 1885). Bas.: Jungermannia eriocaula Hook., Musci Exot. 1: tab. 72, 1818 (Hooker 1818). Riccardia lavovirens Furuki, J. Hattori Bot. Lab. 70: 333, 1991 (Furuki 1991). Riccardia fruticosa (Steph.) Furuki, Nat. Hist. Res. 5 (1): 1, 1998 (Furuki 1998). Bas.: Aneura fruticosa Steph., Sp. Hepat. (Stephani) 6: 27, 1917 (Stephani 1917a). Riccardia glauca Furuki, J. Hattori Bot. Lab. 70: 352, 1991 (Furuki 1991). Riccardia kodamae Mizut. et S.Hatt., J. Hattori Bot. Lab. 18: 57, 1957 (Mizutani and Hattori 1957). 399 Riccardia albomarginata is possibly conspeciic with Riccardia canaliculata (Söderström et al. 2010a). 440 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Riccardia latifrons (Lindb.) Lindb., Acta Soc. Sci. Fenn. 10: 513, 1875 (Lindberg 1875). Bas.: Aneura latifrons Lindb., Bot. Not. 26: 62, 1873 (Anonymous 1873). ** Riccardia latifrons subsp. arctica R.M.Schust. et Damsh., J. Hattori Bot. Lab. 62: 303, 1987 (Schuster 1987d). ** Riccardia latifrons var. miyakeana (Schifn.) Furuki, J. Hattori Bot. Lab. 70: 375, 1991 (Furuki 1991). Bas.: Riccardia miyakeana Schifn., Österr. Bot. Z. 49 (11): 388, 1899 (Schifner 1899c). ** Riccardia nagasakiensis (Steph.) S.Hatt., Bull. Tokyo Sci. Mus. 11: 164, 1944 (Hattori 1944d). Bas.: Aneura nagasakiensis Steph., Sp. Hepat. (Stephani) 6: 34, 1917 (Stephani 1917a). *** Riccardia palmata (Hedw.) Carruth., J. Bot. 3: 302, 1865 (Carruthers 1865). Bas.: Jungermannia palmata Hedw., heoria generat.: 87, 1784 (Hedwig 1784), nom. conserv. ** Riccardia planilora (Steph.) S.Hatt., Bull. Tokyo Sci. Mus. 11: 164, 1944 (Hattori 1944d). Bas.: Aneura planilora Steph., Sp. Hepat. (Stephani) 6: 38, 1917 (Stephani 1917a). ** Riccardia planilora var. aequatorialis Furuki, Nat. Hist. Res. 4 (2): 77, 1997 (Furuki 1997). ** Riccardia pseudodendroceros R.M.Schust., Phytologia 56 (7): 452, 1985 (Schuster 1985c). *** Riccardia pumila Furuki, J. Hattori Bot. Lab. 70: 361, 1991 (Furuki 1991). ** Riccardia pusilla Grolle, J. Jap. Bot. 41 (8): 231, 1966 (Grolle 1966d). Nom. nov. pro Riccardia nana Mizut. et S.Hatt., J. Hattori Bot. Lab. 18: 53, 1957 (Mizutani and Hattori 1957), nom. illeg. *** Riccardia subalpina Furuki, J. Hattori Bot. Lab. 70: 367, 1991 (Furuki 1991). ** Riccardia tamariscina (Steph.) Schifn., Consp. Hepat. Arch. Ind.: 58, 1898 (Schifner 1898b). Bas.: Aneura tamariscina Steph., Hedwigia 32 (1): 27, 1893 (Stephani 1893a). *** Riccardia vitrea Furuki, J. Hattori Bot. Lab. 70: 327, 1991 (Furuki 1991). ** sect. Alcicornia Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 17, 1972 (Hässel 1972a). ** Riccardia alcicornis (Hook.f. et Taylor) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 431, 1877 (Trevisan 1877). Bas.: Jungermannia alcicornis Hook.f. et Taylor, London J. Bot. 3: 479, 1844 (Hooker and Taylor 1844b). ** Riccardia conimitra (Steph.) A.Evans, Trans. Connecticut Acad. Arts 25 (2): 156, 1921 (Evans 1921a). Bas.: Aneura conimitra Steph., Bull. Herb. Boissier 7 (10): 749 (259), 1899 (Stephani 1899f). ** Riccardia corralensis (Steph.) A.Evans, Trans. Connecticut Acad. Arts 25 (2): 143, 1921 (Evans 1921a). Bas.: Aneura corralensis Steph., Bull. Herb. Boissier 7 (10): 742 (252), 1899 (Stephani 1899f). *** Riccardia furtiva E.A.Br. et Braggins, J. Hattori Bot. Lab. 66: 35, 1989 (Brown and Braggins 1989). World checklist of hornworts and liverworts 441 ** Riccardia fuscobrunnea (Steph.) A.Evans, Trans. Connecticut Acad. Arts 25 (2): 152, 1921 (Evans 1921a). Bas.: Aneura fuscobrunnea Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 7, 1911 (Stephani 1911b). ** Riccardia longioleata Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 30, 1972 (Hässel 1972a). *** Riccardia multicorpora E.A.Br., J. Hattori Bot. Lab. 66: 40, 1989 (Brown and Braggins 1989). * Riccardia umida E.A.Br., J. Hattori Bot. Lab. 66: 38, 1989 (Brown and Braggins 1989). ** sect. Crassantia Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 41, 1972 (Hässel 1972a). *** Riccardia aequitexta (Steph.) E.A.Br., J. Hattori Bot. Lab. 66: 78, 1989 (Brown and Braggins 1989). Bas.: Aneura aequitexta Steph., J. Linn. Soc., Bot. 29 (201): 263, 1892 (Stephani 1892b). *** Riccardia alba (Colenso) E.A.Br., J. Hattori Bot. Lab. 66: 66, 1989 (Brown and Braggins 1989). Bas.: Aneura alba Colenso, Trans. & Proc. New Zealand Inst. 16: 357, 1884 (Colenso 1884). ** Riccardia amnicola Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 92, 1972 (Hässel 1972a). ** Riccardia bipinnatiida (Colenso) Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 93, 1970 (Hewson 1970a). Bas.: Aneura bipinnatiida Colenso, Trans. & Proc. New Zealand Inst. 16: 358, 1884 (Colenso 1884). ** Riccardia calva (Schifn.) A.Evans, Trans. Connecticut Acad. Arts 25 (2): 134, 1921 (Evans 1921a). Bas.: Aneura calva Schifn., Leberm., Forschungsr. Gazelle 4 (4): 42, 1890 (Schifner 1890). *** Riccardia colensoi (Steph.) W.Martin, Trans. & Proc. Roy. Soc. New Zealand 78 (4): 499, 1950 (Martin 1950). Bas.: Aneura colensoi Steph., J. Linn. Soc., Bot. 29 (201): 264, 1892 (Stephani 1892b). *** Riccardia crassa (Schwägr.) C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 254, 1885 (Massalongo 1885). Bas.: Jungermannia crassa Schwägr., Hist. Musc. Hepat. Prodr.: 31, 1814 (Schwägrichen 1814). ** Riccardia crassicrispa (Steph.) A.Evans, Trans. Connecticut Acad. Arts 25 (2): 194, 1921 (Evans 1921a). Bas.: Aneura crassicrispa Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 6, 1911 (Stephani 1911b). ** Riccardia diderma Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 56, 1972 (Hässel 1972a). ** Riccardia diversilora A.Evans, Trans. Connecticut Acad. Arts 25 (2): 167, 1921 (Evans 1921a). ** Riccardia diversilora subsp. paucigyna R.M.Schust., J. Hattori Bot. Lab. 67: 97, 1989 (Schuster 1989). ** Riccardia duriuscula Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 82, 1972 (Hässel 1972a). 442 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Riccardia falsiloribunda Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 119, 1972 (Hässel 1972a). ** Riccardia loribunda (Steph.) A.Evans, Trans. Connecticut Acad. Arts 25 (2): 182, 1921 (Evans 1921a). Bas.: Aneura loribunda Steph., Bull. Herb. Boissier 7 (10): 749 (259), 1899 (Stephani 1899f). ** Riccardia luvigena Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 73, 1972 (Hässel 1972a). ** Riccardia georgiensis (Steph.) Hässel, Lindbergia 1 (1/2): 80, 1971 [1972] (Grolle 1971a). Bas.: Aneura georgiensis Steph., Wiss. Ergebn. Schwed. Südpolar-Exped. [1901–1903] 4 (1): 2, 1905 (Stephani 1905e). ** Riccardia georgiensis subsp. sympodea R.M.Schust., J. Hattori Bot. Lab. 67: 83, 1989 (Schuster 1989). *** Riccardia graefei (Steph.) Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 118, 1970 (Hewson 1970a). Bas.: Aneura graefei Steph., Hedwigia 32 (1): 21, 1893 (Stephani 1893a). *** Riccardia marginata (Colenso) Pearson, Univ. Calif. Publ. Bot. 10 (4): 309, 1923 (Pearson 1923). Bas.: Aneura marginata Colenso, Trans. & Proc. New Zealand Inst. 18: 253, 1886 (Colenso 1886b). *** Riccardia marginata var. paciica Furuki, J. Hattori Bot. Lab. 70: 380, 1991 (Furuki 1991). ** Riccardia mycophora A.Evans, Trans. Connecticut Acad. Arts 25 (2): 175, 1921 (Evans 1921a). ** Riccardia negeri (Steph.) A.Evans, Trans. Connecticut Acad. Arts 25 (2): 172, 1921 (Evans 1921a). Bas.: Aneura negeri Steph., Bull. Herb. Boissier 7 (10): 747 (257), 1899 (Stephani 1899f). ** Riccardia nitida (Colenso) E.A.Hodgs., Trans. Roy. Soc. New Zealand, Bot. 3 (4): 92, 1965 (Hodgson 1965). Bas.: Aneura nitida Colenso, Trans. & Proc. New Zealand Inst. 18: 253, 1886 (Colenso 1886b). ** Riccardia papillosa (C.Massal. et Steph.) Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 60, 1972 (Hässel 1972a). Bas.: Aneura papillosa C.Massal. et Steph., Atti Reale Ist. Veneto Sci. Lett. Arti 87 (2): 241, 1928 (Massalongo 1928). ** Riccardia regularis (Steph.) Kühnem., Revista Centro Estud. Doct. Ci. Nat. 1: 171, 1937 (Kühnemann 1937). Bas.: Aneura regularis Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 9, 1911 (Stephani 1911b). ** Riccardia rivularis Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 52, 1972 (Hässel 1972a). ** Riccardia saxicola Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 137, 1972 (Hässel 1972a). ** Riccardia spectabilis (Steph.) A.Evans, Trans. Connecticut Acad. Arts 25 (2): 140, 1921 (Evans 1921a). Bas.: Aneura spectabilis Steph., Bull. Herb. Boissier 7 (10): 746 (256), 1899 (Stephani 1899f). World checklist of hornworts and liverworts 443 ** Riccardia spegazziniana C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 254, 1885 (Massalongo 1885). ** Riccardia tenax (Steph.) A.Evans, Trans. Connecticut Acad. Arts 25 (2): 186, 1921 (Evans 1921a). Bas.: Aneura tenax Steph., Bull. Herb. Boissier 7 (10): 755 (265), 1899 (Stephani 1899f). ** Riccardia tenerrima (Steph.) A.Evans, Trans. Connecticut Acad. Arts 25 (2): 164, 1921 (Evans 1921a). Bas.: Aneura tenerrima Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 9, 1911 (Stephani 1911b). ** Riccardia theliophora Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 96, 1972 (Hässel 1972a). ** sect. Pallidevirida Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 142, 1972 (Hässel 1972a). ** Riccardia granulata (Steph.) A.Evans, Trans. Connecticut Acad. Arts 25 (2): 192, 1921 (Evans 1921a). Bas.: Aneura granulata Steph., Hedwigia 32 (1): 21, 1893 (Stephani 1893a). ** Riccardia intercellula E.A.Br., J. Hattori Bot. Lab. 66: 58, 1989 (Brown and Braggins 1989). ** Riccardia opuntiiformis S.W.Arnell, Arch. Soc. Zool. Bot. Fenn. “Vanamo” 9: 52, 1954 (Arnell 1954b). ** Riccardia pallidevirens (Steph.) A.Evans, Trans. Connecticut Acad. Arts 25 (2): 189, 1921 (Evans 1921a). Bas.: Aneura pallidevirens Steph., Bull. Herb. Boissier 7 (10): 762 (272), 1899 (Stephani 1899f). *** Riccardia pennata E.A.Br., J. Hattori Bot. Lab. 66: 53, 1989 (Brown and Braggins 1989). *** Riccardia perspicua E.A.Br., J. Hattori Bot. Lab. 66: 55, 1989 (Brown and Braggins 1989). ** Riccardia xylophila Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 156, 1972 (Hässel 1972a). ** sect. Riccardia *** Riccardia australis (Lehm.) Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 110, 1970 (Hewson 1970a). Bas.: Sarcomitrium australe Lehm., Nov. Stirp. Pug. 10: 19, 1857 (Lehmann 1857). ** Riccardia autoica (Steph.) A.Evans, Trans. Connecticut Acad. Arts 25 (2): 159, 1921 (Evans 1921a). Bas.: Aneura autoica Steph., Bull. Herb. Boissier 7 (9): 691 (232), 1899 (Stephani 1899e). ** Riccardia breviala E.A.Br., J. Hattori Bot. Lab. 66: 46, 1989 (Brown and Braggins 1989). ** Riccardia breviramosa (Steph.) A.Evans, Nat. Hist. Juan Fernandez (Botany) 2 (20): 559, 1930 (Evans 1930a). Bas.: Aneura breviramosa Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 6, 1911 (Stephani 1911b). 444 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Riccardia ilicina (Colenso) E.A.Hodgs., Rec. Domin. Mus. 4 (11): 129, 1962 (Hodgson 1962a). Bas.: Aneura ilicina Colenso, Trans. & Proc. New Zealand Inst. 16: 358, 1884 (Colenso 1884). ** Riccardia leptostachya A.Evans, Nat. Hist. Juan Fernandez (Botany) 2 (20): 570, 1930 (Evans 1930a). ** Riccardia lobulata (Colenso) E.A.Hodgs., Trans. Roy. Soc. New Zealand, Bot. 3 (4): 92, 1965 (Hodgson 1965). Bas.: Zoopsis lobulata Colenso, Trans. & Proc. New Zealand Inst. 18: 250, 1886 (Colenso 1886b). ** Riccardia mejlandii S.W.Arnell, Results Norweg. Sci. Exped. Tristan da Cunha 42: 30, 1958 (Arnell 1958b). *** Riccardia multiida (L.) Gray, Nat. Arr. Brit. Pl. 1: 684, 1821 (Gray 1821). Bas.: Jungermannia multiida L., Sp. Pl. 1: 1136, 1753 (Linnaeus 1753). *** Riccardia multiida subsp. decrescens (Steph.) Furuki, J. Hattori Bot. Lab. 70: 341, 1991 (Furuki 1991). Bas.: Aneura decrescens Steph., Bull. Herb. Boissier 7 (9): 686 (227), 1899 (Stephani 1899e). ** Riccardia multiida subsp. synoica R.M.Schust., J. Hattori Bot. Lab. 62: 319, 1987 (Schuster 1987d). ** Riccardia multioleata Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 195, 1972 (Hässel 1972a). *** Riccardia papulosa (Steph.) E.A.Br., J. Hattori Bot. Lab. 66: 49, 1989 (Brown and Braggins 1989). Bas.: Aneura papulosa Steph., Hedwigia 32 (1): 25, 1893 (Stephani 1893a). ** Riccardia patens Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 165, 1972 (Hässel 1972a). ** Riccardia polyclada (Mitt.) Hässel, Bryologist 109 (1): 34, 2006 (Hässel 2006b). Bas.: Aneura polyclada Mitt., Trans. Linn. Soc. London, Bot. 2 (13): 297, 1887 (Mitten 1887). ** Riccardia statensis Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 177, 1972 (Hässel 1972a). ** Riccardia thaxteri A.Evans, Trans. Connecticut Acad. Arts 25 (2): 126, 1921 (Evans 1921a). ** Riccardia tristaniana S.W.Arnell, Results Norweg. Sci. Exped. Tristan da Cunha 42: 31, 1958 (Arnell 1958b). ** subg. Spinella (Schifn.) Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 214, 1972 (Hässel 1972a). Bas.: Spinella Schifn., Leberm., Forschungsr. Gazelle 4 (4): 41, 1890 (Schifner 1890). ** Riccardia spinulifera C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 254, 1885 (Massalongo 1885). ** subg. hornoneura Furuki, J. Hattori Bot. Lab. 70: 382, 1991 (Furuki 1991). World checklist of hornworts and liverworts 445 * Riccardia baumannii Hürl., Bauhinia 5 (4): 203, 1976 (Hürlimann 1976). 400 ** Riccardia deguchii Furuki et K.T.Yong, Hikobia 16 (3): 285, 2013 (Furuki et al. 2013). *** Riccardia grossitexta (Steph.) Furuki, J. Hattori Bot. Lab. 70: 382, 1991 (Furuki 1991). Bas.: Aneura grossitexta Steph., Sp. Hepat. (Stephani) 6: 29, 1917 (Stephani 1917a). ** Riccardia inconspicua (Steph.) Reeb et Bardat, Cryptog. Bryol. 35 (1): 61, 2014 (Reeb and Bardat 2014). Bas.: Aneura inconspicua Steph., Hedwigia 32 (1): 23, 1893 (Stephani 1893a). ** subg. Trichothallia Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 224, 1972 (Hässel 1972a). ** Riccardia hyalitricha Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 225, 1972 (Hässel 1972a). Incertae sedis ** Riccardia aberrans (Steph.) Gradst., J. Hattori Bot. Lab. 45: 129, 1979 (Gradstein and Hekking 1979). Bas.: Aneura aberrans Steph., Hedwigia 32 (1): 18, 1893 (Stephani 1893a). ** Riccardia agumana Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 82, 1970 (Hewson 1970a). ** Riccardia algoides (Taylor) Meenks, J. Hattori Bot. Lab. 62: 168, 1987 (Meenks 1987). Bas.: Metzgeria algoides Taylor, London J. Bot. 5: 410, 1846 (Taylor 1846b). *** Riccardia amazonica (Spruce) Schifn. ex Gradst. et Hekking, J. Hattori Bot. Lab. 45: 129, 1979 (Gradstein and Hekking 1979). Bas.: Aneura amazonica Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 545, 1885 (Spruce 1885). *** Riccardia andina (Spruce) Herzog, Svensk Bot. Tidskr. 46 (1): 65, 1952 (Herzog 1952e). Bas.: Aneura andina Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 548, 1885 (Spruce 1885). ** Riccardia angustata Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 126, 1934 (Horikawa 1934). ** Riccardia angustealata (Steph.) Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 111, 1970 (Hewson 1970a). Bas.: Aneura angustealata Steph., Sp. Hepat. (Stephani) 6: 20, 1917 (Stephani 1917a). ** Riccardia angustissima (Steph.) H.A.Mill., Phytologia 47 (4): 323, 1981 (Miller 1981). Bas.: Aneura angustissima Steph., Sp. Hepat. (Stephani) 6: 20, 1917 (Stephani 1917a). ** Riccardia aspera (Steph.) Grolle, J. Hattori Bot. Lab. 30: 117, 1967 (Grolle 1967b). Bas.: Aneura aspera Steph., Sp. Hepat. (Stephani) 6: 21, 1917 (Stephani 1917a). 400 Riccardia baumannii is possibly conspeciic with Riccardia grossitexta or Riccardia tenuicostata (Furuki 1991, Söderström et al. 2010a). 446 * ** ** ** ** ** ** ** ** ** ** *** ** ** ** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Riccardia baldwinii (Steph.) H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 527, 1963 (Miller 1963). Bas.: Aneura baldwinii Steph., Bull. Herb. Boissier 7 (10): 743 (253), 1899 (Stephani 1899f). 401 Riccardia barbilora (Steph.) Piippo, J. Hattori Bot. Lab. 68: 134, 1990 (Piippo 1990). Bas.: Aneura barbilora Steph., Mém. Soc. Nat. Sci. Nat. Math. Cherbourg 29: 209, 1894 (Stephani 1894b). Riccardia bogotensis (Gottsche) Pagán, Bryologist 42 (1): 6, 1939 (Pagán 1939a). Bas.: Pseudoneura bogotensis Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 184, 1864 (Gottsche 1864). Riccardia boliviensis (Steph.) Meenks, J. Hattori Bot. Lab. 62: 170, 1987 (Meenks 1987). Bas.: Aneura boliviensis Steph., Biblioth. Bot. 87 (2): 176, 1916 (Stephani 1916a). Riccardia bongeriana Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 101, 1970 (Hewson 1970a). Riccardia brunnea (Steph.) S.Hatt., Bull. Tokyo Sci. Mus. 11: 164, 1944 (Hattori 1944d). Bas.: Aneura brunnea Steph., Sp. Hepat. (Stephani) 6: 21, 1917 (Stephani 1917a). Riccardia calcarea (Steph.) Meenks, J. Hattori Bot. Lab. 62: 170, 1987 (Meenks 1987). Bas.: Aneura calcarea Steph., Bull. Herb. Boissier 7 (10): 756 (266), 1899 (Stephani 1899f). Riccardia capillacea (Steph.) Meenks et C.De Jong, Cryptog. Bryol. Lichénol. 6 (1): 6, 1985 (Meenks and de Jong 1985). Bas.: Aneura capillacea Steph., Biblioth. Bot. 87 (2): 174, 1916 (Stephani 1916a). Riccardia capillacea var. dentata Meenks, J. Hattori Bot. Lab. 62: 170, 1987 (Meenks 1987). Riccardia cardotii (Steph.) Pandé et S.C.Srivast., Biol. Mem. 1 (1/2): 131, 1976 (Srivastava and Udar 1976). Bas.: Aneura cardotii Steph., Bull. Soc. Roy. Bot. Belgique, Mém. 41 (1): 118, 1904 [1905] (Stephani 1904d). Riccardia cataractarum (Spruce) Schifn., Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 18, 1964 (Schifner and Arnell 1964). Bas.: Aneura cataractarum Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: cxcv, 1889 [1890] (Spruce 1889). Riccardia cervicornis (Spruce) Herzog ex Gradst. et Hekking, J. Hattori Bot. Lab. 45: 129, 1979 (Gradstein and Hekking 1979). Bas.: Aneura cervicornis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 550, 1885 (Spruce 1885). Riccardia changbaishanensis C.Gao, Fl. Hepat. Chin. Boreali-Orient.: 209, 1981 (Gao and Chang 1981). Riccardia chinensis C.Gao, Fl. Hepat. Chin. Boreali-Orient.: 209, 1981 (Gao and Chang 1981). Riccardia ciliolata (Spruce) Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 1: 198, 1933 (Horikawa 1933). Bas.: Aneura ciliolata Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 547, 1885 (Spruce 1885). 401 Riccardia baldwinii is possibly conspeciic with Riccardia laccida (Miller et al. 1983). World checklist of hornworts and liverworts 447 ** Riccardia columbica (Steph.) Hässel ex Gradst. et Hekking, J. Hattori Bot. Lab. 45: 130, 1979 (Gradstein and Hekking 1979). Bas.: Aneura columbica Steph., Sp. Hepat. (Stephani) 6: 22, 1917 (Stephani 1917a). ** Riccardia comata (Steph.) H.A.Mill., Phytologia 47 (4): 323, 1981 (Miller 1981). Bas.: Aneura comata Steph., Sp. Hepat. (Stephani) 6: 22, 1917 (Stephani 1917a). ** Riccardia compacta (Steph.) S.W.Arnell, Bot. Not. 105: 141, 1952 (Arnell 1952b). Bas.: Aneura compacta Steph., Hedwigia 32 (1): 19, 1893 (Stephani 1893a). ** Riccardia comptonii (Pearson) H.A.Mill., Phytologia 47 (4): 323, 1981 (Miller 1981). Bas.: Aneura comptonii Pearson, J. Linn. Soc., Bot. 46 (305): 17, 1922 (Pearson 1922b). ** Riccardia costata (Steph.) Hürl., Bauhinia 5 (4): 206, 1976 (Hürlimann 1976). Bas.: Aneura costata Steph., Sp. Hepat. (Stephani) 6: 23, 1917 (Stephani 1917a). *** Riccardia crassicaulis (Steph.) Meenks et C.De Jong, Cryptog. Bryol. Lichénol. 6 (1): 8, 1985 (Meenks and de Jong 1985). Bas.: Aneura crassicaulis Steph., Biblioth. Bot. 87 (2): 174, 1916 (Stephani 1916a). ** Riccardia crassiretis Schifn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 173, 1898 (Schifner 1898a). ** Riccardia crenulata Schifn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 173, 1898 (Schifner 1898a). ** Riccardia crenuliformis R.M.Schust., J. Hattori Bot. Lab. 67: 72, 1989 (Schuster 1989). * Riccardia decolyana Schifn., J. Indian Bot. Soc. 38 (4): 538, 1959 [1960] (Schifner et al. 1959). 402 ** Riccardia densiramea (Steph.) S.Hatt., Bull. Tokyo Sci. Mus. 11: 164, 1944 (Hattori 1944d). Bas.: Aneura densiramea Steph., Sp. Hepat. (Stephani) 6: 24, 1917 (Stephani 1917a). ** Riccardia devexa Schifn., Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 12, 1964 (Schifner and Arnell 1964). ** Riccardia diablotina (Spruce) Pagán, Bryologist 45 (4): 80, 1942 (Pagán 1942b). Bas.: Aneura diablotina Spruce, J. Linn. Soc., Bot. 30 (210): 366, 1895 (Gepp 1895b). *** Riccardia digitiloba (Spruce) Pagán, Bryologist 42 (1): 6, 1939 (Pagán 1939a). Bas.: Aneura digitiloba Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: cci, 1889 [1890] (Spruce 1889). ** Riccardia dilatata (Spruce) Schäf.-Verw. et Pócs, Cryptog. Bryol. 31 (4): 389, 2010 (Schäfer-Verwimp 2010). Bas.: Aneura dilatata Spruce, J. Linn. Soc., Bot. 30 (210): 368, 1895 (Gepp 1895b). * Riccardia diminuta Schifn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 170, 1898 (Schifner 1898a). 403 * Riccardia diminuta var. thermarum Schifn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 172, 1898 (Schifner 1898a). 402 Riccardia decolyana is possibly conspeciic with Riccardia multiida (Srivastava and Udar 1976). 403 Riccardia diminuta is possibly conspeciic with Riccardia elata (Söderström et al. 2010a). 448 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Riccardia distans (Spruce) Pagán, Bryologist 45 (4): 80, 1942 (Pagán 1942b). Bas.: Aneura distans Spruce, J. Linn. Soc., Bot. 30 (210): 367, 1895 (Gepp 1895b). ** Riccardia elata (Steph.) Schifn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 169, 1898 (Schifner 1898a). Bas.: Aneura elata Steph., Hedwigia 32 (1): 19, 1893 (Stephani 1893a). * Riccardia elata var. laccida Schifn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 170, 1898 (Schifner 1898a). * Riccardia elata var. intercedens Schifn., Denkschr. Kaiserl. Akad. Wiss., Math.Naturwiss. Kl. 67: 170, 1898 (Schifner 1898a). ** Riccardia elegans (Steph.) Hürl., Bauhinia 5 (4): 196, 1976 (Hürlimann 1976). Bas.: Aneura elegans Steph., Sp. Hepat. (Stephani) 6: 25, 1917 (Stephani 1917a). ** Riccardia elisabethae houvenot et Reeb, Telopea 17: 229, 2014 (houvenot and Reeb 2014). *** Riccardia emarginata (Steph.) K.G.Hell, Bol. Fac. Filos. Univ. São Paulo, Bot. 25: 100, 1969 (Hell 1969). Bas.: Aneura emarginata Steph., Hedwigia 32 (1): 20, 1893 (Stephani 1893a). ** Riccardia erosa (Steph.) E.W.Jones, Trans. Brit. Bryol. Soc. 3 (1): 83, 1956 (Jones 1956). Bas.: Aneura erosa Steph., Hedwigia 30 (6): 269, 1891 (Stephani 1891c). ** Riccardia fastigiata (Lehm.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 431, 1877 (Trevisan 1877). Bas.: Jungermannia fastigiata Lehm., Linnaea 4: 370, 1829 (Lehmann 1829). ** Riccardia fendleri (Steph.) Pagán, Bryologist 45 (4): 80, 1942 (Pagán 1942b). Bas.: Aneura fendleri Steph., Hedwigia 32 (1): 20, 1893 (Stephani 1893a). ** Riccardia laccida (Steph.) S.Hatt., Bull. Tokyo Sci. Mus. 11: 164, 1944 (Hattori 1944d). Bas.: Aneura laccida Steph., Sp. Hepat. (Stephani) 6: 26, 1917 (Stephani 1917a). ** Riccardia laccidissima Schifn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 167, 1898 (Schifner 1898a). ** Riccardia lagellaris (A.Gepp) H.A.Mill., Phytologia 47 (4): 323, 1981 (Miller 1981). Bas.: Aneura lagellaris A.Gepp, J. Linn. Soc., Bot. 39 (270): 194, 1909 (Gibbs 1909). ** Riccardia lagellifrons C.Gao, Fl. Hepat. Chin. Boreali-Orient.: 209, 1981 (Gao and Chang 1981). ** Riccardia leischeri (Steph.) H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 527, 1963 (Miller 1963). Bas.: Aneura leischeri Steph., Sp. Hepat. (Stephani) 6: 26, 1917 (Stephani 1917a). ** Riccardia foliacea Meenks et C.De Jong, Cryptog. Bryol. Lichénol. 6 (1): 8, 1985 (Meenks and de Jong 1985). ** Riccardia formosensis (Steph.) Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 125, 1934 (Horikawa 1934). Bas.: Aneura formosensis Steph., Sp. Hepat. (Stephani) 6: 27, 1917 (Stephani 1917a). *** Riccardia fucoidea (Sw.) C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 256, 1885 (Massalongo 1885). Bas.: Jungermannia fucoidea Sw., Prodr. (Swartz): 145, 1788 (Swartz 1788). World checklist of hornworts and liverworts 449 ** Riccardia geniana Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 76, 1970 (Hewson 1970a). *** Riccardia glaziovii (Spruce) Meenks, J. Hattori Bot. Lab. 62: 173, 1987 (Meenks 1987). Bas.: Aneura glaziovii Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: cci, 1889 [1890] (Spruce 1889). ** Riccardia gogolensis (Steph.) Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 102, 1970 (Hewson 1970a). Bas.: Aneura gogolensis Steph., Bull. Herb. Boissier 7 (9): 689 (230), 1899 (Stephani 1899e). ** Riccardia gracilis (Steph.) R.M.Schust., J. Hattori Bot. Lab. 26: 295, 1963 (Schuster 1963b). Bas.: Aneura gracilis Steph., Bull. Herb. Boissier 7 (10): 752 (262), 1899 (Stephani 1899f). ** Riccardia grandilora (Steph.) H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 528, 1963 (Miller 1963). Bas.: Aneura grandilora Steph., Sp. Hepat. (Stephani) 6: 29, 1917 (Stephani 1917a). *** Riccardia grollei Furuki, Haussknechtia, Beih. 9: 139, 1999 (Furuki 1999). ** Riccardia grossidens (Steph.) Pagán, Bryologist 45 (4): 80, 1942 (Pagán 1942b). Bas.: Aneura grossidens Steph., Hedwigia 32 (1): 23, 1893 (Stephani 1893a). ** Riccardia gunniana (Steph.) H.A.Mill., Phytologia 47 (4): 323, 1981 (Miller 1981). Bas.: Aneura gunniana Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 96, 1914 (Stephani and Watts 1914). ** Riccardia hamatilora (Steph.) H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 526, 1963 (Miller 1963). Bas.: Aneura hamatilora Steph., Bull. Herb. Boissier 5 (10): 844, 1897 (Stephani 1897c). *** Riccardia hans-meyeri (Steph.) Meenks et C.De Jong, Cryptog. Bryol. Lichénol. 6 (1): 12, 1985 (Meenks and de Jong 1985). Bas.: Aneura hans-meyeri Steph., Sp. Hepat. (Stephani) 6: 29, 1917 (Stephani 1917a). ** Riccardia hans-meyeri var. dentata Meenks, J. Hattori Bot. Lab. 62: 173, 1987 (Meenks 1987). ** Riccardia hawaica (Steph.) H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 528, 1963 (Miller 1963). Bas.: Aneura hawaica Steph., Sp. Hepat. (Stephani) 6: 30, 1917 (Stephani 1917a). ** Riccardia hebridensis (Steph.) H.A.Mill., Phytologia 47 (4): 323, 1981 (Miller 1981). Bas.: Aneura hebridensis Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 96, 1914 (Stephani and Watts 1914). ** Riccardia herzogiana (Steph.) Meenks et C.De Jong, Cryptog. Bryol. Lichénol. 6 (1): 12, 1985 (Meenks and de Jong 1985). Bas.: Aneura herzogiana Steph., Biblioth. Bot. 87 (2): 175, 1916 (Stephani 1916a). ** Riccardia heteroclada Schifn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 175, 1898 (Schifner 1898a). * Riccardia hirtilora (Steph.) Schifn., Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 19, 1964 (Schifner and Arnell 1964). Bas.: Aneura hirtilora Steph., Arch. Mus. Nac. Rio de Janeiro 13: 116, 1905 (Stephani 1905c). 404 404 Riccardia hirtilora is possibly conspeciic with Riccardia emarginata (Gradstein and Costa 2003). 450 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Riccardia humilis (Gottsche) O.Yano, J. Hattori Bot. Lab. 56: 530, 1984 (Yano 1984). Bas.: Pseudoneura humilis Gottsche, Mexik. Leverm.: 260, 1863 (Gottsche 1863). ** Riccardia hyalina (Steph.) H.A.Mill., Phytologia 47 (4): 324, 1981 (Miller 1981). Bas.: Aneura hyalina Steph., Sp. Hepat. (Stephani) 6: 31, 1917 (Stephani 1917a). ** Riccardia hydra Hürl., Bauhinia 5 (4): 210, 1976 (Hürlimann 1976). ** Riccardia hymenophylloides Schifn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 175, 1898 (Schifner 1898a). * Riccardia hymenophylloides var. laccida Schifn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 176, 1898 (Schifner 1898a). *** Riccardia hymenophytoides (Spruce) Meenks, Beih. Nova Hedwigia 88: 101, 1987 (Schultze-Motel and Menzel 1987). Bas.: Aneura hymenophytoides Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 549, 1885 (Spruce 1885). ** Riccardia hypipamensis Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 97, 1970 (Hewson 1970a). ** Riccardia ibana Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 78, 1970 (Hewson 1970a). *** Riccardia incurvata Lindb., Helsingf. Dagbl. 1878 (315, 18 Nov.): 2, 1878 (Lindberg 1878). ** Riccardia innovans (Steph.) Pagán, Bryologist 45 (4): 80, 1942 (Pagán 1942b). Bas.: Aneura innovans Steph., Symb. Antill. 2: 470, 1901 (Stephani 1901f). ** Riccardia insularis Schifn., Deutsche Südpolar-Exped. 1901-1903, 8 (bot.) 1: 66, 1906 (Schifner 1906a). ** Riccardia intricata (Steph.) H.A.Mill., Phytologia 47 (4): 324, 1981 (Miller 1981). Bas.: Aneura intricata Steph., Bot. Jahrb. Syst. 23 (1/2, 3): 301, 1896 (Stephani 1896a). ** Riccardia jackii Schifn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 165, 1898 (Schifner 1898a). * Riccardia jackii var. densa Schifn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 165, 1898 (Schifner 1898a). ** Riccardia judithae Meenks et C.De Jong, Cryptog. Bryol. Lichénol. 6 (1): 12, 1985 (Meenks and de Jong 1985). *** Riccardia jugata R.M.Schust., J. Hattori Bot. Lab. 62: 305, 1987 (Schuster 1987d). * Riccardia karstenii (Steph.) Schifn., Denkschr. Kaiserl. Akad. Wiss., Math.Naturwiss. Kl. 67: 177, 1898 (Schifner 1898a). Bas.: Aneura karstenii Steph., Hedwigia 32 (1): 23, 1893 (Stephani 1893a). ** Riccardia laticostata (Spruce) Schifn., Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 12, 1964 (Schifner and Arnell 1964). Bas.: Aneura laticostata Spruce, J. Linn. Soc., Bot. 30 (210): 367, 1895 (Gepp 1895b). ** Riccardia latifrondoides Schifn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 168, 1898 (Schifner 1898a). ** Riccardia lepidomitra (Spruce) Gradst., J. Hattori Bot. Lab. 45: 130, 1979 (Gradstein and Hekking 1979). Bas.: Aneura lepidomitra Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 549, 1885 (Spruce 1885). World checklist of hornworts and liverworts 451 ** Riccardia leptophylla (Spruce) Herzog, Svensk Bot. Tidskr. 46 (1): 65, 1952 (Herzog 1952e). Bas.: Aneura leptophylla Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 544, 1885 (Spruce 1885). ** Riccardia leptothallus R.M.Schust., J. Hattori Bot. Lab. 67: 87, 1989 (Schuster 1989). ** Riccardia levieri Schifn., Österr. Bot. Z. 49 (4): 130, 1899 (Schifner 1899b). ** Riccardia lichenoides (Steph.) H.A.Mill., Phytologia 47 (4): 324, 1981 (Miller 1981). Bas.: Aneura lichenoides Steph., Bot. Jahrb. Syst. 23 (1/2, 3): 301, 1896 (Stephani 1896a). ** Riccardia ligulata (Steph.) Pócs et Schäf.-Verw., Cryptog. Bryol. 31 (4): 390, 2010 (Schäfer-Verwimp 2010). Bas.: Aneura ligulata Steph., Sp. Hepat. (Stephani) 6: 32, 1917 (Stephani 1917a). ** Riccardia lilliena (Steph.) H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 527, 1963 (Miller 1963). Bas.: Aneura lilliena Steph., Sp. Hepat. (Stephani) 6: 33, 1917 (Stephani 1917a). ** Riccardia limbata (Steph.) E.W.Jones, Trans. Brit. Bryol. Soc. 3 (1): 79, 1956 (Jones 1956). Bas.: Aneura limbata Steph., Hedwigia 30 (5): 203, 1891 (Stephani 1891a). ** Riccardia loefgrenii Schifn., Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 14, 1964 (Schifner and Arnell 1964). ** Riccardia longilora (Steph.) Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 88, 1970 (Hewson 1970a). Bas.: Aneura longilora Steph., Bull. Herb. Boissier 7 (10): 746 (256), 1899 (Stephani 1899f). ** Riccardia longispica (Steph.) Pearson, Forh. Vidensk.-Selsk. Kristiania 1892 (14): 4, 1893 (Pearson 1893). Bas.: Aneura longispica Steph., Bot. Gaz. 15 (11): 281, 1890 (Stephani 1890c). ** Riccardia loriana (Steph.) H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 528, 1963 (Miller 1963). Bas.: Aneura loriana Steph., Bull. Herb. Boissier 7 (10): 733 (243), 1899 (Stephani 1899f). ** Riccardia macdonaldiana Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 93, 1970 (Hewson 1970a). ** Riccardia macrantha (Pearson) H.A.Mill., Phytologia 47 (4): 324, 1981 (Miller 1981). Bas.: Aneura macrantha Pearson, J. Linn. Soc., Bot. 46 (305): 17, 1922 (Pearson 1922b). *** Riccardia magnicellularis Furuki, J. Hattori Bot. Lab. 100: 93, 2006 (Furuki 2006a). *** Riccardia metzgeriiformis (Steph.) R.M.Schust., J. Hattori Bot. Lab. 26: 295, 1963 (Schuster 1963b). Bas.: Aneura metzgeriiformis Steph., Bull. Herb. Boissier 7 (10): 753 (263), 1899 (Stephani 1899f). ** Riccardia microscopica (Nees) Kuntze, Revis. Gen. Pl. 2: 838, 1891 (Kuntze 1891). Bas.: Aneura microscopica Nees, Syn. Hepat. 4: 500, 1846 (Gottsche et al. 1846). ** Riccardia minuta (Steph.) W.Martin, Trans. & Proc. Roy. Soc. New Zealand 78 (4): 499, 1950 (Martin 1950). Bas.: Aneura minuta Steph., Sp. Hepat. (Stephani) 6: 34, 1917 (Stephani 1917a). 452 * ** ** ** ** ** ** ** ** ** ** *** ** * ** ** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Riccardia multiidoides Schifn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 166, 1898 (Schifner 1898a). 405 Riccardia multispica (Steph.) S.Hatt., Bull. Tokyo Sci. Mus. 11: 164, 1944 (Hattori 1944d). Bas.: Aneura multispica Steph., Sp. Hepat. (Stephani) 6: 34, 1917 (Stephani 1917a). Riccardia nadeaudii (Steph.) Hürl., Bauhinia 5 (4): 201, 1976 (Hürlimann 1976). Bas.: Aneura nadeaudii Steph., Bull. Herb. Boissier 7 (10): 750 (260), 1899 (Stephani 1899f). Riccardia newellana (Steph.) H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 528, 1963 (Miller 1963). Bas.: Aneura newellana Steph., Sp. Hepat. (Stephani) 6: 35, 1917 (Stephani 1917a). Riccardia nigra (Steph.) H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 527, 1963 (Miller 1963). Bas.: Aneura nigra Steph., Sp. Hepat. (Stephani) 6: 35, 1917 (Stephani 1917a). Riccardia nobilis (Steph.) Schifn., Consp. Hepat. Arch. Ind.: 56, 1898 (Schifner 1898b). Bas.: Aneura nobilis Steph., Hedwigia 32 (1): 24, 1893 (Stephani 1893a). Riccardia novo-amstelodamensis Schifn., Deutsche Südpolar-Exped. 1901-1903, 8 (bot.) 1: 65, 1906 (Schifner 1906a). Riccardia nudilora (Steph.) Grolle, Bryophyt. Biblioth. 48: 130, 1995 (Grolle 1995). Bas.: Aneura nudilora Steph., Bot. Gaz. 15 (11): 282, 1890 (Stephani 1890c). Riccardia obtusa S.W.Arnell, Bot. Not. 105: 142, 1952 (Arnell 1952b). Riccardia obtusifrons (Steph.) H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 527, 1963 (Miller 1963). Bas.: Aneura obtusifrons Steph., Sp. Hepat. (Stephani) 6: 36, 1917 (Stephani 1917a). Riccardia omkaliensis Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 97, 1970 (Hewson 1970a). Riccardia pallida (Spruce) Meenks et C.De Jong, Cryptog. Bryol. Lichénol. 6 (1): 15, 1985 (Meenks and de Jong 1985). Bas.: Aneura pallida Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 547, 1885 (Spruce 1885). Riccardia palmatiida (Steph.) H.A.Mill., Phytologia 47 (4): 324, 1981 (Miller 1981). Bas.: Aneura palmatiida Steph., Sp. Hepat. (Stephani) 6: 36, 1917 (Stephani 1917a). Riccardia palmatiformis Schifn., J. Indian Bot. Soc. 38 (4): 538, 1959 [1960] (Schifner et al. 1959). 406 Riccardia papillata (Gottsche) Hässel ex Gradst. et Hekking, J. Hattori Bot. Lab. 45: 130, 1979 (Gradstein and Hekking 1979). Bas.: Pseudoneura papillata Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 185, 1864 (Gottsche 1864). Riccardia paramorum Meenks, J. Hattori Bot. Lab. 62: 176, 1987 (Meenks 1987). 405 Riccardia multiidoides is conspeciic with Riccardia multiida in Meijer (1959), but that was not based on examination of type material (Söderström et al. 2010a). 406 Riccardia palmatiformis was treated as a doubtful taxon with some ainity with Riccardia levieri by Srivastava and Udar (1976). World checklist of hornworts and liverworts 453 *** Riccardia parasitans (Steph.) Meenks et C.De Jong, Cryptog. Bryol. Lichénol. 6 (1): 17, 1985 (Meenks and de Jong 1985). Bas.: Aneura parasitans Steph., Biblioth. Bot. 87 (2): 175, 1916 (Stephani 1916a). ** Riccardia parvula Schifn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 172, 1898 (Schifner 1898a). ** Riccardia pauciramea (Steph.) H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 527, 1963 (Miller 1963). Bas.: Aneura pauciramea Steph., Bull. Herb. Boissier 5 (10): 845, 1897 (Stephani 1897c). ** Riccardia paulensis Schifn., Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 16, 1964 (Schifner and Arnell 1964). ** Riccardia pectinata (Austin) H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 526, 1963 (Miller 1963). Bas.: Aneura pectinata Austin, Bull. Torrey Bot. Club 5 (3): 15, 1874 (Austin 1874). ** Riccardia pellucida Piippo, J. Hattori Bot. Lab. 68: 134, 1990 (Piippo 1990). Nom. nov. pro Aneura pellucida Steph., Sp. Hepat. (Stephani) 6: 37, 1917 (Stephani 1917a), nom. illeg. ** Riccardia pembaiensis (Steph.) Hürl., Bauhinia 5 (4): 205, 1976 (Hürlimann 1976). Bas.: Aneura pembaiensis Steph., Sp. Hepat. (Stephani) 6: 37, 1917 (Stephani 1917a). ** Riccardia pengagensis Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 108, 1970 (Hewson 1970a). ** Riccardia perssonii S.C.Srivast. et Udar, Lindbergia 4 (1/2): 127, 1977 (Srivastava and Udar 1977). *** Riccardia philippinensis Furuki, J. Hattori Bot. Lab. 100: 94, 2006 (Furuki 2006a). ** Riccardia phleganiana Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 80, 1970 (Hewson 1970a). ** Riccardia plana (Steph.) Hürl., Bauhinia 5 (4): 205, 1976 (Hürlimann 1976). Bas.: Aneura plana Steph., Sp. Hepat. (Stephani) 6: 38, 1917 (Stephani 1917a). * Riccardia plana var. minor (Pearson) Hürl. ex H.A.Mill., H.Whittier et B.Whittier, Bryophyt. Biblioth. 25: 303, 1983 (Miller et al. 1983). Bas.: Aneura plana var. minor Pearson, J. Linn. Soc., Bot. 46 (305): 16, 1922 (Pearson 1922b). ** Riccardia planifrons (Spruce) Pagán, Bryologist 45 (4): 80, 1942 (Pagán 1942b). Bas.: Aneura planifrons Spruce, J. Linn. Soc., Bot. 30 (210): 368, 1895 (Gepp 1895b). *** Riccardia plumiformis (Spruce) Hässel ex Meenks, Beih. Nova Hedwigia 88: 101, 1987 (Schultze-Motel and Menzel 1987). Bas.: Aneura plumiformis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 548, 1885 (Spruce 1885). ** Riccardia plumosa (Mitt.) E.O.Campb., J. Roy. Soc. New Zealand 1 (1): 24, 1971 (Campbell 1971). Bas.: Sarcomitrium plumosum Mitt., Bonplandia 10 (2): 19, 1862 (Mitten 1862). *** Riccardia poeppigiana (Lehm. et Lindenb.) Hässel ex Meenks et C.De Jong, Cryptog. Bryol. Lichénol. 6 (1): 17, 1985 (Meenks and de Jong 1985). Bas.: Jungermannia poeppigiana Lehm. et Lindenb., Nov. Stirp. Pug. 6: 23, 1834 (Lehmann 1834). 454 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Riccardia porcina (Hewson) L.Söderstr., Phytotaxa 202 (1): 70, 2015 (Söderström et al. 2015c). Bas.: Riccardia bliklika var. porcina Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 84, 1970 (Hewson 1970a). ** Riccardia portoricensis (Steph.) Pagán, Bryologist 42 (1): 7, 1939 (Pagán 1939a). Bas.: Aneura portoricensis Steph., Bull. Herb. Boissier 7 (10): 739 (249), 1899 (Stephani 1899f). ** Riccardia punahuina (Steph.) H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 526, 1963 (Miller 1963). Bas.: Aneura punahuina Steph., Sp. Hepat. (Stephani) 6: 39, 1917 (Stephani 1917a). ** Riccardia ramosissima (Steph.) Grolle, Bryophyt. Biblioth. 48: 130, 1995 (Grolle 1995). Bas.: Aneura ramosissima Steph., Bull. Soc. Roy. Bot. Belgique, Compt. Rend. 30 (2): 196, 1891 [1892] (Stephani 1891b). ** Riccardia regina Meenks et C.De Jong, Cryptog. Bryol. Lichénol. 6 (1): 17, 1985 (Meenks and de Jong 1985). *** Riccardia regnellii (Ångstr.) K.G.Hell, Bol. Fac. Filos. Univ. São Paulo, Bot. 25: 110, 1969 (Hell 1969). Bas.: Pseudoneura regnellii Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 33 (7): 90, 1876 [1877] (Ångström 1876). ** Riccardia reyesiana Meenks, Acta Bot. Hung. 32 (1/4): 207, 1986 (Meenks 1986). ** Riccardia riccioides Pearson, Univ. Calif. Publ. Bot. 10 (4): 310, 1923 (Pearson 1923). * Riccardia rigida Schifn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 172, 1898 (Schifner 1898a). ** Riccardia robbinsii Hewson et Grolle, J. Hattori Bot. Lab. 29: 72, 1966 (Grolle 1966i). ** Riccardia robusta (Steph.) H.A.Mill., Phytologia 47 (4): 324, 1981 (Miller 1981). Bas.: Aneura robusta Steph., Sp. Hepat. (Stephani) 6: 40, 1917 (Stephani 1917a). ** Riccardia rockii (Steph.) H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 527, 1963 (Miller 1963). Bas.: Aneura rockii Steph., Sp. Hepat. (Stephani) 6: 41, 1917 (Stephani 1917a). ** Riccardia rupicola (Steph.) Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 99, 1970 (Hewson 1970a). Bas.: Aneura rupicola Steph., Sp. Hepat. (Stephani) 6: 41, 1917 (Stephani 1917a). ** Riccardia russellii R.M.Schust., J. Hattori Bot. Lab. 67: 93, 1989 (Schuster 1989). ** Riccardia saccatilora (Steph.) S.W.Arnell, Bot. Not. 105: 144, 1952 (Arnell 1952b). Bas.: Aneura saccatilora Steph., Bot. Gaz. 15 (11): 282, 1890 (Stephani 1890c). ** Riccardia santapaui Udar et S.C.Srivast., Rev. Bryol. Lichénol. 39 (1): 155, 1973 (Udar and Srivastava 1973). *** Riccardia schwaneckei (Steph.) Pagán, Bryologist 42 (1): 7, 1939 (Pagán 1939a). Bas.: Aneura schwaneckei Steph., Hedwigia 27 (11/12): 278, 1888 (Stephani 1888c). ** Riccardia singapurensis Schifn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 165, 1898 (Schifner 1898a). ** Riccardia smaragdina Meenks et C.De Jong, Cryptog. Bryol. Lichénol. 6 (1): 20, 1985 (Meenks and de Jong 1985). *** Riccardia sprucei (Steph.) Meenks et C.De Jong, Cryptog. Bryol. Lichénol. 6 (1): 22, 1985 (Meenks and de Jong 1985). Bas.: Aneura sprucei Steph., Bull. Herb. Boissier 5 (10): 844, 1897 (Stephani 1897c). World checklist of hornworts and liverworts 455 ** Riccardia squamifera Schifn., Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 19, 1964 (Schifner and Arnell 1964). * Riccardia stipatilora (Steph.) Pagán, Bryologist 45 (4): 81, 1942 (Pagán 1942b). Bas.: Aneura stipatilora Steph., Hedwigia 32 (1): 27, 1893 (Stephani 1893a). ** Riccardia stricta R.M.Schust., J. Hattori Bot. Lab. 62: 326, 1987 (Schuster 1987d). ** Riccardia subantarctica Grolle et L.Söderstr., Phytotaxa 202 (1): 70, 2015 (Söderström et al. 2015c). Nom. nov. pro Riccardia pauciramea R.M.Schust., J. Hattori Bot. Lab. 67: 102, 1989 (Schuster 1989), nom. illeg. ** Riccardia subexalata Schifn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 163, 1898 (Schifner 1898a). * Riccardia subexalata var. procera Schifn., Denkschr. Kaiserl. Akad. Wiss., Math.Naturwiss. Kl. 67: 164, 1898 (Schifner 1898a). * Riccardia submersa (Hook.f. et Taylor) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 431, 1877 (Trevisan 1877). Bas.: Jungermannia multiida var. submersa Hook.f. et Taylor, London J. Bot. 4: 94, 1845 (Hooker and Taylor 1845). ** Riccardia submultiida Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 128, 1934 (Horikawa 1934). ** Riccardia subpalmata (Steph.) Hürl., Bauhinia 5 (4): 194, 1976 (Hürlimann 1976). Bas.: Aneura subpalmata Steph., Sp. Hepat. (Stephani) 6: 43, 1917 (Stephani 1917a). ** Riccardia subsimplex (Steph.) Pagán, Bryologist 45 (4): 81, 1942 (Pagán 1942b). Bas.: Aneura subsimplex Steph., Hedwigia 32 (1): 27, 1893 (Stephani 1893a). ** Riccardia sumatrana Schifn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 173, 1898 (Schifner 1898a). ** Riccardia tahitensis (Steph.) Hürl., Bauhinia 5 (4): 201, 1976 (Hürlimann 1976). Bas.: Aneura tahitensis Steph., Bull. Herb. Boissier 7 (10): 728 (238), 1899 (Stephani 1899f). * Riccardia tenella Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 95, 1970 (Hewson 1970a). *** Riccardia tenuicula (Spruce) Meenks, Beih. Nova Hedwigia 88: 101, 1987 (Schultze-Motel and Menzel 1987). Bas.: Aneura tenuicula Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 545, 1885 (Spruce 1885). ** Riccardia tenuis (Steph.) Schifn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 172, 1898 (Schifner 1898a). Bas.: Aneura tenuis Steph., Hedwigia 32 (1): 28, 1893 (Stephani 1893a). * Riccardia tjibodensis Schifn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 165, 1898 (Schifner 1898a). *** Riccardia trichomanoides (Spruce) Hässel ex Meenks, Beih. Nova Hedwigia 88: 101, 1987 (Schultze-Motel and Menzel 1987). Bas.: Aneura trichomanoides Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 547, 1885 (Spruce 1885). ** Riccardia trukensis H.A.Mill. et Bonner, Beih. Nova Hedwigia 11: 70, 1963 (Miller et al. 1963). ** Riccardia tumbareriensis Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 87, 1970 (Hewson 1970a). 456 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Riccardia upoluna (Steph.) Grolle, J. Hattori Bot. Lab. 36: 77, 1972 [1973] (Grolle and Schultze-Motel 1972). Bas.: Aneura upoluna Steph., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 91: 165, 1915 (Stephani 1915a). ** Riccardia valida (Steph.) J.J.Engel, Bryologist 78 (3): 362, 1975 (Engel 1975). Bas.: Aneura valida Steph., Sp. Hepat. (Stephani) 6: 44, 1917 (Stephani 1917a). ** Riccardia venosa (Steph.) Hürl., Bauhinia 5 (4): 208, 1976 (Hürlimann 1976). Bas.: Aneura venosa Steph., Sp. Hepat. (Stephani) 6: 45, 1917 (Stephani 1917a). * Riccardia villosa (Steph.) Pandé et S.C.Srivast., Biol. Mem. 1 (1/2): 129, 1976 (Srivastava and Udar 1976). Bas.: Aneura villosa Steph., Sp. Hepat. (Stephani) 6: 45, 1917 (Stephani 1917a). ** Riccardia virens (Steph.) Hürl., Bauhinia 5 (4): 209, 1976 (Hürlimann 1976). Bas.: Aneura virens Steph., Sp. Hepat. (Stephani) 6: 45, 1917 (Stephani 1917a). ** Riccardia virgata (Gottsche) Pagán, Bryologist 42 (1): 7, 1939 (Pagán 1939a). Bas.: Aneura virgata Gottsche, Hedwigia 27 (11/12): 277, 1888 (Stephani 1888c). ** Riccardia wallisii (Steph.) Gradst., J. Hattori Bot. Lab. 45: 130, 1979 (Gradstein and Hekking 1979). Bas.: Aneura wallisii Steph., Hedwigia 32 (1): 28, 1893 (Stephani 1893a). ** Riccardia wettsteinii Schifn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 162, 1898 (Schifner 1898a). * Riccardia wettsteinii var. angustilimbia Schifn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 163, 1898 (Schifner 1898a). * Riccardia wettsteinii var. crassa Schifn., Denkschr. Kaiserl. Akad. Wiss., Math.Naturwiss. Kl. 67: 163, 1898 (Schifner 1898a). * Riccardia wettsteinii var. procera Schifn., Denkschr. Kaiserl. Akad. Wiss., Math.Naturwiss. Kl. 67: 162, 1898 (Schifner 1898a). * Riccardia wettsteinii var. tenuiretis Schifn., Denkschr. Kaiserl. Akad. Wiss., Math.Naturwiss. Kl. 67: 162, 1898 (Schifner 1898a). ** Riccardia womersleyana Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 98, 1970 (Hewson 1970a). ** Verdoornia R.M.Schust., J. Hattori Bot. Lab. 26: 291, 1963 (Schuster 1963b). *** Verdoornia succulenta R.M.Schust., J. Hattori Bot. Lab. 26: 291, 1963 (Schuster 1963b). *** Metzgeriaceae H.Klinggr. by D. P. Costa he treatment of Metzgeriaceae follows what was outlined in Hewson (1982), Kuwahara (1986), Crandall-Stotler and Stotler (2000), and Costa (2008). World checklist of hornworts and liverworts 457 *** Metzgeria Raddi, Jungermanniogr. Etrusca: 34, 1818 (Raddi 1818a). 407 *** Metzgeria acuminata Steph., Bull. Herb. Boissier 7 (12): 934 (282), 1899 (Stephani 1899g). *** Metzgeria adscendens Steph. ex K.I.Goebel, Flora 77: 427, 1893 (Goebel 1893a). *** Metzgeria agnewiae Kuwah., Bryologist 76 (4): 569, 1973 (Kuwahara 1973b). 408 *** Metzgeria albinea Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: cci, 1889 [1890] (Spruce 1889). *** Metzgeria albinea var. aberrans Schifn., Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 27, 1964 (Schifner and Arnell 1964). *** Metzgeria albinea var. angusta (Steph.) D.P.Costa et Gradst., Bryologist 103 (4): 757, 2000 [2001] (Costa and Gradstein 2000). Bas.: Metzgeria angusta Steph., Bull. Herb. Boissier 7 (12): 944 (292), 1899 (Stephani 1899g). *** Metzgeria allionii Steph., Sp. Hepat. (Stephani) 6: 47, 1917 (Stephani 1917a). *** Metzgeria alpina R.M.Schust. et J.J.Engel, Brittonia 40 (2): 203, 1988 (Engel and Schuster 1988). *** Metzgeria americana Masuzaki, Hikobia 15 (4): 441, 2010 (Masuzaki et al. 2010a). *** Metzgeria attenuata Steph., Biblioth. Bot. 87 (2): 177, 1916 (Stephani 1916a). *** Metzgeria aurantiaca Steph., Bull. Herb. Boissier 7 (12): 938 (286), 1899 (Stephani 1899g). *** Metzgeria auriculata Grolle et Kuwah., Bryophyt. Biblioth. 28: 194, 1986 (Kuwahara 1986). *** Metzgeria bahiensis Schifn., Österr. Bot. Z. 61 (7/8): 262, 1911 (Schifner 1911). *** Metzgeria bartlettii Kuwah., Mem. New York Bot. Gard. 45: 561, 1987 (Kuwahara 1987). *** Metzgeria bischlerae Kuwah., J. Hattori Bot. Lab. 40: 264, 1976 (Kuwahara 1976a). *** Metzgeria bracteata Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 553, 1885 (Spruce 1885). *** Metzgeria brasiliensis Schifn., Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 22, 1964 (Schifner and Arnell 1964). *** Metzgeria chilensis Steph., Bull. Herb. Boissier 7 (12): 937 (285), 1899 (Stephani 1899g). *** Metzgeria ciliata Raddi, Critt. Brasil.: 17, 1822 (Raddi 1822). 409 *** Metzgeria clavilora Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 556, 1885 (Spruce 1885). 407 Metzgeria was divided into many subdivisions proposed by Kuwahara in a series of publications, but they are artiicial and not accepted here. 408 Metzgeria agnewiae is conspeciic with Metzgeria consanguinea in So (2004), but it was accepted by Costa (2008). 409 Metzgeria ciliata is conspeciic with Metzgeria furcata in Grolle (2002) and with Metzgeria dichotoma in Gradstein and Costa (2003), but it was accepted by Costa (2008). 458 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Metzgeria cleeii Kuwah., Proc. Kon. Ned. Akad. Wetensch. C 85 (3): 360, 1982 (Kuwahara 1982). ** Metzgeria comata Steph., Bull. Herb. Boissier 7 (12): 939 (287), 1899 (Stephani 1899g). *** Metzgeria conjugata Lindb., Acta Soc. Sci. Fenn. 10: 495, 1875 (Lindberg 1875). 410 *** Metzgeria consanguinea Schifn., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 60 (2): 271, 1893 (Schifner 1893a). *** Metzgeria convoluta Steph., Bull. Herb. Boissier 7 (12): 940 (288), 1899 (Stephani 1899g). ** Metzgeria coorgensis S.C.Srivast. et S.Srivast., Phytotaxonomy 4: 81, 2004 (Srivastava and Srivastava 2004). ** Metzgeria corralensis Steph., Bull. Herb. Boissier 7 (12): 933 (281), 1899 (Stephani 1899g). *** Metzgeria crassipilis (Lindb.) A.Evans, Rhodora 11 (130): 188, 1909 (Evans 1909). Bas.: Metzgeria furcata subsp. crassipilis Lindb., Acta Soc. Fauna Fl. Fenn. 1 (2): 42, 1877 (Lindberg 1877b). *** Metzgeria cratoneura Schifn., Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 24, 1964 (Schifner and Arnell 1964). *** Metzgeria cylindra Kuwah., Bryologist 81 (3): 405, 1978 (Kuwahara 1978). ** Metzgeria decrescens Steph., Bull. Herb. Boissier 7 (12): 932 (280), 1899 (Stephani 1899g). *** Metzgeria dichotoma (Sw.) Nees, Syn. Hepat. 4: 504, 1846 (Gottsche et al. 1846). Bas.: Jungermannia dichotoma Sw., Prodr. (Swartz): 145, 1788 (Swartz 1788). ** Metzgeria divaricata A.Evans, Proc. Amer. Acad. Arts 58 (7): 288, 1923 (Evans 1923a). *** Metzgeria dorsipara (Herzog) Kuwah., J. Hattori Bot. Lab. 40: 269, 1976 (Kuwahara 1976a). Bas.: Metzgeria violacea var. dorsipara Herzog, Svensk Bot. Tidskr. 51 (1): 187, 1957 (Herzog 1957a). *** Metzgeria duricosta Steph., Sp. Hepat. (Stephani) 6: 50, 1917 (Stephani 1917a). ** Metzgeria engelii Kuwah., Hikobia 8 (3/4): 275, 1980 (Kuwahara 1980a). ** Metzgeria epiphylla A.Evans, Proc. Amer. Acad. Arts 58 (7): 303, 1923 (Evans 1923a). *** Metzgeria ilicina Mitt., Hooker’s J. Bot. Kew Gard. Misc. 3: 361, 1851 (Mitten 1851). ** Metzgeria lavovirens Colenso, Trans. & Proc. New Zealand Inst. 21: 79, 1889 (Colenso 1889). * Metzgeria foliicola Schifn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 181, 1898 (Schifner 1898a). 411 *** Metzgeria francana Steph., Sp. Hepat. (Stephani) 6: 51, 1917 (Stephani 1917a). 410 Metzgeria conjugata is a complex taxon. Fuselier et al. (2009) demonstrated the occurrence of two clearly separated lineages, a northern North American lineage (possibly corresponding to Metzgeria conjugata s.str.) and a southern North American and European lineage (possibly corresponding to Metzgeria simplex). 411 Metzgeria foliicola is conspeciic with Metzgeria albinea in Kuwahara (1986), but it was accepted by So (2002c). World checklist of hornworts and liverworts 459 ** Metzgeria frontipilis Lindb., Acta Soc. Fauna Fl. Fenn. 1 (2): 14, 1877 (Lindberg 1877b). *** Metzgeria fruticola Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 554, 1885 (Spruce 1885). *** Metzgeria furcata (L.) Corda, Gen. hepat.: 654, 1829 (Corda 1829). Bas.: Jungermannia furcata L., Sp. Pl. 1: 1136, 1753 (Linnaeus 1753). 412 * Metzgeria furcata var. expansa Douin, Rev. Bryol. 30 (3): 47, 1903 (Douin 1903). * Metzgeria furcata var. paciica Brinkm., Bryologist 34 (2): 15, 1931 (Brinkman 1931). *** Metzgeria grandilora A.Evans, Torreya 16 (3): 68, 1916 (Evans 1916). ** Metzgeria hasselii Kuwah., J. Hattori Bot. Lab. 40: 509, 1976 (Kuwahara 1976c). ** Metzgeria hebridensis Steph., Sp. Hepat. (Stephani) 6: 51, 1917 (Stephani 1917a). *** Metzgeria hegewaldii Kuwah., Nova Hedwigia 34: 784, 1981 (Kuwahara 1981). *** Metzgeria herminieri Schifn., Österr. Bot. Z. 61 (7/8): 261, 1911 (Schifner 1911). * Metzgeria heteroramea Steph., Biblioth. Bot. 87 (2): 178, 1916 (Stephani 1916a). ** Metzgeria imberbis J.B.Jack et Steph., Hedwigia 34 (6): 316, 1895 (Jack and Stephani 1895). *** Metzgeria inlata Steph., Bull. Herb. Boissier 7 (12): 936 (284), 1899 (Stephani 1899g). *** Metzgeria jamesonii Kuwah., Bryophyt. Biblioth. 28: 157, 1986 (Kuwahara 1986). ** Metzgeria kanaii Kuwah., Fl. E. Himalaya 2: 240, 1971 (Hattori 1971a). *** Metzgeria kinabaluensis Masuzaki, Hikobia 16 (1): 59, 2011 (Masuzaki 2011). Based on: Apometzgeria pubescens var. kinabaluensis Kuwah., J. Hattori Bot. Lab. 28: 166, 1965 (Kuwahara 1965), nom. inval. ** Metzgeria kuwaharae Piippo, Acta Bot. Fenn. 143: 8, 1991 (Piippo 1991). ** Metzgeria laciniata Kuwah., Bryologist 81 (3): 406, 1978 (Kuwahara 1978). *** Metzgeria lechleri Steph., Bull. Herb. Boissier 7 (12): 942 (290), 1899 (Stephani 1899g). *** Metzgeria leptoneura Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 555, 1885 (Spruce 1885). ** Metzgeria leptoneura var. breviseta (Schifn.) O.Yano, J. Hattori Bot. Lab. 56: 532, 1984 (Yano 1984). Bas.: Metzgeria hamata var. breviseta Schifn., Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 27, 1964 (Schifner and Arnell 1964). ** Metzgeria leptoneura var. polychaeta R.M.Schust., Phytotaxa 202 (1): 70, 2015 (Söderström et al. 2015c). Based on: Metzgeria leptoneura var. polychaeta R.M.Schust., J. Hattori Bot. Lab. 70: 150, 1991 (Schuster 1991b), nom. inval. *** Metzgeria liebmaniana Lindenb. et Gottsche, Syn. Hepat. 4: 505, 1846 (Gottsche et al. 1846). *** Metzgeria lindbergii Schifn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 182, 1898 (Schifner 1898a). ** Metzgeria litoralis J.J.Engel et Kuwah., Bryologist 76 (2): 293, 1973 (Engel and Kuwahara 1973). 412 Metzgeria furcata includes several cryptic species as demonstrated by Fuselier et al. (2009). 460 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Metzgeria longitexta Steph., Bull. Herb. Boissier 7 (12): 940 (288), 1899 (Stephani 1899g). ** Metzgeria macrospora Kuwah., J. Hattori Bot. Lab. 32: 17, 1969 (Kuwahara 1969a). ** Metzgeria macveanii Kuwah., Rev. Bryol. Lichénol. 36 (3/4): 539, 1969 [1970] (Kuwahara 1969b). *** Metzgeria maegdefraui Kuwah., Hikobia 8 (3/4): 269, 1980 (Kuwahara 1980b). ** Metzgeria magellanica Schifn., Leberm., Forschungsr. Gazelle 4 (4): 43, 1890 (Schifner 1890). *** Metzgeria metaensis Kuwah., Proc. Kon. Ned. Akad. Wetensch. C 85 (3): 375, 1982 (Kuwahara 1982). *** Metzgeria mexicana Steph., Sp. Hepat. (Stephani) 6: 55, 1917 (Stephani 1917a). 413 ** Metzgeria monoica Kuwah. et J.J.Engel, Hikobia 8 (3/4): 284, 1980 (Kuwahara 1980a). *** Metzgeria myriopoda Lindb., Acta Soc. Fauna Fl. Fenn. 1 (2): 22, 1877 (Lindberg 1877b). *** Metzgeria neotropica Kuwah., Nova Hedwigia 34: 779, 1981 (Kuwahara 1981). *** Metzgeria nudifrons Steph., Hedwigia 31 (3): 126, 1892 (Stephani 1892d). *** Metzgeria parviinvolucrata Kuwah., Nova Hedwigia 34: 774, 1981 (Kuwahara 1981). ** Metzgeria patagonica Steph., Bull. Herb. Boissier 7 (12): 940 (288), 1899 (Stephani 1899g). *** Metzgeria polytricha Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 553, 1885 (Spruce 1885). *** Metzgeria procera Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 166, 1855 (Mitten 1855). *** Metzgeria psilocraspeda Schifn., Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 25, 1964 (Schifner and Arnell 1964). *** Metzgeria pubescens (Schrank) Raddi, Jungermanniogr. Etrusca: 35, 1818 (Raddi 1818a). Bas.: Jungermannia pubescens Schrank, Prim. Fl. Salisb.: 231, 1792 (Schrank 1792). *** Metzgeria pulvinata Steph., Biblioth. Bot. 87 (2): 179, 1916 (Stephani 1916a). ** Metzgeria quadrifaria Steph., Bull. Herb. Boissier 7 (12): 953 (301), 1899 (Stephani 1899g). ** Metzgeria raoi S.C.Srivast. et S.Srivast., Phytotaxonomy 4: 83, 2004 (Srivastava and Srivastava 2004). ** Metzgeria rigida Lindb., Acta Soc. Fauna Fl. Fenn. 1 (2): 43, 1877 (Lindberg 1877b). ** Metzgeria robinsonii Steph., Sp. Hepat. (Stephani) 6: 60, 1917 (Stephani 1917a). 413 Metzgeria mexicana is conspeciic with Metzgeria subundulata in Schuster (1992b), but it was accepted by Costa (2008). World checklist of hornworts and liverworts 461 ** Metzgeria roivainenii Kuwah., J. Hattori Bot. Lab. 40: 516, 1976 (Kuwahara 1976c). *** Metzgeria rufula Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 555, 1885 (Spruce 1885). ** Metzgeria saccata Mitt., J. Linn. Soc., Bot. 22 (145): 241, 1886 (Mitten 1886a). ** Metzgeria saxbyi Pearson, Ann. Cryptog. Exot. 4 (2): 70, 1931 (Pearson 1931a). ** Metzgeria scobina Mitt., J. Linn. Soc., Bot. 22 (145): 243, 1886 (Mitten 1886a). *** Metzgeria scyphigera A.Evans, Trans. Connecticut Acad. Arts 18 (5): 299, 1914 (Evans 1914c). *** Metzgeria senjoana Masuzaki, Hikobia 15 (4): 445, 2010 (Masuzaki et al. 2010a). ** Metzgeria setigera R.M.Schust. ex Crand.-Stotl. et L.Söderstr., Phytotaxa 202 (1): 69, 2015 (Söderström et al. 2015c). Based on: Metzgeria furcata var. setigera R.M.Schust., J. Hattori Bot. Lab. 70: 149, 1991 (Schuster 1991b), nom. inval. ** Metzgeria sikkimensis S.C.Srivast. et K.K.Rawat, Geophytology 31 (1/2): 71, 2001 [2003] (Srivastava and Rawat 2001). * Metzgeria simplex Lorb. ex Müll.Frib., Hedwigia 80 (1/2): 115, 1941 (Müller 1941). *** Metzgeria sinuata Loitl., Diagn. pl. nov.: 25, 1894 (Loitlesberger 1894). ** Metzgeria sparrei Kuwah., Hikobia 8 (3/4): 278, 1980 (Kuwahara 1980a). *** Metzgeria spindleri Steph., Biblioth. Bot. 87 (2): 179, 1916 (Stephani 1916a). *** Metzgeria subaneura Schifn., Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 22, 1964 (Schifner and Arnell 1964). ** Metzgeria submarginata M.L.So, New Zealand J. Bot. 40 (2): 201, 2002 (So 2002b). ** Metzgeria subundulata (Lindb.) Kuwah., Bryologist 86 (3): 276, 1983 [1984] (Kuwahara 1983). Bas.: Metzgeria furcata subsp. subundulata Lindb., Acta Soc. Fauna Fl. Fenn. 1 (2): 42, 1877 (Lindberg 1877b). ** Metzgeria temperata Kuwah., J. Hattori Bot. Lab. 40: 219, 1976 (Kuwahara 1976b). *** Metzgeria uncigera A.Evans, Ann. Bot. (Oxford) 24 (2): 276, 1910 (Evans 1910). *** Metzgeria undulata Kuwah., Nova Hedwigia 34: 792, 1981 (Kuwahara 1981). *** Metzgeria violacea (Ach.) Dumort., Recueil Observ. Jungerm.: 26, 1835 (Dumortier 1835). Bas.: Jungermannia violacea Ach., Beitr. Naturk. (Weber & Mohr) 1: 77, 1805 (Acharius 1805). ** Steereella Kuwah., Amer. J. Bot. 60 (6): 602, 1973 (Kuwahara 1973a). *** Steereella lilliana (Steph.) Kuwah., Bryophyt. Biblioth. 28: 179, 1986 (Kuwahara 1986). Bas.: Metzgeria lilliana Steph., Sp. Hepat. (Stephani) 6: 53, 1917 (Stephani 1917a). *** Steereella linearis (Sw.) Kuwah., Amer. J. Bot. 60 (6): 604, 1973 (Kuwahara 1973a). Bas.: Jungermannia linearis Sw., Prodr. (Swartz): 145, 1788 (Swartz 1788). ** Vandiemenia Hewson, J. Hattori Bot. Lab. 52: 163, 1982 (Hewson 1982). ** Vandiemenia ratkowskiana Hewson, J. Hattori Bot. Lab. 52: 163, 1982 (Hewson 1982). 462 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Pleuroziales Schljakov *** Pleuroziaceae Müll.Frib. by B. hiers *** Pleurozia Dumort., Recueil Observ. Jungerm.: 15, 1835 (Dumortier 1835). ** subg. Constantifolia B.M.hiers, Bryologist 96 (4): 526, 1993 (hiers 1993). *** Pleurozia conchifolia (Hook. et Arn.) Austin, Bull. Torrey Bot. Club 5 (3): 16, 1874 (Austin 1874). Bas.: Jungermannia conchifolia Hook. et Arn., Bot. Beechey Voy. 3: 110, 1832 (Hooker and Arnott 1832). ** Pleurozia conchifolia var. papillosa B.M.hiers, Bryologist 96 (4): 528, 1993 (hiers 1993). *** Pleurozia purpurea Lindb., Acta Soc. Fauna Fl. Fenn. 1 (2): 27, 1877 (Lindberg 1877b). Based on: Jungermannia purpurea Lightf., Fl. Scot. 2: 778, 1777 (Lightfoot 1777), nom. illeg. ** subg. Diversifolia B.M.hiers, Bryologist 96 (4): 531, 1993 (hiers 1993). *** Pleurozia acinosa (Mitt.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 412, 1877 (Trevisan 1877). Bas.: Physiotium acinosum Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 102, 1860 [1861] (Mitten 1860c). *** Pleurozia articulata (Lindb.) Lindb. et Lackström, Hepat. Scand. Exsicc.: 5 (adnot.), 1874 (Lindberg and Lackström 1874). Bas.: Physiotium articulatum Lindb., Öfvers. Förh. Finska Vetensk.-Soc. 12 (2): 78, 1870 (Lindberg 1870). *** Pleurozia caledonica (Gottsche) Steph., Rev. Bryol. 33 (2): 29 (Paris 1906a). Bas.: Physiotium caledonicum Gottsche, Hedwigia 25 (2/3): 81, 1886 (Jack 1886). *** Pleurozia curiosa B.M.hiers, Bryologist 96 (4): 537, 1993 (hiers 1993). *** Pleurozia heterophylla Steph. ex Fulford, Mem. New York Bot. Gard. 23: 842, 1972 (Fulford 1972). *** Pleurozia johannis-winkleri Herzog, Mitt. Inst. Allg. Bot. Hamburg 7 (3): 195, 1931 (Herzog 1931a). *** Pleurozia paradoxa (J.B.Jack) Schifn., Hepat. (Engl.-Prantl): 115, 1893 (Schifner 1893b). Bas.: Physiotium paradoxum J.B.Jack, Hedwigia 25 (2/3): 85, 1886 (Jack 1886). *** Pleurozia subinlata (Austin) Austin, Bull. Torrey Bot. Club 5 (3): 16, 1874 (Austin 1874). Bas.: Physiotium subinlatum Austin, Proc. Acad. Nat. Sci. Philadelphia 21: 224, 1869 (Austin 1869). ** subg. Pleurozia *** Pleurozia gigantea (F.Weber) Lindb., Hepat. Scand. Exsicc.: no. 5, 1874 (Lindberg and Lackström 1874). Bas.: Jungermannia gigantea F.Weber, Hist. Musc. Hepat. Prodr.: 57, 1815 (Weber 1815). World checklist of hornworts and liverworts 463 Incertae sedis ** Pleurozia pocsii Frank Müll., Polish Bot. J. 58 (1): 50, 2013 (Müller 2013). Pelliidae He-Nygrén, Juslén, Ahonen, Glenny et Piippo Fossombroniales Schljakov Calyculariineae He-Nygrén, Juslén, Ahonen, Glenny et Piippo *** Calyculariaceae He-Nygrén, Juslén, Ahonen, Glenny et Piippo *** Calycularia Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 122, 1860 [1861] (Mitten 1860c). *** Calycularia crispula Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 122, 1860 [1861] (Mitten 1860c). *** Calycularia laxa Lindb. et Arnell, Kongl. Svenska Vetensk.-Akad. Handl. (n.ser.) 23 (5): 66, 1889 (Lindberg and Arnell 1889). Fossombroniineae R.M.Schust. ex Stotler et Crand.-Stotl. ** Allisoniaceae Schljakov *** Allisonia Herzog, Hedwigia 80 (1/2): 77, 1941 (Herzog 1941a). *** Allisonia cockaynei (K.I.Goebel) R.M.Schust., J. Hattori Bot. Lab. 26: 294, 1963 (Schuster 1963b). Bas.: Moerckia cockaynei K.I.Goebel, Flora 96: 190, 1906 (Goebel 1906). *** Fossombroniaceae Hazsl. nom. conserv. by R. Stotler, B. J. Crandall-Stotler and D. C. Cargill *** Fossombronia Raddi, Jungermanniogr. Etrusca: 29, 1818 (Raddi 1818a). *** Fossombronia alaskana Steere et Inoue, Bryologist 77 (1): 66, 1974 (Steere and Inoue 1974). *** Fossombronia alata G.A.M.Scott et D.C.Pike, J. Hattori Bot. Lab. 56: 340, 1984 (Scott and Pike 1984). *** Fossombronia altilamellosa G.A.M.Scott et D.C.Pike, Fl. Austral. Suppl. 21: 114, 2003 (McCarthy 2003). Based on: Fossombronia altilamellosa G.A.M.Scott et D.C.Pike, J. Hattori Bot. Lab. 62: 367, 1987 (Scott and Pike 1987), nom. inval. *** Fossombronia angulifolia Perold, Bothalia 28 (2): 159, 1998 (Perold 1998b). *** Fossombronia angulosa (Dicks.) Raddi, Jungermanniogr. Etrusca: 29, 1818 (Raddi 1818a). Bas.: Jungermannia angulosa Dicks., Fasc. Pl. Crypt. Brit. 1: 7, 1785 (Dickson 1785). 464 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Fossombronia areolata G.A.M.Scott et D.C.Pike, J. Hattori Bot. Lab. 62: 368, 1987 (Scott and Pike 1987). ** Fossombronia auricolor G.A.M.Scott et D.C.Pike, Fl. Austral. Suppl. 21: 114, 2003 (McCarthy 2003). Based on: Fossombronia auricolor G.A.M.Scott et D.C.Pike, J. Hattori Bot. Lab. 62: 370, 1987 (Scott and Pike 1987), nom. inval. *** Fossombronia australis Mitt., J. Linn. Soc., Bot. 15 (82): 73, 1876 (Mitten 1876a). *** Fossombronia caespitiformis (Raddi) De Not. ex Rabenh., Hepat. Eur., Leberm. 13-14: no. 123, 1860 [1861] (Rabenhorst 1860). Bas.: Fossombronia angulosa var. caespitiformis Raddi, Jungermanniogr. Etrusca: 30, 1818 (Raddi 1818a). *** Fossombronia caespitiformis subsp. multispira (Schifn.) J.R.Bray et Cargill, Bryologist 106 (1): 131, 2003 (Stotler et al. 2003). Bas.: Fossombronia caespitiformis var. multispira Schifn., Österr. Bot. Z. 67 (4/5): 152, 1918 (Schifner 1918). *** Fossombronia caledonica Steph., Sp. Hepat. (Stephani) 6: 71, 1917 (Stephani 1917a). *** Fossombronia cederbergensis Perold, Bothalia 28 (1): 1, 1998 (Perold 1998c). *** Fossombronia cerebriformis G.A.M.Scott et D.C.Pike, J. Hattori Bot. Lab. 56: 343, 1984 (Scott and Pike 1984). *** Fossombronia crassifolia Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 527, 1885 (Spruce 1885). *** Fossombronia crispa Nees, Syn. Hepat. 4: 469, 1846 (Gottsche et al. 1846). * Fossombronia crispula (Brot.) R.M.Schust., Hepat. Anthocerotae N. Amer. 5: 383, 1992 (Schuster 1992b). Bas.: Jungermannia crispula Brot., Fl. lusit. 2: 422, 1804 [1805] (Brotero 1804). 414 *** Fossombronia cristula Austin, Proc. Acad. Nat. Sci. Philadelphia 21: 228, 1869 (Austin 1869). *** Fossombronia cultriformis G.A.M.Scott et D.C.Pike, J. Hattori Bot. Lab. 62: 371, 1987 (Scott and Pike 1987). *** Fossombronia densa G.A.M.Scott et D.C.Pike, J. Hattori Bot. Lab. 62: 372, 1987 (Scott and Pike 1987). *** Fossombronia densilamellata S.W.Arnell, Bot. Not. 105: 317, 1952 (Arnell 1952a). *** Fossombronia echinata Macvicar, Rev. Bryol. 38 (4): 73, 1911 (Macvicar 1911). *** Fossombronia elsieae Perold, Bothalia 29 (1): 25, 1999 (Perold 1999c). *** Fossombronia fernandeziensis Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n. ser.) 46 (9): 15, 1911 (Stephani 1911b). *** Fossombronia imbriata Paton, J. Bryol. 8 (1): 1, 1974 (Paton 1974). *** Fossombronia leischeri Osterwald ex Loeske, Verh. Bot. Vereins Prov. Brandenburg 70: 125, 1928 (Loeske 1928). *** Fossombronia foveolata Lindb., Helsingf. Dagbl. 1873 (353, 28 Dec): 2, 1873 (Lindberg 1873b). ** Fossombronia fuhreri G.A.M.Scott et D.C.Pike, J. Hattori Bot. Lab. 62: 374, 1987 (Scott and Pike 1987). 414 Fossombronia crispula is based on the description most likely Fossombronia pusilla, but no material has been found. World checklist of hornworts and liverworts 465 *** Fossombronia gemmifera Perold, Bothalia 27 (1): 19, 1997 (Perold 1997a). *** Fossombronia glenii Perold, Bothalia 27 (1): 20, 1997 (Perold 1997a). ** Fossombronia grandis Steph., Mém. Herb. Boissier 16: 29 (383), 1900 (Stephani 1900a). * Fossombronia gregaria Colenso, Trans. & Proc. New Zealand Inst. 20: 252, 1888 (Colenso 1888). 415 * Fossombronia grossepapillata Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 105, 1914 (Stephani and Watts 1914). 416 *** Fossombronia hamatohirta Steph., Hedwigia 33 (1): 8, 1894 (Stephani 1894a). ** Fossombronia hewsoniae G.A.M.Scott et D.C.Pike, J. Hattori Bot. Lab. 62: 375, 1987 (Scott and Pike 1987). *** Fossombronia himalayensis Kashyap, New Phytol. 14 (1): 4, 1915 (Kashyap 1915). *** Fossombronia hyalorhiza Perold, Bothalia 29 (1): 83, 1999 (Perold 1999d). *** Fossombronia incurva Lindb., Helsingf. Dagbl. 1873 (273, 7 Oct): 2, 1873 (Lindberg 1873c). *** Fossombronia indica Steph., Sp. Hepat. (Stephani) 6: 73, 1917 (Stephani 1917a). * Fossombronia integerrima Steph., Mém. Herb. Boissier 16: 40 (394), 1900 (Stephani 1900a). 417 * Fossombronia integrifolia Steph., Sp. Hepat. (Stephani) 6: 73, 1917 (Stephani 1917a). 418 *** Fossombronia intestinalis Taylor, London J. Bot. 5: 408, 1846 (Taylor 1846b). *** Fossombronia japonica Schifn., Österr. Bot. Z. 49 (11): 389, 1899 (Schifner 1899c). ** Fossombronia laciniata G.A.M.Scott et D.C.Pike, J. Hattori Bot. Lab. 62: 375, 1987 (Scott and Pike 1987). *** Fossombronia lamellata Steph., Hedwigia 33 (1): 9, 1894 (Stephani 1894a). *** Fossombronia leucoxantha (Lehm.) Lehm. et Lindenb., Syn. Hepat. 4: 469, 1846 (Gottsche et al. 1846). Bas.: Jungermannia leucoxantha Lehm., Linnaea 4: 368, 1829 (Lehmann 1829). *** Fossombronia longiseta (Austin) Austin, Hepat. bor.-amer.: 30, 1873 (Austin 1873). Bas.: Androcryphia longiseta Austin, Proc. Acad. Nat. Sci. Philadelphia 21: 228, 1869 (Austin 1869). *** Fossombronia lophoclada Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 529, 1885 (Spruce 1885). ** Fossombronia lophoscypha Hässel, Beih. Nova Hedwigia 131: 16, 2007 (Hässel and Villagrán 2007). *** Fossombronia luetzelburgiana K.I.Goebel, Flora 105: 55, 1912 (Goebel 1912). *** Fossombronia macrocalyx Steph., Sp. Hepat. (Stephani) 6: 74, 1917 (Stephani 1917a). 415 Fossombronia gregaria is possibly conspeciic with Fossombronia australis (Scott and Pike 1988b). 416 Fossombronia grossepapillata may be conspeciic with Fossombronia papillata, but the type specimen is too immature to allow determination (Scott and Pike 1984). 417 Fossombronia integerrima is possibly conspeciic with Fossombronia australis. 418 Fossombronia integrifolia is possibly conspeciic with Fossombronia foveolata (Scott and Pike 1984). 466 * ** *** ** *** ** *** *** *** *** * ** *** *** *** *** ** *** *** *** *** *** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Fossombronia macrophylla Colenso, Trans. & Proc. New Zealand Inst. 18: 285, 1886 (Colenso 1886a). Fossombronia magnaspora G.A.M.Scott et D.C.Pike, Fl. Austral. Suppl. 21: 114, 2003 (McCarthy 2003). Based on: Fossombronia magnaspora G.A.M.Scott et D.C.Pike, J. Hattori Bot. Lab. 62: 377, 1987 (Scott and Pike 1987), nom. inval. Fossombronia marindae Perold, Bothalia 29 (1): 86, 1999 (Perold 1999d). Fossombronia maritima (Paton) Paton, J. Bryol. 18 (2): 367, 1994 (Paton 1994). Bas.: Fossombronia pusilla var. maritima Paton, J. Bryol. 7 (3): 244, 1973 (Paton 1973). Fossombronia marshii J.R.Bray et Stotler, Phytologia 92 (2): 230, 2010 (Stotler et al. 2010). Fossombronia microlamellata G.A.M.Scott et D.C.Pike, J. Hattori Bot. Lab. 56: 347, 1984 (Scott and Pike 1984). Fossombronia mittenii Tind., J. Bot. 36: 44, 1898 (Tindall 1898). Fossombronia montaguensis S.W.Arnell, Bot. Not. 105: 316, 1952 (Arnell 1952a). Fossombronia monticola Perold, Bothalia 29 (1): 87, 1999 (Perold 1999d). Fossombronia mylioides Inoue, J. Hattori Bot. Lab. 37: 296, 1973 (Inoue 1973). Fossombronia nigricaulis Colenso, Trans. & Proc. New Zealand Inst. 18: 248, 1886 (Colenso 1886b). 419 Fossombronia nyikaensis Perold, Bothalia 31 (1): 48, 2001 (Perold 2001c). Fossombronia papillata Steph., Hedwigia 28 (3): 157, 1889 (Stephani 1889d). Fossombronia paranapanemae Schifn., Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 34, 1964 (Schifner and Arnell 1964). Fossombronia peruviana Gottsche et Hampe, Linnaea 27 (5): 555, 1854 (Hampe 1854). Fossombronia porphyrorhiza (Nees) Prosk., Bryologist 58 (3): 197, 1955 (Proskauer 1955). Bas.: Jungermannia porphyrorhiza Nees, Fl. Bras. (Martius) 1 (1): 343, 1833 (Nees 1833a). Fossombronia pulvinata Steph., Wiss. Ergebn. Deut. Zentr.-Afr. Exped. (190708), Bot. 2: 113, 1911 (Stephani 1911a). Fossombronia punctata G.A.M.Scott et D.C.Pike, J. Hattori Bot. Lab. 56: 341, 1984 (Scott and Pike 1984). Fossombronia purpureospora G.A.M.Scott et D.C.Pike, Fl. Austral. Suppl. 21: 114, 2003 (McCarthy 2003). Based on: Fossombronia purpureospora G.A.M.Scott et D.C.Pike, J. Hattori Bot. Lab. 62: 379, 1987 (Scott and Pike 1987), nom. inval. Fossombronia pusilla (L.) Nees, Naturgesch. Eur. Leberm. 3: 319, 1838 (Nees 1838b). Bas.: Jungermannia pusilla L., Sp. Pl. 1: 1136, 1753 (Linnaeus 1753). Fossombronia renateae Perold, Bothalia 29 (1): 77, 1999 (Perold 1999b). Fossombronia reticulata Steph., Hedwigia 33 (1): 9, 1894 (Stephani 1894a). 419 Fossombronia nigricaulis is possibly conspeciic with Fossombronia australis (Scott and Pike 1984). World checklist of hornworts and liverworts * ** *** ** *** *** *** * *** *** * *** ** *** *** *** ** ** *** *** *** ** 467 Fossombronia rosulata Colenso, Trans. & Proc. New Zealand Inst. 18: 248, 1886 (Colenso 1886b). Fossombronia rudis G.A.M.Scott et D.C.Pike, Beih. Nova Hedwigia 90: 110, 1988 (Scott and Pike 1988a). Fossombronia ruminata Cargill, Phytotaxa 65: 45, 2012 (Cargill et al. 2012). Nom. nov. pro Fossombronia maritima G.A.M.Scott et D.C.Pike, Fl. Austral. Suppl. 21: 114, 2003 (McCarthy 2003), nom. illeg. Fossombronia rupestris G.A.M.Scott et D.C.Pike, Fl. Austral. Suppl. 21: 114, 2003 (McCarthy 2003). Based on: Fossombronia rupestris G.A.M.Scott et D.C.Pike, J. Hattori Bot. Lab. 62: 381, 1987 (Scott and Pike 1987), nom. inval. Fossombronia rwandaensis Perold, Bothalia 28 (1): 45, 1998 (Perold 1998a). Fossombronia scrobiculata G.A.M.Scott et D.C.Pike, J. Hattori Bot. Lab. 56: 341, 1984 (Scott and Pike 1984). Fossombronia spinifolia Steph., Mém. Herb. Boissier 16: 35 (389), 1900 (Stephani 1900a). Fossombronia spinosa Perold, Bothalia 29 (1): 29, 1999 (Perold 1999c). Fossombronia stephanii Schifn. ex Steph., Mém. Herb. Boissier 16: 27 (381), 1900 (Stephani 1900a). Fossombronia straussiana Perold, Bothalia 27 (1): 24, 1997 (Perold 1997a). Fossombronia subsaccata Steph., Sp. Hepat. (Stephani) 6: 75, 1917 (Stephani 1917a). 420 Fossombronia swaziensis Perold, Bothalia 28 (2): 162, 1998 (Perold 1998b). Fossombronia tesselata G.A.M.Scott et D.C.Pike, J. Hattori Bot. Lab. 62: 382, 1987 (Scott and Pike 1987). Fossombronia texana Lindb., Acta Soc. Sci. Fenn. 10: 533, 1875 (Lindberg 1875). Fossombronia truncata G.A.M.Scott et D.C.Pike, Fl. Austral. Suppl. 21: 114, 2003 (McCarthy 2003). Based on: Fossombronia truncata G.A.M.Scott et D.C.Pike, J. Hattori Bot. Lab. 62: 383, 1987 (Scott and Pike 1987), nom. inval. Fossombronia tumida Mitt., J. Linn. Soc., Bot. 16 (91): 193, 1877 (Mitten 1877). Fossombronia valparaisiana Hässel, Beih. Nova Hedwigia 131: 14, 2007 (Hässel and Villagrán 2007). Fossombronia vermiculata G.A.M.Scott et D.C.Pike, J. Hattori Bot. Lab. 56: 345, 1984 (Scott and Pike 1984). Fossombronia wattsii Steph., Sp. Hepat. (Stephani) 6: 75, 1917 (Stephani 1917a). Fossombronia wondraczekii (Corda) Dumort. ex Lindb., Helsingf. Dagbl. 1873 (273, 7 Oct): 2, 1873 (Lindberg 1873c). Bas.: Jungermannia wondraczekii Corda, Deutschl. Fl. (Sturm), Abt. 2, Cryptog.: 30, 1830 (Corda 1830). Fossombronia wrightii Austin, Bot. Bull. (Hanover) 1 (8): 36, 1876 (Austin 1876a). Fossombronia zuurbergensis Perold, Bothalia 31 (1): 25, 2001 (Perold 2001b). 420 Fossombronia subsaccata has a sterile and unidentiiable type specimen (Scott and Pike 1988b). 468 *** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Petalophyllaceae Stotler et Crand.-Stotl. by R. Stotler and B.J. Crandall-Stotler *** Petalophyllum Nees et Gottsche, Nov. Stirp. Pug. 8: 29, 1844 (Lehmann 1844). *** Petalophyllum americanum C.H.Ford et Crand.-Stotl., Novon 12 (3): 335, 2002 (Crandall-Stotler et al. 2002). *** Petalophyllum hodgsoniae C.H.Ford et Crand.-Stotl., Novon 12 (3): 336, 2002 (Crandall-Stotler et al. 2002). *** Petalophyllum indicum Kashyap, J. Indian Bot. Soc. 7: 14, 1928 (Kashyap 1928). *** Petalophyllum preissii Gottsche, Nov. Stirp. Pug. 8: 30, 1844 (Lehmann 1844). *** Petalophyllum ralfsii (Wilson) Nees et Gottsche, Nov. Stirp. Pug. 8: 30, 1844 (Lehmann 1844). Bas.: Jungermannia ralfsii Wilson, Suppl. Engl. Bot. 4: tab. 2874, 1849 (Borrer et al. 1849). *** Sewardiella Kashyap, New Phytol. 14 (1): 5, 1915 (Kashyap 1915). *** Sewardiella tuberifera Kashyap, New Phytol. 14 (1): 5, 1915 (Kashyap 1915). Makinoiineae He-Nygrén, Juslén, Ahonen, Glenny et Piippo *** Makinoaceae Nakai by R. Stotler and B.J. Crandall-Stotler *** Makinoa Miyake, Bot. Mag. (Tokyo) 13 (144): 23, 1899 (Miyake 1899). *** Makinoa crispata (Steph.) Miyake, Bot. Mag. (Tokyo) 13 (144): 23, 1899 (Miyake 1899). Bas.: Pellia crispata Steph., Bull. Herb. Boissier 5 (2): 103, 1897 (Stephani 1897b). Pallaviciniales W.Frey et M.Stech Pallaviciniineae R.M.Schust. *** Hymenophytaceae R.M.Schust. by R. Stotler and B.J. Crandall-Stotler *** Hymenophyton Dumort., Recueil Observ. Jungerm.: 25, 1835 (Dumortier 1835). *** Hymenophyton labellatum (Labill.) Dumort. ex Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 430, 1877 (Trevisan 1877). Bas.: Jungermannia labellata Labill., Nov. Holl. Pl. 2: 109, 1806 (Labillardière 1806). *** Hymenophyton leptopodum (Hook.f. et Taylor) A.Evans, Trans. Connecticut Acad. Arts 8 (16): 274, 1893 (Evans 1893). Bas.: Jungermannia leptopoda Hook.f. et Taylor, London J. Bot. 3: 571, 1844 (Hooker and Taylor 1844d). World checklist of hornworts and liverworts 469 *** Hymenophyton pedicellatum Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n. ser.) 46 (9): 11, 1911 (Stephani 1911b). *** Moerckiaceae K.I.Goebel ex Stotler et Crand.-Stotl. by B.J. Crandall-Stotler and R. Stotler Mamontov et al. (2015) showed that the family Moerckiaceae is heterogeneous and they moved Hattorianthus and Moerckia lotoviana to their new family Cordaeaceae. However, the type of Moerckiaceae (Moerckia hibernica) was not included in the study and only provisionally left in Moerckia. We prefer to follow Crandall-Stotler & Stotler (2007) until M. hibernica is further studied. *** Hattorianthus R.M.Schust. et Inoue, Bull. Natl. Sci. Mus. Tokyo, B 1 (3): 103, 1975 (Schuster and Inoue 1975). *** Hattorianthus erimonus (Steph.) R.M.Schust. et Inoue, Bull. Natl. Sci. Mus. Tokyo, B 1 (3): 106, 1975 (Schuster and Inoue 1975). Bas.: Pallavicinia erimona Steph., Bull. Herb. Boissier 5 (2): 102, 1897 (Stephani 1897b). *** Moerckia Gottsche, Hepat. Eur., Leberm. 13-14: no. 121, 1860 [1861] (Rabenhorst 1860). *** Moerckia blyttii (Mørch) Brockm., Arch. Vereins Freunde Naturgesch. Mecklenburg 17: 190, 1863 (Brockmüller 1863). Bas.: Jungermannia blyttii Mørch, Fl. Danica 12: 6, 1830 (Hornemann 1830). *** Moerckia lotoviana (Nees) Schifn., Österr. Bot. Z. 51 (2): 43, 1901 (Schifner 1901). Bas.: Cordaea lotoviana Nees, Flora 16 (26): 405, 1833 (Nees 1833b). *** Moerckia hibernica (Hook.) Gottsche, Hepat. Eur., Leberm. 13-14: no. 121, 1860 [1861] (Rabenhorst 1860). Bas.: Jungermannia hibernica Hook., Brit. Jungermann.: tab. 78, 1816 (Hooker 1816a). *** Pallaviciniaceae Mig. by R. Stotler and B.J. Crandall-Stotler ** Pallavicinioideae Mig. ex Grolle *** Jensenia Lindb., Not. Sällsk. Fauna Fl. Fenn. Förh. 9: 13, 1868 (Lindberg 1868a). *** Jensenia connivens (Colenso) Grolle, Rev. Bryol. Lichénol. 33 (1/2): 228, 1964 [1965] (Grolle 1964j). Bas.: Symphyogyna connivens Colenso, Trans. & Proc. New Zealand Inst. 20: 254, 1888 (Colenso 1888). 470 * *** *** *** *** *** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Jensenia crassifrons (Steph.) S.Schuette et Stotler, J. Hattori Bot. Lab. 97: 300, 2005 (Schuette and Stotler 2005). Bas.: Pallavicinia crassifrons Steph., Mém. Herb. Boissier 11: 21 (325), 1900 (Stephani 1900c). 421 Jensenia decipiens (Mitt.) Grolle, Rev. Bryol. Lichénol. 33 (1/2): 228, 1964 [1965] (Grolle 1964j). Bas.: Steetzia decipiens Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 123, 1860 [1861] (Mitten 1860c). Jensenia diformis (Nees) Grolle, Rev. Bryol. Lichénol. 33 (1/2): 228, 1964 [1965] (Grolle 1964j). Bas.: Jungermannia diformis Nees, Fl. Bras. (Martius) 1 (1): 329, 1833 (Nees 1833a). Jensenia lorschuetzii Gronde, Proc. Kon. Ned. Akad. Wetensch. C 83 (3): 273, 1980 (Van der Gronde 1980). Jensenia spinosa (Lindenb. et Gottsche) Grolle, Acta Bot. Fenn. 133: 65, 1986 (Grolle and Piippo 1986). Bas.: Symphyogyna spinosa Lindenb. et Gottsche, Syn. Hepat. 5: 786, 1847 (Gottsche et al. 1847). Jensenia wallisii (J.B.Jack et Steph.) Grolle, Rev. Bryol. Lichénol. 33 (1/2): 228, 1964 [1965] (Grolle 1964j). Bas.: Pallavicinia wallisii J.B.Jack et Steph., Hedwigia 31 (1): 23, 1892 (Jack and Stephani 1892). *** Pallavicinia Gray, Nat. Arr. Brit. Pl. 1: 775, 1821 (Gray 1821) nom. conserv. *** Pallavicinia ambigua (Mitt.) Steph., Mém. Herb. Boissier 11: 8 (312), 1900 (Stephani 1900a). Bas.: Steetzia ambigua Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 123, 1860 [1861] (Mitten 1860c). ** Pallavicinia baldwinii (Austin) A.Evans, Trans. Connecticut Acad. Arts 8 (15): 259, 1891 (Evans 1891). Bas.: Steetzia baldwinii Austin, Bull. Torrey Bot. Club 6 (52): 303, 1879 (Austin 1879). * Pallavicinia bipinnata Steph., Sp. Hepat. (Stephani) 6: 62, 1917 (Stephani 1917a). ** Pallavicinia camisassai Gola, Mem. Reale Accad. Sci. Torino (ser. 2) 65 (1): 4, 1916 (Gola 1916). ** Pallavicinia cylindrica (Austin) A.Evans, Trans. Connecticut Acad. Arts 8 (15): 259, 1891 (Evans 1891). Bas.: Steetzia cylindrica Austin, Bull. Torrey Bot. Club 5 (3): 17, 1874 (Austin 1874). ** Pallavicinia himalayensis Schifn., Mém. Herb. Boissier 11: 13 (317), 1900 (Stephani 1900c). *** Pallavicinia indica Schifn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 183, 1898 (Schifner 1898a). *** Pallavicinia levieri Schifn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 184, 1898 (Schifner 1898a). *** Pallavicinia lyellii (Hook.) Gray, Nat. Arr. Brit. Pl. 1: 775, 1821 (Gray 1821). Bas.: Jungermannia lyellii Hook., Brit. Jungermann.: tab. 77, 1816 (Hooker 1816a). ** Pallavicinia pilifera Steph., Hedwigia 30 (6): 271, 1891 (Stephani 1891c). ** Pallavicinia purpurea Steph., Sp. Hepat. (Stephani) 6: 64, 1917 (Stephani 1917a). 421 Jensenia crassifrons is possibly conspeciic with Jensenia diformis. World checklist of hornworts and liverworts 471 ** Pallavicinia ridleyi Steph., Sp. Hepat. (Stephani) 6: 64, 1917 (Stephani 1917a). *** Pallavicinia rubristipa Schifn., Österr. Bot. Z. 56 (1): 24, 1906 (Schifner 1906b). *** Pallavicinia subciliata (Austin) Steph., Mém. Herb. Boissier 11: 9 (313), 1900 (Stephani 1900c). Bas.: Steetzia subciliata Austin, Bull. Torrey Bot. Club 6 (52): 303, 1879 (Austin 1879). *** Pallavicinia xiphoides (Hook.f. et Taylor) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 427, 1877 (Trevisan 1877). Bas.: Jungermannia xiphoides Hook.f. et Taylor, London J. Bot. 3: 569, 1844 (Hooker and Taylor 1844d). *** Podomitrium Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 164, 1855 (Mitten 1855). *** Podomitrium malaccense (Steph.) Campb., Amer. J. Bot. 2 (5): 199, 1915 (Campbell 1915). Bas.: Hymenophyton malaccense Steph., Hedwigia 34 (2): 46, 1895 (Stephani 1895c). *** Podomitrium marginatum (Steph.) Hürl., Bauhinia 4 (1): 78, 1968 [1969] (Hürlimann 1968). Bas.: Hymenophyton marginatum Steph., Sp. Hepat. (Stephani) 6: 61, 1917 (Stephani 1917a). *** Podomitrium phyllanthus (Hook.) Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 164, 1855 (Mitten 1855). Bas.: Jungermannia phyllanthus Hook., Musci Exot. 1: tab. 95, 1818 (Hooker 1818). ** Symphyogynoideae R.M.Schust. ex Grolle ** Greeneothallus Hässel, J. Bryol. 11 (1): 115, 1980 (Hässel 1980). *** Greeneothallus gemmiparus Hässel, J. Bryol. 11 (1): 115, 1980 (Hässel 1980). *** Seppeltia Grolle, J. Hattori Bot. Lab. 60: 276, 1986 (Grolle and Seppelt 1986). *** Seppeltia succuba Grolle, J. Hattori Bot. Lab. 60: 276, 1986 (Grolle and Seppelt 1986). *** Symphyogyna Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 5: 66, 1836 (Nees and Montagne 1836). *** Symphyogyna apiculispina Steph., Biblioth. Bot. 87 (2): 180, 1916 (Stephani 1916a). *** Symphyogyna aspera Steph. ex F.A.McCormick, Bot. Gaz. 58 (5): 403, 1914 (McCormick 1914). ** Symphyogyna atronervia Taylor, London J. Bot. 5: 409, 1846 (Taylor 1846b). ** Symphyogyna bogotensis Steph., Mém. Herb. Boissier 11: 42 (346), 1900 (Stephani 1900c). Based on: Symphyogyna hymenophyllum var. bogotensis Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 181, 1864 (Gottsche 1864), nom. inval. * Symphyogyna boliviensis Steph., Biblioth. Bot. 87 (2): 180, 1916 (Stephani 1916a). *** Symphyogyna brasiliensis Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 5: 67, 1836 (Nees and Montagne 1836). Nom. nov. pro Jungermannia brasiliensis Nees, Enum. Pl. Crypt. Javae: 11, 1830 (Nees 1830), nom. illeg. ** Symphyogyna brasiliensis var. angustior (Gottsche, Lindenb. et Nees) Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 183, 1864 (Gottsche 1864). Bas.: Symphyogyna brasiliensis 472 ** *** *** *** ** *** *** ** ** ** ** ** *** ** *** *** *** ** * *** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) β angustior Gottsche, Lindenb. et Nees, Syn. Hepat. 4: 484, 1846 (Gottsche et al. 1846). Symphyogyna brasiliensis var. subsinuata Schifn., Österr. Akad. Wiss., Math.Naturwiss. Kl., Denkschr. 111: 32, 1964 (Schifner and Arnell 1964). Symphyogyna brongniartii Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 265, 1843 (Montagne 1843). Symphyogyna circinata Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 5: 69, 1836 (Nees and Montagne 1836). Symphyogyna digitisquama Steph., Mém. Herb. Boissier 11: 31 (335), 1900 (Stephani 1900c). Symphyogyna fuscovirens A.Evans, Trans. Connecticut Acad. Arts 28 (6): 320, 1927 (Evans 1927). Symphyogyna hochstetteri Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 5: 68, 1836 (Nees and Montagne 1836). Symphyogyna hymenophyllum (Hook.) Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 5: 68, 1836 (Nees and Montagne 1836). Bas.: Jungermannia hymenophyllum Hook., Musci Exot. 1: tab. 14, 1818 (Hooker 1818). Symphyogyna hymenophyllum var. heterogenum Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 536, 1885 (Spruce 1885). Symphyogyna ignambiensis Hürl., Bauhinia 4 (1): 79, 1968 [1969] (Hürlimann 1968). Symphyogyna interrupta Carrington et Pearson, Proc. Linn. Soc. New South Wales (ser. 2) 2 (4): 1053, 1888 (Carrington and Pearson 1888a). Symphyogyna irregularis Steph., Mém. Herb. Boissier 11: 29 (333), 1900 (Stephani 1900c). Symphyogyna lacerosquama Steph., Sp. Hepat. (Stephani) 6: 67, 1917 (Stephani 1917a). Symphyogyna leptothelia Taylor, London J. Bot. 5: 408, 1846 (Taylor 1846b). Symphyogyna lindmanii A.Evans, Trans. Connecticut Acad. Arts 28 (6): 316, 1927 (Evans 1927). Symphyogyna luetzelburgii Herzog, Repert. Spec. Nov. Regni Veg. 21 (1/7): 22, 1925 (Herzog 1925a). Symphyogyna marginata Steph., Mém. Herb. Boissier 11: 30 (334), 1900 (Stephani 1900c). Symphyogyna mexicana Steph., Rev. Bryol. 36 (6): 140, 1909 (Stephani 1909b). Symphyogyna multilora Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 133, 1914 (Stephani and Watts 1914). Symphyogyna paucidens Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 13, 1911 (Stephani 1911b). 422 Symphyogyna podophylla (hunb.) Nees et Mont., Flora 29 (9): 135, 1846 (Krauss 1846). Bas.: Jungermannia podophylla hunb., Prodr. Pl. Cap. 2: 174, 1800 (hunberg 1800). 422 Symphyogyna paucidens is a doubtful taxon. Hässel and Rubies (2009) could not ind the type specimen. World checklist of hornworts and liverworts * *** ** *** *** ** ** *** *** *** *** * *** ** 473 Symphyogyna purpureolimbata E.A.Hodgs., Trans. Roy. Soc. New Zealand, Bot. 3 (4): 95, 1965 (Hodgson 1965). Symphyogyna rectidens Grolle, Acta Bot. Fenn. 133: 68, 1986 (Grolle and Piippo 1986). Symphyogyna rhodina (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 4: 487, 1846 (Gottsche et al. 1846). Bas.: Jungermannia rhodina Hook.f. et Taylor, London J. Bot. 4: 93, 1845 (Hooker and Taylor 1845). Symphyogyna rubescens Steph., Mém. Herb. Boissier 11: 29 (333), 1900 (Stephani 1900c). Symphyogyna rubritincta A.Evans, Trans. Connecticut Acad. Arts 27 (1): 38, 1925 (Evans 1925). Symphyogyna semi-involucrata Austin, Bull. Torrey Bot. Club 5 (3): 15, 1874 (Austin 1874). Symphyogyna similis Grolle, Acta Bot. Fenn. 133: 70, 1986 (Grolle and Piippo 1986). Symphyogyna sinuata (Sw.) Nees et Mont., Voy. Amér. Mérid., Bot. 7 (1): 61, 1839 (Montagne 1839b). Bas.: Jungermannia sinuata Sw., Prodr. (Swartz): 145, 1788 (Swartz 1788). Symphyogyna subsimplex Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 166, 1855 (Mitten 1855). Symphyogyna tenuinervis (Hook.f. et Taylor) Grolle, J. Hattori Bot. Lab. 61: 253, 1986 [1987] (Grolle 1986a). Bas.: Jungermannia tenuinervis Hook.f. et Taylor, London J. Bot. 3: 570, 1844 (Hooker and Taylor 1844d). Symphyogyna trivittata Spruce, J. Linn. Soc., Bot. 30 (210): 365, 1895 (Gepp 1895b). Symphyogyna ulvoides (Reinw., Blume et Nees) Nees, Syn. Hepat. 4: 487, 1846 (Gottsche et al. 1846). Bas.: Jungermannia ulvoides Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 196, 1824 [1825] (Reinwardt et al. 1824a). 423 Symphyogyna undulata Colenso, Trans. & Proc. New Zealand Inst. 16: 356, 1884 (Colenso 1884). Symphyogyna volkensii Steph., Mém. Herb. Boissier 11: 35 (339), 1900 (Stephani 1900c). ** Symphyogynopsis Grolle, Acta Bot. Fenn. 133: 72, 1986 (Grolle and Piippo 1986). *** Symphyogynopsis gottscheana (Mont. et Nees) Grolle, J. Hattori Bot. Lab. 63: 441, 1987 (Grolle 1987). Bas.: Symphyogyna gottscheana Mont. et Nees, Syn. Hepat. 4: 484, 1846 (Gottsche et al. 1846). ** Xenothallus R.M.Schust., J. Hattori Bot. Lab. 26: 293, 1963 (Schuster 1963b). *** Xenothallus vulcanicola R.M.Schust., J. Hattori Bot. Lab. 26: 293, 1963 (Schuster 1963b). 423 Symphyogyna ulvoides is probably a ilmy fern (Schifner 1900a), but he did not see any type material. 474 ** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Sandeothallaceae R.M.Schust. ** Sandeothallus R.M.Schust., Nova Hedwigia 36: 10, 1982 (Schuster 1982). ** Sandeothallus japonicus (Inoue) Crand.-Stotl. et Stotler, Beih. Nova Hedwigia 131: 58, 2007 (Crandall-Stotler and Stotler 2007). Bas.: Moerckia japonica Inoue, Bull. Natl. Sci. Mus. Tokyo, B 11 (1): 8, 1985 (Inoue 1985). ** Sandeothallus radiculosus (Schifn.) R.M.Schust., Nova Hedwigia 36: 11, 1982 (Schuster 1982). Bas.: Moerckia radiculosa Schifn., Österr. Bot. Z. 51 (2): 48, 1901 (Schifner 1901). Phyllothalliineae R.M.Schust. *** Phyllothalliaceae E.A.Hodgs. ex T.Katag. *** Phyllothallia E.A.Hodgs., Trans. Roy. Soc. New Zealand, Bot. 2 (19): 247, 1964 (Hodgson 1964). *** Phyllothallia fuegiana R.M.Schust., Trans. Brit. Bryol. Soc. 5 (2): 284, 1967 (Schuster 1967d). *** Phyllothallia nivicola E.A.Hodgs., Trans. Roy. Soc. New Zealand, Bot. 2 (19): 247, 1964 (Hodgson 1964). Pelliales He-Nygrén, Juslén, Ahonen, Glenny et Piippo *** Noterocladaceae W.Frey et M.Stech Frey and Stech (2005a) proposed Noterocladaceae as a monogeneric family based on molecular and morphological evidence, which was later corroborated by CrandallStotler et al. (2010). *** Noteroclada Taylor ex Hook.f. et Wilson, London J. Bot. 3: 166, 1844 (Hooker and Wilson 1844). *** Noteroclada conluens Taylor, London J. Bot. 3: 166, 1844 (Hooker and Wilson 1844). * Excluded from the genus Noteroclada longiuscula Colenso, Trans. & Proc. New Zealand Inst. 19: 299, 1887 (Colenso 1887). 424 424 Noteroclada longiuscula is neither a Noteroclada nor a Fossombronia species. he type specimen could not be found (Crandall-Stotler et al. 2010). World checklist of hornworts and liverworts *** 475 Pelliaceae H.Klinggr. Frey and Stech (2005b) recognized Pelliaceae as a monogeneric family, which was later supported by Crandall-Stotler et al. (2010). ** Pellia Raddi, Jungermanniogr. Etrusca: 38, 1818 (Raddi 1818a) nom. conserv. *** subg. Apopellia Grolle, J. Bryol. 12 (3): 427, 1983 (Grolle 1983a). ** Pellia alpicola R.M.Schust. ex L.Söderstr., A.Hagborg et von Konrat, Phytotaxa 76 (3): 39, 2013 (Söderström et al. 2013d). Based on: Pellia endiviifolia subsp. alpicola R.M.Schust., J. Hattori Bot. Lab. 70: 145, 1991 (Schuster 1991b), nom. inval. *** Pellia endiviifolia (Dicks.) Dumort., Recueil Observ. Jungerm.: 27, 1835 (Dumortier 1835). Bas.: Jungermannia endiviifolia Dicks., Fasc. Pl. Crypt. Brit. 4: 19, 1801 (Dickson 1801). ** Pellia megaspora R.M.Schust., J. Bryol. 11 (3): 419, 1981 (Schuster 1981b). *** subg. Pellia ** Pellia appalachiana R.M.Schust., Phytotaxa 76 (3): 39, 2013 (Söderström et al. 2013d). Based on: Pellia appalachiana R.M.Schust., J. Hattori Bot. Lab. 70: 145, 1991 (Schuster 1991b), nom. inval. *** Pellia epiphylla (L.) Corda, Gen. hepat.: 654, 1829 (Corda 1829). Bas.: Jungermannia epiphylla L., Sp. Pl. 1: 1135, 1753 (Linnaeus 1753). ** Pellia epiphylla subsp. borealis (Lorb.) Messe, Bull. Soc. Roy. Bot. Belgique 114 (1): 13, 1981 (Messe 1981). Bas.: Pellia borealis Lorb., Jahrb. Wiss. Bot. 80: 697, 1934 (Lorbeer 1934). *** Pellia neesiana (Gottsche) Limpr., Hedwigia 15 (2): 18, 1876 (Limpricht 1876). Bas.: Pellia epiphylla f. neesiana Gottsche, Hedwigia 6 (5): 69, 1867 (Gottsche 1867). * * * * * Incertae sedis Pellia cordaeana Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 433, 1877 (Trevisan 1877). Pellia crispa P.Kumm., Führer Leberm.: 60, 1875 (Kummer 1875). Pellia gottscheana Kreh, Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 90 (4): 237, 1909 (Kreh 1909). Pellia longifolia P.Kumm., Führer Leberm.: 60, 1875 (Kummer 1875). Pellia undulata P.Kumm., Führer Leberm.: 60, 1875 (Kummer 1875). 476 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) MARCHANTIOPSIDA Cronquist, Takht. et W.Zimm. Blasiidae He-Nygrén, Juslén, Ahonen, Glenny et Piippo Blasiales Stotler et Crand.-Stotl. *** Blasiaceae H.Klinggr. by L. Söderström *** Blasia L., Sp. Pl. 1: 1138, 1753 (Linnaeus 1753). *** Blasia pusilla L., Sp. Pl. 1: 1138, 1753 (Linnaeus 1753). *** Cavicularia Steph., Bull. Herb. Boissier 5 (2): 87, 1897 (Stephani 1897b). *** Cavicularia densa Steph., Bull. Herb. Boissier 5 (2): 87, 1897 (Stephani 1897b). Marchantiidae Engl. Lunulariales H.Klinggr. *** Lunulariaceae H.Klinggr. by D.G. Long *** Lunularia Adans., Fam. Pl. (Adanson) 2: 15, 1763 (Adanson 1763). *** Lunularia cruciata (L.) Dumort. ex Lindb., Not. Sällsk. Fauna Fl. Fenn. Förh. 9: 298, 1868 (Lindberg 1868b). Bas.: Marchantia cruciata L., Sp. Pl. 1: 1137, 1753 (Linnaeus 1753). * Lunularia cruciata subsp. thaxteri (A.Evans et Herzog) R.M.Schust., Hepat. Anthocerotae N. Amer. 6: 91, 1992 (Schuster 1992d). Bas.: Lunularia thaxteri A.Evans et Herzog, Arch. Esc. Fárm. Fac. Ci. Méd. Córdoba 7: 5, 1938 (Herzog and Hosseus 1938). 425 Marchantiales Limpr. *** Aytoniaceae Cavers by D.G. Long he genera of Aytoniaceae follow the treatment of Bischler-Causse et al. (2005) with the exception of Asterella and Mannia which were re-deined by Schill et al. (2010). 425 Lunularia cruciata subsp. thaxteri was synonymized with subsp. cruciata by Boisselier-Dubayle et al. (1995), but their subsp. thaxteri specimen was identiied with some doubt and originated far from the known range. World checklist of hornworts and liverworts 477 *** Asterella P.Beauv., Dict. Sci. Nat. [F. Cuvier] 3: 257, 1805 (Palisot de Beauvois 1805a) nom. conserv. 426 ** subg. Asterella ** sect. Asterella *** Asterella tenella (L.) P.Beauv., Dict. Sci. Nat. [F. Cuvier] 3: 258, 1805 (Palisot de Beauvois 1805a). Bas.: Marchantia tenella L., Sp. Pl. 1: 1137, 1753 (Linnaeus 1753). ** sect. Brachyblepharis (Nees) D.G.Long, J. Bryol. 22 (2): 113, 2000 (Grolle and Long 2000). Bas.: Fimbraria subg. Brachyblepharis Nees, Syn. Hepat. 4: 569, 1846 (Gottsche et al. 1846). *** Asterella abyssinica (Gottsche) Grolle, Explor. Hydrobiol. Lac Bangweolo Luapula: 170, 1972 (Vanden Berghen 1972b). Bas.: Fimbraria abyssinica Gottsche, Syn. Hepat. 4: 569, 1846 (Gottsche et al. 1846). *** Asterella africana (Mont.) Underw. ex A.Evans, Contr. U.S. Natl. Herb. 20: 250, 1920 (Evans 1920). Bas.: Fimbraria africana Mont., Hist. Nat. Îles Canaries 3 (2): 61, 1840 (Montagne 1840b). *** Asterella blumeana (Nees) Kachroo, J. Gauhati India Univ. 3: 130, 1952 (Kachroo 1952). Bas.: Fimbraria blumeana Nees, Syn. Hepat. 4: 564, 1846 (Gottsche et al. 1846). *** Asterella chilensis (Nees et Mont.) A.Evans, Bull. Torrey Bot. Club 46 (12): 469, 1919 (Evans 1919b). Bas.: Fimbraria chilensis Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 9: 41, 1838 (Montagne 1838). *** Asterella cruciata (Steph.) Horik., Hikobia 1 (2): 79, 1951 (Horikawa 1951c). Bas.: Fimbraria cruciata Steph., Sp. Hepat. (Stephani) 6: 12, 1917 (Stephani 1917a). *** Asterella dissoluta (Steph.) Grolle, Wiss. Z. Friedrich-Schiller-Univ. Jena, Math.Naturwiss. Reihe 38 (2): 237, 1989 (Grolle 1989b). Bas.: Fimbraria dissoluta Steph., Planzenw. Ost-Afrikas C: 62, 1895 (Stephani 1895d). *** Asterella dominicensis S.W.Arnell, Bryologist 61 (2): 140, 1958 (Arnell 1958c). *** Asterella khasyana (Grif.) Grolle, Khumbu Himal 1 (4): 267, 1966 (Grolle 1966k). Bas.: Octokepos khasyanus Grif., Not. pl. asiat. 2: 343, 1849 (Griith 1849). *** Asterella leptophylla (Mont.) Grolle, Feddes Repert. 87 (3/4): 246, 1976 (Grolle 1976a). Bas.: Fimbraria leptophylla Mont., Ann. Sci. Nat. Bot. (sér. 2) 18: 16, 1842 (Montagne 1842b). *** Asterella limbata D.G.Long et Grolle, J. Bryol. 18 (2): 287, 1994 (Long and Grolle 1994). ** Asterella shimizuana Inoue, Bull. Natl. Sci. Mus. Tokyo (n.ser.) 10 (3): 361, 1967 (Inoue 1967b). 427 426 Asterella includes Fimbraria and Hypenantron, but a few taxa have neither been transferred nor synonymized. hey are listed in the “Names in genera not currently accepted” section below. 427 Asterella shimizuana is conspeciic with Asterella khasyana in Grolle and Piippo (1984), but it was accepted by Long (2006). 478 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Asterella tenera (Mitt.) R.M.Schust., J. Hattori Bot. Lab. 26: 298, 1963 (Schuster 1963b). Bas.: Fimbraria tenera Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 170, 1855 (Mitten 1855). * Asterella tenerrima (Steph.) H.A.Mill., Phytologia 47 (4): 320, 1981 (Miller 1981). Bas.: Fimbraria tenerrima Steph., Sp. Hepat. (Stephani) 6: 17, 1917 (Stephani 1917a). 428 *** Asterella venosa (Lehm. et Lindenb.) A.Evans, Contr. U.S. Natl. Herb. 20: 286, 1920 (Evans 1920). Bas.: Fimbraria venosa Lehm. et Lindenb., Nov. Stirp. Pug. 4: 29, 1832 (Lehmann 1832). ** subg. Phragmoblepharis Grolle, Feddes Repert. 87 (3/4): 246, 1976 (Grolle 1976a). *** Asterella australis (Hook.f. et Taylor) Verd., Ann. Bryol. 5: 126, 1932 (Verdoorn 1932c). Bas.: Fimbraria australis Hook.f. et Taylor, London J. Bot. 3: 573, 1844 (Hooker and Taylor 1844d). *** Asterella bachmannii (Steph.) S.W.Arnell, Hepat. South Africa: 62, 1963 (Arnell 1963b). Bas.: Fimbraria bachmannii Steph., Hedwigia 33 (1): 7, 1894 (Stephani 1894a). *** Asterella bolanderi (Austin) Underw., Bot. Gaz. 20 (2): 61, 1895 (Underwood 1895). Bas.: Fimbraria bolanderi Austin, Proc. Acad. Nat. Sci. Philadelphia 21: 230, 1869 (Austin 1869). * Asterella caucasica (Steph.) H.Buch, A.Evans et Verd., Ann. Bryol. 10: 8, 1937 [1938] (Buch et al. 1937). Bas.: Fimbraria caucasica Steph., Bull. Herb. Boissier 7 (3): 206 (132), 1899 (Stephani 1899b). 429 *** Asterella conocephala (Steph.) R.M.Schust., J. Hattori Bot. Lab. 26: 298, 1963 (Schuster 1963b). Bas.: Fimbraria conocephala Steph., Bull. Herb. Boissier 7 (3): 205 (131), 1899 (Stephani 1899b). ** Asterella coronata (Steph.) H.A.Mill., Phytologia 47 (4): 319, 1981 (Miller 1981). Bas.: Fimbraria coronata Steph., Sp. Hepat. (Stephani) 6: 12, 1917 (Stephani 1917a). ** Asterella dioica (Steph.) H.A.Mill., Phytologia 47 (4): 319, 1981 (Miller 1981). Bas.: Fimbraria dioica Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 104, 1914 (Stephani and Watts 1914). * Asterella dognyensis H.A.Mill., Phytologia 47 (4): 319, 1981 (Miller 1981). Nom. nov. pro Fimbraria umbonata Steph., Sp. Hepat. (Stephani) 6: 17, 1917 (Stephani 1917a), nom. illeg. 430 *** Asterella drummondii (Taylor) R.M.Schust. ex D.G.Long, J. Bryol. 21 (1): 76, 1999 (Long 1999b). Bas.: Fimbraria drummondii Taylor, London J. Bot. 5: 412, 1846 (Taylor 1846b). *** Asterella echinella (Gottsche) Underw., Bot. Gaz. 20 (2): 62, 1895 (Underwood 1895). Bas.: Fimbraria echinella Gottsche, Mexik. Leverm.: 271, 1863 (Gottsche 1863). 428 Asterella tenerrima is probably conspeciic with Asterella tenera. 429 Asterella caucasica is probably conspeciic with Asterella whiteleggeana (Grolle 1975c, Scott and Bradshaw 1985). 430 Asterella dognyensis is possibly conspeciic with Asterella heterolora. World checklist of hornworts and liverworts 479 *** Asterella elegans (Spreng.) Trevis., Rendiconti Reale Ist. Lombardo Sci. (ser. 2) 7: 785, 1874 (Trevisan 1874). Bas.: Fimbraria elegans Spreng. Syst. Veg. (ed. 16) [Sprengel] 4 (1): 235, 1827 (Sprengel 1827a). *** Asterella heterolora (Steph.) H.A.Mill., Phytologia 47 (4): 320, 1981 (Miller 1981). Bas.: Fimbraria heterolora Steph., Sp. Hepat. (Stephani) 6: 14, 1917 (Stephani 1917a). *** Asterella innovans (Austin) H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 529, 1963 (Miller 1963). Bas.: Marchantia innovans Austin, Bull. Torrey Bot. Club 5 (3): 14, 1874 (Austin 1874). *** Asterella lateralis M.Howe, Bull. Torrey Bot. Club 25 (4): 189, 1898 (Howe 1898b). *** Asterella lindenbergiana (Corda ex Nees) Lindb. ex Arnell, Lebermoosstud. nördl. Norwegen: 2, 1892 (Arnell 1892). Bas.: Fimbraria lindenbergiana Corda ex Nees, Naturgesch. Eur. Leberm. 4: 266, 1838 (Nees 1838a). *** Asterella linearis (Steph.) M.Howe, Bull. Torrey Bot. Club 25 (4): 191, 1898 (Howe 1898b). Bas.: Fimbraria linearis Steph., Bot. Jahrb. Syst. 20 (3): 302, 1895 (Stephani 1895a). *** Asterella longebarbata (Steph.) H.A.Mill., Phytologia 47 (4): 320, 1981 (Miller 1981). Bas.: Fimbraria longebarbata Steph., Hedwigia 28 (3): 156, 1889 (Stephani 1889d). *** Asterella macropoda (Spruce) A.Evans, Bull. Torrey Bot. Club 46 (12): 472, 1919 (Evans 1919b). Bas.: Fimbraria macropoda Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 564, 1885 (Spruce 1885). *** Asterella marginata (Nees) S.W.Arnell, Hepat. South Africa: 63, 1963 (Arnell 1963b). Bas.: Fimbraria marginata Nees, Horae Phys. Berol.: 44, 1820 (Nees 1820). ** Asterella muelleri (Gottsche) R.M.Schust., J. Hattori Bot. Lab. 26: 298, 1963 (Schuster 1963b). Bas.: Fimbraria muelleri Gottsche, Bull. Herb. Boissier 7 (3): 203 (129), 1899 (Stephani 1899b). *** Asterella multilora (Steph.) Kachroo, J. Hattori Bot. Lab. 19: 3, 1958 (Kachroo 1958). Bas.: Fimbraria multilora Steph., Bull. Herb. Boissier 7 (3): 198 (124), 1899 (Stephani 1899b). *** Asterella mussuriensis (Kashyap) Verd., Ann. Bryol. 8: 156, 1935 (Verdoorn 1935). Bas.: Fimbraria mussuriensis Kashyap, J. Bombay Nat. Hist. Soc. 24 (2): 345, 1916 (Kashyap 1916). *** Asterella mussuriensis subsp. crassa (Shimizu et S.Hatt.) D.G.Long, Lindbergia 26 (1): 44, 2001 (Long 2001). Bas.: Asterella crassa Shimizu et S.Hatt., J. Hattori Bot. Lab. 8: 48, 1952 (Shimizu and Hattori 1952). *** Asterella pappii (Gola) Grolle, Feddes Repert. 87 (3/4): 246, 1976 (Grolle 1976a). Bas.: Fimbraria pappii Gola, Ann. Bot. (Rome) 13 (1): 65, 1914 (Gola 1914a). *** Asterella persica (Steph.) M.Howe, Bull. Torrey Bot. Club 25 (4): 191, 1898 (Howe 1898b). Bas.: Fimbraria persica Steph., Hedwigia 33 (1): 7, 1894 (Stephani 1894a). 480 * ** *** ** *** *** *** *** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Asterella preussii (Schifn.) M.Howe, Bull. Torrey Bot. Club 25 (4): 191, 1898 (Howe 1898b). Bas.: Fimbraria preussii Schifn., Bot. Jahrb. Syst. 20 (3): 303, 1895 (Stephani 1895a). 431 Asterella setisquama (Steph.) R.M.Schust., J. Hattori Bot. Lab. 26: 298, 1963 (Schuster 1963b). Bas.: Fimbraria setisquama Steph., Hedwigia 28 (3): 156, 1889 (Stephani 1889d). Asterella syngenesica (Bory) Grolle, Lindbergia 2 (3/4): 230, 1974 (Grolle and Onraedt 1974). Bas.: Marchantia syngenesica Bory, Voy. îles Afrique 2: 95, 1804 (Bory 1804). Asterella tasmanica (Steph.) R.M.Schust., J. Hattori Bot. Lab. 26: 298, 1963 (Schuster 1963b). Bas.: Fimbraria tasmanica Steph., Bull. Herb. Boissier 7 (3): 206 (132), 1899 (Stephani 1899b). Asterella versicolor A.Evans, Contr. U.S. Natl. Herb. 20: 307, 1920 (Evans 1920). Asterella vulcanica (Schifn.) Kachroo et Bapna, J. Indian Bot. Soc. 56 (1): 75, 1977 (Kachroo et al. 1977). Bas.: Hypenantron vulcanicum Schifn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 155, 1898 (Schifner 1898a). Asterella whiteleggeana (Steph.) R.M.Schust., J. Hattori Bot. Lab. 26: 298, 1963 (Schuster 1963b). Bas.: Fimbraria whiteleggeana Steph., Hedwigia 28 (3): 155, 1889 (Stephani 1889d). Asterella wilmsii (Steph.) S.W.Arnell, Hepat. South Africa: 62, 1963 (Arnell 1963b). Bas.: Fimbraria wilmsii Steph., Hedwigia 31 (3): 122, 1892 (Stephani 1892d). ** subg. Saccatae (Grolle) D.G.Long, J. Bryol. 22 (2): 113, 2000 (Grolle and Long 2000). Bas.: Asterella subg. Phragmoblepharis sect. Saccatae Grolle, Feddes Repert. 87 (3/4): 346, 1976 (Grolle 1976a). *** Asterella alpina (Steph.) D.G.Long, J. Hattori Bot. Lab. 93: 9, 2003 (Gradstein et al. 2003). Bas.: Fimbraria alpina Steph., Bull. Herb. Boissier 7 (3): 211 (137), 1899 (Stephani 1899b). *** Asterella grollei D.G.Long, Bryologist 102 (2): 169, 1999 (Long 1999a). *** Asterella muscicola (Steph.) S.W.Arnell, Mitt. Bot. Staatssamml. München 2 (16): 263, 1957 (Arnell 1957c). Bas.: Fimbraria muscicola Steph., Hedwigia 31 (3): 121, 1892 (Stephani 1892d). *** Asterella palmeri (Austin) Underw., Bot. Gaz. 20 (2): 63, 1895 (Underwood 1895). Bas.: Fimbraria palmeri Austin, Bull. Torrey Bot. Club 6 (7): 47, 1875 (Austin 1875c). *** Asterella pringlei Underw., Bot. Gaz. 20 (2): 64, 1895 (Underwood 1895). *** Asterella rugosa A.Evans, Contr. U.S. Natl. Herb. 20: 289, 1920 (Evans 1920). *** Asterella saccata (Wahlenb.) A.Evans, Contr. U.S. Natl. Herb. 20: 276, 1920 (Evans 1920). Bas.: Marchantia saccata Wahlenb., Mag. Neuesten Entdeck. Gesammten Naturk. Ges. Naturf. Freunde Berlin 5: 296, 1811 (Wahlenberg 1811). 431 Asterella preussii was provisionally placed in synonymy with Asterella bachmannii (Wigginton and Grolle 1996). World checklist of hornworts and liverworts 481 ** subg. Wallichianae D.G.Long, Lindbergia 26 (1): 43, 2001 (Long 2001). ** sect. Californicae D.G.Long, J. Hattori Bot. Lab. 97: 257, 2005 (Long 2005). *** Asterella californica (Hampe ex Austin) Underw., Bot. Gaz. 20 (2): 60, 1895 (Underwood 1895). Bas.: Fimbraria californica Hampe ex Austin, Hepat. bor.-amer.: 33, 1873 (Austin 1873). ** sect. Wallichianae D.G.Long, Phytotaxa 173 (1): 87, 2014 (Long et al. 2014). *** Asterella wallichiana (Lehm. et Lindenb.) Grolle, Khumbu Himal 1 (4): 262, 1966 (Grolle 1966k). Bas.: Fimbraria wallichiana Lehm. et Lindenb., Nov. Stirp. Pug. 4: 4, 1832 (Lehmann 1832). ** Cryptomitrium Austin ex Underw., Bull. Illinois State Lab. Nat. Hist. 2 (1): 36, 1884 (Underwood 1884). *** Cryptomitrium himalayense Kashyap, New Phytol. 14 (1): 2, 1915 (Kashyap 1915). *** Cryptomitrium oreades Perold, Bothalia 24 (2): 149, 1994 (Perold 1994). *** Cryptomitrium tenerum (Hook.) Austin ex Underw., Bull. Illinois State Lab. Nat. Hist. 2 (1): 36, 1884 (Underwood 1884). Bas.: Marchantia tenera Hook., Syn. Pl. (Kunth) 1: 45, 1822 (Kunth 1822). *** Mannia Corda, Gen. hepat.: 646, 1829 (Corda 1829) nom. conserv. ** subg. Mannia *** Mannia androgyna (L.) A.Evans, Chron. Bot. 4: 224, 1938 (Evans 1938a). Bas.: Marchantia androgyna L., Sp. Pl. 1: 1138, 1753 (Linnaeus 1753). *** Mannia californica (Gottsche) L.C.Wheeler, Bryologist 37 (5): 88, 1934 [1935] (Wheeler 1934). Bas.: Grimaldia californica Underw., Bot. Gaz. 13 (5): 114, 1888 (Underwood 1888). *** Mannia controversa (Meyl.) D.B.Schill, Edinburgh J. Bot. 65 (1): 36, 2008 (Schill et al. 2008). Bas.: Grimaldia controversa Meyl., Beitr. Kryptogamenl. Schweiz 6 (4): 87, 1924 (Meylan 1924). *** Mannia controversa subsp. asiatica D.B.Schill et D.G.Long, Edinburgh J. Bot. 65 (1): 45, 2008 (Schill et al. 2008). *** Mannia fragrans (Balb.) Frye et L.Clark, Univ. Wash. Publ. Biol. 6 (1): 62, 1937 (Frye and Clark 1937). Bas.: Marchantia fragrans Balb., Mem. Acad. Sci. Turin, Sci. Phys. 15: 76, 1804 (Balbis 1804). ** Mannia fragrans subsp. orientalis R.M.Schust., Hepat. Anthocerotae N. Amer. 6: 201, 1992 (Schuster 1992d). Nom. nov. pro Mannia barbifrons Shimizu et S.Hatt., J. Hattori Bot. Lab. 10: 49, 1953 (Shimizu and Hattori 1953b). * Mannia perssonii Udar et V.Chandra, Canad. J. Bot. 43 (1): 150, 1965 (Udar and Chandra 1965). 432 432 Mannia perssonii is possibly conspeciic with Mannia sibirica. 482 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Mannia sibirica (Müll.Frib.) Frye et L.Clark, Univ. Wash. Publ. Biol. 6 (1): 66, 1937 (Frye and Clark 1937). Bas.: Grimaldia pilosa var. sibirica Müll.Frib., Lebermoose 1 (5): 265, 1907 (Müller 1907a). ** subg. Neesiella (Schifn.) D.B.Schill et D.G.Long, Bryologist 113 (1): 175, 2010 (Schill et al. 2010). Bas.: Neesiella Schifn., Hepat. (Engl.-Prantl): 32, 1893 (Schifner 1893b). *** Mannia gracilis (F.Weber) D.B.Schill et D.G.Long, Bryologist 113 (1): 173, 2010 (Schill et al. 2010). Bas.: Marchantia gracilis F.Weber, Hist. Musc. Hepat. Prodr.: 105, 1815 (Weber 1815). * Mannia hegewaldii Bischl., Fl. Neotrop. Monogr. 97: 184, 2005 (Bischler-Causse et al. 2005). 433 *** Mannia pilosa (Hornem.) Frye et L.Clark, Univ. Wash. Publ. Biol. 6 (1): 64, 1937 (Frye and Clark 1937). Bas.: Marchantia pilosa Hornem., Fl. Danica 8 (24): 7, tab. 1426, 1810 (Hornemann 1810). *** Mannia triandra (Scop.) Grolle, J. Bryol. 8 (4): 487, 1975 (Grolle 1975d). Bas.: Marchantia triandra Scop., Fl. Carniol. (ed. 2) 2: 354, 1772 (Scopoli 1772). * Incertae sedis Mannia paradoxa R.M.Schust., Phytologia 57 (6): 410, 1985 (Schuster 1985b). 434 *** Plagiochasma Lehm., Nov. Stirp. Pug. 4: 13, 1832 (Lehmann 1832) nom. conserv. * Plagiochasma megacarpon (Grif.) Steph., Bull. Herb. Boissier 6 (10): 789 (86), 1898 (Stephani 1898c). Bas.: Antrocephalus megacarpon Grif., Not. pl. asiat. 2: 338, 1849 (Griith 1849). 435 ** subg. Micropylum Bischl., Rev. Bryol. Lichénol. 43 (1): 103, 1977 (Bischler 1977). *** Plagiochasma rupestre (J.R.Forst. et G.Forst.) Steph., Bull. Herb. Boissier 6 (10): 783 (80), 1898 (Stephani 1898c). Bas.: Aytonia rupestris J.R.Forst. et G.Forst., Char. gen. pl., ed. 2: 148, 1776 (Forster and Forster 1776). *** Plagiochasma rupestre var. volkii Bischl., Rev. Bryol. Lichénol. 44 (3): 289, 1978 (Bischler 1978). ** subg. Plagiochasma *** Plagiochasma appendiculatum Lehm. et Lindenb., Nov. Stirp. Pug. 4: 14, 1832 (Lehmann 1832). *** Plagiochasma argentinicum Bischl., Rev. Bryol. Lichénol. 45 (3): 301, 1979 (Bischler 1979a). 433 Mannia hegewaldii is possibly conspeciic with Mannia triandra. 434 Mannia paradoxa is possibly a Reboulia species. 435 Plagiochasma megacarpon may not be a Plagiochasma species. Bischler (1979b) could not ind any specimen and did not know where to refer it. World checklist of hornworts and liverworts 483 *** Plagiochasma beccarianum Steph., Bull. Herb. Boissier 6 (10): 781 (78), 1898 (Stephani 1898c). *** Plagiochasma cordatum Lehm. et Lindenb., Nov. Stirp. Pug. 4: 13, 1832 (Lehmann 1832). *** Plagiochasma crenulatum Gottsche, Mexik. Leverm.: 266, 1863 (Gottsche 1863). *** Plagiochasma cuneatum A.Evans, Amer. J. Bot. 19 (7): 627, 1932 (Evans 1932a). *** Plagiochasma eximium (Schifn.) Steph., Bull. Herb. Boissier 6 (10): 781 (78), 1898 (Stephani 1898c). Bas.: Aytonia eximia Schifn., Bot. Jahrb. Syst. 20 (3): 300, 1895 (Stephani 1895a). *** Plagiochasma intermedium Lindenb. et Gottsche, Syn. Hepat. 4: 513, 1846 (Gottsche et al. 1846). *** Plagiochasma jamaicense (Haynes) A.Evans, Bull. Torrey Bot. Club 42 (5): 292, 1915 (Evans 1915). Bas.: Aytonia jamaicensis Haynes, Bull. Torrey Bot. Club 34 (2): 58, 1907 (Haynes 1907). *** Plagiochasma japonicum (Steph.) C.Massal., Hepat. Shen-si: 47, 1897 (Massalongo 1897). Bas.: Aytonia japonica Steph., Bull. Herb. Boissier 5 (2): 84, 1897 (Stephani 1897b). *** Plagiochasma landii A.Evans, Bull. Torrey Bot. Club 42 (5): 298, 1915 (Evans 1915). *** Plagiochasma microcephalum (Steph.) Steph., Bull. Herb. Boissier 6 (10): 781 (78), 1898 (Stephani 1898c). Bas.: Aytonia microcephala Steph., Bot. Jahrb. Syst. 20 (3): 301, 1895 (Stephani 1895a). *** Plagiochasma microcephalum var. tunesicum Bischl., Rev. Bryol. Lichénol. 44 (3): 247, 1978 (Bischler 1978). *** Plagiochasma muenchianum Steph., Sp. Hepat. (Stephani) 6: 9, 1917 (Stephani 1917a). *** Plagiochasma pterospermum C.Massal., Hepat. Shen-si: 46, 1897 (Massalongo 1897). *** Plagiochasma wrightii Sull., Musc. Hepat. U.S.: 688, 1856 (Sullivant 1856). Incertae sedis ** Plagiochasma udarii A.Alam et S.C.Srivast., Indian J. Forest. 32 (4): 631, 2009 (Alam and Srivastava 2009). *** Reboulia Raddi, Opusc. Sci. 2 (6): 357, 1818 (Raddi 1818b) nom. conserv. *** Reboulia hemisphaerica (L.) Raddi, Opusc. Sci. 2 (6): 357, 1818 (Raddi 1818b). Bas.: Marchantia hemisphaerica L., Sp. Pl. 1: 1138, 1753 (Linnaeus 1753). ** Reboulia hemisphaerica subsp. acrogyna (R.M.Schust.) R.M.Schust., Hepat. Anthocerotae N. Amer. 6: 168, 1992 (Schuster 1992d). Bas.: Asterella bolanderi subsp. acrogyna R.M.Schust., Phytologia 57 (6): 410, 1985 (Schuster 1985b). ** Reboulia hemisphaerica subsp. australis R.M.Schust., Phytologia 56 (7): 460, 1985 (Schuster 1985c). ** Reboulia hemisphaerica subsp. dioica R.M.Schust., Phytologia 56 (7): 462, 1985 (Schuster 1985c). ** Reboulia hemisphaerica var. issisquama Herzog, Symb. Sin. 5: 5, 1930 (Nicholson et al. 1930). 484 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Reboulia hemisphaerica subsp. orientalis R.M.Schust., Phytologia 56 (7): 461, 1985 (Schuster 1985c). ** Reboulia hemisphaerica var. turkestanica C.E.O.Jensen ex Herzog, Symb. Sin. 5: 5, 1930 (Nicholson et al. 1930). *** Cleveaceae Cavers by D.G. Long he genera of Cleveaceae were re-deined by Rubasinghe et al. (2011a) based on molecular evidence. *** Athalamia Falc., Ann. Mag. Nat. Hist. (ser. 2) 1 (5): 375, 1848 (Anonymous 1848). * Athalamia dioica Kashyap, J. Bombay Nat. Hist. Soc. 24 (2): 348, 1916 (Kashyap 1916). *** Athalamia pinguis Falc., Ann. Mag. Nat. Hist. (ser. 2) 1 (5): 375, 1848 (Anonymous 1848). * Athalamia pulcherrima (Steph.) S.Hatt., J. Hattori Bot. Lab. 12: 54, 1954 (Shimizu and Hattori 1954). Bas.: Clevea pulcherrima Steph., Bot. Jahrb. Syst. 20 (3): 303, 1895 (Stephani 1895a). *** Clevea Lindb., Not. Sällsk. Fauna Fl. Fenn. Förh. 9: 289, 1868 (Lindberg 1868b). *** Clevea hyalina (Sommerf.) Lindb., Not. Sällsk. Fauna Fl. Fenn. Förh. 9: 291, 1868 (Lindberg 1868b). Bas.: Marchantia hyalina Sommerf., Mag. Naturvidensk. 11 (2): 234, 1833 (Sommerfeldt 1833). * Clevea hyalina var. californica M.Howe, Mem. Torrey Bot. Club 7: 38, 1899 (Howe 1899). * Clevea pedicellata (Grif.) Lindb., Acta Soc. Fauna Fl. Fenn. 2 (3): 11, 1882 (Lindberg 1882). Bas.: Plagiochasma pedicellatum Grif., Not. pl. asiat. 2: 331, 1849 (Griith 1849). *** Clevea pusilla (Steph.) Rubas. et D.G.Long, J. Bryol. 33 (2): 167, 2011 (Rubasinghe et al. 2011b). Bas.: Gollaniella pusilla Steph., Hedwigia 44 (2): 74, 1905 (Stephani 1905h). *** Clevea spathysii (Lindenb.) Müll.Frib., Hedwigia 79 (1/2): 75, 1940 (Müller 1940). Bas.: Marchantia spathysii Lindenb., Syn. hepat. eur: 104, 1829 (Lindenberg 1829). *** Peltolepis Lindb., Morgonbladet (Helsinki) 1876 (106, 9 May): 1, 1876 (Elfving 1876). ** Peltolepis japonica (Shimizu et S.Hatt.) S.Hatt., J. Hattori Bot. Lab. 14: 103, 1955 (Shimizu and Hattori 1955). Bas.: Peltolepis quadrata var. japonica Shimizu et S.Hatt., J. Hattori Bot. Lab. 12: 69, 1954 (Shimizu and Hattori 1954). *** Peltolepis quadrata (Saut.) Müll.Frib., Hedwigia 79 (1/2): 74, 1940 (Müller 1940). Bas.: Sauteria quadrata Saut., Flora 43 (22): 351, 1860 (Sauter 1860). World checklist of hornworts and liverworts 485 *** Sauteria Nees, Naturgesch. Eur. Leberm. 4: 139, 1838 (Nees 1838a). ** sect. Sauchia (Kashyap) R.M.Schust., Phytologia 57 (6): 411, 1985 (Schuster 1985b). Bas.: Sauchia Kashyap, J. Bombay Nat. Hist. Soc. 24 (2): 347, 1916 (Kashyap 1916). * Sauteria japonica (Shimizu et S.Hatt.) S.Hatt., J. Hattori Bot. Lab. 12: 62, 1954 (Shimizu and Hattori 1954). Bas.: Sauchia japonica Shimizu et S.Hatt., J. Hattori Bot. Lab. 9: 32, 1953 (Shimizu and Hattori 1953a). 436 *** Sauteria spongiosa (Kashyap) S.Hatt., J. Hattori Bot. Lab. 12: 62, 1954 (Shimizu and Hattori 1954). Bas.: Sauchia spongiosa Kashyap, J. Bombay Nat. Hist. Soc. 24 (2): 347, 1916 (Kashyap 1916). ** sect. Sauteria *** Sauteria alpina (Nees) Nees, Naturgesch. Eur. Leberm. 4: 143, 1838 (Nees 1838a). Bas.: Lunularia alpina Nees, Flora 13 (25): 399, 1830 (Nees and Bischof 1830). * Sauteria inlata C.Gao et K.C.Chang, Acta Bot. Yunnan. 3 (4): 389, 1981 (Gao et al. 1981). 437 * Incertae sedis Sauteria chilensis (Lindenb.) Grolle, J. Hattori Bot. Lab. 58: 200, 1985 (Grolle 1985b). Bas.: Grimaldia chilensis Lindenb., Voy. Amér. Mérid. 7 (2): 53, 1839 (Montagne 1839a). Sauteria crassipes Austin, Proc. Acad. Nat. Sci. Philadelphia 21: 229, 1869 (Austin 1869). Sauteria nyikaensis Perold, Bothalia 33 (2): 167, 2003 (Perold 2003). *** Conocephalaceae Müll.Frib. ex Grolle * * by D.G. Long *** Conocephalum Hill, Gener. Nat. Hist. 2 Hist. pl. (ed. 2): 118, 1773 (Hill 1773) nom. conserv. ** subg. Conocephalum *** Conocephalum conicum (L.) Dumort., Commentat. Bot. (Dumortier): 115, 1822 (Dumortier 1822). Bas.: Marchantia conica L., Sp. Pl. 1: 1138, 1753 (Linnaeus 1753). *** Conocephalum salebrosum Szweyk., Buczk. et Odrzyk., Pl. Syst. Evol. 253 (1/4): 146, 2005 (Szweykowski et al. 2005). 436 Sauteria japonica is possibly conspeciic with Sauteria spongiosa. 437 Sauteria inlata is possibly conspeciic with Sauteria alpina (Schuster 1992d). 486 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** subg. Sandea (Lindb.) Inoue, Ill. Jap. Hep. 2: 192, 1976 (Inoue 1976a). Bas.: Sandea Lindb., Acta Soc. Fauna Fl. Fenn. 2 (5): 3, 1884 (Lindberg 1884). *** Conocephalum japonicum (hunb.) Grolle, J. Hattori Bot. Lab. 55: 501, 1984 (Grolle 1984a). Bas.: Lichen japonicus hunb., Fl. Jap. (hunberg): 344, 1784 (hunberg 1784). ** Corsiniaceae Engl. by D.G. Long ** Corsinioideae Schiffn. *** Corsinia Raddi, Opusc. Sci. 2 (6): 354, 1818 (Raddi 1818b). *** Corsinia coriandrina (Spreng.) Lindb., Hepaticol. Utveckl.: 30, 1877 (Lindberg 1877c). Bas.: Riccia coriandrina Spreng. Anleit. Kenntn. Gew. 3: 320, 1804 (Sprengel 1804). *** Cronisioideae R.M.Schust. *** Cronisia Berk., Introd. crypt. bot.: 434, 1857 (Berkeley 1857). *** Cronisia imbriata (Nees) Whittem. et Bischl., Cryptog. Bryol. 22 (3): 170, 2001 (Bischler and Whittemore 2001). Bas.: Riccia imbriata Nees, Fl. Bras. (Martius) 1 (1): 301, 1833 (Nees 1833a). *** Cronisia weddellii (Mont.) Grolle, J. Bryol. 9 (4): 532, 1977 [1978] (Grolle 1977a). Bas.: Boschia weddellii Mont., Ann. Sci. Nat. Bot. (sér. 4) 5: 352, 1856 (Montagne 1856c). *** Cyathodiaceae Stotler et Crand.-Stotl. by D.G. Long *** Cyathodium Kunze, Nov. Stirp. Pug. 6: 17, 1834 (Lehmann 1834). *** Cyathodium aureonitens (Grif.) Mitten, J. Linn. Soc., Bot. 22 (146): 327 (Mitten 1886b). Bas.: Synhymenium aureonitens Grif., Not. pl. asiat. 2: 344, 1849 (Grifith 1849). *** Cyathodium bischlerianum N.Salazar, Bryologist 104 (1): 141, 2001 (Salazar 2001). *** Cyathodium cavernarum Kunze, Nov. Stirp. Pug. 6: 18, 1834 (Lehmann 1834). ** Cyathodium denticulatum Udar et S.C.Srivast., Geophytology 1 (2): 166, 1971 (Udar and Srivastava 1971). World checklist of hornworts and liverworts 487 *** Cyathodium foetidissimum Schifn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 154, 1898 (Schifner 1898a). *** Cyathodium indicum Udar et D.K.Singh, J. Bryol. 10 (2): 139, 1978 [1979] (Udar and Singh 1978). *** Cyathodium mehranum D.K.Singh, Misc. Bryol. Lichenol. 9 (8): 173, 1983 (Singh 1983b). *** Cyathodium smaragdinum Schifn., Ann. Jard. Bot. Buitenzorg, suppl. 3: 480, 1910 (Schifner 1910c). *** Cyathodium spruceanum Prosk., Bryologist 54 (4): 243, 1951 [1952] (Proskauer 1951b). * Cyathodium spurium (Dicks.) Lindb. ex Braithw., J. Bot. 16: 55, 1878 (Braithwaite 1878). Bas.: Riccia spuria Dicks., Fasc. Pl. Crypt. Brit. 4: 20, 1801 (Dickson 1801). 438 *** Cyathodium steerei Hässel, Rev. Bryol. Lichénol. 30 (3/4): 223, 1961 (Hässel 1961). *** Cyathodium tuberculatum Udar et D.K.Singh, Bryologist 79 (2): 235, 1976 (Udar and Singh 1976). *** Cyathodium tuberosum Kashyap, New Phytol. 13 (6/7): 210, 1914 (Kashyap 1914a). *** Dumortieraceae D.G.Long by D.G. Long *** Dumortiera Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 410, 1824 [1825] (Reinwardt et al. 1824b). *** Dumortiera hirsuta (Sw.) Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 410, 1824 [1825] (Reinwardt et al. 1824b). Bas.: Marchantia hirsuta Sw., Prodr. (Swartz): 145, 1788 (Swartz 1788). 439 * Dumortiera hirsuta subsp. nepalensis (Taylor) R.M.Schust., Hepat. Anthocerotae N. Amer. 6: 386, 1992 (Schuster 1992d). Bas.: Hygrophila nepalensis Taylor, Trans. Linn. Soc. London 17 (3): 392, 1836 (Taylor 1836b). * Dumortiera hirsuta subsp. tatunoi Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 6: 38, 1951 (Horikawa 1951a). *** Exormothecaceae Müll.Frib. ex Grolle by D.G. Long *** Aitchisoniella Kashyap, New Phytol. 13 (6/7): 219, 1914 (Kashyap 1914a). 438 Cyathodium spurium may be conspeciic with Cyathodium cavernarum (Braithwaite 1878). he type specimen is from Scotland, but it is probably mislabelled as no Cyathodium is known from boreal Europe. 439 Dumortiera hirsuta is a species complex (Forrest et al. 2011). 488 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Aitchisoniella himalayensis Kashyap, New Phytol. 13 (6/7): 219, 1914 (Kashyap 1914a). *** Exormotheca Mitt., Nat. hist. Azores: 325, 1870 (Mitten 1870). ** subg. Corbierella (Douin et Trab.) Schifn., Hedwigia 81 (1/2): 71, 1942 (Schifner 1942). Bas.: Corbierella Douin et Trab., Rev. Gén. Bot. 31: 326, 1919 (Douin and Trabut 1919). *** Exormotheca bischlerae Furuki et Higuchi, Cryptog. Bryol. 27 (1): 98, 2006 (Furuki and Higuchi 2006). *** Exormotheca holstii Steph., Bull. Herb. Boissier 7 (3): 219 (145), 1899 (Stephani 1899b). *** Exormotheca welwitschii Steph., Bull. Herb. Boissier 7 (3): 220 (146), 1899 (Stephani 1899b). ** subg. Exormotheca *** Exormotheca pustulosa Mitt., Nat. hist. Azores: 326, 1870 (Mitten 1870). Incertae sedis *** Exormotheca bulbigena Bornefeld, O.H.Volk et R.Wolf, Bothalia 26 (2): 159, 1996 (Bornefeld et al. 1996). *** Exormotheca ceylonensis Meijer, J. Hattori Bot. Lab. 16: 72, 1956 (Meijer 1956). * Exormotheca gollanii Steph., Sp. Hepat. (Stephani) 6: 18, 1917 (Stephani 1917a). *** Exormotheca tuberifera Kashyap, New Phytol. 13 (9): 309, 1914 (Kashyap 1914b). *** Stephensoniella Kashyap, New Phytol. 13 (9): 312, 1914 (Kashyap 1914b). *** Stephensoniella brevipedunculata Kashyap, New Phytol. 13 (9): 312, 1914 (Kashyap 1914b). *** Marchantiaceae Lindl. by D.G. Long he treatment of Marchantiaceae is mainly following Bischler (1984, 1989b) and Bischler-Causse (1993). ** Bucegioideae R.M.Schust. *** Bucegia Radian, Bull. Herb. Inst. Bot. Bucarest 3-4: 3, 1903 (Radian 1903). *** Bucegia romanica Radian, Bull. Herb. Inst. Bot. Bucarest 3-4: 4, 1903 (Radian 1903). World checklist of hornworts and liverworts ** 489 Marchantioideae Schiffn. *** Marchantia L., Sp. Pl. 1: 1137, 1753 (Linnaeus 1753). ** subg. Chlamidium (Corda) Bischl., Cryptog. Bryol. Lichénol. 3 (4): 362, 1982 (Bischler 1982). Bas.: Chlamidium Corda, Gen. hepat.: 647, 1829 (Corda 1829). *** Marchantia breviloba A.Evans, Trans. Connecticut Acad. Arts 21 (3): 265, 1917 (Evans 1917a). *** Marchantia inlexa Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 9: 43, 1838 (Montagne 1838). ** sect. Chlamidium (Corda) Nees, Naturgesch. Eur. Leberm. 4: 101, 1838 (Nees 1838a). Bas.: Chlamidium Corda, Gen. hepat.: 647, 1829 (Corda 1829). *** Marchantia chenopoda L., Sp. Pl. 1: 1137, 1753 (Linnaeus 1753). *** Marchantia crenata Austin, Bull. Torrey Bot. Club 5 (3): 14, 1874 (Austin 1874). *** Marchantia foliacea Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 168, 1855 (Mitten 1855). *** Marchantia formosana Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 121, 1934 (Horikawa 1934). *** Marchantia globosa Brid., Hist. Musc. Hepat. Prodr.: 102, 1815 (Weber 1815). *** Marchantia hexaptera Reichardt, Verh. K.K. Zool.-Bot. Ges. Wien 16: 957, 1866 (Reichardt 1866). *** Marchantia linearis Lehm. et Lindenb., Nov. Stirp. Pug. 4: 8, 1832 (Lehmann 1832). *** Marchantia miqueliana Lehm., Nov. Stirp. Pug. 10: 20, 1857 (Lehmann 1857). *** Marchantia novoguineensis Bischl., Bryophyt. Biblioth. 38: 130, 1989 (Bischler 1989b). *** Marchantia pappeana Lehm., Nov. Stirp. Pug. 10: 21, 1857 (Lehmann 1857). ** Marchantia pappeana subsp. robusta (Steph.) Bischl., Bryophyt. Biblioth. 45: 91, 1993 (Bischler-Causse 1993). Bas.: Marchantia robusta Steph., Candollea 14: 111, 1953 (Bonner 1953b). *** Marchantia pileata Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 169, 1855 (Mitten 1855). *** Marchantia pinnata Steph., Candollea 14: 109, 1953 (Bonner 1953b). *** Marchantia rubribarba Steph., Bull. Herb. Boissier 7 (5): 400 (172), 1899 (Stephani 1899c). *** Marchantia vitiensis Steph., Bull. Herb. Boissier 7 (7): 520 (182), 1899 (Stephani 1899d). ** sect. Paleaceae Bischl., Bryophyt. Biblioth. 38: 90, 1989 (Bischler 1989b). *** Marchantia paleacea Bertol., Opusc. Sci. 1: 242, 1817 (Bertoloni 1817). *** Marchantia paleacea subsp. diptera (Nees et Mont.) Inoue, J. Jap. Bot. 64 (7): 194, 1989 (Inoue 1989c). Bas.: Marchantia diptera Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 243, 1843 (Montagne 1843). 490 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** sect. Papillatae Bischl., Cryptog. Bryol. Lichénol. 10 (1): 69, 1989 (Bischler 1989a). *** Marchantia debilis K.I.Goebel, Organogr. Pl., ed. 2, 2 (1): 901, 1915 (Goebel 1915). *** Marchantia emarginata Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 192, 1824 [1825] (Reinwardt et al. 1824a). *** Marchantia emarginata subsp. lecordiana (Steph.) Bischl., Cryptog. Bryol. Lichénol. 10 (1): 78, 1989 (Bischler 1989a). Bas.: Marchantia lecordiana Steph., Bull. Herb. Boissier 7 (7): 525 (187), 1899 (Stephani 1899d). *** Marchantia emarginata subsp. tosana (Steph.) Bischl., Cryptog. Bryol. Lichénol. 10 (1): 77, 1989 (Bischler 1989a). Bas.: Marchantia tosana Steph., Bull. Herb. Boissier 5 (2): 99, 1897 (Stephani 1897b). *** Marchantia papillata Raddi, Critt. Brasil.: 20, 1822 (Raddi 1822). *** Marchantia papillata subsp. grossibarba (Steph.) Bischl., Cryptog. Bryol. Lichénol. 10 (1): 78, 1989 (Bischler 1989a). Bas.: Marchantia grossibarba Steph., Mém. Soc. Nat. Sci. Nat. Math. Cherbourg 29: 221, 1894 (Stephani 1894b). ** subg. Marchantia *** Marchantia plicata Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 9: 43, 1838 (Montagne 1838). ** sect. Berteroanae R.M.Schust., Hepat. Anthocerotae N. Amer. 6: 310, 1992 (Schuster 1992d). *** Marchantia berteroana Lehm. et Lindenb., Nov. Stirp. Pug. 6: 21, 1834 (Lehmann 1834). ** sect. Marchantia *** Marchantia polymorpha L., Sp. Pl. 1: 1137, 1753 (Linnaeus 1753). *** Marchantia polymorpha subsp. montivagans Bischl. et Boissel.-Dub., J. Bryol. 16 (3): 364, 1991 (Bischler-Causse and Boisselier-Dubayle 1991). *** Marchantia polymorpha subsp. ruderalis Bischl. et Boissel.-Dub., J. Bryol. 16 (3): 364, 1991 (Bischler-Causse and Boisselier-Dubayle 1991). ** subg. Protomarchantia R.M.Schust., Phytologia 57 (6): 410, 1985 (Schuster 1985b). ** sect. Protomarchantia (R.M.Schust.) L.Söderstr., Phytotaxa 202 (1): 69, 2015 (Söderström et al. 2015c). Bas.: Marchantia subg. Protomarchantia R.M.Schust., Phytologia 57 (6): 410, 1985 (Schuster 1985b). *** Marchantia acaulis Steph., Bull. Herb. Boissier 7 (7): 533 (195), 1899 (Stephani 1899d). *** Marchantia antiqua Steph., Candollea 14: 103, 1953 (Bonner 1953b). *** Marchantia carrii Bischl., Bryophyt. Biblioth. 38: 256, 1989 (Bischler 1989b). *** Marchantia geminata Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 194, 1824 [1825] (Reinwardt et al. 1824a). World checklist of hornworts and liverworts 491 *** Marchantia hartlessiana Steph., Candollea 14: 107, 1953 (Bonner 1953b). *** Marchantia macropora Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 169, 1855 (Mitten 1855). *** Marchantia philippinensis Bischl., Bryophyt. Biblioth. 38: 245, 1989 (Bischler 1989b). *** Marchantia solomonensis Bischl., Bryophyt. Biblioth. 38: 281, 1989 (Bischler 1989b). *** Marchantia streimannii Bischl., Bryophyt. Biblioth. 38: 250, 1989 (Bischler 1989b). *** Marchantia subintegra Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 125, 1860 [1861] (Mitten 1860c). *** Marchantia treubii Schifn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 160, 1898 (Schifner 1898a). *** Marchantia wallisii J.B.Jack et Steph., Bull. Herb. Boissier 7 (7): 520 (182), 1899 (Stephani 1899d). ** sect. Subgeminatae Bischl., Bryophyt. Biblioth. 38: 219, 1989 (Bischler 1989b). *** Marchantia subgeminata Steph., Bull. Herb. Boissier 7 (7): 530 (192), 1899 (Stephani 1899d). * * * * * * * * Incertae sedis Marchantia assamica Grif., Not. pl. asiat. 2: 327, 1849 (Griith 1849). 440 Marchantia balboi Gola, Mem. Reale Accad. Sci. Torino (ser. 2) 65 (1): 2, 1916 (Gola 1916). 441 Marchantia balboi var. acutisquamata Gerola, Lav. Bot. Ist. Bot. Univ. Padova 12: 472, 1947 (Gerola 1947). 442 Marchantia cagnii Gola, Ann. Bot. (Rome) 6 (2): 271, 1907 (Gola 1907). 443 Marchantia cengiana Gerola, Lav. Bot. Ist. Bot. Univ. Padova 12: 473, 1947 (Gerola 1947). 444 Marchantia friedrichsthaliana Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 438, 1877 (Trevisan 1877). Marchantia keniae Gola, Mem. Reale Accad. Sci. Torino (ser. 2) 65 (1): 3, 1916 (Gola 1916). 445 Marchantia papyracea Gola, Ann. Bot. (Rome) 6 (2): 271, 1907 (Gola 1907). 446 440 Marchantia assamica is a doubtful taxon. Bischler (1989b) could not ind the type specimen. 441 Marchantia balboi is a doubtful taxon. Bischler-Causse (1993) could not ind the type specimen. 442 Marchantia balboi var. acutisquamata is a doubtful taxon, Bischler-Causse (1993) could not ind the type specimen and did not know what it is. 443 Marchantia cagnii is a doubtful taxon. Bischler-Causse (1993) could not ind the type specimen, but the description points to Marchantia paleacea. 444 Marchantia cengiana is a doubtful taxon. Bischler-Causse (1993) could not ind the type specimen, but the description points to Marchantia pappeana. 445 Marchantia keniae is a doubtful taxon. Bischler-Causse (1993) could not ind the type specimen, but the description points to Marchantia pappeana. 446 Marchantia papyracea is a doubtful taxon. Bischler-Causse (1993) could not ind the type specimen, but the description points to Marchantia pappeana. 492 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) * Marchantia quadriloba Steph., Candollea 14: 110, 1953 (Bonner 1953b). * Marchantia sellae Gola, Ann. Bot. (Rome) 6 (2): 271, 1907 (Gola 1907). 447 ** Marchantia stoloniscyphulus (C.Gao et K.C.Chang) Piippo, J. Hattori Bot. Lab. 68: 134, 1990 (Piippo 1990). Bas.: Marchantiopsis stoloniscyphulus C.Gao et K.C.Chang, Bull. Bot. Res., Harbin 2 (4): 114, 1982 (Gao and Chang 1982). * Marchantia trilocularis Roth, Tent. Fl. Germ. 1: 487, 1788 (Roth 1788). 448 * Marchantia tusui Gola, Mem. Reale Accad. Sci. Torino (ser. 2) 65 (1): 3, 1916 (Gola 1916). 449 ** Preissia Corda, Gen. hepat.: 647, 1829 (Corda 1829). *** Preissia quadrata (Scop.) Nees, Naturgesch. Eur. Leberm. 4: 135, 1838 (Nees 1838a). Bas.: Marchantia quadrata Scop., Fl. Carniol. (ed. 2) 2: 355, 1772 (Scopoli 1772). ** Preissia quadrata subsp. hyperborea R.M.Schust., Phytologia 57 (6): 410, 1985 (Schuster 1985b). *** Monocleaceae A.B.Frank by D.G. Long *** *** *** *** Monoclea Hook., Musci Exot. 2: tab. clxxiv, 1820 (Hooker 1820). Monoclea forsteri Hook., Musci Exot. 2: tab. clxxiv, 1820 (Hooker 1820). Monoclea gottschei Lindb., Rev. Bryol. 13 (6): 102, 1886 (Lindberg 1886). Monoclea gottschei subsp. elongata Gradst. et Mues, Pl. Syst. Evol. 180 (1/2): 133, 1992 (Gradstein et al. 1992). *** Monosoleniaceae Inoue by D.G.Long *** Monosolenium Grif., Not. pl. asiat. 2: 341, 1849 (Griith 1849). *** Monosolenium tenerum Grif., Not. pl. asiat. 2: 341, 1849 (Griith 1849). 447 Marchantia sellae is a doubtful taxon. Bischler-Causse (1993) could not ind the type specimen, but the description points to Marchantia pappeana or Marchantia paleacea. 448 Marchantia trilocularis is a doubtful taxon which Bischler-Causse (1993) could not place. he name may have priority once its identity is determined. 449 Marchantia tusui is a doubtful taxon. Bischler-Causse (1993) could not ind the type specimen, but the description points to Marchantia pappeana. World checklist of hornworts and liverworts *** 493 Oxymitraceae Müll.Frib. ex Grolle by D.G. Long *** Oxymitra Bisch. ex Lindenb., Syn. hepat. eur: 124, 1829 (Lindenberg 1829). *** Oxymitra cristata Garside, Bothalia 23 (2): 211, 1993 (Perold 1993). Based on: Oxymitra cristata Garside, J. S. African Bot. 24: 83, 1958 (Garside 1958), nom. inval. *** Oxymitra incrassata (Brot.) Sérgio et Sim-Sim, J. Bryol. 15 (4): 662, 1989 (Sérgio and Sim-Sim 1989). Bas.: Riccia incrassata Brot., Fl. lusit. 2: 428, 1804 [1805] (Brotero 1804). *** Ricciaceae Rchb. by R. Stotler, B.J. Crandall-Stotler and D.C. Cargill ** Riccia L., Sp. Pl. 1: 1138, 1753 (Linnaeus 1753) nom. conserv. ** subg. Chartaceae Perold, Bothalia 16 (1): 29, 1986 (Volk and Perold 1986c). *** Riccia schelpei O.H.Volk et Perold, Bothalia 16 (1): 29, 1986 (Volk and Perold 1986c). ** subg. Leptoriccia R.M.Schust., Phytologia 56 (2): 72, 1984 (Schuster 1984). *** Riccia membranacea Gottsche et Lindenb., Syn. Hepat. 4: 608, 1846 (Gottsche et al. 1846). ** subg. Riccia *** Riccia albida Sull. ex Austin, Proc. Acad. Nat. Sci. Philadelphia 21: 231, 1869 (Austin 1869). *** Riccia albopunctata Jovet-Ast, Cryptog. Bryol. Lichénol. 12 (3): 237, 1991 (JovetAst 1991). *** Riccia australis Steph., Bull. Herb. Boissier 6 (4): 337 (29), 1898 (Stephani 1898a). *** Riccia boliviensis Jovet-Ast, Cryptog. Bryol. Lichénol. 12 (3): 242, 1991 (Jovet-Ast 1991). *** Riccia brasiliensis Schifn., Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 6, 1964 (Schifner and Arnell 1964). *** Riccia breutelii Hampe, Bull. Herb. Boissier 6 (4): 325 (17), 1898 (Stephani 1898a). *** Riccia brittonii M.Howe, Ann. Missouri Bot. Gard. 2 (1/2): 50, 1915 (Britton 1915). * Riccia chudoana Steph., Sp. Hepat. (Stephani) 6: 1, 1917 (Stephani 1917a). *** Riccia coracina Jovet-Ast, Cryptog. Bryol. 24 (3): 212, 2003 (Jovet-Ast 2003). *** Riccia corrugata Jovet-Ast, Cryptog. Bryol. 21 (4): 308, 2000 (Jovet-Ast 2000). *** Riccia crassivenia Jovet-Ast, Cryptog. Bryol. 21 (4): 312, 2000 (Jovet-Ast 2000). *** Riccia cubensis S.W.Arnell, Bryologist 61 (2): 142, 1958 (Arnell 1958c). 494 *** *** *** *** *** *** *** *** *** ** *** *** *** * *** *** *** *** ** *** ** *** *** *** *** *** *** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Riccia discolor Lehm. et Lindenb., Nov. Stirp. Pug. 4: 1, 1832 (Lehmann 1832). Riccia ekmanii S.W.Arnell, Bryologist 61 (2): 143, 1958 (Arnell 1958c). Riccia elliottii Steph., Bull. Herb. Boissier 6 (4): 324 (16), 1898 (Stephani 1898a). Riccia enyae Jovet-Ast, Cryptog. Bryol. Lichénol. 12 (3): 230, 1991 (Jovet-Ast 1991). Riccia erythrocarpa Jovet-Ast, Cryptog. Bryol. Lichénol. 12 (3): 257, 1991 (JovetAst 1991). Riccia fruchartii Steph., Bull. Herb. Boissier 6 (4): 330 (22), 1898 (Stephani 1898a). Riccia gangetica Ahmad ex L.Söderstr., A.Hagborg et von Konrat, Phytotaxa 65: 57, 2012 (Söderström et al. 2012f). Based on: Riccia gangetica Ahmad, Curr. Sci. 11 (11): 433, 1942 (Ahmad 1942), nom. inval. Riccia grandis Nees, Fl. Bras. (Martius) 1 (1): 300, 1833 (Nees 1833a). Riccia helenae Jovet-Ast, J. Hattori Bot. Lab. 74: 96, 1993 (Jovet-Ast 1993a). Riccia hirta (Austin) Underw., Bot. Gaz. 19 (7): 274, 1894 (Underwood 1894). Bas.: Riccia arvensis var. hirta Austin, Proc. Acad. Nat. Sci. Philadelphia 21: 232, 1869 (Austin 1869). Riccia horrida Jovet-Ast, Cryptog. Bryol. Lichénol. 12 (3): 226, 1991 (Jovet-Ast 1991). Riccia hortorum Bory, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 18 (1): 435, 1836 [1837] (Lindenberg 1836). Riccia howellii M.Howe, Proc. Calif. Acad. Sci. (ser. 4) 21 (17): 202, 1934 (Howe 1934). Riccia ianthina Jovet-Ast, Rev. Bryol. Lichénol. 44 (4): 418, 1978 (Jovet-Ast 1978). 450 Riccia inlexa Taylor, London J. Bot. 5: 417, 1846 (Taylor 1846b). Riccia iodocheila M.Howe, Proc. Calif. Acad. Sci. (ser. 4) 21 (17): 200, 1934 (Howe 1934). Riccia lanceolata Steph., Hedwigia 27 (3/4): 110, 1888 (Stephani 1888d). Riccia lindmanii Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 23 (III, 2): 29, 1897 (Stephani 1897a). Riccia macallisteri M.Howe, Bryologist 20 (3): 35, 1917 (Howe 1917). Riccia macrospora Steph., Bull. Herb. Boissier 6 (4): 328 (20), 1898 (Stephani 1898a). Riccia mamillata Trab. ex Steph., Rev. Bryol. 16 (5): 65, 1889 (Stephani 1889b). Riccia mauryana Steph., Bull. Herb. Boissier 6 (4): 327 (19), 1898 (Stephani 1898a). Riccia olgensis Na-halang, Brunonia 3 (1): 100, 1980 (Na-halang 1980). Riccia planobiconvexa Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 23 (III, 2): 29, 1897 (Stephani 1897a). Riccia ridleyi A.Gepp, J. Linn. Soc., Bot. 27 (181): 74, 1890 (Gepp 1890). Riccia sanguineisporis Jovet-Ast, Cryptog. Bryol. Lichénol. 12 (3): 253, 1991 (Jovet-Ast 1991). Riccia squamata Nees, Fl. Bras. (Martius) 1 (1): 302, 1833 (Nees 1833a). 450 Riccia ianthina seems identical to glabrous phases of Riccia atromarginata (Schuster 1992d). It is related to Riccia iodocheila and Riccia violacea (Bischler-Causse et al. 2005), but it is only known from the type specimen. World checklist of hornworts and liverworts 495 *** Riccia subdepilata Jovet-Ast, Cryptog. Bryol. Lichénol. 12 (3): 228, 1991 (Jovet-Ast 1991). *** Riccia subplana Steph., Symb. Antill. (Urban) 3 (2): 275, 1902 (Stephani 1902e). *** Riccia taeniiformis Jovet-Ast, Cryptog. Bryol. Lichénol. 12 (3): 270, 1991 (Jovet-Ast 1991). *** Riccia viannae Jovet-Ast, Cryptog. Bryol. Lichénol. 12 (3): 261, 1991 (Jovet-Ast 1991). *** Riccia vitalii Jovet-Ast, Mem. New York Bot. Gard. 45: 285, 1987 (Jovet-Ast 1987). *** Riccia weinionis Steph., Bull. Herb. Boissier 6 (4): 326 (18), 1898 (Stephani 1898a). ** sect. Pilifer O.H.Volk, Mitt. Bot. Staatssamml. München 19: 455, 1983 (Volk 1983). *** Riccia alatospora O.H.Volk et Perold, Bothalia 15 (3/4): 534, 1985 (Volk and Perold 1985). *** Riccia albomarginata Bisch. ex C.Krauss, Flora 29 (9): 135, 1846 (Krauss 1846). *** Riccia albovestita O.H.Volk, Mitt. Bot. Staatssamml. München 17: 245, 1981 (Volk 1981). *** Riccia ampullacea Perold, Bothalia 20 (2): 168, 1990 (Perold 1990e). *** Riccia concava Bisch. ex C.Krauss, Flora 29 (9): 135, 1846 (Krauss 1846). *** Riccia elongata Perold, Bothalia 20 (2): 167, 1990 (Perold 1990e). *** Riccia furfuracea Perold, Bothalia 20 (2): 176, 1990 (Perold 1990b). *** Riccia hantamensis Perold, Bothalia 19 (2): 157, 1989 (Perold 1989b). *** Riccia hirsuta O.H.Volk et Perold, Bothalia 16 (2): 187, 1986 (Volk and Perold 1986a). *** Riccia namaquensis Perold, Bothalia 20 (2): 180, 1990 (Perold 1990b). *** Riccia parvoareolata O.H.Volk et Perold, Bothalia 15 (1/2): 117, 1984 (Volk and Perold 1984). *** Riccia pulveracea Perold, Bothalia 20 (2): 185, 1990 (Perold 1990c). *** Riccia radicosa Pearson, Natuurw. Tijdschr. 4 (5/6): 142, 1922 (Pearson 1922a). *** Riccia simii Perold, Bothalia 20 (1): 36, 1990 (Perold 1990a). *** Riccia trachyglossa Perold, Bothalia 20 (2): 172, 1990 (Perold 1990e). *** Riccia villosa Steph., Akad. Wiss. Wien, Math.-Naturwiss. Kl., Denkschr. 88: 724, 1913 (Stephani 1913b). *** Riccia vitrea Perold, Bothalia 20 (2): 178, 1990 (Perold 1990b). ** sect. Riccia *** Riccia albolimbata S.W.Arnell, Mitt. Bot. Staatssamml. München 2 (16): 264, 1957 (Arnell 1957c). *** Riccia alboporosa Perold, Bothalia 19 (1): 12, 1989 (Perold 1989c). *** Riccia albornata O.H.Volk et Perold, Bothalia 18 (2): 160, 1988 (Volk et al. 1988). *** Riccia angolensis Steph., Bull. Herb. Boissier 6 (4): 323 (15), 1898 (Stephani 1898a). *** Riccia argenteolimbata O.H.Volk et Perold, Bothalia 18 (2): 155, 1988 (Volk et al. 1988). ** Riccia atlantica Sérgio et Perold, J. Bryol. 17 (1): 127, 1992 (Sérgio and Perold 1992). *** Riccia atromarginata Levier, Nuovo Giorn. Bot. Ital. 21 (2): 291, 1889 (Martelli 1889). 496 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Riccia atromarginata var. jovet-astiae Rauh et Buchloh, Rev. Bryol. Lichénol. 30 (1/2): 77, 1961 (Rauh and Buchloh 1961). *** Riccia atropurpurea Sim, Trans. Roy. Soc. South Africa 15 (1): 11, 1926 (Sim 1926). *** Riccia beyrichiana Hampe, Nov. Stirp. Pug. 7: 1, 1838 (Lehmann 1838). *** Riccia bicarinata Lindb., Rev. Bryol. 4 (3): 41, 1877 (Lindberg 1877d). *** Riccia bicolorata Perold, Bothalia 20 (2): 188, 1990 (Perold 1990c). *** Riccia bifurca Hofm., Deutschl. Fl., heil 2 (Hofm.): 95, 1795 [1796] (Hofmann 1795). *** Riccia billardierei Mont. et Nees, Syn. Hepat. 4: 602, 1846 (Gottsche et al. 1846). *** Riccia breidleri Jur. ex Steph., Hedwigia 24 (1): 6, 1885 (Stephani 1885e). *** Riccia californica Austin, Bull. Torrey Bot. Club 6 (7): 46, 1875 (Austin 1875c). *** Riccia campbelliana M.Howe, Mem. Torrey Bot. Club 7: 26, 1899 (Howe 1899). *** Riccia ciliata Hofm., Deutschl. Fl., heil 2 (Hofm.): 95, 1795 [1796] (Hofmann 1795). *** Riccia ciliifera Link, Syn. hepat. eur: 119, 1829 (Lindenberg 1829). * Riccia congoana Steph., Bull. Herb. Boissier 6 (4): 328 (20), 1898 (Stephani 1898a). 451 *** Riccia crinita Taylor, London J. Bot. 5: 415, 1846 (Taylor 1846b). *** Riccia crozalsii Levier, Rev. Bryol. 29 (4): 73, 1902 (Levier 1902). * Riccia crustata Trab., Bull. Soc. Hist. Nat. Afrique N. 7: 87, 1916 (Trabut 1916). 452 ** Riccia dictyospora M.Howe, Bull. Torrey Bot. Club 28 (3): 163, 1901 (Howe 1901a). *** Riccia glauca L., Sp. Pl. 1: 1139, 1753 (Linnaeus 1753). ** Riccia glauca var. ciliaris Warnst., Verh. Bot. Vereins Prov. Brandenburg 27 (1): 87, 1886 (Warnstorf 1886). ** Riccia gothica Damsh. et Hallingb., Lindbergia 12 (2/3): 100, 1986 [1987] (Damsholt and Hallingbäck 1986). *** Riccia gougetiana Durieu et Mont., Ann. Sci. Nat. Bot. (sér. 3) 11: 35, 1849 (Montagne 1849). ** Riccia gougetiana var. armatissima Levier ex Müll.Frib., Lebermoose 1 (3): 161, 1907 (Müller 1907b). *** Riccia lamellosa Raddi, Opusc. Sci. 2 (6): 351, 1818 (Raddi 1818b). *** Riccia ligula Steph., Bull. Herb. Boissier 6 (4): 315 (7), 1898 (Stephani 1898a). *** Riccia limbata Bisch. ex C.Krauss, Flora 29 (9): 135, 1846 (Krauss 1846). *** Riccia macrocarpa Levier, Bull. Soc. Bot. Ital. 1894: 114, 1894 (Levier 1894). *** Riccia mammifera O.H.Volk et Perold, Bothalia 16 (2): 176, 1986 (Volk and Perold 1986b). *** Riccia melitensis C.Massal., Bull. Soc. Bot. Ital. 1913 (2/3): 52, 1913 (Massalongo 1913). *** Riccia michelii Raddi, Opusc. Sci. 2 (6): 352, 1818 (Raddi 1818b). *** Riccia microciliata O.H.Volk et Perold, Bothalia 16 (2): 173, 1986 (Volk and Perold 1986b). 451 Riccia congoana is possibly conspeciic with Riccia billardierei (Perold 1989a). 452 Riccia crustata may be conspeciic with Riccia albida (Jovet-Ast 1986). World checklist of hornworts and liverworts 497 *** Riccia montana Perold, Bothalia 19 (1): 9, 1989 (Perold 1989c). *** Riccia natalensis Sim, Trans. Roy. Soc. South Africa 15 (1): 9, 1926 (Sim 1926). *** Riccia nigrella DC., Fl. Franç. (DC. & Lamarck), 5 (6): 193, 1815 (De Candolle and Lamarck 1815). *** Riccia okahandjana S.W.Arnell, Mitt. Bot. Staatssamml. München 2 (16): 268, 1957 (Arnell 1957c). ** Riccia ozarkiana McGregor, Bryologist 63 (1): 30, 1960 (McGregor 1960). *** Riccia papillosa Moris, Stirp. Sard. Elench.: 18, 1829 (Moris 1829). *** Riccia pottsiana Sim, Trans. Roy. Soc. South Africa 15 (1): 10, 1926 (Sim 1926). *** Riccia rosea O.H.Volk et Perold, Bothalia 16 (2): 181, 1986 (Volk and Perold 1986d). * Riccia runssorensis Steph., Bull. Herb. Boissier 6 (4): 330 (22), 1898 (Stephani 1898a). 453 *** Riccia sommieri Levier, Isola Giglio: 119, 1900 (Bottini et al. 1900). *** Riccia sorocarpa Bisch., Bem. Leberm.: 145, 1835 (Bischof 1835). ** Riccia sorocarpa var. heegii Schifn., Hedwigia 53 (1/2): 36, 1912 (Schifner 1912a). *** Riccia subbifurca Warnst. ex Croz., Rev. Bryol. 30 (4): 62, 1903 (Crozals 1903b). ** Riccia tenella D.L.Jacobs, Bryologist 52 (4): 168, 1949 [1950] (Jacobs 1949). *** Riccia trabutiana Steph., Rev. Bryol. 16 (5): 65, 1889 (Stephani 1889b). *** Riccia violacea M.Howe, Ann. Missouri Bot. Gard. 2 (1/2): 51, 1915 (Britton 1915). *** Riccia violacea var. laevis Jovet-Ast, Cryptog. Bryol. Lichénol. 10 (2): 100, 1989 (Jovet-Ast 1989). *** Riccia warnstorii Limpr. ex Warnst., Verh. Bot. Vereins Prov. Brandenburg 27 (1): 85, 1886 (Warnstorf 1886). ** subg. Ricciella (A.Braun) Boulay, Musc. France 2: 198, 1904 (Boulay 1904). Bas.: Ricciella A.Braun, Flora 4 (2): 756, 1821 (Braun 1821). *** Riccia cancellata Taylor, London J. Bot. 5: 414, 1846 (Taylor 1846b). *** Riccia cincta Jovet-Ast, Cryptog. Bryol. 21 (4): 303, 2000 (Jovet-Ast 2000). *** Riccia cruciata Kashyap, J. Bombay Nat. Hist. Soc. 24 (2): 349, 1916 (Kashyap 1916). *** Riccia eburnea Jovet-Ast, Cryptog. Bryol. 21 (4): 300, 2000 (Jovet-Ast 2000). *** Riccia hasskarliana Steph., Bull. Herb. Boissier 6 (5): 374 (49), 1898 (Stephani 1898b). *** Riccia junghuhniana Nees et Lindenb., Syn. Hepat. 4: 609, 1846 (Gottsche et al. 1846). ** Riccia junghuhniana var. simplex Schifn., Hep. Fl. Buitenzorg: 14, 1900 (Schifner 1900a). *** Riccia mangalorica Ahmad ex Jovet-Ast, Cryptog. Bryol. 24 (3): 223, 2003 (JovetAst 2003). Based on: Riccia mangalorica Ahmad, Curr. Sci. 11 (11): 433, 1942 (Ahmad 1942), nom. inval. *** Riccia multiida (Steph.) Steph., Bull. Herb. Boissier 6 (5): 365 (40), 1898 (Stephani 1898b). Bas.: Ricciella multiida Steph., Hedwigia 28 (4): 273, 1889 (Stephani 1889c). 453 Riccia runssorensis is possibly conspeciic with Riccia macrospora (Seppelt 1998). 498 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) ** Riccia polycarpa (Trab.) Jelenc, Bull. Trimestriel Geogr. Archeol. Oran 73 (228): 88, 1950 (Jelenc 1950). Bas.: Ricciella polycarpa Trab., Mém. Soc. Hist. Nat. Afrique N. 3: 36, 1933 (Maire 1933). *** Riccia porosa Taylor, London J. Bot. 5: 416, 1846 (Taylor 1846b). *** Riccia pullulans Jovet-Ast, Cryptog. Bryol. Lichénol. 18 (3): 183, 1997 (Jovet-Ast 1997). ** sect. Ricciella (A.Braun) Bisch., Bem. Leberm.: 160, 1835 (Bischof 1835). Bas.: Ricciella A.Braun, Flora 4 (2): 756, 1821 (Braun 1821). * Riccia bahiensis Steph., Bull. Herb. Boissier 6 (5): 375 (50), 1898 (Stephani 1898b). 454 *** Riccia canaliculata Hofm., Deutschl. Fl., heil 2 (Hofm.): 96, 1795 [1796] (Hofmann 1795). *** Riccia chiapasensis Jovet-Ast, Cryptog. Bryol. Lichénol. 14 (3): 235, 1993 (Jovet-Ast 1993b). *** Riccia crassifrons Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 570, 1885 (Spruce 1885). *** Riccia duplex Lorb. ex Müll.Frib., Hedwigia 80 (1/2): 100, 1941 (Müller 1941). ** Riccia duplex var. megaspora Na-halang, Brunonia 3 (1): 128, 1980 (Na-halang 1980). *** Riccia dussiana Steph., Symb. Antill. (Urban) 3 (2): 275, 1902 (Stephani 1902e). *** Riccia luitans L., Sp. Pl. 1: 1139, 1753 (Linnaeus 1753). *** Riccia frostii Austin, Bull. Torrey Bot. Club 6 (3): 17, 1875 (Austin 1875b). * Riccia frostii var. crystallinoides Schifn., Ann. K. K. Naturhist. Hofmus. 27: 503, 1913 (Schifner 1913). *** Riccia geissleriana Jovet-Ast, Cryptog. Bryol. Lichénol. 14 (3): 236, 1993 (JovetAst 1993b). *** Riccia hegewaldiana Jovet-Ast, Cryptog. Bryol. Lichénol. 14 (3): 238, 1993 (JovetAst 1993b). *** Riccia huebeneriana Lindenb., Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 18 (1): 504d, 1836 [1837] (Lindenberg 1836). * Riccia huebeneriana subsp. sullivantii (Austin) R.M.Schust., Hepat. Anthocerotae N. Amer. 6: 457, 1992 (Schuster 1992d). Bas.: Riccia sullivantii Austin, Proc. Acad. Nat. Sci. Philadelphia 21: 233, 1869 (Austin 1869). *** Riccia jovet-astiae E.Vianna, Bol. Inst. Bioci. Univ. Fed. Rio Grande do Sul 38: 165, 1985 (Vianna 1985). *** Riccia limicola Jovet-Ast, Rev. Bryol. Lichénol. 44 (4): 422, 1978 (Jovet-Ast 1978). *** Riccia paraguayensis Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: cxcvi, 1889 [1890] (Spruce 1889). *** Riccia paranaensis Hässel, Opera Lilloana 7: 228, 1962 [1963] (Hässel 1962). 455 *** Riccia perennis Steph., Bull. Herb. Boissier 6 (5): 372 (47), 1898 (Stephani 1898b). 454 Riccia bahiensis is conspeciic with Riccia cavernosa in Jovet-Ast (1965), but it was accepted by Bischler-Causse et al. (2005). It is only known from the type specimen. 455 Riccia paranaensis may be conspeciic with Riccia huebeneriana (Schuster 1992d), but it was accepted by Hässel and Rubies (2009). World checklist of hornworts and liverworts *** ** ** *** 499 Riccia purpurascens Lehm., Linnaea 4: 371, 1829 (Lehmann 1829). Riccia rhenana Lorb. ex Müll.Frib., Hedwigia 80 (1/2): 94, 1941 (Müller 1941). Riccia rhenana var. violacea M.F.Boiko, Chornom. Bot. J. 7: 93, 2011 (Boiko 2011). Riccia stricta (Lindenb.) Perold, Bothalia 20 (2): 197, 1990 (Perold 1990d). Bas.: Riccia luitans var. stricta Lindenb., Nova Acta Phys.-Med. Acad. Caes. Leop.Carol. Nat. Cur. 18 (1): 444, 1836 [1837] (Lindenberg 1836). ** sect. Spongodes Nees, Naturgesch. Eur. Leberm. 4: 391, 1838 (Nees 1838a). *** Riccia bullosa Link, Syn. hepat. eur: 119, 1829 (Lindenberg 1829). *** Riccia cavernosa Hofm., Deutschl. Fl., heil 2 (Hofm.): 95, 1795 [1796] (Hofmann 1795). *** Riccia crystallina L., Sp. Pl. 1: 1138, 1753 (Linnaeus 1753). *** Riccia cupulifera A.V.Duthie, Trans. Roy. Soc. South Africa 24 (2): 116, 1936 (Duthie and Garside 1936). *** Riccia garsidei Sim, Trans. Roy. Soc. South Africa 15 (1): 13, 1926 (Sim 1926). *** Riccia moenkemeyeri Steph., Bot. Jahrb. Syst. 8 (2): 95, 1886 (Stephani 1886d). *** Riccia rubricollis Garside et A.V.Duthie ex Perold, Bothalia 21 (1): 51, 1991 (Perold 1991a). *** Riccia volkii S.W.Arnell, Mitt. Bot. Staatssamml. München 2 (16): 271, 1957 (Arnell 1957c). *** Riccia vulcanicola Eb.Fisch., Trop. Bryol. 8: 70, 1993 (Fischer 1993). ** subg. hallocarpus (Lindb.) Jovet-Ast, Cryptog. Bryol. Lichénol. 14 (3): 220, 1993 (Jovet-Ast 1993b). Bas.: hallocarpus Lindb., Not. Sällsk. Fauna Fl. Fenn. Förh. 13: 377, 1874 (Lindberg 1874a). *** Riccia curtisii (Austin) Austin, Bull. Torrey Bot. Club 6 (52): 305, 1879 (Austin 1879). Bas.: Cryptocarpus curtisii Austin, Proc. Acad. Nat. Sci. Philadelphia 21: 231, 1869 (Austin 1869). ** Riccia leptothallus R.M.Schust., J. Hattori Bot. Lab. 71: 271, 1992 (Schuster 1992c). *** Riccia perssonii Sultan Khan, Svensk Bot. Tidskr. 49 (3): 433, 1955 (Kahn 1955). ** subg. Triseriata Jovet-Ast, Cryptog. Bryol. Lichénol. 17 (2): 132, 1996 (JovetAst 1996). *** Riccia singularis Jovet-Ast, Cryptog. Bryol. Lichénol. 17 (2): 127, 1996 (Jovet-Ast 1996). Incertae sedis ** Riccia abuensis Bapna, Trans. Brit. Bryol. Soc. 4 (2): 249, 1962 (Bapna 1962). ** Riccia acutisulca Steph., Sp. Hepat. (Stephani) 6: 1, 1917 (Stephani 1917a). * Riccia amboinensis Schifn., Leberm., Forschungsr. Gazelle 4 (4): 44, 1890 (Schifner 1890). ** Riccia aravalliensis Pandé et Udar, J. Indian Bot. Soc. 36 (3): 249, 1957 (Pandé and Udar 1957). 500 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) * Riccia arnellii Sultan Khan, Bryologist 60 (1): 29, 1957 (Khan 1957). ** Riccia asprella Carrington et Pearson, Proc. Linn. Soc. New South Wales (ser. 2) 2 (4): 1059, 1888 (Carrington and Pearson 1888a). * Riccia asservanda De Not. ex Lamothe, Rech. Anat. Taxinom. Gamét. Marchantiales: 138, 1919 (Lamothe 1919). ** Riccia attenuata Pandé, Proc. Natl. Inst. Sci. India B 25 (2): 92, 1959 (Pandé and Udar 1959). * Riccia balansae Steph., Bull. Herb. Boissier 6 (5): 370 (45), 1898 (Stephani 1898b). * Riccia bialbistrata Hässel, Opera Lilloana 7: 243, 1962 [1963] (Hässel 1962). 456 ** Riccia biokoensis Perold, Nova Hedwigia 64 (1/2): 244, 1997 (Perold 1997b). ** Riccia blackii Na-halang, Brunonia 3 (1): 81, 1980 (Na-halang 1980). *** Riccia caroliniana Na-halang, Brunonia 3 (1): 72, 1980 (Na-halang 1980). ** Riccia cartilaginosa Steph., Hedwigia 28 (4): 272, 1889 (Stephani 1889c). * Riccia chartacea K.I.Goebel, Organogr. Pl., ed. 2, 2 (1): 630, 1915 (Goebel 1915). ** Riccia chinensis Herzog, Symb. Sin. 5: 1, 1930 (Nicholson et al. 1930). ** Riccia collata Na-halang, Brunonia 3 (1): 122, 1980 (Na-halang 1980). ** Riccia compacta Garside, Trans. Roy. Soc. South Africa 27 (1): 17, 1939 (Duthie and Garside 1939). ** Riccia convexa Steph., Sp. Hepat. (Stephani) 6: 2, 1917 (Stephani 1917a). * Riccia coronata Sim, Trans. Roy. Soc. South Africa 15 (1): 9, 1926 (Sim 1926). 457 ** Riccia crassa Steph., Bull. Herb. Boissier 6 (5): 376 (51), 1898 (Stephani 1898b). ** Riccia crenatodentata O.H.Volk, Nova Hedwigia 46 (1/2): 27, 1988 (Volk 1988). ** Riccia delavayi Steph., Bull. Herb. Boissier 6 (5): 367 (42), 1898 (Stephani 1898b). ** Riccia deserticola Steph., Bull. Herb. Boissier 6 (5): 373 (48), 1898 (Stephani 1898b). *** Riccia erubescens Perold, J. Bryol. 16 (3): 371, 1991 (Perold 1991b). ** Riccia esulcata Steph., Sp. Hepat. (Stephani) 6: 2, 1917 (Stephani 1917a). ** Riccia fertilissima Steph., Sp. Hepat. (Stephani) 6: 2, 1917 (Stephani 1917a). * Riccia gemmifera O.H.Volk, Nova Hedwigia 39: 131, 1984 (Volk 1984). 458 ** Riccia grollei Udar, Curr. Sci. 34 (4): 126, 1965 (Udar 1965). Nom. nov. pro Riccia tuberculata Pandé et Udar, Proc. Natl. Inst. Sci. India B 24 (2): 83, 1958 (Pandé and Udar 1958), nom. illeg. ** Riccia handelii Schifn., Symb. sin. 2: 81, 1937 (Schifner 1937). ** Riccia hawaiiensis Hürl., Phytologia 61 (5): 339, 1986 (Hürlimann 1986). ** Riccia indica Udar et A.Gupta, Proc. V Indian Geophytol. Conf., Special Publ.: 307, 1984 (Udar and Gupta 1984). ** Riccia indira-gandhiensis Dabhade et A.Hasan, J. Bombay Nat. Hist. Soc. 83 (2): 400, 1986 (Dabhade and Hasan 1986). 456 Riccia bialbistrata is a doubtful taxon closely related to Riccia lindmanii (Bischler-Causse et al. 2005). 457 Riccia coronata is only known from the type specimen, which could not be traced. he description is very brief and it is suspected that it refers to smaller plants of Riccia natalensis (Perold 1999a). 458 Riccia gemmifera is possibly conspeciic with Riccia atropurpurea (Perold 1999a). World checklist of hornworts and liverworts * ** ** *** ** ** ** ** * ** ** ** *** * ** ** ** * *** ** *** ** ** ** 501 Riccia intermedia Roum., Mém. Soc. Arts Sci. Carcassonne 5: 198, 1888 (Roumeguère 1888). 459 Riccia jodhpurensis Bapna, Bot. Not. 114 (2): 181, 1961 (Bapna 1961). Riccia kirinensis C.Gao et K.C.Chang, Acta Phytotax. Sin. 16 (4): 117, 1978 (Gao and Chang 1978). Riccia laxisquamata (Steph.) Steph., Bull. Herb. Boissier 6 (5): 371 (46), 1898 (Stephani 1898b). Bas.: Ricciella laxisquamata Steph., Bot. Jahrb. Syst. 20 (3): 299, 1895 (Stephani 1895a). Riccia liaoningensis C.Gao et K.C.Chang, Acta Phytotax. Sin. 16 (4): 113, 1978 (Gao and Chang 1978). Riccia linearis (Schifn.) Steph., Bull. Herb. Boissier 6 (5): 371 (46), 1898 (Stephani 1898b). Bas.: Ricciella linearis Schifn., Leberm., Forschungsr. Gazelle 4 (4): 43, 1890 (Schifner 1890). Riccia luticola Na-halang, Brunonia 3 (1): 123, 1980 (Na-halang 1980). Riccia mamrensis Perold, Cryptog. Bryol. 26 (1): 68, 2005 (Perold 2005). Riccia marginata Lindb., Meddel. Soc. Fauna Fl. Fenn. 1: 106, 1876 [1877] (Lindberg 1876b). 460 Riccia melanospora Kashyap, Liverworts W. Himal. 1: 94, 1929 (Kashyap 1929). Riccia miyakeana Schifn., Österr. Bot. Z. 49 (11): 386, 1899 (Schifner 1899c). Riccia muscicola Steph., Hedwigia 24 (1): 4, 1885 (Stephani 1885e). Riccia nigerica E.W.Jones, Trans. Brit. Bryol. Soc. 3 (2): 225, 1957 (Jones 1957a). Riccia nigrescens Mont., Voy. Amér. Mérid., Bot. 7 (1): 15, 1839 (Montagne 1839b). 461 Riccia nipponica S.Hatt., J. Hattori Bot. Lab. 9: 38, 1953 (Shimizu and Hattori 1953a). Riccia novo-hannoverana Schifn., Leberm., Forschungsr. Gazelle 4 (4): 44, 1890 (Schifner 1890). Riccia numeensis Steph., Bull. Herb. Boissier 6 (4): 343 (35), 1898 (Stephani 1898a). Riccia obtusa Meijer, J. Hattori Bot. Lab. 20: 113, 1958 (Meijer 1958). 462 Riccia oerstediana Lindenb. et Hampe, Linnaea 24 (3): 304, 1851 [1852] (Hampe 1851b). Riccia pandei Udar, J. Indian Bot. Soc. 38 (1): 149, 1959 (Udar 1959). Riccia papillispora Steph., Bull. Herb. Boissier 6 (4): 334 (26), 1898 (Stephani 1898a). Riccia papulosa (Steph.) Steph., Bull. Herb. Boissier 6 (5): 377 (52), 1898 (Stephani 1898b). Bas.: Ricciella papulosa Steph., Hedwigia 28 (4): 273, 1889 (Stephani 1889c). Riccia papulosa var. variabilis Na-halang, Brunonia 3 (1): 112, 1980 (Na-halang 1980). Riccia pathankotensis Kashyap, J. Bombay Nat. Hist. Soc. 24 (2): 349, 1916 (Kashyap 1916). 459 Riccia intermedia (type from France) has neither been recognized in any recent treatment nor synonymized. he name may have priority once its identity is determined. 460 Riccia marginata is possibly conspeciic with Riccia beyrichiana (Potemkin and Ahti 2012). 461 Riccia nigrescens is an Anthoceros species (Hässel and Rubies 2009). 462 Riccia obtusa may be conspeciic with Riccia gangetica (Söderström et al. 2010a). 502 * * ** ** *** ** * ** * ** * *** ** *** ** * ** *** ** *** ** * * * ** * Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Riccia perthiana Steph. ex K.I.Goebel, Organogr. Pl., ed. 2, 2 (1): 630, 1915 (Goebel 1915). Riccia prominens Meijer, J. Hattori Bot. Lab. 20: 111, 1958 (Meijer 1958). 463 Riccia pseudoluitans C.Gao et K.C.Chang, Acta Phytotax. Sin. 16 (4): 116, 1978 (Gao and Chang 1978). Riccia pubescens S.Hatt., Nat. Sci. Mus. 14 (6): 141, 1943 (Hattori 1943a). Riccia radiata Perold, Bothalia 34 (1): 23, 2004 (Perold 2004). Riccia rechingeri Steph., Akad. Wiss. Wien, Math.-Naturwiss. Kl., Denkschr. 81: 288, 1907 (Stephani 1907a). Riccia reticulatula Udar, Bull. Bot. Soc. Univ. Saugar 13: 49, 1961 (Udar 1961). Riccia rorida Na-halang, Brunonia 3 (1): 101, 1980 (Na-halang 1980). Riccia saharensis Steph. ex Jovet-Ast, Rev. Bryol. Lichénol. 26 (1/2): 62, 1957 (Gillett and Jovet-Ast 1957). 464 Riccia satoi S.Hatt., Bot. Mag. (Tokyo) 62 (733/734): 109, 1949 (Hattori 1949). Riccia schroederi Steph., 52 (5): 304, 1912 (Stephani 1912a). Riccia schweinfurthii Steph., Bull. Herb. Boissier 6 (4): 339 (31), 1898 (Stephani 1898a). Riccia sibayenii Perold, Bothalia 31 (1): 151, 2001 (Perold 2001a). Riccia somaliensis Perold, J. Bryol. 16 (3): 367, 1991 (Perold 1991b). Riccia spongiosula Na-halang, Brunonia 3 (1): 113, 1980 (Na-halang 1980). Riccia subtilis (Steph.) Steph., Bull. Herb. Boissier 6 (5): 364 (39), 1898 (Stephani 1898b). Bas.: Ricciella subtilis Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 23 (III, 2): 31, 1897 (Stephani 1897a). Riccia sumatrana Meijer, J. Hattori Bot. Lab. 20: 114, 1958 (Meijer 1958). Riccia symoensii Vanden Berghen, Explor. Hydrobiol. Lac Bangweolo Luapula: 191, 1972 (Vanden Berghen 1972b). Riccia tasmanica Steph. ex Rodway, Tasm. Bryoph.: 4, 1917 (Rodway 1917b). Riccia tomentosa O.H.Volk et Perold, Bothalia 20 (1): 25, 1990 (Volk and Perold 1990). Riccia treubiana Steph., Bull. Herb. Boissier 6 (4): 323 (15), 1898 (Stephani 1898a). Riccia treubiana var. subrubescens Schifn., Hep. Fl. Buitenzorg: 16, 1900 (Schifner 1900a). Riccia triangularis Steph., Bull. Mus. Natl. Hist. Nat. 18 (2): 116, 1912 (Corbière 1912). Riccia tuberculata Poir., Encycl. (Lamarck) 6: 199, 1804 (Lamarck and Poiret 1804). 465 Riccia udarii Kanwal, J. Indian Bot. Soc. 58 (3): 282, 1979 (Kanwal 1979). Riccia velenovskyi Kavina, Arch. Přír. Výzk. Čech 16 (2): 75, 1915 (Kavina 1915). 466 463 Riccia prominens may be conspeciic with Riccia junghuhniana (Söderström et al. 2010a). 464 Riccia saharensis is possibly conspeciic with Riccia argenteolimbata (Perold 1995). 465 Riccia tuberculata (type from France) has neither been recognized in any recent European treatment nor synonymized. he name may have priority once its identity is determined. 466 Riccia velenovskyi (type from Czech Republic) has neither been recognized in any recent European treatment nor synonymized. he name may have priority once its identity is determined. World checklist of hornworts and liverworts 503 ** Riccia velimalaiana A.E.D.Daniels et P.Daniel, Bull. Bot. Surv. India 44 (1/4): 139, 2002 [2003] (Daniels and Daniel 2002). ** Riccia victoriensis Steph., Bull. Herb. Boissier 6 (5): 370, 1898 (Stephani 1898b). ** Riccia weymouthiana Steph. ex Rodway, Tasm. Bryoph.: 5, 1917 (Rodway 1917b). ** Riccia wichurae Steph., Bull. Herb. Boissier 6 (4): 330 (22), 1898 (Stephani 1898a). ** Ricciocarpos Corda, Gen. hepat.: 651, 1829 (Corda 1829). *** Ricciocarpos natans (L.) Corda, Gen. hepat.: 651, 1829 (Corda 1829). Bas.: Riccia natans L., Syst. Nat., ed. 10., 2: 1339, 1759 (Linnaeus 1759). *** Targioniaceae Dumort. by D.G. Long *** Targionia L., Sp. Pl. 1: 1136, 1753 (Linnaeus 1753). ** subg. Prototargionia R.M.Schust., Hepat. Anthocerotae N. Amer. 6: 69, 1992 (Schuster 1992d). ** Targionia stellaris (Müll.Frib.) Hässel, Opera Lilloana 7: 74, 1962 [1963] (Hässel 1962). Bas.: Grimaldia stellaris Müll.Frib., Feddes Repert. Spec. Nov. Regni Veg. 58: 61, 1955 (Müller 1955). ** subg. Targionia *** Targionia hypophylla L., Sp. Pl. 1: 1136, 1753 (Linnaeus 1753). ** Targionia hypophylla subsp. linealis W.Frey et Kürschner, Nova Hedwigia 57 (1/2): 127, 1993 (Frey and Kürschner 1993). ** Targionia lorbeeriana Müll.Frib., Hedwigia 79 (1/2): 78, 1940 (Müller 1940). * * * * * Incertae sedis Targionia dioica Schifn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 154, 1898 (Schifner 1898a). 467 Targionia elongata Bisch., Syn. Hepat. 4: 576, 1846 (Gottsche et al. 1846). Targionia iorii Gola, Ann. Bot. (Rome) 13 (1): 62, 1914 (Gola 1914a). Targionia formosica Horik., J. Jap. Bot. 11: 499, 1935 (Horikawa 1935). Targionia indica Udar et A.Gupta, Geophytology 13 (1): 83, 1983 (Udar and Gupta 1983). 467 Targionia dioica is probably conspeciic with Targionia hypophylla (Söderström et al. 2010a). 504 *** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Wiesnerellaceae Inoue by D.G.Long *** Wiesnerella Schifn., Österr. Bot. Z. 46 (3): 86, 1896 (Schifner 1896b). *** Wiesnerella denudata (Mitt.) Steph., Bull. Herb. Boissier 7 (5): 382 (154), 1899 (Stephani 1899c). Bas.: Dumortiera denudata Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 125, 1860 [1861] (Mitten 1860c). * Wiesnerella fasciaria C.Gao et K.C.Chang, Acta Bot. Yunnan. 3 (4): 391, 1981 (Gao et al. 1981). Neohodgsoniales D.G.Long *** Neohodgsoniaceae D.G.Long by D.G. Long *** Neohodgsonia Perss., Bot. Not. 107 (1): 40, 1954 (Persson 1954). Nom. nov. pro Hodgsonia Perss., Hodgsonia Leal. Stockholm: 1, 1953 (Persson 1953). *** Neohodgsonia mirabilis (Perss.) Perss., Bot. Not. 107 (1): 40, 1954 (Persson 1954). Bas.: Hodgsonia mirabilis Perss., Hodgsonia Leal. Stockholm: 1, 1953 (Persson 1953). Sphaerocarpales Cavers *** Monocarpaceae D.J.Carr ex Schelpe by D.G. Long *** Monocarpus D.J.Carr, Austral. J. Bot. 4 (2): 176, 1956 (Carr 1956). *** Monocarpus sphaerocarpus D.J.Carr, Austral. J. Bot. 4 (2): 176, 1956 (Carr 1956). *** Riellaceae Engl. by J. G. Segarra-Moragues and F. Puche *** Austroriella Cargill et J.Milne, Polish Bot. J. 58 (1): 72, 2013 (Cargill and Milne 2013). *** Austroriella salta J.Milne et Cargill, Polish Bot. J. 58 (1): 72, 2013 (Cargill and Milne 2013). World checklist of hornworts and liverworts 505 *** Riella Mont., Ann. Sci. Nat. Bot. (sér. 3) 18: 11, 1852 (Montagne 1852). *** subg. Riella *** Riella alatospora Wigglesw., J. Linn. Soc., Bot. 51 (339): 317, 1937 (Wigglesworth 1937). *** Riella americana M.Howe et Underw., Bull. Torrey Bot. Club 30 (4): 218, 1903 (Howe and Underwood 1903). * Riella battandieri Trab., Rev. Bryol. 13 (3): 35, 1886 (Trabut 1886). *** Riella bialata Trab., Rev. Bryol. 35 (4): 96, 1908 (Trabut 1908). *** Riella capensis Cavers, Rev. Bryol. 30 (5): 81, 1903 (Cavers 1903). *** Riella choconensis Hässel, Symp. Biol. Hung. 35: 341, 1987 (Hässel 1987). * Riella cyrenaica Maire, Bull. Soc. Hist. Nat. Afrique N. 30 (5): 312, 1939 (Maire and Weiler 1939). *** Riella echinospora Wigglesw., J. Linn. Soc., Bot. 51 (339): 321, 1937 (Wigglesworth 1937). * Riella gallica Balansa ex Trab., Rev. Gén. Bot. 3 (35): 450, 1891 (Trabut 1891). *** Riella halophila Banwell, Trans. Brit. Bryol. Soc. 1 (5): 475, 1951 (Banwell 1951). *** Riella helicophylla (Bory et Mont.) Mont., Ann. Sci. Nat. Bot. (sér. 3) 18: 12, 1852 (Montagne 1852). Bas.: Duriaea helicophylla Bory et Mont., Ann. Sci. Nat. Bot. (sér. 3) 1: 229, 1844 (Bory and Montagne 1844). * Riella helicophylla var. macrocarpa P.Allorge, Sched. Br. Iber. (ser. 2): 4, 1929 (Allorge 1929). * Riella indica Steph. ex Kashyap, J. Bombay Nat. Hist. Soc. 25 (2): 279, 1917 (Kashyap 1917). *** Riella notarisii (Mont.) Mont., Ann. Sci. Nat. Bot. (sér. 3) 18: 12, 1852 (Montagne 1852). Bas.: Sphaerocarpos notarisii Mont., Ann. Sci. Nat. Bot. (sér. 2) 9: 39, 1838 (Montagne 1838). 468 ** Riella numidica Trab., Bull. Soc. Hist. Nat. Afrique N. 25 (9): 391, 1934 [1935] (Trabut 1934). *** Riella pampae Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 5 (9): 207, 1979 (Hässel 1979). *** Riella parisii Gottsche, Hepat. Eur., Leberm. 38-39: no. 375, 1867 (Gottsche and Rabenhorst 1867). *** Riella purpureospora Wigglesw., J. Linn. Soc., Bot. 51 (339): 312, 1937 (Wigglesworth 1937). * Riella reuteri Mont., Ann. Sci. Nat. Bot. (sér. 3) 18: 12, 1852 (Montagne 1852). * Riella sersuensis Trab., Bull. Soc. Hist. Nat. Afrique N. 25 (9): 392, 1934 [1935] (Trabut 1934). ** Riella spiculata J.Taylor, Kew Bull. 9 (1): 45, 1954 (Taylor 1954). 468 Riella notarisii is a species complex with poorly deined taxonomic boundaries including Riella battandieri, Riella cyrenaica, Riella gallica, Riella indica, Riella reuteri and Riella sersuensis. 506 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) *** Riella trigonospora Segarra et Puche, S. African J. Bot. 94: 175, 2014 (Segarra-Moragues and Puche 2014). *** Riella undulata Hässel, Symp. Biol. Hung. 35: 341, 1987 (Hässel 1987). *** subg. Trabutiella Porsild, Bot. Tidsskr. 24 (3): 327, 1902 (Porsild 1902). *** Riella ainis M.Howe et Underw., Bull. Torrey Bot. Club 30 (4): 221, 1903 (Howe and Underwood 1903). *** Riella cossoniana Trab., Atlas l. Alger 1: 6, 1886 (Battandier and Trabut 1886). *** Riella echinata (Müll.Frib.) Segarra, Puche et Sabovlj., Phytotaxa 159 (3): 165, 2014 (Segarra-Moragues et al. 2014). Bas.: Riella cossoniana var. echinata Müll. Frib., Rev. Bryol. Lichénol. 22 (3/4): 132, 1953 [1954] (Müller 1953). *** Riella gamundiae Hässel, Rev. Bryol. Lichénol. 38 (3/4): 580, 1972 [1973] (Hässel 1972b). *** Riella heliospora Segarra, Puche et Sabovlj., Syst. Bot. 37 (2): 315, 2012 (Segarra-Moragues et al. 2012). *** Riella mediterranea Segarra, Puche, Sabovlj., M.Infante et Heras, Phytotaxa 159 (3): 170, 2014 (Segarra-Moragues et al. 2014). *** Sphaerocarpaceae Heeg by D. Long *** Geothallus Campb., Bot. Gaz. 21 (1): 13, 1896 (Campbell 1896). *** Geothallus tuberosus Campb., Bot. Gaz. 21 (1): 13, 1896 (Campbell 1896). *** Sphaerocarpos Boehm., Def. gen. pl., ed. 3: 501, 1760 (Ludwig 1760). ** subg. Austrosphaerocarpos R.M.Schust., Hepat. Anthocerotae N. Amer. 5: 813, 1992 (Schuster 1992b). *** Sphaerocarpos stipitatus Bisch. ex Lindenb., Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 18 (1): 504i, 1836 [1837] (Lindenberg 1836). ** subg. Sphaerocarpos *** Sphaerocarpos michelii Bellardi, App. l. pedem.: 52, 1792 (Bellardi 1792). ** Sphaerocarpos texanus Austin, Bull. Torrey Bot. Club 6 (30): 158, 1877 (Austin 1877). Incertae sedis *** Sphaerocarpos cristatus M.Howe, Mem. Torrey Bot. Club 7: 66, 1899 (Howe 1899). *** Sphaerocarpos donnellii Austin, Bull. Torrey Bot. Club 6 (30): 157, 1877 (Austin 1877). *** Sphaerocarpos drewiae Wigglesw., Univ. Calif. Publ. Bot. 16 (3): 129, 1929 (Wigglesworth 1929). World checklist of hornworts and liverworts 507 *** Sphaerocarpos europaeus Lorb., Jahrb. Wiss. Bot. 80: 665, 1934 (Lorbeer 1934). 469 *** Sphaerocarpos hians Haynes, Bull. Torrey Bot. Club 37 (5): 225, 1910 (Haynes 1910). ** Sphaerocarpos muccilloi E.Vianna, Lindbergia 7 (1): 58, 1981 (Vianna 1981). Names in genera not currently accepted he following taxa in unsupported genera are all poorly understood. We list them here rather than making new combinations for names we do not know the status of. Acrostolia Dumort., Recueil Observ. Jungerm.: 26, 1835 (Dumortier 1835). 470 * Acrostolia alata (Gottsche et Rabenh.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 431, 1877 (Trevisan 1877). Bas.: Pseudoneura alata Gottsche et Rabenh., Hepat. Eur., Leberm. 56-57: no. 560, 1873 (Gottsche and Rabenhorst 1873b). * Acrostolia brevifolia (Gottsche et Rabenh.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 431, 1877 (Trevisan 1877). Bas.: Pseudoneura brevifolia Gottsche et Rabenh., Hepat. Eur., Leberm. 56-57: no. 560, 1873 (Gottsche and Rabenhorst 1873b). Aphanolejeunea A.Evans, Bull. Torrey Bot. Club 38 (6): 272, 1911 (Evans 1911). * Aphanolejeunea lancifera R.M.Schust., Phytologia 45 (5): 434, 1980 (Schuster 1980b). 471 * Aphanolejeunea minima Tixier, Ann. Fac. Sci. Yaoundé 20: 7, 1975 (Tixier 1975b). 472 Aspiromitus Steph., Sp. Hepat. (Stephani) 5: 957, 1916 (Stephani 1916b). 473 * Aspiromitus asper Schifn., Arch. Hydrobiol., suppl. 21 (3/4): 402, 1955 (Schifner 1955). * Aspiromitus bullosus Schifn., Arch. Hydrobiol., suppl. 21 (3/4): 403, 1955 (Schifner 1955). * Aspiromitus crenatifrons Steph., Sp. Hepat. (Stephani) 5: 968, 1916 (Stephani 1916b). * Aspiromitus lobatus Schifn., Arch. Hydrobiol., suppl. 21 (3/4): 405, 1955 (Schifner 1955). * Aspiromitus squamulosus Schifn., Arch. Hydrobiol., suppl. 21 (3/4): 407, 1955 (Schifner 1955). 469 470 471 472 Sphaerocarpos europaeus is clearly distinct from Sphaerocarpos texanus (Bell et al. 2013). Acrostolia is congeneric with Riccardia, but a few taxa have neither been transferred nor synonymized. Aphanolejeunea lancifera is a Drepanolejeunea species (Pócs et al. 2014). Aphanolejeunea minima seems to be close to Cololejeunea gracilis based on the drawing in the original description, but the type specimen (not available for study) seems to be very fragmentary, having only reduced leaves, so its identity is uncertain (Pócs and Bernecker 2009). 473 Aspiromitus is congeneric with Anthoceros, but some taxa have neither been transferred nor synonymized. 508 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Crossotolejeunea (Spruce) Schifn., Hepat. (Engl.-Prantl): 127, 1893 (Schifner 1893b). Bas.: Lejeunea subg. Crossotolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 161, 1884 (Spruce 1884). 474 * Crossotolejeunea curvifolia Steph., Hedwigia 35 (3): 75, 1896 (Stephani 1896b). 475 Eulejeunea Steph., Hedwigia 27 (2): 60, 1888 (Stephani 1888a). 476 * Eulejeunea setulosa Steph., Sp. Hepat. (Stephani) 6: 421, 1923 (Stephani 1923). * Eulejeunea subpililoba Steph., Sp. Hepat. (Stephani) 6: 420, 1923 (Stephani 1923). 477 Euosmolejeunea (Spruce) Steph., Hedwigia 28 (3): 170, 1889 (Stephani 1889d). Bas.: Lejeunea subg. Euosmolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 241, 1884 (Spruce 1884). 478 * Euosmolejeunea parvistipula (Lindenb. et Gottsche) Steph., Hedwigia 29 (1): 80, 1890 (Stephani 1890a). Bas.: Lejeunea parvistipula Lindenb. et Gottsche, Syn. Hepat. 5: 761, 1847 (Gottsche et al. 1847). * Euosmolejeunea tenerrima (Nees) Steph., Sp. Hepat. (Stephani) 5: 589, 1914 (Stephani 1914b). Bas.: Jungermannia sordida var. tenerrima Nees, Fl. Bras. (Martius) 1 (1): 363, 1833 (Nees 1833a). 479 Fimbraria Nees, Horae Phys. Berol.: 44, 1820 (Nees 1820) nom. illeg. 480 * Fimbraria gigantea Steph., Bull. Herb. Boissier 7 (2): 93 (106), 1899 (Stephani 1899a). 481 * Fimbraria incrassata Steph., Bull. Herb. Boissier 7 (2): 87 (100), 1899 (Stephani 1899a). 482 * Fimbraria kamerunensis Steph., Sp. Hepat. (Stephani) 6: 14, 1917 (Stephani 1917a). 483 * Fimbraria pirottae Gola, Ann. Bot. (Rome) 13 (1): 64, 1914 (Gola 1914a). 474 Crossotolejeunea is congeneric with Lejeunea, but one taxon has neither been transferred nor synonymized. 475 Crossotolejeunea curvifolia has doubtful status, the type specimen could not be found in G (ReinerDrehwald and Goda 2000). 476 Eulejeunea is congeneric with Lejeunea subg. Lejeunea, but a few taxa have neither been transferred nor synonymized. 477 Eulejeunea subpililoba is morphologically similar to Lejeunea spiniloba. 478 Euosmolejeunea is here treated as a subgenus of Cheilolejeunea, but a few taxa have neither been transferred nor synonymized. 479 Euosmolejeunea tenerrima is a doubtful taxon described with three syntypes, two from Brazil and one from Java. he Java material is probably not the same species as the material from Brazil. 480 Fimbraria is congeneric with Asterella, but the following taxa have neither been transferred nor synonymized, and their status is doubtful. 481 Fimbraria gigantea belongs to Asterella subg. Asterella sect. Brachyblepharis. 482 Fimbraria incrassata was provisionally placed in synonymy with Asterella abyssinica (Wigginton and Grolle 1996), and was listed as an unassigned Asterella species by Wigginton (2009). 483 Fimbraria kamerunensis was provisionally placed in synonymy with Asterella abyssinica (Wigginton and Grolle 1996), and was listed as an unassigned Asterella species by Wigginton (2009). World checklist of hornworts and liverworts 509 Hygrolejeunea (Spruce) Schifn., Hepat. (Engl.-Prantl): 124, 1893 (Schifner 1893b). Bas.: Lejeunea subg. Hygrolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 230, 1884 (Spruce 1884). 484 * Hygrolejeunea cubensis Steph., Sp. Hepat. (Stephani) 5: 533, 1914 (Stephani 1914b). * Hygrolejeunea harpaphylla Steph., Sp. Hepat. (Stephani) 5: 552, 1914 (Stephani 1914b). * Hygrolejeunea paciica Steph., Sp. Hepat. (Stephani) 6: 411, 1923 (Stephani 1923). * Hygrolejeunea parvicalycina Steph., Hedwigia 35 (3): 103, 1896 (Stephani 1896b). * Hygrolejeunea parvistipula Steph., Sp. Hepat. (Stephani) 5: 568, 1914 (Stephani 1914b). * Hygrolejeunea patellirostris Steph., Hedwigia 35 (3): 103, 1896 (Stephani 1896b). 485 * Hygrolejeunea staudtiana Steph., Sp. Hepat. (Stephani) 5: 528, 1914 (Stephani 1914b). 486 Hypenantron Corda, Gen. hepat.: 648, 1829 (Corda 1829). 487 * Hypenantron brachypus Steph. ex Lamothe, Rech. Anat. Taxinom. Gamét. Marchantiales: 107, 1919 (Lamothe 1919). * Hypenantron brasiliense Steph. ex Lamothe, Rech. Anat. Taxinom. Gamét. Marchantiales: 104, 1919 (Lamothe 1919). Jamesoniella (Spruce) Carrington, Cat. Brit. Moss. Hepat.: 25, 1881 (Carrington 1881). Bas.: Jungermannia subg. Jamesoniella Spruce, J. Bot. 14: 230, 1876 (Spruce 1876a). 488 * Jamesoniella convoluta Steph., Sp. Hepat. (Stephani) 6: 433, 1924 (Stephani 1924). Kingiolejeunea H.Rob., Bryologist 70 (1): 53, 1967 (Robinson 1967). 489 * Kingiolejeunea guayanensis H.Rob., Bol. Soc. Venez. Ci. Nat. 32 (132/133): 259, 1976 (Robinson 1976b). 484 Hygrolejeunea is congeneric with Lejeunea, but some taxa have neither been transferred nor synonymized. 485 Hygrolejeunea patellirostris is conspeciic with Lejeunea acuta in Tixier (1995b) and Wigginton and Grolle (1996), but it was kept as an unassigned Lejeunea species by Wigginton (2009). 486 Hygrolejeunea staudtiana is likely to be either Lejeunea brenanii or Lejeunea isophylla (Wigginton and Grolle 1996). It was kept as an unassigned Lejeunea species by Wigginton (2009). 487 Hypenantron is congeneric with Asterella, but a few taxa have neither been transferred nor synonymized. 488 Jamesoniella is congeneric with Syzygiella (Feldberg et al. 2010a), but one taxon has neither been transferred nor synonymized. he holotype was burned in B (Grolle 1971b) and it is not clear where it belongs. 489 Kingiolejeunea is congeneric with Lepidolejeunea, but one taxon has neither been transferred nor synonymized. 510 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Leptocolea (Spruce) A.Evans, Bull. Torrey Bot. Club 38 (6): 261, 1911 (Evans 1911). Bas.: Lejeunea sect. Leptocolea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 294, 1884 (Spruce 1884). 490 * Leptocolea sumatrana Herzog, Ann. Bryol. 5: 96, 1932 (Herzog 1932a). 491 Mastigobryum (Nees) Lindenb. et Gottsche, Syn. Hepat. 2: 214, 1845 (Gottsche et al. 1845a) nom. illeg. Bas.: Herpetium sect. Mastigobryum Nees, Naturgesch. Eur. Leberm. 3: 43, 1838 (Nees 1838b). 492 * Mastigobryum aberrans Steph., Sp. Hepat. (Stephani) 6: 486, 1924 (Stephani 1924). * Mastigobryum asperum Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 121, 1914 (Stephani and Watts 1914). * Mastigobryum deningeri Herzog, Beih. Bot. Centralbl. 38 (2): 322, 1921 (Herzog 1921). * Mastigobryum karstenii Steph., Bull. Herb. Boissier (sér. 2) 8 (12): 952 (502), 1908 (Stephani 1908b). * Mastigobryum ledermannii Steph., Sp. Hepat. (Stephani) 6: 486, 1924 (Stephani 1924). * Mastigobryum londbergii Steph., Sp. Hepat. (Stephani) 6: 487, 1924 (Stephani 1924). * Mastigobryum longifolium Steph., Sp. Hepat. (Stephani) 6: 487, 1924 (Stephani 1924). * Mastigobryum minutitextum Steph., Sp. Hepat. (Stephani) 6: 487, 1924 (Stephani 1924). * Mastigobryum multidens Steph., Sp. Hepat. (Stephani) 6: 487, 1924 (Stephani 1924). * Mastigobryum muscicola Steph., Sp. Hepat. (Stephani) 6: 486, 1924 (Stephani 1924). * Mastigobryum nigricans Herzog, Beih. Bot. Centralbl. 38 (2): 322, 1921 (Herzog 1921). * Mastigobryum nipuranum Steph., Sp. Hepat. (Stephani) 6: 485, 1924 (Stephani 1924). * Mastigobryum palmicola Steph., Sp. Hepat. (Stephani) 6: 488, 1924 (Stephani 1924). * Mastigobryum rajanum Herzog, Mitt. Inst. Allg. Bot. Hamburg 7 (3): 190, 1931 (Herzog 1931a). * Mastigobryum ribehanum Steph., Sp. Hepat. (Stephani) 6: 488, 1924 (Stephani 1924). * Mastigobryum ruicaule Beauverd, Sp. Hepat. (Stephani) 6: 485, 1924 (Stephani 1924). * Mastigobryum schraderbergii Steph., Sp. Hepat. (Stephani) 6: 489, 1924 (Stephani 1924). * Mastigobryum squamulistipum Steph., Sp. Hepat. (Stephani) 6: 480, 1924 (Stephani 1924). * Mastigobryum subhyalinum Steph., Sp. Hepat. (Stephani) 6: 482, 1924 (Stephani 1924). * Mastigobryum venezuelanum Molk., Syn. hepat. jav.: 104, 1856 [1857] (Sande Lacoste 1856b). * Mastigobryum vermiculare Herzog, Hedwigia 66 (6): 339, 1926 (Herzog 1926). 490 Leptocolea is congeneric with Cololejeunea, but one taxon has neither been transferred nor synonymized. 491 Leptocolea sumatrana is possibly conspeciic with Cololejeunea equialbi. 492 Mastigobryum is congeneric with Bazzania, but many species have not been studied recently and thus not transferred. It is possible that some prove to be older names of existing taxa when their identities are known. Grolle and Piippo (1984) could not study Mastigobryum ledermannii, Mastigobryum londbergii and Mastigobryum longifolium since the types were destroyed in B. World checklist of hornworts and liverworts 511 Nemoursia Mérat, Ann. Agric. Franç. (ser. 4) 2 (7): 10, 1840 (Mérat 1840). * Nemoursia tuberculata Mérat, Ann. Agric. Franç. (ser. 4) 2 (7): 10, 1840 (Mérat 1840). 493 Physocolea (Spruce) Steph., Sp. Hepat. (Stephani) 5: 863, 1916 (Stephani 1916b). Bas.: Lejeunea sect. Physocolea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 292, 1884 (Spruce 1884). 494 * Physocolea tambillensis (Loitl.) Steph., Sp. Hepat. (Stephani) 5: 885, 1916 (Stephani 1916b). Bas.: Lejeunea tambillensis Loitl., Diagn. pl. nov.: 20, 1894 (Loitlesberger 1894). 495 Plectocolea (Mitt.) Mitt., Fl. vit.: 405, 1871 [1873] (Mitten 1871). Bas.: Solenostoma subg. Plectocolea Mitt., J. Linn. Soc., Bot. 8 (31): 156, 1864 [1865] (Mitten 1864a). 496 * Plectocolea subamoena S.Winkl., Rev. Bryol. Lichénol. 42 (3): 821, 1976 (Winkler 1976). 497 Polyotus Gottsche, Syn. Hepat. 2: 244, 1845 (Gottsche et al. 1845a) nom. illeg. 498 * Polyotus peckianus Austin, Proc. Acad. Nat. Sci. Philadelphia 21: 224, 1869 (Austin 1869). Schisma Dumort., Commentat. Bot. (Dumortier): 114, 1822 (Dumortier 1822) nom. illeg. 499 * Schisma orizabense (Gottsche) Steph., Sp. Hepat. (Stephani) 4: 19, 1909 (Stephani 1909d). Bas.: Sendtnera orizabensis Gottsche, Mexik. Leverm.: 139, 1863 (Gottsche 1863). * Schisma uleanum Steph., Hedwigia 44 (4): 225, 1905 (Stephani 1905a). Strepsilejeunea (Spruce) Schifn., Hepat. (Engl.-Prantl): 127, 1893 (Schifner 1893b). Bas.: Lejeunea sect. Strepsilejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 168, 1884 (Spruce 1884). 500 493 494 495 496 497 498 499 500 Nemoursia tuberculata is a Conocephalum species. Physocolea is congeneric with Cololejeunea, but one taxon has neither been transferred nor synonymized. Physocolea tambillensis is possibly conspeciic with Myriocoleopsis minutissima. Plectocolea is now regarded a subgenus of Solenostoma, but one taxon has neither been transferred nor synonymized. Plectocolea subamoena is probably conspeciic with Solenostoma amoena. Polyotus is a superluous name for Lepidolaena, but a few taxa have neither been transferred nor synonymized. Schisma is congeneric with Herbertus, but a few taxa have neither been transferred nor synonymized. Strepsilejeunea is congeneric with Cheilolejeunea, but some taxa have neither been transferred nor synonymized. 512 * * * * * * * * * * * Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Strepsilejeunea apollinea (Gottsche) Steph., Sp. Hepat. (Stephani) 5: 291, 1913 (Stephani 1913a). Bas.: Lejeunea apollinea Gottsche, Fragm. (Mueller): 64, 1880 (Gottsche 1880). Strepsilejeunea durelii Schifn., Österr. Bot. Z. 49 (6): 206, 1899 (Schifner 1899a). Strepsilejeunea hamatifolia Steph., Sp. Hepat. (Stephani) 6: 396, 1923 (Stephani 1923). Strepsilejeunea lanceolata (Gottsche) Steph., Sp. Hepat. (Stephani) 5: 283, 1913 (Stephani 1913a). Bas.: Lejeunea lanceolata Gottsche, Syn. Hepat. 3: 353, 1845 (Gottsche et al. 1845b). Strepsilejeunea muscicola Herzog, Hedwigia 74 (2): 96, 1934 (Herzog 1934a). Strepsilejeunea novae-guineae Steph., Sp. Hepat. (Stephani) 6: 397, 1923 (Stephani 1923). Strepsilejeunea obtusistipula Steph., Biblioth. Bot. 87 (2): 258, 1916 (Stephani 1916a). Strepsilejeunea papulifolia Steph., Biblioth. Bot. 87 (2): 259, 1916 (Stephani 1916a). Strepsilejeunea pectiniformis (Gottsche) Steph., Sp. Hepat. (Stephani) 5: 285, 1913 (Stephani 1913a). Bas.: Lejeunea pectiniformis Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 156, 1864 (Gottsche 1864). Strepsilejeunea renistipula Steph., Sp. Hepat. (Stephani) 5: 289, 1913 (Stephani 1913a). Strepsilejeunea vatovae Gerola, Lav. Bot. Ist. Bot. Univ. Padova 12: 479, 1947 (Gerola 1947). hysanolejeunea (Spruce) Steph., Hedwigia 31 (1): 20, 1892 (Jack and Stephani 1892). Bas.: Lejeunea subg. hysanolejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 105, 1885 (Spruce 1885). * hysanolejeunea africana Sim, Trans. Roy. Soc. South Africa 15 (1): 50, 1926 (Sim 1926). 501 Trachylejeunea (Spruce) Steph., Hedwigia 28 (4): 262, 1889 (Stephani 1889c) nom. rejic. Bas.: Lejeunea subg. Trachylejeunea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 180, 1884 (Spruce 1884). * Trachylejeunea conifera Steph., Sp. Hepat. (Stephani) 5: 302, 1913 (Stephani 1913a). * Trachylejeunea cristulilora Steph., Hedwigia 35 (3): 137, 1896 (Stephani 1896b). * Trachylejeunea englishii Steph., Bull. Misc. Inform. Kew 1899 (151/152): 126, 1899 (MacGregor 1899). 502 * Trachylejeunea jamaicensis Pearson, Ann. Bryol. 4: 98, 1931 (Pearson 1931b). * Trachylejeunea kusaiensis Inoue et H.A.Mill., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 8 (2): 147, 1965 (Inoue and Miller 1965). 501 hysanolejeunea africana may be conspeciic with Caudalejeunea africana. 502 Trachylejeunea englishii is not a Trachylejeunea species (see Zhu and So 1999a). he status needs further study. World checklist of hornworts and liverworts 513 Alphabetic list of accepted taxa *** *** *** ** *** *** *** ** *** ** ** *** ** ** * *** *** *** *** ** *** *** *** *** ** *** ** *** *** *** ** ** *** *** *** *** *** *** *** ** *** *** *** *** *** *** *** *** *** Acanthocoleus aberrans (Lindenb. et Gottsche) Kruijt ..................................................................297 Acanthocoleus aberrans var. laevis Gradst. .....................................................................................297 Acanthocoleus chrysophyllus (Lehm.) Kruijt...................................................................................297 Acanthocoleus elgonensis Gyarmati et Pócs ....................................................................................297 Acanthocoleus gilvus (Gottsche) Kruijt .........................................................................................297 Acanthocoleus javanicus (Steph.) Kruijt ........................................................................................297 Acanthocoleus juddii Kruijt ..........................................................................................................297 Acanthocoleus madagascariensis (Steph.) Kruijt .............................................................................297 Acanthocoleus trigonus (Nees et Mont.) Gradst. ...........................................................................297 Acanthocoleus yoshinaganus (S.Hatt.) Kruijt .................................................................................298 Acrobolbus africanus (Pearson) Briscoe...........................................................................................93 Acrobolbus anisodontus (Hook.f. et Taylor) Briscoe........................................................................94 Acrobolbus antillanus R.M.Schust. .................................................................................................94 Acrobolbus azoricus (Grolle et Perss.) Briscoe .................................................................................94 Acrobolbus bispinosus (J.B.Jack et Steph.) Steph. ............................................................................96 Acrobolbus caducifolius R.M.Schust. ..............................................................................................94 Acrobolbus ciliatus (Mitt.) Schifn. .................................................................................................94 Acrobolbus cinerascens (Lehm. et Lindenb.) Bastow .......................................................................94 Acrobolbus concinnus (Mitt.) Grolle ...............................................................................................94 Acrobolbus cuneifolius (Steph.) Briscoe...........................................................................................94 Acrobolbus diversifolius R.M.Schust. ..............................................................................................94 Acrobolbus epiphytus (Colenso) Briscoe ..........................................................................................94 Acrobolbus lavicans (J.J.Engel et Grolle) Briscoe et J.J.Engel .........................................................94 Acrobolbus gradsteinii (Grolle) Briscoe ...........................................................................................94 Acrobolbus integrifolius (A.Evans) Briscoe ......................................................................................94 Acrobolbus knightii (Mitt.) Briscoe ................................................................................................94 Acrobolbus kunkelii (Hässel et Solari) Briscoe et J.J.Engel ..............................................................94 Acrobolbus laxus (Lehm. et Lindenb.) Briscoe ................................................................................94 Acrobolbus limbatus (Steph.) Briscoe et J.J.Engel ...........................................................................95 Acrobolbus lophocoleoides (Mitt.) Mitt. ...........................................................................................95 Acrobolbus madeirensis (Grolle et Perss.) Briscoe ............................................................................95 Acrobolbus mittenii Steph. .............................................................................................................95 Acrobolbus ochrophyllus (Hook.f. et Taylor) R.M.Schust................................................................95 Acrobolbus papillosus (J.J.Engel et Glenny) Briscoe ........................................................................95 Acrobolbus perpusillus (Colenso) Briscoe ........................................................................................95 Acrobolbus perpusillus var. denticulatus (J.J.Engel et Glenny) Briscoe .............................................95 Acrobolbus plagiochiloides (J.J.Engel et Glenny) Briscoe .................................................................95 Acrobolbus pseudosaccatus (Grolle) Briscoe .....................................................................................95 Acrobolbus renifolius (Hässel et Solari) Briscoe et J.J.Engel ............................................................95 Acrobolbus ruwenzorensis (S.W.Arnell) Briscoe ..............................................................................95 Acrobolbus saccatus (Hook.) Trevis. ...............................................................................................95 Acrobolbus setulosus (Mitt.) Briscoe ................................................................................................95 Acrobolbus spinifolius R.M.Schust..................................................................................................95 Acrobolbus sumatranus (Schifn.) Briscoe .......................................................................................96 Acrobolbus surculosus (Nees) Trevis. ...............................................................................................96 Acrobolbus tenellus (Taylor) Trevis.................................................................................................96 Acrobolbus tenellus var. diversifolius (E.A.Hodgs.) Briscoe ..............................................................96 Acrobolbus urvilleanus (Mont.) Trevis. ..........................................................................................96 Acrobolbus viridis (Mitt.) Briscoe et J.J.Engel ................................................................................96 514 *** ** *** ** * *** *** ** *** * *** *** *** *** *** * *** ** *** *** *** *** *** *** *** *** *** *** *** *** ** *** *** *** *** *** *** *** ** ** ** ** *** ** *** *** *** ** *** ** * *** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Acrobolbus wilsonii Nees ................................................................................................................96 Acrobolbus wilsonii var. andinus Spruce .........................................................................................96 Acrochila biserialis (Lehm. et Lindenb.) Grolle ............................................................................219 Acrochila caledonica (Steph.) Inoue .............................................................................................219 Acrolejeunea abnormis (Gottsche) Pearson ...................................................................................401 Acrolejeunea allisonii Gradst. .......................................................................................................400 Acrolejeunea arcuata (Nees) Grolle et Gradst. ..............................................................................399 Acrolejeunea arcuata subsp. gradsteinii M.A.M.Renner ................................................................399 Acrolejeunea aulacophora (Mont.) Steph. .....................................................................................400 Acrolejeunea comptonii Pearson....................................................................................................401 Acrolejeunea crassicaulis (Steph.) Jian Wang bis et Gradst. ...........................................................401 Acrolejeunea emergens (Mitt.) Steph. ............................................................................................399 Acrolejeunea emergens var. confertissima (Steph.) Gradst. ..............................................................399 Acrolejeunea fertilis (Reinw., Blume et Nees) Schifn. ..................................................................399 Acrolejeunea heterophylla (A.Evans) Grolle et Gradst. ..................................................................399 Acrolejeunea inlexa (Gottsche) Pearson .......................................................................................401 Acrolejeunea infuscata (Mitt.) Jian Wang bis et Gradst. ...............................................................401 Acrolejeunea meghalayensis (Ajit P.Singh et V.Nath) Jian Wang bis et Gradst. .............................401 Acrolejeunea mollis (Hook.f. et Taylor) Schifn. ...........................................................................400 Acrolejeunea parvula (Mizut.) Gradst. .........................................................................................399 Acrolejeunea pusilla (Steph.) Grolle et Gradst. .............................................................................400 Acrolejeunea pycnoclada (Taylor) Schifn......................................................................................399 Acrolejeunea pycnoclada subsp. latistipula Gradst. ........................................................................399 Acrolejeunea recurvata Gradst. .....................................................................................................400 Acrolejeunea sandvicensis (Gottsche) Steph. .................................................................................399 Acrolejeunea securifolia (Nees) Steph............................................................................................400 Acrolejeunea securifolia subsp. caledonica (Steph.) Gradst. ............................................................400 Acrolejeunea securifolia subsp. hartmannii (Steph.) Gradst. ..........................................................400 Acrolejeunea securifolia subsp. pallida (Ångstr.) Gradst. ...............................................................400 Acrolejeunea sikkimensis (Mizut.) Gradst. ....................................................................................400 Acrolejeunea sinensis (Jian Wang bis, R.L.Zhu et Gradst.) Jian Wang bis et Gradst. ....................401 Acrolejeunea tjibodensis (Verd.) Grolle et Gradst. .........................................................................400 Acrolejeunea torulosa (Lehm. et Lindenb.) Schifn. ......................................................................399 Acrolophozia fuegiana R.M.Schust. ..............................................................................................110 Acrolophozia pectinata R.M.Schust. .............................................................................................110 Acrolophozia sulcata Hässel ..........................................................................................................110 Acromastigum adaptatum Hürl. ...................................................................................................134 Acromastigum anisostomum (Lehm. et Lindenb.) A.Evans............................................................143 Acromastigum anisostomum var. minutum E.A.Hodgs. .................................................................144 Acromastigum aurescens A.Evans ..................................................................................................144 Acromastigum bancanum (Sande Lac.) A.Evans ...........................................................................144 Acromastigum brotheri (Steph.) A.Evans ......................................................................................144 Acromastigum caledonicum (Steph.) Grolle ..................................................................................143 Acromastigum capillare (Steph.) A.Evans .....................................................................................144 Acromastigum cavifolium R.M.Schust. .........................................................................................143 Acromastigum colensoanum (Mitt.) A.Evans ex Reimers ...............................................................144 Acromastigum cunninghamii (Steph.) A.Evans .............................................................................144 Acromastigum curtilobum A.Evans ...............................................................................................144 Acromastigum divaricatum (Nees) A.Evans ex Reimers ................................................................144 Acromastigum echinatiforme (De Not.) A.Evans...........................................................................144 Acromastigum echinatum (Gottsche) A.Evans ..............................................................................144 Acromastigum exiguum (Steph.) A.Evans .....................................................................................144 World checklist of hornworts and liverworts ** *** *** ** ** ** ** *** *** ** ** ** ** ** ** ** ** *** ** *** *** ** ** ** ** *** ** ** ** *** *** *** *** * * *** *** *** *** *** *** ** *** *** *** *** *** *** *** ** ** *** 515 Acromastigum exile (Lindenb.) A.Evans .......................................................................................144 Acromastigum ilum (Steph.) A.Evans ..........................................................................................144 Acromastigum ilum var. papillosum N.Kitag. ...............................................................................144 Acromastigum imbriatum (Steph.) A.Evans .................................................................................144 Acromastigum fumosum E.A.Br. et M.A.M.Renner ......................................................................145 Acromastigum furcatifolium (Steph.) E.A.Br. ...............................................................................145 Acromastigum herzogii Grolle ......................................................................................................143 Acromastigum homodictyon (Herzog) Grolle ................................................................................143 Acromastigum inaequilaterum (Lehm. et Lindenb.) A.Evans ........................................................145 Acromastigum integrifolium (Austin) A.Evans ..............................................................................143 Acromastigum interstisiale E.A.Br. et M.A.M.Renner ...................................................................145 Acromastigum laetevirens (Sande Lac. ex Steph.) A.Evans.............................................................145 Acromastigum laevigatum A.Evans ...............................................................................................145 Acromastigum leptophyllum Herzog..............................................................................................145 Acromastigum linganum (De Not.) A.Evans ................................................................................145 Acromastigum lobuliferum A.Evans ..............................................................................................145 Acromastigum longirete Grolle......................................................................................................145 Acromastigum marginatum E.A.Hodgs. .......................................................................................145 Acromastigum microstictum A.Evans ............................................................................................145 Acromastigum mooreanum (Steph.) E.A.Hodgs. ...........................................................................145 Acromastigum moratii N.Kitag. ...................................................................................................145 Acromastigum obliquatum (Mitt.) A.Evans ..................................................................................145 Acromastigum prismaticale E.A.Br. et M.A.M.Renner..................................................................145 Acromastigum pusillum N.Kitag...................................................................................................145 Acromastigum rigidum R.M.Schust..............................................................................................145 Acromastigum stellare N.Kitag. ....................................................................................................143 Acromastigum stenophyllum R.M.Schust. .....................................................................................143 Acromastigum subechinatiforme Hürl. ..........................................................................................145 Acromastigum tenax (Steph.) A.Evans ..........................................................................................145 Acromastigum verticale (Steph.) E.A.Hodgs. ................................................................................143 Acroscyphella iwatsukii (N.Kitag.) N.Kitag. et Grolle .....................................................................99 Acroscyphella phoenicorhiza (Grolle) N.Kitag. et Grolle..................................................................99 Acroscyphella tjiwideiensis (Sande Lac.) N.Kitag. et Grolle .............................................................99 Acrostolia alata (Gottsche et Rabenh.) Trevis. .............................................................................507 Acrostolia brevifolia (Gottsche et Rabenh.) Trevis. .......................................................................507 Adelanthus aureomarginatus R.M.Schust. ......................................................................................44 Adelanthus carabayensis (Mont.) Grolle .........................................................................................44 Adelanthus falcatus (Hook.) Mitt...................................................................................................44 Adelanthus gemmiparus (R.M.Schust.) E.A.Hodgs.........................................................................44 Adelanthus integerrimus Grolle ......................................................................................................44 Adelanthus lindenbergianus (Lehm.) Mitt. .....................................................................................44 Adelanthus lingulatus J.J.Engel et Váňa..........................................................................................44 Adelanthus occlusus (Hook.f. et Taylor) Carrington .......................................................................44 Adelanthus pittieri (Steph.) Grolle .................................................................................................44 Adelanthus squarrosus Grolle..........................................................................................................44 Adelanthus tenuis J.J.Engel et Grolle..............................................................................................44 Aitchisoniella himalayensis Kashyap .............................................................................................488 Allisonia cockaynei (K.I.Goebel) R.M.Schust. ..............................................................................463 Allisoniella nigra (Rodway) R.M.Schust. .......................................................................................65 Allisoniella nigra var. acutiloba J.J.Engel ........................................................................................65 Allisoniella nigra subsp. novaezelandiae R.M.Schust. .....................................................................65 Allisoniella recurva R.M.Schust. ....................................................................................................65 516 *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** ** *** *** * *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Allisoniella scottii (R.M.Schust.) R.M.Schust. ................................................................................65 Allisoniella subbipartita (C.Massal.) R.M.Schust. et J.J.Engel ........................................................65 Allisoniella tasmanica R.M.Schust. ................................................................................................65 Alobiella husnotii (Spruce) Schifn. ................................................................................................56 Alobiellopsis acroscypha (Spruce) R.M.Schust. ................................................................................56 Alobiellopsis dominicensis (Spruce) Fulford .....................................................................................56 Alobiellopsis heteromorpha (Lehm.) R.M.Schust. ............................................................................56 Alobiellopsis parvifolia (Steph.) R.M.Schust. ..................................................................................57 Alobiellopsis pillansii (Sim) R.M.Schust. ........................................................................................57 Amazoopsis diplopoda (Pócs) J.J.Engel et G.L.Merr. ....................................................................183 Amazoopsis dissotricha (Spruce) J.J.Engel et G.L.Merr. ................................................................183 Amazoopsis gracilis J.J.Engel et G.L.Merr. ...................................................................................183 Amphicephalozia africana Váňa et M.Wigginton ...........................................................................65 Amphicephalozia amplexicaulis R.M.Schust. ..................................................................................65 Amphicephalozia geisslerae Pócs et Váňa.........................................................................................65 Anastrepta orcadensis (Hook.) Schifn. ...........................................................................................50 Anastrophyllopsis involutifolia (Mont. ex Gottsche, Lindenb. et Nees) Váňa et L.Söderstr. .............65 Anastrophyllopsis revoluta (Steph.) Váňa et L.Söderstr. ...................................................................65 Anastrophyllopsis subcomplicata (Lehm. et Lindenb.) Váňa et L.Söderstr. .......................................66 Anastrophyllum alpinum Steph. .....................................................................................................50 Anastrophyllum assimile (Mitt.) Steph. ...........................................................................................50 Anastrophyllum auritum (Lehm.) Steph. ........................................................................................50 Anastrophyllum ciliatum Steph. .....................................................................................................50 Anastrophyllum donnianum (Hook.) Steph. ...................................................................................50 Anastrophyllum ellipticum Inoue ....................................................................................................50 Anastrophyllum esenbeckii (Mont.) Steph. ......................................................................................50 Anastrophyllum issum Steph. .........................................................................................................50 Anastrophyllum joergensenii Schifn................................................................................................50 Anastrophyllum lignicola D.B.Schill et D.G.Long ..........................................................................50 Anastrophyllum michauxii (F.Weber) H.Buch ...............................................................................50 Anastrophyllum nigrescens (Mitt.) Steph. ........................................................................................51 Anastrophyllum obtusum Herzog ....................................................................................................51 Anastrophyllum piligerum (Nees) Steph. ........................................................................................51 Anastrophyllum squarrosum Herzog ...............................................................................................51 Anastrophyllum stellatum R.M.Schust. ...........................................................................................51 Anastrophyllum tubulosum (Nees) Grolle .......................................................................................51 Andrewsianthus aberrans (Nees et Mont.) Grolle ...........................................................................77 Andrewsianthus bidens (Mitt. ex Steph.) R.M.Schust. ....................................................................77 Andrewsianthus bilobus (Mitt.) Grolle ...........................................................................................77 Andrewsianthus carinatus Grolle ....................................................................................................77 Andrewsianthus cavifolius Grolle et Váňa .......................................................................................77 Andrewsianthus chimbuensis R.M.Schust. ......................................................................................77 Andrewsianthus hodgsoniae (R.M.Schust.) R.M.Schust. ex J.J.Engel ..............................................78 Andrewsianthus kinabaluensis N.Kitag. ..........................................................................................77 Andrewsianthus koponenii Váňa et Piippo ......................................................................................77 Andrewsianthus marionensis (S.W.Arnell) Grolle ...........................................................................78 Andrewsianthus mizutanii N.Kitag. ...............................................................................................77 Andrewsianthus papillosus N.Kitag. ................................................................................................78 Andrewsianthus perigonialis (Hook.f. et Taylor) R.M.Schust. ........................................................78 Andrewsianthus puniceus (Nees) R.M.Schust. ex Grolle .................................................................78 Andrewsianthus recurvifolius (Nees) R.M.Schust. ...........................................................................78 Andrewsianthus scabrellus (C.Massal.) R.M.Schust. ex J.J.Engel ....................................................78 World checklist of hornworts and liverworts ** *** *** * * * *** ** * ** *** * ** * * ** ** ** ** * ** ** * *** * ** ** ** * ** * ** *** *** *** ** *** * ** *** ** ** ** * ** * ** * ** * * * 517 Andrewsianthus sphenoloboides (R.M.Schust.) R.M.Schust. ex J.J.Engel .........................................78 Andrewsianthus sundaicus (Schifn.) R.M.Schust. ..........................................................................78 Andrewsianthus zantenii Váňa .......................................................................................................78 Aneura amboinensis Steph. ..........................................................................................................435 Aneura augustae Steph.................................................................................................................435 Aneura bilora Colenso ................................................................................................................435 Aneura blasioides (Horik.) Furuki ................................................................................................435 Aneura brasiliensis (Ångstr.) Steph...............................................................................................435 Aneura brevissima Steph. .............................................................................................................435 Aneura cerebrata Hewson ............................................................................................................435 Aneura crateriformis Furuki et D.G.Long ....................................................................................435 Aneura crinita C.Massal. .............................................................................................................436 Aneura crumii L.Söderstr., A.Hagborg et von Konrat ..................................................................436 Aneura densa Steph. ....................................................................................................................436 Aneura denticulata Mitt. ex hurn ..............................................................................................436 Aneura eachamensis Hewson........................................................................................................436 Aneura erronea Steph. .................................................................................................................436 Aneura eskuchei Hässel ................................................................................................................436 Aneura gemmifera Furuki ............................................................................................................436 Aneura giangena Hewson ............................................................................................................436 Aneura gibbsiana Steph. ..............................................................................................................436 Aneura glaucescens Steph. ............................................................................................................436 Aneura goebeliana Steph. .............................................................................................................436 Aneura hirsuta Furuki .................................................................................................................436 Aneura hunsteinii Steph. .............................................................................................................436 Aneura imbricata Colenso ...........................................................................................................436 Aneura kaguaensis Hewson ..........................................................................................................436 Aneura keniae Gola .....................................................................................................................436 Aneura latemultiida Steph. .........................................................................................................436 Aneura latissima Spruce...............................................................................................................436 Aneura ledermannii Steph. ..........................................................................................................436 Aneura macrostachya Spruce ........................................................................................................436 Aneura marianensis Furuki ..........................................................................................................436 Aneura maxima (Schifn.) Steph..................................................................................................436 Aneura mirabilis (Malmb.) Wickett et Goinet...........................................................................436 Aneura novaecaledoniae R.M.Schust. ...........................................................................................436 Aneura novaguineensis Hewson....................................................................................................437 Aneura nymannii Steph. ..............................................................................................................437 Aneura pellucida Colenso ............................................................................................................437 Aneura pinguis (L.) Dumort. .......................................................................................................437 Aneura polyantha Colenso ...........................................................................................................437 Aneura punctata Colenso ............................................................................................................437 Aneura rodwayi Hewson .............................................................................................................437 Aneura roraimensis Steph. ...........................................................................................................437 Aneura rotangicola Steph. ............................................................................................................437 Aneura serrulata Gottsche ex Steph. ............................................................................................437 Aneura sharpii Inoue et N.G.Mill. ..............................................................................................437 Aneura singalangana (Schifn.) Steph. .........................................................................................437 Aneura subcanaliculata R.M.Schust. ............................................................................................437 Aneura subledermannii Steph. .....................................................................................................437 Aneura subtenerrima Steph. .........................................................................................................437 Aneura vincentina Steph..............................................................................................................437 518 *** *** ** *** * *** *** * *** *** * ** *** *** ** * ** ** ** *** ** * * *** *** *** ** ** ** * ** * ** * * *** *** *** *** ** * * ** *** ** *** ** ** *** * *** *** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Anoplolejeunea conferta (C.F.W.Meissn. ex Spreng.) A.Evans ......................................................361 Anthelia julacea (L.) Dumort. .......................................................................................................98 Anthelia juratzkana (Limpr.) Trevis. .............................................................................................98 Anthoceros adscendens Lehm. et Lindenb. ......................................................................................27 Anthoceros aethyopicus Gola ...........................................................................................................30 Anthoceros agrestis Paton nom. conserv.(p. 000) Anthoceros alpinus Steph................................................................................................................27 Anthoceros angustifolius Gottsche, Lindenb. et Nees ......................................................................28 Anthoceros angustus Steph. .............................................................................................................28 Anthoceros bharadwajii Udar et A.K.Asthana.................................................................................28 Anthoceros brunnthaleri Steph. ......................................................................................................30 Anthoceros buettneri Steph. ............................................................................................................28 Anthoceros capricornii Cargill et G.A.M.Scott ................................................................................28 Anthoceros caucasicus Steph............................................................................................................28 Anthoceros cavernosus Steph. ..........................................................................................................28 Anthoceros chambensis Kashyap ......................................................................................................28 Anthoceros chungii Khanna ............................................................................................................28 Anthoceros crispatus Grif. ..............................................................................................................28 Anthoceros dimorphus Sim .............................................................................................................28 Anthoceros erectus Kashyap ............................................................................................................28 Anthoceros expansus (Steph.) J.C.Villarreal et Cargill .....................................................................28 Anthoceros ferdinandi-muelleri Steph. ............................................................................................28 Anthoceros loribundus Steph. ........................................................................................................30 Anthoceros fragilis Steph. ...............................................................................................................28 Anthoceros fusiformis Austin ...........................................................................................................28 Anthoceros fusiformis var. taiwanensis J.Haseg. ...............................................................................28 Anthoceros gasongorii Gola .............................................................................................................28 Anthoceros granulatus Gottsche......................................................................................................28 Anthoceros harrisanus (Steph.) Parihar ...........................................................................................28 Anthoceros helmsii Steph. ...............................................................................................................28 Anthoceros jamesonii Taylor ex Mitt. .............................................................................................28 Anthoceros javanicoides H.A.Mill. ..................................................................................................29 Anthoceros jungermannioides Schwein. ...........................................................................................29 Anthoceros kajumas (K.I.Goebel) Prosk. ........................................................................................29 Anthoceros koshii Khanna ..............................................................................................................29 Anthoceros lamellatus Steph. ..........................................................................................................29 Anthoceros laminifer Steph.............................................................................................................29 Anthoceros macounii M.Howe .......................................................................................................29 Anthoceros macrosporus Steph. .......................................................................................................29 Anthoceros maritimus Steph. ..........................................................................................................29 Anthoceros megasporus Meijer.........................................................................................................29 Anthoceros mildbraedii Steph. ........................................................................................................30 Anthoceros muscoides Colenso ........................................................................................................29 Anthoceros myriandroecius Steph. ...................................................................................................29 Anthoceros natalensis Steph. ...........................................................................................................29 Anthoceros neesii Prosk. .................................................................................................................29 Anthoceros niger Steph. ..................................................................................................................29 Anthoceros orizabensis (Steph.) Hässel ............................................................................................29 Anthoceros pandei Udar et A.K.Asthana .........................................................................................29 Anthoceros parvifrons Steph. ..........................................................................................................30 Anthoceros patagonicus Hässel ........................................................................................................29 Anthoceros patagonicus subsp. gremmenii J.C.Villarreal, J.J.Engel et Váňa ......................................29 World checklist of hornworts and liverworts ** ** * *** * * *** *** *** ** ** ** ** *** *** *** *** ** * * *** *** *** *** ** *** ** ** *** *** ** ** ** *** *** *** * *** ** ** ** *** * ** ** ** ** *** *** *** *** ** 519 Anthoceros peruvianus Steph. .........................................................................................................29 Anthoceros pinnatus Steph. ............................................................................................................29 Anthoceros pseudocostus Steph. .......................................................................................................30 Anthoceros punctatus L. ..................................................................................................................29 Anthoceros pusillus Colenso............................................................................................................30 Anthoceros rossoi Gola ....................................................................................................................30 Anthoceros rosulans J.Haseg. ..........................................................................................................30 Anthoceros sambesianus Steph. .......................................................................................................30 Anthoceros scariosus Austin.............................................................................................................30 Anthoceros schroederi Steph. ...........................................................................................................30 Anthoceros serratus Steph. ..............................................................................................................30 Anthoceros simulans M.Howe ........................................................................................................30 Anthoceros spongiosus Steph. ..........................................................................................................30 Anthoceros subtilis Steph. ...............................................................................................................30 Anthoceros telaganus Steph. ............................................................................................................30 Anthoceros tristanianus J.C.Villarreal, J.J.Engel et Váňa .................................................................30 Anthoceros tuberculatus Lehm. et Lindenb. ....................................................................................30 Anthoceros venosus Lindenb. et Gottsche........................................................................................30 Aphanolejeunea lancifera R.M.Schust. .........................................................................................507 Aphanolejeunea minima Tixier ....................................................................................................507 Aphanotropis saxicola Herzog .......................................................................................................317 Aponardia huerlimannii (Váňa et Grolle) Váňa ...........................................................................123 Apotreubia hortoniae Konstant. .....................................................................................................42 Apotreubia nana (S.Hatt. et Inoue) S.Hatt. et Mizut. ....................................................................42 Apotreubia pusilla (R.M.Schust.) Grolle ........................................................................................42 Apotreubia yunnanensis Higuchi ....................................................................................................43 Archeophylla paradoxa R.M.Schust. .............................................................................................253 Archeophylla pungens (Herzog) R.M.Schust. ................................................................................253 Archeophylla schusteri (E.A.Hodgs. et Allison) R.M.Schust. .........................................................253 Archilejeunea abbreviata (Mitt.) Vanden Berghen .......................................................................401 Archilejeunea africana Steph. .......................................................................................................401 Archilejeunea alata Steph.............................................................................................................401 Archilejeunea amakawana Inoue ..................................................................................................402 Archilejeunea auberiana (Mont.) Steph........................................................................................402 Archilejeunea autoica Vanden Berghen.........................................................................................402 Archilejeunea badia (Spruce) Steph..............................................................................................402 Archilejeunea bilabiata (Mitt.) Steph. ..........................................................................................402 Archilejeunea bischleriana Gradst. ................................................................................................402 Archilejeunea bongardii Steph. .....................................................................................................402 Archilejeunea brachyantha J.B.Jack et Steph.................................................................................402 Archilejeunea brevilobula Steph. ..................................................................................................402 Archilejeunea crispistipula (Spruce) Steph. ...................................................................................402 Archilejeunea eberhardtii Steph. ...................................................................................................402 Archilejeunea elobulata Steph.......................................................................................................402 Archilejeunea gradsteinii X.Q.Shi et R.L.Zhu ..............................................................................402 Archilejeunea incrassata Steph. .....................................................................................................402 Archilejeunea jonesii Vanden Berghen..........................................................................................402 Archilejeunea juliformis (Nees) Gradst. ........................................................................................402 Archilejeunea kiushiana (Horik.) Verd. ........................................................................................402 Archilejeunea ludoviciana (De Not.) P.Geissler et Gradst. ...........................................................402 Archilejeunea ludoviciana subsp. porelloides (Spruce) Gradst. .......................................................402 Archilejeunea nebeliana Gradst. et Schäf.-Verw. ..........................................................................403 520 * *** * *** ** *** ** ** *** *** ** * * * * * *** *** *** *** *** *** *** *** * *** *** ** *** ** *** * *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** ** *** *** *** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Archilejeunea negrensis Steph. ......................................................................................................403 Archilejeunea olivacea (Hook.f. et Taylor) Steph..........................................................................403 Archilejeunea ovata Herzog..........................................................................................................403 Archilejeunea parvilora (Nees) Steph. .........................................................................................403 Archilejeunea planifolia (Horik.) Mizut. ......................................................................................403 Archilejeunea planiuscula (Mitt.) Steph........................................................................................403 Arctoscyphus fuegiensis (C.Massal.) Hässel ....................................................................................123 Arctoscyphus ronsmithii Hässel .....................................................................................................123 Arnellia fennica (Gottsche) Lindb. ................................................................................................99 Ascidiota blepharophylla C.Massal. ...............................................................................................417 Ascidiota blepharophylla subsp. alaskana Steere et R.M.Schust. ....................................................417 Aspiromitus asper Schifn. ............................................................................................................507 Aspiromitus bullosus Schifn. ........................................................................................................507 Aspiromitus crenatifrons Steph. ....................................................................................................507 Aspiromitus lobatus Schifn. .........................................................................................................507 Aspiromitus squamulosus Schifn. .................................................................................................507 Asterella abyssinica (Gottsche) Grolle ...........................................................................................477 Asterella africana (Mont.) Underw. ex A.Evans ...........................................................................477 Asterella alpina (Steph.) D.G.Long..............................................................................................480 Asterella australis (Hook.f. et Taylor) Verd. .................................................................................478 Asterella bachmannii (Steph.) S.W.Arnell ....................................................................................478 Asterella blumeana (Nees) Kachroo..............................................................................................477 Asterella bolanderi (Austin) Underw. ...........................................................................................478 Asterella californica (Hampe ex Austin) Underw. .........................................................................481 Asterella caucasica (Steph.) H.Buch, A.Evans et Verd. .................................................................478 Asterella chilensis (Nees et Mont.) A.Evans ..................................................................................477 Asterella conocephala (Steph.) R.M.Schust. ..................................................................................478 Asterella coronata (Steph.) H.A.Mill. ...........................................................................................478 Asterella cruciata (Steph.) Horik. .................................................................................................477 Asterella dioica (Steph.) H.A.Mill. ...............................................................................................478 Asterella dissoluta (Steph.) Grolle .................................................................................................477 Asterella dognyensis H.A.Mill. ......................................................................................................478 Asterella dominicensis S.W.Arnell.................................................................................................477 Asterella drummondii (Taylor) R.M.Schust. ex D.G.Long ...........................................................478 Asterella echinella (Gottsche) Underw..........................................................................................478 Asterella elegans (Spreng.) Trevis. ................................................................................................479 Asterella grollei D.G.Long ...........................................................................................................480 Asterella heterolora (Steph.) H.A.Mill. ........................................................................................479 Asterella innovans (Austin) H.A.Mill. ..........................................................................................479 Asterella khasyana (Grif.) Grolle .................................................................................................477 Asterella lateralis M.Howe ...........................................................................................................479 Asterella leptophylla (Mont.) Grolle .............................................................................................477 Asterella limbata D.G.Long et Grolle ..........................................................................................477 Asterella lindenbergiana (Corda ex Nees) Lindb. ex Arnell ...........................................................479 Asterella linearis (Steph.) M.Howe ..............................................................................................479 Asterella longebarbata (Steph.) H.A.Mill. .....................................................................................479 Asterella macropoda (Spruce) A.Evans ..........................................................................................479 Asterella marginata (Nees) S.W.Arnell .........................................................................................479 Asterella muelleri (Gottsche) R.M.Schust.....................................................................................479 Asterella multilora (Steph.) Kachroo ...........................................................................................479 Asterella muscicola (Steph.) S.W.Arnell ........................................................................................480 Asterella mussuriensis (Kashyap) Verd. .........................................................................................479 World checklist of hornworts and liverworts *** *** *** *** * *** *** *** ** ** *** ** *** *** * *** *** *** *** *** *** * *** * *** *** *** *** *** ** *** *** *** *** *** *** *** *** ** *** *** *** ** *** *** *** ** ** *** *** *** *** 521 Asterella mussuriensis subsp. crassa (Shimizu et S.Hatt.) D.G.Long ..............................................479 Asterella palmeri (Austin) Underw. ..............................................................................................480 Asterella pappii (Gola) Grolle ......................................................................................................479 Asterella persica (Steph.) M.Howe ...............................................................................................479 Asterella preussii (Schifn.) M.Howe ............................................................................................480 Asterella pringlei Underw.............................................................................................................480 Asterella rugosa A.Evans...............................................................................................................480 Asterella saccata (Wahlenb.) A.Evans ...........................................................................................480 Asterella setisquama (Steph.) R.M.Schust. ....................................................................................480 Asterella shimizuana Inoue ..........................................................................................................477 Asterella syngenesica (Bory) Grolle................................................................................................480 Asterella tasmanica (Steph.) R.M.Schust. .....................................................................................480 Asterella tenella (L.) P.Beauv........................................................................................................477 Asterella tenera (Mitt.) R.M.Schust. ............................................................................................478 Asterella tenerrima (Steph.) H.A.Mill. .........................................................................................478 Asterella venosa (Lehm. et Lindenb.) A.Evans ..............................................................................478 Asterella versicolor A.Evans ..........................................................................................................480 Asterella vulcanica (Schifn.) Kachroo et Bapna ...........................................................................480 Asterella wallichiana (Lehm. et Lindenb.) Grolle .........................................................................481 Asterella whiteleggeana (Steph.) R.M.Schust. ...............................................................................480 Asterella wilmsii (Steph.) S.W.Arnell ...........................................................................................480 Athalamia dioica Kashyap ...........................................................................................................484 Athalamia pinguis Falc. ...............................................................................................................484 Athalamia pulcherrima (Steph.) S.Hatt........................................................................................484 Aureolejeunea aurifera R.M.Schust. .............................................................................................304 Aureolejeunea lumae (Herzog) van Slageren .................................................................................304 Aureolejeunea paramicola (Herzog) R.M.Schust. .........................................................................304 Aureolejeunea quinquecarinata R.M.Schust. ................................................................................304 Aureolejeunea tonduzana (Steph.) Gradst.....................................................................................304 Austrolophozia andina R.M.Schust. ...............................................................................................96 Austrolophozia camensis (Steph.) Grolle ex Hässel et Solari ............................................................96 Austrolophozia paradoxa R.M.Schust. ............................................................................................96 Austroriella salta J.Milne et Cargill ..............................................................................................504 Balantiopsis asymmetrica (Herzog) J.J.Engel...................................................................................99 Balantiopsis bisbiida (Steph.) Steph. ...........................................................................................100 Balantiopsis brasiliensis Steph.......................................................................................................100 Balantiopsis cancellata (Nees) Steph. ............................................................................................100 Balantiopsis ciliaris S.Hatt. ..........................................................................................................100 Balantiopsis ciliaris subsp. novoguineensis S.Hatt. .........................................................................100 Balantiopsis convexiuscula Berggr. ................................................................................................100 Balantiopsis crocea Herzog ...........................................................................................................100 Balantiopsis diplophylla (Hook.f. et Taylor) Mitt. ........................................................................100 Balantiopsis diplophylla var. hockenii (Berggr.) J.J.Engel et G.L.Merr. ..........................................100 Balantiopsis erinacea (Hook.f. et Taylor) Mitt. ............................................................................100 Balantiopsis lingulata R.M.Schust................................................................................................100 Balantiopsis montana (Colenso) J.J.Engel et G.L.Merr. ...............................................................100 Balantiopsis neocaledonica Pearson ...............................................................................................100 Balantiopsis paucidens Steph. .......................................................................................................100 Balantiopsis purpurata Mitt. ........................................................................................................100 Balantiopsis rosea Berggr. .............................................................................................................100 Balantiopsis splendens (Steph.) J.J.Engel et G.L.Merr. ..................................................................100 Balantiopsis tumida Berggr. .........................................................................................................100 522 *** *** *** *** *** *** *** *** ** ** ** *** ** *** ** ** *** ** ** ** ** ** ** *** ** ** ** ** ** * ** *** ** ** ** ** *** ** ** *** ** ** *** ** ** ** ** *** ** *** ** *** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Balantiopsis verrucosa J.J.Engel et G.L.Merr. ...............................................................................100 Barbilophozia barbata (Schmidel ex Schreb.) Loeske .....................................................................51 Barbilophozia hatcheri (A.Evans) Loeske........................................................................................51 Barbilophozia lycopodioides (Wallr.) Loeske ...................................................................................51 Barbilophozia rubescens (R.M.Schust. et Damsh.) Kartt. et L.Söderstr. ..........................................51 Barbilophozia sudetica (Nees ex Huebener) L.Söderstr., De Roo et Hedd. .....................................51 Bazzania acanthostipa Spruce ......................................................................................................146 Bazzania accreta (Lehm. et Lindenb.) Trevis. ..............................................................................146 Bazzania acinaciformis Steph.......................................................................................................146 Bazzania acuminata (Lindenb. et Gottsche) Trevis. ....................................................................146 Bazzania acutifolia (Steph.) Schifn. ............................................................................................146 Bazzania adnexa (Lehm. et Lindenb.) Trevis. .............................................................................146 Bazzania adnexa var. aucklandica (Lindenb. et Gottsche) J.J.Engel et G.L.Merr. ........................146 Bazzania ainis (Lindenb. et Gottsche) Trevis. ...........................................................................146 Bazzania albifolia Horik. ............................................................................................................146 Bazzania ambigua (Lindenb.) Trevis. ..........................................................................................146 Bazzania amblyphylla Meagher ...................................................................................................146 Bazzania aneityensis (Steph.) Tixier .............................................................................................146 Bazzania angusta (Steph.) Herzog ...............................................................................................146 Bazzania angustifalcata Herzog ...................................................................................................146 Bazzania angustifolia Horik. .......................................................................................................147 Bazzania angustisedens (Steph.) N.Kitag. .....................................................................................147 Bazzania angustistipula N.Kitag. .................................................................................................147 Bazzania appendiculata (Mitt.) S.Hatt. .......................................................................................147 Bazzania approximata Onr..........................................................................................................147 Bazzania arcuata (Lindenb. et Gottsche) Trevis. .........................................................................147 Bazzania arcuata var. mamillosa Gradst. et A.R.Benitez ..............................................................147 Bazzania armatistipula (Steph.) Fulford ......................................................................................147 Bazzania asperrima Steph............................................................................................................147 Bazzania asymmetrica (Steph.) N.Kitag. ......................................................................................147 Bazzania aterrima (Steph.) N.Kitag. ...........................................................................................147 Bazzania aurescens Spruce ...........................................................................................................147 Bazzania avia Meagher ...............................................................................................................147 Bazzania azorica H.Buch et Perss. ..............................................................................................147 Bazzania baldwinii Austin ..........................................................................................................147 Bazzania bernieri (Steph.) Inoue et H.A.Mill. .............................................................................147 Bazzania bescherellei Steph. .........................................................................................................147 Bazzania bhutanica N.Kitag. et Grolle ........................................................................................147 Bazzania bicrenata N.Kitag. ........................................................................................................147 Bazzania bidens (Gottsche et Lindenb.) Trevis. ...........................................................................148 Bazzania bidens var. heterodonta Spruce ......................................................................................148 Bazzania bidentula (Steph.) Yasuda.............................................................................................148 Bazzania bilobata N.Kitag. .........................................................................................................148 Bazzania borneensis N.Kitag........................................................................................................148 Bazzania brasiliensis (Gottsche et Lindenb.) Trevis. ....................................................................148 Bazzania brighamii (Austin) A.Evans ..........................................................................................148 Bazzania cadens N.Kitag. ............................................................................................................148 Bazzania calcarata (Sande Lac.) Schifn. .....................................................................................148 Bazzania callida (Sande Lac. ex Steph.) Abeyw. ..........................................................................148 Bazzania canelensis (Steph.) Fulford ............................................................................................148 Bazzania caudata (Steph.) Herzog ..............................................................................................148 Bazzania caudistipula (Steph.) Inoue et H.A.Mill. ......................................................................148 World checklist of hornworts and liverworts ** ** *** ** *** ** ** * ** ** ** ** ** * ** ** * ** ** *** * ** ** *** ** ** ** ** ** *** ** ** *** ** ** ** ** * *** ** ** ** ** ** ** ** ** *** ** *** *** ** 523 Bazzania ceylanica (Mitt.) Steph. ................................................................................................148 Bazzania chilensis (Steph.) Fulford ..............................................................................................148 Bazzania chimantensis Fulford.....................................................................................................148 Bazzania cincinnata (De Not.) Trevis. ........................................................................................149 Bazzania citharodes Meagher .......................................................................................................149 Bazzania combinata (J.B.Jack et Steph.) Steph. ...........................................................................149 Bazzania commutata (Lindenb. et Gottsche) Schifn. ..................................................................149 Bazzania comorensis Steph...........................................................................................................149 Bazzania confertifolia (Steph.) Herzog.........................................................................................149 Bazzania conistipula (Steph.) H.A.Mill. ......................................................................................149 Bazzania conophylla (Sande Lac.) Schifn. ...................................................................................149 Bazzania consanguinea (Hampe et Lindenb.) Trevis. ...................................................................149 Bazzania consociata (Steph.) H.A.Mill. .......................................................................................149 Bazzania corbieri (Steph.) Meagher .............................................................................................149 Bazzania crassidentata Fulford.....................................................................................................149 Bazzania crassitexta Steph. ..........................................................................................................149 Bazzania crenata (Trevis.) Trevis. ...............................................................................................149 Bazzania cubensis (Gottsche ex Steph.) Pagán .............................................................................149 Bazzania cucullata Onr. ..............................................................................................................150 Bazzania cuneistipula (Gottsche et Lindenb.) Trevis. ..................................................................150 Bazzania curvidens Steph. ...........................................................................................................150 Bazzania debilis N.Kitag. ............................................................................................................150 Bazzania deciduifolia Onr. ..........................................................................................................150 Bazzania decrescens (Lehm. et Lindenb.) Trevis. ..........................................................................150 Bazzania decrescens var. dentistipula Kiaer et Pearson...................................................................150 Bazzania decrescens subsp. molleri (Steph.) E.W.Jones .................................................................150 Bazzania decrescens subsp. pumila (Mitt.) Pócs ............................................................................150 Bazzania densa (Sande Lac.) Schifn............................................................................................150 Bazzania densa var. connata (Sande Lac.) Schifn. .......................................................................150 Bazzania denticulata (Lindenb. et Gottsche) Trevis. ...................................................................150 Bazzania denticulifera Mägd. ......................................................................................................150 Bazzania denudata (Lindenb. et Gottsche) Trevis. ......................................................................150 Bazzania deplanchei (Gottsche) Jovet-Ast ....................................................................................150 Bazzania deplanchei var. ilamentosa Tixier ..................................................................................150 Bazzania desciscens (Steph.) Meijer ..............................................................................................150 Bazzania didericiana (Gottsche ex Steph.) Steph. ........................................................................151 Bazzania diminuta Herzog..........................................................................................................151 Bazzania distans (Nees) Trevis. ...................................................................................................151 Bazzania diversicuspis Spruce ......................................................................................................151 Bazzania drepanophylla Herzog ...................................................................................................151 Bazzania dulitensis Herzog ..........................................................................................................151 Bazzania dulongensis L.P.Zhou et Li Zhang ................................................................................151 Bazzania eggersiana (Steph.) Pagán .............................................................................................151 Bazzania elmeri (Steph.) N.Kitag. ...............................................................................................151 Bazzania elongata Fulford ...........................................................................................................151 Bazzania emarginata (Steph.) C.M.Cooke ..................................................................................151 Bazzania engelii Glenny ..............................................................................................................151 Bazzania erosa (Reinw., Blume et Nees) Trevis. ..........................................................................151 Bazzania erosa var. pulopenangensis (Lindenb. et Gottsche) Schifn. ............................................151 Bazzania exempta J.J.Engel .........................................................................................................151 Bazzania falcata (Lindenb.) Trevis. .............................................................................................151 Bazzania falcifolia (Steph.) H.A.Mill. .........................................................................................151 524 * *** ** ** *** ** ** * ** *** ** *** *** *** ** ** ** ** ** ** ** ** ** ** ** *** *** *** ** ** ** ** ** ** ** * ** ** ** ** ** *** ** ** ** ** ** ** * ** ** ** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Bazzania fallax (Sande Lac.) Schifn. ..........................................................................................152 Bazzania fasciculata (Steph.) Meagher.........................................................................................152 Bazzania fauriana (Steph.) S.Hatt. .............................................................................................152 Bazzania iliformis Steph. ............................................................................................................152 Bazzania laccida (Dumort.) Grolle.............................................................................................152 Bazzania lavescens (Sande Lac. ex Steph.) Schifn. ......................................................................152 Bazzania leischeri (Steph.) Abeyw. .............................................................................................152 Bazzania francana (Steph.) N.Kitag. ...........................................................................................152 Bazzania friabilis N.Kitag. et T.Kodama.....................................................................................152 Bazzania fuhreri Meagher ...........................................................................................................152 Bazzania fuscescens A.Evans .........................................................................................................152 Bazzania gamscottii Meagher.......................................................................................................152 Bazzania gedeana (Steph.) Meijer................................................................................................152 Bazzania gracilis (Hampe et Gottsche) Steph. .............................................................................152 Bazzania grandiretis (Steph.) Herzog ..........................................................................................152 Bazzania griithiana (Steph.) Mizut. ..........................................................................................152 Bazzania gunniana (Steph.) H.A.Mill. ........................................................................................152 Bazzania hainanensis L.P.Zhou et Li Zhang................................................................................153 Bazzania halconiensis (Steph.) N.Kitag. .......................................................................................153 Bazzania hamatifolia (Steph.) H.A.Mill. .....................................................................................153 Bazzania harpago (De Not.) Schifn............................................................................................153 Bazzania hebridensis (Steph.) H.A.Mill. ......................................................................................153 Bazzania herminieri (Gottsche ex Steph.) Pagán .........................................................................153 Bazzania herzogiana Meijer .........................................................................................................153 Bazzania heterostipa (Steph.) Fulford ..........................................................................................153 Bazzania himalayana (Mitt.) Schifn. ..........................................................................................153 Bazzania hochstetteri (Reichardt) E.A.Hodgs. ..............................................................................153 Bazzania hookeri (Lindenb.) Trevis. ............................................................................................153 Bazzania horridula Schifn. .........................................................................................................153 Bazzania inaequabilis Steph. .......................................................................................................153 Bazzania inaequitexta Steph. .......................................................................................................153 Bazzania incrassata (Steph.) N.Kitag. ..........................................................................................153 Bazzania indica (Gottsche et Lindenb.) Trevis. ...........................................................................153 Bazzania indigenarum (Steph.) N.Kitag. .....................................................................................154 Bazzania insignis (De Not.) Trevis. .............................................................................................154 Bazzania intermedia (Gottsche et Lindenb.) Trevis. ....................................................................154 Bazzania intermedia var. sarawakiana (De Not.) Schifn. ............................................................154 Bazzania involuta (Mont.) Trevis................................................................................................154 Bazzania involuta var. submutica (Lindenb. et Gottsche) J.J.Engel et G.L.Merr. .........................154 Bazzania involutiformis (De Not.) Trevis. ...................................................................................154 Bazzania irregularis (Steph.) Schifn............................................................................................154 Bazzania jamaicensis (Lehm. et Lindenb.) Trevis. .......................................................................154 Bazzania japonica (Sande Lac.) Lindb. ........................................................................................154 Bazzania japonica var. sumatrana Herzog ....................................................................................154 Bazzania javanica (Sande Lac.) Schifn. ......................................................................................154 Bazzania kernii Steph. ................................................................................................................154 Bazzania kokawana N.Kitag. et T.Kodama .................................................................................154 Bazzania latifolia Steph...............................................................................................................154 Bazzania lehmanniana (Lindenb.) Trevis. ...................................................................................155 Bazzania leratii (Beauverd) H.A.Mill. .........................................................................................155 Bazzania lessonii (Steph.) H.A.Mill. ............................................................................................155 Bazzania levieri (Steph.) N.Kitag. ...............................................................................................155 World checklist of hornworts and liverworts ** ** ** *** ** *** ** *** *** ** ** ** ** ** ** ** ** * ** * ** * ** ** ** ** ** ** *** ** *** *** ** ** ** ** ** ** ** ** ** ** *** ** ** ** ** ** *** ** ** ** 525 Bazzania liebmaniana (Lindenb. et Gottsche) Trevis. .................................................................155 Bazzania linearis Herzog .............................................................................................................155 Bazzania linguiformis (Sande Lac.) Trevis. ..................................................................................155 Bazzania longa (Nees) Trevis. .....................................................................................................155 Bazzania longa var. papillata (Steph.) Fulford .............................................................................155 Bazzania longicaulis (Sande Lac.) Schifn. ...................................................................................155 Bazzania longicaulis var. latiareata Herzog ..................................................................................155 Bazzania longistipula (Lindenb.) Trevis. ......................................................................................155 Bazzania loricata (Reinw., Blume et Nees) Trevis. ......................................................................155 Bazzania lowii (Sande Lac. ex Steph.) Schifn. ............................................................................155 Bazzania luzonensis (Steph.) Del Ros. .........................................................................................156 Bazzania macgregorii Steph. ........................................................................................................156 Bazzania macrostipula Fulford ....................................................................................................156 Bazzania magna Horik................................................................................................................156 Bazzania magnistipula N.Kitag. ..................................................................................................156 Bazzania malaccensis (Steph.) Tixier ...........................................................................................156 Bazzania manillana (Gottsche ex Steph.) S.Hatt. ........................................................................156 Bazzania marginata (Steph.) N.Kitag. .........................................................................................156 Bazzania marginella (Herzog) N.Kitag. et T.Kodama .................................................................156 Bazzania mascarena (Steph.) Herzog ...........................................................................................156 Bazzania mayebarae S.Hatt. ........................................................................................................156 Bazzania menzelii E.D.Cooper ...................................................................................................156 Bazzania merrillana (Steph.) Inoue ex Bonner ............................................................................156 Bazzania minuta (Austin) A.Evans ..............................................................................................156 Bazzania minutidens (Steph.) Inoue et H.A.Mill. ........................................................................156 Bazzania minutiserra (Steph.) N.Kitag. .......................................................................................156 Bazzania missionum (Herzog) Jovet-Ast ......................................................................................157 Bazzania mittenii (Steph.) Steph. ................................................................................................157 Bazzania monilinervis (Lehm. et Lindenb.) Trevis.......................................................................157 Bazzania morokensis (Steph.) Grolle ............................................................................................157 Bazzania nitida (F.Weber) Grolle ...............................................................................................157 Bazzania novae-zelandiae (Mitt.) Besch. et C.Massal...................................................................157 Bazzania nudicaulis A.Evans .......................................................................................................157 Bazzania nuuanuensis C.M.Cooke ..............................................................................................157 Bazzania obcuneata (Steph.) H.A.Mill. .......................................................................................157 Bazzania obtusata (Mitt.) Abeyw. ...............................................................................................157 Bazzania okaritana Meagher et Glenny .......................................................................................157 Bazzania orbanii Pócs .................................................................................................................157 Bazzania ovistipula (Steph.) Abeyw. ............................................................................................157 Bazzania pallidevirens (Steph.) Fulford .......................................................................................157 Bazzania papillosa S.W.Arnell .....................................................................................................157 Bazzania paradoxa (Sande Lac.) Steph. .......................................................................................157 Bazzania parisii (Steph.) N.Kitag. ...............................................................................................157 Bazzania parvitexta Steph. ..........................................................................................................158 Bazzania patens (Mont.) Trevis. ..................................................................................................158 Bazzania patentistipa (Sande Lac.) Schifn...................................................................................158 Bazzania paucidens (Steph.) H.A.Mill. ........................................................................................158 Bazzania pearsonii Steph. ............................................................................................................158 Bazzania pectinata (Lindenb. et Gottsche) Schifn. .....................................................................158 Bazzania perfalcata N.Kitag. .......................................................................................................158 Bazzania perrotana E.W.Jones ....................................................................................................158 Bazzania peruviana (Nees) Trevis. ..............................................................................................158 526 *** *** ** ** *** * ** ** ** ** ** ** ** * ** ** ** ** *** ** *** ** ** *** *** ** ** *** * ** * ** ** ** * ** ** ** ** *** * ** ** ** ** ** ** ** *** ** ** * Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Bazzania phyllobola Spruce .........................................................................................................158 Bazzania placophylla (Taylor) Grolle ...........................................................................................158 Bazzania platycnema (Schwägr. ex Steph.) H.A.Mill. ..................................................................158 Bazzania pompeana (Sande Lac.) Mitt. .......................................................................................158 Bazzania praerupta (Reinw., Blume et Nees) Trevis. ...................................................................158 Bazzania praerupta var. obliquata (Nees) Schifn. ........................................................................158 Bazzania praerupta var. thermalis Schifn. ...................................................................................158 Bazzania pseudovittata N.Kitag. et T.Kodama ............................................................................159 Bazzania pusilla (Mitt.) Steph. ....................................................................................................159 Bazzania pycnophylla (Taylor) Trevis. .........................................................................................159 Bazzania quadratistipula H.A.Mill. .............................................................................................159 Bazzania rabenhorstii (Steph.) Abeyw. ........................................................................................159 Bazzania recurva (Mont.) Trevis. ................................................................................................159 Bazzania recurva var. major (Sande Lac.) Schifn. .......................................................................159 Bazzania relexa (Gottsche) Steph. ..............................................................................................159 Bazzania reinwardtii (Sande Lac.) Schifn. ..................................................................................159 Bazzania renistipula Steph. .........................................................................................................159 Bazzania revoluta (Steph.) N.Kitag. ............................................................................................159 Bazzania rimosa Meagher............................................................................................................159 Bazzania roccatii Gola.................................................................................................................159 Bazzania roraimensis (Steph.) Fulford .........................................................................................159 Bazzania sandvicensis (Gottsche ex Steph.) Steph. .......................................................................159 Bazzania sauropoda Meagher ......................................................................................................159 Bazzania scalaris Meagher ...........................................................................................................159 Bazzania schlimiana (Gottsche) Fulford ......................................................................................159 Bazzania schultze-motelii N.Kitag................................................................................................159 Bazzania schusterana N.Kitag. .....................................................................................................160 Bazzania schwaneckiana (Hampe et Gottsche) Trevis..................................................................160 Bazzania scutigera (Nees et Mont.) Trevis. ..................................................................................160 Bazzania semicordata (Lindenb. et Gottsche) Kuntze ..................................................................160 Bazzania serpentina (Nees) Trevis. ..............................................................................................160 Bazzania serrapiculata Inoue et H.A.Mill. ...................................................................................160 Bazzania serrata Fulford .............................................................................................................160 Bazzania serrulatoides Horik. ......................................................................................................160 Bazzania sikkimensis (Steph.) Herzog ..........................................................................................160 Bazzania spinosa S.Okamura .......................................................................................................160 Bazzania spiralis (Reinw., Blume et Nees) Meijer .......................................................................160 Bazzania spruceana Steph............................................................................................................160 Bazzania squarrosa (Steph.) H.A.Mill. ........................................................................................160 Bazzania stolonifera (Sw.) Trevis. ................................................................................................160 Bazzania stolonifera var. granatensis (Gottsche) Fulford ...............................................................160 Bazzania stresemannii (Herzog) N.Kitag. ....................................................................................160 Bazzania subacuta (Mitt.) Steph..................................................................................................161 Bazzania subaequitexta (Steph.) N.Kitag. ....................................................................................161 Bazzania subintegra (Steph.) L.Söderstr. et A.Hagborg ...............................................................161 Bazzania sublonga Fulford...........................................................................................................161 Bazzania subserrifolia (Beauverd) H.A.Mill. ................................................................................161 Bazzania subserrulata A.Evans .....................................................................................................161 Bazzania subtilis (Sande Lac.) Trevis. ..........................................................................................161 Bazzania succulenta N.Kitag. ......................................................................................................161 Bazzania sumatrana (Sande Lac. ex Steph.) Steph. ......................................................................161 Bazzania sumbavensis (Gottsche ex Steph.) Steph........................................................................161 World checklist of hornworts and liverworts *** *** ** *** ** *** ** *** *** ** *** ** *** ** ** ** ** ** *** ** ** ** * ** ** ** ** ** *** *** *** *** *** *** *** *** *** *** ** ** ** *** ** *** *** *** *** *** *** *** *** ** 527 Bazzania taleana (Gottsche) Fulford ...........................................................................................161 Bazzania tayloriana (Mitt.) Kuntze .............................................................................................161 Bazzania temariana (Steph.) H.A.Mill. .......................................................................................161 Bazzania tessellata Meagher .........................................................................................................161 Bazzania tiaoloensis Mizut. et K.C.Chang ...................................................................................161 Bazzania tricrenata (Wahlenb.) Lindb. .......................................................................................161 Bazzania tricrenata var. fulfordiae W.S.Hong ..............................................................................161 Bazzania tridens (Reinw., Blume et Nees) Trevis. .......................................................................162 Bazzania tridens var. assamica (Steph.) Pócs ................................................................................162 Bazzania tridens var. cornutistipula (Steph.) Pócs ........................................................................162 Bazzania trilobata (L.) Gray ........................................................................................................162 Bazzania trilobata var. depauperata (Müll.Frib.) Grolle ...............................................................162 Bazzania uncigera (Reinw., Blume et Nees) Trevis. .....................................................................162 Bazzania uncigera var. brevifolia Herzog .....................................................................................162 Bazzania uncigera var. gibba (Sande Lac.) Meijer ........................................................................162 Bazzania undulata Herzog ..........................................................................................................162 Bazzania vietnamica Pócs............................................................................................................162 Bazzania vitiana Mitt. ................................................................................................................162 Bazzania vittata (Gottsche) Trevis. .............................................................................................162 Bazzania vittata var. luxurians (De Not.) Schifn. .......................................................................162 Bazzania wallichiana (Lindenb.) Trevis. .....................................................................................162 Bazzania watanabei Inoue ..........................................................................................................162 Bazzania wattsiana (Steph.) Meagher..........................................................................................162 Bazzania wiltensii (Sande Lac. ex Steph.) Schifn. .......................................................................163 Bazzania wooroonooran Meagher ................................................................................................163 Bazzania wrightii (Gottsche ex Steph.) Steph..............................................................................163 Bazzania yoshinagana (Steph.) Yasuda.........................................................................................163 Bazzania zollingeri (Lindenb.) Trevis. .........................................................................................163 Bazzania zonulata Meagher ........................................................................................................163 Biantheridion undulifolium (Nees) Konstant. et Vilnet ..................................................................52 Blasia pusilla L. ...........................................................................................................................476 Blepharidophyllum densifolium (Hook.) Ångstr. ex C.Massal. ......................................................103 Blepharidophyllum vertebrale (Gottsche) Ångstr. ex C.Massal. .....................................................103 Blepharolejeunea chimantaensis van Slageren et Kruijt ..................................................................298 Blepharolejeunea fuegiana (Besch. et C.Massal.) Gradst. ..............................................................298 Blepharolejeunea incongrua (Lindenb. et Gottsche) van Slageren et Kruijt ...................................298 Blepharolejeunea saccata (Steph.) van Slageren et Kruijt ...............................................................298 Blepharolejeunea securifolia (Steph.) R.M.Schust. ........................................................................298 Blepharostoma arachnoideum M.Howe ........................................................................................137 Blepharostoma indicum G.Asthana, M.Saxena et Maurya.............................................................137 Blepharostoma minor Horik. ........................................................................................................137 Blepharostoma trichophyllum (L.) Dumort. ..................................................................................137 Blepharostoma trichophyllum subsp. brevirete (Bryhn et Kaal.) R.M.Schust. .................................137 Brachiolejeunea conduplicata (Steph.) Gradst. ..............................................................................298 Brachiolejeunea fernandeziana S.W.Arnell ...................................................................................298 Brachiolejeunea laxifolia (Taylor) Schifn. ....................................................................................298 Brachiolejeunea leiboldiana (Gottsche et Lindenb.) Schifn. .........................................................298 Brachiolejeunea phyllorhiza (Nees) Kruijt et Gradst. ....................................................................298 Brachiolejeunea spruceana (C.Massal.) Schifn. ............................................................................298 Bragginsella anomala R.M.Schust. ...............................................................................................189 Brevianthus lavus (Grolle) J.J.Engel et R.M.Schust.....................................................................137 Brevianthus lavus subsp. crenulatus J.J.Engel...............................................................................137 528 * *** *** *** *** * *** *** *** *** *** *** *** * *** ** ** ** *** ** ** *** *** ** ** ** * ** *** ** ** ** ** ** *** ** *** ** ** ** *** *** ** ** ** *** ** ** ** *** *** ** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Brevianthus hypocanthidium M.A.M.Renner et J.J.Engel .............................................................137 Bromeliophila helenae Gradst. ......................................................................................................350 Bromeliophila natans (Steph.) R.M.Schust...................................................................................351 Bryopteris difusa (Sw.) Nees ........................................................................................................403 Bryopteris ilicina (Sw.) Nees .......................................................................................................403 Bryopteris issiloba Steph. .............................................................................................................403 Bryopteris gaudichaudii Gottsche .................................................................................................403 Bucegia romanica Radian.............................................................................................................488 Calatholejeunea lamii Mizut. .......................................................................................................317 Calatholejeunea paradoxa (Schifn.) K.I.Goebel ...........................................................................317 Calycularia crispula Mitt. ............................................................................................................463 Calycularia laxa Lindb. et Arnell .................................................................................................463 Calypogeia aeruginosa Mitt. .........................................................................................................106 Calypogeia amazonica (Spruce) Steph. .........................................................................................106 Calypogeia andicola Bischl. ..........................................................................................................104 Calypogeia angusta Steph. ............................................................................................................106 Calypogeia annabonensis Steph. ...................................................................................................104 Calypogeia apiculata (Steph.) Steph. ............................................................................................106 Calypogeia arguta Nees et Mont. .................................................................................................104 Calypogeia asakawana S.Hatt. ex Inoue .......................................................................................106 Calypogeia azorica Bischl. ............................................................................................................104 Calypogeia azurea Stotler et Crotz ...............................................................................................104 Calypogeia bidentula (F.Weber) Nees ..........................................................................................104 Calypogeia ceylanica S.Hatt. et Mizut. .........................................................................................106 Calypogeia contracta Inoue ..........................................................................................................106 Calypogeia cuspidata (Steph.) Steph. ............................................................................................106 Calypogeia decurrens (Steph.) Steph. ............................................................................................106 Calypogeia falcata Bischl. .............................................................................................................104 Calypogeia issa (L.) Raddi ...........................................................................................................104 Calypogeia issa subsp. neogaea R.M.Schust..................................................................................104 Calypogeia formosana Horik. .......................................................................................................106 Calypogeia fujisana Inoue ............................................................................................................106 Calypogeia goebelii (Schifn.) Steph..............................................................................................104 Calypogeia goebelii var. siamensis N.Kitag. ...................................................................................104 Calypogeia grandistipula (Steph.) Steph. ......................................................................................104 Calypogeia granulata Inoue..........................................................................................................106 Calypogeia integristipula Steph.....................................................................................................104 Calypogeia japonica Steph............................................................................................................106 Calypogeia khasiana Ajit P.Singh et V.Nath ................................................................................106 Calypogeia latissima Steph. ..........................................................................................................106 Calypogeia laxa Gottsche et Lindenb. ..........................................................................................104 Calypogeia lechleri (Steph.) Steph. ...............................................................................................104 Calypogeia lechleri var. densifolia (Steph.) Bischl. .........................................................................104 Calypogeia longifolia Steph. .........................................................................................................104 Calypogeia lophocoleoides Steph. ...................................................................................................104 Calypogeia lunata Mitt. ...............................................................................................................106 Calypogeia marginella Mitt. .........................................................................................................107 Calypogeia mascarenensis Bischl. ..................................................................................................105 Calypogeia microstipula (Steph.) Steph. .......................................................................................105 Calypogeia miquelii Mont. ex Gottsche, Lindenb. et Nees ...........................................................105 Calypogeia muelleriana (Schifn.) Müll.Frib.................................................................................105 Calypogeia muelleriana subsp. blomquistii R.M.Schust. ...............................................................105 World checklist of hornworts and liverworts *** ** *** ** *** *** ** *** * *** ** *** ** *** ** ** ** ** ** *** ** ** *** *** ** *** *** ** *** ** * *** ** ** ** *** *** *** *** *** *** *** ** *** *** *** *** *** * *** *** *** 529 Calypogeia neesiana (C.Massal. et Carestia) Müll.Frib. ................................................................105 Calypogeia neesiana subsp. subalpina (Inoue) Inoue .....................................................................105 Calypogeia oblata Herzog ............................................................................................................105 Calypogeia obovata R.M.Schust. ..................................................................................................107 Calypogeia peruviana Nees et Mont. ............................................................................................105 Calypogeia rhombifolia (Spruce) Steph. ........................................................................................105 Calypogeia rhombifolia var. colombiana Bischl..............................................................................105 Calypogeia sphagnicola (Arnell et J.Perss.) Warnst. et Loeske .......................................................105 Calypogeia steyermarkii Fulford ...................................................................................................107 Calypogeia subintegra (Gottsche, Lindenb. et Nees) Bischl. .........................................................105 Calypogeia subintegra var. dussiana (Steph.) Bischl.......................................................................105 Calypogeia suecica (Arnell et J.Perss.) Müll.Frib. ..........................................................................106 Calypogeia sullivantii Austin ........................................................................................................104 Calypogeia tenax (Spruce) Steph. .................................................................................................106 Calypogeia tosana (Steph.) Steph. ................................................................................................107 Calypogeia udarii Sudipa Das et D.K.Singh ................................................................................107 Calypogeia uncinulatula Herzog...................................................................................................106 Capillolejeunea geisslerae (Pócs) R.L.Zhu, Qiong He, Y.M.Wei et Pócs .......................................353 Capillolejeunea mascarena S.W.Arnell..........................................................................................353 Castanoclobos julaceus (Hatcher ex J.J.Engel) J.J.Engel et Glenny ................................................254 Caudalejeunea acutifolia Gerola ...................................................................................................404 Caudalejeunea africana (Steph.) Schifn. .....................................................................................404 Caudalejeunea cristiloba (Steph.) Gradst. .....................................................................................404 Caudalejeunea cristiloba subsp. samoana (Steph.) Gradst. ............................................................404 Caudalejeunea dusenii Steph........................................................................................................404 Caudalejeunea grolleana Gradst. ..................................................................................................403 Caudalejeunea hanningtonii (Mitt.) Schifn. ................................................................................404 Caudalejeunea katangensis Vanden Berghen ................................................................................404 Caudalejeunea lehmanniana (Gottsche) A.Evans .........................................................................404 Caudalejeunea lewallei Vanden Berghen ......................................................................................404 Caudalejeunea mauritiana Horik.................................................................................................404 Caudalejeunea reniloba (Gottsche) Steph.....................................................................................404 Caudalejeunea streimannii Gyarmati ...........................................................................................404 Caudalejeunea tridentata R.L.Zhu, Y.M.Wei et Qiong He ..........................................................404 Caudalejeunea yangambiensis (Vanden Berghen) E.W.Jones ........................................................404 Cavicularia densa Steph. .............................................................................................................476 Cephalantholejeunea temnanthoides (R.M.Schust.) R.M.Schust. ..................................................405 Cephalojonesia incuba Grolle et Vanden Berghen ..........................................................................66 Cephalojonesia incuba subsp. mexicana Burghardt, Gradst. et Váňa ...............................................66 Cephalolejeunea parvilobula Mizut...............................................................................................405 Cephalomitrion aterrimum (Steph.) R.M.Schust. ...........................................................................66 Cephalozia acuminata (Herzog) Váňa............................................................................................57 Cephalozia acutiloba (Inoue) Váňa ................................................................................................57 Cephalozia albula Steph. ...............................................................................................................57 Cephalozia ambigua C.Massal. ......................................................................................................57 Cephalozia austrigena R.M.Schust. ex J.J.Engel .............................................................................57 Cephalozia badia (Gottsche) Steph................................................................................................57 Cephalozia bicuspidata (L.) Dumort. .............................................................................................57 Cephalozia bicuspidata subsp. lammersiana (Huebener) R.M.Schust. ............................................57 Cephalozia chilensis (J.J.Engel et R.M.Schust.) R.M.Schust. ..........................................................57 Cephalozia conchata (Grolle et Váňa) Váňa ...................................................................................57 Cephalozia crossii Spruce ...............................................................................................................57 530 *** *** *** *** *** * * * *** *** *** *** *** *** * *** ** * ** ** *** * *** *** *** *** *** *** ** *** *** *** ** ** *** *** *** *** *** *** ** *** ** *** ** ** *** ** ** *** *** ** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Cephalozia darjeelingensis Udar et D.Kumar..................................................................................57 Cephalozia drucei (R.M.Schust.) Váňa ..........................................................................................58 Cephalozia fuegiensis Váňa .............................................................................................................58 Cephalozia hamatiloba Steph. ........................................................................................................58 Cephalozia hamatiloba subsp. siamensis (N.Kitag.) Váňa ...............................................................58 Cephalozia hians Steph. .................................................................................................................59 Cephalozia indica Udar et D.Kumar .............................................................................................59 Cephalozia kodaikanalensis G.Asthana et Saumya Srivast. ..............................................................59 Cephalozia lacinulata (J.B.Jack ex Gottsche et Rabenh.) Spruce ....................................................58 Cephalozia lucens (A.Evans) Steph. ................................................................................................58 Cephalozia macgregorii (Steph.) Váňa ............................................................................................58 Cephalozia macounii (Austin) Austin.............................................................................................58 Cephalozia maxima Steph. ............................................................................................................58 Cephalozia mollusca (De Not.) Váňa .............................................................................................58 Cephalozia neesiana Steph. ............................................................................................................58 Cephalozia nishimurae (N.Kitag.) Váňa .........................................................................................58 Cephalozia pachygyna R.M.Schust. ex J.J.Engel .............................................................................58 Cephalozia parvifolia Arnell...........................................................................................................59 Cephalozia physocaula (Hook.f. et Taylor) Steph. ..........................................................................58 Cephalozia schusteriana J.J.Engel ...................................................................................................58 Cephalozia stolonacea (Herzog) Váňa .............................................................................................58 Cephalozia tricuspidata (Nees) Trevis. ...........................................................................................59 Cephalozia tubulata (Hook.f. et Taylor) Trevis. ............................................................................58 Cephaloziella acanthophora (S.Hatt.) Horik. ..................................................................................71 Cephaloziella aenigmatica R.M.Schust. ..........................................................................................66 Cephaloziella anthelioides S.W.Arnell ............................................................................................66 Cephaloziella antillana (Besch. et Spruce) Fulford .........................................................................70 Cephaloziella arctogena (R.M.Schust.) Konstant. ...........................................................................66 Cephaloziella arenaria (Steph.) R.M.Schust. ..................................................................................66 Cephaloziella aspericaulis Jørg. .......................................................................................................66 Cephaloziella baumgartneri Schifn. ...............................................................................................66 Cephaloziella biokoensis Váňa et Frank Müll. .................................................................................71 Cephaloziella breviperianthia C.Gao ..............................................................................................66 Cephaloziella brinkmannii Douin ..................................................................................................66 Cephaloziella calyculata (Durieu et Mont.) Müll.Frib. ...................................................................70 Cephaloziella capensis (Sim) S.W.Arnell.........................................................................................66 Cephaloziella capillaris (Steph.) Douin ..........................................................................................67 Cephaloziella crassigyna (R.M.Schust.) R.M.Schust. ......................................................................67 Cephaloziella crispata N.Kitag. ......................................................................................................67 Cephaloziella densifolia R.M.Schust. ..............................................................................................67 Cephaloziella densifolia var. dubia R.M.Schust. .............................................................................67 Cephaloziella dentata (Raddi) Steph. .............................................................................................70 Cephaloziella dentifolia Udar et Ad.Kumar ....................................................................................71 Cephaloziella divaricata (Sm.) Schifn. ..........................................................................................67 Cephaloziella divaricata var. scabra (M.Howe) Haynes ..................................................................67 Cephaloziella dusenii Steph. ...........................................................................................................67 Cephaloziella elachista (J.B.Jack ex Gottsche et Rabenh.) Schifn. ..................................................67 Cephaloziella elachista var. spinophylla (C.Gao) C.Gao ..................................................................67 Cephaloziella elegans (Heeg) Schifn. .............................................................................................67 Cephaloziella exigua R.M.Schust. ..................................................................................................67 Cephaloziella exililora (Taylor) Douin ..........................................................................................67 Cephaloziella ilum (Trevis.) Steph. ...............................................................................................71 World checklist of hornworts and liverworts * ** *** *** *** *** *** *** ** *** ** *** ** * *** *** ** *** ** ** *** *** ** *** * *** *** *** ** *** ** ** *** *** *** *** *** ** ** * *** *** * ** *** *** *** ** *** *** ** *** 531 Cephaloziella lexuosa C.Gao et K.C.Chang...................................................................................71 Cephaloziella fragillima (Spruce) Fulford .......................................................................................67 Cephaloziella garsidei S.W.Arnell...................................................................................................67 Cephaloziella granatensis (J.B.Jack ex Steph.) Fulford ....................................................................71 Cephaloziella grandiretis (R.M.Schust.) R.M.Schust. .....................................................................67 Cephaloziella grimsulana (J.B.Jack ex Gottsche et Rabenh.) Lacout. ..............................................68 Cephaloziella grisea R.M.Schust. ...................................................................................................71 Cephaloziella hampeana (Nees) Schifn. ex Loeske .........................................................................68 Cephaloziella hebridensis Steph. .....................................................................................................68 Cephaloziella herzogiana (Pandé et K.P.Srivast.) Udar et D.Kumar ...............................................68 Cephaloziella heteroica (C.M.Cooke) Douin..................................................................................68 Cephaloziella hirta (Steph.) R.M.Schust. .......................................................................................70 Cephaloziella hyalina Douin ..........................................................................................................68 Cephaloziella hyalina var. rappii (Douin) R.M.Schust. ..................................................................68 Cephaloziella inaequalis R.M.Schust..............................................................................................68 Cephaloziella integerrima (Lindb.) Warnst.....................................................................................70 Cephaloziella intricata Schifn. ex Douin .......................................................................................71 Cephaloziella invisa R.M.Schust. ...................................................................................................68 Cephaloziella kilohanensis (C.M.Cooke) Douin .............................................................................68 Cephaloziella levieri Schifn. ex Douin...........................................................................................72 Cephaloziella longii Váňa ...............................................................................................................68 Cephaloziella lycopodioides (Sim) S.W.Arnell .................................................................................68 Cephaloziella mammillifera R.M.Schust. et Damsh. ......................................................................68 Cephaloziella massalongi (Spruce) Müll.Frib. .................................................................................68 Cephaloziella meghalayensis Udar et Ad.Kumar..............................................................................71 Cephaloziella microphylla (Steph.) Douin ......................................................................................71 Cephaloziella muelleriana R.M.Schust. ..........................................................................................68 Cephaloziella natalensis (Sim) S.W.Arnell ......................................................................................68 Cephaloziella nicholsonii Douin .....................................................................................................68 Cephaloziella nothogena R.M.Schust. .............................................................................................68 Cephaloziella obtusilobula R.M.Schust. ..........................................................................................68 Cephaloziella patulifolia (Steph.) Douin ........................................................................................69 Cephaloziella pellucida R.M.Schust. ..............................................................................................72 Cephaloziella phyllacantha (C.Massal. et Carestia) Müll.Frib. ........................................................69 Cephaloziella polystratosa (R.M.Schust. et Damsh.) Konstant. .......................................................69 Cephaloziella pseudocrassigyna R.M.Schust. ex J.J.Engel.................................................................69 Cephaloziella pulcherrima R.M.Schust. ..........................................................................................69 Cephaloziella pulcherrima subsp. sphagnicola R.M.Schust. .............................................................69 Cephaloziella pungens Steph. ex Fulford.........................................................................................69 Cephaloziella pygmaea (Spruce) Váňa ............................................................................................72 Cephaloziella rubella (Nees) Warnst. .............................................................................................69 Cephaloziella schelpei S.W.Arnell ...................................................................................................69 Cephaloziella secundifolia Pearson ..................................................................................................72 Cephaloziella sinensis Douin ..........................................................................................................72 Cephaloziella spegazziniana (C.Massal.) Douin .............................................................................69 Cephaloziella spinicaulis Douin......................................................................................................69 Cephaloziella spinigera (Lindb.) Jørg..............................................................................................69 Cephaloziella squarrosula (Trevis.) Steph. ......................................................................................71 Cephaloziella stellulifera (Taylor ex Carrington et Pearson) Croz. ..................................................69 Cephaloziella stephanii Schifn. ex Douin ......................................................................................69 Cephaloziella stolonifera R.M.Schust..............................................................................................69 Cephaloziella subspinosa R.M.Schust. ............................................................................................71 532 ** *** *** *** *** *** *** *** *** ** ** *** *** *** * ** ** *** *** *** *** *** *** *** *** ** *** ** *** ** *** *** ** *** *** *** *** *** *** *** *** ** *** *** ** *** * * *** * *** ** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Cephaloziella subtilis (Lindenb. et Gottsche) Steph........................................................................69 Cephaloziella sumatrana Schifn. ex Douin ....................................................................................69 Cephaloziella tabularis S.W.Arnell .................................................................................................69 Cephaloziella tenuissima (Lehm.) Steph. ........................................................................................71 Cephaloziella transvaalensis S.W.Arnell..........................................................................................69 Cephaloziella triplicata S.W.Arnell ................................................................................................69 Cephaloziella turneri (Hook.) Müll.Frib. .......................................................................................71 Cephaloziella umtaliensis S.W.Arnell .............................................................................................70 Cephaloziella uncinata R.M.Schust................................................................................................70 Cephaloziella uncinata var. brevigyna R.M.Schust. et Damsh.........................................................70 Cephaloziella uncinata var. sphagnicola R.M.Schust. ......................................................................70 Cephaloziella vaginans Steph. ........................................................................................................70 Cephaloziella varians (Gottsche) Steph. .........................................................................................70 Cephaloziella verrucosa Steph.........................................................................................................70 Cephaloziella villaumei (Steph.) Váňa ............................................................................................70 Cephaloziella violacea Schljakov.....................................................................................................70 Cephaloziella welwitschii (Steph.) Douin .......................................................................................70 Cephaloziopsis intertexta (Gottsche) R.M.Schust............................................................................72 Ceramanus centipes (Lindenb. et Gottsche) E.D.Cooper..............................................................169 Ceramanus clatritexta (Steph.) E.D.Cooper .................................................................................169 Ceramanus elegans (Colenso) E.D.Cooper ...................................................................................169 Ceramanus grossiseta (Steph.) E.D.Cooper...................................................................................169 Ceramanus perfragilis (J.J.Engel et G.L.Merr.) E.D.Cooper.........................................................169 Ceramanus pruinosa (Herzog) E.D.Cooper .................................................................................169 Ceramanus tuberifera (J.J.Engel et R.M.Schust.) E.D.Cooper .....................................................169 Ceratolejeunea aliena Herzog .......................................................................................................303 Ceratolejeunea andringitrae Pócs ..................................................................................................301 Ceratolejeunea atlantica Alvarenga et Ilk.-Borg. ...........................................................................301 Ceratolejeunea belangeriana (Gottsche) Steph. .............................................................................301 Ceratolejeunea beninensis E.W.Jones et Vanden Berghen .............................................................301 Ceratolejeunea brevinervis (Spruce) A.Evans ................................................................................301 Ceratolejeunea ceratantha (Nees et Mont.) Schifn. ......................................................................301 Ceratolejeunea coalita (Ångstr.) Steph. .........................................................................................301 Ceratolejeunea coarina (Gottsche) Schifn. ...................................................................................301 Ceratolejeunea confusa R.M.Schust. .............................................................................................302 Ceratolejeunea cornuta (Lindenb.) Steph. ....................................................................................302 Ceratolejeunea cubensis (Mont.) Schifn. ......................................................................................302 Ceratolejeunea dentistipula Steph. ................................................................................................302 Ceratolejeunea desciscens (Sande Lac.) Schifn. .............................................................................303 Ceratolejeunea fallax (Lehm. et Lindenb.) Bonner .......................................................................302 Ceratolejeunea ilaria (Taylor) Steph. ..........................................................................................302 Ceratolejeunea loribunda Steph. ..................................................................................................302 Ceratolejeunea globulifera Herzog ................................................................................................303 Ceratolejeunea grandiloba J.B.Jack et Steph. ................................................................................303 Ceratolejeunea grandiloba subsp. inlata (Mizut.) Gradst. ............................................................303 Ceratolejeunea guianensis (Nees et Mont.) Steph. ........................................................................302 Ceratolejeunea karstenii Steph......................................................................................................302 Ceratolejeunea kuerschneri Eb.Fisch. et Vanderp. .........................................................................302 Ceratolejeunea laetefusca (Austin) R.M.Schust. ............................................................................302 Ceratolejeunea ledermannii Steph. ...............................................................................................302 Ceratolejeunea malleigera (Spruce) Steph. ....................................................................................302 Ceratolejeunea maranhensis Silva Brito et Ilk.-Borg. .....................................................................302 World checklist of hornworts and liverworts ** *** ** ** *** *** *** *** *** *** * ** *** *** *** *** ** ** *** *** *** *** ** ** *** ** ** *** ** *** *** ** ** *** ** ** *** ** ** ** ** ** ** ** *** ** *** *** ** *** ** ** 533 Ceratolejeunea minor Mizut. ........................................................................................................302 Ceratolejeunea minuta G.Dauphin ..............................................................................................302 Ceratolejeunea moniliata Herzog .................................................................................................302 Ceratolejeunea oculata (Gottsche) Steph. .....................................................................................302 Ceratolejeunea oxygonia Steph......................................................................................................302 Ceratolejeunea papulilora Steph. .................................................................................................303 Ceratolejeunea patentissima (Hampe et Gottsche) A.Evans ..........................................................303 Ceratolejeunea pungens Steph.......................................................................................................303 Ceratolejeunea rubiginosa Steph. ..................................................................................................303 Ceratolejeunea saroltae Pócs .........................................................................................................303 Ceratolejeunea sinensis P.C.Chen et P.C.Wu ...............................................................................303 Ceratolejeunea singapurensis (Lindenb.) Schifn. ..........................................................................303 Ceratolejeunea spinosa (Gottsche) Steph. .....................................................................................303 Ceratolejeunea szyszylowiczii (Loitl.) Steph...................................................................................303 Ceratolejeunea temnantha (Spruce) M.E.Reiner ...........................................................................303 Ceratolejeunea umbonata Steph. ..................................................................................................303 Ceratolejeunea vitiensis Steph.......................................................................................................303 Ceratolejeunea zenkeri Steph........................................................................................................303 Chaetocolea palmata Spruce .........................................................................................................254 Chaetophyllopsis whiteleggei (Carrington et Pearson) R.M.Schust. ex Hamlin ................................72 Chandonanthus squarrosus (Menzies) Mitt. ....................................................................................52 Cheilolejeunea acanthina (Spruce) Gradst. et Ilk.-Borg. ...............................................................305 Cheilolejeunea aciculifera R.M.Schust. .........................................................................................311 Cheilolejeunea acutangula (Nees) Grolle ......................................................................................311 Cheilolejeunea adnata (Kunze ex Lehm.) Grolle ..........................................................................305 Cheilolejeunea adnata var. autoica Gradst. et Ilk.-Borg. ...............................................................305 Cheilolejeunea albovirens (Hook.f. et Taylor) E.A.Hodgs.............................................................305 Cheilolejeunea aneogyna (Spruce) A.Evans ...................................................................................305 Cheilolejeunea ascensionis (Hook.f. et Taylor) Grolle ...................................................................305 Cheilolejeunea asperilora (Spruce) Gradst. et Ilk.-Borg. ...............................................................305 Cheilolejeunea asperrima (Steph.) Grolle ......................................................................................305 Cheilolejeunea australis Solari ......................................................................................................311 Cheilolejeunea baumannii Hürl. ..................................................................................................305 Cheilolejeunea beyrichii (Lindenb.) M.E.Reiner ...........................................................................305 Cheilolejeunea birmensis (Steph.) Mizut. ......................................................................................311 Cheilolejeunea boninensis Mizut. ..................................................................................................305 Cheilolejeunea caducifolia (Gradst. et Schäf.-Verw.) W.Ye et R.L.Zhu .........................................306 Cheilolejeunea camerunensis S.W.Arnell .......................................................................................306 Cheilolejeunea campbelliensis (Steph.) R.M.Schust. ......................................................................311 Cheilolejeunea cedercreutzii (H.Buch et Perss.) Grolle ..................................................................311 Cheilolejeunea celebensis (Steph.) Mizut. ......................................................................................306 Cheilolejeunea ceylanica (Gottsche) R.M.Schust. et Kachroo .......................................................314 Cheilolejeunea chenii R.L.Zhu et M.L.So.....................................................................................306 Cheilolejeunea choachina (Gottsche) Gradst. ................................................................................311 Cheilolejeunea clausa (Nees et Mont.) R.M.Schust. .....................................................................311 Cheilolejeunea clavata Mizut. ......................................................................................................311 Cheilolejeunea clypeata (Schwein.) W.Ye et R.L.Zhu ...................................................................306 Cheilolejeunea comans (Spruce) R.M.Schust. ...............................................................................312 Cheilolejeunea compressa (Herzog) Grolle ....................................................................................306 Cheilolejeunea conchifolia (A.Evans) W.Ye et R.L.Zhu ................................................................306 Cheilolejeunea convexa (S.W.Arnell) S.W.Arnell ..........................................................................312 Cheilolejeunea cookiensis (Steph.) R.M.Schust. et Kachroo...........................................................312 534 *** ** ** * *** *** ** ** ** ** *** *** ** *** * * ** ** ** ** ** ** *** ** ** ** ** * * ** ** ** *** ** *** *** ** *** ** ** ** ** ** *** *** *** *** ** ** * ** *** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Cheilolejeunea cordigera (Steph.) Grolle .......................................................................................306 Cheilolejeunea cordistipula (Steph.) Grolle ex E.W.Jones .............................................................312 Cheilolejeunea coronalis (Gottsche) R.M.Schust. ..........................................................................306 Cheilolejeunea curvatilobula (Herzog) Grolle ...............................................................................306 Cheilolejeunea decursiva (Sande Lac.) R.M.Schust. ......................................................................306 Cheilolejeunea discoidea (Lehm. et Lindenb.) R.M.Schust. et Kachroo ........................................306 Cheilolejeunea diversifolia Augier .................................................................................................306 Cheilolejeunea ecarinata Vanden Berghen ....................................................................................306 Cheilolejeunea erostrata R.M.Schust. ...........................................................................................306 Cheilolejeunea eximia (Jovet-Ast et Tixier) R.L.Zhu et M.L.So....................................................314 Cheilolejeunea exinnovata E.W.Jones ...........................................................................................306 Cheilolejeunea falsinervis (Sande Lac.) R.M.Schust. et Kachroo ...................................................314 Cheilolejeunea ischeri Malombe ..................................................................................................312 Cheilolejeunea fragrantissima (Spruce) R.M.Schust. .....................................................................306 Cheilolejeunea fukiensis (P.C.Chen et P.C.Wu) Piippo ................................................................307 Cheilolejeunea galliotii Steph. ......................................................................................................307 Cheilolejeunea gaoi R.L.Zhu, M.L.So et Grolle............................................................................307 Cheilolejeunea gardneri (Mitt.) Mizut. .........................................................................................314 Cheilolejeunea germanii (Besch. et Spruce) Grolle........................................................................307 Cheilolejeunea ghatensis G.Asthana, S.C.Srivast. et A.K.Asthana ..................................................307 Cheilolejeunea gigantea (Steph.) R.M.Schust. et Kachroo ............................................................314 Cheilolejeunea gottscheana C.J.Bastos ...........................................................................................315 Cheilolejeunea gradsteinii (Grolle et Piippo) W.Ye et R.L.Zhu.....................................................307 Cheilolejeunea grandibracteata Steph. ..........................................................................................307 Cheilolejeunea hallingii B.M.hiers .............................................................................................312 Cheilolejeunea hamlinii Grolle .....................................................................................................307 Cheilolejeunea hawaica Steph. .....................................................................................................307 Cheilolejeunea herzogiana Steph...................................................................................................307 Cheilolejeunea heteroclada (Spruce) Schifn. .................................................................................315 Cheilolejeunea huerlimannii Tixier ..............................................................................................314 Cheilolejeunea hyalomarginata R.L.Zhu et Frank Müll. ...............................................................314 Cheilolejeunea implexicaulis (Hook.f. et Taylor) R.M.Schust. ......................................................307 Cheilolejeunea incisa (Gottsche) R.M.Schust. et Kachroo ............................................................314 Cheilolejeunea incisa var. teretilora B.M.hiers............................................................................314 Cheilolejeunea inlexa (Hampe) Grolle.........................................................................................312 Cheilolejeunea insecta Grolle et Gradst.........................................................................................307 Cheilolejeunea insignis Jovet-Ast et Tixier ....................................................................................307 Cheilolejeunea intertexta (Lindenb.) Steph. ..................................................................................312 Cheilolejeunea intricata (Steph.) J.J.Engel ....................................................................................307 Cheilolejeunea invaginata R.M.Schust. ........................................................................................307 Cheilolejeunea jamaicensis Steph. .................................................................................................307 Cheilolejeunea japonica (Horik.) W.Ye et R.L.Zhu ......................................................................307 Cheilolejeunea kitagawae W.Ye et R.L.Zhu..................................................................................307 Cheilolejeunea krakakammae (Lindenb.) R.M.Schust. .................................................................312 Cheilolejeunea lacerata C.J.Bastos et Gradst.................................................................................307 Cheilolejeunea laciniata D.F.Peralta et M.E.Reiner......................................................................312 Cheilolejeunea laevicalyx (J.B.Jack et Steph.) Grolle .....................................................................307 Cheilolejeunea laeviuscula (Mitt.) Steph. ......................................................................................307 Cheilolejeunea larsenii Mizut. ......................................................................................................308 Cheilolejeunea laurentii Steph. .....................................................................................................308 Cheilolejeunea leptophylla (Ångstr.) Steph. ...................................................................................308 Cheilolejeunea lindenbergii (Gottsche) Mizut. ..............................................................................312 World checklist of hornworts and liverworts ** * ** ** ** ** ** *** ** *** ** ** * ** ** ** *** *** ** *** ** ** ** *** ** ** *** ** ** ** ** ** *** *** ** ** ** ** ** ** ** ** ** ** ** ** ** ** ** *** ** *** 535 Cheilolejeunea longidens (Steph.) R.M.Schust. et Kachroo ...........................................................314 Cheilolejeunea longilora (Taylor) R.M.Schust. ............................................................................308 Cheilolejeunea longispina (Herzog) R.M.Schust. ..........................................................................308 Cheilolejeunea loriana (Steph.) W.Ye et R.L.Zhu ........................................................................308 Cheilolejeunea ludoviciae Steph. ...................................................................................................308 Cheilolejeunea lurida (Lindenb.) Steph. .......................................................................................308 Cheilolejeunea macroloba (Herzog) Grolle ...................................................................................308 Cheilolejeunea malaccensis (G.Hofm.) Xiao L.He .......................................................................308 Cheilolejeunea mammifera R.M.Schust. .......................................................................................308 Cheilolejeunea mariana (Gottsche) B.M.hiers et Gradst. ...........................................................308 Cheilolejeunea mexicana Steph.....................................................................................................308 Cheilolejeunea meyeniana (Nees, Lindenb. et Gottsche) R.M.Schust. et Kachroo ........................314 Cheilolejeunea micholitzii (Steph.) R.M.Schust. et Kachroo.........................................................308 Cheilolejeunea microscypha (Hook.f. et Taylor) M.Wigginton .....................................................309 Cheilolejeunea mimosa (Hook.f. et Taylor) R.M.Schust. ..............................................................312 Cheilolejeunea minutilobula Amakawa .........................................................................................315 Cheilolejeunea mizutanii W.Ye et R.L.Zhu ..................................................................................309 Cheilolejeunea montagnei (Gottsche ex Mont.) R.M.Schust. ........................................................314 Cheilolejeunea nana R.M.Schust. ................................................................................................312 Cheilolejeunea neblinensis Ilk.-Borg. et Gradst. ............................................................................309 Cheilolejeunea ngongensis Malombe et Pócs .................................................................................312 Cheilolejeunea nietneri (Steph.) Mizut. ........................................................................................309 Cheilolejeunea nipponica (S.Hatt.) S.Hatt. ...................................................................................312 Cheilolejeunea norisiae G.Dauphin et Gradst. ..............................................................................309 Cheilolejeunea norrisii (Grolle) M.A.M.Renner ...........................................................................315 Cheilolejeunea novaezelandiae R.M.Schust...................................................................................309 Cheilolejeunea obcordata Herzog ..................................................................................................309 Cheilolejeunea obtruncata (Mont.) Solari .....................................................................................309 Cheilolejeunea obtusa (Herzog) Solari ..........................................................................................309 Cheilolejeunea obtusifolia (Steph.) S.Hatt. ...................................................................................309 Cheilolejeunea obtusilobula (S.Hatt.) S.Hatt. ...............................................................................314 Cheilolejeunea occlusa (Herzog) T.Kodama et N.Kitag. ...............................................................309 Cheilolejeunea omphalogastria Pócs ..............................................................................................313 Cheilolejeunea oncophylla (Ångstr.) Grolle et M.E.Reiner ............................................................313 Cheilolejeunea orientalis (Gottsche) Mizut. ..................................................................................309 Cheilolejeunea ornata C.J.Bastos ..................................................................................................313 Cheilolejeunea oscilla M.A.M.Renner...........................................................................................315 Cheilolejeunea osumiensis (S.Hatt.) Mizut. ...................................................................................313 Cheilolejeunea ovistipula Steph. ...................................................................................................309 Cheilolejeunea panurensis (Spruce) Steph. ....................................................................................309 Cheilolejeunea papillata Solari .....................................................................................................309 Cheilolejeunea papulosa Schifn. ...................................................................................................315 Cheilolejeunea paroica Mizut. ......................................................................................................309 Cheilolejeunea parvidens B.M.hiers ............................................................................................314 Cheilolejeunea pirilora Schifn. ...................................................................................................309 Cheilolejeunea pluriplicata (Pearson) R.M.Schust. .......................................................................313 Cheilolejeunea pocsii E.W.Jones ...................................................................................................313 Cheilolejeunea polyantha A.Evans ................................................................................................313 Cheilolejeunea polyantha var. caduciloba R.M.Schust. ..................................................................313 Cheilolejeunea polystachya (Spruce) Gradst. et Ilk.-Borg. ..............................................................310 Cheilolejeunea renneri (G.Hofm.) Xiao L.He..............................................................................310 Cheilolejeunea revoluta (Herzog) Gradst. et Grolle ......................................................................313 536 *** *** *** *** ** ** ** ** ** ** ** ** * *** ** ** *** *** *** * * *** ** ** ** ** ** ** ** * *** ** * *** ** *** ** ** ** * ** * ** ** ** ** * * ** ** ** ** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Cheilolejeunea rigidula (Nees ex Mont.) R.M.Schust. ..................................................................313 Cheilolejeunea riparia (Steph.) M.E.Reiner..................................................................................315 Cheilolejeunea rotalis (Hook.f. et Taylor) M.Wigginton ..............................................................315 Cheilolejeunea rotundistipula (Lindenb. ex Lehm.) Malombe.......................................................310 Cheilolejeunea rufescens (Lindenb.) Grolle ...................................................................................310 Cheilolejeunea ruwenzorensis (S.W.Arnell) R.M.Schust................................................................313 Cheilolejeunea ryukyuensis Mizut. ................................................................................................313 Cheilolejeunea sandvicensis (Prantl) Steph. ...................................................................................305 Cheilolejeunea stenoschiza (Ångstr.) A.Evans ................................................................................310 Cheilolejeunea streimannii Pócs et Ninh ......................................................................................315 Cheilolejeunea subcrenulata (Spruce) R.M.Schust. .......................................................................310 Cheilolejeunea subopaca (Mitt.) Mizut. ........................................................................................313 Cheilolejeunea suborbicularis (Herzog) H.A.Mill., Bonner et Bischl. ............................................310 Cheilolejeunea suprema (Grolle et Piippo) W.Ye et R.L.Zhu ........................................................310 Cheilolejeunea surrepens (Mitt.) E.W.Jones ..................................................................................310 Cheilolejeunea tenerrima (Steph.) C.J.Bastos................................................................................315 Cheilolejeunea trapezia (Nees) Kachroo et R.M.Schust. ...............................................................315 Cheilolejeunea trifaria (Reinw., Blume et Nees) Mizut. ...............................................................313 Cheilolejeunea turgida (Mitt.) W.Ye et R.L.Zhu ..........................................................................310 Cheilolejeunea udarii G.Asthana, S.C.Srivast. et A.K.Asthana......................................................313 Cheilolejeunea ulugurica Malombe, Eb.Fisch. et Pócs ..................................................................310 Cheilolejeunea unciloba (Lindenb.) Malombe ..............................................................................310 Cheilolejeunea upoluensis S.W.Arnell ...........................................................................................310 Cheilolejeunea urubuensis (Zartman et I.L.Ackerman) R.L.Zhu et Y.M.Wei ................................316 Cheilolejeunea usambarana (Steph.) Grolle ..................................................................................313 Cheilolejeunea valenciae (Steph.) Xiao L.He ................................................................................310 Cheilolejeunea ventricosa (Schifn. ex P.Syd.) Xiao L.He ..............................................................315 Cheilolejeunea verrucosa Steph. ....................................................................................................313 Cheilolejeunea virescens (Lehm. et Lindenb.) Steph. .....................................................................311 Cheilolejeunea viridis Steph. ........................................................................................................311 Cheilolejeunea vittata (Steph. ex G.Hofm.) R.M.Schust. et Kachroo ..........................................315 Cheilolejeunea warnstorii (Steph.) Solari ....................................................................................311 Cheilolejeunea wrightii Steph. ......................................................................................................311 Cheilolejeunea xanthocarpa (Lehm. et Lindenb.) Malombe ..........................................................311 Cheilolejeunea xanthophylla (Lindenb.) Steph. .............................................................................311 Chiastocaulon dendroides (Nees) Carl...........................................................................................219 Chiloscyphus acutus Steph. ...........................................................................................................190 Chiloscyphus alpicola J.J.Engel .....................................................................................................190 Chiloscyphus beesleyanus Pearson ..................................................................................................190 Chiloscyphus biidus Schifn. ........................................................................................................190 Chiloscyphus breviculus B.Y.Yang et W.C.Lee ..............................................................................190 Chiloscyphus brevistipulus Steph. ..................................................................................................190 Chiloscyphus chinnarensis Manju, K.P.Rajesh et Madhus. ............................................................190 Chiloscyphus confertifolius Steph...................................................................................................190 Chiloscyphus confertus Steph. .......................................................................................................190 Chiloscyphus cornutistipulus Steph. ...............................................................................................190 Chiloscyphus durus (Steph.) Hässel...............................................................................................190 Chiloscyphus ernstianus Steph. .....................................................................................................190 Chiloscyphus etesseanus Steph. ......................................................................................................190 Chiloscyphus francanus Steph. ......................................................................................................190 Chiloscyphus graefeanus Steph. ....................................................................................................190 Chiloscyphus greenwelliae (H.A.Mill.) H.A.Mill. ..........................................................................190 World checklist of hornworts and liverworts ** * ** ** ** ** ** ** ** ** ** ** *** * *** * ** ** ** *** ** ** ** ** ** ** ** ** * ** * ** ** ** ** ** ** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 537 Chiloscyphus hookeri J.J.Engel ......................................................................................................190 Chiloscyphus hookeri var. constantifolius J.J.Engel .........................................................................190 Chiloscyphus integerrimus Schifn. ................................................................................................190 Chiloscyphus kashyapii A.Srivast. et S.C.Srivast. ...........................................................................189 Chiloscyphus kehdingianus (Steph.) N.Kitag. ................................................................................190 Chiloscyphus kilauensis Steph. ......................................................................................................190 Chiloscyphus koeppensis (Gottsche) Steph. ....................................................................................190 Chiloscyphus laceratus Steph.........................................................................................................191 Chiloscyphus lambertonii H.A.Mill...............................................................................................191 Chiloscyphus latistipus Steph. .......................................................................................................191 Chiloscyphus lepervanchei (Steph.) J.J.Engel et R.M.Schust. .........................................................191 Chiloscyphus longiissus Steph. ......................................................................................................191 Chiloscyphus pallescens (Ehrh.) Dumort. ......................................................................................189 Chiloscyphus pallescens var. fragilis (Roth) Müll.Frib. ...................................................................189 Chiloscyphus polyanthos (L.) Corda ..............................................................................................189 Chiloscyphus polyanthos var. rivularis (Schrad.) Lindb. et Arnell ...................................................189 Chiloscyphus propagulifer Schifn. ................................................................................................191 Chiloscyphus purpureus Steph.......................................................................................................191 Chiloscyphus quadricilius Steph. ...................................................................................................191 Chiloscyphus quadridentatus (Spruce) J.J.Engel et R.M.Schust. ....................................................191 Chiloscyphus rotundifolius Mitt. ...................................................................................................191 Chiloscyphus rotundiphyllus H.A.Mill...........................................................................................191 Chiloscyphus scaberulus Spruce .....................................................................................................191 Chiloscyphus septatus J.J.Engel .....................................................................................................191 Chiloscyphus similis Steph. ...........................................................................................................191 Chiloscyphus skottsbergianus H.A.Mill. .........................................................................................191 Chiloscyphus subacuminatus Herzog .............................................................................................191 Chiloscyphus subsimilis Steph. ......................................................................................................191 Chiloscyphus tridens Steph. ..........................................................................................................191 Chiloscyphus trigonifolius Steph....................................................................................................191 Chiloscyphus venustulus Colenso ..................................................................................................191 Chonecolea acutiloba (Schifn.) R.M.Schust. ................................................................................138 Chonecolea andina Grolle et Váňa ...............................................................................................138 Chonecolea doellingeri (Nees) Grolle ............................................................................................138 Chonecolea ruwenzorensis E.W.Jones ...........................................................................................138 Chonecolea schusteri Udar et Ad.Kumar .......................................................................................138 Chonecolea verae Potemkin..........................................................................................................138 Clandarium clandestinum (Mont.) R.M.Schust. ..........................................................................103 Clandarium gottscheanum (Grolle) R.M.Schust. ..........................................................................103 Clandarium xiphophyllum (Grolle) R.M.Schust. ..........................................................................103 Clasmatocolea bisexualis Glenny et J.J.Engel ................................................................................191 Clasmatocolea crassiretis (Herzog) Grolle......................................................................................191 Clasmatocolea ctenophylla (Schifn.) Grolle ..................................................................................191 Clasmatocolea cucullistipula (Steph.) Grolle .................................................................................192 Clasmatocolea fasciculata (Nees) Grolle ........................................................................................192 Clasmatocolea fulvella (Hook.f. et Taylor) Grolle.........................................................................192 Clasmatocolea gayana (Mont.) Grolle ..........................................................................................192 Clasmatocolea humilis (Hook.f. et Taylor) Grolle ........................................................................192 Clasmatocolea humilis var. polymorpha J.J.Engel...........................................................................192 Clasmatocolea humilis var. suspecta (C.Massal.) J.J.Engel .............................................................192 Clasmatocolea inlexispina (Hook.f. et Taylor) J.J.Engel...............................................................192 Clasmatocolea marginata (Steph.) Grolle .....................................................................................192 538 *** *** *** *** *** *** *** *** *** *** *** *** *** *** * * *** *** ** ** *** ** *** *** ** ** *** ** ** * *** ** *** *** *** *** ** *** *** ** * ** * ** *** ** ** *** * ** ** *** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Clasmatocolea minutiretis J.J.Engel et Grolle................................................................................192 Clasmatocolea moniliformis J.J.Engel............................................................................................192 Clasmatocolea navistipula (Steph.) Grolle ....................................................................................192 Clasmatocolea navistipula var. parceramosa J.J.Engel ....................................................................192 Clasmatocolea notophylla (Hook.f. et Taylor) Grolle ....................................................................192 Clasmatocolea obvoluta (Hook.f. et Taylor) Grolle .......................................................................192 Clasmatocolea obvoluta var. cookiana (C.Massal.) J.J.Engel ..........................................................192 Clasmatocolea puccioana (De Not.) Grolle ...................................................................................193 Clasmatocolea rigens (Hook.f. et Taylor) J.J.Engel .......................................................................193 Clasmatocolea strongylophylla (Hook.f. et Taylor) Grolle ..............................................................193 Clasmatocolea trachyopa (Hook.f. et Taylor) Grolle .....................................................................193 Clasmatocolea vermicularis (Lehm.) Grolle...................................................................................193 Clasmatocolea verrucosa J.J.Engel .................................................................................................193 Clevea hyalina (Sommerf.) Lindb. ...............................................................................................484 Clevea hyalina var. californica M.Howe .......................................................................................484 Clevea pedicellata (Grif.) Lindb. .................................................................................................484 Clevea pusilla (Steph.) Rubas. et D.G.Long.................................................................................484 Clevea spathysii (Lindenb.) Müll.Frib. .........................................................................................484 Cololejeunea abnormis Mizut. ......................................................................................................332 Cololejeunea acuminata Mizut. ....................................................................................................325 Cololejeunea adhaesiva (Mitt.) R.M.Schust. .................................................................................332 Cololejeunea adnata Tixier...........................................................................................................332 Cololejeunea aequabilis (Sande Lac.) Schifn. ...............................................................................326 Cololejeunea africana (Steph.) R.M.Schust. .................................................................................332 Cololejeunea albodentata P.C.Chen et P.C.Wu ............................................................................320 Cololejeunea altimontana Pócs .....................................................................................................326 Cololejeunea amaniensis Pócs .......................................................................................................326 Cololejeunea ambeliensis Tixier ....................................................................................................332 Cololejeunea amieuensis Tixier .....................................................................................................332 Cololejeunea amoena Benedix ......................................................................................................337 Cololejeunea amphibola B.M.hiers .............................................................................................324 Cololejeunea andapania Tixier .....................................................................................................332 Cololejeunea angulata (Steph.) Mizut. .........................................................................................332 Cololejeunea angustibracteata Schifn. ex P.Syd. ...........................................................................326 Cololejeunea angustilora (Steph.) Mizut. .....................................................................................323 Cololejeunea ankaiana Tixier .......................................................................................................332 Cololejeunea antillana Pócs..........................................................................................................324 Cololejeunea apiculata (E.W.Jones) R.M.Schust. .........................................................................326 Cololejeunea appressa (A.Evans) Benedix......................................................................................337 Cololejeunea armata Tixier ..........................................................................................................320 Cololejeunea arrectifolia (Mitt.) Steph..........................................................................................326 Cololejeunea attilana Pócs............................................................................................................332 Cololejeunea augieri Tixier...........................................................................................................324 Cololejeunea aurantia (Tixier) houvenot....................................................................................326 Cololejeunea auriculata (E.W.Jones) R.M.Schust.........................................................................332 Cololejeunea australis Tixier .........................................................................................................319 Cololejeunea autoica (Steph.) Grolle ............................................................................................332 Cololejeunea azorica V.Allorge et Jovet-Ast ..................................................................................324 Cololejeunea bachmaensis Tixier ...................................................................................................338 Cololejeunea bandamiae Tixier ....................................................................................................326 Cololejeunea bebourensis Tixier ....................................................................................................326 Cololejeunea bekkeri Tixier ..........................................................................................................332 World checklist of hornworts and liverworts ** *** *** ** ** ** *** *** ** * *** ** ** * * ** *** ** *** * ** *** *** *** *** * *** *** * ** *** *** *** ** ** *** ** ** *** *** *** * ** ** ** * ** *** ** ** ** * 539 Cololejeunea bergmansiana Tixier ................................................................................................326 Cololejeunea berneckerae Pócs ......................................................................................................317 Cololejeunea bhutanica Grolle et Mizut. ......................................................................................320 Cololejeunea biddlecomiae (Austin) A.Evans.................................................................................321 Cololejeunea bidentula (Steph.) E.W.Jones ..................................................................................326 Cololejeunea bifalcata Pócs ..........................................................................................................326 Cololejeunea bischleriana Tixier ...................................................................................................332 Cololejeunea blepharophylla Pócs..................................................................................................326 Cololejeunea bolovenensis Tixier ...................................................................................................332 Cololejeunea bontocensis Tixier .....................................................................................................338 Cololejeunea borbonica Tixier ......................................................................................................332 Cololejeunea borhidiana Pócs .......................................................................................................326 Cololejeunea bosseriana Tixier ......................................................................................................326 Cololejeunea brunelii Tixier .........................................................................................................333 Cololejeunea caihuaella But et P.C.Wu ........................................................................................321 Cololejeunea calcarata E.W.Jones ................................................................................................321 Cololejeunea calcarea (Lib.) Steph. ...............................................................................................326 Cololejeunea caledonica Gottsche .................................................................................................319 Cololejeunea camillii (Lehm.) A.Evans .........................................................................................324 Cololejeunea camusii Tixier..........................................................................................................326 Cololejeunea capuronii Tixier .......................................................................................................321 Cololejeunea cardiocarpa (Mont.) A.Evans ...................................................................................333 Cololejeunea ceatocarpa (Ångstr.) Steph. ......................................................................................326 Cololejeunea ceratilobula (P.C.Chen) R.M.Schust. ......................................................................319 Cololejeunea ceylanica Onr. .........................................................................................................326 Cololejeunea chamlongiana Tixier ................................................................................................327 Cololejeunea chenii Tixier ............................................................................................................327 Cololejeunea chinii Tixier ............................................................................................................320 Cololejeunea chittagongensis Tixier ...............................................................................................333 Cololejeunea chuahiana Pócs........................................................................................................337 Cololejeunea ciliata Pócs ..............................................................................................................327 Cololejeunea cingens (Herzog) Bernecker et Pócs..........................................................................317 Cololejeunea cocoscola Tixier ........................................................................................................333 Cololejeunea comptonii (Pearson) H.A.Mill..................................................................................327 Cololejeunea conchifolia (Gottsche) Gradst. .................................................................................340 Cololejeunea contractiloba A.Evans...............................................................................................324 Cololejeunea cookei A.Evans .........................................................................................................327 Cololejeunea cordilora Steph. ......................................................................................................327 Cololejeunea cornuta E.W.Jones...................................................................................................324 Cololejeunea cornutissima (R.M.Schust.) Stotler et Crand.-Stotl. .................................................317 Cololejeunea costaricensis (Bern.-Lück.) Bernecker et Pócs............................................................317 Cololejeunea crassipapillata Tixier ................................................................................................338 Cololejeunea crateris Pócs.............................................................................................................327 Cololejeunea cremersii Tixier ........................................................................................................333 Cololejeunea crenata (A.Evans) Pócs ............................................................................................324 Cololejeunea crenulata (Pearson) H.A.Mill. .................................................................................327 Cololejeunea cristata (Steph.) R.M.Schust. ...................................................................................333 Cololejeunea cubensis Pócs ...........................................................................................................317 Cololejeunea cucullifolia (Herzog) E.A.Hodgs. .............................................................................327 Cololejeunea cuneata (Lehm. et Lindenb.) Herzog .......................................................................333 Cololejeunea cuneifolia Steph. ......................................................................................................327 Cololejeunea dadeuniana Tixier ...................................................................................................327 540 ** ** ** ** ** ** * * *** * *** *** ** ** *** ** *** ** *** ** * *** * *** *** ** ** * *** *** ** *** ** ** *** ** * ** ** * ** *** * ** ** ** * ** ** ** ** * Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Cololejeunea dankiaensis Tixier ....................................................................................................340 Cololejeunea dauphinii R.L.Zhu ..................................................................................................337 Cololejeunea decemplicata (Steph.) Tixier.....................................................................................327 Cololejeunea decliviloba Steph......................................................................................................327 Cololejeunea dentata (E.W.Jones) R.M.Schust. ............................................................................327 Cololejeunea denticulata (Horik.) S.Hatt. ....................................................................................327 Cololejeunea dentilobula (Steph.) R.M.Schust..............................................................................327 Cololejeunea deroinii Tixier .........................................................................................................333 Cololejeunea desciscens Steph. .......................................................................................................319 Cololejeunea deslooveri Vanden Berghen ......................................................................................333 Cololejeunea dianae M.Wigginton ...............................................................................................327 Cololejeunea diaphana A.Evans....................................................................................................324 Cololejeunea dilatata (Steph.) Mizut. ...........................................................................................328 Cololejeunea dinghuiana R.L.Zhu et Y.F.Wang ...........................................................................321 Cololejeunea diplasiolejeunoides Tixier ..........................................................................................328 Cololejeunea discilora Tixier........................................................................................................337 Cololejeunea distalopapillata (E.W.Jones) R.M.Schust. ................................................................328 Cololejeunea dolichodonta Tixier ..................................................................................................321 Cololejeunea dozyana (Sande Lac.) Schifn. ..................................................................................321 Cololejeunea drepanolejeuneoides (Horik.) R.M.Schust. ................................................................324 Cololejeunea dzumacensis Tixier ...................................................................................................333 Cololejeunea ecuadoriensis Pócs ....................................................................................................333 Cololejeunea efusa (Mitt.) Steph. ................................................................................................328 Cololejeunea elegans Steph. ..........................................................................................................321 Cololejeunea elephantorum Tixier.................................................................................................328 Cololejeunea ellipsoidea R.M.Schust. ............................................................................................328 Cololejeunea ensifera Tixier ..........................................................................................................340 Cololejeunea epiphylla G.Asthana et A.Shukla ..............................................................................333 Cololejeunea equialbi Tixier .........................................................................................................328 Cololejeunea erostrata (Herzog) Bernecker et Pócs .......................................................................324 Cololejeunea eustacei Pócs ............................................................................................................338 Cololejeunea falcata (Horik.) Benedix ..........................................................................................338 Cololejeunea falcata var. madecassa Tixier ....................................................................................338 Cololejeunea falcidentata R.M.Schust. .........................................................................................321 Cololejeunea ilicis (Herzog) Piippo .............................................................................................328 Cololejeunea ilidens Benedix .......................................................................................................321 Cololejeunea ischeri Tixier ...........................................................................................................328 Cololejeunea issilobula Herzog ....................................................................................................333 Cololejeunea lavicans (Steph.) Mizut...........................................................................................328 Cololejeunea lavida P.C.Wu et J.S.Lou .......................................................................................338 Cololejeunea lavovittata Pócs ......................................................................................................338 Cololejeunea loccosa (Lehm. et Lindenb.) Schifn. .......................................................................338 Cololejeunea loccosa var. amoenoides Tixier ..................................................................................338 Cololejeunea loccosa var. angustibracteata Tixier ..........................................................................338 Cololejeunea loccosa var. aurita Benedix ......................................................................................338 Cololejeunea loccosa var. conivens Benedix ...................................................................................338 Cololejeunea loccosa var. ocellata Tixier .......................................................................................338 Cololejeunea loccosa var. plicata Tixier.........................................................................................338 Cololejeunea loccosa var. trivittata Tixier .....................................................................................338 Cololejeunea lorencei Tixier .........................................................................................................333 Cololejeunea foliicola S.C.Srivast. et G.Srivast. .............................................................................333 Cololejeunea frahmii Tixier ..........................................................................................................321 World checklist of hornworts and liverworts * ** *** ** *** ** ** *** ** ** ** *** *** *** *** *** ** *** ** *** *** ** *** ** *** *** *** * ** ** *** *** ** *** *** *** *** ** ** *** ** ** ** ** *** *** *** * *** *** ** ** 541 Cololejeunea fredericii Onr...........................................................................................................328 Cololejeunea fructumarginata Tixier.............................................................................................333 Cololejeunea furcilobulata (Berrie et E.W.Jones) R.M.Schust. ......................................................333 Cololejeunea fusca (Steph.) Mizut. ...............................................................................................328 Cololejeunea geissleriana Tixier ....................................................................................................333 Cololejeunea georgiana Tixier .......................................................................................................334 Cololejeunea gottschei (Steph.) Pandé, K.P.Srivast. et Ahmad .......................................................328 Cololejeunea gracilis (Jovet-Ast) Pócs ...........................................................................................318 Cololejeunea gracilis var. linearifolia (R.M.Schust.) Pócs ..............................................................318 Cololejeunea gradsteinii M.J.Lai et R.L.Zhu.................................................................................325 Cololejeunea gresicola Tixier .........................................................................................................338 Cololejeunea grolleana Pócs ..........................................................................................................321 Cololejeunea grossepapillosa (Horik.) N.Kitag. ..............................................................................318 Cololejeunea grossestyla M.Wigginton ..........................................................................................328 Cololejeunea grushvitzkiana Pócs .................................................................................................328 Cololejeunea guadelupensis Tixier .................................................................................................334 Cololejeunea gynophthalma Benedix .............................................................................................338 Cololejeunea hainanensis R.L.Zhu................................................................................................328 Cololejeunea hamata Steph. .........................................................................................................340 Cololejeunea harrisii Pócs ............................................................................................................328 Cololejeunea haskarliana (Lehm.) Schifn. ...................................................................................321 Cololejeunea hattoriana Mizut. et Pócs ........................................................................................324 Cololejeunea hebridensis Tixier .....................................................................................................334 Cololejeunea herzogii K.I.Goebel .................................................................................................340 Cololejeunea hildebrandii (Austin) Steph. ....................................................................................328 Cololejeunea hinidumae Onr........................................................................................................334 Cololejeunea hirta Steph. .............................................................................................................328 Cololejeunea hoabinhiana Tixier ..................................................................................................334 Cololejeunea hodgsoniae (Herzog) E.A.Hodgs. .............................................................................328 Cololejeunea hoeana Tixier ..........................................................................................................334 Cololejeunea horikawana (S.Hatt.) Mizut. ...................................................................................329 Cololejeunea huerlimannii Tixier .................................................................................................329 Cololejeunea hungii Tixier ...........................................................................................................334 Cololejeunea hyalina G.Asthana et S.C.Srivast. ............................................................................321 Cololejeunea indosinica Tixier ......................................................................................................334 Cololejeunea inlata Steph............................................................................................................338 Cololejeunea inlectens (Mitt.) Benedix .........................................................................................324 Cololejeunea inlexifolia R.M.Schust. ...........................................................................................329 Cololejeunea inoueana Mizut. ......................................................................................................334 Cololejeunea iradieri M.Infante et Heras .....................................................................................329 Cololejeunea irianensis Tixier .......................................................................................................329 Cololejeunea iwatsukiana (Pócs) Pócs ..........................................................................................318 Cololejeunea jamesii (Austin) M.E.Reiner et Pócs ........................................................................340 Cololejeunea japonica (Schifn.) Mizut. ........................................................................................334 Cololejeunea johannis-winkleri (Herzog) R.L.Zhu ........................................................................329 Cololejeunea jonesii Pócs ..............................................................................................................334 Cololejeunea jovetastiana (Pócs) Pócs ...........................................................................................318 Cololejeunea kahuziensis Tixier ....................................................................................................321 Cololejeunea kapingaensis H.A.Mill. ............................................................................................334 Cololejeunea karnatakensis G.Asthana et S.C.Srivast. ...................................................................321 Cololejeunea kegelii Steph. ...........................................................................................................329 Cololejeunea khanii Tixier ...........................................................................................................329 542 ** ** ** * *** ** *** ** ** *** *** ** ** *** *** *** *** *** ** ** *** *** *** ** ** * *** *** ** *** *** *** ** *** ** ** *** * *** ** ** * ** *** * *** ** * *** *** *** *** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Cololejeunea khiavensis Tixier ......................................................................................................338 Cololejeunea kiriromensis Tixier ...................................................................................................334 Cololejeunea kodamae Kamim. ....................................................................................................321 Cololejeunea kohkongensis Tixier ..................................................................................................329 Cololejeunea kolombangarae Pócs .................................................................................................321 Cololejeunea kolombangarae subsp. sepikensis Pócs .......................................................................321 Cololejeunea konratii Pócs ...........................................................................................................321 Cololejeunea koponenii (Pócs) Pócs ..............................................................................................325 Cololejeunea koratensis Tixier.......................................................................................................339 Cololejeunea kuciana Pócs et Schäf.-Verw. ...................................................................................321 Cololejeunea kulenensis Tixier ......................................................................................................334 Cololejeunea lacinulata Benedix ...................................................................................................320 Cololejeunea laevigata (Mitt.) R.M.Schust. ..................................................................................334 Cololejeunea lanceolata E.W.Jones ...............................................................................................325 Cololejeunea lanciloba Steph. .......................................................................................................334 Cololejeunea latilobula (Herzog) Tixier ........................................................................................334 Cololejeunea latistyla R.L.Zhu .....................................................................................................334 Cololejeunea leloutrei (E.W.Jones) R.M.Schust. ...........................................................................334 Cololejeunea leloutrei var. microlobulata Tixier .............................................................................334 Cololejeunea leloutrei var. ulugurica Pócs ex Tixier .......................................................................334 Cololejeunea lemuriana Tixier......................................................................................................334 Cololejeunea lichenyae R.D.Porley, N.G.Hodgetts et M.Wigginton.............................................329 Cololejeunea linopteroides H.Rob. ................................................................................................319 Cololejeunea lisowskii (Pócs) Pócs ................................................................................................318 Cololejeunea littoralis Tixier.........................................................................................................334 Cololejeunea lobulilineata Tixier ..................................................................................................329 Cololejeunea longiana Grolle et Mizut. ........................................................................................321 Cololejeunea longifolia (Mitt.) Benedix ex Mizut. ........................................................................329 Cololejeunea longistylis A.Evans....................................................................................................335 Cololejeunea macounii (Spruce) A.Evans ......................................................................................322 Cololejeunea madeirensis Schifn. .................................................................................................318 Cololejeunea madothecoides (Steph.) Benedix ...............................................................................320 Cololejeunea magillii Pócs ............................................................................................................322 Cololejeunea magna (Tixier) M.Infante et Heras ..........................................................................329 Cololejeunea magniica Pócs .........................................................................................................329 Cololejeunea magnilobula (Horik.) S.Hatt. ..................................................................................329 Cololejeunea magnistyla (Horik.) Mizut. ......................................................................................335 Cololejeunea malaccensis Tixier ....................................................................................................335 Cololejeunea malanjae Steph. .......................................................................................................322 Cololejeunea malayana Tixier ......................................................................................................335 Cololejeunea mamillata (Ångstr.) E.A.Hodgs. ..............................................................................322 Cololejeunea manlinensis Tixier....................................................................................................339 Cololejeunea maquilingensis Tixier ...............................................................................................339 Cololejeunea marginata (Lehm. et Lindenb.) Pearson ..................................................................335 Cololejeunea maritima Tixier .......................................................................................................335 Cololejeunea metzgeriopsis (K.I.Goebel) Gradst., R.Wilson, Ilk.-Borg. et Heinrichs .....................329 Cololejeunea micrandroecia (Spruce) M.Menzel ...........................................................................317 Cololejeunea micronesica H.A.Mill. et Bonner..............................................................................329 Cololejeunea microscopica (Taylor) Schifn. ..................................................................................318 Cololejeunea microscopica var. africana (Pócs) Pócs et Bernecker ..................................................318 Cololejeunea microscopica var. exigua (A.Evans) Pócs....................................................................318 Cololejeunea minuscula Pócs ........................................................................................................318 World checklist of hornworts and liverworts *** ** *** ** *** ** *** ** ** ** ** *** *** * *** ** *** * *** * ** *** *** ** ** ** *** ** *** ** ** *** *** *** ** ** ** *** *** ** *** *** * ** ** ** *** *** *** *** ** ** 543 Cololejeunea minutilobula Herzog ...............................................................................................335 Cololejeunea mizutaniana Udar et G.Srivast. ...............................................................................322 Cololejeunea mocambiquensis S.W.Arnell .....................................................................................322 Cololejeunea mooreaensis Tixier....................................................................................................329 Cololejeunea moralesiae (Bern.-Lück.) Bernecker et Pócs..............................................................325 Cololejeunea moramangae Tixier ..................................................................................................318 Cololejeunea morobensis (Pócs) Pócs.............................................................................................325 Cololejeunea mouensis (Tixier) H.A.Mill. .....................................................................................324 Cololejeunea mutabilis Benedix ....................................................................................................339 Cololejeunea nakajimae S.Hatt. ...................................................................................................339 Cololejeunea nanhutashanensis J.D.Yang et S.H.Lin .....................................................................322 Cololejeunea nigerica (E.W.Jones) R.M.Schust. ...........................................................................335 Cololejeunea nilgiriensis G.Asthana et S.C.Srivast. .......................................................................322 Cololejeunea ninguana Tixier .......................................................................................................329 Cololejeunea norrisii (Pócs) Pócs ..................................................................................................318 Cololejeunea obcordata (Austin) A.Evans......................................................................................329 Cololejeunea obliqua (Nees et Mont.) Schifn. .............................................................................330 Cololejeunea oblongiperianthia (P.C.Wu et J.S.Lou) Piippo .........................................................330 Cololejeunea obtusifolia (E.W.Jones) Tixier..................................................................................325 Cololejeunea obtusifolia var. madecassa (Tixier) Pócs ....................................................................325 Cololejeunea occidentalis (E.W.Jones) Vanden Berghen ...............................................................335 Cololejeunea ocellata (Horik.) Benedix .........................................................................................339 Cololejeunea ocelloides (Horik.) Mizut. ........................................................................................339 Cololejeunea oleana Sim...............................................................................................................330 Cololejeunea ombrophila Tixier ....................................................................................................320 Cololejeunea onraedtii Tixier .......................................................................................................335 Cololejeunea ornata A.Evans ........................................................................................................322 Cololejeunea ovalifolia A.Evans ....................................................................................................330 Cololejeunea paciica Pócs ............................................................................................................335 Cololejeunea panamensis G.Dauphin et Pócs................................................................................335 Cololejeunea panchoana Tixier .....................................................................................................330 Cololejeunea paniensis (Tixier) Grolle ..........................................................................................330 Cololejeunea papilliloba (Steph.) Steph. .......................................................................................325 Cololejeunea papillosa (K.I.Goebel) Mizut. ..................................................................................318 Cololejeunea papulilora Steph. ....................................................................................................330 Cololejeunea papulosa R.M.Schust. ..............................................................................................325 Cololejeunea parva Vanden Berghen ............................................................................................322 Cololejeunea paucifolia (Spruce) Bernecker et Pócs ......................................................................325 Cololejeunea paucimarginata Tixier .............................................................................................335 Cololejeunea pentagona (Mitt.) Steph...........................................................................................330 Cololejeunea peponiformis Mizut. .................................................................................................325 Cololejeunea perainis (Schifn.) Schifn. .....................................................................................339 Cololejeunea perainis var. ciconiae Tixier ....................................................................................339 Cololejeunea perainis var. elegans Benedix...................................................................................339 Cololejeunea perainis var. serrulata Schifn. ex Benedix...............................................................339 Cololejeunea perakensis Tixier ......................................................................................................335 Cololejeunea plagiochiliana Tixier ................................................................................................330 Cololejeunea plagiophylla Benedix ................................................................................................335 Cololejeunea planilora Benedix ...................................................................................................322 Cololejeunea planissima (Mitt.) Abeyw.........................................................................................335 Cololejeunea planissima var. chagosensis Pócs ................................................................................335 Cololejeunea planiuscula Tixier ....................................................................................................330 544 *** *** * ** ** ** *** ** *** ** ** *** *** *** *** * ** ** * *** *** *** ** * ** *** *** * *** ** ** ** *** *** ** *** ** *** ** ** *** *** ** *** ** *** *** ** *** *** *** *** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Cololejeunea platyneura (Spruce) A.Evans ....................................................................................322 Cololejeunea pluridentata P.C.Wu et J.S.Lou ...............................................................................322 Cololejeunea polisiana Tixier ........................................................................................................339 Cololejeunea praeruptorum Tixier ................................................................................................335 Cololejeunea pretiosa Benedix.......................................................................................................322 Cololejeunea producta (Mitt.) S.Hatt. ..........................................................................................335 Cololejeunea pseudocristallina P.C.Chen et P.C.Wu .....................................................................322 Cololejeunea pseudocuspidata Tixier .............................................................................................325 Cololejeunea pseudoloccosa (Horik.) Benedix ...............................................................................339 Cololejeunea pseudoplagiophylla P.C.Wu et J.S.Lou......................................................................322 Cololejeunea pseudoschmidtii Tixier..............................................................................................322 Cololejeunea pseudoserrata Tixier .................................................................................................330 Cololejeunea pseudostephanii Tixier ..............................................................................................339 Cololejeunea pseudostipulata Schifn. ex P.Syd..............................................................................320 Cololejeunea pterocolea Herzog ....................................................................................................325 Cololejeunea pteroporum Tixier ....................................................................................................330 Cololejeunea pulchella (Mitt.) R.M.Schust. ..................................................................................330 Cololejeunea pulchella var. stylifera R.M.Schust. ...........................................................................330 Cololejeunea punctata (Gottsche) Pearson ....................................................................................336 Cololejeunea pusilla Steph. ...........................................................................................................325 Cololejeunea quadridentata (S.Hatt.) Grolle.................................................................................330 Cololejeunea raduliloba Steph. .....................................................................................................336 Cololejeunea ramromensis Pócs.....................................................................................................322 Cololejeunea reineckeana Steph. ...................................................................................................336 Cololejeunea retusula (Mitt.) H.A.Mill.........................................................................................330 Cololejeunea rosellata Mizut. ........................................................................................................322 Cololejeunea rossettiana (C.Massal.) Schifn. ................................................................................322 Cololejeunea rotundilobula (P.C.Wu et P.J.Lin) Piippo ...............................................................319 Cololejeunea runssorensis (Steph.) Pócs .........................................................................................322 Cololejeunea salgadoi Onr. ...........................................................................................................330 Cololejeunea saltuum Tixier .........................................................................................................336 Cololejeunea sambiroana Tixier....................................................................................................336 Cololejeunea sanctae-helenae M.Wigginton ..................................................................................330 Cololejeunea saroltae Pócs ............................................................................................................336 Cololejeunea schaeferi Grolle ........................................................................................................323 Cololejeunea schmidtii Steph. .......................................................................................................323 Cololejeunea schmidtii var. acutepapillosa Pócs .............................................................................323 Cololejeunea schusteri Pócs ...........................................................................................................336 Cololejeunea schwabei Herzog ......................................................................................................336 Cololejeunea selaginellicola Tixier .................................................................................................323 Cololejeunea selangorensis Tixier...................................................................................................336 Cololejeunea serrata (Steph.) Benedix ..........................................................................................323 Cololejeunea serrulata Steph.........................................................................................................330 Cololejeunea setiloba A.Evans .......................................................................................................331 Cololejeunea setosa Mizut. ............................................................................................................339 Cololejeunea sharpii Mizut. ..........................................................................................................339 Cololejeunea shibiensis Mizut. ......................................................................................................336 Cololejeunea shikokiana (Horik.) S.Hatt. .....................................................................................323 Cololejeunea shimizui N.Kitag. ....................................................................................................320 Cololejeunea shimizui var. phangngana N.Kitag. ..........................................................................320 Cololejeunea siamensis Steph. .......................................................................................................339 Cololejeunea siangensis G.Asthana et S.C.Srivast. .........................................................................331 World checklist of hornworts and liverworts *** *** *** ** *** *** *** *** ** ** * *** ** ** *** *** ** *** ** *** *** ** *** *** *** *** ** ** ** ** ** *** ** ** ** *** ** *** *** ** * ** ** * ** *** * ** *** ** ** ** 545 Cololejeunea sicifolia (Gottsche ex A.Evans) Pócs et Bernecker ....................................................319 Cololejeunea sicifolia subsp. jamaicensis (R.M.Schust.) Bernecker et Pócs.....................................318 Cololejeunea sigmoidea Jovet-Ast et Tixier....................................................................................319 Cololejeunea sigmoidea var. dubia Tixier ......................................................................................319 Cololejeunea sintenisii (Steph.) Pócs .............................................................................................319 Cololejeunea skottsbergii Herzog ...................................................................................................331 Cololejeunea smitinandii Tixier ....................................................................................................336 Cololejeunea societatis Tixier ........................................................................................................331 Cololejeunea sophiana Tixier ........................................................................................................319 Cololejeunea spathulata Jovet-Ast .................................................................................................325 Cololejeunea spathulifolia (Steph.) H.A.Mill. ...............................................................................331 Cololejeunea sphaerodonta Mizut. ................................................................................................339 Cololejeunea spinosa (Horik.) Pandé et R.N.Misra .......................................................................323 Cololejeunea spruceana Tixier ......................................................................................................331 Cololejeunea standleyi Herzog ......................................................................................................319 Cololejeunea stellaris Pócs ............................................................................................................323 Cololejeunea stenophylla Herzog ...................................................................................................331 Cololejeunea stephanii Schifn. ex Benedix ...................................................................................339 Cololejeunea stoniana Tixier ........................................................................................................336 Cololejeunea stotleriana Gradst., Ilk.-Borg. et Vanderp. ...............................................................320 Cololejeunea streimannii Pócs ......................................................................................................331 Cololejeunea streimannii subsp. solomonensis Pócs ........................................................................331 Cololejeunea stylilobula Tixier ......................................................................................................320 Cololejeunea stylosa Steph. ...........................................................................................................336 Cololejeunea subalpina Pócs .........................................................................................................331 Cololejeunea subcardiocarpa Tixier ...............................................................................................336 Cololejeunea subcristata A.Evans ..................................................................................................331 Cololejeunea subloccosa Mizut. ....................................................................................................339 Cololejeunea subinlata Tixier ......................................................................................................336 Cololejeunea subkodamae Mizut. ..................................................................................................323 Cololejeunea sublatistyla Jian Wang bis et R.L.Zhu ......................................................................340 Cololejeunea submarginata Tixier.................................................................................................336 Cololejeunea subminutilobula Mizut. ...........................................................................................336 Cololejeunea subocelloides Mizut. .................................................................................................339 Cololejeunea subscariosa (Spruce) Pócs .........................................................................................336 Cololejeunea subsphaeroidea (R.M.Schust.) Pócs ..........................................................................319 Cololejeunea subtriapiculata Tixier ...............................................................................................336 Cololejeunea succinea Tixier .........................................................................................................336 Cololejeunea surinamensis Tixier ..................................................................................................336 Cololejeunea tahitensis Tixier .......................................................................................................336 Cololejeunea takamakae Tixier .....................................................................................................331 Cololejeunea tamasii Schäf.-Verw. ...............................................................................................331 Cololejeunea tamatavensis Tixier ..................................................................................................336 Cololejeunea tamdaoensis Tixier ...................................................................................................340 Cololejeunea tanneri Pócs ............................................................................................................323 Cololejeunea tanzaniae Pócs.........................................................................................................331 Cololejeunea taprobanea Tixier ....................................................................................................337 Cololejeunea taurifolia Inoue et H.A.Mill. ...................................................................................325 Cololejeunea tenella Benedix ........................................................................................................323 Cololejeunea tenella var. dentiloba Onr.........................................................................................323 Cololejeunea tenuiparietata Tixier ................................................................................................331 Cololejeunea teurnoumensis Tixier ................................................................................................331 546 * ** ** *** ** ** *** ** * *** * ** * * * ** ** *** ** *** *** *** ** ** ** ** *** *** *** *** ** *** *** ** *** *** ** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Cololejeunea thailandensis Tixier ..................................................................................................337 Cololejeunea thiersiae (Pócs) Pócs ................................................................................................319 Cololejeunea thiersiana Tixier ......................................................................................................323 Cololejeunea timoi Pócs ...............................................................................................................331 Cololejeunea tixieri Onr. ..............................................................................................................340 Cololejeunea touwii Pócs..............................................................................................................331 Cololejeunea tranninhiana Tixier .................................................................................................331 Cololejeunea triapiculata (Herzog) Tixier .....................................................................................337 Cololejeunea tribracteata Tixier ....................................................................................................331 Cololejeunea trichomanis (Gottsche) Besch. .................................................................................331 Cololejeunea tridentata Tixier ......................................................................................................337 Cololejeunea tuiwawana Pócs ......................................................................................................323 Cololejeunea tuksapiana Tixier .....................................................................................................331 Cololejeunea uchimae Amakawa ...................................................................................................337 Cololejeunea variifolia (Mitt.) Steph. ...........................................................................................340 Cololejeunea veillonii Tixier .........................................................................................................319 Cololejeunea verdoornii (S.Hatt.) S.Hatt. .....................................................................................340 Cololejeunea verrucosa Steph. .......................................................................................................323 Cololejeunea verrucosa var. rectispina (Herzog) Benedix ...............................................................323 Cololejeunea verwimpii Tixier ......................................................................................................337 Cololejeunea vesicaria (Sande Lac.) Schifn. .................................................................................324 Cololejeunea vidaliana Tixier .......................................................................................................337 Cololejeunea vietnamensis Tixier ..................................................................................................337 Cololejeunea virotana Tixier ........................................................................................................319 Cololejeunea vitaliana Tixier........................................................................................................337 Cololejeunea vulcania Tixier ........................................................................................................331 Cololejeunea wightii Steph. ..........................................................................................................325 Cololejeunea winkleri (M.I.Morales et A.Lücking) Pócs ...............................................................319 Cololejeunea yakusimensis (S.Hatt.) Mizut. ..................................................................................337 Cololejeunea yelitzae Pócs et Bernecker ........................................................................................325 Cololejeunea yipii R.L.Zhu ..........................................................................................................340 Cololejeunea yoshinagana (S.Hatt.) Mizut. ...................................................................................331 Cololejeunea zangii R.L.Zhu et M.L.So .......................................................................................320 Cololejeunea zantenorum Pócs .....................................................................................................340 Cololejeunea zenkeri (Steph.) E.W.Jones ......................................................................................323 Colura acroloba (Prantl) Jovet-Ast ...............................................................................................344 Colura amboinensis Steph. ...........................................................................................................344 Colura andoi Gradst. et Jovet-Ast ................................................................................................341 Colura ari (Steph.) Steph. ...........................................................................................................342 Colura australiensis Jovet-Ast .......................................................................................................342 Colura berghenii Jovet-Ast ...........................................................................................................340 Colura bicornis Jovet-Ast .............................................................................................................343 Colura bisvoluta Herzog et Jovet-Ast ...........................................................................................343 Colura brevistyla Herzog .............................................................................................................342 Colura calderae Pócs ....................................................................................................................342 Colura calyptrifolia (Hook.) Dumort. ..........................................................................................341 Colura clementis Grolle................................................................................................................342 Colura conica (Sande Lac.) K.I.Goebel ........................................................................................342 Colura corniantha Grolle .............................................................................................................344 Colura corynophora (Nees, Lindenb. et Gottsche) Trevis. ............................................................342 Colura crenulata Grolle ...............................................................................................................342 Colura crispiloba Jovet-Ast ..........................................................................................................342 World checklist of hornworts and liverworts *** ** *** ** *** ** *** *** *** *** *** *** *** *** *** *** *** *** ** *** *** *** *** *** *** ** *** ** ** *** ** *** ** *** *** ** *** *** *** *** *** *** *** *** ** *** *** *** *** *** *** *** 547 Colura cristata Jovet-Ast ..............................................................................................................341 Colura cylindrica Herzog .............................................................................................................342 Colura cymbalifera Herzog et Jovet-Ast .......................................................................................342 Colura denticulata Jovet-Ast ........................................................................................................344 Colura digitalis (Mitt.) Steph. .....................................................................................................342 Colura digitalis var. mucronata Pócs ............................................................................................342 Colura dusenii Steph. ..................................................................................................................342 Colura fastigiata Jovet-Ast ...........................................................................................................342 Colura istulosa Jovet-Ast .............................................................................................................342 Colura galeata Jovet-Ast ..............................................................................................................344 Colura greig-smithii Jovet-Ast ......................................................................................................341 Colura hattoriana Pócs ................................................................................................................342 Colura hedbergiana Pócs..............................................................................................................341 Colura heimii Jovet-Ast ...............................................................................................................342 Colura hemisphaerica Jovet-Ast ....................................................................................................342 Colura herzogii Jovet-Ast .............................................................................................................342 Colura hirta Steph.......................................................................................................................344 Colura humbertii Jovet-Ast ..........................................................................................................341 Colura imperfecta Steph...............................................................................................................344 Colura inlata K.I.Goebel ............................................................................................................341 Colura inornata Jovet-Ast ............................................................................................................344 Colura inuii Horik. .....................................................................................................................342 Colura irrorata (Spruce) Heinrichs, Y.Yu, Schäf.-Verw. et Pócs ..................................................341 Colura itatyana Steph..................................................................................................................343 Colura jovet-astiae Grolle ............................................................................................................341 Colura junghuhniana (Prantl) Steph. ...........................................................................................341 Colura karstenii K.I.Goebel .........................................................................................................344 Colura koponenii Pócs .................................................................................................................342 Colura mauritiana Pócs ...............................................................................................................342 Colura maxima Jovet-Ast ............................................................................................................342 Colura medusa J.Eggers et Pócs ...................................................................................................341 Colura meijeri Jovet-Ast ..............................................................................................................341 Colura mizutanii Pócs .................................................................................................................341 Colura mosenii Steph...................................................................................................................342 Colura naumannii (Schifn.) Steph..............................................................................................343 Colura norrisii Pócs .....................................................................................................................342 Colura obesa Jovet-Ast .................................................................................................................343 Colura obvoluta Jovet-Ast ............................................................................................................341 Colura ornata K.I.Goebel ............................................................................................................341 Colura ornithocephala Herzog......................................................................................................343 Colura palawanensis Jovet-Ast .....................................................................................................341 Colura pallida Steph. ..................................................................................................................343 Colura pluridentata Jovet-Ast ......................................................................................................343 Colura pulcherrima Jovet-Ast.......................................................................................................343 Colura pulcherrima var. bartlettii Jovet-Ast ..................................................................................343 Colura queenslandica B.M.hiers.................................................................................................344 Colura rhynchophora Jovet-Ast ....................................................................................................341 Colura saccophylla E.A.Hodgs. et Herzog ....................................................................................343 Colura saroltae Pócs.....................................................................................................................344 Colura schusteri Grolle.................................................................................................................343 Colura siamensis Jovet-Ast ...........................................................................................................344 Colura speciosa Jovet-Ast .............................................................................................................343 548 ** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** ** *** *** *** *** *** *** * *** *** *** ** *** ** *** *** *** *** ** ** *** *** *** * * *** ** ** * *** *** *** *** ** * Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Colura streimannii Pócs ...............................................................................................................343 Colura strophiolata Jovet-Ast .......................................................................................................344 Colura superba (Mont.) Steph. ....................................................................................................343 Colura tenuicornis (A.Evans) Steph..............................................................................................341 Colura thomeensis Pócs ................................................................................................................343 Colura tixieri Jovet-Ast................................................................................................................344 Colura tortifolia (Nees et Mont.) Trevis. .....................................................................................343 Colura tutuilana (Pearson) H.A.Mill. ..........................................................................................343 Colura ulei Jovet-Ast ...................................................................................................................343 Colura usambarica E.W.Jones .....................................................................................................344 Colura valida Jovet-Ast ...............................................................................................................341 Colura verdoornii Herzog et Jovet-Ast .........................................................................................341 Colura vietnamensis Jovet-Ast et Tixier ........................................................................................344 Colura vitiensis Pócs et J.Eggers ...................................................................................................343 Conocephalum conicum (L.) Dumort. ..........................................................................................485 Conocephalum japonicum (hunb.) Grolle ...................................................................................486 Conocephalum salebrosum Szweyk., Buczk. et Odrzyk..................................................................485 Conoscyphus koponenii Piippo, Mamontov et Potemkin...............................................................193 Conoscyphus trapezioides (Sande Lac.) Schifn. .............................................................................193 Corsinia coriandrina (Spreng.) Lindb. .........................................................................................486 Cronisia imbriata (Nees) Whittem. et Bischl. .............................................................................486 Cronisia weddellii (Mont.) Grolle ................................................................................................486 Crossocalyx hellerianus (Nees ex Lindenb.) Meyl. ...........................................................................52 Crossocalyx tenuis (Harry Williams) Schljakov ...............................................................................52 Crossotolejeunea curvifolia Steph. .................................................................................................508 Cryptocolea imbricata R.M.Schust. ..............................................................................................123 Cryptocoleopsis imbricata Amakawa..............................................................................................110 Cryptolophocolea aculeata (Mitt.) L.Söderstr. ...............................................................................193 Cryptolophocolea chiloscyphoidea (Lindenb.) L.Söderstr. et Crand.-Stotl. ......................................193 Cryptolophocolea ciliolata (Nees) L.Söderstr., Crand.-Stotl., Stotler et Váňa ................................193 Cryptolophocolea compacta (Mitt.) L.Söderstr...............................................................................193 Cryptolophocolea connata (Sw.) L.Söderstr. et Váňa .....................................................................193 Cryptolophocolea connatifolia (J.J.Engel) L.Söderstr. ....................................................................194 Cryptolophocolea costata (Nees) L.Söderstr. ..................................................................................194 Cryptolophocolea edentata (J.J.Engel) L.Söderstr. .........................................................................194 Cryptolophocolea explanata (Mitt.) Váňa et Crand.-Stotl..............................................................194 Cryptolophocolea leischeri (Steph.) L.Söderstr. .............................................................................194 Cryptolophocolea guadalupensis (Steph.) L.Söderstr. et Váňa ........................................................194 Cryptolophocolea helmsiana (Steph.) L.Söderstr. ...........................................................................194 Cryptolophocolea leucophylla (Hook.f. et Taylor) L.Söderstr. ........................................................194 Cryptolophocolea levieri (Schifn.) L.Söderstr. ..............................................................................194 Cryptolophocolea lilliena (Steph.) L.Söderstr. ...............................................................................194 Cryptolophocolea martiana (Nees) L.Söderstr., Crand.-Stotl. et Stotler.........................................194 Cryptolophocolea martiana subsp. bidentula (Nees) L.Söderstr., Crand.-Stotl. et Stotler ...............194 Cryptolophocolea martiana var. perissodonta (Spruce) Gradst. .......................................................194 Cryptolophocolea massalongoana (Schifn.) L.Söderstr...................................................................195 Cryptolophocolea mitteniana (Colenso) L.Söderstr........................................................................195 Cryptolophocolea mitteniana var. obtusa (J.J.Engel) L.Söderstr. ....................................................195 Cryptolophocolea mitteniana var. symmetrica (J.J.Engel) L.Söderstr. .............................................195 Cryptolophocolea pallida (Mitt.) L.Söderstr. .................................................................................195 Cryptolophocolea pallidovirens (Hook.f. et Taylor) L.Söderstr. .....................................................195 Cryptolophocolea proteus (Herzog) L.Söderstr. ..............................................................................195 World checklist of hornworts and liverworts * *** *** * *** * *** ** *** * *** *** *** *** *** *** *** *** *** *** *** *** *** ** *** *** *** *** *** * *** *** *** *** *** *** * *** * *** *** * *** *** *** *** *** *** *** *** *** ** 549 Cryptolophocolea pycnophylla (Spruce) L.Söderstr. ........................................................................195 Cryptolophocolea regularis (Steph.) L.Söderstr. .............................................................................195 Cryptolophocolea spinifera (Hook.f. et Taylor) L.Söderstr.............................................................195 Cryptolophocolea stephanii (Schifn.) L.Söderstr. ..........................................................................195 Cryptolophocolea subopposita (J.J.Engel) L.Söderstr. .....................................................................195 Cryptolophocolea thermarum (Schifn.) L.Söderstr. .......................................................................195 Cryptolophocolea trialata (Gottsche) L.Söderstr............................................................................196 Cryptolophocolea tricorata (Hässel) Crand.-Stotl. et Stotler ..........................................................196 Cryptolophocolea tuberculata (J.J.Engel) L.Söderstr. .....................................................................196 Cryptolophocolea whittieriana (Inoue et H.A.Mill.) L.Söderstr. ....................................................196 Cryptomitrium himalayense Kashyap ............................................................................................481 Cryptomitrium oreades Perold ......................................................................................................481 Cryptomitrium tenerum (Hook.) Austin ex Underw.....................................................................481 Cuspidatula contracta (Reinw., Blume et Nees) Steph. ..................................................................45 Cuspidatula laccida (Steph.) K.Feldberg, Váňa, Hentschel et Heinrichs .......................................45 Cuspidatula lexicaulis (Nees) Váňa et L.Söderstr...........................................................................45 Cuspidatula kirkii (Steph.) K.Feldberg, Váňa, Hentschel et Heinrichs ..........................................46 Cuspidatula monodon (Taylor) Steph.............................................................................................46 Cuspidatula orbicularis (Grolle) Váňa et L.Söderstr. ......................................................................46 Cuspidatula robusta (Austin) Váňa et L.Söderstr............................................................................46 Cyathodium aureonitens (Grif.) Schifn.......................................................................................486 Cyathodium bischlerianum N.Salazar ...........................................................................................486 Cyathodium cavernarum Kunze ...................................................................................................486 Cyathodium denticulatum Udar et S.C.Srivast. ............................................................................486 Cyathodium foetidissimum Schifn. ..............................................................................................487 Cyathodium indicum Udar et D.K.Singh .....................................................................................487 Cyathodium mehranum D.K.Singh..............................................................................................487 Cyathodium smaragdinum Schifn. ..............................................................................................487 Cyathodium spruceanum Prosk. ...................................................................................................487 Cyathodium spurium (Dicks.) Lindb. ex Braithw. ........................................................................487 Cyathodium steerei Hässel ............................................................................................................487 Cyathodium tuberculatum Udar et D.K.Singh .............................................................................487 Cyathodium tuberosum Kashyap ..................................................................................................487 Cyclolejeunea accedens (Gottsche) A.Evans ...................................................................................351 Cyclolejeunea chitonia (Taylor) A.Evans.......................................................................................351 Cyclolejeunea convexistipa (Lehm. et Lindenb.) A.Evans ..............................................................351 Cyclolejeunea ecuadorensis Steph. .................................................................................................351 Cyclolejeunea foliorum (Nees) Grolle ...........................................................................................351 Cyclolejeunea integerrima (Steph.) Steph. .....................................................................................351 Cyclolejeunea luteola (Spruce) Grolle ...........................................................................................351 Cyclolejeunea peruviana (Lehm. et Lindenb.) A.Evans .................................................................351 Cyclolejeunea spectabilis Steph......................................................................................................351 Cylindrocolea abyssinica (Gola) Váňa .............................................................................................73 Cylindrocolea andersonii R.M.Schust. ............................................................................................72 Cylindrocolea brasiliensis D.P.Costa, N.D.Santos et Váňa..............................................................72 Cylindrocolea chevalieri (Steph.) R.M.Schust. ................................................................................72 Cylindrocolea gittinsii (E.W.Jones) R.M.Schust. ............................................................................73 Cylindrocolea kiaeri (Austin) Váňa.................................................................................................73 Cylindrocolea madagascariensis (Steph.) R.M.Schust. .....................................................................73 Cylindrocolea nigerica (E.W.Jones) R.M.Schust. ............................................................................73 Cylindrocolea novae-caledoniae (Grolle) R.M.Schust. .....................................................................73 Cylindrocolea obtusifolia Fulford ....................................................................................................73 550 *** *** ** *** *** *** *** *** *** *** *** *** ** ** *** * *** ** ** *** ** ** *** *** * ** *** ** *** ** *** * *** * *** ** ** *** *** *** *** ** *** * * ** *** ** *** ** ** *** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Cylindrocolea planifolia (Steph.) R.M.Schust. ................................................................................73 Cylindrocolea recurvifolia (Steph.) Inoue........................................................................................73 Cylindrocolea reticulata Udar et Ad.Kumar ....................................................................................73 Cylindrocolea rhizantha (Mont.) R.M.Schust. ...............................................................................73 Cylindrocolea sanctae-helenae M.Wigginton ...................................................................................73 Cylindrocolea sprucei R.M.Schust. .................................................................................................73 Cylindrocolea tagawae (N.Kitag.) R.M.Schust. ..............................................................................73 Cylindrocolea ugandica (E.W.Jones) R.M.Schust. ..........................................................................73 Cyrtolejeunea holostipa (Spruce) A.Evans .....................................................................................316 Cystolejeunea lineata (Lehm. et Lindenb.) A.Evans ......................................................................316 Dactylophorella muricata (Gottsche) R.M.Schust.........................................................................300 Deceptifrons plagiochiloides J.J.Engel et Váňa ...............................................................................196 Delavayella serrata Steph. ............................................................................................................117 Delavayella serrata var. purpurea P.C.Chen .................................................................................117 Dendroceros acutilobus Steph. ........................................................................................................32 Dendroceros adglutinatus (Hook.f. et Taylor) Gottsche, Lindenb. et Nees .....................................33 Dendroceros africanus Steph. ..........................................................................................................33 Dendroceros allionii Steph. ............................................................................................................33 Dendroceros australis Steph. ...........................................................................................................33 Dendroceros borbonicus Steph. .......................................................................................................32 Dendroceros breutelii Nees .............................................................................................................33 Dendroceros breutelii var. surinamensis Lindenb. et Gottsche .........................................................33 Dendroceros cavernosus J.Haseg. .....................................................................................................32 Dendroceros cichoraceus (Mont.) Gottsche .....................................................................................33 Dendroceros crassicostatus Steph. ....................................................................................................33 Dendroceros crassinervis (Nees) Gottsche........................................................................................33 Dendroceros crispatus (Hook.) Nees ...............................................................................................33 Dendroceros crispatus var. simplicior Spruce ....................................................................................33 Dendroceros crispus (Sw.) Nees.......................................................................................................32 Dendroceros cucullatus Steph..........................................................................................................33 Dendroceros diicilis Steph.............................................................................................................32 Dendroceros exalatus Steph. ...........................................................................................................33 Dendroceros foliicola J.Haseg. ........................................................................................................32 Dendroceros gracilis Steph. .............................................................................................................33 Dendroceros granulatus Mitt. .........................................................................................................33 Dendroceros herasii M.Infante........................................................................................................33 Dendroceros humboldtensis Hürl. ...................................................................................................33 Dendroceros japonicus Steph. .........................................................................................................32 Dendroceros javanicus (Nees) Nees ................................................................................................32 Dendroceros muelleri Steph. ...........................................................................................................32 Dendroceros ogeramnangus Piippo ..................................................................................................32 Dendroceros paivae C.A.Garcia, Sérgio et J.C.Villarreal .................................................................33 Dendroceros pedunculatus Steph. ....................................................................................................32 Dendroceros rarus Steph.................................................................................................................34 Dendroceros reticulus Herzog .........................................................................................................34 Dendroceros rigidus Steph. .............................................................................................................34 Dendroceros seramensis J.Haseg. .....................................................................................................34 Dendroceros subdiicilis S.Hatt. .....................................................................................................32 Dendroceros subplanus Steph..........................................................................................................32 Dendroceros subtropicus C.J.Wild...................................................................................................34 Dendroceros tahitensis Ångstr. ........................................................................................................34 Dendroceros tubercularis S.Hatt. ....................................................................................................32 World checklist of hornworts and liverworts *** ** ** *** *** *** *** *** * *** *** *** *** *** *** *** ** *** *** *** ** *** *** ** ** ** * *** ** ** *** *** ** *** * * ** *** *** * ** * *** ** * ** ** *** ** ** *** *** 551 Dendroceros validus Steph..............................................................................................................33 Dendroceros vesconianus Gottsche ..................................................................................................33 Dendroceros wattsianus Steph. .......................................................................................................33 Dendrolembidium dendroides (Carrington et Pearson) Herzog .....................................................163 Dendrolembidium tenax (Grev.) Herzog ......................................................................................164 Dendromastigophora lagellifera (Hook.) R.M.Schust. ..................................................................217 Denotarisia linguifolia (De Not.) Grolle ........................................................................................46 Dicranolejeunea axillaris (Nees et Mont.) Schifn. .......................................................................299 Dicranolejeunea bovonei Gola ......................................................................................................299 Dinckleria fruticella (Hook.f. et Taylor) J.J.Engel et Heinrichs....................................................219 Dinckleria pleurata (Hook.f. et Taylor) Trevis.............................................................................219 Diplasiolejeunea alata Jovet-Ast ...................................................................................................344 Diplasiolejeunea albifolia (Taylor) E.W.Jones ..............................................................................347 Diplasiolejeunea andringitrae Schäf.-Verw. ..................................................................................346 Diplasiolejeunea armatiloba Steph. ..............................................................................................345 Diplasiolejeunea aulae E.W.Jones ................................................................................................347 Diplasiolejeunea auriculata Tixier ................................................................................................347 Diplasiolejeunea borhidiana Reyes Montoya ................................................................................346 Diplasiolejeunea brunnea Steph. ..................................................................................................346 Diplasiolejeunea buckii Grolle......................................................................................................347 Diplasiolejeunea caribea Tixier .....................................................................................................346 Diplasiolejeunea cavifolia Steph. ..................................................................................................346 Diplasiolejeunea cobrensis Steph. ..................................................................................................346 Diplasiolejeunea cobrensis subsp. antsirananae Pócs ......................................................................346 Diplasiolejeunea cogoensis M.Infante, Heras et Pócs .....................................................................346 Diplasiolejeunea columbica Tixier ................................................................................................347 Diplasiolejeunea comorensis Tixier ................................................................................................347 Diplasiolejeunea cornuta Steph.....................................................................................................346 Diplasiolejeunea cubensis Tixier....................................................................................................346 Diplasiolejeunea cyanguguensis Tixier ...........................................................................................347 Diplasiolejeunea deslooveri Vanden Berghen.................................................................................347 Diplasiolejeunea eggersii Pócs .......................................................................................................345 Diplasiolejeunea ensifera Tixier ....................................................................................................345 Diplasiolejeunea erostrata Schäf.-Verw. ........................................................................................347 Diplasiolejeunea evansii Tixier .....................................................................................................347 Diplasiolejeunea glaziovii Tixier ...................................................................................................346 Diplasiolejeunea gradsteinii Tixier ................................................................................................347 Diplasiolejeunea grandirostrata Schäf.-Verw. ................................................................................347 Diplasiolejeunea grolleana Reyes Montoya ...................................................................................347 Diplasiolejeunea guadalupensis Steph. ..........................................................................................345 Diplasiolejeunea hamata Tixier ....................................................................................................347 Diplasiolejeunea heimii Jovet-Ast .................................................................................................345 Diplasiolejeunea ingekarolae Schäf.-Verw. ....................................................................................346 Diplasiolejeunea insignis Tixier ....................................................................................................347 Diplasiolejeunea integerrima Tixier ..............................................................................................347 Diplasiolejeunea involuta S.Winkl. ..............................................................................................345 Diplasiolejeunea involuta subsp. andicola Pócs .............................................................................345 Diplasiolejeunea johnsonii A.Evans...............................................................................................345 Diplasiolejeunea johnsonii var. mexicana Jovet-Ast .......................................................................345 Diplasiolejeunea jonesii Tixier ......................................................................................................346 Diplasiolejeunea jovet-astiae Grolle ..............................................................................................346 Diplasiolejeunea kraussiana (Lindenb.) Steph. .............................................................................346 552 *** *** *** ** ** ** *** *** * *** *** ** *** *** *** *** ** *** *** *** *** ** *** *** ** ** *** ** *** *** *** *** *** *** *** *** *** *** * * *** ** ** * * *** * *** *** ** ** *** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Diplasiolejeunea lanceolata Grolle ................................................................................................347 Diplasiolejeunea latipuensis Tixier ................................................................................................348 Diplasiolejeunea leiocarpa Jovet-Ast .............................................................................................346 Diplasiolejeunea lemuriana Tixier ................................................................................................348 Diplasiolejeunea longilobula Herzog .............................................................................................348 Diplasiolejeunea magnistipula Tixier ............................................................................................348 Diplasiolejeunea malleiformis (A.Evans) Tixier .............................................................................346 Diplasiolejeunea mayaykuensis Schäf.-Verw. et Heinrichs .............................................................348 Diplasiolejeunea montecristensis S.Winkl. .....................................................................................345 Diplasiolejeunea onraedtii Grolle .................................................................................................348 Diplasiolejeunea ornata Pócs et Schäf.-Verw. ...............................................................................346 Diplasiolejeunea palustrium Tixier ...............................................................................................346 Diplasiolejeunea papilionacea R.M.Schust. ...................................................................................345 Diplasiolejeunea patelligera Herzog ..............................................................................................347 Diplasiolejeunea pauckertii (Nees) Steph. .....................................................................................345 Diplasiolejeunea pellucida (C.F.W.Meissn. ex Spreng.) Schifn. ...................................................346 Diplasiolejeunea phyllarthronii Tixier ...........................................................................................346 Diplasiolejeunea plicatiloba (Hook.f. et Taylor) Grolle ................................................................347 Diplasiolejeunea pluridentata Schäf.-Verw. ..................................................................................345 Diplasiolejeunea pocsii Reyes Montoya.........................................................................................346 Diplasiolejeunea pusilla Grolle .....................................................................................................347 Diplasiolejeunea ramicola Tixier ..................................................................................................348 Diplasiolejeunea ranomafanae Pócs ..............................................................................................347 Diplasiolejeunea replicata (Spruce) Steph. ....................................................................................345 Diplasiolejeunea riclefgrollei Schäf.-Verw. ....................................................................................348 Diplasiolejeunea rudolphiana Steph..............................................................................................345 Diplasiolejeunea runssorensis Steph. ..............................................................................................345 Diplasiolejeunea subcornuta Tixier ...............................................................................................348 Diplasiolejeunea symoensii Vanden Berghen .................................................................................348 Diplasiolejeunea unidentata (Lehm. et Lindenb.) Schifn. ............................................................345 Diplasiolejeunea utriculata Steph. ................................................................................................344 Diplasiolejeunea villaumei Steph. .................................................................................................347 Diplasiolejeunea zakiae Tixier ......................................................................................................347 Diplocolea sikkimensis Amakawa ..................................................................................................123 Diplophyllum africanum S.W.Arnell ..............................................................................................83 Diplophyllum albicans (L.) Dumort. ..............................................................................................83 Diplophyllum andicola R.M.Schust................................................................................................83 Diplophyllum andrewsii A.Evans....................................................................................................83 Diplophyllum androgynum J.J.Engel et G.L.Merr. .........................................................................83 Diplophyllum angustifolium J.J.Engel et G.L.Merr. ........................................................................83 Diplophyllum apiculatum (A.Evans) Steph. ....................................................................................83 Diplophyllum dioicum R.M.Schust. ...............................................................................................83 Diplophyllum exiguum Steph. ........................................................................................................83 Diplophyllum gemmiparum J.J.Engel et G.L.Merr. ........................................................................83 Diplophyllum incrassatum J.J.Engel et G.L.Merr............................................................................84 Diplophyllum nanum Herzog ........................................................................................................84 Diplophyllum novum J.J.Engel et G.L.Merr. ..................................................................................84 Diplophyllum obtusatum (R.M.Schust.) R.M.Schust......................................................................84 Diplophyllum obtusifolium (Hook.) Dumort..................................................................................84 Diplophyllum obtusifolium subsp. domesticum (Gottsche) Váňa......................................................84 Diplophyllum recurvifolium C.Massal. ...........................................................................................83 Diplophyllum serrulatum (Müll.Frib.) Steph. .................................................................................84 World checklist of hornworts and liverworts *** *** ** *** *** *** *** *** ** *** ** *** *** *** *** *** *** ** ** *** ** *** ** *** *** ** ** ** *** ** ** ** ** *** *** *** *** ** *** *** *** ** ** * * * ** ** ** *** *** ** 553 Diplophyllum squarrosum Steph. ....................................................................................................83 Diplophyllum taxifolium (Wahlenb.) Dumort................................................................................84 Diplophyllum taxifolium var. mucronatum R.M.Schust. .................................................................84 Diplophyllum trollii Grolle ............................................................................................................84 Diplophyllum verrucosum R.M.Schust. ..........................................................................................83 Douinia imbricata (M.Howe) Konstant. et Vilnet .........................................................................84 Douinia ovata (Dicks.) H.Buch.....................................................................................................84 Douinia plicata (Lindb.) Konstant. et Vilnet .................................................................................84 Drepanolejeunea actinogyna Inuthai, R.L.Zhu et Chantanaorr. ....................................................359 Drepanolejeunea aculeata Bischl...................................................................................................353 Drepanolejeunea anderssonii (Ångstr.) A.Evans ............................................................................353 Drepanolejeunea andina Herzog ..................................................................................................353 Drepanolejeunea angustifolia (Mitt.) Grolle..................................................................................353 Drepanolejeunea ankasica E.W.Jones ...........................................................................................353 Drepanolejeunea anoplantha (Spruce) Steph. ...............................................................................354 Drepanolejeunea appalachiana R.M.Schust. .................................................................................354 Drepanolejeunea araucariae Steph................................................................................................354 Drepanolejeunea araucariae var. chilensis Herzog .........................................................................354 Drepanolejeunea aucklandica Steph. ............................................................................................354 Drepanolejeunea aurita Bischl......................................................................................................354 Drepanolejeunea bakeri Herzog ...................................................................................................354 Drepanolejeunea bidens (Prantl) A.Evans .....................................................................................354 Drepanolejeunea bidoupensis Pócs ................................................................................................360 Drepanolejeunea biocellata A.Evans..............................................................................................354 Drepanolejeunea bischlerae (Grolle) Grolle et R.L.Zhu.................................................................360 Drepanolejeunea blumei Steph. ....................................................................................................354 Drepanolejeunea brunnea Mizut. .................................................................................................354 Drepanolejeunea caledonica Steph. ...............................................................................................354 Drepanolejeunea campanulata (Spruce) Steph. .............................................................................354 Drepanolejeunea canceroides H.A.Mill. et Bonner ........................................................................354 Drepanolejeunea capulata (Taylor) J.B.Jack et Steph. ...................................................................354 Drepanolejeunea capulata var. lagellifera S.W.Arnell ...................................................................354 Drepanolejeunea ciliata Mizut. ....................................................................................................354 Drepanolejeunea commutata Grolle et R.L.Zhu ...........................................................................360 Drepanolejeunea crassiretis A.Evans ..............................................................................................354 Drepanolejeunea crucianella (Taylor) A.Evans ..............................................................................354 Drepanolejeunea cultrella (Mitt.) Steph........................................................................................354 Drepanolejeunea cutervoensis (Loitl.) Grolle .................................................................................354 Drepanolejeunea cyclops (Sande Lac.) Grolle et R.L.Zhu ..............................................................360 Drepanolejeunea dactylophora (Nees, Lindenb. et Gottsche) J.B.Jack et Steph..............................360 Drepanolejeunea dactylophora var. submuricata Herzog ................................................................360 Drepanolejeunea decurviloba Steph. .............................................................................................355 Drepanolejeunea dentistipula Steph. .............................................................................................353 Drepanolejeunea deslooveri Vanden Berghen ................................................................................355 Drepanolejeunea devendrae Sushil K.Singh et M.Dey ..................................................................360 Drepanolejeunea dissitifolia A.Evans.............................................................................................355 Drepanolejeunea elegans Herzog ...................................................................................................355 Drepanolejeunea erecta (Steph.) Mizut. ........................................................................................355 Drepanolejeunea evansii Bischl. ex L.Söderstr., A.Hagborg et von Konrat ....................................355 Drepanolejeunea issicornua Steph. ...............................................................................................359 Drepanolejeunea leischeri (Steph.) Grolle et R.L.Zhu ..................................................................360 Drepanolejeunea foliicola Horik. ..................................................................................................360 554 *** ** *** ** *** ** *** * ** *** ** *** ** ** *** * *** *** ** ** * *** *** * ** ** *** ** *** ** ** ** *** * * ** ** *** *** *** ** *** ** ** *** *** ** ** ** ** ** ** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Drepanolejeunea fragilis Bischl. ex L.Söderstr., A.Hagborg et von Konrat ....................................355 Drepanolejeunea fulfordiae L.Söderstr. .........................................................................................355 Drepanolejeunea granatensis (Prantl) Bischl..................................................................................355 Drepanolejeunea grandis Herzog ..................................................................................................355 Drepanolejeunea grollei M.E.Reiner et Schäf.-Verw. ....................................................................355 Drepanolejeunea grossidens Steph. ................................................................................................355 Drepanolejeunea hamatifolia (Hook.) Schifn. .............................................................................355 Drepanolejeunea hampeana Steph. ...............................................................................................359 Drepanolejeunea hamulata Steph. ................................................................................................355 Drepanolejeunea helenae Pócs ......................................................................................................355 Drepanolejeunea herzogii R.L.Zhu et M.L.So ..............................................................................355 Drepanolejeunea inchoata (C.F.W.Meissn.) Steph. ......................................................................356 Drepanolejeunea inchoata var. palmicola Pócs ..............................................................................356 Drepanolejeunea inchoata var. roraimae (Steph. ex Zwickel) Bischl. .............................................356 Drepanolejeunea infundibulata (Spruce) A.Evans .........................................................................356 Drepanolejeunea integerrima Herzog ............................................................................................360 Drepanolejeunea integribracteata Bischl........................................................................................356 Drepanolejeunea intermedia Zwickel ............................................................................................359 Drepanolejeunea laciniata Qiong He et R.L.Zhu .........................................................................359 Drepanolejeunea laevis (Mitt.) Steph............................................................................................356 Drepanolejeunea lancifolia (Gottsche) J.B.Jack et Steph. ..............................................................356 Drepanolejeunea levicornua Steph. ...............................................................................................356 Drepanolejeunea lichenicola (Spruce) Steph..................................................................................356 Drepanolejeunea longicornua (Herzog) Mizut. .............................................................................359 Drepanolejeunea longicruris (Steph.) Grolle et R.L.Zhu ...............................................................360 Drepanolejeunea longii Grolle et R.L.Zhu ....................................................................................356 Drepanolejeunea lyrata Grolle ......................................................................................................359 Drepanolejeunea macrodonta (Mitt.) Steph. .................................................................................356 Drepanolejeunea madagascariensis (Steph.) Grolle ........................................................................359 Drepanolejeunea mawtmiana Ajit P.Singh et V.Nath ...................................................................356 Drepanolejeunea microcarpa Pearson............................................................................................356 Drepanolejeunea moluccensis Herzog ............................................................................................356 Drepanolejeunea mosenii (Steph.) Bischl. .....................................................................................356 Drepanolejeunea nymanii Steph. ..................................................................................................359 Drepanolejeunea obliqua Steph. ...................................................................................................356 Drepanolejeunea obtriangulata T.Kodama ...................................................................................356 Drepanolejeunea obtusifolia T.Yamag. ..........................................................................................357 Drepanolejeunea orthophylla (Nees et Mont.) Bischl. ...................................................................357 Drepanolejeunea palmifolia (Nees) Schifn. ..................................................................................357 Drepanolejeunea pentadactyla (Mont.) Steph. ..............................................................................357 Drepanolejeunea perissodonta (Spruce) Bischl. ..............................................................................357 Drepanolejeunea physifolia (Gottsche) Pearson .............................................................................357 Drepanolejeunea pinnatiloba Schifn. ...........................................................................................357 Drepanolejeunea pleiodictya Herzog .............................................................................................357 Drepanolejeunea pocsii Grolle ......................................................................................................359 Drepanolejeunea polyrhiza (Nees) Grolle et R.L.Zhu ...................................................................360 Drepanolejeunea propagulifera Herzog .........................................................................................357 Drepanolejeunea pseudoneura (A.Evans) Grolle ............................................................................357 Drepanolejeunea pterocalyx (Herzog) Bischl. ................................................................................357 Drepanolejeunea pulla (Mitt.) Grolle ...........................................................................................360 Drepanolejeunea pungens Bischl. ..................................................................................................357 Drepanolejeunea ramentilora Steph. ............................................................................................357 World checklist of hornworts and liverworts * *** * *** ** *** ** ** ** * ** ** ** ** ** * ** * *** ** *** ** *** *** *** ** *** ** ** ** ** ** ** ** *** ** *** *** *** * * *** *** ** ** ** *** *** *** * * * 555 Drepanolejeunea ruandensis Vanden Berghen...............................................................................357 Drepanolejeunea senticosa Bischl. .................................................................................................357 Drepanolejeunea serricalyx Herzog ...............................................................................................360 Drepanolejeunea siamensis (Bischl.) Grolle et R.L.Zhu.................................................................360 Drepanolejeunea sikkimensis (Udar et U.S.Awasthi) Grolle ..........................................................357 Drepanolejeunea spicata (Steph.) Grolle et R.L.Zhu.....................................................................360 Drepanolejeunea spinistipula Herzog ............................................................................................353 Drepanolejeunea spinosa Herzog ..................................................................................................358 Drepanolejeunea spinosocornuta Steph. .........................................................................................358 Drepanolejeunea subdissitifolia Herzog .........................................................................................358 Drepanolejeunea submuricata R.M.Schust. ..................................................................................358 Drepanolejeunea subquadrata (Mitt.) Steph. ................................................................................358 Drepanolejeunea subvittata (Herzog) Grolle .................................................................................358 Drepanolejeunea symoensii Vanden Berghen et Grolle ..................................................................359 Drepanolejeunea symoensii var. minor Tixier ................................................................................359 Drepanolejeunea tenax K.I.Goebel ...............................................................................................358 Drepanolejeunea tenera K.I.Goebel ..............................................................................................358 Drepanolejeunea tenera var. litoceras Herzog ................................................................................358 Drepanolejeunea ternatensis (Gottsche) Schifn. ...........................................................................358 Drepanolejeunea teysmannii (Gottsche) Steph. .............................................................................358 Drepanolejeunea thwaitesiana (Mitt.) Steph. ................................................................................360 Drepanolejeunea thwaitesiana var. zhengii R.L.Zhu......................................................................360 Drepanolejeunea tibetana (P.C.Wu et J.S.Lou) Grolle et R.L.Zhu ...............................................360 Drepanolejeunea trematodes (Nees) Bischl. ...................................................................................359 Drepanolejeunea tricornua Herzog ...............................................................................................360 Drepanolejeunea tridactyla (Gottsche) Steph. ...............................................................................358 Drepanolejeunea trigonophylla Steph. ...........................................................................................358 Drepanolejeunea tristaniana S.W.Arnell.......................................................................................358 Drepanolejeunea tuyamae S.Hatt. ................................................................................................358 Drepanolejeunea ualanensis Inoue et H.A.Mill. ............................................................................358 Drepanolejeunea ungulata (Steph.) Grolle ....................................................................................358 Drepanolejeunea urceolata R.M.Schust. .......................................................................................359 Drepanolejeunea valiae Jovet-Ast .................................................................................................359 Drepanolejeunea vandenberghenii Buchb. et Eb.Fisch. .................................................................359 Drepanolejeunea vesiculosa (Mitt.) Steph......................................................................................359 Drepanolejeunea yulensis Steph. ...................................................................................................359 Drepanolejeunea yunnanensis (P.C.Chen) Grolle et R.L.Zhu .......................................................360 Drucella integristipula (Steph.) E.A.Hodgs. .................................................................................163 Dumortiera hirsuta (Sw.) Nees ....................................................................................................487 Dumortiera hirsuta subsp. nepalensis (Taylor) R.M.Schust...........................................................487 Dumortiera hirsuta subsp. tatunoi Horik. ....................................................................................487 Echinolejeunea papillata (Mitt.) R.M.Schust. ex Hamlin .............................................................362 Endogemma caespiticia (Lindenb.) Konstant., Vilnet et A.V.Troitsky ..........................................109 Enigmella thallina G.A.M.Scott et K.G.Beckm. ............................................................................93 Eocalypogeia quelpaertensis (S.Hatt. et Inoue) R.M.Schust. ..........................................................107 Eocalypogeia schusterana (S.Hatt. et Mizut.) R.M.Schust. ............................................................107 Eotrichocolea furukii T.Katag.......................................................................................................256 Eotrichocolea polyacantha (Hook.f. et Taylor) R.M.Schust...........................................................256 Eremonotus myriocarpus (Carrington) Lindb. et Kaal. ex Pearson.................................................118 Eulejeunea setulosa Steph. ............................................................................................................508 Eulejeunea subpililoba Steph. .......................................................................................................508 Euosmolejeunea parvistipula (Lindenb. et Gottsche) Steph. ..........................................................508 556 * *** *** *** *** * *** *** *** *** * * * * *** ** ** *** ** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** ** ** *** *** *** *** *** *** *** *** * *** *** *** *** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Euosmolejeunea tenerrima (Nees) Steph. ......................................................................................508 Evansianthus georgiensis (Gottsche) R.M.Schust. et J.J.Engel .......................................................196 Exormotheca bischlerae Furuki et Higuchi ....................................................................................488 Exormotheca bulbigena Bornefeld, O.H.Volk et R.Wolf ..............................................................488 Exormotheca ceylonensis Meijer ....................................................................................................488 Exormotheca gollanii Steph. .........................................................................................................488 Exormotheca holstii Steph. ...........................................................................................................488 Exormotheca pustulosa Mitt. ........................................................................................................488 Exormotheca tuberifera Kashyap ..................................................................................................488 Exormotheca welwitschii Steph. ....................................................................................................488 Fimbraria gigantea Steph. ...........................................................................................................508 Fimbraria incrassata Steph. .........................................................................................................508 Fimbraria kamerunensis Steph. ....................................................................................................508 Fimbraria pirottae Gola ...............................................................................................................508 Folioceros amboinensis (Schifn.) Piippo .........................................................................................31 Folioceros apiahynus (Steph.) Hässel ..............................................................................................31 Folioceros argillaceus (Steph.) J.C.Villarreal et Cargill ....................................................................31 Folioceros assamicus D.C.Bharadwaj ..............................................................................................31 Folioceros dilatatus (Steph.) J.C.Villarreal et Cargill .......................................................................31 Folioceros dixitianus (Mahab.) D.C.Bharadwaj ..............................................................................31 Folioceros fuciformis (Mont.) D.C.Bharadwaj ................................................................................31 Folioceros glandulosus (Lehm. et Lindenb.) D.C.Bharadwaj ...........................................................31 Folioceros incurvus (Steph.) D.C.Bharadwaj...................................................................................31 Folioceros indicus D.C.Bharadwaj ..................................................................................................31 Folioceros kashyapii S.C.Srivast. et A.K.Asthana.............................................................................31 Folioceros mangaloreus (Steph.) D.C.Bharadwaj .............................................................................31 Folioceros paliformis D.K.Singh .....................................................................................................31 Folioceros physocladus D.C.Bharadwaj............................................................................................31 Folioceros pinnilobus (Steph.) D.C.Bharadwaj................................................................................31 Folioceros satpurensis D.C.Bharadwaj et K.P.Srivast. ......................................................................31 Folioceros udarii A.K.Asthana et S.C.Srivast. .................................................................................32 Folioceros verruculosus (J.Haseg.) R.L.Zhu et M.J.Lai ....................................................................32 Fossombronia alaskana Steere et Inoue .........................................................................................463 Fossombronia alata G.A.M.Scott et D.C.Pike ..............................................................................463 Fossombronia altilamellosa G.A.M.Scott et D.C.Pike ...................................................................463 Fossombronia angulifolia Perold ...................................................................................................463 Fossombronia angulosa (Dicks.) Raddi .........................................................................................463 Fossombronia areolata G.A.M.Scott et D.C.Pike .........................................................................464 Fossombronia auricolor G.A.M.Scott et D.C.Pike ........................................................................464 Fossombronia australis Mitt. ........................................................................................................464 Fossombronia caespitiformis (Raddi) De Not. ex Rabenh. .............................................................464 Fossombronia caespitiformis subsp. multispira (Schifn.) J.R.Bray et Cargill ..................................464 Fossombronia caledonica Steph. ....................................................................................................464 Fossombronia cederbergensis Perold...............................................................................................464 Fossombronia cerebriformis G.A.M.Scott et D.C.Pike ..................................................................464 Fossombronia crassifolia Spruce ....................................................................................................464 Fossombronia crispa Nees .............................................................................................................464 Fossombronia crispula (Brot.) R.M.Schust....................................................................................464 Fossombronia cristula Austin ........................................................................................................464 Fossombronia cultriformis G.A.M.Scott et D.C.Pike ....................................................................464 Fossombronia densa G.A.M.Scott et D.C.Pike .............................................................................464 Fossombronia densilamellata S.W.Arnell ......................................................................................464 World checklist of hornworts and liverworts *** *** *** *** *** *** ** *** *** ** * * *** ** *** *** *** *** * * *** *** ** *** *** *** *** ** *** *** * ** *** ** *** ** *** *** *** *** * ** *** *** *** *** ** *** *** *** *** *** 557 Fossombronia echinata Macvicar ..................................................................................................464 Fossombronia elsieae Perold ..........................................................................................................464 Fossombronia fernandeziensis Steph. .............................................................................................464 Fossombronia imbriata Paton ......................................................................................................464 Fossombronia leischeri Osterwald ex Loeske ................................................................................464 Fossombronia foveolata Lindb. .....................................................................................................464 Fossombronia fuhreri G.A.M.Scott et D.C.Pike ...........................................................................464 Fossombronia gemmifera Perold ...................................................................................................465 Fossombronia glenii Perold...........................................................................................................465 Fossombronia grandis Steph. ........................................................................................................465 Fossombronia gregaria Colenso ....................................................................................................465 Fossombronia grossepapillata Steph. ..............................................................................................465 Fossombronia hamatohirta Steph..................................................................................................465 Fossombronia hewsoniae G.A.M.Scott et D.C.Pike ......................................................................465 Fossombronia himalayensis Kashyap .............................................................................................465 Fossombronia hyalorhiza Perold ...................................................................................................465 Fossombronia incurva Lindb. .......................................................................................................465 Fossombronia indica Steph. ..........................................................................................................465 Fossombronia integerrima Steph. ..................................................................................................465 Fossombronia integrifolia Steph. ...................................................................................................465 Fossombronia intestinalis Taylor ...................................................................................................465 Fossombronia japonica Schifn. ....................................................................................................465 Fossombronia laciniata G.A.M.Scott et D.C.Pike ........................................................................465 Fossombronia lamellata Steph. .....................................................................................................465 Fossombronia leucoxantha (Lehm.) Lehm. et Lindenb. .................................................................465 Fossombronia longiseta (Austin) Austin ........................................................................................465 Fossombronia lophoclada Spruce...................................................................................................465 Fossombronia lophoscypha Hässel..................................................................................................465 Fossombronia luetzelburgiana K.I.Goebel .....................................................................................465 Fossombronia macrocalyx Steph. ...................................................................................................465 Fossombronia macrophylla Colenso ..............................................................................................466 Fossombronia magnaspora G.A.M.Scott et D.C.Pike ....................................................................466 Fossombronia marindae Perold.....................................................................................................466 Fossombronia maritima (Paton) Paton .........................................................................................466 Fossombronia marshii J.R.Bray et Stotler .....................................................................................466 Fossombronia microlamellata G.A.M.Scott et D.C.Pike ...............................................................466 Fossombronia mittenii Tind. ........................................................................................................466 Fossombronia montaguensis S.W.Arnell ........................................................................................466 Fossombronia monticola Perold ....................................................................................................466 Fossombronia mylioides Inoue ......................................................................................................466 Fossombronia nigricaulis Colenso .................................................................................................466 Fossombronia nyikaensis Perold ....................................................................................................466 Fossombronia papillata Steph. ......................................................................................................466 Fossombronia paranapanemae Schifn. .........................................................................................466 Fossombronia peruviana Gottsche et Hampe................................................................................466 Fossombronia porphyrorhiza (Nees) Prosk. ...................................................................................466 Fossombronia pulvinata Steph. .....................................................................................................466 Fossombronia punctata G.A.M.Scott et D.C.Pike ........................................................................466 Fossombronia purpureospora G.A.M.Scott et D.C.Pike ................................................................466 Fossombronia pusilla (L.) Nees .....................................................................................................466 Fossombronia renateae Perold.......................................................................................................466 Fossombronia reticulata Steph. .....................................................................................................466 558 * ** *** ** *** *** *** * *** *** * *** ** *** *** *** ** ** *** *** *** ** *** *** * ** *** ** * * ** ** * ** ** *** *** ** ** ** *** ** ** * ** ** ** ** ** *** *** ** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Fossombronia rosulata Colenso.....................................................................................................467 Fossombronia rudis G.A.M.Scott et D.C.Pike ..............................................................................467 Fossombronia ruminata Cargill ....................................................................................................467 Fossombronia rupestris G.A.M.Scott et D.C.Pike .........................................................................467 Fossombronia rwandaensis Perold .................................................................................................467 Fossombronia scrobiculata G.A.M.Scott et D.C.Pike ....................................................................467 Fossombronia spinifolia Steph. .....................................................................................................467 Fossombronia spinosa Perold ........................................................................................................467 Fossombronia stephanii Schifn. ex Steph. ....................................................................................467 Fossombronia straussiana Perold ...................................................................................................467 Fossombronia subsaccata Steph. ....................................................................................................467 Fossombronia swaziensis Perold ....................................................................................................467 Fossombronia tesselata G.A.M.Scott et D.C.Pike..........................................................................467 Fossombronia texana Lindb. .........................................................................................................467 Fossombronia truncata G.A.M.Scott et D.C.Pike .........................................................................467 Fossombronia tumida Mitt. ..........................................................................................................467 Fossombronia valparaisiana Hässel ...............................................................................................467 Fossombronia vermiculata G.A.M.Scott et D.C.Pike ....................................................................467 Fossombronia wattsii Steph. .........................................................................................................467 Fossombronia wondraczekii (Corda) Dumort. ex Lindb. ..............................................................467 Fossombronia wrightii Austin .......................................................................................................467 Fossombronia zuurbergensis Perold ...............................................................................................467 Frullania acicularis Hentschel et von Konrat ...............................................................................292 Frullania aculeata Taylor ............................................................................................................287 Frullania ainis Nees et Mont.....................................................................................................293 Frullania akiyamae S.Hatt...........................................................................................................267 Frullania albertii Steph. ..............................................................................................................266 Frullania allanii E.A.Hodgs. .......................................................................................................281 Frullania allionii Steph. ..............................................................................................................293 Frullania alpina Steph.................................................................................................................293 Frullania alstonii Verd. ...............................................................................................................292 Frullania alternans Nees ..............................................................................................................293 Frullania amamiensis Kamim. .....................................................................................................272 Frullania ambronnii Steph. .........................................................................................................287 Frullania amplicrania Steph. .......................................................................................................272 Frullania ampullifera J.B.Jack et Steph. .......................................................................................272 Frullania anderssonii Ångstr. .......................................................................................................272 Frullania angulata Mitt. ..............................................................................................................289 Frullania angulata var. laciniata Vanden Berghen .......................................................................289 Frullania angustistipa Steph.........................................................................................................272 Frullania anomala E.A.Hodgs. ....................................................................................................282 Frullania antaresensis S.Hatt........................................................................................................266 Frullania aoshimensis Horik. .......................................................................................................292 Frullania apertilobula Gerola.......................................................................................................293 Frullania apicalis Mitt. ................................................................................................................289 Frullania apicalis var. camerunensis Vanden Berghen ...................................................................289 Frullania apiculata (Reinw., Blume et Nees) Nees.......................................................................267 Frullania apiculata var. goebelii Schifn........................................................................................267 Frullania aposinensis S.Hatt. et P.J.Lin ........................................................................................273 Frullania appalachiana R.M.Schust.............................................................................................284 Frullania appendiculata Steph. ....................................................................................................292 Frullania appendistipula S.Hatt. ..................................................................................................273 World checklist of hornworts and liverworts ** ** * *** ** *** *** * ** *** *** ** *** * *** *** *** *** * ** ** *** *** ** *** ** *** *** *** ** *** * *** * *** ** * * ** * * *** * *** *** *** * *** *** ** ** * 559 Frullania appendistipula var. spinifera S.Hatt. ..............................................................................273 Frullania armata Herzog et L.Clark ............................................................................................293 Frullania armatifolia Verd...........................................................................................................267 Frullania armitiana Steph. ..........................................................................................................267 Frullania armitiana var. inlexula S.Hatt. ....................................................................................268 Frullania arsenii Steph. ...............................................................................................................266 Frullania asagrayana Mont..........................................................................................................292 Frullania astrolabea Steph. ..........................................................................................................285 Frullania aterrima (Hook.f. et Taylor) Gottsche, Lindenb. et Nees .............................................268 Frullania atrata (Sw.) Nees ex Mont. ..........................................................................................287 Frullania attenuata Steph. ...........................................................................................................268 Frullania auriculata S.Hatt. ........................................................................................................273 Frullania azorica Sim-Sim, Sérgio, Mues et Kraut .......................................................................284 Frullania baileyana Steph. ...........................................................................................................282 Frullania baladina Gottsche ........................................................................................................282 Frullania baumannii S.Hatt. .......................................................................................................271 Frullania beauverdii Steph. .........................................................................................................265 Frullania bella Steph. ..................................................................................................................268 Frullania belmorensis Steph. ........................................................................................................282 Frullania benjaminiana Inoue .....................................................................................................273 Frullania bergmanii S.Hatt..........................................................................................................273 Frullania berthoumieui Steph. .....................................................................................................273 Frullania beyrichiana (Lehm. et Lindenb.) Lehm. et Lindenb. ....................................................287 Frullania bhutanensis S.Hatt. ......................................................................................................273 Frullania bicornistipula Spruce ....................................................................................................287 Frullania blastopetala S.Hatt. ......................................................................................................273 Frullania blepharozia Spruce .......................................................................................................265 Frullania bogotensis Steph............................................................................................................265 Frullania bolanderi Austin ...........................................................................................................294 Frullania bonariensis M.E.Reiner ................................................................................................266 Frullania bonincola S.Hatt. .........................................................................................................273 Frullania borbonica Lindenb. ......................................................................................................289 Frullania boveana C.Massal.........................................................................................................294 Frullania brachycarpa Spruce.......................................................................................................265 Frullania brasiliensis Raddi ..........................................................................................................287 Frullania brasiliensis var. elegantula Spruce ..................................................................................287 Frullania breuteliana Gottsche ....................................................................................................287 Frullania brevicalycina Steph. ......................................................................................................273 Frullania brittoniae A.Evans ........................................................................................................273 Frullania brotheri Steph. .............................................................................................................286 Frullania brunea (Spreng.) Drège ................................................................................................272 Frullania bullata Steph................................................................................................................273 Frullania caespitans Beauverd ......................................................................................................294 Frullania cafraria Steph. ............................................................................................................273 Frullania calcarata Ångstr. ..........................................................................................................273 Frullania calcarifera Steph. ..........................................................................................................292 Frullania caldensis Ångstr. ...........................................................................................................287 Frullania californica (M.Howe) A.Evans .....................................................................................292 Frullania campanulata Sande Lac. ...............................................................................................282 Frullania campanulata var. caduca Verd. .....................................................................................282 Frullania campanulata var. malesiaca (Verd.) S.Hatt. ..................................................................282 Frullania canaliculata Gottsche ...................................................................................................294 560 *** ** ** *** * *** * ** ** *** * * ** * *** ** ** *** * *** *** *** * * *** *** ** * *** * * *** ** * * *** * * * ** *** ** *** ** * *** ** *** ** ** ** * Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Frullania capensis Gottsche .........................................................................................................289 Frullania capillaris Steph. ............................................................................................................294 Frullania carrii Kamim. et S.Hatt. ..............................................................................................273 Frullania catalinae A.Evans .........................................................................................................284 Frullania cataractarum Steph. .....................................................................................................282 Frullania caulisequa (Nees) Mont................................................................................................272 Frullania cavallii Gola.................................................................................................................294 Frullania changii S.Hatt. et C.Gao ..............................................................................................268 Frullania chenii S.Hatt. et P.J.Lin ...............................................................................................273 Frullania chevalieri (R.M.Schust.) R.M.Schust. ..........................................................................291 Frullania chiapasana Steph. .........................................................................................................294 Frullania chilcootiensis Steph. ......................................................................................................294 Frullania chilensis Steph. .............................................................................................................273 Frullania chiovendae Gola ...........................................................................................................294 Frullania chodatii Beauverd.........................................................................................................273 Frullania ciliata Lindenb. et Gottsche .........................................................................................294 Frullania cinchonae Gottsche ......................................................................................................294 Frullania clavata (Hook.f. et Taylor) Gottsche, Lindenb. et Nees ...............................................281 Frullania claviloba Steph. ............................................................................................................268 Frullania clemensiana Verd. ........................................................................................................292 Frullania cobrensis Gottsche ........................................................................................................273 Frullania colliculosa von Konrat, Braggins, Hentschel et Heinrichs .............................................268 Frullania compacta Gottsche .......................................................................................................287 Frullania complicata Steph. .........................................................................................................294 Frullania confertiloba Steph. ........................................................................................................265 Frullania congesta Gottsche, Lindenb. et Nees.............................................................................271 Frullania consociata Steph. ..........................................................................................................273 Frullania contracta Steph. ...........................................................................................................273 Frullania convoluta Lindenb. et Hampe ......................................................................................289 Frullania convoluta var. ampliata Herzog ....................................................................................289 Frullania cordaeana Lindenb. ......................................................................................................294 Frullania cordistipula (Reinw., Blume et Nees) Nees ...................................................................268 Frullania cordistipula var. dentistipula S.Hatt. .............................................................................268 Frullania cordistipula var. mutica (Gottsche, Lindenb. et Nees) Schifn. ......................................268 Frullania cornuta Steph. ..............................................................................................................273 Frullania crassitexta Steph. ..........................................................................................................273 Frullania crawfordii Steph. ..........................................................................................................282 Frullania crenatiloba Steph..........................................................................................................268 Frullania crenulifolia J.B.Jack et Steph. .......................................................................................287 Frullania crispiloba Steph. ...........................................................................................................288 Frullania crispiplicata Yuzawa et S.Hatt. .....................................................................................273 Frullania cristata S.Hatt. .............................................................................................................274 Frullania cuencensis Taylor ..........................................................................................................266 Frullania cuneatistipula Steph. ....................................................................................................294 Frullania cuneiloba Nees .............................................................................................................274 Frullania curvilobula Schäf.-Verw., D.F.Peralta et S.M.Siqueira .................................................267 Frullania curviramea Steph. ........................................................................................................288 Frullania curvistipula Steph.........................................................................................................274 Frullania curvistipula var. falcatidentata S.Hatt. ..........................................................................274 Frullania curvistipula var. lamii Verd. .........................................................................................274 Frullania cuspidifolia Steph. ........................................................................................................274 Frullania cuspiloba Steph. ...........................................................................................................294 World checklist of hornworts and liverworts ** *** *** ** *** *** * *** *** ** ** *** *** ** *** *** ** ** * *** ** *** ** *** *** ** ** ** *** ** ** *** * * * *** * *** ** * ** *** * ** * ** *** ** * ** * * 561 Frullania cyparioides (Schwägr.) Nees ..........................................................................................274 Frullania darwinii Gradst. et Uribe.............................................................................................289 Frullania davurica Hampe ex Gottsche, Lindenb. et Nees...........................................................274 Frullania debilis Steph. ex S.Hatt. ...............................................................................................274 Frullania decidua Spruce .............................................................................................................265 Frullania delexa Mitt..................................................................................................................285 Frullania degelii S.W.Arnell ........................................................................................................268 Frullania densiloba Steph. ex A.Evans .........................................................................................292 Frullania dentata S.Hatt..............................................................................................................282 Frullania dentata var. secernens S.Hatt. ........................................................................................282 Frullania dentifera S.Hatt. et Streimann......................................................................................268 Frullania dentiloba S.Hatt. ..........................................................................................................268 Frullania deplanata Mitt. ............................................................................................................285 Frullania deppii Lehm. ................................................................................................................274 Frullania depressa Mitt. ...............................................................................................................266 Frullania dilatata (L.) Dumort. ...................................................................................................284 Frullania dilatata subsp. asiatica S.Hatt. .....................................................................................284 Frullania diptera (Lehm.) Drège .................................................................................................274 Frullania dispar Nees ..................................................................................................................294 Frullania diversitexta Steph. ........................................................................................................272 Frullania donnellii Austin............................................................................................................289 Frullania dulimensis Uribe ..........................................................................................................289 Frullania durifolia Steph. ............................................................................................................268 Frullania dusenii Steph. ..............................................................................................................266 Frullania duthiana Steph. ...........................................................................................................274 Frullania duthiana var. appendiculata S.Hatt...............................................................................274 Frullania duthiana var. laevis S.Hatt. ..........................................................................................274 Frullania duthiana var. szechuanensis S.Hatt. et C.Gao................................................................274 Frullania eboracensis Lehm. .........................................................................................................284 Frullania echinantha S.Hatt. .......................................................................................................274 Frullania echinatella S.Hatt. ........................................................................................................274 Frullania ecklonii (Spreng.) Gottsche, Lindenb. et Nees ..............................................................265 Frullania ecklonii var. robustior (Gottsche, Lindenb. et Nees) Sim...............................................265 Frullania ecklonii var. rufescens Gottsche, Lindenb. et Nees .........................................................265 Frullania ecklonii var. tenerior (Gottsche, Lindenb. et Nees) Sim ................................................265 Frullania ecuadorensis Steph. .......................................................................................................288 Frullania elegans Lehm. ...............................................................................................................274 Frullania elephantum S.Hatt. ......................................................................................................274 Frullania epiphylla S.Hatt. ..........................................................................................................274 Frullania epiphylla subsp. ijiensis S.Hatt. ....................................................................................274 Frullania eplicata Steph. ..............................................................................................................289 Frullania ericoides (Nees) Mont...................................................................................................284 Frullania ericoides var. laxa (Gottsche, Lindenb. et Nees) Schifn. ...............................................284 Frullania ericoides var. verrucosa (Kamim.) Hentschel et von Konrat ...........................................284 Frullania ericoides var. minor Kamim. .........................................................................................274 Frullania erostrata S.Hatt. ...........................................................................................................274 Frullania errans Verd. .................................................................................................................282 Frullania errans var. angulistipula S.Hatt. ....................................................................................283 Frullania esenbeckiana Beauverd..................................................................................................274 Frullania evelynae S.Hatt. et haithong .......................................................................................274 Frullania evelynae var. devendrae Sushil K.Singh et Barbhuiya.....................................................274 Frullania evelynae var. srivastavae Sushil K.Singh et Barbhuiya....................................................274 562 ** ** *** ** *** * ** ** ** ** *** * * ** *** *** ** *** ** ** ** ** ** *** ** ** *** *** ** ** ** *** ** *** * ** ** *** * *** ** *** *** ** *** * * * *** ** ** ** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Frullania evoluta Mitt. ................................................................................................................287 Frullania exilis Taylor .................................................................................................................268 Frullania eymae S.Hatt. ...............................................................................................................274 Frullania eymae var. crispidentata S.Hatt. et Streimann ...............................................................274 Frullania falciloba Lehm. ............................................................................................................274 Frullania fallax Gottsche .............................................................................................................285 Frullania falsicornuta S.Hatt. ......................................................................................................274 Frullania fauriana Steph. ............................................................................................................274 Frullania fengyangshanensis R.L.Zhu et M.L.So ...........................................................................275 Frullania ferdinandi-muelleri Steph. ............................................................................................275 Frullania fertilis De Not. .............................................................................................................290 Frullania lammea Taylor ............................................................................................................264 Frullania lexuosa S.Hatt. ............................................................................................................275 Frullania formosa Spruce .............................................................................................................288 Frullania fragilifolia (Taylor) Gottsche, Lindenb. et Nees ...........................................................284 Frullania franciscana M.Howe ....................................................................................................292 Frullania fuegiana Steph. ............................................................................................................284 Frullania fugax (Hook.f. et Taylor) Gottsche, Lindenb. et Nees..................................................283 Frullania fulfordiae S.Hatt. .........................................................................................................283 Frullania fuscovirens Steph. .........................................................................................................275 Frullania fuscovirens var. gemmipara (R.M.Schust. et S.Hatt.) S.Hatt. et P.J.Lin .........................275 Frullania gabonensis Vanden Berghen .........................................................................................268 Frullania gaoligongensis X.L.Bai et C.Gao....................................................................................275 Frullania gaudichaudii (Nees et Mont.) Nees et Mont. ...............................................................291 Frullania gaudichaudii var. ceylanica (Nees) S.Hatt. ....................................................................292 Frullania gemmulosa S.Hatt. et haithong...................................................................................275 Frullania gibbosa Nees ................................................................................................................265 Frullania gigantea Steph. .............................................................................................................275 Frullania giraldiana C.Massal. ....................................................................................................275 Frullania giraldiana var. handelii (Verd.) S.Hatt. ........................................................................275 Frullania globosa S.Hatt. et Streimann ........................................................................................275 Frullania glomerata (Lehm. et Lindenb.) Nees et Mont...............................................................283 Frullania gracilicaulis S.Hatt. ......................................................................................................267 Frullania gracilis (Reinw., Blume et Nees) Nees ..........................................................................268 Frullania gracilis var. brevior (Gottsche, Lindenb. et Nees) Schifn..............................................269 Frullania gracilis var. vittata S.Hatt. ............................................................................................269 Frullania gracilis subsp. zennoskei S.Hatt. et haithong ...............................................................269 Frullania gradsteinii Yuzawa, Mues et S.Hatt. .............................................................................266 Frullania granatensis Gottsche .....................................................................................................288 Frullania grandifolia Steph. .........................................................................................................289 Frullania grandilobula S.Hatt. et Piippo......................................................................................275 Frullania grandistipula Lindenb. .................................................................................................275 Frullania griithsiana Gottsche ...................................................................................................288 Frullania grossiclava Steph. ..........................................................................................................271 Frullania grossifolia Steph. ...........................................................................................................272 Frullania guadalupensis Gottsche.................................................................................................288 Frullania gualaquizana Steph. .....................................................................................................288 Frullania guatemalensis Steph. .....................................................................................................294 Frullania haematocysta Spruce .....................................................................................................266 Frullania hainanensis S.Hatt. et P.J.Lin .......................................................................................275 Frullania hamata Steph. ..............................................................................................................286 Frullania hamaticoma Steph. .......................................................................................................281 World checklist of hornworts and liverworts ** ** ** ** *** ** ** ** ** *** * *** *** ** ** ** * *** *** ** *** ** * ** * *** ** ** ** ** ** * * *** ** *** ** *** ** ** ** *** ** ** *** ** *** ** * *** ** ** 563 Frullania hamatiloba Steph. ........................................................................................................275 Frullania hamilora Herzog et L.Clark ........................................................................................288 Frullania handelii Verd. ..............................................................................................................275 Frullania handel-mazzettii S.Hatt................................................................................................275 Frullania hasskarliana Lindenb. ..................................................................................................269 Frullania hasskarliana var. gracilis S.Hatt. ...................................................................................269 Frullania hasskarliana var. parvidentata S.Hatt. ...........................................................................269 Frullania hattoriana J.D.Godfrey et G.Godfrey ..........................................................................284 Frullania hattoriantha Udar et V.Nath ........................................................................................275 Frullania hattorii von Konrat et Braggins ....................................................................................269 Frullania hebridensis Steph. .........................................................................................................275 Frullania hedrantha S.Hatt. et Kamim. .......................................................................................292 Frullania heteromorpha Schifn. ...................................................................................................286 Frullania hicksiae S.Hatt. ............................................................................................................275 Frullania higuchii Yuzawa, Koike et S.Hatt. ................................................................................275 Frullania hiroshii S.Hatt. ............................................................................................................275 Frullania hirtilora Spruce ...........................................................................................................275 Frullania hodgsoniae von Konrat, Braggins, Hentschel et Heinrichs ............................................269 Frullania holostipula S.Hatt. et D.G.Griin ................................................................................265 Frullania hottana S.Hatt. ............................................................................................................269 Frullania howeana Steph. ............................................................................................................276 Frullania huerlimannii S.Hatt. ....................................................................................................286 Frullania huerlimannii var. dioica S.Hatt. ...................................................................................286 Frullania humbertii Vanden Berghen ..........................................................................................267 Frullania humilis Spruce .............................................................................................................276 Frullania hypoleuca Nees .............................................................................................................272 Frullania hypoleucula S.Hatt. ......................................................................................................267 Frullania ignatovii Sofronova, Mamontov et Potemkin ...............................................................276 Frullania imerinensis Steph..........................................................................................................289 Frullania immersa Steph. ............................................................................................................292 Frullania incisoduthiana S.Hatt. ..................................................................................................276 Frullania incisoduthiana var. parva S.Hatt. ..................................................................................276 Frullania incisostipula Steph. .......................................................................................................276 Frullania inconstans Verd. ...........................................................................................................269 Frullania inconstans var. grossedentata Kamim. et S.Hatt. ............................................................269 Frullania incumbens Mitt. ...........................................................................................................283 Frullania incurva S.Hatt. ............................................................................................................267 Frullania inlata Gottsche ...........................................................................................................294 Frullania inlata var. communis R.M.Schust. ...............................................................................294 Frullania inlata var. dioica S.Hatt. et haithong ........................................................................294 Frullania inlata var. mayebarae (S.Hatt.) K.Yamada ...................................................................294 Frullania inlexa Mitt. .................................................................................................................283 Frullania inlexiloba S.Hatt. ........................................................................................................276 Frullania inouei S.Hatt. ..............................................................................................................276 Frullania integristipula (Nees) Nees.............................................................................................285 Frullania integristipula var. emarginata Verd. ..............................................................................285 Frullania intermedia (Reinw., Blume et Nees) Nees ....................................................................286 Frullania intermedia subsp. morokensis (Steph.) S.Hatt................................................................286 Frullania intermedia var. non-apiculata S.Hatt. ...........................................................................286 Frullania intumescens (Lehm. et Lindenb.) Lehm. et Lindenb. ....................................................288 Frullania involvens S.Hatt. et Kamim..........................................................................................286 Frullania iriomotensis S.Hatt. ......................................................................................................286 564 *** ** *** ** * *** *** ** *** ** ** ** ** ** ** ** ** *** ** *** ** * ** * * * * * ** *** * * ** ** ** * *** * *** ** *** *** ** * * ** * * ** ** ** *** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Frullania irregularis S.Hatt. et Piippo..........................................................................................276 Frullania iwatsukii S.Hatt. ..........................................................................................................293 Frullania jackii Gottsche .............................................................................................................276 Frullania jacobsii S.Hatt. .............................................................................................................276 Frullania jacquinotii Gottsche .....................................................................................................276 Frullania jelskii Loitl. ..................................................................................................................266 Frullania johnsonii Steph.............................................................................................................269 Frullania jovetiana von Konrat et Hentschel ...............................................................................276 Frullania junghuhniana Gottsche ................................................................................................291 Frullania junghuhniana var. bisexualis S.Hatt. .............................................................................291 Frullania junghuhniana var. tenella (Sande Lac.) Grolle et S.Hatt. ..............................................291 Frullania kagoshimensis Steph. .....................................................................................................276 Frullania kagoshimensis subsp. hunanensis (S.Hatt.) S.Hatt. et P.J.Lin .........................................276 Frullania kalimantanensis S.Hatt. ................................................................................................276 Frullania kashyapii Verd. ............................................................................................................276 Frullania kitagawana S.Hatt. ......................................................................................................276 Frullania klotzschii Nees..............................................................................................................267 Frullania knightbridgei von Konrat et de Lange ...........................................................................290 Frullania koponenii S.Hatt. .........................................................................................................284 Frullania kunzei (Lehm. et Lindenb.) Lehm. et Lindenb.............................................................289 Frullania kunzei var. maritima R.M.Schust. ................................................................................290 Frullania laetevirens Hampe ex Gottsche, Lindenb. et Nees ........................................................294 Frullania laeviperiantha X.L.Bai et C.Gao ...................................................................................276 Frullania lancistyla Steph. ...........................................................................................................276 Frullania larjiana Sushil K.Singh et D.K.Singh ..........................................................................294 Frullania larjiana var. didyhatii S.N.Srivast. et M.Rai .................................................................294 Frullania laticaulis Spruce ...........................................................................................................288 Frullania latilora Spruce.............................................................................................................276 Frullania latogaleata Herzog........................................................................................................276 Frullania laxilora Spruce ............................................................................................................266 Frullania laxilora var. crossii Spruce ............................................................................................276 Frullania leana Austin .................................................................................................................294 Frullania leeuwenii Verd. ............................................................................................................286 Frullania lepida S.Hatt. et Piippo................................................................................................276 Frullania letestui Vanden Berghen ...............................................................................................267 Frullania levieri Steph. ................................................................................................................276 Frullania lindenbergii Lehm. .......................................................................................................272 Frullania lindenbergii var. fusca Gottsche, Lindenb. et Nees ........................................................272 Frullania lindmanii Steph. ..........................................................................................................264 Frullania linii S.Hatt. .................................................................................................................276 Frullania lobatohastata Steph. .....................................................................................................288 Frullania lobulata (Hook.) Hook. et Nees ...................................................................................290 Frullania longistipula Steph. ........................................................................................................290 Frullania longistipula var. apiculata Demaret et Vanden Berghen ................................................288 Frullania longistyla Yuzawa et S.Hatt. .........................................................................................276 Frullania loricata Pearson ............................................................................................................272 Frullania loricata var. laxa Pearson ..............................................................................................272 Frullania ludoviciae Steph. ..........................................................................................................277 Frullania lushanensis S.Hatt. et P.J.Lin........................................................................................277 Frullania macgregorii Steph. ........................................................................................................269 Frullania macgregorii var. rostellula (S.Hatt.) S.Hatt. ...................................................................269 Frullania macrocephala (Lehm. et Lindenb.) Lehm. et Lindenb...................................................288 World checklist of hornworts and liverworts ** * * ** ** *** ** ** * * ** ** ** *** ** *** ** *** ** ** ** *** * *** ** *** *** * ** ** *** *** ** ** ** *** *** ** *** * ** *** ** * ** ** ** ** *** ** ** ** 565 Frullania macrophylla S.Hatt. ......................................................................................................285 Frullania macularis Taylor ..........................................................................................................277 Frullania madagascariensis Gottsche ............................................................................................294 Frullania madens Steph. ..............................................................................................................269 Frullania madothecoides Spruce ...................................................................................................288 Frullania magellanica F.Weber et Nees .......................................................................................290 Frullania magellanica subsp. tristaniana (S.W.Arnell) Váňa et J.J.Engel ......................................290 Frullania mammillosa S.Hatt.......................................................................................................286 Frullania matafaoica H.A.Mill. ...................................................................................................290 Frullania mauritiana Austin ........................................................................................................294 Frullania maymyoensis Svihla .......................................................................................................277 Frullania mcveanii S.Hatt. ..........................................................................................................291 Frullania media (E.A.Hodgs.) S.Hatt. .........................................................................................283 Frullania megalostipa Spruce .......................................................................................................265 Frullania mehrana S.Hatt. ..........................................................................................................269 Frullania meijeri S.Hatt. .............................................................................................................287 Frullania meridana Steph. ...........................................................................................................288 Frullania meyeniana Lindenb. .....................................................................................................290 Frullania meyeniana var. dioica S.Hatt. .......................................................................................290 Frullania microauriculata Verd....................................................................................................277 Frullania microauriculata var. rotundior Verd. .............................................................................277 Frullania microcaulis Gola ...........................................................................................................291 Frullania microcephala Gottsche..................................................................................................288 Frullania microphylla (Gottsche) Pearson ....................................................................................293 Frullania microrhyncha L.Clark et Svihla .....................................................................................277 Frullania microscopica Pearson ....................................................................................................291 Frullania mirabilis J.B.Jack et Steph............................................................................................294 Frullania miradorensis Lindenb. et Gottsche ...............................................................................267 Frullania mizoramensis Sushil K.Singh et Barbhuiya ...................................................................277 Frullania mizutanii Kamim. et S.Hatt. .......................................................................................283 Frullania moniliata (Reinw., Blume et Nees) Mont. ...................................................................293 Frullania monocera (Hook.f. et Taylor) Gottsche, Lindenb. et Nees............................................281 Frullania monocera var. acutiloba (Mitt.) Hentschel et von Konrat ..............................................282 Frullania monocera var. depauperata S.Hatt. ................................................................................282 Frullania monocera var. schifneri (Verd.) S.Hatt. ........................................................................282 Frullania monocera var. subhampeana (E.A.Hodgs.) Hentschel et von Konrat .............................282 Frullania monocera var. undulata (Kamim.) Hentschel et von Konrat .........................................282 Frullania monoica Steph..............................................................................................................295 Frullania montagnei Gottsche .....................................................................................................288 Frullania montana Steph. ............................................................................................................277 Frullania moritziana Lindenb. et Gottsche..................................................................................288 Frullania morobensis S.Hatt. et Streimann ...................................................................................285 Frullania motoyana Steph. ...........................................................................................................269 Frullania multilacera Steph. ........................................................................................................269 Frullania multilacera subsp. gracilior S.Hatt. ...............................................................................269 Frullania multilacera var. lacerissima S.Hatt. ...............................................................................269 Frullania multilaceroides S.Hatt...................................................................................................269 Frullania multituberculata Hentschel et von Konrat ....................................................................277 Frullania muscicola Steph. ...........................................................................................................284 Frullania mutilata Steph. ............................................................................................................277 Frullania nadeaudii Steph. ..........................................................................................................277 Frullania neocaledonica J.J.Engel .................................................................................................291 566 ** *** *** *** ** * *** ** *** ** *** ** ** ** ** *** ** ** *** *** *** ** * ** * ** ** ** *** ** *** * ** ** ** ** ** ** * ** ** ** ** ** *** ** *** ** * *** *** *** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Frullania neosheana S.Hatt. .........................................................................................................269 Frullania nepalensis (Spreng.) Lehm. et Lindenb. ........................................................................277 Frullania neurota Taylor .............................................................................................................266 Frullania nicholsonii E.A.Hodgs. .................................................................................................277 Frullania nigricaulis (Reinw., Blume et Nees) Nees .....................................................................277 Frullania nigricaulis var. elongata Verd. .......................................................................................277 Frullania nisquallensis Sull...........................................................................................................293 Frullania nivimontana S.Hatt. ....................................................................................................277 Frullania nobilis Steph. ...............................................................................................................277 Frullania nobilis var. cochleata (Steph.) S.Hatt. ...........................................................................277 Frullania nodulosa (Reinw., Blume et Nees) Nees .......................................................................286 Frullania notarisii Steph. .............................................................................................................287 Frullania novocurvirostris S.Hatt. ................................................................................................277 Frullania novoguineensis Schifn. .................................................................................................286 Frullania oahuensis Hampe ex Gottsche, Lindenb. et Nees ..........................................................278 Frullania oakesiana Austin ..........................................................................................................284 Frullania oakesiana subsp. takayuensis (Steph.) R.M.Schust.........................................................284 Frullania obovata S.Hatt. ............................................................................................................278 Frullania obscura (Sw.) Mont. .....................................................................................................265 Frullania obscura var. spiniloba (Steph.) Hentschel et von Konrat ...............................................265 Frullania obscurifolia Mitt. ..........................................................................................................283 Frullania obtusangula Hentschel et von Konrat ...........................................................................278 Frullania ocanniensis Steph..........................................................................................................295 Frullania ocellata S.Hatt. et Kamim. ...........................................................................................271 Frullania odontostipa Spruce........................................................................................................267 Frullania okinawensis Kamim. .....................................................................................................278 Frullania onraedtii Vanden Berghen ...........................................................................................290 Frullania orbicularis Austin .........................................................................................................278 Frullania orientalis Sande Lac......................................................................................................278 Frullania orinocensis Spruce.........................................................................................................278 Frullania ornithocephala (Reinw., Blume et Nees) Nees ..............................................................278 Frullania ornithocephala var. major (Nees) Schifn.......................................................................278 Frullania ornithocephala var. pilosa Verd. ....................................................................................278 Frullania ornithocephala var. tuberculosa S.Hatt. ..........................................................................278 Frullania osculatiana De Not. .....................................................................................................288 Frullania osumiensis (S.Hatt.) S.Hatt...........................................................................................282 Frullania pachyderma S.Hatt. ......................................................................................................278 Frullania pallidevirens Steph. ......................................................................................................278 Frullania pallidula S.Hatt. ..........................................................................................................282 Frullania pancheri Gottsche ........................................................................................................292 Frullania papillata Steph. ............................................................................................................269 Frullania papillilobula S.Hatt. .....................................................................................................292 Frullania papuana Verd. .............................................................................................................290 Frullania papulosa Steph. ............................................................................................................287 Frullania paradoxa Lehm. et Lindenb. ........................................................................................288 Frullania paranensis Steph. ..........................................................................................................264 Frullania parhamii (R.M.Schust.) R.M.Schust. ex von Konrat, L.Söderstr. et A.Hagborg ...........291 Frullania pariharii S.Hatt. et haithong .....................................................................................278 Frullania parvifolia Steph. ...........................................................................................................278 Frullania parvistipula Steph. .......................................................................................................285 Frullania patagonica Steph. .........................................................................................................283 Frullania patula Mitt. .................................................................................................................285 World checklist of hornworts and liverworts ** ** ** ** ** ** *** *** *** *** *** ** ** ** *** *** *** *** * *** ** ** ** *** ** ** ** ** ** ** * ** ** ** ** ** *** ** ** ** ** *** ** *** * ** ** *** ** *** ** ** 567 Frullania pauciramea Steph. ........................................................................................................278 Frullania pauciramea var. pauciramella S.Hatt. et Piippo ............................................................278 Frullania paucirameoides S.Hatt. et Piippo ..................................................................................278 Frullania pearceana Steph. ..........................................................................................................288 Frullania pedicellata Steph. .........................................................................................................278 Frullania pentapleura Taylor .......................................................................................................278 Frullania peruviana Gottsche ......................................................................................................289 Frullania phalangilora Steph. .....................................................................................................289 Frullania physantha Mitt. ............................................................................................................278 Frullania pilibracteola S.Hatt. .....................................................................................................267 Frullania pilistipula Steph. ..........................................................................................................278 Frullania piptophylla S.Hatt. .......................................................................................................278 Frullania piptophylla var. minor S.Hatt........................................................................................278 Frullania piptophylloides S.Hatt. ..................................................................................................278 Frullania pittieri Steph. ...............................................................................................................288 Frullania plana Sull.....................................................................................................................285 Frullania planifolia Steph. ...........................................................................................................266 Frullania platycalyx Herzog .........................................................................................................295 Frullania plicata Hentschel et von Konrat ...................................................................................278 Frullania pluricarinata Gottsche .................................................................................................266 Frullania pocsantha haithong et S.Hatt. ....................................................................................279 Frullania polyptera Taylor ...........................................................................................................283 Frullania polyptera var. angustata (Mitt.) S.Hatt. .........................................................................283 Frullania polysticta Lindenb. .......................................................................................................293 Frullania ponapensis S.Hatt. et Koike ..........................................................................................272 Frullania pran-nathii M.Dey et D.K.Singh .................................................................................279 Frullania pringlei Fulford et Sharp ..............................................................................................279 Frullania probosciphora Taylor ....................................................................................................283 Frullania prominula S.Hatt. et Streimann ...................................................................................279 Frullania propaginea S.Hatt. et Streimann...................................................................................279 Frullania pseudericoides S.Hatt. ...................................................................................................279 Frullania pseudoalstonii Tsudo et J.Haseg. ...................................................................................293 Frullania pseudomeyeniana S.Hatt. ..............................................................................................291 Frullania pseudomonocera S.Hatt. ................................................................................................282 Frullania pseudoschensiana S.Hatt. ..............................................................................................279 Frullania pseudoschensiana var. darjeelingensis S.Hatt. ..................................................................279 Frullania ptychantha Mont. .........................................................................................................271 Frullania pulchella Herzog ..........................................................................................................283 Frullania pullei Verd. ..................................................................................................................279 Frullania pulogensis Steph............................................................................................................269 Frullania punctata Reimers .........................................................................................................293 Frullania purpurea Steph.............................................................................................................269 Frullania pusilla Mitt. .................................................................................................................279 Frullania pycnantha (Hook.f. et Taylor) Gottsche, Lindenb. et Nees ..........................................279 Frullania pyricalycina Steph. .......................................................................................................279 Frullania queenslandica Steph. ....................................................................................................279 Frullania quillotensis (Nees et Mont.) Nees et Mont. ..................................................................295 Frullania ramuligera (Nees) Mont...............................................................................................267 Frullania recurvistipula S.Hatt.....................................................................................................279 Frullania relexistipula Sande Lac. ...............................................................................................279 Frullania relexistipula var. squarrosa S.Hatt. et Piippo ................................................................279 Frullania regularis Schifn. ..........................................................................................................286 568 ** ** ** ** *** ** ** * *** ** ** *** ** ** ** *** *** *** ** *** *** ** ** ** ** ** ** * ** ** ** ** ** ** *** ** *** ** *** ** ** ** ** ** * ** *** ** ** ** ** ** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Frullania reicheana Steph. ...........................................................................................................295 Frullania reimersii Verd. .............................................................................................................270 Frullania remotidens S.Hatt. ........................................................................................................279 Frullania remotiloba Steph. .........................................................................................................286 Frullania repandistipula Sande Lac. .............................................................................................271 Frullania repandistipula subsp. queenslandica S.Hatt. ..................................................................271 Frullania repandistipula subsp. spinibractea S.Hatt. .....................................................................271 Frullania reptans Mitt. ................................................................................................................279 Frullania retusa Mitt. ..................................................................................................................279 Frullania retusa var. gymnantha S.Hatt. et haithong ..................................................................279 Frullania retusa var. hirsuta S.Hatt. et haithong ........................................................................279 Frullania rhystocolea Herzog ........................................................................................................279 Frullania rhytidantha S.Hatt. ......................................................................................................279 Frullania rigescens Spruce ............................................................................................................288 Frullania rigida Steph. ................................................................................................................279 Frullania ringens Spruce ..............................................................................................................265 Frullania rio-janeirensis (Raddi) Ångstr. ......................................................................................265 Frullania riparia Hampe .............................................................................................................279 Frullania rizalii Piippo et S.Hatt. ...............................................................................................280 Frullania rostellata Mitt...............................................................................................................280 Frullania rostrata (Hook.f. et Taylor) Gottsche, Lindenb. et Nees ..............................................291 Frullania rubella Gottsche...........................................................................................................280 Frullania rubella var. elongata (Steph.) S.Hatt. ............................................................................280 Frullania rudoliana S.Hatt. ........................................................................................................287 Frullania rupicola Steph. .............................................................................................................280 Frullania sabahana S.Hatt. ..........................................................................................................285 Frullania sabaliana R.M.Schust. .................................................................................................285 Frullania sabanetica Gottsche ......................................................................................................295 Frullania sackawana Steph. .........................................................................................................285 Frullania saepisdentata S.Hatt. et Streimann ...............................................................................270 Frullania saipanensis S.Hatt. et Koike..........................................................................................280 Frullania sandvicensis Ångstr. ......................................................................................................266 Frullania sarawakensis S.Hatt. .....................................................................................................285 Frullania scalaris S.Hatt. .............................................................................................................291 Frullania scandens Mont. ............................................................................................................285 Frullania schaefer-verwimpii Yuzawa et S.Hatt. ...........................................................................293 Frullania schensiana C.Massal. ....................................................................................................280 Frullania schifneri Verd. .............................................................................................................270 Frullania schimperi Nees .............................................................................................................290 Frullania schimperi var. laciniata Vanden Berghen ......................................................................290 Frullania schusterana S.Hatt. .......................................................................................................270 Frullania schusteri S.Hatt. ...........................................................................................................280 Frullania scottiana S.Hatt............................................................................................................280 Frullania selwyniana Pearson.......................................................................................................293 Frullania semienana Gola ............................................................................................................295 Frullania semivillosa Lindenb. et Gottsche ..................................................................................285 Frullania sergiae Sim-Sim, Fontinha, Mues et Lion .....................................................................293 Frullania seriata Gottsche ...........................................................................................................282 Frullania seriatifolia Steph...........................................................................................................270 Frullania serrata Gottsche ...........................................................................................................270 Frullania serrata var. ceramensis S.Hatt. .......................................................................................270 Frullania serrata subsp. grolleana (S.Hatt.) S.Hatt. ......................................................................270 World checklist of hornworts and liverworts ** ** ** ** ** ** *** ** ** ** ** ** ** ** *** *** ** * ** ** ** *** ** *** ** ** ** *** ** *** *** ** *** ** ** ** *** ** ** ** ** * ** ** ** ** ** ** * ** ** ** 569 Frullania serrata var. hamatispina (S.Hatt.) S.Hatt. .....................................................................270 Frullania serrata var. pertenuis (Nees) Schifn. .............................................................................270 Frullania serrata subsp. spinistipula S.Hatt. .................................................................................270 Frullania serrifolia Steph. ............................................................................................................295 Frullania setacea S.Hatt. ..............................................................................................................270 Frullania setchellii Pearson ..........................................................................................................280 Frullania setigera Steph. ..............................................................................................................288 Frullania shanensis Svihla ............................................................................................................280 Frullania sharpantha Udar et Ad.Kumar .....................................................................................280 Frullania sharpii S.Hatt...............................................................................................................280 Frullania sheana S.Hatt. ..............................................................................................................270 Frullania simmondsii Steph. ........................................................................................................270 Frullania sinensis Steph. ..............................................................................................................280 Frullania sinosphaerantha S.Hatt. et P.J.Lin ................................................................................280 Frullania sinskeana J.J.Engel et B.C.Tan .....................................................................................283 Frullania sinuata Sande Lac. .......................................................................................................270 Frullania socotrana Mitt. .............................................................................................................283 Frullania solanderiana Colenso ...................................................................................................283 Frullania speciosa Herzog ............................................................................................................289 Frullania spegazzinii M.E.Reiner.................................................................................................264 Frullania sphaerantha S.Hatt. ......................................................................................................280 Frullania sphaerocephala Spruce ..................................................................................................265 Frullania sphaerolobulata S.H.Lin ...............................................................................................280 Frullania spinifera Taylor ............................................................................................................280 Frullania spinigastria S.Hatt. .......................................................................................................280 Frullania spiniplica S.Hatt...........................................................................................................280 Frullania spinistipula Steph. ........................................................................................................282 Frullania spongiosa Steph. ...........................................................................................................280 Frullania squamuligera Spruce.....................................................................................................280 Frullania squarrosula (Hook.f. et Taylor) Gottsche, Lindenb. et Nees .........................................280 Frullania standaertii Steph. .........................................................................................................266 Frullania steereana S.Hatt. ..........................................................................................................270 Frullania stenostipa Spruce ..........................................................................................................280 Frullania stipatiloba Steph...........................................................................................................270 Frullania streimannii S.Hatt........................................................................................................282 Frullania stylifera (R.M.Schust.) R.M.Schust. .............................................................................285 Frullania subarctica Vilnet, Borovich. et Bakalin .........................................................................293 Frullania subcaduca S.Hatt..........................................................................................................280 Frullania subclavata Steph...........................................................................................................280 Frullania subdentata Steph. .........................................................................................................270 Frullania subdentata var. latistipula (S.Hatt.) S.Hatt. ..................................................................270 Frullania subdilatata C.Massal. ...................................................................................................285 Frullania subincumbens S.Hatt. ...................................................................................................283 Frullania sublignosa Steph. ..........................................................................................................292 Frullania submultilacera S.Hatt. et Koike ....................................................................................270 Frullania subnigricaulis S.Hatt. ...................................................................................................280 Frullania subnigricaulis var. subtruncata S.Hatt. ..........................................................................280 Frullania subocellata S.Hatt. ........................................................................................................270 Frullania subpedicellata S.Hatt. ...................................................................................................280 Frullania subpilibracteola S.Hatt. ................................................................................................267 Frullania subpyricalycina Herzog .................................................................................................295 Frullania subsquarrosa S.Hatt. .....................................................................................................280 570 ** * ** ** ** *** *** * *** ** *** *** *** ** *** ** *** *** ** * ** *** * * *** ** ** ** * *** ** ** ** ** ** *** ** *** ** *** *** ** *** * ** ** ** ** * ** ** ** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Frullania subtilissima (Nees ex Mont.) Lindenb. .........................................................................267 Frullania subtropica Steph. ..........................................................................................................283 Frullania subtruncata Steph.........................................................................................................295 Frullania subvalida S.Hatt. et haithong.....................................................................................281 Frullania supradecomposita (Lehm. et Lindenb.) Lehm. et Lindenb. ............................................289 Frullania svihlana S.Hatt. ...........................................................................................................281 Frullania tagawana (S.Hatt. et haithong) S.Hatt.......................................................................287 Frullania taiheizana Horik. .........................................................................................................281 Frullania tamarisci (L.) Dumort. .................................................................................................293 Frullania tamsuina Steph. ...........................................................................................................281 Frullania taradakensis Steph. .......................................................................................................281 Frullania taxodiocola R.M.Schust. ...............................................................................................270 Frullania tenerifae (F.Weber) Nees .............................................................................................293 Frullania tenuirostris Steph. .........................................................................................................281 Frullania ternatensis Gottsche......................................................................................................270 Frullania ternatensis var. non-appendiculata S.Hatt. .....................................................................270 Frullania tetraptera Nees et Mont. ..............................................................................................266 Frullania thiersiae S.Hatt. ...........................................................................................................287 Frullania tixieri S.Hatt. ...............................................................................................................287 Frullania tjibodensis S.Hatt. et haithong....................................................................................283 Frullania togashiana S.Hatt. ........................................................................................................281 Frullania toropuku von Konrat, de Lange et Larraín ....................................................................290 Frullania trianae Gottsche ..........................................................................................................289 Frullania tricarinata Sande Lac. ..................................................................................................272 Frullania trichodes Mitt. ..............................................................................................................271 Frullania trigona L.Clark, Jovet-Ast et Frye .................................................................................293 Frullania trinervis (Lehm.) Drège ................................................................................................266 Frullania triquetra Lindenb. et Gottsche .....................................................................................281 Frullania trollii Herzog ...............................................................................................................289 Frullania truncatistyla von Konrat, Hentschel, Heinrichs et Braggins ..........................................291 Frullania tubercularis S.Hatt. et P.J.Lin.......................................................................................281 Frullania tunguraguana L.Clark et Frye.......................................................................................265 Frullania turfosa Lindenb. et Gottsche ........................................................................................295 Frullania tuyamae S.Hatt. et haithong ......................................................................................284 Frullania udarii V.Nath et Ajit P.Singh ......................................................................................295 Frullania uleana Steph. ...............................................................................................................289 Frullania umbonata Mitt.............................................................................................................285 Frullania usambarana Schifn......................................................................................................272 Frullania usambarana var. reducta Vanden Berghen ....................................................................272 Frullania usamiensis Steph...........................................................................................................281 Frullania utriculata Steph. ..........................................................................................................285 Frullania vaga Mitt. ....................................................................................................................271 Frullania vaginata (Sw.) Nees .....................................................................................................271 Frullania vaginata var. nigricans (Gottsche, Lindenb. et Nees) Schifn. .......................................271 Frullania valdiviensis J.B.Jack et Steph. .......................................................................................281 Frullania valida Steph. ................................................................................................................281 Frullania valparaisiana Lehm. .....................................................................................................295 Frullania vandenberghenii Pócs ...................................................................................................267 Frullania van-zantenii Kamim. et S.Hatt. ...................................................................................271 Frullania variegata Steph. ...........................................................................................................281 Frullania venusta S.Hatt..............................................................................................................271 Frullania verdoorniana S.Hatt. ....................................................................................................271 World checklist of hornworts and liverworts * *** *** ** *** ** ** ** ** ** ** *** *** *** *** ** ** ** ** *** *** *** *** *** *** *** *** *** *** ** *** *** * *** ** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** ** 571 Frullania victoriensis Steph. .........................................................................................................281 Frullania virginica Lehm. ............................................................................................................285 Frullania vitalii Yuzawa et S.Hatt. ..............................................................................................271 Frullania vittata S.Hatt. ..............................................................................................................271 Frullania vittiana S.Hatt. ............................................................................................................281 Frullania vivipara Pócs ...............................................................................................................271 Frullania wagneri Steph. .............................................................................................................295 Frullania wairua von Konrat et Braggins ....................................................................................271 Frullania wangii S.Hatt. et P.J.Lin ..............................................................................................281 Frullania warnckeana S.Hatt. ......................................................................................................271 Frullania warnckeana var. dentosa S.Hatt. ...................................................................................271 Frullania weberbaueri Steph. .......................................................................................................289 Frullania winteri Steph. ..............................................................................................................266 Frullania winteri var. vanderhammenii (Haarbrink) Yuzawa ........................................................266 Frullania yuennanensis Steph. ......................................................................................................281 Frullania yuennanensis var. siamensis (N.Kitag., haithong et S.Hatt.) S.Hatt. et P.J.Lin ............281 Frullania yuzawana S.Hatt..........................................................................................................281 Frullania zangii S.Hatt. et P.J.Lin ...............................................................................................281 Frullania zennoskeana S.Hatt. .....................................................................................................281 Frullanoides bahamensis (A.Evans) van Slageren ..........................................................................405 Frullanoides corticalis (Lehm. et Lindenb.) van Slageren ..............................................................405 Frullanoides densifolia Raddi .......................................................................................................405 Frullanoides densifolia subsp. grandidentata (L.Clark) van Slageren .............................................405 Frullanoides laciniatilora (Loitl.) van Slageren ............................................................................405 Frullanoides liebmaniana (Lindenb. et Gottsche) van Slageren ....................................................405 Frullanoides mexicana van Slageren .............................................................................................405 Frullanoides tristis (Steph.) van Slageren ......................................................................................405 Fulfordianthus evansii (Fulford) Gradst. ......................................................................................405 Fulfordianthus pterobryoides (Spruce) Gradst. ..............................................................................405 Fuscocephaloziopsis ainis (Lindb. ex Steph.) Váňa et L.Söderstr. ...................................................59 Fuscocephaloziopsis africana (Váňa) Váňa et L.Söderstr. .................................................................59 Fuscocephaloziopsis albescens (Hook.) Váňa et L.Söderstr. ..............................................................59 Fuscocephaloziopsis albescens var. islandica (Nees) Váňa et L.Söderstr. ............................................59 Fuscocephaloziopsis baldwinii (C.M.Cooke) Váňa et L.Söderstr. ....................................................59 Fuscocephaloziopsis biloba (Herzog) Fulford ...................................................................................59 Fuscocephaloziopsis catenulata (Huebener) Váňa et L.Söderstr........................................................59 Fuscocephaloziopsis catenulata subsp. nipponica (S.Hatt.) Váňa et L.Söderstr..................................59 Fuscocephaloziopsis connivens (Dicks.) Váňa et L.Söderstr. .............................................................60 Fuscocephaloziopsis connivens subsp. issa (Steph.) Váňa et L.Söderstr. ............................................60 Fuscocephaloziopsis connivens subsp. sandwicensis (Mont.) Váňa et L.Söderstr. ...............................60 Fuscocephaloziopsis crassifolia (Lindenb. et Gottsche) Váňa et L.Söderstr. ......................................60 Fuscocephaloziopsis gollanii (Steph.) Váňa et L.Söderstr. ................................................................60 Fuscocephaloziopsis leucantha (Spruce) Váňa et L.Söderstr..............................................................60 Fuscocephaloziopsis loitlesbergeri (Schifn.) Váňa et L.Söderstr. .......................................................60 Fuscocephaloziopsis lunulifolia (Dumort.) Váňa et L.Söderstr. ........................................................60 Fuscocephaloziopsis macrostachya (Kaal.) Váňa et L.Söderstr. ..........................................................60 Fuscocephaloziopsis macrostachya subsp. australis (R.M.Schust.) Váňa et L.Söderstr........................60 Fuscocephaloziopsis macrostachya subsp. macrostachya var. spinilora (Schifn.) Váňa et L.Söderstr. .60 Fuscocephaloziopsis monticola (J.D.Godfrey) Váňa et L.Söderstr.....................................................60 Fuscocephaloziopsis pachycaulis (R.M.Schust.) Váňa et L.Söderstr. .................................................60 Fuscocephaloziopsis pleniceps (Austin) Váňa et L.Söderstr. ..............................................................61 Fuscocephaloziopsis pleniceps var. caroliniana (R.M.Schust.) Váňa et L.Söderstr..............................61 572 *** * *** *** *** *** ** *** *** *** ** *** *** *** ** ** *** *** *** *** *** *** *** *** *** *** *** *** *** *** ** *** *** *** *** ** *** ** *** * *** *** *** *** *** *** *** *** *** *** *** *** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Fuscocephaloziopsis pulvinata (Steph.) Fulford ...............................................................................61 Fuscocephaloziopsis schusteri (Sushil K.Singh et D.K.Singh) Váňa ..................................................61 Fuscocephaloziopsis subintegra Gradst. et Váňa ...............................................................................61 Fuscocephaloziopsis zoopsioides (Horik.) Váňa et L.Söderstr. ...........................................................61 Gackstroemia alpina R.M.Schust. ................................................................................................416 Gackstroemia hariotiana (Besch. et C.Massal.) Grolle ..................................................................415 Gackstroemia ljungneri (Herzog) Grolle .......................................................................................415 Gackstroemia magellanica (Lam.) Trevis. .....................................................................................415 Gackstroemia novae-zelandiae R.M.Schust. et J.J.Engel ...............................................................415 Gackstroemia patagonica (Steph.) Grolle ......................................................................................416 Gackstroemia schwabei (Herzog) Grolle .......................................................................................416 Gackstroemia weindorferi (Herzog) Grolle ...................................................................................415 Geocalyx caledonicus Steph...........................................................................................................109 Geocalyx graveolens (Schrad.) Nees ..............................................................................................109 Geocalyx lancistipulus (Steph.) S.Hatt. .........................................................................................109 Geocalyx orientalis Besch. et Spruce .............................................................................................109 Geothallus tuberosus Campb. .......................................................................................................506 Gerhildiella rossneriana Grolle .......................................................................................................78 Goebeliella cornigera (Mitt.) Steph...............................................................................................415 Goebelobryum grossitextum (Steph.) Grolle.....................................................................................97 Goebelobryum unguiculatum (Hook.f. et Taylor) Grolle ................................................................97 Goebelobryum vermiculare J.J.Engel et Glenny...............................................................................97 Gongylanthus dusenii Steph. ........................................................................................................135 Gongylanthus ericetorum (Raddi) Nees.........................................................................................135 Gongylanthus granatensis (Gottsche) Steph. .................................................................................135 Gongylanthus himalayensis Grolle ................................................................................................135 Gongylanthus javanicus Grolle .....................................................................................................135 Gongylanthus liebmanianus (Lindenb. et Gottsche) Steph............................................................135 Gongylanthus limbatus (Herzog) Grolle et Váňa ..........................................................................135 Gongylanthus muelleri (Gottsche) Steph. .....................................................................................135 Gongylanthus oniscoides (Spruce) Steph........................................................................................135 Gongylanthus richardsii E.W.Jones ..............................................................................................135 Gottschelia maxima (Steph.) Grolle ...............................................................................................74 Gottschelia schizopleura (Spruce) Grolle .........................................................................................74 Greeneothallus gemmiparus Hässel ...............................................................................................471 Grollea antheliopsis R.M.Schust. ..................................................................................................138 Gymnocolea borealis (Frisvoll et Moen) R.M.Schust. .....................................................................52 Gymnocolea fascinifera Potemkin ...................................................................................................52 Gymnocolea inlata (Huds.) Dumort. ............................................................................................52 Gymnocolea inlata subsp. acutiloba (Schifn.) R.M.Schust. et Damsh. ex L.Söderstr. et Váňa .......52 Gymnocoleopsis capensis (S.W.Arnell) R.M.Schust. ........................................................................74 Gymnocoleopsis cylindriformis (Mitt.) R.M.Schust. .........................................................................74 Gymnomitrion adustum Nees.......................................................................................................110 Gymnomitrion africanum (Steph.) Horik. ....................................................................................110 Gymnomitrion alpinum (Gottsche ex Husn.) Schifn. ..................................................................110 Gymnomitrion asperulatum R.M.Schust. .....................................................................................110 Gymnomitrion atroilum Váňa .....................................................................................................110 Gymnomitrion bolivianum (Steph.) Váňa ....................................................................................110 Gymnomitrion brevissimum (Dumort.) Warnst. ...........................................................................110 Gymnomitrion commutatum (Limpr.) Schifn. .............................................................................111 Gymnomitrion concinnatum (Lightf.) Corda ................................................................................111 Gymnomitrion corallioides Nees ...................................................................................................111 World checklist of hornworts and liverworts *** *** *** ** *** *** *** ** *** ** *** *** *** *** *** *** ** *** *** *** ** ** *** *** *** *** *** *** *** *** ** *** *** *** *** *** *** *** *** * *** ** *** ** ** ** ** *** ** * *** ** 573 Gymnomitrion crenatilobum Grolle..............................................................................................111 Gymnomitrion crenulatum Gottsche ex Carrington......................................................................111 Gymnomitrion crystallocaulon (Grolle) Váňa, Crand.-Stotl. et Stotler ..........................................111 Gymnomitrion incompletum (Gottsche) R.M.Schust. ex Váňa .....................................................111 Gymnomitrion laceratum (Steph.) Horik. ....................................................................................111 Gymnomitrion miniatum Lindenb. et Gottsche ...........................................................................111 Gymnomitrion minutulum (Hässel) Váňa ....................................................................................111 Gymnomitrion moralesae Váňa.....................................................................................................111 Gymnomitrion mucronulatum (N.Kitag.) N.Kitag. ......................................................................111 Gymnomitrion mucrophorum R.M.Schust....................................................................................111 Gymnomitrion nigrum (Grolle et Váňa) Váňa ..............................................................................111 Gymnomitrion noguchianum S.Hatt. ...........................................................................................111 Gymnomitrion obtusilobum N.Kitag. ...........................................................................................111 Gymnomitrion obtusum Lindb. ....................................................................................................111 Gymnomitrion paciicum Grolle ...................................................................................................111 Gymnomitrion revolutum (Nees) H.Philib. ..................................................................................111 Gymnomitrion revolutum subsp. novoguineanensis (R.M.Schust.) Váňa, Crand.-Stotl. et Stotler ..112 Gymnomitrion rubidum (Mitt.) Váňa, Crand.-Stotl. et Stotler ....................................................112 Gymnomitrion setaceum Grolle et Váňa .......................................................................................112 Gymnomitrion sinense Müll.Frib. .................................................................................................112 Gymnomitrion strictum (Berggr.) R.M.Schust..............................................................................112 Gymnomitrion strictum var. inaequale R.M.Schust.......................................................................112 Gymnomitrion subintegrum (S.W.Arnell) Váňa ............................................................................112 Gymnomitrion truncatoapiculatum Herzog ..................................................................................112 Gymnomitrion verrucosum W.E.Nicholson ..................................................................................112 Gyrothyra underwoodiana M.Howe .............................................................................................115 Haesselia acuminata Gradst. ..........................................................................................................64 Haesselia roraimensis Grolle et Gradst............................................................................................64 Hamatostrepta concinna Váňa et D.G.Long ...................................................................................52 Haplolejeunea cucullata (Steph.) Grolle .......................................................................................348 Haplolejeunea sticta Grolle ..........................................................................................................348 Haplomitrium blumei (Nees) R.M.Schust. ....................................................................................41 Haplomitrium gibbsiae (Steph.) R.M.Schust. .................................................................................42 Haplomitrium hookeri (Lyell ex Sm.) Nees ....................................................................................42 Haplomitrium hookeri var. minutum (E.O.Campb.) Barthol.-Began ..............................................42 Haplomitrium intermedium Berrie .................................................................................................42 Haplomitrium mnioides (Lindb.) R.M.Schust. ...............................................................................41 Haplomitrium monoicum J.J.Engel ................................................................................................42 Haplomitrium ovalifolium R.M.Schust. .........................................................................................42 Harpalejeunea acuta S.Winkl. .....................................................................................................362 Harpalejeunea ancistrodes (Spruce) Schifn. .................................................................................362 Harpalejeunea buenaventurae Herzog ..........................................................................................362 Harpalejeunea cinchonae (Nees) Schifn.......................................................................................362 Harpalejeunea cinchonae var. strigulosa Herzog ............................................................................363 Harpalejeunea decurvicuspis (Besch. et C.Massal.) P.Syd..............................................................363 Harpalejeunea emarginata Jovet-Ast ............................................................................................363 Harpalejeunea exocellata Herzog ..................................................................................................363 Harpalejeunea grandis Grolle et M.E.Reiner ................................................................................362 Harpalejeunea grandistipula R.M.Schust. ....................................................................................363 Harpalejeunea grossearmata Steph. ...............................................................................................364 Harpalejeunea harpaphylla (Herzog) Bischl. ................................................................................363 Harpalejeunea herzogii Jovet-Ast .................................................................................................363 574 ** ** *** ** *** *** ** ** * ** *** * * *** ** * *** ** * * *** *** *** *** *** ** *** ** *** *** *** *** ** ** ** *** ** ** ** ** *** *** ** * ** ** ** ** ** ** *** *** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Harpalejeunea longibracteata (Spruce) Steph................................................................................363 Harpalejeunea marginalis (Hook.f. et Taylor) Steph. ...................................................................363 Harpalejeunea molleri (Steph.) Grolle ..........................................................................................363 Harpalejeunea molleri subsp. integra (R.M.Schust.) Damsh. ........................................................363 Harpalejeunea oxyphylla (Nees et Mont.) Steph. ..........................................................................363 Harpalejeunea parasitica (Hook.f. et Taylor) Steph. ....................................................................363 Harpalejeunea pinaundensis Grolle ..............................................................................................363 Harpalejeunea relexula A.Evans ..................................................................................................363 Harpalejeunea renneri Herzog .....................................................................................................364 Harpalejeunea scabra Gradst. et Schäf.-Verw. ..............................................................................363 Harpalejeunea schifneri S.W.Arnell .............................................................................................363 Harpalejeunea solitaria (Gottsche) Steph. ....................................................................................363 Harpalejeunea spruceana Steph. ...................................................................................................364 Harpalejeunea stricta (Lindenb. et Gottsche) Steph. ....................................................................364 Harpalejeunea subacuta A.Evans ..................................................................................................364 Harpalejeunea tenuicuspis (Spruce) Schifn. .................................................................................364 Harpalejeunea tridens (Besch. et Spruce) Steph............................................................................364 Harpalejeunea uncinata Steph. ....................................................................................................364 Harpalejeunea uncinata var. setulosa Herzog ................................................................................364 Harpalejeunea vitrea Herzog .......................................................................................................364 Harpanthus drummondii (Taylor) Grolle .....................................................................................116 Harpanthus lotovianus (Nees) Nees ............................................................................................116 Harpanthus scutatus (F.Weber et D.Mohr) Spruce ......................................................................116 Hattoria yakushimensis (Horik.) R.M.Schust. ................................................................................53 Hattorianthus erimonus (Steph.) R.M.Schust. et Inoue ................................................................469 Hattoriolejeunea akiyamae Mizut. ................................................................................................364 Hepatostolonophora abnormis (Besch. et C.Massal.) J.J.Engel et R.M.Schust. ...............................196 Hepatostolonophora conica (Steph.) Hässel ...................................................................................196 Hepatostolonophora paucistipula (Rodway) J.J.Engel ....................................................................196 Hepatostolonophora rotata (Hook.f. et Taylor) J.J.Engel...............................................................196 Hepatostolonophora rotata var. perssonii (R.M.Schust.) J.J.Engel ..................................................197 Herbertus aduncus (Dicks.) Gray .................................................................................................138 Herbertus arcticus (Inoue et Steere) Schljakov ..............................................................................139 Herbertus armitanus (Steph.) H.A.Mill........................................................................................139 Herbertus asparus Tixier ..............................................................................................................139 Herbertus bivittatus Spruce ..........................................................................................................139 Herbertus borealis Crundw. .........................................................................................................139 Herbertus buchii Juslén ................................................................................................................139 Herbertus ceylanicus (Steph.) Abeyw. ...........................................................................................139 Herbertus circinatus (Steph.) H.A.Mill. .......................................................................................139 Herbertus delavayi (Steph.) Steph. ...............................................................................................139 Herbertus dicranus (Gottsche, Lindenb. et Nees) Trevis. .............................................................139 Herbertus durandii (Steph.) Herzog ............................................................................................139 Herbertus evittatus (Steph.) H.A.Mill. .........................................................................................139 Herbertus gaochienii X.Fu ............................................................................................................139 Herbertus gracilis (Mont.) Steph. .................................................................................................139 Herbertus guangdongii P.J.Lin ex Piippo ......................................................................................139 Herbertus hawaiiensis H.A.Mill. ..................................................................................................139 Herbertus helleri (Steph.) W.E.Nicholson....................................................................................139 Herbertus herpocladioides E.B.Scott et H.A.Mill. .........................................................................139 Herbertus hutchinsiae (Gottsche et Rabenh.) A.Evans ..................................................................140 Herbertus juniperoideus (Sw.) Grolle ............................................................................................140 World checklist of hornworts and liverworts *** *** ** ** ** ** ** ** *** ** ** ** ** ** ** *** ** *** ** * *** ** * *** *** *** *** *** *** *** *** *** ** ** ** ** ** *** ** *** ** ** ** *** ** ** * * ** ** *** ** 575 Herbertus juniperoideus subsp. acanthelius (Spruce) K.Feldberg et Heinrichs ...............................140 Herbertus juniperoideus subsp. pensilis (Taylor) K.Feldberg et Heinrichs......................................140 Herbertus kurzii (Steph.) R.S.Chopra ..........................................................................................140 Herbertus leratii (Steph.) H.A.Mill. .............................................................................................140 Herbertus lonchobasis H.A.Mill. ...................................................................................................140 Herbertus longiissus Steph. ..........................................................................................................140 Herbertus longispinus J.B.Jack et Steph. .......................................................................................140 Herbertus mauritianus N.G.Hodgetts ..........................................................................................140 Herbertus norenus D.G.Long, D.Bell et H.H.Blom .....................................................................140 Herbertus oldieldianus (Steph.) Rodway ......................................................................................140 Herbertus pilifer Schifn...............................................................................................................140 Herbertus pocsii N.G.Hodgetts ....................................................................................................140 Herbertus pumilus Steph. .............................................................................................................140 Herbertus ramosus (Steph.) H.A.Mill. ..........................................................................................140 Herbertus runcinatus (Taylor) Trevis. ..........................................................................................140 Herbertus sendtneri (Nees) Lindb.................................................................................................140 Herbertus spicatus N.G.Hodgetts .................................................................................................140 Herbertus stramineus (Dumort.) Trevis. .......................................................................................141 Herbertus streimannii M.L.So ......................................................................................................141 Herbertus subrotundatus X.Fu et Y.J.Yi ........................................................................................141 Herbertus tenuis A.Evans .............................................................................................................141 Herbertus udarii D.Kumar et N.Manocha ...................................................................................141 Herzogianthus sanguineus R.M.Schust. ........................................................................................434 Herzogianthus vaginatus (Herzog) R.M.Schust. ...........................................................................434 Herzogiaria teres (Steph.) Fulford ex Hässel.................................................................................254 Herzogobryum atrocapillum (Hook.f. et Taylor) Grolle ..................................................................74 Herzogobryum iliforme R.M.Schust. .............................................................................................74 Herzogobryum molle Grolle ...........................................................................................................74 Herzogobryum vermiculare (Schifn.) Grolle...................................................................................74 Heterogemma capitata (Hook.) Konstant. et Vilnet .......................................................................79 Heterogemma laxa (Lindb.) Konstant. et Vilnet .............................................................................80 Heterogemma patagonica (Herzog et Grolle) L.Söderstr. et Váňa ...................................................80 Heteroscyphus acutangulus (Schifn.) Schifn. ...............................................................................197 Heteroscyphus allodontus (Hook.f. et Taylor) J.J.Engel et R.M.Schust. .........................................197 Heteroscyphus amboinensis (Schifn.) Schifn. ...............................................................................197 Heteroscyphus ammophilus (Colenso) R.M.Schust. .......................................................................197 Heteroscyphus ammophilus var. obtusifolius J.J.Engel et G.L.Merr. ................................................197 Heteroscyphus argutus (Reinw., Blume et Nees) Schifn................................................................197 Heteroscyphus argutus var. brevidens (Schifn.) Herzog et Nog......................................................197 Heteroscyphus aselliformis (Reinw., Blume et Nees) Schifn. .........................................................197 Heteroscyphus assurgentifolius J.J.Engel .........................................................................................197 Heteroscyphus baduinus (Nees) Schifn.........................................................................................197 Heteroscyphus balnetii (Herzog) Grolle ........................................................................................197 Heteroscyphus billardierei (Schwägr.) Schifn................................................................................197 Heteroscyphus billardierei var. clasmatocoleoides (J.J.Engel et G.L.Merr.) J.J.Engel ........................197 Heteroscyphus brassii (Grolle) Grolle ............................................................................................198 Heteroscyphus caesius (Schifn.) Schifn. .......................................................................................198 Heteroscyphus caledonicus (Steph.) Schifn....................................................................................198 Heteroscyphus chlorophyllus (Hook.f. et Taylor) Schifn................................................................198 Heteroscyphus ciliatus (Steph.) Schifn..........................................................................................198 Heteroscyphus coalitus (Hook.) Schifn. ........................................................................................198 Heteroscyphus coalitus var. simplicifolius J.J.Engel .........................................................................198 576 ** *** *** *** ** ** *** ** ** ** ** ** *** ** ** * * *** ** ** ** ** * ** ** ** ** ** ** ** ** ** ** *** ** ** *** ** ** ** ** *** *** ** ** *** ** ** *** ** ** ** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Heteroscyphus combinatus (Nees) Schifn......................................................................................198 Heteroscyphus conjugatus (Mitt.) J.J.Engel et R.M.Schust.............................................................198 Heteroscyphus contortuplicatus (Nees et Mont.) Grolle .................................................................198 Heteroscyphus cuneistipulus (Steph.) Schifn. ................................................................................198 Heteroscyphus darjeelingensis A.Srivast. et S.C.Srivast. ..................................................................198 Heteroscyphus deceptifrons J.J.Engel ..............................................................................................198 Heteroscyphus dentammophilus J.J.Engel et G.L.Merr...................................................................198 Heteroscyphus denticulatus (Mitt.) Schifn. ...................................................................................199 Heteroscyphus deplanchei (Steph.) Schifn. ...................................................................................199 Heteroscyphus diestianus (Sande Lac.) Piippo ...............................................................................199 Heteroscyphus divergenticiliatus (Steph.) Fulford ..........................................................................199 Heteroscyphus dubius (Gottsche) Schifn. .....................................................................................199 Heteroscyphus echinellus (Lindenb. et Gottsche) J.J.Engel et Xiao L.He .......................................199 Heteroscyphus echinellus var. hyalinus J.J.Engel .............................................................................199 Heteroscyphus elliottii (Steph.) Pagán ...........................................................................................199 Heteroscyphus elliottii var. portoricensis Fulford.............................................................................199 Heteroscyphus falcifolius (Steph.) Schifn. .....................................................................................199 Heteroscyphus issistipus (Hook.f. et Taylor) Schifn. ....................................................................199 Heteroscyphus issistipus var. multispinus (E.A.Hodgs. et Allison) J.J.Engel ...................................199 Heteroscyphus issistipus var. repandus J.J.Engel.............................................................................199 Heteroscyphus laccidus (Mitt.) A.Srivast. et S.C.Srivast. ...............................................................199 Heteroscyphus leischeri (Steph.) D.G.Long et Rubas. ...................................................................199 Heteroscyphus fragilicilius (Schifn.) Schifn. ................................................................................200 Heteroscyphus furcistipulus (E.A.Hodgs.) J.J.Engel et R.M.Schust. ...............................................200 Heteroscyphus giganteus (Steph.) Hürl. .........................................................................................200 Heteroscyphus graefei (J.B.Jack et Steph.) Grolle..........................................................................200 Heteroscyphus grandilorus (Steph.) Hürl. .....................................................................................200 Heteroscyphus grandistipus (Steph.) Schifn. .................................................................................200 Heteroscyphus gunnianus (Mitt.) J.J.Engel et R.M.Schust.............................................................200 Heteroscyphus hastatus (E.A.Hodgs.) J.J.Engel et R.M.Schust. .....................................................200 Heteroscyphus hebridensis (Steph.) Schifn. ...................................................................................200 Heteroscyphus heterophyllus (Steph.) J.J.Engel et R.M.Schust. ......................................................200 Heteroscyphus hyalinus (Steph.) A.Srivast. et S.C.Srivast. .............................................................200 Heteroscyphus integrifolius (Lehm. et Lindenb.) Fulford ...............................................................200 Heteroscyphus iwatsukii (S.Hatt.) Piippo ......................................................................................200 Heteroscyphus jackii (Steph.) Schifn. ...........................................................................................200 Heteroscyphus knightii (Steph.) Grolle..........................................................................................201 Heteroscyphus levieri (Steph.) Schifn. ..........................................................................................201 Heteroscyphus limosus (Carrington et Pearson) Schifn. ................................................................201 Heteroscyphus lingulatus (Colenso) J.J.Engel et R.M.Schust. ........................................................201 Heteroscyphus lophocoleoides S.Hatt. .............................................................................................201 Heteroscyphus lyallii (Mitt.) R.M.Schust. .....................................................................................201 Heteroscyphus magellanicus (Steph.) J.J.Engel et R.M.Schust. ......................................................201 Heteroscyphus mamillatus Piippo..................................................................................................201 Heteroscyphus marginatus (Steph.) Fulford ...................................................................................201 Heteroscyphus menziesii (Mitt.) J.J.Engel......................................................................................201 Heteroscyphus merapiensis (Steph.) Piippo ....................................................................................201 Heteroscyphus miradorensis (Steph.) Schifn. ................................................................................201 Heteroscyphus mononuculus J.J.Engel ...........................................................................................201 Heteroscyphus mononuculus var. ammophilopsis J.J.Engel ..............................................................201 Heteroscyphus mononuculus var. bilobus J.J.Engel .........................................................................201 Heteroscyphus montagnei (Steph.) Fulford ....................................................................................201 World checklist of hornworts and liverworts ** ** ** ** ** ** *** ** ** ** ** *** ** *** *** ** ** ** ** *** ** ** *** ** *** *** ** ** *** ** *** ** ** ** *** ** ** ** ** ** *** ** * * * * * * * *** ** *** 577 Heteroscyphus multiidus (Steph.) J.J.Engel et R.M.Schust. ..........................................................202 Heteroscyphus multiidus var. subintegerrimus J.J.Engel .................................................................202 Heteroscyphus nadeaudii (Steph.) Schifn. ....................................................................................202 Heteroscyphus orbiculatus A.Srivast. et S.C.Srivast. .......................................................................202 Heteroscyphus palniensis A.Srivast. et S.C.Srivast. .........................................................................202 Heteroscyphus pandei S.C.Srivast. et A.Srivast. .............................................................................202 Heteroscyphus parallelifolius J.J.Engel ...........................................................................................202 Heteroscyphus parvulus (Schifn.) Schifn. ....................................................................................202 Heteroscyphus parvus A.Srivast. et S.C.Srivast. .............................................................................202 Heteroscyphus perfoliatus (Mont.) Schifn. ....................................................................................202 Heteroscyphus pertusus (Lehm.) Fulford........................................................................................202 Heteroscyphus planiusculus (Hook.f. et Taylor) J.J.Engel ..............................................................202 Heteroscyphus planus (Mitt.) Schifn. ...........................................................................................202 Heteroscyphus polyblepharis (Spruce) Schifn. ...............................................................................202 Heteroscyphus polychaetus (Spruce) Hentschel et Heinrichs ..........................................................202 Heteroscyphus polycladus (Hook.f. et Lév.) R.M.Schust. ...............................................................202 Heteroscyphus rectangulatus (Herzog) Piippo ................................................................................203 Heteroscyphus saccogynoides Herzog ..............................................................................................203 Heteroscyphus sarawaketanus Piippo .............................................................................................203 Heteroscyphus sinuosus (Hook.) Schifn. .......................................................................................203 Heteroscyphus spectabilis (Steph.) Schifn. ....................................................................................203 Heteroscyphus spinifer C.Gao, T.Cao et Y.H.Wu .........................................................................203 Heteroscyphus splendens (Lehm. et Lindenb.) Grolle.....................................................................203 Heteroscyphus splendidus (E.A.Hodgs.) J.J.Engel et R.M.Schust. ..................................................203 Heteroscyphus stolonifer J.J.Engel..................................................................................................203 Heteroscyphus succulentus (Gottsche) Schifn. ...............................................................................203 Heteroscyphus supinus (Hook.f. et Taylor) R.M.Schust. ...............................................................203 Heteroscyphus tener (Steph.) Schifn. ............................................................................................203 Heteroscyphus thraustus (Spruce) Fulford .....................................................................................203 Heteroscyphus timppae Piippo ......................................................................................................203 Heteroscyphus triacanthus (Hook.f. et Lév.) Schifn. .....................................................................203 Heteroscyphus triacanthus var. magnistipulatus J.J.Engel ...............................................................203 Heteroscyphus tridentatus (Sande Lac.) Grolle ..............................................................................203 Heteroscyphus turgidus (Schifn.) Schifn. .....................................................................................204 Heteroscyphus valdiviensis (Mont.) Schifn. ..................................................................................204 Heteroscyphus varians (Steph.) J.J.Engel .......................................................................................204 Heteroscyphus wettsteinii (Schifn.) Schifn. ..................................................................................204 Heteroscyphus zollingeri (Gottsche) Schifn. .................................................................................204 Hyalolepidozia bicuspidata (C.Massal.) S.W.Arnell ex Grolle .......................................................183 Hygrobiella intermedia Bakalin et Vilnet......................................................................................116 Hygrobiella laxifolia (Hook.) Spruce............................................................................................116 Hygrobiella squamosa Bakalin et Vilnet ........................................................................................116 Hygrolejeunea cubensis Steph. ......................................................................................................509 Hygrolejeunea harpaphylla Steph..................................................................................................509 Hygrolejeunea paciica Steph. .......................................................................................................509 Hygrolejeunea parvicalycina Steph................................................................................................509 Hygrolejeunea parvistipula Steph..................................................................................................509 Hygrolejeunea patellirostris Steph. ................................................................................................509 Hygrolejeunea staudtiana Steph....................................................................................................509 Hygrolembidium acrocladum (Berggr.) R.M.Schust......................................................................164 Hygrolembidium andinum (Herzog) R.M.Schust. ........................................................................164 Hygrolembidium australe (Steph.) Grolle .....................................................................................164 578 *** ** *** *** ** *** *** *** * * *** ** *** *** *** *** *** ** *** *** * ** ** *** ** *** ** * *** ** *** ** * *** *** *** *** *** ** ** *** ** *** ** ** *** *** * *** * *** ** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Hygrolembidium boschianum (Sande Lac.) R.M.Schust................................................................164 Hygrolembidium isophyllum R.M.Schust......................................................................................164 Hygrolembidium rigidum R.M.Schust. et J.J.Engel ......................................................................164 Hygrolembidium triquetrum J.J.Engel et R.M.Schust. ..................................................................164 Hygrolembidium ventrosum (Mitt.) Grolle ...................................................................................164 Hymenophyton labellatum (Labill.) Dumort. ex Trevis. ...............................................................468 Hymenophyton leptopodum (Hook.f. et Taylor) A.Evans ..............................................................468 Hymenophyton pedicellatum Steph. ..............................................................................................469 Hypenantron brachypus Steph. ex Lamothe ..................................................................................509 Hypenantron brasiliense Steph. ex Lamothe .................................................................................509 Isolembidium anomalum (Rodway) Grolle ...................................................................................164 Isolembidium anomalum var. cucullatum (E.A.Hodgs.) J.J.Engel et R.M.Schust. .........................164 Isopaches alboviridis (R.M.Schust.) Schljakov ................................................................................53 Isopaches bicrenatus (Schmidel ex Hofm.) H.Buch........................................................................53 Isopaches decolorans (Limpr.) H.Buch ............................................................................................53 Isopaches pumicicola (Berggr.) Bakalin ...........................................................................................53 Isophyllaria attenuata (Rodway) E.A.Hodgs. ...............................................................................254 Isophyllaria fuegiana (Hässel) R.M.Schust. ..................................................................................254 Isotachis armata (Nees) Gottsche .................................................................................................101 Isotachis aubertii (Schwägr.) Mitt. ...............................................................................................101 Isotachis boliviensis Gottsche ........................................................................................................101 Isotachis chinensis C.Gao, T.Cao et J.Sun ....................................................................................101 Isotachis erythrorhiza (Lehm. et Lindenb.) Spruce........................................................................101 Isotachis fragilis Steph. .................................................................................................................101 Isotachis grandis Carrington et Pearson ........................................................................................101 Isotachis grossidens Steph. .............................................................................................................101 Isotachis hastatistipula (Steph.) J.J.Engel ......................................................................................101 Isotachis hians Steph. ...................................................................................................................101 Isotachis humectata (Hook.f. et Taylor) Steph..............................................................................101 Isotachis inlata Steph. .................................................................................................................101 Isotachis intortifolia (Hook.f. et Taylor) Gottsche ........................................................................101 Isotachis japonica Steph. ..............................................................................................................101 Isotachis lacustris Herzog .............................................................................................................101 Isotachis lopezii (R.M.Schust.) Gradst..........................................................................................101 Isotachis lyallii Mitt. ....................................................................................................................101 Isotachis minima Pearson .............................................................................................................101 Isotachis montana Colenso ...........................................................................................................101 Isotachis multiceps (Lindenb. et Gottsche) Gottsche.....................................................................102 Isotachis multiceps var. fendleri Gottsche ......................................................................................102 Isotachis nigella Herzog ...............................................................................................................102 Isotachis obtusa Steph. .................................................................................................................102 Isotachis olivacea R.M.Schust.......................................................................................................102 Isotachis plicata J.J.Engel .............................................................................................................102 Isotachis pusilla Steph. .................................................................................................................102 Isotachis riparia Rodway ..............................................................................................................102 Isotachis serrulata (Sw.) Gottsche .................................................................................................102 Isotachis spegazziniana C.Massal. .................................................................................................102 Isotachis sprucei Beauverd ............................................................................................................102 Isotachis stephanii E.S.Salmon .....................................................................................................102 Isotachis vexans Steph. .................................................................................................................102 Isotachis westlandica (E.A.Hodgs.) R.M.Schust. ..........................................................................102 Jackiella angustifolia Herzog ........................................................................................................116 World checklist of hornworts and liverworts ** *** *** ** ** ** ** ** ** * * *** * *** *** *** *** *** *** ** ** ** *** ** ** *** *** *** *** ** * *** *** * * * * * * * * *** ** * *** * * * ** * * * 579 Jackiella ceylanica Schifn. ex Steph. ............................................................................................116 Jackiella curvata E.A.Hodgs. et Allison........................................................................................117 Jackiella javanica Schifn. ............................................................................................................117 Jackiella javanica var. cavifolia Schifn. ........................................................................................117 Jackiella javanica var. cordifolia Schifn. ......................................................................................117 Jackiella renifolia Schifn. ............................................................................................................117 Jackiella sinensis (W.E.Nicholson) Grolle ....................................................................................117 Jackiella singapurensis Schifn. .....................................................................................................117 Jackiella singapurensis var. philippinensis N.Kitag. ........................................................................117 Jackiella unica Steph. ..................................................................................................................117 Jamesoniella convoluta Steph........................................................................................................509 Jensenia connivens (Colenso) Grolle.............................................................................................469 Jensenia crassifrons (Steph.) S.Schuette et Stotler .........................................................................470 Jensenia decipiens (Mitt.) Grolle ..................................................................................................470 Jensenia diformis (Nees) Grolle ...................................................................................................470 Jensenia lorschuetzii Gronde .......................................................................................................470 Jensenia spinosa (Lindenb. et Gottsche) Grolle ............................................................................470 Jensenia wallisii (J.B.Jack et Steph.) Grolle ..................................................................................470 Jubula blepharophylla Grolle........................................................................................................295 Jubula hattorii Udar et V.Nath ...................................................................................................295 Jubula hattorii var. muthukuzhiana A.E.D.Daniels et P.Daniel ...................................................295 Jubula himalayensis S.C.Srivast. et D.Sharma ..............................................................................296 Jubula hutchinsiae (Hook.) Dumort. ...........................................................................................296 Jubula hutchinsiae subsp. australiae Pócs et A.Cairns ...................................................................296 Jubula hutchinsiae subsp. bogotensis (Steph.) Verd. ......................................................................296 Jubula hutchinsiae subsp. caucasica Konstant. et Vilnet ...............................................................296 Jubula hutchinsiae subsp. japonica (Steph.) Horik. et Ando .........................................................296 Jubula hutchinsiae subsp. javanica (Steph.) Verd. ........................................................................296 Jubula hutchinsiae subsp. pennsylvanica (Steph.) Verd. ................................................................296 Jubula kwangsiensis C.Gao et K.C.Chang....................................................................................296 Jungermannia amentacea Bertol. ..................................................................................................119 Jungermannia atrovirens Dumort. ...............................................................................................118 Jungermannia borealis Damsh. et Váňa........................................................................................118 Jungermannia brasiliensis Raddi...................................................................................................119 Jungermannia chinensis Osbeck ...................................................................................................119 Jungermannia cordata Vill. ..........................................................................................................119 Jungermannia crenulata Schmidel................................................................................................119 Jungermannia creutzeri Kremer ...................................................................................................119 Jungermannia digitata C.F.W.Meissn. ex Spreng. .......................................................................119 Jungermannia dubioides H.A.Mill. ..............................................................................................119 Jungermannia erectii Ajit P.Singh et V.Nath ...............................................................................118 Jungermannia exsertifolia Steph. ..................................................................................................118 Jungermannia exsertifolia subsp. cordifolia (Dumort.) Váňa .........................................................118 Jungermannia fernandeziana Mitt. ..............................................................................................119 Jungermannia gollanii Steph. .......................................................................................................118 Jungermannia hexagona Schwägr. ................................................................................................119 Jungermannia holandriana Kremer ..............................................................................................119 Jungermannia incerta Gottsche ....................................................................................................119 Jungermannia konstantinovae Bakalin et Vilnet............................................................................118 Jungermannia laterilora Hampe ex Gottsche ..............................................................................119 Jungermannia lescuriana Austin ...................................................................................................119 Jungermannia longiretis Besch. et Spruce .....................................................................................119 580 * * * * *** * *** *** * * * * * * * * * * * ** ** *** ** ** ** * ** *** *** *** *** ** * *** *** ** ** *** ** ** ** *** ** ** ** ** ** *** ** ** ** ** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Jungermannia mastigophora Spreng. ............................................................................................119 Jungermannia michelii Mérat ......................................................................................................119 Jungermannia minima Scop. .......................................................................................................119 Jungermannia odorata With. .......................................................................................................119 Jungermannia ovatotrigona (Steph.) Grolle ..................................................................................118 Jungermannia peltata Schmidel ...................................................................................................119 Jungermannia polaris Lindb. ........................................................................................................118 Jungermannia pumila With. ........................................................................................................118 Jungermannia quadridigitata Grif. ..............................................................................................120 Jungermannia sauteri De Not. ex Rabenh. ...................................................................................120 Jungermannia secunda Hampe ex Gottsche .................................................................................120 Jungermannia stereocaulis Bory ....................................................................................................120 Jungermannia submersa Kremer ...................................................................................................120 Jungermannia sullivantiana Austin ..............................................................................................120 Jungermannia supina Hofm........................................................................................................120 Jungermannia tenuis Ehrh. ..........................................................................................................120 Jungermannia uncifolia Steph. .....................................................................................................120 Jungermannia vernicosa Cass. ex Mérat ........................................................................................120 Kingiolejeunea guayanensis H.Rob. ..............................................................................................509 Kurzia abbreviata Mizut. ............................................................................................................166 Kurzia abietinella (Herzog) Grolle ..............................................................................................166 Kurzia bisetula (Steph.) Grolle ....................................................................................................166 Kurzia borneensis Mizut. .............................................................................................................166 Kurzia brasiliensis (Steph.) Grolle................................................................................................166 Kurzia brevicalycina (Steph.) Grolle ............................................................................................166 Kurzia caduciloba R.M.Schust.....................................................................................................166 Kurzia calcarata (Steph.) Grolle ..................................................................................................165 Kurzia capillaris (Sw.) Grolle ......................................................................................................164 Kurzia capillaris subsp. capillaris var. verrucosa (Steph.) Pócs ......................................................165 Kurzia capillaris subsp. paramicola Pócs ......................................................................................165 Kurzia capillaris subsp. stephanii (Renauld ex Steph.) Pócs ..........................................................165 Kurzia compacta (Steph.) Grolle ..................................................................................................165 Kurzia cucullifolia (Steph.) R.M.Schust.......................................................................................166 Kurzia lagellifera (Steph.) Grolle ................................................................................................166 Kurzia fragilifolia R.M.Schust. ....................................................................................................166 Kurzia fragillima (Herzog) Grolle ...............................................................................................166 Kurzia geniculata Mizut. .............................................................................................................166 Kurzia gonyotricha (Sande Lac.) Grolle........................................................................................165 Kurzia hawaica (C.M.Cooke) Grolle ..........................................................................................166 Kurzia helophila R.M.Schust. ......................................................................................................165 Kurzia helophila var. laccida R.M.Schust. ex J.J.Engel ................................................................165 Kurzia hippurioides (Hook.f. et Taylor) Grolle ............................................................................165 Kurzia hippurioides var. ornata J.J.Engel et G.L.Merr. .................................................................165 Kurzia hispida (Steph.) Grolle .....................................................................................................166 Kurzia irregularis (Steph.) Grolle ................................................................................................165 Kurzia lateconica (Steph.) Grolle .................................................................................................167 Kurzia lineariloba Mizut. ............................................................................................................167 Kurzia longicaulis Piippo .............................................................................................................167 Kurzia makinoana (Steph.) Grolle...............................................................................................167 Kurzia mauiensis (H.A.Mill.) H.A.Mill. ......................................................................................167 Kurzia mollis (Steph.) J.J.Engel et R.M.Schust. ...........................................................................167 Kurzia moniliformis J.J.Engel ......................................................................................................165 World checklist of hornworts and liverworts ** *** *** ** *** *** ** ** ** ** ** ** *** ** ** ** ** *** ** *** *** *** *** *** *** *** *** *** *** *** ** *** ** *** ** *** *** ** ** *** *** *** *** ** ** ** *** *** ** *** ** *** 581 Kurzia nemoides (Hook.f. et Taylor) Grolle.................................................................................165 Kurzia nivicola (R.M.Schust.) E.D.Cooper .................................................................................167 Kurzia pallescens Grolle ...............................................................................................................167 Kurzia pallida Piippo ..................................................................................................................167 Kurzia paucilora (Dicks.) Grolle.................................................................................................166 Kurzia quinquespina J.J.Engel et G.L.Merr. ................................................................................167 Kurzia reversa (Carrington et Pearson) Grolle .............................................................................167 Kurzia saddlensis (Besch. et C.Massal.) Grolle .............................................................................165 Kurzia setiformis (De Not.) J.J.Engel et R.M.Schust....................................................................167 Kurzia sexida (Steph.) Grolle......................................................................................................167 Kurzia sinensis K.C.Chang ..........................................................................................................167 Kurzia sylvatica (A.Evans) Grolle ................................................................................................167 Kurzia tasmanica (Steph.) E.D.Cooper .......................................................................................167 Kurzia tayloriana (H.A.Mill.) H.A.Mill. .....................................................................................167 Kurzia tenerrima (Mitt.) Grolle ...................................................................................................168 Kurzia touwii N.Kitag.................................................................................................................168 Kurzia trichoclados (Müll.Frib.) Grolle ........................................................................................168 Kurzia trilobata (R.M.Schust.) R.M.Schust. ...............................................................................168 Kurzia verticellata (Carrington) Grolle ........................................................................................168 Kymatocalyx africanus Váňa et M.Wigginton.................................................................................74 Kymatocalyx dominicensis (Spruce) Váňa........................................................................................74 Kymatocalyx madagascariensis (Steph.) Gradst. et Váňa..................................................................74 Kymatocalyx rhizomaticus (Herzog) Gradst. et Váňa ......................................................................74 Kymatolejeunea bartlettii Grolle ...................................................................................................362 Lamellocolea granditexta (Steph.) J.J.Engel ..................................................................................204 Lamellocolea integrostia J.J.Engel et Glenny .................................................................................204 Leiolejeunea grandilora A.Evans ..................................................................................................362 Leiomitra breviseta (Steph.) R.M.Schust. .....................................................................................256 Leiomitra capillata Lindb. ...........................................................................................................256 Leiomitra elegans (Lehm.) Hässel .................................................................................................256 Leiomitra elliottii (Steph.) R.M.Schust. .......................................................................................256 Leiomitra laccida Spruce.............................................................................................................257 Leiomitra hirticaulis R.M.Schust. ................................................................................................257 Leiomitra lanata (Hook.) R.M.Schust. ........................................................................................257 Leiomitra mastigophoroides R.M.Schust. ......................................................................................257 Leiomitra merrillana (Steph.) T.Katag. ........................................................................................257 Leiomitra paraphyllina Spruce .....................................................................................................257 Leiomitra robusta (Steph.) R.M.Schust. .......................................................................................257 Leiomitra smaragdina Hässel .......................................................................................................257 Leiomitra tomentosa (Sw.) Lindb. ................................................................................................257 Leiosporoceros dussii (Steph.) Hässel ...............................................................................................41 Lejeunea abyssinica (Gola) Cufod. ...............................................................................................365 Lejeunea acanthogona Spruce .......................................................................................................365 Lejeunea acuminata (Lehm. et Lindenb.) Lehm...........................................................................365 Lejeunea acuta Mitt. ....................................................................................................................365 Lejeunea acutata (Steph.) Solari...................................................................................................365 Lejeunea adpressa Nees ................................................................................................................365 Lejeunea aethiopica E.W.Jones ....................................................................................................365 Lejeunea alaskana (R.M.Schust. et Steere) Inoue et Steere ...........................................................365 Lejeunea alata Gottsche ..............................................................................................................365 Lejeunea alata var. patriciae Pócs .................................................................................................365 Lejeunea albescens (Steph.) Mizut. ...............................................................................................365 582 * ** ** ** ** * ** * *** ** *** * ** * *** * ** ** ** * *** ** ** ** * ** ** *** * ** ** ** ** *** ** ** ** ** *** ** ** *** ** ** *** ** ** * *** *** *** ** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Lejeunea albilora Colenso ...........................................................................................................365 Lejeunea aloba Sande Lac. ...........................................................................................................365 Lejeunea alobifolia H.A.Mill........................................................................................................365 Lejeunea amaniensis E.W.Jones ...................................................................................................365 Lejeunea ambigua Lindenb. et Gottsche ......................................................................................365 Lejeunea amentulifera Steph. .......................................................................................................365 Lejeunea androgyna R.M.Schust. .................................................................................................365 Lejeunea angulifolia Mitt. ............................................................................................................365 Lejeunea angusta (Lehm. et Lindenb.) Mont. ..............................................................................366 Lejeunea anisophylla Mont. .........................................................................................................366 Lejeunea anomala Lindenb. et Gottsche ......................................................................................366 Lejeunea antillana Steph. ............................................................................................................366 Lejeunea aphanes Spruce..............................................................................................................366 Lejeunea apiahyna (Steph.) Sushil K.Singh ..................................................................................381 Lejeunea apiculata Sande Lac. .....................................................................................................366 Lejeunea aptycta Gottsche ...........................................................................................................381 Lejeunea aquatica Horik..............................................................................................................366 Lejeunea aquatica var. apiculata S.Hatt. ......................................................................................366 Lejeunea armitii (Steph.) Steph. ..................................................................................................366 Lejeunea asperifolia Steph. ...........................................................................................................366 Lejeunea asperrima Spruce ...........................................................................................................366 Lejeunea asperula (Steph.) Mizut. ................................................................................................366 Lejeunea asprella Spruce ..............................................................................................................366 Lejeunea asthenica Spruce ............................................................................................................381 Lejeunea atheatostipa Spruce ........................................................................................................366 Lejeunea balazsii (Pócs) R.M.Schust............................................................................................380 Lejeunea barbata (Herzog) R.L.Zhu et M.J.Lai ...........................................................................366 Lejeunea bermudiana (A.Evans) R.M.Schust. ..............................................................................366 Lejeunea bethanica Gottsche........................................................................................................381 Lejeunea bidentula Herzog ..........................................................................................................366 Lejeunea biformis Gottsche ..........................................................................................................366 Lejeunea blepharogona Spruce ......................................................................................................366 Lejeunea blomquistii R.M.Schust. ................................................................................................366 Lejeunea boliviensis (Steph.) R.L.Zhu et M.E.Reiner ...................................................................366 Lejeunea bombonasensis Spruce ....................................................................................................381 Lejeunea bornmuelleri (Steph.) M.E.Reiner .................................................................................381 Lejeunea boryana Mont. ..............................................................................................................366 Lejeunea brenanii E.W.Jones .......................................................................................................366 Lejeunea calcicola R.M.Schust. ....................................................................................................366 Lejeunea calcicola var. mexicana R.M.Schust. ..............................................................................367 Lejeunea canariensis (Steph.) Steph..............................................................................................367 Lejeunea cancellata Nees et Mont. ...............................................................................................367 Lejeunea candida (Pócs) R.M.Schust. ..........................................................................................381 Lejeunea cantabrigiensis E.W.Jones ..............................................................................................367 Lejeunea capensis Gottsche ..........................................................................................................367 Lejeunea caracensis Lindenb.........................................................................................................381 Lejeunea caripensis Lindenb. et Gottsche .....................................................................................381 Lejeunea caroliniana Austin .........................................................................................................367 Lejeunea catinulifera Spruce ........................................................................................................367 Lejeunea caulicalyx (Steph.) M.E.Reiner et Goda ........................................................................367 Lejeunea cavifolia (Ehrh.) Lindb..................................................................................................367 Lejeunea caviloba (Steph.) Besch. ................................................................................................367 World checklist of hornworts and liverworts *** ** ** ** ** ** ** ** ** *** *** ** ** ** *** ** ** ** ** ** ** ** ** *** ** *** *** ** ** ** ** ** *** ** *** ** ** ** *** ** ** ** ** ** ** ** *** * ** *** ** ** 583 Lejeunea cerina (Lehm. et Lindenb.) Lehm. et Lindenb...............................................................367 Lejeunea chaishanensis S.H.Lin ....................................................................................................367 Lejeunea chamissonis Lindenb. .....................................................................................................381 Lejeunea chimborazensis Spruce ...................................................................................................381 Lejeunea cladogyna A.Evans .........................................................................................................367 Lejeunea clavata Lindenb. ...........................................................................................................367 Lejeunea claviformis Lindenb. ex Steph........................................................................................367 Lejeunea cochleata Spruce ............................................................................................................367 Lejeunea cocoes Mitt. ...................................................................................................................367 Lejeunea colensoana (Steph.) M.A.M.Renner ...............................................................................367 Lejeunea combuensis O.S.Moura, Ilk.-Borg. et M.E.Reiner ..........................................................381 Lejeunea compacta (Steph.) Steph. ...............................................................................................367 Lejeunea concava Lindenb. et Gottsche .......................................................................................381 Lejeunea concinnula Spruce et Steph. ..........................................................................................367 Lejeunea conformis Nees et Mont. ...............................................................................................381 Lejeunea connatistipula (Steph.) Steph. ........................................................................................367 Lejeunea contracta Mizut. ............................................................................................................368 Lejeunea controversa Gottsche......................................................................................................368 Lejeunea convexiloba M.L.So et R.L.Zhu .....................................................................................368 Lejeunea corcovadae (Steph.) Bischl. ............................................................................................368 Lejeunea cordilora Spruce ...........................................................................................................368 Lejeunea cordistipula Lindenb. et Gottsche ..................................................................................381 Lejeunea corralensis A.Evans ........................................................................................................368 Lejeunea corynantha Spruce .........................................................................................................368 Lejeunea crassiretis Mitt. ..............................................................................................................368 Lejeunea cristulata (Steph.) M.E.Reiner et Goda .........................................................................368 Lejeunea cristulilora (Steph.) M.E.Reiner et Goda ......................................................................368 Lejeunea curviloba Steph. ............................................................................................................380 Lejeunea cuspidistipula (Steph.) Steph. ex Watts ..........................................................................368 Lejeunea cyanomontana R.M.Schust. ...........................................................................................368 Lejeunea cyanophora R.M.Schust. ................................................................................................368 Lejeunea cyathearum E.W.Jones ..................................................................................................368 Lejeunea cyathophora Mitt. ..........................................................................................................368 Lejeunea cyrtotis Spruce ...............................................................................................................381 Lejeunea debilis (Lehm. et Lindenb.) Nees et Mont. ....................................................................381 Lejeunea denticalyx (Steph.) Steph. ..............................................................................................368 Lejeunea denticuspis (Steph.) Mizut. ............................................................................................368 Lejeunea denudata (Pearson) J.J.Engel .........................................................................................368 Lejeunea deplanata Nees ..............................................................................................................368 Lejeunea deplanata var. cuspidata (Steph.) M.E.Reiner ................................................................368 Lejeunea devendrae (Sushil K.Singh) P.K.Verma et K.K.Rawat ...................................................382 Lejeunea diaphana Spruce ...........................................................................................................368 Lejeunea dictyocalyx Spruce .........................................................................................................382 Lejeunea dimorpha T.Kodama .....................................................................................................369 Lejeunea dipterocarpa E.W.Jones .................................................................................................369 Lejeunea dipterota (Eifrig) G.E.Lee ..............................................................................................382 Lejeunea discreta Lindenb. ...........................................................................................................369 Lejeunea disjecta Spruce...............................................................................................................369 Lejeunea diversicuspis Spruce .......................................................................................................369 Lejeunea drehwaldii Heinrichs et Schäf.-Verw. ............................................................................382 Lejeunea drummondii Taylor .......................................................................................................382 Lejeunea duncaniae (Sim) M.E.Reiner .........................................................................................382 584 ** ** ** ** * * * * *** ** *** ** ** * ** ** ** ** ** *** ** ** ** ** ** *** ** ** ** *** ** ** ** ** ** *** ** * ** ** ** ** ** ** ** ** ** *** * *** ** *** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Lejeunea ecarinata (Steph.) J.M.Coult., Barnes et Arthur ............................................................369 Lejeunea eckloniana Lindenb. ......................................................................................................369 Lejeunea edentata L.Söderstr. ......................................................................................................382 Lejeunea eifrigii Mizut.................................................................................................................369 Lejeunea elegans Gottsche ............................................................................................................385 Lejeunea elongella Gottsche .........................................................................................................369 Lejeunea emarginulilora Gottsche ex Steph. ................................................................................382 Lejeunea epibrya Taylor ...............................................................................................................382 Lejeunea erostrata M.E.Reiner et Goda........................................................................................369 Lejeunea estrellamontana M.A.M.Renner et Pócs.........................................................................382 Lejeunea exilis (Reinw., Blume et Nees) Grolle............................................................................369 Lejeunea exilis var. abnormis (Herzog) G.E.Lee............................................................................369 Lejeunea falcata (Pócs et J.Eggers) Pócs .......................................................................................381 Lejeunea fawcettiae D.J.Carr........................................................................................................382 Lejeunea fernandeziana S.W.Arnell .............................................................................................369 Lejeunea irma Mitt.....................................................................................................................369 Lejeunea issistipula (Steph.) Steph...............................................................................................369 Lejeunea laccida Lindenb. et Gottsche ........................................................................................382 Lejeunea lagellaris Spruce ...........................................................................................................369 Lejeunea lava (Sw.) Nees ............................................................................................................369 Lejeunea lava subsp. moorei (Lindb.) R.M.Schust.......................................................................369 Lejeunea lava subsp. orientalis R.M.Schust. ................................................................................369 Lejeunea lava var. pellucida Lindenb. et Gottsche .......................................................................370 Lejeunea lava subsp. tabularis (Spreng.) S.W.Arnell ...................................................................370 Lejeunea lavida Mitt. .................................................................................................................382 Lejeunea lavovirens Ångstr. .........................................................................................................370 Lejeunea leischeri (Steph.) Mizut. ...............................................................................................370 Lejeunea lorida Spruce................................................................................................................382 Lejeunea loridana A.Evans ..........................................................................................................370 Lejeunea fulfordiae (Jovet-Ast) R.L.Zhu ......................................................................................370 Lejeunea fulva Spruce ..................................................................................................................382 Lejeunea fusagasugana Gottsche...................................................................................................370 Lejeunea galeata Spruce ...............................................................................................................370 Lejeunea gayana Gottsche............................................................................................................370 Lejeunea gibbiloba (Steph.) H.A.Mill. .........................................................................................370 Lejeunea glaucescens Gottsche ......................................................................................................370 Lejeunea glaucescens var. acrogyna R.M.Schust. ............................................................................370 Lejeunea glaucescens var. obsoleta R.M.Schust. .............................................................................370 Lejeunea globosilora (Steph.) Steph. ............................................................................................370 Lejeunea gomphocalyx Spruce.......................................................................................................370 Lejeunea gottscheana Lindenb. .....................................................................................................382 Lejeunea gracilicaulis Spruce ........................................................................................................370 Lejeunea gracilipes (Taylor) Spruce ..............................................................................................370 Lejeunea gracilis Steph. ................................................................................................................370 Lejeunea gradsteiniana Pócs .........................................................................................................370 Lejeunea gradsteinii G.E.Lee, Damanhuri et Latif ......................................................................370 Lejeunea graminicolor Spruce.......................................................................................................382 Lejeunea grolleana (Bernecker) R.L.Zhu et W.Ye ........................................................................382 Lejeunea grossecristata (Steph.) E.W.Jones ...................................................................................370 Lejeunea grossiretis (Steph.) M.E.Reiner et Goda .........................................................................371 Lejeunea grossistipula Steph. ........................................................................................................371 Lejeunea grossitexta (Steph.) M.E.Reiner et Goda ........................................................................371 World checklist of hornworts and liverworts * ** * ** ** ** *** *** ** ** * ** ** ** ** ** * ** ** ** * *** *** *** ** ** ** *** ** ** * ** ** ** ** ** *** *** ** ** *** ** ** * ** ** ** * ** * ** ** 585 Lejeunea grossiuscula Gottsche ex Steph. ......................................................................................371 Lejeunea hahnii Solari .................................................................................................................371 Lejeunea haitica Nees et Mont. ...................................................................................................371 Lejeunea hawaikiana M.A.M.Renner et de Lange .......................................................................371 Lejeunea helenae Pearson .............................................................................................................371 Lejeunea helmsiana (Steph.) Steph. ..............................................................................................371 Lejeunea hepaticola (Steph.) Steph. ..............................................................................................371 Lejeunea herminieri (Steph.) R.L.Zhu .........................................................................................382 Lejeunea heterocheila Taylor ........................................................................................................383 Lejeunea hibernica Bischl., H.A.Mill. et Bonner ex Grolle ...........................................................371 Lejeunea hieronymii Spruce .........................................................................................................385 Lejeunea hodgsoniana Grolle ex R.J.Lewington, Bever. et M.A.M.Renner ...................................371 Lejeunea howeana Grolle .............................................................................................................371 Lejeunea hui R.L.Zhu .................................................................................................................371 Lejeunea humefacta Spruce ..........................................................................................................371 Lejeunea hyalina (Steph.) L.Söderstr. et A.Hagborg.....................................................................371 Lejeunea hygrophila Gottsche ......................................................................................................383 Lejeunea ibadana A.J.Harr. et E.W.Jones ....................................................................................371 Lejeunea immersa Spruce .............................................................................................................371 Lejeunea increscens Spruce ...........................................................................................................371 Lejeunea inlatiloba (Steph.) H.A.Mill., Bonner et Bischl. ...........................................................371 Lejeunea inlexiloba Prantl ...........................................................................................................372 Lejeunea intricata Prantl..............................................................................................................372 Lejeunea isophylla E.W.Jones .......................................................................................................372 Lejeunea japonica Mitt. ...............................................................................................................372 Lejeunea jardinii Spruce ..............................................................................................................372 Lejeunea julacea Steph. ................................................................................................................372 Lejeunea jungneri (Steph.) Steph. ................................................................................................372 Lejeunea juruana Gradst. et M.E.Reiner .....................................................................................380 Lejeunea kashyapii M.Dey, D.K.Singh et D.Singh ......................................................................372 Lejeunea kilimandjarensis Steph...................................................................................................372 Lejeunea kinabalensis Mizut. .......................................................................................................372 Lejeunea kodamae Ikegami et Inoue ............................................................................................372 Lejeunea konosensis Mizut. ...........................................................................................................372 Lejeunea koponenii (Pócs et J.Eggers) Pócs ..................................................................................381 Lejeunea kuerschneriana Pócs.......................................................................................................372 Lejeunea laeta (Lehm. et Lindenb.) Lehm. et Lindenb. ...............................................................372 Lejeunea laetevirens Nees et Mont. ..............................................................................................380 Lejeunea laevicalyx Gottsche ........................................................................................................383 Lejeunea laii R.L.Zhu .................................................................................................................372 Lejeunea lamacerina (Steph.) Schifn. ..........................................................................................372 Lejeunea lamacerina subsp. gemminata R.M.Schust. ....................................................................372 Lejeunea latilobula (Herzog) R.L.Zhu et M.L.So.........................................................................372 Lejeunea laxa (Nees) Lindenb. ....................................................................................................383 Lejeunea leiantha Spruce .............................................................................................................383 Lejeunea leptalea Spruce ..............................................................................................................372 Lejeunea leptoscypha Spruce .........................................................................................................383 Lejeunea leratii (Steph.) Mizut. ...................................................................................................372 Lejeunea leucosis Besch. et Spruce ................................................................................................373 Lejeunea litoralis Steph. ...............................................................................................................373 Lejeunea lomana E.W.Jones ........................................................................................................373 Lejeunea longicollis S.W.Arnell ....................................................................................................373 586 ** ** *** ** *** ** ** ** ** ** ** ** *** ** ** ** *** ** ** *** *** ** ** ** *** ** *** * ** * ** ** ** * ** ** ** ** ** ** ** * *** ** * ** *** *** ** ** ** ** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Lejeunea longilobula Pócs ............................................................................................................373 Lejeunea lowriana Steph. .............................................................................................................373 Lejeunea lumbricoides (Nees) Nees ..............................................................................................373 Lejeunea lunatigastria Tixier ........................................................................................................373 Lejeunea lusoria (Lindenb. et Gottsche) Steph. ............................................................................383 Lejeunea luzonensis (Steph.) R.L.Zhu et M.J.Lai .........................................................................383 Lejeunea lyratilora Prantl ............................................................................................................373 Lejeunea macrorhyncha Spruce.....................................................................................................383 Lejeunea magohukui Mizut. .........................................................................................................373 Lejeunea malangensis (Herzog) R.L.Zhu et Y.M.Wei ...................................................................373 Lejeunea mandonii (Steph.) Müll.Frib. ........................................................................................373 Lejeunea marasmodes Spruce........................................................................................................383 Lejeunea masoalae Pócs................................................................................................................373 Lejeunea massalongoana (Schifn. ex P.Syd) Solari ........................................................................373 Lejeunea megalantha Spruce ........................................................................................................373 Lejeunea mehrana M.Dey, D.K.Singh et D.Singh .......................................................................373 Lejeunea meridensis Ilk.-Borg.......................................................................................................373 Lejeunea micholitzii Mizut. .........................................................................................................373 Lejeunea microloba Taylor ...........................................................................................................373 Lejeunea mimula Hürl.................................................................................................................373 Lejeunea minutiloba A.Evans .......................................................................................................373 Lejeunea minutiloba var. heterogyna R.M.Schust. .........................................................................373 Lejeunea mizutanii Grolle ...........................................................................................................373 Lejeunea molkenboeriana Sande Lac.............................................................................................373 Lejeunea monimiae (Steph.) Steph. ..............................................................................................374 Lejeunea morobensis (Grolle) M.A.M.Renner et Pócs...................................................................383 Lejeunea multidentata M.E.Reiner et Mustelier ...........................................................................374 Lejeunea musae (Spreng.) Gottsche, Lindenb. et Nees .................................................................374 Lejeunea muscicola Spruce ...........................................................................................................374 Lejeunea muscicola var. palmicola Spruce .....................................................................................374 Lejeunea neelgherriana Gottsche ..................................................................................................374 Lejeunea nemoralis Gottsche ........................................................................................................374 Lejeunea nepalensis Steph. ...........................................................................................................374 Lejeunea nesiotica Besch. et Spruce ..............................................................................................374 Lejeunea neumanniana Nees........................................................................................................374 Lejeunea nietneri (Steph.) Steph. ex Watts...................................................................................374 Lejeunea notata Gottsche ............................................................................................................374 Lejeunea novoguineensis Schifn. ..................................................................................................383 Lejeunea nymannii Steph.............................................................................................................374 Lejeunea obfusca Mitt. .................................................................................................................374 Lejeunea obidensis Spruce ............................................................................................................374 Lejeunea obscura Mitt. .................................................................................................................374 Lejeunea obtusangula Spruce........................................................................................................383 Lejeunea obtusata Gottsche..........................................................................................................374 Lejeunea oerstediana Lindenb. et Hampe .....................................................................................383 Lejeunea okomuensis E.W.Jones ...................................................................................................374 Lejeunea oligoclada Spruce...........................................................................................................374 Lejeunea oracola M.A.M.Renner .................................................................................................374 Lejeunea osculatiana De Not. ......................................................................................................374 Lejeunea otiana S.Hatt. ...............................................................................................................374 Lejeunea ovalifolia Steph. ............................................................................................................375 Lejeunea paciica Mont. ...............................................................................................................375 World checklist of hornworts and liverworts ** ** ** * *** ** ** * ** ** ** ** *** *** *** *** ** *** * * *** * *** ** *** ** ** ** *** * ** * ** ** *** *** *** *** *** ** ** *** ** ** *** ** * ** *** ** *** ** 587 Lejeunea pallescens Mitt. ..............................................................................................................375 Lejeunea pallida Lindenb. et Gottsche .........................................................................................375 Lejeunea pallidevirens S.Hatt. ......................................................................................................380 Lejeunea pallidissima Gola ...........................................................................................................375 Lejeunea papilionacea Prantl ........................................................................................................375 Lejeunea papuana (Pócs) R.M.Schust. .........................................................................................381 Lejeunea paraensis Spruce ............................................................................................................383 Lejeunea paratropa Spruce ...........................................................................................................375 Lejeunea parva (S.Hatt.) Mizut. ..................................................................................................380 Lejeunea parviloba Ångstr............................................................................................................383 Lejeunea patagonica (Steph.) Steph. .............................................................................................375 Lejeunea patens Lindb. ................................................................................................................375 Lejeunea patersonii (Steph.) Steph. ..............................................................................................375 Lejeunea patriciae Schäf.-Verw. ...................................................................................................375 Lejeunea paucidentata (Steph.) Grolle .........................................................................................375 Lejeunea pectinella Mizut. ...........................................................................................................375 Lejeunea perigonialis Gottsche .....................................................................................................375 Lejeunea perpapillosa M.E.Reiner et K.C.Pôrto ...........................................................................375 Lejeunea pertusa (Corda ex Nees et Mont.) Gottsche, Lindenb. et Nees ......................................375 Lejeunea pleidereri Sushil K.Singh ..............................................................................................383 Lejeunea phyllobola Nees et Mont................................................................................................375 Lejeunea phyllobola var. turgidula Spruce .....................................................................................375 Lejeunea planiloba A.Evans .........................................................................................................375 Lejeunea polilloensis Steph. ..........................................................................................................375 Lejeunea polyantha Mont. ...........................................................................................................380 Lejeunea praetervisa Steph. ..........................................................................................................375 Lejeunea primordialis (Hook.f. et Taylor) Gottsche, Lindenb. et Nees.........................................375 Lejeunea princeps (Steph.) Mizut. ................................................................................................376 Lejeunea prionoides Spruce ..........................................................................................................380 Lejeunea proboscidea Gottsche .....................................................................................................385 Lejeunea procumbens Mitt. ..........................................................................................................376 Lejeunea prominula Gottsche ......................................................................................................383 Lejeunea propagulifera Gradst. .....................................................................................................376 Lejeunea pteridis Besch. et Spruce ................................................................................................376 Lejeunea pterigonia (Lehm. et Lindenb.) Mont............................................................................384 Lejeunea ptosimophylla C.Massal. ................................................................................................376 Lejeunea puiggariana Steph. ........................................................................................................376 Lejeunea pulverulenta (Steph.) M.E.Reiner ..................................................................................376 Lejeunea pulvinata Nees et Mont. ...............................................................................................376 Lejeunea quinqueumbonata Spruce ..............................................................................................384 Lejeunea quinqueumbonata var. rotundata (Herzog) Sushil K.Singh ............................................384 Lejeunea raddiana Lindenb. ........................................................................................................376 Lejeunea radicans Lindenb. et Gottsche .......................................................................................376 Lejeunea ramosissima Steph. ........................................................................................................376 Lejeunea ramulosa Spruce ............................................................................................................380 Lejeunea rara Steph. ....................................................................................................................376 Lejeunea ravenelii Austin .............................................................................................................376 Lejeunea recurva M.E.Reiner .......................................................................................................376 Lejeunea relexistipula (Lehm. et Lindenb.) Lehm. et Lindenb. ....................................................376 Lejeunea relexistipula var. costaricensis (Steph.) M.E.Reiner.........................................................376 Lejeunea reinerae Ilk.-Borg. .........................................................................................................376 Lejeunea remotifolia Hampe ex Steph. .........................................................................................384 588 ** ** *** ** *** ** * *** *** ** ** * ** ** ** ** ** ** ** ** ** ** ** ** *** ** *** *** ** ** * ** ** ** * ** ** * * *** ** ** ** ** ** *** ** ** *** ** *** *** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Lejeunea resupinata (Steph.) Steph. .............................................................................................377 Lejeunea reticulata Herzog ..........................................................................................................377 Lejeunea rhigophila M.A.M.Renner .............................................................................................377 Lejeunea rhodesiae (Sim) R.M.Schust. .........................................................................................377 Lejeunea rionegrensis Spruce ........................................................................................................377 Lejeunea riparia Mitt. .................................................................................................................377 Lejeunea rothii (Schwägr.) Gottsche, Lindenb. et Nees ................................................................377 Lejeunea rotundifolia Mitt. ..........................................................................................................377 Lejeunea ruthii (A.Evans) R.M.Schust. ........................................................................................384 Lejeunea ruthii var. alata R.M.Schust. .........................................................................................384 Lejeunea sanctae-helenae M.Wigginton ........................................................................................377 Lejeunea scabrilora Loitl. ............................................................................................................385 Lejeunea schusteri Grolle ..............................................................................................................377 Lejeunea semiscabrida Gottsche ...................................................................................................377 Lejeunea semperi Steph. ...............................................................................................................377 Lejeunea seriata Lindenb. et Gottsche .........................................................................................377 Lejeunea sessililora (Steph.) Grolle ..............................................................................................377 Lejeunea setacea (Steph.) Steph. ...................................................................................................377 Lejeunea setiloba Spruce ..............................................................................................................377 Lejeunea sharpii (R.M.Schust.) R.M.Schust. ...............................................................................377 Lejeunea siccata Spruce ................................................................................................................377 Lejeunea sikorae (Steph.) Steph. ..................................................................................................384 Lejeunea silvatica Gottsche ..........................................................................................................378 Lejeunea smaragdina Besch. et Spruce .........................................................................................378 Lejeunea soae R.L.Zhu, Y.M.Wei, L.Söderstr., A.Hagborg et von Konrat....................................378 Lejeunea solanicola Spruce ...........................................................................................................378 Lejeunea sordida (Nees) Nees ......................................................................................................378 Lejeunea spiniloba Lindenb. et Gottsche ......................................................................................378 Lejeunea spinulilora Spruce.........................................................................................................378 Lejeunea sporadica Besch. et Spruce .............................................................................................378 Lejeunea squarrosa (Steph.) Steph. ...............................................................................................378 Lejeunea squarrosula (Herzog) Solari ...........................................................................................378 Lejeunea srivastavae P.K.Verma et K.K.Rawat .............................................................................384 Lejeunea stenodentata M.A.M.Renner et Pócs..............................................................................384 Lejeunea stephaniana Mizut. ........................................................................................................378 Lejeunea stevensiana (Steph.) Mizut.............................................................................................378 Lejeunea subacuta Mitt. ...............................................................................................................378 Lejeunea subaquatica Schifn. ......................................................................................................378 Lejeunea subbiida Steph. ex Duss ...............................................................................................378 Lejeunea subelobata Carrington et Pearson ..................................................................................384 Lejeunea subigiensis (Steph.) Steph. .............................................................................................378 Lejeunea subolivacea Mizut. .........................................................................................................378 Lejeunea subplana (Steph.) C.J.Bastos .........................................................................................378 Lejeunea subrufula Spruce ...........................................................................................................378 Lejeunea subsessilis Spruce............................................................................................................378 Lejeunea subspathulata Spruce .....................................................................................................379 Lejeunea succulenta Herzog .........................................................................................................379 Lejeunea sufruticola Spruce .........................................................................................................379 Lejeunea sulphurea (Lehm. et Lindenb.) Spruce...........................................................................384 Lejeunea syoshii Inoue..................................................................................................................379 Lejeunea talamancensis M.E.Reiner et Schäf.-Verw......................................................................379 Lejeunea tamasii M.E.Reiner, N.Salazar et C.Chung...................................................................379 World checklist of hornworts and liverworts ** *** ** ** ** ** ** *** *** ** ** *** * ** ** ** ** ** *** ** ** ** ** *** ** ** ** ** * ** *** * *** *** *** ** *** *** *** *** *** *** *** *** *** *** *** *** *** ** *** *** 589 Lejeunea tamaspocsii G.E.Lee ......................................................................................................379 Lejeunea tapajosensis Spruce.........................................................................................................379 Lejeunea tarapotensis Spruce ........................................................................................................379 Lejeunea tenella Taylor ................................................................................................................379 Lejeunea tenera (Sw.) Gottsche, Lindenb. et Nees .......................................................................380 Lejeunea terricola Spruce .............................................................................................................384 Lejeunea thallophora (Eifrig) Gradst. ...........................................................................................379 Lejeunea tonduzana (Steph.) M.E.Reiner ....................................................................................379 Lejeunea topoensis Gradst. et M.E.Reiner.....................................................................................380 Lejeunea touwii (Pócs) R.M.Schust. ............................................................................................381 Lejeunea trachygona Spruce .........................................................................................................379 Lejeunea trinitensis Lindenb. .......................................................................................................379 Lejeunea trochantha Spruce .........................................................................................................385 Lejeunea trukensis H.A.Mill. et Bonner .......................................................................................379 Lejeunea tuberculosa Steph. .........................................................................................................379 Lejeunea tumida Mitt. .................................................................................................................379 Lejeunea tunquiniensis M.E.Reiner et Drehwald ..........................................................................365 Lejeunea uleana (Steph.) Steph. ...................................................................................................379 Lejeunea umbilicata (Nees) Nees .................................................................................................379 Lejeunea urbanii (Steph.) Steph. .................................................................................................384 Lejeunea utriculata (Steph.) Mizut. .............................................................................................379 Lejeunea venezuelana (R.M.Schust.) R.L.Zhu et W.Ye ................................................................384 Lejeunea vesicata Mitt. ................................................................................................................379 Lejeunea villaumei (Steph.) Grolle...............................................................................................379 Lejeunea viridis R.M.Schust. ex L.Söderstr. et A.Hagborg ...........................................................384 Lejeunea vojtkoi Pócs ...................................................................................................................379 Lejeunea vulgariformis Gottsche ..................................................................................................379 Lejeunea wattsiana (Steph.) H.A.Mill., Bonner et Bischl. ............................................................380 Lejeunea wichurae (Steph.) Steph. ...............................................................................................380 Lejeunea wightii Lindenb. ...........................................................................................................380 Lejeunea xiphophylla (Herzog) M.E.Reiner ..................................................................................380 Lejeunea zacuapana (Steph.) Prantl .............................................................................................385 Lembidium berggrenii Herzog......................................................................................................168 Lembidium longifolium R.M.Schust. ...........................................................................................168 Lembidium nutans (Hook.f. et Taylor) Mitt. ...............................................................................168 Lembidium nutans var. lagelliferum E.A.Hodgs. .........................................................................168 Lepicolea attenuata (Mitt.) Steph. ................................................................................................142 Lepicolea magellanica (Gola) Solari ..............................................................................................142 Lepicolea norrisii Piippo ..............................................................................................................142 Lepicolea ochroleuca (Spreng.) Spruce ..........................................................................................142 Lepicolea pruinosa (Taylor) Spruce ..............................................................................................142 Lepicolea ramentiissa Herzog ......................................................................................................142 Lepicolea rara (Steph.) Grolle ......................................................................................................142 Lepicolea rigida (De Not.) E.B.Scott ...........................................................................................142 Lepicolea scolopendra (Hook.) Dumort. ex Trevis. .......................................................................142 Lepicolea yakusimensis (S.Hatt.) S.Hatt........................................................................................142 Lepidogyna hodgsoniae (Grolle) R.M.Schust.................................................................................416 Lepidogyna menziesii (Hook.) R.M.Schust...................................................................................416 Lepidolaena berggrenii E.A.Hodgs. ..............................................................................................416 Lepidolaena brachyclada (Lehm.) Trevis. .....................................................................................416 Lepidolaena clavigera (Hook.) Dumort. ex Trevis. .......................................................................416 Lepidolaena novae-zelandiae (E.A.Hodgs. et S.W.Arnell) von Konrat, L.Söderstr. et A.Hagborg .416 590 *** *** *** ** *** *** *** *** ** *** *** *** *** *** *** *** *** *** *** ** ** ** *** ** ** ** *** ** ** ** ** ** *** ** ** * ** ** ** ** ** ** ** ** ** * ** ** ** *** *** ** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Lepidolaena palpebrifolia (Hook.) Dumort. ex Trevis. .................................................................416 Lepidolaena reticulata (Hook.f. et Taylor) Trevis.........................................................................416 Lepidolaena taylorii (Gottsche) Trevis. ........................................................................................417 Lepidolejeunea auriculata Schäf.-Verw. et Heinrichs ....................................................................391 Lepidolejeunea bidentula (Steph.) R.M.Schust. ............................................................................391 Lepidolejeunea bidentula var. novae-caledoniae Piippo ..................................................................391 Lepidolejeunea borneensis (Steph.) R.M.Schust. ...........................................................................392 Lepidolejeunea cordiissa (Taylor) M.E.Reiner ..............................................................................391 Lepidolejeunea cuspidata (Gottsche) Heinrichs et Schäf.-Verw. ....................................................392 Lepidolejeunea delessertii (Nees et Mont.) Grolle..........................................................................392 Lepidolejeunea eluta (Nees) R.M.Schust. .....................................................................................392 Lepidolejeunea falcata (Herzog) R.M.Schust. ...............................................................................391 Lepidolejeunea graefei (J.B.Jack et Steph.) R.M.Schust................................................................392 Lepidolejeunea grossepapulosa (Steph.) Piippo...............................................................................391 Lepidolejeunea integristipula (J.B.Jack et Steph.) R.M.Schust.......................................................392 Lepidolejeunea involuta (Gottsche) Grolle ...................................................................................391 Lepidolejeunea longilobula (Amakawa) Xiao L.He ........................................................................392 Lepidolejeunea serrulata (Steph.) Grolle .......................................................................................392 Lepidolejeunea sullivantii (Gottsche) M.E.Reiner ........................................................................392 Lepidozia acantha J.J.Engel .........................................................................................................169 Lepidozia aequiloba Steph. ..........................................................................................................170 Lepidozia africana Steph. ............................................................................................................170 Lepidozia alstonii Fulford ............................................................................................................170 Lepidozia ambigua De Not..........................................................................................................170 Lepidozia andicola Beauverd........................................................................................................170 Lepidozia appressifolia Steph. .......................................................................................................170 Lepidozia armata Steph. ..............................................................................................................170 Lepidozia asymmetrica Steph. ......................................................................................................170 Lepidozia auriculata Mitt. ...........................................................................................................170 Lepidozia australis (Lehm. et Lindenb.) Mitt. ..............................................................................170 Lepidozia bidens J.J.Engel............................................................................................................170 Lepidozia biloba Herzog ..............................................................................................................170 Lepidozia bisbiida Steph. ............................................................................................................170 Lepidozia borneensis Steph. ..........................................................................................................170 Lepidozia bragginsiana E.D.Cooper et M.A.M.Renner................................................................170 Lepidozia brasiliensis Steph. .........................................................................................................170 Lepidozia brevidentata Mitt. ........................................................................................................170 Lepidozia brevifolia Mitt. ............................................................................................................170 Lepidozia brevifolia var. planifolia Schifn....................................................................................170 Lepidozia brotheri Steph. .............................................................................................................170 Lepidozia bufalona Steph............................................................................................................170 Lepidozia bursifera S.Hatt. et Grolle............................................................................................170 Lepidozia caespitosa Spruce ..........................................................................................................170 Lepidozia caledonica Steph. .........................................................................................................170 Lepidozia caledonica var. tenuisecta Hürl......................................................................................170 Lepidozia ceramensis Herzog ........................................................................................................170 Lepidozia cherydrion Hürl. ..........................................................................................................170 Lepidozia chiloensis Steph. ...........................................................................................................170 Lepidozia chordulifera Taylor ......................................................................................................171 Lepidozia cladorhiza (Reinw., Blume et Nees) Nees ....................................................................171 Lepidozia coilophylla Taylor.........................................................................................................171 Lepidozia coilophylla var. apiculiloba (Steph.) Fulford ..................................................................171 World checklist of hornworts and liverworts ** *** ** * ** *** ** * * ** ** ** *** *** ** ** ** ** * ** *** ** ** ** ** ** * *** *** ** * ** ** ** *** ** * *** *** ** ** *** ** ** *** ** ** *** ** *** ** ** 591 Lepidozia communis Steph. ..........................................................................................................171 Lepidozia concinna Colenso.........................................................................................................171 Lepidozia cordata Lindenb. .........................................................................................................171 Lepidozia cordistipula Steph. .......................................................................................................171 Lepidozia crassitexta Steph. ..........................................................................................................171 Lepidozia cupressina (Sw.) Lindenb. ............................................................................................171 Lepidozia cupressina subsp. natalensis (Steph.) Pócs .....................................................................171 Lepidozia cupressina subsp. pinnata (Hook.) Pócs ........................................................................171 Lepidozia cupressina subsp. quinqueida (Steph.) Pócs..................................................................171 Lepidozia decaisnei Steph.............................................................................................................171 Lepidozia dendritica Spruce .........................................................................................................171 Lepidozia densa Herzog ...............................................................................................................171 Lepidozia digitata Herzog............................................................................................................171 Lepidozia eenii S.W.Arnell ..........................................................................................................171 Lepidozia elobata R.M.Schust......................................................................................................171 Lepidozia erosa Steph. .................................................................................................................171 Lepidozia erronea Herzog ............................................................................................................171 Lepidozia everettii Steph. .............................................................................................................172 Lepidozia everettii var. javensis Herzog.........................................................................................172 Lepidozia fauriana Steph. ............................................................................................................172 Lepidozia ferdinandi-muelleri Steph.............................................................................................172 Lepidozia ilamentosa (Lehm. et Lindenb.) Lehm. et Lindenb......................................................172 Lepidozia istulosa Mitt. ...............................................................................................................172 Lepidozia lexuosa Mitt. ...............................................................................................................172 Lepidozia fuegiensis Steph. ...........................................................................................................172 Lepidozia fugax J.J.Engel .............................................................................................................172 Lepidozia gedena Steph................................................................................................................172 Lepidozia glaucescens J.J.Engel .....................................................................................................172 Lepidozia glaucophylla (Hook.f. et Taylor) Gottsche, Lindenb. et Nees .......................................172 Lepidozia grandifolia Steph. ........................................................................................................172 Lepidozia griseola Herzog ............................................................................................................172 Lepidozia groenlandica Lehm. ......................................................................................................172 Lepidozia gwamii Piippo .............................................................................................................172 Lepidozia hampeana Lindenb. .....................................................................................................172 Lepidozia haskarliana (Gottsche, Lindenb. et Nees) Steph. .........................................................172 Lepidozia hastatistipula Steph. .....................................................................................................172 Lepidozia hexiloba Pearson ..........................................................................................................176 Lepidozia hirta Steph. .................................................................................................................172 Lepidozia holorhiza (Reinw., Blume et Nees) Nees ......................................................................172 Lepidozia holorhiza var. laxa (Nees) Schifn.................................................................................172 Lepidozia inaequalis (Lehm. et Lindenb.) Lehm. et Lindenb. ......................................................172 Lepidozia incurvata Lindenb. ......................................................................................................172 Lepidozia infuscata Mitt. .............................................................................................................173 Lepidozia integrifolia Doei...........................................................................................................173 Lepidozia jamaicensis Steph. ........................................................................................................173 Lepidozia kashyapii D.Singh et D.K.Singh ..................................................................................173 Lepidozia kinabaluensis Mizut. ....................................................................................................173 Lepidozia kirkii Steph. ................................................................................................................173 Lepidozia lacerifolia Steph. ..........................................................................................................173 Lepidozia laevifolia (Hook.f. et Taylor) Gottsche, Lindenb. et Nees ............................................173 Lepidozia laevifolia var. acutiloba J.J.Engel ..................................................................................173 Lepidozia laevifolia var. alpina R.M.Schust. et J.J.Engel ..............................................................173 592 ** ** ** ** *** * *** ** ** * ** ** * * *** ** ** *** *** ** *** * * * ** *** * ** ** *** ** *** * ** *** * *** *** ** *** ** ** ** * ** ** ** *** ** *** *** ** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Lepidozia lindigiana Steph. .........................................................................................................173 Lepidozia loheri Steph. ................................................................................................................173 Lepidozia longifolia Steph. ...........................................................................................................173 Lepidozia loriana Steph. ..............................................................................................................173 Lepidozia macrocolea Spruce ........................................................................................................173 Lepidozia massartiana Schifn. ex Steph.......................................................................................173 Lepidozia microphylla (Hook.) Lindenb.......................................................................................173 Lepidozia microstipula Steph. ......................................................................................................173 Lepidozia minima Steph. .............................................................................................................173 Lepidozia minor (Gottsche, Lindenb. et Nees) Solari...................................................................173 Lepidozia miqueliana Sande Lac. .................................................................................................173 Lepidozia montana Steph. ...........................................................................................................173 Lepidozia newtonii Steph.............................................................................................................173 Lepidozia nova Steph...................................................................................................................174 Lepidozia novae-zelandiae Steph. .................................................................................................174 Lepidozia novae-zelandiae var. heterostipa R.M.Schust. ................................................................174 Lepidozia novae-zelandiae var. minima R.M.Schust. ....................................................................174 Lepidozia obtusiloba Steph...........................................................................................................174 Lepidozia obtusiloba var. parvula J.J.Engel ...................................................................................174 Lepidozia omeiensis P.C.Chen ex Mizut. et K.C.Chang ...............................................................174 Lepidozia ornata J.J.Engel ...........................................................................................................174 Lepidozia pallida Steph. ..............................................................................................................174 Lepidozia palmicola Steph. ..........................................................................................................174 Lepidozia parvistipa Taylor..........................................................................................................176 Lepidozia paschalis Steph. ............................................................................................................174 Lepidozia patens Lindenb. ...........................................................................................................174 Lepidozia paucifolia Steph. ..........................................................................................................174 Lepidozia paupercula Steph. ........................................................................................................174 Lepidozia pearsonii Spruce ...........................................................................................................174 Lepidozia pendulina (Hook.) Lindenb. ........................................................................................174 Lepidozia peruviensis Steph. .........................................................................................................174 Lepidozia pinnaticruris Spruce ex Steph.......................................................................................174 Lepidozia plumula Herzog ...........................................................................................................174 Lepidozia portoricensis Fulford .....................................................................................................174 Lepidozia procera Mitt. ................................................................................................................174 Lepidozia pseudocupressina Schifn. ..............................................................................................174 Lepidozia pumila J.J.Engel ..........................................................................................................174 Lepidozia quadridens (Nees) Nees ...............................................................................................175 Lepidozia quadriida Lindenb. .....................................................................................................175 Lepidozia reptans (L.) Dumort. ...................................................................................................175 Lepidozia richardsii Herzog .........................................................................................................175 Lepidozia rigida Steph. ................................................................................................................175 Lepidozia robusta Steph. ..............................................................................................................175 Lepidozia rufescens Steph. ............................................................................................................175 Lepidozia sandvicensis Lindenb. ...................................................................................................175 Lepidozia schwabei Herzog ..........................................................................................................175 Lepidozia sellingiana H.A.Mill. ...................................................................................................175 Lepidozia septemida Steph. .........................................................................................................175 Lepidozia serpens Spruce ..............................................................................................................175 Lepidozia serrulata J.J.Engel ........................................................................................................175 Lepidozia setigera Steph. ..............................................................................................................175 Lepidozia sikkimensis Steph. ........................................................................................................175 World checklist of hornworts and liverworts *** * *** ** ** ** ** ** ** ** ** ** ** * ** ** ** *** * ** ** *** *** ** ** ** * *** ** ** *** ** ** * ** * * ** ** ** ** *** * *** *** ** ** * ** ** ** ** 593 Lepidozia spinosissima (Hook.f. et Taylor) Mitt. ..........................................................................175 Lepidozia squamifolia Steph. .......................................................................................................175 Lepidozia squarrosa Steph. ...........................................................................................................175 Lepidozia stahlii Steph.................................................................................................................175 Lepidozia stuhlmannii Steph........................................................................................................175 Lepidozia stuhlmannii var. carnosa (Steph.) Pócs .........................................................................175 Lepidozia stuhlmannii subsp. pulvinata (Steph.) Pócs ..................................................................175 Lepidozia subdichotoma Spruce ....................................................................................................175 Lepidozia subintegra Lindenb. .....................................................................................................175 Lepidozia subtransversa Steph. .....................................................................................................175 Lepidozia subtrichodes Steph. .......................................................................................................175 Lepidozia succida Mitt. ................................................................................................................175 Lepidozia supradecomposita Lindenb. ...........................................................................................175 Lepidozia supradecomposita var. falcifolia Herzog .........................................................................175 Lepidozia suyungii C.Gao et X.L.Bai ...........................................................................................176 Lepidozia terricola Steph..............................................................................................................176 Lepidozia triangulifolia Steph. .....................................................................................................176 Lepidozia trichodes (Reinw., Blume et Nees) Nees .......................................................................176 Lepidozia tricuspidata Steph. .......................................................................................................176 Lepidozia tunguraguae Steph. ......................................................................................................176 Lepidozia ubangiensis Steph. ........................................................................................................176 Lepidozia udarii S.C.Srivast., D.Kumar et D.Sharma ..................................................................176 Lepidozia ulothrix (Schwägr.) Lindenb. .......................................................................................176 Lepidozia vitrea Steph. ................................................................................................................176 Lepidozia wattsiana Steph. ..........................................................................................................176 Lepidozia weymouthiana Steph. ...................................................................................................176 Leptocolea sumatrana Herzog.......................................................................................................510 Leptolejeunea amphiophthalma Zwickel .......................................................................................394 Leptolejeunea apiculata (Horik.) S.Hatt. ......................................................................................394 Leptolejeunea arunachalensis Sudipa Das et D.K.Singh ................................................................394 Leptolejeunea astroidea (Mitt.) Steph. ..........................................................................................394 Leptolejeunea australis Steph. .......................................................................................................394 Leptolejeunea balansae Steph. ......................................................................................................394 Leptolejeunea borneensis Herzog ...................................................................................................394 Leptolejeunea brasiliensis Bischl. ...................................................................................................394 Leptolejeunea convexistipa Bischl. .................................................................................................394 Leptolejeunea curvatifolia Steph. ..................................................................................................394 Leptolejeunea denticulata Steph....................................................................................................394 Leptolejeunea dentistipula Steph. ..................................................................................................394 Leptolejeunea diversilobulata Bischl. .............................................................................................394 Leptolejeunea dolabriformis Pearson .............................................................................................394 Leptolejeunea elliptica (Lehm. et Lindenb.) Besch. .......................................................................395 Leptolejeunea emarginata (Horik.) S.Hatt. ...................................................................................395 Leptolejeunea epiphylla (Mitt.) Steph. ..........................................................................................395 Leptolejeunea exocellata (Spruce) A.Evans ....................................................................................395 Leptolejeunea foliicola Steph. .......................................................................................................395 Leptolejeunea integristipula Steph.................................................................................................395 Leptolejeunea jamaicensis R.M.Schust. .........................................................................................395 Leptolejeunea lancifolia Steph. .....................................................................................................395 Leptolejeunea latifolia Herzog ......................................................................................................395 Leptolejeunea lepinii Steph...........................................................................................................395 Leptolejeunea ligulata Herzog ......................................................................................................395 594 *** * ** ** ** *** *** * *** * * ** ** ** ** ** *** *** * ** ** * *** *** *** *** *** *** *** *** ** *** *** *** ** *** ** * *** ** *** *** *** *** *** *** *** * *** *** *** *** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Leptolejeunea maculata (Mitt.) Schifn.........................................................................................395 Leptolejeunea massartiana Schifn. ex Herzog...............................................................................395 Leptolejeunea micronesica Inoue et H.A.Mill. ...............................................................................395 Leptolejeunea minima Herzog ......................................................................................................395 Leptolejeunea mirikana M.Dey et D.K.Singh ..............................................................................395 Leptolejeunea moniliata Steph......................................................................................................395 Leptolejeunea obfuscata (Spruce) Steph. .......................................................................................395 Leptolejeunea punctata Herzog.....................................................................................................395 Leptolejeunea radicosa (Nees ex Mont.) Grolle .............................................................................396 Leptolejeunea renneri Herzog .......................................................................................................396 Leptolejeunea revoluta P.C.Chen .................................................................................................396 Leptolejeunea rosulans Steph. .......................................................................................................396 Leptolejeunea serratifolia Schifn. .................................................................................................396 Leptolejeunea serrulata Herzog .....................................................................................................396 Leptolejeunea spinistipula (Mizut.) Xiao L.He ..............................................................................396 Leptolejeunea subdentata Schifn. ex Herzog ................................................................................396 Leptolejeunea subrotundifolia Herzog ...........................................................................................396 Leptolejeunea tridentata Bischl. ....................................................................................................396 Leptolejeunea trigonostipa (Spruce) Steph. ....................................................................................396 Leptolejeunea tripuncta (Mitt.) Steph. ..........................................................................................396 Leptolejeunea truncatifolia Steph. .................................................................................................396 Leptolejeunea udarii M.Dey et D.K.Singh ...................................................................................396 Leptolejeunea vitrea (Nees) Schifn. .............................................................................................396 Leptophyllopsis laxa (Mitt.) R.M.Schust. ex Hamlin ....................................................................204 Leptoscyphopsis paradoxa R.M.Schust...........................................................................................204 Leptoscyphus aequatus (Hook.f. et Taylor) Mitt. ..........................................................................206 Leptoscyphus amphibolius (Nees) Grolle .......................................................................................207 Leptoscyphus antarcticus (C.Massal.) Solari...................................................................................205 Leptoscyphus australis (Gottsche, Lindenb. et Nees) R.M.Schust. .................................................205 Leptoscyphus autoicus (J.J.Engel et Gradst.) Vanderp. et Gradst. ..................................................207 Leptoscyphus beckettianus (Steph.) R.M.Schust. ex J.J.Engel.........................................................207 Leptoscyphus belmoranus (Steph.) J.J.Engel ..................................................................................205 Leptoscyphus chilensis (De Not.) Hässel ........................................................................................206 Leptoscyphus cleeii Fulford ..........................................................................................................207 Leptoscyphus compactus (Colenso) J.J.Engel..................................................................................207 Leptoscyphus cuneifolius (Hook.) Mitt. .........................................................................................204 Leptoscyphus cuneifolius subsp. fragilis (J.B.Jack et Steph.) Grolle.................................................205 Leptoscyphus diicilis (Steph.) Fulford..........................................................................................206 Leptoscyphus diversifolius (Gottsche) Grolle..................................................................................206 Leptoscyphus erraticus (W.Martin et E.A.Hodgs.) J.J.Engel ..........................................................208 Leptoscyphus excipulatus (Steph.) J.J.Engel ...................................................................................205 Leptoscyphus expansus (Lehm.) Grolle ..........................................................................................206 Leptoscyphus gibbosus (Taylor) Mitt. ............................................................................................205 Leptoscyphus gradsteinii Vanderp., Schäf.-Verw. et D.G.Long......................................................205 Leptoscyphus hedbergii (S.W.Arnell) R.M.Schust. ........................................................................207 Leptoscyphus hexagonus (Nees) Grolle ..........................................................................................205 Leptoscyphus horizontalis (Hook.) Kühnem. .................................................................................208 Leptoscyphus huidobroanus (Mont.) Gottsche ...............................................................................207 Leptoscyphus huonicus Piippo .......................................................................................................207 Leptoscyphus infuscatus (Mitt.) E.W.Jones ex Grolle ....................................................................207 Leptoscyphus innovatus (E.A.Hodgs.) J.J.Engel.............................................................................205 Leptoscyphus intermedius Grolle ...................................................................................................206 World checklist of hornworts and liverworts *** *** *** *** ** *** *** ** *** *** *** *** *** ** *** ** *** ** *** ** *** *** * *** *** *** *** ** ** *** *** ** ** ** ** *** *** *** *** * *** ** ** ** *** ** ** *** ** ** ** 595 Leptoscyphus jackii (Steph.) Grolle ...............................................................................................205 Leptoscyphus lambinonii Vanderp., Schäf.-Verw. et D.G.Long ....................................................206 Leptoscyphus longistipulus (Steph.) J.J.Engel .................................................................................205 Leptoscyphus magellanicus (Gola) Hässel ......................................................................................207 Leptoscyphus normalis (Steph.) J.J.Engel ......................................................................................208 Leptoscyphus obcordatus (Spruce) Grolle .......................................................................................206 Leptoscyphus ovatus (Spruce) Grolle .............................................................................................206 Leptoscyphus physanthus (Hook.f. et Taylor) J.J.Engel ..................................................................208 Leptoscyphus physocalyx (Hampe et Gottsche) Gottsche ...............................................................205 Leptoscyphus porphyrius (Nees) Grolle ..........................................................................................206 Leptoscyphus porphyrius subsp. azoricus (H.Buch et Perss.) Vanderp. et Heinrichs .......................206 Leptoscyphus sotiauxii Vanderp., Schäf.-Verw. et D.G.Long ........................................................206 Leptoscyphus spectabilis (Steph.) Grolle ........................................................................................207 Leptoscyphus subemarginatus (Hook.f. et Taylor) J.J.Engel ...........................................................208 Leptoscyphus trapezoïdes (Mont.) L.Söderstr. ................................................................................208 Lethocolea congesta (Lehm.) S.W.Arnell .........................................................................................97 Lethocolea glossophylla (Spruce) Grolle ...........................................................................................97 Lethocolea indica G.Asthana et Maurya .........................................................................................97 Lethocolea javanica (Schifn.) Grolle ..............................................................................................97 Lethocolea naruto-toganensis Furuki ...............................................................................................97 Lethocolea pansa (Taylor) G.A.M.Scott et K.G.Beckm. .................................................................97 Lethocolea radicosa (Lehm. et Lindenb.) Grolle .............................................................................97 Lethocolea repens S.Winkl. .............................................................................................................97 Lindigianthus cipaconeus (Gottsche) Kruijt et Gradst...................................................................299 Liochlaena lanceolata Nees...........................................................................................................117 Liochlaena subulata (A.Evans) Schljakov .....................................................................................118 Lobatiriccardia alterniloba (Hook.f. et Taylor) Furuki .................................................................437 Lobatiriccardia alterniloba var. gigantea (Steph.) Nebel................................................................437 Lobatiriccardia alterniloba var. robusta (Rodway) Nebel ..............................................................437 Lobatiriccardia athertonensis (Hewson) Furuki ............................................................................438 Lobatiriccardia coronopus (De Not.) Furuki .................................................................................438 Lobatiriccardia coronopus subsp. australis (R.M.Schust.) Nebel, Preussing, Schäf.-Verw. et D.Quandt...................................................................................................................................438 Lobatiriccardia oberwinkleri Nebel, Preussing, Schäf.-Verw. et D.Quandt...................................438 Lobatiriccardia subaquatica (R.M.Schust.) Nebel ........................................................................438 Lobatiriccardia verdoornioides Nebel, Preussing, Schäf.-Verw. et D.Quandt ................................438 Lobatiriccardia yakusimensis (S.Hatt.) Furuki ..............................................................................438 Lobatiriccardia yunnanensis Furuki et D.G.Long .........................................................................438 Lophocolea aberrans Lindenb. et Gottsche ...................................................................................208 Lophocolea aequifolia Nees et Mont. ............................................................................................208 Lophocolea angustistipula Steph. ..................................................................................................208 Lophocolea anisoloba (J.J.Engel et Glenny) L.Söderstr..................................................................208 Lophocolea anomala Steph. ..........................................................................................................208 Lophocolea anomoda (Mont.) Steph. ............................................................................................208 Lophocolea apalachicola R.M.Schust. ...........................................................................................208 Lophocolea aperticaulis (J.J.Engel) L.Söderstr. ..............................................................................208 Lophocolea aphelophylla (Hässel) Váňa .........................................................................................208 Lophocolea apophylla (Hässel) Váňa .............................................................................................209 Lophocolea appalachiana R.M.Schust...........................................................................................209 Lophocolea ascensionis Steph. .......................................................................................................209 Lophocolea asperrima Steph. ........................................................................................................209 Lophocolea atra Gola ...................................................................................................................209 596 *** *** ** ** ** ** ** *** ** *** ** ** ** *** ** *** ** *** *** ** ** * ** ** ** *** * ** ** ** * *** *** *** ** *** ** ** *** *** ** ** ** ** ** ** *** ** ** ** *** ** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Lophocolea attenuata Steph. .........................................................................................................209 Lophocolea australis Gottsche.......................................................................................................209 Lophocolea autoica Steph. ............................................................................................................209 Lophocolea baldwinii Steph. ........................................................................................................209 Lophocolea bartlettii H.A.Mill. ....................................................................................................209 Lophocolea bewsii (Sim) Grolle ....................................................................................................209 Lophocolea bicuspidata Steph. ......................................................................................................209 Lophocolea bidentata (L.) Dumort. ..............................................................................................209 Lophocolea biidistipula Steph. .....................................................................................................209 Lophocolea bispinosa (Hook.f. et Taylor) Gottsche, Lindenb. et Nees ..........................................209 Lophocolea bootanensis Steph. ......................................................................................................209 Lophocolea boulyana Steph. .........................................................................................................209 Lophocolea bowiena Steph. ..........................................................................................................209 Lophocolea brookwoodiana Paton et Sheahan ...............................................................................209 Lophocolea caespitans Steph. ........................................................................................................209 Lophocolea calcarea Steph. ...........................................................................................................209 Lophocolea caledonica Steph.........................................................................................................209 Lophocolea canaliculata (Gottsche, Lindenb. et Nees) Steph. .......................................................210 Lophocolea canaliculata var. concava (J.J.Engel) L.Söderstr. .........................................................210 Lophocolea cervicornis Steph. .......................................................................................................210 Lophocolea ciliifera Steph.............................................................................................................210 Lophocolea coadunata (Sw.) Mont. ..............................................................................................210 Lophocolea concreta Mont. ...........................................................................................................210 Lophocolea convexula Mitt. ..........................................................................................................210 Lophocolea corticola Steph............................................................................................................210 Lophocolea decurrens Herzog........................................................................................................210 Lophocolea deningeri Herzog........................................................................................................210 Lophocolea dentilora Steph..........................................................................................................210 Lophocolea diformis Nees ............................................................................................................210 Lophocolea discedens (Lehm. et Lindenb.) Lehm. et Lindenb. ......................................................210 Lophocolea dusenii Steph. ............................................................................................................210 Lophocolea erosa (J.J.Engel) L.Söderstr.........................................................................................210 Lophocolea excisifolia Steph..........................................................................................................210 Lophocolea fertilis (J.J.Engel) L.Söderstr.......................................................................................210 Lophocolea lavicans Steph. ..........................................................................................................210 Lophocolea loribunda Steph. .......................................................................................................210 Lophocolea foliicola Spruce...........................................................................................................210 Lophocolea fragillima Steph. ........................................................................................................210 Lophocolea fragmentissima R.M.Schust. .......................................................................................211 Lophocolea fragrans (Moris et De Not.) Gottsche, Lindenb. et Nees ............................................211 Lophocolea glaziovii Steph. ..........................................................................................................211 Lophocolea gollanii (Steph.) Váňa ................................................................................................211 Lophocolea granatensis Gottsche...................................................................................................211 Lophocolea griithiana Steph. ......................................................................................................211 Lophocolea hahnii Steph. .............................................................................................................211 Lophocolea haskarliana Gottsche..................................................................................................211 Lophocolea hattorii (J.J.Engel) L.Söderstr.....................................................................................211 Lophocolea hawaica Steph............................................................................................................211 Lophocolea heterodonta Steph.......................................................................................................211 Lophocolea heteromorpha Steph. ...................................................................................................211 Lophocolea heterophylla (Schrad.) Dumort. ..................................................................................211 Lophocolea heterophylla subsp. cladogyna R.M.Schust...................................................................211 World checklist of hornworts and liverworts ** ** * ** ** ** ** ** * ** ** *** * ** *** *** *** ** *** ** ** ** *** ** * *** ** * * ** *** * ** *** *** ** ** ** ** *** ** *** *** ** ** * ** ** ** ** ** ** 597 Lophocolea horikawana S.Hatt. ....................................................................................................211 Lophocolea howeana Steph. ..........................................................................................................211 Lophocolea humifusa (Hook.f. et Taylor) Gottsche, Lindenb. et Nees ..........................................211 Lophocolea humistrata (Hook.f. et Taylor) Gottsche, Lindenb. et Nees .......................................211 Lophocolea itoana Inoue ..............................................................................................................211 Lophocolea javanica Schifn. ........................................................................................................211 Lophocolea koponenii (Piippo) Váňa ............................................................................................211 Lophocolea kurzii Sande Lac. .......................................................................................................212 Lophocolea kurzii var. siamensis N.Kitag. .....................................................................................212 Lophocolea laceristipula Steph. .....................................................................................................212 Lophocolea latistipula Steph. ........................................................................................................212 Lophocolea lauterbachii Steph. .....................................................................................................212 Lophocolea laxissima Herzog ........................................................................................................212 Lophocolea ledermannii Steph. .....................................................................................................212 Lophocolea lenta (Hook.f. et Taylor) Gottsche, Lindenb. et Nees ................................................212 Lophocolea leptantha (Hook.f. et Taylor) Gottsche, Lindenb. et Nees..........................................212 Lophocolea liebmanniana Gottsche ..............................................................................................212 Lophocolea lindmannii Steph. ......................................................................................................212 Lophocolea longiciliata Herzog .....................................................................................................212 Lophocolea lucida (Spreng.) Mont. ..............................................................................................212 Lophocolea madagascariensis Gottsche ..........................................................................................212 Lophocolea magna (Udar et V.Nath) Váňa...................................................................................212 Lophocolea mediinfrons (J.J.Engel et Braggins) L.Söderstr. ...........................................................212 Lophocolea micronesica Inoue et H.A.Mill. ..................................................................................212 Lophocolea microstipula Steph. .....................................................................................................212 Lophocolea minor Nees ................................................................................................................212 Lophocolea minutistipula Steph. ...................................................................................................212 Lophocolea mollis (Nees) Nees .....................................................................................................212 Lophocolea morobeana Piippo ......................................................................................................213 Lophocolea muhavurensis (S.W.Arnell) S.W.Arnell ex Pócs ..........................................................213 Lophocolea muricata (Lehm.) Nees ..............................................................................................213 Lophocolea muricata var. major Pearson .......................................................................................213 Lophocolea nakajimae S.Hatt. et Inoue ........................................................................................213 Lophocolea novae-zeelandiae (Lehm. et Lindenb.) Nees................................................................213 Lophocolea novae-zeelandiae var. meridionalis (Steph.) L.Söderstr. ...............................................213 Lophocolea orbigniana Nees et Mont. ..........................................................................................213 Lophocolea papulimarginata H.A.Mill..........................................................................................213 Lophocolea parca (Gottsche) Fulford et Sharp ..............................................................................213 Lophocolea parva Steph. ..............................................................................................................213 Lophocolea parvispinea (J.J.Engel) L.Söderstr. ..............................................................................213 Lophocolea parvistipula Steph. .....................................................................................................213 Lophocolea patulistipa Steph. .......................................................................................................213 Lophocolea perpusilla (Hook.f. et Taylor) Gottsche, Lindenb. et Nees .........................................213 Lophocolea piacenzai (Gola) Váňa ................................................................................................213 Lophocolea pilistipula Steph. ........................................................................................................213 Lophocolea pinnatistipula Steph. ..................................................................................................213 Lophocolea platensis C.Massal. .....................................................................................................214 Lophocolea purpurea Steph. .........................................................................................................214 Lophocolea pusilla Steph. .............................................................................................................214 Lophocolea randii S.W.Arnell ......................................................................................................214 Lophocolea rara Steph..................................................................................................................214 Lophocolea rectangularis Herzog...................................................................................................214 598 ** *** *** ** ** *** *** ** ** ** * ** *** ** ** *** ** ** ** *** ** ** ** *** ** *** ** *** * *** ** *** ** ** * *** ** *** *** *** *** *** *** ** *** *** *** *** ** ** *** *** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Lophocolea rectangulata Mitt. ......................................................................................................214 Lophocolea rupicola Steph. ...........................................................................................................214 Lophocolea sabuletorum (Hook.f. et Taylor) Gottsche, Lindenb. et Nees .....................................214 Lophocolea salacensis Steph. .........................................................................................................214 Lophocolea savesiana Steph. .........................................................................................................214 Lophocolea semiteres (Lehm.) Mitt. ..............................................................................................214 Lophocolea semiteres var. retusa (J.J.Engel) L.Söderstr...................................................................214 Lophocolea serrata Mitt. ...............................................................................................................214 Lophocolea siamensis Steph. .........................................................................................................214 Lophocolea sikkimensis (Steph.) Herzog et Grolle .........................................................................214 Lophocolea silvestris Gottsche .......................................................................................................214 Lophocolea steetziae De Not. ........................................................................................................214 Lophocolea striatella (C.Massal.) Schifn. .....................................................................................214 Lophocolea subbidentata Herzog ..................................................................................................214 Lophocolea subcostata Steph. ........................................................................................................214 Lophocolea subporosa Mitt. ..........................................................................................................214 Lophocolea subporosa var. inlexifolia (Steph.) L.Söderstr. .............................................................215 Lophocolea subviridis (Hook.f. et Taylor) Gottsche, Lindenb. et Nees .........................................215 Lophocolea sumatrana Schifn. .....................................................................................................215 Lophocolea sylvatica Mitt. ............................................................................................................215 Lophocolea tenera Ångstr. ............................................................................................................215 Lophocolea tenerrima Spruce ........................................................................................................215 Lophocolea teptepensis Piippo .......................................................................................................215 Lophocolea textilis (Hook.f. et Taylor) Gottsche, Lindenb. et Nees ..............................................215 Lophocolea textiloidea J.J.Engel ....................................................................................................215 Lophocolea trichocoleoides (Glenny, J.J.Engel et He-Nygrén) L.Söderstr. ......................................215 Lophocolea tricuspidata Herzog ....................................................................................................215 Lophocolea tristaniana S.W.Arnell ...............................................................................................215 Lophocolea undulata Mont. .........................................................................................................215 Lophocolea villosa Mitt. ...............................................................................................................215 Lophocolea wacei (S.W.Arnell ex J.J.Engel et Váňa) Váňa et L.Söderstr. ......................................215 Lophocolea wambana Piippo ........................................................................................................215 Lophocolea werthii (J.J.Engel et R.M.Schust.) Váňa et L.Söderstr. ...............................................215 Lophocolea widgrenii Steph. .........................................................................................................215 Lopholejeunea aberrantia Horik. ..................................................................................................408 Lopholejeunea applanata (Reinw., Blume et Nees) Schifn. ..........................................................406 Lopholejeunea borbonica Steph. ...................................................................................................406 Lopholejeunea borneensis (Steph.) Verd. .......................................................................................406 Lopholejeunea ceylanica Steph. .....................................................................................................407 Lopholejeunea colensoi Steph. .......................................................................................................407 Lopholejeunea erugata B.M.hiers ...............................................................................................406 Lopholejeunea eulopha (Taylor) Schifn. .......................................................................................406 Lopholejeunea evansiana Verd. .....................................................................................................406 Lopholejeunea grandicrista Steph..................................................................................................408 Lopholejeunea grollei R.L.Zhu et Gradst. .....................................................................................406 Lopholejeunea herzogiana Verd. ...................................................................................................406 Lopholejeunea hispidissima Steph. ................................................................................................407 Lopholejeunea horticola Schifn. ...................................................................................................407 Lopholejeunea jonesii Vanden Berghen.........................................................................................406 Lopholejeunea laciniata E.W.Jones ..............................................................................................406 Lopholejeunea latilobula Verd. .....................................................................................................407 Lopholejeunea leioptera Gyarmati .................................................................................................408 World checklist of hornworts and liverworts ** *** *** ** *** * *** * ** * *** *** * * *** ** ** *** ** *** *** ** * ** ** * ** *** * *** *** *** *** *** * ** *** * ** ** ** ** * ** * * *** *** * ** *** * 599 Lopholejeunea lepidoscypha Kiaer et Pearson .................................................................................408 Lopholejeunea loheri Steph. ..........................................................................................................407 Lopholejeunea magna Mizut. .......................................................................................................407 Lopholejeunea minima Vanden Berghen ......................................................................................406 Lopholejeunea minuta R.L.Zhu et Gradst. ...................................................................................407 Lopholejeunea multilacera Steph. .................................................................................................408 Lopholejeunea nigricans (Lindenb.) Schifn. .................................................................................407 Lopholejeunea obtusilacera Herzog ...............................................................................................406 Lopholejeunea onraedtii Vanden Berghen.....................................................................................408 Lopholejeunea paramultilacera Vanden Berghen ..........................................................................406 Lopholejeunea plicatiscypha (Hook.f. et Taylor) Steph..................................................................407 Lopholejeunea pocsii Gyarmati .....................................................................................................408 Lopholejeunea proxima Steph. ......................................................................................................407 Lopholejeunea quinquecarinata Vanden Berghen .........................................................................406 Lopholejeunea recurvata Mizut.....................................................................................................407 Lopholejeunea renistipula (Mitt.) Steph. .......................................................................................406 Lopholejeunea revoluta E.W.Jones ...............................................................................................406 Lopholejeunea soae R.L.Zhu et Gradst. ........................................................................................407 Lopholejeunea sphaerophora (Lehm. et Lindenb.) Steph. ..............................................................408 Lopholejeunea streimannii B.M.hiers et Gradst. .........................................................................407 Lopholejeunea subfusca (Nees) Schifn. .........................................................................................407 Lopholejeunea subfusca var. elongata Vanden Berghen ..................................................................407 Lopholejeunea tixieriana Vanden Berghen ...................................................................................408 Lopholejeunea udarii M.Dey et D.K.Singh ..................................................................................408 Lopholejeunea utriculata Steph. ...................................................................................................408 Lopholejeunea vietnamica Tixier ..................................................................................................408 Lopholejeunea vojtkoana Gyarmati ...............................................................................................407 Lopholejeunea wiltensii Steph.......................................................................................................407 Lopholejeunea yapensis Steph. ......................................................................................................407 Lopholejeunea zollingeri (Steph.) Schifn. .....................................................................................407 Lophonardia jamesonii (Mont.) L.Söderstr. et Váňa .......................................................................75 Lophonardia laxifolia (Mont.) L.Söderstr. et Váňa.........................................................................75 Lophonardia tristaniana (S.W.Arnell) L.Söderstr. et Váňa .............................................................75 Lophozia ascendens (Warnst.) R.M.Schust. ....................................................................................79 Lophozia austrosibirica Bakalin ......................................................................................................79 Lophozia ciliata Damsh., L.Söderstr. et H.Weibull .......................................................................79 Lophozia guttulata (Lindb. et Arnell) A.Evans ...............................................................................79 Lophozia jamaicensis (Nees) Steph. ................................................................................................79 Lophozia lacerata N.Kitag. ............................................................................................................79 Lophozia lantratoviae Bakalin ........................................................................................................79 Lophozia murmanica Kaal. ............................................................................................................79 Lophozia paciica Bakalin ..............................................................................................................79 Lophozia pallida (Steph.) Grolle ....................................................................................................79 Lophozia savicziae Schljakov..........................................................................................................79 Lophozia schusterana Schljakov ......................................................................................................79 Lophozia serpens (Dumort.) Dumort. ............................................................................................80 Lophozia silvicola H.Buch .............................................................................................................79 Lophozia silvicoloides N.Kitag. .......................................................................................................79 Lophozia subapiculata R.M.Schust. et Damsh. ..............................................................................80 Lophozia udarii S.Srivast., S.C.Srivast. et K.K.Rawat ....................................................................80 Lophozia ventricosa (Dicks.) Dumort.............................................................................................80 Lophozia wenzelii (Nees) Steph. ....................................................................................................80 600 *** ** ** ** *** ** *** ** *** ** * ** *** * *** *** *** *** *** *** *** *** ** *** * * * *** *** *** *** *** * * * *** *** * *** * *** *** *** *** *** *** *** *** * *** *** *** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Lophoziopsis excisa (Dicks.) Konstant. et Vilnet .............................................................................80 Lophoziopsis excisa var. elegans (R.M.Schust.) Konstant. et Vilnet ..................................................80 Lophoziopsis excisa var. infuscata (R.M.Schust. et Damsh.) Konstant. et Vilnet..............................80 Lophoziopsis excisa var. succulenta (R.M.Schust. et Damsh.) Konstant. et Vilnet ............................80 Lophoziopsis longidens (Lindb.) Konstant. et Vilnet .......................................................................81 Lophoziopsis longidens subsp. arctica (R.M.Schust.) Váňa et L.Söderstr..........................................81 Lophoziopsis pellucida (R.M.Schust.) Konstant. et Vilnet...............................................................81 Lophoziopsis pellucida var. minor (R.M.Schust.) L.Söderstr. et Váňa ..............................................81 Lophoziopsis polaris (R.M.Schust.) Konstant. et Vilnet ..................................................................81 Lophoziopsis polaris var. sphagnorum (R.M.Schust.) Konstant. et Vilnet.........................................81 Lophoziopsis propagulifera (Gottsche) Konstant. et Vilnet ..............................................................81 Lophoziopsis rubrigemma (R.M.Schust.) Konstant. et Vilnet ..........................................................81 Lunularia cruciata (L.) Dumort. ex Lindb. ..................................................................................476 Lunularia cruciata subsp. thaxteri (A.Evans et Herzog) R.M.Schust. ...........................................476 Luteolejeunea herzogii (Buchloh) Piippo ......................................................................................304 Macrocolura sagittistipula (Spruce) R.M.Schust. ..........................................................................348 Makinoa crispata (Steph.) Miyake ...............................................................................................468 Mannia androgyna (L.) A.Evans ..................................................................................................481 Mannia californica (Gottsche) L.C.Wheeler ................................................................................481 Mannia controversa (Meyl.) D.B.Schill ........................................................................................481 Mannia controversa subsp. asiatica D.B.Schill et D.G.Long.........................................................481 Mannia fragrans (Balb.) Frye et L.Clark ......................................................................................481 Mannia fragrans subsp. orientalis R.M.Schust. ............................................................................481 Mannia gracilis (F.Weber) D.B.Schill et D.G.Long ....................................................................482 Mannia hegewaldii Bischl. ...........................................................................................................482 Mannia paradoxa R.M.Schust. ....................................................................................................482 Mannia perssonii Udar et V.Chandra ..........................................................................................481 Mannia pilosa (Hornem.) Frye et L.Clark ...................................................................................482 Mannia sibirica (Müll.Frib.) Frye et L.Clark ...............................................................................482 Mannia triandra (Scop.) Grolle ..................................................................................................482 Marchantia acaulis Steph. ...........................................................................................................490 Marchantia antiqua Steph. ..........................................................................................................490 Marchantia assamica Grif. ..........................................................................................................491 Marchantia balboi Gola...............................................................................................................491 Marchantia balboi var. acutisquamata Gerola ..............................................................................491 Marchantia berteroana Lehm. et Lindenb. ...................................................................................490 Marchantia breviloba A.Evans .....................................................................................................489 Marchantia cagnii Gola ...............................................................................................................491 Marchantia carrii Bischl. .............................................................................................................490 Marchantia cengiana Gerola ........................................................................................................491 Marchantia chenopoda L. .............................................................................................................489 Marchantia crenata Austin ..........................................................................................................489 Marchantia debilis K.I.Goebel .....................................................................................................490 Marchantia emarginata Reinw., Blume et Nees ...........................................................................490 Marchantia emarginata subsp. lecordiana (Steph.) Bischl. ............................................................490 Marchantia emarginata subsp. tosana (Steph.) Bischl. ..................................................................490 Marchantia foliacea Mitt. ............................................................................................................489 Marchantia formosana Horik.......................................................................................................489 Marchantia friedrichsthaliana Trevis............................................................................................491 Marchantia geminata Reinw., Blume et Nees ..............................................................................490 Marchantia globosa Brid. .............................................................................................................489 Marchantia hartlessiana Steph. ....................................................................................................491 World checklist of hornworts and liverworts *** *** * *** *** *** *** *** *** *** *** *** ** * *** *** *** *** *** *** *** * *** * *** ** *** *** *** *** * * *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 601 Marchantia hexaptera Reichardt ..................................................................................................489 Marchantia inlexa Nees et Mont. ...............................................................................................489 Marchantia keniae Gola ..............................................................................................................491 Marchantia linearis Lehm. et Lindenb.........................................................................................489 Marchantia macropora Mitt.........................................................................................................491 Marchantia miqueliana Lehm. ....................................................................................................489 Marchantia novoguineensis Bischl. ...............................................................................................489 Marchantia paleacea Bertol..........................................................................................................489 Marchantia paleacea subsp. diptera (Nees et Mont.) Inoue ..........................................................489 Marchantia papillata Raddi .........................................................................................................490 Marchantia papillata subsp. grossibarba (Steph.) Bischl. ..............................................................490 Marchantia pappeana Lehm. .......................................................................................................489 Marchantia pappeana subsp. robusta (Steph.) Bischl. ...................................................................489 Marchantia papyracea Gola .........................................................................................................491 Marchantia philippinensis Bischl. .................................................................................................491 Marchantia pileata Mitt. .............................................................................................................489 Marchantia pinnata Steph. ..........................................................................................................489 Marchantia plicata Nees et Mont. ...............................................................................................490 Marchantia polymorpha L. ...........................................................................................................490 Marchantia polymorpha subsp. montivagans Bischl. et Boissel.-Dub.............................................490 Marchantia polymorpha subsp. ruderalis Bischl. et Boissel.-Dub. .................................................490 Marchantia quadriloba Steph. .....................................................................................................492 Marchantia rubribarba Steph. .....................................................................................................489 Marchantia sellae Gola ................................................................................................................492 Marchantia solomonensis Bischl. ..................................................................................................491 Marchantia stoloniscyphulus (C.Gao et K.C.Chang) Piippo .........................................................492 Marchantia streimannii Bischl. ....................................................................................................491 Marchantia subgeminata Steph. ...................................................................................................491 Marchantia subintegra Mitt. ........................................................................................................491 Marchantia treubii Schifn. .........................................................................................................491 Marchantia trilocularis Roth........................................................................................................492 Marchantia tusui Gola ................................................................................................................492 Marchantia vitiensis Steph. ..........................................................................................................490 Marchantia wallisii J.B.Jack et Steph...........................................................................................491 Marchesinia bongardiana (Lehm. et Lindenb.) Trevis. .................................................................409 Marchesinia brachiata (Sw.) Schifn. ...........................................................................................409 Marchesinia deslooveri Vanden Berghen.......................................................................................409 Marchesinia excavata (Mitt.) Schifn. ..........................................................................................409 Marchesinia languida (Nees et Mont.) Steph. ..............................................................................409 Marchesinia mackaii (Hook.) Gray..............................................................................................408 Marchesinia nobilis (Gottsche) X.Q.Shi, R.L.Zhu et Gradst. .......................................................409 Marchesinia robusta (Mitt.) Schifn. ............................................................................................409 Marsupella alata S.Hatt. et N.Kitag. ...........................................................................................112 Marsupella andreaeoides (Lindb.) Müll.Frib.................................................................................112 Marsupella apiculata Schifn. .......................................................................................................112 Marsupella aquatica (Lindenb.) Schifn. ......................................................................................112 Marsupella arctica (Berggr.) Bryhn et Kaal. .................................................................................112 Marsupella boeckii (Austin) Lindb. ex Kaal. .................................................................................112 Marsupella bolanderi (Austin) Underw. .......................................................................................112 Marsupella condensata (Ångstr. ex C.Hartm.) Lindb. ex Kaal. .....................................................113 Marsupella disticha Steph. ...........................................................................................................113 Marsupella emarginata (Ehrh.) Dumort. .....................................................................................113 602 ** ** ** ** *** *** ** *** *** *** *** *** ** *** *** *** ** *** ** *** * * * * * * * * * * * * * * * * * * * * * *** ** *** *** *** ** *** *** *** *** *** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Marsupella emarginata subsp. tubulosa (Steph.) N.Kitag..............................................................113 Marsupella emarginata subsp. tubulosa var. apertifolia (Steph.) N.Kitag. ......................................113 Marsupella emarginata subsp. tubulosa var. patens N.Kitag. .........................................................113 Marsupella emarginata subsp. tubulosa var. tubulosa (Steph.) N.Kitag. ex Váňa et L.Söderstr. .....113 Marsupella funckii (F.Weber et D.Mohr) Dumort. .....................................................................113 Marsupella microphylla R.M.Schust. ............................................................................................113 Marsupella minutissima N.Kitag. .................................................................................................113 Marsupella neesii Sande Lac. ex Schifn. ......................................................................................113 Marsupella paroica R.M.Schust. ..................................................................................................113 Marsupella profunda Lindb..........................................................................................................113 Marsupella pseudofunckii S.Hatt. .................................................................................................113 Marsupella sparsifolia (Lindb.) Dumort. ......................................................................................113 Marsupella sparsifolia subsp. childii R.M.Schust. .........................................................................113 Marsupella sphacelata (Giesecke ex Lindenb.) Dumort. ...............................................................113 Marsupella spiniloba R.M.Schust. et Damsh................................................................................113 Marsupella sprucei (Limpr.) Bernet ..............................................................................................114 Marsupella stableri Spruce ...........................................................................................................114 Marsupella stoloniformis N.Kitag. ................................................................................................114 Marsupella stoloniformis subsp. vermiformis R.M.Schust. .............................................................114 Marsupella yakushimensis (Horik.) S.Hatt....................................................................................114 Mastigobryum aberrans Steph. .....................................................................................................510 Mastigobryum asperum Steph.......................................................................................................510 Mastigobryum deningeri Herzog...................................................................................................510 Mastigobryum karstenii Steph. .....................................................................................................510 Mastigobryum ledermannii Steph. ................................................................................................510 Mastigobryum londbergii Steph. ...................................................................................................510 Mastigobryum longifolium Steph. .................................................................................................510 Mastigobryum minutitextum Steph. .............................................................................................510 Mastigobryum multidens Steph. ...................................................................................................510 Mastigobryum muscicola Steph. ....................................................................................................510 Mastigobryum nigricans Herzog ...................................................................................................510 Mastigobryum nipuranum Steph. .................................................................................................510 Mastigobryum palmicola Steph.....................................................................................................510 Mastigobryum rajanum Herzog ...................................................................................................510 Mastigobryum ribehanum Steph...................................................................................................510 Mastigobryum ruicaule Beauverd ................................................................................................510 Mastigobryum schraderbergii Steph. .............................................................................................510 Mastigobryum squamulistipum Steph. ..........................................................................................510 Mastigobryum subhyalinum Steph. ...............................................................................................510 Mastigobryum venezuelanum Molk. .............................................................................................510 Mastigobryum vermiculare Herzog ...............................................................................................510 Mastigolejeunea auriculata (Wilson et Hook.) Steph....................................................................409 Mastigolejeunea auriculata var. rhodesica (Vanden Berghen) Sukkharak et Gradst. .......................409 Mastigolejeunea calcarata (Steph.) Verd. ......................................................................................409 Mastigolejeunea lorea (Mitt.) Paris ..............................................................................................409 Mastigolejeunea frauenfeldii (Reichardt) Verd. .............................................................................409 Mastigolejeunea gradsteinii Sukkharak .........................................................................................409 Mastigolejeunea humilis (Gottsche) Schifn. .................................................................................410 Mastigolejeunea indica Steph. ......................................................................................................410 Mastigolejeunea innovans (Spruce) Steph. ....................................................................................410 Mastigolejeunea ligulata (Lehm. et Lindenb.) Schifn...................................................................410 Mastigolejeunea nigra Steph. ........................................................................................................410 World checklist of hornworts and liverworts *** *** *** *** ** *** ** ** ** ** * * ** *** ** ** * * * * ** ** * ** *** ** *** ** *** *** * *** * ** *** ** *** ** * * *** *** ** * * * *** *** ** ** *** * 603 Mastigolejeunea plicatilora (Spruce) Steph. .................................................................................410 Mastigolejeunea recondita (Steph.) Mizut. ....................................................................................410 Mastigolejeunea repleta (Taylor) A.Evans .....................................................................................410 Mastigolejeunea truncata Mizut. ..................................................................................................410 Mastigolejeunea turgida Steph. .....................................................................................................410 Mastigolejeunea virens (Ångstr.) Steph. nom. conserv. ...................................................................410 Mastigopelma fragile (Steph.) N.Kitag. ........................................................................................163 Mastigopelma pulvinulatum (De Not.) Grolle ..............................................................................163 Mastigopelma simplex Mitt. .........................................................................................................163 Mastigopelma subissum Grolle .....................................................................................................163 Mastigophora appendiculata Steph. ..............................................................................................218 Mastigophora attenuata (Taylor) Trevis. ......................................................................................218 Mastigophora caledonica Steph. ....................................................................................................218 Mastigophora diclados (Brid. ex F.Weber) Nees ...........................................................................218 Mastigophora diclados var. borneensis (De Not.) Schifn. ..............................................................218 Mastigophora diclados var. ramentiissa Herzog.............................................................................218 Mastigophora diclados var. villosa Herzog .....................................................................................218 Mastigophora guineensis Steph. ....................................................................................................218 Mastigophora humillima (Taylor) Trevis. .....................................................................................218 Mastigophora pyramidana Steph. .................................................................................................218 Mastigophora sepikiana Piippo .....................................................................................................218 Mastigophora tuberculata D.H.Mill. et H.A.Mill. ........................................................................218 Mastigophora valida Steph. ..........................................................................................................218 Mastigophora viridula (Nees) Trevis. ...........................................................................................218 Mastigophora woodsii (Hook.) Nees.............................................................................................218 Megaceros aneuriformis Steph. .......................................................................................................34 Megaceros austronesophilus Cargill et Seppelt .................................................................................34 Megaceros ciliatus K.I.Goebel ........................................................................................................34 Megaceros denticulatus (Lehm.) Steph. ...........................................................................................34 Megaceros lagellaris (Mitt.) Steph..................................................................................................34 Megaceros lavens (Spruce) Campb. ...............................................................................................34 Megaceros gracilis (Reichardt) Steph. .............................................................................................34 Megaceros jamesonii (Taylor) Steph. ..............................................................................................34 Megaceros leptohymenius (Hook.f. et Taylor) Steph........................................................................34 Megaceros pellucidus (Colenso) E.A.Hodgs. ...................................................................................34 Megaceros tjibodensis Campb. ........................................................................................................34 Megalembidium insulanum (W.Martin et E.A.Hodgs.) R.M.Schust. ...........................................168 Meinungeria mouensis Frank Müll. ..............................................................................................143 Mesoceros mesophoros Piippo ..........................................................................................................38 Mesoceros porcatus Piippo ..............................................................................................................38 Mesoptychia badensis (Gottsche ex Rabenh.) L.Söderstr. et Váňa .................................................120 Mesoptychia bantriensis (Hook.) L.Söderstr. et Váňa ....................................................................120 Mesoptychia bantriensis subsp. wallischii (Ştefănuţ) L.Söderstr. et Váňa ......................................120 Mesoptychia chichibuensis (Inoue) L.Söderstr. et Váňa .................................................................120 Mesoptychia collaris (Nees) L.Söderstr. et Váňa ............................................................................120 Mesoptychia itzgeraldiae (Paton et A.R.Perry) L.Söderstr. et Váňa ..............................................121 Mesoptychia gillmanii (Austin) L.Söderstr. et Váňa ......................................................................121 Mesoptychia heterocolpos (hed. ex Hartm.) L.Söderstr. et Váňa...................................................121 Mesoptychia heterocolpos var. arctica (S.W.Arnell) L.Söderstr. et Váňa..........................................121 Mesoptychia heterocolpos var. harpanthoides (Bryhn et Kaal.) L.Söderstr. et Váňa..........................121 Mesoptychia igiana (S.Hatt.) L.Söderstr. et Váňa .........................................................................121 Mesoptychia mamatkulovii (Duda) L.Söderstr. et Váňa ................................................................121 604 *** *** * *** ** *** *** *** * *** ** ** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** ** *** *** *** ** ** *** *** *** ** *** ** *** *** ** ** *** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Mesoptychia mayebarae (S.Hatt.) L.Söderstr. et Váňa...................................................................121 Mesoptychia morrisoncola (Horik.) L.Söderstr. et Váňa ................................................................121 Mesoptychia polymorpha Stotler, Crand.-Stotl. et Bakalin ............................................................121 Mesoptychia rutheana (Limpr.) L.Söderstr. et Váňa......................................................................121 Mesoptychia rutheana var. laxa (Schifn. ex Burrell) L.Söderstr. et Váňa ......................................121 Mesoptychia sahlbergii (Lindb. et Arnell) A.Evans ........................................................................121 Mesoptychia subcrispa (Herzog) L.Söderstr. et Váňa .....................................................................121 Mesoptychia turbinata (Raddi) L.Söderstr. et Váňa ......................................................................122 Mesoptychia ussuriensis (Bakalin) L.Söderstr. et Váňa ...................................................................122 Metacalypogeia alternifolia (Nees) Grolle .....................................................................................107 Metacalypogeia cordifolia (Steph.) Inoue ......................................................................................107 Metalejeunea crassitexta (J.B.Jack et Steph.) Pócs .........................................................................300 Metalejeunea cucullata (Reinw., Blume et Nees) Grolle ...............................................................300 Metalejeunea winkleri R.L.Zhu et Grolle .....................................................................................300 Metzgeria acuminata Steph. ........................................................................................................457 Metzgeria adscendens Steph. ex K.I.Goebel ..................................................................................457 Metzgeria agnewiae Kuwah..........................................................................................................457 Metzgeria albinea Spruce .............................................................................................................457 Metzgeria albinea var. aberrans Schifn. .......................................................................................457 Metzgeria albinea var. angusta (Steph.) D.P.Costa et Gradst. ......................................................457 Metzgeria allionii Steph...............................................................................................................457 Metzgeria alpina R.M.Schust. et J.J.Engel ...................................................................................457 Metzgeria americana Masuzaki ....................................................................................................457 Metzgeria attenuata Steph. ..........................................................................................................457 Metzgeria aurantiaca Steph. ........................................................................................................457 Metzgeria auriculata Grolle et Kuwah. ........................................................................................457 Metzgeria bahiensis Schifn. .........................................................................................................457 Metzgeria bartlettii Kuwah. .........................................................................................................457 Metzgeria bischlerae Kuwah. ........................................................................................................457 Metzgeria bracteata Spruce ..........................................................................................................457 Metzgeria brasiliensis Schifn. ......................................................................................................457 Metzgeria chilensis Steph. ............................................................................................................457 Metzgeria ciliata Raddi................................................................................................................457 Metzgeria clavilora Spruce ..........................................................................................................457 Metzgeria cleeii Kuwah. ..............................................................................................................458 Metzgeria comata Steph. ..............................................................................................................458 Metzgeria conjugata Lindb...........................................................................................................458 Metzgeria consanguinea Schifn....................................................................................................458 Metzgeria convoluta Steph. ..........................................................................................................458 Metzgeria coorgensis S.C.Srivast. et S.Srivast. ...............................................................................458 Metzgeria corralensis Steph. .........................................................................................................458 Metzgeria crassipilis (Lindb.) A.Evans ..........................................................................................458 Metzgeria cratoneura Schifn. ......................................................................................................458 Metzgeria cylindra Kuwah. ..........................................................................................................458 Metzgeria decrescens Steph. ..........................................................................................................458 Metzgeria dichotoma (Sw.) Nees ..................................................................................................458 Metzgeria divaricata A.Evans.......................................................................................................458 Metzgeria dorsipara (Herzog) Kuwah. .........................................................................................458 Metzgeria duricosta Steph. ...........................................................................................................458 Metzgeria engelii Kuwah..............................................................................................................458 Metzgeria epiphylla A.Evans ........................................................................................................458 Metzgeria ilicina Mitt. ................................................................................................................458 World checklist of hornworts and liverworts ** * *** ** *** *** * * *** ** ** *** *** * ** *** *** ** *** ** ** *** *** ** ** *** *** ** *** ** ** *** ** *** *** ** *** *** *** *** ** *** *** *** *** *** ** ** ** ** ** *** 605 Metzgeria lavovirens Colenso ......................................................................................................458 Metzgeria foliicola Schifn............................................................................................................458 Metzgeria francana Steph. ...........................................................................................................458 Metzgeria frontipilis Lindb. .........................................................................................................459 Metzgeria fruticola Spruce ...........................................................................................................459 Metzgeria furcata (L.) Corda .......................................................................................................459 Metzgeria furcata var. expansa Douin ..........................................................................................459 Metzgeria furcata var. paciica Brinkm. ........................................................................................459 Metzgeria grandilora A.Evans .....................................................................................................459 Metzgeria hasselii Kuwah. ............................................................................................................459 Metzgeria hebridensis Steph. ........................................................................................................459 Metzgeria hegewaldii Kuwah. ......................................................................................................459 Metzgeria herminieri Schifn........................................................................................................459 Metzgeria heteroramea Steph. ......................................................................................................459 Metzgeria imberbis J.B.Jack et Steph............................................................................................459 Metzgeria inlata Steph................................................................................................................459 Metzgeria jamesonii Kuwah. ........................................................................................................459 Metzgeria kanaii Kuwah..............................................................................................................459 Metzgeria kinabaluensis Masuzaki................................................................................................459 Metzgeria kuwaharae Piippo........................................................................................................459 Metzgeria laciniata Kuwah. .........................................................................................................459 Metzgeria lechleri Steph. ..............................................................................................................459 Metzgeria leptoneura Spruce ........................................................................................................459 Metzgeria leptoneura var. breviseta (Schifn.) O.Yano...................................................................459 Metzgeria leptoneura var. polychaeta R.M.Schust. ........................................................................459 Metzgeria liebmaniana Lindenb. et Gottsche ...............................................................................459 Metzgeria lindbergii Schifn. ........................................................................................................459 Metzgeria litoralis J.J.Engel et Kuwah..........................................................................................459 Metzgeria longitexta Steph. ..........................................................................................................460 Metzgeria macrospora Kuwah. .....................................................................................................460 Metzgeria macveanii Kuwah. .......................................................................................................460 Metzgeria maegdefraui Kuwah. ....................................................................................................460 Metzgeria magellanica Schifn......................................................................................................460 Metzgeria metaensis Kuwah. ........................................................................................................460 Metzgeria mexicana Steph. ..........................................................................................................460 Metzgeria monoica Kuwah. et J.J.Engel .......................................................................................460 Metzgeria myriopoda Lindb. ........................................................................................................460 Metzgeria neotropica Kuwah. .......................................................................................................460 Metzgeria nudifrons Steph. ..........................................................................................................460 Metzgeria parviinvolucrata Kuwah...............................................................................................460 Metzgeria patagonica Steph. ........................................................................................................460 Metzgeria polytricha Spruce .........................................................................................................460 Metzgeria procera Mitt. ...............................................................................................................460 Metzgeria psilocraspeda Schifn. ...................................................................................................460 Metzgeria pubescens (Schrank) Raddi ...........................................................................................460 Metzgeria pulvinata Steph. ..........................................................................................................460 Metzgeria quadrifaria Steph. .......................................................................................................460 Metzgeria raoi S.C.Srivast. et S.Srivast. .......................................................................................460 Metzgeria rigida Lindb. ...............................................................................................................460 Metzgeria robinsonii Steph. .........................................................................................................460 Metzgeria roivainenii Kuwah. ......................................................................................................461 Metzgeria rufula Spruce...............................................................................................................461 606 ** ** ** *** *** ** ** * *** ** *** *** ** ** ** *** *** *** *** ** ** ** ** ** ** *** * ** ** ** ** ** *** ** *** *** *** ** *** ** ** ** ** *** ** * ** ** * *** * ** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Metzgeria saccata Mitt. ................................................................................................................461 Metzgeria saxbyi Pearson .............................................................................................................461 Metzgeria scobina Mitt. ...............................................................................................................461 Metzgeria scyphigera A.Evans .......................................................................................................461 Metzgeria senjoana Masuzaki .......................................................................................................461 Metzgeria setigera R.M.Schust. ex Crand.-Stotl. et L.Söderstr......................................................461 Metzgeria sikkimensis S.C.Srivast. et K.K.Rawat ..........................................................................461 Metzgeria simplex Lorb. ex Müll.Frib. .........................................................................................461 Metzgeria sinuata Loitl. ...............................................................................................................461 Metzgeria sparrei Kuwah. ............................................................................................................461 Metzgeria spindleri Steph.............................................................................................................461 Metzgeria subaneura Schifn. .......................................................................................................461 Metzgeria submarginata M.L.So ..................................................................................................461 Metzgeria subundulata (Lindb.) Kuwah. ......................................................................................461 Metzgeria temperata Kuwah. .......................................................................................................461 Metzgeria uncigera A.Evans .........................................................................................................461 Metzgeria undulata Kuwah..........................................................................................................461 Metzgeria violacea (Ach.) Dumort. ..............................................................................................461 Microlejeunea acutifolia Steph. ....................................................................................................385 Microlejeunea africana Steph. ......................................................................................................385 Microlejeunea aligera (Mitt.) Steph..............................................................................................385 Microlejeunea ankasica E.W.Jones ...............................................................................................385 Microlejeunea aphanella (Spruce) Steph. ......................................................................................385 Microlejeunea atsuana Steph. .......................................................................................................385 Microlejeunea bischlerae (B.M.hiers) B.M.hiers .......................................................................385 Microlejeunea bullata (Taylor) Steph. ..........................................................................................385 Microlejeunea byssoides (Gottsche) Pearson ..................................................................................385 Microlejeunea capillaris (Gottsche) Steph. ...................................................................................385 Microlejeunea cochlearifolia Steph. ...............................................................................................386 Microlejeunea colombiana Bischl. .................................................................................................386 Microlejeunea constricta (Grolle) Grolle .......................................................................................386 Microlejeunea crenulifolia (Gottsche) Steph. ................................................................................386 Microlejeunea cystifera Herzog .....................................................................................................386 Microlejeunea dispar Jovet-Ast .....................................................................................................386 Microlejeunea epiphylla Bischl......................................................................................................386 Microlejeunea ilicuspis (Steph.) Heinrichs, Schäf.-Verw., Pócs et S.Dong ...................................386 Microlejeunea ischeri (Tixier) Heinrichs, Schäf.-Verw., Pócs et S.Dong ......................................386 Microlejeunea issistipula Steph. ...................................................................................................386 Microlejeunea globosa (Spruce) Steph. ..........................................................................................386 Microlejeunea herzogiana Steph. ..................................................................................................386 Microlejeunea indica (Udar et U.S.Awasthi) Y.M.Wei et R.L.Zhu...............................................386 Microlejeunea inlata Steph. ........................................................................................................386 Microlejeunea kinabaluensis (Mizut.) Grolle ................................................................................386 Microlejeunea latitans (Hook.f. et Taylor) Heinrichs, Schäf.-Verw., Pócs et S.Dong ...................386 Microlejeunea lunulatiloba Horik. ...............................................................................................386 Microlejeunea magnilobula Gola ..................................................................................................386 Microlejeunea mammillosa (Mizut.) Grolle ..................................................................................387 Microlejeunea minutissima (Mizut.) Grolle ..................................................................................387 Microlejeunea minutistipula Steph. ..............................................................................................387 Microlejeunea moniliata (Mizut.) R.L.Zhu et Y.M.Wei ...............................................................387 Microlejeunea nepalensis Steph. ....................................................................................................387 Microlejeunea nyandaruensis Pócs ................................................................................................387 World checklist of hornworts and liverworts ** ** * ** ** ** ** ** *** ** *** * ** *** * ** * *** ** *** *** *** *** *** *** ** ** ** ** *** ** ** *** * ** *** *** *** *** * * * * *** *** ** *** *** *** *** ** ** 607 Microlejeunea oblongistipula (Gottsche) Pearson ..........................................................................387 Microlejeunea ocellata (Herzog) Grolle ........................................................................................387 Microlejeunea ovistipula Steph. ....................................................................................................387 Microlejeunea papulosa (Gottsche) Pearson ..................................................................................387 Microlejeunea perpusilla (Spruce) Steph. ......................................................................................387 Microlejeunea punctiformis (Taylor) Steph. ..................................................................................387 Microlejeunea pyriformis (Lindenb. et Gottsche) Steph. ...............................................................387 Microlejeunea spinosa (Mizut.) Grolle ..........................................................................................387 Microlejeunea squarrosa (Steph.) Heinrichs, Schäf.-Verw., Pócs et S.Dong ..................................387 Microlejeunea strasbergii Bardat et Ah-Peng .................................................................................387 Microlejeunea subulistipa Steph. ..................................................................................................388 Microlejeunea szechuanensis P.C.Chen .........................................................................................388 Microlejeunea udarii P.K.Verma et S.C.Srivast. ...........................................................................388 Microlejeunea ulicina (Taylor) Steph. ..........................................................................................388 Microlejeunea usambarensis Steph. ...............................................................................................388 Microlejeunea valenciana Steph. ..................................................................................................388 Microlejeunea victoriensis D.J.Carr...............................................................................................388 Microlejeunea wallichiana (Lehm.) R.L.Zhu et Y.M.Wei ............................................................388 Micropterygium angustistipulum Spruce .......................................................................................181 Micropterygium bialatum Fulford ................................................................................................181 Micropterygium bolivarense Fulford .............................................................................................181 Micropterygium campanense Spruce .............................................................................................181 Micropterygium carinatum (Grev.) Reimers .................................................................................181 Micropterygium conchifolium Reimers ..........................................................................................181 Micropterygium duidae Reimers ...................................................................................................181 Micropterygium exalatum Steph. ..................................................................................................181 Micropterygium grandistipulum Steph. .........................................................................................181 Micropterygium laeve H.Rob. ......................................................................................................181 Micropterygium lechleri Reimers ..................................................................................................181 Micropterygium leiophyllum Spruce ..............................................................................................181 Micropterygium longicellulatum Uribe et E.L.Linares ...................................................................182 Micropterygium parvistipulum Spruce ..........................................................................................182 Micropterygium pterygophyllum (Nees) Trevis. .............................................................................182 Micropterygium pterygophyllum var. lancifolium Reimers ..............................................................182 Micropterygium reimersianum Herzog..........................................................................................182 Micropterygium steyermarkii Fulford ............................................................................................182 Micropterygium tatei Reimers ......................................................................................................182 Micropterygium tenax (Steph.) Grolle ..........................................................................................182 Micropterygium trachyphyllum Reimers ........................................................................................182 Micropterygium trachyphyllum var. brasiliense Reimers .................................................................182 Micropterygium trachyphyllum var. cubense Reimers .....................................................................182 Micropterygium trachyphyllum var. guadeloupense Reimers ...........................................................182 Micropterygium trachyphyllum var. jamaicense Reimers ................................................................182 Micropterygium tumidulum Fulford .............................................................................................182 Mizutania riccardioides Furuki et Z.Iwats....................................................................................107 Mnioloma bolivianum (Fulford) R.M.Schust. ..............................................................................108 Mnioloma caespitosum (Spruce) R.M.Schust. ...............................................................................108 Mnioloma cellulosum (Spreng.) R.M.Schust. ...............................................................................108 Mnioloma crenulatum (Bischl.) R.M.Schust. ...............................................................................108 Mnioloma cyclostipum (Spruce) R.M.Schust. ...............................................................................108 Mnioloma elliottii (Steph.) R.M.Schust. ......................................................................................108 Mnioloma issistipulum (Bischl.) R.M.Schust. ..............................................................................108 608 *** *** ** *** ** *** ** *** *** *** *** *** *** *** *** ** *** *** *** *** *** *** *** *** *** ** *** *** *** *** *** *** *** *** * * *** *** *** * * *** ** ** *** *** * * *** *** *** *** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Mnioloma fuscum (Lehm.) R.M.Schust. ......................................................................................108 Mnioloma nephrostipum (Spruce) R.M.Schust. ............................................................................108 Mnioloma novaezelandiae J.J.Engel .............................................................................................108 Mnioloma parallelogramum (Spruce) R.M.Schust. .......................................................................108 Mnioloma retusum (Bischl.) R.M.Schust. ....................................................................................108 Mnioloma rhynchophyllum Herzog ..............................................................................................108 Mnioloma stamatotonum M.A.M.Renner et E.A.Br. ....................................................................108 Mnioloma venezuelanum (Fulford) R.M.Schust. ..........................................................................108 Moerckia blyttii (Mørch) Brockm. ...............................................................................................469 Moerckia lotoviana (Nees) Schifn. .............................................................................................469 Moerckia hibernica (Hook.) Gottsche ..........................................................................................469 Monocarpus sphaerocarpus D.J.Carr .............................................................................................504 Monoclea forsteri Hook................................................................................................................492 Monoclea gottschei Lindb. ............................................................................................................492 Monoclea gottschei subsp. elongata Gradst. et Mues ......................................................................492 Monodactylopsis monodactyla (Spruce) R.M.Schust. .....................................................................183 Monosolenium tenerum Grif. ......................................................................................................492 Mylia anomala (Hook.) Gray ......................................................................................................258 Mylia taylorii (Hook.) Gray ........................................................................................................258 Mylia verrucosa Lindb. ................................................................................................................258 Mylia verrucosa subsp. nuda (Inoue et B.Y.Yang) Potemkin et Kazan. .........................................259 Myriocoleopsis luviatilis (Steph.) M.E.Reiner et Gradst. ..............................................................348 Myriocoleopsis gymnocolea (Spruce) M.E.Reiner et Gradst. ...........................................................348 Myriocoleopsis minutissima (Sm.) R.L.Zhu, Y.Yu et Pócs .............................................................348 Myriocoleopsis minutissima subsp. myriocarpa (Nees et Mont.) R.L.Zhu, Y.Yu et Pócs .................348 Myriocoleopsis vuquangensis (Pócs et Ninh) Pócs ..........................................................................349 Mytilopsis albifrons Spruce ...........................................................................................................182 Nanomarsupella xenophylla (R.M.Schust.) R.M.Schust. ex A.Hagborg, L.Söderstr. et von Konrat......110 Nardia arnelliana Grolle .............................................................................................................114 Nardia assamica (Mitt.) Amakawa...............................................................................................114 Nardia breidleri (Limpr.) Lindb. .................................................................................................114 Nardia compressa (Hook.) Gray...................................................................................................114 Nardia lagelliformis Inoue ..........................................................................................................114 Nardia geoscyphus (De Not.) Lindb. ............................................................................................114 Nardia geoscyphus var. dioica Bakalin ...........................................................................................114 Nardia geoscyphus var. suberecta (Lindb. ex Kaal.) Váňa ...............................................................114 Nardia grollei Váňa et D.G.Long ................................................................................................115 Nardia insecta Lindb. ..................................................................................................................115 Nardia japonica Steph. ................................................................................................................115 Nardia kamtschatica Arnell et C.E.O.Jensen ...............................................................................115 Nardia leptocaulis C.Gao.............................................................................................................115 Nardia lescurii (Austin) Underw. ................................................................................................115 Nardia minutifolia Furuki ...........................................................................................................115 Nardia nuda (Lindenb. et Gottsche) Váňa ..................................................................................115 Nardia poeltii Váňa .....................................................................................................................115 Nardia scalaris Gray ....................................................................................................................115 Nardia scalaris var. botryoidea (R.M.Schust.) Váňa ......................................................................115 Nardia scalaris var. harae (Amakawa) Váňa .................................................................................115 Nardia subclavata (Steph.) Amakawa ..........................................................................................115 Nardia succulenta (A.Rich.) Spruce .............................................................................................115 Nardia unispiralis Amakawa ........................................................................................................115 Neesioscyphus allionii (Steph.) Grolle ...........................................................................................102 World checklist of hornworts and liverworts *** *** *** *** * *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** ** ** ** ** *** 609 Neesioscyphus argillaceus (Nees) Grolle.........................................................................................102 Neesioscyphus bicuspidatus (Steph.) Grolle....................................................................................102 Neesioscyphus carneus (Nees) Grolle .............................................................................................102 Neesioscyphus homophyllus (Nees) Grolle ......................................................................................102 Nemoursia tuberculata Mérat .......................................................................................................511 Neogrollea notabilis E.A.Hodgs....................................................................................................183 Neohattoria herzogii (S.Hatt.) Kamim. ........................................................................................296 Neohodgsonia mirabilis (Perss.) Perss. ..........................................................................................504 Neolepidozia aubertii (Jovet-Ast) E.D.Cooper .............................................................................176 Neolepidozia autoica (J.J.Engel et G.L.Merr.) E.D.Cooper ..........................................................176 Neolepidozia capilligera (Schwägr.) Fulford et J.Taylor ................................................................176 Neolepidozia consobrina (J.J.Engel et G.L.Merr.) E.D.Cooper .....................................................176 Neolepidozia cuneifolia (Steph.) Fulford et J.Taylor .....................................................................177 Neolepidozia disparata (J.J.Engel et G.L.Merr.) E.D.Cooper .......................................................177 Neolepidozia disticha (Steph.) Fulford et J.Taylor ........................................................................177 Neolepidozia gibbsiana (Steph.) E.D.Cooper ...............................................................................177 Neolepidozia heterotexta (Steph.) E.D.Cooper .............................................................................177 Neolepidozia hodgsoniae (J.J.Engel et G.L.Merr.) E.D.Cooper .....................................................177 Neolepidozia longitudinalis (Herzog) E.D.Cooper .......................................................................177 Neolepidozia mamillosa (Schifn.) E.D.Cooper ............................................................................177 Neolepidozia meridiana (E.A.Hodgs.) E.D.Cooper......................................................................177 Neolepidozia oligophylla (Lehm. et Lindenb.) Fulford et J.Taylor ................................................177 Neolepidozia ophiria (Gottsche) E.D.Cooper ..............................................................................177 Neolepidozia palmata (J.J.Engel et G.L.Merr.) E.D.Cooper ........................................................177 Neolepidozia paludicola (E.A.Hodgs.) E.D.Cooper ......................................................................177 Neolepidozia papulosa (Steph.) Fulford et J.Taylor ......................................................................177 Neolepidozia parvifolia (Steph.) Fulford et J.Taylor .....................................................................178 Neolepidozia patentissima (Hook.f. et Taylor) E.D.Cooper..........................................................178 Neolepidozia patentissima var. ampliata (J.J.Engel et G.L.Merr.) E.D.Cooper .............................178 Neolepidozia patentissima var. zebrina (J.J.Engel et G.L.Merr.) E.D.Cooper................................178 Neolepidozia pennata (J.J.Engel et G.L.Merr.) E.D.Cooper .........................................................178 Neolepidozia planifolia (Steph.) E.D.Cooper ...............................................................................178 Neolepidozia praenitens (Lehm. et Lindenb.) E.D.Cooper ...........................................................178 Neolepidozia praenitens var. dentifolia (J.J.Engel et G.L.Merr.) E.D.Cooper ................................178 Neolepidozia quadristipula (Steph.) E.D.Cooper .........................................................................178 Neolepidozia rectangularis (R.M.Schust.) E.D.Cooper .................................................................178 Neolepidozia seriatitexta (Steph.) Fulford.....................................................................................178 Neolepidozia tetrapila (Hook.f. et Taylor) E.D.Cooper ...............................................................178 Neolepidozia tetrapila var. cancellata (Colenso) E.D.Cooper ........................................................178 Neolepidozia tetrapila var. roseana (Steph.) E.D.Cooper ..............................................................179 Neolepidozia tridactylis (Lehm. et Lindenb.) E.D.Cooper ............................................................179 Neolepidozia triida (Steph.) E.D.Cooper ....................................................................................179 Neolepidozia verruculosa (J.J.Engel et G.L.Merr.) E.D.Cooper ....................................................179 Neolepidozia wallichiana (Gottsche) Fulford et J.Taylor ..............................................................179 Neoorthocaulis attenuatus (Mart.) L.Söderstr., De Roo et Hedd.....................................................53 Neoorthocaulis binsteadii (Kaal.) L.Söderstr., De Roo et Hedd. .....................................................53 Neoorthocaulis loerkei (F.Weber et D.Mohr) L.Söderstr., De Roo et Hedd. ..................................53 Neoorthocaulis hyperboreus (R.M.Schust.) L.Söderstr., De Roo et Hedd. .......................................53 Neotrichocolea bissetii (Mitt.) S.Hatt............................................................................................434 Nephelolejeunea bidentata B.M.hiers ex L.Söderstr. et A.Hagborg .............................................349 Nephelolejeunea carcharias M.A.M.Renner ..................................................................................349 Nephelolejeunea conchophylla Grolle ............................................................................................349 610 ** *** *** *** *** ** *** *** ** *** *** ** *** *** *** *** *** *** * *** *** *** ** *** *** ** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** ** *** *** *** *** *** *** *** *** *** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Nephelolejeunea fragilis (R.M.Schust.) L.Söderstr. et A.Hagborg .................................................349 Nephelolejeunea hamata Grolle ....................................................................................................349 Nephelolejeunea hispida R.M.Schust. ex L.Söderstr. et A.Hagborg ...............................................349 Nephelolejeunea jarmaniana Grolle ex L.Söderstr. et A.Hagborg .................................................349 Nephelolejeunea nudipes (Hook.f. et Taylor) L.Söderstr. et A.Hagborg ........................................349 Nephelolejeunea occidentalis Pócs ex L.Söderstr. et A.Hagborg .....................................................349 Nephelolejeunea papillosa Glenny.................................................................................................349 Nephelolejeunea radulifolia (C.Massal.) L.Söderstr. et A.Hagborg ................................................349 Nephelolejeunea secunda M.A.M.Renner ex L.Söderstr. et A.Hagborg .........................................349 Nephelolejeunea talinayi (S.W.Arnell) Grolle ...............................................................................349 Neurolejeunea breutelii (Gottsche) A.Evans .................................................................................299 Neurolejeunea breutelii var. africana Pócs.....................................................................................299 Neurolejeunea catenulata (Nees) Schifn. .....................................................................................299 Neurolejeunea sastreana Gradst. ...................................................................................................299 Neurolejeunea seminervis (Spruce) Schifn....................................................................................300 Nipponolejeunea pilifera (Steph.) S.Hatt. .....................................................................................296 Nipponolejeunea subalpina (Horik.) S.Hatt..................................................................................296 Noteroclada conluens Taylor .......................................................................................................474 Noteroclada longiuscula Colenso ..................................................................................................474 Nothoceros aenigmaticus J.C.Villarreal et K.D.McFarland ..............................................................35 Nothoceros canaliculatus (Pagán) J.C.Villarreal, Hässel et N.Salazar ...............................................35 Nothoceros endiviifolius (Mont.) J.Haseg. ex J.C.Villarreal, Hässel et N.Salazar .............................35 Nothoceros fuegiensis (Steph.) J.C.Villarreal....................................................................................35 Nothoceros giganteus (Lehm. et Lindenb.) J.Haseg. ex J.C.Villarreal, Hässel et N.Salazar ...............35 Nothoceros minarum (Nees) J.C.Villarreal .....................................................................................35 Nothoceros renzagliensis J.C.Villarreal, L.V.Campos et Uribe .........................................................35 Nothoceros schizophyllus (Steph.) J.C.Villarreal ..............................................................................35 Nothoceros superbus J.C.Villarreal, Hässel et N.Salazar...................................................................35 Nothoceros vincentianus (Lehm. et Lindenb.) J.C.Villarreal............................................................35 Nothogymnomitrion erosum (Carrington et Pearson) R.M.Schust...................................................75 Nothostrepta biida (Steph.) R.M.Schust........................................................................................46 Nothostrepta longissima (Steph.) R.M.Schust. ................................................................................46 Notoscyphus lutescens (Lehm. et Lindenb.) Mitt. ..........................................................................122 Notothylas anaporata Udar et D.K.Singh .......................................................................................37 Notothylas breutelii (Gottsche) Gottsche........................................................................................37 Notothylas decurva (Mitt.) Steph. ..................................................................................................37 Notothylas depressispora J.Haseg. ....................................................................................................37 Notothylas dissecta Steph. ...............................................................................................................37 Notothylas labellata Steph. ............................................................................................................37 Notothylas galapagensis M.Howe....................................................................................................37 Notothylas himalayensis Udar et D.K.Singh ...................................................................................37 Notothylas indica Kashyap .............................................................................................................37 Notothylas irregularis Chantanaorr.................................................................................................37 Notothylas javanica (Sande Lac.) Gottsche.....................................................................................37 Notothylas kashyapii D.K.Singh .....................................................................................................38 Notothylas khasiana Udar et D.K.Singh.........................................................................................37 Notothylas nepalensis D.K.Singh ....................................................................................................38 Notothylas orbicularis (Schwein.) Sull. ...........................................................................................37 Notothylas pandei Udar et V.Chandra ...........................................................................................37 Notothylas pleidereri Udar et D.K.Singh .......................................................................................37 Notothylas temperata J.Haseg. ........................................................................................................38 Notothylas udarii D.K.Singh et Semwal.........................................................................................38 World checklist of hornworts and liverworts * *** ** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** ** *** ** ** *** *** *** *** *** *** *** *** * *** ** *** *** *** *** ** *** ** * *** ** *** ** ** ** *** *** *** *** 611 Notothylas verdoornii Khanna ........................................................................................................38 Notothylas vitalii Udar et D.K.Singh .............................................................................................38 Notothylas yunnanensis T.Peng et R.L.Zhu ....................................................................................38 Nowellia aciliata (P.C.Chen et P.C.Wu) Mizut. ...........................................................................62 Nowellia borneensis (De Not.) Schifn. ..........................................................................................61 Nowellia curvifolia (Dicks.) Mitt. ..................................................................................................62 Nowellia dominicensis Steph. .........................................................................................................61 Nowellia evansii Grolle..................................................................................................................61 Nowellia langii Pearson .................................................................................................................61 Nowellia pusilla Grolle ..................................................................................................................61 Nowellia reedii H.Rob...................................................................................................................61 Nowellia wrightii (Gottsche ex Spruce) Steph. ex Duss .................................................................61 Nowellia yunckeri Fulford .............................................................................................................61 Obtusifolium obtusum (Lindb.) S.W.Arnell ...................................................................................75 Odontolejeunea decemdentata (Spruce) Steph. ..............................................................................229 Odontolejeunea lunulata (F.Weber) Schifn. ................................................................................229 Odontolejeunea rhomalea (Spruce) Steph. ....................................................................................229 Odontoschisma biidum (Fulford) Gradst., S.C.Aranda et Vanderp. ...............................................63 Odontoschisma brasiliense Steph. ....................................................................................................62 Odontoschisma cleeii Gradst., S.C.Aranda et Vanderp. ..................................................................62 Odontoschisma denudatum (Mart.) Dumort. .................................................................................62 Odontoschisma denudatum subsp. naviculare (Steph.) Gradst., S.C.Aranda et Vanderp..................62 Odontoschisma denudatum subsp. sandvicense (Ångstr.) Gradst., S.C.Aranda et Vanderp. ..............62 Odontoschisma elongatum (Lindb.) A.Evans ...................................................................................62 Odontoschisma engelii Gradst. et Burghardt ...................................................................................62 Odontoschisma luitans (Nees) L.Söderstr. et Váňa .........................................................................63 Odontoschisma francisci (Hook.) L.Söderstr. et Váňa .....................................................................62 Odontoschisma grosseverrucosum Steph. ..........................................................................................63 Odontoschisma jishibae (Steph.) L.Söderstr. et Váňa ......................................................................63 Odontoschisma longilorum (Taylor) Trevis. ...................................................................................63 Odontoschisma macounii (Austin) Underw. ...................................................................................63 Odontoschisma obcordatum (Spruce) Steph. ...................................................................................64 Odontoschisma portoricense (Hampe et Gottsche) Steph. ................................................................63 Odontoschisma pseudogrosseverrucosum Gradst., S.C.Aranda et Vanderp.........................................63 Odontoschisma purpuratum Herzog ...............................................................................................63 Odontoschisma soratamum Fulford ................................................................................................63 Odontoschisma sphagni (Dicks.) Dumort. ......................................................................................64 Odontoschisma spinosum (Fulford) Gradst., S.C.Aranda et Vanderp. .............................................63 Odontoschisma steyermarkii Gradst. et Ilk.-Borg. ...........................................................................63 Odontoschisma variabile (Lindenb. et Gottsche) Trevis..................................................................63 Odontoschisma zhui Gradst., S.C.Aranda et Vanderp. ...................................................................63 Odontoseries chimantana Fulford .................................................................................................183 Oleolophozia perssonii (H.Buch et S.W.Arnell) L.Söderstr., De Roo et Hedd. ...............................75 Omphalanthus baracoensis Mustelier, M.E.Reiner et Gradst. .......................................................316 Omphalanthus iliformis (Sw.) Nees .............................................................................................316 Omphalanthus iliformis var. platycoleus (Herzog) Gradst. ............................................................316 Omphalanthus iliformis var. wallisii (Prantl) Gradst. ...................................................................316 Omphalanthus huanucensis (Gottsche) Gradst. ............................................................................316 Omphalanthus jackii (Prantl) Gradst. ..........................................................................................316 Omphalanthus ovalis (Lindenb. et Gottsche) Gradst. ...................................................................316 Omphalanthus roccatii (Gola) R.M.Schust...................................................................................316 Orthocaulis atlanticus (Kaal.) H.Buch............................................................................................53 612 * * *** * *** *** *** *** ** *** *** *** *** *** ** *** *** *** *** *** *** *** ** * ** *** ** ** ** ** *** ** * ** ** ** *** *** *** ** ** ** *** *** *** *** *** *** ** *** ** ** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Orthocaulis cavifolius H.Buch et S.W.Arnell..................................................................................53 Otigoniolejeunea crenulata Steph. ................................................................................................304 Otigoniolejeunea huctumalcensis (Lindenb. et Gottsche) Y.M.Wei, R.L.Zhu et Gradst. ...............304 Otigoniolejeunea ledermannii Steph. ............................................................................................304 Otigoniolejeunea portoricensis (Hampe et Gottsche) Y.M.Wei, R.L.Zhu et Gradst. ......................304 Otolejeunea australiensis B.M.hiers ............................................................................................392 Otolejeunea hoana (Tixier) Grolle................................................................................................392 Otolejeunea moniliata Grolle .......................................................................................................393 Otolejeunea philippinensis R.L.Zhu et M.L.So .............................................................................393 Otolejeunea rabenorii Tixier ........................................................................................................392 Otolejeunea schmidii (Tixier) Grolle ............................................................................................392 Otolejeunea schnellii (Tixier) R.L.Zhu et M.L.So.........................................................................393 Otolejeunea semperiana (Steph.) Grolle........................................................................................393 Otolejeunea streimannii Grolle.....................................................................................................393 Otolejeunea subana Pócs ..............................................................................................................393 Otolejeunea zantenii Grolle .........................................................................................................393 Otoscyphus crassicaulis (Steph.) J.J.Engel, Bardat et houvenot ....................................................216 Oxymitra cristata Garside ............................................................................................................493 Oxymitra incrassata (Brot.) Sérgio et Sim-Sim .............................................................................493 Pachyglossa austrigena (Hook.f. et Taylor) L.Söderstr. .................................................................216 Pachyglossa austrigena subsp. okaritana (Steph.) L.Söderstr. .........................................................216 Pachyglossa boveana (C.Massal.) L.Söderstr. ................................................................................216 Pachyglossa dissitifolia Herzog et Grolle .......................................................................................216 Pachyglossa exilis (Herzog et Grolle) Hässel et Solari ....................................................................216 Pachyglossa issa (Mitt.) Herzog et Grolle .....................................................................................216 Pachyglossa gottscheoides (Besch. et C.Massal.) L.Söderstr.............................................................216 Pachyglossa grolleana Váňa ...........................................................................................................216 Pachyglossa otiphylla (Hook.f. et Taylor) Váňa.............................................................................216 Pachyglossa spegazziniana (C.Massal.) Herzog et Grolle ...............................................................216 Pachyglossa tenacifolia (Hook.f. et Taylor) Herzog et Grolle ........................................................216 Pallavicinia ambigua (Mitt.) Steph..............................................................................................470 Pallavicinia baldwinii (Austin) A.Evans ......................................................................................470 Pallavicinia bipinnata Steph........................................................................................................470 Pallavicinia camisassai Gola.........................................................................................................470 Pallavicinia cylindrica (Austin) A.Evans ......................................................................................470 Pallavicinia himalayensis Schifn..................................................................................................470 Pallavicinia indica Schifn. ..........................................................................................................470 Pallavicinia levieri Schifn. ..........................................................................................................470 Pallavicinia lyellii (Hook.) Gray ..................................................................................................470 Pallavicinia pilifera Steph............................................................................................................470 Pallavicinia purpurea Steph. ........................................................................................................470 Pallavicinia ridleyi Steph. ............................................................................................................471 Pallavicinia rubristipa Schifn......................................................................................................471 Pallavicinia subciliata (Austin) Steph. .........................................................................................471 Pallavicinia xiphoides (Hook.f. et Taylor) Trevis. ........................................................................471 Paracromastigum denticulatum (Steph.) E.D.Cooper ...................................................................183 Paracromastigum drucei (R.M.Schust.) R.M.Schust.....................................................................184 Paracromastigum dusenii (Steph.) R.M.Schust. ............................................................................184 Paracromastigum iordlandiae R.M.Schust. et J.J.Engel................................................................184 Paracromastigum furcifolium (Steph.) R.M.Schust. ......................................................................184 Paracromastigum granatense (Gottsche) R.M.Schust. ...................................................................184 Paracromastigum longiscyphum (Taylor) R.M.Schust. et J.J.Engel ................................................184 World checklist of hornworts and liverworts ** ** ** *** ** ** ** ** ** ** *** *** *** *** *** ** *** *** *** ** ** ** * * *** *** ** * * ** *** * ** *** ** *** *** *** *** *** ** ** ** *** *** *** *** ** *** *** *** *** 613 Paracromastigum macrostipum (Steph.) R.M.Schust. ...................................................................184 Paracromastigum micromera (Spruce) R.M.Schust. ......................................................................184 Paracromastigum microphyllum (R.M.Schust. ex J.J.Engel) E.D.Cooper ......................................184 Paracromastigum pachyrhizum (Nees) Fulford .............................................................................184 Paracromastigum ryszardii Váňa, Bedn.-Ochyra et Cykowska......................................................184 Paracromastigum stipulatum (Herzog) Fulford.............................................................................184 Paracromastigum subsimplex (Steph.) Fulford et J.Taylor .............................................................184 Paracromastigum succulentum (Sim) J.J.Engel et G.L.Merr. .........................................................184 Paracromastigum tristanianum (R.M.Schust.) J.J.Engel et R.M.Schust. .......................................184 Paracromastigum vastilobum (Steph.) J.J.Engel et G.L.Merr. .......................................................185 Paramomitrion paradoxum R.M.Schust. ......................................................................................110 Paraphymatoceros diadematus Hässel..............................................................................................38 Paraphymatoceros hallii (Austin) Hässel .........................................................................................38 Paraphymatoceros pearsonii (M.Howe) J.C.Villarreal et Cargill ......................................................38 Paraphymatoceros proskaueri (Stotler, Crand.-Stotl. et W.T.Doyle) J.C.Villarreal et Cargill...........38 Pedinophyllopsis abdita (Sull.) R.M.Schust. et Inoue....................................................................219 Pedinophyllum autoicum (Steph.) Inoue ......................................................................................219 Pedinophyllum interruptum (Nees) Kaal. .....................................................................................219 Pedinophyllum monoicum (Steph.) Grolle ....................................................................................219 Pedinophyllum truncatum (Steph.) Inoue.....................................................................................219 Pellia alpicola R.M.Schust. ex L.Söderstr., A.Hagborg et von Konrat ..........................................475 Pellia appalachiana R.M.Schust. .................................................................................................475 Pellia cordaeana Trevis. ...............................................................................................................475 Pellia crispa P.Kumm. .................................................................................................................475 Pellia endiviifolia (Dicks.) Dumort. ............................................................................................475 Pellia epiphylla (L.) Corda ...........................................................................................................475 Pellia epiphylla subsp. borealis (Lorb.) Messe ...............................................................................475 Pellia gottscheana Kreh ................................................................................................................475 Pellia longifolia P.Kumm.............................................................................................................475 Pellia megaspora R.M.Schust. ......................................................................................................475 Pellia neesiana (Gottsche) Limpr. ................................................................................................475 Pellia undulata P.Kumm. ............................................................................................................475 Peltolepis japonica (Shimizu et S.Hatt.) S.Hatt. ...........................................................................484 Peltolepis quadrata (Saut.) Müll.Frib. ..........................................................................................484 Perdusenia rheophila Hässel .........................................................................................................217 Petalophyllum americanum C.H.Ford et Crand.-Stotl. ................................................................468 Petalophyllum hodgsoniae C.H.Ford et Crand.-Stotl. ...................................................................468 Petalophyllum indicum Kashyap ..................................................................................................468 Petalophyllum preissii Gottsche ....................................................................................................468 Petalophyllum ralfsii (Wilson) Nees et Gottsche ..........................................................................468 Phaeoceros austroandinus Hässel.....................................................................................................38 Phaeoceros bolusii (Sim) S.W.Arnell...............................................................................................38 Phaeoceros brevicapsulus (Steph.) Hässel.........................................................................................39 Phaeoceros carolinianus (Michx.) Prosk. .........................................................................................39 Phaeoceros delicatus E.O.Campb. et Outred ..................................................................................39 Phaeoceros dendroceroides (Steph.) Hässel.......................................................................................39 Phaeoceros engelii Cargill et Fuhrer ................................................................................................39 Phaeoceros erectus Udar et D.K.Singh ............................................................................................39 Phaeoceros evanidus (Steph.) Cargill et Fuhrer ...............................................................................39 Phaeoceros exiguus (Steph.) J.Haseg. ..............................................................................................39 Phaeoceros lexivalvis (Gottsche et Nees) Hässel .............................................................................39 Phaeoceros fulvisporus (Steph.) J.Haseg. .........................................................................................39 614 *** ** *** *** *** *** *** ** * *** *** *** *** *** ** ** * *** ** *** *** ** *** ** ** *** *** *** *** *** *** *** *** ** *** ** *** *** *** *** * *** *** ** *** *** *** ** *** *** *** *** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Phaeoceros gemmifer (Horik.) J.Haseg. ...........................................................................................39 Phaeoceros gualaquizanus (Steph.) Gradst. .....................................................................................39 Phaeoceros himalayensis (Kashyap) Prosk. ex Bapna et G.G.Vyas ...................................................39 Phaeoceros huebschmannii Hässel ...................................................................................................39 Phaeoceros inlatus (Steph.) Cargill et Fuhrer .................................................................................39 Phaeoceros kashyapii A.K.Asthana et S.C.Srivast. ...........................................................................40 Phaeoceros laevis (L.) Prosk. ...........................................................................................................40 Phaeoceros maranguensis (Steph.) Bapna ........................................................................................40 Phaeoceros microsporus (Steph.) Hässel ...........................................................................................40 Phaeoceros minutus (Mitt.) S.W.Arnell ..........................................................................................40 Phaeoceros mohrii (Austin) Hässel..................................................................................................40 Phaeoceros oreganus (Austin) Hässel ...............................................................................................40 Phaeoceros parvulus (Schifn.) J.Haseg. ..........................................................................................40 Phaeoceros perpusillus Chantanaorr. ...............................................................................................40 Phaeoceros pichinchensis (Spruce) Hässel ........................................................................................40 Phaeoceros propagulifer (Steph.) Prosk. ..........................................................................................40 Phaeoceros striatisporus J.Haseg ......................................................................................................40 Phaeoceros tenuis (Spruce) Hässel...................................................................................................40 Phaeoceros tigrinus (Gola) J.C.Villarreal.........................................................................................40 Phaeoceros tuberosus (Taylor) Prosk. ..............................................................................................40 Phaeoceros udarii A.K.Asthana et V.Nath ......................................................................................40 Phaeoceros wrightii (Steph.) Hässel ................................................................................................40 Phaeolejeunea amicorum (Hürl.) Pócs ..........................................................................................410 Phaeolejeunea etesseana (Steph.) Mizut. .......................................................................................410 Phaeolejeunea inermis (Steph.) Mizut. .........................................................................................410 Phaeolejeunea latistipula (Schifn. ex P.Syd.) Mizut. ....................................................................410 Phaeomegaceros coriaceus (Steph.) R.J.Duf, J.C.Villarreal, Cargill et Renzaglia .............................35 Phaeomegaceros imbriatus (Gottsche) R.J.Duf, J.C.Villarreal, Cargill et Renzaglia .......................35 Phaeomegaceros foveatus (J.Haseg.) J.C.Villarreal ...........................................................................35 Phaeomegaceros hirticalyx (Steph.) R.J.Duf, J.C.Villarreal, Cargill et Renzaglia.............................36 Phaeomegaceros plicatus (Mitt.) J.C.Villarreal, J.J.Engel et Váňa ....................................................36 Phaeomegaceros skottsbergii (Steph.) R.J.Duf, J.C.Villarreal, Cargill et Renzaglia ..........................36 Phaeomegaceros squamuliger (Spruce) J.C.Villarreal .......................................................................36 Phaeomegaceros squamuliger subsp. hasselii J.C.Villarreal, Cargill et Goinet .................................36 Phycolepidozia exigua R.M.Schust. ................................................................................................75 Phycolepidozia indica Gradst., J.-P.Frahm et U.Schwarz ................................................................75 Phyllothallia fuegiana R.M.Schust. ..............................................................................................474 Phyllothallia nivicola E.A.Hodgs. ................................................................................................474 Phymatoceros bulbiculosus (Brot.) Stotler, W.T.Doyle et Crand.-Stotl. ..........................................36 Phymatoceros phymatodes (M.Howe) R.J.Duf, J.C.Villarreal, Cargill et Renzaglia .........................36 Physocolea tambillensis (Loitl.) Steph. ..........................................................................................511 Pictolejeunea levis Grolle et M.E.Reiner.......................................................................................300 Pictolejeunea mizutanii Grolle .....................................................................................................301 Pictolejeunea piconii Pócs ............................................................................................................301 Pictolejeunea picta (Steph.) Grolle ...............................................................................................301 Pictolejeunea reginae Ilk.-Borg. ....................................................................................................301 Pictolejeunea sprucei Grolle ..........................................................................................................301 Pigafettoa crenulata C.Massal. .....................................................................................................217 Pisanoa chilensis Hässel .................................................................................................................46 Plagiochasma appendiculatum Lehm. et Lindenb. ........................................................................482 Plagiochasma argentinicum Bischl. ...............................................................................................482 Plagiochasma beccarianum Steph. ................................................................................................483 World checklist of hornworts and liverworts *** *** *** *** *** *** *** *** * *** *** *** *** *** *** ** *** *** ** * ** ** ** *** * ** *** *** *** * * ** *** * ** *** * ** ** * ** *** * ** ** *** *** ** ** ** *** ** 615 Plagiochasma cordatum Lehm. et Lindenb. ..................................................................................483 Plagiochasma crenulatum Gottsche ..............................................................................................483 Plagiochasma cuneatum A.Evans ..................................................................................................483 Plagiochasma eximium (Schifn.) Steph. ......................................................................................483 Plagiochasma intermedium Lindenb. et Gottsche .........................................................................483 Plagiochasma jamaicense (Haynes) A.Evans..................................................................................483 Plagiochasma japonicum (Steph.) C.Massal. .................................................................................483 Plagiochasma landii A.Evans........................................................................................................483 Plagiochasma megacarpon (Grif.) Steph. .....................................................................................483 Plagiochasma microcephalum (Steph.) Steph. ...............................................................................483 Plagiochasma microcephalum var. tunesicum Bischl.......................................................................483 Plagiochasma muenchianum Steph. ..............................................................................................483 Plagiochasma pterospermum C.Massal. .........................................................................................483 Plagiochasma rupestre (J.R.Forst. et G.Forst.) Steph. ...................................................................483 Plagiochasma rupestre var. volkii Bischl. .......................................................................................483 Plagiochasma udarii A.Alam et S.C.Srivast. .................................................................................483 Plagiochasma wrightii Sull. ..........................................................................................................483 Plagiochila abietina (Nees) Mont. et Nees ...................................................................................232 Plagiochila abrupta Lehm. et Lindenb. ........................................................................................232 Plagiochila abscendens Gottsche ...................................................................................................238 Plagiochila acanthocaulis Sull. ......................................................................................................224 Plagiochila aculeata (Hook.f. et Taylor) Gottsche, Lindenb. et Nees ...........................................238 Plagiochila acuta Steph. ...............................................................................................................238 Plagiochila adianthoides (Sw.) Lindenb. .......................................................................................220 Plagiochila adianthoides var. aspergillifera Spruce .........................................................................220 Plagiochila aequatorialis Gottsche ................................................................................................233 Plagiochila aerea Taylor ...............................................................................................................225 Plagiochila africana Steph............................................................................................................233 Plagiochila akiyamae Inoue..........................................................................................................233 Plagiochila albertii Steph. ............................................................................................................238 Plagiochila aliena Gottsche ..........................................................................................................238 Plagiochila allionii Steph. ............................................................................................................238 Plagiochila alternans Lindenb. et Gottsche ..................................................................................223 Plagiochila ambigua Lindenb. et Hampe .....................................................................................238 Plagiochila amboynensis Taylor ....................................................................................................226 Plagiochila amicta Steph. .............................................................................................................226 Plagiochila ampliata Steph...........................................................................................................238 Plagiochila andicola Mont. et Gottsche .......................................................................................238 Plagiochila andina Steph. ............................................................................................................238 Plagiochila angolensis Steph. ........................................................................................................238 Plagiochila angulata Steph. ..........................................................................................................224 Plagiochila angusta Lindenb. .......................................................................................................233 Plagiochila angusteoblonga Steph..................................................................................................238 Plagiochila angustisedens Steph. ...................................................................................................238 Plagiochila angustispina Steph. ....................................................................................................238 Plagiochila angustitexta Steph. .....................................................................................................233 Plagiochila annotina Lindenb. .....................................................................................................231 Plagiochila ansata (Hook.f. et Taylor) Gottsche, Lindenb. et Nees ..............................................223 Plagiochila apicalis Gottsche ........................................................................................................238 Plagiochila appalachiana Inoue ....................................................................................................239 Plagiochila arbuscula (Brid. ex Lehm. et Lindenb.) Lindenb. .......................................................233 Plagiochila arbuscula var. rekohuensis J.J.Engel et G.L.Merr. ........................................................233 616 * ** * ** ** * * * ** ** *** *** ** ** *** ** ** * ** *** ** *** *** * * *** ** * *** *** ** ** * ** ** * * ** ** ** *** *** * *** ** ** ** *** *** ** ** *** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Plagiochila arcta S.Winkl. ...........................................................................................................239 Plagiochila arctica Bryhn et Kaal. ................................................................................................228 Plagiochila arctica var. intermedia R.M.Schust.............................................................................228 Plagiochila arcuata Lindenb. .......................................................................................................239 Plagiochila arenacea Schifn. ........................................................................................................239 Plagiochila arnelliana Steph.........................................................................................................220 Plagiochila artsii Pócs ..................................................................................................................239 Plagiochila aspera Steph...............................................................................................................233 Plagiochila aspericaulis Grolle et M.L.So......................................................................................227 Plagiochila aspleniformis R.M.Schust. ..........................................................................................233 Plagiochila asplenioides (L.) Dumort. ...........................................................................................228 Plagiochila assamica Steph. ..........................................................................................................225 Plagiochila atrovirens Taylor ........................................................................................................239 Plagiochila austinii A.Evans .........................................................................................................232 Plagiochila badia Mitt. ................................................................................................................220 Plagiochila balansae Gottsche ......................................................................................................239 Plagiochila baldwinii Austin ........................................................................................................239 Plagiochila bamingensis Steph. .....................................................................................................239 Plagiochila bancroftii Steph. ........................................................................................................239 Plagiochila banksiana Gottsche....................................................................................................223 Plagiochila banksiana var. echinophora Inoue et R.M.Schust. .......................................................223 Plagiochila bantamensis (Reinw., Blume et Nees) Mont...............................................................222 Plagiochila barbadensis Steph.......................................................................................................239 Plagiochila barbata Steph. ...........................................................................................................239 Plagiochila barbeyi Steph. ............................................................................................................239 Plagiochila barteri Mitt. ..............................................................................................................220 Plagiochila barteri var. valida (Steph.) Vanden Berghen ..............................................................220 Plagiochila batava Steph. .............................................................................................................239 Plagiochila baylisii Inoue et R.M.Schust. .....................................................................................231 Plagiochila bazzanioides J.J.Engel et G.L.Merr. ...........................................................................231 Plagiochila beddomei Steph. .........................................................................................................233 Plagiochila benitoi Inoue ex Piippo ..............................................................................................225 Plagiochila berggrenii Steph. ........................................................................................................239 Plagiochila beskeana Steph. ..........................................................................................................239 Plagiochila bialata Mitt. ..............................................................................................................239 Plagiochila biapiculata Steph. ......................................................................................................239 Plagiochila bicaudata Steph. ........................................................................................................239 Plagiochila biciliata Steph............................................................................................................239 Plagiochila bicornis Hampe et Gottsche .......................................................................................239 Plagiochila bicornuta Steph..........................................................................................................224 Plagiochila bicuspidata Gottsche ..................................................................................................230 Plagiochila bidens Gottsche .........................................................................................................220 Plagiochila bidentula Steph..........................................................................................................239 Plagiochila bifaria (Sw.) Lindenb. ...............................................................................................220 Plagiochila bifaria var. rosea (R.M.Schust.) Heinrichs .................................................................220 Plagiochila binghamiae A.Evans ...................................................................................................239 Plagiochila binominata Steph. .....................................................................................................239 Plagiochila biondiana C.Massal. ..................................................................................................229 Plagiochila bischleriana Grolle et M.L.So.....................................................................................228 Plagiochila bitexta Dugas .............................................................................................................239 Plagiochila blepharobasis Herzog ..................................................................................................239 Plagiochila blepharophora (Nees) Lindenb. ..................................................................................222 World checklist of hornworts and liverworts ** *** * * *** ** *** * ** ** *** *** *** ** * ** ** ** ** ** * *** * ** *** ** ** * ** *** ** ** *** ** ** ** *** * ** * *** ** *** ** ** * ** *** ** *** ** *** 617 Plagiochila bogotensis Gottsche ....................................................................................................239 Plagiochila boivinii Steph. ...........................................................................................................233 Plagiochila boliviana Spruce ........................................................................................................239 Plagiochila borneensis Steph. ........................................................................................................239 Plagiochila boryana Gottsche .......................................................................................................226 Plagiochila brassii Inoue et Grolle................................................................................................240 Plagiochila breuteliana Lindenb. ..................................................................................................226 Plagiochila brevicalycina Lindenb. et Gottsche.............................................................................240 Plagiochila britannica Paton ........................................................................................................228 Plagiochila brunneola Steph. ........................................................................................................240 Plagiochila bryhnii Steph. ............................................................................................................240 Plagiochila bryopteroides Spruce ...................................................................................................233 Plagiochila buchtiniana Steph. .....................................................................................................225 Plagiochila bunburii Taylor .........................................................................................................240 Plagiochila byssacea Hampe .........................................................................................................240 Plagiochila caducidentata R.M.Schust..........................................................................................240 Plagiochila caducifolia Inoue et R.M.Schust. ...............................................................................240 Plagiochila caduciloba H.L.Blomq. ..............................................................................................230 Plagiochila caldana Steph. ...........................................................................................................240 Plagiochila callaensis Steph. .........................................................................................................240 Plagiochila camusii Steph. ...........................................................................................................240 Plagiochila canelensis Steph. .........................................................................................................226 Plagiochila capillicaulis Steph. .....................................................................................................240 Plagiochila caribbeania R.M.Schust. ............................................................................................240 Plagiochila carringtonii (Balf. ex Carrington) Grolle ....................................................................229 Plagiochila carringtonii subsp. lobuchensis Grolle .........................................................................229 Plagiochila caulimammillosa Grolle et M.L.So .............................................................................227 Plagiochila cava Steph. ................................................................................................................240 Plagiochila ceylanica Mitt. ...........................................................................................................240 Plagiochila chacabucensis Steph. ...................................................................................................220 Plagiochila chacoënsis Herzog.......................................................................................................240 Plagiochila chauviniana Mont. ....................................................................................................222 Plagiochila chenii Grolle et M.L.So .............................................................................................231 Plagiochila chiloënsis Steph. .........................................................................................................240 Plagiochila chimborazensis Spruce ................................................................................................230 Plagiochila chinantlana Gottsche .................................................................................................240 Plagiochila chinensis Steph. ..........................................................................................................228 Plagiochila chiovendae Gola .........................................................................................................240 Plagiochila chonotica Taylor ........................................................................................................223 Plagiochila cinchonae Steph. ........................................................................................................240 Plagiochila circinalis (Lehm. et Lindenb.) Lehm. .........................................................................231 Plagiochila circinalis var. hemicardia (Hook.f. et Taylor) J.J.Engel et G.L.Merr. ..........................231 Plagiochila circumdentata Steph...................................................................................................228 Plagiochila circumdentata var. carinata J.J.Engel et G.L.Merr. .....................................................228 Plagiochila circumserrata Inoue et Grolle .....................................................................................228 Plagiochila circumvoluta Gerola ...................................................................................................240 Plagiochila clavatosaccata Steph. ..................................................................................................222 Plagiochila cleeii Inoue ...............................................................................................................240 Plagiochila cobana Steph. ............................................................................................................222 Plagiochila colensoi Hook.f. et Taylor ..........................................................................................231 Plagiochila colensoi var. quinquespina (Steph.) J.J.Engel et G.L.Merr. ..........................................231 Plagiochila colorans Steph. ...........................................................................................................220 618 ** * ** * ** ** ** ** ** ** *** ** *** ** ** * *** ** ** *** ** ** *** *** * *** * *** ** *** *** ** *** ** *** * *** ** ** ** ** ** *** *** ** ** ** *** *** * * *** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Plagiochila columbiana A.Evans ...................................................................................................228 Plagiochila concava Nees .............................................................................................................240 Plagiochila conduplicata Steph. ....................................................................................................240 Plagiochila conferta Steph. ...........................................................................................................240 Plagiochila confertissima Steph. ....................................................................................................240 Plagiochila connata Lindenb. et Gottsche ....................................................................................240 Plagiochila contigua Gottsche ......................................................................................................233 Plagiochila contorta Lindenb. et Hampe ......................................................................................240 Plagiochila convoluta Steph..........................................................................................................240 Plagiochila convolutifolia Steph. ...................................................................................................240 Plagiochila corrugata (Nees) Nees et Mont. .................................................................................233 Plagiochila corticola Steph. ..........................................................................................................231 Plagiochila corymbulosa Pearson...................................................................................................241 Plagiochila costariensis Horik. ......................................................................................................241 Plagiochila crispabilis Lindenb. ....................................................................................................241 Plagiochila crispabilis var. minima Lindenb. ................................................................................241 Plagiochila cristata (Sw.) Lindenb. ...............................................................................................220 Plagiochila cristophylla Steph. ......................................................................................................241 Plagiochila crozetensis Kaal...........................................................................................................224 Plagiochila cuatrecasii H.Rob. .....................................................................................................241 Plagiochila cubensis Steph. ...........................................................................................................241 Plagiochila cucullata Lindenb. et Gottsche...................................................................................241 Plagiochila cucullifolia J.B.Jack et Steph. .....................................................................................226 Plagiochila cucullifolia var. anomala Heinrichs et Gradst. ............................................................226 Plagiochila cuervina Gottsche ......................................................................................................241 Plagiochila cumingiana Steph. .....................................................................................................221 Plagiochila cuneata Lindenb. et Gottsche ....................................................................................230 Plagiochila cuspidata Steph. .........................................................................................................233 Plagiochila cymata Inoue et Grolle...............................................................................................233 Plagiochila debilis Mitt. ...............................................................................................................230 Plagiochila delexa Mont. et Gottsche ..........................................................................................230 Plagiochila delexirama Taylor .....................................................................................................233 Plagiochila defolians Grolle et M.L.So .........................................................................................230 Plagiochila delapsa Inoue .............................................................................................................241 Plagiochila delavayi Steph. ...........................................................................................................228 Plagiochila delognei Steph. ...........................................................................................................241 Plagiochila deltoidea Lindenb. .....................................................................................................224 Plagiochila deltoidea var. densa J.J.Engel et G.L.Merr. .................................................................224 Plagiochila denigrata Inoue..........................................................................................................241 Plagiochila densa Herzog .............................................................................................................241 Plagiochila densilora Herzog .......................................................................................................241 Plagiochila densiramosa Steph. .....................................................................................................241 Plagiochila denticulata Mitt. ........................................................................................................222 Plagiochila dependula Taylor .......................................................................................................225 Plagiochila deppeana Steph. .........................................................................................................241 Plagiochila desciscens Steph. .........................................................................................................241 Plagiochila detecta M.L.So et Grolle ............................................................................................222 Plagiochila devexa Steph. .............................................................................................................227 Plagiochila dichotoma (P.Beauv.) Nees et Mont. ..........................................................................233 Plagiochila dichotoma var. luitans Spruce ....................................................................................233 Plagiochila dilatata Steph. ...........................................................................................................241 Plagiochila dimorpha Lindenb. et Gottsche .................................................................................226 World checklist of hornworts and liverworts *** * *** ** ** *** *** * ** *** *** * ** *** ** ** ** *** ** ** * *** *** * * ** ** ** * *** * * *** *** ** *** ** * ** ** * ** *** * ** ** ** * ** ** ** ** 619 Plagiochila dissecta Steph. ............................................................................................................233 Plagiochila distans Colenso ..........................................................................................................241 Plagiochila disticha (Lehm. et Lindenb.) Lindenb. .......................................................................234 Plagiochila distinctifolia Lindenb. ................................................................................................234 Plagiochila divaricata Lindenb.....................................................................................................241 Plagiochila divergens Steph. .........................................................................................................234 Plagiochila diversifolia Lindenb. et Gottsche ................................................................................225 Plagiochila doerleri Steph. ...........................................................................................................241 Plagiochila dolichoblasta Herzog ..................................................................................................241 Plagiochila dominicensis Taylor ....................................................................................................226 Plagiochila drepanophylla Sande Lac. ...........................................................................................234 Plagiochila drepanophylla var. minor Gottsche .............................................................................234 Plagiochila dura De Not..............................................................................................................224 Plagiochila durelii Schifn. ...........................................................................................................227 Plagiochila durelii subsp. guizhouensis Grolle et M.L.So ..............................................................227 Plagiochila dusenii Steph. ............................................................................................................224 Plagiochila dussiana Steph. ..........................................................................................................241 Plagiochila duthiana Steph. .........................................................................................................229 Plagiochila echinata R.M.Schust. .................................................................................................241 Plagiochila echinella Gottsche ......................................................................................................241 Plagiochila ecuadorensis Steph. .....................................................................................................241 Plagiochila ecuadorica (Inoue) L.Söderstr.....................................................................................226 Plagiochila efusa Steph................................................................................................................234 Plagiochila efusa var. decurrens Steph. .........................................................................................234 Plagiochila efuseintricata Steph. ..................................................................................................242 Plagiochila eggersii Inoue .............................................................................................................242 Plagiochila ekmanii S.W.Arnell ...................................................................................................242 Plagiochila elata Taylor ...............................................................................................................224 Plagiochila electa Steph. ...............................................................................................................242 Plagiochila elegans Mitt. ..............................................................................................................228 Plagiochila elegantula Herzog ......................................................................................................242 Plagiochila emarginatobidentula Steph. ........................................................................................242 Plagiochila emeiensis Grolle et M.L.So .........................................................................................220 Plagiochila ensiformis Taylor ........................................................................................................226 Plagiochila equitans Gottsche ......................................................................................................223 Plagiochila ericicola Steph. ...........................................................................................................234 Plagiochila erlangensis M.L.So .....................................................................................................229 Plagiochila erythraeae Herzog ......................................................................................................242 Plagiochila estrellensis Herzog ......................................................................................................242 Plagiochila eurydictyon Herzog ....................................................................................................242 Plagiochila excisa Steph. ..............................................................................................................242 Plagiochila exesa Lindenb. et Gottsche.........................................................................................242 Plagiochila exigua (Taylor) Taylor ...............................................................................................230 Plagiochila exilis Colenso .............................................................................................................242 Plagiochila exinnovata Steph........................................................................................................234 Plagiochila expansa Gottsche .......................................................................................................242 Plagiochila facallonia Steph. ........................................................................................................242 Plagiochila falcato-oblonga Steph. ................................................................................................242 Plagiochila fallax Lindenb. et Hampe ..........................................................................................242 Plagiochila fasciculata Lindenb. ...................................................................................................231 Plagiochila fastigiata Lindenb. et Gottsche ..................................................................................234 Plagiochila faxinensis Schifn. ......................................................................................................242 620 ** * ** *** *** * ** ** * ** *** ** ** *** * ** * *** ** * ** * * ** *** *** *** *** ** ** *** *** * ** ** * ** ** *** * *** ** ** * * *** ** * *** ** ** *** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Plagiochila fendleri Mont.............................................................................................................242 Plagiochila ilicicola Steph. ..........................................................................................................242 Plagiochila issidentoides Taylor....................................................................................................242 Plagiochila labellata Steph. .........................................................................................................234 Plagiochila labellilora Steph. ......................................................................................................226 Plagiochila labellifrons Spruce .....................................................................................................242 Plagiochila lava Steph.................................................................................................................242 Plagiochila lavescens (Gottsche, Lindenb. et Nees) Gottsche .......................................................242 Plagiochila lavorufescens Steph. ...................................................................................................242 Plagiochila lexicaulis Mont. ........................................................................................................225 Plagiochila lexuosa Mitt. .............................................................................................................232 Plagiochila loridana A.Evans.......................................................................................................234 Plagiochila footei A.Evans ............................................................................................................242 Plagiochila fordiana Steph. ..........................................................................................................234 Plagiochila formosa Nees..............................................................................................................242 Plagiochila fracta Pócs .................................................................................................................230 Plagiochila fragilis Taylor ............................................................................................................221 Plagiochila fragillima Steph. ........................................................................................................222 Plagiochila fragmentata R.M.Schust. ...........................................................................................243 Plagiochila fragmentissima Inoue et R.M.Schust. .........................................................................223 Plagiochila francana Steph. ..........................................................................................................234 Plagiochila frausa Gottsche ..........................................................................................................243 Plagiochila frausa var. boliviana Spruce .......................................................................................243 Plagiochila frayjorgensis Hässel .....................................................................................................243 Plagiochila frondescens (Nees) Lindenb. .......................................................................................225 Plagiochila fruticosa Mitt. ............................................................................................................225 Plagiochila furcifolia Mitt. ...........................................................................................................234 Plagiochila fusca Sande Lac. .........................................................................................................231 Plagiochila fuscella (Hook.f. et Taylor) Gottsche, Lindenb. et Nees .............................................231 Plagiochila fuscella var. novae-zelandiae (E.A.Hodgs.) J.J.Engel et G.L.Merr................................231 Plagiochila fuscolutea Taylor ........................................................................................................225 Plagiochila fusifera Taylor............................................................................................................234 Plagiochila gafatensis Gottsche ex Schweinf. ...............................................................................243 Plagiochila gaudichaudii Mont. et Gottsche.................................................................................243 Plagiochila geniculata Lindenb.....................................................................................................243 Plagiochila geppii Steph. ..............................................................................................................243 Plagiochila germana Gottsche ......................................................................................................243 Plagiochila germanii Steph............................................................................................................24, Plagiochila ghatiensis Steph. .........................................................................................................230 Plagiochila gibbilora Steph..........................................................................................................243 Plagiochila gigantea Lindenb. ......................................................................................................223 Plagiochila gigantea var. inermis J.J.Engel et G.L.Merr. ...............................................................223 Plagiochila gittinsii Inoue ............................................................................................................243 Plagiochila glauca Carl.................................................................................................................243 Plagiochila gracilicaulis Spruce.....................................................................................................243 Plagiochila gracilis Lindenb. et Gottsche......................................................................................232 Plagiochila gracillima Austin........................................................................................................243 Plagiochila granatensis Gottsche...................................................................................................243 Plagiochila grandicrista Steph. .....................................................................................................220 Plagiochila granditexta Steph. ......................................................................................................243 Plagiochila grateloupii Mont. .......................................................................................................243 Plagiochila gregaria (Hook.f. et Taylor) Gottsche, Lindenb. et Nees............................................223 World checklist of hornworts and liverworts ** *** ** ** ** ** *** ** ** *** ** *** *** * ** ** ** *** ** ** * *** ** ** * * * *** ** ** ** * ** ** * ** * ** ** ** ** * ** ** *** ** * ** * ** ** *** 621 Plagiochila grollei Inoue...............................................................................................................227 Plagiochila grossa Grolle et M.L.So ..............................................................................................230 Plagiochila grossispina Steph. .......................................................................................................222 Plagiochila guatemalensis Steph. ...................................................................................................243 Plagiochila guevarii H.Rob. .........................................................................................................226 Plagiochila guttisquama Inoue et Grolle .......................................................................................243 Plagiochila gymnocalycina (Lehm. et Lindenb.) Mont. et Nees.....................................................230 Plagiochila gymnocalycina var. surinamensis (Molk.) Heinrichs et D.S.Rycroft .............................230 Plagiochila gymnocalyx Inoue .......................................................................................................243 Plagiochila gymnoclada Sande Lac................................................................................................232 Plagiochila haeseliae Inoue ...........................................................................................................243 Plagiochila hakkodensis Steph. .....................................................................................................228 Plagiochila hampeana Gottsche ...................................................................................................234 Plagiochila hans-meyeri Steph. .....................................................................................................243 Plagiochila harlingii S.W.Arnell ..................................................................................................243 Plagiochila haumanii Herzog .......................................................................................................244 Plagiochila hawaica Steph. ..........................................................................................................244 Plagiochila herminieri Steph. .......................................................................................................220 Plagiochila heteracantha Steph. ....................................................................................................244 Plagiochila heterodonta (Hook.f. et Taylor) Gottsche, Lindenb. et Nees ......................................224 Plagiochila heterofolia Steph. .......................................................................................................244 Plagiochila heteromalla (Lehm. et Lindenb.) Lindenb. .................................................................220 Plagiochila heterophylla Lindenb. .................................................................................................225 Plagiochila heterophylla var. beauverdii (Steph.) Heinrichs ...........................................................225 Plagiochila heterospina Steph. ......................................................................................................234 Plagiochila heterostipa Steph. .......................................................................................................234 Plagiochila hieronymii Steph. .......................................................................................................244 Plagiochila himalayana Schifn. ...................................................................................................232 Plagiochila hiroshiana Pócs ..........................................................................................................244 Plagiochila hoehnei Herzog ..........................................................................................................244 Plagiochila hoei Inoue..................................................................................................................227 Plagiochila holstii Steph. ..............................................................................................................244 Plagiochila hookeriana Lindenb. ..................................................................................................223 Plagiochila horrida Gottsche ........................................................................................................244 Plagiochila huatuscana Gottsche ..................................................................................................244 Plagiochila huerlimannii Inoue ....................................................................................................244 Plagiochila humboldtiana Gottsche ..............................................................................................244 Plagiochila husnotii Steph. ...........................................................................................................226 Plagiochila hyalina Lindenb.........................................................................................................224 Plagiochila hyalodermica Grolle et M.L.So ...................................................................................227 Plagiochila incerta Gottsche.........................................................................................................234 Plagiochila incisa Dugas ..............................................................................................................244 Plagiochila incurvicolla (Hook.f. et Taylor) Gottsche, Lindenb. et Nees ......................................232 Plagiochila incurvicolla var. lonchoscypha (Herzog) J.J.Engel et G.L.Merr. ...................................232 Plagiochila indica Mitt. ex Steph. ................................................................................................234 Plagiochila inlata Steph. .............................................................................................................244 Plagiochila informifolia Steph. .....................................................................................................244 Plagiochila infuscata Mitt. ...........................................................................................................244 Plagiochila injasutiensis S.W.Arnell..............................................................................................244 Plagiochila inouei Grolle..............................................................................................................244 Plagiochila insularia Mitt. ...........................................................................................................244 Plagiochila integerrima Steph. ......................................................................................................222 622 * ** * ** ** ** ** ** ** *** ** *** * * ** ** *** ** ** ** ** ** * *** ** ** * * * *** * ** ** *** ** ** ** * ** * * * ** ** * * * ** * * ** *** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Plagiochila integrilobula Schifn. ..................................................................................................222 Plagiochila intertexta Hook.f. et Taylor .......................................................................................244 Plagiochila inuensis Steph. ...........................................................................................................244 Plagiochila invisa (R.M.Schust.) R.M.Schust. .............................................................................244 Plagiochila irregularis Gottsche ....................................................................................................244 Plagiochila itatiajensis Steph. .......................................................................................................244 Plagiochila jacquinotii Mont........................................................................................................244 Plagiochila jamaicensis Lindenb. et Hampe..................................................................................244 Plagiochila jaramillii H.Rob. .......................................................................................................244 Plagiochila javanica (Sw.) Nees et Mont......................................................................................234 Plagiochila johannis-winkleri Herzog ...........................................................................................222 Plagiochila junghuhniana Sande Lac. ...........................................................................................235 Plagiochila karstenii Steph. ..........................................................................................................244 Plagiochila kaulfussiana Steph. ....................................................................................................244 Plagiochila keckiana Steph. ..........................................................................................................244 Plagiochila kerneriana S.W.Arnell ...............................................................................................245 Plagiochila khasiana Mitt. ...........................................................................................................235 Plagiochila kiaeri Gottsche ..........................................................................................................235 Plagiochila kiaeri var. capensis (Steph.) M.Wigginton et Grolle ...................................................235 Plagiochila kiaeri var. myriocarpa (Pearson) Pócs .........................................................................235 Plagiochila koponenii Inoue et Piippo ..........................................................................................245 Plagiochila korthalsiana Molk. .....................................................................................................228 Plagiochila kuhliana Sande Lac....................................................................................................228 Plagiochila kunmingensis Piippo ..................................................................................................235 Plagiochila kurokawae Inoue .......................................................................................................235 Plagiochila kurzii Steph. ..............................................................................................................223 Plagiochila lacerifolia Steph. ........................................................................................................245 Plagiochila lachenaudii Steph. .....................................................................................................245 Plagiochila laciniosa Dugas ..........................................................................................................245 Plagiochila laetevirens Lindenb. ...................................................................................................245 Plagiochila laevifolia Gola............................................................................................................245 Plagiochila lamellistipula Spruce ..................................................................................................235 Plagiochila lansbergii Gottsche.....................................................................................................245 Plagiochila lastii Mitt. .................................................................................................................235 Plagiochila latifolia Steph. ...........................................................................................................235 Plagiochila latifrons Gottsche et Hampe ......................................................................................224 Plagiochila latitrigona Schifn. .....................................................................................................245 Plagiochila laxiramea Steph. ........................................................................................................245 Plagiochila lechleri Gottsche ........................................................................................................225 Plagiochila lecontei Steph. ............................................................................................................245 Plagiochila ledermanniana Beauverd ............................................................................................245 Plagiochila ledieui Steph. .............................................................................................................245 Plagiochila lehmanniana S.W.Arnell ............................................................................................245 Plagiochila liebmanniana Lehm. et Lindenb. ...............................................................................245 Plagiochila lignicola Spruce..........................................................................................................245 Plagiochila lindaui Steph. ............................................................................................................245 Plagiochila lindigiana Gottsche ...................................................................................................245 Plagiochila lingua Steph. .............................................................................................................245 Plagiochila loloënsis Steph. ...........................................................................................................235 Plagiochila longiissa Steph...........................................................................................................245 Plagiochila longilora Mont. .........................................................................................................227 Plagiochila longiramea Steph. ......................................................................................................225 World checklist of hornworts and liverworts *** ** ** ** * *** * ** ** * ** * *** ** ** ** *** ** ** * ** ** ** ** ** ** ** ** ** ** * ** ** ** ** ** ** * ** ** ** ** *** ** ** * ** ** ** * *** ** 623 Plagiochila longispina Lindenb. et Gottsche .................................................................................225 Plagiochila lophocoleoides Mont. ..................................................................................................227 Plagiochila loriana Steph. ............................................................................................................228 Plagiochila loriloba Herzog ex Carl ..............................................................................................230 Plagiochila lotsyana Steph. ...........................................................................................................245 Plagiochila lunata S.W.Arnell ......................................................................................................221 Plagiochila luteola Steph. .............................................................................................................245 Plagiochila lutescens Steph. ..........................................................................................................245 Plagiochila luzonensis Grolle et M.L.So .......................................................................................245 Plagiochila macra Taylor .............................................................................................................246 Plagiochila macrifolia Taylor .......................................................................................................246 Plagiochila macrifolia var. angustifolia Spruce ..............................................................................246 Plagiochila macrostachya Lindenb. ...............................................................................................226 Plagiochila maderensis Gottsche ex Steph.....................................................................................230 Plagiochila magna Inoue..............................................................................................................227 Plagiochila manillana Mont. et Gottsche.....................................................................................235 Plagiochila martiana (Nees) Lindenb...........................................................................................246 Plagiochila massalongoana Schifn. ...............................................................................................235 Plagiochila mastigophoroides Inoue ...............................................................................................235 Plagiochila matanga Steph. ..........................................................................................................246 Plagiochila mauiensis Steph. ........................................................................................................225 Plagiochila maunakeana Steph.....................................................................................................246 Plagiochila maximiliana Gottsche................................................................................................246 Plagiochila meghalayensis K.K.Rawat et S.C.Srivast. ....................................................................246 Plagiochila meridana Gottsche ....................................................................................................246 Plagiochila metcalii Steph. ..........................................................................................................233 Plagiochila microdentata M.L.So .................................................................................................228 Plagiochila microdictyon Mitt.......................................................................................................232 Plagiochila microdonta Mitt. ........................................................................................................246 Plagiochila micropteryx Gottsche..................................................................................................235 Plagiochila mildbreadiana Beauverd ............................................................................................247 Plagiochila minarum Herzog .......................................................................................................247 Plagiochila minutiretis Reimers ....................................................................................................247 Plagiochila minutula (Hook.f. et Taylor) Gottsche, Lindenb. et Nees..........................................224 Plagiochila miqueliana Lehm. et Lindenb. ...................................................................................246 Plagiochila miradorensis Gottsche ................................................................................................235 Plagiochila miradorensis var. convoluta R.M.Schust. .....................................................................235 Plagiochila moenkemeyeri Steph. ..................................................................................................235 Plagiochila molliuscula Inoue .......................................................................................................246 Plagiochila mollusca Lehm. ..........................................................................................................246 Plagiochila moniliformis R.M.Schust. ..........................................................................................246 Plagiochila monospiris Inoue et Grolle .........................................................................................232 Plagiochila montagnei Nees..........................................................................................................235 Plagiochila morobensis Inoue et Piippo.........................................................................................236 Plagiochila muelleriana Gottsche .................................................................................................246 Plagiochila multilora Steph. ........................................................................................................246 Plagiochila multipinnula Herzog et S.Hatt. .................................................................................236 Plagiochila mundaliensis Steph.....................................................................................................228 Plagiochila mutila Gottsche .........................................................................................................246 Plagiochila naranjoënsis Steph......................................................................................................246 Plagiochila neckeroidea Mitt. .......................................................................................................236 Plagiochila neesiana Lindenb. ......................................................................................................246 624 * * * *** * *** *** ** * * ** ** 7* ** ** *** ** ** ** ** * *** *** * * * *** * ** ** ** ** ** ** *** ** *** *** * ** * * ** *** *** *** ** ** ** ** ** * Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Plagiochila neglecta Steph. ...........................................................................................................246 Plagiochila negrensis Spruce .........................................................................................................246 Plagiochila negrii Gola.................................................................................................................246 Plagiochila nepalensis Lindenb. ....................................................................................................236 Plagiochila nigricaulis Steph. .......................................................................................................247 Plagiochila nitens Inoue ...............................................................................................................232 Plagiochila nobilis Gottsche .........................................................................................................224 Plagiochila norfolkiensis Steph......................................................................................................236 Plagiochila notha Steph. ..............................................................................................................247 Plagiochila nova Steph.................................................................................................................247 Plagiochila nudifolia (Steph.) Grolle ............................................................................................247 Plagiochila nutans Steph. .............................................................................................................247 Plagiochila obliquetruncata Steph. ...............................................................................................247 Plagiochila oblonga Inoue ............................................................................................................247 Plagiochila obovata Steph. ...........................................................................................................224 Plagiochila obtusa Lindenb. .........................................................................................................236 Plagiochila oerstediana Lindenb. et Hampe..................................................................................247 Plagiochila olivacea Steph. ...........................................................................................................247 Plagiochila orbicularis (S.Hatt.) S.Hatt. .......................................................................................228 Plagiochila oresitropha Spruce ......................................................................................................230 Plagiochila ornata Wilson ex Lindenb. ........................................................................................247 Plagiochila ovalifolia Mitt. ...........................................................................................................228 Plagiochila ovata Lindenb. et Gottsche ........................................................................................224 Plagiochila ovato-obconica Steph. .................................................................................................247 Plagiochila ovifolia Steph. ............................................................................................................247 Plagiochila owaihiensis Nees et Lindenb. .....................................................................................234 Plagiochila pachyloma Taylor .......................................................................................................221 Plagiochila pachyloma var. elatior Spruce .....................................................................................221 Plagiochila paciica Mitt. .............................................................................................................234 Plagiochila palangiensis S.C.Srivast., K.K.Rawat et P.K.Verma ....................................................247 Plagiochila panamensis Inoue .......................................................................................................247 Plagiochila papillifolia Steph. .......................................................................................................221 Plagiochila parallela Steph. ..........................................................................................................236 Plagiochila paramicola Herzog .....................................................................................................247 Plagiochila paraphyllina Herzog...................................................................................................225 Plagiochila paraphyllosa Grolle et M.L.So ....................................................................................227 Plagiochila parvifolia Lindenb. ....................................................................................................234 Plagiochila parviramifera Inoue ...................................................................................................221 Plagiochila parvitexta Steph. ........................................................................................................247 Plagiochila parvivittata Inoue ......................................................................................................247 Plagiochila parvula Steph. ...........................................................................................................247 Plagiochila pastasensis Steph.........................................................................................................247 Plagiochila patentispina Steph. .....................................................................................................247 Plagiochila patentissima Lindenb. ................................................................................................247 Plagiochila patriciae Heinrichs et H.Anton ..................................................................................226 Plagiochila patula (Sw.) Lindenb. ................................................................................................236 Plagiochila patula var. brevifolia Gottsche....................................................................................236 Plagiochila patuloides Schifn. ......................................................................................................247 Plagiochila paucidens Steph. ........................................................................................................236 Plagiochila paucidentata Mont. et Gottsche .................................................................................247 Plagiochila paupercula Gottsche...................................................................................................247 Plagiochila pearceana Steph. ........................................................................................................247 World checklist of hornworts and liverworts *** ** ** ** ** ** ** ** *** ** ** ** *** ** ** *** ** *** ** ** *** ** * *** ** *** * *** ** *** ** ** *** ** *** ** *** ** * *** ** *** ** ** ** *** *** *** ** *** *** *** 625 Plagiochila pectinata Lindenb. .....................................................................................................230 Plagiochila peculiaris Schifn. .......................................................................................................227 Plagiochila pellucida Lindenb. et Gottsche...................................................................................247 Plagiochila pensilis Spruce............................................................................................................236 Plagiochila peradenyensis Schifn. .................................................................................................236 Plagiochila perdentata M.L.So et Grolle.......................................................................................236 Plagiochila perrottetiana Mont. et Gottsche .................................................................................248 Plagiochila perrottetiana var. minor Gottsche ...............................................................................248 Plagiochila perserrata Herzog .......................................................................................................227 Plagiochila philippinensis Steph....................................................................................................227 Plagiochila pilifera Steph. ............................................................................................................248 Plagiochila pinnatidens Steph.......................................................................................................248 Plagiochila pinnilora Steph. ........................................................................................................236 Plagiochila pittieri Steph..............................................................................................................248 Plagiochila platyphylla Herzog .....................................................................................................248 Plagiochila poeltii Inoue et Grolle ................................................................................................229 Plagiochila polopolensis Herzog ....................................................................................................248 Plagiochila porelloides (Torr. ex Nees) Lindenb. ...........................................................................229 Plagiochila porelloides var. norvegica (H.H.Blom et Holten) Schumacker et Váňa ........................229 Plagiochila porelloides var. subarctica (Jørg.) Lammes ...................................................................229 Plagiochila praemorsa Steph. ........................................................................................................236 Plagiochila propinqua Sande Lac..................................................................................................236 Plagiochila prostrata Steph. ..........................................................................................................244 Plagiochila pseudoattenuata S.W.Arnell........................................................................................221 Plagiochila pseudocapillaris Inoue.................................................................................................232 Plagiochila pseudoirma Herzog ...................................................................................................232 Plagiochila pseudopatula Herzog ..................................................................................................248 Plagiochila pseudopoeltii Inoue .....................................................................................................227 Plagiochila pseudoradicans Herzog ...............................................................................................248 Plagiochila pseudorenitens Schifn. ...............................................................................................222 Plagiochila ptychanthoidea Steph..................................................................................................248 Plagiochila pulchella Steph...........................................................................................................248 Plagiochila pulcherrima Horik. ....................................................................................................225 Plagiochila pulvinata Steph..........................................................................................................248 Plagiochila punctata (Taylor) Taylor............................................................................................221 Plagiochila purpurascens Steph. ....................................................................................................236 Plagiochila raddiana Lindenb. .....................................................................................................236 Plagiochila radiculosa Mitt...........................................................................................................232 Plagiochila ragazzii Gola .............................................................................................................236 Plagiochila ramosissima (Hook.) Lindenb. ...................................................................................224 Plagiochila ratkowskiana Inoue ....................................................................................................248 Plagiochila recurvata (W.E.Nicholson) Grolle .............................................................................229 Plagiochila regeliana Steph. .........................................................................................................248 Plagiochila reischeckiana Steph. ...................................................................................................223 Plagiochila remyana Steph. ..........................................................................................................248 Plagiochila renauldii Steph. .........................................................................................................221 Plagiochila renitens (Nees) Lindenb. ............................................................................................227 Plagiochila repanda (Schwägr.) Lindenb. .....................................................................................236 Plagiochila repanda var. perrotana (Steph.) Vanden Berghen .......................................................237 Plagiochila retrorsa Gottsche........................................................................................................221 Plagiochila retrospectans Lindenb. ................................................................................................224 Plagiochila retusa Mitt. ................................................................................................................229 626 *** ** ** *** ** *** * *** *** * ** * *** *** *** ** * ** *** ** ** ** *** * *** ** * * * ** * ** ** *** ** ** * *** ** ** ** ** ** *** *** ** * *** *** ** * ** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Plagiochila revolvens Mitt. ...........................................................................................................248 Plagiochila rigidula Lindenb. et Gottsche ....................................................................................248 Plagiochila riparia Steph. ............................................................................................................224 Plagiochila rodriguezii Steph. .......................................................................................................237 Plagiochila rosana Steph. .............................................................................................................248 Plagiochila rubescens (Lehm. et Lindenb.) Lindenb. .....................................................................221 Plagiochila rubricaulis Steph. .......................................................................................................221 Plagiochila rudischusteri H.Rob. ..................................................................................................225 Plagiochila rudolii Pócs...............................................................................................................237 Plagiochila ruifolia Steph. ...........................................................................................................248 Plagiochila rufoviridis Spruce .......................................................................................................248 Plagiochila rusbyi Spruce .............................................................................................................248 Plagiochila rutilans Lindenb. .......................................................................................................230 Plagiochila rutilans var. moritziana (Lindenb. et Gottsche) Heinrichs..........................................230 Plagiochila rutilans var. standleyi (Herzog ex Carl) Heinrichs et D.S.Rycroft ...............................230 Plagiochila rutlandii Steph. .........................................................................................................224 Plagiochila sabensis Steph. ...........................................................................................................248 Plagiochila sachapatensis Steph.....................................................................................................248 Plagiochila salacensis Gottsche .....................................................................................................237 Plagiochila salazariae Inoue .........................................................................................................248 Plagiochila saltuensis Spruce ex Steph. .........................................................................................248 Plagiochila saltuensis var. spinosissima Herzog ..............................................................................249 Plagiochila salvadorica Steph. ......................................................................................................237 Plagiochila samoana Steph. ..........................................................................................................249 Plagiochila sandei Dozy ex Sande Lac. .........................................................................................223 Plagiochila sarmentosa (Lehm. et Lindenb.) Lindenb. ..................................................................249 Plagiochila schifneri Steph. .........................................................................................................249 Plagiochila schinzei Steph. ...........................................................................................................249 Plagiochila schmidtii Steph. .........................................................................................................249 Plagiochila schoieldiana Inoue .....................................................................................................229 Plagiochila schraderbergii Steph. ..................................................................................................249 Plagiochila schubertiana Steph. ....................................................................................................249 Plagiochila schusteri Inoue et Grolle.............................................................................................249 Plagiochila sciophila Nees ............................................................................................................223 Plagiochila sciophila subsp. ciliigera (R.M.Schust.) L.Söderstr. .....................................................223 Plagiochila scissifolia Steph...........................................................................................................249 Plagiochila scotica Macvicar ex Steph. ..........................................................................................249 Plagiochila secretifolia Mitt. .........................................................................................................229 Plagiochila semiamplexicaulis Steph. ............................................................................................249 Plagiochila semidecurrens (Lehm. et Lindenb.) Lindenb. ..............................................................227 Plagiochila semidecurrens var. alaskana (A.Evans) Inoue...............................................................227 Plagiochila semiermis Dugas ........................................................................................................249 Plagiochila serrialata Herzog........................................................................................................237 Plagiochila shangaica Steph..........................................................................................................237 Plagiochila sichuanensis Grolle et M.L.So ....................................................................................221 Plagiochila silvatica Gottsche .......................................................................................................249 Plagiochila similis Steph. .............................................................................................................249 Plagiochila simplex (Sw.) Lindenb. ..............................................................................................237 Plagiochila singularis Schifn. .......................................................................................................222 Plagiochila sisparensis Steph. ........................................................................................................249 Plagiochila slateri Steph. ..............................................................................................................249 Plagiochila solitaria Gottsche .......................................................................................................249 World checklist of hornworts and liverworts ** *** ** *** * *** ** ** * * *** *** * ** ** * ** *** *** ** *** ** * * ** ** ** * ** ** ** ** ** * * *** ** ** ** ** *** *** * ** * ** *** *** ** ** ** ** 627 Plagiochila spathulifolia Mitt. ......................................................................................................232 Plagiochila spinifera Ångstr. .........................................................................................................249 Plagiochila spinosa M.L.So ..........................................................................................................229 Plagiochila spinulosa (Dicks.) Dumort. ........................................................................................221 Plagiochila sprucei Steph. .............................................................................................................249 Plagiochila squamulosa Mitt. .......................................................................................................237 Plagiochila squamulosa var. crispulocaudata (Gottsche) Vanden Berghen......................................237 Plagiochila squamulosa var. sinuosa (Mitt.) Vanden Berghen ........................................................237 Plagiochila staudtiana Steph. .......................................................................................................249 Plagiochila stephanii Schifn. .......................................................................................................223 Plagiochila stephensoniana Mitt....................................................................................................232 Plagiochila stevensiana Steph. ......................................................................................................222 Plagiochila steyermarkii H.Rob. ...................................................................................................231 Plagiochila sticticola Mont. et Gottsche .......................................................................................221 Plagiochila stipata Steph. .............................................................................................................249 Plagiochila stolzii Steph. ..............................................................................................................249 Plagiochila streimannii Inoue.......................................................................................................237 Plagiochila stricta Lindenb...........................................................................................................221 Plagiochila strictifolia Steph. ........................................................................................................237 Plagiochila strombifolia Taylor .....................................................................................................231 Plagiochila subbidentata Taylor ...................................................................................................225 Plagiochila subcontigua Herzog ....................................................................................................249 Plagiochila subconvoluta Gottsche................................................................................................249 Plagiochila subdenudata Steph. ....................................................................................................249 Plagiochila subedentata Steph. .....................................................................................................250 Plagiochila sublabellata Colenso .................................................................................................237 Plagiochila subfragilis Inoue .........................................................................................................250 Plagiochila subhyalina Steph. .......................................................................................................250 Plagiochila subjavanica M.L.So ...................................................................................................250 Plagiochila subligulata Steph........................................................................................................250 Plagiochila sublyallii M.L.So........................................................................................................250 Plagiochila subpectinata Besch. et C.Massal. ................................................................................224 Plagiochila subplana Lindenb. .....................................................................................................223 Plagiochila subrara Herzog ..........................................................................................................250 Plagiochila subrotundifolia Steph. ................................................................................................250 Plagiochila subtropica Steph. ........................................................................................................237 Plagiochila subundulata Lindenb. ................................................................................................250 Plagiochila sullivantii Gottsche ....................................................................................................232 Plagiochila sullivantii var. spinigera R.M.Schust. .........................................................................232 Plagiochila sumatrana Schifn. .....................................................................................................229 Plagiochila superba (Nees ex Spreng.) Mont. et Nees ...................................................................226 Plagiochila superba var. macrotricha (Spruce) Heinrichs ...............................................................226 Plagiochila suringarii Steph. ........................................................................................................250 Plagiochila sylvicultrix Spruce ......................................................................................................250 Plagiochila symmetrica Steph. ......................................................................................................250 Plagiochila tabinana Gottsche .....................................................................................................250 Plagiochila tabinensis Steph. ........................................................................................................225 Plagiochila tagawae Inoue............................................................................................................222 Plagiochila taiwanensis Inoue.......................................................................................................229 Plagiochila talinayi S.W.Arnell ....................................................................................................250 Plagiochila tamariscina Steph. .....................................................................................................237 Plagiochila tambillensis Loitl. .......................................................................................................250 628 ** ** ** * ** * *** *** ** * ** ** ** ** * ** * *** * * *** ** ** *** * ** ** ** *** ** *** ** ** ** *** ** ** * ** ** * ** ** ** ** * * * * *** * *** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Plagiochila tamiensis Steph. .........................................................................................................250 Plagiochila tarapotensis Steph.......................................................................................................250 Plagiochila tecta Inoue et Grolle ..................................................................................................237 Plagiochila tenera Steph. ..............................................................................................................250 Plagiochila tenuis Lindenb. ..........................................................................................................250 Plagiochila tenuispica Steph. ........................................................................................................250 Plagiochila terebrans Nees et Mont. .............................................................................................237 Plagiochila teysmannii Sande Lac. ................................................................................................237 Plagiochila thamniopsis Spruce.....................................................................................................250 Plagiochila thollonii Steph. ..........................................................................................................250 Plagiochila thrausta Inoue et Grolle .............................................................................................250 Plagiochila thyoides Spruce...........................................................................................................237 Plagiochila tixieri Inoue ...............................................................................................................222 Plagiochila tocarema Gottsche .....................................................................................................237 Plagiochila tonduzana Steph. .......................................................................................................250 Plagiochila tortuosa Lindenb. et Gottsche ....................................................................................250 Plagiochila tovarina Gottsche ......................................................................................................250 Plagiochila trabeculata Steph. ......................................................................................................232 Plagiochila trabeculatospinosa Herzog ..........................................................................................250 Plagiochila trabutii Steph. ...........................................................................................................250 Plagiochila trapezoidea Lindenb. ..................................................................................................229 Plagiochila trianae Gottsche ........................................................................................................250 Plagiochila trichomanes Spruce ....................................................................................................250 Plagiochila trichostoma Gottsche ..................................................................................................231 Plagiochila tricuspis Steph. ...........................................................................................................250 Plagiochila tristaniana Váňa et J.J.Engel ......................................................................................251 Plagiochila tristis Steph. ...............................................................................................................251 Plagiochila trollii Herzog .............................................................................................................251 Plagiochila tronadoris Herzog ......................................................................................................221 Plagiochila truncata Gottsche ......................................................................................................251 Plagiochila turgida Herzog ..........................................................................................................226 Plagiochila ulata Inoue et Grolle .................................................................................................237 Plagiochila uleana Steph. .............................................................................................................251 Plagiochila umbrosioides L.Söderstr. .............................................................................................251 Plagiochila uncialis (Hook.f. et Taylor) Gottsche, Lindenb. et Nees ............................................221 Plagiochila undata Sull. ...............................................................................................................237 Plagiochila undata subsp. crispata (Gottsche) R.M.Schust. ..........................................................237 Plagiochila unduavensis Steph. .....................................................................................................251 Plagiochila ungarangana Sande Lac. ............................................................................................238 Plagiochila uniformis Mitt. ..........................................................................................................229 Plagiochila usambarana Steph. ....................................................................................................251 Plagiochila validissima Steph. ......................................................................................................225 Plagiochila variedentata Steph. ....................................................................................................251 Plagiochila vastifolia Steph. .........................................................................................................251 Plagiochila velata Inoue et Piippo ................................................................................................251 Plagiochila venezuelana Steph. .....................................................................................................251 Plagiochila ventricosotrigona Steph. ..............................................................................................251 Plagiochila verrucosa Steph. .........................................................................................................251 Plagiochila vetustisilva Steph........................................................................................................251 Plagiochila vexans Schifn. ex Steph. ............................................................................................227 Plagiochila viminea Spruce ..........................................................................................................251 Plagiochila vincentina Lindenb. ...................................................................................................226 World checklist of hornworts and liverworts *** ** ** * ** ** ** ** ** ** *** * ** ** ** * ** *** ** *** *** ** ** * * ** ** *** ** *** *** ** *** ** *** ** ** ** ** ** *** ** *** ** *** * *** *** *** *** ** *** 629 Plagiochila virginica A.Evans .......................................................................................................238 Plagiochila virginica var. caroliniana R.M.Schust.........................................................................238 Plagiochila virginica var. euryphylla R.M.Schust. .........................................................................238 Plagiochila viridis Steph. .............................................................................................................251 Plagiochila viridonigra (E.A.Hodgs.) Inoue .................................................................................251 Plagiochila vitiensis Mitt. .............................................................................................................223 Plagiochila vittiana Inoue ............................................................................................................251 Plagiochila vittifolia Steph. ..........................................................................................................251 Plagiochila vulcanica Steph. .........................................................................................................251 Plagiochila wacei S.W.Arnell ex Váňa et J.J.Engel .......................................................................251 Plagiochila wallichiana Steph. .....................................................................................................229 Plagiochila wallisiana Steph.........................................................................................................251 Plagiochila wangii Inoue .............................................................................................................229 Plagiochila wattsiana J.J.Engel et G.L.Merr. ................................................................................252 Plagiochila wettsteiniana S.W.Arnell ............................................................................................252 Plagiochila weymouthiana Steph. .................................................................................................252 Plagiochila wiemanniana S.W.Arnell ...........................................................................................252 Plagiochila wightii Nees...............................................................................................................238 Plagiochila wilhelmina Inoue .......................................................................................................238 Plagiochila wilmsiana Steph. .......................................................................................................221 Plagiochila winteri Steph. ............................................................................................................252 Plagiochila wrightii Steph. ...........................................................................................................252 Plagiochila xalapensis Gottsche ....................................................................................................252 Plagiochila xanthochroma Spruce .................................................................................................252 Plagiochila yoshinagana Steph. .....................................................................................................252 Plagiochila yulungensis Piippo ......................................................................................................229 Plagiochila zacuapana Gottsche ...................................................................................................252 Plagiochila zangii Grolle et M.L.So .............................................................................................228 Plagiochila zantenii Inoue ...........................................................................................................238 Plagiochila zhuensis Grolle et M.L.So ..........................................................................................222 Plagiochila zonata Steph. .............................................................................................................228 Plagiochilidium bidentulum (Steph.) Grolle .................................................................................252 Plagiochilion braunianum (Nees) S.Hatt. ....................................................................................252 Plagiochilion combinatum (Mitt.) Inoue ......................................................................................252 Plagiochilion conjugatum (Hook.) R.M.Schust. ...........................................................................252 Plagiochilion imbriatum (Mitt.) Inoue ........................................................................................252 Plagiochilion giulianettii (Steph.) Inoue .......................................................................................252 Plagiochilion herzogii Inoue .........................................................................................................252 Plagiochilion intermedium R.M.Schust. .......................................................................................252 Plagiochilion mayebarae S.Hatt. ...................................................................................................253 Plagiochilion oppositum (Reinw., Blume et Nees) S.Hatt. ............................................................253 Plagiochilion pachycephalum (De Not.) Inoue ..............................................................................253 Plagiochilion proliferum (Mitt.) R.M.Schust. ...............................................................................253 Plagiochilion theriotanum (Steph.) Inoue .....................................................................................253 Platycaulis renifolius R.M.Schust. ................................................................................................218 Plectocolea subamoena S.Winkl. ...................................................................................................511 Pleurozia acinosa (Mitt.) Trevis. ..................................................................................................462 Pleurozia articulata (Lindb.) Lindb. et Lackström .......................................................................462 Pleurozia caledonica (Gottsche) Steph. ........................................................................................462 Pleurozia conchifolia (Hook. et Arn.) Austin................................................................................462 Pleurozia conchifolia var. papillosa B.M.hiers .............................................................................462 Pleurozia curiosa B.M.hiers .......................................................................................................462 630 *** *** *** *** ** *** *** *** ** ** *** *** *** *** *** * ** ** ** ** * ** * *** ** ** ** ** *** *** * * ** ** ** ** ** ** ** ** ** ** * ** * ** *** * ** ** ** ** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Pleurozia gigantea (F.Weber) Lindb. ...........................................................................................462 Pleurozia heterophylla Steph. ex Fulford ......................................................................................462 Pleurozia johannis-winkleri Herzog .............................................................................................462 Pleurozia paradoxa (J.B.Jack) Schifn. .........................................................................................462 Pleurozia pocsii Frank Müll. ........................................................................................................463 Pleurozia purpurea Lindb. ...........................................................................................................462 Pleurozia subinlata (Austin) Austin ............................................................................................462 Plicanthus birmensis (Steph.) R.M.Schust. .....................................................................................54 Plicanthus diicilis (Steph.) L.Söderstr. et Váňa .............................................................................54 Plicanthus giganteus (Steph.) R.M.Schust. .....................................................................................54 Plicanthus hirtellus (F.Weber) R.M.Schust. ...................................................................................54 Podomitrium malaccense (Steph.) Campb. ...................................................................................471 Podomitrium marginatum (Steph.) Hürl......................................................................................471 Podomitrium phyllanthus (Hook.) Mitt. ......................................................................................471 Poeltia campylata Grolle ..............................................................................................................114 Polyotus peckianus Austin ............................................................................................................511 Porella abyssinica (Nees) Trevis. ..................................................................................................417 Porella abyssinica var. hoehnelii (Steph.) Pócs...............................................................................417 Porella acutifolia (Lehm. et Lindenb.) Trevis. ..............................................................................417 Porella acutifolia var. hattoriana (Pócs) S.Hatt.............................................................................417 Porella acutifolia var. linguifolia (Steph.) M.L.So.........................................................................417 Porella acutifolia subsp. tosana (Steph.) S.Hatt. ...........................................................................417 Porella andica (Gottsche) Hässel .................................................................................................417 Porella arboris-vitae (With.) Grolle..............................................................................................418 Porella arboris-vitae subsp. nitidula (C.Massal.) S.Hatt. ..............................................................418 Porella baueri (Schifn.) C.E.O.Jensen ........................................................................................418 Porella bolanderi (Austin) Pearson ...............................................................................................418 Porella borellii (Gola) Parihar ......................................................................................................418 Porella brachiata (Taylor) Spruce ................................................................................................418 Porella brasiliensis (Raddi) Schifn. ..............................................................................................418 Porella brasiliensis var. ciliata (Gottsche, Lindenb. et Nees) Schifn. ............................................418 Porella brasiliensis var. laevior (Gottsche, Lindenb. et Nees) Schifn. ...........................................418 Porella caespitans (Steph.) S.Hatt.................................................................................................418 Porella caespitans var. cordifolia (Steph.) S.Hatt. ex T.Katag. et T.Yamag. ...................................418 Porella caespitans subsp. latior (S.Hatt.) S.Hatt............................................................................418 Porella caespitans var. nipponica S.Hatt. .......................................................................................418 Porella caespitans var. relexigastria (Pócs) S.Hatt. ........................................................................419 Porella campylophylla (Lehm. et Lindenb.) Trevis. .......................................................................419 Porella campylophylla subsp. lancistipula (Steph.) S.Hatt. ............................................................419 Porella campylophylla var. ligulifera (Taylor) S.Hatt. ....................................................................419 Porella campylophylla var. tixieri (Pócs) S.Hatt. ...........................................................................419 Porella canariensis (F.Weber) Underw. ........................................................................................419 Porella capehorniensis Swails ........................................................................................................419 Porella capensis (Gottsche) Mitt...................................................................................................419 Porella caucasica Steph. ...............................................................................................................419 Porella chenii S.Hatt....................................................................................................................419 Porella chilensis (Lehm. et Lindenb.) Trevis. ................................................................................419 Porella chilensis var. antucensis (Gottsche) Hässel .........................................................................419 Porella chilensis var. fernandeziensis (Herzog) Swails ....................................................................419 Porella chilensis var. microloba (Herzog) Swails ............................................................................419 Porella chinensis (Steph.) S.Hatt. .................................................................................................419 Porella chinensis var. crispata Udar et Shaheen .............................................................................420 World checklist of hornworts and liverworts ** ** ** ** *** ** *** ** ** ** ** ** ** ** ** *** ** ** ** ** ** ** ** ** ** ** * ** ** ** ** ** ** *** ** ** * *** ** *** ** ** *** ** ** ** ** ** ** *** ** *** 631 Porella chinensis var. decurrens (Steph.) S.Hatt.............................................................................420 Porella chinensis var. hattorii Udar et Shaheen .............................................................................420 Porella chinensis var. irregularis (Steph.) S.Hatt. ..........................................................................420 Porella circinnata Lindb. .............................................................................................................420 Porella cordaeana (Huebener) Moore ..........................................................................................420 Porella cranfordii Steph. ..............................................................................................................420 Porella crispata (Hook.) Trevis. ...................................................................................................420 Porella cucullistipula Steph. .........................................................................................................420 Porella densifolia (Steph.) S.Hatt. ................................................................................................420 Porella densifolia subsp. andamana S.Hatt. ..................................................................................420 Porella densifolia subsp. appendiculata (Steph.) S.Hatt.................................................................420 Porella densifolia var. oviloba (Steph.) N.Kitag. ...........................................................................420 Porella densifolia var. paraphyllina (P.C.Chen) Pócs ....................................................................420 Porella densifolia var. pilosa S.Hatt. et K.C.Chang .......................................................................420 Porella densifolia var. robusta (Steph.) S.Hatt...............................................................................420 Porella elegantula (Mont.) E.A.Hodgs. ........................................................................................421 Porella faurieri (Steph.) S.Hatt. ...................................................................................................421 Porella fengii P.C.Chen et S.Hatt. ...............................................................................................421 Porella geheebii (Steph.) S.Hatt. ..................................................................................................421 Porella gracillima Mitt.................................................................................................................421 Porella grandifolia (Steph.) S.Hatt. ..............................................................................................421 Porella grandiloba Lindb. ............................................................................................................421 Porella grollei S.Hatt. ..................................................................................................................421 Porella handelii S.Hatt. ...............................................................................................................421 Porella hattorii Udar et Shaheen ..................................................................................................421 Porella hoeana S.Hatt. .................................................................................................................421 Porella imbricata Lour. ................................................................................................................421 Porella inaequalis (Gottsche) Perss. .............................................................................................421 Porella japonica (Sande Lac.) Mitt. ..............................................................................................421 Porella japonica subsp. appalachiana R.M.Schust. .......................................................................421 Porella japonica var. densespinosa S.Hatt. et M.X.Zhang ..............................................................421 Porella javanica (Gottsche) Inoue................................................................................................421 Porella latifolia J.S.Lou et Q.Li ...................................................................................................421 Porella leiboldii (Lehm.) Trevis. ..................................................................................................421 Porella longifolia (Steph.) S.Hatt. ................................................................................................421 Porella madagascariensis (Nees et Mont.) Trevis. .........................................................................421 Porella maxima (Steph.) M.L.So .................................................................................................422 Porella mexicana (Hampe ex Gottsche, Lindenb. et Nees) Trevis. ...............................................422 Porella montantii (Steph.) E.W.Jones ..........................................................................................422 Porella navicularis (Lehm. et Lindenb.) Pfeif. .............................................................................422 Porella nitens (Steph.) S.Hatt. .....................................................................................................422 Porella oblongifolia S.Hatt. ..........................................................................................................422 Porella obtusata (Taylor) Trevis. ..................................................................................................422 Porella obtusata var. macroloba (Steph.) S.Hatt. et M.X.Zhang....................................................422 Porella obtusiloba S.Hatt. ............................................................................................................422 Porella perrottetiana (Mont.) Trevis.............................................................................................422 Porella perrottetiana var. angustifolia Pócs ....................................................................................422 Porella perrottetiana var. ciliatodentata (P.C.Chen et P.C.Wu) S.Hatt. ........................................422 Porella perrottetiana var. triciliata (Steph.) Pócs ...........................................................................422 Porella pinnata L. ........................................................................................................................422 Porella planifolia J.S.Lou .............................................................................................................422 Porella platyphylla (L.) Pfeif........................................................................................................422 632 *** ** ** ** ** *** ** ** ** *** ** ** *** ** ** ** ** ** *** *** *** ** ** ** ** ** ** ** ** ** ** ** ** *** *** ** *** *** *** * *** *** *** *** *** *** *** *** *** * *** *** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Porella platyphylloidea (Schwein.) Lindb. .....................................................................................423 Porella plicata J.S.Lou .................................................................................................................423 Porella plumosa (Mitt.) Parihar ....................................................................................................423 Porella prolixa (Gottsche) E.W.Jones ..........................................................................................423 Porella pulcherrima Herzog et S.Hatt. .........................................................................................423 Porella relexa (Lehm. et Lindenb.) Trevis. ..................................................................................423 Porella revoluta (Lehm. et Lindenb.) Trevis. ................................................................................423 Porella revoluta var. propinqua (C.Massal.) S.Hatt. ......................................................................423 Porella roellii Steph. ....................................................................................................................423 Porella saccata M.L.So .................................................................................................................423 Porella sichuanensis S.Hatt. et K.C.Chang ...................................................................................423 Porella spinulosa (Steph.) S.Hatt..................................................................................................423 Porella squamulifera (Taylor) Trevis. ...........................................................................................423 Porella stephaniana (C.Massal.) S.Hatt. .......................................................................................423 Porella subdentata (Mitt.) Steph. .................................................................................................423 Porella subdentata var. camerunensis E.W.Jones ...........................................................................423 Porella subobtusa (Steph.) S.Hatt.................................................................................................423 Porella subparaphyllina J.S.Lou....................................................................................................423 Porella subsquarrosa (Nees et Mont.) Trevis. ...............................................................................424 Porella swailsii Grolle ..................................................................................................................424 Porella swartziana (F.Weber) Trevis. ...........................................................................................424 Porella triquetra (Steph.) E.W.Jones............................................................................................424 Porella truncata J.S.Lou...............................................................................................................424 Porella ulophylla (Steph.) S.Hatt..................................................................................................424 Porella undatorevoluta J.S.Lou .....................................................................................................424 Porella urceolata S.Hatt. ..............................................................................................................424 Porella urogea (C.Massal.) S.Hatt. ...............................................................................................424 Porella vallis-gratiae (Gottsche) E.W.Jones ..................................................................................424 Porella variabilis (Kashyap et R.S.Chopra) Parihar ......................................................................424 Porella vernicosa Lindb. ...............................................................................................................424 Porella viridissima (Mitt.) Grolle .................................................................................................424 Porella wataugensis (Sull.) Underw. ex M.Howe ..........................................................................424 Prasanthus jamalicus Potemkin....................................................................................................114 Prasanthus suecicus (Gottsche) Lindb...........................................................................................114 Preissia quadrata (Scop.) Nees .....................................................................................................492 Preissia quadrata subsp. hyperborea R.M.Schust. .........................................................................492 Prionolejeunea aemula (Gottsche) A.Evans ..................................................................................351 Prionolejeunea ampliretis Herzog .................................................................................................351 Prionolejeunea arguta (Nees) Steph..............................................................................................351 Prionolejeunea corbisieri Pearson ..................................................................................................351 Prionolejeunea decora (Taylor) Steph. ..........................................................................................351 Prionolejeunea denticulata (F.Weber) Schifn...............................................................................351 Prionolejeunea diversitexta (Hampe et Gottsche) Steph................................................................352 Prionolejeunea exauriculata A.Evans ............................................................................................352 Prionolejeunea galliotii Steph. ......................................................................................................352 Prionolejeunea grata (Gottsche) Schifn. ......................................................................................352 Prionolejeunea grollei Ilk.-Borg. et Schäf.-Verw. ..........................................................................352 Prionolejeunea guadalupensis (Lindenb.) Steph. ...........................................................................352 Prionolejeunea limpida Herzog ....................................................................................................352 Prionolejeunea maculata Herzog ..................................................................................................353 Prionolejeunea magnistipula Herzog.............................................................................................352 Prionolejeunea meissneri (Gottsche) Steph. ..................................................................................353 World checklist of hornworts and liverworts *** *** *** *** *** *** *** *** ** ** ** *** *** *** *** *** *** ** *** ** *** ** *** *** *** ** ** ** *** *** *** *** ** ** ** *** *** *** *** ** * ** *** *** *** *** *** *** ** ** *** ** 633 Prionolejeunea mucronata (Sande Lac.) Steph. .............................................................................352 Prionolejeunea muricatoserrulata (Spruce) Steph. .........................................................................352 Prionolejeunea principensis Vanden Berghen ................................................................................352 Prionolejeunea recurvula (Spruce) Steph. .....................................................................................352 Prionolejeunea scaberula (Spruce) Steph.......................................................................................352 Prionolejeunea schlimiana (Gottsche) Steph. ................................................................................352 Prionolejeunea trachyodes (Spruce) Steph. ....................................................................................352 Protocephalozia ephemeroides (Spruce) K.I.Goebel .......................................................................182 Protolophozia androgyna R.M.Schust. ex Váňa et L.Söderstr. .........................................................76 Protolophozia autoica (R.M.Schust.) Váňa et L.Söderstr. ...............................................................76 Protolophozia crispata (R.M.Schust.) Váňa et L.Söderstr................................................................76 Protolophozia druceae (Grolle et E.A.Hodgs.) Váňa et L.Söderstr. .................................................76 Protolophozia elongata (Steph.) Schljakov ......................................................................................76 Protolophozia herzogiana (E.A.Hodgs. et Grolle) Váňa et L.Söderstr. ............................................76 Protolophozia lancistipa (Grolle) Váňa et L.Söderstr. .....................................................................76 Protolophozia leucorhiza (Mitt.) Váňa et L.Söderstr. ......................................................................76 Protolophozia longilora (Herzog) L.Söderstr. et Váňa ....................................................................76 Protolophozia monoica (E.A.Hodgs.) Váňa et L.Söderstr. ...............................................................76 Protolophozia multicuspidata (Hook.f. et Taylor) Váňa et L.Söderstr. ............................................76 Protolophozia nivicola (R.M.Schust.) Váňa et L.Söderstr. ..............................................................76 Protolophozia perssoniana (H.A.Mill.) Váňa et L.Söderstr. .............................................................76 Protolophozia subalpina (R.M.Schust.) Váňa et L.Söderstr.............................................................76 Protolophozia tasmanica (R.M.Schust.) Váňa et L.Söderstr. ...........................................................76 Protolophozia verruculosa (R.M.Schust.) Váňa et L.Söderstr. .........................................................76 Protosyzygiella pseudoconnexa (Herzog) R.M.Schust. ......................................................................46 Pseudocephalozia cucullata J.J.Engel et R.M.Schust. ....................................................................168 Pseudocephalozia lepidozioides R.M.Schust...................................................................................169 Pseudocephalozia leptodictyon R.M.Schust....................................................................................169 Pseudocephalozia lobulata (Herzog) R.M.Schust. .........................................................................168 Pseudocephalozia paludicola R.M.Schust. .....................................................................................169 Pseudocephalozia quadriloba (Steph.) R.M.Schust. ......................................................................168 Pseudocephaloziella epiphytica R.M.Schust. ....................................................................................81 Pseudoisotachis pocsii Váňa.............................................................................................................99 Pseudolepicolea andoi (R.M.Schust.) Inoue ..................................................................................254 Pseudolepicolea fryei (Perss.) Grolle et Ando.................................................................................254 Pseudolepicolea grolleana (R.M.Schust.) Grolle ............................................................................254 Pseudolepicolea kuehnemannii (R.M.Schust.) Hässel ....................................................................254 Pseudolepicolea quadrilaciniata (Sull.) Fulford et J.Taylor ............................................................254 Pseudolepicolea temnomoides (R.M.Schust.) Váňa et J.J.Engel ......................................................255 Pseudolepicolea trollii (Herzog) Grolle et Ando ............................................................................255 Pseudolepicolea trollii var. darjeelingensis S.Hatt. et Mizut. ...........................................................255 Pseudolophocolea denticulata R.M.Schust. et J.J.Engel .................................................................253 Pseudomarsupidium aureocinctum (R.M.Schust.) J.J.Engel ............................................................45 Pseudomarsupidium borneensis (Grolle) Váňa, L.Söderstr., A.Hagborg et von Konrat ....................45 Pseudomarsupidium decipiens (Hook.) Grolle ................................................................................45 Pseudomarsupidium piliferum (Steph.) Herzog ex Grolle ...............................................................45 Pseudotritomaria heterophylla (R.M.Schust.) Konstant. et Vilnet ...................................................85 Psiloclada clandestina Mitt...........................................................................................................185 Psiloclada clandestina subsp. melanesica R.M.Schust. ...................................................................185 Psiloclada clandestina subsp. spinosa (S.W.Arnell) R.M.Schust. ...................................................185 Pteropsiella frondiformis Spruce ...................................................................................................185 Pteropsiella metzgeriiformis R.M.Schust. ......................................................................................185 634 *** *** *** ** * *** ** * ** * * *** * *** *** ** ** * *** *** ** * *** ** ** * *** ** ** ** *** *** ** ** *** ** ** *** ** ** ** ** ** *** *** ** *** ** * ** *** ** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Ptilidium californicum (Austin) Pearson ......................................................................................434 Ptilidium ciliare (L.) Hampe .......................................................................................................435 Ptilidium pulcherrimum (Weber) Vain. .......................................................................................435 Ptychanthus africanus Steph. ........................................................................................................411 Ptychanthus stephensonianus (Mitt.) Steph. ..................................................................................411 Ptychanthus striatus (Lehm. et Lindenb.) Nees ............................................................................411 Ptychanthus striatus var. intermedius (Gottsche) Verd. .................................................................411 Pycnolejeunea anotomensis Tixier .................................................................................................396 Pycnolejeunea borneensis Steph. ....................................................................................................396 Pycnolejeunea cavistipula (Steph.) Mizut. .....................................................................................397 Pycnolejeunea connivens Schifn. ex P.Syd. ...................................................................................397 Pycnolejeunea contigua (Nees) Grolle ...........................................................................................397 Pycnolejeunea convexifolia (Mitt.) Steph. .....................................................................................397 Pycnolejeunea decurviloba Steph. .................................................................................................397 Pycnolejeunea densistipula (Lehm. et Lindenb.) Steph. .................................................................397 Pycnolejeunea gradsteinii Ilk.-Borg. ..............................................................................................397 Pycnolejeunea grandiocellata Steph. ..............................................................................................397 Pycnolejeunea grossiloba Steph......................................................................................................397 Pycnolejeunea macroloba (Nees et Mont.) Schifn. .......................................................................397 Pycnolejeunea monophthalma (R.M.Schust.) Xiao L.He ...............................................................397 Pycnolejeunea novae-caledoniae (Steph.) Horik. ...........................................................................397 Pycnolejeunea palmicola Steph. ....................................................................................................397 Pycnolejeunea papillosa Xiao L.He ...............................................................................................397 Pycnolejeunea porrectilobula C.J.Bastos et O.Yano .......................................................................397 Pycnolejeunea retusa R.M.Schust. ................................................................................................397 Pycnolejeunea schlimiana Steph....................................................................................................397 Pycnolejeunea schwaneckei (Steph.) Schifn. ex P.Syd. ..................................................................398 Pycnolejeunea sphaeroides (Sande Lac.) J.B.Jack et Steph. .............................................................398 Radula acuminata Steph..............................................................................................................426 Radula acuta Mitt. ......................................................................................................................427 Radula acutangula Steph. ............................................................................................................430 Radula acutiloba Steph. ...............................................................................................................428 Radula aguirrei R.M.Schust. .......................................................................................................426 Radula allisonii Castle .................................................................................................................427 Radula amentulosa Mitt...............................................................................................................425 Radula amoena Herzog ...............................................................................................................427 Radula anceps Sande Lac. ............................................................................................................427 Radula aneurismalis (Hook.f. et Taylor) Gottsche, Lindenb. et Nees...........................................425 Radula angulata Steph. ...............................................................................................................430 Radula anisotoma M.A.M.Renner ...............................................................................................426 Radula ankeinensis Gottsche .......................................................................................................429 Radula antilleana Castle ..............................................................................................................429 Radula appressa Mitt. ..................................................................................................................428 Radula aquilegia (Hook.f. et Taylor) Gottsche, Lindenb. et Nees................................................428 Radula assamica Steph.................................................................................................................426 Radula auriculata Steph. .............................................................................................................426 Radula australiana K.Yamada......................................................................................................426 Radula australis Austin ................................................................................................................428 Radula bipinnata Mitt.................................................................................................................426 Radula bogotensis Steph. ..............................................................................................................430 Radula bolanderi Gottsche ..........................................................................................................430 Radula boninensis Furuki et K.Yamada ........................................................................................430 World checklist of hornworts and liverworts ** *** ** *** *** ** ** *** *** ** *** ** *** ** ** ** ** *** ** ** *** ** * ** *** *** *** * *** ** ** ** ** *** ** ** ** ** ** ** ** ** ** *** ** ** ** ** *** *** ** ** 635 Radula borneensis Steph...............................................................................................................428 Radula boryana (F.Weber) Nees ex Mont....................................................................................426 Radula brasilica K.Yamada ..........................................................................................................430 Radula brunnea Steph. ................................................................................................................426 Radula buccinifera (Hook.f. et Taylor) Gottsche, Lindenb. et Nees .............................................426 Radula caduca K.Yamada ............................................................................................................428 Radula caespitosa Steph. ..............................................................................................................430 Radula campanigera Mont...........................................................................................................426 Radula campanigera subsp. obiensis (S.Hatt.) K.Yamada ..............................................................426 Radula campanulata Lindenb. et Gottsche ..................................................................................430 Radula carringtonii J.B.Jack ........................................................................................................428 Radula castlei Grolle....................................................................................................................430 Radula cavifolia Hampe ex Gottsche, Lindenb. et Nees ..............................................................430 Radula ceylanica K.Yamada .........................................................................................................425 Radula chinensis Steph. ...............................................................................................................426 Radula cochabambaensis K.Yamada .............................................................................................430 Radula comorensis Steph. .............................................................................................................429 Radula complanata (L.) Dumort..................................................................................................428 Radula conferta Lindenb. et Gottsche ..........................................................................................430 Radula constricta Steph................................................................................................................428 Radula cordata Mitt. ...................................................................................................................430 Radula costaricensis Gottsche .......................................................................................................431 Radula crenulata Schifn..............................................................................................................427 Radula cubensis K.Yamada ..........................................................................................................429 Radula curvilobula M.L.So ..........................................................................................................431 Radula cuspidata Steph................................................................................................................427 Radula decora Gottsche ...............................................................................................................427 Radula decurrens Mitt. ................................................................................................................431 Radula demissa M.A.M.Renner ...................................................................................................426 Radula densifolia Castle ...............................................................................................................431 Radula diaphana K.I.Goebel .......................................................................................................431 Radula diversifolia Steph. ............................................................................................................429 Radula dolabrata K.Yamada ........................................................................................................431 Radula eggersii K.Yamada ............................................................................................................429 Radula elliottii Castle ..................................................................................................................431 Radula emarginata K.Yamada et Piippo ......................................................................................428 Radula episcia Spruce ..................................................................................................................429 Radula evansii Castle...................................................................................................................426 Radula evelynae K.Yamada ..........................................................................................................428 Radula falcata Steph....................................................................................................................431 Radula fauriana Steph.................................................................................................................431 Radula fendleri Gottsche .............................................................................................................428 Radula fernandezana Steph. ........................................................................................................431 Radula issifolia Steph..................................................................................................................425 Radula laccida Lindenb. et Gottsche ..........................................................................................426 Radula laccida var. brachycalyx Spruce ........................................................................................426 Radula lavifolia (Hook.f. et Taylor) Gottsche, Lindenb. et Nees ................................................431 Radula loridana Castle ...............................................................................................................429 Radula foricata M.A.M.Renner ..................................................................................................426 Radula formosa (C.F.W.Meissn. ex Spreng.) Nees .......................................................................425 Radula fujitae Furuki ..................................................................................................................431 Radula fulvifolia (Hook.f. et Taylor) Gottsche, Lindenb. et Nees ................................................429 636 ** ** *** ** ** ** *** ** ** ** *** ** *** * *** *** * *** ** *** ** *** *** *** *** *** ** ** ** *** ** ** ** ** *** ** ** ** *** ** ** ** *** *** *** ** * ** ** ** *** *** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Radula galapagona Steph. ............................................................................................................431 Radula gedena Gottsche ..............................................................................................................431 Radula gottscheana Taylor ...........................................................................................................426 Radula gracilis Mitt. ....................................................................................................................431 Radula gradsteinii K.Yamada .......................................................................................................431 Radula grandifolia Steph. ............................................................................................................431 Radula grandis Steph. ..................................................................................................................428 Radula grevilleana Taylor ............................................................................................................426 Radula grollei K.Yamada et Piippo ..............................................................................................431 Radula guyanensis K.Yamada .......................................................................................................431 Radula hastata Steph. ..................................................................................................................429 Radula hattorii K.Yamada ...........................................................................................................431 Radula hawaiica M.L.So .............................................................................................................431 Radula hedingeri K.I.Goebel .......................................................................................................431 Radula helix (Hook.f. et Taylor) Gottsche, Lindenb. et Nees ......................................................425 Radula hicksiae K.Yamada ...........................................................................................................425 Radula holstiana Steph. ...............................................................................................................429 Radula holtii Spruce ....................................................................................................................430 Radula husnotii Castle .................................................................................................................430 Radula imposita M.A.M.Renner ..................................................................................................427 Radula inlexa Gottsche ..............................................................................................................430 Radula inouei K.Yamada .............................................................................................................431 Radula involvens Spruce ..............................................................................................................431 Radula iwatsukiana K.Yamada ....................................................................................................431 Radula iwatsukii K.Yamada.........................................................................................................425 Radula jamaicensis Pearson..........................................................................................................431 Radula jamesonii Taylor ..............................................................................................................431 Radula japonica Gottsche ............................................................................................................428 Radula javanica Gottsche ............................................................................................................428 Radula jonesii Bouman, Dirkse et K.Yamada...............................................................................428 Radula jovetiana K.Yamada .........................................................................................................427 Radula kegelii Gottsche ex Steph. ................................................................................................430 Radula kilgourii M.A.M.Renner..................................................................................................427 Radula kinabaluensis K.Yamada ..................................................................................................431 Radula kitagawae K.Yamada .......................................................................................................431 Radula kojana Steph. ..................................................................................................................428 Radula koponenii K.Yamada et Piippo .........................................................................................431 Radula kurzii Steph. ...................................................................................................................431 Radula lacerata Steph. .................................................................................................................428 Radula laxiramea Steph...............................................................................................................431 Radula leiboldii Steph. ................................................................................................................432 Radula lewisii K.Yamada .............................................................................................................432 Radula ligula Steph. ....................................................................................................................432 Radula lindenbergiana Gottsche ex C.Hartm...............................................................................428 Radula lingulata Gottsche ...........................................................................................................432 Radula longiloba K.Yamada .........................................................................................................432 Radula longispica Steph. ..............................................................................................................432 Radula loriana Castle ..................................................................................................................427 Radula macroloba Steph. .............................................................................................................430 Radula madagascariensis Gottsche ...............................................................................................428 Radula mammosa Spruce .............................................................................................................427 Radula marginata Gottsche, Lindenb. et Nees ............................................................................432 World checklist of hornworts and liverworts ** *** ** ** ** ** ** *** ** ** ** * *** ** *** ** ** *** *** ** *** *** ** *** * ** ** ** ** ** ** * *** ** ** ** ** *** ** ** * *** *** ** *** ** ** *** * *** ** ** 637 Radula marojezica E.W.Jones ......................................................................................................428 Radula mauiensis M.L.So ............................................................................................................432 Radula mazarunensis K.Yamada ..................................................................................................430 Radula mexicana Lindenb. et Gottsche .......................................................................................430 Radula microloba Gottsche ..........................................................................................................430 Radula microlobula Castle ...........................................................................................................432 Radula minutilobula K.Yamada et Piippo ....................................................................................432 Radula mittenii Steph..................................................................................................................427 Radula mizutanii K.Yamada .......................................................................................................432 Radula morobeana K.Yamada et Piippo .......................................................................................425 Radula multiamentula E.A.Hodgs. ..............................................................................................425 Radula multilora Gottsche ex Schifn. ........................................................................................429 Radula myriopoda M.A.M.Renner...............................................................................................427 Radula neotropica Castle..............................................................................................................430 Radula nigra Pearson ..................................................................................................................432 Radula nilgiriensis Udar et D.Kumar ...........................................................................................432 Radula norrisii K.Yamada et Piippo ............................................................................................432 Radula notabilis M.A.M.Renner..................................................................................................427 Radula novae-hollandiae Hampe .................................................................................................428 Radula novivrieseana K.Yamada ..................................................................................................432 Radula novocaledonica Hürl. et K.Yamada...................................................................................432 Radula novocaledoniensis K.Yamada.............................................................................................432 Radula novoguineensis K.Yamada et Piippo..................................................................................429 Radula nudicaulis Steph. .............................................................................................................430 Radula nudicaulis var. delicatula P.Allorge et V.Allorge ...............................................................430 Radula nymannii Steph. ..............................................................................................................427 Radula obconica Sull....................................................................................................................429 Radula obovata Castle .................................................................................................................432 Radula obscura Mitt. ...................................................................................................................432 Radula obtusiloba Steph. .............................................................................................................429 Radula obtusiloba subsp. polyclada (A.Evans) S.Hatt....................................................................429 Radula oceania Castle ..................................................................................................................429 Radula ocellata K.Yamada ...........................................................................................................428 Radula okamurana Steph. ...........................................................................................................432 Radula onraedtii K.Yamada .........................................................................................................432 Radula opaciuscula (Spruce) Castle ..............................................................................................432 Radula oreopsis M.A.M.Renner ...................................................................................................429 Radula ornata E.A.Br. et Pócs .....................................................................................................425 Radula ovalilobula K.Yamada ......................................................................................................432 Radula oyamensis Steph. ..............................................................................................................432 Radula paganii Castle ..................................................................................................................432 Radula pallens (Sw.) Nees ex Mont. ............................................................................................432 Radula pandei Udar et Dh.Kumar...............................................................................................432 Radula patens K.Yamada .............................................................................................................433 Radula perrottetii Gottsche ..........................................................................................................426 Radula peruviana K.Yamada .......................................................................................................433 Radula philippinensis K.Yamada ..................................................................................................433 Radula physoloba Mont. ..............................................................................................................425 Radula pinnulata Mitt.................................................................................................................433 Radula plicata Mitt. ....................................................................................................................428 Radula pocsii K.Yamada ..............................................................................................................430 Radula portoricensis Steph. ..........................................................................................................429 638 *** ** ** ** *** ** ** ** *** ** ** *** ** *** ** *** *** *** *** * ** ** *** ** ** ** ** ** ** * ** *** * ** ** ** *** ** ** ** ** ** ** ** *** ** ** ** ** ** *** * Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Radula prolifera Arnell ................................................................................................................429 Radula protensa Lindenb. ............................................................................................................427 Radula protensa var. erectilobula Schifn.......................................................................................427 Radula pseudolaccida E.W.Jones .................................................................................................427 Radula pseudoscripta M.A.M.Renner ...........................................................................................425 Radula pseudostachya Spruce........................................................................................................433 Radula psychosis M.A.M.Renner ..................................................................................................427 Radula pugioniformis M.A.M.Renner ..........................................................................................428 Radula pulchella Mitt. .................................................................................................................428 Radula punctata Steph. ...............................................................................................................433 Radula pusilla Spruce ..................................................................................................................433 Radula quadrata Gottsche ...........................................................................................................429 Radula queenslandica K.Yamada ..................................................................................................425 Radula ratkowskiana K.Yamada ..................................................................................................427 Radula recubans Taylor ...............................................................................................................430 Radula relexa Nees et Mont. ......................................................................................................429 Radula retrolexa Taylor ..............................................................................................................428 Radula rhombiloba Steph. ...........................................................................................................433 Radula robinsonii Steph...............................................................................................................427 Radula rupicola K.Yamada ..........................................................................................................433 Radula saccatiloba Steph..............................................................................................................430 Radula santacruziana K.Yamada et Gradst. .................................................................................433 Radula scariosa Mitt. ...................................................................................................................425 Radula schaefer-verwimpii K.Yamada...........................................................................................430 Radula schoieldiana K.Yamada ...................................................................................................430 Radula sharpii K.Yamada ............................................................................................................429 Radula silvestris Gottsche ............................................................................................................429 Radula sinskeana K.Yamada ........................................................................................................433 Radula sinuata Gottsche ex Steph. ..............................................................................................433 Radula socorana Gerola ...............................................................................................................433 Radula sonsonensis Steph. ............................................................................................................433 Radula splendida M.A.M.Renner et Devos ..................................................................................425 Radula squarrosa K.Yamada ........................................................................................................425 Radula stellatogemmipara C.Gao et Y.H.Wu ...............................................................................433 Radula stenocalyx Mont. ..............................................................................................................427 Radula stipatilora Steph..............................................................................................................430 Radula strangulata Hook.f. et Taylor ..........................................................................................427 Radula striata Mitt. .....................................................................................................................430 Radula subinlata Lindenb. et Gottsche .......................................................................................430 Radula subsimplex Steph..............................................................................................................433 Radula subsquarrosa S.W.Arnell ..................................................................................................433 Radula sullivantii Austin .............................................................................................................430 Radula sumatrana Steph..............................................................................................................429 Radula tabularis Steph. ...............................................................................................................433 Radula tasmanica Steph...............................................................................................................428 Radula taylorii Steph. ..................................................................................................................433 Radula tectiloba Steph. ................................................................................................................433 Radula tenax Lindb. ....................................................................................................................426 Radula tenera Mitt. .....................................................................................................................430 Radula tenuis K.Yamada..............................................................................................................433 Radula thiersiae K.Yamada ..........................................................................................................425 Radula tjibodensis K.I.Goebel ......................................................................................................427 World checklist of hornworts and liverworts ** ** *** * *** ** ** *** ** *** ** *** ** *** *** ** ** *** *** ** ** ** ** ** ** * *** *** * * * * *** ** *** ** ** *** *** ** *** * ** ** *** ** *** ** ** ** *** ** 639 Radula tokiensis Steph. ................................................................................................................429 Radula underwoodii Castle ..........................................................................................................429 Radula uvifera (Hook.f. et Taylor) Gottsche, Lindenb. et Nees ...................................................425 Radula vagans Steph....................................................................................................................425 Radula van-zantenii K.Yamada ...................................................................................................429 Radula varilobula Castle..............................................................................................................429 Radula venezuelensis K.Yamada ...................................................................................................433 Radula ventricosa Steph. ..............................................................................................................427 Radula verrucosa K.Yamada.........................................................................................................425 Radula vieillardii Gottsche ..........................................................................................................433 Radula visianica C.Massal. ..........................................................................................................433 Radula voluta Taylor ...................................................................................................................430 Radula vrieseana Sande Lac. ........................................................................................................433 Radula weymouthiana Steph. .......................................................................................................428 Radula wichurae Steph. ...............................................................................................................429 Radula wrightii Castle .................................................................................................................433 Radula xalapensis Nees et Mont. .................................................................................................434 Radula yanoella R.M.Schust. .......................................................................................................427 Reboulia hemisphaerica (L.) Raddi ...............................................................................................483 Reboulia hemisphaerica subsp. acrogyna (R.M.Schust.) R.M.Schust. ............................................483 Reboulia hemisphaerica subsp. australis R.M.Schust. ....................................................................483 Reboulia hemisphaerica subsp. dioica R.M.Schust. .......................................................................483 Reboulia hemisphaerica var. issisquama Herzog............................................................................483 Reboulia hemisphaerica subsp. orientalis R.M.Schust....................................................................484 Reboulia hemisphaerica var. turkestanica C.E.O.Jensen ex Herzog................................................484 Rectolejeunea colombiana R.M.Schust. .........................................................................................393 Rectolejeunea emarginulilora (Schifn.) A.Evans ..........................................................................393 Rectolejeunea lagelliformis A.Evans ..............................................................................................393 Rectolejeunea lagelliformis subsp. hamata R.M.Schust. ................................................................393 Rectolejeunea lindenbergii Steph...................................................................................................393 Rectolejeunea lindigiana Steph. ....................................................................................................394 Rectolejeunea pachyderma R.M.Schust. ........................................................................................394 Rectolejeunea queenslandica (B.M.hiers) Xiao L.He ...................................................................393 Rectolejeunea truncatilobula C.J.Bastos ........................................................................................393 Rectolejeunea versifolia (Schifn.) L.Söderstr. et A.Hagborg ..........................................................393 Riccardia aberrans (Steph.) Gradst. .............................................................................................445 Riccardia aequicellularis (Steph.) Hewson....................................................................................439 Riccardia aequitexta (Steph.) E.A.Br. ...........................................................................................441 Riccardia aeruginosa Furuki .........................................................................................................439 Riccardia agumana Hewson ........................................................................................................445 Riccardia alba (Colenso) E.A.Br. .................................................................................................441 Riccardia albomarginata (Steph.) Schifn. ....................................................................................439 Riccardia alcicornis (Hook.f. et Taylor) Trevis. ............................................................................440 Riccardia algoides (Taylor) Meenks..............................................................................................445 Riccardia amazonica (Spruce) Schifn. ex Gradst. et Hekking ......................................................445 Riccardia amnicola Hässel ...........................................................................................................441 Riccardia andina (Spruce) Herzog ...............................................................................................441 Riccardia angustata Horik. ..........................................................................................................445 Riccardia angustealata (Steph.) Hewson ......................................................................................445 Riccardia angustissima (Steph.) H.A.Mill. ....................................................................................445 Riccardia arcuata Furuki .............................................................................................................439 Riccardia argentolimbata Hewson et Grolle .................................................................................438 640 ** ** *** ** * ** * ** ** ** ** ** ** ** ** ** ** ** ** ** ** *** *** ** ** ** *** *** ** ** ** ** ** ** ** *** *** ** ** ** ** * ** ** ** ** ** *** ** * * ** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Riccardia aspera (Steph.) Grolle...................................................................................................445 Riccardia asperulata R.M.Schust..................................................................................................439 Riccardia australis (Lehm.) Hewson ............................................................................................439 Riccardia autoica (Steph.) A.Evans ..............................................................................................443 Riccardia baldwinii (Steph.) H.A.Mill. ........................................................................................446 Riccardia barbilora (Steph.) Piippo.............................................................................................446 Riccardia baumannii Hürl. ..........................................................................................................445 Riccardia bipinnatiida (Colenso) Hewson ..................................................................................441 Riccardia bogotensis (Gottsche) Pagán ..........................................................................................446 Riccardia boliviensis (Steph.) Meenks ..........................................................................................446 Riccardia bongeriana Hewson ......................................................................................................446 Riccardia breviala E.A.Br. ...........................................................................................................443 Riccardia breviramosa (Steph.) A.Evans .......................................................................................443 Riccardia brunnea (Steph.) S.Hatt. ..............................................................................................446 Riccardia calcarea (Steph.) Meenks..............................................................................................446 Riccardia calva (Schifn.) A.Evans ...............................................................................................441 Riccardia canaliculata (Nees) Kuntze...........................................................................................439 Riccardia capillacea (Steph.) Meenks et C.De Jong......................................................................446 Riccardia capillacea var. dentata Meenks ......................................................................................446 Riccardia cardotii (Steph.) Pandé et S.C.Srivast. ..........................................................................446 Riccardia cataractarum (Spruce) Schifn. .....................................................................................446 Riccardia cervicornis (Spruce) Herzog ex Gradst. et Hekking .......................................................446 Riccardia chamedryfolia (With.) Grolle ........................................................................................439 Riccardia changbaishanensis C.Gao ..............................................................................................439 Riccardia chinensis C.Gao............................................................................................................446 Riccardia ciliolata (Spruce) Horik................................................................................................446 Riccardia cochleata (Hook.f. et Taylor) Kuntze............................................................................439 Riccardia colensoi (Steph.) W.Martin...........................................................................................441 Riccardia columbica (Steph.) Hässel ex Gradst. et Hekking .........................................................447 Riccardia comata (Steph.) H.A.Mill.............................................................................................447 Riccardia compacta (Steph.) S.W.Arnell.......................................................................................447 Riccardia comptonii (Pearson) H.A.Mill. .....................................................................................447 Riccardia conimitra (Steph.) A.Evans...........................................................................................440 Riccardia corralensis (Steph.) A.Evans ..........................................................................................440 Riccardia costata (Steph.) Hürl. ...................................................................................................447 Riccardia crassa (Schwägr.) C.Massal. ..........................................................................................441 Riccardia crassicaulis (Steph.) Meenks et C.De Jong ....................................................................447 Riccardia crassicrispa (Steph.) A.Evans .........................................................................................441 Riccardia crassiretis Schifn. .........................................................................................................447 Riccardia crenulata Schifn. .........................................................................................................447 Riccardia crenuliformis R.M.Schust. ............................................................................................447 Riccardia decolyana Schifn. .........................................................................................................447 Riccardia deguchii Furuki et K.T.Yong ........................................................................................445 Riccardia densiramea (Steph.) S.Hatt...........................................................................................447 Riccardia devexa Schifn. .............................................................................................................447 Riccardia diablotina (Spruce) Pagán ............................................................................................447 Riccardia diderma Hässel.............................................................................................................441 Riccardia digitiloba (Spruce) Pagán .............................................................................................447 Riccardia dilatata (Spruce) Schäf.-Verw. et Pócs .........................................................................447 Riccardia diminuta Schifn. .........................................................................................................447 Riccardia diminuta var. thermarum Schifn..................................................................................447 Riccardia distans (Spruce) Pagán..................................................................................................448 World checklist of hornworts and liverworts ** ** ** ** * * ** ** *** *** ** ** ** ** ** ** ** ** ** *** ** ** ** ** ** *** *** ** *** ** ** ** ** *** *** ** ** *** ** ** *** ** *** ** ** *** ** *** ** ** ** ** 641 Riccardia diversilora A.Evans ......................................................................................................441 Riccardia diversilora subsp. paucigyna R.M.Schust. .....................................................................441 Riccardia duriuscula Hässel .........................................................................................................441 Riccardia elata (Steph.) Schifn....................................................................................................448 Riccardia elata var. laccida Schifn. .............................................................................................448 Riccardia elata var. intercedens Schifn. ........................................................................................448 Riccardia elegans (Steph.) Hürl. ...................................................................................................448 Riccardia elisabethae houvenot et Reeb ......................................................................................448 Riccardia emarginata (Steph.) K.G.Hell ......................................................................................448 Riccardia eriocaula (Hook.) C.Massal. .........................................................................................439 Riccardia erosa (Steph.) E.W.Jones ..............................................................................................448 Riccardia falsiloribunda Hässel ...................................................................................................442 Riccardia fastigiata (Lehm.) Trevis. .............................................................................................448 Riccardia fendleri (Steph.) Pagán .................................................................................................448 Riccardia ilicina (Colenso) E.A.Hodgs. ......................................................................................444 Riccardia laccida (Steph.) S.Hatt. ...............................................................................................448 Riccardia laccidissima Schifn......................................................................................................448 Riccardia lagellaris (A.Gepp) H.A.Mill. ......................................................................................448 Riccardia lagellifrons C.Gao ........................................................................................................448 Riccardia lavovirens Furuki .........................................................................................................439 Riccardia leischeri (Steph.) H.A.Mill...........................................................................................448 Riccardia loribunda (Steph.) A.Evans .........................................................................................442 Riccardia luvigena Hässel............................................................................................................442 Riccardia foliacea Meenks et C.De Jong ......................................................................................448 Riccardia formosensis (Steph.) Horik. ...........................................................................................448 Riccardia fruticosa (Steph.) Furuki...............................................................................................439 Riccardia fucoidea (Sw.) C.Massal. ..............................................................................................448 Riccardia fuegiensis C.Massal. ......................................................................................................439 Riccardia furtiva E.A.Br. et Braggins ...........................................................................................440 Riccardia fuscobrunnea (Steph.) A.Evans......................................................................................441 Riccardia geniana Hewson...........................................................................................................449 Riccardia georgiensis (Steph.) Hässel ............................................................................................442 Riccardia georgiensis subsp. sympodea R.M.Schust. .......................................................................442 Riccardia glauca Furuki ...............................................................................................................439 Riccardia glaziovii (Spruce) Meenks ............................................................................................449 Riccardia gogolensis (Steph.) Hewson ...........................................................................................449 Riccardia gracilis (Steph.) R.M.Schust. ........................................................................................449 Riccardia graefei (Steph.) Hewson ..............................................................................................442 Riccardia grandilora (Steph.) H.A.Mill. ......................................................................................449 Riccardia granulata (Steph.) A.Evans...........................................................................................443 Riccardia grollei Furuki ...............................................................................................................449 Riccardia grossidens (Steph.) Pagán ..............................................................................................449 Riccardia grossitexta (Steph.) Furuki ............................................................................................445 Riccardia gunniana (Steph.) H.A.Mill. ........................................................................................449 Riccardia hamatilora (Steph.) H.A.Mill. .....................................................................................449 Riccardia hans-meyeri (Steph.) Meenks et C.De Jong ..................................................................449 Riccardia hans-meyeri var. dentata Meenks ..................................................................................449 Riccardia hattorii Furuki .............................................................................................................439 Riccardia hawaica (Steph.) H.A.Mill. ..........................................................................................449 Riccardia hebridensis (Steph.) H.A.Mill. ......................................................................................449 Riccardia herzogiana (Steph.) Meenks et C.De Jong ....................................................................449 Riccardia heteroclada Schifn. ......................................................................................................449 642 * ** ** ** ** ** * *** ** ** ** *** ** ** ** ** ** * ** *** * ** ** ** *** ** ** ** ** ** ** ** ** ** ** ** ** ** ** ** ** ** ** ** *** *** *** ** ** *** ** ** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Riccardia hirtilora (Steph.) Schifn. ............................................................................................449 Riccardia humilis (Gottsche) O.Yano ..........................................................................................450 Riccardia hyalina (Steph.) H.A.Mill. ...........................................................................................450 Riccardia hyalitricha Hässel .........................................................................................................445 Riccardia hydra Hürl. ..................................................................................................................450 Riccardia hymenophylloides Schifn. .............................................................................................450 Riccardia hymenophylloides var. laccida Schifn............................................................................450 Riccardia hymenophytoides (Spruce) Meenks ................................................................................450 Riccardia hypipamensis Hewson ...................................................................................................450 Riccardia ibana Hewson..............................................................................................................450 Riccardia inconspicua (Steph.) Reeb et Bardat..............................................................................445 Riccardia incurvata Lindb. ..........................................................................................................450 Riccardia innovans (Steph.) Pagán ...............................................................................................450 Riccardia insularis Schifn............................................................................................................450 Riccardia intercellula E.A.Br. .......................................................................................................443 Riccardia intricata (Steph.) H.A.Mill. .........................................................................................450 Riccardia jackii Schifn. ...............................................................................................................450 Riccardia jackii var. densa Schifn. ...............................................................................................450 Riccardia judithae Meenks et C.De Jong .....................................................................................450 Riccardia jugata R.M.Schust. ......................................................................................................450 Riccardia karstenii (Steph.) Schifn. .............................................................................................450 Riccardia kodamae Mizut. et S.Hatt. ...........................................................................................439 Riccardia laticostata (Spruce) Schifn. ..........................................................................................450 Riccardia latifrondoides Schifn. ...................................................................................................450 Riccardia latifrons (Lindb.) Lindb................................................................................................440 Riccardia latifrons subsp. arctica R.M.Schust. et Damsh. .............................................................440 Riccardia latifrons var. miyakeana (Schifn.) Furuki .....................................................................440 Riccardia lepidomitra (Spruce) Gradst. ........................................................................................450 Riccardia leptophylla (Spruce) Herzog..........................................................................................451 Riccardia leptostachya A.Evans .....................................................................................................444 Riccardia leptothallus R.M.Schust. ...............................................................................................451 Riccardia levieri Schifn. ..............................................................................................................451 Riccardia lichenoides (Steph.) H.A.Mill. ......................................................................................451 Riccardia ligulata (Steph.) Pócs et Schäf.-Verw. ..........................................................................451 Riccardia lilliena (Steph.) H.A.Mill. ............................................................................................451 Riccardia limbata (Steph.) E.W.Jones .........................................................................................451 Riccardia lobulata (Colenso) E.A.Hodgs......................................................................................444 Riccardia loefgrenii Schifn. .........................................................................................................451 Riccardia longilora (Steph.) Hewson ...........................................................................................451 Riccardia longioleata Hässel .........................................................................................................441 Riccardia longispica (Steph.) Pearson ...........................................................................................451 Riccardia loriana (Steph.) H.A.Mill.............................................................................................451 Riccardia macdonaldiana Hewson ...............................................................................................451 Riccardia macrantha (Pearson) H.A.Mill. ....................................................................................451 Riccardia magnicellularis Furuki ..................................................................................................451 Riccardia marginata (Colenso) Pearson .......................................................................................442 Riccardia marginata var. paciica Furuki ......................................................................................442 Riccardia marionensis R.M.Schust. ..............................................................................................438 Riccardia mejlandii S.W.Arnell ...................................................................................................444 Riccardia metzgeriiformis (Steph.) R.M.Schust. ...........................................................................451 Riccardia microscopica (Nees) Kuntze ..........................................................................................451 Riccardia minuta (Steph.) W.Martin ...........................................................................................451 World checklist of hornworts and liverworts *** *** *** ** * ** ** ** ** ** ** ** ** ** ** ** ** ** ** ** ** *** ** *** ** * ** ** *** ** *** ** ** ** ** ** ** ** ** ** *** *** ** *** ** ** ** * ** ** ** *** 643 Riccardia multicorpora E.A.Br. ....................................................................................................452 Riccardia multiida (L.) Gray .......................................................................................................444 Riccardia multiida subsp. decrescens (Steph.) Furuki ...................................................................444 Riccardia multiida subsp. synoica R.M.Schust. ............................................................................444 Riccardia multiidoides Schifn. ....................................................................................................452 Riccardia multioleata Hässel ........................................................................................................444 Riccardia multispica (Steph.) S.Hatt. ...........................................................................................452 Riccardia mycophora A.Evans.......................................................................................................442 Riccardia nadeaudii (Steph.) Hürl. ..............................................................................................452 Riccardia nagasakiensis (Steph.) S.Hatt. .......................................................................................440 Riccardia negeri (Steph.) A.Evans ................................................................................................442 Riccardia newellana (Steph.) H.A.Mill. .......................................................................................452 Riccardia nigra (Steph.) H.A.Mill. ..............................................................................................452 Riccardia nitida (Colenso) E.A.Hodgs.........................................................................................442 Riccardia nobilis (Steph.) Schifn. ................................................................................................452 Riccardia novo-amstelodamensis Schifn. .......................................................................................452 Riccardia nudilora (Steph.) Grolle ..............................................................................................452 Riccardia obtusa S.W.Arnell ........................................................................................................452 Riccardia obtusifrons (Steph.) H.A.Mill. ......................................................................................452 Riccardia omkaliensis Hewson .....................................................................................................452 Riccardia opuntiiformis S.W.Arnell ..............................................................................................443 Riccardia pallida (Spruce) Meenks et C.De Jong .........................................................................452 Riccardia pallidevirens (Steph.) A.Evans ......................................................................................443 Riccardia palmata (Hedw.) Carruth. ...........................................................................................440 Riccardia palmatiida (Steph.) H.A.Mill. .....................................................................................452 Riccardia palmatiformis Schifn. ..................................................................................................452 Riccardia papillata (Gottsche) Hässel ex Gradst. et Hekking .......................................................452 Riccardia papillosa (C.Massal. et Steph.) Hässel...........................................................................442 Riccardia papulosa (Steph.) E.A.Br. .............................................................................................444 Riccardia paramorum Meenks .....................................................................................................452 Riccardia parasitans (Steph.) Meenks et C.De Jong .....................................................................453 Riccardia parvula Schifn.............................................................................................................453 Riccardia patens Hässel ................................................................................................................444 Riccardia pauciramea (Steph.) H.A.Mill. .....................................................................................453 Riccardia paulensis Schifn. ..........................................................................................................453 Riccardia pectinata (Austin) H.A.Mill. ........................................................................................453 Riccardia pectinata var. fasciculata (Steph.) Hürl. ........................................................................438 Riccardia pellucida Piippo ...........................................................................................................453 Riccardia pembaiensis (Steph.) Hürl.............................................................................................453 Riccardia pengagensis Hewson......................................................................................................453 Riccardia pennata E.A.Br.............................................................................................................443 Riccardia perspicua E.A.Br. ..........................................................................................................443 Riccardia perssonii S.C.Srivast. et Udar ........................................................................................453 Riccardia philippinensis Furuki ....................................................................................................453 Riccardia phleganiana Hewson ....................................................................................................453 Riccardia pindensis Hewson .........................................................................................................453 Riccardia plana (Steph.) Hürl......................................................................................................453 Riccardia plana var. minor (Pearson) Hürl. ex H.A.Mill., H.Whittier et B.Whittier ....................453 Riccardia planilora (Steph.) S.Hatt. ............................................................................................440 Riccardia planilora var. aequatorialis Furuki ...............................................................................440 Riccardia planifrons (Spruce) Pagán .............................................................................................453 Riccardia plumiformis (Spruce) Hässel ex Meenks........................................................................453 644 ** *** ** ** ** *** ** *** ** ** ** ** *** ** ** ** * ** ** ** ** ** ** ** ** ** *** ** ** ** ** ** *** *** ** ** * ** *** ** ** * * ** ** ** ** ** ** ** * ** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Riccardia plumosa (Mitt.) E.O.Campb. .......................................................................................453 Riccardia poeppigiana (Lehm. et Lindenb.) Hässel ex Meenks et C.De Jong ................................453 Riccardia polyclada (Mitt.) Hässel................................................................................................444 Riccardia porcina (Hewson) L.Söderstr. .......................................................................................454 Riccardia portoricensis (Steph.) Pagán ..........................................................................................454 Riccardia prehensilis (Hook.f. et Taylor) C.Massal. ......................................................................438 Riccardia pseudodendroceros R.M.Schust. .....................................................................................440 Riccardia pumila Furuki ..............................................................................................................440 Riccardia punahuina (Steph.) H.A.Mill. ......................................................................................454 Riccardia pusilla Grolle ...............................................................................................................440 Riccardia ramosissima (Steph.) Grolle ..........................................................................................454 Riccardia regina Meenks et C.De Jong ........................................................................................454 Riccardia regnellii (Ångstr.) K.G.Hell ..........................................................................................454 Riccardia regularis (Steph.) Kühnem. ..........................................................................................442 Riccardia reyesiana Meenks..........................................................................................................454 Riccardia riccioides Pearson..........................................................................................................454 Riccardia rigida Schifn. ..............................................................................................................454 Riccardia rivularis Hässel.............................................................................................................442 Riccardia robbinsii Hewson et Grolle ..........................................................................................454 Riccardia robusta (Steph.) H.A.Mill. ...........................................................................................454 Riccardia rockii (Steph.) H.A.Mill. ..............................................................................................454 Riccardia rupicola (Steph.) Hewson .............................................................................................454 Riccardia russellii R.M.Schust......................................................................................................454 Riccardia saccatilora (Steph.) S.W.Arnell ....................................................................................454 Riccardia santapaui Udar et S.C.Srivast. ......................................................................................454 Riccardia saxicola Hässel..............................................................................................................442 Riccardia schwaneckei (Steph.) Pagán ..........................................................................................454 Riccardia singapurensis Schifn. ....................................................................................................454 Riccardia smaragdina Meenks et C.De Jong ................................................................................454 Riccardia spectabilis (Steph.) A.Evans ..........................................................................................442 Riccardia spegazziniana C.Massal. ...............................................................................................443 Riccardia spinulifera C.Massal. ....................................................................................................444 Riccardia spongiosa Furuki ...........................................................................................................439 Riccardia sprucei (Steph.) Meenks et C.De Jong ..........................................................................454 Riccardia squamifera Schifn. .......................................................................................................455 Riccardia statensis Hässel .............................................................................................................444 Riccardia stipatilora (Steph.) Pagán ............................................................................................455 Riccardia stricta R.M.Schust........................................................................................................455 Riccardia subalpina Furuki ..........................................................................................................440 Riccardia subantarctica Grolle et L.Söderstr. ................................................................................455 Riccardia subexalata Schifn.........................................................................................................455 Riccardia subexalata var. procera Schifn. .....................................................................................455 Riccardia submersa (Hook.f. et Taylor) Trevis. ............................................................................455 Riccardia submultiida Horik. ......................................................................................................455 Riccardia subpalmata (Steph.) Hürl. ............................................................................................455 Riccardia subsimplex (Steph.) Pagán ............................................................................................455 Riccardia sumatrana Schifn. .......................................................................................................455 Riccardia tahitensis (Steph.) Hürl. ...............................................................................................455 Riccardia tamariscina (Steph.) Schifn. ........................................................................................440 Riccardia tenax (Steph.) A.Evans .................................................................................................443 Riccardia tenella Hewson.............................................................................................................455 Riccardia tenerrima (Steph.) A.Evans...........................................................................................443 World checklist of hornworts and liverworts *** ** ** ** * *** ** ** ** * ** ** ** * ** ** *** ** ** * * * * ** ** ** ** *** *** *** *** *** *** *** *** * *** *** ** *** * ** * ** *** ** *** ** *** * * *** 645 Riccardia tenuicula (Spruce) Meenks ...........................................................................................455 Riccardia tenuis (Steph.) Schifn. .................................................................................................455 Riccardia thaxteri A.Evans ...........................................................................................................444 Riccardia theliophora Hässel ........................................................................................................443 Riccardia tjibodensis Schifn. ........................................................................................................455 Riccardia trichomanoides (Spruce) Hässel ex Meenks ...................................................................455 Riccardia tristaniana S.W.Arnell .................................................................................................444 Riccardia trukensis H.A.Mill. et Bonner ......................................................................................455 Riccardia tumbareriensis Hewson .................................................................................................455 Riccardia umida E.A.Br. ..............................................................................................................441 Riccardia upoluna (Steph.) Grolle................................................................................................456 Riccardia valida (Steph.) J.J.Engel ...............................................................................................456 Riccardia venosa (Steph.) Hürl. ...................................................................................................456 Riccardia villosa (Steph.) Pandé et S.C.Srivast. ............................................................................456 Riccardia virens (Steph.) Hürl. ....................................................................................................456 Riccardia virgata (Gottsche) Pagán ..............................................................................................456 Riccardia vitrea Furuki ................................................................................................................440 Riccardia wallisii (Steph.) Gradst. ...............................................................................................456 Riccardia wettsteinii Schifn. ........................................................................................................456 Riccardia wettsteinii var. angustilimbia Schifn. ............................................................................456 Riccardia wettsteinii var. crassa Schifn. ........................................................................................456 Riccardia wettsteinii var. procera Schifn. .....................................................................................456 Riccardia wettsteinii var. tenuiretis Schifn. ..................................................................................456 Riccardia womersleyana Hewson ..................................................................................................456 Riccardia xylophila Hässel............................................................................................................443 Riccia abuensis Bapna ..................................................................................................................499 Riccia acutisulca Steph.................................................................................................................500 Riccia alatospora O.H.Volk et Perold ..........................................................................................495 Riccia albida Sull. ex Austin ........................................................................................................493 Riccia albolimbata S.W.Arnell .....................................................................................................495 Riccia albomarginata Bisch. ex C.Krauss......................................................................................495 Riccia alboporosa Perold ..............................................................................................................495 Riccia albopunctata Jovet-Ast.......................................................................................................493 Riccia albornata O.H.Volk et Perold ...........................................................................................493 Riccia albovestita O.H.Volk ........................................................................................................495 Riccia amboinensis Schifn. ..........................................................................................................500 Riccia ampullacea Perold .............................................................................................................495 Riccia angolensis Steph.................................................................................................................495 Riccia aravalliensis Pandé et Udar................................................................................................500 Riccia argenteolimbata O.H.Volk et Perold .................................................................................495 Riccia arnellii Sultan Khan ..........................................................................................................500 Riccia asprella Carrington et Pearson ...........................................................................................500 Riccia asservanda De Not. ex Lamothe ........................................................................................500 Riccia atlantica Sérgio et Perold ..................................................................................................495 Riccia atromarginata Levier .........................................................................................................496 Riccia atromarginata var. jovet-astiae Rauh et Buchloh ................................................................496 Riccia atropurpurea Sim...............................................................................................................496 Riccia attenuata Pandé ................................................................................................................500 Riccia australis Steph. ..................................................................................................................493 Riccia bahiensis Steph. .................................................................................................................498 Riccia balansae Steph...................................................................................................................500 Riccia beyrichiana Hampe ...........................................................................................................496 646 * *** *** *** *** ** ** *** *** *** *** *** *** *** *** *** *** *** ** *** * *** ** * *** *** *** ** ** *** * ** *** * *** ** *** *** ** *** *** *** * *** *** *** *** ** ** ** *** *** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Riccia bialbistrata Hässel .............................................................................................................500 Riccia bicarinata Lindb. ..............................................................................................................496 Riccia bicolorata Perold ...............................................................................................................496 Riccia bifurca Hofm. ..................................................................................................................496 Riccia billardierei Mont. et Nees .................................................................................................496 Riccia biokoensis Perold ...............................................................................................................500 Riccia blackii Na-halang............................................................................................................500 Riccia boliviensis Jovet-Ast ...........................................................................................................493 Riccia brasiliensis Schifn. ............................................................................................................493 Riccia breidleri Jur. ex Steph........................................................................................................496 Riccia breutelii Hampe ................................................................................................................493 Riccia brittonii M.Howe .............................................................................................................493 Riccia bullosa Link ......................................................................................................................499 Riccia californica Austin ..............................................................................................................496 Riccia campbelliana M.Howe ......................................................................................................496 Riccia canaliculata Hofm. ..........................................................................................................498 Riccia cancellata Taylor ...............................................................................................................497 Riccia caroliniana Na-halang .....................................................................................................500 Riccia cartilaginosa Steph.............................................................................................................500 Riccia cavernosa Hofm. ..............................................................................................................499 Riccia chartacea K.I.Goebel .........................................................................................................500 Riccia chiapasensis Jovet-Ast ........................................................................................................498 Riccia chinensis Herzog................................................................................................................500 Riccia chudoana Steph. ................................................................................................................493 Riccia ciliata Hofm. ...................................................................................................................496 Riccia ciliifera Link .....................................................................................................................496 Riccia cincta Jovet-Ast .................................................................................................................497 Riccia collata Na-halang ............................................................................................................500 Riccia compacta Garside ..............................................................................................................500 Riccia concava Bisch. ex C.Krauss................................................................................................495 Riccia congoana Steph. ................................................................................................................496 Riccia convexa Steph....................................................................................................................500 Riccia coracina Jovet-Ast .............................................................................................................493 Riccia coronata Sim .....................................................................................................................500 Riccia corrugata Jovet-Ast ............................................................................................................493 Riccia crassa Steph. ......................................................................................................................500 Riccia crassifrons Spruce...............................................................................................................498 Riccia crassivenia Jovet-Ast ..........................................................................................................493 Riccia crenatodentata O.H.Volk ..................................................................................................500 Riccia crinita Taylor ....................................................................................................................496 Riccia crozalsii Levier...................................................................................................................496 Riccia cruciata Kashyap ...............................................................................................................497 Riccia crustata Trab. ....................................................................................................................496 Riccia crystallina L. ......................................................................................................................499 Riccia cubensis S.W.Arnell ...........................................................................................................493 Riccia cupulifera A.V.Duthie .......................................................................................................499 Riccia curtisii (Austin) Austin ......................................................................................................499 Riccia delavayi Steph. ..................................................................................................................500 Riccia deserticola Steph. ...............................................................................................................500 Riccia dictyospora M.Howe ..........................................................................................................496 Riccia discolor Lehm. et Lindenb. ................................................................................................494 Riccia duplex Lorb. ex Müll.Frib. ................................................................................................498 World checklist of hornworts and liverworts ** *** *** *** *** *** *** *** *** ** ** *** *** * *** *** *** *** *** * *** ** ** *** ** *** ** ** *** *** ** *** *** *** ** *** *** *** *** * * ** ** *** * *** ** *** *** ** ** *** 647 Riccia duplex var. megaspora Na-halang .....................................................................................498 Riccia dussiana Steph. .................................................................................................................498 Riccia eburnea Jovet-Ast ..............................................................................................................497 Riccia ekmanii S.W.Arnell...........................................................................................................494 Riccia elliottii Steph.....................................................................................................................494 Riccia elongata Perold ..................................................................................................................495 Riccia enyae Jovet-Ast ..................................................................................................................494 Riccia erubescens Perold ...............................................................................................................500 Riccia erythrocarpa Jovet-Ast........................................................................................................494 Riccia esulcata Steph. ...................................................................................................................500 Riccia fertilissima Steph. ..............................................................................................................500 Riccia luitans L. ..........................................................................................................................498 Riccia frostii Austin .....................................................................................................................498 Riccia frostii var. crystallinoides Schifn.........................................................................................498 Riccia fruchartii Steph. ................................................................................................................494 Riccia furfuracea Perold ...............................................................................................................495 Riccia gangetica Ahmad ex L.Söderstr., A.Hagborg et von Konrat ...............................................494 Riccia garsidei Sim.......................................................................................................................499 Riccia geissleriana Jovet-Ast .........................................................................................................498 Riccia gemmifera O.H.Volk.........................................................................................................500 Riccia glauca L. ...........................................................................................................................496 Riccia glauca var. ciliaris Warnst..................................................................................................496 Riccia gothica Damsh. et Hallingb. ..............................................................................................496 Riccia gougetiana Durieu et Mont. ..............................................................................................496 Riccia gougetiana var. armatissima Levier ex Müll.Frib.................................................................496 Riccia grandis Nees ......................................................................................................................494 Riccia grollei Udar .......................................................................................................................500 Riccia handelii Schifn. ................................................................................................................500 Riccia hantamensis Perold ............................................................................................................495 Riccia hasskarliana Steph. ............................................................................................................497 Riccia hawaiiensis Hürl. ..............................................................................................................500 Riccia hegewaldiana Jovet-Ast ......................................................................................................498 Riccia helenae Jovet-Ast ...............................................................................................................494 Riccia hirsuta O.H.Volk et Perold ...............................................................................................495 Riccia hirta (Austin) Underw. .....................................................................................................494 Riccia horrida Jovet-Ast ...............................................................................................................494 Riccia hortorum Bory...................................................................................................................494 Riccia howellii M.Howe ..............................................................................................................494 Riccia huebeneriana Lindenb. ......................................................................................................498 Riccia huebeneriana subsp. sullivantii (Austin) R.M.Schust. ........................................................498 Riccia ianthina Jovet-Ast .............................................................................................................494 Riccia indica Udar et A.Gupta.....................................................................................................501 Riccia indira-gandhiensis Dabhade et A.Hasan ............................................................................501 Riccia inlexa Taylor ....................................................................................................................494 Riccia intermedia Roum. .............................................................................................................501 Riccia iodocheila M.Howe ...........................................................................................................494 Riccia jodhpurensis Bapna ............................................................................................................501 Riccia jovet-astiae E.Vianna .........................................................................................................498 Riccia junghuhniana Nees et Lindenb..........................................................................................497 Riccia junghuhniana var. simplex Schifn. ....................................................................................497 Riccia kirinensis C.Gao et K.C.Chang .........................................................................................501 Riccia lamellosa Raddi .................................................................................................................496 648 *** *** ** ** *** *** *** *** ** ** ** *** *** ** *** ** *** * *** ** *** *** *** *** ** *** *** *** ** *** *** *** *** * ** ** ** * *** *** *** ** ** *** *** ** ** *** *** *** ** *** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Riccia lanceolata Steph. ...............................................................................................................494 Riccia laxisquamata (Steph.) Steph. .............................................................................................501 Riccia leptothallus R.M.Schust. ....................................................................................................499 Riccia liaoningensis C.Gao et K.C.Chang ....................................................................................501 Riccia ligula Steph. ......................................................................................................................496 Riccia limbata Bisch. ex C.Krauss................................................................................................496 Riccia limicola Jovet-Ast ..............................................................................................................498 Riccia lindmanii Steph. ...............................................................................................................494 Riccia linearis (Schifn.) Steph. ....................................................................................................501 Riccia luticola Na-halang...........................................................................................................501 Riccia macallisteri M.Howe .........................................................................................................494 Riccia macrocarpa Levier .............................................................................................................496 Riccia macrospora Steph. .............................................................................................................494 Riccia mamillata Trab. ex Steph. .................................................................................................494 Riccia mammifera O.H.Volk et Perold ........................................................................................496 Riccia mamrensis Perold ..............................................................................................................501 Riccia mangalorica Ahmad ex Jovet-Ast .......................................................................................497 Riccia marginata Lindb. ..............................................................................................................501 Riccia mauryana Steph. ...............................................................................................................494 Riccia melanospora Kashyap.........................................................................................................501 Riccia melitensis C.Massal............................................................................................................496 Riccia membranacea Gottsche et Lindenb. ...................................................................................493 Riccia michelii Raddi ...................................................................................................................496 Riccia microciliata O.H.Volk et Perold........................................................................................497 Riccia miyakeana Schifn. ............................................................................................................501 Riccia moenkemeyeri Steph. .........................................................................................................499 Riccia montana Perold .................................................................................................................497 Riccia multiida (Steph.) Steph. ...................................................................................................498 Riccia muscicola Steph. ................................................................................................................501 Riccia namaquensis Perold ...........................................................................................................495 Riccia natalensis Sim....................................................................................................................497 Riccia nigerica E.W.Jones ............................................................................................................501 Riccia nigrella DC. ......................................................................................................................497 Riccia nigrescens Mont. ................................................................................................................501 Riccia nipponica S.Hatt. ..............................................................................................................501 Riccia novo-hannoverana Schifn. ................................................................................................501 Riccia numeensis Steph. ...............................................................................................................501 Riccia obtusa Meijer ....................................................................................................................501 Riccia oerstediana Lindenb. et Hampe .........................................................................................501 Riccia okahandjana S.W.Arnell ...................................................................................................497 Riccia olgensis Na-halang ...........................................................................................................494 Riccia ozarkiana McGregor .........................................................................................................497 Riccia pandei Udar ......................................................................................................................501 Riccia papillispora Steph. .............................................................................................................501 Riccia papillosa Moris ..................................................................................................................497 Riccia papulosa (Steph.) Steph. ....................................................................................................501 Riccia papulosa var. variabilis Na-halang ...................................................................................502 Riccia paraguayensis Spruce .........................................................................................................498 Riccia paranaensis Hässel .............................................................................................................499 Riccia parvoareolata O.H.Volk et Perold .....................................................................................495 Riccia pathankotensis Kashyap .....................................................................................................502 Riccia perennis Steph. ..................................................................................................................499 World checklist of hornworts and liverworts *** * *** ** *** *** * ** ** *** *** *** *** *** ** * ** ** *** ** *** *** * * *** ** *** * *** ** *** *** *** *** *** ** ** *** *** *** *** *** * ** *** *** ** ** *** *** *** ** 649 Riccia perssonii Sultan Khan ........................................................................................................499 Riccia perthiana Steph. ex K.I.Goebel .........................................................................................502 Riccia planobiconvexa Steph. .......................................................................................................494 Riccia polycarpa (Trab.) Jelenc .....................................................................................................498 Riccia porosa Taylor ....................................................................................................................498 Riccia pottsiana Sim ....................................................................................................................497 Riccia prominens Meijer...............................................................................................................502 Riccia pseudoluitans C.Gao et K.C.Chang ..................................................................................502 Riccia pubescens S.Hatt. ...............................................................................................................502 Riccia pullulans Jovet-Ast ............................................................................................................498 Riccia pulveracea Perold ..............................................................................................................495 Riccia purpurascens Lehm. ...........................................................................................................499 Riccia radiata Perold ...................................................................................................................502 Riccia radicosa Pearson ................................................................................................................495 Riccia rechingeri Steph.................................................................................................................502 Riccia reticulatula Udar ...............................................................................................................502 Riccia rhenana Lorb. ex Müll.Frib. ..............................................................................................499 Riccia rhenana var. violacea M.F.Boiko .......................................................................................499 Riccia ridleyi A.Gepp...................................................................................................................494 Riccia rorida Na-halang ............................................................................................................502 Riccia rosea O.H.Volk et Perold ..................................................................................................497 Riccia rubricollis Garside et A.V.Duthie ex Perold .......................................................................499 Riccia runssorensis Steph. .............................................................................................................497 Riccia saharensis Steph. ex Jovet-Ast ............................................................................................502 Riccia sanguineisporis Jovet-Ast ....................................................................................................494 Riccia satoi S.Hatt. ......................................................................................................................502 Riccia schelpei O.H.Volk et Perold ..............................................................................................493 Riccia schroederi Steph. ................................................................................................................502 Riccia schweinfurthii Steph. .........................................................................................................502 Riccia sibayenii Perold .................................................................................................................502 Riccia simii Perold.......................................................................................................................495 Riccia singularis Jovet-Ast ............................................................................................................499 Riccia somaliensis Perold ..............................................................................................................502 Riccia sommieri Levier .................................................................................................................497 Riccia sorocarpa Bisch. .................................................................................................................497 Riccia sorocarpa subsp. sorocarpa var. heegii Schifn......................................................................497 Riccia spongiosula Na-halang .....................................................................................................502 Riccia squamata Nees ..................................................................................................................495 Riccia stricta (Lindenb.) Perold ...................................................................................................499 Riccia subbifurca Warnst. ex Croz. ..............................................................................................497 Riccia subdepilata Jovet-Ast .........................................................................................................495 Riccia subplana Steph. .................................................................................................................495 Riccia subtilis (Steph.) Steph. ......................................................................................................502 Riccia sumatrana Meijer ..............................................................................................................502 Riccia symoensii Vanden Berghen ................................................................................................502 Riccia taeniiformis Jovet-Ast ........................................................................................................495 Riccia tasmanica Steph. ex Rodway .............................................................................................502 Riccia tenella D.L.Jacobs .............................................................................................................497 Riccia tomentosa O.H.Volk et Perold ..........................................................................................502 Riccia trabutiana Steph. ..............................................................................................................497 Riccia trachyglossa Perold .............................................................................................................495 Riccia treubiana Steph. ................................................................................................................502 650 * * * ** * ** *** ** *** *** *** *** *** *** *** *** *** ** ** *** *** *** *** * *** *** *** *** * *** *** * *** *** *** * *** * *** *** ** *** *** *** * * ** *** *** ** *** *** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Riccia treubiana var. subrubescens Schifn.....................................................................................502 Riccia triangularis Steph. .............................................................................................................502 Riccia tuberculata Poir. ................................................................................................................502 Riccia udarii Kanwal ...................................................................................................................502 Riccia velenovskyi Kavina .............................................................................................................503 Riccia velimalaiana A.E.D.Daniels et P.Daniel............................................................................503 Riccia viannae Jovet-Ast ..............................................................................................................495 Riccia victoriensis Steph. ..............................................................................................................503 Riccia villosa Steph. .....................................................................................................................495 Riccia violacea M.Howe ..............................................................................................................497 Riccia violacea var. laevis Jovet-Ast ..............................................................................................497 Riccia vitalii Jovet-Ast .................................................................................................................495 Riccia vitrea Perold .....................................................................................................................495 Riccia volkii S.W.Arnell ..............................................................................................................499 Riccia vulcanicola Eb.Fisch. .........................................................................................................499 Riccia warnstorii Limpr. ex Warnst. ...........................................................................................497 Riccia weinionis Steph. ................................................................................................................495 Riccia weymouthiana Steph. ex Rodway.......................................................................................503 Riccia wichurae Steph..................................................................................................................503 Ricciocarpos natans (L.) Corda .....................................................................................................503 Riella ainis M.Howe et Underw. ...............................................................................................506 Riella alatospora Wigglesw...........................................................................................................505 Riella americana M.Howe et Underw. ........................................................................................505 Riella battandieri Trab. ...............................................................................................................505 Riella bialata Trab. .....................................................................................................................505 Riella capensis Cavers...................................................................................................................505 Riella choconensis Hässel ..............................................................................................................505 Riella cossoniana Trab. ................................................................................................................506 Riella cyrenaica Maire..................................................................................................................505 Riella echinata (Müll.Frib.) Segarra, Puche et Sabovlj..................................................................506 Riella echinospora Wigglesw.........................................................................................................505 Riella gallica Balansa ex Trab. .....................................................................................................505 Riella gamundiae Hässel ..............................................................................................................506 Riella halophila Banwell ..............................................................................................................505 Riella helicophylla (Bory et Mont.) Mont.....................................................................................505 Riella helicophylla var. macrocarpa P.Allorge ................................................................................505 Riella heliospora Segarra, Puche et Sabovlj. ..................................................................................506 Riella indica Steph. ex Kashyap ...................................................................................................505 Riella mediterranea Segarra, Puche, Sabovlj., M.Infante et Heras ................................................506 Riella notarisii (Mont.) Mont. .....................................................................................................505 Riella numidica Trab. ..................................................................................................................505 Riella pampae Hässel ...................................................................................................................505 Riella parisii Gottsche .................................................................................................................505 Riella purpureospora Wigglesw.....................................................................................................505 Riella reuteri Mont. .....................................................................................................................505 Riella sersuensis Trab. ..................................................................................................................505 Riella spiculata J.Taylor ...............................................................................................................505 Riella trigonospora Segarra et Puche .............................................................................................506 Riella undulata Hässel .................................................................................................................506 Rivulariella gemmipara (A.Evans) D.H.Wagner ..........................................................................122 Ruizanthus venezuelanus R.M.Schust. .........................................................................................103 Saccobasis polita (Nees) H.Buch ....................................................................................................85 World checklist of hornworts and liverworts ** * ** * *** *** ** ** *** ** ** ** ** *** ** ** *** * * * * * *** *** *** *** ** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** ** ** *** ** *** * *** *** ** *** 651 Saccobasis polymorpha (R.M.Schust.) Schljakov .............................................................................85 Saccogyna ligulata Steph. .............................................................................................................122 Saccogyna subacuta Steph. ...........................................................................................................122 Saccogyna tridens Steph. ..............................................................................................................122 Saccogyna viticulosa (L.) Dumort. ..................................................................................................98 Saccogynidium australe (Mitt.) Grolle ............................................................................................98 Saccogynidium caldense (Ångstr.) Grolle.........................................................................................98 Saccogynidium chiloscyphoides R.M.Schust. ....................................................................................98 Saccogynidium decurvum (Mitt.) Grolle .........................................................................................98 Saccogynidium goebelii (Herzog) Grolle .........................................................................................98 Saccogynidium irregularospinum C.Gao, T.Cao et M.J.Lai.............................................................98 Saccogynidium muricellum (De Not.) Grolle..................................................................................98 Saccogynidium rigidulum (Nees) Grolle .........................................................................................98 Saccogynidium vasculosum (Hook.f. et Taylor) Grolle ....................................................................98 Sandeothallus japonicus (Inoue) Crand.-Stotl. et Stotler ...............................................................474 Sandeothallus radiculosus (Schifn.) R.M.Schust. .........................................................................474 Sauteria alpina (Nees) Nees ........................................................................................................485 Sauteria chilensis (Lindenb.) Grolle .............................................................................................485 Sauteria crassipes Austin ..............................................................................................................485 Sauteria inlata C.Gao et K.C.Chang ..........................................................................................485 Sauteria japonica (Shimizu et S.Hatt.) S.Hatt. ............................................................................485 Sauteria nyikaensis Perold ............................................................................................................485 Sauteria spongiosa (Kashyap) S.Hatt. ...........................................................................................485 Scapania aequiloba (Schwägr.) Dumort. ........................................................................................87 Scapania americana Müll.Frib. ......................................................................................................87 Scapania ampliata Steph. ..............................................................................................................85 Scapania ampliata subsp. queenslandica M.L.Hicks .......................................................................85 Scapania apiculata Spruce .............................................................................................................87 Scapania aspera M.Bernet et Bernet ..............................................................................................87 Scapania bhutanensis Amakawa .....................................................................................................88 Scapania bolanderi Austin .............................................................................................................85 Scapania brevicaulis Taylor............................................................................................................90 Scapania calcicola (Arnell et J.Perss.) Ingham ................................................................................90 Scapania carinthiaca J.B.Jack ex Lindb. .........................................................................................87 Scapania carinthiaca var. massalongi Müll.Frib. .............................................................................87 Scapania ciliata Sande Lac.............................................................................................................88 Scapania ciliata subsp. hawaiica (Müll.Frib.) Potemkin ................................................................88 Scapania ciliatospinosa Horik.........................................................................................................86 Scapania compacta (Roth) Dumort................................................................................................88 Scapania contorta Mitt. .................................................................................................................85 Scapania crassiretis Bryhn ..............................................................................................................90 Scapania curta (Mart.) Dumort. ....................................................................................................88 Scapania curta var. grandiretis R.M.Schust. ...................................................................................88 Scapania curta var. isoloba R.M.Schust. .........................................................................................88 Scapania cuspiduligera (Nees) Müll.Frib. .......................................................................................89 Scapania cuspiduligera var. diplophyllopsis R.M.Schust. ..................................................................89 Scapania davidii Potemkin ............................................................................................................86 Scapania degenii Schifn. ex Müll.Frib...........................................................................................90 Scapania diplophylloides Amakawa et S.Hatt. .................................................................................88 Scapania esterhuyseniae S.W.Arnell ................................................................................................88 Scapania ferruginaeoides T.Cao, C.Gao et J.Sun ............................................................................86 Scapania ferruginea (Lehm. et Lindenb.) Lehm. et Lindenb. .........................................................87 652 *** *** ** * *** *** *** * *** ** *** ** *** *** *** *** *** ** *** *** *** *** *** *** ** *** * *** *** *** *** *** *** ** *** *** ** ** ** *** *** *** *** *** *** *** *** *** *** *** *** ** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Scapania fulfordiae W.S.Hong ......................................................................................................88 Scapania gamundiae R.M.Schust. ..................................................................................................88 Scapania gaochii X.Fu ex T.Cao ....................................................................................................86 Scapania geppii Steph. ...................................................................................................................86 Scapania gigantea Horik. ...............................................................................................................91 Scapania glaucocephala (Taylor) Austin .........................................................................................92 Scapania glaucocephala var. saxicola (R.M.Schust.) Potemkin ........................................................92 Scapania glaucoviridis Horik. ........................................................................................................90 Scapania gracilis Lindb. .................................................................................................................85 Scapania grandiloba Steph. ............................................................................................................91 Scapania griithii Schifn. .............................................................................................................92 Scapania grossidens Steph. ex Müll.Frib. ........................................................................................91 Scapania gymnostomophila Kaal. ....................................................................................................90 Scapania harae Amakawa ..............................................................................................................86 Scapania hedbergii S.W.Arnell .......................................................................................................91 Scapania helvetica Gottsche ...........................................................................................................89 Scapania hians Steph. ex Müll.Frib. ..............................................................................................87 Scapania hians subsp. salishensis J.D.Godfrey et G.Godfrey ...........................................................87 Scapania himalayica Müll.Frib. ex Herzog ....................................................................................90 Scapania hirosakiensis Steph. ex Müll.Frib. ....................................................................................88 Scapania hollandiae W.S.Hong .....................................................................................................88 Scapania hyperborea Jørg. ..............................................................................................................90 Scapania integerrima Steph............................................................................................................91 Scapania irrigua (Nees) Nees.........................................................................................................89 Scapania irrigua subsp. rufescens (Loeske) R.M.Schust. .................................................................89 Scapania javanica Gottsche ...........................................................................................................92 Scapania javanica var. scabra Schifn. ............................................................................................92 Scapania karl-muelleri Grolle ........................................................................................................90 Scapania kaurinii Ryan .................................................................................................................88 Scapania komagadakensis Amakawa ...............................................................................................91 Scapania koponenii Potemkin ........................................................................................................88 Scapania lepida Mitt. ....................................................................................................................88 Scapania ligulata Steph. ................................................................................................................92 Scapania ligulata subsp. stephanii (Müll.Frib.) Potemkin, Piippo et T.J.Kop.................................92 Scapania ligulifolia R.M.Schust. ....................................................................................................90 Scapania lingulata H.Buch ............................................................................................................89 Scapania lingulata var. microphylla (Warnst.) R.M.Schust. ............................................................89 Scapania macroparaphyllia T.Cao, C.Gao et J.Sun ........................................................................85 Scapania magadanica S.S.Choi, Bakalin et B.Y.Sun.......................................................................89 Scapania matveyevae Potemkin ......................................................................................................91 Scapania maxima Horik. ...............................................................................................................85 Scapania microdonta (Mitt.) Müll.Frib. .........................................................................................87 Scapania mucronata H.Buch .........................................................................................................89 Scapania nemorea (L.) Grolle ........................................................................................................91 Scapania nimbosa Taylor ...............................................................................................................86 Scapania nipponica (Amakawa et S.Hatt.) Amakawa .....................................................................85 Scapania obcordata (Berggr.) S.W.Arnell .......................................................................................89 Scapania obscura (Arnell et C.E.O.Jensen) Schifn. .......................................................................91 Scapania orientalis Steph. ex Müll.Frib..........................................................................................87 Scapania ornithopodioides (With.) Waddell ...................................................................................86 Scapania paludicola Loeske et Müll.Frib. .......................................................................................90 Scapania paludicola var. viridigemma R.M.Schust. ........................................................................90 World checklist of hornworts and liverworts ** ** ** ** *** *** ** ** ** *** *** *** *** *** *** *** *** ** ** ** * *** *** *** *** *** *** *** *** *** ** ** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** 653 Scapania paludosa (Müll.Frib.) Müll.Frib. .....................................................................................91 Scapania paraphyllia T.Cao, C.Gao, J.Sun et B.R.Zuo..................................................................85 Scapania parvidens Steph...............................................................................................................91 Scapania parvifolia Warnst. ...........................................................................................................89 Scapania parvitexta Steph. .............................................................................................................91 Scapania portoricensis Hampe et Gottsche .....................................................................................86 Scapania portoricensis var. boliviensis (Steph.) Herzog ....................................................................86 Scapania portoricensis var. organensis (Herzog) Herzog ...................................................................86 Scapania portoricensis var. roraimensis Warnst. ...............................................................................86 Scapania praetervisa Meyl. .............................................................................................................89 Scapania pseudocalcicola R.M.Schust. ............................................................................................90 Scapania pseudocontorta Potemkin.................................................................................................87 Scapania rigida Nees .....................................................................................................................91 Scapania rotundifolia W.E.Nicholson............................................................................................86 Scapania ruidula Warnst. .............................................................................................................91 Scapania sandei Schifn. ex Müll.Frib. ...........................................................................................88 Scapania scandica (Arnell et H.Buch) Macvicar .............................................................................89 Scapania scandica var. argutedentata H.Buch .................................................................................89 Scapania scandica var. dimorpha R.M.Schust. ................................................................................89 Scapania scandica var. grandiretis (Schljakov) Schljakov.................................................................89 Scapania scapanioides (C.Massal.) Grolle .......................................................................................92 Scapania schljakovii Potemkin .......................................................................................................90 Scapania secunda Steph. ................................................................................................................86 Scapania serrulata R.M.Schust. .....................................................................................................91 Scapania simmonsii Bryhn et Kaal. ................................................................................................92 Scapania sinikkae Potemkin ..........................................................................................................87 Scapania sphaerifera H.Buch et Tuom...........................................................................................92 Scapania spiniloba Potemkin .........................................................................................................87 Scapania spitsbergensis (Lindb.) Müll.Frib......................................................................................88 Scapania subalpina (Nees ex Lindenb.) Dumort. ...........................................................................91 Scapania subalpina var. haynesiae Frye et L.Clark ..........................................................................91 Scapania subalpina var. muddiae C.D.Bird et W.S.Hong ..............................................................91 Scapania subnimbosa Steph. ..........................................................................................................85 Scapania tundrae (Arnell) H.Buch.................................................................................................90 Scapania udarii S.C.Srivast. et A.Srivast. .......................................................................................92 Scapania uliginosa (Lindenb.) Dumort. .........................................................................................89 Scapania umbrosa (Schrad.) Dumort. ............................................................................................87 Scapania undulata (L.) Dumort. ...................................................................................................91 Scapania valdonii Váňa, Bedn.-Ochyra et Cykowska.....................................................................89 Scapania verrucosa Heeg ................................................................................................................92 Scapania zemliae S.W.Arnell .........................................................................................................89 Scapania zhukovae Potemkin.........................................................................................................86 Schifneria hyalina Steph. ..............................................................................................................64 Schifneriolejeunea altimontana Vanden Berghen .........................................................................411 Schifneriolejeunea amazonica Gradst. ..........................................................................................411 Schifneriolejeunea cumingiana (Mont.) Gradst. ...........................................................................412 Schifneriolejeunea ferruginea (Steph.) Gradst. .............................................................................412 Schifneriolejeunea fragilis Gradst. et E.W.Jones...........................................................................411 Schifneriolejeunea madagascariensis (Steph.) Gradst. ...................................................................411 Schifneriolejeunea occulta (Steph.) Gradst. ..................................................................................412 Schifneriolejeunea omphalanthoides Verd. ....................................................................................412 Schifneriolejeunea parviloba (Steph.) Gradst. ..............................................................................411 654 *** *** *** *** *** *** *** *** * * ** *** *** *** * *** ** *** *** * *** *** *** *** *** ** ** *** *** *** *** *** *** *** ** *** ** ** *** *** ** ** *** ** *** ** ** ** *** *** ** *** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Schifneriolejeunea tumida (Nees) Gradst. ....................................................................................412 Schifneriolejeunea tumida var. haskarliana (Gottsche) Gradst. et Terken.....................................412 Schifneriolejeunea nymannii (Steph.) Gradst. et Terken ..............................................................412 Schifneriolejeunea pappeana (Nees) Gradst..................................................................................411 Schifneriolejeunea pappeana var. bidentata Gradst. et Vanden Berghen .......................................411 Schifneriolejeunea pappeana var. integra Gradst. et Vanden Berghen ...........................................411 Schifneriolejeunea polycarpa (Nees) Gradst. .................................................................................412 Schifneriolejeunea pulopenangensis (Gottsche) Gradst. .................................................................412 Schisma orizabense (Gottsche) Steph. ..........................................................................................511 Schisma uleanum Steph. ..............................................................................................................511 Schistochila acuminata Steph. ......................................................................................................259 Schistochila aequiloba Steph.........................................................................................................259 Schistochila alata (Lehm.) Schifn. ...............................................................................................259 Schistochila aligera (Nees et Blume) J.B.Jack et Steph. .................................................................259 Schistochila aligera var. laxa (Nees) Schifn. .................................................................................259 Schistochila altissima E.A.Hodgs. .................................................................................................259 Schistochila altissima subsp. polystratosa R.M.Schust. et J.J.Engel .................................................259 Schistochila antara Grolle ............................................................................................................259 Schistochila appendiculata (Hook.) Dumort. ex Trevis. ................................................................259 Schistochila baileyana Steph. ........................................................................................................259 Schistochila balfouriana (Hook.f. et Taylor) Steph. ......................................................................259 Schistochila beccariana (De Not.) Trevis. .....................................................................................260 Schistochila berggrenii (J.J.Engel et R.M.Schust.) Xiao L.He et Glenny .......................................260 Schistochila berteroana (Hook.) Steph. .........................................................................................260 Schistochila blumei (Nees) Trevis. ................................................................................................260 Schistochila brassii Grolle.............................................................................................................260 Schistochila caledonica Steph. .......................................................................................................260 Schistochila carnosa (Mitt.) Steph. ...............................................................................................260 Schistochila caudata R.M.Schust. et J.J.Engel ..............................................................................260 Schistochila childii (R.M.Schust. et J.J.Engel) Xiao L.He et Glenny ............................................260 Schistochila chlorophylla (Colenso) E.A.Hodgs. ............................................................................260 Schistochila ciliata (Mitt.) Steph. .................................................................................................260 Schistochila compacta (Colenso) E.A.Hodgs. ................................................................................260 Schistochila conchophylla Herzog ex E.A.Hodgs. et Allison...........................................................260 Schistochila conchophylla var. multidentata (J.J.Engel) Xiao L.He et Glenny.................................260 Schistochila congoana Steph. ........................................................................................................260 Schistochila cookei (H.A.Mill.) R.M.Schust..................................................................................260 Schistochila crinita Grolle ............................................................................................................260 Schistochila cristata Steph. ...........................................................................................................261 Schistochila cunninghamii Steph. .................................................................................................261 Schistochila doriae (De Not.) Trevis. ...........................................................................................261 Schistochila engleriana Steph........................................................................................................261 Schistochila exalata Herzog ..........................................................................................................261 Schistochila ijiensis H.Buch et Herzog.........................................................................................261 Schistochila glaucescens (Hook.) A.Evans ......................................................................................261 Schistochila hattorii Grolle ...........................................................................................................261 Schistochila integerrima Steph. .....................................................................................................261 Schistochila isotachyphylla (J.J.Engel et R.M.Schust.) Xiao L.He et Glenny..................................261 Schistochila kirkiana Steph. .........................................................................................................261 Schistochila kunkelii S.W.Arnell ..................................................................................................261 Schistochila lacerata Steph. ..........................................................................................................261 Schistochila lamellata (Hook.) Dumort. ex A.Evans .....................................................................261 World checklist of hornworts and liverworts *** *** *** *** ** ** *** *** *** ** *** *** *** *** *** *** *** *** *** *** *** *** *** ** *** *** *** *** ** ** *** *** *** *** *** *** ** *** ** *** *** *** *** *** *** *** *** ** *** ** *** *** 655 Schistochila laminigera (Hook.f. et Taylor) A.Evans ....................................................................261 Schistochila latiloba (R.M.Schust. et J.J.Engel) Xiao L.He et Glenny ...........................................261 Schistochila lehmanniana (Lindenb.) Steph. .................................................................................261 Schistochila leucophylla (Lehm. ex Gottsche, Lindenb. et Nees) Steph. ........................................261 Schistochila macrodonta W.E.Nicholson ......................................................................................261 Schistochila minor C.Gao et Y.H.Wu ..........................................................................................261 Schistochila monticola R.M.Schust. ..............................................................................................262 Schistochila muricata E.A.Hodgs. et Allison.................................................................................262 Schistochila nadeaudiana Steph....................................................................................................262 Schistochila neesii (Mont.) Lindb. ................................................................................................262 Schistochila nitidissima R.M.Schust. ............................................................................................262 Schistochila nivicola (R.M.Schust. et J.J.Engel) Xiao L.He et Glenny ..........................................262 Schistochila nobilis (Hook.) Trevis. ..............................................................................................262 Schistochila nuda Horik. ..............................................................................................................262 Schistochila pachyphylla (Lehm.) Steph. .......................................................................................262 Schistochila papillifera R.M.Schust. .............................................................................................262 Schistochila parvistipula Rodway..................................................................................................262 Schistochila pellucida R.M.Schust. et J.J.Engel .............................................................................262 Schistochila piligera Steph. ...........................................................................................................262 Schistochila pinnatifolia (Hook.) Trevis. ......................................................................................262 Schistochila pluriciliata R.M.Schust. et J.J.Engel..........................................................................262 Schistochila pseudociliata R.M.Schust. .........................................................................................262 Schistochila quadriida A.Evans....................................................................................................262 Schistochila ramentacea Steph. .....................................................................................................262 Schistochila relexa (Mont.) Steph. ...............................................................................................262 Schistochila relexistipula J.J.Engel et R.M.Schust. .......................................................................263 Schistochila reinwardtii (Nees) Schifn. ........................................................................................263 Schistochila repleta (Hook.f. et Taylor) Steph. .............................................................................263 Schistochila rubriseta Steph. .........................................................................................................263 Schistochila schultzei Steph. .........................................................................................................263 Schistochila sciophila R.M.Schust. ................................................................................................263 Schistochila sciurea (Nees) Schifn................................................................................................263 Schistochila simulans (C.Massal.) Xiao L.He et Yu Sun ................................................................263 Schistochila spegazziniana (C.Massal.) Steph................................................................................263 Schistochila sphagnoides (Schwägr.) Lindb. ex Steph.....................................................................263 Schistochila splachnophylla (Hook.f. et Taylor) Steph. ..................................................................263 Schistochila stratosa (Mont.) A.Evans ...........................................................................................263 Schistochila subhyalina R.M.Schust. ............................................................................................263 Schistochila subhyalina var. grandidentata (J.J.Engel et R.M.Schust.) Xiao L.He et Glenny..........263 Schistochila subimmersa J.J.Engel et R.M.Schust..........................................................................263 Schistochila succulenta (J.J.Engel et R.M.Schust.) Xiao L.He et Glenny .......................................263 Schistochila tasmanica Steph. .......................................................................................................264 Schistochila trispiralis R.M.Schust................................................................................................264 Schistochila tuloides (Hook.f. et Taylor) Steph. ............................................................................264 Schistochila undulatifolia Piippo ..................................................................................................264 Schistochila virescens R.M.Schust. ................................................................................................264 Schistochila vitreocincta (Herzog) Xiao L.He et Glenny ...............................................................264 Schistochila volans Grolle .............................................................................................................264 Schistochila yakushimensis N.Ohnishi et Deguchi ........................................................................264 Schistochila zantenii Grolle ..........................................................................................................264 Schistochilopsis cornuta (Steph.) Konstant. .....................................................................................93 Schistochilopsis grandiretis (Lindb. ex Kaal.) Konstant. ...................................................................93 656 ** *** * ** *** ** *** ** *** *** *** *** *** *** *** ** ** *** *** *** ** *** *** *** *** *** *** *** *** *** *** *** *** ** *** *** * *** *** *** *** *** *** *** *** *** *** ** *** *** *** *** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Schistochilopsis hyperarctica Konstant. et L.Söderstr. ......................................................................93 Schistochilopsis incisa (Schrad.) Konstant. ......................................................................................93 Schistochilopsis nakanishii (Inoue) Konstant...................................................................................93 Schistochilopsis opacifolia (Culm. ex Meyl.) Konstant. ....................................................................93 Schistochilopsis setosa (Mitt.) Konstant. ..........................................................................................93 Schizophyllopsis aristata (Herzog ex N.Kitag.) Váňa et L.Söderstr. .................................................54 Schizophyllopsis bidens (Reinw., Blume et Nees) Váňa et L.Söderstr...............................................54 Schizophyllopsis lanciloba (Steph.) Váňa et L.Söderstr. ...................................................................54 Schizophyllopsis papillosa (J.J.Engel et Braggins) Váňa et L.Söderstr. ..............................................54 Schizophyllopsis sphenoloboides (R.M.Schust.) Váňa et L.Söderstr. ..................................................54 Schljakovia kunzeana (Huebener) Konstant. et Vilnet ...................................................................55 Schljakovianthus quadrilobus (Lindb.) Konstant. et Vilnet .............................................................55 Schusterolejeunea inundata (Spruce) Grolle ..................................................................................350 Seppeltia succuba Grolle...............................................................................................................471 Sewardiella tuberifera Kashyap ....................................................................................................468 Siphonolejeunea elegantissima (Steph.) Grolle ...............................................................................350 Siphonolejeunea neesii (Mont.) Bischl. .........................................................................................350 Siphonolejeunea schifneri (Schifn.) Herzog .................................................................................350 Solenostoma amoenum (Lindenb. et Gottsche) R.M.Schust. ex Váňa, Hentschel et Heinrichs......129 Solenostoma amplexifolium (Hampe) Váňa et Schäf.-Verw. ..........................................................129 Solenostoma appalachianum R.M.Schust. ex Bakalin....................................................................129 Solenostoma appressifolium (Mitt.) Váňa et D.G.Long .................................................................129 Solenostoma appressifolium var. minor (Amakawa) Váňa et D.G.Long ..........................................129 Solenostoma appressifolium var. nigricans (Amakawa) Váňa et D.G.Long .....................................129 Solenostoma ariadne (Taylor) R.M.Schust. ex Váňa et D.G.Long ................................................129 Solenostoma atrobrunneum (Amakawa) Váňa et D.G.Long ..........................................................129 Solenostoma atrorevolutum (Grolle ex Amakawa) Váňa et D.G.Long ...........................................129 Solenostoma atrovirens Steph........................................................................................................129 Solenostoma balfourii (Váňa) Váňa, Hentschel et Heinrichs .........................................................125 Solenostoma baueri (Schifn.) Steph. ............................................................................................130 Solenostoma bengalense (Amakawa) Váňa et D.G.Long ................................................................130 Solenostoma bilobum (S.Hatt. ex Amakawa) Potemkin et Nyushko .............................................123 Solenostoma borneense (Amakawa) Váňa, Hentschel et Heinrichs.................................................125 Solenostoma brevilorum Kashyap et R.S.Chopra..........................................................................130 Solenostoma caeleste (Inoue et Váňa) Váňa, Hentschel et Heinrichs .............................................130 Solenostoma callithrix (Lindenb. et Gottsche) Steph. ...................................................................125 Solenostoma caoi (C.Gao et X.L.Bai) Váňa et D.G.Long..............................................................130 Solenostoma caucasicum (Váňa) Konstant.....................................................................................130 Solenostoma champawatense (S.N.Srivast. et Amakawa) Váňa et D.G.Long .................................125 Solenostoma chenianum (C.Gao, Y.H.Wu et Grolle) Váňa et D.G.Long......................................130 Solenostoma clavellatum Mitt. ex Steph. .......................................................................................130 Solenostoma comatum (Nees) C.Gao ............................................................................................125 Solenostoma comatum var. novae-guineae (Váňa) Váňa, Hentschel et Heinrichs ...........................125 Solenostoma confertissimum (Nees) Schljakov ...............................................................................130 Solenostoma conilorum (Schifn.) Steph.......................................................................................130 Solenostoma crassulum (Nees et Mont.) Steph. .............................................................................130 Solenostoma crenuliforme (Austin) Steph. .....................................................................................125 Solenostoma cryptogynum R.M.Schust. ex J.J.Engel ......................................................................130 Solenostoma cyclops (S.Hatt.) R.M.Schust. ...................................................................................130 Solenostoma decolor (Schifn.) R.M.Schust. ex Váňa, Hentschel et Heinrichs ...............................125 Solenostoma diversiclavellatum (Amakawa et Grolle) R.M.Schust. ex Váňa et D.G.Long .............130 Solenostoma dulongense Váňa et D.G.Long ..................................................................................130 World checklist of hornworts and liverworts *** *** *** *** *** * *** *** *** *** *** *** *** * *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** ** *** ** *** ** *** * *** *** *** *** * * *** ** * ** *** * 657 Solenostoma dusenii (Steph.) Váňa, Hentschel et Heinrichs .........................................................125 Solenostoma emarginatum (Amakawa) Váňa, Hentschel et Heinrichs ...........................................124 Solenostoma erectum (Amakawa) C.Gao.......................................................................................125 Solenostoma exsertum (A.Evans) Steph. ........................................................................................131 Solenostoma faurieanum (Beauverd) R.M.Schust. ........................................................................131 Solenostoma lagellalioides C.Gao .................................................................................................125 Solenostoma lagellare (Amakawa) Váňa et D.G.Long ..................................................................131 Solenostoma lagellatum (S.Hatt.) Váňa et D.G.Long ...................................................................124 Solenostoma lavialbicans (Amakawa et Grolle) Váňa et D.G.Long ..............................................126 Solenostoma lavorevolutum (Váňa) Váňa et D.G.Long ................................................................131 Solenostoma fossombronioides (Austin) R.M.Schust. .....................................................................126 Solenostoma fusiforme (Steph.) R.M.Schust..................................................................................124 Solenostoma glaucum (Amakawa) Váňa et D.G.Long ...................................................................126 Solenostoma gongshanense (C.Gao et J.Sun) Váňa et D.G.Long....................................................126 Solenostoma gracillimum (Sm.) R.M.Schust. ................................................................................124 Solenostoma grollei (D.G.Long et Váňa) K.Feldberg, Hentschel, Bombosch, D.G.Long, Váňa et Heinrichs ................................................................................................................................... 131 Solenostoma grosseverrucosum (Amakawa et S.Hatt.) Váňa, Crand.-Stotl. et Stotler ......................131 Solenostoma handelii (Schifn.) Müll.Frib. ...................................................................................124 Solenostoma haskarlianum (Nees) R.M.Schust. ex Váňa et D.G.Long ..........................................126 Solenostoma hattorianum (Amakawa) Potemkin et Nyushko........................................................126 Solenostoma heterolimbatum (Amakawa) Váňa et D.G.Long ........................................................131 Solenostoma hewsoniae (Amakawa et Grolle) R.M.Schust. ex Váňa, Hentschel et Heinrichs ........131 Solenostoma hirticalyx (Steph.) R.M.Schust. ex Váňa, Hentschel et Heinrichs .............................126 Solenostoma hiugaense Amakawa ..................................................................................................131 Solenostoma hokkaidense (Váňa) Váňa, Hentschel et Heinrichs ....................................................124 Solenostoma horikawanum (Amakawa) Váňa, Hentschel et Heinrichs ..........................................126 Solenostoma hyalinum (Lyell) Mitt...............................................................................................126 Solenostoma indrodayanum (Sushil K.Singh et D.K.Singh) Váňa et D.G.Long ............................131 Solenostoma infuscum (Mitt.) Hentschel ......................................................................................126 Solenostoma infuscum var. ovicalyx (Steph.) Potemkin et Sofronova .............................................126 Solenostoma inundatum (Hook.f. et Taylor) Mitt. ex Steph. ........................................................131 Solenostoma javanicum (Schifn.) Steph. ......................................................................................131 Solenostoma kanaii (Amakawa) Váňa et D.G.Long ......................................................................131 Solenostoma kashyapii (S.C.Srivast., S.Srivast. et D.Sharma) Váňa et D.G.Long ..........................131 Solenostoma kurilense (Bakalin) Váňa...........................................................................................126 Solenostoma lanigerum (Mitt.) Váňa et D.G.Long .......................................................................132 Solenostoma lignicola (Schifn.) Váňa, Hentschel et Heinrichs .....................................................124 Solenostoma limbatifolium (Amakawa) Váňa et D.G.Long ...........................................................124 Solenostoma lixingjiangii (C.Gao et X.L.Bai) Váňa et D.G.Long .................................................126 Solenostoma macrocarpum (Schifn. ex Steph.) Váňa et D.G.Long ...............................................132 Solenostoma mamatkulovii (Váňa et Zerov) Váňa, Hentschel et Heinrichs ...................................132 Solenostoma marginatum (S.Hatt.) R.M.Schust. ..........................................................................126 Solenostoma micranthum (Mitt.) Váňa, Hentschel et Heinrichs ...................................................127 Solenostoma microphyllum C.Gao ................................................................................................132 Solenostoma microrevolutum (C.Gao et X.L.Bai) Váňa et D.G.Long ............................................132 Solenostoma mildbraedii (Steph.) R.M.Schust. .............................................................................132 Solenostoma montanum (Steph.) Váňa .........................................................................................127 Solenostoma multicarpum (C.Gao et J.Sun) Váňa et D.G.Long ...................................................132 Solenostoma nilgiriense (A.Alam, Ad.Kumar et S.C.Srivast.) Váňa et D.G.Long...........................127 Solenostoma niveum (Grolle) R.M.Schust. ex Váňa, Hentschel et Heinrichs ................................132 Solenostoma novazelandiae R.M.Schust........................................................................................132 658 *** *** *** *** *** * *** *** *** *** *** *** ** *** *** *** *** *** ** *** *** *** ** *** *** *** *** *** *** *** *** *** *** * *** *** *** *** *** *** ** *** *** *** *** *** *** *** *** * Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Solenostoma obliquifolium (Schifn.) R.M.Schust. ex Váňa, Hentschel et Heinrichs .....................127 Solenostoma obovatum (Nees) C.Massal. ......................................................................................124 Solenostoma obscurum (A.Evans) R.M.Schust. .............................................................................124 Solenostoma ohbae (Amakawa) C.Gao .........................................................................................132 Solenostoma onraedtii (Váňa) Váňa, Hentschel et Heinrichs ........................................................127 Solenostoma orbicularifolium (Piippo ex C.Gao et Bai) Váňa .......................................................127 Solenostoma orbiculatum (Colenso) R.M.Schust. .........................................................................132 Solenostoma otianum (S.Hatt.) R.M.Schust. ................................................................................127 Solenostoma ovalifolium (Amakawa) Váňa ...................................................................................127 Solenostoma paroicum (Schifn.) R.M.Schust. ..............................................................................127 Solenostoma parvitextum (Amakawa) Váňa et D.G.Long .............................................................132 Solenostoma patoniae (Grolle, D.B.Schill et D.G.Long) K.Feldberg, Hentschel, Bombosch, D.G.Long, Váňa et Heinrichs .....................................................................................................132 Solenostoma philippinense Váňa....................................................................................................124 Solenostoma plagiochilaceum (Grolle) Váňa et D.G.Long .............................................................127 Solenostoma pocsii (Váňa) Bakalin................................................................................................132 Solenostoma poeltii (Amakawa) Váňa et D.G.Long ......................................................................133 Solenostoma polyrhizoides (Grolle ex Amakawa) Váňa et D.G.Long .............................................127 Solenostoma pseudocyclops (Inoue) Váňa et D.G.Long ..................................................................133 Solenostoma pseudopyrilorum Bakalin et Vilnet............................................................................133 Solenostoma purpuratum (Mitt.) Steph. .......................................................................................133 Solenostoma pyrilorum Steph.......................................................................................................133 Solenostoma pyrilorum var. gracillimum (Amakawa) Váňa et D.G.Long ......................................133 Solenostoma pyrilorum var. major (S.Hatt.) Bakalin.....................................................................133 Solenostoma pyrilorum var. minutissimum (Amakawa) Bakalin ....................................................133 Solenostoma radicellosum Mitt. ....................................................................................................127 Solenostoma raujeanum (Grolle ex Amakawa) Váňa et D.G.Long ................................................133 Solenostoma renauldii (Steph.) Váňa, Hentschel et Heinrichs ......................................................127 Solenostoma ricleii Váňa et D.G.Long.........................................................................................133 Solenostoma rigidulum (S.Hatt.) R.M.Schust. ..............................................................................127 Solenostoma rosulans (Steph.) Váňa et D.G.Long .........................................................................127 Solenostoma rotundatum Amakawa ..............................................................................................127 Solenostoma rubripunctatum (S.Hatt.) R.M.Schust. .....................................................................127 Solenostoma rubrum (Gottsche) R.M.Schust................................................................................125 Solenostoma rupicola (Amakawa) Váňa et D.G.Long ...................................................................133 Solenostoma sanguinolentum (Grif.) Steph...................................................................................133 Solenostoma schaulianum (Steph.) Váňa et D.G.Long ..................................................................133 Solenostoma schusteranum (J.D.Godfrey et G.Godfrey) Váňa, Hentschel et Heinrichs .................124 Solenostoma shimizuanum (S.Hatt. ex Váňa) Váňa, Hentschel et Heinrichs .................................133 Solenostoma sikkimense (Schifn. ex Steph.) Váňa et D.G.Long ....................................................127 Solenostoma speciosum (Horik.) Hentschel, K.Feldberg, Bombosch, D.G.Long, Váňa et Heinrichs....133 Solenostoma speciosum subsp. villosum (R.M.Schust.) Hentschel, K.Feldberg, Bombosch, D.G.Long, Váňa et Heinrichs .......................................................................................................................134 Solenostoma sphaerocarpum (Hook.) Steph. .................................................................................134 Solenostoma stephanii (Schifn.) Steph. ........................................................................................134 Solenostoma stoloniferum (Steph.) S.W.Arnell ..............................................................................134 Solenostoma strictum (Schifn.) Váňa, Hentschel et Heinrichs......................................................134 Solenostoma subacutum (Herzog) Váňa, Crand.-Stotl. et Stotler ..................................................134 Solenostoma suborbiculatum (Amakawa) Váňa et D.G.Long ........................................................125 Solenostoma subrubrum (Schifn. ex Steph.) Váňa et D.G.Long ...................................................134 Solenostoma subtilissimum (Schifn.) R.M.Schust. ........................................................................124 Solenostoma sunii Bakalin et Vilnet..............................................................................................134 World checklist of hornworts and liverworts *** * *** * *** ** *** *** *** * *** *** * *** * *** *** *** *** *** *** *** *** *** *** ** *** ** *** *** *** *** *** ** *** *** *** *** *** *** *** *** * *** *** *** *** *** *** *** *** *** 659 Solenostoma tetragonum (Lindenb.) R.M.Schust. ex Váňa et D.G.Long .......................................127 Solenostoma tetragonum var. kodaikanalense A.Alam, D.Sharma et So.Yadav................................127 Solenostoma torticalyx (Steph.) C.Gao..........................................................................................134 Solenostoma totopapillosum (E.A.Hodgs.) R.M.Schust. ................................................................134 Solenostoma truncatum (Nees) R.M.Schust. ex Váňa et D.G.Long ..............................................134 Solenostoma truncatum var. setulosum (Herzog) Váňa et D.G.Long..............................................134 Solenostoma tuberculiferum (Herzog) Váňa, Hentschel et Heinrichs ............................................127 Solenostoma udarii (S.C.Srivast. et P.Singh) Váňa et D.G.Long ..................................................134 Solenostoma unispire (Amakawa) Váňa, Hentschel et Heinrichs ...................................................127 Solenostoma ventroversum (Grolle) Váňa et D.G.Long .................................................................134 Solenostoma virgatum (Mitt.) Váňa et D.G.Long .........................................................................127 Solenostoma vulcanicola (Schifn.) Nyushko ex Potemkin et Sofronova........................................127 Solenostoma zangmuii (C.Gao et X.L.Bai) Váňa et D.G.Long .....................................................127 Solenostoma zantenii (Amakawa) R.M.Schust. ex Váňa et D.G.Long ..........................................134 Solenostoma zengii (C.Gao et X.L.Bai) Váňa et D.G.Long ...........................................................134 Southbya gollanii Steph. ..............................................................................................................135 Southbya nigrella (De Not.) Henriq. ...........................................................................................135 Southbya organensis Herzog .........................................................................................................136 Southbya tophacea (Spruce) Spruce ..............................................................................................136 Sphaerocarpos cristatus M.Howe ..................................................................................................506 Sphaerocarpos donnellii Austin .....................................................................................................506 Sphaerocarpos drewiae Wigglesw. .................................................................................................506 Sphaerocarpos europaeus Lorb. .....................................................................................................507 Sphaerocarpos hians Haynes .........................................................................................................507 Sphaerocarpos michelii Bellardi ....................................................................................................506 Sphaerocarpos muccilloi E.Vianna ................................................................................................507 Sphaerocarpos stipitatus Bisch. ex Lindenb. ..................................................................................506 Sphaerocarpos texanus Austin .......................................................................................................506 Sphenolobopsis pearsonii (Spruce) R.M.Schust................................................................................55 Sphenolobus austroamericanus (Váňa) Váňa....................................................................................55 Sphenolobus minutus (Schreb. ex D.Crantz) Berggr. ......................................................................55 Sphenolobus saxicola (Schrad.) Steph..............................................................................................55 Spruceanthus macrostipulus (Steph.) Gradst. ................................................................................412 Spruceanthus mamillilobulus (Herzog) Verd.................................................................................412 Spruceanthus pluriplicatus (Steph.) Gradst. ..................................................................................412 Spruceanthus polymorphus (Sande Lac.) Verd. ..............................................................................412 Spruceanthus semirepandus (Nees) Verd. ......................................................................................412 Spruceanthus sulcatus (Nees) Gradst. ...........................................................................................413 Spruceanthus theobromae (Spruce) Gradst. ...................................................................................413 Spruceanthus thozetianus (Gottsche et F.Muell.) B.M.hiers et Gradst. .......................................413 Steereella lilliana (Steph.) Kuwah. ...............................................................................................461 Steereella linearis (Sw.) Kuwah. ...................................................................................................461 Stephaniella boliviensis Steph. ......................................................................................................136 Stephaniella hamata Steph. ..........................................................................................................136 Stephaniella paraphyllina J.B.Jack ................................................................................................136 Stephaniella rostrata U.Schmitt ...................................................................................................136 Stephaniella uncifolia S.Winkl. ....................................................................................................136 Stephaniellidium sleumeri (Müll.Frib.) S.Winkl. ex Grolle ...........................................................136 Stephensoniella brevipedunculata Kashyap ....................................................................................488 Stictolejeunea balfourii (Mitt.) E.W.Jones ....................................................................................300 Stictolejeunea balfourii var. bekkeri Gradst. ..................................................................................300 Stictolejeunea iwatsukii Mizut......................................................................................................300 660 *** ** *** ** *** ** ** * * * * * * * * * * * *** *** *** *** *** *** *** ** ** * *** ** ** *** *** *** ** *** *** ** ** ** ** ** *** ** *** *** *** ** * *** * *** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Stictolejeunea squamata (Willd.) Schifn. .....................................................................................300 Stolonivector clasmatocoleoides J.J.Engel .......................................................................................127 Stolonivector iordlandiae (E.A.Hodgs.) J.J.Engel .........................................................................127 Stolonivector iordlandiae var. nodulosus J.J.Engel .........................................................................127 Stolonivector gremmenii (Váňa) Váňa ...........................................................................................127 Stolonivector obtusilobus J.J.Engel ................................................................................................127 Stolonivector waipouensis J.J.Engel ...............................................................................................127 Strepsilejeunea apollinea (Gottsche) Steph....................................................................................512 Strepsilejeunea durelii Schifn.......................................................................................................512 Strepsilejeunea hamatifolia Steph. ................................................................................................512 Strepsilejeunea lanceolata (Gottsche) Steph. .................................................................................512 Strepsilejeunea muscicola Herzog ..................................................................................................512 Strepsilejeunea novae-guineae Steph..............................................................................................512 Strepsilejeunea obtusistipula Steph. ...............................................................................................512 Strepsilejeunea papulifolia Steph. .................................................................................................512 Strepsilejeunea pectiniformis (Gottsche) Steph. .............................................................................512 Strepsilejeunea renistipula Steph. ..................................................................................................512 Strepsilejeunea vatovae Gerola ......................................................................................................512 Symbiezidium barbilorum (Lindenb. et Gottsche) A.Evans .........................................................398 Symbiezidium dentatum Herzog ..................................................................................................398 Symbiezidium madagascariense Steph. ..........................................................................................398 Symbiezidium transversale (Sw.) Trevis. .......................................................................................398 Symbiezidium transversale var. hookerianum (Gottsche, Lindenb. et Nees) Gradst. et J.Beek .......398 Symphyogyna apiculispina Steph. .................................................................................................471 Symphyogyna aspera Steph. ex F.A.McCormick ...........................................................................471 Symphyogyna atronervia Taylor....................................................................................................471 Symphyogyna bogotensis Steph. .....................................................................................................471 Symphyogyna boliviensis Steph. ....................................................................................................471 Symphyogyna brasiliensis Nees et Mont. .......................................................................................471 Symphyogyna brasiliensis var. angustior (Gottsche, Lindenb. et Nees) Gottsche ............................471 Symphyogyna brasiliensis var. subsinuata Schifn. ..........................................................................472 Symphyogyna brongniartii Mont. .................................................................................................472 Symphyogyna circinata Nees et Mont. ..........................................................................................472 Symphyogyna digitisquama Steph. ................................................................................................472 Symphyogyna fuscovirens A.Evans .................................................................................................472 Symphyogyna hochstetteri Nees et Mont. ......................................................................................472 Symphyogyna hymenophyllum (Hook.) Nees et Mont. ..................................................................472 Symphyogyna hymenophyllum var. heterogenum Spruce .................................................................472 Symphyogyna ignambiensis Hürl. ..................................................................................................472 Symphyogyna interrupta Carrington et Pearson ............................................................................472 Symphyogyna irregularis Steph. ....................................................................................................472 Symphyogyna lacerosquama Steph. ...............................................................................................472 Symphyogyna leptothelia Taylor ....................................................................................................472 Symphyogyna lindmanii A.Evans ..................................................................................................472 Symphyogyna luetzelburgii Herzog ...............................................................................................472 Symphyogyna marginata Steph. ....................................................................................................472 Symphyogyna mexicana Steph. .....................................................................................................472 Symphyogyna multilora Steph. ....................................................................................................472 Symphyogyna paucidens Steph. .....................................................................................................472 Symphyogyna podophylla (hunb.) Nees et Mont. ........................................................................472 Symphyogyna purpureolimbata E.A.Hodgs. ..................................................................................473 Symphyogyna rectidens Grolle .......................................................................................................473 World checklist of hornworts and liverworts ** *** *** ** ** *** *** *** *** * *** ** *** *** *** *** *** *** ** *** ** *** *** *** *** *** *** *** *** *** ** *** *** *** *** *** *** *** * *** *** *** *** *** *** *** *** *** *** * * * 661 Symphyogyna rhodina (Hook.f. et Taylor) Gottsche, Lindenb. et Nees ........................................473 Symphyogyna rubescens Steph. ......................................................................................................473 Symphyogyna rubritincta A.Evans ................................................................................................473 Symphyogyna semi-involucrata Austin ..........................................................................................473 Symphyogyna similis Grolle ..........................................................................................................473 Symphyogyna sinuata (Sw.) Nees et Mont. ...................................................................................473 Symphyogyna subsimplex Mitt. .....................................................................................................473 Symphyogyna tenuinervis (Hook.f. et Taylor) Grolle ....................................................................473 Symphyogyna trivittata Spruce .....................................................................................................473 Symphyogyna ulvoides (Reinw., Blume et Nees) Nees ...................................................................473 Symphyogyna undulata Colenso ...................................................................................................473 Symphyogyna volkensii Steph........................................................................................................473 Symphyogynopsis gottscheana (Mont. et Nees) Grolle ....................................................................473 Syzygiella acinacifolia (Hook.f. et Taylor) K.Feldberg, Váňa, Hentschel et Heinrichs ....................47 Syzygiella anomala (Lindenb. et Gottsche) Steph. ..........................................................................47 Syzygiella autumnalis (DC.) K.Feldberg, Váňa, Hentschel et Heinrichs .........................................48 Syzygiella bilobata Inoue................................................................................................................47 Syzygiella campanulata Herzog ......................................................................................................48 Syzygiella ciliata Gradst. et A.R.Benitez .........................................................................................47 Syzygiella colorata (Lehm.) K.Feldberg, Váňa, Hentschel et Heinrichs ...........................................48 Syzygiella colorata var. collenchymata J.J.Engel et Váňa ..................................................................48 Syzygiella concreta (Gottsche) Spruce .............................................................................................47 Syzygiella contigua Steph. ..............................................................................................................48 Syzygiella eatonii (Austin) Inoue ....................................................................................................49 Syzygiella elongella (Taylor) K.Feldberg, Váňa, Hentschel et Heinrichs .........................................48 Syzygiella jacquinotii (Mont.) Hentschel, K.Feldberg, Váňa et Heinrichs ......................................48 Syzygiella macrocalyx (Mont.) Spruce ............................................................................................48 Syzygiella manca (Mont.) Steph.....................................................................................................47 Syzygiella nigrescens (Steph.) K.Feldberg, Váňa, Hentschel et Heinrichs ........................................47 Syzygiella nipponica (S.Hatt.) K.Feldberg, Váňa, Hentschel et Heinrichs ......................................48 Syzygiella oenops (Lindenb. et Gottsche) K.Feldberg, Váňa, Hentschel et Heinrichs ......................49 Syzygiella ovalifolia Inoue ..............................................................................................................47 Syzygiella paludosa (Steph.) K.Feldberg, Váňa, Hentschel et Heinrichs..........................................47 Syzygiella pectiniformis Spruce .......................................................................................................47 Syzygiella perfoliata (Sw.) Spruce ...................................................................................................49 Syzygiella pseudocclusa (E.A.Hodgs.) K.Feldberg, Váňa, Hentschel et Heinrichs ............................47 Syzygiella purpurascens (Steph.) K.Feldberg, Váňa, Hentschel et Heinrichs....................................49 Syzygiella rubricaulis (Nees) Steph. ................................................................................................49 Syzygiella securifolia (Nees) Inoue ..................................................................................................48 Syzygiella setulosa Steph. ................................................................................................................49 Syzygiella sonderi (Gottsche) K.Feldberg, Váňa, Hentschel et Heinrichs ........................................47 Syzygiella spegazziniana (Spruce ex C.Massal.) K.Feldberg, Váňa, Hentschel et Heinrichs.............47 Syzygiella subintegerrima (Reinw., Blume et Nees) Spruce .............................................................48 Syzygiella tasmanica (Hook.f. et Taylor) K.Feldberg, Váňa, Hentschel et Heinrichs ......................48 Syzygiella teres (Carrington et Pearson) Váňa .................................................................................49 Syzygiella tonduzana Steph. ...........................................................................................................47 Syzygiella trigonifolia (Steph.) Herzog............................................................................................47 Syzygiella uleana Steph. .................................................................................................................49 Syzygiella undata (Mont.) K.Feldberg, Váňa, Hentschel et Heinrichs ............................................49 Targionia dioica Schifn. .............................................................................................................503 Targionia elongata Bisch..............................................................................................................503 Targionia iorii Gola ...................................................................................................................503 662 * *** ** * ** ** ** ** ** ** ** ** ** ** ** * ** ** * * * * * ** ** ** ** * ** * ** ** ** ** ** * * ** * * * ** * ** ** ** ** * * ** ** * Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Targionia formosica Horik. ..........................................................................................................503 Targionia hypophylla L. ...............................................................................................................503 Targionia hypophylla subsp. linealis W.Frey et Kürschner ............................................................503 Targionia indica Udar et A.Gupta ...............................................................................................503 Targionia lorbeeriana Müll.Frib. .................................................................................................503 Targionia stellaris (Müll.Frib.) Hässel..........................................................................................503 Taxilejeunea acutifolia (Steph.) Steph. .........................................................................................388 Taxilejeunea antillana Steph........................................................................................................388 Taxilejeunea apiculata (Gottsche) J.B.Jack et Steph. ....................................................................388 Taxilejeunea argentina Steph. ......................................................................................................388 Taxilejeunea arsenii Steph. ..........................................................................................................388 Taxilejeunea auriculata Steph. .....................................................................................................388 Taxilejeunea berteroana Steph. ....................................................................................................388 Taxilejeunea beyrichiana Steph. ...................................................................................................388 Taxilejeunea biapiculata Steph. ...................................................................................................388 Taxilejeunea boliviana Steph. ......................................................................................................388 Taxilejeunea brasiliensis Steph. ....................................................................................................388 Taxilejeunea coilantha Herzog .....................................................................................................388 Taxilejeunea compressiuscula Steph. .............................................................................................389 Taxilejeunea convoluta Herzog ....................................................................................................389 Taxilejeunea cuervi (Gottsche) Steph. ..........................................................................................389 Taxilejeunea cuneistipula Steph. ..................................................................................................389 Taxilejeunea cuspidata Steph. ......................................................................................................389 Taxilejeunea decurrens Steph. ......................................................................................................389 Taxilejeunea delexa Steph. ..........................................................................................................389 Taxilejeunea densilora A.Evans ...................................................................................................389 Taxilejeunea diaphana (Lehm.) Steph. ........................................................................................389 Taxilejeunea dissitifolia Steph. .....................................................................................................389 Taxilejeunea eggersiana Schifn. ...................................................................................................389 Taxilejeunea elobulata Sim ..........................................................................................................389 Taxilejeunea issistipula Steph. .....................................................................................................389 Taxilejeunea foliicola Steph. ........................................................................................................389 Taxilejeunea furcicornuta Grolle ..................................................................................................389 Taxilejeunea fuscorufa Steph. .......................................................................................................389 Taxilejeunea galapagensis Onr......................................................................................................389 Taxilejeunea ghatensis P.K.Verma et S.C.Srivast. .........................................................................389 Taxilejeunea giulianettii Steph. ....................................................................................................389 Taxilejeunea gomphocalyx Herzog ................................................................................................389 Taxilejeunea grandifolia Steph. ....................................................................................................389 Taxilejeunea grandistipula Steph..................................................................................................389 Taxilejeunea hamatifolia Steph. ...................................................................................................389 Taxilejeunea himalayensis Herzog ................................................................................................389 Taxilejeunea immersa Eifrig .........................................................................................................389 Taxilejeunea irregularis Steph. .....................................................................................................389 Taxilejeunea jamaicensis A.Evans .................................................................................................389 Taxilejeunea jeringii Steph. .........................................................................................................389 Taxilejeunea killipii Herzog.........................................................................................................389 Taxilejeunea laevis (Gottsche) Steph............................................................................................340 Taxilejeunea langiana Pearson .....................................................................................................340 Taxilejeunea lindbergiana Steph. .................................................................................................340 Taxilejeunea linguifolia Steph. .....................................................................................................340 Taxilejeunea maxima Steph. ........................................................................................................340 World checklist of hornworts and liverworts ** * * ** * * ** ** ** ** ** * * * ** ** * ** * *** ** ** * ** ** ** * ** ** * ** * * * ** *** *** ** *** ** *** *** *** *** *** *** *** *** *** *** *** ** 663 Taxilejeunea mexicana Steph. ......................................................................................................340 Taxilejeunea microstipula Steph. ..................................................................................................340 Taxilejeunea mucronata Steph. ....................................................................................................340 Taxilejeunea multilora Steph. .....................................................................................................340 Taxilejeunea muscicola Steph. ......................................................................................................340 Taxilejeunea nilgiriensis P.K.Verma et S.C.Srivast. ......................................................................340 Taxilejeunea nymannii Steph. ......................................................................................................340 Taxilejeunea obtusifolia Steph. .....................................................................................................340 Taxilejeunea papulilora Steph. ....................................................................................................340 Taxilejeunea parvibracteata Steph. ...............................................................................................340 Taxilejeunea parvistipula Steph....................................................................................................340 Taxilejeunea paucidens Steph. ......................................................................................................340 Taxilejeunea pendula Steph. ........................................................................................................340 Taxilejeunea peruviana Steph. .....................................................................................................340 Taxilejeunea planilobula Herzog ..................................................................................................340 Taxilejeunea pulchrilora Pearson .................................................................................................340 Taxilejeunea pusilla Steph. ..........................................................................................................340 Taxilejeunea renistipula (Lindenb.) Steph. ...................................................................................340 Taxilejeunea rufescens Steph. .......................................................................................................340 Taxilejeunea serpillifolioides (Raddi) D.P.Costa ...........................................................................340 Taxilejeunea setchellii Pearson .....................................................................................................340 Taxilejeunea speciosa Herzog .......................................................................................................340 Taxilejeunea splendida Eifrig .......................................................................................................340 Taxilejeunea stephanii Eifrig ........................................................................................................340 Taxilejeunea steyermarkii H.Rob. ................................................................................................340 Taxilejeunea surinamensis (Lindenb. et Gottsche) Steph. .............................................................340 Taxilejeunea suringarii Steph. ......................................................................................................341 Taxilejeunea tenerrima Steph.......................................................................................................341 Taxilejeunea tenuiplica Steph. .....................................................................................................341 Taxilejeunea tjibodensis (Steph.) Eifrig.........................................................................................341 Taxilejeunea tonduzana Steph. ....................................................................................................341 Taxilejeunea uleana Steph. ..........................................................................................................341 Taxilejeunea umbonata Steph. .....................................................................................................341 Taxilejeunea urbanii Steph. .........................................................................................................341 Taxilejeunea vallis-gratiae Steph. .................................................................................................341 Telaranea anomala R.M.Schust. ex J.J.Engel et G.L.Merr. ..........................................................186 Telaranea apiahyna (Steph.) Fulford ...........................................................................................185 Telaranea azorica (H.Buch et Perss.) Pócs ...................................................................................187 Telaranea bicruris (Steph.) M.Howe ...........................................................................................185 Telaranea bischleriana Pócs .........................................................................................................186 Telaranea blepharostoma (Steph.) Fulford ....................................................................................185 Telaranea breviseta (Herzog) J.J.Engel et G.L.Merr. ....................................................................185 Telaranea chaetophylla (Spruce) Schifn. ......................................................................................185 Telaranea coactilis (Spruce) J.J.Engel et G.L.Merr. ......................................................................186 Telaranea confervoides J.J.Engel et G.L.Merr. ..............................................................................186 Telaranea diacantha (Mont.) J.J.Engel et G.L.Merr. ...................................................................186 Telaranea europaea J.J.Engel et G.L.Merr. ..................................................................................185 Telaranea fragilis Mizut...............................................................................................................186 Telaranea granulata J.J.Engel et G.L.Merr. .................................................................................186 Telaranea herzogii (E.A.Hodgs.) E.A.Hodgs................................................................................186 Telaranea inaequalis R.M.Schust. ex J.J.Engel et G.L.Merr. ........................................................187 Telaranea indica (S.C.Srivast. et P.K.Verma) A.E.D.Daniels et P.Daniel ....................................187 664 *** ** ** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** ** ** ** ** ** *** *** *** *** *** ** ** *** ** ** ** *** *** *** *** *** ** *** *** *** *** *** ** *** *** ** *** *** *** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Telaranea longifolia (M.Howe) J.J.Engel et G.L.Merr. ................................................................186 Telaranea major (Herzog) J.J.Engel et G.L.Merr. ........................................................................187 Telaranea maorensis Pócs .............................................................................................................187 Telaranea microstipulata R.M.Schust. .........................................................................................187 Telaranea monocera (Mitt. ex R.M.Schust. et Grolle) J.J.Engel et G.L.Merr. ..............................187 Telaranea nematodes (Gottsche ex Austin) M.Howe ....................................................................186 Telaranea panchoi Del Ros. .........................................................................................................186 Telaranea pecten (Spruce) J.J.Engel et G.L.Merr. ........................................................................187 Telaranea pellucida J.J.Engel et G.L.Merr. ..................................................................................186 Telaranea pseudozoopsis (Herzog) Fulford ....................................................................................186 Telaranea redacta (Steph.) J.J.Engel et G.L.Merr. .......................................................................186 Telaranea rosarioana H.A.Mill. ...................................................................................................186 Telaranea sejuncta (Ångstr.) S.W.Arnell ......................................................................................187 Telaranea setosa J.J.Engel et G.L.Merr.........................................................................................186 Telaranea tenuifolia J.J.Engel et G.L.Merr. .................................................................................187 Telaranea trisetosa (Steph.) Grolle ...............................................................................................186 Temnoma angustifolium R.M.Schust. ..........................................................................................255 Temnoma chaetophyllum R.M.Schust. .........................................................................................255 Temnoma palmatum (Lindb. ex Pearson) R.M.Schust. ................................................................255 Temnoma palmatum var. cuneatum R.M.Schust. .........................................................................255 Temnoma palmatum var. laxifolium R.M.Schust. ........................................................................255 Temnoma palmatum var. pseudospiniferum R.M.Schust. ..............................................................255 Temnoma patagonicum R.M.Schust. ...........................................................................................255 Temnoma paucisetigerum R.M.Schust..........................................................................................255 Temnoma pilosum (A.Evans) R.M.Schust. ...................................................................................255 Temnoma pulchellum (Hook.) Mitt. ............................................................................................255 Temnoma quadriidum (Mitt.) Mitt. ex E.A.Hodgs. et Allison ....................................................255 Temnoma quadripartitum (Hook.) Mitt. .....................................................................................255 Temnoma quadripartitum var. pseudopungens R.M.Schust. ..........................................................255 Temnoma quadripartitum var. randii (S.W.Arnell) R.M.Schust. .................................................256 Temnoma setigerum (Lindenb.) R.M.Schust. ...............................................................................256 Temnoma setigerum var. hawaiicum Inoue ...................................................................................256 Temnoma townrowii R.M.Schust. ...............................................................................................256 Tetracymbaliella comptonii (Pearson) Grolle ................................................................................137 Tetracymbaliella cymbalifera (Hook.f. et Taylor) Grolle ..............................................................137 Tetracymbaliella decipiens (Gottsche) Grolle ................................................................................138 Tetracymbaliella subsimplex (Austin) J.J.Engel .............................................................................138 Tetralophozia cavallii (Gola) Váňa ................................................................................................55 Tetralophozia iliformis (Steph.) Urmi ...........................................................................................55 Tetralophozia pilifera (Steph.) R.M.Schust. ...................................................................................55 Tetralophozia setiformis (Ehrh.) Schljakov .....................................................................................55 hysananthus aculeatus Herzog ....................................................................................................413 hysananthus amazonicus (Spruce) Schifn...................................................................................413 hysananthus anguiformis (Hook.f. et Taylor) Gottsche, Lindenb. et Nees ..................................413 hysananthus appendiculatus Steph. .............................................................................................413 hysananthus ciliaris (Sande Lac.) Sukkharak ..............................................................................413 hysananthus comosus Lindenb. ...................................................................................................413 hysananthus convolutus Lindenb. ...............................................................................................413 hysananthus convolutus var. laceratus (Steph.) Sukkharak ...........................................................413 hysananthus discretus Sukkharak et Gradst. ................................................................................413 hysananthus fruticosus (Lindenb. et Gottsche) Schifn. ...............................................................414 hysananthus gottschei (J.B.Jack et Steph.) Steph. ........................................................................413 World checklist of hornworts and liverworts ** *** *** *** *** ** *** * *** * * * * * *** ** ** ** ** *** *** *** *** *** *** ** ** *** ** ** ** *** ** *** * ** ** *** *** *** *** ** *** *** *** *** * *** ** ** ** *** 665 hysananthus gottschei var. continuus Sukkharak ..........................................................................413 hysananthus mollis Steph. ..........................................................................................................414 hysananthus montanus Gradst., Xiao L.He et Piippo..................................................................414 hysananthus pancheri (Steph.) Hürl. ..........................................................................................413 hysananthus retusus (Reinw., Blume et Nees) B.M.hiers et Gradst. ..........................................414 hysananthus retusus subsp. sellingii (Herzog) Sukkharak.............................................................414 hysananthus spathulistipus (Reinw., Blume et Nees) Lindenb. ....................................................414 hysanolejeunea africana Sim .......................................................................................................512 Trabacellula tumidula Fulford .......................................................................................................64 Trachylejeunea conifera Steph. .....................................................................................................512 Trachylejeunea cristulilora Steph. ................................................................................................512 Trachylejeunea englishii Steph......................................................................................................512 Trachylejeunea jamaicensis Pearson ..............................................................................................512 Trachylejeunea kusaiensis Inoue et H.A.Mill. ...............................................................................512 Treubia insignis K.I.Goebel ...........................................................................................................43 Treubia insignis subsp. bracteata (Steph.) R.M.Schust. et G.A.M.Scott .........................................43 Treubia insignis subsp. caledonica R.M.Schust. et G.A.M.Scott .....................................................43 Treubia insignis subsp. vitiensis R.M.Schust. et G.A.M.Scott ........................................................43 Treubia lacunosoides T.Pfeif., W.Frey et M.Stech .........................................................................43 Treubia lacunosa (Colenso) Prosk..................................................................................................43 Treubia pygmaea R.M.Schust. .......................................................................................................43 Treubia scapanioides R.M.Schust...................................................................................................43 Treubia tahitensis (Nadeaud) Besch. ..............................................................................................43 Treubia tasmanica R.M.Schust. et G.A.M.Scott ............................................................................43 Triandrophyllum eophyllum (R.M.Schust.) Gradst. ......................................................................141 Triandrophyllum fernandeziense (S.W.Arnell) Grolle ex Fulford et Hatcher .................................141 Triandrophyllum heterophyllum (Steph.) Grolle ...........................................................................141 Triandrophyllum subtriidum (Hook.f. et Taylor) Fulford et Hatcher ..........................................141 Triandrophyllum subtriidum var. triidum (Gottsche) Solari........................................................141 Triandrophyllum symmetricum J.J.Engel ......................................................................................141 Trichocolea argentea Herzog ........................................................................................................257 Trichocolea breviissa Steph. .........................................................................................................257 Trichocolea comptonii Pearson .....................................................................................................257 Trichocolea ilicaulis Steph. ..........................................................................................................257 Trichocolea loccosa Herzog et Hatcher ........................................................................................257 Trichocolea geniculata Pearson .....................................................................................................257 Trichocolea gracillima Austin .......................................................................................................257 Trichocolea hatcheri E.A.Hodgs. ..................................................................................................257 Trichocolea iriomotensis T.Katag. .................................................................................................257 Trichocolea japonica T.Katag. ......................................................................................................257 Trichocolea magna T.Katag. ........................................................................................................257 Trichocolea minutifolia Steph. .....................................................................................................257 Trichocolea mollissima (Hook.f. et Taylor) Gottsche ....................................................................257 Trichocolea pluma (Reinw., Blume et Nees) Mont.......................................................................258 Trichocolea rigida R.M.Schust. ....................................................................................................258 Trichocolea rudimentaris Steph. ...................................................................................................258 Trichocolea sprucei Steph. ............................................................................................................258 Trichocolea tomentella (Ehrh.) Dumort. ......................................................................................258 Trichocolea udarii D.K.Singh ......................................................................................................258 Trichocolea wattsiana Steph. ........................................................................................................258 Trichocoleopsis sacculata (Mitt.) S.Okamura ................................................................................434 Tricholepidozia chaetocarpa (Pearson) E.D.Cooper ......................................................................179 666 *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** *** ** *** ** *** ** ** *** * * *** ** ** ** ** *** ** ** ** *** *** ** *** *** *** *** Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Tricholepidozia fernandeziensis (Steph.) E.D.Cooper ...................................................................179 Tricholepidozia ferruginea (J.J.Engel et G.L.Merr.) E.D.Cooper ..................................................179 Tricholepidozia issifolia (Steph.) E.D.Cooper..............................................................................179 Tricholepidozia jowettiana (H.A.Mill.) E.D.Cooper ....................................................................179 Tricholepidozia kaindina (Grolle) E.D.Cooper ............................................................................179 Tricholepidozia kogiana (Steph.) E.D.Cooper ..............................................................................179 Tricholepidozia lawesii (Steph.) E.D.Cooper ...............................................................................179 Tricholepidozia leratii (Steph.) E.D.Cooper ................................................................................180 Tricholepidozia lindenbergii (Gottsche) E.D.Cooper ...................................................................180 Tricholepidozia lindenbergii var. complanata (J.J.Engel et G.L.Merr.) E.D.Cooper ......................180 Tricholepidozia lindenbergii var. mellea (J.J.Engel et G.L.Merr.) E.D.Cooper ..............................180 Tricholepidozia lindenbergii var. papillata (J.J.Engel et G.L.Merr.) E.D.Cooper ..........................180 Tricholepidozia marginata (J.J.Engel et G.L.Merr.) E.D.Cooper .................................................180 Tricholepidozia martinii (E.A.Hodgs.) E.D.Cooper.....................................................................180 Tricholepidozia melanesica (H.A.Mill.) E.D.Cooper ....................................................................180 Tricholepidozia neesii (Lindenb.) E.D.Cooper .............................................................................180 Tricholepidozia octoloba (Del Ros.) E.D.Cooper ..........................................................................180 Tricholepidozia plumulosa (Lehm. et Lindenb.) E.D.Cooper .......................................................180 Tricholepidozia pulcherrima (Steph.) E.D.Cooper .......................................................................180 Tricholepidozia pulcherrima var. mooreana (Steph.) E.D.Cooper .................................................180 Tricholepidozia quadriseta (Steph.) E.D.Cooper ..........................................................................181 Tricholepidozia remotifolia (E.A.Hodgs.) E.D.Cooper .................................................................181 Tricholepidozia semperiana (Steph.) E.D.Cooper .........................................................................181 Tricholepidozia tetradactyla (Hook.f. et Taylor) E.D.Cooper .......................................................181 Tricholepidozia trichocoleoides (Herzog) E.D.Cooper ...................................................................181 Trichotemnoma corrugatum (Steph.) R.M.Schust. .......................................................................136 Trilophozia quinquedentata (Huds.) Bakalin .................................................................................81 Trilophozia quinquedentata var. asymmetrica (Horik.) L.Söderstr. et Váňa.....................................82 Tritomaria exsecta (Schmidel) Schifn. ex Loeske ...........................................................................82 Tritomaria exsecta subsp. novaezelandiae J.J.Engel .........................................................................82 Tritomaria exsectiformis (Breidl.) Schifn. ex Loeske ......................................................................82 Tritomaria exsectiformis subsp. arctica R.M.Schust. .......................................................................82 Tritomaria exsectiformis subsp. camerunensis S.W.Arnell ex Váňa ..................................................82 Tritomaria ferruginea (Grolle) Váňa ..............................................................................................82 Tritomaria koreana Bakalin, S.S.Choi et B.Y.Sun .........................................................................82 Tritomaria mexicana Bakalin.........................................................................................................82 Tritomaria scitula (Taylor) Jørg.....................................................................................................82 Tuyamaella angulistipa (Steph.) R.M.Schust. et Kachroo ............................................................350 Tuyamaella borneensis Tixier .......................................................................................................350 Tuyamaella hattorii Tixier ...........................................................................................................350 Tuyamaella jackii (Steph.) Tixier ................................................................................................350 Tuyamaella molischii (Schifn.) S.Hatt. .......................................................................................350 Tuyamaella molischii var. brevistipa P.C.Wu et P.J.Lin ................................................................350 Tuyamaella molischii var. taiwanensis R.L.Zhu et M.L.So............................................................350 Tuyamaella serratistipa S.Hatt. ....................................................................................................350 Tuzibeanthus chinensis (Steph.) Mizut. ........................................................................................414 Vanaea plagiochiloides (Inoue et Gradst.) Inoue et Gradst. ............................................................49 Vandiemenia ratkowskiana Hewson.............................................................................................461 Verdoornia succulenta R.M.Schust. ..............................................................................................456 Verdoornianthus griinii Gradst...................................................................................................414 Verdoornianthus marsupiifolius (Spruce) Gradst. ..........................................................................414 Vetaforma dusenii (Steph.) Fulford et J.Taylor ............................................................................142 World checklist of hornworts and liverworts *** ** *** *** *** *** *** * ** ** *** *** *** *** *** *** *** *** *** ** ** *** ** *** ** *** *** ** *** *** * *** ** *** * *** *** ** *** ** 667 Vitalianthus bischlerianus (K.C.Pôrto et Grolle) R.M.Schust. et Giancotti ..................................361 Vitalianthus guangxianus R.L.Zhu, Qiong He et Y.M.Wei..........................................................361 Wettsteinia densiretis (Herzog) Grolle ............................................................................................45 Wettsteinia inversa (Sande Lac.) Schifn.........................................................................................45 Wettsteinia rotundifolia (Horik.) Grolle .........................................................................................45 Wettsteinia schusteriana Grolle.......................................................................................................45 Wiesnerella denudata (Mitt.) Steph..............................................................................................504 Wiesnerella fasciaria C.Gao et K.C.Chang ...................................................................................504 Xenocephalozia navicularis (Steph.) R.M.Schust. .........................................................................217 Xenochila integrifolia (Mitt.) Inoue..............................................................................................253 Xenothallus vulcanicola R.M.Schust. ............................................................................................473 Xylolejeunea aquarius (Spruce) Xiao L.He et Grolle .....................................................................398 Xylolejeunea crenata (Nees et Mont.) Xiao L.He et Grolle ...........................................................398 Xylolejeunea grolleana (Pócs) Xiao L.He et Grolle ........................................................................398 Xylolejeunea muricella Xiao L.He et Grolle ..................................................................................398 Zantenia borneensis (Herzog) Váňa et J.J.Engel .............................................................................56 Zantenia denticulata (Grolle) Váňa et J.J.Engel .............................................................................56 Zantenia karstenii (Schifn.) Váňa et J.J.Engel ...............................................................................56 Zantenia prionophylla (S.Hatt.) Váňa et J.J.Engel ..........................................................................56 Zoopsidella antillana (Steph.) H.Rob...........................................................................................187 Zoopsidella antillana subsp. jamaicensis R.M.Schust. ...................................................................187 Zoopsidella caledonica (Steph.) R.M.Schust. ................................................................................187 Zoopsidella cynosurandra (Steph.) R.M.Schust. ............................................................................187 Zoopsidella integrifolia (Spruce) R.M.Schust................................................................................188 Zoopsidella macella (Spruce) R.M.Schust. ....................................................................................188 Zoopsidella serra (Spruce) R.M.Schust. ........................................................................................188 Zoopsis argentea (Hook.f. et Taylor) Gottsche, Lindenb. et Nees .................................................188 Zoopsis argentea var. lagelliformis (Colenso) R.M.Schust. ............................................................188 Zoopsis bicruris Glenny et E.A.Br. ...............................................................................................188 Zoopsis ceratophylla (Spruce) Hamlin...........................................................................................188 Zoopsis ciliata Colenso ................................................................................................................189 Zoopsis leitgebiana (Carrington et Pearson) Bastow .....................................................................188 Zoopsis liukiuensis Horik. ............................................................................................................188 Zoopsis macrophylla R.M.Schust. .................................................................................................188 Zoopsis martinicensis Steph. .........................................................................................................189 Zoopsis matawaia M.A.M.Renner ...............................................................................................188 Zoopsis nitida Glenny, Braggins et R.M.Schust. ..........................................................................188 Zoopsis setigera K.I.Goebel ..........................................................................................................189 Zoopsis setulosa Leitg....................................................................................................................188 Zoopsis uleana Steph....................................................................................................................189 Acknowledgements he Early Land Plants Today project has been a community driven and collaborative efort that has involved numerous dedicated individuals, many institutions, and funding agencies that have helped provide partial funding. he high quality of the data and information could not have been achieved without rapid responses to our requests for information regarding publications, PDFs or other forms of bibliographic data from committed individuals. hese include Afroz Alam, Dulip Daniels, Rajendra Lavate, Shuvadeep Majum- 668 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) dar, Alexey Potemkin, Ana Séneca, Mafalda Silva, Devendra K. Singh, Sushil K. Singh, Louis houvenot, Praveen Verma, and Gerhard Winter. We wish to especially thank the many individuals who have provided a wealth of logistical support that has been broad ranging from access to data, assistance with translation, access to unpublished molecular results and information, access to type material or herbaria support, library support, advice on Latin usage as well as many other forms of support. hese have included Silvia Aranda, Riccardo Baldini, Fred Barrie, Laura Campos, Tong Cao, Xia-Fang Cheng, Patricia Eckel, Len Ellis, Lars Hedenäs, Christine Giannoni, Qiong He, Nijole Kalinauskaitė, Lucia Kawasaki, Peter de Lange, Itambi Malombe, John McNeill, David Meagher, Chiara Nepi, Michelle Price, Qiong Qui, Steven Rogstad, Cecilia Sérgio, Blanka Shaw, Jon Shaw, Akiko Shinya, Lei Shu, Sorin Ştefănuţ, Alain Vanderpoorten, Anna Vilnet, Yu-Mei Wei, Ying Yu, and Lan-Ping Zhou. We gratefully acknowledge the Biodiversity Heritage Library as a resource and the Conservatoire et Jardin botaniques de la Ville de Genève for providing access to the Franz Stephani database and specimens and Missouri Botanical Garden for access to the TROPICOS database. hroughout the years, the project has been generously supported in part by the Global Biological Information Facility (GBIF) Seed Money Award No.2007-41 and supported in part by funding from the Biodiversity Synthesis Center of the Encyclopedia of Life. Funding from the National Science Foundation (DBI0749762, DBI 1057418, DBI 1458300, DEB- 1145898, EF-0531730, EF-1115116) has also partially supported aspects of the ELPT project. We also wish to express our gratitude to the editors and publication team of Phytotaxa that published the important series of notes that helped lead to the current publication and to the tireless eforts of the editorial and production team of PhytoKeys that led to the current publication. References Abeywickrama BA (1959) he genera of liverworts of Ceylon. Ceylon Journal of Science. Biological Sciences 2 (1): 33–81. Acharius E (1805) Jungermannia violacea, eine neue Art aus Dusky Bay. Beiträge zur Naturkunde (Weber & Mohr) 1: 76–78. doi: 10.5962/bhl.title.45015 Adanson M (1763) Familles des plantes, II partie. Vincent, Paris, 640 pp. doi: 10.5962/bhl. title.271 Ahmad S (1942) hree new species of Riccia from India. Current Science 11 (11): 433–434. Ah-Peng C, Bardat J (2011) Microlejeunea strasbergii sp. nov. (Lejeuneaceae) from Réunion Island (Mascarenes). Bryologist 114 (4): 668–673. doi: 10.1639/0007-2745-114.4.668 Alam A, Srivastava SC (2009) Current status of genus Plagiochasma in Nilgiri and Palni Hills with SEM details of spores. Indian Journal of Forestry 32 (4): 623–634. Alam A, Kumar A, Srivastava SC (2007) Jungermannia nilgiriensis, a new species from Nilgiri Hills (Western Ghats) India. Bulletin of the Botanical Survey of India 49 (1/4): 219–224. Alam A, Sharma D, Yadav S (2012) Solenostoma tetragonum (Lindenb.) R. M. Schust. ex Váňa et D. G.Long var. kodaikanalensis var. nov. Phytotaxonomy 12: 68–71. Allorge P (1929) Schedae ad Bryothecam Ibericam, 2e série, Nos. 51-100, Espagne. Paris, 27 pp. World checklist of hornworts and liverworts 669 Allorge P, Allorge V (1950) Hépatiques récoltées par P. et V. Allorge aux îles Açores en 1937. Revue Bryologique et Lichénologique 19 (1/2): 90–118. Allorge V, Jovet-Ast S (1955) Cololejeunea azorica V. All. et Jovet-Ast, Lejeunéacée nouvelle de l’Île San Miguel. Mitteilungen der hüringischen Botanischen Gesellschaft 1 (2/3): 17–22. Amakawa T (1954) Notes on Japanese hepaticae (1). Journal of Japanese Botany 29 (6): 177–180. Amakawa T (1956) Notes on Japanese hepaticae (2). Journal of Japanese Botany 31 (2): 47–50. Amakawa T (1957a) Notes on Japanese hepaticae (4). Journal of Japanese Botany 32 (6): 165–169. Amakawa T (1957b) Notes on Japanese hepaticae (6). Journal of Japanese Botany 32 (10): 307–312. Amakawa T (1957c) Notes on Japanese hepaticae (3). Journal of Japanese Botany 32 (2): 34–41. Amakawa T (1957d) Notes on Japanese hepaticae (5). Journal of Japanese Botany 32 (7): 215–219. Amakawa T (1958a) Notes on Japanese hepaticae (7). Journal of Japanese Botany 33 (5): 142–145. Amakawa T (1958b) Notes on Japanese hepaticae (8). Journal of Japanese Botany 33 (11): 338–343. Amakawa T (1959a) Family Jungermanniaceae of Japan. I. Journal of the Hattori Botanical Laboratory 21: 248–291. Amakawa T (1959b) Notes on Japanese hepaticae (9). Journal of Japanese Botany 34 (4): 111–116. Amakawa T (1960a) Notes on Japanese hepaticae (11). Journal of Japanese Botany 35 (12): 363–368. Amakawa T (1960b) Family Jungermanniaceae of Japan, II. Journal of the Hattori Botanical Laboratory 22: 1–90. Amakawa T (1962) A new genus Diplocolea (Hepaticae) from eastern Himalayas. Journal of Japanese Botany 37 (9): 274. Amakawa T (1963) New or little known Asiatic species of the family Jungermanniaceae. I. Journal of the Hattori Botanical Laboratory 26: 20–26. Amakawa T (1964a) A short revision of Himalayan Scapania (Hepaticae). Journal of the Hattori Botanical Laboratory 27: 1–19. Amakawa T (1964b) Notes on Japanese hepaticae (13). Journal of Japanese Botany 39 (5): 135–139. Amakawa T (1965) Notes on Japanese hepaticae (14). Journal of Japanese Botany 40 (10): 307–310. Amakawa T (1966) New or little known Asiatic species of the family Jungermanniaceae. II. Journal of the Hattori Botanical Laboratory 29: 253–266. Amakawa T (1967a) Scapania sect. Gracilidae of the areas around the North Paciic Ocean. Journal of the Hattori Botanical Laboratory 30: 315–327. Amakawa T (1967b) New or little known Asiatic species of the family Jungermanniaceae. III. Journal of the Hattori Botanical Laboratory 30: 181–198. Amakawa T (1968a) New or critical species of the genus Scapania of Japan I. Journal of the Hattori Botanical Laboratory 31: 94–100. 670 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Amakawa T (1968b) New or little known Asiatic species of the family Jungermanniaceae. IV. Journal of the Hattori Botanical Laboratory 31: 101–112. Amakawa T (1970) New or little known Asiatic species of the family Jungermanniaceae. VI. Jungermannia comata Nees and its allies. Journal of the Hattori Botanical Laboratory 33: 153–160. Amakawa T, Hattori S (1953) A revision of the Japanese species of Scapaniaceae. I. Journal of the Hattori Botanical Laboratory 9: 45–62. Amakawa T, Hattori S (1954) A revision of the Japanese species of Scapaniaceae. II. Journal of the Hattori Botanical Laboratory 12: 91–112. Amakawa T, Hattori S (1955) A revision of the Japanese species of Scapaniaceae. III. Journal of the Hattori Botanical Laboratory 14: 71–90. Ando H (1963) A Pseudolepicolea found in the middle Honshu of Japan. Hikobia 3 (3): 177–183. Ångström J (1872) Förteckning och beskrifning öfver mossor, samlade af Professor N.J. Anderson under Fregatten Eugenies verldsomsegling åren 1851-53. I-III. Öfversigt af Kongl. Vetenskaps-Akademiens Förhandlingar 29 (4): 3–29. Ångström J (1873) Mossor från Tahiti och Eimeo. Öfversigt af Kongl. Vetenskaps-Akademiens Förhandlingar 30 (5): 113–151. Ångström J (1876) Primae lineae muscorum cognoscendorum, qui ad Caldas Brasiliae sunt collecti. Continnuatio. II. Hepaticae. Öfversigt af Kongl. Vetenskaps-Akademiens Förhandlingar 33 (7): 77–92. Anonymous (1848) Proceedings of learned societies. Linnean Society. Annals and Magazine of Natural History (ser. 2) 1 (5): 375–387. Anonymous (1873) Smärre notiser. Lärda sällskaps sammanträden. Botaniska Notiser 26: 59–62. Aranda SC, Gradstein SR, Patiño J, Laenen B, Désamoré A, Vanderpoorten A (2014) Phylogeny, classiication and species delimitation in the liverwort genus Odontoschisma (Cephaloziaceae). Taxon 63 (5): 1008–1025. doi: 10.12705/635.12 Ariyanti NS, Gradstein SR, Sporn SG, Angelika R, Tan BC (2009) Catalogue of the bryophytes of Sulawesi. Supplement 1: new species records. Blumea 54 (1/3): 287–289. doi: 10.3767/000651909X476300 Arnell HW (1892) Lebermoosstudien im nördlichen Norwegen. C. J. Lundgrens Enkas Tryckeri, Jönköping, 44 pp. Arnell HW (1898) Musci novi. Revue Bryologique 25 (1): 1–9. Arnell HW (1902) Novae species generis Kantiae. Revue Bryologique 29 (2): 26–32. Arnell HW (1903) Martinellia calcicola Arnell et Persson nova sp. Revue Bryologique 30 (6): 97–98. Arnell HW (1913) Zur Mooslora des Lena-Tales. Arkiv för Botanik 13 (2): 1–94. Arnell HW (1921) Martinellia tundrae Arnell, nova species. Botaniska Notiser 74: 289–291. Arnell HW (1927) Lebermoose aus Kamtschatka. Hedwigia 67 (1/2): 110–112. Arnell HW (1928) Levermossor (in: Hartmans Handbok i Skandinaviens Flora redigerad av Otto R. Holmberg). P. A. Norstedt & Söner Förlag, Stockholm, 224 pp. Arnell SW (1947) Contribution to the knowledge of the hepatics of Novaya Zemlya. Svensk Botanisk Tidskrift 41: 209–217. Arnell SW (1950) Leiocolea arctica, nov. spec. Svensk Botanisk Tidskrift 44 (2): 374–378. Arnell SW (1952a) Hepaticae collected in South and West Africa 1951. Botaniska Notiser 105: 307–329. World checklist of hornworts and liverworts 671 Arnell SW (1952b) South African species of Riccardia. Botaniska Notiser 105: 138–156. Arnell SW (1953a) Hepaticae collected in South Africa 1951. Svensk Botanisk Tidskrift 47 (1): 107–118. Arnell SW (1953b) Hepaticae collected in South Africa 1951. New and little known species. III. Botaniska Notiser 106: 271–289. Arnell SW (1953c) List of hepaticae collected in Marion Island by Mr. R. W. Rand Dec. 1951-April 1952. Svensk Botanisk Tidskrift 47 (3): 411–425. Arnell SW (1954a) New species of hepaticae from South Africa. Revue Bryologique et Lichénologique 23 (1/2): 173–179. Arnell SW (1954b) Species of Riccardia collected by H. Roivainen in Tierra del Fuego and west Patagonia, 1928-29. Archivum Societatis Zoologicae Botanicae Fennicae “Vanamo” 9: 48–55. Arnell SW (1954c) A new species of Lepidozia from South Africa. Botaniska Notiser 107: 427–429. Arnell SW (1955a) A new species of Cololejeunea from Moçambique. Mitteilungen der hüringischen Botanischen Gesellschaft 1 (1): 7–8. Arnell SW (1955b) Notes on South African hepatics. II. Botaniska Notiser 108: 309–313. Arnell SW (1956a) A new species of Cheilolejeunea from Samoa. Svensk Botanisk Tidskrift 50 (3): 516–517. Arnell SW (1956b) Illustrated moss lora of Fennoscandia. I. Hepaticae. Nordic Bryological Society, Lund, 314 pp. Arnell SW (1956c) Hepaticae collected in Cuba and Hispaniola by E. L. Ekman. Bryologist 59 (4): 271–276. doi: 10.2307/3240336 Arnell SW (1956d) Hepaticae collected by Dr. and Mrs. Carl Skottsberg on Cerro Talinay, Prov. Coquimbo, Chile, 1955. Svensk Botanisk Tidskrift 50 (2): 308–312. Arnell SW (1956e) Hepaticae collected by O. Hedberg et al. on the East African Mountains. Arkiv för Botanik (n.ser.) 3 (16): 517–562. Arnell SW (1957a) Notes on South African hepaticae. IV. Botaniska Notiser 110 (1): 17–27. Arnell SW (1957b) Hepaticae collected during Dr. and Mrs. Skottsberg’s second expedition to the Juan Fernández Islands, Dec. 1954-March 1955. Arkiv för Botanik (n.ser.) 4 (1): 1–21. Arnell SW (1957c) Hepaticae collected in southwest Africa by Dr. O. H. Volk. Mitteilungen der Botanischen Staatssammlung München 2 (16): 262–272. Arnell SW (1957d) Notes on South African hepaticae. V. Botaniska Notiser 110: 399–405. Arnell SW (1958a) New hepaticae from Cameroon Mountain. Svensk Botanisk Tidskrift 52 (1): 63–67. Arnell SW (1958b) Hepatics from Tristan da Cunha. Results of the Norwegian Scientiic Expedition to Tristan da Cunha, 1937-38 42: 1–76. Arnell SW (1958c) New species of Asterella and Riccia from Sto. Domingo and Cuba. Bryologist 61 (2): 140–143. doi: 10.2307/3240214 Arnell SW (1959) Hepatics collected by Dr. G. Degelius on Martinique and in French Guiana in 1958. Svensk Botanisk Tidskrift 53 (4): 499–506. Arnell SW (1961) Some new American species of hepatics. Svensk Botanisk Tidskrift 55 (1): 205–210. 672 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Arnell SW (1962a) Notes on South African hepaticae. VII. Botaniska Notiser 115: 203–207. Arnell SW (1962b) Contribution to the knowledge of the hepaticae of Ecuador. Svensk Botanisk Tidskrift 56 (2): 334–350. Arnell SW (1963a) Two new hepatics from Queensland. Svensk Botanisk Tidskrift 57 (2): 190–192. Arnell SW (1963b) Hepaticae of South Africa. Swedish Natural Science Research Council, Stockholm, 411 pp. Arnell SW (1965) Hepaticae collected by Mr. Gillis Een in Mauritius and Réunion in 1962. Svensk Botanisk Tidskrift 59 (1): 65–84. Arnell HW, Buch H (1921) Martinellia scandica nov. spec. Botaniska Notiser 74: 1–2. Arnell HW, Jensen C (1907) Die Moose des Sarekgebietes. Erste Abteilung. In: Hamberg A (Ed.) Naturwissenschaftliche Untersuchungen des Sarekgebirges in Schwedisch-Lappland Bd. 3. Botanik. C. E. Fritzes, Stockholm, 71–132. Arnell SW, Mårtensson O (1959) A contribution to the knowledge of the bryophyte lora of W. Spitsbergen, and Kongsjorden (King’s Bay, 79°N.) in particular. Arkiv för Botanik (n.ser.) 4 (6): 105–164. Asakawa Y (1998) Biologically active compounds from bryophytes. Journal of the Hattori Botanical Laboratory 84: 91–104. Asthana G, Maurya M (2014) Lethocolea indica: A new liverwort from India. National Academy Science Letters 37 (6): 535–539. doi: 10.1007/s40009-014-0274-z Asthana AK, Nath V (1993) A new Phaeoceros from western Himalayas, Phaeoceros udarii sp. nov. Proceedings of the National Academy of Sciences of India. Section B, Biological Sciences 63 (4): 461–464. Asthana G, Shukla A (2010) A new epiphyllous species of Cololejeunea (Lejeuneaceae) from India. Cryptogamie, Bryologie 31 (3): 217–221. Asthana AK, Srivastava SC (1986) A new Folioceros from Silent Valley. Cryptogamie: Bryologie, Lichénologie 7 (2): 149–153. Asthana AK, Srivastava SC (1991) Indian hornworts (a taxonomic study). Bryophytorum Bibliotheca 42: 1–159. Asthana G, Srivastava SC (2003) Indian Cololejeunea: A taxonomic study. Bryophytorum Bibliotheca 60: 1–155. Asthana G, Srivastava S (2013) Cephalozia kodaikanalensis sp. nov. (Cephaloziaceae) from Palni Hills, Tamil Nadu, India. Geophytology 43 (1): 63–67. Asthana G, Srivastava SC, Asthana AK (1995) he genus Cheilolejeunea in India. Lindbergia 20 (2/3): 125–143. Asthana G, Saxena M, Maurya M (2013) A new species of Blepharostoma, B. indica sp. nov. from the western Himalaya, India, with observations on the closely allied B. trichophyllum (L.) Dumort. Journal of Bryology 35 (4): 266–269. doi: 10.1179/0373668713Z.00000000076 Augier J (1972) Cheilolejeunea diversifolia sp. nov. Annales, Faculté des Sciences, Université Fédérale du Cameroun 11: 65–70. Austin CF (1869) Characters of some new hepaticae. Proceedings of the Academy of Natural Sciences of Philadelphia 21: 218–234. World checklist of hornworts and liverworts 673 Austin CF (1872) New hepaticae. Bulletin of the Torrey Botanical Club 3 (3): 9–18. doi: 10.2307/2476500 Austin CF (1873) Hepaticae boreali-americanae. Closter, New Jersey, 1–48. Austin CF (1874) Sandwich Island hepaticae. Bulletin of the Torrey Botanical Club 5 (3): 14–18. doi: 10.2307/2475623 Austin CF (1875a) Notes on the Anthocerotaceae of North America, with descriptions of several new species. Bulletin of the Torrey Botanical Club 6 (4): 25–29. doi: 10.2307/2476484 Austin CF (1875b) New hepaticae. Bulletin of the Torrey Botanical Club 6 (3): 17–21. doi: 10.2307/2475861 Austin CF (1875c) New hepaticae. Bulletin of the Torrey Botanical Club 6 (7): 46–47. doi: 10.2307/2475383 Austin CF (1876a) Notes on hepaticology. Botanical Bulletin. Hanover 1 (8): 35–36. Austin CF (1876b) Notes on hepaticology. Botanical Bulletin. Hanover 1 (7): 31–32. Austin CF (1876c) Notes and criticism on Hepaticae Boreali-Americanae Exsiccatae. Bulletin of the Torrey Botanical Club 6 (16): 85. doi: 10.2307/2476773 Austin CF (1877) New hepaticae. Bulletin of the Torrey Botanical Club 6 (30): 157–158. doi: 10.2307/2475830 Austin CF (1878) Notes on hepaticology. Botanical Gazette 3 (1): 6–7. doi: 10.1086/325111 Austin CF (1879) Notes on hepaticology. Bulletin of the Torrey Botanical Club 6 (52): 301– 306. doi: 10.2307/2476817 Bai X-L, Gao C (1999) A new species of the genus Frullania taxa from Yunnan, China. Hikobia 13 (1): 87–88. Bai X-L, Gao C (2000) Frullania laevi-periantha (Frullaniaceae, Hepaticae). A new species from Yunnan, China. Nova Hedwigia 70 (1/2): 135–137. Bakalin VA (2003) Notes on Lophozia. IV. Some new taxa of Lophozia sensu stricto. Annales Botanici Fennici 40 (1): 47–52. Bakalin VA (2005) Monograicheskaia obrabotka roda Lophozia (Dumort.) Dumort. s. str. [Monograph of the genus Lophozia (Dumort.) Dumort. s. str.]. Nauka, Moscow, 240 pp. Bakalin VA (2007) New liverwort records from Sakhalin Province. 2. Southern Kuril Islands. Arctoa 16: 202–209. doi: 10.15298/arctoa.16.03 Bakalin VA (2008a) New liverwort records from Sakhalin Province. 3 [Novye nahodki pechenochnikov v Sahalinskoj oblasti]. Arctoa 17: 226–230. doi: 10.15298/arctoa.17.16 Bakalin VA (2008b) New data on distribution and taxonomy of some species in Lophoziaceae (Hepaticae) [Novye svedeniia o rasprostranenii i taksonomii nekotoryh vidov semejstva Lophoziaceae (Hepaticae)]. Arctoa 17: 161–164. doi: 10.15298/arctoa.17.12 Bakalin VA (2008c) On taxonomy of some hepatics from Primorsky Territory (Russian Far East), with the list of taxa of the territory [O taksonomii nekotoryh pechenonikov Primorskogo kraia (Rossijskij Dal’nij Vostok) so spiskom taksonov, izvestnyh v krae]. Arctoa 17: 101–108. doi: 10.15298/arctoa.17.08 Bakalin VA (2011) Notes on Lophozia VI. Taxonomy and distribution of Lophozia and Schistochilopsis (Lophoziaceae) in North America north of Mexico. Bryologist 114 (2): 298– 315. doi: 10.1639/0007-2745.114.2.298 674 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Bakalin VA (2013) New taxa of Solenostoma and Plectocolea and other taxonomic novelties based on study of collections in the New York Botanical Garden herbarium. Polish Botanical Journal 58 (1): 127–142. doi: 10.2478/pbj-2013-0014 Bakalin VA (2014) he study of type collection in Conservatoire et Jardin Botanique de la Ville de Genève (G): the hepatic genera Jungermannia, Solenostoma and Plectocolea. Arctoa 23: 91–136. doi: 10.15298/arctoa.23.10 Bakalin VA, Vilnet AA (2009) Two new species of Jungermanniaceae from Asiatic Russia [Dva novyh vida Jungermanniaceae iz aziatskoj chasti Rossii]. Arctoa 18: 151–162. doi: 10.15298/arctoa.18.08 Bakalin VA, Vilnet AA (2014) Two new species of the liverwort genus Hygrobiella Spruce (Marchantiophyta) described from the North Paciic based on integrative taxonomy. Plant Systematics and Evolution 300 (10): 2277–2291. doi: 10.1007/s00606-014-1050-8 Bakalin VA, Choi SS, Sun B-Y (2009a) A new species of Tritomaria (Lophoziaceae) from the Korean peninsula [Novyj vid Tritomaria (Lophoziaceae) s Korejskogo Poluostrova]. Arctoa 18: 163–166 . doi: 10.15298/arctoa.18.09 Bakalin VA, Čerdanceva VY, Ignatov MS, Ignatova EA, Nûshko TI (2009b) Bryophyte lora of the South Kuril Islands (East Asia) [Flora mohoobraznykh iuzhnykh Kurilskikh ostrovov (Vostochnaia Asiia)]. Arctoa 18: 69–114. doi: 10.15298/arctoa.18.03 Bakalin VA, Vilnet AA, Furuki T, Katagiri T (2014) Taxonomic novelties in Solenostoma– Plectocolea complex (Solenostomataceae, Hepaticae) in East Asia. Botanica Paciica 3 (2): 3–18. doi: 10.17581/bp.2014.03201 Balbis GB (1804) Sur trois nouvelles espèces d’hépatiques à ajouter à la lore du Piémont. Mémoires de l’Académie des Sciences, Littérature et Beaux-arts de Turin, Sciences Physiques et Mathématiques 15: 73–77. Balmford A, Crane P, Dobson A, Green RE, Mace GM (2005) he 2010 challenge: data availability, information needs and extraterrestrial insights. Philosophical Transactions of the Royal Society of London. Series B 360 (1454): 221–228. doi: 10.1098/rstb.2004.1599 Banwell AD (1951) A new species of Riella from Australia. Transactions of the British Bryological Society 1 (5): 475–478. doi: 10.1179/006813851804878463 Bapna KR (1961) A new species of Riccia from Jodhpur (India). Botaniska Notiser 114 (2): 181–184. Bapna KR (1962) A new species of Riccia from Mount Abu (India). Transactions of the British Bryological Society 4 (2): 249–253. doi: 10.1179/006813862804812599 Bapna KR (1966) A review of Indian Anthocerotales. University of Udaipur Research Studies 3: 127–145. Bapna KR, Vyas GG (1962) Studies in the liverworts of Mount Abu (India). I. A preliminary account. Journal of the Hattori Botanical Laboratory 25: 81–90. Bartholomew-Began SE (1991) A morphogenetic re-evaluation of Haplomitrium Nees (Hepaticopsida). Bryophytorum Bibliotheca 41: 1–297. Bastos CJ (2011) Cheilolejeunea ornata (Lejeuneaceae), a new species from Brazilian Atlantic Forest. Journal of Bryology 33 (1): 86–92. doi: 10.1179/1743282010Y.0000000005 Bastos CJ (2012a) Nomenclatural notes on the genus Rectolejeunea A.Evans (Lejeuneaceae, Marchantiophyta). Journal of Bryology 34 (2): 144–145. doi: 10.1179/1743282011Y.0000000032 World checklist of hornworts and liverworts 675 Bastos CJ (2012b) Type studies on Cheilolejeunea (Spruce) Schifn. (Lejeuneaceae): Brazilian species described by Stephani. Journal of Bryology 34 (4): 315–318. doi: 10.1179/1743282012Y.0000000028 Bastos CJ (2014) On Trachylejeunea subplana Steph. (Lejeuneaceae, Marchantiophyta). Journal of Bryology 36 (3): 249–250. doi: 10.1179/1743282014Y.0000000093 Bastos CJ, Gradstein SR (2006) Two new species of Cheilolejeunea (Spruce) Schifn. (Lejeuneaceae) from Brazil: C. lacerata sp. nov. and C. rupestris sp. nov. Journal of Bryology 28 (2): 133–138. doi: 10.1179/174328206X105425 Bastos CJ, Yano O (2002) Pycnolejeunea porrectilobula (Lejeuneaceae), a new species from Brazil. Nova Hedwigia 74 (3/4): 439–444. doi: 10.1127/0029-5035/2002/0074-0439 Bastow RA (1888) Tasmanian hepaticae. Papers and Proceedings of the Royal Society of Tasmania 1887: 209–289. Battandier JA, Trabut LC (1886) Atlas de la lore d’Alger, fasc. 1. Adolph Jordan, Alger, 1–16. Beckmann KG, Scott GAM (1992) A new thallose genus of leafy liverworts from Australia. Journal of Bryology 17 (2): 297–305. doi: 10.1179/jbr.1992.17.2.297 Bell D, Long DG, Forrest AD, Hollingsworth ML, Blom HH, Hollingsworth PM (2012) DNA barcoding of European Herbertus (Marchantiopsida, Herbertaceae) and the discovery and description of a new species. Molecular Ecology Resources 12: 36–47. doi: 10.1111/j.1755-0998.2011.03053.x Bell D, Long D, Hollingsworth P (2013) he use of DNA barcoding to address major taxonomic problems for rare British bryophytes. Royal Botanic Garden, Edinburgh, 69 pp. Bellardi CAL (1792) Appendix ad loram pedemontanum. Augustae Taurinorum, Torino, 80 pp. Benedix EH (1953) Indomalayische Cololejeuneen. Feddes Repertorium Specierum Novarum Regni Vegetabilis. Beiheft 134: 1–88. Berggren S (1875) Musci et hepaticae spetsbergenses. Kongliga Svenska Vetenskaps-Akademiens Handlingar (n.ser.) 13 (7): 1–103. Berggren S (1898) On New Zealand hepaticae. I. E. Malmström, Lund, 48 pp. Berkeley MJ (1857) Introduction to cryptogamic botany. H. Bailliere, London, 604 pp. doi: 10.5962/bhl.title.45843 Bernal R, Gradstein SR, Celis M (2015) New names and new combinations for the catalogue of the plants and lichens of Colombia. Phytoneuron 2015 (22): 1–6. Bernecker-Lücking A (1998) New species of Lejeuneaceae (Hepaticae) from Costa Rica – I. Aphanolejeunea. Nova Hedwigia 66 (1/2): 163–171. Bernecker-Lücking A (1999) New species of Lejeuneaceae (Hepaticae) from Costa Rica. II. Oryzolejeunea. Haussknechtia, Beiheft 9: 37–40. Bernet H (1888) Catalogue des hépatiques du sud-ouest de la Suisse. H. Georg, Genève, Bale et Lyon, 135 pp. Berrie GK (1963) Australian liverworts. I. Haplomitrium intermedium sp. nov. (Calobryales). Proceedings of the Linnean Society of New South Wales (ser. 2) 87 (399): 191–195. Bertoloni A (1817) Fine delle osservazioni botaniche. Opuscoli Scientiici 1: 229–243. Bertoloni A (1862) Miscellanea botanica XXIII. Memorie della Reale. Accademia della Scienze dell’Istituto di Bologna (Ser. 2) 1: 215–232. Bescherelle É (1892) Énumération des hépatiques récoltées au Tonkin par M. Balansa et déterminées par M. Stephani. Revue Bryologique 19 (1): 13–15. 676 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Bescherelle É (1898) Énumération des hépatiques connues dans les îles de la Société (principalement à Tahiti) et dans les îles Marquises. Journal de Botanique (Morot) 12: 136–150. Bescherelle É, Massalongo CB (1886) Hepaticae novae americanae-australes. Bulletin Mensuel de la Société Linnéenne de Paris 1 (79): 626–640. Bescherelle É, Massalongo CB (1889) Muscinées. I. Hépatiques. In: Hariot PA, Petit P, Muller d’Argovie J, Bescherelle E, Massalongo C, Franchet A (Eds) Mission scientiique du Cap Horn, Tome V. Botanique. Gauthier-Villars et ils, Paris, 201–252. doi: 10.5962/bhl.title.2480 Bescherelle É, Spruce R (1889) Hépatiques nouvelles des colonies Françaises. Bulletin de la Société Botanique de France (Congrès de Botanique) 36: clxxvii–clxxxix. Bharadwaj DC (1971) On Folioceros, a new genus of Anthocerotales. Geophytology 1 (1): 6–15. Bharadwaj DC (1975) Validation of some new combinations under Folioceros Bharadwaj. Geophytology 5 (2): 227–228. Bharadwaj DC (1978) Studies in Indian Anthocerotaceae V. Morphotaxonomy of some Indian species of Folioceros Bharadwaj. Geophytology 8 (1): 111–119. Bisang I (1991) Biosystematische Studien an Lophozia subg. Schistochilopsis (Hepaticae). Bryophytorum Bibliotheca 43: 1–187. Bischler H (1962a) he genus Calypogeja Raddi in Central and South America. II. Subgenus Calypogeja subgroups 1, 2 and 3. Candollea 18: 53–93. Bischler H (1962b) he genus Calypogeja Raddi in Central and South America. I. Introduction and subgenera Mnioloma and Caracoma. Candollea 18: 19–51. Bischler H (1962c) he genus Calypogeja Raddi in Central and South America III. Subgenus Calypogeja, subgroups 4 and 5. Candollea 18: 95–128. Bischler H (1964) Le genre Drepanolejeunea Steph. en Amérique Centrale et Méridionale. Revue Bryologique et Lichénologique 33 (1/2): 15–179. Bischler H (1967) Le genre Drepanolejeunea Steph. en Amérique Centrale et Méridionale. II. Revue Bryologique et Lichénologique 35 (1/4): 95–134. Bischler H (1968) Monographie du genre Rhaphidolejeunea Herzog. Revue Bryologique et Lichénologique 36 (1/2): 56–104. Bischler H (1969) Le genre Leptolejeunea (Spruce) Steph. en Amérique. Nova Hedwigia 17: 265–350. Bischler H (1970) Les espèces du genre Calypogeia sur le continent africain et les îles africaines. Revue Bryologique et Lichénologique 37 (1): 63–134. Bischler H (1977) Plagiochasma Lehm. et Lindenb. I. Le genre et ses subdivisions. Revue Bryologique et Lichénologique 43 (1): 67–109. Bischler H (1978) Plagiochasma Lehm. & Lindenb. II. Les taxa européens et africains. Revue Bryologique et Lichénologique 44 (3): 223–300. Bischler H (1979a) Plagiochasma Lehm. et Lindenb. IV. Les taxa américains. Revue Bryologique et Lichénologique 45 (3): 255–333. Bischler H (1979b) Plagiochasma Lehm. et Lindenb. III. Les taxa d’Asie et d’Océanie. Journal of the Hattori Botanical Laboratory 45: 25–79. World checklist of hornworts and liverworts 677 Bischler H (1982) Marchantia L.: morphologie sporale, germination et rang taxonomique des sections Marchantia et Chlamidium (Corda) Nees. Cryptogamie: Bryologie, Lichénologie 3 (4): 351–364. Bischler H (1984) Marchantia L. he New World species. Bryophytorum Bibliotheca 26: 1–228. Bischler H (1989a) Marchantia L.: subg. Chlamidium (Nees) Bischl. sect. Papillatae Bischl. sect. nov. en Asie et en Océanie. Cryptogamie: Bryologie, Lichénologie 10 (1): 61–79. Bischler H (1989b) Marchantia L. he Asiatic and Oceanic taxa. Bryophytorum Bibliotheca 38: 1–317. Bischler H, Whittemore AT (2001) Cronisia and its two species, C. imbriata and C. weddellii. Cryptogamie, Bryologie 22 (3): 167–174. doi: 10.1016/S1290-0796(01)90001-X Bischler H, Miller HA, Bonner CEB (1961) Studies in Lejeuneaceae III. A historical account of Lejeunea cucullata (Reinwardt, Blume & Nees) Nees and its varieties. Nova Hedwigia 3 (4): 445–462. Bischler H, Bonner CEB, Miller HA (1963) Studies in Lejeuneaceae VI: he genus Microlejeunea Steph. in Central and South America. Nova Hedwigia 5 (1/2): 359–411. Bischler-Causse H (1993) Marchantia L. he European and African taxa. Bryophytorum Bibliotheca 45: 1–129. Bischler-Causse H, Boisselier-Dubayle MC (1991) Lectotypiication of Marchantia polymorpha L. Journal of Bryology 16 (3): 361–365. doi: 10.1179/jbr.1991.16.3.361 Bischler-Causse H, Gradstein SR, Jovet-Ast S, Long DG, Salazar Allen N (2005) Marchantiidae. Flora Neotropica, Monograph 97: 1–267. Bischof GW (1835) Bemerkungen über die Lebermoose. Halle, 1–180. Bizot M, Pócs T (1979) East African bryophytes, III. Acta Botanica Academiae Scientiarum Hungaricae 25 (3/4): 223–261. Blom HH, Holten JI (1988) Plagiochila norvegica, a new hepatic from west Norway. Lindbergia 14 (1): 8–11. Blomquist HL (1939) A new species of Plagiochila from southern Appalachian Mountains. Bryologist 42 (5): 113–117. doi: 10.2307/3239827 Bluf MJ, Fingerhuth CA (1831) Compendium lorae Germaniae. Tomus III. J. L. Schlag, Norimbergae [Nürnberg] 654 pp. Blume CF, Nees CG (1823) Pugillus plantarum iavanicarum e cryptogamicarum variis ordinibus selectus. Nova Acta Physico-Medica Academiae Caesareae Leopoldino-Carolinae Naturae Curiosorum 11: 119–140. Boiko MF (2011) Syntrichia ruralis var. polysporogonica and Riccia rhenana var. violacea – new infraspeciic bryophyte taxa. Chornomorski Botanical Journal 7: 92–93. Boisselier-Dubayle MC, de Chaldée M, Guérin L, Lambourdière J, Bischler H (1995) Genetic variability in western European Lunularia (Hepaticae, Lunulariaceae). Fragmenta Floristica et Geobotanica 40 (1): 379–391. Bonner CEB (1953a) De hepaticis. III. A contribution to the study of the genus Ceratolejeunea (Spruce) Schifner. Candollea 14: 163–252. Bonner CEB (1953b) De hepaticis. II. An unpublished section of volume 6 of Stephani’s Species hepaticarum: the genus Marchantia. Candollea 14: 101–112. 678 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Bonner CEB (1963) Index hepaticarum. Pars III. Barbilophozia to Ceranthus. J. Cramer, Weinheim, 636 pp. Bornefeld T, Volk OH, Wolf R (1996) Exormotheca bulbigena sp. nov. (Hepaticae, Marchantiales) and its relation to E. holstii in southern Africa. Bothalia 26 (2): 159–165. doi: 10.4102/abc.v26i2.703 Borrer W, Babington CC, Berkeley MJ, Wilson W (1849) Supplement to the English botany of the late sir. J. E. Smith and Mr. Sowerby, vol. IV. J. Davis, London, 2868–2960. Bory JBGM (1804) Voyage dans les quatre principales îles des mers d’Afrique. Tome Second. F. Buisson, Paris, 431 pp. Bory JBGM, Montagne JFC (1844) Sur un nouveau genre de la famille des hépatiques. Annales des Sciences Naturelles; Botanique (sér. 3) 1: 223–235. Bottini A, Massalongo CB, Levier E (1900) Muscinee dell’Isola del Giglio. In: Sommier S (Ed.) Isola dell’Giglio. C. Clausen, Torino, 103–124. Boulay JN (1904) Muscinées de la France. Deuxième partie. Hépatiques. Paul Klincksieck, Paris, 224 pp. Bouman AC, Dirkse GM, Yamada K (1988) Radula jonesii spec. nov. (Hepaticae) a new species from Tenerife. Journal of Bryology 15 (1): 161–164. doi: 10.1179/jbr.1988.15.1.161 Braithwaite R (1878) Riccia spuria, Dickson. Journal of Botany, British and Foreign 16: 55. Braun A (1821) Bemerkungen über einige Lebermoose. Flora 4 (2): 754–757. Breidler J (1894) Die Lebermoose Steiermarks. Mitteilungen des Naturwissenschaftlichen Vereines für Steiermark 30: 256–357. Brinkman AH (1931) Notes on some Canadian hepatics, no. 2. Bryologist 34 (2): 13–16. doi: 10.2307/3239812 Briscoe L, Engel JJ, Söderström L, Hagborg A, von Konrat M (2015) Notes on Early Land Plants Today. 66. Nomenclatural notes on Acrobolbaceae. Phytotaxa 202 (1): 58–62. doi: 10.11646/phytotaxa.202.1.8 Brito ES, Ilkiu-Borges AL (2012) A new species of Ceratolejeunea Jack & Steph. (Lejeuneaceae, Jungermanniopsida) from a remnant of Amazonian forest in Maranhão, Brazil. Nova Hedwigia 95 (3/4): 423–428. doi: 10.1127/0029-5035/2012/0066 Britton NL (1915) he vegetation of Mona Island. Annals of the Missouri Botanical Garden 2 (1/2): 33–58. doi: 10.2307/2990028 Brockmüller H (1863) Beiträge zur Kryptogamenlora Mecklenburgs. Archiv des Vereins der Freunde der Naturgeschichte in Mecklenburg 17: 162–256. Brotero FA (1804) Flora lusitanica, pars II. Typographia Regia, Olissopone [Lisboa], 607 pp. Brotherus VF (1872) Musci fenniae exsiccatae, fasc. 2. Helsinki, no. 51–100. Brown EA, Braggins JE (1989) A revision of the genus Riccardia S. F. Gray in New Zealand with notes on the genus Aneura Dum. Journal of the Hattori Botanical Laboratory 66: 1–132. Brown EA, Pócs T (2001) A new species of Radula sect. Cavifolium (Radulaceae: Hepaticae) from Queensland, Australia. Telopea 9 (3): 435–438. doi: 10.7751/telopea20024000 Brown EA, Renner MAM (2014) he genus Acromastigum in Australia. Telopea 17: 251–293. doi: 10.7751/telopea20147708 Brown EA, Söderström L, Hagborg A, von Konrat MJ (2012) Notes on Early Land Plants Today. 10. A new combination in Acromastigum. Phytotaxa 65: 58. World checklist of hornworts and liverworts 679 Bryhn N (1892) Scapania crassiretis sp. nov. Revue Bryologique 19 (1): 7–8. Bryhn N (1906) Bryophyta in itinere polari norvagorum. Report of the Second Norwegian Arctic Expedition in the “Fram” 1898–1902 11: 1–260. Buch H (1916) Studien über die Scapanien Fenno-Scandias – 1. Scapania Curta-Gruppe. Meddelanden af Societas pro Fauna et Flora Fennica 42: 85–96. Buch H (1927) Cladopodiella statt Cladopus. Memoranda Societatis pro Fauna et Flora Fennica 1: 89–90. Buch H (1928) Die Scapanien Nordeuropas und Sibiriens – Systematischer Teil. Akademische Buchhandlung, Helsingfors, 1–177. Buch H (1929) Eine neue moossystematische Methodik. In: Winge Ø (Ed.) Beretning om, det 18. Skandinaviske naturforskermøde i København 26.-31. August 1929. Frederiksberg Bogtrykkeri, København, 225–229. Buch H (1932) Vorarbeiten zu einer Lebermooslora Fenno-scandias. I. Ein Versuch zur Aufspaltung der Gattungen Lophozia Dum. und Sphenolobus Steph. Memoranda Societatis pro Fauna et Flora Fennica 8: 282–297. Buch H (1942) Vorarbeiten zur eine Lebermooslora Fenno-Scandias. VII. Über die verwandtschaftlichen Beziehungen zwischen den Arten der Gattungen Sphenolobus St. sens. lat. und Lophozia Dum. sens. lat. Memoranda Societatis pro Fauna et Flora Fennica 17: 283–290. Buch H (1944) Lophozia perssonii Buch et S. Arnell, spec. nov. Botaniska Notiser 97: 381– 387. Buch H (1951) Orthocaulis cavifolius Buch und S. Arnell, ein für die Wissenschaft neues Lebermoose. Memoranda Societatis pro Fauna et Flora Fennica 26: 71–74. Buch H, Persson H (1941) Bryophyten von den Azoren und Madeira. Akademische Buchhandlung, Helsingfors, 1–16. Buch H, Tuomikoski R (1936) Scapania sphaerifera spec. nova auctore Buch et Tuomikoski. Memoranda Societatis pro Fauna et Flora Fennica 11: 227–229. Buch H, Evans AW, Verdoorn F (1937) A preliminary check list of the hepaticae of Europe and America (north of Mexico). Annales Bryologici 10: 3–8. Buchbender V, Fischer E (2004) Drepanolejeunea vandenberghenii (Jungermanniopsida: Lejeuneaceae), a previously overlooked new species from Rwanda including comments on D. physaefolia and Harpalejeunea ischeri. Journal of Bryology 26 (4): 273–283. doi: 10.1179/174328204X19469 Buchloh G (1961) Einige species novae und Neufunde von Moosen aus den Anden von Peru. Nova Hedwigia 3 (4): 507–516. Buczkowska K (2010) Morphological diferentiation of Calypogeia muelleriana (Jungermanniales, Hepaticae) in Poland. Biodiversity: Research and Conservation 17: 23–32. doi: 10.2478/v10119-010-0004-4 Buczkowska K, Bączkiewicz A (2011) New taxon of the genus Calypogeia (Jungermanniales, Hepaticae) in Poland. Acta Societatis Botanicorum Poloniae 80 (4): 327–333. doi: 10.5586/asbp.2011.039 680 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Buczkowska K, Sawicki J, Szczecińska M, Rozadziński S, Rabska M, Bączkiewicz A (2011) Two morphologically distinct groups of the Calypogeia issa complex were found in Europe. Biodiversity: Research and Conservation 23: 29–41. doi: 10.2478/v10119-011-0014-x Buczkowska K, Sawicki J, Szczecińska M, Klama H, Bączkiewicz A (2012a) Allopolyploid speciation of Calypogeia sphagnicola (Jungermanniopsida, Calypogeiaceae) based on isozyme and DNA markers. Plant Systematics and Evolution 298 (3): 549–560. doi: 10.1007/ s00606-011-0565-5 Buczkowska K, Sawicki J, Szczecińska M, Klama H, Bączkiewicz A (2012b) Isozyme and DNA markers reveal a new genetically distinct taxon of Calypogeia sphagnicola (Jungermanniopsida, Calypogeiaceae). Polish Botanical Journal 57 (1): 95–107. Burghardt M, Gradstein SR (2008) A revision of Tylimanthus (Acrobolbaceae, Marchantiophyta) in Tropical America, Africa, and Macaronesia. Fieldiana: Botany (n.ser.) 47: 199– 210. doi: 10.3158/0015-0746-47.1.199 Burghardt M, Gradstein SR, Váňa J (2006) Discovery of the African liverwort genus Cephalojonesia (Cephaloziellaceae) in Mexico. Journal of the Hattori Botanical Laboratory 100: 35–39. Burrell WH (1911) Lophozia schultzii (Nees) Schifn. var. nov. laxa. Journal of Botany, British and Foreign 49: 217–219. Campbell DH (1896) A new Californian liverwort. Botanical Gazette 21 (1): 9–13. doi: 10.1086/327288 Campbell DH (1907) Studies on some Javanese Anthocerotaceae. I. Annals of Botany. Oxford 21 (4): 467–486. Campbell DH (1915) he morphology and systematic position of Podomitrium. American Journal of Botany 2 (5): 199–210. doi: 10.2307/2435241 Campbell EO (1971) Liverworts collected in Fiji by A. C. Smith and W. Greenwood. Journal of the Royal Society of New Zealand 1 (1): 7–30. doi: 10.1080/03036758.1971.10419353 Campbell EO (1982) Notes on some anthocerotae of New Zealand (3). Tuatara 26 (1): 20–26. Campbell EO (1987) Steereomitrium minutum gen. et sp. nov. (Calobryales). Memoirs of the New York Botanical Garden 45: 569–574. Campbell EO, Outred HA (1995) Phaeoceros delicatus a new species of anthocerotae from New Zealand. New Zealand Journal of Botany 33 (3): 285–290. doi: 10.1080/0028825X.1995.10412956 Campelo MJA, Pôrto KC (2007) Briolora epíita e epíila da RPPN Frei Caneca, Jaqueira, PE, Brasil. Acta Botanica Brasilica 21 (1): 185–192. doi: 10.1590/S0102-33062007000100017 Cao T, Sun J, Yu J, Song G-Y (2003) Scapania gaochii, a new species of hepaticae from Yunnan, China. Acta Botanica Yunnanica 25 (5): 541–543. Cao T, Gao C, Sung J, Yu J, Song G-Y, Zuo B-R (2004) Scapania macroparaphyllia, a new species of Scapania (Scapaniaceae) from Xizang, China. Acta Phytotaxonomica Sinica 42 (2): 180–182. Cao T, Váňa J, Söderström L, Hagborg A, von Konrat M (2013) Notes on Early Land Plants Today. 34. Validation of Scapania gaochii (Scapaniaceae, Marchantiophyta). Phytotaxa 97 (2): 26. doi: 10.11646/phytotaxa.97.2.1 World checklist of hornworts and liverworts 681 Cargill DC, Fuhrer BA (2008) Taxonomic Studies of the Australian Anthocerotophyta II: he genus Phaeoceros. Fieldiana: Botany (n.ser.) 47: 239–253. doi: 10.3158/0015-074647.1.239 Cargill DC, Milne J (2013) A new terrestrial genus and species within the aquatic liverwort family Riellaceae (Sphaerocarpales) from Australia. Polish Botanical Journal 58 (1): 71–80. doi: 10.2478/pbj-2013-0008 Cargill DC, Scott GAM (1997) Taxonomic studies of the Australian Anthocerotales. I. Journal of the Hattori Botanical Laboratory 82: 47–60. Cargill DC, Söderström L, Hagborg A, von Konrat M (2012) Notes on Early Land Plants Today. 3. Fossombronia ruminata Cargill, a new name for Fossombronia maritima G.A.M.Scott et D.C.Pike ex Cargill, nom. illeg. Phytotaxa 65: 45. Cargill DC, Vella NGF, Sharma I, Miller JT (2013a) Cryptic speciation and species diversity among Australian and New Zealand hornwort taxa of Megaceros (Dendrocerotaceae). Australian Systematic Botany 26 (5): 356–377. doi: 10.1071/SB13030 Cargill DC, Söderström L, Hagborg A, von Konrat M (2013b) Notes on Early Land Plants Today. 23. A new synonym in Anthoceros (Anthocerotaceae, Anthocerotophyta). Phytotaxa 76 (3): 48–49. doi: 10.11646/phytotaxa.76.3.11 Carl H (1931a) Morphologische Studien an Chiastocaulon Carl, einer neuen Lebermoosgattung. Flora 126: 45–60. Carl H (1931b) Die Arttypen und die systematische Gliederung der Gattung Plagiochila Dum. Annales Bryologici, suppl. 2: 1–170. Carr DJ (1956) Contributions to Australian bryology. I. he structure, development, and systematic ainities of Monocarpus sphaerocarpus gen. et spec. nov. (Marchantiales). Australian Journal of Botany 4 (2): 175–191. doi: 10.1071/BT9560175 Carr DJ (2004) Lejeuneaceae (Hepaticae) of Victoria; additions and a restitution. Proceedings of the Royal Society of Victoria 116 (2): 221–232. Carr DJ (2005) Two new bryophytes in Victoria. Proceedings of the Royal Society of Victoria 117 (2): 319–325. Carrington B (1863) Irish hepaticae. Transactions of the Botanical Society of Edinburgh 7 (3): 441–458. doi: 10.1080/03746606309467881 Carrington B (1870) On two new British hepaticae. Transactions of the Botanical Society of Edinburgh 10: 378–382. doi: 10.1080/03746607009468722 Carrington B (1881) Hepaticae. In: Lees FA (Ed.) he London catalogue of British mosses and hepatics, second edition, London, 21–26. Carrington B, Pearson WH (1878) Hepaticae britannicae exsiccatae, fasc. I. Manchester, no. 1–75. Carrington B, Pearson WH (1879) Hepaticae britannicae exsiccatae, fasc. II. Manchester, no. 76–150. Carrington B, Pearson WH (1888a) List of hepaticae collected by Mr homas Whitelegge in New South Wales, 1884-5. Proceedings of the Linnean Society of New South Wales (ser. 2) 2 (4): 1035–1060. 682 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Carrington B, Pearson WH (1888b) Description of new or rare Tasmanian hepaticae (collected by R.A. Bastow, Esq., F.L.S.). Papers and Proceedings of the Royal Society of Tasmania 1887: 1–12. Carrington B, Pearson WH (1889) A new hepatic. Journal of Botany, British and Foreign 27: 225. Carruthers W (1865) On the nomenclature of the British hepaticae. Journal of Botany, British and Foreign 3: 297–302. Castle H (1938) Radula evansii. Annales Bryologici 11: 37–39. Castle H (1959) A revision of the genus Radula. Part II. Subgenus Acroradula. Section 3. Dichotomae. Journal of the Hattori Botanical Laboratory 21: 1–52. Castle H (1962) A revision of the genus Radula. Part II. Subgenus Acroradula. Section 7. Lingulatae. Revue Bryologique et Lichénologique 31 (3/4): 139–151. Castle H (1965) A revision of the genus Radula. Part II. Subgenus Acroradula. Section 9. Densifoliae. Revue Bryologique et Lichénologique 33 (3/4): 328–398. Castle H (1968) Radula loridana sp. nov. Revue Bryologique et Lichénologique 36 (1/2): 1–4. Cavers F (1903) A new species of Riella (R. capensis) from South Africa. Revue Bryologique 30 (5): 81–84. Chang K-C (1988) Notes on the Porellaceae of China. Bulletin of Botanical Research. Harbin 8 (2): 43–47. Chang K-C, Gao C (1984) Plantae novae hepaticarum sinarum. Bulletin of Botanical Research. Harbin 4 (3): 83–99. Chantanaorrapint S (2009) Phaeoceros perpusillus (Notothyladaceae), a new species of hornwort from hailand. Acta Botanica Hungarica 51 (1/2): 29–33. doi: 10.1556/ ABot.51.2009.1-2.6 Chantanaorrapint S (2014) Notothylas irregularis (Notothyladaceae, Anthocerotophyta), a new species of hornwort from northern hailand. Acta Botanica Hungarica 56 (3/4): 269–274. doi: 10.1556/ABot.56.2014.3–4.3 Chantanaorrapint S, Pócs T (2014) Southern hailand bryophytes I, with description of Cololejeunea ramromensis. In: Telnov D (Ed.) Biodiversity, biogeography and nature conservation in Wallacea and New Guinea, volume II. Entomological Society of Latvia, Riga, 113–122. Chantanaorrapint S, Peng T, Zhu R-L (2014) Reappraisal of Dendroceros cucullatus (Dendrocerotaceae, Anthocerotophyta). Phytotaxa 167 (1): 145–149. doi: 10.11646/phytotaxa.167.1.14 Chen P-C (1955) Bryophyta nova sinica. Feddes Repertorium Specierum Novarum Regni Vegetabilis 58: 23–52. Chen P-C, Wu P-C (1964) [Study on epiphyllous liverworts of China. (1)]. Acta Phytotaxonomica Sinica 9 (3): 213–276. Chen P-C, Wu P-C (1965) he preliminary study of the bryophytes of Mt. Hwangshan. In: NN (Ed.) Observationes ad lorulam Hwangshanicum. Scientiic and Technical Publishers, Shanghai, 1–59. Choi SS, Bakalin VA, Sun B-Y (2012) Scapania and Macrodiplophyllum in the Russian Far East. Botanica Paciica 1: 31–95. doi: 10.17581/bp.2012.01104 World checklist of hornworts and liverworts 683 Chopra RS (1943) A census catalogue of Indian hepatics. Journal of the Indian Botanical Society 22: 237–259. Churchill SP, Sanjines NN, Fuentes A, Lozano R (2008) Additions to the Bolivian bryophyte lora. Tropical Bryology 29: 54–59. Clark L (1953) Some hepaticae from Galápagos, Cocos and other Paciic coast islands. Proceedings of the California Academy of Sciences (ser. 4) 27 (18): 593–624. Clark L (1954) Frullania armata. Bryologist 57 (1): 36–39. doi: 10.2307/3240074 Clark L, Frye TC (1952) Frullania tunguraguana. Bryologist 55 (2): 133–136. doi: 10.2307/3240201 Clark L, Schultz MR (1953) Frullania hamilora. Bryologist 56 (3): 180–182. doi: 10.2307/3240116 Clark L, Svihla RD (1950) A new Frullania from Panama. Bryologist 53 (1): 63–66. doi: 10.2307/3240261 Clark L, Jovet-Ast S, Frye TC (1947) A new Frullania from Guadeloupe. Bryologist 50 (1): 52–55. doi: 10.2307/3239584 Colenso W (1884) A further contribution toward making known the botany of New Zealand. Transactions and Proceedings of the New Zealand Institute 16: 325–363. Colenso W (1885) A description of some newly-discovered and rare indigenous plants; being a further contribution toward making known the botany of New Zealand. Transactions and Proceedings of the New Zealand Institute 17: 237–265. Colenso W (1886a) A descriptions of some newly-discovered and rare indigenous plants. Transactions and Proceedings of the New Zealand Institute 18: 256–287. Colenso W (1886b) A description of some newly-discovered cryptogamic plants, being a further contribution towards the making known the botany of New Zealand. Transactions and Proceedings of the New Zealand Institute 18: 219–255. Colenso W (1887) A description of some newly-discovered cryptogamic plants, being a further contribution toward making known the botany of New Zealand. Transactions and Proceedings of the New Zealand Institute 19: 271–301. Colenso W (1888) On new indigenous cryptogams of the orders Lycopodiaceae, Musci and Hepaticae. Transactions and Proceedings of the New Zealand Institute 20: 234–254. Colenso W (1889) A description of some newly-discovered cryptogamic plants; being a further contribution toward making known the botany of New Zealand. Transactions and Proceedings of the New Zealand Institute 21: 43–80. Colenso W (1892) A list of new species of hepaticae Novae Zealandiae, named by F. Stephani, Leipzig. Transactions and Proceedings of the New Zealand Institute 24: 398–400. Cooke CM (1904) he Hawaiian hepaticae of the tribe Trigonantheae. Transactions of the Connecticut Academy of Arts and Sciences 12 (1): 1–44. Cooper ED (2013) Notes on Early Land Plants Today. 37. Towards a stable, informative classiication of the Lepidoziaceae (Marchantiophyta). Phytotaxa 97 (2): 44–51. doi: 10.11646/phytotaxa.97.2.4 Cooper ED, Renner MAM (2014) Lepidozia bragginsiana, a new species from New Zealand. Phytotaxa 173 (2): 117–126. doi: 10.11646/phytotaxa.173.2.2 684 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Cooper ED, Shaw AJ, Shaw B, Henwood MJ, Heslewood MM, Brown EA (2011) A multi-locus molecular phylogeny of the Lepidoziaceae: Laying the foundations for a stable classiication. Molecular Phylogenetics and Evolution 59 (2): 489–509. doi: 10.1016/j. ympev.2011.02.006 Cooper ED, Henwood MJ, Brown EA (2012a) A molecular phylogeny of the Lepidozia generic complex supports re-circumscription of the Lepidozioideae. Molecular Phylogenetics and Evolution 65 (1): 10–22. doi: 10.1016/j.ympev.2012.05.008 Cooper ED, Henwood MJ, Brown EA (2012b) Are the liverworts really that old? Cretaceous origins and Cenozoic diversiications in Lepidoziaceae relect a recurrent theme in liverwort. Journal of the Linnean Society. Biology 107 (2): 425–441. doi: 10.1111/j.10958312.2012.01946.x Cooper ED, Söderström L, Hagborg A, von Konrat MJ (2013) Notes on Early Land Plants Today. 38. New combinations and synonyms in Lepidoziaceae (Marchantiophyta). Phytotaxa 97 (2): 52–62. doi: 10.11646/phytotaxa.97.2.5 Cooper ED, Söderström L, Hagborg A, von Konrat M (2014) Notes on Early Land Plants Today. 54. A transfer in Lepidoziaceae (Marchantiophyta). Phytotaxa 167 (2): 218–219. doi: 10.11646/phytotaxa.167.2.13 Corbière L (1912) Nouvelles muscinées de l’Afrique tropicale. Bulletin du Muséum National d’Histoire Naturelle 18 (2): 108–121. Corda AJC (1829) Genera Hepaticarum. In: Opiz PM (Ed.) Beiträge zur Naturgeschichte als Fortsetzung des Naturalientausches No. 12. C.W. Enders, Praha, 643–655. Corda AJC (1830) Deutschlands Jungermannien, Heft 19–20. In: Sturm J (Ed.) Deutschlands Flora in Abbildungen nach der Natur mit Beschreibungen. Abtheilung II Cryptogamie. Privately published by author, Nürnberg, 1–62. Costa DP (2008) Metzgeriaceae. Flora Neotropica, Monograph 102: 1–169. Costa DP (2009) Crittogame brasiliane, a review of Guiseppe Raddi bryophyte collections in the state of Rio de Janeiro. Journal of Bryology 31 (4): 222–233. doi: 10.1179/03736680 9X12495600997402 Costa DP, Gradstein SR (2000) On the status of Metzgeria angusta, a poorly known neotropical taxon. Bryologist 103 (4): 757–759. doi: 10.1639/0007-2745(2000)103[0757:OTSOMA] 2.0.CO;2 Costa DP, Santos ND, Váňa J (2008) A new species of Cylindrocolea (Cephaloziellaceae) from Brazil. Bryologist 111 (4): 667–669. doi: 10.1639/0007-2745-111.4.667 Coulter JM, Barnes CR, Arthur JC (1890) General index. Botanical Gazette 15 (12): 347–352. Coxson DS (1991) Nutrient release from epiphytic bryophytes in tropical montane rain forest (Guadeloupe). Canadian Journal of Botany 69 (10): 2122–2129. doi: 10.1139/b91-266 Crandall-Stotler BJ, Stotler RE (2000) Morphology and classiication of the Marchantiophyta. In: Shaw AJ, Goinet B (Eds) Bryophyte biology. Cambridge University Press, Cambridge, 21–70. doi: 10.1017/cbo9781139171304.003 Crandall-Stotler BJ, Stotler RE (2007) On the identity of Moerckia hibernica (Hook.) Gottsche (Moerckiaceae fam. nov., Marchantiophyta). Beihefte zur Nova Hedwigia 131: 41–59. Crandall-Stotler BJ, Stotler RE, Ford CH (2002) Contributions toward a monograph of Petalophyllum (Marchantiophyta). Novon 12 (3): 334–337. doi: 10.5962/bhl.title.744 World checklist of hornworts and liverworts 685 Crandall-Stotler BJ, Stotler RE, Doyle WT, Forrest LL (2008a) Phaeoceros proskaueri sp. nov., a new species of the Phaeoceros hallii (Austin) Prosk. –Phaeoceros pearsonii (M. Howe) Prosk. complex and the systematic ainities of Paraphymatoceros Hässel. Fieldiana: Botany (n.ser.) 47: 213–238. doi: 10.3158/0015-0746-47.1.213 Crandall-Stotler BJ, Stotler R, Long DG (2008b) Morphology and classiication of the Marchantiophyta. In: Goinet B, Shaw J (Eds) Bryophyte Biology, 2nd ed. Cambridge University Press, Cambridge, 1–54. Crandall-Stotler BJ, Stotler RE, Long DG (2009) Phylogeny and classiication of the Marchantiophyta. Edinburgh Journal of Botany 66 (1): 155–198. doi: 10.1017/S0960428609005393 Crandall-Stotler BJ, Stotler R, Zhang L, Forrest L (2010) On the morphology, systematics and phylogeny of Noteroclada (Noterocladaceae, Marchantiophyta). Nova Hedwigia 91 (3/4): 421–450. doi: 10.1127/0029-5035/2010/0091-0421 Crandall-Stotler BJ, Stotler R, Bakalin VA, Doyle WT (2013) A new species of Mesoptychia (Lindb.) A. Evans from California. Polish Botanical Journal 58 (1): 81–89. doi: 10.2478/pbj-2013-0009 Crane PR (2004) Documenting plant diversity: uninished business. Philosophical Transactions of the Royal Society of London. Series B 359 (1444): 735–737. doi: 10.1098/ rstb.2003.1441 Crantz D (1770) Fortsättning af historien om Grönland. Johan Georg Lange, Stockholm, 380 pp. (non vidi) Cremer KW, Mount AB (1965) Early stages of plant succession following the complete felling and burning of Eucalyptus regnans forest in the Florentine Valley, Tasmania. Australian Journal of Botany 13 (2): 303–322. doi: 10.1071/BT9650303 Cronquist A (1988) Schuster and Wallace’s line. Beihefte zur Nova Hedwigia 90: 39–40. Crosby MR, Engel JJ (2006) Index of hepatics 1974–2000. Hattori Botanical Laboratory, Nichinan, 368 pp. Crosby MR, Magill RE (2005) Index of bryophytes 2001-2004. Missouri Botanical Garden, St. Louis, 31 pp. Crosby MR, Magill RE (2006) Index of bryophytes 2005. Missouri Botanical Garden, St. Louis, 12 pp. Crozals A (1903a) Flore bryologique de Roquehaute (Hérault). Revue Bryologique 30 (2): 17–32. Crozals A (1903b) Riccia subbifurca Warn., in litt. Revue Bryologique 30 (4): 62–64. Crum HA, Bruce J (1996) A new species of Cryptothallus from Costa Rica. Bryologist 99 (4): 433–438. doi: 10.2307/3244107 Crundwell AC (1970) Herberta borealis, a new species from Scotland and Norway. Transactions of the British Bryological Society 6 (1): 41–49. doi: 10.1179/006813870804146464 Cufodontis G (1952) Enumeratio plantarum aethiopiae, II (Bryophyta: Hepaticae). Phyton (Horn) 4: 72–82. Dabhade GT, Hasan A (1986) New species of Riccia –Riccia indira-gandhiensis sp. nov. Journal of the Bombay Natural History Society 83 (2): 398–401. Damsholt K (2002) Illustrated lora of Nordic liverworts and hornworts. Nordic Bryological Society, Lund, 837 pp. Damsholt K, Hallingbäck T (1986) Riccia gothica, a new species of hepaticae from Sweden. Lindbergia 12 (2/3): 100–102. 686 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Damsholt K, Váňa J (1977) he genus Jungermannia L. emend. Dumort. (Hepaticae) in Greenland. Lindbergia 4 (1/2): 1–26. Daniels AED, Daniel P (2002) Two new species of Riccia L. (Hepaticae: Marchantiales) from the Western Ghats of Tamil Nadu. Bulletin of the Botanical Survey of India 44 (1/4): 135–140. Daniels AED, Daniel P (2007) Name changes in two Indian liverworts. Bulletin of the Botanical Survey of India 49 (1/4): 231–232. Daniels AED, Daniel P (2013) he bryolora of the southernmost Western Ghats, India. Bishen Singh Madendra Pal Singh, Dehra Dun, 352 pp. Das S, Singh DK (2008) Leptolejeunea arunachalensis (Lejeuneaceae, Hepaticae) – a new species from India. Journal of Japanese Botany 83 (6): 343–346. Das S, Singh DK (2011) A new species of Calypogeia Raddi (Marchantiophyta : Calypogeiaceae) from Eastern Himalaya, India. Nelumbo 53: 194–196. Dauphin G (2003) he genus Ceratolejeunea Jack & Steph. (Hepaticae: Lejeuneaceae) in Tropical America. Flora Neotropica, Monograph 90: 1–86. Dauphin G (2005) Catalogue of Costa Rican hepaticae and anthocerotae. Tropical Bryology 26: 141–218. Dauphin G, Gradstein SR (2003) A new species of Cheilolejeunea (Spruce) Schifn. from Panamá. Journal of Bryology 25 (4): 259–261. doi: 10.1179/037366803225013128 Dauphin G, Pócs T, Villarreal JC, Salazar N (2006) Nuevos registros de hepáticas y anthocerotóitas para Panamá. Tropical Bryology 27: 73–85. Dauphin G, Morales T, Moreno EJ (2008) Catálogo preliminar de Lejeuneaceae (Hepaticae) de Venezuela. Cryptogamie, Bryologie 29 (3): 215–265. Davis EC (2004) A molecular phylogeny of leafy liverworts (Jungermnannidae: Marchantiophyta). Monographs in Systematic Botany from the Missouri Botanical Garden 98: 61–86. De Candolle A-P, Lamarck JBAP (1815) Flore française, tome cinquième, ou sixième volume. Desray, Paris, 662 pp. De Lucia EH, Turnbull MH, Walcroft AS, Grifen KL, Tissue DT, Glenny D, McSeveny TM, Whitehead D (2003) he contribution of bryophytes to the carbon exchange for a temperate rainforest. Global Change Biology 9 (8): 1158–1170. doi: 10.1046/j.13652486.2003.00650.x De Notaris G (1838) Primitiae hepaticologiae italicae. Memorie della reale accademia delle scienze di Torino (ser. 2) 1: 287–354. De Notaris G (1857) Jungermanniearum americanarum pugillus. Memorie della reale accademia delle scienze di Torino (ser. 2) 16: 211–238. De Notaris G (1859) Appunti per un nuovo censimento delle epatiche italiane. Memorie della reale accademia delle scienze di Torino (ser. 2) 18: 457–498. De Notaris G (1874) Epatiche di Borneo. Stamperia reale di G. B. Paravia E. C., Torino, 42 pp. de Roo RT, Hedderson TA, Söderström L (2007) Molecular insights into the phylogeny of the leafy liverwort family Lophoziaceae Cavers. Taxon 56 (2): 301–314. Del Rosario RM (1971) New and noteworthy Philippine liverworts, II. Philippine Journal of Science 100 (3/4): 227–242. World checklist of hornworts and liverworts 687 Del Rosario RM (1977) Philippine liverworts. III. Calobryales and Herbertales of the Philippines. Philippine Journal of Science 104 (3/4): 93–209. Demaret F, Vanden Berghen C (1950) Révision du quelques espèces africaines du genre Frullania Raddi, sous-genre Meteoriopsis Spruce. Bulletin du Jardin Botanique de l’État, Bruxelles 20 (1): 1–10. doi: 10.2307/3666581 Devos N, Renner MAM, Gradstein SR, Shaw AJ, Vanderpoorten A (2011) Molecular data challenge traditional subgeneric divisions in the leafy liverwort Radula. Taxon 60 (6): 1623–1632. Dey M, Singh DK (2008) Frullania pran-nathii – a new epiphyllous liverwort from Darjeeling, India. Journal of Japanese Botany 83 (5): 280–283. Dey M, Singh DK (2010) Two new epiphyllous Leptolejeunea (Hepaticae: Lejeuneaceae) from Eastern Himalaya, India. Taiwania 55 (4): 355–362. Dey M, Singh DK (2011) A new Lopholejeunea (Spruce) Schifn. (Hepaticae: Lejeuneaceae) from India. Nelumbo 53: 197–200. Dey M, Singh DK, Singh D (2008) Two new species of Lejeunea Lib. (Hepaticae: Lejeuneaceae) from Sikkim, India. Journal of Bryology 30 (2): 126–132. doi: 10.1179/174328208X300651 Dickson J (1785) Fasciculus plantarum cryptogamicarum britanniae. G. Nichol, London, 26 pp. Dickson J (1790) Fasciculus secundus plantarum cryptogamicarum britanniae. G. Nichol, London, 31 pp. Dickson J (1793) Fasciculus tertius plantarum cryptogamicarum britanniae. G. Nichol, London, 24 pp. Dickson J (1801) Fasciculus quartus plantarum cryptogamicarum britanniae. G. Nichol, London, 28 pp. Dixon HN, Schifner V, Verdoorn, F (1932) Bryophyta nova (1–5). Annales Bryologici 5: 157–164. Doei H (1987) A new species of Lepidozia (Hepaticae) from Seram Island, Indonesia. Journal of the Hattori Botanical Laboratory 63: 421–424. Dong S, Schäfer-Verwimp A, Meinecke P, Feldberg K, Bombosch A, Pócs T, Schmidt AR, Reitner J, Schneider H, Heinrichs J (2012) Tramps, narrow endemics and morphologically cryptic species in the epiphyllous liverwort Diplasiolejeunea. Molecular Phylogenetics and Evolution 65 (2): 582–594. doi: 10.1016/j.ympev.2012.07.009 Dong S, Schäfer-Verwimp A, Pócs T, Feldberg K, Czumaj A, Schmidt AR, Schneider H, Heinrichs J (2013) Size doesn’t matter – recircumscription of Microlejeunea (Lejeuneaceae, Porellales) based on molecular and morphological evidence. Phytotaxa 85 (2): 41–55. doi: 10.11646/phytotaxa.85.2.2 Douin C (1903) Le Sphaerocarpus terrestris Sm. Revue Bryologique 30 (3): 44–57. Douin C (1913a) Cephaloziella obtusa P. Culmann sp. nov. Revue Bryologique 40 (5): 65–71. Douin C (1913b) L’inlorescence des Céphaloziellacées. Revue Bryologique 40 (6): 81–87. Douin C (1914) Les mélanges d’espèces chez les Céphaloziellacées. Revue Bryologique 41 (1): 1–8. Douin C (1920) La famille des Céphaloziellacées. Mémoires de la Société Botanique de France 29: 1–90. Douin C, Trabut ML (1919) Deux hépatiques peu connues. Revue Générale de Botanique 31: 321–328. 688 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Drège JF (1843) Alphabetisches Verzeichniss der von I. F. Drège in Südafrika gesammelten Planzen, nebst Angabe der Fundorte. Flora, Beigabe 26: 161–230. Duda J (1970) Lophozia (subgenus Leiocolea) mamatkulovii Duda sp. nov. von Pamir (SSSR). Transactions of the British Bryological Society 6 (1): 82–85. doi: 10.1179/006813870804146428 Duf RJ, Villarreal JC, Cargill DC, Renzaglia KS (2007) Progress and challenges toward developing a phylogeny and classiication of the hornworts. Bryologist 110 (2): 214–243. doi: 10.1639/0007-2745(2007)110[214:PACTDA]2.0.CO;2 Dugas M (1928) Contribution à l’étude du genre Plagiochila Dum. Masson et Cie, Paris, 1–199. Dumortier BCJ (1822) Commentationes botanicae. Ch. Casterman-Dien, Tournay, 118 pp. doi: 10.5962/bhl.title.10534 Dumortier BCJ (1831) Sylloge Jungermannidearum Europae indigenarum. J. Casterman, Tournay, 100 pp. doi: 10.5962/bhl.title.22343 Dumortier BCJ (1835) Recueil d’Observations sur les Jungermanniacées. J.-A. Blanquart, Tournay, 27 pp. doi: 10.5962/bhl.title.731 Dumortier BCJ (1874) Jungermannideae Europae. Bulletin de la Société Royale de Botanique de Belgique 13: 5–203. Duss A (1903) Énumération méthodique des muscinées des Antilles françaises. I. – Hépatiques. Lucien Declume, Lons-le-Saunier, 41 pp. Duthie AV, Garside S (1936) Studies in South African Ricciaceae I. hree annual species: R. plana Taylor, R. cupulifera sp. nov., and R. Curtisii T. P. James. Transactions of the Royal Society of South Africa 24 (2): 93–133. doi: 10.1080/00359193609520543 Duthie AV, Garside S (1939) Studies in South African Ricciaceae II. he annual species of the section Ricciella (concluded): R. compacta sp. nov., and R. Rautanenii Steph. Transactions of the Royal Society of South Africa 27 (1): 17–28. doi: 10.1080/00359193909519784 Ehrhart JF (1783) Meine Reise nach der Grafschaft Bentheim, un von da nach Holland, nebst der Retour nach Herrenhausen (Fortsetzung). Hannoverisches Magazin 21 (18): 273–288. Ehrhart JF (1784) Botanische Bemerkungen. Hannoverisches Magazin 22 (8): 113–144. Ehrhart JF (1789) Kennzeichen seltener und unbestimmter Planzen. Beiträge zur Naturkunde (Ehrhart) 4: 42–47. doi: 10.5962/bhl.title.44806 Eifrig H (1937) Monographische Studien über die indomalayischen Arten von Taxilejeunea. Friedrich-Schiller-Universität, Jena, 73–114. Elfving F (1876) Sällskapets pro Fauna et Flora Fennica sammanträde den 6 dennes. Morgonbladet (Helsinki) 1876 (106, 9 May): 1. Endlicher SFL (1833) Prodromus lorae Norfolkicae. Friedrich Beck Universitatis Bibliopolam, Vindobonae [Wien], 100 pp. doi: 10.5962/bhl.title.6703 Engel JJ (1968) A taxonomic monograph of the genus Balantiopsis (Hepaticae). Nova Hedwigia 16: 83–130. Engel JJ (1972) Chiloscyphus hookeri n. sp. and nomenclatural changes in the genus Clasmatocolea. Journal of the Hattori Botanical Laboratory 36: 150–156. World checklist of hornworts and liverworts 689 Engel JJ (1975) Hepaticae and anthocerotae collected by Dr. Harold E. Moore, Jr. in New Caledonia, Seychelles, Mauritius and Reunion in 1972. Bryologist 78 (3): 361–362. doi: 10.2307/3241895 Engel JJ (1976a) he southern South American hepaticae and anthocerotae collected by H. Roivainen in 1969–1970, with new taxa and notes on range extensions. Annales Botanici Fennici 13 (3): 132–136. Engel JJ (1976b) Austral hepaticae. VI. Some new species and new combinations of taxa from southern Chile. Bryologist 79 (4): 514–515. doi: 10.2307/3241949 Engel JJ (1978) A taxonomic and phytogeographic study of Brunswick Peninsula (Strait of Magellan) Hepaticae and Anthocerotae. Fieldiana: Botany 41: 1–319. doi: 10.5962/bhl.title.2426 Engel JJ (1979a) Austral hepaticae. XI. Lophocoleaceae: new taxa, new combinations and realignments. Phytologia 41 (5): 309–312. Engel JJ (1979b) Austral hepaticae. X. A revision of Hepatostolonophora Engel et Schust., nom. nov. (Hepaticae). Journal of the Hattori Botanical Laboratory 46: 91–108. Engel JJ (1980a) A monograph of Clasmatocolea (Hepaticae). Fieldiana: Botany (n.ser.) 3: 1–229. Engel JJ (1980b) Austral hepaticae. XII. A new species of Clasmatocolea (Hepaticae) from Tasmania. Bryologist 83 (2): 220–223. doi: 10.2307/3242138 Engel JJ (1981) Haplomitrium monoicum, a remarkable new species of Calobryales (Hepaticae) from New Caledonia, together with a reclassiication of subg. Haplomitrium. Annals of the Missouri Botanical Garden 68 (4): 668–676. doi: 10.2307/2398895 Engel JJ (1990a) Studies on Geocalycaceae (Hepaticae). I. he taxonomic position of Chiloscyphus amplectens (Mitt.) Engel & Schust. together with reinement in Heteroscyphus Schifn. Journal of the Hattori Botanical Laboratory 68: 303–315. Engel JJ (1990b) Falkland Islands (Islas Malvinas) hepaticae and Anthocerotophyta: A taxonomic and phytogeographic study. Fieldiana: Botany (n.ser.) 25: 1–209. Engel JJ (1991a) Studies on Geocalycaceae (Hepaticae). II. Stolonivector, a new genus from New Zealand. Journal of the Hattori Botanical Laboratory 69: 79–86. Engel JJ (1991b) Studies on Geocalycaceae (Hepaticae). VI. Comments on Leptoscyphus in Australasia, together with reinements in Clasmatocolea. Bryologist 94 (4): 435–437. doi: 10.2307/3243840 Engel JJ (1991c) Studies on Geocalycaceae (Hepaticae). IV. Lamellocolea, a new genus of Leptoscyphoideae from New Zealand. Journal of the Hattori Botanical Laboratory 70: 63–78. Engel JJ (1993) Studies on Geocalycaceae. IX. Chiloscyphus hattorii Engel, a new species from New Zealand, together with nomenclatural reinements in Australasian Heteroscyphus and Leptoscyphus. Journal of the Hattori Botanical Laboratory 74: 29–33. Engel JJ (1997) Studies on Geocalycaceae (Hepaticae). X. New taxa and new combinations in Chiloscyphus Corda for Australasia. Phytologia 83 (1): 42–46. Engel JJ (1999a) Austral hepaticae. 26. he identity, taxonomic position, and ecology of Trichocolea julacea Hatcher (Trichocoleaceae). Novon 9 (1): 25–28. doi: 10.5962/bhl.title.744 Engel JJ (1999b) Austral hepaticae. 30. A critical new species of Triandrophyllum (Herbertaceae) from New Zealand. Haussknechtia, Beiheft 9: 115–120. 690 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Engel JJ (2001) Studies on Geocalycaceae. XII. Heteroscyphus mononuculus Engel, a new species of hepaticae from New Zealand. Journal of the Hattori Botanical Laboratory 90: 241–244. Engel JJ (2003) Studies on Geocalycaceae (Hepaticae). XIII. he genus Stolonivector Engel, including a new species from New Zealand. Journal of the Hattori Botanical Laboratory 93: 69–77. Engel JJ (2004a) Austral hepaticae. 36. A new species of Lepidozia from New Zealand, together with an assessment of subg. Austrolepidozia (Schust.) Schust. Journal of the Hattori Botanical Laboratory 96: 273–279. Engel JJ (2004b) Studies on Geocalycaceae. XV. Chiloscyphus aperticaulis Engel, an interesting new species of hepaticae from New Zealand. Journal of the Hattori Botanical Laboratory 95: 229–234. Engel JJ (2005) Austral hepaticae. 39. Kurzia moniliformis, an interesting new species of Hepaticae from New Zealand, belonging to a new section, Kurzia sect. Moniliformes Engel. Cryptogamie, Bryologie 26 (1): 73–78. Engel JJ (2006a) Austral hepaticae. 42. he austral species of Mnioloma (Calypogeiaceae), together with a new species, Mnioloma novaezelandiae n. sp. Cryptogamie, Bryologie 27 (1): 111–117. Engel JJ (2006b) Austral hepaticae. 40. Tritomaria (Jungermanniaceae subfam. Lophozioideae) new to the south temperate, together with a new subspecies, T. exsecta subsp. novaezelandiae subsp. nov. Bryologist 109 (1): 60–67. doi: 10.1639/0007-2745(2006)109[0060:AH TJSL]2.0.CO;2 Engel JJ (2006c) Austral hepaticae. 41. Bazzania exempta Engel, an interesting new species from New Zealand, belonging to a new section, Bazzania sect. Exemptae Engel. Journal of the Hattori Botanical Laboratory 99: 197–205. Engel JJ (2007) Studies of New Zealand hepaticae. 20-38. A miscellanea of new taxa and combinations. Novon 17 (3): 310–314. doi: 10.3417/1055-3177(2007)17[310:SONZH A]2.0.CO;2 Engel JJ (2009) Studies on Lophocoleaceae (Hepaticae). XVIII. Stolonivector Engel in New Zealand, including two new species, together with comments on generic endemism of austral hepaticae. Nova Hedwigia 88 (3/4): 335–346. doi: 10.1127/0029-5035/2009/0088-0335 Engel JJ (2010) Austral hepaticae. 45. A Monograph of the Genus Chiloscyphus Corda (Lophocoleaceae) for Australasia. Fieldiana: Botany (n.ser.) 48: 1–209. doi: 10.3158/0015-0746-48.1.1 Engel JJ (2011) Studies of New Zealand hepaticae. 56–68. A miscellanea of new taxa and combinations. Nova Hedwigia 93 (3/4): 401–410. doi: 10.1127/0029-5035/2011/0093-0401 Engel JJ (2013a) Austral hepaticae 50. Gackstroemia in New Zealand, together with an interesting new species. Phytotaxa 118 (1): 9–21. doi: 10.11646/phytotaxa.118.1.2 Engel JJ (2013b) Studies on Lophocoleaceae XXII. New taxa and combinations in New Zealand Heteroscyphus Schifn. Polish Botanical Journal 58 (1): 95–106. doi: 10.2478/ pbj-2013-0011 Engel JJ (2014) Studies on Lophocoleaceae XXIII. Novelties in Heteroscyphus Schifn. together with reinements in Cryptolophocolea L.Söderstr., Crand.-Stotl., Stotler & Váňa and Leptoscyphus Mitt. Nova Hedwigia 99 (1/2): 157–170. doi: 10.1127/0029-5035/2014/0208 Engel JJ (2015a) Studies on Lophocoleaceae. XXV. A conspectus of Heteroscyphus Schifn. in temperate Australasia together with nomenclatural changes in Chiloscyphus Corda and World checklist of hornworts and liverworts 691 Leptoscyphus Mitt., reinements and a range extension in Clasmatocolea Spruce, and a range extension in Stolonivector J.J.Engel. Nova Hedwigia 100 (3/4): 553–582. doi: 10.1127/ nova_hedwigia/2015/0260 Engel JJ (2015b) Studies on Lophocoleaceae. XXIV. Chiloscyphus alpicola J.J.Engel, an interesting new liverwort species from New Zealand together with nomenclatural changes in Tetracymbaliella Grolle. Phytotaxa 207 (2): 181–186. doi: 10.11646/phytotaxa.207.2.4 Engel JJ, Braggins JE (1998) Austral hepaticae. 27. he genus Anastrophyllum (Spruce) Steph. (Jungermanniales) in Australasia, with a synopsis of austral taxa. Journal of Bryology 20 (2): 371–388. doi: 10.1179/jbr.1998.20.2.371 Engel JJ, Glenny DS (2007) Austral hepaticae. 43. Castanoclobos, a new genus of Trichocoleaceae from New Zealand. Novon 17 (4): 424–428. doi: 10.3417/1055-3177(2007)17[424:AHCANG]2.0.CO;2 Engel JJ, Glenny DS (2008a) A lora of the liverworts and hornworts of New Zealand, Volume 1. Missouri Botanical Garden, St. Louis, 897 pp. Engel JJ, Glenny DS (2008b) Studies on Lophocoleaceae. XVI. Chiloscyphus anisolobus, an interesting new species from New Zealand. Bryologist 111 (1): 118–123. doi: 10.1639/0007-2745(2008)111[118:SOLXCA]2.0.CO;2 Engel JJ, Glenny DS (2008c) Austral hepaticae. 44. A revision of Marsupidium Mitt. for New Zealand. Nova Hedwigia 87 (3/4): 277–313. doi: 10.1127/0029-5035/2008/0087-0277 Engel JJ, Glenny DS (2011) Studies on Lophocoleaceae. XX. A new species of Lamellocolea J. J. Engel from New Zealand. Bryologist 114 (1): 23–27. doi: 10.1639/0007-2745-114.1.23 Engel JJ, Glenny DS (2012) Austral hepaticae. 48. Goebelobryum Grolle (Acrobolbaceae). Nova Hedwigia 95 (3/4): 319–336. doi: 10.1127/0029-5035/2012/0066 Engel JJ, Gradstein SR (2003) Studies in Geocalycaceae XIV. Physotheca J. J. Engel & Gradst., a new genus of hepaticae from Ecuador, belonging to a new subfamily, Geocalycaceae subfam. Physothecoideae J. J. Engel & Gradst. Taxon 52 (4): 763–773. doi: 10.2307/3647350 Engel JJ, Grolle R (1971) Marsupidium in South America. Journal of the Hattori Botanical Laboratory 34: 437–444. Engel JJ, He X (2010) Studies on Lophocoleaceae. XIX. he systematic identity of Cyanolophocolea R. M. Schust., an intriguing liverwort from New Zealand and Australia, based on morphological and molecular evidence. Bryologist 113 (1): 149–163. doi: 10.1639/0007-2745-113.1.149 Engel JJ, Heinrichs J (2008) Studies of New Zealand hepaticae. 39. Dinckleria Trevis. an older name for Proskauera Heinrichs and J.J.Engel. Cryptogamie, Bryologie 29 (2): 193–194. Engel JJ, Kuwahara Y (1973) Metzgeria litoralis sp. nov. and Apometzgeria from southern South America. Bryologist 76 (2): 293–296. doi: 10.2307/3241332 Engel JJ, Schuster RM (1975) Austral hepaticae. III. Stolonophora, a new genus of Geocalycaceae. Fieldiana: Botany 36 (11): 111–124. Engel JJ, Schuster RM (1981) Austral hepaticae. XV. Brevianthaceae, fam. nov. and Brevianthus, gen. nov. from Tasmania. Phytologia 47 (4): 317–318. Engel JJ, Schuster RM (1983) Austral hepaticae. XVIII. Studies towards a revision of Telaranea subg. Neolepidozia (Lepidoziaceae). Fieldiana: Botany (n.ser.) 14: 1–7. 692 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Engel JJ, Schuster RM (1984) An overview and evaluation of the genera of Geocalycaceae subfamily Lophocoleoideae (Hepaticae). Nova Hedwigia 39: 385–463. Engel JJ, Schuster RM (1988) Studies of New Zealand hepaticae. 1-6. Brittonia 40 (2): 200–207. doi: 10.2307/2807005 Engel JJ, Schuster RM (2001) Austral hepaticae. 32. A revision of the genus Lepidozia (Hepaticae) for New Zealand. Fieldiana: Botany (n.ser.) 42: 1–107. Engel JJ, Smith Merrill GL (1994) Studies of New Zealand hepaticae. 8-13. Bazzania and Acromastigum. Bryologist 97 (3): 313–320. doi: 10.2307/3243464 Engel JJ, Smith Merrill GL (1995) Austral hepaticae. 23. New taxa and new combinations in Telaranea Spruce ex Schifn. (Lepidoziaceae). Phytologia 79 (3): 250–253. Engel JJ, Smith Merrill GL (1996) Studies of New Zealand hepaticae. 14-19. Kurzia and Lepicolea. Journal of the Hattori Botanical Laboratory 80: 217–231. Engel JJ, Smith Merrill GL (1997) Austral hepaticae. 22. he genus Balantiopsis in New Zealand, with observations on extraterritorial taxa and a phylogeny of Balantiopsis and the family Balantiopsaceae (Jungermanniales). Fieldiana: Botany (n.ser.) 37: 1–62. Engel JJ, Smith Merrill GL (1998) Austral hepaticae. 25. Krunodiplophyllum Grolle and a revision of the Australasian species of Diplophyllum (Dum.) Dum. (Scapaniaceae, Jungermanniales). Journal of the Hattori Botanical Laboratory 84: 241–283. Engel JJ, Smith Merrill GL (1999a) Austral hepaticae. 29. More new taxa and combinations in Telaranea (Lepidoziaceae) and a new name for Frullania caledonica (Schust.) Schust. (Frullaniaceae) from New Caledonia. Novon 9 (3): 339–344. doi: 10.5962/bhl.title.744 Engel JJ, Merrill GL (1999b) Austral hepaticae. 28. Plagiochila bazzanioides Engel & Merrill, a remarkable new species of Plagiochilaceae from New Zealand. Novon 9 (1): 29–31. doi: 10.5962/bhl.title.744 Engel JJ, Smith Merrill GL (2001) Austral hepaticae. 33. Paracromastigum succulentum, comb. nov. (Lepidoziaceae, Jungermanniales). Bryologist 104 (1): 151–153. doi: 10.1639/0007-2745(2001)104[0151:AHPSCN]2.0.CO;2 Engel JJ, Smith Merrill GL (2004) Austral hepaticae. 35. A taxonomic and phylogenetic study of Telaranea (Lepidoziaceae), with a monograph of the genus in temperate Australasia and commentary on extra-Australasian taxa. Fieldiana: Botany (n.ser.) 44: 1–265. doi: 10.3158/0015-0746(2004)44[1:AHATAP]2.0.CO;2 Engel JJ, Smith Merrill GL (2009) Austral hepaticae. 46. he identity of Plagiochila retrospectans (Nees ex Spreng.) Lindenb. and P. fuscella (Hook.f. & Taylor) Gottsche, Lindenb. & Nees, two misunderstood Australasian species. Nova Hedwigia 89 (3/4): 287–301. doi: 10.1127/0029-5035/2009/0089-0287 Engel JJ, Smith Merrill GL (2010) Studies on New Zealand hepaticae. 39-55. More new taxa, combinations, typiications and synonymy in Plagiochila from New Zealand (Plagiochilaceae). Nova Hedwigia 91 (3/4): 501–517. doi: 10.1127/0029-5035/2010/00910501 Engel JJ, Smith Merrill GL (2013) Austral hepaticae. 49. New section names and synonymy in Plagiochila, with reference to New Zealand species [Plagiochilaceae (Jörg.) K. Müll.]. Nova Hedwigia 96 (3/4): 399–408. doi: 10.1127/0029-5035/2013/0085 World checklist of hornworts and liverworts 693 Engel JJ, Bardat J, houvenot L (2012) Studies on Lophocoleaceae. XXI. Otoscyphus J.J.Engel, Bardat et houvenot, a new liverwort genus from New Caledonia with an unusual morphology. Cryptogamie, Bryologie 33 (3): 279–289. doi: 10.7872/cryb.v33.iss3.2012.279 Engel JJ, Váňa J, Söderström L, Hagborg A, von Konrat M (2014) Notes on Early Land Plants Today. 64. Nomenclatural notes in Balantiopsaceae. Phytotaxa 183 (4): 299–300. doi: 10.11646/phytotaxa.183.4.13 Evans AW (1891) A provisional list of the hepaticae of the Hawaiian Islands. Transactions of the Connecticut Academy of Arts and Sciences 8 (15): 253–261. Evans AW (1892a) Hepaticae. In: Millspaugh CF (Ed.) Preliminary catalogue of the lora of West Virginia. Moses W. Donally, Charleston, West Va., 495–498. Evans AW (1892b) List of liverworts from Southern Patagonia. Contributions from the United States National Herbarium 1 (5): 140–142. Evans AW (1893) An arrangement of the genera of hepaticae. Transactions of the Connecticut Academy of Arts and Sciences 8 (16): 262–280. Evans AW (1896) Notes on the North American species of Plagiochila. Botanical Gazette 21 (4): 185–194. doi: 10.1086/327330 Evans AW (1898) An enumeration of the hepaticae collected by John B. Hatcher in southern Patagonia. Bulletin of the Torrey Botanical Club 25 (8): 407–431. doi: 10.2307/2477911 Evans AW (1899) A revision of the North American species of Frullania, a genus of Hepaticae. Transactions of the Connecticut Academy of Arts and Sciences 10 (1): 1–39. Evans AW (1900a) he Hawaiian hepaticae of the tribe Jubuloideae. Transactions of the Connecticut Academy of Arts and Sciences 10 (8): 387–462. Evans AW (1900b) A new genus of hepaticae from the Hawaiian Islands. Bulletin of the Torrey Botanical Club 27 (3): 97–104. doi: 10.2307/2478313 Evans AW (1900c) Notes on hepaticae collected in Alaska. Proceedings of the Washington Academy of Sciences 2: 287–314. Evans AW (1902a) he Lejeuneae of the United States and Canada. Memoirs of the Torrey Botanical Club 8 (2): 113–183. doi: 10.5962/bhl.title.97556 Evans AW (1902b) A new hepatic from the eastern United States. Botanical Gazette 34 (5): 372–375. doi: 10.1086/328299 Evans AW (1902c) Hepaticae of Puerto Rico. I. he species of Leptolejeunea including an account of their vegetative reproduction. Bulletin of the Torrey Botanical Club 29 (8): 496–510. doi: 10.2307/2478871 Evans AW (1903a) Yukon hepaticae. Ottawa Naturalist 17: 13–24. Evans AW (1903b) Hepaticae of Puerto Rico. II. Drepanolejeunea. Bulletin of the Torrey Botanical Club 30 (1): 19–41. doi: 10.2307/2478645 Evans AW (1903c) Hepaticae of Puerto Rico. III. Harpalejeunea, Cyrtolejeunea, Euosmolejeunea and Trachylejeunea. Bulletin of the Torrey Botanical Club 30 (10): 544–563. doi: 10.2307/2478516 Evans AW (1904a) Hepaticae of Puerto Rico. IV. Odontolejeunea, Cyclolejeunea and Prionolejeunea. Bulletin of the Torrey Botanical Club 31 (4): 183–226. doi: 10.2307/2478687 Evans AW (1904b) Notes on New England Hepaticae — II (cont.). Rhodora 6 (69): 185–190. 694 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Evans AW (1905a) Hepaticae of Puerto Rico. V. Ceratolejeunea. Bulletin of the Torrey Botanical Club 32 (6): 273–290. doi: 10.2307/2478810 Evans AW (1905b) New or noteworthy hepaticae from Florida. Bulletin of the Torrey Botanical Club 32 (4): 179–192. doi: 10.2307/2478831 Evans AW (1905c) A remarkable Ptilidium from Japan. Revue Bryologique 32 (4): 57–60. Evans AW (1906a) Hepaticae of Puerto Rico. VI. Cheilolejeunea, Rectolejeunea, Cystolejeunea, and Pycnolejeunea. Bulletin of the Torrey Botanical Club 33 (1): 1–25. doi: 10.2307/2478618 Evans AW (1906b) Notes on Japanese hepaticae. Proceedings of the Washington Academy of Sciences 8: 141–166. Evans AW (1906c) he hepaticae of Bermuda. Bulletin of the Torrey Botanical Club 33 (3): 129–135. doi: 10.2307/2478823 Evans AW (1907a) Hepaticae of Puerto Rico. VIII. Symbiezidium, Marchesinia, Mastigolejeunea, Caudalejeunea and Bryopteris. Bulletin of the Torrey Botanical Club 34 (11): 533–568. doi: 10.2307/2479269 Evans AW (1907b) Hepaticae of Puerto Rico. VII. Stictolejeunea, Neurolejeunea, Omphalanthus, and Lopholejeunea. Bulletin of the Torrey Botanical Club 34 (1): 1–34. doi: 10.2307/2479163 Evans AW (1908a) Hepaticae of Puerto Rico. IX. Brachiolejeunea, Ptychocoleus, Archilejeunea, Leucolejeunea, and Anoplolejeunea. Bulletin of the Torrey Botanical Club 35 (4): 155–179. doi: 10.2307/2478953 Evans AW (1908b) New West Indian Lejeuneae. Bulletin of the Torrey Botanical Club 35 (8): 371–389. doi: 10.2307/2479284 Evans AW (1909) Notes on New England hepaticae, – VII. Rhodora 11 (130): 185–195. Evans AW (1910) Vegetative reproduction in Metzgeria. Annals of Botany. Oxford 24 (2): 271–303. Evans AW (1911) Hepaticae of Puerto Rico. X. Cololejeunea, Leptocolea and Aphanolejeunea. Bulletin of the Torrey Botanical Club 38 (6): 251–286. doi: 10.2307/2479100 Evans AW (1912a) Notes on North American hepaticae. III. Bryologist 15 (4): 54–63. doi: 10.2307/3237690 Evans AW (1912b) Notes on New England hepaticae, – IX. Rhodora 14 (157): 1–18. Evans AW (1912c) Hepaticae of Puerto Rico. XI. Diplasiolejeunea. Bulletin of the Torrey Botanical Club 39 (5): 209–225. doi: 10.2307/2479376 Evans AW (1912d) New West Indian Lejeuneae – II. Bulletin of the Torrey Botanical Club 39 (12): 603–611. doi: 10.2307/2479587 Evans AW (1913) Notes on North American hepaticae. IV. Bryologist 16 (4): 49–55. doi: 10.2307/3238381 Evans AW (1914a) Report on the hepaticae of Alaska. Bulletin of the Torrey Botanical Club 41 (12): 577–616. doi: 10.2307/2479541 Evans AW (1914b) Notes on New England hepaticae, – XI. Rhodora 16 (184): 62–76. Evans AW (1914c) Hepaticae: Yale Peruvian Expedition of 1911. Transactions of the Connecticut Academy of Arts and Sciences 18 (5): 291–345. World checklist of hornworts and liverworts 695 Evans AW (1915) he genus Plagiochasma and its North American species. Bulletin of the Torrey Botanical Club 42 (5): 259–308. doi: 10.2307/2479766 Evans AW (1916) A new species of Metzgeria from the Galapagos Islands. Torreya 16 (3): 67–70. Evans AW (1917a) he American species of Marchantia. Transactions of the Connecticut Academy of Arts and Sciences 21 (3): 205–313. Evans AW (1917b) Notes on North American hepaticae. VII. Bryologist 20 (2): 17–28. doi: 10.2307/3238917 Evans AW (1917c) Notes on the genus Herberta, with a revision of the species known from Europe, Canada and the United States. Bulletin of the Torrey Botanical Club 44 (4): 191–222. doi: 10.2307/2479530 Evans AW (1917d) A new Lejeunea from Bermuda and the West Indies. Bulletin of the Torrey Botanical Club 44 (11): 525–528. doi: 10.2307/2479506 Evans AW (1918) Noteworthy Lejeuneae from Florida. American Journal of Botany 5 (3): 131–150. doi: 10.2307/2435178 Evans AW (1919a) Notes on New England hepaticae, – XV. Rhodora 21 (249): 149–169. Evans AW (1919b) hree South American species of Asterella. Bulletin of the Torrey Botanical Club 46 (12): 469–480. doi: 10.2307/2480391 Evans AW (1920) he North American species of Asterella. Contributions from the United States National Herbarium 20: 247–312. Evans AW (1921a) he genus Riccardia in Chile. Transactions of the Connecticut Academy of Arts and Sciences 25 (2): 93–209. Evans AW (1921b) Taxilejeunea pterogonia and certain allied species. Bulletin of the Torrey Botanical Club 48 (4): 107–136. doi: 10.2307/2480341 Evans AW (1922) Notes on North American hepaticae. IX. Bryologist 25 (2): 25–33. doi: 10.2307/3237567 Evans AW (1923a) he Chilean species of Metzgeria. Proceedings of the American Academy of Arts and Sciences 58 (7): 271–324. doi: 10.2307/20025995 Evans AW (1923b) Notes on North American hepaticae. X. Bryologist 26 (6): 55–67. doi: 10.2307/3238024 Evans AW (1925) he lobate species of Symphyogyna. Transactions of the Connecticut Academy of Arts and Sciences 27 (1): 1–50. Evans AW (1927) A further study of the American species of Symphyogyna. Transactions of the Connecticut Academy of Arts and Sciences 28 (6): 301–353. Evans AW (1930a) he thallose hepaticae of the Juan Fernández Islands. In: Skottsberg C (Ed.) he Natural History of Juan Fernández and Easter Island. Botany 2. Almquist & Wiksell, Stockholm, 551–586. doi: 10.5962/bhl.title.25662 Evans AW (1930b) Two species of Lejeunea from Chile. Annales Bryologici 3: 83–88. Evans AW (1932a) A new species of Plagiochasma from Texas. American Journal of Botany 19 (7): 627–631. doi: 10.2307/2436254 Evans AW (1932b) Some representative species of Bazzania from Sumatra. Papers of the Michigan Academy of Science, Arts and Letters 17: 69–118. Evans AW (1934) A revision of the genus Acromastigum. Annales Bryologici, suppl. 3: 1–178. doi: 10.1007/978-94-015-3499-4 696 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Evans AW (1938a) he invalidity of the genus Grimaldia of Raddi. Chronica Botanica 4: 223–225. Evans AW (1938b) A new species of Chiloscyphus from Utah. Bryologist 41 (3): 50–57. doi: 10.2307/3238980 Evans AW (1938c) Notes on the genus Cololejeunea. Bryologist 41 (4): 71–82. doi: 10.2307/3239339 Fang Y-M, Enroth J, Koponen T, Piippo S (1998) he bryophytes of Jianxi province, China: An annotated checklist. Hikobia 12: 343–363. Feldberg K, Heinrichs J (2006) A taxonomic revision of Herbertus (Jungermanniidae: Herbertaceae) in the Neotropics based on nuclear and chloroplast DNA and morphology. Botanical Journal of the Linnean Society 151: 309–332. doi: 10.1111/j.1095-8339.2006.00534.x Feldberg K, Hentschel J, Bombosch A, Long DG, Váňa J, Heinrichs J (2009) Transfer of Gottschelia grollei, G. patoniae and Scaphophyllum speciosum to Solenostoma based on chloroplast DNA rbcL sequences. Plant Systematics and Evolution 280 (3/4): 243–250. doi: 10.1007/s00606-009-0187-3 Feldberg K, Váňa J, Long DG, Shaw AJ, Hentschel J, Heinrichs J (2010a) A phylogeny of Adelanthaceae (Jungermanniales, Marchantiophyta) based on nuclear and chloroplast DNA markers, with comments on classiication, cryptic speciation and biogeography. Molecular Phylogenetics and Evolution 55 (1): 293–304. doi: 10.1016/j.ympev.2009.11.009 Feldberg K, Váňa J, Hentschel J, Heinrichs J (2010b) Currently accepted species and new combinations in Jamesonielloideae (Adelanthaceae, Jungermanniales). Cryptogamie, Bryologie 31 (2): 141–146. Feldberg K, Váňa J, Schulze C, Bombosch A, Heinrichs J (2011) Morphologically similar but genetically distinct: on the diferentiation of Syzygiella concreta and S. perfoliata (Adelanthaceae subfam. Jamesonielloideae). Bryologist 114 (4): 686–695. doi: 10.1639/0007-2745-114.4.686 Feldberg K, Heinrichs J, Schmidt AR, Váňa J, Schneider H (2013) Exploring the impact of fossil constraints on the divergence time estimates of derived liverworts. Plant Systematics and Evolution 299 (3): 585–681. doi: 10.1007/s00606-012-0745-y Fischer E (1993) Taxonomic results of the BRYOTROP expedition to Zaire and Rwanda. 8. Riccia vulcanicola E. Fischer (subgenus Ricciella, sectio Cavernosae), a new species from the Virunga volcanoes, Rwanda. Tropical Bryology 8: 69–74. Fischer E, Vanderpoorten A (2010) New records for the liverwort lora of Gabon, with a description of Ceratolejeunea kuerschneri, sp. nov. (Lejeuneaceae, Jungermanniopsida). Beihefte zur Nova Hedwigia 138: 85–97. Forrest LL, Davis EC, Long DG, Crandall-Stotler BJ, Clark A, Hollingsworth ML (2006) Unravelling the evolutionary history of the liverworts (Marchantiophyta): multiple taxa, genomes and analyses. Bryologist 109 (3): 303–334. doi: 10.1639/0007-2745(2006)109[303:UT EHOT]2.0.CO;2 Forrest LL, Salazar N, Gudiño JA, Korpelainen H, Long DG (2011) Molecular and morphological evidence for distinct species in Dumortiera (Dumortieraceae). Bryologist 114 (1): 102–115. doi: 10.1639/0007-2745-114.1.102 Forster JR, Forster G (1776) Characteres generum plantarum, ed. 2. White, Cadell et Elmsly, London, 150 pp. doi: 10.5962/bhl.title.4448 World checklist of hornworts and liverworts 697 Frahm J-P (1990) Bryophyte phytomass in tropical ecosystems. Journal of the Linnean Society. Botany 104 (1/3): 23–33. doi: 10.1111/j.1095-8339.1990.tb02209.x Frahm J-P (2006) Frullania tamarisci var. azorica (Jubulaceae, Marchantiopsida), a new taxon from the Azores. Tropical Bryology 27: 101–105. Frahm J-P, Eggers J (2001) Lexikon deutschsprachiger Bryologen. Books on Demand, Norderstedt, 672 pp. Frahm J-P, Frey W, Kürschner H, Menzel M (1990) Mosses and liverworts of Mt. Kinabalu. Sabah Parks Publication 12: 1–91. Frey W, Kürschner H (1993) Targionia hypophylla L. ssp. linealis (Marchantiidae, Hepaticae), eine neue Unterart aus Saudi Arabien sowie weitere Neufunde. Studien an arabischen Bryophyten 17. Nova Hedwigia 57 (1/2): 127–133. Frey W, Stech M (2005a) A morpho-molecular classiication of the Anthocerotophyta (hornworts). Nova Hedwigia 80 (3/4): 541–545. doi: 10.1127/0029-5035/2005/0080-0541 Frey W, Stech M (2005b) A morpho-molecular classiication of the liverworts (Hepaticophytina, Bryophyta). Nova Hedwigia 81 (1/2): 55–78. doi: 10.1127/0029-5035/2005/0081-0055 Frey W, Stech M (2008) New suprageneric taxa of liverworts (Marchantiophyta) and mosses (Bryophyta). Nova Hedwigia 87 (1/2): 261–267. doi: 10.1127/0029-5035/2008/00870261 Frisvoll AA, Moen A (1980) Lophozia borealis sp. nov., a rich fen hepatic from Fennoscandia. Lindbergia 6 (2): 137–146. Frye TC, Clark L (1937) Hepaticae of North America. University of Washington Publications in Biology 6 (1): 1–161. Frye TC, Clark L (1946) Hepaticae of North America, part IV. University of Washington Publications in Biology 6 (4): 565–733. Fulford M (1941) Studies on American hepaticae I. Revision of the genus hysananthus. Bulletin of the Torrey Botanical Club 68 (1): 32–42. doi: 10.2307/2481253 Fulford M (1946) he genus Bazzania in Central and South America. Annales Cryptogamici et Phytopathologici 3: 1–175. Fulford M (1959a) Studies on American hepaticae. 9-11. A supplement to “he genus Bazzania in Central and South America”, II. Tridentatae (3-5). Bulletin of the Torrey Botanical Club 86 (6): 394–412. doi: 10.2307/2482645 Fulford M (1959b) Studies on American hepaticae. 7-8. A supplement to “he genus Bazzania in Central and South America.” Part 1. Introduction and the subgenus Bidentatae. Bulletin of the Torrey Botanical Club 86 (5): 308–341. doi: 10.2307/2482503 Fulford M (1960) Bazzania chimantensis, a new species from Venezuela. Bryologist 63 (2): 88–93. doi: 10.2307/3240878 Fulford M (1962a) Segregate genera of the Lepidozia complex (Hepaticae). Part 3. Microlepidozia Jörg. and Micrisophylla gen. nov. Brittonia 14 (1): 121–136. doi: 10.2307/2805325 Fulford M (1962b) Additional notes on the Vetaformaceae (Hepaticae). Nova Hedwigia 4 (1/2): 81–85. Fulford M (1963) Segregate genera of the Lepidozia complex (Hepaticae). Part 4. Telaranea and a review of the Lepidoziaceae. Brittonia 15 (1): 65–85. doi: 10.2307/2805041 698 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Fulford M (1966) Manual of the leafy hepaticae of Latin America II. Memoirs of the New York Botanical Garden 11 (2): 173–276. Fulford M (1967) Hepaticae. In: Steyermark JA (Ed.) Flora del Auyan-tepui. Acta Botánica Venezuelica 2 (5/8): 5–370. Fulford M (1968) Manual of the leafy hepaticae of Latin America III. Memoirs of the New York Botanical Garden 11 (3): 277–392. Fulford M (1972) Hepaticae. In: Steyermark JA (Ed.) Maguire B, he lora of the Meseta del Cerro Jaua. Memoirs of the New York Botanical Garden 23: 833–892. Fulford M (1976) Manual of the leafy hepaticae of Latin America IV. Memoirs of the New York Botanical Garden 11 (4): 395–535. Fulford M, Hatcher RE (1958) Triandrophyllum, a new genus of leafy hepaticae. Bryologist 61 (4): 276–285. doi: 10.2307/3240159 Fulford M, Hatcher RE (1961) he genus Triandrophyllum – some nomenclatural changes. Bryologist 64 (4): 348–351. doi: 10.2307/3240856 Fulford M, Sharp AJ (1990) he leafy hepaticae of Mexico: one hundred and twenty-seven years after C. M. Gottsche. Memoirs of the New York Botanical Garden 63: 1–86. Fulford M, Taylor J (1959a) he segregate genera of the Lepidozia complex (Hepaticae). Part 1. Sprucella Steph. and Neolepidozia gen. nov. Brittonia 11 (2): 77–85. doi: 10.2307/2805175 Fulford M, Taylor J (1959b) Two new families of leafy hepaticae: Vetaformaceae and Pseudolepicoleaceae from southern South America. Nova Hedwigia 1 (3/4): 405–422. Fulford M, Taylor J (1961) Segregate genera of the Lepidozia complex (Hepaticae). Part 2. Two new genera, Bonneria and Paracromastigum. Brittonia 13 (4): 334–339. doi: 10.2307/2805411 Furuki T (1991) A taxonomical revision of the Aneuraceae (Hepaticae) of Japan. Journal of the Hattori Botanical Laboratory 70: 293–397. Furuki T (1994a) Taxonomic studies of Asiatic species of Aneuraceae (Hepaticae). I. Riccardia subg. Corioneura Furuki. Journal of the Hattori Botanical Laboratory 75: 257–261. Furuki T (1994b) Aneura marianensis sp. nov. (Hepaticae) of the northern Mariana Islands. Bryologist 97 (1): 87–88. doi: 10.2307/3243357 Furuki T (1997) Taxonomic studies of Asiatic species of Aneuraceae (Hepaticae). V. Riccardia planilora (Steph.) Hatt. var. aequatorialis Furuki var. nov. Natural History Research 4 (2): 77–79. Furuki T (1998) Taxonomic studies of Asiatic species of Aneuraceae (Hepaticae). VI. Riccardia fruticosa (Steph.) Furuki, comb. nov., described from New Guinea and its related species. Natural History Research 5 (1): 1–10. Furuki T (1999) Taxonomic studies of Asiatic Species of Aneuraceae (Hepaticae). VII. Riccardia grollei Furuki, sp. nov. from southeast Asia. Haussknechtia, Beiheft 9: 139–142. Furuki T (2001) Lethocolea naruto-toganensis, a new hepatic from swamps of Japan. Bryologist 104 (2): 306–309. doi: 10.1639/0007-2745(2001)104[0306:LNTANH]2.0.CO;2 Furuki T (2006a) Taxonomical studies of the family Aneuraceae (Hepaticae) based on the Philippine collections made by Dr. and Mrs. A. J. Sharp and Dr. Z. Iwatsuki. Journal of the Hattori Botanical Laboratory 100: 89–99. World checklist of hornworts and liverworts 699 Furuki T (2006b) A new species of Nardia (Jungermanniaceae, Hepaticae) from Japan. Bryological Research 9 (3): 73–77. Furuki T (2007) Radula fujitae Furuki, sp. nov. (Radulaceae, Hepaticae) from Okinawa Island, southern Japan. Bryological Research 9 (5): 143–146. Furuki T, Higuchi M (2006) A new species of Exormotheca (Exormothecaceae, Hepaticae) from China. Cryptogamie, Bryologie 27 (1): 97–102. Furuki T, Iwatsuki Z (1989) Mizutania riccardioides, gen. et sp. nov. (Mizutaniaceae, fam. nov.), a unique liverwort from Tropical Asia. Journal of the Hattori Botanical Laboratory 67: 291–296. Furuki T, Long DG (1994) Aneura crateriformis, a new liverwort species from the East Himalaya and China. Journal of Bryology 18 (2): 281–286. doi: 10.1179/jbr.1994.18.2.281 Furuki T, Long DG (2007) Lobatiriccardia yunnanensis, sp. nov. (Metzgeriales, Aneuraceae) from Yunnan, China. Journal of Bryology 29 (3): 161–164. doi: 10.1179/174328207X227429 Furuki T, Yamada K (1986) he genus Radula (Hepaticae) of the Bonin and Volcano Islands. Journal of Japanese Botany 61 (10): 311–314. Furuki T, Yong K-T, Mohamed AMH (2013) Riccardia deguchii Furuki & K.T.Yong (Marchantiophyta, Aneuraceae), sp. nov. from Malaysia. Hikobia 16 (3): 285–288. Fuselier L, Davison PG, Clements M, Shaw B, Devos N, Heinrichs J, Hentschel J, Sabovljević M, Szövényi P, Schuette S, Hofbauer W, Shaw AJ (2009) Phylogeographic analyses reveal distinct lineages of the liverworts Metzgeria furcata (L.) Dumort. and Metzgeria conjugata Lindb. (Metzgeriaceae) in Europe and North America. Journal of the Linnean Society. Biology 98 (4): 745–756. doi: 10.1111/j.1095-8312.2009.01319.x Fuselier LC, Shaw B, Engel JJ, von Konrat M, Costa DP, Devos N, Shaw AJ (2011) he status and phylogeography of the liverwort genus Apometzgeria Kuwah. (Metzgeriaceae). Bryologist 114 (1): 92–101. doi: 10.1639/0007-2745-114.1.92 Gao C (2003) Flora bryophytorum Sinicorum 9. Takakiales, Calobryales, Jungermanniales. Science Press, Beijing, 323 pp. Gao C, Bai X-L (2001) A synoptic revision of family Jungermanniaceae (Hepaticae) in China including some taxa nova. Philippine Scientist 38: 111–170. Gao C, Bai X-L (2002) Lepidozia suyungii (Lepidoziaceae, Hepaticae), a new species from southwestern China, with discussion of the species of Lepidozia in China. Journal of the Hattori Botanical Laboratory 92: 191–197. Gao C, Chang KC (1978) Species novae Ricciacearum Chinae boreali-orientalis. Acta Phytotaxonomica Sinica 16 (4): 113–118. Gao C, Chang KC (1981) Flora hepaticarum Chinae boreali-orientalis. Science Press, Beijing, 220 pp. Gao C, Chang K (1982) Genus nova Bryophytarum [sic]. Bulletin of Botanical Research. Harbin 2 (4): 113–121. Gao C, Wu Y-H (2004) On seven species of the Schistochilaceae (Hepaticae) in China, including one new species and one new combination. Journal of the Hattori Botanical Laboratory 95: 263–270. 700 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Gao C, Wu Y-H (2005) Radula stellatogemmipara (Radulaceae, Hepaticae), a new species from Fujian and Guangxi, China. Nova Hedwigia 80 (1/2): 237–240. doi: 10.1127/00295035/2005/0080-0237 Gao C, Chang K-C, Cao T (1981) Taxa nova bryophytarum Tibeticarum. Acta Botanica Yunnanica 3 (4): 389–399. Gao C, Cao T, Lai MJ (2001) he genus Saccogynidium (Geocalycaceae, Hepaticae) in China. Bryologist 104 (1): 126–129. doi: 10.1639/0007-2745(2001)104[0126:TGSGHI]2.0.CO;2 Gao C, Cao T, Sun J (2002) he genus Isotachis (Hepaticae, Balantiopsidaceae) in China. Bryologist 105 (4): 693–698. doi: 10.1639/0007-2745(2002)105[0693:TGIHBI]2.0.CO;2 Gao C, Wu Y-H, Grolle R (2003) Jungermannia cheniana, a new liverwort with 2-3-stratose leaves from Yunnan, China. Nova Hedwigia 77 (1/2): 189–193. doi: 10.1127/00295035/2003/0077-0189 Gao C, Cao T, Wu Y, Yu J, Chen Y (2004) A new species and three new records of Heteroscyphus (Jungermanniopsida: Geocalycaceae) to China. Journal of Bryology 26 (2): 97–102. doi: 10.1179/037366804225021047 Garcia C, Sérgio C, Villarreal JC, Sim-Sim M, Lara F (2012) he hornworts Dendroceros Nees and Megaceros Campb. in São Tomé e Príncipe (Africa, Gulf of Guinea) with the description of Dendroceros paivae sp. nov. Cryptogamie, Bryologie 33 (1): 3–21. doi: 10.7872/ cryb.v33.iss1.2012.003 Garside S (1958) Studies in South African Ricciaceae. III. A new species of Oxymitra. Journal of South African Botany 24: 83–87. Gaudichaud C (1827) Voyage autour du monde entrepris par ordre du Roi, exécuté sur les corvettes l’Uranie et la Physicienne. Botanique, fasc. 4. Arthus Bertrand, Paris, 121–160. doi: 10.5962/bhl.title.15862 Geissler P, Gradstein SR (1994) On the identity of Phragmicoma lehmanniana Nees and other species described in the genus Phragmicoma (Studies in Lejeuneaceae subfam. Ptychanthoideae. 23). Journal of the Hattori Botanical Laboratory 75: 201–209. Gepp A (1890) Musci, Hepaticae (in: Ridley HN, Notes on the Botany of Fernando Noronha). Journal of the Linnean Society. Botany 27 (181): 74–75. doi: 10.1111/j.1095-8339.1890. tb00800.x Gepp A (1895a) Additional notes on Mr. W. R. Elliott’s hepaticae. Journal of Botany, British and Foreign 33: 82–84. Gepp A (1895b) Hepaticae Elliottianae, insulis Antillanis St Vincentii et Dominica a clar. W. R. Elliott, annis 1891-92, lectae, Richard Spruce determinatae. Journal of the Linnean Society. Botany 30 (210): 331–372. doi: 10.1111/j.1095-8339.1894.tb02416.x Gerola FM (1947) Epatiche dell’Abissinia meridionale. Lavori di Botanica, Istituto di Botanica e di Fisiologia Vegetale dell’Università de Padova 12: 471–485. Gerson U (1982) Bryophytes and invertebrates. In: Smith AJE (Ed.) Bryophyte Ecology. Chapman & Hall, London, 291–332. doi: 10.1007/978-94-009-5891-3_9 Gibbs LS (1909) A contribution to the montane lora of Fiji (including Cryptogams), with ecological notes. Journal of the Linnean Society. Botany 39 (270): 130–212. doi: 10.1111/ j.1095-8339.1909.tb01192.x World checklist of hornworts and liverworts 701 Gignac LD (2001) Bryophytes as indicators of climate change. Bryologist 104 (3): 410–420. doi: 10.1639/0007-2745(2001)104[0410:BAIOCC]2.0.CO;2 Gillett JM, Jovet-Ast S (1957) Deux Riccia de l’Aïr (Territoire du Niger). Revue Bryologique et Lichénologique 26 (1/2): 62–66. Giordano S, Sorbo S, Adamo P, Basile A, Spagnuolo V, Cobianchi RC (2004) Biodiversity and trace element content of epiphytic bryophytes in urban and extraurban sites of southern Italy. Plant Ecology 170 (1): 1–14. doi: 10.1023/B:VEGE.0000019025.36121.5d Glenny DS (1996) Nephelolejeunea papillosa, a new liverwort species from New Zealand, with notes on the distribution of Kymatolejeunea bartlettii Grolle. New Zealand Journal of Botany 34 (2): 195–198. doi: 10.1080/0028825X.1996.10410683 Glenny DS, Bartlett R (2007) A new Bazzania species (Lepidoziaceae) from Stockton Plateau, Nelson, South Island, New Zealand. Fieldiana: Botany (n.ser.) 47: 175–180. doi: 10.3158/0015-0746-47.1.175 Glenny D, Engel JJ (2013) A new species of Clasmatocolea (Marchantiophyta: Lophocoleaceae), and a new record of Tasmanian species, both from New Zealand. New Zealand Journal of Botany 51 (1): 22–30. doi: 10.1080/0028825X.2012.736390 Glenny DS, Braggins JE, Schuster RM (1997) Zoopsis nitida (Hepaticae: Lepidoziaceae), a new species from New Zealand. Journal of Bryology 19 (4): 775–780. doi: 10.1179/ jbr.1997.19.4.775 Glenny DS, Engel JJ, He-Nygrén X-L (2009) he systematic identity of Chiloscyphus trichocoleoides, a new liverwort species from New Zealand, uncovered by morphological and molecular evidence. Journal of Bryology 31 (2): 93–105. doi: 10.1179/174328209X427524 Godfrey JD (1976) Schoieldia, a new hepaticae from the Paciic northwest. Bryologist 79 (3): 314–320. doi: 10.2307/3242370 Godfrey JD, Godfrey GA (1978) Scapania hians in Shensi, China and British Columbia, Canada. Bryologist 81 (3): 357–367. doi: 10.2307/3242238 Godfrey JD, Godfrey GA (1979) Jungermannia schusterana, a new hepatic from the Paciic coast of North America. Journal of the Hattori Botanical Laboratory 46: 109–117. Godfrey JD, Godfrey GA (1980) Frullania hattoriana, a new hepatic from British Columbia, Canada. Journal of the Hattori Botanical Laboratory 48: 321–327. Goebel KI (1888) Morphologische und biologische Studien. I. Ueber epiphytische Farne und Muscineen. Annales du Jardin Botanique de Buitenzorg 7 (1): 1–73. Goebel KI (1889) Über die Jugendzustände der Planzen. Flora 72 (1): 1–45. Goebel KI (1890) Morphologische und biologische Studien. IV. Ueber Javanische Lebermoose. Annales du Jardin Botanique de Buitenzorg 9 (1): 1–40. Goebel KI (1891) Planzenbiologische Schilderungen, Zweiter Teil, Erste Lieferung. N. G. Elwert’sche Verlagsbuchhandlung, Marburg, 160 pp. doi: 10.5962/bhl.title.1665 Goebel KI (1893a) Archegoniatenstudien. 5. Die Blattbildung der Lebermoose und ihre biologische Bedeutung. Flora 77: 423–459. Goebel KI (1893b) Archegoniatenstudien. 3. Rudimentäre Lebermoose. Flora 77 (2): 82–103. Goebel KI (1906) Archegoniatenstudien. X. Beiträge zur Kenntnis australischer und neuseeländischer Bryophyten. Flora 96: 1–202. 702 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Goebel KI (1912) Archegoniatenstudien. XV. Die Homologie der Antheridien- und Archegonienhüllen bei den Lebermoosen. Flora 105: 53–70. Goebel KI (1915) Organographie der Planzen, ed. 2, vol. 2(1), Spezielle Organographie, Bryophyten. Gustav Fischer Verlag, Stuttgart, Jena, New York, 515–902. doi: 10.5962/bhl. title.20547 Goebel KI (1928) Morphologische und Biologische Studien XII-XV. Annales du Jardin Botanique de Buitenzorg 39: 1–232. Gola G (1907) Species novae in excelsis Ruwenzori in expeditione Ducis Aprutii Lectae. III: Hepaticae. Annali di Botanica. Roma 6 (2): 271–276. Gola G (1914a) Epatiche dell’Abissinia. Annali di Botanica. Roma 13 (1): 59–75. Gola G (1914b) Epatiche del Kashmir raccolte dalla Spedizione Piacenza. Atti della Reale Accademia delle Scienze di Torino. Classe di Scienze, Fisiche, Matematiche e Naturali 49: 757–761. Gola G (1916) Le epatiche della regione del Kenia. Memorie della reale accademia delle scienze di Torino (ser. 2) 65 (1): 1–11. Gola G (1920) Contributo alla conoscenza delle Epatiche del Katanga (Congo Belga). Nuovo Giornale Botanico Italiano (n.ser.) 27 (2/4): 244–250. Gola G (1922) Le epatiche raccolte dal Dott. G.B. de Gasperi nella Terra del Fuoco sudoccidentale. Nuovo Giornale Botanico Italiano (n.ser.) 29 (1/4): 162–173. Gottsche CM (1853) Muscorum hepaticorum species novae javanenses. Natuurkundig Tijdschrift voor Nederlandsch-Indië 4: 573–576. Gottsche CM (1856) Plantae Muellerianae: Hepaticae Australiae. Linnaea 28 (5): 547–561. Gottsche CM (1857) Pugillus novarum hepaticarum. Annales des Sciences Naturelles; Botanique (sér. 4) 8: 318–348. Gottsche CM (1858) Uebersicht und kritische Würdigung der seit dem Erscheinen der Synopsis Hepaticarum bekannt gewordenen Leistungen in der Hepaticologie. Botanische Zeitung. Berlin, Beilage 16: 1–54. Gottsche CM (1863) De Mexikanske Levermosser. Bianco Lunos Bogtrykkeri, Kjøbenhavn, 285 pp. Gottsche CM (1864) Hepaticae. In: Triana JJ, Planchon JE (Eds) Prodromus lorae novo-granatensis. Annales des Sciences Naturelles; Botanique (sér. 5) 1: 95–198. Gottsche CM (1867) Einige Bemerkungen zu hom. Jensen, Conspectus Hepaticarum Daniae eller Beskrivelse af de danske halvmosser. Hedwigia 6 (5): 65–77. Gottsche CM (1880) Musci hepaticae (Adans., Hedw.) sive lichenastri (Dill., Wallr.) Australiani. In: Müller F (Eds) Supplementum ad volumen undecimum Fragmentorum Phytographiae Australiae. Gobierni Coloniae Victoriae, Melbourne, 53–69. Gottsche CM (1882) Reliquiae Rutenbergianae. Lebermoose. Abhandlungen Herausgegeben vom Naturwissenschaftlichen Vereins zu Bremen 7: 338–365. Gottsche CM (1890) Die Lebermoose Süd-Georgiens. In: Neumayer GB (Eds) Die Internationale Polarforschung 1882-83. Die Deutschen Expeditionen und ihre Ergebnisse. II Beschreibende Naturwissenschaften. A. Asher & Co., Berlin, 449–454. Gottsche CM, Rabenhorst GL (1863) Hepaticae Europaeae. Die Lebermoose Europa’s unter Mitwirkung mehrerer namhafter Botaniker, decades 29-30. Dresden, no. 281–300. Gottsche CM, Rabenhorst GL (1867) Hepaticae Europaeae. Die Lebermoose Europa’s unter Mitwirkung mehrerer namhafter Botaniker, decades 38-39. Dresden, no. 371–390. World checklist of hornworts and liverworts 703 Gottsche CM, Rabenhorst GL (1868) Hepaticae Europaeae. Die Lebermoose Europa’s unter Mitwirkung mehrerer namhafter Botaniker, decades 42-44. Dresden, no. 411–440. Gottsche CM, Rabenhorst GL (1873a) Hepaticae Europaeae. Die Lebermoose Europa’s unter Mitwirkung mehrerer namhafter Botaniker, decades 58-59. Dresden, no. 571–590. Gottsche CM, Rabenhorst GL (1873b) Hepaticae Europaeae. Die Lebermoose Europa’s unter Mitwirkung mehrerer namhafter Botaniker, decades 56-57. Dresden, no. 551–570. Gottsche CM, Rabenhorst GL (1877) Hepaticae Europaeae. Die Lebermoose Europa’s unter Mitwirkung mehrerer namhafter Botaniker, decades 62-64. Dresden, no. 611–640. Gottsche CM, Lindenberg JBW, Nees CG (1843) Hepaticae. In: Meyen FJF (Ed.) Observationes botanicas. Novorum Actorum Academiae Caesareae Leopoldino-Carolinae Naturae Curiosorum 19 [suppl. 1]: 1–512. Gottsche CM, Lindenberg JBW, Nees CG (1844) Synopsis hepaticarum, fasc. 1. Meissner, Hamburg, 1–144. doi: 10.5962/bhl.title.15221 Gottsche CM, Lindenberg JBW, Nees CG (1845a) Synopsis hepaticarum, fasc. 2. Meissner, Hamburg, 145–304. doi: 10.5962/bhl.title.15221 Gottsche CM, Lindenberg JBW, Nees CG (1845b) Synopsis hepaticarum, fasc. 3. Meissner, Hamburg, 305–464. doi: 10.5962/bhl.title.15221 Gottsche CM, Lindenberg JBW, Nees CG (1846) Synopsis hepaticarum, fasc. 4. Meissner, Hamburg, 465–624. doi: 10.5962/bhl.title.15221 Gottsche CM, Lindenberg JBW, Nees CG (1847) Synopsis hepaticarum, fasc. 5. Meissner, Hamburg, 625–834. doi: 10.5962/bhl.title.15221 Gottsche CM, Rabenhorst GL (1872) Hepaticae Europaeae. Die Lebermoose Europa’s unter Mitwirkung mehrerer namhafter Botaniker, Decades 53-55. C. Heinrichs, Dresden, tab. 530–550. Gradstein SR (1974a) Studies on Lejeuneaceae subfam. Ptychanthoideae. I. Nomenclature and taxonomy of Ptychocoleus, Acrolejeunea and Schifneriolejeunea. Journal of the Hattori Botanical Laboratory 38: 327–336. Gradstein SR (1974b) Studies on Lejeuneaceae subfam. Ptychanthoideae (Hepaticae). II. Two remarkable species of Caudalejeunea: C. grolleana spec. nov. and C. cristiloba (Steph.) comb. nov. Acta Botanica Neerlandica 23 (3): 333–343. Gradstein SR (1975) A taxonomic monograph of the genus Acrolejeunea (Hepaticae), with an arrangement of the Genera of Ptychanthoideae. Bryophytorum Bibliotheca 4: 1–162. Gradstein SR (1977) Studies on Lejeuneaceae subfam. Ptychanthoideae (Hepaticae) IV. Verdoornianthus, a new genus from Amazonas, Brazil. Bryologist 80 (4): 606–611. doi: 10.2307/3242417 Gradstein SR (1985a) A guide to the holostipous Lejeuneaceae. Beihefte zur Nova Hedwigia 80: 13–29. Gradstein SR (1985b) A revision of the genus Stictolejeunea (Spruce) Schifn. Beihefte zur Nova Hedwigia 80: 195–220. Gradstein SR (1986) he genus Colura (Hepaticae) in the Galápagos Islands. Hikobia 9: 353–356. Gradstein SR (1989) A key to the hepaticae and anthocerotae of Puerto Rico and the Virgin Islands. Bryologist 92 (3): 329–348. doi: 10.2307/3243402 Gradstein SR (1991) Diversity and distribution of Asian Lejeuneaceae subfamily Ptychanthoideae. Tropical Bryology 4: 1–16. 704 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Gradstein SR (1992a) he genera hysananthus, Dendrolejeunea, and Fulfordianthus gen. nov. (Studies on Lejeuneaceae subfamily Ptychanthoideae. XXI). Bryologist 95 (1): 42–51. doi: 10.2307/3243784 Gradstein SR (1992b) What is Lejeunea trigona? (Studies on Lejeuneaceae subfam. Ptychanthoideae XXII). Contributions from the University of Michigan Herbarium 18: 99–103. Gradstein SR (1994) Lejeuneaceae: Ptychantheae, Brachiolejeuneae. Flora Neotropica, Monograph 62: 1–216. Gradstein SR (1997) Bromeliophila helenae, a new species of Lejeuneaceae from the Neotropics. Cryptogamie: Bryologie, Lichénologie 18 (3): 217–221. Gradstein SR (1999) Hepatics. In: Luteyn JL (Ed.) Páramos, a checklist of plant diversity, geographical distribution and botanical literature. Memoirs of the New York Botanical Garden 84: 1–278. Gradstein SR (2013a) Notes on Early Land Plants Today. 22. New combinations and new synonymy in Omphalanthus and Aureolejeunea (Lejeuneaceae, Marchantiophyta). Phytotaxa 76 (3): 45–47. doi: 10.11646/phytotaxa.76.3.10 Gradstein SR (2013b) Notes on Early Land Plants Today. 21. On Archilejeunea herminieri (Lejeuneaceae, Marchantiophyta). Phytotaxa 76 (3): 43–44. doi: 10.11646/phytotaxa.76.3.9 Gradstein SR (2013c) A classiication of Lejeuneaceae (Marchantiophyta) based on molecular and morphological evidence. Phytotaxa 100 (1): 6–20. doi: 10.11646/phytotaxa.100.1.2 Gradstein SR (2013d) Notes on Early Land Plants Today. 26. Miscellaneous synonyms in liverworts (Marchantiophyta). Phytotaxa 81 (1): 3–7. doi: 10.11646/phytotaxa.81.1.2 Gradstein SR, Benitez A (2014) Two new taxa of leafy liverworts (Marchantiophyta: Jungermanniidae) from Cerro Plateado, Cordillera del Cóndor, Ecuador. Nova Hedwigia 99 (1/2): 111–118. doi: 10.1127/0029-5035/2014/0181 Gradstein SR, Burghardt M (2008) A new species of Odontoschisma (Cephaloziaceae, Marchantiophyta) from South America. Fieldiana: Botany (n.ser.) 47: 193–198. doi: 10.3158/00150746-47.1.193 Gradstein SR, Buskes GM (1985) A revision of neotropical Archilejeunea (Spruce) Schifn. Beihefte zur Nova Hedwigia 80: 89–112. Gradstein SR, Costa DP (2003) he hepaticae and anthocerotae of Brazil. Memoirs of the New York Botanical Garden 87: 1–316. Gradstein SR, Florschütz-de Waard J (1989) Results of a botanical expedition to Mt. Roraima, Guyana. 1. Bryophytes. Tropical Bryology 1: 25–54. Gradstein SR, Geissler P (1997) Notes on the genus Leucolejeunea (Hepaticae). Cryptogamie: Bryologie, Lichénologie 18 (3): 177–182. Gradstein SR, Hekking WHA (1979) Studies on Colombian cryptogams. IV. A catalogue of the hepaticae of Colombia. Journal of the Hattori Botanical Laboratory 45: 93–144. Gradstein SR, Ilkiu-Borges AL (2009) Guide to the plants of central French Guiana, part 4. Liverworts and hornworts. Memoirs of the New York Botanical Garden 76 (4): 1–140. Gradstein SR, Ilkiu-Borges AL (2015) A taxonomic revision of the genus Odontoschisma (Marchantiophyta: Cephaloziaceae). Nova Hedwigia 100 (1/2): 15–100. doi: 10.1127/ nova_hedwigia/2014/0219 World checklist of hornworts and liverworts 705 Gradstein SR, Reiner-Drehwald ME (2007) he status of Neopotamolejeunea (Lejeuneaceae) and description of a new species from Ecuador and southern Brazil. Systematic Botany 32 (3): 487–492. doi: 10.1600/036364407782250571 Gradstein SR, Schäfer-Verwimp A (2011) A new species of Harpalejeunea (Spruce) Schifn. (Lejeuneaceae) from montane cloud forest of Colombia and Costa Rica. Cryptogamie, Bryologie 32 (2): 101–106. doi: 10.7872/cryb.v32.iss1.2011.101 Gradstein SR, Schäfer-Verwimp A (2012) A new species of Archilejeunea (Spruce) Schifn. (Lejeuneaceae) from Ecuador. Cryptogamie, Bryologie 33 (2): 107–112. doi: 10.7872/ cryb.v33.iss2.2012.107 Gradstein SR, Terken L (1981) Studies on Lejeuneaceae subfam. Ptychanthoideae VI. A revision of Schifneriolejeunea sect. Saccatae from Asia. Occasional Papers of the Farlow Herbarium of Cryptogamic Botany 16: 71–81. Gradstein SR, van Beek J (1985) A revision of the genus Symbiezidium Trevis. Beihefte zur Nova Hedwigia 80: 221–249. Gradstein SR, Váňa J (1999) On the taxonomy of Kymatocalyx and Stenorrhipis (Cephaloziellaceae). Haussknechtia, Beiheft 9: 155–170. Gradstein SR, Vanden Berghen C (1985) Schifneriolejeunea sect. Pappeanae en Afrique. Beihefte zur Nova Hedwigia 80: 173–193. Gradstein SR, Cleef AM, Fulford M (1977) Studies on Colombian cryptogams. IIA-C. Hepaticae – oil body structure and ecological distribution of selected species of tropical Andean Jungermanniales. Proceedings, Koninklijke Nederlandse Akademie van Wetenschappen. Series C, biological and medical sciences 80: 377–420. Gradstein SR, Klein R, Kraut L, Mues R, Spörle J, Becker H (1992) Phytochemical and morphological support for the existence of two species in Monoclea (Hepaticae). Plant Systematics and Evolution 180 (1/2): 115–135. doi: 10.1007/BF00940401 Gradstein SR, Grolle R, Schäfer-Verwimp A (1993) Two interesting species of Lejeuneaceae from Brazil. Journal of the Hattori Botanical Laboratory 74: 59–70. Gradstein SR, Lücking A, Morales MI, Dauphin G (1994) Additions to the hepatic lora of Costa Rica. Lindbergia 19 (2/3): 73–86. Gradstein SR, Churchill SP, Salazar Allen N (2001a) Guide to the bryophytes of tropical America. Memoirs of the New York Botanical Garden 86: 1–577. Gradstein SR, Griin D, Morales MI, Nadkarni N (2001b) Diversity and habitat diferentiation of mosses and liverworts in the cloud forest of Monteverde, Costa Rica. Caldasia 23 (1): 203–212. Gradstein SR, He X, Piippo S, Mizutani M (2002) Bryophyte lora of the Huon Peninsula, Papua New Guinea. LXVIII. Lejeuneaceae subfamily Ptychanthoideae (Hepaticae). Acta Botanica Fennica 174: 1–88. Gradstein SR, Meneses RI, Allain B (2003) Catalogue of the hepaticae and anthocerotae of Bolivia. Journal of the Hattori Botanical Laboratory 93: 1–67. Gradstein SR, Wilson R, Ilkiu-Borges AL, Heinrichs J (2006) Phylogenetic relationships and neotenic evolution of Metzgeriopsis (Lejeuneaceae) based on chloroplast DNA sequences and morphology. Botanical Journal of the Linnean Society 151 (3): 293–308. doi: 10.1111/j.1095-8339.2006.00531.x 706 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Gradstein SR, Ilkiu-Borges AL, Vanderpoorten A (2011) Habitat specialization triggers the evolution of unusual morphologies: the case of Cololejeunea stotleriana sp. nov. from Ecuador. Bryologist 114 (1): 9–22. doi: 10.1639/0007-2745-114.1.9 Gradstein SR, Laenen B, Frahm J-P, Schwarz U, Crandall-Stotler BJ, Engel JJ, von Konrat M, Stotler RE, Shaw B, Shaw AJ (2014a) On the taxonomic status of the enigmatic Phycolepidoziaceae (Marchantiophyta: Jungermanniales) with description of a new species, Phycolepidozia indica. Taxon 63 (3): 498–508. doi: 10.12705/633.17 Gradstein SR, Aranda SC, Vanderpoorten A (2014b) Notes on Early Land Plants Today. 47. Transfer of Iwatsukia to Odontoschisma (Cephaloziaceae, Marchantiophyta). Phytotaxa 162 (4): 232–233. doi: 10.11646/phytotaxa.162.4.6 Gray SF (1821) Natural arrangement of British plants. Vol. 1. Baldwin, Craddock, and Joy, London, 824 pp. doi: 10.5962/bhl.title.43804 Gray A (1846) Sullivant’s muscology. American Journal of Science and Arts (ser. 2) 1 (1): 70–81. Gray A (1848) A manual of the botany of the northern United States. John Chapman, London, 710 pp. doi: 10.5962/bhl.title.10392 Greville RK (1825) Descriptions of some new plants belonging to the order musci and hepaticae. Annals of the Lyceum of Natural History of New York 1 (2): 271–278. Griith W (1849) Notulae ad plantas asiaticas. Part II. On the higher cryptogamous plants. Bishop’s College Press, Calcutta, 257–628. doi: 10.5962/bhl.title.70352 Grolle R (1956) Revision der Clasmatocolea-Arten. Revue Bryologique et Lichénologique 25 (3/4): 288–303. Grolle R (1958) Über Jungermannia placophylla Taylor. Revue Bryologique et Lichénologique 27 (1/2): 52–54. Grolle R (1959a) Lophozia (Massula) patagonica Herz. & Grolle n. sp. Revue Bryologique et Lichénologique 28 (3/4): 343–345. Grolle R (1959b) Beitrag zur Kenntnis der afrikanischen Lophocoleen. Transactions of the British Bryological Society 3 (4): 582–598. doi: 10.1179/006813859804829171 Grolle R (1959c) Über Herpocladium issum Mitt. Ein Nachtrag zu “Was ist Pachyglossa?”. Revue Bryologique et Lichénologique 28 (3/4): 346–350. Grolle R (1960a) Notulae hepaticologicae I.-II.-III. Revue Bryologique et Lichénologique 29 (3/4): 207–211. Grolle R (1960b) Über Plagiochila monoica St., P. carnosa Herz. und P. lobata Kaal. und ihre Beziehungen zu Pedinophyllum. Nova Hedwigia 2: 287–291. Grolle R (1960c) Nachtrag zur “Revision der Clasmatocolea-Arten”. Revue Bryologique et Lichénologique 29 (1/2): 68–91. Grolle R (1960d) Über Saccogyna Dum. und Saccogynidium, eine neue Lebermoosgattung. Journal of the Hattori Botanical Laboratory 23: 41–67. Grolle R (1961a) Notulae hepaticologicae IV.-V.-VI. Revue Bryologique et Lichénologique 30 (1/2): 80–84. Grolle R (1961b) Tetracymbaliella, eine neue Lebermoosgattung. Nova Hedwigia 3 (1): 47–53. Grolle R (1962a) Monographie der Lebermoosgattung Leptoscyphus Mitt. Nova Acta Leopoldina (n.ser.) 25 (161): 1–143. World checklist of hornworts and liverworts 707 Grolle R (1962b) Goebelobryum, eine neue marsupiale Lebermoosgattung. Journal of the Hattori Botanical Laboratory 25: 135–144. Grolle R (1962c) Eine bemerkenswerte neue Lophozia aus Neuseeland. Revue Bryologique et Lichénologique 31 (3/4): 152–156. Grolle R (1963a) Zwei Gattungen der Lophoziaceae neu für Afrika. Transactions of the British Bryological Society 4 (3): 437–445. doi: 10.1179/006813863804812363 Grolle R (1963b) Über Kurzia v. Martens. Revue Bryologique et Lichénologique 32 (3/4): 166–180. Grolle R (1963c) Ein neuer Tylimanthus aus Tasmanien. Nova Hedwigia 6 (3/4): 391–394. Grolle R (1963d) Notulae hepaticologicae VII-IX. Revue Bryologique et Lichénologique 32 (3/4): 157–165. Grolle R (1964a) Miscellanea hepaticologica 11-20. Journal of Japanese Botany 39 (6): 173–178. Grolle R (1964b) Eine Nardia aus Africa. Botanical Magazine, Tokyo 77 (914): 297–299. doi: 10.15281/jplantres1887.77.297 Grolle R (1964c) Notulae hepatilogicae XV. Neue Notizen über Kurzia v. Mart. und Verwandte. Journal of Japanese Botany 39 (3): 79–81. Grolle R (1964d) Eine neue Echinocolea auf Celebes. Botanical Magazine, Tokyo 77 (915): 333–335. doi: 10.15281/jplantres1887.77.333 Grolle R (1964e) Neesioscyphus – eine neue Lebermoosgattung mit gedrehten Sporogonklappen. Österreichische Botanische Zeitschrift 111 (1): 19–36. doi: 10.1007/BF01373318 Grolle R (1964f) Miscellanea hepaticologica 1-10. Österreichische Botanische Zeitschrift 111 (2/3): 185–192. doi: 10.1007/BF01373763 Grolle R (1964g) Miscellanea hepaticologica 21-30. Journal of Japanese Botany 39 (8): 236–241. Grolle R (1964h) Über neue bemerkenswerte Acromastigum-arten. Österreichische Botanische Zeitschrift 111 (2/3): 240–256. doi: 10.1007/BF01373767 Grolle R (1964i) Jamesoniella carringtonii – eine Plagiochila in Nepal mit Perianth. Transactions of the British Bryological Society 4 (4): 653–663. doi: 10.1179/006813864804812227 Grolle R (1964j) Notulae hepaticologicae X-XIII. Revue Bryologique et Lichénologique 33 (1/2): 224–229. Grolle R (1965a) Die Lebermoosgattungen Blepharidophyllum Ångstr. und Krunodiplophyllum nov. gen. (Scapaniaceae). Journal of the Hattori Botanical Laboratory 28: 55–74. Grolle R (1965b) Lebermoose aus Neuguinea. 1. Journal of the Hattori Botanical Laboratory 28: 43–54. Grolle R (1965c) Über Gymnanthe concinna und Lethocolea. Botanical Magazine, Tokyo 78 (921): 79–84. doi: 10.15281/jplantres1887.78.79 Grolle R (1965d) Miscellanea hepaticologica 41–50. Journal of Japanese Botany 40 (7): 206–219. Grolle R (1965e) Miscellanea hepaticologica 31–40. Journal of the Hattori Botanical Laboratory 28: 101–106. Grolle R (1965f) Wettsteinia Schifn. Journal of the Hattori Botanical Laboratory 28: 94–100. Grolle R (1965g) Harpanthus drummondii – ein Lebermoosendemit des östlichen Nordamerika. Österreichische Botanische Zeitschrift 112 (3): 268–284. doi: 10.1007/BF01372951 Grolle R (1966a) Eine neue Lebermoosgattung (Lophoziaceae) aus Nepal. Revue Bryologique et Lichénologique 34 (1/2): 187–190. 708 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Grolle R (1966b) Über Diplasiolejeunea in Asien. Feddes Repertorium 73 (2): 78–89. doi: 10.1002/fedr.19660730203 Grolle R (1966c) Notulae hepaticologicae XIV. Zwei weitere Neesioscyphus-Arten. Revue Bryologique et Lichénologique 34 (1/2): 182–186. Grolle R (1966d) Miscellanea hepaticologica 61-70. Journal of Japanese Botany 41 (8): 225–232. Grolle R (1966e) Lebermoose aus Neuguinea. 3. Stenolejeunea. Journal of the Hattori Botanical Laboratory 29: 75–78. Grolle R (1966f) Gymnomitrion crenulatum und Verwandte. Transactions of the British Bryological Society 5 (1): 86–94. doi: 10.1179/006813866804804567 Grolle R (1966g) Lebermoose aus Neuguinea. 5. Telaranea. Journal of the Hattori Botanical Laboratory 29: 279–289. Grolle R (1966h) Lebermoose aus Neuguinea. 4. Schistochila. Journal of the Hattori Botanical Laboratory 29: 238–252. Grolle R (1966i) Lebermoose aus Neuguinea. 2. Zweite Fundliste. Journal of the Hattori Botanical Laboratory 29: 70–74. Grolle R (1966j) Herzogobryum – eine beblätterte Lebermoosgattung mit dorsal verzahnten Merophyten. Österreichische Botanische Zeitschrift 113 (2): 220–234. doi: 10.1007/ BF01441035 Grolle R (1966k) Die Lebermoose Nepals. Khumbu Himal, Ergebnisse des Forschungsunternehmens Nepal Himalaya 1 (4): 262–298. Grolle R (1967a) Monographie der Lepidolaenaceae. Journal of the Hattori Botanical Laboratory 30: 1–53. Grolle R (1967b) Lebermoose aus Neuguinea. 6. Dritte Fundliste. Journal of the Hattori Botanical Laboratory 30: 113–118. Grolle R (1968a) Lebermoose aus Neuguinea. 7. Vierte Fundliste. Journal of the Hattori Botanical Laboratory 31: 1–12. Grolle R (1968b) Monographie der Gattung Nowellia. Journal of the Hattori Botanical Laboratory 31: 20–49. Grolle R (1968c) Gottschelia – eine neue Jungermanniales-Gattung der Paläotropis. Journal of the Hattori Botanical Laboratory 31: 13–19. Grolle R (1968d) Einige ostmalesische Lebermoose. Nova Hedwigia 16: 147–159. Grolle R (1969a) Miscellanea hepaticologica 101-110. Österreichische Botanische Zeitschrift 117 (1): 1–6. doi: 10.1007/BF01376933 Grolle R (1969b) Miscellanea Hepaticologica (91-100). Transactions of the British Bryological Society 5 (4): 766–774. doi: 10.1179/006813869804146682 Grolle R (1969c) Novae Guineae hepaticae Schusteranae. I. Journal of the Hattori Botanical Laboratory 32: 140–144. Grolle R (1970a) Radula castlei sp. nov. und Anmerkungen zur Gattung Radula. Bryologist 73 (4): 662–668. doi: 10.2307/3241278 Grolle R (1970b) Eine neuen Jackiella aus Tasmanien. Journal of the Hattori Botanical Laboratory 33: 222–224. Grolle R (1970c) Lebermoose aus Neuguinea. 8. Mastigopelma. Journal of the Hattori Botanical Laboratory 33: 36–40. World checklist of hornworts and liverworts 709 Grolle R (1971a) Die Lebermoose der Crozet-Inseln (Subantarktis). Lindbergia 1 (1/2): 80–82. Grolle R (1971b) Jamesoniella und Verwandte. Feddes Repertorium 82 (1): 1–100. doi: 10.1002/fedr.19710820102 Grolle R (1971c) Lebermoose aus Neuguinea. 9. Jungermannia nivea sp. nov. Miscellanea Bryologica et Lichenologica 6 (1): 1–3. Grolle R (1971d) Hepaticopsida. In: van Zinderen Bakker Sr EM (Eds) Winterbottom JM, Dyer RA (eds.), Marion and Prince Edward Islands. Report on the South African Biological and Geological Expedition 1965–1966. A. A. Balkema, Cape Town, 228–236. Grolle R (1972a) Miscellanea hepaticologica 121-130. Journal of the Hattori Botanical Laboratory 36: 547–551. Grolle R (1972b) Bazzania in Europa und Makaronesien. Zur Taxonomie und Verbreitung. Lindbergia 1 (3/4): 193–204. Grolle R (1972c) Zur Kenntnis von Adelanthus Mitt. Journal of the Hattori Botanical Laboratory 35: 325–370. Grolle R (1973a) Miscellanea hepaticologica 131-140. Herzogia 3: 75–82. Grolle R (1973b) Nephelolejeunea - eine neue Gattung der Tuyamaelloideae. Journal of the Hattori Botanical Laboratory 37: 251–261. Grolle R (1974a) Eine neue Taxilejeunea aus Madagaskar und Reunion. Journal of Bryology 8 (1): 93–96. doi: 10.1179/jbr.1974.8.1.93 Grolle R (1974b) Lebermoose aus Neuguinea. 12. Rhaphidolejeunea. Journal of the Hattori Botanical Laboratory 38: 651–655. Grolle R (1975a) Diplasiolejeunea in Australasien. Feddes Repertorium 86 (1/2): 75–82. doi: 10.1002/fedr.19750860108 Grolle R (1975b) Haplolejeunea aus Madagascar – Eine weitere neue Gattung der Tuyamaelloideae. Journal of the Hattori Botanical Laboratory 39: 203–210. Grolle R (1975c) Miscellanea hepaticologica 151-160. Lindbergia 3 (1/2): 47–56. Grolle R (1975d) Miscellanea hepaticologica 141-150. Journal of Bryology 8 (4): 483–492. doi: 10.1179/jbr.1975.8.4.483 Grolle R (1976a) Verzeichnis der Lebermoose Europas und benachbarter Gebiete. Feddes Repertorium 87 (3/4): 171–279. doi: 10.1002/fedr.19760870303 Grolle R (1976b) Eine weitere Siphonolejeunea –S. elegantissima (Steph.) comb. nov. aus Australien. Journal of the Hattori Botanical Laboratory 41: 405–409. Grolle R (1976c) Drepanolejeunea subg. Kolpolejeunea – eine neue Untergattung aus der Palaeotropis. Journal of the Hattori Botanical Laboratory 40: 191–216. Grolle R (1977a) Miscellanea hepaticologica 161-170. Journal of Bryology 9 (4): 529–538. doi: 10.1179/jbr.1977.9.4.529 Grolle R (1977b) Pictolejeunea – eine neue Gattung der Lejeuneoideae aus der Neotropis und Borneo. Feddes Repertorium 88 (4): 247–256. doi: 10.1002/fedr.19770880402 Grolle R (1978a) Eine neue Diplasiolejeunea-Art aus Sri Lanka. Feddes Repertorium 89 (5/6): 301–305. doi: 10.1002/fedr.19780890502 Grolle R (1978b) Lebermoose aus Neuguinea. 16. Acromastigum. Journal of the Hattori Botanical Laboratory 44: 1–15. 710 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Grolle R (1979a) Lebermoose aus Neuguinea. 17. Harpalejeunea. Journal of the Hattori Botanical Laboratory 46: 43–47. Grolle R (1979b) Miscellanea hepaticologica 171-180. Journal of Bryology 10 (3): 263–272. doi: 10.1179/jbr.1979.10.3.263 Grolle R (1979c) Miscellanea hepaticologica 181-190. Journal of the Hattori Botanical Laboratory 45: 173–183. Grolle R (1979d) Miscellanea hepaticologica 191-200. Journal of the Hattori Botanical Laboratory 46: 337–355. Grolle R (1980a) Nephelolejeunea in Australasien. Journal of the Hattori Botanical Laboratory 48: 161–170. Grolle R (1980b) Schusterolejeunea Grolle nom. nov. statt Cladocolea Schust. 1963, non van Tieghem 1895. Journal of Bryology 11 (1): 105–106. doi: 10.1179/jbr.1980.11.1.105 Grolle R (1980c) Miscellanea hepaticologica 201-210. Journal of Bryology 11 (2): 325–334. doi: 10.1179/jbr.1980.11.2.325 Grolle R (1980d) Zur Kenntnis der Lebermoose von Samoa. I. Wissenschaftliche Zeitschrift der Friedrich-Schiller-Universität Jena/hüringen. Mathematisch-naturwissenschaftliche Reihe 29 (4): 637–648. Grolle R (1981) Miscellanea hepaticologica 211-220. Journal of the Hattori Botanical Laboratory 49: 85–92. Grolle R (1982) Übersicht der Lejeuneaceae in Tasmanien. Wissenschaftliche Zeitschrift der Friedrich-Schiller-Universität Jena/hüringen. Mathematisch-naturwissenschaftliche Reihe 31 (2): 207–227. Grolle R (1983a) Hepatics of Europe including the Azores: an annotated list of species, with synonyms from the recent literature. Journal of Bryology 12 (3): 403–459. doi: 10.1179/ jbr.1983.12.3.403 Grolle R (1983b) Nomina generica hepaticarum; references, types and synonymies. Acta Botanica Fennica 121: 1–62. Grolle R (1984a) Miscellanea hepaticologica 221-230. Journal of the Hattori Botanical Laboratory 55: 501–511. Grolle R (1984b) Zur Kenntnis der Lejeuneoideae in Cuba (1): Cyclolejeunea. Wissenschaftliche Zeitschrift der Friedrich-Schiller-Universität Jena/hüringen. Mathematisch-naturwissenschaftliche Reihe 33: 759–764. Grolle R (1984c) Kymatolejeunea Grolle – eine neue Gattung der Lejeuneoideae aus Neuseeland. Wissenschaftliche Zeitschrift der Friedrich-Schiller-Universität Jena/hüringen. Mathematisch-naturwissenschaftliche Reihe 32 (6): 1005–1012. Grolle R (1985a) Zur Kenntnis der Lebermoosgattung Otolejeunea. Haussknechtia 2: 45–56. Grolle R (1985b) Miscellanea hepaticologica 231-240. Journal of the Hattori Botanical Laboratory 58: 197–202. Grolle R (1986a) Miscellanea hepaticologica 241-250. Journal of the Hattori Botanical Laboratory 61: 249–255. Grolle R (1986b) Cololejeunea schaeferi spec. nov., ein verkanntes Lebermoos in Makaronesien. Journal of Bryology 13 (4): 487–495. doi: 10.1179/jbr.1985.13.4.487 World checklist of hornworts and liverworts 711 Grolle R (1987) Miscellanea hepaticologica 251-260. Journal of the Hattori Botanical Laboratory 63: 437–443. Grolle R (1988a) Verzeichnis der Lebermoose von Ascension Island nebst Beschreibung von Cheilolejeunea ascensionis (Hook. f. et Tayl.) Grolle, comb. nov. Haussknechtia 4: 43–49. Grolle R (1988b) Zur Kenntnis der Lejeuneoideae in Cuba (2): Lejeunea subg. Macrolejeunea Spruce. Wissenschaftliche Zeitschrift der Friedrich-Schiller-Universität Jena/hüringen. Mathematisch-naturwissenschaftliche Reihe 37: 169–176. Grolle R (1988c) Miscellanea hepaticologica 261-270. Journal of the Hattori Botanical Laboratory 65: 403–410. Grolle R (1989a) Marsupidium in Guyana. Journal of the Hattori Botanical Laboratory 66: 337–342. Grolle R (1989b) Über Asterella subg. Brachyblepharis in Lateinamerika. Wissenschaftliche Zeitschrift der Friedrich-Schiller-Universität Jena/hüringen. Mathematisch-naturwissenschaftliche Reihe 38 (2): 231–239. Grolle R (1989c) A technically new lectotypiication of Harpalejeunea (Hepaticae). Taxon 38 (1): 88–90. doi: 10.2307/1220901 Grolle R (1989d) Adelanthus on Mt. Roraima. Journal of the Hattori Botanical Laboratory 67: 243–247. Grolle R (1989e) Two new species of Cololejeunea from Bhutan. Journal of Bryology 15 (2): 281–287. doi: 10.1179/jbr.1988.15.2.281 Grolle R (1991) Miscellanea hepaticologica 281-290. Journal of the Hattori Botanical Laboratory 69: 185–194. Grolle R (1992a) Zwei neue Arten der Lebermoosgattung Diplasiolejeunea aus der Neotropis. Beiträge zur Phytotaxonomie 15: 105–110. Grolle R (1992b) Austrolejeunea jarmaniana, a new species of hepaticae from Tasmania. Nova Hedwigia 55 (1/2): 111–117. Grolle R (1995) he hepaticae and anthocerotae of the East African Islands. An annotated catalogue. Bryophytorum Bibliotheca 48: 1–178. Grolle R (2001) Miscellanea hepaticologica 291-300. Haussknechtia 8: 59–69. Grolle R (2002) Two new synonyms of Metzgeria furcata (L.) Dumort. Cryptogamie, Bryologie 23 (3): 209–210. Grolle R, Gradstein SR (1988) Haesselia, a new genus of Cephaloziaceae (Hepaticae) from Mt. Roraima, Guyana. Journal of the Hattori Botanical Laboratory 64: 327–334. Grolle R, Heinrichs J (1999) Redescription and synonymy of Plagiochila aerea Taylor 1846 (Hepaticae), irst described as Lycopodium pinnatum by Lamarck 1792. Nova Hedwigia 68: 511–526. Grolle R, Long DG (2000) An annotated check-list of the hepaticae and anthocerotae of Europe and Macaronesia. Journal of Bryology 22 (2): 103–140. doi: 10.1179/jbr.2000.22.2.103 Grolle R, Onraedt M (1974) Lebermoose aus Madagaskar und den Maskarenen (1). Lindbergia 2 (3/4): 230–233. Grolle R, Persson H (1966) Die Gattung Tylimanthus auf den Atlantischen Inseln. Svensk Botanisk Tidskrift 60 (1): 164–174. 712 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Grolle R, Piippo S (1984) Annotated catalogue of Western Melanesian bryophytes. I. Hepaticae and Anthocerotae. Acta Botanica Fennica 125: 1–86. Grolle R, Piippo S (1986) Bryophyte lora of the Huon Peninsula, Papua New Guinea. XVI. Pallaviciniaceae (Hepaticae). Acta Botanica Fennica 133: 59–79. Grolle R, Piippo S (1990) Bryophyte lora of the Huon Peninsula, Papua New Guinea. XXXVII. Leucolejeunea (Lejeuneaceae, Hepaticae). Annales Botanici Fennici 27 (2): 119–129. Grolle R, Reiner-Drehwald ME (1997) Cheilolejeunea oncophylla (Ångstr.) Grolle & Reiner comb. nov. (Lejeuneaceae), from the neotropics. Journal of Bryology 19 (4): 781–785. doi: 10.1179/jbr.1997.19.4.781 Grolle R, Reiner-Drehwald ME (1999) Review of the genus Harpalejeunea (Lejeuneaceae) including the description of H. grandis, a new species from the páramos of Colombia. Journal of Bryology 21 (1): 31–45. doi: 10.1179/jbr.1999.21.1.31 Grolle R, Reiner-Drehwald ME (2005) Pictolejeunea levis, a new species of Lejeuneaceae (Jungermanniopsida) from Cuba. Journal of Bryology 27 (3): 281–283. doi: 10.1179/174328205X70038 Grolle R, Schultze-Motel W (1972) Vorläuiges Verzeichnis der Lebermoose von Samoa. Journal of the Hattori Botanical Laboratory 36: 75–89. Grolle R, Seppelt RD (1986) Seppeltia, a new leafy genus of Metzgeriales from Macquarie Island. Journal of the Hattori Botanical Laboratory 60: 275–282. Grolle R, So ML (1997a) Plagiochila bischleriana, a new species from Nepal. Cryptogamie: Bryologie, Lichénologie 18 (3): 191–193. Grolle R, So ML (1997b) Studies on Plagiochila in China. I. Bryologist 100 (4): 467–477. doi: 10.2307/3244409 Grolle R, So ML (1998a) Studies on Plagiochila in China. II. Bryologist 101 (2): 282–294. doi: 10.2307/3244206 Grolle R, So ML (1998b) Notes on Plagiochila sect. Caducilobae in East Asia. Systematic Botany 23 (4): 459–465. doi: 10.2307/2419377 Grolle R, So ML (1998c) Plagiochila caulimammillosa, a peculiar new species from Yunnan, China. Journal of Bryology 20 (1): 41–49. Grolle R, So ML (1999a) Studies of Plagiochila sect. Subtropicae in Asia. Bryologist 102 (1): 67–75. doi: 10.2307/3244462 Grolle R, So ML (1999b) On the Plagiochila species of sect. Zonatae with paraphyllia or mamillose stems (Hepaticae). Systematic Botany 24 (3): 297–310. doi: 10.2307/2419690 Grolle R, So ML (1999c) Studies on Plagiochila in China. III. Bryologist 102 (2): 200–207. doi: 10.2307/3244360 Grolle R, So ML (2000) Notes on Plagiochila section Firmae (Hepaticae). Systematic Botany 25 (1): 5–14. doi: 10.2307/2666668 Grolle R, Váňa J (1992) Eine neue Jungermannia (Hepaticae, Jungermanniaceae) aus Nepal. Fragmenta Floristica et Geobotanica 37 (1): 3–6. Grolle R, Vanden Berghen C (1970) Une genre nouveau pour la famille Cephaloziellaceae: Cephalojonesia Grolle. Revue Bryologique et Lichénologique 37 (4): 763–768. Grolle R, Zhu R-L (1999) Drepanolejeunea longii (Lejeuneaceae, Hepaticae), a new species from Bhutan. Annales Botanici Fennici 36 (2): 115–118. World checklist of hornworts and liverworts 713 Grolle R, Zhu R-L (2000) A study of Drepanolejeunea subg. Rhaphidolejeunea (Herzog) Grolle & R.L.Zhu, stat. nov. (Hepaticae, Lejeuneaceae) in China with notes on its species elsewhere. Nova Hedwigia 70 (3/4): 373–396. Grolle R, Zhu R-L (2002) On Macrocolura and the subdivision of Colura (Lejeuneaceae, Hepaticae). Journal of the Hattori Botanical Laboratory 92: 181–190. Grolle R, Zhu R-L, Gradstein SR (2001) On Cyrtolejeunea A. Evans (Lejeuneaceae, Hepaticae). Taxon 50 (4): 1067–1074. doi: 10.2307/1224721 Grolle R, Schill DB, Long DG (2003) Notes on Gottschelia (Jungermanniales, Lophoziaceae), with a description of G. patoniae, a new species from the East Himalaya. Journal of Bryology 25 (1): 3–6. doi: 10.1179/037366803125002608 Haarbrink J (1981) Studies on Colombian cryptogams. XI. High Andean species of Frullania subg. Chonanthelia (Hepaticae). Lindbergia 7 (1): 47–57. Hagborg A, Söderström L, von Konrat M (2013) Notes on Early Land Plants Today. 42. Validation of Nanomarsupella (Gymnomitriaceae, Marchantiophyta). Phytotaxa 112 (1): 16–17. doi: 10.11646/phytotaxa.112.1.3 Hallingbäck T, Hodgetts N (2000) Mosses, liverworts, and hornworts. Status survey and conservation action plan for bryophytes. International Union for Conservation of Nature, Gland, 106 pp. Hamlin BG (1972) Hepaticae of New Zealand, parts I and II. Index of binominals and preliminary checklist. Records of the Dominion Museum 7: 243–366. Hampe E (1836) Prodromus lorae Hercyniae. Gebauerschen Buchdruckerei, Halle, 90 pp. doi: 10.5962/bhl.title.6713 Hampe E (1847) Berichte über die Hepaticae, welche Hr. Moritz in Columbien sammelte und dem königlichen Herbarium in Schönberg überlieferte, nach Synopsis hepaticarum und den Moritzschen Nummern aufgefürt. Linnaea 20 (3): 321–336. Hampe E (1851a) Hepaticae Oerstedianae (continuatio et inis). Linnaea 24 (6): 640–641. Hampe E (1851b) Hepaticae Oerstedianae. Linnaea 24 (3): 300–304. Hampe E (1854) Plantae quaedem Lechlerianae. Enumeratio hepaticarum, quae in Sectione II. plantarum chilensium et in pl. peruvianis W. Lechler collectis et a R. J. Hohenacker editis occurrunt. Linnaea 27 (5): 553–556. Hampe E, Gottsche CM (1852) Expositio hepaticarum Portoricensium, quas collegit Schwanecke, hortulanus. Linnaea 25 (3): 337–358. Hartman CJ (1838) Handbok i Skandinaviens lora, tredje upplagan. Sednare delen: loran. Zacharias Hæggströms förlag, Stockholm, 350 pp. Hartman CJ (1864) Handbok i Skandinaviens lora, nionde upplagan, sednare delen: mossor. Zacharias Hæggströms förlag, Stockholm, 120 pp. doi: 10.5962/bhl.title.9746 Hartman CJ (1871) Handbok i Skandinaviens lora, tionde upplagan, sednare delen: mossor. Zacharias Hæggströms förlag, Stockholm, 179 pp. Hartmann FA, Wilson R, Gradstein SR, Schneider H, Heinrichs J (2006) Testing hypothesis on species delimitations and disjunctions in the liverwort Bryopteris (Jungermanniopsida: Lejeuneaceae). International Journal of Plant Sciences 167 (6): 1205–1214. doi: 10.1086/508023 Hasegawa J (1979) Taxonomical studies on Asian anthocerotae. I. Acta Phytotaxonomica et Geobotanica 30 (1/3): 15–30. 714 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Hasegawa J (1980) Taxonomical studies on Asian anthocerotae. II. Some Asian species of Dendroceros. Journal of the Hattori Botanical Laboratory 47: 287–309. Hasegawa J (1984) Taxonomical studies on Asian anthocerotae. IV. A revision of the genera Anthoceros, Phaeoceros and Folioceros in Japan. Journal of the Hattori Botanical Laboratory 57: 241–272. Hasegawa J (1986a) A collection of the Anthocerotae from Seram and Ambon. Acta Phytotaxonomica et Geobotanica 37 (1/3): 9–15. Hasegawa J (1986b) he anthocerotae collected by Drs. Z. Iwatsuki and N. Kitagawa in New Caledonia and Fiji. Journal of the Hattori Botanical Laboratory 60: 379–390. Hasegawa J (1993a) Taxonomic results of the BRYOTROP expedition to Zaire and Rwanda. 5. Anthocerotae. Tropical Bryology 8: 51–52. Hasegawa J (1993b) Taxonomical studies on Asian anthocerotae. V. A short revision of Taiwanese anthocerotae. Acta Phytotaxonomica et Geobotanica 44 (2): 97–112. Hasegawa J (1994a) A remarkable new species of Phaeoceros (Anthocerotae) with canaliculatestriate spore surface. Journal of the Hattori Botanical Laboratory 75: 267–273. Hasegawa J (1994b) New classiication of anthocerotae. Journal of the Hattori Botanical Laboratory 76: 21–34. Hasegawa J (2001) A new species of Phaeoceros with remarkable spore features from southeast Asia. Bryological Research 7 (12): 373–377. Hässel GG (1961) Una nueva especie de un género nuevo para Argentina: Cyathodium steerei. Revue Bryologique et Lichénologique 30 (3/4): 223–231. Hässel GG (1962) Estudio de las Anthocerotales y Marchantiales de la Argentina. Opera Lilloana 7: 1–298. Hässel GG (1972a) Revisión taxonómica del género Riccardia (Hepaticae). Especies andinopatagonias y subantarticas incluyendo las Islas Juan Fernández, Malvinas, Georgias del Sur, etc. Revista del Museo Argentino de Ciencias Naturales “Bernardino Rivadavia” e Instituto Nacional de Investigación de las Ciencias Naturales: Ciencias Botánicas 4 (1): 1–242. Hässel GG (1972b) Riella gamundiae Hässel n. sp. (Hepaticae) la segunda especie del género hallada en Sudamérica. Revue Bryologique et Lichénologique 38 (3/4): 579–586. Hässel GG (1974) Hepaticae Fuegianae II-III. Comunicaciones del Museo Argentino de Ciencias Naturales “Bernardino Rivadavia” e Instituto Nacional de Investigación de las Ciencias Naturales: Ciencias Botánicas 2 (9): 45–51. Hässel GG (1979) Riella pampae Hässel n. sp. (Hepaticae) la tercera especie del género hallada en Sudamérica. Revista del Museo Argentino de Ciencias Naturales “Bernardino Rivadavia” e Instituto Nacional de Investigación de las Ciencias Naturales: Ciencias Botánicas 5 (9): 205–211. Hässel GG (1980) Liverworts new to South Georgia II. Journal of Bryology 11 (1): 107–128. doi: 10.1179/jbr.1980.11.1.107 Hässel GG (1981) Patagonian bryophytes. 2. On Herzogiaria teres (Steph.) Fulf. ex Hässel. Lindbergia 7 (1): 23–26. Hässel GG (1983) Informaciónes nomenclaturales sobre las especies del género Plagiochila (Hepaticae) de Argentina y Chile. Boletín de la Sociedad Argentina de Botánica 22 (1/4): 87–129. World checklist of hornworts and liverworts 715 Hässel GG (1986a) Leiosporoceros Hässel n. gen. and Leiosporocerotaceae Hässel n. fam. of the Anthocerotopsida. Journal of Bryology 14 (2): 255–259. doi: 10.1179/jbr.1986.14.2.255 Hässel GG (1986b) Neue Lebermoosfunde aus dem Nordosten Argentiniens – New indings of hepaticae from northeastern Argentina – Hallazgos nuevos de hepáticas en el nordeste de la Argentina. Veröfentlichungen des Geobotanischen Institutes der ETH, Stiftung Rübel, Zürich 91: 293–304. Hässel GG (1987) Progress with knowledge of the submerged genus Riella (Hepaticae) in Argentina. Symposia Biologica Hungarica 35: 335–342. Hässel GG (1988) Patagonian bryophytes. 10. Pisanoa chilensis Hässel, a new genus and species of Jungermanniaceae (Hepatophyta). Lindbergia 14 (3): 179–182. Hässel GG (1989a) Las especies de Phaeoceros (Anthocerotophyta) de América del Norte, Sud y Central; la ornamentación de sus esporas y taxonomía. Candollea 44 (2): 715–739. Hässel GG (1989b) Perdusenia rheophila Hässel, a new genus and species of Hepatophyta from southern South America. Revista del Museo Argentino de Ciencias Naturales “Bernardino Rivadavia” e Instituto Nacional de Investigación de las Ciencias Naturales: Ciencias Botánicas 7 (2): 11–17. Hässel GG (1990a) Patagonian bryophytes. 11. On Arctoscyphus ronsmithii Hässel (Hepatophyta), a new genus and species, with comments on Pedinophyllopsis Schust. et Inoue. Lindbergia 16 (4): 133–137. Hässel GG (1990b) Las especies de Anthoceros y Folioceros (Anthocerotophyta) de América del Norte, Sud y Central: la ornamentación de sus esporas y taxonomía. Candollea 45 (1): 201–220. Hässel GG (1996) La ubicación taxonómica de Leioscyphus repens Mitt. var. β fuegiensis Mass. (Hepaticophyta). Cryptogamie: Bryologie, Lichénologie 17 (3): 163–170. Hässel GG (1999) Chiloscyphus subgenus Phaeochiloscyphus (Hepatophyta, Geocalycaceae) from southern South America. Revista del Museo Argentino de Ciencias Naturales (Nueva serie) 1 (2): 121–127. Hässel GG (2000) Parachiloscyphus Hässel, a new subgenus of Chiloscyphus Corda (Hepatophyta) from southern South America, including a new species. Nova Hedwigia 70 (3/4): 451–460. Hässel GG (2001) Revision of the genus Leptoscyphus Mitt. (Hepatophyta) from southern South America. Journal of the Hattori Botanical Laboratory 91: 205–227. Hässel GG (2002) One new combination and one new species in Leiomitra (Trichocoleaceae, Hepatophyta) from southern South America. Novon 12 (4): 465–470. doi: 10.5962/bhl. title.744 Hässel GG (2005) On Chiloscyphus aphelophyllus Hässel, sp. nov. and C. apophyllus Hässel, sp. nov. (Marchantiophyta, Geocalycaceae) from Chile. Journal of the Hattori Botanical Laboratory 98: 123–129. Hässel GG (2006a) Paraphymatoceros Hässel, gen. nov. (Anthocerotophyta). Phytologia 88 (2): 208–211. Hässel GG (2006b) Aneura polyclada, A. polyptera and A. denticulata (Aneuraceae) from South America, overlooked names since 1886. Bryologist 109 (1): 33–37. doi: 10.1639/0007-2745(2006)109[0033:APAPAA]2.0.CO;2 716 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Hässel GG (2009) Andinopatagonian species of Plagiochila (Plagiochilaceae, Marchantiophyta) Sectio I. Duseniae Carl emend. Hässel and sectio II. Jacquinotiae Hässel sect. nov. Nova Hedwigia 89 (1/2): 71–95. doi: 10.1127/0029-5035/2009/0089-0071 Hässel GG, Rubies MF (2009) Catalogue of Marchantiophyta and Anthocerotophyta of southern South America. Beihefte zur Nova Hedwigia 134: 1–672. Hässel GG, Solari SS (1970) Consideraciones sobre el género Austrolophozia (Hepaticae). Revista del Museo Argentino de Ciencias Naturales “Bernardino Rivadavia” e Instituto Nacional de Investigación de las Ciencias Naturales: Ciencias Botánicas 3 (6): 239–247. Hässel GG, Solari SS (1972) Sinopsis de las especies andinopatagónicas del género Tylimanthus (Hepaticae). Darwiniana 17: 568–591. Hässel GG, Solari SS (1975) Bryophyta Hepaticopsida: Calobryales, Jungermanniales: Vetaformaceae, Balantiopsidaceae. In: Guarrera SA (Ed.) Gamundi de Amos I, Rabinovich de Halperin D, Flora Criptogámica de Tierra del Fuego. 15, Fasc. 1. Fundación para la Educación, la Ciencia y la Cultura, Buenos Aires, 7–181. Hässel GG, Solari SS (1985) Catálogo de las hepáticas. In: Boelcke O, Moore DM, Roig FA (Eds) Transecta Botánica de la Patagonia Austral. CONICET, Buenos Aires, 299–342. Hässel GG, Villagrán C (2007) New species of Fossombronia (Hepatophyta, Fossombroniopsida) from Chile. Beihefte zur Nova Hedwigia 131: 13–20. Hatcher RE (1957) he genus Trichocolea in North, Central and South America – (Hepaticae). Lloydia 20 (3): 139–185. Hattori S (1941) Notulae de hepaticis japonicis (I). Journal of Japanese Botany 17: 457–466. Hattori S (1942) Notulae de hepaticis japonicis (IV). Journal of Japanese Botany 18 (11): 653–660. Hattori S (1943a) [Ricciaceae nipponicae]. Natural Science and Museum 14 (6): 138–143. Hattori S (1943b) Notulae de hepaticis japonicis (VI). Journal of Japanese Botany 19 (11): 345–356. Hattori S (1943c) Notulae de hepaticis japonicis (V). Journal of Japanese Botany 19 (7): 197– 202. Hattori S (1944a) Hepaticarum species novae et minus cognitae Nipponenses. III. Botanical Magazine, Tokyo 58 (686): 38–46. doi: 10.15281/jplantres1887.58.38 Hattori S (1944b) Hepaticarum species novae et minus cognitae Nipponenses. II. Botanical Magazine, Tokyo 58 (685): 1–7. doi: 10.15281/jplantres1887.58.1 Hattori S (1944c) Notula de hepaticis japonicis (VII). Journal of Japanese Botany 20: 103–112. Hattori S (1944d) Contributio ad loram hepaticarum austro-kiushiuensem. Bulletin of the Tokyo Science Museum 11: 1–203. Hattori S (1946) Hepaticarum species novae et minus cognitae nipponenses. V. Botanical Magazine, Tokyo 59 (693/694): 25–31. doi: 10.15281/jplantres1887.59.25 Hattori S (1947a) [Five new genera of Hepaticae]. Biosphaera 1 (1): 3–7. Hattori S (1947b) Contributio ad loram hepaticarum yakusimensem. II. Journal of the Hattori Botanical Laboratory 2: 1–26. Hattori S (1948a) Hepaticarum species novae vel minus cognitae Nipponenses. VI. Journal of the Hattori Botanical Laboratory 3: 37–52. World checklist of hornworts and liverworts 717 Hattori S (1948b) Contributio ad loram hepaticarum yakusimensem. III. Journal of the Hattori Botanical Laboratory 3: 1–35. Hattori S (1949) [A new species of Riccia found in prov. Shansi, north China]. Botanical Magazine, Tokyo 62 (733/734): 109. doi: 10.15281/jplantres1887.62.109 Hattori S (1950) Contributio ad loram hepaticarum yakusimensem. IV. Journal of the Hattori Botanical Laboratory 4: 49–70. Hattori S (1951a) Contributio ad loram hepaticarum yakusimensem. VI. Journal of the Hattori Botanical Laboratory 6: 7–23. Hattori S (1951b) Oil bodies of Japanese hepaticae (1). Journal of the Hattori Botanical Laboratory 5: 69–97. Hattori S (1951c) On a small collection of hepaticae from Dutch New Guinea. Botanical Magazine, Tokyo 64 (755/756): 112–119. doi: 10.15281/jplantres1887.64.112 Hattori S (1951d) Contributio ad loram hepaticarum yakusimensem. V. Journal of the Hattori Botanical Laboratory 5: 43–68. Hattori S (1952a) Hepaticarum species novae et minus cognitae nipponenses. VII. Botanical Magazine, Tokyo 65 (763/764): 13–17. doi: 10.15281/jplantres1887.65.13 Hattori S (1952b) Notulae de hepaticis japonicis (15). Journal of Japanese Botany 27 (2): 53–59. Hattori S (1952c) Hepaticae of Shikoku and Kyushu, southern Japan (2). Journal of the Hattori Botanical Laboratory 8: 21–46. Hattori S (1953a) Notes on little known Japanese species of hepaticae (3). Journal of Japanese Botany 28 (8): 231–235. Hattori S (1953b) Notulae de hepaticis japonicis (17). Journal of Japanese Botany 28 (6): 181–185. Hattori S (1953c) Hepaticae novae vel minus cognitae Nipponenses (8). Journal of the Hattori Botanical Laboratory 10: 35–48. Hattori S (1954) Hepaticae japonicae exsiccatae, ser. 1-6. Journal of the Hattori Botanical Laboratory 12: 76–90. Hattori S (1955) A remarkable Frullania species from northern Japan. Feddes Repertorium Specierum Novarum Regni Vegetabilis 58: 53–54. Hattori S (1956a) Notula de hepaticis japonicis, 18. Journal of Japanese Botany 31 (7): 201–204. Hattori S (1956b) Hepaticae japonicae exsiccatae, ser. 8. Journal of the Hattori Botanical Laboratory 17: 75. Hattori S (1957a) Hepaticae of Hayachine Mountain with special reference to Hepaticae occurring on serpentine rocks. Journal of the Hattori Botanical Laboratory 18: 106–118. Hattori S (1957b) Hepaticae of Hokkaido. II: Rishiri and Rebun Islands. Journal of the Hattori Botanical Laboratory 18: 78–92. Hattori S (1957c) [A short review of some genera of Japanese Lejeuneaceae]. Miscellanea Bryologica et Lichenologica 1 (14): 1–2. Hattori S (1964) A remarkable Saccogynidium (liverwort) from north Borneo. Journal of Japanese Botany 39 (7): 206–208. Hattori S (1966a) A remarkable Anastrophyllum (Hepaticae) from New Guinea. Botanical Magazine, Tokyo 79 (937): 342–344. doi: 10.15281/jplantres1887.79.342 718 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Hattori S (1966b) A remarkable Balantiopsis found in tropical Asia. Journal of Japanese Botany 41 (5): 129–133. Hattori S (1966c) Anthocerotae and hepaticae. In: Hara H (Ed.) he lora of Eastern Himalaya. Results of the botanical expedition to eastern Himalaya organized by the University of Tokyo 1960 and 1963. University of Tokyo Press, Tokyo, 501–536. Hattori S (1966d) Hepaticae and anthocerotae of Mt. Chokai, northern Japan. Journal of the Hattori Botanical Laboratory 29: 267–278. Hattori S (1967) Studies of the Asiatic species of the genus Porella (Hepaticae). I. Some new or little known Asiatic species of Porella. Journal of the Hattori Botanical Laboratory 30: 129–151. Hattori S (1968) Hepaticae collected by F. Schmid in Ceylon and Pakistan. Candollea 23: 287–294. Hattori S (1969) Studies of the Asiatic species of the genus Porella (Hepaticae). II. Journal of the Hattori Botanical Laboratory 32: 319–359. Hattori S (1970) Studies of the Asiatic species of the genus Porella (Hepaticae). III. Journal of the Hattori Botanical Laboratory 33: 41–87. Hattori S (1971a) Hepaticae. In: Hara H (Ed.) Flora of Eastern Himalaya, second report. Bulletin, University Museum, University of Tokyo 2: 222–240. Hattori S (1971b) Studies of the Asiatic species of the genus Porella (Hepaticae). IV. Journal of the Hattori Botanical Laboratory 34: 411–428. Hattori S (1972a) Frullania tamarisci complex and the species concept. Journal of the Hattori Botanical Laboratory 35: 202–251. Hattori S (1972b) Novae Guineae hepaticae Schusteranae. III. Species novae Frullaniae. Journal of the Hattori Botanical Laboratory 36: 411–443. Hattori S (1972c) Notes on the Asiatic species of the genus Frullania, Hepaticae. I. Journal of the Hattori Botanical Laboratory 36: 109–140. Hattori S (1973a) Notes on the Asiatic species of the genus Frullania, Hepaticae. II. Journal of the Hattori Botanical Laboratory 37: 55–84. Hattori S (1973b) Notes on the Asiatic species of the genus Frullania, Hepaticae. IV. Journal of the Hattori Botanical Laboratory 37: 121–152. Hattori S (1973c) Notes on the Asiatic species of the genus Frullania, Hepaticae. III. Journal of the Hattori Botanical Laboratory 37: 85–120. Hattori S (1974a) Frullania taxa in the Philippine collection made by Dr. and Mrs. A. J. Sharp and Dr. Z. Iwatsuki. Journal of the Hattori Botanical Laboratory 38: 169–183. Hattori S (1974b) A peculiar Bornean species of Frullania, Hepaticae. Bulletin of the National Science Museum, Tokyo (n.ser.) 17 (4): 307–310. Hattori S (1974c) Notes on the Asiatic species of the genus Frullania, hepaticae. VI. Journal of the Hattori Botanical Laboratory 38: 223–274. Hattori S (1974d) Notes on the Asiatic species of the genus Frullania, hepaticae. V. Journal of the Hattori Botanical Laboratory 38: 185–221. Hattori S (1975a) A remarkable species of Frullania (Hepaticae) from Java. Journal of Japanese Botany 50 (6): 161–163. Hattori S (1975b) Mr. M. Togashi’s collection of Frullania (Hepaticae), made in Malay Peninsula. Bulletin of the National Science Museum, Tokyo. Series B, Botany 1 (3): 109–120. World checklist of hornworts and liverworts 719 Hattori S (1975c) Studies of the Asiatic species of the genus Porella (Hepaticae). V. Journal of the Hattori Botanical Laboratory 39: 269–276. Hattori S (1975d) Notes on the Asiatic species of the genus Frullania, hepaticae. VII. Journal of the Hattori Botanical Laboratory 39: 277–313. Hattori S (1975e) Bryophyta, anthocerotae & hepaticae. In: Ohashi H (Ed.) Flora of Eastern Himalaya. Bulletin, University Museum, University of Tokyo 8: 1–458. Hattori S (1975f) Notes on the Asiatic species of the genus Frullania, hepaticae. VIII. Bulletin of the National Science Museum, Tokyo. Series B, Botany 1 (4): 141–163. Hattori S (1975g) A revision of Indian species of Frullania (Hepaticae) published by W. Mitten in 1861. Bulletin of the National Science Museum, Tokyo. Series B, Botany 1 (2): 73–81. Hattori S (1976a) A remarkable New Caledonian species of Frullania (Hepaticae). Journal of Japanese Botany 51 (7): 193–198. Hattori S (1976b) Notes on some species of the New Caledonian Frullaniaceae (Hepaticae). Bulletin of the National Science Museum, Tokyo. Series B, Botany 2 (3): 79–86. Hattori S (1976c) A new species of Porella (Hepaticae) from Hawaii. Miscellanea Bryologica et Lichenologica 7 (5): 85–87. Hattori S (1976d) Notes on the Asiatic species of the genus Frullania, hepaticae. X. Journal of the Hattori Botanical Laboratory 40: 461–507. Hattori S (1976e) A new species of Frullania (Hepaticae) from the Philippines. In: Kachroo P (Ed.) Recent advances in Botany. Bishen Singh Madendra Pal Singh, Dehra Dun, 66–69. Hattori S (1976f) Studies of the Asiatic species of the genus Porella (Hepaticae). VI. Journal of the Hattori Botanical Laboratory 40: 121–138. Hattori S (1977a) Frullania Memoranda, I. Miscellanea Bryologica et Lichenologica 7 (8): 162–163. Hattori S (1977b) Dr. H. Hürlimann’s collections of New Caledonian Frullaniaceae. Journal of the Hattori Botanical Laboratory 43: 409–438. Hattori S (1978a) Studies of the Asiatic species of the genus Porella (Hepaticae). VII. A synopsis of Asiatic Porellaceae. Journal of the Hattori Botanical Laboratory 44: 91–120. Hattori S (1978b) Notes on the Asiatic species of the genus Frullania, hepaticae. XI. Journal of the Hattori Botanical Laboratory 44: 525–554. Hattori S (1979a) A revision of the Australasian species of the genus Frullania, hepaticae. II. Journal of the Hattori Botanical Laboratory 46: 119–153. Hattori S (1979b) A revision of the Australasian species of the genus Frullania, hepaticae. I. Journal of the Hattori Botanical Laboratory 45: 323–363. Hattori S (1979c) Corrections of names of Porella taxa. Miscellanea Bryologica et Lichenologica 8 (4): 79. Hattori S (1980a) A revision of the subgenus Homotropantha of the genus Frullania, hepaticae. Journal of the Hattori Botanical Laboratory 47: 165–236. Hattori S (1980b) Dr. H. Inoue’s Frullania collection made in Formosa. Bulletin of the National Science Museum, Tokyo. Series B, Botany 6 (1): 33–40. Hattori S (1980c) Notes on Frullania species of Iriomote and Ishigaki Islands, the Ryukyu Archipelago. Journal of Japanese Botany 55 (5): 132–135. Hattori S (1980d) Notes on the Asiatic species of the genus Frullania, hepaticae. XII. Journal of the Hattori Botanical Laboratory 47: 85–125. 720 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Hattori S (1981a) Notes on the Asiatic species of the genus Frullania, hepaticae. XIII. Journal of the Hattori Botanical Laboratory 49: 147–168. Hattori S (1981b) Notes on the Paciic species of Frullaniaceae (Hepaticae), I. Journal of the Hattori Botanical Laboratory 49: 359–383. Hattori S (1982a) Corrections and additions to “A synopsis of New Guinean Frullania”. Miscellanea Bryologica et Lichenologica 9 (6): 123–125. Hattori S (1982b) Asian taxa of the Frullania dilatata complex. Journal of Japanese Botany 57 (9): 257–260. Hattori S (1982c) A small collection of Frullania from Yunnan and Sichuan (=Szechwan). Bulletin of the National Science Museum, Tokyo. Series B, Botany 8 (3): 93–100. Hattori S (1982d) A synopsis of New Guinean Frullania, hepaticae. Journal of the Hattori Botanical Laboratory 51: 203–271. Hattori S (1983) A revision of the Australasian species of the genus Frullania, hepaticae. III. Journal of the Hattori Botanical Laboratory 54: 133–182. Hattori S (1984a) New Caledonian Frullaniaceae. Journal of the Hattori Botanical Laboratory 57: 405–426. Hattori S (1984b) A new species of Frullania from Bonin Islands. Journal of Japanese Botany 59 (10): 308–311. Hattori S (1984c) Dr. Marie L. Hicks’ Frullania collection made in northern Queensland. Cryptogamie: Bryologie, Lichénologie 5 (1/2): 177–189. Hattori S (1985) A small collection of Frullania and Porella (Hepaticae) made on Viti Levu, Fiji Islands. Bulletin of the National Science Museum, Tokyo. Series B, Botany 11 (1): 11–17. Hattori S (1986a) New Zealand Porella species. Bulletin of the National Science Museum, Tokyo. Series B, Botany 12 (1): 31–36. Hattori S (1986b) Frullania collection made by Dr. H. Akiyama on Seram Island. Journal of the Hattori Botanical Laboratory 60: 239–253. Hattori S (1986c) Notes on the Asiatic species of the genus Frullania, hepaticae, XIV. Bulletin of the National Science Museum, Tokyo. Series B, Botany 12 (4): 127–138. Hattori S (1986d) Porellaceae from New Caledonia. Journal of Japanese Botany 61 (6): 185–192. Hattori S (1986e) A synopsis of New Caledonian Frullaniaceae. Journal of the Hattori Botanical Laboratory 60: 203–237. Hattori S (1987a) New or little-known species of Frullania (Frullaniaceae) from Queensland and New South Wales, Australia. Memoirs of the New York Botanical Garden 45: 544–555. Hattori S (1987b) he Frullania lora of Lord Howe Island. Bryologist 90 (4): 365–370. doi: 10.2307/3243098 Hattori S (1988a) Frullania lora of Mt. Albert Edward, Papua New Guinea. Journal of the Hattori Botanical Laboratory 65: 411–453. Hattori S (1988b) Frullania collections made by Dr. Barbara M. hiers in Queensland and New South Wales, Australia. Beihefte zur Nova Hedwigia 90: 147–158. Hattori S, Gao C (1985) Two new Frullania taxa from China. Journal of Japanese Botany 60 (1): 1–4. Hattori S, Griin III DG (1978) A new species of Frullania from the Andes of Venezuela. Miscellanea Bryologica et Lichenologica 8 (3): 47–48. World checklist of hornworts and liverworts 721 Hattori S, Inoue H (1954) On a new liverwort family Treubiaceae. Journal of the Hattori Botanical Laboratory 11: 99–102. Hattori S, Kamimura M (1971) A new genus of Frullaniaceae (Hepaticae) from Borneo. Journal of the Hattori Botanical Laboratory 34: 429–436. Hattori S, Kamimura M (1973) Some new or little-known Asiatic species of Frullania (Hepaticae), I. Journal of the Hattori Botanical Laboratory 37: 519–533. Hattori S, Kodama T (1953) A new Cololejeunea from Kyoto. Journal of the Hattori Botanical Laboratory 10: 57–58. Hattori S, Lin P-J (1985a) A preliminary study of Chinese Frullania lora. Journal of the Hattori Botanical Laboratory 59: 123–169. Hattori S, Lin PJ (1985b) Two new species of Chinese Frullania (Hepaticae). Journal of Japanese Botany 60 (4): 106–110. Hattori S, Lin PJ (1986) A new species of Frullania from Hainan Island, China. Journal of Japanese Botany 61 (10): 307–309. Hattori S, Mizutani M (1958) A revision of the Japanese species of the family Lepidoziaceae. Journal of the Hattori Botanical Laboratory 19: 76–118. Hattori S, Mizutani M (1967) Metacalypogeia schusterana and Metacalypogeia quelpartensis. Miscellanea Bryologica et Lichenologica 4 (8): 121–124. Hattori S, Mizutani M (1968) Asiatic species of Pseudolepicolea (Hepaticae). Journal of the Hattori Botanical Laboratory 31: 251–259. Hattori S, Mizutani M (1969) Studies in the lora of hailand. 59. Hepaticae. Dansk Botanisk Arkiv 27 (1): 91–98. Hattori S, Piippo S (1986) Bryophyte lora of the Huon Peninsula, Papua New Guinea. XV. Frullania (Frullaniaceae, Hepaticae). Acta Botanica Fennica 133: 25–58. Hattori S, Streimann H (1985) A collection of Frullania taxa from Papua New Guinea. Journal of the Hattori Botanical Laboratory 59: 101–121. Hattori S, haithong O (1977) A new species of Frullania (Hepaticae) from Java. Journal of Japanese Botany 52 (10): 289–292. Hattori S, haithong O (1978a) Dr. Parihar’s collection of Indian Frullania (Hepaticae). Journal of Japanese Botany 53 (5): 129–133. Hattori S, haithong O (1978b) Indian Frullania collection made by Prof. and Mrs. A. J. Sharp and Dr. Z. Iwatsuki. Journal of the Hattori Botanical Laboratory 44: 177–193. Hattori S, haithong O (1978c) A Frullania collection made by Dr. T. Tuyama in Laos. Journal of Japanese Botany 53 (6): 172–177. Hattori S, Zhang M-X (1985) Porellaceae of Shensi Province, China. Journal of Japanese Botany 60 (11): 321–326. Hattori S, Hong W, Inoue H (1962) On a small collection of hepaticae made on Quelpart Island (Korea). Journal of the Hattori Botanical Laboratory 25: 126–134. Hattori S, Sharp AJ, Mizutani M, Iwatsuki Z (1966) he systematic position and distribution of Treubia nana. Bryologist 69 (4): 488–492. doi: 10.2307/3240581 Hattori S, haithong O, Kitagawa N (1977) he genus Frullania in hailand. Journal of the Hattori Botanical Laboratory 43: 439–457. 722 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Haynes CC (1907) Two new species of Aytonia from Jamaica. Bulletin of the Torrey Botanical Club 34 (2): 57–60. doi: 10.2307/2478915 Haynes CC (1909) An enumeration of the Washington and Oregon hepaticae collected by Mr. A. S. Foster. Bryologist 12 (4): 65–71. doi: 10.2307/3238886 Haynes CC (1910) Sphaerocarpos hians sp. nov., with a revision of the genus and illustration of the species. Bulletin of the Torrey Botanical Club 37 (5): 215–230. doi: 10.2307/2479405 He X (1996a) Type studies on Pycnolejeunea (Lejeuneaceae, Hepaticae). II. Annales Botanici Fennici 33 (1): 51–58. He X (1996b) Type studies on Pycnolejeunea (Lejeuneaceae, Hepaticae), III. Two Asiatic species described by Hofmann. Annales Botanici Fennici 33 (1): 59–64. He X (1997) Type studies on Pycnolejeunea (Lejeuneaceae, Hepaticae). V. On the identity of Pycnolejeunea spinistipula Mizut. and Lepidolejeunea queenslandica hiers. Annales Botanici Fennici 34 (2): 127–132. He X (1999) A taxonomic monograph of the genus Pycnolejeunea (Lejeuneaceae, Hepaticae). Acta Botanica Fennica 163: 1–77. He X, Glenny DS (2010) Perssoniella and the genera of Schistochilaceae: a new classiication based on molecular phylogenies. Australian Systematic Botany 23 (4): 229–238. doi: 10.1071/SB10007 He X, Grolle R (2001) Xylolejeunea, a new genus of the Lejeuneaceae (Hepaticae) from the Neotropics, Madagascar and the Seychelles. Annales Botanici Fennici 38 (1): 25–44. He X, Sun Y (2013) Multigene evidence reveals the systematic position of Pleurocladopsis simulans (C. Massal.) R. M. Schust. within Schistochila Dumort., Schistochilaceae. Polish Botanical Journal 58 (2): 467–474. doi: 10.2478/pbj-2013-0060 He Q, Zhu R-L, Chantanaorrapint S, Kornochalert S, Printarakul N (2012a) Drepanolejeunea laciniata (Lejeuneaceae), a new species from northern hailand. Cryptogamie, Bryologie 33 (3): 291–298. doi: 10.7872/cryb.v33.iss3.2012.291 He Q, Wei Y-M, Zhu R-L (2012b) Vitalianthus guangxianus Zhu RL, He Q, Wei YM (Lejeuneaceae), sp. nov. from Guangxi, China. Journal of Bryology 34 (1): 32–36. doi: 10.1179/1743282011Y.0000000038 He Q, Liu L-J, Liu C-X, Liu G-L, Ma Y, Gao D-L, Wei Y-M, Shi R-P, Shu L, Zhu R-L (2013) Synonymy, taxonomic notes, and range extensions for several noteworthy Asian liverworts. Journal of Bryology 35 (2): 123–128. doi: 10.1179/174328213X13663649362415 He Q, Wei Y-M, Pócs T, Zhu R-L (2014a) he reappraisal of Capillolejeunea S.W.Arnell (Marchantiophyta, Lejeuneaceae). Phytotaxa 175 (3): 166–170. doi: 10.11646/phytotaxa.175.3.7 He X, Glenny D, Söderström L, Hagborg A, von Konrat M (2014b) Notes on Early Land Plants Today. 58. Historical circumscription of Schistochilaceae (Marchantiophyta) and a new combination in Schistochila. Phytotaxa 173 (1): 91–94. doi: 10.11646/phytotaxa.173.1.10 Hedwig J (1784) heoria generationis et fructiicationis. Typis Academiae imper. scientarum, Petropoli [St. Petersburg], 164 pp. Heeg M (1893) Hepaticarum species novae. Revue Bryologique 20 (5): 81–83. Heinrichs J (2002) A taxonomic revision of Plagiochila sect. Hylacoetes, sect. Adiantoidea and sect. Fuscoluteae in the Neotropics with a preliminary subdivision of neotropical Plagiochilaceae into nine lineages. Bryophytorum Bibliotheca 58: 1–184. World checklist of hornworts and liverworts 723 Heinrichs J, Gradstein SR, Grolle R (1998) A revision of the neotropical species of Plagiochila (Dumort.) Dumort. (Hepaticae) described by Olof Swartz. Journal of the Hattori Botanical Laboratory 85: 1–32. Heinrichs J, Groth H, Gradstein SR, Rycroft DS, Cole WJ, Anton H (2001) Plagiochila rutilans (Hepaticae): A poorly known species from tropical America. Bryologist 104 (3): 350–361. doi: 10.1639/0007-2745(2001)104[0350:PRHAPK]2.0.CO;2 Heinrichs J, Groth H, Holz H, Rycroft DS, Renker C, Pröschold T (2002) he systematic position of Plagiochila moritziana, P. trichostoma, and P. delexa based on ITS sequence variation of nuclear ribosomal DNA, morphology, and lipophilic secondary metabolites. Bryologist 105 (2): 189–203. doi: 10.1639/0007-2745(2002)105[0189:TSPOPM]2.0.CO;2 Heinrichs J, Gradstein SR, Groth H, Lindner M (2003) Plagiochila cucullifolia var. anomala var. nov. from Ecuador, with notes on discordant molecular and morphological variation in Plagiochila. Plant Systematics and Evolution 242 (1/4): 205–216. doi: 10.1007/s00606-003-0063-5 Heinrichs J, Lindner M, Pócs T (2004) nrDNA internal transcribed spacer data reveal that Rhodoplagiochila R. M. Schust. (Marchantiophyta: Jungermanniales) is a member of Plagiochila sect. Arrectae Carl. Organisms, Diversity and Evolution 4 (1/2): 109–118. doi: 10.1016/j.ode.2004.01.001 Heinrichs J, Lindner M, Gradstein SR, Groth H, Buchbender V, Solga A, Fischer E (2005) Origin and subdivision of Plagiochila (Jungermanniidae: Plagiochilaceae) in tropical Africa based on evidence from nuclear and chloroplast DNA sequences and morphology. Taxon 54 (2): 317–333. doi: 10.2307/25065360 Heinrichs J, Rycroft DS, Feldberg K, Lindner M, Hartmann FA (2006) he systematic position of Plagiochila surinamensis inferred from nrDNA ITS sequences, morphology and phytochemistry. Journal of the Hattori Botanical Laboratory 100: 135–142. Heinrichs J, Hentschel J, Wilson R, Feldberg K, Schneider H (2007) Evolution of leafy liverworts (Jungermanniidae, Marchantiophyta): estimating divergence times from chloroplast DNA sequences using penalized likelihood with integrated fossil evidence. Taxon 56 (1): 31–44. doi: 10.2307/25065733 Heinrichs J, Hentschel J, Bombosch A, Fiebig A, Reise J, Edelmann M, Kreier H-P, SchäferVerwimp A, Caspari S, Schmidt AR, Zhu R-L, von Konrat M, Shaw B, Shaw AJ (2010) One species or at least eight? Delimitation and distribution of Frullania Dumort. s. l. (Jungermanniopsida, Porellales) inferred from nuclear and chloroplast DNA markers. Molecular Phylogenetics and Evolution 56 (3): 1105–1114. doi: 10.1016/j.ympev.2010.05.004 Heinrichs J, Bombosch A, Feldberg K, Kreier H-P, Hentschel J, Eckstein J, Long D, Zhu R-L, Schäfer-Verwimp A, Schmidt AR, Shaw B, Shaw AJ, Váňa J (2012a) A phylogeny of the northern temperate leafy liverwort genus Scapania (Scapaniaceae, Jungermanniales). Molecular Phylogenetics and Evolution 62 (3): 973–985. doi: 10.1016/j.ympev.2011.11.029 Heinrichs J, Dong S, Feldberg K, Schäfer-Verwimp A, Schmidt AR (2012b) Sphaerolejeunea (Lejeuneaceae, Porellales) is a synonym of Lejeunea. Phytotaxa 69: 7–15. Heinrichs J, Dong S, Yu Y, Schäfer-Verwimp A, Pócs T, Feldberg K, Hentschel J, Schmidt AR, Schneider H (2012c) A 150 year-old mystery solved: Transfer of the rheophytic endemic liverwort Myriocolea irrorata to Colura. Phytotaxa 66: 55–64. 724 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Heinrichs J, Dong S, Schäfer-Verwimp A, Pócs T, Feldberg K, Czumaj A, Schmidt AR, Reitner J, Renner MAM, Hentschel J, Stech M, Schneider H (2013) Molecular phylogeny of the leafy liverwort Lejeunea (Porellales): evidence for a Neotropical origin, uneven distribution of sexual systems and insuicient taxonomy. PLOS one 8 (12): 1–14. doi: 10.1371/ journal.pone.0082547 Heinrichs J, Schäfer-Verwimp A, Czumay A, Dong S, Scheben A, Feldberg K, Schneider H (2014) Towards a monophyletic classiication of Lejeuneaceae I: subtribe Leptolejeuneinae subtr. nov. Phytotaxa 156 (3): 165–174. doi: 10.11646/phytotaxa.165.3.7 Heinrichs J, Feldberg K, Bechteler J, Scheben A, Czumaj A, Pócs T, Schneider H, Schäfer-Verwimp A (2015) Integrative taxonomy of Lepidolejeunea (Jungermanniopsida: Porellales): Ocelli allow the recognition of two neglected species. Taxon 64 (2): 216–228. doi: 10.12705/642.5 Hell KG (1969) Brióitas talosas dos arredores da cidade de São Paulo (Brasil). Boletim de Faculdade de Filosoia, Ciências e Letras, Universidade de São Paulo, botánica 25: 1–190. Hendry TA, Wang B, Yang Y, Davis EC, Braggins JE, Schuster RM, Qui Y-L (2007) Evaluating phylogenetic positions of four liverworts from New Zealand, Neogrollea notabilis, Jackiella curvata, Goebelobryum unguiculatum and Herzogianthus vaginatus, using three chloroplast genes. Bryologist 110 (4): 738–751. doi: 10.1639/0007-2745(2007)110[738:EPPOFL ]2.0.CO;2 Henriques J (1886) Hepáticas colhidas em Portugal. Boletim da Sociedade Broteriana 4: 234–249. Hentschel J, Wilson R, Burghardt M, Zündorf H-J, Schneider H, Heinrichs J (2006) Reinstatement of Lophocoleaceae (Jungermanniopsida) based on chloroplast gene rbcL data: exploring the importance of female involucres for the systematics of Jungermanniales. Plant Systematics and Evolution 258 (2): 211–226. doi: 10.1007/s00606-006-0408-y Hentschel J, Paton JA, Schneider H, Heinrichs J (2007a) Acceptance of Liochlaena Nees and Solenostoma Mitt., the systematic position of Eremonotus Pearson and notes on Jungermannia L. s. l. (Jungermanniidae) based on chloroplast DNA sequence data. Plant Systematics and Evolution 268 (1/4): 147–157. doi: 10.1007/s00606-007-0549-7 Hentschel J, Zhu R-L, Long DG, Davison PG, Schneider H, Gradstein SR, Heinrichs J (2007b) A phylogeny of Porella (Porellaceae, Jungermanniopsida) based on nuclear and chloroplast DNA sequences. Molecular Phylogenetics and Evolution 45 (2): 693–705. doi: 10.1016/j.ympev.2007.05.005 Hentschel J, Feldberg K, Zündorf H-J, Hellwig FH, Schneider H, Heinrichs J (2007c) he systematic position of Pachyglossa and Clasmatocolea (Jungermanniopsida: Lophocoleaceae) inferred from nrDNA ITS sequences and morphology. Taxon 56 (4): 1136–1142. doi: 10.2307/25065908 Hentschel J, von Konrat MJ, Pócs T, Schäfer-Verwimp A, Shaw AJ, Schneider H, Heinrichs J (2009) Molecular insights into the phylogeny and subgeneric classiication of Frullania Raddi (Frullaniaceae, Porellales). Molecular Phylogenetics and Evolution 52 (1): 142–156. doi: 10.1016/j.ympev.2008.12.021 Hentschel J, von Konrat M, Söderström L, Hagborg A, Larraín J, Sukkharak P, Uribe J, Zhang L (2015) Notes on Early Land Plants Today. 72. Infrageneric classiication and new combinations, new names, new synonyms in Frullania. Phytotaxa 220 (2): 127-142 doi: 10.11646/phytotaxa.220.2.3. World checklist of hornworts and liverworts 725 He-Nygrén X, Piippo S (2003) Phylogenetic relationships of the generic complex ChiloscyphusLophocolea-Heteroscyphus (Geocalycaceae, Hepaticae): Insights from three chloroplast genes and morphology. Annales Botanici Fennici 40 (5): 317–329. He-Nygrén X, Juslén A, Ahonen I, Glenny D, Piippo S (2006) Illuminating the evolutionary history of liverworts (Marchantiophyta) - towards a natural classiication. Cladistics 22 (1): 1–31. doi: 10.1111/j.1096-0031.2006.00089.x Herzog T (1920) Die Bryophyten meiner zweiten Reise durch Bolivia (Nachtrag). Bibliotheca Botanica 88: 1–31. Herzog T (1921) Die Lebermoose der 2. Freiburger Molukkenexpedition und einige neue Arten der engeren Indomalaya. Beihefte zum Botanischen Centralblatt 38 (2): 318–332. Herzog T (1925a) Neue Bryophyten aus Brasilien. Repertorium Specierum Novarum Regni Vegetabilis 21 (1/7): 22–33. doi: 10.1002/fedr.19250210104 Herzog T (1925b) Contribuições ao conhecimento da lora briologica do Brasil. Archivos de Botanica do Estado de São Paulo 1 (2): 27–105. Herzog T (1926) Bryophyten der weiteren Indomalaya. Hedwigia 66 (6): 337–358. Herzog T (1927) Zwei Bryophytensammlungen aus Südamerika. Hedwigia 67 (6): 249–268. Herzog T (1928) Scapania portoricensis Hpe et G. Annales Bryologici 1: 91–112. Herzog T (1930a) Mnioloma Herzog nov. gen. hepaticarum. Annales Bryologici 3: 115–120. Herzog T (1930b) Studien über Drepanolejeunea I. Annales Bryologici 3: 126–149. Herzog T (1931a) Hepaticae. In: Irmscher E (Ed.) Beiträge zur lora von Borneo. Mitteilungen aus dem Institut für allgemeine Botanik in Hamburg 7 (3): 179–181. Herzog T (1931b) Hepaticae philippinenses a cl. C. J. Baker lectae. Annales Bryologici 4: 79–94. Herzog T (1932a) Neue Hepaticae aus der weiteren Indomalaya. Annales Bryologici 5: 83–98. Herzog T (1932b) Neue und bemerkenswerte Bryophyten, von H. Burgef 1927/28 auf Java und den Philippinen gesammelt. Annales Bryologici 5: 69–82. Herzog T (1932c) Beiträge zur Kenntnis der Gattung Plagiochila 1. Neotropische Arten. Hedwigia 72 (6): 195–242. Herzog T (1934a) Die Bryophyten der Andenreisen von C. Troll (Bolivia, Colombia, Panamá). Hedwigia 74 (2): 79–114. Herzog T (1934b) Studien über Drepanolejeunea II. Annales Bryologici 7: 57–94. Herzog T (1936a) Studien über Drepanolejeunea III. Annales Bryologici 9: 115–130. Herzog T (1936b) Descriptions of new species of New Zealand hepatics. Transactions and Proceedings of the Royal Society of New Zealand 65 (3): 350–356. Herzog T (1937) Neue Bryophyten vom Ruwenzori und aus dem Patagonischen Inlandeisgebiet. Repertorium Specierum Novarum Regni Vegetabilis 41 (14/25): 285–292. doi: 10.1002/fedr.19370411410 Herzog T (1938a) Hepaticae standleyanae costaricenses et hondurenses. Revue Bryologique et Lichénologique 11 (1): 5–30. Herzog T (1938b) Sphaerolejeunea, eine neue Gattung der Lejeuneaceae Schizostipae. Annales Bryologici 11: 86–89. Herzog T (1938c) Descriptions of new species of New Zealand hepatics, II. Transactions and Proceedings of the Royal Society of New Zealand 68 (1): 40–46. 726 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Herzog T (1938d) Beiträge zur Kenntnis der Gattung Plagiochila II. Palaeotropische Arten. Hedwigia 78 (3/4): 222–244. Herzog T (1939a) Studien über Drepanolejeunea IV. Annales Bryologici 12: 98–122. Herzog T (1939b) Zwei Bryophytensammlungen aus dem Sikkim-Himalaya. Annales Bryologici 12: 71–97. Herzog T (1939c) Zur Bryophytenlora Südchiles. I. Verzeichnis der gesammelten Bryophyten. Beihefte zum Botanischen Centralblatt 60B (1/2): 1–35. Herzog T (1940) Die Moose der Expedition Ljunger nach Patagonien, 1932-34. Arkiv för Botanik 29A (21): 1–17. Herzog T (1941a) Allisonia Herz., eine neue Gattung of the Haplolaenaceae. Hedwigia 80 (1/2): 77–83. Herzog T (1941b) Byssolejeunea, eine neue Gattung der Lejeuneaceae. Hedwigia 80 (1/2): 84–86. Herzog T (1942a) Die foliosen Lebermoose der Juan Fernández-Insel und der Osterinsel. In: Skottsberg C (Ed.) he Natural History of Juan Fernández and Easter Island. Botany 2. Almquist & Wiksell, Stockholm, 697–752. doi: 10.5962/bhl.title.25662 Herzog T (1942b) Lebermoose aus Sumatra. Annalen des Naturhistorischen Museums in Wien 53 (1): 358–373. Herzog T (1942c) Revision der Lebermoosgattung Leptolejeunea Spr. in der Indomalaya. Flora 135: 377–434. Herzog T (1942d) Beiträge zur Kenntnis neotropischer Bryophyten. Beihefte zum Botanischen Centralblatt 61B (3): 559–590. Herzog T (1942e) Drei neue Ceratolejeunea-Arten aus der Neotropis. Revue Bryologique et Lichénologique 13: 20–24. Herzog T (1943a) Eine kleine Lebermoos-Sammlung aus Chile. Acta Horti Gotoburgensis 15: 157–162. Herzog T (1943b) Micropterygium reimersianum Herz. Eine neue Art der Gattung aus Südbrasilien. Hedwigia 81 (5/6): 225–228. Herzog T (1943c) Rhaphidolejeunea Herz., eine neue Lejeuneaceengattung der Indomalaya. Mittheilungen des hüringischen Botanischen Vereins 50: 100–105. Herzog T (1947) Hepaticae von der Comoreninsel Johanna. Botaniska Notiser 100 (4): 317–334. Herzog T (1948) Studien über kritische und neue Lejeuneaceae der Indomalaya. Svensk Botanisk Tidskrift 42 (3): 230–241. Herzog T (1949a) Descriptions of new species of New Zealand hepatics III. Transactions and Proceedings of the Royal Society of New Zealand 77 (2): 253–256. Herzog T (1949b) Four new hepatics from Australia. Transactions of the British Bryological Society 1 (3): 181–189. doi: 10.1179/006813849804879096 Herzog T (1950a) Hepaticae borneenses (Oxford University expeditions to Sarawak, 1932). Transactions of the British Bryological Society 1 (4): 275–326. doi: 10.1179/006813850804878680 Herzog T (1950b) Miscellanea bryologica. II. Paleotropica. Memoranda Societatis pro Fauna et Flora Fennica 26: 37–66. World checklist of hornworts and liverworts 727 Herzog T (1950c) Miscellanea bryologica. I. Neotropica. Memoranda Societatis pro Fauna et Flora Fennica 25: 43–72. Herzog T (1951a) Hepaticae standleyanae costaricenses et hondurenses. Pars II. Revue Bryologique et Lichénologique 20 (1/2): 126–175. Herzog T (1951b) Revision der Lebermoos Gattung Lembidium Mitt. Arkiv för Botanik (n. ser.) 1 (13): 471–502. Herzog T (1952a) Nachtrag zu ‘Hepaticae borneenses (Oxford University Expedition to Sarawak, 1932)’. Transactions of the British Bryological Society 2 (1): 71–73. doi: 10.1179/006813852804878435 Herzog T (1952b) Aphanotropis Herz., eine neue Gattung der Lejeuneaceae aus Borneo. Transactions of the British Bryological Society 2 (1): 62–65. doi: 10.1179/006813852804878255 Herzog T (1952c) Miscellanea bryologica. III. Memoranda Societatis pro Fauna et Flora Fennica 27: 92–110. Herzog T (1952d) Drei neue Lebermoose aus West Patagonien. Revue Bryologique et Lichénologique 21 (3/4): 256–261. Herzog T (1952e) Hepaticae ecuadorienses a Cl. D:re Gunnar Harling annis 1946–1947 lectae. Svensk Botanisk Tidskrift 46 (1): 62–108. Herzog T (1952f) Eine neue Lebermoosgattung aus Westpatagonien, Chondrophyllum Herz. Revue Bryologique et Lichénologique 21 (1/2): 46–49. Herzog T (1952g) Perssoniella Herz. nov. gen. hepaticarum. Arkiv för Botanik (n.ser.) 2 (4): 265–269. Herzog T (1952h) Beiträge zur Kenntnis der argentinischen Bryophytenlora. Feddes Repertorium Specierum Novarum Regni Vegetabilis 55 (1): 1–27. doi: 10.1002/fedr.19520550102 Herzog T (1953a) Lebermoose aus Neukaledonien gesammelt von Dr. O. H. Selling. Arkiv för Botanik (n.ser.) 3 (3): 43–61. Herzog T (1953b) Eine neue Lebermossgattung, Pseudomarsupidium Herz. n. gen. aus Westpatagonien. Svensk Botanisk Tidskrift 47 (1): 34–42. Herzog T (1954) Zur Bryophytenlora Chiles. Revue Bryologique et Lichénologique 23 (1/2): 27–99. Herzog T (1955) Hepaticae aus Columbia und Peru. Feddes Repertorium Specierum Novarum Regni Vegetabilis 57 (1/2): 156–203. doi: 10.1002/fedr.4880570105 Herzog T (1957a) Lebermoose aus Ecuador gesammelt von Dr. E. Asplund. Svensk Botanisk Tidskrift 51 (1): 187–196. Herzog T (1957b) Eine Bryophytensammlung aus dem argentischen Nationalpark Nahuel-Huapi. Darwiniana 11 (2): 207–222. Herzog T (1960) Weitere Beiträge zur Bryophytenlora von Chile. Revue Bryologique et Lichénologique 29 (3/4): 183–206. Herzog T, Grolle R (1958) Was ist Pachyglossa? Revue Bryologique et Lichénologique 27 (3/4): 147–165. Herzog T, Hosseus CC (1938) Contribución al conocimiento de la lora brióita del sur de Chile. Archivos de la Escuela de Farmacia de la Facultad de Ciencias Médicas de Córdoba 7: 1–95. 728 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Herzog T, Noguchi A (1955) Beitrag zur Kenntnis der Bryophytenlora von Formosa und den benachbarten Inseln Botel Tobago und Kwashyoto. Journal of the Hattori Botanical Laboratory 14: 29–70. Herzog T, hériot I, Buch H, Schifner V, Verdoorn F (1933) Bryophyta nova (6-16). Annales Bryologici 6: 124–135. Heslewood MM, Brown EA (2007) A molecular phylogeny of the liverwort family Lepidoziaceae Limpr. in Australasia. Plant Systematics and Evolution 265 (3/4): 193–219. doi: 10.1007/ s00606-006-0512-z Hewson HJ (1970a) he family Aneuraceae in Australia and New Guinea: II. he genus Riccardia. Proceedings of the Linnean Society of New South Wales (ser. 2) 95 (1): 60–121. Hewson HJ (1970b) he family Aneuraceae in Australia and New Guinea: I. he genus Aneura. Proceedings of the Linnean Society of New South Wales (ser. 2) 94 (2): 173–193. Hewson HJ (1982) Vandiemeniaceae: a new family in the Metzgeriales. Journal of the Hattori Botanical Laboratory 52: 163–165. Hicks ML (1991) Scapania in Queensland, Australia. Journal of the Hattori Botanical Laboratory 69: 129–132. Hiepko P, Schultze-Motel W (1981) Floristische und ethnobotanische Untersuchungen im Eipomek-Tal, Irian Jaya (West-Neuguinea), Indonesien. Schriftenreihe Mensch, Kultur und Umwelt im zentralen Bergland West-Neuguinea 7: 1–75. Higuchi M (1998) A new species of Apotreubia (Treubiaceae, Hepaticae) from China. Cryptogamie: Bryologie, Lichénologie 19 (4): 321–328. Hill J (1773) A general natural history. Volume 2. A history of plants. Edition 2. London, 644 pp. Hodgetts NG (2008) A morphological revision of the genus Herbertus S.Gray (Herbertaceae, Marchantiophyta) in Africa, including the East African islands. Journal of Bryology 30 (4): 239–263. doi: 10.1179/174328208X300714 Hodgson EA (1941) Review of the New Zealand species of Schistochila, with notes on Colenso’s species. Transactions and Proceedings of the Royal Society of New Zealand 71 (3): 181–194. Hodgson EA (1943) A review of the New Zealand species of the genus Chiloscyphus. Transactions and Proceedings of the Royal Society of New Zealand 73 (1): 27–52. Hodgson EA (1944) A review of New Zealand species of Plagiochila (New Zealand hepaticae III). Transactions and Proceedings of the Royal Society of New Zealand 73 (4): 270–296. Hodgson EA (1949) New Zealand hepaticae (liverworts). VI. A review of the New Zealand species of the genus Frullania. Transactions and Proceedings of the Royal Society of New Zealand 77 (3): 361–389. Hodgson EA (1952) New Zealand hepaticae (liverworts) – VII. A review of the New Zealand species of the genus Lophocolea with notes on Chiloscyphus. Transactions of the Royal Society of New Zealand 80 (3/4): 329–358. Hodgson EA (1954) New Zealand hepaticae (liverworts) – VIII. A review of the New Zealand species of the genera Bazzania and Acromastigum. Transactions of the Royal Society of New Zealand 82 (1): 7–24. World checklist of hornworts and liverworts 729 Hodgson EA (1956) New Zealand hepaticae (liverworts) – IX. A review of the New Zealand species of the genus Lepidozia. Transactions of the Royal Society of New Zealand 83 (4): 589–620. Hodgson EA (1958) New Zealand hepaticae (liverworts) – X. Marsupial genera of New Zealand. Transactions of the Royal Society of New Zealand 85 (4): 565–584. Hodgson EA (1959) New Zealand hepaticae (liverworts) – XI. A review of New Zealand species of the genus Lepidolaena. Transactions of the Royal Society of New Zealand 87 (3/4): 199–210. Hodgson EA (1962a) Hepatics from the subantarctic islands of New Zealand including “Cape Expedition” collections from the Auckland and Campbell Islands. Records of the Dominion Museum 4 (11): 101–132. Hodgson EA (1962b) New Zealand hepaticae (liverworts). XIV. Drucella, a new genus of the Lepidoziaceae. Transactions of the Royal Society of New Zealand. Botany 2 (3): 45–47. Hodgson EA (1964) New Zealand hepaticae (liverworts) – XV: A new monotypic family of the thalloid Hepaticae Phyllothalliaceae Hodgson fam. nov. Transactions of the Royal Society of New Zealand. Botany 2 (19): 247–250. Hodgson EA (1965) New Zealand hepaticae (liverworts) – XVI. A miscellany of new genera, new species and notes, Part I. Transactions of the Royal Society of New Zealand. Botany 3 (4): 67–97. Hodgson EA (1967) New Zealand hepaticae (liverworts) – XVII A miscellany of taxonomic notes. Part. 2. Transactions of the Royal Society of New Zealand. Botany 3 (11): 175–198. Hodgson EA (1970) New Zealand hepaticae (liverworts) – XIX. Adelanthus Mitten, a new New Zealand genus. Transactions of the Royal Society of New Zealand. Biological Sciences 11 (18): 239–242. Hodgson EA (1972) New Zealand hepaticae (liverworts) – XX: A miscellany taxonomic notes, Part 3. Journal of the Royal Society of New Zealand 2 (1): 109–118. doi: 10.1080/03036758.1972.10423309 Hodgson EA, Allison KW (1962) New Zealand hepaticae (liverworts). XIII. A review of the New Zealand species of the genus Temnoma and Anoplostoma, a new genus. Transactions of the Royal Society of New Zealand. Botany 1 (12): 139–154. Hodgson EA, Sainsbury GOK (1948) Bryophytes collected by G. E. Du Rietz on the Antipodes Islands. Svensk Botanisk Tidskrift 42 (3): 273–280. Hofman G (1935) Monographische Studien über die indomalayischen Arten von Pycnolejeunea. Annales Bryologici 8: 80–129. Hofmann GF (1795) Deutschlands Flora. Cryptogamie. Johan Jakob Palm, Erlangen, 200 pp. Hong WS (1980) he genus Scapania in western North America. II. Taxonomic treatment. Bryologist 83 (1): 40–59. doi: 10.2307/3242392 Hong WS (1988) he family Lepidoziaceae in North America west of the hundreth meridian. Bryologist 91 (4): 326–333. doi: 10.2307/3242771 Hooker WJ (1812) British Jungermanniae, part 1–8. Longmans, London, tab. 1–32. Hooker WJ (1813) British Jungermanniae, part. 9–15. Longmans, London, tab. 33–60. Hooker WJ (1814) British Jungermanniae, part. 16–17. Longmans, London, tab. 61–68. Hooker WJ (1815) British Jungermanniae, part. 18–19. Longmans, London, tab. 69–76. Hooker WJ (1816a) British Jungermanniae, part. 20–22. Longmans, London, tab. 77–84. 730 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Hooker WJ (1816b) Plantae cryptogamicae. IDC, Leiden, 4 pp. Hooker WJ (1818) Musci exotici, vol. I. Longmans, London, 96 tab. doi: 10.5962/bhl.title.10721 Hooker WJ (1820) Musci exotici, vol. II. Longmans, London, 176 tab. doi: 10.5962/bhl. title.10721 Hooker WJ (1830) Botanical miscellany, vol. I. John Murray, London, 1–236. Hooker WJ (1831) Botanical miscellany, vol. II. John Murray, London, 129–421. Hooker JD (1867) Handbook of the New Zealand lora, part II. Reeve, London, 393–798. Hooker WJ, Arnott GAW (1832) he botany of Captain Beechey’s voyage, part. 3. Henry G Bohn, London, 97–144. doi: 10.5962/bhl.title.246 Hooker JD, Taylor T (1844a) Hepaticae Antarcticae. I. Species of Lord Auckland’s and Campbell’s Islands. London Journal of Botany 3: 366–400. Hooker JD, Taylor T (1844b) Hepaticae Antarcticae. II. Species of the Falkland Islands, Cape Horn, and of Kerguelen’s Land. London Journal of Botany 3: 454–481. Hooker JD, Taylor T (1844c) Hepaticae Novae-Zelandiae et Tasmaniae. IV. Species Tasmaniae. London Journal of Botany 3: 577–582. Hooker JD, Taylor T (1844d) Hepaticae Novae-Zelandiae et Tasmaniae. III. Species of New Zealand. London Journal of Botany 3: 556–577. Hooker JD, Taylor T (1845) Hepaticae Antarcticae, supplementum. London Journal of Botany 4: 79–97. Hooker WJ, Wilson W (1844) Enumeration of the mosses and hepaticae. London Journal of Botany 3: 149–167. Horikawa Y (1929a) Studies on the hepaticae of Japan. II. Science Reports of the Tôhoku Imperial University. Series 4, Biology 4 (2): 395–429. Horikawa Y (1929b) Studies on the hepaticae of Japan. I. Science Reports of the Tôhoku Imperial University. Series 4, Biology 4 (1): 37–72. Horikawa Y (1929c) Studies on the hepaticae of Japan. III. Science Reports of the Tôhoku Imperial University. Series 4, Biology 5 (4): 623–650. Horikawa Y (1931a) Studies on the hepaticae of Japan. V. Journal of Science of the Hiroshima University: Series B, Division 2 (Botany) 1: 55–76. Horikawa Y (1931b) Studies on the hepaticae of Japan. IV. Journal of Science of the Hiroshima University: Series B, Division 2 (Botany) 1: 13–35. Horikawa Y (1932a) Studies on the hepaticae of Japan. VI. Journal of Science of the Hiroshima University: Series B, Division 2 (Botany) 1: 77–94. Horikawa Y (1932b) Die epiphyllen Lebermoose von Japan. Botanical Magazine, Tokyo 46 (544): 176–184. doi: 10.15281/jplantres1887.46.176 Horikawa Y (1932c) Studies on the hepaticae of Japan. VII. Journal of Science of the Hiroshima University: Series B, Division 2 (Botany) 1: 121–134. Horikawa Y (1933) Studies on the hepaticae of Japan. VIII. Journal of Science of the Hiroshima University: Series B, Division 2 (Botany) 1: 197–205. Horikawa Y (1934) Monographia hepaticarum australi-japonicarum. Journal of Science of the Hiroshima University: Series B, Division 2 (Botany) 2: 101–325. Horikawa Y (1935) Contributions to the bryological lora of eastern Asia II. Journal of Japanese Botany 11: 499–508. World checklist of hornworts and liverworts 731 Horikawa Y (1936) Symbolae lorae bryophytae Orientali-Asiae et Micronesiae X. Botanical Magazine, Tokyo 50 (598): 556–561. doi: 10.15281/jplantres1887.50.556 Horikawa Y (1939) Contributions to the bryological lora of Eastern Japan. VI. Journal of Japanese Botany 15: 359–368. Horikawa Y (1943) Notulae hepaticologicae. Acta Phytotaxonomica et Geobotanica 13: 212–214. Horikawa Y (1951a) Tropical bryophytes in the Japanese Archipelago. Journal of Science of the Hiroshima University: Series B, Division 2 (Botany) 6: 27–38. Horikawa Y (1951b) Distributions of important bryophytes in the Japanese Archipelago (1). Hikobia 1 (2): 100–108. Horikawa Y (1951c) Symbolae lorae bryophytae orientali-asiae et micronesiae. XII. Hikobia 1 (2): 78–100. Horikawa Y, Ando H (1954) Taxonomic and ecological studies of Jubula in the Japanese archipelago. Journal of Science of the Hiroshima University: Series B, Division 2 (Botany) 6: 297–314. Hornemann JW (1810) Flora Danica vol. 8 heft 24 (tab. MCCCLXXXI-MCCCCXL). E. A. H. Mölleri Aulae Regiae & Universitatis Typographi, Haunie [Copenhagen], 1–8. Hornemann JW (1830) Flora Danica vol. 12 heft 34 (tab. MDCCCCLXXXI-MMXL). E. A. H. Mölleri Aulae Regiae & Universitatis Typographi, Haunie [Copenhagen], 1–14. Howe MA (1894) Notes on Californian bryophytes.–1. Erythea 2 (6): 97–101. Howe MA (1897a) Gyrothyra, a new genus of hepaticae. Bulletin of the Torrey Botanical Club 24 (4): 201–205. doi: 10.2307/2478334 Howe MA (1897b) he North American species of Porella. Bulletin of the Torrey Botanical Club 24 (11): 512–528. doi: 10.2307/2478073 Howe MA (1898a) he Anthocerotaceae of North America. Bulletin of the Torrey Botanical Club 25 (1): 1–24. doi: 10.2307/2478007 Howe MA (1898b) New American hepaticae. Bulletin of the Torrey Botanical Club 25 (4): 183–192. doi: 10.2307/2477701 Howe MA (1899) he hepaticae and anthocerotae of California. Memoirs of the Torrey Botanical Club 7: 1–208. Howe MA (1901a) Riccia beyrichiana and Riccia dictyospora. Bulletin of the Torrey Botanical Club 28 (3): 161–165. doi: 10.2307/2477946 Howe MA (1901b) An enumeration of the hepaticae collected by R. S. Williams, 1898–1899. Contribution to the botany of the Yukon Territory. Bulletin of the New York Botanical Garden 2 (6): 101–105. Howe MA (1902) Notes on North American hepaticae. Bulletin of the Torrey Botanical Club 29 (5): 281–289. doi: 10.2307/2478722 Howe MA (1917) Notes on North American species of Riccia. Bryologist 20 (3): 33–36. doi: 10.2307/3237525 Howe MA (1934) he hepaticae (chiely Riccia and Anthocerotaceae) of the Galapagos Islands and the coast and islands of Central America and Mexico. Proceedings of the California Academy of Sciences (ser. 4) 21 (17): 199–209. 732 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Howe MA, Underwood LM (1903) he genus Riella, with descriptions of new species from North America and the Canary Islands. Bulletin of the Torrey Botanical Club 30 (4): 214–224. doi: 10.2307/2478779 Hübener JWP (1832) Beschreibungen einiger neuen deutschen Jungermannien. Flora 15 (20): 305–308. Hübener JWP (1834) Hepaticologia Germanica. Schwann and Götz, Mannheim, 314 pp. Hudson W (1762) Flora Anglica, ed. 1. J. Nourse, London, 506 pp. Hudson W (1778) Flora Anglica, ed. 2, tom. 2. J. Nourse, London, 335–690. Hürlimann H (1960) Hepaticae aus dem Gebiete des südlichen Paziik. I. Bauhinia 1 (3): 251–260. Hürlimann H (1968) Hepaticae aus dem Gebiete des südlichen Paziik. II. Bauhinia 4 (1): 73–84. Hürlimann H (1976) Hepaticae aus dem Gebiete des südlichen Paziik. IV. Bauhinia 5 (4): 191–213. Hürlimann H (1983) Hepaticae aus dem Gebiete des südlichen Paziik. VII. Bauhinia 7 (4): 259–268. Hürlimann H (1985) Hepaticae aus dem Gebiete des südlichen Paziik. VIII. Bauhinia 8 (2): 101–118. Hürlimann H (1986) Riccia hawaiiensis Hürl., species nova. Phytologia 61 (5): 339–342. Hürlimann H (1987) Hepaticae aus dem Gebiete des südlichen Paziik. IX. Bauhinia 8 (4): 221–234. Hürlimann H (1989) Hepaticae aus dem Gebiete des südlichen Paziik. X. Bauhinia 9 (2): 153–170. Hürlimann H (1991) Hepaticae aus dem Gebiete des südlichen Paziik. XI. Bauhinia 9 (4): 257–264. Hürlimann H (1993) Hepaticae aus dem Gebiete des südlichen Paziik. XII. Bauhinia 11 (1): 3–17. Hürlimann H (1995) Hepaticae aus dem Gebiete des südlichen Paziik. XIII. Bauhinia 11 (3): 159–175. Hürlimann H (1998) Hepaticae aus dem Gebiete des südlichen Paziik. XIV. Bauhinia 12 (1/2): 109–119. Hürlimann H, Yamada K (1979) A new species of Radula (Hepaticae) from New Caledonia. Journal of Japanese Botany 54 (8): 238–240. Husnot T (1875) Hepaticologia gallica, fasc. 1. F. Savy, Paris, 1–30. Ilkiu-Borges AL (2002) Pictolejeunea reginae, a new species of Lejeuneaceae (Hepaticae) from Venezuela. Brittonia 54 (4): 318–321. doi: 10.1663/0007-196X(2003)54[318:PRANSO ]2.0.CO;2 Ilkiu-Borges AL (2005) A taxonomic revision of Echinocolea (Lejeuneaceae, Hepaticae). Nova Hedwigia 80 (1/2): 45–71. doi: 10.1127/0029-5035/2005/0080-0045 Ilkiu-Borges AL (2006) A taxonomic monograph of the genus Prionolejeunea (Lejeuneaceae, Jungermanniopsida). Cuvillier Verlag, Göttingen, 191 pp. Ilkiu-Borges AL (2011) On Pycnolejeunea gradsteinii (Lejeuneaceae), a new species from Brazil. Boletim do Instituto de Botânica. São Paulo 21 (1): 1–3. World checklist of hornworts and liverworts 733 Ilkiu-Borges AL, Pereira Alvarenga LD (2008) On Ceratolejeunea atlantica, a new species of Lejeuneaceae from Brazil. Nova Hedwigia 86 (1/2): 237–241. doi: 10.1127/00295035/2008/0086-0237 Ilkiu-Borges AL, Gradstein SR (2008) A new species of Cheilolejeunea (Spruce) Schifn. (Lejeuneaceae) from Cerro de la Neblina, Venezuela. Nova Hedwigia 87 (3/4): 521–528. doi: 10.1127/0029-5035/2008/0087-0521 Ilkiu-Borges AL, Schäfer-Verwimp A (2005) On Prionolejeunea grollei, a new species from the West Indies (Lejeuneaceae, Hepaticae). Cryptogamie, Bryologie 26 (1): 29–35. Infante M (2010) Notes on the genus Dendroceros in West Africa and south Atlantic Islands. Journal of Bryology 32 (4): 283–287. doi: 10.1179/037366810X12814321877426 Infante M, Heras P (1999) Bryophytes from the Republic of Equatorial Guinea (West Central Africa). VI. Cololejeunea iradieri sp. nov., Cololejeunea magna stat. nov. and their relations to similar species. Tropical Bryology 17: 13–18. Infante M, Heras P, Pócs T (1999) Bryophytes from the Republic of Equatorial Guinea (West Central Africa). V. Diplasiolejeunea cogoensis sp. nov. Tropical Bryology 17: 9–12. Ingham W (1904) Scapania calcicola, a new British hepatic. Naturalist. London, Huddersield, Hull, etc. 564: 11–12. Inoue H (1956) A new and additional descriptions of hepaticae from Titibu district. Journal of Japanese Botany 31 (11): 340–344. Inoue H (1958) he family Plagiochilaceae of Japan and Formosa. II. Journal of the Hattori Botanical Laboratory 20: 54–106. Inoue H (1959a) Hepatics from Isl. Palau, Caroline. Journal of Japanese Botany 34 (9): 267–271. Inoue H (1959b) On Metacalypogeia, a new genus of Hepaticae. Journal of the Hattori Botanical Laboratory 21: 231–235. Inoue H (1959c) A review of Japanese species of Lophocolea Dum. Journal of the Hattori Botanical Laboratory 21: 214–230. Inoue H (1960) Contributions to the knowledge of Plagiochilaceae of southern Asia. II. On some Himalayan species of Plagiochila. Journal of the Hattori Botanical Laboratory 23: 29–36. Inoue H (1961a) A new Lejeunea (Hepaticae) from Japan. Journal of Japanese Botany 36 (1): 7–10. Inoue H (1961b) An interesting new species of Lophozia from limestone districts of ChichibuOkutama mountains. Journal of Japanese Botany 36 (2): 41–45. Inoue H (1962a) Notes on Plagiochila eatonii Aust., a Hawaiian liverwort. Journal of Japanese Botany 37 (12): 357–360. Inoue H (1962b) Miscellaneous notes on hepatics of Japan (3). Journal of Japanese Botany 37 (4): 101–105. Inoue H (1962c) Two new species of Plagiochila Dum. Journal of Japanese Botany 37 (6): 187–190. Inoue H (1963) Contributions to the knowledge of the Plagiochilaceae of southeastern Asia. IV. he genus Xenochila. Bulletin of the National Science Museum, Tokyo (n.ser.) 6 (4): 372–377. Inoue H (1964a) Miscellaneous notes on hepatics of Japan (4). Journal of Japanese Botany 39 (4): 105–108. 734 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Inoue H (1964b) he genus Plagiochilion. Journal of the Hattori Botanical Laboratory 27: 51–72. Inoue H (1965a) Contributions to the knowledge of the Plagiochilaceae of southeastern Asia. VII. Synopsis of Plagiochila Dum. in the Himalayan region. Bulletin of the National Science Museum, Tokyo (n.ser.) 8 (3): 375–403. Inoue H (1965b) Some nomenclatural changes and synonyms in the genera Syzygiella and Plagiochila. Bryologist 68 (2): 217–219. doi: 10.2307/3241017 Inoue H (1965c) Contributions to the knowledge of the Plagiochilaceae of southern Asia. VI. Studies on the Plagiochila semidecurrens complex. Journal of the Hattori Botanical Laboratory 28: 209–218. Inoue H (1966a) New and noteworthy liverworts from Formosa. Bulletin of the National Science Museum, Tokyo (n.ser.) 9 (1): 37–41. Inoue H (1966b) Contributions to the knowledge of Plagiochilaceae of southeastern Asia, IX. Plagiochila autoica Steph. with a review of Pedinophyllum. Bulletin of the National Science Museum, Tokyo (n.ser.) 9 (4): 571–575. Inoue H (1966c) A monograph of the hepatic genus Syzygiella Spruce. Journal of the Hattori Botanical Laboratory 29: 171–213. Inoue H (1966d) Miscellaneous notes on hepatics of Japan (5). Journal of Japanese Botany 41 (1): 14–16. Inoue H (1967a) Studies on oil bodies of some Malayan liverworts. Journal of the Hattori Botanical Laboratory 30: 54–70. Inoue H (1967b) Asterella shimizuana, a new liverwort from New Guinea. Bulletin of the National Science Museum, Tokyo (n.ser.) 10 (3): 361–363. Inoue H (1967c) hree new hepatics from Malay Peninsula. Bulletin of the National Science Museum, Tokyo (n.ser.) 10 (2): 155–162. Inoue H (1967d) Taxonomic miscellany on the Plagiochilaceae. Journal of Japanese Botany 42 (6): 182–187. Inoue H (1968a) Taxonomic miscellany on the genera Plagiochila and Syzygiella. Bulletin of the National Science Museum, Tokyo (n.ser.) 11 (3): 299–307. Inoue H (1968b) A new species of Calypogeia Raddi. Journal of Japanese Botany 43 (10/11): 468–472. Inoue H (1969) Contributions to the knowledge of Plagiochilaceae of southern Asia XIII. New species of Plagiochila with notes on sect. Abietinae and sect. Peculiares. Journal of the Hattori Botanical Laboratory 32: 99–115. Inoue H (1970a) Novae Guineae hepaticae Schusteranae. II. Plagiochilae species novae. Journal of the Hattori Botanical Laboratory 33: 317–330. Inoue H (1970b) Plagiochilaceae notes, I. On some species of Plagiochila from South Paciic Islands. Journal of the Hattori Botanical Laboratory 33: 305–316. Inoue H (1971a) Notes on the Calypogeia neesiana complex in Japan. Memoirs of the National Science Museum 4: 55–58. Inoue H (1971b) Nardia lagelliformis (sp. nov.) with reference to the subgeneric and sectional revision of the genus Nardia Gray. Journal of Japanese Botany 46 (1): 1–7. World checklist of hornworts and liverworts 735 Inoue H (1971c) Contributions to the knowledge of the Plagiochilaceae of southern Asia XVII. A new species of Plagiochilion from Sumatra. Bulletin of the National Science Museum, Tokyo (n.ser.) 14 (2): 269–272. Inoue H (1972a) Miscellaneous notes on hepatics of Japan (7). Journal of Japanese Botany 47 (11): 347–350. Inoue H (1972b) Four new species of Plagiochila from New Guinea. Journal of the Hattori Botanical Laboratory 36: 487–496. Inoue H (1972c) Plagiochilaceae notes part 3. Plagiochila major and Plagiochila rhizophora in the paciic northwest of North America. Bulletin of the National Science Museum, Tokyo (n.ser.) 15 (1): 181–190. Inoue H (1973) he genus Fossombronia Raddi in Japan. Journal of the Hattori Botanical Laboratory 37: 293–297. Inoue H (1974a) Contributions to the knowledge of the Plagiochilaceae of southeastern Asia. XV. Enumeration of Plagiochila species from hailand. Journal of the Hattori Botanical Laboratory 38: 555–563. Inoue H (1974b) Temnoma setigerum (Lindenb.) Schust. from the Hawaiian Islands. Bulletin of the National Science Museum, Tokyo (n.ser.) 17 (3): 227–231. Inoue H (1975a) Notes on the Plagiochilaceae, V. Descriptions of some new species of Plagiochila Dum. Bulletin of the National Science Museum, Tokyo. Series B, Botany 1 (3): 83–100. Inoue H (1975b) Two new species of Calypogeia Raddi from Japan. Bulletin of the National Science Museum, Tokyo. Series B, Botany 1 (4): 135–140. Inoue H (1976a) Illustrations of Japanese hepaticae. 2. Tokyo, 193 pp. Inoue H (1976b) Notes on the Plagiochilaceae, V. Studies in the genus Plagiochila (Dum.) Dum. in the Hawaiian Islands. Journal of the Hattori Botanical Laboratory 40: 411–440. Inoue H (1976c) Contributions to the knowledge of the Plagiochilaceae of south-eastern Asia, part 16. On Plagiochila hattori new species from Japan. Bulletin of the National Science Museum, Tokyo. Series B, Botany 2 (3): 69–75. Inoue H (1977a) Notes on the Plagiochilaceae. VIII. Two new species of the genus Plagiochila (Dum.) Dum. from Mexico. Bulletin of the National Science Museum, Tokyo. Series B, Botany 3 (4): 135–142. Inoue H (1977b) A remarkable new species of the genus Lejeunea Libert from southern Japan. Bulletin of the National Science Museum, Tokyo. Series B, Botany 3 (4): 142–148. Inoue H (1977c) Notes on the Plagiochilaceae. VII. Four new species of the genus Plagiochila (Dum.) Dum. from the Galápagos Islands. Bulletin of the National Science Museum, Tokyo. Series B, Botany 3 (2): 45–54. Inoue H (1978a) Studies on Taiwan hepaticae. III. Subord. Herbertinae and subord. Ptilidiinae. Bulletin of the National Science Museum, Tokyo. Series B, Botany 4 (3): 93–100. Inoue H (1978b) Anastrophyllum ellipticum Inoue (sp. nov., Hepaticae) from Japan. Bulletin of the National Science Museum, Tokyo. Series B, Botany 4 (1): 13–17. Inoue H (1979a) Contributions to the knowledge of the Plagiochilaceae of southeastern Asia. XVIII. A review of the genus Plagiochila (Dum.) Dum. in Ceylon. Journal of the Hattori Botanical Laboratory 46: 193–233. 736 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Inoue H (1979b) hree new hepatics from Papua New Guinea. In: Kurokawa S (Ed.) Studies on Cryptogams of Papua New Guinea. Academia Scientiic Book, Tokyo, 11–18. Inoue H (1979c) Contributions to the knowledge of the Plagiochilaceae of southeastern Asia. XVII. Descriptions of ive new species of Plagiochila (Dum.) Dum. Bulletin of the National Science Museum, Tokyo. Series B, Botany 5 (1): 25–36. Inoue H (1980) A new species of Plagiochila (Hepaticae) from Tasmania. Brunonia 3 (1): 141–144. doi: 10.1071/BRU9800141 Inoue H (1982) Studies on Taiwan hepaticae. IV. Plagiochilaceae. Bulletin of the National Science Museum, Tokyo. Series B, Botany 8 (4): 125–144. Inoue H (1984a) Notes on the Plagiochilaceae. XII. Notes on some species of Plagiochila (Dum.) Dum. from southern Chile and Argentina. In: Inoue H (Ed.) Studies on Cryptogams in Southern Chile. Kensei-sha, Tokyo, 97–108. Inoue H (1984b) he genus Plagiochila (Dum.) Dum. in southeast Asia. Academia Scientiic Book, Tokyo, 142 pp. Inoue H (1985) Studies on Pallaviciniaceae and Allisoniaceae (Metzgeriales) in Japan. 2. he genus Moerckia Gott. Bulletin of the National Science Museum, Tokyo. Series B, Botany 11 (1): 1–10. Inoue H (1986) Contributions to the knowledge of the Plagiochilaceae of southeastern Asia. XIX. he genus Plagiochila from Isl. Seram. Bulletin of the National Science Museum, Tokyo. Series B, Botany 12 (3): 73–84. Inoue H (1987a) Steereochila, a new genus of the Plagiochilaceae from the neotropics. Memoirs of the New York Botanical Garden 45: 279–282. Inoue H (1987b) Contributions to the knowledge of the Plagiochilaceae of Southeastern Asia XX. Studies on collections made by Dr. D. G. Long in Bhutan. Bulletin of the National Science Museum, Tokyo. Series B, Botany 13 (2): 41–51. Inoue H (1987c) Notes on the Plagiochilaceae. XIV. New species of Plagiochila (Dum.) Dum. from the neotropics. In: Inoue H (Ed.) Studies on the Cryptogams in Southern Peru. Tokai University Press, Tokyo, 95–105. Inoue H (1988a) Two new species of Plagiochila from the neotropics. Beihefte zur Nova Hedwigia 90: 171–175. Inoue H (1988b) Notes on the Plagiochilaceae, XV. he gneus [sic] Steereochila Inoue and Plagiochila eggersii, sp. nov. Bulletin of the National Science Museum, Tokyo. Series B, Botany 14 (4): 135–141. Inoue H (1988c) Two new species of Plagiochila from Papua New Guinea. Journal of Japanese Botany 63 (11): 365–369. Inoue H (1989a) Two new species of Plagiochila (Dum.) Dum. from Panamá, Central America. Bulletin of the National Science Museum, Tokyo. Series B, Botany 15 (3): 91–96. Inoue H (1989b) he bryophytes of Sabah (north Borneo) with special reference to the BRYOTROP transect of Mount Kinabalu V. Plagiochila (Plagiochilaceae, Hepaticae). Willdenowia 18 (2): 555–567. Inoue H (1989c) Taxonomic miscellany on hepatics 6. Journal of Japanese Botany 64 (7): 193–198. World checklist of hornworts and liverworts 737 Inoue H, Gradstein SR (1988) A remarkable new species of Anastrophyllum (Spruce) Steph. (Hepaticae) from Mt. Roraima, Guyana. Bulletin of the National Science Museum, Tokyo. Series B, Botany 14 (3): 87–91. Inoue H, Miller HA (1965) Hepaticae from Kusaie, Caroline Islands. Bulletin of the National Science Museum, Tokyo (n.ser.) 8 (2): 139–160. Inoue H, Miller HA (1968) Noteworthy hepaticae from Micronesia. Bulletin of the National Science Museum, Tokyo (n.ser.) 11 (1): 1–12. Inoue H, Miller NG (1985) A new Aneura Dum. (Hepaticae, Aneuraceae) from eastern North America. Bulletin of the National Science Museum, Tokyo. Series B, Botany 11 (3): 95–101. Inoue H, Schuster RM (1971) A monograph of New Zealand and Tasmanian Plagiochilaceae. Journal of the Hattori Botanical Laboratory 34: 1–225. Inoue H, Yang B-Y (1966) he genus Mylia Gray in Taiwan. Taiwania 12 (1): 35–38. Inuthai J, Zhu R-L, Chantanaorrapint S (2014) Drepanolejeunea actinogyna (Lejeuneaceae), a new species from southern hailand. Bryologist 117 (2): 165–169. doi: 10.1639/00072745-117.2.165 Iwatsuki Z, Hattori S (1956) Studies of the epiphytic moss lora of Japan. 2. he bryophytic community on Machilus japonica in a dense hard-wood forest in a valley at Inohae in southern Kyūshū. Journal of the Hattori Botanical Laboratory 16: 83–90. Jack JB (1881) Die Europäischen Radula-Arten (Schluss). Flora 64 (25): 385–400. Jack JB (1886) Monographie der Lebermoosgattung Physiotium. Hedwigia 25 (2/3): 49–87. Jack JB (1894) Stephaniella paraphyllina Jack nov. gen. hepaticarum. Hedwigia 33 (1): 11–14. Jack JB, Stephani F (1892) Hepaticae Wallisianae. Hedwigia 31 (1): 11–27. Jack JB, Stephani F (1894) Hepaticae in insulis Vitiensibus et Samoanis a Dre Ed. Graefe anno 1864 lectae. Botanisches Centralblatt 60 (4): 97–109. Jack JB, Stephani F (1895) Hepaticae Lorentzianae. Hedwigia 34 (6): 313–318. Jacobs DL (1949) Two new Riccias from Georgia. Bryologist 52 (4): 167–172. doi: 10.2307/3239471 Jelenc F (1950) Muscinées de l’Afrique du Nord (suite). Bulletin Trimestriel de Geographie et d’Archeologie [Oran] 73 (228): 59–89. Jensen CEO (1915) Danmarks mosser. I. Hepaticales, Anthocerotales og Sphagnales. Gyldendal, Copenhagen, 317 pp. Jones EW (1953a) African hepatics. III. Cololejeunea and Leptocolea with dentate leaves. Transactions of the British Bryological Society 2 (2): 158–163. doi: 10.1179/006813853804878191 Jones EW (1953b) African hepatics. II. Leptocolea with hyaline-margined leaves. Transactions of the British Bryological Society 2 (2): 144–157. doi: 10.1179/006813853804878137 Jones EW (1954a) African hepatics. VIII. Diplasiolejeunea albifolia (Taylor) E. W. Jones comb. nov. Transactions of the British Bryological Society 2 (3): 393–395. doi: 10.1179/006813854804830075 Jones EW (1954b) African hepatics VII. he genus Cheilolejeunea. Transactions of the British Bryological Society 2 (3): 380–392. doi: 10.1179/006813854804830129 Jones EW (1954c) African hepatics. X. Leptocolea and Cololejeunea. Transactions of the British Bryological Society 2 (3): 408–438. doi: 10.1179/006813854804830156 738 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Jones EW (1956) African hepatics. XI. he genus Riccardia in tropical Africa. Transactions of the British Bryological Society 3 (1): 74–84. doi: 10.1179/006813856804829741 Jones EW (1957a) African hepatics. XIII. he Ricciaceae in tropical Africa. Transactions of the British Bryological Society 3 (2): 208–227. doi: 10.1179/006813857804829524 Jones EW (1957b) African hepatics. XII. Some little-known Lejeuneaceae. Transactions of the British Bryological Society 3 (2): 191–207. doi: 10.1179/006813857804829533 Jones EW (1958) African hepatics. XIV. Some Cephaloziellas of lowland tropical Africa. Transactions of the British Bryological Society 3 (3): 430–440. doi: 10.1179/006813858804829398 Jones EW (1962) African hepatics XV. Plagiochila in Tropical Africa. Transactions of the British Bryological Society 4 (2): 254–325. doi: 10.1179/tbbs.1962.4.2.254 Jones EW (1963) African hepatics. XVI. Porella in tropical Africa. Transactions of the British Bryological Society 4 (3): 446–461. doi: 10.1179/006813863804812327 Jones EW (1964) African hepatics. XVII. Gongylanthus in tropical Africa. Transactions of the British Bryological Society 4 (4): 649–652. doi: 10.1179/006813864804812137 Jones EW (1968) African hepatics. XIX. he Lejeunea lava complex. Transactions of the British Bryological Society 5 (3): 548–562. doi: 10.1179/006813868804146863 Jones EW (1969) African hepatics. XXI. Microlejeunea, Chaetolejeunea and Pleurolejeunea. Transactions of the British Bryological Society 5 (4): 775–789. doi: 10.1179/006813869804146637 Jones EW (1972) African hepatics. XXIII. Some species of Lejeunea. Journal of Bryology 7 (1): 23–45. doi: 10.1179/jbr.1972.7.1.23 Jones EW (1973) African hepatics. XXIV. Lejeuneaceae: some new or little-known species and extensions of range. Journal of Bryology 7 (4): 545–561. doi: 10.1179/jbr.1973.7.4.545 Jones EW (1974) African hepatics. XXVI. he Lejeunea eckloniana complex. Journal of Bryology 8 (1): 77–91. doi: 10.1179/jbr.1974.8.1.77 Jones EW (1975) African hepatics. XXVII. Bazzania. Journal of Bryology 8 (3): 299–316. doi: 10.1179/jbr.1975.8.3.299 Jones EW (1976) African hepatics. XXIX. Some little-known species and extensions of range. Journal of Bryology 9 (1): 43–54. doi: 10.1179/jbr.1976.9.1.43 Jones EW (1977) African hepatics. XXX. he genus Radula Dumortier. Journal of Bryology 9 (4): 461–504. doi: 10.1179/jbr.1977.9.4.461 Jones EW (1979) African hepatics. XXXI. Rare or little-known Lejeuneaceae and extensions of range. Journal of Bryology 10 (4): 387–400. doi: 10.1179/jbr.1979.10.4.387 Jones EW (1982) African hepatics. XXXIII. Some new Lejeuneaceae. Journal of Bryology 12 (1): 37–48. doi: 10.1179/jbr.1982.12.1.37 Jones EW (1985) African hepatics. XXXIV. Little-known or new Lejeuneaceae. Journal of Bryology 13 (3): 385–398. doi: 10.1179/jbr.1985.13.3.385 Jones EW (1986) African hepatics. XXXV. Some new or little-known species and some noteworthy records. Journal of Bryology 13 (4): 497–508. doi: 10.1179/jbr.1985.13.4.497 Jones EW (1988) African hepatics XXXVIII. Cheilolejeunea subg. Strepsilejeunea (Spruce) Schust., with special reference to East Africa. Journal of Bryology 15 (1): 149–160. doi: 10.1179/jbr.1988.15.1.149 Jones EW (1989) African hepatics. XXXIX. Some dioecious species of Lejeunea. Journal of Bryology 15 (4): 665–673. doi: 10.1179/jbr.1989.15.4.665 World checklist of hornworts and liverworts 739 Jones EW (1992) African hepatics. XLII. Radula marojezica E.W.Jones, a new species from Madagascar. Journal of Bryology 17 (2): 307–311. doi: 10.1179/jbr.1992.17.2.307 Jones EW, Harrington AJ (1983) he hepatics of Sierra Leone and Ghana. Bulletin of the British Museum (Natural History), Botany 11 (3): 215–289. Jones EW, Pócs T (1987) African hepatics. XXXVI. hree new species of Colura. Journal of Bryology 14 (3): 495–501. Jørgensen E (1894) Om loraen i Nord-reisen og tilstødende dele af Lyngen. Forhandlinger i Videnskabs-Selskabet i Kristiania 1894 (8): 1–104. Jørgensen E (1922) Notiser til Norges levermoslora 1. Bergens Museums Aarbok 1919/20 (7): 1–6. Jørgensen E (1934) Norges levermossor. Bergens Museums Skrifter (n.ser.) 16: 1–343. Jovet-Ast S (1947a) A propos du Lepidozia Aubertii Jovet-Ast (=L. Wallichii St. ms.). Candollea 11: 31–35. Jovet-Ast S (1947b) Hépatiques des Antilles françaises récoltées par P. et V. Allorge en 1936. I. Revue Bryologique et Lichénologique 16 (1/2): 17–46. Jovet-Ast S (1948) Hépatiques des Antilles françaises récoltées par P. et V. Allorge en 1936 II. Revue Bryologique et Lichénologique 17 (1/4): 24–34. Jovet-Ast S (1949a) Hépatiques des Antilles françaises récoltées par P. et V. Allorge en 1936 III. Revue Bryologique et Lichénologique 18 (1/2): 35–42. Jovet-Ast S (1949b) Récoltes de J. T. Buchholz en Nouvelle-Calédonie. Revue Bryologique et Lichénologique 18 (1/2): 83. Jovet-Ast S (1951) Hépatiques des Nouvelles-Hebrides. Récoltes de E. Aubert de la Rüe, 1934. Revue Bryologique et Lichénologique 20 (1/2): 96–98. Jovet-Ast S (1953) Le genre Colura, Hépatiques, Lejeunéacées, Diplasiae. Revue Bryologique et Lichénologique 22 (2/3): 206–312. Jovet-Ast S (1954) Le genre Colura, Hépatiques, Lejeunéacées, Diplasiae (supplément). Revue Bryologique et Lichénologique 23 (1/2): 1–22. Jovet-Ast S (1955) Harpalejeunea herzogii S. J.-A., Lejeunéacée nouvelle de la Guadeloupe. Feddes Repertorium Specierum Novarum Regni Vegetabilis 58: 19–22. Jovet-Ast S (1956) Deux Colura nouveaux de Madagascar. Revue Bryologique et Lichénologique 25 (3/4): 272–276. Jovet-Ast S (1958) Hépatiques du Cambodge récoltées par le Professeur Roger Heim. Revue Bryologique et Lichénologique 27 (1/2): 24–30. Jovet-Ast S (1959) Un Microlejeunea nouveau de l’île de la Réunion. Revue Bryologique et Lichénologique 27 (3/4): 191–194. Jovet-Ast S (1960) Muscinées du Mexique récoltées par Roger Heim en 1956 et 1959. Revue Bryologique et Lichénologique 29 (1/2): 30–43. Jovet-Ast S (1961) Colura de Sud-Vietnam (Récoltes de Pierre Tixier). Revue Bryologique et Lichénologique 30 (1/2): 5–12. Jovet-Ast S (1965) Riccia crystallina L. emend. Raddi et Riccia cavernosa Hofm. emend. Raddi (Note préliminaire). Revue Bryologique et Lichénologique 33 (3/4): 459–483. Jovet-Ast S (1967a) Compléments à l’étude des Colura: localités nouvelles; description d’une espèce nouvelle de Borneo. Revue Bryologique et Lichénologique 35 (1/4): 143–148. 740 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Jovet-Ast S (1967b) Colura récoltes du Pakistan aux Philippines par Pierre Tixier. Revue Bryologique et Lichénologique 35 (1/4): 138–142. Jovet-Ast S (1976) Compléments à la connaissance des Colura: espèces et localités nouvelles. Revue Bryologique et Lichénologique 42 (4): 909–922. Jovet-Ast S (1978) Riccia des îles Galápagos. Revue Bryologique et Lichénologique 44 (4): 411–428. Jovet-Ast S (1980) La section Oidocrorys S. J.-A. du genre Colura Dum. est-elle agée de plus de 100 millions d’années? Cryptogamie: Bryologie, Lichénologie 1 (3): 277–287. Jovet-Ast S (1983) Colura nouveaux d’origines diverses (Hépatiques, Lejeunéacées). Cryptogamie: Bryologie, Lichénologie 4 (3): 205–216. Jovet-Ast S (1986) Les Riccia de la région méditerannéenne. Cryptogamie: Bryologie, Lichénologie 7 (3, suppl.): 283–431. Jovet-Ast S (1987) Une Riccia nouveau du Nord-est du Brésil: Riccia vitalii (Ricciaceae, sous-genre Riccia). Memoirs of the New York Botanical Garden 45: 283–288. Jovet-Ast S (1989) Un complexe de taxons dans le genre Riccia. Cryptogamie: Bryologie, Lichénologie 10 (2): 95–117. Jovet-Ast S (1991) Riccia (Hépatiques, Marchantiales) d’Amérique latine. Taxons du sousgenre Riccia. Cryptogamie: Bryologie, Lichénologie 12 (3): 189–370. Jovet-Ast S (1993a) Riccia (sous-genre Riccia) de l’île de la Réunion. Journal of the Hattori Botanical Laboratory 74: 95–104. Jovet-Ast S (1993b) Riccia L. (Hépatiques, Marchantiales) d’Amérique latine. Taxons des sous-genres hallocarpus, Leptoriccia, Ricciella. Cryptogamie: Bryologie, Lichénologie 14 (3): 219–301. Jovet-Ast S (1996) Riccia Triseriata subg. nov. et R. singularis sp. nov., taxons nouveaux d’Australie. Cryptogamie: Bryologie, Lichénologie 17 (2): 127–133. Jovet-Ast S (1997) Une Riccia prolifère d’Australie: Riccia pullulans sp. nov. (sous-genre Ricciella). Cryptogamie: Bryologie, Lichénologie 18 (3): 183–189. Jovet-Ast S (2000) Documents pour la connaissance des Riccia australiens (Hépatiques, Marchantiales) – nouvelles récoltes. Taxons nouveaux. Commentaires morphologiques et écologiques. Cryptogamie, Bryologie 21 (4): 289–343. doi: 10.1016/S12900796(00)01042-7 Jovet-Ast S (2003) Riccia des sous-genres Riccia et Ricciella récoltées en Indie et en Indonésie. Cryptogamie, Bryologie 24 (3): 209–228. Jovet-Ast S, Tixier P (1958) Hépatiques du Viet-Nam – I. Revue Bryologique et Lichénologique 27 (3/4): 201–210. Jovet-Ast S, Tixier P (1962) Hépatiques du Viet-Nam – II. Revue Bryologique et Lichénologique 31 (1/2): 23–33. Juslén A (2006a) Revision of Asian Herbertus (Herbertaceae, Marchantiophyta). Annales Botanici Fennici 43 (6): 409–436. Juslén A (2006b) Phylogeny of Vetaformaceae, Lepicoleaceae and Herbertaceae (including Mastigophoraceae) inferred from chloroplast trnL-F, nuclear ITS2, and morphology. Annales Botanici Fennici 43 (5): 349–362. Kaalaas B (1893a) Levermosernes udbredelse i Norge. Nyt Magazin for Naturvidenskaberne 33 (1): 1–192. World checklist of hornworts and liverworts 741 Kaalaas B (1893b) Levermosernes udbredelse i Norge. Nyt Magazin for Naturvidenskaberne 33 (4/5): 289–490. Kaalaas B (1896) Scapania gymnostomophila n. sp. Botaniska Notiser 49: 21–22. Kaalaas B (1898) Beiträge zur Lebermooslora Norwegens. Skrifter udgivne af Videnskabs-Selskabet i Christiana. Mathematisk-Naturvidenskabelig Klasse 1898 (9): 1–28. Kaalaas B (1902) Cephaloziae species duae novae. Revue Bryologique 29 (1): 8–10. Kaalaas B (1911) Bryophyten aus den Crozet-inseln. I. Nyt Magazin for Naturvidenskaberne 49 (2/3): 81–98. Kachroo P (1952) Notes on Assam hepaticae. A preliminary note. Journal of Gauhati India University 3: 129–131. Kachroo P (1958) Morphology of Rebouliaceae. III. Development of sex organs, sporangium and interrelationships of the various genera. Journal of the Hattori Botanical Laboratory 19: 1–24. Kachroo P (1968) History of the genus Cheilolejeunea. Ceylon Journal of Science. Biological Sciences 8 (1): 1–10. Kachroo P, Schuster RM (1961) he genus Pycnolejeunea and its ainities to Cheilolejeunea, Euosmolejeunea, Nipponolejeunea, Tuyamaella, Siphonolejeunea and Strepsilejeunea. Journal of the Linnean Society. Botany 56 (368): 475–511. doi: 10.1111/j.1095-8339.1961. tb02542.x Kachroo P, Bapna KR, Dhar GL (1977) Hepaticae of India – A taxonomic survey and census. V. Fossombroniaceae through anthocerotae. Journal of the Indian Botanical Society 56 (1): 62–86. Kahn SA (1955) Riccia perssonii S. A. Kahn: a new and interesting species from East Pakistan. Svensk Botanisk Tidskrift 49 (3): 433–436. Kamimura M (1955) Two new species of Lejeuneaceae Astipae from Japan. Feddes Repertorium Specierum Novarum Regni Vegetabilis 58: 55–59. Kamimura M (1961) A monograph of Japanese Frullaniaceae. Journal of the Hattori Botanical Laboratory 24: 1–109. Kamimura M (1962) On the genus Neohattoria Kamim. nom. nov. Journal of Japanese Botany 37 (7): 218. Kamimura M (1970) Some addenda to the Frullaniaceae of Japan and neighboring areas. I. Bulletin of Kochi Gakuen Junior College 1: 51–53. Kamimura M (1971) Some addenda to the Frullaniaceae of Japan and neighboring areas. II. Bulletin of Kochi Gakuen Junior College 2: 21–25. Kamimura M (1982) Some addenda to the Frullania of Japan and neighbouring areas III. Miscellanea Bryologica et Lichenologica 9 (4): 87–93. Kanwal HS (1979) A new species of Riccia L., R. udarii sp. nov. from Kumaon (Western Himalayas) India. Journal of the Indian Botanical Society 58 (3): 281–285. Kashyap SR (1914a) Morphological and biological notes on new and little known West Himalayan Liverworts. I. New Phytologist 13 (6/7): 206–226. doi: 10.1111/j.1469-8137.1914.tb05751.x Kashyap SR (1914b) Morphological and biological notes on new and little known West Himalayan liverworts. II. New Phytologist 13 (9): 308–323. doi: 10.1111/j.1469-8137.1914.tb05760.x 742 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Kashyap SR (1915) Morphological and biological notes on new and little known west Himalayan liverworts III. New Phytologist 14 (1): 1–18. doi: 10.1111/j.1469-8137.1915.tb07159.x Kashyap SR (1916) Liverworts of the western Himalayas and the Punjab, with notes on known species and descriptions of new species I. Journal of the Bombay Natural History Society 24 (2): 343–350. Kashyap SR (1917) Liverworts of the western Himalayas and the Punjab, with notes on known species and descriptions of new species II. Journal of the Bombay Natural History Society 25 (2): 279–281. Kashyap SR (1928) A new species of Petalophyllum, P. indicum Kash. Journal of the Indian Botanical Society 7: 14. Kashyap SR (1929) Liverworts of the western Himalayas and the Panjab plain, part 1. University of the Panjab, Lahore, 129 pp. Kashyap SR, Chopra RS (1932) Liverworts of the western Himalayas and the Panjab plain, part 2. University of the Panjab, Lahore, 137 pp. Kashyap SR, Dutt NL (1925) he genus Notothylas in India. Proceedings of the Lahore Philosophical Society 4: 49–56. Katagiri T (2013) A tropical American species, Trichocolea ilicaulis, new to Papua New Guinea. Hikobia 16 (3): 293–298. Katagiri T, Deguchi H (2012) Taxonomic studies of the Trichocoleaceae in southeast Asia I. he genus Leiomitra Lindb. Bryologist 115 (4): 475–492. doi: 10.1639/0007-2745115.4.475 Katagiri T, Yamaguchi T (2011) [Validation of Porella caespitans var. cordifolia]. Bryological Research 10 (5): 133–134. Katagiri T, Miyauchi H, Deguchi H (2011) Trichocolea japonica (Trichocoleaceae), a new species from Japan. Bryologist 114 (4): 744–748. doi: 10.1639/0007-2745-114.4.744 Katagiri T, Suleiman M, Deguchi H (2012) Taxonomic studies of the Trichocoleaceae in southeast Asia II. A new species of Eotrichocolea from Malaysia. Bryologist 115 (4): 518–522. doi: 10.1639/0007-2745-115.4.518 Katagiri T, Sadamitsu A, Miyauchi H, Tsubota H, Deguchi H (2013) Taxonomic studies of the Trichocoleaceae in Southeast Asia. Ill. he genus Trichocolea Dumort. Hattoria 4: 1–42. Kavina K (1915) Monograie českých jatrovek. Díl. I.Jatrovky lupenité (Hepaticae frondosae Bohemiae). Archiv pro Přírodovědecký výzkum Čech 16 (2): 1–284. Khan SA (1957) Studies in Ricciaceae of East Pakistan. I. New and little known species of Riccia. Bryologist 60 (1): 28–32. doi: 10.2307/3240049 Khanna LP (1933) A new species of Notothylas from Rangoon. Revue Bryologique et Lichénologique 6: 116–118. Khanna LP (1936) On Indian species of the genus Anthoceros Linn. with a description of a new species from Travancore. Journal of the Indian Botanical Society 15 (4): 235–240. Khanna LP (1938) On two species of Anthoceros from China. Journal of the Indian Botanical Society 17 (5/6): 311–323. Kitagawa N (1959) New or interesting species of Gymnomitrion (Hepaticae) in Japan. Acta Phytotaxonomica et Geobotanica 18 (2/3): 33–38. World checklist of hornworts and liverworts 743 Kitagawa N (1960a) Notes on the hepatic lora of Island of Yakushima. Acta Phytotaxonomica et Geobotanica 18 (7): 187–192. Kitagawa N (1960b) New or interesting species of Marsupella in Japan. Memoirs of the College of Science, Kyoto Imperial University. Series B. Biology 27: 75–82. Kitagawa N (1962a) Status of Gymnomitrion concinnatum var. mucronulatum. Acta Phytotaxonomica et Geobotanica 19 (2/3): 53. Kitagawa N (1962b) [Hepaticae of Mt. Tsurugi in Shikoku, Japan]. Acta Phytotaxonomica et Geobotanica 19 (2/3): 54–66. Kitagawa N (1963a) Lophoziaceae of Taiwan (Formosa). Hikobia 3 (3): 169–176. Kitagawa N (1963b) A revision of the family Marsupellaceae of Japan. Journal of the Hattori Botanical Laboratory 26: 76–118. Kitagawa N (1964) A new genus of hepaticae from North Borneo. Journal of the Hattori Botanical Laboratory 27: 178–182. Kitagawa N (1965) A revision of the family Lophoziaceae of Japan and its adjacent regions. I. Journal of the Hattori Botanical Laboratory 28: 239–291. Kitagawa N (1967a) Studies on the hepaticae of hailand. I. he genus Bazzania, with general introduction. Journal of the Hattori Botanical Laboratory 30: 249–270. Kitagawa N (1967b) Marsupellae of Mt. Kinabalu, North Borneo. Journal of the Hattori Botanical Laboratory 30: 199–204. Kitagawa N (1969a) A new species of Andrewsianthus from North Borneo. Journal of the Hattori Botanical Laboratory 32: 307–310. Kitagawa N (1969b) A new species of Cololejeunea (Chondriolejeunea) from Malay Peninsula. Acta Phytotaxonomica et Geobotanica 23 (5/6): 184–188. Kitagawa N (1969c) Studies on the hepaticae of hailand. II. Cephalozia and Cephaloziella. Journal of the Hattori Botanical Laboratory 32: 290–306. Kitagawa N (1970) Lophoziaceae of North Borneo. Journal of the Hattori Botanical Laboratory 33: 203–221. Kitagawa N (1972) Miscellaneous notes on little-known species of hepaticae, 1-25. Journal of the Hattori Botanical Laboratory 36: 444–454. Kitagawa N (1973) Miscellaneous notes on little-known species of hepaticae, 26-50. Journal of the Hattori Botanical Laboratory 37: 263–273. Kitagawa N (1977) Studies on Asian species of Bazzania, Hepaticae, I. Bulletin of Nara University of Education. Series B, Natural Sciences 26 (2): 73–82. Kitagawa N (1978) he d of hailand collected by Dr. A. Touw (I). Acta Phytotaxonomica et Geobotanica 29 (1/5): 47–64. Kitagawa N (1979a) Studies on Asian species of Bazzania, hepaticae, II. Bulletin of Nara University of Education. Series B, Natural Sciences 28 (2): 71–83. Kitagawa N (1979b) he hepaticae of hailand collected by Dr. A. Touw (II). Acta Phytotaxonomica et Geobotanica 30 (1/3): 31–40. Kitagawa N (1980) New Guinean species of the genus Bazzania, I. Journal of the Hattori Botanical Laboratory 47: 127–143. Kitagawa N (1981a) Miscellaneous notes on little-known species of hepaticae, 51-70. Hikobia, suppl. 1: 67–72. 744 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Kitagawa N (1981b) Two interesting species of the hepaticae from the Philippines. Miscellanea Bryologica et Lichenologica 9 (1): 8–10. Kitagawa N (1982) A remarkable new species of Hygrobiella from Japan. Miscellanea Bryologica et Lichenologica 9 (4): 69–72. Kitagawa N (1984) A new genus of the hepaticae from New Caledonia. Acta Phytotaxonomica et Geobotanica 35 (1/3): 1–6. Kitagawa N (1985) A study of the genus Acromastigum (Hepaticae) of New Caledonia. Acta Phytotaxonomica et Geobotanica 36 (4/6): 107–122. Kitagawa N (1988) Studies on the hepaticae of hailand. V. he Family Calypogeiaceae. Beihefte zur Nova Hedwigia 90: 163–170. Kitagawa N, Grolle R (1985) A new name for Acroscyphus N. Kitag., Hepaticae. Acta Phytotaxonomica et Geobotanica 36 (1/3): 58. Kitagawa N, Grolle R (1986) A new Acromastigum-like species of Bazzania S. Gray from Bhutan. Journal of the Hattori Botanical Laboratory 61: 269–272. Kitagawa N, Kodama T (1973) Enumeration of hepaticae collected by Drs. S. Kokawa and M. Hotta in Sabah (North Borneo) I. Bulletin of the Osaka Museum of Natural History 27: 11–21. Kitagawa N, Kodama T (1974) Enumeration of hepaticae collected by Drs. S. Kokawa and M. Hotta in Sabah (North Borneo) II. Bulletin of the Osaka Museum of Natural History 28: 33–48. Kitagawa N, Kodama T (1975a) A remarkable new species of Bazzania (Hepaticae) with endogenous gemmae. Journal of Japanese Botany 50 (1): 11–14. Kitagawa N, Kodama T (1975b) Two new species of Bazzania with an unusual habitat in Sabah (North Borneo). Journal of the Hattori Botanical Laboratory 39: 67–70. Knapp S, McNeill J, Turland N (2011) Changes to publication requirements made at the XVIII International Botanical Congress in Melbourne: What does e-publication mean for you? Phytotaxa 28: 1–5. Kodama T (1976) hree new species of Lejeuneaceae from the Kinabalu National Park, Sabah (North Borneo). Journal of the Hattori Botanical Laboratory 41: 381–387. Koike N (1994) Frullania taxa of some islands of Micronesia. Journal of the Hattori Botanical Laboratory 75: 183–192. Konstantinova NA (2000) Redkie pechenochniki (Hepaticae) Murmansko oblasti i podhody k ih ohrane [Rare liverworts (Hepaticae) of Murmansk province and some approaches to their protection]. Botanicheskij Zhurnal. Moscow & Leningrad 85 (10): 122–135. Konstantinova NA, Potemkin AD (1994) Studies on Scapania sphaerifera (Hepaticae). Annales Botanici Fennici 31 (2): 121–126. Konstantinova NA, Vasil’ev AN (1994) On the hepatic lora of Sayan Mountains (south Siberia) [K lore pečenočnikov Saân (ûžnaâ Sibir’)]. Arctoa 3: 123–132. doi: 10.15298/arctoa.03.07 Konstantinova NA, Vilnet AA (2009) New taxa and new combinations in Jungermanniales (Hepaticae). Arctoa 18: 65–67. doi: 10.15298/arctoa.18.02 Konstantinova NA, Vilnet AA (2011) Jubula hutchinsiae subsp. caucasica subsp. nov. (Jubulaceae, Marchantiophyta) – a new taxon from the western Caucasus. Arctoa 20: 227–238. doi: 10.15298/arctoa.20.18 World checklist of hornworts and liverworts 745 Konstantinova NA, Potemkin AD, Shliakov RN (1992) Checklist of the hepaticae and anthocerotae of the former USSR. Arctoa 1: 87–127. doi: 10.15298/arctoa.01.02 Konstantinova NA, Bakalin VA, Andreeva EN, Bezgodov AG, Borovichev EA, Dulin MA, Mamontov YS (2009) Checklist of liverworts (Marchantiophyta) of Russia. Arctoa 18: 1–64. doi: 10.15298/arctoa.18.01 Konstantinova NA, Vilnet AA, Söderström L, Hagborg A, von Konrat MJ (2013a) Notes on Early Land Plants Today. 14. Transfer of two Macrodiplophyllum species to Douinia (Scapaniaceae, Marchantiophyta). Phytotaxa 76 (3): 31–32. doi: 10.11646/phytotaxa.76.3.2 Konstantinova NA, Söderström L, Hagborg A, von Konrat MJ (2013b) Notes on Early Land Plants Today. 15. Apotreubia hortoniae validated (Treubiaceae, Marchantiophyta). Phytotaxa 76 (3): 33. doi: 10.11646/phytotaxa.76.3.3 Konstantinova NA, Vilnet AA, Söderström L, Hagborg A, von Konrat M (2014a) Notes on Early Land Plants Today. 53. Hygrobiellaceae (Marchantiophyta) validated. Phytotaxa 167 (2): 217. doi: 10.11646/phytotaxa.167.2.12 Konstantinova NA, Söderström L, Hagborg A, von Konrat M (2014b) Notes on Early Land Plants Today. 51. Validation of Schistochilopsis hyperarctica (Scapaniaceae, Marchantiophyta). Phytotaxa 162 (4): 240. doi: 10.11646/phytotaxa.162.4.10 Koponen TJ, Isoviita P, Lammes T (1977) he bryophytes of Finland: an annotated checklist. Flora Fennica 6: 1–77. Koponen TJ, Järvinen I, Isoviita P (1978) Bryophytes from the Soviet Far East, mainly the Khabarovsk territory. Annales Botanici Fennici 15 (2): 107–121. Koponen TJ, Cao T, Huttunen S, Juslén A, Peng C, Piippo S, Rao P, Váňa J, Virtanen V (2004) Bryophyte lora of Hunan Province, China. 3. Bryophytes from Taoyuandong and Yankou nature reserves and Badagongshan and Hupingshan national nature reserves, with additions to loras of Mangshan Nature Reserve and Wulingyuan Global cultural Heritage Area. Acta Botanica Fennica 177: 1–47. Krauss F (1846) Planzen des Cap- und Natal-Landes. Div. II. Plantae cellulares. Flora 29 (9): 129–138. Kreh W (1909) Über die Regeneration der Lebermoose. Nova Acta Academiae Caesareae LeopoldinoCarolinae Germanicae Naturae Curiosorum 90 (4): 213–301. Kreier H-P, Feldberg K, Mahr F, Bombosch A, Schmidt AR, Zhu R-L, von Konrat M, Shaw B, Shaw AJ, Heinrichs J (2010) Phylogeny of the leafy liverwort Ptilidium: Cryptic speciation and shared haplotypes between the Northern and Southern Hemispheres. Molecular Phylogenetics and Evolution 57 (3): 1260–1267. doi: 10.1016/j. ympev.2010.10.002 Krémer J-P (1837) Monographie des hépatiques de la Moselle. Metz, 44 pp. Kruijt RC (1988) A monograph of the genera Dicranolejeunea and Acanthocoleus. Bryophytorum Bibliotheca 36: 1–135. Kruijt RC, Gradstein SR (1985) A new genus of Lejeuneaceae from Tropical America: Lindigianthus gen. nov. Beihefte zur Nova Hedwigia 80: 165–172. Kruijt RC, Gradstein SR (1986) Studies on Lejeuneaceae subfam. Ptychanthoideae X. On Brachiolejeunea phyllorhiza (Nees) Kruijt & Gradstein comb. nov. (Hepaticae). Nova Hedwigia 43 (3/4): 299–309. 746 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Kühnemann O (1937) Contribución al catálogo briológico argentino. I. Revista del Centro de Estudiantes del Doctorado en Ciéncias Naturales 1: 155–179. Kumar D, Manocha N (1999) Herbertus udarii Kumar et Manocha, a new species from India. Geophytology 29 (1/2): 105–109. Kummer P (1875) Der Führer in die Lebermoose und die Gefässkryptogamen. Springer Verlag, Berlin, 141 pp. doi: 10.1007/978-3-642-91857-5 Kunth CS (1822) Synopsis plantarum. Tomus primus. F. G. Levrault, Paris, 491 pp. doi: 10.5962/bhl.title.638 Kuntze O (1891) Revisio generum plantarum. Pars II. Arthur Felix, Leipzig, 375–1011. doi: 10.5962/bhl.title.327 Kuwahara Y (1965) he Metzgeriaceae of Mt. Kinabalu, North Borneo. Journal of the Hattori Botanical Laboratory 28: 166–170. Kuwahara Y (1969a) An addendum to the Himalayan Metzgeriaceae. Journal of the Hattori Botanical Laboratory 32: 17–20. Kuwahara Y (1969b) Taxonomic and phytogeographic accounts of three new species of the hepatic genus Metzgeria from higher altitudes of New Guinea, the Philippines and Japan. Revue Bryologique et Lichénologique 36 (3/4): 531–542. Kuwahara Y (1973a) Steereella, a new genus of hepaticae from the West Indies. American Journal of Botany 60 (6): 602–606. doi: 10.2307/2441386 Kuwahara Y (1973b) Further notes on the production of vegetative thallus structures by female involucres of Metzgeria, and a new species of Metzgeria. Bryologist 76 (4): 566–571. doi: 10.2307/3241426 Kuwahara Y (1976a) Studies of genus Metzgeria of Colombia collected by Mme. Hélène Bischler, 1956–1959. Journal of the Hattori Botanical Laboratory 40: 259–290. Kuwahara Y (1976b) Metzgeria temperata, a new holarctic species of hepaticae. Journal of the Hattori Botanical Laboratory 40: 217–220. Kuwahara Y (1976c) Four new Metzgeria species from South America. Journal of the Hattori Botanical Laboratory 40: 509–519. Kuwahara Y (1978) Elaterophores observed in two new species of Metzgeria from Peru and considerations of the elaterophore in the hepaticae. Bryologist 81 (3): 404–410. doi: 10.2307/3242242 Kuwahara Y (1980a) Four new species of the Metzgeriaceae (Hepaticae) from the subantarctic region, with enumeration of previously reported taxa of the family. Hikobia 8 (3/4): 274–296. Kuwahara Y (1980b) Metzgeria maegdefraui, spec. nova from the neotropics. Hikobia 8 (3/4): 269–273. Kuwahara Y (1981) Studies of Peruvian collections of the genus Metzgeria made by P. & E. Hegewald in 1973 and 1977. Nova Hedwigia 34: 769–815. Kuwahara Y (1982) Studies on Colombian cryptogams. XV. On high Andean Metzgeria collected by Dr. Antoine M. Cleef in 1972 and 1973. Proceedings, Koninklijke Nederlandse Akademie van Wetenschappen. Series C, biological and medical sciences 85 (3): 357–380. Kuwahara Y (1983) Metzgeria subundulata (Aust. ex Lindb.) Kuw., stat. nov. from Texas. Bryologist 86 (3): 276–277. doi: 10.2307/3242721 World checklist of hornworts and liverworts 747 Kuwahara Y (1986) he Metzgeriaceae of the neotropics. Bryophytorum Bibliotheca 28: 1–254. Kuwahara Y (1987) Two new species of Metzgeria from New Zealand, with a key to the New Zealand species of the family Metzgeriaceae. Memoirs of the New York Botanical Garden 45: 561–568. Labillardière JJH (1806) Novae Hollandiae plantarum specimen, tomus secundus. Dominae Huzard, Paris, 130 pp. Lacouture C (1905) Hépatiques de la France. Paul Klincksieck, Paris, 58 pp. Lai M-J, Zhu R-L, Chantanaorrapint S (2008) Liverworts and hornworts of hailand: an updated checklist and bryoloristic accounts. Annales Botanici Fennici 45 (5): 321–341. Lamarck JB (1789) Encyclopédie méthodique, botanique, tome 3 (1). Panckoucke, Paris, 1–360. doi: 10.5962/bhl.title.824 Lamarck JB, Poiret JLM (1804) Encylopédie méthodique, botanique. Tome sixième, part 1. H. Agasse, Paris, 384 pp. doi: 10.5962/bhl.title.824 Lamothe A (1919) Recherches anatomiques et taxinomiques sur le gamétophyte des Marchantiales. hèses de la Faculté des sciences de l’Université de Lyon, Lyon, 1–181. Lange J (1871) Floræ danicæ iconum fasciculus vol. 16 heft 48 (Tab. MMDCCCXXI-MMDCCCLXXX). København, 1–26. Larraín J, Carter B, Shaw B, Hentschel J, Strozier LS, Furuki T, Heinrichs J, Crandall-Stotler B, Engel J, von Konrat M (2015) he resurrection of Neohattoria Kamim. (Jubulaceae, Marchantiophyta): a six decade systematic conlict resolved through a molecular perspective. PhytoKeys 50: 101–122. doi: 10.3897/phytokeys.50.4940 Lawrence GHM, Bucheim AFG, Daniels GS, Dolezal H (1968) Botanico Periodicum Huntianum. Hunt Institute for Botanical Documentation, Pittsburgh, 1063 pp. Lee GE (2013) A systematic revision of the genus Lejeunea Lib. (Marchantiophyta: Lejeuneaceae) in Malaysia. Cryptogamie, Bryologie 34 (4): 381–484. doi: 10.7872/cryb.v34. iss4.2013.381 Lee GE, Gradstein SR (2013) Distribution and habitat of the Malaysian species of Lejeunea (Marchantiophyta: Lejeuneaceae), with description of Lejeunea tamaspocsii sp. nov. Polish Botanical Journal 58 (1): 59–69. doi: 10.2478/pbj-2013-0007 Lee GE, Pócs T, Damanhuri A, Latif A (2010) Lejeunea gradsteinii (Lejeuneaceae), a new liverwort species from Mt. Kinabalu, Sabah. Acta Biologica Plantarum Agriensis 1: 29–36. Lehmann JGC (1829) Hepaticarum capensium a C.F. Linnaea 4: 357–371. Lehmann JGC (1831) Novarum et minus cognitarum stirpium pugillus tertius. Meissner, Hamburg, 58 pp. doi: 10.5962/bhl.title.45011 Lehmann JGC (1832) Novarum et minus cognitarum stirpium pugillus quartus. Meissner, Hamburg, 64 pp. doi: 10.5962/bhl.title.45011 Lehmann JGC (1833) Novarum et minus cognitarum stirpium pugillus quintus. Meissner, Hamburg, 28 pp. doi: 10.5962/bhl.title.45011 Lehmann JGC (1834) Novarum et minus cognitarum stirpium pugillus sextus. Meissner, Hamburg, 72 pp. doi: 10.5962/bhl.title.45011 Lehmann JGC (1835) Cel. Endlicher. Linnaea 9 (4): 421–427. 748 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Lehmann JGC (1838) Novarum et minus cognitarum stirpium pugillus septimus. Meissner, Hamburg, 41 pp. doi: 10.5962/bhl.title.45011 Lehmann JGC (1844) Novarum et minus cognitarum stirpium pugillus octavus. Meissner, Hamburg, 56 pp. doi: 10.5962/bhl.title.45011 Lehmann JGC (1857) Novarum et minus cognitarum stirpium pugillus decimus. Meissner, Hamburg, 34 pp. doi: 10.5962/bhl.title.45011 Leitgeb H (1876) Ueber Zoopsis. Mitteilungen des Naturwissenschaftlichen Vereines für Steiermark 13: 21–28. León Y, Pócs T, Rico RR (1998) Registros para la briolora de Los Andes Venezolanos, I. Cryptogamie: Bryologie, Lichénologie 19 (1): 1–25. Levier E (1894) Tessellina pyramidata e Riccia macrocarpa (in Adunza della sede di Firenze 11 Marzo 1894). Bullettino della Società Botanica Italiana 1894: 114–115. Levier E (1902) Riccia crozalsii Lev. nov. spec. Revue Bryologique 29 (4): 73–76. Levier E (1906) Muscinee raccolte nello Schen-Si (Cina) dal Rev. Guiseppe Giraldi (continuazione e ine). Nuovo Giornale Botanico Italiano (n.ser.) 13 (4): 347–356. Lewington RJ, Beveridge P, Renner MAM (2013) Lejeunea hodgsoniana, a newly described, long recognised Lejeunea (Jungermanniopsida, Lejeuneaceae) from lowland coastal forest habitats in New Zealand. PhytoKeys 29: 1–15. doi: 10.3897/phytokeys.29.5376 Li X-J (1985) Bryolora of Xizang. Science Press, Beijing, 581 pp. Li J, Zhang L, Zhou L (2011) Phylogenetic position of the genus Hattorioceros (Anthoceratophyta). Taxon 60 (6): 1633–1636. Libert M-A (1820) Sur un genre nouveau d’hépatiques, Lejeunia. Annales générales des sciences physiques 6: 372–374. Lightfoot J (1777) Flora scotica, vol. 2. B. White, London, 531–1151. Limpricht KG (1876) Schlesische Lebermoose. Hedwigia 15 (2): 17–19. Limpricht KG (1879) Neue und kritische Lebermoose. Jahresbericht der Schlesischen Gesellschaft für Vaterländische Cultur 57: 311–317. Limpricht KG (1881) Ueber Gymnomitrium adustum N. v. E. Flora 64 (5): 71–76. Limpricht KG (1884) Über einige neue Arten und Formen bei den Laub- und Lebermoosen. Jahresbericht der Schlesischen Gesellschaft für Vaterländische Cultur 61: 204–225. Lin S-H, Chen Y-Y (1997) A taxonomic study of Frullaniaceae from Taiwan. Tunghai Journal 38: 79–125. Lin S-H, Yang C-S (1992) Bryophytes of Chaishan, Taiwan. Yushania 9: 1–12. Lin P-J, Piippo S, Koponen TJ, Wu P-C (1992) Bryophyte lora of Jianfengling Mts., Hainan Island, China. Bryobrothera 1: 195–214. Lindberg SO (1866) Förteckning öfver mossor, insamlade under de svenska expeditionerna till Spitsbergen 1858 och 1861. Öfversigt af Kongl. Vetenskaps-Akademiens Förhandlingar 23 (10): 535–561. Lindberg SO (1868a) En liten profbit på namnförbistring. Notiser ur Sällskapets pro Fauna et Flora Fennica Förhandlingar 9: 3–18. Lindberg SO (1868b) Musci novi scandinavici. Notiser ur Sällskapets pro Fauna et Flora Fennica Förhandlingar 9: 253–299. World checklist of hornworts and liverworts 749 Lindberg SO (1870) Nya mossor. Öfversigt af Finska Vetenskaps-Societetens Förhandlingar 12 (2): 70–84. Lindberg SO (1871) Revisio critica iconum. Societatis litterariae fennicae, Helsinki, 1–118. Lindberg SO (1872a) On Zoopsis H.f. & T. Journal of the Linnean Society. Botany 13 (67): 188–203. doi: 10.1111/j.1095-8339.1872.tb02401.x Lindberg SO (1872b) Contributio ad loram cryptogamam Asiae boreali-orientalis. Acta Societatis Scientiarum Fennicae 10: 221–280. Lindberg SO (1873a) Sällskapet pro Fauna & Flora Fennica sammanträde 6 dennes. Morgonbladet (Helsinki) 1873 (286, 9 Dec): 1–2. Lindberg SO (1873b) Sällskapet pro Fauna et Flora Fennicas sammanträde den 6 december. Helsingfors Dagblad 1873 (353, 28 Dec): 2. Lindberg SO (1873c) Referat från Sällskapet pro Fauna et Flora Fennicas möte 4 Okt. 1873. Helsingfors Dagblad 1873 (273, 7 Oct): 2–3. Lindberg SO (1874a) Manipulus muscorum secundus. Notiser ur Sällskapets pro Fauna et Flora Fennica Förhandlingar 13: 351–417. Lindberg SO (1874b) Sällskapet pro Fauna et Flora Fennicas sammanträde den 7 februari. Helsingfors Dagblad 1874 (45, 16 Feb): 2. Lindberg SO (1875) Hepaticae in Hibernia mense Julii 1873 lectae. Acta Societatis Scientiarum Fennicae 10: 465–559. Lindberg SO (1876a) Sällskapets pro Fauna et Flora fennica sammanträde den 4 november 1876. Helsingfors Dagblad 1876 (323, 26 Nov.): 2. Lindberg SO (1876b) Meddelanden från Sällskapets förhandlingar I. Meddelanden af Societas pro Fauna et Flora Fennica 1: 91–119. Lindberg SO (1876c) Sällskapet pro Fauna & Flora Fennica sammanträde 2 dennes. Morgonbladet (Helsinki) 1876 (287, 10 Dec.): 1. Lindberg SO (1877a) Sällskapets pro Fauna et Flora fennica sammanträde den 3 dennes. Morgonbladet (Helsinki) 1877 (30, 6 Feb): 2. Lindberg SO (1877b) Monographia Metzgeriae. Acta Societatis pro Fauna et Flora Fennica 1 (2): 1–49. Lindberg SO (1877c) Hepaticologiens utveckling från äldsta tider till och med Linné. J.C. Frenckell, Helsinki, 51 pp. Lindberg SO (1877d) Riccia bicarinata n. sp. Lindb. Revue Bryologique 4 (3): 41–42. Lindberg SO (1878) Sällskapet pro Fauna et Flora Fennicas sammanträde den 9 november. Helsingfors Dagblad 1878 (315, 18 Nov.): 2–3. Lindberg SO (1879) Musci Scandinavici. Jessaie Edquist, Uppsala, 50 pp. Lindberg SO (1880a) Distinctio Scapaniae carinthiacae e Sc. apiculata. Revue Bryologique 7 (4): 77–78. Lindberg SO (1880b) Sällskapet pro Fauna et Flora Fennicas sammanträde den 6 november. Helsingfors Dagblad 1880 (311, 15 Nov.): 2–3. Lindberg SO (1882) Monographia praecursoria Peltolepidi, Sauteriae et Cleveae. Acta Societatis pro Fauna et Flora Fennica 2 (3): 1–15. Lindberg SO (1884) Sandea et Myriorrhynchus. Acta Societatis pro Fauna et Flora Fennica 2 (5): 1–9. 750 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Lindberg SO (1886) Sur la morphologie des mousses (suite). Revue Bryologique 13 (6): 100–109. Lindberg SO (1887a) Bidrag till nordens mosslora. I. J. Simelii arfvingar, Helsingfors, 63–77. Lindberg SO (1887b) Hepaticae novae lusitanicae. Revue Bryologique 14 (2): 19–21. Lindberg SO, Arnell HW (1889) Musci asiae borealis. Erster heil: Lebermoose. Kongliga Svenska Vetenskaps-Akademiens Handlingar (n.ser.) 23 (5): 1–69. Lindberg SO, Lackström EF (1874) Hepaticae Scandinavicae exsiccata quarum specimina, fasc. 1. Helsinki, no. 1–25. Lindenberg JBW (1829) Synopsis hepaticarum europaearum. Eduard Weberum, Bonnae, 1–133. Lindenberg JBW (1836) Monographie der Riccieen. Nova Acta Physico-Medica Academiae Caesareae Leopoldino-Carolinae Naturae Curiosorum 18 (1): 361–504. Lindenberg JBW (1839) Species hepathicarum, fasc. 1. Henry & Cohen, Bonn, 1–36. Lindenberg JBW (1840) Species hepaticarum, fasc. 2–4. Henry & Cohen, Bonn, 37–120. Lindenberg JBW (1843) Species hepaticarum, fasc. 5. Henry & Cohen, Bonn, 121–164. Lindenberg JBW (1844) Monographia hepaticarum generis Plagiochilae. Henry & Cohen, Bonn, i–xxvix. Lindenberg JBW, Gottsche CM (1851a) Plantae Kegelianae. Linnaea 24 (6): 625–639. Lindenberg JBW, Gottsche CM (1851b) Species hepaticarum, fasc. 8–11. Henry & Cohen, Bonn, 1–118. Link HF (1823) Neuigkeiten. Flora 6 (2): 27–32. Linnaeus C (1753) Species plantarum, ed. 1. Laurentii Salvii, Holmiae [Stockholm], 1200 pp. Linnaeus C (1759) Systema naturae. Tomus II. Editio decima, reformata. Laurentii Salvii, Holmiae [Stockholm], 825–1384. Linnaeus C (1763) Species plantarum, tomus II, editio secunda. Laurentii Salvii, Holmiae [Stockholm], 785–1648. Loeske L (1903) Mooslora des Harzes. Bornträger, Leipzig, 350 pp. Loeske L (1905) Bryologisches vom Harze und aus anderen Gebieten. Verhandlungen des Botanischen Vereins der Provinz Brandenburg 47: 317–344. Loeske L (1908) Bryologische Beobachtungen aus der Algäuer Alpen von Loeske und Osterwald. Verhandlungen des Botanischen Vereins der Provinz Brandenburg 49 (1): 30–65. Loeske L (1909) Zur Mooslora der Zillertaler Alpen. Hedwigia 49 (1/2): 1–53. Loeske L (1928) Fossombronia leischeri Osterwald. Verhandlungen des Botanischen Vereins der Provinz Brandenburg 70: 125–127. Loitlesberger K (1894) Hepaticae. In: Szyszylowicz I (Ed.) Diagnoses plantarum novarum a cl. d. Const. Jelski in Peruvia lectarum, pars prima. Sumptibus Academiae, Cracoviae, 18–25. Long DG (1999a) Studies on the genus Asterella. IV. Asterella grollei sp. nov., a new species from eastern Asia related to the American A. palmeri. Bryologist 102 (2): 169–178. doi: 10.2307/3244357 Long DG (1999b) Validation of the combination Asterella drummondii. Journal of Bryology 21 (1): 76. doi: 10.1179/jbr.1999.21.1.76 Long DG (2001) Studies on the genus Asterella (Aytoniaceae). V. Miscellaneous notes on Asiatic Asterella. Lindbergia 26 (1): 43–45. World checklist of hornworts and liverworts 751 Long DG (2005) Studies on the genus Asterella (Aytoniaceae) VI. Infrageneric classiication in Asterella. Journal of the Hattori Botanical Laboratory 97: 249–261. Long DG (2006) Revision of the genus Asterella P. Beauv. in Eurasia. Bryophytorum Bibliotheca 63: 1–299. Long DG, Grolle R (1990) Hepaticae of Bhutan II. Journal of the Hattori Botanical Laboratory 68: 381–440. Long DG, Grolle R (1994) Studies on the genus Asterella P. Beauv. II. Asterella limbata, a new species from Sumatra and Sabah. Journal of Bryology 18 (2): 287–295. doi: 10.1179/ jbr.1994.18.2.287 Long DG, Rubasinghe SCK (2014) Liverworts and hornworts of Sri Lanka: a revised checklist. Ceylon Journal of Science. Biological Sciences 43 (1): 1–36. doi: 10.4038/cjsbs.v43i1.7280 Long DG, Váňa J (2007) he genus Gottschelia Grolle (Jungermanniopsida, Lophoziaceae) in China, with a description of G. grollei, sp. nov. Journal of Bryology 29 (3): 165–168. doi: 10.1179/174328207X227410 Long DG, Söderström L, Hagborg A, von Konrat M (2014) Notes on Early Land Plants Today. 56. Validation of Asterella sect. Wallichianae (Aytoniaceae, Marchantiophyta). Phytotaxa 173 (1): 87. doi: 10.11646/phytotaxa.173.1.8 Longton RE (1992) he role of bryophytes and lichens in terrestrial ecosystems. In: Bates JW, Farmer AM (Eds) Bryophytes and lichens in a changing environment. Clarendon Press, Oxford, 32–76. Lorbeer G (1934) Die Zytologie der Lebermoose mit besonderer Berücksichtigung allgemeiner Chromosomenfragen. I. Teil. Jahrbücher für Wissenschaftliche Botanik 80: 567–817. Lou J-S (1987) hree new species of Porella from Mountain Hengduan, China. Acta Phytotaxonomica Sinica 25 (6): 482–485. Lou J-S, Wu P-C (1980) A preliminary report on the new bryophytes of Xizang (Tibet). Acta Phytotaxonomica Sinica 18 (1): 119–125. Lou J-S, Wang M-Z (1983) [he mountain Hengduan, the distributioned centre for east-asiatic Porellaceae (Hepaticae)]. In: Sun HL (Ed.) Qing Zang gao yuan yan jiu Heng Duanshan kaocha zhuan ji [he collective papers of study on Qing-Zang Plateau and Heng-Duan Mt. exploration 1]. Yunnan People’s Publishing House, Kunming, 270–279. Loureiro J (1790) Flora cochinchinensis, tomus II. Academicis, Ulyssipone [Lisboa], 357–745. Ludwig CG (1760) Deinitiones generum plantarum, ed. 3. Joh. Frideric. Gleditschii, Leipzig, 516 pp. Lughadha EN (2004) Towards a working list of all known plant species. Philosophical Transactions of the Royal Society of London. Series B 359 (1444): 681–687. doi: 10.1098/ rstb.2003.1446 Lye KA, Pócs T (1997) New records to the hepatic lora of Uganda. Lidia 4 (1): 13–36. MacGregor WM (1899) Flora of British New Guinea. Bulletin of Miscellaneous Information, Royal Gardens, Kew 1899 (151/152): 95–126. doi: 10.2307/4111436 Macvicar SM (1911) Fossombronia echinata nov. sp. Revue Bryologique 38 (4): 73–76. Macvicar SM (1926) he student’s handbook of British hepatics, ed. 2. Wheldon & Wesley, London, 464 pp. 752 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Mägdefrau K (1983) he bryophyte vegetation of the forests and páramos of Venezuela and Colombia. Nova Hedwigia 38: 1–63. Mahabale TS (1941) A long-lost liverwort from South India: Aspiromitus, a rare member of the anthocerotes. Current Science 10 (12): 530–533. Maire R (1933) Études sur lore et végétation du Sahara central. Mémoires de la Société d’Histoire Naturelle de l’Afrique du Nord 3: 1–272. Maire R, Weiler M (1939) Contributions à l’étude de la lore de Libye. Bulletin de la Société d’Histoire Naturelle de l’Afrique du Nord 30 (5): 255–314. Malombe I (2009) Studies on African Cheilolejeunea (Lejeuneaceae) I: New species and new combinations. Acta Botanica Hungarica 51 (3/4): 315–328. doi: 10.1556/ ABot.51.2009.3-4.8 Malombe I, Fischer E, Pócs T (2010) Cheilolejeunea ulugurica (Lejeuneaceae, Marchantiophyta), a new species from Tanzania. Acta Biologica Plantarum Agriensis 1: 23–28. Mamontov YS, Vilnet AA (2013) Anastrophyllum ellipticum Inoue (Jungermanniales, Marchantiophyta), a new species for Russian liverwort lora [Anastrophyllum ellipticum Inoue (Jungermanniales, Marchantiophyta) – novyj vid dlâ lory pečenočnikov Rossii]. Arctoa 22: 151–158. doi: 10.15298/arctoa.22.24 Mamontov YS, Konstantinova NA, Vilnet AA, Bakalin VA (2015) On the phylogeny and taxonomy of Pallaviciniales (Marchantiophyta), with overview of Russian species. Arctoa 24 (1): 98–123. doi: http://dx.doi.org/10.15298/arctoa24.12 Manju CN, Rajesh KP, Madhusoodanan PV (2011) Chiloscyphus chinnarensis (Lophocoleaceae: Marchantiophyta): A new species from the Western Ghats of India. Acta Botanica Hungarica 53 (1/2): 151–157. doi: 10.1556/ABot.53.2011.1-2.14 Martelli M (1889) Una nuova specie di Riccia. Nuovo Giornale Botanico Italiano 21 (2): 290–292. Martin W (1950) he bryophytes of Stewart Island – part II. Transactions and Proceedings of the Royal Society of New Zealand 78 (4): 485–501. Martius CFP (1817) Flora cryptogamica erlangensis. J.L.Schrag, Norinbergae, 508 pp. Massalongo C (1879) Hepaticologia veneta, fasc. 1. P. Prosperini, Padova, 1–68. Massalongo C (1881) Duae species novae e genere Lejeunia, quas circa Buenos-Ayres legit C. Spegazzini. Nuovo Giornale Botanico Italiano 13 (2): 122–124. Massalongo C (1885) Epatiche raccolte alla Tierra del Fuoco. Nuovo Giornale Botanico Italiano 17 (3): 201–277. Massalongo C (1889) Nuova specie di Lejeunea scoperta dal Dott. C. Rossetti in Toscana. Nuovo Giornale Botanico Italiano 21 (3): 485–487. Massalongo C (1897) Hepaticae in provincia Schen-si. G. Franchini, Verona, 3–63. Massalongo C (1898) Due nuovi generi di Epatiche. Nuovo Giornale Botanico Italiano (n.ser.) 5 (2): 255–260. Massalongo C (1903) Le epatiche dell’erbario crittogamico italiano. G. Bresciani, Ferrara, 20 pp. Massalongo C (1904) Intorno alla Radula Visianica sp. nov. Annali di Botanica. Roma 1 (4): 297–300. World checklist of hornworts and liverworts 753 Massalongo C (1906a) Epatiche della Repubblica Argentina raccolte dal Prof. C. Spegazzini. Atti della Accademia delle Scienze Mediche e Naturali di Ferrara 80 (3/4): 1–14. Massalongo C (1906b) Di una nova specie di Madotheca della China. Bullettino della Società Botanica Italiana 1906: 141–141. Massalongo C (1907) La specie italiane del genere Cephalozia Dmrt. emend. Malpighia 21 (7/8): 289–339. Massalongo C (1913) Nuovi rappresentanti nella Flora Italica del genere Riccia. Bullettino della Società Botanica Italiana 1913 (2/3): 50–53. Massalongo C (1917) Nuova specie del genere Aneura D. Mort. Bullettino della Società Botanica Italiana 1917 (8/9): 80–82. Massalongo C (1928) Revisio critica hepaticarum in Argentina. Atti del Reale Istituto Veneto di Scienze, Lettere ed Arti 87 (2): 215–251. Massalongo C, Carestia A (1880) Epatiche delle Alpi Pennine. Nuovo Giornale Botanico Italiano 12 (4): 306–366. Masuzaki H (2011) A nomenclatural change for Metzgeria kinabaluensis (Metzgeriaceae, Marchantiophyta). Hikobia 16 (1): 59–60. Masuzaki H, Tsubota H, Shimamura M, Yamaguchi T, Deguchi H (2010a) A taxonomic revision of the genus Apometzgeria (Metzgeriaceae, Marchantiophyta). Hikobia 15 (4): 427–452. Masuzaki H, Shimamura M, Furuki T, Tsubota H, Yamaguchi T, Majid HMA, Deguchi H (2010b) Systematic position of the enigmatic liverwort Mizutania (Mizutaniaceae, Marchantiophyta) inferred from molecular phylogenetic analyses. Taxon 59 (2): 448–458. McCarthy PM (2003) Catalogue of Australian liverworts and hornworts. Flora of Australia Supplementary Series 21: 1–137. McCormick FA (1914) A study of Symphyogyna aspera. Botanical Gazette 58 (5): 401–418. doi: 10.1086/331430 McGregor RL (1960) A new species of Riccia from the Ozarks. Bryologist 63 (1): 30–31. doi: 10.2307/3241169 McNeill J, Barrie FR, Buck WR, Demoulin V, Greuter W, Hawksworth DL, Herendeen PS, Knapp S, Marhold K, Prado J, Prud’homme van Reine WF, Smith GF, Wiersema JH, Turland NJ (2012) International Code of Nomenclature for algae, fungi and plants (Melbourne Code) adopted by the Eighteenth International Botanical Congress Melbourne, Australia, July 2011. Regnum Vegetabile 154: 1–240. Meagher D (2002) New bryophyte records for Australia. Australasian Bryological Newsletter 46: 6. Meagher D (2005a) Bazzania sauropoda D. Meagher (Marchantiophyta: Lepidoziaceae), a new species from tropical Queensland. Austrobaileya 7 (1): 129–133. Meagher D (2005b) New and interesting bryophyte records from New South Wales, Queensland and Victoria. Australasian Bryological Newsletter 50: 6–9. Meagher D (2006) Bazzania scalaris sp. nov. (Marchantiophyta: Lepidoziaceae) from Papua New Guinea. Telopea 11 (3): 246–251. doi: 10.7751/telopea20065724 Meagher D (2008) Studies on Bazzania 1. Some new and little known species from Australasia. Nova Hedwigia 86 (3/4): 477–495. doi: 10.1127/0029-5035/2008/0086-0477 754 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Meagher D (2010) Studies on Bazzania 2. Seven poorly known species from Australia. Nova Hedwigia 90 (3/4): 395–411. doi: 10.1127/0029-5035/2010/0090-0395 Meagher D (2011) Studies on Bazzania (Lepidoziaceae, Marchantiophyta) 3. Four new species from Australia. Nova Hedwigia 92 (3/4): 487–495. doi: 10.1127/0029-5035/2011/00920487 Meagher D (2012) Studies on Bazzania (Marchantiophyta, Lepidoziaceae). 5. Bazzania lehmanniana (Lindenb.) Trevis., from Nova Hollandia but South America, not Australia. Nova Hedwigia 94 (1/2): 271–274. doi: 10.1127/0029-5035/2012/0094-0271 Meagher D (2013) Studies on Bazzania (Marchantiophyta: Lepidoziaceae) 7. Bazzania avia sp. nov. from Tasmania. Nova Hedwigia 97 (3/4): 529–532. doi: 10.1127/0029-5035/2013/0133 Meagher D (2015) Studies on Bazzania (Marchantiophyta: Lepidoziaceae) 8. Bazzania wooroonooran sp. nov. and seven other rare species from tropical Australia. Nova Hedwigia 100 (3/4): 535–552. doi: 10.1127/nova_hedwigia/2015/0247 Meagher D, Glenny DS (2007) A new species of Bazzania (Lepidoziaceae) from New Zealand. Journal of Bryology 29 (1): 60–63. doi: 10.1179/174328207X175184 Meenks JLD (1986) Riccardia reyesiana, a new liverwort from Cuba. Acta Botanica Hungarica 32 (1/4): 207–208. Meenks JLD (1987) Studies on Colombian cryptogams. XXVIII. A guide to the tropical Andean species of Riccardia (Hepaticae). Journal of the Hattori Botanical Laboratory 62: 161–182. Meenks JLD, De Jong C (1985) Light microscope studies on the oil bodies of Andean Aneuraceae (Hepaticae). Cryptogamie: Bryologie, Lichénologie 6 (1): 1–24. Meijer W (1956) A new species of Exormotheca from Ceylon. Journal of the Hattori Botanical Laboratory 16: 72–74. Meijer W (1957) Notes on some Malaysian species of Anthoceros L. (Hepaticae) – II. Journal of the Hattori Botanical Laboratory 18: 1–13. Meijer W (1958) Notes on species of Riccia from the Malaysian region. Journal of the Hattori Botanical Laboratory 20: 107–118. Meijer W (1959) Notes on species of Riccardia from their type localities in western Java. Journal of the Hattori Botanical Laboratory 21: 61–78. Meijer W (1960) Notes on the species of Bazzania (Hepaticae) mainly of Java. Blumea 10 (2): 367–384. Meissner CF (1848) Hepaticae Javanicae a Zollingero collectae. Botanische Zeitung. Berlin 6 (25): 462–463. Menzel M (1984) Katalog der Lebermoose von Peru. Willdenowia 14: 473–523. Mérat FV (1821) Nouvelle lore des environs de Paris (ed. 2), vol. 1 Cryptogamie. MéquignonMarvis, Paris, 292 pp. Mérat FV (1840) Notice sur une hépatique regardée comme l’individu mâle du Marchantia conica L. Annales de l’Agriculture Françoise (ser. 4) 2 (7): 1–12. Messe V (1981) Pellia borealis Lorbeer en Belgique. Bulletin de la Société Royale de Botanique de Belgique 114 (1): 3–14. Meylan C (1924) Les hépatiques de la Suisse. Beiträge zur Kryptogamenlora der Schweiz 6 (4): 1–318. World checklist of hornworts and liverworts 755 Meylan C (1926) Note sur une nouvelle espèce de Scapania. Jahresbericht der Naturforschenden Gesellschaft Graubündens (n.f.) 64: 363–366. Meylan C (1939) Localités nouvelles pour la lore des muscinées de la Suisse. Bulletin de la Société Vaudoise des Sciences Naturelles 60 (249): 261–276. Michaux A (1803) Flora boreali-americana, tomus secundus. Caroli Crapelet, Paris, 340 pp. Miller HA (1956) Cryptogams of Kapingamarangi Atoll, Carolina Islands. I. Bryophyta. Bryologist 59 (3): 167–173. doi: 10.2307/3239943 Miller HA (1960) A preliminary list of Micronesian Bryophytes. Bryologist 63 (2): 116–125. doi: 10.2307/3240888 Miller HA (1963) Notes on Hawaiian hepaticae. V. Collections from recent Swedish expeditions. Arkiv för Botanik (n.ser.) 5 (2): 489–531. Miller HA (1965) A review of Herberta in the tropical Paciic and Asia. Journal of the Hattori Botanical Laboratory 28: 299–412. Miller HA (1967) Oddments of Hawaiian bryology. Journal of the Hattori Botanical Laboratory 30: 271–276. Miller HA (1970) Some circum-paciic Schistochilaceae. Phytologia 20 (5): 315–323. Miller HA (1981) Notulae hepaticarum Polynesiae. Phytologia 47 (4): 319–324. Miller HA (1986) Paciic bryophytes. 3. An overview of Telaranea, Hepatophyta. Journal of Bryology 14 (2): 231–244. doi: 10.1179/jbr.1986.14.2.231 Miller DH, Miller HA (1994) A new Mastigophora (Hepatophyta) from New Guinea. Journal of the Hattori Botanical Laboratory 75: 179–182. Miller HA, Bonner CEB, Bischler H (1962) Studies in Lejeuneaceae V. Microlejeunea in Paciic Oceania. Nova Hedwigia 4: 551–561. Miller HA, Whittier HO, Bonner CEB (1963) Bryolora of the atolls of Micronesia. Beihefte zur Nova Hedwigia 11: 1–89. Miller HA, Bonner CEB, Bischler H (1967) Studies in Lejeuneaceae VIII. Microlejeunea in Asia and Australia. Nova Hedwigia 14 (1): 61–67. Miller HA, Whittier HO, Whittier BA (1983) Prodromus lorae hepaticarum Polynesiae. Catalogue of hepaticae and anthocerotae. Bryophytorum Bibliotheca 25: 1–423. Mitten W (1851) Catalogue of cryptogamic plants collected by Professor W. Jameson in the vicinity of Quito (conclusion). Hooker’s Journal of Botany and Kew Gardens Miscellany 3: 351–361. Mitten W (1854) Nat. Ord. CI. Hepaticae, L. In: Hooker JD (Ed.) he botany of the antarctic voyage. II. Flora Novae Zealandiae. Part II. Flowerless plants. Reeve, London, 125–160. doi: 10.5962/bhl.title.16029 Mitten W (1855) Nat. Ord. CI. Hepaticae, L. In: Hooker JD (Ed.) he botany of the antarctic voyage. II. Flora Novae Zealandiae. Part II. Flowerless plants. Reeve, London, 161–172. doi: 10.5962/bhl.title.16029 Mitten W (1860a) On some new species of musci and hepaticae in the herbarium of sir W. J. Hooker, collected in tropical Africa, chiely by the late Dr. Vogel and Mr. Barger. Transactions of the Linnean Society of London 23 (1): 51–58. doi: 10.1111/j.1096-3642.1860. tb00117.x 756 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Mitten W (1860b) Hepaticae. In: Hooker JD (Ed.) he Botany of the Antarctic Voyage of H. M. discovery ships Erebus and Terror in the years 1839-43. III. Flora Tasmaniae 2. Reeve, London, 221–241. Mitten W (1860c) Hepaticae Indiae Orientalis. Journal of the Proceedings of the Linnean Society. Botany 5 (18): 89–128. doi: 10.1111/j.1095-8312.1860.tb01045.x Mitten W (1861) Musci et hepaticae vitiensis. Bonplandia 9 (24): 365–367. Mitten W (1862) Musci et Hepaticae vitiensis. Bonplandia 10 (2): 19. Mitten W (1863) On the musci and hepaticae from the Cameroons mountain and from the River Niger. Journal of the Proceedings of the Linnean Society. Botany 7 (27): 147–169. doi: 10.1111/j.1095-8312.1863.tb01066h.x Mitten W (1864a) On some species of musci and hepaticae, additional to the lora of Japan and the coast of China. Journal of the Linnean Society. Botany 8 (31): 148–158. doi: 10.1111/j.1095-8312.1864.tb01081.x Mitten W (1864b) A new genus of hepaticae. Journal of the Proceedings of the Linnean Society. Botany 7 (28): 243–244. doi: 10.1111/j.1095-8312.1864.tb01067e.x Mitten W (1864c) he “Bryologia” of the survey of the 48th parallel of latitude. Journal of the Proceedings of the Linnean Society. Botany 8 (29): 12–55. doi: 10.1111/j.10958312.1864.tb01071.x Mitten W (1867) Hepaticae. In: Hooker JD (Ed.) Handbook of the New Zealand lora, part II. Reeve, London, 497–755. Mitten W (1870) Hepaticae. In: Godman F (Ed.) Natural history of the Azores, or Western Islands. John Van Voorst, London, 316–328. Mitten W (1871) Jungermanniae and Marchantiae. In: Seemann B (Ed.) Flora vitiensis, part 10. Reeve, London, 404–419. doi: 10.5962/bhl.title.455 Mitten W (1875) Hepaticae. In: Melliss JC (Ed.) St. Helena. Reeve, London, 366–373. Mitten W (1876a) he musci and hepaticae collected by H. N. Moseley, M. A., naturalist to H. M. S. “Challenger”. Journal of the Linnean Society. Botany 15 (82): 59–73. doi: 10.1111/j.1095-8339.1876.tb00223.x Mitten W (1876b) A list of musci and hepaticae collected in Kerguelen’s Island by the Rev. A. E. Eaton. Journal of the Linnean Society. Botany 15 (84): 193–197. doi: 10.1111/j.10958339.1876.tb00238.x Mitten W (1877) List of hepaticae collected by the Rev. A. E. Eaton at the Cape of Good Hope. Journal of the Linnean Society. Botany 16 (91): 187–197. doi: 10.1111/j.10958339.1877.tb02316a.x Mitten W (1879) Hepaticae. Philosophical Transactions of the Royal Society of London 168: 396–401. doi: 10.1098/rstl.1879.0038 Mitten W (1884a) Tristan da Cunha. Musci et Hepaticae. In: Hemsley W (Ed.) Report on the Scientiic Results of the voyage of H. M. S. Challenger. Botany. Vol. 1. Second Part. Her Majesty’s Stationary Oice, London, 170–178. Mitten W (1884b) Juan Fernández and Masafuera. Musci et hepaticae. In: Hemsley W (Ed.) Report on the Scientiic Results of the voyage of H. M. S. Challenger. Botany. Vol. 1. III. part 1. Her Majesty’s Stationary Oice, London, 78–89. World checklist of hornworts and liverworts 757 Mitten W (1886a) Some new species of the genus Metzgeria. Journal of the Linnean Society. Botany 22 (145): 241–243. doi: 10.1111/j.1095-8339.1886.tb00467.x Mitten W (1886b) he mosses and hepaticae collected in Central Africa. Journal of the Linnean Society. Botany 22 (146): 298–329. doi: 10.1111/j.1095-8339.1886.tb00649.x Mitten W (1887) Musci. In: hurn EF (Ed.) he botany of the Roraima expedition of 1884. Transactions of the Linnean Society of London. Botany 2 (13): 249–299. Mitten W (1888) Muscineae, hepaticae. In: Balfour IB (Ed.) he botany of Socotra. Transactions of the Royal Society of Edinburgh 31: 1–446. Mitten W (1891) An enumeration of all species of musci and hepaticae recorded from Japan. Transactions of the Linnean Society of London. Botany 3 (3): 153–206. doi: 10.1111/ j.1095-8339.1891.tb00626.x Miyake K (1899) Makinoa, a new genus of hepaticae. Botanical Magazine, Tokyo 13 (144): 21–24. doi: 10.15281/jplantres1887.13.144_21 Mizutani M (1961) A revision of Japanese Lejeuneaceae. Journal of the Hattori Botanical Laboratory 24: 115–302. Mizutani M (1963) On some Indian species of the family Lejeuneaceae described by W. Mitten. Journal of the Hattori Botanical Laboratory 26: 171–184. Mizutani M (1964a) Studies of little known Asiatic species of hepaticae in the Stephani Herbarium. 1. On some little known southeast Asiatic species of the family Lejeuneaceae. Journal of the Hattori Botanical Laboratory 27: 139–148. Mizutani M (1964b) A small collection of New Caledonian hepatics. Journal of the Hattori Botanical Laboratory 27: 131–132. Mizutani M (1965) Studies of little known Asiatic species of hepaticae in the Stephani Herbarium. 2. On some little known southeast Asiatic species of the genus Cololejeunea. Journal of the Hattori Botanical Laboratory 28: 107–121. Mizutani M (1966) Epiphyllous species of Lejeuneaceae from Sabah (North Borneo). Journal of the Hattori Botanical Laboratory 29: 153–170. Mizutani M (1967) Studies of the Himalayan species of Bazzania. Journal of the Hattori Botanical Laboratory 30: 71–90. Mizutani M (1968) Studies of little known Asiatic species of hepaticae in the Stephani Herbarium. 4. Phaeolejeunea, a new genus of Lejeuneaceae. Journal of the Hattori Botanical Laboratory 31: 130–134. Mizutani M (1969) Lejeuneaceae subfamily Ptychanthoideae from Sabah (North Borneo). Journal of the Hattori Botanical Laboratory 32: 129–139. Mizutani M (1970) Lejeuneaceae, subfamilies Lejeuneoideae and Cololejeuneoideae, from Sabah (North Borneo). Journal of the Hattori Botanical Laboratory 33: 225–265. Mizutani M (1971a) Lejeunea from the Himalayan region. Journal of the Hattori Botanical Laboratory 34: 445–457. Mizutani M (1971b) On some Japanese species of Lejeunea. Miscellanea Bryologica et Lichenologica 5 (10/12): 177–180. Mizutani M (1972a) Studies of little known Asiatic species of hepaticae in the Stephani Herbarium. 8. Some little known species of the subfamily Lejeuneoideae of the Lejeuneaceae. Journal of the Hattori Botanical Laboratory 36: 157–162. 758 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Mizutani M (1972b) Studies of little known Asiatic species of hepaticae in the Stephani Herbarium. 7. Some little known species of the subfamily Lejeuneoideae of the Lejeuneaceae. Journal of the Hattori Botanical Laboratory 35: 399–411. Mizutani M (1973) he genus Harpalejeunea from Sabah (North Borneo). Journal of the Hattori Botanical Laboratory 37: 191–203. Mizutani M (1974) Lepidoziaceae, subfamily Lepidozioideae from Sabah (North Borneo). Journal of the Hattori Botanical Laboratory 38: 371–385. Mizutani M (1975) Epiphyllous species of Lejeuneaceae from the Philippines. Journal of the Hattori Botanical Laboratory 39: 255–262. Mizutani M (1976a) Lepidoziaceae, subfamily Lepidozioideae from the Philippines. Journal of the Hattori Botanical Laboratory 40: 447–451. Mizutani M (1976b) Studies of little known Asiatic species of hepaticae in the Stephani Herbarium. 9. Some little known species of the family Lejeuneaceae. Journal of the Hattori Botanical Laboratory 40: 441–446. Mizutani M (1977) Lejeunea magohukui Mizut., sp. nov. Miscellanea Bryologica et Lichenologica 7 (7): 132–134. Mizutani M (1978) Lejeuneaceae from the Ishigaki and Iriomote Islands of Ryukyu Archipelago. Journal of the Hattori Botanical Laboratory 44: 121–136. Mizutani M (1979a) Hepatics from eastern Nepal collected by Himalayan Expedition of Chiba University in 1977. Journal of the Hattori Botanical Laboratory 46: 311–325. Mizutani M (1979b) Notes on the Lejeuneaceae. 2. Some peculiar Asiatic species in the Rijksherbarium, Leiden. Journal of the Hattori Botanical Laboratory 46: 357–372. Mizutani M (1979c) Hepatics from central Nepal collected by the Kochi Himalaya Expedition, 1976. Journal of the Hattori Botanical Laboratory 46: 385–392. Mizutani M (1980) On Cheilolejeunea ontakensis. Miscellanea Bryologica et Lichenologica 8 (7): 146–149. Mizutani M (1981) Notes on the Lejeuneaceae. 5. Some Asiatic species of the genus Ceratolejeunea. Journal of the Hattori Botanical Laboratory 49: 305–318. Mizutani M (1982) Notes on the Lejeuneaceae. 6. Japanese species of the genus Cheilolejeunea. Journal of the Hattori Botanical Laboratory 51: 151–173. Mizutani M (1984a) Notes on the Lejeuneaceae. 7. Calatholejeunea paradoxa, C. lamii (sp. nov.) and Plagiolejeunea zantenii (gen. et sp. nov.). Journal of the Hattori Botanical Laboratory 56: 331–338. Mizutani M (1984b) Notes on the Lejeuneaceae. 8. Japanese species of the subgenus Taeniolejeunea of the genus Cololejeunea. Journal of the Hattori Botanical Laboratory 57: 153–170. Mizutani M (1984c) Notes on the Lejeuneaceae. 9. Cololejeunea lanciloba and its related species in Japan. Journal of the Hattori Botanical Laboratory 57: 427–442. Mizutani M (1985) On some hepatic species from Yakushima I., Japan. Proceedings of the Bryological Society of Japan 4 (3): 35–36. Mizutani M (1986a) Notes on the Lejeuneaceae. 11. Cololejeunea spinosa and its related species in Japan. Journal of the Hattori Botanical Laboratory 60: 439–450. Mizutani M (1986b) Lejeuneaceae from Seram Island, Indonesia. Journal of the Hattori Botanical Laboratory 61: 299–308. World checklist of hornworts and liverworts 759 Mizutani M (1986c) Notes on the Lejeuneaceae. 12. Mastigolejeunea humilis and its related species from Asia. Journal of the Hattori Botanical Laboratory 61: 281–297. Mizutani M (1990) Notes on the Lejeuneaceae. 16. Drepanolejeunea thwaitesiana and its related species from Asia. Journal of the Hattori Botanical Laboratory 68: 367–380. Mizutani M (1992) Notes on the Lejeuneaceae. 17. Lejeunea curviloba and its related species from Japan. Journal of the Hattori Botanical Laboratory 71: 123–132. Mizutani M (1993) Notes on the Lejeuneaceae. 18. Japanese species of the genus Archilejeunea. Journal of the Hattori Botanical Laboratory 73: 175–182. Mizutani M (1994) he second species of Japanese Nowellia. Hikobia 11: 469–470. Mizutani M, Chang KC (1986) A preliminary study of Chinese Lepidoziaceae lora. Journal of the Hattori Botanical Laboratory 60: 419–437. Mizutani M, Hattori S (1957) An etude on the systematics of Japanese Riccardias. Journal of the Hattori Botanical Laboratory 18: 27–64. Montagne JFC (1838) Centurie de plantes cellulaires exotiques nouvelles (suite). Annales des Sciences Naturelles; Botanique (sér. 2) 9: 38–57. Montagne JFC (1839a) lorula boliviensis. In: d’Orbigny A (Ed.) Voyage dans l’Amérique Méridionale. Tome Septième. Bertrand, Paris, 1–119. Montagne JFC (1839b) Sertum patagonicum. In: d’Orbigny A (Ed.) Voyage dans l’Amérique méridionale. Tome septième. Bertrand, Paris, 1–19. Montagne JFC (1839c) Cryptogamae brasilienses. Annales des Sciences Naturelles; Botanique (sér. 2) 12: 42–55. Montagne JFC (1840a) Seconde centurie de plantes cellulaires exotiques nouvelles. Décades VI, VII et VIII. Annales des Sciences Naturelles; Botanique (sér. 2) 14: 321–350. Montagne JFC (1840b) Plantae cellulares. In: Webb PB , Berthelot S (Eds) Histoire Naturelle des Îles Canaries, Tome troisième. Deuxième partie. Phytographia Canariensis, section ultima. Béthuen, Paris, 9–160. Montagne JFC (1842a) Botanique. Plantes cellulaires. In: de la Sagra RDJ (Ed.) Histoire Physique, Politique et Naturelle de l’Ile de Cuba. Arthus Bertrand, Paris, 427–492. doi: 10.5962/bhl.title.51128 Montagne JFC (1842b) Cryptogamae nilgheriensis. Annales des Sciences Naturelles; Botanique (sér. 2) 18: 12–23. Montagne JFC (1843) Quatrième centurie de plantes cellulaires exotiques nouvelles, décades I-VI. Annales des Sciences Naturelles; Botanique (sér. 2) 19: 238–266. Montagne JFC (1844a) Diagnoses muscorum quorundam javanicorum. London Journal of Botany 3: 632–634. Montagne JFC (1844b) Quatrième centurie de plantes cellulaires exotiques nouvelles. Décade VII. Annales des Sciences Naturelles; Botanique (sér. 2) 20: 294–306. Montagne JFC (1845a) Plantae cellulares quas in insulis philippinensibus. London Journal of Botany 4: 3–11. Montagne JFC (1845b) Cinquième centurie de plantes cellulaires exotiques nouvelles. Décades VII à X. Annales des Sciences Naturelles; Botanique (sér. 3) 4: 346–367. 760 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Montagne JFC (1845c) Plantes cellulaires. In: Hombron MM, Jacquinot H (Eds) Voyage au Pole Sud et dans l’Océanie sur les corvettes l’Astrolabe et la Zélée. Botanique, vol. 1. Gide & Cie, Paris, 1–349. doi: 10.5962/bhl.title.6485 Montagne JFC (1846) Botanique, Tome premiere, Cryptogames cellulaires et vasculaires. In: Gaudichaud C (Ed.) Voyage autour du monde execute pendant les années 1836 et 1837 sur la corvette la Bonite commandée par M. Vaillant. Arthus Bertrand, Paris, 1–355. Montagne JFC (1848) Sixième centurie de plantes cellulaires exotiques nouvelles. Décades I et II. Annales des Sciences Naturelles; Botanique (sér. 3) 10: 106–136. Montagne JFC (1849) Sixième centurie de plantes cellulaires nouvelles, tant indigènes qu’exotiques. Décades III à VI. Annales des Sciences Naturelles; Botanique (sér. 3) 11: 33–66. Montagne JFC (1852) Note sur le genre Riella et description d’une espèce nouvelle R. reuteri. Annales des Sciences Naturelles; Botanique (sér. 3) 18: 11–13. Montagne JFC (1855) Cryptogamia guayensis. Annales des Sciences Naturelles; Botanique (sér. 4) 3 (5): 311–329. Montagne JFC (1856a) Huitième centurie de plantes cellulaires nouvelles tant indigènes qu’exotiques, décades I a III. Annales des Sciences Naturelles; Botanique (sér. 4) 6: 179–199. Montagne JFC (1856b) Sylloge generum specierumque cryptogamarum. Sumptibus J.-B. Ballière, Paris, 498 pp. Montagne JFC (1856c) Septième centurie de plantes cellulaires nouvelles. Annales des Sciences Naturelles; Botanique (sér. 4) 5: 331–374. Montagne JFC (1860) Neuvième centurie de plantes cellulaires nouvelles tant indigènes qu’exotiques, décades I et II. Annales des Sciences Naturelles; Botanique (sér. 4) 14: 167–185. Moore D (1877) On irish hepaticae. Proceedings of the Royal Irish Academy (ser. 2) 2: 591–672. Morales MI, Dauphin G (1998) A new species of Cololejeunea (Lejeuneaceae: Cololejeuneoideae) from Panama. Tropical Bryology 14: 133–136. Morales MI, Lücking A (1995) Aphanolejeunea winkleri Morales & A. Lücking, a new species of Lejeuneaceae (Hepaticae, Cololejeuneoideae) from Costa Rica. Nova Hedwigia 60 (1/2): 119–124. Moris JH (1829) Stirpium sardoarum elenchus, fasc. III. Carali, Taurini, 26 pp. Moris JH, De Notaris G (1839) Florula caprariae. Typographia Regia, Torino, 244 pp. Moura O, Ilkiu-Borges AL, Reiner-Drehwald ME (2012) A new species of Lejeunea Lib. (Lejeuneaceae) from Low Várzea forest in lower Amazon (Pará, Brazil). Nova Hedwigia 95 (1/2): 197–202. doi: 10.1127/0029-5035/2012/0033 Müller K (1899) Eine neue Lepidozia-art. Hedwigia 38 (4): 196–200. Müller K (1901a) Vorarbeiten zu einer Monographie der Gattung Scapania. Bulletin de l’Herbier Boissier (sér. 2) 1 (6): 593–614. Müller K (1901b) Ueber die im Jahre 1900 in Baden gesammelten Lebermoose. Beihefte zum Botanischen Centralblatt 10 (4/5): 213–223. Müller K (1902) Neue Bürger der Badischen Lebermoos-Flora. Mitteilungen des Badischen Botanischen Vereins 4 (182/183): 283–288. Müller K (1903) Neue und kritische Lebermoose. Bulletin de l’Herbier Boissier (sér. 2) 3 (1): 34–44. World checklist of hornworts and liverworts 761 Müller K (1904) Über die in Baden in den Jahren 1902 und 1903 gesammelten Lebermoose. Beihefte zum Botanischen Centralblatt 17 (2): 211–233. Müller K (1905) Monographie der Lebermoosgattung Scapania Dum. Nova Acta Academiae Caesareae Leopoldino-Carolinae Germanicae Naturae Curiosorum 83: 1–312. Müller K (1907a) Die Lebermoose (Dr. L. Rabenhorst’s Kryptogamen-Flora von Deutschland, Oesterreich und der Schweiz, 2 Aul., 6 Band), 1 Abth., 5 Lieferung. Eduard Kummer, Leipzig, 257–320. Müller K (1907b) Die Lebermoose (Dr. L. Rabenhorst’s Kryptogamen-Flora von Deutschland, Oesterreich und der Schweiz, 2 Aul., 6 Band), 1 Abth., 3 Lieferung. Eduard Kummer, Leipzig, 129–192. Müller K (1912) Die Lebermoose (Dr. L. Rabenhorst’s Kryptogamen-Flora von Deutschland, Oesterreich und der Schweiz, 2 Aul., 6 Band), 2 Abth., 16 Lieferung. Eduard Kummer, Leipzig, 81–144. Müller K (1913a) Die Lebermoose (Dr. L. Rabenhorst’s Kryptogamen-Flora von Deutschland, Oesterreich und der Schweiz, 2 Aul., 6 Band), 2 Abth., 17 Lieferung. Eduard Kummer, Leipzig, 145–208. Müller K (1913b) Die Lebermoose (Dr. L. Rabenhorst’s Kryptogamen-Flora von Deutschland, Oesterreich und der Schweiz, 2 Aul., 6 Band), 2 Abth., 18 Lieferung. Eduard Kummer, Leipzig, 209–272. Müller K (1915a) Die Lebermoose (Dr. L. Rabenhorst’s Kryptogamen-Flora von Deutschland, Oesterreich und der Schweiz, 2 Aul., 6 Band), 2 Abth., 22 Lieferung. Eduard Kummer, Leipzig, 465–528. Müller K (1915b) Die Lebermoose (Dr. L. Rabenhorst’s Kryptogamen-Flora von Deutschland, Oesterreich und der Schweiz, 2 Aul., 6 Band), 2 Abth., 21 Lieferung. Eduard Kummer, Leipzig, 385–464. Müller K (1916) Die Lebermoose (Dr. L. Rabenhorst’s Kryptogamen-Flora von Deutschland, Oesterreich und der Schweiz, 2 Aul., 6 Band), 2 Abth., 27 Lieferung. Eduard Kummer, Leipzig, 785–848. Müller K (1940) Beiträge zur Systematik der Lebermoose. Hedwigia 79 (1/2): 72–80. Müller K (1941) Beiträge zur Systematik der Lebermoose II. Hedwigia 80 (1/2): 90–118. Müller K (1942) Revision der europäischen Arten der Lebermoosgattung Chiloscyphus auf Grund des Chromosomensatzes und von Kulturen. Berichte der Deutschen Botanischen Gesellschaft 59 (10): 428–436. Müller K (1947) Morphologische Untersuchungen zur Aufklärung einiger europäischer Lebermoose. Beiträge zur Kryptogamenlora der Schweiz 10 (2): 1–55. Müller K (1951a) Bestimmungs- und Nomenklaturberichtigungen zum Schifnerischen Exsikkatenwerk Hepaticae Europaeae exsiccatae. Feddes Repertorium Specierum Novarum Regni Vegetabilis 54 (2/3): 207–222. doi: 10.1002/fedr.19510540205 Müller K (1951b) Neue Lebermoose. Revue Bryologique et Lichénologique 20 (1/2): 176–178. Müller K (1953) Hepatikologische Notizen. Revue Bryologique et Lichénologique 22 (3/4): 131–140. Müller K (1955) Lebermoose aus Südamerika. Feddes Repertorium Specierum Novarum Regni Vegetabilis 58: 59–74. 762 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Müller K (1958) Die Lebermoose Europas (Dr. L. Rabenhorst’s Kryptogamen-Flora von Deutschland, Österreich und der Schweiz, 3 Aul., 6 Band), 2 Abth., 9 Lieferung. Akademische Verlagsgesellschaft, Leipzig, 1221–1365. Müller F (2007) Meinungeria mouensis (Lepidoziaceae), a new genus and species from New Caledonia. Bryologist 110 (3): 494–499. doi: 10.1639/0007-2745(2007)110[494:MML ANG]2.0.CO;2 Müller F (2013) Pleurozia pocsii sp. nov. (Pleuroziaceae) from New Caledonia. Polish Botanical Journal 58 (1): 49–53. doi: 10.2478/pbj-2013-0005 Müller F, Pócs T (2007) A contribution to the knowledge of epiphyllous bryophytes of Bioko Island (Equatorial Guinea), including additional remarks on non-epiphyllous species. Journal of Bryology 29 (2): 81–94. doi: 10.1179/174328207X186803 Mutke J, Barthlott W (2005) Patterns of vascular plant diversity at continental to global scales. Biologiske Skrifter 55: 521–531. Nadeaud J (1873) Énumération des plantes indigènes de l’île de Tahiti. F. Savy, Paris, 86 pp. Nath V, Singh AP (2006) Frullania udarii sp. nov. – A new species from Meghalaya, India. Current Science 91 (6): 744–746. Na-halang O (1980) A revision of the genus Riccia (Hepaticae) in Australia. Brunonia 3 (1): 61–140. doi: 10.1071/BRU9800061 Nebel M, Söderström L, Hagborg A, von Konrat MJ (2013) Notes on Early Land Plants Today. 28. Transfers of some taxa to Lobatiriccardia (Aneuraceae, Marchantiophyta). Phytotaxa 81 (1): 10–11. doi: 10.11646/phytotaxa.81.1.4 Nees CG (1820) Horae physicae berolinenses. Adolphi Marcus, Bonnae [Bonn], 123 pp. Nees CG (1830) Enumeratio plantarum cryptogamicarum Javae et insularum adiacentium. Fasciculus prior, hepaticas complectens, ab editore illustratas. Grass, Barth & Co., Breslau, 86 pp. Nees CG (1831) Berichtigungen zur Enumeratio plantarum cryptogamicarum Javae. Linnaea 6 (4): 602–623. Nees CG (1833a) Hepaticae Hedw. In: Martius CFP (Ed.) Flora brasiliensis. Sumptibus J. G. Cottae, Stuttgart, 294–390. doi: 10.5962/bhl.title.454 Nees CG (1833b) Beiträge zur Naturgeschichte der deutschen Lebermoose (Schluss). Flora 16 (26): 401–414. Nees CG (1833c) Naturgeschichte der Europäischen Lebermoose, vol. 1. August Rücker, Berlin, 347 pp. Nees CG (1836) Naturgeschichte der Europäischen Lebermoose, vol. 2. August Rücker, Berlin, 499 pp. Nees CG (1838a) Naturgeschichte der Europäischen Lebermoose, vol. 4. Grass, Barth & Co., Breslau, 539 pp. Nees CG (1838b) Naturgeschichte der Europäischen Lebermoose, vol. 3. Grass, Barth & Co., Breslau, 593 pp. Nees CG, Bischof GW (1830) Lunularia alpina und Corsinia lamellosa, zwei neue europäische Lebermoose. Flora 13 (25): 393–404. Nees CG, Montagne JFC (1836) Jungermanniearum herbarii Montagneani species. Annales des Sciences Naturelles; Botanique (sér. 2) 5: 52–72. World checklist of hornworts and liverworts 763 Nicholson WE (1925) Notes on some New Zealand species of Frullania. Bryologist 28 (2): 17–19. doi: 10.2307/3238374 Nicholson WE (1942) Some hepatics from the Hawaiian Islands. Revue Bryologique et Lichénologique 13: 142–144. Nicholson WE, Herzog T, Verdoorn F (1930) Hepaticae. In: Handel-Mazzetti HM (Ed.) Symbolae Sinicae, Part 5. Hepaticae. Springer Verlag, Berlin, 1–60. Nordstedt O (1874) Lärda sällskaps sammanträden. Botaniska Notiser 1874: 155–157. Ohnishi N, Deguchi H (2003) A new species of Schistochila (Hepaticae) from East Asia. Bryologist 106 (3): 451–453. doi: 10.1639/12 Okamura S (1911) Neue Beiträge zur Mooslora Japans III. Botanical Magazine, Tokyo 25 (293): 159–162. doi: 10.15281/jplantres1887.25.293_159 Okamura S (1916) Contributiones novae ad loram bryophyton japonicam, pars secunda. Journal of the College of Science, Imperial University of Tokyo 38 (4): 1–100. O’Neill KP (2000) Role of bryophyte-dominated ecosystems in the global carbon budget. In: Shaw AJ, Goinet B (Eds) Bryophyte biology. Cambridge University Press, Cambridge, 344–368. doi: 10.1017/cbo9781139171304.012 Onraedt M (1977) Bryophytes des îles mascareno-malgaches et Seychelles. III. Hepaticae: Bazzania. Bulletin du Jardin Botanique National de Belgique 47 (1/2): 139–144. doi: 10.2307/3667988 Onraedt M (1978) Bryophytes de Sri Lanka (Ceylon). Revue Bryologique et Lichénologique 44 (1): 77–82. Onraedt M (1979) Bryophytes de Sri Lanka (Ceylan) III. Cololejeunea ceylanica Onr. et Cololejeunea hinidumae Onr., espèces nouvelles. Acta Botanica Academiae Scientiarum Hungaricae 25 (1/2): 107–111. Onraedt M (1982) Une nouvelle espèce de Taxilejeunea des îles Galápagos. Miscellanea Bryologica et Lichenologica 9 (6): 117–119. Onraedt M (1989) Quatre Cololejeunea (Hepaticae) nouveaux des Philippines. Bulletin du Jardin Botanique National de Belgique 59 (3/4): 433–438. doi: 10.2307/3668357 Osbeck P (1757) Dagbok öfwer en Ostindisk resa åren 1750, 1751, 1752. Lor. Ludv. Greing, Stockholm, 376 pp. Pagán FM (1939a) A preliminary list of the hepaticae of Puerto Rico including Vieques and Mona Island. Bryologist 42 (1): 1–12. doi: 10.2307/3239058 Pagán FM (1939b) A preliminary list of the hepaticae of Puerto Rico including Vieques and Mona Island (continued). Bryologist 42 (2): 37–50. doi: 10.2307/3239008 Pagán FM (1942a) A new species of Dendroceros from Puerto Rico. Bryologist 45 (4): 111–115. doi: 10.2307/3239434 Pagán FM (1942b) Catalogue of the hepaticae of Guadeloupe. Bryologist 45 (4): 76–110. doi: 10.2307/3239433 Palisot de Beauvois AMFJ (1805a) Asterella. In: Cuvier F (Ed.) Dictionnaire des sciences naturelles dans lequel on traite méthodiquement des diferents êtres de la nature, fasc. 3. F. G. Levrault, Paris, 257–258. Palisot de Beauvois AMFJ (1805b) Flore d’Oware, vol. 1 (3). Fain et Compagnie, Paris, 17–82. 764 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Pandé SK (1960) he Anthocerotales, some aspects of their systematics and morphology. Proceedings of the Indian Science Congress Association 47 (2): 90–104. Pandé SK, Misra RN (1943) Studies in Indian hepaticae. II. On epiphyllous liverworts of India and Ceylon (1). Journal of the Indian Botanical Society 22 (2/4): 159–169. Pandé SK, Srivastava KP (1955) On a species of Cephalozia Dum. from India: C. herzogiana Pandé et Srivastava sp. nov. Feddes Repertorium Specierum Novarum Regni Vegetabilis 58: 75–79. Pandé SK, Udar R (1957) A species of Riccia, R. aravalliensis Pandé et Udar sp. nov., from Mt. Abu, Rajasthan, India. Journal of the Indian Botanical Society 36 (3): 248–253. Pandé SK, Udar R (1958) Genus Riccia in India - II. Species of Riccia from South India with description of a new species and notes on the synonyms of some recently described ones. Proceedings of the National Institute of Science of India, part B, Biological Sciences 24 (2): 79–88. Pandé SK, Udar R (1959) Genus Riccia in India. III. Species of Riccia from the East Himalaya Territory with description of a new species, R. attenuata Pandé sp. nov. Proceedings of the National Institute of Science of India, part B, Biological Sciences 25 (2): 90–100. Pandé SK, Srivastava KP, Ahmad S (1957) Epiphyllous liverworts of India and Ceylon. II. Journal of the Indian Botanical Society 36 (3): 335–347. Parihar NS (1962) An annotated revised census of Indian hepatics. University of Allahabad Studies, Botany Section 1961-2: 1–56. Paris EG (1906a) Hépatiques de Nouvelle-Calédonie. Revue Bryologique 33 (2): 27–29. Paris EG (1906b) Muscinées de l’Afrique occidentale française. Revue Bryologique 33 (3): 38–42. Paris EG (1907) Muscinées de l’Asie orientale (6e article). Revue Bryologique 34 (3): 41–49. Parolly G, Kürschner H, Schäfer-Verwimp A, Gradstein SR (2004) Cryptogams of the Reserva Biológica San Francisco (Province Zamora-Chinchipe, southern Ecuador). III. Bryophytes – Additions and new species. Cryptogamie, Bryologie 25 (3): 271–289. Paton JA (1973) Taxonomic studies in the genus Fossombronia Raddi. Journal of Bryology 7 (3): 243–252. doi: 10.1179/jbr.1973.7.3.243 Paton JA (1974) Fossombronia imbriata sp. nov. Journal of Bryology 8 (1): 1–4. doi: 10.1179/ jbr.1974.8.1.1 Paton JA (1979a) Anthoceros agrestis, a new name for A. punctatus var. cavernosus sensu Prosk. 1958, non (Nees) Gottsche et al. Journal of Bryology 10 (3): 257–261. doi: 10.1179/ jbr.1979.10.3.257 Paton JA (1979b) Plagiochila britannica. a new species in the British Isles. Journal of Bryology 10 (3): 245–256. doi: 10.1179/jbr.1979.10.3.245 Paton JA (1994) Fossombronia pusilla (L.) Nees var. maritima Paton elevated to the rank of species. Journal of Bryology 18 (2): 366–368. doi: 10.1179/jbr.1994.18.2.366 Paton AJ (2013) From working list to online lora of all known plants—looking forward with hind-sight. Annals of the Missouri Botanical Garden 99 (2): 206–213. doi: 10.3417/2011115 Paton JA, Perry AR (1995) Leiocolea itzgeraldiae sp. nov. in Britain and Ireland. Journal of Bryology 18 (3): 469–478. doi: 10.1179/jbr.1995.18.3.469 World checklist of hornworts and liverworts 765 Paton JA, Sheahan MC (2006) Lophocolea brookwoodiana (Jungermanniales: Geocalycaceae), a new species in Britain. Journal of Bryology 28 (3): 163–166. doi: 10.1179/174328206X119952 Paton AJ, Brummitt N, Govaerts R, Harman K, Hinchclife S, Allkin R, Lughadha EN (2008) Towards Target 1 of the Global Strategy for Plant Conservation: A working list of all known plant species—Progress and prospects. Taxon 57 (2): 602–611. doi: 10.2307/25066027 Pätsch R, Hentschel J, Linares-Palomino R, Zhu R-L, Heinrichs J (2010) Diversiication and taxonomy of the liverwort Jubula Dumort. (Jungermanniopsida: Porellales) inferred from nuclear and chloroplast DNA sequences. Systematic Botany 35 (1): 6–12. doi: 10.1600/036364410790862515 Pearson WH (1886) Hepaticae natalensis. Forhandlinger i Videnskabs-Selskabet i Kristiania 1886 (3): 1–20. Pearson WH (1887a) Blepharostoma palmatum Lindb. Journal of Botany, British and Foreign 25: 193–195. Pearson WH (1887b) Hepaticae knysnanae. Forhandlinger i Videnskabs-Selskabet i Kristiania 1887 (9): 1–16. Pearson WH (1890) List of Canadian hepaticae. WM. Foster Brown & Co., Montreal, 28 pp. doi: 10.5962/bhl.title.17936 Pearson WH (1891) Frullaniae madagascariensis. Forhandlinger i Videnskabs-Selskabet i Kristiania 1890 (2): 1–9. Pearson WH (1892) Lejeuneae madagascariensis. Forhandlinger i Videnskabs-Selskabet i Kristiania 1892 (8): 3–9. Pearson WH (1893) Hepaticae madagascariensis. Forhandlinger i Videnskabs-Selskabet i Kristiania 1892 (14): 1–12. Pearson WH (1894) Frullania microphylla. Journal of Botany, British and Foreign 32: 328–330. Pearson WH (1900) he hepaticae of the British Isles, vol. I text (fasc. 6-15). Lovell Reeve & Co. Ltd., London, 97–288. Pearson WH (1922a) Notes on a collection of hepaticae from the Belgian Congo (ex herb R. Naveau, Antwerp). Natuurwetenschappelijk Tijdschrift 4 (5/6): 118–143. Pearson WH (1922b) A systematic account of the plants collected in New Caledonia and Isle of Pines. Part III. Cryptogams (Hepaticae-Fungi). Hepaticae. Journal of the Linnean Society. Botany 46 (305): 13–44. doi: 10.1111/j.1095-8339.1922.tb00474.x Pearson WH (1923) Notes on a collection of New Zealand hepatics. University of California Publications in Botany 10 (4): 307–370. Pearson WH (1924a) Hepaticae. In: Setchell WA (Ed.) American Samoa: Part. 1. Vegetation of Tutuila Island. , Washington, 131–151. Pearson WH (1924b) Notes on a collection of hepaticae from Mount Elgon, East Africa, made by Dr. G. Lindblom in 1920. Arkiv för Botanik 19 (5): 1–16. Pearson WH (1931a) Notes on a collection of hepaticae made by Mr. Saxby on the West Coast of Africa. Annales de Cryptogamie Exotique 4 (2): 61–71. Pearson WH (1931b) Notes on a collection of hepaticae from Jamaica. Annales Bryologici 4: 95–112. 766 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Pearson LC (1995) he diversity and evolution of plants. CRC Press, New York, 646 pp. Peck CH (1866) List of mosses of the state of New York. Report (Annual) of the Regents of the University of the State of New York on the Condition of the State Cabinet of Natural History 19: 42–70. doi: 10.5962/bhl.title.59550 Peng T, Zhu R-L (2014) A revision of the genus Notothylas (Notothyladaceae, Anthocerotophyta) in China. Phytotaxa 156 (3): 156–164. doi: 10.11646/phytotaxa.156.3.6 Peralta DF, Reiner-Drehwald ME (2013) Cheilolejeunea laciniata (Lejeuneaceae, Marchantiophyta), a new species from Southeastern Brazil. Bryologist 116 (1): 53–57. doi: 10.1639/0007-2745-116.1.053 Perold SM (1989a) Spore-wall ornamentation as an aid in identifying the southern African species of Riccia. Journal of the Hattori Botanical Laboratory 67: 109–201. Perold SM (1989b) Studies in the genus Riccia (Marchantiales) from southern Africa. 13. A new species, R. hantamensis, in section Pilifer, and a new record for R. alatospora. Bothalia 19 (2): 157–160. doi: 10.4102/abc.v19i2.953 Perold SM (1989c) Studies in the genus Riccia (Marchantiales) from southern Africa. 11. R. montana and R. alboporosa, a further two new white-scaled species of the group ‘Squamatae’. Bothalia 19 (1): 9–16. doi: 10.4102/abc.v19i1.934 Perold SM (1990a) Studies in the genus Riccia (Marchantiales) from southern Africa. 16. R. albomarginata and R. simii sp. nov. Bothalia 20 (1): 31–39. doi: 10.4102/abc.v20i1.891 Perold SM (1990b) Studies in the genus Riccia (Marchantiales) from southern Africa. 18. New species in the section Pilifer from NW Cape: R. furfuracea, R. vitrea and R. namaquensis. Bothalia 20 (2): 175–183. doi: 10.4102/abc.v20i2.912 Perold SM (1990c) Studies in the genus Riccia (Marchantiales) from southern Africa. 19. Two new species: R. pulveracea, section Pilifer and R. bicolorata, section Riccia, group ‘Squamatae’. Bothalia 20 (2): 185–190. doi: 10.4102/abc.v20i2.913 Perold SM (1990d) Studies in the genus Riccia (Marchantiales) from southern Africa. 21. R. stricta, R. purpurascens and R. luitans, subgenus Ricciella. Bothalia 20 (2): 197–206. doi: 10.4102/abc.v20i2.915 Perold SM (1990e) Studies in the genus Riccia (Marchantiales) from southern Africa. 17. hree new species in the section Pilifer: R. elongata, R. ampullacea and R. trachyglossum. Bothalia 20 (2): 167–174. doi: 10.4102/abc.v20i2.911 Perold SM (1991a) Studies in the genus Riccia (Marchantiales) from southern Africa. 22. R. rubricollis now validated, typiied and described. Bothalia 21 (1): 51–54. doi: 10.4102/abc.v21i1.858 Perold SM (1991b) Two new species of the liverwort genus Riccia L. from tropical Africa: R. somaliensis and R. erubescens. Journal of Bryology 16 (3): 367–377. doi: 10.1179/ jbr.1991.16.3.367 Perold SM (1993) Studies in the Marchantiales (Hepaticae) from Southern Africa.2. he genus Athalamia and A. spathysii; the genus Oxymitra and O. cristata. Bothalia 23 (2): 207–214. doi: 10.4102/abc.v23i2.804 Perold SM (1994) Studies in the Marchantiales (Hepaticae) from southern Africa. 7. he genus Cryptomitrium (Aytoniaceae) and C. oreades sp. nov. Bothalia 24 (2): 149–152. doi: 10.4102/abc.v24i2.764 World checklist of hornworts and liverworts 767 Perold SM (1995) A survey of the Ricciaceae of tropical Africa. Fragmenta Floristica et Geobotanica 40 (1): 53–91. Perold SM (1997a) Studies in the liverwort genus Fossombronia (Metzgeriales) from southern Africa. 1. hree new species from Northern Province, Gauteng and Mpumalanga. Bothalia 27 (1): 17–27. doi: 10.4102/abc.v27i1.649 Perold SM (1997b) A new species of the liverwort genus Riccia L. from Bioko Island, R. biokoensis Perold. Nova Hedwigia 64 (1/2): 243–248. Perold SM (1998a) Fossombronia rwandaensis, a new species from tropical Africa. Bothalia 28 (1): 45–49. doi: 10.4102/abc.v28i1.614 Perold SM (1998b) Studies in the liverwort genus Fossombronia (Metzgeriales) from southern Africa. 6. New species from Lesotho, Swaziland and Mpumalanga and new records from Lesotho. Bothalia 28 (2): 159–165. doi: 10.4102/abc.v28i2.633 Perold SM (1998c) Studies in the liverwort genus Fossombronia (Metzgeriales) from southern Africa. 5. A new species from Northern and Western Cape. Bothalia 28 (1): 1–5. doi: 10.4102/abc.v28i1.606 Perold SM (1999a) Flora of Southern Africa. Hepatophyta. Part 1: Marchantiopsida. Fascicle 1: Marchantiidae. National Botanical Institute, Pretoria, 252 pp. Perold SM (1999b) Studies in the liverwort genus Fossombronia (Metzgeriales) from southern Africa. 9. A new species from Mpumalanga and KwaZulu-Natal, with notes on other species. Bothalia 29 (1): 77–82. doi: 10.4102/abc.v29i1.573 Perold SM (1999c) Studies in the liverwort genus Fossombronia (Metzgeriales) from southern Africa. 8. F. elsieae and F. spinosa, two new Western Cape species with spinose spores. Bothalia 29 (1): 25–33. doi: 10.4102/abc.v29i1.569 Perold SM (1999d) Studies in the liverwort genus Fossombronia (Metzgeriales) from southern Africa. 10. hree new species from Northern and Western Cape. Bothalia 29 (1): 83–93. doi: 10.4102/abc.v29i1.574 Perold SM (2001a) Studies in the genus Riccia (Marchantiales) from southern Africa. 25. A new species in subgenus Ricciella, section Ricciella. Bothalia 31 (1): 151–154. doi: 10.4102/abc. v31i2.513 Perold SM (2001b) Studies in the liverwort genus Fossombronia (Metzgeriales) from southern Africa. 11. F. zuurbergensis, a new species from Eastern Cape and new records for the area. Bothalia 31 (1): 25–29. doi: 10.4102/abc.v31i1.495 Perold SM (2001c) Notes on African plants: Fossombroniaceae. Fossombronia nyikaensis, a new species from Malawi. Bothalia 31 (1): 48–52. doi: 10.4102/abc.v31i1.506 Perold SM (2003) Sauteria nyikaensis, a new liverwort species from Malawi. Bothalia 33 (2): 165–171. doi: 10.4102/abc.v33i2.450 Perold SM (2004) Studies in the genus Riccia (Marchantiales) from southern Africa. 26. A new species in section Pilifer, Riccia radiata, is described. Bothalia 34 (1): 23–26. doi: 10.4102/ abc.v34i1.402 Perold SM (2005) Riccia mamrensis, a new species from Western Cape, South Africa. Cryptogamie, Bryologie 26 (1): 67–72. Persson H (1946) Some Alaskan and Yukon bryophytes. Bryologist 49 (2): 41–58. doi: 10.2307/3239692 768 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Persson H (1953) Hodgsonia nov. gen. (Hepaticae). Stockholm, 1 pp. Persson H (1954) On Neohodgsonia H. Perss. the new hepatic genus from new Zealand and Tristan da Cunha. Botaniska Notiser 107 (1): 39–44. Persson H (1955) Remarks on the Porella pinnata group. Archivum Societatis Zoologicae Botanicae Fennicae “Vanamo”, suppl. 9: 225–231. Persson H, Grolle R (1961) Roivainenia Persson, eine neue Gattung der Lophoziaceae. Nova Hedwigia 3 (1): 43–46. Pfeifer LKG (1855) Flora von Niederhessen und Münden, Zweiter Band. heodor Fischer, Kassel, 252 pp. Pfeifer T, Frey W, Stech M (2002) A new species of Treubia (Treubiaceae, Hepaticophytina) from New Zealand based on molecular evidence. Studies in austral temperate rain forest bryophytes 20. Nova Hedwigia 75 (1/2): 241–253. doi: 10.1127/0029-5035/2002/0075-0241 Philibert H (1890) Sur la fructiication du Marsupella revoluta Dumortier. Revue Bryologique 17 (3): 33–34. Piippo S (1984a) Bryophyte lora of the Huon Peninsula, Papua New Guinea. III. Haplomitriaceae, Lepicoleaceae, Herbertaceae, Pseudolepicoleaceae, Trichocoleaceae, Schistochilaceae, Balantiopsidaceae, Pleuroziaceae and Porellaceae (Hepaticae). Annales Botanici Fennici 21 (1): 21–48. Piippo S (1984b) Bryophyte lora of the Huon Peninsula, Papua New Guinea. VI. Lepidoziaceae subfam. Lepidozioideae, Calypogeiaceae, Adelanthaceae, Cephaloziaceae subfam. Cephalozioideae and subfam. Odontoschismatoideae and Jubulaceae (Hepaticae). Annales Botanici Fennici 21 (4): 309–335. Piippo S (1985a) Bryophyte lora of the Huon Peninsula, Papua New Guinea. XII. Geocalycaceae (Hepaticae). Acta Botanica Fennica 131: 129–167. Piippo S (1985b) Bryophyte lora of the Huon Peninsula, Papua New Guinea. XIII. Arachniopsis and Kurzia (Lepidoziaceae subfam. Lepidozioideae, Hepaticae). Acta Botanica Fennica 131: 169–179. Piippo S (1986a) A monograph of Lepidolejeunea and Luteolejeunea (Lejeuneaceae: Hepaticae). Acta Botanica Fennica 132: 1–69. Piippo S (1986b) Bryophytes from Frieda River, East and West Sepik provinces, Papua New Guinea. II. Hepaticae (Haplomitriaceae-Frullaniaceae). Annales Botanici Fennici 23 (1): 1–10. Piippo S (1987) Hepatics from Papua New Guinea collected by Dr. G. Shea. Journal of the Hattori Botanical Laboratory 63: 493–498. Piippo S (1988) Bryophyte lora of the Huon Peninsula, Papua New Guinea. XXI. Lepicolea norrisii (Lepicoleaceae, Hepaticae). Annales Botanici Fennici 25 (1): 55–57. Piippo S (1989a) he bryophytes of Sabah (North Borneo) with special reference to the BRYOTROP transect of Mount Kinabalu. III. Geocalycaceae (Hepaticae). Willdenowia 18 (2): 513–527. Piippo S (1989b) Bryophyte lora of the Huon Peninsula, Papua New Guinea. XXX. Plagiochilaceae (Hepaticae). Annales Botanici Fennici 26 (2): 183–236. Piippo S (1990) Annotated catalogue of Chinese hepaticae and anthocerotae. Journal of the Hattori Botanical Laboratory 68: 1–192. World checklist of hornworts and liverworts 769 Piippo S (1991) Bryophyte lora of the Huon Peninsula, Papua New Guinea. XXXIX. Fossombronia (Fossombroniaceae) and Metzgeria (Metzgeriaceae). Acta Botanica Fennica 143: 1–22. Piippo S (1992) Bryophyte lora of the Huon peninsula, Papua New Guinea. LI. Additions and corrections to the Geocalycaceae (Hepaticae). Annales Botanici Fennici 29 (3): 243–248. Piippo S (1993a) Hepatics from the Solomon Islands. 1. Nova Hedwigia 56 (3/4): 355–365. Piippo S (1993b) Bryophyte lora of the Huon Peninsula, Papua New Guinea. LIV. Anthocerotophyta. Acta Botanica Fennica 148: 27–51. Piippo S (1993c) On the taxonomy and nomenclature of SW Asiatic Geocalycaceae (Hepaticae). Annales Botanici Fennici 30 (3): 195–203. Piippo S (1997) Two new Plagiochila species (Plagiochilaceae, Hepaticae) from Yunnan, China. Annales Botanici Fennici 34 (4): 281–284. Piippo S (1998) Bryophytes from Frieda River, East and West Sepik Provinces, Papua New Guinea. IV. Chiloscyphus koponenii sp. nov. (Geocalycaceae). Annales Botanici Fennici 35 (1): 55–57. Piippo S (1999) Mesoceros porcatus, a new hornwort from Yunnan. Haussknechtia, Beiheft 9: 279–282. Piippo S, Tan BC (1992) Novelties for the Philippine hepatic lora. Journal of the Hattori Botanical Laboratory 72: 117–126. Piippo S, He X, Juslén A, Tan BC, Murphy DH, Pócs T (2002) Hepatic and hornwort lora of Singapore. Annales Botanici Fennici 39 (2): 101–127. Piippo S, Mamontov Y, Potemkin AD (2014) Bryophyte lora of the Huon Peninsula, Papua New Guinea. LXXVI. Conoscyphus koponenii, spec. nov. (Hepaticae, Jungermanniaceae, Chandonanthoideae). Acta Bryolichenologica Asiatica 5: 19–24. Pitard L, Corbière L (1907) Hépatiques des îles Canaries. Mémoires de la Société Botanique de France 7: 28–44. Pitcairn CER, Fowler D, Grace J (1995) Deposition of ixed atmospheric nitrogen and foliar nitrogen content of bryophytes and Calluna vulgaris (L.) Hull. Environmental Pollution 88 (2): 193–205. doi: 10.1016/0269-7491(95)91444-P Pócs T (1968) he genus Porella in Vietnam. Journal of the Hattori Botanical Laboratory 31: 65–93. Pócs T (1969) A short survey of the Bazzania of North Viet-Nam. Journal of the Hattori Botanical Laboratory 32: 79–94. Pócs T (1971) Issledovanie gornoj briolory provincii Hažhang (DRV). Hepaticae. [An investigation of the mountain bryolora of the Hashang province (Democratic Republic of Vietnam). Hepaticae]. Botanicheskij Zhurnal. Moscow & Leningrad 56 (5): 670–677. Pócs T (1975) New or little-known epiphyllous liverworts I. Cololejeunea from tropical Africa. Acta Botanica Academiae Scientiarum Hungaricae 21 (3/4): 353–375. Pócs T (1980a) he epiphytic biomass and its efect on the water balance of two rain forest types in the Uluguru Mountains (Tanzania, East Africa). Acta Botanica Academiae Scientiarum Hungaricae 26 (1/2): 143–167. 770 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Pócs T (1980b) New or little known epiphyllous liverworts. II. hree new Cololejeunea from East Africa. Journal of the Hattori Botanical Laboratory 48: 305–320. Pócs T (1984a) New or little known epiphyllous liverworts. III. he genus Aphanolejeunea Evans in Tropical Africa. Cryptogamie: Bryologie, Lichénologie 5 (3): 239–267. Pócs T (1984b) Synopsis of the African Lepidoziaceae K. Müll. In: Váňa J (Ed.) Proceedings of the third meeting of the bryologists from Central and East Europe. Univerzita Karlova, Praha, 107–119. Pócs T (1985) East african bryophytes, VII. he hepaticae of the Usambara rain forest project expedition, 1982. Acta Botanica Hungarica 31 (1/4): 113–133. Pócs T (1988) Plagiochila rudolii sp. nov., from Tanzania, East Africa. Beihefte zur Nova Hedwigia 90: 223–233. Pócs T (1993) New or little known epiphyllous liverworts. IV. Two new Cololejeuneoideae from the Comoro Archipelago. Journal of the Hattori Botanical Laboratory 74: 45–58. Pócs T (1994a) East African bryophytes, XIII. Bryophytes from the Bale Mountains, SE Ethiopia. 2. Hepaticae. Fragmenta Floristica et Geobotanica 39 (1): 221–233. Pócs T (1994b) Taxonomic results of the BRYOTROP expedition to Zaïre and Rwanda. 28. Lejeuneaceae, a ramicolous collection. Tropical Bryology 9: 131–136. Pócs T (1994c) Taxonomic results of the BRYOTROP expedition to Zaïre and Rwanda. 27. Lepidoziaceae. Tropical Bryology 9: 123–130. Pócs T (1994d) New or little known epiphyllous liverworts. V. Aphanolejeunea collected by Barbara M. hiers in Australia and Papua New Guinea. Hikobia 11: 457–462. Pócs T (1995) East African bryophytes, XIV. Hepaticae from the Indian Ocean islands. Fragmenta Floristica et Geobotanica 40 (1): 251–277. Pócs T (1997a) New or little known epiphyllous liverworts. VII. Two new Lejeuneaceae species from the Mascarene Islands. Cryptogamie: Bryologie, Lichénologie 18 (3): 195–205. Pócs T (1997b) New or little known epiphyllous liverworts, VI. Papillolejeunea gen. nov. from Papua New Guinea. Tropical Bryology 13: 1–18. Pócs T (1999) Trachylejeunea grolleana, a new representative of the neotropical subgenus Hygrolejeuneopsis in Madagascar. Haussknechtia, Beiheft 9: 283–290. Pócs T (2001) East African bryophytes, XVI. New taxa of Lejeuneoideae (Lejeuneaceae) collected in Manongarivo Special Reserve, NW Madagascar. Candollea 56 (1): 69–78. Pócs T (2002a) East African bryophytes, XVII. Cololejeunea attilana sp. nov. from Tanzania. In: Salamon-Albert É (Ed.) Magyar Botanikai Kutatások az Ezredfordulón Tanulmányok Borhidi Attila 70. Születésnapja Tiszteletére. PTE Növénytani Tanszék, Pécs, 185–192. Pócs T (2002b) New or little known epiphyllous liverworts. IX. Two new neotropical Cololejeunea species. Acta Botanica Hungarica 44 (3/4): 371–382. doi: 10.1556/ABot.44.2002.3-4.13 Pócs T (2002c) East African bryophytes. XVIII. Two new Lejeuneaceae (Hepaticae) from Aberdare Mountains (Kenya). Polish Botanical Journal 47 (1): 11–20. Pócs T (2004) New or little known epiphyllous liverworts. XI. Otolejeunea subana sp. nov. from Madagascar. Acta Academiae Paedagogicae Agriensis, Sectio Biologiae 25: 49–57. Pócs T (2006a) East African bryophytes. XXII. Two new species, Plagiochila artsii and Plagiochila hiroshiana (Plagiochilaceae, Marchantiophyta). Journal of the Hattori Botanical Laboratory 100: 333–350. World checklist of hornworts and liverworts 771 Pócs T (2006b) New or little-known epiphyllous liverworts. X. On two new neotropical Diplasiolejeunea species. Bryologist 109 (3): 408–414. doi: 10.1639/0007-2745(2006)109[408:NOLELX]2.0.CO;2 Pócs T (2006c) East African bryophytes. XXI. Two new species of Telaranea, sect. Tenuifoliae and records on Amazoopsis (Lepidoziaceae) from the Indian Ocean islands. Acta Botanica Hungarica 48 (1/2): 119–137. doi: 10.1556/ABot.48.2006.1-2.14 Pócs T (2006d) Contributions to the bryolora of Australia. II. On the Australasian “Tuyamaelloideae” (Lejeuneaceae), with the description of Austrolejeunea occidentalis. Journal of the Hattori Botanical Laboratory 99: 185–195. Pócs T (2007) A sixth species of Pictolejeunea (Jungermanniopsida) from Venezuelan Guyana. Acta Botanica Hungarica 49 (1/2): 109–119. doi: 10.1556/ABot.49.2007.1–2.12 Pócs T (2008a) Bryophytes from the Fiji Islands, IV. he genus Frullania Raddi (Jungermanniopsida), I., with description of F. vivipara Pócs, spec. nov. Fieldiana: Botany (n.ser.) 47: 147–158. doi: 10.3158/0015-0746-47.1.147 Pócs T (2008b) Bryophytes from the Fiji Islands, III. he genus Phaeolejeunea Mizut. (Lejeuneaceae), with detailed description of P. amicorum (Hürl.) Pócs, stat. nov. Fieldiana: Botany (n.ser.) 47: 139–145. doi: 10.3158/0015-0746-47.1.139 Pócs T (2010a) Myriocoleopsis in southeast Asia. Tropical Bryology 31: 123–125. Pócs T (2010b) On some new or less known Lejeunea (Lejeuneaceae, Jungermanniopsida) species in tropical Africa. East African bryophytes, XXVII. Beihefte zur Nova Hedwigia 138: 99–116. Pócs T (2010c) Bazzania orbanii (Lepidoziaceae), a new species from Madagascar. East African bryophytes, XXVIII. Acta Biologica Plantarum Agriensis 1: 15–22. Pócs T (2010d) What is Cladolejeunea Zwickel? New or little known epiphyllous liverworts. XV. Acta Biologica Plantarum Agriensis 1: 53–62. Pócs T (2011a) Type studies of some African Lejeuneaceae. Acta Botanica Hungarica 53 (1/2): 181–192. doi: 10.1556/ABot.53.2011.1-2.18 Pócs T (2011b) New or little known epiphyllous liverworts. XIV. he genus Colura (Lejeuneaceae) in São Tomé Island, with the description of Colura thomeensis sp. nov. Bryologist 114 (2): 362–366. doi: 10.1639/0007-2745-114.2.362 Pócs T (2011c) East African bryophytes XXIX. he Ceratolejeunea (Lejeuneaceae) species of the Indian Ocean islands. Polish Botanical Journal 56 (2): 131–153. Pócs T (2012a) Notes on the representatives of genus Kurzia G. Martens (Lepidoziaceae, Jungermanniopsida) in the Colombian Andes. Acta Biologica Plantarum Agriensis 2: 101–113. Pócs T (2012b) Bryophytes from the Fiji Islands, VI. he genus Cololejeunea Raddi (Jungermanniopsida), with the description of seven new species. Acta Botanica Hungarica 54 (1/2): 145–188. doi: 10.1556/ABot.54.2012.1–2.16 Pócs T (2013) he genus Colura (Lejeuneaceae) in New Guinea and in the neighboring areas. Chenia 11: 12–38. Pócs T (2015a) Validation of Plagiochila fracta nomen nudum (Jungermanniopsida). East African bryophytes XXXI. Phytotaxa 195 (2): 183–187. doi: 10.11646/phytotaxa.195.2.8 Pócs T (2015b) Contribution to the bryolora of Australia, V. Colura streimannii sp. nov. from Queensland. Polish Botanical Journal 60 (1): 7–11. doi: 10.1515/pbj-2015-0006 772 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Pócs T, Bernecker A (2009) Knowledge of Aphanolejeunea (Jungermanniopsida) after 25 years. Polish Botanical Journal 54 (1): 1–11. Pócs T, Bernecker A (2013) New or little known epiphyllous liverworts, XIX Cololejeunea yelitzae, a new neotropical species. Acta Botanica Hungarica 55 (3/4): 385–391. doi: 10.1556/ ABot.55.2013.3-4.13 Pócs T, Cairns A (2008) Contributions to the bryolora of Australia, III. he genus Jubula Dumort., with the description of J. hutchinsiae Hook. subsp. nov. australiae (Jubulaceae, Jungermanniopsida). Nova Hedwigia 86 (1/2): 231–236. doi: 10.1127/00295035/2008/0086-0231 Pócs T, Eggers J (1999) New or little known epiphyllous liverworts. VIII. Two new Papillolejeunea species from Papua New Guinea. Bryobrothera 5: 159–164. Pócs T, Eggers J (2007) Bryophytes from the Fiji Islands, II. An account of the genus Colura, with a description of C. vitiensis sp. nov. Polish Botanical Journal 52 (2): 81–92. Pócs T, Luke Q (2007) East African bryophytes, XXV. Bryological records from the Chyulu range, Kenya. Journal of East African Natural History 96 (1): 27–46. doi: 10.2982/0012-8317(2007)96[27:EABXBR]2.0.CO;2 Pócs T, Lye KA (1999) New records and additions to the hepatic lora of Uganda 2. Tropical Bryology 17: 23–33. Pócs T, Ninh T (2005) Contribution to the bryolora of Vietnam. VI. On the liverwort lora of Vu Quang Nature Reserve. Acta Botanica Hungarica 47 (1/2): 151–171. doi: 10.1556/ ABot.47.2005.1-2.14 Pócs T, Piippo S (1999) Bryophyte lora of the Huon Peninsula, Papua New Guinea. LXIV. Aphanolejeunea (Lejeuneaceae, Hepaticae). Acta Botanica Fennica 165: 85–102. Pócs T, Piippo S (2011) Bryophyte lora of the Huon Peninsula, Papua New Guinea. LXXIV. Cololejeunea (Lejeuneaceae, Hepaticae). Acta Bryolichenologica Asiatica 4: 59–137. Pócs T, Schäfer-Verwimp A (2006) East African bryophytes. XXIII. hree new species of Diplasiolejeunea (Lejeuneaceae, Jungermanniopsida) from Madagascar. Cryptogamie, Bryologie 27 (4): 439–452. Pócs T, Schäfer-Verwimp A (2012) Cololejeunea kuciana (Lejeuneaceae, Marchantiophyta), another new species from southern Ecuador. Polish Botanical Journal 57 (1): 51–53. Pócs T, Váňa J (2001) A new species of Amphicephalozia (Hepaticae) from Madagascar. Polish Botanical Journal 46 (2): 145–150. Pócs T, Mizutani M, Piippo S (1994) Bryophyte lora of the Huon Peninsula, Papua New Guinea. LXV. Preliminary contributions on Lejeuneaceae (Hepaticae). 1. Annales Botanici Fennici 31 (3): 179–190. Pócs T, Sass-Gyarmati A, Naikatini A, Tuiwawa M, Braggins JE, Pócs S, von Konrat MJ (2011) New liverwort (Marchantiophyta) records for the Fiji Islands. Telopea 13 (3): 455–494. Pócs T, Luong T-T, Ho B-C (2013) New or little known epiphyllous liverworts, XVIII. Records from the Bidoup-Núi Bà National Park, Vietnam, with the description of Drepanolejeunea bidoupensis, sp. nov. Cryptogamie, Bryologie 34 (3): 287–298. doi: 10.7872/cryb.v34. iss3.2013.287 Pócs T, Bernecker A, Tixier P (2014) Synopsis of the neotropical species of Cololejeunea (Spruce) Schifn. Acta Botanica Hungarica 56 (1/2): 185–226. doi: 10.1556/ABot.56.2014.1–2.14 World checklist of hornworts and liverworts 773 Pócs T, Zhu R-L, Reiner-Drehwald E, Söderström L, Hagborg A, von Konrat M (2015a) Notes on Early Land Plants Today. 71. New synonyms, new names and new combinations in Lejeuneaceae (Marchantiophyta). Phytotaxa 208 (1): 97–102. doi: 10.11646/phytotaxa.208.1.10 Pócs T, Müller F, Shevock JR (2015b) Additions to the liverwort and hornwort lora of São Tomé and Príncipe II, with Neurolejeunea, a genus new to Africa. Herzogia 28 (1): 50–69. Pócs T, Zhu R-L, Söderström L, Hagborg A, von Konrat M (2015c) Notes on Early Land Plants Today. 67. Notes on Lejeuneaceae subtribus Cololejeuneinae (Marchantiophyta). Phytotaxa 202 (1): 63–68. doi: 10.11646/phytotaxa.202.1.9 Polakowski H (1877) Bryophytas et cormophytas costaricensis anno 1875 lectas. Journal of Botany, British and Foreign 15: 225–231. Porsild MP (1902) Sur une nouvelle espèce de Riella (subg. nov.: Trabutiella) de l’Asie centrale. Botanisk Tidsskrift 24 (3): 323–327. Pôrto KC, Grolle R (1987) Drepanolejeunea bischleriana sp. nov. du Brésil. Cryptogamie: Bryologie, Lichénologie 8 (4): 301–304. Potemkin AD (1992) A new species of Prasanthus (Hepaticae, Gymnomitriaceae) from the Yamal Peninsula, West Siberian arctic. Annales Botanici Fennici 29 (4): 319–323. Potemkin AD (1993) he hepaticae of the Yamal Peninsula, West Siberian Arctic. Arctoa 2: 57–101. doi: 10.15298/arctoa.02.04 Potemkin AD (1998) On the origin, evolution and classiication of the genus Scapania (Dum.) Dum. (Hepaticae). Journal of the Hattori Botanical Laboratory 85: 33–61. Potemkin AD (1999) An analysis of the practical taxonomy of some critical northern species of Scapania (Scapaniaceae, Hepaticae). Bryologist 102 (1): 32–38. doi: 10.2307/3244456 Potemkin AD (2000a) An updated list of liverworts of the Severnaya Zemlya Archipelago (East Siberian high arctic) with description of a new species, Scapania matveyevae. Arctoa 9: 95–100. Potemkin AD (2000b) hree new species of Scapania (Hepaticae) collected by Dr. David G. Long in Nepal. Arctoa 9: 115–122. doi: 10.15298/arctoa.09.12 Potemkin AD (2000c) Bryophyte lora of Hunan Province, China. 2. Scapania koponenii sp. nova (Scapaniaceae, Hepaticae). Annales Botanici Fennici 37 (1): 41–44. Potemkin AD (2001) hree new species of Scapania (Hepaticae) from India and China. Annales Botanici Fennici 38 (2): 83–89. Potemkin AD (2002) Phylogenetic system and classiication of the family Scapaniaceae Mig. emend. Potemkin (Hepaticae). Annales Botanici Fennici 39 (4): 309–334. Potemkin AD (2005) Pečenočiki i antocerotovye Rossii: Taksonomičeskij sostav i perspektivy dal’nejših issledovanij [Liverworts and hornworts of Russia: taxonomic composition and prospects of further studies]. In: Afonina OM, Potemkin AD, Czernyadjeva IV (Eds) Actual Problems in Bryology: Proceedings of the International Meeting Devoted to the 90-th Anniversary of A. L. Abramova (Saint Petersburg, November 22-25, 2005). , St. Petersburg, 164–171. Potemkin AD, Ahti T (2012) On Riccia marginata and related species (Ricciaceae, Marchantiophyta) [O Riccia marginata i rodstvennyh vidah (Ricciaceae, Marchantiophyta)]. Novosti Sistematiki Nizših Rastenij 46: 298–305. Potemkin AD, Kazanovsky SG (1993) On the genus Mylia S. Gray (Hepaticae, Jungermanniaceae, Mylioideae). Arctoa 2: 1–11. doi: 10.15298/arctoa.02.01 774 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Potemkin AD, Sofronova EV (2009) Pečënočniki i antocerotov’e Rossii, Tom 1 [Liverworts and Hornworts of Russia, Vol. 1]. Russian Academy of Sciences, V. L. Komarov Botanical Institute, St. Petersburg, 368 pp. Potemkin AD, Piippo S, Koponen TJ (2004) Bryophyte lora of Hunan Province, China. 4. Diplophyllaceae and Scapaniaceae (Hepaticae). Annales Botanici Fennici 41 (6): 415–427. Prantl K (1890) Verzeichniss der in diesem Bande erwähnten Planzen (Kryptogamen). Hedwigia 29: vii–xxix. Prantl K (1892) Verzeichniss der in diesem Bande erwähnten Planzen (Kryptogamen). Hedwigia 31: viii–xxvii. Preussing M, Olsson S, Schäfer-Verwimp A, Wickett NJ, Wicke S, Quandt D, Nebel M (2010) New insights in the evolution of the liverwort family Aneuraceae (Metzgeriales, Marchantiophyta), with emphasis on the genus Lobatiriccardia. Taxon 59 (5): 1424–1440. Proskauer J (1951a) Studies on Anthocerotales. III. Bulletin of the Torrey Botanical Club 78 (4): 331–349. doi: 10.2307/2481996 Proskauer J (1951b) Notes on hepaticae. II. Bryologist 54 (4): 243–266. doi: 10.2307/3240002 Proskauer J (1955) Notes on hepaticae. III. Bryologist 58 (3): 192–200. doi: 10.2307/3239904 Proskauer J (1958) Nachtrag zur Familie Anthocerotaceae. In: Müller K (Ed.) Die Lebermoose Europas (Dr. L. Rabenhorst’s Kryptogamen-Flora von Deutschland, Österreich und der Schweiz, 3 Aul., 6 Band), 2 Abth., 9 Lieferung. Akademische Verlagsgesellschaft, Leipzig, 1303–1319. Proskauer J (1960) Studies on Anthocerotales VI. Phytomorphology 10: 1–19. Proskauer J (1964) Riccia tuberosa Taylor = Anogramma leptophylla (L.) Link, or the importance of being bryophytic. Journal of the Indian Botanical Society 42A: 185–188. Qiu YL, Li L, Wang B, Chen Z, Dombrovska O, Lee J, Kent L, Li R, Jobson RW, Hendry TA, Taylor DW, Testa CM, Ambros M (2007) Nonlowering land plant phylogeny inferred from nucleotide sequences of seven chloroplast, mitochondrial, and nuclear genes. International Journal of Plant Sciences 168 (5): 691–708. doi: 10.1086/513474 Qiu YL, von Konrat M, Engel JJ (2013) Rudolf Mathias Schuster. 1921-2012. Plant Science Bulletin 59 (4): 165–168. Qui Q, Wei Y-M, Cheng X-F, Zhu R-L (2014) Notes on Early Land Plants Today. 57. Cheilolejeunea boliviensis and Cheilolejeunea savesiana, two new synonyms in Lejeunea (Marchantiophyta, Lejeuneaceae). Phytotaxa 173 (1): 88–90. doi: 10.11646/phytotaxa.173.1.9 Rabenhorst GL (1857) Jungermannia sauteri De N. Hedwigia 1 (20): 121. Rabenhorst GL (1859) Hepaticae Europaeae. Die Lebermoose Europa’s unter Mitwirkung mehrerer namhafter Botaniker, Decades 9-10. Dresden, tab. 81–100. Rabenhorst GL (1860) Hepaticae Europaeae. Die Lebermoose Europa’s unter Mitwirkung mehrerer namhafter Botaniker, Decades 13-14. Dresden, tab. 121–140. Rabenhorst GL (1862) Hepaticae Europaeae. Die Lebermoose Europa’s unter Mitwirkung mehrerer namhafter Botaniker, Decades 21-22. Dresden, tab. 201–220. Raddi G (1818a) Jungermanniograia etrusca. Modena, 45 pp. Raddi G (1818b) Novarum vel rariorum ex cryptogamia stirpium in agro lorentino collectarum. Opuscoli Scientiici 2 (6): 349–361. World checklist of hornworts and liverworts 775 Raddi G (1822) Crittogame brasiliane. Società Tipograica, Modena, 33 pp. doi: 10.5962/bhl. title.65673 Radian SS (1903) Sur le Bucegia, nouveau genre d’hépatique à thalle. Bulletin de l’Herbier de l’Institut Botanique de Bucarest 3-4: 3–7. Ramaiya M, Johnson MG, Shaw J, Heinrichs J, Hentschel J, von Konrat M, Davison PG, Shaw AJ (2010) Morphologically cryptic biological species within the liverwort Frullania asagrayana. American Journal of Botany 97 (10): 1707–1718. doi: 10.3732/ajb.1000171 Rao DN (1982) Responses of bryophytes to air pollution. In: Smith AJE (Ed.) Bryophyte Ecology. Chapman & Hall, London, 445–471. doi: 10.1007/978-94-009-5891-3_12 Raoul EFL (1846) Choix de plantes de la Nouvelle-Zélande. Fortin, Masson et Cie, Paris, 56 pp. Rauh W, Buchloh G (1961) Riccia atromarginata Levier var. jovet-astii var. nov. Rauh & Buchloh. Revue Bryologique et Lichénologique 30 (1/2): 74–79. Rawat KK, Srivastava SC (2005) Plagiochila meghalayensis sp. nov. from Meghalaya, India. Geophytology 35 (1/2): 49–53. Reeb C, Bardat J (2014) Studies on African Riccardia types and related material. Cryptogamie, Bryologie 35 (1): 47–75. doi: 10.7872/cryb.v35.iss1.2014.47 Reichardt HW (1866) Diagnose der neuen Arten von Lebermoosen, welche die Novara-Expedition mitbrachte. Verhandlungen der Kaiserlich-Königlichen Zoologisch-Botanischen Gesellschaft in Wien 16: 957–960. Reimers H (1925) Hepaticae novae chilenses. Repertorium Specierum Novarum Regni Vegetabilis 21 (8/20): 264–265. doi: 10.1002/fedr.19250210817 Reimers H (1931) Beiträge zur Mooslora Chinas. I. Hedwigia 71 (1/2): 1–77. Reimers H (1933) Revision der Lebermoosgattung Micropterygium. Hedwigia 73 (3/4): 133–204. Reimers H (1936) Nachtrag zur Revision der Lebermoosgattung Micropterygium. Hedwigia 76 (3): 165–168. Reiner ME (1988) Contribución al conocimiento de las hepáticas del Noreste de la provincia de Buenos Aires (Argentina). Frullaniaceae (Jungermanniales). Boletín de la Sociedad Argentina de Botánica 25 (3/4): 301–325. Reiner-Drehwald ME (2000) On Potamolejeunea and Neopotamolejeunea gen. nov. (Lejeuneaceae, Hepaticae). Nova Hedwigia 71 (3/4): 447–464. Reiner-Drehwald ME (2005a) On Amphilejeunea and Cryptogynolejeunea, two small genera of Lejeuneaceae (Jungermanniopsida), and two common neotropical Lejeunea species. Nova Hedwigia 81 (3/4): 395–411. doi: 10.1127/0029-5035/2005/0081-0395 Reiner-Drehwald ME (2005b) Taxilejeunea pulverulenta (Lejeuneaceae, Jungermanniopsida), a poorly known species from the neotropics, is transferred to Lejeunea. Cryptogamie, Bryologie 26 (1): 59–65. Reiner-Drehwald ME (2006) Type studies on neotropical Lejeuneaceae (Jungermanniopsida). Cheilolejeunea and Lepidolejeunea. Nova Hedwigia 83 (3/4): 473–482. doi: 10.1127/00295035/2006/0083-0473 Reiner-Drehwald ME (2010) A taxonomic revision of Lejeunea deplanata (Lejeuneaceae, Marchantiophyta) from tropical America. Nova Hedwigia 91 (3/4): 519–532. doi: 10.1127/0029-5035/2010/0091-0519 776 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Reiner-Drehwald ME (2011) Studies on neotropical Lejeuneaceae (Jungermanniopsida). New synonyms and Ceratolejeunea temnantha (Spruce) comb. nov. Cryptogamie, Bryologie 32 (2): 95–100. doi: 10.7872/cryb.v32.iss1.2011.095 Reiner-Drehwald ME (2015) Lejeunea tunquiniensis (Lejeuneaceae, Marchantiophyta), a new species from a humid montane forest in the Yungas region, Bolivia. Nova Hedwigia 100 (3/4): 583–588. doi: 10.1127/nova_hedwigia/2015/0249 Reiner-Drehwald ME, Goda A (2000) Revision of the genus Crossotolejeunea (Lejeuneaceae, Hepaticae). Journal of the Hattori Botanical Laboratory 89: 1–54. Reiner-Drehwald ME, Gradstein SR (1997) New combinations and synonyms in Myriocoleopsis Schifn. (Lejeuneaceae). Journal of Bryology 19 (3): 638–640. doi: 10.1179/ jbr.1997.19.3.638 Reiner-Drehwald ME, Grolle R (2012) Review of the genus Rectolejeunea (Lejeuneaceae, Marchantiophyta). Nova Hedwigia 95 (3/4): 451–482. doi: 10.1127/0029-5035/2012/0063 Reiner-Drehwald ME, Mustelier Martínez K (2004) On Lejeunea multidentata, a new species from Cuba (Jungermanniopsida: Lejeuneaceae). Journal of Bryology 26 (2): 103–106. doi: 10.1179/037366804225021056 Reiner-Drehwald ME, Cavalcanti Pôrto K (2007) Lejeunea perpapillosa (Lejeuneaceae), a new species from north-eastern Brazil. Nova Hedwigia 85 (3/4): 541–546. doi: 10.1127/00295035/2007/0085-0541 Reiner-Drehwald ME, Schäfer-Verwimp A (1996) Drepanolejeunea grollei (Lejeuneaceae, Hepaticae) a new species from south-eastern Brazil. Candollea 51 (2): 475–482. Reiner-Drehwald ME, Schäfer-Verwimp A (2008) On Inflatolejeunea, Lejeunea species with eplicate perianths and Lejeunea talamancensis sp. nov. from Costa Rica (Lejeuneaceae). Nova Hedwigia 87 (3/4): 387–420. doi: 10.1127/0029-5035/2008/00870387 Reiner-Drehwald ME, Mustelier Martínez K, Gradstein SR (2007) A new species of Omphalanthus (Lejeuneaceae) from Cuba. Journal of Bryology 29 (2): 95–97. doi: 10.1179/174328207X171881 Reiner-Drehwald ME, Salazar Allen N, Chung CC (2013) New combinations and synonyms in neotropical Lejeuneaceae (Marchantiophyta), with description of Lejeunea tamasii, a new species from Barro Colorado Island, Panama. Polish Botanical Journal 58 (2): 419–426. doi: 10.2478/pbj-2013-0041 Reinwardt CGC, Blume CL, Nees von Esenbeck CG (1824a) Hepaticae iavanicae. Nova Acta Physico-Medica Academiae Caesareae Leopoldino-Carolinae Naturae Curiosorum 12 (1): 181–238. Reinwardt CGC, Blume CL, Nees von Esenbeck CG (1824b) Hepaticarum iavanicarum, supplementum. Nova Acta Physico-Medica Academiae Caesareae Leopoldino-Carolinae Naturae Curiosorum 12 (1): 409–417. Renauld F, Cardot J (1891) Contribution à la lore des muscinées des îles austro-africaines et de l’Océan Indien. Revue Bryologique 18 (4): 55–60. Renauld F, Cardot J (1893) Musci exotici novi vel minus cogniti. V. Bulletin de la Société Royale de Botanique de Belgique, Mémoires 32: 101–121. World checklist of hornworts and liverworts 777 Renner MAM (2006) Plants seen and unseen: Radula pseudoscripta sp. nov. and Radula uvifera (Radulaceae: Jungermanniopsida). New Zealand Journal of Botany 44 (3): 337–348. doi: 10.1080/0028825X.2006.9513026 Renner MAM (2010) Another new species of Austrolejeunea (Lejeuneaceae) from New Zealand’s subalpine environs. Bryologist 113 (4): 781–787. doi: 10.1639/0007-2745-113.4.781 Renner MAM (2012) Further insight into lobule teeth homology in Lejeuneaceae subf. Lejeuneoideae from Cheilolejeunea oscilla, a new species from Australia. Bryologist 115 (4): 536–556. doi: 10.1639/0007-2745-115.4.536 Renner MAM (2013) A new subspecies of Acrolejeunea arcuata, and notes on typiication, synonymy, and distribution of other Australasian Lejeuneaceae. Phytotaxa 83 (1): 39–53. doi: 10.11646/phytotaxa.83.1.2 Renner MAM (2014) Radula subg. Radula in Australasia and the Paciic (Jungermanniopsida). Telopea 17: 107–167. doi: 10.7751/telopea20147553 Renner MAM, Brown EA (2008) Mnioloma (Calypogeiaceae: Jungermanniopsida) in Australasia: how many species are there? Fieldiana: Botany (n.ser.) 47: 159–174. doi: 10.3158/00150746-47.1.159 Renner MAM, de Lange PJ (2009) Radula multilora Gottsche ex Schifn. – a new record for the Chatham Islands, New Zealand. Australasian Bryological Newsletter 57: 12–13. Renner MAM, de Lange PJ (2011) Additions to the Lejeuneaceae lora of New Zealand: new species from the Kermadec Islands and range extensions of New Zealand species into the South Paciic. New Zealand Journal of Botany 49 (3): 421–433. doi: 10.1080/0028825X.2011.580765 Renner MAM, Brown EA, Glenny DS (2006) Two new Zoopsis species and their relationships to other zoopsids (Jungermanniopsida: Lepidoziaceae). Journal of Bryology 28 (4): 331–344. doi: 10.1179/174328206X136278 Renner MAM, Brown EA, Wardle GM (2009) Evidence for species recognition on the basis of a single specimen: Nephelolejeunea carcharias sp. nov. (Lejeuneaceae: Jungermanniopsida). Systematic Botany 34 (4): 615–624. doi: 10.1600/036364409790139754 Renner MAM, Devos N, Shaw AJ (2010a) Radula splendida sp. nov. (Radulaceae: Marchantiophyta), a polymorphic species from New Zealand. Nova Hedwigia 90 (1/2): 105–122. doi: 10.1127/0029-5035/2010/0090-0105 Renner MAM, Brown EA, Wardle GM (2010b) he Lejeunea tumida species group (Lejeuneaceae: Jungermanniopsida) in New Zealand. Australian Systematic Botany 23 (6): 443– 462. doi: 10.1071/SB10037 Renner MAM, Devos N, Patiño J, Brown EA, Orme A, Elgey M, Wilson TC, Gray LJ, von Konrat MJ (2013a) Integrative taxonomy resolves the cryptic and pseudo-cryptic Radula buccinifera complex (Porellales, Jungermanniopsida), including two reinstated and ive new species. PhytoKeys 27: 1–113. doi: 10.3897/phytokeys.27.5523 Renner MAM, Söderström L, Hagborg A, von Konrat MJ (2013b) Notes on Early Land Plants Today. 29. A new combination in Radula (Radulaceae, Marchantiophyta). Phytotaxa 81 (1): 12–14. doi: 10.11646/phytotaxa.81.1.5 Renner MAM, Devos N, Brown EA, von Konrat MJ (2013c) New records, replacements, reinstatements and four new species in the Radula parvitexta and R. ventricosa species groups (Jungermanniopsida) in Australia: cases of mistaken identity. Australian Systematic Botany 26 (4): 298–345. doi: 10.1071/SB13027 778 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Renner MAM, Pócs T, Söderström L, Hagborg A, von Konrat MJ (2013d) Notes on Early Land Plants Today. 27. Transfer of some taxa from Stenolejeunea (Lejeuneaceae, Marchantiophyta). Phytotaxa 81 (1): 8–9. doi: 10.11646/phytotaxa.81.1.3 Renner MAM, Söderström L, Hagborg A, von Konrat M (2014) Notes on Early Land Plants Today. 50. Radula sainsburiana is a synonym of R. helix (Radulaceae, Marchantiophyta). Phytotaxa 162 (4): 239. doi: 10.11646/phytotaxa.162.4.9 Renner MAM, Engel JJ, Patzak SDF, Heinrichs J (2015) A new species of Brevianthus (Brevianthaceae, Marchantiophyta) from New Caledonia with unusual underleaf production. PhytoKeys 50: 43–60. doi: 10.3897/phytokeys.50.4998 Reyes DM (1982) El género Diplasiolejeunea en Cuba. Acta Botanica Academiae Scientiarum Hungaricae 28 (1/2): 145–180. Robinson H (1964) New taxa and new records of bryophytes from Mexico and Central America. Bryologist 67 (4): 446–458. doi: 10.2307/3240770 Robinson H (1965) A new species of Plagiochila from Venezuela. Bryologist 68 (1): 93–94. doi: 10.2307/3240990 Robinson H (1967) Preliminary studies on the bryophytes of Colombia. Bryologist 70 (1): 1–61. doi: 10.2307/3241138 Robinson H (1970) Notes on the genus Nowellia. Bryologist 73 (1): 150–152. doi: 10.2307/3241596 Robinson H (1976a) A new species of Taxilejeunea from Venezuela. Phytologia 34 (1): 67–68. Robinson H (1976b) Hepaticae. In: Steyermark JA, Brewer-Carías C (Eds) La vegetación de la cima del Macizo de Jaua. Boletín de la Sociedad Venezolana de Ciencias Naturales 32 (132/133): 179–405. Robinson H (1988) Plagiochila rudischusteri, a new species from the Paria Peninsula of Venezuela. Beihefte zur Nova Hedwigia 90: 199–202. Rodway L (1917a) Additions to the bryophyte lora. Papers and Proceedings of the Royal Society of Tasmania 1916: 44–47. Rodway L (1917b) Tasmanian Bryophyta. II. Hepatics. Royal Society of Tasmania, Hobart, 95 pp. Röll J (1891) Vorläuige Mittheilung über die von mir im Jahre 1888 in Nord-Amerika gesammelten neuen Arten der Lebermoose. Botanisches Centralblatt 45: 203–204. Roth A (1788) Tentamen lorae germanicae. tomus 1. Bibliophilio I. G. Mülleriano, Lipsiae, 568 pp. Roth A (1800) Tentamen lorae germanicae, tomus III, pars I. Bibliopolio. I.G. Mülleriano, Lipsiae [Leipzig], 581 pp. doi: 10.5962/bhl.title.6694 Roumeguère C (1888) Hépatiques de l’Aude. Mémoires de la Société des arts et des sciences de Carcassonne 5: 105–206. Rubasinghe SCK, Milne R, Forrest LL, Long DG (2011a) Realignment of the genera of Cleveaceae (Marchantiopsida, Marchantiidae). Bryologist 114 (1): 116–127. doi: 10.1639/0007-2745-114.1.116 Rubasinghe SCK, Long DG, Milne R (2011b) A new combination and three new synonyms in the genus Clevea Lindb. (Marchantiopsida, Cleveaceae). Journal of Bryology 33 (2): 166–168. doi: 10.1179/1743282011Y.0000000005 World checklist of hornworts and liverworts 779 Ryan E (1889) Scapania kaurinii n. sp. Botaniska Notiser 42: 210–211. Salazar Allen N (2001) Cyathodium bischlerianum, sp. nov. (Marchantiales) a new species from the Neotropics. Bryologist 104 (1): 141–145. doi: 10.1639/0007-2745(2001)104[0141:CBSNMA]2.0.CO;2 Salmon ES (1901) Isotachis Stephanii sp. nov. Revue Bryologique 28 (4): 75–76. Sande Lacoste CM (1854) Novae species hepaticarum ex insula Java detexit Dr. F. Junghuhn. Nederlandsch Kruidkundig Archief. Verslagen en Mededelingen der Nederlandsche Botanische Vereeniging 3: 415–424. Sande Lacoste CM (1855) Speciebus novis hepaticarum javanicarum herbarii Junghuhniani. Nederlandsch Kruidkundig Archief. Verslagen en Mededelingen der Nederlandsche Botanische Vereeniging 3: 521–522. Sande Lacoste CM (1856a) Hepaticae. In: De Vriese WH (Ed.) Plantae indiae batavae orientalis, fasc. 1. E. J. Brill, Lugdunum Batavorum, 18–26. Sande Lacoste CM (1856b) Synopsis hepaticarum javanicarum. C. G. van der Post, Amsterdam, 1–112. Sande Lacoste CM (1856c) Hepaticae. In: Dozy F (Ed.) Plagiochila sandei Dz. Jac. Hazenberg Corn. il., Leiden, 5–12. Sande Lacoste CM (1864) Hepaticae. Jungermanniae archipelagi indici. Annales Musei Botanici Lugduno-Batavi 1: 287–314. Sande Lacoste CM (1867) Musci et hepaticae. In: Miquel FAG (Ed.) Prolusio lorae japonicae. Vander Post, Amsterdam, 209–0. Sass-Gyarmati A (2001) Lopholejeunea leioptera Gyarmati (Lejeuneaceae, subfam. Ptychanthoideae), une nouvelle espèce récoltée dans la Réserve Spéciale de Manongarivo (Nord-Ouest de Madagascar). Candollea 56 (1): 79–83. Sass-Gyarmati A (2002) Ptychanthoideae (subfam. of Lejeuneaceae, hepaticae) from Vanuatu, with the description of Caudalejeunea streimannii Gyarmati sp. n. Tropical Bryology 22: 125–134. Sass-Gyarmati A (2005) Lopholejeunea pocsii Gyarmati (Lejeuneaceae, subfam. Ptychanthoideae), a new species of subgenus Pholianthus B. hiers & Gradst. from the Fiji-Islands. Cryptogamie, Bryologie 26 (4): 403–410. Sass-Gyarmati A (2008) Bryophytes from the Fiji Islands, V. Lopholejeunea vojtkoana Gyarmati sp. nov. from Fiji Islands (Lejeuneaceae, subfam. Ptychanthoideae). Nova Hedwigia 87 (3/4): 479–486. doi: 10.1127/0029-5035/2008/0087-0479 Sass-Gyarmati A, Pócs T (2014) Acanthocoleus elgonensis Gyarmati et Pócs, sp. nov. from Mount Elgon (Uganda). Cryptogamie, Bryologie 35 (2): 119–125. doi: 10.7872/cryb. v35.iss2.2014.119 Sauter AE (1860) Berichtigung. Flora 43 (22): 351. Schäfer-Verwimp A (2001a) On Lejeunea patriciae, nom. nov. for Lejeunea pilifera Tixier. Candollea 56 (1): 63–67. Schäfer-Verwimp A (2001b) Diplasiolejeunea pluridentata (Lejeuneaceae, Marchantiopsida), eine neue Art aus Costa Rica. Haussknechtia 8: 71–78. Schäfer-Verwimp A (2004) he genus Diplasiolejeunea (Lejeuneaceae, Marchantiopsida) in the tropical Andes, with description of two new species. Cryptogamie, Bryologie 25 (1): 3–17. 780 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Schäfer-Verwimp A (2005) Diplasiolejeunea riclefgrollei (Lejeuneaceae, Hepaticae), a remarkable new species from Cuba. Cryptogamie, Bryologie 26 (1): 37–40. Schäfer-Verwimp A (2006) A new species of Diplasiolejeunea (Lejeuneaceae, Jungermanniopsida) from Sumatra, and a key for the genus in Asia. Herzogia 19: 239–244. Schäfer-Verwimp A (2010) A checklist of the liverworts and hornworts of Dominica, West Indies. Cryptogamie, Bryologie 31 (4): 313–415. Schäfer-Verwimp A (2012) Cololejeunea tamasii (Lejeuneaceae, Marchantiophyta), a new species from Panama. Phytotaxa 60: 9–12. Schäfer-Verwimp A, Pócs T (2009) Contributions to the hepatic lora of the Dominican Republic, West Indies. Acta Botanica Hungarica 51 (3/4): 367–425. doi: 10.1556/ ABot.51.2009.3–4.13 Schäfer-Verwimp A, Reiner-Drehwald ME (2009) Some additions to the bryophyte lora of Guadeloupe, West Indies, and new synonyms in the genera Diplasiolejeunea and Lejeunea (Lejeuneaceae). Cryptogamie, Bryologie 30 (3): 357–375. Schäfer-Verwimp A, Peralta DF, Costa Siqueira SM (2012) Frullania curvilobula (Frullaniaceae, Marchantiophyta), a new species from Brazil. Phytotaxa 57 (4): 27–30. Schäfer-Verwimp A, Nebel M, Heinrichs J (2013) Diplasiolejeunea mayaykuensis (Lejeuneaceae, Marchantiophyta), a new liverwort species from southern Ecuador. Polish Botanical Journal 58 (1): 143–148. doi: 10.2478/pbj-2013-0015 Schäfer-Verwimp A, Feldberg K, Dong S, van Melick H, Peralta DF, Schmidt AR, Schneider H, Heinrichs J (2014) Towards a monophyletic classiication of Lejeuneaceae III: the systematic position of Leiolejeunea. Phytotaxa 170 (3): 187–198. doi: 10.11646/phytotaxa.170.3.4 Schifner V (1890) Lebermoose (Hepaticae) mit Zugrundelegung der von Dr. A.C.M. Gottsche ausgeführten Vorarbeiten. In: Engler A (Ed.) Die Forschungsreise S. M. S. “Gazelle”, IV. heil. Botanik 4. Ernst Siegfried Mittler und Sohn, Berlin, 1–45. doi: 10.5962/bhl.title.984 Schifner V (1893a) Ueber exotische Hepaticae, hauptsächlich aus Java, Amboina und Brasilien, nebst einigen morphologischen und kritischen Bemerkungen über Marchantia. Nova Acta Academiae Caesareae Leopoldino-Carolinae Germanicae Naturae Curiosorum 60 (2): 219–316. Schifner V (1893b) Hepaticae. In: Engler A, Prantl K (Eds) Die Natürlichen Planzenfamilien, Teil. I, Abt. 3. Engelmann, Leipzig, 1–144. Schifner V (1894) Revision der Gattungen Bryopteris, hysananthus, Ptychanthus und Phragmicoma im Herbarium des Berliner Museums. Hedwigia 33 (4): 170–189. Schifner V (1896a) Neue Beiträge zur Bryologie Nordböhmens und des Riesengebirges. Sitzungsberichte des deutschen naturwissenschaftlich-medicinischen Vereins für Böhmen “Lotos” in Prag 44 (8): 267–289. Schifner V (1896b) Wiesnerella, eine neue Gattung der Marchantiaceen. Österreichische Botanische Zeitschrift 46 (3): 82–88. doi: 10.1007/BF01677836 Schifner V (1897) Revision der Gattungen Omphalanthus und Lejeunea im Herbarium des Berliner Museums. Botanische Jahrbücher für Systematik, Planzengeschichte und Planzengeographie 23 (5): 578–600. World checklist of hornworts and liverworts 781 Schifner V (1898a) Expositio plantarum in itinere suo indico annis 1893/94 suscepto collectarum. Denkschriften der Kaiserlichen Akademie der Wissenschaften. Mathematisch-Naturwissenschaftliche Klasse 67: 153–203. Schifner V (1898b) Conspectus hepaticarum archipelagi indici. Staatsdruckerei, Batavia, 382 pp. Schifner V (1898c) Eine neue Planzengattung der indo-malayischen Flora. Annales du Jardin botanique de Buitenzorg, suppl. 2: 39–46. Schifner V (1899a) Beiträge zur Lebermooslora von Bhutan (Ost-Indien), II. Österreichische Botanische Zeitschrift 49 (6): 203–207. doi: 10.1007/BF01790595 Schifner V (1899b) Beiträge zur Lebermooslora von Bhutan (Ost-Indien). Österreichische Botanische Zeitschrift 49 (4): 127–132. doi: 10.1007/BF01794513 Schifner V (1899c) Ueber einige Hepaticae aus Japan. Österreichische Botanische Zeitschrift 49 (11): 385–395. doi: 10.1007/BF01672323 Schifner V (1900a) Die Hepaticae der Flora von Buitenzorg. I. Band. E. J. Brill, Leiden, 220 pp. Schifner V (1900b) Hepaticae massartianae javanicae. Hedwigia 39 (4): 191–208. Schifner V (1900c) Exposito plantarum in itinere suo indico annis 1893/94 suspecto collectarum, II. Denkschriften der Kaiserlichen Akademie der Wissenschaften. MathematischNaturwissenschaftliche Klasse 70: 155–218. Schifner V (1900d) Nachweis einiger für die böhmische Flora neuer Bryophyten nebst Bemerkungen über einzelne bereits daselbst nachgewiesene Formen. Sitzungsberichte des deutschen naturwissenschaftlich-medicinischen Vereins für Böhmen “Lotos” in Prag 48: 320–354. Schifner V (1901) Untersuchungen über Mörckia Flotowiana und über das Verhältnis der Gattungen Mörckia Gott. und Calycularia Mitt. zu einander. Österreichische Botanische Zeitschrift 51 (2): 41–51. doi: 10.1007/BF01672735 Schifner V (1902) Neue Materialen zur Kenntnis der Bryophyten der atlantischen Inseln. Hedwigia 41 (5): 269–294. Schifner V (1903a) Studien über kritische Arten der Gattungen Gymnomitrium und Marsupella (Schluss). Österreichische Botanische Zeitschrift 53 (7): 280–284. doi: 10.1007/ BF01791102 Schifner V (1903b) Studien über kritische Arten der Gattungen Gymnomitrium und Marsupella (Fortsetzung). Österreichische Botanische Zeitschrift 53 (6): 246–252. doi: 10.1007/ BF01679557 Schifner V (1906a) Die Lebermoose der deutschen Südpolar-Expedition 1901–1903. In: Drygalski ED (Ed.) Deutsche Südpolar-Expedition 1901-1903, Band. 8 (Botanik). Georg Reimers, Berlin, 59–80. Schifner V (1906b) Bryologische Fragmente XXVII-XXXIII. Österreichische Botanische Zeitschrift 56 (1): 20–27. doi: 10.1007/BF01791661 Schifner V (1906c) Die bisher bekannt gewordenen Lebermoose Dalmatiens, nebst Beschreibung und Abbildung von zwei neuen Arten. Verhandlungen der Kaiserlich-Königlichen Zoologisch-Botanischen Gesellschaft in Wien 56 (3): 263–280. Schifner V (1908a) Kritische Bemerkungen über die europäischen Lebermoose mit Bezug auf die Exemplare des Exsiccatenwerkes, Hepaticae europaeae exsiccatae 5 (201-250). Bericht des Naturwissenschaftlich-medizinischen Vereins Innsbruck 31 [Beilage]: 1–69. Schifner V (1908b) Bryologische Fragmente. XLIX-LII. Österreichische Botanische Zeitschrift 58 (10): 377–382. doi: 10.1007/BF01792481 782 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Schifner V (1909a) Bemerkungen über zwei kritische Hepaticae der europäischen Flora. Hedwigia 48 (3): 184–190. Schifner V (1909b) Hepaticae. In: Handel-Mazzetti HM (Ed.) Ergebnisse einer botanische Reise in des Pontische Randgebirge in Sandschak Trapezunt. Annalen des Naturhistorischen Museums in Wien 23: 6–212. Schifner V (1910a) Über die Gattungen Chiloscyphus und Heteroscyphus n. gen. Österreichische Botanische Zeitschrift 60 (5): 169–173. doi: 10.1007/BF01631854 Schifner V (1910b) Eine neue europäische Art der Gattung Anastrophyllum. Hedwigia 49 (4): 396–399. Schifner V (1910c) Studien über die Rhizoïden der Marchantiales. Annales du Jardin botanique de Buitenzorg, suppl. 3: 473–492. Schifner V (1910d) Kritische Bemerkungen über die europäischen Lebermoose mit Bezug auf die Exemplare des Exsiccatenwerkes: Hepaticae europaeae exsiccatae VIII (363-376). Lotos. Zeitschrift für Naturwissenschaften 58: 314–333. Schifner V (1911) Über einige neotropische Metzgeria-Arten, Schluß. Österreichische Botanische Zeitschrift 61 (7/8): 261–264. doi: 10.1007/BF01634594 Schifner V (1912a) Über eine kritische Form von Riccia sorocarpa und Riccia pseudopapillosa. Hedwigia 53 (1/2): 36–40. Schifner V (1912b) Bryologische Fragmente LXVI-LXXI. Österreichische Botanische Zeitschrift 62 (1): 8–15. doi: 10.1007/BF01644055 Schifner V (1912c) Kritik der Europäischen Formen der Gattung Chiloscyphus auf phylogenetischer Grundlage. Beihefte zum Botanischen Centralblatt 29: 74–116. Schifner V (1913) Bryophyta aus Mesopotamien und Kurdistan, Syrien, Rhodos, Mytilini und Prinkipo. Annalen des K. K. Naturhistorischen Hofmuseums 27: 472–504. Schifner V (1914a) Cephalozia-Studien. Hedwigia 54 (6): 311–327. Schifner V (1914b) Lebermoose aus Ungarn und Kroatien. IV. Beitrag. Magyar Botanikai Lapok 13: 302–309. Schifner V (1918) Hepaticae baumgartnerianae dalmaticae. III. Serie. Österreichische Botanische Zeitschrift 67 (4/5): 147–156. doi: 10.1007/BF01648499 Schifner V (1919) Kritische Bemerkungen über die europäischen Lebermoose mit Bezug auf die Exemplare des Exsiccatenwerkes: Hepaticae europaeae exsiccatae XIV. Privately published, Prague, 1–20. Schifner V (1929) Über epiphylle Lebermoose aus Japan nebst einigen Beobachtungen über Rhizoiden, Elateren und Brutkörper. Annales Bryologici 2: 87–106. Schifner V (1937) Nachträge und Berichtigungen zu Teil V, soweit solche nicht an leicht auindbaren Stellen von anderen veröfentlicht sind. In: Keissler K, Lohwag H (Ed.) Symbolae Sinicae, part 2. Fungi. Springer Verlag, Berlin, 81–0. Schifner V (1942) Monographie der Gattung Exormotheca Mitt. Hedwigia 81 (1/2): 40–74. Schifner V (1944) Myriocoleopsis, eine neue Gattung der Jubuleae. Hedwigia 81 (5/6): 234–237. Schifner V (1955) Die Lebermoose der Deutschen Limnologischen Sunda-Expedition. Archiv für Hydrobiologie, suppl. 21 (3/4): 382–407. Schifner V, Arnell SW (1964) Ergebnisse der Botanischen Expedition der kaiserlichen Akademie der Wissenschaften nach Südbrasilien 1901. II. hallophyta und Bryophyta. Hepaticae World checklist of hornworts and liverworts 783 (Lebermoose). Österreichische Akademie der Wissenschaften. Mathematisch-Naturwissenschaftliche Klasse, Denkschriften 111: 1–156. Schifner V, Pandé SK, Srivastava KP (1959) he genus Riccardia Gray in India. II. Riccardia palmatiformis Schfn. and R. decolyana Schfn. Journal of the Indian Botanical Society 38 (4): 538–542. Schill D, Long DG (2002) Anastrophyllum lignicola (Lophoziaceae), a new species from the SinoHimalaya, and A. hellerianum new to China. Annales Botanici Fennici 39 (2): 129–132. Schill DB, Long DG, Moeller M, Squirrell J (2004) Phylogenetic relationships between Lophoziaceae and Scapaniaceae based on chloroplast sequences. Monographs in Systematic Botany from the Missouri Botanical Garden 98: 141–149. Schill DB, Long DG, Köckinger H (2008) Taxonomy of Mannia controversa (Marchantiidae, Aytoniaceae) including a new subspecies from East Asia. Edinburgh Journal of Botany 65 (1): 35–47. doi: 10.1017/S0960428608004939 Schill DB, Long DG, Forrest LL (2010) A molecular phylogenetic study of Mannia (Marchantiophyta, Aytoniaceae) using chloroplast and nuclear markers. Bryologist 113 (1): 164–179. doi: 10.1639/0007-2745-113.1.164 Schmidel CC (1760) Dissertatio inauguralis botanica de Jungermanniae charactere. Tezschnerianis, Erlangae [Erlangen], 29 pp. Schmitt U, Winkler S (1968) Systematische Untersuchungen über die foliose Lebermoosgattung Stephaniella Jack. Österreichische Botanische Zeitschrift 115 (2): 120–133. doi: 10.1007/BF01373533 Schrader HA (1797) Systematische Sammlung cryptogamischer Gewächse, Zweyte Lieferung. J. C. Dietrich, Göttingen, 16 pp. Schrader HA (1802) Plantae cryptogamicae nouae, rariores aut minus cognitae. Journal für die Botanik 5: 66–80. Schrank FP (1792) Primitiae lorae salisburgensis dissertatione praevia de discrimine plantarum ab animalibus. Varrentrapp et Wenner, Frankfurt a. M., 240 pp. Schreber JCD (1771) Spicilegium lorae lipsicae. Bibliopolio Dykiano, Lipsiae [Leipzig], 148 pp. Schuette SW, Stotler RE (2005) A conspectus of the liverwort genus Jensenia in Latin America. Journal of the Hattori Botanical Laboratory 97: 299–308. Schultze-Motel W, Menzel M (1987) Die Lebermooslora im BRYOTROP-Transekte von Peru. Beihefte zur Nova Hedwigia 88: 61–104. Schumacker R, Váňa J (2005) Identiication keys to the liverworts and hornworts of Europe and Macaronesia, ed. 2. Sorus, Poznań, 209 pp. Schuster RM (1950) Notes on nearctic hepaticae. II. he hepaticae in east coast of Hudson Bay. Bulletin of the National Museum of Canada 122: 1–62. Schuster RM (1952) Notes on nearctic hepaticae. V. he status of Lophozia gracillima Buch and its relation to Lophozia longidens, Lophozia porphyroleuca and Sphenolobus ascendens. Bryologist 55 (3): 173–185. doi: 10.2307/3239846 Schuster RM (1953) Boreal hepaticae. A manual of the liverworts of Minnesota and adjacent regions. American Midland Naturalist 49 (2): 257–684. Schuster RM (1956a) North American Lejeuneaceae. V. Schizostipae: Ceratolejeunea. Journal of the Elisha Mitchell Scientiic Society 72 (2): 292–316. 784 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Schuster RM (1956b) Notes on American Lejeuneaceae. IV. Paradoxae: Cololejeunea (concl.), Diplasiolejeunea. Journal of the Elisha Mitchell Scientiic Society 72 (1): 87–125. Schuster RM (1956c) Aphanolejeunea cornutissima nom. nov. Bryologist 59 (3): 217–218. doi: 10.2307/3239952 Schuster RM (1957a) North American Lejeuneaceae. VI. Lejeunea. Introduction and keys; subgenus Lejeunea. Journal of the Elisha Mitchell Scientiic Society 73 (1): 122–197. Schuster RM (1957b) Notes on nearctic hepaticae. XII. Marsupella paroica n. sp. Bryologist 60 (2): 145–151. doi: 10.2307/3240015 Schuster RM (1957c) Notes on Nearctic hepaticae, IX. A study of Plagiochila yokogurensis Steph. Journal of the Hattori Botanical Laboratory 18: 14–26. Schuster RM (1957d) North American lejeuneaceae. VI. Lejeunea: subgenus Lejeunea (II, concluded). Journal of the Elisha Mitchell Scientiic Society 73 (2): 388–443. Schuster RM (1958a) Notes on nearctic hepaticae. VI. Phytogeographical relationships of critical species in Minnesota and adjacent areas of the Great Lakes. Rhodora 60 (717): 243–256. Schuster RM (1958b) Notes on nearctic hepaticae. XIII: Tritomaria (Lophoziaceae) in Arctic. Canadian Journal of Botany 36 (2): 269–288. doi: 10.1139/b58-023 Schuster RM (1959a) A monograph of the nearctic Plagiochilaceae. Part. I. Introduction and sectio I. Asplenioides. American Midland Naturalist 62 (1): 1–166. doi: 10.2307/2422546 Schuster RM (1959b) A monograph of the nearctic Plagiochilaceae. Part. II. Sectio Zonatae through Sectio Parallelae. American Midland Naturalist 62 (2): 257–395. doi: 10.2307/2422533 Schuster RM (1959c) Studies on hepaticae. I Temnoma. Bryologist 62 (4): 233–242. doi: 10.2307/3240148 Schuster RM (1960a) Notes on nearctic hepaticae. XIX. he relationships of Blepharostoma, Temnoma and Lepicolea, with description of Lophochaete and Chandonanthus subg. Tetralophozia, subg. n. Journal of the Hattori Botanical Laboratory 23: 192–210. Schuster RM (1960b) Studies on hepaticae. II. he new family Chaetophylliopsidaceae. Journal of the Hattori Botanical Laboratory 23: 68–76. Schuster RM (1960c) A monograph of the nearctic Plagiochilaceae. Part. III. Sectio Contiguae to conclusion. American Midland Naturalist 63 (1): 1–130. doi: 10.2307/2422932 Schuster RM (1961a) Studies in Lophoziaceae. 1. he genera Anastrophyllum and Sphenolobus and their segregates. Revue Bryologique et Lichénologique 30 (1/2): 55–73. Schuster RM (1961b) Notes on nearctic hepaticae. XVIII. New Lophoziaceae from the arctic archipelago of Canada. Canadian Journal of Botany 39 (4): 965–992. doi: 10.1139/b61-081 Schuster RM (1962a) North American Lejeuneaceae. VIII. Lejeunea subgenera Microlejeunea and Chaetolejeunea. Journal of the Hattori Botanical Laboratory 25: 1–80. Schuster RM (1962b) Notes on American Lejeuneaceae. VII. Lejeunea (Lejeunea) blomquistii sp. nov. Journal of the Elisha Mitchell Scientiic Society 78 (1): 64–68. Schuster RM (1963a) An annotated synopsis of the genera and subgenera of Lejeuneaceae. I. Introduction; annotated keys to subfamilies and genera. Beihefte zur Nova Hedwigia 9: 1–203. Schuster RM (1963b) Studies on antipodal hepaticae. I. Annotated keys to the genera of antipodal Hepaticae with special reference to New Zealand and Tasmania. Journal of the Hattori Botanical Laboratory 26: 185–309. World checklist of hornworts and liverworts 785 Schuster RM (1963c) Studies on hepaticae XI-XIII. On Temnoma, Vetaforma and Lophochaete (Blepharostomaceae; Hepaticae). Nova Hedwigia 5: 27–46. Schuster RM (1964a) Studies on antipodal hepaticae. IV. Metzgeriales. Journal of the Hattori Botanical Laboratory 27: 183–216. Schuster RM (1964b) Studies on hepaticae. XXII-XXV. Pleurocladopsis Schust., gen. n., Eoisotachis Schust., gen. n., Grollea Schust., gen. n., with critical notes on Anthelia Dumort. Nova Hedwigia 8 (3/4): 275–296. Schuster RM (1965a) North American Lejeuneaceae. IX. Taxilejeunea. Journal of the Elisha Mitchell Scientiic Society 81 (1): 32–50. Schuster RM (1965b) Studies on hepaticae. XXVI. he Bonneria-Paracromastigum-Pseudocephalozia-Hyalolepidozia-Zoopsis-Pteropsiella complex and its allies – a phylogenetic study, part 1. Nova Hedwigia 10 (1/2): 19–61. Schuster RM (1965c) Studies on antipodal hepaticae. II. Archeophylla Schuster and Archeochaete Schuster, new genera of Blepharostomataceae. Transactions of the British Bryological Society 4 (5): 801–817. doi: 10.1179/006813865804811975 Schuster RM (1966a) Studies on hepaticae, VII-X. On Adelanthus Mitten and Calyptrocolea Schuster, gen. n. Revue Bryologique et Lichénologique 34 (3/4): 676–703. Schuster RM (1966b) Studies in Lophoziaceae. 1. he genera Anastrophyllum and Sphenolobus and their segregates. 2. Cephalolobus gen. n., Acrolophozia gen. n. and Protomarsupella gen. n. Revue Bryologique et Lichénologique 34 (1/2): 241–287. Schuster RM (1966c) A memoir on the family Blepharostomataceae, II. Candollea 21 (2): 241–355. Schuster RM (1966d) he hepaticae and anthocerotae of North America. I. Columbia University Press, New York, 802 pp. Schuster RM (1966e) A memoir on the family Blepharostomataceae, I. Candollea 21 (1): 59–136. Schuster RM (1966f) Studies on hepaticae, XXVIII. On Phycolepidozia, a new, highly reduced genus of Jungermanniales of questionable ainity. Bulletin of the Torrey Botanical Club 93 (6): 437–449. doi: 10.2307/2483417 Schuster RM (1967a) North American Lejeuneaceae. X. Harpalejeunea, Drepanolejeunea, and Leptolejeunea. Journal of the Elisha Mitchell Scientiic Society 83 (4): 192–229. Schuster RM (1967b) A note on the genus Gymnocolea Dum. Bryologist 70 (1): 111–112. doi: 10.2307/3241147 Schuster RM (1967c) Studies on hepaticae. XV. Calobryales. Nova Hedwigia 13 (1/2): 1–76. Schuster RM (1967d) Studies on antipodal hepaticae. IX. Phyllothalliaceae. Transactions of the British Bryological Society 5 (2): 283–288. doi: 10.1179/006813867804804296 Schuster RM (1968a) Studies on antipodal hepaticae. X. Subantarctic Scapaniaceae, Balantiopsidaceae and Schistochilaceae. Bulletin of the National Science Museum, Tokyo (n.ser.) 11 (1): 13–31. Schuster RM (1968b) Studies on the hepaticae, XXIX-XLIV. A miscellany of new taxa and new range extensions. Nova Hedwigia 15: 437–529. Schuster RM (1969a) Studies on hepaticae XLVI-XLVII. On Alobiella (Spr.) Schifn. and Alobiellopsis Schust. Bulletin of the National Science Museum, Tokyo (n.ser.) 12 (3): 659– 683. 786 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Schuster RM (1969b) he hepaticae and anthocerotae of North America. II. Columbia University Press, New York, 1062 pp. Schuster RM (1969c) Problems of the antipodal distribution in lower land plants. Taxon 18 (1): 46–91. doi: 10.2307/1218591 Schuster RM (1970a) Studies on antipodal hepaticae, III. Jubulopsis Schuster, Neohattoria Kamimura and Amphijubula Schuster. Journal of the Hattori Botanical Laboratory 33: 266–304. Schuster RM (1970b) Studies on hepaticae, XLIX-LIII. New Lejeuneaceae from Dominica and Jamaica. Bulletin of the Torrey Botanical Club 97 (6): 336–352. doi: 10.2307/2483854 Schuster RM (1971a) he ecology and distribution of hepaticae in a mahogany hammock in tropical Florida. Castanea 36 (2): 90–111. Schuster RM (1971b) Studies on Cephaloziellaceae. Nova Hedwigia 22: 121–265. Schuster RM (1971c) Studies of antipodal Schistochilaceae and Scapaniaceae. Bulletin of the National Science Museum, Tokyo (n.ser.) 14 (4): 609–660. Schuster RM (1971d) Two new antipodal species of Haplomitrium (Calobryales). Bryologist 74 (2): 131–143. doi: 10.2307/3241827 Schuster RM (1972) Phylogenetic and taxonomic studies in Jungermanniidae. Journal of the Hattori Botanical Laboratory 36: 321–405. Schuster RM (1974) he hepaticae and anthocerotae of North America. III. Columbia University Press, New York, 880 pp. Schuster RM (1978a) Studies on Venezuelan hepaticae. I. Phytologia 39 (4): 239–251. Schuster RM (1978b) Studies on Venezuelan hepaticae. II. Phytologia 39 (6): 425–432. Schuster RM (1980a) Studies on hepaticae. LIV-LVIII. Kurzia v. Mart. [Microlepidozia (Spr.) Joerg.], Megalembidium Schust., Psiloclada Mitt., Drucella Hodgs. and Isolembidium Schust. Journal of the Hattori Botanical Laboratory 48: 337–421. Schuster RM (1980b) New combinations and taxa of hepaticae I. Phytologia 45 (5): 415–437. Schuster RM (1980c) he hepaticae and anthocerotae of North America. IV. Columbia University Press, New York, 1334 pp. Schuster RM (1981a) Austral hepaticae. VIII. Tuyamaelloideae. Phytologia 47 (4): 301–308. Schuster RM (1981b) Evolution and speciation in Pellia, with special reference to the Pellia megaspora-endiviifolia complex (Metzgeriales). I. Taxonomy and distribution. Journal of Bryology 11 (3): 411–431. doi: 10.1179/jbr.1981.11.3.411 Schuster RM (1982) Studies on hepaticae, LIX. On Sandeothallus Schust., gen. n. and the classiication of the Metzgeriales. Nova Hedwigia 36: 1–16. Schuster RM (1983a) Phytogeography of the Bryophyta. In: Schuster RM (Ed.) New Manual of Bryology, vol. 1. Hattori Botanical Laboratory, Nichinan, 463–626. Schuster RM (1983b) Notes on nearctic hepaticae. XVI. New taxa of Frullania from eastern North America. Phytologia 53 (5): 364–366. Schuster RM (1984) Diagnoses of some new taxa of hepaticae. Phytologia 56 (2): 65–74. Schuster RM (1985a) Studies in Porellineae: New taxa of Jubulaceae. Phytologia 57 (5): 369–373. Schuster RM (1985b) Some new taxa of hepaticae. Phytologia 57 (6): 408–414. Schuster RM (1985c) Austral hepaticae. XIX. Some taxa new to New Zealand and New Caledonia. Phytologia 56 (7): 449–464. World checklist of hornworts and liverworts 787 Schuster RM (1986) Gymnocolea borealis (Frisvoll & Moen) Schust. [Lophozia (Leiocolea) borealis Frisvoll & Moen] in North America. Lindbergia 12 (1): 5–8. Schuster RM (1987a) Phylogenetic studies on Jungermanniidae. II. Mastigophoraceae and Chaetophyllopsidaceae. Memoirs of the New York Botanical Garden 45: 733–748. Schuster RM (1987b) Preliminary studies on anthocerotae. Phytologia 63 (3): 193–201. Schuster RM (1987c) On Aureolejeunea Schust. and Brachiolejeunea paramicola Herzog. Phytologia 61 (7): 445–447. Schuster RM (1987d) Studies on Metzgeriales: I. North American Aneuraceae. Journal of the Hattori Botanical Laboratory 62: 299–329. Schuster RM (1989) Studies on the hepatic lora of Prince Edward Islands. I. Aneuraceae. Journal of the Hattori Botanical Laboratory 67: 59–108. Schuster RM (1991a) On neotenic species of Radula. Journal of the Hattori Botanical Laboratory 70: 51–62. Schuster RM (1991b) Diagnoses of new taxa of hepaticae. I. Jungermanniidae. Journal of the Hattori Botanical Laboratory 70: 143–150. Schuster RM (1992a) he oil-bodies of the hepaticae. II. Lejeuneaceae (part 2). Journal of the Hattori Botanical Laboratory 72: 163–359. Schuster RM (1992b) he hepaticae and anthocerotae of North America. V. Columbia University Press, New York, 854 pp. Schuster RM (1992c) Studies on Marchantiales. I-III. Journal of the Hattori Botanical Laboratory 71: 267–287. Schuster RM (1992d) he hepaticae and anthocerotae of North America. VI. Columbia University Press, New York, 937 pp. Schuster RM (1993a) Studies on hepaticae, LXII-LXIV. Lepidoziaceae subfamily Zoopsidoideae (1). Nova Hedwigia 56 (1/2): 35–59. Schuster RM (1993b) On Cephalozia pachycaulis sp. nov. and the perimeters of Cephalozia. Bryologist 96 (4): 619–625. doi: 10.2307/3243994 Schuster RM (1994) Studies on Lejeuneaceae. I. Preliminary studies on new genera of Lejeuneaceae. Journal of the Hattori Botanical Laboratory 75: 211–235. Schuster RM (1995a) Phylogenetic and taxonomic studies of Jungermanniidae. III. Calypogeiaceae. Fragmenta Floristica et Geobotanica 40 (2): 825–888. Schuster RM (1995b) Studies on Cephaloziellaceae. III. On Cephalomitrion Schust., gen. n. Nova Hedwigia 61 (3/4): 547–559. Schuster RM (1995c) he hepaticae of Prince Edward Islands. II. On Gymnocoleopsis (Schust.) Schust., Lophozia cylindriformis (Mitt.) Steph. and the subgeneric classiication of the genus Lophozia Dumort. Journal of the Hattori Botanical Laboratory 78: 119–135. Schuster RM (1995d) On a new species of Gymnomitrion, G. mucrophorum Schust., sp. n. Bryologist 98 (2): 242–245. doi: 10.2307/3243310 Schuster RM (1996a) Studies on antipodal hepaticae. XII. Gymnomitriaceae. Journal of the Hattori Botanical Laboratory 80: 1–147. Schuster RM (1996b) On Olgantha Schust., gen. n. Isophylly and evolution of Jungermanniales. Nova Hedwigia 63 (3/4): 529–543. 788 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Schuster RM (1996c) Studies on Lejeuneaceae II. Neotropical taxa of Drepanolejeunea (Spr.) Schifn. Nova Hedwigia 62 (1/2): 1–46. Schuster RM (1996d) Studies on Cephaloziellaceae. IV. On New Zealand taxa. Nova Hedwigia 63 (1/2): 1–61. Schuster RM (1997a) On Bragginsella, a new genus of Jungermanniales from New Zealand. Bryologist 100 (3): 362–367. doi: 10.2307/3244506 Schuster RM (1997b) On a new, microphyllous New Caledonean Acromastigum (Lepidoziaceae). Nova Hedwigia 64 (3/4): 613–620. Schuster RM (1998a) On Lejeunea (Papillolejeunea) pocsii Schust., sp. n. of New Zealand. Journal of the Hattori Botanical Laboratory 85: 83–87. Schuster RM (1998b) On the genus Scaphophyllum (Jungermanniaceae). Bryologist 101 (3): 428–434. doi: 10.2307/3244182 Schuster RM (1999a) Studies on hepaticae. LXVII-LXVIII. Lepidoziaceae subfamily Zoopsidoideae (4): Monodactylopsis and Pteropsiella. Nova Hedwigia 69 (3/4): 517–540. Schuster RM (1999b) Studies on hepaticae LXVI. Lepidoziaceae subfamily Zoopsidoideae (3): Zoopsidella. Nova Hedwigia 69 (1/2): 101–150. Schuster RM (1999c) Harpalejeunea (Spr.) Schifn. I. Studies on a new Andean species of Harpalejeunea. Journal of the Hattori Botanical Laboratory 87: 287–294. Schuster RM (1999d) Studies on hepaticae, LXV. Lepidoziaceae subfamily Zoopsidioideae (2): Zoopsis. Nova Hedwigia 68 (1/2): 1–64. Schuster RM (2000a) Austral hepaticae. Part 1. Beihefte zur Nova Hedwigia 118: 1–524. Schuster RM (2000b) Studies on Lejeuneaceae. III. Revisionary studies on Stenolejeunea Schust. Journal of the Hattori Botanical Laboratory 89: 151–170. Schuster RM (2000c) Studies on Lejeuneaceae. II. Rectolejeunea Evs. emend. Schust. (Lejeuneoideae). Journal of the Hattori Botanical Laboratory 89: 113–150. Schuster RM (2001a) Revisionary studies on austral Acrobolbaceae, I. Journal of the Hattori Botanical Laboratory 90: 97–166. Schuster RM (2001b) Studies on hepaticae. LXI. Trichocoleaceae. Nova Hedwigia 73 (3/4): 461–486. Schuster RM (2002a) Revisionary studies of the Chandonanthoideae (Jungermanniales, Jungermanniaceae). Nova Hedwigia 74 (3/4): 465–496. doi: 10.1127/0029-5035/2002/0074-0465 Schuster RM (2002b) Austral hepaticae. Part II. Beihefte zur Nova Hedwigia 119: 1–606. Schuster RM (2006) Studies on Lejeuneaceae. V. Leucolejeunea and allies. Journal of the Hattori Botanical Laboratory 100: 361–406. Schuster RM, Damsholt K (1974) he hepaticae of west Greenland from ca. 66° N to 72° N. Meddelelser om Grønland 199 (1): 1–373. Schuster RM, Damsholt K (1987) Some new taxa of Jungermanniales. Phytologia 63 (5): 325–328. Schuster RM, Engel JJ (1973) Austral hepaticae. II. Evansianthus, a new genus of Geocalycaceae. Bryologist 76 (4): 516–520. doi: 10.2307/3241410 Schuster RM, Engel JJ (1974) A monograph of the genus Pseudocephalozia (Hepaticae). Journal of the Hattori Botanical Laboratory 38: 665–701. Schuster RM, Engel JJ (1975) Austral hepaticae. V. Studies on Schistochilaceae. Phytologia 30 (4): 241–250. World checklist of hornworts and liverworts 789 Schuster RM, Engel JJ (1977) Austral hepaticae. V. he Schistochilaceae of South America. Journal of the Hattori Botanical Laboratory 42: 273–423. Schuster RM, Engel JJ (1981) Austral hepaticae. XIII. Two new genera of Geocalycaceae (Lophocoleaceae). Phytologia 47 (4): 309–312. Schuster RM, Engel JJ (1982) Austral hepaticae. XVI. Gondwanalandic Leptoscyphoideae (Geocalycaceae). Lindbergia 8 (2): 65–74. Schuster RM, Engel JJ (1985) Austral hepaticae. V(2). Temperate and subantarctic Schistochilaceae of Australasia. Journal of the Hattori Botanical Laboratory 58: 255–539. Schuster RM, Engel JJ (1987a) Austral hepaticae. XX. New species of Hygrolembidium (Lepidoziaceae). Phytologia 62 (1): 9–12. Schuster RM, Engel JJ (1987b) A monograph of Lepidoziaceae subfam. Lembidioideae (Hepaticae). Journal of the Hattori Botanical Laboratory 63: 247–350. Schuster RM, Engel JJ (1996) Austral hepaticae. XXI. Paracromastigum iordlandiae (sp. nov.) and the delimitation of Paracromastigum and Hyalolepidozia (Lepidoziaceae). Brittonia 48 (2): 165–173. doi: 10.2307/2807810 Schuster RM, Engel JJ (1997) Austral hepaticae. XXIV. A revision of Isotachis Mitt. (Balantiopsaceae: Isotachidoideae) in New Zealand. Journal of the Hattori Botanical Laboratory 83: 187–227. Schuster RM, Giancotti C (1993) On Vitalianthus Schust. & Giancotti, a new genus of Lejeuneaceae. Nova Hedwigia 57 (3/4): 445–456. Schuster RM, Hattori S (1954) he oil bodies of the hepaticae. II. he Lejeuneaceae. Journal of the Hattori Botanical Laboratory 11: 11–86. Schuster RM, Inoue H (1975) Studies on Pallaviciniaceae and Allisoniaceae (Metzgeriales) in Japan. 1. Introduction and genus Hattorianthus, gen. nov. Bulletin of the National Science Museum, Tokyo. Series B, Botany 1 (3): 101–107. Schuster RM, Konstantinova NA (1995) Studies on Treubiales, I. On Apotreubia Hatt. et al. and A. hortonae Schust. & Konstantinova, sp. n. Journal of the Hattori Botanical Laboratory 78: 41–61. Schuster RM, Scott GAM (1969) A study of the family Treubiaceae (Hepaticae; Metzgeriales). Journal of the Hattori Botanical Laboratory 32: 219–268. Schuster RM, Steere WC (1958) Hygrolejeunea alaskana sp. n., a critical endemic of northern Alaska. Bulletin of the Torrey Botanical Club 85 (3): 188–196. doi: 10.2307/2483215 Schuster RM, Steere WC, homson JW (1959) he terrestrial cryptogams of northern Ellesmere Island. Bulletin of the National Museum of Canada 164: 1–132. Schwägrichen CF (1814) Historiae muscorum hepaticarum, prodromus. Joannis Ambrosii Barth, Lipsiae [Leipzig], 39 pp. Schweinfurth G (1866) Beitrag zur Flora Aethiopiens, Erste Abtheilung. Georg Reimers, Berlin, 311 pp. Schweinitz LD (1821) Specimen lorae americae septentrionalis cryptogamicae. J. Gales, Raleigh 27 pp. doi: 10.5962/bhl.title.62456 Scopoli JA (1772) Flora carniolica. Tom. II. Editio secunda. Joannis Pauli Krauss, Viennae [Wien], 496 pp. Scott EB (1960) A monograph of the genus Lepicolea (Hepaticae). Nova Hedwigia 2: 129–172. Scott GAM, Beckmann K (1987) he biology of Lethocolea squamata. Symposia Biologica Hungarica 35: 209–214. 790 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Scott GAM, Bradshaw JA (1985) Australian liverworts (Hepaticae): Annotated list of binominals and check-list of published species with bibliography. Brunonia 8 (1): 1–171. doi: 10.1071/BRU9850001 Scott EB, Miller HA (1959) Notes on Hawaiian hepaticae IV. Herberta herpocladioides sp. nov. Bryologist 62 (2): 116–118. doi: 10.2307/3240029 Scott GAM, Pike DC (1984) New species of Fossombronia from Australia. Journal of the Hattori Botanical Laboratory 56: 339–349. Scott GAM, Pike DC (1987) Studies on Fossombronia in Australia. II. Fourteen more new species. Journal of the Hattori Botanical Laboratory 62: 367–386. Scott GAM, Pike DC (1988a) A new species of Fossombronia from Australia. Beihefte zur Nova Hedwigia 90: 109–112. Scott GAM, Pike DC (1988b) Revisionary notes on Fossombronia. Bryologist 91 (3): 193–201. doi: 10.2307/3243219 Seaward MRD, Ellis LT, Pócs T, Wigginton MJ (2006) Bryophyte lora of the Chagos Archipelago. Journal of Bryology 28 (1): 11–19. doi: 10.1179/174328206X90440 Segarra-Moragues JG, Puche F (2014) Advances in the knowledge of South African Riella (Sphaerocarpales) and a new endemic species, Riella trigonospora. South African Journal of Botany 94: 166–176. doi: 10.1016/j.sajb.2014.06.013 Segarra-Moragues JG, Puche F, Sabovljević M (2012) Riella heliospora (Riellaceae) a new monoecious species of Riella subgenus Trabutiella from California. Systematic Botany 37 (2): 307–319. doi: 10.1600/036364412X635368 Segarra-Moragues JG, Puche F, Sabovljević M, Infante M, Heras P (2014) Taxonomic revision of Riella subgenus Trabutiella (Riellaceae, Sphaerocarpales). Phytotaxa 159 (3): 131–174. doi: 10.11646/phytotaxa.159.3.1 Seppelt RD (1998) he genus Riccia (Marchantiales: Ricciaceae), in South Australia. Hikobia 12: 317–341. Sérgio C, Perold SM (1992) A new species of Riccia L. from the island of Madeira, Riccia atlantica, sp. nov. Journal of Bryology 17 (1): 127–132. doi: 10.1179/jbr.1992.17.1.127 Sérgio C, Sim-Sim M (1989) Riccia incrassata Brotero, a neglected name of Oxymitra paleacea Bisch. ex Lindenb. Journal of Bryology 15 (4): 659–663. doi: 10.1179/jbr.1989.15.4.659 Sharma D, Srivastava SC (1993) Indian Lepidoziinae (a taxonomic revision). Bryophytorum Bibliotheca 47: 1–353. Shaw B, Crandall-Stotler B, Váňa J, Stotler RE, von Konrat M, Engel JJ, Davis CE, Long DG, Sova P, Shaw AJ (2015) Phylogenetic relationships and morphological evolution in a major clade of leafy liverworts (phylum Marchantiophyta, order Jungermanniales): suborder Jungermanniineae. Systematic Botany 40 (1): 27–45. doi: 10.1600/036364415X686314 Shi X-Q, Gradstein SR, Zhu R-L (2015a) Phylogeny and taxonomy of Archilejeunea (Marchantiophyta: Lejeuneaceae: Ptychanthoideae) based on molecular markers and morphology. Taxon 64 (5): 881–892. doi: 10.12705/645.1 Shi X-Q, Zhu R-L (2015b) A revision of Archilejeunea s.str. (Lejeuneaceae, Marchantiophyta). Nova Hedwigia 100 (3/4): 589–601. doi: 10.1127/nova_hedwigia/2015/0246 World checklist of hornworts and liverworts 791 Shi X-Q, Gradstein SR, Zhu R-L (2015) Type studies on Archilejeunea (Lejeuneaceae, Marchantiophyta): six new synonyms and a new combination. Phytotaxa 195 (3): 248–250. doi: 10.11646/phytotaxa.195.3.5 Shimizu D, Hattori S (1952) Studies on the Japanese species of Asterella (1). Journal of the Hattori Botanical Laboratory 8: 46–54. Shimizu D, Hattori S (1953a) Marchantiales of Japan, I. Journal of the Hattori Botanical Laboratory 9: 32–44. Shimizu D, Hattori S (1953b) Marchantiales of Japan (2). Journal of the Hattori Botanical Laboratory 10: 49–55. Shimizu D, Hattori S (1954) Marchantiales of Japan. III. Journal of the Hattori Botanical Laboratory 12: 53–75. Shimizu D, Hattori S (1955) Marchantiales of Japan. IV. Journal of the Hattori Botanical Laboratory 14: 91–107. Shliakov RN (1971) Dve novye kruppokletochinye razpovidnosti pechenochnikov [Varietates duae novae magnicellulares hepaticarum]. Novosti Sistematiki Nizših Rastenij 8: 327–334. Shliakov RN (1973) Sistematicheskie zametkino semejtvo Lophoziaceae Cavers [Notulae systematicae in Lophoziaceis Cavers]. Novosti Sistematiki Nizših Rastenij 10: 287–302. Shliakov RN (1975) Dopolneniâ k lore pechenochnikov Sovetskoj arktiki [Additamenta ad loram hepaticarum arcticae URSS]. Novosti Sistematiki Nizših Rastenij 12: 318–323. Shliakov RN (1976) Novye kombinacii i novye taksony pechenochnyh mkhov (anthocerotae i hepaticae) [Combinationes et taxa anthocerotarum et hepaticarum nova]. Novosti Sistematiki Nizših Rastenij 13: 225–229. Shliakov RN (1978) Novye vidy pechninochnikov iz Sibiri i dal’nego vostoka [Hepaticarum species novae e Sibiria et oriente extremo]. Novosti Sistematiki Nizših Rastenij 15: 242–247. Shliakov RN (1979) Novye dopolneniia k lore pechenochnikov severnykh rajonov SSSR [Additamenta nova ad loram hepaticarum regionum septentrionalium URSS]. Novosti Sistematiki Nizših Rastenij 16: 201–208. Shliakov RN (1980a) Pechenochnye Mkhi Severa SSSR. 3. Nauka, Leningrad, 188 pp. Shliakov RN (1980b) Novye interesnye nakhodki pechenochnikov iz severnykh rajonov SSSR [De hepaticis in regionibus borealibus URSS inventis novis et curiosis. Novosti Sistematiki Nizših Rastenij 17: 235–241. Shliakov RN (1981) Pechenochnye Mkhi Severa SSSR. 4. Nauka, Leningrad, 221 pp. Shliakov RN (1982) Novye kombinacii i novye nazvaniia mokhoobraznykh [Combinationes et nomina bryophytorum novae]. Novosti Sistematiki Nizših Rastenij 19: 209–210. Shu L, Zhu RL (2014) Notes on Early Land Plants Today. 59. Leptolejeunea himalayensis and Leptocolea mirpurensis (Marchantiophyta, Lejeuneaceae) synonymous with Lejeunea cocoes and Cololejeunea raduliloba, respectively. Phytotaxa 173 (1): 95–96. doi: 10.11646/phytotaxa.173.1.11 Sim TR (1926) he bryophyta of South Africa. Transactions of the Royal Society of South Africa 15 (1): 1–475. doi: 10.1080/00359192609519311 Sim-Sim M, Sérgio C, Mues R, Kraut L (1995) A new Frullania species (Trachycolea) from Portugal and Macaronesia, Frullania azorica sp. nov. Cryptogamie: Bryologie, Lichénologie 16 (2): 111–123. 792 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Sim-Sim M, Fontinha S, Mues R, Lion U (2000) A new Frullania species (subg. Frullania) from Deserta Grande, Madeira archipelago, Frullania sergiae sp. nov. Nova Hedwigia 71 (1/2): 185–193. Singh DK (1983a) Trichocolea udarii Singh – a new hepatic from Jaintia Hills (Meghalaya), India. Bulletin of the Botanical Survey of India 25: 177–180. Singh DK (1983b) Cyathodium mehranum Singh, sp. nov., from Arunachal Pradesh, India. Miscellanea Bryologica et Lichenologica 9 (8): 172–177. Singh DK (1987a) A new species of Notothylas Sull. (Bryophyta) from Nepal. Journal of the Bombay Natural History Society 84 (3): 650–653. Singh DK (1987b) A new species of Folioceros Bharad. (Anthocerotaceae) from Arunachal Pradesh, India. Bulletin of the Botanical Survey of India 29: 176–180. Singh SK (2011) A new Rectolejeunea from Indian botanic garden, India. Indian Journal of Forestry 34 (3): 341–344. Singh SK (2013) New combinations in Lejeunea with a new name to Otigoniolejeunea indica. Phytotaxa 96 (1): 63–64. doi: 10.11646/phytotaxa.96.1.3 Singh SK, Barbhuiya HA (2012) Contribution to the hepaticae and anthocerotae of Mizoram V. hree New Taxa of Frullania from India. Taiwania 57 (2): 106–116. Singh SK, Dey M (2012) A new species of Drepanolejeunea (Marchantiophyta: Lejeuneaceae) from India. Nelumbo 54: 20–23. Singh AP, Nath V (2007a) A new Calypogeia Raddi from India. Taiwania 52 (4): 320–323. Singh AP, Nath V (2007b) Hepaticae of Khasi and Jaintia Hills: Eastern Himalayas. Bishen Singh Madendra Pal Singh, Dehra Dun, 382 pp. Singh AP, Nath V (2008) A new species of Trocholejeunea (Hepaticae: Ptychanthoideae) from Meghalaya, Eastern Himalayas, India. Journal of Japanese Botany 83 (1): 1–6. Singh DK, Semwal RC (2000) A new species of Notothylas Sull. (Bryophyta) from Uttaranchal, India. Indian Journal of Forestry 23 (4): 386–389. Singh DK, Semwal RC (2001) A new species of Notothylas Sull. from Dehradun, India. Phytotaxonomy 1: 35–39. Singh SK, Singh DK (2005) Frullania larjiana (Jungermanniopsida: Frullaniaceae), a new species from India. Journal of Bryology 27 (2): 105–108. doi: 10.1179/037366805X53004 Singh SK, Singh DK (2007a) Cephalozia schusteri (Cephaloziaceae, Hepaticae) – a new species from India, with note on the Indian species of the genus. Lindbergia 32 (1): 1–4. Singh SK, Singh DK (2007b) Jungermannia indrodayana (Jungermanniaceae, Hepaticae) – a new species from India. Cryptogamie, Bryologie 28 (2): 103–108. Singh D, Singh DK (2012) A new species of the genus Lepidozia (Marchantiophyta, Lepidoziaceae) from Sikkim, India. Nova Hedwigia 94 (1/2): 221–225. doi: 10.1127/00295035/2012/0094-0221 Smith JE, Sowerby J (1800) English botany, vol. 10. J. Davis, London, 649–720. Smith JE, Sowerby J (1806) English botany, vol. 23. J. Davis, London, 1585–1656. Smith JE, Sowerby J (1811) English botany, vol. 32. J. Davis, London, 2233–2304. Smith JE, Sowerby J (1814) English botany, vol. 36. J. Davis, London, 2521–2592. So ML (1999) Studies on Plagiochila sect. Poeltiae (Plagiochilaceae, Hepaticae). Haussknechtia, Beiheft 9: 347–358. World checklist of hornworts and liverworts 793 So ML (2000a) Plagiochila sect. Contiguae (Hepaticae) in Australasia and the Paciic, with description of Plagiochila subjavanica sp. nov. Australian Systematic Botany 13 (5): 803–815. doi: 10.1071/SB00011 So ML (2000b) Plagiochila sect. Plagiochila (Hepaticae) in SE Asia and Melanesia, with description of two new species. New Zealand Journal of Botany 38 (3): 425–432. doi: 10.1080/0028825X.2000.9512694 So ML (2001a) Recognised species of Plagiochila in Australasia and some island groups of the Paciic Ocean. New Zealand Journal of Botany 39 (3): 395–421. doi: 10.1080/0028825X.2001.9512745 So ML (2001b) On several little known species of Plagiochila in Australasia and the Paciic with description of Plagiochila sublyallii sp. nov. from Papua New Guinea. New Zealand Journal of Botany 39 (1): 109–114. doi: 10.1080/0028825X.2001.9512718 So ML (2002a) he genus Porella (Porellaceae, Hepaticae) in Australasia and the South Paciic. Systematic Botany 27 (1): 4–13. doi: 10.1043/0363-6445-27.1.4 So ML (2002b) Metzgeria submarginata sp. nov., a “new” species from Australia and New Zealand. New Zealand Journal of Botany 40 (2): 201–205. doi: 10.1080/0028825X.2002.9512783 So ML (2002c) Metzgeria (Hepaticae) in Australasia and the Paciic. New Zealand Journal of Botany 40 (4): 603–627. doi: 10.1080/0028825X.2002.9512818 So ML (2003) he genus Herbertus (Hepaticae) in Australasia and the South Paciic. Systematic Botany 28 (1): 12–23. doi: 10.1043/0363-6445-28.1.12 So ML (2004) Metzgeria (Metzgeriaceae, Marchantiophyta) in Africa. New Zealand Journal of Botany 42 (2): 271–292. doi: 10.1080/0028825X.2004.9512904 So ML (2005a) Radula (Radulaceae, Marchantiophyta) in Hawaii. Journal of the Hattori Botanical Laboratory 98: 175–191. So ML (2005b) Porella (Porellaceae, Marchantiophyta) in Latin America. New Zealand Journal of Botany 43 (1): 301–321. doi: 10.1080/0028825X.2005.9512956 So ML (2006) Radula (Radulaceae, Marchantiophyta) in the South Paciic. Journal of the Hattori Botanical Laboratory 99: 207–232. So ML, Grolle R (2000a) Checklist of Plagiochila (Hepaticae) in Asia. Journal of the Hattori Botanical Laboratory 88: 199–243. So ML, Grolle R (2000b) Description of Plagiochila detecta sp. nov. (Hepaticae) from East Asia based on morphological and RAPD evidence. Nova Hedwigia 71 (3/4): 387–394. So ML, Grolle R (2001) On Plagiochila subgenus Plagiochila section Abietinae (Hepaticae). Systematic Botany 26 (3): 459–469. doi: 10.1043/0363-6445-26.3.459 So ML, Wang J (2006) Frullania (Frullaniaceae, Marchantiophyta) in Hawaii. Journal of the Hattori Botanical Laboratory 100: 419–430. So ML, Zhu R-L (1998) On six species of the genus Lejeunea in China, including one new species. Bryologist 101 (1): 137–143. doi: 10.2307/3244086 Söderström L, Karttunen K, Hedenäs L (1992) Nomenclatural notes on Fennoscandian bryophytes. Annales Botanici Fennici 29 (2): 119–122. Söderström L, Weibull H, Damsholt K (2000) A new species of Lophozia (subg. Protolophozia) from Fennoscandia. Lindbergia 25 (1): 3–7. 794 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Söderström L, Urmi E, Váňa J (2002) Distribution of hepaticae and anthocerotae in Europe and Macaronesia. Lindbergia 27 (1): 3–47. Söderström L, Hagborg A, von Konrat M, Renner MAM (2008) Early Land Plants Today: Liverwort checklist of checklists. Fieldiana: Botany (n.ser.) 47: 105–130. doi: 10.3158/0015-0746-47.1.105 Söderström L, Gradstein SR, Hagborg A (2010a) Checklist of the hornworts and liverworts of Java. Phytotaxa 9: 53–149. doi: 10.11646/phytotaxa.9.1.7 Söderström L, de Roo RT, Hedderson TAJ (2010b) Taxonomic novelties resulting from recent reclassiication of the Lophoziaceae/Scapaniaceae clade. Phytotaxa 3: 47–53. doi: 10.11646/phytotaxa.3.1.7 Söderström L, Hagborg A, Pócs T, Sass-Gyarmati A, Brown E, von Konrat MJ, Renner MAM (2011a) Checklist of hornworts and liverworts of Fiji. Telopea 13 (3): 405–454. Söderström L, Hagborg A, Váňa J, von Konrat M (2011b) Land of wood and water: A checklist of liverworts and hornworts of Jamaica. Bryologist 114 (1): 67–91. doi: 10.1639/00072745-114.1.67 Söderström L, Hagborg A, von Konrat MJ (2012a) Notes on Early Land Plants Today. 1. A new name for Cryptothallus hirsutus. Phytotaxa 65: 43. Söderström L, Hagborg A, Crosby MR, von Konrat MJ (2012b) Early Land Plants Today: Index of Liverworts & Hornworts 2009–2010. Phytotaxa 63: 21–68. Söderström L, Hagborg A, von Konrat MJ (2012c) Notes on Early Land Plants Today. 5. Validation of two Drepanolejeunea species. Phytotaxa 65: 47–48. Söderström L, Hagborg A, von Konrat M (2012d) Notes on Early Land Plants Today. 6. he correct name for Cephaloziella divaricata var. asperifolia. Phytotaxa 65: 49–50. Söderström L, von Konrat M, Hagborg A, Séneca A (2012e) Early Land Plants Today – a community driven approach. Studia Botanica Hungarica 43: 47–58. Söderström L, Hagborg A, von Konrat MJ (2012f) Notes on Early Land Plants Today. 9. Validation of Riccia gangetica Ahmad. Phytotaxa 65: 57. Söderström L, Hagborg A, von Konrat M (2012g) he friendly islands – a checklist of hornworts and liverworts of Tonga. Polish Botanical Journal 57 (1): 129–135. Söderström L, Váňa J, Hagborg A, von Konrat M (2013a) Notes on Early Land Plants Today. 31. Lophonardia replaces Hypolophozia (Cephaloziellaceae, Marchantiophyta). Phytotaxa 81 (1): 19–21. doi: 10.11646/phytotaxa.81.1.7 Söderström L, Crandall-Stotler B, Stotler RE, Váňa J, Hagborg A, von Konrat M (2013b) Notes on Early Land Plants Today. 36. Generic treatment of Lophocoleaceae (Marchantiophyta). Phytotaxa 97 (2): 36–43. doi: 10.11646/phytotaxa.97.2.3 Söderström L, Váňa J, Hagborg A, von Konrat M (2013c) Notes on Early Land Plants Today. 35. Notes on Lophoziaceae (Marchantiophyta). Phytotaxa 97 (2): 27–35. doi: 10.11646/ phytotaxa.97.2.2 Söderström L, Hagborg A, von Konrat MJ (2013d) Notes on Early Land Plants Today. 19. Validation of two names in Pellia (Pelliaceae, Marchantiophyta). Phytotaxa 76 (3): 39–40. doi: 10.11646/phytotaxa.76.3.7 Söderström L, Hagborg A, von Konrat M, Séneca A (2013e) he Guaraní Land – checklist of hornworts (Anthocerotophyta) and liverworts (Marchantiophyta) of Paraguay. Polish Botanical Journal 58 (1): 267–277. doi: 10.2478/pbj-2013-0027 World checklist of hornworts and liverworts 795 Söderström L, Váňa J, Crandall-Stotler B, Stotler RE, Hagborg A, von Konrat M (2013f) Notes on Early Land Plants Today. 43. New combinations in Lophocoleaceae (Marchantiophyta). Phytotaxa 112 (1): 18–32. doi: 10.11646/phytotaxa.112.1.4 Söderström L, Hagborg A, von Konrat M (2014) Early Land Plants Today: Index of liverworts & hornworts 2011-2012. Phytotaxa 170 (2): 61–85. doi: 10.11646/phytotaxa.170.2.1 Söderström L, Stotler RE, Gradstein SR, Barrie FR, Hagborg A, Crandall-Stotler BJ, von Konrat M (2015a) Notes on Early Land Plants Today. 73. Genera of Lejeuneaceae established in the period 1884–1893: dates of validation and implications. Phytotaxa 220 (2): 143– 198. doi: http://dx.doi.org/10.11646/phytotaxa.220.2.4 Söderström L, Hagborg A, von Konrat M (2015b) Notes on Early Land Plants Today. 69. Circumscription of Plagiochilaceae (Marchantiophyta) with a preliminary infrageneric subdivision of Plagiochila. Phytotaxa 208 (1): 75–91. doi: 10.11646/phytotaxa.208.1.8 Söderström L, Váňa J, Crandall-Stotler B, Hentschel J, Hagborg A, von Konrat MJ (2015c) Notes on Early Land Plants Today. 68. Miscellaneous notes on Marchantiophyta. Phytotaxa 202 (1): 69–72. doi: 10.11646/phytotaxa.202.1.10 Söderström L, Pócs T, Váňa J, Hagborg A (2015d) Notes on Early Land Plants Today. 74. Validations of a few names in Marchantiophyta. Phytotaxa 220 (2): 199–200. doi: 10.11646/ phytotaxa.220.2.5 Sofronova EV, Mamontov IuS, Potemkin AD (2013) Frullania ignatovii (Porellales, Marchantiophyta) — Novyj vid iz Sibiri [Frullania ignatovii, a new species from Siberia]. Novosti Sistematiki Nizših Rastenij 47: 334–343. Solari SS (1973) Miscelánea briológica (Hepaticae) I. Boletín de la Sociedad Argentina de Botánica 15 (2/3): 197–203. Solari SS (1976) Miscelánea briológica (Hepaticae). III. Darwiniana 20 (3/4): 387–390. Solari SS (1981) Miscelánea briológica (Hepaticae). IV. Novedades en Lejeuneaceae. Comunicaciones del Museo Argentino de Ciencias Naturales “Bernardino Rivadavia” e Instituto Nacional de Investigación de las Ciencias Naturales: Ciencias Botánicas 2 (11): 67–75. Solari SS (1983a) Lejeuneaceae – catálogo de especies andinopatagónicas. Journal of the Hattori Botanical Laboratory 54: 533–553. Solari SS (1983b) Patagonian bryophytes. 8. On the identity of Lepicolea scolopendra (Hook.) Trev. var. magellanica Gola and Sendtnera ochroleuca (Spreng.) Nees var. piligera De Not. Lindbergia 9 (2): 86–88. Solari SS (1986) Miscelánea briológica (Hepaticae). V. Consideraciones sobre el género Leptoscyphus. Cryptogamie: Bryologie, Lichénologie 7 (3): 219–223. Solari SS, Hässel GG (1983) Las hepáticas de Spegazzini, parte I. Bollettino del Museo Civico di Storia Naturale di Verona 10: 193–209. Sommerfeldt SC (1833) Bidrag til Spitsbergens og Beeren-Eilands lora, efter herbarier, medbragte af M. Keilhaue. Magazin for Naturvidenskaberne 11 (2): 234–252. Sprengel C (1804) Anleitung zur Kenntnis der Gewächse, Dritte Sammlung. Karl August Kümmel, Halle, 374 pp. Sprengel C (1809) Observationes de Jungermanniis aut plane nondum aut minus bene delineatis. Annalen der Wetterauischen Gesellschaft für die Gesammte Naturkunde 1: 21–26. Sprengel C (1821) Neue Entdeckungen im ganzen Umfang der Planzenkunde, Zweyter Band. Friedrich Fleischer, Leipzig, 363 pp. 796 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Sprengel C (1827a) Systema vegetabilium. Editio decima sexta, voluminis IV, pars I. Dieterich, Göttingen, 592 pp. doi: 10.5962/bhl.title.822 Sprengel C (1827b) Systema vegetabilium. Editio decima sexta, voluminis IV, pars II. Dieterich, Göttingen, 410 pp. doi: 10.5962/bhl.title.822 Spruce R (1847) Hepaticae pyrenaicae. London, no. 1–77. Spruce R (1849) he musci and hepaticae of the Pyrenees (cont.). Annals and Magazine of Natural History (ser. 2) 3 (18): 478–503. Spruce R (1850) he musci and hepaticae of the Pyrenees. Transactions of the Botanical Society of Edinburgh 3 (1/4): 103–216. doi: 10.1080/03746605009467580 Spruce R (1876a) On Anomoclada, a new genus of hepaticae, and on its allied genera, Odontoschisma and Adelanthus (concluded). Journal of Botany, British and Foreign 14: 230–235. Spruce R (1876b) On Anomoclada, a new genus of hepaticae, and on its allied genera, Odontoschisma and Adelanthus (continued). Journal of Botany, British and Foreign 14: 161–170. Spruce R (1881a) On Marsupella stableri n. sp. and some allied species of European Hepaticae. Revue Bryologique 8 (6): 89–104. Spruce R (1881b) Musci praeteriti. Journal of Botany, British and Foreign 19: 33–40. Spruce R (1882) On Cephalozia. Slater, Malton, 99 pp. doi: 10.5962/bhl.title.46289 Spruce R (1884) Hepaticae amazonicae et andinae. I. Transactions and Proceedings of the Botanical Society of Edinburgh 15: 1–308. Spruce R (1885) Hepaticae amazonicae et andinae. II. Transactions and Proceedings of the Botanical Society of Edinburgh 15: 309–588. Spruce R (1887a) Lejeunea holtii a new hepatic from Killarney. Journal of Botany, British and Foreign 25: 33–39. Spruce R (1887b) On a new Irish hepatic. Journal of Botany, British and Foreign 25: 209–211. Spruce R (1889) Hepaticae novae americanae tropicae et aliae. Bulletin de la Société Botanique de France (Congrès de Botanique) 36: clxxxix–ccvii. Spruce R (1890) Hepaticae bolivianae, in Andibus boliviae orientalis. Memoirs of the Torrey Botanical Club 1 (3): 113–140. Srivastava KP (1960) On a species of Anthoceros, A. satpurensis Sriv. sp. nov. Proceedings of the Indian Science Congress Association 47 (3): 337–338. Srivastava SN, Amakawa T (1991) A new species of Jungermannia from western Himalayas, India. Proceedings of the National Academy of Sciences of India. Section B, Biological Sciences 61 (2): 205–208. Srivastava SC, Asthana AK (1989) Two species of Folioceros from India, including F. kashyapii, sp. nov. Bryologist 92 (2): 219–224. doi: 10.2307/3243949 Srivastava SN, Rai M (2011) Frullania largiana var. didyhatii var. nov. from Kumaun region in Western Himalaya. Geophytology 41 (1/2): 109–112. Srivastava SC, Rawat KK (2001) Metzgeria sikkimensis sp. novo from Sikkim Himalayas, India. Geophytology 31 (1/2): 71–73. Srivastava SC, Sharma D (1990) A new species of Jubula Dumort. from Milam in Kumaon (western Himalaya). Proceedings of the Indian Academy of Sciences. Plant Sciences 100 (2): 85–89. World checklist of hornworts and liverworts 797 Srivastava SC, Singh P (1995) Some species of Jungermannia subgenus Solenostoma (Mitt.) Amak. from eastern Himalaya. In: Kumar SS (Ed.) Recent studies in Indian bryophytes. Bishen Singh Madendra Pal Singh, Dehra Dun, 147–156. Srivastava SC, Srivastava G (1989a) Two species of Cololejeunea from South India. Proceedings of the Indian Academy of Sciences. Plant Sciences 99 (2): 83–90. Srivastava SC, Srivastava A (1989b) he genus Heteroscyphus Schifn. in the western Himalayas. Lindbergia 15 (6): 195–202. Srivastava SC, Srivastava A (1993) A remarkable Scapania (Scapaniaceae) from Manali (Himachal Pradesh: Western Himalaya). Journal of the Indian Botanical Society 72: 237–240. Srivastava A, Srivastava SC (2002) Indian Geocalycaceae (Hepaticae). A taxonomic study. Bishen Singh Madendra Pal Singh, Dehra Dun, 246 pp. Srivastava SC, Srivastava S (2004) Two new Metzgerias from peninsular India. Phytotaxonomy 4: 79–86. Srivastava SC, Udar R (1976) Indian Aneuraceae. A monographic study. Biological Memoirs 1 (1/2): 121–154. Srivastava SC, Udar R (1977) Riccardia perssonii sp. nov. and R. tenuicostata Schifn., two hepatics new to India. Lindbergia 4 (1/2): 127–131. Srivastava SC, Verma PK (2004) Genus Arachniopsis Spruce – new to India with A. indica sp. nov. as new to science. National Academy Science Letters 27 (7/8): 269–272. Srivastava SC, Kumar D, Sharma D (1988) A new Lepidozia from eastern Himalayas, India. Cryptogamie: Bryologie, Lichénologie 9 (3): 235–240. Srivastava SC, Srivastava S, Dharma D (2003) A new Jungermannia (Solenostoma) from the Valley of Flowers, India. Lindbergia 28 (3): 129–133. Srivastava SC, Rawat KK, Verma PK (2006) An interesting Plagiochila from Kodaikanal (India). National Academy Science Letters 29 (7/8): 267–270. Srivastava S, Srivastava SC, Rawat KK (2013) Status of family Lophoziaceae (Hepaticae) in India. Nelumbo 55: 113–152. Staleu FA, Cowan R (1976-2009) Taxonomic literature: a selective guide to botanical publications and collections with dates, commentaries and types (TL2). Regnum Vegetabile 94, 98, 105, 110, 112, 115, 116, 125, 130, 132, 134, 135, 137, 149, 150. doi: 10.5962/bhl.title.48631 Stech M, Konstantinova NA, Frey W (2002) Molecular divergence between Treubia Goebel and Apotreubia S. Hatt. & Mizut., the two genera of the archaic liverwort class Treubiopsida (Hepaticophytina): Studies in austral temperate rain forest bryophytes 19. Nova Hedwigia 75 (1/2): 91–100. doi: 10.1127/0029-5035/2002/0075-0091 Steere WC, Inoue H (1974) Fossombronia alaskana, a new hepatic from arctic Alaska. Bryologist 77 (1): 63–71. doi: 10.2307/3241778 Steere WC, Inoue H (1978) he hepaticae of arctic Alaska. Journal of the Hattori Botanical Laboratory 44: 251–345. Steere WC, Schuster RM (1960) he hepatic genus Ascidiota Massalongo new to North America. Bulletin of the Torrey Botanical Club 87 (3): 209–215. doi: 10.2307/2482767 Ştefănuţ S (2008) he hornwort and liverwort atlas of Romania. Ars Docendi, Bucharest, 510 pp. Stephani F (1883) Zwei neue Lebermoose. Hedwigia 22 (10): 145–148. Stephani F (1884a) Die Gattung Radula (Fortsetzung). Hedwigia 23 (10): 145–159. 798 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Stephani F (1884b) Die Gattung Radula. Hedwigia 23 (8): 113–137. Stephani F (1884c) Die Gattung Radula (Fortsetzung). Hedwigia 23 (9): 129–137. Stephani F (1885a) Hepaticarum species novae vel minus cognitae. III. Hedwigia 24 (5): 214–218. Stephani F (1885b) Hepaticarum species novae vel minus cognitae. IV. Hedwigia 24 (6): 246–250. Stephani F (1885c) Hepaticae (in Contributiones para o estudo da Flora d’Africa). Boletim da Sociedade Broteriana 4: 170–184. Stephani F (1885d) Di una nuova specie di Plagiochila di F. Stephani. Annuario del Reale Istituto Botanico di Roma 2: 86. Stephani F (1885e) Neue und kritische Arten der Gattung Riccia. Hedwigia 24 (1): 2–7. Stephani F (1885f) Hepaticarum species novae vel minus cognitae 2. Hedwigia 24 (4): 166–168. Stephani F (1886a) Hepaticae von der Halbinsel Alaska, gesammelt 1881/82 von den Doctoren Arthur und Aurel Krause. Botanische Jahrbücher für Systematik, Planzengeschichte und Planzengeographie 8 (2): 96–99. Stephani F (1886b) Hepaticarum species novae vel minus cognitae. VI. Hedwigia 25 (4): 133–134. Stephani F (1886c) Hepaticae species novae vel minus cognitae VIII. Hedwigia 25 (6): 233–249. Stephani F (1886d) Hepaticae africanae. Botanische Jahrbücher für Systematik, Planzengeschichte und Planzengeographie 8 (2): 79–95. Stephani F (1886e) Hepaticae species novae vel minus cognitae V. Hedwigia 25 (1): 5–9. Stephani F (1886f) Hepaticarum species novae vel minus cognitae. VII. Hedwigia 25 (5): 202–208. Stephani F (1887) Ueber einige Lebermoose Portugals. Hedwigia 26 (1): 1–6. Stephani F (1888a) Hepaticae africanae. Hedwigia 27 (2): 59–63. Stephani F (1888b) Westindische Hepaticae II. Hepaticae ex insulis St. Domingo et Dominica, quas collegit Eggers. Hedwigia 27 (11/12): 299–302. Stephani F (1888c) Westindische Hepaticae I. Hepaticae portoricenses. Hedwigia 27 (11/12): 276–299. Stephani F (1888d) Hepaticae africanae. Hedwigia 27 (3/4): 106–113. Stephani F (1889a) Hepaticae australiae 1. Hedwigia 28 (2): 128–135. Stephani F (1889b) Deux nouvelles espèces du genre Riccia. Revue Bryologique 16 (5): 65–67. Stephani F (1889c) Hepaticae australiae III. Hedwigia 28 (4): 257–278. Stephani F (1889d) Hepaticae australiae II. Hedwigia 28 (3): 155–175. Stephani F (1890a) Die Gattung Lejeunea im Herbarium Lindenberg. Hedwigia 29 (1): 1–23. Stephani F (1890b) Die Gattung Lejeunea im Herbarium Lindenberg (Fortsetzung). Hedwigia 29 (2): 68–99. Stephani F (1890c) Hepaticae africanae novae in insulis Bourbon, Maurice et Madagascar lectae. Botanical Gazette 15 (11): 281–292. doi: 10.1086/326585 Stephani F (1890d) Die Gattung Lejeunea im Herbarium Lindenberg (Schluss). Hedwigia 29 (3): 133–142. Stephani F (1891a) Hepaticae africanae. Hedwigia 30 (5): 201–217. Stephani F (1891b) Hepaticae. In: Renauld F, Cardot J (Eds) Musci exotici novi vel minus cogniti. II. Bulletin de la Société Royale de Botanique de Belgique, Comptes-rendus des Séances 30 (2): clxxxi–ccvii. Stephani F (1891c) Hepaticae africanae. Hedwigia 30 (6): 265–272. World checklist of hornworts and liverworts 799 Stephani F (1892a) Hepaticae novae caucasicae. Botanisches Centralblatt 50 (3): 70–72. Stephani F (1892b) A revision of Colenso’s hepaticae with descriptions of new species collected by him. Journal of the Linnean Society. Botany 29 (201): 263–280. doi: 10.1111/j.10958339.1892.tb02036.x Stephani F (1892c) Hepaticae. In: Renauld F, Cardot J (Eds) Musci exotici novi vel minus cogniti. III. Bulletin de la Société Royale de Botanique de Belgique, Comptes-rendus des Séances 31: 100–123. Stephani F (1892d) Hepaticae africanae. Hedwigia 31 (3): 120–130. Stephani F (1892e) Hepaticae. In: Durand T, Pittier HF (Eds) Primitiae lorae costaricensis, vol. 1, fasc. 2. Jardin Botanique de l’État. Jardin Botanique de l’État : 0–151. doi: 10.5962/bhl.title.51686 Stephani F (1892f) he North American Lejeuneae. Botanical Gazette 17 (6): 170–173. doi: 10.1086/326806 Stephani F (1892g) Hepaticae africanae. Hedwigia 31 (4): 165–174. Stephani F (1893a) Hepaticarum species novae. Pars I. Hedwigia 32 (1): 17–29. Stephani F (1893b) Hepaticarum species novae. Pars II. Hedwigia 32 (3): 137–147. Stephani F (1893c) Hepaticarum species novae III. Hedwigia 32 (4): 204–214. Stephani F (1893d) Hepaticarum species novae IV. Hedwigia 32 (5): 315–327. Stephani F (1893e) Hepaticae. In: Renauld F, Cardot J (Eds) Musci exotici novi vel minus cogniti. IV. Bulletin de la Société Royale de Botanique de Belgique, Comptes-rendus des Séances 32 (2): 8–40. Stephani F (1894a) Hepaticarum species novae V. Hedwigia 33 (1): 1–10. Stephani F (1894b) Hepaticae chinenses. Mémoires de la Société Nationale des Sciences Naturelles et Mathématiques de Cherbourg 29: 207–228. Stephani F (1894c) Eine neue Lebermoos-Gattung. Österreichische Botanische Zeitschrift 44 (1): 1–5. doi: 10.1007/BF01789755 Stephani F (1894d) Hepaticarum species novae VI. Hedwigia 33 (3): 137–169. Stephani F (1895a) Hepaticae africanae. Botanische Jahrbücher für Systematik, Planzengeschichte und Planzengeographie 20 (3): 299–321. Stephani F (1895b) Hepaticarum species novae VIII. Hedwigia 34 (5): 232–253. Stephani F (1895c) Hepaticarum species novae VII. Hedwigia 34 (2): 43–65. Stephani F (1895d) Hepaticae. In: Engler A (Ed.) Die Planzenwelt Ost-Afrikas und der Nachbargebiete. heil C. Verzeichniss der bis jetzt aus Ost-Afrika bekannt gewordenen Planzen. Dietrich Reimer Verlag, Berlin, 62–66. doi: 10.5962/bhl.title.587 Stephani F (1896a) Hepaticae. In: Reinecke F (Ed.) Die Flora der Samoa Inseln. Botanische Jahrbücher für Systematik, Planzengeschichte und Planzengeographie 23 (1/2, 3): 237–368. Stephani F (1896b) Hepaticarum species novae IX. Hedwigia 35 (3): 73–140. Stephani F (1897a) Die Lebermoose der ersten Regnell’schen expedition nach Südamerika. Bihang till Kongliga Svenska Vetenskaps-Akademiens Handlingar 23 (III, 2): 1–36. Stephani F (1897b) Hepaticae japonicae. Bulletin de l’Herbier Boissier 5 (2): 76–108. Stephani F (1897c) Hepaticae sandvicenses. Bulletin de l’Herbier Boissier 5 (10): 840–849. Stephani F (1898a) Species hepaticarum 1. Bulletin de l’Herbier Boissier 6 (4): 309–343. doi: 10.5962/bhl.title.95494 800 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Stephani F (1898b) Species hepaticarum 1. Bulletin de l’Herbier Boissier 6 (5): 361–378. doi: 10.5962/bhl.title.95494 Stephani F (1898c) Species hepaticarum 1. Bulletin de l’Herbier Boissier 6 (10): 757–799. doi: 10.5962/bhl.title.95494 Stephani F (1899a) Species hepaticarum 1. Bulletin de l’Herbier Boissier 7 (2): 84–110. doi: 10.5962/bhl.title.95494 Stephani F (1899b) Species hepaticarum 1. Bulletin de l’Herbier Boissier 7 (3): 198–225. doi: 10.5962/bhl.title.95494 Stephani F (1899c) Species hepaticarum 1. Bulletin de l’Herbier Boissier 7 (5): 381–407. doi: 10.5962/bhl.title.95494 Stephani F (1899d) Species hepaticarum 1. Bulletin de l’Herbier Boissier 7 (7): 518–533. doi: 10.5962/bhl.title.95494 Stephani F (1899e) Species hepaticarum 1. Bulletin de l’Herbier Boissier 7 (9): 655–695. doi: 10.5962/bhl.title.95494 Stephani F (1899f) Species hepaticarum 1. Bulletin de l’Herbier Boissier 7 (10): 727–764. doi: 10.5962/bhl.title.95494 Stephani F (1899g) Species hepaticarum 1. Bulletin de l’Herbier Boissier 7 (12): 927–956. doi: 10.5962/bhl.title.95494 Stephani F (1899h) Hepaticae. In: Renauld F, Cardot J (Eds) Musci exotici novi vel minus cogniti. IX. Bulletin de la Société Royale de Botanique de Belgique 38 (1): 7–48. Stephani F (1900a) Species hepaticarum 1. Mémoires de l’Herbier Boissier 11: 1–46. doi: 10.5962/bhl.title.95494 Stephani F (1900b) Beiträge zur Lebermooslora Westpatagoniens und des südlichen Chiles. Bihang till Kongliga Svenska Vetenskaps-Akademiens Handlingar 26 (III, 6): 1–69. Stephani F (1900c) Species hepaticarum 1. Mémoires de l’Herbier Boissier 11: 1–49. doi: 10.5962/bhl.title.95494 Stephani F (1901a) Species hepaticarum 2. Bulletin de l’Herbier Boissier (sér. 2) 1 (5): 477–521. doi: 10.5962/bhl.title.95494 Stephani F (1901b) Lebermoose der Magellansländer, mit einer Einleitung von P. Dusén. Bihang till Kongliga Svenska Vetenskaps-Akademiens Handlingar 26 (III, 17): 1–36. Stephani F (1901c) Species hepaticarum 2. Bulletin de l’Herbier Boissier (sér. 2) 1 (10): 1022–1151. doi: 10.5962/bhl.title.95494 Stephani F (1901d) Hepatics. In: Hiern WP (Ed.) Catalogue of the African Plants collected by Dr. Friedrich Welwitsch in 1853-61. Volume II, Part. II. Cryptogamia. British Museum (Natural History), London, 310–321. Stephani F (1901e) Hepaticae. In: Brown NE (Ed.) Reports on two botanical collections made by Messrs, F. V. McConnell & J. J. Quelch at Mount Roraima in British Guiana. Transactions of the Linnean Society of London. Botany 6 (1): 1–107. Stephani F (1901f) Hepaticae novae Dussianae. In: Urban I (Ed.) Symbolae antillanae, Vol. 2. Bornträger, Leipzig, 469–472. doi: 10.5962/bhl.title.144 Stephani F (1901g) Hepaticae (in: Engler HGA, Beitrag zur lora von Afrika XXII). In: Engler A (Ed.) Beitrag zur lora von Afrika. XXII. Berichte über die botanischen Ergebnisse der Nyassa-See- und Kinga-Gebirgs-Expedition. Botanische Jahrbücher für Systematik, Planzengeschichte und Planzengeographie 30 (2): 239–445. World checklist of hornworts and liverworts 801 Stephani F (1901h) Species hepaticarum 2. Bulletin de l’Herbier Boissier (sér. 2) 1 (2): 140–177. doi: 10.5962/bhl.title.95494 Stephani F (1902a) Species hepaticarum 2. Bulletin de l’Herbier Boissier (sér. 2) 2 (8): 657–688. doi: 10.5962/bhl.title.95494 Stephani F (1902b) Hepaticae. In: Schmidt J (Ed.) Flora of Koh Chang. V. Botanisk Tidsskrift 24 (3): 241–280. Stephani F (1902c) Species hepaticarum 2. Bulletin de l’Herbier Boissier (sér. 2) 2 (1): 35–48. doi: 10.5962/bhl.title.95494 Stephani F (1902d) Species hepaticarum 2. Bulletin de l’Herbier Boissier (sér. 2) 2 (2): 157–179. doi: 10.5962/bhl.title.95494 Stephani F (1902e) Hepaticae novae Dussianae II. In: Urban I (Ed.) Symbolae antillanae, Vol. 3. Bornträger, Leipzig, 275–279. doi: 10.5962/bhl.title.144 Stephani F (1902f) Species hepaticarum 2. Bulletin de l’Herbier Boissier (sér. 2) 2 (5): 454–474. doi: 10.5962/bhl.title.95494 Stephani F (1902g) Species hepaticarum 2. Bulletin de l’Herbier Boissier (sér. 2) 2 (10): 857–888. doi: 10.5962/bhl.title.95494 Stephani F (1902h) Species hepaticarum 2. Bulletin de l’Herbier Boissier (sér. 2) 2 (12): 969–987. doi: 10.5962/bhl.title.95494 Stephani F (1903a) Species hepaticarum 2. Bulletin de l’Herbier Boissier (sér. 2) 3 (11): 959–974. doi: 10.5962/bhl.title.95494 Stephani F (1903b) Species hepaticarum 2. Bulletin de l’Herbier Boissier (sér. 2) 3 (2): 98–129. doi: 10.5962/bhl.title.95494 Stephani F (1903c) Species hepaticarum 2. Bulletin de l’Herbier Boissier (sér. 2) 3 (4): 326–341. doi: 10.5962/bhl.title.95494 Stephani F (1903d) Species hepaticarum 2. Bulletin de l’Herbier Boissier (sér. 2) 3 (6): 522–537. doi: 10.5962/bhl.title.95494 Stephani F (1903e) Species hepaticarum 2. Bulletin de l’Herbier Boissier (sér. 2) 3 (7): 596–611. doi: 10.5962/bhl.title.95494 Stephani F (1903f) Species hepaticarum 2. Bulletin de l’Herbier Boissier (sér. 2) 3 (10): 873–888. doi: 10.5962/bhl.title.95494 Stephani F (1904a) Species hepaticarum 2. Bulletin de l’Herbier Boissier (sér. 2) 4 (10): 973–988. doi: 10.5962/bhl.title.95494 Stephani F (1904b) Species hepaticarum 2. Bulletin de l’Herbier Boissier (sér. 2) 4 (6): 586–601. doi: 10.5962/bhl.title.95494 Stephani F (1904c) Species hepaticarum 2. Bulletin de l’Herbier Boissier (sér. 2) 4 (8): 775–790. doi: 10.5962/bhl.title.95494 Stephani F (1904d) Hepaticae. In: Renauld F, Cardot J (Ed.) Musci exotici novi vel minus cogniti. X. Bulletin de la Société Royale de Botanique de Belgique 41 (1): 7–122. Stephani F (1904e) Species hepaticarum 2. Bulletin de l’Herbier Boissier (sér. 2) 4 (4): 345–360. doi: 10.5962/bhl.title.95494 Stephani F (1904f) Species hepaticarum 2. Bulletin de l’Herbier Boissier (sér. 2) 4 (2): 153–168. doi: 10.5962/bhl.title.95494 Stephani F (1904g) Species hepaticarum 2. Bulletin de l’Herbier Boissier (sér. 2) 4 (1): 18–32. doi: 10.5962/bhl.title.95494 802 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Stephani F (1904h) Species hepaticarum 2. Bulletin de l’Herbier Boissier (sér. 2) 4 (12): 1197–1214. doi: 10.5962/bhl.title.95494 Stephani F (1904i) Hepaticarum species novae X. Hedwigia 44 (1): 14–15. Stephani F (1905a) Hepaticae amazonicae ab Ernesto Ule collectae. Hedwigia 44 (4): 223– 229. Stephani F (1905b) Species hepaticarum 3. Bulletin de l’Herbier Boissier (sér. 2) 5 (12): 1129–1144. doi: 10.5962/bhl.title.95494 Stephani F (1905c) Tableau des hépatiques. In: Dusén P (Ed.) Sur la lore de la Serra do Itatiaya au Brésil. Archivos do Museu Nacional do Rio de Janeiro 13: 1–119. Stephani F (1905d) Species hepaticarum 2. Bulletin de l’Herbier Boissier (sér. 2) 5 (2): 175–190. doi: 10.5962/bhl.title.95494 Stephani F (1905e) Hepaticae gesammelt von C. Skottsberg während der schwedischen Südpolarexpedition, 1901–1903. Wissenschaftliche Ergebnisse der Schwedischen Südpolar-Expedition 1901–1903 4 (1): 1–11. Stephani F (1905f) Species hepaticarum 2. Bulletin de l’Herbier Boissier (sér. 2) 5 (10): 917–946. doi: 10.5962/bhl.title.95494 Stephani F (1905g) Species hepaticarum 2. Bulletin de l’Herbier Boissier (sér. 2) 5 (9): 885–900. doi: 10.5962/bhl.title.95494 Stephani F (1905h) Hepaticae species novae XI. Hedwigia 44 (2): 72–75. Stephani F (1905i) Species hepaticarum 2. Bulletin de l’Herbier Boissier (sér. 2) 5 (8): 736–751. doi: 10.5962/bhl.title.95494 Stephani F (1905j) Species hepaticarum 2. Bulletin de l’Herbier Boissier (sér. 2) 5 (4): 351–366. doi: 10.5962/bhl.title.95494 Stephani F (1906a) Species hepaticarum 3. Bulletin de l’Herbier Boissier (sér. 2) 6 (5): 377–392. doi: 10.5962/bhl.title.95494 Stephani F (1906b) Zwei neue irländische Plagiochilen. Hedwigia 45 (4): 213–214. Stephani F (1906c) Species hepaticarum 3. Bulletin de l’Herbier Boissier (sér. 2) 6 (11): 935–966. doi: 10.5962/bhl.title.95494 Stephani F (1906d) Species hepaticarum 3. Bulletin de l’Herbier Boissier (sér. 2) 6 (10): 872–889. doi: 10.5962/bhl.title.95494 Stephani F (1906e) Species hepaticarum 3. Bulletin de l’Herbier Boissier (sér. 2) 6 (9): 781–796. doi: 10.5962/bhl.title.95494 Stephani F (1906f) Species hepaticarum 3. Bulletin de l’Herbier Boissier (sér. 2) 6 (7): 535–550. doi: 10.5962/bhl.title.95494 Stephani F (1906g) Species hepaticarum 3. Bulletin de l’Herbier Boissier (sér. 2) 6 (3): 217–232. doi: 10.5962/bhl.title.95494 Stephani F (1906h) Species hepaticarum 3. Bulletin de l’Herbier Boissier (sér. 2) 6 (8): 649–664. doi: 10.5962/bhl.title.95494 Stephani F (1907a) Hepaticae. In: Rechinger KH (Ed.) Botanische und zoologische Ergebnisse einer wissenschaftlichen Forschungsreise nach den Samoainseln. I. Teil. Denkschriften der Akademie der Wissenschaften in Wien. Mathematisch- Naturwissenschaftliche Klasse 81: 197–317. doi: 10.5962/bhl.title.12030 Stephani F (1907b) Species hepaticarum 3. Bulletin de l’Herbier Boissier (sér. 2) 7 (10): 837–852. doi: 10.5962/bhl.title.95494 World checklist of hornworts and liverworts 803 Stephani F (1907c) Species hepaticarum 3. Bulletin de l’Herbier Boissier (sér. 2) 7 (1): 59–72. doi: 10.5962/bhl.title.95494 Stephani F (1907d) Species hepaticarum 3. Bulletin de l’Herbier Boissier (sér. 2) 7 (4): 297–312. doi: 10.5962/bhl.title.95494 Stephani F (1907e) Species hepaticarum 3. Bulletin de l’Herbier Boissier (sér. 2) 7 (8): 683–698. doi: 10.5962/bhl.title.95494 Stephani F (1908a) Species hepaticarum 3. Bulletin de l’Herbier Boissier (sér. 2) 8 (11): 837–866. doi: 10.5962/bhl.title.95494 Stephani F (1908b) Species hepaticarum 3. Bulletin de l’Herbier Boissier (sér. 2) 8 (12): 941–966. doi: 10.5962/bhl.title.95494 Stephani F (1908c) Species hepaticarum 3. Bulletin de l’Herbier Boissier (sér. 2) 8 (10): 745–776. doi: 10.5962/bhl.title.95494 Stephani F (1908d) Species hepaticarum 3. Bulletin de l’Herbier Boissier (sér. 2) 8 (9): 661–696. doi: 10.5962/bhl.title.95494 Stephani F (1908e) Species hepaticarum 3. Bulletin de l’Herbier Boissier (sér. 2) 8 (8): 561–608. doi: 10.5962/bhl.title.95494 Stephani F (1908f) Species hepaticarum 3. Bulletin de l’Herbier Boissier (sér. 2) 8 (7): 483–514. doi: 10.5962/bhl.title.95494 Stephani F (1908g) Species hepaticarum 3. Bulletin de l’Herbier Boissier (sér. 2) 8 (6): 426–436. doi: 10.5962/bhl.title.95494 Stephani F (1908h) Species hepaticarum 3. Bulletin de l’Herbier Boissier (sér. 2) 8 (3): 205–220. doi: 10.5962/bhl.title.95494 Stephani F (1908i) Species hepaticarum 3. Bulletin de l’Herbier Boissier (sér. 2) 8 (2): 125–148. doi: 10.5962/bhl.title.95494 Stephani F (1908j) Species hepaticarum 3. Bulletin de l’Herbier Boissier (sér. 2) 8 (4): 267–282. doi: 10.5962/bhl.title.95494 Stephani F (1908k) Species hepaticarum 3. Bulletin de l’Herbier Boissier (sér. 2) 8 (1): 49–64. doi: 10.5962/bhl.title.95494 Stephani F (1908l) Hépatiques de la Nouvelle-caledonie et du Tonkin. Revue Bryologique 35 (2): 28–35. Stephani F (1909a) Species hepaticarum 3. George & Cie, Genève & Bale, 517–693. doi: 10.5962/bhl.title.95494 Stephani F (1909b) Hepaticae mexicanae novae. Revue Bryologique 36 (6): 138–140. Stephani F (1909c) Dendroceros, eine Gattung der Lebermoose. Sitzungsberichte der Naturforschenden Gesellschaft zu Leipzig 36: 11–20. Stephani F (1909d) Species hepaticarum 4. George & Cie, Genève & Bale, 1–96. doi: 10.5962/ bhl.title.95494 Stephani F (1910a) Hepaticae. In: Rechinger KH (Ed.) Botanische und zoologische Ergebnisse einer wissenschaftlichen Forschungsreise nach den Samoainseln. III. Teil. Denkschriften der Akademie der Wissenschaften in Wien. Mathematisch-Naturwissenschaftliche Klasse 85: 175–432. Stephani F (1910b) Species hepaticarum 4. George & Cie, Genève & Bale, 97–448. doi: 10.5962/bhl.title.95494 804 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Stephani F (1911a) Hepaticae. In: Mildbraed J (Ed.) Wissenschaftliche Ergebnisse der Deutschen Zentral-Afrika-Expedition, 1907-1908. Band II: Botanik, Lief. 2. Klinkhart & Biermann, Leipzig, 111–134. doi: 10.5962/bhl.title.7048 Stephani F (1911b) Botanische Ergebnisse der schwedischen Expedition nach Patagonien und dem Feuerlande 1907–1909. II. Die Lebermoose. Kungliga Svenska Vetenskapsakademiens Handlingar (n.ser.) 46 (9): 1–92. Stephani F (1911c) Eine neue Gattung der Hepaticae. Hedwigia 51 (1): 61–64. Stephani F (1911d) Hepaticae samoanae, II Nachtrag. In: Rechinger KH (Ed.) Botanische und zoologische Ergebnisse einer wissenschaftlichen Forschungsreise nach den Samoainseln. IV. Teil. Denkschriften der Akademie der Wissenschaften in Wien. Mathematisch-Naturwissenschaftliche Klasse 88: 1–65. Stephani F (1911e) Species hepaticarum 4. George & Cie, Genève & Bale, 449–752. doi: 10.5962/bhl.title.95494 Stephani F (1912a) Hepaticae. In: Schröder B (Ed.) Zellplanzen Ostafrikas, gesammelt auf der Akademischen Studienfahrt 1910. Hedwigia 52 (5): 288–315. Stephani F (1912b) Species hepaticarum 4. George & Cie, Genève & Bale, 753–824. doi: 10.5962/bhl.title.95494 Stephani F (1912c) Species hepaticarum 5. George & Cie, Genève & Bale, 1–176. doi: 10.5962/bhl.title.95494 Stephani F (1913a) Species hepaticarum 5. George & Cie, Genève & Bale, 177–448. doi: 10.5962/bhl.title.95494 Stephani F (1913b) Hepaticae. In: Brunnthaler J (Ed.) Ergebnisse einer botanischen Forschungsreise nach Deutsch-Ostafrika und Südafrika (Kapland, Natal und Rhodesien). I heil. Denkschriften der Kaiserlichen Akademie der Wissenschaften. Mathematisch-Naturwissenschaftliche Klasse 88: 711–744. Stephani F (1914a) Six new liverworts. Lealets of Philippine Botany 6: 2287–2290. Stephani F (1914b) Species hepaticarum 5. George & Cie, Genève & Bale, 449–704. doi: 10.5962/bhl.title.95494 Stephani F (1914c) Hepaticae. In: Sarasin F, Roux J (Eds) Nova Caledonia, Forschungen in Neu-Caledonien und auf den Loyalty-Inseln, B. Botanik 1. C. W. Kreidels Verlag, Weisbaden, 19–0. Stephani F (1915a) Nachtrag zu dem Hepaticae der Samoainseln. In: Rechinger KH (Ed.) Botanische und zoologische Ergebnisse einer wissenschaftlichen Forschungsreise nach den Samoainseln. VI. Teil. Denkschriften der Akademie der Wissenschaften in Wien. Mathematisch-Naturwissenschaftliche Klasse 91: 1–75. Stephani F (1915b) Species hepaticarum 5. George & Cie, Genève & Bale, 705–832. doi: 10.5962/bhl.title.95494 Stephani F (1916a) Hepaticae. In: Herzog T (Ed.) Die Bryophyten meiner zweiten Reise durch Bolivia. Bibliotheca Botanica 87: 1–347. Stephani F (1916b) Species hepaticarum 5. George & Cie, Genève & Bale, 833–1008. doi: 10.5962/bhl.title.95494 Stephani F (1917a) Species hepaticarum 6. George & Cie, Genève & Bale, 1–128. doi: 10.5962/bhl.title.95494 World checklist of hornworts and liverworts 805 Stephani F (1917b) Species hepaticarum 5. George & Cie, Genève & Bale, 1009–1044. doi: 10.5962/bhl.title.95494 Stephani F (1918) Species hepaticarum 6. George & Cie, Genève & Bale, 129–176. doi: 10.5962/bhl.title.95494 Stephani F (1921) Species hepaticarum 6. George & Cie, Genève & Bale, 177–240. doi: 10.5962/bhl.title.95494 Stephani F (1922) Species hepaticarum 6. George & Cie, Genève & Bale, 241–368. doi: 10.5962/bhl.title.95494 Stephani F (1923) Species hepaticarum 6. George & Cie, Genève & Bale, 369–432. doi: 10.5962/bhl.title.95494 Stephani F (1924) Species hepaticarum 6. George & Cie, Genève & Bale, 433–622. doi: 10.5962/bhl.title.95494 Stephani F, Watts WW (1914) Hepaticae australes. Journal and Proceedings of the Royal Society of New South Wales 48 (1/2): 94–135. Stotler RE (1969) he genus Frullania subgenus Frullania in Latin America. Nova Hedwigia 18: 397–555. Stotler RE, Crandall-Stotler BJ (1977) A checklist of the liverworts and hornworts of North America. Bryologist 80 (3): 405–428. doi: 10.2307/3242017 Stotler RE, Crandall-Stotler B (2005) A revised classiication of the Anthocerotophyta and a checklist of the hornworts of North America, north of Mexico. Bryologist 108 (1): 16–26. doi: 10.1639/0007-2745(2005)108[16:ARCOTA]2.0.CO;2 Stotler RE, Crotz DK (1983) On Mnium trichomanis Linnaeus (Hepatophyta). Taxon 32 (1): 64–75. doi: 10.2307/1219851 Stotler RE, Bray JR Jr, Cargill DC, Krayesky D, Crandall-Stotler BJ (2003) Typiications in the genus Fossombronia (Marchantiophyta). Bryologist 106 (1): 130–142. doi: 10.1639/0007-2745(2003)106[0130:TITGFM]2.0.CO;2 Stotler RE, Doyle WT, Crandall-Stotler BJ (2005) Phymatoceros Stotler, W. T. Doyle & Crand.-Stotl., gen. nov. (Anthocerotophyta). Phytologia 87 (2): 113–116. Stotler RE, Crandall-Stotler BJ, Bray JR Jr (2010) Fossombronia marshii (Marchantiophyta), a new liverwort species from Arkansas. Phytologia 92 (2): 230–232. Sukkharak P (2014) Studies on the genus Mastigolejeunea (Marchantiophyta: Lejeuneaceae): Mastigolejeunea gradsteinii Sukkharak sp. nov. Journal of Bryology 36 (1): 56–60. doi: 10.1179/1743282013Y.0000000082 Sukkharak P (2015) A systematic monograph of the genus hysananthus (Lejeuneaceae, Marchantiophyta). Phytotaxa 193 (1): 1–81. doi: 10.11646/phytotaxa.193.1.1 Sukkharak P, Gradstein SR (2010) Studies on the genus hysananthus (Marchantiophyta: Lejeuneaceae) 1. hysananthus discretus Sukkharak et Gradst. sp. nov. Cryptogamie, Bryologie 31 (2): 113–119. Sukkharak P, Gradstein SR (2014) A taxonomic revision of the genus Mastigolejeunea (Marchantiophyta: Lejeuneaceae). Nova Hedwigia 99 (3/4): 279–345. doi: 10.1127/00295035/2014/0206 Sukkharak P, Gradstein SR, Stech M (2011) Phylogeny, taxon circumscriptions, and character evolution in the core Ptychanthoideae (Lejeuneaceae, Marchantiophyta). Taxon 60 (6): 1607–1622. 806 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Sullivant WS (1849) Contributions to the bryology and hepaticology of North America, 2. Memoirs of the American Academy of Arts and Sciences (n.ser.) 4: 169–176. Sullivant WS (1850) Notices of several new species of mosses and hepaticae from Tierra del Fuego. Hooker’s Journal of Botany and Kew Gardens Miscellany 2: 315–318. Sullivant WS (1856) he musci and hepaticae of the United States east of the Mississippi river. In: Gray A (Ed.) Manual of the Botany of the northern United States, second edition. George P. Putnam & Co., New York, 607–737. doi: 10.5962/bhl.title.50405 Sun J, Duan Y-X (2007) he rectiication of three illegal names in the genus Jungermannia (Hepaticae, Jungermanniaceae). Bulletin of Botanical Research. Harbin 27 (2): 139–140. Sun J, Cao T, Gao C, Yu J (2004) A new species of Scapania (Scapaniaceae, Hepaticae) from Sichuan, China. Guihaia 24 (1): 23–24. Svihla RD (1957) Frullaniaceae of Burma. II. Frullania shanensis n. sp. Bryologist 60 (4): 359–363. doi: 10.2307/3239883 Svihla RD (1958) Frullaniaceae of Burma IV. Frullania maymyoensis n. sp. Bryologist 61 (4): 376–379. doi: 10.2307/3240172 Swails Jr LF (1970) he genus Porella in Latin America. Nova Hedwigia 19: 201–291. Swartz O (1781) Methodus muscorum illustrata. Joh. Edman, Upsaliae [Uppsala], 38 pp. Swartz O (1788) Nova genera et species plantarum prodromus. Bibliopolio. I.G. Mülleriano, Lipsiae [Leipzig], 152 pp. Swartz O (1806) Flora indiae occidentalis, tomus 3. J. Palm, Erlangae, 1231–2018. doi: 10.5962/bhl.title.434 Sydow P (1894) Moose. Just’s botanischer Jahresbericht 19: 223–249. Szweykowski J, Buczkowska K, Odrzykoski IJ (2005) Conocephalum salebrosum (Marchantiopsida, Conocephalaceae) – a new holarctic liverwort species. Plant Systematics and Evolution 253 (1/4): 133–158. doi: 10.1007/s00606-005-0301-0 Szyszylowicz I (1894) Diagnoses plantarum novarum a cl. d. Const. Jelski in Peruvia lectarum, pars prima. Sumptibus Academiae, Cracoviae, 1–25. Tan BC, Engel JJ (1986) An annotated checklist of Philippine Hepaticae. Journal of the Hattori Botanical Laboratory 60: 283–355. Taylor T (1836a) Flora hibernica. Part second, comprising the musci, hepaticae and lichenes. William Curry Jun and Co., Dublin, 279 pp. doi: 10.5962/bhl.title.6699 Taylor T (1836b) De Marchantieis. Transactions of the Linnean Society of London 17 (3): 375–395. doi: 10.1111/j.1095-8339.1834.tb00030.x Taylor T (1843) Descriptions of two species of British Jungermanniae. Annals and Magazine of Natural History 12 (76): 172–173. doi: 10.1080/03745484309442506 Taylor T (1844a) Descriptions of Jungermannia ulicina, (Taylor), and of J. Lyoni, (Taylor). Transactions of the Botanical Society of Edinburgh 1 (1/4): 115–116. doi: 10.1080/03746604409467519 Taylor T (1844b) On two new species of Jungermannia, and another new to Britain. Transactions of the Botanical Society of Edinburgh 1: 179–181. doi: 10.1080/03746604409467527 Taylor T (1846a) New hepaticae. London Journal of Botany 5: 258–284. Taylor T (1846b) New hepaticae. London Journal of Botany 5: 365–417. World checklist of hornworts and liverworts 807 Taylor T (1847a) Diagnostic characters of ive new species of cryptogamic plants from Jamaica. Annals and Magazine of Natural History 20 (135): 379–381. doi: 10.1080/037454809496074 Taylor T (1847b) Descriptions of new musci and hepaticae, collected by Professor William Jameson on Pichincha, near Quito. London Journal of Botany 6: 328–342. Taylor T (1848a) On some new musci, collected by professor W. Jameson on Pichincha. London Journal of Botany 7: 187–199. Taylor T (1848b) On the speciic characters of certain new cryptogamic plants, lately received from, and collected by, professor William Jameson, on Pichincha, near Quito. London Journal of Botany 7: 278–285. Taylor J (1954) A new Australian hepatic. Kew Bulletin 9 (1): 45–47. doi: 10.2307/4108927 Taylor T, Hooker JD (1845) XXXIV Hepaticae, Juss. In: Hooker JD (Ed.) he Botany of the Antarctic Voyage of H. M. discovery ships Erebus and Terror in the years 1839-43. I. Flora Antarctica I. Lord Auckland’s Group and Campbell’s Island. Reeve, London, 144–169. haithong O, Hattori S (1977) A new species of Frullania from North Vietnam and the Indian F. physantha Mitt. Bulletin of the National Science Museum, Tokyo. Series B, Botany 3 (4): 149–154. hiers BM (1983) Type studies in the Lejeuneaceae. II. Pteryganthus, a new subgenus of Lopholejeunea. Brittonia 35 (1): 81–86. doi: 10.2307/2806055 hiers BM (1984) Studies in Lejeuneaceae, III. Lopholejeunea erugata, a new name for Ptychocoleus inermis. Brittonia 36 (2): 174–177. doi: 10.2307/2806626 hiers BM (1985) Austrolejeunea bidentata, a new species of Lejeuneaceae subfamily Tuyamaelloideae from Australia. Bryologist 88 (4): 350–352. doi: 10.2307/3242671 Thiers BM (1987a) Lepidolejeunea queenslandica (Lejeuneaceae subfamily Lejeuneoideae): a new species from Australia. Memoirs of the New York Botanical Garden 45: 556–560. hiers BM (1987b) A preliminary account of Colura (Hepaticae, Lejeuneaceae) in Australia. Brittonia 39 (2): 175–179. doi: 10.2307/2807369 hiers BM (1988) he Australian species of Cololejeunea. Beihefte zur Nova Hedwigia 90: 113–146. hiers BM (1992a) New species of Cheilolejeunea and Otolejeunea (Hepaticae, Lejeuneaceae) from Australia. Brittonia 44 (2): 160–165. doi: 10.2307/2806830 hiers BM (1992b) A re-evaluation of Cheilolejeunea subgenus Xenolejeunea. Tropical Bryology 5: 11–21. hiers BM (1993) A monograph of Pleurozia (Hepaticae, Pleuroziaceae). Bryologist 96 (4): 517–554. doi: 10.2307/3243984 hiers BM (1997a) Cheilolejeunea in Australia: description of new taxa and key. Journal of the Hattori Botanical Laboratory 82: 321–328. hiers BM (1997b) Lejeunea bischlerae, a new species of Lejeunea subgenus Microlejeunea from Australia. Cryptogamie: Bryologie, Lichénologie 18 (3): 223–226. hiers BM, Gradstein SR (1989) Lejeuneaceae (Hepaticae) of Australia. I. Subfamily Ptychanthoideae. Memoirs of the New York Botanical Garden 52: 1–79. 808 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) hiers BM, Söderström L, Hagborg A, von Konrat MJ (2012) Notes on Early Land Plants Today. 11. Microlejeunea bischlerae (B.M.hiers) comb. nov. Phytotaxa 65: 59. houvenot L, Reeb C (2014) Riccardia elisabethae, a new species of Aneuraceae (Marchantiophyta) from New Caledonia. Telopea 17: 229–232. doi: 10.7751/telopea20147824 houvenot L, Gradstein SR, Hagborg A, Söderström L, Bardat J (2011) Checklist of the liverworts and hornworts of New Caledonia. Cryptogamie, Bryologie 32 (4): 287–390. doi: 10.7872/cryb.v32.iss4.2011.287 hunberg CP (1784) Flora iaponica. J. G. Müller, Leipzig, 418 pp. hunberg CP (1800) Prodromus plantarum capensium, pars posterior. Joh. Edman, Upsaliae [Uppsala], 85–191. hurn EF (1886) Notes of the plants observed during the Roraima expedition of 1884. Timehri 5: 145–223. Tindall EM (1898) Fossombronia mittenii n. sp. Journal of Botany, British and Foreign 36: 44–45. Tixier P (1962) A propos du genre Tuyamaella Hatt. dans le Sud-est asiatique: Tuyamaella hattorii n. sp. Revue Bryologique et Lichénologique 31 (3/4): 187–189. Tixier P (1967) Bryophytae indosinicae. A contribution to the knowledge of bryophytes in western Indochina. Dacca University Studies. Part B, Sciences 15: 1–14. Tixier P (1969) Cololejeunea de l’Asie du sud-est. I. –Leonidentes et espèces aines. Revue Bryologique et Lichénologique 36 (3/4): 543–594. Tixier P (1970a) Bryophytae Indosinicae. Bryophytes from hai Tenasserim. Natural History Bulletin of the Siam Society 23 (4): 541–559. Tixier P (1970b) Contribution à la connaissance du genre Cololejeunea en Asie du Sud-Est. II. La section Radulae (nov. sect.) du sous-genre Lasiolejeunea. Annales de la Faculté des Sciences, Université de Phnom Penh 3: 177–190. Tixier P (1971) A Contribution to the Bryological Knowledge of Fraser’s Hill station (Malaysia). Gardens’ Bulletin, Singapore 25 (3): 335–353. Tixier P (1972a) Contribution à l’étude des Lejeunéacées du Paciique. Bulletin de la Société d’Histoire Naturelle de l’Afrique du Nord 63: 9–12. Tixier P (1972b) Mount Maquiling bryolora (Luzon). Gardens’ Bulletin, Singapore 26 (1): 137–153. Tixier P (1973a) Le genre Tuyamaella Hatt. (Lejeunéacées) Monographie. Revue Bryologique et Lichénologique 39 (2): 221–244. Tixier P (1973b) Contribution to the knowledge of genus Cololejeunea in south-east Asia. III. Some new species. Natural History Bulletin of the Siam Society 24 (3/4): 439–447. Tixier P (1973c) Jovetastella (Lejeuneaceae, Paradoxae) genre nouveau. Revue Bryologique et Lichénologique 39 (4): 661–663. Tixier P (1973d) Bryophytes exotiques. Bulletin du Muséum National d’Histoire Naturelle (sér. 3), Botanique 10 (190): 73–86. Tixier P (1974) Bryophytae Indosinicae. À propos de quelques espèces nouvelles récoltées au Vietnam. Annales Historico-Naturales Musei Nationalis Hungarici 66: 87–100. Tixier P (1975a) Contribution à l’étude du genre Cololejeunea V. Quelques espèces de la region indopaciique. Botaniska Notiser 128: 425–431. World checklist of hornworts and liverworts 809 Tixier P (1975b) Contribution à l’étude de l’hépaticologie africaine. I. Récoltes en bordure du Golfe de Guinée (Cameroun et Gabon). Annales de la Faculté des Sciences de Yaoundé 20: 3–10. Tixier P (1977a) La famille Cololejeunoideae (Grolle) dans l’Ocean Indien Occidental – Essai Monographique. Bulletin de l’Académie Malgache (n.ser.) 55 (1/2): 173–247. Tixier P (1977b) Espèces nouvelles malgaches du genre Diplasiolejeunea (Spruce) Schifn. (Hepaticae). Lindbergia 4 (1/2): 117–125. Tixier P (1979a) Bryogéographie du Mont Bokor (Cambodge) (Bryophyta indosinica XXIV). Bryophytorum Bibliotheca 18: 5–121. Tixier P (1979b) Contribution à l’étude du genre Cololejeunea. Les Cololejeunoidées de Nouvelle Calédonie. Nova Hedwigia 31: 721–787. Tixier P (1979c) Nouvelles espèces malgaches de Diplasiolejeunea (Diplasiae) II. Revue Bryologique et Lichénologique 45 (2): 209–226. Tixier P (1979d) Contribution to the knowledge of the genus Cololejeunea VIII. Some new species of Malagasy Cololejeunea. Bryologist 82 (4): 602–608. doi: 10.2307/3242002 Tixier P (1980a) Contribution à l’étude du genre Cololejeunea (Lejeuneaceae) IX. Espèces nouvelles du sous genre Pedinolejeunea (Ben.) Mizutani en région néotropicale. Bradea 3 (6): 35–44. Tixier P (1980b) Deux nouveaux genres de Lejeunéacées, Otolejeunea Grolle et P. Tx. et Allorgella P. Tx. Nova Hedwigia 32: 607–622. Tixier P (1981) La notion d’espèce chez le genre Cololejeunea. La complexe Cololejeunea loccosa (Lehm. & Lindenb.) Schifn. Cryptogamie: Bryologie, Lichénologie 2 (1): 47–76. Tixier P (1982) Une nouvelle espèce pour le genre Jovetastella: J. aurantia. Cryptogamie: Bryologie, Lichénologie 3 (1): 29–31. Tixier P (1983a) La notion d’espèce dans le genre Diplasiolejeunea. 2. Diplasiolejeunea pauckertii (Nees) Steph. et D. columbica sp. nov. Lejeunéacées corticoles des “paramos” andins. Cryptogamie: Bryologie, Lichénologie 4 (3): 231–236. Tixier P (1983b) Contributions to the knowledge of Paciic Lejeuneaceae. II. Cheilolejeunea (Xenolejeunea) huerlimannii sp. nov. Miscellanea Bryologica et Lichenologica 9 (9): 184–185. Tixier P (1984) Contribution à l’étude du genre Diplasiolejeunea (Spruce) Schifn. 4. La Section Villaumeae sur la côte est de Madagascar. Acta Botanica Hungarica 30 (1/2): 11–26. Tixier P (1985a) Contribution à la connaissance des Cololejeunoideae. Bryophytorum Bibliotheca 27: 1–439. Tixier P (1985b) A propos du genre Bazzania en Nouvelle-Calédonie récoltes de H. S. Mac Kee. Cryptogamie: Bryologie, Lichénologie 6 (2): 177–180. Tixier P (1988) Le domaine lémuro-australasien intérêt biogéographique de deux espèces nouvelles. Nova Hedwigia 46 (3/4): 373–383. Tixier P (1989) Contribution à l’étude du genre Cololejeunea (Lejeuneaceae, Hepaticae) X. Espèces nouvelles récoltées par Sten Bergman en Irian Jaya (1949). Bulletin du Jardin Botanique National de Belgique 59 (3/4): 439–444. doi: 10.2307/3668358 Tixier P (1991) Bryophyta exotica – 9. Quelques Lejeuneaceae (Hépatiques) nouvelles pour l’Amerique du Sud. Candollea 46 (2): 267–302. 810 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Tixier P (1993) Contribution à la genre Cololejeunea (Lejeuneaceae, Hepaticae) XI. De quelques Pedinolejeunea austreaux. Cryptogamie: Bryologie, Lichénologie 14 (3): 353–360. Tixier P (1995a) Rectiications nomenclaturales – Typiications. Cryptogamie: Bryologie, Lichénologie 16 (3): 229–230. Tixier P (1995b) Résultats taxonomiques de l’expédition BRYOTROP au Zaire et Rwanda. 30. Bryophytes épiphylles (récoltes de E. Fischer). Tropical Bryology 11: 11–76. Trabut ML (1886) Riella battandieri sp. nov. Revue Bryologique 13 (3): 35. Trabut ML (1891) Revision des espèces du genre Riella et description d’une espèce nouvelle. Revue Générale de Botanique 3 (35): 449–459. Trabut ML (1908) Une nouvelle Riella d’Algerie (Riella bialata). Revue Bryologique 35 (4): 96. Trabut ML (1916) Observations sur deux Ricciacées de la lore d’Algerie. Bulletin de la Société d’Histoire Naturelle de l’Afrique du Nord 7: 87. Trabut ML (1934) Hépatiques Nord-Africaines inédites. Bulletin de la Société d’Histoire Naturelle de l’Afrique du Nord 25 (9): 391–393. Trelease W (1897) Botanical observations on the Azores. Annual Report of the Missouri Botanical Garden 8: 77–220. doi: 10.2307/2992160 Trevisan VBA (1853) Herbarium cryptogamicum trevisianum. Num. II (Musci, Hepaticae). Imprimerie Bianchi, Padova, 16 pp. (non vidi) Trevisan VBA (1874) Nuova censo delle epatiche italiane. Rendiconti dell Reale Istituto Lombardo di Scienze e Lettere (ser. 2) 7: 776–786. Trevisan VBA (1877) Schema di una nuova classiicazione delle Epatiche. Memorie del Reale Istituto Lombardo de Scienze e Lettere (Serie 3), Classe di Scienze Matematiche e Naturali 4 (13): 383–451. Tsudo S, Hasegawa J (2006) A new species of Frullania (Frullaniaceae, Hepaticae) found in Miyazaki, southern Japan. Bryological Research 9 (3): 43–46. Tuba Z, Slack NG, Stark LR (2011) Bryophyte ecology and climate change. Cambridge University Press, Cambridge, 528 pp. Udar R (1959) Genus Riccia in India – IV. A new Riccia, Riccia pandei Udar sp. nov. from Garhwal with a note on the species of the genus from the West Himalayan territory. Journal of the Indian Botanical Society 38 (1): 146–159. Udar R (1961) Genus Riccia in India – V. A new Riccia, R. reticulatula Udar, sp. nov., from Pilani with a note on the species of Riccia from the central India zone, Gangetic plains, Panjab and Rajasthan. Bulletin of the Botanical Society of the University of Saugar 13: 46–55. Udar R (1965) Riccia grollei Udar nom. nov. a correction for Riccia tuberculata Pandé et Udar from India. Current Science 34 (4): 126. Udar R, Asthana AK (1985a) A new Anthoceros from Nagaland. Journal of the Indian Botanical Society 64: 303–305. Udar R, Asthana AK (1985b) Anthoceros bharadwajii – a new species from India. Proceedings of the Indian National Science Academy. Part B, Biological Sciences 51 (4): 483–489. Udar R, Awasthi US (1979) A new species of Leptolejeunea from India. Miscellanea Bryologica et Lichenologica 8 (6): 115–117. World checklist of hornworts and liverworts 811 Udar R, Awasthi US (1981) A new species of Lejeunea from India. Cryptogamie: Bryologie, Lichénologie 2 (3): 345–348. Udar R, Chandra S (1965) On two new species of Mannia, M. foreaui Udar et Chandra and M. perssonii Udar et Chandra, with a note on the genus and its Indian. Canadian Journal of Botany 43 (1): 147–160. doi: 10.1139/b65-017 Udar R, Chandra S (1977) A new species of Notothylas Sull., N. pandei Udar et Chandra from India. Geophytology 7 (2): 142–146. Udar R, Gupta A (1983) Diferentiation of the genus Targionia L. in India-II. he East Himalayan and South Indian complex and description of a new species of Targionia. Geophytology 13 (1): 83–87. Udar R, Gupta A (1984) A new Riccia (Mich.) L. from Deoban western Himalayas, India. In: NN (Ed.) Proceedings of the V Indian Geophytological Conference, Lucknow 14–16 November, 1983, Special Publication. Paleobotanical Society, Lucknow, 307–311. Udar R, Kumar D (1976) Genus Cephalozia in Eastern Himalayas. Geophytology 6 (1): 35–45. Udar R, Kumar A (1982a) A new Chonecolea from India. Bryologist 85 (3): 315–318. doi: 10.2307/3243051 Udar R, Kumar D (1982b) he genus Radula in India – I. Journal of the Indian Botanical Society 61: 177–182. Udar R, Kumar A (1982c) A remarkable Cylindrocolea Schust. from India. Lindbergia 8 (3): 181–184. Udar R, Kumar A (1982d) Two new species of Cephaloziella from India. Lindbergia 8 (1): 30–34. Udar R, Kumar A (1983a) A new Frullania from India. Miscellanea Bryologica et Lichenologica 9 (9): 192–194. Udar R, Kumar D (1983b) Radula pandei, a new hepatic from South India. Lindbergia 9 (2): 133–136. Udar R, Nath V (1976) A new species of Cephaloziella Spruce, C. magna Udar et Nath. sp. nov. from Sheetlakhet, Almora (western Himalayas) India. Geophytology 6 (1): 105–107. Udar R, Nath V (1978) A new species of Jubula Dum. from India. Miscellanea Bryologica et Lichenologica 8 (3): 49–52. Udar R, Nath V (1981) Frullania hattoriantha sp. nov. from India. Miscellanea Bryologica et Lichenologica 9 (2): 44–47. Udar R, Shaheen F (1982) A new Porella, P. chinensis (St.) Hatt. var. crispata, var. nov. from Himalayas. Miscellanea Bryologica et Lichenologica 9 (4): 74–76. Udar R, Shaheen F (1983a) Porella chinensis (St.) Hatt. var. hattori var. nov. from Himalayas. Miscellanea Bryologica et Lichenologica 9 (7): 146–148. Udar R, Shaheen F (1983b) Porella hattorii sp. nov. from India. Lindbergia 9 (1): 70–72. Udar R, Singh DK (1976) A new Cyathodium from India. Bryologist 79 (2): 234–238. doi: 10.2307/3241921 Udar R, Singh DK (1978) Cyathodium indicum Udar et Singh, sp. nov. from western Himalayas India. Journal of Bryology 10 (2): 139–142. doi: 10.1179/jbr.1978.10.2.139 Udar R, Singh DK (1979a) Notothylas pleidereri sp. nov. from Mangalore, India. Lindbergia 5 (1): 28–30. 812 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Udar R, Singh DK (1979b) On a new species of Notothylas Sull. from Western Ghats, India (1). Revue Bryologique et Lichénologique 45 (2): 201–208. Udar R, Singh DK (1980) An interesting Notothylas from Brasil. Miscellanea Bryologica et Lichenologica 8 (9): 173–178. Udar R, Singh DK (1981a) A new species of the genus Notothylas Sull., N. himalayensis Udar et Singh, from India. Journal of Bryology 11 (3): 451–457. doi: 10.1179/jbr.1981.11.3.451 Udar R, Singh DK (1981b) Notothylas khasiana Udar et Singh sp. nov. from Shillong, India. Journal of the Indian Botanical Society 60: 112–117. Udar R, Singh DK (1981c) Some new combinations under Phaeoceros Proskauer. Geophytology 11 (2): 257–258. Udar R, Srivastava SC (1971) A new species of Cyathodium Kunze, C. denticulatum Udar et Srivastava sp. nov., from Darjeeling (Eastern Himalayas), India. Geophytology 1 (2): 165–169. Udar R, Srivastava SC (1973) On a species of Riccardia, R. santapaui Udar et Srivastava from Chindwara (Madhya Pradesh), India. Revue Bryologique et Lichénologique 39 (1): 155–161. Udar R, Srivastava G (1983) A new Cololejeunea from India. Miscellanea Bryologica et Lichenologica 9 (7): 137–139. Underwood LM (1884) Descriptive catalogue of the North American hepaticae, north of Mexico. Bulletin of the Illinois State Laboratory of Natural History 2 (1): 1–133. Underwood LM (1888) Some undescribed hepaticae from California. Botanical Gazette 13 (5): 112–114. doi: 10.1086/326267 Underwood LM (1890) A new North American Lejeunea. Bulletin of the Torrey Botanical Club 17 (10): 258–259. doi: 10.2307/2476586 Underwood LM (1891) A preliminary list of Paciic Coast hepaticae. Zoe 1 (12): 361–367. Underwood LM (1894) Notes on our hepaticae II. he genus Riccia. Botanical Gazette 19 (7): 273–278. doi: 10.1086/327066 Underwood LM (1895) Notes on our hepaticae III. he distribution of North American Marchantiaceae. Botanical Gazette 20 (2): 59–71. doi: 10.1086/327151 Uribe J (2004) Type studies on Frullania subgenus Meteoriopsis (Hepaticae). IV. A new species from the Galápagos Islands. Cryptogamie, Bryologie 25 (4): 295–299. Uribe J (2006) Type studies on Frullania subgenus Meteoriopsis (Hepaticae). V. Frullania dulimensis sp. nov. from Colombia. Cryptogamie, Bryologie 27 (3): 309–312. Uribe J, Linares EL (2015) Micropterygium longicellulatum (Lepidoziaceae, Marchantiophyta), a new species from Colombia. Phytotaxa 213 (3): 296–299. doi: 10.11646/ phytotaxa.213.3.10 Urmi E (1983) Tetralophozia iliformis (Steph.) comb. nov. in Europe. Journal of Bryology 12 (3): 393–401. doi: 10.1179/jbr.1983.12.3.393 Vainio E (1878) Itä-Hämeen kasvistosta [Florula tavastiae orientalis]. Meddelanden af Societas pro Fauna et Flora Fennica 3: 1–121. Van der Gronde K (1980) Studies on Colombian cryptogams. VIII. he genus Jensenia Lindb. (Hepaticae). Proceedings, Koninklijke Nederlandse Akademie van Wetenschappen. Series C, biological and medical sciences 83 (3): 271–278. World checklist of hornworts and liverworts 813 van Slageren M (1985) A taxonomic monograph of the genera Brachiolejeunea and Frullanoides. Mededelingen van het Botanisch Museum en Herbarium van de Rijks Universiteit te Utrecht 544: 7–205. van Slageren M, Kruijt C (1985) A review of the genus Blepharolejeunea S. Arn. Beihefte zur Nova Hedwigia 80: 113–154. Váňa J (1970a) Jungermannia caucasica sp. n. (Hepaticae). Preslia 42: 96–97. Váňa J (1970b) Kymatocalyx in Mittelamerika. Österreichische Botanische Zeitschrift 118 (5): 572–579. doi: 10.1007/BF01376264 Váňa J (1970c) Eine neue Lebermoosart aus Neukaledonien. Österreichische Botanische Zeitschrift 118 (3): 233–236. doi: 10.1007/BF01377860 Váňa J (1972a) Miscellaneous notes on the Asiatic Jungermannioideae. Journal of the Hattori Botanical Laboratory 35: 312–318. Váňa J (1972b) Miscellaneous notes on the Asiatic Jungermannioideae II. Journal of the Hattori Botanical Laboratory 36: 57–74. Váňa J (1973a) Lebermoose aus Neuguinea. 10. Jungermannia. Journal of the Hattori Botanical Laboratory 37: 185–190. Váňa J (1973b) Studien über die Jungermannioideae (Hepaticae). 1. Allgemeine Charakteristik. Folia Geobotanica et Phytotaxonomica 8 (2): 181–208. doi: 10.1007/BF02854563 Váňa J (1973c) Studien über die Jungermannioideae (Hepaticae). 2. Jungermannia subg. Jungermannia. Folia Geobotanica et Phytotaxonomica 8 (3): 255–309. doi: 10.1007/ BF02852828 Váňa J (1974a) Studien über die Jungermannioideae (Hepaticae). 5. Jungermannia subg. Plectocolea und subg. Solenostoma: Afrikanische Arten. Folia Geobotanica et Phytotaxonomica 9 (3): 277–312. doi: 10.1007/BF02853150 Váňa J (1974b) Lebermoose aus Neuguinea. 11. Andrewsianthus. Journal of the Hattori Botanical Laboratory 38: 639–649. Váňa J (1974c) Studien über die Jungermannioideae (Hepaticae). 4. Jungermannia subg. Plectocolea und subg. Solenostoma: Allgemeines, süd- und mittelamerikanische Arten. Folia Geobotanica et Phytotaxonomica 9 (2): 179–208. doi: 10.1007/BF02852200 Váňa J (1975a) Miscellaneous notes on the Asiatic Jungermannioideae IV. Journal of the Hattori Botanical Laboratory 39: 211–214. Váňa J (1975b) Studien über die Jungermannioideae (Hepaticae). 9. Jungermannia subg. Plectocolea und subg. Solenostoma in Hawaii: Ergänzungen und Synopsis der Gattung Jungermannia. Folia Geobotanica et Phytotaxonomica 10 (4): 357–382. doi: 10.1007/ BF02854974 Váňa J (1976a) Lebermoose aus Neuguinea. 13. Gymnomitriaceae. Journal of the Hattori Botanical Laboratory 40: 185–189. Váňa J (1976b) Drei neue Gymnomitriaceen aus Südamerika. Journal of the Hattori Botanical Laboratory 41: 411–417. Váňa J (1976c) Studien über die Jungermannioideae (Hepaticae). 10. Nardia. Folia Geobotanica et Phytotaxonomica 11 (4): 367–425. doi: 10.1007/BF02853183 Váňa J (1977) Jungermannia (Solenostoma) mamatkulovii – eine neue Lebermoose aus Tadshikistan (UdSSR). Preslia 49: 181–182. 814 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Váňa J (1980) Some new South and Central American hepatics. Journal of the Hattori Botanical Laboratory 48: 225–234. Váňa J (1988) Cephalozia (Dum.) Dum. in Africa, with notes on the genus (Notes on some African hepatic genera 10). Beihefte zur Nova Hedwigia 90: 179–198. Váňa J (1992) A new species of Cephaloziella (Bryophyta) from the Himalayas. Folia Geobotanica et Phytotaxonomica 27 (2): 193–195. doi: 10.1007/BF02856254 Váňa J (1993) Taxonomic results of the BRYOTROP expedition to Zaire and Rwanda. 11. Cephaloziaceae, Cephaloziellaceae, Gymnomitriaceae, Jungermanniaceae, Lophoziaceae. Tropical Bryology 8: 99–103. Váňa J (2003) Notes on Gymnomitriaceae (subf. Gymnomitrioidae) in Latin America. Acta Academiae Paedagogicae Agriensis, Sectio Biologiae 24: 109–128. Váňa J (2013) Pseudoisotachis pocsii Váňa, a new genus and species of liverwort from the subantarctic Marion Island. Polish Botanical Journal 58 (1): 55–58. doi: 10.2478/pbj-2013-0006 Váňa J, Engel JJ (2013) he liverworts and hornworts of the Tristan da Cunha group of islands in the south Atlantic Ocean. Memoirs of the New York Botanical Garden 105: 1–138. Váňa J, Gremmen N (2005) Hepatics of Heard Island. Cryptogamie, Bryologie 26 (1): 79–90. Váňa J, Long DG (2008) Hamatostrepta concinna gen. et sp. nov. (Jungermanniopsida, Scapaniaceae), a new Asiatic leafy liverwort from the Sino-Burmese border. Fieldiana: Botany (n.ser.) 47: 133–138. doi: 10.3158/0015-0746-47.1.133 Váňa J, Long DG (2009) Jungermanniaceae of the Sino-Himalayan region. Nova Hedwigia 89 (3/4): 485–517. doi: 10.1127/0029-5035/2009/0089-0485 Váňa J, Müller F (2003) Cephaloziella biokoensis sp. nov. (Marchantiopsida, Cephaloziellaceae), from the island of Bioko (Equatorial Guinea). Tropical Bryology 24: 1–4. Váňa J, Piippo S (1989) Bryophyte lora of the Huon Peninsula, Papua New Guinea. XXXI. Cephaloziaceae subfam. Allobielloideae, Cephaloziellaceae, Antheliaceae and Lophoziaceae (Hepaticae). Annales Botanici Fennici 26 (3): 263–290. Váňa J, Wigginton MJ (2008) Taxonomic results of the BRYOTROP expedition to Zaire and Rwanda. 34. A new species of Amphicephalozia, A. africana sp. nov. Journal of Bryology 30 (1): 55–58. doi: 10.1179/174328208X281996 Váňa J, Bednarek-Ochyra H, Cykowska B (2009) Two new species of liverworts from the subantarctic Prince Edward Islands. Nova Hedwigia 89 (1/2): 121–129. doi: 10.1127/00295035/2009/0089-0121 Váňa J, Hentschel J, Heinrichs J (2010a) New combinations in Jungermanniales: transfer of 32 taxa to Solenostoma Mitt. Cryptogamie, Bryologie 31 (2): 135–139. Váňa J, Söderström L, Hagborg A, von Konrat MJ, Engel JJ (2010b) Early Land Plants Today: Taxonomy, systematics and nomenclature of Gymnomitriaceae. Phytotaxa 11: 1–80. doi: 10.11646/phytotaxa.11.1 Váňa J, Söderström L, Hagborg A, von Konrat MJ (2010c) Nomenclatural novelties and lectotypiications in Gymnomitriaceae. Novon 20 (2): 225–227. doi: 10.3417/2009082 Váňa J, Hentschel J, Müller J, Heinrichs J (2012a) Taxonomic novelties in Scapania. PhytoKeys 10: 13–17. doi: 10.3897/phytokeys.10.2654 Váňa J, Söderström L, Hagborg A, von Konrat MJ (2012b) Notes on Early Land Plants Today. 8. New combinations and some lectotypiications in Mesoptychia. Phytotaxa 65: 52–56. World checklist of hornworts and liverworts 815 Váňa J, Söderström L, Hagborg A, von Konrat MJ (2012c) Notes on Early Land Plants Today. 4. Aponardia gen. et stat. nov. Phytotaxa 65: 46. Váňa J, Söderström L, Hagborg A, von Konrat MJ (2012d) Notes on Early Land Plants Today. 2. Two new combinations in Solenostoma. Phytotaxa 65: 44. Váňa J, Grolle R, Long DG (2012e) Taxonomic realignments and new records of Gongylanthus and Southbya (Marchantiophyta: Southbyaceae) from the Sino-Himalayan region. Nova Hedwigia 95 (1/2): 183–196. doi: 10.1127/0029-5035/2012/0042 Váňa J, Söderström L, Hagborg A, von Konrat MJ (2012f) Notes on Early Land Plants Today. 12. Pseudomarsupidium borneensis (Grolle) comb. nov. Phytotaxa 65: 60. Váňa J, Söderström L, Hagborg A, von Konrat MJ (2013a) Notes on Early Land Plants Today. 33. Notes on Anastrophyllaceae (Marchantiophyta). Phytotaxa 81 (1): 26–32. doi: 10.11646/phytotaxa.81.1.9 Váňa J, Söderström L, Hagborg A, von Konrat MJ (2013b) Notes on Early Land Plants Today. 20. New synonyms in Gymnocoleopsis (Cephaloziellaceae, Marchantiophyta). Phytotaxa 76 (3): 41–42. doi: 10.11646/phytotaxa.76.3.8 Váňa J, Söderström L, Hagborg A, von Konrat MJ (2013c) Notes on Early Land Plants Today. 44. Comments on sexuality in Solenostoma (Solenostomataceae, Marchantiophyta) and on some newly described taxa. Phytotaxa 152 (1): 33–47. doi: 10.11646/phytotaxa.152.1.3 Váňa J, Söderström L, Hagborg A, von Konrat MJ (2013d) Notes on Early Land Plants Today. 18. Transfers of some taxa in Nardia (Gymnomitriaceae, Marchantiophyta). Phytotaxa 76 (3): 37–38. doi: 10.11646/phytotaxa.76.3.6 Váňa J, Söderström L, Hagborg A, von Konrat MJ (2013e) Notes on Early Land Plants Today. 40. Notes on Cephaloziellaceae (Marchantiophyta). Phytotaxa 112 (1): 1–6. doi: 10.11646/phytotaxa.112.1.1 Váňa J, Söderström L, Hagborg A, von Konrat MJ (2013f) Notes on Early Land Plants Today. 45. A correction. Phytotaxa 152 (1): 48–49. doi: 10.11646/phytotaxa.152.1.4 Váňa J, Söderström L, Hagborg A, von Konrat MJ (2013g) Notes on Early Land Plants Today. 41. New combinations and new synonyms in Cephaloziaceae (Marchantiophyta). Phytotaxa 112 (1): 7–15. doi: 10.11646/phytotaxa.112.1.2 Váňa J, Söderström L, Hagborg A, von Konrat MJ (2013h) Notes on Early Land Plants Today. 17. Transfers of some taxa in Adelanthaceae (Marchantiophyta). Phytotaxa 76 (3): 35–36. doi: 10.11646/phytotaxa.76.3.5 Váňa J, Söderström L, Hagborg A, von Konrat MJ (2013i) Notes on Early Land Plants Today. 16. Validation of Metahygrobiella subg. Apohygrobiella (Cephaloziaceae, Marchantiophyta). Phytotaxa 76 (3): 34. doi: 10.11646/phytotaxa.76.3.4 Váňa J, Söderström L, Hagborg A, von Konrat MJ (2013j) Notes on Early Land Plants Today. 13. New names and synonyms in Diplophyllum (Scapaniaceae, Marchantiophyta). Phytotaxa 76 (3): 28–30. doi: 10.11646/phytotaxa.76.3.1 Váňa J, Söderström L, Hagborg A, von Konrat MJ (2013k) Notes on Early Land Plants Today. 30. Transfer of some taxa from Anastrophyllum (Anastrophyllaceae, Marchantiophyta). Phytotaxa 81 (1): 15–18. doi: 10.11646/phytotaxa.81.1.6 Váňa J, Söderström L, Hagborg A, von Konrat MJ (2013l) Notes on Early Land Plants Today. 24. What is Protolophozia? (Cephaloziellaceae, Marchantiophyta). Phytotaxa 76 (3): 50–54. doi: 10.11646/phytotaxa.76.3.12 816 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Váňa J, Söderström L, Hagborg A, von Konrat MJ (2013m) Notes on Early Land Plants Today. 32. New synonyms in Andrewsianthus and a transfer to Tritomaria (Lophoziaceae, Marchantiophyta). Phytotaxa 81 (1): 22–25. doi: 10.11646/phytotaxa.81.1.8 Váňa J, Söderström L, Hagborg A, von Konrat MJ (2014a) Notes on Early Land Plants Today. 61. New synonyms and new combinations in Cephaloziaceae and Cephaloziellaceae (Marchantiophyta). Phytotaxa 183 (4): 290–292. doi: 10.11646/phytotaxa.183.4.10 Váňa J, Söderström L, Hagborg A, von Konrat MJ (2014b) Notes on Early Land Plants Today. 52. Validation of Tritomaria camerunensis (Lophoziaceae, Marchantiophyta). Phytotaxa 167 (2): 215–216. doi: 10.11646/phytotaxa.167.2.11 Váňa J, Söderström L, Hagborg A, von Konrat MJ (2014c) Notes on Early Land Plants Today. 60. Circumscription of Gymnomitriaceae (Marchantiophyta). Phytotaxa 183 (4): 287–289. doi: 10.11646/phytotaxa.183.4.9 Váňa J, Feldberg K, Heinrichs J, Söderström L, Hagborg A, von Konrat M (2014d) Notes on Early Land Plants Today. 55. New taxa and synonyms in Adelanthaceae (Marchantiophyta). Phytotaxa 173 (1): 80–86. doi: 10.11646/phytotaxa.173.1.7 Váňa J, Söderström L, Hagborg A, von Konrat MJ (2015) Notes on Early Land Plants Today. 65. New synonyms in Scapaniaceae (Marchantiophyta). Phytotaxa 202 (1): 57. doi: 10.11646/phytotaxa.202.1.7 Vanden Berghen C (1950) Contribution à l’étude des espèces africaines du genre Lopholejeunea (Spr.) Schifn. Bulletin du Jardin Botanique de l’État, Bruxelles 20 (2): 161–179. doi: 10.2307/3666689 Vanden Berghen C (1951a) Note sur trois hépatiques du Congo Belge. Bulletin de la Société Royale de Botanique de Belgique 84: 61–64. Vanden Berghen C (1951b) Contribution a l’étude des espèces africaines du genre Ceratolejeunea (Spruce) Schifn. Bulletin du Jardin Botanique de l’État, Bruxelles 21 (1/2): 61–81. doi: 10.2307/3666810 Vanden Berghen C (1951c) Contribution à l’étude des espèces africaines du genre Archilejeunea (Spr.) Schifn. Revue Bryologique et Lichénologique 20 (1/2): 112–121. Vanden Berghen C (1951d) Contribution à l’étude des espèces africaines du genre Brachiolejeunea (Spruce) Schifn. Bulletin du Jardin Botanique de l’État, Bruxelles 21 (1/2): 87–94. doi: 10.2307/3666812 Vanden Berghen C (1957) Flore générale de Belgique, Bryophytes, Vol. 1, Fasc. 3. Jardin Botanique de l’État, Bruxelles, 271–389. Vanden Berghen C (1960a) Hépatiques récoltées en Afrique par M. Monod. Revue Bryologique et Lichénologique 29 (1/2): 50–67. Vanden Berghen C (1960b) Hépatiques récoltées par Dr J.-J. Symoens dans la région péritanganyikaise. Bulletin de la Société Royale de Botanique de Belgique 92: 111–138. Vanden Berghen C (1961) Hépatiques récoltées par le Dr J.-J. Symoens dans la région péri-tanganyikaise. Bulletin de la Société Royale de Botanique de Belgique 93: 55–74. Vanden Berghen C (1972a) Hépatiques épiphylles récoltées au Burundi par J. Lewalle. Bulletin du Jardin Botanique National de Belgique 42 (4): 431–494. doi: 10.2307/3667667 World checklist of hornworts and liverworts 817 Vanden Berghen C (1972b) Hépatiques et anthocérotées. In: Symoens J-J (Ed.) Exploration hydrobiologique du bassin du Lac Bangweolo ed du Luapula, vol. VIII, fasc. 1. Cercle Hydrobiologique de Bruxelles, Bruxelles, 1–202. Vanden Berghen C (1976a) Deux Lejeunéacées holostipées nouvelle pour la lore africaine. Revue Bryologique et Lichénologique 42 (4): 923–929. Vanden Berghen C (1976b) Frullaniaceae (Hepaticae) africanae. Bulletin du Jardin Botanique National de Belgique 46 (1/2): 1–220. doi: 10.2307/3667414 Vanden Berghen C (1977) Hépatiques épiphylles récoltées par J L De Sloover au Kivu, Zaïre, au Ruanda et au Burundi. Bulletin du Jardin Botanique National de Belgique 47 (1/2): 199–246. doi: 10.2307/3667993 Vanden Berghen C (1978) Hépatiques épiphylles récoltées en Rhodésie. Revue Bryologique et Lichénologique 44 (4): 443–452. Vanden Berghen C (1981) Le genre Plagiochila (Dum.) Dum. (Hepaticae) à Madagascar et aux Mascareignes, principalement d’après les récoltes de M. Onraedt. Bulletin du Jardin Botanique National de Belgique 51 (1/2): 41–103. doi: 10.2307/3667735 Vanden Berghen C (1984a) Le genre Lopholejeunea (Spruce) Schifn. (Lejeuneaceae, Hepaticae) en Afrique. Bulletin du Jardin Botanique National de Belgique 54 (3/4): 393–464. doi: 10.2307/3667852 Vanden Berghen C (1984b) Lejeuneaceae (Hepaticae) nouvelles de La Réunion: Marchesinia obtusifolia sp. nov. et Cheilolejeunea ecarinata sp. nov. Bulletin du Jardin Botanique National de Belgique 54 (1/2): 7–14. doi: 10.2307/3667862 Vanderpoorten A, Schäfer-Verwimp A, Heinrichs J, Devos N, Long DG (2010) he taxonomy of the leafy liverwort genus Leptoscyphus (Lophocoleaceae) revisited. Taxon 59 (1): 176–186. Vanderpoorten A, Désamoré A, Laenen B, Gradstein SR (2012) Striking autapomorphic evolution in Physotheca J.J.Engel & Gradst. (Marchantiophyta: Lophocoleaceae) blurred its actual relationships with Leptoscyphus Mitt. Journal of Bryology 34 (4): 251–256. doi: 10.1179/1743282012Y.0000000018 Verdoorn F (1928a) Kritische Bemerkungen ueber ostasiatische und ozeanische FrullaniaArten aus dem subgenus Homotropantha (De Frullaniaceis III). Revue Bryologique et Lichénologique 1: 109–122. Verdoorn F (1928b) Ueber einige amerikanische Frullaniaceae, De Frullaniaceis II. Annales de Cryptogamie Exotique 1 (2): 213–220. Verdoorn F (1929a) V Schifner – Expositio plantarum in itinere suo indico annis 1893-94 suscepto collectorum speciminibusque exsiccatis distributarum, adjectis descriptionibus novarum. Series tertia (no. 1473–2460). Frullaniaceas continens (De Frullaniaceis IV). Annales Bryologici 2: 117–154. Verdoorn F (1929b) Einige morphologische Notizen über Frullania. De Frullaniaceis VI. Annales du Jardin Botanique de Buitenzorg 40: 139–145. Verdoorn F (1930a) Revision der von Ozeanien angeführten Frullaniaceae (De Frullaniaceis VIII). Nederlandsch Kruidkundig Archief. Verslagen en Mededelingen der Nederlandsche Botanische Vereeniging 40 (2): 155–175. 818 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Verdoorn F (1930b) Frullaniaceae. Nova Guinea: Résultats des expéditions scientiiques à la Nouvelle Guinée 14: 540–548. Verdoorn F (1930c) Die Frullaniaceae der Indomalesischen Inseln (De Frullaniaceis VII). Annales Bryologici, suppl. 1: 1–187. doi: 10.1007/978-94-015-5385-8 Verdoorn F (1932a) Neue Beiträge zur Kenntnis indomalesischer Frullaniaceae (De Frullaniaceis IX). Bulletin du Jardin Botanique de Buitenzorg (sér. 3) 12 (1): 53–64. Verdoorn F (1932b) Ueber einige neue Frullania-sammlungen (De Frullaniaceis X). Nederlandsch Kruidkundig Archief. Verslagen en Mededelingen der Nederlandsche Botanische Vereeniging (ser. 3) 42 (2): 484–500. Verdoorn F (1932c) Hepaticae selectae et criticae, series III et IV (1932). Annales Bryologici 5: 125–144. Verdoorn F (1933a) Hepaticae selectae et criticae, series V et VI. Annales Bryologici 6: 95–104. Verdoorn F (1933b) Die von V. Schifner (1893–1894) und von Fr. Verdoorn (1930) auf den indomalesischen Inseln gesammelten Lejeuneaceae Holostipae. De Frullaniaceis XI. Recueil des Travaux Botaniques Néerlandais 30: 212–233. Verdoorn F (1933c) Ueber zwei neue Gattungen der Lebermoose. De Frullaniaceis XIII. Annales Bryologici 6: 88–91. Verdoorn F (1934a) De Frullaniaceis XV. Die Lejeuneaceae Holostipae der Indomalaya unter Berücksichtung sämtlicher aus Asien, Australien, Neuseeland und Ozeanien angeführten Arten. Annales Bryologici, suppl. 4: 40–192. Verdoorn F (1934b) De Frullaniaceis XIV. Revision der von Ozeanien, Australien und Neuseeland angeführten Lejeuneaceae Holostipae. Blumea 1 (1): 216–240. Verdoorn F (1934c) Bryologie und Hepaticologie, ihre Methodik und Zukunft. Annales Bryologici, suppl. 4: 1–39. Verdoorn F (1934d) Lejeuneaceae Holostipae. Nova Guinea: Résultats des expéditions scientiiques à la Nouvelle Guinée 18: 1–8. Verdoorn F (1935) Hepaticae selectae et criticae, series VII (1934) et series VIII (1935) and Musci selecti et critici, series I (1934) et series II (1935). Annales Bryologici 8: 150–158. Verdoorn F (1936) Hepaticae Selecti et Critici (ser. 9). Utrecht, 401–450. (non vidi) Verma PK, Rawat KK (2013) Lejeunea srivastavae sp. nov. (Marchantiophyta: Lejeuneaceae), from Nilgiri hills of Western Ghats (India). Taiwania 58 (1): 7–11. Verma PK, Rawat KK (2014) Present status of genus Rectolejeunea A.Evans (Marchantiophyta) in India. Journal of Bryology 36 (2): 160–162. doi: 10.1179/1743282014Y.0000000097 Verma PK, Srivastava SC (2007) Diversity of genus Taxilejeunea (Spr.) Schifn. in Western Ghats (India). Proceedings of the National Academy of Sciences of India. Section B, Biological Sciences 77 (2): 206–214. Verma PK, Srivastava SC (2011) Species diversity of genus Microlejeunea Steph. (Lejeuneaceae, Hepaticae) in Nilgiri Hills, Western Ghats, Tamil Nadu, India. Journal of the Bombay Natural History Society 108 (2): 120–125. Vianna EC (1981) Sphaerocarpos muccilloi, a new hepatic from Brazil. Lindbergia 7 (1): 58–60. World checklist of hornworts and liverworts 819 Vianna EC (1985) Flora illustrada do Rio Grande do Sul / 15. Marchantiales. Boletim do Instituto de Biociências, Universidade Federal do Rio Grande do Sul 38: 1–213. Villarreal JC, Renner SS (2012). Hornwort pyrenoids, carbon-concentrating structures, evolved and were lost at least ive times during the last 100 million years. Proceedings of the National Academy of Science U.S.A. 109 (46): 18873–18878. doi: 10.1073/pnas.1213498109 Villarreal JC, Renner SS (2014) A review of molecular clock calibrations and substitution rates in liverworts, mosses, and hornworts, and a timeframe for a taxonomically cleaned-up genus Nothoceros. Molecular Phylogenetics and Evolution 78: 25–35. doi: 10.1016/j.ympev.2014.04.014 Villarreal JC, Hässel GG, Salazar Allen N (2007) Nothoceros superbus (Dendrocerotaceae), a new hornwort from Costa Rica. Bryologist 110 (2): 279–285. doi: 10.1639/0007-2745(2007)110[279:NSDANH]2.0.CO;2 Villarreal JC, Cargill DC, Goinet B (2010a) Phaeomegaceros squamuliger subspecies hasselii (Dendrocerotaceae, Anthocerotophyta), a new taxon from the southern hemisphere. Nova Hedwigia 91 (3/4): 349–360. doi: 10.1127/0029-5035/2010/0091-0349 Villarreal JC, Goinet B, Duf RJ, Cargill DC (2010b) Phylogenetic delineation of Nothoceros and Megaceros (Dendrocerotaceae). Bryologist 113 (1): 106–113. doi: 10.1639/00072745-113.1.106 Villarreal JC, Campos LV, Uribe J, Goinet B (2012) Parallel evolution of endospory within hornworts: Nothoceros renzagliensis (Dendrocerotaceae), sp. nov. Systematic Botany 37 (1): 31–37. doi: 10.1600/036364412X616594 Villarreal JC, Cargill C, Söderström L, Hagborg A, von Konrat M (2015) Notes on Early Land Plants Today. 70. Nomenclatural noted in hornworts (Anthocerotophyta). Phytotaxa 208 (1): 92–96. doi: 10.11646/phytotaxa.208.1.9 Villars D (1789) Histoire des plantes de Dauphiné, Tome troisième, parte 2. Prevost, Paris, 581–1089. Vilnet AA, Konstantinova NA (2007) [Molecular phylogeny of Scapaniaceae Mig. (Hepaticae)]. Proceedings of Conference on plant morphology and taxonomy dedicated to 300th anniversary of Carl Linnaeus. KMK, Moscow, 53–55. Vilnet AA, Konstantinova NA, Troitsky AV (2007a) On molecular phylogeny of Gymnomitriaceae H. Klinggr. (Hepaticae). Computational phylogenetics and molecular systematics “CPMS’ 2007”. Conference proceedings. KMK, Moscow, 24–26. Vilnet AA, Milyutina IA, Konstantinova NA, Ignatov MS, Troitsky AV (2007b) Phylogeny of the Genus Lophozia (Dumort.) Dumort. s. str. inferred from nuclear and chloroplast sequences ITS1-2 and TRNL-F. Russian Journal of Genetics 43 (11): 1306–1313. doi: 10.1134/S1022795407110142 Vilnet AA, Konstantinova NA, Troitsky VA (2008) Phylogeny and systematics of the genus Lophozia s. str. (Dumort.) Dumort. (Hepaticae) and related taxa from nuclear ITS1–2 and chloroplast trnL-F sequences. Molecular Phylogenetics and Evolution 47 (1): 403–418. doi: 10.1016/j.ympev.2007.12.013 Vilnet AA, Konstantinova NA, Troitsky AV (2010) Molecular insight on phylogeny and systematics of the Lophoziaceae, Scapaniaceae, Gymnomitriaceae and Jungermanniaceae. Arctoa 19: 31–50. 820 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Vilnet AA, Konstantinova NA, Troitsky AV (2011) Taxonomical rearrangements of Solenostomataceae (Marchantiophyta) with description of a new family Endogemmataceae based on trnL-F cpDNA analysis. Folia Cryptogamica Estonica 48: 125–133. Vilnet AA, Konstantinova NA, Troitsky AV (2012) Molecular phylogeny and systematics of the suborder Cephaloziineae with special attention to the family Cephaloziaceae s.l. (Jungermanniales, Marchantiophyta). Arctoa 21: 113–132. Vilnet AA, Borovichev EA, Bakalin VA (2014) Frullania subarctica - a new species of the Frullania tamarisci complex (Frullaniaceae, Marchantiophyta). Phytotaxa 173 (1): 61–72. doi: 10.11646/phytotaxa.173.1.5 Volk OH (1981) Beiträge zur Kenntnis der Lebermoose (Hepaticae) aus Südwestafrika (Namibia), II. Mitteilungen der Botanischen Staatssammlung München 17: 245–252. Volk OH (1983) Vorschläge für eine Neugliederung der Gattung Riccia L. Mitteilungen der Botanischen Staatssammlung München 19: 453–465. Volk OH (1984) Beiträge zur Kenntnis der Lebermoose (Marchantiales) aus Südwestafrika (Namibia). IV. Zur Biologie einiger Hepaticae mit besonderer Berücksichtigung der Gattung Riccia. Nova Hedwigia 39: 117–143. Volk OH (1988) Riccia crenatodentata (Marchantiales) sp. nov. aus Arabien. Nova Hedwigia 46 (1/2): 27–35. Volk OH, Perold SM (1984) Studies in the liverwort genus Riccia (Marchantiales) from the southwest Cape. Bothalia 15 (1/2): 117–124. doi: 10.4102/abc.v15i1 & 2.1110 Volk OH, Perold SM (1985) Studies in the genus Riccia (Marchantiales) from southern Africa. 1. Two new species of the section Pilifer: R. duthieae and R. alatospora. Bothalia 15 (3/4): 531–539. doi: 10.4102/abc.v15i3 & 4.1837 Volk OH, Perold SM (1986a) Studies in the genus Riccia (Marchantiales) from southern Africa. 6. R. hirsuta, a new species, in a new section. Bothalia 16 (2): 187–191. doi: 10.4102/ abc.v16i2.1086 Volk OH, Perold SM (1986b) Studies in the genus Riccia (Marchantiales) from southern Africa. 4. hree endemic species, R. natalensis Sim, R. microciliata sp. nov. and R. mammifera sp. nov. Bothalia 16 (2): 169–180. doi: 10.4102/abc.v16i2.1084 Volk OH, Perold SM (1986c) Studies in the genus Riccia (Marchantiales) from southern Africa. 3. R. schelpei, a new species, in the new subgenus Chartacea. Bothalia 16 (1): 29–33. doi: 10.4102/abc.v16i1.1057 Volk OH, Perold SM (1986d) Studies in the genus Riccia (Marchantiales) from southern Africa. 5. R. rosea, a new species. Bothalia 16 (2): 181–185. doi: 10.4102/abc.v16i2.1085 Volk OH, Perold SM (1990) Studies in the genus Riccia (Marchantiales) from southern Africa. 15. R. hirsuta and R. tomentosa, sp. nov., two distinct species previously treated as one. Bothalia 20 (1): 23–29. doi: 10.4102/abc.v20i1.890 Volk OH, Perold SM, Bornefeld T (1988) Studies in the genus Riccia (Marchantiales) from southern Africa. 10. Two new white-scaled species of the group ‘Squamatae’: R. argenteolimbata and R. albornata. Bothalia 18 (2): 155–163. doi: 10.4102/abc.v18i2.998 von Flotow JCG, von Göppert R, Nees CG (1842) Ueber Páo Pereira und mehrere darauf wachsende kryptogamische Planzen. Repertorium für die Pharmacie 76: 32–55. World checklist of hornworts and liverworts 821 von Konrat MJ, Braggins JE (2003) A new and unusual species of Frullania (Jubulaceae) from Tasmania, Australia. New Zealand Journal of Botany 41 (1): 55–62. doi: 10.1080/0028825X.2003.9512831 von Konrat MJ, Braggins JE (2005) Frullania wairua, a new and seemingly rare liverwort species from Northland, New Zealand. New Zealand Journal of Botany 43 (4): 885–893. doi: 10.1080/0028825X.2005.9512998 von Konrat M, Renner MAM, Söderström L, Hagborg A, Mutke J (2008a) Early Land Plants Today: Liverwort species diversity and the relationship with higher taxonomy and higher plants. Fieldiana: Botany (n.ser.) 47: 91–104. doi: 10.3158/0015-0746-47.1.91 von Konrat MJ, Hagborg A, Söderström L, Mutke J, Renner M, Gradstein SR, Engel JJ, Zhu R-L, Pickering J (2008b) Early Land Plants Today: Global patterns of liverwort diversity, distribution, and loristic knowledge. In: Mohamed H, Baki BB (Eds) Nasrulhaq-Boyce A, Lee PKY, Bryology in the New Millennium. University of Malaya, Kuala Lumpur, 425–438. von Konrat M, Söderström L, Renner MAM, Hagborg AH, Briscoe L, Engel JJ (2010a) Early Land Plants Today (ELPT): How many liverwort species are there?. Phytotaxa 9: 22–40. doi: 10.11646/phytotaxa.9.1.5 von Konrat MJ, Hentschel J, Heinrichs J, Braggins JE, Pócs T (2010b) Forty-one degrees below and sixty years in the dark: Frullania sect. Inconditum, a new section of Australasian Frullania species including F. colliculosa, sp. nov., F. hodgsoniae, nom. and stat. nov., F. aterrima, and F. hattorii (Frullaniaceae, Marchantiophyta). Nova Hedwigia 91 (3/4): 471–500. doi: 10.1127/0029-5035/2010/0091-0471 von Konrat M, Söderström L, Hagborg A, Crosby MR, Engel JJ (2010c) Early Land Plants Today: Index of liverworts & hornworts 2006-2008. Cryptogamie, Bryologie 31 (1): 3–30. von Konrat M, Söderström L, Hagborg A (2010d) he Early Land Plants Today project: A community-driven efort and a new partnership with Phytotaxa. Phytotaxa 9: 11–21. doi: 10.11646/phytotaxa.9.1.4 von Konrat MJ, Hentschel J, Heinrichs J, Braggins JE (2011) Deep southern hemisphere connections: A revision of Frullania sect. Amphijubula. Bryologist 114 (1): 52–66. doi: 10.1639/0007-2745-114.1.52 von Konrat MJ, Söderström L, Hagborg A (2012a) Notes on Early Land Plants Today. 7. Transfer of Jubulopsis novae-zelandiae to Lepidolaena. Phytotaxa 65: 51. von Konrat MJ, de Lange P, Greif M, Strozier L, Hentschel J, Heinrichs J (2012b) Frullania knightbridgei, a new liverwort (Frullaniaceae, Marchantiophyta) species from the deep south of Aotearoa-New Zealand based on an integrated evidence-based approach. PhytoKeys 8: 13–36. doi: 10.3897/phytokeys.8.2496 von Konrat M, de Lange P, Larraín J, Hentschel J, Carter B, Shaw J, Shaw B. (2013) A small world: uncovering hidden diversity in Frullania – a new species from Aotearoa - New Zealand. Polish Botanical Journal 58 (2): 437–447. doi: 10.2478/pbj-2013-0056 von Malmborg S (1933) Cryptothallus nov. gen. Ein saprofytisches Lebermoos. Annales Bryologici 6: 122–123. Voronov’ IuN (1914) Materìalny k’ lore pechenochnikov’ Kavkaza. Izvěstìâ Kavkazskago Muzeâ 8: 81–88. 822 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Waddell CH (1897) Moss exchange club catalogue of British hepaticae. W. Wesley and Son, London, 7 pp. Wagner DH (2013) Rivulariella, gen. nov. (Jungermanniaceae), endemic to western North America. Phytoneuron 2013 (10): 1–9. Wahlenberg G (1811) Kamtschadalische Laub- und Lebermoose, gesammelt auf der russischen Entdeckungsreise von dem Herrn Hofrath Tilesius. Magazin für die neuesten Entdeckungen in der gesammten Naturkunde, Gesellschaft Naturforschender Freunde zu Berlin 5: 289–297. Wahlenberg G (1812) Flora lapponica. Taberna Libraria Scholae Realis, Berlin, 550 pp. Wahlenberg G (1814) Flora carpatorum principalium. Impensis Vandenhock et Ruprecht, Gottingae, 408 pp. Wang J, Zhu R-L (2008) Lejeunea laii nom. nov. for Lejeunea ramulosa (Herzog) R. M. Schust. (Jungermanniopsida: Lejeuneaceae) from Taiwan. Journal of Bryology 30 (2): 173–174. doi: 10.1179/174328208X300624 Wang J, Lai M-J, Zhu R-L (2011) Liverworts and hornworts of Taiwan: an updated checklist and loristic accounts. Annales Botanici Fennici 48 (5): 369–395. Wang J, Zhou L-Y, Zhu R-L (2014a) A new epiphyllous species of Cololejeunea (Lejeuneaceae, Marchantiophyta) from China. Phytotaxa 161 (2): 165–168. doi: 10.11646/phytotaxa.161.2.8 Wang J, Gradstein SR, Shi X-Q, Zhu R-L (2014b) Phylogenetic position of Trocholejeunea and a new infrageneric classiication of Acrolejeunea (Lejeuneaceae, Marchantiophyta). Bryophyte Diversity and Evolution 36 (1): 31–44. doi: 10.11646/bde.36.1.3 Wang J, Gradstein SR, Cheng X-F, Zhu R-L (2014c) A new species of Trocholejeunea (Lejeuneaceae, Marchantiophyta) from China. Phytotaxa 174 (5): 296–300. doi: 10.11646/phytotaxa.174.5.7 Warnstorf C (1886) Mooslora der Provinz Brandenburg. Verhandlungen des Botanischen Vereins der Provinz Brandenburg 27 (1): 1–94. Warnstorf C (1902) Kryptogamenlora der Mark Brandenburg, Erster Band, Leber- und Torfmoose, 2 Heft. Gebrüder Borntraeger, Leipzig, 113–288. Warnstorf C (1913) Zur Bryogeographie des Russischen Reiches. Hedwigia 53 (3): 184–320. Warnstorf C (1916) Bryophyta nova europaea et exotica. Hedwigia 57 (1/2): 62–131. Warnstorf C (1917) Die europäischen Artengruppen der Gattung Calypogeia Raddi (1820). Bryologische Zeitschrift 1 (7): 97–114. Warnstorf C (1921) Die Unterfamilie der Scapanioideen (Spruce, 1885). Hedwigia 63 (2): 58–116. Watts WW (1902) Notes on some New South Wales hepatics. Proceedings of the Linnean Society of New South Wales (ser. 2) 26 (104): 633–634. Watts WW (1903) Notes on some New South Wales hepatics. Proceedings of the Linnean Society of New South Wales (ser. 2) 27 (108): 493–494. Weber GH (1778) Spicilegium lorae goettingensis. Sumptibus Ettingeri, Gothae [Göttingen], 288 pp. Weber F (1815) Historiae muscorum hepaticarum prodromus. Aug. Hesse, Academiae bibliopolae, Kiel, 160 pp. World checklist of hornworts and liverworts 823 Weber F, Mohr DMH (1807) Botanisches Taschenbuch auf das Jahr 1807. Deutschlands Kryptogamische Gewächse. Erste Abtheilung. Filices, Musci frondosi et Hepatici. Akademischen Buchhandlung, Kiel, 509 pp. Wei Y-M, Zhu R-L (2013a) Transfer of two Asiatic taxa from Lejeunea to Microlejeunea (Lejeuneaceae, Marchantiophyta). Cryptogamie, Bryologie 34 (3): 307–311. doi: 10.7872/cryb. v34.iss3.2013.307 Wei Y-M, Zhu R-L (2013b) Notes on Early Land Plants Today. 39. Microlejeunea indica (Marchantiophyta, Lejeuneaceae). Phytotaxa 97 (2): 63–64. doi: 10.11646/phytotaxa.97.2.6 Wei Y-M, He Q, Gradstein SR, Campos LV, Zhu R-L (2013) Notes on Early Land Plants Today. 46. Transfer of Vitalianthus urubuensis (Marchantiophyta, Lejeuneaceae) to Cheilolejeunea. Phytotaxa 152 (1): 50–52. doi: 10.11646/phytotaxa.152.1.5 Wei Y-M, Zhu R-L, Gradstein SR (2014) Notes on Early Land Plants Today. 49. On Lejeunea huctumalcensis and the resurrection of Otigoniolejeunea, an older name for Physantholejeunea (Marchantiophyta, Lejeuneaceae). Phytotaxa 162 (4): 236–238. doi: 10.11646/phytotaxa.162.4.8 Wellman CH, Osterlof PL, Mohiuddin U (2003) Fragments of the earliest land plants. Nature 425 (6955): 282–285. doi: 10.1038/nature01884 Wheeler LC (1934) he genus Grimaldia. Bryologist 37 (5): 87–88. doi: 10.2307/3239514 Wickett NJ, Goinet B (2008) Origin and relationships of the myco-heterotrophic liverwort Cryptothallus mirabilis Malmb. (Metzgeriales, Marchantiophyta). Botanical Journal of the Linnean Society 156 (1): 1–12. doi: 10.1111/j.1095-8339.2007.00743.x Wigginton MJ (2002) Checklist and distribution of the liverworts and hornworts of sub-Saharan Africa, including the East African Islands. Tropical Bryology Research Reports 3: 1–88. Wigginton MJ (2006) Bryophytes of St Helena, south Atlantic Ocean. 1. hree new species of Cololejeunea (Jungermanniales, Lejeuneaceae), C. dianae sp. nov., C. sanctaehelenae sp. nov. and C. grossestyla sp. nov. Journal of Bryology 28 (4): 363–373. doi: 10.1179/174328206X152298 Wigginton MJ (2007) Bryophytes of St Helena, south Atlantic Ocean. 2. Lejeunea sanctahelenae sp. nov. (Jungermanniales, Lejeuneaceae), an unusual species with a hooded lobule. Journal of Bryology 29 (1): 12–17. doi: 10.1179/174328207X160595 Wigginton MJ (2009) Checklist and distribution of the liverworts and hornworts of sub-Saharan Africa, including the East African Islands (edition 3, 24 January 2009). Tropical Bryology Research Reports 8: 1–116. Wigginton MJ (2012) Bryophytes of St Helena, south Atlantic Ocean. 6. Cheilolejeunea (Spruce) Schifn. (Jungermanniales: Lejeuneaceae), including C. microscypha (Hook.f. & Taylor) M.Wigginton, comb. nov. and C. rotalis (Hook.f. & Taylor) M.Wigginton, comb. nov. Journal of Bryology 34 (4): 268–276. doi: 10.1179/1743282012Y.0000000027 Wigginton MJ (2013) Bryophytes of St. Helena, South Atlantic Ocean. 7. Cylindrocolea (Jungermanniales, Cephaloziellaceae): C. sanctae-helenae M. Wigginton sp. nov. Polish Botanical Journal 58 (1): 107–115. doi: 10.2478/pbj-2013-0012 Wigginton MJ, Grolle R (1996) Catalogue of the hepaticae and anthocerotae of Sub-saharan Africa. Bryophytorum Bibliotheca 50: 1–267. 824 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Wigginton MJ, Porley RD, Hodgetts NG (2007) British Bryological Society expedition to Mulanje Mountain, Malawi. 18. Cololejeunea lichenyae, a new species of Lejeuneaceae (Jungermanniopsida) from Malawi. Journal of Bryology 29 (1): 7–11. doi: 10.1179/174328207X160586 Wigglesworth G (1929) A new California species of Sphaerocarpos. University of California Publications in Botany 16 (3): 129–137. Wigglesworth G (1937) South African species of Riella, including an account of the developmental stages of three of the species. Journal of the Linnean Society. Botany 51 (339): 309–332. doi: 10.1111/j.1095-8339.1937.tb01910.x Wild CJ (1893) On the occurrence of a new Dendroceros in Queensland. Transactions of the Natural History Society of Queensland 1: 49–50. Williams H (1968) A new member of Lophoziaceae from Ontario. Bryologist 71 (1): 34–38. doi: 10.2307/3240650 Wilson W (1841) Musci americani. Warrington, 1–180. (non vidi) Wilson R, Gradstein SR, Schneider H, Heinrichs J (2007) Unravelling the phylogeny of Lejeuneaceae (Jungermanniopsida): evidence for four main lineages. Molecular Phylogenetics and Evolution 43 (1): 270–282. doi: 10.1016/j.ympev.2006.10.017 Winkler S (1967) Die epiphyllen Moose der Nebelwälder von El Salvador, C. A. Revue Bryologique et Lichénologique 35 (1/4): 303–369. Winkler S (1969) Systematisch-anatomische Untersuchungen über die marsupialen Lebermoose der Sierra Nevada de Santa Marta in Kolumbien. Mitteilungen aus dem Instituto Colombo-Alemán de Investigaciones Cientíicas 3: 59–76. Winkler S (1976) Die Hepaticae der Sierra Nevada de Santa Marta, Kolumbien. I. Terrestrische, epixyle und epipetrische Arten. Revue Bryologique et Lichénologique 42 (3): 789–827. Withering W (1776) A botanical arrangement of all the vegetables naturally growing in Great Britain, vol. II. M. Swinney, Birmingham, 385–838. Withering W (1796) An arrangement of British Plants, ed. 3. M. Swinney, Birmingham, 513– 920. Wu P-C, But PP (2009) Hepatic lora of hong Kong. North-Eastern Forestry University Press, Harbin, 194 pp. Wu P-C, Lin P-J (1978) A preliminary observation on the hepaticae of the Island Hainan, China and their phytogeographical relationships. Acta Phytotaxonomica Sinica 16 (2): 56–71. Wu P-C, Lou J-S (1978) [Studyies [sic!] on the epiphyllous liverworts of China (II) he epiphyllous liverworts from Tibet]. Acta Phytotaxonomica Sinica 16 (4): 102–112. Yamada K (1973a) A study of Radula from Mt. Kinabalu, north Borneo, collected by Drs. Z. Iwatsuki and M. Mizutani. Journal of Japanese Botany 48 (5): 133–137. Yamada K (1973b) A new species of Radula from Mt. Kinabalu, north Borneo. Miscellanea Bryologica et Lichenologica 6 (6): 97–98. Yamada K (1974) Notes on Frullania inlata complex from Japan. Miscellanea Bryologica et Lichenologica 6 (9): 162–164. Yamada K (1975a) Notes on Radula from Tanzania, East Africa. Journal of Japanese Botany 50 (4): 115–118. World checklist of hornworts and liverworts 825 Yamada K (1975b) Radula collections made by Dr. H. Inoue in Ceylon. Journal of Japanese Botany 50 (12): 372–378. Yamada K (1979a) A new species of Radula (Hepaticae) from Ceylon. Miscellanea Bryologica et Lichenologica 8 (6): 113–114. Yamada K (1979b) A revision of Asian taxa of Radula, Hepaticae. Journal of the Hattori Botanical Laboratory 45: 201–322. Yamada K (1982a) Some new species of Radula (Hepaticae). Journal of the Hattori Botanical Laboratory 51: 323–328. Yamada K (1982b) Notes on Latin American species of the genus Radula, hepaticae 1. Miscellanea Bryologica et Lichenologica 9 (6): 121–123. Yamada K (1983) Four new species of Radula from Cuba. Journal of the Hattori Botanical Laboratory 54: 241–249. Yamada K (1984a) A Radula collection made by Dr. M. L. Hicks in Queensland, Australia. Cryptogamie: Bryologie, Lichénologie 5 (1/2): 191–199. Yamada K (1984b) Tasmanian species of Radula (Hepaticae) collected by Mrs. A. V. Ratkowsky. Journal of Japanese Botany 59 (3): 91–96. Yamada K (1985a) hree new species of Radula (Hepaticae) from Papua New Guinea. Journal of Japanese Botany 60 (9): 257–264. Yamada K (1985b) Radula collections made by Drs. Z. Iwatsuki and N. Kitagawa in New Caledonia. Journal of the Hattori Botanical Laboratory 58: 111–130. Yamada K (1987) A preliminary study of the genus Radula from Queensland, Australia. Journal of the Hattori Botanical Laboratory 62: 191–200. Yamada K (1990) Two new species of Radula from (Hepaticae) from Australia and Brazil. Journal of Japanese Botany 65 (1): 1–6. Yamada K (1993) Four new species of Radula from Neotropics. Journal of the Hattori Botanical Laboratory 74: 35–44. Yamada K (1997) Two new Radula species (Hepaticae, Radulaceae) from the Neotropica. Journal of the Hattori Botanical Laboratory 82: 337–342. Yamada K, Gradstein SR (1991) he genus Radula (Hepaticae) in the Galapagos Islands. Tropical Bryology 4: 63–68. Yamada K, Piippo S (1989) Bryophyte lora of the Huon Peninsula, Papua New Guinea. XXXII. Radula (Radulaceae, Hepaticae). Annales Botanici Fennici 26 (4): 349–387. Yamaguchi T (1984) Drepanolejeunea obtusifolia sp. nov. and Euosmolejeunea fuscobrunnea Horik. from the Yaeyama Islands, Japan. Journal of Japanese Botany 59 (11): 332–336. Yang BY, Lee WC (1964) Bryophytic lora of Chi-Tou. Botanical Bulletin of Academia Sinica (n.ser.) 5 (2): 181–194. Yang J-D, Lin S-H (2014) Cololejeunea nanhutashanensis (Lejeuneaceae), a new species from Taiwan. Phytotaxa 177 (1): 56–60. doi: 10.11646/phytotaxa.177.1.5 Yano O (1984) Checklist of Brazilian liverworts and hornworts. Journal of the Hattori Botanical Laboratory 56: 481–548. Yasuda A (1911) Shokubutsugaku kakuron: Inkabu [Botany essays: Cryptogams]. Hakubunkan, Tokyo, 1230 pp. 826 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Yatsentyuk SP, Konstantinova NA, Ignatov MS, Hyvönen J, Troitsky AV (2004) On phylogeny of Lophoziaceae and related families (Hepaticae, Jungermanniales) based on trnLtrnF intron-spacer sequences of chloroplast DNA. Monographs in Systematic Botany from the Missouri Botanical Garden 98: 151–167. Ye W, Zhu R-L (2010) Leucolejeunea, a new synonym of Cheilolejeunea (Lejeuneaceae), with special reference to new combinations and nomenclature. Journal of Bryology 32 (4): 279– 282. doi: 10.1179/037366810X12814321877507 Ye W, Zhu R-L, Shaw AJ, Gradstein SR (2011) Proposal to conserve the name Cheilolejeunea against Omphalanthus (Lejeuneaceae). Taxon 60 (2): 588–589. Ye W, Zhu RL, Long DG (2013a) Range extension and description for the rare Cheilolejeunea chenii (Lejeuneaceae, Marchantiophyta), with reference to the Cyrtolejeunea clade. Journal of Bryology 35 (2): 143–147. doi: 10.1179/1743282013Y.0000000050 Ye W, Wei Y-M, Schäfer-Verwimp A, Zhu R-L (2013b) Phylogenetic position of Oryzolejeunea (Lejeuneaceae, Marchantiophyta): Evidence from molecular markers and morphology. Journal of Systematics and Evolution 51 (4): 468–475. doi: 10.1111/j.17596831.2012.00238.x Ye W, Gradstein SR, Shaw AJ, Ho BC, Schäfer-Verwimp A, Pócs T, Heinrichs J, Zhu R.-L. (2015) Phylogeny and classiication of Lejeuneaceae subtribe Cheilolejeuneinae (Marchantiophyta) based on nuclear and plastid molecular markers. Cryptogamie, Bryologie 36(4): 313–333. doi: 10.7872/cryb/v36.iss4.2015.313 Yi Y-J, Fu X, Gao C (2001) A new species of Herbertus (Hepaticae) from China. Acta Phytotaxonomica Sinica 39 (1): 89–91. Yu Y, Pócs T, Schäfer-Verwimp A, Heinrichs J, Zhu R-L, Schneider H (2013) Evidence for rampant homoplasy in the phylogeny of the epiphyllous liverwort genus Cololejeunea (Lejeuneaceae). Systematic Botany 38 (3): 553–563. doi: 10.1600/036364413X670304 Yu Y, Pócs T, Zhu R-L (2014) Notes on Early Land Plants Today. 62. A synopsis of Myriocoleopsis (Lejeuneaceae, Marchantiophyta) with special reference to transfer of Cololejeunea minutissima to Myriocoleopsis. Phytotaxa 183 (4): 293–297. doi: 10.11646/phytotaxa.183.4.11 Yuzawa Y (1991) A monograph of subg. Chonanthelia of gen. Frullania (Hepaticae) of the world. Journal of the Hattori Botanical Laboratory 70: 181–291. Yuzawa Y, Hattori S (1983) A new Frullania species from Fukushima, Japan. Journal of Japanese Botany 58 (2): 43–45. Yuzawa Y, Hattori S (1988a) A new species of Frullania from Brazil. Journal of Japanese Botany 63 (1): 30–32. Yuzawa Y, Hattori S (1988b) A new species of Frullania from Haiti, West Indies. Journal of Japanese Botany 63 (11): 361–364. Yuzawa Y, Hattori S (1989) A new species of Frullania (Hepaticae) from Brazil. Journal of Japanese Botany 64 (2): 37–40. Yuzawa Y, Koike N (1994) A Frullania (Hepaticae) collection made by Dr. M. Higuchi in Nepal. Journal of the Hattori Botanical Laboratory 75: 193–199. Yuzawa Y, Mues R, Hattori S (1987) Morphological and chemical studies on the taxonomy of 14 Frullania species, subgenus Chonanthelia. Journal of the Hattori Botanical Laboratory 63: 425–436. World checklist of hornworts and liverworts 827 Zartman CE, Ackerman IL (2002) A new species of Vitalianthus (Lejeuneaceae, Hepaticae) from the Brazilian Amazon. Bryologist 105 (2): 267–269. doi: 10.1639/0007-2745(2002)105[0267:ANSOVL]2.0.CO;2 Zhou L-P, Zhang L, Xing F-W (2012) he genus Bazzania in China and adjacent regions. 1. Bazzania dulongensis L.-P.Zhou & L.Zhang sp. nov. and Bazzania hainanensis L.-P.Zhou & L.Zhang sp. nov. Journal of Bryology 34 (1): 22–31. doi: 10.1179/1743282011Y.0000000039 Zhu R-L (1995) Notes on some species of the genus Cololejeunea (Lejeuneaceae, Hepaticae) in China. Journal of the Hattori Botanical Laboratory 78: 83–109. Zhu R-L (2006a) Taxonomy and distribution of Cheilolejeunea krakakammae (Lejeuneaceae, Jungermanniopsida, Marchantiophyta), with a description and illustrations of Cheilolejeunea laevicalyx from Bolivia, Colombia and Ecuador. Nova Hedwigia 83 (1/2): 187–198. doi: 10.1127/0029-5035/2006/0083-0187 Zhu R-L (2006b) Cololejeunea dauphinii nom. nov. for Cololejeunea tixieri M. Morales & G. Dauphin from Panamá (Jungermanniopsida: Lejeuneaceae). Journal of Bryology 28 (3): 277. doi: 10.1179/174328206X157202 Zhu R-L, Cheng X-F (2008) he status of Amblyolejeunea (Lejeuneaceae) from Ecuador and Guadeloupe. Systematic Botany 33 (4): 617–620. doi: 10.1600/036364408786500181 Zhu R-L, Gradstein SR (2004) Lopholejeunea minuta R.L. Zhu & Gradst. (Lejeuneaceae, Hepaticae), a new species from the Solomon Islands, Paciic Oceania. Nova Hedwigia 78 (3/4): 435–438. doi: 10.1127/0029-5035/2004/0078-0435 Zhu R-L, Gradstein SR (2005) Monograph of Lopholejeunea (Spruce) Schifn. (Lejeuneaceae, Hepaticae) in Asia. Monographs in Systematic Botany from the Missouri Botanical Garden 74: 1–98. Zhu R-L, Grolle R (2002) Metalejeunea winkleri R.-L. Zhu & Grolle (Lejeuneaceae, Hepaticae), a new species from Borneo. Nova Hedwigia 74 (3/4): 497–500. doi: 10.1127/00295035/2002/0074-0497 Zhu R-L, Müller F (2012) Cheilolejeunea hyalomarginata, a remarkable new species of Lejeuneaceae (Marchantiophyta) from New Caledonia. Bryologist 115 (2): 217–221. doi: 10.1639/0007-2745-115.2.217 Zhu R-L, Reiner-Drehwald ME (2004) Lejeunea boliviensis, a remarkable species with bizarre underleaves and eplicate perianths. Bryologist 107 (2): 237–241. doi: 10.1639/0007-2745(2004)107[0237:LBARSW]2.0.CO;2 Zhu R-L, So ML (1997a) A new record of the genus Otolejeunea (Hepaticae, Lejeuneaceae) in subtropical China. Annales Botanici Fennici 34 (4): 285–289. Zhu R-L, So ML (1997b) Frullania fengyangshanensis (Hepaticae), a new species from China. Bryologist 100 (3): 356–358. doi: 10.2307/3244504 Zhu R-L, So ML (1998) A new species of Otolejeunea (Hepaticae, Lejeuneaceae) from the Philippines. Systematic Botany 23 (2): 231–234. doi: 10.2307/2419590 Zhu R-L, So ML (1999a) A note on Trachylejeunea chinensis (Hepaticae, Lejeuneaceae). Taxon 48 (3): 489–492. doi: 10.2307/1224560 Zhu R-L, So ML (1999b) Cololejeunea zangii (Hepaticae, Lejeuneaceae), a new epiphyllous species from China. Systematic Botany 24 (4): 501–505. doi: 10.2307/2419638 Zhu R-L, So ML (2000) he genus Tuyamaella (Hepaticae, Lejeuneaceae) in China. Nova Hedwigia 70 (1/2): 185–192. 828 Lars Söderström et al. / PhytoKeys 59: 1–828 (2016) Zhu R-L, So ML (2001) Epiphyllous liverworts of China. Beihefte zur Nova Hedwigia 121: 1–418. Zhu R-L, So ML (2002) Notes on Taxilejeunea latilobula Herzog (Marchantiopsida, Lejeuneaceae). Journal of Bryology 24 (2): 168–170. doi: 10.1179/037366802125001097 Zhu R-L, Wang Y (1992) [A preliminary revision of epiphyllous liverworts from Dinghushan]. Journal of East China Normal University: Natural Science Edition 2: 90–97. Zhu R-L, Wei Y-M (2012) Taxonomic notes on Trachylejeunea malangensis Herzog (Lejeuneaceae) known from Borneo. Journal of Bryology 34 (4): 318–321. doi: 10.1179/1743282012Y.0000000010 Zhu R-L, Hu R-L, Zhang G-Z (1994) Epiphyllous liverworts from Baishanzu Nature Reserve, Zhejiang Province, China. Hikobia 11: 543–547. Zhu R-L, So ML, Cao T, Gao Q (1999) Neurolejeunea fukiensis belongs to Cheilolejeunea (Lejeuneaceae, Hepaticae). Taxon 48 (4): 663–666. doi: 10.2307/1223637 Zhu R-L, So ML, Grolle R (2000) Cheilolejeunea gaoi (Hepaticae, Lejeuneaceae), a new species from Guanxi, China. Bryologist 103 (3): 499–502. doi: 10.1639/0007-2745(2000)103[0499:CG HLAN]2.0.CO;2 Zhu R-L, Zheng M, Zhao X (2004) Taxonomic studies on the genus Cololejeunea (Lejeuneaceae, Hepaticae). I. Notes on several little known species of Asia and Oceania. Nova Hedwigia 79 (3/4): 527–535. doi: 10.1127/0029-5035/2004/0079-0527 Zhu R-L, Zheng M, Nan Z, Shi X-Q (2005) he genus Ceratolejeunea (Lejeuneaceae, Hepaticae) in China. Cryptogamie, Bryologie 26 (1): 91–96. Zhu R-L, Wei Y-M, He Q (2011) Caudalejeunea tridentata, a remarkable new species of Lejeuneaceae (Marchantiophyta) from China. Bryologist 114 (3): 469–473. doi: 10.1639/00072745-114.3.469 Zhu R-L, Wei Y-M, Söderström L, Hagborg A, von Konrat MJ (2013) Notes on Early Land Plants Today. 25. Lejeunea soae, a new name for Lejeunea chinensis, hom. illeg. (Lejeuneaceae, Marchantiophyta). Phytotaxa 81 (1): 1–2. doi: 10.11646/phytotaxa.81.1.1 Zuo B-R, Cao T, Guo S-L (2007a) Comparison and assessment of three East-Asian species of the genus Scapania (Hepaticae: Scapaniaceae). Acta Phytotaxonomica Sinica 45 (5): 742–750. doi: 10.1360/aps06028 Zuo B-R, Cao T, Gao C, Sun J (2007b) Scapania paraphyllia Cao T, Gao C, Sun J, Zuo BR, a new species of Hepaticae (Scapaniaceae) from Zhejiang, China. Acta Phytotaxonomica Sinica 45 (3): 311–314. doi: 10.1360/aps06029 Zwickel W (1933) Zwei neue Gattungen, einige neue Arten und Umstellungen bei den Lejeuneaceen. Annales Bryologici 6: 105–121.

RELATED PAPERS

RELATED TOPICS

Log In

World checklist of hornworts and liverworts

World checklist of hornworts and liverworts

Related Papers

RELATED PAPERS

RELATED TOPICS