Important Plant Areas of Ukraine
Editor: V.A. Onyshchenko
Selecion criteria
УДК 502.75(477)
В12
Aconitum pseudanthora
Błocki ex Pacz.
LC
*
*
-
R Kamiani Mohyly
*
-
-
-
-
*
-
-
-
-
V Medobory; Kniazhpilskyi
Lis; Sovyi Yar; Vasylivskyi
i Rozkopynskyi Yary;
Shebutynskyi Yar; Ternava
– Dnister
V Probabyn, Vymushiv
*
3
RDBU
RBEB
© V.A. Onyshchenko (editor), 2017
© M.G. Kholodny Insitute of Botany, 2017
© Alterpress, 2017
Res 6
ISBN 978-966-542-622-6
Achillea glaberrima
Klokov
Aconitum besserianum
Andrz. ex Trautv.
BC
Reviewers: S.L. Mosyakin, O.M. Bayrak
Taxon
IUCN
db 2015-4
Edited by V.A. Onyshchenko
Co-edited (English text and plant names) by S.L. Mosyakin
Table 1. Criterion A species and corresponding selected areas.
A(iv)
Approved by the Academic Council of the M.G. Kholodny Insitute of Botany,
Naional Academy of Sciences of Ukraine
A(iii)
Authors:
V.A. Onyshchenko, V.P. Kolomiychuk, I.I. Chorney, R.Ya. Kish, A.I. Tokariuk, V.V. Budzhak,
O.O. Orlov, H.I. Oliiar, O.V. Bezrodnova, O.M. Derkach, N.O. Stetsiuk, N.V. Shumska,
M.V. Banik, H.V. Kolomiets, L.H. Lubinska, V.V. Datsiuk, M.M. Peregrym, I.M. Danylyk,
O.M. Popova, O.T. Kuzyarin, O.V. Holovko, O.I. Pryadko, L.T. Horbnyak, V.A. Kolodiy,
L.V. Markivska, O.V. Lukash, N.B. Saidakhmedova, H.P. Protsiv, O.Yu. Umanets,
T.D. Solomakha, V.M. Virchenko, V.V. Rasevich
A(i)
Важливі ботанічні території України / за ред. В.А. Онищенка. – Київ: Альтерпрес,
2017. – 376 с.
ISBN 978-966-542-622-6
Книга містить описи 173 Важливих ботанічних територій України. Дані про кожну
територію включають її площу, географічні координати, критерії виділення, площі
оселищ за класифікацією EUNIS, характеристику рослинності, загрози, види людської
діяльності, інформацію про природно-заповідні території, список літератури і карту на
основі космічної фотографії.
The aim of the Important Plant Areas (IPAs) programme is to idenify and protect
a network of the best sites for plant conservaion throughout Europe and the rest of the
world, using consistent criteria (Anderson, 2002). The ideniicaion of IPAs is based on
three criteria. Criterion A – Presence of threatened plant species: the site holds signiicant
populaions of one or more species that are of global or regional conservaion concern.
Criterion B – Presence of botanical richness: the site has an excepionally rich lora in a
regional context in relaion to its biogeographic zone. Criterion C – Presence of threatened
habitats: the site is an outstanding example of a habitat or vegetaion type of global or
regional plant conservaion and botanical importance. "IPA" is not an oicial designaion.
IPAs are selected scieniically using criteria supported by expert scieniic judgement.
IPA criteria were published in 2001 (Palmer & Smart 2001). Since then IPA were
selected in many countries. In Ukraine, irst six IPAs were ideniied in 2008 (Onyshchenko et
al. 2008). In 2012 16 new areas were selected in the Sea of Azov region (Kolomiychuk et. al.
2012). The present book includes informaion about 173 IPAs ideniied within the territory
of Ukraine. These data are available also in the IPA database online. IPAs were ideniied in
Ukraine mainly using criteria A and C.
Criterion A includes subcriteria A(i) (species with categories EX, CR, En and VU in the
IUCN database version 2015–4), A(ii) (species listed in Appendix I of the Bern Convenion,
Resoluion 6 of the Steering Commitee of the Bern Convenion, or the Red Data Book of
European bryophytes), A(iii) (species with categories "endangered" and "vulnerable" in the
Red Data Book of Ukraine (2009) that are considered naional endemics of Ukraine), A(iv)
(limited range species with categories "endangered" and "vulnerable" in the Red Data Book
of Ukraine (2009) that are not naional endemics). Totally the Criterion A list for Ukraine
has 185 species (table 1): subcriterion A(i) – 23 species, A(ii) – 96, A(iii) – 33, and A(iv) – 65
species. 145 species were used as criteria.
A(ii)
Important Plant Areas of Ukraine / V.A. Onyshchenko (editor). – Kyiv: Alterpress, 2017.
– 376 p.
The book contains descripions of 173 Important Plant Areas of Ukraine. Data on each site
include its area, geographical coordinates, selecion criteria, areas of EUNIS habitat types,
characterizaion of vegetaion, threats, human aciviies, informaion about protected areas, references, and a map on the satelite image background.
IPAs where the taxon is a
criterion of selecion
-
-
-
- Byriuchyi Ostriv;
Arabatska Strilka; Peresyp
Aktaskoho Ozera;
Kazantyp
- Kozachelaherska Arena;
Nyzhniodniprovski Pisky;
Biriuchyi Ostriv; Dnipro
– Orlil
R Zaplava Prypiai; Shatski
Ozera; Dunaiski Plavni
E Molochnyi Lyman;
Kazantyp; Bereh
Syvashyka
R Molochnyi Lyman;
Tendrivska Zatoka;
Askania-Nova; Bereh
Syvashyka
V Nyzhniodniprovski
Pisky; Dnipro – Oril;
Triokhizbenskyi Step;
Volyzhyn Lis
V Skhidnyi Churiuk;
Tendrivska Zatoka
V –
*
Aldrovanda vesiculosa L.
*
Allium pervesitum
Klokov
*
*
*
*
Allium savranicum
(Nyman) Oxner
Allium sphaeropodum
Klokov
Alyssum borzaeanum
Nyár. (Odontarrhena
borzaeana (Nyár.)
D.A.German)
Alyssum gymnopodum
P.A.Smirn. (Odontarrhena
gymnopoda (P.A.Smirn.)
D.A.German)
Alyssum savranicum
Andrz. ex Besser
(Odontarrhena savranica
(Andrz. ex Besser)
D.A.German)
-
-
-
En
*
*
-
En
-
-
-
-
*
Allium scythicum Zoz
En
*
*
*
-
*
-
*
-
-
-
DD
-
-
-
-
-
-
-
-
*
-
-
V Peresyp Aktaskoho Ozera;
Tendrivska Zatoka
*
-
-
-
-
V Kreidiana Flora; Oskilski
Skhyly; Petro-Ivanivka;
Marina Hora
*
-
-
-
-
E Nyzhniodniprovski Pisky;
Kozachelaherska Arena;
Volyzhyn Lis
4
Astragalus setosulus
Gontsch.
Astragalus tanaiticus C.
Koch
Astragalus zingeri Korsh.
Barbula enderesii Garov.
Betula oycoviensis Besser
Botrychium
matricarifolium A.Br. ex
Koch
Botrychium muliidum
(S.G.Gmel.) Rupr.
Brassica taurica (Tzvelev)
Tzvelev
Buxbaumia viridis (Moug.
ex Lam. et DC.) Brid. ex
Moug. et Nestl.
Caldesia parnassifolia
(L.) Parl.
Calophaca wolgarica
(Moug.) Moug. et Nestl.
Campanula abieina
Griseb. et Schenk
RBEB
En
-
E
V Chornohora
-
-
-
-
*
-
*
*
-
E Vovchanski Skyly; Oskilski
Skhyly; Petro-Ivanivka
- –
*
-
-
-
V
V –
*
-
-
-
-
V Krymski Hory; Mehanom;
Tepe-Oba; Karadah
*
-
-
-
-
Vu
*
*
-
E Ak-Monaiskyi Step;
Askania-Nova; Mehanom;
Tepe-Oba
E Krymski Hory; Mehanom
Vu
*
*
-
R Kamiani Mohyly
*
Vu
-
*
-
V
-
E
E
*
-
*
-
-
*
EN
*
*
-
R Chyvchyny; Marmaroski
Hory
V Ayu-Dah; Mys Marian
*
-
*
*
V
- Chornohora
*
LC
*
*
-
E –
LC
-
-
-
V Troitska Balka
-
*
-
-
- Chyvchyny; Marmaroski
Hory; Svydovets;
Chornohora
*
Anacamptodon
splachnoides (Brid.) Brid.
Androsace kosopoljanskii Ovcz.
Angelica palustris
(Besser) Hofm.
Anoectangium handelii
Schiffn.
Astracantha arnacantha
(M.Bieb.) Podlech
(Astragalus arnacantha
M.Bieb.)
Astragalus reduncus Pall.
*
*
*
*
*
*
*
*
*
*
*
*
5
RDBU
- –
Res 6
-
BC
*
IUCN
db 2015-4
-
-
A(iv)
-
-
A(iii)
*
Taxon
A(i)
RBEB
- Sinozhai, Dziurkach
IPAs where the taxon is a
criterion of selecion
A(ii)
Res 6
-
RDBU
BC
*
A(iv)
-
A(ii)
-
A(iii)
*
*
Agropyron dasyanthum
Ledeb.
Allium regelianum
A.Becker ex Iljin
IUCN
db 2015-4
Adenophora lilifolia (L.)
Ledeb.
Agrimonia pilosa
Ledebour
Agropyron cimmericum
Nevski
A(i)
Taxon
IPAs where the taxon is a
criterion of selecion
–
–
–
Chyvchyny; Svydovets;
Chornohora
-
-
*
-
-
-
-
*
Vu
*
*
-
*
-
-
-
-
*
-
-
-
-
*
-
-
-
-
V Kozachelaherska Arena;
Nyzhniodniprovski Pisky
V Krymski Hory
-
-
-
-
V Sviai Hory
*
*
-
-
-
-
*
-
-
-
-
E Kurylivka; Kuchuhury
E Mykhailo-Laryne;
Kovalivka
E Mishkovo-Pohorilove
*
-
-
-
-
-
-
-
-
E Bobrovyi Kut;
Novohredneve
E –
-
-
-
-
E Halitsynove
Ex
*
*
-
R Kamiani Mohyly
Carlina cirsioides Klokov
Carlina onopordifolia
Besser ex Szafer, Kulcz.
et Pawł.
Centaurea appendicata
Klokov
Centaurea breviceps Iljin
*
Centaurea comperiana
Steven
Centaurea donetzica
Klokov
Centaurea konkae Klokov
Centaurea margaritaalba Klokov
Centaurea margaritacea *
Ten.
Centaurea paczoskii
Kotov ex Klokov
Centaurea protogerberi
Klokov
Centaurea
protomargaritacea
Klokov
Centaurea
*
pseudoleucolepis
Kleopow
Centaurea sarandinakiae
N.B.Illar.
Centaurea taliewii
Kleopow (Rhaponicoides
taliewii (Kleopow)
M.V.Agab. & Greuter)
*
*
*
*
*
V Dunaiski Plavni; Nyzhnii
Dnipro; Hrakove
V Chortova Hora; Kasova
Hora; Horodnytski Tovtry
V Velyki Holdy; Lysa Hora i
Sinka; Vasylivka; Pidlyska
Hora
E Lysohirka
*
-
-
-
-
V Krymski Hory; Karadah
*
-
-
-
-
V Tarkhankut; Askania-Nova
6
RBEB
*
-
-
-
E Krymski Hory; Karadah
-
-
-
-
E –
*
-
-
-
-
*
-
-
-
-
*
-
-
-
-
*
-
-
-
-
V Hranitno-Stepove
Pobuzhia
V Yelanetskyi Step;
Mykhailivskyi Step; Zelena
Balka
V Kadubivska Sinka;
Probabyn; Medobory
V Krymski Hory
*
*
En
*
*
-
E Kolivska Ulohovyna
*
*
LC
*
*
-
-
*
-
-
V Skhyly Kohylnyka;
Kuchurhan
E Krymski Hory
DD
-
-
-
DD
*
*
-
*
-
-
-
-
*
-
-
-
-
*
-
-
-
-
-
-
*
-
-
-
-
-
V Osovynskyi Step; ТепеОба; Karadah
V Khomutovskyi Step;
Mykhailivskyi Step
V Karadah
-
*
*
-
E –
E Krymski Hory
*
Chamaecyisus graniicus
(Rehmann.) Rothm.
Chamaecyisus podolicus
(Błocki) Klásková
Chamaecyisus wulii
(V.Krecz.) Klásková
Cochlearia polonica
*
Frohl.
Colchicum fominii Bordz.
*
Comperia comperiana
(Steven) Asch. et Graebn.
Crambe aspera M.Bieb.
Crambe grandilora DC.
*
Crambe koktebelica
(Junge) N.Busch
Crambe mitridais Juz.
*
*
*
Crambe pinnaiida
W.T.Aiton
Crambe steveniana Rupr.
Crambe tataria Sebeok
Crataegus pojarkovae
Kossych
Crocus banaicus Gay
Cyclamen kuznetzovii
Kotov et Czernowa
*
*
*
*
*
7
RDBU
-
*
BC
-
Res 6
-
-
Cephalaria demetrii
Bobrov
Cephalaria litvinovii
Bobrov
Cerasus klokovii Sobko
A(iv)
-
IUCN
db 2015-4
-
*
Caragana scythica
(Kom.) Pojark.
Carex secalina Wahlenb.
- Chyvchyny; Marmaroski
Hory; Chornohora;
Dzhohul
V Korsak-Mohyla
A(iii)
-
Taxon
A(i)
RBEB
*
IPAs where the taxon is a
criterion of selecion
A(ii)
Res 6
-
*
RDBU
BC
LC
A(iv)
A(ii)
IUCN
db 2015-4
Campanula serrata (Kit.)
Hendrych
A(iii)
A(i)
Taxon
IPAs where the taxon is a
criterion of selecion
V Tepe-Oba; Opuk; Karadah
V Osovynskyi Step;
Karalarskyi Step;
Kazantyp; Ak-Monaiskyi
Step; Karadah
R Tepe-Oba; Opuk; Karadah
V Ak-Monaiskyi Step;
Kazantyp; Karalarskyi
Step; Tarkhankut; Opuk
V Kazantyp; Karadah
-
-
-
*
-
*
*
-
*
-
-
-
V
- –
*
-
-
-
-
E Dunaiski Plavni
*
Vu
*
*
-
Dianthus
pseudoseroinus Błocki
Dicranum viride (Sull. et
Lesq.) Lindb.
Diplotaxis cretacea Kotov
-
-
-
-
-
*
*
V
-
-
-
-
Dracocephalum
austriacum L.
Dracocephalum
ruyschiana L.
Echium russicum
G.F.Gmel.
Eleocharis carniolica
W.D.J.Koch
Eremurus tauricus Steven
Erysimum hungaricum
*
Zapał.
Erysimum krynkense
Lavrenko
Erysimum ucrainicum
J.Gay
*
-
*
*
-
*
-
*
-
-
*
-
-
*
-
V Dolyna Mertvovodu;
Hranitno-Stepove
Pobuzhia; Dolyna Inhulu
V Levkivski Lisy; Shatski
Ozera
- Chyvchyny; Marmaroski
Hory; Chornohora
V Striltsivskyi Step;
Kreidiana Flora; Oskilski
Skhyly; Marina Hora
V Horodnytski Tovtry;
Hostra Skelia
I Potashnianski Lisy;
Kolivska Ulohovyna
- –
*
LC
*
*
-
V –
Vu
-
-
-
E Krymski Hory; Karadah
- –
*
-
-
-
-
E –
*
-
-
-
-
V Oskilski Skhyly;
Balakyrivka
*
*
*
*
*
8
Genista tetragona Besser *
Gladiolus palustris Gaud.
Goniolimon
graminifolium (Ait.)
Boiss.
Goniolimon rubellum
(S.G.Gmel.) Klokov
*
*
*
-
-
*
DD
*
*
-
*
Vu
DD
-
*
-
*
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
*
*
-
*
-
-
-
-
E Kreidiani Vidslonennia
*
-
-
-
-
E Novobila; Sharivka
-
-
-
E
R Svydovets; Chornohora
*
-
*
*
-
V Krymski Hory; Karadah
*
-
-
-
-
V Naholnyi Kriazh
*
-
-
*
-
V Sviai Hory
-
-
*
-
- Irpinskyi Lis, Ternava –
Dnister; Pohorylivka;
Kadubivska Sinka
*
*
*
Gymnospermium
odessanum (DC.) Takht.
*
Gypsophila thyraica
Krasnova
Hamatocaulis
verniculosus (Mit.)
Hedenas
Hedysarum cretaceum
Fisch.
Hedysarum ucrainicum
Kaschm.
Heterophyllium affine
(Mitt.) Fleisch.
Himantoglossum
caprinum (M.Bieb.)
C.Koch
Hyacinthella pallasiana
(Steven) Losinsk.
Iris humilis Georgi (Iris
pineicola Klokov)
Iris hungarica Waldst.
& Kit.
-
*
*
*
*
*
9
RDBU
*
*
-
*
*
RBEB
Daphne taurica Kotov
Delphinium rossicum Litv.
Delphinium sergii
Wissjul.
Dendranthema zawadskii
(Herbich) Tzvelev
Desmatodon cernuus
(Hüb.) B. et S.
Dianthus bessarabicus
Klokov
Dianthus hypanicus
*
Andrz.
Festuca porcii Hack.
Fritillaria montana
Hoppe
Res 6
-
BC
-
IUCN
db 2015-4
-
*
A(iv)
En
*
Ferula orientalis L.
A(iii)
*
V Romosh; Tsetsyno,
Krymski Hory; Pidlyska
Hora
E Vovchanski Skyly;
Kolodiazne
E Krymski Hory
V Dnipro-Oril
V Provalskyi Step; Dolyna
Mertvovodu
R –
Taxon
A(i)
RBEB
-
IPAs where the taxon is a
criterion of selecion
A(ii)
Res 6
*
*
RDBU
BC
*
A(iv)
IUCN
db 2015-4
Daphne sophia Kalen.
-
A(iii)
A(i)
Cypripedium calceolus L.
A(ii)
Taxon
IPAs where the taxon is a
criterion of selecion
- Molochnyi Lyman;
Askania-Nova
V Chyvchyny; Chornohora
E Shebutynskyi Yar;
Vasylivskyi i Rozkopynskyi
Yary; Kaplivka; Podvirivka
E Trosianets
E –
V Nyzhniodniprovski Pisky;
Volyzhyn Lis
V Arabatska Strilka;
Mytrofanivskyi Pivostriv;
Bereh Syvashyka
V Hranitno-Stepove
Pobuzhia, Mykhailivskyi
Step
V Verbetska Tovtra;
Kadubivska Sinka;
Tovtrivska Sinka;
Pohorylivka; Probabyn
- Marmaroski Hory;
Svydovets; Chornohora
-
-
-
-
E Kedryn
DD
-
-
-
Cr
*
*
-
V Arabatska Strilka; Skhidnyi
Churiuk; Mytrofanivskyi
Pivostriv
E Tepe-Oba
-
-
*
-
V Chyvchyny; Bushchanske
Boloto
-
-
-
-
V Tiup-Tarkhan i Kalynivskyi
Pivostriv
*
Lindernia procumbens
(Krock.) Borb.
Liparis loeselii (L.) Rich.
*
LC
*
-
-
*
-
*
*
-
Mannia triandra (Scop.)
Grolle
Marsilea quadrifolia L.
Medicago kotovii Wissjul. *
Medicago saxailis
*
M.Bieb.
Meesia longiseta Hedw.
Moehringia hypanica
*
Grynj et Klokov
*
-
-
*
-
*
LC
Vu
En
-
*
-
-
Vu
*
-
*
*
-
-
*
R Ex –
- R Dolyna Mertvovodu;
Hranitno-Stepove
Pobuzhia
- - –
*
*
*
*
*
*
-
10
-
-
-
V Kreidiana Flora;
Riznykivka; Balakyrivka;
Skhyly Krasnoi i Kobylky
- Dnipro – Oril
V Bushchanske Boloto; Bir
na Merli
- Chornohora; Uholka –
Shyrokyi Luh
V –
- Byriuchyi Ostriv
I Krymski Hory
RBEB
V Krymski Hory
*
-
Neckera pennata Hedw.
*
-
-
-
V
-
-
-
-
-
-
-
-
*
-
-
-
-
*
-
-
-
-
*
-
*
*
-
*
-
*
-
-
*
-
*
-
-
*
*
-
*
*
-
-
-
-
-
-
E Krymski Hory
E Krymski Hory; Tepe-Oba;
Karadah
E –
*
-
-
-
E
- –
*
-
*
*
-
*
-
-
-
V
V Krymski Hory;
Khomutovskyi Step;
Striltsivskyi Step; Vitrohon
- –
*
-
-
-
-
E Kolivska Ulohovyna
*
-
-
-
-
V –
Nigritella carpaica
(Zapał.) Teppner, Klein
et Zagulski (Gymnadenia
carpaica (Zapał.)
Teppner & E.Klein)
Onobrychis pallasii
(Willd.) M.Bieb.
Onobrychis vassilczenkoi
Grossh.
Onosma graniicola
Klokov
Onosma polyphylla
Ledeb.
Ophrys oestrifera
M.Bieb.
Ophrys taurica
(Aggeenko) Nevski
Orchis provincialis Balb.
Orchis punctulata Steven
ex Lindl.
Ornithogalum
amphibolum Zahar.
Orthotrichum scanicum
Gronv.
Paeonia tenuifolia L.
Pallavicinia lyelii (Hook.)
Carruth.
Pinguicula bicolor Woł.
(Pinguicula vulgaris L.
ssp. bicolor (Woł.) A. et
D.Löve)
Pinus cretacea (Kalen.)
Kondr.
*
*
*
*
11
RDBU
-
*
A(iv)
*
-
A(iii)
*
*
A(i)
Vu
Narcissus angustifolius
Curt.
Taxon
A(ii)
- –
RDBU
-
Lepidium turczaninowii
*
Lipsky
Ligularia sibirica (L.)
Cass. (L. bucovinensis
Nakai)
Limonium tschurjukiense
(Klokov) Lavrenko ex
Klolkov
Linaria cretacea Fisch. ex
Spreng.
Moehringia lateriflora
(L.) Fenzl.
*
*
Res 6
*
-
BC
*
-
IPAs where the taxon is a
criterion of selecion
IUCN
db 2015-4
*
RBEB
*
Res 6
*
BC
A(iv)
A(ii)
A(iii)
*
IUCN
db 2015-4
Jurinea cyanoides (L.)
Rchb.
Lagoseris purpurea
(Willd.) Boiss. (Crepis
purpurea (Willd.)
M.Bieb.)
Larix decidua Mill.
subsp. polonica (Racib.)
A.E.Murray
Lepidium syvaschicum
Kleopow
A(i)
Taxon
IPAs where the taxon is a
criterion of selecion
V Dolyna Nartsysiv;
Marmaroski Hory;
Svydovets
- Svydovets; Chornohora;
Uholka – Shyrokyi Luh
E Chyvchyny; Dzhohul
V Krymski Hory; Mehanom;
Tepe-Oba; Karadah
V –
E Provalskyi Step; HranitnoStepove Pobuzhia
V Krymski Hory; Mehanom;
Karadah
E Krymski Hory; Kazantyp;
Mys Marian; Tepe-Oba;
Karadah
E Krymski Hory
*
-
-
*
-
- Marmaroski Hory;
Svydovets
*
-
-
-
V
R –
*
-
-
-
V
- –
*
LC
*
*
-
Pulsailla patens (L.) Mill.
(P. laifolia Rupr.)
Pyramidula tetragona
(Brid.) Brid.
Ranunculus lapponicus L.
Rheum rhaponicum L.
Rhus coriaria L.
*
Rhododendron luteum
Sweet
Salvinia natans (L.) All.
*
-
*
*
-
V Sinozhai, Horodnytski
Tovtry; Kasova Hora;
Horaivka; Pohorylivka
I Levkivski Lisy; Sinozhai
*
-
-
-
V
- –
*
*
*
Vu
-
-
*
*
*
-
-
*
I
*
-
-
Saxifraga hirculus L.
Schivereckia podolica
(Besser) Andrz. & Besser
ex DC. (Draba podolica
(Besser) Rupr.)
Senecio besserianus
Minder.
Serratula lycopifolia
(Vill.) A.Kern. (Klasea
lycopifolia (Vill.) Á.Löve
& D.Löve)
Serratula tanaitica
P.Smirn.
*
*
LC
*
*
*
*
-
*
-
-
–
–
Krymski Hory; Karadah
Olevski Lisy; Yemilchynski
Lisy
I Dunaiski Plavni; Nyzhnii
Dnipro
V Bushchanske Boloto
I Hostra Skelia; Medobory;
Usia – Shutnivtsi; Ternava
– Dnister; Marina Hora
-
-
V –
*
DD
*
*
-
- Podvirivka, Vyshnivka
*
-
*
*
-
R –
12
*
-
*
*
-
RDBU
RBEB
*
Res 6
Silene cretacea Fisch. ex
Spreng.
BC
V Krymski Hory
IUCN
db 2015-4
-
A(iv)
-
A(iii)
RBEB
-
Taxon
A(i)
Res 6
Vu
IPAs where the taxon is a
criterion of selecion
A(ii)
BC
*
RDBU
IUCN
db 2015-4
A(ii)
A(iv)
Pinus stankewiczii
*
(Sukacz.) Fomin (P. bruia
Ten. var. pityusa (Steven)
Silba; P. pityusa Steven
var. stankewiczii Sukacz.)
Poa graniica BraunBlanq. (Poa deylii Chrtek
et V.Jirasek)
Pterygoneurum kozlovii
Lazar.
Pterygoneurum
lamellatum (Lindb.) Jur.
Pulsatilla grandis Wend.
A(iii)
A(i)
Taxon
IPAs where the taxon is a
criterion of selecion
V Kreidiana Flora;
Riznykivka; Skhyly Krasnoi
i Kobylky
V Hranitno-Stepove
Pobuzhia
V Krymski Hory
Silene hypanica Klokov
*
-
-
-
-
Silene jailensis N.I.
Rubtzov
Silene sytnikii Krytzka,
Novosad et Protopopova
*
-
-
-
-
*
-
-
-
-
DD
*
-
-
LC
-
-
*
-
-
-
*
*
-
- –
E Zhyzhava; Ternava –
Dnister
E Krymski Hory
*
-
-
-
-
E –
*
*
-
*
*
-
-
-
*
-
-
-
*
*
*
*
-
V Karadah
V Mehanom; Tepe-Oba;
Karadah
I Kreidiana Flora; Vitrohon
V Kreidiana Flora; Marina
Hora
V –
- –
-
-
-
-
Sisymbrium confertum
Steven ex Turcz.
Sisymbrium supinum L.
Spiraea polonica Błocki
Steveniella satyrioides
(Steven) Schlechter
Sipa anomala
P.A.Smirn.
Sipa poëica Klokov
Stipa syreistschikowii
P.A.Smirn.
Stipa zalesskii Wilensky
Syrenia talievi Klokov
Syringa josikaea Jacq. fil.
Thesium ebracteatum
Hayne
Thymus litoralis Klokov
et Des.-Shost.
*
*
*
*
*
*
*
*
*
*
*
Tilia dasystyla Steven
Tortula handelii Schiffn.
Tozzia carpathica Woł.
*
*
DD
-
*
V
-
Trapa natans L.
*
LC
*
-
-
*
13
V Dolyna Mertvovodu;
Hranitno-Stepove
Pobuzhia
- Krymski Hory
V Karalarskyi Step;
Arabatska Strilka; Peresyp
Aktaskoho Ozera; Opuk
E Krymski Hory; Karadah
- –
- Chyvchyny; Marmaroski
Hory; Chornohora
I Dunaiski Plavni; Nyzhnii
Dnipro
BC
Res 6
RBEB
-
Tulipa scythica Klokov
et Zoz
Typha minima Funk
Typha shutleworthii
W.D.J.Koch et Sond.
Ulota rehmanii Jur.
Zostera marina L.
*
-
-
-
-
V Hranitno-Stepove
Pobuzhia; Yelanetskyi
Step; Mykhailivskyi Step;
Dolyna Inhulu; Zelena
Balka
V Askania-Nova
*
*
-
*
*
-
-
E Dunaiski Plavni
- Chyvchyny
*
*
LC
*
-
E
-
- –
- Molochnyi Lyman;
Biriuchyi Ostriv; Arabatska
Strilka; Obyichna Kosa
RDBU
IUCN
db 2015-4
-
A(iv)
-
A(ii)
-
A(iii)
*
A(i)
Tulipa hypanica Klokov
et Zoz
Taxon
IPAs where the taxon is a
criterion of selecion
Abbreviaions: IUCN db 2015 – version 2015-4 of the IUCN database, BC – Appendix I of the Bern
Convenion, Res6 - Resoluion 6 of the Bern Convenion (1998, year of revision 2011), RBEB – Red
Data Book of European bryophytes (1995), RDBU – Red Data Book of Ukraine (2009).
Note. The scientific names of plants are given mostly according to the checklist of vascular plants of
Ukraine (Mosyakin & Fedoronchuk 1999), with necessary taxonomic and nomenclatural adjustments.
However, to avoid confusion, we mainly preserved as "main" names of protected species those
names which are listed in the official documents mentioned above, even in the cases when new
taxonomic and phylogenetic data dictate changes in their nomenclature. For example, some species
of Centaurea are currently placed in segregate genera (e.g., Rhaponticoides, Psephellus), some species
of Alyssum were recently transferred to Odontarrhena, etc. In all such cases, the names currently
accepted by taxonomists are provided after the names used in the IUCN database, Appendix I of
the Bern Convention, Resolution 6 of the Steering Committee of the Bern Convention, the Red Data
Book of Ukraine, the Red Data Book of European bryophytes, or in other relevant red lists. Since
the present book is not a taxonomic treatment, such usage of plant names should not be viewed as
endorsement of alternative taxonomic or nomenclatural decisions.
Criterion B was applied only for the EUNIS level 2 habitat G1 Broadleaved deciduous woodland. The indicator species checklist of this habitat contains 77 species. All these
species have their opimum in this habitat type and constancy <3% in vegetaion data plots
of deciduous woods. Most of them are species with small extent of occurence. Analysis of
distribuion of indicator species number in deciduous forests of Ukraine shows that the richest 10% of forests are Crimean forests and some forests of the western part of Ukraine with
calcareous rocks (Onyshchenko et al. 2008). Six areas have been selected by this criterion:
Krymski Hory, Uholka – Shyrokyi Luh, Verkhniobuzki Lisy, Medobory, Karadah, and Mys Marian. Selected areas contain 6% of the habitat G1 in Ukraine.
1.
2.
3.
4.
5.
6.
7.
8.
9.
10.
11.
12.
13.
14.
15.
16.
17.
18.
19.
20.
21.
22.
23.
24.
25.
26.
27.
28.
29.
30.
31.
32.
14
Checklist of species for assessment of the botanical richness of habitats type
G1: Broadleaved deciduous woodland
Taxaceae
Taxus baccata L.
Ranunculaceae
Aconitum besserianum Andrz. ex Trautv.
Aconitum degenii Gayer
Aconitum gracile (Rchb.) Gayer
Aconitum lasiostomum Rchb.
Aconitum nemorosum M.Bieb. ex Rchb.
Aconitum variegatum L.
Aquilegia vulgaris L
Cimicifuga europaea Schipcz. (Actaea europaea (Schipcz.) J.Compton)
Delphinium pallasii Nevski
Helleborus purpurascens Waldst. et Kit.
Uricaceae
Urica kioviensis Rogov.
Caryophyllaceae
Cerasium sylvaicum Waldst. et Kit.
Brassicaceae
Cardamine tenera S.G. Gmel. ex C.A. Mey.
Hesperis candida Kit. ex Müggenb., Kanitz et Knapp
Hesperis matronalis L.
Hesperis steveniana DC.
Hesperis sibirica L.
Hesperis voronovii N.Busch
Primulaceae
Cyclamen kuznetzovii Kotov et Czernowa s.str. (Cyclamen coum Mill. s.l.)
Lysimachia vericillaris Spreng.
Thymeleaceae
Daphne sophia Kalen.
Daphne taurica Kotov
Tiliaceae
Tilia dasystyla Stev.
Fabaceae
Lathyrus laevigatus (Waldst. et Kit.) Fritsch
Lathyrus transsilvanicus (Spreng.) Rchb.
Lathyrus venetus (Mill.) Wohlf.
Rutaceae
Dictamnus gymnostylis Stev.
Aceraceae
Acer stevenii Pojark.
Apiaceae
Laserpiium laifolium L.
Celastraceae
Euonymus nanus M.Bieb.
Rhamnaceae
Rhamnus inctoria Waldst. et Kit.
15
33.
34.
35.
36.
37.
38.
39.
40.
41.
42.
43.
44.
45.
46.
47.
48.
49.
50.
51.
52.
53.
54.
55.
56.
57.
58.
59.
60.
61.
62.
63.
64.
65.
66.
67.
68.
Santalaceae
Thesium ebracteatum Hayne
Oleaceae
Syringa josikaea Jacq. il.
Rubiaceae
Asperula propinqua Pobed.
Boraginaceae
Solenanthus biebersteinii DC.
Solanaceae
Scopolia carniolica Jacq.
Scrophulariaceae
Scrophularia vernalis L.
Campanulaceae
Adenophora liliifolia (L.) A. DC.
Liliaceae
Colchicum umbrosum Steven
Erythronium dens-canis L.
Friillaria ruthenica Wikstr.
Nectaroscordum meliophilum (Juz.) Zahar. (Allium siculum Ucria subsp.
dioscoridis (Sm.) K.Richt.)
Ruscus hypoglossum L.
Veratrum nigrum L.
Amaryllidaceae
Galanthus elwesii Hook. il
Leucojum vernum L.
Iridaceae
Crocus banaicus J. Gay
Iris graminea L.
Iris hungarica Waldst. et Kit.
Iris variegata L.
Orchidaceae
Anacampis pyramidalis (L.) Rich.
Comperia comperiana (Steven) Asch. et Graebn.
Corallorhiza triida Châtel.
Cypripedium calceolus L.
Dactylorhiza romana (Seb. et Mauri) Soó
Epipacis atrorubens (Hofm ex Bernh.) Schult.
Epipacis microphylla (Ehrh.) Sw.
Epipacis purpurata Smith
Epipogium aphyllum (F.W.Schmidt) Sw.
Himantoglossum caprinum (M.Bieb.) K.Koch
Limodorum aborivum (L.) Sw.
Ophrys apifera Huds.
Ophrys oestrifera M.Bieb.
Ophrys taurica (Aggeenko) Nevski
Orchis mascula (L.) L.
Orchis picta Loisel.
Orchis provincialis Balb.
16
69.
70.
71.
72.
73.
74.
75.
76.
77.
Orchis punctulata Stev. ex. Lindl.
Orchis purpurea Huds.
Orchis simia Lam.
Steveniella satyrioides (Steven) Schltr.
Cyperaceae
Carex depauperata Curt. ex With.
Carex strigosa Huds.
Poaceae
Festuca drymeia Mert. et Koch
Araceae
Arum albispathum Steven ex Ledeb.
Arum orientale M.Bieb.
Criterion C in our analysis was the presence of habitats from Resluion 4 of the
Steering Commitee of the Bern Convenion, 2014 version (table 2). In addiion, one
priority habiatat from the Habitats Direcive was used: 4070 Bushes with Pinus mugo and
Rhododendron hirsutum (Mugo-Rhododendretum hirsui). The Ukrainian subtype of this
habitat (F2.46) is not present in Resoluion 4. For most habitat types we selected maximum
ive areas. Main excepions are some habitats correspoding to the priority habitats of the
Habiatats Direcive: E1.2 Perennial calcareous grassland and basic steppes (includes 62C0
Ponto-Sarmaic steppes and 6240 Sub-Pannonic steppic grasslands), F3.247 Ponto-Sarmatic
deciduous thickets (includes priority habitat type 40C0 Ponto-Sarmatic deciduous thickets),
G1.6 Fagus woodland (includes priority habitat type 9150 Medio-European limestone
beech forests of the Cephalanthero-Fagion), G1.7 Thermophilous deciduous woodland
(includes priority habitat types 91H0 Pannonian woods with Quercus pubescens and 91I0
Euro-Siberian steppic woods with Quercus spp.). The largest number of IPAs are selected for
habitat E1.2. Former (and potenial) area of habitat E1.2 in Ukraine is a few tens of imes
larger than its current area. At present, the habitat is highly fragmented so ive best sites
include only 10% of its area in Ukraine. Besides, ive sites cannot represent the geographical
and ecological diversity paterns of steppes.
Table 2. Criterion C habitats and corresponding areas.
Code
A1.11
A1.22
A1.44
A2.2
A2.3
A2.4
A2.5
Name in Resoluion 4
IPAs where the habitat is a
criterion of selecion
Mussel and/or barnacle
–
comminiies
Mussels and fucoids on moderately –
exposed shores
Communiies of litoral caves and
overhangs
Litoral sand and muddy sand
Littoral mud
Littoral mixed sediments
Coastal saltmarshes and saline
reedbeds
17
–
–
–
–
Molochnyi Lyman; Dzharylhach;
Tendrivska Zatoka
Code
A2.61
A3
A4
A5
B1.1
B1.3
B1.4 (*2130)
B1.8
B2.1
B2.3
B3.3
C1.1
C1.222
C1.223
C1.224
C1.225
C1.226
C1.25
C1.32
C1.33
C1.3411
Name in Resoluion 4
IPAs where the habitat is a
criterion of selecion
Seagrass beds on litoral sediments
Infralitoral rock and other hard
substrata
Circalitoral rock and other hard
substrata
Sublittoral sediment
–
Tarkhankut; Mehanom; Tepe-Oba;
Opuk
–
Tendrivska Zatoka; Dzharylhach;
Opuk; Mehanom
Sand beach dritline
Arabatska Strilka; Biriuchyi Ostriv;
Obyichna Kosa
Shifting coastal dunes
Tendrivska Zatoka; Dzharylhach;
Arabatska Strilka; Biriuchyi Ostriv
Coastal stable dune grassland (grey Tendrivska Zatoka; Dzharylhach;
dunes)
Arabatska Strilka
Moist and wet dune slacks
–
–
Shingle beach driftlines
Upper shingle beaches with open –
vegetaion
Rock clifs, ledges and shores, with Kazantyp; Krymski Hory;
angiosperms
Mehanom; Tepe-Oba; Opuk
Permanent oligotrophic lakes,
Shatski Ozera; Bile Ozero;
ponds and pools
Svydovets; Chornohora
Floaing Hydrocharis morsus-ranae –
rats
Floating Stratiotes aloides rafts
Zaplava Prypiai; Zaplava Desny;
Zaplavy Dnipra i Sozha
Floaing Utricularia australis and
Zaplava Desny; Dunaiski Plavni;
Utricularia vulgaris colonies
Zaplavy Dnipra i Sozha; Nyzhnii
Dnipro
Floating Salvinia natans mats
Dunaiski Plavni; Nyzhnii Dnipro
Floating Aldrovanda vesiculosa
Zaplava Prypiai; Shatski Ozera;
communities
Dunaiski Plavni
Charophyte submerged carpets in Shatski Ozera
mesotrophic waterbodies
Free-loaing vegetaion of
–
eutrophic waterbodies
Rooted submerged vegetaion of
–
eutrophic waterbodies
Ranunculus communiies in shallow Zaplava Desny; Dunaiski Plavni;
water
Nyzhnii Dnipro; Zaplavy Dnipra i
Sozha
18
Code
C1.3413
C1.44
C1.5
C1.66
C1.67
C2.12 (*7220)
C2.18
C2.19
C2.1A
C2.1B
C2.25
C2.26
C2.27
C2.28
C2.33
C2.34
C3.4
C3.51
C3.55
C3.62
Name in Resoluion 4
Hotonia palustris beds in shallow
water
Charophyte submerged carpets in
dystrophic waterbodies
Permanent inland saline and
brackish lakes, ponds and pools
Temporary inland saline and
brackish waters
Turlough and lake-botom meadows
Hard water springs
Acid oligotrophic vegetaion of
spring brooks
Lime-rich oligotrophic vegetaion of
spring brooks
Mesotrophic vegetaion of spring
brooks
Eutrophic vegetaion of spring
brooks
Acid oligotrophic vegetaion of fastlowing streams
Lime-rich oligotrophic vegetaion of
fast-lowing streams
Mesotrophic vegetaion of fastlowing streams
Eutrophic vegetaion of fast-lowing
streams
Mesotrophic vegetaion of slowlowing rivers
Eutrophic vegetaion of slowlowing rivers
Species-poor beds of low-growing
water-fringing or amphibious
vegetaion
Euro-Siberian dwarf annual
amphibious swards (but excluding
C3.5131 Toad-rush swards)
Sparsely vegetated river gravel
banks
Unvegetated river gravel banks
19
IPAs where the habitat is a
criterion of selecion
–
–
–
–
–
Krymski Hory; Chyvchyny;
Marmaroski Hory; Chornohora
–
–
–
–
–
–
–
Zaplava Tysy
Zaplava Desny; Zaplavy Dnipra i
Sozha
Nyzhnii Dnipro; Dunaiski Plavni;
Zaplava Tysy
Zaplava Desny; Zaplavy Dnipra i
Sozha; Kanivski Ostrovy; Zaplava
Tysy; Ubort – Bolotnytsia
Zaplava Desny; Zaplavy Dnipra i
Sozha; Kanivski Ostrovy; Zaplava
Tysy
–
–
Code
Name in Resoluion 4
D2.226
Peri-Danubian black-white-star
sedge fens
D2.3
Transiion mires and quaking bogs
D4.1
D5.2
D6.1 (*1340)
E1.11 (*6110)
E1.12
E1.13
E1.2 (incl.
*6240, *62C0)
E1.2 (6210,
*Important
orchid sites)
E1.3 (*6220)
IPAs where the habitat is a
criterion of selecion
Chyvchyny, Chornohora; Svydovets;
Marmaroski Hory
Perebrody; Somyno; Syra Pohonia;
Bile Ozero; Ubort – Bolotnytsia;
Cheremske Boloto
Rich fens, including eutrophic tall- Pecheniiske Boloto; Zolochivka;
herb fens and calcareous lushes
Chyvchyny; Bushchanske Boloto;
and soaks
Kolivska Ulohovyna; Markovychi
Beds of large sedges normally
Zaplava Desny; Zaplava Prypiai;
without free-standing water
Zaplavy Dnipra i Sozha; Nyzhnii
Dnipro; Dunaiski Plavni
Inland saline and brackish marshes Tendrivska zatoka
and reedbeds
Euro-siberian pioneer rock debris Ternava – Dnister; Zhyzhava;
swards
Hranitno-Stepove Pobuzhia;
Dolyna Inhulu; Dolyna Mertvovodu
Euro-Siberian pioneer calcareous
–
sand swards
Coninental dry rocky steppic
Kreidiana Flora; Novobila; Oskilski
grasslands and dwarf scrub on
Skhyly; Vovchanski Skyly
chalk outcrops
Perennial calcareous grassland and Krymski Hory; Tarkhankut;
basic steppes
Askania-Nova; Karalarskyi Step;
Medvezhanka; Osovynskyi Step;
Naholnyi Kriazh; Kalmiuskyi
Step; Novobila; Polovetskyi Step;
Oskilski Skhyly, Mykhailivskyi Step;
Yelanetskyi Step; Davydo-Mykilske;
Hranitno-Stepove Pobuzhia;
Zelena Balka; Khomutovskyi Step;
Dzharylhach; Dolyna Mertvovodu;
Dobrianski Hory; Obyichna Kosa;
Skhidnyi Churiuk; Striltsivskyi
Step; Mehanom; Tepe-Oba;
Vitrohon; Provalskyi Step; Ternava
– Dnister; Tulyntsi – Makedony;
Karadah; Vyshnivka; Kasova Hora;
Pohorylivka; Chorna Hora; Yulivski
Hory
Perennial calcareous grassland and Zholoby; Dubovetski Skhyly
basic steppes (Important orchid
sites)
Mediterranean xeric grassland
–
20
Code
Name in Resoluion 4
E1.71 (overlaps Nardus stricta swards
with *6230)
E1.9
Open non-Mediterranean dry acid
and neutral grassland, including
inland dune grassland
E2.2
E2.3
E3.4
E3.5
E4.11
E4.12
E4.3 (overlaps
with *6230)
E4.4
E5.4
E5.5
E6.2
F2.22
F2.26
F2.46 (*4070)
F3.16
F3.241 (*40A0)
IPAs where the habitat is a
criterion of selecion
Polonyna Borzhava; Chyvchyny
Nyzhniodniprovski Pisky;
Triokhizbenskyi Step;
Kozachelaherska Arena; Dnipro –
Oril
Low and medium alitude hay
Zaplava Desny; Hadiatski Luky;
meadows
Zaplavy Dnipra i Sozha; Zaplava
Siverskoho Dintsia; Stuzhytsia
Mountain hay meadows
Chyvchyny
Moist or wet eutrophic and
Zaplava Desny; Zaplavy Dnipra
mesotrophic grassland
i Sozha; Zaplava Prypiai;
Kremenchutski Plavni
Moist or wet oligotrophic grassland Zaplava Prypiai; Zaplava Desny;
Zaplavy Dnipra i Sozha; Dubrova
Boreo-alpine acidocline snow-patch Svydovets; Chornohora; Polonyna
grassland and herb habitats
Borzhava, Marmaroski Hory
Boreo-alpine calcicline snow-patch Svydovets; Chornohora
grassland and herb habitats
Svydovets; Chornohora;
Acid alpine and subalpine grassland Marmaroski Hory; Polonyna
Borzhava, Chyvchyny
Calcareous alpine and subalpine
Chornohora; Svydovets;
grassland
Chyvchyny; Marmaroski Hory;
Moist or wet tall-herb and fern
Zaplava Desny; Kanivski Ostrovy;
fringes and meadows
Zaplavy Dnipra i Sozha; Zaplava
Tysy
Subalpine moist or wet tall-herb
Chornohora; Chyvchyny;
and fern stands
Svydovets; Marmaroski Hory
Continental inland salt steppes
Tendrivska Zatoka; Molochnyi
Lyman; Dunaiski Plavni; Hrakove;
Samarskyi Lis
Alpide acidocline Rhododendron
Svydovets; Chornohora;
heaths
Marmaroski Hory; Chyvchyny
Bruckenthalia heaths
–
Carpathian Pinus mugo scrub
Chornohora; Chyvchyny; Grofa;
Gorgany; Ihrovets - Tavpishyrka
Medobory
Juniperus communis scrub
Central European subcontinental
Chorna Hora
thickets
21
Code
F3.247 (*40C0)
F4.2
F5.13
F7
F9.1
Name in Resoluion 4
IPAs where the habitat is a
criterion of selecion
Krymski Hory; Striltsivskyi Step;
Hranitno-Stepove Pobuzhia;
Ternava – Dnister; Osovynskyi
Ponto-Sarmatic deciduous thickets
Step; Kreidiana Flora; Karalarskyi
Step; Kamiani Mohyly; Dolyna
Mertvovodu; Zhyzhava
Dry heaths
Polonyna Borzhava; Chyvchyny;
Svydovets; Chornohora
–
Juniper matorral
Spiny Mediterranean heaths
(phrygana, hedgehog-heaths and
related coastal clif vegetaion)
Riverine scrub
F9.3
Southern riparian galleries and
thickets
G1.11 (overlaps Riverine Salix woodland
with *91E0)
Name in Resoluion 4
G1.6 (excluding Fagus woodland (excluding MedioG1.66)
European limestone Fagus forests)
G1.66 (*9150)
Medio-European limestone Fagus
forests
G1.7 (incl.
*91H0, *91I0)
Thermophilous deciduous
woodland
G1.8
Acidophilous Quercus-dominated
woodland
Quercus – Fraxinus – Carpinus
betulus woodland on eutrophic and
mesotrophic soils
Ravine and slope woodland
Krymski Hory; Tarkhankut;
Karadah; Mehanom; Tepe-Oba
Zaplava Desny; Zaplavy Dnipra
i Sozha; Kanivski Ostrovy;
Kremenchutski Plavni
Mehanom
Zaplava Desny; Zaplavy Dnipra
i Sozha; Kremenchutski Plavni;
Zaplava Tysy; Kanivski Ostrovy
G1.12 (included Boreo-alpine riparian galleries
Chyvchyny; Marmaroski Hory;
in *91E0)
Svydovets; Chornohora
G1.21 (included Riverine Fraxinus – Alnus woodland, Zaplava Desny; Zaplavy Dnipra
in *91E0)
wet at high but not at low water
i Sozha; Koncha-Zaspivskyi Lis;
Dubrova
Zaplava Siverskoho Dintsia;
Mixed Quercus – Ulmus – Fraxinus Samarskyi Lis; Zaplava Latorystsi;
G1.22
woodland of great rivers
Zaplavni Lisy na Borzhavi; Zaplava
Desny; Dnipro – Oril
G1.3
Mediterranean riparian woodland Zaplava Tysy; Kremenchutski
Plavni; Dnipro – Oril; Samarskyi
Lis; Kanivski Ostrovy; Zaplava
Siverskoho Dintsia
G1.4115
Eastern Carpathian Alnus gluinosa –
swamp woods
G1.414
Steppe swamp Alnus gluinosa
Zaplava Siverskoho Dintsia;
woods
Samarskyi Lis; Zaplavni Lisy na
Borzhavi; Sviai Hory; Volyzhyn Lis
G1.51
Sphagnum Betula woods
Somyno; Morochno-2; Cheremske
Boloto
22
Code
G1.A1
G1.A4 (incl.
*9180)
G3.1B
G3.1F
G3.25
G3.4232
G3.4E
G3.5 (incl.
*9530)
G3.7
G3.9 (incl.
*9560)
G3.E (*9120)
Alpine and Carpathian subalpine
Picea forests
Enclave Picea abies forests
Carpathian Larix and Pinus cembra
forests
Sarmaic steppe Pinus sylvestris
forests
Ponto-Caucasian Pinus sylvestris
forests
Pinus pallasiana and Pinus banaica
forests
Lowland to montain mediterranean
Pinus woodland (excluding Pinus
nigra)
Coniferous woodland dominated by
Cupressaceae or Taxaceae
Nemoral bog conifer woodland
23
IPAs where the habitat is a
criterion of selecion
Krymski Hory; Uholka – Shyrokyi
Luh; Skhidne Roztochia; Stuzhytsia;
Svydovets
Verkhniobuzky Lisy;
Tsetsyno; Holohirskyi Lis;
Uholka – Shyrokyi Luh
Krymski Hory; Savranskyi Lis (91I0);
Serbyno (91I0); Yulivski Hory
(91H0, 91I0, 91M0); Chorna Hora
(91H0, 91M0); Kreidiana Flora
(91I0); Potashnianski Lisy (91I0);
Dubrova (91I0); Ternava-Dnister
(91I0); Levkivski Lisy (91I0); Ardov
(91H0)
Slovechanskyi Kriazh; Polonskyi Lis;
Horodnytskyi Lis; Yemilchynski Lisy
Medobory; Chornyi Lis;
Homilshanskyi Lis; Brytavskyi Lis;
Sviai Hory
Medobory; Verkhniobuzki Lisy;
Krymski Hory; Marmaroski Hory;
Stuzhytsia
Chyvchyny; Chornohora; Gorgany;
Svydovets; Grofa
Cherevaskyi Lis; Rys; Cheremske
Boloto; Shatski Ozera
Gorgany; Yaiko Perehinske; Kedryn;
Ihrovets - Tavpishyrka
Sviai Hory; Samarskyi Lis; KonchaZaspivskyi Lis; Bir na Merli
Krymski Hory
Krymski Hory; Mys Marian
Krymski Hory
Krymski Hory; Mys Marian;
Karahah
Syra Pohonia; Ubort – Bolotnytsia;
Turova Dacha; Morochno-2;
Vtenske Boloto
Code
H1
Name in Resoluion 4
H2.6 (incl.
*8160)
Terrestrial underground caves, cave
systems, passages and waterbodies
Temperate-montane acid siliceous
screes
Temperate-montane calcareous and
ultra-basic screes
Acid siliceous screes of warm
exposures
Calcareous and ultra-basic screes of
warm exposures
H3.1
Acid siliceous inland clifs
H3.2
Basic and ultra-basic inland clifs
H3.511 (*8240)
X01
X02
X03
X04 (incl. *7110)
Limestone pavements
Estuaries
Saline coastal lagoons
Brackish coastal lagoons
Raised bog complexes
X18
X29
X35
Wooded steppe
Salt lake islands
Inland Sand Dunes
H2.3
H2.4 (incl.
*8160)
H2.5
IPAs where the habitat is a
criterion of selecion
–
Gorgany; Svydovets; Chornohora;
Marmaroski Hory
Chyvchyny
Арап Р.Я. Методичні аспекти впровадження міжнародної програми "Важливі ботанічні
території" в Україні. – К.: Арістей, 2008. – 43 с.]
Palmer M., Smart J. Guidelines to the selecion of Important Plant Areas in Europe.
Planta Europa, 2001.
Red Data Book of European bryophytes, Trondheim, 1995. — 291 pp.
Red Data Book of Ukraine. Vegetable kingdom / ed. by Ya.P. Didukh. – Kyiv:
Globalconsuling, 2009. – 900 pp. (in Ukr.) [Червона книга України. Рослинний світ / за
ред. Я.П. Дідуха. – К.: Глобалконсалтинг, 2009. – 900 с.]
–
Krymski Hory; Ternava – Dnister;
Zhyzhava
Hranitno-Stepove Pobuzhia;
Dolyna Mertvovodu; Kamiani
Mohyly
Krymski Hory; Karadah; Ayu-Dah;
Tepe-Oba; Ternava – Dnister
Krymski Hory; Kazantyp; Chyvchyny
–
–
–
Syra Pohonia; Ubort – Bolotnytsia;
Morochno-2
–
–
Kozachelaherska Arena;
Triokhizbenskyi Step; Zaplava
Prypiai
Literature
Anderson S. Idenifying Important Plant Areas: a site selecion manual for Europe, and
a basis for developing guidelines for other regions of the world. – Plantlife, 2002. – 52 pp.
Idenifying and protecing the World's most important plant areas. – Plantlife, 2004. –
7 pp.
Kolomiychuk V.P., Onyshchenko V.A., Peregrym M.M. Important plant areas of Azov
Region / Ed. T.L. Andrienko. – Kyiv: Alterpress, 2012. – 42 pp. (in Ukr.) [Коломійчук В.П.,
Онищенко В.А., Перегрим М.М. Важливі ботанічні території Приазов'я / під ред.
Т.Л. Андрієнко. – Київ: Альтерпрес, 2012. – 42 с.]
Mosyakin S.L., Fedoronchuk M.M. Vascular plants of Ukraine: A nomenclatural checklist.
Kiev, 1999. – xxiv + 346 pp.
Onyshchenko V.A., Andrienko T.L., Chorney I.I., Kolomiychuk V.P., Didukh Ya.P., Oliiar
H.I., Pryadko O.I. Methodical aspects of introducion of internaional program "Important
Plant Areas" in Ukraine. – Kyiv: Aristei, 2008 – 43 pp. (in Ukr.) [Онищенко В.А., Андрієнко
Т.Л., Чорней І.І., Коломійчук В.П., Дідух Я.П., Оліяр Г.І., Прядко О.І., Вірченко В.М.,
24
25
Descripions of IPAs
Totally, 173 IPAs were ideniied (maps 1, 2). Their total area is 1 300 774 ha
(2.29% of Ukraine with the Territorial sea). The largest areas are “Krymski Hory” (“Crimean
Mountains”, 301 150 ha) and “Zaplava Desny” (“Floodplain of the Desna”, 185 741 ha). The
smallest area is 4.4 ha. About 55.8% of the total area of IPAs belong to naional and regional
(subnaional) protected areas. Overlapping with Emerald Sites is 86.6%.
Map 2. Important Plant Areas in Ukraine
Aiu-Dah
Map 1. Centres of Important Plant Areas in Ukraine
26
V.A. Onyshchenko
Ukrainian name: Аю-Даг.
Transliteraion/Translaion variants: Ayu-Dag, Ayuv Dağ (Crimean Tatar), Vedmid’-Hora
(Ukrainian), Medved’-Gora (Russian).
Area: 636.0 ha.
Altitude: 0–571 m.
Latitude: 44°33’29” N (44.5581°).
Longitude: 34°20’09” E (34.3357°).
Administrative regions. Autonomous Republic of Crimea: Alushta city (mainly), Yalta city;
Territorial waters of Ukraine.
Ownership: state.
Biogeographic regions: steppic.
Habitats. Level 1. A – 15.3%; B – 1.7%; E – 10.0%; F – 40.0%; G – 25.0%; H – 8.0%.
Habitats. Level 2. A1 Littoral rock and other hard substrata – 0.5%; A2 Littoral sediment –
1.1%; A3 Infralittoral rock and other hard substrata –7.0%; A5 Sublittoral sediment – 6.7%;
B2 Coastal shingle – 1.2%; B3 Rock cliffs, ledges and shores, including the supralittoral
– 0.5%; E1 Dry grasslands – 10.0%; F3 Temperate and mediterranean-montane scrub –
27
15.0%; F5 Maquis, arborescent matorral and thermo-Mediterranean bushes – 25.0%; F6
Garrigue; G1 Broadleaved deciduous woodland – 25.0%; H2 Screes – 1.9%; H3 Inland cliffs,
rock pavements and outcrops – 6.0%; H5 Miscellaneous inland habitats with very sparse
or no vegetation – 0.1%.
basic inland clifs; H5.6 Trampled areas.
Land use: nature conservaion and research – 100%; tourism/recreaion – major
Protected areas: includes Aiu-Dah state landscape reserve (527 ha), overlaps (about 100
ha) with regional hydrological nature monument “Pryberezhnyi Kompleks Bilia Hory Aiudah”.
Threats: development (recreaion/tourism) – low, eutrophicaion – low.
General description. The IPA is located in the southern part of the Crimea. It includes Mount
Ayu-Dah and a 200-m wide strip of the sea. The main rock type is gabbro. A large area is
occupied by low (4–5 m) Quercus pubescens and Carpinus orientalis woods with some presence of Fraxinus angusifoliua, Juniperus excelsa, Sorbus domesica. Low shrubby plants of
Ruscus ponicus oten form a dense layer. Major species of the herb layer are Poa sterilis
and Luzula forsteri. On the top of the mount, main dominant of the tree layer is Quercus
petraea. In the herb layer, Poa sterilis and Luzula forsteri are dominants. The Southern slope
is covered by low thickets of Juniperus excelsa, Juniperus oxycedrus, Cistus tauricus, Quercus
pubescens. Considerable areas are occupied by screes and outcrops. Litoral habitats are
represented by shingle beach and rocks.
Botanical significance. Main locality of endemic species Brassica taurica (>80% of global
populaion). Important area for conservaion of basic outcrops and termophilous deciduous
woods.
Criterion A(ii)
• Brassica taurica (Tzvelev) Tzvelev (Brassica sylvestris Mill. subsp. taurica Tzvelev); A(i);
A(ii); abundance: 190 individuals; trend: unknown; species data quality: good; trend
data quality: poor.
Criterion C
• H3.2 Basic and ultra-basic inland cliffs; area: 40 ha; trend: stable; area data quality: poor;
trend data quality: medium.
Literature
1. Червона книга України. Рослинний світ / за ред. Я.П. Дідуха. – К.: Глобалконсалтинг, 2009. – 900 с.
2. Шеляг-Сосонко Ю.Р., Дидух Я.П., Молчанов Е.Ф. Государственный заповедник
“Мыс Мартьян”. – Киев: Наук. думка, 1985. – 260 с.
Ak-Monaiskyi Step
Futher habitat description. A2.1 Litoral coarse sediment; B2.1 Shingle beach dritlines;
B2.2 Unvegetated mobile shingle beaches above the dritline; B2.3 Upper shingle beaches
with open vegetaion; B3.1 Supralitoral rock (lichen or splash zone); B3.2 Unvegetated
rock clifs, ledges, shores and islets; B3.3 Rock clifs, ledges and shores, with angiosperms;
B3.4 Sot sea-clifs, oten vegetated; E1.1 Inland sand and rock with open vegetation; E1.3
Mediterranean xeric grassland; F3.2 Submediterranean deciduous thickets and brushes;
F5.2 Maquis; F5.3 Pseudomaquis; F6.4 Black Sea garrigues; G1.7 Thermophilous deciduous
woodland; H2.6 Calcareous and ultra-basic screes of warm exposures; H3.2 Basic and ultra-
28
V.P. Kolomiychuk, V.A. Onyshchenko
Ukrainian name: Ак-Монайський степ.
Transliteraion/Translaion variants: Ak-Monaiskii Steppe.
Area: 468.0 ha.
Altitude: 0–37 m.
Latitude: 45°18’53” N (45.3146°).
Longitude: 35°36’38” E (35.6104°).
Administrative regions. Autonomous Republic of Crimea: Lenine raion; Territorial waters
of Ukraine.
Ownership: state.
Biogeographic regions: steppic.
Habitats. Level 1. A – 16.0%; B – 3.0%; E – 78.0%; H – 3.0%.
29
Habitats. Level 2. A1 Littoral rock and other hard substrata – 0.2%; A2 Littoral sediment
– 1.3%; A3 Infralittoral rock and other hard substrata – 2.5%; A5 Sublitoral sediment –
12.0%; B1 Coastal dunes and sandy shores – 0.1%; B2 Coastal shingle – 2.4%; B3 Rock clifs,
ledges and shores, including the supralittoral – 0.5%; E1 Dry grasslands – 78.0%; H2 Screes
– 0.3%; H3 Inland clifs, rock pavements and outcrops – 1.7%; H5 Miscellaneous inland
habitats with very sparse or no vegetation – 1.0%.
Criterion A
• Astragalus reduncus Pall.; A(iv); abundance: occasional; trend: unknown; species data
quality: medium; trend data quality: poor.
• Crambe grandilora DC.; A(iv); abundance: occasional; trend: unknown; species data
quality: medium; trend data quality: poor.
• Crambe mitridais Juz.; A(iv); abundance: occasional; trend: unknown; species data
quality: medium; trend data quality: poor.
Conservaion proposals. Create a state botanical reserve.
Arabatska Strilka
Futher habitat description. A2.1 Litoral coarse sediment; A3.2 Atlanic and Mediterranean
low energy infralitoral rock; A5.1 Sublitoral coarse sediment; A5.2 Sublitoral sand; A5.5
Sublitoral macrophyte-dominated sediment; B1.2 Sand beaches above the dritline; B2.2
Unvegetated mobile shingle beaches above the dritline; B2.3 Upper shingle beaches with
open vegetaion; B3.1 Supralitoral rock (lichen or splash zone); B3.2 Unvegetated rock
clifs, ledges, shores and islets; B3.3 Rock clifs, ledges and shores, with angiosperms;
B3.4 Sot sea-clifs, oten vegetated; E1.1 Inland sand and rock with open vegetation; E1.2
Perennial calcareous grassland and basic steppes; H2.6 Calcareous and ultra-basic screes
of warm exposures; H3.2 Basic and ultra-basic inland clifs; H5.3 Sparsely- or un-vegetated
habitats on mineral substrates not resulting from recent ice activity; H5.6 Trampled areas.
Land use: agriculture (animals) – 30%; tourism/recreaion – minor.
Protected areas: overlaps (446 ha) with Emerald Site “Ak-Monaiskyi Steppe”.
Threats: –.
General description. A part of the southern shore of the Sea of Azov. Major vegetaion
type is the steppe dominated by Festuca valesiaca, Koeleria cristata, Sipa brauneri, S.
capillata, S. pulcherrima. Other fequent species are Artemisia taurica, Bellevalia sarmatica,
Convolvulus cantabrica, Dactylis glomerata, Euphorbia seguierana, Gypsophilla paniculata,
Ornithogalum flavescens, Poa bulbosa, Securigera varia, Teucrium chamaedrys, Thalictrum
minus, Tragopogon major, Verbascum phlomoides, Xeranthemum annum. By the sea,
Crambe pontica, Eringium maritimum, Leymus sabulosus occasionally prevail. There are
many calcareous clifs and screes.
Botanical significance. Important area for some steppe species, in paricular Astragalus
reduncus, Crambe grandilora, Crambe mitridais.
V.P. Kolomiychuk, V.A. Onyshchenko
Ukrainian name: Арабатська стрілка.
Transliteraion/Translaion variants: Arabatska Spit, Arabatska Bar.
Area: 12377.0 ha.
Altitude: 0–5 m.
Latitude: 45°31’26” N (45.5238°).
Longitude: 35°11’00” E (35.1831°).
Administraive regions. Autonomous Republic of Crimea: Lenine raion. Territorial waters
of Ukraine.
Ownership: state.
Biogeographic regions: steppic.
Habitats. Level 1. A – 76.5%; B – 2.0%; C – 2.3%; D – 1.0%; E – 17.2%; H – 1.0%.
Habitats. Level 2. A2 Littoral sediment – 10.9%; A5 Sublittoral sediment – 65.6%; B1 Coastal
dunes and sandy shores – 1.4%; B2 Coastal shingle – 0.6%; C1 Surface standing waters
– 2.3%; D6 Inland saline and brackish marshes and reedbeds – 1.0%; E1 Dry grasslands –
13.2%; E6 Salt steppes – 4.0%; H5 Miscellaneous inland habitats with very sparse or no
vegetation – 1.0%.
Futher habitat description. A2.1 Littoral coarse sediment; A2.2 Littoral sand and muddy
sand; A2.3 Littoral mud; A2.4 Littoral mixed sediments; A2.5 Coastal saltmarshes and
saline reedbeds; B1.1 Sand beach driftlines; B1.2 Sand beaches above the driftline; B1.3
Shifting coastal dunes; B1.4 Coastal stable dune grassland (grey dunes); B2.1 Shingle beach
driftlines; B2.2 Unvegetated mobile shingle beaches above the driftline; C1.5 Permanent
inland saline and brackish lakes, ponds and pools; C1.6 Temporary lakes, ponds and pools;
D6.1 Inland saltmarshes; D6.2 Inland saline or brackish species-poor helophyte beds
normally without free-standing water; E1.2 Perennial calcareous grassland and basic
steppes (E1.2G); E6.2 Coninental inland salt steppes; H5.6 Trampled areas.
Land use: agriculture (animals) – 20%; forestry – minor; hunting – minor; mowing/hay
making – minor; nature conservation and research – major; tourism/recreaion – minor.
Protected areas: includes Arabatskyi state landscape reserve (600 ha), included in Ramsar
Site “Eastern Syvash”, included in Emeral Site “Eastern Syvash”.
Threats: climate change/ sea level rise – medium, development (recreaion/tourism) – low,
natural events: erosion – medium.
General description. A bar consising of sand and shell with the adjacent waters of the
Sea of Azov and Syvash Bay. The sand steppe occupies elevated areas. It is dominated by
Agropyron cimmericum, Ephedra distachya, Euphorbia seguierana, Marrubium peregrinum,
30
31
Medicago kotovii, Stipa borysthenica, Sipa capillata, Tortula ruralis. Dominants of halophyic vegetaion are Boloboschoenus mariimus, Elytrigia elongata, Halimione pedunculata, Halimione verrucifera, Halocnemum strobilaceum, Juncus mariimus, Limonium meyeri,
Plantago salsa, Puccinellia gigantea, Salicornia perennans, Suaeda prostrata, Tripolium
pannonicum, Phragmites australis. On the supralitoral sand and shell, Leymus sabulosus,
Crambe mariima, Сakile euxina, Centaurea odessana, Carex colchica, Eryngium maritimum
prevail. Large areas of the Syvash are ocuupied by Zostera marina.
Botanical significance. Important area for conservaion of sand steppes and brackish
aquaic vegetaion.
•
Goniolimon rubellum (S.G.Gmel.) Klokov; A(iii); abundance: occasional; trend: stable;
species data quality: medium; trend data quality: poor.
• Lepidium syvaschicum Kleopow; A(iii); abundance: occasional; trend: stable; species
data quality: medium; trend data quality: poor.
• Thymus litoralis Klokov et Shost.; A(iii); abundance: occasional; trend: stable; species
data quality: medium; trend data quality: poor.
• Zostera marina L.; A(ii); abundance: abundant; trend: stable; species data quality: good;
trend data quality: poor.
Criterion C
• B1.1 Sand beach driftlines; area: 10 ha; trend: stable; area data quality: medium; trend
data quality: medium.
• B1.3 Shifting coastal dunes; area: 50 ha; trend: stable; area data quality: medium; trend
data quality: medium.
• B1.4 Coastal stable dune grassland (grey dunes); area: 90 ha; trend: stable; area data
quality: medium; trend data quality: medium.
Conservaion proposals. Increase area of Arabatskyi state landscape reserve to include all
of the IPA.
Literature
1. Коломійчук В.П., Абкадирова Е.Ш. Ботанічна і ландшафтна характеристика проектованого ландшафтного заказника “Арабатський степ” // Проблеми фундаментальної і прикладної екології, екол. геології та раціон. природокористування : мат-ли міжнарод. наук.-практ. конфер. – Кривий Ріг: Видавничий Дім, 2009. – С. 290–293.
2. Корженевский В.В., Клюкин А.А. Растительность абразионных и аккумулятивных
форм рельефа морских побережий и озер Крыма. – Ялта: ГНБС, 1990. – 108 c. – Деп. в
ВИНИТИ 10.07.90, № 3822-В90.
3. Лоскот Н.П. О растительном покрове южной части Арабатской стрелки / Вопросы
физиологии, биохимии, цитологии и флоры Украины. – К.: Наукова думка, 1974. –
С. 13–16.
Ardov
Criterion A
• Agropyron cimmericum Nevski; A(i); abundance: occasional; trend: stable; species data
quality: medium; trend data quality: poor.
32
R.Ya. Kish
Ukrainian name: Ардов.
Area: 31.0 ha.
Altitude: 113–240 m.
Latitude: 48°13’59” N (48.2330°).
Longitude: 22°39’05” E (22.6514°).
Administrative regions. Zakarpatska region: Berehove raion.
Ownership: state.
Biogeographic regions: pannonian.
Habitats. Level 1. E – 7.0%; F – 3.4%; G – 88.6%; H – 1.0%.
Habitats. Level 2. E1.2 Perennial calcareous grassland and basic steppes – 7.0%; F3
Temperate and mediterranean-montane scrub – 3.4%; G1 Broadleaved deciduous
woodland – 88.6%; H3 Inland cliffs, rock pavements and outcrops – 0.0%; H5 Miscellaneous
inland habitats with very sparse or no vegetation – 1.0%.
Futher habitat description. F3.1 Temperate thickets and scrub; F3.2 Submediterranean
33
deciduous thickets and brushes; G1.7 Thermophilous deciduous woodland; G1.A Mesoand eutrophic Quercus, Carpinus, Fraxinus, Acer, Tilia, Ulmus and related woodland; H5.6
Trampled areas.
Land use: forestry – 90%, nature conservation and research – 81%.
Protected areas: includes regional botanical reserve “Ardov” (24.0 ha).
Threats: abandonment/reducion of land management – medium.
General description. A hill composed of rhyoliic tuf, andesite, basalt, and sedimentary
rocks. The main vegetaion type is termophilous oak woods. Dominant of the tree layer
is Quercus dalechampii. The shrub layer consists of Cornus mas, Corylus avellana, Euonymus europaeus, Ligustrum vulgare, Pyrus communis, Spiraea media, Swida sanguinea (Cornus sanguinea). In the herb layer, there are Achillea millefolium, Anthericum ramosum,
Anthoxanthum odoratum, Brachypodium pinnatum, Campanula patula, Campanula persicifolia, Carex caryophyllea, Carex michelii, Chondrilla juncea, Clinopodium vulgare, Euphorbia cyparissias, Hylotelephium polonicum, Inula ensifolia, Laser trilobum, Melica uniflora,
Melittis melissophyllum, Millium efusum, Muscari comosum, Poa nemoralis, Polygonatum
odoratum, Potentilla argentea, Potentilla recta, Veronica teucrium, Vincetoxicum
hirundinaria, Viscaria viscosa. There are also dry grasslands (with Dianthus carthusianorum, Festuca rupicola, Galathella linosyris, Galium campanulatum, Geranium sanguineum,
Melica transsilvanica, Phleum phleoides, Rosa gallica, Seseli annuum, Veronica spicata) and
shrubs (Crataegus monogyna s. l., Prunus spinosa, Rosa sp., Spiraea media).
Botanical significance. The largest locaion of Habitat Direcive priority habitat 91H0 in
Ukraine.
Criterion C
• G1.7 Thermophilous deciduous woodland (*91H0); area: 24 ha; trend: stable; area data
quality: good; trend data quality: medium.
34
Askania-Nova
V.A. Onyshchenko
Ukrainian name: Асканія-Нова.
Area: 11024.0 ha.
Altitude: 19–32 m.
Latitude: 46°27’47” N (46.4630°).
Longitude: 33°57’46” E (33.9628°).
Administrative regions. Kherson region: Chaplynka raion.
Ownership: state.
Biogeographic regions: steppic.
Habitats. Level 1. C – 0.2%; E – 99.8%.
Habitats. Level 2. C1 Surface standing waters – 0.1%; C3 Littoral zone of inland surface
waterbodies – 0.1%; E1 Dry grasslands – 88.0%; E2 Mesic grasslands – 11.0%; E3 Seasonally
wet and wet grasslands – 0.8%.
Futher habitat description. E1.2 Perennial calcareous grassland and basic steppes, E2.5
Meadows of the steppe zone, E3.4 Moist or wet eutrophic and mesotrophic grassland.
Land use: nature conservation and research – 100%; agriculture (animals) – 20%; tourism/
recreation – minor.
Protected areas: included in Askania-Nova biosphere reserve (naional category), AskaniaNova biosphere reserve (UNESCO), and Emerald Site “Askaniia-Nova Biosphere Reserve”.
Threats: abandonment / reduction of land management – low, burning of vegetaion – low.
General description. One of the largest steppe areas in Ukraine. It represents the southern
European steppes on chestnut soils. Main dominants of steppe communities are Stipa
ucrainica, Sitpa capillata, Stipa lessingiana, Festuca valesiaca. Species of high constancy:
Dianthus andrzejowskianus, Euphorbia seguierana, Falcaria vulgaris, Galium ruthenicum,
Goniolimon tataricum, Hylotelephium stepposum, Limonium sareptanum, Phlomoides hybrida (Phlomis hybrida), Phlomis pungens, Potentilla laciniosa, Seseli tortuosum, Verbascum
phoeniceum. In depressions, there are mesic grasslands dominated by Poa angustifolia,
35
Alopecurus pratensis, Carex praecox with presence of Achillea micranthoides, Allium
regelianum, Euphorbia virgata, Phalacrachena inuloides.
In more humid habitats, Elytrigia pseudocaesia and Catex melanostachya dominate. In years
with floods they are replaced with Beckmannia eruciformis, Butomus umbellatus, Gratiola
officinalis, Lythrum virgatum, Schoenoplectus lacustris.
Botanical significance. This area is one of the most important for conservation of steppe
vegetation in Europe.
Criterion A
• Allium regelianum A.Becker ex Iljin; A(ii); abundance: occasional; trend: fluctuating;
species data quality: medium; trend data quality: medium. A large part of national
population.
• Astragalus reduncus Pall.; A(iv); abundance: rare; trend: unknown; species data quality:
medium; trend data quality: poor.
• Centaurea taliewii Kleopow (Rhaponicoides taliewii (Kleopow) M.V.Agab. & Greuter);
A(iv); abundance: occasional; trend: unknown; species data quality: medium; trend data
quality: poor.
• Ferula orientalis L.; A(ii); abundance: occasional; trend: unknown; species data quality:
medium; trend data quality: poor.
• Tulipa scythica Klokov et Zoz; A(iii); abundance: frequent; trend: fluctuating; species
data quality: medium; trend data quality: medium. A large part of national population.
Criterion C
• E1.2 Perennial calcareous grassland and basic steppes; area: 9617 ha; trend: stable;
area data quality: good; trend data quality: good.
Literature
1. Веденьков Е.П. Флора заповедника “Аскания-Нова” (аннотированный список
цветковых растений заповедной степи). – М., 1989. – 52 с.
2. Дрогобыч Н.Е., Шаповал В.В. Распространение редких, исчезающих и эндемичных
видов флоры цветковых в заповедной степи “Аскания-Нова”. 2. Эндемичное ядро //
Вісті Біосферного заповідника “Асканія-Нова”. – Асканія-Нова, 2004. – Т. 6. – С. 6–13.
3. Ткаченко В.С. Особливості автогенезу асканійського степу // Укр. ботан. журн. –
1990. – 47, № 4. – С. 20–25.
4. Ткаченко В.С., Шаповал В.В. Сукцесії фітосистем ділянки “Північна”
новоасканійського заповідного степу у другій половині ХХ і на початку ХХІ ст. // Вісті
Біосферного заповідника “Асканія-Нова”. – 2010. – Т. 12. – С. 21–32.
5. Шаповал В.В. БЗ Асканія-Нова імені Ф.Е. Фальц-Фейна // Фіторізноманіття
заповідників і національних природних парків України. Ч. 1. Біосферні заповідники.
Природні заповідники / під ред. В.А. Онищенка і Т.Л. Андрієнко. – Київ: Фітосоціоцентр,
2012. – С. 13–30.
Balakyrivka
Ukrainian name: Балакирівка.
Area: 266.0 ha.
Altitude: 57–140 m.
Latitude: 49°20'57" N (49.3490°).
Longitude: 38°55'08" E (38.9188°).
Administrative regions. Luhansk region: Starobilsk raion.
Ownership: state.
Biogeographic regions: steppic.
V.A. Onyshchenko
Habitats. Level 1. E – 78.5%; F – 2.0%; G – 13.9%; H – 5.6%.
Habitats. Level 2. E1 Dry grasslands – 78.5%; F3 Temperate and mediterranean-montane
scrub – 2.0%; G1 Broadleaved deciduous woodland – 13.9%; H2 Screes – 5.6%.
Futher habitat description. E1.1 Inland sand and rock with open vegetation; E1.2 Perennial
36
37
calcareous grassland and basic steppes; F3.2 Submediterranean deciduous thickets and
brushes; G1.7 Thermophilous deciduous woodland; G1.A Meso- and eutrophic Quercus,
Carpinus, Fraxinus, Acer, Tilia, Ulmus and related woodland; G1.C Highly artificial
broadleaved deciduous forestry plantations; H2.6 Calcareous and ultra-basic screes of
warm exposures.
Land use: agriculture (animals) – major; forestry – minor; nature conservaion and research
– major.
Protected areas: includes Balakyrivskyi regional landscape reserve (204 ha), included in
Emerald Site "Balakyrivskyi".
Threats: agricultural intensification/expansion (arable) – low, extracion (minerals/quarries)
– low; forestry (aforestaion) – medium.
General description. Slopes of the valleys of the Aidar Rivers with the steppe vegetaion,
chalk outcrops, and oak woods. Steppes are dominated by Sipa capillata, Sipa lessingiana,
Festuca valesiaca s.l. Dominants of chalk outcrops are Hyssopus cretaceus, Pimpinella
titanophila, Thymus cretaceus.
Botanical significance. Important area for conservaion of chalk outcrops with endemic
species.
Criterion A
• Erysimum ucrainicum J.Gay.; A(iv); abundance: unknown; trend: unknown; species data
quality: poor; trend data quality: poor.
• Linaria cretacea Fisch. ex Spreng.; A(iv); abundance: unknown; trend: unknown; species
data quality: poor; trend data quality: poor.
Conservaion proposals. Do not allow aforestaion. Change the status of regional
landscape reserve to state botanical reserve.
Land use: agriculture (animals) – major; nature conservaion and research – major
Protected areas: included in Pryazovskyi naional nature park, overlaps (about 670 ha) with
Syvashyk state landscape reserve, included in Emerald Site "Pryazovskyi Naional Nature
Park".
Bereh Syvashyka
V.P. Kolomiychuk, V.A. Onyshchenko
Ukrainian name: Берег Сивашика.
Transliteraion/Translaion variants: Bereg Sivashika, Coast of Syvashyk.
Area: 1646.0 ha.
Altitude: 0–8 m.
Latitude: 46°22'43" N (46.3786°).
Longitude: 35°05'30" E (35.0916°).
Administrative regions. Zaporizhia region: Yakymivka raion .
Ownership: state (major), private.
Biogeographic regions: steppic.
Habitats. Level 1. A – 30.7%; C – 23.9%; E – 45.3%; H – 0.1%.
Habitats. Level 2. A2 Littoral sediment – 3.9%; A5 Sublittoral sediment – 26.8%; C1 Surface
standing waters – 18.0%; C3 Littoral zone of inland surface waterbodies – 5.9%; E1 Dry
grasslands – 3.0%; E6 Salt steppes – 42.3%; H5 Miscellaneous inland habitats with very
sparse or no vegetation – 0.1%.
Futher habitat description. A2.3 Littoral mud; A2.5 Coastal saltmarshes and saline
reedbeds; A5.3 Subittoral mud; C3.2 Water-fringing reedbeds and tall helophytes other
than canes; E1.2 Perennial calcareous grassland and basic steppes; E6.2 Continental inland
salt steppes; H5.6 Trampled areas.
38
Threats: burning of vegetaion – low.
General description. Syvashyk is a lagoon of the Sea of Azov. On the litoral of the lagoon
and saline lakes, there are salt marshes with Halocnemum strobilaceum, Artemisia santonica, Petrosimonia brachiata, P. oposiifolia, Suaeda salsa, Puccinellia fominii, P. distans. At
slightly higher levels, there are salt grasslands dominated by Elytrigia elongata, Limonium
meyeri, Juncus gerardii, Apera mariima. On rather steep slopes and on the plateau, there
are typical steppes and salt steppes. The typical steppes are dominated by Festuca valesiaca, Koeleria cristata, Sipa capillata, S. ucrainica. Major dominants of the salt steppes
are Agropyron pecinatum, Artemisia taurica, Festuca valesiaca s.l., Kochia prostrata.
Botanical significance. One of 10 known localiies of Allium regelianum in Ukraine. One of 7
known localiies of Allium pervesitum in Ukraine.
39
Criterion A
• Allium pervesitum Klokov; A(i), A(iv); abundance: rare; trend: unknown; species data
quality: medium; trend data quality: poor.
• Allium regelianum A.Becker ex Iljin.; A(ii); abundance: rare; trend: unknown; species
data quality: medium; trend data quality: poor.
• Goniolimon rubellum (S.G.Gmel.) Klokov; A(iii); abundance: rare; trend: stable; species
data quality: medium; trend data quality: poor.
Literature
1. Костильов О.В., Устименко П.М., Попович С.Ю. Рослинний покрив заказника
"Сивашик" // Укр. ботан. журн. – 1994. – 51, № 5. – С. 115–120.
2. Коломійчук В.П. Ключові ботанічні території Північного Приазов'я // Заповідна
справа в Україні. – 2008. – Т. 14, вип. 1. – С. 61–66.
3. Коломійчук В.П., Яровий С.О. Конспект флори судинних рослин Приазовського
національного природного парку. – К.: Альтерпрес, 2011. – 296 с.
Protected areas: overlaps (8051 ha) with Rivnenskyi nature reserve, overlaps (8051) with
Emerald Site "Rivnenskyi Nature Reserve".
Bile Ozero
V.A. Onyshchenko
Ukrainian name: Біле Озеро.
Transliteraion/Translaion variants: White Lake
Area: 11126 ha.
Altitude: 154–165 m.
Latitude: 51°30'03" N (51.5008°).
Longitude: 25°42'42" E (25.7116°).
Administrative regions. Rivne region: Volodymyrets, Zarichne raion; Volynska region:
Manevychi raion.
Ownership: state, private (minor).
Biogeographic regions: continental.
Habitats. Level 1. C – 4.2%; D – 38%; E – 0.1%; F – 1.0%; G – 56.7%.
Habitats. Level 2. C1 Surface standing waters – 4.2%; D2 – Valley mires, poor fens and
transition mires – 25%; D5 Sedge and reedbeds, normally without free-standing water
– 13%; E2 Mesic grasslands – 0.1%; F9 Riverine and fen scrubs – 1%; G1 Broadleaved
deciduous woodland – 10%; G3 Coniferous woodland – 40.0%; G4 Mixed deciduous and
coniferous woodland – 6.7%.
Futher habitat description. C1.1 Permanent oligotrophic lakes, ponds and pools; C1.2
Permanent mesotrophic lakes, ponds and pools; D2.3 Transition mires and quaking bogs;
D5.1 Reedbeds normally without free-standing water; D5.2 Beds of large sedges normally
without free-standing water; G1.5 Broadleaved swamp woodland on acid peat; G1.8
Acidophilous Quercus-dominated woodland; G1.9 Non-riverine woodland with Betula,
Populus tremula or Sorbus aucuparia; G1.A Meso- and eutrophic Quercus, Carpinus,
Fraxinus, Acer, Tilia, Ulmus and related woodland; G1.B Non-riverine Alnus woodland; G3.4
Pinus sylvestris woodland south of the taiga; G3.E Nemoral bog conifer woodland, G4.1
Mixed swamp woodland; G4.4 Mixed Pinus sylvestris – Betula woodland; G4.7 Mixed Pinus
sylvestris – acidophilous Quercus woodland.
Land use: forestry – minor, nature conservation and research – 72%; tourism/recreaion –
minor.
40
Threats: eutrophicaion – low; water (drainage) – low.
General description. Major habitats types are Pinus sylvestris forests and transiion mires.
The largest area is occupied by mesic pine forests dominated by Vaccinium myrillus
and Pleurozium shreberi. Constant species are Betula pendula, Dicranum polysetum,
Melampyrum pratense. On richer soils, diferenial species are Luzula pilosa, Maianthemum
bifolium, Quercus robur. Dry pine forests are dominanted by Cladonia gracilis, Cladonia
miis, Corynephorus canescens. There also occur swamped Alnus gluinosa woods (oten
with dominance of Sphagnum palustre or S. squarrosum), swamped Betula pubescens
41
woods, swamped Pinus sylvestris woods and bogs, wet Alnus gluinosa forests, small
areas of Picea abies forests and Carpinus betulus forests. Mires are represented by both
mesotrophic and eutrophic ones. The most typical are communiies dominated by Carex
lasiocarpa and Carex rostrata with a layer of Sphagnum species (S. cuspidatum, S. fallax,
S. lexuosum, S. subsecundum). In the places with a higher water level, Potenilla palustris,
Menyanthes trifoliata, Sphagnum teres dominate. Vegetaion of the oligo-mesotrophic karst
Lake Bile (453 ha) includes communiies of Chara delicatula, Isoetes lacustris, Myriophyllum
spicatum, Nymphaea candida, Phragmites australis, Polygonum amphibium, Potamogeton
gramineus, Potamogeton pusillus, Schoenoplectus lacustris (Scirpus lacustris).
Botanical significance. Important for conservaion of transiion mires and oligotrophic lakes.
Criterion C
• C1.1 Permanent oligotrophic lakes, ponds and pools; area: 400 ha; trend: decreasing;
area data quality: poor; trend data quality: poor.
• D2.3 Transition mires and quaking bogs; area: 1600 ha; trend: stable; area data quality:
medium; trend data quality: medium.
Conservaion proposals. Reduce the intensity of recreaion by Bile Lake. Create state
reserves in northeastern and southwestern parts of the IPA that are not yet included in
Rivnenskyi nature reserve.
Literature
1. Андрієнко Т.Л. ПЗ Рівненський // Фіторізноманіття заповідників і національних
природних парків України. Ч. 1. Біосферні заповідники. Природні заповідники / під
ред. В.А. Онищенка і Т.Л. Андрієнко. – Київ: Фітосоціоцентр, 2012. – С. 313–324.
2. Андрієнко Т.Л., Прядко О.І., Онищенко В.А. Раритетна компонента флори
Рівненського природного заповідника // Укр. ботан. журн. – 2006. – 63, № 2. – С. 220–
228.
3. Андриенко Т.Л., Шеляг-Сосонко Ю.Р. Растительный мир Украинского Полесья в
аспекте его охраны. – Киев: Наук. думка, 1983. – 215 с.
4. Водно-болотні угіддя України / під ред. Г.Б. Марушевського та І.С. Жарук. – К.:
Чорноморська програма Ветландс Інтернешнл, 2006. – 312 с.
5. Орлов О.О., Якушенко Д.М., Борисова О.В. Синтаксономія рослинності озера Біле
(Рівненcький природний заповідник) // Збереження та відтворення біорізноманіття
природо-заповідних територій: матеріали міжнар. наук.-практ. конф., присвяч.
10-річчю Рівненського природного заповідника (м. Сарни, 11 – 13 черв. 2009 р.) / ред.
кол.: М. Д. Будз та ін. – Рівне, 2009. – С. 258–266.
6. Фіторізноманіття Українського Полісся та його охорона / Під заг. ред.
Т.Л. Андрієнко. – Київ: Фітосоціоцентр, 2006. – 316 с.
Longitude: 35°14'00" E (35.2335°).
Administrative regions. Kharkiv region: Krasnokutsk raion.
Ownership: state.
Biogeographic regions: coninental.
Habitats. Level 1: C – 0.4%, %, D – 0.2%, F – 0.3%, G – 98.1%, H – 1.0%.
Habitats. Level 2: C1 Surface standing waters – 0.2%; C3 Littoral zone of inland surface
waterbodies – 0.2%; D2 Valley mires, poor fens and transition mires – 0.1%; D5 Sedge and
reedbeds, normally without free-standing water – 0.1%; F9 Riverine and fen scrubs – 0.3%;
G1 Broadleaved deciduous woodland – 8.1%, G3 Coniferous woodland – 75.0%; G4 Mixed
deciduous and coniferous woodland – 15.0%; H5 Miscellaneous inland habitats with very
sparse or no vegetation – 1.0%.
Futher habitat descripion: C1.2 Permanent mesotrophic lakes, ponds and pools; C3.2
Water-fringing reedbeds and tall helophytes other than canes; D2.3 Transiion mires and
quaking bogs; F9.2 Salix carr and fen scrub; G1.4 Broadleaved swamp woodland not on acid
peat; G1.7 Thermophilous deciduous woodland; G1.B Non-riverine Alnus woodland; G3.4
Pinus sylvestris woodland south of the taiga; G4.C Mixed Pinus sylvestris – thermophilous
Quercus woodland; H5.6 Trampled areas.
Bir na Merli
O.V. Bezrodnova, N.B. Saidakhmedova
Ukrainian name: Бір на Мерлі.
Transliteraion/Translaion variants: Pinewoods on the Merla River
Area: 2924.0 ha.
Altitude: 110–145 m.
Latitude: 50°03'44" N (50.0622°).
Land use: forestry – minor; nature conservation and research – 100%.
Protected areas: included in Slobozhanskyi naional nature park, included in Emerald Site
42
43
"Slobozhanskyi Naional Nature Park".
Threats: forestry (intensiied forest management) – low.
General description. Terrace of the Merla river with woods and swamps. Pinus sylvestris
woods (both natural and plantaions) occupies the largest area. Frequent species are
Calamagrosis epigeios, Chamaecyisus ruthenicus, Euonymus verrucosus, Frangula alnus,
Genista inctoria, Helichrysum arenarium, Polygonatum odoratum, Sambucus racemosa,
Solidago virgaurea, Sorbus aucuparia. In habitats with higher moisture and richness of soil,
the fracion of deciduous trees is higher. Typical species are Acer campestre, A. platanoides,
A. tataricum, Clinopodium vulgare, Dryopteris carthusiana, Dryopteris ilix-mas, Corylus
avellana, Euonymus europaeus, Fragaria vesca, Fraxinus excelsior, Geranium roberianum,
Padus avium, Poa nemoralis, Rubus saxailis, Quercus robur, Sambucus nigra, Tilia cordata,
Ulmus glabra. In moderately wet Alnus gluinosa woods, frequent species are Aegopodium
podagraria, Glechoma hederacea, Galium aparine, Padus avium, Sambucus nigra, Viburnum
opulus. The tree layer of swamped woods consists of Alnus gluinosa, Pinus sylvestris, Betula
pendula, Betula pubescens, Populus tremula. Dominants of the herb and shrub layers are
Carex vesicaria, Filipendula ulmaria, Molinia caerulea, Ribes nigrum, Salix aurita, Salix cinerea,
Scirpus sylvaica, Thelypteris palustris, Urica galeopsifolia. In mesotrophic habitats, there
occur Carex lasiocarpa, Eriophorum angusifolium, Eriophorum vaginatum, Lycopodium
annoinum, Spagnum auriculatum, S. fallax, S. lexuosum, S. girgensohnii, S. palustre,
S. subsecundum. Eutrophic and mesotrophic mires without the tree layer occupy smaller
areas. Frequent species of sandy glades are Artemisia marschalliana, Psephellus sumensis
(Centaurea sumensis), Helichrysum arenarium, Jurinea cyanoides, Koeleria sabuletorum,
Oenothera biennis, Rumex acetosella, Thymus pallasianus, Cladonia spp.
Botanical significance. One of the largest protected steppe pine woods in Ukraine. Swamps
with species that are rare in the forest-steppe zone. There are 6 species from the list of
criterion A species: Dracocephalum ruyschiana, Iris humilis, Iris hungarica, Jurinea cyanoides,
Liparis loeselii, Pulsailla patens.
Criterion A
• Liparis loeselii (L.) Rich.; A(ii); rare; trend: decreasing; species data quality: poor; trend
data quality: poor.
Criterion C
• G3.4232 Sarmaic steppe Pinus sylvestris forests; area: 1500 ha; trend: stable; area data
quality: medium; trend data quality: medium.
Literature
1. Безроднова О.В., Саідахмедова Н.Б. Збереження популяції Diphasiastrum
complanatum (L.) Holub у НПП "Слобожанський" // Природоохоронні території
в минулому, сучасному й майбутньому світі (до 130-річчя створення "Пам’ятки
Пеняцької" – першої природоохоронної території у Європі): Матеріали Другої
міжнародної наукової конференції (Львів – Броди – Пеняки, 26-27 жовтня 2016 року). –
Львів: Ліга-Прес, 2016. – С. 39–42.
2. Саідахмедова Н.Б., Безроднова О.В. Профілі та трансекти // Літопис природи
Національного природного парку "Слобожанський" (за 2015 рік). Т. 4. – Краснокутськ,
2016. – С. 13–20 [Рукопис].
3. Філатова О.В., Саідахмедова Н.Б., Клімов О.В. НПП Слобожанський //
Фіторізноманіття заповідників і національних природних парків України. Ч. 2.
Національні природні парки / під ред. В.А. Онищенка і Т.Л. Андрієнко. – Київ:
Фітосоціоцентр, 2012. – С. 486–495.
44
Bobrovyi Kut
V.P. Kolomiychuk
Ukrainian name: Бобровий Кут.
Area: 30.5 ha.
Altitude: 7–23 m.
Latitude: 47°05'16" N (47.0877°).
Longitude: 32°55'33" E (32.9258°).
Administrative regions. Kherson region: Velyka Oleksandrivka raion.
Ownership: state.
Biogeographic regions: steppic.
Habitats. Level 1. E – 91.5%; G – 3.5%; H – 2.0%; J – 3.0%.
Habitats. Level 2. E1 Dry grasslands – 91.5%; G1 Broadleaved deciduous woodland – 3.5%;
H5 Miscellaneous inland habitats with very sparse or no vegetaion – 2.0%; J3 Acive
opencast mineral extracion sites, including quarries – 3.0%.
Futher habitat description. E1.9 Open non-Mediterranean dry acid and neutral grassland,
including inland dune grassland; G1.C Highly artificial broadleaved deciduous forestry
plantations; H5.6 Trampled areas; J3.2 Active opencast mineral extraction sites, including
quarries.
Land use: agriculture (animals) – major, extracion (minerals) – minor, mowing/hay making
– minor.
Protected areas: –
Threats: agricultural intensiicaion/expansion (grazing) – low, burning of vegetaion – low,
45
extracion (mineral/quarries) – medium.
General description. Grassland on the sand terrace of the Inhulets River. Dominant
species are Artemisia campestris, Festuca beckeri, Poa bulbosa. Other frequent species are
Agropyron lavrenkoanum, Chondrilla juncea, Dianthus platyodon, Echinops ritro, Eragrostis
minor, Jurinea sp., Helicrysum arenarium, Koeleria glauca, Plantago scabra, Tortula ruralis,
Cladonia sp. Small parts of the area are ariicial wood stands with Robinia pseudoacacia,
Ulmus minor, Elaeagnus angusifolia and an illegal sand quarry.
Botanical significance. Important area for the narrow endemic species Centaurea paczoskii
(one of 2 localiies).
Criterion A
• Centaurea paczoskii Kotov ex Klokov; A(iii); abundance: occasional; trend: unknown;
species data quality: good; trend data quality: poor.
Conservaion proposals. Create a state botanical reserve and an Emerald Site.
Literature
1.Мойсієнко І.І., Мельник Р.М. Волошка Пачоського Centaurea paczoskii Klokov (C.
margaritacea Ten. subsp. paczoskii (Kotov et Klokov) Dostál) / Червона книга України.
рослинний світ. / за ред. Я.П. Дідуха. – К.: Глобалконсалтинг, 2009. – С. 306.
2.Крицька Л.І., Деркач О.М. Сучасний стан популяцій видів ряду Margaritaceae Klok.
(Centaurea L.) // Укр. ботан. журн. – 1991. – 48, № 3. – С. 78–80.
layer, there prevail Carex brevicollis, Carex pilosa, Corydalis solida, Lamium galeobdolon,
sometimes also Convallaria majalis, Hedera helix, Vinca minor. Typical species are Acer
campestre, Acer platanoides, Acer tataricum, Aegonychon purpureocaeruleum, Arum
besseranum, Cerasus avium, Cornus mas, Euphorbia amygdaloides, Iris graminea, Hedera
helix, Melica uniflora, Scutellaria altissima, Stellaria holostea, Swida sanguinea (Cornus
sanguinea), Viburnum lantana.
Brytavskyi Lis
V.A. Onyshchenko
Ukrainian name: Бритавський ліс.
Transliteraion/Translaion variants: Brytavskyi Forest.
Area: 4407 ha.
Altitude: 190–281 m.
Latitude: 48°10'36" N (48.1767°).
Longitude: 29°08'24" E (29.1400°).
Administrative regions. Vinnytsia region: Chechelnyk raion.
Ownership: state.
Biogeographic regions: coninental.
Habitats. Level 1. G – 99.5%; H – 0.5%.
Habitats. Level 2. G1 Broadleaved deciduous woodland – 99.5%; H5 Miscellaneous inland
habitats with very sparse or no vegetation – 0.5% .
Futher habitat description. G1.7 Thermophilous deciduous woodland; G1.A Meso- and
eutrophic Quercus, Carpinus, Fraxinus, Acer, Tilia, Ulmus and related woodland; H5.6
Trampled areas.
Land use: forestry – minor, nature conservation and research – major.
Protected areas: includes Brytavskyi state botanical reserve (3259 ha), included in
Karmeliukove Podillia naional nature park, included in Emerald Site "Karmeliukove Podillia
Naional Nature Park".
Threats: development (recreaion/tourism) – low.
General description. Almost the entire territory is occupied by the broadleaved forest. The
tree layer is dominated by Quercus petraea, Quercus robur, Carpinus betulus. In the herb
Botanical significance. Well preserved oak-hornbeam forest.
Criterion C
• G1.A1 Quercus – Fraxinus – Carpinus betulus woodland on eutrophic and mesotrophic
soils; area: 4350 ha; trend: stable; area data quality: good; trend data quality: good.
Literature
1. Куземко А.А., Яворська О.Г., Ворона Є.І., Чорна Г.А., Федорончук М.М. Ключові
території національного рівня на території Вінницької області та їх значення для
оптимізації мережі природно-заповідного фонду // Заповідна справа в Україні. – 2010.
– т. 16, вип. 1. – С. 88–93.
46
47
Ownership: state, private (minor).
Biogeographic regions: coninental.
Habitats. Level 1. C – 0.7%; D – 27.7%; E – 19.0%; F – 7.0%; G – 45.6%.
Habitats. Level 2. C1 Surface standing waters – 0.1%; C2 Surface running waters – 0.2%;
C3 Littoral zone of inland surface waterbodies – 0.4%; D4 Base-rich fens and calcareous
spring mires – 15.9%; D5 Sedge and reedbeds, normally without free-standing water –
11.8%; E3 Seasonally wet and wet grasslands – 17.6%; E5 Woodland fringes and clearings
and tall forb stands – 1.4%; F9 Riverine and fen scrubs – 7.0%; G1 Broadleaved deciduous
woodland – 33.3%; G3 Coniferous woodland – 8.5%; G4 Mixed deciduous and coniferous
woodland – 3.8%.
Futher habitat descripion. D4.1 Rich fens, including eutrophic tall-herb fens and
calcareous flushes and soaks; D5.1 Reedbeds normally without free-standing water; D5.2
Beds of large sedges normally without free-standing water; E3.4 Moist or wet eutrophic
and mesotrophic grassland; E5.4 Moist or wet tall-herb and fern fringes and meadows;
F9.2 Salix carr and fen scrub; G1.2 Mixed riparian floodplain and gallery woodland; G1.4
Broadleaved swamp woodland not on acid peat; G3.4 Pinus sylvestris woodland south of
the taiga; G4.4 Mixed Pinus sylvestris – Betula woodland; G4.7 Mixed Pinus sylvestris –
acidophilous Quercus woodland.
Land use: mowing/hay making – 6%; forestry – minor; nature conservation and research
– major.
Protected areas: overlaps (about 773 ha) with Dermansko-Ostrozkyi naional nature park,
overlaps (265 ha) with Bushchanskyi state botanical reserve; includes Zaplava Zbytynky
regional botanical reserve (112.5 ha), includes Boloto Kruhliak regional botanical reserve
(32.6 ha), includes Pivdennomosivskyi regional landscape reserve (17.7 ha), overlaps
(about 773 ha) with Emerald Site "Dermansko-Ostrozkyi Naional Nature Park".
Threats: abandonment/reducion of land management – low.
General description. Botom of the Zbytynka river valley with woods and fens. The main
vegetaion type is swamp forest dominated mainly by Alnus gluinosa. In the herb layer,
there prevail Carex appropinquata, Carex acuiformis, Filipendula vulgaris. Other typical
species are Ranunculus repens, Peucedanum palustre, Stachys palustris. Dominants of
non-forest mires are Carex rostrata, Carex lasiocarpa, Carex lava, Aulacomium palustre,
Drepanocladus aduncus, Hamatocaulis vernicosus, Calliergonella cuspidata. Mires with
dominance of Carex lasiocarpa, Sphagnum fallax, S. squarrosum, S. warnstorii occupy
a small area. Rich fens are dominated by Schoenus ferrugineus, Carex davalliana, Carex
hosiana, Carex lava, Molinia caerulea. On higher ground there is a Pinus sylvestris forest.
Botanical significance. Important area for rich fens and Ligularia sibirica, Liparis loeselii,
Saxifraga hirculus.
Criterion A
• Ligularia sibirica (L.) Cass. (L. bucovinensis Nakai); A(ii); rare; trend: unknown; species
data quality: medium; trend data quality: poor.
• Liparis loeselii (L.) Rich.; A(ii); rare; trend: unknown; species data quality: poor; trend
data quality: poor.
• Saxifraga hirculus L.; A(ii); rare; trend: unknown; species data quality: poor; trend data
quality: poor.
Criterion C
• D4.1 Rich fens, including eutrophic tall-herb fens and calcareous flushes and soaks;
area: 140 ha; trend: stable; area data quality: medium; trend data quality: medium.
Literature
1. Андрієнко Т.Л., Онищенко В.А., Дацюк В.В. НПП Дермансько-Острозький
// Фіторізноманіття національних природних парків України / Під заг. ред.
Т.Л. Андрієнко та В.А. Онищенка. – Київ: Науковий світ, 2003. – С. 206–214.
2. Головко О.В., Діхтярук В.М., Кальчук Г.В. Аналіз стану популяції Swertia perennis
L. (Gentianaceae) на території національного природного парку "ДерманськоОстрозький" // Труды Гос. Никит. ботан. сада. – 2013. – Т. 135. – С. 132–137.
3. Пашкевич Н.А., Блінкова О.І., Козинятко Т.А. Еколого-ценотичні особливості
популяції Schoenus ferrugineus L. на території Дермансько-Острозького національного
природного парку // Заповідна справа в Україні. – 2013. – 19 (1). – C. 86–88.
48
49
2. Польовий Є.В., Давидов Д.А. Синузія весняних ефемероїдів Бритавського лісу //
Наукові записки НаУКМА. – 2016. – Т. 184. – С. 67–71.
3. Яворська О.Г. НПП Кармелюкове Поділля // Фіторізноманіття заповідників і
національних природних парків України. Ч. 2. Національні природні парки / під ред.
В.А. Онищенка і Т.Л. Андрієнко. – Київ: Фітосоціоцентр, 2012. – С. 266–272.
Bushchanske Boloto
O.V. Holovko, V.A.Onyshchenko
Ukrainian name: Бущанське болото.
Transliteraion/Translaion variants: Bushchanske Fen.
Area: 880.0 ha.
Altitude: 206–230 m.
Latitude: 50°17'55" N (50.2987°).
Longitude: 26°12'40" E (26.2057°).
Administrative regions. Rivne region: Ostroh raion, Zdolbuniv raion
Byriuchyi Ostriv
V.P. Kolomiychuk, V.A. Onyshchenko
Ukrainian name: Бирючий острів.
Transliteraion/Translaion variants: Byryuchiy Island, Biriuchii Ostrov.
Area: 13406.1 ha.
Altitude: 0–5 m.
Latitude: 46°08'59" N (46.1498°).
Longitude: 35°08'24" E (35.1400°).
Administrative regions. Kherson region: Henichesk raion. Zaporizhia region: Yakymivka
raion. Territorial waters of Ukraine.
Ownership: state (major), private.
Biogeographic regions: steppic.
Habitats. Level 1. A– 44.3%; B – 1.2%; C – 4.0%; D – 11.0%; E – 37.0%; G – 1.0%; H – 1.5%.
Habitats. Level 2. A2 Littoral sediment – 5.0%; A5 Sublittoral sediment – 39.3%; B1 Coastal
dunes and sandy shores – 0.7%; B2 Coastal shingle – 0.5%; C1 Surface standing waters –
3.0%; C3 Littoral zone of inland surface waterbodies – 1.0%; D6 Inland saline and brackish
marshes and reedbeds – 11.0%; E1 Dry grasslands – 35.0%; E6 Salt steppes – 2.0%; G1
Broadleaved deciduous woodland – 1.0%; H5 Miscellaneous inland habitats with very
sparse or no vegetation – 1.5%.
Futher habitat description. A2.1 Littoral coarse sediment; A2.2 Littoral sand and muddy
sand; A2.4 Littoral mixed sediments; A2.5 Coastal saltmarshes and saline reedbeds;
B1.1 Sand beach driftlines; B1.2 Sand beaches above the driftline; B1.3 Shifting coastal
dunes; B1.4 Coastal stable dune grassland (grey dunes); B2.1 Shingle beach driftlines; B2.2
Unvegetated mobile shingle beaches above the driftline; C1.5 Permanent inland saline
and brackish lakes, ponds and pools; C1.6 Temporary lakes, ponds and pools; D6.2 Inland
saline or brackish species-poor helophyte beds normally without free-standing water; E1.2
Perennial calcareous grassland and basic steppes (E1.2G); E6.2 Coninental inland salt
steppes; G1.C Highly ariicial broadleaved deciduous forestry plantations, H5.6 Trampled
areas.
Land use: agriculture (animals) – major; forestry – minor; hunting – minor; mowing/hay
making – 3%; nature conservation and research – major; tourism/recreaion – minor.
Protected areas: overlaps (about 10407 ha) with Azovo-Syvaskyi naional nature park,
overlaps (about 1220 ha) with Pryazovskyi naional nature park, overlaps (about 10407
ha) with Emeral Site "Azovo-Syvaskyi Naional Nature Park", overlaps (about 1220 ha) with
Emeral Site "Pryazovskyi Naional Nature Park".
Threats: climate change/ sea level rise – low, development (recreaion/tourism) – low,
forestry (aforestaion) – low, natural events: spit erosion – medium.
General description. A large spit and adjacent sea. The spit consists of sand and shell. The
northwestern side is occupied by Phragmites australis (mainly), Bolboschoenus mariimus,
Juncus maritimus. A large area is covered by saline meadows dominated by Puccinellia
distans, Puccinellia gigantea, Lepidium laifolium, Elytrigia elongata, Elytrigia pseudocaesia,
Tripolium vulgaris, Juncus gerardii. On the elevated areas, there are sand steppes with
prevailng Festuca beckeri, Carex colchica, Ephedra distachya, Marrubium peregrinum,
Teucrium polium, Stipа borysthenica, Astragalus borysthenicus, Medicago kotovii. Other
species of high constancy are Agropyron lavrenkoanum, Agropyron pectinatum, Alyssum
hirsutum, Androsace elongata, Сentaurea diffusa, Cynodon dactylon, Secale sylvestre,
50
Silene subconica, Thymus dimorphus, Verbascum pinnatifidum. On the litoral, there
dominate Cakile euxina and Salsola ponica. On the beach ridges, dominant species are
Leymus sabulosus, Eryngium mariimum, Crambe maritima, Argusia sabirica, Euphorbia
seguieriana. In the most saline habitats, there dominate Salicornia perennans (mainly),
Suaeda prostrata, Limonium caspium, Halocnemum strobilaceum, Limonium meyeri. The
major dominant of the aquaic vegetaion is Zostera marina. It co-dominates with Zostera
noltii (Nanozostera nolii), Zannichellia pedunculata, Potamogeton pectinatum (Stuckenia
pecinata).
Botanical significance. Important area for conservaion of sand steppes, psammophyic
species, and brackish aquaic vegetaion.
51
Criterion A
• Agropyron cimmericum Nevski; A(i); abundance: occasional; trend: stable; species data
quality: medium; trend data quality: poor.
• Agropyron dasyanthum Ledeb.; A(i); abundance: occasional; trend: stable; species data
quality: medium; trend data quality: poor.
• Мedicago kotovii Wissjul.; A(i); abundance: occasional; trend: decreasing; species data
quality: medium; trend data quality: medium.
• Zostera marina L.; A(iv); abundance: abundant; trend: stable; species data quality:
good; trend data quality: poor.
Criterion C
• B1.1 Sand beach driftlines; area: 10 ha; trend: stable; area data quality: medium; trend
data quality: medium.
• B1.3 Shifting coastal dunes; area: 30 ha; trend: stable; area data quality: medium; trend
data quality: medium.
Conservaion proposals. Reduce the number of ungulates.
Literature
1. Dubyna D.V., Neuhauslova Z., Shelyag-Sosonco Ju.R. Coastal vegetation of the "Birjucij
Island" Spit in the Azov Sea, Ukraine // Preslia. – Praha, 1994. – Vol. 66. – P. 193–216.
2. Dubyna D.V., Neuhauslova Z., Shelyag-Sosonco Ju.R. Vegetaion of the "Birjucij Island"
Spit in the Azov Sea. Sand Steppe Vegetaion // Folia Geobot. Phytotax. – Praha, 1995. – Vol.
30. – P. 1–31.
3. Dubyna D.V., Neuhäuslová Z., Shelyag-Sosonco Ju.R Salt meadows of the Birjuchij
Islands Spit in the Azov Sea, Ukraine // Preslia. – Praha, 1996. – Vol. 68. – P. 51–62.
4. Dubyna D.V., Neuhauslova Z., Shelyag-Sosonco Ju.R. Salt meadows (FestucoPuccinellietea) of the "Birjucij Island" Spit in the Azov Sea, Ukraine // Preslia. – Praha, 2000.
– Vol. 72. – P. 31–48.
5. Dubyna D.V., Neuhäuslová Z. Salt meadows of the Birjuchij Islands Spit in the Azov
Sea. Classes Juncetea maritimi and Bolboschoenetea maritimi // Acta Bot. Croat. – 2000. –
Vol. 59, №1. – P. 167–178.
6. Dubyna D.V., Neuhäuslová Z. The vegetation of the Azov-Sivash National Nature Park.
Class Thero-Salicornietea (S. Pignatti 1953) R.Tx. in R. Tx. et Oberd. 1958 // Thaiszia. – J. Bot.
(Kosice). – 2003. – Vol. 13, №1. – P. 1–30.
7. Коломійчук В.П. Ключові ботанічні території Північного Приазов'я // Заповідна
справа в Україні. – 2008. – Т. 14, вип. 1. – С. 61–66.
8. Коломійчук В.П. НПП Приазовський // Фіторізноманіття заповідників і
національних природних парків України. Ч. 2. Національні природні парки / під ред.
В.А. Онищенка і Т.Л. Андрієнко. – Київ: Фітосоціоцентр, 2012. – С. 5–26.
9. Тищенко О.В. Рослинність приморських кіс північного узбережжя Азовського
моря. – Київ: Фітосоціоцентр, 2006. – 156 с.
10. Фіторізноманіття національних природних парків України / Під заг. ред.
Т.Л. Андрієнко та В.А. Онищенка. – Київ: Науковий світ, 2003. – 143 с.
52
Cheremske Boloto
V.A. Onyshchenko
Ukrainian name: Черемське болото.
Transliteraion/Translaion variants: Cheremske Mire.
Area: 4205 ha.
Altitude: 157–160 m.
Latitude: 51°31’52” N (51.5310°).
Longitude: 25°33’20” E (25.5555°).
Administrative regions. Rivne region: Volodymyrets raion; Volynska region: Manevychi
raion.
Ownership: state.
Biogeographic regions: coninental.
Habitats. Level 1. C – 0.5%; D – 34%; F – 1%; G – 64.5%.
Habitats. Level 2. C1 Surface standing waters – 0.5%; D1 Raised and blanket bogs – 1%;
D2 Valley mires, poor fens and transition mires – 32%; D5 Sedge and reedbeds, normally
without free-standing water – 1%; F9 Riverine and fen scrubs – 1%; G1 Broadleaved
deciduous woodland – 15%; G3 Coniferous woodland – 45.5%; G4 Mixed deciduous and
coniferous woodland – 4%.
Futher habitat description. C1.2 Permanent mesotrophic lakes, ponds and pools; C1.4
Permanent dystrophic lakes, ponds and pools; C3.2 Water-fringing reedbeds and tall
helophytes other than canes; D1.1 Raised bogs; D2.3 Transiion mires and quaking bogs;
53
D5.1 Reedbeds normally without free-standing water; D5.2 Beds of large sedges normally
without free-standing water; F9.2 Salix carr and fen scrub; G1.4 Broadleaved swamp
woodland not on acid peat; G1.5 Broadleaved swamp woodland on acid peat; G1.9 Nonriverine woodland with Betula, Populus tremula or Sorbus aucuparia; G1.B Non-riverine
Alnus woodland; G3.1 Abies and Picea woodland; G3.4 Pinus sylvestris woodland south of
the taiga; G3.E Nemoral bog conifer woodland; G4.1 Mixed swamp woodland; G4.4 Mixed
Pinus sylvestris – Betula woodland; G4.7 Mixed Pinus sylvestris – acidophilous Quercus
woodland.
Land use: forestry – minor, nature conservation and research – 71%.
Protected areas: includes Cheremskyi nature reserve (2975.7 ha), includes Emerald Site
“Cheremskyi Nature Reserve” (2975.7 ha).
Threats: burning of vegetaion – low, water (drainage) – low.
General description. Transiion mire and surrounding forest. The mire is dominated by Carex
rostrata, Menyanthes trifoliata, Vaccinium oxycoccos (Oxycoccus palustris), Phragmites australis, Carex appropinquata, Carex lasiocarpa, Sphagnum fallax, oten with the open layer
of Betula pubescens. Raised bogs with oppressed Pinus sylvestris occupy about 50 ha. Pinus
sylvestris forests occupy the largest area. Typical dominants are Vaccinium myrillus, Pleurozium schreberi, Dicranum polysetum. In drier habitats, there dominate lichens (Cladonia alpestre, C. rangiferina, C. sylvaica, Cetraria islandica). In wet habitats the main dominant of
the herb layer is Molinia caerulea. In swamp pine forests, there dominate Eriophorum vaginatum, Ledum palustre, Vaccinium oxycoccos (Oxycoccus palustris), Sphagnum nemoreum,
S. fallax, S, centrale. Swampy Alnus gluinosa forests are dominated by Carex riparia, Carex
acuiformis, Scirpus sylvaicus, Phragmites australis. In more mesic alder forests, dominant
species are Athyrium ilix-femina, Vaccinium myrillus, Deschampsia cespitosa. The IPA includes mesotrophic lakes Redychi (14 ha) and Cheremske (7.7 ha).
Botanical significance. One of the largest transiion mires in Ukraine.
Criterion C
• D2.3 Transition mires and quaking bogs; area: 900 ha; trend: stable; area data quality:
medium; trend data quality: poor.
• G1.51 Sphagnum Betula woods; area: 500 ha; trend: stable; area data quality: poor;
trend data quality: poor.
• G3.1F Enclave Picea abies forests; area: 15 ha; trend: stable; area data quality: medium;
trend data quality: poor.
Literature
1. Андриенко Т.Л.,Коніщук В.В. Карта рослинності Черемського природного заповідника // Укр. ботан. журн. – 2003. – 60, № 6. – С. 659–669.
2. Коніщук В.В. Рідкісні види рослин Черемського природного заповідника // Укр.
ботан. журн. – 2003. – 60, № 3. – С. 264–272.
3. Коніщук В.В. Хвойні ліси Черемського природного заповідника // Рослинність
хвойних лісів України (матеріали робочої наради, Київ, листопад 2003 р). – К.: Фітосоціоцентр, 2003. – С. 111–122.
4. Коніщук В.В. Нові місцезнаходження рідкісних рослин у Черемському природному заповіднику // Заповідна справа в Україні. – Канів, 2004. – Т. 10, Вип. 1–2. – С. 18–23.
5. Коніщук В.В. Раритетна компонента біорізноманіття Черемського природного заповідника // Науковий вісник Волинського державного університету імені Лесі Українки № 11 (Ч. ІІ). – Луцьк: РВВ “Вежа”, 2007 р. – С. 125–132.
6. Коніщук В.В. ПЗ Черемський // Фіторізноманіття заповідників і національних
природних парків України. Ч.1. Біосферні заповідники. Природні заповідники / під ред.
54
В.А. Онищенка і Т.Л. Андрієнко. – Київ, 2012. – С. 379–389
7. Коніщук В.В., Коновальчук В.К., Парчук Г.В. Черемське болото – потенційне Рамсарське угіддя: проблеми і перспективи охорони // Вісник національного університету водного господарства та природокористування (збірник наукових праць), Випуск 1
(33). – Рівне, 2006. – С. 28–35.
8. Прядко Е.И. Растительность и стратиграфия болота Черемошского в Западном
Полесье / Вопросы физиологии, биохимии, цитологии и флоры Украины. – К.: Наук.
думка, 1974. – C. 28–53.
9. Фіторізноманіття Українського Полісся та його охорона / Під заг. ред. Т.Л. Андрієнко. – Київ: Фітосоціоцентр, 2006. – 316 с.
Cherevaskyi Lis
V.A. Onyshchenko
Ukrainian name: Череваський ліс.
Transliteraion/Translaion variants: Cherevaskyi Forest.
Area: 466.0 ha.
Altitude: 171–188 m.
Latitude: 51°17’35” N (51.29305°).
Longitude: 25°26’05” E (25.43465°).
Administrative regions. Volynska region: Manevychi raion.
Ownership: state.
Biogeographic regions: coninental.
Habitats. Level 1. E – 1.0%; G – 98.0%; H – 0.8%; J – 0.2%.
Habitats. Level 2. G1 Broadleaved deciduous woodland – 13.0%; G3 Coniferous woodland
– 70.0%; G4 Mixed deciduous and coniferous woodland – 15.0%; H5 Miscellaneous inland
habitats with very sparse or no vegetation – 0.8%; J4 Transport networks and other
constructed hard-surfaced areas – 0.2%.
Futher habitat descripion. G1.4 Broadleaved swamp woodland not on acid peat; G1.9
Non-riverine woodland with Betula, Populus tremula or Sorbus aucuparia; G3.1 Abies and
Picea woodland; G3.4 Pinus sylvestris woodland south of the taiga; G4.1 Mixed swamp
woodland; G4.4 Mixed Pinus sylvestris – Betula woodland; H5.6 Trampled areas.
Land use: forestry – major.
Protected areas: included in Emerald Site “Cherevaskyi Lis”.
Threats: forestry (intensiied forest management) – medium.
General description. Forest dominated by Pinus sylvestris, Betula pendula, Alnus gluinosa,
Picea abies.
Botanical significance. The largest locaion of Picea abies forests in the latland part of
Ukraine.
Criterion C
• G3.1F Enclave Picea abies forests; area: 200 ha; trend: decreasing; area data quality:
poor; trend data quality: poor.
Conservaion proposals. Create a state botanical reserve (or a state forest reserve).
55
scrub – 0.3%; G1 Broadleaved deciduous woodland – 98.1%; H2 Screes – 0.0%; H3 Inland
cliffs, rock pavements and outcrops – 0.0%; H5 Miscellaneous inland habitats with very
sparse or no vegetation – 0.5%.
Literature
1. Мельник В.І. Острівні ялинники Українського Полісся. – К.: Наукова думка, 1993.
– 104 с.
Chorna Hora
R.Ya. Kish, V.A. Onyshchenko
Ukrainian name: Чорна гора.
Transliteraion/Translaion variants: Black Mount, Black Hill.
Area: 810 ha.
Altitude: 170–565 m.
Latitude: 48°09’14” N (48.1538°).
Longitude: 23°04’18” E (23.0716°).
Administrative regions. Zakarpatska region: Vynohradiv raion.
Ownership: state.
Biogeographic regions: pannonian.
Habitats. Level 1. E – 1.1%; F – 0.3%; G – 98.1%; H – 0.5%.
Habitats. Level 2. E1 Dry grasslands – 1.1%; F3 Temperate and mediterranean-montane
56
Futher habitat description. E1.2 Perennial calcareous grassland and basic steppes –
0.3%; F3.1 Temperate thickets and scrub; F3.2 Submediterranean deciduous thickets and
brushes; G1.6 Fagus woodland; G1.7 Thermophilous deciduous woodland; G1.A Meso- and
eutrophic Quercus, Carpinus, Fraxinus, Acer, Tilia, Ulmus and related woodland; H2.3 Temperate-montane acid siliceous screes; H3.1 Acid siliceous inland clifd; H5.6 Trampled areas.
Land use: forestry – minor, nature conservation and research – 93%.
Protected areas: overlaps (747 ha) with Carpathian biosphere reserve (UNESCO) (includes
cluster “Chorna Hora” of the biosphere reserve), overlaps (747 ha) with the Karpatskyi
biosphere reserve (naional category), includes state botanical reserve “Chorna Hora” (747
ha, belongs to the biosphere reserve), overlaps (747 ha) with Emerald Site “Carpathian Biosphere Reserve.
Threats: abandonment/reducion of land management – low.
General description. A hill composed of tuf, andesite, rhyolite, and sedimentary rocks. On
the southern slopes, the major vegetaion type is the termophilous oak wood. The tree
layer is composed of Quercus petraea, someimes with Quercus dalechampii, Quercus polycarpa, Tilia tomentosa, Fraxinus ornus (single locality in Ukraine). In the shrub layer, there
prevail Cornus mas, Crataegus monogyna s. l., Euonymus europaeus, Ligustrum vulgare.
Typical species of the herb layer are Aegonychon purpureo-caeruleum, Ajuga genevensis,
Anthericum ramosum, Astragalus glycyphyllos, Brachypodium pinnatum, Carex michelii,
Clinopodium vulgare, Euphorbia cyparissias, Euphorbia polychroma, Hylotelephium polonicum, Hypericum perforatum, Inula hirta, Lathyrus niger, Melica picta, Melica unilora,
57
Melittis melissophyllum, Poa nemoralis, Polygonatum odoratum, Stellaria holostea, Veronica chamaedrys, Veronica oicinalis, Vincetoxicum hirundinaria. On the nothern slopes there
are Fagus sylvaica and mixed Fagus sylvaica – Quercus petraea forests (with Asarum europaeum, Brachypodium sylvaicum, Carex pilosa, Dentaria bulbifera, Galium odoratum, Geranium roberthianum, Lamium galeobdolon, Mercurialis perennis). Grasslands occupy a small
area. Dominant species are Festuca pseudodalmaica and Brachypodium pinnatum, there
are also Anchusa barellieri, Carduus collinus, Cruciata pedemontana, Ferulago sylvaica,
Galium campanulatum, Melica transsilvanica, Phleum phleoides, Potenilla recta, Scabiosa
ochroleuca, Sipa transcarpathica, Teucrium chamaedrys, Thymus pannonicus, Tordylium
maximum, Trifolium alpestre, Veronica spicata.
Botanical significance. The largest locaion of steppe vegetaion and one of the largest locaions of termophilous oak woods in the Transcarpathian part of Ukraine.
Criterion C
• E1.2 Perennial calcareous grassland and basic steppes (*6240); area: 9 ha; trend:
decreasing; area data quality: medium; trend data quality: medium.
• F3.241 Central European subcontinental thickets; area: 5 ha; trend: stable; area data
quality: medium; trend data quality: medium.
• G1.7 Thermophilous deciduous woodland (*91H0, 91M0); area: 400 ha; trend: stable;
area data quality: medium; trend data quality: medium.
Conservaion proposals. Do not allow the expansion of shrubs and trees in grasslands.
Literature
1. Стойко С.М. Дубові ліси Українських Карпат: екологічні особливості, відтворення,
охорона. – Львів, 2009. – 220 с.
2. Фіторізноманіття заповідників і національних природних парків України.
Ч.1. Біосферні заповідники. Природні заповідники / Колектив авторів під ред. В.А.
Онищенка і Т.Л. Андрієнко. – Київ: Фітосоціоцентр, 2012. – 406 с.
pavements and outcrops – 0.0%; H5 Miscellaneous inland habitats with very sparse or no
vegetation – 0.5%.
Chornohora
V.A. Onyshchenko, V.M. Virchenko
Ukrainian name: Чорногора.
Area: 43124.0 ha.
Altitude: 570–2061 m.
Latitude: 48°08’03” N (48.1341°).
Longitude: 24°32’45” E (24.5458°).
Administrative regions. Ivano-Frankivsk region: Verkhovyna raion, Yaremche city;
Zakarpatska region: Rakhiv raion.
Ownership: state.
Biogeographic regions: alpine.
Habitats. Level 1. C – 0.2%; D – 0.1%; E – 16.0%; F – 6.2%; G – 77.0%; H – 0.5%.
Habitats. Level 2. C1 Surface standing waters – 0.0%; C2 Surface running waters – 0.2%; E1
Dry grasslands – 0.1%; E2 Mesic grasslands – 0.1%; E3 Seasonally wet and wet grasslands
– 0.1%; E4 Alpine and subalpine grasslands stands – 10.1%; E5 Woodland fringes and
clearings and tall forb stands – 5.7%; F2 Arctic, alpine and subalpine scrub – 6.2%; G1
Broadleaved deciduous woodland – 6.5%; G3 Coniferous woodland – 35.3%; G4 Mixed
deciduous and coniferous woodland – 35.2%; H2 Screes – 0.0%; H3 Inland cliffs, rock
58
Futher habitat description. C1.1 Permanent oligotrophic lakes, ponds and pools; C1.2
Permanent mesotrophic lakes, ponds and pools; C2.1 Springs, spring brooks and geysers;
C2.2 Permanent non-tidal, fast, turbulent watercourses; E1.7 Non-Mediterranean dry acid
and neutral closed grassland; E2.2 Low and medium alitude hay meadows; E3.4 Moist or
wet eutrophic and mesotrophic grassland; E4.1 Vegetated snow-patch; E4.2 Moss and
lichen dominated mountain summits, ridges and exposed slopes; E4.3 Acid alpine and
subalpine grassland; E4.4 Calcareous alpine and subalpine grassland; E5.4 Moist or wet tallherb and fern fringes and meadows; E5.5 Subalpine moist or wet tall-herb and fern stands;
F2.2 Evergreen alpine and subalpine heath and scrub; F2.3 Subalpine deciduous scrub; F2.4
Conifer scrub close to the tree limit; G1.1 Riparian and gallery woodland, with dominant
Alnus, Betula, Populus or Salix; G1.6 Fagus woodland; G3.1 Abies and Picea woodland; G4.6
Mixed Abies – Picea – Fagus woodland; H2.3 Temperate-montane acid siliceous screes;
H2.4 Temperate-montane calcareous and ultra-basic screes; H3.1 Acid silicious inland clifs;
59
H3.2 Basic and ultra-basic inland clifs; H5.6 Trampled areas.
Land use: agriculture (animals) – minor, forestry – minor, mowing/hay making – minor,
nature conservation and research – major, tourism/recreaion – minor.
Protected areas: overlaps (about 14537 ha) with Karpatskyi (Carpathian) biosphere reserve
(naional category), overlaps (about 14537 ha) with Carpathian biosphere reserve (UNESCO),
overlaps (about 14537 ha) with Emerald Site “Carpathian Biosphere Reserve”, overlaps
(about 26269 ha) with Karparskyi (Carpathian) naional nature park, overlaps (15402 ha)
with World Heritage Site “Primeval Beech Forests of the Carpathians and the Ancient Beech
Forests of Germany” (core zone: 2477 ha, bufer zone: 12925 ha), overlaps (about 26269 ha)
with Emerald Site “Carpathian Naional Nature Park”.
Threats: climate change/sea level rise – low, development (recreaion/tourism) – low.
General description. The area includes the highest part of the Ukrainian Carpathians. Major
vegetaion type is mixed Picea abies – Fagus sylvaica – Abies alba forests. The herb layer is
dominated by Athyrium ilix-femina, Dryopteris dilatata, D. carthusiana, D. ilix-mas, Lamium
galeobdolon, Calamagrosis arundinacea, Oxalis acetosella, Rubus hirtus. Other frequent
species are Anemone nemorosa, Dentaria glandulosa (Cardamine glandulosa), Geniana
asclepiadea, Homogyne alpina, Luzula luzuloides, Luzula sylvaica, Symphytum cordatum,
Vaccinium myrillus. Pure Fagus sylvaica forests of similar species composiion occupy a
smaller area. Natural pure Picea abies forests predominate at higher alitudes and form
the upper limit of the forest vegetaion. Dominants of their herb layer are Calamagrosis
arundinacea, Luzula sylvaica, Oxalis acetosella, Vaccinium myrillus. Other frequent species
are Athyrium distenifolium, Homogyne alpina, Leucanthemum rotundifolium, Rubus
idaeus, Senecio ovatus, Soldanella hungarica, Sorbus aucuparia, Streptopus amplexifolium,
Vaccinium viis-idaea. The moss layer is dominated by Dicranum scoparium, Pleurozium
schreberi, Polytrichum formosum. In the grasslands of the forest belt, there predominate
Agrosis tenuis, Festuca rubra and Nardus stricta. There occur poor and rich fens. In the
subalpine belt, the largest area is occupied by Pinus mugo communiies. Besides there are
shrubs dominated by Duschekia alnobetula (Alnus alnobetula), Juniperus sibirica. Other
important subalpine dominants are Nardus stricta, Deschampsia cespitosa, Vaccinium
myrillus, Festuca airoides (F. supina), Calamagrosis villosa. Rhododendron myrifolium
heaths are less common. In the wet habitats, there occur communiies of Adenostyles
alliariae, Cirsium waldsteinii. There are small areas of bogs, transiion mires, poor and
rich fens. On the silicate rocks, there prevail Festuca picta and Poa deylii. Calcareous rocky
grasslands dominated by Festuca inarmata (Festuca amethysina s.l.) or Festuca versicolor
cover small areas. At the highest alitudes (>1800 m), the typical dominants are Oreojuncus
triidus (Juncus triidus), Carex curvula, Carex sempervirens.
Botanical significance. The most important area for subalpine habitats and species in
Ukraine. Important for oligotrophic aquaic habitats and some species of mosses.
Criterion A
• Anacamptodon splachnoides (Brid.) Brid.; A(ii); abundance: rare; trend: unknown;
species data quality: medium; trend data quality: poor.
• Botrychium matricariifolium (A. Braun ex Döll) W.D.J.Koch; A(ii); abundance: rare; trend:
unknown; species data quality: medium; trend data quality: poor.
• Buxbaumia viridis (Moug. ex Lam. et DC.) Brid. ex Moug. et Nestl. (B. indusiata Brid.);
A(ii); abundance: rare; trend: unknown; species data quality: poor; trend data quality:
poor.
• Campanula abieina Griseb. & Schenk; A(ii); abundance: occasional; trend: stable;
species data quality: medium; trend data quality: poor.
60
•
Campanula serrata (Kit.) Hendrych; A(ii); abundance: rare; trend: unknown; species
data quality: medium; trend data quality: poor.
• Dicranum viridae (Sull. et Lesq.) Lindb.; A(ii); abundance: occasional; trend: unknown;
species data quality: medium; trend data quality: poor.
• Festuca porcii Hack.; A(iv); abundance: rare; trend: unknown; species data quality:
medium; trend data quality: poor.
• Hamatocaulis vernicosus (Mitt.) Hedenas (Drepanocladus vernicosus (Mitt.) Warnst.);
A(ii); abundance: occasional; trend: unknown; species data quality: medium; trend data
quality: poor.
• Heterophyllium affine (Mitt.) Fleisch; A(ii); abundance: rare; trend: unknown; species
data quality: medium; trend data quality: poor.
• Mannia triandra (Scop.) Grolle; A(ii); abundance: rare; trend: unknown; species data
quality: poor; trend data quality: poor.
• Neckera pennata Hedw.; A(ii); abundance: rare; trend: unknown; species data quality:
poor; trend data quality: poor.
• Tozzia carpathica Woł.; A(ii); abundance: rare; trend: unknown; species data quality:
medium; trend data quality: poor.
Criterion C
• C1.1 Permanent oligotrophic lakes, ponds and pools; area: 1 ha; trend: stable; area data
quality: medium; trend data quality: medium.
• C2.12 Hard water springs; area: 0.1 ha; trend: stable; area data quality: poor; trend data
quality: poor.
• D2.226 Peri-Danubian black-white-star sedge fens; area: 5 ha; trend: stable; area data
quality: poor; trend data quality: poor.
• E4.11 Boreo-alpine acidocline snow-patch grassland and herb habitats; area: 10 ha;
trend: stable; area data quality: poor; trend data quality: poor.
• E4.12 Boreo-alpine calcicline snow-patch grassland and herb habitats; area: 1 ha; trend:
stable; area data quality: poor; trend data quality: poor.
• E4.3 Acid alpine and subalpine grassland; area: 4000 ha; trend: stable; area data quality:
medium; trend data quality: poor.
• E4.4 Calcareous alpine and subalpine grassland; area: 400 ha; trend: stable; area data
quality: poor; trend data quality: poor.
• E5.5 Subalpine moist or wet tall-herb and fern stands; area: 2500 ha; trend: stable; area
data quality: medium; trend data quality: poor.
• F2.22 Alpide acidocline Rhododendron heaths; area: 30 ha; trend: stable; area data
quality: poor; trend data quality: poor.
• F2.46 (*4070) Carpathian Pinus mugo scrub; area: 2100 ha; trend: stable; area data
quality: good; trend data quality: poor.
• F4.2 Dry heaths; area: 10 ha; trend: stable; area data quality: poor; trend data quality:
medium.
• G1.12 Boreo-alpine riparian galleries; area: 100 ha; trend: stable; area data quality:
medium; trend data quality: medium.
• G3.1B Alpine and Carpathian subalpine Picea forests; area: 2000 ha; trend: increasing;
area data quality: medium; trend data quality: medium.
• H2.3 Temperate-montane acid siliceous screes; area: 40 ha; trend: stable; area data
quality: poor; trend data quality: medium.
Literature
1. Андриенко Т.Л., Каркуциев Г.Н., Прядко Е.И. Гидрофильная растительность
61
верховьев р. Прут // Гидробиол. журн. – 1991. – 27, № 5. – С. 16–22.
2.Біорізноманіття Карпатського біосферного заповідника / Кол. авт., Ред. рада:
Я.І. Мовчан, Ф.Д. Гамор та ін. – К.: Інтерекоцентр, 1997. – 711 с.
3.Гамор Ф.Д., Волощук М.І., Антосяк Т.М., Козурак А.В. БЗ Карпатський //
Фіторізноманіття заповідників і національних природних парків України. Ч. 1. Біосферні
заповідники. Природні заповідники / під ред. В.А. Онищенка і Т.Л. Андрієнко. – Київ:
Фітосоціоцентр, 2012. – С. 45–72.
4.Карпатський національний природний парк // Киселюк О.І., Приходько М.М.,
Яворський А.І. та ін. / За ред. М.М.Приходька, О.І. Киселюка, А.І. Яворського. – ІваноФранківськ: Фоліант, 2009. – 672 с.
5.Малиновський К.А. Рослинність високогір’я Українських Карпат. – К.: Наук. думка,
1980. – 280 с.
6.Малиновський К.А., Крічфалушій В.В. Рослинні угруповання високогір’я
Українських Карпат. – Ужгород, 2002. – 244 с.
7.Нестерук Ю. Рослинний світ Українських Карпат: Чорногора. Екологічні мандрівки.
– Львів: БаК, 2003. – 520 с.
8.Онищенко В.А., Данилик І.І. НПП Карпатський // Фіторізноманіття заповідників і
національних природних парків України. Ч. 2. Національні природні парки / під ред.
В.А. Онищенка і Т.Л. Андрієнко. – Київ: Фітосоціоцентр, 2012. – С. 273–292.
9.Природа Карпатського національного парку. – К.: Наук. думка, 1993. – 216 с.
10. Флора і рослинність Карпатського заповідника / Стойко С.М., Тасенкевич Л.О.,
Мілкіна Л.І. та ін. – К.: Наук. думка, 1982. – 220 с.
Threats: forestry (intensiied forest management) – low; development (recreaion/tourism)
– low.
Chornyi Lis
V.A. Onyshchenko
Ukrainian name: Чорний ліс.
Transliteraion/Translaion variants: Black Woods, Black Forest.
Area: 6909 ha.
Altitude: 168–218 m.
Latitude: 48°46’20” N (48.7721°).
Longitude: 32°34’21” E (32.5726°).
Administrative regions. Kirovohrad region: Znamianka raion.
Ownership: state.
Biogeographic regions: coninental.
Habitats. Level 1. C – 0.1%; D – 0.1%; E – 0.1%; F – 0.2%; G – 98.3%; H – 1.0%; J – 0.2%.
Habitats. Level 2. C1 Surface standing waters – 0.1%; D5 Sedge and reedbeds, normally
without free-standing water – 0.1%; E2 Mesic grasslands – 0.1%; G1 Broadleaved deciduous
woodland – 98.3%; H5 Miscellaneous inland habitats with very sparse or no vegetation –
1.0%; J4 Transport networks and other constructed hard-surfaced areas – 0.2%.
Futher habitat description. G1.A Meso- and eutrophic Quercus, Carpinus, Fraxinus, Acer,
Tilia, Ulmus and related woodland; H5.6 Trampled areas.
Land use: forestry – major, nature conservation and research – 46%.
Protected areas: overlaps with Chornoliskyi state landscape reserve (about 3200 ha),
includes Chornoliska state hydrological nature monument (2 ha), included in Emerald Site
“Znamianskyi Chornyi Lis”.
62
General description. One of the largest woods in the southern forest-steppe zone of
Ukraine. Almost the entire territory is occupied by the dark broadleaved forest. The tree
layer is dominated by Quercus robur, Fraxinus excelsior, Carpinus betulus, sometimes
by Acer campestre, Acer platanoides, Tilia cordata. Main dominants of the herb layer in
summer are Aegopodium podagraria, Stellaria holostea, on slopes oten also Carex pilosa.
In spring there prevail Allium ursinum, Anemone ranunculoides, Corydalis cava, Corydalis
solida, Ficaria verna. Other typical species are Alliaria petiolata, Asarum europaeum, Corylus
avellana, Euonymus europaeus, Euonymus verrucosus, Galium aparine, Gagea lutea, Galium
odoratum, Geum urbanum, Polygonatum multiflorum, Pulmonaria obscura, Scilla siberica,
Ulmus glabra, Urtica dioica, Viola odorata. An interesting object in this area is a small mire
which is one of the southernmost localities of Sphagnum in Ukraine (Chornoliska state
hydrological nature monument).
Botanical significance. Rather well preserved broadleaved forest, one of the best in Ukraine.
Criterion C
63
•
G1.A1 Quercus – Fraxinus – Carpinus betulus woodland on eutrophic and mesotrophic
soils; area: 5900 ha; trend: stable; area data quality: good; trend data quality: good.
Conservaion proposals. Create an Emerald Site. Do not allow clearcuing.
Literature
1. Мирза-Сіденко В.М., Андрієнко Т.Л., Онищенко В.А., Прядко О.І. Флора і рослинність проектованого Чорнолісько-Дмитрівського національного природного парку //
Укр. ботан. журн. – 2008. – 65, № 3. – С. 352–369.
2. Наумович Г.О., Ходосовцев О.Є. Лишайники Чорного лісу (Знам’янський район,
Кіровоградська область) // Чорноморський ботанічний журнал. – 2008. – 4, № 1. –
С. 7–13.
3. Онищенко В.А., Андрієнко Т.Л. Майбутній національний парк // Заповідні куточки Кіровоградської землі. – К.: Арктур-A, 1999. – С. 131–134.
4. Онищенко В.А., Сіденко В.М. Класифікація лісової рослинності ур. Чорний ліс
(Знам’янський район Кіровоградської області) // Наук. вісник Чернівецького університету. – Вип. 145. – Сер.: Біологія, 2002. – С. 178–194.
Chortova Hora
Ukrainian name: Чортова гора.
Transliteraion/Translaion variants: Devil’s Hill.
Area: 12 ha.
V.A. Onyshchenko, N.V. Shumska
Ownership: state.
Biogeographic regions: coninental
Habitats. Level 1. E – 100.0%; H – 0.0%.
Habitats. Level 2. E1 Dry grasslands – 99.0%; E7 Sparsely wooded grasslands – 1.0%; H3
Inland cliffs, rock pavements and outcrops – 0.0%.
Futher habitat descripion. E1.2 Perennial calcareous grassland and basic steppes; E7.2
Sub-coninental parkland; H3.2 Basic and ultra-basic inland cliffs.
Land use: nature conservation and research – major.
Protected areas: same as Chortova Hora state botanical nature monument.
Threats: abandonment/reducion of land management – low.
General description. A hill with grasslands dominated by Brachypodium pinnatum, Elytrigia
repens, Festuca valesiaca, with presence of Filipendula vulgaris, Fragaria viridis, Salvia
pratensis.
Botanical significance. The largest populaion of Carlina cirsioides in Ukraine.
Criterion A
• Carlina cirsioides Klokov; A(iv); abundance: frequent (5000 individuals); trend: stable;
species data quality: good; trend data quality: poor.
Literature
1. Куковиця Г.С., Дідух Я.П., Шеляг-Сосонко Ю.Р., Абдулоєва О.С. Синтаксономія
лучних степів пам’яток природи республіканського значення гг. Касова та Чортова //
Укр. фітоцен. зб. – 1998. – Сер. А, Вип. 2 (11). – Київ: Фітосоціоцентр. – С. 42–62.
2. Шумська Н.В., Дмитраш І.І. Поширення та еколого-ценотичні особливості Carlina
cirsioides Klok. і Carlina onopordifolia Besser ex Szafer, Kulcz. et Pawł. на Бурштинському
Опіллі // Флорологія та фітосозологія. – Т. 2. – Київ: Фітон, 2011. – С. 77–81.
Chyvchyny
Altitude: 290–350 m.
Latitude: 49°24’06” N (49.4017°).
Longitude: 24°39’54” E (24.6649°).
Administrative regions. Ivano-Frankivsk region: Rohatyn raion.
64
I.I. Chorney, V.A. Onyshchenko
Ukrainian name: Чивчини.
Area: 23177.0 ha.
Altitude: 970–1769 m.
Latitude: 47°48’23” N (47.8063°).
Longitude: 24°52’41” E (24.8780°).
Administrative regions. Chernivtsi region: Putyla raion; Ivano-Frankivsk region: Verkhovyna
raion.
Ownership: state (major), private.
Biogeographic regions: alpine.
Habitats. Level 1. C – 0.3%; D – 1.0%; E – 15.4%; F – 2.8%; G – 80.2%; H – 0.3%.
Habitats. Level 2. C2 Surface running waters – 0.3%; D4 Base-rich fens and calcareous
spring mires – 0.8%; D5 Sedge and reedbeds, normally without free-standing water – 0.2%;
E1 Dry grasslands – 4.8%; E2 Mesic grasslands – 1.4%; E4 Alpine and subalpine grasslands
stands – 2.0%; E5 Woodland fringes and clearings and tall forb stands – 7.2%; F2 Arctic,
alpine and subalpine scrub – 2.8%; G1 Broadleaved deciduous woodland – 1.0%; G3
Coniferous woodland – 76.2%; G4 Mixed deciduous and coniferous woodland – 3.0%; H2
Screes – 0.1%; H3 Inland cliffs, rock pavements and outcrops – 0.0%; H5 Miscellaneous
inland habitats with very sparse or no vegetation – 0.2%.
65
Futher habitat description. C2.1 Springs, spring brooks and geysers; C2.2 Permanent
non-tidal, fast, turbulent watercourses; E1.7 Non-Mediterranean dry acid and neutral
closed grassland (E1.71 Nardus stricta swards, E1.72 Agrosis – Festuca grassland); D4.1
Rich fens, including eutrophic tall-herb fens and calcareous lushes and soaks; D5.2 Beds
of large sedges normally without free-standing water; E3.4 Moist or wet eutrophic and
mesotrophic grassland; E4.1 Vegetated snow-patch; E4.2 Moss and lichen dominated
mountain summits, ridges and exposed slopes; E4.3 Acid alpine and subalpine grassland;
E4.4 Calcareous alpine and subalpine grassland; E5.4 Moist or wet tall-herb and fern fringes
and meadows; E5.5 Subalpine moist or wet tall-herb and fern stands; F2.2 Evergreen alpine
and subalpine heath and scrub; F2.3 Subalpine deciduous scrub; F2.4 Conifer scrub close to
the tree limit; G1.1 Riparian and gallery woodland, with dominant Alnus, Betula, Populus
or Salix; G3.1 Abies and Picea woodland; G4.6 Mixed Abies – Picea – Fagus woodland; H2.3
Temperate-montane acid siliceous screes; H2.4 Temperate-montane calcareous and ultrabasic screes; H3.1 Acid silicious inland cliffs; H3.2 Basic and ultra-basic inland cliffs; H3.5
Almost bare rock pavements, including limestone pavements; H5.6 Trampled areas.
Land use: agriculture (animals) – minor, forestry – minor, nature conservation and research
– major.
Protected areas: includes Verkhovynskyi national nature park (12022.9 ha), overlaps (about
6540 ha) with Cheremoskyi national nature park, includes Chornyi Dil state landscape
reserve (263 ha), overlaps (about 6300 ha) with Chyvchyno-Hryniavskyi regional landscape
reserve, overlaps (about 5890 ha) with Emerald Site “Cheremoskyi”, overlaps (about 14000
ha) with Emerald Site “Verkhovynskyi”.
Threats: abandonment/reducion of land management – low, climate change/sea level rise
– low.
General description. The major vegetaion type is pure Picea abies forest. On rich soils,
there dominate Anemone nemorosa, Dentaria glandulosa (Cardamine glandulosa),
Calamagrosis arundinacea, Calamagrosis villosa, Oxalis acetosella. Species of signiicant
constancy in spruce forests are Athyrium distenifolium, Athyrium ilix-femina, Dicranum
scoparium, Dryopteris dilatata, Dryopteris ilix-mas, Euphorbia carniolica, Gymnocarpium
dryopteris, Homogyne alpina, Leucanthemum roundifolium, Lonicera nigra, Phegopteris
connecilis, Polytrichum formosum, Senecio ovatus, Solidago virgaurea, Sorbus aucuparia,
Stellaria nemorum, Streptopus amplexifolius, Symphytum cordatum, Vaccinium myrillus.
In some places over the Picea abies belt, there is Pinus mugo scrub. Mesic grasslands of
the forest belt are dominated by Cynosurus cristatus, Festuca rubra, Festuca pratensis,
Agrostis capillaris, Anthoxanthum odoratum, Dactylis glomerata. Their typical species are
Gymnadenia conopsea, Dactylorhiza majalis, Listera ovata (Neoia ovata), Traunsteinera
globosa, Campanula serrata. At higher alitudes, the main dominant of grasslands is Nardus
stricta, species of high constancy are Agrostis capillaris, Arnica montana, Avenella flexuosa,
Deschampsia cespitosa, Hypochaeris uniflora, Luzula luzuloides, Scorzonera rosea, Sieglingia
decumbens, Vaccinium myrtillus, Vaccinium vitis-idaea. A common vegetaion type is
subalpine Deschampsia caespitosa grasslands (with Arnica montana, Bistorta officinalis,
Calamagrostis arundinacea, Festuca rubra, Geranium alpestre, Hypericum maculatum, Luzula
luzuloides, Phleum alpinum, Poa chaixii, Scorzonera rosea, Viola declinata). Smaller areas are
dominated by Petasites kablikianus (along streams), Festuca saxailis and Festuca carpaica
(on stony calcareous soil). There occur calcareous outcrops with rare species (Aster alpinus,
Saxifraga luteoviridis, Saxifraga aizoides, Silenananthe zawadskii, Ptarmica tenuifolia,
Festuca saxatilis, Silene dubia, Jovibarba preissiana, Campanula kladniana, Cystopteris
alpina), rich fens dominated by Carex paniculata and Carex lava; raised bogs, subalpine
small shrubs communiies dominated by Juniperus sibirica, Vaccinium myrtyllus, Vaccinium
uliginosum; hard water springs and streams with Doronicum carpaicum; acid springs and
streams with Cardamine opizii and Saxifraga stellaris (Micranthes stellaris).
Botanical significance. Rich in endemic species part of the Carpathians. Important area for
mountain spruce forests and subalpine vegetaion.
Criterion A
• Botrychium matricariifolium (A. Braun ex Döll) W.D.J.Koch; A(ii); abundance: rare; trend:
unknown; species data quality: medium; trend data quality: poor.
• Botrychium multifidum (S.G.Gmel.) Rupr.; A(ii); abundance: rare; trend: unknown;
species data quality: medium; trend data quality: poor.
• Campanula abieina Griseb. & Schenk; A(ii); abundance: occasional; trend: stable;
species data quality: medium; trend data quality: poor.
• Campanula serrata (Kit.) Hendrych; A(ii); abundance: occasional; trend: unknown;
species data quality: medium; trend data quality: poor.
• Dicranum viride (Sull. et Lesq.) Lindb.; A(ii); abundance: occasional; trend: unknown;
species data quality: poor; trend data quality: poor.
• Festuca porcii Hack.; A(iv); abundance: rare; trend: unknown; species data quality:
medium; trend data quality: poor.
• Ligularia sibirica (L.) Cass. (L. bucovinensis Nakai); A(ii); rare; trend: unknown; species
66
67
data quality: medium; trend data quality: poor.
Nigritella carpatica (Zapał.) Teppner, Klein et Zagulski (Gymnadenia carpatica (Zapał.)
Teppner et Klein); A(ii); abundance: rare; trend: unknown; species data quality: medium;
trend data quality: poor.
• Tozzia carpathica Woł.; A(ii); abundance: occasional; trend: unknown; species data
quality: medium; trend data quality: poor.
• Typha schuttleworthii W.D.J.Koch et Sond.; A(ii); abundance: rare; trend: unknown;
species data quality: poor; trend data quality: poor.
Criterion C
• C2.12 Hard water springs; area: 0.1 ha; trend: stable; area data quality: poor; trend data
quality: poor.
• D2.226 Peri-Danubian black-white-star sedge fens; area: 15 ha; trend: stable; area data
quality: poor; trend data quality: poor.
• D4.1 Rich fens, including eutrophic tall-herb fens and calcareous lushes and soaks; area:
150 ha; trend: stable; area data quality: poor; trend data quality: poor.
• E1.71 Nardus stricta swards; area: 130 ha; trend: stable; area data quality: poor; trend
data quality: poor.
• E2.3 Mountain hay meadows; area: 100 ha; trend: stable; area data quality: medium;
trend data quality: poor.
• E4.3 Acid alpine and subalpine grassland; area: 350 ha; trend: stable; area data quality:
medium; trend data quality: poor.
• E4.4 Calcareous alpine and subalpine grassland; area: 3 ha; trend: stable; area data
quality: poor; trend data quality: poor.
• E5.5 Subalpine moist or wet tall-herb and fern stands; area: 1000 ha; trend: stable; area
data quality: medium; trend data quality: poor.
• F2.22 Alpide acidocline Rhododendron heaths; area: 1 ha; trend: stable; area data
quality: poor; trend data quality: poor.
• F2.46 (*4070) Carpathian Pinus mugo scrub; area: 670 ha; trend: stable; area data
quality: good; trend data quality: poor.
• F4.2 Dry heaths; area: 10 ha; trend: stable; area data quality: poor; trend data quality:
medium.
• G1.12 Boreo-alpine riparian galleries; area: 100 ha; trend: stable; area data quality:
poor; trend data quality: medium.
• G3.1B Alpine and Carpathian subalpine Picea forests; area: 20000 ha; trend: increasing;
area data quality: medium; trend data quality: medium.
• H2.4 Temperate-montane calcareous and ultra-basic screes; area: 1 ha; trend: stable;
area data quality: poor; trend data quality: medium.
• H3.511 Limestone pavements; area: 1 ha; trend: stable; area data quality: poor; trend
data quality: medium.
Literature
1.Pawłowski В., Walas J. Les associations des plantes vesculaires des Monts de Czywczyn
// Bull. Int. Acad. pol. B. – 1949. – 1. – P. 1–181.
2.Pawłowski В. Ogόlna charakterystyka geobotaniczna Gόr Czywczyńskich // Rozprawy
Wydziału Mat.-Przyrodniczego. – Krakόw, 1948. – P. 1–72.
3.Андрієнко Т.Л., Чорней І.І., Онищенко В.А., Буджак В.В. Флора та рослинність
проектованого міждержавного україно-румунського біосферного резервату
“Мармароські та Чивчино-Гринявські гори” // Укр. ботан. журн. – 2005. – 62, № 4. –
С. 589–596.
4.Величко М.В., Чорней І.І. Ботанічна характеристика урочища Мокринів Камінь у
Чивчинських горах (Українські Карпати) // Заповідна справа в Україні. – 2003. – Т. 9,
вип. 2. – С. 16–18.
5.Величко М.В., Чорней І.І., Буджак В.В. Інвентаризаційний список судинних рослин
Чивчинських гір (Українські Карпати) // Наук. вісник Чернівецького ун-ту: Зб. наук.
праць. Вип. 223: Біологія. – Чернівці: Рута, 2004. – С. 152–161.
6.Горбик В.П. Лучна рослинність Чивчинських та Гринявcьких гір // Укр. ботан. журн.
– 1968. – 25, № 4. – С. 11–17.
7.Горбик В.П. Лісова та чагарникова рослинність Чивчинcьких та Гринявcьких гір //
Укр. ботан. журн. – 1968. – 25, № 6. – С. 87–92.
8.Горбик В.П. Ботанічні об’єкти Чивчинських і Гринявcьких гір, що потребують
охорони // Мат-ли 5 з’їзду Укр. ботан. тов. – Ужгород, 1972. – С. 133–134.
9.Горбик В.П., Андрієнко Т.Л. Болота Чивчин // Укр. ботан. журн. – 1969. – 26, № 3. –
С. 40–44.
10. Загульський М.М., Чорней І.І. Нове місцезнаходження Nigritella nigra (L.) Rich.
(Orchidaceae) в Українських Карпатах // Укр. ботан. журн. – 1993. – 50, № 2. – С. 125–129.
11. Малиновський К.А., Крічфалушій В.В. Рослинні угруповання високогір’я
Українських Карпат – Ужгород, 2002. – 244 с.
12. Червона книга України. Рослинний світ / за ред. Я.П. Дідуха. – К.:
Глобалконсалтинг, 2009. – 900 с.
13. Чопик B.I. Ботаніко-географічна характеристика Чивчино-Гринявських гір в
Українських Карпатах // Укр. ботан. журн. – 1969. – 26, № 6. – С. 26–33.
14. Чорней І.І. До питання про ботанічну цінність Чивчинських гір (Українські
Карпати) // Наук. вісн. Чернівецького ун-ту: Зб. наук. праць. – Вип. 298: Біологія. –
Чернівці: Рута, 2006. – С. 269–273.
15. Чорней І.І., Буджак В.В. Нове місцезнаходження раритетних видів флори у
Чивчинських горах (Українські Карпати) // Укр. ботан. журн. – 2003. – 60, № 1. – С. 53–57.
16. Чорней І.І., Буджак В.В., Андрієнко Т.Л. Болота Буковинських Карпат // Укр.
ботан. журн. – 2008. – 65, № 2. – С. 180–188.
17. Чорней I.I., Загульський М.М., Смолінська М.О., Королюк B.I. Стан та
перспективи охорони рідкісних видів флори у верхів’ї р. Білий Черемош // Екологічні
основи оптимізації режиму охорони і використання природно-заповідного фонду. Тез.
доп. наук.-практ. конф. – Рахів, 1993. – С. 70–72.
18. Чорней I.I., Токарюк А.І., Буджак В.В. НПП Верховинський // Фіторізноманіття
заповідників і національних природних парків України. Ч. 2. Національні природні
парки / під ред. В.А. Онищенка і Т.Л. Андрієнко. – Київ: Фітосоціоцентр, 2012. – С. 74–92.
19. Чорней I.I., Токарюк А.І., Буджак В.В. НПП Черемоський // Фіторізноманіття
заповідників і національних природних парків України. Ч. 2. Національні природні
парки / під ред. В.А. Онищенка і Т.Л. Андрієнко. – Київ: Фітосоціоцентр, 2012. –
С. 530–547.
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•
Davydo-Mykilske
V.A. Onyshchenko, M.M. Peregrym
Ukrainian name: Давидо-Микільське.
70
Area: 1580.0 ha.
Altitude: 25–150 m.
Latitude: 48°30’49” N (48.5136°).
Longitude: 39°51’00” E (39.8499°).
Administrative regions. Luhansk region: Sorokyne (Krasnodon) raion.
Ownership: state.
Biogeographic regions: steppic.
Habitats. Level 1. E – 70.5%; F – 3.0%; G – 26.0%; H – 0.5%.
Habitats. Level 2. E1 Dry grasslands – 70.5%; F3 Temperate and mediterranean-montane
scrub – 3.0%; G1 Broadleaved deciduous woodland – 25.9%; G3 Coniferous woodland –
0.1%; H2 Screes – 0.5%.
Futher habitat description. E1.1 Inland sand and rock with open vegetation; E1.2
Perennial calcareous grassland and basic steppes; F3.1 Temperate thickets and scrub;
F3.2 Submediterranean deciduous thickets and brushes; G1.7 Thermophilous deciduous
woodland; G1.A Meso- and eutrophic Quercus, Carpinus, Fraxinus, Acer, Tilia, Ulmus and
related woodland; G1.C Highly artificial broadleaved deciduous forestry plantations; H2.6
Calcareous and ultra-basic screes of warm exposures.
Land use: agriculture (animals) – major; forestry – minor.
Protected areas: included in Emerald Site “Sukhodilskyi Regional Landscape Park”.
Threats: agricultural intensification/expansion (arable) – low, agricultural intensiicaion/
expansion (grazing) – low, extracion (minerals/quarries) – low, forestry (aforestaion) –
medium.
General description. Slope of the valley of the Siverskyi Donets river with the steppe
vegetaion, forests, and chalk outcrops. Dominants of steppe vegetaion are 4 species
of Sipa (S. pulcherrima, S. lessingiana, S. ucrainica and S. capillata), Festuca valesiaca,
Caragana frutex, Amygdalus nana and Bothriochloa ischaemum. Signiicant components
of the steppe communiies are Salvia nutans, Phlomis pungens, Marrubium praecox,
Psephellus marschallianus (Centaurea marschalliana). Plant communiies on chalk are
formed by Onosma tanaiica, Artemisia salsoloides, Linum czernjajevii, Ephedra distachya,
Euphorbia cretophila, Astragalus albicaulis, Hedysarum grandilorum. Dominants of natural
woods are Quercus robur, Fraxinus excelior, Ulmus suberosa, Euonymus verrucosus, Galium
aparine, Geum urbanum, Chelidonium majus. There are also a lot of ephemeroids: Ficaria
stepporum, Corydalis solida, C. marschalliana, Anemone ranunculoides, Friillaria ruthenica,
Tulipa quercetorum, Scilla siberica, Gagea lutea, Gagea minima.
Botanical significance. Important area for conservaion of the steppe vegetaion and chalk
outcrops.
Criterion C
• E1.2 Perennial calcareous grassland and basic steppes; area: 1120 ha; trend: stable;
area data quality: good; trend data quality: poor.
Conservaion proposals. Do not allow aforestaion. Create a state botanical reserve an
Emerald Site.
Literature:
1. Природно-заповідний фонд Луганської області // О.А. Арапов (заг. ред.), Т.B. Сова,
О.А. Cавенко, В.Б. Фєрєнц, Н.У. Кравець, Л.Л. Зятьков, Л.О. Морозова. Довідник. – 3-є
вид., доп. і перероб. – Луганськ: “Луганська правда”, 2013. – 224 с.
2. Перегрим М.М., Мойсієнко І.І., Перегрим Ю.С., Мельник В.О. Tulipa gesneriana L.
(Liliaceae) в Україні. – К.: Видавничо- поліграфічній центр “Київський університет”, 2009.
– 135 с.
71
Administrative regions. Dnipropetrovsk region: Dnipropetrovsk raion, Petrykivka raion.
Ownership: state.
Biogeographic regions: steppic.
Habitats. Level 1. C – 17.0%; D – 1.0%; E – 39.4%; F – 0.2%; G – 40.8%; H – 0.5%; I – 1.1%.
Habitats. Level 2. C1 Surface standing waters – 8.0%; C2 Surface running waters – 8.0%;
C3 Littoral zone of inland surface waterbodies – 1.0%; D5 Sedge and reedbeds, normally
without free-standing water – 1.0%; E1 Dry grasslands – 16.7%; E2 Mesic grasslands – 5.9%;
E3 Seasonally wet and wet grasslands – 15.8%; E6 Inland salt steppes – 1.0%; F9 Riverine
and fen scrubs – 0.2%; G1 Broadleaved deciduous woodland – 28.9% G3 Coniferous
woodland – 11.9%; H5 Miscellaneous inland habitats with very sparse or no vegetation –
0.5%; I1 Arable land and market gardens – 1.1%.
Futher habitat description. C1.2 Permanent mesotrophic lakes, ponds and pools; C2.3
Permanent non-tidal, smooth-flowing watercourses; C3.2 Water-fringing reedbeds and
tall helophytes other than canes; D5.1 Reedbeds normally without free-standing water;
D5.2 Beds of large sedges normally without free-standing water; D6.1 Inland saltmarshes;
E1.2 Perennial calcareous grassland and basic steppes; E1.9 Open non-Mediterranean dry
acid and neutral grassland including inland dune grassland; E2.5 Meadows of the steppe
zone; E3.4 Moist or wet eutrophic and mesotrophic grassland; E6.2 Coninental inland salt
steppes; F9.2 Salix carr and fen scrub; G1.1 Riparian and gallery woodland, with dominant
Alnus, Betula, Populus or Salix; G1.2 Mixed riparian floodplain and gallery woodland; G1.5
Broadleaved swamp woodland on acid peat; G3.4 Pinus sylvestris woodland south of the
taiga; G3.F Highly artificial coniferous plantations.
Land use: nature conservation and research – 100%; agriculture (arable) – 1%; mowing /
hay making – minor.
Protected areas: same as Dniprovsko-Orilskyi Nature Reserve and Emerald Site “DniprovskoOrilskyi Nature Reserve”.
Threats: abandonment / reduction of land management – low, burning of vegetaion – low,
water (dredging / canalizaion) – low.
General description. This is a part of the loodplain and sand terrace over the loodplain on
the let bank of the Dnipro in the steppe zone. The loodplain has its looding regime close
to natural. Vegetaion of this area is diverse. Forest vegetaion is represented by loodplain
Quercus robur woods (400 ha), Populus alba and Populus nigra woods (400 ha), Salix alba
woods (80 ha), swamped Alnus gluinosa woods (1–2 ha), Pinus sylvestris and Pinus nigra
subsp. pallasiana plantaions (450 ha). Typical species of Quercus robur woods are Ulmus
minor (dominantes in the second sublayer of the tree layer), Acer tataricum, Sambucus
nigra, Crataegus curvisepala, Alliaria peiolata, Galium aparine, Chaerophyllum temulum,
Convallaria majalis, Geum urbanum, Glechoma hederacea, Stellaria media (Alsine media),
Torilis japonica, Urica dioica. Typical species of Populus woods are Amorpha fruicosa (invasive), Frangula alnus, Swida sanguinea (Cornus sanguinea), Rubus caesius, Viburnum opulus. Typical species of Salix alba woods: Populus nigra (frequent co-dominant), Cardamine
amara, Carex riparia, Equisetum arvense, Iris pseudacorus, Stachys sylvaica. Typical species of Alnus gluinosa woods are Carex acuiformis, Thelypteris palustris. Shrub vegetaion
includes communiies of Salix acuifolia (on sands) and Salix cinerea (in waterlogged parts
of loodplain). Dry grasslands (670 ha) include sand steppe and open sand grassland. They
are dominated by Festuca beckeri, Sipa borysthenica, Artemisia marschalliana, Euphorbia
seguieriana, Gypsophila paniculata. Typical species: Allium savranicum, Chamaecyisus borysthenicus, Helichrysum arenarium, Jasione montana, Minuaria piskunovii, Mollugo cerviana, Polygonum arenarium, Salix acuifolia, Jacobaea borysthenica (Senecio borysthenicus),
Seseli tortuosum. Floodplain periodically wet grasslands usually are dominated by Festuca
regeliana. Typical species: Agrosis stolonifera, Alopecurus pratensis, Calamagrosis canescens, Carex otrubae, Elytrigia repens, Galium physocarpum, Hieracium umbellatum, Inula
britannica, Lathyrus pratensis, Poa pratensis, Poa remota, Thalictrum lavum, Vicia tenuifolia, Vicia terasperma. Mesic grasslands are dominated by Poa angusifolia. Saltmarshes are
dominated by Elytrigia elongata, Glaux mariima, Limonium hypanicum, Salicornia perennans, Spergularia media. Eutrophic mires are represented mainly by reedbeds dominated
by Phragmites australis, Typha laifolia, Typha angusifolia and eutrophic sedge mires dominated by Carex riparia, Carex acuta. Aquaic vegetaion is very diverse. Main dominants:
Lemna minor, Lemna trisulca, Nymphaea alba, Nuphar lutea, Salvinia natans, Straiotes
aloides, Spirodela polyrrhiza, Trapa borysthenica (T. natans s. l.), Wolia arrhiza. Smaller
areas are covered by Ceratophyllum demersum, Potamogeton crispus, Potamogeton lucens,
Potamogeton pecinatus (Stuckenia pecinata), Potamogeton perfoliatus.
Botanical significance. This area is one of the most important for conservation of loodplain
complexes in Ukraine.
72
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Dnipro – Oril
V.A. Onyshchenko
Ukrainian name: Дніпро – Оріль.
Area: 3766.2 ha.
Altitude: 50–73 m.
Latitude: 48°30’30” N (48.5083°).
Longitude: 34°47’54” E (34.7983°).
Criterion A
• Agropyron dasyanthum Ledeb.; A(i); abundance: occasional; trend: stable; species data
quality: medium; trend data quality: poor.
• Allium savranicum Besser; A(iv); abundance: occasional; trend: unknown; species data
quality: medium; trend data quality: poor.
• Delphinium rossicum Litv.; A(iv); abundance: rare; trend: unknown; species data quality:
medium; trend data quality: poor.
• Lindernia procumbens (Krock.) Borb.; A(ii); abundance: rare; trend: unknown; species
data quality: medium; trend data quality: poor.
Criterion C
• E1.9 Open non-Mediterranean dry acid and neutral grassland including inland dune
grassland; area: 500 ha; trend: unknown; area data quality: medium; trend data
quality: poor.
• G1.22 Mixed Quercus – Ulmus – Fraxinus woodland of great rivers; area: 400 ha; trend:
stable; area data quality: medium; trend data quality: poor.
• G1.3 Mediterranean riparian woodland; area: 400 ha; trend: unknown; area data
quality: medium; trend data quality: poor.
Literature
1. Манюк В.В. Эколого-флористические особенности дубрав ДнепровскоОрельського природного заповедника // Питання степового лісознавства та лісової
рекультивації земель. – Д.: ДДУ, 1998. – С. 139–146.
2. Манюк В.В. Нарис рослинності Дніпровсько-Орельського природного
заповідника // Заповідна справа в Україні. – Канів, 2000. – Т. 6, вип. 1–2. – С. 7–14.
3. Манюк В.В. Структура, типологія, динаміка і відновлення дібров ДніпровськоОрільського природного заповідника. Дис. … канд. биол. наук: 03.00.16. – Д., 2005. –
373 с.
4. Манюк В.В. ПЗ Дніпровсько-Орільський // Фіторізноманіття заповідників і
національних природних парків України. Ч. 1. Біосферні заповідники. Природні
заповідники / під ред. В.А. Онищенка і Т.Л. Андрієнко. – Київ: Фітосоціоцентр, 2012. –
С. 102–114.
5. Придюк М.П. Мікобіота Дніпровсько-Орельського природного заповідника /
Автореф. дис... канд. біол. наук: 03.00.21. – К., 1999. – 19 с.
Dobrianski Hory
V.A. Onyshchenko
Ukrainian name: Добрянські гори.
Transliteraion/Translaion variants: Dobrianski Hills.
Area: 1048.0 ha.
Altitude: 100–190 m.
Latitude: 48°21’28” N (48.3577°).
Longitude: 39°23’01” E (39.3836°).
Administrative regions. Luhansk region: Lutuhyne raion.
Ownership: state.
Biogeographic regions: steppic.
Habitats. Level 1. C – 0.1%; E – 93.3%; F – 2.0%; G – 4.2%; H – 0.3%; J – 0.1%.
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Habitats. Level 2. C1 Surface standing waters – 0.1%; E1 Dry grasslands – 92.0%; E2
Mesic grasslands – 1.3%; F3 Temperate and mediterranean-montane scrub – 2.0%; G1
Broadleaved deciduous woodland – 4.2%; H5 Miscellaneous inland habitats with very
sparse or no vegetation – 0.3%; J4 Transport networks and other constructed hard-surfaced
areas – 0.1%.
Futher habitat description. E1.2 Perennial calcareous grassland and basic steppes; F3.2
Submediterranean deciduous thickets and brushes; G1.7 Thermophilous deciduous
woodland; H5.6 Trampled areas.
Land use: agriculture (animals) – major, forestry – minor.
Protected areas: includes proposed botanical reserve “Dobrianski Hory” (about 100 ha),
overlaps with Emerald Site “Dobrianski Hory” (about 108 ha).
Threats: agricultural intensification/expansion (arable) – low, agricultural intensiicaion/
expansion (grazing) – low, extracion (minerals/quarries) – low, forestry (aforestaion) –
low.
General description. A slope of the Luhanchyk river valley. The major habitat type is the
steppe. It is dominated by Festuca valesiaca s.l., Sipa capillata, Sipa dasyphylla, Sipa
pennata, Caragana frutex. Forests are dominated by Quercus robur, Fraxinus excelior,
Ulmus suberosa, Euonymus verrucosus, Galium aparine, Geum urbanum, Ficaria stepporum,
Corydalis solida. Prevailng species of the shrub vegetaion are Cerasus fruicosa, Cerasus
stepposa, Rosa sp. There are outcrops of sandstone, slate, and coal.
Botanical significance. Important area for conservaion of the steppe vegetaion.
Criterion C
• E1.2 Perennial calcareous grassland and basic steppes; area: 930 ha; trend: stable; area
data quality: good; trend data quality: medium.
Conservaion proposals. Create a state botanical reserve including the enire IPA.
Literature
1. Перегрим М.М. Флора та рослинність проектованого заповідного урочища
“Добрянські гори” (Луганська область) // Теоретичні та прикладні аспекти інтродукції
рослин і зеленого будівництва: Матеріали ІІ Міжнародної наукової конференції
молодих дослідників. – Київ: Фітосоціоцентр, 2002 – С. 65–66.
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Dolyna Inhulu
V.A. Onyshchenko
Ukrainian name: Долина Інгулу.
Transliteraion/Translaion variants: Ingul Valley, Ingul River Valley.
Area: 1243.0 ha.
Altitude: 39–81 m.
Latitude: 47°46’24” N (47.7734°).
Longitude: 32°22’42” E (32.3782°).
Administrative regions. Kirovohrad region: Ustynivka raion; Mykolaiv region: Novyi Buh
raion.
Ownership: state.
Biogeographic regions: steppic.
Habitats. Level 1. C – 19.6%; D – 0.2%; E – 74.2%; F – 1.0%; G – 4.0%; H – 1.0% .
Habitats. Level 2. C1 Surface standing waters – 19.0%; C2 Surface running waters – 0.5%;
C3 Littoral zone of inland surface waterbodies – 0.1%; D5 Sedge and reedbeds, normally
without free-standing water – 0.2%; E1 Dry grasslands – 71.0%; E2 Mesic grasslands – 3.0%;
E3 Seasonally wet and wet grasslands – 0.2%; F3 Temperate and mediterranean-montane
scrub – 1.0%; G1 Broadleaved deciduous woodland – 4.0%; H2 Screes – 0.0; H3 Inland cliffs,
rock pavements and outcrops – 0.4%; H5 Miscellaneous inland habitats with very sparse
or no vegetation – 0.6%.
Futher habitat description. C1.2 Permanent mesotrophic lakes, ponds and pools; C1.3
Permanent eutrophic lakes, ponds and pools; C2.2 Permanent non-idal, fast, turbulent
watercourses; C2.3 Permanent non-idal, smooth-lowing watercourses; E1.1 Inland sand
and rock with open vegetation; E1.2 Perennial calcareous grassland and basic steppes; F3.1
Temperate thickets and scrub; F3.1 Temperate thickets and scrub; F3.2 Submediterranean
deciduous thickets and brushes; G1.C Highly ariicial broadleaved deciduous forestry
plantaions; H3.1 Acid silicious inland clifs; H5.6 Trampled areas.
Land use: agriculture (animals) – major; forestry – minor; nature conservaion and research
– major; water management – minor.
Protected areas: overlaps with Pryinhulskyi regional landscape park (1031 ha) and Emerald
Site “Pryinhulskyi Regional Landscape Park “ (1031 ha), overlaps with regional hydrological
reserve “Soiivske Vodoskhovyshche” (174 ha); includes state zoological reserve “Polozova
Balka” (27 ha), regional botanical reserve “Pelaheivskyi” (123.5 ha).
Threats: agricultural intensiicaion/expansion (grazing) – low; forestry (aforestaion) – low.
General description. The Inhul valley with steppe vegetaion and silicate outcrops. The
steppe vegetaion is dominated by Botriochloa ischaemum, Bromopsis inermis, Caragana
frutex, Elytrigia intermedia, Elytrigia repens, Carex praecox, Festuca valesiaca, Galatella
villosa, Poa angusifolia, Poa bulbosa (in areas with intensive grazing), Spiraea hypericifolia,
Sipa capillata, Sipa lessingiana, Sipa pennata, someimes Sipa dasyphylla, Sipa irsa. The
most frequent species are Eryngium campestre, Euphorbia seguierana, Festuca valesiaca,
Koeleria cristata, Medicago falcata, Potenilla impolita, Teucrium chamaedrys. Petrophyic
steppes difer with presence of Achillea ochroleuca, Ceratodon purpureus, Eremogone
biebersteinii, Erophila verna, Minuaria setacea, Potenilla incana, Pulsailla pratensis,
Sedum acre, Sedum pallasii, Tortula ruralis, Sipa graniicola, meadow steppes – with
Caragana frutex, Filipendula vulgaris, Fragaria viridis, Galium verum, Spiraea hypericifolia,
Thalictrum minus, true steppes - with Herniaria besseri, Marrubium praecox, Phlomis
76
pungens, Sipa lessingiana, Teucrium polium. Diferenial species of overgrazed steppes are
Anthemis ruthenica, Chenopodium album, Grindelia squarrosa (invasive), Poa bulbosa, Salvia
aethiopis, Sisymbrium alissimum. In mesic grasslands, there prevail Calamagrosis epigeios,
Elytrigia repens, Poa angusifolia, Poa pratensis. The shrub vegetaion is represented mainly
by communiies of Acer tataricum and Prunus spinosa s.l.
Botanical significance. Important for conservaion of steppe vegetaion, silicious outcrops,
endemic petrophyic species.
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Criterion A
• Dianthus hypanicus Andrz.; A(i), A(ii), A(iii), abundance: rare; trend: unknown; species
data quality: medium; trend data quality: poor.
• Tulipa hypanica Klokov et Zoz; A(iii); abundance: occasional; trend: unknown; species
data quality: medium; trend data quality: poor.
Criterion C
• E1.11 Euro-siberian pioneer rock debris swards; area: 2 ha; trend: unknown; area data
quality: poor; trend data quality: poor.
• E1.2 Perennial calcareous grassland and basic steppes; area: 900 ha; trend: stable; area
data quality: medium; trend data quality: medium.
• F3.247 Ponto-Sarmatic deciduous thickets; area: 3 ha; trend: stable; area data quality:
medium; trend data quality: medium.
• H3.1 Acid siliceous inland cliffs; area: 4 ha; trend: stable; area data quality: medium;
trend data quality: good.
Literature
1. Винокуров Д.С. Синтаксономія ксеротермної рослинності р. Інгул (клас FestucoBroetea). Частина І. Петрофітно-степова рослинність. до флори Правобережного степу
України // Укр. бот. журн. – 2014. – 71, № 2. – С. 148–160.
2. Винокуров Д.С. Синтаксономія ксеротермної рослинності долини р. Інгул (клас
Festuco-Brometea). Частина 2. Лучно-степова, чагарниково-степова, справжньостепова
рослинність // Укр. бот. журн. – 2014. – 71, № 5. – С. 538–549.
3. Винокуров Д. Созофіти долини р. Інгул і завдання їх охорони // Вісник Львівського
університету. Сер. Біологічна. – 2014. – 65. – С. 135–150.
4. Винокуров Д.С. Синтаксономія вищої водної рослинності долини р. Інгул //
Чорноморський бот. журн. – 2011. – 7, № 1. – С. 26–40.
5. Винокуров Д.С. Ключові території Інгульського регіонального екокоридору:
характеристика, зв'язки, оптимізація // Чорноморський бот. журн. – 2011. – 7, № 4. –
С. 329–346.
6. Екомережа степової зони України: принципи створення, структура, елементи /
під ред. Д.В. Дубини і Я.І. Мовчана. – К., 2013. – 409 с.
without free-standing water – 0.6%; E1 Dry grasslands – 61.2%; E2 Mesic grasslands –
0.3%; F3 Temperate and mediterranean-montane scrub – 2.0%; G1 Broadleaved deciduous
woodland – 30.6%; H2 Screes – 0.1%; H3 Inland cliffs, rock pavements and outcrops – 2.9%;
H5 Miscellaneous inland habitats with very sparse or no vegetation – 0.2% .
Futher habitat description. C1.2 Permanent mesotrophic lakes, ponds and pools; C1.3
Permanent eutrophic lakes, ponds and pools; C2.2 Permanent non-idal, fast, turbulent
watercourses; C2.3 Permanent non-idal, smooth-lowing watercourses; E1.1 Inland sand
and rock with open vegetation; E1.2 Perennial calcareous grassland and basic steppes; F3.1
Temperate thickets and scrub; F3.2 Submediterranean deciduous thickets and brushes;
G1.7 Thermophilous deciduous woodland; G1.C Highly ariicial broadleaved deciduous
forestry plantaions; H3.1 Acid silicious inland clifs.
Land use: agriculture (animals) – major; forestry – 20%; nature conservaion and research
Dolyna Mertvovodu
V.A. Onyshchenko
Ukrainian name: Долина Мертвоводу.
Transliteraion/Translaion variants: Mertvovod Valley, Mertvovod River Valley.
Area: 1571.0 ha.
Altitude: 23–100 m.
Latitude: 47°42’47” N (47.7130°).
Longitude: 31°27’21” E (31.4557°).
Administrative regions. Mykolaiv region: Arbuzynka raion, Bratske raion, Voznesensk raion.
Ownership: state, unknown.
Biogeographic regions: steppic.
Habitats. Level 1. C – 2.1%; D – 0.6%; E – 61.5%; F – 2.0%; G – 30.6%; H – 3.2% , J – 0.0%.
Habitats. Level 2. C1 Surface standing waters – 0.9%; C2 Surface running waters – 0.4%;
C3 Littoral zone of inland surface waterbodies – 0.8%; D5 Sedge and reedbeds, normally
78
– 100%.
Protected areas: included in Buzkyi Hard Naional Nature Park and Emerald Site “Bugzkyi
Gard Naional Nature Park”.
Threats: development (recreaion/tourism) – low; forestry (aforestaion) – medium;
habitat fragmentaion/isolaion – low.
General description. The Mertvovod valley with high (10–60 m) steep slopes and ganite
clifs. Large areas are covered by the steppe vegetaion, natural Quercus robur termophilous
woods, ariicial Quercus robur, Robinia pseudoacacia woods, steppic and loodplain scrub.
Natural deciduous termophilous woods. Dominants: Quercus robur (major), Acer campestre,
Fraxinus excelsior, Tilia cordata, Ulmus minor; Acer tataricum, Coinus coggygria, Crataegus
79
fallacina, Euonymus europaeus, Eunymus verrucosus, Swida sanguinea (Cornus sanguinea).
Typical species: Malus sylvestris, Pyrus communis; Chelidonium major, Galium aparine,
Geum urbanum, Urica dioica.
Shrubs. Dominant: Crataegus praearmata, Spiraea crenata, Prunus stepposa.
Dry grasslands (steppes). Dominants: Sipa capillata, Sipa lessingiana, Festuca valesiaca,
Botriochloa ischaemum
Silicious outcrops and screes. Typical species: Achillea ochroleuca, Alyssum murale, Aurinia
saxailis, Rumex fascilobus, Sedum acre, Sedum borissovae, Sempervivum ruthenicum,
Thymus dimorphus.
Botanical significance. Important for conservaion of silicious outcrops with endemic
species (Dianthus hypanicus Andrz., Moehringia hypanica Grynj et Klokov, Sedum borissovae
Balk., Silene sytnikii Krytzka, Novosad et Protopopova, Tulipa hypanica Klokov et Zoz),
termophilous oak woods and steppic scrub.
Criterion A
• Delphinium sergii Wissjul.; A(iv); abundance: rare; trend: unknown; species data quality:
medium; trend data quality: poor.
• Dianthus hypanicus Andrz.; A(i), A(ii), A(iii); abundance: occasional; trend: stable;
species data quality: medium; trend data quality: poor.
• Moehringia hypanica Grynj et Klokov; A(i); abundance: rare (one locality); trend:
decreasing; species data quality: good; trend data quality: medium.
• Silene sytnikii Krytzka, Novosad et Protopopova; A(iii); abundance: occasional; trend:
stable; species data quality: good; trend data quality: medium.
Criterion C
• E1.11 Euro-siberian pioneer rock debris swards; area: 1 ha; trend: unknown; area data
quality: poor; trend data quality: poor.
• E1.2 Perennial calcareous grassland and basic steppes; area: 970 ha; trend: stable; area
data quality: medium; trend data quality: medium.
• F3.247 Ponto-Sarmatic deciduous thickets; area: 15 ha; trend: stable; area data quality:
poor; trend data quality: poor.
• H3.1 Acid siliceous inland cliffs; area: 45 ha; trend: stable; area data quality: medium;
trend data quality: good.
Literature
1. Гревцова Г.Т. Кизильники гранітно-степового Побужжя // Збірник наукових праць
Полтавського пед. ун-ту. – 2003. – Вип.4 (31). – С. 54–61.
2. Драбинюк Г.В. НПП Бузький Гард // Фіторізноманіття заповідників і національних
природних парків України. Ч. 2. Національні природні парки / під ред. В.А. Онищенка і
Т.Л. Андрієнко. – Київ: Фітосоціоцентр, 2012. – С. 49–60.
3. Новосад В.В., Крицкая Л.И., Протопопова В.В. Новий для науки ендемічний
вид Гранітно-степового Побужжя смілка Ситника (Silene sytnikii Krytzka, Novosad
et Protopopova), його таксономічні, еколого-ценотичні, хорологічні, генезисні та
созологічні особливості // Укр. бот. журн. – 1996. – 53, № 5. – С. 578–585.
4. Партика Л.Я., Вірченко В.М., Нипорко С.О. До бріофлори регіонального
ландшафтного парку “Гранітно-степове Побужжя” // Чорномор. ботан. журн. – 2006. –
2, № 1. – С. 116–122.
5. Соломаха В.А., Драбинюк Г.В., Вініченко Т.С., Мойсієнко І.І., Деркач О.М. Адаптивні
особливості південнобузьких ендемів Dianthus hypanicus Andrz. та Moehringia hypanica
Grynj et Klok. // Укр. фітоцен. зб. – Сер. С. – 2006. – Вип. 24. – С. 70–86.
6. Соломаха В.А., Соломаха Т.Д., Драбинюк Г.В., Мойсієнко І.І. Знахідка Asplenium
V.A. Onyshchenko
Ukrainian name: Долина Нарцисів.
Transliteraion/Translaion variants: Narcissus Valley.
Area: 257.6 ha.
Altitude: 169–180 m.
Latitude: 48°10’57” N (48.1823°).
Longitude: 23°21’28” E (23.3575°).
Administrative regions. Zakarpatska region: Khust raion.
Ownership: state.
Biogeographic regions: pannonian.
Habitats. Level 1. C – 0.4%; D – 12.0%; E – 79.3%; F – 6.0%; G – 2.0%; J – 0.3%.
Habitats. Level 2. C2 Surface running waters – 0.4%; C3.2 Water-fringing reedbeds
and tall helophytes other than canes; D2 Valley mires, poor fens and transition mires
– 9.0%; D5 Sedge and reedbeds, normally without free-standing water – 3.0%; E1 Dry
grasslands – 4.0%; E2 Mesic grasslands – 27.0%; E3 Seasonally wet and wet grasslands –
48.0%; E5 Woodland fringes and clearings and tall forb stands – 0.3%; F3 Temperate and
mediterranean-montane scrub – 0.2%; F9 Riverine and fen scrubs – 5.8%; G1 Broadleaved
deciduous woodland – 2.0%; J4 Transport networks and other constructed hard-surfaced
areas – 0.3%.
Futher habitat description. C2.3 Permanent non-tidal, smooth-flowing watercourses;
D5.1 Reedbeds normally without free-standing water; E2.2 Low and medium alitude hay
meadows; E3.4 Moist or wet eutrophic and mesotrophic grassland; E3.5 Moist or wet
oligotrophic grassland; E5.4 Moist or wet tall-herb and fern fringes and meadows; F3.1
Temperate thickets and scrub; F9.1 Riverine scrub; F9.2 Salix carr and fen scrub; G1.1
Riparian and gallery woodland, with dominant Alnus, Betula, Populus or Salix; G1.2 Mixed
riparian floodplain and gallery woodland; J4.2 Road networks.
Land use: mowing/hay making – major, nature conservation and research – major, tourism/
recreaion – minor.
Protected areas: included in Karpatskyi (Carpathian) biosphere reserve (naional category),
included in Carpathian biosphere reserve (UNESCO), included in Emerald Site “Carpathian
Biosphere Reserve “.
Threats: –.
General description. Floodplain of the river Khustets. The main vegetaion type is
80
81
x alternifolium Wulfen у степовій зоні України // Укр. бот. журн. – 2006. – 63, № 9. –
C. 515–517.
7. Червона книга України. Рослинний світ / за ред. Я.П. Дідуха. – К.: Глобалконсалтинг,
2009. – 900 с.
8. Щербакова О.Ф., Новосад В.В., Крицька Л.І. Раритетний флорофонд КодимоЄланецького Побужжя (ЧКУ, 2009): популяційні та флоросозологічні аспекти. Рослинний
світ у Червоній книзі України: впровадження Глобальної стратегії збереження рослин.
Мат-ли міжнар. конф. (11–15 жовтня 2010 р., м.Київ). – Київ: Альтерпрес, 2010. – С.
210–214.
Dolyna Nartsysiv
meadows dominated by Molinia caerulea, Deschampsia cespitosa, Narcissus angusifolius,
Anthoxanthum odotatum, Festuca ovina, Festuca rubra, Agrostis capillaris, with presence of
Betonica officinalis, Centaurea jacea, Filipendula vulgaris, Poa pratensis, Potentilla erecta,
Ranunculus acris, Ranunculus repens, Sanguisorba officinalis, Leucanthemum vulgare. Besides
there are communniies of Phragmites australis, Iris pseudacorus, Juncus efusus. Along
watercourses, there prevail Salix alba, Salix capraea, Salix cinerea, Salix viminalis.
Botanical significance. The area includes the largest populaion of Narcissus angusifolius in
Ukraine. Narcissus angusifolius dominates in the area of 30 ha.
Criterion A
Мовчан, Ф.Д. Гамор та ін. – К.: Інтерекоцентр, 1997. – 711 с.
2.Стойко С.М., Тасєнкевич Л.О., Мілкіна Л.І. та ін. Флора і рослинність Карпатського
заповідника. – Київ : Наукова думка, 1982 . – 219 с.
3.Устименко П.М., Дубина Д.В., Гамор Ф.Д. Рослинність заповідного масиву “Долина
Нарцисів”: сучасний стан та динамічні тенденції // Укр. бот. журн. – 2007. – 64, № 2. –
C. 195–205.
Dubovetski Skhyly
Ukrainian name: Дубовецькі схили.
Transliteraion/Translaion variants: Dubovetski Slopes.
Area: 4.4 ha.
Alitude: 265–315 m.
Laitude: 49°05’07” N (49.0854°).
Longitude: 24°48’37” E (24.8103°).
Subnaional regions. Ivano-Frankivsk region: Halych raion.
Ownership: state.
•
Narcissus angustifolius Curtis; A(ii); abundance: abundant; trend: stable; species data
quality: good; trend data quality: medium.
Literature
1.Біорізноманіття Карпатського біосферного заповідника / Кол. авт., Ред. рада: Я.І.
82
I.M. Danylyk
Biogeographic regions: coninental.
Habitats. Level 1. E – 86.0%; F – 10.0%; G – 4.0%.
Habitats. Level 2. E1 Dry grasslands –81.0%; E7 Sparsely wooded grasslands – 5.0%; F3
Temperate and mediterranean-montane scrub – 10.0%; G1 Broadleaved deciduous
woodland – 4.0%.
Futher habitat descripion. E1.2 Perennial calcareous grassland and basic steppes; F3.1
83
Temperate thickets and scrub.
Land use: mowing/hay making (major), forestry – minor.
Protected areas: same as proposed state botanical reserve “Ofrys”
Threats: abandonment/reducion of land management – high, forestry (aforestaion) – low.
General descripion. Semi-dry grasslands, natural deciduous woods and shrubs (Frangula
alnus, Prunus spinosa, Crataegus spp.). Grasslands are dominated by Inula ensifolia, Carex
humilis, Peucedanum cervaria, Brachypodium pinnatum. Other frequent species: Geranium
sanguineum, Salvia pratensis, Filipendula vulgaris, Carex montana, Stachis recta, Trifolium
medium, Cirsium pannonicum, Eryngium planum, Prunella grandilora, Thalictrum minus,
Agrimonia eupatoria, Asperula cynanchica, Campanula sibirica, Teucrium chamaedrys.
Botanical signiicance. The only populaion of Ophrys apifera in Ukraine outside the Crimea.
Criterion C
• E1.2 Perennial calcareous grassland and basic steppes (6210, *Important orchid site);
area: 3.5 ha; trend: decreasing; area data quality: good; trend data quality: good. Size
of populaion of Ophrys apifera: 30–100 individuals.
Conservaion proposals. Do not allow expansion of trees and shrubs, mow grasslands,
non-intensive grazing. Create a state botanical reserve. Include in Halytskyi naional nature
park and Emerald Site “Halytskyi Naional Nature Park”.
Literature
1. Данилик І.М., Борсукевич Л.М. Нове місцезнаходження Ophrys apifera Huds.
(Orchidaceae) в Україні // Укр. ботан. журн. – 2011. – 68, № 1. – С. 58-64.
2. Червона книга України. Рослинний світ / за ред. Я.П. Дідуха. – К.: Глобалконсалтинг,
2009. – 912 с.
Protected areas: includes regional preserve (zapovidne urochyshche) “Dubrovy” (10.0 ha),
includes regional preserve (zapovidne urochyshche) “Mochar” (8.0 ha), includes proposed
regional botanical reserve “Bushtynskyi Park Liodovykovoho Periodu”.
Threats: water (drainage) – medium.
General description. A complex of deciduous forests, swamps and wet meadows. Woods
Dubrova
are represented by moist acidophilous Quercus robur forests (with Agrosis canina, Betonica
oicinalis, Betula pubescens, Carex brizoides (dom.), Crataegus monogyna s. l., Deschampsia
cespitosa, Euonymus europaeus, Frangula alnus, Molinia caerulea (dom.), Malus sylvestris,
Populus tremula, Swida sanguinea (Cornus sanguinea) and hygrophylous species that
are typical of alder swamps), mesic species rich Quercus robur forests (with Anemone
nemorosa, Anemone ranunculoides, Carex curvata, Clemais recta, Corylus avellana,
Frangula alnus, Fraxinus excelsior, Lathyrus niger, Malus sylvestris, Melica unilora, Populus
nigra, Potenilla alba, Prunus spinosa, Polygonatum mulilorum, Pulmonaria oicinalis,
Ranunculus polyanthemos, Serratula inctoria, Vicia cassubica, Vinca minor and less oten
Crocus banaicus, Pilosella auraniaca (Hieracium auraniacum), Muscari transsilvanicum,
Phyteuma vagneri, Rosa gallica, Veratrum album), wet Alnus gluinosa woods (with Carex
elongata, Carex riparia, Carex vesicaria, Galium palustre, Glyceria maxima, Iris pseudacorus,
Lycopus europaeus, Lythrum salicaria, Rubus caesius, Salix cinerea, Solanum dulcamara,
Stachys palustris). Former fens and bogs are replaced by Molinia caerulea abandoned
pastures (predominantly) and Alopecurus pratensis hay meadows.
Botanical significance. The area is important for conservaion of termophilous oak forests
and hygrophilous deciduous forests.
Criterion C
• E3.5 Moist or wet oligotrophic grassland; area: 350 ha; trend: stable; area data quality:
medium; trend data quality: medium.
• G1.21 Riverine Fraxinus – Alnus woodland (*91E0), wet at high but not at low water;
area: 100 ha; trend: stable; area data quality: medium; trend data quality: medium.
R.Ya. Kish, V.A. Onyshchenko
Ukrainian name: Дуброва.
Area: 762.0 ha.
Altitude: 202–219 m.
Latitude: 48°05’13” N (48.0869°).
Longitude: 23°28’48” E (23.4800°).
Administrative regions. Zakarpatska region: Tiachiv raion.
Ownership: state, private (minor).
Biogeographic regions: pannonian.
Habitats. Level 1. C – 0.1%; D – 1.0%; E – 51.3%; G – 47.1%; H – 0.5%.
Habitats. Level 2. C2 Surface running waters – 0.1%, D2 – Valley mires, poor fens and
transition mires – 0.0%; D5 Sedge and reedbeds, normally without free-standing water –
1.0%; E3 Seasonally wet and wet grasslands – 51.3%; G1 Broadleaved deciduous woodland
– 47.1%; H5 Miscellaneous inland habitats with very sparse or no vegetation – 0.5%.
Futher habitat description. D2.3 Transition mires and quaking bogs; D5.2 Beds of large
sedges normally without free-standing water; G1.2 Mixed riparian floodplain and gallery
woodland; G1.4 Broadleaved swamp woodland not on acid peat; G1.7 Thermophilous
deciduous woodland; G1.8 Acidophilous Quercus-dominated woodland; H5.6 Trampled
areas.
Land use: forestry – minor, agriculture (animals) – minor; mowing/hay making – minor.
84
85
•
G1.7 Thermophilous deciduous woodland (*91I0); area: 50 ha; trend: stable; area data
quality: medium; trend data quality: medium.
Conservaion proposals. Raise the groundwater table. Create a botanical reserve and an
Emerald Site.
Lemna trisulca, Myriophyllum vericillatum, Myriophllum spicatum, Potamogeton pecinatus
(Stuckenia pecinata), Salvinia natans, Spirodela polyrrhiza, Trapa natans. In brackish water
Dunaiski Plavni
V.A. Onyshchenko
Ukrainian name: Дунайські плавні.
Transliteraion/Translaion variants: Danube Wetlands.
Area: 43601.0 ha.
Altitude: 0–13 m.
Latitude: 45°23’40” N (45.3945°).
Longitude: 29°41’00” E (29.6833°).
Administrative regions. Odesa region: Kiliya raion; Territorial waters of Ukraine .
Ownership: state.
Biogeographic regions: steppic.
Habitats. Level 1. A – 11.2%; C – 10.7%; D – 71.1%; E – 5.0%; F – 0.4%; G – 1.6%.
Habitats. Level 2. A2 Littoral sediment – 0.1%; A5 Sublittoral sediment – 11.1%; C1 Surface
standing waters – 3.6%; C2 Surface running waters – 2.3%; C3 Littoral zone of inland
surface waterbodies – 4.8%; D5 Sedge and reedbeds, normally without free-standing water
– 70.9%; D6 Inland saline and brackish marshes and reedbeds – 0.2%; E1 Dry grasslands
– 0.2%; E3 Seasonally wet and wet grasslands – 1.5%; E6 Inland salt steppes – 3.3%; F9
Riverine and fen scrubs – 0.4%; G1 Broadleaved deciduous woodland – 0.4% G3 Coniferous
woodland – 1.2%.
Futher habitat description. C1.2 Permanent mesotrophic lakes, ponds and pools; C2.3
Permanent non-tidal, smooth-flowing watercourses; C3.2 Water-fringing reedbeds and tall
helophytes other than canes; D5.1 Reedbeds normally without free-standing water; D5.2
Beds of large sedges normally without free-standing water; D6.1 Inland saltmarshes; E1.9
Open non-Mediterranean dry acid and neutral grassland including inland dune grassland;
E3.4 Moist or wet eutrophic and mesotrophic grassland; E6.2 Coninental inland salt
steppes; F9.1 Riverine scrub; G1.1 Riparian and gallery woodland, with dominant Alnus,
Betula, Populus or Salix; G1.C Highly artificial broadleaved deciduous forestry plantations;
G3.F Highly artificial coniferous plantations.
Land use: isheries/aquaculture – minor; forestry – 2%; nature conservation and research –
76%; mowing/hay making – minor; urban/industrial/transport – 1%.
Protected areas: overlaps (about 35747 ha) with Danube Delta transboundary biosphere
reserve and Emerald Site “Danube Biosphere Reserve” (about 35747 ha).
Threats: forestry (aforestaion) – medium.
General description. The Ukrainian part of the Danube delta. It includes branches of the
river, large reed marsches, lakes, meadows, halophyic vegetaion, 1 km strip of the Black
Sea, a sand ridge with pine plantaions and dry sand grasslands. The dominant vegetaion
type is freshwater marshes. The main dominant is Phragmites australis (a frequent codominant is Schoenoplectus lacustris), other important dominants are Typha angusifolia,
Carex acuiformis, Carex elata, Carex pseudocyperus. Freshwater aquaic vegetaion is
usually dominanted by Ceratophyllum demersum, Hydrocharis morsus-ranae, Lemna minor,
86
in the sea and lagoons, Zostera marina, Zostera nolii (Nanozostera nolii), Ruppia mariima
predominate. Halophyic vegetaion is represented by communiies of annuals Salicornia
perennans, Suaeda prostrara, Halimione pedunculata and perennials Aeluropus litoralis,
Bolboschoenus mariimus, Carex distans, Carex extensa, Juncus gerardii, Juncus mariimus,
Puccinellia gigantea. Broadleaved forests are represented mainly by riverine Salix alba
woods, both natural and ariicial. On the largest area of dry sands (about 640 ha), there are
plantaions of Pinus pallasiana and grasslands dominated by Calamagrosis epigeios and
Carex colchica. On the sea beach sand ridges, dominants are Artemisia arenaria, Leymus
sabulosus, Secale sylvestre. Psammophyic communiies contain a narrow endemic species,
Dianthus bessarabicus.
87
Botanical significance. This area is richest in aquaic vascular plant species in Ukraine.
Criterion A
• Aldrovanda vesiculosa L.; A(i), A(ii); abundance: occasional; trend: unknown; species
data quality: medium; trend data quality: poor.
• Carex secalina Wahlenb.; A(ii); abundance: occasional; trend: stable; species data
quality: medium; trend data quality: poor.
• Dianthus bessarabicus Klokov; A(iii); abundance: rare; trend: stable; species data quality:
medium; trend data quality: medium.
• Salvinia natans (L.) All.; A(ii); abundance: frequent; trend: stable; species data quality:
good; trend data quality: medium.
• Trapa natans L.; A(ii); abundance: occasional; trend: stable; species data quality:
medium; trend data quality: medium.
• Typha minima Funk; A(ii); abundance: occasional; trend: unknown; species data quality:
medium; trend data quality: poor.
Criterion C
• C1.224 Floaing Utricularia australis and Utricularia vulgaris colonies; area: 10 ha; trend:
stable; area data quality: poor; trend data quality: medium.
• C1.225 Floaing Salvinia natans mats; area: 10 ha; trend: stable; area data quality: poor;
trend data quality: medium.
• C1.226 Floating Aldrovanda vesiculosa communities; area: 0.1 ha; trend: unknown; area
data quality: poor; trend data quality: poor.
• C1.3411 Ranunculus communities in shallow water; area: 1 ha; trend: unknown; area
data quality: poor; trend data quality: poor.
• C2.34 Eutrophic vegetation of slow-flowing rivers; area: 50 ha; trend: stable; area data
quality: poor; trend data quality: medium.
• D5.2 Beds of large sedges normally without free-standing water; area: 400 ha; trend:
unknown; area data quality: good; trend data quality: poor.
• E6.2 Continental inland salt steppes; area: 1400 ha; trend: stable; area data quality:
poor; trend data quality: poor.
Conservaion proposals. Restore sand grasslands on the areas occupied by Pinus
plantaions.
Literature
1. Біорізноманітність Дунайського біосферного заповідника, збереження та
управління / Гол. ред. Ю.Р. Шеляг-Сосонко. – К.: Інтерекоцентр, 1999. – 702 с.
2. Дворецький Т.В. Вплив викошування на рослинність засолених луків Дунайського
біосферного заповідника (ДБЗ) // Укр. фітоцен. зб. – К., 1999. – Сер. С, вип. 1 (15). –
С. 68–78.
3. Дубина Д.В., Жмуд О.І. БЗ Дунайський // Фіторізноманіття заповідників і
національних природних парків України. Ч. 1. Біосферні заповідники. Природні
заповідники / під ред. В.А. Онищенка і Т.Л. Андрієнко. – Київ: Фітосоціоцентр, 2012. –
С. 31–44.
4. Дубына Д.В., Шеляг-Сосонко Ю.Р. Плавни Причерноморья. – К.: Наук. думка,
1989. – 272 с.
5. Дубина Д.В., Шеляг-Сосонко Ю.Р., Жмуд О.І., Жмуд М.Є., Дворецький Т.В.,
Дзюба Т.П., Тимошенко П.А. Дунайський біосферний заповідник. Рослинний світ. – К.:
Фітосоціоцентр, 2003. – 459 с.
6. Зеров К.К. Водная растительность Килийской дельты Дуная // Дунай и
придунайские водоемы в пределах СССР. – Тр. Ин-та гидробиологии АН УССР. – 36. –
88
С. 37–48.
7. Червона книга України. Рослинний світ / за ред. Я.П. Дідуха. – К.: Глобалконсалтинг,
2009. – 900 с.
8. Шеляг-Сосонко Ю.Р., Дубына Д.В. Государственный заповедник “Дунайские
плавни”. – К.: Наук. думка, 1984. – 286 с.
Dzharylhach
V.A. Onyshchenko, V.P. Kolomiychuk
Ukrainian name: Джарилгач.
Area: 10555.0 ha.
Altitude: 0–3 m.
Latitude: 46°02’09” N (46.0358°).
Longitude: 32°52’48” E (32.8800°).
Administrative regions. Kherson region: Skadovsk raion; Territorial waters of Ukraine.
Ownership: state.
Biogeographic regions: steppic.
Habitats. Level 1. A – 63.0%; B – 0.8%; C – 1.0%; E – 34.5%; G – 0.4%; H – 0.3%.
Habitats. Level 2. A2 Littoral sediment – 20.0%; A5 Sublittoral sediment – 43.0%; B1 Coastal
dunes and sandy shores – 0.6%; B2 Coastal shingle – 0.2%; C1 Surface standing waters
– 1.0%; E1 Dry grasslands – 31.5%; E2 Mesic grasslands – 1.0%; E6 – Inland salt steppes –
2.0%; G1 Broadleaved deciduous woodland – 0.4%; H5 Miscellaneous inland habitats with
very sparse or no vegetation – 0.3%.
Futher habitat description. A2.5 Coastal saltmarshes and saline reedbeds; A5.2 Sublitoral
sand; A5.4 Sublitoral mixed sediments; A5.5 Sublitoral macrophyte-dominated sediment;
B1.1 Sand beach driftlines; B1.2 Sand beaches above the driftline; B1.3 Shifting coastal
dunes; B1.4 Coastal stable dune grassland (grey dunes); B1.8 Moist and wet dune slacks;
B2.1 Shingle beach driftlines; B2.2 Unvegetated mobile shingle beaches above the driftline;
C1.5 Permanent inland saline and brackish lakes, ponds and pools; D6.2 Inland saline or
89
brackish species-poor helophyte beds normally without free-standing water; E1.9 Open
non-Mediterranean dry acid and neutral grassland including inland dune grassland; E6.2
Coninental inland salt steppes; G1.C Highly ariicial broadleaved deciduous forestry
plantations; H5.3 Sparsely- or un-vegetated habitats on mineral substrates not resulting
from recent ice activity; H5.6 Trampled areas.
Land use: nature conservation and research – 95%; forestry – minor, tourism/recreation –
minor.
Protected areas: includes Dzharylhatskyi Naional Nature Park (10000 ha) and Emerald Site
“Dzharylhatskyi Naional Nature Park” (10000 ha), included in Ramsar Site “Karkinitska and
Dzharylgatska Bays”.
Threats: climate change/sea level rise – low, forestry (afforestation) – medium, natural
events (spit erosion) – low.
General description. The IPA includes Dzharylhach accumulaive island with adjacent areas
of the Black Sea. The island is composed of sand and shells. Its southern bank is steeper,
with low dunes (1–2 m). The nothern bank is very low, with numerous lagoons. A large part
of the island is occupied by dry grasslands in complex with saltmarshes and saline lakes.
Aquaic vegetaion. Dominants: Zostera nolii (Nanozostera nolii), Zostera marina, Ruppia
cirrhoza, Zannichellia palustris, Zannicelia pedunculata, Potamogeton pecinatus (Stuckenia
pecinata).
Tall helophytes communiies. Dominants: Phragmites australis, Cladium mariscus.
Sand beach dritline vegetaion. Dominants: Cakile maritima, Euphorbia peplis, Salsola soda
(Soda inermis), Polygonum mariimum.
Sand dunes vegetaion. Dominants: Leymus sabulosus, Crambe pontica. Typical species:
Argusia sibirica, Artemisia arenaria, Asperula graveolens, Carex colchica, Centaurea majorovii,
Centaurea odessana, Cynanchum acutum, Eryngium mariimum, Euphorbia seguierana,
Lactuca tatarica, Polygonum mesembricum, Secale sylvestre.
Halophyic vegetaion. Dominants: Aeluropus litoralis, Bolbochoenus mariimus, Elytrigia
elongata, Juncus gerardii, Juncus mariimus, Limonium caspium, Limonium meyeri,
Puccinellia distans, Puccinellia fominii, Salicornia perennans, Suaeda prostrata.
Dry grasslands on sands and shells. Dominants: Carex colchica, Carex praecox, Centaurea
majorovii, Cynodon dactylon, Ephedra distachya, Euphorbia seguierana, Festuca beckeri,
Festuca valesiaca, Poa angusifolia, Poa bulbosa, Secale sylvestre, Sipa borysthenica, Sipa
capillata. Typical species: Artemisia arenaria, Asperula setulosa, Centaurea odessana.
Mesic grasslands. Dominants: Calamagrosts epigeios, Elytrigia repens, Festuca pratensis,
Poa pratensis.
Botanical significance. Important area for conservaion of sand grasslands, litoral
vegetaion, mesic and wet halophyic vegetaion, vegetaion of salt waters.
Criterion C
• A2.5 Coastal saltmarshes and saline reedbeds; area: 2000 ha; trend: stable; area data
quality: medium; trend data quality: medium.
• A5 Sublitoral sediment; area: 4500 ha; trend: stable; area data quality: good; trend
data quality: good.
• B1.3 Shifting coastal dunes; area: 20 ha; trend: stable; area data quality: medium; trend
data quality: medium.
• B1.4 Coastal stable dune grassland (grey dunes); area: 20 ha; trend: stable; area data
quality: medium; trend data quality: medium.
• E1.2 Perennial calcareous grassland and basic steppes; area: 1000 ha; trend: stable;
area data quality: poor; trend data quality: medium.
90
Literature
1. Биоразнообразие Джарылгача: современное состояние и пути сохранения /
Котенко Т.И., Ардамацкая Т.Б., Дубына Д.В. и др. / Науч. ред. Т.И. Котенко, Ю.Р. ШелягСосонко. – Вестн. зоологии. – 2000. – Спец. выпуск. – 240 с.
2. Дубина Д.В., Дзюба Т.П., Ємельянова С.М. НПП Джарилгацький // Фіторізноманіття
заповідників і національних природних парків України. Ч. 2. Національні природні
парки / під ред. В.А. Онищенка і Т.Л. Андрієнко. – Київ: Фітосоціоцентр, 2012. –
С. 230–239.
3. Дубина Д.В., Дзюба Т.П. Фітоценотична різноманітність острова Джарилгач
(Херсонська обл.) // Укр. ботан. журн. – 2005. – 62, № 2. – С. 128–142.
4. Дубина Д.В., Тимошенко П.А. Особливості флористичного різноманіття острова
Джарилгач // Укр. ботан. журн. – 2004. – 61. № 3. – С. 61–72.
Dzhohul
I.I. Chorney, A.I. Tokaryuk, V.V. Budzhak
Ukrainian name: Джогуль.
Area: 263 ha.
Altitude: 944–1185 m.
Latitude: 47°48’35” N (47.8096°).
Longitude: 25°06’14” E (25.1038°).
Administrative regions. Chernivtsi region: Putyla raion.
Ownership: state (major), private.
Biogeographic regions: alpine.
Habitats. Level 1: C – 0.1%, E – 74.4%, G – 25.0%, H – 0.5%.
Habitats. Level 2: C2 Surface running waters – 0.1%, E1 Dry grasslands – 29.0%; E2 Mesic
grasslands – 41.4%, E3 Seasonally wet and wet grasslands – 1.0%; E7 Sparsely wooded
grasslands – 3.0%; G1 Broadleaved deciduous woodland – 10.0%; G3 Coniferous woodland
– 15.0%; H5 Miscellaneous inland habitats with very sparse or no vegetation – 0.5%.
Futher habitat description: C2.2 Permanent non-tidal, fast, turbulent watercourses; E1.7
Non-Mediterranean dry acid and neutral closed grassland; E2.1 Permanent mesotrophic
pastures and aftermath-grazed meadows; E2.2 Low and medium alitude hay meadows;
E3.4 Moist or wet eutrophic and mesotrophic grassland; G1.6 Fagus woodland; G3.1 Abies
and Picea woodland; H5.6 Trampled areas.
Land use: agriculture (animals) – minor; forestry – minor; mowing/hay making – major.
Protected areas: –.
Threats: abandonment/reducion of land management – medium; agricultural
intensification/expansion (grazing) – low.
General description. A complex of grasslands with small Picea abies woods, Fagus sylvaica
woods and sparse trees. Grasslands are dominated by Helictotrichon praeustum, Agrostis
tenuis, Anthoxanthum odoratum, Luzula luzuloides, Sieglingia decumbens, Lerchenfeldia
flexuosa with some presence of Arnica montana, Astrantia major, Botrychium lunaria,
Campanula serrata, Arabidopsis arenosa (Cardaminopsis arenosa), Carex umbrosa, Carlina
acaulis, Coeloglossum viride, Dianthus compactus, Euphorbia carniolica, Festuca pratensis,
Gentiana asclepiadea, Gymnadenia conopsea, Hypochaeris uniflora (Achyrophorus
uniflorus), Leucorchis albida, Lilium martagon, Listera ovata (Neoia ovata), Luzula
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multiflora, Melampyrum saxosum, Potentilla aurea, Potentilla erecta, Pyrethrum clusii,
Scorzonera rosea, Soldanella montana, Thesium alpinum, Thymus pulegioides, Traunsteinera
globosa, Trifolium montanum, Trollius europaeus, Vaccinium myrtillus, Vaccinium vitisidaea, Viola declinata.
– P. 169–187.
2. Чорней І.І., Буджак В.В., Токарюк А.І. Сторінками Червоної книги України
(рослинний світ). Чернівецька область. – Чернівці: ДрукАрт, 2010. – 452 с.
Dziurkach
I.I. Chorney, A.I. Tokaryuk, V.V. Budzhak
Ukrainian name: Дзюркач.
Area: 25.3 ha.
Altitude: 398–448 m.
Latitude: 48°18’03” N (48.3008°).
Longitude: 25°46’30” E (25.7748°).
Administrative regions. Chernivtsi region: Kitsman raion.
Ownership: state, private.
Biogeographic regions: alpine, coninental.
Habitats. Level 1: E – 59.0%, G – 39.5%, H – 1.5%.
Habitats. Level 2: E2 Mesic grasslands – 56.0%; G1 Broadleaved deciduous woodland –
39.5%; E7 Sparsely wooded grasslands – 3.0%; H5 Miscellaneous inland habitats with very
sparse or no vegetation – 1.5%.
Botanical significance. The area includes about 5% of naional populaion of narrow
endemic species Nigritella carpaica (Gymnadenia carpatica). Another Criterion A species
Campanula serrata is frequent here. A good example of species rich mountain meadows
with signiicant constancies of several Orchidaceae species.
Criterion A
• Nigritella carpatica (Zapał.) Teppner, Klein et Zagulski (Gymnadenia carpatica (Zapał.)
Teppner et Klein); A(iv); abundance: rare; trend: unknown; species data quality:
medium; trend data quality: poor.
• Campanula serrata (Kit. ex Schult.) Hendrych; A(ii); abundance: frequent; trend:
unknown; species data quality: good; trend data quality: poor.
Conservaion proposals. Maintain mowing of the meadows. Create a state botanical
reserve and an Emerald Site.
Literature
1. Teppner H., Klein Е., Drescher A., Zahulskij М. Nigritella carpatica (Orchidaceae) –
ein Reliktendemit der Ost-Karpaten // Phyton. Annales rei botanicae. – 1994. – 34, № 2.
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Futher habitat description: E2.2 Low and medium alitude hay meadows; G1.6 Fagus
woodland; H5.6 Trampled areas.
Land use: forestry – minor; mowing/hay making – major.
Protected areas: included in Chernivetskyi regional landscape park and Emerald Site
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“Chernivetskyi regional landscape park”.
Threats: –.
General description. Complex of species-rich hay meadows with small areas of broadleaved
woods and sparse trees. Meadows are dominated by Arrhenatherum elatior, Anthoxanthum
odoratum, Brachypodium pinnatum, Briza media, Dactylis glomerata, Cynosurus cristatus,
Festuca pratensis, Festuca rubra, Molinia caerulea agg, Trisetum flavescens. Other frequent
species: Adenophora liliifolia, Centaurea jacea, Crepis sibirica, Ferulago sylvatica, Filipendula
vulgaris, Galium verum, Laserpitium latifolium, Leucanthemum vulgare, Pedicularis exaltata,
Peucedanum oreoselinum, Pteridium aquilinum, Pyrethrum corymbosum, Serratula
tinctoria, Trifolium montanum, Trifolium pannonicum, Veratrum nigrum.
Botanical significance. Important area for Adenophora liliifolia. Extreme species rich
grassland.
Criterion A
• Adenophora liliifolia; A(ii); abundance: frequent; trend: unknown; species data quality:
good; trend data quality: poor.
Literature
1. Roleček J., Čornej I.I., Tokarjuk A.I. Understanding the extreme species richness of
semi-dry grasslands in east-central Europe: a comparaive approach // Preslia. – 2014. – 86:
1–XX. – P. 13–34.
Evergreen alpine and subalpine heath and scrub; F2.3 Subalpine deciduous scrub; F2.4
Conifer scrub close to the tree limit; G1.1 Riparian and gallery woodland, with dominant
Alnus, Betula, Populus or Salix; G1.6 Fagus woodland; G3.1 Abies and Picea woodland; G4.6
Mixed Abies – Picea – Fagus woodland; H2.3 Temperate-montane acid siliceous screes; H3.1
Acid silicious inland clifs; H5.6 Trampled areas.
Land use: forestry – minor, mowing/hay making – minor, nature conservation and research
– major.
Protected areas: includes Gorgany nature reserve (5344.2 ha), overlaps (about 3429 ha)
with Karparskyi naional nature park, includes Emerald Site “Gorgany Nature Reserve”
(5344.2 ha), overlaps (about 3429 ha) with Emerald Site “Carpathian Naional Nature Park”.
Threats: climate change/sea level rise – low, development (recreaion/tourism) – low
Gorgany
V.A. Onyshchenko
Ukrainian name: Горгани.
Transliteraion/Translaion variants: Horhany mountains.
Area: 9217.0 ha.
Altitude: 690–1755 m.
Latitude: 48°24’11” N (48.4030°).
Longitude: 24°25’03” E (24.4176°).
Administrative regions. Ivano-Frankivsk region: Nadvirna raion, Yaremche city.
Ownership: state (major), private.
Biogeographic regions: alpine.
Habitats. Level 1. C – 0.2%; E – 1.3%; F – 5.3%; G – 89.7%; H – 3.5%.
Habitats. Level 2. C2 Surface running waters – 0.2%; E1 Dry grasslands – 0.4%; E2 Mesic
grasslands – 0.3%; E4 Alpine and subalpine grasslands stands – 0.5%; E5 Woodland fringes
and clearings and tall forb stands – 0.1%; F2 Arctic, alpine and subalpine scrub – 5.3%; G1
Broadleaved deciduous woodland – 2.0%; G3 Coniferous woodland – 62.7%; G4 Mixed
deciduous and coniferous woodland – 25.0%; H2 Screes – 3.3%; H3 Inland cliffs, rock
pavements and outcrops – 0.1%; H5 Miscellaneous inland habitats with very sparse or no
vegetation – 0.1%.
Futher habitat description. C2.1 Springs, spring brooks and geysers; C2.2 Permanent nontidal, fast, turbulent watercourses; E1.7 Non-Mediterranean dry acid and neutral closed
grassland; E2.2 Low and medium alitude hay meadows; E3.4 Moist or wet eutrophic and
mesotrophic grassland; E4.2 Moss and lichen dominated mountain summits, ridges and
exposed slopes; E4.3 Acid alpine and subalpine grassland; E5.4 Moist or wet tall-herb and
fern fringes and meadows; E5.5 Subalpine moist or wet tall-herb and fern stands; F2.2
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General description. The major vegetaion type is Picea abies forest (pure or co-dominated by
Abies alba, Pinus cembra or Pinus sylvestris). Dominants of the herb layer are Calamagrosis
villosa, Luzula luzuloides, Vaccinium myrillus. Typical species of the herb layer are
Dryopteris dilatata, Homogyne alpina, Luzula sylvaica, Oxalis acetosella, Vaccinium viisidaea. The moss layer is formed by Polytrichum formosum, Pleurozium schreberi, Dicranum
scoparium, Bazzania trilobata, Leucobryum juniperoides. 10–15% of the coniferous forest
is the primeval forest. A large area is covered by mixed Fagus sylvaica – Abies alba – Picea
abies forests. Its herb layer consists of Anemone nemorosa, Dentaria glandulosa, Dryopteris
ilix-mas, Euphorbia amygdaloides, Galium odoratum, Glechoma hirsuta, Mercurialis perennis,
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Oxalis acetosella, Polygonatum vericillatum, Senecio ovatus, Symphytum cordatum, Viola
reichenbachiana. On the mountain tops, there are large areas of unvegetated coarse
sandstone screes and Pinus mugo thickets. Besides there are communities of Duschekia
alnobetula (Alnus alnobetula) and Juniperus sibirica.
Botanical significance. Important area for mountain Picea abies forests and Carpathian
Pinus cembra forests.
Criterion C
• F2.46 (*4070) Carpathian Pinus mugo scrub; area: 400 ha; trend: stable; area data
quality: medium; trend data quality: poor.
• G3.1B Alpine and Carpathian subalpine Picea forests; area: 5800 ha; trend: stable; area
data quality: medium; trend data quality: medium.
• G3.25 Carpathian Larix and Pinus cembra forests; area: 90 ha; trend: decreasing; area
data quality: medium; trend data quality: medium.
• H2.3 Temperate-montane acid siliceous screes; area: 300 ha; trend: stable; area data
quality: medium; trend data quality: good.
Literature
1.Берко Й.М. Фітоценотичний нарис смерекових лісів (Piceeta abietis) на Горганах //
Укр. ботан. журн. – 1970. – 28, № 5. – С. 608–613.
2.Клімук Ю.В., Міскевич У.Д., Якушенко Д.М., Чорней І.І., Буджак В.В., Нипорко С.О.,
Шпільчак М.Б., Чернявський М.В., Токарюк А.І. та ін. Природний заповідник “Горгани”.
Рослинний світ. – Природно-заповідні території України. Рослинний світ. Вип. 6. – К.:
Фітосоціоцентр, 2006. – 400 с.
3.Стойко С.М., Третяк П.Р., Бойчук І.І. Сосна кедрова (Pinus cembra L.) на верхній межі
лісу у Ґорґанах: хорологія, екологія, фенологія // Науковий вісник ДЛГУ: Дослідження,
охорона та збагачення біорізноманіття. – Львів: Вид-во ЛДГУ, 1999. – Вип. 99. –
С. 173–179.
4.Чорней І.І., Токарюк І.І., Буджак В.В. ПЗ Горгани // Фіторізноманіття заповідників
і національних природних парків України. Ч. 1. Біосферні заповідники. Природні
заповідники / під ред. В.А. Онищенка і Т.Л. Андрієнко. – Київ: Фітосоціоцентр, 2012. –
С. 94–101.
alpine and subalpine scrub – 16.0%; G1 Broadleaved deciduous woodland – 0.3%; G3
Coniferous woodland – 78.9%; G4 Mixed deciduous and coniferous woodland – 0.1%; H2
Screes – 4.0%; H3 Inland cliffs, rock pavements and outcrops – 0.1%; H5 Miscellaneous
inland habitats with very sparse or no vegetation – 0.2%.
Grofa
V.A. Onyshchenko
Ukrainian name: Грофа.
Transliteraion/Translaion variants: Hrofa Mount.
Area: 5610.0 ha.
Altitude: 815–1748 m.
Latitude: 48°35’19” N (48.5885°).
Longitude: 23°55’46” E (23.9293°).
Administrative regions. Ivano-Frankivsk region: Rozhniaiv raion (major); Zakarpatska
region: Tiachiv raion.
Ownership: state.
Biogeographic regions: alpine.
Habitats. Level 1. C – 0.1%; E – 0.3%; F – 16.0%; G – 79.3%; H – 4.3%.
Habitats. Level 2. C2 Surface running waters – 0.1%; E1 Dry grasslands – 0.3%; F2 Arctic,
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Futher habitat description. C2.1 Springs, spring brooks and geysers; C2.2 Permanent nontidal, fast, turbulent watercourses; E1.7 Non-Mediterranean dry acid and neutral closed
grassland; F2.2 Evergreen alpine and subalpine heath and scrub; F2.3 Subalpine deciduous
scrub; F2.4 Conifer scrub close to the tree limit; G1.6 Fagus woodland; G3.1 Abies and
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Picea woodland; G4.6 Mixed Abies – Picea – Fagus woodland; H2.3 Temperate-montane
acid siliceous screes; H3.1 Acid silicious inland clifs; H5.6 Trampled areas.
Land use: forestry – major; nature conservation and research – major, tourism/recreaion
– minor.
Protected areas: overlaps (about 2450 ha) with Grofa state landscape reserve, overlaps
(5375 ha) with Emerald Site “Dolynsko-Rozhniatynskyi”.
Threats: forestry (intensiied forest management) – low.
General description. The major vegetaion type is the mountain Picea abies forest on acid
soil. A part of spruce forests (>500 ha) has some admixture of Pinus cembra. Besides there
are large areas of Pinus mugo scrub and coarse sandtone screes in the subalpine belt.
Botanical significance. Important area for mountain Picea abies forests.
Criterion C
• F2.46 (*4070) Carpathian Pinus mugo scrub; area: 850 ha; trend: stable; area data
quality: medium; trend data quality: poor.
• G3.1B Alpine and Carpathian subalpine Picea forests; area: 4450 ha; trend: stable; area
data quality: good; trend data quality: poor.
Literature
1. Природно-заповідні території та об’єкти Івано-Франківщини. – Івано-Франківськ,
2000. – 272 с.
Perennial calcareous grassland and basic steppes; E1.9 Open non-Mediterranean dry acid
and neutral grassland including inland dune grassland; E2.2 Low and medium alitude
Hadiatski Luky
N.O. Stetsiuk, V.A. Onyshchenko
Ukrainian name: Гадяцькі луки.
Transliteraion/Translaion variants: Hadiatski Meadows, Hadiach Meadows, Gadiach
Meadows.
Area: 15123 ha.
Altitude: 93–108 m.
Latitude: 50°18’40” N (50.3111°).
Longitude: 34°02’04” E (34.0343°).
Administrative regions. Poltava region: Hadiach raion (major), Myrhorod raion; Sumy
region: Lebedyn raion.
Ownership: state (major), private.
Biogeographic regions: coninental.
Habitats. Level 1. C – 2.5%; D – 5.0%; E – 65.6%; F – 0.2%; G – 26.7%.
Habitats. Level 2. C1 Surface standing waters – 0.5%; C2 Surface running waters – 1.9%;
C3 Littoral zone of inland surface waterbodies – 0.1%; D5 Sedge and reedbeds, normally
without free-standing water – 5.0%; E1 Dry grasslands – 1.0%; E2 Mesic grasslands – 46.5%;
E3 Seasonally wet and wet grasslands – 16.1%; E6 Salt steppes – 2.0%; F9 Riverine and fen
scrubs – 0.2%; G1 Broadleaved deciduous woodland – 25.7%; G3 Coniferous woodland –
0.7%; G4 Mixed deciduous and coniferous woodland – 0.3%.
Futher habitat descripion. C1.2 Permanent mesotrophic lakes, ponds and pools; C1.6
Temporary lakes, ponds and pools; C2.3 Permanent non-tidal, smooth-flowing watercourses;
C3.2 Water-fringing reedbeds and tall helophytes other than canes; C3.5 Periodically
inundated shores with pioneer and ephemeral vegetation; D5.1 Reedbeds normally without
free-standing water; D5.2 Beds of large sedges normally without free-standing water; E1.2
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hay meadows; E3.4 Moist or wet eutrophic and mesotrophic grassland; E6.2 Coninental
inland salt steppes; F9.1 Riverine scrub; G1.1 Riparian and gallery woodland, with dominant
Alnus, Betula, Populus or Salix; G1.2 Mixed riparian floodplain and gallery woodland; G1.3
Mediterranean riparian woodland; G1.4 Broadleaved swamp woodland not on acid peat;
G1.A Meso- and eutrophic Quercus, Carpinus, Fraxinus, Acer, Tilia, Ulmus and related
woodland; G3.4 Pinus sylvestris woodland south of the taiga; G4.7 Mixed Pinus sylvestris –
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acidophilous Quercus woodland.
Land use: forestry – 30%; mowing/hay making – major, nature conservation and research
– major.
Protected areas: overlaps (about 9437 ha) with Hadiatskyi regional landscape park,
includes Pisotsko-Konkove regional landscape reserve (204 ha), includes Ternovyi Kushch
regional reserve (303 ha), includes Velyky Lis regional botanical reserve (182 ha), includes
Zozulyntsevi Luky regional botanical reserve (45 ha), overlaps (about 9437 ha) with Emerald
Site “Hadiatskyi Regional Landscape Park”.
Threats: abandonment/reducion of land management – low; agricultural intensiicaion/
expansion (general) – low; forestry (intensiied forest management) – low; development
(recreaion/tourism) – low.
General description. A part of the loodplain of the Psel river. The pastures are dominated
by Poa pratensis, Festuca pratensis, Dactylis glomerata, Elytrigia repens, Phleum pretense,
Bromopsis inermis, Agrosis gigantea, Lolium perrene, Calamogrosis epigeios. Their constant
species are Trifolium repens, T. pratens, Medicago lupulina, M. falcata, Vicia cracca, V.
villosa, V. tetrasperma, V. grandilora, Plantago media, Ranunculus polyanthemos, R. acris,
Rumex thyrsilorus, Equisetum pratense, Glechoma hederacea, Lysimachia nummulara,
Prunella vulgaris, Potenilla anserina. The largest area is occupied by mesic hay meadows.
They are dominated by Poa pratensis, Festuca pratensis, Alopecurus pratensis, Dactylis
glomerata, Elytrigia repens, Arhrenantherum elaius. Other species of high constancy are
Bromopsis inermis, Plantago lanceolata, Potenilla argentea, Galium ruthenicum, Cerasium
holosteoides, Medicago falcata, Psammophiliella muralis, Achillea submillefolium, Echium
vulgare, Verbascum lychniis, Veronica spicata, Hypericum perforatum, Centaurium
erythraea, Silene viscaria (Steris viscaria, Viscaria viscosa), Trifolium alpestre, Fragaria
viridis. In drier habitats, there dominate Festuca rubra, Festuca pseudovinae, Festuca beckeri,
Festuca valesiaca, Poa angusifolia, Anthoxanthum odoratum. In the moist grasslands, there
prevail Poa palustris, Beckmannia eruciformis, Agrosis stolonifera, in saline grasslands –
Festuca orientalis, Alopecurus arundinaceae, Carex distans, Juncus gerardii, Eleocharis
uniglumis, Plantago salsa, Boulboschoenus mariimus. Sedge and reedbeds are dominated
by Phragmites australis, Typha angusifolia, T. laifolia, Glyceria maxima, G. luitans,
Carex acuta, C. acuiformis, C. riparia, C. elata, Schoenoplectus lacustris, Phalaroides
arundinacea, Sium laifolium. Main aquaic dominants are Hydrocharis morsus-ranae,
Spirodella polyrhyza, Lemna minor, L. trisulca, Batrachium aquaile (Ranunculus aquailis),
B. rionii (Ranunculus rionii), Nuphar lutea, Ceratophyllum demersum, Potamogeton natans,
P. crispus, P. perfoliatus, P. pecinatum (Stuckenia pecinata), P. lucens. Mesic deciduous
forests on the slopes of the valley and in the loodplain are dominated by Quercus robur,
Tilia cordata, Acer platanoides, Acer campestre. Dominants of the shrub layer are Corylus
avellana (major), Euonymus europaeus, E. verrucosus, Frangula alnus (in moist places),
dominants of the herb layer – Stellaria holostea, Aegopodium podagraria, Convallaria
majalis. Forests dominanted by Alnus gluinosa have the herb layer consising mainly of
Urica galeopsifolia, Eupatorium cannabinum, Galium aparine, Carex acuiformis. Besides
there are forests dominated by Populus nigra, P. tremula, P. alba. Their typical species are
Aegopodium podagraria, Convallaria majalis, Frangula alnus, Glechoma hederacea, Poa
palustris, Rubus caesius, Sambucus nigra, Urica dioica.
Botanical significance. Important area for mesic loodplain hay meadows.
Criterion C
• E2.2 Low and medium alitude hay meadows; area: 6000 ha; trend: stable; area data
quality: medium; trend data quality: medium.
100
Literature
1. Стецюк Н.А., Ханнанова О.Р. Флоросозологическая характеристика
проектированного регионального ландшафтного парка “Гадячский” (Украина,
Полтавская область) // Структурно-функциональная организация и динамика
растительного покрова: материалы Всероссийской науч.-практ. конф. с международным
участием. – Самара, 2011. – С. 56–59.
2. Ханнанова О.Р. Раритетна флора регіонального ландшафтного парку “Гадяцький”
(Полтавська область)” // Актуальні проблеми ботаніки та екології: мaт-ли Міжнарод.
конф. молодих учених. – Умань, 2014. – С. 68–69.
Halitsynove
V.P. Kolomiychuk
Ukrainian name: Галіцинове.
Area: 7.4 ha.
Altitude: 8–13 m.
Latitude: 46°48’10” N (46.8028°).
Longitude: 31°57’25” E (31.9569°).
Administrative regions. Mykolaiv region: Vitovka (Zhovtnevyi) raion.
Ownership: state.
Biogeographic regions: steppic.
Habitats. Level 1. E – 97.0%; H – 2.0%; J – 1.0%.
Habitats. Level 2. E1 Dry grasslands – 97.0%; H5 Miscellaneous inland habitats with very
sparse or no vegetation – 2.0%; J4 Transport networks & other contructed hard-surfaced
areas – 1.0%.
Futher habitat description. E1.9 Open non-Mediterranean dry acid and neutral grassland,
including inland dune grassland; H5.6 Trampled areas.
Land use: nature conservation and research – major.
Protected areas: overlaps (about 7 ha) with Starohalicynivska regional botanical nature
monument.
Threats: abandonment/reducion of land management – low, burning of vegetaion – low.
General description. Grassland on the sand terrace of the Southern Bug (Buh) river.
Dominant species are Artemisia marschalliana, Thymus pallasianum, Secale sylvestre.
Other frequent species are Carex colchica, Chondrilla juncea, Euphorbia seguieriana, Rumex
acetosella, Jacobaea borysthenica (Senecio borysthenicus), Seseli tortuosum. Populaion of
Centaurea protomargaritacea occupies minimum 4 ha. Density of the species is between 2
and 30 specimens per 100 m2.
Botanical significance. Important area for the narrow endemic species Centaurea
protomargaritacea (one of two localiies).
Criterion A
• Centaurea protomargaritacea Klokov; A(iii); abundance: frequent; trend: unknown;
species data quality: good; trend data quality: poor.
Conservaion proposals: Do not allow aforestaion. Change the status of regional botanical
nature monument to state botanical nature monument.
Literature
1. Коломієць Г.В. Перлинні волошки секції Pseudophalolepis Klok. ряду Margaritacea
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Holohirskyi Lis
V.A. Onyshchenko
Ukrainian name: Гологірський ліс.
Transliteraion/Translaion variants: Holohirsky Forest, Gologirskyi Forest.
Area: 609.0 ha.
Klok. Питання систематики та охорони // Укр. фітоценологічний зб. – Вип. 1–2 (12–13).
– К.: Фітосоціоцентр, 1999. – С. 165–169.
2.Крицька Л.І., Деркач О.М. Сучасний стан популяцій видів ряду Margaritaceae Klok.
(Centaurea L.) // Укр. ботан. журн. – 1991. – 48, № 3. – С. 78 – 80.
3.Крицька Л.І., Деркач О.М. Волошка первинноперлинна Centaurea protomargaritacea
Klokov (C. margaritacea Ten. subsp. protomargaritacea (Klokov) Dostál) / Червона книга
України. Рослинний світ / за ред. Я.П. Дідуха. – К.: Глобалконсалтинг, 2009. – С. 308.
4.Перлини піщаної флори у пониззях Південного Бугу та Інгулу. Cерія: Збереження
біорізноманіття в Приморсько-степовому екокоридорі / Під ред. Г.В. Коломієць. – К.:
Громадська організація “Веселий Дельфін”, 2008. – 60 c.
102
Altitude: 277–461 m.
Latitude: 49°46'10" N (49.7693°).
Longitude: 24°39'40" E (24.6610°).
Administrative regions. Lviv region: Zolochiv raion.
Ownership: state.
Biogeographic regions: coninental.
Habitats. Level 1. E – 0.2%; G – 98.8%; H – 1.0%.
Habitats. Level 2. G1 Broadleaved deciduous woodland – 94.5%; G4 Mixed deciduous and
coniferous woodland – 4.3%; H5 Miscellaneous inland habitats with very sparse or no
vegetation – 1.0%.
Futher habitat descripion. G1.6 Fagus woodland; G1.A Meso- and eutrophic Quercus,
Carpinus, Fraxinus, Acer, Tilia, Ulmus and related woodland; H5.6 Trampled areas.
Land use: forestry – minor, nature conservation and research – 100%.
Protected areas: included in Pivnichne Podillia naional nature park, includes state
complex nature monument "Hora Vapniarka" (309.8 ha), includes regional preserve "Lis Pid
Trudovachem" (33 ha), includes regional geological nature monument "Velykyi Kamin" (0.03
ha), included in Emerald Site "Pivnichne Podillia".
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Threats: –.
General description. Hills covered by broadleaved forest. A large area is occupied by a
species-rich Fagus sylvaica forest on calcium rich soil.
Botanical significance. Important area for limestone beech forest.
Criterion C
• G1.66 Medio-European limestone Fagus forests (*9150); area: 200 ha; trend: stable;
area data quality: poor; trend data quality: medium.
Homilshanskyi Lis
V.A. Onyshchenko
Ukrainian name: Гомільшанський ліс.
Transliteraion/Translaion variants: Homilshansky Forest, Gomil'shansky Forest.
Area: 5450.0 ha.
Altitude: 80–200 m.
Latitude: 49°35'14" N (49.5872°).
Longitude: 36°18'28" E (36.3076°).
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Administrative regions. Kharkiv region: Zmiiv raion.
Ownership: state .
Biogeographic regions: steppic.
Habitats. Level 1. C – 1.0%; E – 0.7%; G – 97.3%; H – 1.0%.
Habitats. Level 2. C1 Surface standing waters – 0.3%; C2 Surface running waters – 0.7%;
E2 Mesic grasslands – 0.5%; E3 Seasonally wet and wet grasslands – 0.2%; G1 Broadleaved
deciduous woodland – 97.3%; H5 Miscellaneous inland habitats with very sparse or no
vegetation – 1.0%.
Futher habitat description. C1.2 Permanent mesotrophic lakes, ponds and pools; C2.3
Permanent non-tidal, smooth-flowing watercourses; E2.2 Low and medium alitude hay
meadows; E3.4 Moist or wet eutrophic and mesotrophic grassland; G1.2 Mixed riparian
floodplain and gallery woodland; G1.4 Broadleaved swamp woodland not on acid peat;
G1.A Meso- and eutrophic Quercus, Carpinus, Fraxinus, Acer, Tilia, Ulmus and related
woodland; H5.6 Trampled areas.
Land use: mowing/hay making – minor; forestry – 98%; nature conservaion and research
– 100%; tourism/recreaion – minor.
Protected areas: included in Homilshanski Lisy naional nature park and Emerald Site
"Gomilshanski Lisy Naional Nature Park".
Threats: forestry (intensiied forest management) – low.
General description. The major vegetaion type is the mesic broadleaved forest dominated
by Quercus robur. Other important species of the tree layer are Fraxinus excelsior, Tilia
cordata, Acer campestre, Acer platanoides. In the herb layer, there prevail Carex pilosa,
Stellaria holostea, Aegopodium podagraria, someimes Asarum europaeum, Mercurialis
perennis, Convallaria majalis, Galium odoratum. In spring there dominate Corydalis
marschalliana, Corydalis solida, Anemone ranunculoides, Ficaria verna, Scilla siberica,
someimes Allium ursinum, Tulipa quercetorum. Other species of high constancy are
Corylus avellana, Euonymus europaeus, Euonymus verrucosus, Gagea lutea, Lathyrus
vernus, Polygonatum mulilorum, Pulmonaria obscura, Ulmus glabra, Viola mirabilis, Viola
odorata. The forest belongs to the zones of strict protecion and regulated recreaion of the
naional nature park "Homilshanski Lisy". The western part of the IPA is the loodplain of
the Siverskyi Donets river. Vegetaion of this area is Quercus robur forests, Alnus gluinosa
forests, meadows and aquaic vegetaion.
Botanical significance. Important for conservaion of mesic deciduous forests.
Criterion C
• G1.A1 Quercus – Fraxinus – Carpinus betulus woodland on eutrophic and mesotrophic
soils; area: 5300 ha; trend: stable; area data quality: good; trend data quality: good.
Literature
1. Байрак О.М. Лишайники Гомольшанського державного природного парку // Укр.
ботан. журн. – 1987. – 44, № 4. – С. 38–42.
2. Вовк О.Г., Клімов О.В., Філатова О.В., Тверетінова В.В. Ботанічна характеристика
проектованого національного природного парку "Гомільшанські ліси" // Біологія та
валеологія: Зб. наук. праць. – Харків: ХДПУ, 2000. – Вип. 2. – С. 167–178.
3. Горелова Л.Н. Национальный природный парк "Гомольшанский" //
Характеристика основних заповідних територій Харківської області. – Харків: Мін-во
освіти і науки України, 2004. – С. 14–23.
4. Горелова Л.Н., Алехин А.А., Друлева И.В., Гамуля Ю.Г. Редкие и исчезающие
растения национального природного парка "Гомольшанские леса". – Харьков:
Видавничий центр ХНУ ім. В.Н. Каразіна, 2007. – 137 с.
105
5. Горєлова Л.М., Альохін О.О., Комариста В.П., Гамуля Ю.Г. Лікарські рослини
Національного природного парку "Гомільшанські ліси". – Харків: Видавничий центр
ХНУ ім. В.Н. Каразіна, 2006. – 122 с.
6. Леонтьев Д.В. Видовой состав миксомицетов (Myxomycota) национального
природного парка "Гомольшанские леса" (Украина) // Микол. и фитопатол. – 2006. –
40, вып. 2. – С. 101–107.
7. Леонтьев Д.В. Фитоценотические связи миксомицетов (Myxomycetes) в
Национальном природном парке "Гомольшанские леса" (Украина) // Экология. – 2007.
– № 2. – C. 1–3.
8. Перспективная сеть заповедных объектов Украины / Ю.Р. Шеляг-Сосонко,
С.М. Стойко, Я.П. Дидух и др. – К.: Наук. думка, 1987. – С. 128–132.
9. Саідахмедова Н.Б. Проблеми збереження фіторізноманіття в НПП "Гомільшанські
ліси" та шляхи їх вирішення // Каразінські прирородознавчі студії: Мат-ли наук.
конф. з міжнар. участю, присвяченої 100-річчю з дня народження Ю.М. Прокудіна і
О.М. Матвієнко – професорів Харківського університету. – Харків: Харк. нац. ун-т
ім. В.Н. Каразіна, 2011. – С. 68–71.
10. Саідахмедова Н.Б., Філатова О.В., Клімов О.В., Прилуцький О.В., Акулов
О.Ю., Біатов А.П. НПП Гомільшанські ліси // Фіторізноманіття заповідників і
національних природних парків України. Ч. 2. Національні природні парки / під ред.
В.А. Онищенка і Т.Л. Андрієнко. – Київ: Фітосоціоцентр, 2012. – С. 152–175.
11. Сивоконь О.В. Гастероїдні базидіоміцети Національного природного парку
"Гомільшанські ліси" // Заповідна справа в Україні. – 2008. – Т. 14, № 2. – С. 56–62.
"Podilski Tovtry Naional Nature Park".
Threats: agricultural intensification/expansion (grazing) – low, development (recreaion/
tourism) – low.
Horaivka
L.H. Lubinska, L.T. Horbnyak
Ukrainian name: Гораївка.
Area: 11.4 ha.
Altitude: 122–244 m.
Latitude: 48°34'57" N (48.5825°).
Longitude: 26°59'55" E (26.9985°).
Administrative regions. Kmelnytskyi region: Kamianets-Podilskyi raion.
Ownership: state (major), private.
Biogeographic regions: coninental.
Habitats. Level 1. E – 22.0%; F – 4.5%; G – 68.2%; H – 5.3%.
Habitats. Level 2. E1 Dry grasslands – 22.0%; F3 Temperate and mediterranean-montane
scrub – 4.5%; G1 Broadleaved deciduous woodland – 68.2%; H2 Screes – 4.0%; H3 Inland
clifs, rock pavements and outcrops – 1.3%.
Futher habitat descripion. E1.1 Inland sand and rock with open vegetation; E1.2
Perennial calcareous grassland and basic steppes; F3.1 Temperate thickets and scrub;
G1.7 Thermophilous deciduous woodland; G1.A Meso- and eutrophic Quercus, Carpinus,
Fraxinus, Acer, Tilia, Ulmus and related woodland; H2.6 Calcareous and ultra-basic screes of
warm exposures; H3.2 Basic and ultra-basic inland cliffs.
Land use: agriculture (animals) – minor, forestry – minor, nature conservation and research
– major, tourism/recreaion – minor.
Protected areas: included in Podilski Tovtry naional nature park, included in Emerald Site
General description. The steep slope of the Dnister valley with deciduous forests, dry
grasslands, shrub vegetaion, calcareous rocks and screes. Steppes are dominated by Sipa
сapillata, Festuca valesiaca, Elytrigia intermedia, Sesleria heuleriana. Typical species are
Leontodon hispidus, Pulsailla grandis, Pulsailla pratenis, Verbascum nigrum. On rocks
there are Allium podolicum, Asperula cynanchica, Aurinia saxailis, Melica transsylvanica,
Potenilla arenaria, Salvia vericillata, Sempervivum ruthenicum, Teucrium chamaedrys,
Thymus moldavicus, Veronica incana. The area belongs to the zone of strict protecion of
naional nature park "Podiliski Tovtry".
Botanical significance. One of the best populaions of Pulsailla grandis in Ukraine.
Criterion A
• Pulsailla grandis Wend.; A(ii); abundance: occasional (1000 individuals, including 300
generaive ones); trend: unknown; species data quality: good; trend data quality: poor.
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Administrative regions. Ternopil region: Pidvolochysk raion.
Ownership: state.
Biogeographic regions: coninental.
Habitats. Level 1. E – 98.8%; F – 1.0%; H – 0.2%.
Habitats. Level 2. E1 Dry grasslands – 90.0%; E2 Mesic grasslands – 8.8%; F3 Temperate and
mediterranean-montane scrub – 1.0%; H2 Screes – 0.1%; H3 Inland clifs, rock pavements
and outcrops – 0.1%
Futher habitat descripion. E1.1 Inland sand and rock with open vegetation; E1.2 Perennial
calcareous grassland and basic steppes; E2.2 Low and medium alitude hay meadows; F3.1
Temperate thickets and scrub; F3.2 Submediterranean deciduous thickets and brushes; H2.6
Calcareous and ultra-basic screes of warm exposures; H3.2 Basic and ultra-basic inland cliffs;
H3.5 Almost bare rock pavements, including limestone pavements.
Land use: nature conservation and research – 100%.
Protected areas: included in Medobory nature reserve, included in Emerald Site "Medobory
Nature Reserve".
Threats: abandonment/reducion of land management – medium, burning of vegetaion
– low.
General description. Three hills with limestone outcrops. The major vegetaion type is
the meadow steppe dominated by Poa angusifolia, Festuca valesiaca s.l., Carex humilis.
Someimes there dominate Calamagrosis epigeios, Brachypodium pinnatum, Festuca
pratensis. Species of high constancy are Achillea millefolium s.l., Acinos arvensis, Artemisia
absinthium, Cerasium arvense, Eryngium planum, Filipendula vulgaris, Fragaria viridis,
Galium verum, Hypericum perforatum, Koeleria cristata, Pedicularis kaufmannii, Plantago
media, Pimpinella saxifraga, Primula veris, Polygala comosa, Potenilla argentea, Ranunculus
polyanthemos, Salvia pratensis s.l. (Salvia dumetorum), Salvia vericillata, Scabiosa
ochroleuca, Jacobaea vulgaris (Senecio jacobaea), Seseli annuum, Thymus marschallianus,
Veronica incana, Viola hirta. Typical species on the outcrops are Asperula cynanchica,
Allium lusitanicum s.l. (Allium montanum auct.), Aurinia saxailis, Arabidopsis arenosa
(Cardaminopsis arenosa), Galium campanulatum, Poa compressa, Potenilla incana, Sedum
acre, Torella tortuosa, Tortula ruralis.
Botanical significance. The largest populaion of Dracocephalum austriacum in Ukraine,
probably one of two largest populaions of Pulsailla grandis in Ukraine. Important area for
conservaion of Carlina cirsioides.
Criterion A
• Carlina cirsioides Klokov; A(iv); abundance: occasional (500 individuals); trend: stable;
species data quality: good; trend data quality: medium.
• Dracocephalum austriacum L.; A(ii); abundance: frequent (3500 individuals); trend:
stable; species data quality: good; trend data quality: medium.
• Pulsailla grandis Wender.; A(ii); abundance: frequent (15000 individuals); trend:
unknown; species data quality: good; trend data quality: poor.
Conservaion proposals: Mow a part of the grasslands.
Literature
1. Szafer W. Geobotaniczne stosunki Miodoborόw Galicyjskich // Rozpr. Wydz.
Matemat.-Przyrod. Akad. Umiejętności. – 1910. – Ser. 3, Dz. B, T. 10. – S. 63–172.
2. Данилків І.С., Рабик І.В. Мохоподібні (Bryophytes) природного заповідника
"Медобори" // Чорноморський ботан. журн. – 2007. – 3, № 1. – С. 85–99.
3. Кондратюк С.Я. Лишайники заповідника "Медобори" // Укр. ботан. журн. – 1995.
– 52, № 1. – С. 141–144.
4. Кондратюк С.Я., Коломієць І.В. Нові для України види лишайників та ліхенофільних
грибів заповідника "Медобори" // Укр. ботан. журн. – 1997. – 54, № 1. – С. 42–47.
5. Оліяр Г. І. Рослини Червоної книги України в природному заповіднику "Медобори"
// Заповідна справа в Україні. – 1995. – Т. 1. – С. 11–12.
6. Оліяр Г. І. Фітораритети природного заповідника "Медобори" з філією
"Кременецькі гори" в міжнародних червоних списках // Природно-заповідний
фонд України – минуле, сьогодення, майбутнє. Матеріали міжнародної науковопрактичної конференції, присвяченої 20-річчю природного заповідника "Медобори"
(смт. Гримайлів, 26–28 травня 2010 р.) – Тернопіль: Підручники і посібники, 2010. –
С. 460–464.
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Horodnytski Tovtry
Ukrainian name: Городницькі товтри.
Transliteraion/Translaion variants: Gorodnytski Tovtry.
Area: 54.0 ha.
Altitude: 343–389 m.
Latitude: 49°24'12" N (49.4032°).
Longitude: 26°04'09" E (26.0691°).
H.I. Oliiar
7. Онищенко В.А. Рослинність карбонатних відслонень природного заповідника
"Медобори" // Укр. фітоцен. зб. – Сер. A. – 2001. – 1 (17). – С. 86–104.
8. Онищенко В.А., Оліяр Г.І. ПЗ Медобори // Фіторізноманіття заповідників і
національних природних парків України. Ч. 1. Біосферні заповідники. Природні
заповідники / під ред. В.А. Онищенка і Т.Л. Андрієнко. – Київ: Фітосоціоцентр, 2012. –
С. 254–276.
9. Смеречинская Т. А. Закономерности распространения лишайников по
фитоценозам природного заповедника "Медоборы" // Заповідна справа в Україні. –
2005. – Т. 11, Випуск 1. – С. 9–15.
Horodnytskyi Lis
V.A. Onyshchenko
Ukrainian name: Городницький ліс.
Transliteraion/Translaion variants: Horodnytskyi Forest, Gorodnytskiy Forest.
Latitude: 50°50'45" N (50.8460°).
Longitude: 27°18'19" E (27.3051°).
Administrative regions. Zhytomyr region: Novohrad-Volynskyi raion.
Ownership: state.
Biogeographic regions: coninental.
Habitats. Level 1. D – 0.1%; E – 0.2%; G – 99.2%; H – 0.5%.
Habitats. Level 2. D2 Valley mires, poor fens and transition mires – 0.1%; E3 Seasonally
wet and wet grasslands – 0.1%; G1 Broadleaved deciduous woodland – 28%; G3 Coniferous
woodland – 10.0%; G4 Mixed deciduous and coniferous woodland – 61.3%; H5 Miscellaneous
inland habitats with very sparse or no vegetation – 0.5%.
Futher habitat description. D2.3 Transiion mires and quaking bogs; G1.8 Acidophilous
Quercus-dominated woodland; G1.9 Non-riverine woodland with Betula, Populus tremula
or Sorbus aucuparia; G3.4 Pinus sylvestris woodland south of the taiga; G3.E Nemoral
bog conifer woodland; G4.4 Mixed Pinus sylvestris – Betula woodland; G4.7 Mixed Pinus
sylvestris – acidophilous Quercus woodland; H5.6 Trampled areas.
Land use: nature conservation and research – 100%; forestry – major.
Protected areas: same as Horodnytskyi state botanical reserve, included in Emerald Site
"Horodnytskyi".
Threats: –
General description. Major vegetaion types are mixed Pinus sylvestris – Quercus robur
forest, acidophilous Quercus robur forest, acidophilous Pinus sylvestris forest. The largest
area is occupied by mixed pine-oak forest with dominance of Rhododendron luteum
in the shrub layer and Vaccinium myrillus in the layer of dwarf shrubs and herbs. In the
lower layers of oak forest, there dominate Corylus avellana, Rhododendron luteum,
Carex brizoides, Vaccinium myrillus. Pine forest is dominated by Vaccinium myrillus and
Pleurozium schreberi. Besides there are small areas of swamped pine woods with Eriophorum
vaginatum, Vaccinium uliginosum, Sphagnum sp. and transiion mires dominated by Carex
lasiocarpa, Carex rostrata, Sphagnum sp.
Botanical significance. A good example of oak and mixed pine-oak acidophilous forests with
Rhododendron luteum.
Criterion C
• G1.8 Acidophilous Quercus-dominated woodland; area: 90 ha (together with mixed
oak-pine forests – 300 ha); trend: stable; area data quality: good; trend data quality:
medium.
Literature
1. Андриенко Т.Л., Шеляг-Сосонко Ю.Р. Растительный мир Украинского Полесья в
аспекте его охраны. – Киев: Наук. думка, 1983. – 215 с.
2. Фіторізноманіття Українського Полісся та його охорона / Під заг. ред.
Т.Л. Андрієнко. – Київ: Фітосоціоцентр, 2006. – 316 с.
Area: 343 ha.
Altitude: 195–202 m.
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Habitats. Level 1. E – 5.0%; F – 75.5%; G – 19.0%; H – 0.5%.
Habitats. Level 2. E1 Dry grasslands – 5.0%; F3 Temperate and mediterranean-montane
scrub – 75.5%; G1 Broadleaved deciduous woodland – 19.0%; H2 Screes – 0.1%; H3 Inland
clifs, rock pavements and outcrops – 0.4%.
Futher habitat descripion. E1.1 Inland sand and rock with open vegetation; E1.2
Perennial calcareous grassland and basic steppes; F3.1 Temperate thickets and scrub; F3.2
Submediterranean deciduous thickets and brushes; G1.A Meso- and eutrophic Quercus,
Carpinus, Fraxinus, Acer, Tilia, Ulmus and related woodland; H2.6 Calcareous and ultra-basic
screes of warm exposures; H3.2 Basic and ultra-basic inland cliffs; H3.5 Almost bare rock
pavements, including limestone pavements.
Land use: nature conservation and research – 100%; tourism/recreaion – minor.
Protected areas: included in Medobory nature reserve, included in Emerald Site "Medobory
Nature Reserve".
Threats: abandonment/reducion of land management – medium.
General description. A hill with limestone outcrops. Major vegetaion type is the shrubs
dominated by Crataegus leiomonogyna, Euonymus europaeus, Swida sanguinea (Cornus
sanguinea). On the northern slope, there is a small area of the broadleaved wood with a
dense shrub layer. On the ridge and in upper part of the southern slope, there is the steppe
vegetaion in complex with limestone outcrops. The stepe vegetaion is dominated by Carex
humilis, Dictamnus albus, Anthericum ramosum, Sipa pennata. Typical species on the
outcrops are Asperula cynanchica, Aurinia saxailis, Centaurea stoebe, Festuca valesiaca,
Galium campanulatum, Koeleria cristata, Melica transsilvanica, Potenilla incana, Sedum
acre, Seseli libanois, Thymus dimorphus, Veronica incana. Main threat is overgrowing by
shrubs and trees. Management includes clearing of shrubs and trees on a part of the area.
Botanical significance. Important area for conservaion of Dracocephalum austriacum and
Schivereckia podolica (Draba podolica).
Criterion A
• Dracocephalum austriacum L.; A(ii); abundance: occasional (230 individuals); trend:
luctuaing; species data quality: good; trend data quality: good.
• Schivereckia podolica (Besser) Andrz. ex DC. (Draba podolica (Besser) Rupr.); A(ii);
abundance: frequent (3000 individuals); trend: decreasing; species data quality: good;
trend data quality: medium.
Conservaion proposals: Do not allow natural aforestaion.
Literature
1. Szafer W. Geobotaniczne stosunki Miodoborόw Galicyjskich // Rozpr. Wydz.
Matemat.-Przyrod. Akad. Umiejętności. – 1910. – Ser. 3, Dz. B, T. 10. – S. 63–172.
2. Данилків І.С., Рабик І.В. Мохоподібні (Bryophytes) природного заповідника
"Медобори" // Чорноморський ботан. журн. – 2007. – 3, № 1. – С. 85–99.
3. Кондратюк С.Я. Лишайники заповідника "Медобори" // Укр. ботан. журн. – 1995.
– 52, № 1. – С. 141–144.
4. Кондратюк С.Я., Коломієць І.В. Нові для України види лишайників та ліхенофільних
грибів заповідника "Медобори" // Укр. ботан. журн. – 1997. – 54, № 1. – С. 42–47.
5. Оліяр Г.І. Зміни у флорі природного заповідника "Медобори" за минуле сторіччя
// Роль природно-заповідних територій Західного Поділля та Юри Ойцовської у
збереженні біологічного та ландшафтного різноманіття, – Гримайлів, 2003. – С. 333–338.
6. Оліяр Г.І. Фітораритети природного заповідника "Медобори" з філією
"Кременецькі гори" в міжнародних червоних списках // Природно-заповідний
фонд України – минуле, сьогодення, майбутнє. Матеріали міжнародної науковопрактичної конференції, присвяченої 20-річчю природного заповідника "Медобори"
(смт. Гримайлів, 26–28 травня 2010 р.) – Тернопіль: Підручники і посібники, 2010. –
С. 460–464.
7. Онищенко В.А. Рослинність карбонатних відслонень природного заповідника
"Медобори" // Укр. фітоцен. зб. – Сер. A. – 2001. – 1 (17). – С. 86–104.
8. Онищенко В.А., Оліяр Г.І. ПЗ Медобори // Фіторізноманіття заповідників і
національних природних парків України. Ч. 1. Біосферні заповідники. Природні
заповідники / під ред. В.А. Онищенка і Т.Л. Андрієнко. – Київ: Фітосоціоцентр, 2012. –
С. 254–276.
9. Смеречинская Т. А. Закономерности распространения лишайников по фитоценозам природного заповедника "Медоборы" // Заповідна справа в Україні. – 2005. – Т. 11,
Вип. 1. – С. 9–15.
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Hostra Skelia
Ukrainian name: Гостра скеля.
Transliteraion/Translaion variants: Gostra Skelya, Sharp Clif, Sharp Rock.
Area: 8.0 ha.
Altitude: 319–356 m.
Latitude: 49°21'24" N (49.3565°).
Longitude: 26°04'39" E (26.0774°).
Administrative regions. Ternopil region: Husiatyn raion.
Ownership: state.
Biogeographic regions: coninental.
H.I. Oliiar
Hrakove
N.O. Stetsiuk
Ukrainian name: Гракове.
Transliteraion/Translaion variants: Grakove.
Area: 768 ha.
Altitude: 80–85 m.
Latitude: 49°34’32” N (49.5757°).
Longitude: 32°54’57” E (32.9157°).
Administrative regions. Poltava region: Semenivka raion.
Ownership: state, municipal.
Biogeographic regions: coninental.
Habitats. Level 1. C – 7.8%; D – 23.4%; E – 68.1%; G – 0.7%.
Habitats. Level 2. C1 Surface standing waters – 1.3%; C3 Littoral zone of inland surface
waterbodies – 6.5%; D6 Inland saline and brackish marshes and reedbeds – 23.4%; E6 Salt
steppes – 68.1%; G1 Broadleaved deciduous woodland – 0.7%.
Land use: mowing/hay making – major, nature conservation and research – major.
Protected areas: includes Hrakove state hydrological reserve (500 ha).
Threats: abandonment/reducion of land management – low; agricultural intensiicaion/
expansion (general) – low.
General description. Major vegetaion types are the halophyic wet and mesic grasslands
dominated by Alopecurus arundinaceus, Beckmannia eruciformis, Carex distans, Juncus
gerardii, Festuca regeliana, Puccinellia distans with presence of Carex secalina, Glaux
mariima, Limonium alutaceum, Plantago salsa, Taraxacum bessarabicum, Triglochin
mariima, Triglochin palustris. Large areas are covered by halophyic non-inundated
Phragmites australis beds. There are marshes dominated by Bolboschoenus mariimus,
Schoenoplectus tabernaemontani. On the most saline soils, there are communiies of
Lepidium crassifolium with Salicornia herbacea, Halimione pedunculata, Halimione
verrucifera. Brackish lakes are overgrown with Phragmites australis and Typha angusifolia.
Botanical significance. One of the best sites with halophyic vegetaion in the coninental
biogeographic region of Ukraine.
Criterion A
• Carex secalina Wahlenb.; A(ii); abundance: abundant; trend: stable; species data quality:
medium; trend data quality: poor.
Criterion C
• E6.2 Coninental inland salt steppes; area: 500 ha; trend: stable; area data quality:
medium; trend data quality: medium.
Literature
1. Байрак О.М., Проскурня М.І., Стецюк Н.О., Слюсар М.В., Томін Є.М.,
Гостудим О.М. Еталони природи Полтавщини. Розповідні про заповідні території
Полтавщини. Науково-популярне видання. – Полтава: Верстка, 2003. – 212 с.
Hranitno-Stepove Pobuzhia
Futher habitat descripion. C1.5 Permanent inland saline and brackish lakes, ponds and
pools; C3.2 Water-fringing reedbeds and tall helophytes other than canes; D6.1 Inland
saltmarshes; D6.2 Inland saline or brackish species-poor helophyte beds normally without
free-standing water; E6.2 Continental inland salt steppes .
V.A. Onyshchenko
Ukrainian name: Гранітно-степове Побужжя.
Transliteraion/Translaion variants: Granitno-Stepove Pobuzhzhya, Granite-Steppe
Southern Buh Area.
Area: 3815.0 ha.
Altitude: 17–120 m.
Latitude: 47°53’25” N (47.8903°).
Longitude: 31°06’19” E (31.1053°).
Administrative regions. Mykolaiv region: Arbuzynka raion, Domanivka raion, Pervomaisk
raion, Voznesensk raion.
Ownership: state, unknown.
Biogeographic regions: steppic.
Habitats. Level 1. C – 21.6%; D – 0.5%; E – 48.8%; F – 2.0%; G – 22.7%; H – 4.4% .
Habitats. Level 2. C1 Surface standing waters – 5.0%; C2 Surface running waters – 16.5%;
C3 Littoral zone of inland surface waterbodies – 0.1%; D5 Sedge and reedbeds, normally
without free-standing water – 0.5%; E1 Dry grasslands – 48.6%; E2 Mesic grasslands – 0.1%;
E3 Seasonally wet and wet grasslands – 0.1%; F3 Temperate and mediterranean-montane
scrub – 2.0%; G1 Broadleaved deciduous woodland – 22.7%; H2 Screes – 0.2; H3 Inland
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cliffs, rock pavements and outcrops – 4.0%; H5 Miscellaneous inland habitats with very
sparse or no vegetation – 0.2%.
Futher habitat description. C1.2 Permanent mesotrophic lakes, ponds and pools; C1.3
Permanent eutrophic lakes, ponds and pools; C2.2 Permanent non-idal, fast, turbulent
watercourses; C2.3 Permanent non-idal, smooth-lowing watercourses; E1.1 Inland sand
and rock with open vegetation; E1.2 Perennial calcareous grassland and basic steppes; F3.1
Temperate thickets and scrub; F3.2 Submediterranean deciduous thickets and brushes;
G1.1 Riparian and gallery woodland, with dominant Alnus, Betula, Populus or Salix; G1.7
Thermophilous deciduous woodland; G1.C Highly ariicial broadleaved deciduous forestry
plantaions; H3.1 Acid silicious inland clifs; H5.6 Trampled areas.
Land use: agriculture (animals) – 15%; agriculture (arable) – minor; forestry – 15%; nature
conservaion and research – 100%; water management – 7%.
Protected areas: included in Buzkyi Hard naional nature park and Emerald Site “Bugzkyi
Gard Naional Nature Park”.
Threats: construcion/impact of dyke/dam/barrage – low; development (recreaion/
tourism) – low; forestry (aforestaion) – low.
General description. The Southern Bug valley with high (10–60 m) steep slopes and clifs
of granite and gneiss. Large areas are covered by the steppe vegetaion, natural Quercus
robur termophilous woods, ariicial Quercus robur woods, steppic and loodplain scrub,
loodplain decidous woods.
Natural deciduous termophilous woods. Dominants: Quercus robur (major), Acer campestre,
Fraxinus excelsior, Tilia cordata, Ulmus minor; Acer tataricum, Coinus coggygria, Crataegus
fallacina, Euonymus europaeus, Eunymus verrucosus, Swida sanguinea. Typical species:
Malus sylvestris, Pyrus communis; Chelidonium major, Galium aparine, Geum urbanum,
Urica dioica.
Floodplain woods. Dominants: Alnus gluinosa, Populus alba, Populus nigra, Salix alba, Salix
fragilis.
Shrubs. Dominant: Crataegus praearmata, Spiraea crenata, Prunus stepposa.
Dry grasslands (steppes). Dominants: Sipa capillata, Sipa lessingiana, Festuca valesiaca,
Botriochloa ischaemum, Koelerua cristata, Galatella villosa. Typical species: Artemisia
marschalliana, Chamaecyisus austriacus, Galium verum, Medicago romanica, Poa
angusifolia, Potenilla incana, Salvia nemorosa, Salvia nutans. Dominants in shrub steppes:
Amygdalus nana, Caragana frutex. In steppes on shallow stony soils, typical species are
Allium podolicum, Asperula rumelica, Genista inctoria, Pilosella oicinarum, Poa bulbosa,
Seseli pallasii, Teucrium polium, Verbascum phoeniceum.
Open silicious outcrops. Typical species: Achillea ochroleuca, Alyssum murale, Aurinia
saxailis, Rumex fascilobus, Sedum acre, Sedum borissovae, Sempervivum ruthenicum,
Thymus dimorphus; Ceratodon purpureus, Grimmia laevigata, Grimmia ovalis, Grimmia
pulvinata, Polytrichum piliferum, Polytrichum juniperinum, Syntrichia ruralis, Tortula muralis.
Shaded silicious outcrops. Dominants: Hypnum cupressiforme, Hedwigia ciliata,
Homalothecium sericeum, Leucodon sciuroides. Typical species: Bryum capillare,
Amblystegium serpens, Leskea polycarpa, Radula complanata.
Litoral and mire vegetaion. Dominants: Agrosis stolonifera, Bolboschienus mariimus,
Carex riparia, Glyceria maxima, Phalaroides arundinacea, Phragmites australis.
Aquaic vegetaion. Dominants: Ceratophyllum demersum, Myriophyllum spicatum, Nuphar
lutea, Potamogeton nodosus, Vallisneria spiralis.
Botanical significance. Important for conservaion of endemic species of silicious outcrops
(Dianthus hypanicus Andrz., Moehringia hypanica Grynj et Klokov, Cerasus klokovii
Sobko, Sedum borissovae Balk., Silene hypanica Klokov, Silene sytnikii Krytzka, Novosad et
Protopopova, Tulipa hypanica Klokov et Zoz, Onosma graniicola Klokov), dry grasslands,
termophilous oak woods and steppic scrub.
Criterion A
• Cerasus klokovii Sobko; A(iii); abundance: occasional; trend: stable; species data quality:
medium; trend data quality: poor.
• Dianthus hypanicus Andrz.; A(i), A(ii), A(iii), abundance: occasional; trend: stable;
species data quality: medium; trend data quality: poor.
• Gymnospermium odessanum (DC.) Takht.; A(iv); abundance: unknown; trend: unknown;
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species data quality: poor; trend data quality: poor.
Moehringia hypanica Grynj et Klokov; A(i), A(ii); abundance: rare (one locality); trend:
decreasing; species data quality: good; trend data quality: medium.
• Onosma graniicola Klokov; A(iv); abundance: 400–500 individuals (one locality, 1.5–2.0
ha); trend: luctuaing; species data quality: good; trend data quality: medium.
• Silene hypanica Klokov; A(iii); abundance: rare; trend: luctuaing; species data quality:
good; trend data quality: medium.
• Silene sytnikii Krytzka, Novosad et Protopopova; A(iii); abundance: occasional; trend:
stable; species data quality: good; trend data quality: medium.
• Tulipa hypanica Klokov et Zoz; A(iii); abundance: frequent; trend: stable; species data
quality: good; trend data quality: medium.
Criterion C
• E1.11 Euro-siberian pioneer rock debris swards; area: 10 ha; trend: unknown; area data
quality: poor; trend data quality: poor.
• E1.2 Perennial calcareous grassland and basic steppes; area: 1500 ha; trend: stable;
area data quality: medium; trend data quality: medium.
• F3.247 Ponto-Sarmatic deciduous thickets; area: 75 ha; trend: stable; area data quality:
medium; trend data quality: medium.
• H3.1 Acid siliceous inland cliffs; area: 150 ha; trend: stable; area data quality: medium;
trend data quality: good.
Literature
1. Mikhailyuk T.I., Demchenko E.M., Kondratyuk S.Ya. Algae of granitе outcrops from
the left bank of Pivdennyi Bug river (Ukraine) // Biologia. – Bratislava, 2003. – 58 (4). –
P. 589–601.
2. Гревцова Г.Т. Кизильники гранітно-степового Побужжя // Збірник наукових праць
Полтавського пед. ун-ту. – 2003. – Вип. 4 (31). – С. 54–61.
3. Гринь Ф.О., Клоков М.В. Новий вид мерингії з гранітів р. Південного Бугу // Бот.
журн. АН УРСР. – 1950. – 7, № 4 – С. 55–60.
4. Деркач О.М. Доповнення до флори Правобережного степу України // Укр. бот.
журн. – 1990. – 47, № 6. – С. 84–85.
5. Драбинюк Г.В. НПП Бузький Гард // Фіторізноманіття заповідників і національних
природних парків України. Ч. 2. Національні природні парки / під ред. В.А. Онищенка і
Т.Л. Андрієнко. – Київ: Фітосоціоцентр, 2012. – С. 49–60.
6. Коломієць Г.В., Таращук С.В. Бузькі брояки // Водно-болотні угіддя України.
Довідник / під ред. Марушевського Г.Б., Жарук І.С. – К.: Чорноморська програма
Ветландс Интернешнл. – 2006. – С. 188–193.
7. Михайлюк Т.И., Дариенко Т.М., Демченко Э.Н. Водоросли гранитных обнажений
регионального ландшафтного парка “Гранитно-степное Побужье” (Николаевская обл.,
Украина) // Новости систематики низших растений. – 2003. – 37. – С. 53–71.
8. Новосад В.В., Крицкая Л.И., Протопопова В.В. Новий для науки ендемічний
вид Гранітно-степового Побужжя смілка Ситника (Silene sytnikii Krytzka, Novosad
et Protopopova), його таксономічні, еколого-ценотичні, хорологічні, генезисні та
созологічні особливості // Укр. бот. журн. – 1996. – 53, № 5. – С. 578–585.
9. Партика Л.Я., Вірченко В.М., Нипорко С.О. 2006. До бріофлори регіонального
ландшафтного парку “Гранітно-степове Побужжя” // Чорномор. ботан. журн. – 2006. –
2, № 1. – С. 116–122.
10. Собко В.Г. Ендемічні та реліктові елементи флори гранітних відслонень
Придніпровської височини // Укр. бот. журн. – 1972. – 29, № 5. – C. 624–630.
11. Соломаха В.А., Драбинюк Г.В., Вініченко Т.С., Мойсієнко І.І., Деркач О.М.
Адаптивні особливості південнобузьких ендемів Dianthus hypanicus Andrz. та
Moehringia hypanica Grynj et Klok. // Укр. фітоцен. зб. – Сер. С. – 2006. – Вип. 24. – С.
70–86.
12. Червона книга України. Рослинний світ / за ред. Я.П. Дідуха. – К.:
Глобалконсалтинг, 2009. – 900 с.
13. Щербакова О.Ф. Особливості популяційної структури чистецю вузьколистого
Stachys angustifolia M. Bieb в Гранітно-Степовому Побужжі та проблеми його охорони
// Актуальні проблеми ботаніки та екології. Збірн. наук. праць. – 2005. – Вип. 1. –
С. 96–104.
14. Щербакова О.Ф., Новосад В.В., Крицька Л.І. Раритетний флорофонд КодимоЄланецького Побужжя (ЧКУ, 2009): популяційні та флоросозологічні аспекти. Рослинний
світ у Червоній книзі України: впровадження Глобальної стратегії збереження рослин.
Мат-ли міжнар. конф. (11–15 жовтня 2010 р., м.Київ). – Київ: Альтерпрес, 2010. –
С. 210–214.
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•
Ihrovets – Tavpishyrka
V.A. Onyshchenko, R.Ya. Kish
Ukrainian name: Ігровець – Тавпіширка.
Area: 6538.0 ha.
Altitude: 1062–1836 m.
Latitude: 48°32’36” N (48.5433°).
Longitude: 24°07’15” E (24.1207°).
Administrative regions. Ivano-Frankivsk region: Bohorodchany, Nadvirna raion, Rozhniaiv;
Zakarpatska region: Tiachiv raion.
Ownership: state.
Biogeographic regions: alpine.
Habitats. Level 1. E – 0.6%; F – 16.8%; G – 78.6%; H – 4.0%.
Habitats. Level 2. F2 Arctic, alpine and subalpine scrub – 16.8%; G1 Broadleaved deciduous
woodland – 0.4%; G3 Coniferous woodland – 77.7%; G4 Mixed deciduous and coniferous
woodland – 0.5%; H2 Screes – 3.8%; H3 Inland cliffs, rock pavements and outcrops – 0.2%.
Futher habitat description. F2.4 Conifer scrub close to the tree limit; G1.6 Fagus woodland;
G3.1 Abies and Picea woodland; G4.6 Mixed Abies – Picea – Fagus woodland; H2.3
Temperate-montane acid siliceous screes; H3.1 Acid silicious inland clifs.
Land use: forestry – minor; nature conservation and research – minor.
Protected areas: overlaps with Tavpyshyrkivskyi state botanical reserve (about 390 ha),
includes state botanical reserve “Gorgany i Tavpishyrka” (248 ha), includes Syvulia regional
nature preserve (17 ha), includes Tavpishyrka regional nature preserve (33 ha), overlaps
with regional nature preserve “Urochyshche Salatruk” (about 100 ha), overlaps (1476 ha)
with Emerald Site “Dolynsko-Rozhniatynskyi”.
Threats: climate change/sea level rise – low, forestry (intensiied forest management) – low.
General description. Forests dominated by Picea abies, Pinus mugo shrub, coarse sandstone.
Dominants of the lower layers of the spruce woods areLower layers are dominated by
Vaccinium myrillus, Homogyne alpina, Oxalis acetosella, Calamagrosis arundinacea. Small
areas are occupied by Pinus cembra woods.
Botanical significance. Important area for communiies of Pinus mugo, Carpathian Pinus
Irpinskyi Lis
Ukrainian name: Ірпінський ліс.
Transliteraion/Translaion variants: Irpin’ Forest.
Area: 307.0 ha
Altitude: 112-165 m.
Latitude: 50°30’42” N (50.5115°).
Longitude: 30°16’37” E (30.2769°).
Administraive regions. Kyiv city: Sviatoshyn raion.
Ownership: state.
Biogeographic regions: coninental.
cembra forests and subspecies Larix decidua ssp. polonica.
Criterion C
• F2.46 (*4070) Carpathian Pinus mugo scrub; area: 1100 ha; trend: stable; area data
quality: good; trend data quality: poor.
• G3.25 Carpathian Larix and Pinus cembra forests; area: 15 ha; trend: decreasing; area
data quality: medium; trend data quality: medium.
120
V.A. Onyshchenko, O.I. Pryadko
Habitats. Level 1. G – 98.0%; H – 2.0%
Habitats. Level 2. G1 Broadleaved deciduous woodland – 42.0%; G3 Coniferous woodland
– 2.0%; G4 Mixed deciduous and coniferous woodland – 50.0%; G5 Lines of trees, small
anthropogenic woodlands, recently felled woodland, early-stage woodland and coppice –
4.0%; H5 Miscellaneous inland habitats with very sparse or no vegetation – 2.0%.
Futher habitat description. G1.7 Thermophilous deciduous woodland; G3.4 Pinus sylvestris
woodland south of the taiga, G4.C Mixed Pinus sylvestris – thermophilous Quercus
woodland, H5.6 Trampled areas.
Land use: forestry – major; nature conservaion and research – major.
Protected areas: included in Holosiivskyi naional nature park, included in Emerald Site
121
“Holosiivskyi Naional Nature Park”.
Threats: unsustainable plant exploitaion – low.
General description. A part of the terrace of the Irpin river over the loodplain. Quercus
robur and mixed Quercus robur - Pinus sylvestris forest with the herb layer dominated by
Carex michelii, Clemais recta, Convallaria majalis. Constant species: Agrosis capillaris,
Convallaria majalis, Campanula bononiensis, Cruciata glabra, Euphorbia cyparissias,
Fallopia dumetorum, Festuca rubra, Fragaria vesca, Frangula alnus, Galeopsis bifida, Galium
mollugo, Geranium roberianum, Geranium sanguineum, Geum urbanum, Hylotelephium
polonicum, Hypericum perforatum, Impaiens parvilora, Malus sylvestris, Melampyrum
nemorosum, Moehringia trinervia, Peucedanum oreoselinum, Pinus sylvestris, Polygonatum
odoratum, Populus tremula, Potenilla alba, Pteridium aquilinum, Pyrus communis, Quercus
robur, Stachys officinalis, Solidago virgaurea, Sorbus aucuparia, Torilis japonica.
Botanical significance. One of the largest populaions of Iris hungarica in Ukraine.
Criterion A
• Iris hungarica Waldst. & Kit.; A(ii); abundance: frequent (~1000 clones); trend: stable;
species data quality: medium; trend data quality: poor.
Conservation proposals: include in the zone of strict protecion of the Holosiivskyi naional
nature park.
Literature
1. Онищенко В.А., Прядко О.І., Арап Р.Я., Дацюк В.В. Світлі дубові ліси СвятошинськоБіличанського відділення НПП “Голосіївський” // Природоохороні території в минулому,
сучасному й майбутньому світі (до 130-річчя створення “Пам’ятки Пеняцької” – першої
природоохоронної території у Європі): Мат-ли Другої міжнародної конференції (Львів
– Броди – Пеняки, 26–27 жовтня 2016 року). – Львів: Ліга-Прес, 2016. – С. 189–192.
dominated by Bothriochloa ischaemum, Carex humilis, Elytrigia intermedia, Festuca
valesiaca, Koeleria cristata, Poa angustifolia, Stipa capillata. Other frequent species are
Achillea submillefolium, Agrimonia eupatoria, Asperula cynanchica, Astragalus austriacus,
A. onobrychis, Chamaecytisus podolicus, Filipendula vulgaris, Fragaria viridis, Galium verum,
Gypsophila thyraica, Inula ensifolia, Knautia arvensis, Onobrychis arenaria, Pimpinella
saxifraga, Potentilla arenaria, Poterium sanguisorba, Ranunculus polyanthemos, Salvia
pratensis, S. verticillata, Securigera varia, Teucrium chamaedrys, T. pannonicum, Thalictrum
minus, Thymus marschallianus, Trifolium montanum, Viola hirta.
Kadubivska Sinka
I.I. Chorney, A.I. Tokaryuk, V.V. Budzhak
Ukrainian name: Кадубівська стінка.
Transliteraion/Translaion variants: Kadubivska Wall, Kadubivska Clif.
Area: 27.0 ha.
Altitude: 226-270 m.
Latitude: 48°34’02” N (48.5672°).
Longitude: 25°45’55” E (25.7653°).
Administraive regions. Chernivtsi region: Zastavna raion.
Ownership: state.
Biogeographic regions: coninental.
Habitats. Level 1. E – 99.0%; F – 0.7%; H – 0.3%
Habitats. Level 2. E1 Dry grasslands – 99.0%; F3 Temperate and mediterranean-montane
scrubs – 0.7%; H3 Inland cliffs, rock pavements and outcrops – 0.3%.
Futherhabitat description: E1.2 Perennial calcareous grassland and basic steppes; F3.2
Submediterranean deciduous thickets and brushes; H3.2 Basic and ultra-basic inland clifs.
Landuse: agriculture (animals) – major; nature conservaion and research – major.
Protected areas: includes Kadubivska Sinka state landscape reserve (22.8 ha).
Threats: agricultural intensiicaion/expansion (grazing) – low.
General description. Slopes with grasslands and gypsum outcrops. Grasslands are
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Botanical significance. One of the largest populaions of Chamaecytisus podolicus, locus
classucus of this species. The area is important also for Gypsophila thyraica and Iris
hungarica.
Criterion A
• Chamaecytisus podolicus (Błocki) Klásk. A(iv); abundance: frequent; trend: unknown;
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species data quality: good; trend data quality: poor.
Gypsophila thyraica A.Krasnova A(iiі); abundance: occasional; trend: unknown; species
data quality: good; trend data quality: poor.
• Iris hungarica Waldst. & Kit.; A(ii); abundance: rare; trend: unknown; species data
quality: good; trend data quality: poor.
Literature
1. Коротченко І.А., Токарюк А.І. Флора та рослинність степів ландшафтного
заказника “Кадубівська стінка” (Чернівецька область) // Наук. вісник Чернівецкого унту: Зб. наук. праць. – Чернівці: Рута, 2004. – Вип. 194. – С. 117–127.
2. Чорней І.І., Буджак В.В., Токарюк А.І. Сторінками Червоної книги України
(рослинний світ). Чернівецька область. – Чернівці: ДрукАрт, 2010. – 452 с.
3. Чорней І.І., Скільський І.В., Коржик В.П., Буджак В.В. Заповідні об’єкти Буковини
загальнодержавного значення як основа регіональної екологічної мережі // Заповідна
справа в Україні. – 2001. – Т. 7, вип. 2. – С. 73–98.
•
major dominants are Festuca valesiaca and Sipa capillata. Typical species are Euphorbia
seguierana, Herniaria kotovii, Artemisia austriaca. On stony soils, there dominates Sipa
graiicola and Thymus graniicus. On rocks, there are Erodium beketowii, Asperula graniicola,
Rumex fascilobus, Jurinea graniicola, Hedysarum grandilorum, Scophularia graniica.
Botanical significance. Important area for conservaion of the steppe vegetaion.
Kalmiuskyi Step
V.P. Kolomiychuk, V.A. Onyshchenko
Ukrainian name: Кальміуський степ
Transliteraion/Translaion variants: Kal’mius Steppe.
Area: 2994.0 ha.
Altitude: 40–130 m.
Latitude: 47°30’39” N (47.5107°).
Longitude: 37°55’33” E (37.9257°)
Administrative regions. Donetsk region: Boikivske (Telmanove) raion.
Ownership: state.
Biogeographic regions: steppic.
Habitats. Level 1. C – 0.8%; E – 96.0%; F – 0.5%; G – 1.7%; H – 1.0%.
Habitats. Level 2. C2 Surface running waters – 0.6%; C3 Littoral zone of inland surface
waterbodies – 0.1%; E1 Dry grasslands – 94.0%; E2 Mesic grasslands – 2.0%; F3 Temperate
and mediterranean-montane scrub – 0.5%; G1 Broadleaved deciduous woodland – 0.1%;
G3 Coniferous woodland – 1.6%; H2 Screes – 0.2%; H3 Inland cliffs, rock pavements and
outcrops – 0.6%; H5 Miscellaneous inland habitats with very sparse or no vegetation –
0.2%.
Futher habitat description. E1.1 Inland sand and rock with open vegetation; E1.2 Perennial
calcareous grassland and basic steppes; E2.5 Meadows of the steppe zone; G3.F Highly
artificial coniferous plantations; H2.5 Acid siliceous screes of warm exposures, H3.1 Acid
silicious inland clifs.
Land use: agriculture (animals) – major; nature conservaion and research – major.
Protected areas: overlaps (579.6 ha) with Ukrainskyi Stepovyi nature reserve (includes
division “Kalmiuske” of the reserve), overlaps (579.6 ha) with Emerald Site “Ukrainskyi
Stepovyi Nature Reserve”.
Threats: agricultural intensification/expansion (arable) – low, agricultural intensiicaion/
expansion (grazing) – low, extracion (minerals/quarries) – low.
General description. The valleys of the Kalmius river and its tributary with the steppe
vegetaion and granite outcrops. The steppe vegetaion occupies the largest area. Its
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Criterion C
• E1.2 Perennial calcareous grassland and basic steppes; area: 2500 ha; trend: stable;
area data quality: good; trend data quality: good
Literature
1. Коломійчук В.П., Остапко В.М., Яровий С.С. ПЗ Український степовий //
Фіторізноманіття заповідників і національних природних парків України. Ч. 1. Біосферні
заповідники. Природні заповідники / під ред. В.А. Онищенка і Т.Л. Андрієнко. – Київ:
Фітосоціоцентр, 2012. – С. 336–378.
2. Ткаченко В.С., Генов А.П. Флороценотична характеристика запропонованого
Кальміуського державного заказника // Укр. ботан. журн. – 1986. – 43, № 5. – С. 92–96.
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Kamiani Mohyly
V.A. Onyshchenko, V.P. Kolomiychuk
Ukrainian name: Кам’яні могили.
Transliteraion/Translaion variants: Kam’yani Mogyly, Rocky Hills.
Area: 438.0 ha.
Altitude: 123–206 m.
Latitude: 47°18’23” N (47.3065°).
Longitude: 37°04’48” E (37.0800°).
Administrative regions. Donetsk region: Volodarske raion. Zaporizhia region: Rozivka raion.
Ownership: state.
Biogeographic regions: steppic.
Habitats. Level 1. C – 0.4%; D – 0.4%; E – 83.7%; F – 5.0%; G – 0.5%; H – 10.0%.
Habitats. Level 2. C3 Littoral zone of inland surface waterbodies – 0.4%; D5 Sedge and
reedbeds, normally without free-standing water – 0.4%; E1 Dry grasslands – 81.1%; E2
Mesic grasslands – 2.6%; F3 Temperate and mediterranean-montane scrub – 5.0%; G1
Broadleaved deciduous woodland – 0.5%; H3 Inland cliffs, rock pavements and outcrops
– 2.0%.
Futher habitat description. E1.2 Perennial calcareous grassland and basic steppes; E2.5
Meadows of the steppe zone; F3.1 Temperate thickets and scrub; F3.2 Submediterranean
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deciduous thickets and brushes; H2.5 Acid siliceous screes of warm exposures, H3.1 Acid
silicious inland clifs; H3.5 Almost bare rock pavements, including limestone pavements.
Land use: agriculture (animals) – minor; nature conservaion and research – major.
Protected areas: includes division “Kamiani Mohyly “ of Ukrainskyi Stepovyi nature reserve
(389.2 ha), overlaps (389.2 ha) with Emerald Site “Ukrainskyi Stepovyi Nature Reserve”.
Threats: –.
General description. Protected area with the steppe vegetaion and hills with outcrops of
granite. Main dominants are Poa angusifolia and Elytrigia trichophora. Species of high
constancy are Achillea setacea, Artemisia absinthium, Euphorbia stepposa, Falcaria vulgaris,
Festuca valesiaca, Galatella villosa, Medicago falcata, Phlomoides tuberosa (Phlomis
tuberosa), Salvia nemorosa, Sipa capillata, Teucrium polium, Thalictrum minus, Verbascum
austriacum. On more stony solils, there are plant communiies dominates by Festuca
valesiaca, Sipa capillata, Sipa lessingiana with presence of Achillea setacea, Cota inctoria
(Anthemis inctoria), Artemisia austriaca, Bromopsis riparia, Centaurea trinervia, Filipendula
vulgaris, Galatella villosa, Hypericum perforatum, Marrubium praecox, Odonites luteus,
Potenilla argentea, Salvia nutans, Scabiosa ochroleuca, Securigera varia, Stachys recta,
Thymus dimorphus, Thymus marschallianus. Prevailng dominant of the shrub vegetaion is
Prunus stepposa. On granite rocks, there are Allium decipiens, Allium lavescens, Asplenium
septentrionale, Asplenium trichomanis, Aurinia saxailis, Centaurea pseodoleucolepis,
Festuca valesiaca, Oites hellmannii, Rosa subpygmaea, Spiraea hypericifolia.
Botanical significance. Two species have all their natural range inside this area: Achillea
glaberrima, Centaurea pseudoleucolepis. Important area for conservaion of steppe thickets
and siliceus outcrops.
Criterion A
• Achillea glaberrima Klokov; A(ii); abundance: frequent (8 000 000 individuals); trend:
stable; species data quality: good; trend data quality: good.
• Astragalus tanaiicus K. Koch.; A(i), A(ii); abundance: rare; trend: luctuaing; species
data quality: medium; trend data quality: poor.
• Centaurea pseudoleucolepis Kleopow.; A(i), A(ii); abundance: frequent (150 000
individuals); trend: decreasing; species data quality: good; trend data quality: good.
Criterion C
• F3.247 Ponto-Sarmatic deciduous thickets; area: 20 ha; trend: increasing; area data
quality: good; trend data quality: good.
• H3.1 Acid siliceous inland cliffs; area: 1 ha; trend: stable; area data quality: medium;
trend data quality: good. Note: On the site, there prevail horizontal granite outcrops
(about 30 ha) that are not meet the habitat type H3.1 including only verical outcrops.
Literature
1. Білик Г.І., Панова Л.С. Поновлення степової рослинності в заповіднику Кам’яні
Могили після припинення випасання // Укр. ботан. журн. – 1970. – 27, № 6. – С. 711–715.
2. Бойко М.Ф. Участь мохоподібних у формуванні фітоценозів заповідника “Кам’яні
Могили” // Укр. ботан. журн. – 1979. – 36, № 5. – С. 478–483.
3. Коломійчук В.П., Остапко В.М., Яровий С.С. ПЗ Український степовий //
Фіторізноманіття заповідників і національних природних парків України. Ч. 1. Біосферні
заповідники. Природні заповідники / під ред. В.А. Онищенка і Т.Л. Андрієнко. – Київ:
Фітосоціоцентр, 2012. – С. 336–378.
4. Лисенко Г.М. Моніторинг фітосистем заповідного степу “Кам’яні Могили” //
Чорноморський ботанічний журнал. – 2008. – 65, № 1 – С. 89–97.
5. Осычнюк В.В., Генов А.П., Генова Л.Ф. Флора Украинского степного заповедника
127
(аннотированный список сосудистых растений). – М., 1988. – 44 с.
6. Ткаченко В.С. Резерватні сукцесії та охоронний режим степової рослинності в
заповіднику “Кам’яні Могили” // Укр. ботан. журн. – 1992. – 49, № 6. – С. 18–22.
7. Ткаченко В.С., Генов А.П., Сіренко В.О. Саморозвиток фітосистем заповідного
степу “Кам’яні Могили” (Донецька область) // Український ботанічний журнал. – 2003.
– 60, № 3. – С. 248–255.
8. Ткаченко В.С., Дідух Я.П., Генов А.П. та ін. Український природний степовий
заповідник. Рослинний світ. – К.: Фітосоціоцентр, 1998. – 280 с.
Polygonum hydropiper, Rubus caesius, Symphytum oicinale. Main dominants of shrub
communiies are Amorpha fruicosa and Salix acuifolia. Psammophyic communiies
dominated by Koeleria glauca s.l., Festuca beckeri, Secale sylvestre, Calamagrosis epigeios,
Carex colchica, Sedum sexangulare and unmanaged mesic and wet grasslands dominated
by Alopecurus pratensis, Arrhenatherum elaius, Carex praecox, Elytrigia repens, Festuca
pratensis, Graiola oicinalis, Lycopus exaltatus, Lysimachia vulgaris, Lythrum salicaria,
Lythrum virgatum, Poa palustris occupy about 200 ha.
Kanivski Ostrovy
V.A. Onyshchenko
Ukrainian name: Канівські острови.
Transliteraion/Translaion variants: Kaniv Islands.
Area: 1136 ha.
Altitude: 81–89 m.
Latitude: 49°42’16” N (49.7045°).
Longitude: 31°55’14” E (31.5873°).
Administrative regions. Cherkasy region: Kaniv raion.
Ownership: state.
Biogeographic regions: continental.
Habitats. Level 1. C – 40%; D – 2%; E – 16%; F – 17%; G – 25%.
Habitats. Level 2. C1 Surface standing waters – 10%; C2 Surface running waters – 28%; C3
Littoral zone of inland surface waterbodies – 2%; D5 Sedge and reedbeds, normally without
free-standing water – 2%; E1 Dry grasslands – 5%; E3 Seasonally wet and wet grasslands –
8%; E5 Woodland fringes and clearings and tall forb stands – 3%; F9 Riverine and fen scrubs
– 17%; G1 Broadleaved deciduous woodland – 25%.
Futher habitat description. C1.2 Permanent mesotrophic lakes, ponds and pools; C1.6
Temporary lakes, ponds and pools; C2.3 Permanent non-tidal, smooth-flowing watercourses;
C3.2 Water-fringing reedbeds and tall helophytes other than canes; C3.4 Species-poor beds
of low-growing water-fringing or amphibious vegetation; C3.5 Periodically inundated shores
with pioneer and ephemeral vegetation; C3.6 Unvegetated or sparsely vegetated shores
with soft or mobile sediments; D5.1 Reedbeds normally without free-standing water; D5.2
Beds of large sedges normally without free-standing water; E1.9 Open non-Mediterranean
dry acid and neutral grassland, including inland dune grassland (× E1.1); E3.4 Moist or wet
eutrophic and mesotrophic grassland; E5.4 Moist or wet tall-herb and fern fringes and
meadows; F9.1 Riverine scrub; G1.1 Riparian and gallery woodland, with dominant Alnus,
Betula, Populus or Salix; G1.3 Mediterranean riparian woodland.
Land use: conservation and research – 44%.
Protected areas: overlaps (686 ha) with Kanivskyi nature reserve, overlaps (686 ha) with
Emerald Site “Kanivskyi Nature Reserve”.
Threats: abandonment/reduction of land management – low.
General description. Several loodplain islands of variable coniguraion in the valley of
the Dnipro. A big area is occupied by riverine woods and shrubs. Tree layer of the woods
is dominated by Populus alba, Populus nigra, Salix alba. Species of high constancy: Acer
negundo, Aristolochia clemaiis, Galium verum, Myosois palustris, Poa nemoralis,
128
Botanical significance. The area is noted for conservaion of comlexes of a big river (forests,
shrubs, litoral and aquaic vegetaion).
Criterion C
• C3.4 Species-poor beds of low-growing water-fringing or amphibious vegetation; area: 1
ha; trend: luctuaing; area data quality: poor; trend data quality: medium
129
•
C3.51 Euro-Siberian dwarf annual amphibious swards; area: 3 ha; trend: unknown; area
data quality: poor; trend data quality: poor.
• E5.4 Moist or wet tall-herb and fern fringes and meadows; area: 30 ha; trend: unknown;
area data quality: poor; trend data quality: poor.
• F9.1 Riverine scrub; area: 180 ha; trend: stable; area data quality: poor; trend data
quality: poor.
• G1.11 Riverine Salix woodland; area: 80 ha; trend: stable; area data quality: poor; trend
data quality: poor.
• G1.3 Mediterranean riparian woodland; area: 200 ha; trend: stable; area data quality:
medium; trend data quality: poor.
Literature
1. Шевчик В.Л. ПЗ Канівський // Фіторізноманіття заповідників і національних
природних парків України. Ч. 1. Біосферні заповідники. Природні заповідники / під
ред. В.А. Онищенка і Т.Л. Андрієнко. – Київ: Фітосоціоцентр, 2012. – С. 151–169.
2. Шевчик В.Л., Соломаха В.А. Синтаксономія рослинності островів Круглик та
Шелестів Канівського природного заповідника // Укр. фітоцен. зб., 1997. – Сер. А, Вип.
1. – С. 12–27.
3. Шевчик В.Л., Соломаха В.А., Войтюк Ю.О. Синтаксономія рослинності та список
флори Канівського природного заповідника // Укр. фітоцен. зб, 1996. – Сер B., Вип. 1. –
С. 1–119.
Brachypodium sylvaicum, Viola mirabilis, Lathyrus vernus, Hedera helix, Hepaica nobilis.
Botanical significance. One of the best populaions of Friillaria montana in Ukraine.
Kaplivka
I.I. Chorney, A.I. Tokaryuk, V.V. Budzhak
Ukrainian name: Каплівка.
Area: 32.8 ha.
Altitude: 192-232 m.
Latitude: 48°25’33” N (48.4258°).
Longitude: 26°33’16” E (26.5545°).
Administraive regions: Chernivtsi region: Khotyn raion.
Ownership: state.
Biogeographiczones: coninental.
Habitats. Level 1. G1 – 100.0%.
Habitats. Level 2. G1 Broadleaved deciduous woodland – 100.0 %;
Futher habitat description: G1.A Meso- and eutrophic Quercus, Carpinus, Fraxinus, Acer,
Tilia, Ulmus and related woodland.
Landuse: forestry – major.
Protected areas: –.
Threats: forestry (intensiied forest management) – medium.
General description. Broadleaved forest with dominance of Quercus robur and signiicant
admixture of Tilia cordata, Carpinus betulus, Fraxinus excelsior, Acer campeste, Acer
pseudoplatanus, Acer platanoides, Cerasus avium. The shrub layer consists of Acer tataricum,
Swida sanguinea (Cornus sanguinea), Corylusa avellana, Sambucus nigra, Euonymus
europaeus, Euonymus verrucosus, Viburnum lantana. Typical species of the herb layer are:
Isopyrum thalictroides, Ficaria verna, Corydalis cava, Corydalis solida, Arum besserianum,
Polygonatum hirtum, Pulmonaria obscura, Aegopodium podagraria, Asarum europaeum,
Criterion A
• Fritillaria montana Hoppe; А (ii); abundance: rare; trend: stable; species dataquality:
good; trend data quality: poor.
Conservaion proposals. Create a state botanical reserve and an Emerald Site.
Literature
1. Каземірська М.А., Чорней І.І. Вікова та просторова структури популяцій Fritillaria
montana Hoppe у Прут-Дністровському межиріччі // Біологічні системи. – 2010. – Т. 2.,
Вип. 2. – С. 62-66.
2. Каземірська М.А., Токарюк А.І., Чорней І.І. Насіннєва продуктивність Fritillaria
montana Hoppe (Liliaceae) в популяціях на північно-східній межі ареалу (середнє ПрутДністров’я) // Заповідна справа в Україні. – 2010. – Т. 16, вип. 2.. – С. 9-14.
3. Каземірська М.А., Чорней І.І. Fritillaria montanaHoppe (Liliaceae): географічна
характеристика, поширення в Україні // Науковий вісник Чернівецького університету.
Біологія (Біологічні системи). – 2010. – Т. 2, Вип. 3. – С. 63-68.
4. Каземірська М.А., Чорней І.І. Стан популяцій Friillaria montana Hoppe (Liliaceae)
на крайній північно-східній межі ареалу // Ботаніка та мікологія: проблеми і
перспективи на 2011-2020 роки (Матеріали Всеукраїнської наукової конференції, Київ,
6-8 квітня 2011 року) / Під. ред. І.О. Дудки та С.Я. Кондратюка. – К.: Інститут ботанікиім.
М.Г. Холодного, 2011. – С. 65-67.
5. Чорней І.І., Буджак В.В., Токарюк А.І. Сторінками Червоної книги України
(рослинний світ). Чернівецька область. – Чернівці: ДрукАрт, 2010. – 452 с.
130
131
Karadah
V.A. Onyshchenko
Ukrainian name: Карадаг.
Transliteraion/Translaion variants: Karadag, Kara Dag, Kara Dağ.
Area: 3001.0 ha.
Altitude: 0–577 m.
Latitude: 44°55’56” N (44.9323°).
Longitude: 35°13’25” E (35.2236°).
Administrative regions. Autonomous Republic of Crimea: Feodosia city; Territorial waters
of Ukraine.
Ownership: state.
Biogeographic regions: steppic.
Habitats. Level 1. A – 24.0%; B – 0.2%; E – 17.5%; F – 3.3%; G – 47.0%; H – 8.0%.
Habitats. Level 2. A1 Littoral rock and other hard substrata – 0.1%; A2 Littoral sediment –
0.1%; A3 Infralittoral rock and other hard substrata – 15.5%; A5 Sublittoral sediment – 8.3%;
132
B2 Coastal shingle – 0.1%; B3 Rock cliffs, ledges and shores, including the supralittoral –
0.1%; E1 Dry grasslands – 17.5%; F3 Temperate and mediterranean-montane scrub – 1.3%;
F7 Spiny Mediterranean heaths – 2.0%; F9 Riverine and fen scrubs – 0.0%; G1 Broadleaved
deciduous woodland – 36.4%; G3 Coniferous woodland – 10.6%; H2 Screes – 1.0%; H3
Inland clifs, rock pavements and outcrops – 6.9%; H5 Miscellaneous inland habitats with
very sparse or no vegetation – 0.1%.
Futher habitat description. A2.1 Litoral coarse sediment; A2.5 Coastal saltmarshes and
saline reedbeds; A3.2 Atlanic and Mediterranean moderate energy infralitoral rock; A3.2
Atlanic and Mediterranean low energy infralitoral rock; A5.1 Sublitoral coarse sediment;
A5.5 Sublitoral macrophyte-dominated sediment; B2.1 Shingle beach dritlines; B2.2
Unvegetated mobile shingle beaches above the dritline; B2.3 Upper shingle beaches
with open vegetaion; B3.1 Supralitoral rock (lichen or splash zone); B3.2 Unvegetated
rock clifs, ledges, shores and islets; B3.3 Rock clifs, ledges and shores, with angiosperms;
B3.4 Sot sea-clifs, oten vegetated; E1.1 Inland sand and rock with open vegetation; E1.2
Perennial calcareous grassland and basic steppes; E1.3 Mediterranean xeric grassland; F3.2
Submediterranean deciduous thickets and brushes; F7.4 Hedgehog-heaths; F9.3 Southern
riparian galleries and thickets; G1.7 Thermophilous deciduous woodland; G1.A1 Quercus –
Fraxinus – Carpinus betulus woodland on eutrophic and mesotrophic soils; G3.9 Coniferous
woodland dominated by Cupressaceae or Taxaceae; G3.F Highly ariicial coniferous forestry
plantations; G5.61 Deciduous shrub woodland; H2.6 Calcareous and ultra-basic screes of
warm exposures; H3.2 Basic and ultra-basic inland clifs; H5.3 Sparsely- or un-vegetated
habitats on mineral substrates not resulting from recent ice activity; H5.6 Trampled areas.
Land use: nature conservaion and research – 95%; tourism/recreaion – minor.
Protected areas: overlaps (2860 ha) with Karadazkyi nature reserve, overlaps (2860 ha)
with Emerald Site “Karadazkyi Nature Reserve”.
Threats: development (recreaion/tourism) – low, eutrophicaion – low.
General description. IPA is located in the eastern part of the Crimean mounains and
includes 809 ha of the Black Sea. A part of the area are typical of Crimea. Mountains of
this territory are composed of limestone. Closer to the sea, there prevail volcanic rocks
(tufs, liparites). The largest area (about 600 ha) is occupied by mesic Quercus pubescens
forests with the shrub layer dominated by Cornus mas. Aegonychon purpureo-caeruleum
prevails in the herb layer. Constant species are Acer campestre, Asparagus vericillatus,
Carex hallerana, Dictamnus gymnostylis, Laser trilobum, Mercurialis perennis, Paeonia
daurica, Physospermum cornubiense, Polygonatum hirtum. Besides there are about 200
ha of dry open Quercus pubescens woods with Paliurus spina-chrisi, Juniperus oxycedrus,
Pyrus eleagnifolia, Jasminum fruicans, Coinus coggygria, Rosa canina s.l., Festuca rupicola,
Festuca callieri, Elytrigia nodosa. At higer alitudes, there are Quercus petraea woods with
dominance or co-dominance of Carpinus betulus, Physospermum cornubiense, Mercurialis
perennis, Galanthus plicatus, Corydalis marschalliana and Fraxinus excelsior woods with
dominance of Physospermum cornubiense and Ranunculus constantinopolitanus in the
herb layer. Open woods of Juniperus excelsa occupy about 37 ha. Steppe vegetaion covers
about 400 ha. Typical dominants of the steppe vegetaion are Festuca valesiaca, Festuca
callieri, Sipa ponica; species of high constancy: Achillea nobilis, Aegilops triuncialis,
Botriochloa ischaemum, Centaurea difusa, Filipendula vulgaris, Galatella villosa,
Helianthemum salicifolium, Inula oculus-chrisi, Jurinea sordida, Koeleria cristata, Thesium
arvense, Teucrium polium, Veronica capsellicarpa, Velezia rigida. Steppes dominated
by Bromopsis cappadocica or Sipa brauneri occur on stony soils. Steppes dominated by
Elytrigia nodosa, Agropyron ponicum, A. pecinatum have high constacies of Alyssum
133
hirsutum, Alyssum umbellatum, Bromus squarrosus, Crupina vulgaris, Eryngium campestre,
Jurinea stoechadifolia, Poa sterilis. Ephemeral grasslands dominated by Aegilops triuncialis,
Anisantha sterilis, Anisantha tectorum occupy about 40 ha. In summer Cynodon dactylon
and Elytrigia repens dominate in these places. Major dominants of marine vegetaion on
sublitoral rock, boulders and pebble are Cladophora dalmaica, Cladostephus vericillatus,
Corallina mediterranea, Cystoseira crinita, Cystosera barbata, Phyllophora nervosa. At
oligotrophic sites at depth 1–5 m, there occur communiies co-dominated by Dilophus fasciola
f. repens, Polysiphonia opaca, Ceramium ciliatum, Enteromorpha compressa. Polysiphonia
elongata and Zanardinia prototypus dominate on sand at depth more than 15 m. Nemalion
helminthoides and Laurencia papillosa dominates on litoral rocks, Enteromorpha linza and
Ulva rigida – on stones at the water’s edge at eutrophic sites. Decreasing trend in biomass
of algae and in area of oligotrophic communiies exists because of eutrophicaion of the sea.
Botanical significance. This area is the richest in narrow endemic species part of Ukraine.
Flora of Karadah includes two narrow endemic species: Cerasium stevenii, Crataegus
pojarkovae. Important area for conservaion of deciduous woods, Juniperus excelsa
woodlands, dry grasslands, tomillares and marine vegetaion.
Criterion A
• Astracantha arnacantha (M.Bieb.) Podlech; A(iv); abundance: frequent, 5000 individuals;
trend: stable; species data quality: good; trend data quality: poor.
• Centaurea sarandinakiae Illar.; A(iv); abundance: occasional, >500 individuals; trend:
unknown; species data quality: good; trend data quality: poor.
• Cephalaria demetrii Bobrov; A(iii); abundance: rare, 150 individuals; trend: unknown;
species data quality: good; trend data quality: poor.
• Crambe aspera M. Bieb.; A(iv); abundance: rare, 100 individuals in 2 localities; trend:
unknown; species data quality: good; trend data quality: poor.
• Crambe koktebelica (Junge) N.Busch; A(ii); abundance: >100 individuals; trend:
unknown; species data quality: good; trend data quality: poor.
• Crambe pinnaiida W.T.Aiton.; A(iv); abundance: rare; trend: unknown; species data
quality: good; trend data quality: poor.
• Crambe steveniana Rupr.; A(iv); abundance: rare; trend: unknown; species data quality:
good; trend data quality: poor.
• Crataegus pojarkovae Kossych; A(iii); abundance: rare, 100 individuals; trend: stable;
species data quality: good; trend data quality: poor.
• Eremurus tauricus Steven; A(iv); abundance: occasional; trend: unknown; species data
quality: good; trend data quality: poor.
• Himantoglossum caprinum (Bieb.) C.Koch.; A(ii), A(iv); abundance: frequent, >8000
individuals; trend: unknown; species data quality: good; trend data quality: poor.
• Onobrychis pallasii (Willd.) M.Bieb.; A(iii); abundance: rare, 150 individuals; trend:
unknown; species data quality: good; trend data quality: poor.
• Onosma polyphylla Ledeb.; A(ii), A(iv); abundance: occasional, 3000 individuals; trend:
unknown; species data quality: good; trend data quality: poor.
• Ophrys oestrifera Bieb.; A(ii); abundance: rare, 120 individuals; trend: unknown; species
data quality: good; trend data quality: poor.
• Orchis punctulata Steven ex Lindl.; A(ii); abundance: rare, 40 individuals; trend:
unknown; species data quality: good; trend data quality: poor.
• Rhus coriaria L.; A(i); abundance: occasional; trend: stable; species data quality:
medium; trend data quality: medium.
• Sipa poëica Klokov; A(iv); abundance: occasional; trend: unknown; species data
quality: poor; trend data quality: poor.
Sipa syreistschikowii P.Smirn.; A(ii), A(iv); abundance: occasional; trend: unknown;
species data quality: medium; trend data quality: poor.
• Tilia dasystyla Stev.; A(iv); abundance: rare, 70 generaive individuals; trend: unknown;
species data quality: good; trend data quality: poor.
Criterion B
• G1 deciduous broadleaved woodland; area 1090 ha; % of indicator species: 30.0%; No
of indicator species: 20; trend: increasing; species data quality: good; area data quality:
good; trend data quality: medium.
Criterion C
• E1.2 Perennial calcareous grassland and basic steppes; area: 300 ha; trend: stable; area
data quality: medium; trend data quality: medium .
• F7 Spiny Mediterranean heaths; area: 40 ha; trend: stable; area data quality: poor;
trend data quality: medium.
• G3.9 Coniferous woodland dominated by Cupressaceae or Taxaceae; area: 37 ha; trend:
stable; area data quality: medium; trend data quality: medium.
• H3.2 Basic and ultra-basic inland cliffs; area: 200 ha; trend: stable; area data quality:
medium; trend data quality: medium.
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Karalarskyi Step
V.P. Kolomiychuk, V.A. Onyshchenko
Ukrainian name: Караларський степ.
Transliteraion/Translaion variants: Karalar Steppe.
Area: 10096.0 ha.
Altitude: 0–156 m.
Latitude: 45°27’07” N (45.4519°).
Longitude: 36°12’49” E (36.2135°).
Administrative regions. Autonomous Republic of Crimea: Lenine raion; Territorial waters
of Ukraine.
Ownership: state.
Biogeographic regions: steppic.
Habitats. Level 1. A – 2.1%; C – 7.8%; B – 0.4%; D – 0.1%; E – 84.4%; F – 0.2%; H – 4.0%;
I – 1.0%.
137
Habitats. Level 2. A1 Littoral rock and other hard substrata – 0.1%; A2 Littoral sediment –
0.1%; A3 Infralittoral rock and other hard substrata – 0.2%; A5 Sublittoral sediment – 1.7%;
B1 Coastal dunes and sandy shores – 0.1%; B2 Coastal shingle – 0.2%; B3 Rock clifs, ledges
and shores, including the supralittoral – 0.1%; D6 Inland saline and brackish marshes and
reedbeds – 0.1%; E1 Dry grasslands – 84.3%; E6 Salt steppes – 0.1%; F3 Temperate and
mediterranean-montane scrub – 0.2%; H2 Screes – 0.1%; H3 Inland clifs, rock pavements
and outcrops – 3.6%; H5 Miscellaneous inland habitats with very sparse or no vegetation –
0.3%; I1 Arable land and market gardens – 1.0%.
Futher habitat description. A2.1 Litoral coarse sediment; A3.2 Atlanic and Mediterranean
low energy infralitoral rock; A5.1 Sublitoral coarse sediment; A5.2 Sublitoral sand; A5.4
Sublitoral mixed sediments; B1.2 Sand beaches above the dritline; B2.2 Unvegetated
mobile shingle beaches above the dritline; B2.3 Upper shingle beaches with open
vegetaion; B3.1 Supralitoral rock (lichen or splash zone); B3.2 Unvegetated rock clifs,
ledges, shores and islets; B3.3 Rock clifs, ledges and shores, with angiosperms; B3.4 Sot
sea-clifs, oten vegetated; D6.1 Inland saltmarshes; E1.1 Inland sand and rock with open
vegetation; E1.2 Perennial calcareous grassland and basic steppes (80% of the IPA); E6.2
Continental inland salt steppes; F3.1 Temperate thickets and scrub; F3.2 Submediterranean
deciduous thickets and brushes; H2.6 Calcareous and ultra-basic screes of warm exposures;
H3.2 Basic and ultra-basic inland clifs; H5.3 Sparsely- or un-vegetated habitats on mineral
substrates not resulting from recent ice activity; H5.6 Trampled areas, I1.5 Bare illed,
fallow or recently abandoned arable land.
Land use: agriculture (animals) – minor, mowing/hay making – minor, nature conservaion
and research – major, tourism/recreaion – minor.
Protected areas: overlaps (about 6440 ha) with Karalarskyi regional landscape park, includes
“Ozero Chokrak” (1000 ha) regional hydrological reserve, overlaps (9885 ha) with Emerald
Site (“Karalarskyi”).
Threats: development (recreaion/tourism) – low.
General description. Major habitat type is the steppe. Dominant species are Sipa brauneri,
Sipa pulcherrima, Sipa capillata, Festuca rupicola, Festuca valesiaca, Festuca pseudovina,
Bromopsis cappadocica. The largest area is dominated by Sipa brauneri. Typical species
are Aegilops cylindrica, Cerastium tauricum, Koeleria cristata, Festuca rupicola, Galatella
villosa, Poa bulbosa. On the tops of the ridges, there is the steppe vegetaion dominated by
Sipa capillata and co-dominated by Artemisia taurica and Galatella villosa. On the gentle
slopes near the inlets, there are the sandy steppes. They are formed by Stipa boryshenica,
Carex colchica, Festuca beckeri, Jurinea longifolia. On the calcareous outcrops and screes,
there prevail Anisantha tectorum, Alyssum calycocarpum, Ephedra distachya, Euphorbia
petrophilla, Poa bulbosa, Pimpinella lithophila. On clay outcrops, there dominate Elytrigia
repens, Bromopsis cappadocica, Dactylis glomerata, Poa bulbosa, Medicago minima. On
calcareous rocks by the sea, there are Puccinellia distans, Kochia prostrata, Halimione
verrucifera, Holosteum umbellatum, Anisantha tectorum, Elytrigia bessarabica. Prevailing
species on the sand and shell beach are Leymus sabulosus, Cakile euxina, Crambe pontica.
Other typical species of this habitat are Euphorbia peplis, Lactuca tatarica, Pleconax
subconica, Salsola kali. subsp. pontica, Xanthium albinum. On the slopes of northern
aspects, there is the shrub vegetaion dominated by Crataegus dipyrena, С. monogyna, С.
taurica, Prunus spinosa, Rosa spinosissima, Sambucus nigra. There occur Swida australis
(Cornus australis), Ligustrum vulgare, Celis glabrata. The herb layer is composed of Arum
elongatum, Anthriscus longirostris, Cerasium tauricum, Galium aparine, Rumex euxinus.
The IPA includes salt Lake Chokrak (850 ha). The lake is shallow (average 0.85 m). In its
litoral, there are communiies of Salicornia perennans, Suaeda prostrata, Spergularia
media, Halocnemum strobilaceum, Halimione verrucifera.
Botanical significance. One of the largest areas of the steppe vegetaion in Ukraine.
Important for two threatned species of Crambe and seaside endemic species Thymus
litoralis.
Criterion A
• Crambe grandilora DC.; A(iv); abundance: rare; trend: stable; species data quality:
medium; trend data quality: poor.
• Crambe mitridais Juz.; A(iv); abundance: rare; trend: stable; species data quality:
medium; trend data quality: poor.
• Thymus litoralis Klokov et Shost.; A(iii); abundance: rare; trend: unknown; species data
quality: medium; trend data quality: poor.
Criterion C
• E1.2 Perennial calcareous grassland and basic steppes; area: 8000 ha; trend: stable;
area data quality: medium; trend data quality: medium
• F3.247 Ponto-Sarmatic deciduous thickets; area: 15 ha; trend: unknown; area data
quality: medium; trend data quality: poor.
Literature
1. Дідух Я.П., Шеляг-Сосонко Ю.Р. Ковиловий степ Чигини (Крим) // Укр. ботан.
журн. – 1980. – 37, № 4. – С. 79–84.
2. Коломійчук В.П. Флора Караларського регіонального ландшафтного парку (АР
Крим) // Укр. ботан. журн. – 2013. – 70, № 3. – С. 326–329.
3. Корженевский В. В., Клюкин А. А. Растительность абразионных и аккумулятивных
форм рельефа морских побережий и озер Крыма. – Ялта: ГНБС, 1990. – 108 c. – Деп. в
ВИНИТИ 10.07.90, № 3822-В90.
4. Котова И.Н. Флора и растительность Керченского полуострова // Тр. Никит. ботан.
сада. Материалы по флоре и растительности Крыма. – 1961. – Т. 35. – С. 64–168.
5. Новосад В.В. Флора Керченско-Таманского региона. – Киев: Наукова думка. –
138
139
1992. – 280 с.
6. Парнікоза І.Ю., Годлевська О.В., Зімнухов Р.О. Шляхи збереження унікальних
степових біотопів та археологічної спадщини у Караларському Приазов’ї // Заповідна
справа в Україні. – 2010. – Т. 16, вип. 2. – С. 99–104.
Kasova Hora
V.A. Onyshchenko, N.V. Shumska
Ukrainian name: Касова гора.
Transliteraion/Translaion variants: Kasova Gora, Kasova Mount, Kasova Hill.
Area: 149.7 ha.
Altitude: 250–335 m.
Latitude: 49°13’17” N (49.2214°).
Longitude: 24°42’18” E (24.7048°).
Administrative regions. Ivano-Frankivsk region: Halych raion.
Ownership: state, private.
Biogeographic regions: coninental
Habitats. Level 1. E – 96.0%; F – 3.8%; G – 0.2%.
Habitats. Level 2. E1 Dry grasslands – 96.0%; F3 Temperate and mediterranean-montane
scrub – 3.8%; G1 Broadleaved deciduous woodland – 0.2%.
Futher habitat descripion. E1.2 Perennial calcareous grassland and basic steppes; F3.1
Temperate thickets and scrub .
Land use: nature conservation and research – major.
Protected areas: overlaps (65 ha) with Halytskyi naional nature park, overlaps (65 ha) with
140
Emerald Site “Halytskyi Naional Nature Park”.
Threats: abandonment/reducion of land management – medium, agricultural
intensification/expansion (grazing) – low, burning of vegetaion – low, development
(recreaion/tourism) – low.
General description. A hill with the steppe vegetaion. Dominant species are Brachypodium
pinnatum, Carex humilis, Elyigia intermedia, Festuca valesiaca, Sipa capillata, Salvia
pratensis, Trifolium montanum, Teucrium chamaedrys. Other typical species are Adonis
vernalis, Anemone sylvestris, Anthericum ramosum, Asperula cynanchica, Aster amellus,
Bupleurum falcatum, Campanula glomerata, Campanula sibirica, Centaurea scabiosa,
Chamaecyisus ruthenicus, Filipendula vulgaris, Galium verum, Geranium sanguineum, Inula
ensifolia, Inula hirta, Peucedanum cervaria, Prunella grandilora, Salvia pratensis, Salvia
vericillata, Thalictrum minus, Veronica spicata. There are gypsum outcrops. A part of the
area belongs to the zone of strict protecion of naional park “Halytskyi”.
Botanical significance. The largest meadow steppe in Western Ukraine. Important for
Carlina cirsioides, Pulsailla grandis and other species of meadow steppes.
Criterion A
• Carlina cirsioides Klokov; A(iv); abundance: occasional (967 individuals); trend:
decreasing; species data quality: good; trend data quality: poor.
• Pulsailla grandis Wend.; A(ii); abundance: frequent (10 000 individuals); trend: stable;
species data quality: good; trend data quality: poor.
Criterion C
• E1.2 Perennial calcareous grassland and basic steppes; area: 140 ha; trend: stable; area
data quality: good; trend data quality: medium.
Literature
1. Куковиця Г.С. Найбільша ділянка ковилового степу на Поділлі // Укр. ботан. журн.
– 1970. – 27, № 1. – С. 111–113.
2. Куковиця Г.С., Мовчан Я.І., Соломаха В.А., Шеляг-Сосонко Ю.Р. Синтаксономія
лучних степів Західного Поділля // Укр. ботан. журн. – 1994. – 51, № 2–3. – С. 35–48.
3. Куковиця Г.С., Дідух Я.П., Шеляг-Сосонко Ю.Р., Абдулоєва О.С. Синтаксономія
лучних степів пам’яток природи республіканського значення гг. Касова та Чортова //
Укр. фітоцен. збірн. Серія А. Фітосоціологія. – 1998. – Вип. 2 (11). – С. 42–61.
4. Скоропляс І.О. Сучасний стан популяцій Carlina cirsioides га горі Касова // Вісник
Дніпропетровського державного аграрно-економічного університету. – 2014. – 33,
№ 1. – С. 143–145.
5. Чуй О.В., Шумська Н.В. Поширення видів роду Pulsatilla Mill. у Галицькому
національному природному парку та на прилеглих територіях // Рослинний світ у
Червоній книзі України: впровадження глобальної стратегії збереження рослин.
Матер. Міжнар. наук. конф. (м. Київ, 11–15 жовтня 2010 р.). – К.: Альтерпрес, 2010. –
С. 205–209.
6. Шеляг-Сосонко Ю.Р., Дідух Я.П., Єременко Л.П. та ін. Рослинність Касової гори
(Опілля) // Укр. ботан. журн. – 1981. – 38, № 3. – С. 60–66.
7. Шумська Н.В., Онищенко В.А., Маланюк В.Б. НПП Галицький // Фіторізноманіття
заповідників і національних природних парків України. Ч. 2. Національні природні
парки / під ред. В.А. Онищенка і Т.Л. Андрієнко. – Київ: Фітосоціоцентр, 2012. –
С. 105–123.
141
Habitats. Level 1. A – 22.0%; B – 1.4%; E – 67.1%; H – 9.5%.
Habitats. Level 2. A1 Littoral rock and other hard substrata – 0.6%; A2 Littoral sediment
– 0.1%; A3 Infralittoral rock and other hard substrata – 8.0%; A5 Sublitoral sediment –
13.3%; B1 Coastal dunes and sandy shores – 0.1%; B2 Coastal shingle – 0.1%; B3 Rock clifs,
ledges and shores, including the supralittoral – 1.2%; E1 Dry grasslands – 67.1%; H2 Screes
– 0.3%; H3 Inland clifs, rock pavements and outcrops – 9.0%; H5 Miscellaneous inland
habitats with very sparse or no vegetation – 0.2%.
Futher habitat description. A2.1 Litoral coarse sediment; A3.2 Atlanic and Mediterranean
low energy infralitoral rock; A5.1 Sublitoral coarse sediment; A5.2 Sublitoral sand; A5.4
Sublitoral mixed sediments; A5.5 Sublitoral macrophyte-dominated sediment; B1.2 Sand
beaches above the dritline; B2.2 Unvegetated mobile shingle beaches above the dritline;
B2.3 Upper shingle beaches with open vegetaion; B3.1 Supralitoral rock (lichen or splash
zone); B3.2 Unvegetated rock clifs, ledges, shores and islets; B3.3 Rock clifs, ledges and
shores, with angiosperms; B3.4 Sot sea-clifs, oten vegetated; E1.1 Inland sand and
rock with open vegetation; E1.2 Perennial calcareous grassland and basic steppes; H2.6
Calcareous and ultra-basic screes of warm exposures; H3.2 Basic and ultra-basic inland
clifs; H5.3 Sparsely- or un-vegetated habitats on mineral substrates not resulting from
recent ice activity; H5.6 Trampled areas.
Land use: nature conservaion and research – major.
Protected areas: includes Kazantypskyi nature reserve (450.1 ha), includes Ramsar Site
“Aquaic-clif complex of cape Kazantyp” (251 ha), overlaps (410 ha) with Emerald Site
“Kazantypskyi Nature Reserve”.
Threats: –.
General description. Major habitat type is the stony steppe. It is dominated by Bromopsis
cappadocica, Crinitaria villosa, Festuca pseudodalmaica, Festuca pseudovina, Festuca
rupicola, Koeleria cristata, Medicago falcata, Sipa capillata, Sipa lessingiana, Sipa
ucrainica, Teucrium polium. On the calcareous outcrops, there prevail Anisantha tectorum,
Alyssum calycocarpum, Asperula stevenii, Cerasium tauricum, Clypeola jonthlaspi, Ephedra
distachya, Euphorbia petrophila, Pimpinella lithophila, Poa bulbosa. On the supralitoral
rocks, there is halophytoc vegetaion with dominance of Puccinellia distans, Kochia
prostrata, Halimione verrucifera, Holosteum umbellatum, Anisantha tectorum, Elytrigia
bessarabica. Major species of the marine vegetaion are Cladophora albida, Cystoseira
barbata, Enteromorpha prolifera, Zannichellia major, Zostera nolii (Nanozostera nolii).
Botanical significance. Important area for some steppe species, in paricular Allium
pervesitum, Crambe grandilora, Crambe grandilora, Crambe mitridais, Crambe
pinnaiida. It harbours a small populaion of Ophrys oestrifera outside its main range. One
of the most important areas for conservaion of sea-clifs in Ukraine. The best site of the
marine vegetaion in the Sea of Azov.
Criterion A
• Agropyron cimmericum Nevski; A(i); abundance: occasional; trend: stable; species data
quality: medium; trend data quality: poor.
• Allium pervesitum Klokov.; A(i), A(iv); abundance: rare; trend: stable; species data
quality: medium; trend data quality: poor.
• Crambe grandilora DC.; A(iv); abundance: rare; trend: unknown; species data quality:
medium; trend data quality: poor.
• Crambe mitridais Juz.; A(iv); abundance: rare; trend: stable; species data quality:
medium; trend data quality: poor.
• Crambe pinnaiida W.T.Aiton.; A(iv); abundance: rare; trend: unknown; species data
quality: medium; trend data quality: poor.
• Ophrys oestrifera M. Bieb.; A(ii); abundance: rare; trend: unknown; species data quality:
medium; trend data quality: poor.
Criterion C
• B3.3 Rock cliffs, ledges and shores, with angiosperms; area: 2 ha; trend: stable; area
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Kazantyp
V.P. Kolomiychuk, V.A. Onyshchenko
Ukrainian name: Казантип.
Area: 522.5 ha.
Altitude: 0–107 m.
Latitude: 45°28’16” N (45.4711°).
Longitude: 35°50’36” E (35.8433°).
Administrative regions. Autonomous Republic of Crimea: Lenine raion; Territorial waters
of Ukraine.
Ownership: state.
Biogeographic regions: steppic.
data quality: poor; trend data quality: good.
H3.511 Limestone pavements; area: 60 ha; trend: stable; area data quality: medium;
trend data quality: good.
Literature
1. Исиков В.П., Корнилова Н.В. Казантипский природный заповедник // Сборник тр.
Никит. ботан. сада. – 2001. – С. 27–40.
2. Корженевський В.В., Квітницька О.А., Садогурський С.Ю. ПЗ Казантипський //
Фіторізноманіття заповідників і національних природних парків України. Ч. 1. Біосферні
заповідники. Природні заповідники / під ред. В.А. Онищенка і Т.Л. Андрієнко. – Київ:
Фітосоціоцентр, 2012. – С. 139–150.
3. Садогурская С.А., Садогурский С.Е., Белич Т.В. Аннотированный список
фитобентоса Казантипского природного заповедника Труды Никит. ботан. сада. – 2006.
– Т. 126. – С. 190–208.
4. Садогурский С.Е., Белич Т.В. Современное состояние макрофитобентоса
Казантипского природного заповедника (Азовское море) // Заповідна справа в Україні.
– 2003. – Т. 9, вип 1. – С. 10–15.
5. Ходосовцев А.Е. Аннотированный список лишайников Казантипского природного
заповедника // Тр. Никит. ботан. сада. – 2006. – Т. 126. – С. 216–221.
•
layer and presence of Betula pendula, Dryopteris dilatata, Fagus sylvaica, Luzula luzuloides,
Sorbus aucuparia. The moss layer consists of Polytrichum formosum, Pleurozium schreberi,
Dicranum scoparium, Bazzania trilobata, Leucobryum juniperoides, Sphagnum spp.
Botanical significance. Important area for Pinus cembra forests with Larix polonica.
Kedryn
R.Ya. Kish, V.A. Onyshchenko
Ukrainian name: Кедрин.
Area: 225.0 ha.
Altitude: 780–1464 m.
Latitude: 48°25’09” N (48.4191°).
Longitude: 24°01’07” E (24.0187°).
Administrative regions. Zakarpatska region: Tiachiv raion.
Ownership: state.
Biogeographic regions: alpine.
Habitats. Level 1. C – 0.2%; E – 0.3%; G – 98.6%; H – 0.9%.
Habitats. Level 2. C2 Surface running waters – 0.2%; E1 Dry grasslands – 0.3%; G3
Coniferous woodland – 95.6%; G4 Mixed deciduous and coniferous woodland – 3.0%; H2
Screes – 0.5%; H3 Inland cliffs, rock pavements and outcrops – 0.3%; H5 Miscellaneous
inland habitats with very sparse or no vegetation – 0.1%.
Futher habitat description. C2.1 Springs, spring brooks and geysers; C2.2 Permanent nontidal, fast, turbulent watercourses; E1.7 Non-Mediterranean dry acid and neutral closed
grassland; G3.1 Abies and Picea woodland; G4.6 Mixed Abies – Picea – Fagus woodland;
G5.6 Early-stage natural and semi-natural woodlands and regrowth; H2.3 Temperatemontane acid siliceous screes; H3.1 Acid silicious inland clifs; H5.6 Trampled areas.
Land use: forestry – minor, nature conservation and research – major.
Protected areas: includes Kedrynskyi state botanical reserve (166 ha).
Threats: climate change/sea level rise – low, forestry (intensiied forest management)) –
low.
General description. Picea abies (predominantly) and Pinus cembra forests with prevailing
of Vaccinium myrillus, Vaccinium viis-idaea, Calamagrosis villosa, Rubus hirtus in the herb
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Criterion A
• Larix decidua Mill. subsp. polonica; A(ii); abundance: frequent; trend: unknown; species
data quality: medium; trend data quality: poor.
Criterion C
• G3.25 Carpathian Larix and Pinus cembra forests; area: 28 ha; trend: decreasing; area
data quality: medium; trend data quality: medium.
Conservation proposals. Do not allow clear fellings including sanitary ones. Create an
Emerald Site. Expand Kedrynskyi state botanical reserve.
Literature
1.Природно-заповідний фонд Закарпатської област. – Ужгород: Карпати, 2011. – 256 с.
2.Стойко С.М., Третяк П.Р., Бойчук І.І. Сосна кедрова (Pinus cembra L.) на верхній межі
лісу у Ґорґанах: хорологія, екологія, фенологія // Науковий вісник ДЛГУ: Дослідження,
охорона та збагачення біорізноманіття. – Львів: Вид-во ЛДГУ, 1999. – Вип. 99. –
С. 173–179.
145
Biogeographic regions: steppic.
Habitats. Level 1. C – 0.7%; E – 97.2%; F – 2.0%; G – 0.1%.
Habitats. Level 2. C1 Surface standing waters – 0.1%; C2 Surface running waters – 0.1%;
C3 Littoral zone of inland surface waterbodies – 0.5%; E1 Dry grasslands – 96.0%; E2
Mesic grasslands – 1.2%; F3 Temperate and mediterranean-montane scrub – 2.0%; G1
Broadleaved deciduous woodland – 0.1%.
Futher habitat description. E1.2 Perennial calcareous grassland and basic steppes; E2.5
Meadows of the steppe zone; F3.2 Submediterranean deciduous thickets and brushes.
Land use: agriculture (animals) – minor; mowing/hay making – minor; nature conservaion
and research – major.
Protected areas: includes division “Khomutovskyi Step” of Ukrainskyi Stepovyi nature
reserve (1030.4 ha), overlaps (1030.4 ha) with Emerald Site “Ukrainskyi Stepovyi Nature
Reserve”.
Threats: abandonment/reducion of land management – low.
General description. The area is protected since 1926. Major vegetaion type is the
steppe. Dominant species are Poa angusifolia, Elytrigia repens, Elytrigia trichophora,
Inula germanica, Vicia tenuifolia, Thalictrum minus, Galatella rossica. Due to the small
biomass of herbivores the vegetaion is dominated by bunch grasses (Festuca valesiaca
s.l., Sipa capillata, Sipa lessingiana) occupies a lesser area. Typical species are Artemisia
austriaca, Artemisia ponica, Caragana frutex, Centaurea adpressa, Euphorbia stepposa,
Galium octonarium, Hypericum elegans, Limonium platyphyllum, Linaria vulgaris, Marrubium
praecox, Nepeta parvilora, Phlomis pungens, Salvia nutans, Securigera varia, Stachys transsilvanica. Major species of the shrub vegetaion are Rhamnus cathartica, Prunus stepposa,
Caragana frutex, Genista scythica.
Botanical significance. Important area for conservaion of the steppe vegetaion.
Criterion A
• Crambe tataria Sebeok.; A(ii); abundance: frequent; trend: unknown; species data
quality: medium; trend data quality: poor.
• Paeonia tenuifolia L.; A(ii); abundance: frequent; trend: stable; species data quality:
medium; trend data quality: medium.
Criterion C
• E1.2 Perennial calcareous grassland and basic steppes; area: 1000 ha; trend: stable;
area data quality: good; trend data quality: good.
Literature
1. Білик Г.І., Осичнюк В.В., Ткаченко В.С. Рослинний покрив Хомутовського степу
за даними крупномасштабного картування // Укр. ботан. журн. – 1975. – 32, № 6. –
С. 747–752.
2. Білик Г.І., Ткаченко В.С. Рослинний покрив абсолютно заповідної ділянки
Хомутовського степу // Укр. ботан. журн. – 1971. – 28, № 3. – С. 337–342.
3. Гелюта В. П., Генов А. П., Ткаченко В. С., Минтер Д. В. Заповедник “Хомутовская
степь”. План управления / Под редакцией В. П. Гелюты. – Киев: Академпериодика,
2002. – 40 с.
4. Гринь Ф.О. Заповідник Хомутовський степ // Укр. ботан. журн. – 1956. – 13, № 2.
– С. 15–30.
5. Коломійчук В.П., Остапко В.М., Яровий С.С. ПЗ Український степовий //
Фіторізноманіття заповідників і національних природних парків України. Ч. 1. Біосферні
заповідники. Природні заповідники / під ред. В.А. Онищенка і Т.Л. Андрієнко. – Київ:
Фітосоціоцентр, 2012. – С. 336–378.
6. Осичнюк В.В. Зміни рослинності заповідника Хомутовський степ за 40 років //
Укр. ботан. журн. – 1966. – 23, № 4. – С. 50–56.
7. Ткаченко В.С. Графічна модель автогенної сукцесії Хомутовського степу // Укр.
ботан. журн. – 1992. – 49, № 2. – С. 16–21.
8. Ткаченко В.С. Фітоценотичний моніторинг резерватних сукцесій в Українському
степовому природному заповіднику. – К.: Фітосоціоцентр, 2004. – 184 с.
9. Ткаченко В.С., Дідух Я.П., Генов А.П. та ін. Український природний степовий
заповідник. Рослинний світ. – К.: Фітосоціоцентр, 1998. – 280 с.
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Khomutovskyi Step
V.A. Onyshchenko, V.P. Kolomiychuk
Ukrainian name: Хомутовський степ.
Transliteraion/Translaion variants: Khomutovskyi Steppe, Homutovsky Steppe.
Area: 1059.0 ha.
Altitude: 19–71 m.
Latitude: 47°17’09” N (47.2857°).
Longitude: 38°11’02” E (38.1839°).
Administrative regions. Donetsk region: Novoazovsk raion.
Ownership: state.
Kniazhpilskyi Lis
Ukrainian name: Княжпільський ліс.
Transliteraion/Translaion variants: Knyazhpil’skyi Forest.
Area: 1032.0 ha.
Altitude: 150–312 m.
Latitude: 48°42’31” N (48.7087°).
Longitude: 26°47’15” E (26.7874°).
Administrative regions. Khmelnytskyi region: Kamianets-Podilskyi raion.
Ownership: state.
V.A. Onyshchenko
Tilia, Ulmus and related woodland; H5.6 Trampled areas.
Land use: forestry – major, nature conservaion and research – major
Protected areas: includes Kniazhpilskyi state botanical reserve (821 ha), included in Podilski
Tovtry naional nature park, included in Emerald Site “Podilski Tovtry Naional Nature Park”.
Threats: –.
General description. A wood with the dominance of Quercus robur and Carpinus betulus
in the tree layer. In the herb layer, there prevail Carex pilosa, Aegopodium podagraria,
Corydalis solida, Anemone ranunculoides.
Botanical significance. Important for endemic species Aconitum besserianum.
Criterion A
• Aconitum besserianum Andrz. ex Trautv.; A(iii); abundance: unknown; trend: unknown;
species data quality: poor; trend data quality: poor.
Literature
1. Червона книга України. Рослинний світ / за ред. Я.П. Дідуха. – К.: Глобалконсалтинг,
2009. – 900 с.
Kolodiazne
Biogeographic regions: coninental.
Habitats. Level 1. C – 0.5%; E – 0.8%; G – 97.7%; H – 1.0%.
Habitats. Level 2. C2 Surface running waters – 0.5%; E2 – Mesic grasslands – 0.8%; G1
Broadleaved deciduous woodland – 96.9%; G3 Coniferous woodland – 0.1%; G4 Mixed
deciduous and coniferous woodland – 0.7%; H5 Miscellaneous inland habitats with very
sparse or no vegetation – 1.0%.
Futher habitat description. G1.A Meso- and eutrophic Quercus, Carpinus, Fraxinus, Acer,
V.A. Onyshchenko, M.V. Banik, O.V. Bezrodnova, V.V. Rasevich
Ukrainian name: Колодязне.
Area: 25.7 ha.
Altitude: 119–181 m
Latitude: 50°00’57” N (50.0159°).
Longitude: 37°39’33” E (37.6592°).
Administrative regions. Kharkiv region: Dvorichna raion.
Ownership: state.
Biogeographic regions: steppic.
Habitats. Level 1. E – 58.0%; F – 2.0%; G – 33.0%; H – 7.0%.
Habitats. Level 2. E1 Dry grasslands – 58.0%; F3 Temperate and mediterranean-montane
scrub – 2.0%; G1 Broadleaved deciduous woodland – 33.0%; H2 Screes – 7.0%.
Futher habitat description. E1.1 Inland sand and rock with open vegetation; E1.2 Perennial
calcareous grassland and basic steppes; F3.2 Submediterranean deciduous thickets and
brushes; G1.7 Thermophilous deciduous woodland; G1.A Meso- and eutrophic Quercus,
Carpinus, Fraxinus, Acer, Tilia, Ulmus and related woodland; G1.C Highly artificial
broadleaved deciduous forestry plantations; H2.6 Calcareous and ultra-basic screes of
warm exposures .
Land use: agriculture (animals) – major; forestry – major.
Protected areas: –.
Threats: burning of vegetaion – low, extracion (minerals/quarries) – low, forestry
(aforestaion) – medium, natural events – low.
General description. Slopes of the valley of the Verkhnia Dvorichna river. The broadleaved
forest and the steppe vegetaion occupy the largest areas. Besides there are chalk outcrops
that are rich in endemic species.
Botanical significance. The largest populaion of Daphne sophia in Ukraine.
Criterion A
• Daphne sophia Kalen.; A(i); A(iv); abundance: 1500 individuals (70% of recorded
148
149
individuals in Ukraine); trend: unknown; species data quality: good; trend data quality:
poor.
Conservaion proposals: Create a state botanical reserve.
Literature
1. Banik M. Daphne’04. Final report. – Kharkiv, 2006. – 20 p. (http://www.rufford.org/
files/127.01.04%20Detailed%Final%20Report.pdf).
2. Банік М.В., Тверетинова В.В., Волкова Р.Є., Атемасова Т.А., Атемасов А.А.,
Брезгунова О.О., Влащенко А.С., Гончаров Г.Л., Коноваленко С.В., Скоробогатов
В.М., Скоробогатов Є.В., Целіщев О.Г. Нові місцезнаходження Daphne sophiae Kalen.,
(Thymeleaceae) в Україні // Укр. ботан. журн. – 2007. – 64, № 4. – С. 565–569.
3. Расевич В.В. Еколого-ценотичні особливості популяцій Daphne sophia Kalen. у
природній флорі України // Укр. ботан. журн. – 2008. – 65, № 1. – С. 90–103.
4. Червона книга України. Рослинний світ / за ред. Я.П. Дідуха. – К.: Глобалконсалтинг,
2009. – 900 с.
Kolivska Ulohovyna
V.A.Onyshchenko
Ukrainian name: Колтівська улоговина.
Area: 324.0 ha.
Altitude: 286–323 m.
150
Latitude: 49°51’20” N (49.8555°).
Longitude: 25°06’09” E (25.1026°).
Administrative regions. Lviv region: Zolochiv raion.
Ownership: state.
Biogeographic regions: coninental.
Habitats. Level 1. D – 48.0%; E – 40.0%; F – 2.0%; G – 10.0%.
Habitats. Level 2. D4 Base-rich fens and calcareous spring mires – 12.0%; D5 Sedge and
reedbeds, normally without free-standing water – 36.0%; E3 Seasonally wet and wet
grasslands – 39.0%; E5 Woodland fringes and clearings and tall forb stands – 1.0%; F9
Riverine and fen scrubs – 2.0%; G1 Broadleaved deciduous woodland – 10.0%.
Futher habitat descripion. D4.1 Rich fens, including eutrophic tall-herb fens and calcareous
flushes and soaks; D5.1 Reedbeds normally without free-standing water; D5.2 Beds of
large sedges normally without free-standing water; E2.2 Low and medium altitude hay
meadows; E3.4 Moist or wet eutrophic and mesotrophic grassland; E5.4 Moist or wet tallherb and fern fringes and meadows; F9.2 Salix carr and fen scrub.
Land use: mowing/hay making – minor, nature conservation and research – major.
Protected areas: same as Verkhniobuzkyi regional landscape reserve, overlaps (about 316
ha) with Pivnichne Podillia naional nature park and Emerald Site “Pivnichne Podillia”.
Threats: abandonment/reducion of land management – low.
General description.
Parially drained mire. It comprises eutrophic reed and sedge communiies, rich fens, wet
meadows. The most frequent dominants are Carex appropinquata, Phragmites australis,
Carex nigra, Carex lava, Deschampsia cespitosa, Molinia caerulea, Carex rostrata,
signiicant area is occupied by communiies of Schoenus ferrugineus and Carex davalliana.
151
In some places there dominate Agrosis stolonifera, Anthoxanthum odoratum, Briza media,
Dactylis glomerata, Cladium mariscus, Carex paniculata, Festuca arundinacea, Festuca
rubra, Filipendula ulmaria, Scirpus sylvaicus. Other frequent species are Eupatorium
cannabinum, Lysimachia vulgaris, Lythrum salicaria, Mentha arvensis, Potenilla erecta.
On low elevaions consising of calcareous sand, there are mesic grasslands dominated by
Molinia caerulea, Festuca ovina, Calluna vulgaris with Astrania major, Carlina cirsioides,
Dracocephalum ruyschiana, Filipendula vulgaris, Genthiane pneumonanthe, Leucanthemum
vulgare, Peucedanum oreoselinum, Pimpinella major, Polygonatum odoratum, Prunella
grandilora, Pulsailla patens, Serratula inctoria, Stachys oicinalis, Thalictrum minus,
Trifolium montanum, Trollius europaeus. Dominants of shrub vegetaion area Salix cinerea
and Betula humilis. The tree layer of woods is formed mainly by Betula pubescens.
Botanical significance. Important area for Cochlearia polonica, Dracocephalum ruyschiana,
Pinguicula bicolor.
Criterion A
• Cochlearia polonica Frohl.; A(i); A(ii); A(iii); abundance: about 300 individuals (100% of
recorded individuals in Ukraine); trend: luctuaing; species data quality: good; trend
data quality: good.
• Dracocephalum ruyschiana L.; A(ii); abundance: rare; trend: unknown; species data
quality: medium; trend data quality: poor.
• Pinguicula bicolor Woł.; A(iv); abundance: rare; trend: unknown; species data quality:
poor; trend data quality: poor.
Criterion C
• D4.1 Rich fens, including eutrophic tall-herb fens and calcareous flushes and soaks;
area: 40 ha; trend: stable; area data quality: poor; trend data quality: medium.
Literature
1. Кагало О.О. Фітосозологічна характеристика болотного масиву у верхів’ї
р. Західний Буг // Укр. ботан. журн. – 1990. – 47, № 1. – С. 80–84.
2. Червона книга України. Рослинний світ / за ред. Я.П. Дідуха. – К.: Глобалконсалтинг,
2009. – 900 с.
0.3%; E3 Seasonally wet and wet grasslands – 0.0%; F9 Riverine and fen scrubs – 2.0%; G1
Broadleaved deciduous woodland – 25.0%; G3 Coniferous woodland – 66.0%; G4 Mixed
deciduous and coniferous woodland – 3.2%; H5 Miscellaneous inland habitats with very
Koncha-Zaspivskyi Lis
V.A. Onyshchenko
Ukrainian name: Конча-Заспівський ліс.
Transliteraion/Translaion variants: Koncha-Zaspa Forest.
Area: 2376.0 ha.
Altitude: 93–123 m.
Latitude: 50°16’57” N (50.2826°).
Longitude: 30°34’51” E (30.5809°).
Administrative regions. Kyiv city: Holosiyivskyi raion.
Ownership: state.
Biogeographic regions: continental.
Habitats. Level 1. C – 1.7%; D – 0.5%; E – 0.6%; F – 2.0%; G – 94.2%; H – 1.0%.
Habitats. Level 2. C1 Surface standing waters – 0.9%; C2 Surface running waters – 0.1%;
C3 Littoral zone of inland surface waterbodies – 0.7%; D5 – sedge and reedbeds, normally
without free-standing water – 0.8%; E1 Dry grasslands – 0.3%; E2 Mesic grasslands –
152
sparse or no vegetation– 1%.
Futher habitat description. E1.1 Inland sand and rock with open vegetation, F9.2 Salix carr
and fen scrub, G1.2 Mixed riparian floodplain and gallery woodland, G1.4 Broadleaved
swamp woodland not on acid peat, G1.7 Thermophilous deciduous woodland, G1.A Mesoand eutrophic Quercus, Carpinus, Fraxinus, Acer, Tilia, Ulmus and related woodland, G1.B
Non-riverine Alnus woodland; G3.4 Pinus sylvestris woodland south of the taiga, G4.C
Mixed Pinus sylvestris – thermophilous Quercus woodland, H5.3 Sparsely- or un-vegetated
153
habitats on mineral substrates not resulting from recent ice activity, H5.6 Trampled areas.
Land use: forestry – minor, nature conservation and research – major, tourism/recreation
– major.
Protected areas: includes Lisnyky Botanical Reserve (1110 ha), included in Holosiivskyi
national nature park and Emerald Site “Holosiivskyi Naional Nature Park”;
Threats: development (recreation/tourism) – low, water (drainage) – medium (urbanization
and drainage of adjacent areas).
Building cottages on a former mire near the IPA.
General description. Forest area in the southern part of Kyiv. The largest porion of this
territory is the sand terrace above the floodplain of the Dnipro river with Pinus sylvestris,
rarely Quercus robur, forests. The herb layer is dominated by Calamagrostis epigeios,
Calamagrostis arundinacea, Festuca ovina, Poa angustifolia. Species of high constancy: Carex
ericetorum, Calamagrostis arundinacea, Calamagrostis epigeios, Chamaecytisus ruthenicus,
Convallaria majalis, Dicranum polysetum, Euphorbia cyparissias, Festuca ovina, Frangula
alnus, Galium verum, Melampyrum pratense, Melica nutans, Peucedanum oreoselinum,
Polygonatum odoratum, Pyrus communis, Pleurozium schreberi, Poa angustifolia, Rubus
saxatilis, Silene nutans, Sorbus aucuparia, Veronica officinalis. Non-forest vegetaion is
represented by sands with vegetaion dominated by Calamagrosis epigeios, Corynephorus
canescens, Festuca beckeri, Cladonia miis.
In the northwestern part of the site, there is a wide (2 km) valley of a small river. Main
vegetation here is mesic and wet forests dominated by Alnus glutinosa, Fraxinus excelsior,
Quercus robur, Acer campeste, Tilia cordata, Ulmus minor; Alnus glutinosa and Salix cinerea
swamps; eutrophic mires with dominance of Carex riparia, Carex acutiformis, Phragmites
australis.
Botanical significance. This area contains good examples of diferent forest types.
Criterion C
• G3.4232 Sarmatic steppe Pinus sylvestris forests; area: 1800 ha; trend: stable; area data
quality: medium; trend data quality: medium.
• G1.21 Riverine Fraxinus – Alnus woodland, wet at high but not at low water; area: 40 ha;
trend: stable; area data quality: poor; trend data quality: poor.
Conservaion proposals: Do not allow clearcuings including coninuous sanitary fellings.
Literature
1. Вірченко В.М. Матеріали до бріофлори НПП “Голосіївський” // Проблеми
відтворення і охорони біорізноманіття України. Мат-ли всеукр. наук.-практ. конф. (21–
22 квітня, 2011 р., м. Полтава). – Полтава: Астрая, 2011. – С. 86–88.
2. Дідух Я., Андрієнко Т., Серебряков В., Вольвач Ф., Філенко А., Лопарьов С.,
Чумак К. Заказник “Лісники” та його проблеми // Ойкумена. – 1994. № 1–2. – С. 116–
127.
3. Дідух Я.П., Чумак К.В., Геоботанічна характеристика заказника “Лісники” (м. Київ)
// Укр. ботан. журн. – 1992. – 49. № 3. – С. 22–27.
4. Онищенко В.А., Прядко О.І., Арап Р.Я. НПП “Голосіївський” // Фіторізноманіття
заповідників і національних природних парків України. Ч. 2. Національні природні
парки / Колектив авторів під ред. В.А. Онищенка і Т.Л. Андрієнко. – Київ: Фітосоціоцентр,
2012. – С. 139–151.
5. Прядко О.І., Арап Р.Я. Поширення та сучасний стан популяцій видів рослин
із Червоної книги України на території НПП “Голосіївський” // Мат-ли Міжнародної
наукової конференції “Рослинний світ у Червоній книзі України: впровадження
Глобальної стратегії збереження рослин”. – Київ, 2010. – С. 297–300.
Habitats. Level 2. E1 Dry grasslands – 94.4%; F3 Temperate and mediterranean-montane
scrub – 0.5%; G1 Broadleaved deciduous woodland – 2.0%; H2 Screes – 0.1%; H3 Inland
cliffs, rock pavements and outcrops – 0.9%; H5 Miscellaneous inland habitats with very
sparse or no vegetation – 2.1%.
Futher habitat description. E1.2 Perennial calcareous grassland and basic steppes; G1.C
154
155
Korsak Mohyla
V.A. Onyshchenko, V.P. Kolomiychuk
Ukrainian name: Корсак-Могила.
Transliteraion/Translaion variants: Korsak Hill.
Area: 72.5 ha.
Altitude: 87–138 m.
Latitude: 46°53’30” N (46.8917°).
Longitude: 36°08’06” E (36.1350°).
Administrative regions. Zaporizhia region: Prymorsk raion .
Ownership: state.
Biogeographic regions: steppic.
Habitats. Level 1. E – 94.4%; F – 0.5%; G – 2.0%; H – 3.1%.
Highly artificial broadleaved deciduous forestry plantations; H2.5 Acid siliceous screes
of warm exposures, H3.1 Acid silicious inland clifs; H3.5 Almost bare rock pavements,
including limestone pavements; H5.6 Trampled areas.
Land use: agriculture (animals) – minor; forestry – minor; nature conservaion and research
– major; tourism/recreaion – minor.
Protected areas: included in regional landscape reserve “Korsak-Mohyla”, included in
Emerald Site “Korsak Mohyla”.
Threats: abandonment/reducion of land management – low.
General description. Six hills with silicate outcrops. Major habitat type is the steppe
dominated by Festuca valesiaca s.l., Sipa capillata, Carex praecox, Caragana frutex, Poa
angusifolia, Elytrigia repens, E. intermedia, Bromopsis riparia. On the outcrops, there
prevail Achillea leptophylla, Carex praecox, С. stenophylla, Ephedra distachya, Teucrium
polium, Thymus dimorphus.
Botanical significance. Important area for Caragana scythica in the most transformed
region of Ukraine.
Criterion A
• Caragana scythica (Kom.) Pojark.; A(iv); abundance: frequent; trend: stable; species
data quality: good; trend data quality: medium.
Literature
1. Коломійчук В.П. Ботанічна характеристика Корсак-Могили (Запорізька область)
// Заповідна справа в Україні. – 2003. – Т. 9, № 1. – С. 1–2.
2. Коломійчук В.П. Psathyrosthachis juncea (Fisch.) Nevski в Северном Приазовье //
Вісник Запорізького національного університету: Зб. наук. статей. Фізико-математичні
науки. Біологічні науки. – Запоріжжя: ЗДУ. – 2005. – № 1. – С. 100–102.
3. Коломійчук В.П. Ключові ботанічні території Північного Приазов’я // Заповідна
справа в Україні. – 2008. – Т. 14, вип. 1. – С. 61–66.Тищенко О.В. Рослинність приморських
кіс північного узбережжя Азовського моря. – Київ: Фітосоціоцентр, 2006. – 156 с.
Kovalivka
O.M. Derkach, H.V. Kolomiets
Ukrainian name: Ковалівка.
Area: 19.9 ha.
Altitude: 5–12 m.
Latitude: 47°18’26” N (47.3071°).
Longitude: 31°43’01” E (31.7169°).
Administrative regions. Mykolaiv region: Mykolaiv raion.
Ownership: state.
Biogeographic regions: steppic.
Habitats. Level 1. E – 59.0%; G – 40.0%; H – 1.0%.
Habitats. Level 2. E1 Dry grasslands – 59.0%; G3 Coniferous woodland – 40.0%; H5
Miscellaneous inland habitats with very sparse or no vegetaion – 1.0%.
Futher habitat description. E1.9 Open non-Mediterranean dry acid and neutral grassland,
including inland dune grassland (× E1.1); G3.F Highly artificial coniferous plantations; H5.6
Trampled areas.
Land use: forestry – major.
156
Protected areas: included in regional preserve (zapovidne urochyshche) “Andriivske”
Threats: forestry (aforestaion) – medium.
General description. A part of sand terrace above the loodplain of the Southern Bug
river. Forest plantaions of Pinus pallasiana and Pinus sylvestris in comlex with dry sand
grasslands. Dominant species of the grasslands are Carex colchica, Artemisia marschalliana,
Festuca beckeri, Koeleria sabuletorum, Thymus pallasianus, Sipa borysthenica. Other
frequent species are Scabiosa ucrainica, Achillea ochroleuca, Helichrysum corymbiforme,
Astragalus varius, Anchusa gmelinii, Chamaecyisus borysthenicus, Allium savranicum,
Jurinea charcoviensis, Tragopogon borysthenicus, Alyssum savranicum. Density of
Centaurea margaritalba is 3 to 30 individuals per 100 m2.
Botanical significance. A locality of narrow endemic species Centaurea margarita-alba s. str.
Criterion A
• Centaurea margarita-alba Klokov; A(iii); abundance: frequent; trend: unknown; species
data quality: good; trend data quality: poor.
Conservaion proposals: Do not allow aforestaion. Restore sand grasslands in adjacent
area. Create a state botanical reserve.
Literature
1.Деркач О.М. Ключові ботанічні території Миколаївщини: сучасний стан та
проблеми збереження // Теорія і практика заповідної справи в Україні. Зб. наук. праць.
– К., 2005. – С. 167–173.
2.Деркач О.М. Андріївське // Південно-Бузький меридіональний екологічний
коридор: стислий огляд біорізноманіття та найцінніші території / під ред. В. Костюшина.
– К., 2007. – С. 49.
3.Коломієць Г.В. Перлинні волошки секції Pseudophalolepis Klok. ряду Margaritacea
Klok. Питання систематики та охорони // Укр. фітоценологічний зб. – Вип. 1–2 (12–13).
– К.: Фітосоціоцентр, 1999. – С. 165–169.
4. Крицька Л.І., Деркач О.М. Сучасний стан популяцій видів ряду Margaritaceae Klok.
157
(Centaurea L.) // Укр. ботан. журн. – 1991. – 48, № 3. – С. 78–80.
5.Крицька Л.І., Деркач О.М., Собко В.Г. Волошка білоперлинна Centaurea margaritaalba Klokov / Червона книга України. Рослинний світ / за ред. Я.П. Дідуха. – К.:
Глобалконсалтинг, 2009. – С. 305.
6.Перлини піщаної флори у пониззях Південного Бугу та Інгулу. Cерія: Збереження
біорізноманіття в Приморсько-степовому екокоридорі / Під ред. Г.В. Коломієць. – К.:
Громадська організація “Веселий Дельфін”, 2008. – 60 c.
typical species are Alyssum minutum, Artemisia marschalliana, Calamagrostis epigeios,
Centaurea breviceps, Cerastium schmalhausenii, Cerastium ucrainicum, Dianthus platyodon,
Erophila verna, Euphorbia seguieriana, Genista sibirica, Helychrysum corymbiforme,
Holosteum umbellatum, Inula sabuletorum, Jurinea laxa, Salix rosmarinifolia, Scabiosa
ucrainiaca, Jacobaea borysthenica (Senecio borysthenicus), Thymus borysthenicus,
Tragopogon borysthenicus, Veronica arvensis, Veronica dillenii. The layer of mosses and
lichens is formed by Tortula ruraliformis, Ceratodon purpureus, Cladonia foliacea, Cladonia
rangiformis, Cetraria aculeata, Neofuscelia pockornii. Dominants of mesic grasslands are
Agrostis gigantea, Calamagrostis epigeios, Cynodon dactylon, Festuca rupicola, Scirpoides
Kozachelaherska Arena
V.A. Onyshchenko
Ukrainian name: Козачелагерська арена.
Transliteraion/Translaion variants: Kozachi Lageri Sands.
Area: 18974.0 ha.
Altitude: 4–25 m.
Latitude: 46°36’07” N (46.6020°).
Longitude: 33°03’19” E (33.0552°).
Administrative regions. Kherson region: Oleshky (Tsiurupynsk) raion.
Ownership: state.
Biogeographic regions: steppic.
Habitats. Level 1. C – 0.02%; D – 0.07%; E – 25.0%; F – 1.0. G – 24.9%; H – 49.0%.
Habitats. Level 2. C1 Surface standing waters – 0.01%; C3 Littoral zone of inland surface
waterbodies – 0.01%; D5 Sedge and reedbeds normally without free-standing water –
0.06%; D6 Inland saline and brackish marshes and reedbeds – 0.01%; E1 Dry grasslands
– 12.5%; E2 Mesic grasslands – 10.0%; E3 Seasonally wet and wet grasslands – 2.5%; E6
Inland salt steppes – 0.01%; F3 Temperate and mediterranean-mountane scrub – 1.0%;
G1 Broadleaved deciduous woodland – 20.9%; G3 Coniferous woodland – 4.0%; H5
Miscellaneous inland habitats with very sparse or no vegetation – 49.0%.
Futher habitat description. E1.2 Perennial calcareous grassland and basic steppes, E1.9
Open non-Mediterranean dry acid and neutral grassland including inland dune grassland
(× E1.1), E2.5 Meadows of the steppe zone, E3.4 Moist or wet eutrophic and mesotrophic
grassland, F3.1 Temperate thickets and scrub; G1.9 Non-riverine woodland with Betula,
Populus tremula or Sorbus aucuparia, G3.F Highly artificial coniferous plantations, H5.3
Sparsely- or un-vegetated habitats on mineral substrates not resulting from recent ice
activity; H5.6 Trampled areas.
Land use: agriculture (animals) – minor, nature conservation and research – 35%; tourism/
recreation – minor.
Protected areas: overlaps (about 6700 ha) with Oleshkivski Pisky national nature park,
included in Emerald Site “Oleshkivski Pisky”.
Threats: forestry (afforestation) – medium.
General description. The largest expanse of unwooded sand in Ukraine. This is a part of
the Dnipro river terrace in the steppe zone. Sand dunes reach a height of 5 m. On a large
part of the area vegetation is sparse, the most typical species are Agropyron dasyanthum,
Chamaecytisus borysthenicus, Linaria dulcis, and non-native species Corynephorus
canescens. More dense sand vegetation (sand steppe) is dominated by Festuca beckeri,
Koeleria sabuletorum, Agropyron lavrenkoanum, Stipa borysthenica, Carex colchica. Its
158
holoschoenus. Waterlogged habitats are occupied by Carex acutiformis, Carex acuta, Carex
riparia, Eupatorium cannabinum, Iris pseudacorus, Juncus conglomeratus, Phragmites
australis. Forest vegetation is represented by small (10–2500 m2) natural groves of Betula
borysthenica, Populus tremula and small plantations of Pinus sylvestris and Pinus pallasiana.
The species composition of Betula borysthenica woods is heterogenous in respect of
159
requirements of species to soil moisture: Agrostis gigantea, Calamagrostis epigeios, Carex
elata, Frangula alnus, Genista sibirica, Hieracium umbellatum, Juncus conglomeratus,
Phragmites australis, Rhamnus cathartica, Rubus caesius, Salix rosmarinifolia, Solidago
virgaurea. Populus tremula woods harbour mainly hygrophilous species
Botanical significance. The largest area of unwooded sands in Europe with endemic species.
Criterion A
• Agropyron dasyanthum Ledeb.; A(i); abundance: frequent; trend: stable; species data
quality: medium; trend data quality: poor.
• Alyssum savranicum Andrz. (Odontarrhena savranica (Andrz. ex Besser) D.A.German);
A(iv); abundance: rare; trend: unknown; species data quality: medium; trend data
quality: poor.
• Centaurea breviceps Iljin; A(iii); abundance: frequent; trend: unknown; species data
quality: medium; trend data quality: poor.
Criterion C
• E1.9 Open non-Mediterranean dry acid and neutral grassland, including inland dune
grassland; area: 2000 ha; trend: stable; area data quality: medium; trend data quality:
good .
• X35 Inland sand dunes; area: 13500 ha; trend: stable; area data quality: medium; trend
data quality: good.
Literature
1. Котенко Т.И., Уманец О.Ю., Селюнина З.В. Природный комплекс Казачьелагерской
арены Нижнеднепровских песков и проблемы его сохранения. Сообщение 1. Общая
характеристика Казачьелагерской арены // Заповідна справа в Україні. – 1999. – Том 5.
Вип. 1. – С. 61–72.
2. Мойсієнко І.І. НПП Олешківські піски // Фіторізноманіття заповідників і
національних природних парків України. Ч. 2. Національні природні парки / під ред.
В.А. Онищенка і Т.Л. Андрієнко. – Київ: Фітосоціоцентр, 2012. – С. 357–372.
3. Соломаха І.В., Воробйов Є.О., Мойсієнко І.І. Рослинний покрив лісів та чагарників
Північного Причорномор’я. - К.: Фітосоціоцентр, 2015. – 387 с.
4. Уманец О.Ю. Природный комплекс Казачьелагерской арены Нижнеднепровских
песков и проблемы его сохранения. Сообщение 2. Изменение флоры и растительности
Казачьелагерской арены за 65 лет // Заповідна справа в Україні. – 1999. – Том 5. Вип.
2. – С. 9–15.
Habitats. Level 1. E – 50.0%; F – 2.0%; G – 40.0%; H – 8.0%.
Habitats. Level 2. E1 Dry grasslands – 50.0%; F3 Temperate and mediterranean-montane
scrub – 2.0%; G1 Broadleaved deciduous woodland – 14.0%; G3 Coniferous woodland –
20.0%; G4 Mixed deciduous and coniferous woodland – 6.0%; H2 Screes – 8.0%.
Futher habitat description. E1.1 Inland sand and rock with open vegetation; E1.2 Perennial
calcareous grassland and basic steppes; F3.2 Submediterranean deciduous thickets and
brushes; G1.7 Thermophilous deciduous woodland; G3.4 Pinus sylvestris woodland south
of the taiga; H2.6 Calcareous and ultra-basic screes of warm exposures; H5.3 Sparsely- or
un-vegetated habitats on mineral substrates not resulting from recent ice activity.
Land use: agriculture (animals) – 15%; nature conservaion and research – 85%.
Protected areas: overlaps (1134 ha) with Ukrainskyi Stepovyi nature reserve (same as its
Kreidiana Flora division), overlaps (1134 ha) with Emerald Site “Ukrainian Steppe Nature
Reserve”.
V.A. Onyshchenko
Ukrainian name: Крейдяна флора.
Transliteraion/Translaion variants: Chalk Flora.
Area: 1347.0 ha.
Altitude: 51–198 m.
Latitude: 48°52’07” N (48.8687°).
Longitude: 37°53’44” E (37.8956°).
Administrative regions. Donetsk region: Sloviansk raion, Lyman (Krasnyi Lyman) raion
Ownership: state.
Biogeographic regions: steppic.
Threats: abandonment/reducion of land management – low.
General description. Slopes with outcrops of chalk, tomillares, steppes, oak and pine
forests. Chalk with very sparse vegetaion occupies about 20 ha. Open vegetaion on chalk
occupies about 100 ha; dominants: Thymus cretaceus, Carex humilis, Festuca valesiaca,
Sipa capillata, Jurinea brachycephala, Artemisia salsoloides, Pimpinella itanophila,
Botriochloa ischaemum, Koeleria cristata, Koeleria talievii. Dominants of grasslands
(about 600 ha): Festuca valesiaca, Poa angusifolia, Sipa capillata; other typical species:
Bromopsis riparia, Calamagrosies epigeios, Carex praecox, Elytrigia repens, Euphorbia
stepposa, Medicago romanica, Salvia nutans. Prevailing species of shrub vegetaion (about
30 ha): Coinus coggygria, Crataegus curvisepala, Crataegus praearmata, Prunus spinosa,
Rhamnus catharica, Rosa corymbifera, Rosa tomentosa, Rosa villosa, Swida sanguinea
(Cornus sanguinea). Typical species of Pinus sylvestris woods (about 300 ha) are Quercus
robur, Swida sanguinea, Ligustrum vulgare, Coinus coggygria, Rosa canina, Chamaecyisus
160
161
Kreidiana Flora
ruthenicus, Euonymus verrucosus, Polygonatum odoratum, Securigera varia, Seseli libaniis
subsp. intermedia. Natural broadleaved forests occupies about 150 ha. Their tree layer is
dominated by Acer campestre, Acer platanoides, Fraxinus excelsior, Quercus robur, Tilia
cordata, Ulmus minor. In the herb layer, there dominates Aegonychon purpureo-coeruleum,
Dactylis glomerata, Elytrigia intermedia, Melica picta, Stellaria holostea. Area of broadleaved
forest is increasing.
Botanical significance. Important area for conservaion of chalk outcrops with endemic
endemic species and termophilous broad-leaved forests.
Criterion A
• Alyssum gymnopodum P.Smirn. (Odontarrhena gymnopoda (P.A.Smirn.) D.A.German);
A(iv); abundance: occasional; trend: stable; species data quality: medium; trend data
quality: poor.
• Diplotaxis cretacea Kotov; A(iv); abundance: occasional; trend: stable; species data
quality: medium; trend data quality: poor.
• Linaria cretacea Fisch. ex Spreng.; A(iv); abundance: rare; trend: stable; species data
quality: medium; trend data quality: poor.
• Silene cretacea Fisch. ex Spreng.; A(ii), A(iv); abundance: occasional; trend: stable;
species data quality: medium; trend data quality: poor.
• Sipa zalesskii Wilensky; A(ii); abundance: occasional; trend: stable; species data quality:
medium; trend data quality: poor.
• Syrenia talievi Klokov; A(iv); abundance: rare; trend: stable; species data quality:
medium; trend data quality: poor.
Criterion C
• E1.13 Coninental dry rocky steppic grasslands and dwarf scrub on chalk outcrops;
area: 200 ha; trend: increasing; area data quality: medium; trend data quality: good.
• F3.247 Ponto-Sarmatic deciduous thickets; area: 36 ha; trend: stable; area data quality:
poor; trend data quality: poor.
• G1.7 Thermophilous deciduous woodland (*91I0); area: 200 ha; trend: increasing; area
data quality: medium; trend data quality: good.
Literature
1. Ткаченко В.С., Генов А.П. Мовчан Я.І. Флористична характеристика Криволуцького
кретофільного степу на Донбасі та необхідність його заповідання // Укр. ботан. журн. –
1987. – 44, № 4. – С. 70–75.
2. Ткаченко В.С., Дідух Я.П., Генов А.П. та ін. Український природний степовий
заповідник. Рослинний світ. – К.: Фітосоціоцентр, 1998. – 280 с.
3. Червона книга України. Рослинний світ / за ред. Я.П. Дідуха. – К.: Глобалконсалтинг,
2009. – 900 с.
Kreidiani Vidslonennia
Ukrainian name: Крейдяні відслонення.
Transliteraion/Translaion variants: Chalk Outcrops.
Area: 43.8 ha.
Altitude: 90–110 m.
Latitude: 49°19’00” N (49.3167°).
162
V.A. Onyshchenko
Longitude: 39°49’55” E (39.8320°).
Administrative regions. Luhansk region: Milove raion.
Ownership: state.
Biogeographic regions: steppic.
Habitats. Level 1. E – 84.0%; H – 16.0%.
Habitats. Level 2. E1 Dry grasslands – 83.0%; E6 Salt steppes – 1.0%; H2 Screes – 16.0%.
Futher habitat description. E1.1 Inland sand and rock with open vegetation; E1.2 Perennial
calcareous grassland and basic steppes; E6.2 Continental inland salt steppes, H2.6
Calcareous and ultra-basic screes of warm exposures.
Land use: agriculture (animals) – major; nature conservaion and research – major.
Protected areas: includes Kreidiani Vidslonennia regional botanical reserve (30 ha), includes
Emerald Site “Kreidiani Vidslonennia” (30 ha).
Threats: agricultural intensification/expansion (arable) – low, extracion (minerals/quarries)
– low; forestry (aforestaion) – low.
General description. Slope of the valley of the Komyshna river with the steppe vegetaion
and outcrops of chalk. Major dominants of the steppe vegetaion are Festuca valesiaca
and Sipa capillata. On the tops of the hills, there are psammophyic steppes dominated
by Festuca beckeri and Cleistogenes squarrosa. Prevailing dominant on the chalk outcrops
is Thymus calcareus. Minor dominants are Artemisia hololeuca, Artemisia salsoloides,
Hyssopus cretaceus, Linaria cretacea, Onosma tanaiica, Pimpinella itanophila, Scrophularia
cretacea.
Botanical significance. Important area for conservaion of Hedysarum cretaceum and other
endemic species of chalk outcrops.
Criterion A
163
•
Hedysarum cretaceum J.Gay.; A(iv); abundance: occasional; trend: unknown; species
data quality: medium; trend data quality: poor.
Literature
1. Природно-заповідний фонд Луганської області // О.А. Арапов (заг. ред.), Т.B. Сова,
О.А. Cавенко, В.Б. Фєрєнц, Н.У. Кравець, Л.Л. Зятьков, Л.О. Морозова. Довідник. – 3-е
вид., доп. і перероб. – Луганськ: “Луганська правда”, 2013. – 224 с.
Kremenchutski Plavni
expansion (grazing) – low.
General description. The area includes parts of the loodplains of the Dnipro and Psel rivers.
A large area is covered by riverine Populus alba, Populus nigra, Salix alba woods, meadows
with dominance of Poa pratensis, Agrosis stolonifera, Elytrigia repens, psammophyic
grasslands dominated by Koeleria glauca s.l., Festuca beckeri, Calamagrosis epigerios,
Artemisia campestris s.l., loodplain Quercus robur woods and swamped Alnus gluinosa
woods with herb layer of Carex riparia and Carex acuiformis. Litoral communiies are
dominated by Typha angusifolia, Typha laifolia, Phragmites australis. Major dominants of
aquaic vegetaion are Certophyllum demersum, Ceratophyllum submersum, Lemna minor,
Myriophyllum spicatum, Potamogeton lucens, Potamogeton crispus, Hydrocharis morsusranae, Nuphar lutea, Trapa borysthenica.
V.A. Onyshchenko
Ukrainian name: Кременчуцькі плавні.
Transliteraion/Translaion variants: Kremenchuk Wetlands.
Area: 7231 ha.
Altitude: 64–70 m.
Latitude: 49°02’59” N (49.0496°).
Longitude: 33°26’49” E (33.4468°).
Administrative regions. Poltava region: Horishni Plavni (Komsomolsk) city, Kremenchuk city,
Kremenchuk raion.
Ownership: state.
Biogeographic regions: continental
Habitats. Level 1. C – 20%; D – 10%; E – 31%; F – 4%; G – 35%.
Habitats. Level 2. C1 Surface standing waters – 5%; C2 Surface running waters – 10%; C3
Littoral zone of inland surface waterbodies – 5%; D5 Sedge and reedbeds, normally without
free-standing water – 10%; E1 Dry grasslands – 1%; E2 Mesic grasslands – 15%; E3 Seasonally
wet and wet grasslands – 14%; E5 Woodland fringes and clearings and tall forb stands – 1%;
F9 Riverine and fen scrubs – 4%; G1 Broadleaved deciduous woodland – 34%; G3 Coniferous
woodland – 1%.
Futher habitat description. C1.2 Permanent mesotrophic lakes, ponds and pools;
C1.6 Temporary lakes, ponds and pools; C2.3 Permanent non-tidal, smooth-flowing
watercourses; C3.2 Water-fringing reedbeds and tall helophytes other than canes; C3.5
Periodically inundated shores with pioneer and ephemeral vegetation; C3.6 Unvegetated or
sparsely vegetated shores with soft or mobile sediments; D5.1 Reedbeds normally without
free-standing water; D5.2 Beds of large sedges normally without free-standing water; E1.9
Open non-Mediterranean dry acid and neutral grassland, including inland dune grassland (×
E1.1); E3.4 Moist or wet eutrophic and mesotrophic grassland; E5.4 Moist or wet tall-herb
and fern fringes and meadows; F9.1 Riverine scrub; G1.1 Riparian and gallery woodland,
with dominant Alnus, Betula, Populus or Salix; G1.2 Mixed riparian floodplain and gallery
woodland; G1.3 Mediterranean riparian woodland; G1.4 Broadleaved swamp woodland
not on acid peat.
Land use: agriculture (animals) – minor, conservation and research – major, forestry – 30%;
mowing / hay making – major, urban/industrial/transport – minor.
Protected areas: includes Biletskivski Plavni state landscape reserve (2980 ha), includes
Kremenchutski Plavni regional landscape park (5080 ha), includes Emerald Site
“Kremenchutski Plavni Regional Landscape Park” (5080 ha), overlaps with Emerald Site
“Dniprodzerzhynske Reservoir” (290 ha).
Threats: abandonment/reduction of land management – low; agricultural intensification/
Botanical significance. The area is noted for conservaion of comlexes of a big river (forests,
shrubs, litoral and aquaic vegetaion, meadows).
Criterion C
• E3.4 Moist or wet eutropic and mesotrophic grassland; area: 1000 ha; trend: stable;
area data quality: medium; trend data quality: medium.
• F9.1 Riverine scrub; area: 500 ha; trend: stable; area data quality: poor; trend data
quality: poor.
• G1.11 Riverine Salix woodland; area: 300 ha; trend: stable; area data quality: poor;
trend data quality: poor.
• G1.3 Mediterranean riparian woodland; area: 1800 ha; trend: stable; area data quality:
poor; trend data quality: poor.
Literature
1. Водно-болотні угіддя України. Довідник / Під ред. Марушевського Г.Б., Жарук І.С.
– К.: Чорноморська програма Ветландс Интернешнл, 2006. – 312 с.
2. Гальченко Н. П. Рослинність регіонального ландшафтного парку “Кременчуцькі
плавні” // Укр. ботан. журн. – 2004. – 61, № 4. – С. 48–55.
3. Гальченко Н.П. Регіональний ландшафтний парк “Кременчуцькі плавні” / Природнозаповідні території України. Рослинний світ. – Вип. 5. – К.: Фітосоціоцентр, 2006. – 176 с.
164
165
Habitats. Level 1. A – 0.1%; B – 0.0%; C – 0.0%; E – 13.0%; F – 3.0%; G – 82.9%; H – 1.0%;
J – 0.0%.
Habitats. Level 2. A1 Littoral rock and other hard substrata sediment – 0.0%; A2 Littoral
sediment – 0.1%; A3 Infralittoral rock and other hard substrata sediment – 0.1%; A5
Sublittoral sediment – 0.0%; B2 Coastal shingle – 0.0%; B3 Rock cliffs, ledges and shores,
including the supralittoral – 0.0%; C1 Surface standing waters – 0.0%; C2 Surface running
waters – 0.0%; C3 Littoral zone of inland surface waterbodies – 0.0%; E1 Dry grasslands –
11.0%; E2 – Mesic grasslands – 2.0%; F3 Temperate and mediterranean-montane scrub
– 2.0%; F5 – Maquis, arborescent matorral and thermo-Mediterraneal brushes; F6 –
Garrigues; F7 Spiny Mediterranean heaths – 1.0%; G1 Broadleaved deciduous woodland
– 77.3%; G3 Coniferous woodland – 3.0%; G4 Mixed deciduous and coniferous woodland
– 2.6%; H2 Screes – 0.1%; H3 Inland clifs, rock pavements and outcrops – 0.8%; H5
Miscellaneous inland habitats with very sparse or no vegetation – 0.1%.
Futher habitat description. A2.1 Litoral coarse sediment; A3.2 Atlanic and Mediterranean
moderate energy infralitoral rock; A3.2 Atlanic and Mediterranean low energy infralitoral
rock; A5.1 Sublitoral coarse sediment; A5.5 Sublitoral macrophyte-dominated sediment;
B2.1 Shingle beach dritlines; B2.2 Unvegetated mobile shingle beaches above the dritline;
B2.3 Upper shingle beaches with open vegetaion; B3.1 Supralitoral rock (lichen or splash
zone); B3.2 Unvegetated rock clifs, ledges, shores and islets; B3.3 Rock clifs, ledges and
shores, with angiosperms; B3.4 Sot sea-clifs, oten vegetated; E1.1 Inland sand and
rock with open vegetaion; E1.2 Perennial calcareous grassland and basic steppes; E1.3
Mediterranean xeric grassland; E2.5 Meadows of the steppe zone; F3.2 Submediterranean
deciduous thickets and brushes; F5.2 Maquis; F5.3 Pseudomaquis; F6.4 Black Sea garrigues;
F7.4 Hedgehog-heaths; G1.6 Fagus woodland; G1.7 Thermophilous deciduous woodland;
G1.A Meso- and eutrophic Quercus, Carpinus, Fraxinus, Acer, Tilia, Ulmus and related
woodland; G3.4 Pinus sylvestris woodland south of the taiga; G3.5 Pinus nigra woodland;
G3.7 Lowland to montane mediterranean Pinus woodland (excluding Pinus nigra); G3.F
Highly ariicial coniferous forestry plantaions; H2.6 Calcareous and ultra-basic screes of
warm exposures; H3.2 Basic and ultra-basic inland clifs; H5.3 Sparsely- or un-vegetated
habitats on mineral substrates not resuling from recent ice acivity; H5.6 Trampled areas.
Land use: agriculture (animals) – 3%; forestry – 65%; nature conservaion and research –
28%; tourism/recreaion – major.
Protected areas. Naional: overlaps (34563 ha) with Krymskyi nature reserve (includes
mountain part of the reserve), includes Yaltynskyi Hirsko-Lisovyi nature reserve (14523
ha), includes Aharmyskyi Lis state complex nature monument (40 ha), overlaps (17954 ha)
with Baydarskyi state landscape reserve, includes Dolhorukivska Yayla regional landscape
reserve (2130 ha), includes Hirskyi Karst Krymu state geological reserve (4316 ha), includes
Hirskyi Masyv Karaul-Oba state compex nature monument (100 ha), includes Hora
Kara-Tau state botanical nature monument (100 ha), includes Kachynskyi Kanyon state
geological reserve (100 ha), includes Karabi-Yaylynska Ulohovyna state botanical nature
monument (32 ha), includes Karstova Shakhta Soldatska state geological nature monument
(10 ha), includes Khapkhalskyi state hydrological reserve (250 ha), includes Kyzyl-Koba
state geological nature monument (33 ha), includes Manhup-Kale state complex nature
monument (90 ha), includes Mys Aia state landscape reserve (1340 ha), includes Novyi
Svit state botanical reserve (477 ha), includes Urochyshche Demerdzhi state geological
nature monument (20 ha), includes Urochyshche Karabi-Yayla state bonaical reserve (491
ha), includes Urochyshche Karasu-Bashi state hydrological nature monument (24 ha),
includes Urochyshche Kubalach state bonaical reserve (526 ha), includes Velyky Kanyon
Krymu state landscape reserve (300 ha). Regional: includes Ai-Petrynska Yayla regional
landscape reserve (1795 ha), includes Ai-Serez regional complex nature monument (5 ha),
includes Bakla regional geological nature monument (5 ha), includes Belbetskyi Tysovyi
Hay regional botanical reserve (20 ha), includes Demerdzhi Yayla regional landscape
reserve (2076 ha), includes regional reserve “Dolyna Richky Satera” (10 ha), includes
regional reserve “Hirsko-Lisovyi Masyv Bilia Sela Topolivka I Sela Kurske” (20 ha), includes
Hora Koshka regional complex nature monument (50 ha), includes Hora Krestova regional
complex nature monument (7 ha), includes regional botanical reserve Karabi-Yayla (2829
ha), includes Papaya-Kaya regional landscape reserve (550 ha), includes Hora Liagushka
regional complex nature monument (5 ha), includes regional geological nature monument
“Pryrodni Sinksy Karalezkoyi Dolyny” (5 ha), includes regional geological nature monument
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Krymski Hory
V.A. Onyshchenko
Ukrainian name: Кримські гори.
Transliteraion/Translaion variants: Crimean Mountains.
Area: 301150.0 ha.
Altitude: 0–1545 m.
Latitude: 44°45’07” N (44.7520°).
Longitude: 34°15’17” E (34.2548°).
Administrative regions. Autonomous Republic of Crimea: Alushta city, Bakhchysarai
raion, Bilohirsk raion, Feodosia city, Kirovske raion, Simferopol raion, Sudak city, Yalta city.
Sevastopol city: Balaklava raion. Territorial waters of Ukraine.
Ownership: state.
Biogeographic regions: steppic.
“Pryrodni Sinksy v Dolyni Richky Churuk-Su” (5 ha), includes Skeli Laspi regional preserve
(18 ha), includes Tepe-Kermen regional geological nature monument (5 ha), includes
Urochyshche Parahilmen regional botanical reserve (225 ha), includes regional landscape
park “Nauchnyi” (965 ha). Internaional: overlaps (22117 ha) with Emerald Site “Baidarskyi
ta Mys Aia”, overlaps (42597 ha) with Emerald Site “Bakhchysaraisko-Alushtynskyi”,
includes (130603 ha) Emerald Site “Bilohirskyi”, overlaps (34563 ha) with Emerald Site
“Crimean Nature Reserve”, overlaps (3651 ha) with Emerald Site “Sevastopolskyi”, includes
Emerald Site “Yaltynskyi Hirsko-Lisovyi Nature Reserve” (14523 ha), includes Emerald Site
“Gora Bila” (1091 ha).
Threats: burning of vegetaion – medium, development (recreaion/tourism) – low,
development (urbanizaion) – low, eutrophicaion – low.
General description. IPA includes major part of the Crimean mounains. Flora and vegetaion
are of Eastern submediterranean type with steppe elements. Forest vegetaion occupies
the largest area. Besides there are a lot of steppe vegetaion and large (>100 meters in
height and several kilometers in length) limestone clifs. Major forest type is mesic Quercus
petraea, Carpinus betulus and Fraxinus excelsior forests with Cornus mas in the shrub
layer and dominance of Mercurialis perennis, Physospermum cornubiense, Ranunculus
constaninopolitanus in the herb layer. At lower alitudes, there are Quercus pubescens
and Carpinus orientalis woods with high paricipaion of Aegonychon purpureo-caeruleum,
Carex hallerana, in drier habitats – open Quercus pubescens woods dominated by Elytrigia
nodosa. Fagus sylvaica ssp. moesiaca forests prevails at alitudes above 700 m. In their
herb layer, Dentaria glandulosa, Galium odoratum, Mercurialis perennis dominates. In
the lower belt, there occur Juniperus excelsa woodlands with paricipaion of Arbutus
andrachne, Cistus tauricus, Jasmnum fruicans, Juniperus oxycedrus, Quercus pubescens. In
the layer of herbs and low shrubs, there dominate of Achnatherum bromoides, Bromopsis
cappadocica, Elytrigia nodosa, Festuca callieri, Helianthemum stevenii, Poa sterilis, Thymus
callieri, Teucrium chamaedrys. On the southern slope of the main mountain chain, near
Yalta city, a considerable area is occupied by Pinus pallasiana (P. nigra ssp. pallasiana) and
Pinus sylvestris var. hamata (Pinus kochiana) forests with dominance of Brachypodium
rupestre, Carex humilis, Laser trilobum, Pteridium aquilinum. Besides there occur Acer
stevenii, Alnus gluinosa, Pinus bruia woods. Steppe vegetaion occupies large areas on the
tops of mountains (“Yayla”) and at low alitudes in outer parts of the mountains. Dominants
of the steppe vegetaion are Agropyron ponicum, Bromopsis cappadocica, Carex humilis,
Elytrigia nodosa, Festuca valesiaca, Sipa brauneri, Sipa lithophila, Sipa ponica, Sipa irsa,
Sipa pulcherrima. Mesic grasslands occur in medium and upper belts. Tomillares are in
all belts. Maximum of their distribuion is on Yayla. Dominaing species are Asphodeline
taurica, Helianthemum stevenii, Teucrium chamaedrys, Teucrium jailae, Thymus callieri,
Thymus hirsutus, Tymus tauricus.
Botanical significance. This area is important for a large number of species and habitats. For
some threatened species it includes all their extent of occurence.
Criterion A
• Astracantha arnacantha (M.Bieb.) Podlech; A(iv); abundance: frequent; trend:
decreasing; species data quality: good; trend data quality: poor.
• Astragalus setosulus Gontsch.; A(i), A(ii), A(iii); abundance: rare; trend: decreasing;
species data quality: medium; trend data quality: poor.
• Centaurea comperiana Steven; A(iii); abundance: occasional; trend: unknown; species
data quality: poor; trend data quality: poor.
• Centaurea sarandinakiae N.B.Illar.; A(iv); abundance: rare; trend: unknown; species
168
•
•
•
•
•
•
•
•
•
•
•
•
•
•
•
•
•
•
•
•
•
•
•
data quality: medium; trend data quality: poor.
Cephalaria demetrii Bobrov; A(iii); abundance: occasional; trend: unknown; species data
quality: medium; trend data quality: poor.
Chamaecyisus wulii (V.Krecz.) Klásková; A(iv); abundance: rare; trend: unknown;
species data quality: medium; trend data quality: poor.
Comperia comperiana (Steven) Asch. et Graebn.; A(ii); abundance: occasional; trend:
decreasing; species data quality: medium; trend data quality: poor.
Cyclamen kuznetzovii Kotov et Czernowa.; A(ii); A(iii); abundance: occasional
(>10 000 000 individuals, area of occupancy 400 ha); trend: stable; species data quality:
good; trend data quality: poor.
Cypripedium calceolus L.; A(ii); abundance: rare; trend: unknown; species data quality:
medium; trend data quality: poor.
Daphne taurica Kotov.; A(iii); abundance: 256 individuals (two localiies); trend: stable;
species data quality: good; trend data quality: medium.
Eremurus tauricus Steven; abundance: A(iv); occasional; trend: unknown; species data
quality: medium; trend data quality: poor.
Himantoglossum caprinum (Bieb.) C.Koch.; A(ii), A(iv); abundance: occasional; trend:
decreasing; species data quality: medium; trend data quality: medium.
Lagoseris purpurea (Willd.) Boiss.; A(i), A(ii), A(iii); abundance: rare; trend: unknown;
species data quality: medium; trend data quality: poor.
Medicago saxailis M.Bieb.; A(i); abundance: rare; trend: unknown; species data quality:
medium; trend data quality: poor.
Onobrychis pallasii (Willd.) M.Bieb.; A(iii); abundance: rare; trend: unknown; species
data quality: medium; trend data quality: poor.
Onosma polyphylla Ledeb.; A(ii), A(iv); abundance: occasional; trend: unknown; species
data quality: medium; trend data quality: poor.
Ophrys oestrifera Bieb.; A(ii); abundance: occasional; trend: decreasing; species data
quality: medium; trend data quality: poor.
Ophrys taurica (Aggeenko) Nevski Bieb.; A(ii); abundance: occasional; trend: decreasing;
species data quality: medium; trend data quality: poor.
Orchis provincialis Balb.; A(ii); abundance: rare; trend: unknown; species data quality:
medium; trend data quality: poor.
Orchis punctulata Steven ex Lindl.; A(ii); abundance: occasional; trend: unknown;
species data quality: medium; trend data quality: poor.
Paeonia tenuifolia L.; abundance: A(ii); frequent (1 000 000 individuals); trend: unknown;
species data quality: good; trend data quality: poor.
Pinus stankewiczii (Sukacz.) Fomin (P. bruia Ten. var. pityusa (Steven) Silba; A(i), A(iv);
abundance: occasional (three natural localiies with high number of individuals); trend:
decreasing; species data quality: good; trend data quality: medium.
Rhus coriaria L.; A(i); abundance: rare; trend: stable; species data quality: medium;
trend data quality: medium.
Silene jailensis N.I.Rubtzov; A(iii); abundance: 470 individuals (in 4 localiies); trend:
stable; species data quality: medium; trend data quality: poor.
Sisymbrium confertum Steven ex Turcz.; A(ii); abundance: rare; trend: unknown; species
data quality: poor; trend data quality: poor.
Steveniella satyrioides (Steven) Schlechter; A(ii); abundance: rare; trend: unknown;
species data quality: poor; trend data quality: poor.
Tilia dasystyla Stev.; A(iv); abundance: occasional; trend: stable; species data quality:
169
medium; trend data quality: poor.
Criterion B
• G1 Broadleaved deciduous woodland; area 240 000 ha; % of indicator species: 54.5%;
No of indicator species: 42; trend: stable; species data quality: good; area data quality:
good; trend data quality: good.
Criterion C
• B3.3 Rock cliffs, ledges and shores, with angiosperms; area: 1 ha; trend: stable; area
data quality: poor; trend data quality: medium.
• C2.12 Hard water springs; area: 0.1 ha; trend: stable; area data quality: poor; trend data
quality: poor.
• E1.2 Perennial calcareous grassland and basic steppes; area: 27 000 ha; trend: stable;
area data quality: medium; trend data quality: medium.
• F3.247 Ponto-Sarmatic deciduous thickets; area: 5000 ha; trend: stable; area data
quality: poor; trend data quality: medium.
• F7 Spiny Mediterranean heaths; area: 800 ha; trend: stable; area data quality: poor;
trend data quality: medium.
• G1.A4 Ravine and slope woodland; area: 100 ha; trend: stable; area data quality: poor;
trend data quality: poor.
• G1.6 Fagus woodland (excluding G1.66); area: 40000 ha; trend: stable; area data quality:
medium; trend data quality: medium.
• G1.7 Thermophilous deciduous woodland; area: 40000 ha; trend: stable; area data
quality: medium; trend data quality: medium.
• G3.4E Ponto-Caucasian Pinus sylvestris forests; area: 2000 ha; trend: decreasing; area
data quality: medium; trend data quality: medium.
• G3.5 Pinus nigra woodland; area: 9000 ha; trend: decreasing; area data quality:
medium; trend data quality: good.
• G3.7 Lowland to montainmediterranean Pinus woodland (excluding Pinus nigra); area:
200 ha; trend: decreasing; area data quality: medium; trend data quality: medium.
• G3.9 Coniferous woodland dominated by Cupressaceae or Taxaceae; area: 5000 ha;
trend: decreasing; area data quality: medium; trend data quality: good.
• H2.6 Calcareous and ultra-basic screes of warm exposures; area: 3000 ha; trend: stable;
area data quality: poor; trend data quality: medium.
• H3.2 Basic and ultra-basic inland cliffs; area: 15 000 ha; trend: stable; area data quality:
medium; trend data quality: good.
• H3.511 Limestone pavements; area: 5000 ha; trend: stable; area data quality: poor;
trend data quality: medium.
Literature
1. Didukh Ya.P. The communities of the class Quercetea pubescenti-petraeae at the
Crimean Mountains // Ukr. Phytosoc. Col. – Kyiv, 1996. – Ser. A., iss. 1. – P. 63–77.
2. Korzhenevsky V.V. Pinus pallasiana forest in the Crimea // Укр. фiтосоцiол. збiрник. –
Київ, 1998. – Сер. А. № 1 (9). – С. 78–97.
3. Дідух Я.П. Томіляри Гірського Криму // Укр. бот. журн. – 1981. – 38, № 4. – С. 84–89.
4. Дидух Я.П. Опыт класификации ксерофильной полукустарниковой и травянистой
растительности Горного Крыма // Ботан. журн., 1983. – 68, № 11. – С. 1456–1466.
5. Дидух Я.П. Растительный покров Горного Крыма (структура, динамика, эволюция
и охрана). – Киев: Наукова думка, 1992 – 256 с.
6. Дідух Я.П. Неморальні ліси Гірського Криму класу Querco-Fagetea Br.-Bl. еt. Vlieg.
1937 // Український фітоценологічний збірник, 1996. – Сер. А, №3. – С. 34–51.
7. Дідух Я.П. Гірські бори (Erico-Pinetea Horvat 1959) України // Рослинність хвойних
лісів України. – К., 2003. С. 43–79.
8. Дідух Я.П., Куземко А.А. Нові синтаксони класу Molinio-Arrhenatheretea з
Гірського Криму // Чорноморський ботан. журн. – 2009. – 5, № 4. – С. 547–562.
9. Дідух Я.П. ПЗ Ялтинський гірсько-лісовий // Фіторізноманіття заповідників
і національних природних парків України. Ч. 1. Біосферні заповідники. Природні
заповідники / під ред. В.А. Онищенка і Т.Л. Андрієнко. – Київ: Фітосоціоцентр, 2012. –
С. 390–405.
10. Зелена книга України / під ред. Я.П. Дідуха – К.: Альтерпрес, 2009. – 448 с.
11. Корженевский В.В. Сосняки из Pinus kochiana на Главной гряде Крымских
гор (синтаксономическое положение) // Классификация растительности СССР
(с использованием флористических критериев). М.: Изд-во Моск. ун-та, 1986. –
С. 102–112.
12. Корженевський В.В., Киселев О.А. Фитоценология восточно-буковых лесов
южного макросклона Главной гряды Крымских гор // Структура растительности и
биоэкология растений Крыма. – Ялта, 1982. – С. 26–35.
13. Корженевський В.В., Руденко М.І., Садогурський С.Ю. ПЗ Кримський //
Фіторізноманіття заповідників і національних природних парків України. Ч. 1. Біосферні
заповідники. Природні заповідники / під ред. В.А. Онищенка і Т.Л. Андрієнко. – Київ:
Фітосоціоцентр, 2012. – С. 198–220.
14. Партика Л.Я. Бріофлора Ялтинського гірсько-лісового природного заповідника
// Укр. ботан. журн. – 1995. – 52, № 2. – С. 260–270.
15. Ходосовцев О.Є., Богдан О.В. Анотований список лишайників Ялтинського
гірсько-лісового заповідника // Чорноморський ботан. журн. – 2005. – 1. – С. 117–132.
16. Рифф Л.Е. Продромус рослинності кам’янистих відслонень Гірського Криму
// Ю.Д. Клеопов та сучасна ботанічна наука. – К: Фітосоціоцентр, 2002. – С. 286–289.
17. Червона книга України. Рослинний світ / за ред. Я.П. Дідуха. – К.:
Глобалконсалтинг, 2009. – 900 с.
170
171
Kuchuhury
V.P. Kolomiychuk
Ukrainian name: Кучугури.
Transliteraion/Translaion variants: Kuchugury.
Area: 75.8 ha.
Altitude: 13–21 m.
Latitude: 47°32’47” N (47.5463°).
Longitude: 35°12’18” E (35.2049°).
Administrative regions. Zaporizhia region: Vasylivka raion.
Ownership: state.
Biogeographic regions: steppic.
Habitats. Level 1. C – 40.0%; E – 30.0%; F – 7.0%; G – 18.0%; H – 5.0%.
Habitats. Level 2. C1 Surface standing waters – 30.0%; C3 Littoral zone of inland surface
waterbodies – 10.0%; E1 Dry grasslands – 30.0%; F3 Temperate and mediterraneanmontane scrub – 3.0%; F9 Riverine and fen scrubs – 4.0%; G1 Broadleaved deciduous
woodland – 18.0%; H5 Miscellaneous inland habitats with very sparse or no vegetation
– 5.0%.
Futher habitat description. E1.9 Open non-Mediterranean dry acid and neutral grassland
including inland dune grassland (× E1.1); F3.1 Temperate thickets and scrub; F9.1 Riverine
scrub; G1.1 Riparian and gallery woodland, with dominant Alnus, Betula, Populus or Salix;
H5.3 Sparsely- or un-vegetated habitats on mineral substrates not resulting from recent
ice activity.
Land use: nature conservaion and research – 100%.
Protected areas: included in Velyky Luh naional nature park (zone of strict protecion);
included in Emerald Site “Velykyi Luh Naional Nature Park”.
Botanical significance. one of two known locality of narrow endemic species Centaurea
konkae.
Criterion A
• Centaurea konkae Klokov; A(iii); abundance: occasional (900 individuals); trend:
unknown; species data quality: good; trend data quality: poor.
Literature
1. Коломійчук В.П. Сучасний список регіонально рідкісних судинних рослин
Запорізької області // Промышленная ботаника. – 2011. – Вып. 11. – С. 105‒111.
2. Коломійчук В.П., Гальченко Н.П. Національний природний парк Великий Луг
// Фіторізноманіття заповідників і національних природних парків України. Ч. 2.
Національні природні парки / Колектив авторів під ред. В.А. Онищенка і Т.Л. Андрієнко.
– Київ: Фітосоціоцентр, 2012. – С. 63–76.
3. Тарасов В.В. Флора Дніпропетровської та Запорізької областей. Судинні рослини.
– Д.: Вид-во ДНУ, 2005. – 276 с.
4. Червона книга України. Рослинний світ / за ред. Я.П. Дідуха. – К.: Глобалконсалтинг,
2009. – 900 с.
Kuchurhan
Threats: natural events – low (erosion of islands).
General description. Sand islands in the Khakovka resrvoir. Dry sand grasslands are
dominated by Artemisia marschalliana and Secale sylvestre. Species of signiicant
constancy: Achillea micrantha, Agropyron dasyanthum, Carex colchica, Chondrilla juncea,
Euphorbia seguieriana, Jurinea paczoskiana, Rumex acetosella, Linaria odora subsp. dulcis,
Jacobaea borysthenica, Seseli tortuosum. Total cover is 50–80%. Besides there are Salix alba
woods, Salix acuifolia scrub and fresh standing waters with aquaic vegetaion and litoral
Phragmites australis communiies.
172
V.A. Onyshchenko, O.M. Popova
Ukrainian name: Кучурган.
Transliteraion/Translaion variants: Kuchurgan.
Area: 1673 ha.
Altitude: 21–160 m.
Latitude: 46°57’29” N (46.9580°).
Longitude: 29°53’28” E (29.8909°).
Administrative regions. Odesa region: Rozdilna raion, Velyka Mykhailivka raion
Ownership: state.
Biogeographic regions: steppic.
Habitats. Level 1. E – 65.0%; F – 2.0%; G – 32.0%; H – 1.0%.
Habitats. Level 2. E1 Dry grasslands – 63.0%; E2 Mesic grasslands – 2.0%; F3 Temperate
and mediterranean-montane scrub – 2.0%; G1 Broadleaved deciduous woodland – 17.0%;
G3 Coniferous woodland – 15.0%; H5 Miscellaneous inland habitats with very sparse or no
vegetation – 1.0%.
Futher habitat description. E1.1 Inland sand and rock with open vegetation; E1.2
Perennial calcareous grassland and basic steppes; E2.5 Meadows of the steppe zone;
F3.2 Submediterranean deciduous thickets and brushes; G1.7 Thermophilous deciduous
woodland; G1.C Highly artificial broadleaved deciduous forestry plantations; G3.F Highly
artificial coniferous plantations; H5.6 Trampled areas.
Land use: agriculture (animals) – 70%; forestry – 24%; mowing/hay making – minor
Protected areas: same as Emerald Site “Kuchurhanskyi”.
Threats: forestry (aforestaion) – medium.
General description. Eroded slopes of the Kuchurgan valley and its right tributary. The
largest area is occupied by steppe vegetaion and forest plantaions. No detailed data on
vegetaion.
Botanical significance. Important for conservaion of Colchicum fominii (~Colchicum
173
Habitats. Level 1. E – 94.0%; F – 2.0%; G – 2.0%; H – 2.0%.
Habitats. Level 2. E1 Dry grasslands – 94.0%; F3 Temperate and mediterranean-montane
scrub – 2.0%; G1 Broadleaved deciduous woodland – 0.4%; G3 Coniferous woodland –
1.6%; H5 Miscellaneous inland habitats with very sparse or no vegetation – 2.0%.
Futher habitat description. E1.9 Open non-Mediterranean dry acid and neutral grassland
including inland dune grassland (× E1.1); F3.1 Temperate thickets and scrub; G3.F Highly
artificial coniferous plantations; H5.6 Trampled areas.
Land use: forestry – 100%.
Protected areas: included in Emerald Site “Dniprovske Reservoir”.
Threats: forestry (aforestaion) – high, burning of vegetaion – low.
General description. IPA is situaued on the sand terrace of the Dnipro river. Major vegenaion
type is the dry sand grasslands dominated by Festuca beckeri, Artemisia marschalliana,
Secale sylvestre with total cover 40–50%. Frequent species are Achillea micrantha,
Agropyron dasyanthum, Carex colchica, Chondrilla juncea, Euphorbia seguieriana, Jurinea
charkoviensis, Rumex acetosella, Linaria odora subsp. dulcis, Jacobaea borysthenica, Seseli
tortuosum. With lower constancies, there occur Astragalus varius, Anchusa gmelinii,
Chamaecyisus borysthenicus, Gypsophilla paniculata. Populaion of Centaurea konkae
occupies about 1 ha. It has several hundred individuals.
Botanical significance. One of two known populaion of narrow endemic species Centaurea
konkae.
arenarium) in Ukraine.
Criterion A
• Colchicum fominii Bordz.; A(ii), A (iv); abundance: occasional; trend: unknown; species
data quality: medium; trend data quality: poor.
Conservaion proposals: Do not allow ariicial aforestaion, restore natural vegetaion.
Kurylivka
Ukrainian name: Курилівка.
Area: 65.4 ha.
Altitude: 55–58 m.
Latitude: 48°33’24” N (48.5566°).
Longitude: 34°37’19” E (34.6220°).
Administrative regions. Dnipropetrovsk region: Petrykivka raion.
Ownership: state.
Biogeographic regions: steppic.
174
V.P. Kolomiychuk
Criterion A
• Centaurea konkae Klokov; A(iii); abundance: occasional; trend: unknown; species data
quality: good; trend data quality: poor.
Conservaion proposals: Create a state botanical reserve.
175
Literature
1. Тарасов В.В. Флора Дніпропетровської та Запорізької областей. Судинні рослини.
– Дніпропетровськ: Вид-во ДНУ, 2005. – 276 с.
2. Червона книга Дніпропетровської області. (Рослинний світ) / Під ред.
А.П. Травлєєва. – Дніпропетровськ: ВКК “Баланс-Клуб”, 2010. – 500 с.
3. Червона книга України. Рослинний світ / за ред. Я.П. Дідуха. – К.: Глобалконсалтинг,
2009. – 900 с.
persicifolia, Centaurea pseudophrygia, Clinopodium vulgare, Maianthemum bifolium,
Meliis sarmaica, Polygonatum odaratum, Potenilla alba, Primula veris, Pulsatilla
patens, Serratula coronata. In the herb layer of Alnus gluinosa woods, there dominate
Carex vesicaria, Scirpus sylvaicus. Constant species are Carex acuiformis, C. elongata, С.
rostrata, Galium uliginosum, Juncus conglomeratus, Lysimachia vulgaris, Lythrum salicaria,
Padus racemosa, Peucedanum palustre, Poa palustris. Prevailing species of meadows are
Arrhenatherum elaius, Carex panicea, Cynosurus cristatus, Festuca rubra.
Levkivski Lisy
O.O. Orlov
Ukrainian name: Левківські ліси.
Transliteraion/Translaion variants: Levkivsky Forest.
Area: 973.0 ha.
Altitude: 176–201 m.
Latitude: 50°13’36” N (50.2265°).
Longitude: 28°53’56” E (28.8987°).
Administrative regions. Zhytomyr region: Zhytomyr raion.
Ownership: state.
Biogeographic regions: coninental.
Habitats. Level 1. C – 0.3%; E – 0.2%; G – 98.5%; H – 1.0%.
Habitats. Level 2. C1 Surface standing waters – 0.2%; C3 Littoral zone of inland surface
waterbodies – 0.1%; E2 Mesic grasslands – 0.2%; G1 Broadleaved deciduous woodland
– 9.5%; G3 Coniferous woodland – 14.5%; G4 Mixed deciduous and coniferous woodland
– 60.0%; G5 Lines of trees, small anthropogenic woodlands, recently felled woodland, earlystage woodland and coppice – 14.5%; H5 Miscellaneous inland habitats with very sparse or
no vegetation – 1.0%.
Futher habitat description. E2.2 Low and medium alitude hay meadows; G1.7
Thermophilous deciduous woodland; G1.9 Non-riverine woodland with Betula, Populus
tremula or Sorbus aucuparia; G3.4 Pinus sylvestris woodland south of the taiga; G4.7 Mixed
Pinus sylvestris – acidophilous Quercus woodland; G5.6 Early-stage natural and semi-natural
woodlands and regrowth; H5.6 Trampled areas.
Land use: forestry – major.
Protected areas: included in Emerald Site “Korostyshivskyi”.
Threats: forestry (intensiied forest management) – medium.
General description. Vegetaion is represented mainly by forests (deciduous, mixed and
coniferous approximately in equal parts). Mesic Pinus sylvestris forests are dominated
by Peucedanum oreoselinum, Dicranum polysetum, Pleurozium schreberi. Other frequnt
species are Campanula rotundifolia, Chamaecyisus ruthenicus, Dianthus pseudosquarrosus,
Gypsophila fasigiata, Luzula pilosa, Melampyrum pratense, Pulsatilla patens, Rumex
acetosella, Thymus serpyllum, Vaccinium viis-idaea. In weter pine forests, dominant
species of the lower layers are Frangula alnus, Vaccinium myrillus, Molinia caerulea,
Pteridium aquilinum, Dicranum polysetum, Pleurozium schreberi. Secondary Betula pendula
woods have species composiion similar to one of pine woods. Dominants of the herb layer
of Quercus robur woods and mixed Pinus sylvestris - Quercus robur woods are Convallaria
majalis and Pteridium aquilinum. Their constant species are Betonica oicinalis, Campanula
176
Botanical significance. Large dense populaion of Pulsailla patens. One of the largest
known localiies of endemic species Dianthus pseudoseroinus. A good example of species
rich oak woods.
Criterion A
• Dianthus pseudoseroinus Błocki; A(ii); abundance: occasional; trend: stable; speciesdata quality: medium; trend data quality: poor.
• Pulsailla patens (L.) Mill.; A(ii); abundance: frequent; trend: stable; species data quality:
good; trend data quality: medium.
Criterion C
• G1.7 Thermophilous deciduous woodland; area: 40 ha; trend: decreasing; area data
quality: medium; trend data quality: medium.
Conservaion proposals. Create a state botanical reserve including the enire IPA.
177
Lysa Hora i Sinka
Ukrainian name: Лиса гора і Стінка.
Transliteraion/Translaion variants: Bald Hill and Clif.
Area: 351 ha.
Altitude: 288–427 m.
Latitude: 49°47’00” N (49.7832°).
Longitude: 24°44’21” E (24.7390°).
Administrative regions. Lviv region: Zolochiv raion.
Ownership: state.
Biogeographic regions: coninental
178
V.A. Onyshchenko, T.D. Solomakha
Habitats. Level 1. E – 36.8%; F – 1.0%; G – 60.5%; H – 1.7%
Habitats. Level 2. E1 Dry grasslands – 36.2%; E5 Woodland fringes and clearings and tall forb
stands – 0.3%; E7 Sparsely wooded grasslands – 0.3%; F3 Temperate and mediterraneanmontane scrub – 1.0%; G1 Broadleaved deciduous woodland – 44.5%; G3 Coniferous
woodland – 12.0%; G4 Mixed deciduous and coniferous woodland – 1.5%; G5 Lines of
trees, small anthropogenic woodlands, recently felled woodland, early-stage woodland
and coppice – 2.5%; H3 Inland cliffs, rock pavements and outcrops – 1.7%.
Futher habitat descripion. E1.2 Perennial calcareous grassland and basic steppes; E7.2
Sub-coninental parkland; F3.1 Temperate thickets and scrub; G1.6 Fagus woodland; G3.4
Pinus sylvestris woodland south of the taiga; G3.F Highly artificial coniferous plantations;
G5.6 Early-stage natural and semi-natural woodlands and regrowth; H3.2 Basic and ultrabasic inland cliffs; H3.5 Almost bare rock pavements, including limestone pavements.
Land use: nature conservation and research – major, forestry – major.
Protected areas: overlaps (196 ha) with state botanical nature monument “Lysa Hora i Hora
Sypukha”, overlaps (about 345 ha) with Pivnichne Podillia naional nature park, included in
Emerald Site “Pivnichne Podillia”.
Threats: abandonment/reducion of land management – medium, burning of vegetaion
– low.
General description. Hills with beech forests, pine plantaions, grasslands and marl
outcrops. The grasslands are dominated by Briza media, Carex lava, Carex humilis,
Dactylis glomerata, Festuca pratensis, Festuca rubra, Festuca rupicola, Helictotrichon
desertorum, Inula ensifolia, Teucrium chamaedrys. Frequent species are Allium lusitanicum
s.l. (Allium montanum auct.), Anemone sylvestris, Asperula cynanchica, Bupleurum
falcatum, Campanula sibirica, Centaurea pannonica, Euphorbia cyparissias, Galium verum,
Lembotropis nigricans, Leontodon hispidus, Linum catharicum, Onobrychis arenaria,
Pimpinella saxifraga, Plantago media, Potenilla incana (P. arenaria), Primula veris, Prunella
grandilora, Ranunculus polyanthemos, Ranunculus zapalowiczii, Scabiosa ochroleuca,
Thymus marschallianus, Thymus pannonicus, Salvia pratensis.
Botanical significance. Important area for Carlina onopordifolia.
Criterion A
• Carlina onopordifolia Besser ex Szafer, Kulcz. et Pawł.; A(i), A(ii), A(iv); abundance:
frequent (2000 individuals); trend: stable; species data quality: good; trend data quality:
medium.
Conservaion proposals: do not allow natural and ariicial aforestaion, expand state
botanical monument “Lysa Hora i Hora Sypukha”.
Literature
1. Дідух Я., Коротченко І. Ксеротермна рослинність північно-західного Поділля //
Вісник Львівського ун-ту. Серія біологічна. – 2003. – 34. – С. 82–91.
2. Кагало О.О., Загульський М.М., Зеленчук А.Т., Сичак Н.М. Судинні рослини
державного заказника “Лиса гора та гора Сипуха” в Золочівському районі Львівської
області // Наукові основи збереження біотичної різноманітності: тематичний збірник
Інституту екології Карпат НАН України. – Вип. 6. – Львів: Ліга-Прес, 2006. – С. 66–81.
3. Мельник В.І., Скоропляс І.О., Баточенко В.М. Сучасний стан популяцій Carlina
onopordifolia (Asteraceae) на західному Волино-Поділлі // Укр. ботан. журн. – 2014. –
71, № 2. – С. 35–48.
4. Мельник В.И., Скоропляс И.О. Современное состояние популяций Сarlina
onopordifolia Besser ex Szafer, Kulcz. & Pawl. в Украине // Austrian Journal of Technical and
Natural Sciences. – 2014. № 9-10. – С. 21–24.
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Lysohirka
V.P. Kolomiychuk
Ukrainian name: Лисогірка.
Transliteraion/Translaion variants: Lysogirka.
Area: 4.8 ha.
Altitude: 32–43 m.
Latitude: 47°39’44” N (47.6621°).
Longitude: 35°06’19” E (35.1052°).
Administrative regions. Zaporizhia region: Zaporizhia raion.
Ownership: state.
Biogeographic regions: steppic.
Habitats. Level 1. E – 65.0%; G – 30.0%; H – 5.0%.
Habitats. Level 2. E1 Dry grasslands – 65.0%; G3 Coniferous woodland – 30.0%; H5
Miscellaneous inland habitats with very sparse or no vegetation – 5.0%.
Futher habitat description. E1.2 Perennial calcareous grassland and basic steppes; E1.9
Open non-Mediterranean dry acid and neutral grassland including inland dune grassland (×
E1.1); G3.F Highly artificial coniferous plantations; H5.3 Sparsely- or un-vegetated habitats
on mineral substrates not resulting from recent ice activity; H5.6 Trampled areas.
Land use: forestry – 100%.
Protected areas: –.
Threats: forestry (aforestaion) – high, burning of vegetaion – low.
General description. Inland dunes on the right bank of the Dnipro river. Dry sand grasslands
are dominated by Artemisia marschalliana, major co-dominant is Secale sylvestre. Species
of signiicant constancy: Festuca beckeri, Koeleria sabuletorum, Dianthus platyodon, Oites
borysthenica, Chondrilla juncea, Gypsophilla paniculata, Erigeron canadensis, Digitaria
sanquinalis, Eragrosis minor, Kochia lanilora, Setaria pycnocoma, Tragus racemosus. Total
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cover is 50–80%. Besides there are forest plantaions of Pinus pallasiana.
Botanical significance. Single known locality of narrow endemic species Centaurea
appendicata.
Criterion A
• Centaurea appendicata Klokov; A(iii); abundance: 270 individuals; trend: unknown;
species data quality: good; trend data quality: poor.
Conservaion proposals. Create a state botanical nature monument.
Literature
1. Коломійчук В.П. Сучасний список регіонально рідкісних судинних рослин
Запорізької області // Промышленная ботаника. – 2011. – вып. 11. – С. 105‒111.
2. Мойсієнко І.І., Коломійчук В.П., Діденко В.І., Костіков І.Ю., Баданіна В.А.,
Захарова М.Я., Овсієнко В.М. Білоперлинні волошки берегів Каховського водосховища:
сучасний стан популяцій та заходи з їх охорони // Чорноморськ. бот. журнал. – 2014. –
Т. 10, № 3. – С. 352–357.
3. Червона книга України. Рослинний світ / за ред. Я.П. Дідуха. – К.: Глобалконсалтинг,
2009. – 900 с.
Marina Hora
V.A. Onyshchenko
Ukrainian name: Мар’їна гора.
Transliteraion/Translaion variants: Mar’yina Gora, Maria’s Hill.
Area: 84.7 ha.
Altitude: 57–114 m.
Latitude: 48°55’42” N (48.9282°).
Longitude: 38°09’16” E (38.1544°).
Administrative regions. Donetsk region: Bakhmut (Artemivsk) raion.
Ownership: state.
Biogeographic regions: steppic.
Habitats. Level 1. E – 96.1%; F – 0.2%; G – 1.5%; H – 2.2%.
Habitats. Level 2. E1 Dry grasslands – 96.1%; F3 Temperate and mediterranean-montane
scrub – 0.2%; G1 Broadleaved deciduous woodland – 1.5%; H2 Screes – 2.0%; H5
Miscellaneous inland habitats with very sparse or no vegetation – 0.2%; H5.6 Trampled
areas.
Futher habitat description. E1.1 Inland sand and rock with open vegetation; E1.2 Perennial
calcareous grassland and basic steppes; F3.2 Submediterranean deciduous thickets and
brushes; G1.7 Thermophilous deciduous woodland; H2.6 Calcareous and ultra-basic screes
of warm exposures.
Land use: agriculture (animals) – major; nature conservaion and research – major;
tourism/recreaion – minor.
Protected areas: overlaps (about 80 ha) with regional botanical nature monument “Marina
Hora”, overlaps (83 ha) with Emerald Site “Kreminski Lisy”).
Threats: –.
General description. Slope of the Siverskyi Donets valley with the steppe vegataion and
chalk outcrops.
Botanical significance. Important area for some endemic species of chalk outcrops.
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Criterion A
• Alyssum gymnopodum P.A.Smirn. (Odontarrhena gymnopoda (P.A.Smirn.) D.A.German);
A(iv); abundance: unknown; trend: unknown; species data quality: poor; trend data
quality: poor.
• Diplotaxis cretacea Kotov; A(i); abundance: unknown: rare; trend: unknown; species
data quality: por; trend data quality: poor.
• Schivereckia podolica Andrz. ex DC. (Draba podolica (Besser) Rupr.); A(ii); abundance:
unknown; trend: unknown; species data quality: poor; trend data quality: poor.
• Syrenia talievi Klokov; A(iv); abundance: unknown; trend: unknown; species data
quality: poor; trend data quality: poor.
calcareous spring mires – 45.0%; D5 Sedge and reedbeds, normally without free-standing
water – 42.6%; F9 Riverine and fen scrubs – 0.3%.
Futher habitat descripion. D4.1 Rich fens, including eutrophic tall-herb fens and calcareous
flushes and soaks; D5.1 Reedbeds normally without free-standing water; D5.2 Beds of large
sedges normally without free-standing water; E3.4 Moist or wet eutrophic and mesotrophic
grassland; F9.2 Salix carr and fen scrub.
Land use: nature conservation and research – 100%.
Protected areas: included in regional hydrological reserve “Luha-Svynoryika”, included in
Emerald Site “Markovychi”.
Threats: abandonment/reducion of land management – low, burning of vegetaion – low.
General description. Major vegetaion type is the rich fens dominated by Schoenus
ferrugineus, Cladium mariscus, Carex lava, Pragmites australis, Carex elata, Eriophorum
laifolioum, with presence of Briza media, Calliergonella cuspidata, Carex panicea, Carex
nigra, Drepanocladus intermedius, Drepanocladus polygamus, Lysimachia vulgaris,
Molinia caerulea, Parnassia palustris, Potenilla erecta, Salix rozmarinifolia. Besides there
are marshes dominated by Carex acuiformis, Carex elata, Phragmites australis without
signiicant presence of calciphile species.
Botanical significance. One of the largest rich fens in Ukraine.
Literature
1. Червона книга України. Рослинний світ / за ред. Я.П. Дідуха. – К.: Глобалконсалтинг,
2009. – 900 с.
Markovychi
V.V. Datsiuk
Ukrainian name: Марковичі.
Area: 51.4 ha.
Altitude: 198–205 m.
Latitude: 50°43’48” N (50.7298°).
Longitude: 24°43’13” E (24.7201°).
Administrative regions. Volynska region: Lokachi raion.
Ownership: state.
Biogeographic regions: coninental.
Habitats. Level 1. C – 0.1%; D – 87.6; E – 12.0%; %; F – 0.3%.
Habitats. Level 2. E3 Seasonally wet and wet grasslands – 12.0%; D4 Base-rich fens and
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Criterion C
• D4.1 Rich fens, including eutrophic tall-herb fens and calcareous flushes and soaks;
area: 22 ha; trend: stable; area data quality: medium; trend data quality: medium.
Literature
1. Дацюк В.В., Андрієнко Т.Л Найбільший в Україні локалітет рідкісних угруповань
Schoenus ferrugineus L. та Сladium mariscus (L.) Pohl (Cyperaceae) на Волинському
лесовому плато ∕∕ Наук. вісник. Східноєвропейського національного університету імені
Лесі Українки. Біологічні науки. – Луцьк. – 2013. – № 14. – С. 8–13.
183
Marmaroski Hory
V.A. Onyshchenko, V.M. Virchenko
Ukrainian name: Мармароські гори.
Transliteraion/Translaion variants: Marmarosh Mountains, Maramureș Mountains.
Area: 8464.0 ha.
Altitude: 435–1940 m.
Latitude: 47°56’41” N (47.9446°).
Longitude: 24°18’14” E (24.3037°).
Administrative regions. Zakarpatska region: Rakhiv raion.
Ownership: state.
Biogeographic regions: alpine.
Habitats. Level 1. C – 0.4%; D – 0.1%; E – 8.4%; F – 1.6%; G – 89.1%; H – 0.4%.
Habitats. Level 2. C2 Surface running waters – 0.4%; E1 Dry grasslands – 0.6%; E4 Alpine
and subalpine grasslands stands – 5.8%; E5 Woodland fringes and clearings and tall forb
stands – 2.0%; F2 Arctic, alpine and subalpine scrub – 1.6%; G1 Broadleaved deciduous
woodland – 7.0%; G3 Coniferous woodland – 14.2%; G4 Mixed deciduous and coniferous
woodland – 67.9%; H2 Screes – 0.1%; H3 Inland cliffs, rock pavements and outcrops – 0.1%;
H5 Miscellaneous inland habitats with very sparse or no vegetation – 0.2%.
Futher habitat description. C2.1 Springs, spring brooks and geysers; C2.2 Permanent nontidal, fast, turbulent watercourses; E1.7 Non-Mediterranean dry acid and neutral closed
grassland (E1.71 Nardus stricta swards, E1.72 Agrosis – Festuca grassland); E2.2 Low and
medium alitude hay meadows; E3.4 Moist or wet eutrophic and mesotrophic grassland;
E4.1 Vegetated snow-patch; E4.2 Moss and lichen dominated mountain summits, ridges
and exposed slopes; E4.3 Acid alpine and subalpine grassland; E4.4 Calcareous alpine and
subalpine grassland; E5.4 Moist or wet tall-herb and fern fringes and meadows; E5.5
Subalpine moist or wet tall-herb and fern stands; F2.2 Evergreen alpine and subalpine heath
and scrub; F2.3 Subalpine deciduous scrub; F2.4 Conifer scrub close to the tree limit; G1.1
Riparian and gallery woodland, with dominant Alnus, Betula, Populus or Salix; G1.6 Fagus
woodland; G3.1 Abies and Picea woodland; G4.6 Mixed Abies – Picea – Fagus woodland;
H2.3 Temperate-montane acid siliceous screes; H2.4 Temperate-montane calcareous and
ultra-basic screes; H3.1 Acid silicious inland clifs; H3.2 Basic and ultra-basic inland clifs;
H5.6 Trampled areas.
Land use: agriculture (animals) – minor, forestry – minor, nature conservation and research
– major.
Protected areas: included in Karpatskyi biosphere reserve (naional category), included in
Carpathian biosphere reserve (UNESCO), included in Emerald Site “Carpathian Biosphere
Reserve”, overlaps (8474.0 ha) with World Heritage Site “Primeval Beech Forests of the
Carpathians and the Ancient Beech Forests of Germany” (core zone: 2243.6 ha, bufer zone:
6230.4 ha).
Threats: climate change/sea level rise – low.
General description. At lower alitudes, there prevails mixed forests co-dominated by Fagus
sylvaica, Picea abies and Abies alba and pure Fagus sylvaica forests. The herb layer is
composed predominantly of Anemone nemorosa, Athyrium ilix-femina, Dryopteris ilixmas, Galium odoratum, Mercurialis perennis, Oxalis acetosella, Symphytum cordatum. At
higher alitudes, there are pure Picea abies forests and subalpine vegetaion. Subalpine
vegetaion is represented mainly by Pinus mugo, Duschekia alnobetua (Alnus alnobetula),
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Juniperus sibirica scrubs, Vaccinium myrillus heaths, grasslands dominated by Nardus stricta
and Deschampsia cespitosa. Besides there are communiies dominated by Rhododendron
myrifolium, Festuca inarmata, Festuca carpaica, Festuca saxailis, Poa deylii.
Botanical significance. Important area for beech, mixed and spruce forests, subalpine
habitats and species.
Criterion A
• Botrychium multifidum (S.G.Gmel.) Rupr.; A(ii); abundance: rare; trend: unknown;
species data quality: medium; trend data quality: poor.
• Campanula abieina Griseb. & Schenk; A(ii); abundance: occasional; trend: stable;
species data quality: medium; trend data quality: poor.
• Campanula serrata (Kit.) Hendrych; A(ii); abundance: rare; trend: unknown; species
data quality: medium; trend data quality: poor.
• Dicranum viride (Sull. et Lesq.) Lindb.; A(ii); abundance: occasional; trend: unknown;
species data quality: medium; trend data quality: poor.
• Hamatocaulis vernicosus (Mitt.) Hedenas (Drepanocladus vernicosus (Mitt.) Warnst.);
A(ii); abundance: rare; trend: unknown; species data quality: medium; trend data
quality: poor.
• Narcissus angustifolius Curtis; A(ii); abundance: rare; trend: unknown; species data
quality: medium; trend data quality: poor.
• Poa graniica Braun.-Blanq. (Poa deylii Chrtek & V.Jira); A(ii); abundance: rare; trend:
unknown; species data quality: medium; trend data quality: poor.
• Tozzia carpathica Woł.; A(ii); abundance: rare; trend: unknown; species data quality:
medium; trend data quality: poor.
Criterion C
• C2.12 Hard water springs; area: 0.1 ha; trend: stable; area data quality: poor; trend data
quality: poor.
• D2.226 Peri-Danubian black-white-star sedge fens; area: 1 ha; trend: unknown; area
data quality: poor; trend data quality: poor.
185
•
E4.11 Boreo-alpine acidocline snow-patch grassland and herb habitats; area: 1 ha;
trend: stable; area data quality: poor; trend data quality: poor.
• E4.3 Acid alpine and subalpine grassland; area: 450 ha; trend: stable; area data quality:
poor; trend data quality: poor.
• E4.4 Calcareous alpine and subalpine grassland; area: 2 ha; trend: stable; area data
quality: poor; trend data quality: poor.
• E5.5 Subalpine moist or wet tall-herb and fern stands; area: 150 ha; trend: stable; area
data quality: poor; trend data quality: poor.
• F2.22 Alpide acidocline Rhododendron heaths; area: 3 ha; trend: stable; area data
quality: poor; trend data quality: poor.
• G1.12 Boreo-alpine riparian galleries; area: 80 ha; trend: stable; area data quality:
medium; trend data quality: medium.
• G1.A4 (*9180) Ravine and slope woodland; area: 30 ha; trend: stable; area data quality:
poor; trend data quality: medium.
• H2.3 Temperate-montane acid siliceous screes; area: 10 ha; trend: stable; area data
quality: poor; trend data quality: medium.
Literature
1.Біорізноманіття Карпатського біосферного заповідника / Кол. авт., Ред. рада:
Я.І. Мовчан, Ф.Д. Гамор та ін. – К.: Інтерекоцентр, 1997. – 711 с.
2.Гамор Ф.Д., Волощук М.І., Антосяк Т.М., Козурак А.В. БЗ Карпатський //
Фіторізноманіття заповідників і національних природних парків України. Ч. 1. Біосферні
заповідники. Природні заповідники / під ред. В.А. Онищенка і Т.Л. Андрієнко. – Київ:
Фітосоціоцентр, 2012. – С. 45–72.
3.Гамор Ф.Д., Довганич Я.О., Покиньчереда В.Ф. та ін. Праліси Закарпаття.
Інвентаризація та менеджмент. – Рахів, 2008. – 86 с.
4.Малиновський К.А. Рослинність високогір’я Українських Карпат. – К.: Наук. думка,
1980.– 280 с.
5.Малиновський К.А., Крічфалушій В.В. Рослинні угруповання високогір’я
Українських Карпат. – Ужгород, 2002. – 244 с.
woodland – 95.5%; G3 Coniferous woodland – 1.0%; H3 Inland clifs, rock pavements and
outcrops – 0.0%; H5 Miscellaneous inland habitats with very sparse or no vegetation – 1.0%
Futher habitat descripion. E1.1 Inland sand and rock with open vegetation; E1.2 Perennial
calcareous grassland and basic steppes; E2.2 Low and medium alitude hay meadows; G1.6
Fagus woodland; G1.A Meso- and eutrophic Quercus, Carpinus, Fraxinus, Acer, Tilia, Ulmus
Medobory
V.A. Onyshchenko
Ukrainian name: Медобори.
Area: 11811 ha.
Altitude: 235–414 m.
Latitude: 49°15’06” N (49.2515°).
Longitude: 26°10’33” E (26.1758°).
Administrative regions. Khmelnytskyi region: Chemerivtsi raion, Horodok raion; Ternopil
region: Husiatyn raion, Pidvolochysk raion.
Ownership: state.
Biogeographic regions: coninental.
Habitats. Level 1. C – 0.1%; D – 0.1%; E – 2.1%; F – 0.2%; G – 96.5%; H – 1.0%.
Habitats. Level 2. C2 Surface running waters – 0.1%; D5 Sedge and reedbeds, normally
without free-standing water – 0.1%; E1 Dry grasslands – 1.1%; E2 Mesic grasslands – 1.0%;
F3 Temperate and mediterranean-montane scrub – 0.2%; G1 Broadleaved deciduous
186
and related woodland; G3.F Highly artificial coniferous forestry plantations; H3.2 Basic and
ultra-basic inland clifs; H3.5 Almost bare rock pavements, including limestone pavements;
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H5.6 Trampled areas.
Land use: agriculture (animals) – minor; forestry – minor, mowing/hay making – minor,
nature conservation and research – 98%.
Protected areas: overlaps with Medobory nature reserve (about 8786 ha), overlaps with
Podilski Tovtry naional nature park (about 2810 ha), includes Satanivskyi state forest
reserve (1015 ha), includes Ivakhnovenskyi state landscape reserve (1778 ha), includes
Velyka and Mala Buhaikha state landscape reserve (66 ha), includes Vilkhovetski Tovtry
regional botanical reserve (47 ha), overlaps with Emerald Site “Medobory Nature Reserve”
(about 8786 ha), overlaps with Emerald Site “Podiliski Tovtry Naional Nature Park” (about
2950 ha).
Threats: abandonment/reducion of land management – low.
General description. A part of the Tovtry hills. Major vegetaion type is the broadleaved
forest. The tree layer is formed mainly by Carpinus betulus, Quercus robur, Fraxinus excelsior. Main dominants of the herb layer are Aegopodium podagraria, Anemone nemorosa,
Anemone ranunculoides, Carex pilosa, Corydalis cava, Isopyrum thalictroides, Lamium galeobdolon. Other typical species are Acer platanoides, Acer pseudoplatanus, Corydalis solida, Euonymus europaeus, Galium odoratum, Glechoma hirsuta, Milium efusum, Mercurialis
perennis, Polygonatum hirtum, Polygonatum mulilorum, Pulmonaria obscura, Ranunculus cassubicus, Sambucus nigra, Stachys sylvaica, Stellaria holostea, Tilia cordata, Urica
dioica. On the stony slopes and tops of the limestone hills, there occur forests dominated
by Fraxinus excelsior and Acer platanoides with diferenial species Arum besseranum, Bromopsis benekenii, Hordelymus europaeus, Hylotelephium polonicum, Lamium maculatum,
Lunaria rediviva, Melandrium dioicum, Scutellaria alissima, Viburum lantana. The area includes the easternmost natural Fagus sylvaica forest in the coninental biogeographical
region. In some places there is the steppe vegetaion in complex with the shrub vegetaion.
Main dominants of the steppes are Carex humilis, Festuca valesiaca s.l., Poa angusifolia.
Prevailng species of shrubs are Chamaecyisus albus, Crataegus leiomonogyna, Juniperus
communis, Prunus spinosa, Swida sanguinea (Cornus sanguinea). Hay medows in the loodplain of the Zbruch river are dominated by Arrhenatherum elaius, Dactylis glomerata, Poa
pratensis. Besides in other places Calamagrosis epigeios meadows occur.
Botanical significance. One of the best oak-hornbeam forests in the latland porion of
Ukraine. Besides this area is important for Chamaecyisus podolicus and communiies of
Juniperus communis. Other important species: Aconitum besseranum, Cypripedium calceolus, Chamaecyisus blockianus, Euphorbia volhynica, Schivereckia podolica (Draba podolica).
Criterion A
• Aconitum besserianum Andrz.; A(iiі); abundance: rare; trend: stable; species data
quality: medium; trend data quality: medium.
• Chamaecytisus podolicus (Błocki) Klásk. A(iv); abundance: rare; trend: unknown; species
data quality: medium; trend data quality: poor.
• Schivereckia podolica (Besser) Andrz. ex DC. (Draba podolica (Besser) Rupr.); A(ii);
abundance: rare; trend: unknown; species data quality: medium; trend data quality:
poor.
Criterion B
• G1 Broadleaved deciduous woodland; area 11100 ha; % of indicator species: 13.0%; No
of indicator species: 10; trend: increasing; species data quality: good; area data quality:
good; trend data quality: good.
Criterion C
• F3.16 Juniperus communis scrub; area: 16 ha; trend: stable; area data quality: medium;
trend data quality: poor.
G1.A1 Quercus – Fraxinus – Carpinus betulus woodland on eutrophic and mesotrophic
soils; area: 9000 ha; trend: increasing; area data quality: medium; trend data quality:
good.
• G1.A4 (*9180) Ravine and slope woodland; area: 200 ha; trend: stable; area data quality:
medium; trend data quality: good.
Literature
1. Onyshchenko V.A. Forests of order Fagetalia sylvaticae in Ukraine. – Kyiv: Alterpress,
2009. – 212 p.
2. Szafer W. Geobotaniczne stosunki Miodoborόw Galicyjskich // Rozpr. Wydz.
Matemat.-Przyrod. Akad. Umiejętności. – 1910. – Ser. 3, Dz. B, T. 10. – S. 63–172.
3. Данилків І.С., Рабик І.В. Мохоподібні (Bryophytes) природного заповідника
“Медобори” // Чорноморський ботан. журн. – 2007. – 3, № 1. – С. 85–99.
4. Коломієць І.В., Дудка І.О., Тихоненко Ю.Я. Іржасті гриби заповідника “Медобори”
// Укр. ботан. журн. – 1996. – 53, № 3. – С. 201–207.
5. Кондратюк С.Я. Лишайники заповідника “Медобори” // Укр. ботан. журн. – 1995.
– 52, № 1. – С. 141–144.
6. Кондратюк С.Я., Коломієць І.В. Нові для України види лишайників та ліхенофільних
грибів заповідника “Медобори” // Укр. ботан. журн. – 1997. – 54, № 1. – С. 42–47.
7. Котов М. Геоботанический очерк буковых лесов по р. Збруч // Журнал Русского
ботанического общества. – 1930. – 15, № 1–2. – P. 139–148.
8. Мельник В.І., Корінько О.М. Букові ліси Подільської височини. – Київ:
Фітосоціоцентр, 2005. – 152 с.
9. Оліяр Г.І. Рослини Червоної книги України в природному заповіднику
“Медобори”// Заповідна справа в Україні. – 1995. – Т. 1. – С. 11–12.
10. Оліяр Г.І. Конспект флори природного заповідника “Медобори” // Наукові
записки ТДПУ. Серія: Біологія, № 2 (17). – 2002. – С. 18–25.
11. Оліяр Г.І. Зміни у флорі природного заповідника “Медобори” за минуле
сторіччя // Роль природно-заповідних територій Західного Поділля та Юри Ойцовської
у збереженні біолог. та ландшафтн. різноманіття, – Гримайлів, 2003. – С. 333–338.
12. Оліяр Г. І. Фітораритети природного заповідника “Медобори” з філією
“Кременецькі гори” в міжнародних червоних списках // Природно-заповідний
фонд України – минуле, сьогодення, майбутнє. Матеріали міжнародної науковопрактичної конференції, присвяченої 20-річчю природного заповідника “Медобори”
(смт. Гримайлів, 26–28 травня 2010 р.) – Тернопіль: Підручники і посібники, 2010. –
С. 460–464.
13. Онищенко В.А. Рослинність карбонатних відслонень природного заповідника
“Медобори” // Укр. фітоцен. зб. – Сер. A. – 2001. – 1 (17). – С. 86–104.
14. Онищенко В.А., Оліяр Г.І. // Рідкісні лісові угруповання природного
заповідника “Медобори” // Укр. ботан. журн. – 1998. – 55, № 4. – С. 413–416.
15. Онищенко В.А., Оліяр Г.І. ПЗ Медобори // Фіторізноманіття заповідників
і національних природних парків України. Ч. 1. Біосферні заповідники. Природні
заповідники / під ред. В.А. Онищенка і Т.Л. Андрієнко. – Київ: Фітосоціоцентр, 2012. –
С. 254–276.
16. Смеречинская Т. А. Закономерности распространения лишайников по
фитоценозам природного заповедника “Медоборы” // Заповідна справа в Україні. –
2005. – Т.11, Випуск 1. – С. 9–15.
188
189
•
Medvezhanka
V.A. Onyshchenko, M.M. Peregrym
Ukrainian name: Медвежанка.
Area: 7849.0 ha.
Altitude: 80–223 m.
Latitude: 48°13’24” N (48.2234°).
Longitude: 39°31’34” E (39.5260°).
Administrative regions. Luhansk region: Dovzhansk (Sverdlovsk) raion, Lutuhyne raion,
Sorokyne (Krasnodon) raion.
Ownership: state.
Biogeographic regions: steppic
ultra-basic inland cliffs; H5.6 Trampled areas; I1.5 Bare illed, fallow or recently abandoned
arable land.
Land use: agriculture (animals) – major; agriculture (arable) – minor; extracion (minerals)
– minor; forestry – minor; nature conservaion and research – minor.
Protected areas: overlaps (about 1135 ha) with regional landscape reserve “Krasnianske
Vodoskhovyshche”, overlaps (7790 ha) with Emerald Site “Vedmezhanka”.
Threats: agricultural intensification/expansion (arable) – medium, agricultural
intensiicaion/expansion (grazing) – low, extracion (minerals/quarries) – low, forestry
(aforestaion) – low.
General description. The area is situated within the Donetsk Ridge. Dominants of the steppe
vegetaion are Festuca valesiaca, Bothriochloa ischaemum, Bromopsis riparia, Bromopsis
inermis, Elytrigia intermedia, Poa angusifolia, Sipa capillata, S. lessingiana, S. irsa, S.
ucrainica. On the neutral outcrops of slate and sandstone, there are Thymus marschallianus,
Pimpinella titanophila, Artemisia marschalliana, Agropyron pectinatum.
Botanical signiicance. One of the largest areas of the steppe vegetaion in Ukraine.
Criterion C
• E1.2 Perennial calcareous grassland and basic steppes; area: 7000 ha; trend: decreasing;
area data quality: good; trend data quality: poor.
Conservaion proposals. Do not allow plowing. Сreate a naional nature park.
Literature
1. Природно-заповідний фонд Луганської області // О.А. Арапов (заг. ред.), Т.B. Сова,
О.А. Cавенко, В.Б. Фєрєнц, Н.У. Кравець, Л.Л. Зятьков, Л.О. Морозова. Довідник. – 3-е
вид., доп. і переробл. – Луганськ: “Луганська правда”, 2013. – 224 с.
Mehanom
Habitats. Level 1. E – 90.0%; F – 1.0%; G – 3.0%; H – 1.0%; I – 5.0%;
Habitats. Level 2. E1 Dry grasslands – 89.9%; E2 Mesic grasslands – 0.1%; F3 Temperate
and mediterranean-montane scrub – 1.0%; G1 Broadleaved deciduous woodland – 2.6%;
G3 Coniferous woodland – 0.4%; H2 Screes – 0.1%; H3 Inland cliffs, rock pavements and
outcrops – 0.1%; H5 Miscellaneous inland habitats with very sparse or no vegetation –
0.8%; I1 Arable land and market gardens – 5.0%.
Futher habitat description. E1.1 Inland sand and rock with open vegetation; E1.2 Perennial
calcareous grassland and basic steppes; F3.2 Submediterranean deciduous thickets and
brushes; G1.7 Thermophilous deciduous woodland; G3.F Highly artificial coniferous
plantations; H2.6 Calcareous and ultra-basic screes of warm exposures; H3.2 Basic and
V.A. Onyshchenko
Ukrainian name: Меганом.
Transliteraion/Translaion variants: Meganom.
Area: 1936.0 ha.
Altitude: 0–358 m.
Latitude: 44°48’55” N (44.8152°).
Longitude: 35°05’34” E (35.0929°).
Administrative regions. Autonomous Republic of Crimea: Sudak city; Territorial waters of
Ukraine.
Ownership: state.
Biogeographic regions: steppic.
Habitats. Level 1. A – 6.6%; B – 1.3%; E – 63.1%; F – 4.7%; G – 20.7%; H – 3.7%; J – 0.1%.
Habitats. Level 2. A1 Littoral rock and other hard substrata – 0.2%; A2 Littoral sediment –
0.2%; A3 Infralittoral rock and other hard substrata – 2.1%; A5 Sublittoral sediment – 4.1%;
B2 Coastal shingle – 0.1%; B3 Rock cliffs, ledges and shores, including the supralittoral –
1.2%; E1 Dry grasslands – 59.9%; E6 Salt steppes – 3.1%; F3 Temperate and mediterraneanmontane scrub – 2.7%; F7 Spiny Mediterranean heaths – 2.0%; F9 Riverine and fen scrubs
– 0.0%; G1 Broadleaved deciduous woodland – 20.7%; H2 Screes – 0.5%; H3 Inland clifs,
rock pavements and outcrops – 3.1%; H5 Miscellaneous inland habitats with very sparse
or no vegetation – 0.1%.
190
191
Futher habitat description. A2.1 Litoral coarse sediment; A2.5 Coastal saltmarshes and
saline reedbeds; A3.2 Atlanic and Mediterranean moderate energy infralitoral rock; A3.2
Atlanic and Mediterranean low energy infralitoral rock; A5.1 Sublitoral coarse sediment;
A5.5 Sublitoral macrophyte-dominated sediment; B2.1 Shingle beach dritlines; B2.2
Unvegetated mobile shingle beaches above the dritline; B2.3 Upper shingle beaches
with open vegetaion; B3.1 Supralitoral rock (lichen or splash zone); B3.2 Unvegetated
rock clifs, ledges, shores and islets; B3.3 Rock clifs, ledges and shores, with angiosperms;
B3.4 Sot sea-clifs, oten vegetated; E1.1 Inland sand and rock with open vegetation; E1.2
Perennial calcareous grassland and basic steppes; E1.3 Mediterranean xeric grassland;
F3.2 Submediterranean deciduous thickets and brushes; F7.4 Hedgehog-heaths; F9.3
Southern riparian galleries and thickets; G1.7 Thermophilous deciduous woodland; G1.C
Highly ariicial broadleaved deciduous forestry plantations; H2.6 Calcareous and ultrabasic screes of warm exposures; H3.2 Basic and ultra-basic inland clifs; H5.3 Sparsely- or
un-vegetated habitats on mineral substrates not resulting from recent ice activity; H5.6
Trampled areas.
Land use: tourism/recreaion – minor, urban/industrial/transport – 2% (wind turbines,
road).
Protected areas: includes Pivostriv Mehanom regional complex nature monument (651.6
ha), included in Emerald Site “Mehanom”.
Threats: development (recreaion/tourism) – low, eutrophicaion – low.
General description. Mehanom is a peninsula in the southeastern part of Crimea. This is the
most arid place in Crimea. Hills are covered chiely with the steppe vegetaion. Maximum
alitude is 358 m. Slope to the sea is steep, with clifs about 100 m high. Adjacent sea is
included. Surface of the shore consists of rocks, boulders, pebble and gravel. Dominants
of the steppe vegetaion are Agropyron pecinatum, Bothriochloa ischaemum, Bromopsis
cappadocica, Festuca valesiaca, Koeleria cristata, Sipa capillata, Sipa ponica, Sipa
ucrainica. Typical are Achillea nobilis, Achillea millefolium, Galatella linosyris, G. villosa,
Cruciata taurica, Inula germanica, Inula oculus-christi, Jurinea roegneri, Centaurea salonitana,
Limonium platyphyllum, Linum austriacum, Linum nervosum, Limonium squamulosum,
Nepeta ucranica, Onosma taurica, Onosma visianii, Phlomis pungens, Poterium polygamum,
Psephellus trinervius, Salvia nemorosa s.l., Scabiosa argentea, Stachys cretica subsp. velata.
Petrophyic steppe difers with presence of Alyssum tortuosum (Odontarrhena tortuosa),
Ephedra distachya, Euphorbia petrophila, Fumana procumbens, Paronychia cephalotes,
Siederitis syriaca, Thymus roegneri, Thymus tauricus. Salt steppes are dominanted by
Artemisia taurica, Artemisia lercheana, Camphorosma monspeliaca, Petrosimonia brachiata.
Dry grasslands of mediterranean type with signiicant paricipaion of annual species
(Aegilops biuncialis, Aegilops cylindrica, Aegilops triuncialis, Alyssum hirsutum, Alyssum
desertorum, Bromus japonicus, Bromus hordeaceus, Bromus sguarrosus, Gaudinopsis
macra, Taeniatherum asperum, Taeniatherum crinitum) occupy about 100 ha. In dry rocky
habitats, there is open vegetaion with Alyssum murale, Asphodeline taurica, Cleistogenes
serotina, Ephedra distachya, Euphorbia petrophila, Fumana procumbens, Hedysarum
candidum, Helianthemum grandiflorum, Helianthemum nummularium, Medicago cretacea,
Melica monticola, Melica taurica, Pimpinella tragium, Poa sterilis, Sideritis syriaca subsp.
catillaris, Sedum acre, Sedum hispanicum, Seseli gummiferum, Teucrium polium, Thymus
roegneri, Thymus tauricus, Veronica multifida. On the sea beach, there occur Crambe
mariima, Cynanchum acutum, Limonium gmelini, Peganum harmala, several species of
Atriplex and Soda. Main species of trees in sparse woods is Quercus pubescens. Besides
there are Carpinus orientalis, Clematis vitalba, Cotoneaster tauricus, Cotinus coggygria,
species of Crataegus, Paliurus spina-christi, Pyrus communis, Pyrus eleagnifolia. The herb
layer consists mainly of steppe and petrophyic species, some wood species occur too. Shrub
communiies are dominated by Rosa canina, R. spinosissima, Paliurus spina-christi, Cotinus
coggygria. In valleys, there are small areas of Tamarix ramosissima, Tamarix tetrandra and
Eleagnus angusifolia stands. In the sea at depths to 3 m, dominant species are Ceramium
ciliatum, Cladostephus vericillatus, Corallina mediterranea, Cystosera barbata, Cystoseira
crinita, Dilophus fasciola, Enteromorpha compressa, Grateloupia dichotoma, Polysiphonia
opaca. At depths 3–10 m, there predominate Chondria tenuissima, Cladophora albida,
Cladophora dalmaica, Cystoseira crinita, Сystoseira barbata, Phyllophora nervosa, Ulva
rigida, Zostera marina, Zostera nolii (Nanozostera nolii).
Protected areas: –.
Botanical significance. Important area for conservaion of dry grasslands, sea and inland
clifs, marine macrophytes. There are threatened endemic species Astragalus reduncus,
Astragalus setosulus, Onobrychis pallasii, Onosma polyphylla, Stipa syreistschikowii.
Criterion A
• Astracantha arnacantha (M.Bieb.) Podlech; A(iv); abundance: frequent; trend:
unknown; species data quality: medium; trend data quality: poor.
• Astragalus reduncus Pall.; A(iv); abundance: rare; trend: unknown; species data quality:
medium; trend data quality: poor.
• Astragalus setosulus Gontsch.; A(i), A(ii), A(iii); abundance: rare; trend: unknown;
species data quality: medium; trend data quality: poor.
• Onobrychis pallasii (Willd.) M.Bieb.; A(iii); abundance: occasional; trend: unknown;
species data quality: medium; trend data quality: poor.
• Onosma polyphylla Ledeb.; A(ii), A(iv); abundance: rare; trend: unknown; species data
quality: medium; trend data quality: poor.
• Stipa syreistschikowii P.Smirn.; A(ii), A(iv); abundance: occasional; trend: unknown;
species data quality: medium; trend data quality: poor.
192
193
Criterion C
• A3 Infralittoral rock and other hard substrata; area: 80 ha; trend: stable; area data
quality: medium; trend data quality: poor.
• A5 Sublitoral sediment; area: 80 ha; trend: stable; area data quality: medium; trend
data quality: medium.
• B3.3 Rock cliffs, ledges and shores, with angiosperms; area: 0.1 ha; trend: stable; area
data quality: poor; trend data quality: medium.
• E1.2 Perennial calcareous grassland and basic steppes; area: 700 ha; trend: stable; area
data quality: medium; trend data quality: medium .
• F7 Spiny Mediterranean heaths; area: 40 ha; trend: stable; area data quality: poor;
trend data quality: medium.
• F9.3 Southern riparian galleries and thickets; area: 1 ha; trend: unknown; area data
quality: poor; trend data quality: poor.
Literature
1. Костенко Н.С., Дикий Е.А., Заклецкий А.А., Марченко В.С. Аквальні комплекси
бухти Лісьей та півострова Меганом – перспективі об'єкти природнозаповідного фонду
// Современные проблемы экологии Азово-Черноморского региона: материалы II
международной конференции (Керчь, 26–27 июня 2006 г.). – Керчь: ЮгНИИРО, 2006. –
С. 103–108
2. Костенко Н.С., Дикий Е.А., Заклецкий А.А., Марченко В.С. Донная растительность
приоритетных акваторий юго-восточного Крыма: современное состояние и
необходимые меры по сохранению // Заповедники Крыма – 2007. Материалы
IV международной научно-практической конференции, посвященной 10-летию
проведения международного семинара "Оценка потребностей сохранения
биоразнообразия Крыма" (Гурзуф, 1997). Часть 1. Ботаника. Общие вопросы охраны
природы. – Симферополь, 2007. – С. 63–68.
3. Миронова Л.П., Шатко В.Г. Полуостров Меганом в юго-восточном Крыму
(природные условия, флора, растительность // Геополитика и экогеодинамика
регионов. – 2013. – Т. 9, Вып. 2, Ч. 2. – С. 26–64.
Trampled areas.
Land use: forestry – major, nature conservation and research – minor.
Protected areas: overlaps with regional forest reserve “Mishkovo-Pohorilove”.
Threats: forestry (aforestaion) – medium, development (urbanization) – low.
General description. Terrace of the Inhul river with Pinus pallasiana and Pinus sylvestris
plantaions and sand grasslands. Sand grasslands are dominated by Carex colchica, Artemisia
marschalliana, Festuca beckeri, Koeleria sabuletorum, Thymus pallasianus. Other frequent
species are Sipa borysthenica, Scabiosa ucrainica, Achillea ochroleuca, Helichrysum
corymbiforme, Astragalus varius, Anchusa gmelinii, Jurinea charcoviensis, Tragopogon
borysthenicus, Alyssum savranicum (Odontarrhena savranica), Secale sylvestre. Density of
Centaurea margaritacea in natural grasslands is 3 to 30 individuals per 100 m2.
Botanical significance. 100% of global populaion of narrow endemic species Centaurea
margaritacea s. str.
Criterion A
• Centaurea margaritacea Ten. s. str.; A(iii); abundance: frequent; trend: unknown;
species data quality: good; trend data quality: poor.
Conservaion proposals. Create a state botanical reserve instead of local forest reserve
“Mishkovo-Pohorilove”. Restore sand grasslands.
Mishkovo-Pohorilove
O.M.Derkach, H.V. Kolomiets
Ukrainian name: Мішково-Погорілове.
Area: 152 ha.
Altitude: 2–41 m.
Latitude: 47°00’54” N (47.0148°).
Longitude: 32°06’11” E (32.1029°).
Administrative regions. Mykolaiv region: Vitovka (Zhovtnevyi) raion.
Ownership: state.
Biogeographic regions: steppic.
Habitats. Level 1. E – 8.0%; G – 90.0%; H – 2.0%.
Habitats. Level 2. E1 Dry grasslands – 8.0%; G3 Coniferous woodland – 90.0%; H5
Miscellaneous inland habitats with very sparse or no vegetaion – 2.0%.
Futher habitat description. E1.9 Open non-Mediterranean dry acid and neutral grassland,
including inland dune grassland (× E1.1); G3.F Highly artificial coniferous plantations; H5.6
194
Literature
1.Деркач О.М. Ключові ботанічні території Миколаївщини: сучасний стан та
проблеми збереження // Теорія і практика заповідної справи в Україні. Зб.наук.праць.
– К., 2005. – С.167–173.
2.Деркач О.М. Нижньоінгульське // Південно-Бузький меридіональний екологічний
коридор: стислий огляд біорізноманіття та найцінніші території / Під ред. В. Костюшина.
– К., 2007. – С. 53.
195
3.Коломієць Г.В. Перлинні волошки секції Pseudophalolepis Klok. ряду Margaritacea
Klok. Питання систематики та охорони // Укр. фітоценологічний зб. – Вип. 1–2 (12–13).
– К.: Фітосоціоцентр, 1999. – С. 165–169.
4.Крицька Л.І., Деркач О.М. Сучасний стан популяцій видів ряду Margaritaceae Klok.
(Centaurea L.) // Укр. ботан. журн. – 1991. – 48, № 3. – С. 78–80.
5.Крицька Л.І., Деркач О.М. Волошка первинноперлинна Centaurea margaritacea Ten.
/ Червона книга України. Рослинний світ / за ред. Я.П. Дідуха. – К.: Глобалконсалтинг,
2009. – С. 304.
6.Перлини піщаної флори у пониззях Південного Бугу та Інгулу. Cерія: Збереження
біорізноманіття в Приморсько-степовому екокоридорі / Під ред. Г.В. Коломієць. – К.:
Громадська організація “Веселий Дельфін”, 2008. – 60 c.
presence of psammophyic species Artemisia marschalliana, Sipa borysthenica, Ephedra
distachya, Helicrysum arenarium. A large area is occupied by salt meadows (dominats:
Molochnyi Lyman
V.P. Kolomiychuk, V.A. Onyshchenko
Ukrainian name: Молочний лиман.
Area: 28196.0 ha.
Altitude: 0–20 m.
Latitude: 46°34’42” N (46.5782°).
Longitude: 35°19’44” E (35.3289°).
Administrative regions. Zaporizhia region: Melitopol raion, Pryazovsk raion, Yakymivka
raion .
Ownership: state.
Biogeographic regions: steppic
Habitats. Level 1. A – 85.2%; C – 0.7%; D – 1.1%; E – 12.9%; H – 0.1%.
Habitats. Level 2. A2 Littoral sediment – 17.0%; A5 Sublittoral sediment – 68.2%; C1
Surface standing waters – 0.1%; C3 Littoral zone of inland surface waterbodies – 0.6%; D5
Sedge and reedbeds, normally without free-standing water – 0.2%; D6 Inland saline and
brackish marshes and reedbeds – 0.9%; E1 Dry grasslands – 1.0%; E6 Salt steppes – 11.9%;
H5 Miscellaneous inland habitats with very sparse or no vegetation – 0.1%.
Futher habitat description. A2.3 Littoral mud; A2.5 Coastal saltmarshes and saline
reedbeds; A5.3 Subittoral mud; C3.2 Water-fringing reedbeds and tall helophytes other
than canes; D5.1 Reedbeds normally without free-standing water; D6.1 Inland saltmarshes;
D6.2 Inland saline or brackish species-poor helophyte beds normally without free-standing
water; E1.2 Perennial calcareous grassland and basic steppes; E6.2 Continental inland salt
steppes; H5.6 Trampled areas.
Land use: agriculture (animals) – minor; nature conservaion and research – major.
Protected areas: overlaps (about 22730 ha) with Pryazovskyi naional nature park, overlaps
(about 22400 ha) with Ramsar Site “Molochnyi Liman”, overlaps (about 22730 ha) with
Emerald Site “Pryazovskyi Naional Nature Park”.
Threats: –.
General description. A lagoon in the northwestern part of the Sea of Azov and the mouth of
the Molochna river in the nothern part of the lagoon. Connecion with the sea is intermitent.
Level of water is unstable. A large area of the botom is exposed in the dry season. The right
bank is rather high. There is the steppe vegetaion dominated by Agropyron pecinatum,
Festuca valesiaca, Koeleria cristata, Sipa lessingiana, S. pulcherrima, S. ucrainica with
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Elytrigia elongata, Puccinellia distans, Aeluropus litoralis, Juncus gerardii) and wet
halophyic communiies (dominants: Phragmites australis, Scirpus tabernaemontani,
Juncus maritimus, Bolboschoenus maritimus). In the most saline habitats, there dominate
Halocnemum strobilaceum, Salicornia perennans, Suaeda prostrata, Halimione pedunculata,
H. verrucifera, Limonium caspium, L. meyeri. On elevated areas of the let bank there are salt
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steppes with dominance of Festuca valesiaca s.l., Camphorosma monspeliaca, Agropyron
pecinatum, Koeleria cristata, Crinitaria villosa. On the southern side of the bar, there are
litoral communiies with Argusia sibirica, Cakile euxina, Leymus sabulosus, Salsola pontica.
In some places, there are aquaic communiies of Zostera marina, Zannichellia major,
Myriophyllum spicatum, Ceratophyllum demersum, Potamogeton pecinatum (Stuckenia
pecinata).
Botanical significance. Important area for halophyic vegetaion, Zostera marina and some
threatened steppe species.
Criterion A
• Allium pervesitum Klokov.; A(i), A(iv); abundance: rare; trend: stable; species data
quality: good; trend data quality: good.
• Allium regelianum A.Becker ex Iljin.; A(ii); abundance: rare (100–150 individuals); trend:
unknown; species data quality: good; trend data quality: poor.
• Ferula orientalis L.; A(ii); abundance: occasional; trend: stable; species data quality:
medium; trend data quality: poor.
• Zostera marina L.; A(ii); abundance: frequent; trend: stable; species data quality:
medium; trend data quality: poor.
Criterion C
• A2.5 Coastal saltmarshes and saline reedbeds; area: 500 ha; trend: unknown; area data
quality: poor; trend data quality: poor.
• E6.2 Coninental inland salt steppes; area: 3300 ha; trend: stable; area data quality:
medium; trend data quality: medium.
Literature
1. Дубина Д.В., Шеляг-Сосонко Ю.Р. Тенденції антропогенних змін плавневоліторальних фітосистем р. Молочної // Укр. ботан. журн. – 1996. – 53, № 1–2. – С. 31–37.
2. Дубовий М.С., Дмитренко Є.М., Коломійчук В.П. Флора і рослинність Молочного
лиману // Зб. доп. VІ Міжнар. науков. конф. аспірантів і студентів “Охорона
навколишнього середовища та раціональне використання природних ресурсів” (17–19
квітня 2007 р., м. Донецьк). – Т. 1. – Донецьк: ДонНТУ, ДонНУ, 2007. – С. 176–177.
3. Коломийчук В.П., Подорожный С.Н. Флора берегов Молочного лимана //
Біологічний вісник Мелітопольського держ. пед. університуту ім. Богдана
Хмельницького. – 2013. – № 2. – С. 128–135.
4. Коломійчук В.П., Тищенко О.В. НПП Приазовський // Фіторізноманіття заповідників
і національних природних парків України. Ч. 2. Національні природні парки / під ред.
В.А. Онищенка і Т.Л. Андрієнко. – Київ: Фітосоціоцентр, 2012. – С. 410–428.
5. Коломійчук В.П., Яровий С.О. Конспект флори судинних рослин Приазовського
національного природного парку. – К.: Альтерпрес, 2011. – 296 с.
6. Тищенко О.В. Рослинність приморських кіс північного узбережжя Азовського
моря. – Київ: Фітосоціоцентр, 2006. – 156 с.
Latitude: 51°37’43” N (51.6285°).
Longitude: 26°17’05” E (26.2846°).
Administrative regions. Rivne region: Zarichne raion, Volodymyrets raion.
Ownership: state.
Biogeographic regions: coninental.
Habitats. Level 1. C – 0.02%; D – 10%; F – 0.1%; G – 90%.
Habitats. Level 2. C1 Surface standing waters – 0.01%; C2 Surface running waters – 0.02%;
D1 Raised and blanket bogs – 1%; D2 Valley mires, poor fens and transition mires – 8%;
D5 Sedge and reedbeds, normally without free-standing water – 1%; F9 Riverine and fen
scrubs – 0.1%; G1 Broadleaved deciduous woodland – 15%; G3 Coniferous woodland –
25%; G4 Mixed deciduous and coniferous woodland – 50%.
Morochno-2
Futher habitat description. C1.2 Permanent mesotrophic lakes, ponds and pools; C2.3
Permanent non-tidal, smooth-flowing watercourses; C3.2 Water-fringing reedbeds and
tall helophytes other than canes; D1.1 Raised bogs; D2.3 Transiion mires and quaking
bogs; D5.1 Reedbeds normally without free-standing water; D5.2 Beds of large sedges
normally without free-standing water; F9.2 Salix carr and fen scrub; G1.1 Riparian and
gallery woodland, with dominant Alnus, Betula, Populus or Salix; G1.5 Broadleaved swamp
V.A. Onyshchenko
Ukrainian name: Морочно-2.
Area: 4487 ha.
Altitude: 148–159 m.
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199
woodland not on acid peat; G3.4 Pinus sylvestris woodland south of the taiga; G3.E Nemoral
bog conifer woodland; G4.1 Mixed swamp woodland.
Land use: forestry – minor, nature conservation and research – 100%.
Protected areas: includes Khynotskyi state botanical reserve (2267 ha), includes
Svarytsevytskyi state botanical reserve (2220 ha), included in Emerald Site “Dubrovytskyi”.
Threats: –.
General description. Wooded raised bogs and transiional mires with smaller areas of mires
without the tree layer. Dominants: Pinus sylvestris, Betula pubescens, Carex lasiocarpa,
Eriophorym vaginatum, Ledum palustre, Vaccinium oxycoccos (Oxycoccus palustris),
Phragmites australis, Sphagnum cuspidatum, Sphagnum fallax, Sphagnum magellanicum,
Sphagnum fuscum.
Botanical significance. The largest area of wooded oligotrophic and mesotrophic mires with
Pinus sylvestris and Betula pubescens in Ukraine.
Criterion C
• G1.51 Sphagnum Betula woods; area: 600 ha; trend: stable; area data quality: poor;
trend data quality: poor.
• G3.E Nemoral bog conifer woodland; area: 1000 ha; trend: stable; area data quality:
poor; trend data quality: poor.
• X04 Raised bog complexes; area: 40 ha; trend: stable; area data quality: poor; trend
data quality: poor.
Literature
1. Фіторізноманіття Українського Полісся та його охорона / Під заг. ред.
Т.Л. Андрієнко. – Київ: Фітосоціоцентр, 2006. – 316 с.
Land use: agriculture (animals) – minor; nature conservaion and research – major.
Protected areas: includes Mykhailivskyi Step regional landscape reserve (1343.1 ha),
included in Emerald Site "Mykhailivskyi Steppe".
Threats: –.
General description. Grasslands dominated by Festuca valesiaca, Sipa capillata, S,
lessingiana, S. pulcherrima, S. ucrainica, Caragana frutex, Amygdalus nana, Koeleria
cristata, Poa angusifolia. On stony calcareous soil, there prevail Sipa asperella, Thymus
Mykhailivskyi Step
V.A. Onyshchenko, O.M. Derkach
Ukrainian name: Михайлівський степ.
Transliteraion/Translaion variants: Myhaylivskyi Steppe.
Area: 1774.0 ha.
Altitude: 12–89 m.
Latitude: 47°24'51" N (47.4140°).
Longitude: 31°37'43" E (31.6284°).
Administrative regions. Mykolaiv region: Voznesensk raion, Nova Odesa raion.
Ownership: state.
Biogeographic regions: steppic.
Habitats. Level 1. E – 95.9%; F – 0.3%; G – 3.3%; H – 0.5%.
Habitats. Level 2. E1 Dry grasslands – 95.0%; E2 Mesic grasslands – 0.9%; F3 Temperate
and mediterranean-montane scrub – 0.3%; G1 Broadleaved deciduous woodland – 3.3%;
H2 Screes – 0.2%; H3 Inland cliffs, rock pavements and outcrops – 0.2%; H5 Miscellaneous
inland habitats with very sparse or no vegetation – 0.1%.
Futher habitat description. E1.1 Inland sand and rock with open vegetation; E1.2 Perennial
calcareous grassland and basic steppes; F3.2 Submediterranean deciduous thickets and
brushes; G1.7 Thermophilous deciduous woodland; G1.C Highly ariicial broadleaved
deciduous forestry plantaions; H2.6 Calcareous and ultra-basic screes of warm exposures;
H3.2 Basic and ultra-basic inland clifs; H5.6 Trampled areas.
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dimorphus, Koeleria brevis, Jurinea brachycephala with Allium lavescens, Astragalus
albidus, A. ucrainicus, A. corniculatus, Centaurea marschalliana, Dianthus pseudoarmeria,
Genista scythica, Gypsophila collina, Hedysarum grandilorum, Linaria macroura, Linum
linearifolium, Pimpinella itanophila, Teucrium polium.
Botanical significance. One of the best steppe areas west to the Dnipro river.
Criterion A
• Chamaecyisus graniicus (Rehmann.) Rothm.; A(iii); abundance: occasional; trend:
stable; species data quality: poor; trend data quality: poor.
• Crambe tataria Sebeok.; A(ii); abundance: frequent; trend: unknown; species data
quality: medium; trend data quality: poor.
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•
Gymnospermium odessanum (DC.) Takht.; A(iv); abundance: unknown; trend: unknown;
species data quality: poor; trend data quality: poor.
• Tulipa hypanica Klokov et Zoz; A(iii); abundance: occasional; trend: stable; species data
quality: poor; trend data quality: poor.
Criterion C
• E1.2 Perennial calcareous grassland and basic steppes; area: 1700 ha; trend: stable;
area data quality: good; trend data quality: good.
Conservaion proposals. Include in Yelanetskyi nature reserve.
Literature
1. Південно-Бузький меридіональний екологічний коридор: стислий огляд
біорізноманіття та найцінніші території / під ред. В. Костюшина. – К., 2007. – 92 с.
Literature
1.Деркач О.М. Ключові ботанічні території Миколаївщини: сучасний стан та
проблеми збереження // Теорія і практика заповідної справи в Україні. Зб.наук.праць.
– К., 2005. – С.167–173.
2.Коломієць Г.В. Перлинні волошки секції Pseudophalolepis Klok. ряду Margaritacea
Klok. Питання систематики та охорони // Укр. фітоценологічний зб. – Вип. 1–2 (12–13).
– К.: Фітосоціоцентр, 1999. – С. 165–169.
Mykhailo-Laryne
O.M.Derkach, H.V. Kolomiets
Ukrainian name: Михайло-Ларине.
Area: 39.6 ha.
Altitude: 10–45 m.
Latitude: 47°08'14" N (47.1373°).
Longitude: 32°13'00" E (32.2165°).
Administrative regions. Mykolaiv region: Vitovka (Zhovtnevyi) raion.
Ownership: state.
Biogeographic regions: steppic.
Habitats. Level 1. E – 30.0%; G – 69.0%; H – 1.0%.
Habitats. Level 2. E1 Dry grasslands – 30.0%; G3 Coniferous woodland – 69.0%; H5
Miscellaneous inland habitats with very sparse or no vegetaion – 1.0%.
Futher habitat description. E1.9 Open non-Mediterranean dry acid and neutral grassland,
including inland dune grassland (× E1.1); G3.F Highly artificial coniferous plantations; H5.6
Trampled areas.
Land use: forestry – major.
Protected areas: includes local botanical reserve "Mykhailo-Larynskyi" (14.8 ha)
Threats: forestry (aforestaion) – medium.
General description. A part of sand terrace above the loodplain of the Inhul river. Forest
plantaions of Pinus pallasiana and Pinus sylvestris in comlex with dry sand grasslands.
Dominant species of the grasslands are Carex colchica, Artemisia marschalliana. Other
typical species are Anchusa gmelinii, Centaurea adpressa, Centaurea margarita-alba,
Dianthus membranaceus, Euphorbia seguieriana, Gypsophila paniculata, Helichrysum
arenarium, Linaria genisifolia, Scabiosa ucrainica, Sipa borysthenica, Thymus pallasianus.
Botanical significance. 40% of global populaion of narrow endemic species Centaurea
margarita-alba s. str.
Criterion A
• Centaurea margarita-alba Klokov; A(iii); abundance: frequent; trend: unknown; species
data quality: good; trend data quality: poor.
Conservaion proposals. Remove trees. Create a state botanical reserve instead of local
botanical reserve "Mykhailo-Larynskyi" including the enire IPA. Restore sand grasslands in
adjacent area.
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3. Крицька Л.І., Деркач О.М. Сучасний стан популяцій видів ряду Margaritaceae Klok.
(Centaurea L.) // Укр. ботан. журн. – 1991. – Т. 48, № 3. – С. 78–80.
4. Крицька Л.І., Деркач О.М., Собко В.Г. Волошка білоперлинна Centaurea margaritaalba Klokov / Червона книга України. Рослинний світ / за ред. Я.П. Дідуха. – К.:
Глобалконсалтинг, 2009. – С. 305.
5.Перлини піщаної флори у пониззях Південного Бугу та Інгулу. Cерія: Збереження
біорізноманіття в Приморсько-степовому екокоридорі / Під ред. Г.В. Коломієць. – К.:
Громадська організація "Веселий Дельфін", 2008. – 60 c.
Mys Marian
Ukrainian name: Мис Март'ян.
Transliteraion/Translaion variants: Cape Marian, Cape Martyan.
Area: 128.0 ha.
Altitude: 0–240 m.
203
V.A. Onyshchenko
Latitude: 44°30'40" N (44.5110°).
Longitude: 34°14'56" E (34.2487°).
Administrative regions. Autonom. Republic of Crimea: Yalta city; Territ. waters of Ukraine.
Ownership: state.
Biogeographic regions: steppic.
Habitats. Level 1. A – 7.0%; B – 1.0%; E – 0.1%; G – 91.4%; H – 0.5%.
Habitats. Level 2. A1 Littoral rock and other hard substrata – 0.1%; A2 Littoral sediment –
0.9%; A3 Infralittoral rock and other hard substrata – 3.0%; A5 Sublittoral sediment – 3.0%;
B2 Coastal shingle – 0.8%; B3 Rock cliffs, ledges and shores, including the supralittoral
– 0.2%; E1 Dry grasslands – 0.1%; G1 Broadleaved deciduous woodland – 33.2%; G3
Coniferous woodland – 35.0%; G4 Mixed deciduous and coniferous woodland – 23.2%;
H2 Screes – 0.1%; H3 Inland clifs, rock pavements and outcrops – 0.2%; H5 Miscellaneous
inland habitats with very sparse or no vegetation – 0.2%.
Futher habitat description. A2.1 Litoral coarse sediment; B2.1 Shingle beach dritlines;
B2.2 Unvegetated mobile shingle beaches above the dritline; B2.3 Upper shingle beaches
with open vegetaion; B3.1 Supralitoral rock (lichen or splash zone); B3.2 Unvegetated
rock clifs, ledges, shores and islets; B3.3 Rock clifs, ledges and shores, with angiosperms;
B3.4 Sot sea-clifs, oten vegetated; E1.1 Inland sand and rock with open vegetation; E1.3
Mediterranean xeric grassland; G1.7 Thermophilous deciduous woodland; G3.5 Pinus
nigra woodland; G3.9 Coniferous woodland dominated by Cupressaceae or Taxaceae; H2.6
Calcareous and ultra-basic screes of warm exposures; H3.2 Basic and ultra-basic inland
clifs; H5.6 Trampled areas.
Land use: nature conservaion and research – 98%; tourism/recreaion – minor.
Protected areas: overlaps (about 126 ha) with Mys Marian nature reserve, overlaps (94
ha) with Emerald Site "Mys Marian".
Threats: burning of vegetaion – medium, development (recreaion/tourism) – low.
General description. The IPA is situated in the southern part of the Crimea. It includes a
slope near the sea with woods at alitudes up to 240 m and 50 m wide strip of the sea.
The largest area is occupied by broadleaved woods dominated by Quercus pubescens with
admixture of Arbutus andrachne, Fraxinus angusifolia, Juniperus excelsa, Pinus pallasiana.
The shrub layer is dominated by Carpinus orientalis, Cornus mas, Juniperus oxycedrus, the
lower shrub sublayer consists of Ruscus ponicus. In the herb layer, there are Achnatherum
bromoides, Aegonychon purpureocaeruleum, Brachypodium rupestre, Carex cuspidata,
Carex hallerana, Elytrigia nodosa, Festuca valesiaca. Juniperus excelsa woods also occupy
a large area. Their shrub layer is composed of Juniperus oxycedrus (predominantly),
Bupleurum fruicosum, Carpinus orientalis, Cornus mas, Paliurus spina-chrisi and lower
shrubs such as Cistus tauricus, Hippocrepis emeroides, Jasminum fruicans, Ruscus ponicus.
In the herb layer, there prevail Achnatherum bromoides, Carex cuspidata, Carex hallerana,
Elytrigia nodosa, Fumana procumbens, Helianthemum canum, Helianthemum stevenii,
Teucrium chamaedrys, Teucrium polium, Thymus roegneri. Smaller area is covered by
Pinus pallasiana forest with dominance of Asperula stevenii, Brachypodium rupestre, Carex
hallerana, Dorycnium herbaceum, Galium mollugo, Laser trilobum, Teucrium chamaedrys.
Substrate types of the beach are unmobile rock and shingle. In the sea, there dominate
Cystoseira crinita, Cystoseira barbata, Zostera marina, Zostera nolii (Nanozostera nolii).
Considerable areas are occupied by screes and outcrops. Litoral habitats are represented
by shingle beach and rocks.
Botanical significance. Important for conservaion of Quercus pubescens woods, Juniperus
excelsa woods, Brassica taurica.
Criterion A
• Brassica taurica (Tzvelev) Tzvelev (Brassica sylvestris Mill. subsp. taurica Tzvel.); A(i),
A(ii); abundance: 30 individuals; trend: luctuaing; species data quality: good; trend
data quality: good.
• Ophrys oestrifera M.Bieb.; A(ii); abundance: 70 individuals; trend: stable; species data
quality: good; trend data quality: medium.
Criterion B
• G1 Broadleaved deciduous woodland; area 75 ha; % of indicator species: 19.5%; No of
indicator species: 15; trend: increasing; species data quality: good; area data quality:
good; trend data quality: medium.
Criterion C
• G3.5 Pinus pallasiana and Pinus banaica forests; area: 5 ha; trend: stable; area data
quality: medium; trend data quality: medium.
• G3.9 Coniferous woodland dominated by Cupressaceae or Taxaceae; area: 40 ha; trend:
stable; area data quality: medium; trend data quality: medium.
Literature
1. Белич Т.В. К изучению бриофлоры заповедника "Мыс Мартьян" // Заповедники
Крыма. Заповедное дело, биоразнообразие, экообразование: Матер. III научн. конф.
(22 апреля 2005 г., Симферополь, Крым). – Ч. I. – Симферополь, 2005. – С. 141–144.
2. Голубева И.В., Крайнюк Е.С. Аннотированный каталог высших растений
заповедника "Мыс Мартьян". – Ялта, ГНБС. – 1987. – 40 с.
3. Шеляг-Сосонко Ю.Р., Дидух Я.П., Молчанов Е.Ф. Государственный заповедник
"Мыс Мартьян". – Киев: Наук. думка, 1985. – 260 с.
4. Григоров А.Н. Естественное возобновление и возрастная структура насаждений
можжевельника высокого в заповеднике "Мыс Мартьян" // Труды Никит. ботан. сада.
– 1982. – Т. 86. – С. 35–44.
5. Ена А.В. Современное состояние крымских популяций земляничника
мелкоплодного // Природоохранные аспекты изучения Горного Крыма. – Симферополь,
204
205
1986. – С. 26–30.
6. Крайнюк Е.С. Редкие виды высших растений в заповеднике "Мыс Мартьян" //
Бюл. Никит. ботан. сада. – 1988. – Вып. 67. – С. 20–25.
7. Крайнюк Е.С. Современное состояние раритетного фитофонда заповедника
"Мыс Мартьян" // Труды Никит. ботан. сада. – 2001. – Т. 120. – С. 63–73.
8. Крайнюк Е.С. Растения Красной книги Украины в природном заповеднике "Мыс
Мартьян" // рослинний світ у Червоній книзі України: впровадження Глобальной
стратегії збереження рослин. Матеріали міжнар. конф. (11 – 15 жовтня 2010 р.), Київ. –
Київ: Альтерпрес, 2010. – С. 274–277.
9. Крайнюк К.С., Маслов І.І. ПЗ Мис Мартьян // Фіторізноманіття заповідників
і національних природних парків України. Ч. 1. Біосферні заповідники. Природні
заповідники / під ред. В.А. Онищенка і Т.Л. Андрієнко. – Київ: Фітосоціоцентр, 2012. –
С. 277–290.
10. Куликов Г.В., Лялин Г.С. Земляничник мелкоплодный в заповеднике "Мыс
Мартьян" // Бюл. Главн. ботан. сада. – 1975. – Вып. 98. – С. 59–63.
11. Ларина Т. Г. Флора и растительность заповедника "Мыс Мартьян" // Труды
Гос. Никит. ботан. сада. – 1976. – Т. 70. – С. 45–62.
12. Лукс Ю.А. Флора орхидных заповедника "Мыс Мартьян" // Труды Гос. Никит.
ботан. сада. – 1976. – Т. 70. – С. 95–104.
13. Маслов И.И. Фитобентос псевдолиторального пояса района Ялты //
Природные экосистемы Южного берега Крыма и их охрана. Сборник науч. тр. – Ялта:
ГНБС. – 1984. – Т. 94. – С. 72–87.
14. Маслов И.И., Куропатов Л.А. К детальному описанию биоценоза цистозиры
заповедника "Мыс Мартьян". – Бюлл. Гос. Никит. ботан. сада. – 1987. – Вып .63. – С.
13–17.
15. Маслов И.И., Саркина И.С., Белич Т.В., Садогурский С.Е. Аннотированный
каталог водорослей и грибов заповедника "Мыс Мартьян". – Ялта, ГНБС. – 1998. – 31 с.
16. Молчанов Е.Ф., Григоров А.Н., Голубева И.В., Ларина Т.Г., Щербатюк Л.К.,
Ругузов И.А., Склонная Л.У., Бескаравайный М.М. Высокоможжевеловые леса Крыма и
проблема их охраны // Гос. Никитск. ботан. сад. – Ялта, 1992. – Деп. В ВИНИТИ 30.12.92.
№ 3706 – В 92. – 296 с.
17. Погребняк И.И., Маслов И.И. К изучению донной растительности района
мыса Мартьян // Труды Гос. Никит. ботан. сада. – 1976. – Т. 70. – С. 105–113.
18. Ходосовцев А.Е., Редченко А.А. Аннотированный список лишайников
заповедника "Мыс Мартьян" (Украина) // Укр. ботан. журн. – 2002. – 59, № 1. – С. 64–
71.
19. Червона книга України. Рослинний світ / за ред. Я.П. Дідуха. – К.: Глобалконсалтинг, 2009. – 900 с.
Mytrofanivskyi Pivostriv
V.P. Kolomiychuk, V.A. Onyshchenko
Ukrainian name: Митрофанівський півострів.
Transliteraion/Translaion variants: Mytrofanivsky Peninsula.
Area: 817.1 ha.
Altitude: 0–19 m
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Latitude: 46°08'27" N (46.1407°).
Longitude: 34°07'05" E (34.1181°).
Administrative regions. Kherson region: Novotroitske raion .
Ownership: state, private.
Biogeographic regions: steppic.
Habitats. Level 1. A – 16.5%; E – 83.5%.
Habitats. Level 2. A2 Littoral sediment – 11.0%; A5 Sublittoral sediment – 5.5%; E1 Dry
grasslands – 40.8%; E6 Salt steppes – 42.7%.
Futher habitat description. A2.3 Littoral mud; A2.5 Coastal saltmarshes and saline
reedbeds; A5.3 Subittoral mud; E1.2 Perennial calcareous grassland and basic steppes; E6.2
Continental inland salt steppes.
Land use: nature conservaion and research – minor.
Protected areas: included in Ramsar Site "Central Syvash", overlaps (about 218 ha) with
Azovo-Syvaskyi naional nature park and Emerald Site "Azovo-Syvaskyi Naional Nature
Park".
Threats: agricultural intensification/expansion (arable) – low, burning of vegetaion – low.
General description. Northern shore of the Central Syvash Bay (saline lagoon of the Sea
of Azov). The IPA includes areas with the steppe vegaion, litoral with the halophyic
vegetaion and shallow water of the bay. The steppe vegetaion is dominated by Agropyron
pecinatum, Artemisia taurica, Festuca valesiaca, Sipa lessingiana, Sipa capillata,
Crinitaria villosa. Other typical species are Achillea setacea, Astragalus asper, Astragalus
reduncus, Limonium platyphyllum, Linaria macroura, Medicago romanica, Phlomis
pungens, Potenilla obscura, Prangos odontalgica. Dominants of the halophyic vegetaion
are Halocnemum strobilaceum, Puccinelia fominii, Petrosimonia brachiata, Limonium
sufruicosum. A part of the steppe vegetaion is thretened by plowing.
Botanical significance. The area includes large parts of global populaions of endemic
species Goniolimon rubellum and Lepidium syvaschicum.
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Criterion A
• Goniolimon rubellum (S.G.Gmel.) Klokov; A(iii); abundance: occasional; trend: stable;
species data quality: medium; trend data quality: poor.
• Lepidium syvaschicum Kleopow; A(iii); abundance: occasional; trend: stable; species
data quality: medium; trend data quality: poor.
Conservaion proposals. Include in Azovo-Syvaskyi naional nature park.
Literature
1. Коломійчук В.П. Сучасне поширення видів судинних рослин описаних
Ю.Д. Клеоповим з Присивашшя // Ю.Д. Клеопов та сучасна ботанічна наука. Мат-ли
читань, присвячені 100-річчю з дня народження Ю.Д. Клеопова (Київ, 10–13 листопада
2002 р.). – Київ: Фітосоціоцентр, 2002. – С. 48–51.
2. Павлов В.В. Північне Присивашшя як елемент Азово-Чорноморського
екокоридору // Укр. ботан. журн. – 2002. – 59, №1. – С. 89–93.
lessingiana. Frequent species are Salvia nutans, Plantago stepposa, Trifolium montanum,
Achillea nobilis, Artemisia austriaca, Echium rubrum, Thalictrum minus. Steppic shrub
communiies are dominated by Caragana frutex, Amygdalus nana, Spiraea hypericifolia,
Cotoneaster melanocarpa. Shale outcrops are characterized by the presence of the local
endemic species Scrophularia donetzica. Woods with dominance of Quercus robur, Fraxinus
excelior, Tilia cordata and Ulmus laevis are in valleys and gulches.
Naholnyi Kriazh
V.A. Onyshchenko, M.M. Peregrym
Ukrainian name: Нагольний кряж.
Transliteraion/Translaion variants: Nagol'nyi Ridge.
Area: 3835.0 ha.
Altitude: 140–249 m.
Latitude: 47°58'48" N (47.9801°).
Longitude: 39°29'59" E (39.4996°).
Administrative regions. Luhansk region: Dovzansk (Sverdlovsk) raion.
Ownership: state.
Biogeographic regions: steppic.
Habitats. Level 1. E – 92.4%; F – 0.5%; G – 1.8%; H – 1.0%; I – 4.3%.
Habitats. Level 2. E1 Dry grasslands – 91.4%; E2 Mesic grasslands – 1.0%; F3 Temperate and
mediterranean-montane scrub – 0.5%; G1 Broadleaved deciduous woodland – 1.8%; H3
Inland cliffs, rock pavements and outcrops – 0.1%; H5 Miscellaneous inland habitats with
very sparse or no vegetation – 0.9%; I1 Arable land and market gardens – 4.3%.
Futher habitat description. E1.2 Perennial calcareous grassland and basic steppes; F3.2
Submediterranean deciduous thickets and brushes; I1.5 Bare illed, fallow or recently
abandoned arable land; G1.C Highly artificial broadleaved deciduous forestry plantations;
H2.6 Calcareous and ultra-basic screes of warm exposures; H3.2 Basic and ultra-basic inland
cliffs; H5.6 Trampled areas.
Land use: agriculture (animals) – major; agriculture (arable) – minor; extracion (minerals)
– minor; nature conservaion and research – minor.
Protected areas: overlaps (about 434 ha) with regional landscape reserve "Naholnyi Kriazh",
included in Emerald Site "Naholny Kriazh".
Threats: agricultural intensification/expansion (arable) – medium, agricultural
intensiicaion/expansion (grazing) – low, extracion (minerals/quarries) – low, forestry
(aforestaion) – low.
General description. Dry grasslands with small areas of sandstone and shale outcrops,
shrubs and deciduous woods. Dominants of the steppe vegetaion are Festuca valesiaca,
Bothriochloa ischaemum, Agropyron pectinatum, Poa angusifolia, Sipa capillata, S.
208
Botanical signiicance. A large area of the steppe vegetaion.
Criterion A
• Hyacinthella pallasiana (Steven) Losinsk.; A(iv); abundance: occasional; trend: unknown;
species data quality: poor; trend data quality: poor.
Criterion C
• E1.2 Perennial calcareous grassland and basic steppes; area: 2800 ha; trend: unknown;
area data quality: good; trend data quality: poor
Conservaion proposals. Do not allow plowing. Create a state botanical reserve including
the enire IPA.
Literature
1. Природно-заповідний фонд Луганської області // О.А. Арапов (заг. ред.), Т.B. Сова,
О.А. Cавенко, В.Б. Фєрєнц, Н.У. Кравець, Л.Л. Зятьков, Л.О. Морозова. Довідник. – 3-е
вид., доп. і перероб. – Луганськ: "Луганська правда", 2013. – 224 с.
209
Novobila
V.A. Onyshchenko, M.V. Banik
Ukrainian name: Новобіла.
Area: 2469.0 ha.
Biogeographic regions: steppic.
Habitats. Level 1. D – 4.3%; E – 77.2%; F – 2.0%; G – 9.4%; H – 5.1%; I – 2.0%.
Habitats. Level 2. D5 Sedge and reedbeds, normally without free-standing water – 4.3%;
E1 Dry grasslands – 76.6%; E2 Mesic grasslands – 0.6%; F3 Temperate and mediterraneanmontane scrub – 2.0%; G1 Broadleaved deciduous woodland – 9.4%; H2 Screes – 5.1%; I1
Arable land and market gardens – 2.0%.
Futher habitat description. E1.1 Inland sand and rock with open vegetation; E1.2 Perennial
calcareous grassland and basic steppes; F3.2 Submediterranean deciduous thickets and
brushes; G1.7 Thermophilous deciduous woodland; G1.A Meso- and eutrophic Quercus,
Carpinus, Fraxinus, Acer, Tilia, Ulmus and related woodland; G1.C Highly artificial
broadleaved deciduous forestry plantations; H2.6 Calcareous and ultra-basic screes of
warm exposures; I1.5 Bare illed, fallow or recently abandoned arable land.
Land use: agriculture (animals) – major; forestry – minor; nature conservaion and research
– major.
Protected areas: overlaps (>2000 ha) with Novobila regional botanical nature monument,
overlaps (2214 ha) with Emerald Site "Novobilskyi".
Threats: abandonment/reducion of land management – low; agricultural intensification/
expansion (arable) – low, extracion (minerals/quarries) – low; forestry (aforestaion) –
low.
General description. Slopes with the steppe vegetaion, outcrops of chalk and oak woods;
small areas of reedbeds and hay meadows. There are many threatened species of steppes
and chalk outcrops.
Botanical significance. The largest locality of Hedysarum ucrainicum in Ukraine. Important
area for conservaion of the steppes and chalk outcrops with endemic species.
Criterion A
• Hedysarum ucrainicum Kaschm.; A(iv); abundance: frequent; trend: stable; species data
quality: good; trend data quality: medium.
Criterion C
• E1.13 Coninental dry rocky steppic grasslands and dwarf scrub on chalk outcrops;
area: 135 ha; trend: stable; area data quality: medium; trend data quality: poor.
• E1.2 Perennial calcareous grassland and basic steppes; area: 2000 ha; trend: stable;
area data quality: good; trend data quality: poor.
Literature
1. Banik M. Daphne'04. Final report. – Kharkiv, 2006. – 20 p. (http://www.rufford.org/
files/127.01.04%20Detailed%Final%20Report.pdf).
Novohredneve
Altitude: 70–160 m.
Latitude: 49°47'03" N (49.7841°).
Longitude: 39°09'34" E (39.1595°).
Administrative regions. Luhansk region: Novopskov raion.
Ownership: state.
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Ukrainian name: Новогредневе.
Transliteraion/Translaion variants: Novogredneve.
Area: 16.2 ha.
Altitude: 7–23 m.
Latitude: 47°09'41" N (47.1613°).
Longitude: 32°59'52" E (32.9978°).
Administrative regions. Kherson region: Velyka Oleksandrivka raion.
211
V.P. Kolomiychuk
Ownership: state.
Biogeographic regions: steppic.
Habitats. Level 1. E – 76.0%; G – 20.0%; H – 2.0%; J – 2.0%.
Habitats. Level 2. E1 Dry grasslands – 76.0%; G1 Broadleaved deciduous woodland – 19.0%;
G3 Coniferous woodland – 1.0%; H5 Miscellaneous inland habitats with very sparse or no
vegetaion – 2.0%; J3 Acive opencast mineral extracion sites, including quarries – 2.0%.
Futher habitat description. E1.9 Open non-Mediterranean dry acid and neutral grassland,
including inland dune grassland (× E1.1); G1.C Highly artificial broadleaved deciduous
forestry plantations; H5.6 Trampled areas; J3.2 Active opencast mineral extraction sites,
including quarries.
Land use: agriculture (animals) – major, extracion (minerals) – minor, forestry – major.
Protected areas: –.
Threats: agricultural intensiicaion/expansion (grazing) – low, burning of vegetaion – low,
Robinia pseudoacacia, Pinus pallasiana, Ulmus minor. Illegal sand quarry occupies a small
part of the site.
Botanical significance. Important area for narrow endemic species Centaurea paczoskii
(one of 2 localiies).
Criterion A
• Centaurea paczoskii Kotov ex Klokov; A(iii); abundance: occasional; trend: unknown;
species data quality: good; trend data quality: poor.
Conservaion proposals. Remove trees. Create a state botanical reserve and an Emerald Site
including the enire IPA.
Literature
1. Крицька Л.І., Деркач О.М. Сучасний стан популяцій видів ряду Margaritaceae Klok.
(Centaurea L.) // Укр. ботан. журн. – 1991. – 48, № 3. – С. 78 – 80.
2. Мойсієнко І.І., Мельник Р.М. Волошка Пачоського Centaurea paczoskii Klokov (C.
margaritacea Ten. subsp. paczoskii (Kotov et Klokov) Dostál) / Червона книга України.
рослинний світ / за ред. Я.П. Дідуха. – К.: Глобалконсалтинг, 2009. – С. 306.
Nyzhnii Dnipro
extracion (mineral/quarries) – medium, forestry (aforestaion) – low.
General description. Grassland on the sand terrace of the Inhulets river. Dominant species
are Carex colchica, Festuca beckeri, Artemisia campestris. Other frequent species are
Eragrostis minor, Chondrilla juncea, Dianthus platyodon, Jurinea sp., Helichrysum arenarium,
Koeleria glauca s.l., Plantago scabra. Moss layer consists of Tortula ruralis (cover 3–10%).
Populaion of Centaurea paczoskii occupies 2–3 ha. A part of the area is ariicial wood with
212
V.A. Onyshchenko
Ukrainian name: Нижній Дніпро.
Transliteraion/Translaion variants: Lower Dnipro, Lower Reaches of the Dnieper.
Area: 43083.0 ha.
Altitude: 0–13 m.
Latitude: 46°36'01" N (46.6004°).
Longitude: 32°38'46" E (32.6461°).
Administrative regions. Kherson region: Beryslav raion, Biloozerka raion, Hola Prystan raion,
Oleshky (Tsiurupynsk) raion, Kherson city; Nova Kakhovka city .
Ownership: state.
Biogeographic regions: steppic.
Habitats. Level 1. C – 30.4%; D – 55.5%; E – 4.2%; F – 0.4%; G – 9.5%.
Habitats. Level 2. C1 Surface standing waters – 14.5%; C2 Surface running waters – 13.8%;
C3 Littoral zone of inland surface waterbodies – 2.1%; D5 Sedge and reedbeds, normally
without free-standing water – 55.5%; E3 Seasonally wet and wet grasslands – 3.9%; E6
Inland salt steppes – 0.3%; F9 Riverine and fen scrubs – 0.4%; G1 Broadleaved deciduous
woodland – 8.3%; G3 Coniferous woodland – 1.2%.
Futher habitat descripion. C1.2 Permanent mesotrophic lakes, ponds and pools; C2.3
Permanent non-idal, smooth-lowing watercourses; C3.2 Water-fringing reedbeds and tall
helophytes other than canes; D5.1 Reedbeds normally without free-standing water; D5.2
Beds of large sedges normally without free-standing water; E1.9 Open non-Mediterranean
dry acid and neutral grassland including inland dune grassland (× E1.1); E3.4 Moist or wet
eutrophic and mesotrophic grassland; E6.2 Coninental inland salt steppes; F9.1 Riverine
scrub; G1.1 Riparian and gallery woodland, with dominant Alnus, Betula, Populus or Salix;
G1.3 Mediterranean riparian woodland; G3.F Highly ariicial coniferous plantaions.
Land use: forestry – 10%; mowing/hay making – minor, urban/industrial/transport – minor,
tourism/recreaion – minor.
Protected areas: overlaps with Nyzhniodniprovskyi naional nature park (about 40000 ha),
213
overlaps (39039 ha) with Emerald Site "Lower Dnipro", overlaps (3823 ha) with Emerald Site
"Dniprovsko-Buzkyi Lyman", overlaps (123 ha) with Emerald Site "Kinburnska Kosa".
Threats: development (recreaion/tourism) – low, development (urbanizaion) – low.
General descripion. Floodplain of the Dnipro river from Nova Kakhovka to the mouth.
Phragmites australis dominated marshes occupy the largest area. Other important
dominants are Typha angusifolia, Carex acuiformis, Carex elata. Large areas are occupied
by aquaic vegetaion dominated by Ceratophyllum demersum, Lemna minor, Myriophyllum
•
C2.34 Eutrophic vegetation of slow-flowing rivers; area: 300 ha; trend: stable; area data
quality: poor; trend data quality: medium.
• D5.2 Beds of large sedges normally without free-standing water; area: 400 ha; trend:
stable; area data quality: poor; trend data quality: poor.
Literature
1. Дубына Д.В., Шеляг-Сосонко Ю.Р. Плавни Причерноморья. – К.: Наук. думка,
1989. – 272 с.
2. Червона книга України. Рослинний світ / за ред. Я.П. Дідуха. – К.: Глобалконсалтинг,
2009. – 900 с.
3. Чинкіна Т.Б. Сучасний стан і завдання охорони рослинного покриву гирлової
області Дніпра // Природничий альманах. Сер. біол. науки, 2002. – Вип. 2(3). –
С. 240–247.
4. Чинкіна Т. Синтаксономічна схема заплавної рослинності гирлової ділянки
Дніпра // Вісник Львівського університету. Сер. біол., 2006. – 42. – С. 32–37.
Nyzhniodniprovski Pisky
vericillatum, Myriophllum spicatum, Nuphar lutea, Nymphaea alba, Potamogeton
pecinatus (Stuckenia pecinata), Potamogeton perfoliatus, Potamogeton crispus,
Potamogeton berchtoldii, Salvinia natans, Spirodela polyrrhiza. Woods cover about 4000
ha. Main dominant is Salix alba, other important species – Salix fragilis, Populus alba,
Populus nigra. Shrub vegetaion is represented mainly by communiies of Salix cinerea,
Salix triandra, Salix acuifolia.
Botanical significance. This area is one of the richest in aquaic vascular plant species in
Ukraine.
Criterion A
• Carex secalina Wahlenb.; A(ii); abundance: occasional; trend: stable; species data
quality: medium; trend data quality: poor.
• Salvinia natans (L.) All.; A(ii); abundance: frequent; trend: stable; species data quality:
good; trend data quality: medium.
• Trapa natans L.; A(ii); abundance: occasional; trend: stable; species data quality:
medium; trend data quality: medium.
Criterion C
• C1.224 Floaing Utricularia australis and Utricularia vulgaris colonies; area: 3 ha; trend:
stable; area data quality: poor; trend data quality: medium.
• C1.225 Floaing Salvinia natans mats; area: 10 ha; trend: stable; area data quality: poor;
trend data quality: medium.
• C1.3411 Ranunculus communities in shallow water; area: 1 ha; trend: unknown; area
data quality: poor; trend data quality: poor.
V.A. Onyshchenko
Ukrainian name: Нижньодніпровські піски.
Transliteraion/Translaion variants: Lower Dnipro Sands.
Area: 10388.0 ha.
Altitude: 0–10 m.
Latitude: 46°27'21" N (46.4559°).
Longitude: 31°58'28" E (31.9744°).
Administrative regions. Kherson region: Hola Prystan raion; Mykolaiv region: Ochakiv raion.
Ownership: state.
Biogeographic regions: steppic.
Habitats. Level 1. A – 1.6%; C – 16.0%; D – 1.6%; E – 67.8%; F – 1.0%; G – 10.0%; H – 2.0%.
Habitats. Level 2. A2 Littoral sediment – 0.2%; A5 Sublittoral sediment – 1.4%; C1 Surface
standing waters – 16.0%; D5 Sedge and reedbeds normally without free-standing water
– 1.5%; D6 Inland saline and brackish marshes and reedbeds – 0.1%; E1 Dry grasslands
– 58.7%; E2 Mesic grasslands – 5.4%; E3 Seasonally wet and wet grasslands – 2.3%; E6
Inland salt steppes – 1.4%; F3 Temperate and mediterranean-mountane scrub – 1.0%; G1
Broadleaved deciduous woodland – 10.0%; H5 Miscellaneous inland habitats with very
sparse or no vegetation – 2.0%.
Futher habitat description. E1.2 Perennial calcareous grassland and basic steppes, E1.9
Open non-Mediterranean dry acid and neutral grassland including inland dune grassland,
(× E1.1) E2.5 Meadows of the steppe zone, E3.4 Moist or wet eutrophic and mesotrophic
grassland, F3.1 Temperate thickets and scrub; G1.7 Thermophilous deciduous woodland,
G1.9 Non-riverine woodland with Betula, Populus tremula or Sorbus aucuparia, H5.3
Sparsely- or un-vegetated habitats on mineral substrates not resulting from recent ice
activity.
Land use: nature conservation and research – major, agriculture (animals) – minor.
Protected areas: overlaps (5397 ha) with Chernomorskiy (Chornomorskyi, Black Sea)
biosphere reserve (UNESCO, core zone of the reserve), overlaps (5397 ha) with Chornomorskyi
biosphere reserve (naional category), overlaps (ca. 270 ha) with Biloberezhia Sviatoslava
214
215
naional nature park, overlaps with Emerald Sites "Black Sea Biosphere Reserve" (5397 ha),
"Biloberezhzhia Sviatoslava Naional Nature Park" (267 ha), "Kinburnska Kosa" (4724 ha).
Threats: forestry (afforestation) – low.
General description. Major habitat type is the dry sand grassland. Other habitats are
sublittoral of Kinburn (Yahorlytska) Bay, mesic grasslands, beds of Carex riparia and Carex
elata, halophytic vegetation, deciduous woods (small groves), freshwater and saline
permanent and temporary lakes. Sand grasslands are dominated by Festuca beckeri, Koeleria
sabuletorum, Agropyron lavrenkoanum, Stipa borysthenica, Artemisia marschalliana. Their
typical species are Achillea micrantha, Agropyron dasyanthum, Anchusa gmelinii, Asperula
graveolens, Dianthus platyodon, Helichrysum corymbiforme, Jacobaea borysthenica (Senecio
borysthenicus), Scabiosa ucrainica, Scorzonera ensifolia, Seseli tortuosum, Tragopogon
borysthenicus. Dominants of the mesic grasslands are Agrostis sabulicola, Calamagrosts
epigeios, Cynodon dactylon, Elytrigia elongata, Elytrigia repens, Phleum phleoides, Poa
angustifolia. Typical species: Achillea euxina, Allium guttatum, Campanula rapunculus,
Carex melanostachya, Centaurea adpressa, Festuca rupicola, Hypericum perforatum, Inula
sabuletorum, Linum perenne, Rumex acetosella, Rumex thyrsiflorus, Veronica steppacea.
Puccinellia gigantea dominates in saline grasslands. Other frequent halophyic specis are
Aeluropus littoralis, Juncus gerardii, Limonium meyeri, Scorzonera parviflora, Spergularia
marina. Woods are represented by groves of Quercus robur, Betula borysthenica, Populus
tremula. Quercus robur woods have the shrub layer (0.2–0.4) consisting of Rhamnus
catharctica, Sambucus nigra, Crataegus fallacina, Crataegus alutacea. The most frequent
species of the herb layer are Acillea euxina, Asparagus tenuifolius, Calamagrostis epigeios,
Carex elata, Convallaria majalis, Dactylis glomerara, Elytrigia repens, Phragmites australis,
Poa sylvicola, Polygonatum odoratum, Scilla bifolia, Viola odorata. The most frequent species
of Betula borysthenica woods: Agrostis sabulicola, Calamagrostis epigeios, Carex elata,
Frangula alnus, Genista borysthenica, Genista sibirica, Hieracium umbellatum, Juncus
atratus, Juncus conglomeratus, Phragmites australis, Rhamnus cathartica, Rubus caesius,
Salix rosmarinifolia, Solidago virgaurea. The most frequent species of Populus tremula
woods: Carex elata, Euphorbia semivillosa, Frangula alnus, Lysimachia vulgaris, Phalaroides
arundinacea, Rubus caesius, Sambucus nigra, Veronica longifolia. Littoral vegetation of
216
lakes is dominated by Phragmites australis, aquaic vegetaion of lakes by Lemna minor and
Lemna trisulca.
Botanical significance. The best area of sand steppe in Ukraine.
Criterion A
• Agropyron dasyanthum Ledeb.; A(i); abundance: frequent; trend: stable; species data
quality: medium; trend data quality: poor.
• Allium savranicum Besser; A(iv); abundance: rare; trend: unknown; species data quality:
medium; trend data quality: poor.
• Alyssum savranicum Andrz. ex Besser (Odontarrhena savranica (Andrz. ex Besser)
D.A.German); A(iv); abundance: rare; trend: unknown; species data quality: medium;
trend data quality: poor.
• Centaurea breviceps Iljin; A(iii); abundance: frequent; trend: unknown; species data
quality: medium; trend data quality: poor.
• Goniolimon graminifolium (Ait.) Boiss.; A(iv); abundance: occasional; trend: unknown;
species data quality: medium; trend data quality: poor
Criterion C
• E1.9 Open non-Mediterranean dry acid and neutral grassland, including inland dune
grassland; area: 6000 ha; trend: stable; area data quality: medium; trend data quality:
medium.
Conservaion proposals. Do not allow ariicial aforestaion.
Literature
1. Андрієнко Т.Л., Кофман І.Ш., Уманець О.Ю., Якушина Л.А. Розподіл рослинності
та її антропогенні зміни на Івано-Рибальчанській ділянці Чорноморського біосферного
заповідника // Укр. ботан. журн. – 1992. – 49, № 2. – С. 22–25.
2. Соломаха І.В., Воробйов Є.О., Мойсієнко І.І. Рослинний покрив лісів та чагарників
Північного Причорномор’я. – К.: Фітосоціоцентр, 2015. – 387 с.
3. Ткаченко В.С., Лисенко Г.М., Маяцький Г.Б., Уманець О.Ю. Структурні зміни
фітоценокомплексів Солоноозерної ділянки Чорноморського біосферного заповідника
за даними періодичного картографування // Укр. ботан. журн. – 1997. – 54, № 3. –
С. 232–239.
4. Ткаченко В.С., Уманець О.Ю. Фітоценотична характеристика Солоноозерної
ділянки Чорноморського біосферного заповедника (Херсонська область, Україна) //
Укр. ботан. журн. – 1993. – 50, № 2. – С. 14–23.
5. Уманець О.Ю. БЗ Чорноморський // Фіторізноманіття заповідників і національних
природних парків України. Ч. 1. Біосферні заповідники. Природні заповідники / під
ред. В.А. Онищенка і Т.Л. Андрієнко. – Київ: Фітосоціоцентр, 2012. – С. 73–93.
6. Уманець О.Ю., Соломаха І.В. Синтаксономія рослинності Чорноморського
біосферного заповідника. III. Ділянка Івано-Рибальчанська // Укр. фітоцен. зб. – Київ,
1999. – Сер. А, вип. 3 (14). – C. 84–102.
217
Obyichna Kosa
V.P. Kolomiychuk, V.A. Onyshchenko
Ukrainian name: Обитічна коса.
Transliteraion/Translaion variants: Obyichna Spit, Obitochna Spit.
Area: 4738.1 ha.
Altitude: 0–2 m.
Latitude: 46°34’10” N (46.5693°).
Longitude: 36°14’00” E (36.2332°).
Administrative regions. Zaporizhia region: Prymorsk raion. Territorial waters of Ukraine.
Ownership: state (major), private.
Biogeographic regions: steppic.
Habitats. Level 1. A– 49.1%; B – 1.7%; C – 9.7%; D – 0.5%; E – 36.7%; G – 2.3%.
Habitats. Level 2. A2 Littoral sediment – 5.3%; A5 Sublittoral sediment – 43.8%; B1 Coastal
dunes and sandy shores – 1.0%; B2 Coastal shingle – 0.7%; C1 Surface standing waters
218
– 9.7%; D6 Inland saline and brackish marshes and reedbeds – 0.5%; E1 Dry grasslands –
34.2%; E6 Salt steppes – 2.5%; G1 Broadleaved deciduous woodland – 2.3%.
Futher habitat description. A2.1 Littoral coarse sediment; A2.2 Littoral sand and muddy
sand; A2.4 Littoral mixed sediments; A2.5 Coastal saltmarshes and saline reedbeds;
B1.1 Sand beach driftlines; B1.2 Sand beaches above the driftline; B1.3 Shifting coastal
dunes; B1.4 Coastal stable dune grassland (grey dunes); B2.1 Shingle beach driftlines; B2.2
Unvegetated mobile shingle beaches above the driftline; C1.5 Permanent inland saline
and brackish lakes, ponds and pools; C1.6 Temporary lakes, ponds and pools; D6.2 Inland
saline or brackish species-poor helophyte beds normally without free-standing water; E1.2
Perennial calcareous grassland and basic steppes (E1.2G); E6.2 Coninental inland salt
steppes; G1.C Highly ariicial broadleaved deciduous forestry plantations.
Land use: forestry – minor; hunting – minor; mowing/hay making – 2%; nature conservation
and research – major.
Protected areas: included in Kosa Obyichna state landscape reserve, includes Ramsar Site
“Obytochna Spit and Obytocna Bay”, overlaps (about 4708 ha) with Emerald ste “Obyichna
Kosa Ta Zatoka”.
Threats: climate change/ sea level rise – low, development (recreaion/tourism) – low,
forestry (aforestaion) – low, natural events: spit erosion – medium
General description. A spit composed of shells and sand with adjacent sea. Major aquaic
species are Zostera marina, Zostera nolii (Nanozostera nolii), Potamogeton pecinatus
(Stuckenia pecinata), Zanichellia major. Litoral vegetaion of the Sea of Azov and saline
lakes is represented by halophyic communiies dominated by Phragmites australis, Juncus
mariimus, Salicornia perennans. On sand, there prevail Leymus sabulosus, Carex colchica,
Crambe mariima, Festuca beckeri, Medicago kotovii; other typical species are Artemisia
santonica, Centaurea adpressa, Centaurea odessana, Euphorbia seguierana.
Botanical significance. Important area for conservaion of sand steppes and brackish
aquaic vegetaion.
Criterion A
• Zostera marina L.; A(ii); abundance: frequent; trend: stable; species data quality:
medium; trend data quality: poor.
Criterion C
• B1.1 Sand beach driftlines; area: 6 ha; trend: stable; area data quality: medium; trend
data quality: medium.
• E1.2 Perennial calcareous grassland and basic steppes; area: 850 ha; trend: decreasing;
area data quality: medium; trend data quality: medium.
Literature
1. Коломійчук В.П. Флора і рослинність островів Обитічної затоки // Укр. ботан.
журн. – 2000. – 57, № 2. – С. 134–141.
2. Коломійчук В.П. Ключові ботанічні території Північного Приазов’я // Заповідна
справа в Україні. – 2008. – Т. 14, вип. 1. – С. 61–66.
3. Ткаченко В.С., Кучерява Л.Ф., Тищенко О.В. Багаторічні зміни та сучасний стан
рослинності Обитічної коси // Укр. ботан. журн. – 1998. – 55, № 6. – С. 639–647.
4. Тищенко О.В. Рослинність приморських кіс північного узбережжя Азовського
моря. – Київ: Фітосоціоцентр, 2006. – 156 с.
219
Olevski Lisy
Ukrainian name: Олевські ліси.
Transliteraion/Translaion variants: Olevskyi Forest.
Area: 4930.0 ha.
Altitude: 182–190 m.
Latitude: 51°17’17” N (51.2879°).
Longitude: 27°30’15” E (27.5042°).
Administrative regions. Zhytomyr region: Olevsk raion.
Ownership: state.
Biogeographic regions: coninental.
Habitats. Level 1. D – 0.4%; F – 1.0%; G – 97.6%; H – 1.0%.
O.O. Orlov
Habitats. Level 2. D2 – Valley mires, poor fens and transition mires – 0.4%; F9 Riverine and
fen scrubs – 1.0%; G1 Broadleaved deciduous woodland – 38.3%; G3 Coniferous woodland
– 29.0%; G4 Mixed deciduous and coniferous woodland – 26.6%; G5.6 Early-stage natural
and semi-natural woodlands and regrowth – 3.7%; H5 Miscellaneous inland habitats with
very sparse or no vegetation – 1.0%.
Futher habitat description. D2.3 Transition mires and quaking bogs; F9.2 Salix carr and
fen scrub; G1.5 Broadleaved swamp woodland on acid peat; G1.8 Acidophilous Quercusdominated woodland; G1.9 Non-riverine woodland with Betula, Populus tremula or Sorbus
220
aucuparia; G1.A Meso- and eutrophic Quercus, Carpinus, Fraxinus, Acer, Tilia, Ulmus
and related woodland; G3.4 Pinus sylvestris woodland south of the taiga; G4.7 Mixed
Pinus sylvestris – acidophilous Quercus woodland; G5 Lines of trees, small anthropogenic
woodlands, recently felled woodland, early-stage woodland and coppice; H5.6 Trampled
areas.
Land use: forestry – major; nature conservaion and research – 20%.
Protected areas: overlaps (about 1000 ha) with Banky regional forest reserve.
Threats: forestry (intensiied forest management) – low.
General description. Acidophilous Pinus sylvestris, Quercus robur, Betula pendula woods,
neutrophilous Quercus robur – Carpinus betulus woods, and Betula pubescens swamps.
Dominants of the wet pine woods are Franula alnus, Molinia caerulea, Pteridium aquilinum,
Vaccinium myrillus, Dicranum polysetum, Pleurozium shreberi. Other frequent species are
Dryopteris carthusiana, Trientalis europaea (Lysimachia europaea), Vaccinium uliginosum,
Vaccinium viis-idaea. On the drier soils, there are species-rich Pinus sylvestris woods with
dominance of Rhododendron luteum. Dominants of the herb layer are Convallaria majalis and
Pteridium aquilinum. Constant species: Calamagrosis arundinacea, Carex montana, Galium
intermedium, Maianthemum bifolium, Polygonatum odoratum, Primula veris, Serratula
coronata. Diferenial species of the swamped pine woods are Andromeda polifolia, Carex
nigra, Ledum palustre, Sphagnum fallax, Sphagnum palustre. In acidophilous Quercus robur
woods, there are the shrub layer consising of Frangula alnus. The herb layer is dominanted
by Anemone nemorosa, Luzula pilosa, Lysimachia vulgaris, Oxalis acetosella. On the richer
soil, there are Quercus robur – Carpinus betulus woods dominated by Anemone nemorosa,
Asarum europaeum, Milium efusum, Aegopodium podagraria, Galium odoratum, Lamium
galeobdolon, Pulmonaria obscura, Viola reichenbachiana.
Botanical significance. Important for conservaion of Rhododendron lutem and acidophilous
oak forests.
Criterion A
• Rhododendron luteum Sweet; A(ii); abundance: frequent; trend: stable; species data
quality: good; trend data quality: medium.
Literature
1. Орлов О.О., Сіренький С.П., Якушенко Д.М., Жижин М.П., Степаненко М.А.,
Тарасевич О.В. Природно-заповідний фонд Житомирської області. Довідник / За заг.
ред. О.О. Орлова. – Житомир – Новоград-Волинський: Вид-во НОВОград, 2015. – 404 с.
2. Орлов О.О., Якушенко Д.М., Воробйов Є.О. Флористична класифікація лісів із
участю Rhododendron luteum Sweet та радіоекологічна оцінка їх асоціацій в Поліссі
України. І. Синтаксономія лісів із участю Rhododendron luteum // Укр. фітоцен. зб. –
2000. - Серія А, Вип. 1 (16). – Київ: Фітосоціоцентр. – С. 94–113.
Opuk
V.A. Onyshchenko
Ukrainian name: Опук.
Area: 2070.0 ha.
Altitude: 0–184 m.
Latitude: 45°02’35” N (45.0429°).
Longitude: 36°12’37” E (36.2103°).
221
Administrative regions. Autonomous Republic of Crimea: Lenine raion; Territorial waters
of Ukraine.
Ownership: state.
Biogeographic regions: steppic.
Habitats. Level 1. A – 24.9%; B – 0.7%; C – 32.0%; D – 0.5%; E – 41.5%; H – 0.4%.
Habitats. Level 2. A1 Littoral rock and other hard substrata – 0.1%; A2 Littoral sediment
– 0.5%; A3 Infralittoral rock and other hard substrata – 4.1%; A5 Sublittoral sediment –
20.1%; B1 Coastal dunes and sandy shores – 0.3%; B2 Coastal shingle – 0.2%; B3 Rock cliffs,
ledges and shores, including the supralittoral – 0.1%; C1 Surface standing waters – 32.0%;
D6 Inland saline and brackish marshes and reedbeds – 0.5%; E1 Dry grasslands – 39.9%;
E2 – Mesic grasslands – 1.5%; E6 Inland salt steppes – 0.1%; H2 Screes – 0.01%; H3 Inland
clifs, rock pavements and outcrops – 0.3%; H5 Miscellaneous inland habitats with very
sparse or no vegetation – 0.1%.
Futher habitat description. A2.1 Litoral coarse sediment; A2.2 Litoral sand and muddy
sand; A5.4 Sublitoral mixed sediments; A5.5 Sublitoral macrophyte-dominated sediment;
A2.5 Coastal saltmarshes and saline reedbeds; A3.2 Atlanic and Mediterranean moderate
energy infralitoral rock; A3.2 Atlanic and Mediterranean low energy infralitoral rock;
A5.1 Sublitoral coarse sediment; A5.2 Sublitoral sand; A5.5 Sublitoral macrophytedominated sediment; B1.1 Sand beach dritline; B1.2 Sand beaches above the dritline;
B2.1 Shingle beach dritlines; B2.2 Unvegetated mobile shingle beaches above the dritline;
B2.3 Upper shingle beaches with open vegetaion; B3.1 Supralitoral rock (lichen or splash
zone); B3.2 Unvegetated rock clifs, ledges, shores and islets; B3.3 Rock clifs, ledges and
shores, with angiosperms; B3.4 Sot sea-clifs, oten vegetated; C1.5 Permanent inland
saline and brackish lakes, ponds and pools; C1.6 Temporary lakes, ponds and pools; D6.1
Inland saltmarshes; D6.2 Inland saline or brackish species-poor helophyte beds normally
without free-standing water; E1.1 Inland sand and rock with open vegetation; E1.2
Perennial calcareous grassland and basic steppes; E1.3 Mediterranean xeric grassland;
E2.7 Unmanaged mesic grassland; E6.2 Coninental inland salt steppes; H2.6 Calcareous
and ultra-basic screes of warm exposures; H3.2 Basic and ultra-basic inland clifs; H5.6
Trampled areas.
Land use: nature conservation and research – 77%.
Protected areas: includes Opukskyi nature reserve (1592.3 ha) and Emerald Site “Opukskyi
Nature Reserve” (1592.3 ha).
Threats: –.
General description. The IPA includes Opuk Hill (184 m over the sea level), salt Lake Koyash
(Koyaske) and adjacent part of the Black Sea. The steppe vegetaion occupies the largest
area, usually it is on shallow stony solis; dominants: Sipa capillata, Sipa pulcherrina,
Festuca rupicola, Festuca valesiaca, Koeleria cristata; typical species: Achillea setacea,
Artemisia taurica, Asperula kotovii, Bromopsis cappadocica, Euphorbia seguierana,
Filipendula vulgaris, Galium tenuissimum, Galium verum, Medicago glandulosa, Phleum
phleoides, Teucrium chamaedrys, Teucrium polium. On limestone outcrops, typical species
are Euphorbia petrophila, Minuaria hypanica, Pimpinella lithophila, Thymus callieri. Salinity
of Lake Koyash is 160–350‰. In summer the lake is the most saline and its peripheral part
is dry. There no macrophytes in aquaic vegetaion in the lake excluding the southern part
where there is iniltraion of less saline sea water; saltmarshes are dominated mainly by
Halocnemum strobilaceum with presence of Limonium caspium, Suaeda salsa, Puccinellia
distans, Salicornia perennans. Near the sea, there occur communiies of Crambe mariima,
Eryngium mariimum, Leymus racemosus, Phragmites australis, Juncus mariimus, Elytrigia
elongata, Aeluropus litoralis. In the sea on hard substrates near capes, there prevail
Ceramium rubrum, Cladostephus vericillatus, Corallina granifera, Cystoseira barbata,
Cystoseira crinita, Dilophus fasciola, Gelidium crinale, Gelidium laifolium, Polysiphonia
subulifera. In bays, Enteromorpha intesinalis and Ceramium ciliatum dominates. On pebble
beach and boulders near the sea, there dominate Crithmum mariimum and Elytrigia
bessarabica; on the slopes of Opuk Hill near the sea, there are communiies of Halimione
verrucifera, Holosteum umbellatum, Kochia prostrata.
Botanical significance. Important area for conservaion of the steppe vegetaion, sea clifs,
and marine macrophytes.
Criterion A
• Crambe aspera M.Bieb.; A(iv); abundance: rare; trend: unknown; species data quality:
poor; trend data quality: poor.
• Crambe koktebelica (Junge) N.Busch; A(ii); abundance: frequent; trend: unknown;
species data quality: medium; trend data quality: poor.
• Crambe mitridais Juz.; A(iv); abundance: rare; trend: unknown; species data quality:
medium; trend data quality: poor.
• Thymus litoralis Klokov et Des.-Shost.; A(iii); abundance: rare; trend: unknown; species
data quality: poor; trend data quality: poor.
Criterion C
• A3 Infralittoral rock and other hard substrata; area: 40 ha; trend: stable; area data
quality: poor; trend data quality: poor.
• A5 Sublitoral sediment; area: 400 ha; trend: stable; area data quality: medium; trend
222
223
data quality: good.
B3.3 Rock cliffs, ledges and shores, with angiosperms; area: 1 ha; trend: stable; area
data quality: poor; trend data quality: medium.
Literature
1. Белич Т.В., Садогурская С.А., Садогурский С.Е. Аннотированный список
фитобентоса Опукского природного заповедника // Труды Никит. ботан. сада. – 2006. –
Т. 126. – С. 74–88.
2. Исиков В.П. Опукский природный заповедник // Сборник тр. Никит. ботан. сада.–
2001. – С. 13–27.
3. Корженевский В.В., Рыфф Л.Э. Анализ флоры высших сосудистых растений
Опукского природного заповедника // Труды Никит. Ботан. сада. – 2006. – Т. 126. –
С. 51–73.
4. Корженевский В.В., Садогурський С.Ю., Квітницька О.І. ПЗ Опукський //
Фіторізноманіття заповідників і національних природних парків України. Ч. 1. Біосферні
заповідники. Природні заповідники / під ред. В.А. Онищенка і Т.Л. Андрієнко. – Київ:
Фітосоціоцентр, 2012. – С. 291–300.
5. Маслов И.И. Макрофитобентос некоторых заповедных акваторий Черного моря
(Украина) // Альгология. – 2002. – Т. 12, № 1. – С. 81–95.
6. Новосад В.В. Флора Керченско-Таманского региона. – Киев: Наукова думка, 1992.
– 278 с.
7. Пузанов И.И. По нехоженому Крыму. – М.: Географгиз, 1960. – 270 с.
8. Садогурская С.А. Cyanophyta морской каменистой супралиторали Крыма: Дис. …
канд. биол. наук: 03.00.05 – Ялта, 2005. – 395 с.
9. Садогурский С.Е., Белич Т.В. Макрофитобентос Опукского природного
заповедника (Черное море) // Мат-лы науч. конф., посв. 180-летию заслуженного
проф. Харьковского ун-та Л.С.Ценковского (Харьков, 4–5 декабря, 2002 г.). – Харьков,
2003. – С. 65–67.
10. Садогурский С.Е. Белич Т.В. Современное состояние макрофитобентоса
Опукского природного заповедника (Черное море) // Альгология. – 2003. – 13, № 2 – С.
185–203.
11. Садогурский С.Е., Белич Т.В., Садогурская С.А., Маслов И.И. Видовой состав
фитобентоса природных заповедников Крыма // Бюлл. ГБС РАН. – 2003. – Вып. 186. –
С. 86–104.
•
Habitats. Level 1. E – 78.6%; F – 2.0%; G – 9.6%; H – 9.8%.
Habitats. Level 2. E1 Dry grasslands – 78.6%; F3 Temperate and mediterranean-montane
scrub – 2.0%; G1 Broadleaved deciduous woodland – 9.6%; H2 Screes – 9.8%
Futher habitat description. E1.1 Inland sand and rock with open vegetation; E1.2
Perennial calcareous grassland and basic steppes; F3.1 Temperate thickets and scrub;
F3.2 Submediterranean deciduous thickets and brushes; G1.7 Thermophilous deciduous
woodland; G1.A Meso- and eutrophic Quercus, Carpinus, Fraxinus, Acer, Tilia, Ulmus and
related woodland; H2.6 Calcareous and ultra-basic screes of warm exposures.
Oskilski Skhyly
V.A. Onyshchenko, O.V. Bezrodnova, M.V. Banik
Ukrainian name: Оскільські схили.
Transliteraion/Translaion variants: Oskil Slopes.
Area: 1525.0 ha.
Altitude: 80–180 m.
Latitude: 49°57’39” N (49.9609°).
Longitude: 37°48’06” E (37.8017°).
Administrative regions. Kharkiv region: Dvorichna raion.
Ownership: state.
Biogeographic regions: steppic.
Land use: agriculture (animals) – major; nature conservaion and research – major.
Protected areas: overlaps (1067 ha) with Dvorichanskyi naional nature park, includes
Chervonyi regional botanical reserve (49.8 ha), includes Korobochkine regional botanical
reserve (29.1 ha), includes Kreidianyi regional botanical reserve (36.9 ha), includes
Konopliane regional botanical reserve (315.9 ha), overlaps (1067 ha) with Emerald Site
“Dvorichanskyi Naional Nature Park”.
224
225
Threats: abandonment/reducion of land management – low; burning of vegetaion –
medium; extracion (minerals/quarries) – low; forestry (aforestaion) – low.
General description. Slopes with outcrops of chalk, steppes, oak forests. Dominants of
communiies on chalk are Asperula tephrocarpa, Bupleurum falcatum, Thymus calcareus, less
frequently – Artemisia hololeuca, Astragalus albicaulis, Carex humilis, Cephalaria uralensis,
Hyssopus cretaceus, Koeleria talievii, Linum ucrainicum, Onosma tanaiica, Scrophularia
cretacea; locally – Artemisia salsoloides, Hedysarum grandilorum. Other typical species are
Androsace koso-poljanskii, Helianthemum cretaceum, Astragalus austriacus, Erucastrum
cretaceum, Euphorbia seguieriana, Genista tanaitica, Gypsophila alissima, Mathiola fragrans,
Meniocus linifolius, Odontites luteus, Polygala cretacea, Polygala sibirica, Silene supina,
Teucrium polium. Steppe vegetaion is dominated by Bromopsis riparia, Bromopsis inermis,
Festuca valesiaca, Koeleria cristata, Poa angusifolia, Salvia nutans, Sipa capillata, Sipa
lessingiana, locally – Sipa pennata, Sipa pulcherrima. There occur steppes with dominance
of shrubs (Caragana frutex, Chamaecyisus austriacus, Chamaecyisus tuthenicus). Typical
species of the steppe vegetaion are also Adonis vernalis, Clemais integrifolia, Euphorbia
stepposa, Galatella villosa, Galatella linosyris, Jurinea calcarea, Teucrium polium, Thalictrum
minus, Vinca herbacea. Shrub vegetaion is formed by Cerasus fruicosa, Crataegus sp.,
Euonymus verrucosus, Malus sylvestris, Prunus spinosa, Rhamnus catharica, Rosa sp., Swida
sanguinea (Cornus sanguinea), Ulmus minor. In the tree layer of forests, there dominate
Quercus robur, Fraxinus excelsior, Acer campestre, Tilia cordata. The herb layer is dominated
by Aegopodium podagraria, Stellaria holostea.
Botanical significance. Important area for conservaion of endemic species of chalk
outcrops.
Criterion A
• Alyssum gymnopodum P.A.Smirn. (Odontarrhena gymnopoda (P.A.Smirn.) D.A.German);
A(iv); abundance: rare; trend: stable; species data quality: medium; trend data quality:
medium.
• Androsace koso-poljanskii Ovcz.; A(iv); abundance: frequent; trend: stable; species data
quality: medium; trend data quality: medium.
• Diplotaxis cretacea Kotov; abundance: rare; trend: unknown; species data quality: poor;
trend data quality: poor.
• Erysimum ucrainicum J.Gay.; A(iv); abundance: rare; trend: stable; species data quality:
medium; trend data quality: medium.
Criterion C
• E1.13 Coninental dry rocky steppic grasslands and dwarf scrub on chalk outcrops;
area: 120 ha; trend: unknown; area data quality: medium; trend data quality: poor.
• E1.2 Perennial calcareous grassland and basic steppes; area: 1200 ha; trend: stable;
area data quality: medium; trend data quality: good.
Literature
1. Безроднова О.В. Раритетные кальцефильные виды в Гербарии Харьковского
национального университета имени В. Н. Каразина (CWU) //Вісник Харківського
національного університету імені В.Н. Каразіна. Серія: біологія. – 2015. Вип. 25. –
С. 16–26.
2. Савченко Г.О., Банік М.В. Ронкін В.І. Червонокнижні види судинних рослин степів
і виходів крейди національного природного парку “Дворічанський” та його околиць //
Рідкісні рослини і гриби України та прилеглих територій: реалізація природоохоронних
стратегій. Матеріали IV Міжнар. конф. 16-20 травня 2016 р. Київ. – К.: друкарня
А.В. Паливоди, 2016. – С. 133-136.
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3. Саідахмедова Н.Б., Банік М.В., Громакова А.Б., Кривохижа М.В. НПП
Дворічанський // Фіторізноманіття заповідників і національних природних парків
України. Ч. 2. Національні природні парки / під ред. В.А. Онищенка і Т.Л. Андрієнко. –
Київ: Фітосоціоцентр, 2012. – С. 191–205.
4. Ткаченко В.С., Парахонська Н.О., Горєлова Л.М. Ботанічний заказник для охорони
природної рослинності Поосколля // Укр. ботан. журн. – 1986. – 43, № 6. – С. 59–63.
5. Червона книга України. Рослинний світ / за ред. Я.П. Дідуха. – К.: Глобалконсалтинг,
2009. – 900 с.
Osovynskyi Step
V.P. Kolomiychuk, V.A. Onyshchenko
Ukrainian name: Осовинський степ.
Transliteraion/Translaion variants: Osovynskyi Steppe.
Area: 5613.0 ha.
Altitude: 0–164 m.
Latitude: 45°25’58” N (45.4327°).
Longitude: 36°28’51” E (36.4807°).
Administrative regions. Autonomous Republic of Crimea: Lenine raion; Territorial waters
of Ukraine.
Ownership: state (major), private.
Biogeographic regions: steppic.
Habitats. Level 1. A – 3.2%; B – 0.6%; D – 0.1%; E – 89.3%; F – 4.2%; G – 0.6%; H – 1.0%;
I – 1.0%.
Habitats. Level 2. A1 Littoral rock and other hard substrata – 0.0%; A2 Littoral sediment –
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0.2%; A3 Infralittoral rock and other hard substrata – 0.0%; A5 Sublittoral sediment – 2.9%;
B1 Coastal dunes and sandy shores – 0.2%; B2 Coastal shingle – 0.4%; B3 Rock clifs, ledges
and shores, including the supralittoral – 0.0%; D6 inland saline and brackish marshes and
reedbeds – <0.1%; E1 Dry grasslands – 89.3%; E6 Salt steppes – <0.1%; F3 Temperate and
mediterranean-montane scrub – 4.2%; H3 Inland clifs, rock pavements and outcrops –
0.1%; H5 Miscellaneous inland habitats with very sparse or no vegetation – 0.9%; I1 Arable
land and market gardens – 1.0%.
Futher habitat description. A2.1 Litoral coarse sediment; A3.2 Atlanic and Mediterranean
low energy infralitoral rock; A5.1 Sublitoral coarse sediment; A5.2 Sublitoral sand; A5.4
Sublitoral mixed sediments; B1.2 Sand beaches above the dritline; B2.2 Unvegetated
mobile shingle beaches above the dritline; B2.3 Upper shingle beaches with open
vegetaion; B3.1 Supralitoral rock (lichen or splash zone); D6.1 Inland saltmarshes; E1.1
Inland sand and rock with open vegetaion; E1.2 Perennial calcareous grassland and basic
steppes (>80% of the IPA); E6.2 Continental inland salt steppes; F3.2 Submediterranean
deciduous thickets and brushes; G1.C Highly ariicial broadleaved deciduous forestry
plantations; H2.6 Calcareous and ultra-basic screes of warm exposures; H3.2 Basic and
ultra-basic inland clifs; H5.3 Sparsely- or un-vegetated habitats on mineral substrates not
resulting from recent ice activity; H5.6 Trampled areas, I1.5 Bare illed, fallow or recently
abandoned arable land.
Land use: agriculture (animals) – minor, mowing/hay making – minor, nature conservaion
and research – minor, tourism/recreaion – minor.
Protected areas: includes regional landscape reserve “Osovynskyi Step” (3472 ha), regional
geological nature monuments “Hriaziova Sopka Vernadskoho” (1.0 ha), “Hriazova Sopka
Andrusova (1.0 ha), “Hriazova Sopka Obrucheva” (1.0 ha), overlaps (5433 ha) with Emerald
Site (“Karalarskyi”).
Threats: agricultural intensiicaion/expansion (general) – low, burning of vegetaion – low,
development (recreaion/tourism) – low, development (urbanization) – low.
General description. Major habitat type is the steppe. Dominant species are Festuca
valesiaca, Dactylis glomerata, Stipa brauneri, S. capillata, Agropyron pecinatum,
Elytrigia intermedia, E. elongata, Melica transsilvanica, Galatella villosa. The most typical
species with lower cover are Achillea setacea, Carduus uncinatus, Centaurea salonitana,
Malabaila graveolens, Phlomis pungens, Potenilla obscura. There are shrub communiies
with prevailing of Rosa corymbifera, R. lapidosa, R. lupulina, R. pygmaea, R. tesquicola,
Crataegus dipyrena, C. stevenii, C. taurica. On slopes to the sea, there are small groves
of Ulmus minor. In their herb layer, Anacamptis pyramidalis, Arum elongatum, Corydalus
paczoskii, Оrchis purpurea, O. picta, Pisum elatius, Scilla bifolia, Viola suavis occur. In
ravines and near mud volcanos there are halophic communiies with predominance of
Lepidium crassifolium, Petrosimonia brachiata, Petrosimonia opposiifolia, Puccinellia
fominii, Artemisia santonica, Eremopyrum triiceum.
Botanical significance. One of the largest areas of the steppe vegetaion in Ukraine. One of
the most important areas for Crambe grandilora and Crambe steveniana.
Criterion A
• Crambe grandilora DC.; A(iv); abundance: occasional; trend: stable; species data
quality: medium; trend data quality: poor.
• Crambe steveniana Rupr.; A(iv); abundance: rare; trend: stable; species data quality:
medium; trend data quality: poor.
Criterion C
• E1.2 Perennial calcareous grassland and basic steppes; area: 4300 ha; trend: stable;
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•
area data quality: medium; trend data quality: poor.
F3.247 Ponto-Sarmatic deciduous thickets; area: 50 ha; trend: unknown; area data
quality: medium; trend data quality: poor.
Pecheniiske Boloto
V.A. Onyshchenko, O.T. Kuzyarin
Ukrainian name: Печенійське болото.
Transliteraion/Translaion variants: Pecheniiske Mire.
Area: 684.0 ha.
Altitude: 234–250 m.
Latitude: 49°46’09” N (49.7692°).
Longitude: 24°21’12” E (24.3532°).
Administrative regions. Lviv region: Pustomyty raion, Zolochiv raion.
Ownership: state, private
Biogeographic regions: coninental.
Habitats. Level 1. C – 0.7%; D – 50.0%; E – 44.3%; F – 2.0%; G – 3.0%.
Habitats. Level 2. C1 Surface standing waters – 0.1%; C3 Littoral zone of inland surface
waterbodies – 0.6%; E2 Mesic grasslands – 1.3%; E3 Seasonally wet and wet grasslands
– 39.7%; E5 Woodland fringes and clearings and tall forb stands – 3.3%; D4 Base-rich
fens and calcareous spring mires – 30.0%; D5 Sedge and reedbeds, normally without freestanding water – 20.0%; F9 Riverine and fen scrubs – 2.0%; G1 Broadleaved deciduous
woodland – 3.0%.
Futher habitat descripion. D4.1 Rich fens, including eutrophic tall-herb fens and calcareous
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flushes and soaks; D5.1 Reedbeds normally without free-standing water; D5.2 Beds of
large sedges normally without free-standing water; E2.2 Low and medium altitude hay
meadows; E3.4 Moist or wet eutrophic and mesotrophic grassland; E5.4 Moist or wet tallherb and fern fringes and meadows; F9.2 Salix carr and fen scrub.
Land use: mowing/hay making – minor.
Protected areas: –.
Threats: abandonment/reducion of land management – low, agricultural intensiicaion/
expanion (general) – medium, water (drainage) – low.
General description. Parially drained fen. Major vegetaion types are rich fens and wet
grasslands. Dominant species of rich fens are Carex davalliana, Carex lava, Cladium
mariscus, Molinia caerulea, Pragmites australis, Schoenus ferrugineus. Other frequent
species are Carex lacca, Carex panicea, Equisetum palustre, Lycopus europaeus, Lysimachia
vulgaris, Lythrum salicaria, Potenilla anserina, Potenilla erecta, Salix cinerea. Meadows are
dominated by Deschampsia cespitosa, Filipendula ulmaria, Molinia caerulea.
Botanical significance. One of the largest rich fens in Ukraine with a large populaion of
Pedicularis sceptrum-carolinum.
Criterion C
• D4.1 Rich fens, including eutrophic tall-herb fens and calcareous flushes and soaks;
area: 200 ha; trend: stable; area data quality: poor; trend data quality: medium.
Conservaion proposals. Do not allow recovery of reclamaion ditches, create a state
reserve, create an Emerald Site.
Literature
1. Кузярін О.Т. Перспективні природоохоронні території басейну верхів’я Західного
Бугу // Наукові записки державного природохнавчого музею. – Львів. – 2012. – Вип. 28.
– С. 121–130.
2. Червона книга України. Рослинний світ / за ред. Я.П. Дідуха. – К.: Глобалконсалтинг,
2009. – 900 с.
Futher habitat description. D1.1 Raised bogs; D2.3 Transition mires and quaking bogs; D5.1
Reedbeds normally without free-standing water; D5.2 Beds of large sedges normally without
free-standing water; E1.7 Closed non-Mediterranean dry acid and neutral grassland; E1.9
Open non-Mediterranean dry acid and neutral grassland, including inland dune grassland;
F9.2 Salix carr and fen scrub; G1.5 Broadleaved swamp woodland on acid peat; G1.9 Nonriverine woodland with Betula, Populus tremula or Sorbus aucuparia; G3.4 Pinus sylvestris
woodland south of the taiga; G3.E Nemoral bog conifer woodland, G4.1 Mixed swamp
woodland; G4.4 Mixed Pinus sylvestris – Betula woodland.
Land use: nature conservation and research – major%.
Perebrody
V.A. Onyshchenko, O.I. Pryadko
Ukrainian name: Переброди.
Area: 21103 ha.
Altitude: 135–145 m.
Latitude: 51°41’51” N (51.6974°).
Longitude: 27°05’45” E (27.0957°).
Administrative regions. Rivne region: Dubrovytsia raion, Rokytne raion.
Ownership: state.
Biogeographic regions: continental.
Habitats. Level 1. C – 0.5%; D – 60%; E – 0.1%; F – 1%; G – 38.4%.
Habitats. Level 2. C1 Surface standing waters – 0.4%; C3 Littoral zone of inland surface
waterbodies – 0.1%; D1 Raised and blanket bogs – 1%; D2 – Valley mires, poor fens and
transition mires – 39%; D5 Sedge and reedbeds, normally without free-standing water –
20%; E1 Dry grasslands – 0.1%; F9 Riverine and fen scrubs – 1%; G1 Broadleaved deciduous
woodland – 10%; G3 Coniferous woodland – 18.4%; G4 Mixed deciduous and coniferous
woodland – 10%.
Protected areas: overlaps (13460 ha) with Rivnenskyi nature reserve, Emerald Sites
“Dubrovytsko-Sarnynskyi” (3719 ha) and “Rivnenskyi Nature Reserve” (13460 ha).
Threats: –.
General description. One of the largest mires in Ukraine. Major vegetaion type is Carex
lasiocarpa and Carex rostrata mires with a peat moss layer (Sphagnum centrale, S. fallax,
S. lexuosum, S. magellanicum, S. obtusum, S. papillosum, S. subsecundum). Constant
species are Betula pubescens, Carex limosa, Equisetum luviaile, Menyanthes trifoliata,
230
231
Peucedanum palustre, Phragmites australis, Potenilla palustris, Vaccinium oxycoccos
(Oxycoccus palustris). In hollows, there dominate Carex limosa, Rhynchospora alba,
Sphagnum cuspidatum. Signiicant area is covered by eutrophic and mesotrophic communiies
of Carex elata. Raised bogs occupy a small area. Main dominants are Eriophorum vaginatum
and Sphagnum fallax. Constant species are Andromeda polifolia, Vaccinium oxycoccos and
Pinus sylvestris. Pinus sylvestris woods are represented by dry lichen pine woods (dominants:
Cladonia miis, Corynephorus canescens), woods dominated by Vaccinium myrillus and
Pleurozium schreberi (constant species: Betula pendula, Frangula alnus, Melampyrum
pratense, Molinia caerulea, Sorbus aucuparia, Vaccinium viis idaea), Sphagnum pine woods.
Besides there are Betula pubescens and Alnus gluinosa woods with a layer of peat mosses.
Botanical significance. The largest transiion mire in Ukraine.
Criterion C
• D2.3 Transition mires and quaking bogs; area: 8000 ha; trend: stable; area data quality:
poor; trend data quality: medium.
Literature
1. Андрієнко Т.Л. ПЗ Рівненський // Фіторізноманіття заповідників і національних
природних парків України. Ч. 1. Біосферні заповідники. Природні заповідники / під
ред. В.А. Онищенка і Т.Л. Андрієнко. – Київ: Фітосоціоцентр, 2012. – С. 313–324.
2. Андрієнко Т.Л., Балашов Л.С., Прядко О.І. Унікальний болотний масив Переброди
на Ровенщині // Укр. ботан. журн. – 1976. – 33, № 5. – С. 532–536.
3. Андрієнко Т.Л., Прядко О.І., Онищенко В.А. Раритетна компонента флори
Рівненського природного заповідника // Укр. ботан. журн. – 2006. – 63, № 2. –
С. 220–228.
4. Онищенко В., Прядко О., Андрієнко Т. Рослинність ділянки Переброди
Рівненського природного заповідника // Науковий вісник Східноєвропейського
національного ун-ту ім. Лесі Українки. Сер. Біологічні науки. – 2015. – 13. – С. 32–49.
5. Фіторізноманіття Українського Полісся та його охорона / Під заг. ред.
Т.Л. Андрієнко. – Київ: Фітосоціоцентр, 2006. – 316 с.
Peresyp Aktaskoho Ozera
dunes and sandy shores – 15.0%; B2 Coastal shingle – 10.0%; E1 Dry grasslands – 22.0%; E6
Salt steppes – 12.0%; H5 Miscellaneous inland habitats with very sparse or no vegetation
– 4.0%.
Futher habitat description. A2.1 Litoral coarse sediment; A5.1 Sublitoral coarse sediment;
A5.2 Sublitoral sand; B1.2 Sand beaches above the dritline; B2.2 Unvegetated mobile
shingle beaches above the dritline; B2.3 Upper shingle beaches with open vegetaion; E1.1
Inland sand and rock with open vegetation; H5.6 Trampled areas.
V.P. Kolomiychuk, V.A. Onyshchenko
Ukrainian name: Пересип Актаського озера.
Transliteraion/Translaion variants: Peresyp of Aktash Lake.
Area: 189.0 ha.
Altitude: 0–3 m.
Latitude: 45°23’18” N (45.3884°).
Longitude: 35°55’05” E (35.9181°).
Administrative regions. Autonomous Republic of Crimea: Lenine raion; Territorial waters
of Ukraine.
Ownership: state.
Biogeographic regions: steppic.
Habitats. Level 1. A – 37.0%; B – 25.0%; E – 34.0%; H – 4.0%.
Habitats. Level 2. A2 Littoral sediment – 4.0%; A5 Sublitoral sediment – 33.0%; B1 Coastal
Land use: nature conservaion and research – minor, tourism/recreaion – minor.
Protected areas: overlaps with state ornithological reserve “Astaninski Plavni” (“Ostaninski
Zaplavy”).
Threats: climate change/ sea level rise – low, development (recreaion/tourism) – medium.
General description. A bar composed of sand and seashells. Vegetaion of the beach is
formed by Leymus sabulosus, Crambe mariima, Сakile euxina, Centaurea odessana, Carex
colchica, Eryngium maritimum. The most elevated places are dominated by Artemisia
marschalliana, Sipa capillata, Ephedra distachya, Agropyron cimmericum. On the side
opposite the sea, there are mesic and wet plant communiies dominated by Elytrigia
elongata, Elyigia repens, Glycyrrhiza glabra, Limonium meyeri.
Botanical significance. Important area for seaside threatened species Alyssum borzaeanum
(Odontarrhena borzaeana) and Thymus litoralis.
Criterion A
• Agropyron cimmericum Nevski; A(i); abundance: occasional; trend: stable; species data
quality: medium; trend data quality: poor.
232
233
•
Alyssum borzaeanum Nyár. (Odontarrhena borzaeana (Nyár.) D.A.German); A(ii);
abundance: rare; trend: unknown; species data quality: medium; trend data quality:
poor.
• Thymus litoralis Klokov et Shost.; A(iii); abundance: rare; trend: unknown; species data
quality: medium; trend data quality: poor.
Conservaion proposals. Create an Emerald Site including the enire IPA.
Literature
1. Корженевский В.В., Волкова Т.А., Клюкин А.А. О синтаксономическом положении
растительности пляжей и формирующихся дюн Азовского побережья Керченского
полуострова // Ботан. журн. – 1984. – Т. 69, № 11. – С. 1462–1467.
2. Котова И.Н. Флора и растительность Керченского полуострова // Тр. Никит. ботан.
сада. Материалы по флоре и растительности Крыма. – 1961. – Т. 35. – С. 64–168.
3. Новосад В.В. Флора Керченско-Таманского региона. – Киев: Наукова думка. –
1992. – 280 с.
cretacea, Scrophularia cretacea, Silene supina, Teucrium polium. Steppe vegetaion is
dominated by Sitpa capillata, Bromopsis riparia, Festuca valesiaca, Koeleria cristata, Poa
angusifolia, Elytrigia repens. There occur steppes with dominance of shrubs (Caragana
frutex, Chamaecyisus austriacus, Chamaecyisus ruthenicus). Typical species of the steppe
vegetaion are also Adonis vernalis, Salvia nutans, Stachys recta, Linum flavum, Thalictrum
minus. Shrub vegetaion is formed by Cerasus fruicosa, Crataegus sp., Prunus spinosa,
Rosa sp.
Petro-Ivanivka
О.V. Bezrodnova
Ukrainian name: Петро-Іванівка.
Area: 188 ha.
Altitude: 90-160 m.
Latitude: 49°55’22” N (49.9226°).
Longitude: 37°40’43”E (37.6785°).
Administrative regions. Kharkiv region: Dvorichna raion.
Ownership: state.
Biogeographic regions: steppic.
Habitats. Level 1. E – 73.3%; F – 2.0%; G – 15.7%; H – 9.0%.
Habitats. Level 2. E1 Dry grasslands – 72.7%; E5 Woodland fringes and clearings and tall forb
stands – 0.6%; F3 Temperate and mediterranean-montane scrub – 2.0%; G1 Broadleaved
deciduous woodland – 15.7%; H2 Screes – 9.0%
Futher habitat description. E1.1 Inland sand and rock with open vegetation; E1.2
Perennial calcareous grassland and basic steppes; F3.1 Temperate thickets and scrub;
F3.2 Submediterranean deciduous thickets and brushes; G1.7 Thermophilous deciduous
woodland; G1.A Meso- and eutrophic Quercus, Carpinus, Fraxinus, Acer, Tilia, Ulmus and
related woodland; G1.C Highly artificial broadleaved deciduous forestry plantations; H2.6
Calcareous and ultra-basic screes of warm exposures.
Land use: agriculture (animals) – major; forestry – minor.
Protected areas: –
Threats: abandonment/reducion of land management – low; extracion (minerals/
quarries) – low.
General description. Slopes with outcrops of chalk and steppes. Dominants of vegetaion
on chalk are Androsace koso-poljanskii, Artemisia hololeuca, Hyssopus cretaceus, Linum
ucrainicum, Thymus calcareus. Other typical species are Asperula tephrocarpa, Astragalus
albicaulis, Astragalus austriacus, Bupleurum falcatum, Carex humilis, Cephalaria uralensis,
Euphorbia stepposa, E. seguierana, Gypsophila fasigiata, Hedysarum grandilorum,
Mathiola fragrans, Odontites luteus, Onosma tanaiica, Pimpinella titanophila, Polygala
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Botanical significance. Important area for conservaion of Androsace koso-poljanskii and
other endemic species of chalk outcrops.
Criterion A
• Alyssum gymnopodum P.A.Smirn. (Odontarrhena gymnopoda (P.A.Smirn.) D.A.German);
A(iv); abundance: rare; trend: unknown; species data quality: poor; trend data quality:
poor.
• Androsace koso-poljanskii Ovcz.; A(iv); abundance: occasional; trend: unknown;
species data quality: medium; trend data quality: poor.
235
Conservaional proposals: create a state botanical reserve and an Emerald Site.
Literature
1. Безроднова О.В. Раритетные кальцефильные виды в Гербарии Харьковского
национального университета имени В. Н. Каразина (CWU) // Вісник Харківського
національного університету імені В.Н. Каразіна. Серія: біологія. – 2015. Вип.25. –
С. 16-26.
Pidlyska Hora
V.A. Onyshchenko, T.D. Solomakha
Ukrainian name: Підлиська гора.
Transliteraion/Translaion variants: Pidlys’ka Gora, Pidlys’ka Hill.
Area: 81.9 ha.
Altitude: 262–374 m.
Latitude: 49°55’56” N (49.9321°).
236
Longitude: 24°50’36” E (24.8432°).
Administrative regions. Lviv region: Zolochiv raion.
Ownership: state.
Biogeographic regions: coninental
Habitats. Level 1. E – 22.8%; F – 1.0%; G – 75.2%; H – 1.0%
Habitats. Level 2. E1 Dry grasslands – 20.5%; E5 Woodland fringes and clearings and
tall forb stands – 0.3%; E7 Sparsely wooded grasslands – 2.0%; F3 Temperate and
mediterranean-montane scrub – 1.0%; G1 Broadleaved deciduous woodland – 8.0%; G3
Coniferous woodland – 17.0%; G4 Mixed deciduous and coniferous woodland – 20.2%;
G5 Lines of trees, small anthropogenic woodlands, recently felled woodland, early-stage
woodland and coppice – 30.0%; H3 Inland cliffs, rock pavements and outcrops – 0.5%; H5
Miscellaneous inland habitats with very sparse or no vegetation – 0.5%.
Futher habitat descripion. E1.2 Perennial calcareous grassland and basic steppes; E7.2
Sub-coninental parkland; F3.1 Temperate thickets and scrub; G1.6 Fagus woodland; G3.4
Pinus sylvestris woodland south of the taiga; G3.F Highly artificial coniferous plantations;
G5.6 Early-stage natural and semi-natural woodlands and regrowth; H3.2 Basic and ultrabasic inland cliffs; H3.5 Almost bare rock pavements, including limestone pavements.
Land use: nature conservation and research – major, forestry – major.
Protected areas: included in regional nature monument “Pidlyska Hora abo Hora Markiana
Shashkevycha”.
Threats: abandonment/reducion of land management – medium, burning of vegetaion
– low.
General description. A hill with the forest, grassland and limestone outcrops. The grassland
vegetaion is dominated by Briza media, Carex lava, Carex humilis, Dactylis glomerata,
Festuca pratensis, Festuca rubra, Festuca rupicola, Inula ensifolia, Teucrium chamaedrys.
Frequent species are Allium lusitanicum s.l. (Allium montanum auct.), Anemone sylvestris,
Asperula cynanchica, Bupleurum falcatum, Campanula sibirica, Centaurea pannonica,
Euphorbia cyparissias, Galium verum, Lembotropis nigricans, Leontodon hispidus, Linum
catharicum, Onobrychis arenaria, Pimpinella saxifraga, Plantago media, Potenilla incana,
Primula veris, Prunella grandilora, Ranunculus polyanthemos, Ranunculus zapalowiczii,
Scabiosa ochroleuca, Thymus marschallianus, Thymus pannonicus, Salvia pratensis.
Botanical significance. Important area for Carlina onopordifolia and Cypripedium calceolus.
Criterion A
• Carlina onopordifolia Besser ex Szafer, Kulcz. et Pawł.; A(i), A(ii), A(iv); abundance:
frequent (289 individuals); trend: stable; species data quality: good; trend data quality:
medium.
• Cypripedium calceolus; A(ii); abundance: frequent; trend: unknown; species data
quality: medium; trend data quality: poor.
Conservaion proposals: do not allow aforestaion.
Literature
1. Дідух Я., Коротченко І. Ксеротермна рослинність північно-західного Поділля //
Вісник Львівського ун-ту. Серія біологічна. – 2003. – 34. – С. 82–91.
2. Мельник В.І., Скоропляс І.О., Баточенко В.М. Сучасний стан популяцій Carlina
onopordifolia (Asteraceae) на західному Волино-Поділлі // Укр. ботан. журн. – 2014. –
71, № 2. – С. 35–48.
3. Мельник В.И., Скоропляс И.О. Современное состояние популяций Сarlina
onopordifolia Besser ex Szafer, Kulcz. & Pawl. в Украине // Austrian Journal of Technical and
Natural Sciences. – 2014. № 9–10. – С. 21–24.
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4. Скоропляс І.О. Географічне поширення Carlina onopordifolia Besser ex Szafer,
Kulcz. et Pawł. (Asteraceae) в Україні // Ботанічні дослідження – 2014: Збірник наукових
праць V Всеукраїнської науково-практичної конференції молодих учених і студентів. –
Житомир: Вид-во ЖДУ ім. І. Франка, 2014. – С. 430–434.
Podvirivka
I.I. Chorney, A.I. Tokaryuk, V.V. Budzhak
Ukrainian name: Подвір’ївка.
Area: 45.4 ha.
Altitude: 172-221 m.
Latitude: 48°20’36” N (48.3432°).
Longitude: 26°48’29” E (26.8079°).
Administraive regions: Chernivtsi region: Kelmentsi raion.
Landuse: agriculture (animals) – major.
Protected areas: –.
Threats: agricultural intensiicaion/expansion (grazing) – low.
General description. Slopes with grasslands dominated by Briza media, Dactylis glomerata,
Elytrigia intermedia, Festuca pratensis, F. valesiaca, Helictotrichon pubescens, Koeleria
cristata, Poa pratensis. Other frequent species are Achillea millefolium, Carex tomentosa,
Centaurea jacea, Filipendula vulgaris, Galium verum, Knautia arvensis, Leontodon hispidus,
Primula veris, Ranunculus polyanthemos, Salvia pratensis, Serratula lycopifolia, Vicia
tetrasperma. The synusium of spring ephemeroids consists of Friillaria montana, Ficaria
stepporum, Bulbocodium versicolor.
Botanical significance. Important populaions of Serratula lycopifolia and Friillaria montana.
Criterion A
• Fritillaria montana Hoppe; А (ii); abundance: occasional; trend: stable; species data
quality: good; trend data quality: medium.
• Serratula lycopifolia (Vill.) A. Kern.; А (ii); abundance: occasional; trend: stable; species
data quality: good; trend data quality: medium.
Conservaion proposals. Create a state botanical reserve and an Emerald Site.
Literature
1. Каземірська М.А., Токарюк А.І., Чорней І.І. Насіннєва продуктивність Fritillaria
montana Hoppe (Liliaceae) в популяціях на північно-східній межі ареалу (середнє ПрутДністров’я) // Заповідна справа в Україні. – 2010. – Т. 16, Вип. 2. – С. 9–14.
2. Чорней І.І., Буджак В.В., Токарюк А.І. Сторінками Червоної книги України
(рослинний світ). Чернівецька область. – Чернівці: ДрукАрт, 2010. – 452 с.
Pohorylivka
Ownership: state.
Biogeographic regions: coninental.
Habitats. Level 1. E – 97.0%; F – 2.0%; G – 1.0%.
Habitats. Level 2. E1 Dry grasslands – 88.0%; E2 Mesic grasslands – 10.0%; E7 Sparsely
wooded grasslands – 1.0%; F3 Temperate and mediterranean-montane scrubs – 2.0%; G1
Broadleaved deciduous woodland – 1.0%.
Futherhabitat description: E1.2 Perennial calcareous grassland and basic steppes; E7.2
Sub-continental parkland.
I.I. Chorney, A.I. Tokaryuk, V.V. Budzhak
Ukrainian name: Погорилівка.
Area: 94.7 ha.
Altitude: 194-258 m.
Latitude: 48°32’26” N (48.5406°).
Longitude: 25°59’18” E (25.9883°).
Administraive regions: Chernivtsi region: Zastavna raion.
Ownership: state (major), municipal.
Biogeographic regions: coninental.
Habitats. Level 1. E – 96.7%; F – 2.0%; H – 0.6%; J – 0.7%
Habitats. Level 2. E1 Dry grasslands – 96.7%; F3 Temperate and mediterranean-montane
scrubs – 2.0%; H1 Terrestrial underground caves, cave systems, passages and waterbodies;
H3 Inland cliffs, rock pavements and outcrops – 0.1%; H5 Miscellaneous inland habitats
with very sparse or no vegetation – 0.5%; J3 Extractive industrial sites – 0.7%.
Futherhabitat description: E1.2 Perennial calcareous grassland and basic steppes; F3.1
Temperate thickets and scrub; H3.5 Almost bare rock pavements, including limestone
pavements; H5.6 Trampled areas.
Landuse: agriculture (animals) – minor; nature conservaion and research – major.
Protected areas: includes Chornopototskyi state karst-spelaean reserve (49 ha), Martynivske
regional nature preserve (zapovidne urochyshche) (26 ha), Pechera Fushteika regional
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geological nature monument (2.5 ha).
Threats: agricultural intensiicaion/expansion (grazing) – low.
General description. Karst area with grasslands and gypsum outcrops. Grasslands are
dominated by Carex humilis, Carex montana, Festuca valesiaca, Koeleria cristata, Stipa
Literature
1. Чорней І.І., Буджак В.В., Токарюк А.І. Сторінками Червоної книги України
(рослинний світ). Чернівецька область. – Чернівці: ДрукАрт, 2010. – 452 с.
2. Чорней І.І., Скільський І.В., Коржик В.П., Буджак В.В. Заповідні об’єкти Буковини
загальнодержавного значення як основа регіональної екологічної мережі // Заповідна
справа в Україні. – 2001. – Т. 7, вип. 2. – С. 73–98.
3. Чорней І.І., Токарюк А.І., Буджак В.В., Скільський І.В. Заповідні урочища Північної
Буковини та Хотинщини: загальний огляд, раритетні флора і фауна // Заповідна справа
в Україні. – 2009. – Т. 15, вип. 1. – С. 82–100.
Polonskyi Lis
V.A. Onyshchenko
Ukrainian name: Полонський ліс.
Transliteraion/Translaion variants: Polons’kyi Forest.
Area: 1704.0 ha.
Altitude: 236–246 m.
Latitude: 50°09’57” N (50.1659°).
Longitude: 27°24’44” E (27.4122°).
Administrative regions. Khmelnytskyi region: Polonne raion, Shepeivka raion.
Ownership: state.
capillata, Stipa pennata, Inula ensifolia, Molinia caerulea agg. There are Adonis vernalis,
Aconitum anthora, Asperula cynanchica, Astragalus austriacus, A. onobrychis, Anthericum
ramosum, Chamaecytisus albus, Filipendula vulgaris, Galium verum, Gypsophila thyraica,
Inula ensifolia, Iris hungarica, Potentilla arenaria, Pulsailla grandis, Pulsailla patens,
Pulsatilla pratensis, Salvia pratensis, S. verticillata, Securigera varia, Teucrium chamaedrys,
T. pannonicum, Thymus marschallianus, Trifolium montanum.
Botanical significance. One of the largest areas of dry grasslands in western Ukraine. There
are many protected species including Cypripedium calceolus, Iris hungarica, Gypsophila
thyraica, Pulsailla grandis, Pulsailla patens, Pulsailla pratensis, Sipa pennata.
Criterion A
• Gypsophila thyraica A.Krasnova; A(iiі); abundance: frequent; trend: unknown; species
data quality: medium; trend data quality: poor.
• Iris hungarica Waldst. & Kit.; A(ii); abundance: frequent; trend: unknown; species data
quality: medium; trend data quality: poor.
• Pulsailla grandis Wend.; A(ii); abundance: rare; trend: unknown; species data quality:
medium; trend data quality: poor.
Criterion C
• E1.2 Perennial calcareous grassland and basic steppes; area: 80 ha; trend: stable; area
data quality: medium; trend data quality: medium.
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Biogeographic regions: coninental.
Habitats. Level 1. C – 2.6%; D – 2.0%; E – 0.1%; G – 94.3%; H – 1.0%.
Habitats. Level 2. C1 Surface standing waters – 0.6%; C2 Surface running waters – 0.0%;
C3 Littoral zone of inland surface waterbodies – 2.0%; D5 Sedge and reedbeds, normally
without free-standing water – 2.0%; E3 Seasonally wet and wet grasslands – 0.1%; G1
Broadleaved deciduous woodland – 73.3%; G3 Coniferous woodland – 10.0%; G4 Mixed
deciduous and coniferous woodland – 11.0%; H5 Miscellaneous inland habitats with very
sparse or no vegetation – 1.0%.
Futher habitat description. C1.2 Permanent mesotrophic lakes, ponds and pools; C2.3
Permanent non-tidal, smooth-flowing watercourses; C3.2 Water-fringing reedbeds and
tall helophytes other than canes; E3.4 Moist or wet eutrophic and mesotrophic grassland;
G1.2 Mixed riparian floodplain and gallery woodland; G1.4 Broadleaved swamp woodland
not on acid peat; G1.8 Acidophilous Quercus-dominated woodland; G1.9 Non-riverine
woodland with Betula, Populus tremula or Sorbus aucuparia; G1.A Meso- and eutrophic
Quercus, Carpinus, Fraxinus, Acer, Tilia, Ulmus and related woodland; G3.4 Pinus sylvestris
woodland south of the taiga; G4.7 Mixed Pinus sylvestris – acidophilous Quercus woodland;
H5.6 Trampled areas.
Land use: forestry – major; nature conservaion and research – major; tourism/recreaion
– minor.
Protected areas: included in Maliovanka regional landscape park, included in Emerald Site
“Maliovanka Regional Landscape Park”.
Threats: forestry (intensiied forest management) – low.
General description. Major vegetaion type is the mesotrophic moist forests dominated
by Quercus robur in the tree layer and Carex brizoides in the herb layer. Usually there is
the shrub layer with prevailng of Frangula alnus. Other species of signiicant constancy
are Betula pendula, Calamagrosis arundinacea, Corylus avellana, Lysimachia vulgaris,
Maianthemum bifolium, Pinus sylvestris, Populus tremula. Other forest types: Quercus robur
forests dominated by Convallaria majalis or Pteridium aquilinum with presence of Betonica
oicinalis, Serratula inctoria; Pinus sylvestris and mixed Pinus sylvestris – Quercus robur
forests dominated by Vaccinium myrillus and Pleurozium shreberi; Alnus gluinosa forests
and swamps.
Botanical significance. Important for conservaion of acidophilous oak forests.
Criterion C
• G1.8 Acidophilous Quercus-dominated woodland; area: 1000 ha; trend: stable; area
data quality: medium; trend data quality: medium.
Conservaion proposals. Create a state botanical reserve. Do not allow clearcuing.
Literature
1. Природа ункального краю – Малого Полісся / Т.Л. Андрієнко, Л.С. Юглічек,
Р.Г. Білик, О.І. Прядко, М.Д. Матвєєв, Л.П. Казімірова, В.А. Онищенко, Л.Г.Безусько,
Р.Я. Арап, М.О. Тарасенко / під ред. Т.Л.Андрієнко. – Кам’янець-Подільський: Вид-во
ПП Мошинського В.С., 2010. – 254 с.
Polonyna Borzhava
V.A. Onyshchenko
Ukrainian name: Полонина Боржава.
Area: 4448.0 ha.
Altitude: 1100–1682 m.
Latitude: 48°38’46” N (48.6460°).
Longitude: 23°13’59” E (23.2332°).
Administrative regions. Zakarpatska region: Mizhhirya raion, Svaliava raion, Volovets raion.
Ownership: state, private.
Biogeographic regions: alpine.
Habitats. Level 1. E – 50.6%; F – 45.2%; G – 3.7%; H – 0.5%.
Habitats. Level 2. E1 Dry grasslands – 44.0%; E4 Alpine and subalpine grasslands stands –
6.3%; E5 Woodland fringes and clearings and tall forb stands – 0.3%; F2 Arctic, alpine and
subalpine scrub – 1.6%; F4 Temperate shrub heathland – 43.6; G1 Broadleaved deciduous
woodland – 3.7%; H3 Inland cliffs, rock pavements and outcrops – 0.0%; H5 Miscellaneous
inland habitats with very sparse or no vegetation – 0.5%.
Futher habitat description. E1.7 Non-Mediterranean dry acid and neutral closed grassland;
E4.1 Vegetated snow-patch; E4.2 Moss and lichen dominated mountain summits, ridges
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and exposed slopes; E4.3 Acid alpine and subalpine grassland; E5.5 Subalpine moist or
wet tall-herb and fern stands; F2.2 Evergreen alpine and subalpine heath and scrub; F2.3
Subalpine deciduous scrub; F4.2 Dry heaths; G1.6 Fagus woodland; H3.1 Acid silicious
inland clifs; H5.6 Trampled areas.
Land use: agriculture (animals) – major.
Protected areas: same as Emerald Site “Polonyna Borzhava”.
Threats: abandonment/reducion of land management – low.
General description. Grasslands and heaths above the upper limit of the forest. Main
dominant species are Nardus stricta and Vaccinium myrillus.
Botanical significance. Important area for heaths and mountain grasslands.
Criterion C
• E1.71 Nardus stricta swards; area: 1400 ha; trend: stable; area data quality: medium;
trend data quality: poor.
• E4.11 Boreo-alpine acidocline snow-patch grassland and herb habitats; area: 1 ha;
trend: stable; area data quality: poor; trend data quality: poor.
• E4.3 Acid alpine and subalpine grassland; area: 220 ha; trend: stable; area data quality:
medium; trend data quality: poor.
• F4.2 Dry heaths; area: 1800 ha; trend: stable; area data quality: medium; trend data
quality: poor.
Conservaion proposals. Create a naional protected area.
Literature
1.Малиновський К.А., Крічфалушій В.В. Рослинні угруповання високогір’я
Українських Карпат. – Ужгород, 2002. – 244 с.
Polovetskyi Step
V.P. Kolomiychuk
Ukrainian name: Половецький степ.
Transliteraion/Translaion variants: Polovtsian Steppe.
Area: 2120.0 ha.
Altitude: 59–135 m.
Latitude: 47°10’47” N (47.1795°).
Longitude: 36°52’26” E (36.8739°).
Administrative regions. Donetsk region: Volodarske raion; Zaporizhia region: Berdiansk
raion, Bilmak raion.
Ownership: state, private.
Biogeographic regions: steppic.
Habitats. Level 1. C – 1.2%; D – 0.9%; E – 91.7%; F – 1.0%; G – 4.2%; H –1.0%.
Habitats. Level 2. C1 Surface standing waters – 0.1%; C2 Surface running waters – 0.1%;
C3 Littoral zone of inland surface waterbodies – 1.0%; D5 Sedge and reedbeds, normally
without free-standing water – 0.9%; E1 Dry grasslands – 90.6%; E2 Mesic grasslands –
1.0%; E5 Woodland fringes and clearings and tall forb stands – 0.1% F3 Temperate and
mediterranean-montane scrub – 1.0%; G1 Broadleaved deciduous woodland – 4.0%; G3
Coniferous woodland – 0.2%; H3 Inland cliffs, rock pavements and outcrops – <0.1%; H5
Miscellaneous inland habitats with very sparse or no vegetation – 1.0%.
Futher habitat description. E1.2 Perennial calcareous grassland and basic steppes; F3.1
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Temperate thickets and scrub; F3.1 Temperate thickets and scrub; F3.2 Submediterranean
deciduous thickets and brushes; G1.1 Riparian and gallery woodland, with dominant Alnus,
Betula, Populus or Salix; G1.3 Mediterranean riparian woodland; G1.C Highly artificial
broadleaved deciduous forestry plantations; G3.F Highly artificial coniferous plantations;
H3.2 Basic and ultra-basic inland cliffs; H5.6 Trampled areas.
Land use: agriculture (animals) – major, mowing/hay making – minor, nature conservation
and research – 47%.
Protected areas: overlaps (about 990 ha) with Meotyda naional nature park and Emerald
Site “Meotyda”.
Threats: abandonment/reducion of land management – low; agricultural intensiicaion/
expansion (grazing) – low, forestry (aforestaion) – medium.
General descripion. Valleys of the Berda river in its middle reaches and the Karaiuk river
with the gulches lowing in them. Major habitat type is the steppe. Dominant species are
Festuca valesiaca, Sipa capillata, S. lessingiana, S. ucrainica, Koeleria cristata, in some places
Sipa pulcherrima. Other constant species are Adonis vernalis, Asrtagalus onobrychis, A.
palescens, Euphorbia stepposa, Jurinea arachnoidea, Haplophyllum suaveolens, Onobrychis
tanaiica, Oxytropis pilosa, Phlomoides tuberosa (Phlomis tuberosa), Plantago stepposa,
Pulsailla pratensis, Stachys transsilvanica, Thymus marschallianus. In more mesic habitats,
rhizomatous grasses Poa angusifolia, Elytrigia repens, Alopecurus pratensis dominate. In
some places there are shrub steppes with co-dominance of Caragana frutex, Amygdalus
nana. Typical species of the steppes on stony soils are Festuca valesiaca, Achillea leptophylla,
Helicrysum arenarium, Thymus dimorphus. Besides there are riverine forests of Salix alba,
Populus alba, P. nigra, plantaions of Robinia pseudoacacia, Pinus pallasiana, Ulmus minor,
Fraxinus lanceolata, shrub communiies with dominance of Prunus spinosa, Rhamnus
catharica, Crataegus fallacina, saline medows with dominance of Festuca regeliana and
Elytrigia elongata, beds of Phragmites australis, granite outcrops.
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Botanical significance. Well-preserved coninental steppe.
Criterion C
• E1.2 Perennial calcareous grassland and basic steppes; area: 1900 ha; trend: stable;
area data quality: medium; trend data quality: medium.
Conservaion proposals. Сreate a state botanical reserve.
Literature
1. Вакаренко Л.П., Мовчан Я.І., Турута О.Є. Рослинні раритети середньої течії
р. Берди // Укр. ботан. журн. – 1996. – 53, № 5. – С. 598–603
2. Вакаренко Л.П., Мосякін С.Л., Генов А.П. Наукове обгрунтування необхідності
створення міжрегіонального ландшафтного парку “Надбердянський степ” (Північне
Приазов’я) // Заповідна справа в Україні. – 2000. – Т. 6. – С. 17–27.
3. Ландшафты, растительный покров и животный мир регионального ландшафтного
парка “Меотида” / Г. Н. Молодан, С. А. Приходько, С. В. Третьяков и др. – Донецк:
Ноулидж, 2010. – 184 с.
europaea), Vaccinium myrillus (dom.), Vaccinium viis-idaea. In termophilous Quercus
robur forests, Convallaria majalis dominates. Other frequent species are Allium lusitanicum
s.l. (Allium montanum auct.), Campanula persicifolia, Centaurea phrygia, Fragaria vesca,
Lathyrus niger, Meliis sarmaica, Polygonatum odoratum, Potenilla alba, Pteridium
aquilinum, Serratula inctoria, Vicia cassubica.
Botanical significance. Large populaion of Dracocephalum ruyschiana. One of the best
species-rich oak woods in the northen Ukraine.
Criterion A
Potashnianski Lisy
O.O. Orlov, V.A. Onyshchenko
Ukrainian name: Поташнянські ліси.
Transliteraion/Translaion variants: Potashnia Forests.
Area: 852.0 ha.
Altitude: 174–190 m.
Latitude: 50°25'32" N (50.4253°).
Longitude: 29°18'56" E (29.3154°).
Administrative regions. Zhytomyr region: Radomyshl raion.
Ownership: state.
Biogeographic regions: coninental.
Habitats. Level 1. D – 1.0%; G – 98.0%; H – 1.0%.
Habitats. Level 2. G1 Broadleaved deciduous woodland – 29.9%; G3 Coniferous woodland
– 29.9%; G4 Mixed deciduous and coniferous woodland – 29.8%; G5 Lines of trees, small
anthropogenic woodlands, recently felled woodland, early-stage woodland and coppice –
8.4%; H5 Miscellaneous inland habitats with very sparse or no vegetation – 1.0%.
Futher habitat description. G1.7 Thermophilous deciduous woodland; G1.9 Non-riverine
woodland with Betula, Populus tremula or Sorbus aucuparia; G3.4 Pinus sylvestris woodland
south of the taiga; G4.7 Mixed Pinus sylvestris – acidophilous Quercus woodland; G5.6
Early-stage natural and semi-natural woodlands and regrowth; H5.6 Trampled areas.
Land use: forestry – major; nature conservaion and research – 46%.
Protected areas: includes Bervy regional botanical reserve (392 ha).
Threats: forestry (intensiied forest management) – medium.
General description. Vegetaion is represented mainly by forests (deciduous, mixed and
coniferous approximately in equal parts). Constatnt species of drier Pinus sylvestris forests
are Сhamaecyisus zingeri, Campanula rotundifolia, Melampyrum pratense, Hieracium
umbellatum, Peucedanum cervaria (dom.), Solidago virgaurea, Pleurozium schreberi (dom.),
Dicranum polysetum (dom.). In weter pine forests, constant species are Frangula alnus
(dom.), Sorbus aucuparia, Dryopteris carthusiana, Luzula pilosa, Melampyrum pratense,
Mollinia caerulea (dom.), Pteridium aquilinum (dom.), Trientalis europaea (Lysimachia
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•
Dracocephalum ruyschiana L.; A(ii); abundance: frequent; trend: unknown; species data
quality: medium; trend data quality: poor.
Criterion C
• G1.7 Thermophilous deciduous woodland; area: 150 ha; trend: stable; area data quality:
medium; trend data quality: poor.
Literature
1. Якушенко Д.М., Вініченко Т.С. Еколого-ценотична характеристика Dracocephalum
ruyschiana L. на південному сході Житомирського Полісся // Вісник Київського
національного університету імені Тараса Шевченка. Інтродукція та збереження
рослинного різноманіття. – 2005. – Вип. 8. – С. 42–44.
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Probabyn
I.I. Chorney, A.I. Tokaryuk, V.V. Budzhak
Ukrainian name: Пробабин.
Area: 4.6 ha.
Altitude: 260–287 m.
Latitude: 48°39'39" N (48.6609°).
Longitude: 25°35'14" E (25.5872°).
valesiaca, Koeleria cristata, Poa angustifolia. Other frequent species: Adonis vernalis,
Asperula cynanchica, Astragalus austriacus, Cephalaria uralensis, Chamaecytisus podolicus,
Filipendula vulgaris, Fragaria viridis, Gypsophila thyraica, Hyacinthella leucophaea, Inula
ensifolia, Muscari botryoides, Potentilla arenaria, Salvia pratensis, Securigera varia,
Teucrium chamaedrys, Thymus marschallianus, Trifolium montanum.
Botanical significance. Important area for endemic species Aconitum pseudanthora,
Chamaecytisus podolicus, Gypsophila thyraica; locus classicus of Chamaecyisus podolicus.
There are other Crierion A species: Iris hungarica, Pulsailla grandis, Pulsailla patens.
Criterion A
• Aconitum pseudanthora Błocki ex Pacz. A(iii); abundance: occasional; trend: unknown;
species data quality: medium; trend data quality: poor.
• Chamaecytisus podolicus (Błocki) Klásk. A(iv); abundance: frequent; trend: unknown;
species data quality: good; trend data quality: medium.
• Gypsophila thyraica A.Krasnova A(iiі); abundance: occasional; trend: unknown; species
data quality: medium; trend data quality: poor.
Conservaion proposals. Create a state botanical nature monument.
Provalskyi Step
Ukrainian name: Провальський степ.
Transliteraion/Translaion variants: Proval'skyi Steppe.
Area: 587.5 ha.
Altitude: 115–231 m.
Latitude: 48°07'55" N (48.1319°).
Administrative regions. Ivano-Frankivsk region: Horodenka raion.
Ownership: state.
Biogeographic regions: coninental.
Habitats. Level 1: E – 98.0%, F – 2.0%.
Habitats. Level 2: E1 Dry grasslands – 98.0%, F3 Temperate and mediterranean-montane
scrub – 2.0%.
Futher habitat description: E1.2 Perennial calcareous grassland and basic steppes; F3.1
Temperate thickets and scrub.
Land use: agriculture (animals) – minor.
Protected areas: –.
Threats: agricultural intensiicaion/expansion (grazing) – low, extracion (mineral/quarries)
– low.
General description. Grasslands dominated by Carex humilis, Festuca rupicola, Festuca
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V.A. Onyshchenko
Longitude: 39°49'09" E (39.8191°).
Administrative regions. Luhansk region: Dovzhansk (Sverdlovsk) raion.
Ownership: state.
Biogeographic regions: steppic.
Habitats. Level 1. E – 77.0%; D – 1.0%; F – 1.0%; G – 20.0%; H – 1.0%.
Habitats. Level 2. E1 Dry grasslands – 76.0%; E2 Mesic grasslands – 1.0%; D5 Sedge and
reedbeds, normally without free-standing water – 1.0%; F3 Temperate and mediterraneanmontane scrub – 1.0%; G1 Broadleaved deciduous woodland – 20.0%; H2 Screes – 0.1%; H3
Inland cliffs, rock pavements and outcrops – 0.7%; H5 Miscellaneous inland habitats with
very sparse or no vegetation – 0.2%.
Futher habitat description. E1.1 Inland sand and rock with open vegetation; E1.2 Perennial
calcareous grassland and basic steppes; E2.5 Meadows of the steppe zone; D5.1 Reedbeds
normally without free-standing water; D5.2 Beds of large sedges normally without freestanding water; F3.1 Temperate thickets and scrub; F3.2 Submediterranean deciduous
thickets and brushes; G1.7 Thermophilous deciduous woodland; H5.3 Sparsely- or unvegetated habitats on mineral substrates not resulting from recent ice activity; H5.6
Trampled areas.
Land use: nature conservaion and research – 100%.
Protected areas: included in Luhanskyi nature reserve (same as division Provalskyi Step of
the reserve), included in Emerald Site "Luhanskyi Nature Reserve".
Threats: abandonment/reducion of land management – low.
General description. Major vegetaion of the area is steppe. Most common are steppes
dominated by Sipa irsa. They are rich in mesic species: Anemone sylvestris, Filipendula
vulgaris, Fragaria viridis, Inula hirta, Melampyrum argyrocomum, Trifolium alpestre,
Trifolium montanum, Myosois popovii. A large area is occupied by the steppes of Sipa
dasyphylla with signiicant admixture of Festuca rupicola, Festuca valesiaca, Elytrigia
intermedia, Bromopsis riparia, Sipa capillata, Caragana frutex, Spiraea hypericifolia. On
eroded soils, a typical dominant is Sipa ucrainica. On shallow gravel soils, there dominates
Sipa pulcherrima, Sipa asperella, Sipa ucrainica, Botriochloa ischaemum with presence
of Artemisia marschalliana, Cota inctoria (Anthemis inctoria), Ephedra distachya,
Festuca valesiaca, Jurinea arachnoidea, Galatella villosa, Pimpinella itanophila, Psephellus
marschallianus (Centaurea marschalliana), Salvia nutans, Tanacetum millefoliom, Teucrium
polium, Thymus dimorphus. On northern slopes and in depressions, a large area is occupied
by the grasslands with dominance of rhizomatous grasses Bromopsis riparia, Bromopsis
inermis, Elytrigia intermedia, Poa angusifolia, Calamagrosis epigeios, Hierochloe repens
with admixture of Galium verum, Inula hirta, Salvia nutans, Salvia tesquicola, Securigera
varia, Phlomis pungens, Phlomoides tuberosa (Phlomis tuberosa), Trifolium alpestre,
Trifolium montanum, Trifolium pratense. Halophyic steppes dominated by Festuca
valesiaca and Galatella dracunculoides occur between ridges, their typical species are
Artemisia santonica, Halimione verrucifera, Kochia prostrata, Limonium bungei, Pimpinella
itanophila, Plantago tenuilora. On outcrops, there is open vegetaion with prevailing
Thymus calcareus, Pimpinella itanophila, Artemisia marschalliana, Agropyron pecinatum.
Typical species of these communiies are Spiraea hypericifolia, Spiraea crenata, Alyssum
tortuosum (Odontarrhena tortuosa), Asperula tephrocarpa, Silene supina, Thymus
dimorphus, Tulipa ophiophylla. Forests are dominated by Quercus robur, Fraxinus excelsior,
Acer campestre; Acer tataricum, Rhamnus catharica, Sambucus nigra, Crataegus fallacina,
Ligustrum vulgare, Euonymus verrucosus, Euonymus czernjaevii, Swida sanguinea (Cornus
sanguinea); Aegopodium podagraria, Ficaria verna, Anemone ranunculoides, Scilla siberica,
Corydalis solida, Corydalis marschalliana, Viola odorata, Melica picta, Poa nemoralis. The
IPA consists of two clusters located at a distance of 4 km from each other.
Botanical significance. Important area for conservaion of steppes.
Criterion A
• Delphinium sergii Wissjul.; A(iv); abundance: rare; trend: stable; species data quality:
medium; trend data quality: poor.
• Onosma graniicola Klokov; A(iv); abundance: rare; trend: stable; species data quality:
medium; trend data quality: medium.
Criterion C
• E1.2 Perennial calcareous grassland and basic steppes; area: 446 ha; trend: stable; area
data quality: good; trend data quality: good.
Literature
1. Біорізноманітність Луганського природного заповідника НАН України / сост.
Т. Сова. – Луганськ: ЛГСІ, 2005. – 218 с.
2. Биоразнообразие Луганского природного заповедника: растительный мир /
Составители: Сова Т. В., Русина Н. В., Гузь Г. В., Боровик Л. П., Шиян-Глотова А. В. –
Луганск: Элтон–2, 2009. – 130 с.
3. Білик Г.І., Ткаченко В.С. Рослинний покрив Провальского степу // Укр. ботан.
журн. – 1971. – 28, № 4. – С. 443–448.
4. Вєтрова З.І. Розподіл Евгленофітових у водоймах заповідника "Провальский
степ" // Укр. ботан. журн. – 1988. – 45, № 4. – С. 58–61.
5. Кондратюк Е.Н., Бурда Р.И., Чуприна Т.Т., Хомяков М.Т. Луганский гocударственный
заповедник. Растительный мир. – Киев: Наук. думка, 1988. – 188 с.
6. Маслова В.Р. Ліхенофлора Провальского степу // Зб. наук. праць "Інтродукція і
акліматизація рослин на Україні". – 1979. – Вип. 15. – С. 51–54.
7. Надеина О.В. Лишайники Провальской степи (Украина) // Ботан. журн. – 2008. –
93, № 1. – С. 3–9.
8. Приходькова Л.П. Синьозелені водорості в грунтах заповідника "Провальський
степ" // Укр. ботан. журн. – 1988. – 45, № 2. – С. 9–13.
9. Русіна Н.В. Історія та сучасний стан дослідження ліхенобіоти Луганського
природного заповідника // Наукові праці Луганського природного заповідника.
Рослинний і тваринний світ та його охорона. – Луганськ. – 2011. – Вип. 2. – С. 47–53
10. Сова Т.В., Боровик Л.П. ПЗ Луганський // Фіторізноманіття заповідників
і національних природних парків України. Ч. 1. Біосферні заповідники. Природні
заповідники / під ред. В.А. Онищенка і Т.Л. Андрієнко. – Київ: Фітосоціоцентр, 2012. –
С. 221–253.
11. Ткаченко В.С. Прогноз змін рослинності заповідника "Провальский степ" //
Укр. ботан. журн. – 1981. – 38, № 1. – С. 83–89.
12. Ткаченко В.С. Темпи демутації та коригування прогнозу розпитку степової
рослинності Провальского степу // Укр. ботан. журн. – 1990. – 47, № 2. – С. 49–54.
13. Ткаченко В.С., Чуприна Т.Т., Бакланов О.В. Заповідник Провальський степ
(сучасний стан і завдання наукових досліджень) // Укр. ботан. журн. – 1979. – 36, № 4.
– С. 352–356.
14. Червона книга України. Рослинний світ / за ред. Я.П. Дідуха. – К.:
Глобалконсалтинг, 2009. – 900 с.
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Riznykivka
Ukrainian name: Різниківка.
Area: 330.0 ha.
Altitude: 100–160 m.
Latitude: 48°50'22" N (48.8393°).
Longitude: 38°01'40" E (38.0278°).
Administrative regions. Donetsk region: Bakhmut (Artemivsk) raion.
Ownership: state.
Biogeographic regions: steppic.
V.A. Onyshchenko
quarries) – low, forestry (aforestaion) – medium.
General description. Slope of the Riznykivska valley. The steppe vegetaion is dominated by
Festuca valesiaca, Sipa capillata, S. lessingiana, S. pennata, S. tirsa, S. ucrainica. Dominants
of the chalk outcrops are Artemisia hololeuca, Hyssopys cretaceus, Thymus cretaceus.
Botanical significance. Important area for some endemic species of chalk outcrops.
Criterion A
• Linaria cretacea Fisch. ex Spreng.; A(iv); abundance: rare; trend: unknown; species data
quality: poor; trend data quality: poor.
• Silene cretacea Fisch. ex Spreng.; A(ii), A(iv); abundance: occasional; trend: unknown;
species data quality: poor; trend data quality: poor.
Conservaion proposals. Create a state botanical reserve.
Literature
1. Остапко В.М., Мулєнкова О.Г., Приходько С.А. Перспективні ботанічні об'єкти
для включення до природно-заповідного фонду Донецької області та формування
регіональної екомережі // Промышленная ботаника. Сборник научных трудов. –
Донецк: Донецкий ботанический сад НАН Украины. – 2013. – Вып. 13. – С. 25–34
2. Червона книга України. Рослинний світ / за ред. Я.П. Дідуха. – К.: Глобалконсалтинг,
2009. – 900 с.
Romosh
Habitats. Level 1. E – 79.6%; F – 2.0%; G – 8.4%; H – 10.0%.
Habitats. Level 2. E1 Dry grasslands – 79.6%; F3 Temperate and mediterranean-montane
scrub – 2.0%; G1 Broadleaved deciduous woodland – 7.7%; G3 Coniferous woodland –
0.7%; H2 Screes – 10.0%.
Futher habitat description. E1.1 Inland sand and rock with open vegetation; E1.2
Perennial calcareous grassland and basic steppes; F3.1 Temperate thickets and scrub;
F3.2 Submediterranean deciduous thickets and brushes; G1.7 Thermophilous deciduous
woodland; G1.C Highly artificial broadleaved deciduous forestry plantations; G3.F Highly
artificial coniferous plantations; H2.6 Calcareous and ultra-basic screes of warm exposures.
Land use: agriculture (animals) – major; forestry – major.
Protected areas: overlaps (about 320.0 ha) with regional botanical reserve "Kreidiana
roslynnist bilia sela Kirove", included in Emerald Site "Riznykivskyi".
Threats: agricultural intensiicaion/expansion (grazing) – low, extracion (minerals/
V.V. Datsiuk, V.A. Onyshchenko
Ukrainian name: Ромош.
Area: 636 ha.
Altitude: 205–235 m.
Latitude: 50°35'12" N (50.5867°).
Longitude: 24°21'12" E (24.3533°).
Administrative regions. Lviv region: Sokal raion.
Ownership: state.
Biogeographic regions: coninental.
Habitats. Level 1: E – 1.2%, G – 97.8%, H – 1.0% .
Habitats. Level 2: E2 Mesic grasslands – 1.2%; G1 Broadleaved deciduous woodland –
69.8%; G3 Coniferous woodland – 14.0%; G4 Mixed deciduous and coniferous woodland
– 14.0%; H5 Miscellaneous inland habitats with very sparse or no vegetation – 1.0%.
Futher habitat description: G1.9 Non-riverine woodland with Betula, Populus tremula or
Sorbus aucuparia; G1.A Meso- and eutrophic Quercus, Carpinus, Fraxinus, Acer, Tilia, Ulmus
and related woodland; G3.4 Pinus sylvestris woodland south of the taiga; G3.F Highly artificial
coniferous plantations; G4.7 Mixed Pinus sylvestris – acidophilous Quercus woodland; G5.6
Early-stage natural and semi-natural woodlands and regrowth; H5.6 Trampled areas.
Land use: forestry – major.
Protected areas: –.
Threats: abandonment/reducion of land management – low; forestry (intensiied forest
management) – low.
General description. The major vegetaion type is deciduous forests dominated by Quercus
robur, Carpinus betulus, Betula pendula. Dominants of the herb layer are Convallaria majalis,
Galium odoratum, Stellaria holostea, Asarum europaeum. Frequent species are Betula
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pendula, Fragaria vesca, Frangula alnus, Maianthemum bifolium, Pulmonaria obscura,
Sanicula europaea, Sorbus aucuparia, Viola mirabilis. Besides there are plantaions of Pinus
sylvestris with dominance of Pleurozium schreberi in the moss later.
Botanical significance. One of the largest populaions of Cypripedium calceolus in Ukraine.
The density of populaion is the highest in Pinus sylvestris wood in a former quarry of chalk
(219 individuald per 0.3 ha in the early 1980th). Typical density is 5–50 individuals per ha.
Cypripedium calceolus is present both in deciduous and coniferous forests.
Criterion A
• Cypripedium calceolus; A(ii); abundance: frequent; trend: decreasing; species data
quality: good; trend data quality: medium.
Conservaion proposals. Create a state botanical reserve.
Literature
1. Вавриш П.О., Собко В.Г. Рідкісна популяція Cypripedium calceolus L. на Волинській
височині // Укр. ботан. журн. – 1984. – 41, № 2. – С. 86–88.
2. Мельник В.І., Логвиненко І.П. Поширення та сучасний стан популяцій Cypripedium
calceolus L. (Orchidaceae) на Волинській височині // Укр. ботан. журн. – 2013. – 70, № 6.
– С. 788–791.
3. Собко В.Г. Орхідеї України. – Київ: Наук. думка, 1989. – С. 191 с.
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Rys
Ukrainian name: Рись.
Area: 320.5 ha.
Altitude: 175–192 m.
Latitude: 51°15'42" N (51.26156°).
Longitude: 25°22'59" E (25.38314°).
Administrative regions. Volynska region: Manevychi raion.
Ownership: state.
Biogeographic regions: coninental.
Habitats. Level 1. G – 99.0%; H – 1.0%.
V.A. Onyshchenko
Habitats. Level 2. G1 Broadleaved deciduous woodland – 10.0%; G3 Coniferous woodland
– 49.0%; G4 Mixed deciduous and coniferous woodland – 40.0%; H5 Miscellaneous inland
habitats with very sparse or no vegetation – 1.0%.
Futher habitat descripion. G1.9 Non-riverine woodland with Betula, Populus tremula or
Sorbus aucuparia; G3.1 Abies and Picea woodland; G3.4 Pinus sylvestris woodland south of
the taiga; G4.1 Mixed swamp woodland; G4.4 Mixed Pinus sylvestris – Betula woodland;
H5.6 Trampled areas.
Land use: forestry – minor, nature conservation and research – minor.
Protected areas: same as Rys state zoological reserve, included in Emerald Site "Cherevaskyi
Lis".
Threats: forestry (intensiied forest management) – low.
General description. Forest dominated by Pinus sylvestris, Betula pendula, Alnus gluinosa,
Picea abies.
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Botanical significance. One of the largest locaions of Picea abies forests in the latland part
of Ukraine.
Criterion C
• G3.1F Enclave Picea abies forests; area: 100 ha; trend: stable; area data quality: poor;
trend data quality: poor.
Conservaion proposals. Do not allow clearcuings. Create an Emerald Site including the
enire IPA.
Samarskyi Lis
V.A. Onyshchenko
Ukrainian name: Самарський ліс.
Transliteraion/Translaion variants: Samara Forest, River Samara Forest.
Area: 14372.0 ha.
Altitude: 51–110 m.
Latitude: 48°42'27" N (48.7075°).
Longitude: 35°31'50" E (35.5306°).
Administrative regions. Dnipropetrovsk region: Novomoskovsk raion, Pavlohrad raion.
Ownership: state.
Biogeographic regions: steppic.
Habitats. Level 1. C – 3.8%; D – 1.5%; E – 17.4%; F – 0.1%; G – 76.7%; H – 0.5%.
Habitats. Level 2. C1 Surface standing waters – 1.4%; C2 Surface running waters – 1.9%;
C3 Littoral zone of inland surface waterbodies – 0.5%; D5 Sedge and reedbeds, normally
without free-standing water – 1.2%; D6 Inland saline and brackish marshes and reedbeds
– 0.1%; E1 Dry grasslands – 0.2%; E2 Low and medium alitude hay meadows – 15.1%; E3
Seasonally wet and wet grasslands – 1.0%; E6 Inland salt steppes – 1.3%; F9 Riverine and
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fen scrubs – 0.1%; G1 Broadleaved deciduous woodland – 41.4% G3 Coniferous woodland
– 31.7%; G4 Mixed deciduous and coniferous woodland – 3.6%; H5 Miscellaneous inland
habitats with very sparse or no vegetation – 0.5%.
Futher habitat descripion. C1.2 Permanent mesotrophic lakes, ponds and pools; C2.3
Permanent non-idal, smooth-lowing watercourses; C3.2 Water-fringing reedbeds and tall
helophytes other than canes; D5.1 Reedbeds normally without free-standing water; D5.2
Beds of large sedges normally without free-standing water; D6.1 Inland saltmarshes; E1.9
Open non-Mediterranean dry acid and neutral grassland including inland dune grassland
(× E1.1); E3.4 Moist or wet eutrophic and mesotrophic grassland; E6.2 Coninental inland
salt steppes; F9.1 Riverine scrub; G1.1 Riparian and gallery woodland, with dominant
Alnus, Betula, Populus or Salix; G1.2 Mixed riparian loodplain and gallery woodland; G1.3
Mediterranean riparian woodland; G1.B Non-riverine Alnus woodland; G3.F Highly ariicial
coniferous plantaions; G1.A Meso- and eutrophic Quercus, Carpinus, Fraxinus, Acer,
Tilia, Ulmus and related woodland; G4.C Mixed Pinus sylvestris – thermophilous Quercus
woodland; H5.3 Sparsely- or un-vegetated habitats on mineral substrates not resulting
from recent ice activity; H5.6 Trampled areas.
Land use: forestry – 80%; mowing/hay making – 10%; tourism/recreaion – minor.
Protected areas: includes state ornithological reserve "Bulakhivskyi lyman" (100 ha),
regional ornithological reserve "Vasylivska kolonia sirykh chapel" (144 ha), botanical nature
monument "Dilianka nasadzhen sosny zvychainoi" (43 ha), regional botanical nature
monument "Vysokoproduktyvni nasadzhennia sosny zvychainoi" (5 ha), regional botanical
nature monument "Dilianka vikovych dubiv Vasylivskoi dachi" (3.4 ha), regional botanical
nature monument "Shtuchni dubovi nasadzhennia" (4.3 ha), regional botanical nature
monument "Shtuchni dubovi nasadzhennia" (2.2 ha), regional botanical nature monument
"Storichni dubovi nasadzhennia pryrodnoho pokhodzhennia" (7.0 ha), regional botanical
nature monument "Storichni dubovi nasadzhennia pryrodnoho pokhodzhennia" (1.9 ha),
regional botanical nature monument "Stolitni dubovi nasadzhennia" (1.8 ha), overlaps
(13136 ha) with Emerald Site "Samarskyi Lis".
Threats: agricultural intensiicaion/expansion (arable) – low; development (recreaion/
tourism) – low; forestry (intensiied forest management) – medium.
General descripion. The valley of the Samara river with broadleaved forests and meadows
in the loodplain and pine forests on the sand terrace. The tree layer of the broadleaved
forests is dominated by Quercus robur (major), Fraxinus excelsior, Tilia cordata, Ulmus
laevis, Ulmus minor. In other layers, there prevail Acer tataricum, Aegopodium podagraria,
Aristolochia clemaiis, Calamagrosis epigeios, Convallaria majalis, Frangula alnus,
Glechoma hederacea, Rubus caesius, Stellaria holostea, Urica dioica. Species of signiicant
constancy are also Anthriscus sylvestris, Asarum europaeum, Geum urbanum, Polygonatum
mulilorum, Pulmonaria obscura, Viola hirta, Viola mirabilis. Interesing are saline
broadleaved woods with presence of Carex melanostachya, Limonium gmelinii, Peucedanum
laifolium, Silaum silaus. Oak forest is also on the slope of the right bank of the Samara
river. Another important forest types are Alnus gluinosa swamp and Populus nigra wood,
oten co-dominated by Populus alba, Salix alba or Salix fragilis. The tree layer of pine forests
consists of Pinus sylvestris with paricipaion of Betula pendula, Quercus robur and Populus
tremula. The pine forest occupies soils of diferent moister: dry (with Festuca beckeri, Koeleria
sabuletorum, Sempervivum ruthenicum, Thymus pallasianus, Cladonia foliacea, Cladonia
furcata, Cladonia rangiferina), mesic (with Anthericum ramosum, Calamagrosis epigeios
(main dominant), Dianthus campestris, Hypericum perforatum, Melampyrum pratense,
Peucedanum oreoselinum, Solidago virgaurea, Pleurozium schreberi, Dicranum polysetum),
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moist (with Molinia caerulea), wet (with Calamagrosis canescens). In depressions on the
sand terrace, among pine forest, there are Betula pubescens woods with the Sphagnum
layer and wet Populus tremula woods. On dry sand, there are grasslands dominated by
Festuca beckeri, Sipa borysthenica with presence of Artemisia marschalliana, Euphorbia
seguieriana, Koeleria sabuletorum. Floodplain meadows are dominated by Bromopsis
inermis, Calamagrosis epigeios, Elytrigia repens, Festuca pratensis with presence of
Achillea millefolium, Carex praecox, Centaurea scabiosa, Convolvulus arvensis, Euphorbia
seguieriana, Euphorbia virgata, Galium verum, Gypsophila paniculata, Lathyrus tuberosus,
Lotus ucrainicus, Poa angusifolia, Poa pratensis, Securigera varia, Vicia cracca. In weter
habitats, Agrosis stolonifera and Beckmannia eruciformis dominate, there are Althaea
oicinalis, Carduus crispus, Euphorbia palustris, Geranium collinum, Inula germanica,
Lysimachia nummularia, Lotus ucrainicus, Lythrum virgatum, Melica alissima, Plantago
cornui, Trifolium pratense. In moderately saline habitats, there dominate Bolboschoenus
mariimus, Festuca regeliana, Juncus gerardii, Puccinellia distans, Triglochin mariima. More
saline habitats are occupied by communiies of Salicornia prostrata, Suaeda prostrata,
Kochia prostrata. Eutrophic mires are dominated by Carex acuta, Carex acuiformis, Carex
riparia, Phragmites australis; litoral vegetaion – Phragmites australis, Bolboschoenus
mariimus, Typha angusifolia, Glyceria maxima; aquaic vegetaion – Potamogeton
perfoliatus, Potamogeton pecinatus (Stuckenia pecinata), Myriophyllum vericillatum.
Botanical significance. This area includes one of the largest and the southernmost natural
steppe Pinus sylvestris wood in Ukraine. It includes also one of the largest loodplain oak
woods. Important for poplar woods and inland halophyic vegetaion.
Criterion C
• E6.2 Continental inland salt steppes; area: 200 ha; trend: stable; area data quality:
poor; trend data quality: poor.
• G1.22 Mixed Quercus – Ulmus – Fraxinus woodland of great rivers; area: 4500 ha; trend:
decreasing; area data quality: medium; trend data quality: medium
• G1.3 Mediterranean riparian woodland; area: 300 ha; trend: unknown; area data
quality: poor; trend data quality: poor.
• G1.414 Steppe swamp Alnus gluinosa woods; area: 300 ha; trend: stable; area data
quality: poor; trend data quality: medium.
• G3.4232 Sarmatic steppe Pinus sylvestris forests; area: 3000 ha; trend: decreasing; area
data quality: medium; trend data quality: medium.
Conservaion proposals. Create an Emerald Site a naional protected area including the
enire IPA.
Literature
1. Бельгард А.Л. Лесная растительность юго-востока УССР, Киев: Изд-во КГУ, 1950. –
264 с.
2. Екомережа степової зони України: принципи створення, структура, елементи /
під ред. Д.В. Дубини і Я.І. Мовчана. – К., 2013. – С. 409 с.
258
Savranskyi Lis
Ukrainian name: Савранський ліс.
Transliteraion/Translaion variants: Savran' Forest.
Area: 8475 ha.
Altitude: 125–205 m.
Latitude: 48°05'10" N (48.0862°).
Longitude: 30°05'36" E (30.0934°).
Administrative regions. Odesa region: Savran raion.
Ownership: state, private (minor).
Biogeographic regions: coninental.
V.A. Onyshchenko
Habitats. Level 1. G – 97.3%; H – 1.1%; I – 1.6%.
Habitats. Level 2. G1 Broadleaved deciduous woodland – 97.3%; H5 Miscellaneous inland
habitats with very sparse or no vegetation – 1.1%; I1 Arable land and market gardens –
1.6%.
Futher habitat description. G1.7 Thermophilous deciduous woodland; G1.A Meso- and
eutrophic Quercus, Carpinus, Fraxinus, Acer, Tilia, Ulmus and related woodland; H5.6
Trampled areas.
Land use: forestry – major, nature conservation and research – major.
Protected areas: includes Savranskyi Lis state landscape reserve (8397 ha), same as Emerald
Site "Savranskyi Lis".
Threats: forestry (intensiied forest management) – medium.
General description. Vegetaion consists of oak-hornbeam forests and termophilous
oak forests. The tree layer is dominated by Quercus robur, Carpinus betulus, Quercus
petraea. Frequent species are Acer campestre, Acer platanoides, Acer tataricum, Anemone
ranunculoides, Asparagus tenuifolius, Cerasus avium, Corydalis solida, Euonymus europaeus,
Euonymus verrucosus, Polygonatum hirtum, Pyrus communis, Scutellaria alissima, Swida
sanguinea (Cornus sanguinea), Tilia cordata, Ulmus glabra, Ulmus minor.
259
Botanical significance. The largest termophilous deciduous forest in latland part of Ukraine.
Criterion C
G1.7 Thermophilous deciduous woodland (*91I0); area: 5000 ha; trend: stable; area data
quality: medium; trend data quality: medium.
Conservaion proposals. Create an Emerald including the enire IPA.
Serbyno
Ukrainian name: Сербино.
Area: 1577 ha.
Altitude: 150–243 m.
Latitude: 47°53'23" N (47.8897°).
Longitude: 29°26'34" E (29.4429°).
Administrative regions. Odesa region: Balta raion.
Ownership: state.
Biogeographic regions: coninental.
V.A. Onyshchenko
Threats: forestry (intensiied forest management) – low.
General description. Termophilous deciduous forest. Main dominants of the tree layer
are Fraxinus excelsior and Quercus robur. Other frequent species are Acer campestre, Acer
platanoides, Acer tataricum, Aegonychon purpureo-caeruleum, Anemone ranunculoides,
Anthriscus sylvestris, Cerasus avium, Corydalis solida, Crataegus curvisepala s.l., Euonymus
europaeus, Euonymus verrucosus, Polygonatum hirtum, Pyrus communis, Scutellaria
alissima, Sambucus nigra, Swida sanguinea (Cornus sanguinea), Tilia cordata, Torilis
japonica, Ulmus minor.
Botanical significance. One of the largest termophilous deciduous forests in latland part
of Ukraine.
Criterion C
G1.7 Thermophilous deciduous woodland (*91I0); area: 1500 ha; trend: stable; area data
quality: medium; trend data quality: medium.
Conservaion proposals. Create a state botanical reserve including the enire IPA.
Sharivka
Ukrainian name: Шарівка.
Area: 44.4 ha.
Altitude: 84–130 m.
Latitude: 49°49’58” N (49.8327°).
Longitude: 38°53’44” E (38.8955°).
Administrative regions. Luhansk region: Bilokurakyne raion.
Ownership: state.
Habitats. Level 1. G – 99.0%; H – 1.0%.
Habitats. Level 2. G1 Broadleaved deciduous woodland – 99.0%; H5 Miscellaneous inland
habitats with very sparse or no vegetation – 1.0%.
Futher habitat description. G1.7 Thermophilous deciduous woodland; H5.6 Trampled
areas.
Land use: forestry – major.
Protected areas: included in Emerald Site "Serbyno".
260
261
V.A. Onyshchenko, M.V. Banik
Biogeographic regions: steppic.
Habitats. Level 1. E – 73.0%; F – 2.0%; G – 5.0%; H – 20.0%.
Habitats. Level 2. E1 Dry grasslands – 73.0%; F3 Temperate and mediterranean-montane
scrub – 2.0%; G1 Broadleaved deciduous woodland – 5.0%; H2 Screes – 20.0%.
Futher habitat description. E1.1 Inland sand and rock with open vegetation; E1.2 Perennial
calcareous grassland and basic steppes; H2.6 Calcareous and ultra-basic screes of warm
exposures.
Land use: agriculture (animals) – major.
Protected areas: –.
Threats: extracion (minerals/quarries) – low; forestry (aforestaion) – medium.
General description. A hill with chalk outcrops and steppe vegetaion.
Botanical significance. Important area for Hedysarum ucrainicum.
Criterion A
• Hedysarum ucrainicum Kaschm.; A(iv); abundance: frequent; trend: stable; species data
quality: good; trend data quality: medium.
Conservaion proposals. Create a state botanical reserve.
Literature
1. Banik M. Daphne’04. Final report. – Kharkiv, 2006. – 20 p. (http://www.rufford.org/
files/127.01.04%20Detailed%Final%20Report.pdf).
woodland not on acid peat; G1.5 Broadleaved swamp woodland on acid peat; G1.7
Thermophilous deciduous woodland; G1.8 Acidophilous Quercus-dominated woodland;
G1.9 Non-riverine woodland with Betula, Populus tremula or Sorbus aucuparia; G1.A
Meso- and eutrophic Quercus, Carpinus, Fraxinus, Acer, Tilia, Ulmus and related woodland;
G1.B Non-riverine Alnus woodland; G3.4 Pinus sylvestris woodland south of the taiga;
G4.4 Mixed Pinus sylvestris – Betula woodland; G4.7 Mixed Pinus sylvestris – acidophilous
Quercus woodland.
Land use: agriculture (animals) – minor, forestry – 30%; mowing / hay making – minor,
nature conservation and research – major, tourism/recreaion – minor.
Shatski Ozera
V.A. Onyshchenko
Ukrainian name: Шацькі озера.
Transliteraion/Translaion variants: Shatsk Lakes.
Area: 25003 ha.
Altitude: 162–176 m.
Latitude: 51°31’06” N (51.5184°).
Longitude: 23°52’20” E (23.8723°).
Administrative regions. Volynska region: Shatsk raion.
Ownership: state.
Biogeographic regions: coninental.
Habitats. Level 1. C – 27%; D – 6%; E – 1%; F – 1%; G – 65%.
Habitats. Level 2. C1 Surface standing waters – 25.5%; C3 littoral zone of inland surface
waterbodies – 1.5%; D2 Valley mires, poor fens and transition mires – 1.8%; D5 Sedge and
reedbeds, normally without free-standing water – 4.3%; F9 Riverine and fen scrubs – 1%;
G1 Broadleaved deciduous woodland – 17%; G3 Coniferous woodland – 42%; G4 Mixed
deciduous and coniferous woodland – 6%.
Futher habitat description. C1.1 Permanent oligotrophic lakes, ponds and pools; C1.2
Permanent mesotrophic lakes, ponds and pools; C1.3 Permanent eutrophic lakes, ponds
and pools; C3.2 Water-fringing reedbeds and tall helophytes other than canes; D2.2 Poor
fens and sot-water spring mires; D2.3 Transiion mires and quaking bogs; D5.1 Reedbeds
normally without free-standing water; D5.2 Beds of large sedges normally without freestanding water; E1.9 Open non-Mediterranean dry acid and neutral grassland including
inland dune grassland; E3.4 Moist or wet eutrophic and mesotrophic grassland; E3.5 Moist
or wet oligotrophic grassland; F9.2 Salix carr and fen scrub; G1.4 Broadleaved swamp
262
Protected areas: overlaps (24581 ha) with Shatskyi naional nature park, included in West
Polesie biosphere reserve, overlaps (24581 ha) with Emerald Site “Shatskyi”.
Threats: agricultural intensification/expansion (general) – low; development (recreaion/
tourism) – low, eutrophicaion – medium.
General description. A complex of 17 lakes with adjacent forests and mires. The largest
lakes are Sviiaz (2750 ha, maximum depth 58 m), Pulemetske (1635 ha, 4.4 m), Luky (675
ha, 3.2 m), Liutsymer (430 ha, 11 m), Ostrivyanske (250 ha, 3.8 m), Peremut (146 ha, 6.7
ha), Krymne (144 ha, 6 m), Pisochne (138 ha, 16 m). They are diferent in trophic indices
(oligo-mesotrophic to eutrophic). Aquaic vegetaion is dominated mainly by Ceratophyllum
demersum, Potamogeton crispus, Potanogeton lucens, Potamogeton pecinatus (Stuckenia
pecinata), Elodea canadensis, Straiotes aloides, Nymphaea alba, Nymphaea candida,
Nuphar lutea. There are charophyte communiies (Chara aspera, Ch. fragilis, Ch. delicatula,
Ch. contraria, Lychnothamnus barbatus, Nitella symcarpa). Litoral vegetaion is dominated
by Schoenoplectus lacustris (Scirpus lacustris), Phragmites australis, Glyceria maxima. Mires
263
belong predominantly to eutrophic and mesotrophic ones. Eutrophic mires are dominated
by Carex elata, C. appropinquata, Phragmites australis, with presence of Potenilla
palustris, Menyanthes trifoliata, Carex rostrata, Carex flava, Equisetum fluviatile, Lysimachia
thyrsiflora. Mesotrophic mires are dominated by Carex lasiocarpa, Phragmites australis,
Sphagnum fallax. Menyanthes trifoliata and Vaccinium oxycoccos (Oxycoccus palustris) are
typical co-dominants. Oten there is an open layer of Pinus sylvestris and Betula bubescens.
Swamp Alnus gluinosa forests occupy a large area. Their herb layer is dominated by
Carex riparia, C. acutiformis, Iris pseudacorus. Typical species are Thelypteris palustris,
Peucedanum palustre, Carex vesicaria, C. pseudocyperus, Stachys palustris. In more acidic
habitats, there is a layer of peat mosses (Sphagnum centrale, S. squarrosum, S. obtusum).
Wet Alnus gluinosa forests oten are co-dominated by Betula pendula, Populus tremula,
Quercus robur, Fraxinus excelsior. Main species of the shrub layer are Frangula alnus and
Corylus avellana. Typical species of the herb layer are Athyrium filix-femina, Urtica dioica,
Festuca gigantea, Ficaria verna, Aegopodium podagraria, Asarum europaeum. Pine forests
occupy the largest area. The most typical dominants of lower layers in mesic and wet Pinus
sylvestris forests are Pleurozium schreberi, Vaccinium myrillus, Molinia caerulea. Constant
species are Betula pendula, Calluna vulgaris, Dicranum polysetum, Frangula alnus, Quercus
robur, Sorbus aucuparia, Vaccinium viis-idaea. In dry habitats, typical species are Thymus
serpyllum, Festuca ovina, Jasione montana, Pilosella oicinarum (Hieracium pilosella).
There occur acidophilous Quercus robur forests, acidophilous mixed oak-pine forests, and
Betula pendula forests.
Botanical significance. Important area for conservaion of diverse freshwater communiies,
Aldrovanda vesiculosa, Dianthus pseudoseroinus.
Criterion A
• Aldrovanda vesiculosa L.; A(i), A(ii); abundance: rare; trend: unknown; species data
quality: poor; trend data quality: poor.
• Dianthus pseudoseroinus Błocki; A(iv); abundance: rare; trend: unknown; species data
quality: poor; trend data quality: poor.
Criterion C
• C1.1 Permanent oligotrophic lakes, ponds and pools; area: 1000 ha; trend: decreasing
(eutrophicaion); area data quality: poor; trend data quality: good.
• C1.226 Floating Aldrovanda vesiculosa communities; area: <0.1 ha; trend: unknown;
area data quality: poor; trend data quality: poor.
• C1.25 Charophyte submerged carpets in mesotrophic waterbodies; area: 10 ha; trend:
decreasing (eutrophicaion); area data quality: poor; trend data quality: medium.
• G3.1F Enclave Picea abies forests; area: 2 ha; trend: stable; area data quality: medium;
trend data quality: poor.
Conservaion proposals. Reduce the intensity of recreaion, the area of arable lands and the
use of ferilizers in adjacent areas.
Literature
1. Андрієнко Т.Л., Кузьмичов А.І., Прядко О.І. Болота в районі Шацьких озер // Укр.
ботан. журн. – 1971. – 26, № 6. – С. 727–733.
2. Андриенко Т.Л., Шеляг-Сосонко Ю.Р. Растительный мир Украинского Полесья в
аспекте его охраны. – Киев: Наук. думка, 1983. – 215 с.
3. Борисова Е.В., Царенко П.М., Якушенко Д.Н. Современное состояние Charales
озера Свитязь (Шацкий национальный природный парк, Волынское Полесье, Украина)
// Альгология. – 2008. – 18, № 4. – С. 449–456
4. Борисова О.В., Гончаренко В.І. До вивчення Charales озер Шацького національного
природного парку (Волинське Полісся) // Вісн. Львів. ун-ту. Сер.біол. – 2007, № 44. –
С. 46–51.
5. Борисова О.В., Якушенко Д.М. Угруповання харових водоростей південнозахідного сектора озера Світязь (Волинське Полісся) // Укр. ботан. журн. – 2008. – 65,
№ 2. – С. 226–233.
6. Вірченко В.М. Бріофлора Шацького національного природного парку // Укр.
ботан. журн. – 1999. – 56, № 1. – С. 67–73.
7. Водно-болотні угіддя України / Під ред. Г.Б. Марушевського, І.С. Жарук. – К.:
Wetlands International Black Sea Programme, 2006. – 312 с.
8. Дідух Я.П., Фіцайло Т.В., Пашкевич Н.А., Мала Ю.І. Екологічні карти
трилатерального біосферного резервату “Західне Полісся” (Шацький НПП) та їх аналіз
// Укр. ботан. журн. – 2013. – Т. 70, № 4. – С. 450–456.
9. Дідух Я.П., Якушенко Д.М., Фіцайло Т.В. Класифікація рослинності та біотопів
Української частини транскордонного біосферного резервату “Західне Полісся” //
Створення транскордонного біосферного резервату та регіональної екологічної мережі
в Поліссі. – К., 2008. – С. 41–56.
10. Карпова Г.О., Зуб Л.М. До поширення Aldrovanda vesiculosa L. на озерах
Шацької групи // Ю.Д. Клеопов та сучасна ботанічна наука. Мат-ли читань, присвячених
100-річчю з дня народження Ю.Д. Клеопова. – К., 2002. – С. 375–378.
11. Прядко О.І., Онищенко В.А. НПП Шацький // Фіторізноманіття заповідників
і національних природних парків України. Ч. 2. Національні природні парки / під ред.
В.А. Онищенка і Т.Л. Андрієнко. – Київ: Фітосоціоцентр, 2012. – С. 548–557.
12. Цурик Є.І., Жижин М.П., Ященко П.Т. Поширення та охорона Picea abies (L.)
Karsten у районі Шацьких озер // Укр. ботан. журн. – 1973. – 36, № 4. – С. 313–315.
13. Шацький національний природний парк. Наукові дослідження 1983–93 рр. –
Світязь, 1994. – 248 с.
14. Шацький національний природний парк. Наукові дослідження 1994–2004 рр.
– Луцьк, 2004. – 224 с.
15. Якушенко Д.М., Фіцайло Т.В., Коротченко І.А. Псамофітні екосистеми
української частини біосферного резервату “Західне Полісся” // Стан і біорізноманіття
екосистем Шацького національного природного парку: Мат-ли наук. конф. – Львів:
СПОЛОМ, 2005. – С. 82–86.
16. Ященко П.Т. Флористична оцінка території Шацького природного
національного парку // Укр. ботан. журн. – 1985. – 42, № 1. – С. 22–23.
17. Ященко П.Т. Судинні рослини Шацького національного природного парку //
Шацький національний природний парк. Наукові дослідження 1983–1993 рр. – Світязь.
– С. 132–163 с.
18. Ященко П.Т., Андрієнко Т.Л., Шеляг-Сосонко Ю.Р., Стойко С.М. Рослинний
покрив Шацького природного парку // Укр. ботан. журн. – 1983. – 50, № 4. – С. 68–72.
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Shebutynskyi Yar
I.I. Chorney, A.I. Tokaryuk, V.V. Budzhak
Ukrainian name: Шебутинський яр.
Transliteraion/Translaion variants: Shebutynskyi Ravine.
Area: 896 ha.
Altitude: 115–275 m.
Latitude: 48°33’03” N (48.5508°).
Longitude: 27°10’49” E (27.1803°).
Administrative regions. Chernivtsi region: Sokyriany raion.
Ownership: state.
Biogeographic regions: coninental.
Habitats. Level 1. E – 1%; G – 98%; H – 1%
Habitats. Level 2. E1 Dry grasslands – 0.5%; E2 Mesic grasslands – 0.5%; G1 – Broadleaved
deciduous woodland – 98.0%; H5 Miscellaneous inland habitats with very sparse or no
vegetation – 1.0%.
Yar”, included in naional nature park “Khotynskyi”, overlaps (299 ha) with Emerald Site
“Khotynskyi Naional Nature Park”.
Threats: development (recreaion/tourism) – low.
General description. Broadleaved forests dominated by Quercus robur with Carpinus
betulus, Acer pseudoplatanus, Acer platanoides, Tilia cordata, Fraxinus excelsior, Cerasus
avium, Sorbus torminalis. The shrub layer is composed of Acer tataricum, Acer campestre,
Corylus avellana, Euonymus verrucosus, Euonymus europaeus, Sambucus nigra, Staphylea
pinnata, Swida sanguinea (Cornus sanguinea), Viburnum lantana. Typical species of the
herb layer are Carex pilosa, Stellaria holostea, Asarum europaeum, Aegopodium podagraria,
Lathyrus vernus, Hedera helix, Hepatica nobilis.
Botanical significance. Important area for Friillaria montana and Aconitum besserianum.
Criterion А
• Aconitum besserianum Andrz. ex Trautv; А (iiі); abundance: rare; trend: stable; species
data quality: medium; trend data quality: poor.
• Fritillaria montana Hoppe; А (ii); abundance: rare (1000 individuals); trend: stable;
species data quality: good; trend data quality: poor.
Literature
1. Волуца О.Д. Fritillaria montana Hoppe у флорі Чернівецької області // Науковий
вісник Чернівецького університету: Збірник наукових праць. – Вип. 293: Біологія. –
Чернівці: Рута, 2006. – С. 31–34.
2. Каземірська М.А. Стан популяції Fritillaria montana Hoppe (Liliaceae) на території
ландшафтного заказника місцевого значення “Шебутинський яр” // Наукові основи
збереження біотичної різноманітності: Матеріали десятої наукової конференції
молодих учених (Львів, 7-8 жовтня 2010 року). – Львів, 2010. – С. 50–52.
3. Каземірська М.А., Чорней І.І. Fritillaria montana Hoppe (Liliaceae): географічна
характеристика, поширення в Україні // Науковий вісник Чернівецького університету.
Біологія (Біологічні системи). – 2010. – Т. 2, Вип. 3. – С. 63–68.
4. Каземірська М.А., Чорней І.І. Fritillaria montana Hoppe на території ландшафтного
заказника місцевого значення “Шебутинський яр” // Молодь і поступ біології: збірник
тез VI Міжнародної наукової конференції студентів і аспірантів (21-24 вересня 2010
року, м. Львів). – Львів, 2010. – С. 56–57.
5. Каземірська М.А., Чорней І.І. Вікова та просторова структури популяцій Fritillaria
montana Hoppe у Прут-Дністровському межиріччі // Біологічні системи. – 2010. – Т. 2.,
Вип. 2. – С. 62–66.
6. Червона книга України. Рослинний світ / за ред. Я.П. Дідуха. – К.: Глобалконсалтинг,
2009. – 900 с.
7. Чорней І.І., Буджак В.В., Токарюк А.І. Сторінками Червоної книги України
(рослинний світ). Чернівецька область. – Чернівці: ДрукАрт, 2010. – C. 85–87.
Sinozhai
Futher habitat description. G1.7 Thermophilous deciduous woodland; G1.A Meso- and
eutrophic Quercus, Carpinus, Fraxinus, Acer, Tilia, Ulmus and related woodland.
Land use: forestry – major, nature conservation and research – major.
Protected areas: overlaps (about 790 ha) with regional landscape reserve “Shebutynskyi
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I.I. Chorney, A.I. Tokaryuk, V.V. Budzhak
Ukrainian name: Сіножаті.
Area: 63.5 ha.
Altitude: 285–325 m.
Latitude: 48°36’05” N (48.6015°).
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Longitude: 25°13’11” E (25.2198°).
Administrative regions. Ivano-Frankivsk region: Horodenka raion, Kolomyia raion.
Ownership: private, state.
Biogeographic regions: coninental.
Habitats. Level 1: E – 92.0%, F – 8.0%.
Habitats. Level 2: E1 Dry grasslands – 75.0%; E2 Mesic grasslands – 17.0%, F3 Temperate
Botanical significance. One of the best areas for Pulsailla grandis. There are other criterion
A species: Adenophora liliifolia, Pulsailla patens.
Criterion A
• Adenophora liliifolia; A(ii); abundance: rare; trend: unknown; species data quality: good;
trend data quality: poor.
• Pulsailla grandis Wend.; A(ii); abundance: frequent; trend: unknown; species data
quality: good; trend data quality: poor.
• Pulsailla patens (L.) Mill.; A(ii); abundance: frequent; trend: unknown; species data
quality: good; trend data quality: poor.
Conservaion proposals. Do not allow land plowing. Create a state botanical reserve and
an Emerald Site.
Skhidne Roztochia
Ukrainian name: Східне Розточчя.
Transliteraion/Translaion variants: Eastern Roztochchia.
Area: 13680 ha.
Altitude: 288–397 m.
Latitude: 49°58’22” N (49.9728°).
Longitude: 23°50’53” E (23.8479°).
Administrative regions. Lviv region: Zhovkva raion, Yavoriv raion.
and mediterranean-montane scrub – 8.0%.
Futher habitat description: E1.2 Perennial calcareous grassland and basic steppes; F3.1
Temperate thickets and scrub.
Land use: agriculture (animals) – minor.
Protected areas: –.
Threats: agricultural intensification/expansion (arable) – low.
General description. Species-rich grassland with dominance of Brachypodium pinnatum,
Calamagrostis arundinacea, Cirsium pannonicum, Festuca rupicola, F. valesiaca, Koeleria
cristata, Molinia caerulea agg. Other frequent species: Adenophora lilifolia, Colchicum
autumnale, Clematis recta, Galium boreale, Geranium sanguineum, Gymnadenia conopsea,
Helictotrichon praeustum, Hypericum maculatum, Hypochaeris maculata (Achyrophorus
maculatus), Inula hirta, Inula salicina, Knautia arvensis, Lathyrus pannonicus, Lilium
martagon, Orchis ustulata, Peucedanum cervaria, Potentilla alba, Prunella grandiflora,
Pulsatilla grandis, P. patens, Salvia pratensis, Sanguisorba officinalis, Scorzonera humilis, S.
purpurea, Traunsteinera globosa, Trifolium alpestre, T. montanum, T. pannonicum.
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V.A. Onyshchenko
Ownership: state.
Biogeographic regions: coninental.
Habitats. Level 1. C – 0.2%; D – 0.4%; E – 0.3%; G – 98.6%; H – 0.5%.
Habitats. Level 2. C1 Surface standing waters – 0.2%; D2 Valley mires, poor fens and
transition mires – 0.1%; D5 Sedge and reedbeds, normally without free-standing water
– 0.3%; E2 Mesic grasslands – 0.3%; G1 Broadleaved deciduous woodland – 77.7%; G3
Coniferous woodland – 2.8%; G4 Mixed deciduous and coniferous woodland – 18.1%.
Futher habitat descripion. D2.3 Transiion mires and quaking bogs; E1.9 Open nonMediterranean dry acid and neutral grassland including inland dune grassland; E2.2
Low and medium alitude hay meadows; E3.4 Moist or wet eutrophic and mesotrophic
grassland; E3.5 Moist or wet oligotrophic grassland; G1.2 Mixed riparian floodplain and
gallery woodland; G1.4 Broadleaved swamp woodland not on acid peat, G1.6 Fagus
woodland; G1.8 Acidophilous Quercus-dominated woodland; G1.9 Non-riverine woodland
with Betula, Populus tremula or Sorbus aucuparia; G1.A Meso- and eutrophic Quercus,
Carpinus, Fraxinus, Acer, Tilia, Ulmus and related woodland; G3.4 Pinus sylvestris woodland
south of the taiga; G3.E Nemoral bog conifer woodland; G4.4 Mixed Pinus sylvestris – Betula
woodland; G4.7 Mixed Pinus sylvestris – acidophilous Quercus woodland; H5.6 Trampled
areas.
Land use: forestry – major, nature conservation and research – major.
Protected areas: includes Roztochia nature reserve (2085 ha), overlaps (about 2920 ha)
with Yavorivskyi naional nature park; includes Zavadivskyi regional landscape reserve (3561
ha), includes Hriada forest reserve (1149 ha), overlaps (about 5790 ha) with Roztochya
biosphere reserve; includes Emerald Site “Roztochia Nature Reserve” (2085 ha), overlaps
(about 2920 ha) with Emerald Site “Yavorivskyi Naional Nature Park”, overlaps (about 7820
ha) with Emerald Site “Zavadivskyi”, overlaps (about 855 ha) with Emerald Site “Roztochia”.
Threats: forestry (intensiied forest management) – low; development (recreaion/tourism)
– low.
General description. Major vegetaion type is the Fagus sylvaica forest. Dominants of
the herb layer are Anemone nemorosa, Asarum europaeum, Carex pilosa, Convallaria
majalis, Lamium galeobdolon, Maianthemum bifolium. Other species of high constancy
are Acer platanoides, Acer pseudoplatanus, Athyrium ilix femina, Carex digitata, Dryopteris
carthusiana, Dryopteris ilix-mas, Melica nutans, Galium odoratum, Polytrichum formosum,
Quercus robur, Rubus hirtus. Signiicant areas are covered by the Carpinus betulus – Quercus
robur forests, Quercus robur – Pinus sylvestris forests, Pinus sylvestris forests. Dominants of
acidophilous Quercus robur – Pinus sylvestris forests are Convallaria majalis, Maianthemum
bifolium, Vaccinium myrillus. Dominants of the Pinus sylvestris forests are Vaccinium
myrillus, Pleurozium schreberi. Other constant species are Dicranum polysetum, Luzula
pilosa, Melampyrum pratense, Trientalis europaea (Lysimachia europaea), Vaccinium viisidaea.
Botanical significance. Important area for latland beech forests.
Criterion C
• G1.6 Fagus woodland (excluding G1.66); area: 8300 ha; trend: increasing; area data
quality: medium; trend data quality: good.
Literature
1. Данилків І.С., Сорока М.І. Мохоподібні державного заповідника “Розточчя”.
Препр. – Львів, 1989. – 78 с.
2. Жижин М.П., Кагало О.О., Чабан Х.І. Рослинність урочища Заливки заповідника
Розточчя // Укр. ботан. журн. – 1988. – 45, № 1. – С. 68–72.
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3. Кагало О.О. ПЗ Розточчя // Фіторізноманіття заповідників і національних
природних парків України. Ч. 1. Біосферні заповідники. Природні заповідники / під
ред. В.А. Онищенка і Т.Л. Андрієнко. – Київ: Фітосоціоцентр, 2012. – С. 325–335.
4. Кагало О.О. НПП Яворівський // Фіторізноманіття заповідників і національних
природних парків України. Ч. 2. Національні природні парки / під ред. В.А. Онищенка і
Т.Л. Андрієнко. – Київ: Фітосоціоцентр, 2012. – С. 558–565.
5. Сорока М.І. Судинні рослини державного заповідника Розточчя. – Львів, 1990. –
С. 81–85.
6. Сорока М.І. Синтаксономія рослинності Українського Розточчя // Науковий вісник
УкрДЛТУ. – Львів, 1998, вип. 7. – С. 37–41.
7. Сорока М.І. Рослинність Українського Розточчя. – Львів: Вид-во “Світ”, 2008. – 432 с.
8. Ткачик В. Рослинність заповідника “Розточчя”: ідентифікація сучасного розмаїття
фітоценозів. – Львів: НТШ, 1997. – 120 с.
9. Ткачик В.П. Рослинність заповідника “Розточчя”: класифікація методом БраунБланке. – Львів: НТШ, 1999. – 198 с.
Skhidnyi Churiuk
V.P. Kolomiychuk, V.A. Onyshchenko
Ukrainian name: Східний Чурюк.
Transliteraion/Translaion variants: Eastern Churyuk.
Area: 2210.0 ha.
Altitude: 0–13 m.
Latitude: 46°07’00” N (46.1166°).
Longitude: 34°15’36” E (34.2600°).
Administrative regions. Kherson region: Novotroitske raion.
Ownership: state.
Biogeographic regions: steppic.
Habitats. Level 1. A – 58.8%; E – 41.2%.
Habitats. Level 2. A2 Littoral sediment – 3.0%; A5 Sublittoral sediment – 55.8%; E1 Dry
grasslands – 37.2%; E6 Salt steppes – 4.0%.
Futher habitat description. A2.3 Littoral mud; A2.5 Coastal saltmarshes and saline
reedbeds; A5.3 Subittoral mud; E1.2 Perennial calcareous grassland and basic steppes; E6.2
Continental inland salt steppes.
Land use: nature conservaion and research – 100%.
Protected areas: included in Azovo-Syvaskyi naional nature park (zone of strict protecion),
included in Ramsar Site “Central Syvash”, included in Emerald Site “Azovo-Syvaskyi Naional
Nature Park”.
Threats: abandonment/reducion of land management – low, natural events: erosion –
medium.
General description. A loess island in the Syvash Bay with adjacent waters. Major vegetaion
type on the Churiuk island is the steppe. Dominant species are Stipa lessingiana, S. capillata,
S. ucrainica, Agropyron pectinatum, Festuca valesiaca. Other frequent species are Achillea
setacea, Crinitaria villosa, Dianthus elongatus, Elisanthe viscosa, Hesperis trisis, Linaria
macroura, Malabaila graveolens, Phlomis pungens, Tanacetum millefolium, Valeriana
tuberosa, Vicia angusifolia, Vicia cracca. In depressions, there dominate Elytrigia repens,
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Elytrigia intermedia, Leymus ramosus, Festuca valesiaca, Poa angusifolia. In the salt steppe,
dominant species are Artemisia santonica, Festuca valesiaca, Caroxylon laricinum, Kochia
prostrata, Ephedra distachya. In mesic and wet halophyic communiies, there prevail
Puccinellia fominii, Limonium meyeri, Artemisia santonica. Peripheral part of the island is
occupied by solonchaks dominated by Halocnemum strobilaceum, Lepidium crassifolium,
Limonium sufruicosum, Petrosimona brachiata, Puccinellia fominii, Salicornia perennans.
2. Коломійчук В.П. Рідкісні види судинних рослин Азово-Сиваського національного
природного парку // Вісті біосферного заповідника “Асканія-Нова”. – 2002. – №4. –
С. 37–44.
3. Коломійчук В.П. Азово-Сиваський національний природний парк //
Фіторізноманіття національних природних парків України / під ред. Т.Л. Андрієнко та
В.А. Онищенка. – К.: Науковий світ, 2003. – C. 19–28.
4. Коломійчук В.П. Рідкісні рослинні угруповання островів Сиваша // Укр. ботан.
журн. – 2003. – 60, № 5 – С. 540–545.
5. Коломійчук В.П. Азово-Сиваський національний природний парк //
Фіторізноманіття національних природних парків України / під ред. Т.Л. Андрієнко та
В.А. Онищенка. – К.: Науковий світ, 2003. – C. 19–28.
6. Коломійчук В. П., Онищенко В. А., Перегрим М. М. Важливі ботанічні території
Приазов’я / за ред. Т.Л. Андрієнко. – К. : Альтерпрес, 2012. – 114 с.
Skhyly Kohylnyka
Ukrainian name: Схили Когильника.
Transliteraion/Translaion variants: Kohyl’nyk Slopes.
Area: 2683 ha.
Altitude: 40–180 m.
Latitude: 46°14’24” N (46.2400°).
Botanical significance. One of the best sites for endemic species Lepidium syvaschicum
(locus classicus, globally ~10 localiies), Allium scythicum (globally ~10 localiies). One of the
best examples of the southern Ukrainian steppes.
Criterion A
• Allium scythicum Zoz.; A(iii); abundance: rare; trend: luctuaing; species data quality:
medium; trend data quality: poor.
• Lepidium syvaschicum Kleopow; A(iii); abundance: occasional; trend: stable; species
data quality: medium; trend data quality: medium.
Criterion C
• E1.2 Perennial calcareous grassland and basic steppes; area: 800 ha; trend: slow
decreasing; area data quality: good; trend data quality: good.
Literature
1. Бойко М.Ф. Раритетне фіторізноманіття Херсонщини (Україна) та його охорона //
Чорноморський ботанічний журнал. – 2005. – 1, №1. – С. 53–59.
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273
V.A. Onyshchenko, O.M. Popova
Longitude: 29°04’26” E (29.0739°).
Administrative regions. Odesa region: Tarutyne raion.
Ownership: state.
Biogeographic regions: continental.
Habitats. Level 1. E – 84%; G – 16%.
Habitats. Level 2. E1 Dry grasslands – 84%; G1 Broadleaved deciduous woodland – 16%.
Futher habitat description. E1.2 Perennial calcareous grassland and basic steppes; G1.7
Thermophilous deciduous woodland (minor); G1.C Highly artificial broadleaved deciduous
forest plantations.
Land use: agriculture (animals) – major, forestry – major.
Protected areas: included in Emerald Site “Besarabskyi Kolkhikum”.
Threats: forestry (aforestaion) – high.
General description.
The area is situated on the slope of the Kohylnyk river’s valley. Major habitat type is the
steppe. A signiicant part of the area is occupied by forest plantaions of Gleditschia
triacanthos and Robinia pseudoacacia. Fragments of natural termophilous Fraxinus
excelsior woods occur.
Botanical significance. The area is noted for conservaion of Colchicum fominii.
Criterion A
• Colchicum fominii Bordz; A(ii), A(iv); abundance: frequent; trend: unknown; species
data quality: medium; trend data quality: poor.
Conservaion proposals. Create an Emerald Site.
Land use: agriculture (animals) – major; forestry – minor; nature conservaion and research
– major.
Protected areas: overlaps (about 350 ha) with Honcharivskyi regional botanical reserve
Skhyly Krasnoi i Kobylky
V.A. Onyshchenko
Ukrainian name: Схили Красної і Кобилки.
Transliteraion/Translaion variants: Krasna and Kobylka Slopes.
Area: 663.9 ha.
Altitude: 80–130 m.
Latitude: 49°30’50” N (49.5138°).
Longitude: 38°08’12” E (38.1367°).
Administrative regions. Luhansk region: Svatove raion.
Ownership: state.
Biogeographic regions: steppic.
Habitats. Level 1. E – 83.7%; F – 2.0%; G – 6.2%; H – 5.1%; I – 3.0%.
Habitats. Level 2. E1 Dry grasslands – 78.7%; E2 Mesic grasslands – 5.0%; F3 Temperate and
mediterranean-montane scrub – 2.0%; G1 Broadleaved deciduous woodland – 6.2%; H2
Screes – 5.1%; I1 Arable land and market gardens – 3.0%.
Futher habitat description. E1.1 Inland sand and rock with open vegetation; E1.2 Perennial
calcareous grassland and basic steppes; E2.2 Low and medium alitude hay meadows; E2.5
Meadows of the steppe zone; F3.1 Temperate thickets and scrub; F3.2 Submediterranean
deciduous thickets and brushes; G1.7 Thermophilous deciduous woodland; G1.A Mesoand eutrophic Quercus, Carpinus, Fraxinus, Acer, Tilia, Ulmus and related woodland; G1.C
Highly artificial broadleaved deciduous forestry plantations; H2.6 Calcareous and ultrabasic screes of warm exposures; I1.5 Bare illed, fallow or recently abandoned arable land.
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Threats: agricultural intensification/expansion (arable) – low, extracion (minerals/quarries)
– low; forestry (aforestaion) – low.
General description. Slopes of the valleys of the Krasna and Kobylka rivers with the steppe
vegetaion, outcrops of chalk and oak woods. Dominants of the chalk outcrops are Artemisia
hololeuca, Artemisia salsoloides, Hyssopus cretaceus, Silene cretacea, Silene supina, Thymus
cretaceus. Other typical species are Asperula tephrocarpa, Bupleurum falcatum, Cephalaria
uralensis, Gypsophila oligosperma, Pimpinella titanophila, Teucrium polium.
Botanical significance. Important area for conservaion of chalk outcrops with endemic
species.
Criterion A
• Linaria cretacea Fisch. ex Spreng.; A(iv); abundance: rare; trend: unknown; species data
quality: poor; trend data quality: poor.
• Silene cretacea Fisch. ex Spreng.; A(ii), A(iv); abundance: occasional; trend: stable;
species data quality: medium; trend data quality: poor.
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Conservaion proposals. Create an Emerald Site including the enire IPA.
Literature
1. Природно-заповідний фонд Луганської області // О.А. Арапов (заг. ред.), Т.B.
Сова, О.А. Cавенко, В.Б. Фєрєнц, Н.У. Кравець, Л.Л. Зятьков, Л.О. Морозова. Довідник.
– 3-е вид., доп. і перероб. – Луганськ: “Луганська правда”, 2013. – 224 с.
Slovechanskyi Kriazh
V.A. Onyshchenko
Ukrainian name: Словечанський кряж.
Transliteraion/Translaion variants: Slovechanskyi Ridge, Slovechno Ridge.
Area: 6119.0 ha.
Altitude: 187–191 m.
Latitude: 51°20’18” N (51.3384°).
Longitude: 28°12’25” E (28.2069°).
Administrative regions. Zhytomyr region: Ovruch raion.
G1 Broadleaved deciduous woodland – 57.3%; G3 Coniferous woodland – 30.9%; G4 Mixed
deciduous and coniferous woodland – 10.7%; H5 Miscellaneous inland habitats with very
sparse or no vegetation – 0.5%; J4 Transport networks and other constructed hard-surfaced
areas – 0.1%.
Futher habitat description. D5.1 Reedbeds normally without free-standing water; D5.2
Beds of large sedges normally without free-standing water; G1.4 Broadleaved swamp
woodland not on acid peat; G1.8 Acidophilous Quercus-dominated woodland; G1.9 Nonriverine woodland with Betula, Populus tremula or Sorbus aucuparia; G3.4 Pinus sylvestris
woodland south of the taiga; G4.7 Mixed Pinus sylvestris – acidophilous Quercus woodland;
H5.6 Trampled areas.
Land use: forestry – major; nature conservaion and research – major.
Protected areas: included in Slovechanskyi Kriazh regional forest reserve, included in
Emerald Site “Slovechanskyi Kriazh”.
Threats: forestry (intensiied forest management) – low.
General description. Major vegetaion types are acidophilous forests dominated by Quercus
robur, Pinus sylvestris, Betula pendula. In the herb layer, there dominate Calamagrosis
arundinacea, Maianthemum bifolium, Oxalis acetosella, Vaccinium myrillus. In pine forests,
there is a moss layer dominated by Pleurozium shreberi and Dicranum polysetum. Smaller
areas are covered by Quercus petraea and Alnus gluinosa forests.
Botanical significance. Important for conservaion of acidophilous oak forests.
Criterion C
• G1.8 Acidophilous Quercus-dominated woodland; area: 2000 ha; trend: stable; area
data quality: medium; trend data quality: medium.
Somyno
Ownership: state.
Biogeographic regions: coninental.
Habitats. Level 1. D – 0.5%; G – 98.9%; H – 0.5%; J – 0.1%.
Habitats. Level 2. D5 Sedge and reedbeds, normally without free-standing water – 0.5%;
V.A. Onyshchenko
Ukrainian name: Сомино.
Area: 10852 ha.
Altitude: 146–160 m.
Latitude: 51°25’22” N (51.4228°).
Longitude: 26°55’11” E (26.9197°).
Administrative regions. Rivne region: Sarny raion.
Ownership: state.
Biogeographic regions: continental.
Habitats. Level 1. C – 1%; D – 60%; F – 1%; G – 38%.
Habitats. Level 2. C1 Surface standing waters – 1%; D2 – Valley mires, poor fens and
transition mires – 59%; D5 Sedge and reedbeds, normally without free-standing water – 1%;
F9 Riverine and fen scrubs – 1%; G1 Broadleaved deciduous woodland – 10% G3 Coniferous
woodland – 18%.
Futher habitat description. C1.2 Permanent mesotrophic lakes, ponds and pools; C3.2
Water-fringing reedbeds and tall helophytes other than canes; D2.3 Transition mires and
quaking bogs; D5.1 Reedbeds normally without free-standing water; D5.2 Beds of large
sedges normally without free-standing water; E1.7 Closed non-Mediterranean dry acid and
neutral grassland; E1.9 Open non-Mediterranean dry acid and neutral grassland, including
inland dune grassland; F9.2 Salix carr and fen scrub; G1.5 Broadleaved swamp woodland on
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acid peat; G1.9 Non-riverine woodland with Betula, Populus tremula or Sorbus aucuparia;
G3.4 Pinus sylvestris woodland south of the taiga; G3.E Nemoral bog conifer woodland; G4.4
Mixed Pinus sylvestris – Betula woodland.
Land use: nature conservation and research – 100%.
•
D2.3 Transition mires and quaking bogs; area: 6400 ha; trend: stable; area data quality:
medium; trend data quality: medium.
• G1.51 Sphagnum Betula woods; area: 2000 ha; trend: stable; area data quality: medium;
trend data quality: poor.
Literature
1. Андрієнко Т.Л. ПЗ Рівненський // Фіторізноманіття заповідників і національних
природних парків України. Ч. 1. Біосферні заповідники. Природні заповідники / під
ред. В.А. Онищенка і Т.Л. Андрієнко. – Київ: Фітосоціоцентр, 2012. – С. 313–324.
2. Андрієнко Т.Л., Прядко О.І. Болотний масив Сомино на Ровенщині, його наукова
і господарська цінність // Укр. ботан. журн. – 1980. – 36, № 4. – С. 65–69.
3. Андрієнко Т.Л., Прядко О.І., Онищенко В.А. Раритетна компонента флори
Рівненського природного заповідника // Укр. ботан. журн. – 2006. – 63, № 2. –
С. 220–228.
4. Андриенко Т.Л., Шеляг-Сосонко Ю.Р. Растительный мир Украинского Полесья в
аспекте его охраны. – Киев: Наук. думка, 1983. – 215 с.
5. Водно-болотні угіддя України / під ред. Г.Б. Марушевського та І.С.Жарук. – К.:
Чорноморська програма Ветландс Інтернешнл, 2006. – 312 с.
6. Онищенко В.А., Андрієнко Т.Л., Прядко О.І. Рослинність ділянки Сомино
Рівненського природного заповідника // Науковий вісник Волинського національного
університету імені Лесі Українки. Біологічні науки. – 2009. – № 9. – С. 173–187.
7. Фіторізноманіття Українського Полісся та його охорона / Під заг. ред. Т.Л.
Андрієнко. – Київ: Фітосоціоцентр, 2006. – 316 с.
Sovyi Yar
Protected areas: included in Rivnenskyi nature reserve, included in Emerald Site “Rivnenskyi
Nature Reserve”.
Threats: –.
General description. Transiion mire dominated by Carex lasiocarpa and peat mosses
(Sphagnum centrale, S. fallax, S. cuspidatum, S. subsecundum, S. magellanicum, S. palustre).
Species of high constancy are Eriophorum gracile, Lysimachia thyrsilora, Lysimachia vulgaris,
Menyanthes trifoliata, Pedicularis palustris, Peucedanum palustre, Phragmites australis,
Potenilla palustris, Rhynchospora alba; in more oligotrophic condiions – Andromeda
polifolia, Drosera rotundifolia, Vaccinium oxycoccos (Oxycoccus palustris); in more eutrophic
condiions – Carex elata, Cnidium dubium, Lythrum salicaria, Mentha arvensis. A porion of
the mire has a tree layer of Betula pubescens. In some places, Carex rostrata, Phragmites
australis, Eriophorum gracile, Carex omskiana dominate. Higher levels are occupied by
Pinus sylvestris woods.
Botanical significance. One of the largest transiion mires in Ukraine.
Criterion C
278
V.A. Onyshchenko
Ukrainian name: Совий яр.
Transliteraion/Translaion variants: Sovii Ravine.
Area: 1081.0 ha.
Altitude: 125–280 m.
Latitude: 48°39’49” N (48.6636°).
Longitude: 26°53’11” E (26.8863°).
Administrative regions. Khmelnytskyi region: Kamianets-Podilskyi raion.
Ownership: state.
Biogeographic regions: coninental.
Habitats. Level 1. C – 0.3%; E – 1.5%; G – 96.2%; H – 1.0%; I – 0.5%; J – 0.5%.
Habitats. Level 2. C2 Surface running waters – 0.3%; E1 – Dry grasslands – 0.7%; E2 – Mesic
grasslands – 0.8%; G1 Broadleaved deciduous woodland – 94.4%; G3 Coniferous woodland
– 0.2%; G5 Lines of trees, small anthropogenic woodlands, recently felled woodland, earlystage woodland and coppice – 1.6%; H5 Miscellaneous inland habitats with very sparse or
no vegetation – 1.0%.
Futher habitat description. G1.A Meso- and eutrophic Quercus, Carpinus, Fraxinus, Acer,
Tilia, Ulmus and related woodland.
Land use: forestry – major, nature conservaion and research – major
Protected areas: includes Sovyi Yar state landscape reserve (827 ha), included in Podilski
279
Tovtry naional nature park, included in Emerald Site “Podilski Tovtry Naional Nature Park”.
Threats: –.
General description. A wood with the dominance of Quercus robur and Carpinus betulus in
the tree layer.
Botanical significance. Important for endemic species Aconitum besserianum.
Criterion A
• Aconitum besserianum Andrz. ex Trautv.; A(iii); abundance: unknown; trend: unknown;
species data quality: poor; trend data quality: poor.
Literature
1. Червона книга України. Рослинний світ / за ред. Я.П. Дідуха. – К.: Глобалконсалтинг,
2009. – 900 с.
Striltsivskyi Step
Ukrainian name: Стрільцівський степ.
Transliteraion/Translaion variants: Stril’tsivs’kyi Steppe.
Area: 1036.5 ha.
Altitude: 120–182 m.
Latitude: 49°17’35” N (49.2930°).
Longitude: 40°04’43” E (40.0786°).
280
V.A. Onyshchenko
Administrative regions. Luhansk region: Milove raion.
Ownership: state.
Biogeographic regions: steppic.
Habitats. Level 1. E – 68.3%; D – 0.03%; F – 31.0%; G – 0.4%; H – 0.2%.
Habitats. Level 2. D5 Sedge and reedbeds, normally without free-standing water – 0.03%;
E1 Dry grasslands – 67.8%; E2 Mesic grasslands – 0.5%; E6 Inland salt steppes – 0.02%;
F3 Temperate and mediterranean-montane scrub – 31.0%; F9 Riverine and fen scrubs –
0.01%; G1 Broadleaved deciduous woodland – 0.4%; H2 Screes – 0.02%; H5 Miscellaneous
inland habitats with very sparse or no vegetation – 0.2%.
Futher habitat description. E1.1 Inland sand and rock with open vegetation; E1.2
Perennial calcareous grassland and basic steppes; E2.5 Meadows of the steppe zone; E6.2
Continental inland salt steppes; D5.1 Reedbeds normally without free-standing water; D5.2
Beds of large sedges normally without free-standing water; F3.1 Temperate thickets and
scrub; F3.2 Submediterranean deciduous thickets and brushes; F9.1 Riverine scrub; G1.1
Riparian and gallery woodland, with dominant Alnus, Betula, Populus or Salix; G1.C Highly
artificial broadleaved deciduous forestry plantations; H2.6 Calcareous and ultra-basic
screes of warm exposures; H5.3 Sparsely- or un-vegetated habitats on mineral substrates
not resulting from recent ice activity; H5.6 Trampled areas.
Land use: nature conservaion and research – 100%.
Protected areas: included in Luhanskyi nature reserve (same as Striltsivskyi Step division of
the reserve), included in Emerald Site “Luhanskyi Nature Reserve”.
Threats: abandonment/reducion of land management – low.
General description. The predominant vegetaion of the area is the steppe, both virgin (446
ha) and old fallows (270 ha). The second major vegetaion type is the steppe shrubs (320 ha).
Among the steppe communiies the shrub steppes have the largest area. The most common
combinaions of dominants are Sipa irsa with Caragana frutex or Amygdalus nana, less
common – with Chamaecyisus ruthenicus. Steppes without co-dominance of shrubs also
occupy a large area. Usually they are dominated by Sipa irsa and Sipa zalesskii. Another
281
important species: Achillea pannonica, Adonis wolgensis, Bromopsis riparia, Elytrigia
intermedia, Elytrigia sipifolia, Elytrigia trichophora, Festuca rupicola, Galatella villosa,
Helictotrichon schellianum, Hierochloe repens, Koeleria cristata, Limonium platyphyllum,
Medicago romanica, Phleum phleoides, Phlomoides tuberosa (Phlomis tuberosa), Poa
angusifolia, Potenilla schuri, Salvia nutans, Salvia stepposa, Sipa capillata, Sipa dasyphylla,
Sipa pennata, Sipa pulcherrima, Veronica spicata. Besides there occur Sipa pulcherrima
steppes with calciphilous species Astragalus albicaulis, Gypsophila oligosperma, Hedysarum
grandilorum, Onosma tanaiica, Rhaponicoides ruthenica (Centaurea ruthenica); sand
steppes dominated by Sipa borysthenica and Festuca beckeri with Artemisia marschalliana,
Astragalus varius, Jurinea centauroides, Potenilla incana, Pulsailla patens, Pulsailla
pratensis; salt steppes dominated by Festuca valesiaca and Galatella dracunculoides with
Artemisia santonica, Bulbocodium versicolor (Colchicum versicolor), Ferula tatrica, Kochia
prostrata, Tulipa ophiophylla. Shrub vegetaion is represented by the steppe shrubs (Prunus
stepposa, Caragana frutex, Amygdalus nana, Chamaecyisus ruthenicus), the wood shrubs
with small trees (Acer tataricum, Euonymus europaeus, Euonymus verrucosus, Malus
praecox, Rhamnus catharica, Sambucus nigra, Swida sanguinea (Cornus sanguineus),
Ulmus laevis, Ulmus pumila) and the loodplain Salix triandra and Salix cinerea shrubs (small
area). Forest vegetaion (4 ha) is represented by ariicial forest belts (Fraxinus lanceolata)
and loodplain Salix alba woods.
Botanical significance. Important area for conservaion of steppe shrubs and steppes.
Criterion A
• Diplotaxis cretacea Kotov; A(iv); abundance: occasional; trend: stable; species data
quality: good; trend data quality: medium.
• Paeonia tenuifolia L.; A(ii); abundance: frequent; trend: stable; species data quality:
good; trend data quality: medium.
Criterion C
• E1.2 Perennial calcareous grassland and basic steppes; area: 710 ha; trend: decreasing;
area data quality: good; trend data quality: good.
• F3.247 Ponto-Sarmatic deciduous thickets; area: 150 ha; trend: increasing; area data
quality: medium; trend data quality: medium.
Literature
1. Білик Г.І., Ткаченко В.С. Рослинний покрив Стрільцівського степу // Укр. ботан.
журн. – 1971. – 28, № 5. – С. 613–617.
2. Біорізноманітність Луганського природного заповідника НАН України / сост.
Т. Сова. – Луганськ: ЛГСІ, 2005. – 218 с.
3. Биоразнообразие Луганского природного заповедника: растительный мир /
Составители: Сова Т. В., Русина Н. В., Гузь Г. В., Боровик Л. П., Шиян-Глотова А. В. –
Луганск: Элтон–2, 2009. – 130 с.
4. Бойко М.Ф. Мохоподібні заповідників “Стрільцівський степ” і “Михайлівська
цілина” // Укр. ботан. журн. – 1981. – 38, № 4. – С. 27–31.
5. Боровик Л.П. Природні та антропогенні фактори демутації перелогів на
території Стрільцівського степу (відділення Луганського природного заповідника) //
Чорноморський ботан. журн. – 2008. – 4, № 1. – С. 98–106.
6. Боровик Л.П. Растительность залежей как важный компонент сохранения
биоразнообразия на востоке Украины (Луганская область) // Вісник Одеського
національного університету. – 2008. – 13, вып. 16. – С. 69–73.
7. Боровик Л.П. Постпирогенная динамика растительных сообществ Стрельцовской
степи // Збірка тез доповідей міжнародної конференції V-ї ботанічні читання пам’яті
Й.К. Пачоського. – Херсон: Айлант. – 2009. – С. 101.
8. Кондратюк Е.Н., Бурда Р.И., Чуприна Т.Т., Хомяков М.Т. Луганский гocударственный
заповедник. Растительный мир. – Киев: Наук. думка, 1988. – 188 с.
9. Маслова В.Р. Ліхенофлора Провальского степу // Зб. наук. праць “Інтродукція і
акліматизація рослин на Україні”. – 1979. – Вип. 15. – С. 51–54.
10. Приходькова Л.П. Синьозелені водорості в грунтах заповідника “Провальський
степ” // Укр. ботан. журн. – 1988. – 45, № 2. – С. 9–13.
11. Русіна Н.В. Історія та сучасний стан дослідження ліхенобіоти Луганського
природного заповідника // Наукові праці Луганського природного заповідника.
Рослинний і тваринний світ та його охорона. – Луганськ. – 2011. – Вип. 2. – С. 47–53
12. Русина Н.В., Ходосовцев А.Е. Лихенобиота Стрельцовской cтепи // VI
міжнародна наукова конференція “Промислова ботаніка: стан та перспективи
розвитку”. – Донецьк. – 2010. – 405–407.
13. Сова Т.В., Боровик Л.П. ПЗ Луганський // Фіторізноманіття заповідників
і національних природних парків України. Ч. 1. Біосферні заповідники. Природні
заповідники / під ред. В.А. Онищенка і Т.Л. Андрієнко. – Київ: Фітосоціоцентр, 2012. –
С. 221–253.
14. Ткаченко В.С., Чуприна Т.Т., Бакланов О.В. Заповідник Провальський степ
(сучасний стан і завдання наукових досліджень) // Укр. ботан. журн. – 1979. – 36, № 4.
– С. 352–356.
15. Ткаченко В.С. Изучение особенностей резерватных сукцессий Стрельцовской
степи по материалам периодического картирования // Геоботаническое
картографирование. – 1989. – С. 47–61.
16. Ткаченко В.С. Особливості екотопічних змін та механізму становлення
потенційних фітоценоструктур Стрільцівського степу // Укр. ботан. журн. – 1996. – 53,
№ 5. – С. 527–535.
17. Ткаченко В.С. Прогноз розвитку рослинності та збереження біорізноманітності
“Стрільцівського степу” // Укр. ботан. журн. – 1996. – 53, № 6. – С. 645–652.
18. Ткаченко В.С. “Стрільцівський степ” в фітоценотичному моніторингу
Старобільських степів // Вісті Біосферного заповідника “Асканія-Нова”. – 2009. – Т. 11. –
С. 6–19.
19. Ткаченко B.C., Боровик Л.П., Сова Т.В., Лисенко Г.М. Структура рослинного
покриву ділянки розширення “Стрільцівського степу” (Луганська обл., Україна) // Вісті
Біосферного заповідника “Асканія-Нова”. – 2009. – Т. 11. – С. 35–47.
20. Ткаченко B.C., Сова Т.В., Боровик Л.П. Еколого-генетичний ряд рослинності
“Стрільцівського степу” на Луганщині // Зб. ст. до 100-річчя д.б.н., проф. Г.І. Білика
“Степові і галофітні екосистеми України”. – К. – 2004. – С. 595–620.
21. Ткаченко В.С., Чуприна Т.Т. Зміни в рослинному покриві Стрільцівського степу
за даними фітоценологічного моніторингу // Укр. ботан. журн. – 1995. – 52, № 2. –
С. 252–259.
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283
Stuzhytsia
Ukrainian name: Стужиця.
Area: 11208 ha (10072.0) ha.
Altitude: 355–1250 m.
Latitude: 49°02’04” N (49.0344°).
Longitude: 22°39’32” E (22.6589°).
Administrative regions. Zakarpatska region: Velykyi Bereznyi raion.
Ownership: state, private.
Biogeographic regions: alpine.
V.A. Onyshchenko
Habitats. Level 1. C – 0.1%; D – 0.1%; E – 24.3%; G – 74.4%; H – 1.0%.
Habitats. Level 2. C2 Surface running waters – 0.1%; D2 Valley mires, poor fens and transition
mires – 0.1%; E1 Dry grasslands – 4.5%; E2 Mesic grasslands – 17.0%; E3 Seasonally wet
and wet grasslands – 0.8%; E5 Woodland fringes and clearings and tall forb stands – 2.0%;
G1 Broadleaved deciduous woodland – 70.0%; G3 Coniferous woodland – 1.3%; G4 Mixed
deciduous and coniferous woodland – 3.1%; H5 Miscellaneous inland habitats with very
sparse or no vegetation – 1.0%.
Futher habitat description. C2.1 Springs, spring brooks and geysers; C2.2 Permanent nontidal, fast, turbulent watercourses; D2.2 Poor fens and soft-water spring mires; E1.7 NonMediterranean dry acid and neutral closed grassland; E2.2 Low and medium alitude hay
meadows; E3.4 Moist or wet eutrophic and mesotrophic grassland; E4.3 Acid alpine and
subalpine grassland; E5.2 Thermophile woodland fringes; E5.4 Moist or wet tall-herb and
fern fringes and meadows; E5.5 Subalpine moist or wet tall-herb and fern stands; G1.6
Fagus woodland; G3.F Highly artificial coniferous plantations; G4.6 Mixed Abies – Picea
284
– Fagus woodland; H2.3 Temperate-montane acid siliceous screes; H5.6 Trampled areas.
Land use: agriculture (animals) – minor, forestry – minor, mowing/hay making – 25%;
nature conservation and research – major.
Protected areas: included in Uzhanskyi naional nature park, overlaps with World Heritage
Site “Primeval Beech Forests of the Carpathians and the Ancient Beech Forests of Germany”
(6147 ha; core zone: 2532 ha, bufer zone: 3615 ha), included in biosphere reserve “East
Carpathians”, included in Emerald Site “Uzhanskyi Naional Nature Park”.
Threats: abandonment/reducion of land management – low, agricultural intensification/
expansion (general) – low.
General description. The largest area is covered by Fagus sylvaica forests. Dominants
of the herb layer are Dentaria glandulosa, Dentaria bulbosa, Dryopteris filix-mas, Galium
odoratum, Lamium galeobdolon, Rubus hirtus, Symphytum cordatum. Other typical species
are Actaea spicata, Anemone nemorosa, Anemone ranunculoides, Corydalis cava, Corydalis
solida, Oxalis acetosella, Polygonatum verticillatum, Salvia glutinosa, Senecio ovatus.
In acidophilous beech forests, the herb layer consists mainly of Luzula luzuloides, Luzula
sylvatica, Prenanthes purpurea, Rubus hirtus, Festuca drymeja, Galium odoratum, Dryopteris
filix-mas, Oxalis acetosella, Maianthemum bifolium. On the steep slopes with stony soils,
there are forests dominated by Acer pseudoplatanus with signiicant paricipaion of Ulmus
glabra. Their typical species are Sambucus nigra, Ribes uva-crispa, Rubus hirtus, Pulmonaria
obscura, Glechoma hirsuta, Polystichum braunii, Phyllitis scolopendrium, Dryopteris filix-mas.
Hay meadows are dominated by Arrhenatherum elatior, Festuca rubra, Cynosurus cristatus.
Other typical species are Campanula patula, Galium mollugo, Knautia arvensis, Crepis
biennis.
Botanical significance. One of the best areas of beech forests in Ukraine. A good example of
mountain low and medium alitude hay meadows.
Criterion C
• E2.2 Low and medium alitude hay meadows; area: 1350 ha; trend: stable; area data
quality: medium; trend data quality: medium.
• G1.6 Fagus woodland; area: 8100 ha; trend: stable; area data quality: medium; trend
data quality: medium.
• G1.A4 (*9180) Ravine and slope woodland; area: 50 ha; trend: stable; area data quality:
medium; trend data quality: poor.
Literature
1. Hadač E., Stoyko S.M., Terray J., Tasenkevich L., Bural M. Notes on plant communities
of the protected complex Stuzhytsia – a part of the trilateral Polish-Slovakian-Ukrainian
biosphere reserve “The Eastern Carpathians” // Укр. ботан. журн. – 1995. – 52, № 5. – С.
686–695.
2. Hadač E., Stoyko S.M., Terray J., Tasenkevich L., Bural M. Notes on the flora and
vegetation of the botanical reserve “Stinka” (Biosfere reserve “The Eastern Carpathians”) //
Укр. ботан. журн. – 1996. – 53, № 1/2. – С. 105–111.
3. Кваковська І.М. НПП Ужанський // Фіторізноманіття заповідників і національних
природних парків України. Ч. 2. Національні природні парки / під ред. В.А. Онищенка і
Т.Л. Андрієнко. – Київ: Фітосоціоцентр, 2012. – С. 506–518.
4. Крічфалушій В.В., Лесьо І.М. Раритетні види рослин Ужанського національного
природного парку // Укр. ботан. журн. – 2004. – 61, № 1. – С. 27–35.
5. Ужанський національний природний парк / Крічфалушій В.В., Луговой О.Є.,
Іванега І.Ю. та ін. – Ужгород: ЕкоЦентр – Тиса, 2001. – 120 с. – (Серія “Збереження
біорізноманіття”: кн. 5).
285
6. Ужанський національний природний парк. Поліфункціональне значення /
С.М. Стойко, Е. Гадач, Л.О. Тасєнкевич та ін.; під ред. С.М. Стойко. – Львів: Меркатор,
2007. – 306 с.
7. Устименко П.М., Попович С.Ю. Продромус рослинності Стужицького масиву
Карпатського біосферного заповідника // Укр. ботан. журн. – 1995. – 52, № 3. – С. 414–
419.
8. Устименко П.М., Попович С.Ю. Висотна диференціація рослинності Стужицького
масиву Карпатського біосферного заповідника // Укр. ботан. журн. – 1996. – 53, № 3. –
С. 703–707.
9. Фіторізноманіття національних природних парків України / Під заг. ред.
Т.Л. Андрієнко та В.А. Онищенка. – Київ: Науковий світ, 2003. – 143 с.
Sviai Hory
Ukrainian name: Святі гори.
Transliteraion/Translaion variants: Svyai Gory, Holy Hills.
Area: 18207.0 ha.
Altitude: 58–211 m.
Latitude: 49°01’52” N (49.0311°).
Longitude: 37°31’30” E (37.5250°).
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V.A. Onyshchenko
Administrative regions. Donetsk region: Sloviansk raion, Lyman (Krasnyi Lyman) raion
Ownership: state.
Biogeographic regions: steppic.
Habitats. Level 1. C – 0.4%; D – 0.1%; E – 1.2%; F – 0.1%; G – 96.5%; H – 1.5% , J – 0.2%
Habitats. Level 2. C1 Surface standing waters – 0.1%; C2 Surface running waters – 0.2%;
C3 Littoral zone of inland surface waterbodies – 0.1%; D5 Sedge and reedbeds, normally
without free-standing water – 0.1%; E1 Dry grasslands – 0.1%; E2 Mesic grasslands – 0.1%;
E3 Seasonally wet and wet grasslands – 1.0%; F3 Temperate and mediterranean-montane
scrub – 0.1%; G1 Broadleaved deciduous woodland – 61.9%; G3 Coniferous woodland
– 26.5%; G4 Mixed deciduous and coniferous woodland – 9.2%; H2 Screes – 0.01%; H5
Miscellaneous inland habitats with very sparse or no vegetation – 1.5%; J4 Transport
networks and other constructed hard-surfaced areas – 0.2%.
Futher habitat description. C1.2 Permanent mesotrophic lakes, ponds and pools; C2.3
Permanent non-tidal, smooth-flowing watercourses; E1.2 Perennial calcareous grassland
and basic steppes; E1.9 Open non-Mediterranean dry acid and neutral grassland, including
inland dune grassland (× E1.1); E3.4 Moist or wet eutrophic and mesotrophic grassland;
F3.1 Temperate thickets and scrub; G1.2 Mixed riparian floodplain and gallery woodland;
G1.4 Broadleaved swamp woodland not on acid peat; G1.7 Thermophilous deciduous
woodland; G1.A Meso- and eutrophic Quercus, Carpinus, Fraxinus, Acer, Tilia, Ulmus and
related woodland; G1.B Non-riverine Alnus woodland; G3.4 Pinus sylvestris woodland
south of the taiga; G4.7 Mixed Pinus sylvestris – acidophilous Quercus woodland; H2.6
Calcareous and ultra-basic screes of warm exposures; H5.3 Sparsely- or un-vegetated
habitats on mineral substrates not resulting from recent ice activity; H5.6 Trampled areas;
J4.2 Road networks; J4.3 Rail networks.
Land use: mowing/hay making – 1%; forestry – 85%; nature conservaion and research –
100%; tourism and recreaion – minor.
Protected areas: included in Sviai Hory naional nature park and Emerald Site “Sviai Hory”
(a part of this park belongs to IPA “Zaplava Siverskoho Dintsia”).
Threats: development (recreaion/tourism) – low; forestry (aforestaion) – low.
General description. The area includes the loodplain of the Siverskyi Donets river (762
ha, medium part of the IPA), sand terrace over the loodplain (7602 ha, northern part) and
plateau (2666 ha, southern part). In the loodplain, there prevail Quercus robur forests,
lesser areas are covered by Alnus gluinosa and Populus alba forests. Floodplain meadows
are dominated by Festuca pratensis, Elytrigia repens, Koeleria delavignei, Phleum pratense,
Poa pratensis, Beckmanioa eruciformis, Carex praecox. Dominants of wet habitats are Carex
riparia, Phalaroides arundinacea, Phragmites australis, Carex melanostachya, Eleocharis
palustris. Sand terrace is occupied by Pinus sylvestris and Pinus silvestris – Quercus robur
forests, in depressions – Alnus gluinosa woods. The major dominant of the herb layer in pine
forests is Calamagrosis epigeios, other species of high constancy are Pilosella oicinarum,
Rumex acetosella, Solidago virgaurea, Tanacetum vulgare. Sand grasslands occupy small
area due to ariicial aforestaion. Their dominants are Artemisia tschernieviana, Carex
colchica, Festuca beckeri, Koeleria sabuletorum, Thymus pallasianus. South of the Siverskyi
Donets river, there are mesic forests dominated by Quercus robur, Fraxinus excelsior, with
co-dominance of Acer campestre, Acer platanoides, Tilia cordata. Other constant species:
Anemone ranunculoides, Asarum europaeum, Corydalis solida, Corylus avellana, Dentaria
quinquefolia, Euonymus europaeus, Euonymus verrucosus, Galium odoratum, Lathyrus
vernus, Polygonatum mulilorum, Pulmonaria obscura, Scilla sibirica, Stellaria holostea,
Ulmus minor. Insigniicant area is occupied by outcrops (screes) of chalk which are rich in
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endemic species.
Condiion of broadleaved forests of the southern porion of the territory is good. A big part
of them belongs to the zone of strict protecion of the naional park, another part – to the
zone of regulated recreaion. Mean age of the tree layer is about 90 years. Condiion of
pine woods is medium, but it is beter than in most other places of Ukraine. The tree layer
is mainly of ariicial origin (plantaion of Pinus sylvestris). Species composiion has natural
character and includes endemic species. Northern part of IPA belongs predominantly to the
zone of regulated recreaion that does not allow forest clearing.
Eforts to establish Pinus sylvestris plantaions prevent forming sand steppes which
previously occupied a much larger area.
Botanical significance. Important for conservaion of steppe pine forests, mesic deciduous
forests and steppe black alder woods.
Criterion A
• Centaurea donetzica Klokov; A(iv); abundance: rare; trend: unknown; species data
quality: medium; trend data quality: poor.
• Iris humilis Georgi (Iris pineicola Klokov); A(ii), A(iv); abundance: frequent; trend: stable;
species data quality: good; trend data quality: medium.
Criterion C
• G1.414 Steppe swamp Alnus gluinosa woods; area: 300 ha; trend: stable; area data
quality: medium; trend data quality: medium.
• G1.A1 Quercus – Fraxinus – Carpinus betulus woodland on eutrophic and mesotrophic
soils; area: 10000 ha; trend: stable; area data quality: good; trend data quality: good.
• G3.4232 Sarmatic steppe Pinus sylvestris forests; area: 4750 ha; trend: stable; area data
quality: medium; trend data quality: medium.
Literature
1. Бурда Р.І., Остапко В.М., Глухов А.З., Шпилева Н.В. Національний природний
парк “Святі Гори”: біологічна різноманітність рослинного покриву // Заповідна справа
в Україні. – 1997. – Т.3, вип. 1. – С. 10–17.
2. Дьякова О.В. Грабово-дубовий ліс в Національному природному парку “Святі
Гори” // Менеджмент екосистем природно-заповідних територій. Мат-ли Всеукр.
наук.-практ. конф., присвяч. 10-річчю створення Національного природного парку
“Подільські Товтри”. – Кам’янець-Подільський: Аксіома, 2006. – С. 93–97.
3. Дьякова О.В. О нахождении Muscari neglectum Guss. на территории НПП “Святые
Горы” // Промислова ботаніка: стан та перспективи розвитку: Мат-ли міжнар. наук.
конф. “Промислова ботаніка: стан та перспективи розвитку” (Донецьк, 24–26 вересня
2007 р.). – Донецьк, 2007. – С.156–157.
4. Дьякова О.В. Рослинність вільхових лісів урочища Святогірський табір (НПП “Святі
Гори”) // Відновлення порушених природних екосистем: Мат-ли Третьої міжнар. наук.
конф. (м. Донецьк, 7–9 жовтня 2008 р.). – Донецьк, 2008. – С. 183–185.
5. Зацепина Д.Я., Хлевная Н.А. О флоре меловых обнажений заказника “Горы
Артема” // Ботан. журн. – 1980. – 65, № 4. – С. 551–555.
6. Мальцева І.А. Грунтові водорості лісів національного природного парку “Святі
Гори” (Донецька обл., Україна) // Мат-ли конф., присвяч. 80-річчю Канівського
природного заповідника “Роль природно-заповідних територій у підтриманні
біорізноманіття” (9–11 вересня 2003 р., м. Канів). – Канів, 2003. – С. 118–119.
7. Мальцева І.А., Супронюк І., Пушкіна О. Грунтові водорості крейдяних
борів національного природного парку “Святі Гори” // 21–30 червня 2004 р., м.
Дніпропетровськ). – Т. 31. Біологічні науки. – Дніпропетровськ: Наука і освіта, 2004. –
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С. 35.
8. Морозова І. Міксоміцети Національного природного пароку “Святі Гори” // Матли ІV Міжн. наук. конф. студентів та аспірантів “Молодь і поступ біології” (м. Львів, 7–10
квітня 2008 р.). – Львів: ЛНУ ім. І. Франка, 2008. – С. 106–107.
9. Надєіна О.В. Лишайники національного природного парку “Святі Гори” //
Чорноморський ботан. журн. – 2007. – 3, № 2. – С. 100–108.
10. Онищенко В.А., Дьякова О.В., Карпенко Ю.О. Лісова рослинність урочищ
Теплинська Дача і Маяцька Дача (національний природний парк “Святі Гори”) //
Чорноморський ботан. журн. – 2007. – 3, № 2. – С. 88–99.
11. Ордынец А.В., Акулов А.Ю. Предварительные данные о биоте
афиллофороидных грибов Национального природного парка “Святые Горы” (Донецкая
обл., Украина) // Актуальные проблемы Экологии и сохранения биоразнообразия. Сб.
статей. – Владикавказ. – СОИТСИ, 2008. – С. 64–71.
12. Остапко В.М., Шпилева Н.В., Дьякова О.В. НПП Святі гори // Фіторізноманіття
заповідників і національних природних парків України. Ч. 2. Національні природні
парки / під ред. В.А. Онищенка і Т.Л. Андрієнко. – Київ: Фітосоціоцентр, 2012. –
С. 440–456.
13. Тихоненко Ю.Я., Дудка І.О. Знахідки іржастих грибів (порядок Uredinales) в
національному природному парку “Святі Гори” // Укр. ботан. журн. – 2005. – 62, № 4. –
С. 495–499.
14. Трискиба С.Д., Полохіна І.І., Сухомлин М.М. Знахідка Grifola frondosa (Fr.) S.
Gray на півночі Донецької області // Укр. ботан. журн. – 2005. – 62, № 9. – С. 87–90.
15. Устименко П.М., Попович С.Ю. Растительность проектируемого
Славяногорского национального парка и зонирование его территории // Бюл. ботан.
сада. – 1992. – Вып.164. – С.76–81.
16. Шпилевая Н.В. Синантропная флора национального природного парка
“Святые Горы” // Відновлення порушених природних екосистем: мат-ли другої міжн.
конф., 6–8 верес. 2005 р.: тезисы докл. – Донецьк: ТОВ “Лебідь”, 2005. – С. 109–110.
17. Шпилева Н.В. Оцінка антропогенної трансформації флори національного
природного парку “Святі Гори” // Синантропізація рослинного покриву України: наук.
конф.: тези. допов. – Київ, Переяслав-Хмельницький, 2006. – С. 220–223.
Svydovets
V.A. Onyshchenko
Ukrainian name: Свидовець.
Area: 21167.0 ha.
Altitude: 570–1883 m.
Latitude: 48°16’05” N (48.2681°).
Longitude: 24°10’30” E (24.1749°).
Administrative regions. Zakarpatska region: Tiachiv raion, Rakhiv raion.
Ownership: state.
Biogeographic regions: alpine.
Habitats. Level 1. C – 0.2%; D – 0.1%; E – 35.6%; F – 9.7%; G – 54.4%; H – 0.5%.
Habitats. Level 2. C1 Surface standing waters – 0.0%; C2 Surface running waters – 0.2%; E4
Alpine and subalpine grasslands stands – 35.0%; E5 Woodland fringes and clearings and tall
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forb stands – 0.6%; F2 Arctic, alpine and subalpine scrub – 9.7%; G1 Broadleaved deciduous
woodland – 29.0%; G3 Coniferous woodland – 22.1%; G4 Mixed deciduous and coniferous
woodland – 3.6%; H2 Screes – 0.0%; H3 Inland cliffs, rock pavements and outcrops – 0.0%;
H5 Miscellaneous inland habitats with very sparse or no vegetation – 0.5%.
Futher habitat description. C1.1 Permanent oligotrophic lakes, ponds and pools; C1.2
Permanent mesotrophic lakes, ponds and pools; C2.1 Springs, spring brooks and geysers;
C2.2 Permanent non-tidal, fast, turbulent watercourses; E4.1 Vegetated snow-patch; E4.2
Moss and lichen dominated mountain summits, ridges and exposed slopes; E4.3 Acid alpine
and subalpine grassland; E4.4 Calcareous alpine and subalpine grassland; E5.4 Moist or wet
tall-herb and fern fringes and meadows; E5.5 Subalpine moist or wet tall-herb and fern
stands; F2.2 Evergreen alpine and subalpine heath and scrub; F2.3 Subalpine deciduous
scrub; F2.4 Conifer scrub close to the tree limit; G1.1 Riparian and gallery woodland, with
dominant Alnus, Betula, Populus or Salix; G1.6 Fagus woodland; G3.1 Abies and Picea
woodland; G4.6 Mixed Abies – Picea – Fagus woodland; H2.3 Temperate-montane acid
siliceous screes; H2.4 Temperate-montane calcareous and ultra-basic screes; H3.1 Acid
silicious inland clifs; H3.2 Basic and ultra-basic inland clifs; H5.6 Trampled areas.
Land use: agriculture (animals) – major, forestry – minor, nature conservation and research
– major, tourism/recreaion – minor.
Protected areas: overlaps (8523 ha) with Karpatskyi biosphere reserve (naional category),
overlaps (8523 ha) with Carpathian biosphere reserve (UNESCO), overlaps (8492 ha) with
Emerald Site “Carpathian Biosphere Reserve”, overlaps (12644 ha) with Emerald Site
“Zakhidnyi Svydovets”, overlaps (8670 ha) with World Heritage Site “Primeval Beech Forests
of the Carpathians and the Ancient Beech Forests of Germany” (core zone: 3030.5 ha, bufer
zone: 5639.5 ha).
Threats: climate change/sea level rise – low, development (recreaion/tourism) – low.
General description. One of the highest parts of the Ukrainian Carpathians. Large areas
are occupied by the spruce forests, beech forests and the subalpine vegetaion. Picea abies
forests prevail on the northeastern slope of the Svydovets Range. Upper limit of the forest
vegetaion is formed by pure Picea abies forests. Typical dominants are Calamagrosis
arundinacea, Luzula sylvaica, Vaccinium myrillus, Pleurozium schreberi, Polytrichum
formosum. At lower alitudes, there are mixed forests dominated by Picea abies, Abies alba
and Fagus sylvaica. Pure Fagus sylvaica forests occupy a large area on the southern slope.
In the herb layer, there predominate Dentaria glandulosa (Cardamine glandulosa), Galium
odoratum, Rubus hirtus. In the subalpine belt, signiicant areas are dominated by Juniperus
sibirica, Duschekia alnobetula (Alnus alnobetula), Nardus stricta, Deschampsia cespitosa,
Vaccinium myrillus, Calamagrosis villosa, Festuca airoides, Oreojuncus triidus (Juncus
triidus). Someimes there dominate Festuca picta, Luzula alpinopininosa, Rhododendron
myrifolium, Vaccinium uliginosum. In the wet habitats, there occur communiies of
Adenostyles alliariae, Cirsium waldsteinii. Calcareous rocky grasslands dominated by Festuca
inarmata (Festuca amethysina s.l.) or Festuca versicolor have small areas.
Botanical significance. Important area for subalpine habitats and species, beech forests and
oligotrophic aquaic habitats.
Criterion A
• Botrychium matricariifolium (A. Braun ex Döll) W.D.J.Koch; A(ii); abundance: rare; trend:
unknown; species data quality: medium; trend data quality: poor.
• Campanula abieina Griseb. & Schenk; A(ii); abundance: occasional; trend: stable;
species data quality: medium; trend data quality: poor.
• Hamatocaulis vernicosus (Mitt.) Hedenas (Drepanocladus vernicosus (Mitt.) Warnst.);
A(ii); abundance: occasional; trend: unknown; species data quality: medium; trend data
quality: poor.
• Heterophyllium affine (Mitt.) Fleisch; A(ii); abundance: rare; trend: unknown; species
data quality: medium; trend data quality: poor.
• Narcissus angustifolius Curtis; A(ii); abundance: rare; trend: unknown; species data
quality: medium; trend data quality: poor.
• Neckera pennata Hedw.; A(ii); abundance: rare; trend: unknown; species data quality:
medium; trend data quality: poor.
• Poa graniica Braun-Blanq. (Poa deylii Chrtek & V.Jirasek); A(ii); abundance: rare; trend:
unknown; species data quality: medium; trend data quality: poor.
Criterion A
• C1.1 Permanent oligotrophic lakes, ponds and pools; area: 2 ha; trend: decrasing; area
data quality: medium; trend data quality: medium.
• D2.226 Peri-Danubian black-white-star sedge fens; area: 3 ha; trend: stable; area data
quality: poor; trend data quality: poor.
• E4.11 Boreo-alpine acidocline snow-patch grassland and herb habitats; area: 10 ha;
trend: stable; area data quality: poor; trend data quality: poor.
• E4.12 Boreo-alpine calcicline snow-patch grassland and herb habitats; area: 1 ha; trend:
stable; area data quality: poor; trend data quality: poor.
• E4.3 Acid alpine and subalpine grassland; area: 7300 ha; trend: stable; area data quality:
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poor; trend data quality: poor.
E4.4 Calcareous alpine and subalpine grassland; area: 10 ha; trend: stable; area data
quality: poor; trend data quality: poor.
• E5.5 Subalpine moist or wet tall-herb and fern stands; area: 400 ha; trend: stable; area
data quality: poor; trend data quality: poor.
• F2.22 Alpide acidocline Rhododendron heaths; area: 30 ha; trend: stable; area data
quality: poor; trend data quality: medium.
• F4.2 Dry heaths; area: 10 ha; trend: stable; area data quality: poor; trend data quality:
medium.
• G1.12 Boreo-alpine riparian galleries; area: 40 ha; trend: stable; area data quality: poor;
trend data quality: medium.
• G3.1B Alpine and Carpathian subalpine Picea forests; area: 2000 ha; trend: stable; area
data quality: medium; trend data quality: medium.
• G1.6 Fagus woodland; area: 5800 ha; trend: stable; area data quality: medium; trend
data quality: medium.
• H2.3 Temperate-montane acid siliceous screes; area: 40 ha; trend: stable; area data
quality: poor; trend data quality: medium.
Conservaion proposals. Create an Emerald Sites including the enire IPA and a state reserve
in the northwestern part of the IPA outside the Carpathian biosphere reserve.
Literature
1.Бедей М.І. Близниці-Драгобрат (флора, рослинність, охорона). – Ужгород: Ліра,
2006. – 108 с.
2.Біорізноманіття Карпатського біосферного заповідника / Кол. авт., Ред. рада:
Я.І. Мовчан, Ф.Д. Гамор та ін. – К.: Інтерекоцентр, 1997. – 711 с.
3.Гамор Ф.Д., Вайнагій І.В., Антосяк В.М. Стан охорони червонокнижних видів
рослин на заповідних територіях Українських Карпат // Укр. ботан. журн. – 1994. – 51,
№ 5. – С. 122–129.
4.Гамор Ф.Д., Волощук М.І., Антосяк Т.М., Козурак А.В. БЗ Карпатський //
Фіторізноманіття заповідників і національних природних парків України. Ч. 1. Біосферні
заповідники. Природні заповідники / під ред. В.А. Онищенка і Т.Л. Андрієнко. – Київ:
Фітосоціоцентр, 2012. – С. 45–72.
5.Гамор Ф.Д., Довганич Я.О., Покиньчереда В.Ф. та ін. Праліси Закарпаття.
Інвентаризація та менеджмент. – Рахів, 2008. – 86 с.
6.Малиновський К.А. Рослинність високогір’я Українських Карпат. – К.: Наук. думка,
1980.– 280 с.
7.Малиновський К.А., Крічфалушій В.В. Рослинні угруповання високогір’я
Українських Карпат. – Ужгород, 2002. – 244 с.
8.Праліси в центрі Європи: Путівник по лісах Карпатського біосферного заповідника
/ Кол. авт., ред.: У-Б. Брендлі, Я. Довганич. – Бірменсдорф: Швейцарський федеральний
ін-т досліджень лісу, снігу і ландшафтів (WSL) – Рахів: Карпатський біосферний
заповідник (КБЗ), 2003. – 192 с.
•
Syra Pohonia
V.A. Onyshchenko
Ukrainian name: Сира Погоня.
Area: 18811 ha.
Altitude: 145–160 m.
Latitude: 51°31’33” N (51.5258°).
Longitude: 27°10’12” E (27.1699°).
Administrative regions. Rivne region: Dubrovytsia raion (minor), Rokytne raion.
Ownership: state, private (minor).
Biogeographic regions: continental.
Habitats. Level 1. D – 51%; G – 49%.
Habitats. Level 2. D1 Raised and blanket bogs – 30%; D2 – Valley mires, poor fens and
transition mires – 21%; G1 Broadleaved deciduous woodland – 10%; G3 Coniferous
woodland – 28%; G4 Mixed deciduous and coniferous woodland – 11%.
Futher habitat description. D1.1 Raised bogs; D2.3 Transition mires and quaking bogs; E1.7
Closed non-Mediterranean dry acid and neutral grassland; E1.9 Open non-Mediterranean
dry acid and neutral grassland, including inland dune grassland; G1.5 Broadleaved swamp
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woodland on acid peat; G3.4 Pinus sylvestris woodland south of the taiga; G3.E Nemoral
bog conifer woodland, G4.1 Mixed swamp woodland, G4.4 Mixed Pinus sylvestris – Betula
woodland.
Land use: nature conservation and research – 100%.
Protected areas: overlaps (9926 ha) with Rivnenskyi nature reserve, overlaps (9926 ha) with
Emerald Site “Rivnenskyi Nature Reserve”.
Threats: water (drainage) – low.
General description. Raised bogs, Pinus sylvestris woods, and transiion mires. Dominants
of the raised bogs are Eriophorum vaginatum, Sphagnum magellanicum, Sphagnum fuscum.
Oten they have a sparse layer of Pinus sylvestris and Betula pubescens. In Sphagnum pine
woods, typical dominants are Ledum palustre, Vaccinium uliginosum, Vaccinium oxycoccos
(Oxycoccus palustris). In hollows, Sphagnum cuspidatum, Scheuchzeria palustris, Carex
limosa, Carex rostrata dominate. Higher levels are occupied by Pinus sylvestris woods.
Botanical significance. The largest raised bog in Ukraine.
Criterion C
• D2.3 Transition mires and quaking bogs; area: 3800 ha; trend: stable; area data quality:
poor; trend data quality: medium.
• G3.E Nemoral bog conifer woodland; area: 3000 ha; trend: stable; area data quality:
medium; trend data quality: medium.
• X04 Raised bog complexes; area: 7000 ha; trend: stable; area data quality: medium;
trend data quality: medium.
Conservaion proposals. Create a state reserve that will include the IPA outside Rivnenskyi
nature reserve.
Literature
1. Андрієнко Т.Л. ПЗ Рівненський // Фіторізноманіття заповідників і національних
природних парків України. Ч. 1. Біосферні заповідники. Природні заповідники / під
ред. В.А. Онищенка і Т.Л. Андрієнко. – Київ: Фітосоціоцентр, 2012. – С. 313–324.
2. Андрієнко Т.Л., Прядко О.І., Онищенко В.А. Раритетна компонента флори
Рівненського природного заповідника // Укр. ботан. журн. – 2006. – 63, № 2. – С. 220–
228.
3. Андриенко Т.Л., Шеляг-Сосонко Ю.Р. Растительный мир Украинского Полесья в
аспекте его охраны. – Киев: Наук. думка, 1983. – 215 с.
4. Водно-болотні угіддя України / під ред. Г.Б. Марушевського та І.С. Жарук. – К.:
Чорноморська програма Ветландс Інтернешнл, 2006. – 312 с.
5. Фіторізноманіття Українського Полісся та його охорона / Під заг. ред.
Т.Л. Андрієнко. – Київ: Фітосоціоцентр, 2006. – 316 с.
Tarkhankut
V.A. Onyshchenko
Ukrainian name: Тарханкут.
Area: 14379.0 ha.
Altitude: 0–146 m.
Latitude: 45°22’16” N (45.3711°).
Longitude: 32°34’09” E (32.5692°).
Administrative regions. Krym: Chornomorske raion; Territorial waters of Ukraine.
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Ownership: state.
Biogeographic regions: steppic.
Habitats. Level 1. A – 21.9%; B – 0.01%; E – 69.2%; F – 1.0%; H – 0.9%; I – 7.0%; J – 0.05%.
Habitats. Level 2. A2 Littoral sediment – 0.02%; A3 Infralittoral rock and other hard substrata
– 15.0%; A4 Circalittoral rock and other hard substrata – 7.6%; A5 Sublittoral sediment –
0.2%; B1 Coastal dunes and sandy shores – 0.001%; B2 Coastal shingle – 0.002%; B3 Rock
cliffs, ledges and shores, including the supralittoral – 0.01%; E1 Dry grasslands – 69.2%; F3
Temperate and mediterranean-montane scrub – 0.02%; F7 Spiny Mediterranean heaths
– 0.9%; H1 Terrestrial underground caves, cave systems, passages and waterbodies, H2
Screes – 0.003%; H3 Inland cliffs, rock pavements and outcrops – 0.04%; H5 Miscellaneous
inland habitats with very sparse or no vegetation – 0.9%; I1 Arable land and market gardens
– 7.0%; J2 Low density buildings – 0.01%; J3 Extractive industrial sites – 0.01%; J4 Transport
networks and other constructed hard-surfaced areas – 0.03%.
Futher habitat description. A2.1 Litoral coarse sediment; A2.2 Litoral sand and muddy
sand; A2.4 Litoral mixed sediments; A2.5 Coastal saltmarshes and saline reedbeds;
A3.2 Atlanic and Mediterranean moderate energy infralitoral rock; A3.3 Atlanic and
Mediterranean low energy infralitoral rock; A3.7 Features of infralitoral rock; A4.3 Atlanic
295
and Mediterranean low energy circalitoral rock; A5.1 Sublitoral coarse sediment; A5.2
Sublitoral sand; A5.4 Sublitoral mixed sediments; A5.5 Sublitoral macrophyte-dominated
sediment; E1.1 Inland sand and rock with open vegetation, E1.2 Perennial calcareous
grassland and basic steppes; F3.1 Temperate thickets and scrub; F3.2 Submediterranean
deciduous thickets and brushes; F7.4 Hedgehog-heaths; H2.6 Calcareous and ultra-basic
screes of warm exposures; H3.2 Basic and ultra-basic inland clifs; H3.5 Almost bare rock
pavements, including limestone pavements; H5.3 Sparsely- or un-vegetated habitats on
mineral substrates not resuling from recent ice acivity; H5.6 Trampled areas; I1.5 Bare
illed, fallow or recently abandoned arable land.
Land use: agriculture (animals) – 30%; nature conservaion and research – 77%; tourism/
recreation – major.
Protected areas: includes Charivna Havan naional nature park (10900 ha); includes Emerald
Site “Charivna Havan’ Naional Nature Park” (10900 ha).
Threats: development (recreation/tourism) – medium; eutrophicain – low.
General description. The IPA includes areas occupied mainly by steppes on stony soils in the
western part of the Crimean Peninsula and adjacent 1-km wide strip of the Black Sea. The
shore is a popular place for sea diving.
Dominants of marine vegetaion are Cladostephus spongiosus, Cystoseira crinita, Cystoseira
barbata, Corallina elongata, Phyllophora crispa. Other typical species: Ceramium virgatum,
Chondrophycus papillosus, Cladophora albida, Cladophora sericea, Dictyota fasciola,
Dilophus fasciola, Laurencia coronopus, Laurencia obtusa, Ulva intesinalis, Ulva linza. Dry
grasslands are dominated by: Agropyron pecinatum, Botriochloa ischaemum, Festuca
rupicola, Festuca valesiaca, Koeleria brevis, Sipa capillata, Sipa lessingiana. Steppes of
more xeric habitats difer by presence of Artemisia lerchiana, Artemisia taurica, Astragalus
henningii, Centaurea orientalis, Dianthus capitatus, Galium ruthenicum, Linum tenuifolium,
Medicago romanica, Scabiosa praemontana, Tanacetum paczoskii. Diferenial species of
more mesic habitats are Ajuga salicifolia, Centaurea difusa, Filipendula vulgaris, Galium
verum, Potenilla argentea, Salvia aethiopis, Salvia nemorosa, Salvia nutans, Veronica
spicata. Artemisia-dominated steppes have been formed as a result of intensive grazing.
Their dominants are Artemisia austriaca, Artemisia lerchiana, Artemisia taurica. Other
important species of Artemisia-dominated steppes: Carthamus lanatus, Centaurea solsiialis,
Euphorbia seguieriana, Salvia aethiopis, Sipa capillata, Sipa lessingiana. Diferenial
species of Artemisia-dominated steppes on shallow rendzina soils are Artemisia lerchiana
(dom.), Artemisia caucasica (dom.), Caragana scythica, Convolvulus cantabrica, Convolvulus
holosericeus, Cota dubia (Anthemis dubia), Crambe aspera, Genista albida, Ononis pusilla,
Teucrium chamaedrys, Teucrium polium, Veronica taurica. Typical species of abandoned
arable lands are Centaurea difusa (dom.), Carduus nutans, Eryngium campestre, Euphorbia
seguieriana, Euphorbia stepposa, Euphorbia virgata, Marrubium peregrinum, Salvia
aethiopis. Tomillares are dominated by Thymus roegneri (Th. callieri), Th. dzevanovskyi,
Asphodeline taurica, Jurinea stoechadifolia, Convolvulus holosericeus, Achillea nobilis with
admixture (10–15%) of steppe species. Small areas are occupied by deciduous thickets and
brushes (Berberis vulgaris, Coinus coggygria, Crataegus monogyna, Euonymus verrucosus,
Genista albida, Hedera helix s.l., Jasminus fruicans, Malus sylvestris, Fraxinus excelsior,
Prunus spinosa, Pyrus communis, Pyrus eleagnifolia, Rhamnus catharica, Rosa spp, Rubus
sp. div., Ulmus minor; in the herb layer: Arum elongatum, Asparagus oicinalis, Asparagus
litoralis, Asparagus vericillatus, Crambe mitridais, Dactylis glomerata, Malva erecta,
Thalictrum minus.
Botanical significance. One of the largest areas of steppe vegetaion and one of the most
valuable sites of seaweed communiies in Ukraine.
Criterion A
• Centaurea taliewii Kleopow; A(iv); abundance: unknown; trend: unknown; species data
quality: poor; trend data quality: poor.
• Crambe mitridais Juz.; A(iv); abundance: unknown; trend: unknown; species data
quality: poor; trend data quality: poor.
Criterion C
• A3 Infralittoral rock and other hard substrata; area: 2000 ha; trend: stable; area data
quality: medium; trend data quality: poor.
• E1.2 Perennial calcareous grassland and basic steppes; area: 9800 ha; trend: increasing;
area data quality: good; trend data quality: good.
• F7 Spiny Mediterranean heaths; area: 100 ha; trend: unknown; area data quality: poor;
trend data quality: poor.
Conservaion proposals. Add coastal part of the sea to the Charivna Havan naional nature
park and Emerald Site “Charivna Havan’ Naional Nature Park”.
Literature
1. Белянина Н.Б., Шатко В.Г. Флористические находки с Тарханкутского полуострова
// Бюл. Главн. ботанич. сада. – 1992. – Вып. 164. – С. 57–63.
2. Белянина Н.Б., Шатко В.Г. Конспект флоры Джангульского побережья (Крым) //
Бюл. Главн. ботанич. сада. – 1999. – Вып. 178. – С. 43–65.
3. Голубева И.В., Маслова И.И. Оценка современного состояния и предложения по
заповедной охране степной растительности в связи с организацией Тарханкутского
заповедника // Фонды Никитс. ботан. сада. – Ялта, 1986. – 4 с.
4. Дідух Я.П., Вакаренко Л.П. Флористичні та ценотичні особливості Тарханкутського
півострова (Крим) // Укр. ботан. журн. – 1987. – 44, № 3. – С. 31–36.
5. Маслов И.И. Фитобентос заповедной акватории “Прибрежный аквальный
комплекс у мыса Атлеш” (Черное море) // Экология моря. – 2001. – Вып. 56. – С. 30–34.
6. Маслов И.И. Макрофитобентос некоторых заповедных акваторий Черного моря
(Украина) // Альгология. – 2002. – 12, № 1 – С. 81–95.
7. Мильчакова Н.А. Макрофитобентос // Современное состояние биоразнообразия
прибрежных вод Крыма (черноморский сектор). – Севастополь: ЭКОСИ-Гидрофизика,
2003. – С. 152–208.
8. Мільчакова Н.П., Вахрушева Л.П., Єпіхін Д.В. НПП Чарівна гавань //
Фіторізноманіття заповідників і національних природних парків України. Ч. 2.
Національні природні парки / під ред. В.А. Онищенка і Т.Л. Андрієнко. – Київ:
Фітосоціоцентр, 2012. – С. 519–529.
9. Подгородецкий П.Д. Тарханкутский национальный природный парк //
Заповедники Крыма – 2007. Мат-лы IV междунар. науч.-практ. конф., 2 нояб. 2007 г.,
Симферополь. – Ч. 1. Ботаника. Общие вопросы охраны природы. – Симферополь,
2007. – С. 341–352.
296
297
Habitats. Level 1. A – 75.0%; B – 0.3%; C – 1.8%; D – 2.0%; E – 20.8%; H – 0.1%.
Habitats. Level 2. A2 Littoral sediment – 4.0%; A5 Sublittoral sediment – 71.0%; B1 Coastal
dunes and sandy shores – 0.3%; B2 Coastal shingle – 0.1%; C1 Surface standing waters
– 1.8%; D6 Inland saline and brackish marshes and reedbeds – 2.0%; E1 Dry grasslands
– 1.5%; E2 – Mesic grasslands – 0.2%; E6 Inland salt steppes – 19.1%; H5 Miscellaneous
inland habitats with very sparse or no vegetation – 0.1%.
Futher habitat description. A2.5 Coastal saltmarshes and saline reedbeds; A5.5 Sublitoral
macrophyte-dominated sediment; B1.1 Sand beach driftlines; B1.2 Sand beaches above the
driftline; B1.3 Shifting coastal dunes; B1.4 Coastal stable dune grassland (grey dunes); B1.8
Moist and wet dune slacks; B2.1 Shingle beach driftlines; B2.2 Unvegetated mobile shingle
beaches above the driftline; C1.5 Permanent inland saline and brackish lakes, ponds and
pools; D6.1 Inland saltmarshes; D6.2 Inland saline or brackish species-poor helophyte beds
normally without free-standing water; E6.2 Continental inland salt steppes; H5.3 Sparsely-
or un-vegetated habitats on mineral substrates not resulting from recent ice activity.
Land use: nature conservation and research – 85%.
Protected areas: overlaps (about 44700 ha) with Chernomorskiy (Chornomorskyi, Black
Sea) biosphere reserve (UNESCO, core zone of the reserve) and, Chornomorskyi biosphere
reserve (naional category), overlaps (about 44700 ha) with Emerald Site “Black Sea
Biosphere Reserve”.
Threats: climate change/sea level rise – low.
General description. The IPA includes Gulf of Tendra, smaller lagoons and salt lakes with
adjacent inland salt marshes, salt steppes, accumulaive islands. Gulf of Tendra is up to 2.6 m
depth. Its salinity is 15–18 ‰. Tendra island is a bar situated south to the gulf. It consists
of sand and shells. Dominants of aquaic vegetaion are Zostera nolii (Nanozostera nolii),
Zostera marina, Ruppia spiralis, Zannichellia maior, Potamogeton pecinatus (Stuckenia
pecinata). On lower part of the beach Cakile maritima, Crambe pontica, Salsola kali subsp.
pontica dominates. Typical species are Argusia sibirica, Lactuca tatarica, Xanthium albinum,
Leymus sabulosus. On sand beaches above the dritline, dominants are Leymus sabulosus,
Crambe pontica, Calamagrosis epigeios, Polygonum euxinum, Elytrigia bessarabica. Other
typical species are Asparagus mariimus, Astrodaucus littoralis, Centaurea odessana,
Cynanchum acutum, Eryngium mariimum, Lactuca tatarica, Polygonum mesembricum.
Halophyic vegetaion is represented by the communiies of Juncus mariimus, Halocnemum
strobilaceum, Salicornia perennans, Halocnemum strobilacеum, Frankenia pulverulenta,
Lepidium pumilum, Puccinellia syvaschica, Puccinellia fomini, Puccinellia bilykiana,
Puccinellia brachylepis (~ P. gigantea), Aeluropus litoralis, Bolbochoenus mariimus,
Tripolium vulgare, Scirpus tabernaemontani. Drier salt habitats (salt steppes) are dominated
by Artemisia santonica, Festuca valesiaca, Stipa capillata, Agropyron pectinatum, Elytrigia
pseudocaesia. Typical species: Achillea setacea, Artemisia austriaca, Bromus mollis, Bromus
squarrosus, Festuca pseudodalmatica, Festuca valesiaca, Halimione verrucifera, Limonium
meyeri, Poa bulbosa, Dianthus guttatus, Milium vernale. Dominant species of dry grasslands
on sands and shells (sand steppes) is Festuca beckeri, other typical species are Artemisia
arenaria, Asperula setulosa, Centaurea odessana. In the mesic grasslands, there prevail
Agrostis sabulicola, Calamagrosts epigeios, Cynodon dactylon, Elytrigia elongata, Elytrigia
repens, Phleum phleoides, Poa angustifolia.
Botanical significance. Important area for conservaion of sand steppes, litoral vegetaion,
sand dunes, halophyic vegetaion, vegetaion of salt waters and endemic psammophyic
and halophyic species.
Criterion A
• Allium regelianum A.Beckerex Iljin; A(ii); abundance: occasional; trend: unknown;
species data quality: medium; trend data quality: poor.
• Allium scythicum Zoz; A(iii); abundance: rare; trend: unknown; species data quality:
medium; trend data quality: poor.
• Alyssum borzaeanum Nyár. (Odontarrhena borzaeana (Nyár.) D.A.German); A(ii);
abundance: occasional; trend: stable; species data quality: medium; trend data quality:
poor.
Criterion C
• A2.5 Coastal saltmarshes and saline reedbeds; area: 200 ha; trend: stable; area data
quality: poor; trend data quality: medium.
• A5 Sublitoral sediment; area: 37000 ha; trend: stable; area data quality: good; trend
data quality: medium.
• B1.3 Shifting coastal dunes; area: 50 ha; trend: stable; area data quality: medium; trend
298
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Tendrivska Zatoka
V.A. Onyshchenko, O.Yu. Umanets
Ukrainian name: Тендрівська затока.
Transliteraion/Translaion variants: Tendra Bay, Gulf of Tendra.
Area: 52622.0 ha.
Altitude: 0–3 m.
Latitude: 46°14’48” N (46.2467°).
Longitude: 31°54’11” E (31.9029°).
Administrative regions. Kherson region: Hola Prystan raion; Territorial waters of Ukraine
Ownership: state.
Biogeographic regions: steppic.
data quality: medium.
B1.4 Coastal stable dune grassland (grey dunes); area: 50 ha; trend: stable; area data
quality: medium; trend data quality: medium.
• D6.1 Inland saltmarshes; area: 600 ha; trend: stable; area data quality: poor; trend data
quality: medium.
• E6.2 Continental inland salt steppes; area: 11150 ha; trend: stable; area data quality:
poor; trend data quality: medium.
Literature
1. Бiлик Г.I., Ткаченко В.С. Рослиннiсть урочища Потiївка Чорноморського
державного заповiдника // Укр. ботан. журн. – 1970. – 27. № 4. – С. 491–496.
2. Войтюк Б.Ю., Уманець О.Ю., Соломаха І.В Синтаксономія галофільної рослинності
Чорноморського біосферного заповідника // Науковий вісник Чернівецького ун-ту:
Збірник наукових праць. – Вип. 193: Біологія. – Чернівці: Рута, 2004. – С. 85–92.
3. Мринський О.П. Рослиннiсть заповiдноi дiлянки Ягорлицького пiвострова /
Охорона, вивчення та збагачення рослинного свiту. – Мiжвiдомчий збiрник. – К: Вища
школа, 1975. – С. 41–44.
4. Ткаченко В.С., Лисенко Г.М., Маяцький Г.Б., Уманець О.Ю. Структурні зміни
фітоценокомплексів Солоноозерної ділянки Чорноморського біосферного заповідника
за даними періодичного картографування // Укр. ботан. журн. – 1997. – 54. № 3. –
С. 232–239.
5. Ткаченко В.С., Маяцький Г.Б. Динамiка рослинностi Потiївськой дiлянки
Чорноморського бiосферного заповiдника пiд впливом iригацiї // Укр. ботан. журн. –
1989. – 46. № 3. – С. 66–71.
6. Уманец О.Ю. Фитоценотическая характеристика сообществ с участием Puccinellia
syvaschica Bilyk на территории Черноморского биосферного заповедника // Заповiдна
справа в Українi. – 2000, Т.6. – Вип.1. – С. 14–16.
7. Уманець О.Ю. БЗ Чорноморський // Фіторізноманіття заповідників і національних
природних парків України. Ч. 1. Біосферні заповідники. Природні заповідники / під
ред. В.А. Онищенка і Т.Л. Андрієнко. – Київ: Фітосоціоцентр, 2012. – С. 73–93.
8. Уманець О.Ю., Войтюк Б.Ю., Соломаха І.В. Синтаксономія рослинності
Чорноморського біосферного заповідника. IV. Ділянка Потіївська // Укр. фітоцен. зб. –
Київ, 2001. – 1 (17). – C. 66–86.
9. Уманець О.Ю., Соломаха I.В. Синтаксономiя рослинностi Чорноморського
бiосферного заповiдника I. Урочище “Ягорлицький Кут” // Укр. фiтоцен. зб. – Київ. –
1998. – Сер. А., вип. 2 (11). – С. 109–127.
10. Уманець О.Ю., Соломаха І.В. Синтаксономія рослинності Чорноморського
біосферного заповідника. II. Острів Тендра // Укр. фітоцен. зб. – Київ. – 1999, Сер.А,
вип.1–2 (12–13). – C. 63–77.
•
Tepe-Oba
V.A. Onyshchenko
Ukrainian name: Тепе-Оба.
Area: 3977.0 ha.
Altitude: 0–287 m.
Latitude: 45°00’09” N (45.0024°).
300
Longitude: 35°19’40” E (35.3276°).
Administrative regions. Autonomous Republic of Crimea: Feodosia city; Territorial waters
of Ukraine.
Ownership: state.
Biogeographic regions: steppic.
Habitats. Level 1. A – 5.9%; B – 0.8%; E – 77.5%; F – 3.8%; G – 8.5%; H – 3.5%.
Habitats. Level 2. A1 Littoral rock and other hard substrata – 0.1%; A2 Littoral sediment
– 0.1%; A3 Infralittoral rock and other hard substrata sediment – 3.5%; A5 Sublittoral
sediment – 2.2%; B2 Coastal shingle – 0.3%; B3 Rock cliffs, ledges and shores, including
the supralittoral – 0.5%; E1 Dry grasslands – 76.5%; E6 Salt steppes – 1.0%; F3 Temperate
and mediterranean-montane scrub – 2.0%; F7 Spiny Mediterranean heaths – 1.8%; G1
Broadleaved deciduous woodland – 2.8%; G3 Coniferous woodland – 5.7%; H2 Screes
– 0.5%; H3 Inland clifs, rock pavements and outcrops – 2.9%; H5 Miscellaneous inland
habitats with very sparse or no vegetation – 0.1%.
Futher habitat description. A2.1 Litoral coarse sediment; A2.5 Coastal saltmarshes and
saline reedbeds; A3.2 Atlanic and Mediterranean moderate energy infralitoral rock; A3.2
Atlanic and Mediterranean low energy infralitoral rock; A5.1 Sublitoral coarse sediment;
A5.5 Sublitoral macrophyte-dominated sediment; B2.1 Shingle beach dritlines; B2.2
Unvegetated mobile shingle beaches above the dritline; B2.3 Upper shingle beaches
with open vegetaion; B3.1 Supralitoral rock (lichen or splash zone); B3.2 Unvegetated
rock clifs, ledges, shores and islets; B3.3 Rock clifs, ledges and shores, with angiosperms;
B3.4 Sot sea-clifs, oten vegetated; E1.1 Inland sand and rock with open vegetation; E1.2
Perennial calcareous grassland and basic steppes; E1.3 Mediterranean xeric grassland; F3.2
Submediterranean deciduous thickets and brushes; F7.4 Hedgehog-heaths; F9.3 Southern
riparian galleries and thickets; G1.7 Thermophilous deciduous woodland; G3.F Highly
301
ariicial coniferous forestry plantations; H2.6 Calcareous and ultra-basic screes of warm
exposures; H3.2 Basic and ultra-basic inland clifs; H5.3 Sparsely- or un-vegetated habitats
on mineral substrates not resulting from recent ice activity; H5.6 Trampled areas.
Land use: agriculture (animals) – major, tourism/recreaion – minor
Protected areas: includes Tepe-Oba regional botanical reserve (1200 ha), overlaps with
Tykha Bukta regional landscape park (about 1499 ha), same as Emerald Site “Tepe-Oba”.
Threats: development (recreaion/tourism) – medium, eutrophicaion – medium.
General description. Tepe-Oba is the easternmost part of the Crimean mountains with
alitudes up to 238 m over the sea level. Major vegetaion type is the steppe. Besides
there are signiicant areas of sparsely vegetated rocks and screes, dry open rock debris
swards, grasslands of submediterranean type with high cover of annuals, woods of Quercus
pubescens, forest plantaions of Pinus. Slope to the sea is steep, with clifs about 100 m high.
Shore consists of rocks, boulders, pebble and gravel. Dominant macrophytes in the sea are
Cystoseira crinita, Cystoseira barbata, Polisiphonia elongata.
Botanical significance. Important area for conservaion of dry grasslands, sea and inland
clifs, marine macrophytes communiies, threatened endemic rock and steppe species.
• Astracantha arnacantha (M.Bieb.) Podlech; A(iv); abundance: unknown; trend:
unknown; species data quality: medium; trend data quality: poor.
• Astragalus reduncus Pall.; A(iv); abundance: unknown; trend: unknown; species data
quality: medium; trend data quality: poor.
• Crambe aspera M. Bieb.; A(iv); abundance: unknown; trend: unknown; species data
quality: medium; trend data quality: poor.
• Crambe koktebelica (Junge) N.Busch; A(ii); abundance: unknown; trend: unknown;
species data quality: medium; trend data quality: poor.
• Crambe steveniana Rupr.; A(iv); abundance: unknown; trend: unknown; species data
quality: medium; trend data quality: poor.
• Lepidium turczaninowii Lipsky; A(i), A(ii), A(iv); abundance: unknown; trend: unknown;
species data quality: medium; trend data quality: poor.
• Onobrychis pallasii (Willd.) M.Bieb.; A(iii); abundance: unknown; trend: unknown;
species data quality: medium; trend data quality: poor.
• Ophrys oestrifera Bieb.; A(ii); abundance: rare; trend: unknown; species data quality:
medium; trend data quality: poor.
• Orchis punctulata Steven ex Lindl.; A(ii); abundance: rare; trend: unknown; species data
quality: medium; trend data quality: poor.
• Stipa syreistschikowii P.Smirn.; A(ii), A(iv); abundance: unknown; trend: unknown;
species data quality: medium; trend data quality: poor.
Criterion C
• A3 Infralittoral rock and other hard substrata; area: 150 ha; trend: decreasing; area
data quality: medium; trend data quality: poor.
• B3.3 Rock cliffs, ledges and shores, with angiosperms; area: 1 ha; trend: stable; area
data quality: poor; trend data quality: medium.
• E1.2 Perennial calcareous grassland and basic steppes; area: 700 ha; trend: stable; area
data quality: medium; trend data quality: medium.
• F7 Spiny Mediterranean heaths; area: 40 ha; trend: stable; area data quality: poor;
trend data quality: medium.
• H3.2 Basic and ultra-basic inland cliffs; area: 40 ha; trend: stable; area data quality: poor;
trend data quality: medium.
302
Literature
1. Костенко Н.С., Дикий Е.А., Алексеева Н.А. Донная растительность Юго–
Восточного Крыма // Сборник научных трудов, посвященных 90–летию Карадагской
научной станции и 25–летию Карадагского природного заповедника НАН Украины. –
Симферополь: СОНАТ, 2004. – С. 66–84.
2. Костенко Н.С., Дикий Е.А., Заклецкий А.А., Марченко В.С. Донная растительность
приоритетных акваторий юго-восточного Крыма: современное состояние и
необходимые меры по сохранению // Заповедники Крыма – 2007. – Симферополь,
2007. – С. 63–68.
3. Шатко В.Г., Миронова Л.П. Конспект флоры хребта Тепе-Оба (Крым) // Бюллетень
Главного ботанического сада. – 2011. – Вып. 197. – С. 43–70.
Ternava – Dnister
V.A. Onyshchenko
Ukrainian name: Тернава – Дністер.
Area: 1601.0 ha.
Altitude: 122–260 m.
Latitude: 48°36’35” N (48.6097°).
Longitude: 26°47’30” E (26.7917°).
Administrative regions. Chernivtsi region: Kelmentsi raion; Kmelnytskyi region: KamianetsPodilskyi raion.
Ownership: state (major), private.
Biogeographic regions: coninental.
Habitats. Level 1. C – 10.9%; E – 24.8%; F – 2.9%; G – 59.5%; H – 1.9%.
Habitats. Level 2. C1 Surface standing waters – 10.9%; E1 Dry grasslands – 24.8%; F3
Temperate and mediterranean-montane scrub – 2.9%; G1 Broadleaved deciduous
woodland – 59.5%; H2 Screes – 0.1%; H3 Inland clifs, rock pavements and outcrops – 1.8%.
Futher habitat descripion. C1.2 Permanent mesotrophic lakes, ponds and pools; E1.1
Inland sand and rock with open vegetation; E1.2 Perennial calcareous grassland and basic
steppes; F3.1 Temperate thickets and scrub; F3.2 Submediterranean deciduous thickets and
brushes; G1.7 Thermophilous deciduous woodland; G1.A Meso- and eutrophic Quercus,
Carpinus, Fraxinus, Acer, Tilia, Ulmus and related woodland; H2.6 Calcareous and ultra-basic
screes of warm exposures; H3.2 Basic and ultra-basic inland cliffs.
Land use: agriculture (animals) – major, forestry – major, nature conservation and research
– major, tourism/recreaion – minor.
Protected areas: overlaps (774 ha) with Podilski Tovtry naional nature park, overlaps
(662 ha) with Khotynskyi naional nature park, includes Kytaihorodske Vidslonennia state
geological nature monument (60 ha), includes Chaplia state botanical reserve (177 ha),
overlaps (774 ha) with Emerald Site “Podilski Tovtry Naional Nature Park”, overlaps (662
ha) with Emerald Site “Khotynskyi Naional Nature Park”.
Threats: agricultural intensification/expansion (grazing) – low, development (recreaion/
tourism) – low.
General description. Steep slopes of the Ternava and Dnister rivers with dry grasslands,
limestone and sandstone rocks, shrub vegetaion and woods. Steppe vegetaion is
dominated by Carex humilis, Festuca valesiaca, Sesleria heuleriana, Sipa capillata,
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Botriochloa ischaemum, Koeleria cristata, Potenilla incana. Typical species are Asperula
cynanchica, Campanula sibirica, Filipendula vulgaris, Galium verum, Medicago falcata,
Salvia vericillata, Scabiosa ochroleuca, Silene eugeniae, Teucrium chamaedrys, Thymus
marschallianus. Shrub vegetaion is represented mainly by Prunus spinosa communiies.
Besides there are Crataegus curvisepala, C. fallacina, C. monogyna, Cerasus mahaleb,
Prunus stepposa, Swida sanguinea (Cornus sanguinea). The tree layer of woods consists of
Quercus robur, Quercus petraea, Sorbus torminalis. In the shrub layer, there occur Cornus
mas, Staphylea pinnata, Viburnum lantana. On the outcrops, there are Allium montanum,
Asplenium ruta-muraria, Aurinia saxailis, Cleistogenes bulgarica, Poa versicolor, Sedum
acre, Sedum sexangulare.
Botanical significance. Important area for conservaion of Podolian steppes, shrubs and
rocks and several endemic species.
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Criterion A
• Aconitum besserianum Andrz.; A(iiі); abundance: rare; trend: unknown; species data
quality: medium; trend data quality: poor.
• Iris hungarica Waldst. & Kit.; A(ii); abundance: frequent; trend: stable; species data
quality: medium; trend data quality: medium.
• Schivereckia podolica Andrz. ex DC. (Draba podolica (Besser) Rupr.); A(ii); abundance:
occasional (1000 individuals); trend: stable; species data quality: good; trend data
quality: medium.
• Spiraea polonica Błocki; A(iii); abundance: rare; trend: stable; species data quality:
medium; trend data quality: medium.
Criterion C
• E1.11 Euro-Siberian rock debris swards; area: 15 ha; trend: stable; area data quality:
medium; trend data quality: good.
• E1.2 Perennial calcareous grassland and basic steppes; area: 370 ha; trend: stable; area
data quality: medium; trend data quality: medium
• F3.247 Ponto-Sarmatic deciduous thickets; area: 40 ha; trend: stable; area data quality:
medium; trend data quality: medium.
• G1.7 Thermophilous deciduous woodland; area: 100 ha; trend: stable; area data quality:
poor; trend data quality: poor.
• H2.6 Calcareous and ultra-basic screes of warm exposures; area: 1 ha; trend: stable;
area data quality: medium; trend data quality: good.
• H3.2 Basic and ultra-basic inland cliffs; area: 25 ha; trend: stable; area data quality:
poor; trend data quality: good.
Literature
1. Волуца О.Д. Spiraea polonica Błocki (Rosaceae) у Чернівецькій області // Актуальні
проблеми ботаніки та екології. Матеріали міжнародної конверенції молодих учених.
Кременець. 11-15 серпня 2009 р. – Тернопіль, 209. – С. 61-62.
2. Волуца О.Д., Токарюк А.І., Чорней І.І., Буджак В.В. Раритетні види флори
національного природного парку “Хотинський”: поширення, ценотична приуроченість
// Прагматичні аспекти діяльності національних природних парків у контексті
збалансованого розвитку : матеріали міжнар.-практ. конф., присвяч. 20-річчю нац.
природ. парку “Вижницький” (17-19 вер. 2015 р., смт Берегомет, Чернівецька обл.,
Україна). – Чернівці: ДрукАрт, 2015. – С. 326–347.
3. Волуца О. Д., Чорней І. І. Родина Зозулинцеві у флорі Північної Бессарабії //
Запов. справа в Україні. – 2009. – Т. 15, вип. 2. – С. 26–31.
4. Заповідні перлини Хмельниччини / під ред. Т.Л.Андрієнко. – Хмельницький:
ПАВФ “Інтрада”, 2006. – 220 с.
5. Червона книга України. Рослинний світ / за ред. Я.П. Дідуха. – К.: Глобалконсалтинг,
2009. – 900 с.
6. Федорончук М. М., Белемець Н. М., Волуца О. Д. Рідкісні види роду Spiraea L.
(Rosaceae) флори України та стан їхньої охорони // Укр. ботан. журн. – 2013. – 70, №
2. – С. 164–167.
7. Чорней І. І., Буджак В. В., Токарюк А. І. Сторінками Червоної книги України
(рослиннийсвіт). Чернівецька область. – Чернівці: ДрукАрт, 2010. – 452 с.
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Tiup-Tarkhan i Kalynivskyi Pivostriv
V.P. Kolomiychuk, V.A. Onyshchenko
Ukrainian name: Тюп-Тархан і Калинівський півострів.
Transliteraion/Translaion variants: Tiup-Tarkhan and Kalynivskyi Peninsula.
Area: 5266.0 ha.
Altitude: 0–4 m.
Latitude: 45°48’07” N (45.8018°).
Longitude: 34°40’40” E (34.6778°).
Administrative regions. Autonomous Republic of Crimea: Dzhankoi raion.
Ownership: state.
Biogeographic regions: steppic.
“Eastern Syvash” (4050 ha).
Threats: natural events: erosion – low.
General description. A part of shores of the Eastern Syvash with the steppe and halophyic
vegetaion. The steppes are dominated by Agropyron pectinatum, Artemisia santonica,
Carex melanostachya, Festuca valesiaca, Kochia prostrata, Poa bulbosa, Sipa capillata,
S. ucrainica. Other frequent species are Atriplex aucheri, Elisanthe viscosa, Limonium
tschurjukiense, Phlomis pungens, Verbascum densilorum. Mesic grasslands are
dominated by Elytrigia elongata and E. repens. Dominants of salt grasslands are Argusia
sibirica, Artemisia santonica, Salsola soda (Soda inermis), Tripolium pannonicum. The
litoral halophyic vegetaion is formed mainly by Halocnemum strobilaceum, Halimione
verrucifera, Salicornia perennans, Puccinellia fominii, Aeluropus litoralis.
Botanical significance. One of the best sites for endemic species Limonium tschurjukiense.
Criterion A
• Limonium tschurjukiense (Klokov) Lavrenko ex Klokov; A(iv); abundance: frequent;
trend: stable; species data quality: medium; trend data quality: medium.
Conservaion proposals. Сreate Syvaskyi national nature park and an Emerald Site including
the enire IPA.
Literature
1. Коломійчук В. П., Онищенко В. А., Перегрим М. М. Важливі ботанічні території
Приазов’я / за ред. Т. Л. Андрієнко. – К. : Альтерпрес, 2012. – 114 с.
2. Котов С.Ф., Вахрушева Л.П. Растительный покров Калиновского регионального
ландшафтного парка (современное состояние и перспективы сохранения
ценогенофонда) // Бюлл. Никитск. ботан. сада, 2003. – Вып. 88. – С. 34–37.
3. Котов С.Ф., Вахрушева Л.П., Бирюлева Э.Г. Научное обоснование о
целесообразности организации в Крыму государственного ботанического заказника
“Калиновский”. – Симферополь: Экоцентр “Синтез НТ”, 1991. – 21 с.
Tovtrivska Sinka
Habitats. Level 1. A – 62.8%; E – 37.2%.
Habitats. Level 2. A2 Littoral sediment – 61.8%; A5 Sublittoral sediment – 1.0%; E1 Dry
grasslands – 11.4%; E6 Salt steppes – 25.8%.
Futher habitat description. A2.3 Littoral mud; A2.4 Littoral mixed sediments; A2.5 Coastal
saltmarshes and saline reedbeds; A5.3 Subittoral mud; E1.2 Perennial calcareous grassland
and basic steppes; D6.1 Inland saltmarshes; E6.2 Continental inland salt steppes.
Land use: agriculture (animals) – major; nature conservaion and research – major.
Protected areas: overlaps (about 3300 ha) with Kalynivskyi regional landscape park,
overlaps (about 3700 ha) with Ramsar Site “Eastern Syvash”, overlaps with Emerald Site
I.I. Chorney, A.I. Tokaryuk, V.V. Budzhak
Ukrainian name: Товтрівська стінка.
Transliteraion/Translaion variants: Tovtry Wall, Tovtry Clif.
Area: 16.1 ha.
Altitude: 196–255 m.
Latitude: 48°33’51” N (48.5642°).
Longitude: 25°52’03” E (25.8675°).
Administrative regions. Chernivtsi region: Zastavna raion.
Ownership: state.
Biogeographic regions: coninental.
Habitats. Level 1: E – 89.0%, F – 1.0%; H – 10.0% .
Habitats. Level 2: E1 Dry grasslands – 89.0%; F3 Temperate and mediterranean-montane
scrub – 1.0%; H3 Inland cliffs, rock pavements and outcrops – 10.0%.
Futher habitat description: E1.1 Inland sand and rock with open vegetaion; E1.2 Perennial
calcareous grassland and basic steppes; F3.1 Temperate thickets and scrub; H3.2 Basic and
ultra-basic inland clifs.
Land use: nature conservation and research – major.
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307
Protected areas: includes Tovtrivska Sinka state botanical reserve .
Threats: –.
Triokhizbenskyi Step
V.A. Onyshchenko
Ukrainian name: Трьохізбенський степ.
Transliteraion/Translaion variants: Triokhizbens’kyi Steppe.
Area: 3281.0 ha.
Altitude: 50–150 m.
Latitude: 48°48’37” N (48.8103°).
Longitude: 38°57’59” E (38.9664°).
Administrative regions. Luhansk region: Slovianoserbsk raion, Novoaidar raion.
Ownership: state.
Biogeographic regions: steppic.
General description. Dry grasslands, gypsum and limestone outcrops. Grasslands are
dominated by Carex humilis, C. montana, Festuca valesiaca, Koeleria cristata, Poa
angustifolia, Sesleria heufleriana, Stipa capillata, S. pennata. Other frequent species
are Achillea submillefolium, Asperula cynanchica, Astragalus austriacus, A. onobrychis,
Anthericum ramosum, Bupleurum falcatum, Galium verum, Gypsophila thyraica, Minuartia
thyraica, Potentilla arenaria, Salvia pratensis, S. verticillata, Securigera varia, Teucrium
chamaedrys, T. pannonicum, Thymus marschallianus, Trifolium montanum.
Botanical significance. Large populaion of endemic species Gypsophila thyraica.
Criterion A
• Gypsophila thyraica A.Krasnova; A(iiі); abundance: frequent; trend: unknown; species
data quality: good; trend data quality: poor.
Literature
1. Чорней І.І., Скільський І.В., Коржик В.П., Буджак В.В. Заповідні об’єкти Буковини
загальнодержавного значення як основа регіональної екологічної мережі // Заповідна
справа в Україні. – 2001. – Т. 7, вип. 2. – С. 73–98.
308
Habitats. Level 1. E – 81.0%; F – 2.0%; G – 16.0%; H – 1.0%.
Habitats. Level 2. E1 Dry grasslands – 76.0%; E3 Seasonly wet and wet grasslands – 5.0%;
F3 Temperate and mediterranean-montane scrub – 2.0%; G1 Broadleaved deciduous
woodland – 8.0%; G3 Coniferous woodland – 8.0%; H5 Miscellaneous inland habitats with
very sparse or no vegetation – 1.0%.
Futher habitat description. E1.9 Open non-Mediterranean dry acid and neutral grassland,
including inland dune grassland (× E1.1); E3.4 Moist or wet eutrophic and mesotrophic
grassland; F3.1 Temperate thickets and scrub; G1.7 Thermophilous deciduous woodland;
G3.4 Pinus sylvestris woodland south of the taiga; H5.3 Sparsely- or un-vegetated habitats
309
on mineral substrates not resulting from recent ice activity; H5.6 Trampled areas.
Land use: nature conservaion and research – 100%.
Protected areas: included in Luhanskyi nature reserve (same as Triokhizbenskyi Step division
of the reserve), included in Emerald Site “Luhanskyi Nature Reserve”.
Threats: abandonment/reducion of land management – low.
General description. A part of sand terrace over the loodplain of the Siverskyi Donets river.
Prevailng vegetaion is open grasslands on sand dominated by Koeleria sabuletorum, Festuca
beckeri, Sipa borysthenica, Carex colchica, Agropyron tanaiicum, Agropyron lavrenkoanum,
Calamagrosis epigeios, Artemisia marschalliana. These habitats are rich in endemic species
(Achillea micrantha, Agropyron tanaiicum, Anchusa popovii, Astragalus varius, Centaurea
majorovii, Jacobaea borysthenica (Senecio borysthenicus), Jurinea thyrsilora, Linaria dulcis,
Linaria genisifolia, Tragopogon tanaiicus, Tragopogon ucrainicus). Besides there are wet
meadows (Juncus ariculatus, Juncus conglomeratus), shrub communiies (Salix acuifolia,
Salix rosmarinifolia), woods (Betula pendula, Populus tremula, Populus nigra, Alnus
gluinosa, Pinus sylvestris).
Botanical significance. Important area for conservaion of sand steppes and endemic
psammophyic species.
Criterion A
• Allium savranicum Besser; A(ii); abundance: occasional; trend: stable; species data
quality: medium; trend data quality: poor.
Criterion C
• E1.9 Open non-Mediterranean dry acid and neutral grassland, including inland dune
grassland; area: 2400 ha; trend: stable; area data quality: medium; trend data quality:
medium.
• X35 Inland Sand Dunes; area: 300 ha; trend: stable; area data quality: medium; trend
data quality: medium.
Literature
1. Русіна Н.В., Ходосовцев О.Є. Ліхенобіота Трьохізбенського полігону як
перспективного відділення Луганського природного заповідника // Наукові праці
Луганського природного заповідника. Рослинний і тваринний світ та його охорона. –
Луганськ. – 2008. – Вип. 1, присвячений 40-річному ювілею Луганського природного
заповідника. – С. 38–43.
2. Русіна Н.В. Історія та сучасний стан дослідження ліхенобіоти Луганського
природного заповідника // Наукові праці Луганського природного заповідника.
Рослинний і тваринний світ та його охорона. – Луганськ. – 2011. – Вип. 2. – С. 47–53
3. Сова Т.В., Боровик Л.П. ПЗ Луганський // Фіторізноманіття заповідників і
національних природних парків України. Ч. 1. Біосферні заповідники. Природні
заповідники / під ред. В.А. Онищенка і Т.Л. Андрієнко. – Київ: Фітосоціоцентр, 2012. –
С. 221–253.
4. Червона книга України. Рослинний світ / за ред. Я.П. Дідуха. – К.: Глобалконсалтинг,
2009. – 900 с.
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Troitska Balka
V.P. Kolomiychuk, V.A. Onyshchenko
Ukrainian name: Троїцька балка.
Transliteraion/Translaion variants: Troits’ka Ravine.
Area: 387.0 ha.
Altitude: 25–81 m.
Latitude: 47°04’07” N (47.0685°).
Longitude: 35°26’35” E (35.4430°).
Administrative regions. Zaporizhia region: Melitopol raion
Ownership: state.
Biogeographic regions: steppic.
Habitats. Level 1. E – 82.3%; F – 0.1%; G – 15.4%; H – 1.8%; J – 0.4%.
Habitats. Level 2. E1 Dry grasslands – 81.3%; E2 Mesic grasslands – 1.0%; F3 Temperate
and mediterranean-montane scrub – 0.1%; G1 Broadleaved deciduous woodland – 15.4%;
H5 Miscellaneous inland habitats with very sparse or no vegetation – 1.8%; J4 Transport
networks and other constructed hard-surfaced areas – 0.4%.
Futher habitat description. E1.2 Perennial calcareous grassland and basic steppes; G1.C
Highly artificial broadleaved deciduous forestry plantations; H5.6 Trampled areas; J4.3 Rail
networks.
Land use: agriculture (animals) – major; forestry – major; nature conservaion and research
– minor; urban/industrial/transport – minor.
Protected areas: includes regional entomological reserve “Tsilynna balka Troitska, dilianka
№1” (17 ha), includes regional entomological reserve “Tsilynna balka Troitska, dilianka
№2” (9 ha), includes regional botanical reserve reserve “Troitska balka” (1 ha), included in
Emerald Site “Troitska Balka”.
311
Threats: forestry (aforestaion) – medium, included in Emerald Site “Troitska Balka”.
General description. A valley with prevailing steppe vegetaion dominated by Sipa
lessingiana, Sipa capillata, Sipa ucrainica, Agropyron pecinatum, Festuca valesiaca.
Botanical significance. Important area for threatened steppe species Calophaca wolgarica,
Tulipa ophiophylla, Cymbochasma borysthenica, Allium pervestitum, Elytrigia stipifolia,
Caragana scythica, Centaurea talievii, Astragalus pallescens, Eremogone rigida, Dianthus
palidilorus.
Criterion A
• Calophaca wolgarica (L.f.) DC.; A(iv); abundance: occasional; trend: stable; species data
quality: medium; trend data quality: poor.
Conservaion proposals. Do not allow ariicial aforestaion. Сreate a state botanical
reserve and an Emerald Site including the enire IPA.
Literature
1. Коломійчук В.П. Ключові ботанічні території Північного Приазов’я // Заповідна
справа в Україні. – 2008. – Т.14, вип. 1. – С. 61–66.
2. Коломійчук В. П., Онищенко В. А., Перегрим М. М. Важливі ботанічні території
Приазов’я / за ред. Т. Л. Андрієнко. – К. : Альтерпрес, 2012. – 114 с.
occupies the largest area. Besides there are termophilous Quercus robur forests, ariicial
plantaions of coniferous and broadleaved trees, outcrops of limestone, mesic meadows
and reed marshes.
Trosianets
V.A. Onyshchenko, O.M. Popova
Ukrainian name: Тростянець.
Area: 665.0 ha.
Altitude: 40–150 m.
Latitude: 47°29’05” N (47.4847°).
Longitude: 29°11’50” E (29.1973°).
Administrative regions. Odesa region: Krasni Okny raion.
Ownership: state.
Biogeographic regions: coninental.
Habitats. Level 1. C – 1.6%; E – 70.9%; F – 2.0%; G – 24.0%; H – 1.5%.
Habitats. Level 2. C3 Littoral zone of inland surface waterbodies – 1.6%; E1 Dry grasslands
– 69.0%; E2 Mesic grasslands – 1.9%; F3 Temperate and mediterranean-montane scrub –
2.0%; G1 Broadleaved deciduous woodland – 12.0%; G3 Coniferous woodland – 12.0%;
H2 Screes – 0.2%; H3 Inland cliffs, rock pavements and outcrops – 1.1%; H5 Miscellaneous
inland habitats with very sparse or no vegetation – 0.2%.
Futher habitat description. E1.1 Inland sand and rock with open vegetation; E1.2
Perennial calcareous grassland and basic steppes; E2.5 Meadows of the steppe zone;
F3.2 Submediterranean deciduous thickets and brushes; G1.7 Thermophilous deciduous
woodland; G1.C Highly artificial broadleaved deciduous forestry plantations; G3.F Highly
artificial coniferous plantations; H5.3 Sparsely- or un-vegetated habitats on mineral
substrates not resulting from recent ice activity; H5.6 Trampled areas.
Land use: agriculture (animals) – 70%; forestry – 24%; mowing/hay making – minor.
Protected areas: same as Emerald Site “Trosianetskyi”.
Threats: abandonment/reducion of land management – low; agricultural intensiicaion/
expansion (grazing) – low; forestry (aforestaion) – low.
General description. Valleys of the Trosianets river and its tributaries. The steppe vegetaion
312
Botanical significance. One of two localiies of Genista tetragona in Ukraine.
Criterion A
• Genista tetragona Besser; A(i), A(ii), A(iv); abundance: occasional; trend: unknown;
species data quality: medium; trend data quality: poor.
Conservaion proposals. Сreate a state botanical reserve and an Emerald Site including the
enire IPA.
313
Tsetsyno
I.I. Chorney, A.I. Tokaryuk, V.V. Budzhak
Ukrainian name: Цецино.
Area: 519 ha.
Altitude: 295–537 m.
Latitude: 48°18’07” N (48.3019°).
Longitude: 25°49’45” E (25.8292°).
Administrative regions. Chernivtsi region: Chernivtsi city, Kitsman raion, Storozhynets raion.
Ownership: state.
Biogeographic regions: coninental.
avellana, Swida sanguinea (Cornus sanguinea), Viburnum opulus, rarely Staphylea pinnata.
Frequent summer species of the herb layer are Galium odoratum, Lamium galeobdolon,
Pulmonaria obscura, Asarum europaeum, Mercurialis perennis, Aegopodium podagraria,
Actaea spicata, Dryopteris ilix-mas, Salvia gluinosa, Symphytum cordatum, Paris quadrifolia,
Stachys sylvaica. Spring ephemeroids are respresented by Allium ursinum, Corydalis
cava, Corydalis solida, Crocus heuffelianus, Ficaria verna, Isopyrum thalictroides. Typical
are calciphilous species Cephalanthera damasonium, C. longifolia, C. rubra, Cypripedium
calceolus, Epipactis helleborine, Epipactis purpurata, Neottia nidus-avis, Platanthera bifolia,
Sorbus torminalis, Taxus baccata.
Botanical significance. Important area for calciphilous beech forests and Cypripedium
calceolus.
Criterion A
• Cypripedium calceolus; A(ii); abundance: occasional; trend: unknown; species data
quality: good; trend data quality: poor.
Criterion C
• G1.66 Medio-European limestone Fagus forests (*9150); area: 300 ha; trend: stable;
area data quality: medium; trend data quality: good.
Literature
1. Чорней І.І., Скільський І.В., Коржик В.П., Буджак В.В. Заповідні об’єкти Буковини
загальнодержавного значення як основа регіональної екологічної мережі // Заповідна
справа в Україні. – 2001. – Т. 7, вип. 2. – С. 73–98.
Tulyntsi – Makedony
Habitats. Level 1: G – 99.2%, H – 0.2%, J – 0.6%.
Habitats. Level 2: G1 Broadleaved deciduous woodland – 99.2%; H5 Miscellaneous inland
habitats with very sparse or no vegetation – 0.2%, J4 Transport networks and other
constructed hard-surfaced areas – 0.6%.
Futher habitat description: G1.6 Fagus woodland; H5.6 Trampled areas, J4.2 Road
networks.
Land use: forestry – minor; nature conservation and research – major.
Protected areas: included in Chernivetskyi regional landscape park, include Tsetsyno state
landscape reserve (430 ha), included in Emerald Site “Chernivetskyi Regional Landscape
Park”.
Threats: forestry (intensiied forest management) – low.
General description. Fagus sylvaica forest with paricipaion of Acer platanoides and Acer
pseudoplatanus. Sparse shrub layer consists Sambucus nigra (predominantly), Corylus
V.A. Onyshchenko
Ukrainian name: Тулинці – Македони.
Area: 529.0 ha.
Altitude: 114–182 m.
Latitude: 49°48’49” N (49.8135°).
Longitude: 31°08’18” E (31.1383°).
Administrative regions. Kyiv region: Myronivka raion.
Ownership: state.
Biogeographic regions: coninental.
Habitats. Level 1. E – 75.1%; F – 0.1%; G – 24.2%; H – 0.6%.
Habitats. Level 2. E1 Dry grasslands – 70.1%; E2 Mesic grasslands – 5.0%; F3 Temperate and
mediterranean-montane scrub – 0.1%; G1 Broadleaved deciduous woodland – 24.2%; H5
Miscellaneous inland habitats with very sparse or no vegetation – 0.6%.
Futher habitat description. E1.2 Perennial calcareous grassland and basic steppes; G1.A
Meso- and eutrophic Quercus, Carpinus, Fraxinus, Acer, Tilia, Ulmus and related woodland.
Land use: agriculture (animals) – major; nature conservaion and research – minor.
Protected areas: includes regional botanical reserve “Tulynetski Perelisky” (88 ha).
Threats: agricultural intensification/expansion (grazing) – medium.
General description. Valleys with the steppe and forest vegetaion. Major dominants of
grasslands are Poa angusifolia, Bromopsis inermis, Botriochloa ischaemum, Calamagrosis
epigeios. Communiies with the dominance of Carex humilis, Festuca rupicola, Festuca
valesiava, Sipa capillata, Sipa pennata occupy smaller areas. Frequent species are Carex
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praecox, Elytrigia intermedia, Elytrigia repens, Medicago falcata, Salvia pratensis. The
grasslands have many species that are character species of meadows steppes: Adonis
vernalis, Anemone sylvestris, Astragalus onobrychis, Astragalus dasyanthus, Bulbocodium
versicolor, Crocus reiculatus, Dianthus membranaceus, Friillaria ruthenica, Inula hirta,
Iris hungarica, Jurinea arachnoidea, Jurinea salicifolia, Jurinea calcarea, Hyacinthella
leucophaea, Pedicularis kaufmannii, Pulsailla pratensis, Salvia nemorosa, Scorzonera
purpurea, Thalictrum minus. In botoms of the valleys there are mesic and wet meadows.
Turova Dacha
V.A. Onyshchenko, N.V. Shumska
Ukrainian name: Турова дача.
Area: 1046.0 ha.
Altitude: 402–420 m.
Latitude: 48°52’02” N (48.8672°).
Longitude: 24°14’36” E (24.2433°).
Administrative regions. Ivano-Frankivsk region: Rozhniaiv raion.
Botanical significance. One of the largest areas of meadow steppes in Ukraine.
Criterion C
• E1.2 Perennial calcareous grassland and basic steppes; area: 350 ha; trend: stable; area
data quality: medium; trend data quality: medium.
Literature
1. Бортняк М.М., Войтюк Ю.О., Любченко В.М., Голяченко Т.В. Флористичні
особливості ділянки ділянки степу Шандра-Тулинці // Укр. ботан. журн. – 1993. – 50, №
2. – С. 122–125.
2. Василюк О., Костюшин В., Норенко А., Плига А., Прекрасна Є., Коломицев Г.,
Фатікова М. Природно-заповідний фонд Київської області. Київ: НЕЦУ, 2012. – 338 с.
3. Кучерява Л. Ф., Шевчик В. Л., Бакаліна М. В., Тищенко О. В. Червонокнижні
види у заказнику “Тулинецькі переліски” (Київська обл.) // Роль природно-заповідних
територій у підтриманні біорізноманіття. Матеріали конф. – Канів, 2003. – С. 116.
4. Кучерява Л. Ф., Шевчик В. Л., Тищенко О. В. Судинні рослини заказника “Тулинецькі
переліски” (Київська обл.) // Заповідна справа в Україні. – 2007. – 13, 1–2. – С. 38–41.
5. Мельник В. І., Гриценко В. В. Росслинний покрив урочища “Тулинецькі переліски”
– еталон лучних степв Київського плато // Збірник наукових праць Полтавського дер.
пед. ун-ту ім. В. Г. Короленка. Серія “Екологія.Біологічні науки”. – Полтава, 2005. – №4
(43) – С. 22–28.
Ownership: state.
Biogeographic regions: alpine.
Habitats. Level 1. C – 0.0%; D – 0.3%; E – 0.1%, G – 98.9%; H – 0.7%.
Habitats. Level 2. C2 Surface running waters – 0.0%; D1 Raised and blanket bogs – 0.2%; D2
Valley mires, poor fens and transition mires – 0.1%; E3 Seasonally wet and wet grasslands
– 0.1%; G1 Broadleaved deciduous woodland – 5.5%; G3 Coniferous woodland – 73.4%; G4
Mixed deciduous and coniferous woodland – 20.0%; H5 Miscellaneous inland habitats with
very sparse or no vegetation – 0.7%.
Futher habitat description. C2.3 Permanent non-tidal, smooth-flowing watercourses; D1.1
Raised bogs; D2.2 Poor fens and sot-water spring mires; D2.3 Transiion mires and quaking
bogs; E3.4 Moist or wet eutrophic and mesotrophic grassland; G1.5 Broadleaved swamp
woodland on acid peat; G1.9 Non-riverine woodland with Betula, Populus tremula or Sorbus
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317
aucuparia; G3.4 Pinus sylvestris woodland south of the taiga; G3.E Nemoral bog conifer
woodland; G4.1 Mixed swamp woodland; G4.4 Mixed Pinus sylvestris – Betula woodland.
Land use: forestry – major, nature conservation and research – 24%.
Protected areas: includes state hydrological reserve “Turova Dacha” (255 ha), included in
Emerald Site “Turova Dacha”.
Threats: burning of vegetaion – low, eutrophicaion – low; water (drainage) – low.
General description. Swamped Pinus sylvestris woods with the layer of Sphagnum and
wet Pinus sylvestris woods. In the layer of herbs and small shrubs of the swamped woods,
there dominate Eriophorum vaginatum, Ledum palustre, Vaccinium oxycoccos (Oxycoccus
palustris), Vaccinium uliginosum. The moss layer is formed by Sphagnum capillifolium,
S. magellanicum, S. recurvum s.l., Polytrichum strictum. In the forests of lower moisture,
typical dominant is Vaccinium myrillus. The area of swamped woods is decreasing because
of drainage works carried out several decades ago.
Botanical significance. The largest Sphagnum pine wood in the alpine biogeographic region
of Ukraine.
Criterion C
• G3.E Nemoral bog conifer woodland; area: 100 ha; trend: decreasing; area data quality:
poor; trend data quality: good.
Conservaion proposals. Increase the groudwater level.
Literature
1. Онищенко В.А., Андрієнко Т.Л. Клас Oxycocco-Sphagnetea Br.-Bl. et Tüxen ex
Westhoff et al. 1946 в Українських Карпатах. – Укр. ботан. журн. – 2015. – 72 (3). –
С. 218–228.
2. Природно-заповідні території та об’єкти Івано-Франківщини. – Івано-Франківськ,
2000. – 272 с.
Permanent non-tidal, smooth-flowing watercourses; C3.2 Water-fringing reedbeds and
tall helophytes other than canes; C3.4 Species-poor beds of low-growing water-fringing
or amphibious vegetation; D1.1 Raised bogs; D2.2 Poor fens and sot-water spring mires;
D2.3 Transiion mires and quaking bogs; D5.1 Reedbeds normally without free-standing
water; D5.2 Beds of large sedges normally without free-standing water; E1.1 Inland sand
and rock with open vegetation; E3.4 Moist or wet eutrophic and mesotrophic grassland;
F9.1 Riverine scrub; F9.2 Salix carr and fen scrub; G1.1 Riparian and gallery woodland,
with dominant Alnus, Betula, Populus or Salix; G1.2 Mixed riparian floodplain and gallery
woodland; G1.4 Broadleaved swamp woodland not on acid peat; G1.5 Broadleaved swamp
woodland on acid peat; G1.8 Acidophilous Quercus-dominated woodland; G1.9 Nonriverine woodland with Betula, Populus tremula or Sorbus aucuparia; G3.4 Pinus sylvestris
woodland south of the taiga (major); G3.E Nemoral bog conifer woodland; G4.1 Mixed
swamp woodland; G4.4 Mixed Pinus sylvestris – Betula woodland.
Ubort – Bolotnytsia
V.A. Onyshchenko
Ukrainian name: Уборть – Болотниця.
Area: 23423 ha.
Altitude: 148–178 m.
Latitude: 51°30’40” N (51.5111°).
Longitude: 27°57’03” E (27.9508°).
Administrative regions. Zhytomyr region: Olevsk raion, Ovruch raion.
Ownership: state
Biogeographic regions: coninental.
Habitats. Level 1. C – 0.2%; D – 11.0%; E – 1.4%; F – 1.0%; G – 85.4%; H – 1.0%.
Habitats. Level 2. C1 Surface standing waters – 0.04%; C2 Surface running waters – 0.03%;
C3 Littoral zone of inland surface waterbodies – 0.13%; D1 Raised and blanket bogs – 1%;
D2 Valley mires, poor fens and transition mires – 9%; D5 Sedge and reedbeds, normally
without free-standing water – 1%; E1 Dry grasslands – 1%; E3 Seasonally wet and wet
grasslands – 0.4%; F9 Riverine and fen scrubs – 1%; G1 Broadleaved deciduous woodland
– 8%; G3 Coniferous woodland – 55.4%; G4 Mixed deciduous and coniferous woodland –
22%; H5 Miscellaneous inland habitats with very sparse or no vegetation – 1%.
Futher habitat description. C1.2 Permanent mesotrophic lakes, ponds and pools; C2.3
318
Land use: forestry – minor, mowing / hay making – minor, nature conservation and research
– 97%.
Protected areas: includes Poliskyi nature resrve (20858 ha), includes Didove Ozero state
hydrological reserve (294 ha), includes Plotnytsia state landscape reserve (460 ha), overlaps
with Emerald Site “Poliskyi” (22966 ha).
Threats: abandonment/reduction of land management – low, burning of vegetaion – low,
water (drainage) – low.
General description. Complex of Pinus sylvestris forests on luvioglacial sands, with
319
oligotrophic, mesotrophic and eutrophic mires. Most constant species of pine forests
are Betula pendula, Dicranum polysetum, Pleurozium schreberi. Oten these species are
dominants. Dry lichen forests difer by presence and dominance of Calluna vulgaris, Cladonia
rangiferina, Cladonia aplestris, Corynephorus canescens, Polytrichum piliferum. Mesic pine
forests have high constancies of Calamagrosis epigeios, Calluna vulgaris, Chamaecyisus
zingeri, Festuca ovina. Constant diferenial species of wet pine forests are Frangula alnus,
Molinia caerulea, Populus tremula, Quercus robur, Sorbus aucuparia, Vaccinium viis-idaea,
Vaccinium myrilus. Swamp pine forests difer from wet ones by presence of Andromeda
polifolia, Eriophorum vaginatum, Polytrichum alpestre, Sphagnum cuspidatum, Sphagnum
fallax, Vaccinium oxycoccos (Oxycoccus palustris), Vaccinium uliginosum. Constant species
of raised bogs are Andromeda polifolia, Betula pubescens, Calluna vulgaris, Eriophorum
vaginatum, Ledum palustre, Vaccinium oxycoccos, Pinus sylvestris, Polytrichum alpestre,
Sphagnum centrale, Sphagnum cuspidatum, Sphagnum fallax, on hummocks – Sphagnum
magellanicum. Usually they have an open tree layer of low Pinus sylvestris and Betula
pubescens. Most frequent dominants of the lower layers are Eriophorum vaginatum,
Sphagnum cuspidatum, Sphagnum fallax. Transiion mires usually are dominated by Carex
lasiocarpa, Sphagnum recurvum s.l., Sphagnum cuspidatum. Oten they have a tree layer of
Pinus sylvestris and Betula pubescens. Other constant species are Eriophorum vaginatum
and Polytrichum alpestre. Eutrophic mires are dominated by Carex elata and Carex rostrata.
Botanical significance. Important for conservation of mires and pine forests.
Criterion C
• C3.4 Species-poor beds of low-growing water-fringing or amphibious vegetation; area:
0.1 ha; trend: stable; area data quality: poor; trend data quality: poor; note: Juncus
bulbosus.
• D2.3 Transition mires and quaking bogs; area: 1000 ha; trend: stable; area data quality:
medium; trend data quality: poor.
• G3.E Nemoral bog conifer woodland; area: 1500 ha; trend: stable; area data quality:
poor; trend data quality: poor.
• X04 Raised bog complexes; area: 200 ha; trend: stable; area data quality: poor; trend
data quality: poor.
Literature
1. Андрієнко Т.Л. ПЗ Полівський // Фіторізноманіття заповідників і національних
природних парків України. Ч.1. Біосферні заповідники. Природні заповідники /
Колектив авторів під ред. В.А. Онищенка і Т.Л. Андрієнко. – Київ, 2012. – С. 301–312.
2. Андриенко Т.Л., Шеляг-Сосонко Ю.Р. Растительный мир Украинского Полесья в
аспекте его охраны. – Киев: Наук. думка, 1983. – 216 с.
3. Андриенко Т.Л., Попович С.Ю., Шеляг-Сосонко Ю.Р. Полесский государственный
заповедник. Растительный мир. – Киев: Наук. думка, 1986. – 208 с.
4. Балашов Л.С., Мошкова Н.О. Синузії деяких водоростей асоціації водяного
горіха (Trapa rossica V.Vassil.) в заплаві р. Уборті // Укр. ботан. журн. – 1973. – 3, № 3. –
С. 360–364.
5. Бумар Г.Й. Chamaedaphne calyculata (L.) Moench. на Житомирському Полiссi //
Укр. ботан. журн. – 1990. – 47, № 4. – С. 73–74.
6. Водно-болотні угіддя України / Під ред. Г.Б. Марушевського, І.С. Жарук. – К.:
Wetlands International Black Sea Programme, 2006. – 312 с.
7. Воробйов Є.О., Балашов Л.С., Соломаха В.А. Синтаксономія рослинності
Поліського природного заповідника // Укр. фітоцен. зб. – 1997. Сер. Б, вип. 1(8). – К.,
1997. – 128 с.
320
8. Маслова В.Р. Лишайники Поліського заповідника // Укр. ботан. журн. – 1977. –
34, № 1. – С. 55–61.
9. Орлов О.О. Рідкісні та зникаючі види судинних рослин Житомирської області. –
Житомир: Волинь, 2005. – 496 с.
10. Парахонська Н.О., Мошкова Н.О. Рослинний покрив болота Волисок у
Поліському заповіднику та деякі його альгосинузії // Укр. ботан. журн. – 1975. – 32,
№ 6. – С. 741–746.
11. Партика Л.Я. До бріофлори Поліського природного заповідника. – Укр. ботан.
журн. – 1974. – 31, № 6. – С. 770–773.
12. Партика Л.Я. Бриофлора // Полесский государственный заповедник.
Растительный мир. – Киев: Наук. думка, 1986. – С. 34–41.
13. Поліському природному заповіднику – 30 років. – Зб. наук. праць. – Вип. 1. –
Житомир, 1999. – 144 с.
14. Попович С.Ю. Флористичні знахідки на території Поліського державного
заповідника // Укр. ботан. журн. – 1983. – 40, № 6. – С. 94–98.
15. Фіторізноманіття Поліського природного заповідника: водорості, мохоподібні,
судинні рослини / за заг. ред. к.б.н. О.О. Орлова. – К.: НВП Інтерсервіс, 2013. – 256 с.
16. Фіторізноманіття Українського Полісся та його охорона / Під заг. ред.
Т.Л. Андрієнко. – Київ: Фітосоціоцентр, 2006. – 316 с.
Uholka – Shyrokyi Luh
V.A. Onyshchenko
Ukrainian name: Уголька – Широкий Луг.
Area: 14485.0 ha.
Altitude: 370–1501 m.
Latitude: 48°17’52” N (48.2979°).
Longitude: 23°41’14” E (23.6873°).
Administrative regions. Zakarpatska region: Khust raion, Tiachiv raion.
Ownership: state (major), private.
Biogeographic regions: alpine.
Habitats. Level 1. C – 0.2%; E – 4.0%; G – 95.7%; H – 0.1%.
Habitats. Level 2. C2 Surface running waters – 0.2%; E1 Dry grasslands – 1.9%; E2 Mesic
grasslands – 0.3%; E4 Alpine and subalpine grasslands stands – 1.8%; G1 Broadleaved
deciduous woodland – 95.6%; G3 Coniferous woodland – 0.1%; H5 Miscellaneous inland
habitats with very sparse or no vegetation – 0.1%.
Futher habitat description. C2.1 Springs, spring brooks and geysers; C2.2 Permanent nontidal, fast, turbulent watercourses; E1.7 Non-Mediterranean dry acid and neutral closed
grassland (E1.71 Nardus stricta swards, E1.72 Agrosis – Festuca grassland); E2.2 Low and
medium alitude hay meadows; G1.6 Fagus woodland; G3.F Highly artificial coniferous
plantations; H2.3 Temperate-montane acid siliceous screes; H2.4 Temperate-montane
calcareous and ultra-basic screes; H3.1 Acid silicious inland clifs; H3.2 Basic and ultra-basic
inland clifs; H5.6 Trampled areas.
Land use: agriculture (animals) – minor, forestry – minor, mowing/hay making – minor,
nature conservation and research – major.
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Protected areas: included in Karpatskyi biosphere reserve (naional category), included in
Carpathian biosphere reserve (UNESCO), included in World Heritage Site “Primeval Beech
Forests of the Carpathians and the Ancient Beech Forests of Germany” (14485 ha; core zone:
11860 ha, bufer zone: 2625 ha), included in Emerald Site “Carpathian Biosphere Reserve”.
Threats: –.
•
Neckera pennata Hedw.; A(ii); abundance: rare; trend: unknown; species data quality:
poor; trend data quality: poor.
Criterion B
• G1 deciduous broadleaved woodland; area 13860 ha; % of indicator species: 20.8%; No
of indicator species: 16; trend: increasing; species data quality: good; area data quality:
good; trend data quality: good.
Criterion C
• G1.6 Fagus woodland (excluding G1.66); area: 13800 ha; trend: stable; area data quality:
good; trend data quality: good.
• G1.66 Medio-European limestone Fagus forests (*9150); area: 10 ha; trend: stable; area
data quality: medium; trend data quality: good.
Literature
1. Brändli U.-B., Dovhanych Y. Virgin forest of Uholka: Nature Guide to the Largest Virgin
Beech Forest of Europe. A UNESCO World Heritage Site. WSL, Rakhiv: Birmensdorf and CBR,
2008 – 23 p.
2. Гамор Ф.Д., Волощук М.І., Антосяк Т.М., Козурак А.В. БЗ Карпатський //
Фіторізноманіття заповідників і національних природних парків України. Ч. 1. Біосферні
заповідники. Природні заповідники / під ред. В.А. Онищенка і Т.Л. Андрієнко. – Київ:
Фітосоціоцентр, 2012. – С. 45–72.
3. Гамор Ф.Д., Довганич Я.О., Покиньчереда В.Ф. та ін. Праліси Закарпаття.
Інвентаризація та менеджмент. – Рахів, 2008. – 86 с.
4. Біорізноманіття Карпатського біосферного заповідника / Кол. авт., Ред. рада: Я.І.
Мовчан, Ф.Д. Гамор та ін. – К.: Інтерекоцентр, 1997. – 711 с.
5. Онищенко В.А. Лісова рослинність верхньої частини басейну Малої Угольки
(Карпатський біосферний заповідник) // Науковий вісник Чернівецького університету.
– 2007. – Вип. 343: Біологія. – С. 130–147.
6. Праліси в центрі Європи: Путівник по лісах Карпатського біосферного заповідника
/ Кол. авт., ред.: У-Б. Брендлі, Я. Довганич. – Бірменсдорф: Швейцарський федеральний
ін-т досліджень лісу, снігу і ландшафтів (WSL) – Рахів: Карпатський біосферний
заповідник (КБЗ), 2003. – 192 с.
7. Флора і рослинність Карпатського заповідника / Стойко С.М., Тасенкевич Л.О.,
Мілкіна Л.І. та ін. – К.: Наук. думка, 1982. – 220 с.
Usia – Shutnivtsi
General description. The beech forest occupies about 95% of the area. Predominantly it is
the neutrophilous forest. The tree layer usually consists of Fagus sylvaica. Constant species
are Acer pseudoplatanus, Athyrium ilix-femona, Dentaria bulbifera (Cardamine bulbifera),
Dentaria glandulosa (Cardamine glandulosa), Dryopteris ilix-mas, Galium odoratum, Rubus
hirtus. Acidophilous beech forests occupies a smaller area. Besides there are calcareous
beech forests. They difer by presence of Tilia platyphyllos, Taxus baccata, Carex digitata,
Hepaica nobilis, Meliis carpaicum, Moehringia muscosa, Primula acaulis, Seseli libanois.
Botanical significance. Probably this is the largest primeval beech forest in Europe.
Criterion A
• Mannia triandra (Scop.) Grolle; A(ii); abundance: rare; trend: unknown; species data
quality: poor; trend data quality: poor.
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L.H. Lubinska, V.A. Kolodiy
Ukrainian name: Устя – Шутнівці.
Area: 46.5 ha.
Altitude: 122–210 m.
Latitude: 48°35’07” N (48.5853°).
Longitude: 26°37’32” E (26.6255°).
Administrative regions. Kmelnytskyi region: Kamianets-Podilskyi raion.
Ownership: state.
Biogeographic regions: coninental.
Habitats. Level 1. E – 9.6%; F – 7.4%; G – 70.0%; H – 13.0%.
Habitats. Level 2. E1 Dry grasslands – 9.6%; F3 Temperate and mediterranean-montane
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scrub – 7.4%; G1 Broadleaved deciduous woodland – 70.0%; H2 Screes – 1.0%; H3 Inland
clifs, rock pavements and outcrops – 12.0%.
Futher habitat descripion. E1.1 Inland sand and rock with open vegetation; E1.2
Vasylivka
V.P. Kolomiychuk, L.V. Markivska
Ukrainian name: Василівка.
Area: 132.5 ha.
Altitude: 153–235 m.
Latitude: 48°14’37” N (48.2436°).
Longitude: 29°18’41” E (29.3114°).
Administrative regions. Vinnytsia region: Chechelnyk raion.
Ownership: state.
Biogeographic regions: coninental
Perennial calcareous grassland and basic steppes; F3.1 Temperate thickets and scrub;
G1.7 Thermophilous deciduous woodland; G1.A Meso- and eutrophic Quercus, Carpinus,
Fraxinus, Acer, Tilia, Ulmus and related woodland; H2.6 Calcareous and ultra-basic screes of
warm exposures; H3.2 Basic and ultra-basic inland cliffs.
Land use: agriculture (animals) – minor, forestry – minor, nature conservation and research
– major, tourism/recreaion – minor.
Protected areas: included in Podilski Tovtry naional nature park (zone of regulated
recreaion), included in Emerald Site “Podilski Tovtry Naional Nature Park”.
Threats: agricultural intensification/expansion (grazing) – low, development (recreaion/
tourism) – low.
General description. Steep slopes of the Smotrych river valley with calcareous rocks,
shrub vegetaion dominated by Prunus spinosa, and deciduous forest. Typical species of
the outcrops are Aurinia saxatilis, Sedum acre, Allium podolicum, Potenilla arenaria, Sipa
capillata, Veronica incana, Festuca valesiaca, Sesleria heuleriana, Teucrium chamaedrys,
T. pannonicum, Thymus moldavicus, Scabiosa ochroleuca, Schivereckia podolica (Draba
podolica), Anthericum ramosum. The area belongs to the zone of regulated recreaion of
naional nature park “Podiliski Tovtry”.
Botanical significance. One of the best populaions of Schivereckia podolica in Ukraine.
Criterion A
• Schivereckia podolica (Besser) Andrz. et Besser ex DC. (Draba podolica (Besser) Rupr.);
A(ii); abundance: frequent; trend: unknown; species data quality: good; trend data
quality: poor.
Habitats. Level 1. E – 95.0%; F – 1.0%; G – 3.0%; H – 1.0%
Habitats. Level 2. E1 Dry grasslands – 94.7%; E7 Sparsely wooded grasslands – 0.3%; G1
Broadleaved deciduous woodland – 1.0%; H5 Miscellaneous inland habitats with very
sparse or no vegetation – 1.0%.
Futher habitat descripion. E1.2 Perennial calcareous grassland and basic steppes.
Land use: nature conservaion and research – minor.
Protected areas: includes regional botanical reserve “Romashkove” 8.7 ha (a part of
Karmeliukove Podillia naional nature park), overlaps with Emerald Site “Karmeliukove
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325
Podillia Naional Nature Park” 8.7 ha).
Threats: abandonment/reducion of land management – low, agricultural intensiicaion/
expansion (grazing) – low, burning of vegetaion – low.
General description. Slopes with the steppe vegetaion dominated by Festuca valesiaca,
Elytrigia intermedia, Poa angusifolia, Sipa capillata, with presence of Botryochloa
ischaemum, Bromopsis inermis, Chamaecyisus austriacus, Convolvolus arvensis, Elytrigia
repens, Euphorbia cyparissias, Galium verum, Inula encifolia, Koeleria cristata, Salvia
pratensis, Salvia vericillata, Securigera varia, Stachus recta, Teucrium chamaedrys.
Botanical significance. Important area for Carlina onopordifolia.
Criterion A
• Carlina onopordifolia Besser ex Szafer, Kulcz. et Pawł.; A(i), A(ii), A(iv); abundance:
occasional (1400 individuals); trend: stable; species data quality: good; trend data
quality: poor.
Conservaional proposals. Include enire IPA in naional nature park “Karmeliukove Podillia”.
Literature
1. Kuzemko A.A., Becker T., Didukh Y.P., Ardelean I.A., Becker U., Beldean M., Dolnik
C., Jeschke M., Naqinezhad A., Ugurlu E., Ünal A., Vassilev K., Vorona E.I., Yavorska O.H.,
Dengler J. Dry grassland vegetation of Central Podolia (Ukraine) – a preliminary overview on
syntaxonomy, ecology and biodiversity // Tuexenia. – 2014. – Vol. 34. – S. 391–430.
2. Дідух Я.П., Кагало О.О., Орлов О.О. Відкасник татарниколистий Carlina
onopordifolia Besser ex Szafer, Kulcz. et Pawł. / Червона книга України. Рослинний світ /
за ред. Я.П. Дідуха. – К.: Глобалконсалтинг, 2009. – C. 279.
3. Мельник В.І., Скоропляс І. О., Ваколюк В. Д. Carlina onopordifolia (Asteraceae) на
Східному Поділлі // Укр. ботан. журн. – 2014. – 71, №3. – С. 324–329.
4. Польовий Є.В., Дідух Я. П. Еколого-територіальна диференціація рослинного
покриву модельного полігону “Ромашково” в долині р. Савранки (Вінницька обл.) //
Укр. ботан. журн. – 2014. – 71, №6. – С. 647–659.
5. Яворська О.Г. НПП Кармелюкове Поділля // Фіторізноманіття заповідників і
національних природних парків України. Ч. 2. Національні природні парки / Колектив
авторів під ред. В.А. Онищенка і Т.Л. Андрієнко. – Київ, 2012. – С. 265–272.
sparse or no vegetation – 0.8%.
Futher habitat description. C2.5 Temporary running waters; E1.2 Perennial calcareous
grassland and basic steppes; G1.7 Thermophilous deciduous woodland; G1.A Meso- and
eutrophic Quercus, Carpinus, Fraxinus, Acer, Tilia, Ulmus and related woodland.
I.I. Chorney, A.I. Tokaryuk, V.V. Budzhak
Ukrainian name: Василівський і Розкопинський яри.
Transliteraion/Translaion variants: Vasylivs’kyi and Rozkopyns’kyi Ravines.
Area: 1962 ha.
Altitude: 84–263 m.
Latitude: 48°30’08” N (48.5021°).
Longitude: 27°25’36” E (27.4263°).
Administrative regions. Chernivtsi region: Sokyriany raion.
Ownership: state.
Biogeographic regions: coninental.
Habitats. Level 1. E – 0.2%; G – 99.0%; H – 0.8%.
Habitats. Level 2. C2 Surface running waters – 0.0%; E1 Dry grasslands – 0.2%; G1 –
Broadleaved deciduous woodland – 99.0%; H5 Miscellaneous inland habitats with very
Land use: forestry – major, nature conservation and research – minor.
Protected areas: includes (about 497 ha) with regional landscape reserve “Vasilivskyi Yar”.
Threats: forestry (intensiied forest management) – medium.
General description. Broadleaved forests dominated by Quercus robur, Carpinus betulus,
Acer pseudoplatanus, Fraxinus excelsior with admixture of Acer campestre, Acer platanoides,
Tilia cordata, Sorbus torminalis. The shrub layer is composed of Acer tataricum, Acer
campestre, Corylus avellana, Euonymus verrucosus, Euonymus europaeus, Sambucus nigra,
Staphylea pinnata, Swida sanguinea (Cornus sanguinea), Viburnum lantana. Typical species
of the herb layer are Carex pilosa, Stellaria holostea, Asarum europaeum, Aegopodium
podagraria, Convallaria majalis, Galium odoratum, Lathyrus vernus, Hedera helix, Hepatica
nobilis. There scatered limestone boulders up to 3 m in diameter, covered by mosses and
Phylliis scolopendrium.
Botanical significance. Important area for Friillaria montana and Aconitum besserianum.
Criterion А
• Fritillaria montana Hoppe; А (ii); abundance: occasional (5000 individuals); trend:
stable; species data quality: good; trend data quality: poor.
• Aconitum besserianum Andrz. ex Trautv; А (iiі); abundance: rare; trend: stable; species
data quality: medium; trend data quality: poor.
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Vasylivskyi i Rozkopynskyi Yary
Literature
1. Волуца О.Д. Fritillaria montana Hoppe у флорі Чернівецької області // Науковий
вісник Чернівецького університету: Збірник наукових праць. – Вип. 293: Біологія. –
Чернівці: Рута, 2006. – С. 31–34.
2. Волуца О. Euonymus nana M. Bieb. (Celastraceae) – новий вид для флори ПрутДністровського межиріччя (в межах України) // Вісник Прикарпатського національного
університету імені Василя Стефаника. Серія Біологія / Матеріали Міжнародної наукової
конференції “Проблеми вивчення та охорони біорізноманіття Карпат та прилеглих
територій” (Івано-Франківськ, 8-9 листопада 2007р.). – Вип. VII-VIII. – Івано-Франківськ:
Гостинець, 2007. – С. 51-52.
3. Волуца О.Д. Нові місцезнаходження видів роду Pulsailla Mill. (Ranunculaceae)
на території Північно-Бессарабського геоботанічного округу (Чернівецька область) //
“Молодь у вирішенні регіональних та транскордонних проблем екологічної безпеки”:
Матеріали Шостої Міжнародної наукової конференції (м. Чернівці, 11–12 травня 2007
року). – Чернівці: Зелена Буковина, 2007. – С. 59–61.
4. Волуца О.Д., Токарюк А.І. Стан популяції Friillaria montana Hoppe (Liliaceae) в
околицях м. Сокиряни (Чернівецька область) // “Молодь та поступ біології”: Збірник
тез IV Міжнародної конференції студентів і аспірантів (7-10 квітня 2008 року, м. Львів).
– Львів, 2008. – С. 119-120.
5. Каземірська М.А., Чорней І.І. Fritillaria montana Hoppe (Liliaceae): географічна
характеристика, поширення в Україні // Науковий вісник Чернівецького університету.
Біологія (Біологічні системи). – 2010. – Т. 2, Вип. 3. – С. 63–68.
6. Каземірська М.А., Чорней І.І. Вікова та просторова структури популяцій Fritillaria
montana Hoppe у Прут-Дністровському межиріччі // Біологічні системи. – 2010. – Т. 2.,
Вип. 2. – С. 62–66.
7. Червона книга України. Рослинний світ / за ред. Я.П. Дідуха. – К.: Глобалконсалтинг,
2009. – 900 с.
8. Чорней І.І., Буджак В.В., Токарюк А.І. Сторінками Червоної книги України
(рослинний світ). Чернівецька область. – Чернівці: ДрукАрт, 2010. – C. 85–87.
trees, small anthropogenic woodlands, recently felled woodland, early-stage woodland
and coppice – 1.5%; H3 Inland cliffs, rock pavements and outcrops – 0.1%.
Futher habitat descripion. E1.2 Perennial calcareous grassland and basic steppes; E7.2
Sub-coninental parkland; F3.1 Temperate thickets and scrub; G1.A Meso- and eutrophic
Quercus, Carpinus, Fraxinus, Acer, Tilia, Ulmus and related woodland; G3.F Highly artificial
coniferous plantations; G5.6 Early-stage natural and semi-natural woodlands and regrowth;
H3.2 Basic and ultra-basic inland cliffs; H3.5 Almost bare rock pavements, including
limestone pavements.
Land use: agriculture (animals) – minor, nature conservation and research – minor, forestry
Velyki Holdy
N.V. Shumska, V.A. Onyshchenko
Ukrainian name: Великі Голди.
Area: 141 ha.
Altitude: 260–347 m.
Latitude: 49°19’32” N (49.3255°).
Longitude: 24°39’54” E (24.6650°).
Administrative regions. Ivano-Frankivsk region: Rohatyn raion.
Ownership: state.
Biogeographic regions: coninental
Habitats. Level 1. E – 21.9%; F – 2.0%; G – 76.0%; H – 0.1%
Habitats. Level 2. E1 Dry grasslands – 21.5%; E5 Woodland fringes and clearings and tall forb
stands – 0.1%; E7 Sparsely wooded grasslands – 0.3%; F3 Temperate and mediterraneanmontane scrub – 2.0%; G1 Broadleaved deciduous woodland – 55.0%; G3 Coniferous
woodland – 19.0%; G4 Mixed deciduous and coniferous woodland – 0.5%; G5 Lines of
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– major, mowing/hay making – minor.
Protected areas: includes state complex nature monument “Velyki Holdy” (24.0 ha),
included in Emerald Site “Velyki Holdy”.
Threats: abandonment/reducion of land management – medium, agricultural
intensiicaion/expansion (grazing) – low, burning of vegetaion – low.
General description. Hills with the deciduous forest, pine plantaions, dry grasslands, shrubs,
and gypsum outcrops. The grasslands are dominated by Brachypodium pinnatum, Briza
media, Elytrigia intermedia. Other typical species are Anthericum ramosum, Bupleurum
falcatum, Carex montana, Dactylis glomerata, Salvia pratensis. In upper parts of the slopes
and on eroded stony slopes, dominant species are Sipa capillata, Sipa pennata, Festuca
valesiaca, Carex humilis, Koeleria cristata, Teucrium chamaedrys, Astragalus onobrychis,
other frequent species are Artemisia austriaca, Euphorbia cyparissias, Inula ensifolia,
Phleum phleoides, Thymus marschallianus. On gypsum outcrops, there prevail Allium
lusitanicum s.l. (Allium montanum auct.), Festuca pallens, Gypsophila fasigiata, Potenilla
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arenaria, Sedum acre. Main dominant of the shrub vegetaion is Prunus spinosa. The area of
trees and shrubs is increasing, this is the main threat for conservaion of Carlina cirsioides
and other species of grasslands.
Botanical significance. Important area for Carlina onopordifolia.
Criterion A
• Carlina onopordifolia Besser ex Szafer, Kulcz. et Pawł.; A(i), A(ii), A(iv); abundance:
frequent (2000 individuals); trend: stable; species data quality: good; trend data quality:
medium.
Conservaional proposals: do not allow natural and ariicial aforestaion, expand area of
nature monument “Velyki Holdy”.
Literature
1. Шумська Н.В., Чуй О.В., Дмитраш І.І. Збереження фіторізноманіття комплексної
пам’ятки природи “Великі Голди” (Рогатинське Опілля) шляхом оптимізації її території
// Науковий вісник національного університету біоресурсів і природокористування
України. Сер.: Лісівництво та декоративно садівництво. – 2012. – 171 (1). – С. 287–290.
Verbetska Tovtra
Ukrainian name: Вербецька товтра.
Area: 10.8 ha.
Altitude: 280–303 m.
Latitude: 48°48’33” N (48.8092°).
Longitude: 26°36’14” E (26.6038°).
Administrative regions. Kmelnytskyi region: Kamianets-Podilskyi raion.
Ownership: state
Biogeographic regions: coninental.
V.A. Onyshchenko
scrub – 1.1%; H2 Screes – 0.1%; H3 Inland clifs, rock pavements and outcrops – 1.2%.
Futher habitat descripion. E1.1 Inland sand and rock with open vegetation; E1.2 Perennial
calcareous grassland and basic steppes; F3.1 Temperate thickets and scrub; H2.6 Calcareous
and ultra-basic screes of warm exposures; H3.2 Basic and ultra-basic inland cliffs.
Land use: nature conservation and research – major.
Protected areas: overlaps (about 8 ha) with Tovtra Verbetska state botanical reserve,
included in Podilski Tovtry naional nature park, included in Emerald Site “Podilski Tovtry
Naional Nature Park”.
Threats: agricultural intensification/expansion (arable) – low, development (recreaion/
tourism) – low.
General description. A small hill with the steppe vegetaion and limestone outcrops.
Steppe vegetaion is dominated by Carex humilis, Festuca valesiaca, Sipa capillata, Sesleria
heuleriana. On the outcrops, there are Allium lavescens, Allium lusitanicum, Allium
podolicum, Sedum acre, Sempervivum ruthenicum, Veronica incana.
Botanical significance. Important area for Gypsophila thyraica.
Criterion A
• Gypsophila thyraica Krasnova; A(iii); abundance: frequent; trend: unknown; species
data quality: poor; trend data quality: poor.
Literature
1. Заповідні перлини Хмельниччини / під ред. Т.Л.Андрієнко. – Хмельницький:
ПАВФ “Інтрада”, 2006. – 220 с.
Verkhniobuzki Lisy
Habitats. Level 1. E – 97.6%; F – 1.1%; H – 1.3%.
Habitats. Level 2. E1 Dry grasslands – 97.6%; F3 Temperate and mediterranean-montane
V.A. Onyshchenko
Ukrainian name: Верхньобузькі ліси.
Transliteraion/Translaion variants: Upper Buh Forests.
Area: 6543.5 ha.
Altitude: 270–437 m.
Latitude: 49°52’56” N (49.8823°).
Longitude: 25°01’30” E (25.0249°).
Administrative regions. Lviv region: Brody raion; Zolochiv raion.
Ownership: state.
Biogeographic regions: coninental.
Habitats. Level 1. E – 2.1%; G – 96.9%; H – 1.0%.
Habitats. Level 2. E2 Mesic grasslands – 1.6%; E3 Seasonally wet and wet grasslands –
0.5%; G1 Broadleaved deciduous woodland – 94.5%; G3 Coniferous woodland – 0.4%; G4
Mixed deciduous and coniferous woodland – 2.0%; H5 Miscellaneous inland habitats with
very sparse or no vegetation – 1.0%.
Futher habitat descripion. E2.2 Low and medium alitude hay meadows; E3.4 Moist or
wet eutrophic and mesotrophic grassland; G1.2 Mixed riparian floodplain and gallery
woodland; G1.6 Fagus woodland; G1.A Meso- and eutrophic Quercus, Carpinus, Fraxinus,
Acer, Tilia, Ulmus and related woodland; G1.B Non-riverine Alnus woodland; G3.4 Pinus
sylvestris woodland south of the taiga; G3.F Highly artificial coniferous forestry plantations;
G4.7 Mixed Pinus sylvestris – acidophilous Quercus woodland; H5.6 Trampled areas.
Land use: forestry – major, nature conservation and research – 83%.
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331
Protected areas: overlaps with Pivnichne Podillia naional nature park (5920 ha), includes
Pamiatka Peniatska regional complex nature monument (35 ha), overlaps (65 ha) with
Pidlyska Hora regional complex nature monument, includes Sasivska regional botanical
nature monument (130 ha), includes Sviata Hora regional complex nature monument (186
ha), includes regional complex nature monument “Zhulytska Hora, Hora Storozhykha, Hora
Vysoka” (261 ha), overlaps with Emerald Site “Pivnichne Podillia” (about 5920 ha).
Ranunculus lanuginosus. Besides there are mixed Quercus robur – Pinus sylvestris forests
and Alnus gluinosa forests.
Botanical significance. Important area for conservaion of species-rich broadleaved forests.
Criterion B
• G1 Broadleaved deciduous woodland; area: 5800 ha; % of indicator species: 13.0%; No
of indicator species: 10; trend: increasing; species data quality: good; area data quality:
good; trend data quality: medium.
Criterion C
• G1.66 Medio-European limestone Fagus forests (*9150); area: 500 ha; trend: stable;
area data quality: poor; trend data quality: good.
• G1.A4 (*9180) Ravine and slope woodland; area: 200 ha; trend: stable; area data quality:
poor; trend data quality: poor.
Literature
1. Кагало О.О. Північне Поділля // Фіторізноманіття заповідників і національних
природних парків України. Ч. 2. Національні природні парки / під ред. В.А. Онищенка і
Т.Л. Андрієнко. – Київ: Фітосоціоцентр, 2012. – С. 381–394.
2. Рало В. М. Матеріали до класифікації букових лісів Верхобузького пасма
(північно-західне Поділля) // Наукові основи збереження біотичної різноманітності /
Тематичний збірник Інституту екології Карпат НАН України. – Вип. 8–9. – Львів, 2009. –
С. 65–102.
3. Рало В.М., Онищенко В.А. Anthrisco nitidi-Aceretum pseudoplatani – нова асоціація
із Західного Поділля // Науковий вісник Волинського національного університету імені
Лесі Українки. Біологічні науки. – 2008. – № 3. – С. 239–249.
Vitrohon
V.A. Onyshchenko
Ukrainian name: Вітрогон.
Area: 688.0 ha.
Altitude: 70–140 m.
Threats: forestry (intensiied forest management) – low.
General description. Hills covered by broadleaved forests. The largest area is occupied by
Fagus sylvaica forests. Main dominants of the herb layer are Aegopodium podagraria,
Anemone nemorosa, Asarum europaeum, Carex pilosa, Galium odoratum, Hedera helix,
Lamium galeobdolon. Constatnt species are Acer platanoides, Acer pseudoplatanus,
Anemone nemorosa, Carex digitata, Dryopteris ilix-mas, Fagus sylvaica, Galium odoratum,
Lamium galeobdolon, Maianthemum bifolium, Polygonatum mulilorum. Calcareous beech
forests difer by constant presence of Cephalanthera damasonium, Campanula persicifolia,
Campanula trachelium, Convallaria majalis, Daphne mezereum, Lilium martagon, Meliis
sarmaica (M. melissophyllum s.l.), Neoia nidus-avis, Swida sanguinea (Cornus sanguinea),
Viola mirabilis. Species comoposiion of oak-hornbeam forests (Quercus robur, Carpinus
betulus) is similar to one of beech forests. On slopes and botoms of valleys and gullies,
there are species-rich forests with Fraxinus excelsior and Acer pseudoplatanus prevailing
in the tree layer. Their characterisic species are Anthrisus niida, Campanula laifolia,
Cerasium sylvaicum, Chaerophyllum aromaicum, Dipsacus pilosus, Primula elaior,
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Latitude: 49°07’59” N (49.1331°).
Longitude: 39°42’25” E (39.7068°).
333
Administrative regions. Luhansk region: Bilovodsk raion.
Ownership: state.
Biogeographic regions: steppic.
Habitats. Level 1. C – 0.2%; E – 96.6%; F – 2.0%; G – 0.6%; H – 0.5%; J – 0.1%.
Habitats. Level 2. C1 Surface standing waters – 0.1%; C2 Surface running waters – 0.1%;
E1 Dry grasslands – 93.0%; E2 Mesic grasslands – 3.6%; F3 Temperate and mediterraneanmontane scrub – 2.0%; G1 Broadleaved deciduous woodland – 0.6%; H5 Miscellaneous
inland habitats with very sparse or no vegetation – 0.5%; J4 Transport networks and other
constructed hard-surfaced areas – 0.1%.
Futher habitat description. E1.1 Inland sand and rock with open vegetation; E1.2 Perennial
calcareous grassland and basic steppes; F3.2 Submediterranean deciduous thickets and
brushes; H5.6 Trampled areas.
Land use: agriculture (animals) – minor.
Protected areas: overlaps (350–400 ha) with proposed regional botanical reserve
“Vitrohon”, overlaps (647 ha) with Emerald Site “Vitrohonskyi”.
Threats: agricultural intensification/expansion (arable) – low, agricultural intensiicaion/
expansion (grazing) – low, forestry (aforestaion) – low.
General description. Small valleys with the steppe vegetaion (abandoned pastures) on
the slopes. Major dominants are Festuca valesiaca s.l., Sipa capillata, Sipa irsa, Sipa
ucrainica, Sipa zalesskii.
Botanical significance. Important area for conservaion of the steppe vegetaion and some
steppe species.
Criterion A
• Paeonia tenuifolia L.; A(ii); abundance: frequent; trend: stable; species data quality:
medium; trend data quality: medium.
• Stipa zalesskii Wilensky; A(ii); abundance: frequent; trend: stable; species data quality:
medium; trend data quality: medium.
Criterion C
• E1.2 Perennial calcareous grassland and basic steppes; area: 600 ha; trend: stable; area
data quality: good; trend data quality: medium.
Conservaion proposals. Сreate an Emerald Site including the enire IPA.
Volyzhyn Lis
V.A. Onyshchenko, O.Yu. Umanets
Ukrainian name: Волижин ліс.
Transliteraion/Translaion variants: Volyzhyn Wood.
Area: 203.0 ha.
Altitude: 0–6 m.
Latitude: 46°32’21” N (46.5391°).
Longitude: 31°43’14” E (31.7205°).
Administrative regions. Mykolaiv region: Ochakiv raion.
Ownership: state.
Biogeographic regions: steppic.
Habitats. Level 1. C – 10.3%; D – 2.6%; E – 74.4%; F – 0.4%; G – 11.8%; H – 0.5%.
Habitats. Level 2. Surface standing waters – 8.4%; C3 Littoral zone of inland surface
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waterbodies – 1.9%; D5 Sedge and reedbeds normally without free-standing water – 2.6%;
E1 Dry grasslands – 52.8%; E2 Mesic grasslands – 21.5%; E6 Inland salt steppes – 0.1%;
F3 Temperate and mediterranean-mountane scrub – 0.4%; G1 Broadleaved deciduous
woodland – 11.8%; H5 Miscellaneous inland habitats with very sparse or no vegetation –
0.5%.
Futher habitat description. D5.1 Reedbeds normally without free-standing water; D5.2 Beds
of large sedges normally without free-standing water; E1.2 Perennial calcareous grassland
and basic steppes; E1.9 Open non-Mediterranean dry acid and neutral grassland including
inland dune grassland (× E1.1); E2.5 Meadows of the steppe zone; E6.2 Coninental inland
salt steppes; F3.1 Temperate thickets and scrub; G1.5 Broadleaved swamp woodland on
acid peat; G1.7 Thermophilous deciduous woodland; G1.9 Non-riverine woodland with
Betula, Populus tremula or Sorbus aucuparia; H5.3 Sparsely- or un-vegetated habitats on
mineral substrates not resulting from recent ice activity.
Land use: mowing/hay making – minor, nature conservation and research – 67%.
Protected areas: included in Chernomorskiy (Chornomorskyi, Black Sea) biosphere reserve
(core zone of the reserve), included in Emerald Site “Black Sea Biosphere Reserve”.
Threats: –.
General description. Major habitat type is the sand steppe. Other important habitats types
are freshwater permanent lakes, mesic grasslands, eutrophic mires, deciduous forests
(swamp and mesic), halophytic vegetation. Sand steppes are dominated by Artemisia
marschalliana, Festuca beckeri, Helichrysum corymbiforme, Koeleria sabuletorum, Thymus
borysthenicus, Stipa borysthenica. Typical species: Agropyron dasyanthum, Goniolimon
graminifolium, Polygonum arenarium. Prevailing species of mesic grasslands are Agrostis
sabulicola, Calamagrosts epigeios, Cynodon dactylon, Elytrigia elongata, Elytrigia repens,
Phleum phleoides, Poa angustifolia. Besides there are communiies of Puccinellia gigantea,
Salicornia perennans, Tripolium vulgare, Carex riparia, Phragmites australis, Lemna
minor, Lemna trisulca. Dominants of the the herb layer of Alder gluinosa swamp woods
are Calamagrosis epigeios, Carex riparia. Constant species: Eupatorium cannabinum,
Lysimachia vulgaris, Urica dioica. In Betula borysthenica woods, the shrub layer consists of
Rhamnus catharica, Sambucus nigra. Dominants of the herb layer are Agrostis sabulicola,
Calamagrostis epigeios, Carex elata, Phragmites australis.
Botanical significance. Important area for conservaion of steppe swamp Alnus gluinosa
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woods and some psammophyic species.
Criterion A
• Allium savranicum Besser; A(iv); abundance: rare; trend: unknown; species data quality:
medium; trend data quality: poor.
• Alyssum savranicum Andrz. ex Besser (Odontarrhena savranica (Andrz. ex Besser)
D.A.German); A(iv); abundance: rare; trend: unknown; species data quality: medium;
trend data quality: poor.
• Goniolimon graminifolium (Ait.) Boiss.; A(iv); abundance: frequent; trend: unknown;
species data quality: medium; trend data quality: poor.
Criterion C
• G1.414 Steppe swamp Alnus gluinosa woods; area: 11 ha; trend: stable; area data
quality: good; trend data quality: medium.
Literature
1. Кузнєцова Г.О., Протопопова В.В., Саричева З.А. Флора і рослинність Волижиного
лісу в Чорноморському заповіднику // Укр. ботан. журн. – 1969. – 26. № 4. – С. 35–40.
2. Уманець О.Ю. БЗ Чорноморський // Фіторізноманіття заповідників і національних
природних парків України. Ч. 1. Біосферні заповідники. Природні заповідники / під
ред. В.А. Онищенка і Т.Л. Андрієнко. – Київ: Фітосоціоцентр, 2012. – С. 73–93.
Threats: abandonment/reducion of land management – low; burning of vegetaion –
medium, extracion (minerals/quarries) – low, forestry (aforestaion) – medium.
General description. Slopes of the valley of the Vovcha river with chalk outcrops. The major
habitat type is the steppe. Dominants of the steppe vegetaion are Bromopsis riparia, Carex
humilis, Caragana frutex, Chamaecyisus austriacus, Chamaecythisus ruthenicus, Festuca
valesiaca, Koeleria cristata, Poa angusifolia, Sipa capillata, Sipa pulcherrima, Sipa pennata,
Salvia nutans. Major dominants of the chalk outcrops are Thymus cretaceus, Artemisia
hololeuca, Asperula tephrocarpa, Onosma tanaiica. Someimes there dominate Androsace
koso-poljanskii, Hyssopus cretaceus, Linum ucrainicum, Onosma tanaiica, Polygala
cretacea, Scutellaria creicola, Ephedra distachia. Other species of the outcrops: Gypsophila
alissima, Helianthemum cretaceum. Natural forests are dominated by Quercus robur, Tilia
cordata, Acer platanoides. There are also plantaions of Pinus sylvestris, Pinus pallasiana,
Quercus robur, Fraxinus excelsior, Tilia cordata, Cotynus coggygria. Rare endemic species
Vovchanski Skhyly
V.A. Onyshchenko, O.V. Bezrodnova, M.V. Banik
Ukrainian name: Вовчанські схили.
Transliteraion/Translaion variants: Vovcha Slopes.
Area: 1131.6 ha.
Altitude: 120–200 m.
Latitude: 50°22’35” N (50.3763°).
Longitude: 37°17’23” E (37.2898°).
Administrative regions. Kharkiv region: Vovchansk raion.
Ownership: state.
Biogeographic regions: coninental.
Habitats. Level 1. E – 72.7%; F – 2.0%; G – 17.3%; H – 8.0%.
Habitats. Level 2. E1 Dry grasslands – 72.7%; F3 Temperate and mediterranean-montane
scrub – 2.0%; G1 Broadleaved deciduous woodland – 15.4%; G3 Coniferous woodland –
1.9%; H2 Screes – 8.0%.
Futher habitat description. E1.1 Inland sand and rock with open vegetation; E1.2
Perennial calcareous grassland and basic steppes; F3.1 Temperate thickets and scrub;
G1.7 Thermophilous deciduous woodland; G1.A Meso- and eutrophic Quercus, Carpinus,
Fraxinus, Acer, Tilia, Ulmus and related woodland; G1.C Highly artificial broadleaved
deciduous forestry plantations; G3.F Highly artificial coniferous plantations; H2.6
Calcareous and ultra-basic screes of warm exposures.
Land use: agriculture (animals) – major; forestry – minor, nature conservaion and research
– minor.
Protected areas: includes Vovchanskyi state botanical reserve (185 ha), overlaps (18 ha)
with Siverskodonetskyi regional landscape reserve, overlaps (about 1050 ha) with Emerald
Site “Siverskodonetskyi”.
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Daphne sophia occurs mainly at forest margins.
Botanical significance. Important area for conservaion of endemic species Daphne sophia
and Androsace koso-poljanskii and vegetaion of chalk outcrops.
Criterion A
• Androsace koso-poljanskii Ovcz.; A(iv); abundance: frequent; trend: stable; species data
quality: good; trend data quality: medium.
• Daphne sophia Kalen.; A(i), A(iv); abundance: rare (675 individuals in 3 localiies); trend:
stable; species data quality: good; trend data quality: medium.
Criterion C
• E1.13 Coninental dry rocky steppic grasslands and dwarf scrub on chalk outcrops;
area: 90 ha; trend: stable; area data quality: medium; trend data quality: medium.
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Literature
1. Banik M. Daphne’04. Final report. – Kharkiv, 2006. – 20 p. (http://www.rufford.org/
files/.01.04%20Detailed%Final%20Report.pdf).
2. Банік М.В., Тверетинова В.В., Волкова Р.Є., Атемасова Т.А., Атемасов А.А.,
Брезгунова О.О., Влащенко А.С., Гончаров Г.Л., Коноваленко С.В., Скоробогатов
В.М., Скоробогатов Є.В., Целіщев О.Г. Нові місцезнаходження Daphne sophiae Kalen.,
(Thymeleaceae) в Україні // Укр. ботан. журн. – 2007. – 64, № 4. – С. 565–569.
3. Ермоленко Е.Д., Горелова Е.Н., Кушнарева Ю.И. К флоре и растительности
меловых обнажений рек Волчьей и Оскол в Харьковской области // Вест. ХГУ. – 1981. –
№ 211. – С. 6–11.
4. Коротченко І.А. Рідкісні види флори степів південної частини Лівобережного
Лісостепу України // Наукові основи збереження біотичної різноманітності / Темат. зб.
ін-ту екології Карпат НАН України. – Вип. 3. – Львів: Ліга-Прес, 2001. – С. 26–36
5. Расевич В.В. Еколого-ценотичні особливості популяцій Daphne sophia Kalen. у
природній флорі України // Укр. ботан. журн. – 2008. – 65, № 1. – С. 90–103.
6. Смолко С.С. Третинний релікт – вовчі ягоди Софії (Daphne sophia Kalen.) на
Середньоросійській височині та його сучасне поширення // Укр. ботан. журн. – 1967. –
24, № 1. – С. 69–75.
7. Червона книга України. Рослинний світ / за ред. Я.П. Дідуха. – К.: Глобалконсалтинг,
2009. – 900 с.
Vtenske Boloto
V.A. Onyshchenko
Ukrainian name: Втенське болото.
Transliteraion/Translaion variants: Vtenske Swamp, Vtenske Bog.
Area: 179 ha.
Altitude: 155–160 m.
Latitude: 51°37’31” N (51.6251°).
Longitude: 23°41’01” E (23.6835°).
Administrative regions. Volynska region: Shatsk raion.
Ownership: state.
Biogeographic regions: continental.
Habitats. Level 1. G – 100%.
Habitats. Level 2. G1 Broadleaved deciduous woodland – 4.0%; G3 Coniferous woodland –
76.0%; G4 Mixed deciduous and coniferous woodland – 20.0%.
Futher habitat description. G1.5 Broadleaved swamp woodland on acid peat; G3.E Nemoral
bog conifer woodland, G4.1 Mixed swamp woodland.
Land use: nature conservation and research – 100%.
Protected areas: includes Vtenskyi state botanical reserve (130 ha), included in biosphere
reserve “West Polesie”, included in Shatskyi naional nature park, included in Emerald Site
“Shatskyi”.
Threats: water (drainage) – low.
General description. Wooded bog. The tree layer is dominated by low Pinus sylvestris, oten
with admixture of Betula pubescens. Dominant species of the lower layers are Sphagnum
magellanicum, Sphagnum angusifolium, Eriophorum vaginatum, Vaccinium oxycoccos
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(Oxycoccus palustris). Other frequent species: Ledum palustre, Andromeda polifolia, Drosera
rotundifolia, Calluna vulgaris.
Botanical significance. Important site of Pinus sylvestris bogs.
Criterion C
• G3.E Nemoral bog conifer woodland; area: 90 ha; trend: stable; area data quality:
medium; trend data quality: medium.
Literature
1. Природно-заповідний фонд Волинської області / Упор.: М. Химин та ін. – Луцьк:
Ініціал, 1999. – 48 с.
2. Фіторізноманіття Українського Полісся та його охорона / Під заг. ред. Т.Л.
Андрієнко. – Київ: Фітосоціоцентр, 2006. – 316 с.
Vymushiv
I.I. Chorney, A.I. Tokaryuk, V.V. Budzhak
Ukrainian name: Вимушів.
Area: 8.8 ha.
Altitude: 258–282 m.
Latitude: 48°38’31” N (48.6418°).
Longitude: 25°39’37” E (25.6602°).
Administrative regions. Chernivtsi region: Zastavna raion.
Ownership: state.
Biogeographic regions: coninental.
Habitats. Level 1: E – 90.0%, F – 9.0%, H – 1.0%.
Habitats. Level 2: E1 Dry grasslands – 90.0%; F3 Temperate and mediterranean-montane
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scrub – 9.0%; H3 Inland cliffs, rock pavements and outcrops – 1.0%.
Futher habitat description: E1.2 Perennial calcareous grassland and basic steppes; F3.1
Temperate thickets and scrub; H3.2 Basic and ultra-basic inland cliffs.
Land use: agriculture (animals) – minor.
Vyshnivka
I.I. Chorney, A.I. Tokaryuk, V.V. Budzhak
Ukrainian name: Вишнівка.
Area: 309 ha.
Altitude: 244–329 m.
Latitude: 48°34’31” N (48.5751°).
Longitude: 25°23’53” E (25.3981°).
Administrative regions. Ivano-Frankivsk region: Horodenka raion, Sniatyn raion.
Ownership: state, private.
Biogeographic regions: coninental.
Protected areas: –.
Threats: abandonment/reducion of land management – low, extracion (mineral/quarries)
– low.
General description. Steep slopes with dry grasslands and outcrops of gypsum and limestone.
Grasslands are dominated by Lembotropis nigricans, Elytrigia intermedia. Frequent species:
Adonis vernalis, Agrimonia eupatoria, Anthericum ramosum, Arrhenatherum elatius,
Asyneuma canescens, Campanula glomerata, Centaurea jacea, Clematis integrifolia,
Crinitaria linosyris, Cruciata glabra, Dactylis glomerata, Eryngium campestre, Falcaria
vulgaris, Filipendula vulgaris, Galium verum, Inula ensifolia, Knautia arvensis, Medicago
romanica, Salvia pratensis, Salvia verticillata, Scabiosa ochroleuca, Securigera varia,
Thalictrum simplex, Trifolium montanum.
Botanical significance. Important area for the endemic species Aconitum pseudanthora.
Criterion A
• Aconitum pseudanthora Błocki ex Pacz. A(iii); abundance: occasional; trend: unknown;
species data quality: medium; trend data quality: poor.
Conservaion proposals. Create a state botanical reserve or nature monument.
Habitats. Level 1: C – 1.7%, E – 97.3%, F – 1.0%.
Habitats. Level 2: C1 Surface standing waters – 1.3%, C3 Littoral zone of inland surface
waterbodies – 0.4%, E1 Dry grasslands – 75.3%; E2 Mesic grasslands – 19.0%, E3 Seasonally
wet and wet grasslands – 3.0%; F3 Temperate and mediterranean-montane scrub – 1.0%.
Futher habitat description: E1.2 Perennial calcareous grassland and basic steppes; E2.2
Low and medium alitude hay meadows; E3.4 Moist or wet eutrophic and mesotrophic
grassland; F3.1 Temperate thickets and scrub.
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341
Land use: agriculture (animals) – minor.
Protected areas: –.
Threats: agricultural intensification/expansion (arable) – medium.
General description. Grasslands dominated by Brachypodium pinnatum with presence of
Arrhenatherum elatius, Briza media, Cirsium pannonicum, Festuca rupicola, Filipendula
vulgaris, Galium boreale, Galium verum, Inula salicina, Iris hungarica, Koeleria cristata,
Lathyrus pannonicus, Potentilla alba, Prunella grandiflora, Pyrethrum corymbosum,
Salvia pratensis, Salvia verticillata, Sanguisorba officinalis, Serratula lycopifolia, Trifolium
montanum.
Botanical significance. One of the largest areas of dry grasslans in western Ukraine with
presence of Pulsailla grandis, Iris hungarica, Serratula lycopifolia.
Criterion A
• Serratula lycopifolia (Vill.) A. Kern.; А (ii); abundance: rare; trend: unknown; species data
quality: medium; trend data quality: poor.
Criterion C
• E1.2 Perennial calcareous grassland and basic steppes; area: 70 ha; trend: stable; area
data quality: good; trend data quality: medium.
Conservaion proposals. Do not allow land plowing. Create a state botanical reserve and
an Emerald Site.
austriacum, Dryopteris dilatata, Dryopteris carthusiana, Geniana asclepiadea, Luzula
pilosa, Luzula sylvaica, Polytrichum formosum, Prenanthes purpurea, Sorbus aucuparia,
Vaccinium viis-idaea. Pinus cembra is the main dominant in an area of 18 ha. Besides there
are Pinus mugo scrub and coarse sandstone screes.
Yaiko Perehinske
V.A. Onyshchenko
Ukrainian name: Яйко Перегінське.
Area: 358.0 ha.
Altitude: 1000–1596 m.
Latitude: 48°40’03” N (48.6674°).
Longitude: 23°54’05” E (23.9014°).
Administrative regions. Ivano-Frankivsk region: Rozhniaiv raion.
Ownership: state.
Biogeographic regions: alpine.
Habitats. Level 1. F – 2.4%; G – 95.0%; H – 2.6%.
Habitats. Level 2. F2 Arctic, alpine and subalpine scrub – 2.4%; G1 Broadleaved deciduous
woodland – 0.5%; G3 Coniferous woodland – 92.0%; G4 Mixed deciduous and coniferous
woodland – 2.5%; H2 Screes – 2.5%; H3 Inland cliffs, rock pavements and outcrops – 0.1%.
Futher habitat description. F2.4 Conifer scrub close to the tree limit; G1.6 Fagus
woodland; G3.1 Abies and Picea woodland; G4.6 Mixed Abies – Picea – Fagus woodland;
H2.3 Temperate-montane acid siliceous screes; H3.1 Acid silicious inland clifs.
Land use: forestry – major; nature conservation and research – major.
Protected areas: includes Yaikivskyi state botanical reserve (270 ha), included in Emerald
Site “Dolynsko-Rozhniatynskyi”.
Threats: forestry (intensiied forest management) – low.
General description.
Predominant vegetaion is Picea abies forests with admixture of Pinus cembra. Lower layers
are dominated by Vaccinium myrillus, Homogyne alpina, Oxalis acetosella, Calamagrosis
arundinacea. Other species of high constancy are Dicranum scoparium, Doronicum
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Botanical significance. Important area for Carpathian Pinus cembra forests.
Criterion C
• G3.25 Carpathian Larix and Pinus cembra forests; area: 18 ha; trend: decreasing; area
data quality: good; trend data quality: medium.
Literature
1. Природно-заповідні території та об’єкти Івано-Франківщини. – Івано-Франківськ,
2000. – 272 с.
Yelanetskyi Step
V.A. Onyshchenko
Ukrainian name: Єланецький степ.
Transliteraion/Translaion variants: Yelanets Steppe.
Area: 1675.7 ha.
Altitude: 30–100 m.
Latitude: 47°33’31” N (47.5586°).
Longitude: 32°01’27” E (32.0242°).
Administrative regions. Mykolaiv region: Yelanets raion, Nova Odesa raion.
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Habitats. Level 1. E – 98.1%; F – 0.2%; G – 1.5%; H – 0.2%
Habitats. Level 2. E1 Dry grasslands – 87.1%; E2 Mesic grasslands – 11.0%; F3 Temperate
and mediterranean-montane scrub – 0.2%; G1 Broadleaved deciduous woodland – 1.5%;
H3 Inland cliffs, rock pavements and outcrops – 0.02%; H5 Miscellaneous inland habitats
with very sparse or no vegetation – 0.2%.
Futher habitat description. E1.2 Perennial calcareous grassland and basic steppes; F3.1
Temperate thickets and scrub; G1.C Highly ariicial broadleaved deciduous forestry
plantaions; I1.5 Bare illed, fallow or recently abandoned arable land.
Land use: agriculture (animals) – minor; nature conservaion and research – 100%.
Protected areas: same as Yelanetskyi Step nature reserve and Emerald Site “Yelanetskyi
Step Nature Reserve”.
Threats: –.
General description. Valleys of intermitent streams, mainly with the steppe vegetaion.
Mesic grasslands occupy botoms of the valleys. There are ourcrops of limestone and granite.
Dry grasslands are dominated by Sipa capillata (major), Sipa lessingiana (major), Festuca
valesiaca (major), Botriochloa ischaemum (major), Sipa ucrainica, Sipa pulcherrima, Sipa
irsa. A signiicant part of steppes (about 75 ha) are co-dominated by Caragana frutex,
rarely by other shrub species (Caragana scythica, Chamaecyisus graniicus). Other typical
specis of the steppe communiies: Achillea nobilis, Bromopsis riparia, Eryngium campestre,
Euphorbia seguierana, Euphorbia stepposa, Koeleria cristata, Koeleria lobata, Marrubium
praecox, Securigera varia, Teucrium polium. In petrophyic variant of the steppe, Thymus
dimorphus co-dominates. Main species of calcareous outcrops and screes are Thymus
dimorphus, Jurinea mulilora, Jurinea brachycephala, Centaurea marschalliana, Koeleria
brevis. Dominants of other vegetaion types: mesic grasslands – Elytrigia repens, Poa
angusifolia, Bromopsis inermis, reedbed – Phragmites australis, Typha laifolia; shrubs –
Crataegus curvisepala s.l., abandoned arable lands – Elytrigia repens, Artemisia absinthium,
Cirsium arvense, Melilotus oicinalis, Bunias orientalis.
Botanical significance. Important for conservaion of Chamaecyisus graniicus and Tulipa
hypanica. One of the best steppe areas west to the Dnipro river.
Criterion A
• Chamaecyisus graniicus (Rehmann.) Rothm.; A(iii); abundance: occasional; trend:
stable; species data quality: medium; trend data quality: poor.
• Tulipa hypanica Klokov et Zoz; A(iii); abundance: occasional; trend: stable; species data
quality: medium; trend data quality: poor.
Criterion C
• E1.2 Perennial calcareous grassland and basic steppes; area: 1300 ha; trend: stable;
area data quality: good; trend data quality: good .
Literature
1. Бойко М.Ф. Анотований список мохоподібних заповідника “Єланецький степ”
та прилеглих територій (Миколаївська область, Україна) // Чорноморськ. бот. журн. –
2009. – 5, № 4. – С. 583–588.
2. Бойко Т.О. Лишайники та ліхенофільні гриби вапнякових відслонень природного
заповідника “Єланецький степ” // Чорноморськ. бот. журн. – 2008. – 4, № 1. – С. 84–89.
3. Бойко Т.О. Анотований список лишайників та ліхенофільних грибів природного
заповідника “Єланецький степ” // Чорноморськ. бот. журн. – 2009. – 5, № 3. –
С. 448–458.
4. Бойко Т.О. Нові та рідкісні для України лишайники з природного заповідника
“Єланецький степ” // Чорноморськ. бот. журн. – 2009. – 5, № 2. – С. 241–247.
5. Воронова С.М. Фітораритети судинних рослин міжнародного та державного
природоохоронного статусу у природному заповіднику “Єланецький степ” // Вісник
Національного науково-природничого музею України. Серія ботанічна. Частина друга.
– 2005–2007. – № 4–5. – С. 298–333.
6. Воронова С.М. Раритетний фітофлорофонд та природно-заповідна мережа
Єланецько-Інгульського регіону // Заповідна справа в Україні. – 2008. – Т. 14, вип. 1. –
С. 66–70.
7. Коломійчук В.П., Мойсієнко І.І., Деркач О.М., Бойко Т.О. ПЗ Єланецький степ //
Фіторізноманіття заповідників і національних природних парків України. Ч. 1. Біосферні
заповідники. Природні заповідники / під ред. В.А. Онищенка і Т.Л. Андрієнко. – Київ:
Фітосоціоцентр, 2012. – С. 124–138.
8. Костильов О.В. Рослинність запроектованого заповідника “Єланецький” // Укр.
ботан. журн. – 1987. – 44, № 2. – С. 77–81.
9. Костылев А.В. Еланецкий (заповедник) / Перспективная сеть заповедных
объектов Украины. – Киев: Наукова думка, 1987. – С. 199–204.
10. Мойсієнко І.І., Соломаха В.А., Драбинюк Г.В., Соломаха Т.Д. Еколого-ценотичні
особливості Scutellaria verna Besser в умовах природного заповідника “Єланецький
степ” // Чорноморський ботанічний журнал. – 2005. – 1, № 2. – С. 83–91.
11. Ткаченко В.С. Структурні зміни в рослинному покриві “Єланецького степу” за
перше десятиліття заповідання // Чорноморський ботанічний журнал. – 2009. – 5, № 3.
– С. 319–332.
12. Ткаченко В.С. Зміни ектопічних характеристик заповідника “Єланецький степ”
в першому десятилітті його існування // Чорноморський ботанічний журнал. – 2009. –
5, № 4. – С. 475–490.
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Ownership: state.
Biogeographic regions: steppic.
13. Ткаченко В.С., Сиротенко П.О. Вихідний стан рослинності “Єланецького
степу” в системі фітоценотичного моніторингу // Укр. ботан. журн. – 1999. – 56, № 6. –
С. 623–629.
Yemilchynski Lisy
Ukrainian name: Ємільчинські ліси.
Transliteraion/Translaion variants: Yemilchyne Forests.
Area: 2645.0 ha.
Altitude: 192–203 m.
Latitude: 50°59’28” N (50.9911°).
Longitude: 27°36’43” E (27.6118°).
Administrative regions. Zhytomyr region: Yemilchyne raion.
Ownership: state.
Biogeographic regions: coninental.
Habitats. Level 1. D – 0.6%; F – 0.5%; G – 97.9%; H – 1.0%.
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O.O. Orlov, V.A. Onyshchenko
Habitats. Level 2. D2 – Valley mires, poor fens and transition mires – 0.6%; F9 Riverine and
fen scrubs – 0.5%; G1 Broadleaved deciduous woodland – 32.0%; G3 Coniferous woodland
– 29.0%; G4 Mixed deciduous and coniferous woodland – 34.2%; G5 Lines of trees, small
anthropogenic woodlands, recently felled woodland, early-stage woodland and coppice –
2.7%; H5 Miscellaneous inland habitats with very sparse or no vegetation – 1.0%.
Futher habitat description. D2.3 Transition mires and quaking bogs; F9.2 Salix carr and
fen scrub; G1.5 Broadleaved swamp woodland on acid peat; G1.8 Acidophilous Quercusdominated woodland; G1.9 Non-riverine woodland with Betula, Populus tremula or Sorbus
aucuparia; G3.4 Pinus sylvestris woodland south of the taiga; G4.7 Mixed Pinus sylvestris –
acidophilous Quercus woodland; G5.6 Early-stage natural and semi-natural woodlands and
regrowth; H5.6 Trampled areas.
Land use: forestry – major; nature conservaion and research – 17%.
Protected areas: includes Yuzykhivka regional forest reserve (439 ha).
Threats: forestry (intensiied forest management) – medium.
General description. Vegetaion is represented by acidophilous Pinus sylvestris, Quercus
robur, Betula pendula woods, and Betula pubescens swamps. Dominants of the wet pine
woods are Franula alnus, Molinia caerulea, Pteridium aquilinum, Vaccinium myrillus,
Dicranum polysetum, Pleurozium shreberi. Other frequent species are Dryopteris
carthusiana, Trientalis europaea (Lysimachia europaea), Vaccinium uliginosum, Vaccinium
viis-idaea. On the drier soils, there are species rich Pinus sylvestris woods with dominance
of Rhododendron luteum in the shrub layer. Dominants of the herb layer are Convallaria
majalis and Pteridium aquilinum. Constant species: Calamagrosis arundinacea, Carex
montana, Galium intermedium, Maianthemum bifolium, Polygonatum odoratum, Primula
veris, Serratula coronata. Diferenial species of the swamped pine woods are Andromeda
polifolia, Carex nigra, Ledum palustre, Sphagnum fallax, Sphagnum palustre. In acidophilous
Quercus robur woods, there are the shrub layer consising of Frangula alnus. The herb layer
is dominanted by Anemone nemorosa, Luzula pilosa, Lysimachia vulgaris, Oxalis acetosella.
Dominants of the mesotrophic swamps are Betula pubescens, Carex lasiocarpa, Juncus
conglomeratus, Molinia caerulea, Sphagnum fallax.
Botanical significance. Important for conservaion of Rhododendron lutem and acidophilous
oak forests.
Criterion A
• Rhododendron luteum Sweet; A(ii); abundance: frequent; trend: stable; species data
quality: good; trend data quality: medium.
Criterion C
• G1.8 Acidophilous Quercus-dominated woodland; area: 800 ha; trend: stable; area data
quality: medium; trend data quality: medium.
Literature
1. Орлов О.О., Сіренький С.П., Якушенко Д.М., Жижин М.П., Степаненко М.А.,
Тарасевич О.В. Природно-заповідний фонд Житомирської області. Довідник / За заг.
ред. О.О. Орлова. – Житомир – Новоград-Волинський: Вид-во НОВОград, 2015. – 404 с.
2. Орлов О.О., Якушенко Д.М., Воробйов Є.О. Флористична класифікація лісів із
участю Rhododendron luteum Sweet та радіоекологічна оцінка їх асоціацій в Поліссі
України. І. Синтаксономія лісів із участю Rhododendron luteum // Укр. фітоцен. зб. –
2000. - Серія А, Вип. 1 (16). – Київ: Фітосоціоцентр. – С. 94–113.
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Yulivski Hory
R.Ya. Kish, V.A. Onyshchenko
Ukrainian name: Юлівські гори.
Transliteraion/Translaion variants: Yulivski Gory, Yulivs’ki Hills.
Area: 601 ha.
Altitude: 129–318 m.
Latitude: 48°01’35” N (48.0265°).
Longitude: 23°05’06” E (23.0849°).
Administrative regions. Zakarpatska region: Vynohradiv raion.
Ownership: state.
Biogeographic regions: pannonian.
biosphere reserve (naional caregory), includes state botanical reserve “Yulivska Hora” (176
ha, belongs to the biosphere reserve), includes regional botanical reserve “Kholmovetska
Hora” (95.4 ha), overlaps (176 ha) with Emerald Site “Carpathian Biosphere Reserve”,
overlaps (227 ha) with Emerald Site “Okli-Hed”, overlaps (32.4 ha) with proposed regional
botanical reserve “Dibrovy Zatysianshchyny”.
Threats: abandonment/reducion of land management – low, consequences of invasive
species – low, forestry (intensified forest management) – low.
General description. Hills composed of tuf, andesite, basalt, and sedimentary rocks. The
major vegetaion type is the termophilous oak wood. The tree layer consists of Quercus
petraea, Cerasus avium, Carpinus betulus, Acer tataricum, someimes with Quercus
dalechampii, Quercus polycarpa, Quercus cerris, Tilia tomentosa. In the shrub layer there are
Acer tataricum, Corylus avellana, Crataegus monogyna s. l., Euonymus europaeus, Sambucus
nigra, Swida sanguinea (Cornus sanguinea). Typical species of the herb layer are Anthericum
ramosum, Brachypodium pinnatum, Campanula persicifolia, Carex michelii, Clinopodium
vulgare, Digitalis grandilora, Hieracium murorum, Hylotelephium polonicum, Inula ensifolia,
Lathyrus niger, Melittis melissophyllum, Poa nemoralis, Polygonatum odoratum, Potenilla
alba, Pyrethrum corymbosum, Veronica teucrium, Vincetoxicum hirundinaria, Viscaria
viscosa. In woods on steep southern slopes, there occur Brachypodium pinnatum, Festuca
valesiaca, Galium campanulatum, Inula hirta, Phleum phleoides, Poa angusifolia, Stachys
recta, Veronica spicata. Besides there are beech forests (with Allium ursinum, Asarum
europaeum, Dentaria bulbifera (Cardamine bulbifera), Lamium galeobdolon, Mercurialis
perennis, Pulmonaria obscura), oak-hornbeam forests and small areas of dry grasslands
(dominated by Festuca rupicola, Festuca pseudodalmaica) and shrubs.
Botanical significance. The largest locaion of termophilous oak woods in Transcarpathian
part of Ukraine.
Criterion C
• G1.7 Thermophilous deciduous woodland (*91H0, *91I0, 91M0); area: 400 ha; trend:
stable; area data quality: medium; trend data quality: medium.
Conservaion proposals. Remove alien tree species. Do not allow the expansion of shrubs in
grasslands. Create proposed botanical reserve “Dibrovy Zatysianshchyny”.
Literature
1. Стойко С.М. Дубові ліси Українських Карпат: екологічні особливості, відтворення,
охорона. – Львів, 2009. – 220 с.
2. Фіторізноманіття заповідників і національних природних парків України.
Ч.1. Біосферні заповідники. Природні заповідники / Колектив авторів під ред. В.А.
Онищенка і Т.Л. Андрієнко. – Київ: Фітосоціоцентр, 2012. – 406 с.
Habitats. Level 1. E – 0.3%; F – 0.1%; G – 99.0%; H – 0.6%.
Habitats. Level 2. E1 Dry grasslands – 0.3%; F3 Temperate and mediterranean-montane
scrub – 0.1%; G1 Broadleaved deciduous woodland – 99.0%; H3 Inland cliffs, rock
pavements and outcrops – 0.0%; H5 Miscellaneous inland habitats with very sparse or no
vegetation – 0.6%.
Futher habitat description. E1.2 Perennial calcareous grassland and basic steppes – 0.3%;
F3.1 Temperate thickets and scrub; F3.2 Submediterranean deciduous thickets and brushes;
G1.6 Fagus woodland; G1.7 Thermophilous deciduous woodland; G1.A Meso- and eutrophic
Quercus, Carpinus, Fraxinus, Acer, Tilia, Ulmus and related woodland; H5.6 Trampled areas.
Land use: forestry – 99%, nature conservation and research – 82%.
Protected areas: overlaps (176 ha) with Carpathian biosphere reserve (UNESCO) (includes
cluster “Yulivskai Hora” of the biosphere reserve), overlaps (176 ha) with the Karpatskyi
V.A. Onyshchenko, O.V. Lukash
Ukrainian name: Заплава Десни.
Transliteraion/Translaion variants: Floodplain of the Desna River.
Area: 185741 ha.
Altitude: 95–130 m.
Latitude: 51°23’17” N (51.3881°).
Longitude: 32°11’12” E (32.1865°).
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Zaplava Desny
Administrative regions. Chernihiv region: Borzna raion, Chernihiv city, Chernihiv raion,
Kozelets raion, Korop raion, Kulykivka raion, Mena raion, Novhorod-Siverskyi city, NovhorodSiverskyi raion, Sosnytsia raion; Kyiv region: Brovary raion, Vyshhorod raion; Sumy region:
Krolevets raion, Shostka raion, Seredyna-Buda raion.
Ownership: state, private.
Biogeographic regions: continental.
Habitats. Level 1. C – 5%; D – 15%; E – 68%; F – 1%; G – 9%; H – 1%; I – 1%.
Habitats. Level 2. C1 Surface standing waters – 2%; C2 Surface running waters – 3%; C3
Littoral zone of inland surface waterbodies – 1%; D5 Sedge and reedbeds, normally without
free-standing water – 15%; E2 Mesic grasslands – 33%; E3 Seasonally wet and wet grasslands
– 35%; E5 Woodland fringes and clearings and tall forb stands – 5.3%; F9 Riverine and fen
scrubs – 1%; G1 Broadleaved deciduous woodland – 6%; G3 Coniferous woodland – 3%; H5
Miscellaneous inland habitats with very sparse or no vegetation – 0.5%; I1 Arable land and
market gardens – 1%.
Futher habitat description. C1.2 Permanent mesotrophic lakes, ponds and pools; C1.6
Temporary lakes, ponds and pools; C2.3 Permanent non-tidal, smooth-flowing watercourses;
C3.2 Water-fringing reedbeds and tall helophytes other than canes; C3.4 Species-poor beds
of low-growing water-fringing or amphibious vegetation; C3.5 Periodically inundated shores
with pioneer and ephemeral vegetation; C3.6 Unvegetated or sparsely vegetated shores
with soft or mobile sediments; D5.1 Reedbeds normally without free-standing water; D5.2
Beds of large sedges normally without free-standing water; E2.1 Permanent mesotrophic
pastures and aftermath-grazed meadows; E2.2 Low and medium alitude hay meadows;
E3.4 Moist or wet eutrophic and mesotrophic grassland; E5.4 Moist or wet tall-herb and
fern fringes and meadows; F9.1 Riverine scrub; F9.2 Salix carr and fen scrub; G1.1 Riparian
and gallery woodland, with dominant Alnus, Betula, Populus or Salix; G1.2 Mixed riparian
floodplain and gallery woodland; G1.4 Broadleaved swamp woodland not on acid peat; G3.4
Pinus sylvestris woodland south of the taiga.
Land use: agriculture (animals) – 15%; agriculture (arable) – 1%; forestry – 9%; mowing/hay
making – 45%; nature conservation and research – 5%; urban/industrial/transport – minor.
Protected areas: overlaps (3743 ha) with Desniansko-Starohutskyi national nature park,
overlaps (14038 ha) with Desnianskyi biosphere reserve, overlaps (about 5007 ha) with
Mezynskyi national nature park, overlaps with Mizhrichynskyi regional landscape park (about
11200 ha), includes Desna River Floodplains Ramsar Site (4270 ha), includes Kamoretskyi
state zoological reserve (515 ha), Obolonskyi state botanical reserve (400 ha), Putyvskyi
state botanical reserve (150 ha), Muravyivska state hydrological nature monument (40 ha),
Ozero Trubyn state hydrological nature monument (40 ha), Vaden’ state hydrological nature
monument (20 ha),
includes Babakove regional hydrological reserve (12 ha), Boloto Kolodlyve regional
hydrological reserve (13.3 ha), Blystovskyi regional landscape reserve (400 ha), Deminka
regional landscape reserve (1431 ha), Fedorove regional hydrological reserve (14 ha),
Horytskyi landscape hydrological reserve (796 ha), Kovchynskyi regional landscape reserve
(311 ha), Kyslyche regional hydrological reserve (178 ha), Lebedynske regional hydrological
reserve (184 ha), Makoshynskyi regional landscape reserve (1533 ha), Mialyne regional
hydrological reserve (102 ha), Oryvtsiove regional hydrological reserve (12 ha), PaikaKryvcha regional hydrological reserve (216 ha) Popovychove regional hydrological reserve
(11 ha), Smolianske regional hydrological reserve (15 ha), Spaskyi regional landscape reserve
(543 ha), Spaskyi–1 regional hydrological reserve (214 ha), Synychne regional hydrological
reserve (10 ha), Urochyshche Kuty regional botanical reserve (122 ha), Vuzke regional
hydrological reserve (11 ha), Zadesniansly regional landscape reserve (940 ha), Zolotynka
regional landscape reserve (527 ha),
includes Ramsar Site “Desna river loodplains” (4270 ha), includes proposed Ramsar Site
“Floodplains between town Oster and village Smolyn”, overlaps (82511 ha) with Emerald Site
“Chernihivske Podesennia”, overlaps (3743 ha) with Emerald Site “Desniansko-Starohutskyi
Naional Nature Park”, overlaps (14038 ha) with Emerald Site “Desnianskyi Biosphere
Reserve”, overlaps overlaps (5340 ha) with Emerald Site “Kyivske Podesennia”, overlaps
(13660 ha) with Emerald Site “Verhnie Podesennia”, overlaps (5007 ha) with Emerald Site
“Mezynskyi Naional Nature Park”, overlaps (1824 ha) with Emerald Site “Dolyna Seimu”,
overlaps (50750 ha) with Emerald Site “Nyzhnie Podesennia”, overlaps (7058 ha) with
Emerald Site “Mizhrichynskyi Regional Landscape Park”, overlaps with proposed naional
nature parks “Prydesnianskyi” and “Shostkynskyi”.
Threats: abandonment/reduction of land management – low, agricultural intensification/
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expansion (general) – low; burning of vegetation – low, development (urbanization) – low,
water (drainage) – low.
General description.
338 km section of the floodplain of the Desna river. This is the largest floodplain with
natural water regime in Ukraine. Its average width is about 5 km. Floods are regular. The
riverbed has natural meanders. There are many oxbows and lakes. Major vegetation types
are mesic meadows dominated by Alopecurus pratensis, Poa pratensis, Festuca pratensis,
Festuca rubra, Agrostis gigantea; drier sandy meadows dominated by Poa angustifolia,
Agrostis vinealis, Calamagrostis epigeios; moist medadows dominated by Deschampsia
cespitosa, Phalaroides arundinacea, Filipendula ulmaria, Geranium palustre; mires and
littoral vegetation dominated by Phragmites auistralis, Glyceria maxima, Carex acuta,
Carex acuiformis, Carex appropinquata, Carex juncella, Carex elata, Carex rostrata, Carex
vesicaria, Carex vulpina. There are forests (Salix alba, Quercus robur, Alnus glutinosa,
Populus nigra, Populus alba, Pinus sylvestris) and shrubs (Salix cinerea, Salix triandra, Salix
acutifolia). Aquatic vegetation is very diverse. Main dominants are Ceratophyllum demersum,
Lemna minor, Lemna trisulca, Hydrocharis morsus-ranae, Nuphar lutea, Nymphaea alba,
Nymphaea candida, Potamogeton natans, Sagittaria sagittifolia, Sparganium emersum,
Stratiotes aloides, Utricualaria vulgaris.
Botanical significance. This area is most important for conservation of floodplain complexes
in Ukraine.
Criterion C
• C1.223 Floating Stratiotes aloides rafts; area: 30 ha; trend: stable; area data quality:
poor; trend data quality: medium.
• C1.224 Floating Utricularia australis and Utricularia vulgaris colonies; area: 20 ha; trend:
stable; area data quality: poor; trend data quality: medium.
• C1.3411 Ranunculus communities in shallow water; area: 1 ha; trend: stable; area data
quality: poor; trend data quality: medium.
• C2.33 Mesotrophic vegetation of slow-flowing rivers; area: 200 ha; trend: stable; area
data quality: poor; trend data quality: medium.
• C3.4 Species-poor beds of low-growing water-fringing or amphibious vegetation ; area:
30 ha; trend: stable; area data quality: poor; trend data quality: medium.
• C3.51 Euro-Siberian dwarf annual amphibious swards; area: 30 ha; trend: stable; area
data quality: poor; trend data quality: medium.
• D5.2 Beds of large sedges normally without free-standing water; area: 25000 ha; trend:
stable; area data quality: poor; trend data quality: medium.
• E2.2 Low and medium alitude hay meadows; area: 30000 ha; trend: stable; area data
quality: medium; trend data quality: medium.
• E3.4 Moist or wet eutropic and mesotrophic grassland; area: 65000 ha; trend: stable;
area data quality: medium; trend data quality: medium.
• E3.5 Moist or wet oligotrophic grassland; area: 1000 ha; trend: stable; area data quality:
poor; trend data quality: poor.
• E5.4 Moist or wet tall-herb and fern fringes and meadows; area: 300 ha; trend: stable;
area data quality: poor; trend data quality: poor.
• F9.1 Riverine scrub; area: 5000 ha; trend: stable; area data quality: poor; trend data
quality: poor.
• G1.11 Riverine Salix woodland; area: 2000 ha; trend: stable; area data quality: poor;
trend data quality: poor.
• G1.21 Riverine Fraxinus – Alnus woodland, wet at high but not at low water; area: 200
352
ha; trend: stable; area data quality: poor; trend data quality: poor.
G1.22 Mixed Quercus – Ulmus – Fraxinus woodland of great rivers; area: 1000 ha; trend:
stable; area data quality: medium; trend data quality: poor.
Conservaion proposals. Сreate Emerald Sites including the enire IPA.
Literature
1. Андриенко Т.Л., Шеляг-Сосонко Ю.Р. Растительный мир Украинского Полесья в
аспекте его охраны. – Киев: Наук. думка, 1983. – 216 с.
2. Водно-болотні угіддя України / Під ред. Г.Б. Марушевського, І.С. Жарук. – К.:
Wetlands International Black Sea Programme, 2006. – 312 с.
3. Деснянський екологічний коридор / під заг. ред. В.Костюшина, Є.Прекрасної. –
К.: НЕЦУ, 2010. – 164 с.
4. Карпенко О.Ю. НПП Мезинський // Фіторізноманіття заповідників і національних
природних парків України. Ч. 2. Національні природні парки / під ред. В.А. Онищенка і
Т.Л. Андрієнко. – Київ: Фітосоціоцентр, 2012. – С. 312–320.
5. Панченко С.М. НПП Деснянсько-Старогутський // Фіторізноманіття заповідників
і національних природних парків України. Ч. 2. Національні природні парки / під ред.
В.А. Онищенка і Т.Л. Андрієнко. – Київ: Фітосоціоцентр, 2012. – С. 215–229.
6. Панченко С.М., Андрієнко Т.Л., Гаврись Г.Г., Кузьменко Ю.В. Екологічна мережа
Новгород-Сіверського Полісся. – Суми: Університетська книга, 2003. – 92 с.
7. Перспективная сеть заповедных объектов Украины / под ред. Ю.Р.ШелягСосонко. – К.: Наук. думка, 1987. – 292 с.
8. Смарагдова мережа в Україні / Колектив авторів під ред. Проценка Л.Д. – Київ:
Хімджест, 2011. – 192 с.
•
Zaplava Latorytsi
V.A. Onyshchenko
Ukrainian name: Заплава Латориці.
Transliteraion/Translaion variants: Floodplain of the Latorytsa River.
Area: 4752 ha.
Altitude: 99–110 m.
Latitude: 48°27’54” N (48.4649°).
Longitude: 22°21’15” E (22.3541°).
Administrative regions. Zakarpatska region: Uzhhorod raion, Mukachevo raion.
Ownership: state.
Biogeographic regions: pannonian.
Habitats. Level 1. C – 2.3%; D – 1.5%; E – 22.0%; F – 6.0%; G – 68.2%.
Habitats. Level 2. C1 Surface standing waters – 0.8%; C2 Surface running waters – 0.9%;
C3 Littoral zone of inland surface waterbodies – 0.6%; D5 Sedge and reedbeds, normally
without free-standing water – 1.5%; E3 Seasonally wet and wet grasslands – 22.0%; F9
Riverine and fen scrubs – 6.0%; G1 Broadleaved deciduous woodland – 68.2%.
Futher habitat descripion. C1.2 Permanent mesotrophic lakes, ponds and pools; C1.6
Temporary lakes, ponds and pools; C2.3 Permanent non-tidal, smooth-flowing watercourses;
C3.2 Water-fringing reedbeds and tall helophytes other than canes; C3.5 Periodically
inundated shores with pioneer and ephemeral vegetation; D5.1 Reedbeds normally without
free-standing water; D5.2 Beds of large sedges normally without free-standing water; E3.4
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Moist or wet eutrophic and mesotrophic grassland; F9.1 Riverine scrub; F9.2 Salix carr
and fen scrub; G1.1 Riparian and gallery woodland, with dominant Alnus, Betula, Populus
or Salix; G1.2 Mixed riparian floodplain and gallery woodland; G1.A Meso- and eutrophic
Quercus, Carpinus, Fraxinus, Acer, Tilia, Ulmus and related woodland.
Land use: agriculture (animals) – minor, forestry – 50%; mowing/hay making – minor, nature
conservation and research – major.
Protected areas: overlaps (about 4752 ha) with Prytysianskyi regional landscape park,
includes Velykodobronskyi state zoological reserve (1736 ha), overlaps (about 4493 ha) with
Emerald Site “Prytysianskyi”.
Threats: –.
General description. A part of the loodplain of the Latorytsia river. Moist Fraxinus pannonica
and Quercus robur forest occupies about half of the territory. There is also a large area of
wet meadows.
Botanical significance. Important area for loodplain ash-oak forests.
Criterion C
• G1.22 Mixed Quercus – Ulmus – Fraxinus woodland of great rivers; area: 3000 ha; trend:
stable; area data quality: medium; trend data quality: poor.
Literature
1. Регіональний ландшафтний парк “Притисянський” – збереження природної
спадщини рівнинного Закарпаття / Р. Кіш, Б. Проць, А. Поляновський, Т. Башта та ін.. –
Ужгород: Мистецька лінія, 2009. – 20 с.
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Zaplava Prypiai
V.A. Onyshchenko
Ukrainian name: Заплава Прип’яті.
Transliteraion/Translaion variants: Floodplain of the Prypiat’ (Pripet) River.
Area: 44488 ha.
Altitude: 137–154 m.
Latitude: 51°51’59” N (51.8664°).
Longitude: 25°25’08” E (25.4188°).
Administrative regions. Rivne region: Zarichne raion; Volynska region: Liubeshiv raion,
Ratno raion.
Ownership: state.
Biogeographic regions: coninental.
Habitats. Level 1. C – 3.5%; D – 59%; E – 15%; F – 13%; G – 9%; I – 0.5%.
Habitats. Level 2. C1 Surface standing waters – 3%; C2 surface running waters – 0.1%; C3
littoral zone of inland surface waterbodies – 0.4%; D2 Valley mires, poor fens and transition
mires – 10%; D5 Sedge and reedbeds, normally without free-standing water – 49%; E2
Mesic grasslands – 3%; E3 Seasonly wet and wet grasslands – 12%; F9 Riverine and fen
scrubs – 13%; G1 Broadleaved deciduous woodland – 6%; G3 Coniferous woodland – 1%;
G4 Mixed deciduous and coniferous woodland – 2%.
Futher habitat description. C1.2 Permanent mesotrophic lakes, ponds and pools; C2.3
Permanent non-tidal, smooth-lowing watercourses; C3.2 Water-fringing reedbeds and tall
helophytes other than canes; D2.2 Poor fens and sot-water spring mires; D2.3 Transiion
mires and quaking bogs; D5.1 Reedbeds normally without free-standing water; D5.2 Beds
of large sedges normally without free-standing water; E1.9 Open non-Mediterranean dry
acid and neutral grassland including inland dune grassland; E2.1 Permanent mesotrophic
pastures and aftermath-grazed meadows; E2.2 Low and medium altitude hay meadows;
E3.4 Moist or wet eutrophic and mesotrophic grassland; E3.5 Moist or wet oligotrophic
grassland; F9.2 Salix carr and fen scrub; G1.1 Riparian and gallery woodland, with dominant
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Alnus, Betula, Populus or Salix; G1.2 Mixed riparian floodplain and gallery woodland; G1.4
Broadleaved swamp woodland not on acid peat; G1.5 Broadleaved swamp woodland
on acid peat; G1.7 Thermophilous deciduous woodland; G1.8 Acidophilous Quercusdominated woodland; G1.9 Non-riverine woodland with Betula, Populus tremula or Sorbus
aucuparia; G1.B Non-riverine Alnus woodland; G3.4 Pinus sylvestris woodland south of
the taiga; G4.4 Mixed Pinus sylvestris – Betula woodland; G4.7 Mixed Pinus sylvestris –
acidophilous Quercus woodland.
Land use: forestry – minor, nature conservation and research – 71%.
Protected areas: overlaps (about 26400 ha) with Prypiat-Stokhid naional nature park;
overlaps (about 11300 ha) with Prypiat-Stokhid regional landscape park (Rivne region);
overlaps (about 23400 ha) with Ramsar site “Prypiat river loodplains”, includes Birkivskyi
regional hydrological reserve (850 ha), Hirkivskyi regional hydrological reserve (400
ha), Zalukhivskyi regional hydrological reserve (839 ha), Velykohlushanskyi regional
hydrological reserve (360 ha), Vetlivskyi regional hydrological reserve (850 ha), Nobelskyi
regional ornithological reserve (510 ha), Prypiatski regional ichthyological reserve (3155
ha), regional hydrological reserve “Prypiatskyi–1” (340 ha), regional hydrological reserve
“Prypiatskyi–2” (220 ha), regional hydrological reserve “Prypiatskyi–3” (320 ha), Tsyrskyi
regional hydrological reserve (210 ha), overlaps (about 26817 ha) with Emerald Site “PrypiatStokhid Naional Nature Park”, overlaps (about 9664 ha) with Emerald Site “ Stokhid-Nobel”.
Threats: –.
General description.
The loodplain of the Prypiat river with fragments of a higher terrace. The area includes
lakes Liubiaz (430 ha), Nobel (470 ha), Volianske (405 ha). Eutrophic sedge mires occupy the
largest area. Main dominants are Carex appropinquata and Carex elata, other typical species
are Potenilla palustris, Lythrum salicaria, Thelypteris palustris, Iris pseudacorus. Large areas
are covered by Alnus gluinosa and Salix cinerea swamps. There occur poor fens with Carex
nigra, Carex panicea, Carex lava, Agrosis canina. Floodplain meadows are dominated
mainly by Deschampsia cespitosa, Molinia caerulea, someimes by Agrosis capillaris,
Poa pratensis, Agrosis stolonifera. The most typical dominants of aquaic vegetaion are
Straiotes aloides, Potamogeton nodosus, Potamogeton lucens. Main dominants of the
litoral vegetaion are Phragmites australis, Typha angusifolia, Glyceria maxima. Pinus
sylvestris and Quercus robur forests on the sand terrace are dominated by Vaccinium
myrillus, Pteridium aquilinum, Molinia caerulea. There are several areas od sand dunes.
There are complexes of unvegetated sands with communiies of Corynephorus canescens
(predominanly), Calamagrosis epigeios, Koeleria glauca, Filago minima, Polytrichum
piliferum with presence of Centaurea stoebe, Helichrysum arenarium, Thymus serpyllum.
Botanical significance. The IPA includes the largest eutrophic sedge mire in Ukraine,
important for conservaion of Aldrovanda vesiculosa.
Criterion A
• Aldrovanda vesiculosa L.; A(i), A(ii); abundance: occasional; trend: unknown; species
data quality: medium; trend data quality: poor.
Criterion C
• C1.223 Floating Stratiotes aloides rafts; area: 50 ha; trend: stable; area data quality:
poor; trend data quality: poor.
• C1.226 Floating Aldrovanda vesiculosa communities; area: <0.1 ha; trend: unknown;
area data quality: poor; trend data quality: poor.
• D5.2 Beds of large sedges normally without free-standing water; area: 15000 ha; trend:
stable; area data medium: poor; trend data quality: poor.
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•
E3.4 Moist or wet eutropic and mesotrophic grassland; area: 5000 ha; trend: stable;
area data quality: medium; trend data quality: medium.
• E3.5 Moist or wet oligotrophic grassland; area: 1000 ha; trend: stable; area data quality:
poor; trend data quality: poor.
• X35 Inland sand dunes; area: 38 ha; trend: stable; area data quality: medium; trend data
quality: medium.
Literature
1. Андрієнко Т.Л., Прядко О.І. Флористичне та ценотичне різноманіття проектованого
національного природного парку “Прип’ять-Стохід” // Науковий вісник Волинського
державного університету імені Лесі Українки. І Міжнародна науково-практична
конференція “Шацький національний природний парк: регіональні аспекти, шляхи та
напрями розвитку”. – 2007. – 11, ч. 2. – С. 132–140.
2. Андрієнко Т.Л., Прядко О.І. НПП Прип’ять-Стохід // Фіторізноманіття заповідників
і національних природних парків України. Ч. 2. Національні природні парки / під ред.
В.А. Онищенка і Т.Л. Андрієнко. – Київ: Фітосоціоцентр, 2012. – С. 429–439.
3. Андрієнко Т.Л., Прядко О.І., Арап Р.Я., Коніщук М.О. Національний природний парк
“Прип’ять-Стохід”. Рослинний світ / Під заг. ред. Т.Л.Андрієнко. – К.: Фітосоціоцентр,
2009. – 86 с.
4. Прядко О.І. Рослинний покрив водно-болотних угідь РЛП “Прип’ять-Стохід” //
Сучасний стан водно-болотних угідь регіонального ландшафтного парку “Прип’ятьСтохід” та їх біорізноманіття. – К:, 2001 – С. 70–75.
Zaplava Siverskoho Dintsia
V.A. Onyshchenko
Ukrainian name: Заплава Сіверського Дінця.
Transliteraion/Translaion variants: Floodplain of the Siverskyi Donets’ River.
Area: 16679 ha.
Altitude: 49–70 m.
Latitude: 48°54’07N (48.9020°).
Longitude: 37°50’05” E (37.8347°).
Administrative regions. Donetsk region: Sloviansk raion, Lyman (Krasnyi Lyman) raion;
Luhansk region: Kreminna raion.
Ownership: state.
Biogeographic regions: steppic
Habitats. Level 1. C – 6.8%; D – 1.9%; E – 20.9%; G – 69.9%; H – 0.5%.
Habitats. Level 2. C1 Surface standing waters – 2.6%; C2 Surface running waters – 3.6%;
C3 Littoral zone of inland surface waterbodies – 0.6%; D5 Sedge and reedbeds, normally
without free-standing water – 1.9%; E2 Mesic grasslands – 19.0%; E3 Seasonally wet and
wet grasslands – 1.9%; G1 Broadleaved deciduous woodland – 68.6%; G3 Coniferous
woodland – 1.2%; G4 Mixed deciduous and coniferous woodland – 0.1%; H5 Miscellaneous
inland habitats with very sparse or no vegetation – 0.5%.
Futher habitat description. C1.2 Permanent mesotrophic lakes, ponds and pools; C2.3
Permanent non-tidal, smooth-flowing watercourses; E3.4 Moist or wet eutrophic and
mesotrophic grassland; G1.2 Mixed riparian floodplain and gallery woodland; G1.3
Mediterranean riparian woodland; G1.4 Broadleaved swamp woodland not on acid
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peat; G3.4 Pinus sylvestris woodland south of the taiga; G4.C Mixed Pinus sylvestris –
thermophilous Quercus woodland; H5.6 Trampled areas.
Land use: agriculture (animals) – 10%; mowing/hay making – 25%; forestry – 70%; nature
quality: medium; trend data quality: medium.
Literature
1. Бурда Р.І., Остапко В.М., Глухов А.З., Шпилева Н.В. Національний природний
парк “Святі Гори”: біологічна різноманітність рослинного покриву // Заповідна справа
в Україні. – 1997. – Т. 3, вип.1. – С.10–17.
2. Морозова І. Міксоміцети Національного природного пароку “Святі Гори” // Матли ІV Міжн. наук. конф. студентів та аспірантів “Молодь і поступ біології” (м. Львів, 7–10
квітня 2008 р.). – Львів: ЛНУ ім. І. Франка, 2008. – С.106–107.
3. Надєіна О.В. Лишайники національного природного парку “Святі Гори” //
Чорноморський ботан. журн. – 2007. – 3, № 2. – С. 100–108.
4. Остапко В.М., Шпилева Н.В., Дьякова О.В. НПП Святі гори // Фіторізноманіття заповідників і національних природних парків України. Ч. 2. Національні природні парки
/ під ред. В.А. Онищенка і Т.Л. Андрієнко. – Київ: Фітосоціоцентр, 2012. – С. 440–456.
Zaplava Tysy
conservaion and research – 34%.
Protected areas: overlaps (about 7403 ha) with Sviai Hory naional nature park and Emerald
Site “Sviai Hory “, overlaps with Emerald Site “Kreminski Lisy” (5252 ha).
Threats: agricultural intensiicaion/expansion (general) – low; development (recreaion/
tourism) – low; forestry (intensified forest management) – low.
General description. The loodplain of the Siverskyi Donets river. Its major vegetaion
is represented by broadleaved forests and meadows. The largest area is occupied
by Quercus robur forests with co-domimanance of Tilia cordata, Populus tremula,
Fraxinus excelsior. Other typical species are Euonymus europaeus, E. verrucosus, Corylus
avellana, Crataegus avellana, Acer tataricum, Aegopodium podagraria (dominant),
Convallaria majalis (dominant), Stellaria holostea (dominant), Anemone ranunculoides,
Corydalis solida, Ficaria verna, Glechoma hederacea, Lysimachia nummularia,
Pulmonaria obscura, Scilla siberica, Tulipa quercetorum, Viola odorata, Viola mirabilis.
Besides there occur Alnus gluinosa and Populus alba forests. Floodplain meadows
are dominated by Festuca pratensis, Elytrigia repens, Koeleria delavignei, Phleum
pratense, Poa pratensis. Dominants of eutrophic mires are Phragmites australis (major),
Carex acuta, Carex acutiformis, Carex riparia, Carex vesicaria. Dominants of litoral
vegetaion: Phragmites australis, Acorus calamus, Schoenoplectus lacustris, Typha
angustifolia, Typha latifolia. Dominants of aquaic vegetaion: Lemna minor, Potamogeton
natans, Nuphar lutea, Nymphaea alba, Salvinia natans.
Botanical significance. Important for conservaion of Quercus robur loodplain forests, black
alder woods and loodplain meadows.
Criterion C
• E2.2 Low and medium alitude hay meadows; area: 2700 ha; trend: stable; area data
quality: good; trend data quality: medium.
• G1.22 Mixed Quercus – Ulmus – Fraxinus woodland of great rivers; area: 9000 ha; trend:
stable; area data quality: medium; trend data quality: medium.
• G1.3 Mediterranean riparian woodland; area: 140 ha; trend: unknown; area data
quality: poor; trend data quality: poor.
• G1.414 Steppe swamp Alnus gluinosa woods; area: 700 ha; trend: stable; area data
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R.Ya. Kish, V.A. Onyshchenko
Ukrainian name: Заплава Тиси.
Transliteraion/Translaion variants: Floodplain of the Tysa (Tisza) River.
Area: 7930 ha.
Altitude: 115–204 m.
Latitude: 48°11’03N (48.1841°).
Longitude: 23°10’57” E (23.1823°).
Administrative regions. Zakarpatska region: Khust city, Khust raion, Tiachiv raion, Vynohradiv
raion.
Ownership: state.
Biogeographic regions: pannonian.
Habitats. Level 1. C – 20.0%; D – 3.0%; E – 16.0%; F – 3.0%; G – 57.0%; H – 1.0%.
Habitats. Level 2. C1 Surface standing waters – 0.3%; C2 Surface running waters – 7.7%;
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C3 Littoral zone of inland surface waterbodies – 12.0%; D5 Sedge and reedbeds, normally
without free-standing water – 3.0%; E2 Mesic grasslands – 2.0%; E3 Seasonally wet and
wet grasslands – 14.0%; F9 Riverine and fen scrubs – 3.0%; G1 Broadleaved deciduous
woodland – 57.0%; H5 Miscellaneous inland habitats with very sparse or no vegetation –
1.0%.
Futher habitat description. C1.2 Permanent mesotrophic lakes, ponds and pools; C1.3
Permanent eutrophic lakes, ponds and pools; C2.2 Permanent non-tidal, fast, turbulent
watercourses; C2.3 Permanent non-tidal, smooth-flowing watercourses; C3.6 Unvegetated
or sparsely vegetated shores with soft or mobile sediments; E3.4 Moist or wet eutrophic
and mesotrophic grassland; G1.1 Riparian and gallery woodland, with dominant Alnus,
Betula, Populus or Salix; G1.2 Mixed riparian floodplain and gallery woodland; G1.4
Broadleaved swamp woodland not on acid peat; H5.6 Trampled areas; G1.A Meso- and
eutrophic Quercus, Carpinus, Fraxinus, Acer, Tilia, Ulmus and related woodland.
Land use: agriculture (animals) – minor; forestry – minor; mowing/hay making – minor;
nature conservaion and research – minor; tourism/recreaion – minor.
Protected areas: overlaps (about 4596 ha) with Prytysianskyi regional landscape park,
includes Ehresh regional botanical reserve (37.4 ha), overlaps (4669 ha) with Emerald Site
“Vynohradivska Tysa”.
Threats: construcion/impact of dike/dam/barrage – low; eutrophicaion – low.
General description. Floodplain of the Tysa river. Vegetaion is represented by Salix alba and
Populus nigra forests, Salix shrubs, moist and mesic grasslands. Signiicant parts of total area
are running waters and unvegetated alluvial deposits (pebble, gravel). Besides there are
standing waters, Alnus gluinosa swamps, sedge beds. Water is predominantly eutrophic.
Botanical significance. Important for loodplain forests, shrubs, aquaic and ephemeral
litoral vegetaion.
Criterion C
• C2.34 : Eutrophic vegetaion of slow-lowing rivers; area: 5 ha; trend: unknown; area
data quality: poor; trend data quality: poor.
• C2.28 : Eutrophic vegetaion of fast-lowing streams; area: 3 ha; trend: unknown; area
data quality: poor; trend data quality: poor.
• C3.4 Species-poor beds of low-growing water-fringing or amphibious vegetaion; area:
10 ha; trend: unknown; area data quality: poor; trend data quality: poor.
• C3.51 Euro-siberian dwarf annual amphibious swards (but excluding C3.5131 Toad-rush
swards); area: 10 ha; trend: unknown; area data quality: poor; trend data quality: poor.
• E5.4 Moist or wet tall-herb and fern fringes and meadows; area: 50 ha; trend: unknown;
area data quality: poor; trend data quality: poor.
• G1.11 Riverine Salix woodland; area: 600 ha; trend: stable; area data quality: poor;
trend data quality: poor.
• G1.3 Mediterranean riparian woodland; area: 3000 ha; trend: unknown; area data
quality: poor; trend data quality: poor.
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Zaplavni Lisy na Borzhavi
R.Ya. Kish, V.A. Onyshchenko
Ukrainian name: Заплавні ліси на Боржаві.
Transliteraion/Translaion variants: Floodplain Forests on the Borzhava.
Area: 2775 ha.
Altitude: 113–130 m.
Latitude: 48°13’00” N (48.2166°).
Longitude: 22°48’13” E (22.8034°).
Administrative regions. Zakarpathia region: Berehove raion, Vynohradiv raion.
Ownership: state.
Biogeographic regions: pannonian.
Habitats. Level 1. C – 0.3%; D – 0.2%; E – 1.8%; F – 0.0%; G – 97.0%; H – 0.7%.
Habitats. Level 2. C1 Surface standing waters – 0.2%; C2 Surface running waters – 0.1%;
C3 Littoral zone of inland surface waterbodies – 0.0%; D5 Sedge and reedbeds, normally
without free-standing water – 0.2%; E2 Mesic grasslands – 0.1%; E3 Seasonally wet and wet
grasslands – 1.7%; F9 Riverine and fen scrubs – 0.0%; G1 Broadleaved deciduous woodland
– 97.0%; H5 Miscellaneous inland habitats with very sparse or no vegetation – 0.7%.
Futher habitat description. F9.2 Salix carr and fen scrub; G1.1 Riparian and gallery
woodland, with dominant Alnus, Betula, Populus or Salix; G1.2 Mixed riparian floodplain
and gallery woodland; G1.4 Broadleaved swamp woodland not on acid peat; G1.A Mesoand eutrophic Quercus, Carpinus, Fraxinus, Acer, Tilia, Ulmus and related woodland; H5.6
Trampled areas.
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Land use: forestry – major, nature conservation and research – minor.
Protected areas: overlaps (2506 ha) with Prytysianskyi regional landscape park, includes
regional preserve (“zapovidne urochyshche”) “Borzhava” (153 ha, included in the regional
landscape park), includes regional botanical reserve “Velykyi Lis” (1.5 ha, included in
regional preserve “Borzhava”), overlaps (2152 ha) with Emerald Site “Ponyzzia Borzhavy”.
Threats: forestry (intensiied forest management) – low.
General description. Major vegetaion types are moist loodplain Quercus robur - Fraxinus
pannonicus forest and mesic Quercus robur - Carpinus betulus forest. Dominants of the herb
layer of the moist forests are Filipendula ulmaria, Glechoma hederacea, Impaiens nolitangere, Phalaroides arundinacea, Rubus caesius, Urica dioica. In the mesic forests, there
prevail Anemone nemorosa, Aegopodium podagraria, Hedera helix, Lamium galeobdolon.
Besides there are Alnus gluinosa wet forests and Alnus gluinosa swamps, small areas of
Salix alba and Salix fragilis woods, sedge and reedbeds, Salix cinerea swamps, meadows.
Botanical significance. One of the best riverine forests in Ukraine and Central Europe.
Criterion C
• G1.22 Mixed Quercus – Ulmus – Fraxinus woodland of great rivers; area: 1400 ha; trend:
stable; area data quality: medium; trend data quality: medium
• G1.414 Steppe swamp Alnus gluinosa woods; area: 40 ha; trend: stable; area data
quality: medium; trend data quality: medium.
Conservaion proposals. Stop clear felling and create a state botanical reserve including the
enire IPA.
pioneer and ephemeral vegetation; F9.1 Riverine scrub; F9.2 Salix carr and fen scrub; G1.1
Riparian and gallery woodland, with dominant Alnus, Betula, Populus or Salix; G1.2 Mixed
riparian floodplain and gallery woodland; G1.4 Broadleaved swamp woodland not on acid
peat; G3.4 Pinus sylvestris woodland south of the taiga.
Zaplavy Dnipra i Sozha
V.A. Onyshchenko, O.V. Lukash
Ukrainian name: Заплави Дніпра і Сожа.
Transliteraion/Translaion variants: Floodplain of the Dnipro and the Sozh.
Area: 34967 ha.
Altitude: 104–120 m.
Latitude: 51°43’50” N (51.7304°).
Longitude: 30°39’05” E (30.6515°).
Administrative regions. Chernihiv region: Chernihiv raion, Ripky raion.
Ownership: state, private.
Biogeographic regions: continental
Habitats. Level 1. C – 11%; D – 15%; E – 44%; F – 8%; G – 22%.
Habitats. Level 2. C1 Surface standing waters – 3.0%; C2 Surface running waters – 7.2%;
C3 Littoral zone of inland surface waterbodies – 0.8%; D5 Sedge and reedbeds, normally
without free-standing water – 15%; E3 Seasonally wet and wet grasslands – 44%; F9 Riverine
and fen scrubs – 8%; G1 Broadleaved deciduous woodland – 18%; G3 Coniferous woodland
– 4%.
Futher habitat description. C1.2 Permanent mesotrophic lakes, ponds and pools; C1.6
Temporary lakes, ponds and pools; C2.3 Permanent non-tidal, smooth-flowing watercourses;
C3.2 Water-fringing reedbeds and tall helophytes other than canes; C3.4 Species-poor beds
of low-growing water-fringing or amphibious vegetation; C3.6 Unvegetated or sparsely
vegetated shores with soft or mobile sediments; D5.1 Reedbeds normally without freestanding water; D5.2 Beds of large sedges normally without free-standing water; E3.4
Moist or wet eutrophic and mesotrophic grassland; C3.5 Periodically inundated shores with
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Land use: agriculture (animals) – 10%; agriculture (arable) – 1%; forestry – 20%; mowing /
hay making – 30%; nature conservation and research – minor%.
Protected areas: includes regional hydrological reserve “Kryvi Hriady” (129.2 ha), includes
regional zoological reserve “Kryvi Hriady – 2” (7 ha), includes regional hydrological reserve
“Ozera Sympol, Sviate ta Prylehli Bolota” (140 ha), includes regional hydrological reserve
“Urochyshche Dor” (166 ha), overlaps (6679 ha) with Emerald Site “Ripkynskyi”, overlaps
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(6406 ha) with Emerald Site “Liubetskyi”, overlaps (3840 ha) with Emerald Site “Pakulskyi”,
overlaps with proposed Liubechskyi national nature park, overlaps with proposed
Dniprovskyi naional nature park.
Threats: abandonment/reduction of land management – low, agricultural intensification/
expansion (general) – low; burning of vegetation – low, development (urbanization) – low.
General description. Ukrainian part of the floodplains of the Dnipro and Sozh rivers length of
104 km. Western boundary runs along the state border between Ukraine and Belarus. This is
one of the largest floodplains with natural water regime in Ukraine. Floods are regular. The
riverbed has natural meanders. There are many oxbows and lakes. Meadows occupy the
largest area. Sandy meadows on high elements of relief are dominated by Calamagrostis
epigeios, Agrostis vinealis, Poa angustifolia. Dominants of meadows of a lower level are
Alopecurus pratensis, Poa pratensis, Festuca pratensis, Festuca rubra, Agrostis gigantea.
Moist meadows are dominated by Deschampsia cespitosa and Phalaroides arundinacea.
Mires and littoral vegetation are dominated by Phragmites auistralis, Glyceria maxima,
Carex acuta, C. elata, Carex juncella, C. appropinquata. There are forests (Quercus robur,
Alnus glutinosa, Salix alba, Pinus sylvestris) and shrubs (Salix cinerea, Salix triandra, Salix
acutifolia). Main dominants of aquatic vegetation are Lemna minor, Spirodela polyrrhiza,
Sagittaria sagittifolia, Hydrocharis morsus-ranae, Nuphar lutea, Nymphaea alba,
Potamogeton perfoliatus, Potamogeton lucens, Stratiotes aloides.
Botanical significance. This area is important for conservation of floodplain complexes in
Ukraine.
Criterion C
• C1.223 Floating Stratiotes aloides rafts; area: 6 ha; trend: stable; area data quality: poor;
trend data quality: poor.
• C1.224 Floating Utricularia australis and Utricularia vulgaris colonies; area: 4 ha; trend:
stable; area data quality: poor; trend data quality: poor
• C1.3411 Ranunculus communities in shallow water; area: 0.2 ha; trend: stable; area data
quality: poor; trend data quality: poor.
• C2.33 Mesotrophic vegetation of slow-flowing rivers; area: 80 ha; trend: stable; area
data quality: poor; trend data quality: poor.
• C3.4 Species-poor beds of low-growing water-fringing or amphibious vegetation; area: 6
ha; trend: stable; area data quality: poor; trend data quality: poor.
• C3.51 Euro-Siberian dwarf annual amphibious swards; area: 6 ha; trend: stable; area
data quality: poor; trend data quality: poor.
• D5.2 Beds of large sedges normally without free-standing water; area: 5000 ha; trend:
stable; area data quality: poor; trend data quality: poor.
• E2.2 Low and medium alitude hay meadows; area: 5000 ha; trend: stable; area data
quality: medium; trend data quality: medium.
• E3.4 Moist or wet eutropic and mesotrophic grassland; area: 10000 ha; trend: stable;
area data quality: medium; trend data quality: medium.
• E3.5 Moist or wet oligotrophic grassland; area: 100 ha; trend: stable; area data quality:
poor; trend data quality: poor.
• E5.4 Moist or wet tall-herb and fern fringes and meadows; area: 6 ha; trend: stable; area
data quality: poor; trend data quality: poor.
• F9.1 Riverine scrub; area: 5000 ha; trend: stable; area data quality: poor; trend data
quality: poor.
• G1.11 Riverine Salix woodland; area: 1000 ha; trend: stable; area data quality: poor;
trend data quality: poor.
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•
G1.21 Riverine Fraxinus – Alnus woodland, wet at high but not at low water; area: 150
ha; trend: stable; area data quality: poor; trend data quality: poor.
Literature
1. Андриенко Т.Л., Шеляг-Сосонко Ю.Р. Растительный мир Украинского Полесья в
аспекте его охраны. – Киев: Наук. думка, 1983. – 216 с.
2. Водно-болотні угіддя України / Під ред. Г.Б. Марушевського, І.С. Жарук. – К.:
Wetlands International Black Sea Programme, 2006. – 312 с.
3. Лукаш А.В., Андриенко Т.Л. Ботанически ценные охраняемые природные
территории Полесья. – Чернигов: Десна Полиграф, 2014. – 104 с.
4. Перспективная сеть заповедных объектов Украины / под ред. Ю.Р.ШелягСосонко. – К.: Наук. думка, 1987. – 292 с.
5. Смарагдова мережа в Україні / Колектив авторів під ред. Проценка Л.Д. – Київ:
Хімджест, 2011. – 192 с.
Zelena Balka
V.A. Onyshchenko
Ukrainian name: Зелена Балка.
Transliteraion/Translaion variants: Green Ravine.
Area: 1339.0 ha.
Altitude: 40–104 m.
Latitude: 47°47’29” N (47.7914°).
Longitude: 33°11’30” E (33.1915°).
Administrative regions. Dnipropetrovsk region: Shyroke raion.
Ownership: state.
Biogeographic regions: steppic.
Habitats. Level 1. C – 3.8%; D – 0.5%; E – 93.7%; F – 0.1%; G – 0.9%; H – 1.0%.
Habitats. Level 2. C1 Surface standing waters – 3.4%; C3 Littoral zone of inland surface
waterbodies – 0.4%; E1 Dry grasslands – 92.1%; E2 Mesic grasslands – 0.5%; E5 Woodland
fringes and clearings and tall forb stands – 0.1%; E7 Sparsely wooded grasslands – 1.0%;
F3 Temperate and mediterranean-montane scrub – 0.1%; G1 Broadleaved deciduous
woodland – 0.9%; H2 Screes – 0.0%; H3 Inland cliffs, rock pavements and outcrops – 0.0%;
H5 Miscellaneous inland habitats with very sparse or no vegetation – 1.0%.
Futher habitat description. C1.2 Permanent mesotrophic lakes, ponds and pools; C3.2
Water-fringing reedbeds and tall helophytes other than canes; D5.1 Reedbeds normally
without free-standing water; E1.1 Inland sand and rock with open vegetation; E1.2
Perennial calcareous grassland and basic steppes; F3.1 Temperate thickets and scrub;
F3.2 Submediterranean deciduous thickets and brushes; G1.C Highly ariicial broadleaved
deciduous forestry plantaions; H2.6 Calcareous and ultra-basic screes of warm exposures;
H3.2 Basic and ultra-basic inland clifs; H5.6 Trampled areas.
Land use: agriculture (animals) – major.
Protected areas: –.
Threats: –.
General description. Grasslands dominated by Festuca valesiaca, Sipa capillata, S.
lessingiana, Botriochloa ischaemum, Koeleria cristata, Sipa ucrainica (rarely) with presence
of Teucrium polium, Euphorbia seguierana, Euphorbia stepposa, Salvia nutans, Eryngium
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campestre, Marrubium praecox, Veronica barellieri, Artemisia austriaca, Medicago falcata. In
more mesic habitats, there prevail Poa angusifolia, Bromopsis inermis, Elytrigia intermedia,
Elytrigia trichophora, Carex praecox, frequent species are Achillea pannonica, Marrubium
Criterion A
• Chamaecyisus graniicus (Rehmann.) Rothm.; A(iii); abundance: occasional; trend:
stable; species data quality: medium; trend data quality: medium.
• Tulipa hypanica Klokov et Zoz; A(iii); abundance: occasional; trend: stable; species data
quality: medium; trend data quality: poor.
Criterion C
• E1.2 Perennial calcareous grassland and basic steppes; area: 1000 ha; trend: stable;
area data quality: medium; trend data quality: good.
Conservaion proposals. Create a state reserve and an Emerald Site..
Literature
1. Красова О.О., Сметана О.М. Матеріали до оцінки перспективних степових
компонентів екомережі Кривбасу (“Балка Зелена”) // Чорноморський ботанічний
журнал. – 2012. – 8, № 4. – С. 459–470.
Zholoby
H.I. Oliiar, H.P. Protsiv
Ukrainian name: Жолоби.
Area: 60.0 ha.
Altitude: 308–387 m.
Latitude: 49°29’04” N (49.4845°).
Longitude: 24°53’20” E (24.8888°).
praecox, Galium ruthenicum, Plantago urvillei. In shrub steppes, Amygdalus nana,
Caragana frutex, Caragana scythica predominate. On stony calcareous soil, there prevail
Jurinea brachycephala, Teucrium chamaedrys, Potenilla incana, Sipa asperella, Galatella
villosa, Elytrigia sipifolia, Chamaecyisus graniicus with presence of Alyssum tortuosum
(Odontarrhena tortuosa), Cephalaria uralensis, Convolvulus lineatus. Communiies of
Elytrigia repens, Calamagrosis epigeios, Festuca regeliana, Galega oicinalis, Puccinellia
distans occupy botoms of the valleys. In weter habitats, dominant species are Phragmites
australis, Typha angusifolia, Schoenoplectus tabaernaemontani, Bolboschoenus mariimus,
Juncus gerardii, Agrosis stolonifera. In the most saline habitats, there are communiies of
Limonium alutaceum and Artemisia santonica. Typical lichens of outcrops are Caloplaca
aurania, C. saxicolsa, C. variabilis, Candellariella aurella, Lecanora dispersa, Verrucaria
nigrescens. Communiies of high shrubs are represented by thickets of Crataegus fallacina,
Prunus stepposa.
Botanical significance. One of the best steppe areas west to the Dnipro river.
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Administrative regions. Ternopil region: Berezhany raion.
Ownership: state.
Biogeographic regions: coninental.
Habitats. Level 1. E – 25.0%; F – 10.0%; G – 64.0%; H – 1.0%.
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Habitats. Level 2. E1 Dry grasslands – 22.0%; E7 Sparsely wooded grasslands – 3.0%;
F3 Temperate and mediterranean-montane scrub – 10.0%; G1 Broadleaved deciduous
woodland – 12.9%; G3 Coniferous woodland – 52.1%; H5 Miscellaneous inland habitats
with very sparse or no vegetation – 1.0%.
Futher habitat descripion. E1.2 Perennial calcareous grassland and basic steppes; F3.1
Temperate thickets and scrub; F3.2 Submediterranean deciduous thickets and brushes;
G3.F Highly artificial coniferous plantations; H5.6 Trampled areas.
Land use: forestry (major), mowing/hay making – minor.
Protected areas: overlaps (60 ha) with Emerald Site “Berezhanske Opillia”, same as proposed
state botanical reserve “Zholoby”.
Threats: abandonment/reducion of land management – high, forestry (afforestation) – low.
General description. Semi-dry grasslands, plantaions of Pinus sylvestris, natural deciduous
woods and shrubs (Prunus spinosa, Crataegus spp.). Grasslands are dominated by
Brachypodium pinnatum, Calamagrosis epigeios, Carex humilis. Other frequent species:
Centaurea stricta, Filipendula vulgaris, Teucrium chamaedrys.
Botanical significance. Large populaion of Orchis militaris.
Criterion C
• E1.2 Perennial calcareous grassland and basic steppes (6210, *Important orchid site);
area: 13 ha; trend: decreasing; area data quality: good; trend data quality: good. Size
of populaion of Orchis militaris: 2000–4000 individuals.
Conservaion proposals: remove all pine trees, do not allow expansion of trees and shrubs,
mow grasslands, non-intensive grazing.
Literature
1. Оліяр Г.І., Проців Г.П. Флористичні особливості деяких урочищ
Бережанського Опілля // Науковий вісник Національного університету біоресурсів і
природокористування України. Серія “Лісівництво та декоративне садівництво” – К.:
ВЦ НУБіП України, 2012. – Вип. 171, ч. 1. – С. 131–134.
Fraxinus, Acer, Tilia, Ulmus and related woodland; H2.6 Calcareous and ultra-basic screes of
warm exposures; H3.2 Basic and ultra-basic inland cliffs.
Zhyzhava
V.A. Onyshchenko
Ukrainian name: Жижава.
Area: 107.0 ha.
Altitude: 146–300 m.
Latitude: 48°41’54” N (48.6983°).
Longitude: 25°39’11” E (25.6531°).
Administrative regions. Ternopil region: Zalishchyky raion.
Ownership: state.
Biogeographic regions: coninental.
Habitats. Level 1. E – 36.0%; F – 13.0%; G – 40.0%; H – 11.0%.
Habitats. Level 2. E1 Dry grasslands – 35.0%; E2 Mesic grasslands – 1.0%; F3 Temperate and
mediterranean-montane scrub – 13.0%; G1 Broadleaved deciduous woodland – 40.0%; H2
Screes – 2.0%; H3 Inland clifs, rock pavements and outcrops – 9.0%.
Futher habitat descripion. E1.1 Inland sand and rock with open vegetation; E1.2
Perennial calcareous grassland and basic steppes; F3.1 Temperate thickets and scrub;
G1.7 Thermophilous deciduous woodland; G1.A Meso- and eutrophic Quercus, Carpinus,
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Land use: agriculture (animals) – minor, forestry – minor, nature conservation and research
– major, tourism/recreaion – minor.
Protected areas: overlaps (37 ha) with Zhyzhavskyi state botanical reserve, included in
naional nature park “Dnistrovskyi Kanion”, included in Emeral Site “Dnistrovskyi Kanion
Naional Nature Park”.
Threats: agricultural intensification/expansion (grazing) – low, development (recreaion/
tourism) – low.
General description. A steep slope of the Dnister valley with broadleaved forest, steppe
vegetaion, shrubs, calcareous rocks and screes. The tree layer of the forest consists of
Carpinus betulus, Quercus robur, Quercus petraea, Fraxinus excelsior. The shrub vegetaion
is dominated by Prunus spinosa, Staphylea pennata, Crataegus sp. Dominants of the steppe
are Carex humilis, Festuca valesiaca, Sipa capillata. There are a lot of protected species:
Chamaecyisus podolicus, Iris hungarica, Pulsailla grandis, Pulsailla pratensis, Spiraea
369
polonica etc.
Botanical significance. Important for narrow endemic species Spiraea polonica (locus
classicus), Podolian steppes, shrubs and rocks with many rare species.
Criterion A
• Spiraea polonica Błocki; A(iii); abundance: occasional (about 200 individuals in one
compact populaion); trend: stable; species data quality: good; trend data quality:
medium.
Criterion C
• E1.11 Euro-Siberian rock debris swards; area: 12 ha; trend: stable; area data quality:
poor; trend data quality: good.
• F3.247 Ponto-Sarmatic deciduous thickets; area: 14 ha; trend: stable; area data quality:
medium; trend data quality: medium.
• H2.6 Calcareous and ultra-basic screes of warm exposures; area: 0.5 ha; trend: stable;
area data quality: poor; trend data quality: good.
Literature
1. Федорончук М.М., Белемець Н.М., Волуца О.Д. Рідкісні види роду Spiraea L.
(Rosaceae) флори України та стан їхньої України // Укр. ботан. журн. – 2013. – 70, № 2. –
С. 164–167.
2. Червона книга України. Рослинний світ / за ред. Я.П. Дідуха. – К.: Глобалконсалтинг,
2009. – 900 с.
Zolochivka
O.T. Kuzyarin, V.A.Onyshchenko
Ukrainian name: Золочівка.
Area: 398.0 ha.
Altitude: 265–280 m.
Latitude: 49°46’43” N (49.7786°).
Longitude: 24°58’20” E (24.9721°).
Administrative regions. Lviv region: Zolochiv raion.
Ownership: state (major), private
Biogeographic regions: coninental.
Habitats. Level 1. C – 0.7%; D – 50.0%; E – 40.3%; F – 1.0%; G – 8.0%.
Habitats. Level 2. C1 Surface standing waters – 0.4%; C3 Littoral zone of inland surface
waterbodies – 0.3%; D4 Base-rich fens and calcareous spring mires – 30.3%; D5 Sedge
and reedbeds, normally without free-standing water – 10.0%; E2 Mesic grasslands – 4.3%;
E3 Seasonally wet and wet grasslands – 36.0%; F9 Riverine and fen scrubs – 1.0%; G1
Broadleaved deciduous woodland – 8.0%.
Futher habitat descripion. D4.1 Rich fens, including eutrophic tall-herb fens and calcareous
flushes and soaks; D5.1 Reedbeds normally without free-standing water; D5.2 Beds of
large sedges normally without free-standing water; E2.2 Low and medium altitude hay
meadows; E3.4 Moist or wet eutrophic and mesotrophic grassland; F9.2 Salix carr and fen
scrub.
Land use: mowing/hay making – minor.
Protected areas: –.
Threats: abandonment/reducion of land management – low, agricultural intensiicaion/
370
expanion (general) – low, water (drainage) – low.
General description. Parially drained fen. Major vegetaion types are rich fens and
wet grasslands. Dominant species of rich fen are Carex davalliana, Carex lacca, Carex
hosiana, Carex panicea, Molinia caerulea, Pragmites australis, Schoenus ferrugineus,
Sesleria caerulea, Calliergonella cuspidata, Campylium stellatum, Limprychia revolvens.
Other frequent species are Epipacis palustris, Eupatorium cannabinum, Lythrum salicaria,
Parnassia palustris, Pinguicula vulgaris, Potenilla erecta, Rinanthus seroinus, Sanguisorba
oicinalis, Toieldia calyculata, Valeriana simplicifolia. Meadows are dominated by Briza
media, Deschampsia cespitosa, Festuca rubra, Molinia caerulea. Besides there are marshes
dominated by Carex acuta and other large sedges without signiicant presence of calciphile
species.
Botanical significance. One of the largest rich fens in Ukraine with rare species (Sesleria
caerulea, Juncus subnodulosus, Dactylorhiza ochroleuca, Dactylorhiza traunsteineri).
Criterion C
• D4.1 Rich fens, including eutrophic tall-herb fens and calcareous flushes and soaks;
area: 140 ha; trend: stable; area data quality: poor; trend data quality: medium.
Conservaion proposals. Do not allow recovery of reclamaion ditches, include in Pivnichne
Podillia naional nature park and Emerald Site “Pivnichne Podillia”.
Literature
1. Кузярін О.Т. Перспективні природоохоронні території басейну верхів’я Західного
Бугу // Наукові записки державного природохнавчого музею. – Львів. – 2012. – Вип. 28.
– С. 121–130.
2. Червона книга України. Рослинний світ / за ред. Я.П. Дідуха. – К.: Глобалконсалтинг,
2009. – 900 с.
371
Contents
Selection criteria ............................................................................................................ 3
Descriptions of IPAs ...................................................................................................... 26
Aiu-Dah (V.A. Onyshchenko) ...................................................................................... 27
Ak-Monaiskyi Step (V.P. Kolomiychuk, V.A. Onyshchenko) ......................................... 29
Arabatska Strilka (V.A. Onyshchenko) ....................................................................... 31
Ardov (R.Ya. Kish) ....................................................................................................... 33
Askania-Nova (V.A. Onyshchenko) .............................................................................. 35
Balakyrivka (V.A. Onyshchenko) ................................................................................. 37
Bereh Syvashyka (V.P. Kolomiychuk, V.A. Onyshchenko) ............................................ 38
Bile Ozero (V.A. Onyshchenko) .................................................................................. 40
Bir na Merli (O.V. Bezrodnova, N.B. Saidakhmedova) ............................................... 42
Bobrovyi Kut (V.P. Kolomiychuk) ................................................................................ 45
Brytavskyi Lis (V.A. Onyshchenko) ............................................................................. 46
Bushchanske Boloto (O.V. Holovko, V.A. Onyshchenko) ............................................ 48
Byriuchyi Ostriv (V.P. Kolomiychuk, V.A. Onyshchenko) ............................................. 50
Cheremske Boloto (V.A. Onyshchenko) ..................................................................... 53
Cherevaskyi Lis (V.A. Onyshchenko) ........................................................................... 55
Chorna Hora (R.Ya. Kish, V.A. Onyshchenko) .............................................................. 56
Chornohora (V.A. Onyshchenko, V.M. Virchenko) ...................................................... 58
Chornyi Lis (V.A. Onyshchenko) ................................................................................. 62
Chortova Hora (V.A. Onyshchenko, N.V. Shumska) .................................................... 64
Chyvchyny (I.I. Chorney, V.A. Onyshchenko) .............................................................. 65
Davydo-Mykilske (V.A. Onyshchenko, M.M. Peregrym) ............................................ 70
Dnipro – Oril (V.A. Onyshchenko) .............................................................................. 72
Dobrianski Hory (V.A. Onyshchenko) ......................................................................... 74
Dolyna Inhulu (V.A. Onyshchenko) ............................................................................ 76
Dolyna Mertvovodu (V.A. Onyshchenko) ................................................................... 78
Dolyna Nartsysiv (V.A. Onyshchenko) ........................................................................ 81
Dubovetski Skhyly (I.M. Danylyk) .............................................................................. 83
Dubrova (R.Ya. Kish, V.A. Onyshchenko) .................................................................... 84
Dunaiski Plavni (V.A. Onyshchenko) ........................................................................... 86
Dzharylhach (V.A. Onyshchenko, V.P. Kolomiychuk) ................................................... 89
Dzhohul (I.I. Chorney, A.I. Tokaryuk, V.V. Budzhak) ................................................... 91
Dziurkach (I.I. Chorney, A.I. Tokaryuk, V.V. Budzhak) ................................................. 93
Gorgany (V.A. Onyshchenko) .................................................................................... 94
Grofa (V.A. Onyshchenko) ......................................................................................... 96
Hadiatski Luky (N.O. Stetsiuk, V.A. Onyshchenko) ..................................................... 98
Halitsynove (V.P. Kolomiychuk) ................................................................................ 101
Holohirskyi Lis (V.A. Onyshchenko) .......................................................................... 103
Homilshanskyi Lis (V.A. Onyshchenko) ..................................................................... 104
Horaivka (L.H. Lubinska, L.T. Horbniak) .................................................................... 106
Horodnytski Tovtry (H.I. Oliiar) ................................................................................ 108
Horodnytskyi Lis (V.A. Onyshchenko) ....................................................................... 110
372
Hostra Skelia (H.I. Oliiar) ..........................................................................................
Hrakove (N.O. Stetsiuk) ...........................................................................................
Hranitno-Stepove Pobuzhia (V.A. Onyshchenko) .....................................................
Ihrovets – Tavpishyrka (V.A. Onyshchenko, R.Ya. Kish) .............................................
Irpinskyi Lis (V.A. Onyshchenko, O.I. Pryadko) .........................................................
Kadubivska Sinka (I.I. Chorney, A.I. Tokaryuk, V.V. Budzhak) ..................................
Kalmiuskyi Step (V.P. Kolomiychuk, V.A. Onyshchenko) ...........................................
Kamiani Mohyly (V.A. Onyshchenko, V.P. Kolomiychuk) ...........................................
Kanivski Ostrovy (V.A. Onyshchenko) .......................................................................
Kaplivka (I.I. Chorney, A.I. Tokaryuk, V.V. Budzhak ) .................................................
Karadah (V.A. Onyshchenko) ....................................................................................
Karalarskyi Step (V.P. Kolomiychuk, V.A. Onyshchenko) ...........................................
Kasova Hora (V.A. Onyshchenko, N.V. Shumska) ......................................................
Kazantyp (V.P. Kolomiychuk, V.A. Onyshchenko) ......................................................
Kedryn (R.Ya. Kish, V.A. Onyshchenko) .....................................................................
Khomutovskyi Step (V.A. Onyshchenko, V.P. Kolomiychuk) ......................................
Kniazhpilskyi Lis (V.A. Onyshchenko) .......................................................................
Kolodiazne (V.A. Onyshchenko, M.V. Banik, O.V. Bezrodnova, V.V. Rasevich) ...........
Kolivska Ulohovyna (V.A. Onyshchenko) ................................................................
Koncha-Zaspivskyi Lis (V.A. Onyshchenko) ...............................................................
Korsak Mohyla (V.A. Onyshchenko, V.P. Kolomiychuk) .............................................
Kovalivka (Derkach O.M., Kolomiets H.V.) ................................................................
Kozachelaherska Arena (V.A. Onyshchenko) ............................................................
Kreidiana Flora (V.A. Onyshchenko) .........................................................................
Kreidiani Vidslonennia (V.A. Onyshchenko) .............................................................
Kremenchutski Plavni (V.A. Onyshchenko) ..............................................................
Krymski Hory (V.A. Onyshchenko) ............................................................................
Kuchuhury (V.P. Kolomiychuk) .................................................................................
Kuchurhan (V.A. Onyshchenko, O.M. Popova) .........................................................
Kurylivka (V.P. Kolomiychuk) ....................................................................................
Levkivski Lisy (O.O. Orlov) ........................................................................................
Lysa Hora i Sinka (V.A. Onyshchenko, T.D. Solomakha) ...........................................
Lysohirka (V.P. Kolomiychuk) ....................................................................................
Marina Hora (V.A. Onyshchenko) .............................................................................
Markovychi (V.V. Datsiuk) ........................................................................................
Marmaroski Hory (V.A. Onyshchenko, V.M. Virchenko) ...........................................
Medobory (V.A. Onyshchenko) ................................................................................
Medvezhanka (V.A. Onyshchenko, M.M. Peregrym) ................................................
Mehanom (V.A. Onyshchenko) ................................................................................
Mishkovo-Pohorilove (O.M. Derkach, H.V. Kolomiets) .............................................
Molochnyi Lyman (V.P. Kolomiychuk, V.A. Onyshchenko) ........................................
Morochno-2 (V.A. Onyshchenko) .............................................................................
Mykhailivskyi Step (V.A. Onyshchenko, O.M. Derkach) ............................................
Mykhailo-Laryne (O.M. Derkach, H.V. Kolomiets) ....................................................
Mys Martyan (V.A. Onyshchenko) ...........................................................................
Mytrofanivskyi Pivostriv (V.P. Kolomiychuk, V.A. Onyshchenko) ..............................
Naholnyi Kriazh (V.A. Onyshchenko, M.M. Peregrym) .............................................
Novobila (V.A. Onyshchenko, M.V. Banik) ................................................................
373
112
114
115
119
121
122
124
126
128
130
132
137
140
142
144
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148
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150
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190
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196
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208
210
Novohredneve (V.P. Kolomiychuk) ...........................................................................
Nyzhnii Dnipro (V.A. Onyshchenko) .........................................................................
Nyzhniodniprovski Pisky (V.A. Onyshchenko) ..........................................................
Obytichna Kosa (V.P. Kolomiychuk, V.A. Onyshchenko) ...........................................
Olevski Lisy (O.O. Orlov) ..........................................................................................
Opuk (V.A. Onyshchenko) ........................................................................................
Oskilski Skhyly (V.A. Onyshchenko, O.V. Bezrodnova, M.V. Banik) ...........................
Osovynskyi Step (V.P. Kolomiychuk, V.A. Onyshchenko) ..........................................
Pecheniiske Boloto (V.A. Onyshchenko, O.T. Kuzyarin) ............................................
Perebrody (V.A. Onyshchenko, O.I. Pryadko) ...........................................................
Peresyp Aktaskoho Ozera (V.P. Kolomiychuk, V.A. Onyshchenko) ............................
Petro-Ivanivka (О.V. Bezrodnova) ............................................................................
Pidlyska Hora (V.A. Onyshchenko, T.D. Solomakha) .................................................
Podvirivka (I.I. Chorney, A.I. Tokaryuk, V.V. Budzhak) ..............................................
Pohorylivka (I.I. Chorney, A.I. Tokaryuk, V.V. Budzhak) ............................................
Polonskyi Lis (V.A. Onyshchenko) .............................................................................
Polonyna Borzhava (V.A. Onyshchenko) ..................................................................
Polovetskyi Step (V.P. Kolomiychuk) ........................................................................
Potashnianski Lisy (O.O. Orlov, V.A. Onyshchenko) ..................................................
Probabyn (I.I. Chorney, A.I. Tokaryuk, V.V. Budzhak) ...............................................
Provalskyi Step (V.A. Onyshchenko) .........................................................................
Riznykivka (V.A. Onyshchenko) ................................................................................
Romosh (V.V. Datsiuk, V.A. Onyshchenko) ...............................................................
Rys (V.A. Onyshchenko) ...........................................................................................
Samarskyi Lis (V.A. Onyshchenko) ............................................................................
Savranskyi Lis (V.A. Onyshchenko) ...........................................................................
Serbyno (V.A. Onyshchenko) ....................................................................................
Sharivka (V.A. Onyshchenko, M.V. Banik) .................................................................
Shatski Ozera (V.A. Onyshchenko) ...........................................................................
Shebutynskyi Yar (I.I. Chorney, A.I. Tokaryuk, V.V. Budzhak) ....................................
Sinozhai (I.I. Chorney, A.I. Tokaryuk, V.V. Budzhak) ................................................
Skhidne Roztochia (V.A. Onyshchenko) ....................................................................
Skhidnyi Churiuk (V.P. Kolomiychuk,V.A. Onyshchenko) ...........................................
Skhyly Kohylnyka (V.A. Onyshchenko, O.M. Popova) ...............................................
Skhyly Krasnoi i Kobylky (V.A. Onyshchenko) ...........................................................
Slovechanskyi Kriazh (V.A. Onyshchenko) ................................................................
Somyno (V.A. Onyshchenko) ....................................................................................
Sovyi Yar (V.A. Onyshchenko) ...................................................................................
Striltsivskyi Step (V.A. Onyshchenko) .......................................................................
Stuzhytsia (V.A. Onyshchenko) ................................................................................
Sviai Hory (V.A. Onyshchenko) ................................................................................
Svydovets (V.A. Onyshchenko) .................................................................................
Syra Pohonia (V.A. Onyshchenko) ............................................................................
Tarkhankut (V.A. Onyshchenko) ...............................................................................
Tendrivska Zatoka (V.A. Onyshchenko, O.Yu. Umanets) ...........................................
Tepe-Oba (V.A. Onyshchenko) .................................................................................
Ternava – Dnister (V.A. Onyshchenko) .....................................................................
Tiup-Tarkhan i Kalynivskyi Pivostriv (V.P. Kolomiychuk, V.A. Onyshchenko) .............
374
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284
286
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293
294
298
300
303
306
Tovtrivska Sinka (I.I. Chorney, A.I. Tokaryuk, V.V. Budzhak) .................................... 307
Triokhizbenskyi Step (V.A. Onyshchenko) ................................................................ 309
Troitska Balka (V.P. Kolomiychuk, V.A. Onyshchenko) .............................................. 311
Trosianets (V.A. Onyshchenko, O.M. Popova) ......................................................... 312
Tsetsyno (I.I. Chorney, A.I. Tokaryuk, V.V. Budzhak) ................................................. 314
Tulyntsi – Makedony (V.A. Onyshchenko) ................................................................ 315
Turova Dacha (V.A. Onyshchenko, N.V. Shumska) .................................................... 317
Ubort – Bolotnytsia (V.A. Onyshchenko) .................................................................. 318
Uholka – Shyrokyi Luh (V.A. Onyshchenko) .............................................................. 321
Usia – Shutnivtsi (L.H. Lubinska, V.A. Kolodiy) ........................................................ 323
Vasylivka (V.P. Kolomiychuk, L.V. Markivska) ............................................................ 325
Vasylivskyi i Rozkopynskyi Yary (I.I. Chorney, A.I. Tokaryuk, V.V. Budzhak) ............... 326
Velyki Holdy (N.V. Shumska, V.A. Onyshchenko) ...................................................... 328
Verbetska Tovtra (V.A. Onyshchenko) ....................................................................... 330
Verkhniobuzki Lisy (V.A. Onyshchenko) .................................................................... 331
Vitrohon (V.A. Onyshchenko) .................................................................................... 333
Volyzhyn Lis (V.A. Onyshchenko, O.Yu. Umanets) .................................................... 334
Vovchanski Skhyly (V.A. Onyshchenko, O.V. Bezrodnova, M.V. Banik) ..................... 336
Vtenske Boloto (V.A. Onyshchenko) ........................................................................ 338
Vymushiv (I.I. Chorney, A.I. Tokaryuk, V.V. Budzhak) ............................................... 339
Vyshnivka (I.I. Chorney, A.I. Tokaryuk, V.V. Budzhak) ............................................... 341
Yaiko Perehinske (V.A. Onyshchenko) ...................................................................... 342
Yelanetskyi Step (V.A. Onyshchenko) ....................................................................... 343
Yemilchynski Lisy (O.O. Orlov, V.A. Onyshchenko) ................................................... 346
Yulivski Hory (R.Ya. Kish, V.A. Onyshchenko) ............................................................ 348
Zaplava Desny (V.A. Onyshchenko, O.V. Lukash) ...................................................... 349
Zaplava Latorytsi (V.A. Onyshchenko) ...................................................................... 353
Zaplava Prypiai (V.A. Onyshchenko) ....................................................................... 355
Zaplava Siverskoho Dintsia (V.A. Onyshchenko) ....................................................... 357
Zaplava Tysy (R.Ya. Kish, V.A. Onyshchenko) ............................................................. 359
Zaplavni Lisy na Borzhavi (R.Ya. Kish, V.A. Onyshchenko) ........................................ 361
Zaplavy Dnipra i Sozha (V.A. Onyshchenko, O.V. Lukash) ......................................... 362
Zelena Balka (V.A. Onyshchenko) ............................................................................. 365
Zholoby (H.I. Oliiar, H.P. Protsiv) .............................................................................. 367
Zhyzhava (V.A. Onyshchenko) .................................................................................. 368
Zolochivka (O.T. Kuzyarin, V.A. Onyshchenko) .......................................................... 370
375
Наукове видання
Важливі ботанічні території України
За редакцією д.б.н. В.А. Онищенка
Підписано до друку 16.05.2017. Формат: 60 x 84/16. Папір офс.
Гарнітура: Calibri. Друк офс. Умов. друк. арк. 21,86
Замовлення № 2017-10
“Альтерпрес”, 01034 Київ, вул. В. Житомирська, 28.
Свідоцтво про реєстрацію ДК №177 від 15.09.2000 р.
Віддруковано: “Альтерпрес”, 04112 Київ, вул. Шамрила, 23
376