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133 Phytocoenologia 34 (1) 133-161 Berlin - Stuttgart, March 18, 2004 Vegetation units of the Argentine Semiarid Chaco: The Toba-Pilagá perception by G.F. Scarpa & P. Arenas, Instituto de Botánica Darwinion (IBODA), San Isidro, Argentina with 3 Figures and 4 Tables Abstract. This paper studies the nomenclature and classification assigned to vegetation units by the Toba-Pilagá Indians of the Gran Chaco. The ethno-botanical research consisted of 12 campaigns in which we worked with 58 informants. The study describes a particular aspect of the traditional knowledge of plants: the perception of the plant environment. The vernacular names of plants, the names of vegetation units, and their respective classification are provided. Of a total of 235 recorded species, 217 taxa form part of some type of vegetation unit, whereas 18 are not part of any type of plant unit. A total of 196 vernacular names referring to 304 vegetation units were recorded. An ethno-taxonomic system by which the Toba-Pilagá classify 277 vegetation units into 27 categories according to attributes such as physiognomy, succession, relative ease of access and visibility, and according to their association with elements of the landscape is revealed. These categories are organised in turn within a hierarchical system with different degrees of inclusiveness. Keywords: Ethnobotany, ethnophytogeography, Toba-Pilagá people, Chaco, Argentina. 1. Introduction The way in which a human group names and classifies different vegetation units in its environment is closely related to the vernacular names given to plants, and the particular concept of space by which members of the group relate to the landscape. The present study is based on an ethnophytogeographic perspective; this discipline organises and encodes empirical knowledge of vegetation units. It also belongs to the field of the ethnosciences, which deal with the way in which Man constructs his world of experience as reflected in the vernacular terminology, and the way in which he organizes, categorizes and classifies its terms (Grebe Vicuña, 1986). Peasant communities and indigenous groups name and classify vegetation units according to their own specific cultural criteria. However, according to a brief review of the recent literature on the ethnobotany and toponymy of hunter-gatherer groups by Johnson (2000), relatively few studies provide an exhaustive list of identified and named ecological types. Among them special mention should be made of studies by Friedberg (1990) for the Bunaq of Timor (Indonesia); and Aldunate et al. (1981) for Andean peoples, Johnson (2000) for the Gitksan of northeast British Columbia, Martin (1974) for the Khmer of Cambodia, and Shepard et al. (2001) 134 for the Matsigenka of the Peruvian Amazon, just to mention a few of the most representative studies. With regard to present-day hunter-gatherer groups in the Gran Chaco, there are no studies providing a complete list of the vernacular names assigned to vegetation units, or showing how they have been traditionally classified. However, a relevant precedent is a brief ethno-ecological lexicon, with emphasis on vegetation, compiled by Filipov (1996). For the same ethnic group Idoyaga Molina (1988-89, 1990-91) also provides a classification of plants by their life-forms, and according to cosmological categories. In addition to these studies are those on toponymy in the native languages of the Chaco. Of these, special mention should be made of the studies by Martínez Crovetto (1987), Wright (1991), Dell’Arciprete (1991), De la Cruz (1993, 1995), and Palmer (1995), in which different types of information referred to in place-names, including the vernacular names of numerous vegetation units, are discussed. Finally, we also drew upon the basic information on Toba-Pilagá plant names compiled by Arenas (1993). In the present study we have compiled a list of denominations given by the Toba-Pilagá to vegetation units in the semi-arid Chaco. On the basis of interviews with informants we have also outlined an approach to the criteria used by these people to classify different areas of their botanical environment. This was done after numerous research campaigns in which we performed detailed ethno-biological surveys with particular emphasis on ethno-botanical aspects, designed to reveal and understand the relationship between the cultural and natural elements of the Toba-Pilagá landscape (Arenas, 1995). We also wish to point out that we addressed the subject matter dealt with in this study only after gaining a thorough knowledge of the ethnography of this human group. 2. Study area The Gran Chaco is a vast alluvial plain formed by quaternary sediments, on which allochthonous rivers have markedly shaped the landscape. Our area of study is located on the wide floodplain of the River Pilcomayo in the semi-arid portion of the Argentine Chaco (Fig. 1). From its Andean origins it meanders along a course punctuated by frequent bends due to the extremely flat terrain. This particular fluviomorphological dynamic has produced numerous circular or crescent-shaped lagoons which originated in changes in the course of the river (Adámoli et al., 1972; Ramella & Spichiger, 1989; Prado, 1993). As a result of these processes the present topographic gradient is made up of: upland areas which correspond to the old alluvial plain; intermediate areas representing the old ridges of abandoned meanders; and low-lying areas comprising the old and modern-day river beds. According to the phytogeographic classification established by Cabrera & Willink (1980) the area is located in the Western District of the Province of Chaco. In the upland areas of the gradient, zonal communities represented by xerophilous forests 135 Fig. 1. The Toba-Pilagá indians area. of mesophanerophytes such as Schinopsis lorentzii, Aspidosperma quebracho-blanco, and Bulnesia sarmientoi are dominant (names of taxa are according to Zuloaga & Morrone, 1996, 1999; Zuloaga et al. 1994). The intermediate area is inhabited by azonal communities represented by xerophilous forests of microphanerophytes such as Prosopis alba, P. nigra, Geoffroea decorticans, and Ruprechtia triflora; woodlands of Mimozyganthus carinatus; and different species of Acacia. The low-lying areas are inhabited by grasslands of Cynodon dactylon, Paspalum laxum, arboreal communities where Tessaria integrifolia and Salix humboldtiana are dominant, Copernicia alba palms, halophyte steppes dominated by Ambrosia tenuifolia and Phyla reptans, and hydrophytic communities (Morello, 1970). Riparian environments are superimposed on a matrix of interfluvial environments all over the region, thus forming a mosaic-like landscape pattern. 136 In the past, woodlands alternated with extended grasslands forming a typical savanna landscape. However, during the first part of the twentieth century the region underwent a process of grassland encroachment by advancing shrub and tree formations, such that the original landscape underwent a complete alteration. Among the physiognomic studies on the vegetation of the Gran Chaco emphasis should be made of those by Castellanos (1958); Morello & Saravia Toledo (1959); Morello & Adámoli (1974), Ramella & Spichiger (1989) and Prado (1993). From a phytosociological point of view we can mention studies by Eskuche (1992); Galán de Mera & Navarro (1992); Navarro (1997); Navarro & Fuentes (1999), Fuentes & Navarro (2000), and Galán de Mera (2002). Navarro (1997); Navarro & Fuentes (1999), and Fuentes & Navarro (2000) establish some kind of parallelism between series of phytosociological units and their native names. These observations highlight the importance of rescuing and interpreting the native forms of perception of the plant environment, and the need to compare it with the scientific classification. 2.1. The Toba-Pilagá people The Toba-Pilagá are an ethnic group belonging to the Guaycurú linguistic family (Métraux, 1946; Mason 1950; Tovar 1961; Censabella, 1999); they are closely related to other Toba groups, as well as to other living and extinct ethnic groups (Métraux, 1937; 1946; Tebboth, 1943). They live in the Province of Formosa in the north of the Argentine Chaco (Fig. 1). The current population is of about 2000 people, in 13 rural villages, and one neighbourhood in the outskirts of the small town of Ing. G. N. Juárez. The Toba-Pilagá have an intimate knowledge of nature. Their perception of the surrounding environment is very concise and is based on explanations of the structure of the universe, representing their cosmovision. The Toba-Pilagá perceive the universe as a structure composed of 7 superimposed layers (some informants refer to only 3 or 5) of which only one corresponds to the terrestrial surface, the stratum on which they live and on which accidents and events involving man – and the vegetation units- occur. Only the shaman knows and visits the other spaces. The common mortals experience life on the terrestrial surface, and this is the layer we will refer to in this study. This is where the woodlands, rivers, lagoons, the village, and other elements that are part of daily existence are found. While knowledge of the cosmos is reserved to initiated or erudite individuals, the knowledge of nature is inherent to any member of the social group. The Toba-Pilagá were gatherers, hunters and fishermen, and practiced a little agriculture. Their activities allowed them to acquire an intimate knowledge of the local geography as well as of the particular characteristics of the flora and fauna. Since the start of the twentieth century they have been in close contact with national society, to such an extent that they have undergone an intense process of cultural change. Nevertheless, they still conserve a great deal of the essential features of their identity, such as their language, social organization, ethical norms, and some of their subsistence habits. 137 3. Data base and ethnophytogeographical methodology Starting in 1993, a total of 12 campaigns were undertaken to investigate the ethnobotany of the Toba-Pilagá. The first data relating to the lexicon on vegetation units were obtained from the ethnobotanical material compiled during the campaigns. In order to carry out in-depth research on the matter, two additional and specific campaigns were made to the Toba-Pilagá villages in 2000 and 2001, and the data was used to verify and expand preliminary data obtained during the first campaigns. The information and materials were collected with the help of 58 informants who were given semi-structured interviews in which they were asked about the vernacular names of plants and vegetation units. The informants were chosen from among individuals of 40 to 80 years of age. Outings to the field were made with each one of them in order to verify in situ the types of communities they referred to, and the vernacular names were recorded. The field notes, the tape recordings of oral information and other documents are now in the archives of the Instituto de Botánica Darwinion. The data were cross-checked with at least two informants. When differing points of view were recorded, only those on which there was general agreement among the interviewees were accepted. During field work inquiries were made into the characterization of the vegetation units referred to according to the procedures used in ethnophytogeography (Martin, 1974) . We asked about their geographical location, land relief, accessibility, structure, spatial pattern, relative extension, associations with elements of the landscape, the nature and general features of these elements, ecological dynamics (or life history of the vegetation unit), and general significances given to them. From the ethnoscientific perspective, the usual methodology was applied in order to broach these questions; the field work revealed the points of view of the society being studied (emic perspective), which were later analysed in the light of the standards of western science (ethic perspective) (Fowler, 1979; Grebe Vicuña, 1986; Winthrop, 1991; Martin, 1995). As a first step in revealing the Toba-Pilagá perception of the vegetation units, a linguistic analysis of the vernacular names assigned was made (Friedberg, 1974; Martin, 1974). This identified the existence in the Toba-Pilagá language of particles which when added to the name of the plant formed collective nouns, which represented minimum units of grouping of the vegetation for these Indians. On the basis of this, an exhaustive questionnaire about the plants that formed these communities was drawn up. As a result of the analysis of the meanings of the intervening particles, basic grouping levels of vegetation units were identified. However, the analysis of the vernacular nomenclature is only a first approach to the study of ethnotaxonomy. The ethnophytogeographic papers consulted all indicate that there is no standardised method to follow when attempting to explain those groupings not derived directly from the nomenclature (Martin, 1974; Friedberg, 1974). 138 Given the multiplicity of reference systems to which Indian taxonomies mention, the only way to understand the ethnophytogeography of a human group seems to consist in distinguishing “the words that the Indian uses to describe the forests, spontaneous reactions and attitudes which acknowledge the extension of the semantic field of certain terms used” (Martin, 1974). On the basis of the characterisation of the vegetation units made by our informants supraordinal grouping levels were specified. The categories within each level were related to each other, and with the basic grouping levels already mentioned, through a loosely inclusive hierarchical system established according to the particular relationships of inclusion expressed by the Indians. The comparison between the vernacular classification of the vegetation units and that derived from phyto-sociological studies is totally beyond the scope of this study, considering the ethnobotanical origin of our data -focused only on the level of plant species-, and because no phytosociological inventory has been made. Nevertheless, a description of Toba-Pilagá categories is made by using the physiognomic perspective,whenever possible, because it was the one that best fits the viewpoint of the Indians. In order to record terms in the Toba-Pilagá language we used their equivalent terms in Spanish, except for those sounds that do not exist in that language. These are “q”: postvelar occlusive voiceless; “β”: fricative bilabial voiced; “#”: occlusive glottal voiceless; “h”: fricative glottal voiceless; “G”: fricative postvelar voiced; “w”: semiconsonant bilabial voiced; and “y”: semiconsonant palatal voiced. For further precision on the phonetic features used, reference should be made to De la Cruz (1993, 1995), and Mendoza & Browne (1995). 4. Results In this study we have used the terms “vegetation unit” and “plant unit” to refer to the collective botanical nouns identified in the Toba-Pilagá discourse. However, these concepts do not always coincide with the meaning given to them in natural sciences. In fact, it was observed that both in the field and in the Toba-Pilagá discourse collective botanical nouns sometimes refer to what is defined in ecology as “populations”, sometimes to “communities”, and sometimes to actual “landscape units”. Firstly, we shall describe the nomenclatural system which vegetation units belong to, and then show how the Toba-Pilagá group them within an ethnoclassificatory system. 4.1. Identification and nomenclature of the vegetation units The Toba-Pilagá identify as a basic plant unit any group of plants of at least five individuals of the same species that share the same space. This benchmark number is the only one informants refer to with any precision when specifically asked about the subject. These units may occupy surface areas ranging from one square meter 139 (especially herbaceous plants) to several square kilometers (some vegetation units of palms); and may refer both to pure stands (monospecific) and to plurispecific stands with a clearly dominant species. In all cases, these basic plant units were defined in relation to a single vernacular species, not including associations of two or more species in any one case. The language elements making up the names of the vegetation units are: Basic Term: The Toba-Pilagá plant name that identifies a particular unit. Suffix: A particle that, when added to a basic term, denotes an indefinite and numerous group of at least five individuals of the same species. There are different types of suffixes, each one bearing a particular meaning. They are: • 'hat/'sat: Indicates that the plant unit is in a higher area of the altitudinal gradient, which is not subject to flooding, whose spatial shape is undefined. Less frequently it refers to plant communities in general (see discussion); • 'hatain'gi: Indicates that the vegetation unit is located in a permanently flooded and rounded lowland; • 'hata'ge: Indicates that the plant unit is located along an elongated and temporarily flooded lowland; • lo’laGae’hat: Indicates that the vegetation unit is in either a young developmental phase or as saplings (only used for tree species). In a few cases the names formed in this way are used together with an attributive that refers to the area covered by the plant formation: o'lek indicates a small area. Table 1 exemplifies the main structure referred to by the nomenclatural system of vegetation units. The vernacular names of the identified vegetation units are enumerated in Table 2 (phonetic variations identified for some of them were excluded from the reckoning), together with the scientific name, and basic term of each species. The first column of the table shows the category the plant unit belongs to (this will be dealt with in the next paragraph). Because of the high number of vegetation units identified by these indigenous people, a valid question would be whether all the plant species mentioned by them form plant units. It was clear that not all species bear a collective botanical name. They explained that these plants “live alone”, or that “many individual plants rarely grow together”, or that, Table 1. Example of the nomenclature system of the vegetation units. Nombre científico Término de base Término de base sufijado Significado Prosopis alba ma'pihat ma’pik mapi'hataingi mapihata'ge ma’pik lola'Gae'hat Comunidad de P. alba de zonas altas Idem anterior alrededor de un bajo circular Idem anterior al lo largo de un bajo alargado Brotal de P. alba 140 Table 2. Vegetation units identified by the Toba-Pilagá people arranged by the ethnoclassificatory categories they belong to. (*: understorey plant). Vegetation unit categories/ Taxa included* Basic term Vegetation unit BI-1. - ' iaq ("Woodland") BI-1.1- ' iaq halo'te BI-1.1.A - ' iaq 'tadaik Achatocarpus praecox* ' lamaqaik ' lamaGae' hat Aspidosperma quebracho-blanco no' dik no' dihat Bromelia hieronymi* ko' taGayak ko' taGayaGa' hat Bromelia serra* ko' taGayak ko' taGayaGa' tha Bromelia serra* kota' ke kotake' pihat; kota' kehat Bulnesia sarmientoi qasa' qaik qasa' qaik' sat Capsicum chacoense* ko' dae ko' daehat Ceiba insignis qope' daGañik qope' daGañihat Deinacanthon urbanianum* kail' te kail' tehat Dyckia sp.* kail' te kail' tehat Dyckia sp.* ne' kota neko' taGahat Schinopsis lorentzii ke' taqaik ke' taGahat Sideroxylon obtusifolium ssp. obtusifolium BI-1.1.B - ' iaq we' daGañik we' daGañihat BI-1.1.B.1 - Unnamed category Caesalpinia paraguariensis wa' laGañik wa' laGañihat Geoffroea decorticans ' taqaik ' taqaehat Parkinsonia aculeata ñie' chik ñi:' chiehat Petiveria alliacea* hepe' toq hepe' toGohat Prosopis aff. alba noqolo' kik noqolo' kihat Prosopis alba ma' pik mapi' hat, mapi' hat o' lek Prosopis elata dela' kik dela' kihat, dela' ki#ehat Prosopis kuntzei ta' de:k ta' de:hat Prosopis nigra pa:' taik pa:ta' hat Prosopis ruscifolia ' laGaya' kaik ' laGaya' hat, ' layaGa' hat Prosopis sp. dela' kik delae' hat Prosopis x elata pana' naik pana' naihat Prosopis x nigra dela' kik delae' hat Ruellia hygrophila* e' daGaik e' daGai' hat Ruellia ciliatiflora* ke' taGaik ke' taGahat Ruellia erythropus* e' daGaik e' daGai' hat Ruellia coerulea* ke' taGaik ke' taGahat Tabebuia nodosa pa' lidokoik pa' lidoGahat Zizyphus mistol ' nalaik ' nalahat ' taqaik ' taqaehatain' gi BI-1.1.B.2 - Unnamed category BI-1.1.B.2.a - Unnamed category Geoffroea decorticans 141 Vegetation unit categories/ Taxa included* Basic term Vegetation unit Parkinsonia aculeata Prosopis nigra Prosopis ruscifolia ñie' chik pa:' taik ' laGaya' kaik ñi:' chiehatain' gi pa:ta' hataingi ' laGayaga' hataingi Tabebuia nodosa pa' lidoqoik pa' lidoGahatain' gi BI-1.1.B.2.b - Unnamed category Geoffroea decorticans ' taqaik ' taqae' hatage Prosopis alba ma' pik mapi' hatage Prosopis nigra pa:' taik pa:ta' hata' ge BI-1.1.C - Unnamed category Albizia inundata ya' talik ya' talihataingi Calycophyllum multiflorum pa' geaGa ' lo#ok paGea' ga' lohat; pa' geaGa' lohata Maclura tinctoria ssp. mora qoBi' ye qoBi' hat; qo' Bihatain' gi Phyllostylon rhamnoides qoBi' ye la' tedae qoBi' ye la' tedaetain' gi Pisonia zapallo na' se:k na' se:hat; na' se:hatain' gi Sapindus saponaria wedaelqo' lakik wedaelqo' laehat Sapium haematospermum ki' chiaGadik ki' chiaGadihatain' gi; ki' chiaGadih BI-1.2.A - Unnamed category Acacia aroma pa' Gaik paGage' hat Acacia furcatispina waGea' qaik, waGea' qait' wayaGa' hat Acacia praecox petegeanaGa' kaik petegeanaGa' hat BI-1.2 - ' iaq haka' lok ✁ waGea'qaik waGeaqai' hat Acanthosyris falcata pa' Gañik pa' Gañihat Arundo donax qoqo' te qoqode' hat Cercidium praecox paGa' chik paGa' chihat Grabowskia duplicata todo' lokik todo' lokihat; todo' lo(Ga)' hat Habranthus sp.* la' chie; la' chi la' chiehat Marsdenia castillonii* howa' qaik howaGa' hat Maytenus vitis-idaea hata' chik hata' chiehat Mimozyganthus carinatus no#legea' kaik no#legeaGa' hat Opuntia anacantha* pichi' ñi pichi' ñiehat Ruprechtia triflora oma' qaik omaGa' hat Synandrospadix vermitoxicus* naqaigea' Ga la' #ta naqaigea' Ga la' taGahat Ximenia americana todo' lokik todo' lokihat; todo' lo(Ga)' hat BI-1.2.B - Unnamed category Acacia aroma pa' Gaik paGage' hataingi Arundo donax qoqo' te qoqode' hataingi Prosopis alba ma' pik mapi' hatain' gi Ruprechtia triflora oma'qaik omaGa' hataingi 142 Vegetation unit categories/ Taxa included* Basic term Vegetation unit βiaq poñio' le BI-1.3 - 'β BI-1.3.A - yaga' ta#a' ge BI-1.3.A.1 - Unnamed category Acacia caven pa' Gaik la' chielik paGaela' chieli' hat Acacia praecox petegeanaGa' kaik petegeanaGa' hat waGea' qaik waGeaqai' hat Bromelia hieronymi* ko' taGayak ko' taGayaGa' hat Bromelia serra* ko' taGayak ko' taGayaGa' hat kota' ke kota' kehat, kotake' pihat Castela coccinea ' helkaik na' maik ' helkaik na' maik' te:hat Celtis spp. chiyaGa' dik chiyaGa' dihat Cyclolepis genistoides wa' BiaGaik wa' BiaGaihat Deinacanthon urbanianum* kail' te kail' tehat Dyckia sp.* kail' te; ne' kota kail' tehat, neko' taGahat Mimosa detinens petigeana' qaik petigeana' qaihat Mimosa detinens waGea' qaik waGea' qaihat Prosopis ruscifolia ' laGaya' kaik ' laGaya' hat, ' layaGa' hat Prosopis sericantha ta' de:k la' chielik;' tade:k ta' de:k la' chielihat; tade:' hat BI-1.3.A.2 - Unnamed category Acacia curvifructa pa' Gaik la' chielik paGaela' chieli' hataingi Acacia caven pa' Gaik la' chielik paGaela' chieli' hataingi Celtis spp. chiyaGa' dik chiyaGa' dihatain' gi Prosopis ruscifolia ' laGaya' kaik ' laGayaga' hataingi BI-1.3.B. Unnamed category BI-1.3.B.1 - Unnamed category Aloysia virgata tanaGana' Ga lepe' naqa' tetanagana' Ga lepe' naqa' tehat Capparis retusa tegea' qaik tegeaGa' hat Capparis salicifolia el' kik eli' hat Capparis speciosa nelo' mik nelomi' hat Capparis tweediana kili' li#i kili' li' hat Lophocarpinia aculeatifolia wo' lege wole' gehat Lycium americanum qo' BioGonaqa' tae qo' BioGonaqa' taehat Lycium cuneatum ' pi#tala' dik pi#tala' dihat Lycium nodosum ' pi#tala' dik pi#tala' dihat Prosopis alba ma' pik ma' pik lol' laGahat Vallesia glabra hama' ñik hamañi:' hat Zizyphus mistol ' nalaik ' nalaik lol' laGahat BI-1.3.B.2 - Unnamed category Capparis retusa tegea' qaik tegeaGa' hataingi 143 Vegetation unit categories/ Taxa included* BI-1.4 - ' iaq ni' yege Cercidium praecox Basic term Vegetation unit ✂ paGa' chik paGa' chihat NO-2. - ' nonaGa ("Field") NO-2.1- ' nonaGa ' tadaik NO-2.1.A - Unnamed category Aristida mendocina awaqa' pi; ma' ñie la' de awaka' pihat; ma' ñie la'hatde Copernicia alba ' chaik chai' hat Digitaria insularis pa' Ga la' wo pa' Ga' lo#ahat Echinochloa cf. polystachya pol' chaq pol' chaGahat Echinochloa cf. polystachya awaka' pi awaka' pihat Elyonurus muticus ' chi#maGa ' chi#maGa' hat Eragrostis lugens awaka' pi awaka' pihat Gouinia latifolia qoqo' ta qoqode' hat Heliotropium procumbens pa' Ga la' wo pa' Ga' lo#ahat Hymenachne amplexicaulis pol' chaq pol' chaGahat Imperata brasiliensis ' na#iky' Got ' na#ikyGode' hat Opuntia chakensis qo' chaq qo' chaGahat Opuntia discolor pichi' ñi; ma' ñik lo' qo pichi' ñiehat; mañieloqo' hat Opuntia elata yate' nak yate' naGahat Opuntia quimilo qadaqa' chik qadaqa' chihat Opuntia sulphurea var. pampeana ma' ñik lo' qo; pichi' ñi mañieloqo' hat; pichi' ñiehat Panicum sp. pol' chaq pol' chaGahat Paspalum conjugatum awaka' pi; pol' chaq awaka' pihat; pol' chaGahat Pennisetum frutescens qoqo' ta qoqode' hat Ruellia hygrophila e' daGaik e' daGai' hat Ruellia ciliatiflora ke' taGaik ke' taGahat Ruellia erythropus e' daGaik e' daGai' hat Ruellia coerulea ke' taGaik ke' taGahat Sporobolus pyramidatus awaka' pi awaka' pihat Stetsonia coryne laGa' dik laGa' dihat Trichloris crinita awaka' pi awaka' pihat Tripogon spicatus awaka' pi awaka' pihat Trithrinax biflabellata laqa' ta laqa' tahat NO-2.1.B.1- Unnamed category Aristida mendocina ma' ñie la' de ma' ñie la' dehataingi Copernicia alba ' chaik ' chaihatain' gi Echinochloa cf. polystachya awaka' pi awaka' pihataingi pol' chaq pol' chaGahatain' gi NO-2.1.B - Unnamed category 144 Vegetation unit categories/ Taxa included* Basic term Vegetation unit Hymenachne amplexicaulis Imperata brasiliensis pol' chaq ' na#iky' Gode pol' chaGahatain' gi ' na#ikyGo' hatain' gi Panicum sp. pol' chaq pol' chaGahatain' gi Paspalum conjugatum Pennisetum frutescens pol' chaq qoqo' ta pol' chaGahatain' gi qoqode' hataingi NO-2.1.B.2 - Unnamed category Copernicia alba ' chaik ' chaihata' ge NO-2.2 - ' nonaGa pe' geat Acanthospermum hispidum ' naGadae ' naGada' hat Alternanthera pungens ta#a' hot ta#aho' hat Bidens subalternans ma' ñie le' lote' ke ma' ñielelote' kehat Boerhavia diffusa var. leiocarpa qo' doGona qo' doGona' hat Cenchrus equinatus i' naGadae i' naGadae' hat Cereus stenogonus ha' Gachik ha' Gadihat Chenopodium ambrosioides ñiaGa' diaGa' leñi ñiaGa' diaGa' lehatain' gi ñiaGa' diaGa' lehat Euphorbia lasiocarpa qo' doGona qo' doGona' hat Euphorbia serpens wol' cha#a; qo' doGona wol' cha#ahat; qo' doGona' hat Harrisia bonplandii ' neko ' nekoGa' hat Nicotiana longiflora maGa' yi leqochia' Ga maGa' yi leqochiaGa' hat Opuntia anacantha pichi' ñi pichi' ñiehat Opuntia chakensis qo' chaq qo' chaGahat Opuntia discolor ' dowoq; ma' ñik lo' qo ' dowoGo' hat; mañieloqo' hat Opuntia elata yate' nak yate' naGahat Opuntia ficus-indica sae' ñik saeñi' hat Opuntia quimilo qadaqa' chik qadaqa' chihat Opuntia salmiana ' dowoq ' dowoGo' hat Opuntia sulphurea var. pampeana ma' ñik lo' qo; pichi' ñi mañieloqo' hat; pichi' ñiehat Opuntia utkilio ' dowoq ' dowoGo' hat Pectis odorata penaGa' di penaGa' dihat Physalis neesiana kochi' maGañik kochi' maGañi' hat Physalis viscosa qoto' ñi qoto' ñiehat Portulaca oleracea ' qato; qo' doGona ' qatoGo' hat; qo' doGona' hat Portulaca sp. ' qato ' qatoGo' hat Quiabentia verticillata ' dowoq la' chik ' dowoq la' chihat Ricinus communis ' wotepe la' chielik ' wotepe la' chielihat Rivina humilis wa' qaw lai' te wa' qaw lai' tehat Selaginella sellowii ' qol ' qolhat Senna morongii ' wayaGa na' maik ' waGayaGa' hat Senna obtusifolia ' wayaGa na' maik ' waGayaGa' hat 145 Vegetation unit categories/ Taxa included* Basic term Vegetation unit Senna occidentalis Senna pendula var. paludicola ' wayaGa na' maik ' wayaGa na' maik ' waGayaGa' hat ' waGayaGa' hat Sida cordifolia pa' Ga la' wo pa' Ga' lo#ahat Solanum argentinum pioGola' dik pioGola' dihat Solanum conditum ta' pañi ta' pañihat Solanum elaeagnifolium Solanum hieronymi ta' pañi ta' pañi la' te# ta' pañihat ta' pañ i la' tehat Solanum sisymbriifolium ' niyaq lae' te ' niyaGalae' tehat, ' niyaq lae' tehat Sonchus asper na' lachi la' ta#a nala' chihat Sphaeralcea bonariensis pa' Ga la' wo pa' Ga' lo#ahat Stetsonia coryne laGa' dik laGa' dihat Tagetes minuta ma' ñiek le' lote' ke ma'ñie lelote' kehat Talinum paniculatum qo' doGona; ' qato qo' doGona' hat; ' qatoGo' hat Talinum polygaloides kowoqo' lek kowoqo' le:hat Talinum triangulare qo' doGona; ' qato qo' doGona' hat; ' qatoGo' hat Tragia hieronymi na' naik lodai' te na' naik lodai' tehat Xanthium spinosum i' naGadae i' naGadae' hat kalgea' Gaik kalgeaGai' hat nol' ke nolke' hat Cyperus sp. ' piaGeana' Gaik ' piaGeanaGai' hat Echinodorus sp. diki' chik diki' chihatain' gi Nicotiana glauca qoin' yila' qaik qoin' yilaqai' hat Phyla reptans kalgea' Gaik kalgeaGai' hat NE-3.1 - ne' dep ' tadaik Nymphaea gardneriana diki' chik diki' chihatain' gi Salix humboldtiana lo' chik lo' chi:hat,o'l chiehat o' lek lo' chik lo' chiehatain' gi ha' laq halaGa' hat; halaGa' hataingi kalgea' Gaik kalgeaGai' hat nol' ke nolke' hat Cyperus sp. ' piaGeana' Gaik ' piaGeanaGai' hat Echinodorus sp. diki' chik diki' chihatain' gi Nicotiana glauca qoin' yila' qaik qoin' yilaqai' hat Phyla reptans kalgea' Gaik kalgeaGai' hat Urera aurantiaca po' qo po' qohat NO-2.3 - ' nonaGa ' dalaGaik Alternanthera paronichyoides ssp. chacoënsis Baccharis salicifolia NE-3.- ne' dep ("Marsh") Tessaria integrifolia NE-3.2. ne' dep qoqo' te Alternanthera paronichyoides ssp. chacoënsis Baccharis salicifolia 146 Vegetation unit categories/ Taxa included* Basic term Vegetation unit NE-3.3 - ne' dep ' lolaGae Geoffroea decorticans ' taqaik ' taqae lola' Gae' hat OC- 4. - ("Out of classification") OC- 4.1. - no' Gop wet#' ot Azolla caroliniana ' qol ' qolhatain' gi; ' qolhat Eichhornia azurea ' molalke' tela molalke'telahatain' gi Lemna sp. ' qol ' qolhatain' gi; ' qolhat Limnobium spongia ssp. laevigatum to' pi to' pihatain' gi Marsilea sp. ' qol ' qolhatain' gi; ' qolhat Nymphoides indica ' molalke' tela molalke' telahatain' gi Pistia stratiotes to' pi to' pihatain' gi OC-4.2 - no' Gop layo' got Ceratophyllum demersum ' qol ' qolhatain' gi; ' qolhat OC-4.3 - Unnamed category Albizia inundata ya' talik ya' talihataingi Baccharis salicifolia nol' ke nol' kehatain' gi Byttneria filipes ' naliem lodai' te ' naliem lodai' tehat Calycophyllum multiflorum pa' geaGa ' lo#ok paGea' ga' lohat, pa' geaGa' lohata Cynodon dactylon awaka' pi ' poleo awka' pihat; awaka' pihataingi Cyperus sp. ' piaGeana' Gaik ' piaGeanaGai' hataingi Eleocharis elegans ' qol ' qolhat; ' qol hatain' gi Maclura tinctoria ssp. mora qoBi' ye qoBi' hat; qo' Bihatain' gi Phyllostylon rhamnoides qoBi' ye la' tedae qoBi' ye la' tedaetain' gi Pisonia zapallo na' se:k na' se:hat; na' se:hatain' gi Polygonum hispidum ta' ha Gal' che tahaGal' chehatain' gi Polygonum hydropiperoides var. hydropiperoides Polygonum punctatum ta' ha Gal' che tahaGal' chehatain' gi ta' ha Gal' che tahaGal' chehatain' gi Sapindus saponaria wedaelqo' lakik wedaelqo' laehat Sapium haematospermum ki' chiaGadik ki' chiaGadihatain' gi; ki' chiaGadih Schoenoplectus californicus na:' te na:' tehataingi Solanum glaucophyllum no' kyet no' kyede' hat; no' kyedehatain' gi Thalia geniculata pe' laq pe' laGahatain' gi Typha domingensis chi#' na chi#nahatain' gi 'naliem lodai' tehatain' gi 147 Vegetation unit categories/ Taxa included* OC-5 - lekeo' Gok OC-5.1 - Unnamed category Arrabidaea corallina Basic term Vegetation unit noqo' lo noqo' loGa' hat, noqo' loahat no' hel no' helhat Canavalia brasiliensis no' hel no' helhat Cissus sicyoides var. sicyoides nal' kyok; no' hel nal' kyoGahat; no' hel hat Clematis montevidensis nal' kyok, noqo' lo, no' helnal' kyoGahat, noqo' lohat, no' helh Cuscuta indecora var. indecora nal' kyok nal' kyoGahat Cynanchum sarcostemma Funastrum clausum kowoqo' lek qope' lañi kowoqo' le:hat qope' laGañihat Funastrum gracile kowa' ka kowaGa' hat Ipomoea bonariensis var. chacoensis nal' kyok; no' hel nal' kyoGahat; no' helhat Mascagnia brevifolia no' hel no' helhat Merremia dissecta na' qaik la' ta#a; no' hel na' qaik la' taGahat, no' helhat Mikania cordifolia noqo' lo, no' hel noqo' lohat,no' helhat Mikania periplocifolia noqo' lo noqo' lohat Morrenia grandiflora ' qalagana' Gae ' qalaganaGae' hat Morrenia odorata ' chiaGadae ' chiaGadae' hat Morrenia variegata ' talo' wai talo' waihat Passiflora foetida ' qop ' qop ' qop' qop(e)' hat Passiflora mooreana piogo' lola piogo' lola' hat Pithecoctenium cynanchoides no' hel; noqo' lo no' helhat; noqo' lohat Schubertia grandiflora yaGa' ñi yaGa' ñi' hat Schubertia schreiteri ' ta#nagana' Ga lote' te ta#nagana' Ga lote' tehat Urvillea chacoensis no' hel no' helhat OC-5.1 - Unnamed category Arrabidaea corallina noqo' lo noqo' loGahatain' gi Funastrum clausum qope' lañi qope' laGahatain' gi Funastrum gracile kowa' ka kowaGa' hataingi OC-6 - ha' palwo Phoradendron hieronymi neta' Gae la' kaik neta' Gae la' kai' hat Phoradendron liga neta' Gae la' kaik neta' Gae la' kai' hat Phoradendron pruinosum neta' Gae la' kaik neta' Gae la' kai' hat Struthanthus angustifolius neta' Gae la' kaik neta' Gae la' kai' hat Tillandsia duratii dapi' chie#e' hat Tillandsia streptocarpa dapi' chie#e, dapi' chie na' dapi'mochie#e, dapi' chie na' dapi'mochie#e, dapi' chie Tripodanthus acutifolius na' neta'mo Gae la' kaik neta' Gae la' kai' hat Usnea dichroa ' qol ' qolhat Usnea sulcata ' qol ' qolhat Tillandsia meridionalis dapi' chie#e' hat dapi' chie#e' hat 148 Vegetation unit categories/ Taxa included* Basic term Vegetation unit "Not categorized" Anemia tomentosa var. anthriscifolia na' maik la' ta#a na' mai la' ta#hat Apium leptophyllum alo' Go aloGo' hataingi, aloGo' hat Calvatia aff. cyathiformis (Fungi) waqachi' ñi la ' tek waqachi' ñi late:' hat Calvatia fragilis (Fungi) waqachi' ñi la' tek waqachi' ñi late:' hat Cheilanthes obducta na' maik la' ta#a na' mai la' ta#hat Eryngium elegans alo' Go aloGo' hat, aloGo' hataingi Erythroxylon sp. coca' BiaGahe, coca' BiaGacoca' BiaGahatain ' gi, coca' BiaGaha Flaveria bidentis qa' do:l nehe' pe:l qa' do:l nehe' pe:lhat Ganoderma aff. lucidum (Fungi) Ganoderma aff. resinaceum (Fungi) e' pak leke' tela e' pak leke' tela e' pakleke' telahat e' pakleke' telahat Heteranthera peduncularis ' molalke' tela molalke' telahatain' gi Hexagona hydnoides (Fungi) e' pak leke' tela e' pakleke' telahat Hexagona papyracea (Fungi) e' pak leke' tela e' pakleke' telahat Holmbergia tweedii wa' BiaGaik wa' BiaGaihat Hymenoxys anthemoides penaGa' di penaGa' dihat Hyptis lappacea i' diaG ata' Gaik ' lawenek i' diaGataGaik ' lawehatain' gi Iresine diffusa ka' tek leko' lak ka' tek leko' laehat Jacaratia corumbensis egea' Ga egeaGa' hat Mycenastrum corium (Fungi) waqachi' ñi la' tek waqachi' ñi late:' hat Pleurotus aff. ostreatus (Fungi) e' pak leke' atel e' pakleke' telahat Pycnoporus sanguineus (Fungi) e' pak leke' tela e' pakleke' telahat Stemodia ericifolia ' qol ' qolhatain' gi, ' qolhat Tulostoma sp. (Fungi) waqachi' ñi la' tek waqachi' ñi late:' hat Vascellum pampeanum (Fungi) waqachi' ñi la' tek waqachi' ñi la te:' hat References: * understorey plant. when they do grow together, the assigned term is halko’te -meaning “numerous” followed by the vernacular name of the plant without an added suffix. A list of these plants is given in Table 3. 4.2. Ethnotaxonomy of the vegetation units The ethnotaxonomic analysis confirmed that the Toba-Pilagá classify the vegetation units enumerated in Table 2 in different vernacular categories. Most of these are related to each other in a kind of loosely inclusive hierarchical system. The main features of those hierarchical levels are described below: 149 Table 3. Plant species not bearing a botanic collective name. Scientific name Vernacular name Battarrea stevenni (Fungi) Bougainvillea praecox ' to#olea' Gaik naco' lik Bulnesia foliosa naco' lik Capparis atamisquea el' kik la' chik Cleistocactus baumannii ' kochia# Cleome sp. paGadele' wo Cyrtopodium punctatum he' dik Echinopsis sp. po' Gok Gymnocalycium mihanovichii po:' Gok Heimia salicifolia pe' naqa' te Jatropha hieronymi poko:dik Monvillea cavendishii ' kochia# Monvillea spegazzinii na' pet, ne' pet Odontocarya salicifolia noqolo kalgueaGaik, na' yok Opuntia colubrina pichiñi Phellorinia sp. (Fungi) ' to#olea' Gaik Pluchea sagittalis o' waGaik Podaxon sp. (Fungi) ' to#olea' Gaik 4.2.1.: The vegetation units are firstly assigned to one of the three large groups of environments which the Toba-Pilagá use to indicate outstanding discontinuities of their surrounding landscape; viz.: 'βiaq (woodland), 'nonaGa (field) and ne' dep(marsh). It should be noted that these categories represent elements of the landscape rather than vegetation units. Their defining criteria are: 'βiaq ("woodland", "monte"): It approximately corresponds to the “dry semideciduous forest” of the well -known phytogeographic classifications (Cabrera, 1971; Hueck, 1978), and also to communities dominated by shrub-species. It comprises a wide spectrum of floristic compositions, and of horizontal and vertical structures, among other characteristics. The Toba-Pilagá, however, emphasize its relative visibility and accessibility as distinctive features. (It is defined as a “closed” land environment opposed to the “open” characteristic of the “field” landscape); ' nonaGa("field"): A place with high visibility and limitless accessibility (land environment defined as “open”). It includes the savanna environments, halophytic steppes, palm forests, seriously degraded areas (called "peladares"), and domestic and peridomestic areas. ne' dep("marsh"): A place with high visibility but limited accessibility (aquatic environment). It includes the great swamp (some 20 kilometers wide) which crosses the region in a NW-SE direction. 150 4.2.2.: The categories identified within the second level have a binominal structure. The first element refers to the first level categories (i.e. woodland, field or marsh), and is followed by an attributive (second element) indicating specificity (i.e. : new, modified field). Many of the expressions used to characterize ' nonaGa ' dalaGaik the first level categories refer to their spatial extension (' tadaik : large; pe' geat, ' qoGot : ). In addition, the small), and to their condition as “new” or “modified” ( ' dalaGaik woodlands were classified according to their relative height and degree of accessibility. These types of categories are frequently found in ethnoecological studies (Martin, 1974; Johnson, 2000). 4.2.3: For some categories of the second classificatory level informants named only one subcategory. However, as the latter did not encompass all the vegetation units included in the category, a complementary subcategory was assigned under the label “unnamed category”. This subcategory was however tacitly evident by opposition to that explicitly named by the native. It is recognised by the people but is described by means of a periphrasis. Berlin et al. (1974) state that in vernacular taxonomies the absence of a name for a given category does not necessarily imply the absence of the category itself. On the contrary, the existence of a defined but unnamed classificatory category (also called "covert intermediate categories") by the human group in question was frequently observed (Berlin et al., 1968; 1974; Friedberg, 1974; Martin, 1974; Shepard et al., 2001). 4.2.4.: In “woodland” as well as in “field” categories, subcategories were identified according to the location of vegetation units in high areas not liable to flooding, or in low areas which are liable to flooding. As in the previous level, generic names were not given for the subcategories, although there is clear evidence of their existence in the type of suffix used in the name given to the units each of them refer to (' hatfor those related to high areas, ' hatain' gi and ' hata' ge for those associated to low areas). 4.2.5.: In the “woodland” as well as in the “field” categories, subcategories referring to the shape of the lowlands (rounded or long), and the duration of flooding (permanent or temporary) were identified for the formations associated with low areas which are liable to flooding. As in the preceding classificatory level, subcategories bear no name, although there is evidence of them in the type of suffix used (' hatain' for gi the rounded formations; ' hata' ge for the elongated ones). 4.2.6. Classification of the vegetation by the Toba-Pilagá Based on the previous levels, there follows the elucidated classification together with a schematic illustration of the most representative categories in each environment (Fig. 2-3). 1. 'β βiaq ("Woodland", "Monte") ("BI") Land environment with poor accessibility and visibility. It comprises trees, shrubs, and their undergrowth. βiaq halo' te("extended, dense, high forest"): Tree formations of more than 1.1. 'β 5 m in height (mesophanerophytes and microphanerophytes). 151 Fig. 2. Schematic illustration of "woodland" categories identified by Toba-Pilagá people (For abbreviation of category names see text and Table 2). Fig. 3. Schematic illustration of "field", "marsh" and "woodland" categories identified by Toba-Pilagá people. "A": Plant formation along a kama:ain' gi(suffix employed: ' hatain' );gi"B": Plant formation along a kama:a' ge(suffix employed: ' hata' ).ge(For abbreviation of category names see text and Table 2). 152 1.1.A. 'β βiaq ' tadaik("large forest"; "great forest"): Tree communities with individuals more than 15 m in height (mesophanerophytes). It is defined as harco#' ge(wide, i.e. it extends over a large area). Accessibility is restricted for fear of individuals not being able to leave the forest. These forests are always located on high lands (komade' lege ). It corresponds to the "quebracho colorado and palo santo forests" sensu Cabrera (1971), Adámoli et al. (1972), and Hueck (1978). βiaq ("woodland"): Tree formation on the intermediate level of 1.1.B. 'β the altitudinal gradient, mostly composed of microphanerophytes (less than 12 m in height). It corresponds to "algarrobo forest" sensu Hueck (1978), thorn woodlands sensu Prado (1993), and "raleras" sensu (Morello & Adámoli, 1974). It differs from 1.1.A in that it is easily accessible. 1.1.B. 1. Unnamed category. It is associated with high (komade' lege ) areas not liable to flooding, and corresponds to "algarrobal-chañaral-duraznillar" sensu Adámoli et al. (1972). 1.1.B.2. Unnamed category. It is asssociated with low areas prone to flooding, and corresponds to "algarrobal-chañaral" sensu Adámoli et al. (1972). 1.1.B.2.a. Unnamed category. It is associated with permanently flooded rounded lowlands (called ka’ma:in' gi). 1.1.B.2.b. Unnamed category. It is associated with temporarily flooded elongated lowlands (called kama:' ge ). 1.1.C. Unnamed category: Trees associated exclusively with bodies of water. They correspond to gallery forests or "selvas de ribera" and "bosquecillos de madrejones" sensu Adámoli et al. (1972), on permanent water courses, and on rounded lowlands with trees taller than 12 m. 1.2. 'β βiaq ha' lokor 'β βiaq haka' lok:Low forests between 2 and 5 m high, including small trees such as Ruprechtia triflora, Acanthosyris falcata, and Ximenia americana. They have a characteristic very dense undergrowth or na' wigemeaning that people are forced to crouch (koyo' qochiñi)to enter the forest; sometimes passage is impossible (ne' hoeñi) , depending on the intensity of degradation caused by domestic animals. These forests are not necessarily associated with a body of water, and correspond to a closed low forest sensu Morello & Adámoli (1974). 1.2.A. Unnamed category. It is associated with high areas not prone to flooding, generally on sandy hills (' we#em) . 1.2.B. Unnamed category. It is associated with low areas prone to flooding. βiaq poñio' le("Woodland like a person"): Shrubs or saplings less than 2 m 1.3. 'β high. This category may vary as to their accessibility. It corresponds to shrublands. (“thorns prevent passage"): Mainly thorny shrubs. 1.3.A. yaga' ta#a' ge 1.3.A.1. Unnamed category. It is associated with high areas not liable to flooding. 1.3.A.2. Unnamed category. It is associated with low areas that are liable to flooding. 1.3.B. Unnamed category: Thornless shrubs or saplings that form dense 153 (na' wige ) stands, i.e. one must crouch (koyo' qochiñi)to enter. 1.3.B.1. Unnamed category. It is associated with high areas not liable to flooding. 1.3.B.2. Unnamed category. It is associated with low areas that are liable to flooding. 1.4. 'β βiaq ' kogotor 'β βiaq ni' yege("small woodland"): Woodland patches in open fields. This subcategory mainly refers to a spatial pattern and the site at which the forest develops. It clearly refers to the woodland-field transition. It corresponds to tree patches in ancient savannas sensu Cabrera (1971) and Morello & Adámoli (1974). βiaq ' dalaGaik. ("New woodland"): Mixed woodland or bush, 1.5. 'β plurispecific, where none of the aforementioned categories are dominant. It might correspond to a secondary forest without well defined vegetation units. Because of this it is also named wo' yema e' paq ("intermingled trees") or yale' Gat 'βiaq ("mixed woodland"). Informants refer to it as deriving from an intensely . Also called "bosques de rehache", in Spanish (Morello & exploited 'βiaq ' tadaik Saravia Toledo, 1959). 2. ' nonaGa("Field", "Open field") ("NO") Land environments with high visibility and accessibility. They include arboreal, shrubby, and herbaceous life-forms. ("Large field"). It is the proper field. It corresponds to the 2.1. ' nonaGa ' tadaik: grassland matrix of ancient savannas, palm groves, and "cardonales", among others, sensu Cabrera (1971) and Adámoli et al. (1972). 2.1.A. Unnamed category. It is associated with high areas not liable to flooding. 2.1.B. Unnamed category. It is associated with low areas that are liable to flooding. 2.1.B.1. Unnamed category. It is associated with permanently flooded rounded lowlands (kama:in' gi ). 2.1.B.2. Unnamed category. It is associated with temporarily flooded elongated lowlands (kama:' ge ). 2.2. ' nonaGa pe' geator ' nonaGa ' kogot("small field"): Peridomestic area. Small field with woodland patches or disperse trees. Located in disturbed areas with modified soils (ditches and road edges, marsh borders, stubble fields, etc.); some of its plant communities “sprout after storms” ( wa' tep qoqo' te) reported the informants. This subcategory refers mainly to a spatial pattern determined in the field-woodland transition. ("New field"): It includes vegetation units developing 2.3. ' nonaGa ' dalaGaik after a marsh has shrunk; this is the reason why it is also named ne' dep qoqo' te ("sons of the marsh"). It includes shrubs and herbaceous plants, and corresponds, in part, to halophytic steppes (sensu Cabrera, 1971). 3. ne' dep("Marsh") ("NE") Aquatic and terrestrial environments with high visibility and constrained accessibility. They are represented by large areas subjected to either permanent or temporary floods. These areas do not have clearly defined boundaries. 154 3.1. ne' dep ' tadaik("Large marsh"): Expanding large marsh. Aquatic environments. ("Small marsh"): Shrinking small marsh. Communities of 3.2. ne' dep qoqo' te herbaceous species and small shrubs colonizing areas left in the wake of a retreating marsh. Related to ' nonaGa ' dalaGaik (field-marsh transition). It refers, in part, to halophytic steppes. βiaq ' lolaGae ("Marsh or woodland saplings"): 3.3. ne' dep ' lolaGaeor 'β Communities formed by saplings of tree species growing after a shrinking marsh. It refers to the woodland-marsh transition). Three vernacular categories and their corresponding subcategories could not be included in the abovementioned system. They are: Plants associated exclusively with water. 4) no' Gop wet#' ot: Acropleustophytic plants. 4.1.) no' Gop ' weta:we: Mesopleustophytic plants. 4.2) no' Gop layo' got: 4.3) Unnamed category: Partially submerged herbaceous plants, shrubs, and trees 5) lekeo' Gok:Vines 5.1.) Unnamed category: On high and dry areas. 5.2.) Unnamed category: Associated with permanently flooded rounded lowlands. 6) ha' palwo:Epiphytic plants. The first column of Table 2 shows the inclusion of 276 vegetation units into one or more of the abovementioned categories. Categories not included within the main classificatory system are referred to as “out of classification” ("OC") (levels 4 to 6, including 70 plant units). Finally, 27 vegetation units have not been ascribed to any category; i.e. nawo' Go ' hat#aingi (tree community with conspicuous flowers, around a rounded body of water); waqachi' ñi late:' hat (group of puffball and stalked puffball fungus species); and e' paq leke' telahat (group of bracket fungus species), among others; they are referred to as "not categorized" in Table 2. 5. Discussion The results of this study were exclusively based on ethnobotanic interviews that dealt with the nomenclature and characterization of vegetation units. The actual information gathered was much wider than that presented in this article. For example, we also obtained data on types of soil, other kinds of water bodies, and faunal and supernatural beings associated with the habitats. At no time did such attributes define specific vegetation units alone, nor did they modify the proposed classificatory system. However, consideration of those features would be relevant for a classification of the overall habitat, of which the present study is just a part. 155 On the other hand, it should be noted that we have deliberately omitted a set of techniques habitually applied in ethnosciences and used in enquiries relating to classificatory systems -such as data cards, images, keys, dichotomic trees, etc. We did not use them because they have been widely criticised for introducing bias into results beforehand (Friedberg, 1968, 1974; Fowler, 1979). 5.1. The nomenclatural system of the formations and their meaning A varied number of the studies on Toba-Pilagá and Pilagá lexicons agree on the use of the suffix ' hatto exclusively denote botanical collective terms (Buckwalter, 1980; Dell' Arciprete , 1991). In these studies the suffixes ' hatain' gi and ' hata' ge are considered mere extensions of the semantic field of collective terms formed with ' hat . As recorded by De la Cruz (1993), it was noticed in the course of the present study that the latter suffix also has a more specific meaning than that given above. In fact it can be observed in Table 2 that the Toba-Pilagá make a clear distinction between the types of collectives formed by each plant species. Thus, there are species that form their collective term by adding only the suffix ' hatto the basic term (i.e. laGa' dihat , of and/or Stetsonia coryne); other species do so by adding the suffixes ' hatand ' hatain' gi (see Table 1); and still others by adding only the suffixes ' hatain' gi (i.e. ' hata' ge diki' chik ' hatain', of gi Nymphaea gardneriana). These observations led us to inquire about the possible existence of collective terms complementary to those named by only one type of suffix; e.g. a collective term formed with ' hatain' gi for S. coryne, or one formed with ' hatfor N. gardneriana. In cases such as S. coryne, the response was negative. According to the informants there are no plant units of S. coryne in low-lying flood-prone areas (“it doesn´t belong to the water”); that is th e reason why the term laGadi' hataingidoes not exist. In fact, it is related to the distinction made between high areas (komade' lege ) and low areas (kama:in' giand kama:a' ge ) in the Toba-Pilagá discourse; the suffixes which refer to low areas (ain’gi and a' ge ) are the same as those used to form collective terms for vegetation units related to low-lying flood-prone areas. However, in those cases such as N. gardneriana, informants stated that the units commonly named with the suffix ' hatain' gi (that “belong t o water”; i.e. they are not found in dry places), can be indistinctly named with the suffix ' hat . In fact, almost all the vernacular names with the suffix ' hatain' gi can also bear the suffix ' hat(except ). In this case, the suffix ' hathas diki' chik; ya' talik; to' pi; molalke' and telata' haGa l' che the general meaning of "plant collective". All in all, we can conclude from an analysis of the nomenclature and location of those units that: (a) when a species only forms collectives with ' hat , it means that the vegetation unit is located in high and dry areas; and (b) when a collective is formed with it means that the plant units are located only in low-lying areas (as is ' hatand ' hatain' , gi the case of the formations grouped in categories BI-1.1.C, NE-3.1; OC-4.1 and OC-4.3), or in high as well as in low areas (as is the case with BI-1.1.B; BI-1.2; BI-1.3.A; BI1.3.B; NO-2.1 and OC-5). 156 5.2. Vegetation units and the classification of the plant environment Although there is no information on how other Gran Chaco peoples classify their plant environment, the Toba-Pilagá’s first classificatory level of the environments (woodland - field - marsh) recalls that used by the Bororo of Southern Brazil. These people also describe their plant universe as a trichotomy: plants of the earth, plants of the water, and plants of the air (Lévi-Strauss, 1994). A similar classification of habitats is made by the Mbya-Guaraní of Paraguay. In a study of the methods employed by this latter group to classify birds, Cebolla Badie (2000) states that the first relevant division is made between birds living in fields and marshes, and those living in forests. The author also applies this kind of division to other cultural matters of the same ethnic group. On the other hand, the classificatory criteria of accessibility, visibility and height for woodlands were also found to fulfil the same functions in the ethnophytogeography of the Khmer of Cambodia and the Bunaq of Timor (the degree of visibility is sometimes expressed as a kind of gradient in vegetation density) (Martin, 1974; Friedberg, 1990). Vegetation units belonging to more than one category, together with grouping categories outside the classification, highlight the variable inclusiveness shown by the classificatory system devised for the plant environment. Other authors have also commented on this feature; e.g. Hallpike (1986) states that although indigenous classifications build hierarchical grouping systems with wide general categories divided into more specific ones, they are based on variable criteria in such a way that mutually overlapping classes coexist with classes that do not converge in any other higher category. Similar observations were made by Berlin et al. (1974), Martin (1974), and Friedberg (1990). Categories of plants based on life-form designated by Filipov (1996) and Idoyaga Molina (1988-89, 1990-91) for the Pilagás were not found to be key grouping criteria for our classification. Only vines and epyphites appeared as categories of that type, but outside the classification. The difference between those studies and ours lies in the fact that the former deal with “classes of plants” whereas we have worked with “types of vegetation units”. The third level woodland category named 'βiaq deserves a detailed analysis. This woodland may be defined as the prototype woodland or “proper” woodland, because the addition of any other descriptive adjective to it in the native language would be unnecessary. The idea of a model or prototype taxon has been described in several studies on ethnosystematics (Friedberg, 1974); it highlights the conceptual importance these types of woodlands have for this human group (in fact, this subcategory contains the tree communities of greatest economic value for them). 157 It is evident that the classification for "woodlands" was more refined than that for either "fields" or "marshes". Whereas five classificatory levels were identified in the former category, only four and two levels were found for “fields” and “marshes” respectively. Furthermore, the classification for woodlands involves the highest number of classificatory categories. However, it would be misleading to conclude that the TobaPilagá consider the "woodland" environments to be more important than the others. Actually, “field” environments have played a most remarkable role in their culture; they were very important as hunting-gathering grounds, and also as a location for the placement of their temporary settlements on the field-woodland boundaries . An explanation of the more comprehensive classification of “woodland” environments compared to “fields” might be sought in the profound ecological alterations that the region underwent in the past, particularly during the process of grassland colonization by woody species referred to earlier. Evidently the present TobaPilagá groups have no accounts of the various “field” environments of the past. The scarcity of information on the “marsh" environments (this category only comprises three subcategories) is also remarkable. Plausibly it is as a consequence of the erratic hydrological regime of the Pilcomayo river. In the past this river had a relatively steady course, and its seasonal floods extended over well delimited but restricted areas. Three decades ago, however, the river left its course, thus producing the great marsh the Toba-Pilagá know today. It is the relative novelty of this marsh environment in the regional landscape that might explain the lack of data on it. Briefly, the occurrence of large physiognomic and hydrological changes in the region during recent decades might explain why woodland, field, and marsh environments are not known to the same extent. Therefore, the characterization of transitional categories between major environments made by the informants is illustrative and interesting in itself. As in other ethnotaxonomies of the plant environment, the Toba-Pilagá referred to a series of taxa which in fact are truly “ecotones” between woodland, field and marsh. These types of Table 4. Transitional taxons. 'βiaq 'βiaq X 'nonaGa ne'dep ' nonaGa pe' geat (Peridomestic area) ne' dep ' lolaGae (tree saplings growing after a shrinking marsh) 'nonaGa ne'dep epaGa' lahoe' hat 'βiaq ni' yege (Woodland patches in (Submerged dead standing trees in marshes) open fields) X ' nonaGa ' dalaGaik (Field in an area left by a shrinking marsh) ne' dep qoqo' te (Shrinking marsh in adjacent field) X 158 categories have been identified as “node taxons” in ethnosystematic studies ( Friedberg, 1974). These taxons are enumerated in the Table 4; they are semantically closer to the elements in the rows than to those in the columns. 6. Conclusion From the results of the interviews it was established that 217 species -92 % of the species investigated- formed some type of vegetation unit, whereas the remaining 18 did not make groups of any type. Later, we were able to confirm that the Toba-Pilagá recognize a total of 304 types of vegetation units, and employ 196 vernacular terms to name them. Although the use of different suffixes generally assigns one species to more than one type of plant unit, the lower number of total collective terms in relation to the number of species they refer to is due to the fact that a particular vernacular name can commonly refer to more than one botanical species. The results of the classification of vegetation show the variety of plant-grouping levels perceived by the Toba-Pilagá. The smallest unit of any group is in all cases a small set of individuals of the same species. Species that do not form any vegetation unit are exceptional: they are perceived as strange plants that “live alone” or are “solitary”. Visibility and accessibility are two core concepts for the Toba-Pilagá classification. Tree height is also one of the basic features used for the discrimination of woodland types. The life-form of species, on the other hand, is not a criterion for grouping plants as it has often been in other ethnotaxonomies, in which plants are grouped as trees, bushes, herbs, vines, etc. (Berlin et al. 1968, 1974; Friedberg; 1990). It is important to note the clear distinction made between vegetation units living in uplands liable to flooding and those in lowlands not prone to flooding, as well as the distinction made in the latter as to their shape, and the duration of floodwater in the lowlands adjacent to the vegetation units. These nomenclatural features show that the Toba-Pilagá perceive the vegetation unit and its associated abiotic characteristics (topography, bodies of water and their shape) as a whole. These attributes are so closely interrelated for these people, that they are inseparable parts of the units' name. As an example of additional dissimilarities with scientific phytosociology, both palm and columnar cactus communities taller than 10 and 6 meters, respectively, are considered as a “field ”( ' nonaGa ). Both the categories and the classificatory system seem to be dynamic and comprehensive enough to account for both temporary transitional environments and important ecological and hydrological changes which historically have affected the physiognomy of the regional landscape. This dynamism is reflected in the nomenclature as well as in the inclusion of relatively new environments in their classificatory system (see Table 4). The results of this study are relevant for a number of reasons. Firstly, the ethnophytogeographic perspective is original as a subject of study because it has not been described for any other ethnic group in the South American Gran Chaco. 159 Secondly, it significantly contributes to elucidating how the Toba Pilagá relate to their plant environment; a very important ethnological issue indeed. Finally, the ethnophytogeographic approach lays down the bases for undertaking an exhaustive comparison with the scientific phytosociology of the region, particularly with regard to the relationship between phytocoenosis, fluviogeomorphology, and plant succession. This information would undoubtedly be of great value for understanding the ecological dynamics of a region which has been subjected to intense environmental fluctuations during the past century. Acknowledgements. We thank Daniel Ginzo, Haydée González and Lawrence Wheeler for their help in preparing the manuscript. 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