Catalogue of Diptera of Colombia

Silvio Nihei

The present catalogue summarizes 154 species and 71 genera occurring in Colombia. All the four subfamilies (Phasiinae, Dexiinae, Tachininae and Exoristinae), and 24 tribes are represented in the Colombian territory.

This considerably small family is poorly known in Colombia, with only two species reported for the genus Sylvicola Harris (1776) so far. We synonymize Neomesochria Amorim & Tozoni (1994) to Mycetobia Meigen (1818), hence transferring the Dominican amber species Neomesochria antillea (Grimaldi 1991) and N. cryptambra (Grimaldi 1991), and the recent Neotropical species N. limanda (Stone 1966) and N. stonei (Lane & d’Andretta 1958) back to the genus Mycetobia. This paper provides new records for Mycetobia and Olbiogaster Osten-Sacken (1886) for Colombia.

The Bibionidae are a family belonging to the suborder Bibionomorpha with four genera and 17 species known from Colombia. This work expands the distribution of these species to other localities in the country.

Zootaxa 4122 (1): 001–949 http://www.mapress.com/j/zt/ Copyright © 2016 Magnolia Press Monograph ISSN 1175-5326 (print edition) ZOOTAXA ISSN 1175-5334 (online edition) http://doi.org/10.11646/zootaxa.4122.1.1 http://zoobank.org/urn:lsid:zoobank.org:pub:8923AFC4-7EED-45EC-AE5F-36D2082FB653 ZOOTAXA 4122 Catalogue of Diptera of Colombia Edited by MARTA WOLFF, SILVIO S. NIHEI & CLAUDIO J. B. DE CARVALHO Magnolia Press Auckland, New Zealand Published: 14 Jun. 2016 Licensed under a Creative Commons Attribution License http://creativecommons.org/licenses/by/3.0 MARTA WOLFF, SILVIO S. NIHEI & CLAUDIO J. B. DE CARVALHO (EDS.) Catalogue of Diptera of Colombia (Zootaxa 4122) 949 pp.; 30 cm. 14 Jun. 2016 ISBN 978-1-77557-961-8 (paperback) ISBN 978-1-77557-962-5 (Online edition) FIRST PUBLISHED IN 2016 BY Magnolia Press P.O. Box 41-383 Auckland 1346 New Zealand e-mail: magnolia@mapress.com http://www.mapress.com/j/zt/ © 2016 Magnolia Press ISSN 1175-5326 (Print edition) ISSN 1175-5334 (Online edition) 2 · Zootaxa 4122 (1) © 2016 Magnolia Press Zootaxa 4122 (1): 003–007 http://www.mapress.com/j/zt/ Table of contents Copyright © 2016 Magnolia Press ISSN 1175-5326 (print edition) ZOOTAXA ISSN 1175-5334 (online edition) http://doi.org/10.11646/zootaxa.4122.1.2 Catalogue of Diptera of Colombia MARTA WOLFF, SILVIO S. NIHEI & CLAUDIO J. B. DE CARVALHO (EDS.) 8 Catalogue of Diptera of Colombia: an introduction MARTA WOLFF, SILVIO S. NIHEI & CLAUDIO J. B. DE CARVALHO 15 Family Anisopodidae DALTON DE SOUZA AMORIM, RAFAELA LOPES FALASCHI & SARAH SIQUEIRA OLIVEIRA 20 Family Bibionidae RAFAELA LOPES FALASCHI, SARAH SIQUEIRA OLIVEIRA & DALTON DE SOUZA AMORIM 26 Family Cecidomyiidae VALÉRIA CID MAIA 41 Family Sciaridae SHEILA PATRÍCIA CARVALHO-FERNANDES 46 Family Rangomaramidae RAFAELA LOPES FALASCHI & DALTON DE SOUZA AMORIM 50 Family Ditomyiidae RAFAELA LOPES FALASCHI 53 Family Diadocidiidae RAFAELA LOPES FALASCHI 56 Family Keroplatidae RAFAELA LOPES FALASCHI 62 Family Lygistorrhinidae SARAH SIQUEIRA OLIVEIRA & DALTON DE SOUZA AMORIM 65 Family Mycetophilidae SARAH SIQUEIRA OLIVEIRA & DALTON DE SOUZA AMORIM 73 Families Tipulidae and Limoniidae GUILHERME C. RIBEIRO & DAUBIAN SANTOS 98 Family Ceratopogonidae GUSTAVO SPINELLI & MARTA WOLFF 142 Family Chironomidae HUMBERTO FONSECA MENDES & LUIZ CARLOS PINHO Published: 14 Jun. 2016 Licensed under a Creative Commons Attribution License http://creativecommons.org/licenses/by/3.0 3 154 Family Simuliidae MARTA I. WOLFF E, DANIEL RAFAEL MIRANDA-ESQUIVEL & LIGIA INES MONCADA-ALVAREZ 178 Family Corethrellidae LEONARDO H. GIL-AZEVEDO, CAIO C. D. CORRÊA & ART BORKENT 182 Family Blephariceridae LEONARDO H. GIL-AZEVEDO & DURVAL DA S. SANTOS 187 Family Psychodidae EDUAR ELÍAS BEJARANO & LUIS GREGORIO ESTRADA 239 Family Scatopsidae DALTON DE SOUZA AMORIM 246 Family Rhagionidae CHARLES MORPHY D. SANTOS & DANIEL D. D. CARMO 249 Family Tabanidae MARTA WOLFF E & DANIEL RAFAEL MIRANDA-ESQUIVEL 302 Family Xylophagidae DIEGO AGUILAR FACHIN 306 Family Pantophthalmidae MARTA WOLFF, YARDANY RAMOS-PASTRANA & FERNANDO VALLEJO 312 Family Stratiomyidae DIEGO AGUILAR FACHIN & CRISTIANE VIEIRA DE ASSIS-PUJOL 342 Family Xylomyidae DIEGO AGUILAR FACHIN & CRISTIANE VIERA DE ASSIS-PUJOL 346 Family Acroceridae JÉSSICA PAULA GILLUNG 350 Family Nemestrinidae CECÍLIA KOSMANN & CARLOS JOSÉ EINICKER LAMAS 353 Family Asilidae MARTA WOLFF & CARLOS JOSÉ EINICKER LAMAS 372 Family Bombyliidae CARLOS JOSÉ EINICKER LAMAS & NEAL L. EVENHUIS 382 Family Mydidae JÚLIA CALHAU & CARLOS JOSÉ EINICKER LAMAS 387 Family Therevidae PAULA FERNANDA MOTTA RODRIGUES & JULIA CALHAU 393 Family Empididae JOSÉ ALBERTINO RAFAEL & JOSENIR TEIXEIRA CÂMARA 4 · Zootaxa 4122 (1) © 2016 Magnolia Press WOLFF ET AL. (EDS.) 397 Family Hybotidae ROSALY ALE-ROCHA & RAFAEL AUGUSTO PINHEIRO DE FREITAS-SILVA 404 Family Dolichopodidae RENATO SOARES CAPELLARI 414 Family Phoridae DANILO CÉSAR AMENT & BRIAN V. BROWN 452 Family Pipunculidae JOSÉ ALBERTINO RAFAEL & HUGO CESAR RODRIGUEZ 457 Family Syrphidae AUGUSTO L. MONTOYA 538 Family Micropezidae STEPHEN MARSHALL, ANDERSON CARDONA & MARTHA WOLFF 549 Family Neriidae TATIANA SEPÚLVEDA & CLAUDIO J. B. DE CARVALHO 555 Family Tanypezidae MARTA WOLFF & DIANA GRISALES 558 Family Syringogastridae ALESSANDRA RUNG 561 Family Conopidae LEONARDO SILVESTRE GOMES ROCHA & CÁTIA ANTUNES DE MELLO-PATIU 566 Family Lonchaeidae JULIANA MORGADO FERNANDES & MÁRCIA SOUTO COURI 576 Family Piophilidae MARTA WOLFF, SANDRA PÉREZ & DIANA GRISALES 579 Family Platystomatidae LISIANE DILLI WENDT 582 Family Pyrgotidae RAMON LUCIANO MELLO & CARLOS JOSÉ EINICKER LAMAS 585 Family Richardiidae LISIANE DILLI WENDT & ROSALY ALE-ROCHA 596 Family Tephritidae MARCOANDRE SAVARIS, LUCIANE MARINONI & ALLEN L. NORRBOM 622 Family Lauxaniidae VERA CRISTINA SILVA 635 Family Ropalomeridae ROSALY ALE-ROCHA Zootaxa 4122 (1) © 2016 Magnolia Press · 5 642 Family Sciomyzidae LUCIANE MARINONI & WILLIAM L. MURPHY 648 Family Sepsidae VERA CRISTINA SILVA 653 Family Clusiidae MARTA WOLFF & DIANA GRISALES 659 Family Agromyzidae VIVIANE RODRIGUES DE SOUSA & MÁRCIA SOUTO COURI 677 Family Periscelididae ALESSANDRA RUNG 681 Family Aulacigastridae ALESSANDRA RUNG 685 Family Sphaeroceridae KARLA P. TEPEDINO 696 Family Chloropidae PAULA RAILE RICCARDI 708 Family Milichiidae JOHN SWANN 716 Family Curtonotidae RAMON LUCIANO MELLO 719 Family Drosophilidae ROSANA TIDON & JONATHAN MENDES DE ALMEIDA 752 Family Ephydridae WAYNE NEILSEN MATHIS, FRANCISCO DE ASSIS RODRIGUES JÚNIOR & MÁRCIA SOUTO COURI 771 Family Hippoboscidae GUSTAVO GRACIOLLI 780 Family Nycteribiidae GUSTAVO GRACIOLLI, CARL W. DICK & RICARDO GUERRERO 784 Family Streblidae CARL W. DICK, GUSTAVO GRACIOLLI & RICARDO GUERRERO 803 Family Anthomyiidae DIANA GRISALES, ANDRÉ CÉSAR LOPES & CLAUDIO J. B. DE CARVALHO 807 Family Fanniidae DIANA GRISALES & CLAUDIO J. B. DE CARVALHO 814 Family Muscidae SANDRA PÉREZ & CLAUDIO J. B. DE CARVALHO 6 · Zootaxa 4122 (1) © 2016 Magnolia Press WOLFF ET AL. (EDS.) 854 Family Scathophagidae MARTA WOLFF, DIANA GRISALES & SILVIO S. NIHEI 856 Families Calliphoridae and Mesembrinellidae MARTA WOLFF & CECILIA KOSMANN 876 Family Oestridae MARTA WOLFF & DIANA GRISALES 881 Family Rhinophoridae SILVIO S. NIHEI 884 Family Sarcophagidae CÁTIA ANTUNES DE MELLO-PATIU 904 Family Tachinidae SILVIO S. NIHEI Zootaxa 4122 (1) © 2016 Magnolia Press · 7 Zootaxa 4122 (1): 008–014 http://www.mapress.com/j/zt/ Copyright © 2016 Magnolia Press Preface ISSN 1175-5326 (print edition) ZOOTAXA ISSN 1175-5334 (online edition) http://doi.org/10.11646/zootaxa.4122.1.3 http://zoobank.org/urn:lsid:zoobank.org:pub:EDA56EB2-BD51-4700-8077-4AF473EB4B29 Catalogue of Diptera of Colombia: an introduction MARTA WOLFF1, SILVIO S. NIHEI2 & CLAUDIO J. B. DE CARVALHO4 1 Grupo de Entomologia, Instituto de Biología, Universidad de Antioquia, Colombia. E-mail: martha.wolff@udea.edu.co Department of Zoology, Institute of Biosciences, Universidade de São Paulo, Rua do Matão, Travessa 14, n.101, CEP 05508-090, São Paulo, Brazil. E-mail: silvionihei@gmail.com 4 Department of Zoology, Universidade Federal do Paraná, Postal box 19020, Curitiba, 81531–580, Brazil. E-mail: cjbcarva@ufpr.br 2 Colombia has an imposing natural wealth due to its topography has many unique characteristics as a consequence of having Caribbean and Pacific shores, as well as sharing part of the Amazon basin and northern Andes mountains. Thus, many natural and biological features are due to the convergence of three biogeographical regions: Pacific, Andes and Amazonia. The Andean uplift created a complex mosaic of mountains and isolated valleys, including eleven biogeographical provinces (Morrone 2006). The Andes dominate the Colombian topography and cross the country south to north. There are three mountain ranges (Western, Central, and Eastern) with a maximum elevation of 5,775 m, and an average elevation of 2,000 m. The Magdalena and Cauca River valleys separate these ranges, that along with the Putumayo and Caquetá Rivers, the Catatumbo watershed, the Darién, Pique Hill, the Orinoquia Region (with its savannas), the Amazon region (with tropical rainforests), and some lower mountain ranges (Macarena and Chiribiquete), have generated the conditions for very high levels of endemism. This variety of conditions has resulted in an extremely diverse plant and animal biota, and in which 48% of the nation remains unexplored. Biodiversity in Colombia is a world treasure, and therefore understanding it is the first step in preserving it. Understanding insect diversity is a challenge, due to the complexity of finding species in such a diverse biological and geological settings, as well as due to the difficulty in identifying the many, often undescribed, species. Historically, Colombia has had many entomologists that have strived towards documenting this diversity and today, continues to education new researchers for this far-from-finished goal. Though there is a long tradition of entomological studies, taxonomic knowledge of insects in Colombia remains fragmented, with the Lepidoptera probably being the best known. Much work remains for the identification of most species in the Coleoptera, Hymenoptera and Diptera. These orders comprise nearly 70% of all known animal species in the country (Andrade, 1996, Amat et al. 1999, Fernández et al. 2004). Diptera in Colombia has mostly been studied in species that are relevant to health and forensic concerns, such as the Culicidae, Simuliidae, Tabanidae, and Calliphoridae. The more conspicuous flies, and larger families, including the Syrphidae, Asilidae and Bombyliidae, remain poorly known and poorly studied. Recently, efforts to collect in relatively undisturbed locations may be one of the main initiatives by the government and universities for better understanding these well-preserved areas and for conservation planning. Here, we provide a catalogue with the primary goal of providing information on Diptera diversity in Colombia. This catalogue is based on a combination of a review of the relevant literature with data from new species that will be presented here for the first time. All genus- and species-group names are listed alphabetically. Valid names are in bold type at the first mention. Synonyms when mentioned subsequently are italicized. Each published name including synonyms is given in a chronological sequence according to the scheme: the name of the species, author(s), year of description, page, genus in the original description (if different from the valid one), type-locality (or localities in the case of syntypes; updated name into brackets), information about the primary types and their deposition (and, if a lectotype, a reference to the author and date of the designation), known geographic distribution. The catalogue contains 3,135 species in 74 families from Colombia (Table 1). This adds another nine families to the Catalogue of the Diptera of the Americas South of the United States between 1966-1984 (edited by Nelson Papavero). Unfortunately, and highlighting the need for additional collecting, the families Dixidae, 8 Accepted by Z.-Q. Zhang: 26 May 2015; published: 14 Jun. 2016 Licensed under a Creative Commons Attribution License http://creativecommons.org/licenses/by/3.0 Chaoboridae, Athericidae, Vermileonidae, Platypezidae, Ulidiidae and Heleomyzidae cannot be included in this catalogue, due to the lack of specialists who can identify species in these families. Also, the Culicidae, with many records in the country, is also not included in this catalogue because of its taxonomic complexity. FIGURE 1. Map of Colombia, with the geopolitical departments. Chapters of this catalogue are organized into an abstract with key words, a brief introduction, the list of species with information on valid names and synonyms for Colombian species, type locality, museum where types are deposited (when available), localities where collected in Colombia and references. Finally, we provide a table that summarizes subfamily and tribe (if applicable), genus, species and author of species and year. CATALOGUE OF DIPTERA OF COLOMBIA: AN INTRODUCTION Zootaxa 4122 (1) © 2016 Magnolia Press · 9 TABLE 1. Summary of Diptera families in Colombia: Number of species in Colombia (based on present catalogue), Number of species in Neotropical region (based on literature), and the Distribution of each family currently known for the Colombian Departments. Family 1. Anisopodidae 2. Bibionidae 3. Cecidomyiidae 4. Sciaridae 5. Rangomaramidae 6. Ditomyiidae 7. Diadocidiidae 8. Keroplatidae 9. Lygisthorrhinidae Colombian species Neotropical species 2 631 17 44 16 Ant, Cun 2 Bog, Met, VCau, 2 Ces, Mag, Vau 2 Bog, Cal, Caq, Cun, Mag. 169 360 193 20 3 Ris 6 undescr 44 2 Nar, NSant 1 undescr 2 1 5 1 undescr 2 Ris 2 151 7 2 Cau, Met, Nar, VCau Vau 2 10. Mycetophilidae 12 991 11a. Tipulidae 38 7182 11b. Limoniidae Distribution in Departments 2 Ama, Bol, Boy, Cau, Cho, Cun, Hui, Mag, Met, Nar, NSant, Put, Vau. Bol, Bog, Boy, Cal, Cau, Ces, Cun, Mag, Met, Tol, VCau, 93 2369 Boy, Cal, Cau, Cho, Cun, Mag, Met, Tol, Gorg Is. 235 6902 Ama, Ant, Ara, Atl, Boy, Caq, Cau, Ces, Cho, Cun, Guj, Guv, Hui, Mag, Met, Nar, Put, Ris, Tol, VCau, Vau, 13. Chironomidae 30 2052 Met. (without others specifics localities) 14. Simuliidae 67 3484 Ama, Ant, Boy, Cal, Cau, Ces, Cun, Guj, Hui, Mag, Met, Nar, NSant, Qui, Tol, Vau, VCau 15. Corethrellidae 9 442 Ama, Boy, Cor, Cun, Met, VCau 16. Blephariceridae 13 632 Ant, Cal, Cun, Guj, Mag, Qui, Tol, VCau 17. Psychodidae 199 3702 All departments (32) 18. Scatopsidae 2 625 VCau 3 6 19. Rhagionidae 20. Tabanidae 21. Xylophagidae 22. Panthophthalmidae 97 ** 7 255 1200 1 282 11 20 Ama, Ant, Atl, Bol, Boy, Cal, Caq, Cas, Cau, Cho, Cor, Cun, Hui, Mag, Met, Nar, Put, Sant, Tol, VCau, Vau Cho 8 Ama, Ant, Boy, Cal, Cho, Cun, Met, Put, Sant, VCau, 7 23. Stratiomyidae 86 987 24. Xylomyidae 1 92 Nar, VCau 25. Acroceridae 4 * Cun 26. Nemestrinidae 27. Asilidae 28. Bombyliidae 29. Mydidae 30. Therevidae 31. Empididae 32. Hybotidae 33. Dolichopodidae 1 72 22 2 4 4 19 29 64 9 Ama, Ant, Atl, Boy, Cal, Cho, Cun, Gorg Is, Mag, Met, Nar, VCau Mag 10 1576 Ama, Ant, Bog, Cal, Cau, Ces, Cun, Cho, Mag, Met, VCau 472 11 ** 116 12 Boy 2 Bol, Mag 500 13 Cau 250 14 Ant, Caq, Hui, Met, Nar, Put, Tol, VCau 147 15 1195 Ant, Atl, Boy, Cau, Cun, Gorg Is, Mag, Met, VCau ...Continued on next page 10 · Zootaxa 4122 (1) © 2016 Magnolia Press WOLFF ET AL. (EDS.) TABLE 1. (Continued) Colombian species Neotropical species 34. Phoridae 225 * 35. Pipunculidae 10 2507 Family 36. Syrphidae 312 Distribution in Departments Ama, Bol, Boy, Caq, Cau, Cho, Cun, Gorg Is, Hui, Mag, Met, Nar, NSant, Put, Ris, Suc, VCau, Vau, Vich 2 1637 2 Ama, Cau, Cun, Nar, Tol, VCau 31 dptos excepto Guv 37. Micropezidae 57 273 Ama, Ant, Atl, Bol, Boy, Caq, Cun, Mag, Met, Nar, Put, Tol, VCau, Vich 38. Neriidae 17 392 Ama, Ant, Atl, Bol, Boy, Cal, Caq, Cau, Cho, Cun, Guj, Hui, Mag, Met, Nar, NSant, Put, Qui, Ris, Sant, Suc, VCau, Vau, Vich 39. Tanypezidae 9 192 Ant, Boy, Cun 40. Syringogastridae 3 9 2 Ama, Cal, Cor, VCau 2 41. Conopidae 16 175 42. Lonchaeidae 44 * 43. Piophilidae 44. Platystomatidae 45. Pyrgotidae 46. Richardiidae 2 1 1 19 Met, VCau Ant, Cau, Cun, NSant, Sant, Tol, VCau 10 7 Ant, Boy, Cun, Ris 26 2 ** 42 2 Ant, Cun 2 Ant, Cal, Cun, Met, Qui, Ris, Sant, Suc, VCau 2 170 47. Tephritidae 93 675 Ama, Ant, Bol, Boy, Cal, Cau, Cor, Cun, NSant, Put, Qui, Sant, Tol, VCau 48. Lauxaniidae 36 3672 Ama, Bol, Boy, Cau, Cho, Cun, Hui, Mag, Met, Nar, NSant, Put, Ris, Tol, Vau, Vich 49. Ropalomeridae 8 3116 Ant, Atl, Bol, Boy, Cal, Caq, Cau, Cho, Hui, Mag, Met, Suc 7 74 2 Cun, Nar, Put, VCau 26 2 Mag 95 2 Ama, Ant, Cun, Mag, Met, Tol, Vau, VCau 17 Ama, Ant, Cal, Cau, Ces, Cun, Met, Nar, VCau 2 Cun, VCau 50. Sciomyzidae 51. Sepsidae 52. Clusiidae 53. Agromyzidae 54. Periscelididae 55. Aulacigastridae 56. Sphaeroceridae 57. Chloropidae 58. Milichiidae 59. Curtonotidae 10 23 74 300 3 14 1 2 25 46 6 3 2 VCau 2 Ama, Ant, Cun, Mag, Met, NSant, VCau 2 Atl, Cal, Cun, Mag 174 430 66 2 VCau 21 2 ** 2 60. Drosophilidae 176 694 Ama, Ant, Atl, Cal, Cau, Cho, Cun, Guj, Hui, Mag, Met, Nar, NSant, Sant, VCau 61. Ephydridae 53 2812 Ant, Atl, Bol, Cau, Cun, Mag, Tol, VCau, 2 62. Hippoboscidae 18 44 63. Nycteribiidae 8 372 7 Ant, Atl, Bog, Bol, Boy, Caq, Cau, Cun, Hui, Mag, Met, NSant, Sant, VCau, Ant, Bol, Caq, Cun, Cho, Met, Put, Sant, Tol 64. Streblidae 73 152 Ant, Bol, Cal, Caq, Cas, Cau, Cor, Cun, Cho, Gui, Hui, Mag, Met, Nar, NSant, Put, Sant, Tol, VCau, Vau, Vich 65. Anthomyiidae 6 1082 Cun 66. Fanniidae 32 79 7 Ama, Ant, Boy, Cal, Cun, Cho, Mag, Met, NSant, Ris, Sant ...Continued on next page CATALOGUE OF DIPTERA OF COLOMBIA: AN INTRODUCTION Zootaxa 4122 (1) © 2016 Magnolia Press · 11 TABLE 1. (Continued) Family 67. Muscidae 68. Scatophagidae Colombian species Neotropical species 108 84618 1 619 Distribution in Departments Ama, Ant, Bol, Boy, Cal, Caq, Cas Cau, Ces, Cor, Cun, Cho, Guj, Hui, Mag, Met, Nar, NSant, Put, Qui, Ris, Sant, Suc, Tol, VCau, Vich Ant 2 69. Calliphoridae (includes Mesembrinellinae) 52 126 Ama, Ant, Atl, Boy, Cal, Caq, Cau, Cho, Cun, Guj, Hui, Mag, Met, Nar, NSant, Put, Qui, Ris, Sant, Suc, Tol, VCau, Vau, Vich. 70. Oestridae (Cuterebridae) 3 3720 Ant, Cun, Hui, 71. Rhinophoridae 1 22 Mag 7 72. Sarcophagidae 102 750 73. Tachinidae 154 286421 Ama, Ant, Boy, Cal, Cau, Cho, Cun, Gor Is, Mag, Met, Nar, Put, Qui, Sant, Suc, Tol, VCau, Vau, Vich Ant, Boy, Cal, Cun, Mag, VCau, * Species not catalogued. ** Without specific localities. Abbreviations used for Colombian Departments: Ama=Amazonas, Ant=Antioquia, Ara=Arauca, Atl=Atlántico, Bol=Bolivar, Boy=Boyacá, Cal=Caldas, Caq=Caquetá, Cas=Casanare, Cau=Cauca, Ces=Cesar, Cho=Chocó, Cor=Córdoba, Cun=Cundinamarca, Gorg Is= Isla Gorgona, Gui=Guainía, Guj=Guajira, Guv=Guaviare, Hui=Huila, Mag=Magdalena, Met=Meta, Nar=Nariño, NSant=Norte de Santander, Put=Putumayo, Qui=Quindío, Ris=Risaralda, Sant=Santander, Suc=Sucre, Tol=Tolima, VCau=Valle del Cauca, Vau=Vaupés, Vich=Vichada. References cited for number of Neotropical species: 1, Hancock & Amorim, 2009; 2, Amorim et al. 2002; 3, Amorim & Falaschi, 2012; 4, Coscaron, Coscaron-Arias & Papavero, 2008; 5, Amorim 2009; 6, Santos 2008; 7, de Carvalho et al. 2012; 8, Papavero, 2009a; 9, Papavero & Benardi, 2009; 10, Papavero, 2009b, 11, Evenhuis & Greathead, 1999; 12, Papavero, 2009c; 13, Yang et al., 2007; 14, Smith, 1967; 15, Yang et al 2006; 16, Prado & Papavero, 2009; 17, Spencer, 1984; 18, de Carvalho et al., 2005; 19, Albuquerque, 1984; 20, Papavero & Guimarães, 2009; 21, Guimarães, 1971. The editors would like to thank the many friends and colleagues who contributed to these chapters and this catalogue. We greatly appreciate their work, dedication and generosity to this task. Thanks to Carolina Henao Sepulveda for her support in the early reviews. MW thanks the Universidad de Antioquia “Proyecto Sostenibilidad 2012-2013 y 2014-2015”, and the Fundação de Apoio à Pesquisa do Distrito Federal (FAPDF), to Instituto de Ciências Biológicas da Universidade de Brasília (UnB). SSN thanks the financial support from FAPESP (proc. n. 2013/05131–6). CJBC is grateful to the “Conselho Nacional de Desenvolvimento Científico e Tecnológico” (CNPq) for fellowship and grant (process number 304713/2011-2). References Albuquerque, D.O. (1968) 96B. Family Scatophagidae. In: Papavero, N. (Ed.), A catalogue of the Diptera of the Americas South of the United States. Museu de Zoologia (Universidad de São Paulo), São Paulo, 4 pp. Amat, G., Andrade, G. & Fernández, F (1999) Insectos de Colombia, Vol II. Academia Colombiana de Ciencias Exactas, Fisicas y Naturales, Coleccion Jorge Alvarez Lleras Nº 13, Bogota, Colombia, 433 pp. Amorim, D.S. (2009) Catalogue of Neotropical Diptera. Scatopsidae. Neotropical Diptera, Ribeirão Preto, 4, 1–18. Amorim, D.S., Silva, V.C. & Balbi, M.I.P.A. (2002) Estado de conhecimento dos Diptera neotropicais. In: C. Costa, S.A. Vanin, J.M. Lobo & A. Melic. (Eds.), Proyeto de Red Iberoamericana de Biogeografia y Entomología sistemática PrIBES 2002. 1a. ed. Vol. 2. Sociedad Entomológica Aragonesa, Zaragoza, pp. 29–36. Amorim, D.S. (2012) Catalogue of Neotropical Diptera. Rangomaramidae. Neotropical Diptera, Ribeirão Preto, 21, 1–7. Andrade, G., Amat, G. & Fernández, F. (1996) Insectos de Colombia, Estudios escogidos. Academia Colombiana de Ciencias Exactas, Fisicas y Naturales, Coleccion Jorge Alvarez Lleras Nº 13, Bogota, Colombia, 541 pp. de Carvalho, C.J.B., Couri, M.S., Pont, A.C., Pamplona, D.M. & Lopes, S.M. (2005) A Catalogue of the Muscidae (Diptera) of the Neotropical Region. Zootaxa, 860, 1–282. de Carvalho, C.J.B., Rafael, J.A., Couri, M.S. & Silva, V. (2012) 40. Diptera. In: Rafael, J.A., Melo, G.A.R., Casari, S. & 12 · Zootaxa 4122 (1) © 2016 Magnolia Press WOLFF ET AL. (EDS.) Costantino, R. (Eds), Insetos do Brasil, Diversidad e Taxonomia. Holos Editora, Ribeirão Preto, pp. 701–743. Coscarón, S., Coscarón-Arias, M.C. & Papavero, N. (2008) Catalogue of Neotropical Diptera. Simuliidae. Neotropical Diptera, Ribeirão Preto, 2, 1–90. Evenhuis, N.L. & Greathead, D.J. (1999) World Catalog of Bee Flies (Diptera: Bombyliidae). Backhuys Publishers, Leiden, 756 pp. Fernández, F., Andrade, G. & Amat, G. (2004) Insectos de Colombia, Vol III. Universidad Nacional de Colombia, Bogota, Colombia, 602 pp. Guimarães, J.H. (1971) 104. Family Tachinidae. In: Papavero, N. (Ed.), A catalogue of the Diptera of the Americas South of the United States. Museu de Zoologia (Universidad de São Paulo), São Paulo, 333 pp. Hancock, G. & Amorim, D.S. (2009) Anisopodidae. In: Brown, B., Borkent, A., Cumming, J.M., Wood, D.M. & Zumbado, M.A. (Eds.), Manual of Central American Diptera. Vol. 1. NRC Research Press, Ottawa, pp. 341–345. Morrone, J.J. (2006) Biogeographic areas and transition zones of Latin America and the Caribean Islands based on panbiogeographic and cladistics analyses of the entomofauna. Annual Review of Entomology, 51, 467–494. Papavero, N. (2009a) Catalogue of Neotropical Diptera. Pantophthalmidae. Neotropical Diptera, Ribeirão Preto, 19, 1–11. Papavero, N. (2009b) Catalogue of Neotropical Diptera. Asilidae. Neotropical Diptera, Ribeirão Preto, 17, 1–178. Papavero, N. (2009c) Catalogue of Neotropical Diptera. Mydidae. Neotropical Diptera, Ribeirão Preto, 14, 1–31. Papavero, N. & Bernardi, N. (2009) Catalogue of Neotropical Diptera. Nemestrinidae. Neotropical Diptera, Ribeirão Preto, 7, 1–16. Papavero, N . & Guimarães, J.H. (2009) Catalogue of Neotropical Diptera. Cuterebridae. Neotropical Diptera, Ribeirão Preto, 11, 1–17. Prado, A.P. & Papavero, N. (2009) Catalogue of Neotropical Diptera. Ropalomeridae. Neotropical Diptera, Ribeirão Preto, 13, 1–8. Santos, C.M.D. (2008) Geographical distribution of Tabanomorpha (Diptera, Brachycera): Athericidae, Austroleptidae, Oreoleptidae, Rhagionidae, and Vermileonidae. EntomoBrasilis, 1, 43–50. Smith, K.G.V. (1967) 39. Family Empididae (Empidae, Hybotidae). In: Papavero, N. (Ed.), A catalogue of the Diptera of the Americas South of the United States. Museu de Zoologia (Universidad de São Paulo), São Paulo, 67 pp. Spencer, K.A. (1984) The Agromyzidae (Diptera) of Colombia, including a new species attacking potato in Bolivia. Revista Colombiana de Entomologia, 10, 3–33 Yang, D., Zhu, Y., Wang, M. & Zhang, L. (2006) World Catalog of Dolichopodidae (lnsecta: Diptera). China Agricultural University Press, Beijing, 740 pp. Yang D., Zhang, K., Yao, G. & Zhang, J. (2007) World Catalog of Empididae (lnsecta: Diptera). China Agricultural University Press, Beijing, 599 pp. List of Contributing Authors Alessandra Rung. California Department of Food and Agriculture, U.S.A. Allen L. Norrbom. United States Department of Agriculture, Agricultural Research, U.S.A. Anderson Cardona. Universidad de Antioquia, Colombia. André César Lopes. Universidade Federal do Paraná, Brazil. Art Borkent. Royal British Columbia Museum, American Museum of Natural History, and Instituto Nacional de Biodiversidad, Canada Augusto L. Montoya. Universidad de Antioquia, Colombia. Brian V. Brown. Natural History Museum of Los Angeles County, U.S.A. Caio C. D. Corrêa. Museu Nacional, Universidade Federal do Rio de Janeiro, Brazil. Carlos José Einicker Lamas. Museu de Zoologia da Universidade de São Paulo, Brazil Carl W. Dick. Western Kentucky University, U.S.A. Cátia Antunes de Mello-Patiu. Museu Nacional, Universidade Federal do Rio de Janeiro, Brazil. Cecília Kosmann. Universidade Estadual Paulista, Brazil. Charles Morphy D. Santos. Universidade Federal do ABC, Brazil. Claudio J. B. de Carvalho. Universidade Federal do Paraná, Brazil. Cristiane Vieira de Assis Pujol-Luz. Universidade Católica de Brasília, Brazil. Dalton de Souza Amorim. Universidade de São Paulo, Brazil. Daniel D. D. Carmo. Universidade de São Paulo, Brazil. Daniel Rafael Miranda-Esquivel. Universidad Industrial de Santander, Colombia. Danilo César Ament. Universidade de São Paulo, Brazil. Daubian Santos. Universidade Federal do ABC. Brazil. CATALOGUE OF DIPTERA OF COLOMBIA: AN INTRODUCTION Zootaxa 4122 (1) © 2016 Magnolia Press · 13 Diana Grisales. Universidade Federal do Paraná, Brazil. Diego Aguilar Fachin. Universidade de São Paulo, Brazil. Durval da S. Santos. Museu Nacional, Universidade Federal do Rio de Janeiro, Brazil. Eduar Elías Bejarano. Universidad de Sucre, Universidad de Cartagena, Colombia. Fernando Vallejo. Universidad de Caldas, Colombia. Francisco de Assis Rodrigues Júnior. Museu Nacional, Universidade Federal do Rio de Janeiro, Brazil. Guilherme C. Ribeiro. Universidade Federal do ABC, Brazil. Gustavo Graciolli. Universidade Federal de Mato Grosso do Sul, Brazil. Gustavo Spinelli. Museo de La Plata, Argentina. Hugo Cesar Rodriguez. Laboratorio de Enemigos Naturales de Plagas Agrícolas del NOA, Argentina. Humberto Fonseca Mendes. Universidade Federal do ABC, Brazil. Jéssica Paula Gillung. University of California Davis, U.S.A. Julia Calhau. Museu de Zoologia da Universidade de São Paulo, Brazil. Juliana Morgado Fernandes. Museu Nacional, Universidade Federal do Rio de Janeiro, Brazil. John Swann. University of Calgary, Canada. Jonathan Mendes de Almeida. Universidade de Brasília, Brazil. José Albertino Rafael. Instituto Nacional de Pesquisas da Amazônia (INPA), Brazil. Josenir Teixeira Câmara. Instituto Nacional de Pesquisas da Amazônia (INPA), Brazil Karla P. Tepedino. Universidade de Brasília, Brazil. Leonardo H. Gil-Azevedo. Museu Nacional, Universidade Federal do Rio de Janeiro, Brazil. Leonardo S. Gomes Rocha. Instituto Federal de Educação, Ciência e Tecnologia do Rio de Janeiro, Brazil. Ligia I. Moncada-Alvarez. Universidad Nacional de Colombia, Colombia. Lisiane Dilli Went. Instituto Nacional de Pesquisas da Amazônia (INPA), Brazil. Luciane Marinoni. Universidade Federal do Paraná, Brazil. Luis Gregorio Estrada. Universidad de Sucre, Colombia. Luiz Carlos Pinho. Universidade Federal de Santa Catarina, Brasil Marcoandre Savaris. Universidade Federal do Paraná, Brazil. Márcia Souto Couri. Museu Nacional, Universidade Federal do Rio de Janeiro, Brazil. Marta Wolff. Universidad de Antioquia, Colombia. Neal L. Evenhuis. Bishop Museum, Hawaii, U.S.A. Paula Fernanda Motta Rodrigues. Museu de Zoologia da Universidade de São Paulo, Brazil. Paula Raile Riccardi. Universidade de São Paulo, Brazil. Rafael Augusto Pinheiro de Freitas-Silva. Instituto Nacional de Pesquisas da Amazônia (INPA), Brazil. Rafaela Lopes Falaschi. Universidade de São Paulo, Brazil. Ramon Luciano Mello. Universidade Federal de Mato Grosso do Sul, Brazil. Renato Soares Capellari. Universidade de São Paulo, Brazil. Rosaly Ale-Rocha. Instituto Nacional de Pesquisas da Amazônia (INPA), Brazil. Ricardo Guerrero. Universidad Central de Venezuela, Venezuela. Rosana Tidon. Universidade de Brasília, Brazil. Sandra Pérez P. Universidad de Antioquia, Colombia. Sarah Siqueira Oliveira. Universidade de São Paulo, Brazil. Sheila Patrícia Carvalho-Fernandes. Museu Nacional, Brazil Silvio Shigueo Nihei. Universidade de São Paulo, Brazil. Stephen Marshall. University of Guelph, Canada. Tatiana Sepúlveda. Universidade Federal do Paraná, Brazil. Valéria Cid Maia. Museu Nacional, Universidade Federal do Rio de Janeiro, Brazil. Vera Cristina Silva. Universidade Estadual Paulista, Brazil. Viviane Rodrigues de Sousa. Museu Nacional, Universidade Federal do Rio de Janeiro, Brazil. Wayne Nielsen Mathis. National Museum of Natural History, U.S.A. William L. Murphy. Smithsonian Institution, U.S.A. Yardany Ramos-Pastrana. Universidad de La Amazonia, Colombia. 14 · Zootaxa 4122 (1) © 2016 Magnolia Press WOLFF ET AL. (EDS.) Zootaxa 4122 (1): 015–019 http://www.mapress.com/j/zt/ Copyright © 2016 Magnolia Press Article ISSN 1175-5326 (print edition) ZOOTAXA ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.4122.1.4 http://zoobank.org/urn:lsid:zoobank.org:pub:CEBCC02B-828F-4AAD-9194-946CA9A9044E FAMILY ANISOPODIDAE DALTON DE SOUZA AMORIM1, RAFAELA LOPES FALASCHI2 & SARAH SIQUEIRA OLIVEIRA3 1 Departamento de Biologia, Faculdade de Filosofia, Ciências e Letras de Ribeirão Preto, Universidade de São Paulo. Av. Bandeirantes, 3900, 14040-901. Ribeirão Preto SP Brazil. E-mail: dsamorim@usp.br 2 Universidade de São Paulo, Museu de Zoologia. Av. Nazaré, 481, Ipiranga. 04263-000. São Paulo SP Brazil. E-mail:rlfalaschi@gmail.com 3 Departamento de Ecologia, Instituto de Ciências Biológicas, Universidade Federal de Goiás, Campus II. Prédio ICB1, 74001-970. Goiânia GO Brazil. E-mail: oliveira.sarahcv@gmail.com Abstract This considerably small family is poorly known in Colombia, with only two species reported for the genus Sylvicola Harris (1776) so far. We synonymize Neomesochria Amorim & Tozoni (1994) to Mycetobia Meigen (1818), hence transferring the Dominican amber species Neomesochria antillea (Grimaldi 1991) and N. cryptambra (Grimaldi 1991), and the recent Neotropical species N. limanda (Stone 1966) and N. stonei (Lane & d’Andretta 1958) back to the genus Mycetobia. This paper provides new records for Mycetobia and Olbiogaster Osten-Sacken (1886) for Colombia. Key words: catalogue, Bibionomorpha, Colombia, distribution Introduction The family Anisopodidae encompasses seven recent genera in the world, with about 120 described species (Amorim & Tozoni 1994, Hancock & Amorim 2009). Five genera and 65 species are known from Neotropics (Hancock & Amorim 2009, Amorim et al. 2002), most of the records from Brazil (Papavero 1967). This family is considerably old in the fossil record. Protorhyphidae, possibly the family sister to the Anisopodidae, has members already in the Triassic (Shcherbakov et al. 2005), while the Triassic genus Crosaphis Evans, originally described as a homopteran, was considered by Kovalev (1983) as close to Mycetobia Meigen (1818), and most likely fits in the subfamily Mycetobiinae (Amorim & Tozoni 1994). In the Lower Jurassic there are species obviously related to Olbiogaster Osten-Sacken (1886). The genus Valeseguya Colless, known from a single male of a recent species in Australia (Colless 1990) and from relatively abundant specimens in Dominican amber (Grimaldi 1991), has been tentatively assumed to be an anisopodid (Colless 1990, Grimaldi 1991, Amorim & Tozoni 1994), but the description of Cretoseguya Amorim & Grimaldi from Myanmar Cretaceous amber allowed to transfer the clade to the Scatopsoidea (Amorim & Grimaldi 2006)—so it does not fit the Anisopodiformia anymore. Some authors have accepted family status for groups taken as subfamilies in other classifications (e.g., Hennig 1948, 1973, Amorim & Tozoni 1994), but we keep here a more conservative approach (see, e.g., Michelsen 1999, Thompson 2006), with the main clades of the group referred as subfamilies within the Anisopodidae s.l. A phylogeny for the family has been proposed by Amorim & Tozoni (1994). The most speciose genus in the Neotropics (see Hancock & Amorim 2009) is Sylvicola Harris (1776), but Olbiogaster has a fairly large number of species as well (see Tozoni 1993, Hancock & Amorim 2009). The Mycetobia-like Neotropical species of the family were included by Amorim & Tozoni (1994) in a separate genus, Neomesochria Amorim & Tozoni (1994). This group has few species described and is considerably rare in collections. Thompson (2006) disagreed from the transference of Mesochria neotropica Grimaldi (1991), a Dominican amber species, to the genus Neomesochria, as proposed by Amorim & Tozoni (1994). It may indeed be the case that, despite similarities with the Afro-Oriental genus Mesochria Enderlein (1910), some of the Accepted by S. Nihei: 10 Jun. 2015; published: 14 Jun. 2016 15 Neotropical species of Mycetobiinae are actually more connected to the Nearctic species of Mycetobia. The question of the Neotropical species of mycetobiines is still not well solved and a reassessment of the phylogeny of the mycetobiines with more characters is necessary. We transfer below the Neotropical species under Neomesochria back to Mycetobia and synonymize both genera. The Neotropical fauna of Anisopodidae is, hence, composed by six genera—Carreraia Corrêa (1947), Sylvicola, Olbiogaster, Lobogaster Philippi (1865), Mycetobia, and Mesochria. In Colombia there are published records of two species of the genus Sylvicola: S. fasciatus (Roeder) and S. marmoratus (Edwards) (Papavero 1967). We add here new records for the genera Olbiogaster and Mycetobia. These flies are more or less slender, with small to medium body size, varying from 2 to 12 mm. The head is small, rounded, three ocelli, with big dichoptic eyes in males and females. The antenna always has 14 flagellomeres, in most cases with slightly elongated flagellomeres. The thorax is convex and the wing membrane has color patterns as in Sylvicola. The male terminalia has the cercus prominent, which is also developed in females (Amorim 1997, Hancock & Amorim 2009). The larvae are saprophagous, found in fermenting substrates, rotten potatoes, leaves and roots decomposed, sappy exudations from trees, wood rotting, and in holes in trees (Hancock & Amorim 2009). In the material studied from Colombia, some specimens were obtained from bat feces. Acronyms used for the depositories IAvH—Alexander von Humboldt Biological Resources Research Institute, Bogota List of abbreviations cat.—catalogue des.—designation distr.—geographic distribution i.e.—example m.—meters mon.—monotypy nom. nov.—nomen novum (new name) n. syn.—new synonym orig. des.—by original designation pp.—pages refs.—references rev.—revision reinst.—reinstated sp., spp.—species ssp.—subspecies subs.—subsequent subg.—subgenus HT—Holotype LT—Lectotype NA—Not applicable NT—Neotype PT—Paratype ST—Syntype T—Type (unspecified) F (f#)—Female M (m#)—Male 16 · Zootaxa 4122 (1) © 2016 Magnolia Press AMORIM ET AL. Catalogue of Anisopodidae of Colombia Family Anisopodidae Knab, 1912 Subfamily Anisopodinae Knab, 1912 Genus Olbiogaster Osten-Sacken Olbiogaster Osten-Sacken, 1886: 20. Type species, Rhyphus taeniatus Bellardi, (Coquillett, 1910: 579). undescribed sp. Distr.: Colombia (Bolivar, Cauca, Meta). Material examined. COLOMBIA: Cauca: PNN [Parque Nacional Natural], Gorgona El Helechal, 2º58’N 78º11’W, 4 males, 30 m, 11.xii.2001-18.i.2002, H. Torres Leg. M.2788, Malaise, (IAvH); Bolivar: SFF [Santuario de Fauna y Flora], Los Colorados, Las Yaya, 09°54’N 75°07’W, 2 males, 280 m, 22.iv-07.v.2001, E. Deulufeut Leg, Malaise, (IAvH); Meta: PNN [Parque Nacional Natural], Sierra de la Macarena Cabaña Cerrillo, 03°21’N 73°56’W, 2 males, 493 m, 25.x-03.xi.2004, W. Willalba Leg., Malaise 3, (IAvH). Genus Sylvicola Harris Sylvicola Harris, 1776: 100. Type species, brevis Harris (des. Coquillett, 1910: 610) = fenestralis (Scopoli). Phryne Meigen, 1800: 16. Type species, Tipula fuscata Fabricius (des. Coquillett, 1910: 589). Suppressed by ICZN, 1963: 339. Anisopus Meigen, 1803: 264. Type species, fuscus Meigen (des. Coquillett, 1910: 507) = fuscata (Fabricius). Rhyphus Latreille, 1804: 188. Type species, Tipula fenestralis Scopoli (as fenestrarum; mon.). fasciatus (Roeder), 1886: 260 (Rhyphus). Type locality: Colombia, Cordillera, Páramo, 3200 m. Distr.: Colombia (Antioquia (Andes (Páramo de Santa Rita)). Refs.: Papavero, 1967: 2 (cat.). marmoratus (Edwards), 1923: 484, (Anisopus). Type locality: Colombia, Bogotá. Distr.: Colombia (Bogotá). Refs.: Papavero, 1967: 3 (cat.). Subfamily Mycetobyiinae Winnertz, 1863 Genus Mycetobia Meigen Mycetobia Meigen, 1818: 229. Type species, pallipes Meigen (Westwood, 1840: 127). Neomesochria Amorim & Tozoni, 1995: 533. Type species, Mycetobia stonei Lane & d’Andretta, 1958. n. syn. Comments. Following his discussion of problems in the phylogeny of the Mycetobiinae in Amorim & Tozoni (1994), Thompson (2006) formally moved the Dominican amber species Neomesochria neotropical (Grimaldi) back to Mesochria. Thompson (2006, page 13) recognized that “[t]he problem of a proper synapomorphy for Mycetobia remains” and “that the status of the Neotropical and New Caledonian species of Mycetobia remains to be clarified.” An investigation of the relationships between the mycetobiines of the world with additional characters is still necessary for a proper understanding of the evolution of the subfamily. Until this is done, it seems advisable to synonymize Neomesochria to Mycetobia, and to reinstate the Neotropical and New Caledonian species described in Mycetobia to their original generic combination—Mycetobia neocaledonica Baylac & Matile, 1988, reinst.; Mycetobia scutelaris Baylac & Matile, 1988, reinst.; Mycetobia antillea Grimaldi, 1991, reinst.; Mycetobia cryptambra Grimaldi, 1991, reinst.; Mycetobia limanda Stone, 1966, reinst.; Mycetobia stonei Lane & d’Andretta, 1958, reinst.. undescribed sp. Distr.: Colombia (Cauca, Bolivar, Risaralda). Material examined. COLOMBIA: Cauca: PNN [Parque Nacional Natural], Gorgona El Helechal, 2º58’N CATALOGUE OF ANISOPODIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 17 78º11’W, 30 m, 4 males, 11.xii.2001-18.i.2002, H. Torres Leg., Malaise (IAvH); Bolivar: SFF [Santurio Fauna y Flora], Los Colorados, Las Yaya, 09°54’N 75°07’W, 280 m, 2 males, 22.iv-07.v.2001, E. Deulufeut Leg., Malaise (IAvH); Risaralda: SFF [Santuario Fauna y Flora], Otún- Quimbaya Cuchilla Camino, 4°44’N 75°35’W, 1960 m, 2 males. 19.i-4.ii.2003, G. López Leg. Malaise (IAvH). Acknowledgements The authors are indebted to Michael Sharkey (National Science Foundation Grant DEB0205982), to whom we are most grateful. Additional information on the project can be found at the web address of the Project (http:// www.sharkeylab.org/biodiversity/static.php?app=colombia&page=index). During the preparation of this paper, the authors received financial support from FAPESP grants 2013/16524-9 (RLF) and 2012/51577-3 (SSO) and CNPq grants 314371/2009-5 (DSA) and 159016/2012-6 (RLF). TABLE 1. Anisopodidae species recorder from Colombia. Subfamily Genus Species Anisopodinae Olbiogaster sp. Sylvicola fasciatus (Roeder, 1886) marmoratus (Edwards, 1923) Mycetobiinae Mycetobia Author sp. References Amorim, D.S. (1997) Anisopodidae. In: Solís, A. (Ed.) Las Familias de insectos de Costa Rica. INBio. Avaiable from: http:// www.inbio.ac.cr/papers/insectoscr/Texto335.html (accessed January 2015) Amorim, D.S. & Grimaldi, D. (2006) Valeseguyidae, a new family of Diptera in the Scatopsoidea, with a new genus in Cretaceous amber from Myanmar. Systematic Entomology, 31 (3), 508–516. Amorim, D.S. & Tozoni, S.H.S. (1994) Phylogenetic and biogeographic analysis of the Anisopodoidea (Diptera, Bibionomorpha), with an area cladogram for intercontinental relationships. 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Paulo, 8 (8), 97–107. Enderlein, G. (1910) Diptera, Mycetophilidae. In: Reports of the Percy Sladen Trust Expedition to the Indian Ocean in 1905, 5, under the leadership of Mr. J. Stanley Gardiner, M.A. The Transactions of the Linnean Society of London, (2) Zoology, 14, 59–81. Edwards, F.W. (1923) Notes on the Dipterous Anisopodidae. Annals and Magazine of Natural History, 9 (12), 475–493. Grimaldi, D.A. (1991) Mycetobiine woodgnats (Diptera: Anisopodidae) from the Oligo-Miocene amber of the Dominican Republic, and Old World affinities. American Museum Novitates, 3014, 1–24. Hancock, E.G. & Amorim, D.S. (2009) Anisopodidae (Wood gnats or Window gnats). In: Brown, B.V., Borkent, A., Cumming, J.M., Wood, D.M., Woodley, N.E. & Zumbado, M.A. (Eds.), Manual of Central American Diptera, Vol. 1. NRC Research Press, Ottawa, pp. 341–345. Harris, M. (1776) An exposition of British Insects. Decad IV: 100–138, pls. 31–40. London. Hennig, W. (1948) Die Larvenformen der Dipteren. 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Pure and Applied Geophysics 162: 6–7, 1051−1076. Stone, A. (1966) Bredin-Archbold-Smithsonian Biological survey of Dominica. 2. New species of Diptera from Dominica (Anisopodidae and Bibionidae). Proceedings of United States National Museum, 121, 1–6 Thompson, C.F. (2006) New Mesochria species (Diptera: Anisopodidae) from Fiji, with notes on the classification of the family. Fiji Arthropods IV. Bishop Museum Occasional Papers, 86, 11–21. Tozoni, S.H.S. (1993) Revision of the genus Olbiogaster Osten-Sacken (Diptera, Anisopodoidea, Olbiogastridae). I. Description of 13 new species, taxonomical notes, and a key to the Neotropical species. Revista Nordestina de Biologia, 8 (2), 119–142. Westwood, J.O. (1840) An introduction to the modern classification of insects. Synopsis of the genera of Bristish Insects. 158 pp. London. Winnertz, J. (1863) Beitrag zu einer Monographie der Pilzmücken. Abhandlungen der Zoologisch-Botanischen Gesellschaft in Wien, 13, 637–964. CATALOGUE OF ANISOPODIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 19 Zootaxa 4122 (1): 020–025 http://www.mapress.com/j/zt/ Copyright © 2016 Magnolia Press Article ISSN 1175-5326 (print edition) ZOOTAXA ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.4122.1.5 http://zoobank.org/urn:lsid:zoobank.org:pub:22BD0DE0-31B2-4ABB-BB3C-CAFA9EA79087 FAMILY BIBIONIDAE RAFAELA LOPES FALASCHI1, SARAH SIQUEIRA OLIVEIRA2 & DALTON DE SOUZA AMORIM3* 1 Universidade de São Paulo, Museu de Zoologia. Av. Nazaré, 481. Ipiranga. 04263-000. São Paulo SP Brazil. E-mail: rlfalaschi@gmail.com 2 Departamento de Ecologia, Instituto de Ciências Biológicas, Universidade Federal de Goiás, Campus II. Prédio ICB1, 74001-970. Goiânia GO Brazil. E-mail: oliveira.sarahcv@gmail.com 3 Departamento de Biologia, Faculdade de Filosofia, Ciências e Letras de Ribeirão Preto, Universidade de São Paulo. Av. Bandeirantes, 3900. 14040-901. Ribeirão Preto SP Brazil. E-mail: dsamorim@usp.br, CNPq Research Fellowship. *Corresponding author: dsamorim@usp.br Abstract The Bibionidae are a family belonging to the suborder Bibionomorpha with four genera and 17 species known from Colombia. This work expands the distribution of these species to other localities in the country. Key words: catalogue, Bibionomorpha, Colombia, distribution Introduction The Bibionidae are a family belonging to the suborder Bibionomorpha, with eight genera and more than 700 species described worldwide (Pinto & Amorim 2000; Fitzgerald 2009). In the Neotropical region, there are records for seven genera and a total of 169 species (Hardy 1959, 1966; Amorim et al. 2002). Within the bibionid genera, only Hesperinus Walker has not been found in Neotropics so far. Hitherto there are records of four genera and 17 species in Colombia (Hardy 1966; Sturm 1990) (Table 1). Some authors accept Hesperinus in the separate family Hesperinidae (Krivosheina 1997; Papp 2010; Wood & Borkent 1989; Nel & Skartveit 2011), but this is not the main approach regarding the family classification. The family Bibionidae is currently organized in four subfamilies, Hesperininae, Penthetriinae, Pleciinae, and Bibioninae, of which the first three are monotypic (Pinto & Amorim 2000; Fitzgerald 2009; Papp 2010). The bibionids are usually black flies, with size ranging from 4.5 to 7.0 mm; some species have large orange portions of the body. The antennae are short and the ocelli are always present. The wing membrane is hyaline to black or brown, the pterostigma often being distinct. R2+3 and M3 are always absent, and R4+5 might be bifurcated or simple (i.e., R4 may be present or not). Males and females are easily recognized by the sexual dimorphism: females are bigger, dichoptic, with eyes widely separated; males are smaller than females, always holoptic. Acronyms used for the depositories. IAvH – Alexander von Humboldt Biological Resources Research Institute, Bogota, Colombia List of abbreviations cat.—catalogue des.—designation distr.—geographic distribution 20 Accepted by S. Nihei: 10 Jun. 2015; published: 14 Jun. 2016 i.e.—example m.—meters mon.—monotypy nom. nov.—nomen novum (new name) orig. des.—by original designation pp.—pages refs.—references rev.—revision sp., spp.—species ssp.—subspecies subs.—Subsequent subg.—subgenus HT—Holotype LT—Lectotype NA—Not applicable NT—Neotype PT—Paratype ST—Syntype T—Type (unspecified) F (f#)—Female M (m#)—Male Catalogue of Bibionidae of Colombia Family Bibionidae Fleming, 1821 Subfamily Penthetriinae Rohdendorf, 1946 Genus Penthetria Meigen Penthetria Meigen, 1803: 264. Type species, funebris Meigen (sub. mon., Meigen, 1804: 104). Eupeitenus Macquart, 1838: 88 (1838: 84). Type species, Penthetria atra Macquart (mon.) = heteroptera (Say). nigerrima (Bellardi), 1859: 14 (Plecia). Type locality: Mexico, Orizaba. Distr.: Mexico, Colombia, Ecuador. Refs.: Hardy, 1945: 388 (rev.); 1966: 2 (cat.). nigrita Perty, 1830: 180. Type locality: Brazil, Piauí. Distr.: Venezuela, Colombia, Ecuador, Peru, Brazil. Refs.: Hardy, 1966: 2 (cat.). Subfamily Pleciinae McAtee, 1921 Genus Plecia Wiedemann Plecia Wiedemann, 1828: 72. Type species, Hirtea fulvicollis Fabricius (Blanchard, 1840: 576). Rhinoplecia Bellardi,1859: 16. Type species, rostrata Bellardi (mon.). Penthera Philippi, 1865: 639. Type species, nigra Philippi (mon.). disparis Hardy, 1942: 108. Type locality: Colombia, Upper Putumayo River. Distr.: Colombia. Refs.: Hardy, 1945: 406 (rev.); 1966: 3 (cat.). CATALOGUE OF BIBIONIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 21 ecuadorensis Hardy, 1942: 108. Type locality: “Ecuador”. Distr.: Ecuador, Colombia. Refs.: Hardy, 1945: 406-407 (rev.); 1966: 3 (cat.). imperialis Schiner, 1868: 22. Type locality: “Colombia”. Distr.: Colombia. Refs.: Hardy, 1945: 411 (rev.), 1956: 86 (comment); 1966: 3 (cat.). (First available name for costalis Walker). paracollaris Hardy, 1961: 88. Type locality: Colombia, Buenaventura. Distr.: Colombia. Refs.: Hardy, 1966: 4 (cat.). plagiata Wiedemann, 1824: 11. Type locality: “Brazil”. Distr.: Widespread throughout the Neotropical region. Refs.: Hardy, 1945: 427 (rev.); 1953: 355 (comment); 1966: 4 (cat.). heteroptera Macquart, 1846: 21. Type locality: Colombia, Santa Fe de Bogotá. vittata Bellardi, (nec Wiedemann) 1862: 4. bellardi Townsend, 1912: 289. rugosa Hardy, 1940: 25. Type locality: Colombia, Bet. Queremal. Distr.: Colombia. Refs.: Hardy, 1945: 436 (rev.); 1966: 5 (cat.). Subfamily Bibioninae Fleming, 1821 Genus Bibio Geoffroy Bibio Geoffroy, 1762: 568. Type species, Tipula hortulana Linnaeus (Latreille, 1810: 442). Validated by ICZN, 1957: 86. Pullata Harris, 1776: 77. Type species, funesta Harris = pomonae (Fabricius) (Coquillett, 1910: 598). Hirtea Fabricius, 1798: 551 (nec Scopoli, 1763) (no type des.). dispar Schiner, 1868: 20. Type locality: “Colombia”. Distr.: Colombia, Argentina. Refs.: Hardy, 1953: 347 (redescription); 1966: 6 (cat.). superfluus Schiner, 1868: 20. Type locality: “Colombia”. Distr.: Colombia. Refs.: Hardy, 1966: 6 (cat.). Genus Dilophus Meigen Philia Meigen, 1800: 20. Type species, Tipula febrilis Linnaeus (Coquillett, 1910: 588). Suppressed by ICZN, 1963: 339. Dilophus Meigen, 1803: 264. Type species, Tipula febrilis Linnaeus (Latreille, 1810: 442). Acanthocnemis Blanchard, 1852: 355 (no type des.). Triploctenia Enderlein, 1934: 181. Type species, Dilophus tenuis Meigen (orig. des.). Tridicroctena Enderlein, 1934: 181. Type species, Dilophus africanus Becker (orig. des.). Dactylodiscia Enderlein, 1934: 181. Type species, Dilophus hiemalis Becker (orig. des.). Cnemidoctenia Enderlein, 1934: 181. Type species, Dilophus crassicrus Lundström (orig. des.). balfouri Edwards, 1935: 20. Type locality: Colombia, Villavicencio, Quatiquia River. Distr.: Colombia. Refs.: Hardy, 1966: 8 (cat.). espeletiae Sturm, 1990: 198. Type locality: Colombia, Pâramo de Monserrate, Bogotá. Distr.: Colombia. giganteus Macquart, 1846: 21. Type locality: “Nova Granada”. Distr.: Colombia. Refs.: Hardy, 1966: 9 (cat.). 22 · Zootaxa 4122 (1) © 2016 Magnolia Press FALASCHI ET AL. lucifer Schiner, 1868: 18. Type locality: “Colombia”. Distr.: Colombia. Refs.: Hardy, 1966: 9 (cat.). pallens (Blanchard), 1852: 357 (Acanthocnemis). Type locality: Chile, Chiloe. Distr.: Colombia, Chile. Refs.: Edwards, 1930: 80 (general morphology comments); Hardy, 1966: 10 (cat.). pictus Schiner, 1868: 19. Type locality: “Colombia”. Distr.: Colombia. Refs.: Hardy, 1966: 10 (cat.). tapir Schiner, 1868: 18. Type locality: “Colombia”. Distr.: Colombia. Refs.: Hardy, 1966: 11 (cat.). Acknowledgements The authors are indebted to Michael Sharkey (National Science Foundation Grant DEB0205982), to whom we are most grateful. Additional information on the project can be found at the web address of the project (http:// www.sharkeylab.org/biodiversity/static.php?app=colombia&page=index). During the preparation of this paper, the authors received financial support from FAPESP grants 2013/16524-9 (RLF) and 2012/51577-3 (SSO) and CNPq grants 159016/2012-6 (RLF) and 314371/2009-5 (DSA). TABLE 1. Bibionidae species recorded from Colombia. Subfamily Genus Species Author Penthetriinae Penthetria nigerrima (Bellardi), 1859 nigrita Perty, 1830 disparis Hardy, 1942 ecuadorensis Hardy, 1942 imperialis Schiner, 1868 paracollaris Hardy, 1961 plagiata Wiedemann, 1824 rugosa Hardy, 1940 dispar Schiner, 1868 superfluus Schiner, 1868 Pleciinae Bibioninae Plecia Bibio Dilophus balfouri Edwards, 1935 espeletiae Sturm, 1990 giganteus Macquart, 1846 lucifer Schiner, 1868 pallens (Blanchard), 1852 pictus Schiner, 1868 tapir Schiner, 1868 References Amorim, D.S., Silva, V.C. & Balbi, M.I.P.A. (2002) Estado do conhecimento dos Diptera neotropicais. In: Costa, C., Vanin, S.A., Lobo, J.M. & Melic, A. (Eds.), Proyecto de Red Iberoamericana de Biogeografia y Entomología Sistemática, PRIBES 2002i. Sociedad Entomológica Aragoneza y CYTED, Zaragoza, pp. 29–36. Bellardi, L. (1859) Saggio di ditterología messicana. Parte. I. - Memoire della Reale Accademia delle Scienze di Torino, Serie 2, 19, 1–80, pls. 1–2. Bellardi, L. (1859) Saggio di ditterología messicana. Appendice. - Memoire della Reale Accademia delle Scienze di Torino, Serie 2, 21, 1–18, pl. 1. Blanchard, E. (1840) Histoire naturelle des insects Orthoptères, Névroptères, Hémiptères, Hyménoptères, Lépidoptères et CATALOGUE OF BIBIONIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 23 Diptères. In: Laporte, F.L. de (Ed.), Histoire naturelle des animaux articulés, 3, pp. 1–672, pls. 1–67. [Paris] Blanchard, E. (1852) Diptera. In: Gay, C. (Ed.), Historia Fisica y Politica de Chile, Fauna Chilena, Insectos. Zoología, 7, pp. 353–358. Coquillett, D.W. (1910) The type-species of North American Genera of Diptera. Proceedings of the United States National Museum, 37, 499–647. http://dx.doi.org/10.5479/si.00963801.37-1719.499 Edwards, F.W. (1930) Bibionidae. In: Diptera of Patagonia and South Chile. Part 2 (Fascicle 3). British Museum (Natural History), London, pp. 77–88. Edwards, F.W. (1935) New Neotropical Bibionidae (Diptera). 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(Ed.), A Catalogue of the Diptera of the Americas south of the United States. Departamento de Zoologia da Secretaria da Agricultura do Estado de São Paulo, São Paulo, pp. 1–20. Harris, M. (1776–1780) An exposition of English insects, with curious observations and remarks, wherein each insect is particularly described; its parts and properties considered; the different sexes distinguished, and the natural history faithfully related. The whole illustrated with copper plates, drawn, engraved, and coloured, by the author. Printed for the Author, London, 166 + [4] pp, plates 1–50. International Commission on Zoological Nomenclature (1957) Opinion 441. Validation under the plenary powers of the names for five genera in the Order Diptera (Class Insecta) published in 1762 by Geoffroy (E.L.) in the work entitled "Histoire abrégée des insectes qui se trouvent aux environs de Paris." (Opinion supplementary to Opinion 228.). 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(1846) Diptères exotiques, nouveaux ou peu connus. Supplèment. Mémoires de la Société Royale des Sciences, de l'Agriculture et des Arts de Lille, I, 5–238 + pls. 1–20. [Roret, Paris] McAtee, W.L. (1921) Notes on Nearctic bibionid flies. Proceedings of United States of National Museum, 60 (11), 1–27. Meigen, J.G. (1800) Nouvelle classification des mooches à deux ailes (Diptera L.) d’après un plan tout nouveau. Perronneau, Paris, 40 pp. Meigen, J.G. (1803) Versuch einer neuen Gattungs Eintheilung der europäischen zweiflügeligen Insekten. Magazin für Insektenkunde, 2, 264. Meigen, J.G. (1804) Klassifikazion und Beschreibung der europäischen zweifügeligen Insekten (Diptera Linn.). Braunschweig, 1, 104. Nel, A. & Skartveit, J. (2011) Occurrence of the genus Hesperinus Walker (Diptera, Hesperinidae) in the Oligocene of France. Studia Dipterologica, 18 (1/2), 85–89. Papp, L. (2010) A study on Hesperinus Walker with description of a new species (Diptera: Hesperinidae). Acta Zoologica 24 · Zootaxa 4122 (1) © 2016 Magnolia Press FALASCHI ET AL. Academiae Scientiarum Hungaricae, 56 (4), 347–370. Perty, J.A.M. (1830) Insecta brasiliensia. In: von Spix, J.B. & von Martius, K.F.P. (Eds.), Delectus animalium articulatorum, quae in itinere per Brasiliam annis MDCCCVII-MDCCCXX jussu et auspiciis Maximiliani Josephi I. Bavariae regis augustissimi. ed. Martius, Munich, pp. 125–224. Philippi, R.A. (1865) Aufzählung der Chilenischen Dipteren. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien. Verhandlungen der Zoologisches-botanisches Gesellschaft Wien, 15, 595–782. Pinto, L.G. & Amorim, D.S. (2000) Bibionidae (Diptera: Bibionomorpha). Morfologia e análise filogenética. Holos, Ribeirão Preto, 98 pp. Schiner, J.R. (1868) Diptera. In: Wüllerstorf-Urbair, B. von (Ed.), Reise der österreichischen Fregatte Novara um die Erde in den Jahren 1857, 1858, 1859 unter den Befehlen des Commodore B. von Wüllerstorf-Urbair. Zoologischer Theil. Vol. 2. Abt. I. K. Gerold’s Son, Wien, pp. 1–388. http://dx.doi.org/10.5962/bhl.title.7913 Scopoli, J.A. (1763) Entomologia Carniolica exhibens insecta Carnioliæ indigena et distributa in ordines, genera, species, varietates. Methodo Linnæana. I.T. Trattner, Vindobonae (= Vienna), (30) + 420 + (1) pp. http://dx.doi.org/10.5962/bhl.title.34434 Sturm, H. (1990) Eine neue Dilophus-Art (Insecta, Diptera, Bibionidae) aus den Hochanden Kolumbiens. Annalen des Naturhistorischen Museums in Wien, 91, 197–204. Townsend, C.H.T. (1912) On the Diptera of Baja California, including some species from adjacent regions. Canadian Entomologist, 44, 289. http://dx.doi.org/10.4039/Ent44287-10 Wiedemann, C.R.W. (1824) Munus rectoris in Academia Christiana Albertina aditurus analecta entomologica ex Museo Regio Havniensi. el regio typographeo scholarum, Kiliae [Kiel], 11 pp. http://dx.doi.org/10.5962/bhl.title.77322 Wiedemann, C.R.W. (1828) Aussereuropäische zweifügelige Insekten. Vol. 1. In der Schulzischen Buchhandlung, Hamm, 608 pp. [pp. 72–81] Wood, D.M. & Borkent, A. (1989) Phylogeny and classification of the Nematocera. In: McAlpine, J.F. & Wood, D.M. (Coords.), Manual of the Neartic Diptera. Vol. 3. Monograph 32. Research Branch, Agriculture Canada, Ottawa, pp. 1333–1370. CATALOGUE OF BIBIONIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 25 Zootaxa 4122 (1): 026–040 http://www.mapress.com/j/zt/ Copyright © 2016 Magnolia Press Article ISSN 1175-5326 (print edition) ZOOTAXA ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.4122.1.6 http://zoobank.org/urn:lsid:zoobank.org:pub:FA1118FE-3B67-403C-AEEE-39A4830AC865 FAMILY CECIDOMYIIDAE VALÉRIA CID MAIA Museu Nacional, Depto. Entomologia, Quinta da Boa Vista, São Cristóvão, 20940-040 Rio de Janeiro, RJ, Brazil; E-mail: maiavcid@acd.ufrj.br Abstract This large family is poorly known in Colombia, where only 44 species have been recorded in 20 genera. All of them are included in Cecidomyiinae, which is the most diverse subfamily of gall midges in number of species and feeding habits, including phytophagous, predaceous and fungivorous species. Most of them are galler. The other subfamilies have never been recorded in this country. Key words: catalogue, Cecidomyiidae, Colombia, distribution, galler, host plant Introduction Cecidomyiidae are a monophyletic family of nematocerous characterized by these apomorphies: lack of tibial spur (adult), reduced larval head, styletiform mandibles (larva) and presence of spatula (larva) (Wood & Borkent 1989). The family is divided into six subfamilies: Catotrichinae, Lestremiinae, Micromyiinae, Porricondylinae, Winnertziinae and Cecidomyiinae (Gagné & Jaschhof 2014). The first five are considered the most primitive groups and include only fungivorous species. Cecidomyiinae are the youngest subfamily, as well as the most diverse, including the majority of the known species (Gagné 1989). Differently from the other subfamilies, Cecidomyiinae comprise not only fungivorous, but also predaceous and phytophagous species, being the majority gall inducer. Adult: slender, thin, and small to medium body 1.0–8.0mm. Head: eyes generally holoptic in both sexes, ocelli present only in the most primitive families; antenna primitively with 14 flagellomeres, most composed by a node and an apical neck, longer in males than in females; antennal sensoria including setae, setulae and circumfila; mouth parts generally reduced and nonfunctional, consisting of labrum, labella, hypopharynx and maxilary palpi. Thorax convex, wing with reduced venation, C continuous around margin, usually with a break near R5 insertion; legs usually long, without tibial spurs; tarsomere 1 of each leg longer than the second one in Lestremiinae, Catotrichinae, and Micromyiini and much shorter in Porricondylinae, Winnertziinae, and Cecidomyiinae; claws simple or toothed, empodia varying in length. Abdomen elongate, tergites 1-8 and sternites 2-7 or 2-8 generally rectangular, with setae, scales and a basal pair of trichoid sensilla; male terminalia consisting of dorsal cerci, hypoproct, gonocoxites, gonostyli and aedeagus; parameres present or absent; ovipositor varying in length and shape; female cerci two-segmented in the most primitive subfamilies and one-segmented in almost all Cecidomyiinae, fused into a single lobe or separate. Pupa: generally exarate; head with antennal horns, spines and papillae present or absent; prothoracic spiracles usually elongate; abdominal spiracles generally short; abdominal tergites with spines in many groups. Larvae: cephalic capsule reduced, eyeless, antennae generally short (excepting in predators), styletiform mandibules; prothorax with spatula (dermal and sclerotized structure used for digging through soil or cutting through plant tissue), nine abdominal segments, and integument with papillae (complement of lateral and terminal papillae taxonomically important). The family is cosmopolitan with about 6,203 described species in 736 genera (Gagné & Jaschhof 2014). In the Neotropical region, 500 species in 170 genera are known (Gagné 1994). In Colombia, only 44 species in 20 genera have been recorded (Table 1). The knowledge of the Colombian fauna results mainly from the works of Edwin Möhn (Senckenberg Research Institute, Germany) and Antonio Wünsche (Max Born Gymnasium, Germany). The 26 Accepted by S. Nihei: 13 Jun. 2015; published: 14 Jun. 2016 former described seven Colombian species (Möhn 1955a; 1955b; 1959; 1960; 1961; 1963; 1964 and 1975) and the latter 24 (Wüsnche 1979), totalizing together about 66% of the known species. Data on Colombian gall midges fauna were obtained from literature and are based mainly on the catalogues of Gagné (2004; 2010) and Gagné & Jaschhof (2014), as well as on the publications of Möhn (1955a; 1955b; 1959; 1960; 1961; 1963; 1964 and 1975) and Wünsche (1979). Acronyms used for the depositories BMNH—The Natural History Museum, London, England MNHNP—Muséum National d´Histoire Naturelle, Paris, France NYSM—USA: New York State Museum, Albany, USA SMNS—Germany: Staatliches Museum für Naturkunde, Stuttgart, Germany TAVC—Tavares Collection, in SMNS or MNHN (Houard Collection), USA USNM—National Museum of Natural History, Smithsonian Institution, Washington, D. C., USA List of abbreviations cat.—catalogue des.— designation distr.—geographic distribution i.e.—exemple m.— meters missp. – misspelling mon.— monotypy orig. des.—by original designation pp.— pages preocc.— preoccupied refs.—references rev.— revision sp., spp.—species subs.—subsequent subg.— subgenus HT—Holotype NT—Neotype LT—Lectotype ST—Syntype T—Type (unspecified) NA—Not applicable F—Female M—Male L —Larva P —Pupa Catalogue of Cecidomyiidae of Colombia Family Cecidomyiidae Newman, 1834 Subfamily Cecidomyiinae Newman, 1834 CATALOGUE OF CECIDOMYIIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 27 Supertribe Cecidomyiidi Rübsaamen & Hedicke, 1926 Tribe Aphidoletini Harris, 1966 Genus Bremia Rondani Bremia Rondani, 1860: 289, as subg. of Diplosis. Type species, Cecidomyia decorata Loew (orig. des.). Comm.: Cosmopolitan; 19 spp. Larvae are predators. The host of only one species is known: eggs of odonata. Refs.: Harris, 1981 (biology, larva); Gagné & Jasschhof, 2014 (cat.). mirifica Gagné, 1994. Type locality, Colombia, Valle del Cauca. HT M (USNM). Distr.: Colombia (Valle del Cauca). Refs.: Gagné, 1994: 176; Gagné, 2004: 94 (cat.); Gagné, 2010: 144 (cat.); Gagné & Jaschhof, 2014: 129 (cat.). Tribe Asphondyliini Rübsaamen & Hedicke, 1926 Subtribe Asphondyliina Gagné, 1994 Genus Asphondylia Loew Asphondylia Loew, 1850: 21, 37, as subg. of Cecidomyia. Type species, sarothamni Loew (subs. des. Karsch, 1877:15). Phyllophaga Rondani, 1856: 199. Type species, Cecidomyia fusca Meigen (orig. des.). Asphondilia Rondani, 1860: 290, missp. of Asphondylia. Cylindrocera Lioy, 1864: 503. Type species, Asphondylia sarothamni Loew (subs. des. Gagné, 2004). Monasphondylia Kieffer, 1913a: 48. Type species, Asphondylia phlomidis Trotter (orig. des.). Eumarchalia (also as Eumarschalia) Del Guercio, 1914: 231, as subg. of Schizomyia. Type-species, Schizomyia gennadii Marchal (orig. des.). Gisonobasis Rübsaamen, 1916a: 432. Type species, tournefortiae Rübsaamen (subs. des. Rübsaamen, 1916b: 12). Ischnonyx Rübsaamen, 1916b: 5. Type species, Cecidomyia verbasci Vallot (subs. des. Möhn 1955a: 147). Gisomobasis Shinji, 1944: 227, missp. of Gisonobasis. Eoasphondylia Möhn, 1960: 214, as subg. of Asphondylia. Type species, Asphondylia convolvuli Möhn (orig. des.). Ussurasphondylia Kovalev, 1964: 438, as subg. of Asphondylia. Type species, Asphondylia aceris Kovalev. Halimodendromyia Marikovskij, 1965: 75. Type species, heptopotamica Marikovskij (orig. des.). Chenasphondylia Mamaev, 1972: 898, as subg. of Asphondylia. Type-species, Asphondylia aelleniae Mamaev (orig. des.). Aspondilia Fedotova & Kovalev, 2001: 547, missp. of Asphondylia. Comm.: Cosmopolitan; 320 spp. All species are gallers. They induce galls on 66 plant families, on many plant parts, mostly on flowers and buds, preventing fruiting. All are associated with symbiotic fungi. Refs.: Möhn, 1961 (rev.,Neotr. spp.); Gagné, 2004 (syn. note.); Tokuda, 2012 (rev.); Gagné & Jaschhof, 2014 (cat.). blechi Wünsch 1979: 31. Type locality: Colombia, Magdalena, Rio Buritaca. HT M (SMNS). Blechum pyramidatum (Acanthaceae). Distr.: Colombia (Magdalena (Rio Buritica)). Refs.: Gagné, 1994: 123; Gagné, 2004: 76 (cat.); Gagné, 2010: 121 (cat.); Gagné & Jaschhof, 2014: 108 (cat.). boerhaaviae Möhn, 1959: 302. Type locality, El Salvador: San Vicente. HT M (SMNS). Boerhaavia erecta; Boerhaavia spp. (Nyctaginaceae). Distr.: Mexico, El Salvador, Colombia (Magdalena (Santa Marta, Punta de Betin, Mamatoco, Bahia Concha, Rio Manzanares, Bonda, Bahia Gairaca, Nenguange)), Jamaica. Refs.: Gagné, 1994: 123; Gagné, 2004: 76 (cat.); Gagné, 2010: 121 (cat.); Gagné & Jaschhof, 2014: 108 (cat.) camarae Möhn, 1959: 340. Type locality: El Salvador, Cuscatlán, 27 km, San Martin. HT M (SMNS). Lantana camara (Verbenaceae). Distr.: El Salvador, Colombia. Refs.: Gagné, 1994: 123; Gagné, 2004: 77 (cat.); Gagné, 2010: 121 (cat.); Gagné & Jaschhof, 2014: 109 (cat.). canavaliae Wünsch, 1979: 33. Type locality: Colombia, Magdalena, Cañaverales. HT F (SMNS). Canavalia maritima (Fabaceae). Distr.: Colombia (Magdalena (Cañaverales)). Refs.: Gagné, 1994: 124; Gagné, 2004: 77 (cat.); Gagné, 2010: 121 (cat.); Gagné & Jaschhof, 2014: 109 (cat.). 28 · Zootaxa 4122 (1) © 2016 Magnolia Press MAIA caprariae Wünsch, 1979: 53. Type locality: Colombia, Magdalena, Bahia Nenguange. HT M (SMNS). Capraria biflora (Scrophulariaceae). Distr.: Colombia (Magdalena (Bahia Nenguange)). Refs.: Gagné, 1994: 124; Gagné, 2004: 77 (cat.); Gagné, 2010: 121 (cat.); Gagné & Jaschhof, 2014: 109 (cat.). duplicornis Wünsch, 1979: 51. Type locality: Colombia, Magdalena, Cañaverales. HT M (SMNS). Melanthera aspera (Asteraceae). Distr.: Colombia (Magdalena (Cañaverales)). Refs.: Gagné, 1994: 124; Gagné, 2004: 78 (cat.); Gagné, 2010: 123 (cat.); Gagné & Jaschhof, 2014: 110 (cat.). evae Wünsch, 1979: 39. Type locality: Colombia, Magdalena, Bahia Gairaca. HT M (SMNS). Chamizoa sp. (Amaranthaceae). Distr.: Colombia (Magdalena (Bahia Gairaca)). Refs.: Gagné, 1994: 124; Gagné, 2004: 79 (cat.); Gagné, 2010: 123; Gagné & Jaschhof, 2014: 110 (cat.). lopezae Wünsch, 1979: 43. Type locality: Colombia: Magdalena, Santa Marta. HT M (SMNS). Amaranthus dubius, A. spinosus, Iresine angustifolia (type host not specified) (Amaranthaceae). Distr.: Colombia (Magdalena (Santa Marta)). Refs.: Gagné, 1994: 125; Gagné, 2004: 81 (cat.); Gagné, 2010: 126; Gagné & Jaschhof, 2014: 113 (cat.). portulacae Möhn, 1959: 308. Type locality: El Salvador, La Libertad, San Diego, La Libertad. HT P (SMNS). Portulaca oleracea (Portulacaceae). Distr.: USA (Florida), El Salvador, Colombia (Magdalena (Cañaverales and Santa Marta)), West Indies, Bolivia, Argentina. Refs.: Gagné, 1994: 126; Gagné, 2004: 82 (cat.); Gagné, 2010: 128; Gagné & Jaschhof, 2014: 114 (cat.). sidae Wünsch, 1979: 36. Type locality: Colombia, Magdalena, Cañaverales. HT M (SMNS). Sida acuta; S. rhombifolia (Malvaceae). Distr.: Colombia (Magdalena (Cañaverales)). Refs.: Gagné, 1994: 126; Gagné, 2004: 84 (cat.); Gagné, 2010: 130; Gagné & Jaschhof, 2014: 116 (cat.). yukawai Wünsch, 1979: 47. Type locality: Colombia: Magdalena, Bahia Nenguange. HT M (SMNS). Arenaria sp. (Caryophyllaceae), Melochia spp. (Sterculiaceae). Distr.: Colombia (Magdalena (Bahia Nenguange)). Refs.: Gagné, 1994: 128; Gagné, 2004: 86 (cat.); Gagné, 2010: 132; Gagné & Jaschhof, 2014: 118 (cat.). Genus Hemiasphondylia Möhn Hemiasphondylia Möhn, 1960: 229. Type species, mimosae Möhn 1960 (orig. des.). Comm.: Neotropical; 2 spp. Both species induces bud galls on Mimosoideae (Fabaceae). mimosae Möhn, 1960: 230. Type locality: El Salvador, La Libertad, N. El Cimarrón. HT M (SMNS). Mimosa albida; M. leiocarpa, Prosopis juliflora (Fabaceae). Distr.: El Salvador, Colombia (Magdalena (Cerro San Fernando)). Refs. Refs.: Gagné, 1994: 130; Gagné, 2004: 164 (cat.); Gagné, 2010: 245; Gagné & Jaschhof, 2014: 221 (cat.). mimosicola Gagné, 1978: 514 (Asphondylia), new name for then sec. hom. Asphondylia mimosae Möhn, 1960. Genus Rhoasphondylia Möhn Rhoasphondylia Möhn, 1960: 224. Type species, Oxasphondylia friburgensis Tavares (orig. des.). Comm.: It comprises three galling species, all associated with Asteraceae. sanpedri Wünsch, 1979: 63. Type locality: Colombia, Magdalena, San Pedro de la Sierra. HT M (SMNS). Chaetospira funcki (Asteraceae). Distr.: Colombia (Magdalena (San Pedro de la Sierra)). Refs.: Gagné, 1994: 131; Gagné, 2004: 237 (cat.); Gagné, 2010: 351; Gagné & Jaschhof, 2014: 317 (cat.). CATALOGUE OF CECIDOMYIIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 29 Subtribe Schizomyiina Gagné, 1994 Genus Schizomyia Kieffer Schizomyia Kieffer, 1889: 183. Type species, galiorum Kieffer, 1889 (mon.). Parasphondylia Kieffer, 1913b: 93. Type species, variicornis Kieffer, 1913b (orig. des.). Schyzomya, Schizomya Del Guercio, 1918: 126, missp. of Schizomyia. Shizomyia Shinji, 1938a: 239, missp. of Schizomyia. Comm.: Cosmopolitan; 55 spp. Schizomyia is associated with 26 plant families. It is an artificial genus that gathers species with needlelike ovipositors, four-segmented palpi, and larvae with four pairs of terminal papillae present. Refs.: Gagné & Menjivar, 2008 (review Neotropical spp.); Gagné & Jaschhof, 2014 (cat.). manihoti Tavares, 1925: 22. Type locality, Brazil (Ceará). ST M, F, P, L (TAVC). Manihot utilissima (Euphorbiaceae). Distr.: Colombia (Magdalena (Santa Marta)), Brazil (Ceará). Refs.: Gagné, 1994: 142; Gagné, 2004: 253 (cat.); Gagné, 2010: 370; Gagné & Jaschhof, 2014: 334 (cat.). Tribe Cecidomyiini Rübsaamen & Hedicke, 1926 Genus Contarinia Rondani Contarinia Rondani, 1860: 289, as subg. of Cecidomyia. Type species, Tipula loti De Geer, 1776 (orig. des.). Eudiplosis Kieffer, 1894a: xxviii. Type species, Contarinia sorbi Kieffer (subs. des., Gagné, 2004: 108). Kieffer, 1895: cclxxx designated Tipula loti as Type species, but it was neither of the two originally included species. Stictodiplosis Kieffer, 1894a: xxviii. Type species, Contarinia picridis Kieffer, 1894 (subs. des., Gagné, 2004: 108). Kieffer, 1895: cclxxx designated Contarinia nubilipennis as Type species, but it was neither of the two originally included species. Contariuia Rübsaamen, 1906: 194, missp. of Contarinia. Syndiplosis Rübsaamen, 1910a: 284. Type species, winnertzi Rübsaamen, 1910a (orig. des. as n. g., n. sp.) = petioli Kieffer. Atylodiplosis Rübsaamen, 1910b: 342. Type species, Diplosis acetosellae Rübsaamen (subs. des. Rübsaamen & Hedicke, 1926: 261). Doxodiplosis Kieffer, 1912a: 1. Type species, picridis Kieffer, 1912a (orig. des.). Dryodiplosis Kieffer, 1912a: 1. Type species, Contarinia subulifex Kieffer, 1897 (orig. des.). Navasodiplosis Tavares, 1920: 65. Type species, camphorosmae Tavares, 1920 (orig. des.). Sissudiplosis Mani, 1943: 44. Type species, chatterjeei Mani, 1943 (orig. des.). Bothriochloamyia Rao & Sharma, 1977: 237. Type species, orientalis Rao & Sharma, 1977 (orig. des.). Contarinomyia Fedotova, 1991: 51. Type species, Contarinia reaumuriae Fedotova, 1991 (orig. des.). Achillinia Fedotova, 1992: 115, as subgenus Contarinia. Type species, Contarinia achilleae Fedotova, 1992 (orig. des.). Comm.: Cosmopolitan; 313 spp. The genus is associated with 60 plant families. Larvae of many species are free-living in flower heads, others induce leaf roll, or less frequently complex galls. Some species are inquilinous in galls. Refs.: Gagné, 1994; 2004 (syn. note.); Gagné & Jaschhof, 2014 (cat.). gossypii Felt, 1908a: 210. Type locality: Antigua. ST M, F (USNM). Gossypium sp. (Malvaceae). Distr.: Colombia, U.S. Virgin Is, Antigua, Montserrat, Barbados. Refs.: Gagné, 1994: 147; Gagné, 2004: 113 (cat.); Gagné, 2010: 172; Gagné & Jaschhof, 2014: 115 (cat.). Genus Prodiplosis Felt Prodiplosis Felt, 1908b: 403. Type species, Cecidomyia floricola Felt, 1908b (orig. des.). Geisenheyneria Rübsaamen, 1910a: 289. Type species, rhenana Rübsaamen, 1910a (orig. des. as n. g., n. sp.). Comm.: Holarctic, Neotropical; 12 spp. The genus is associated with 16 plant families. Many larvae live in flowers and buds. Some species are polyphagous and economically important. Ref.: Gagné, 1986 (revision); Gagné & Jaschhof, 2014 (cat.). longifila Gagné, 1986: 240. Type locality: USA, Florida, Monroe Co., Marco I. HT M (USNM). Gossypium sp. (Malvaceae); Lycopersicon esculentum & Solanum tuberosum (Solanaceae), Citrus aurantifolia (Rutaceae), Medicago sativa & Phaseolus sp. (Fabaceae), Chenopodium ambrosioides (Chenopodiaceae), & Ricinus communis (Euphorbiaceae). Distr.: USA (Virginia, Florida), Colombia, Ecuador, Peru. Refs.: Gagné, 1994: 149; Gagné, 2004: 226 (cat.); Gagné, 2010: 335; Gagné & Jaschhof, 2014: 303 (cat.). 30 · Zootaxa 4122 (1) © 2016 Magnolia Press MAIA Tribe Lestodiplosini Harris, 1966 Genus Feltiella Rübsaamen Feltiella Rübsaamen, 1910a: 285. Type species, tetranychi Rübsaamen, 1910a (orig. des. as n. g., n. sp.) = acarisuga Vallot. Therodiplosis Kieffer, 1912a: 2. Type species, persicae Kieffer, 1912a (orig. des.) = acarisuga Vallot, 1827. Acaroletes Kieffer, 1913c: 229. Type species, Arthrocnodax tetranychi Kieffer (orig. des.). Acroletes Mani, 1946: 215, missp. of Acaroletes. Comm.: Cosmopolitan; 10 spp. Larvae of this cosmopolitan genus prey almost exclusively on red spider mites. Refs.: Gagné, 1995 (rev.); 2004 (syn. note); Gagné & Jaschhof, 2014 (cat.). insularis (Felt, 1913a: 305) (Mycodiplosis). Type locality: Puerto Rico, Rio Piedras. LT (Gagné, 1995:26) M F L (NYSM). Tetranychus sp.; many tetranychids (Acarina). Distr.: USA (Illinois, New Jersey, Florida), Jamaica, Puerto Rico, Trinidad, Colombia, Argentina. Refs.: Gagné, 1994: 161; Gagné, 2004: 153 (cat.); Gagné, 2010: 228. carolina Felt, 1913b: 488 (Arthrocnodax). USA: South Carolina, Batesburg; (Gagné, 1995: 26), LT M (NYSM). constricta Felt, 1914a: 481 (Arthrocnodax). Puerto Rico: Rio Piedras; (Gagné, 1995: 26), LT M (NYSM). venatoria Felt, 1917: 195 (Feltiella). USA: Illinois, Chicago; LT (Gagné, 1995: 26) M [NYSM]. Genus Lestodiplosis Kieffer Leptodiplosis Kieffer, 1894a: xxviii. Type species, Lestodiplosis septemguttata Kieffer (subs. des. Kieffer, 1895: cclxxx). Replaced by Lestodiplosis as valid original spelling (ICZN, 1958: 291). Lestodiplosis Kieffer, 1894b: 84. Type species, Cecidomyia septemguttata Kieffer (subs. des. Kieffer, 1895: cclxxx). Coprodiplosis Kieffer, 1894b: 84. Type species, Cecidomyia polypori Loew (subs. des. Kieffer, 1895: cclxxx). Hemidiplosis Kieffer, 1894c: 9 (publ'n not seen). Type species, Lestodiplosis rosea Kieffer, 1894c (mon.). Adelgimyza Del Guercio, 1918: 231. Type species, strobilobii Del Guercio, 1918 (orig. des. as n.g, n.sp.). Cecidomyella Del Guercio, 1918: 247. Type species, aulacaspidis Del Guercio, 1918 (orig. des. as n. g., n. sp.). Moreschiella Del Guerico, 1918: 157. Type species, moricola Del Guerico, 1918 (mon.) = moricolata Gagné, 2004. Theatodiplosis Tavares, 1922a: 148. Type species, quercina Tavares, 1922a (orig. des.). Cecidomyela Felt, 1925: 163, missp. of Cecidomyella. Cecidomyiella Mani, 1934: 428, missp. of Cecidomyella. Chiliodiplosis Möhn, 1955b: 415. Type species, vasta Möhn (orig. des.). Phonodiplosis Möhn, 1955b: 420. Type species, casta Möhn (orig. des.). Coplodiplosis Shinji, 1944: 245, missp. of Coprodiplosis. Comm.: Cosmopolitan and copal fossil; 181 spp. Larvae are predators of several arthropods (Diplopoda, Acarina, Coleoptera, Diptera, Hemiptera, Hymenoptera, Lepidoptera, Thysanoptera). Many are found in inflorescences. They are associated with at least 18 plant families. According to gagné & jaschhof, 2014, this genus contains many apparent synonyms. Ref.: Gagné & Jaschhof, 2014 (cat.). gagnei Baylac, 1987: 125. Type locality: Colombia, San-Alberto. HT M (MNHN). Elaeidobius subvittatus (Coleoptera: Curculionidae). Distr.: Colombia (Cesar (San-Alberto)). Refs.: Gagné, 1994: 163; Gagné, 2004: 187 (cat.); Gagné, 2010: 278; Gagné & Jaschhof, 2014: 250 (cat.). Genus Trisopsis Kieffer Trisopsis Kieffer, 1912b: 171. Type species, oleae Kieffer, 1912b (orig. des.). Plagiodiplosis Kieffer, 1913a: 55. Type species, Lestodiplosis nana Kieffer, 1913a (orig. des.). Comm.: Cosmopolitan; 25 spp., presumably all larvae are predators although most were caught In flight. According to Gagné & Jaschhof, 2014, this genus may not be monophyletic. Ref.: Gagné & Jaschhof, 2014 (cat.). incisa (Felt, 1907: 43) (Cecidomyia). Type locality: USA, New York, Albany. HT M (NYSM). Orig. unknow; from laboratory colonies of Anopheles sp. (Culicidae), Perileucoptera coffeella (Lepidoptera: Lyonetiidae), Pedronia sp. (Hemiptera: Pseudococcidae), on shoots of Sorghum sp., on leaves of Cyrtandra sp. (Gesneriaceae), in fruiting bodies of fungi, and various insect larvae in flower heads of Chromolaena odorata (Asteraceae) (cf. to list under CATALOGUE OF CECIDOMYIIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 31 Lestodiplosis callipus).Distr.: USA (Washington, New York, District of Columbia), Trinidad, Colombia, Hawaiian Is (T. oleae of Hardy), Greece, Israel. Refs.: Gagné, 1997; Gagné, 1994: 165; Gagné, 2004: 269 (cat.); Gagné, 2010: 396 (cat.); Gagné & Jaschhof, 2014: 357 (cat.). oleae Kieffer, 1912b: 171. Type locality: South Africa, Wellington. ST M, F (?). Distr.: South Africa, New Zealand, Colombia. Refs.: Gagné, 1994: 165; Gagné, 2004: 269 (cat.); Gagné, 2010: 396; Gagné & Jaschhof, 2014: 358 (cat.). Genus Diadiplosis Felt Diadiplosis Felt, 1911a: 54. Type species, cocci Felt, 1911a (orig. des.). Coccodiplosis Meijere, 1917: 238. Type species, pseudococci Meijere, 1917 (mon.) = smithi Felt, 1915. Cleodiplosis Felt, 1922: 1. Type species, aleyrodici Felt, 1922 (orig. des.). Ghesquierinia Barnes, 1939: 327. Type species, megalamellae Barnes, 1939 (orig. des.). Kalodiplosis Felt, 1915: 229. Type species, Dicrodiplosis multifila Felt, 1915 (orig. des.). Schizobremia Felt, 1926: 183. Type species, formosana Felt, 1926 (orig. des.). Olesicoccus Borgmeier, 1931: 186. Type species, costalimai Borgmeier, 1931 (orig. des.) = coccidivora Felt, 1914b. Cieodiplosis Shinji, 1944: 232, missp. of Cleodiplosis. Nipponodiplosis Harris, 1968: 458. Type species, Diadiplosis hirticornis Felt (orig. des.). Golanudiplosis Grover & Prasad, 1968: 213. Type species, japonica Grover & Prasad, 1968 (mon.). Phagodiplosis Blanchard, 1938: 345. Type species, haywardi Blanchard, 1938 (orig. des.) = coccidivora Felt, 1914b. Vincentodiplosis Harris, 1968: 473. Type species, Lobodiplosis coccidarum Felt, 1911b (orig. des.) = coccidarum Cockerell, 1892. Comm.: Cosmopolitan, chiefly tropical; 30 spp. All larvae are predaceous on Hemiptera (mainly Pseudococcidae, but also Aleyrodidae, Coccidae, Dactylopiidae, Diaspididae, Eriococcidae, and Margarodidae). Refs.: Harris, 1968 (rev., illus.); Gagné, 1994 (review); Gagné & Jaschhof, 2014 (cat.). coccidarum (Cockerell, 1892: 181) (Diplosis). Type locality: Jamaica. HT M F (USNM). Orig. from a box containing various Hemiptera: Aleurodes sp. (Aleyrodidae); Aonidiella aurantii & Dactylopius sp. (Diaspididae); and Ferrisia sp., Dysmicoccus spp., Nipaecoccus sp., Paracoccus marginatus, Phenacoccus spp., Pseudococcus spp., & Saccharicoccus spp. (Pseudococcidae). Distr.: USA (Florida), West Indies (Cuba to Trinidad), Colombia, Guyana, Peru. Refs.: Gagné, 1994: 184; Gagné, 2004: 145 (cat.); Gagné, 2010: 213; Gagné & Jaschhof, 2014: 191 (cat.). coccidarum Felt, 1911b: 195 (Lobodiplosis). St. Vincent; ST M F (NYSM). cocci Felt, 1913a: 304 (Karschomyia). Puerto Rico: Patillas; STM F (lost). vaupedis (Harris, 1968: 450) (Ghesquierinia). Type locality: Colombia, Vaupés. HT M (BMNH). Undet. sp. of coccoid; Planococcus sp. (Hemiptera: Pseudococcidae). Distr.: Colombia (Vaupés (Gadaloupe)), Guadaloupe. Refs.: Gagné, 1994: 185; Gagné, 2004: 146 (cat.); Gagné, 2010: 213 (cat.); Gagné & Jaschhof, 2014: 192 (cat.). Genus Moehniella Wünsch Moehniella Wünsch, 1979: 66. Type species, fernandi Wünsch 1979 (orig. des.). Comm.: Neotropical; 1 sp. The genus is known for only one species, probably an inquiline in fruits of Mimosa (Fabaceae). Ref.: Gagné & Jaschhof, 2014 (cat.). fernandi Wünsch, 1979: 67. Type locality: Colombia, Magdalena, Cerro San Fernando. HT F (SMNS). Mimosa leiocarpa (Fabaceae). Distr.: Colombia (Magdalena (Cerro San Fernando)). Refs.: Gagné, 1994: 192; Gagné, 2004: 202 (cat.); Gagné, 2010: 299 (cat.); Gagné & Jaschhof, 2014: 270 (cat.). Supertribe Lasiopteridi Rübsaamen & Hedicke, 1926 Tribe Alycaulini Rübsaamen & Hedicke, 1926 32 · Zootaxa 4122 (1) © 2016 Magnolia Press MAIA Genus Alycaulus Rübsaamen Alycaulus Rübsaamen, 1916a: 476. Type species, mikaniae Rübsaamen 1916a (mon.). Comm.: Neotropical; 3 spp., all induce galls on Mikania (Asteraceae). Ref.: Gagné & Jaschhof, 2014 (cat.). trilobatus Möhn, 1964: 583. Type locality: El Salvador, Cuscatlán, NE Cojutepeque. HT L (SMNS). Mikania micrantha; M. cordifolia (Asteraceae). Distr.: El Salvador, Colombia (Magdalena (Rio Sevilla, Rio Buritaca)). Refs.: Gagné, 1994: 65; Gagné, 2004: 66 (cat.); Gagné, 2010: 106 (cat.); Gagné & Jaschhof, 2014: 95 (cat.). Genus Geraldesia Tavares Geraldesia Tavares, 1917: 134. Type species, eupatorii Tavares (orig. des.). Comm.: Neotropical; 3 spp., associated with Asteraceae (2 spp.) and Polygonaceae (01 sp.). All induce blister leaf galls. Ref.: Gagné & Jaschhof, 2014 (cat.). polygonarum Wünsch, 1979: 111. Type locality: Colombia, Magdalena, Cerro San Fernando. HT M (SMNS). Triplaris, Symmeria and Ruprechtia sp. (Polygonaceae). Distr.: Colombia (Magdalena (Cerro San Fernando)). Refs.: Gagné, 1994: 68; Gagné, 2004: 155 (cat.); Gagné, 2010: 232 (cat.); Gagné& Jaschhof, 2014: 210 (cat.). Genus Meunieriella Kieffer Meunieria Rübsaamen, 1905: 137, preocc. Kieffer, 1904. Type species, dalechampiae Rübsaamen, 1905 (orig. des.). Meunieriella Kieffer, 1909: 35, new name for Meunieria Rübsaamen 1905. Dolicholabis Tavares 1918: 72. Type species, lantanae Tavares 1918 (orig. des.). Comm.: Neotropical, Nearctic; 21 spp. Most species are Neotropical and inquilinous in galls of other Cecidomyiidi but at least two are gall makers. They are associated at least with 11 plant families. Refs.: Möhn, 1975 (rev.); Gagné & Jaschhof, 2014 (cat.). datxeli Wünsch, 1979: 123. Type locality: Colombia, Magdalena, Bahia Nenguange. HT M (SMNS). Triplaris, Symmeria and Ruprechtia sp. (Polygonaceae). Distr.: Colombia (Magdalena (Bahia Nenguange)). Refs.: Gagné, 1994: 81; Gagné, 2004: 200 (cat.); Gagné, 2010: 268 (cat.); Gagné & Jaschhof, 2014: 268 (cat.). gairae Wünsch, 1979: 127. Type locality: Colombia: Magdalena, Bella-Vista. HT M (SMNS). Cordia subtruncata (Boraginaceae). Distr.: Colombia (Magdalena (Bella-Vista)). Refs.: Gagné, 1994: 81; Gagné, 2004: 200 (cat.); Gagné, 2010: 268 (cat.); Gagné & Jaschhof, 2014: 268 (cat.). magdalenae Wünsch, 1979: 119. Type locality: Colombia, Magdalena, Bahia Nenguange. HT M (SMNS). Distr.: Colombia (Magdalena (Bahia Nenguange)). Coccoloba candolleana (Polygonaceae). Refs.: Gagné, 1994: 82; Gagné, 2004: 201 (cat.); Gagné, 2010: 268 (cat.); Gagné & Jaschhof, 2014: 268 (cat.). Genus Neolasioptera Felt Neolasioptera Felt, 1908b: 330. Type species, Lasioptera vitinea Felt, 1907 (subs. des. Coquillett, 1910: 575). Luisieria Tavares, 1922b: 43. Type species, fariae Tavares, 1922b (orig. des.). Dilasioptera Möhn, 1964: 570, as subg. of Neolasioptera. Type species, Neolasioptera serrata Möhn, 1964 (orig. des.). Neurolasioptera Brèthes, 1922: 138. Type species, baezi Brèthes, 1922 (mon.). Physalidicola Brèthes, 1917: 240. Type species, argentata Brèthes, 1917 (orig. des.). Comm.: Nearctic, Neotropical; 134 spp. The genus is associated with 44 plant families, mostly Asteraceae. The species induce mainly stem galls, but also petiole, leaf vein, or flower galls. Refs.: Gagné, 1994 (review); Gagné & Jaschhof, 2014 (cat.). camarae Möhn, 1964: 565. Type locality: El Salvador, San Diego and La Libertad. HT M (SMNS). Lantana camara (Verbenaceae). Distr.: Colombia (Magdalena (La Togrera, Minca), El Salvador. Refs.: Gagné, 1994: 71; Gagné, 2004: 201 (cat.); Gagné, 2010: 306 (cat.); Gagné & Jaschhof, 2014: 276 (cat.). CATALOGUE OF CECIDOMYIIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 33 cruttwellae Gagné, 1977: 115. Type locality: Trinidad, Simla. HT F (USNM). Chromolaena odorata (Asteraceae). Distr.: Colombia, Trinidad. Refs.: Gagné, 1994: 72; Gagné, 2004: 206 (cat.); Gagné, 2010: 307 (cat.); Gagné & Jaschhof, 2014: 277 (cat.). odorati Wünsch, 1979: 100 (Neolasioptera). Colombia, Magdalena (Cañaverales); HT M (SMNS). cusani Wünsch, 1979: 104. Type locality: Colombia, Magdalena, Donama. HT L (SMNS). Undet. sp. of Amaranthaceae. Distr.: Colombia (Magdalena). Refs.: Gagné, 1994: 72; Gagné, 2004: 207 (cat.); Gagné, 2010: 307 (cat.); Gagné & Jaschhof, 2014: 277 (cat.). diclipterae Wünsch, 1979: 78. Type locality: Colombia, Magdalena, Rio Manzanares. HT M (SMNS). Dicliptera assurgens (Acanthaceae). Distr.: Colombia (Magdalena (Rio Manzanares). Refs.: Gagné, 1994: 72; Gagné, 2004: 207 (cat.); Gagné, 2010: 307 (cat.); Gagné & Jaschhof, 2014: 277 (cat.). donamae Wünsch, 1979: 97. Type locality: Colombia, Magdalena, Donama. HT L (SMNS). Iresine angustifolia (Amaranthaceae). Distr.: Colombia (Magdalena (Donama). Refs.: Gagné, 1994: 72; Gagné, 2004: 207 (cat.); Gagné, 2010: 307 (cat.); Gagné & Jaschhof, 2014: 277 (cat.). mincae Wünsch, 1979: 95. Type locality: Colombia, Magdalena, Minca. HT L (SMNS). Melanthera aspera (Asteraceae). Distr.: Colombia (Magdalena (Minca). Refs.: Gagné, 1994: 74; Gagné, 2004: 209 (cat.); Gagné, 2010: 309 (cat.); Gagné & Jaschhof, 2014: 279 (cat.). olivae Wünsch, 1979: 92. Type locality: Colombia, Magdalena, Donama. HT F (SMNS). Calea caracasana (Asteraceae). Distr.: Colombia (Magdalena, Donama). Refs.: Gagné, 1994: 74; Gagné, 2004: 209 (cat.); Gagné, 2010: 310 (cat.); Gagné & Jaschhof, 2014: 279 (cat.). portulacae (Cook, 1906): 251. Type locality: Cuba. SYN gall depos. unknown. Portulaca oleracea (Portulacaceae). Distr.: Colombia (Santa Marta, Bahia Nenguange), Cuba, El Salvador, Jamaica, Mexico, USA (Florida), West Indies. Refs.: Gagné, 1994: 74; Gagné, 2004: 210 (cat.); Gagné, 2010: 310 (cat.); Gagné & Jaschhof, 2014: 279 (cat.). samariae Wünsch, 1979: 89. Type locality: Colombia, Magdalena, Santa Marta. HT M (SMNS). Adenocalymma dugandi (Bignoniaceae). Distr.: Colombia (Magdalena (Santa Marta)). Refs.: Gagné, 1994: 75; Gagné, 2004: 210 (cat.); Gagné, 2010: 310 (cat.); Gagné & Jaschhof, 2014: 280 (cat.). tribulae Wünsch, 1979: 81. Type locality: Colombia, Magdalena, Santa Marta. HT M (SMNS). Tribulus cistoides (Zygophyllaceae). Distr.: Colombia (Magdalena (Santa Marta)). Refs.: Gagné, 1994: 75; Gagné, 2004: 211 (cat.); Gagné, 2010: 311 (cat.); Gagné & Jaschhof, 2014: 280 (cat.). Tribe Camptoneuromyiini Möhn, 1975 Genus Camptoneuromyia Felt Camptoneuromyia Felt, 1908b: 334. Type species, Dasineura virginica Felt, 1907 (subs. des. Coquillett, 1910: 518). Comm.: Neotropical (9 spp.) and Nearctic 6 spp.). Most are inquilines in galls of other Cecidomyinae. Ref.: Gagné & Jaschhof, 2014 (cat.). boerhaaviae Möhn, 1975: 22. Type locality: El Salvador, San Vicente, km 81, nr Rio San Felipe. HT M (SMNS). Boerhaavia erecta; B. caribaea (Nyctaginaceae Distr.: El Salvador, Colombia (Magdalena (Bahia Nenguange)). Refs.: Gagné, 1994: 78; Gagné, 2004: 98 (cat.); Gagné, 2010: 150 (cat.); Gagné & Jaschhof, 2014: 135 (cat.). 34 · Zootaxa 4122 (1) © 2016 Magnolia Press MAIA Genus Domolasioptera Möhn Domolasioptera Möhn, 1975: 31. Type species, adversaria Möhn, 1975 (orig. des.). Comm.: Neotropical; 7 spp., larvae are inquilines in galls of other Cecidomyiinae. Ref.: Gagné & Jaschhof, 2014 (cat.). acuario Wünsch, 1979: 115. Type locality: Colombia Magdalena, Rio Buritaca. HT M (SMNS). Mikania cordifolia (Asteraceae). Distr.: Colombia (Magdalena (Rio Buritaca)). Refs.: Gagné, 1994: 79; Gagné, 2004: 148 (cat.); Gagné, 2010: 217 (cat.); Gagné & Jaschhof, 2014: 196 (cat.). Tribe Rhopalomyiini Rübsaamen & Hedicke, 1926 Genus Rhopalomyia Rübsaamen Rhopalomyia Rübsaamen, 1892: 370. Type species, Oligotrophus tanaceticola Karsch, 1879 (subs. des. Kieffer, 1896: 89). Diarthronomyia Felt, 1908b: 339. Type species, artemisiae Felt (orig. des.) = pomum Gagné, 1975. Diathronomyia Coquillett, 1910: 532, missp. of Diarthronomyia. Calopedila Kieffer, 1913a: 49. Type species, Rhopalomyia herbsti Kieffer (orig. des.). Misospatha Kieffer, 1913b: 48. Type species, Rhopalomyia globifex Kieffer & Jörgensen, 1910 (orig. des.). Panteliola Kieffer, 1913a: 49. Type species, Rhopalomyia haasi Kieffer, 1905 (orig. des.). Boucheella Rübsaamen, 1914: 93. Type species, Cecidomyia artemisiae Bouché, 1834 (orig. des.). Dichelonyx Rubsaamen, 1914: 94. Type species, Cecidomyia foliorum Loew, 1850 (orig. des.). Dictyomyia Tavares, 1919a: 25. Type species, navasina Tavares, 1919a (orig. des.). Navasia Tavares, 1919a: 34, preocc. Kirby, 1914. Type-species, santolinae Tavares, 1919a (orig. des.). Navasiella Tavares, 1919b: 93, new name for Navasia Tavares, 1919a . Eudictyomyia Tavares, 1920: 55. Type species, Rhopalomyia navasi Tavares, 1904 (orig. des.). Rhopalomya Shinji, 1938b: 1063, missp. of Rhopalomyia. Misospatna Shinji, 1939: 588, missp. of Misospatha. Phopalomyia Shinji, 1939: 588, missp. of Rhopalomyia. Ropalomyia Shinji, 1939: 588, missp. of Rhopalomyia. Calopedia Shinji, 1944: 323, missp. of Calopedila. Mesospatha Shinji, 1944: 191, missp. of Misospatha. Mesospathia Shinji, 1944: 29, missp. of Misospatha. Mesospathi Shinji, 1944: 42, missp. of Misospatha. Rhapalomyia Shinji, 1944: 162, missp. of Rhopalomyia. Artemisiobia Kovalev, 1967: 102. Type species, globosa Kovalev, 1967 (orig. des.). Calopedilla Gagné, 1994: 88, missp. of Calopedila. Dracunculomyia Fedotova, 1999a: 834. Type species, kashkarovi Fedotova, 1999a (orig. des.). Arenaromyia Fedotova, 1999b: 588. Type species, caspica Fedtova, 1999b (orig. des.). Absinthomyia (also as Absinhomyia), as subg. of Dracunculomyia Fedotova, 1999a: 835. Type-species, Dracunculomyia bergi Fedotova, 1999a (orig. des.). Pupascleromyia (also as Pupascleromtyia) Fedotova, 1999a: 844, nomen nudum. Seriphidomyia Fedotova, 1999a: 841, nomen nudum. Polynomyia Fedotova, 1999a: 846, nomen nudum. Seriphidomyia Fedotova, 2000: 1421. Type species, butakovi Fedotova, 2000 (orig. des.). Polynomyia, as subg. of Seriphidomyia Fedotova, 2001a: 59. Type species, Seriphidomyia tarbagataica Fedotova, orig. des. Yukawyx, as subg of Dichelonyx Fedotova, 2001b: 954. Type species, Dichelonyx ustjurtensis Fedotova, 1999a (orig. des.). Pupascleromyia Fedotova, 2001c: 1084. Type species, iliensis (orig. des.). Comm.: Cosmopolitan; 267 spp. They are associated with only 8 plant families. Most species induce complex galls on Asteraceae, chiefly on Anthemideae. Ref.: Gagné & Jaschhof, 2014 (cat.). chrysanthemi (Ahlberg, 1939: 276) (Diarthronomyia). Type locality: Europe (origin unknown, presumably from Asia with chrysanthemum trade). LT (Gagné, 1983: 68; USA: MI, East Lansing) M (NYSM). Chrysanthemum spp. (Asteraceae). Distr.: widespread Nearctic, widespread northern Europe, Chile, Colombia, Argentina, Hawaiian Is, and New Zealand. Refs.: Gagné, 1994: 93; Gagné, 2004: 241 (cat.); Gagné, 2010: 355 (cat.); Gagné & Jaschhof, 2014: 321 (cat.). CATALOGUE OF CECIDOMYIIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 35 Tribe Trotteriini Rübsaamen & Hedicke, 1926 Genus Trotteria Rübsaamen Choristoneura Rübsaamen, 1892: 342, preocc. Lederer, 1859. Type species, Cecidomyia obtusa Loew, 1850. Trotteria Kieffer, 1902: 561, new name for Choristoneura Rübsaamen. Comm.: Holarctic, Neotropical, Afrotropical; 23 spp. Most species are inquilines in galls induced by Asphondyliini. At least one specie is associated with Contarinia and other with Youngomyia. Ref.: Gagné & Jaschhof, 2014 (cat.). rivinae Wünsch, 1979: 143. Type locality: Colombia, Magdalena, Bahia Concha. HT F (SMNS). Rivina humilis (Phytolaccaceae). Distr.: Colombia (Magdalena (Bahia Concha)). Refs.: Gagné, 1994: 99; Gagné, 2004: 270 (cat.); Gagné, 2010: 399 (cat.); Gagné & Jaschhof, 2014: 360 (cat.). salvadorensis Möhn, 1963: 3. Type locality: El Salvador, San Vicente, km 81, nr Rio San Felipe, HT F (SMNS). Boerhaavia erecta (Sterculiaceae) and Ayenia pusilla (Nyctaginaceae). Distr.: El Salvador; Colombia (Magdalena (Mamatoco, Bahia Concha, Bahia Nenguange)). Refs.: Gagné, 1994: 99; Gagné, 2004: 270 (cat.); Gagné, 2010: 399 (cat.); Gagné & Jaschhof, 2014: 360 (cat.). TABLE 1. Cecidomyiidae species recorded from Colombia. Subfamily Genus Species Epithet Author Bremia mirifica Gagné, 1994 Asphondylia blechi Wünsch, 1979 boerhaaviae Möhn, 1959 camarae Möhn, 1959 canavaliae Wünsch, 1979 caprariae Wünsch, 1979 duplicornis Wünsch, 1979 evae Wünsch, 1979 lopezae Wünsch, 1979 portulacae Möhn, 1959 sidae Wünsch, 1979 Cecidomyiinae yukawai Wünsch, 1979 Hemiasphondylia mimosae Möhn, 1960 Rhoasphondylia sanpedri Wünsch, 1979 Schizomyia manihoti Tavares, 1925 Contarinia gossypii Felt, 1908a Prodiplosis longifila Gagné, 1986 Feltiella insularis (Felt, 1913a) Lestodiplosis gagnei Baylac, 1987 Trisopsis incisa (Felt, 1907) oleae Kieffer, 1912b coccidarum Cockerell, 1892 vaupedis (Harris, 1968) fernandi Wünsch, 1979 Alycaulus trilobatus Möhn, 1964 Geraldesia polygonarum Wünsch, 1979 Meunieriella datxeli Wünsch, 1979 gairae Wünsch, 1979 Diadiplosis Moehniella ...Continued on next page 36 · Zootaxa 4122 (1) © 2016 Magnolia Press MAIA TABLE 1. (Continued) Subfamily Genus Neolasioptera Species Epithet Author magdalenae Wünsch, 1979 cruttwellae Gagné, 1977 camarae Möhn, 1964 cusani Wünsch, 1979 diclipterae Wünsch, 1979 donamae Wünsch, 1979 mincae Wünsch, 1979 olivae Wünsch, 1979 portulacae (Cook, 1906) samariae Wünsch, 1979 tribulae Wünsch, 1979 boerhaaviae Möhn, 1975 Domolasioptera acuario Wünsch, 1979 Rhopalomyia chrysanthemi (Ahlberg, 1939) Trotteria rivinae Wünsch, 1979 salvadorensis Möhn, 1963 Camptoneuromyia References Ahlberg, O. (1939) Diarthronomyia chrysanthemi nom. nov. (hypogaea Felt nec Löw). Entomologisk Tidskrift, 60, 274–278. Barnes, H.F. (1939) Some gall midge species and their host plant range. Annals of Applied Biology, 26, 318–347, pl.XXII. http://dx.doi.org/10.1111/j.1744-7348.1939.tb06974.x Baylac, M. (1987) Description d'un Lestodiplosis nouveau, prédateur d'Elaeidobius subvittatus (Col. Curculionidae), pollinisateur du palmier à huile Elaeis guineensis en Colombie (Dipt. Cecidomyiidae). 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Research Branch, Agriculture Canada, Hull, Quebec, pp. 1333–1370. Wünsch, A. (1979) Gallenerzeugende Insekten nordkolumbiens, speziell Asphondyliidi und Lasiopteridi (Diptera, Cecidomyiidae) aus dem Küstenbereich um Santa Marta. Wünsch, Ludwigsburg-Hoheneck, 238 pp. 40 · Zootaxa 4122 (1) © 2016 Magnolia Press MAIA Zootaxa 4122 (1): 041–045 http://www.mapress.com/j/zt/ Copyright © 2016 Magnolia Press Article ISSN 1175-5326 (print edition) ZOOTAXA ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.4122.1.7 http://zoobank.org/urn:lsid:zoobank.org:pub:7F481715-CFDC-4141-8620-2CA934E6FBC6 FAMILY SCIARIDAE SHEILA PATRÍCIA CARVALHO-FERNANDES Department of Entomology, Museu Nacional, Universidade Federal do Rio de Janeiro, Quinta da Boa Vista, São Cristóvão, 20940040, Rio de Janeiro, RJ, Brazil. E-mail: sheilapcfernandes@gmail.com Abstract Sciaridae are a widely distributed family with high number of species. They are known as black fungus gnats due to their dark color and feeding activity. This catalogue presents 17 species from Colombia distributed in eight genera, and for each species the geographical distribution is provided. Key words: black fungus gnats, catalogue, Colombia, Sciaridae Introduction Sciaride are a large and cosmopolitan family, commonly known as “black fungus gnats”. Of the 2400 described species, about 29 genera and 193 species are found in the neotropics (Amorim et al. 2002). They are tinny and fragile flies, 1.0–6.0 mm long, usually blackish, yellowish or brownish. Head ovoid, compound eyes meeting above the base of antennae forming dorsal bridge. Antennae long with 14 flagellomeres, scape and pedicel rounded, three ocelli present; palpus usually with three segments. Wing hyaline, brownish or darkwish. C short, Sc weak, veins M and Cu thinner, one cross-vein r-m, and CuA stem short. Foretibia with apical spur, mid and hindtibia with two apical spurs. Tarsal claws simple or toothed. Abdomen cylindrical, tapered posteriorly in females, and an exposed genitalia in males. Larvae have soft skin, white, with a translucent 12-segmented body, and a black head (Mohrig & Menzel 2009). The immatures feed upon many organic matter in the soil and plant litter, besides mine in stem or leaves, and live under the bark of live trees. Some species have been found in caves, animal burrows, ant colonies, and bird nests. Some genera are known as pests in mushroom and plant cultivation (Mohrig & Menzel 2009). Sciaridae represent a monophyletic family based on characteristics from larval head, but the classification of the family into subfamilies is controversial among the authors and is not used here. Sciaridae are probably a sister group of Mycetophilidae rather than of Cecidomyiidae due to similarity of the larval mouthparts and the loss of the eighth abdominal spiracle in the larvae (Mohrig & Menzel 2009). In this catalogue all Sciaridae species recorded from Colombia are summarized and listed (Table 1). The catalogue of Amorim (1992) was adopted for the classification and nomenclature. Recent publications concerning taxonomic changes were considered. All the distributional data available are provided. Acronyms used for the depositories BMNH—British Museum (Natural History), London, United Kingdom. BZM—Berliner Zoologishes Museum, Berlin, Germany. CUC—Cornell University Collection, Cornell, USA. IZV—Instytut Zoologii, Warsaw, Poland. NMW—Naturhistorisches Museum, Vienna, Austria. Accepted by S. Nihei: 25 May 2015; published: 14 Jun. 2016 41 List of abbreviations cat.—catalogue distr.—geographic distribution mon.—monotypy n. comb.—new combination nom. nov.—nomen novum (new name) orig. des.—by original designation preocc.—preoccupied redescr.—redescription refs.—references spp.—species subg.—subgenus subs. des.—subsequent designation syn.—synonym HT—Holotype LT—Lectotype PL—Paralectotype ST—Syntype T—Type F (f#)—Female M (m#)—Male Catalogue of Sciaridae of Colombia Family Sciaridae Billberg, 1820 Genus Bradysia Winnertz Bradysia Winnertz, 1867: 180. Type species, Bradysia angustipennis Winnertz (subs. des. by Enderlein, 1911: 127). Refs.: Tuomikoski, 1960: 110 (redefinition); Lane, 1959a (Key to spp.) Amorim, 1992: 57 (cat.). Neosciara Pettey, 1918: 320. Type species, Sciara coprophila Lindner (orig. des.) (= Bradysia amoena (Winnertz)). aequalis (Rübsaamen), 1894: 36 (Sciara). Type locality: Colombia, Bogotá. HT F (BZM). Distr.: Colombia (Bogotá). Refs.: Amorim, 1992: 57 (cat). columbia (Pettey), 1918: 342 (Neosciara). Type locality: Colombia, La Sierra. HT M (CUC). Distr.: Colombia (Santa Marta (La Sierra)). Refs.: Amorim, 1992: 57 (cat.). Genus Hybosciara Rübsaamen Hybosciara Rübsaamen, 1894: 28. Type species, Hybosciara gigantea Rübsaamen (mon.) (= Sciara gigantea Macquart). Refs.: Amorim, 1992: 61 (cat.). gigantea (Macquart), 1844: 147 (1846: 19) (Sciara). Type locality: “Nouvelle-Grenade”. ST FF (not located (“De la collection de M. Bigot e de la mienne”)). Refs.: Enderlein, 1911: 146 (as synonym of H. gigantea Rübsaamen) Amorim, 1992: 61 (cat.). gigantea Rübsaamen, 1894: 28. Type locality: “Colombia”. LT F (BZM). 42 · Zootaxa 4122 (1) © 2016 Magnolia Press CARVALHO-FERNANDES Genus Merianina Frey Merianina Frey, 1942: 29. Type species, Merianina americana Frey (orig. des.). Refs.: Amorim, 1992: 62 (cat.). leptogaster (Schiner), 1868: 13 (Sciara). Type locality: “Colombia”. LT F (NMW). Refs.: Amorim, 1992: 62 (cat.). prosthioxantha (Enderlein), 1911: 167 (Lycoria). Type locality: Colombia, Rio Magdalena. HT F (IZV). Distr.: Colombia (Magdalena (Rio Magdalena)). Refs.: Amorim, 1992: 62 (cat.). Genus Novaniarella Özdikmen & Başar Aniarella Enderlein, 1911; preocc. Bolivar, 1906. Type species, Aniarella pelluscens Enderlein, 1911 (orig. des.). Novaniarella Özdikmen & Başar, 2011: 15 (nom. nov. for Aniarella Enderlein). Refs.: Özdikmen & Başar, 2011 brevis (Rübsaamen, 1894): 36 (Sciara). Type locality: Colombia, Facatativa. LT M (BZM). Distr.: Colombia (Cundinamarca (Facatativa)). Refs.: Özdikmen & Başar, 2011: 15. pimpliformis Enderlein, 1911: 162 (Lycoriella); Refs.: Amorim, 1992: 56 (syn.) nigrina Rubsaamen, 1894: 35 (Sciara). Refs.: Enderlein, 1911: 141 (n. comb.); Amorim, 1992: 56 (syn.). Genus Pseudosciara Schiner Pseudosciara Schiner, 1868: 13; Lane, 1959b. Type species, Pseudosciara hirtella Schiner (mon.). Refs.: Edwards, 1934: 367 (synonym of Megalosphys); Amorim, 1992: 63 (cat.). Megalosphys Enderlein, 1911: 129. Type species, Megalosphys luteicoxa Enderlein (orig. des.). hirtella Schiner, 1868: 14. Type locality: “Colombia”. T M, lost. Refs.: Amorim, 1992: 64 (cat.). hirtipes (Enderlein), 1911: 132 (Megalosphys). Type locality: Colombia, Hacienda Pehlke (Depto. de Caldas). LT F (IZV). Distr.: Colombia (Caldas (Victoria, Hacienda Pehlke)), Panama. Refs.: Amorim, 1992: 64 (cat.). nigricolor (Enderlein), 1911: 133 (Megalosphys). Type locality: “Colombia” LT F (IZV). Refs.: Amorim, 1992: 64 (cat.). paradoxa (Pettey), 1918: 341, (Sciara); Lengersdorf, 1940: 247 (as a Pseudosciara). Type locality: Colombia, Caquetá, Valle de Papas. HT M (CUC). Distr.: Colombia (Caquetá (Valle de Las Papas)). Refs.: Amorim, 1992: 64 (cat.). Genus Rhynchosciara Rübsaamen Rhynchosciara Rübsaamen, 1894: 29. Type species, Rhynchosciara villosa Rübsaamen (subs. des. by Coquillett, 1910: 601) (= cognata Walker, 1848). Refs.: Breuer, 1969 (rev.), Amorim, 1992: 65 (cat.). busaccai Breuer, 1969: 183. Type locality: Costa Rica, Irazú. HT F (BMNH). Distr.: Colombia, Mexico, Guatemala, Nicaragua, El Salvador, Costa Rica. americana of Osten-Sacken (non Wiedemann), 1886: 1 (part, F). cognata (Walker), 1848: 103 (Sciara). Type locality: Colombia, Bogotá. HT F (BMNH). Distr.: Colombia (Bogotá), Ecuador. Refs.: Amorim, 1992: 65 (cat.). americana of Wulp (non Wiedemann), 1891: 193 (F). villosa of Rübsaamen, 1894: 29; Breuer, 1969 (redescr.). LT F (BZM). CATALOGUE OF SCIARIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 43 pleciodes (Enderlein), 1911: 158 (Lycoria). Type locality: Ecuador, Balzapamba. LT F (IZV). Distr.: Colombia, Ecuador. Refs.: Amorim, 1992: 66 (cat.). vespertilio (Schiner), 1868: 12. Type locality: “Venezuela”. LT M (NMW). Distr.: Venezuela, Colombia. Refs.: Amorim, 1992: 66 (cat.). Genus Schwenkfeldina Frey Neosciara, subg. Schwenkfeldina Frey, 1942: 32; Type species, Sciara carbonaria Meigen (orig. des.). Refs.: Amorim, 1992: 66 (cat.). Frey, 1948: 51 (as a subgenus of Bradysia); Tuomikoski, 1960: 29 (as a genus). moebiusi (Rübsaamen), 1894: 33 (Sciara). Type locality: “Colombia”. HT F (BZM). Distr.: Venezuela, Colombia. Refs.: Amorim, 1992: 67 (cat.). Genus Sciara Meigen Lycoria Meigen, 1800: 17. Type species, Tipula thomae Linnaeus (subs. des. by Coquillett, 1910: 563). Supressed by I.C.Z.N. (1963: 339). Sciara Meigen, 1803: 263. Type species, Tipula thomae Linnaeus (mon.). Refs.: Amorim, 1992: 67 (cat.). columbiana (Enderlein), 1911: 173 (Lycoria). Type locality: “Colombia”. LT M, PL F (IZV). Distr.: Colombia (Rio Magdalena). Refs.: Amorim, 1992: 67 (cat.). Unplaced species of Sciaridae marginalis Röder, 1886: 258 (Sciara). Type locality: Colombia, Bogotá. Type not located. Unrecognizable. Distr.: Colombia (Bogotá). Refs.: Amorim, 1992: 69 (cat.). TABLE 1. Sciaridae species recorded from Colombia. Genus Species Author Bradysia aequalis (Rübsaamen, 1894) columbia (Pettey, 1918) Hybosciara gigantea (Macquart, 1844) Merianina leptogaster (Schiner, 1868) prosthioxantha (Enderlein, 1911) Novaniarella brevis (Rübsaamen, 1894) Pseudosciara hirtella Schiner, 1868 hirtipes (Enderlein, 1911) nigricolor (Enderlein, 1911) paradoxa (Pettey, 1918) busaccai Breuer, 1969 cognata (Walker, 1848) pleciodes (Enderlein, 1911) vespertilio (Schiner, 1868) Schwenkfeldina moebiusi (Rübsaamen, 1894) Sciara columbiana (Enderlein, 1911) Unplaced marginalis Röder, 1886 Rhynchosciara 44 · Zootaxa 4122 (1) © 2016 Magnolia Press CARVALHO-FERNANDES References Amorim, D.S. (1992) A catalogue of the family Sciaridae (Diptera) in the Americas South of the United States. Revista Brasileira de Entomologia, 36 (1), 55–77. Amorim, D.S., Silva, V.C. & Balbi, M.I.P.A. (2002) Estado do conhecimento dos Diptera Neotropicais. 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(1934) New Neotropical Mycetophilidae (III) (Diptera). Revista de Entomologia, 4, 354–372. Frey, R. (1942) Entwurf einer neuen Klassifikation der Mückenfamilie Sciaridae (Lycoriidae). Notulae Entomologicae, 22, 5–44. Frey, R. (1948) Entwurf einer neuen Klassifikation der Mückenfamilie Sciaridae (Lycoriidae). II. Die nordeuropäischen Arten. Notulae Entomologicae, 27, 33–112. Lane, J. (1959a) On Neotropical Neosciara (Diptera, Sciaridae). Studia Entomologica, 2, 69–104. Lane, J. (1959b) On Neotropical “Pseudosciara” (Diptera, Sciaridae). Revista Brasileira de Biologia, 19, 287–298. Lengersdorf, F. (1940) Beitragzur Kenntnis der Lycoriiden (Sciariden) in Mexiko. 1 Teil. Arbeiten über morphologische und taxonomische Entomologie, 7, 245–251. Macquart, J. (1844) Diptères exotiques nouveaux ou peu connus. Mémoires de la Société royale de sciences, de l'agriculture et des arts de Lille, 1844, 133–164. [1845] Meigen, J.W. (1800) Nouvelle classification des mooches à deu xailes (Diptera L.) d’apres um plant tout nouveau. Paris, 40 pp. Meigen, J.W. (1803) Versuch einer neuen Gattungseintheilung der europäischen zweiflügeligen Insekten. Magazine fur Insektenkunde, 2, 259–281. Mohrig, W. & Menzel, F. (2009) Sciaridae (Black Fungus Gnats). In: Brown, B.V., Borkent, A., Cumming, J.M., Wood, D.M., Woodley, N.E. & Zumbado, M.A. (Eds.), Manual of Central American Diptera. Vol. 1. NRC Research Press, Ottawa, pp. 279–292. Özdikmen, H. & Başar, K. (2011) A new name for the Neotropical genus Aniarella Enderlein (Diptera, Sciaridae). Revista Brasileira de Entomologia, 55 (1), 15–16. Pettey, F.W. (1918) A revision of the genus Sciara of the family Mycetophilidae (Diptera). Annals of the Entomological Society of America, 11, 319–343. http://dx.doi.org/10.1093/aesa/11.4.319 Röder, V. von (1886) Dipteren von den Cordilleren in Collumbien gesammelt durch Herrn Dr. Alpheris Stübel. Stettiner Entomologische Zeitung, 47, 257–270. Rübsaamen, E.H. (1894) Die aussereuropäischen Trauermücken des Königl. Museums für Naturkunde zu Berlin. Berliner Entomologische Zeitschrift, 39, 17–42. http://dx.doi.org/10.1002/mmnd.18940390105 Schiner, J.R. (1868) Diptera. In: Wüllerstorf-Urbair, B. von (Ed.), Reise der österreichischen Fregatte Novara um die Erde in den Jahren 1857, 1858, 1859 unter den Befehlen des Commodore B. von Wüllerstorf-Urbair. Zoologischer Theil., Vol. 2, Abt. I. (Sect.). B. K. Gerold’s Son, Wien, pp. 388. Tuomikoski, R. (1960) Zur Kenntnis der Sciariden (Dipt.) Finnlands. Suom. El-inia Kasvitiet Seura Vanamo. El-intiet. Julkaisu. Annales Zoologici Societatis Zoologicae Botanicae Fennicae Vanamo, 21 (4), 1–164. Walker, F. (1848–1849) List of the specimens of dipterous insects in the collection of the British Museum. Vol. 1–4. Printed by order of the Trustees, London, 1172 pp. [Vol. 1., pp. 1–229 (1848); Vol. 2., pp. 231–484 (1849); Vol. 3., pp. 485–687 (1849); Vol. 4., pp. 689–1172 (1849)] Winnertz, J. (1867) Beitrag zu einer Monographie der Sciarinen. Verhandlungen des Zoologischbotanische Gesellschaft in Wien, 17, 1–187. CATALOGUE OF SCIARIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 45 Zootaxa 4122 (1): 046–049 http://www.mapress.com/j/zt/ Copyright © 2016 Magnolia Press Article ISSN 1175-5326 (print edition) ZOOTAXA ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.4122.1.8 http://zoobank.org/urn:lsid:zoobank.org:pub:31511FCC-9362-4A29-8AB1-863581B22BEB FAMILY RANGOMARAMIDAE RAFAELA LOPES FALASCHI1* & DALTON DE SOUZA AMORIM2 1 Universidade de São Paulo, Museu de Zoologia. Av. Nazaré, 481. Ipiranga. 04263-000. São Paulo SP Brazil. Departamento de Biologia, Faculdade de Filosofia, Ciências e Letras de Ribeirão Preto, Universidade de São Paulo. Av. Bandeirantes, 3900. 14040-901. Ribeirão Preto SP Brazil. dsamorim@usp.br, CNPq Research Fellowship. *Corresponding author: rlfalaschi@gmail.com 2 Abstract This small family of Diptera is poorly known in Colombia. In the present catalogue, we feature a single report for the country, of a species of the genus Eratomyia. Key words: Bibionomorpha, Catalogue, Colombia, distribution, diversity, Rangomaramidae Introduction The family Rangomaramidae was erected by Jaschhof and Didham (2002) for five New Zealand species of the genus Rangomarama Jaschhof and Didham. The concept of the family was revised by Amorim and Rindal (2007) based on a phylogenetic study of the Mycetophiliformia. A number of genera of controversial position in Bibionomorpha fit, in the analysis, in three clades that gathered with Rangomarama. The Rangomaramidae now would include Rangomarama in a subfamily of its own, Rangomaraminae, Heterotricha Loew as the single genus of Heterotrichinae, and two other subfamilies, Chiletrichinae, including Chiletricha Chandler and related genera, and Ohakuneinae, including Ohakunea Tonnoir and Edwards and related genera. These clades are obviously related in terms of plesiomorphies to a number of genera known from Jurassic deposits and to the family Archizelmiridae (Grimaldi et al. 2003). The study of Amorim and Rindal (2007) was carefully reviewed by Jaschhof (2011) and many of the inferences were questioned based on arguments concerning methods and characters. But a reanalysis of the higher Bibionomorpha with more data, taxa and another supposedly more thorough method is still wanted. Six genera and twenty species have been recorded for the Neotropics in the family (Amorim & Falaschi 2012). They belong to the genera Chiletricha, Eratomyia Amorim and Rindal, Ohakunea, Colonomyia Colless, Cabamofa Jaschhof and Rogambara Jaschhof. In the Ohakuneinae, Cabamofa and Rogambara are both monotypic and known only from Costa Rica (Jaschhof & Hippa 2003), there is one species of Ohakunea know from Chile, and there are seven species of Colonomyia from Chile, Argentina, Brazil, and Costa Rica (Matile & Duret 1994; Hippa & Jaschhof 2004; Amorim & Rindal 2007). In the Chiletrichinae, there are eight species of Chiletricha, known from Chile, Argentina, and southern Brazil, and two species of Eratomyia, the type-species from Ecuador, and an additional species from Colombia (Amorim & Falaschi 2010). The flies of this family have body size ranging from 1.7 to 6.0 mm and wing length varying from 2.0 to 5.5 mm. The body color is usually shining or dull brown, sometimes with yellowish areas. Usually there are three ocelli, but sometimes the lateral ocelli are displaced laterally, but never come into contact with the eye margin—the ocelli are absent in Rogambara and Cabamofa. An eye-bridge is absent or incomplete in most genera, being complete only in Rogambara and Cabamofa. The antenna has 14 flagellomeres, the first article usually 1.5 the length of the second flagellomere, with setae haphazardly distributed. The maxillary palpus has the palpiger and four additional palpomeres, the last segment much longer than preceding ones in the Chiletrichinae, Ohakunea, and Colonomyia. The thorax is more or less elongate (not so much in Cabamofa). The primary anapleural suture and 46 Accepted by S. Nihei: 10 Apr. 2015; published: 14 Jun. 2016 midpleural pit are present and the katepisternum is not particularly developed. Anepisternal and katepisternal setae are absent, with few exceptions. The mesopleurotrochantin is not visible. The mesepimeron is usually high and slender, in some cases obliterated in the ventral half due to fusion between katepisternum and laterotergite. Mesepimeral setae are present only in Eratomyia. The laterotergite is convex posteriorly into the mediotergite, slightly projected laterally, and laterotergite setae are present only in Heterotricha. The mediotergite is high, slightly curved, entirely bare. The metepisternum is normally produced, small, the postnotal phragma not being particularly reduced. The halter has a long pedicel. All coxae are elongated, with the tibial spurs 1:2:2, but the spurs not as long as seen in ditomyiids or most mycetophilids. The front tibia has a differentiated area with a regular row of setulae, except in Chiletricha. The wing is elongated, the macrotrichia on the membrane being either present or absent. Sc is incomplete, with the possible exception of Rangomarama. The base of Rs is oblique or transverse, long only in Rangomarama, while r-m can be nearly transverse (e.g., in Chiletricha), oblique backwards (e.g., Eratomyia or Rhynchoheterotricha Freeman) or longitudinal (in Ohakuneinae); r-m is particularly long in the Eratomyia and in the Ohakuneinae, but very short in Heterotricha, and entirely absent in Rangomarama. R4 always absent; R1 is usually long, reaching C before the mid of the wing length only in Colonomyia and Rogambara. M1+2 forks beyond the mid of the wing, the medial fork being particularly long in Heterotricha, Ohakunea, and Colonomyia. In most genera m-cu reaches CuA typically close to the wing base, with tb (M1+2/M3+4) longitudinal. The first sector of CuA is usually short, longer only in Eratomyia. A1 never reaches the wing margin, being entirely absent in Colonomyia, Cabamofa, and Rogambara. The abdomen is only slightly more slender at its base. The abdominal tergites and sternites 1-7 are normally sclerotized. Segment 8 is well developed, only slightly shorter than preceding segments. The male gonocoxites, gonostyles, tergite 9, and cercus are considerably variable between the subfamily and genera. In females, the first cercomere is shorter than the second in Heterotricha and Chiletricha, but longer than the second in Rangomarama. Two sclerotized spermathecae are present in all genera for which information is available. Acronyms used for the depositories IAvH—Alexander von Humboldt Biological Resources Research Institute, Bogota, Colombia. MZSP—Museu de Zoologia da Universidade de São Paulo, São Paulo, Brazil. List of abbreviations cat.—catalogue des.—designation distr.—geographic distribution i.e.—example m.—meters mon.—monotypy nom. nov.—nomen novum (new name) orig. des.—by original designation pp.—pages refs.—references rev.—revision sp., spp.—species ssp.—subspecies subs.—subsequent subg.—subgenus HT—Holotype LT—Lectotype NA—Not applicable NT—Neotype CATALOGUE OF RANGOMARAMIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 47 PT—Paratype ST—Syntype T—Type (unspecified) F (f#)—Female M (m#)—Male Catalogue of Rangomaramidae of Colombia Family Rangomaramidae Jaschhof & Didham, 2002 Subfamily Chiletrichinae Amorim & Rindal, 2007 Genus Eratomyia Amorim & Rindal Eratomyia Amorim & Rindal, 2007: 14. Type species, magnifica Amorim & Rindal, 2007 (orig. des.). risaralda Amorim & Falaschi, 2010: 56, (M terminalia), (F terminalia). Type locality: Colombia, Risaralda SFF, Otún Quimbaya, El Molinillo, 2,220 m, HT M (IavH). Refs.: Amorim & Falaschi, 2012: 3 (cat.). Material examined. COLOMBIA: Risaralda: SFF [Santuario de Fauna y Flora], Otún, Quimbaya, El Molinillo, 04º43’S 75º34’W, 2220 m, 1 male HT, 2 females PT, 17.ii–04.iii.2003, G. López Leg., (IavH, MZSP). Acknowledgements The authors are indebted to Michael Sharkey (National Science Foundation Grant DEB0205982), to whom we are most grateful. Additional information on the project can be found at the web address of the project (http:// www.sharkeylab.org/biodiversity/static.php?app=colombia&page=index). During the preparation of this paper, the authors received financial support from FAPESP grant 2013/16524-9 (RLF) and CNPq grants 159016/2012-6 (RLF) and 314371/2009-5 (DSA). TABLE 1. Rangomaramidae species recorded from Colombia. Subfamily Genus Species Author Chiletrichinae Eratomyia risaralda Amorim & Falaschi, 2010 References Amorim, D.S. & Rindal, E. (2007) A phylogenetic study of the Mycetophiliformia, with creation of the subfamilies Heterotrichinae,Ohakuneinae, and Chiletrichinae for the Rangomaramidae (Diptera, Bibionomorpha). Zootaxa, 1535, 1–92. Amorim, D.S. & Falaschi, R.L. (2010) Second known species of Eratomyia Amorim & Rindal (Diptera, Rangomaramidae, Chiletrichinae), from Colombia. Zootaxa, 2641, 55–61. Amorim, D.S. & Falaschi, R.L. (2012) Catalogue of Neotropical Diptera. Rangomaramidae. Neotropical Diptera, 21, 1–7. Freeman, P. (1951) Mycetophilidae. In: Diptera of Patagonia and South Chile. Part III (Fascicle 1). The British Museum (Natural History), London, pp. 138, plates I–XLIX. Grimaldi, D., Amorim, D.S. & Blagoderov, V. (2003) The Mesozoic family Archizelmiridae (Diptera: Insecta). Journal of Paleontology, 77 (2), 368–381. http://dx.doi.org/10.1666/0022-3360(2003)077%3C0368:TMFADI%3E2.0.CO;2 Hippa, H. & Jaschhof, M. (2004) A re-evaluation of the genus Colonomyia Colless (Diptera, Sciaroidea), with the description of two new species. Insect Systematics and Evolution, 35, 335–352. http://dx.doi.org/10.1163/187631204788920220 48 · Zootaxa 4122 (1) © 2016 Magnolia Press FALASCHI & AMORIM Jaschhof, M. (2011) Phylogeny and classification of the Sciaroidea (Diptera: Bibionomorpha): Where do we stand after Amorim & Rindal (2007). Beiträge zur Entomologie, 61 (2), 455–463. Jaschhof, M. & Didham, R.K. (2002) Rangomaramidae fam. nov. from New Zealand and implications for the phylogeny of the Sciaroidea (Diptera: Bibionomorpha). Studia Dipterologica, 11 (Supplement), 1–60. Jaschhof, M. & Hippa, H. (2003) Sciaroid but not sciarid: a review of the genus Ohakunea Tonnoir & Edwards, with the description of two new species (Insecta: Diptera: Bibionomorpha). Entomologische Abhandlungen, 60, 23–44. Matile, L. & Duret, J.P. (1994) Le genre Colonomyia Colless découvert en région Néotropicale (Diptera, Mycetophiloidea). Revue Française d'Entomologie (n.s.), 16 (4), 143–147. CATALOGUE OF RANGOMARAMIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 49 Zootaxa 4122 (1): 050–052 http://www.mapress.com/j/zt/ Copyright © 2016 Magnolia Press Article ISSN 1175-5326 (print edition) ZOOTAXA ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.4122.1.9 http://zoobank.org/urn:lsid:zoobank.org:pub:45B943B9-E679-4507-9D5D-8D9D2361BE82 FAMILY DITOMYIIDAE RAFAELA LOPES FALASCHI Universidade de São Paulo, Museu de Zoologia.Av. Nazaré, 481. Ipiranga. 04263-000. São Paulo SP Brazil. E-mail: rlfalaschi@gmail.com Abstract The Ditomyiidae are a family belonging to the suborder Bibionomorpha, until now without records to Colombia. This work reports the first record of the genera Rhipidita and Calliceratomyia for the Colombian region. Key words: Bibionomorpha, Catalogue, Colombia, distribution Introduction Ditomyiidae are a family of gnats in the suborder Bibionomorpha, with eight genera and more than 100 species described worldwide (Vockeroth 2009), except in the Afrotropical region. In the Neotropical region, four genera and 35 species are recorded (Munroe 1974, Papavero 1978; Amorim et al. 2002; Huerta & Amorim 2016), none of them known from Colombia. For the first time the genus Rhipidita Edwards was collected with two new species (Falaschi & Amorim, in prep.) and the genus Calliceratomyia Lane, with four new species (Falaschi et al., in prep.). Adults are found on vegetation in primary forests. As pointed out by Munroe (1974), the wood- or hard fungus-boring habits of the larvae make extremely difficult to study the life history of these organisms in detail. Immature stages of ditomyiids are currently known only for the Holarctic genera Symmerus Walker and Ditomyia Winnertz, and the Chilean-Australian genus Australosymmerus Freeman (Matile 1990). Larvae are associated with bracket fungus (Basidiomycota: Polyporaceae) in rotten wood (Munroe 1974), where they build their galleries. Ditomyiids are medium-size gnats (3.0-8.0 mm), and can be recognized by the following characters: postpronotum with one or more long bristles, veins M1+2 and M3+4 basally connected (m-cu), basal portion of M absent and R4 present, long, well over half the length of the apical part of R5, Sc weak and ending free. Acronyms used for the depositories IAvH—Alexander von Humboldt Biological Resources Research Institute, Bogota, Colombia MZSP—Museu de Zoologia da Universidade de São Paulo, São Paulo, Brazil List of abbreviations cat.—catalogue col.—collector des.—designation distr.—geographic distribution i.e.—example m.—meters mon.—monotypy 50 Accepted by S. Nihei: 10 Apr. 2015; published: 14 Jun. 2016 nom. nov.—nomen novum (new name) orig. des.—by original designation pp.—pages refs.—references rev.—revision sp., spp.—species ssp.—subspecies subs.—subsequent subg.—subgenus HT—Holotype NT—Neotype PT—Paratype LT—Lectotype ST—Syntype T—Type (unspecified) NA—Not applicable F (f#)—Female M (m#)—Male Catalogue of Ditomyiidae of Colombia Family Ditomyiidae Keilin, 1919 Genus Rhipidita Edwards Rhipidita Edwards, 1940: 441. Type species, fusca (orig. des.). undescribed sp. 1.. Distr.: Colombia (Nariño). Material examined. COLOMBIA: Nariño: RN [Reserva Natural], La Planada, Parcela Permanente, 01º15’N 78º15’W, 2 males, 1885 m, 01–16.xii.2001, G. Oliva Leg., M. 2740, Malaise (IAvH, MZSP). undescribed sp. 2.. Distr.: Colombia (Norte de Santander). Material examined. COLOMBIA: Norte de Santander: ANU [Área Natural Única], Los Estoraques, Bosque Piritama, 08º14’N 73º15’W, 1850 m, 2 males, 27.iv–25.v.2004, J. Vargas & E. Bayona Leg., Malaise (IAvH, MZSP). Genus Calliceratomyia Lane Calliceratomyia Lane, 1946: 244. Type species, pectinata (orig. des.). undescribed sp. 1.. Distr.: Colombia (Risaralda). Material examined. COLOMBIA: Risaralda: SFF [Santuario de Fauna y Flora], Otún, Quimbaya, Cuchilla Camino, 04º43’N 73º35’W, 2050 m, 3 males, 04–14.iv.2003, G. López Leg., Malaise (IAvH, MZSP). undescribed sp. 2.. Distr.: Colombia (Risaralda). Material examined. COLOMBIA: Risaralda: SFF [Santuario de Fauna y Flora], Otún, Quimbaya, El Molinillo, 04º43’N 75º34’W, 14 males, 2220 m, 04–17.ii.2003, G. López Leg., Malaise (IAvH, MZSP). undescribed sp. 3.. Distr.: Colombia (Norte de Santander). Material examined. COLOMBIA: Norte de Santander: ANU [Área Natural Única], Los Estoraques, CATALOGUE OF DITOMYIIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 51 Bosque Piritama, 08º14’N 73º15’W, 1850 m, 3 males, 27.iv–25.v.2004, J. Vargas & E. Bayona Leg., Malaise (IAvH, MZSP). undescribed sp. 4.. Distr.: Colombia (Risaralda). Material examined. COLOMBIA: Risaralda: SFF [Santuario de Fauna y Flora], Otún, Quimbaya, Cuchilla Camino, 04º43’N 73º35’W, 2050 m, 2 males, 04–14.iv.2003, G. López Leg., Malaise (IAvH, MZSP). Acknowledgements The author is indebted to Michael Sharkey (National Science Foundation Grant DEB0205982), to whom I am most grateful. Additional information on the project can be found at the web address of the project (http:// www.sharkeylab.org/biodiversity/static.php?app=colombia&page=index). During the preparation of this paper, the author received financial support from FAPESP grant 2013/16524-9 and CNPq grant 159016/2012-6. TABLE 1. Ditomyiidae species recorded from Colombia. Genus Species Rhipidita sp. 1 Author sp. 2 Calliceratomyia sp. 1 sp. 2 sp. 3 sp. 4 References Amorim, D.S., Silva, V.C. & Balbi, M.I.P.A. (2002) Estado do conhecimento dos Diptera neotropicais. In: Costa, C., Vanin, S.A., Lobo, J.M. & Melic, A. (Eds.), Proyecto de Red Iberoamericana de Biogeografia y Entomología Sistemática, PRIBES 2002i. Sociedad Entomológica Aragoneza y CYTED, Zaragoza, pp. 29–36. Edwards, F.W. (1940) A note on the subfamily Ditomyiinae, with description of new recent and fossil forms. Annals and Magazines of Natural History, 9 (7), 431–437. Falaschi, R.L. & Amorim, D.S. (2016) A Revision of the Neotropical genus Rhipidita Edwards, 1940 (Diptera: Ditomyiidae). [in preparation] Falaschi, R.L., Riccardi, P.R. & Amorim, D.S. (2016) Taxonomic revision of the genus Calliceratomyia Lane, 1946 (Diptera: Ditomyiidae), with the description of nine new species. [in preparation] Huerta, H. & Amorim, D.S. (2016) Three new species of Calliceratomyia Lane (Diptera: Ditomyiidae) from Neotropical Mexico. Zootaxa, 4072 (4), 430–440. http://dx.doi.org/10.11646/zootaxa.4072.4.2 Keilin, D. (1919) On the structure of the larvae and the systematic position of the genera Mycetobia, Mg., Ditomyia, Winn., and Symmerus, Walk. (Diptera Nematocera). Annals and magazine of natural history, 3 (9), 33–42. Lane, J. (1946) “Mycetophilidae” do Brasil (Diptera, Nemocera). Revista Brasileira de Zoologia, 8 (2), 247–254. Matile, L. (1990) Recherches sur la systématique et l’évolution des Keroplatidae (Diptera, Mycetophiloidea). Mémoires du Muséum national d’Histoire naturelle Paris, Série A (Zoologie), 148, 1–682. Munroe, D.D. (1974) The systematics, phylogeny, and zoogeography of Symmerus Walker and Australosymmerus Freeman (Diptera: Mycetophilidae: Ditomyiidae). Memoirs of the Entomological Society of Canada, 92, 1–183. http://dx.doi.org/10.4039/entm9217fv Papavero, N. (1978) 19B. Family Ditomyiidae. In: Papavero, N. (Ed.), A Catalogue of the Diptera of the Americas south of the United States. Departamento de Zoologia da Secretaria da Agricultura do Estado de São Paulo, São Paulo, pp. 6. Vockeroth, J.R. (2009) Ditomyiidae (Ditomyiid fungus gnats). In: Brown, B.V., Borkent, A., Cumming, J.M., Wood, D.M., Woodley, N.E. & Zumbado, M.A. (Eds.), Manual of Central American Diptera. Vol. 1. NRC Research Press, Ottawa, Ontario, pp. 253–255. 52 · Zootaxa 4122 (1) © 2016 Magnolia Press FALASCHI Zootaxa 4122 (1): 053–055 http://www.mapress.com/j/zt/ Copyright © 2016 Magnolia Press Article ISSN 1175-5326 (print edition) ZOOTAXA ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.4122.1.10 http://zoobank.org/urn:lsid:zoobank.org:pub:0FD00F12-697F-41B8-97CB-EAE1F0B380A0 FAMILY DIADOCIDIIDAE RAFAELA LOPES FALASCHI Universidade de São Paulo, Museu de Zoologia. Av. Nazaré, 481. Ipiranga. 04263-000. São Paulo SP Brazil. E-mail: rlfalaschi@gmail.com Abstract Diadocidiidae are a small family belonging to the suborder Bibionomorpha. Three species were described for the Neotropical region, two from Costa Rica and one from Brazil, and one additional undescribed species from Colombia is reported here for the first time. Key words: Bibionomorpha, catalogue, Colombia, distribution Introduction The Diadocidiidae correspond to a small family of Mycetophiliformia, in the suborder Bibionomorpha, with 24 extant species in two genera, including the living genus Diadocidia Ruthe, 1831, and a fossil genus from the Cretaceous amber from Katchin (Myamar), with one species, Docidiadia burmitica (Blagoderov & Grimaldi, 2004). The genus Diadocidia is divided into three subgenera: Adidocidia Laštovka and Matile, 1972, Diadocidia and Taidocidia Papp and Ševčík, 2005 (Blagoderov 2007; Bechev & Chandler 2011). The living species of the family are found in the Holarctic, Oriental, Australasian/Oceanian, and Neotropical regions. Little is known about the biology of these flies. Adults are found in shaded places in forests along streams and on tree trunks (Rindal & Gammelmo 2007). Larvae were reported to live in mucous tubes under rotting logs, associated with Polyporaceae fungi (Hutson et al. 1980, Sasakawa 2004), similarly to some Keroplatidae (Matile 1997). In such habitats, larvae feed on the hymenium of these fungi (Zaitzev 1994) and Chandler (1993) reported that a larva of Diadocidia ferruginosa (Meigen, 1830) was feeding on Peniophora sp., a Basidiomycota fungus belonging to the Peniophoraceae. Diadocidiids are medium size flies, with body length of 2.5–5.6 mm. Diagnostic features for the family are mostly plesiomorphies: three ocelli, antennae with scape and pedicel short, flagellum about 1.5 times as long as the combined length of head and thorax; antennae with 14 flagellomeres; thorax humpbacked; and macrotrichia widely distributed over the wing membrane. Gnats are easily recognized, however, by the wing vein pattern, unique in the Bibionomorpha: r-m and the stem of M3–M4 are almost perfectly aligned transversally in the wing. Diadocidia is widely distributed in the world, even though the number of species is not considered high. For the Neotropical region, Papavero (1977) mentioned a single species from Brazil and an undescribed species from Chile (Falaschi & Amorim, 2014). Jaschhof & Jaschhof (2007) recently added two species from Costa Rica (D. similis Jaschhof & Jaschhof, 2007 and D. hybrida Jaschhof & Jaschhof, 2007), whereas Vockeroth (2009) referred to an undescribed species from Mexico. The occurrence of the genus Diadocidia in Colombia is reported here for the first time (Table 1). Acronyms used for the depositories IAvH—Alexander von Humboldt Biological Resources Research Institute, Bogota, Colombia MZSP—Museu de Zoologia da Universidade de São Paulo, São Paulo, Brazil Accepted by S. Nihei: 10 Apr. 2015; published: 14 Jun. 2016 53 List of abbreviations cat.—catalogue col.—collector des.—designation distr.—geographic distribution i.e.—example m.—meters mon.—monotypy nom. nov. —— nomen novum (new name) orig. des.—by original designation pp.—pages refs.—references rev.—revision sp., spp.—species ssp.—subspecies subs.—subsequent subg.—subgenus HT—Holotype NT—Neotype PT—Paratype LT—Lectotype ST—Syntype T—Type (unspecified) NA—Not applicable F (f#)—Female M (m#)—Male Catalogue of Diadocidiidae of Colombia Family Diadocidiidae Winnertz, 1864 Genus Diadocidia Ruthe Diadocidia Ruthe, 1831: 1210. Type species, flavicans Ruthe, 1831 [= Mycetobia ferruginosa Meigen, 1830], by mon. Macronevra Macquart, 1834: 146. Type species, Macronevra winthemi Macquart, 1834 [= Diadocidia ferruginosa (Meigen, 1830)], by mon. Syn.: Winnertz (1852). Aclada Loew, 1850: 33, 35. Type species, Diadocidia parallela Evenhuis, 1994, subsequent designation. Aclada originally proposed without included species. The type species des. was from the first species included within the genus in accordance with the ICZN. (see Evenhuis, 1994 and Bechev & Chandler, 2011) Macroneura Rondani, 1856: 197, 214 (unjustified emendation of Macronevra Macquart, 1834). Palaeodocidia Sasakawa, 2004: 208. Type species, Palaeodocidia ishizakii Sasakawa, 2004, orig. des. undescribed sp. Distr.: Colombia (Risaralda). Material examined: COLOMBIA: Risaralda: SFF [Santuario Fauna y Flora], Otún, Quimbaya, Cuchilla Camino, 04º43’N 73º35’W, 3 males, 2050 m, 04–14.iv.2003, G. López col., Malaise, (IAvH, MZSP). Acknowledgements The author is indebted to Michael Sharkey (National Science Foundation Grant DEB0205982), to whom I am most grateful. Additional information on the project can be found at the web address of the project (http:// www.sharkeylab.org/biodiversity/static.php?app=colombia&page=index). During the preparation of this paper, the author received financial support from FAPESP grant 2013/16524-9 and CNPq grant 159016/2012-6. 54 · Zootaxa 4122 (1) © 2016 Magnolia Press FALASCHI TABLE 1. Diadocidiidae species recorded from Colombia. Genus Species Diadocidia sp. Author References Bechev, D. & Chandler, P. (2011) Catalogue of the Bolitophilidae and Diadocidiidae of the World (Insecta: Diptera). Zootaxa, 2741, 38–58. Blagoderov, V. & Grimaldi, D. (2004) Fossil Sciaroidea (Diptera) in Cretaceous ambers, exclusive of Cecidomyiidae, Sciaridae, and Keroplatidae. American Museum Novitates, 3433, 1–76. http://dx.doi.org/10.1206/0003-0082(2004)433%3C0001:FSDICA%3E2.0.CO;2 Blagoderov, V. (2007) Fungus Gnats Online. Available from: http://sciaroidea.info/content/diadocidiidae-0 (accessed 30 January 2014) Chandler, P. (1993) New rearing records of fungus gnats (Diptera: Mycetophilidae and allied families). Dipterists Digest, 13, 29–34. Evenhuis, N.L. (1994) Catalogue of the fossil flies of the world (Insecta: Diptera). Backhuys Publishers, Leiden, viii + 600 pp. Falaschi, R.L. & Amorim, D.S. (2014) 79. Diadocidiidae. In: Roig-Juñent, S.A., Claps, L.E., & Morrone, J.J. (Eds.), Biodiversidad de Artrópodos Argentinos. Vol. 4. 1st Edition. Editorial INSUE - Universidad Nacional de Tucumán, San Miguel de Tucumán, pp. 367–368. Hutson, A.M., Ackland, D.M. & Kidd, L.N. (1980) Mycetophilidae (Bolitophilinae, Ditomyiinae, Diadocidiinae, Keroplatinae, Sciophilinae and Manotinae) Diptera, Nematocera. Handbooks for the Identification of British Insects, 9 (3), 1–111. Jaschhof, M. & Jaschhof, C. (2007) On the genus Diadocidia (Diptera, Sciaroidea, Diadocidiidae) in Costa Rica. Zootaxa, 1586, 33–38. Laštovka, P. & Matile, L. (1972) Révision des Diadocidia Holarctiques [Dipt. Mycetophilidae]. Annales de la Societé entomologique de France, New Series, 8 (1), 205–223. Loew, H. (1850) Ueber den Bernstein und die Bernsteinfauna. Programm K. Realschule zu Meseritz, 1850, 1–44. Macquart, J. (1834) Histoire naturelle des insectes. Diptères. Ouvrage accompagné de planches. Tome première. Roret, N.E., Paris, 578 pp. +12 pls. Matile, L. (1997) Phylogeny and evolution of the larval diet in the Sciaroidea (Diptera, Bibionomorpha) since the Mesozoic. In: Grandcolas, P. (Ed.), The origin of biodiversity in insects: Phylogenetic tests of evolutionary scenarios. Mémoires du Muséum National d'Histoire Naturelle, Paris, 173, 273–303. Meigen, J.W. (1830) Systematische Beschreibung der bekannten europäischen zweiflügeligen Insekten. Sechster Theil. Schulz-Wundermann, Hamburg, xi + 401 + [3] pp. Papavero, N. (1977) 19A. Family Diadocidiidae. In: Papavero, N. (Ed.), A Catalogue of the Diptera of the Americas south of the United States. Departamento de Zoologia da Secretaria da Agricultura do Estado de São Paulo, São Paulo, pp. 3. Papp, L. & Ševčík, J. (2005) New taxa of Diadocidiidae (Diptera) from the Oriental region. Acta Zoologica Academiae Hungaricae, 51 (4), 329–341. Rindal, E. & Gammelmo, O. (2007) On the family Diadocidiidae (Diptera, Sciaroidea) in Norway. Norwegian Journal of Entomology, 54, 69–74. Rondani, C. (1856) Dipterologiae Italicae prodromus. Vol. I. Genera Italica ordinis dipterorum ordinatim disposita et distincta et in familias et stirpes aggregata. A. Stoschi, Parmae [= Parma], 226 + [2] pp. Ruthe, J.F. von (1831) Einige Bemerkungen und Nachträge zu Meigen's "Systematischer Beschreibung der europäischen zweiflügeligen Insecten". Isis (Oken's), 1831, 1203–1222. Sasakawa, M. (2004) Diadocidiidae and Borboropsidae (Insecta: Diptera) of Japan, with descriptions of two new species. Species Diversity, 9, 207–214. Vockeroth, J.R. (2009) Diadocidiidae (diadocidiid fungus gnats). In: Brown, B.V., Borkent, A., Cumming, J.M., Wood, D.M., Woodley, N.E. & Zumbado, M.A. (Eds.), Manual of Central American Diptera. Vol. 1. NRC Research Press, Ottawa, Ontario, pp. 671–694. Winnertz, J. (1852) Dipterologisches. Stettiner Entomologische Zeitung, 13, 49–58. Winnertz, J. (1864) Beitrag zu einer Monographie der Pilzmücken. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien, 13 [1863], 637–964. Zaitzev, A.I. (1994) Fungus gnats of the fauna of Russia and adjacent regions. Part 1. Nauka, Moscow, 288 pp. CATALOGUE OF DIADOCIDIIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 55 Zootaxa 4122 (1): 056–061 http://www.mapress.com/j/zt/ Copyright © 2016 Magnolia Press Article ISSN 1175-5326 (print edition) ZOOTAXA ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.4122.1.11 http://zoobank.org/urn:lsid:zoobank.org:pub:E1E57B31-5E72-4388-AD16-129821BD81EA FAMILY KEROPLATIDAE RAFAELA LOPES FALASCHI Universidade de São Paulo, Museu de Zoologia. Av. Nazaré, 481. Ipiranga. 04263-000. São Paulo SP Brazil. E-mail: rlfalaschi@gmail.com. Abstract Currently, 86 genera are ascribed to the Keroplatidae, with about 950 known species. The family includes three subfamilies: Arachnocampinae, Macrocerinae, and Keroplatinae. Both Macrocerinae and Keroplatinae are found in Colombia, with five described species distributed in four genera. The available information about Colombian keroplatids is compiled, with new records for five genera. Key words: Bibionomorpha, catalogue, Colombia, distribution Introduction Keroplatidae presently comprise 86 genera, and approximately 950 described species (Papavero 1978; Matile 1990; Evenhuis 1994, 2006; Falaschi 2012), present in all biogeographic regions. In the fossil record, they are known from the Upper Jurassic (about 150 million y.b.p.), while the Orfeliini appear in the fossil record only in the Cenozoic (about 65 million y.b.p.) (Matile 1990; Evenhuis 1994). The family is divided into three subfamilies: Arachnocampinae, Macrocerinae and Keroplatinae (Matile 1981). Papp and Ševčik (2005) proposed a new subfamily, Sciarokeroplatinae, based on a single species, Sciarokeroplatus pileatus Papp & Ševčík, from the Oriental region, but its validity was contested by Amorim et al. (2008). The family is distributed worldwide and commonly found in moist tropical forests, but also in other ecosystems, usually associated with fungi. Adults are often found in dark and humid places, like caves. They can be collected with sweeping in low vegetation, under hanging rocks, trunks and along banks of streams. They are also frequently found in Malaise traps, some species being collected in light traps with ultra-violet lamps and occasionally in yellow pan traps (Evenhuis 2006). Larvae are present in moist and dark places, like caves, trunks, slits in stones, and have different alimentary habits. Predaceous larvae are seen in Arachnocampinae, Macrocerinae and Keroplatinae (Mansbridge 1933) while mycophagy is present only in Keroplatinae (Evenhuis 2006). Keroplatids are medium-size flies (2.0–9.0 mm), with vein R4 present ending at C or R1 (except for a few species of Robsonomyiini and the keroplatini genus Chetoneura Colless), fusion of base of M1+2 with the base of RS, resulting in the r-m disappearance (although in the genera Pseudoplatyura Skuse and Platyura Meigen this condition is seemingly reverse) (Amorim & Rindal 2007). The Colombian fauna of keroplatids is composed by subfamilies Macrocerinae and Keroplatinae. In Macrocerinae, there are only two species, both belonging to the tribe Macrocerini. In Keroplatinae, both Keroplatini and Orfeliini are recorded with eight species (new record for five genera and species). The former is represented by two genera, and two species, whereas Orfeliini has five genera, with six species (Falaschi in prep.) (Table 1). Acronyms used for the depositories IAvH—Alexander von Humboldt Biological Resources Research Institute, Bogota, Colombia 56 Accepted by S. Nihei: 10 Apr. 2015; published: 14 Jun. 2016 MZSP—Museu de Zoologia da Universidade de São Paulo, São Paulo, Brazil NMW—Naturhistorisches Museum, Vienna, Austria USNM—National Museum of Natural History, Washington, D.C., USA List of abbreviations aut.—automatic cat.—catalogue col.—collector des.—designation distr.—geographic distribution i.e.—exemple m. —meters mon.— monotypy nom. nov.— nomen novum (new name) orig. des.—by original designation pp.— pages refs.—references rev.— revision sp., spp.—species ssp.—subspecies subs.—subsequent subg.—subgenus HT—Holotype NT—Neotype PT—Paratype LT—Lectotype ST—Syntype T—Type (unspecified) NA—Not applicable F (f#)—Female M (m#)—Male Catalogue of Keroplatidae of Colombia Family Keroplatidae Rondani, 1856 Subfamily Macrocerinae Rondani, 1856 Tribe Macrocerini Winnertz, 1863 Genus Macrocera Meigen Euphrosyne Meigen, 1800: 16. Type species, Macrocera lutea Meigen, 1804, by subs. des. (Coquillett, 1910: 542). [Suppressed by I.C.Z.N. (1963: 339).] Macrocera Meigen, 1803: 261. Type species, Macrocera lutea Meigen, 1804, by subs. des. (Guérin, 1826: 7). Geneja Lioy, 1863: 229 (unjustified new replacement name for Macrocera Meigen). Type species, Macrocera lutea Meigen, 1804, by subs. des. Promacrocera Speiser, 1913: 134. Type species, Promacrocera interrogationis Speiser, 1913, by orig. des. Promacrocera Armbruster, 1938: 118. Type species, Promacrocera archaica Armbruster, 1938, by mon. [Preoccupied by Speiser, 1913.] Macroura: Incorrect subs. spelling of Macrocera (Berendt, 1945: 57). CATALOGUE OF KEROPLATIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 57 matilei Papavero, 1978: 12 (new replacement name for nobilis Edwards, 1934: 354, preoccupied by Johnson, 1922.). Type locality: Colombia, Cordillera Central, Paso del Quindin, 3000 m. HT ♀ [NMW]. Distr.: Colombia. Ref.: Evenhuis, 2006: 42 (cat.). Genus Paramacrocera Tonnoir & Edwards Paramacrocera Tonnoir & Edwards, 1927: 779. Type species, Paramacrocera brevicornis Edwards, 1927, by orig. des. Subgenus Freemaniola Matile Freemaniola Matile, 1990: 174. Type species, Paramacrocera anomala Freeman, 1951, by orig. des. lanei Matile, 1990: 175. Type locality: “Colombia”. HT ♂ [MZSP]. Distr.: Colombia. Refs.: Evenhuis, 2006: 49 (cat.). Subfamily Keroplatinae Rondani, 1856 Tribe Orfeliini Malloch, 1917 Genus Isoneuromyia Brunetti Isoneuromyia Brunetti, 1912: 66. Type species, Isoneuromyia annandalei Brunetti (subs. des.). (Edwards, 1925: 166; see Evenhuis 2006: 131). undescribed sp. Distr.: Colombia (Valle del Cauca). Material examined. COLOMBIA: Valle del Cauca: PNN [Parque Nacional Natural], Farallones de Cali Cgto., La Meseta, 03°34'N 76°40'W, 2 males, 1960 m, 10–25.ii.2003, S. Sarria & M. Losso Leg., M.4543, Malaise (IAvH). Genus Lyprauta Edwards Lyprauta Edwards, 1931: 77 (as Platyura subgenus). Type species, Platyura defecta Edwards, 1931, by orig. des. zeteki (Lane), 1950: 64, (Platyura). Type locality: Panama. HT ♂ [USNM]. Distr.: Brazil (Piauí), Colombia, Panama, Trinidad. Refs.: Evenhuis, 2006: 79 (cat.). Genus Neoditomyia Lane & Sturm Neoditomyia Lane & Sturm, 1958: 199. Type species, Neoditomyia colombiana Lane & Sturm, 1958, by orig. des. Neoditomya: incorrect subsequent spelling of Neoditomyia (Papavero, 1978: 5). andina Lane & Sturm, 1958: 201. Type locality: Colombia, Sierra de la Macarena, Villavicencio, Río Zanza, 400–500 m. HT ♂ [MZSP]. Distr.: Colombia. Refs.: Evenhuis, 2006: 84 (cat.). colombiana Lane & Sturm, 1958: 204. Type locality: Colombia, Resina (w. Andes, between Garzón and Florencia) 1600–2000 m. HT ♂ [MZSP]. Distr.: Colombia. Refs.: Evenhuis, 2006: 84 (cat.). 58 · Zootaxa 4122 (1) © 2016 Magnolia Press FALASCHI Genus Neoplatyura Malloch Neoplatyura Malloch, 1928: 601. Type species, Platyura setiger Johannsen, 1910, by orig. des. undescribed sp. Distr.: Colombia (Nariño). Material examined. COLOMBIA: Nariño: RN [Reserva Natural], La Planada, Mun. Ricaurte Fca., Santa Rosa, Rio Miraflores, 01°15'N 78°15'W, 2 males, 1500 m, 11.viii.2004, D. Arias Leg., M.4986, (IAvH). Genus Proceroplatus Edwards Proceroplatus Edwards, 1925: 523. Type species, Platyura pictipennis Williston, 1896, by orig. des. Calliplatyura Malloch, 1928: 600. Type species, Platyura elegans Coquillett, 1895, by orig. des. Calloplatyura: incorrect original spelling of Calliplatyura (Malloch, 1928: 601). undescribed sp. Distr.: Colombia (Cauca). Material examined. COLOMBIA: Cauca: PNN [Parque Nacional Natural], Gorgona El Roble, 2°58'N 78°11'W, 2 males, 130 m, 28.ix–22.x.2001, H.Torres Leg., M.2461, Malaise, (IAvH). Tribe Keroplatini Rondani, 1856 Genus Keroplatus Bosc Keroplatus Bosc, 1792: 42. Type species, Keroplatus tipuloides Bosc, 1792, by mon.. Ceroplatus Fabricius, 1798: 550 (unjustified emendation of Keroplatus Bosc). Type species, Keroplatus tipuloides Bosc, 1792, aut. Ceroplates: incorrect subs. spelling of Ceroplatus. undescribed sp. Distr.: Colombia (Cauca). Material examined. COLOMBIA: Cauca: PNN [Parque Nacional Natural], Gorgona El Saman, 2°58'N 78°11'W, 2 males, 5 m, 06–23.iii.2002, R. Duque Leg., M.3088, Malaise, (IAvH). Genus Neoceroplatus Edwards Neoceroplatus Edwards, 1941: 304. (as Ceroplatus subgenus). Type species, Ceroplatus minimax Edwards, 1934, by orig. des. undescribed sp. Distr.: Colombia (Nariño). Material examined. COLOMBIA: Nariño: RN [Reserva Natural], La Planada, Centro Científico, 01°15'N 78°15'W, 2 males, 1700 m, 09–12.viii.2004, D. Arias Leg., M.4898, (IAvH). Acknowledgements The author is indebted to Michael Sharkey (National Science Foundation Grant DEB0205982), to whom I am most grateful. Additional information on the project can be found at the web address of the project (http:// www.sharkeylab.org/biodiversity/static.php?app=colombia&page=index). During the preparation of this paper, the author received financial support from FAPESP grant 2013/16524-9 and CNPq grant 159016/2012-6. CATALOGUE OF KEROPLATIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 59 TABLE 1. Keroplatidae species recorded from Colombia. Subfamily Tribe Genus Species Author Macrocerinae Macrocerini Macrocera matilei Papavero, 1978 Paramacrocera lanei Matile, 1990 Isoneuromyia sp. Lyprauta zeteki (Lane, 1950) Neoditomyia andina Lane & Sturm, 1958 colombiana Lane & Sturm, 1958 Keroplatinae Orfeliini Neoceroplatus sp. Proceroplatus sp. Keroplatus sp. Neoceroplatus sp. Keroplatini References Amorim, D.S. & Rindal, E. (2007) A phylogenetic study of the Mycetophiliformia, with creation of the subfamilies Heterotrichinae,Ohakuneinae, and Chiletrichinae for the Rangomaramidae (Diptera, Bibionomorpha). Zootaxa, 1535, 1–92. Amorim, D.S., Niu, C., Li, X., Lei, C. & Clarke, A.K. (2008) Chetoneura shennonggongensis, a new species of cave–dwelling Keroplatini from China (Diptera: Keroplatidae), with a discussion of the position of Chetoneura. Zootaxa, 1716, 59–68. Armbruster, L. 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Illinois State Laboratory of Natural History Bulletin, 12, 161–409. http://dx.doi.org/10.5962/bhl.title.9383 Malloch, J.R. (1928) Notes on Australian Diptera. No. xvii. Proceedings of the Linnean Society of New South Wales, 53, 598–617. Mansbridge, G.H. (1933) On the biology of some Ceroplatinae and Macrocerinae (Diptera, Mycetophilidae). With an appendix on the chemical nature of the web fluid in larvae of Ceroplatinae by H.W. Buston. Transactions of the Royal Entomological Society of London, 81, 75–92. http://dx.doi.org/10.1111/j.1365-2311.1933.tb00400.x Matile, L. (1981) Description d’un Keroplatidae du crétacé moyen et données morphologiques et taxinomiques sur les Mycetophiloidea (Diptera). Annales de la Société Entomologique de France, New Series, 17, 99–123. Matile, L. (1990) Recherches sur la systématique et l’évolution des Keroplatidae (Diptera, Mycetophiloidea). Mémoires du Muséum national d’Histoire naturelle Paris, Series A, 148, 1–682. Meigen, J.G. (1800) Nouvelle classification des mooches à deux ailes (Diptera L.) d’après un plan tout nouveau. J.J. Fuchs, Paris, 40 pp. Meigen, J.G. (1803) Versuch einer neuen Gattungs Eintheilung der europäischen zweiflügeligen Insekten. Magazin für Insektenkunde, 2, 264. Meigen, J.G. (1804) Klassifikazion und Beschreibung der europäischen zweifügeligen Insekten (Diptera Linn.). Braunschweig, 1, 104. Papavero, N. (1978) 19C. Family Keroplatidae (Ceroplatidae, incl. Macroceridae). In: Papavero, N. (Ed.), A Catalogue of the Diptera of the Americas south of the United States. Departamento de Zoologia da Secretaria da Agricultura do Estado de São Paulo, São Paulo, pp. 1–22. Papp, L. & Ševčík, J. (2005) Sciarokeroplatinae, a new subfamily of Keroplatidae (Diptera). Acta Zoologica Academiae Scientiarum Hungaricae, 51 (2), 113–123. Rondani, C. (1856) Dipterologiae Italicae Prodromus. Vol. 1. Genera Italica ordinis dipterorum ordinatim disposita et distincta et in familias et stirpes aggregata. Parma, 228 pp. Speiser, P. (1913) Beiträge zur Dipterenfauna von Kamerun. Deutsche Entomologische Zeitschrift, 1913 (2), 131–146. Tonnoir, A.L. & Edwards, F.W. (1927) New Zealand fungus gnats (Diptera, Mycetophilidae). Transactions of the New Zealand Institute, 57, 747–878. Williston, S.W. (1896) On the Diptera of St. Vincent (West Indies). Transactions of the Entomological Society of London, 1896, 253–446. Winnertz, J. (1863) Beitrag zu einer Monographie der Pilzmücken (Mycetophilidae). Verhandlungen der ZoologischBotanischen Gesellschaft in Wien, 13, 637–964. [1863] CATALOGUE OF KEROPLATIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 61 Zootaxa 4122 (1): 062–064 http://www.mapress.com/j/zt/ Copyright © 2016 Magnolia Press Article ISSN 1175-5326 (print edition) ZOOTAXA ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.4122.1.12 http://zoobank.org/urn:lsid:zoobank.org:pub:569BC41C-D32B-40EF-86FF-405602B8CA97 FAMILY LYGISTORRHINIDAE SARAH SIQUEIRA OLIVEIRA1 & DALTON DE SOUZA AMORIM2* 1 Departamento de Ecologia, Instituto de Ciências Biológicas, Universidade Federal de Goiás, Campus II. Prédio ICB1, 74001-970. Goiânia GO Brazil. E-mail: oliveira.sarahcv@gmail.com 2 Departamento de Biologia, Faculdade de Filosofia, Ciências e Letras de Ribeirão Preto, Universidade de São Paulo. Av. Bandeirantes, 3900. 14040-901. Ribeirão Preto SP Brazil. E-mail: dsamorim@usp.br *Corresponding author: dsamorim@usp.br Abstract The Lygistorrhinidae are a family belonging to the suborder Bibionomorpha, with no previous record from Colombia. This paper refers for the first time to the occurrence of the family in the country, an undetermined species of the genus Lygistorrhina (Probolaeus) Williston. Key words: catalogue, Colombia, Distribution, Lygistorrhinidae. Introduction Lygistorrhinidae are a family widely distributed in tropical and subtropical regions of the world, with a total of 15 genera, of which eight are extant and seven known only from fossils (Grimaldi & Blagoderov 2001, Hippa et al. 2005, Pape et al. 2011). Detailed information on the biology of the group and immature stages habitat is still wanting. The genus Lygistorrhina Skuse encompasses 21 species worldwide, divided in two subgenera—Lygistorrhina s.str. and Probolaeus Williston. Phylogenies for the family have been proposed by Grimaldi & Blagoderov (2001), Blagoderov & Grimaldi (2004), Hippa et al. (2005), and Blagoderov et al. (2009). Despite the robust hypothesis of monophyly of Lygistorrhina s.l., there is still no indication that the set of species gathered under the subgenus L. (Lygistorrhina) composes a monophyletic arrangement. Hence, neither generic nor subgeneric status to Probolaeus correspond to true solutions for the problem of the group. A complete revision of Lygistorrhina s.l. must be undertaken before the genus could be split into two or more groups, independently of the rank (Oliveira & Amorim 2012). A catalogue of the Neotropical Lygistorrhinidae was published by Papavero (1977, 2002), and an updated version is now available Oliveira & Amorim (2012). Brief descriptions, photos of the types of some species, and distributional data are also available in Fungus Gnats Online (2013). Unfortunately, in the material available for our examination, only females were present, so this allowed establishing here the first record of the family for Colombia (Table 1), but it is not much helpful for species level determination. Acronyms used for the depositories IAvH—Alexander von Humboldt Biological Resources Research Institute, Bogota, Colombia. 62 Accepted by S. Nihei: 10 Jun. 2015; published: 14 Jun. 2016 List of abbreviations det.—determine distr.—geographic distribution m.—meters mon.—by monotypy refs.—references Catalogue of Lygistorrhinidae of Colombia Family Lygistorrhinidae Genus Lygistorrhina Skuse Lygistorrhina Skuse, 1890: 598. Type species, L. insignis Skuse, 1890: 600 (mon.) Subgenus Lygistorrhina (Probolaeus) Williston Probolaeus Williston, 1896: 261. Type species, P. singularis Williston (mon.) (as a genus). Refs.: Thompson (1975), Grimaldi & Blagoderov (2001), Hippa et al., (2005). undetermined sp. Distr.: Colombia (Vaupés). Material examined. COLOMBIA: Vaupés: Estación Biológica Mosiro-Itajura (Caparú) Igapo, 01°04’S 69°31’W, 60 m., 2 females, 1–9.ii.2003, J. Pinzó, Malaise, Oliveira, S.S. det. 2013 (IAvH); 1 female, 4–11.iii.2003. J. Pinzón, Malaise, Leg. M.3623. Oliveira, S.S. det. 2013 (IAvH); 1 female, 18–27.iii.2003. L. Benavides, Malaise Leg. M.3626. Oliveira, S.S. det. 2013 (IAvH). Acknowledgements We are thankful to Dr. Michael Sharkey that kindly made available the material for this study. Additional information can be found in the project web address http://www.sharkeylab.org/biodiversity/static.php?app=colombia&page=index. We are grateful to Marta Wolff, Silvio Shigueo Nihei, Sandra Patricia Pérez Pareja, and Claudio José Barros de Carvalho, the editors of the “Catalogue of Diptera of Colombia”. The first author has a fellowship from the Fundação de Amparo à Pesquisa do Estado de São Paulo (FAPESP, grant 2012/51577-3) and the second author has a fellowship from the Brazilian National Science Research Agency (CNPq, grant 314371/2009-5). TABLE 1. Lygistorrhinidae species recorded from Colombia. Genus Subgenus Species Author Lygistorrhina Probolaeus sp. Williston, 1896 References Blagoderov, V. & Grimaldi, D. (2004) Fossil Sciaroidea (Diptera) in Cretaceous ambers, exclusive of Cecidomyiidae, Sciaridae, and Keroplatidae. American Museum Novitats, 3433, 1–76. http://dx.doi.org/10.1206/0003-0082(2004)433%3C0001:FSDICA%3E2.0.CO;2 Blagoderov, V., Hippa, H. & Ševčík, J. (2009) Asiorrhina, a new Oriental genus of Lygistorrhinidae (Diptera: Sciaroidea) and its phylogenetic position. Zootaxa, 2295, 31–45. Fungus Gnats Online (2013) Available from: http://sciaroidea.info/taxonomy/41555 (accessed 22 August 2013) Grimaldi, D. & Blagoderov, V. (2001) A new genus of Lygistorrhinidae from Vietnam (Diptera: Sciaroidea), and CATALOGUE OF LYGISTORRHINIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 63 phylogenetic relationships in the family. Studia Dipterologica, 8, 43–57. Hippa, H., Mattsson, I. & Vilkamaa, P. (2005) New taxa of the Lygistorrhinidae (Diptera: Sciaroidea) and their implications for a phylogenetic analysis of the family. Zootaxa, 960, 1–34. Oliveira, S.S. & Amorim, D.S. (2012) Catalogue of Neotropical Diptera. Lygistorrhinidae. Neotropical Diptera, 22, 1–4. Papavero, N. (1977) 19D. Family Lygistorrhinidae. In: Papavero, N. (Ed.), A catalogue of the Diptera of the Americas South of the United States. Departamento de Zoologia da Secretaria da Agricultura do Estado de São Paulo, São Paulo, pp. 1. Papavero, N. (2002) Insecta, Diptera, Lygistorrhinidae. Fauna da Amazônia Brasileira, 10, 1. Pape, T., Blagoderov, V. & Mostovski, M.B. (2011) Order Diptera Linnaeus, 1758. In: Zhang, Z.-Q. (Ed.), Animal biodiversity: An outline of higher-level classification and survey of taxonomic richness. Zootaxa, 3148, pp. 222–229. Skuse, F.A.A. (1890) Diptera of Australia. Nematocera, Supplemment II. Proceedings of the Linnean Society New South Wales, 2 (5), 595–640. Thompson, F.C. (1975) Notes on the genus Lygistorrhina Skuse with the description of the first nearctic species (Diptera: Mycetophiloidea). Proceedings of the Entomological Society of Washington, 77 (4), 434–445. Williston, S.W. (1896) On the Diptera of St. Vincent (West Indies). Transactions of the Entomological Society of London, 1896, 253–449. http://dx.doi.org/10.1111/j.1365-2311.1896.tb00965.x 64 · Zootaxa 4122 (1) © 2016 Magnolia Press OLIVEIRA & AMORIM Zootaxa 4122 (1): 065–072 http://www.mapress.com/j/zt/ Copyright © 2016 Magnolia Press Article ISSN 1175-5326 (print edition) ZOOTAXA ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.4122.1.13 http://zoobank.org/urn:lsid:zoobank.org:pub:5845FE68-E84E-4C55-9854-26C82492A2AF FAMILY MYCETOPHILIDAE SARAH SIQUEIRA OLIVEIRA1 & DALTON DE SOUZA AMORIM2 1 Departamento de Ecologia, Instituto de Ciências Biológicas, Universidade Federal de Goiás, Campus II. Prédio ICB1, 74001-970. Goiânia GO Brazil. E-mail: oliveira.sarahcv@gmail.com 2 Departamento de Biologia, Faculdade de Filosofia, Ciências e Letras de Ribeirão Preto, Universidade de São Paulo. Av. Bandeirantes, 3900. 14040-901. Ribeirão Preto SP Brazil. E-mail: dsamorim@usp.br *Corresponding author: oliveira.sarahcv@gmail.com Abstract The Mycetophilidae include small fungus-gnats which life cycle is associated with fungi, especially of the larvae. The known diversity of the family in the Neotropical region is 1,145 species, but only some very few papers have been published on the Colombian species of Mycetophilidae, with records for the genera Docosia Winnertz, Paraleia Tonnoir, and Dziedzickia Johannsen. This catalogue gathers the information available on mycetophilids from Colombia, including genera and some species that for the first time are mentioned to occur in the country—as Leiella unicincta Edwards and Leiella zonalis Edwards. Key words: catalogue, Colombia, distribution, Mycetophilidae Introduction Mycetophilids are small fungus-gnats (Diptera, Mycetophilidae), usually with a combination of brown, black and yellow colors in adults (Hutson et al. 1980). Adults are often found in dark, damp woods, tunnels and root system of trees, and are particularly common in cavities under tree roots and overhanging stream banks (Hutson et al. 1980, Ševčík 2010). Larvae generally feed on mycelium and sporophores penetrating hyphae. But there are species with diverging biology, being associated to decaying plant organic material. Details of immatures and the biology of most species in the family remain unknown (Hutson et al. 1980, Ševčík 2010). In a study of forest stratification in the Appalachians, a non-random distribution of different species of Mycetophilidae was found in three regions of Tsuga canadensis Cup (L.), associated with incidence of natural light (Coots et al. 2012). Only adults were collected, indicating possible migratory behavior to areas with more suitable temperature, moisture, food sources and protection (Coots et al. 2012). Mycetophilids are common in forests and continuity between forest fragments has a strong impact on the faunal composition of the family. The life cycle, particularly immatures, is connected to fungi with fruit bodies, so precipitation, wood composition and wood abundance correspond to important factors in population density (Økland 1994, 1996, Økland et al. 2005). The Mycetophilidae belong to the suborder Bibionomorpha (in the sense of Amorim & Yeates 2006), together composing with seven other recent families the Mycetophiliformia (Hennig 1973, Amorim & Rindal 2007). There are conflicts between authors about the phylogeny of the group (Søli 1997, Tozoni 1998, Rindal et al. 2009a), but the monophyly of Bibionomorpha and Mycetophiliformia is well corroborated (Wood & Borkent 1989, Amorim & Rindal 2007). Different hypotheses of relationship of the family within the higher Bibionomorpha can be found in Amorim & Rindal (2007: 55, 69), but and there is little doubt about Lygistorrhinidae being the sister group of Mycetophilidae. The Mycetophilidae are widely accepted as a monophyletic group (Søli 1997, Tozoni 1998, Rindal et al. 2009a). The family includes clades recently treated as subfamilies: Sciophilinae, Gnoristinae, Mycomyiinae, Leiinae, Manotinae, Allactoneurinae, and Mycetophilinae (Tozoni 1998, Chandler 2009, Rindal et al. 2009a, Amorim & Oliveira 2008, Oliveira & Amorim 2010a, b, 2012). Some phylogenetic studies were performed for the Accepted by S. Nihei: 10 Jun. 2015; published: 14 Jun. 2016 65 Mycetophilidae subfamilies (e.g., Hippa et al. 2004, Rindal & Søli 2006, Rindal et al. 2007, 2009b, Borkent & Wheeler 2013, Oliveira 2013, Ševčík et al. 2013), even though the taxonomic sampling of some of these studies may consist a limitation to the reliability of the conclusions. A wide phylogeny of the family, however, including all or almost all of its genera is still waiting. A total of 233 genera and about 4,500 species of Mycetophilidae were described so far from all biogeographic regions (Pape et al. 2011). Records of species belonging to the crown-group of the family are known from fossil at least from the Cretaceous (Evenhuis 1994, Amorim & Silva 2002). For the Neotropical region, 54 genera and 1,145 species of Mycetophilidae have been described so far (Papavero 1978, Oliveira & Amorim 2014). The type series of most species is allocated at the Natural History Museum (NHM), London (mostly F.W Edwards and John Lane’s species) (Amorim & Oliveira 2013), at the Museu de Zoologia da Universidade de São Paulo, São Paulo (MZUSP) (mostly John Lane’s species), and at the Museum national d’Histoire Naturelle, Paris (mostly J.P. Duret’s species). Species of Mycetophilidae have been described based on material from Colombian type localities only very recently (Oliveira & Amorim 2011, 2012). Their holotypes and paratypes are located at the Alexander von Humboldt Biological Resources Research Institute (IAvH), Bogota, Colombia, and at the MZUSP. Colombia is one of the countries in the Neotropical region with particularly special diversity not only in terms of number of species, but also in terms of the biogeographical components to which the species belong. Higher altitude species of Mycetophilidae in the Andes of Colombia in many cases belong to genera with amphinotic distribution (i.e., present in Chile and Australia and/or New Zealand—Amorim et al. 2009), with tropical connections (i.e., present in Amazonia, tropical forests in Central America and in the Atlantic Forest), as well as nearctic elements, sometimes also found in higher areas in Central America (e.g., Costa Rica) (Oliveira et al. 2007). The distribution given for the species in this catalogue in many cases is that one stated in published papers. However, experience shows that, although some cases actually may correspond to widely distributed species, many species have been misidentified and wide distributions may be mistaken for some species. This can be corrected only with detailed studies within each genus. Acronyms used for the depositories IAvH—Alexander von Humboldt Biological Resources Research Institute, Bogota, Colombia MTD—Museum für Tierkunde, Dresden, Germany MZUSP—Museu de Zoologia da Universidade de São Paulo, São Paulo, Brazil NHM—Natural History Museum, London, United Kingdom List of abbreviations aut.—automatic det.—determined distr.—geographic distribution fig.—figure e.g.—example HT—holotype leg.—collector m.—meters mon.—monotypy nom. nov.—nomen novum (new name) orig. des.—by original designation pp.—pages preocc.—preoccupied refs.—references 66 · Zootaxa 4122 (1) © 2016 Magnolia Press OLIVEIRA & AMORIM sp., spp.—species syn.—synonym F (f#)—Female M (m#)—Male Catalogue of Mycetophilidae of Colombia Family Mycetophilidae Newman, 1834 Subfamily Gnoristinae Edwards, 1925 Genus Docosia Winnertz Docosia Winnertz, 1863: 802. Type species, Mycetophila sciarina Meigen (Johannsen, 1909: 92). adusta Oliveira & Amorim, 2011: 689, 690. (fig. 1, habitus), 691 (fig. 2, thorax), 692 (fig. 3, asa), 693 (figs. 4-6, M terminalia), 694 (fig. 7, F terminalia). Type locality: Colombia Cundinamarca, Parque Nacional Natural Chingaza Alto de La Bandera. HT M (IAvH). Distr.: Colombia (Cundinamarca). Genus Dziedzickia Johannsen Hertwigia Dziedzicki, 1885: 166 (preocc. Schmidt 1880). Type species, H. marginata Dziedzicki (mon.). Dziedzickia Johannsen, 1909: 44 (nom. nov. for Hertwigia Dziedzicki). Type species, Hertwigia marginata Dziedzicki (aut.). Syntemna, authors, not Winnertz. intermedia Lane, 1954a: 102, fig. 8 (M terminalia). Type locality: Brazil, São Paulo, Campos do Jordão. HT M (MZUSP). Distr.: Costa Rica (Alajuela, San Mateo), Colombia (Vaupés), Brasil (Amapá, Sergipe, Minas Gerais, São Paulo). Refs.: Oliveira, 2009 (redescription, identification key, and distr.). jurupari Lane, 1961: 249. Type locality: Brazil, Amapá, Rio Felício. HT M (MZUSP). Distr.: Colombia (Amazonas, Nariño, Putumayo, Vaupés), Brasil (Amapá, Acre). Refs.: Oliveira, 2009 (redescription, identification key, and distr.). Subfamily Leiinae Hendel, 1936 Genus Leiella Enderlein Leiella Enderlein, 1910a: 200. Type species, ochreocalcar Enderlein (orig. des.). Refs.: Lane, 1954b (key to spp.). unicincta Edwards, 1933: 319. Type locality: Peru, mouth of Río Pachitea, 150 m. HT M (MTD). Distr.: Peru (Pasco/Huanaco), Colombia (Meta, Vaupés). Refs.: Lane, 1952: 137, fig. 5 (male terminalia), 1954b: 450, fig. 1 (M terminalia). Material examined. COLOMBIA:. Meta: PNN [Parque Nacional Natural] Sierra de La Macarena, Cabaña Cerrillo, 03°21’N 73°56’W, 460 m, 1 female, 10.xi–21.xii.2002. A. Herrera & W. Villalba, Malaise, Leg. M.2982. Oliveira, S.S. det. ix.2013 (IAvH); Vaupés: Estación Ecológica Mosiro-Itajura (Caparú), Centro Ambiental, 01°04’S 69°31’W, 60m, 1 female, 24.xi-01.xii.2003. J. Pinzón, Malaise, Leg. M.4439. Oliveira, S.S. det. ix.2013 (IAvH). zonalis Edwards, 1931: 259. Type locality: British Guyana, Berbice. HT F (NHM). Distr.: Argentina (Tucuman, Salta), British Guyana (Berbice), Chile (Santa Fé, Chaco), Colombia (Magdalena, Meta, Bolivar, Vaupés, Norte de Santander, Cauca, Chocó, Putumayo). Refs.: Lane, 1954c (distr.), 1958 (distr.). CATALOGUE OF MYCETOPHILIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 67 Material examined: COLOMBIA: Magdalena: PNN [Parque Nacional Natural] Tayrona, Cañaveral, 11°20’N 74°02’W, 30 m, 6 females, 03–22.xi.2000. R. Henriquez Leg. M.943. Oliveira, S.S. det. ix.2013, (IavH); Meta: PNN [Parque Nacional Natural] Sierra de La Macarena, Cabaña Cerrillo, 03°21’N 73°56’W, 460 m, 1 female, 21.xii.2002–04.i.2003, Malaise, A. Herrera & W. Villalba Leg. M.2983. Oliveira, S.S. det. ix.2013 (IAvH); Bolivar: SFF [Santuario de Fauna y Flora] Los Colorados, Vila Roca, 09°54’N 75°07’W, 180m, 1 female, 23.v–07.vi.2001, Malaise E. Deulufeut Leg. M.1723. Oliveira, S.S. det. ix.2013, (IAvH); Vaupés: Estación Ecológica Mosiro-Itajura (Caparú), Igapo, 01°04’S 69°31’W, 60m, 1 male, 1–9.ii.2003. J. Pinzón Leg. M.3637, Malaise, Oliveira, S.S. det. ix.2013 (IAvH); same data, except 1 female, 22.x-06.xi.2002, L. Benevides Leg. M.3396 (IavH); same data, except 1 male, 18–27.iii.2003, M.3635 (IAvH); same data, except Centro Ambiental, 1 female, 10–17.iv.2003, M.3614 (IAvH); same data, except Centro Ambiental, 1 female, 03–10.iv.2003, M.3638 (IAvH); Norte de Santander: ÁNU [Área Natural Única] Los Estoraques, Qda. La Media, 08°13’N 73°15’W, 1480m, 1 female, 15–29.x.2003, J. Vargas Leg. M.4089, Malaise, Oliveira, S.S. det. ix.2013 (IAvH); same data, except 1 female, 08°13’N 73°14’W, 01–16.xii.2003, M.4606 (IAvH); same data, except Ciudad Perdida, 2females, 08°13’N 73°14’W, 22.xi–05.xii.2003, E. Bayona Leg. M.4607 (IAvH); same data, except Platanillo, 3 females, 08°14’N 73°14’W, 1516 m, 28.xi–13.xii.2003, E. Bayona Leg. M.4600 (IAvH); Cauca: PNN [Parque Nacional Natural] Gorgona El Saman, 02°58’N 78°11’W, 5 m, 1 female, 06-23.iii.2002. R. Duque Leg. M.3088, Malaise, Oliveira, S.S. det. ix.2013 (IAvH); Chocó: PNN [Parque Nacional Natural] Los Katios, Centro Adm. Sautatá Dentro del bosque, 07°51’N 77°08’W, 30m, 1 female, 13–29.vi.2003. P. López Leg. M.3764, Malaise, Oliveira, S.S. det. ix.2013 (IAvH); Putumayo: PNN [Parque Nacional Natural] La Paya, Resguardo Cecilio Cocha, 00°11’N 74°55’W, 200m, 1 male, 1 female, 20–24.i.2003. C. Sarmiento Leg. M.3421, Malaise, Oliveira, S.S. det. ix.2013 (IAvH). Genus Paraleia Tonnoir Paraleia Tonnoir, 1929: 611. Type species, P. fulvescens Tonnoir (orig. des.). Selkirkius Enderlein, 1940: 670. Type species, S. nephrodops Enderlein (orig. des.). Refs.: Freeman, 1954: 26 (syn.). Refs.: Oliveira & Amorim, 2012 (new species and identification key). bolivari Oliveira & Amorim, 2012: 4, 6 (fig. 2, thorax), 7 (figs. 3–4, wings), 8 (fig. 13, male terminalia), 9 (fig. 14, female terminalia), 10 (fig. 15, female terminalia). Type locality: Colombia, Cundinamarca, Parque Nacional Natural Chingaza. HT M (IAvH). Distr.: Colombia (Cundinamarca). denticulata Oliveira & Amorim, 2012: 12, 7 (figs. 7–8, wings), 14 (fig. 18, male terminalia), 15 (fig. 19, female terminalia). Type locality: Colombia, Cundinamarca, Parque Nacional Natural Sumapaz Bocatoma, Cerro El Zapato. HT M (IAvH). Distr.: Colombia (Cundinamarca). fumosa Oliveira & Amorim, 2012: 6, 5 (fig. 1, habitus), 7 (figs. 5-6, wings), 11 (fig. 16, male terminalia), 13 (fig. 17, female terminalia). Type locality: Ecuador, Pichincha, E. Papallacta (Quito/Baeza). HT M (MZUSP). Distr.: Colombia (Huilla), Ecuador (Pichincha). nidorosa Oliveira & Amorim, 2012: 17, 7 (fig. 10, wing), 18 (fig. 21, female terminalia). Type locality: Colombia, Huilla, Parque Nacional Natural Cueva de lós Guácharos. HT F (IAvH). Distr.: Colombia (Huilla). sharkeyi Oliveira & Amorim, 2012: 19, 7 (fig. 11, wing), 20 (fig. 22, female terminalia). Type locality: Colombia, Cundinamarca, Parque Nacional Natural Chingaza, Alto de La Bandera. HT F (IAvH). Distr.: Colombia (Cundinamarca). tonnoiri Oliveira & Amorim, 2012: 15, 7 (fig. 9, wing), 16 (fig. 20, male terminalia). Type locality: Colombia, Magdalena, Parque Nacional Natural Santa Marta, San Lorenzo. HT M (IAvH). Distr.: Colombia (Magdalena). undetermined sp. Oliveira & Amorim, 2012: 19, 7 (fig. 12, wing), 21 (fig. 23, female terminalia). Type locality: Colombia, Cundinamarca, Parque Nacional Natural Chingaza, Bosque Palacio. F (IAvH, MZUSP). Distr.: Colombia (Cundinamarca, Boyacá, Huilla). 68 · Zootaxa 4122 (1) © 2016 Magnolia Press OLIVEIRA & AMORIM First records of Mycetophilidae genera in Colombia Dr. Michael Sharkey (National Science Foundation Grant DEB-0205982) project of collections in Colombia with Malaise traps resulted in an amazing amount of material with invaluable importance to our understanding of the Diptera diversity. Additional information on the project can be found at the web address of the project (http:// www.sharkeylab.org/biodiversity/static.php?app=colombia&page=index, last access in September, 05th, 2013). Many genera of Mycetophilidae were recorded for the first time in Colombia after this project. Some of the new generic occurrences were already mentioned (Oliveira et al. 2007, Oliveira 2009), but we should add here records for other genera whose species still wait to be published/described. Based on the Dr. Michael Sharkey material, housed at the IAvH collection, the following genera are first recorded for Colombia: Acnemia Winnertz (1863), Allocotocera Mik (1886), Cluzobra Edwards (1940), Duretophragma Borkent (2013), Eudicrana Loew (1869), Megalopelma Enderlein (1910a), Monoclona Mik (1886), Neoaphelomera Miller (1945), Neuratelia Rondani (1856), and Sciophila Meigen (1818)—Subfamily Sciophilinae Rondani; Austrosynapha Tonnoir (1929), Synapha Meigen (1818), Tetragoneura Winnertz (1846)—Subfamily Gnoristinae Edwards; Mycomya Rondani (1856) and Neoempheria Osten-Sacken (1878)—Subfamily Mycomyinae Edwards; Aphrastomyia Coher & Lane (1949), Leia Meigen (1818), Megophthalmidia Dziedzicki (1889), and Procycloneura Edwards (1932) - Subfamily Leiinae Edwards; Manota Williston (1896)—Subfamily Manotinae Edwards; Cordyla Meigen (1803)—which three species are being described by Kurina & Oliveira (in press) —, Epicypta Winnertz (1863), Exechiopsis Tuomikoski (1966), Mycetophila Meigen (1803), Neallodia Edwards (1932), Platurocypta Enderlein (1910b), Phronia Winnertz (1863), Rymosia Winnertz (1863), Trichonta Winnertz (1863), and Zygomyia Winnertz (1863)—Subfamily Mycetophilinae Newman. The updated version of the Neotropical catalogue of Mycetophidae (Oliveira & Amorim, 2014: 25) mistakenly referes to the species Leia amabilis (Williston) as distributed in Colombia. This species was described for the region of the Vera Cruz, near Medellin, in Mexico, and not in Colombia. Hence, Leia is first recorded from Colombia after the inventory project of the country fauna, as mentioned above. Acknowledgements The authors are deeply thankful to Dr. Michael Sharkey that kindly made available the material for this study. SSO benefited from FAPESP (grant 2012/51577-3) and DSA has a fellowship from CNPq (314371/2009-5). Special thanks to the editors of the “Catalogue of Diptera of Colombia”: Marta Wolff, Silvio Shigueo Nihei, Sandra Patricia Pérez Pareja, and Claudio José Barros de Carvalho. TABLE 1. Mycetophilidae species recorded from Colombia. Subfamily Genus Species Author Gnoristinae Docosia adusta Oliveira & Amorim, 2011 Dziedzickia intermedia Lane ,1954a jurupari Lane, 1961 unicincta Edwards, 1933 zonalis Edwards, 1931 Leiinae Leiella Paraleia CATALOGUE OF MYCETOPHILIDAE OF COLOMBIA bolivari Oliveira & Amorim, 2012 denticulata Oliveira & Amorim, 2012 fumosa Oliveira & Amorim, 2012 nidorosa Oliveira & Amorim, 2012 sharkeyi Oliveira & Amorim, 2012 tonnoiri Oliveira & Amorim, 2012 undetermined sp. Oliveira & Amorim, 2012 Zootaxa 4122 (1) © 2016 Magnolia Press · 69 References Amorim, D.S. & Oliveira, S.S. (2013) Types of Neotropical Mycetophilidae (Diptera) at the Natural History Museum collection, London. Zootaxa, 3726 (1), 1–119. http://dx.doi.org/10.11646/zootaxa.3726.1.1 Amorim, D.S. & Oliveira, S.S. (2008) Eleven new species of the genus Cluzobra Edwards (Diptera, Mycetophilidae, Sciophilinae) from the Atlantic Forest of Brazil. Zootaxa, 1920, 1–28. Amorim, D.S. & Rindal, E. 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Research Branch Monograph 29, Agriculture Canada, Ottawa, Ontario, pp. 1333–1370. 72 · Zootaxa 4122 (1) © 2016 Magnolia Press OLIVEIRA & AMORIM Zootaxa 4122 (1): 073–097 http://www.mapress.com/j/zt/ Copyright © 2016 Magnolia Press Article ISSN 1175-5326 (print edition) ZOOTAXA ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.4122.1.14 http://zoobank.org/urn:lsid:zoobank.org:pub:C2E36D43-DAEC-4FEB-9321-2E9DE5A82C44 FAMILIES TIPULIDAE AND LIMONIIDAE GUILHERME C. RIBEIRO* & DAUBIAN SANTOS *Universidade Federal do ABC, Centro de Ciências Naturais e Humanas, Av. dos Estados, 5001. Bairro Bangu, 09210-580, Santo André, SP, Brazil. *Corresponding author: guilherme.ribeiro@ufabc.edu.br Abstract A catalogue of the Colombian crane flies (Tipulomorpha, families Tipulidae and Limoniidae) is provided, based on updated information from the Catalogue of the Crane flies of World, with additional data on the geographical distribution of the species in Colombia taken from the primary literature. A total of 131 valid species are recorded for Colombia, 38 in the family Tipulidae and 93 in the family Limoniidae. Key words: catalogue, Colombia, Tipulidae, Limoniidae Introduction The crane flies–Tipulomorpha (Starý 1992, Amorim & Yeates 2006)—include the families Limoniidae, Tipulidae, Pediciidae and Cylindrotomidae. Over 15,000 extant species in ca. 500 genera are included in this group, which has a worldwide distribution (Oosterbroek 2015). Only the families Limoniidae and Tipulidae occur in Colombia. Adults of crane flies are found in a variety of different environments, such as forests, grasslands, cultivated fields, urban yards, and deserts. The immature stages of most species live in aquatic or semi aquatic habitats, but many species with terrestrial larvae are also known (Alexander & Byers 1981, Gelhaus 2005, De Jong et al. 2008). The detailed review of Pritchard (1983) is still the most complete account on the biology of crane flies. Morphologically, the adults of Tipulomorpha can be recognized by their slender bodies with narrow wings and long legs. However, a lot of variation may occur and species with relatively more stout bodies are common, especially in the Limoniidae. The ocelli is absent in all members of the group, which also have a typical “V”shaped transverse suture in the mesonotum. Crane flies can be small or large flies with wing lengths spanning from less than 3 mm to over 30 mm. Members of the Tipulidae typically have larger sizes as compared with the Limoniidae. In Tipulidae, the terminal segment of the maxillary palps is slender and longer than the penultimate segment, and the antennae are normally 13-segmented. In the Limoniidae, the terminal segment of the maxillary palps is as short as the others, and the antennae usually have 14 to 16 segments. Crane fly larvae are typically elongate with distinct but retractable head capsules. A recent detailed diagnosis for the group is provided by Gelhaus (2009). The classification of crane flies is conflicting regarding the taxonomic ranks attributed to its higher taxa. According to the system of classification adopted here, which follows Starý (1992) and Amorim & Yeates (2006), the Tipulomorpha includes four families: Tipulidae, Limoniidae, Pediciidae and Cylindrotomidae. Another classification groups the crane flies into a single family, the Tipulidae sensu lato, with the taxa treated here as families ranked as subfamilies (Tipulinae, Limoniinae and Cylindrotominae) and tribe (Pediciini, subordinated to the subfamily Limoniinae) (e.g., Alexander & Alexander 1970, Gelhaus 2009). While the monophyly of the entire Tipulomorpha seems well supported by phylogenetic studies with both morphological and molecular data (Oosterbroek & Theowald 1991, Starý 1992, Ribeiro 2008, Petersen et al. 2010), neither of the competing classifications can be considered as natural. The Pediciidae is probably the sister-group to the rest of the Tipulomorpha. Tipulidae and Cylindrotomidae are likely to be monophyletic sister groups. However, the large and Accepted by S. Nihei: 17 Apr. 2015; published: 14 Jun. 2016 73 heterogeneous Limoniidae is almost certainly paraphylectic with respect to the Tipulidae + Cylindrotomidae clade (Ribeiro 2008, Petersen et al. 2010). For the student of the crane flies, a very especial and valuable source of information is available on the internet: the Catalogue of the Crane flies of the World (CCW), by Pjotr Oosterbroek (Oosterbroek 2015, available at http://ccw.naturalis.nl/index.php). This database covers all 17,638 genus-group and species-group taxa of the four crane fly families, and has been regularly updated. CCW brings not only the taxonomic information included in traditional catalogues, but also illustrations of many taxa (by the time this chapter is being written, 5,138 images of 1,664 species were included), and a vast reprint library on crane fly taxonomy, biology, systematics and biogeography (PDF files) which includes all the papers of Charles Paul Alexander, among other authors. The catalogue presented here for Colombia follows the same format and is primarily based on the information provided by CCW, with additions regarding type-localities and precise distributions of the species in Colombia as provided by the primary literature, mainly the original descriptions of the taxa. For some taxa, however, detailed information on distribution is simply not available. Also, many taxa are known exclusively from their type-localities. Another important source of information on type-localities and distributions is the catalogue of the Neotropical Region (Alexander & Alexander 1970). The reader must be aware that the classification adopted here follows that used in CCW, and differs from the system adopted by Alexander & Alexander (1970). Also, the status of some taxa, synonyms, and spellings of some taxon names may differ in these two sources, the most updated one being what is found in CCW. The crane fly species fauna of Colombia belongs to genera and subgenera typically found in the tropical parts of the Neotropical Region. All the genera and subgenera known to occur in Colombia are included in the detailed treatment of Gelhaus (2009) of the Central American fauna, which includes a richly illustrated identification key to the genera and subgenera (adults) and a synopsis of the fauna with useful information for each taxon. A total of 131 valid species of crane flies are recorded for Colombia, 38 in the family Tipulidae and 93 in the Limoniidae (Table 1). Most species (81 species, or 62% of the fauna) are known exclusively from Colombia, and many are known only from their type-localities. However, considering our fragmentary knowledge about the Neotropical fauna in general, it is likely that the total number of Colombian species is much larger and that some of the “endemic” taxa may also occur elsewhere in Central and South America. Most of the crane flies known from Colombia were described by the same author, the great expert on crane fly taxonomy Charles Paul Alexander (1889-1981). As for the Neotropical Region in general, the identification of Colombian crane flies to the level of species can be very difficult, even for experts in crane fly taxonomy. There is no comprehensive identification tool available for the species, and the original descriptions are often very superficial. Often, only within the context of a taxonomic revision of the genus in question can the identification of the species in hand be done accurately. Up to now, the most recent taxonomic revision that included species occurring in Colombia is that of Ribeiro (2003) for the genus Styringomyia. Thus, it seems unnecessary to say that additional revisionary studies including members of the local fauna are much needed. Additional sampling efforts are also necessary in order to improve our knowledge about the Colombian fauna, as for the Neotropical fauna in general. Catalogue of Tipulidae and Limoniidae of Colombia Family Tipulidae Subfamily Tipulinae Genus Brachypremna Osten Sacken Brachypremna Osten Sacken, 1887: 166. Type species, Tipula dispellens Walker, 1861 (designation: Coquillett, 1910). dispellens (Walker) 1861: 334 (Tipula). Type locality: “Mexico”. Distr.: USA, Brazil, Colombia (Boyacá, Muzo: Alexander, 1938a), Costa Rica, Guatemala, Mexico, Panama, Suriname, Trinidad, Venezuela. 74 · Zootaxa 4122 (1) © 2016 Magnolia Press RIBEIRO & SANTOS Genus Leptotarsus Guerin-Meneville Leptotarsus Guerin-Meneville, 1831. Type species, Leptotarsus macquartii Guerin-Meneville, 1831 (monotypic). Semnotes Westwood, 1876: 501 (as genus). Type species, Semnotes imperatoria Westwood, 1876 (designation: Pierre, 1926). Subgenus Pehlkea Enderlein Pehlkea Enderlein, 1912: 15 (as genus). Type species, Pehlkea columbiana Enderlein, 1912 (original designation). columbianus (Enderlein), 1912: 15 (Pehlkea). Type locality: “Colombia”. Distr.: Colombia. regina (Alexander), 1914a: 214 (Tanypremna). Type locality: Colombia, Cordillera Central. Distr.: Colombia (La Vega, Cordillera Central: Alexander, 1914a). Subgenus Tanypremna Osten Sacken Tanypremna Osten Sacken, 1886: 19 (as genus). Type species, Tanypremna opilio Osten Sacken, 1886 (monotypic). Stegasmonotus Enderlein, 1912: 19 (as genus). Type species, Tanypremna opilio Osten Sacken, 1886 (monotypic). elegantior (Alexander), 1945a: 51 (Tanypremna). Type locality: “Colombia”. Distr.: Colombia (Alexander, 1945a). fuscitarsis (Alexander), 1919: 149 (Tanypremna). Type locality: Colombia, Magdalena, Mt. Santa Marta. Distr.: Colombia (Magdalena, Mt. Santa Marta: Alexander, 1919). longipes (Fabricius), 1805: 25 (Tipula). Type locality: “South America”. Distr.: Brazil, Colombia, Guyana, Peru. Genus Nephrotoma Meigen Pales Meigen, 1800. Type species, Tipula dorsalis Fabricius, 1781 (designation: Hendel, 1908). Name suppressed by I.C.Z.N., 1963: 20: Opinion 678. Nephrotoma Meigen, 1803: 262 (as genus). Type species, Tipula dorsalis Fabricius, 1781 (monotypic). Pachyrrhina Macquart, 1834: 88 (as genus). Type species, Tipula crocata Linnaeus, 1758 (designation: Westwood, 1840). alleni (Alexander), 1913a: 206 (Pachyrhina). Type locality: Colombia, Cauca, Valle de Las Papas. Distr.: Colombia (Cauca,Valle de Las Papas: Alexander, 1913a), Ecuador. globosa (Alexander), 1969a: 200 (Nephrotoma). Type locality: “Colombia”. Distr.: Colombia (Caldas, Manizales: Alexander, 1969a). medioligula (Alexander), 1945b: 67 (Nephrotoma). Type locality: Venezuela, Aragua, Rancho Grande. Distr.: Colombia, Venezuela. Genus Ozodicera Macquart Ozodicera Macquart, 1834: 92. Type species, Ozodicera ochracea Macquart, 1834 (original designation) [= pectinata (Wiedemann, 1828)] Hemicteina Westwood, 1835: 450 (as genus). Type species, Hemicteina gracilis Westwood, 1835 (monotypic). Subgenus Dihexaclonus Enderlein Dihexaclonus Enderlein, 1912: 28 (as genus). Type species, Ozodicera apicalis Macquart, 1838 (original designation). CATALOGUE OF TIPULIDAE AND LIMONIIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 75 telestyla (Alexander), 1969a: 210 (Ozodicera). Type locality: “Colombia”. Distr.: Colombia (Meta, Puerto Lopez: Alexander, 1969a). Genus Tipula Linnaeus Tipula Linnaeus, 1758: 585. Type species, Tipula oleracea Linnaeus, 1758 (designation: Latreille, 1810). Subgenus Bellardina Edwards Bellardina Edwards, 1931: 82. Type species, Tipula craverii Bellardi, 1859 (original designation, as cravieri). obliquefasciata (Macquart), 1846: 143(15) (Tipula). Type locality: Colombia [New Grenada], temperate region. Distr.: Colombia. Subgenus Eumicrotipula Alexander Eumicrotipula Alexander, 1923: 74 (as subgenus of Microtipula). Type species, Microtipula macrotrichiata Alexander, 1923 (original designation). armillata (Alexander), 1916: 30 (Tipula). Type locality: Colombia, Magdalena, La Cumbre. Distr.: Colombia (Magdalena, La Cumbre, Mt. Santa Marta; Boyaca, Popayan: Alexander, 1916), Bolivia, Ecuador, Venezuela. bogotana (Alexander), 1938b: 327 (Tipula). Type locality: Colombia, Bogotá. Distr.: Colombia (Bogotá: Alexander, 1938b). carizona (Alexander), 1913a: 208 (Tipula). Type locality: Colombia, Cauca, Valle de Las Papas. Distr.: Colombia (Cauca, Valle de Las Papas: Alexander, 1913a). foersteriana (Alexander), 1967a: 461 (Tipula). Type locality: Colombia, Manizales. Distr.: Colombia (Caldas, Manizales: Alexander, 1967a), Ecuador, Peru. laterosetosa (Alexander), 1931: 623 (Tipula). Type locality: Colombia, Magdalena, Mt. Santa Marta, Vista Nieve. Distr.: Colombia (Magdalena, Vista Nieve, Mt. Santa Marta: Alexander, 1931). mecoglossa (Alexander), 1967a: 464 (Tipula). Type locality: Colombia, Cundinamarca, Monterredondo. Distr.: Colombia (Cundinamarca, Monterredondo: Alexander, 1967a). miranha (Alexander), 1913a: 210 (Tipula). Type locality: Colombia, Cauca, Valle de Las Papas. Distr.: Colombia (Cauca, Valle de Las Papas: Alexander, 1913a). mitua (Alexander, 1916). Alexander, 1916: 29 (Tipula). Type locality: Colombia, Cauca, Valle de Las Papas. Distr.: Colombia (Cauca, Valle de Las Papas: Alexander, 1916). mocoa (Alexander), 1913a: 210 (Tipula). Type locality: Colombia, Cauca, Valle de Las Papas. Distr.: Colombia (Cauca, Valle de Las Papas: Alexander, 1913a). moniliferoides (Alexander), 1920a: 221 (Tipula). Type locality: Argentina, Buenos Aires, Monte Veloz. Distr.: Colombia, Argentina, Brazil. moniliformis (Röder), 1886: 259 (Tipula). Type locality: Colombia, Cerro del Altar. Distr.: Colombia. 76 · Zootaxa 4122 (1) © 2016 Magnolia Press RIBEIRO & SANTOS ornaticornis (Wulp), 1891: 195 (Tipula). Type locality: “Colombia” [New Grenada]. Distr.: Colombia. pallidineuris (Macquart), 1846: 144(16) (Tipula). Type locality: “Colombia” [New Grenada]. Distr.: Colombia. redunca (Alexander), 1967a: 469 (Tipula). Type locality: Colombia, Manizales. Distr.: Colombia (Caldas, Manizales: Alexander, 1967a). riveti tolimensis (Alexander), 1941: 244 (Tipula). Type locality: Colombia, Cordillera Central, Mt. Tolima. Distr.: Colombia (Mt. Tolima, Cordillera Central: Alexander, 1941). Subgenus Microtipula Alexander Microtipula Alexander, 1912a: 360 (genus). Type species: Microtipula amazonica Alexander, 1912 (original designation). Nephrotomodes Alexander, 1946: 224 (as subgenus of Tipula). Type species: Tipula smilodon Alexander, 1940 (original designation). colombicola (Alexander), 1929a: 768 (Tipula). Type locality: Colombia, Chiriguana, Lake Sapatoza Region. Distr.: Colombia (Cesar, Lake Zapatosa, Chiriguana: Alexander, 1929a) erostrata (Alexander), 1945c: 424 (Tipula). Type locality: Colombia, Boyacá, Muzo. Distr.: Colombia (Boyacá, Muzo: Alexander, 1945c). multimoda (Alexander), 1938b: 328 (Tipula). Type locality: Colombia, Puerto Boyacá, Magdalena Basin. Distr.: Colombia (Boyaca, Puerto Boyacá, Magdalena: Alexander, 1938b). perangustula (Alexander), 1938a: 431 (Tipula). Type locality: Colombia, Boyacá, Muzo. Distr.: Colombia (Boyacá, Muzo: Alexander, 1938a). tenuicula (Enderlein), 1912: 7 (Tipula). Type locality: “Colombia”. Distr.: Colombia. urophora (Alexander), 1938a: 429 (Tipula). Type locality: Ecuador, Los Rios, Playas de Montalvo. Distributions: Colombia (Boyacá, Muzo: Alexander, 1938a), Ecuador. Genus Zelandotipula Alexander Zelandotipula Alexander, 1922a: 522 (as subgenus of Holorusia). Type species, Tipula novarae Schiner, 1868 (original designation). austrofurcifera (Alexander), 1969a: 236 (Zelandotipula). Type locality: Colombia, Monterredondo, Cundinamarca. Distr.: Colombia (Cundinamarca, Monterredondo: Alexander, 1969a), Ecuador. corynostyla (Alexander), 1969a: 237 (Zelandotipula). Type locality: Colombia, Monterredondo, Cundinamarca. Distr.: Colombia (Cundinamarca, Monterredondo: Alexander, 1969a). longitarsis (Macquart), 1846: 145 (Tipula). Type locality: “Venezuela”. Distr.: Colombia, Venezuela. orophila (Alexander), 1914b: 354 (Holorusia). Type locality: Colombia, San Lorenzo Mountain. Distr.: Colombia (Magdalena, Sierra San Lorenzo: Alexander, 1914b). retrorsa (Alexander), 1969a: 243 (Zelandotipula). Type locality: Colombia, Monterredondo, Cundinamarca. Distr.: Colombia (Cundinamarca, Monterredondo: Alexander, 1969a). CATALOGUE OF TIPULIDAE AND LIMONIIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 77 wardiana (Alexander), 1981a: 9 (Zelandotipula). Type locality: Colombia, Magdalena, 18 km east of San Pedro. Distr.: Colombia (Magdalena: Alexander, 1981). Family Limoniidae Subfamily Chioneinae Genus Atarba Osten Sacken Atarba Osten Sacken, 1869: 127. Type species, Atarba picticornis Osten Sacken, 1869 (monotypic). Subgenus Atarba Osten Sacken Atarba Osten Sacken, 1869: 127 (as genus). Type species, Atarba picticornis Osten Sacken, 1869 (monotypic). brunneicornis (Alexander), 1916: 15 (Atarba). Type locality: Colombia, Magdalena, La Cumbre. Distr.: Colombia (Magdalena, La Cumbre, Mt. Santa Marta: Alexander, 1916). columbiana (Alexander), 1913a: 199 (Atarba). Type locality: Colombia, Cauca, Almaguer. Distr.: Colombia (Cauca, Almaguer: Alexander 1913a). Subgenus Ischnothrix Bigot Lachnocera Philippi, 1866: 615 (as genus). Type species, Lachnocera delicatula Philippi, 1866 (monotypic). Ichnothrix Bigot, 1888: 7 (as genus). Type species, Ischnothrix aetherea Bigot, 1888 (monotypic). Oromyia Alexander, 1913a: 203 (as genus). Type species, Oromyia lloydi Alexander, 1913a (original designation). Orolimnophila Alexander, 1921a: 178. Type species, Oromyia lloydi Alexander, 1913a. lloydi (Alexander), 1913a: 204 (as Oromyia). Type locality: Colombia, Cauca, Valle de Las Papas. Distr.: Colombia (Cauca, Valle de Las Papas: Alexander, 1913a). Genus Erioptera Meigen Polymeda Meigen, 1800. Type species, Erioptera lutea Meigen, 1804 (designation: Coquillett, 1910). Name suppressed by I.C.Z.N., 1963: 20: Opinion 678. Erioptera Meigen, 1803: 262. Type species, Erioptera lutea Meigen, 1804 (designation: Coquillett, 1910). Subgenus Erioptera Meigen Polymeda Meigen, 1800. Type species, Erioptera lutea Meigen, 1804 (designation: Coquillett, 1910). Name suppressed by I.C.Z.N., 1963: 20: Opinion 678. Erioptera Meigen, 1803: 262 (as genus). Type species, Erioptera lutea Meigen, 1804 (designation: Coquillett, 1910). Polyraphia Meigen, 1818: 111 (as genus). Type species, Erioptera taenionota Meigen, 1818 (monotypic). Polyrhaphia Agassiz, 1847: 304. Type species, Erioptera taenionota Meigen, 1818 (monotypic). Octavia Bigot, 1854: 456, 474 (as genus). Type species, Erioptera taenionota Meigen, 1818 (designation: Coquillett, 1910). Chemalida Rondani, 1856: 180 (as genus), Type species, Erioptera taenionota Meigen, 1818 (original designation). Ilisophila Rondani, 1856: 180 (as genus), Type species, Erioptera lutes Meigen, 1804 (original designation). Limnoea Rondani, 1856: 180 (as genus), Type species, Erioptera flavescens Meigen, 1804 (original designation). Limnoica Rondani, 1861: 180 (as genus), Type species, Erioptera flavescens Meigen, 1804 (original designation). andina (Alexander), 1913a: 200 (Erioptera). Type locality: Colombia, Cauca, Valle de Las Papas. Distr.: Argentina, Bolivia, Colombia (Cauca, Valle de Las Papas: Alexander, 1913a). 78 · Zootaxa 4122 (1) © 2016 Magnolia Press RIBEIRO & SANTOS annulipes (Williston), 1896: 294 (Erioptera). Type locality: Lesser Antilles, St. Vincent. Distr.: Brazil, Colombia, Guatemala, Paraguay, Peru, St. Vincent. quinquecincta (Alexander), 1927a: 278 (Erioptera). Type locality: Colombia, Caldas. Distr.: Colombia (Caldas: Alexander, 1927a). Genus Eugnophomyia Alexander Eugnophomyia Alexander, 1947a: 72 (as subgenus of Gnophomyia). Type species, Gnophomyia luctuosa Osten Sacken, 1860 (original designation). luctuosa (Osten Sacken), 1860: 224 (Gnophomyia). Type locality: U.S.A., Florida. Distr.: USA, Colombia, Guatemala, México. Genus Gnophomyia Osten Sacken Gnophomyia Osten Sacken, 1860: 223 (as genus). Type species, Gnophomyia tristissima Osten Sacken, 1860 (designation: Coquillett, 1910). acricula (Alexander), 1931: 637 (Gnophomyia). Type locality: Colombia, Magdalena, Mt. Santa Marta, Vista Nieve. Distr.: Colombia (Magdalena, Mt. Santa Marta, Vista Nieve: Alexander, 1931). vilis (Alexander), 1931: 638 (Gnophomyia). Type locality: Colombia, Orihueca. Distr.: Colombia (Magdalena, Orihueca: Alexander, 1931). Genus Gonomyia Meigen Gonomyia Meigen, 1818: 147 (as genus). Type species, Limnobia tenella Meigen, 1818 (monotypic). Taphrosia Rondani, 1856: 182 (as genus). Type species, Limnobia tenella Meigen, 1818 (original designation). Goniomyia Osten Sacken, 1869: 176. Type species, Limnobia tenella Meigen, 1818 (monotypic). Subgenus Gonomyia Meigen Gonomyia Meigen, 1818: 147 (as genus). Type species, Limnobia tenella Meigen, 1818 (monotypic). Taphrosia Rondani, 1856: 182 (as genus). Type species, Limnobia tenella Meigen, 1818 (original designation). Goniomyia Osten Sacken, 1869: 176. Type species, Limnobia tenella Meigen, 1818 (monotypic). andicola (Alexander) 1913a: 199 (Gonomyia). Type locality: Colombia, Cauca, Valle de Las Papas. Distr.: Colombia (Cauca, Valle de Las Papas: Alexander, 1913a). methodica (Alexander), 1931: 635 (as Gonomyia). Type locality: Colombia, Magdalena, Mt. Santa Marta, Vista Nieve. Distr.: Colombia (Magdalena, Vista Nieve, Mt. Santa Marta: Alexander, 1931). Subgenus Leiponeura Skuse Leiponeura Skuse, 1890: 795 (as genus). Type species, Leiponeura gracilis Skuse, 1890 (designation: Alexander, 1913b) [= skusei Alexander 1919]. Lipophleps Bergroth, 1915: 55 (as [unnecessary] new name for Leiponeura Skuse, 1890 [not preoccupied by Liponeura Loew, 1844]). Type species, Leiponeura gracilis Skuse, 1890. CATALOGUE OF TIPULIDAE AND LIMONIIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 79 cervaria (Alexander), 1938b: 334 (Gonomyia). Type locality: Colombia, Cundinamarca, Fusagasugá. Distr.: Colombia (Cundinamarca, Fusagasugá: Alexander, 1938b). Subgenus Paralipophleps Alexander Paralipophleps Alexander, 1947a: 97 (as subgenus of Gonomyia). Type species, Atarba pleuralis Williston, 1896 (original designation). lemniscata (Alexander), 1931: 634 (Gonomyia). Type locality: Colombia, Magdalena, Mt. Santa Marta, Vista Nieve. Distr.: Colombia (Magdalena, Vista Nieve, Mt. Santa Marta: Alexander, 1931). Genus Molophilus Curtis Monophilus Curtis, 1833: 444 (as genus). Type species, Molophilus brevipennis Curtis, 1833 (original designation) [= ater Meigen, 1804)]. Archimolophilus Enderlein, 1938: 669 (as genus). Type species, Archimolophilus selkirkianus Enderlein, 1938 (monotypic). Subgenus Molophilus Curtis Monophilus Curtis, 1833: 444 (as genus). Type species, Molophilus brevipennis Curtis, 1833 (original designation). Archimolophilus Enderlein, 1938: 669 (as genus). Type species, Archimolophilus selkirkianus Enderlein, 1938 (monotypic). capricornis (Alexander), 1916: 19 (Molophilus). Type locality: Colombia, Magdalena, La Cumbre. Distr.: Colombia (Magdalena, La Cumbre, Mt. Santa Marta: Alexander, 1916), Peru. conscriptus (Alexander), 1938b: 337 (Molophilus). Type locality: Colombia, Cundinamarca, Fusagasugá. Distr.: Colombia (Cundinamarca, Fusagasugá: Alexander, 1938b). flexilistylus (Alexander), 1931: 639 (Molophilus). Type locality: Colombia, Magdalena, Mt. Santa Marta, La Cumbre. Distr.: Colombia (Magdalena, La Cumbre, Mt. Santa Marta: Alexander, 1931). lictor (Alexander), 1938b: 336 (Molophilus). Type locality: Colombia, Cundinamarca, Fusagasugá. Distr.: Colombia (Cundinamarca, Fusagasugá: Alexander, 1938b). luxuriosus (Alexander), 1938b: 335 (Molophilus). Type locality: Colombia, Usme. Distr.: Colombia (Bogota, Usme: Alexander, 1938b). marthae (Alexander), 1931: 641 (Molophilus). Type locality Colombia (Magdalena, Mt. Santa Marta, Vista Nieve. Distr.: Colombia (Magdalena, Vista Nieve, Mt. Santa Marta: Alexander, 1931). perseus (Alexander), 1913a: 201 (Molophilus). Type locality: Colombia, Cauca, Valle de Las Papas. Distr.: Colombia (Cauca, Valle de Las Papas: Alexander, 1913a). subtenebricosus (Alexander), 1931: 640 (Molophilus): Type locality: Colombia (Magdalena, Mt. Santa Marta, Vista Nieve. Distr.: Colombia (Magdalena, Vista Nieve, Mt. Santa Marta: Alexander, 1931). tenebricosus (Alexander), 1916: 20 (Molophilus). Type locality: Colombia, Magdalena, Mt. Santa Marta, La Cumbre. Distr.: Colombia (Magdalena, La Cumbre, Mt. Santa Marta: Alexander, 1916). walkeri (Alexander), 1931: 640 (Molophilus). Type locality: Colombia, Magdalena, Mt. Santa Marta, Vista Nieve. Distr.: Colombia (Magdalena, Vista Nieve, Mt. Santa Marta: Alexander, 1931). 80 · Zootaxa 4122 (1) © 2016 Magnolia Press RIBEIRO & SANTOS Genus Neognophomyia Alexander Neognophomyia Alexander, 1926: 391 (as subgenus of Gnophomyia). Type species, Gnophomyia immaculipennis Alexander, 1926 (original designation). colombicola (Alexander), 1931: 636 (Neognophomyia). Type locality: Colombia, Magdalena, St. Marta, Vista Nieve. Distr.: Colombia (Magdalena, Vista Nieve, Mt. Santa Marta: Alexander, 1931). Genus Rhabdomastix Skuse Palaeogonomyia Meunier, 1899: 359 (as genus). Type species, Gonomyia pulcherrima Meunier, 1906. Rhabdomastix Skuse, 1890: 828 (as genus). Type species, Rhabdomastix ostensackeni Skuse, 1890 (monotypic). Sacandaga Alexander, 1911: 349 (as genus). Type species, Sacandaga flava Alexander, 1911. Subgenus Rhabdomastix Skuse Palaeogonomyia Meunier, 1899: 359 (as genus). Type species, Gonomyia pulcherrima Meunier, 1906. Rhabdomastix Skuse, 1890: 828 (as genus). Type species, Rhabdomastix ostensackeni Skuse, 1890 (monotypic). Sacandaga Alexander, 1911: 349 (as genus). Type species, Sacandaga flava Alexander, 1911. tantilla (Alexander), 1938a: 440 (Rhabdomastix). Type locality: Colombia, Meta, Restrepo. Distr.: (Meta, Restrepo: Alexander, 1938a). Genus Styringomyia Loew Styringomyia Loew, 1845: 6 (as genus). Type species, Styringomyia venusta Loew, 1845 (monotypic) (fossil). Idiophlebia Grunberg, 1903: 524 (as genus). Type species, Idiophlebia pallida Grunberg, 1903 (monotypic). Pycnocrepis Enderlein, 1912: 65 (as genus). Type species, Pycnocrepis annulipes Enderlein, 1912 (original designation). Syringomyia Alexander, 1972: 841 (unjustified emendation of Styringomyia). maya Ribeiro, 2003: 22. Type locality: Ecuador, Azuay, 18 km SE Jesus Maria. Distr.: Belize, Bolivia, Colombia (Valle del Cauca, Sevilla: Ribeiro, 2003), Costa Rica, Ecuador, Guyana, Honduras, Panama, Suriname, Venezuela. Genus Symplecta Meigen Helobia Lepeletier and Serville, 1828: 585 (as genus). Type species, Limnobia punctipennis Meigen, 1818 (monotypic). Symplecta Meigen, 1830: 282 (as genus). Type species, Limnobia punctipennis Meigen, 1818 (designation: Westwood, 1840) [= hybrida Meigen, 1804)]. Idioneura Philippi, 1866: 615 (as genus). Type species, Idioneura macroptera Philippi, 1866 (monotypic). Subgenus Symplecta Meigen Helobia Lepeletier and Serville, 1828: 585 (as genus). Type species, Limnobia punctipennis Meigen, 1818 (monotypic). Symplecta Meigen, 1830: 282 (as genus). Type species, Limnobia punctipennis Meigen, 1818 (designation: Westwood, 1840) [= hybrida Meigen, 1804)]. Idioneura Philippi, 1866: 615 (as genus). Type species, Idioneura macroptera Philippi, 1866 (monotypic). colombiana colombiana (Alexander), 1937: 616 (Helobia). Type locality: Colombia, Usme. Distr.: Colombia (Bogota, Usme: Alexander, 1937). CATALOGUE OF TIPULIDAE AND LIMONIIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 81 Genus Teucholabis Osten Sacken Teucholabis Osten, 1860: 222 (as genus). Type species, Teucholabis complexa Osten Sacken, 1860 (monotypic). Subgenus Paratropesa Schiner Paratropesa Schiner, 1866: 932 (as genus). Type species, Paratropesa singularis Schiner, 1868 (original designation). chalybeia (Alexander), 1927b: 18 (Teucholabis). Type locality: Colombia: El Líbano. Distr.: Colombia (Madgalena, El Libano: Alexander, 1927b). Subgenus Teucholabis Osten Sacken Teucholabis Osten Sacken, 1860: 222 (as genus). Type species, Teucholabis complexa Osten Sacken, 1860 (monotypic). jocosa (Alexander), 1913c: 440 (Teucholabis). Type locality: Bolivia, Yungas de Coroico. Distr.: Argentina, Bolivia, Brazil, Colombia (La Cumbre, Mt. Santa Marta; Cali: Alexander, 1913c), Ecuador, Peru, Venezuela. marticola (Alexander), 1931: 633 (Teucholabis). Type locality: Colombia, Magdalena, Mt. Santa Marta. Distr.: Colombia (Magdalena, Mt. Santa Marta: Alexander, 1931). megaphallus (Alexander), 1968: 219 (Teucholabis). Type locality: Colombia, Monterredondo. Distr.: Colombia (Cundinamarca). salti (Alexander), 1930: 738 (Teucholabis). Type locality Colombia, Magdalena, Rio Frío. Distr.: Colombia (Magdalena, Rio Frio; Sevilla; Orihueca: Alexander, 1930). spinigera (Schiner), 1868: 44 (Teucholabis). Type locality: “Venezuela”. Distr.: Colombia, Peru, Venezuela. trifasciata trifasciata (Enderlein), 1912: 69 (Teucholabis). Type locality: “South America”. Distr.: Colombia (Rio Magdalena: Enderlein, 1912), Ecuador, Guyana, México, Panamá, Peru, Venezuela. venezuelensis (Macquart), 1846: 147 (Limnobia). Type locality: “South America”. Distr.: Colombia, Venezuela. Subfamily Limnophilinae Genus Epiphragma Osten Sacken Epiphragma Osten Sacken, 1860: 238 (as subgenus of Limnophila). Type species, Limnophila pavonina Osten Sacken, 1860 (designation: Coquillett, 1910) [= fasciapenne (Say, 1823)]. Subgenus Epiphragma Osten Sacken Epiphragma Osten Sacken, 1860: 238 (as subgenus of Limnophila). Type species, Limnophila pavonina Osten Sacken, 1860 (designation: Coquillett, 1910) [= fasciapenne (Say, 1823)]. cordillerense (Alexander), 1913a: 202 (Epiphragma). Type locality: Colombia, Cauca, Popayán. Distr.: Colombia (Cauca, Valle de las Papas; Popayán: Alexander, 1913a), Bolivia, Peru. 82 · Zootaxa 4122 (1) © 2016 Magnolia Press RIBEIRO & SANTOS delicatulum (Osten Sacken), 1888: 208 (Epiphragma). Type locality: “Colombia”. Distr.: Colombia, Venezuela. gracilicorne (Alexander), 1916: 21 (Epiphragma). Type locality: Colombia, Magdalena, La Cumbre. Distr.: Colombia (Magdalena, La Cumbre, Mt. Santa Marta: Alexander, 1916). Genus Hexatoma Latreille Hexatoma Latreille, 1809: 260 (as genus). Type species, Hexatoma nigra Latreille, 1809 (monotypic). Nematocera Meigen, 1818: 209. Type species, Nematocera bicolor Meigen, 1818 (designation: Savchenko et al., 1992). Anisomera Meigen, 1818: 210. Type species, Anisomera obscura Meigen, 1818 (monotypic). Peronecera Curtis, 1836: 589 Type species, Peronecera fuscipennis Curtis, 1836 (monotypic). Trimacromera Enderlein, 1936: 23. Type species, Nematocera nubeculosa Burmeister, 1829 (monotypic). Subgenus Eriocera Macquart Eriocera Macquart, 1838: 74 (as genus). Type species, Limnobia nigra (Macquart, 1838), misidentification, not Wiedemann, 1828 (monotypic) [= macquarti (Enderlein, 1912)]. Caloptera Guérin-Méneville, 1831 (as genus). Type species, Caloptera fasciata Guerin-Meneville, 1831 (original designation). Arrhenica Osten Sacken, 1860: 243 (as genus). Type species, Arrhenica spinosa Osten Sacken, 1860 (designation: Coquillett, 1910). Penthoptera Schiner, 1863: 220 (as genus). Type species, Tipula chirotheca Scopoli, 1763 (original designation). antennata (Alexander), 1930: 735 (Eriocera). Type locality: Colombia, Magdalena, Sevilla. Distr.: Colombia (Magdalena, Sevilla: Alexander, 1930). bequaertiana (Alexander), 1938a: 437 (Hexatoma). Type locality: Colombia, Boyacá, Muzo. Distr.: Colombia (Boyacá, Muzo: Alexander, 1938). braconides (Enderlein), 1912: 47 (Eriocera). Type locality: “Colombia”. Distr.: Colombia. columbiana (Alexander), 1919: 147 (Eriocera). Type locality: Colombia, Bolívar. Distr.: Colombia (Bolívar: Alexander, 1919), Costa Rica ohausiana (Enderlein), 1912: 45 (Eriocera). Type locality: Ecuador, Sabanilla. Distr.: Bolivia, Colombia, Ecuador, Peru. perlaeta (Alexander), 1922b: 54 (Eriocera): Type locality: “Colombia”. Distr.: Brazil, Colombia. sanctaemartae (Alexander), 1919: 148 (Penthoptera). Type locality: Colombia, Magdalena, Mt. Santa Marta. Distr.: Colombia (Magdalena, Mt. Santa Marta: Alexander, 1919). schineri (Alexander), 1922b: 60 (Eriocera). Type locality: “Colombia”. Distr.: Colombia, Venezuela. virgulativentris (Enderlein), 1912: 47 (Eriocera). Type locality: Colombia, Tolima, Natagaima. Distr.: Colombia (Tolima, Natagaima: Enderlein, 1912). Genus Lecteria Osten Sacken Lecteria Osten Sacken, 1888: 206 (as genus). Type species, Tipula armillaris Fabricius, 1805 (original designation). CATALOGUE OF TIPULIDAE AND LIMONIIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 83 Subgenus Lecteria Osten Sacken Lecteria Osten Sacken, 1888: 206 (as genus). Type species, Tipula armillaris Fabricius, 1805 (original designation). retrorsa (Alexander), 1969b: 337 (Lecteria). Type locality: Colombia, Sierra San Lorenzo. Distr.: Colombia (Magdalena, Sierra San Lorenzo: Alexander, 1969b). Genus Limnophila Macquart. Limnophila Macquart, 1834: 95 (as genus). Type species, Limnobia pictipennis Meigen, 1818 (designation: Westwood, 1840). Limnomya Rondani, 1861: 11 (as [unnecessary] new name for Limnophila Macquart, 1834 [not preoccupied in zoology]). Type species, Limnobia pictipennis Meigen, 1818 (autom.). Poecilostola Schiner, 1863: 222. Type species, Limnobia pictipennis Meigen, 1818 (original designation). Subgenus Limnophila Macquart Limnophila Macquart, 1834: 95 (as genus). Type species, Limnobia pictipennis Meigen, 1818 (designation: Westwood, 1840). Limnomya Rondani, 1861: 11 (as genus) (as [unnecessary] new name for Limnophila Macquart, 1834 [not preoccupied in zoology]). Type species, Limnobia pictipennis Meigen, 1818 (autom.). Poecilostola Schiner, 1863: 222 (as genus). Type species, Limnobia pictipennis Meigen, 1818 (original designation). lloydi (Alexander), 1913a: 205 (Limnophila) Type locality: Colombia, Cauca, Valle de Las papas. Distr.: Colombia (Cauca, Valle de Las Papas: Alexander, 1913a). Genus Polymera Wiedemann. Polymera Wiedemann, 1820: 40 (as genus). Type species, Chironomus hirticornis Fabricius, 1805 (monotypic). Subgenus Polymera Wiedemann Polymera Wiedemann, 1820: 40 (as genus). Type species, Chironomus hirticornis Fabricius, 1805 (monotypic). ominosa (Alexander), 1938b: 332 (as Polymera). Type locality: Colombia, Cundinamarca, Fusagasugá. Distr.: Colombia (Cundinamarca, Fusagasugá: Alexander, 1938b). sordidipes (Alexander), 1938b: 332 (as Polymera). Type locality: Colombia, Cundinamarca, Fusagasugá. Distr.: Colombia (Cundinamarca, Fusagasugá: Alexander, 1938b). Genus Shannonomyia Alexander Shannonomyia Alexander, 1929: 142 (as genus). Type species, Limnophila lenta Osten Sacken, 1860 (original designation). Subgenus Shannonomyia Alexander Shannonomyia Alexander, 1929: 142 (as genus). Type species, Limnophila lenta Osten Sacken, 1860 (original designation). bogotensis (Alexander), 1938b: 331 (Shannonomyia). Type locality: Colombia, Usme. Distr.: Colombia (Bogota, Usme: Alexander, 1938b) 84 · Zootaxa 4122 (1) © 2016 Magnolia Press RIBEIRO & SANTOS lathraea (Alexander), 1926: 386 (Pilaria). Type locality: Colombia, San Lorenzo, Cincinnati. Distr.: Colombia (Magdalena, Hacienda Cincinnati, Sierra San Lorenzo: Alexander, 1926), Venezuela. orophila (Alexander), 1913a: 205 (Limnophila). Type locality: Colombia, Cauca, Almaguer. Distr.: Colombia (Cauca, Almaguer: Alexander, 1913a). Subfamily Limoniinae Genus Dicranomyia Stephens Furcomyia Meigen, 1818: 133 (as genus). Type species, Limnobia lutea Meigen, 1818 (monotypic). Dicranomyia Stephens, 1829: 53 (as genus). Type species, Limnobia modesta Meigen, 1818 (designation: Coquillett, 1910). Telecephala Pierre, 1922: 149 (as genus). Type species, Limnobia longicollis Macquart, 1846 (monotypic). Tedotea Santos Abreu, 1923: 143 (as genus). Type species, Tedotea domestica Santos Abreu, 1923 (monotypic). Subgenus Dicranomyia Stephens Furcomyia Meigen, 1818: 133 (as genus). Type species, Limnobia lutea Meigen, 1818 (monotypic). Dicranomyia Stephens, 1829: 53 (as genus). Type species, Limnobia modesta Meigen, 1818 (designation: Coquillett, 1910). Telecephala Pierre, 1922: 149 (as genus). Type species, Limnobia longicollis Macquart, 1846 (monotypic). Tedotea Santos Abreu, 1923: 143 (as genus). Type species, Tedotea domestica Santos Abreu, 1923 (monotypic). elegantula (Alexander), 1913a: 195. (Dicranomyia). Type locality: Colombia, Cauca, Almaguer. Distr.: Colombia (Cauca, Almaguer: Alexander, 1913a) insignifica (Alexander), 1912b: 363 (Furcomyia). Type locality: Peru, Iquico. Distr.: Colombia, Peru. longiventris (Alexander), 1913a: 198. (Dicranomyia). Type locality: Colombia, Cauca, Valle de Las Papas. Distr.: Colombia (Cauca, Valle de Las Papas: Alexander, 1913a). pampoecila (Alexander), 1922b: 1 (Limonia). Type locality: Costa Rica, Tiribi. Distr.: Brazil, Colombia (Chinquiquita: Alexander, 1938a), Costa Rica, Ecuador, Panamá, Peru, Venezuela. quadrituberculata (Alexander), 1937: 601 (Limonia). Type locality: Colombia, Usme near Bogotá. Distr.: Colombia (Bogota, Usme: Alexander, 1937). subandicola (Alexander), 1947b: 60 (Limonia). Type locality: Mexico, Mexico D.F., El Junquito. Distr.: Bolivia, Colombia (Cauca, Almaguer: Alexander, 1947b), México, Peru, Venezuela. teinoterga (Alexander), 1967b: 303 (Limonia). Type locality: Colombia, Mt Redondo. Distr.: Colombia (Cundinamarca, Monterredondo: Alexander, 1967b). Subgenus Neoglochina Alexander Neoglochina Alexander, 1967b: 316 (as subgenus of Limonia). Type species, Limonia felix Alexander, 1950 (original designation). capnora metae (Alexander), 1938a: 434 (Limonia). Type locality: Colombia, Meta, Restrepe. Distr.: Colombia (Meta, Restrepo: Alexander, 1938a). CATALOGUE OF TIPULIDAE AND LIMONIIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 85 Subgenus Neolimnobia Alexander Neolimnobia Alexander, 1928a: 20 (as subgenus of Dicranomyia). Type species, Limnobia diva Schiner, 1868 (original designation). diva (Schiner), 1868: 46 (as Limnobia). Type locality: “Brazil”. Distr.: Bolivia, Brazil, Colombia (Meta, Restrepo: Alexander, 1938a), Cuba, Jamaica, México, Panamá, Puerto Rico, Venezuela. Subgenus Peripheroptera Schiner Peripheroptera Schiner, 1868: 47 (as genus). Type species, Peripheroptera nitens Schiner, 1868 (original designation). nitens (Schiner), 1868: 47 (Peripheroptera). Type locality: Colombia, Venezuela. Distr.: Colombia, Panamá, Peru, Venezuela. Genus Elephantomyia Osten Sacken Elephantomyia Osten Sacken, 1860: 220 (as genus). Type species, Limnobiorhynchus canadensis Westwood, 1835, sensu Osten Sacken, 1860 (original designation) [= westwoodi Osten Sacken, 1869]. Subgenus Elephantomyia Osten Sacken Elephantomyia Osten Sacken, 1860: 220 (as genus). Type species, Limnobiorhynchus canadensis Westwood, 1835, sensu Osten Sacken, 1860 (original designation) [= westwoodi Osten Sacken, 1869]. angustissima (Alexander), 1931: 632 (Elephantomyia). Type locality: Colombia, Magdalena, Mt. Santa Marta, Vista Nieve. Distr.: Colombia (Magdalena, Vista Nieve: Mt. Santa Marta: Alexander, 1931), México. humilis (Alexander), 1931: 632 (Elephantomyia). Type locality: Colombia, Magdalena, Mt. Santa Marta, Vista Nieve. Distr.: Colombia (Magdalena, Vista Nieve: Mt. Santa Marta: Alexander, 1931). Genus Geranomyia Haliday. Geranomyia Haliday, 1833: 154 (as genus). Type species, Geranomyia unicolor Haliday, 1833 (monotypic). Limnobiorhynchus Westwood, 1836: 683 (as genus). Type species, Limnobiorhynchus brailiensis Westwood, 1836 (designation: Coquillett, 1910). Aporosa Macquart, 1839: 100 (as genus). Type species, Aporosa maculipennis Macquart, 1839 (designation: Coquillett, 1910). Pletussa Philippi, 1866: 597 (as genus). Type species, Pletussa virescens Philippi, 1866 (designation: Coquillett, 1910). Triphana Skuse, 1890: 777 (as subgenus of Geranomyia). Type species, Geranomyia lutulenta Skuse, 1890 (designation: Alexander, 1920b). Tetraphana Skuse, 1890: 778 (as subgenus of Geranomyia). Type species, Geranomyia fusca Skuse, 1890 (monotypic). Monophana Edwards, 1912: 200 (as subgenus of Geranomyia). Type species, Geranomyia immaculata Edwards, 1912 (monotypic). Pseudaporosa Alexander, 1924: 177 (as subgenus of Geranomyia). Type species, Geranomyia venustithorax Alexander, 1924 (original designation). Parageranomyia Santos Abreu, 1923: 100 (as genus). Type species, Parageranomyia palmensis Santos Abreu, 1923 (monotypic). eurygramma eurygramma (Alexander), 1928b: 107 (as Geranomyia). Type species, México, Veracruz, Córdoba, Barranca of the Rio San Antonio. Distr.: Colombia, Dominica, Ecuador, México, Panamá. 86 · Zootaxa 4122 (1) © 2016 Magnolia Press RIBEIRO & SANTOS lacteitarsis (Alexander), 1922b: 34 (Geranomyia). Type locality: Colombia, Condoto. Distr.: Colombia (Choco, Condoto: Alexander, 1922b). laudanda (Alexander), 1938b: 330 (Limonia). Type locality: Colombia, Usme. Distr.: Colombia (Bogota, Usme), Venezuela. lineata (Enderlein), 1912: 73 (Geranomyia). Type locality: Colombia, Hacienda Pehlke. Distr.: Colombia (Tolima, Hacienda Pehlke: Enderlein, 1912). marthae (Alexander), 1930: 726 (Limonia). Type locality: Colombia, Magdalena, Mt, Santa Marta, Vista Nieve. Distr.: Bolivia, Colombia (Magdalena, Vista Nieve, Mt. Santa Marta: Alexander, 1930), Venezuela. neogaudens (Alexander), 1942: 784 (Limonia). Type locality: Peru, Ayacucho, Huanta, Lake Rasvilca. Distr.: Colombia, Peru. neonumenius (Alexander), 1930: 729 (Limonia). Type locality: Colombia, Magdalena, Mt, Santa Marta, Vista Nieve. Distr.: Bolivia, Colombia (Magdalena, Vista Nieve, Mt. Santa Marta). plumbeipleura (Alexander), 1916: 10 (Geranomyia). Type locality: Ecuador, Huigra. Distr.: Brazil, Colombia (Caldas; Magdalena, La Cumbre, Mt. Santa Marta), Dominica, Ecuador, Panama, Peru, Trinidad, Venezuela. recondita recondita (Alexander), 1921b: 42 (Geranomyia). Type locality: Peru, Iquitos. Distr.: Brazil, Colombia (Gorgona Is), Ecuador, Guyana, Panamá, Peru, Puerto Rico. versuta (Alexander), 1930: 728 (Limonia). Type locality: Colombia, Magdalena, Mt, Santa Marta, Vista Nieve. Distr.: Colombia (Magdalena, Vista Nieve, Mt. Santa Marta: Alexander, 1930), México. walkeri (Alexander), 1930: 730 (Limonia). Type locality: Colombia, Magdalena, Mt, Santa Marta, Vista Nieve. Distr.: Colombia (Magdalena, Vista Nieve, Mt. Santa Marta: Alexander, 1930), Brazil, Venezuela. Genus Helius Lepeletier & Serville, 1828 Megarhina Lepeletier and Serville, 1828: 585 (as genus). Type species, Limnobia longirostris Meigen, 1818 (monotypic). Helius Latreille et al., 1828: 655 (as genus). Type species, Limnobia longirostris Meigen, 1818 (autom.). Leptorhina Stephens, 1829: 243 (as genus). Type species, Limnobia longirostris Meigen, 1818 (monotypic). Rhamphidia Meigen, 1830: 281 (as genus). Type species, Limnobia longirostris Meigen, 1818 (designation: Westwood, 1840). Subgenus Helius Lepeletier & Serville, 1828 Megarhina Lepeletier and Serville, 1828: 585 (as genus). Type species, Limnobia longirostris Meigen, 1818 (monotypic). Helius Latreille et al., 1828: 655 (as genus). Type species, Limnobia longirostris Meigen, 1818 (autom.). Leptorhina Stephens, 1829: 243 (as genus). Type species, Limnobia longirostris Meigen, 1818 (monotypic). Rhamphidia Meigen, 1830: 281 (as genus). Type species, Limnobia longirostris Meigen, 1818 (designation: Westwood, 1840). albitarsis albitarsis (Osten Sacken), 1888: 184 (Rhamphidia). Type locality: “Puerto Rico”. Distr.: Colombia, Cuba, Dominica, Guyana, Jamaica, Peru, Puerto Rico, St. Vincent, Venezuela. Genus Limonia Meigen Amphinome Meigen, 1800 (as genus). Type species, Tipula tripunctata Fabricius, 1781 (designation: Coquillett, 1910). Name suppressed by I.C.Z.N., 1963: 20 (Opinion 678). CATALOGUE OF TIPULIDAE AND LIMONIIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 87 Limonia Meigen, 1803: 262 (as genus). Type species, Tipula tripunctata Fabricius, 1781 (autom.) [= phragmitidis (Schrank, 1781)]. Limnobia Meigen, 1818: 116 (as [unnecessary] new name for Limonia Meigen, 1803. Type species: Tipula tripunctata Fabricius, 1781. Unomyia Meigen, 1818: 135 (as genus). Type species, Tipula tripunctata Fabricius, 1781 (designation: Coquillett, 1910). Limnomyza Rondani, 1856: 185 (as genus). Type species, Tipula tripunctata Fabricius, 1781 (original designation). ocellata (Röder), 1886: 259 (Limnobia). Type locality: Colombia, Cerro del Altar. Distr.: Colombia. Genus Neolimonia Alexander Neolimonia Alexander, 1964: 28 (as subgenus of Limonia). Type species, Furcomyia eiseni Alexander, 1912 (original designation). cordillerensis (Alexander), 1913a: 196 (Dicranomyia). Type locality: Colombia, Cauca, Valle de Las Papas. Distr.: Colombia (Cauca, Valle de Las Papas: Alexander, 1913a). sanctaemartae (Alexander), 1930: 722 (Limonia). Type locality: Colombia, Magdalena, Mt, Santa Marta, La Cumbre. Distr.: Colombia (Magdalena, La Cumbre, Mt. Santa Marta: Alexander, 1930), Peru. Genus Orimarga Osten Sacken Orimarga Osten Sacken 1869: 120 (as genus). Type species, Limnobia alpina Zetterstedt, 1851 (designation: Coquillett, 1910) [= attenuata (Walker, 1848)]. Ninguis Wallengren, 1882: 183, 206 (as genus). Type species, Limnobia virgo Zetterstedt, 1851 (designation: Coquillett, 1910). Subgenus Orimarga Osten Sacken Orimarga Osten Sacken 1869: 120 (as genus). Type species, Limnobia alpina Zetterstedt, 1851 (designation: Coquillett, 1910) [= attenuata (Walker, 1848)]. Ninguis Wallengren, 1882: 183, 206 (as genus). Type species, Limnobia virgo Zetterstedt, 1851 (designation: Coquillett, 1910). andina (Alexander), 1916: 14 (Orimarga). Type locality: Colombia, Magdalena, Mt, Santa Marta, La Cumbre. Distr.: Colombia (Magdalena, La Cumbre, Mt. Santa Marta: Alexander, 1916). Genus Rhipidia Meigen Rhipidia Meigen, 1818: 153. Type species, Rhipidia maculata Meigen, 1818 (monotypic). Ceratostephanus Brunetti, 1911: 271. Type species, Ceratostephanus antennatus Brunetti, 1911 (monotypic). Arhipidia Alexander, 1912c: 6 (as subgenus of Rhipidia). Type species, Rhipidia domestica Osten Sacken, 1860 (designation: Alexander, 1950). Monorhipidia Alexander, 1912c: 6 (as subgenus of Rhipidia). Type species, Rhipidia domestica Osten Sacken, 1860 (designation: Alexander, 1950). Conorhipidia Alexander, 1914c: 117 (as subgenus of Rhipidia). Type species, Rhipidia conica Alexander, 1914c (original designation). Subgenus Rhipidia Meigen Rhipidia Meigen, 1818: 153 (as genus). Type species, Rhipidia maculata Meigen, 1818 (monotypic). Ceratostephanus Brunetti, 1911: 271 (as genus). Type species, Ceratostephanus antennatus Brunetti, 1911 (monotypic). Arhipidia Alexander, 1912c: 6 (as subgenus of Rhipidia). Type species, Rhipidia domestica Osten Sacken, 1860 (designation: Alexander, 1950). Monorhipidia Alexander, 1912c: 6 (as subgenus of Rhipidia). Type species, Rhipidia domestica Osten Sacken, 1860 88 · Zootaxa 4122 (1) © 2016 Magnolia Press RIBEIRO & SANTOS (designation: Alexander, 1950). Conorhipidia Alexander, 1914c: 117 (as subgenus of Rhipidia). Type species, Rhipidia conica Alexander, 1914c (original designation). annulicornis (Enderlein), 1912: 80 (Rhipidia). Type locality: Colombia, Hacienda Pehlke. Distr.: Brazil, Colombia (Tolima, Hacienda Pehlke: Enderlein, 1912), Costa Rica, Ecuador, Guyana, Paraguay, Peru, Trinidad, Venezuela. luxuriosa (Alexander), 1929c: 247 (Limonia). Type locality: Colombia, Magdalena, La Cumbre. Distr.: Colombia (Magdalena, La Cumbre). mystica (Alexander), 1931: 627 (Limonia). Type locality: Colombia, Magdalena, Mt, Santa Marta, Vista Nieve. Distr.: Colombia (Magdalena, Vista Nieve, Mt. Santa Marta: Alexander, 1931), Honduras. nigrorostrata muzoensis (Alexander), 1938a: 434 (Limonia). Type locality: Colombia, Boyacá, Muzo. Distr.: Colombia (Boyacá, Muzo: Alexander, 1938a). stonei (Alexander), 1938b: 328 (Limonia). Type locality: Colombia, Usme. Distr.: Colombia (Bogota, Usme: Alexander, 1938b). subpectinata pleuralis (Alexander), 1912c: 12 (Rhipidia). Type locality: Guatemala, Trece Aguas, Cacao, Alta Vera, Paz. Distr.: Colombia, Guatemala, Mexico, Panama. tripectinata (Alexander), 1931: 625 (Limonia). Type locality: Colombia, Magdalena, Mt, Santa Marta, Vista Nieve. Distr.: Colombia (Magdalena, Vista Nieve, Mt. Santa Marta: Alexander, 1931). Genus Toxorhina Loew Toxorhina Loew, 1850: 36. Type species, Toxorhina fragilis Loew, 1851 (designation: Osten Sacken, 1869). Neoceratocheilus Wesche, 1910: 356. Type species, Neoceratocheilus grahami Wesche, 1910 (monotypic). Subgenus Ceratocheilus Wesche Ceratocheilus Wesche, 1910 (as genus). Type species, Ceratocheilus winnsampsoni Wesche, 1910 (designation: Brunetti, 1920) [= cornigera (Speiser, 1908)]. Neostyringomyia Alexander, 1912d: 85 (as subgenus of Styringomyia Loew, 1845). Type species, Styringomyia cornigera Speiser, 1908 (original designation). Conithorax Brunetti, 1918: 298 (as genus). Type species, Conithorax latifrons Brunetti, 1918 (original designation). americana (Alexander), 1913d: 49 (Ceratocheilus). Type locality: Panamá, Culebra. Distr.: Brazil, Colombia, Costa Rica, Guyana, Panama, Peru. leucomelanopus (Enderlein), 1912: 78 (Aporosa). Type locality: Colombia, Hacienda Pelhke. Distr.: Colombia (Tolima, Hacienda Pehlke: Enderlein, 1912). niveitarsis (Alexander), 1922b: 49 (Ceratocheilus). Type locality: Colombia, Boca del Condoto. Distr.: Colombia (Choco, Boca del Condoto: Alexander, 1922b). Subgenus Toxorhina Loew Toxorhina Loew, 1850: 36 (as genus). Type species, Toxorhina fragilis Loew, 1851 (designation: Osten Sacken, 1869). Neoceratocheilus Wesche, 1910: 356 (as genus). Type species, Neoceratocheilus grahami Wesche, 1910 (monotypic). atripes (Alexander), 1922b: 48 (Toxorhina). Type locality: Colombia, Condoto. Distr.: Colombia (Choco, Condoto: Alexander, 1922b). CATALOGUE OF TIPULIDAE AND LIMONIIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 89 TABLE 1. Tipulidae and Limoniidae species recorded from Colombia (according to the Catalogue of Crane flies of the World by Oosterbroek 2015). Family Subfamily Genus Tipulidae Tipulinae Brachypremna Leptotarsus Subgenus Pehlkea Tanypremna Nephrotoma Species Subspecies dispellens (Walker, 1861) columbianus (Enderlein, 1912) regina (Alexander, 1914) elegantior (Alexander, 1945) fuscitarsis (Alexander, 1919) longipes (Fabricius, 1805) alleni (Alexander, 1913) globosa Alexander, 1969 medioligula Alexander, 1945 Ozodicera Dihexaclonus telestyla Alexander, 1969 Tipula Bellardina obliquefasciata Macquart, 1846 Eumicrotipula armillata Alexander, 1916 bogotana Alexander, 1938 carizona Alexander, 1913 foersteriana Alexander, 1967 laterosetosa Alexander, 1931 mecoglossa Alexander, 1967 miranha Alexander, 1913 mitua Alexander, 1916 mocoa Alexander, 1913 moniliferoides Alexander, 1920a moniliformis von Roder, 1886 ornaticornis van der Wulp, 1891 pallidineuris Macquart, 1846 redunca riveti Microtipula Zelandotipula Limoniidae Chioneinae Author Atarba Atarba Ischnothrix Erioptera Erioptera Alexander, 1967 tolimensis Alexander, 1941 colombicola Alexander, 1929 erostrata Alexander, 1945 multimoda Alexander, 1938 perangustula Alexander, 1938 tenuicula Enderlein, 1912 urophora Alexander, 1938 austrofurcifera Alexander, 1969 corynostyla Alexander, 1969 longitarsis (Macquart, 1846) orophila (Alexander, 1914) retrorsa Alexander, 1969 wardiana Alexander, 1981 brunneicornis Alexander, 1916 columbiana Alexander, 1913 lloydi (Alexander, 1913) andina Alexander, 1913 annulipes Williston, 1896 ...Continued on next page 90 · Zootaxa 4122 (1) © 2016 Magnolia Press RIBEIRO & SANTOS TABLE 1. (Continued) Family Subfamily Genus Subgenus Species Author quinquecincta Alexander, 1927 Eugnophomyia luctuosa (Osten Sacken, 1860) Gnophomyia acricula Alexander, 1931 vilis Alexander, 1931 andicola Alexander, 1913 methodica Alexander, 1931 cervaria Alexander, 1938 Gonomyia Gonomyia Leiponeura Molophilus Paralipophleps lemniscata Alexander, 1931 Molophilus capricornis Alexander, 1916 conscriptus Alexander, 1938 flexilistylus Alexander, 1931 lictor Alexander, 1938 luxuriosus Alexander, 1938 marthae Alexander, 1931 perseus Alexander, 1913 subtenebricosus Alexander, 1931 tenebricosus Alexander, 1916 walkeri Alexander, 1931 colombicola Alexander, 1931 Rhabdomastix tantilla Alexander, 1938 Symplecta Symplecta colombiana Teucholabis Paratropesa chalybeia Alexander, 1927 Teucholabis jocosa Alexander, 1913 marticola Alexander, 1931 megaphallus Alexander, 1968 salti Alexander, 1930 spinigera Schiner, 1868 Neognophomyia Rhabdomastix Styringomyia maya Ribeiro, 2003 trifasciata Limnophilinae Subspecies Epiphragma Hexatoma Epiphragma Eriocera colombiana trifasciata (Alexander, 1937) Enderlein, 1912 venezuelensis (Macquart, 1846) cordillerense Alexander, 1913 delicatulum Osten Sacken, 1888 gracilicorne Alexander, 1916 antennata (Alexander, 1930) bequaertiana Alexander, 1938 braconides (Enderlein, 1912) columbiana (Alexander, 1919) ohausiana (Enderlein, 1912) perlaeta (Alexander, 1922) sanctaemartae (Alexander, 1919) schineri (Alexander, 1922) virgulativentris (Enderlein, 1912) Lecteria Lecteria retrorsa Alexander, 1969 Limnophila Limnophila lloydi Alexander, 1913 ...Continued on next page CATALOGUE OF TIPULIDAE AND LIMONIIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 91 TABLE 1. (Continued) Family Subfamily Genus Subgenus Polymera Polymera Shannonomyia Limoniinae Dicranomyia Elephantomyia Species Subspecies Author ominosa Alexander, 1938 sordidipes Alexander, 1938 Shannonomyia bogotensis Alexander, 1938 Dicranomyia lathraea (Alexander, 1926) orophila (Alexander, 1913) elegantula Alexander, 1913 insignifica (Alexander, 1912) longiventris Alexander, 1913 pampoecila Alexander, 1922 quadrituberculata (Alexander, 1937) subandicola (Alexander, 1947) teinoterga (Alexander, 1967) Neoglochina capnora Neolimnobia diva metae (Alexander, 1938) (Schiner, 1868) Peripheroptera nitens (Schiner, 1868) Elephantomyia angustissima Alexander, 1931 humilis Geranomyia eurygramma Alexander, 1931 eurygramma Alexander, 1928 lacteitarsis Alexander, 1922 laudanda (Alexander, 1938) lineata Enderlein, 1912 marthae (Alexander, 1930) neogaudens (Alexander, 1942) neonumenius (Alexander, 1930) plumbeipleura recondita Alexander, 1916 recondita Alexander, 1921 versuta (Alexander, 1930) walkeri (Alexander, 1930) Helius Helius albitarsis Limonia Uncertain ocellata (von Roder, 1886) cordillerensis (Alexander, 1913) sanctaemartae (Alexander, 1930) Neolimonia albitarsis (Osten Sacken, 1888) Orimarga Orimarga andina Alexander, 1916 Rhipidia Rhipidia annulicornis Enderlein, 1912 luxuriosa (Alexander, 1929) mystica (Alexander, 1931) nigrorostrata muzoensis stonei subpectinata Toxorhina Ceratocheilus Toxorhina 92 · Zootaxa 4122 (1) © 2016 Magnolia Press (Alexander, 1938) (Alexander, 1938) subpectinata Alexander, 1912 tripectinata (Alexander, 1931) americana (Alexander, 1913) leucomelanopus (Enderlein, 1912) niveitarsis (Alexander, 1922) atripes Alexander, 1922 RIBEIRO & SANTOS References Agassiz, J.L.R. 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Transactions of the Royal Entomological Society of London, 1896, 253–446. http://dx.doi.org/10.1111/j.1365-2311.1896.tb00965.x Wulp, F.M. van der (1891) Eenige uitlandsche Diptera. Tijdschrift voor Entomologie, 34, 193–218. Zetterstedt, J.W. (1851) Diptera Scandinaviae, disposita et descripta. Vol 10. Officina Lundbergiana, Lundae [= Lund], 3711–4090. CATALOGUE OF TIPULIDAE AND LIMONIIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 97 Zootaxa 4122 (1): 098–141 http://www.mapress.com/j/zt/ Copyright © 2016 Magnolia Press Article ISSN 1175-5326 (print edition) ZOOTAXA ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.4122.1.15 http://zoobank.org/urn:lsid:zoobank.org:pub:F75443F4-82D3-4B00-B21A-71A13050D0E3 FAMILY CERATOPOGONIDAE GUSTAVO SPINELLI1,3 & MARTA WOLFF2 1 División Entomología, Museo de La Plata, Paseo del Bosque s/n, 1900 La Plata, Argentina. E-mail: spinelli@fcnym.unlp.edu.ar Grupo de Entomologia, Instituto de Biologia, Universidad de Antioquia, Medellín-Colombia. E-mail: martha.wolff@udea.edu.co 3 Corresponding author: E-mail: spinelli@fcnym.unlp.edu.ar 2 Abstract Two hundred and thirty five species of Ceratopogonids arranged in 28 genera are listed for Colombia, mostly Culicoides (112 species) and Forcipomyia (50 species). For each listed species, information on type locality, type category and sex, location of type material, species distribution, localities for Colombia and main references, is provided. Localities are not specified for 17 species in which the literature searching was negative. Forcipomyia squamitibia Lutz and Bezzia clavipennis are excluded from the list. Key words: catalogue, Ceratopogonidae, Colombia, distribution, localities, species list Introduction The Ceratopogonidae are commonly known as biting midges, no-see-ums or punkies, in Spanish as manta blanca, polvorines, or jejenes (which also may refer to Simuliidae), and in Portuguese as maruim or mosquito pólvora. Adult females of Leptoconops Skuse Culicoides Latreille and Forcipomyia Meigen (only in the subgenus Lasiohelea Kieffer) are haematophagous on vertebrates, needing the blood to develop their eggs. They are nasty biters that pester humans and domestic animals and, in some instances, transmit harmful diseases (Borkent & Spinelli 2007). On the other hand, this group of flies provides important services in a wide array of ecosystems. Some species are important pollinators of such plants as cacao and rubber trees, and the larvae of many are important predators of other organisms in semiaquatic and aquatic habitats. The adults of most biting midges actually suck blood from other insects and may even be important vectors of viruses that kill caterpillars (Borkent et al. 2009). Borkent and Spinelli (2007), in their catalog of the Neotropical ceratopogonids, listed 1093 species and 132 new species were subsequently described. From the 1225 species that inhabit the Neotropics, 235 arranged in 28 genera are reported from Colombia, mostly Culicoides (112 species) and Forcipomyia (50 species) (Table 1). We are presenting here the updated catalog of the Ceratopogonidae of Colombia (Table 1). Under the heading for each generic name, citations for keys to species or the most recent revision of the species in that group are provided. In some instances these are only for a species group or a limited geographical area. When a reference is not given, the most recent references are those of the genus and/or species themselves. For each listed species, the following information is included: type locality, type category and sex, location of type material, species distribution, localities for Colombia (between brackets) and main references. The latter include the original description, those mentioning Colombian localities, lists and catalogs. The literature searching for localities of 17 species reported for Colombia was negative, and for these species the first report for the country is included. Specific epithets of synonyms are accompanied by type locality, type category and sex, and location of type material. Two species were erroneously recorded for Colombia, Forcipomyia squamitibia Lutz by Marino & Spinelli (2002) and Bezzia clavipennis in the original description by Spinelli & Wirth (1989b), and they are excluded from the list. 98 Accepted by S. Nihei: 25 May 2015; published: 14 Jun. 2016 Acronyms used for the depositories BMNH—Department of Entomology, Natural History Museum, London, SW7 5BD, United Kingdom. CAIM—Colección de Insectos con Importancia Médica. Laboratorio de Entomología, InDRE, Francisco de P. Miranda No. 177, Col. Unidad Lomas Plateros, México D.F. 11340, Mexico. CASC—Department of Entomology, California Academy of Sciences, Golden Gate Park, San Francisco, California, 94118, USA. CNCI—Canadian National Collection of Insects, Eastern Cereal and Oilseed Research Centre, Agriculture 8 Canada, K.W. Neatby Building, Ottawa, Ontario, K1A 0C6, Canada. CUIC—Department of Entomology, Comstock Hall, Cornell University, Ithaca, New York, 14853 –0999, USA. DEFS—Departamento de Epidemiologia, Faculdade de Saúde Pública, Universidade de São Paulo, Av. Dr. Arnaldo, 715, 01.255, São Paulo, SP, Brazil. FIOC—Instituto Oswaldo Cruz, Departamento de Entomologia, Caixa Postal 926, 20010 Rio de Janeiro, Rio de Janeiro, Brazil. FSCA—Florida State Collection of Arthropods, Florida Department of Agriculture and Consumer Services Division of Plant Industry, 1911 SW 34th Street, Gainesville, Florida, 32608-1268, USA. HNHM—Zoological Dept., Hungarian Natural History Museum, Baross utca 13, H –1088, Budapest, Hungary. ICNC—Universidad Nacional de Colombia. Carrera 30#45-03. Código postal 111321. Instituto de Ciencias Naturales. Edificio 425. INHC—Instituto Nacional de Higiene, Caracas, Venezuela. INMA—Departamento de Artropodología Sanitaria, Instituto Nacional de Microbiología “Dr. Carlos G. Malbrán”, Ciudad de Buenos Aires, Argentina. MCZC—Entomology Department, Museum of Comparative Zoology, Harvard University, 26 Oxford St., Cambridge, Massachusetts, 02138, USA. MHNL—Museo de Historia Natural La Salle, Caracas, Venezuela. MLPA—Museo de la Plata, Division Entomologia, Universidad de La Palata, Paseo del Bosque, 1900 La Plata, Argentina. MNHN—Museum National d'Histoire Naturelle, 45, rue Buffon, Paris, 75005, France. MUNS—Museo Nacional, Santiago, Chile. UNCE—Universidad Central del Ecuador. Cuidadela Universitaria, Quito, Ecuador. UNPR—School of Tropical Medicine, University of Puerto Rico, San Juan, Puerto Rico. USNM—United States National Entomological Collection, Dept. of Entomology, U.S. National Museum of Natural History, Washington, DC 20560, USA. ZMHB—Museum für Naturkunde der Humboldt Universität zu Berlin, Invalidenstrasse 43, Berlin 1040, Germany. List of abbreviations cat.—catalogue comm.—comments distr.—Geographical Distribution m—meters km—kilometers mon.—monotypy orig. des.—original designation preocc.—preoccupied refs.—references rev.—revision Rt.—route sub. des.—subsequent designation F (f#)—Female HT—Holotype CATALOGUE OF CERATOPOGONIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 99 LT—Lectotype M (m#)—Male NW—northwest ST—Syntype T—Type Catalogue of Ceratopogonidae of Colombia (The names of those species of Leptoconops and Culicoides which are known to bite humans are underlined). Family Ceratopogonidae Newman, 1834 Subfamily Leptoconopinae Noè, 1907 Genus Leptoconops Skuse Leptoconops Skuse, 1889: 288. Type species, Leptoconops stygius Skuse (mon.). Refs.: Ronderos, 1988 (key to females from Argentina); Ronderos & Spinelli, 1992 (key to females); Wirth & Atchley, 1973 (key to subgenera of females, other than Proleptoconops; key to Nearctic females). Tersesthes Townsend, 1893: 370. Type species, Tersesthes torrens Townsend (orig. des.). Mycterotypus Noè, 1905: 114. Type species, Mycterotypus bezzii Noè, sub. des. by Carter, 1921:3. Protersesthes Kieffer, 1921b: 107. Type species, Tersesthes brasiliensis Lutz (orig. des.). Subgenus Megaconops Wirth & Atchley Megaconops Wirth & Atchley, 1973: 18 (as subgenus of Leptoconops). Type species, Leptoconops floridensis Wirth (orig. des.). floridensis Wirth, 1951a: 282. Type locality: USA, Florida, Escambia Co., Santa Rosa Island. T F (UNFL). Distr.: USA (Florida), Jamaica, Colombia (Valle del Cauca (Bahia Buenaventura, “Bella Vista”)). Refs.: Wirth, 1951a: 282; Wirth & Atchley, 1973: 18; Borkent & Spinelli, 2007: 44. Subfamily Forcipomyiinae Lenz, 1934 Genus Atrichopogon Kieffer Atrichopogon Kieffer, 1906a: 53 (as subgenus of Ceratopogon). Type species, Ceratopogon exilis Coquillett (= Ceratopogon levis Coquillett), Coquillett, 1910: 512 (des.). Refs.: Huerta, 1996 (key to males and females from Mexico); Ingram & Macfie, 1931 (key males and females from Patagonia and southern Chile); Macfie, 1939b (key to males and females from the Neotropical Region). Subgenus Atrichopogon Kieffer auricoma Kieffer, 1917b: 300. Type locality: Colombia, Magdalena, Aracataca. T F (HNHM, lost). Distr.: Colombia (Magdalena (Aracataca)). Refs.: Kieffer 1917b: 300; Borkent & Spinelli, 2007: 44; Borkent, 2015: 18 (cat.). columbianus Kieffer, 1917b: 300). Type locality: Colombia, Magdalena, Aracataca. T F (HNHM, lost). Distr.: Colombia (Magdalena (Aracataca)). Refs.: Kieffer 1917b: 300; Borkent & Spinelli, 2007: 45; Borkent, 2015: 20 (cat.). 100 · Zootaxa 4122 (1) © 2016 Magnolia Press SPINELLI & WOLFF Genus Forcipomyia Meigen Forcipomyia Meigen, 1818: 73, 75. Type species, Tipula bipunctata Linnaeus, Westwood, 1840: 126 (des.). Generic name first published in synonymy with Ceratopogon but available under I.C.Z.N Code Article 11(e). Refs.: Debenham, 1987 (key to males and females to subgenera from Australasian Region which includes all Neotropical subgenera other than Rhynchoforcipomyia and Saliohelea); Ingram & Macfie, 1931 (key to males and females from Patagonia and southern Chile); Wirth & Ratanaworabhan, 1978 (key to females to subgenera; their genus A near Lepidohelea = Pedilohelea). Tetraphora Philippi, 1865: 630. Type species, Tetraphora fusca Philippi (mon.). Subgenus Caloforcipomyia Saunders Caloforcipomyia Saunders, 1957: 680 (as subgenus of Forcipomyia). Type species, Forcipomyia caerulea Saunders (orig. des.). Refs.: Utmar & Wirth, 1976 (key to New World males and females). caerulea Saunders, 1957: 681. Type locality: Brazil, Rio de Janeiro. HT F (# 6504 CNCI). Distr.: Panama, Colombia (Antioquia (Río Anorí), Valle del Cauca (Río Raposo)), Dominica, Brazil (Pará, Rio de Janeiro). Refs.: Saunders, 1957: 681; Utmar & Wirth, 1976: 115; Borkent & Spinelli, 2007: 48. eukosma Macfie, 1939b: 148. Type locality: Brazil, Santa Catarina, Nova Teutonia. LT M (BMNH). Distr.: Colombia (Valle del Cauca (Río Raposo)), Dominica, Brazil (Pará, São Paulo, Río de Janeiro, Santa Catarina). Refs.: Macfie, 1939b: 148; Utmar & Wirth, 1976: 119; Borkent & Spinelli, 2007: 48. glauca Macfie, 1934: 144. Type locality: Great Britain, Norfolk, Hickling Broad. HT M (BMNH). Distr.: Europe, North America (Alberta to Nova Scotia to Arkansas and Florida) to Colombia (?) and southern Brazil. Refs.: Macfie, 1934: 144; Utmar & Wirth, 1976: 123; Borkent & Spinelli, 2007: 48. hatoensis Utmar & Wirth, 1976: 131. Type locality: Panama, Chiriqui, El Hato. HT M (# 72216 USNM). Distr.: Mexico (Sinaloa), Costa Rica, Panama, Colombia (Antioquia (Río Anorí; Río Truando), Caquetá (Tres Esquinas)). Refs.: Utmar & Wirth, 1976: 131; Borkent & Spinelli, 2007: 48. hermosa Utmar & Wirth, 1976: 117. Type locality: Brazil, Pará, Belém. HT M (DEFS). Distr.: Costa Rica, Panama, Colombia (Antioquia (Río Anorí), Valle del Cauca (Río Raposo)), Jamaica, Panama, Brazil (Amazonas, Pará). Refs.: Utmar & Wirth, 1976: 117; Borkent & Spinelli, 2007: 48. remigera Utmar & Wirth, 1976: 121. Type locality: Colombia, Valle del Cauca (Río Raposo). HT M (# 72212 USNM). Distr.: Panama, Colombia (Valle del Cauca (Río Raposo)), French Guiana, Brazil (Pará). Refs.: Utmar & Wirth, 1976: 121; Borkent & Spinelli, 2007: 48; Borkent, 2015: 30 (cat.). sabalitensis Utmar & Wirth, 1976: 127. Type locality: Costa Rica, Puntarenas, Sabalito. HT M (# 72215 USNM). Distr.: Honduras to Colombia (Antioquia (Camp Teresita, Río Truando)), Brazil (Paráná, Santa Catarina), northeastern Argentina. Refs.: Utmar & Wirth, 1976: 127; Borkent & Spinelli, 2007: 48. Subgenus Euprojoannisia Brèthes Euprojoannisia Brèthes, 1914: 155. Type species, Euprojoannisia platensis Brèthes (orig. des.). Refs.: Bystrak & Wirth, 1978 (key to Nearctic larvae, pupae, males, females); Huerta, 1996 (key to males and females from Mexico). blantoni Soria & Bystrak, 1975: 3. Type locality: Brazil, Bahia, Ilhéus (CEPEC). HT M (DEFS). Distr.: Eastern USA to Colombia (Valle del Cauca (Palmira)), Ecuador, and Brazil (Bahia). Refs.: Soria & Bystrak, 1975: 3; Soria & Wirth, 1979: 79; Borkent & Spinelli, 2007: 49. Subgenus Forcipomyia Meigen Refs.: Huerta, 1996 (key to females from Mexico); Marino & Spinelli, 1999 (key to species groups); Marino & Spinelli, 2001 CATALOGUE OF CERATOPOGONIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 101 (key to males and females from Patagonia); Marino & Spinelli, 2002 (key to squamitibia group); Wirth, 1982a (key to argenteola group). argenteola Species Group argenteola Macfie, 1939b: 146. Type locality: Brazil, Santa Catarina, Nova Teutonia. ST F (BMNH). Distr.: Colombia (Valle del Cauca (Palmira)); Brazil (Rio de Janeiro, Santa Catarina), Paraguay, Argentina (Misiones). Refs.: Macfie, 1939b: 146; Soria & Wirth, 1979: 79; Borkent & Spinelli, 2007: 49. calatheae Wirth, 1982a: 573. Type locality: Dominica, Calabishie, swamp at mouth of Hodges river. HT F (# 76582 USNM). Distr.: Colombia (Antioquia (San Jeronimo)), Dominica, Brazil (Bahia, Santa Catarina), northern Argentina. Refs.: Wirth, 1982a: 573; Borkent & Spinelli, 2007: 49. genualis Species Group genualis (Loew), 1866: 128 (Ceratopogon). Type locality: Cuba. HT F (MCZC). Distr.: USA (Louisiana to Florida), Bahamas, Mexico (Yucatán, Chiapas) to Colombia. Locality in Colombia (Valle del Cauca (Palmira)), Galápagos Islands, Cayman Islands, Cuba to Argentina (Buenos Aires province); São Tomé. Refs.: Loew, 1866: 128; Wirth & Soria, 1975: 22; Soria & Wirth, 1979: 79; Borkent & Spinelli, 2007: 50. propinqua (Williston), 1896: 279 (Ceratopogon). Type locality: St. Vincent. HT M (BMNH). raleighi Macfie, 1938: 160. Type locality: Trinidad and Tobago, St. Augustine. LT M (BMNH). harpegonata Wirth & Soria, 1975: 19. Type locality: Puerto Rico, Mayaguez, Penn Farm. HT M (# 69493 USNM). Distr.: Costa Rica, Bahamas, Jamaica, Puerto Rico, Colombia (Chocó (Río Truando, Camp Teresita); Valle del Cauca (Río Raposo; Palmira)), Trinidad and Tobago to Brazil (Amazonas, Pará, Bahia). Refs.: Wirth & Soria, 1975: 19; Soria & Wirth, 1979: 79; Borkent & Spinelli, 2007: 50. squamitibia Species Group catarinensis Marino & Spinelli, 2002: 309. Type locality: Brazil, Santa Catarina, Nova Teutonia. HT M (MLPA). Distr.: Mexico, Costa Rica, Colombia (Antioquia (Tamesis)), Brazil. Refs.: Marino & Spinelli, 2002: 309; Borkent & Spinelli, 2007: 51. Subgenus Lasiohelea Kieffer Centrorhynchus Lutz, 1913: 62 (preocc. Centrorhynchus Steven or Fisher Waldheim, 1829). Type species, Centrorhynchus stylifer Lutz (orig. des.). Lasiohelea Kieffer, 1921b: 115. Type species, Atrichopogon pilosipennis Kieffer (= Ceratopogon velox Winnertz) (orig. des.). Refs.: Aitken et al., 1975 (species in Trinidad and Tobago); Ronderos & Spinelli, 1999 (rev. of Neotropical species). stylifer (Lutz), 1913: 63 (Centrorhynchus). Type locality: Brazil, São Paulo, Tatuí. LT F (# 3699 FIOC). Distr.: Belize to Colombia (Valle del Cauca (Palmira)), Ecuador, Trinidad and Tobago, Venezuela to Northeastern Argentina. Refs.: Lutz, 1913: 63; Soria & Wirth, 1979: 79; Borkent & Spinelli, 2007: 52. Subgenus Lepidohelea Kieffer Lepidohelea Kieffer, 1917b: 364. Type species, Ceratopogon chrysolophus Kieffer (orig. des.). Refs.: Wirth, 1991c (key to species groups; key to males and females of bicolor group); Wirth & Spinelli, 1992 (males and females of seminole subgroup); Wirth & Spinelli, 1993 (key to males and some females of annulatipes group). annulatipes Macfie, 1939b: 154. Type locality: Brazil, Santa Catarina, Nova Teutonia. HT M (BMNH). Distr.: 102 · Zootaxa 4122 (1) © 2016 Magnolia Press SPINELLI & WOLFF Colombia (Antioquia (near Río Anorí), Valle del Cauca (Río Raposo)), Brazil (Pará, Bahia, Santa Catarina). Refs.: Macfie, 1939b: 154; Wirth & Spinelli, 1993: 111; Borkent & Spinelli, 2007: 52. convexipenis Wirth & Spinelli, 1993: 116. Type locality: Colombia, Valle del Cauca (Río Raposo). HT M (USNM). Distr.: Colombia (Antioquia (Turbo, Río León), Valle del Cauca (Río Raposo)); Refs.: Wirth & Spinelli, 1993: 116; Borkent & Spinelli, 2007: 52; Borkent, 2015: 47 (cat.). euthystyla Wirth & Spinelli, 1993: 117. Type locality: Colombia, Valle del Cauca (Río Raposo). HT M (USNM). Distr.: Panama, Colombia (Antioquia (Turbo, Río León; near Río Anorí), (Valle del Cauca (Río Raposo)). Refs.: Wirth & Spinelli, 1993: 117; Borkent & Spinelli, 2007: 52; Borkent, 2015: 47 (cat.). hobbsi Wirth & Spinelli, 1993: 121. Type locality: Dominica, 2 km W Pont Casse. HT M (USNM). Distr.: Mexico (Tabasco) to Colombia (Meta (Finca Barbascal)), Jamaica to Grenada. Refs.: Wirth & Spinelli, 1993: 121; Borkent & Spinelli, 2007: 53. luteigenua Wirth & Spinelli, 1992: 353. Type locality: Costa Rica, Limón, Siquirres, Hacienda Theobroma. HT F (USNM). Distr.: USA (Florida) to Colombia (Antioquia (Tamesis)) and Brazil (Bahia). Refs.: Wirth & Spinelli, 1992: 353; Borkent & Spinelli, 2007: 53. Subgenus Microhelea Kieffer Microhelea Kieffer, 1917b: 364. Type species, Atrichopogon microtomus Kieffer, sub. des. by Kieffer, 1921a: 7, with explanatory notation on erroneous previous type designation. Refs.: Clastrier & Wirth, 1995 (key to females in ixodoides group); Huerta, 1996 (key to males and females from Mexico); Wirth, 1972 (key to females in fuliginosa group). Phasmidohelea Mayer, 1937: 233. Type species, Phasmidohelea crudelis Mayer (= Ceratopogon fuliginosus Meigen) (orig. des.). antioquiae Clastrier & Wirth, 1995: 142. Type locality: Colombia, Antioquia (near Río Anorí). HT F (USNM). Distr.: Colombia (Antioquia (near Río Anorí)). Refs.: Clastrier & Wirth, 1995: 142; Borkent & Spinelli, 2007: 54; Borkent, 2015: 49 (cat.). belemensis Clastrier & Wirth, 1995: 140. Type locality: Brazil, Pará, Belém, APEG Forest. HT F (FIOC). Distr.: Colombia (Valle del Cauca (Río Raposo; Oleoducto de Pacific, 10 km NW Dagua)), Brazil (Pará). Refs.: Wirth, 1971: 239 (record of F. obesa from Colombia); Clastrier & Wirth, 1995: 140; Borkent & Spinelli, 2007: 54. colombiana Clastrier & Wirth, 1995: 102. Type locality: Colombia, Valle del Cauca (Río Raposo). HT F (USNM). Distr.: Costa Rica, Colombia (Antioquia (near Río Anorí), Valle del Cauca (Río Raposo)). Refs.: Clastrier & Wirth, 1995: 102; Borkent & Spinelli, 2007: 54; Borkent, 2015: 50 (cat.). fuliginosa (Meigen), 1818: 86 (Ceratopogon). Type locality: Germany, T?. Distr.: Widespread in the Palaearctic, Oriental, Afrotropical, Nearctic and Neotropical regions. In the Neotropics south to Argentina and Uruguay. Locality in Colombia (Valle del Cauca (Palmira)). Refs.: Meigen, 1818: 86; Soria & Wirth, 1979: 79; Borkent & Spinelli, 2007: 54. crudelis Knab, 1914: 66 (preocc. by Forcipomyia crudelis (Karsch), 1886). Type locality: Mexico, La Oaxagena, near Santa Lucrecia. HT F (# 18420 USNM). tropica (Kieffer), 1917b: 297 (Ceratopogon). Type locality: Costa Rica, San Mateo, Higueto. T F (HNHM, lost). insignipalpis Macfie, 1949: 109. Type locality: Mexico, Chiapas, San Cristobal, north of Aurora. HT F (BMNH). Distr.: Mexico (Chiapas), Costa Rica, Panama, Colombia (Valle del Cauca (Oleoducto de Pacífic, 10 km NW. Dagua), Caquetá (Tres Esquinas)), Brazil (Pará). Refs.: Macfie, 1949: 109; Wirth, 1972: 573; Borkent & Spinelli, 2007: 54. CATALOGUE OF CERATOPOGONIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 103 penultimata Wirth, 1972: 573. Type locality: Brazil, Santa Catarina, Nova Teutonia. HT F (# 71148 USNM). Distr.: Belize and Jamaica to Colombia (Valle del Cauca (Río Raposo)), Brazil (Santa Catarina). Refs.: Wirth, 1972: 573; Borkent & Spinelli, 2007: 55. raposoensis Clastrier & Wirth, 1995: 144. Type locality: Colombia, Valle del Cauca (Río Raposo). HT F (USNM). Distr.: Colombia (Valle del Cauca (Río Raposo)). Refs.: Clastrier & Wirth, 1995: 144; Borkent & Spinelli, 2007: 55; Borkent, 2015: 51 (cat.). squamosa Lutz, 1914: 87. Type locality: Brazil, Río de Janaeiro, Manguinhos. ST M (FIOC). Distr.: Colombia (Valle del Cauca (Río Raposo; Oleoducto de Pacífic, 10 km NW Dagua)), Bolivia, Brazil (Amazonas, Bahia, Rio de Janeiro, Santa Catarina), Argentina (Mendoza). Refs.: Lutz, 1914: 87; Wirth, 1972: 569; Borkent & Spinelli, 2007: 55. valleensis Clastrier & Wirth, 1995: 145. Type locality: Colombia, Valle del Cauca (Río Raposo). HT F (USNM). Distr.: Colombia (Valle del Cauca (Río Raposo)). Refs.: Clastrier & Wirth, 1995: 145; Borkent & Spinelli, 2007: 55; Borkent, 2015: 51 (cat.). vernoni Clastrier & Wirth, 1995: 138. Type locality: Colombia, Valle del Cauca (Río Raposo). HT F (USNM). Distr.: Colombia (Valle del Cauca (Río Raposo)). Refs.: Clastrier & Wirth, 1995: 138; Borkent & Spinelli, 2007: 55; Borkent, 2015: 52 (cat.). willistoni Wirth, 1971: 242. Type locality: Puerto Rico, El Verde, Luquillo Mts. HT F (# 71146 USNM). Distr.: Jamaica, Puerto Rico, Colombia (Cauca, Tambo, Finca Carpenteria), Brazil (Santa Catarina). Refs.: Wirth, 1971: 242; Clastrier & Wirth, 1995: 134; Borkent & Spinelli, 2007: 55. Subgenus Pedilohelea de Meillon & Wirth Pedilohelea de Meillon & Wirth, 1980: 9 (as subgenus of Forcipomyia). Type species, Forcipomyia clastrieri Dessart (orig. des.). Refs.: de Meillon & Wirth, 1980 (key to males, some females). aitkeni de Meillon & Wirth, 1980: 12. Type locality: Brazil, Pará, Belém. HT M (# 67551 USNM). Distr.: Colombia (Valle del Cauca (Río Raposo)), Brazil (Pará). Refs.: de Meillon & Wirth, 1980: 12; Borkent & Spinelli, 2007: 55. raposoi de Meillon & Wirth, 1980: 21. Type locality: Brazil, Pará, Belém. HT M (# 67237 USNM). Distr.: Colombia (Valle del Cauca (Río Raposo)), Brazil (Pará). Refs.: de Meillon & Wirth, 1980: 21; Borkent & Spinelli, 2007: 55; Borkent, 2015: 53 (cat.). spangleri de Meillon & Wirth, 1980: 21. Type locality: Guatemala, Suchitepequez, San Antonio Suchitepequez. HT M (# 65711 USNM). Distr.: Mexico (Yucatán), Guatemala, Panama, Colombia (Valle del Cauca (Río Raposo)), Ecuador. Refs.: de Meillon & Wirth, 1980: 21; Borkent & Spinelli, 2007: 55. spilmani de Meillon & Wirth, 1980: 22. Type locality: Dominica, Clarke Hall. HT M (# 73565 USNM). Distr.: Panama, Colombia (Valle del Cauca (Río Raposo)), Dominica, Trinidad and Tobago, Brazil (Santa Catarina). Refs.: de Meillon & Wirth, 1980: 22; Borkent & Spinelli, 2007: 55. Subgenus Pterobosca Macfie Pterobosca Macfie, 1932a: 266. Unavailable name; proposed after 1930 without type species designation. Pterobosca Macfie, 1940d: 16. Type species, Ceratopogon aeschnosuga de Meijere (orig. des.). incubans (Macfie), 1937: 111 (Pterobosca). Type locality: Belize, Cayo District, Mtn. Pine Ridge, on Argia ulmeca. HT F (BMNH). Distr.: Mexico (San Luis Potosí, Veracruz), Puerto Rico, Colombia (Risaralda (La Virginia en Pantano)) to Argentina (Buenos Aires). Refs.: Macfie, 1937: 111; Wirth, 1956: 363; Borkent & Spinelli, 2007: 56. 104 · Zootaxa 4122 (1) © 2016 Magnolia Press SPINELLI & WOLFF Subgenus Rhynchoforcipomyia Wirth & Dow Rhynchoforcipomyia Wirth & Dow, 1972: 863 (as subgenus of Forcipomyia). Type species, Forcipomyia messersmithi Wirth & Dow (orig. des.). Refs.: Wirth & Dow, 1972 (key to females). brachyrhyncha Wirth & Dow, 1972: 867. Type locality: Colombia, Valle del Cauca (Río Raposo). HT F (# 71168 USNM). Distr.: Colombia (Valle del Cauca (Río Raposo)), Trinidad and Tobago. Refs.: Wirth & Dow, 1972: 867; Borkent & Spinelli, 2007: 56; Borkent, 2015: 54 (cat.). messersmithi Wirth & Dow, 1972: 864. Type locality: El Salvador, Cuscatlán, San Pedro Perulapán. HT F (# 71167 USNM). Distr.: Mexico (Veracruz), Jamaica, Central America to Colombia (Valle del Cauca (Río Raposo)). Refs.: Wirth & Dow, 1972: 864; Borkent & Spinelli, 2007: 57. puracensis Wirth & Dow, 1972: 867. Type locality: Colombia, Cauca, Parámo de Purace, 28 km S Purace. HT F (# 71165 USNM). Distr.: Colombia (Cauca (Parámo de Purace, 28 km S Purace). Refs.: Wirth & Dow, 1972: 867; Borkent & Spinelli, 2007: 57; Borkent, 2015: 54 (cat.). zeteki Wirth & Dow, 1972: 870. Type locality, Panama, Canal Zone, Barro Colorado I. HT F (# 71166 USNM). Distr.: Panama, Colombia (Valle del Cauca (Río Raposo)). Refs.: Wirth & Dow, 1972: 870; Borkent & Spinelli, 2007: 57. Subgenus Saliohelea Wirth & Ratanaworabhan Saliohelea Wirth & Ratanaworabhan, 1978: 494 (as subgenus of Forcipomyia). Type species, Forcipomyia leei Wirth & Ratanaworabhan (orig. des.). leei Wirth & Ratanaworabhan, 1978: 498. Type locality: Colombia (Valle del Cauca (Río Raposo)). HT F (# 70440 USNM). Distr.: Eastern USA and Caribbean to Colombia (Antioquia (Río Anorí), Valle del Cauca (Río Raposo)) and southern Brazil (Amazonas, Pará, Bahia, Rio de Janeiro). Refs.: Wirth & Ratanaworabhan, 1978: 498; Borkent & Spinelli, 2007: 57; Borkent, 2015: 55 (cat.). Subgenus Thyridomyia Saunders Thyridomyia Saunders, 1925: 268. Type species, Thyridomyia palustris Saunders (= Ceratopogon monilicornis Coquillett) (orig. des.). Refs.: Wirth, 1970 (key to males and females). colombiae Wirth, 1970: 435. Type locality: Colombia, Valle del Cauca (Río Raposo). HT M (# 70662 USNM). Distr.: Mexico (Veracruz), Costa Rica, Panama, Trinidad and Tobago, Colombia (Valle del Cauca (Río Raposo)). Refs.: Wirth, 1970: 435; Borkent & Spinelli, 2007: 58; Borkent, 2015: 56 (cat.). jipajapae Wirth, 1970: 437. Type locality: Panama, Canal Zone, Gamboa, rain forest on “Pipeline road”. HT M (# 70661 USNM). Distr.: Costa Rica to Colombia (Valle del Cauca (Río Raposo)) and Ecuador, Trinidad and Tobago. Refs.: Wirth, 1970: 437; Borkent & Spinelli, 2007: 58. nana (Macfie), 1939b: 171 (Lasiohelea). Type locality: Brazil, Santa Catarina, Nova Teutonia. HT F (BMNH). Distr.: Mexico (Veracruz), Jamaica through Caribbean to Colombia (Valle del Cauca (Río Raposo)) and southern Brazil (Santa Catarina). Refs.: Macfie, 1939b: 171; Wirth, 1970: 432; Borkent & Spinelli, 2007: 58. nodosa Saunders, 1959: 43. Type locality: Costa Rica, Finca Theobroma, near Siquirres. HL (CNCI). Distr.: USA (Arizona, Kansas, Mississippi, Florida), Mexico (Baja California, Sonora, Sinaloa, San Luis Potosí), Costa Rica, Colombia (Valle del Cauca (Río Raposo)). Refs.: Saunders, 1959: 43; Wirth, 1970: 434; Borkent & Spinelli, 2007: 58. CATALOGUE OF CERATOPOGONIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 105 Subgenus Warmkea Saunders Warmkea Saunders, 1957: 671 (as subgenus of Forcipomyia). Type species, Forcipomyia bicolor Saunders (= Forcipomyia lesliei Wirth) (orig. des.). Refs.: Wirth & Soria, 1980 (key to males and females). aeria Saunders, 1957: 675. Type locality: Puerto Rico, Mayaguez. HT F (CNCI). Distr.: Puerto Rico, Colombia (Antioquia (Río Anorí), Valle del Cauca (Río Raposo)), Argentina (Tucumán). Refs.: Saunders, 1957: 675; Wirth & Soria, 1980: 143; Borkent & Spinelli, 2007: 59. galindoi Wirth & Soria, 1980: 146. Type locality: Panama, Canal Zone, Barro Colorado island. HT M (#76102 USNM). Distr.: Panama, Colombia (Antioquia (Río Anorí), Valle del Cauca (Río Raposo)), Ecuador, Brazil (Pará). Refs.: Wirth & Soria, 1980: 146; Borkent & Spinelli, 2007: 59. lesliei Wirth, 1974: 12. New name for bicolor. Distr.: Jamaica, Haiti to Venezuela, Colombia (Valle del Cauca (Río Raposo; Oleoducto de Pacifico, 10 km NW Dagua)). Refs.: Saunders, 1957: 672; Wirth & Soria, 1980: 148; Borkent & Spinelli, 2007: 59. bicolor Saunders, 1957: 672 (preocc. Forcipomyia bicolor Lutz, 1914). Type locality: Puerto Rico, Mayaguez. HT F (CNCI). louriei (Macfie), 1935a: 49 (Lasiohelea). Type locality: Brazil, Piauí, Tutóia. HT F (BMNH). Distr.: Colombia (Antioquia (Río Anorí), Valle del Cauca (Río Raposo)), Bolivia, Trinidad and Tobago, Brazil (Pará, Maranhão, Santa Catarina). Refs.: Macfie, 1935: 49; Wirth & Soria, 1980: 151; Borkent & Spinelli, 2007: 59. narthekophora Macfie, 1939b: 166. Type locality: Brazil, Santa Catarina, Nova Teutonia. HT M (BMNH). spinosa Saunders, 1957: 676. Type locality: Puerto Rico, Mayaguez. HT F (CNCI). Distr.: Puerto Rico, Dominica, Trinidad and Tobago to Colombia (Valle del Cauca (Río Raposo)), Guyana and Brazil (Pará, Bahia, Rio de Janeiro). Refs.: Saunders, 1957: 676; Wirth & Soria, 1980: 153; Borkent & Spinelli, 2007: 59. tuberculata Saunders, 1957: 677. Type locality: Trinidad and Tobago, North Range. HT F (CNCI). Distr.: Costa Rica, Colombia (Antioquia (Río Anorí), Valle del Cauca (Río Raposo)), Trinidad and Tobago and Tobago, Brazil (Amazonas, Pará), Argentina (Buenos Aires). Refs.: Saunders, 1957: 677; Wirth & Soria, 1980: 157; Borkent & Spinelli, 2007: 59. Subfamily Dasyheleinae Lenz, 1934 Genus Dasyhelea Kieffer Dasyhelea Kieffer, 1911c: 5. Type species, Dasyhelea halophila Kieffer (mon.). Refs.: Huerta, 1996 (key to males and females from Mexico); Ingram & Macfie, 1931 (key males and females from Patagonia and southern Chile); Spinelli & Wirth, 1984c (key to males and females of cincta group); Díaz et al., 2010 (rev. of patagonica group); Díaz et al., 2014 (phylogeny of mutabilis group in the Neotropics). Pseudoculicoides Malloch, 1915a: 309. Type species, Ceratopogon mutabilis Coquillett, (orig. des.). columbiana (Kieffer), 1917b: 304 (Culicoides). Type locality: Colombia, Magdalena, Aracataca. T F (HNHM, lost). Distr.: Colombia (Magdalena (Aracataca)). Refs.: Kieffer 1917b: 304; Borkent & Spinelli, 2007: 60; Borkent, 2015: 63 (cat.). santaemarthae (Kieffer), 1917b: 306 (Culicoides). Type locality: Colombia, Magdalena, Santa Marta. T F (HNHM, lost). Distr.: Colombia (Magdalena (Santa Marta)). Refs.: Kieffer 1917b: 306; Borkent & Spinelli, 2007: 61; Borkent, 2015: 71 (cat.). 106 · Zootaxa 4122 (1) © 2016 Magnolia Press SPINELLI & WOLFF Subfamily Ceratopogoninae Newman, 1834 Tribe Culicoidini Kieffer, 1911 Genus Culicoides Latreille Culicoides Latreille, 1809: 251. Type species, Culicoides punctatus Latreille (= Ceratopogon punctatus Meigen) (mon). Refs.: Aitken et al., 1975 (key to females from Trinidad and Tobago); Blanton & Wirth, 1979 (key to larvae, males and females from Florida (USA)); Forattini, 1957 (key to females of all species); Huerta, 1996 (key to females from Mexico); Ortíz & León, 1955 (key to females from Ecuador); Ronderos, 1988 (key to females from Argentina); Ronderos & Spinelli, 1998 (key to males and females from the Yacyretá Dam, between Paraguay and Argentina); Spinelli & Wirth, 1986b (key to females south of the Amazon Basin); Wirth & Blanton, 1959 (key to females and some males from Panama); Wirth & Blanton, 1973 (key to females and some males from Amazon Basin); Wirth & Blanton, 1974 (key to females and some males from the Caribbean); Wirth et al., 1988 (some meristic character states and photographs of wings of all species); Spinelli et al., 2005 (key to males and females from Argentina). Padrosia Rafinesque, 1815: 130 (unnecessary replacement name for Culicoides Latreille). Type species, Culicoides punctatus Latreille (= Ceratopogon punctatus Meigen), automatic. Comm.: There is no reliable key to the subgenera and species groups of Neotropical Culicoides. For species outside of local areas for which revisions are available (as listed above), the easiest guide to species identification is that of Wirth et al. (1988). From there readers may refer to keys to subgenera or species groups referred to below. Subgenus Anilomyia Vargas Anilomyia Vargas, 1960: 37 (as subgenus of Culicoides). Type species, Culicoides covagarciai Ortíz (orig. des.). Refs.: Wirth & Blanton, 1956b (key to males and females of covagarciai group); Wirth & Blanton, 1970c (key to key to males and females of nigrigenus group). ameliae Browne, 1980: 543. Type locality: Colombia, Cauca, Munchique, 2000 m. HT F (USNM). Distr.: Colombia (Cauca (Munchique)). Refs.: Browne, 1980: 543; Wirth et al., 1988: 20; Borkent & Spinelli, 2007: 62; Spinelli et al., 2009: 87; Borkent, 2015: 76 (cat.). covagarciai Ortíz, 1950a: 457. Type locality: Venezuela, Caracas. HT F (INHC). Distr.: Honduras to Colombia (Valle del Cauca (Cali, Pance, Estación El Topacio, Peñas Blancas (Finca El Rosal)), Venezuela. Refs.: Ortiz, 1950a: 457; Barreto, 1986: 142; Borkent & Spinelli, 2007: 62; Spinelli et al., 2009: 87. beebei Fox, 1952: 366. Type locality: Venezuela, Rancho Grande. HT F (UNPR). efferus Fox, 1952: 365. Type locality: Peru, Río Charape. HT F (UNPR). Distr.: Guatemala to Colombia (Valle del Cauca (Cali, Pance, Estacion El Topacio, Peñas Blancas)), Peru and Bolivia. Refs.: Fox, 1952: 365; Barreto, 1986: 142; Borkent & Spinelli, 2007: 62; Spinelli et al., 2009: 87. inermis Spinelli, Santamaría, Cabrera, Ronderos & Suárez, 2009: 84. Type locality: Colombia, Cauca, López, Río Micay. HT F (MLPA). Distr.: Colombia (Cauca (López, Río Micay)). Refs.: Spinelli et al., 2009: 84; Borkent, 2015: 88 (cat.). marshi Wirth & Blanton, 1956b: 220. Type locality: Panama, Bocas del Toro, Almirante. HT F (# 62957 USNM). Distr.: Costa Rica, Panama, Colombia (Valle del Cauca (Río Raposo; Cali, Pance, El Topacio)). Refs.: Wirth & Blanton, 1956b: 220; Barreto, 1986: 142; Borkent & Spinelli, 2007: 63; Spinelli et al., 2009: 87. metagonatus Wirth & Blanton, 1956b: 221. Type locality: Panama, Cerro Campana. HT F (# 62957 USNM). Distr.: Nicaragua to Colombia (Valle del Cauca (Río Raposo)) and Ecuador. Refs.: Wirth & Blanton, 1956b: 221; Barreto, 1986: 142; Borkent & Spinelli, 2007: 63; Spinelli et al., 2009: 87. CATALOGUE OF CERATOPOGONIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 107 nigrifemur Spinelli, Santamaría, Cabrera, Ronderos & Suárez, 2009: 85. Type locality: Colombia, Cauca, Parámo de Puracé, 28.4 km E Puracé, 3100 m. HT F (MLPA). Distr.: Colombia (Cauca (Parámo de Puracé, 28.4 km E Puracé, 3100 m)). Refs.: Spinelli et al., 2009: 85; Borkent, 2015: 94 (cat.). nigrigenus Wirth & Blanton, 1956b: 222. Type locality: Panama, Bocas del Toro, Almirante. HT F (# 62824 USNM). Distr.: Mexico (Veracruz) to Colombia (Valle del Cauca (Río Raposo)), Trinidad and Tobago, Argentina (Salta). Refs.: Wirth & Blanton, 1970c: 151; Barreto, 1986: 143; Borkent & Spinelli, 2007: 63; Spinelli et al., 2009: 87. popayanensis Wirth & Lee, 1967: 10. Distr.: Type locality: Colombia, Cauca, 28.4 km E Puracé, 3100 m. HT F (# 69399 USNM). Distr.: Colombia (Cauca (28.4 km E Puracé, 3100 m)). Refs.: Wirth & Lee, 1967: 10; Barreto, 1986: 143; Wirth et al., 1988: 22; Borkent & Spinelli, 2007: 63; Spinelli et al., 2009: 87; Borkent, 2015: 98 (cat.). trapidoi Wirth & Barreto, 1978: 554. Type locality: Colombia, Valle del Cauca (Río Raposo). HT F (# 71153 USNM). Distr.: Costa Rica, Colombia (Valle del Cauca (Río Raposo)), Brazil. Refs.: Wirth & Barreto, 1978: 554; Barreto, 1986: 143; Wirth et al. 1988: 22; Borkent & Spinelli, 2007: 63; Spinelli et al., 2009: 87; Borkent, 2015: 105 (cat.). Subgenus Avaritia Fox Avaritia Fox, 1955: 218 (as subgenus of Culicoides). Type species, Ceratopogon obsoletus Meigen (orig. des.). Refs.: Rodríguez & Wirth, 1986 (rev. of andicola group); Wirth & Mullens, 1992 (key to males and females in pusillus group). andicola Wirth & Lee, 1967: 5. Type locality: Colombia, Cauca, Parámo de Puracé, Lago Buey, 3250 m. HT F (# 69396 USNM). Distr.: Colombia (Cauca (Parámo de Puracé, Lago Buey, 3250 m; 28.4 km E Puracé, 3100 m)). Refs.: Wirth & Lee, 1967: 5; Barreto, 1986: 143; Wirth et al., 1988: 14; Borkent & Spinelli, 2007: 63; Spinelli et al., 2009: 87; Borkent, 2015: 76 (cat.). orjuelai Wirth & Lee, 1967: 6. Type locality: Colombia, Cauca, Parámo de Puracé, 18 km E. Puracé, 3320 m. HT F (# 69397 USNM). Distr.: Colombia (Cauca (Parámo de Puracé, Lago Buey, 3250 m; 28.4 km E Puracé, 3100 m)). Refs.: Wirth & Lee, 1967: 6; Barreto, 1986: 143; Wirth et al., 1988: 14; Borkent & Spinelli, 2007: 63; Spinelli et al., 2009: 87; Borkent, 2015: 96 (cat.). puracensis Wirth & Lee, 1967: 7. Type locality: Colombia, Cauca, Parámo de Puracé, Lago Buey, 3250 m. HT F (# 69398 USNM). Distr.: Colombia (Cauca (Parámo de Puracé, Lago Buey, 3250 m; 28.4 km E Puracé, 3100 m; 18 km E Puracé, 3320 m)). Refs.: Wirth & Lee, 1967: 7; Barreto, 1986: 143; Wirth et al., 1988: 14; Borkent & Spinelli, 2007: 63; Spinelli et al., 2009: 87; Borkent, 2015: 99 (cat.). pusillus Lutz, 1913: 52. Type locality: Brazil, Río de Janeiro, Manguinhos. ST F, M (FIOC). Distr.: USA (Florida), Mexico (Chiapas) to northeastern Argentina. Locality in Colombia (Valle del Cauca (Candelaria, Finca San Luis)). Refs.: Lutz, 1913: 52; Barreto, 1986: 143; Borkent & Spinelli, 2007: 63; Spinelli et al., 2009: 87. suarezi Rodríguez & Wirth, 1986: 313. Type locality: Colombia, Cundinamarca, Bogotá, D.E., Monserrate, 3230 m. HT F (USNM). Distr.: Colombia (Cundinamarca (Bogotá, D.E., Monserrate, 3230 m)). Refs.: Rodríguez & Wirth, 1986: 313; Barreto, 1986: 143; Wirth et al., 1988: 14; Borkent & Spinelli, 2007: 63; Spinelli et al., 2009: 87; Borkent, 2015: 103 (cat.). Subgenus Cotocripus Brèthes Cotocripus Brèthes, 1912: 451. Type species, Cotocripus caridei Brèthes (mon.). bambusicola Lutz, 1913: 62. Type locality: Brazil, Rio de Janeiro, Petrópolis. HT F (# 2982 FIOC). Distr.: Colombia (Valle del Cauca (área de Cali, Pance; Candelaria)), Venezuela, Brazil (Bahia, Espírito Santo, Rio de 108 · Zootaxa 4122 (1) © 2016 Magnolia Press SPINELLI & WOLFF Janeiro, São Paulo), Argentina (Misiones, Buenos Aires). Refs.: Lutz, 1913: 62; Lee, 1968: 81; Barreto, 1986: 145; Borkent & Spinelli, 2007: 64; Spinelli et al., 2009: 87. bahiensis Barbosa, 1947: 11. Type locality: Brazil, Bahia. ST F,M (USNM). gabrieli Spinelli, Santamaría, Cabrera, Ronderos & Suárez, 2009: 82. HT M (MLPA). Type locality: Peru, Cuzco, Kirigueti. Distr.: Colombia (Chocó (Pié de Pepe)), Peru. Refs.: Spinelli et al., 2009: 82; Borkent, 2015: 85 (cat.). raposoensis Wirth & Barreto, 1978: 561. Type locality: Colombia, Valle del Cauca (Río Raposo). HT F (# 71157 USNM). Distr.: Colombia (Valle del Cauca (Río Raposo; Río Micay, Casa de Suárez)). Refs.: Wirth & Barreto, 1978: 561; Barreto, 1986: 148 (as raposensis); Wirth et al., 1988: 54; Borkent & Spinelli, 2007: 64; Spinelli et al., 2009: 87; Borkent, 2015: 100 (cat.). Subgenus Diphaomyia Vargas Diphaomyia Vargas, 1960: 40 (as subgenus of Culicoides). Type species, Culicoides baueri Hoffman (orig. des.). iriartei Fox, 1952: 368. Type locality: Venezuela, Zulia, La Salina. HT F (UNPR). Distr.: Guatemala to Colombia (?), Venezuela, Tobago, Brazil (Pará). Refs.: Fox, 1952: 368; Wirth & Blanton, 1973: 441; Barreto, 1986: 143; Borkent & Spinelli, 2007: 64; Spinelli et al., 2009: 87. vargasi Wirth & Blanton, 1953a: 74. Type locality: Panama, Los Santos, Las Tablas. HT F (# 61502 USNM). lisicarruni Moncada, Carrasquilla & Spinelli, in Moncada et al., 2010: 978. Type locality: Colombia, Cundinamarca, Chingaza National Natural Park, Monterredondo Station edge of Babilonia River, 3200 m. HT F (ICNC). Distr.: Colombia (Cundinamarca (Chingaza National Natural Park, Monterredondo Station edge of Babilonia River, 3200 m)). Refs.: Spinelli et al., 2010: 978; Borkent, 2015: 91 (cat.). marinkellei Wirth & Lee, 1967: 13. Type locality: Colombia, Cauca, Parámo de Puracé, Lago Buey, 3250 m. HT F (# 69400 USNM). Distr.: Colombia (Cauca (Parámo de Puracé, Lago Buey, 3250 m; 18 km E Puracé, 3320 m)). Refs.: Wirth & Lee, 1967: 13; Barreto, 1986: 143; Wirth et al., 1988: 32; Borkent & Spinelli, 2007: 64; Spinelli et al., 2009: 87; Borkent, 2015: 92 (cat.). mirsae Ortíz, 1953: 801. Type locality, Venezuela, ST F (INHC). Distr.: Panama, Colombia (Boyacá (Otanche, Cortaderal)), Venezuela, Trinidad and Tobago. Refs.: Ortíz, 1953: 801; Borkent & Spinelli, 2007: 65; Spinelli et al., 2009: 87. Subgenus Drymodesmyia Vargas Drymodesmyia Vargas, 1960: 40 (as subgenus of Culicoides). Type species, Culicoides copiosus Root & Hoffman (orig. des.). Refs.: Wirth & Hubert, 1960: 646 (key to females and some males, as copiosus group). jamaicensis Edwards, 1922: 165 (as variety of loughnani. Type locality: Jamaica, Kingston. ST F, M (BMNH). Distr.: USA (Texas, Florida), Mexico (Yucatán), Central America and Caribbean to Colombia (?) and Venezuela. Refs.: Edwards, 1922: 165; Wirth et al., 1988: 26; Borkent & Spinelli, 2007: 65; Spinelli et al., 2009: 87. pilosus Wirth & Blanton, 1959: 332. Type locality: Panama, Bocas del Toro, Almirante. HT F (# 63164 USNM). Distr.: Costa Rica, Panama, Colombia (?) Brazil (Pará). Refs.: Wirth & Blanton, 1959: 332; Wirth & Blanton, 1973: 445; Barreto, 1986: 143; Borkent & Spinelli, 2007: 65; Spinelli et al., 2009: 88. poikilonotus Macfie, 1948: 82. Type locality: Mexico, Chiapas, El Vergel. HT F (BMNH). Distr.: Mexico (Chiapas), Central America, Colombia (?), Venezuela and Trinidad and Tobago to Brazil (Bahia). Refs.: Macfie, 1948: 82; Aitken et al., 1975: 136; Barreto, 1986: 143; Borkent & Spinelli, 2007: 65; Spinelli et al., 2009: 88. cacozelus Macfie, 1948: 85. Type locality: Mexico, HT F (BMNH). hertigi Wirth & Blanton, 1953b: 229. Type locality: Panama, Cocle, Río Hato. HT F (# 61504 USNM). CATALOGUE OF CERATOPOGONIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 109 Subgenus Haematomyidium Goeldi Haematomyidium Goeldi, 1905: 137. Type species, Haematomyidium Paráense Goeldi (orig. des). Refs.: Vitale et al., 1981 (key to females of debilipalpis group); Wirth & Felippe-Bauer, 1989 (key to paraensis group). debilipalpis Lutz, 1913: 60. Type locality: Brazil, São Paulo, Anhemby and Formoso (Serra da Bocaina). ST F (FIOC, lost). Distr.: Widespread from USA (Maryland, Kentucky, Nebraska south to Lousiana and Florida), Guatemala and Belize to Argentina. Localities in Colombia (Antioquia (?), Magdalena (?), Valle del Cauca (Río Raposo)). Refs.: Lutz, 1913: 60; Wirth & Blanton, 1974: 15; Barreto, 1986: 146; Borkent & Spinelli, 2007: 66; Spinelli et al. 2009: 88. ichesi Ronderos & Spinelli, 1995: 77. Type locality: Argentina, Misiones, Posadas, Arroyo Zaimán. HT F (MLPA). eldridgei Wirth & Barreto, 1978: 561. Type locality: Colombia, Valle del Cauca (Río Raposo). HT F (# 71157 USNM). Distr.: Colombia (Valle del Cauca (Río Raposo)). Refs.: Wirth & Barreto, 1978: 561; Wirth et al., 1988: 46; Barreto, 1986: 146; Borkent & Spinelli, 2007: 66; Spinelli et al., 2009: 88; Borkent, 2015: 83 (cat.). filiductus Wirth, in Vitale et al., 1981: 155. Type locality: Panama, Panamá Prov., Bayano Field Station. HT F (# 76107 USNM). Distr.: Belize to Colombia (Amazonas (Leticia)). Refs.: Vitale et al., 1981: 155; Borkent & Spinelli, 2007: 66; Spinelli et al., 2009: 86. germanus Macfie, 1940c: 27. Type locality: Guyana, New River. HT F (BMNH). Distr.: Costa Rica, Colombia (Tolima (Melgar, El Aguila)), Guyana. Refs.: Macfie, 1940c: 27; Borkent & Spinelli, 2007: 66; Spinelli et al., 2009: 86. ginesi Ortíz, 1951: 586. Type locality: Venezuela, Yaracuy, San Felipe. HT F (INHC). Distr.: El Salvador to Panama, Colombia (?), Venezuela, Trinidad and Tobago, Brazil (Pará), Northeastern Argentina. Refs.: Ortíz, 1951: 586; Wirth et al., 1988: 48; Borkent & Spinelli, 2007: 66; Spinelli et al., 2009: 88. glabrior Macfie, 1940c: 27 (as variety of debilipalpis). Type locality: Guyana, New River. HT F (BMNH). Distr.: Honduras to Colombia (?), Ecuador, Guyana, Surinam, Trinidad and Tobago, Brazil (Pará). Refs.: Macfie, 1940c: 27; Barreto, 1986: 147; Borkent & Spinelli, 2007: 66; Spinelli et al., 2009: 88. grahambelli Forattini, 1956: 35. Type locality, Panama, Cerro Cefa. HT F (DEFS). insinuatus Ortíz & León, 1955: 577. Type locality: Ecuador, Napo-Pastaza, Nuevo Rocafuerte. HT F (UNCE). Distr.: Colombia (Amazonas (Leticia, Santa Sofía, Río Arará y Puerto Arica, Río Putumayo), Vaupés (Mitú; Caño Cuduyarí)), Ecuador, Peru, Trinidad and Tobago, Guyana, Brazil (Amazonas, Pará, (?) São Paulo). Refs.: Ortíz & León, 1955: 577; Tidwell et al., 1980a: 125; Tidwell & Tidwell, 1982: 1139; Barreto, 1986: 147; Borkent & Spinelli, 2007: 67; Spinelli et al., 2009: 88. paraensis (Goeldi), 1905: 137 (Haematomyidium). Type locality: Brazil, Pará, Belém. NT F (FIOC). Distr.: USA (Colorado, Nebraska, Pennsylvania, Wisconsin to Louisiana and Florida) to Argentina. Localities in Colombia (Amazonas (Leticia; Arará, vic. Leticia), Antioquia (Río Anorí Valley, Buenos Aires, 690 m; Providencia; Tiroteos, 590 m), Caquetá (Tres Esquinas), Meta (Villavicencio, El Buque), Tolima (Mariquita, Florazul; Vereda, La Guardia, Finca Bélgica; Melgar, El Aguila), Valle del Cauca (Cali; Buchitolo; Navarro, Vaupés (San José de Inambú; San Pablo; Ortega)). Refs.: Goeldi, 1905: 137; Barreto, 1974: 124; Wirth & Blanton, 1973: 443; Barreto, 1986: 147; Wirth & Felippe-Bauer, 1989: 553; Borkent & Spinelli, 2007: 67; Spinelli et al., 2009: 88. undecimpunctatus Kieffer, 1917b: 307. Type locality: Argentina, San Pablo. T F (HNHM, lost). quasiparaensis Clastrier, 1971: 286. Type locality: French Guiana, Cayena, Bosque Cabassou. HT F (MNHN). Distr.: Honduras and El Salvador to Colombia (Antioquia (Río Anorí Valley, Buenos Aires, 690 m; La Tirana, 320 m), Putumayo (Puerto Leguizamo, Santa Cecilia), Valle del Cauca (Candelaria, Finca San Luis)), Peru, French Guiana, Brazil (Amazonas, Rondônia). Refs.: Clastrier, 1971: 286; Barreto, 1986: 148; Wirth & Felippe Bauer, 1989: 561; Borkent & Spinelli, 2007: 67; Spinelli et al., 2009: 88. 110 · Zootaxa 4122 (1) © 2016 Magnolia Press SPINELLI & WOLFF youngi Wirth & Barreto, 1978: 562. Type locality: Colombia, Valle del Cauca (Río Raposo). HT F (# 71159 USNM). Distr.: Colombia (Valle del Cauca (Río Raposo)). Refs.: Wirth & Barreto, 1978: 562; Barreto, 1986: 148; Wirth et al., 1988: 50; Borkent & Spinelli, 2007: 67; Spinelli et al., 2009: 88; Borkent, 2015: 107 (cat.). Subgenus Hoffmania Fox Hoffmania Fox, 1948: 21 (as subgenus of Culicoides). Type species, Culicoides inamollae Fox & Hoffman (= Culicoides insignis Lutz) (orig. des.). Refs.: Spinelli et al., 1993 (key to species groups; key to males and females of guttatus group); Wirth & Blanton, 1968a (key to males and females in hylas group); Felippe-Bauer et al., 2009 (key to females in hylas group). antioquiensis Spinelli, Santamaría, Cabrera, Ronderos & Suárez, 2009: 82. Type locality: Colombia, Antioquia, near Río Anorí. HT M (MLPA). Distr.: Colombia (Antioquia (near Río Anorí)). Refs.: Spinelli et al., 2009: 82; Borkent, 2015: 76 (cat.). batesi Wirth & Blanton, 1973: 426. Type locality: Brazil, Pará, Belém, APEG Forest. HT F (USNM). Distr.: Guatemala, Colombia (Cauca (López, Río Micay, Casa de Suárez), Chocó (Rt. 25; Curiche; Teresita), Valle del Cauca (Río Raposo)), Ecuador, Bolivia, Brazil (Pará). Refs.: Wirth & Blanton, 1973: 426; Wirth et al., 1988: 14; Spinelli et al., 1993: 18; Borkent & Spinelli, 2007: 67; Spinelli et al., 2009: 88. sanmartini Wirth & Barreto, 1978: 553. Type locality: Colombia, Valle del Cauca (Río Raposo). HT F (# 71152 USNM). brownei Spinelli, in Spinelli et al. 1993: 24. Type locality: Colombia, Valle del Cauca (Río Raposo). HT F (USNM). Distr.: Colombia (Boyacá (Palagua, Puerto Boyacá), Valle del Cauca (Río Raposo)). Refs.: Spinelli et al. 1993: 24; Borkent & Spinelli, 2007: 68; Spinelli et al., 2009: 88; Borkent, 2015: 79 (cat.). coutinhoi Barretto, 1944: 96. Type locality: Brazil, São Paulo, Juquiá, Prainha. HT M (DEFS). Distr.: Colombia (Caquetá (San Vicente del Caguán, Tres Esquinas)), French Guiana, Brazil (Amazonas, Pará, São Paulo). Refs.: Barretto, 1944: 96; Borkent & Spinelli, 2007: 68; Spinelli et al., 2009: 86. davidi Spinelli, in Spinelli et al., 1993: 30. Type locality: Colombia, Chocó, Rt. 25. HT F (USNM). Distr.: Costa Rica, Colombia (Chocó (Rt. 25)), Trinidad and Tobago. Refs.: Spinelli et al., 1993: 30; Borkent & Spinelli, 2007: 68; Spinelli et al., 2009: 88; Borkent, 2015: 82 (cat.). diabolicus Hoffman, 1925: 294. Type locality: Panama, Cabima. ST F (USNM). Distr.: Mexico to Venezuela, Colombia (Cauca (López, Río Micay, Casa de Suárez), Chocó (Rt. 25; Curiche), Meta (Puerto López; San Martín, Barbascal), Boyacá (Puerto Boyacá)) and Ecuador. Refs.: Hoffman, 1925: 294; Macfie, 1932b: 487; Barreto, 1986: 143; Wirth et al., 1988: 14; Spinelli et al., 1993: 32; Borkent & Spinelli, 2007: 68; Spinelli et al., 2009: 88. fernandoi Tavares & Souza, 1979: 611. Type locality: Brazil, Rio de Janeiro, Sumidouro, Sitio Peri-Ceci. HT M (FIOC). Distr.: Colombia (Meta (Finca Barbascal)), Brazil (Espírito Santo, Rio de Janeiro, Santa Catarina), northeastern Argentina, Uruguay. Refs.: Tavares & Souza, 1979: 611; Spinelli et al., 1993: 36; Borkent & Spinelli, 2007: 68; Spinelli et al., 2009: 88. filarifer Hoffman, 1939: 172. Type locality: Mexico, Chiapas, El Vergel, near Huixtla. HT F (USNM). Distr.: Mexico (Veracruz, Chiapas) to northern Brazil. Localities in Colombia (?). Refs.: Hoffman, 1939: 172; Barreto, 1986: 144 Spinelli et al., 1993: 37; Borkent & Spinelli, 2007: 68; Spinelli et al., 2009: 88. foxi Ortíz, 1950b: 461. Type locality: Puerto Rico, Camp Tortuguero. HT M (UNPR). Distr.: Mexico (Chiapas) to Bolivia, Puerto Rico to northeastern Argentina; Colombia (Caquetá (Los Alicángaros), Cauca (Tambo, Finca Carpenteria), Chocó (Teresita), Magdalena (?); Meta (Finca Barbascal; Restrepo, El Retiro, Refugio Macarena), Putumayo (Puerto Leguizamo; Puerto López), Valle del Cauca (Pance; Río Raposo; oleoducto de Pacific, 10 km NW Dagua)). Refs.: Ortíz, 1950b: 461; Forattini, 1957: 205; Wirth & Blanton, 1973: 434; Wirth & Blanton, 1974: 14; Barreto, 1986: 144; Spinelli et al., 1993: 41; Borkent & Spinelli, 2007: 68; Spinelli et al., 2009: 88. CATALOGUE OF CERATOPOGONIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 111 franklini Spinelli, in Spinelli et al., 1993: 45. Type locality: Panama, Canal Zone, Summit Gardens. HT F (USNM). Distr.: Mexico (Guerrero) to Colombia (Chocó (Teresita), Valle del Cauca (Río Raposo)), Bolivia, Brazil (Pará). Refs.: Spinelli et al., 1993: 45; Borkent & Spinelli, 2007: 68; Spinelli et al., 2009: 88. fusipalpis Wirth & Blanton, 1973: 435. Type locality: Brazil, Pará, Belém. HT F (USNM). Distr.: El Salvador to Colombia (Caquetá (Los Alicángaros), Putumayo (Puerto Leguizamo), Valle del Cauca (Río Raposo)), Ecuador, Bolivia, French Guiana, Guyana, Brazil (Amazonas, Pará, Bahia, Rio de Janeiro). Refs.: Wirth & Blanton, 1973: 435; Aitken et al., 1975: 125; Barreto, 1986: 144; Borkent & Spinelli, 2007: 68; Spinelli et al., 2009: 88. heliconiae Fox & Hoffman, 1944: 108. Type locality: Venezuela, Maracay. HT F (UNPR). Distr.: Belize to Colombia (Antioquia (?), Caquetá (Tres Esquinas, Los Alicángaros), Huila (Suaza, San Calixto), Valle del Cauca (Cali, Pance, Peñas Blancas; Rio Raposo)), Ecuador, Venezuela, Grenada, Trinidad and Tobago, Brazil (Amazonas, Pará). Refs.: Fox & Hoffman, 1944: 108; Wirth & Blanton, 1968: 207; Wirth & Blanton, 1973: 439; Wirth & Blanton, 1974: 14; Barreto, 1986: 144; Wirth et al., 1988: 18; Borkent & Spinelli, 2007: 68; Spinelli et al., 2009: 88. rozeboomi Barbosa, 1947: 26. Type locality: Panama, Balboa, Canal Zone. HT M (USNM). hylas Macfie, 1940c: 26. Type locality: Guyana, New River. ST F (BMNH). Distr.: Mexico (Veracruz) to Colombia (Caquetá (Tres Esquinas), Cauca (López, Río Micay, San Antonio, Casa de Suárez), Valle del Cauca (Río Raposo)), Peru, Brazil (Pará). Refs.: Macfie, 1940c: 26; Wirth & Blanton, 1968: 203; Wirth & Blanton, 1973: 439; Barreto, 1986: 144; Borkent & Spinelli, 2007: 68; Spinelli et al., 2009: 88. insignis Lutz, 1913: 51. Type locality: Brazil, Bahia and Rio de Janeiro. LT M (# 5092 FIOC). Distr.: USA (Alabama, Georgia, Florida), Mexico (Yucatán, Chiapas), Central America and Caribbean to central Argentina. Localities in Colombia (Amazonas (Sofía), Antioquia (El Hatillo, Barbosa; Caucasia, Hacienda La Candelaria), Boyacá (Palagua, Puerto Boyacá), Cauca (11 km N Santander de Quilchao; Puerto López), Chocó (Condoto, Opogodó), César (San Alberto, 120 km N Bucaramanga), Huila (San Agustín), Magdalena (La Gloria), Meta (Finca Barbascal; Restrepo, El Retiro, Refugio Macarena), Putumayo (Puerto Leguizamo), Calderón), Valle del Cauca (Cali, Candelaria, Pance; Palmira; Río Raposo)). Refs.: Lutz, 1913: 51; Wirth & Blanton, 1956: 319; Forattini, 1957: 223; Wirth & Blanton, 1973: 440; Wirth & Soria, 1981: 79; Barreto, 1986: 144; Spinelli et al., 1993: 53; Borkent & Spinelli, 2007: 68; Spinelli et al., 2009: 88. inamollae Fox & Hoffman, 1944: 110. Type locality: Puerto Rico, Mayaguez. HT F (UNPR). painteri Fox, 1946: 257. Type locality: Honduras, Puerto Castilla. HT F (UNPR). lutzi Costa Lima, 1937: 419. Type locality: Brazil, Pará, Abaeté. ST F (# 2408 FIOC). Distr.: Colombia (Amazonas (Sofía), Boyacá (Palagua, Puerto Boyacá), Meta (Finca Barbascal; Refugio Macarena), Putumayo (Puerto Leguizamo)) to French Guiana, northeastern Argentina, Brazil (Roraima, Amazonas, Pará, Mato Grosso, Goiás, São Paulo, Rio de Janeiro, Santa Catarina). Refs.: Costa Lima, 1937: 419; Wirth & Blanton, 1973: 442; Barreto, 1986: 144; Spinelli et al., 1993: 59; Borkent & Spinelli, 2007: 68; Spinelli et al., 2009: 88. ocumarensis Ortíz, 1950a: 455. Type locality: Venezuela, Miranda, Ocumare del Tuy. HT F (INHC). Distr.: Mexico (Oaxaca, Tabasco) to Colombia (Amazonas (Sofía), Antioquia (Caucasia, Finca La Candelaria; Barbosa, El Establo; Turbo, Río León), Boyacá (Puerto Boyacá), Caquetá (Tres Esquinas), Cauca (López, Río Micay, San Antonio, Casa de Suárez; Puerto López), Cesar (San Alberto, 120 km N Bucaramanga), Meta (Finca Barbascal), Valle del Cauca (Río Raposo)), Venezuela, Peru and northern Brazil (Pará, Rondônia). Refs.: Ortíz, 1950a: 455; Spinelli et al., 1993: 63; Borkent & Spinelli, 2007: 69; Spinelli et al., 2009: 88. palpalis Macfie, 1948: 78. Type locality: Mexico, Chiapas, San Cristobal. HT F (BMNH). Distr.: Mexico (Chiapas) to Colombia (Caquetá (Los Alicángaros)), Venezuela, Peru, Brazil (Amazonas). Refs.: Macfie, 1948: 78; Wirth & Blanton, 1973: 443; Barreto, 1986: 144; Borkent & Spinelli, 2007: 69; Spinelli et al., 2009: 88. paraignacioi Spinelli, in Spinelli et al., 1993: 66. Type locality: Colombia, Antioquia, near Río Anorí. HT F (USNM). Distr.: Belize to Peru, Colombia (Antioquia (near Río Anorí), Caquetá (Los Alicángaros), Chocó (Curiche; Rt. 25; Teresita), Guaviare (San José del Guaviare, Trocha Oriental), Putumayo (Puerto Leguizamo)), 112 · Zootaxa 4122 (1) © 2016 Magnolia Press SPINELLI & WOLFF French Guiana, Brazil (Amazonas, Pará). Refs.: Spinelli et al., 1993: 66; Borkent & Spinelli, 2007: 69; Spinelli et al., 2009: 88; Borkent, 2015: 97 (cat.). plaumanni Spinelli, in Spinelli et al., 1993: 69. Type locality: Argentina, Chaco, Parque Nacional El Chaco. HT F (MLPA). Distr.: Colombia (Meta (Villavicencio, El Buque)), Peru, Bolivia, Brazil (Amazonas) northeastern Argentina. Refs.: Spinelli et al., 1993: 69; Borkent & Spinelli, 2007: 69; Spinelli et al., 2009: 91. polypori Wirth & Blanton, 1968: 212. Type locality: Panama, Canal Zone, Barro Colorado Island. HT F (# 69916 USNM). Distr.: Honduras to Colombia (Valle del Cauca (Río Raposo)), Brazil (Amazonas). Refs.: Wirth & Blanton, 1968: 212; Barreto, 1986: 144; Wirth et al., 1988: 20; Borkent & Spinelli, 2007: 69; Spinelli et al., 2009: 88. pseudodiabolicus Fox, 1946: 256. Type locality: Trinidad and Tobago, Cumuto Village. HT F (UNPR). Distr.: Mexico (Veracruz) to Colombia (Antioquia (near Río Anorí), Caquetá (Los Alicángaros; Tres Esquinas), Cauca (López, Río Micay), Chocó (Curiche; Rt. 25; Teresita), Meta, Boyacá (Puerto Boyacá), Putumayo (Puerto Leguizamo), Valle del Cauca (Río Raposo)), Venezuela, Peru and Northern Brazil. Refs.: Fox, 1946: 256; Wirth & Blanton, 1973: 445; Barreto, 1986: 144; Spinelli et al., 1993: 71; Borkent & Spinelli, 2007: 69; Spinelli et al., 2009: 88. ruizi Forattini, 1954: 189. Type locality: Brazil, Goiás, Aruana. HT F (DEFS). Distr.: Colombia (Intendencia de Arauca), Brazil (Amazonas, Goiás). Refs.: Forattini, 1954: 189; Wirth et al., 1988: 18; Spinelli et al., 1993: 74; Borkent & Spinelli, 2007: 69; Spinelli et al., 2009: 88. tidwelli Spinelli, in Spinelli et al. 1993: 74. Type locality: Colombia, Valle del Cauca (Río Raposo). HT F (USNM). Distr.: Honduras to Colombia (Cauca (López, Río Micay), Chocó (Curiche; Teresita), Valle del Cauca (Río Raposo; Oleoducto de Pacífico, 9 km N Dagua)), Ecuador. Refs.: Spinelli et al., 1993: 74; Borkent & Spinelli, 2007: 69; Spinelli et al., 2009: 88; Borkent, 2015: 104 (cat.). trinidadensis Hoffman, 1925: 286. Type locality: Trinidad and Tobago, Port of Spain, Caronia River. HT F (USNM). Distr.: Coastal; Honduras and El Salvador to Colombia (Antioquia (Currulao), Chocó (Curiche), Valle del Cauca (Río Raposo, Bahía Buenaventura, Bella Vista)), Ecuador to Surinam, Cuba and Cayman Islands to Trinidad and Tobago. Refs.: Hoffman, 1925: 286; Wirth & Blanton, 1974: 14; Barreto, 1986: 145; Wirth et al., 1988: 18; Spinelli et al., 1993: 78; Borkent & Spinelli, 2007: 69; Spinelli et al., 2009: 89. oliveri Fox & Hoffman, 1944: 108. Type locality: Haiti, Mariani. HT M (UNPR). wokei Barbosa, 1947: 28 (preocc. Culicoides wokei Fox, 1947). Type locality: Panama, Balboa. LT M (USNM). diminutus Barbosa, 1951: 163. New name for wokei. verecundus Macfie, 1948: 76. Type locality: Mexico, Chiapas, El Vergel. ST F, M (BMNH). Distr.: Mexico (Chiapas) to Panama, Colombia (Valle del Cauca (Río Raposo)), Ecuador and Brazil (Amazonas, Pará). Refs.: Macfie, 1948: 76; Barreto, 1986: 145; Borkent & Spinelli, 2007: 69; Spinelli et al., 2009: 89. Subgenus Macfiella Fox Macfiella Fox, 1955: 217 (as subgenus of Culicoides). Type species: Ceratopogon phlebotomus Williston (orig. des). phlebotomus (Williston), 1896: 281 (Ceratopogon). Type locality: St. Vincent. HT F (BMNH). Distr.: Coastal; Mexico (Sinaloa) to Colombia (Valle del Cauca (Río Raposo)), Ecuador, Jamaica to Brazil (Maranhão, Ceará, Pernambuco, Goiás). Refs.: Williston, 1896: 281; Barreto, 1986: 145; Borkent & Spinelli, 2007: 69; Spinelli et al., 2009: 89. amazonius Macfie, 1935: 52. Type locality: Brazil, Piauí, Tutóia. ST FM (BMNH). CATALOGUE OF CERATOPOGONIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 113 Subgenus Mataemyia Vargas Mataemyia Vargas, 1960: 43 (as subgenus of Culicoides). Type species, Culicoides mojingaensis Wirth & Blanton (orig.des.). Refs.: Wirth & Soria, 1981 (key males and females to some species, as discrepans group). dalessandroi Wirth & Barreto, 1978: 556. Type locality: Colombia, Valle del Cauca (Río Raposo). HT F (# 71154 USNM). Distr.: Costa Rica, Panama, Colombia (Antioquia (Río Anorí), Boyacá (Puerto Boyacá), Meta (Finca Barbascal), Valle del Cauca (Río Raposo)). Refs.: Wirth & Barreto, 1978: 556; Barreto, 1986: 146; Wirth et al., 1988: 36; Borkent & Spinelli, 2007: 70; Spinelli et al., 2009: 89; Borkent, 2015: 81 (cat.). dicrourus Wirth & Blanton, 1955b: 123. Type locality: Panama, Canal Zone, Loma Borracho. HT F (# 62810 USNM). Distr.: Costa Rica to Colombia (Valle del Cauca (Río Raposo)), Ecuador. Refs.: Wirth & Blanton, 1955b: 123; Wirth & Soria, 1981: 110; Barreto, 1986: 146; Borkent & Spinelli, 2007: 70; Spinelli et al., 2009: 89. volcanensis Wirth & Blanton, 1959: 389. Type locality: Panama, Chiriquí, Volcán. HT F (# 63168 USNM). Distr.: Panama, Colombia (?). Refs.: Wirth & Blanton, 1959: 389; Wirth, 1974: 38; Borkent & Spinelli, 2007: 70; Spinelli et al., 2009: 89. Subgenus Oecacta Poey Oecacta Poey, 1853: 238. Type species, Oecacta furens Poey (mon.). alahialinus Barbosa, 1952: 11 (1953: 12). Type locality: Ecuador, Esmeralda, Limones. HT F (UNCE). Distr.: Costa Rica, Panama, Colombia (Valle del Cauca (Río Raposo)), Ecuador. Refs.: Barbosa, 1952: 11; Barreto, 1986: 145; Borkent & Spinelli, 2007: 70; Spinelli et al., 2009: 89. barbosai Wirth & Blanton, 1956a: 161. Type locality: Panama, Canal Zone, Mojinga Swamp. HT F (# 63157 USNM). Distr.: USA (Florida) to Colombia (Valle del Cauca (Río Raposo)), Ecuador.Refs.: Wirth & Blanton, 1956a: 161; Wirth & Blanton, 1974: 15; Barreto, 1986: 145; Borkent & Spinelli, 2007: 70; Spinelli et al., 2009: 89. furens (Poey), 1853: 238 (Oecacta). Type locality: Cuba, north coast, probably La Habana. Type most probably lost. Distr.: USA (Massachusetts to Florida and Texas), Mexico (Campeche, Sinaloa, Yucatán, Veracruz) and Caribbean to Colombia (Magdalena (?); Valle del Cauca (Río Raposo)), Ecuador and coastal Brazil. Refs.: Poey, 1853: 238; Wirth & Blanton, 1974: 15; Barreto, 1986: 145; Borkent & Spinelli, 2007: 70; Spinelli et al., 2009: 89. maculithorax (Williston), 1896: 277 (Ceratopogon). Type locality: St. Vincent. HT F (BMNH). birabeni Cavalieri, 1966: 59. Venezuela, Sucre, Cariaguito. HT F (INMA). gorgasi Wirth & Blanton, 1953b: 232. Type locality: Panama, Los Santos, Las Tablas. HT F (# 61507 USNM). Distr.: Costa Rica to Colombia (?). Refs.: Wirth & Blanton, 1953b: 232; Wirth, 1974: 42; Borkent & Spinelli, 2007: 70; Spinelli et al., 2009: 89. Subgenus Psychophaena Philippi Psychophaena Philippi, 1865: 628. Type species, Psychophaena pictipennis Philippi (= Culicoides venezuelensis Ortíz & Mirsa) (mon.). venezuelensis Ortíz & Mirsa, 1950: 137. Type locality, Venezuela, Miranda, Los Chorros. HT F (INHC). Distr.: Costa Rica to Chile and Central Argentina; Colombia (Valle del Cauca (Cali, Pance; Río Raposo)). Refs.: Ortíz & Mirsa, 1950: 137; Barreto, 1986: 148; Borkent & Spinelli, 2007: 71; Spinelli et al., 2009: 89. pictipennis (Philippi), 1865: 628 (Psychophaena) (preocc. Culicoides pictipennis (Staeger, 1839). Type locality: Chile, Santiago, San Cristóbal. LHT F (MUNS). ortizi Fox, 1952: 366. Type locality, Venezuela, Zulia, La Salina. HT F (UNPR). 114 · Zootaxa 4122 (1) © 2016 Magnolia Press SPINELLI & WOLFF Subgenus unplaced, acotylus Species Group acotylus Lutz, 1913: 69. Type locality: Brazil, Mato Grosso, Río Tapajós, Salto Augusto. HT F (# 2984 FIOC). Distr.: Mexico to Colombia (?), Venezuela, Trinidad and Tobago, Surinam, Brazil (Mato Grosso, Pará). Refs.: Lutz, 1913: 69; Wirth, 1974: 27; Barreto, 1986: 145; Borkent & Spinelli, 2007: 71; Spinelli et al., 2009: 89. carsiomelas Wirth & Blanton, 1955a: 100. Type locality: Panama, Canal Zone, Mojinga Swamp. HT F (# 62806 USNM). Distr.: Panama, Colombia (Valle del Cauca (Río Raposo)), Brazil (Pará). Refs.: Wirth & Blanton, 1955a: 100; Barreto, 1986: 146; Wirth et al., 1988: 38; Borkent & Spinelli, 2007: 71; Spinelli et al., 2009: 89. teretipalpis Wirth & Barreto, 1978: 557. Type locality: Colombia, Valle del Cauca (Río Raposo). HT F (# 71156 USNM). Distr.: Colombia (Antioquia (Río Anorí), Valle del Cauca (Río Raposo)). Refs.: Wirth & Barreto, 1978: 557; Barreto, 1986: 148; Wirth et al., 1988: 38; Borkent & Spinelli, 2007: 71; Spinelli et al., 2009: 89; Borkent, 2015: 104 (cat.). Subgenus unplaced, carpenteri Species Group belemensis Wirth & Blanton, 1973: 427. Type locality: Brazil, Pará, Belém. HT F (USNM). Distr.: Colombia (?), Brazil (Amazonas, Pará). Refs.: Wirth & Blanton, 1973: 427; Wirth et al., 1988: 32; Borkent & Spinelli, 2007: 71; Spinelli et al., 2009: 89. camposi Ortíz & León, 1955: 580. Type locality: Ecuador, Pichincha, San Miguel de los Colorados, Zapallo Grande. HT F (UNCE). Distr.: Costa Rica, Panama, Colombia (Putumayo (Puerto Leguizamo), Valle del Cauca (Río Raposo)), Ecuador. Refs.: Ortíz & León, 1955: 580; Wirth & Blanton, 1973: 428; Barreto, 1986: 145; Wirth et al., 1988: 32; Borkent & Spinelli, 2007: 71; Spinelli et al., 2009: 89. fairchildi Wirth & Blanton, 1955a: 102. Type locality: Panama, Canal Zone, Madden Dam. HT F (# 62801 USNM). Subgenus unplaced, daedalus Species Group beaveri Wirth & Barreto, 1978: 557. Type locality: Colombia, Valle del Cauca (Río Raposo). HT F (# 71155 USNM). Distr.: Colombia (Valle del Cauca (Río Raposo)). Refs.: Wirth & Barreto, 1978: 557; Barreto, 1986: 145; Wirth et al., 1988: 28; Borkent & Spinelli, 2007: 71; Spinelli et al., 2009: 89. crescentis Wirth & Blanton, 1959: 317. Type locality: Panama, Canal Zone, Mojinga Swamp. HT F (# 63159 USNM). Distr.: Mexico (Chiapas) to Colombia (?), northeastern Argentina. Refs.: Wirth & Blanton, 1959: 317; Wirth et al., 1988: 28; Borkent & Spinelli, 2007: 71; Spinelli et al., 2009: 89. daedaloides Wirth & Blanton, 1959: 330. Type locality: Panama, Bocas del Toro, Amirante. HT F (# 63163 USNM). Distr.: Panama, Colombia (Valle del Cauca (Río Raposo)), northwestern Argentina. Refs.: Wirth & Blanton, 1959: 330; Barreto, 1986: 148; Borkent & Spinelli, 2007: 71; Spinelli et al., 2009: 89. daedalus Macfie, 1948: 83. Type locality, Mexico, Chiapas, El Vergel. HT M (BMNH). Distr.: USA (Arizona, New Mexico), Mexico (Chiapas) to Colombia (?). Refs.: Macfie, 1948: 83; Wirth et al., 1988: 28; Borkent & Spinelli, 2007: 71; Spinelli et al., 2009: 89. Subgenus unplaced, dasyophrus Species Group dasyophrus Macfie, 1940c: 27. Type locality: Guyana, New River. ST M, F (BMNH). Distr.: Colombia (Meta (Refugio Macarena)), Ecuador, Venezuela, Guyana, Brazil (Amazonas, Mato Grosso, Pará). Refs.: Macfie, 1940c: 27; Wirth & Blanton, 1973: 430; Barreto, 1986: 146; Wirth et al., 1988: 50; Borkent & Spinelli, 2007: 72; Spinelli et al., 2009: 89. CATALOGUE OF CERATOPOGONIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 115 Subgenus unplaced, eublepharus Species Group caldasi Browne, 1980: 535. Type locality: Colombia, Valle del Cauca, Cali, Peñas Blancas, Finca El Rosal, 1800 m. HT F (USNM). Distr.: Colombia (Valle del Cauca (Cali, Peñas Blancas, Finca El Rosal, 1800 m)). Refs.: Browne, 1980: 535; Barreto, 1986: 145; Wirth et al., 1988: 42; Borkent & Spinelli, 2007: 72; Spinelli et al., 2009: 89; Borkent, 2015: 79 (cat.). caucaensis Wirth & Lee, 1967: 19. Type locality: Colombia, Cauca, 28.4 km E Puracé, 3100 m. HT F (# 69403 USNM). Distr.: Colombia (Cauca (28.4 km E Puracé, 3100 m)). Refs.: Wirth & Lee, 1967: 19; Barreto, 1986: 146; Wirth et al., 1988: 42; Borkent & Spinelli, 2007: 72; Spinelli et al., 2009: 89; Borkent, 2015: 80 (cat.). eublepharus Macfie, 1948: 86. Type locality: Guyana, New River. HT F (lost). Distr.: Mexico (Chiapas), Costa Rica, Colombia (Antioquia (Río Anorí), Caquetá (Tres Esquinas)) to Ecuador, Venezuela, northern Brazil. Refs.: Macfie, 1948: 86; Wirth & Blanton, 1973: 431; Barreto, 1986: 146; Wirth et al., 1988: 44; Borkent & Spinelli, 2007: 72; Spinelli et al., 2009: 89. transferrans Ortíz, 1953: 801. Type locality: Venezuela, Amazonas, head Río Orinoco. HT F (INHC). florenciae Messersmith, 1972: 167. Type locality: Colombia, Caquetá, Florencia, Ríohacha, 1000 m. HT F (# 73147 USNM). Distr.: Colombia (Caquetá (Florencia), Guajira (Ríohacha, 1000 m), Valle del Cauca (Cali, Peñas Blancas, Finca El Rosal, 1800 m)). Refs.: Messersmith, 1972: 167; Browne, 1980: 533; Barreto, 1986: 146; Wirth et al., 1988: 44; Borkent & Spinelli, 2007: 72; Spinelli et al., 2009: 89; Borkent, 2015: 84 (cat.). micayensis Spinelli, Santamaría, Cabrera, Ronderos & Suárez, 2009: 85. HT F (MLPA). Type locality: Colombia, Cauca, López, Río Micay. Distr.: Colombia (Cauca (López, Río Micay)). Refs.: Spinelli et al., 2009: 85; Borkent, 2015: 93 (cat.). pabloi Browne, 1980: 541. Type locality: Colombia, Valle del Cauca, Cali, Pance, 1600 m. HT F (USNM). Distr.: Colombia (Valle del Cauca (Cali, Pance, 1600 m; Peñas Blancas)). Refs.: Browne, 1980: 541; Barreto, 1986: 147; Wirth et al., 1988: 44; Borkent & Spinelli, 2007: 72; Spinelli et al., 2009: 89; Borkent, 2015: 96 (cat.). rangeli Ortíz & Mirsa, 1952b: 126. Type locality: Venezuela, Miranda, Los Chorros. HT F (INHC). Distr.: Mexico (Oaxaca) to Colombia. Locality in Colombia (Valle del Cauca (Cali, Peñas Blancas; Río Raposo)), Ecuador, Bolivia, Venezuela, Guadeloupe, Trinidad and Tobago, Brazil (Amazonas). Refs.: Ortíz & Mirsa, 1952b: 126; Barreto, 1986: 148; Borkent & Spinelli, 2007: 72; Spinelli et al., 2009: 89. donajii Vargas, 1954: 28. Type locality: Mexico, Oaxaca. HT M (CAIM). patulipalpis Wirth & Blanton, 1959: 421. Type locality: Panama, Panama prov., Cerro Campana. HT F (# 63170 USNM). tamboensis Wirth & Lee, 1967: 20. Type locality: Colombia, Cauca, El Tambo, Finca Carpenteria, 2500 m. HT F (# 69404 USNM). Distr.: Colombia (Cauca (El Tambo, Finca Carpenteria, 2500 m), Valle del Cauca (Cali, Pance)). Refs.: Wirth & Lee, 1967: 20; Barreto, 1986: 148; Wirth et al., 1988: 44; Borkent & Spinelli, 2007: 72; Spinelli et al., 2009: 89; Borkent, 2015: 104 (cat.). Subgenus unplaced, fluvialis Species Group balsapambensis Ortíz & León, 1955: 569 (as variety of pifanoi). Type locality: Ecuador, Bolivar, Balsapamba. ST F (UNCE). Distr.: Costa Rica to Colombia (Valle del Cauca (Cali, Pance)), Ecuador, Brazil. Refs.: Ortíz & León, 1955: 569; Barreto, 1986: 145; Borkent & Spinelli, 2007: 72; Spinelli et al., 2009: 89. castillae Fox, 1946: 251. Type locality: Honduras, Puerta Castilla. HT F (UNPR). Distr.: Guatemala to Colombia (Boyacá (Pauna, Topo Grande), Caquetá (Solano)), Ecuador, Venezuela, Trinidad and Tobago. Refs.: Fox, 1946: 251; Borkent & Spinelli, 2007: 72; Spinelli et al., 2009: 86. gibsoni Wirth, 1952: 246. Type locality: Guatemala, Chimaltenango, San Pedro Yepocapa. HT F (# 61407 USNM). flochabonnenci Ortíz & Mirsa, 1952a: 267. Type locality: Venezuela, Miranda, Los Chorros. HT F (INHC). 116 · Zootaxa 4122 (1) © 2016 Magnolia Press SPINELLI & WOLFF fluvialis Macfie, 1940c: 25. Type locality: Guyana, New River. HT F (BMNH). Distr.: Honduras to Colombia (?), Venezuela, Trinidad and Tobago, Guyana, Brazil (Amazonas, Pará). Refs.: Macfie, 1940c: 25; Wirth et al., 1988: 44; Borkent & Spinelli, 2007: 72; Spinelli et al., 2009: 86. leopoldoi Ortíz, 1951: 579. Type locality: Venezuela, Miranda, Ocumare del Tuy. HT F (INHC). Distr.: Guatemala and Belize to Bolivia and northeastern Argentina, Trinidad and Tobago, Colombia (Valle del Cauca (Río Raposo)). Refs.: Ortíz, 1951: 579; Barreto, 1986: 147; Borkent & Spinelli, 2007: 72; Spinelli et al., 2009: 90. Subgenus unplaced, leoni Species Group fieldi Wirth & Blanton, 1956c: 50. Type locality: Honduras, Lancetilla. HT F (# 63130 USNM). Distr.: Honduras, Costa Rica, Panama, Colombia (Valle del Cauca (Río Raposo)). Refs.: Wirth & Blanton, 1956c: 50; Barreto, 1986: 146; Borkent & Spinelli, 2007: 73; Spinelli et al., 2009: 90. gabaldoni Ortíz, 1954: 221. Type locality: Venezuela, Yaracuy, San Felipe. HT F (INHC). Distr.: Mexico (Tabasco) to Colombia (Valle del Cauca (Río Raposo)), Ecuador, Venezuela, Trinidad and Tobago, Brazil (Bahia), Paraguay, northeastern Argentina. Refs.: Ortíz, 1954: 221; Barreto, 1986: 146; Borkent & Spinelli, 2007: 73; Spinelli et al., 2009: 90. glabellus Wirth & Blanton, 1956c: 47. Type locality: Panama, Bocas del Toro, Almirante. HT F (# 63129 USNM). Distr.: Honduras to Colombia (Putumayo (Puerto Leguizamo), Valle del Cauca (Río Raposo)), Ecuador, Trinidad and Tobago, Brazil (Bahia, Pará). Refs.: Wirth & Blanton, 1956c: 47; Wirth & Blanton, 1973: 437; Barreto, 1986: 147; Borkent & Spinelli, 2007: 73; Spinelli et al., 2009: 90. leoni Barbosa, 1952: 17 (1953: 19). Type locality: Ecuador, Santo Domingo. HT F (UNCE). Distr.: Colombia (Boyacá (Pauna, Topo Grande; Aguasal)), Ecuador. Refs. : Barbosa, 1952, 17 (1953: 19); Borkent & Spinelli, 2007: 73; Spinelli et al., 2009: 87. Subgenus unplaced, limai Species Group galindoi Wirth & Blanton, 1953a: 73. Type locality: Panama, Canal Zone, Mojinga Swamp. HT F (# 61501 USNM). Distr.: Costa Rica, Panama, Colombia (Valle del Cauca (Río Raposo)). Refs.: Wirth & Blanton, 1953a: 73; Barreto, 1986: 147; Borkent & Spinelli, 2007: 73; Spinelli et al., 2009: 87; Borkent, 2015: 85 (cat.). limai Barretto, 1944: 99. Type locality, Brazil, São Paulo, Mojí das Cruzes. HT M (DEFS). Distr.: El Salvador to Colombia (Meta (Restrepo, El Retiro)), Ecuador, Brazil (Pará, Mato Grosso, São Paulo, Rio de Janeiro, Santa Catarina) to northeastern Argentina. Refs.: Barretto, 1944: 99; Forattini, 1954: 223; Forattini, 1957: 335; Barreto, 1986: 147; Borkent & Spinelli, 2007: 73; Spinelli et al., 2009: 90. santanderi Browne, 1980: 536. Type locality: Colombia, Valle del Cauca, Cali, Pance, Finca Topacio, 1600 m. HT F (USNM). Distr.: Colombia (Valle del Cauca (Cali, Pance, Finca Topacio, 1600 m)). Refs.: Browne, 1980: 536; Barreto, 1986: 148; Wirth et al., 1988: 34; Borkent & Spinelli, 2007: 73; Spinelli et al., 2009: 90; Borkent, 2015: 101 (cat.). vernoni Wirth & Blanton, 1973: 448. Type locality: Brazil, Pará, Belém. HT F (USNM). Distr.: Costa Rica, Colombia (Valle del Cauca (Cali, Peñas Blancas, El Rosal 1850 m; Río Raposo)), Bolivia, Brazil (Pará). Refs.: Wirth & Blanton, 1973: 448; Barreto, 1986: 148; Borkent & Spinelli, 2007: 73; Spinelli et al., 2009: 90. Subgenus unplaced, monticola Species Group andinus Wirth & Lee, 1967: 17. Type locality: Colombia, Cauca, El Tambo, Finca Carpentería, 2500 m. HT F (# 69402 USNM). Distr.: Colombia (Cauca (El Tambo, Finca Carpentería, 2500 m; 18 km E Puracé, 3320 m; 28.4 km E Puracé, 3100 m)). Refs.: Wirth & Lee, 1967: 17; Barreto, 1986: 145; Wirth et al., 1988: 32; Borkent & Spinelli, 2007: 73; Spinelli et al., 2009: 90; Borkent, 2015: 76 (cat.). CATALOGUE OF CERATOPOGONIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 117 monticola Wirth & Lee, 1967: 15. Type locality: Colombia, Cauca, El Tambo, Finca Carpentería, 2500 m. HT F (# 69401 USNM). Distr.: Costa Rica, Panama, Colombia (Cauca (El Tambo, Finca Carpentería, 2500 m), Valle del Cauca (Peñas Blancas, 1800 m; Pance)), Ecuador. Refs.: Wirth & Lee, 1967: 15; Barreto, 1986: 147; Wirth et al., 1988: 34; Borkent & Spinelli, 2007: 73; Spinelli et al., 2009: 90; Borkent, 2015: 93 (cat.). pichindensis Browne, 1980: 538. Type locality: Colombia, Valle, Peñas Blancas, 1800 m. HT F (USNM). Subgenus unplaced, pachymerus Species Group caprilesi Fox, 1952: 364. Type locality: Venezuela, Mount Marahuaca. HT F (UNPR). Distr.: Panama, Colombia (Vaupés (Mitú; Caño Cuduyarí)), Venezuela, Brazil (Pará, Mato Grosso). Refs.: Fox, 1952: 364; Marinkelle & Germán, 1970: 107; Wirth & Blanton, 1973: 428; Barreto, 1986: 145; Wirth et al., 1988: 52; Borkent & Spinelli, 2007: 74; Spinelli et al., 2009: 90. kintzi Wirth & Blanton, 1953a: 72. Type locality: Panama, Canal Zone, Fort Sherman, Mojinga Swamp. HT F (# 61498 USNM). obnoxius Fox, 1952: 365. Type locality: Venezuela, Mount Marahuaca. HT F (UNPR). Distr.: Colombia (Antioquía (Río Anorí)), Venezuela. Refs.: Fox, 1952: 365; Wirth et al., 1988: 54; Borkent & Spinelli, 2007: 74; Spinelli et al., 2009: 90. pachymerus Lutz, 1914: 83. Type locality: Brazil, Manaus, Río Negro. ST F (FIOC). Distr.: Guatemala to Colombia (Huila (Neiva), Tolima (Honda)), Brazil (Amazonas). Refs.: Lutz, 1914: 83; Barreto, 1986: 147; Borkent & Spinelli, 2007: 74; Spinelli et al., 2009: 90. uniradialis Wirth & Blanton, 1953a: 70. Type locality: Panama, Canal Zone, Fort Sherman, Mojinga Swamp. HT F (# 61497 USNM). Distr.: Panama, Colombia (Antioquia (Río Anorí), Valle del Cauca (Río Raposo)). Refs.: Wirth & Blanton, 1953a: 70; Wirth & Blanton, 1973: 430; Barreto, 1986: 148; Wirth et al., 1988: 54 Borkent & Spinelli, 2007: 74; Spinelli et al., 2009: 90. Subgenus unplaced, reticulatus Species Group diplus Santarem & Felippe-Bauer, 2014: 261. Type locality: Colombia, Valle, Rio Raposo. HT F (CCER). Distr.: Colombia (Valle del Cauca (Rio Raposo)), Panama. Refs.: Santarem & Felippe-Bauer, 2014: 261; Borkent, 2015: 82 (cat.). macrostigma Wirth & Blanton, 1953b: 230. Type locality: Panama, Canal Zone, Fort Sherman, Mojinga Swamp. HT F (# 61505 USNM). Distr.: Costa Rica to Colombia (?). Refs.: Wirth & Blanton, 1953b: 230; Wirth et al., 1988: 40; Borkent & Spinelli, 2007: 74; Spinelli et al., 2009: 90. paucienfuscatus Barbosa, 1947: 23. Type locality: Brazil, Amazonas, Manaus. HT F (USNM). Distr.: Costa Rica to Peru and Bolivia,Venezuela, Trinidad and Tobago, Brazil (Amazonas, Pará). Localities in Colombia (Caquetá (Tres Esquinas), Meta (Refugio Macarena)). Refs.: Barbosa, 1947: 23; Wirth & Blanton, 1973: 444; Barreto, 1986: 147; Borkent & Spinelli, 2007: 74; Spinelli et al., 2009: 90. pifanoi Ortíz, 1951: 588. Type locality: Venezuela, Maracay, San Felipe. HT F (INHC). Distr.: Belize to Colombia (Valle del Cauca (Río Raposo)), Venezuela, Trinidad and Tobago, Brazil (Pará, Bahia), Paraguay. Refs.: Ortíz, 1951: 588; Barreto, 1986: 147; Borkent & Spinelli, 2007: 74; Spinelli et al., 2009: 90. tricoloratus Wirth & Blanton, 1953b: 233. Type locality: Panama, Panamá Prov., Pacora. HT F (# 61499 USNM). Subgenus unplaced, stigmalis Species Group Refs.: Felippe-Bauer & Wirth, 1987 (key to males and females). fluviatilis (Lutz), 1914: 82 (Johannseniella). Type locality: Brazil, Amazonas, Río Negro, São Gabriel. ST F (FIOC). 118 · Zootaxa 4122 (1) © 2016 Magnolia Press SPINELLI & WOLFF Distr.: Colombia (Vaupés (Mitú; Caño Cuduyarí)), Ecuador, Bolivia, Venezuela, Trinidad and Tobago, Brazil (Amazonas). Refs.: Lutz, 1914: 82; Barreto, 1986: 149; Borkent & Spinelli, 2007: 75; Spinelli et al., 2009: 90. scorzai Ortíz, 1956: 93. Type locality: Venezuela, Bolivar, Río Guayaraca. HT F (INHC). Miscellaneous Unplaced Species of Culicoides arubae Fox & Hoffman, 1944: 109. Type locality: Aruba. HT F (UNPR). Distr.: USA (Texas), Mexico (Yucatán), Aruba and Grenada, to Colombia (Magdalena (?)) and Venezuela. Refs.: Fox & Hoffman, 1944: 109; Wirth & Blanton, 1974: 15; Barreto, 1986: 148; Borkent & Spinelli, 2007: 75; Spinelli et al., 2009: 90. pancensis Browne, 1980: 537. Type locality, Colombia, Valle del Cauca, Cali, Pance, Finca Topacio, 1600 m. HT F (USNM). Distr.: Colombia (Valle del Cauca (Cali, Pance, Finca Topacio, 1600 m). Refs.: Browne, 1980: 537; Barreto, 1986: 149; Wirth et al. 1988: 58; Borkent & Spinelli, 2007: 75; Spinelli et al., 2009: 90; Borkent, 2015: 97 (cat.). Tribe Ceratopogonini Genus Allohelea Kieffer Allohelea Kieffer, 1917b: 364. Type species, Sphaeromias pulchripennis Kieffer (orig. des.). neotropica Wirth, 1991a: 503. Type locality: Jamaica, Westmoreland Parish, Negril, Crystal Waters. HT F (USNM). Distr.: Belize, Panama, Colombia (Valle del Cauca (Río Raposo)), Jamaica. Refs.: Wirth 1991a: 503; Borkent & Spinelli, 2007: 76. Genus Alluaudomyia Kieffer Alluaudomyia Kieffer, 1913: 12. Type species, Alluaudomyia imparunguis Kieffer (mon.). Refs.: Huerta, 1996 (key to males and females from Mexico); Spinelli & Wirth, 1984b (key to males and females). Neoceratopogon Malloch, 1915b: 310. Type species, Ceratopogon bellus Coquillett (orig. des.). Isoecacta Garrett, 1925: 9. Type species, Isoecacta poeyi Garrett (= Ceratopogon bellus Coquillett) (orig. des.). caribbeana Spinelli & Wirth, 1984b: 678. Type locality: Belize, Nattieville. HT F (USNM). Distr.: USA (California to Texas, Florida), Central America, circum-Caribbean from Mexico (Yucatán, Morelos, Sinaloa) to Colombia (Meta (Puerto López)) and Venezuela. Refs.: Spinelli & Wirth, 1984c: 678; Borkent & Spinelli, 2007: 76. distispinulosa Spinelli & Wirth, 1984b: 682. Type locality: Brazil, Amazonas, Manaus. HT F (USNM). Distr.: Colombia (Valle del Cauca (Río Raposo)), Bolivia, Venezuela, Brazil (Amazonas). Refs.: Spinelli & Wirth, 1984b: 682; Borkent & Spinelli, 2007: 76. leei Spinelli & Wirth, 1984b: 687. Type locality: Colombia, Meta, Finca Barbascal. HT F (USNM). Distr.: Colombia (Meta (Finca Barbascal)), Bolivia. Refs.: Spinelli & Wirth, 1984b: 687; Borkent & Spinelli 2007: 76; Borkent, 2015: 111 (cat.). nubeculosa Spinelli & Wirth, 1984b: 689. Type locality: Brazil, Amazonas, Río Solimões. HT F (USNM). Distr.: Colombia (Valle del Cauca (Río Raposo)), Brazil (Amazonas, Pará). Refs.: Spinelli & Wirth, 1984b: 689; Borkent & Spinelli, 2007: 76. sexpunctata Spinelli & Wirth, 1984b: 695. Type locality: Colombia, Valle del Cauca (Río Raposo). HT F CATALOGUE OF CERATOPOGONIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 119 (USNM]). Distr.: Colombia (Valle del Cauca (Río Raposo)). Refs.: Spinelli & Wirth, 1984b: 695; Borkent & Spinelli, 2007: 77; Borkent, 2015: 113 (cat.). spinellii Wirth & Grogan, 1988: 21. New name for tripunctata. Distr.: Colombia (Valle del Cauca (Río Raposo)). Refs.: Wirth & Grogan, 1988: 21; Borkent & Spinelli, 2007: 77; Borkent, 2015: 113 (cat.). tripunctata Spinelli & Wirth, 1984b: 697 (preocc. Alluaudomyia tripunctata Chaudhuri, Das Gupta & Chaudhuri, 1972). Type locality: Colombia, Valle del Cauca (Río Raposo). HT F (USNM). youngi Spinelli & Wirth, 1984b: 699. Type locality: Colombia, Antioquia (near Río Anorí). HT F (USNM). Distr.: Panama, Colombia (Antioquia, (near río Anorí)). Refs.: Spinelli & Wirth, 1984b: 699; Borkent & Spinelli, 2007: 77; Borkent, 2015: 114 (cat.). Genus Baeohelea Wirth & Blanton Baeohelea Wirth & Blanton, 1970b: 95. Type species, Baeohelea nana Wirth & Blanton (orig. des.). nana Wirth & Blanton, 1970b: 95. Type locality: Dominica, Pont Casse. HT F (# 70647 USNM). Distr.: Costa Rica to Ecuador, St. Kitts, Dominica. Locality in Colombia (Valle del Cauca (Río Raposo)). Refs.: Wirth & Blanton, 1970b: 95; Borkent & Spinelli, 2007: 78. Genus Brachypogon Kieffer Brachypogon Kieffer, 1899: 69. Type species, Ceratopogon vitiosus Winnertz (orig. des.). Refs.: Spinelli & Wirth, 1993 (key to subgenera as part of larger key to genera). Subgenus Brachypogon Kieffer Refs.: Spinelli & Grogan, 1998 (key to males and females). bimaculatus Spinelli & Grogan, 1998: 67. Type locality: Colombia, Valle del Cauca (Río Raposo). HT M (FSCA). Distr.: Mexico (Yucatán), Colombia (Valle del Cauca (Río Raposo)), Peru, Brazil (Mato Grosso). Refs.: Spinelli & Grogan, 1998: 67; Borkent & Spinelli, 2007: 78; Borkent, 2015: 116 (cat.). ecuadorensis Spinelli & Grogan, 1998: 68. Type locality: Ecuador, Loja, Catamayo. HT M (USNM). Distr.: Belize, Colombia (Valle del Cauca (Río Raposo)), Costa Rica, Ecuador, Jamaica, Panama. Refs.: Spinelli & Grogan, 1998: 68; Borkent & Spinelli, 2007: 78. impar (Johannsen), 1938: 223 (Ceratopogon). Type locality: Puerto Rico, El Yunque trial. HT F (CUIC). Distr.: Colombia (Meta (Finca Barbascal)), Venezuela, Puerto Rico, Brazil (Santa Catarina). Refs.: Johannsen, 1938: 223; Spinelli & Grogan, 1998: 69; Borkent & Spinelli, 2007: 79. monicae Spinelli & Grogan, 1998: 70. Type locality: Colombia, Valle del Cauca (Río Raposo). HT M (USNM). Distr.: Colombia (Valle del Cauca (Río Raposo)). Refs.: Spinelli & Grogan, 1998: 70; Borkent & Spinelli, 2007: 79; Borkent, 2015: 117 (cat.). Subgenus Isohelea Kieffer Isohelea Kieffer, 1917b: 295. Type species, Ceratopogon lacteipennis Zetterstedt (misidentified, =Psilohelea sociabilis Goetghebuer) (orig. des.). Refs.: Spinelli & Grogan, 1994 (key to males and females). hugoi Spinelli & Grogan, 1994: 2. Type locality: Colombia, Nariño, 20 km E Pasto. HT F (CASC). Distr.: Colombia (Nariño (20 km E Pasto)). Refs.: Spinelli & Grogan, 1994: 2; Borkent & Spinelli, 2007: 79; Borkent, 2015: 118 (cat.). 120 · Zootaxa 4122 (1) © 2016 Magnolia Press SPINELLI & WOLFF Genus Downeshelea Wirth & Grogan Downeshelea Wirth & Grogan, 1988: 50. Type species, Monohelea stonei Wirth (orig. des.). Refs.: Lane & Wirth, 1964 (key to males and females). chiapasi (Lane & Wirth), 1964: 219 (Monohelea). Type locality: Nicaragua, Villa Somoza. HT M (# 66348 USNM). Distr.: Nicaragua to Colombia (Valle del Cauca (Río Raposo)). Refs.: Lane & Wirth, 1964: 219; Borkent & Spinelli, 2007: 80. colombiae (Lane & Wirth), 1964: 220 (Monoheleae). Type locality: Colombia, Valle del Cauca (Río Raposo). HT M (# 67563 USNM). Distr.: Colombia (Valle del Cauca (Río Raposo)). Refs.: Lane & Wirth, 1964: 220; Borkent & Spinelli, 2007: 80; Borkent, 2015: 123 (cat.). fuscipennis (Lane & Wirth), 1964: 221 (Monohelea). Type locality: Colombia, Valle del Cauca (Río Raposo). HT F (# 67564 USNM). Distr.: Colombia (Valle del Cauca (Río Raposo)). Refs.: Lane & Wirth, 1964: 221; Borkent & Spinelli, 2007: 80; Borkent, 2015: 123 (cat.). grogani Huerta, Felippe-Bauer & Spinelli, 2012: 64. Type locality: Mexico, Oaxaca, Municipio San Juan Guichicovi, Localidad El Zacatal. HT M (CAIM). Distr.: Mexico (Veracruz, Oaxaca, Quintana Roo), Belize and Colombia (Valle del Cauca (Río Raposo)). Refs.: Huerta et al., 2012: 64. stonei (Wirth), 1953: 148 (Monohelea). Type locality: USA, Louisiana, Baton Rouge. HT M (# 61094 USNM). Distr.: Eastern North America (Ontario, Massachusetts to Florida), Caribbean to Colombia (Valle del Cauca (Río Raposo)) and Trinidad and Tobago. Refs.: Wirth, 1953: 148; Lane & Wirth, 1964: 222; Borkent & Spinelli, 2007: 81. Genus Echinohelea Macfie Echinohelea Macfie, 1940b: 187. Type species: Echinohelea ornatipennis Macfie (orig. des.). Refs.: Wirth, 1994 (key to males and females). Subgenus Echinohelea Macfie lanei Wirth, 1951b: 319. Type locality: USA, Virginia, Falls Church. HT M (# 60973 USNM). Distr.: USA (Virginia, Michigan and Massachetts to Florida), Panama, Colombia ((Valle del Cauca (Río Raposo; Río Micay, Casa de Suárez)), Trinidad and Tobago, Brazil (Pará, Rondônia). Refs.: Wirth, 1951b: 319; Wirth, 1994: 234; Borkent & Spinelli, 2007: 81. leei Wirth, 1994: 237. Type locality: Colombia, Valle del Cauca (Río Raposo). HT F (USNM). Distr.: Colombia (Valle del Cauca (Río Raposo)). Refs.: Wirth, 1994: 237; Borkent & Spinelli, 2007: 81; Borkent, 2015: 124 (cat.). neotropica Wirth, 1994: 237. Type locality: Colombia, Valle del Cauca (Río Raposo). HT F (USNM). Distr.: Panama, Colombia (Antioquia (near río Anorí), Chocó (Camp Teresita, Río Truando), Valle del Cauca (Río Micay, Casa de Suárez; Rio Raposo)), Ecuador, Brazil (Amazonas, Pará, Rio de Janeiro). Refs.. Wirth, 1994: 237; Borkent & Spinelli, 2007: 81; Borkent, 2015:124 (cat.). smarti Macfie, 1940b: 190. Type locality: Guyana, Mazaruni. HS F (BMNH). Distr.: Colombia (Valle del Cauca (Río Raposo)), Guyana. Refs.: Macfie, 1940b: 190; Wirth, 1994: 241; Borkent & Spinelli, 2007: 81. Genus Leptohelea Wirth & Blanton Leptohelea Wirth & Blanton, 1970a: 12. Type species, Leptohelea micronyx Wirth & Blanton (orig. des.). CATALOGUE OF CERATOPOGONIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 121 micronyx Wirth & Blanton, 1970a: 12. Type locality: Colombia, Valle del Cauca (Río Raposo). HT F (# 70652 USNM). Distr.: Costa Rica, Colombia (Valle del Cauca (Río Raposo)). Refs.: Wirth & Blanton, 1970a: 12; Borkent & Spinelli, 2007: 82; Borkent, 2015: 127 (cat.). Genus Monohelea Kieffer Monohelea Kieffer, 1917b: 294. Type species, Monohelea hieroglyphica Kieffer, (orig. des.). Refs.: Lane & Wirth, 1964 (key to males and females). mayeri Ortíz, 1950c: 202. Type locality: Venezuela, Miranda, Ocumare del Tuy. HT F (MHNL). Distr.: Costa Rica to Colombia (Valle del Cauca (Río Raposo)), Venezuela, Trinidad and Tobago. Refs.: Ortiz, 1950c: 202; Lane & Wirth 1964: 230; Borkent & Spinelli, 2007: 83. Genus Nannohelea Grogan & Wirth Nannohelea Grogan & Wirth, 1980: 374. Type species, Ceratopogon bouríoni Clastrier (orig. des.). clastrieri Grogan & Wirth, 1980: 381. Type locality: Colombia, Valle del Cauca (Río Raposo). HT M (# 76099 USNM). Distr.: Colombia (Valle del Cauca (Río Raposo)). Refs.: Grogan & Wirth, 1980: 381; Borkent & Spinelli, 2007: 83; Borkent, 2015: 130 (cat.). Genus Parabezzia Malloch Parabezzia Malloch, 1915a: 358. Type species, Parabezzia petiolata Malloch, (orig. des.). Refs.: Spinelli & Grogan, 1987 (key to females and some males). unguis Wirth, 1965: 228. Type locality: USA, Arizona, Pima County, Sabino Canyon. HT F (# 66669 USNM). Distr.: USA (Arizona, Texas) to Colombia (Valle del Cauca (oleoducto de Pacific, 10 km NW Dagua)). Refs.: Wirth, 1965: 228; Spinelli & Grogan, 1987: 32; Borkent & Spinelli, 2007: 84. Genus Parastilobezzia Wirth & Blanton Parastilobezzia Wirth & Blanton, 1970a: 10. Type species, Parastilobezzia leei Wirth & Blanton (orig. des.). leei Wirth & Blanton, 1970a: 10. Type locality: Colombia, Valle del Cauca (Río Raposo). HT F (# 70651 USNM). Distr.: Costa Rica, Colombia (Valle del Cauca (Río Raposo)). Refs.: Wirth & Blanton, 1970a: 12; Borkent & Spinelli, 2007: 84; Borkent, 2015: 132 (cat.). Genus Stilobezzia Kieffer Stilobezzia Kieffer, 1911a: 118. Type species, Stilobezzia festiva Kieffer (orig. des.). Refs.: Huerta, 1996 (key to males and females from Mexico); Ingram &Macfie, 1931 (key males and females from Patagonia and southern Chile); Lane & Forattini, 1961 (key to males and females from Neotropical Region north of Patagonia and southern Chile); Wirth & Grogan, 1988 (key to subgenera, as part of generic key). Hartomyia Malloch, 1915a: 339. Type species, Ceratopogon pictus Coquillett (= Stilobezzia coquilletti Kieffer) (orig. des.). Subgenus Acanthohelea Kieffer Acanthohelea Kieffer, 1917a: 198. Type species, Acanthohelea pruinosa Kieffer (mon.). 122 · Zootaxa 4122 (1) © 2016 Magnolia Press SPINELLI & WOLFF maia Lane & Forattini, 1958: 204. Type locality: Panama, Prov. Panamá, Almirante, Bocas del Toro. HT M (DEFS). Distr.: Panama, Colombia (?). Refs.: Lane & Forattini, 1958: 204; Borkent & Spinelli, 2007: 85. Tribe Heteromyiini Wirth, 1962 Genus Clinohelea Kieffer Clinohelea Kieffer, 1917b: 295. Type species, Ceratopogon variegatus Winnertz (= Ceratopogon unimaculata Macquart) (orig. des.). Subgenus Ceratobezzia Kieffer Ceratobezzia Kieffer, 1917b: 326. Type species, Ceratobezzia fallax Kieffer (orig. des.). fallax Kieffer, 1917b: 326. Type locality: Paraguay. T F (HNHM, lost). Distr.: El Salvador to Colombia (Meta (Indostán)), Venezuela, Brazil (Amazonas, Pará, Mato Grosso, Sãom Paulo), Paraguay, Northeastern Argentina. Refs.: Kieffer, 1917b: 326; Spinelli & Duret, 1993: 41; Borkent & Spinelli, 2007: 88. barrettoi Lane & Duret, 1954: 249. Type locality: Brazil, São Paulo, Ribeirao Preto. HT F (# 10433 DEFS). Subgenus Clinohelea Kieffer Refs.: Spinelli & Duret, 1993 (key to males and females). horacioi Lane, 1944: 257. Type locality: Brazil, São Paulo, São Roque. HT F (# 4984 DEFS). Distr.: Mexico (Yucatán, Tabasco, Morelos) to northeastern Argentina. Localities in Colombia (Valle del Cauca (Río Raposo), Meta (Indostán), Chocó (Vuelta)). Refs.: Lane, 1944: 257; Spinelli & Duret, 1993: 45; Borkent & Spinelli, 2007: 88. muzoni Spinelli & Duret, 1993: 46. Type locality: Colombia, Valle del Cauca (Río Raposo). HT F (USNM). Distr.: Colombia (Valle del Cauca (Río Raposo), Chocó (Río Sucio)). Refs.: Spinelli & Duret, 1993: 46; Borkent & Spinelli, 2007: 88; Borkent, 2015: 143 (cat.). nigripes Macfie, 1939b: 205. Type locality: Brazil, Santa Catarina, Nova Teutonia. HT F (BMNH). Distr.: Costa Rica to Colombia (Boyacá (Puerto Boyacá), Valle del Cauca (Río Raposo)), Brazil (São Paulo, Santa Catarina), northeastern Argentina. Refs.: Macfie, 1939b: 205; Spinelli & Duret, 1993: 47; Borkent & Spinelli, 2007: 88. Genus Heteromyia Say Heteromyia Say, 1825: plate 35. Type species, Heteromyia fasciata Say (mon.). Refs.: Duret & Lane, 1955 (key to males and females). Pachyleptus Walker, 1856: 426. Type species, Pachyleptus fasciatus Walker (= Heteromyia nigra Kieffer) (mon.). clavata Williston, 1900: 225 (female; Mexico). Type locality: Mexico, Veracruz. HT F (BMNH). Distr.: Mexico (Veracruz) to Colombia (Atlántico (Barranquilla)), Brazil (Amazonas), northeastern Argentina. Refs.: Williston, 1900: 225; Borkent & Spinelli, 2007: 89. rufa Kieffer, 1917b: 325. Type locality: Colombia (Barranquilla). T F (HNHM, lost). caloptera Kieffer, 1919: 192. Unnecessary new name for H. rufa Kieffer, not Loew, 1861. Tribe Sphaeromiini Newman, 1834 Genus Pellucidomyia Macfie CATALOGUE OF CERATOPOGONIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 123 Pellucidomyia Macfie, 1939a: 99. Type species, Pellucidomyia ugandae Macfie (orig. des.). Refs.: Wirth, 1960 (World key to males and females). Macfiehelea Lane, 1946: 208. Type species, Macfiehelea oliveirai Lane (orig. des.). lanei Wirth & Ratanaworabhan, 1971: 446. Type locality: Colombia, Valle del Cauca (Río Raposo). HT F (# 70678 USNM). Distr.: Colombia (Valle del Cauca (Río Raposo)), Jamaica. Refs.: Wirth & Ratanaworabhan, 1971: 446; Borkent & Spinelli, 2007: 89; Borkent, 2015: 145 (cat.). wirthi (Lane), 1956b: 437 (Macfiehelea). Type locality: Panama, Canal Zone, Mindi Dairy. HT F (USNM). Distr.: USA (Texas) to Colombia (Meta (Finca Barbascal)). Refs.: Lane, 1956b: 437; Wirth & Ratanaworabhan, 1971b: 446; Borkent & Spinelli, 2007: 89. Genus Lanehelea Wirth & Blanton Lanehelea Wirth & Blanton, 1972: 433. Type species, Lanehelea leei Wirth and Blanton (orig. des.). leei Wirth & Blanton, 1972: 434. Type locality: Colombia, Valle del Cauca (Río Raposo). HT F (# 71181 USNM). Distr.: Colombia (Valle del Cauca (Río Raposo)). Refs.: Wirth & Blanton,1972: 434; Borkent & Spinelli, 2007: 91; Borkent, 2015: 154 (cat.). spinifemur Wirth & Blanton, 1972: 436. Type locality: Colombia, Valle del Cauca (Río Raposo). HT F (# 71181 USNM). Distr.: Colombia (Valle del Cauca (Río Raposo)). Refs.: Wirth & Blanton, 1972: 436; Borkent & Spinelli, 2007: 91; Borkent, 2015: 154 (cat.). Genus Neobezzia Wirth & Ratanaworabhan Neobezzia Wirth & Ratanaworabhan, 1972a: 477. Type species: Neobezzia clavipes Wirth & Ratanaworabhan (orig. des.). Refs.: Wirth & Ratanaworabhan, 1972a (key to females). amnicola (Macfie), 1940c: 30 (Bezzia). Type locality: Guyana, New River. HT F (BMNH). Distr.: Belize to Colombia (Valle del Cauca (Río Raposo)), Brazil (Amazonas, Rondônia, Mato Grosso, São Paulo). Refs.: Macfie, 1940c: 30; Wirth & Ratanaworabhan, 1972a: 481; 1972a: 434; Borkent & Spinelli, 2007: 91. blantoni Wirth & Ratanaworabhan, 1972a: 482. Type locality: Panama, Gamboa, Canal Zone (Río Agua Salud). HT F (# 71476 USNM). Distr.: Nicaragua to Colombia (Valle del Cauca (Río Raposo)), Brazil (Amazonas, Pará, Rondônia), northeastern Argentina. Refs.: Wirth & Ratanaworabhan, 1972a: 482; Borkent & Spinelli, 2007: 91. Tribe Palpomyiini Enderlein, 1936 Genus Amerohelea Grogan & Wirth Amerohelea Grogan & Wirth, 1981: 1280. Type species, Amerohelea galindoi Grogan & Wirth (orig. des.). Refs.: Grogan & Wirth, 1981 (key to females); Huerta, 1996 (key to females from Mexico). fasciata Grogan & Wirth, 1981: 1283. Type locality: Belize, Augustine. HT F (# 76569 USNM). Distr.: Mexico (Sonora, Morelos, Tabasco, Chiapas) to Colombia (Valle del Cauca (Río Raposo)). Refs.: Grogan & Wirth, 1981: 1283; Borkent & Spinelli, 2007: 92. frontispina (Dow & Turner), 1976: 138 (Bezzia). Type locality: USA, Texas, Gillespie Co., Fredericksburg. HT F (# 76578 USNM). Distr.: USA (California to Texas) south to Colombia (Valle del Cauca (Río Raposo)) and Venezuela. Refs.: Dow & Turner, 1976: 138; Grogan & Wirth, 1981: 1301; Borkent & Spinelli, 2007: 92. 124 · Zootaxa 4122 (1) © 2016 Magnolia Press SPINELLI & WOLFF galindoi Grogan & Wirth, 1981: 1294. Type locality: Colombia Valle del Cauca (Río Raposo). HT F (# 76574 USNM). Distr.: Panama, Colombia (Valle del Cauca (Río Raposo)), Venezuela, Guadeloupe, northern Argentina. Refs.: Grogan & Wirth, 1981: 1294; Borkent & Spinelli, 2007: 92; Borkent, 2015: 157 (cat.). ronderosi Grogan & Wirth, 1981: 1298. Type locality: Colombia, Valle del Cauca (Río Raposo). HT F (# 76576 USNM). Distr.: Colombia (Valle del Cauca (Río Raposo)). Refs.: Grogan & Wirth, 1981: 1298; Borkent & Spinelli, 2007: 92; Borkent, 2015: 157 (cat.). spinellii Grogan & Wirth, 1981: 1292. Type locality: Colombia, Valle del Cauca (Río Micay, Casa de Suárez). HT F (# 76573 USNM). Distr.: Colombia (Valle del Cauca (Río Micay, Casa de Suárez, Río Raposo)). Refs.: Grogan & Wirth, 1981: 1292; Borkent & Spinelli, 2007: 92; Borkent, 2015: 157 (cat.). Genus Bezzia Kieffer Bezzia Kieffer, 1899: 69. Type species, Ceratopogon ornatus Meigen (orig. des.). Refs.: Huerta, 1996 (key to males and females from Mexico); Spinelli & Wirth, 1989a (key to subgenera and species groups for males and females, key to males and females of nobilis and punctipennis groups); Spinelli & Wirth, 1989b (key to females and some males of glabra and brevicornis groups); Spinelli & Wirth, 1990 (key to males and females of gibbera group); Spinelli & Wirth, 1991 (key to males and females of dentifemur and venustula groups). dentifemur Spinelli & Wirth, 1991: 3. Type locality: Colombia, Valle del Cauca (Río Raposo). HT F (USNM). Distr.: Belize, Colombia (Valle del Cauca (Río Raposo)), Trinidad and Tobago, Brazil (Pará). Refs.: Spinelli & Wirth, 1991: 3; Borkent & Spinelli, 2007: 93; Borkent, 2015: 159 (cat.). filiducta Spinelli & Wirth, 1991: 3 (as filiductus). Type locality: Colombia, Valle del Cauca (Río Raposo). HT F (USNM) Distr.: Honduras, Colombia (Valle del Cauca (Río Raposo)). Refs.: Spinelli & Wirth, 1991: 3; Borkent & Spinelli, 2007: 93; Borkent, 2015: 160 (cat.). fusca Spinelli & Wirth, 1991: 9. Type locality: Colombia, Valle del Cauca (Río Raposo). HT F (USNM). Distr.: Honduras, Colombia (Valle del Cauca (Río Raposo)). Refs.: Spinelli & Wirth, 1991: 9; Borkent & Spinelli, 2007: 93; Borkent, 2015: 160 (cat.). grogani Spinelli & Wirth, 1990: 19. Type locality: Panama, Canal Zone, Fort Davis. HT F (USNM). Distr.: Mexico (Morelos), Panama, Colombia (Valle del Cauca (Río Raposo), Meta (Finca Barbascal)). Refs.: Spinelli & Wirth, 1990: 19; Borkent & Spinelli, 2007: 94. jubata Spinelli & Wirth, 1990: 20. Type locality: Colombia, Valle del Cauca (Río Raposo). HT F (USNM). Distr.: Colombia (Antioquia (near Río Anorí), Valle del Cauca (Río Raposo; Bahia, Buenaventura, Bella Vista)). Refs.: Spinelli & Wirth, 1990: 20; Borkent & Spinelli, 2007: 94; Borkent, 2015: 161 (cat.). leei Spinelli & Wirth, 1990: 21. Type locality, Colombia, Valle del Cauca (Río Raposo). HT F (USNM). Distr.: Colombia (Valle del Cauca (Río Raposo)). Refs.: Spinelli & Wirth, 1990: 21; Borkent & Spinelli, 2007: 94; Borkent, 2015: 161 (cat.). megatheca Spinelli & Wirth, 1990: 22. Type locality: Colombia, Valle del Cauca (Río Raposo). HT F (USNM). Distr.: Colombia (Valle del Cauca (Río Raposo)), Brazil (Rondônia). Refs.: Spinelli & Wirth, 1990: 22; Borkent & Spinelli, 2007: 94; Borkent, 2015: 162 (cat.). raposoensis Spinelli & Wirth, 1991: 4. Type locality: Colombia, Valle del Cauca (Río Raposo). HT F (USNM). Distr.: Mexico (Yucatán), Colombia (Valle del Cauca (Río Raposo)). Refs.: Spinelli & Wirth, 1991: 4; Borkent & Spinelli, 2007: 94; Borkent, 2015: 163 (cat.). setigera Spinelli & Wirth, 1990: 24. Type locality: Colombia, Valle del Cauca, Oleoducto Pacífico (10 km NW CATALOGUE OF CERATOPOGONIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 125 Dagua). HT F (USNM). Distr.: El Salvador, Colombia (Antioquia (near Río Anorí), Valle del Cauca (Oleoducto Pacífico (10 NW Dagua; Río Raposo)). Refs.: Spinelli & Wirth, 1990: 24; Borkent & Spinelli, 2007: 94; Borkent, 2015: 164 (cat.). snowi Lane, 1958: 34. Type locality: Guatemala. HT F (# 11769 DEFS). Distr.: Guatemala, Panama, Colombia (Valle del Cauca (Río Raposo)), Trinidad and Tobago, Brazil (Pará). Refs.: Lane, 1958: 34; Spinelli & Wirth, 1991: 10; Borkent & Spinelli, 2007: 94. venustula (Williston), 1896: 278 (Ceratopogon). Type locality: St. Vincent. HT F (BMNH). Distr.: Widespread, from Mexico (Veracruz, Morelos, Chiapas) to Argentina. Localities in Colombia (Valle del Cauca (Oleoducto Pacífico (10 NW Dagua; Río Raposo)). Refs.: Williston, 1896: 278; Spinelli & Wirth, 1991: 7; Borkent & Spinelli, 2007: 94. concoloripes Macfie, 1940c: 31. Type locality: Guyana, New River. HT F (BMNH). Genus Pachyhelea Wirth Pachyhelea Wirth, 1959: 50. Type species, Ceratopogon magnus Coquillett (= Ceratopogon pachymerus Williston) (orig. des.). albidiventris (Kieffer), 1917b: 316 (Sphaeromias). Type locality: Colombia, Atlántico, Barranquilla. T F (HNHM, lost). Distr.: Colombia (Atlántico (Barranquilla)). Refs.: Kieffer, 1917b: 316; Borkent & Spinelli, 2007: 95; Borkent, 2015: 165 (cat.). Genus Palpomyia Meigen Palpomyia Meigen, 1818: 82. Type species, Ceratopogon flavipes Meigen (mon.). Generic name first published in synonymy with Ceratopogon but available under ICZN Code Article 11(e). Refs.: Ingram & Macfie, 1931 (key males and females from Patagonia and southern Chile); Lane, 1960 (key to females). columbiana Kieffer, 1917b: 323. Type locality: Colombia, Magdalena (Sierra San Lorenzo). T F (HNHM, lost). Distr.: Colombia (Magdalena (Sierra San Lorenzo)). Refs.: Kieffer, 1917b: 323; Borkent & Spinelli, 2007: 96; Borkent, 2015: 167 (cat.). hispida Borkent, in Borkent & Wirth, 1997: 132. New name for multispinosa. Distr.: El Salvador to Colombia (Valle del Cauca (Río Raposo)). Refs.: Borkent & Wirth, 1997: 132; Borkent & Spinelli, 2007: 96. multispinosa Spinelli & Grogan, 1989: 6 (preocc. Palpomyia multispinosa (Pierce), 1966). Type locality: El Salvador, San Vicente, Santo Domingo. HT F (USNM). microchela Kieffer, 1917b: 321. Type locality: Colombia, Magdalena (Sierra San Lorenzo). T F (HNHM, lost). Distr.: Colombia (Magdalena (Sierra San Lorenzo)). Refs.: Kieffer, 1917b: 321; Borkent & Spinelli, 2007: 96; Borkent, 2015: 169 (cat.). Genus Phaenobezzia Haeselbarth Phaenobezzia Haeselbarth, 1965: 297. Type species, Probezzia pistiae Ingram & Macfie (orig. des.). Refs.: Spinelli & Wirth, 1986 (key to males and females). ateles (Macfie), 1940a: 78 (Bezzia). Type locality, Brazil, Bahia, Mundo Novo. HT M (BMNH). Distr.: Colombia (Valle del Cauca (Río Raposo)), Ecuador, Brazil (Bahia). Refs.: Macfie, 1940a: 78; Spinelli & Wirth, 1986: 232; Borkent & Spinelli, 2007: 97. astyla Spinelli & Wirth, 1986: 232. Type locality: Colombia, Valle del Cauca (Río Raposo). HT F (USNM). 126 · Zootaxa 4122 (1) © 2016 Magnolia Press SPINELLI & WOLFF Tribe Stenoxenini Coquillett, 1899 Genus Paryphoconus Enderlein Paryphoconus Enderlein, 1912: 57. Type species, Paryphoconus angustipennis Enderlein (orig. des.). Refs.: Huerta, 1996 (key to females from Mexico); Spinelli & Wirth, 1984a (key to females). angustipennis Enderlein, 1912: 57. Type locality, Brazil, Santa Catarina. HT F (ZMHB). Distr.: Colombia (Valle del Cauca (Río Raposo)), Peru, Venezuela, Brazil (Goiás, São Paulo, Santa Catarina), northeastern Argentina. Refs.: Enderlein, 1912: 57; Wirth & Ratanaworabhan, 1972b: 1371; Borkent & Spinelli, 2007: 97. anomalicornis Kieffer, 1917b: 333. Type locality: Colombia, Magdalena, Santa Marta and Paraguay (Puerto Max). T F (HNHM, lost). Distr.: Mexico (San Luis Potosí) to Colombia (Magdalena (Santa Marta), Valle del Cauca (Río Raposo)), Venezuela, Brazil (Amazonas), Paraguay, northeastern Argentina. Refs.: Kieffer, 1917b: 333: Wirth & Ratanaworabhan, 1972b: 1372; Borkent & Spinelli, 2007: 97; Borkent, 2015: 173 (cat.). brunneipennis Spinelli & Wirth, 1984a: 890. Type locality: Brazil, Amazonas, Río Solimões. HT F (USNM). Distr.: Colombia (Valle del Cauca (Río Raposo)), Brazil (Amazonas). Refs.: Spinelli & Wirth, 1984a: 890; Borkent & Spinelli, 2007: 98. latipennis Spinelli & Wirth, 1984a: 898. Type locality: Colombia, Valle del Cauca (Río Raposo). HT F (USNM). Distr.: Colombia (Valle del Cauca (Río Raposo)), Venezuela. Refs.: Spinelli & Wirth, 1984a: 898; Borkent & Spinelli, 2007: 98; Borkent, 2015: 173 (cat.). leei Spinelli & Wirth, 1984a: 898. Type locality, Colombia, Valle del Cauca (Río Raposo). HT F (USNM). Distr.: Colombia (Valle del Cauca (Río Raposo)), Venezuela. Refs.: Spinelli & Wirth, 1984a: 898; Borkent & Spinelli, 2007: 98; Borkent, 2015: 173 (cat.). oliveirai Lane, 1956a: 303. Type locality: Brazil, Pará, Cachimbo. HT F (# 11526 FIOC). Distr.: Colombia (Amazonas (Leticia)), Brazil (Amazonas, Pará). Refs.: Lane, 1956a: 303; Spinelli & Wirth, 1984a: 902; Borkent & Spinelli, 2007: 98. subflavus Macfie, 1940c: 23. Type locality: Guyana, New River. HT F (BMHN). Distr.: Colombia (Valle del Cauca (Río Raposo)), Guyana, Brazil (Amazonas, Pará, Rondônia). Refs.: Macfie, 1940c: 23; Wirth & Ratanaworabhan, 1972b: 1376; Borkent & Spinelli, 2007: 98. travassosi Lane, 1956a: 304. Type locality, Brazil, Pará, Cachimbo. HT F (# 11525 FIOC). Acknowledgements To Estrategia de Sostenibilidad 2013-2014 University of Antioquia for the grants (MW). Thanks to Biologist Carolina Henao Sepulveda for her dedication and support to this project. TABLE 1. Ceratopogonidae species recorded from Colombia. Subfamily Tribe Genus Subgenus Leptoconopinae Leptoconops Megaconops Forcipomyiinae Atrichopogon Atrichopogon Forcipomyia Caloforcipomyia Species Group Species Author floridensis Wirth, 1951 auricoma Kieffer, 1917 columbianus Kieffer, 1917 caerulea Saunders, 1957 eukosma Macfie, 1939 ...Continued on next page CATALOGUE OF CERATOPOGONIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 127 TABLE 1. (Continued) Subfamily Tribe Genus Subgenus Species Group Euprojoannisia Forcipomyia argenteola genualis Species Author glauca Macfie, 1934 hatoensis Utmar & Wirth, 1976 hermosa Utmar & Wirth, 1976 remigera Utmar & Wirth, 1976 sabalitensis Utmar & Wirth, 1976 blantoni Soria & Bystrak, 1975 argenteola Macfie, 1939 calatheae Wirth, 1982 genualis (Loew, 1866) harpegonata Wirth & Soria, 1975 squamitibia catarinensis Marino & Spinelli, 2002 Lasiohelea stylifer (Lutz, 1913) Lepidohelea annulatipes Macfie, 1939 convexipenis Wirth & Spinelli, 1993 eusthystyla Wirth & Spinelli, 1993 hobbsi Wirth & Spinelli, 1993 luteigenua Wirth & Spinelli, 1992 antioquiae Clastrier & Wirth, 1995 belemensis Clastrier & Wirth, 1995 colombiana Clastrier & Wirth, 1995 fuliginosa (Meigen, 1818) insignipalpis Macfie, 1949 penultimata Wirth, 1972 raposoensis Clastrier & Wirth, 1995 squamosa Lutz, 1914 valleensis Clastrier & Wirth, 1995 vernoni Clastrier & Wirth, 1995 willistoni Wirth, 1971 aitkeni Microhelea Pedilohelea Pterobosca incubans de Meillon & Wirth, 1980 de Meillon & Wirth, 1980 de Meillon & Wirth, 1980 de Meillon & Wirth, 1980 (Macfie, 1937) Rhynchoforcipom yia brachyrhyncha Wirth & Dow, 1972 messersmithi Wirth & Dow, 1972 puracensis Wirth & Dow, 1972 zeteki Wirth & Dow, 1972 leei Wirth & Ratanaworabhan, 1978 ...Continued on next page raposoi spangleri spilmani Saliohelea 128 · Zootaxa 4122 (1) © 2016 Magnolia Press SPINELLI & WOLFF TABLE 1. (Continued) Subfamily Tribe Genus Subgenus Thyridomyia Warmkea Dasyheleinae Ceratopogoninae Culicoidini Dasyhelea Culicoides Anilomyia Avaritia Cotocripus Diphaomyia Drymodesmyia Haematomyidium Species Group Species Author colombiae Wirth, 1970 jipajapae Wirth, 1970 nana (Macfie, 1939) nodosa Saunders, 1959 aeria Saunders, 1957 galindoi Wirth & Soria, 1980 lesliei Wirth, 1974 louriei (Macfie, 1935) spinosa Saunders, 1957 tuberculata Saunders, 1957 columbiana (Kieffer, 1917) santaemarthae (Kieffer, 1917) ameliae Browne, 1980 covagarciai Ortiz, 1950 efferus Fox, 1952 inermis Spinelli et al., 2009 marshi Wirth & Blanton, 1956 metagonatus Wirth & Blanton, 1956 nigrifemur Spinelli et al., 2009 nigrigenus Wirth & Blanton, 1956 popayanensis Wirth & Lee, 1967 trapidoi Wirth & Barreto, 1978 andicola Wirth & Lee, 1967 orjuelai Wirth & Lee, 1967 puracensis Wirth & Lee, 1967 pusillus Lutz, 1913 suarezi bambusicola Rodriguez & Wirth, 1986 Lutz, 1913 gabrieli Spinelli et al., 2009 raposoensis Wirth & Barreto, 1978 iriartei Fox, 1952 lisicarruni Moncada et al., 2010 marinkellei Wirth & Lee, 1967 mirsae Ortiz, 1953 jamaicensis Edwards, 1922 pilosus Wirth & Blanton, 1959 poikilonotus Macfie, 1948 debilipalpis Lutz, 1913 eldridgei Wirth & Barreto, 1978 filiductus Wirth, 1981 germanus Macfie, 1940 ...Continued on next page CATALOGUE OF CERATOPOGONIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 129 TABLE 1. (Continued) Subfamily Tribe Genus Subgenus Species Group Hoffmania Species Author ginesi Ortiz, 1951 glabrior Macfie, 1940 insinuatus Ortiz & Leon, 1955 paraensis (Goeldi, 1905) quasiparaensis Clastrier, 1971 youngi Wirth & Barreto, 1978 antioquensis Spinelli et al., 2009 batesi Wirth & Blanton, 1973 brownei Spinelli et al., 1993 coutinhoi Barreto, 1944 davidi Spinelli et al., 1993 diabolicus Hoffman, 1925 fernandoi Tavares & Souza, 1979 filarifer Hoffman, 1939 foxi Ortiz, 1950 franklini Spinelli et al., 1993 fusipalpis Wirth & Blanton, 1973 heliconiae Fox & Hoffman, 1944 hylas Macfie, 1940 insignis Lutz, 1913 lutzi Costa-Lima, 1937 ocumarensis Ortiz, 1950 palpalis Macfie, 1948 paraignacioi Spinelli et al., 1993 plaumanni Spinelli et al., 1993 polypori Wirth & Blanton, 1968 pseudodiabolicu Fox, 1946 s ruizi Forattini, 1954 tidwelli Spinelli et al., 1993 trinidadensis Hoffman, 1925 verecundus Macfie, 1948 Macfiella phlebotomus (Williston, 1896) Mataemyia dalessandroi Wirth & Barreto, 1978 dicrourus Wirth & Blanton, 1955 volcanensis Wirth & Blanton, 1959 alahialinus Barbosa, 1952 barbosai Wirth & Blanton, 1953 furens (Poey, 1853) gorgasi Wirth & Blanton, 1953 venezuelensis Ortiz & Mirsa, 1950 acotylus Lutz, 1913 Oecacta Psychophaena unplaced acotylus ...Continued on next page 130 · Zootaxa 4122 (1) © 2016 Magnolia Press SPINELLI & WOLFF TABLE 1. (Continued) Subfamily Tribe Genus Subgenus unpalced unplaced Species Group carpenteri daedalus Species Author carsiomelas Wirth & Blanton, 1955 teretipalpis Wirth & Barreto, 1978 belemensis Wirth & Blanton, 1973 camposi Ortiz & Leon, 1955 beaveri Wirth & Barreto, 1978 crescentis Wirth & Blanton, 1959 daedaloides Wirth & Blanto, 1959 daedalus Macfie, 1948 unplaced dasyophrus dasyophrus Macfie, 1940 unplaced eublepharus caldasi Browne, 1986 unplaced unplaced unplaced unplaced unplaced unplaced fluvialis leoni limai monticola caucaensis Wirth & Lee, 1967 eublepharus Macfie, 1948 florenciae Messersmith, 1972 micayaensis Spinelli et al., 2009 pabloi Browne, 1980 rangeli Ortiz & Mirsa, 1952 tamboensis Wirth & Lee, 1967 balsapambensis Ortiz & leon, 1955 castillae Fox, 1946 fluvialis Macfie, 1940 leopoldoi Ortiz, 1951 fieldi Wirth & Blanton, 1956 gabaldoni Ortiz, 1954 glabellus Wirth & Blanton, 1956 leoni Barbosa, 1952 galindoi Wirth & Blanton, 1953 limai barretto, 1944 santanderi Browne, 1980 vernoni Wirth & Blanton, 1973 andinus Wirth & Lee, 1967 monticola Wirth & Lee, 1967 pachymerus caprilesi Fox, 1952 obnoxius Fox, 1952 pachymerus Lutz, 1914 reticulatus uniradialis Wirth & Blanton, 1953 diplus Santarem & FelippeBauer, 2014 Wirth & Blanton, 1953 macrostigma paucienfuscatus Barbosa, 1947 miscelaneous unplaced pifanoi Ortiz, 1951 reticulatus Lutz, 1913 arubae Fox & Hoffman, 1944 ...Continued on next page CATALOGUE OF CERATOPOGONIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 131 TABLE 1. (Continued) Subfamily Tribe Genus Subgenus Species Author pancensis Browne, 1980 neotropica Wirth, 1991 caribbeana Spinelli & Wirth, 1984 distispinulosa Spinelli & Wirth, 1984 leei Spinelli & Wirth, 1984 nubeculosa Spinelli & Wirth, 1984 sexpunctata Spinelli & Wirth, 1984 spinellii Wirth & Grogan, 1988 youngi Spinelli & Wirth, 1984 nana Wirth & Blanton, 1970 bimaculatus Spinelli & Grogan, 1998 ecuadorensis Spinelli & Grogan, 1998 impar (Johansen, 1938) monicae Spinelli & Grogan, 1998 hugoi Spinelli & Grogan, 1998 chiapasi (Lane & Wirth, 1964) colombiae (Lane & Wirth, 1964) fuscipennis (Lane & Wirth, 1964) grogani Huerta et al., 2012 stonei (Wirth, 1953) lanei Wirth, 1951 leei Wirth, 1994 neotropica Wirth, 1994 smarti Macfie, 1940 Leptohelea micronyx Wirth & Blanton, 1970 Monohelea mayeri Ortiz, 1950 Nannohelea clastrieri Grogan & Wirth, 1980 Ceratopogoni Allohelea ni Alluaudomyia Baeohelea Brachypogon Brachypogon Isohelea Downeshelea Echinohelea Echinohelea Species Group Parabezzia unguis Wirth, 1965 Parastilobezzi a Stilobezzia Acanthohelea leei Wirth & Blandon, 1970 maia Lane & Forattini, 1958 fallax kieffer, 1917 Heteromyiini Clinohelea Ceratobezzia Clinohelea Heteromyia Sphaeromiini Pellucidomyia Lanehelea Neobezzia horacioi Lane, 1944 muzoni Spinelli & Duret, 1993 nigripes Macfie, 1939 clavata Williston, 1900 lanei wirthi Wirth & Ratamaworabham, 1971 (Lane, 1956) leei Wirth & Blandon, 1972 spinifemur Wirth & Blandon, 1972 amnicola (Macfie, 1940) ...Continued on next page 132 · Zootaxa 4122 (1) © 2016 Magnolia Press SPINELLI & WOLFF TABLE 1. (Continued) Subfamily Tribe Palpomyiini Genus Subgenus Species Author blantoni fasciata Wirth & Ratamaworabham, 1972 Grogan & Wirth, 1981 frontispina (Dow & Turner, 1976) galindoi Grogan & Wirth, 1981 ronderosi Grogan & Wirth, 1981 spinellii Grogan & Wirth, 1981 dentifemur Spinelli & Wirth, 1991 filiducta Spinelli & Wirth, 1991 fusca Spinelli & Wirth, 1991 grogani Spinelli & Wirth, 1990 jubata Spinelli & Wirth, 1990 leei Spinelli & Wirth, 1990 megatheca Spinelli & Wirth, 1990 raposoensis Spinelli & Wirth, 1991 setigera Spinelli & Wirth, 1990 snowi Lane, 1958 venustula (Williston, 1896) Pachyhelea albidiventris (Kieffer, 1917) Palpomyia columbiana Kieffer, 1917 hispida Borkent, 1997 microchela Kieffer, 1917 Phaenobezzia ateles (Macfie, 1940) Paryphoconus angustipennis Enderlein, 1912 anomalicornis Kieffer, 1917 brunneipennis Spinelli & Wirth, 1984 latipennis Spinelli & Wirth, 1984 leei Spinelli & Wirth, 1984 oliverai Lane, 1956 subflavus Macfie, 1940 Amerohelea Bezzia Stenoxenini Species Group References Aitken, T.H.G., Wirth, W.W., Williams, R.W, Davies, J.B. & Tikasingh, E.S. 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(1955b) Studies in Panama Culicoides (Diptera, Heleidae) V. Descriptions of three new species of the subgenus Oecacta Poey. Bulletin of the Brooklyn Entomological Society, 50, 121–127. Wirth, W.W. & Blanton, F.S. (1956a) A new species of salt-marsh sand fly from Florida, the Bahamas, Panama and Ecuador: its distribution and taxonomic differentiation from Culicoides furens (Poey) (Diptera, Heleidae). Florida Entomologist, 39, 157–162. http://dx.doi.org/10.2307/3492592 Wirth, W.W. & Blanton, F.S. (1956b) Studies in Panama Culicoides VII. The species of the pulicaris and cova-garciai group (Diptera, Heleidae). Proceedings of the Entomological Society of Washington, 58, 211–227. Wirth, W.W. & Blanton, F.S. (1956c) Studies in Panama Culicoides (Diptera, Heleidae). IX. Two new species related to leoni Barbosa and reevesi Wirth. Bulletin of the Brooklyn Entomological Society, 51, 45–52. Wirth, W.W. & Blanton, F.S. (1959) Biting midges of the genus Culicoides from Panama (Diptera: Heleidae). Proceedings of the United States National Museum, 109, 237–482. http://dx.doi.org/10.5479/si.00963801.109-3415.237 140 · Zootaxa 4122 (1) © 2016 Magnolia Press SPINELLI & WOLFF Wirth, W.W. & Blanton, F.S. (1968) A revision of the neotropical biting midges of the hylas group of Culicoides (Diptera: Ceratopogonidae). Florida Entomologist, 51, 201–215. http://dx.doi.org/10.2307/3493420 Wirth, W.W. & Blanton, F.S. (1970a) New genera of Neotropical Ceratopogonidae (Diptera). Florida Entomologist, 53, 7–14. http://dx.doi.org/10.2307/3493107 Wirth, W.W. & Blanton, F.S. (1970b) Notes on Brachypogon Kieffer (Diptera, Ceratopogonidae), a new species, and two new Neotropical genera of the tribe Ceratopogonini. Florida Entomologist, 53, 93–104. http://dx.doi.org/10.2307/3493452 Wirth, W.W. & Blanton, F.S. (1970c) A review of the Culicoides nigrigenus group, with two new species (Diptera: Ceratopogonidae). Entomological News, 81, 141–151. Wirth, W.W. & Blanton, F.S. (1972) Lanehelea, a new neotropical genus of Sphaeromiini (Diptera: Ceratopogonidae). Studia Entomologica, 15, 433–438. Wirth, W.W. & Blanton, F.S. (1973) A review of the maruins or biting midges of the genus Culicoides (Diptera: Ceratopogonidae) in the Amazon Basin. Amazoniana, 4, 405–470. Wirth, W.W. & Blanton, F.S. (1974) The West Indian sandflies of the genus Culicoides (Diptera: Ceratopogonidae). Agricultural Research Service, United States Department of Agriculture, Technical Bulletin, 1474, iv + 98 pp. Wirth, W.W. & Dow, M.I. (1972) Studies on the genus Forcipomyia 4. Rhynchoforcipomyia, a new neotropical subgenus in the Trichohelea complex (Diptera: Ceratopogonidae). Annals of the Entomological Society of America, 65, 862–872. http://dx.doi.org/10.1093/aesa/65.4.862 Wirth, W.W., Dyce, A.L. & Spinelli, G.R. (1988) An atlas of wing photographs, with a summary of the numerical characters of the Neotropical species of Culicoides (Diptera: Ceratopogonidae). Contributions of the American Entomological Institute, 25, 1–72 Wirth, W.W. & Felippe-Bauer, M.L. (1989) The neotropical biting midges related to Culicoides paraensis (Diptera: Ceratopogonidae). Memórias do Instituto Oswaldo Cruz, 84, 551–565. http://dx.doi.org/10.1590/S0074-02761989000800096 Wirth, W.W. & Grogan, W.L. (1988) The predaceous midges of the world (Diptera: Ceratopogonidae; Tribe Ceratopogonini). Flora and Fauna Handbook. 4. E.J. Brill, Leiden, xv + 160 pp. Wirth, W.W. & Hubert, A.A. (1960) Ceratopogonidae (Diptera) reared from cacti, with a review of the copiosus group of Culicoides. Annals of the Entomological Society of America, 53, 639–658. http://dx.doi.org/10.1093/aesa/53.5.639 Wirth, W.W. & Lee, V.H. (1967) New species of Culicoides from high altitudes in the Colombian Andes (Diptera: Ceratopogonidae). Proceedings of the United States National Museum 124, 1–22. http://dx.doi.org/10.5479/si.00963801.124-3626.1 Wirth, W.W. & Ratanaworabhan, N.C. (1971) Notes on Neotropical Pellucidomyia (Diptera, Ceratopogonidae). Annals of the Entomological Society of America, 64, 446–448. http://dx.doi.org/10.1093/aesa/64.2.446 Wirth, W.W. & Ratanaworabhan, N.C. (1972a) Neobezzia, a new neotropical biting midge genus of the tribe Sphaeromiini (Diptera: Ceratopogonidae). Journal of the Kansas Entomological Society, 45, 476–490. Wirth, W.W. & Ratanaworabhan, N.C. (1972b) A revision of the tribe Stenoxenini (Diptera: Ceratopogonidae). Annals of the Entomological Society of America, 65, 1368–1388. http://dx.doi.org/10.1093/aesa/65.6.1368 Wirth, W.W. & Ratanaworabhan, N.C. (1978) Studies on the genus Forcipomyia. V. Key to subgenera and description of a new subgenus related to Euprojoannisia Brèthes (Diptera: Ceratopogonidae). Proceedings of the Entomological Society of Washington, 80, 493–507. Wirth, W.W. & Soria, S.J. (1975) A new Neotropical Forcipomyia midge closely related to F. (F.) genualis (Loew) (Diptera: Ceratopogonidae). Revista Theobroma, 5, 19–27. Wirth, W.W. & Soria, S.J. (1980) Studies on the genus Forcipomyia VI. The Neotropical species of the subgenus Warmkea (Diptera: Ceratopogonidae). Revista Theobroma, 9, 137–161. Wirth, W.W. & Soria, S.J. (1981) Two Culicoides biting midges reared from inflorescences of Calathea in Brazil and Colombia, and a key to the species of the discrepans group (Diptera: Ceratopogonidae). Revista Theobroma, 11, 107–117. Wirth, W.W. & Spinelli, G.R. (1992) Immature stages of Forcipomyia seminole Wirth and a related new neotropical species (Diptera: Ceratopogonidae). Florida Entomologist, 75, 349–356. http://dx.doi.org/10.2307/3495856 Wirth, W.W. & Spinelli, G.R. (1993) The American species of the annulatipes group of the subgenus Lepidohelea, genus Forcipomyia (Diptera: Ceratopogonidae). Insecta Mundi, 6, 109–125. [1992] CATALOGUE OF CERATOPOGONIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 141 Zootaxa 4122 (1): 142–153 http://www.mapress.com/j/zt/ Copyright © 2016 Magnolia Press Article ISSN 1175-5326 (print edition) ZOOTAXA ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.4122.1.16 http://zoobank.org/urn:lsid:zoobank.org:pub:728FB3F8-46D0-4D00-B875-6539604C66D2 FAMILY CHIRONOMIDAE HUMBERTO FONSECA MENDES1 & LUIZ CARLOS PINHO2* 1 Universidade Federal de Alfenas, Instituto de Ciências da Natureza, Rua Gabriel Monteiro da Silva, 700, CEP 37130-000, Alfenas— MG. Brazil. E-mail: orthocladiinae@gmail.com 2 Universidade Federal de Santa Catarina, Campus Trindade, CEP 88040-900, Florianópolis—SC. Brazil. E-mail: luiz.pinho@ufsc.br. *Corresponding Author: Email: luiz.pinho@ufsc.br Abstract Only 30 chironomid species distributed in 16 genera and three subfamilies are formally recorded to Colombia. Another 32 genera and two subfamilies have been recorded on larval stage, with no identification at species level. Many new records, genera and species are expected when focusing on systematics of chironomids from Colombia. Key words: catalogue, Chironomidae, Colombia, Distribution, taxonomy Introduction According to Ashe & O’Connor (2009), there are eleven subfamilies of Chironomidae, ten of which are known to occur in the Neotropical region and the remaining subfamily is recorded from a single species from the Usambara mountains in Tanzania (namely Usambaromyiinae). Among the ten subfamilies from the Neotropical region, one is endemic and known from a single species from Chile (Chilenomyiinae), all the remaining subfamilies are more species-rich and occur in at least two of the major biogeographical regions. There are five subfamilies recorded to Colombia, which are Tanypodinae, Orthocladiinae and Chironominae with described species, and some records of unidentified Podonominae and Diamesinae (Riss & Ospina 2000; Meza et al. 2012; Posada-García et al. 2008), but at least three subfamilies that occur in specialized habitats such as marine, lowland and highland taxa should be expected (namely Telmatogetoninae, Prodiamesinae, Buchonomyinae). The present Catalogue of Chironomidae of Colombia accounts a total of 30 species distributed in 16 genera and three subfamilies as occurring in the country (Table 1). Since the catalog compiled by Spies & Reiss (1996), not much has been added to Colombia. The knowledge of Neotropical chironomid named species increased by 15% in less than 10 years (Spies et al. 2009), but these records are sort of concentrated in Argentina, Brazil, Chile, Costa Rica, Mexico and Venezuela with sporadic new records and species from Ecuador, Honduras, Jamaica, Nicaragua, Puerto Rico and Uruguay. Due to the fact that Colombia has a wide range of altitudes, from sea level up to over 3900 meters above sea level and with shores on both Atlantic and Pacific oceans, the number registered is for sure underestimated. Ecological studies have been recording 32 genera and two subfamilies new to the country (Table 2), which means that at least 62 chironomid species occur in Colombia. The case of Brazilian Orthocladiinae diversity is something that exemplify what may be going on in Colombia soon (where only Cricotopus crassimanus is known): Seven species were recorded in 1996 (Spies & Reiss 1996), 37 in 2007 (Mendes et al. 2007) and 122 nowadays (Mendes & Pinho, 2014). We hope that this Colombian chironomid diversity’s scenario depicted in this catalogue stimulate students to begin researches on Systematics of Neotropical Chironomidae, with focus on its probable megadiverse fauna from Colombia. 142 Accepted by S. Nihei: 16 Apr. 2015; published: 14 Jun. 2016 Acronyms used for the depositories ANSP—Academy of Natural Sciences of Philadelphia, Pennsylvania, U.S.A. CUIC—Cornell University Insect Collection, Ithaca, New York, U.S.A. INPA—Instituto Nacional de Pesquisas da Amazônia, Manaus, Amazonas, Brazil NHMW—Naturhistorisches Museum Wien, Austria Townes—H. K. and M. C. Townes Collection, Takoma Park, Maryland, U.S.A. UFMT—Collection of Federal University of Mato Grosso, Cuiabá, Mato Grosso, Brazil USNM—National Museum of Natural History, Washington D.C., U.S.A. ZSM—Zoologische Staatssammlung München, Germany List of abbreviations cat.—catalogue comb.—combination desc.—description distr.—geographical distribution mon.—monotypy misident.—misidentification orig. desig.—original designation orig. desc.—original description redesc.—redescription refs.—references rev.—revision rec.—record spec.—species subs. des.—subsequent designation HT—Holotype NT—Neotype F (f#)—Female M (m#)—Male Catalogue of Chironomidae of Colombia Family Chironomidae Newman, 1834 Subfamily Tanypodinae Kieffer, 1906 Genus Ablabesmyia Johannsen Ablabesmyia Johannsen, 1905: 135. Type species, Tipula monilis Linnaeus (subs. des. Johanssen, 1907: 400). Refs.: See Ashe & O’Connor, 2009: 112 for synonyms; Spies & Reiss, 1996: 80 (cat.); Oliveira et al. 2013: 5 (rev., key to Neotropical species). Subgenus Sartaia Roback Sartaia Roback, 1983: 236. Type species, Ablabesmyia (Sartaia) metica Roback, 1983 (orig. desig.). Refs.: Ashe & O’Connor, 2009: 124 (cat.); Oliveira et al., 2013: 7 (new spec. key to Neotropical species). metica Roback, 1983: 239. Type locality: “Laguna Mozambique, 16 km SW of Puerto Lopez, Dept. del Meta, Colombia”. HT M (ANSP). Distr.: Brazil, Colombia. Refs.: Spies & Reiss, 1996: 80 (cat.); Oliveira et al., 2008: 62 (redesc. male, desc. immatures, key, new rec.); Ashe & O’Connor, 2009: 124 (cat.); Oliveira et al. 2013: 7 (key). CATALOGUE OF CHIRONOMIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 143 Genus Labrundinia Fittkau Labrundinia Fittkau, 1962: 372. Type species, Tanypus longipalpis Goetghebuer, 1921 (orig. desig.). Refs.: Spies & Reiss, 1996: 81 (cat.); Ashe & O’Connor, 2009: 164 (cat.); Silva et al., 2014 (rev., key). fera Roback, 1987: 218. Type locality: “Laguna Mozambique, 16 km SW of Puerto Lopez, Dept. del Meta, Colombia”. HT M (ANSP). Distr.: Colombia. Refs.: Spies & Reiss, 1996: 81 (cat.); Ashe & O’Connor, 2009: 164 (cat.); Silva et al., 2014: 31 (redesc., key). fosteri Roback, 1987: 218. Type locality: “Laguna Mozambique, 16 km SW of Puerto Lopez, Dept. del Meta, Colombia”. HT M (ANSP). Distr.: Colombia. Refs.: Spies & Reiss, 1996: 81 (cat.); Ashe & O’Connor, 2009: 164 (cat.); Silva et al. 2014: 44 (redesc., key). hirsuta Roback, 1987: 221. Type locality: “Laguna Mozambique, 16 km SW of Puerto Lopez, Dept. del Meta, Colombia”. HT M (ANSP). Distr.: Colombia. Refs.: Spies & Reiss, 1996: 81 (cat.); Ashe & O’Connor, 2009: 165 (cat.); Silva et al., 2014: 51 (redesc., key). meta Roback, 1987: 220. Type locality: “Laguna Mozambique, 16 km SW of Puerto Lopez, Dept. del Meta, Colombia”. HT M (ANSP). Distr.: Colombia. Refs.: Spies & Reiss, 1996: 81 (cat.); Ashe & O’Connor, 2009: 165 (cat.); Silva et al. 2014: 79 (redesc., key). opela Roback, 1987: 218. Type locality: “Laguna Mozambique, 16 km SW of Puerto Lopez, Dept. del Meta, Colombia”. HT M (ANSP). Distr.: Colombia. Refs.: Spies & Reiss, 1996: 81 (cat.); Ashe & O’Connor, 2009: 165 (cat.); Silva et al. 2014: 97 (redesc., key). parabecki Roback, 1987: 215. Type locality: “Laguna Mozambique, 16 km SW of Puerto Lopez, Dept. del Meta, Colombia”. HT M (ANSP). Distr.: Colombia. Refs.: Spies & Reiss, 1996: 81 (cat.); Ashe & O’Connor, 2009: 165 (cat.); Silva et al. 2014: 101 (redesc., key). tenata Roback, 1987: 214. Type locality: “Laguna Mozambique, 16 km SW of Puerto Lopez, Dept. del Meta, Colombia”. HT M (ANSP). Distr.: Colombia. Refs.: Spies & Reiss, 1996: 81 (cat.); Ashe & O’Connor, 2009: 166 (cat.); Silva et al. 2014: 163 (redesc., key). Genus Monopelopia Fittkau Monopelopia Fittkau, 1962: 394. Type species, Tanypus tenuicalcar Kieffer (orig. desig.). Refs.: Spies & Reiss, 1996: 81 (cat.); Serpa-Filho et al., 1997: 47 (new spec.); Mendes et al., 2003: 1 (new spec., key); Ashe & O’Connor, 2009: 179 (cat.); Oliveira et al., 2010: 54 (new spec., key). boliekae Beck & Beck, 1966: 335. Type locality: [U.S.A.] “Leon County, Florida, Boliek’s Pond”. HT M (USNM). Distr.: Canada, U.S.A., Cuba, Colombia. Refs.: Spies & Reiss, 1996: 81 (cat.); Mendes et al., 2003: 2 (key); Ashe & O’Connor, 2009: 179 (cat.); Oliveira et al., 2010: 60 (key). Genus Naelotanypus Roback Naelotanypus Roback, 1982: 135. Type species: Naelotanypus viridis Roback (orig. desig.). Refs.: Spies & Reiss, 1996: 81 (cat.); Ashe & O’Connor, 2009: 181 (cat.). viridis Roback, 1982: 135. Type locality: “Hacienda Mozambique 16 km SW of Puerto Lopez, Dept. del Meta, Colombia”. HT M (ANSP). Distr.: Colombia, Surinam. Refs.: Spies & Reiss, 1996: 81 (cat.); Ashe & O’Connor, 2009: 181 (cat.). 144 · Zootaxa 4122 (1) © 2016 Magnolia Press MENDES & PINHO Genus Procladius Roback Procladius Skuse, 1889: 283. Type species, Procladius paludicola Skuse, 1889 (subs. des. Coquillett 1910:594). Refs: Spies & Reiss, 1996: 82 (cat.); Ashe & O’Connor, 2009: 196 (cat.); Saether, 2010; 36 (new spec., keys, redesc.). Subgenus Procladius Skuse Procladius Skuse, 1889: 283. Type species: Procladius paludicola Skuse, 1889 (subs. des. Coquillett, 1910:594). Refs.: Ashe & O’Connor, 2009: 208 (cat.); Saether, 2010; 36 (keys, redesc.). mozambique Roback, 1982: 124. Type locality: “Hacienda Mozambique 16 km SW of Puerto Lopez, Dept. del Meta, Colombia”. HT M (ANSP). Distr.: Colombia. Refs.: Spies & Reiss, 1996: 82 (cat.); Ashe & O’Connor, 2009: 208 (cat.). Subgenus Psilotanypus Kieffer Psilotanypus Kieffer, 1906: 39. Type species, Tanypus bellus Loew (subs. des. Coquillett, 1910: 597). Refs.: See Ashe & O’Connor, 2009: 210 (cat.) for synonyms Saether, 2010; 36 (keys, redesc.). etatus Roback, 1982: 124. Type locality: “Hacienda Mozambique 16 km SW of Puerto Lopez, Dept. del Meta, Colombia”. HT M (ANSP). Distr.: Colombia, Costa Rica. Refs.: Spies & Reiss, 1996: 82 (cat.); Ashe & O’Connor, 2009: 211 (cat.). stroudi Roback, 1982: 122. Type locality: “Hacienda Mozambique 16 km SW of Puerto Lopez, Dept. del Meta, Colombia”. HT M (ANSP). Distr.: Colombia. Refs.: Spies & Reiss, 1996: 82 (cat.); Ashe & O’Connor, 2009: 213 (cat.). Subfamily Orthocladiinae Kieffer, 1911 Genus Cricotopus Wulp Cricotopus Wulp, 1874: LXX, LXXI. Type species, Chironomus tibialis Meigen, 1804 (subs. des. Coquillett, 1910: 528). Refs.: Ashe & O’Connor, 2012: 205 (cat.)Spies & Reiss, 1996: 76 (cat.). crassimanus (Kieffer, 1917: 362) (Trichocladius). Type locality: “Colombia”. Depository data unavailable. Distr.: Colombia. Refs.: Spies & Reiss, 1996: 76 (cat.). Subfamily Chironominae Macquart, 1838 Genus Chironomus Meigen Chironomus Meigen, 1803: 260. Type species, Tipula plumosa Linnaeus (subs. des. Latreille 1810: 42). Refs.: Ashe, 1983: 16 for synonyms; Reiss, 1974b: 4 (desc. Amazonian species); Trivinho-Strixino & Strixino, 1981: 333 (new spec.); Spies & Reiss, 1996: 69 (cat.); Correia et al., 2005: 30 (new spec.); Correia et al., 2006: 58 (new spec.); Correia & TrivinhoStrixino, 2007: 54 (new spec.). calligraphus Goeldi, 1905: 134. Type locality: [Brazil] “Belém, Pará”. NT M (INPA). Distr.: Refs.: Fittkau, 1965: 220 (redesc., new rec.); Spies & Reiss, 1996: 69 (cat.); Spies et al., 2002: 93 (rev.; new rec.). columbiensis Wülker, Sublette, Morath & Martin, 1989: 123. Type locality: “San Fernando (suburb of Cali, Columbia)”. HT M (ZSM). Distr. Guatemala, Colombia: Refs.: Wülker et al., 1989: 123 (orig. descr.); Spies & Reiss, 1996: 69 (cat.). CATALOGUE OF CHIRONOMIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 145 gigas Reiss, 1974: 4. Type locality: [Brazil] “Tiefe, Lago Muratú”. HT M (INPA). Distr.: Colombia, Brazil: Refs.: Reiss, 1974: 4 (orig. desc.); Spies & Reiss, 1996: 69 (cat.). latistylus Reiss, 1974: 7. Type locality: “Lago Cabaliana bei Manacapurú”, Brazil. HT M (INPA). Distr.: Colombia, Brazil. Refs.: Reiss, 1974: 7 (orig. desc.); Spies & Reiss, 1996: 69 (cat.). Genus Cladopelma Kieffer Cladopelma Kieffer, 1921: 274. Type species, Chironomus virescens Meigen, 1818 (subs. des. Harnisch, 1923: 304). Refs.: See Ashe, 1983: 17 for synonyms; Spies & Reiss, 1996: 69 (cat.). forcipis (Rempel, 1939: 211) (Chironomus). Type locality: [Brazil] “Ac. ‘Doublo’”. HT M (ZSM). Distr.: Mexico, Guatemala, Nicaragua, Costa Rica, Panama, Colombia, Brazil. Refs.: Rempel, 1939: 211 (orig. desc.); Spies & Reiss 1996: 69 (cat.). Genus Dicrotendipes Kieffer Dicrotendipes Kieffer, 1913: 23. Type species, Dicrotendipes pictipennis Kieffer (mon.). Refs.: See Epler, 1988:6 for synonyms; Spies & Reiss, 1996: 70 (cat.). californicus (Johannsen, 1905: 217) (Chironomus). Type locality: [U.S.A.] “Pasadena, California”. HT M (CUIC). Distr.: U.S.A., Mexico, Costa Rica, Panamá, Colombia, Peru, Chile: Refs.: Epler, 1987: 22 (redesc.); Epler, 1988: 61 (distr.); Spies & Reiss, 1996: 70 (cat.). sinoposus Epler, 1987: 152. Type locality: “Mexico, Hidalgo, Otongo”. Distr.: Mexico, Colombia, Brazil Refs.: Epler, 1987: 152 (orig. desc.); Epler, 1988: 64 (distr.); Spies & Reiss, 1996: 70 (cat.). Genus Fissimentum Cranston et Nolte Fissimentum Cranston & Nolte, 1996: 1. Type species, Fissimentum desiccatum Cranston & Nolte (orig. desig.). Refs.: Cranston & Nolte, 1996: 2 (orig. desc.); Spies & Reiss, 1996: 70 (cat.); Pinho & Froehlich, 2011: 145 (new spec., key). desiccatum Cranston & Nolte, 1996: 2. Type locality: “Brazil, Mato Grosso, Rio Bento Gomes”. HT M (UFMT). Distr.: Colombia, Brazil. Refs.: Cranston & Nolte, 1996: 4 (orig. desc.); Spies & Reiss, 1996: 70 (cat.); Nazarova et al., 2004: 284 (new rec. to Colombia); Pinho & Froehlich, 2011: 145 (key). Genus Goeldichironomus Fittkau Goeldichironomus Fittkau, 1965: 210. Type species, Chironomus holoprasinus Goeldi (orig. desig.). Refs.: See Ashe, 1983: 25 for synonyms; Fittkau, 1968: 260 (new spec., as Siolimyia); Reiss, 1974a: 86 (new spec.); Contreras-Lichtenberg, 1982: 175 (comb.); Trivinho-Strixino & Strixino, 1991: 593 (new spec.); Spies & Reiss, 1996: 70 (cat.).; Trivinho-Strixino & Strixino, 1998: 271 (new spec.); Trivinho-Strixino & Strixino, 2005: 441 (new spec.). carus (Townes, 1945: 118) (Tendipes). Type locality: “La Mucuy, Merida, Venezuela”. HT M (Townes). Distr.: Nicaragua, Costa Rica, Panamá, Venezuela, Colombia, Brazil. Refs.: Townes, 1945: 118 (orig. desc., as Tendipes (Tendipes) carus); Lichtenberg, 1979: 93 (desc. pupa, as Goeldichironomus pseudopictus); Contreras-Lichtenberg, 1982: 175 for synonyms and desc. larva; Spies & Reiss, 1996: 70 (cat.); Nazarova et al., 2004: 284 (new rec. to Colombia). devineyae (Beck, 1961) (Chironomus). Type locality: [U.S.A.] “St. James City, Florida”. HT M (USNM). Distr.: U.S.A., Colombia: Refs.: Nazarova et al., 2004: 284 (new rec. to Colombia). 146 · Zootaxa 4122 (1) © 2016 Magnolia Press MENDES & PINHO Genus Oukuriella Epler Oukuriella Epler, 1986: 157. Type species, Oukuriella albistyla Epler (orig. desig.). Refs.: Epler, 1986: 157 (orig. desc.); Epler, 1996: 4 (new spec.); Spies & Reiss, 1996: 71 (cat.); Messias & Fittkau, 1997: 255 (new spec., key); Messias & Oliveira, 1999: 260 (new spec.); Messias et al., 2000: 162 (new spec.); Trivinho-Strixino & Messias, 2005: 285 (new spec.); Fusari et al., 2013: 3 (rev. species associated to freshwater sponges); Fusari et al. 2014: 419 (new spec.). simulatrix Epler, 1986: 160. Type locality: “Colombia, Valle Rio Raposo”. HT M (USNM). Distr. Colombia: Refs.: Epler, 1986: 160 (orig. desc.); Spies & Reiss, 1996: 71 (cat.); Messias & Fittkau, 1997: 255 (key). Genus Parachironomus Lenz Parachironomus Lenz, 1921: 160. Type species, Chironomus cryptomonus Kieffer (subs. desig. Townes, 1945: 152). Refs.: See Ashe, 1983: 39 for synonyms; Spies et al., 1994: 63 (rev. Neotropical species, key); Spies & Reiss, 1996: 71 (cat.). camajura Spies, Fittkau & Reiss, 1994: 69. Type locality: “Colombia, Meta, Mozambique”. HT M (ZSM). Distr.: Colombia, Brazil. Refs.: Spies et al., 1994: 69 (orig. desc., key); Spies & Reiss, 1996: 71 (cat.). cayapo Spies, Fittkau & Reiss, 1994: 70. Type locality: “Colombia, Meta, Mozambique”. HT M (ZSM). Distr.: Costa Rica, Nicaragua, Colombia, Surinam, Brazil. Refs.: Spies et al., 1994: 70 (orig. desc., key); Spies & Reiss, 1996: 71 (cat.). Genus Stenochironomus Kieffer Stenochironomus Kieffer in Kieffer & Thienemann, 1919: 44. Type species, Tipula flexilis Linnaeus (subs. desig. Goetghebuer in Goetghebuer & Lenz: 12) (misident. Tipula gibbus Fabricius). Refs.: Borkent, 1984: 33 (rev., key); Sublette & Sasa, 1994: 51 (dist.); Spies & Reiss, 1996: 73 (cat.); Pinho et al., 2005: 38 (new spec.); Andersen et al., 2008: 2 (new spec.); Dantas et al., 2010: 48 (new spec.); Reis et al., 2013: 450 (new spec., new rec.). palliaculeatus Borkent, 1984: 130. Type locality: [Brazil] “Amazonien, Raum oberer Rio Negro, Rio Marauia, Mission S. Antonio”. HT M (INPA). Distr.: Colombia, Brazil. Refs.: Borkent, 1984: 130 (orig. desc., key); Spies & Reiss, 1996: 73 (cat.). Genus Tanytarsus van der Wulp Tanytarsus van der Wulp, 1874: 134. Type species, Chironomus signatus van der Wulp (subs. des. I.C.Z.N 1961: 361 – Opinion 616). Refs.: See Ashe, 1983: 51 for synonyms; Fittkau & Reiss, 1973: 3 (new spec.); Spies & Reiss, 1996: 73 (cat.); Ekrem & Reiss, 1999: 206 (new spec.); Sanseverino et al., 2002: 453 (new spec.); Sanseverino & Wiedenbrug, 2000: 209 (desc. pupa, new rec.); Trivinho-Strixino & Sanseverino, 2003: 2 (comb., redesc.); Wiedenbrug & OspinaTorres, 2005: 319 (key to pupa); Trivinho-Strixino & Sonoda, 2006: 2 (new spec.); Sanseverino & Fittkau, 2006: 2 (new spec.); Trivinho-Strixino & Strixino, 2007: 62 (new spec.); Vinogradova et al., 2009: 12 (new spec.); Giłka & Zakrzewska, 2013: 454 (new spec.). curvicristatus Contreras-Lichtenberg, 1988: 101. Type locality: [Colombia] “El Cederal, Forellenzuchstation im Bereich des Rio Otún, 2115 m SH”. HT M (NHMW). Distr.: Colombia. Refs.: Contreras-Lichtenberg, 1988: 101 (orig. desc.); Spies & Reiss, 1996: 73 (cat.). Genus Zavreliella Kieffer Zavreliella Kieffer, 1920: 334. Type species, Chironomus clavatricus Kieffer [=Chironomus marmoratus van der Wulp] (orig. desig., mon.). Refs: Reiss, 1990: 85 (rev., key); Spies & Reiss, 1996: 74 (cat.). CATALOGUE OF CHIRONOMIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 147 acuta Reiss, 1990: 96. Type locality: “Surinam, Parmaribo, l’Hermitage”. HT M (ZSM). Distr.: Colombia, Surinam, Brazil. Refs.: Reiss, 1990: 96 (rev.); Spies & Reiss, 1996: 74 (cat.). TABLE 1. Chironomidae species recorded from Colombia, and its described life stages and sexes (M, male; F, female; P, pupa; L, larva). Subfamily Genus Species Author Stages TANYPODINAE Ablabesmyia metica Roback, 1983 M,F,P,L Labrundinia fera Roback, 1987 M fosteri Roback, 1987 M hirsuta Roback, 1987 M meta Roback, 1987 M opela Roback, 1987 M parabechi Roback, 1987 M tenata Roback, 1987 M Monopelopia boliekae Beck & Beck, 1966 M,F,P,L Naelotanypus viridis Roback, 1982 M Procladius mozambique Roback, 1982 M,F etatus Roback, 1982 M stroudi Roback, 1982 M,F ORTHOCLADIINAE Cricotopus crassimanus (Kieffer, 1917) F CHIRONOMINAE Cladopelma forcipis (Rempel, 1939) M Chironomus caligraphus Goeldi, 1905 M,F,P,L columbiensis Wülker et al., 1989 M,L gigas Reiss, 1974 M,P,L latistylus Reiss, 1974 M,P,L californicus (Johannsen, 1905) M sinoposus Epler, 1987 M Fissimentum desiccatum Cranston & Nolte, 1996 M,F,P,L Goeldichironomus carus (Townes, 1945) M,F,P devineyae (Beck, 1961) M,P,L simulatrix Epler, 1986 M Dicrotendipes Oukuriella Parachironomus camajura Spies, Fittkau & Reiss, 1994 M cayapo Spies, Fittkau & Reiss, 1994 M Stenochironomus palliaculeatus Borkent, 1984 M Tanytarsus curvicristatus Contreras-Lichtenberg, 1988 M Zavreliella acuta Reiss, 1990 M,P TABLE 2. Larval records of Chironomidae genera from Colombia. Subfamily Genus Reference PODONOMINAE Podonomus Philippi Ruiz-Moreno et al., 2000a; Meza et al., 2012 Parochlus Enderlein Ruiz-Moreno et al., 2000a; Meza et al., 2012 DIAMESINAE Paraheptagyia Brundin Ruiz-Moreno et al., 2000a; Posada-García et al., 2008 TANYPODINAE Apsectrotanypus Fittkau Riss & Ospina, 2000 ...Continued on next page 148 · Zootaxa 4122 (1) © 2016 Magnolia Press MENDES & PINHO TABLE 2. (Continued) Subfamily ORTHOCLADIINAE CHIRONOMINAE Genus Reference Clinotanypus Kieffer Nazarova et al., 2004 Coelotanypus Kieffer Nazarova et al., 2004 Djalmabatista Fittkau Ruiz-Moreno et al., 2000a Krenopelopia Fittkau Ruiz-Moreno et al., 2000a Pentaneura Philippi Riss & Ospina, 2000 Rheopelopia Fittkau Ruiz-Moreno et al., 2000a Tanypus Meigen Nazarova et al., 2004 Telmatopelopia Fittkau Riss & Ospina, 2000 Zavrelimyia Fittkau Riss & Ospina, 2000 Barbadocladius Cranston & Krosch Prat et al., 2013 Corynoneura Winnertz Riss & Ospina, 2000 Metriocnemus van der Wulp Posada-García et al., 2008 Orthocladius van der Wulp Riss & Ospina, 2000 Parametriocnemus Goetghebuer Meza et al., 2012 Paratrichocladius Santos Abreu Riss & Ospina, 2000 Pseudosmittia Goetghebuer Posada-García et al., 2008 Thienemanniella Kieffer Riss & Ospina, 2000 Apedilum Townes Nazarova et al., 2004 Beardius Reiss & Sublette Nazarova et al., 2004 Caladomyia Säwedal Riss & Ospina, 2000; Sanseverino & Fittkau, 2007 Endotribelos Grodhaus, 1987 Ruiz-Moreno et al., 2000b Lauterborniella Thienemann & Bause Nazarova et al., 2004 Paracladopelma Harnisch Ruiz-Moreno et al., 2000b Paratanytarsus Thienemann et Bause Riss & Ospina, 2000; Ruiz-Moreno et al., 2000b Polypedilum Kieffer Nazarova et al., 2004; Ruiz-Moreno et al., 2000b; Meza et al., 2012 Pseudochironomus Malloch Ruiz-Moreno et al., 2000b Rheotanytarsus Thienemann & Bause Riss & Ospina, 2000; Mesa-S et al., 2012 Xestochironomus Sublette & Wirth Ruiz-Moreno et al., 2000b References Andersen, T., Mendes, H.F. & Pinho, L.C. 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CATALOGUE OF CHIRONOMIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 153 Zootaxa 4122 (1): 154–177 http://www.mapress.com/j/zt/ Copyright © 2016 Magnolia Press Article ISSN 1175-5326 (print edition) ZOOTAXA ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.4122.1.17 http://zoobank.org/urn:lsid:zoobank.org:pub:446C7E51-6B93-470E-A9FD-F815516CFD96 FAMILY SIMULIIDAE MARTA I. WOLFF1, DANIEL RAFAEL MIRANDA-ESQUIVEL2 & LIGIA INES MONCADA-ALVAREZ3 1 Grupo de Entomologia, Instituto de Biología, Universidad de Antioquia, Medellín-Colombia. Laboratorio de Sistematica y Biogeografía, Escuela de Biología, Universidad Industrial de Santander Bucaramanga Colombia. 3 Universidad Nacional de Colombia E-mail: martha.wolff@udea.edu.co; dmiranda@uis.edu.co; limoncadaa@unal.edu.co * Corresponding author: limoncadaa@unal.edu.co 2 Abstract This catalogue presents 70 species distributed in two genera for Colombia: Gigantodax with 5 groups, and Simulium with 6 subgenera and 17 groups. Three species are recorded for the first time: Gigantodax vulcanius, G. gracilis, and Simulium pautense, and the distributional range is expanded to S.ochraceum, S. paynei, S. metallicum, and S. callidum. Key words: Catalogue, distribution, Simuliidae, taxonomy Introduction Adults have a robust appearance, measuring from 1.2 to 5.5 mm in length; black, dark brown or yellow color. Head large, round; ocelli absent, compound eyes attached dorsally in males and separated in females; male have dicoptic eyes and females have holoptic, antennas thick and compacts with 7 to 9 flagellomeres; mouth parts short and thick adapted to cut skin and blood sucking. Thorax big and arched; wings rounded at the base, with thickened anterior veins and subsequent weak; legs strong and short. Abdomen robust. Larvae are characterized by their bottle shape; cylindrical and widened on the back, living mainly in lotic aquatic environments (water currents), from large rivers to streams, submerged and anchored to materials such as rocks, logs, leaves and even artificial substrate, where they feed by filtering particles obtained directly from the water. Pupae have the same ecological habit than the larvae, the body is enclosed in a cocoon; head and thorax forming a cephalothorax, bearing a pair of gills that have multiple forms, from tree-like shaped to globose (Gigantodax-Simuliini); the cocoon and gill forms are important taxonomic characters, that could be used to define even at species level. Most simulids females are hematophagous (females Pedrowygomyia have maxilla and mandible obsolescent), its bloodsucking behavior makes some species responsible for the transmission to humans of the filarial nematode Onchocerca volvulus in Africa, central and northern South America, a disease called river blindness, because the nematode could lodge in the eye and causes blindness. They are known as black-flies, Piuns, Jejenes, Borrachudos and are diurnal, the female by biting leaves a red dot and is very easy to observe while feeding, as it is on the host for a long time. The family is monophyletic (Currie 1988), and most of the analyses divide the family into two monophyletic subfamilies Simuliinae and Parasimiliinae (Currie 1988). The oldest known fossil from the Cretraceous is an undescribed species (Cretaceosimulium Vulcano 1985, nomen nudum). Black flies are known for being involved as an active or mechanic transmitter of some agents; in Colombia they are involved in transmission of two nematodes to humans: Mansonella ozardi (Tidwell et al. 1980; Tidewll & Tidwell 1982) and Onchocerca volvulus (recently eliminated of the country) (WHO 2010; WHO 2011). In addition, these insects have been found as vectors of Leucocytozoon, an avian blood protozoa, in other regions of the world. (Khan & Fallis 1970; Sato et al. 2009; Murdock et al. 2013). In Colombia, the role as vectors of these parasites has been suggested by Matta et al. (2014). 154 Accepted by S. Nihei: 25 May 2015; published: 14 Jun. 2016 The breeding places of blackflies are running water and larvae usually live aggregated in the most oxygenated areas. Taking into account that Colombia is one of the richest countries in the world in water sources, in addition to the fact that the country has three different cordilleras, which give to the territory a great variety of climates, as a consequence a lot of possible breeding places can be found, both in high and in low altitudes. Especially in paramo region, as Díaz et al. (2015) stated that “High mountains of Andes look like water factories”. Larvae are important filter feedings in streams, with critical role in resource turnover in steams (McCreadie & Adler 2012). This article collects the information concerning geographical distribution and altitudinal records of genus Simulium and Gigantodax present in Colombia. These are the unique genera occurring in the country. This catalogue is primarily based upon the monograph of Coscarón, Coscarón Arias & Papavero (2008) and the classification adopted is based on the revision by Adler & Crosskey (2015). The types of species and the museum are not presented. The Colombian departments are underline and specific localities are in brackets. At the end we present a table summarizing all species found in Colombia, including rank and author (Table 1). List of abbreviations aut.—automatic cat.—catalogue des.—designation distr.—geographic distribution i.e.—example inv.—inventory m.—meters mon.—monotypy nom. nov.—nomen novum (new name) orig. des.—by original designation pp.—pages preocc.—preoccupied redescr.—redescription Refs.—references rev.—revision sp., spp.—species ssp.—subspecies subs.—subsequent subg.—subgenus subst.—substituted unav.—unavailable Catalogue of Simuliidae of Colombia Family Simuliidae Newman, 1834 Genus Gigantodax Enderlein Gigantodax Enderlein, 1925: 205. Type species, Gigantodax matebranchia Enderlein, 1925 (orig. des.). Refs.: Wygodzinsky & Coscarón, 1989 (rev.); Coscarón et al., 2008: 9 (cat.); Adler & Crosskey, 2015: 23 (inv.). Archicnesia Enderlein, 1934a: 273. Type species, Simulium (Gigantodax) femineum Edwards, 1931 (orig. des.). CATALOGUE OF SIMULIIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 155 brophyi-species-group chilensis (Philippi), 1865: 634. Type locality: unknown, “The probable type is a female deposited in Museo Nacional de Historia Natural of Santiago, Chile, with one label "Corral 1859" and another "Philippi." Wygodzinsky & Coscarón, 1989: 107 Distr.: Colombia (Cundinamarca (Parque Natural Nacional Chingaza, Sector Palacio 04°42'2.6"N 73°50'39.0"W; Monterredondo Sector, Monterredondo stream, 04°37'41.1"N 73°43'27.4"W). Ref: Díaz et al., 2015: 553. rufinotum Edwards, 1931: 140 ( Simulium). multituberculatus Wygodzinsky & Coscarón, 1989: 123. Type locality: Colombia, Cundinamarca, Buenos Aires, near La Siberia, E. of La Calera, 3050m. Distr.: Colombia ((Boyacá (Cerca al costado N. del lago de Tota; Tota quebrada Guachata; Tota Aquitania quebrada Pozos), Cauca (Popayán to Páramo de Puracé road; E. Silvia), Cundinamarca (Zipaquirá to Pacho road; Subachoque; Albán; Páramo de Guasca; E. La Calera; Cachipay; Bogota to Choachí road; Sibaté to Aguadita road; Chibaté; Bogota, Tolima (Páramo de los Valles)), Ecuador. Refs.: Muñoz de Hoyos, 1994: 416; Muñoz de Hoyos, 1995: 609; Muñoz de Hoyos & Miranda-Esquivel, 1997: 302; Coscarón et al., 2008: 78 (cat.); Adler & Crosskey, 2015: 23 (inv.). ortizi Wygodzinsky, 1973a: 246. Type locality: Venezuela, Mérida, near Apartaderos-Santo Domingo road, 34003500m. Distr.: Colombia ((Boyacá (Páramo de Tota; Páramo de Pisba), Cundinamarca (E. Bogotá; río San Francisco; Subachoque; Páramo de Guasca; E. of La Calera; Bogotá to Choachí road; Sibaté; Chipaque; Páramo de Chisacá; 5km S. of Usme; Santa Helena, quebrada El Oso; El Brasil, quebrada Hoya Honda)), Venezuela. Refs.: Torres-Fernández et al., 1991: 256; Muñoz de Hoyos, 1994: 417; Muñoz de Hoyos et al., 1994: 403; Muñoz de Hoyos, 1995: 610; Muñoz de Hoyos & Miranda-Esquivel, 1997: 302; Coscarón et al., 2008: 8 (cat.); Adler & Crosskey, 2015: 23 (inv.). osornorum Muñoz de Hoyos, Martínez, Mejía & Bueno, in Muñoz de Hoyos, 1995: 610. Type locality: Colombia, Cundinamarca, Santa Helena, Quebrada del Oso, Páramo de Chisacá, 3410m. Distr.: Colombia (Cundinamarca (Chisacá)). Refs.: Muñoz de Hoyos & Miranda-Esquivel, 1997: 302; Coscarón et al., 2008: 7 (cat.); Adler & Crosskey, 2015: 23 (inv.). paramorum Wygodzinsky & Coscarón, 1989: 112. Type locality: Colombia, Cundinamarca, Buenos Aires near La Siberia, W. of La Calera, 3050m. Distr.: Colombia (Cundinamarca (Páramo de Guasca; W. La Calera; Sibaté to Aguadita road; Páramo de Chisacá; Usme, río Tunjuelo; Parque Natural Nacional Chingaza, Sector Palacio 04°42'2.6"N 73°50'39.0"W; Monterredondo Sector, Monterredondo stream, 04°37'41.1"N 73°43'27.4"W)). Refs.: Muñoz de Hoyos, 1994: 418; Muñoz de Hoyos, 1995: 614; Muñoz de Hoyos & Miranda-Esquivel, 1997: 302; Coscarón et al., 2008: 7 (cat.); Adler & Crosskey, 2015: 23 (inv.); Díaz et al., 2015: 553. zumbahue Wygodzinsky & Coscarón, 1989: 131. Type locality: Ecuador, Cotopaxi, road from Pujili to Quevedo, río Zumbahua, 2500m. Distr.: Colombia (Tolima (Páramo de los Valles)), Ecuador. Refs.: Muñoz de Hoyos, 1995: 616; Muñoz de Hoyos & Miranda-Esquivel, 1997: 301; Coscarón et al., 2008: 7 (cat.); Adler & Crosskey, 2015: 23 (inv.). cilicinus-species-group arrarteorum Wygodzinsky & Coscarón, 1989: 161. Type locality: Perú, Junín. Distr.: Perú, Colombia (Risaralda (W side of Cordillera Central Andes, Nevado de Santa Isabel)). Ref.: Mantilla et al., 2013: 427. basinflatus Wygodzinsky & Coscarón, 1989: 147. Type locality: Colombia, Cundinamarca, Páramo de Chisacá (or Sumapáz), 3700m. Distr.: Colombia (Cundinamarca (Páramo de Guasca; Bogotá; Choachí; Páramo de Chisacá, río Chisacá; Bogotá to Choachí road; Santa Helena, Quebrada El Oso), Boyacá (Pàramo de Pisba), Risaralda (W. side of Cordillera Central Andes, Nevado de Santa Isabel, Lake Otún 4°47'41.9”N 75º24'29.1”W, 3950m), Tolima (Páramo de Los Valles)), Ecuador. Refs.: Muñoz de Hoyos, 1994: 414; Muñoz de Hoyos et al., 1994: 404; Muñoz 156 · Zootaxa 4122 (1) © 2016 Magnolia Press WOLFF ET AL. de Hoyos, 1995: 616; Muñoz de Hoyos & Miranda-Esquivel, 1997: 302; Coscarón et al., 2008: 7 (cat.); Adler & Crosskey, 2015: 23 (inv.). destitutus Wygodzinsky & Coscarón, 1989: 144. Type locality: Colombia, Cundinamarca, 2km SE. of Albán, 2400m. Distr.: Colombia (Cundinamarca (Albán, Gachetá, Santandercito Zoológico Santacruz)), Venezuela. Refs.: Muñoz de Hoyos, 1994: 416; Muñoz de Hoyos, 1995: 616; Muñoz de Hoyos & Miranda-Esquivel, 1997: 302; Coscarón et al., 2008: 78 (cat.); Adler & Crosskey, 2015: 23 (inv.). cormonsi-species-group brevis Wygodzinsky & Coscarón, 1989: 183. Type locality: Colombia, Cundinamarca, near road from Usme to Páramo de Chisaca (or Sumapaz), 3300m. Distr.: Colombia (Cundinamarca (La Calera; Bogotá; Choachí; Páramo de Chisacá, río Chisacá; Bogotá to Choachí road; Zipaquirá to Pacho road; E. of La Siberia; Santa Helena Quebrada El Oso; Quebrada Hoya Honda); Boyacá (Páramo de Tota; Páramo de Pisba; Tota, La Punta, quebrada Guachaca). Refs.: Muñoz de Hoyos, 1994: 414; Muñoz de Hoyos et al., 1994: 405; Muñoz de Hoyos, 1995: 617; Muñoz de Hoyos & Miranda-Esquivel, 1997: 302; Coscarón et al., 2008: 8 (cat.); Adler & Crosskey, 2015: 24 (inv.). gracilis Wygodzinsky & Coscarón, 1989: 175. Type locality: Ecuador, Pichincha, Quito-Papallacta road, 3300m. Colombia (Nariño (Pupiales Quebrada Agua-Caliente; 0° 54´ N, 77° 39´W)). New record to Colombia. leonorum Wygodzinsky & Coscarón, 1989: 193. Type locality: Ecuador, Pichincha, Quito-Papallacta road, 3300m. Distr.: Colombia (Boyacá (Sogamoso), Valle del Cauca), Ecuador. Refs.: Muñoz de Hoyos, 1994: 416; Muñoz de Hoyos, 1995: 617; Coscarón et al., 2008: 8 (cat.); Adler & Crosskey, 2015: 24 (inv.). misitu Wygodzinsky & Coscarón, 1989: 179. Type locality: Ecuador, Cotopaxi, NW. slope of Mt. Cotopaxi, 3500m. Distr.: Colombia (Cauca (Popayán to Páramo de Puracé road), Cundinamarca (Bogotá to Choachí road; Páramo de Chisacá; río Chicasá), Risaralda (W. side of Cordillera Central Andes, Nevado de Santa Isabel, Lake Otún (4°47'41.9”N 75º24'29.1”W), 3950m), Tolima (Páramo de los Valles)), Ecuador. Refs.: Muñoz de Hoyos, 1994: 416; Muñoz de Hoyos et al., 1994: 406; Muñoz de Hoyos, 1995: 617; Muñoz de Hoyos & Miranda-Esquivel, 1997: 302; Coscarón et al., 2008: 8 (cat.); Mantilla et al, 2013: 427; Adler & Crosskey, 2015: 24 (inv.). siberianus Wygodzinsky & Coscarón, 1989: 208. Type locality: Colombia, Cundinamarca, Buenos Aires, near La Siberia, W. of La Calera, 3050m. Distr.: Colombia (Cundinamarca (Buenos Aires near La Siberia, W. La Calera; Bogotá to Choachí road; Páramo de Chisacá; río Chisacá)). Refs.: Muñoz de Hoyos, 1994: 418; Muñoz de Hoyos et al., 1994: 406; Muñoz de Hoyos, 1995: 618, Muñoz de Hoyos & Miranda-Esquivel, 1997: 302; Coscarón et al., 2008: 8 (cat.); Adler & Crosskey, 2015: 24 (inv.). vulcanius Wygodzinsky & Coscarón, 1989: 202. Type locality: Ecuador, Pichincha, Quito-Papallacta road, 3300m. Distr. Colombia Nariño (Pupiales, quebrada Agua Caliente 0° 54´ N, 77° 39´W). Ref.: New record to Colombia. vulcanius Takaoka, Hirai & Tada, 1988: 544 (unav., nomen nudum). wygodzinskyi Moncada, Muñoz de Hoyos & Bueno, 1981: 302. Type locality: Colombia, Cundinamarca, Cordillera Oriental, E. of La Calera, río Teusacá. Distr.: Colombia (Boyacá (Páramo de Pisba), Cundinamarca (La Calera, río Teusacá; Buenos Aires near La Siberia E. of La Calera; Subachoque, La Pradera; 2km SE. Albán; La Calera; Bogotá to Choachí road; Sibaté to Aguadita road; Páramo de Chisacá; Usme to Páramo de Chisacá road), Tolima (Páramo de los Valles), Norte de Santander (Pamplona, río Pamplonita). Refs.: Wygodzinsky & Coscarón, 1989: 187; Muñoz de Hoyos, 1994: 419; Muñoz de Hoyos, 1995: 618; Muñoz de Hoyos & Miranda-Esquivel, 1997: 302; Coscarón et al., 2008: 8 (cat.), Adler & Crosskey, 2015: 24 (inv.). wygozinsky incorrect spelling CATALOGUE OF SIMULIIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 157 multifilis-species-group multifilis Wygodzinsky & Coscarón, 1989: 63. Type locality: Ecuador, Pichincha, highway from Aloag to Santo Domingo de los Colorados, western slope of mountain range, 2650m. Distr.: Colombia (Tolima (Páramo de los Valles)), Ecuador. Refs.: Muñoz de Hoyos, 1995: 618; Muñoz de Hoyos & Miranda-Esquivel, 1997: 302; Coscarón et al., 2008: 9 (cat.); Adler & Crosskey, 2015: 24 (inv.). wrighti-species-group cervicornis Wygodzinsky, 1973b: 245 (as cervicorne). Type locality: Colombia, Cundinamarca, 2km SE. of Albán, 2400m. Distr.: Colombia (Cauca (Popayán to Páramo de Puracé road), Cundinamarca (Albán; Páramo de Guasca; La Calera; Ubaque; Sibaté to Aguadita road; Chipaque), Quindío (Ibagué to Calarcá road), Norte de Santander (Pamplona); Tolima (Páramo de Los Valles)), Ecuador, Venezuela. Refs.: Wygodzinsky & Coscarón, 1989: 226; Muñoz de Hoyos, 1994: 416; Muñoz de Hoyos, 1995: 619; Muñoz de Hoyos & Miranda-Esquivel, 1997: 302; Coscarón et al., 2008: 9 (cat.); Adler & Crosskey, 2015: 24 (inv.). cypellus Wygodzinsky & Coscarón, 1989: 262. Type locality: Ecuador, Cotopaxi, río Zumbahua, road from Pujili to Quevedo, 3500m. Distr.: Colombia (Cauca (Puracé to La Plata road, quebrada Cocuy), Ecuador. Refs.: Muñoz de Hoyos, 1994: 416; Muñoz de Hoyos, 1995: 619; Coscarón et al., 2008: 9 (cat.); Adler & Crosskey, 2015: 24 (inv.). cypellus Takaoka et al., 1988: 544 (unav., nomen nudum) nasutus Wygodzinsky & Coscarón, 1989: 240. Type locality: Colombia, Cundinamarca, Alto de San Miguel, on road from Sibate to Fusagasugá, 2800m. Distr.: Colombia (Cundinamarca (Sibaté; Páramo de Chisacá, río Chisacá; Alto de San Miguel)). Refs.: Muñoz de Hoyos, 1994: 417; Muñoz de Hoyos et al., 1994: 406; Muñoz de Hoyos, 1995: 619; Muñoz de Hoyos & Miranda-Esquivel, 1997: 302; Coscarón et al., 2008: 9 (cat.); Adler & Crosskey, 2015: 25 (inv.). septenarius Wygodzinsky & Coscarón, 1989: 220. Type locality: Colombia, Cundinamarca, 2km SW. of Albán, 2400m. Distr. Colombia (Cundinamarca (2km SW. Albán). Refs.: Muñz de Hoyos, 1994: 418; Muñoz de Hoyos, 1995: 619; Muñoz de Hoyos & Miranda-Esquivel, 1997: 302; Coscarón et al., 2008: 10 (cat.); Adler & Crosskey, 2015: 25 (inv.). Genus Simulium Latreille 1802 Subgenus Aspathia Enderlein Aspathia Enderlein, 1935: 359 (as genus). Type species, Simulium hunteri Malloch, 1914 (orig. des.). Refs.: Coscarón et al., 1999: 543-604 (as Simulium (Simulium)) and Coscarón & Coscarón Arias, 2007: 471, 472-483 (keys to spp., based on females, males, pupae and larvae. Jalacingomyia Py-Daniel in Py-Daniel & Moreira Sampaio, 1994: 127. Type species, Simulium anduzei Vargas & Díaz Nájera, 1948: 328 (orig. des.). matteabranchium Anduze 1947: 374. Type locality: Venezuela, Aragua, Colonia Tovar. Distr.: Colombia (Cundinamarca (Albán to Sasaima road; Santandercito Zoológico Santacruz), ?Mexico, Venezuela. Refs.: Coscarón, 1987: 33 (cat.); Muñoz de Hoyos, 1994: 434 (as matteabranchium); Muñoz de Hoyos & MirandaEsqueivel, 1997: 302 (as matteabranchium); Muñoz de Hoyos & Coscarón, 1999: 183; Coscarón et al., 2008: 14 (cat.); Adler & Crosskey, 2015: 31 (inv.). metallicum Bellardi, 1859: 13. Type locality: “Mexico”. Distr.: Colombia (Antioquia (río Medellín; Medellín, vereda Las Palmas), Cauca (Paniquita, 20km N. Popayán; Silvia), Cesar (Donacui; Ariguaní), Cundinamarca 158 · Zootaxa 4122 (1) © 2016 Magnolia Press WOLFF ET AL. (Albán; Albán to Sasaima road; Aguadita; Guaduas; La Vega, río Tabacal; San Francisco, río San Miguel; Sasaima; Guayabal de Siquima, río Siquima; Gachetá; Cachipay; Santandercito, Zoológico Santacruz; Ubaque, río El Palmar; Chipaque to Caquezá road; Choachí to Villavicencio road; Une, Fusagasugá; Pasca), Huila (Pitalito, quebrada Charguayaco; San Agustín), Magdalena (Ariguaní, Donacuí), Quindío (Montenegro to Armenia farm), Tolima (Anaime, Gualanday), Valle del Cauca (río Pance between Saladito to Dagua road; El Queremal; 10km W. of Delfina; Pichindé; río Dagua to río Bitaco farm; Antucela; Cisneros; Anchicayá, Peñas Blancas; El Carmen)), Belize, Brasil, Costa Rica, Ecuador, Guatemala, Mexico, Venezuela. Refs.: Barreto, 1969: 33; Muñoz de Hoyos et al., 1982: 144; Coscarón, 1987: 33 (cat.); Conn et al., 1989: 1217; Muñoz de Hoyos, 1994 434; Muñoz de Hoyos & Miranda-Esqueivel, 1997: 302; Muñoz de Hoyos & Coscarón, 1999: 183; Coscarón & Coscarón Arias, 2007: 495; Coscarón et al., 2008: 14 (cat.); Adler & Crosskey, 2015: 31 (inv.). riveti Roubaud, 1906: 108. nitidum Malloch, 1912: 652. versicolor Lutz & Núñez Tovar, 1928: 47. avidum Hoffmann, 1930: 51. violacescens Enderlein, 1934a: 285 (Odagmia). cytoform A Conn et al., 1989: 1221. cytoform B Conn et al., 1989: 1224. cytoform C Conn et al., 1989: 1225. cytoform D Conn et al., 1988 : 1225 cytoform E Conn et al., 1989: 1226. cytoform F Conn et al., 1989: 1228. cytoform G Conn et al., 1989: 1229. cytoform I Conn et al., 1989: 1230. cytoform J Conn et al., 1989: 1231. cytoform K Conn et al., 1989: 1233. cytoform L Arteaga & Muñoz de Hoyos, 1999: 133. Subgenus Notolepria Enderlein, 1930 Notolepria Enderlein, 1930: 95 (as genus). Type species, Simulium exiguum Roubaud, 1906 (orig. des.). Refs.: Coscarón & Coscarón Arias, 2007: 224, 225-228 (redescr.; keys to spp. based on females, males, pupae and larvae). exiguum Roubaud, 1906b: 108. Type locality: Venezuela, Alto Sarate. Distr.: Colombia (Antioquia (Barbosa, El Hatillo; río Arquía, Puerto Palacio), Boyacá (Guateque; Toguí; río Ubasa); Cauca (río 20km N. of Piendamó; Santander de Quilichao; San Antonio-López de Micay, quebrada Nayita; río Micay), Cesar (Valledupar, Meollaca), Chocó (Río Arquía, Puerto Palacio; Tagachi río Tagachi; Quibdó río Bebará), Cudinamarca (Guaduas, Villeta, La Vega, San Francisco, Gachetá, Une), Guajira (Palomino, río Jeréz), Magdalena (Ariguaní), Meta (Restrepo; El Retiro), Quindío (between Montenegro and Armenia; Armenia), Norte de Santander (río Peralonso, bocas del río San Miguel), Tolima (Mariquita-Honda quebrada de Padilla), Valle del Cauca (between río Dagua and río Bitaco; Atuncela; Dagua; Saladito to Anchicayá road; Cisneros; Valle río Pance; río Pance; Cali; Pichindé, Jamundí), Argentina, Bolivia, Brazil Ecuador, Guatemala, Mexico, Panama, Perú, Venezuela. Refs.: Vulcano, 1967: 11 (cat.); Barreto, 1969: 32; Tidwell et al., 1980: 377 (as vector of Onchocerca volvulus); Tidwell et al., 1980: 119, 121, 126; Muñoz de Hoyos et al., 1982: 143; Muñoz de Hoyos, 1994: 431; Muñoz de Hoyos & Miranda-Esquivel, 1997: 301; Muñoz de Hoyos & Coscarón, 1999: 183; Coscarón & Coscarón Arias, 2007: 228; Coscarón et al., 2008: 37 (cat.); Adler & Crosskey, 2015: 68 (inv.). glaucophthalmum Knab, 1914: 123. delpontei Paterson & Shannon, 1927: 739. urubambanum Enderlein, 1934a: 286 . morphoform ‘C’ Shelley, Nunes de Mello & Rees, 1976: 328. cytoform ‘Aguarico’ Procunier, Shelley & Arzube, 1985, in: Charalambous, Shelley & Arzube, 1997: 153. cytoform ‘Bucay’ Procunier, Shelley & Arzube, 1985, in: Charalambous, Shelley & Arzube, 1997: 153. cytoform ‘Cayapa’ Procunier, Shelley & Arzube, 1985, in: Charalambous, Shelley & Arzube, 1997: 153. cytoform ‘Quevedo’ Procunier, Shelley & Arzube, 1985, in: Charalambous, Shelley & Arzube, 1997: 153. CATALOGUE OF SIMULIIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 159 cytoform ‘N’ Charalambous, Shelley & Arzube, 1997: 155 cytoform ‘Huatarac’ Charalambous, Arzube & Lowell, 1998: 251. cytoform ‘Napo’ Charalambous, Arzube & Lowell, 1998: 249. Subgenus Psaroniocompsa Enderlein Psaroniocompsa Enderlein 1934b : 192 (as genus). Type species, incrustatum, as opalinifrons (orig. des.). Refs.: Coscarón & Coscarón Arias, 2007: 295, 296-301; Coscarón et al., 2008: 38 (cat.); Adler & Crosskey, 2015: 71 (inv.). Pliodasina Enderlein, 1936 : 124 (as genus). Type species, guttatum (original designation) Cerqueirellum Py-Daniel, 1983: 165 (as subgenus of Simulium). Type species, amazonicum (orig. des.). Coscaroniellum Py-Daniel, 1983: 183 (as subgenus of Simulium). Type species, quadrifidum (orig. des.) Inaequalium Coscarón & Wygodzinsky, 1984: 78 (as subgenus of Simulium). Type species, inaequale (orig. des.) Shelleyellum Py-Daniel & Pessoa, 2005: 474 (as genus) (unav., no designated type species) Shelleyellum Py-Daniel & Pessoa, 2006: 133 (as genus). Type species, siolii (orig. des.) amazonicum species-group amazonicum Goeldi, 1905: 138. Type locality: Brazil, Amazonas, Tefé, Purus and Acre. Distr.: Colombia (Amazonas (Arara; Santa Sofía, río Amazonas 40km NW. Leticia; Puerto Arica, río Putumayo), Cesar (Valledupar), Vaupés (Santa Marta, Mitú)), Brazil, Venezuela. Refs.: Muñoz de Hoyos et al., 1982: 143; Tidwell & Tidwell, 1982: 1137; Py-Daniel, 1983: 167; Coscarón, 1987: 24 (cat.); Muñoz de Hoyos, 1994: 419; Coscarón & Coscarón Arias, 2007: 341; Coscarón et al., 2008: 16 (cat.); Adler & Crosskey, 2015: 72 (inv.). amazonense Lutz, 1917: 64, 1922: 71 morphoform ‘R. Mari’ Shelley, Hernández, Maia-Herzog, Luna Dias & Luz, 2006: 20. morphoform ‘R. Ituxi’ Shelley, Hernández, Maia-Herzog, Luna Dias & Luz, 2006: 20. tallaferroae Ramírez Pérez, 1971: 339. argentiscutum Shelley & Dias, 1980: 105. Type locality: Brazil, Rondônia, Cachoeira Teotônio, río Madeira, 8º50’S, 64º05’W, 60m. Distr.: Colombia (Amazonas (Arara; Santa Sofía, río Amazonas near Leticia)), Brazil, Perú. Refs.: Tidwell & Tidwell, 1982: 1137; Coscarón, 1987: 24 (cat.); Muñoz de Hoyos, 1994: 419; Coscarón & Coscarón Arias, 2007: 346; Coscarón et al., 2008: 17 (cat.); Adler & Crosskey, 2015: 72 (inv.). ‘sp.n’ Shelley, Luna Dias & Moraes, 1980: 785 (Simulium) ‘E’ Tidwell et al., 1981: 85 (morphoform) ‘Madeira’ Shelley, Pinter & Moraes, 1982: 27 (Simulium sp) [amazonicum: Cerqueira, 1959: 886, not Goeldi] [urubambanum: Enderlein, 1934a: 286 [1933] (part)] oyapockense Floch & Abonnenc, 1946a: 4. Type locality: Franch Guiana, Oyapock, Maripa, “près du saut Cafésoca”. Distr. Colombia (Vaupés (Santa Marta, near Mitú), Argentina, Brazil, Ecuador, French Guiana, Guyana, Paraguay, Uruguay, Venezuela. Refs.: Coscarón, 1987: 25 (cat.); Muñoz de Hoyos, 1994: 420; Coscarón & Coscarón Arias, 2007: 335; Coscarón et al., 2008: 17 (cat.); Adles & Crosskey, 2012: 70 (inv.). Refs.: Coscarón et al., 2008: 17 (cat.); Adles & Crosskey, 2015: 72 (inv.). pseudosanguineum Ramírez Pérez & Peterson, 1981: 154. pseudoamazonicum Ramírez Pérez & Peterson, 1981: 151 cuasisanguineum Ramírez Pérez et al., 1982: 36. sanchezi Ramírez Pérez et al., 1982: 71. pydanieli Pessoa et al., 2008 : 571. ‘A’ Tidwell et al., 1981: 84 (morphoform). ‘B’ Tidwell et al., 1981: 84 (morphoform). ‘C’ Tidwell et al., 1981: 85 (morphoform). 160 · Zootaxa 4122 (1) © 2016 Magnolia Press WOLFF ET AL. ‘Barbacoas’ Shelley et al., 1982: 27. ‘Bem Querer’ Shelley et al., 2006: 7. [amazonicum: authors, not Goeldi] [haematopotum: Smart, 1940, not Malloch] [minusculum: authors, not Lutz] sanguineum Knab, 1915: 279. Type locality: Colombia, Chocó, Boca de Arquia, río Atrato. Distr.: Colombia (Cauca (Santander de Quilichao), Tolima (Honda, río Magdalena), Chocó (Boca de Arquía, río Atrato; río Tagachi, near río Atrato), Tolima, Vaupés (Mitú)), Panama. Refs.: Vulcano, 1967: 20 (cat.); Barreto, 1969: 33; Muñoz de Hoyos et al., 1982: 145 (as sangineum, incorrect spelling); Muñoz de Hoyos, 1994: 420; Muñoz de Hoyos & Miranda-Esquivel, 1997: 302; Tidwell et al., 1980: 1209; Tidwell & Tidwell, 1982: 1137; Muñoz de Hoyos & Coscarón, 1999: 183; Coscarón & Coscarón Arias, 2007: 343; Coscarón et al., 2008: 18 (cat.); Adles & Crosskey, 2015: 72 (inv.). auristriatum species-group schmidtmummi Wygodzinsky, 1973a: 2. Type locality: Colombia, Boyacá, Páramo de Tota (west slope), 3400m. Distr.: Colombia (Boyacá (Aquitania, quebrada El Tejar; Páramo de Los Curies; Tota), Cundinamarca (in the Páramo area between 3050-3450m; La Calera, río Teusacá; Bogotá; Sibaté; río Chisacá; Sibaté to Aguadita road; La Siberia)). Refs.: Coscarón & Wygodzinsky, 1984: 77; Coscarón, 1987: 23 (cat.), 1991: 193; Py-Daniel, 1988: 308; Muñoz de Hoyos et al., 1982: 145 (as schmidtmmumi; incorrect spelling); Muñoz de Hoyos et al., 1994: 403; Muñoz de Hoyos & Miranda-Esquivel, 1997: 302; Coscarón & Coscarón Arias, 1997: 115; Crosskey & Howard, 1997: 57; Coscarón & Coscarón Arias, 2007: 318; Coscarón et al., 2008:40 (cat.); Adler & Crosskey, 2015: 73 (inv.). inaequale-species-group subnigrum Lutz, 1910: 239, fig. 7. Type locality: Brazil, São Paulo, São Paulo (ríos Pacaembu e Anhangabaú) e Rio de Janeiro: Mendes and Petrópolis. Distr.: Colombia (Meta (Restrepo)), Venezuela, Trinidad, Brazil, Paraguay, Argentina. Refs.: Vulcano, 1967: 21 (cat.); Barreto, 1969: 34; Muñoz de Hoyos et al., 1982: 145; Muñoz de Hoyos, 1994: 431; Coscarón & Coscarón Arias, 2007: 278; Coscarón et al., 2008: 35 (cat.); Adler & Crosskey, 2015: 73 (inv.). diversifurcatum Lutz, 1910: 258. subclavibranchium Lutz, 1910: 260. nogueirai d’Andretta & Dolores Gonzalez, 1964: 103. mbarigui Coscarón & Wygodzinsky, 1973b: 142. beaupertuyi Ramírez-Pérez, Rassi & Ramírez, 1977: 165. nahimi Py-Daniel, 1984: 125. leopoldense (Strieder & Py-Daniel), 2000: 15. incrustatum-species-group incrustatum Lutz, 1910: 243. Type locality: Brazil, Rio de Janeiro, Petrópolis and Mendes; São Paulo, Serra da Bocaina (Fazenda do Bonito) and São Paulo. LectoType locality: Brazil, Rio de Janeiro, Mendes. Distr.: Colombia (Cauca (Santander de Quilichao), Guajira (Fómeca, río Caña), Meta (Restrepo: Guacavía, Caibe, Caney, La Floresta, Los Medios, Sardinata; caño de Los Micos), Valle del Cauca (Jamundí)), Venezuela, Ecuador, Brazil, Paraguay, Argentina. Refs.: Barreto, 1969: 32; Coscarón & Wygodzinsky, 1973a: 170; Muñoz de Hoyos et al., 1982: 144; Coscarón & Wygodzinsky, 1984: 62; Coscarón, 1987: 23 (cat.), 1991: 184; Shelley, 1991: 310; Muñoz de Hoyos, 1994: 432; Coscarón, 2003: 5; Coscarón & Coscarón Arias, 2007: 311; Hernandez et al., 2007: 22; Coscarón et al., 2008: 39 (cat.); Adler & Crossky, 2015: 74 (inv.). CATALOGUE OF SIMULIIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 161 aequifurcatum Lutz, 1910: 259, 263. angrense Pinto, 1931: 683. opalinifrons (Enderlein, 1934b): 192. yarzabali Ramírez-Pérez, 1980: 66. jujuyense Paterson & Shannon, 1927: 739, 741. Type locality: Argentina, Jujuy, Zapla. Distr.: Colombia (Cundinamarca (Fusagasugá)), Argentina, Bolivia, Ecuador, Perú. Refs.: Muñoz de Hoyos et al., 1982: 144; Crosskey & Howard, 1997: 57; Coscarón & Coscarón Arias, 2007: 306 Hernandez et al., 2007: 25; Coscarón et al., 2008: 39 (cat.); Adler & Crossky, 2015: 74 (inv.). exiguum Lutz, 1910: 234 (preocc.) minimum Surcouf & Gonzalez-Rincones, 1911: 290 (subst. name, preocc., for exiguum). lutzi Knab, 1913: 155 (subst. name, for exiguum). lutzi Malloch, 1914: 14 (subst. name, preoccu., for exiguum). auripellitum Enderlein, 1934a: 286. mendozanum (Enderlein), 1936: 123 (Psaroniocompsa) guttatum (Enderlein), 1936: 124 (Pliodasina) bonaerense Coscarón & Wygodzinsky, 1984: 59. minuanum Strieder & Coscarón, 2000: 105. clavigerum Amaral-Calvao & Maia-Herzog, 2003: 266. quadrifidum-species-group quadrifidum Lutz, 1917: 66. Type locality: Brazil, Rondônia, Estrada de Ferro Madeira-Mamoré. Distr.: Colombia (Vaupés (Santa Marta, near Mitú), Venezuela, Guyana, Suriname, French Guiana, Ecuador, Brazil, Bolivia. Refs.: Muñoz de Hoyos, 1994: 421; Coscarón & Coscarón Arias, 2007: 354; Coscarón et al., 2008: 21 (cat.); Adler & Crosskey, 2015: 74 (inv.). rassii Ramírez Pérez, 1980: 60 (Simulium). torrealbai Ramírez Pérez, 1980: 64. quadrivittatum-species-group quadrivittatum Loew, 1862: 56. Type locality: “Cuba”. Distr.: Colombia (Chocó (El Tigre)), Belize, Costa Rica, Cuba, Dominican Republic, Ecuador, Guatemala, Mexico, Nicaragua, Panama, Puerto Rico, Venezuela. Refs.: Muñoz de Hoyos, 1994: 433; Coscarón & Coscarón Arias, 2007: 365; Coscarón et al., 2008: 22 (cat.); Adler & Crosskey, 2015: 75 (inv.). mallochi Enderlein, 1925: 208 (Wilhelmia; unnecessary nom. nov. for quadrivittatum of Malloch, 1914). fairchildi Vargas, 1942: 458 (nom. nov. for haematopotum Malloch of Fairchild, 1940). benjamini Dalmat, 1952: 339. [haematopotum: Fairchild, 1940: 704, not Malloch] [metallicum: Malloch, 1914: 48, nor Bellardi] Subgenus Psilopelmia Enderlein Psilopelmia Enderlein, 1934a: 283 (as genus). Type species, Psilopelmia rufidorsum Enderlein, 1934 (orig. des.) = Simulium escomeli Roubaud, 1909. Refs.: Coscarón et al., 2008: 41 (cat.); Adler & Crosskey, 2012: 73 (inv.). Ectemnaspis Enderlein, 1934a: 281 (as genus). Type species, bicoloratum Malloch, 1912, as macca (orig. des.) Lanea Vargas, Martínez Palacios & Díaz Nájera, 1946: 107, (as subgenus of Simulium). Type species, haematopotum Malloch, 1914 (orig. des.). 162 · Zootaxa 4122 (1) © 2016 Magnolia Press WOLFF ET AL. bicoloratum-species-group arcabucense Coscarón, 1990: 123. Type locality: Colombia, Boyacá, valley of río Pomeca (NW. of Arcabuco), 2400 m. Distr.: Colombia (Boyacá (Valle río Pomeca NW. Arcabuco). Refs.: Muñoz de Hoyos, 1994: 421; Crosskey & Howard, 1997: 29; Coscarón & Coscarón Arias, 2007: 390; Coscarón et al., 2008: 22 (cat.); Adler & Crosskey, 2015: 75 (inv.). bicoloratum Malloch, 1912: 649. Type locality: Perú, río Charape. Distr.: Colombia (Cauca (Popayán to Páramo de Puracé road), Cundinamarca (Sibaté to Aguadita road; Chipaque), Tolima (Ibagué to Armenia road), Valle del Cauca (Sevilla, quebrada Peñas Blancas)), Bolivia, Ecuador, Perú, Venezuela. Refs.: Vulcano, 1967: 7 (cat.); Barreto, 1969: 31; Muñoz de Hoyos et al., 1982: 143; Muñoz de Hoyos, 1994: 421; Crosskey & Howard, 1997: 29; Muñoz de Hoyos & Coscarón, 1999: 183; Coscarón & Coscarón Arias, 2007: 400; Coscarón et al., 2008: 22 (cat.); Adler & Crosskey, 2015: 75 (inv.). gaudeatum Knab, 1914: 81. macca (Enderlein), 1934a: 276 (Dasypelmoza). macca (Enderlein), 1934a: 281 (Ectemaspis; preocc. Enderlein, 1934a: 276). molli Vargas, 1943: 142 (nom. nov. for macca Enderlein, 1934a: 281). gandeatum Iriarte, 1946: 460, error. bicornutum Wygodzinsky & Coscarón, 1982: 2. Type locality: Colombia, Cundinamarca, road from Usme to Páramo de Chisacá, 3200m. Distr.: Colombia (Boyacá (Páramo de Pisba; Páramo de Los Curies, quebrada El Machito), Cundinamarca (Usme to Páramo de Chisacá road; Páramo de Chisacá, río Chisacá; Santa Helena, quebrada El Oso), Tolima (Páramo de los Valles). Refs.: Muñoz de Hoyos, 1994: 421; Muñoz de Hoyos et al., 1994: 395; Muñoz de Hoyos & Miranda-Esquivel, 1997: 302; Crosskey & Howard, 1997: 29; Muñoz de Hoyos & Coscarón, 1999: 183; Coscarón & Coscarón Arias, 2007: 397; Coscarón et al., 2008: 23 (cat.); Adler & Crosskey, 2015: 75 (inv.). callidum (Dyar & Shannon), 1927: 12, 16 (Eusimulium). Type locality: Mexico, Veracruz, Córdoba. Distr.: Colombia (Antioquia, Valle del Cauca (Sevilla, quebrada Peñas Blancas; río Pance)), Belize, Guatemala, Mexico. Refs.: Muñoz de Hoyos et al., 1982: 143; Crosskey & Howard, 1997: 58; Coscarón & Coscarón Arias, 2007: 461; Hernandez et al., 2007: 26; Coscarón et al., 2008: 41 (cat.); Adler & Crosskey, 2015: 75 (inv.). mooseri (Dampf), 1927: 127. costaricensis Smart, 1944: 132 rufidorsum (Enderlein) 1936: 119 (preocc.) cormonsi Wygodzinsky, 1971: 31. Type locality: Venezuela, Mérida. Distr.: Colombia Cundinamarca (Parque Natural Nacional Chingaza; La Cortadera stream 04°37932.3”N, 73°43940.8”W, 3040m; Monteredondo (04°37946.3”N, 73°43928.8”W, 3072m), Peñas Blancas (04°42904.2”N 73°50927.1”W, 3008m; río Chuza 04°38940.6”N, 73°44944.1”W). Refs.: Moncada et al, 2013:103; Adler & Crosskey, 2015: 75 (inv.); Díaz et al., 2015:553. furcillatum Wygodzinsky & Coscarón, 1982: 6. Type locality: Colombia, Boyacá, Páramo de Tota, 3400m. Distr.: Colombia (Boyacá (Páramo de Tota; Santuario de Fauna y Flora de Iguaque), Cundinamarca (Tunjaque, SE. La Calera; Zipaquirá to Pacho road; Bogotá, río San Francisco bajo; Usme to Páramo de Chisacá road; Usme, río Tunjuelo; Subachoque; La Calera; Bogotá to Choachí road; Chipaque; Páramo de Chisacá; Santa Helena, Quebrada El Oso; El Brasil, quebrada Hoya Honda)). Refs.: Campo-Gaona & Muñoz de Hoyos, 1990: 715; Muñoz de Hoyos, 1994: 422; Muñoz de Hoyos et al., 1994: 397; Crosskey & Howard, 1997: 29; Muñoz de Hoyos & Miranda-Esquivel, 1997: 302; Muñoz de Hoyos & Coscarón, 1999: 183; Coscarón & Coscarón Arias, 2007: 398; Coscarón et al., 2008: 23 (cat.); Adler & Crosskey, 2015: 75 (inv.); Diaz et al., 2015: 553. ignescens Roubaud, 1906: 107. Type locality: Ecuador, Napo. Distr.: Colombia (Boyacá (Arcabuco to Villa de Leyva road), Cauca (next to Puracé; río Piendamó, E. Silvia; Silvia; Paniquita, 20km N. of Popayán), CATALOGUE OF SIMULIIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 163 Cundinamarca (Albán; Albán to Sasaima road; Sibaté to Aguadita road; Aguadita, río Blanco; Aguadita to Pasca road, quebrada La Pradera; near Salto Tequendama; Guaduas; Zipaquirá to Pacho road; Villeta; La Vega; San Francisco; Subachoque, La Pradera; Chipaque to Caquezá road, Munar, río Une; Guasca; río frio W. Guasca; La Calera, Teusacá river; Santandercito Zoólogico Santacruz; Choachí; Sibaté; Parque Natural Nacional Chingaza, sector Palacio; Usme river Tunjuelo; Tunjaque SE.; Páramo de Chisacá; Pasca), Huila (San Agutín quebrada El Playón), Norte de Santander (Pamplona), Risaralda (Parque Natural Los Nevados, Ucumarí; Tolima (Ibagué to Armenia road; Anaime; Gualanday), Valle del Cauca (2km W. Felidia)), Bolivia, Ecuador, Perú, Venezuela. Refs.: Duque et al., 1988: 300 (polytene chrosomosomes); Muñoz de Hoyos et al., 1982: 143; Torres-Fernández et al., 1991: 256; Muñoz de Hoyos, 1994: 422; Muñoz de Hoyos et al., 1994: 398; Crosskey & Howard, 1997: 29; Muñoz de Hoyos & Miranda-Esquivel, 1997: 302; Muñoz de Hoyos & Coscarón, 1999: 183; Coscarón & Coscarón Arias, 2007: 387; Coscarón et al., 2008: 23 (cat.); Adler & Crosskey, 2015: 75 (inv.); Esquivel et al., 2015: 92. pautense Coscarón & Takaoka, 1989: 191. Type locality: Ecuador: Paute, Azuay, Distr.: Colombia (Cundinamarca (Chingaza National Natural Park, streams Babilonia (04°37932.3”N 73°43940.8”W, 3040m above sea level, Palacio sector La Cortadera (04°42907.7”N 73°50936.3”W, 2996m)). Ref.: Adler & Crosskey, 2015: 76 (inv.) ; Diaz et al., 2015 : 541. pifanoi Ramírez Pérez, 1971: 346. Type locality: Venezuela, Mérida, Santo Domingo. Distr.: Colombia (Cauca (near Puracé), Cundinamarca (Albán; Sibaté to Aguadita road; 1km W. Sibaté; Santandercito Zoolológico Santacruz; Aguadita; Salto del Tequendama road under)), Venezuela. Refs.: Crosskey & Howard, 1997: 29; Muñoz de Hoyos, 1994: 424; Muñoz de Hoyos & Miranda-Esquivel, 1997: 302; Muñoz de Hoyos & Coscarón, 1999: 183; Coscarón & Coscarón Arias, 2007: 388; Coscarón et al., 2008: 23 (cat.); Adler & Crosskey, 2015: 76 (inv.). tolimaense Coscarón, 1984: 301 (nom. nov. for simplex Wygodzinsky & Coscarón, 1982). Type locality: Colombia, Tolima, Ibague-Armenia highway, near La Linea, E. slope of hillside. Distr.: Colombia (Tolima (Ibagué to Armenia road). Refs.: Coscarón, 1987: 29 (cat.), 1990: 128; Muñoz de Hoyos, 1994: 424; Crosskey & Howard, 1997: 30; Coscarón & Coscarón Arias, 2007: 397; Coscarón et al., 2008: 23 (cat.); Adler & Crosskey, 2015: 76 (inv.). simplex Wygodzinsky & Coscarón 1982: 11. tunja Coscarón, 1990: 126. Type locality: Colombia, Boyacá, 2km W. of Tunja, 2900m. Distr.: Colombia (Boyacá (2km W. Tunja; near lago de Tota), Cundinamarca (Subachoque; Albán; Guasca; La Calera río Teusacá; Sibaté; Aguadita; Páramo de Chisacá; Pasca; Sibaté to Aguadita road; Usme to Páramo de Chisacá road; Subachoque, La pradera; Santa Helena, Quebrada El Oso)). Refs.: Muñoz de Hoyos, 1994: 425; Muñoz de Hoyos et al., 1994: 400; Crosskey & Howard, 1997: 30; Muñoz de Hoyos & Miranda-Esquivel, 1997: 302; Muñoz de Hoyos & Coscarón, 1999: 183, Coscarón & Coscarón Arias, 2007: 394; Coscarón et al., 2008: 23 (cat.); Adler & Crosskey, 2015: 76 (inv.). “C” Duque et al., 1988: 306 (Simulium sp.) “A” Lewis & Lee-Potter, 1964: 97 (Simulium sp.) ‘B’ Lewis & Lee-Potter, 1964: 98 (Simulium sp.) dinellii-species-group dinellii (Joan), 1912: 363 (reprint, p. 5) (Melusina). Type locality: Argentina, Tucumán, Montes de Apachirí. Distr.: Colombia (“Tierra caliente”; Risaralda (stream near the road between Otún-Quimbaya refuge 4°43’N, 75°33’W, 1885m; El Cedral aquaculture station 4°42’N 75°31’W, 2119m)), Argentina, Bolivia, Brazil, Ecuador, Perú. Refs.: Vulcano, 1967: 10 (cat.); Barreto, 1969: 31; Muñoz de Hoyos et al., 1982: 143; Muñoz de Hoyos, 1994: 427; Crosskey & Howard, 1997: 58; Coscarón & Coscarón, Arias 2007: 414; Coscarón et al., 2008: 24 (cat.); Adler & Crosskey, 2015: 76 (inv.); Mantilla et al., 2015: 217. miniatum (Enderlein, 1934a: 275 (Dasypelmoza) miniatum (Enderlein, 1934b: 194) (Trichodagmia) (2) (preocc.) 164 · Zootaxa 4122 (1) © 2016 Magnolia Press WOLFF ET AL. martinezi Vargas, 1943: 137 (nom. nov. for Trichodagmia miniata Enderlein, 1934). ochraceum Lane & Porto, 1939: 171, Lane & Vulcano, 1933: 434, not Walker. ochraceum Walker, 1861: 332. Type locality: Mexico, Chiapas, Huixtla, Morelos. Distr.: Colombia (Antioquia, Cundinamarca, Valle del Cauca (Anchicayá)), Brazil, Costa Rica, Cuba, Dominican Republic, Ecuador, Guatemala, Mexico, Panama, Perú, Puerto Rico, Trinidad, Venezuela, Virgin Islands. Refs.: Vulcano, 1967: 17 (cat.); Barreto, 1969: 34; Muñoz de Hoyos, 1994: 427; Crosskey & Howard, 1997: 58; Coscarón et al., 2008: 24 (cat.); Adler & Crosskey, 2015: 76 (inv.). bipunctatum Malloch, 1912: 650. antillarum Jennings, 1915: 200. scutellatum Lane & Porto, 1940: 192. wolcotti Fox, 1953: 138. pseudoantillarum Ramírez-Pérez & Vulcano, 1973: 379. ‘D’ Lewis, 1963: 53 (Simulium sp.) cytoform ‘A’ Hirai & Uemoto, 1983, In: Procunier, 1989: 563 cytoform ‘B’ Hirai & Uemoto, 1983, In: Procunier, 1989: 563 cytoform ‘C’ Hirai & Uemoto, 1983, In: Procunier, 1989: 563 shewellianum Coscarón, 1984: 318. Type locality: Ecuador, río Palenque, 0º 35’ S, 70º 20’ W. Distr.: Colombia (Cundinamarca (Guaduas), Valle del Cauca (10km W. Delfina, valle río Dagua; Saladito to Anchicayá road; Anchicayá 70km E. Buenaventura), Ecuador, Brazil. Refs.: Muñoz de Hoyos, 1994: 428; Muñoz de Hoyos & Miranda-Esquivel, 1997: 301; Crosskey & Howard, 1997: 58; Muñoz de Hoyos & Coscarón, 1999: 183; Coscarón & Coscarón Arias, 2007: 415; Coscarón et al., 2008: 24 (cat.); Adler & Crosskey, 2015: 76 (inv.). perflavum-species-group rorotaense Floch & Abonnenc, 1946b: 4. Type locality: French Guiana, Plateau Rorota. Distr.: Colombia (Vaupes (Santa Marta near Mitú), Venezuela, French Guiana, Brazil. Refs.: Muñoz de Hoyos, 1994: 427; Crosskey & Howard, 1997: 59; Hamada & Adler, 1998: 221, 2001: 119 (biol.); Shelley et al., 2004: 455; Coscarón & Coscarón Arias, 2007: 411, figs. 84C, 85M, 86R, 87L, 89M, 90H, K. Coscarón et al., 2008: 26 (cat.); Adler & Crosskey, 2015: 77 (inv.). fulvinotum Cerqueira & Nunes de Mello, 1967: 36 [in Cerqueira] [Brazil (Amazonas)] romanai-species-group albanense Coscarón, 1990: 135. Type locality: Colombia, Cundinamarca., 2km W. of Albán. Distr.: Boyacá (Lago de Tota; Arcabuco to Villa de Leyva road; Valle río Pomeca), Cundinamarca (Albán; 3km S. Chipaque). Refs.: Muñoz de Hoyos, 1994: 425; Crosskey & Howard, 1997: 60; Muñoz de Hoyos & Coscarón, 1999: 183; Coscarón & Coscarón Arias, 2007: 406; Coscarón et al., 2008: 64 (cat.); Adler & Crosskey, 2015: 77 (inv.). anaimense Coscarón & Muñoz de Hoyos, 1995: 587. Type locality: Colombia, Tolima, arroyo Potosí, 17km from Anaime, 2300m. Distr.: Colombia (Tolima, on the Cordillera Central, Anaime). Refs.: Crosskey & Howard, 1997: 60; Muñoz de Hoyos & Miranda-Esquivel, 1997: 302; Muñoz de Hoyos & Coscarón, 1999: 183 (key); Coscarón & Coscarón Arias, 2007: 402; Coscarón et al., 2008: 22 (cat.); Adler & Crosskey, 2015: 77 (inv.). escomeli Roubaud, 1909: 428. Type locality: Perú, Arequipa, Tingo. Distr.: Colombia, Ecuador, Chile, Perú. Refs.: Crosskey & Howard, 1997: 58; Coscarón Arias, 2003: 195, 198, 204, 209, 215; Coscarón & Coscarón Arias, 2007: 449; Coscarón et al., 2008: 42 (cat.); Adler & Crosskey, 2015: 77 (inv.). rufidorsum (Enderlein), 1934a: 283 [samboni: Dampf, 1943: 34, not Jennings] CATALOGUE OF SIMULIIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 165 gabaldoni Ramírez Pérez, 1971: 352. Type locality: Venezuela, Monagas, San Antonio. Distr.: Colombia (Cesar (Meollaca), Valle del Cauca (Valle río Pance, Hda san Pedro)), Venezuela. Refs.: Muñoz de Hoyos, 1994: 425; Crosskey & Howard, 1997: 60; Coscarón & Coscarón Arias, 2007: 405; Coscarón et al., 2008: 27 (cat.); Adler & Crosskey, 2015: 78 (inv.). ‘B’ Lewis, 1963: 53 (Simulium sp) haematopotum Malloch, 1914: 62. Type locality: Mexico, Veracruz, Santa Lucrecia & Cuba, Cayamas. Distr.: Colombia, Costa Rica, Cuba, Guatemala, Gutyana, Mexico, Panama, Puerto Rico, Venezuela. Refs.: Vulcano, 1967: 12 (cat.); Muñoz de Hoyos, 1994: 433; Crosskey & Howard, 1997: 59; Coscarón & Coscarón Arias, 2007: 448; Coscarón et al., 2008: 42 (cat.); Adler & Crosskey, 2015: 78 (inv.). pseudohaematopotum Hoffmann, 1930: 295. marathrumi Fairchild, 1940: 715. boydi De León, 1945: 76. lutzianum Pinto, 1931: 748 (rubrithorax Lutz of Lutz & Nuñez Tovar, 1928). Type locality: “Venezuela”. Distr.: Colombia (Cauca (river 20km of Piendamó; Cerro Munchique, Tambo), Cundinamarca (Guaduas; Villeta; 1km W. Sibaté; Sasaima; Subachoque; Albán; Santandercito Zoólogico Santacruz; Sibaté; Aguadita; Pasca), Norte de Santander (río Peralonso), Tolima (Anaime, Gualanday), Valle del Cauca (río Cali; Saladito to Anchicayá road; Anchicayá; Valle río Pance; río Blanco), Panama, Venezuela, Ecuador, Bolivia, Brazil, Argentina. Refs.: Barreto, 1969: 33; Muñoz de Hoyos et al., 1982: 144; Muñoz de Hoyos, 1994: 426; Crosskey & Howard, 1997: 60; Muñoz & Miranda-Esquivel, 1997: 302; Muñoz de Hoyos & Coscarón, 1999: 183; Coscarón & Coscarón Arias, 2007: 404; Coscarón et al., 2008: 263 (cat.); Adler & Crosskey, 2015: 78 (inv.). lewisi Ramírez Pérez, 1971: 349. alirioi Ramírez Pérez & Vulcano, 1973: 383. iguazuense Coscarón, 1976: 147. cytoform ‘A’ Lewis, 1963: 53 (Simulium sp.) cytoform ‘C’ Lewis, 1963: 53 (Simulium sp.) machetorum Mantilla et al., 2013: 426. Type locality: Colombia, Department of Risaralda, W side of Cordillera Central Andes, Nevado de Santa Isabel, Lake Otún (4°47'41.9”N 75o24'29.1”W), 3950 masl. Distr.: Colombia (Risaralda). Refs: Adler & Crosskey, 2015: 78 (inv.). romanai Wygodzinsky, 1951: 229 (as romañai). Type locality: Argentina, Tucumán, Cerro San Javier, Parque Aconquija. Distr.: Colombia (Cauca (20km N. Piendamó), Cundinamarca (Albán to Sasaima road; Guadua; Villeta; La Vega; San Francisco; Sasaima; Guayabal de Siquima río Siquima; Albán; Santandercito Zoológico Santacruz; Sibaté to Aguadita road; Sibaté to Fusagasuga road; Pasca), Tolima (Anaime, Gualanday), Valle del Cauca (Saladito to Anchicayá road; río Pance), Venezuela, Ecuador, Perú, Argentina. Refs.: Crosskey & Howard, 1997: 60; Muñoz de Hoyos, 1994: 426; Muñoz de Hoyos & Miranda-Esquivel, 1997: 302; Muñoz de Hoyos & Coscarón, 1999: 183; Coscarón & Coscarón Arias, 2007: 403; Coscarón et al., 2008: 26 (cat.); Adler & Crosskey, 2015: 78 (inv.). adolfolutzi Wygodzinsky, 1951: 234, figs. 1-5, 50, 52, 98-103. samboni Jennings, 1915: 199. Type locality: Panama, Canal Zone, Empire, Comacho river. Distr.: Colombia (Tolima (Mariquita-Honda quebrada de Padilla), Guajira (Riohacha, río Jeréz), Belize, Guatemala, Honduras, Mexico, Panama, Trinidad, Venezuela. Refs.: Muñoz de Hoyos et al., 1982: 145; Muñoz de Hoyos, 1994: 434; Muñoz de Hoyos & Miranda-Esquivel, 1997: 302; Muñoz de Hoyos & Coscarón, 1999: 183; Coscarón et al., 2008: 43 (cat.); Adler & Crosskey, 2015: 78 (inv.). colvini Dalmat, 1952: 344. ‘E’ Lewis, 1963: 53 (Simulium sp.) 166 · Zootaxa 4122 (1) © 2016 Magnolia Press WOLFF ET AL. Subgenus Pternaspatha Enderlein, 1930 Pternaspatha Enderlein, 1930: 84. Type species, Pternaspatha nigristrigata Enderlein, 1930 (orig. des.). Ref.: Coscarón et al., 2008: 45 (cat.); Adler & Crosskey, 2012: 77 (inv.). Acropogon Enderlein, 1934a: 276. Type species, Acropogon barbatipes Enderlein, 1934 (orig. des.). Dasypelmoza Enderlein, 1934a: 275. Type species, Simulium varipes Philippi, 1865 (orig. des.). nigristrigatum-species-group quimbayium Mantilla et al., 2013: 424. Figs. 1, 3A-C, 4A-D-5. Type locality: Colombia, Department of Risaralda, W side of Cordillera Central Andes, Nevado de Santa Isabel, Lake Otún (4°47'41.9”N 75º24'29.1”W), 3950m. Distr.: Colombia (Risaralda). Refs.: Adler & Crosskey, 2015: 80 (inv.). Subgenus Trichodagmia Enderlein Trichodagmia Enderlein, 1934a: 288 (as genus). Type species, Trichodagmia latitarsis Enderlein, 1934 (orig. des.) = Simulium townsendi Malloch, 1912. Refs.: Adler & Crosskey, 2015: 106 (inv.). Thyrsopelma Enderlein, 1934a: 284 (as genus). Type species, orbitale (Lutz, 1910) (as brasiliense), (orig. des.) Hemicnetha Enderlein, 1934b:190 (as genus). Type species, paynei (Vargas, 1942), as mexicana Enderlein (orig. des.) Dyarella Vargas et al., 1946: 105 (as subgenus of Simulium). Type species, Simulium mexicanum Bellardi, 1862 (orig. des.) Hearlea Vargas et al., 1946: 104 (as subgenus of Simulium). Type species, canadense Hearle, 1932 (orig. des.) Obuchovia Rubtsov, 1947: 90 (as subgenus of Simulium). Type species, albellum (mon.) Grenieriella Vargas & Díaz Nájera, 1951: 141 (as subgenus of Simulium). Type species, Eusimilium lahillei, 1927 (orig. des.) Hagenomyia Shewell, 1959: 83 (as subgenus of Simulium) (preocc.). Type species, pictipes Hagen, 1880 (orig. des.) Shewellomyia Peterson, 1975: 111 (new name for Hagenomyia Shewell, therefore same type species). orbitale species group muiscorum Bueno et al., 1979: 581. Type locality: Colombia, Cundinamarca (Cordillera Oriental), La Calera, río Teusacá. Distr.: Colombia Bogotá, Distrito Capital (Cundinamarca, La Calera, río Teusacá; Parque Natural Nacional Chingaza, sector Palacio 4°41′N, 73°50′W; 2,438m). Refs.: Coscarón, 1987: 37 (cat.); Muñoz de Hoyos, Martínez, Mejía & Bueno, 1994: 401; Crosskey & Howard, 1997: 79; Muñoz de Hoyos & Coscarón, 1999: 183; Miranda Esquivel & Coscarón, 2001: 429, 435; Coscarón & Coscarón Arias, 2007: 608; Adler & Crosskey, 2015: 106 (inv.); Esquivel et al., 2015: 92. nigrimanum Macquart, 1838: 88 (as nigrimana). Type locality: Brazil, São Paulo. Distr.: Colombia (Vaupés), Brazil, Paraguay, Venezuela. Refs.: Crosskey & Howard, 1997: 79; Eaton et al., 1998: 123-124; Miranda Esquivel & Coscarón, 2001: 429, 435; Coscarón & Coscarón Arias 2007: 609; Coscarón et al., 2008: 49 (cat.); Adler & Crosskey, 2015: 106 (inv.). pruinosum Lutz, 1910: 250. spadicidorsum (Enderlein), 1934b: 194. orbitale Lutz, 1910: 231, 251, figs. 35, 36, 39, 40 (nom. nov. for nigrimanum Macquart of Lutz, 1909). Type locality: Brazil, São Paulo, río Tietê, Salto de Itapura; Franca, Fazenda Norda do Mato; and Benjamin Constant, río Paraíba. Distr.: Colombia (Guajira (río Jeréz)), Brazil, Argentina, Panama. Refs.: Muñoz de Hoyos et al., 1982: 144; Adler & Crosskey, 2015: 106 (inv.). brasiliense (Enderlein), 1934a: 284 ((Thyrsopelma) CATALOGUE OF SIMULIIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 167 albopictum Lane & Porto, 1940: 193 [nigrimanum: Lutz, 1909: 137-138, 145-146, 1910: 215 / Pinto, 1931: 705-706, 726, 760, not Macquart] [paraguayense: Lane & Vulcano, 1943: 432, not Schrottky] scutistriatum Lutz, 1909: 133. Type locality: Brazil, Rio de Janeiro, Itaguaí and Duque de Caxias. Distr.: Colombia, Brazil, Venezuela. Refs.: Crosskey & Howard, 1997: 79; Coscarón & Coscarón Arias, 2007: 625; Coscarón et al., 2008: 49 (cat.); Adler & crosskey, 2015: 106 (inv.). sumapazense Coscarón & Py-Daniel, 1989: 321. Type locality: Colombia, Cundinamarca, Páramo de Sumapaz, 3700m. Distr.: Colombia (Cundimarca (Páramo de Guasca; Páramo de Sumapaz; Bogotá to Choachí road; Chisacá; río Teusacá)). Refs.: Muñoz de Hoyos, 1994: 429; Muñoz de Hoyos, 1996: 142; Crosskey & Howard, 1997: 79; Muñoz de Hoyos & Miranda-Esquivel, 1997: 302; Muñoz de Hoyos & Coscarón, 1999: 183; Coscarón & Coscarón Arias, 2007: 609.; Coscarón et al., 2008: 50 (cat.); Adler & Crosskey, 2015: 106 (inv.). townsendi Malloch, 1912: 651–652. Type locality: Perú, río Charape; 13.vii.1911. Distr.: Colombia (Boyacá, Cundinamarca; Norte de Santander), Perú, Bolivia, Ecuador. Refs.: Hernández & Shelley, 2005: 12; Coscarón & Coscarón-Arias, 2007: 601; Adler & Crosskey, 2015: 106 (inv.). chalcocoma, Knab, 1914a: 85. ecuadoriense (Enderlein), 1934b: 193. latitarse (Enderlein), 1934a: 289 (Trichodagmia). strigatum (Enderlein), 1934a: 290 (Trichodagmia). muiscorum Bueno et al., 1979: 581–591. tarsatum species-group paynei Vargas, 1942: 246 (nom. nov. for mexicanum Enderlein, 1934b). Type locality: “Mexico”. Distr.: U. S. A., México, Guatemala, Colombia (Cundinamarca, Magdalena (Minka)) Venezuela, Ecuador, Perú. Refs.: Crosskey & Howard, 1997: 38; Muñoz de Hoyos & Coscarón, 1999: 183; Coscarón & Coscarón Arias, 2007: 531, 531; Coscarón et al., 2008: 32 (cat); Adler & Crosskey, 2015: 107 (inv.). mexicanum (Enderlein, 1934b): 190 (Hemicnetha, preocc. Bellardi, 1862) mathesoni Vargas, 1943: 360. bilimekae Smart, 1944: 132. acatenangoense Dalmat, 1951: 31. conviti Ramírez-Pérez & Vulcano, 1973: 376. biuxinisa Coscarón & Ibáñez-Bernal, 1993: 64. pulverulentum Knab, 1915: 177. Type locality: Belize, Punta Gorda. Distr.: Colombia (Caldas (La Victoria), Cundinamarca (La Vega, San Francisco), Tolima (Honda río Gualí, Gualanday)), Mexico, Belize, Guatemala, Honduras, Costa Rica, El Salvador, Panama, Venezuela. Refs.: Muñoz de Hoyos, 1994: 430; Muñoz de Hoyos & Miranda-Esquivel, 1997: 302; Muñoz de Hoyos & Coscarón, 1999: 183; Coscarón et al., 2008: 30 (cat.); Adler & Crosskey, 2015: 107 (inv.). rubrithorax Lutz, 1909: 132. Type locality: Brazil, São Paulo, Serra da Bocaina (Fazenda do Bonito), 1500m, and Batatais. Type locality: Brazil, Minas Gerais, Juiz de Fora, 21o 47’S, 43o22’W. Distr.: Colombia (Meta (Restrepo: Guacavía, Caibe, Caney, La Floresta, Los Medios; El retiro)), Venezuela, Guyana, Bolívia, Brazil, Argentina. Refs.: Barreto, 1969: 33; Muñoz de Hoyos et al., 1982: 145 (as rubithorax, incorrect spelling); Coscarón, 1987: 36 (cat.); Muñoz de Hoyos, 1994: 430; Crosskey & Howard, 1997: 38; Coscarón & Coscarón Arias, 2007: 533, figs. 113A-C, 114A-B, 115A, L, O, 116A, 117A, D, H, K, N; Coscarón et al., 2008: 32 (cat.); Adler & Crosskey, 2015: 107 (inv). magnum Lane & Porto, 1940: 192. mutucuna Mello & Silva, 1974: 69. 168 · Zootaxa 4122 (1) © 2016 Magnolia Press WOLFF ET AL. tarsatum Macquart, 1846: 20. Type locality: “Colombia” [as Nouvelle Grenade]. Refs.: Vulcano, 1967: 21 (cat.); Barreto, 1969: 34; Muñoz de Hoyos et al., 1982: 145 (list.); Muñoz de Hoyos, 1994: 435; Hernández & Shelley, 2005: 6; Coscarón & Coscarón Arias, 2007: 560; Coscarón et al., 2008: 33 (cat.); Adler & Crosskey, 2015: 107 (inv.). mexicanum Bellardi, 1862: 6 [1864/5] seriatum Knab, 1914: 84. aureopunctatum Malloch, 1914: 27. placidum Knab, 1915: 281. lugubre Lutz & Nuñes Továr, 1928: 46. turgidum (Hoffmann), 1930: 293, 298-300, 303 (Eusimulium). laticalx (Enderlein), 1934a: 291 (Trichodagmia). latidigitus (Enderlein), 1936: 129 (Trichodagmia). bellardii (Py-Daniel & Moreira Sampaio), 1994: 149 (unnecessary subst. name for mexicanum Bellardi). virgatum Coquillett, 1902: 97. Type locality: U. S. A., New Mexico, Las Vegas, Hot Springs. Distr.: Colombia (Cundinamarca (Fusagasugá)), U.S.A. Mexico, El Salvador, Guatemala, Panama. Refs.: Muñoz de Hoyos et al., 1982: 145 (list); Coscarón, 1987: 36 (cat.); Muñoz de Hoyos, 1994: 431; Crosskey & Howard, 1997: 38; Coscarón & Coscarón Arias, 2007: 537, figs. 114C, 115C, 116B; Coscarón et al., 2008: 32 (cat.); Adler & Crosskey, 2015: 108 (inv.). cinereum Bellardi, 1859: 13 (preocc. Macquart, 1834). tephrodes Speiser, 1904: 148. rubicundulum Knab, 1915: 178. chiapanense Hoffmann, 1930: 295. Acknowledgments To Estrategia de Sostenibilidad 2014-2015 University of Antioquia for the grants (MW). Thanks to Biologist Carolina Henao Sepulveda for her dedication and support to this project, to people who helped in the collection of the material, specially to Jorge Luis de las Salas Ali and Arelis Coral (UNAL). TABLE 1. Simuliidae species recorded from Colombia. Genus Subgenus Group Species Author brophyi chilensis Philippi, 1865 multituberculatus Wygodzynsky & Coscarón, 1989 ortizi Wygodzynsky, 1973 osornorum Muñoz de Hoyos, Martínez, Mejia & Bueno, 1994 paranomorum Wygodzynsky & Coscarón, 1989 zumbahue Wygodzynsky & Coscarón, 1989 arrarteorum Wygodzynsky & Coscarón, 1989 basinflatus Wygodzynsky & Coscarón, 1989 destitutus Wygodzynsky & Coscarón, 1989 brevis Wygodzynsky & Coscarón, 1989 gracilis Wygodzynsky & Coscarón, 1987 leonorum Wygodzynsky & Coscarón, 1989 misitu Wygodzynsky & Coscarón, 1989 Gigantodax cilicinus cormonsi ...Continued on next page CATALOGUE OF SIMULIIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 169 TABLE 1. (Continued) Genus Subgenus Group Species Author siberianus Wygodzynsky & Coscarón, 1989 vulcanicus Wygodzynsky & Coscarón, 1987 wygodzynsky Moncada, Muñoz de Hoyos & Bueno, 1981 multifilis multifilis Wygodzynsky & Coscarón, 1989 wrighti cervicornis Wygodzynsky, 1973 cypellus Wygodzynsky & Coscarón, 1989 nasutus Wygodzynsky & Coscarón, 1989 septenarius Wygodzynsky & Coscarón, 1989 matteabranchium Anduze, 1947 metallicum Bellardi, 1859 exiguum Roubaud, 1906 amazonicum Goeldi, 1905 argentiscutum Shelley & Dias, 1980 oyapokense Floch & Abonnec, 1946 sanguineum Knab, 1915 auristriatum schmidtmummi Wygodzynsky, 1973 inaequale subnigrum Lutz, 1910 incrustatum incrustatum Lutz, 1910 jujuyense Paterson & Shannon, 1928 quadrifidum quadrifidum Lutz, 1917 quadrivittatum quadrivittatum Loew, 1862 bicoloratum arcabucense Coscarón, 1990 bicoloratum Malloch, 1912 bicornutum Wygodzynsky & Coscarón, 1982 cormonsi Wygodzynsky, 1971 callidum (Dyar & Shannon, 1927) furcillatum Wygodzynsky & Coscarón, 1982 ignescens Roubaud, 1906 pautense Coscarón & Takaoka, 1989 pifanoi Ramírez Pérez, 1971 tolimaense Coscarón, 1984 tunja Coscarón, 1990 dinellii (Joan, 1912) ochraceum Walker, 1861 shewellianum Coscarón, 1984 perflavum rorotaense Floch &Abonnenc, 1946 romanai albanense Coscarón, 1990 anaimense Coscarón & Muñoz de Hoyos, 1995 escomeli Roubaud, 1909 gabaldoni Rámirez Pérez, 1971 Simulium Aspathia Notolepria Psaroniocompsa Psilopelmia amazonicum dinellii ...Continued on next page 170 · Zootaxa 4122 (1) © 2016 Magnolia Press WOLFF ET AL. TABLE 1. (Continued) Genus Subgenus Group Species Author haemotopotum Malloch, 1914 lutzianum Pinto, 1931 machetorum Mantilla, Moncada, Matta & Adler, 2013 romanai Wygodzynsky, 1951 samboni Jennings, 1915 Pternaspatha nigristatum quimbayium Mantilla et al., 2013 Trichodagmia orbitale muiscorum Bueno, Moncada & Muñoz de Hoyos, 1979 nigrimanum Macquart, 1838 orbitale Lutz, 1910 scutistriatum Lutz, 1909 sumapazense Coscarón & Py-Daniel, 1989 townsendi Malloch, 1912 paynei Vargas, 1942 pulverulentum Knab, 1915 rubrithorax Lutz, 1909 tarsatum Macquart, 1846 virgatum Coquillett, 1902 tarsatum References Adler, P.H. & Crosskey, R.W. (2015) World black flies (Diptera: Simuliidae): A comprehensive revision of the taxonomic and geographical inventory [2015]. 123 pp. Available from: http://www.clemson.edu/cafls/biomia/pdfs/blackflyinventory.pdf (accessed 10 March 2015) d’Andretta, C. Jr. & González, D. (1964) Tres especies brasileiras de Simuliidae (Diptera, Nemtocera). 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Bulletin of the American Museum of Natural History, 189, 1–269. CATALOGUE OF SIMULIIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 177 Zootaxa 4122 (1): 178–181 http://www.mapress.com/j/zt/ Copyright © 2016 Magnolia Press Article ISSN 1175-5326 (print edition) ZOOTAXA ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.4122.1.18 http://zoobank.org/urn:lsid:zoobank.org:pub:CA3D354F-6AAD-446F-B8F5-23B266C1C59B FAMILY CORETHRELLIDAE LEONARDO H. GIL-AZEVEDO1*, CAIO C. D. CORRÊA1 & ART BORKENT2 1 Departamento de Entomologia, Museu Nacional, Universidade Federal do Rio de Janeiro, Rio de Janeiro, RJ, Brasil. E-mail: lhgazevedo@mn.ufrj.br. 2 Research Associate of the Royal British Columbia Museum, and American Museum of Natural History, 691-8th Ave. SE, Salmon Arm, British Columbia, V1E 2C2, Canada. E-mail: artborkent@telus.net. *Corresponding author: lhgazevedo@mn.ufrj.br Abstract Corethrellidae are poorly known in Colombia, with few specimens deposited in collections, and only nine named species of Corethrella Coquillett are reported for the country. An additional six undescribed species have been recently collected, indicating that the number of recorded species in Colombia is underestimated. Key words: catalogue, Colombia, distribution, frog-biting midges, Neotropical Region Introduction Corethrellidae Edwards, 1932 are found on all continents other than Antarctica, with the majority of species restricted to latitudes between 30°N and 30°S and altitudes below 1,500 meters. There are 107 extant and seven fossils species known worldwide, all in the single genus Corethrella Coquillett, 1902, with nine of the extant species present in Colombia (Borkent 2008, 2014, Borkent & Grafe 2012, Amaral & Pinho 2015, Yu et al. 2013). Adults may be recognized by their slender and very small body with a wing length of 0.6–2.5 mm. Head small, round, with large dichoptic eyes, and ocelli absent; antenna with globular pedicel, flagellum plumose in male and merely setose in female. Thorax convex, with many setae, wing with R four-branched, M two-branched, Cu twobranched, with branches of Rs and M nearly parallel to each other but with R1 closer to Sc and lying nearly midway between Sc and R2. Legs with midfemur thicker than that of the hind leg. Abdomen tubular and elongate, with setae and microtrichia (Cook 1981, Borkent 2008, 2009). The immature stages are aquatic, and inhabit tree holes, leaf axils, old bamboo stems and epiphytic plants, small pools isolated on the margins of streams, and sometimes in heavily vegetated margins of larger aquatic habitats (Borkent 2008, 2009). This catalogue is based on Borkent (2008, 2014), the departments are underlined and specific localities are in brackets (Table 1). Borkent (2008, 2014) noted the presence of further undescribed species in Colombia and a recent collection of eight specimens indicated a further six undescribed species. This indicates that it is very likely that numerous further species will be discovered in Colombia. Acronyms used for the depositories CNCI—Canadian National Collection of Insects, Ottawa, Ontario, Canada. FSP-USP—Departamento de Epidemiologia, Faculdade de Saúde Pública, Universidade de São Paulo, São Paulo, SP, Brazil. INMA—Departamento de Arthropodologia Sanitaria, Instituto Nacional de Microbiologia, "Dr. Carlos G. Malbran", Buenos Aires, Argentina. USNM—United States National Museum, Smithsonian Institution, Washington, D.C., USA. 178 Accepted by S. Nihei: 10 Apr. 2015; published: 14 Jun. 2016 List of abbreviations aut.—automatic cat.—catalogue col.—collector des—designation distr.—geographic distribution i.e.—example m.—meters mon.—monotypy nom. nov.—nomen novum (new name) orig. des.—by original designation pp.—pages preocc.—preoccupied refs.—references rev.—revision sp., spp.—species ssp.—subspecies subs.—subsequent subg.—subgenus HT—Holotype LT—Lectotype NA—Not applicable NT—Neotype ST—Syntype T—Type (unspecified) F (f#)—Female M (m#)—Male Catalogue of Corethrellidae of Colombia Family Corethrellidae Edwards, 1932 Genus Corethrella Coquillett Corethrella Coquillett, 1902: 191. Type species, Corethra brakeleyi Coquillett, 1902 (= Corethrella brakeleyi), (by orig. des.). Ramcia Annandale, 1911: 188. Type species, Ramcia inepta Annandale, 1911, (by mon.). Lutzomiops Lane, 1942: 127 (as subgenus). Type species, Corethrella nigra Lane (= C. davisi Shannon & Del Ponte, 1928 (by orig. des.). Corethrellae Shannon, 1930: 499, misspelling. amazonica Lane, 1939a: 110. Type locality: Brazil, Rondônia, Porto Velho, Rio Madeira. HT F (location unknown). Distr.: Colombia (Amazonas (Leticia), Boyacá (10km E. Puerto Boyacá, 170 m)), Mexico, Nicaragua, Costa Rica, Trinidad and Tobago, Guyana, French Guiana, and Brazil. Refs.: Borkent, 2008: 86. coutinhoi Lane, 1942: 131. appendiculata Grabham, 1906: 343. Type locality: Jamaica, Kingston. LT M (USNM) (des. by Belkin et al., 1970: 242). Distr.: Colombia (Meta (Puerto Lopez, about 50 km NE. of Remolino, about 400m from Río Meta, 160 m)), USA, Mexico, Guatemala, Nicaragua, Costa Rica, Panama, Jamaica, Dominican Republic, Dominica, St. Lucia, St. Vincent, Trinidad and Tobago, Venezuela, French Guiana, Brazil, and Argentina.). Refs.: Heinemann & Belkin, 1978: 505; Borkent, 2008: 139. arborealis Shannon & Del Ponte, 1928: 102. confusa Lane, 1939a: 109. CATALOGUE OF CORETHRELLIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 179 blanda Dyar, 1928: 79. Type locality: Panama, Las Sabanas. HT M (USNM). Distr.: Colombia (Meta (Canos Negros, 10 km E. Villavicencio, 350m; Villavicencio, Finca La Corocora, 450m; Villavicencio, Finca Porvenir, 450m), Amazonas, (Leticia)), Costa Rica, Panama, Trinidad and Tobago, French Guiana, and Brazil. Refs.: Borkent, 2008: 81. iridescens Lane, 1939b: 389 colombiana Borkent, 2008: 111. Type locality: San Isidro, Colombia. HT F (USNM). Distr.: Colombia (Cundinamarca (San Isidro, 1,465m)). Refs.: Borkent, 2008: 111. davisi Shannon & Del Ponte, 1928: 102. Type locality: Argentina, Tucuman, 5 km W. of Concepcion. LT F (des. by Casal in Belkin et al., 1968: 17 in (INMA)), (USNM). Distr.: Colombia (Meta (Villavicencio, Outskirts of Finca Porvenir, 450m; Villavicencio, Outskirts of Finca La Corocora, 450m)), Honduras, Haiti, Costa Rica, Trinidad and Tobago, Venezuela, Ecuador, French Guiana, and Brazil. Refs.: Borkent, 2008: 88. nigra Lane, 1939a: 105. nigrescens Lane, 1942: 129. edwardsi Lane 1942: 126. Type locality: Brazil, Mato Grosso, Salobra. HT F (FSP-USP). Distr.: Colombia (Cordoba (13 km NE., Monteria, 10 m)), Costa Rica, Panama, Guyana, and Brazil. Refs.: Borkent, 2008: 153. pallida Lane, 1942: 129. Type locality: Panama, Porto Bello. HT M (USNM). Distr.: Colombia (Valle del Cauca (Buenaventura, Leticia 80m)), Costa Rica, Panama, and Venezuela. Refs.: Borkent, 2008: 79. peruviana Lane, 1939a: 110. Type locality: Peru, Iquitos. HT F (FSP-USP). Distr.: Colombia (Cordoba (13 km NE., Monteria), Boyacá (10 km E. Puerto Boyacá, 170m)), Costa Rica, Panama, Trinidad and Tobago, Guyana, French Guiana, Ecuador, Peru, Brazil. Refs.: Borkent, 2008: 72. juquiana Lane, 1939a: 109 similans Lane & Aitken, 1956: 537. quadrivittata Shannon & Del Ponte, 1928: 101. Type locality: Argentina, Salta, Tres Pozos, near Embarcación. LT M (des. by Casal in Belkin et al., 1968: 16) (INMA). Distr.: Colombia (Cordoba (13 km NE. Monteria, 10m); Meta (Villavicencio, Finca La Corocora, 450m), Amazonas (PNN [Parque Nacional Natural] Amacayacu, Sector MataMata, 150 m)), Mexico, Costa Rica, Panama, Trinidad & Tobago, Venezuela, Suriname, Guyana, Peru, Brazil, Argentina. Refs.: Borkent, 2008: 155. kummi Lane, 1942: 121. dyari Lane, 1942: 125. izquierdoi Vargas, 1952: 59. TABLE 1. Corethrellidae species recorded from Colombia. Genus Species Author Corethrella amazonica Lane, 1939a appendiculata Grabham, 1906 blanda Dyar, 1928 colombiana Borkent, 2008 davisi Shannon & Del Ponte, 1928 edwardsi Lane, 1942 pallida Lane, 1942 peruviana Lane, 1939a quadrivittata Shannon & Del Ponte, 1928 180 · Zootaxa 4122 (1) © 2016 Magnolia Press GIL-AZEVEDO ET AL. References Amaral, A.P. & Pinho, L.C. (2015) New species and records of frog-biting midges from southern Brazil (Diptera: Corethrellidae). Zootaxa, 3946 (2), 274–284. http://dx.doi.org/10.11646/zootaxa.3946.2.9 Annandale, N. 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(1932) Diptera family Culicidae. Genera Insectorum, 194, 1–258. Grabham, M. (1906) A new Corethrella from Jamaica. Entomological News, 17, 343–345. Heinemann, S.J. & Belkin, J.N. (1978) Collection records of the project “Mosquitoes of Middle America” 12. Colombia (COA, COB, COL, COM). Mosquito Systematics, 10, 493–539. Lane, J. (1939a) Notes on non hematophagous Culicidae. Boletim Biologico, Clube Zoologico do Brasil, 4, 99–113. Lane, J. (1939b) Non-hematophagous Culicidae. (second paper). Boletim Biologico, Clube Zoologico do Brasil, 4, 386–393. Lane, J. (1942) Dixinae e Chaoborinae. Revisão das espécies neotropicas (Diptera, Culicidae). Revista de Entomologia, 13, 81–148, estampa 1–4. Lane, J. & Aitken, T.H.G. (1956) Chaoborinae from Trinidad, B.W.I. (Diptera, Culicidae). Annals of the Entomological Society of America, 49, 530–543. http://dx.doi.org/10.1093/aesa/49.6.530 Shannon, R.C. & Del Ponte, E. (1928) Los Culicidos en la Argentina. Revista del Instituto Bacteriologico, 5, 29–147. Shannon, R.C. (1930) List of species of Argentine Culicidae. Sexta Reunion de la Sociedad Argentina de Patologia Regional del Norte, 494–500. Vargas, L. (1952) Dos nuevas especies Mexicanas de Corethrella (Diptera: Culicidae). Revista de la Sociedad Mexicana de Historia Natural, 13, 57–62. Wood, D.M. & Borkent, A. (1986) The phylogenetic relationships among families of Nematocera. In: Abstracts of the First International Congress of Dipterology, 17–24 August 1986, Budapest, pp. 262. Wood, D.M. & Borkent, A. (1989) Phylogeny and classification of the Nematocera. Chapter 114. In: McAlpine, J.F. & Wood, D.M. (Eds.), Manual of Nearctic Diptera. Vol. 3. Research Branch Monograph 29. Agriculture Canada, Ottawa, pp. 1333–1370. Yu, Y.-X., Huang, E.-J. & Zhang, J.-Q. (2013) A new species of Corethrellidae in China – Corethrella feipengi sp. nov. (Diptera: Corethrelliidae). [in Chinese, English summary]. Acta Parasitologica et Medica Entomologica Sinica, 20, 261–264. CATALOGUE OF CORETHRELLIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 181 Zootaxa 4122 (1): 182–186 http://www.mapress.com/j/zt/ Copyright © 2016 Magnolia Press Article ISSN 1175-5326 (print edition) ZOOTAXA ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.4122.1.19 http://zoobank.org/urn:lsid:zoobank.org:pub:AAFD8661-13DB-4E21-94B0-772FE84A3B67 FAMILY BLEPHARICERIDAE LEONARDO H. GIL-AZEVEDO* & DURVAL DA S. SANTOS Departamento de Entomologia, Museu Nacional, Universidade Federal do Rio de Janeiro, Rio de Janeiro, RJ, Brasil. *Corresponding author: lhgazevedo@mn.ufrj.br Abstract Blephariceridae are poorly known in Colombia, being reasonably well sampled only in Antioquia Department. In the present catalogue, five species of Limonicola Lutz and eight species of Paltostoma Schiner are reported for the country. Key words: Blepharicerinae, catalogue, Colombia, distribution, Neotropical Region Introduction Blephariceridae Loew (1861) are found in all continents (except Antarctica), and 320 species distributed in 27 genera are reported in the world (Courtney 2009). In the Neotropical Region 76 species in five genera are known, with 13 species in two genera in Colombia. Adults may be recognized by the slender and small to medium body 3.0–13.0 mm. Head small, round with large and conspicuous eyes dichoptic or subholoptic, three ocelli, antenna with 11–13 flagellomeres. Thorax convex, wing with netlike pattern of folds, legs long and thin. Abdomen somewhat compressed, tergites and sternites simple quadrate sclerites, with sparse short setae (Hogue 1981, Courtney 2009). The immature stages are aquatic, inhabit a variety of streams with fast flowing, often torrential. Larvae are scrapers, feeding on thin films of periphyton, and attach to smooth rock substrata through the action of six ventral suctorial discs, which function as suckers (Wagner et al. 2008, Courtney 2009). The classification adopted here is based on Zwick (1977) (Table 1), specific localities are in brackets. Blephariceridae are poorly known in all South America, and in Colombia is only reasonably well sampled in Antioquia Department. Acronyms used for the depositories BMNH—Natural History Museum, London, U.K. ICN—Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Bogotá, Colombia. LACM—Natural History Museum of Los Angeles County, Los Angeles, USA. NHMW—Naturhistorisches Museum, Vienna, Autria. USNM—National History Museum of Natural History, Washington, D.C., USA. List of abbreviations des.—designation distr.—geographic distribution m.—meters mon.—monotypy orig. des.—by original designation 182 Accepted by S. Nihei: 10 Apr. 2015; published: 14 Jun. 2016 pp.—pages refs.—references rev.—revision sp., spp.—species ssp.—subspecies subs.—subsequent subg.—subgenus HT—Holotype LT—Lectotype PT—Paratype sex?—sex unknow F(f#)—Female M (m#)—Male Catalogue of Blephariceridae of Colombia Family Blephariceridae Loew, 1861 Subfamily Blepharicerinae Loew Tribe Paltostomatini Bezzi, 1913 Genus Limonicola Lutz Limonicola Lutz, 1928: 67. Type species, Limonicola plurivectis Lutz, by mon. astylis Hogue, 1989, in Hogue & Bedoya, 1989: 45. Type locality: Colombia, Antioquia, Quebrada Salado. HT M (LACM). Distr.: Colombia (Antioquia (Quebrada Salado, Quebrada Alarcona, Quebrada El Salado, 24 km E. Santuario), Valle del Cauca (Quebrada Honda, Peñas Blancas, 1.800 m)). Refs.: Hogue & Bedoya, 1989: 45. davila Hogue, 1982: 3. Type locality: Colombia, Magdalena, Sierra Nevada de Santa Marta, Río Gallina at Finca California, 1.500m. HT M (ICN), PT (ICN, USNM, BMNH, LACM). Distr.: Colombia (La Guajira (Sierra Nevada de Santa Marta, Río Ancho and Higway Nº 2), Magdalena (Sierra Nevada de Santa Marta, Río Gallina at Finca California, 1,500m, Quebrada Donama, 5 km NE. Mamatoco, Quebrada Minca, Minca, Río Sevilla, 8.7 km SW. Palmar), Antioquia (49.9 km E. Marinilla; 7.6 km W. Porce, at km 57). Refs.: Hogue, 1982: 3; Hogue & Bedoya, 1989: 47. kolomeros Hogue, 1989, in Hogue & Bedoya, 1989: 51. Type locality: Colombia, Antioquia, Quebrada Alarcona. HT M (LACM). Colombia (Antioquia (Quebrada Alarcona). Refs.: Hogue & Bedoya, 1989: 51. lichanos Hogue & Bedoya, 1989: 39. Type locality: Colombia, Antioquia, Quebrada La Hundida, 16.8 km E. Santuario. HT M (LACM). Distr.: Colombia (Antioquia (Quebrada La Hundida, 16.8 km E. Santuario, Quebrada Salado, Quebrada Santo Domingo, 7.6 km S. Porce, 7.6 km W. Porce, 7.6 km W. Porce at km 57, Río Medellín, Primavera), Caldas (5 km N. Supia), Cundinamarca (Río Caqueza, 9 km NW. Caqueza, tributary to Río Caqueza, 1 km NE. exit of Carrera 78 to Une), Quindío (Río Quindío, 10 km N. Armenia, Boquia, Quebrada Salado, 4 km E. Calarcá), Valle del Cauca (km 23 Cali-Dagua Higway)). Refs.: Hogue & Bedoya, 1989: 39. medius Hogue, 1989, in Hogue & Bedoya: 47. Type locality: Colombia, Antioquia, Quebrada Alarcona. HT M (LACM). Distr.: Colombia (Antioquia (Quebrada Alarcona, Quebrada La Agudelo, 2.5 km S. El Retiro)). Refs.: Hogue & Bedoya, 1989: 47. CATALOGUE OF BLEPHARICERIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 183 Genus Paltostoma Schiner Paltostoma Schiner, 1866: 931. Type species, Paltostoma superbiens Schiner, by orig. des. andeana Hogue & Bedoya, 1989: 35. Type locality: Colombia, Antioquia, Quebrada La Londoña, 9.3 km NW. Palmitas. HT M (LACM). Distr.: Colombia (Antioquia (Quebrada La Londoña, 9.3 km NW. Palmitas, Quebrada La Cristalina, 31 km W. Doradal, Río Araña, 20 km S. La Pintada, 49.9 km E. Marinilla, Quebrada La Habana, 72.4 km E. Marinilla, Quebrada La Habana, 72.4 km E. Marinilla, 78.1 km E. Marinilla, 13 km N. Palmitas, 7.4 km W. Porce at km 54, 5.1 km E. Río Calderas, Marinilla-Doradal, Quebrada La Hundida, 16.8 km E. Santuario; Quebrada La Cascada, 65.9 km E. Santuario), Caldas (Salto Lucio, 1 km SE. Filadelfia, 6 km N. Manizales, 5 km N. Supia), Cundinamarca (tributary to Río Sumapaz, km 86 Boquerón-Melgar Hwy), Tolima (3 km W. Boquerón)). Refs.: Hogue & Bedoya, 1989: 35. aspergonympha Hogue, 1989, in Hogue & Bedoya: 14. Type locality: Colombia, Antioquia, Quebrada El Treinta and nearby stream, 2 km N. Palmitas. HT M (LACM). Distr.: Colombia (Antioquia (Quebrada El Treinta and nearby stream, 2 km N. Palmitas, 1 km SE. Paso de Boquerón)). Refs.: Hogue & Bedoya, 1989: 14. charadrae Hogue, 1989, in Hogue & Bedoya: 18. Type locality: Colombia, Antioquia, 1km SE. Paso de Boquerón. HT M (LACM). Distr.: Colombia (Antioquia (1km SE. Paso de Boquerón, 12 km W. Piedra Blanca, Guarne, Quebrada La Cascada, 9.1 km E. Medellín, Quebrada El Treinta, 2 km N. Palmitas), Quindío (Quebrada Salado, 4 km E. Calarcá)). Refs.: Hogue & Bedoya, 1989: 18. eldorado Hogue, 1989, in Hogue & Bedoya: 24. Type locality: Colombia, Antioquia, Quebrada Piedra Blanca, 12 km W. Guarne. HT M (LACM). Distr.: Colombia (Antioquia (Quebrada Piedra Blanca, 12 km W. Guarne; Salto Tequendamita), Caldas (6 km N. Manizales)). Refs.: Hogue & Bedoya, 1989: 24. inca Hogue, 1989, in Hogue & Bedoya, 1989: 28. Type locality: Bolivia, La Paz, small tributary to south of Río Unduari, 21.2 km W. Puente Villa. HT M (LACM). Distr.: Colombia (Antioquia (Quebrada La Espadera, 7.7 km E. Medellín; 3 km N. Palmitas), Valle del Cauca (Quebrada Honda, Peñas Blancas, 1.800 m)), Bolivia, Ecuador, Peru. Refs.: Hogue & Bedoya, 1989: 28. roldani Hogue & Bedoya, 1989: 6. Type locality: Colombia, Quindío, Quebrada Salado, 4 km E. Calarcá. HT M (LACM). Distr.: Colombia (Antioquia (Quebrada Iguaná, Quebrada La Cascada, 9.1 km Medellín, Quebrada Piedra Blanca, 12 km W. Guare, Quebrada La Espadera, 7.7 km E. Medellín, Quebrada El Treinta, 2 km N. Palmitas; 1 km SE. Paso de Boquerón, Quebrada La Hundida, 16.8 km E. Santuario; Quebrada Cadavid, 17.9 km E. Santurio), Quindío (Quebrada Salado, 4 km E. Calarcá), Valle del Cauca (Quebrada Honda, Peñas Blancas, 1.800 m)). Refs.: Hogue & Bedoya, 1989: 6. saltana Edwards, 1929: 71. Type locality: Argentina, Salta, Quebrada San Lorenzo. LT F des. by Zwick, 2007: 114, (BMNH). Distr.: Colombia (Antioquia (1 km SE. Paso de Boquerón), Cundinamarca (Río Frio, road from Zipaquira to Pacho, 2,900m)), Argentina. Refs.: Hogue, 1971: 8.7; Hogue & Bedoya, 1989: 27; Zwick, 2007: 114. superbiens Schiner, 1866: 931. Type locality: Colombia. HT Sex? (NHMW). Distr.: Colombia (exact locality unknown). Refs.: Schiner, 1866: 931, Schiner, 1868: 27. Acknowledgements Weiping Xie (LACM), Gregory Courtney (Iowa State University), and Claudia Alejandra Medina Uribe (Instituto de Investigación de Recursos Biológicos Alexander von Humboldt) for information about Charles Hogue types. 184 · Zootaxa 4122 (1) © 2016 Magnolia Press GIL-AZEVEDO & SANTOS TABLE 1. Blephariceridae species recorded from Colombia. Subfamily Tribe Genus Species Author Blepharicerinae Paltostomatini Limonicola astylis Hogue, 1989 davila Hogue, 1982 kolomeros Hogue, 1989 lichanos Hogue & Bedoya, 1989 medius Hogue, 1989 andeana Hogue & Bedoya, 1989 aspergonympha Hogue, 1989 charadrae Hogue, 1989 eldorado Hogue, 1989 inca Hogue, 1989 roldani Hogue & Bedoya, 1989 saltana Edwards, 1929 superbiens Schiner, 1866 Paltostoma References Bezzi, M. (1913) Blefaroceridi italiani con descrizione di una nuova forma e di due specie esotiche. Bollettino della Società Entomologica Italiana, 44, 3–114. Brauer, F. (1869) Kurze Charakteristik der Dipteren-Larven zur Bekräftigung des neuer von Dr. Schiner entworfenen Dipteren-Systems. Abhandlungen der Zoologisch-Botanischen Gesellschaft in Wien, 19, 843–852. Courtney, G.W. (2009) Blephariceridae (Net-winged midges). In: Brown, B.V., Borkent, A., Cumming, J.M., Wood, D.M., Woodley, N.E. & Zumbado, M.A. (Eds.), Manual of Central American Diptera. Vol. 1. NRC Research Press, Ottawa, pp. 237–243. Edwards, F.W. (1929) Blepharoceridae. In: Diptera of Patagonia and South Chile. Part 2, Fascicle 2. British Museum (Natural History), London, pp. 1–75. Hogue, C.L. (1971) 8. Family Blephariceridae. In: Papavero, N. (Ed.), Catalogue of the Diptera of the Americas South of the United States. Museu de Zoologia, Universidade de São Paulo, São Paulo, pp. 12. Hogue, C.L. (1981) Blephariceridae. In: McAlpine, J.F., Peterson, B.V., Shewell, G.E., Teskey, H.J., Vockeroth, J.R. & Wood, D.M. (Eds.), Manual of Neartic Diptera. Vol. 1. Research Branch Monograph 27. Agriculture Canada, Ottawa, pp. 191–197. Hogue, C.L. (1982) Description of a new species of net-winged midge (Diptera: Blephariceridae) from the Sierra Nevada de Santa Maria, Colombia. Contributions in Science, 340, 1–10 Hogue, C.L. & Bedoya, I. (1989) The net-winged midge fauna (Diptera: Blephariceridae) of Antioquia Department, Colombia. Contributions in Science, 413, 1–57. Loew, H. (1861) Ueber die Dipterenfauna des Bernsteins. Amtlicher Bericht über die fünf und dreissigste Versammlung Deutscher Naturforscher und Ärzte in Königsberg in Preussen im September, 1860 (1861), 88–98. Loew, H. (1877) Revision der Blepharoceridae. Zeitschrift für Entomologie, Wroclaw, 6, 54–98. Rondani, C. (1856) Dipterologiae Italicae prodromus. Vol. I. Genera Italica. A. Stoschi, Parma, 226 pp. http://dx.doi.org/10.5962/bhl.title.8160 Schiner, J.R. (1866) Bericht über die von Weltumseglungsreise der k. Fregatte Novara mitgebrachten Dipteren. Verhandlungen der Kaiserlich-Königlichen Zoologisch-Botanischen Gesellschaft in Wien, 16, 927–934. Schiner, J.R. (1868) Diptera. In: Wüllerstorf-Urbair, B. von (Ed.), Reise der österreichischen Fregatte Novara um die Erde in den Jahren 1857, 1858, 1859 unter den Befehlen des Commodore B. von Wüllerstorf-Urbair. Zoologischer Theil. Vol. 2. Abt. I. B. K. Gerold’s Sohn, Wien, pp. i–vi + 1–388, pls. 1–4. Wagner, R., Barták, M., Borkent, A., Courtney, G., Goddeeris, B., Haenni, J.P., Knutson, L., Pont, A., Rotheray, G.E., Rozknosný, R., Sinclair, B., Woodley, N., Zatwarnicki, T. & Zwick, P. (2008) Global diversity of dipteran families (Insecta Diptera) in freshwater (excluding Simulidae, Culicidae, Chironomidae, Tipulidae and Tabanidae). Hydrobiologia, 595, 489–519. http://dx.doi.org/10.1007/s10750-007-9127-9 CATALOGUE OF BLEPHARICERIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 185 Zwick, P. (1977) Autralian Blephariceridae (Diptera). Australian Journal of Zoology, Supplementary Series, 46, 1–121. http://dx.doi.org/10.1071/AJZS046 Zwick, P. (2007) Some net-winged midges from Argentina (Diptera: Blephariceridae). Studies on Neotropical Fauna and Environment, 42, 109–119.} http://dx.doi.org/10.1080/01650520600908436 186 · Zootaxa 4122 (1) © 2016 Magnolia Press GIL-AZEVEDO & SANTOS Zootaxa 4122 (1): 187–238 http://www.mapress.com/j/zt/ Copyright © 2016 Magnolia Press Article ISSN 1175-5326 (print edition) ZOOTAXA ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.4122.1.20 http://zoobank.org/urn:lsid:zoobank.org:pub:EF68238F-5B32-488B-B965-1B9FE0308AB8 FAMILY PSYCHODIDAE EDUAR ELÍAS BEJARANO1,2 & LUIS GREGORIO ESTRADA1 1 Investigaciones Biomédicas, Universidad de Sucre, Cra. 14 No. 16B-32, Sincelejo, Sucre, CP 700003, Colombia. E-mail: eduarelias@yahoo.com* 2 Universidad de Cartagena, Calle 30 No. 48-152, Cartagena de Indias, Bolívar, CP 130001, Colombia *Corresponding author: eduarelias@yahoo.com. Abstract A catalogue is presented of the species of haematophagous and non-haematophagous psychodids recorded in Colombia. The list comprises 199 species distributed among five subfamilies and 16 genera, as follows: Subfamily Bruchomyiinae, genus Nemopalpus Macquart, 1838 (4 species); subfamily Phlebotominae, genera Brumptomyia França & Parrot, 1921 (8 species), Lutzomyia França, 1924 (153 species) and Warileya Hertig, 1948 (2 species); subfamily Psychodinae, genera Arisemus Satchell, 1955 (3 species), Australopericoma Vaillant, 1975 (1 species), Balbagathis Quate, 1996 (1 species), Clogmia Enderlein, 1937 (1 species), Didicrum Enderlein, 1937 (1 species), Feuerborniella Vaillant, 1971 (1 species), Lepidiella Enderlein, 1937 (1 species), Maruina Müller, 1895 (4 species), Paramormia Enderlein, 1935 (1 species), Parasetomima Duckhouse, 1968 (1 species) and Psychoda Latreille, 1796 (7 species); subfamily Sycoracinae, genus Sycorax Haliday, 1839 (5 species); and subfamily Trichomyiinae, genus Trichomyia Haliday, 1839 (5 species). Key words: catalogue, Colombia, Diptera, moth flies, Neotropical Region, sand flies Introduction This cosmopolitan family of Nematocera comprises more than 3,000 described species (Pape et al. 2011), distributed among six subfamilies, i.e., Bruchomyiinae, Horaiellinae, Phlebotominae, Psychodinae, Sycoracinae and Trichomyiinae. The adults are small, measuring 1–5 mm, the body being densely covered with hairs and scales. The wings have 9–12 longitudinal veins and lack crossveins in the distal half. The psychodids are holometabolous insects whose life cycle takes place in aquatic, semiaquatic or terrestrial ecosystems. The immatures develop in diverse microhabitats that range from still or running bodies of freshwater or wastewater to the moist soil at the bases of trees, rotten tree trunks and domestic microhabitats such as the drains of bathrooms and kitchens. The adults tend to rest in protected and relatively humid microhabitats, such as under bark and in tree holes, the undersides of leaves, crevices in rocks, burrows, stables, caves and the interior walls of buildings. Although the adults feed on plant materials such as nectar, the females of some species and, in rare cases, also certain males, feed on blood, which they obtain by biting vertebrates including man (Young & Duncan 1994). Although the psychodids per se constitute a group of taxonomically interesting insects that have sometimes been considered as potential indicators of habitat quality in tropical forests (Barrett et al. 1996, Alexander et al. 2001b), most of the studies carried out on this family revolve around the vectorial role of the Phlebotominae, which together with the Psychodinae constitute the subfamilies with the highest species richness as well as the ones of greatest interest from the medical point of view. The females of some species of sand flies can transmit protozoans, bacteria or viruses that cause diseases in man (Maroli et al. 2013), most notably leishmaniasis that causes 20,000–40,000 deaths each year (Alvar et al. 2012). Furthermore, due to their adaptation to synanthropic habitats some species of moth flies can become a sanitary nuisance, producing massive infestations in human dwellings and even hospitals (Faulde & Spiesberger 2012). Even more important, several of these species have been identified as causative agents of different types of myiasis, as potential mechanical vectors of bacterial pathogens associated with nosocomial infections or as sources of allergens in cases of asthma (Kino et al. 1987, Tu et al. 2007, Oğuz et al. 2012, Faulde & Spiesberger 2013). Accepted by S. Nihei: 25 May 2015; published: 14 Jun. 2016 187 Knowledge of the diversity of the non-haematophagous psychodids of the Neotropics remains limited, although in the last two decades there has been renewed interest in this group of insects, leading to the description of an increasing number of new taxa. Colombia is no exception to this situation and research on this family remains at an incipient level, with the exception of the phlebotomines which have been the object of extensive studies and reviewed in previous compilations (Osorno-Mesa et al. 1972b, Young 1979, Ferro & Morales 1988, Ferro & Morales 1998, Montoya-Lerma & Ferro 1999, Bejarano 2006a). Due to the renewed interest in the nonhaematophagous species as well as the public health relevance of the phlebotomines, an updated catalogue of the psychodids of Colombia is badly needed. This could be made available to taxonomists, researchers, medical entomologists and public health personnel, providing updated information on the distribution of species within Colombian territory. The present catalogue includes the haematophagous and non-haematophagous psychodids recorded in Colombia (Table 1). This list of species is based on a bibliographic review, on examination of specimens in some entomological collections and field samples made during studies undertaken in collaboration with Colombian entomologists. The taxa are organized hierarchically and in alphabetical order, under the categories of subfamily, genus, subgenus, and species or subspecies. Type species with type fixation criteria are given for genera and subgenera. Some genera are included that were recorded in the country by authors who did not name the species involved. We also include some species of sand flies that have not been formally described but which in practice are accepted as species by taxonomists. The subtribes of Psychodinae, objects of often heated debates among taxonomists, are not approached in this catalogue, nor are the categories of subgenus or species groups of phlebotomines. The valid names are highlighted in bold type, with the synonyms and the names that are not accepted in italics. Each entry lists the author’s name, year and the page number of the publication in which the name was first proposed as new. For those species that were initially described within another genus, the original generic names are given in parentheses and italics. The developmental stages and sexes currently known are then provided. The type locality is then presented, followed by the type material, which includes the type category, the sex described, and in parentheses the acronym of the museum where the type specimens are deposited. For those sand fly species described until 1994, information is provided only for the holotype and allotype; detailed information on the remaining type material can be consulted on the web page of the CIPA group (http://cipa.snv.jussieu.fr). Comments are provided for each genus on its distribution, the number of species it incorporates and citations of taxonomic revisions dealing with these taxa. In addition, the geographic distribution of each species by country is shown, and in the case of Colombia the distribution by department or district presented. Finally, bibliographic references are compiled for each species. Acronyms used for the depositories AMNH—American Museum of Natural History, New York, USA. BMNH—Natural History Museum, London, England. CEUA—Colección Entomológica de la Universidad de Antioquia, Medellín, Colombia. CIDEIM—Centro Internacional de Entrenamiento e Investigaciones Médicas, Cali, Colombia. CNRFV—Centro Nacional de Referencia de Flebótomos y otros Vectores, Universidad de Carabobo, Maracay, Venezuela. CPqRR—Centro de Pesquisas René Rachou, Belo Horizonte, Brazil. CZUS—Colección Zoológica de la Universidad de Sucre, Sincelejo, Colombia. DZUP—Coleção Entomológica Padre Jesus Santiago Moure, Universidade Federal do Paraná, Curitiba, Brazil. FMUSP—Faculdade de Medicina, Universidade de São Paulo, Brazil. FSCA—Florida State Collection of Arthropods, Gainesville, USA. FSPUSP—Faculdade de Saúde Pública da Universidade de São Paulo, São Paulo, Brazil. ICGES—Instituto Conmemorativo Gorgas de Estudios de la Salud, Panama. ICNMHN—Instituto de Ciencias Naturales, Museo de Historia Natural, Universidad Nacional de Colombia, Bogotá, Colombia. IEC—Instituto Evandro Chagas, Belém, Brazil. 188 · Zootaxa 4122 (1) © 2016 Magnolia Press BEJARANO & ESTRADA IEXA—Colección Entomológica del Instituto de Ecología, Xalapa, Mexico. IMT—Instituto de Medicina Tropical, Universidad Central de Venezuela, Caracas, Venezuela. INBIO—Instituto Nacional de Biodiversidad, Santo Domingo, Costa Rica. INH—Instituto Nacional de Higiene, Caracas, Venezuela. INHG—Instituto Nacional de Higiene, Guayaquil, Ecuador. INPA—Instituto Nacional de Pesquisas da Amazônia, Manaus, Brazil. INS—Instituto Nacional de Salud, Bogotá, Colombia. IOC—Instituto Oswaldo Cruz, Rio de Janeiro, Brazil. IP—Institut Pasteur, Paris, France. IPG—Institut Pasteur de la Guyane, Cayenne, French Guiana. IRD—Institut de Recherche pour le Développement, Montpellier, France. LACM—Natural History Museum of Los Angeles County, Los Angeles, USA. MBUCV—Museo de Biología, Universidad Central de Venezuela, Caracas, Venezuela. MHNSM—Museo de Historia Natural, Universidad Nacional Mayor de San Marcos, Lima, Peru. MNHN—Muséum National d'Histoire Naturelle, Paris, France. MNHNB—Museo Nacional de Historia Natural, La Paz, Bolivia. MZUSP—Museu de Zoologia da Universidade de São Paulo, São Paulo, Brazil. NMSA—KwaZulu-Natal Museum, Pietermaritzburg, South Africa. PMNH—Peabody Museum of Natural History, Yale University, New Haven, USA. RW—Collection of Rüdiger Wagner, Limnologische Flußstation, Schlitz, Germany. UA—University of Adelaide, Adelaide, Australia. ULA—Universidad de Los Andes, Mérida, Venezuela. ULMG—University of Leipzig, Leipzig, Germany. USNM—National Museum of Natural History, Smithsonian Institution, Washington, D.C., USA. UVS—Colección de Artrópodos de Importancia Médica, Universidad del Valle, Cali, Colombia. VHET—Colección de Vectores y Hospedadores Intermediarios de Enfermedades Tropicales, Programa de Estudio y Control de Enfermedades Tropicales, Universidad de Antioquia, Medellín, Colombia. List of abbreviations biol.—biology cat.—catalogue comm.—comments dev. biol.—developmental biology distr.—geographic distribution ecol.—ecology incl.—including leg.—legit,—collected mol. biol.—molecular biology mol. tax.—molecular taxonomy mon.—monotypy nat. inf.—natural infections nom. nov.—nomen novo, new name orig. des.—by original designation refs.—bibliographic references rev.—revision sub. des.—by subsequent designation tax.—taxonomy var.—variety, varietas. AT—Allotype HT—Holotype CATALOGUE OF PSYCHODIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 189 LT—Lectotype PLT—Paralectotype PT—Paratype E.—Egg L.—Larvae P.—Pupae F (f#)—Female M (m#)—Male Catalogue of Psychodidae of Colombia Family Psychodidae Newman, 1834 Subfamily Bruchomyiinae Alexander, 1920 Genus Nemopalpus Macquart Nemopalpus Macquart, 1838a: 219; 1838b: 223; 1839: 102. Type species, Nemopalpus flavus Macquart, 1838 (mon.). Nemapalpus Macquart, 1838a: 81; 1838b: 85. Rejected original spelling (Sabrosky et al. 1999). Nygmatodes Loew, 1845: 9. Unavailable name. Palaeosycorax Meunier, 1905: 50. Type species, Palaeosycorax tertiariae Meunier, 1905 (mon.). Comm.: This cosmopolitan genus is comprised of 42 species, 29 of which have been described from the Neotropics (Curler & Jacobson 2012, Santos et al. 2009a, 2013b), generally associated with primary forests (Wagner & Stuckenberg 2012). Quate & Alexander (2000) presented a synopsis of the New World species, including a key for the identification of males, which was followed by a revision of the genus worldwide by Williams (2003). For this reason only references published after the latter catalogue are cited below. Five species are recorded from Colombia. cancer Wagner & Stuckenberg, 2012: 365. M. Type locality: Colombia, Valle del Cauca, Peñas Blancas. Type material: HT M (NMSA). Distr.: Colombia (Valle del Cauca). mopani De León, 1950: 9. M., F. Type locality: Guatemala, Petén, Mopán and Poptún. Type material: HT M, PT F (current location unknown). Distr.: Colombia (Norte de Santander (Alexander 1987)), Belize (Ibáñez-Bernal 2001), Guatemala, Mexico. Refs.: Williams 2003: 6 (full refs. to that date, world cat.); Bejarano 2006a: 49 (Colombian Psychodidae checklist); Bravo & Barata 2012: 67 (western hemisphere checklist). patriciae Alexander, 1987: 377. M., F. Type locality: Colombia, Norte de Santander, Arboledas, Siravita. Type material: HT M, AT F (FSCA); 7 PT M (BMNH, FSCA, INS, UA, USNM). Distr.: Colombia (Norte de Santander). Refs.: Williams 2003: 7 (full refs. to that date, world cat.); Bejarano 2006a: 49 (Colombian Psychodidae checklist); Bravo & Barata 2012: 67 (western hemisphere checklist). phoenimimos Quate & Alexander, 2000: 186. M. Type locality: Colombia, Valle del Cauca, Cali, Alto Aguacatal. Type material: HT M (USNM); 4 PT M (BMNH, MZUSP); 1 PT M (LACM). Distr.: Colombia (Valle del Cauca), Venezuela (Cazorla et al., 2014). Refs.: Williams 2003: 7 (full refs. to that date, world cat.); Bejarano 2006a: 49 (Colombian Psychodidae checklist); Bravo & Barata 2012: 67 (western hemisphere checklist). Subfamily Phlebotominae Rondani, 1840 Genus Brumptomyia França & Parrot Phlebotomus, subg. Brumptomyia França & Parrot, 1921: 281. Type species, Brumptomyia brumpti Larrousse, 1920 (sub. des., Dyar 1929: 112). Comm.: This constitutes a taxonomically stable genus which comprises 26 species distributed from Mexico to 190 · Zootaxa 4122 (1) © 2016 Magnolia Press BEJARANO & ESTRADA Argentina (Galati et al. 2007), eight of which have been recorded in Colombia. As in other phlebotomines, the females of Brumptomyia exhibit haematophagous habits but they are clearly zoophilic, without any role having been attributed to these species in the transmission of pathogens. avellari (Costa Lima), 1932: 48 (Phlebotomus). E., L., P., M., F. Type locality: Brazil, Minas Gerais, Lassance. Type material: ST M (IOC). Distr.: Colombia (Boyacá, Córdoba), Argentina (Spinelli et al. 1999), Brazil, Bolivia, Panama, Paraguay (Salomón et al. 2003), Peru, Venezuela. Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 49 (Colombian Psychodidae checklist). beaupertuyi (Ortíz), 1954: 235 (Phlebotomus). M., F. Type locality: Venezuela, Lara, Duaca. Type material: HT M (MBUCV). Distr.: Colombia (Antioquia (Contreras-Gutiérrez et al. 2014), Caldas (Contreras-Gutiérrez et al. 2014), Norte de Santander, Santander, Valle del Cauca (Barreto et al. 2007)), Venezuela. Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 49 (Colombian Psychodidae checklist). galindoi (Fairchild & Hertig), 1947a: 615 (Phlebotomus). L., P., M., F. Type locality: Panama, Chiriquí, Boquete. Type material: HT M (FSCA). Distr.: Colombia (?Antioquia, ?Bolívar, ?Boyacá, ?Caquetá), Bolivia, Brazil, ?Costa Rica, Ecuador, ?Nicaragua (Valle & Rivera 1995), Panama, Paraguay (Young 1979), Peru. Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 49 (Colombian Psychodidae checklist). hamata (Fairchild & Hertig), 1947a: 614 (Phlebotomus). M., F. Type locality: Panama, Panamá, Chilibre, Chilibrillo. Type material: HT M (FSCA). Distr.: Colombia (Antioquia (Vivero et al. 2010b), Chocó, Santander, Sucre (Pérez-Doria et al. 2009)), Belize, Costa Rica, Ecuador (Martins et al. 1978), Mexico, Panama, Peru (Cáceres & Galati 2001). Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 49 (Colombian Psychodidae checklist). leopoldoi (Rodríguez), 1953: 52 (Phlebotomus). M., F. Type locality: Ecuador, Los Ríos, Quevedo. Type material: HT M, AT F (?INHG). Distr.: Colombia (Antioquia, Chocó, Nariño (Alexander et al. 1995), Valle del Cauca), Belize, Ecuador, Panama. Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 49 (Colombian Psychodidae checklist). mesai Sherlock, 1962: 332. M., F. Type locality: Colombia, Santander, San Vicente de Chucurí. Type material: HT M (current location unknown). Distr.: Colombia (Antioquia (Vivero et al. 2010b), Chocó (Vivero et al. 2011b), Santander, Sucre (Bejarano et al. 2007a)), Belize, Honduras, Mexico. Refs.: Ibáñez-Bernal 1999: 70 (revalidated from the synonymy of Brumptomyia galindoi, distr.); Bejarano 2006a: 49 (Colombian Psychodidae checklist). pentacantha (Barretto), 1947: 239 (Flebotomus). M, ?F. Type locality: Brazil, Pará, Belém, Aurá. Type material: HT M (?FMUSP). Distr.: Colombia (Amazonas), Bolivia, Brazil, Ecuador, Peru. Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 49 (Colombian Psychodidae checklist). pintoi (Costa Lima), 1932: 48 (Phlebotomus). M., F. Type locality: Brazil, Minas Gerais, Lassance. Type material: ST M (?IOC). Distr.: Colombia (Meta), Argentina, Bolivia, Brazil, French Guiana, Suriname, Venezuela. Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 49 (Colombian Psychodidae checklist). Genus Lutzomyia França Phlebotomus, subg. Lutzomyia França, 1924: 10 (nom. nov. for Lutzia França, 1920). Type species, Phlebotomus longipalpis Lutz & Neiva, 1912 (orig. des.). Lutzia França, 1920: 234 (not Theobald, 1903). Preoccupied name. Fransaia Dyar & Núñez-Tovar, 1927: 155 (nom. nov. for Lutzia França, 1920, not Theobald, 1903). Lutziomyia Cordero, Vogelsang & Cossio, 1928: 649 (nom. nov. for Lutzia França, 1920, not Theobald, 1903). Lutziola Strand, 1932: 195 (nom. nov. for Lutzia França, 1920; not Theobald, 1903). Comm.: Due to its recognized medical importance, this is the New World psychodid genus that has been most studied. Its distribution extends from Canada to Argentina (Young & Duncan 1994), although the greatest diversity of species is CATALOGUE OF PSYCHODIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 191 concentrated in the Neotropics, particularly in Brazil and Colombia. Although these species have been the object of numerous revisions (Barretto 1955b, 1962, Theodor 1965, Forattini 1973, Abonnenc & Leger 1976, Lewis et al. 1977, Martins et al. 1978, Young 1979, Artemiev 1991, Young & Duncan 1994, Galati 1995, CIPA group 1999), its taxonomy remains as controversial as ever. Two taxonomic proposals are currently extant, one of which suggests that Lutzomyia is a single genus comprised of 26 subgenera and species groups (Young & Duncan 1994) and the other dividing the taxon into 19 genera (Galati 1995, 2003a). The present catalogue adopts the former approach, without giving any consideration to the putative subgenera of Lutzomyia. However, there is clearly a need for detailed molecular studies that involve representatives of all the subgenera or genera described to allow its evolutionary history to be unravelled (Bejarano 2002). Young & Duncan (1994), CIPA group (1999) and Galati (2003a,b) all carried out detailed revisions of the taxonomy and geographic distribution of this group of insects. Of the almost 500 species described from the Neotropics, 153 have been recorded from Colombia, besides six that have not been formally described, i.e., Lu. sp. no. 1 from Araracuara Morales & Minter, 1981, Lu. sp. no. 2 from Araracuara Morales & Minter, 1981, Lu. sp. from Pichindé Young, 1979, Lu. sp. from Tres Esquinas Young, 1979, Lu. sp. from Anchicayá Young, 1979 and Lu. sp. from Baduel (Floch & Abonnenc, 1945a). The two undescribed species from Araracuara, known from both males and females, have only been collected from the Department of Caquetá, whereas Lu. sp. of Pichindé, for which both sexes are also known, has been recorded from Norte de Santander (Gálvis-Ovallos et al. 2013), Risaralda, Santander (Sandoval et al. 2006) and Valle del Cauca. The undescribed species Lu. sp. of Tres Esquinas, known only from female specimens, has been reported from Caquetá, Guaviare (Cabrera et al. 2009), Meta (Cabrera et al. 2009) and Putumayo. Two of the six undescribed species have also been collected outside Colombia: Lu. sp. of Anchicayá, known only from the male, which has been recorded from Ecuador (Young & Duncan 1994) as well as Valle del Cauca, Colombia; and Lu. sp. from Baduel, considered by some authors to be conspecific to Lu. sericea (Floch & Abonnenc, 1944a) (Leon et al. 2009), reported from Brazil, French Guiana and Suriname as well as the Colombian departments of Guaviare (Cabrera et al. 2009) and Vichada. Note: Bibliographic citations are only given for those geographic records not included by the CIPA group (1999) or Bejarano (2006a), as well as for unavailable names. Similarly, only literature published since the latter article is cited, dealing with changes in the taxonomic status of the species. abonnenci (Floch & Chassignet), 1947b: 1 (Phlebotomus). M., F. Type locality: French Guiana, Cayenne, Baduel. Type material: HT M (IP). Distr.: Colombia (Antioquia, Bolívar, Boyacá, Caldas, Caquetá, Chocó, Meta (Molina et al. 1997), Nariño, Norte de Santander, Santander, Sucre (Romero-Ricardo et al. 2013)), Bolivia, Brazil, Ecuador, French Guiana, Panama, Peru, Suriname, Venezuela. Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 52 (Colombian Psychodidae checklist). abunaensis Martins, Falcão & Silva, 1965: 17. M., F. Type locality: Brazil, Rondônia, Guajará-Mirim, Abunã. Type material: HT M (?CPqRR). Distr.: Colombia (Amazonas), Bolivia (Galati 2003a), Brazil, Ecuador, Peru. Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 50 (Colombian Psychodidae checklist). aclydifera (Fairchild & Hertig), 1952: 511 (Phlebotomus). L., P., M., F. Type locality: Panama, Canal Zone, Mojinga swamp, near Gatún. Type material: HT M, AT F (FSCA). Distr.: Colombia (Antioquia, Chocó, Valle del Cauca), Belize (Ibáñez-Bernal 2001), Bolivia (Williams 1999), Costa Rica, Ecuador, Guatemala (Rowton et al. 1991), Honduras, Mexico, Nicaragua, Panama. Refs.: Rosabal 1954: 11 (Phlebotomus sp. no. 1); CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 50 (Colombian Psychodidae checklist). amazonensis (Root), 1934: 244 (Phlebotomus). M., F. Type locality: Peru, Loreto, Iquitos. Type material: LT F (USNM). Distr.: Colombia (Amazonas, Caquetá, Meta, Putumayo), Bolivia, Brazil, Ecuador, French Guiana, Peru, Suriname, Trinidad and Tobago, Venezuela. Refs.: CIPA group 1999 (tax., distr., nat. inf., ecol., refs.); Bejarano 2006a: 52 (Colombian Psychodidae checklist). robini Abonnenc, Arias, Léger & Young, 1980: 711. andina Osorno, Osorno-Mesa & Morales, 1972c: 2. M., F. Type locality: Colombia, Cundinamarca, Soacha, El Charquito. Type material: HT M, AT F (INS). Distr.: Colombia (Cundinamarca). Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 54 (Colombian Psychodidae checklist). antioquiensis (Wolff & Galati), 2002: 318 (Pintomyia). M. Type locality: Colombia, Antioquia, Montebello, Savanitas. Type material: HT M (CEUA). Distr.: Colombia (Antioquia). Refs.: Bejarano 2006a: 54 (Colombian Psychodidae checklist). 192 · Zootaxa 4122 (1) © 2016 Magnolia Press BEJARANO & ESTRADA antunesi (Coutinho), 1939: 181 (Flebotomus). E., L., M., F. Type locality: Brazil, Amazonas, Manacapurú, Rio Amazonas. Type material: HT M (FSPUSP). Distr.: Colombia (Amazonas, Arauca (Turriago et al. 2007), Caquetá, Casanare, Guainía (Bejarano et al. 2007a), Guaviare (Cabrera et al. 2009), ?La Guajira (Osorno-Mesa et al. 1972b), Meta, Putumayo, Santander (Sandoval et al. 2006), Vaupés, Vichada), Bolivia, Brazil, French Guiana, Peru, Suriname, Trinidad and Tobago, Venezuela. Refs.: CIPA group 1999 (tax., distr., nat. inf., ecol., refs.); Bejarano 2006a: 51 (Colombian Psychodidae checklist). balourouensis (Floch & Abonnenc), 1944b: 1 (Phlebotomus). machicouensis (Floch & Abonnenc), 1944d: 8 (Phlebotomus). aragaoi (Costa Lima), 1932: 48 (Phlebotomus). M., F. Type locality: Brazil, Minas Gerais, Lassance. Type material: ST M (IOC). Distr.: Colombia (Amazonas, Antioquia, Caquetá, Casanare, Chocó, Guaviare (Cabrera et al. 2009), Guainía (Flórez & Ferro 2007), Huila (Barreto et al. 2006), Meta, Nariño, Putumayo, Valle del Cauca, Vichada), Bolivia, Brazil, Costa Rica, Ecuador, French Guiana, Panama, Paraguay, Peru, Suriname (Kent et al. 2013), Trinidad and Tobago, Venezuela. Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 50 (Colombian Psychodidae checklist). heckenrothi (Floch & Abonnenc), 1942: 8 (Phlebotomus). araracuarensis Morales & Minter, 1981: 98. M. Type locality: Colombia, Caquetá, Araracuara, left margin of Río Caquetá. Type material: HT M (INS). Distr.: Colombia (Amazonas, Caquetá), Brazil. Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 50 (Colombian Psychodidae checklist). atroclavata (Knab), 1913: 135 (Phlebotomus). L., M., F. Type locality: Trinidad and Tobago, Gasparee Island. Type material: LT M, PLT F (USNM). Dist.: Colombia (Bolívar (incl. Isla Fuerte (Montoya-Lerma 1996)), Boyacá, Caldas, Chocó (Vivero et al. 2011b), Cundinamarca, Huila, La Guajira, Magdalena, Norte de Santander, Risaralda (Bejarano et al. 2007c), Santander, Sucre, Tolima), Costa Rica, Guadeloupe, Martinique, Panama, Trinidad and Tobago, Venezuela, Virgin Islands. Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 51 (Colombian Psychodidae checklist). tejerae (Larrousse), 1922: 41 (Phlebotomus). guadeloupensis (Floch & Abonnenc), 1945a: 1 (Phlebotomus). auraensis (Mangabeira), 1942a: 161 (Flebotomus). M., F. Type locality: Brazil, Pará, Belém, Aurá. Type material: HT M (IOC). Distr.: Colombia (Caquetá, Putumayo), Argentina (Spinelli et al. 1999), Bolivia, Brazil, Peru, Suriname, Venezuela. Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 53 (Colombian Psychodidae checklist). ayrozai (Barretto & Coutinho), 1940: 131 (Phlebotomus). M., F. Type locality: Brazil, São Paulo, Horto Florestal da Cantareira. Type material: ST M (FSPUSP). Distr.: Colombia (Amazonas, Antioquia, Boyacá, Caquetá, Casanare (Molina et al. 2008), Cauca, Caldas (Barreto et al. 2006), Chocó, Guainía (Flórez & Ferro 2007), Meta, Nariño, Putumayo, Risaralda, Santander, Valle del Cauca, Vichada (Vivero et al. 2010a)), Bolivia, Brazil, Ecuador, French Guiana, Panama, Peru, Suriname (Kent et al. 2013) Trinidad and Tobago, Venezuela. Refs.: Lewis 1975: 502 (Lutzomyia Turure sp.); CIPA group 1999 (tax., distr., nat. inf., ecol., refs.); Bejarano 2006a: 52 (Colombian Psychodidae checklist). tintinnabula Christensen & Fairchild, 1971: 301. barrettoi barrettoi (Mangabeira), 1942a: 148 (Flebotomus). M., F. Type locality: Brazil, Pará, Belém, Aurá. Type material: HT M (IOC). Distr.: Colombia (Caquetá, Guaviare (Cabrera et al. 2009), Meta (Vásquez-Trujillo et al. 2013)), Bolivia (Le Pont et al. 1992), Brazil, Ecuador, French Guiana, Peru, Suriname, Trinidad and Tobago. Refs.: Floch & Abonnenc 1946: 3 (Phlebotomus sp. from Maripa); CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 50 (Colombian Psychodidae checklist). barrettoi majuscula Young, 1979: 129. L., M., F. Type locality: Panama, Canal Zone, Madden Forest Reserve, Cruces Trail. Type material: HT M, AT F (FSCA). Distr.: Colombia (Antioquia, Chocó, Cauca (Barreto et al. 2006), Guaviare (Cabrera et al. 2009), Meta (Vásquez-Trujillo et al. 2013), Nariño, Santander (Sandoval et al. CATALOGUE OF PSYCHODIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 193 2006), Tolima, Valle del Cauca), Costa Rica, Ecuador, El Salvador, Honduras, Nicaragua, Panama. Refs.: Rosabal 1954: 10 (Phlebotomus sp. no. 6); CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 50 (Colombian Psychodidae checklist). begonae (Ortíz & Torres), 1975: 101 (Phlebotomus). M., F. Type locality: Venezuela, Amazonas, El Gavilán. Type material: HT M (?IMT). Distr.: Colombia (Guaviare (Cabrera et al. 2009), Meta (Vásquez-Trujillo et al. 2013)), Brazil, Venezuela. Refs.: CIPA group 1999 (tax., distr., ecol., refs.). bernalei (Osorno-Mesa, Morales & Osorno), 1967: 30 (Phlebotomus). M., F. Type locality: Colombia, Caquetá, Araracuara, left margin of Río Caquetá. Type material: HT M (INS). Distr.: Colombia (Caquetá, Guainía (Flórez & Ferro 2007)), Bolivia (Galati 2013), Brazil, Venezuela. Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 52 (Colombian Psychodidae checklist). bettinii Feliciangeli, Ramírez-Pérez & Ramírez, 1988: 52. M., F. Type locality: Venezuela, Amazonas, Cerro La Neblina. Type material: HT M, AT F (CNRFV). Distr.: Colombia (Guainía (Flórez & Ferro 2007), Vichada (Vivero et al. 2010a)), Brazil (Fé et al. 1998), Venezuela. Refs.: CIPA group 1999 (tax., distr., ecol., refs.). bicolor Fairchild & Theodor, 1971: 157. E., L., P., M., F. Type locality: Panama, Canal Zone, Ft. Sherman Reservation, Mojinga Swamp. Type material: HT M, AT F (FSCA). Distr.: Colombia (Amazonas, Antioquia, Boyacá, Caquetá, Casanare, Cauca (incl. Isla Gorgona (Barreto et al. 1997)), Chocó, Cundinamarca, Guainía (Bejarano et al. 2007a), Guaviare (Cabrera et al. 2009), La Guajira, Magdalena, Meta, Nariño, Norte de Santander, Putumayo, Santander, Sucre (Romero-Ricardo et al. 2013), Tolima, Valle del Cauca), Brazil, Costa Rica, Ecuador, Panama, Peru, Venezuela. Refs.: CIPA group 1999 (tax., distr., nat. inf., ecol., refs.); Williams 1999: 473 (raised to the category of species); Bejarano 2006a: 51 (Colombian Psychodidae checklist). bifoliata Osomo-Mesa, Morales, Osorno & Muñoz, 1970: 8. M., F. Type locality: Colombia, Boyacá, Puerto Boyacá, El Terminal. Type material: HT M (INS). Distr.: Colombia (Antioquia, Boyacá, Caldas, Chocó, Risaralda, Santander, Tolima, Valle del Cauca (Jaramillo & Montoya-Lerma 1991)), Ecuador (Hashiguchi et al. 2014). Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 50 (Colombian Psychodidae checklist). bispinosa (Fairchild & Hertig), 1951a: 410 (Phlebotomus). M., F. Type locality: Panama, Cerro Jefe, La Victoria. Type material: HT M, AT F (FSCA). Distr.: Colombia (Amazonas, Caquetá, Chocó, Putumayo, Vaupés), Belize, Brazil, Costa Rica, Ecuador, French Guiana, Guatemala, Honduras, Nicaragua, Panamá, Suriname. Refs.: CIPA group 1999 (tax., distr., nat. inf., ecol., refs.); Bejarano 2006a: 53 (Colombian Psychodidae checklist). bursiformis (Floch & Abonnenc), 1944c: 5 (Phlebotomus). M., F. Type locality: French Guiana, Cayenne, Baduel. Type material: HT F (IP). Distr.: Colombia (Boyacá), Brazil, Ecuador, French Guiana, Venezuela. Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Galati 2003a: 37 (tax., distr.). baityi (Damasceno, Causey & Arouck), 1945: 22 (Flebotomus). campbelli (Damasceno, Causey & Arouck), 1945: 26 (Flebotomus). M., F. Type locality: Brazil, Pará, Monte Alegre, Machirá. Type material: HT M (USNM). Distr.: Colombia (Guaviare (Cabrera et al. 2009), Bolivia, Brazil, French Guiana, Peru, Venezuela. Refs.: CIPA group 1999 (tax., distr., ecol., refs.). camposi (Rodríguez), 1950: 26 (Phlebotomus). L., M., F. Type locality: Ecuador, Los Ríos, Quevedo. Type material: HT M (?INHG). Distr.: Colombia (Antioquia, Atlántico (Maestre-Serrano et al. 2010), Boyacá, Caldas, Chocó, Huila (Carvajal et al. 2003), Nariño, Santander, Sucre (Pérez-Doria et al. 2008), Tolima, Valle del Cauca), Costa Rica, Ecuador, Nicaragua (Collantes & Martínez-Ortega 1998), Panama. Refs.: Floch 1953: 58 (Phlebotomus sp. o); Rosabal 1954: 10 (Phlebotomus sp. no. 4); CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 52 (Colombian Psychodidae checklist). acanthobasis (Fairchild & Hertig), 1952: 508 (Phlebotomus). 194 · Zootaxa 4122 (1) © 2016 Magnolia Press BEJARANO & ESTRADA caprina Osorno-Mesa, Morales & Osorno, 1972a: 437. M., F. Type locality: Colombia, Caldas, La Victoria, El Llano. Type material: HT M, AT F (INS). Distr.: Colombia (Antioquia, Boyacá, Caldas, Chocó, Córdoba, Santander, Valle del Cauca), Brazil (Oliveira et al. 2013), Costa Rica, Honduras, Nicaragua (Collantes & MartínezOrtega 1998), Panama, Peru. Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 54 (Colombian Psychodidae checklist). carpenteri (Fairchild & Hertig), 1953: 28 (Phlebotomus). L., M., F. Type locality: Panama, Canal Zone, Chiva Chiva. Type material: HT M, AT F (FSCA). Distr.: Colombia (Antioquia, Caldas (Contreras-Gutiérrez et al. 2014), Chocó, Cundinamarca, Guainía (Flórez & Ferro 2007), Guaviare (Cabrera et al. 2009), Huila (Carvajal et al. 2003), Magdalena (Bejarano et al. 2015), Santander (Arismendi-Solano 2005), Sucre (Pérez-Doria et al. 2008), Tolima (Marín-Casas et al. 2008)), Belize (Williams 1999), Costa Rica, Mexico, Nicaragua (Valle & Rivera 1995), Panama. Refs.: Rosabal 1954: 11 (Phlebotomus sp. no. 2); CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 50 (Colombian Psychodidae checklist). carrerai (Barretto), 1946: 286 (Flebotomus). E., L., M., F. Type locality: Colombia, Meta, Restrepo, Vega Grande. Type material: HT M, AT F (FSPUSP). Distr.: Colombia (Amazonas, Caquetá, Guaviare (Cabrera et al. 2009), Meta, Nariño (Barreto et al. 1989), Putumayo, Risaralda (Delgado et al. 2011), Valle del Cauca (Jaramillo & Montoya-Lerma 1991)), Bolivia, Brazil, Ecuador, Peru, Venezuela. Refs.: CIPA group 1999 (tax., distr., nat. inf., ecol., refs.); Bejarano 2006a: 53 (Colombian Psychodidae checklist); Carvalho et al. 2006: 131 (raised to the category of species). pessoana (Barretto), 1955a: 247 (Sergentomyia). cayennensis cayennensis (Floch & Abonnenc), 1941a: 13 (Phlebotomus). L., P., M., F. Type locality: French Guiana, Cayenne, Montjoly. Type material: HT M (IP). Distr.: Colombia (Antioquia, Atlántico (Maestre-Serrano et al. 2010), Bolívar, Boyacá (Osorno-Mesa et al. 1970), Caldas, Cesar, Chocó (Carrillo et al. 2011), Córdoba, Cundinamarca, Huila, La Guajira, Magdalena, Meta, Nariño, Norte de Santander, Risaralda (Barreto et al. 1997), Santander, Sucre, Tolima), Belize, Brazil, Costa Rica, Ecuador, El Salvador, French Guiana, Honduras, Mexico, Nicaragua (Valle & Rivera 1995), Panama, Peru, Trinidad and Tobago, Venezuela. Refs.: CIPA group 1999 (tax., distr., nat. inf., ecol., refs.); Bejarano 2006a: 51 (Colombian Psychodidae checklist). ceferinoi (Ortíz & Álvarez), 1963: 285 (Phlebotomus). M, ?F. Type locality: Venezuela, Portuguesa, road to Biscucuy. Type material: HT M (current location unknown). Distr.: Colombia (Norte de Santander (Galati & Cáceres 1994), Tolima (Ferro et al. 2009)), Venezuela. Refs.: CIPA group 1999 (tax., distr., ecol., refs.). cellulana Young, 1979: 142. M, ?F. Type locality: Colombia, Caquetá, Tres Esquinas. Type material: HT M, AT F (FSCA). Distr.: Colombia (Antioquia (Zuluaga et al. 2012), Caquetá, Guaviare (Cabrera et al. 2009), Meta, Putumayo), Ecuador. Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 53 (Colombian Psychodidae checklist). cerqueirai (Causey & Damasceno), 1945b: 645 (Flebotomus). M., F. Type locality: Brazil, Pará, Belém, Utinga. Type material: HT M (USNM). Distr.: Colombia (Santander (Sandoval et al. 2006), Tolima), Brazil, Peru (Fernández et al. 1994). Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 50 (Colombian Psychodidae checklist). chagasi (Costa Lima), 1941: 12 (Flebotomus). M., F. Type locality: Brazil, Amazonas, Rio Negro. Type material: HT F (IOC). Distr.: Colombia (Amazonas, Caquetá, Meta (Vásquez-Trujillo et al. 2013), Putumayo, Santander (Sandoval et al. 2006), Vaupés), Bolivia, Brazil, Peru, Venezuela. Refs.: CIPA group 1999 (tax., distr., nat. inf., ecol., refs.); Bejarano 2006a: 53 (Colombian Psychodidae checklist). unisetosus (Mangabeira), 1941b: 238 (Flebotomus). chassigneti (Floch & Abonnenc), 1944a: 3 (Phlebotomus). M., F. Type locality: Frech Guiana, Cayenne, Baduel. Type material: HT M, AT F (IP). Distr.: Colombia (Amazonas, Antioquia, Chocó), Brazil, French Guiana, CATALOGUE OF PSYCHODIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 195 Suriname. Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 52 (Colombian Psychodidae checklist). choti (Floch & Abonnenc), 1941a: 7 (Phlebotomus). P., M., F. Type locality: French Guiana, Cayenne, Montjoly. Type material: HT M, AT F (IP). Distr.: Colombia (Guaviare, Meta), Bolivia, Brazil, Ecuador, French Guiana, Peru, Suriname. Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 52 (Colombian Psychodidae checklist). christenseni Young & Duncan, 1994: 272. L., P., M., F. Type locality: Panama, Canal Zone, Las Cruces Trail. Type material: HT M, AT F (FSCA). Distr.: Colombia (Amazonas, Antioquia, Boyacá, Caquetá, Chocó, Meta, Santander (Sandoval et al. 2006), Tolima (Morales et al. 1981)), Brazil, Panama, Trinidad and Tobago, Venezuela. Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 52 (Colombian Psychodidae checklist). cirrita Young & Porter, 1974: 321. M., F. Type locality: Colombia, Antioquia, 24 km SW of Zaragoza, Río Anorí. Type material: HT M, AT F (FSCA). Distr.: Colombia (Antioquia, Caldas, Risaralda, Valle del Cauca). Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 50 (Colombian Psychodidae checklist). claustrei Abonnenc, Léger & Fauran, 1979: 80. M., F. Type locality: French Guiana, Maripasoula. Type material: HT M (MNHN). Distr.: Colombia (Guainía (Flórez & Ferro 2007), Meta, Putumayo, Vichada (Vivero et al. 2010a)), Bolivia, Brazil, French Guiana, Peru, Suriname, Venezuela. Refs.: CIPA group 1999 (tax., distr., nat. inf., ecol., refs.); Bejarano 2006a: 53 (Colombian Psychodidae checklist). columbiana (Ristorcelli & Van Ty), 1941: 263 (Phlebotomus). E., M., F. Type locality: Colombia, Nariño, El Capulí. Type material: ST F (current location unknown). Distr.: Colombia (Antioquia, Caldas, Cauca, Huila, ?La Guajira, Nariño, Risaralda, Tolima, Valle del Cauca). Refs.: CIPA group 1999 (tax., distr., nat. inf., ecol., refs.); Bejarano 2006a: 54 (Colombian Psychodidae checklist). incarum (Ristorcelli & Van Ty), 1941: 266 (as Phlebotomus monticolus var. incarum). conviti Ramírez-Pérez, Martins & Ramírez, 1976: 599. M., F. Type locality: Venezuela, Amazonas, Ocamo. Type material: HT M, AT F (CPqRR). Distr.: Colombia (Meta), Brazil (Pinheiro et al. 2010), Venezuela. Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 54 (Colombian Psychodidae checklist). cuzquena Martins, Llanos & Silva, 1975: 650. M. Type locality: Peru, Cusco, Paucartambo, Kosñipata, Pilcopata, left margin of Río Tono, Santa Alicia. Type material: HT M (CPqRR). Distr.: Colombia (Amazonas), Brazil, Peru, Venezuela. Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 52 (Colombian Psychodidae checklist). damascenoi (Mangabeira), 1941d: 369 (Flebotomus). M., F. Type locality: Brazil, Pará, Belém, Aurá. Type material: HT M, AT F (?IOC). Distr.: Colombia (Caquetá (Molina et al. 2000), Casanare (Molina et al. 2008), Meta), Brazil, French Guiana, Suriname (Kent et al. 2013). Refs.: CIPA group 1999 (tax., distr., nat. inf., ecol., refs.); Bejarano 2006a: 52 (Colombian Psychodidae checklist). spinosus (Floch & Abonnenc), 1942: 1 (Phlebotomus). dasymera (Fairchild & Hertig), 1961: 242 (Phlebotomus). L., M., F. Type locality: Panama, Canal Zone, Ft. Sherman Reservation, Mojinga Swamp. Type material: HT M, AT F (FSCA). Distr.: Colombia (Antioquia, Boyacá, Caldas (Contreras-Gutiérrez et al. 2014), Chocó, Nariño, Norte de Santander, Santander (Sandoval et al. 2006), Valle del Cauca), Belize, Brazil, Costa Rica, Ecuador, Mexico, Nicaragua, Panama, Venezuela. Refs.: Rosabal 1954: 11 (Phlebotomus sp. no. 16); CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 52 (Colombian Psychodidae checklist). davisi (Root), 1934: 242 (Phlebotomus). E., L., M., F. Type locality: Brazil, Pará, Fordlândia. Type material: HT M (USNM). Distr.: Colombia (Amazonas, Antioquia (Zuluaga et al. 2012), Caquetá, Guainía (Bejarano et al. 2007a), 196 · Zootaxa 4122 (1) © 2016 Magnolia Press BEJARANO & ESTRADA Guaviare (Cabrera et al. 2009), Meta, Norte de Santander, Putumayo, Santander (Sandoval et al. 2006), Vaupés), Bolivia, Brazil, Ecuador, French Guiana, Peru, Suriname, Venezuela. Refs.: CIPA group 1999 (tax., distr., nat. inf., ecol., refs.); Bejarano 2006a: 53 (Colombian Psychodidae checklist). rooti (Mangabeira), 1942a: 112 (Flebotomus). dendrophyla (Mangabeira), 1942a: 139 (Flebotomus). E., M., F. Type locality: Brazil, Pará, Belém, Aurá. Type material: HT M (IOC). Distr.: Colombia (Amazonas, Caquetá, Meta, Nariño (Travi et al. 1988), Putumayo, Valle del Cauca (Barreto et al. 1989), Vaupés), Bolivia, Brazil, Ecuador, French Guiana, Peru, Suriname, Venezuela. Refs.: CIPA group 1999 (tax., distr., nat. inf., ecol., refs.); Bejarano 2006a: 52 (Colombian Psychodidae checklist). disiuncta Morales, Osorno & Osorno-Mesa, 1974: 446. M., F. Type locality: Colombia, Cundinamarca, Suba. Type material: HT M, AT F (INS). Distr.: Colombia (Cesar, Cundinamarca). Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 54 (Colombian Psychodidae checklist). dreisbachi (Causey & Damasceno), 1945b: 649 (Flebotomus). M., F. Type locality: Brazil, Amazonas, Coari, Rio Solimões. Type material: HT M (USNM). Distr.: Colombia (Amazonas, Caquetá), Bolivia (Le Pont et al. 1992), Brazil, Ecuador (Jones et al. 2010), French Guiana, Peru, Suriname (Burgos & Hudson 1994), Venezuela. Refs.: Floch & Abonnenc 1945b: 9 (Phlebotomus sp. from Crique Anguille); CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 50 (Colombian Psychodidae checklist). dubitans (Sherlock), 1962: 324 (Phlebotomus). M., F. Type locality: Colombia, Santander, San Vicente de Chucurí. Type material: HT M (USNM). Distr.: Colombia (Antioquia, Atlántico (Maestre-Serrano et al. 2010), Arauca (Turriago et al. 2007), Bolívar (Cortés 2006), Boyacá, Córdoba, Cundinamarca, Huila, La Guajira, Magdalena (Bejarano et al. 2015), Nariño (Travi et al. 1988), Norte de Santander, Santander, Sucre, Tolima, Valle del Cauca), Brazil, Costa Rica, Panama, Trinidad and Tobago, Venezuela. Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 51 (Colombian Psychodidae checklist). duncanae Le Pont, Martínez, Torrez-Espejo & Durjardin, 1998: 170. M., F. Type locality: Bolivia, La Paz, Nor Yungas, Carrasco, Unión Berea III. Type material: HT M, AT F, 3 PT (1 M and 2 F) (MNHNB); 5 PT (2 M and 3 F) (IRD). Distr.: Colombia (Casanare (Young & Morales 1987)), Bolivia, Peru (Galati 2013). Refs.: Velasco 1973: 88 (Lutzomyia sp. D); Young & Morales 1987: 662 (Lutzomyia sp. 1). dysponeta (Fairchild & Hertig), 1952: 505 (Phlebotomus). M., F. Type locality: Panama, Canal Zone, Juan Mina, Río Chagres between Gamboa and Madden Dam. Type material: HT M, AT F (FSCA). Distr.: Colombia (Boyacá, Chocó, Guaviare (Cabrera et al. 2009), Magdalena (Bejarano et al. 2015), Meta, Nariño, Tolima (Romero-Ricardo et al. 2012)), Brazil, Costa Rica, Ecuador, Panama, Venezuela. Refs.: Rosabal 1954: 10 (Phlebotomus sp. no. 3); CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 52 (Colombian Psychodidae checklist). emberai Bejarano, Duque & Vélez, 2004: 837. F. Type locality: Colombia, Chocó, Bahía Solano, El Brazo. Type material: HT F, PT F (VHET). Distr.: Colombia (Chocó). Refs.: Bejarano 2006a: 54 (Colombian Psychodidae checklist). erwindonaldoi (Ortíz), 1978: 205 (Phlebotomus). M., F. Type locality: Venezuela, Zulia, Rómulo Gallegos, Caja Seca. Type material: HT M (current location unknown). Distr.: Colombia (Huila (Pardo-Puentes 2006), Santander (Sandoval et al. 2006), Tolima (Marín-Casas et al. 2008)), Peru (Cáceres & Galati 2001), Venezuela. Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 50 (Colombian Psychodidae checklist). evangelistai Martins & Frahia, 1971: 361. M., F. Type locality: Brazil, Pará, Belém, Utinga. Type material: HT M, PT F (CPqRR). Distr.: Colombia (Amazonas, Guaviare (Cabrera et al. 2009), Putumayo), Bolivia, Brazil, Peru. Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 51 (Colombian Psychodidae checklist). CATALOGUE OF PSYCHODIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 197 evansi (Núñez-Tovar), 1924: 44 (Phlebotomus). E., L., P., M., F. Type locality: Venezuela, Carabobo, Mariara. Type material: ST M (current location unknown). Distr.: Colombia (Antioquia, Atlántico (Maestre-Serrano et al. 2010), Bolívar (incl. Isla Fuerte (Gallego & Vélez 1994)), Casanare, Cesar (Estrada et al. 2015), Córdoba, La Guajira, Magdalena, Meta (Molina 1997), Norte de Santander, Santander, Sucre), Costa Rica, El Salvador, Guatemala, Honduras, Mexico (Ibáñez-Bernal et al. 2004), Nicaragua, ?Peru (Cabanillas et al. 2001), Venezuela. Refs.: CIPA group 1999 (tax., distr., nat. inf., ecol., refs.); Bejarano 2006a: 54 (Colombian Psychodidae checklist). Material examined. COLOMBIA: Cesar: Mpio. [Municipio] Valledupar, Balneario Hurtado, 7 F., 6.vii.2013, O.A. Aponte leg., CDC light trap (CZUS); 2 M., 5 F., same data except 19.vii.2013. fairtigi Martins, 1970: 279. M., F. Type locality: Colombia, Meta, Villavicencio. Type material: HT M (FSCA). Distr.: Colombia (Caquetá, Casanare, Guaviare (Cabrera et al. 2009), Meta, Vaupés). Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 53 (Colombian Psychodidae checklist). falcata Young, Morales & Ferro in Young & Duncan, 1994: 61. M., F. Type locality: Colombia, Amazonas, Leticia. Type material: HT M, AT F (INS). Distr.: Colombia (Amazonas), Brazil, Ecuador, Peru (Valdivia et al. 2012). Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 51 (Colombian Psychodidae checklist). ferroae Young & Morales, 1987: 651. M., F. Type locality: Colombia, Norte de Santander, Arboledas, Potreros. Type material: HT M, AT F (INS). Distr.: Colombia (Norte de Santander). Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 53 (Colombian Psychodidae checklist). flaviscutellata (Mangabeira), 1942a: 144 (Flebotomus). E., L., P., M., F. Type locality: Brazil, Pará, Belém, Aurá. Type material: HT M (IOC). Distr.: Colombia (Amazonas, Antioquia, Casanare, Chocó, Guaviare (Cabrera et al. 2009), Meta, Norte de Santander, Putumayo, Vichada (Vivero et al. 2010a)), Bolivia, Brazil, Ecuador, French Guiana, Peru, Suriname, Trinidad and Tobago, Venezuela. Refs.: CIPA group 1999 (tax., distr., nat. inf., ecol., refs.); Bejarano 2006a: 51 (Colombian Psychodidae checklist). apicalis (Floch & Abonnenc), 1943: 25 (Phlebotomus). furcata (Mangabeira), 1941c: 256 (Flebotomus). ?L,?P, M, F. Type locality: Brazil, Pará, Belém, Aurá. Type material: HT M (?IOC). Distr.: Colombia (Amazonas, Caquetá, Guaviare (Cabrera et al. 2009), Meta, Nariño, Santander, Valle del Cauca), Bolivia, Brazil, Costa Rica, Ecuador (Alexander et al. 1992), French Guiana, Peru, Venezuela. Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 54 (Colombian Psychodidae checklist). arborealis (Floch & Abonnenc), 1944b: 8 (Phlebotomus). geniculata (Mangabeira), 1941b: 245 (Flebotomus). L., P., M., F. Type locality: Brazil, Pará, Belém, Aurá. Type material: not extant (Shimabukuro & Galati 2011). Distr.: Colombia (Amazonas, Caquetá, Chocó, Guaviare (Cabrera et al. 2009), Meta, Putumayo), Belize, Bolivia, Brazil, Costa Rica, Ecuador, French Guiana, Guatemala, Nicaragua (Zeledón & Murillo 1983, Collantes & Martínez-Ortega 1998), Panama, Peru, Venezuela. Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 53 (Colombian Psychodidae checklist). gomezi (Nitzulescu), 1931: 247 (Phlebotomus). E., L., P., M., F. Type locality: Venezuela, Táchira, San Cristobal. Type material: HT F (current location unknown). Distr.: Colombia (Amazonas, Antioquia, Atlántico (MaestreSerrano et al. 2010), Arauca (Turriago et al. 2007), Bolívar (incl. Isla Fuerte (Gallego & Vélez 1994)), Boyacá, Caldas, Caquetá, Casanare (Manotas-Solano et al. 2012), Cesar, Chocó, Córdoba, Cundinamarca, Guaviare (Cabrera et al. 2009), Huila, La Guajira, Magdalena, Meta, Nariño, Norte de Santander, Putumayo, Quindío (Contreras-Gutiérrez et al. 2014), Risaralda, Santander, Sucre, Tolima, Valle del Cauca, Vichada), Bolivia, Brazil, Costa Rica, Ecuador, El Salvador, French Guiana, Guatemala (Vivero et al. 2011a), Honduras, Mexico (IbáñezBernal 1999), Nicaragua, Panama, Peru, Trinidad and Tobago, Venezuela. Refs.: CIPA group 1999 (tax., distr., nat. inf., ecol., refs.); Bejarano 2006a: 51 (Colombian Psychodidae checklist). suis (Rozeboom), 1940: 8 (Phlebotomus). japignyi (Floch & Abonnenc), 1944c: 2 (Phlebotomus). 198 · Zootaxa 4122 (1) © 2016 Magnolia Press BEJARANO & ESTRADA gorbitzi (Blancas), 1959: 126 (Phlebotomus). P., M., F. Type locality: Peru, Ancash, Cañon del Pato. Type material: HT M, AT F (?ICGES). Distr.: Colombia (Nariño, Sucre (Vivero et al. 2013)), Costa Rica, Ecuador, Panama, Peru. Refs.: Rosabal 1954: 10 (Phlebotomus sp. no. 11); CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 50 (Colombian Psychodidae checklist). hansoni (Fairchild & Hertig), 1961: 244 (Phlebotomus). hartmanni (Fairchild & Hertig), 1957: 328 (Phlebotomus). L., P., M., F. Type locality: Panama, Panamá, Cerro Campana. Type material: HT M, AT F (FSCA). Distr.: Colombia (Antioquia, Boyacá, Caldas, Cauca (incl. Isla Gorgona (Montoya-Lerma & Baena 1995)), Chocó, Cundinamarca, Nariño, Norte de Santander (Gálvis-Ovallos et al. 2013), Risaralda, Santander, Tolima, Valle del Cauca), Costa Rica, Ecuador, Panama, Peru. Refs.: CIPA group 1999 (tax., distr., nat. inf., ecol., refs.); Bejarano 2006a: 50 (Colombian Psychodidae checklist). hernandezi (Ortíz), 1965c: 412 (Phlebotomus). L., M., F. Type locality: Venezuela, Trujillo, Pan-American Highway between Betijoque, Isnotú and Caja Seca. Type material: HT M (?INH). Distr.: Colombia (Norte de Santander), Peru (Pérez et al. 2011), Venezuela. Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 51 (Colombian Psychodidae checklist). hirsuta (Mangabeira), 1942a: 116 (Flebotomus). M., F. Type locality: Brazil, Pará, Abaeté, Piratuba. Type material: HT M (?IOC). Distr.: Colombia (Amazonas, Caquetá, Guaviare (Cabrera et al. 2009), Meta, Putumayo, Vaupés), Bolivia, Brazil, Ecuador, French Guiana, Peru, Suriname. Refs.: Floch 1947: 152 (Phlebotomus sp. II); Floch & Chassignet 1948: 1 (Phlebotomus sp. 780); Velasco 1973: 84 (Lutzomyia sp. C); CIPA group 1999 (tax., distr., nat. inf., ecol., refs.); Bejarano 2006a: 53 (Colombian Psychodidae checklist); Carvalho et al. 2006: 131 (raised to the category of species). colasbelcouri (Floch & Chassignet), 1947a: 1 (Phlebotomus). howardi Young, 1979: 144. M., F. Type locality: Colombia, Amazonas, 17 km W of Leticia. Type material: HT M (FSCA). Distr.: Colombia (Amazonas, Putumayo), Brazil (Galati 2013), Peru (Pérez et al. 1991, Cáceres et al. 2000). Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 53 (Colombian Psychodidae checklist). ignacioi Young, 1972: 312. M., F. Type locality: Venezuela, Mérida, Mérida. Type material: HT M, AT F (FSCA). Distr.: Colombia (Cundinamarca), Venezuela. Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 54 (Colombian Psychodidae checklist). infraspinosa (Mangabeira), 1941a: 216 (Flebotomus). M., F. Type locality: Brazil, Pará, Belém, Aurá. Type material: HT M (IOC). Distr.: Colombia (Amazonas, Santander), Bolivia, Brazil, French Guiana, ?Peru (Young et al. 1985), Suriname, Venezuela (Galati 2013). Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 50 (Colombian Psychodidae checklist). isovespertilionis (Fairchild & Hertig), 1958: 516 (Phlebotomus). L., P., M., F. Type locality: Panama, Canal Zone, km 19 road on a tributary of Río Cocoli. Type material: HT M, AT F (FSCA). Distr.: Colombia (Antioquia, Chocó), Costa Rica, Panama. Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 50 (Colombian Psychodidae checklist). lerayi Le Pont, Martínez, Torrez-Espejo & Dujardin, 1998: 166. M. Type locality: Bolivia, La Paz, Sud Yungas, Cajuata, Río Suri. Type material: HT M, 2 PT M (MNHNB); 2 PT M (IRD). Distr.: Colombia (Tolima (MarínCasas et al. 2008)), Bolivia. lichyi (Floch & Abonnenc), 1950: 1 (Phlebotomus). E., L., P., M., F. Type locality: Venezuela, Carabobo, Río Borburata. Type material: HT F (?IPG). Distr.: Colombia (Antioquia, Caldas, Chocó, Cundinamarca, Huila, Magdalena (Young 1979), Meta, Nariño, Norte de Santander, Quindío (Alexander et al. 2001a), Risaralda, Santander, Tolima, Valle del Cauca), Brazil, Costa Rica, Ecuador, French Guiana, Panama, Peru, Trinidad and CATALOGUE OF PSYCHODIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 199 Tobago, Venezuela. Refs.: CIPA group 1999 (tax., distr., nat. inf., ecol., refs.); Bejarano 2006a: 51 (Colombian Psychodidae checklist). foliatus (Mirsa & Ortíz), 1952: 249 (Phlebotomus). vexillarius (Fairchild & Hertig), 1952: 514 (Phlebotomus). limafalcaoae (Wolff & Galati), 2002: 317. M., F. (Pintomyia). Type locality: Colombia, Antioquia, Montebello, Savanitas. Type material: HT M, AT F (CEUA). Distr.: Colombia (Antioquia). Refs.: Bejarano 2006a: 54 (Colombian Psychodidae checklist). longiflocosa Osorno-Mesa, Morales, Osorno & Muñoz, 1970: 1. E., M., F. Type locality: Colombia, Huila, Tello, San Andrés, Romero. Type material: HT M, AT F (INS). Distr.: Colombia (Cundinamarca (Ferro et al. 2007), Huila, Norte de Santander (Cárdenas et al. 2005), Tolima). Refs.: CIPA group 1999 (tax., distr., nat. inf., ecol., refs.); Bejarano 2006a: 54 (Colombian Psychodidae checklist). longipalpis (Lutz & Neiva), 1912: 90 (Phlebotomus). E., L., P., M., F. Type locality: Brazil (several localities cited in original description: São Paulo, São Paulo, Bosque da Saúde; Minas Gerais, Benjamin Constant, and Cordisburgo, Gruta do Maquiné). Type material: ST F, M (?IOC). Distr.: Colombia (Antioquia, Caldas, Cauca (Barreto et al. 2006), Córdoba, Cundinamarca, Huila, La Guajira, Norte de Santander, Santander, Sucre, Tolima), Argentina, Bolivia, Brazil, Costa Rica, El Salvador, Guatemala, Honduras, Mexico, Nicaragua, Panama, Paraguay, Uruguay (Salomón et al. 2011), Venezuela. Refs.: CIPA group 1999 (tax., distr., nat. inf., ecol., refs.); Bejarano 2006a: 51 (Colombian Psychodidae checklist). almazani (Galliard), 1934b: 193 (Phlebotomus). otamae (Núñez-Tovar), 1924: 44 (Phlebotomus). longispina (Mangabeira), 1942a: 186 (Flebotomus). E., L., M., F. Type locality: Brazil, Belém, Pará, Aurá. Type material: HT M (IOC). Distr.: Colombia (Caquetá, Meta (Molina et al. 1997)), Brazil, French Guiana, Paraguay, Venezuela. Refs.: CIPA group 1999 (tax., distr., nat. inf., ecol., refs.); Bejarano 2006a: 53 (Colombian Psychodidae checklist). lutziana (Costa Lima), 1932: 48 (Phlebotomus). M., F. Type locality: Brazil, Minas Gerais, Lassance. Type material: ST M (?IOC). Distr.: Colombia (Amazonas, Meta (Bejarano et al. 2007b)), Bolivia, Brazil, French Guiana, Paraguay, Peru, Suriname, Venezuela. Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 52 (Colombian Psychodidae checklist). marinkellei Young, 1979: 59. M., F. Type locality: Colombia, Caquetá, Tres Esquinas. Type material: HT M, AT F (FSCA). Distr.: Colombia (Amazonas, Caquetá), Brazil. Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 51 (Colombian Psychodidae checklist). martinezi Young & Morales, 1987: 653. M., F. Type locality: Colombia, Meta, Río Iteviare. Type material: HT M, AT F (INS). Distr.: Colombia (Meta), ?Ecuador (Gomez et al. 2014). Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 53 (Colombian Psychodidae checklist). micropyga (Mangabeira), 1942a: 132 (Flebotomus). P., M., F. Type locality: Brazil, Pará, Belém, Aurá. Type material: ST M (IOC). Distr.: Colombia (Amazonas, Antioquia, Bolívar, Boyacá, Caldas, Casanare, Cesar (Estrada et al. 2015), Chocó, Córdoba, Cundinamarca, Guaviare (Cabrera et al. 2009), La Guajira, Magdalena, Meta, Nariño, Norte de Santander, Putumayo, Santander, Sucre, Tolima, Valle del Cauca), Bolivia, Brazil, Costa Rica, Ecuador, French Guiana, Panama, Peru, Trinidad and Tobago, Venezuela. Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 51 (Colombian Psychodidae checklist). Material examined. COLOMBIA: Cesar: Mpio. [Municipio] Valledupar, Balneario Hurtado, 3 F., 6.vii.2013, O.A. Aponte leg., CDC light trap (CZUS). migonei (França), 1920: 230 (Phlebotomus). E., L., P., M., F. Type locality: Paraguay, Asunción. Type material: ST 200 · Zootaxa 4122 (1) © 2016 Magnolia Press BEJARANO & ESTRADA M (current location unknown). Distr.: Colombia (Casanare (Molina et al. 1997), Córdoba, La Guajira (ZambranoHernández et al. 2005), Magdalena, Sucre (Le Pape 1992)), Argentina, Bolivia, Brazil, Ecuador (Gomez et al. 2014), Paraguay, Peru, Trinidad and Tobago, Venezuela. Refs.: CIPA group 1999 (tax., distr., nat. inf., ecol., refs.); Bejarano 2006a: 51 (Colombian Psychodidae checklist). rangeli (Núñez-Tovar), 1924: 45 (Phlebotomus). araozi (Paterson & Shannon), 1926: 236 (Phlebotomus). monstruosa (Floch & Abonnenc), 1944a: 1 (Phlebotomus). M., F. Type locality: French Guiana, Baduel. Type material: HT M (IP). Distr.: Colombia (Amazonas, Caquetá), Brazil, French Guiana, Peru (Pérez et al. 2008), Suriname, Venezuela. Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 50 (Colombian Psychodidae checklist). falciformis (Floch & Abonnenc), 1944c: 8 (Phlebotomus). moralesi Young, 1979: 80. M., F. Type locality: Colombia, Valle del Cauca, 15 km SW of Cali, Río Pance. Type material: HT M, AT F (FSCA). Distr.: Colombia (Valle del Cauca). Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 54 (Colombian Psychodidae checklist). nematoducta Young & Arias, 1984: 428. M., F. Type locality: Brazil, Amazonas, 26 km E of Manaus, Reserva Ducke. Type material: HT M, AT F (USNM). Distr.: Colombia (Guaviare (Cabrera et al. 2009), Meta (VásquezTrujillo et al. 2013)), Brazil, Peru (Valdivia et al. 2012). Refs.: CIPA group 1999 (tax., distr., ecol., refs.). nevesi (Damasceno & Arouck), 1956: 9 (Flebotomus). M., F. Type locality: Brazil, Pará, Capim. Type material: HT M (FSPUSP). Distr.: Colombia (Amazonas, Caquetá, Meta, Putumayo), Bolivia, Brazil, Ecuador, Peru. Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 54 (Colombian Psychodidae checklist). nocticola Young, 1973: 109. M., F. Type locality: Colombia, Antioquia, 24 km SW of Zaragoza, Río Anorí. Type material: HT M, AT F (FSCA). Distr.: Colombia (Amazonas, Antioquia, Casanare (Molina et al. 1997), Putumayo, Santander (Sandoval et al. 2006)), Bolivia, Ecuador, French Guiana, Mexico (Cruz-Ruíz et al. 1994), Panama, Peru. Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 53 (Colombian Psychodidae checklist). nuneztovari (Ortíz), 1954: 232 (Phlebotomus). M., F. Type locality: Venezuela, Lara, Duaca. Type material: HT M (?MBUCV). Distr.: Colombia (Antioquia, Boyacá, Caldas, Cundinamarca, Huila, Norte de Santander, Santander (Sandoval et al. 2006), Tolima), Bolivia, Guatemala, Honduras, Panama, Peru, Venezuela. Refs.: Velasco 1973: 78 (Lutzomyia sp. I); CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 54 (Colombian Psychodidae checklist). anglesi Le Pont & Desjeux, 1984: 277 (as Lutzomyia nuneztovari anglesi). oresbia (Fairchild & Hertig), 1961: 240 (Phlebotomus). M, F. Type locality: Panama, Chiriquí, Volcán, Santa Clara. Type material: HT M, AT F (USNM). Distr.: Colombia (Huila (Pardo-Puentes 2006), Norte de Santander (Gálvis-Ovallos et al. 2013), Tolima (Marín-Casas et al. 2008)), Costa Rica, Panama. Refs.: CIPA group 1999 (tax., distr., ecol., refs.). osornoi (Ristorcelli & Van Ty), 1941: 260 (Phlebotomus). M., F. Type locality: Colombia, Nariño, El Capulí. Type material: HT F (current location unknown). Distr.: Colombia (Nariño), Bolivia, Ecuador, Peru. Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 50 (Colombian Psychodidae checklist). montoyai (Sherlock), 1962: 328 (Phlebotomus). ovallesi (Ortíz), 1952: 155 (Phlebotomus). E., L., P., M., F. Type locality: Venezuela, Lara, Duaca. Type material: HT M (?INH). Distr.: Colombia (Amazonas, Antioquia, Boyacá, Caldas, Casanare, Chocó, Cundinamarca, Magdalena, Norte de Santander, Santander, Sucre, Tolima (Morales et al. 1981)), Belize, Costa Rica, Guatemala, Honduras, Mexico, Nicaragua, Panama, Trinidad and Tobago, Venezuela. Refs.: CIPA group 1999 (tax., distr., nat. inf., ecol., refs.); Bejarano 2006a: 54 (Colombian Psychodidae checklist). CATALOGUE OF PSYCHODIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 201 pabloi Barreto, Burbano & Young, 2002: 601. M., F. Type locality: Colombia, Putumayo, Villagarzón, San Vicente. Type material: HT M, AT F, 6 PT M (UVS). Distr.: Colombia (Putumayo), Ecuador (Jones et al. 2010). Refs.: Bejarano 2006a: 53 (Colombian Psychodidae checklist). pajoti Abonnenc, Léger & Fauran, 1979: 75 (as Lutzomyia yuilli pajoti). M., F. Type locality: French Guiana, Saül. Type material: HT M, AT F (IP). Distr.: Colombia (Guainía (Flórez & Ferro 2007), Guaviare (Cabrera et al. 2009), Putumayo), Brazil, Ecuador (Terán-Soto 2006), French Guiana, Peru, Suriname. Refs.: Floch & Abonnenc 1944d: 9 (Phlebotomus sp. de Souvenir); CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 51 (Colombian Psychodidae checklist). panamensis (Shannon), 1926: 192 (Phlebotomus). E., L., P., M., F. Type locality: Panama, Canal Zone, Caño Saddle. Type material: HT M, F (USNM). Distr.: Colombia (Antioquia, Atlántico (Maestre-Serrano et al. 2010), Bolívar (Gallego et al. 1994), Boyacá, Caldas, Caquetá, Casanare, Cauca, Chocó, Córdoba, Guaviare (Cabrera et al. 2009), La Guajira, Magdalena (Bejarano et al. 2015), Meta, Nariño, Norte de Santander, Putumayo, Risaralda, Santander, Sucre, Tolima, Valle del Cauca), Belize, Brazil, Costa Rica, Ecuador, French Guiana, Guatemala, Honduras, Mexico, Nicaragua, Panama, ?Peru, Venezuela. Refs.: CIPA group 1999 (tax., distr., nat. inf., ecol., refs.); Bejarano 2006a: 53 (Colombian Psychodidae checklist). Material examined. COLOMBIA: Magdalena: Mpio. [Municipio] Santa Marta, Minca, 600m, 2 M., 4 F., 1–2.xi.2008, A. Pérez-Doria leg., CDC light trap (CZUS). paraensis (Costa Lima), 1941: 7 (Flebotomus). E., L., M., F. Type locality: Brazil, Pará, Belém, Aurá. Type material: HT M (IOC). Distr.: Colombia (Amazonas, Caquetá, Putumayo), Bolivia (Le Pont et al. 1992), Brazil, Ecuador, French Guiana, Peru, Suriname, Venezuela. Refs.: CIPA group 1999 (tax., distr., nat. inf., ecol., refs.); Bejarano 2006a: 53 (Colombian Psychodidae checklist). pia (Fairchild & Hertig), 1961: 248 (Phlebotomus). M., F. Type locality: Panama, Chiriquí, El Volcán, Santa Clara. Type material: HT M, AT F (FSCA). Distr.: Colombia (Antioquia, Boyacá, Caldas, Cundinamarca, Huila, Meta, Nariño, Norte de Santander, Risaralda, Tolima, Valle del Cauca), Bolivia, Costa Rica, Panama, Peru, Venezuela. Refs.: CIPA group 1999 (tax., distr., nat. inf., ecol., refs.); Bejarano 2006a: 54 (Colombian Psychodidae checklist). pilosa (Damasceno & Causey), 1944: 342 (Flebotomus). M., F. Type locality: Brazil, Pará, Belém, Santa Isabel. Type material: HT M (USNM). Distr.: Colombia (Amazonas, Antioquia, Bolívar, Boyacá, Caquetá, Chocó, Cundinamarca, Huila, Nariño (Alexander et al. 1995), Norte de Santander, Putumayo, Santander, Tolima), Brazil, Costa Rica, French Guiana, Panama, Trinidad and Tobago, Venezuela. Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 52 (Colombian Psychodidae checklist). preclara Young & Arias, 1984: 429. M., F. Type locality: Colombia, Amazonas, 9 km N of Leticia. Type material: HT M (FSCA). Distr.: Colombia (Amazonas, Guaviare (Cabrera et al. 2009), Meta, Vichada (Vivero et al. 2010a)), Bolivia, Brazil (Gomes et al. 2009), Peru. Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 53 (Colombian Psychodidae checklist). punctigeniculata (Floch & Abonnenc), 1944b: 5 (Phlebotomus). M., F. Type locality: French Guiana, Cayenne, Baduel. Type material: ST M, F (IP). Distr.: Colombia (Atlántico (Maestre-Serrano et al. 2010), Bolívar, Boyacá, Caldas, Caquetá, Casanare, Chocó, Cundinamarca, Huila, La Guajira, Magdalena, Meta (Vásquez-Trujillo et al. 2013), Nariño, Putumayo, Tolima), Argentina (Salomón 1994), Bolivia, Brazil, Ecuador (Jones et al. 2010), French Guiana, Panama, Peru, Suriname, Venezuela. Refs.: CIPA group 1999 (tax., distr., nat. inf., ecol., refs.); Bejarano 2006a: 52 (Colombian Psychodidae checklist). christophersoni (Damasceno & Causey), 1944: 347 (Flebotomus). quasitownsendi Osorno, Osorno-Mesa & Morales, 1972c: 10. M., F. Type locality: Colombia, Santander, Barbosa, El Banqueo. Type material: HT M, AT F (INS). Distr.: Colombia (Boyacá, Cundinamarca (Santamaría et al. 2003), 202 · Zootaxa 4122 (1) © 2016 Magnolia Press BEJARANO & ESTRADA Norte de Santander, Santander). Refs.: CIPA group 1999 (tax., distr., nat. inf., ecol., refs.); Bejarano 2006a: 54 (Colombian Psychodidae checklist). rangeliana (Ortíz), 1952: 153 (Phlebotomus). M., F. Type locality: Venezuela, Lara, Duaca, Reserva Forestal El Buco. Type material: HT M (?INH). Distr.: Colombia (Atlántico (Maestre-Serrano et al. 2010), Bolívar, Córdoba, Cesar (Estrada et al. 2015), Cundinamarca, La Guajira, Magdalena, Sucre, Tolima), Panama, Trinidad and Tobago, Venezuela. Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 54 (Colombian Psychodidae checklist). Material examined. COLOMBIA: Cesar: Mpio. [Municipio] Valledupar, Balneario Hurtado, 1 F., 6.vii.2013, O.A. Aponte leg., CDC light trap (CZUS). reburra (Fairchild & Hertig), 1961: 246 (Phlebotomus). M., F. Type locality: Panama, Panamá, Cerro Azul, La Zumbadora. Type material: HT M, AT F (FSCA). Distr.: Colombia (Antioquia (Vergara et al. 2008a), Cauca, Chocó, Nariño, Valle del Cauca), Costa Rica, Ecuador, Panama. Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 53 (Colombian Psychodidae checklist). recurva Young, 1973: 106. M., F. Type locality: Colombia, Chocó, 3 km SE of the mouth of Río Curiche and 1 km inland from Humboldt Bay. Type material: HT M, AT F (FSCA). Distr.: Colombia (Chocó), Panama, Peru (Pérez et al. 2008). Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 53 (Colombian Psychodidae checklist). reducta Feliciangeli, Ramírez-Pérez & Ramírez, 1988: 54 (as Lutzomyia olmeca reducta). M., F. Type locality: Venezuela, Amazonas, Cerro La Neblina. Type material: HT M, PT F (CNRFV). Distr.: Colombia (Amazonas), Brazil, Peru (Ogusuku et al. 2001), Venezuela. Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 51 (Colombian Psychodidae checklist). richardwardi Ready & Fraiha, 1981: 705. E., M., F. Type locality: Brazil, Pará, km 43 of the Trans-Amazonian Highway, between Altamira and Itaituba. Type material: HT F, PT M (IEC). Distr.: Colombia (Amazonas, Guaviare (Cabrera et al. 2009), Meta, Putumayo, Vaupés), Bolivia, Brazil, Ecuador (Jones et al. 2010), Peru. Refs.: Ward & Killick-Kendrick 1974: 216 (Lutzomyia sp no. 260.44); CIPA group 1999 (tax., distr., nat. inf., ecol., refs.); Bejarano 2006a: 51 (Colombian Psychodidae checklist). rorotaensis (Floch & Abonnenc), 1944e: 4 (Phlebotomus). M., F. Type locality: French Guiana, Rorota. Type material: ST M, F (IPG). Distr.: Colombia (Antioquia, Chocó, Meta (Cabrera et al. 2009), Nariño, Valle del Cauca), Brazil, French Guiana, Panama, Peru, Suriname, Venezuela. Refs.: Floch & Abonnenc 1941b: 10 (Phlebotomus sp. from Rorota); CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 52 (Colombian Psychodidae checklist). rosabali (Fairchild & Hertig), 1956: 310 (Phlebotomus). M., F. Type locality: Panama, Chiriquí, Puerto Armuelles. Type material: HT M, AT F (FSCA). Distr.: Colombia (Antioquia, Cauca, Nariño, Valle del Cauca), Costa Rica, Panama. Refs.: Rosabal 1954: 10 (Phlebotomus sp. no. 9); CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 50 (Colombian Psychodidae checklist). ruii Arias & Young, 1982: 249. M., F. Type locality: Brazil, Amazonas, 26 km E of Manaus, Reserva Ducke. Type material: HT M, AT F (INPA). Distr.: Colombia (Amazonas), Brazil. Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 53 (Colombian Psychodidae checklist). runoides (Fairchild & Hertig), 1953: 30 (Phlebotomus). L., M., F. Type locality: Panama, Bocas del Toro, Almirante. Type material: HT M, AT F (FSCA). Distr.: Colombia (Amazonas, Antioquia, Caquetá, Nariño, Valle del Cauca, Vichada (Vivero et al. 2010a)), Brazil, Costa Rica, Ecuador (Jones et al. 2010), Panama, Peru. Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 50 (Colombian Psychodidae checklist). CATALOGUE OF PSYCHODIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 203 saltuosa Young, 1979: 148. M. Type locality: Colombia, Amazonas, 17 km W of Leticia. Type material: HT M (FSCA). Distr.: Colombia (Amazonas). Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 53 (Colombian Psychodidae checklist). sanguinaria (Fairchild & Hertig), 1957: 332 (Phlebotomus). E., L., P., M., F. Type locality: Panama, Bocas del Toro, Almirante. Type material: HT M, AT F (FSCA). Distr.: Colombia (Chocó, Valle del Cauca), Costa Rica, Ecuador (Hashiguchi et al. 2014), Honduras, Nicaragua, Panama. Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 50 (Colombian Psychodidae checklist). saulensis (Floch & Abonnenc), 1944a: 11 (Phlebotomus). L., M., F. Type locality: French Guiana, Haute Maná, Saül. Type material: HT M (IP). Distr.: Colombia (Amazonas, Antioquia, Caquetá, Cauca, Chocó, Guaviare (Cabrera et al. 2009), Meta, Nariño, Santander, Tolima, Vichada (Vivero et al. 2010a), Valle del Cauca), Bolivia, Brazil, Costa Rica, Ecuador, French Guiana, Panama, Peru, Venezuela. Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 53 (Colombian Psychodidae checklist). pinealis (Floch & Abonnenc), 1944b: 11 (Phlebotomus). sauroida Osorno-Mesa, Morales & Osorno, 1972a: 433. M., F. Type locality: Colombia, Boyacá, Santa Ana, San Pedro. Type material: HT M, AT F (INS). Distr.: Colombia (Boyacá, Norte de Santander), Venezuela. Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 54 (Colombian Psychodidae checklist). scaffi (Damasceno & Arouck), 1956: 6 (Flebotomus). M., F. Type locality: Brazil, Pará, Capim. Type material: HT M (FSPUSP). Distr.: Colombia (Amazonas, Caquetá, Guaviare (Cabrera et al. 2009)), Bolivia (Le Pont et al. 1992), Brazil, Ecuador (Terán-Soto 2006), French Guiana, Peru, Suriname. Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 52 (Colombian Psychodidae checklist). scorzai (Ortíz), 1965a: 28 (Phlebotomus). M., F. Type locality: Venezuela, Aragua, Rancho Grande. Type material: HT M, AT F (current location unknown). Distr.: Colombia (Antioquia (Vergara et al. 2008a), Caldas, Cauca, Cundinamarca, Nariño, Norte de Santander (Gálvis-Ovallos et al. 2013), Risaralda, Tolima, Valle del Cauca), Venezuela, Peru (Cáceres et al. 2000). Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 50 (Colombian Psychodidae checklist). sericea (Floch & Abonnenc), 1944a: 6 (Phlebotomus). M., F. Type locality: French Guiana, Baduel. Type material: HT M, PT M (IP). Distr.: Colombia (Guaviare (Cabrera et al. 2009)), Brazil, Ecuador, French Guiana, Suriname, Venezuela. Refs.: CIPA group 1999 (tax., distr., ecol., refs.). deanei (Damasceno, Causey & Arouck), 1945: 4 (Flebotomus). serrana (Damasceno & Arouck), 1949: 843 (Flebotomus). L., P., M., F. Type locality: Brazil, Pará, Vizeu, Piriábas. Type material: HT M (FSPUSP). Distr.: Colombia (Antioquia, Bolívar, Boyacá, Caldas, Chocó, Córdoba, Cundinamarca, Meta, Nariño, Norte de Santander, Santander, Sucre, Tolima, Valle del Cauca), Belize (IbáñezBernal 2001), Bolivia, Brazil, Costa Rica, Ecuador, French Guiana, Guatemala, Honduras, Mexico, Nicaragua, Panama, Peru, Venezuela. Refs.: Rosabal 1954: 11 (Phlebotomus sp. no. 5); CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 54 (Colombian Psychodidae checklist). guayasi (Rodríguez), 1956: 77 (Phlebotomus). shannoni (Dyar), 1929: 121 (Phlebotomus). E., L., P., M., F. Type locality: Panama, Canal Zone, Lago Gatún, Caño Saddle. Type material: LT M (USNM). Distr.: Colombia (Amazonas, Antioquia, Atlántico (Maestre-Serrano et al. 2010), Bolívar, Boyacá, Caldas, Caquetá, Chocó, Córdoba, Cundinamarca, Guaviare (Cabrera et al. 2009), La Guajira, Magdalena, Meta, Nariño, Norte de Santander, Quindío (Contreras-Gutiérrez et al. 2014), Santander, Sucre, Tolima, Valle del Cauca), Argentina, Belize, Bolivia, Brazil, Costa Rica, Ecuador, French Guiana, Guatemala, Honduras, Mexico, Nicaragua, Panama, Paraguay, Peru, Suriname, Trinidad and Tobago, United States of America, Venezuela. Refs.: CIPA group 1999 (tax., distr., nat. inf., ecol., refs.); Bejarano 2006a: 52 (Colombian Psychodidae checklist); Sábio et al. 2014: 332 (tax.). 204 · Zootaxa 4122 (1) © 2016 Magnolia Press BEJARANO & ESTRADA microcephalus (Barretto & Duret), 1953: 341 (Phlebotomus). pifanoi (Ortíz), 1972: 21 (Phlebotomus). shawi Fraiha, Ward & Ready, 1981: 699. E., M., F. Type locality: Brazil, Pará, Marabá, Serra dos Carajás. Type material: HT F, PT M (IEC). Distr.: Colombia (Amazonas, Guaviare, Meta, Vaupés), Bolivia, Brazil, Peru. Refs.: Ward et al. 1973: 178 (Lutzomyia sp. no. 260.43); CIPA group 1999 (tax., distr., nat. inf., ecol., refs.); Bejarano 2006a: 51 (Colombian Psychodidae checklist). sherlocki Martins, Silva & Falcão, 1971: 415. M., F. Type locality: Brazil, Mato Grosso, Vila Bela. Type material: HT M (CPqRR). Distr.: Colombia (Amazonas), Bolivia, Brazil, Ecuador (Young & Duncan 1994), Peru. Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 51 (Colombian Psychodidae checklist). sipani Fernández, Carbajal, Alexander & Need, 1994: 167. M. Type locality: Peru, Loreto, Maynas, San Antonio. Type material: HT M (MHNSM). Distr.: Colombia (Putumayo), Brazil (Figueira et al. 2013), Peru. Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 50 (Colombian Psychodidae checklist). sordellii (Shannon & Del Ponte), 1927: 730 (Phlebotomus). M., F. Type locality: Argentina, Chaco, Resistencia. Type material: LT M, F (USNM). Distr.: Colombia (Amazonas, Antioquia, Boyacá, Caldas (Alexander et al. 2001a), Caquetá, Casanare (Manotas-Solano et al. 2012), Cauca (Barreto et al. 2006), Chocó, Guaviare (Cabrera et al. 2009), Meta, Nariño, Norte de Santander, Putumayo, Santander (Sandoval et al. 1998), Tolima, Valle del Cauca), Argentina, Bolivia, Brazil, Costa Rica, Ecuador, French Guiana, Panama, Peru, Suriname (Kent et al. 2013), Trinidad and Tobago, Venezuela. Refs.: CIPA group 1999 (tax., distr., nat. inf., ecol., refs.); Bejarano 2006a: 53 (Colombian Psychodidae checklist). nordestinus (Mangabeira), 1942b: 327 (Flebotomus). longicornutus (Floch & Abonnenc), 1943: 6 (Phlebotomus). spinicrassa Morales, Osorno-Mesa, Osorno & Muñoz, 1969: 383. E., M., F. Type locality: Colombia, Boyacá, Almeida, Umbabita. Type locality: HT M, AT F (INS). Distr.: Colombia (Boyacá, Norte de Santander, Sucre), Venezuela. Refs.: CIPA group 1999 (tax., distr., nat. inf., ecol., refs.); Bejarano 2006a: 54 (Colombian Psychodidae checklist). strictivilla Young, 1979: 206. M., F. Type locality: Colombia, Antioquia, 24 km SW of Zaragoza, Río Anorí. Type locality: HT M, AT F (FSCA). Distr.: Colombia (Antioquia), Ecuador (Alexander et al. 1992), Venezuela (Feliciangeli 1988). Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 50 (Colombian Psychodidae checklist). suapiensis Le Pont, Torrez-Espejo & Dujardin, 1997: 56. F. Type locality: Bolivia, La Paz, Nor Yungas, Suapi. Type material: HT F, 2 PT F (MNHNB); 7 PT F (IRD). Distr.: Colombia (Tolima (Contreras et al. 2012)), Bolivia, Peru (Cáceres et al. 2001). thula Young, 1979: 180 (as Lutzomyia carrerai thula). E., L., M., F. Type locality: Panama, Panamá, Cerro Campana. Type material: HT M, AT F (FSCA). Distr.: Colombia (Antioquia, Caldas (Contreras-Gutiérrez et al. 2014), Cauca, Chocó, Nariño, Risaralda (Contreras-Gutiérrez et al. 2014), Valle del Cauca), Costa Rica, Ecuador, Honduras, Panama. Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 53 (Colombian Psychodidae checklist); Carvalho et al. 2006: 131 (raised to the category of species). tihuiliensis Le Pont, Torrez-Espejo & Dujardin, 1997: 56. F. Type locality: Bolivia, La Paz, Nor Yungas, Suapi. Type material: HT F, 2 PT F (MNHNB); 6 PT F (IRD). Distr.: Colombia (Antioquia), Bolivia, Peru (Cáceres & Galati 2001). Refs.: Bejarano 2006a: 54 (Colombian Psychodidae checklist). tolimensis Carrasquilla, Munstermann, Marín, Ocampo & Ferro, 2012: 994. M., F. Type locality: Colombia, Tolima, Chaparral, Agua Bonita. Type material: HT M, AT F, 7 PT (3 M and 4 F) (INS); 6 PT (3 M and 3 F) CATALOGUE OF PSYCHODIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 205 (PMNH, CIDEIM). Distr.: Colombia (Tolima). tortura Young & Rogers, 1984: 605. M., F. Type locality: Ecuador, Napo, Limoncocha. Type material: HT M, AT F (FSCA). Distr.: Colombia (Amazonas, Putumayo), Bolivia (Le Pont et al. 1992), Ecuador. Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 50 (Colombian Psychodidae checklist). torvida Young, Morales & Ferro in Young & Duncan, 1994: 180. M., F. Type locality: Colombia, Cundinamarca, Reventones. Type material: HT M, AT F (INS). Distr.: Colombia (Cundinamarca) Refs.: Ferro & Morales 1988: 69 (Lutzomyia sp. from Reventones); Kreutzer et al. 1990: 2 (Lutzomyia sp. A); CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 54 (Colombian Psychodidae checklist). townsendi (Ortíz), 1960: 23 (Phlebotomus). E., M., F. Type locality: Venezuela, Aragua, Rancho Grande. Type material: not extant (Feliciangeli et al. 1987). Distr.: Colombia (Valle del Cauca), Venezuela. Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 54 (Colombian Psychodidae checklist). trapidoi (Fairchild & Hertig), 1952: 524 (Phlebotomus). E., L., P., M., F. Type locality: Panama, Bocas del Toro, Almirante. Type material: HT M, AT F (FSCA). Distr.: Colombia (Antioquia, Boyacá, Caldas, Cauca (incl. Isla Gorgona (Barreto et al. 1997)), Chocó, Meta, Nariño, Norte de Santander, Risaralda, Santander, Tolima, Valle del Cauca), Costa Rica, Ecuador, Guatemala, Honduras, Nicaragua, Panama. Refs.: Rosabal 1954: 10 (Phlebotomus sp. no. 7); CIPA group 1999 (tax., distr., nat. inf., ecol., refs.); Bejarano 2006a: 52 (Colombian Psychodidae checklist). triacantha (Mangabeira), 1942a: 119 (Flebotomus). E., L., P., M., F. Type locality: Brazil, Pará, Belém, Aurá. Type material: HT M (IOC). Distr.: Colombia (Chocó, Meta, Nariño (Travi et al. 1988)), Brazil, Ecuador, French Guiana, Peru, Venezuela. Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 52 (Colombian Psychodidae checklist). trinidadensis (Newstead), 1922: 47 (Phlebotomus). E., L., P., M., F. Type locality: Trinidad. Type material: LT M, PLT F (BMNH). Distr.: Colombia (Antioquia, Atlántico (Maestre-Serrano et al. 2010), Bolívar (incl. Isla Fuerte (Gallego & Vélez 1994)), Boyacá, Caldas, Casanare (Manotas-Solano et al. 2012), Cesar, Chocó, Córdoba, Cundinamarca, Huila, La Guajira, Magdalena, Meta (Osorno-Mesa et al. 1972b), Nariño, Norte de Santander, Santander, Sucre, Tolima, Vichada (Vivero et al. 2010a)), Belize, Bolivia, Brazil, Costa Rica, Ecuador, French Guiana, Guatemala, Honduras, Mexico, Nicaragua, Panama, Peru, Suriname, Trinidad and Tobago, Venezuela. Refs.: CIPA group 1999 (tax., distr., nat. inf., ecol., refs.); Bejarano 2006a: 52 (Colombian Psychodidae checklist). yucatanensis (Galliard), 1934a: 1 (Phlebotomus). baduelensis (Floch & Abonnenc), 1941a: 4 (as Phlebotomus yucatanensis var. baduelensis). baduelensis (Floch & Abonnenc), 1944e: 1 (Phlebotomus). triramula (Fairchild & Hertig), 1952: 517 (Phlebotomus). L., P., M., F. Type locality: Panama, Colón, Río del Medio, near Río Gatún. Type material: HT M, AT F (FSCA). Distr.: Colombia (Antioquia, Caldas (Vergara et al. 2008b), Chocó, Nariño, Santander, Tolima, Valle del Cauca), Belize, Costa Rica, Ecuador, Guatemala, Mexico (Galati 2003a, Ibáñez-Bernal et al. 2011), Panama. Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 53 (Colombian Psychodidae checklist). tuberculata (Mangabeira), 1941c: 251 (Flebotomus). M., F. Type locality: Brazil, Pará, Belém, Aurá. Type material: HT M (IOC). Distr.: Colombia (Antioquia, Caquetá, Chocó, Santander (Sandoval et al. 2006), Guaviare (Cabrera et al. 2009), Meta, Nariño, Valle del Cauca), Bolivia (Le Pont et al. 1992), Brazil, Ecuador (Arrivillaga et al. 2014), French Guiana, Panama, Peru (Ogusuku et al. 1997), Suriname, Venezuela. Refs.: Floch & Abonnenc 1944b: 10 (Phlebotomus sp. x); CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 54 (Colombian Psychodidae checklist). munangai Wijers & Huisenga, 1967: 395. 206 · Zootaxa 4122 (1) © 2016 Magnolia Press BEJARANO & ESTRADA ubiquitalis (Mangabeira), 1942a: 158 (Flebotomus). M., F. Type locality: Brazil, Pará, Belém, Aurá. Type material: ST M (?IOC). Distr.: Colombia (Caquetá, Chocó, Meta), Bolivia (Le Pont et al. 1992), Brazil, Ecuador, French Guiana, Peru, Suriname, Venezuela. Refs.: CIPA group 1999 (tax., distr., nat. inf., ecol., refs.); Bejarano 2006a: 53 (Colombian Psychodidae checklist). basispinosus (Barretto & Coutinho), 1943: 185 (Phlebotomus). cauchensis (Floch & Abonnenc), 1943: 22 (Phlebotomus). umbratilis Ward & Fraiha, 1977: 314. L., M., F. Type locality: Brazil, Pará, Almeirim, Monte Dourado. Type material: HT F (IEC). Distr.: Colombia (Amazonas, ?Boyacá (Young 1979), Caquetá, Putumayo), Bolivia, Brazil, Ecuador (Williams 1999; Balbino et al. 2001), French Guiana, Peru, Suriname, Venezuela. Refs.: Ward et al. 1973: 178 (Lutzomyia sp. no. 260.31); CIPA group 1999 (tax., distr., nat. inf., ecol., refs.); Bejarano 2006a: 52 (Colombian Psychodidae checklist). undulata (Fairchild & Hertig), 1950: 527 (Phlebotomus). M., F. Type locality: Guatemala, Escuintla. Type material: HT M (FSCA). Distr.: Colombia (Boyacá, Chocó, Nariño, Santander), Belize, Bolivia, Costa Rica, Ecuador, El Salvador, Guatemala, Honduras, Mexico, Panama. Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 52 (Colombian Psychodidae checklist). humboldti (Vargas & Díaz-Nájera), 1959: 143 (Phlebotomus). vattierae Le Pont & Desjeux, 1992: 263. M., F. Type locality: Bolivia, Beni, Yucumo. Type material: HT F, AT M (IRD). Distr.: Colombia (Guaviare (Cabrera et al. 2009), Meta (Bejarano et al. 2006)), Bolivia, Peru (Ogusuku et al. 2001). Refs.: CIPA group 1999 (tax., distr., ecol., refs.). velezi Bejarano, Vivero & Uribe, 2010: 322. M. Type locality: Colombia, Antioquia, San Francisco, Reserva Natural Cañón del Río Claro “El Refugio”. Type material: HT M, 2 PT M (VHET). Distr.: Colombia (Antioquia). venezuelensis (Floch & Abonnenc), 1948: 1 (Phlebotomus). E., M., F. Type locality: Venezuela, Zulia, Selva de Tamanaco. Type material: HT M (current location unknown). Distr.: Colombia (Antioquia, Bolívar (Cortés 2006), Cundinamarca, Cesar (Estrada et al. 2015), La Guajira, Magdalena, Meta, Norte de Santander, Santander, Sucre, Vaupés), Venezuela. Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 51 (Colombian Psychodidae checklist). zuliaensis (Floch & Abonnenc), 1948: 5 (Phlebotomus). Material examined. COLOMBIA: Cesar: Mpio. [Municipio] Valledupar, Balneario Hurtado, 4 M., 1 F., 6.vii.2013, O.A. Aponte leg., CDC light trap (CZUS). vespertilionis (Fairchild & Hertig), 1947b: 617 (Phlebotomus). L., P., M., F. Type locality: Panama, Panamá, Cerro Campana. Type material: HT M, AT F (FSCA). Distr.: Colombia (Antioquia, Bolívar, Boyacá, Caldas, Chocó (Young 1979), Nariño, Santander, Tolima, Valle del Cauca), Costa Rica, Ecuador, Nicaragua (Zeledón & Murillo 1983), Panama. Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 50 (Colombian Psychodidae checklist). wagleyi (Causey & Damasceno), 1945a: 25 (Flebotomus). M., F. Type locality: Brazil, Amazonas, São Paulo de Olivença, Rio Solimões. Type material: HT M (USNM). Distr.: Colombia (Caquetá), Bolivia, Brazil, Venezuela. Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 53 (Colombian Psychodidae checklist). walkeri (Newstead), 1914: 188 (Phlebotomus). E., L., P., M., F. Type locality: Bolivia-Brazil boundary, Rio Abuná. Type material: LT M, PLT F (BMNH). Distr.: Colombia (Amazonas, Antioquia, Bolívar (Cortés 2006), Boyacá, Caldas (Contreras-Gutiérrez et al. 2014), Caquetá, Casanare, Cundinamarca, Guaviare (Morales et al. 1984), Huila, Meta, Norte de Santander, Putumayo, Santander, Tolima, Vaupés, Vichada (Vivero et al. 2010a)), Bolivia, Brazil, Ecuador, French Guiana, Panama, Paraguay, Peru, Trinidad and Tobago, Venezuela. Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 51 (Colombian Psychodidae checklist). marajoensis (Damasceno & Causey), 1944: 339 (Phlebotomus). CATALOGUE OF PSYCHODIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 207 gasti (Sherlock), 1962: 326 (Phlebotomus). witoto Young & Morales, 1987: 655. M. Type locality: Colombia, Amazonas, 8 km N of Leticia. Type material: HT M (INS). Distr.: Colombia (Amazonas, Guaviare (Cabrera et al. 2009), Putumayo), Ecuador (Jones et al. 2010). Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 53 (Colombian Psychodidae checklist). yencanensis (Ortíz), 1965b: 207 (Phlebotomus). M., F. Type locality: Venezuela, Aragua, Rancho Grande, Parque Nacional Henri Pittier. Type material: not designated. Distr.: Colombia (Norte de Santander, Santander (Sandoval et al. 2006)), Venezuela. Refs.: CIPA group 1999 (tax., distr., nat. inf., ecol., refs.); Bejarano 2006a: 51 (Colombian Psychodidae checklist). ylephiletor (Fairchild & Hertig), 1952: 518 (Phlebotomus). E., L., P., M., F. Type locality: Panama, Bocas del Toro, Almirante. Type material: HT M, AT F (FSCA). Distr.: Colombia (Bolívar (Cortés 2006), Chocó, Nariño, Valle del Cauca), Belize, Costa Rica, Ecuador (Arzube 1960), Guatemala, Honduras, Mexico, Nicaragua, Panama. Refs.: Rosabal 1954: 10 (Phlebotomus sp. no. 14); CIPA group 1999 (tax., distr., nat. inf., ecol., refs.); Bejarano 2006a: 52 (Colombian Psychodidae checklist). youngi Feliciangeli & Murillo, 1987: 141. E., L., M., F. Type locality: Venezuela, Trujillo, Las Calderas. Type material: HT M, AT F (CNRFV). Distr.: Colombia (Antioquia, Caldas, Valle del Cauca), Costa Rica, Venezuela. Refs.: CIPA group 1999 (tax., distr., nat. inf., ecol., refs.); Bejarano 2006a: 54 (Colombian Psychodidae checklist). yucumensis (Le Pont, Caillard, Tibayrenc & Desjeux), 1986: 79 (Psychodopygus). M., F. Type locality: Bolivia, Beni, Serranía del Pilón, km 4 road Yucumo-Rurrénabaque. Type material: HT M, AT F (IRD). Distr.: Colombia (Putumayo), Bolivia, Brazil, Peru. Refs.: Biancardi et al. 1982: 177 (Psychodopygus sp. no. 2); CIPA group 1999 (tax., distr., nat. inf., ecol., refs.); Bejarano 2006a: 53 (Colombian Psychodidae checklist). yuilli Young & Porter, 1972: 524. E., M., F. Type locality: Colombia, Antioquia, 24 km SW of Zaragoza, Rio Anorí. Type material: HT M, AT F (USNM). Distr.: Colombia (Amazonas, Antioquia, Bolívar (Cortés 2006), Boyacá (Santamaría et al. 2006), Caldas (Alexander et al. 2001a), Caquetá, Chocó, Guainía (Flórez & Ferro 2007), Guaviare, Meta, Putumayo, Santander, Vichada (Vivero et al. 2010a)), Bolivia, Brazil, Ecuador, Peru, Venezuela. Refs.: CIPA group 1999 (tax., distr., nat. inf., ecol., refs.); Bejarano 2006a: 52 (Colombian Psychodidae checklist). fraihai Martins, Falcão & Silva, 1979: 357. Genus Warileya Hertig Warileya Hertig, 1948: 12. Type species, Warileya phlebotomanica Hertig, 1948 (orig. des.). Hertigia Fairchild, 1949: 81. Type species, Hertigia hertigi Fairchild, 1949 (orig. des.). Comm.: Unlike Lutzomyia and Brumptomyia that present a wide diversity and distribution in the Neotropics, Warileya includes only nine described species, distributed in Costa Rica, Panama, Colombia, Peru, Ecuador, Bolivia and French Guiana (CIPA group 1999). The genus, which is associated principally with forest habitats, is represented in Colombia by the following two species, the last of which has been observed biting man in this country (Young 1979). nigrosaccula Fairchild & Hertig, 1951b: 428. M., F. Type locality: Panama, Panamá, Cerro Campana. Type material: HT F (FSCA). Distr.: Colombia (Chocó, Valle del Cauca), Panama. Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 55 (Colombian Psychodidae checklist). rotundipennis Fairchild & Hertig, 1951b: 424. L., M., F. Type locality: Panama, Panamá, Cerro Campana. Type material: HT M, AT F (FSCA). Distr.: Colombia (Antioquia, Chocó, Norte de Santander, Risaralda, Tolima (Ferro et al. 2011), Valle del Cauca), Bolivia, Costa Rica, Panama, Peru (Ogusuku et al. 1997). Refs.: CIPA group 1999 (tax., distr., ecol., refs.); Bejarano 2006a: 55 (Colombian Psychodidae checklist). 208 · Zootaxa 4122 (1) © 2016 Magnolia Press BEJARANO & ESTRADA Subfamily Psychodinae Rondani, 1856 Genus Arisemus Satchell Telmatoscopus, subg. Arisemus Satchell, 1955: 88. Type species, Psychoda grabhamana Dyar, 1926 (by synonymy with originally designated type species, Telmatoscopus maculosus Satchell, 1955). Bazara Vaillant, 1986: 337. Type species, Arisemus tetradactylus Botosaneanu & Vaillant, 1970 (orig. des.). Comm.: Arisemus is a genus that encompasses 31 species distributed between Guatemala and Brazil, including several Caribbean islands (Quate & Brown 2004, Bravo & Araújo 2013b). A revision of the genus and key for the identification of species is found in Quate & Brown (2004). Three Arisemus species have been recorded from Colombia. guhli Wagner & Joost, 1994: 76. M. Type locality: Colombia, Huila, San Agustín, Río Magdalena. Type material: HT M, 7 PT (2 M and 5 F) (ULMG). Distr.: Colombia (Huila). Refs.: Quate & Brown 2004: 74 (rev., key); Bejarano 2006a: 48 (Colombian Psychodidae checklist). martinezi Wagner & Joost, 1994: 79. M. Type locality: Colombia, Bogotá, Quebrada Chicó. Type material: HT M, 2 PT (M and F) (ULMG); 2 PT (M and F) (RW). Distr.: Colombia (Bogotá). Refs.: Quate & Brown 2004: 78 (rev., key); Bejarano 2006a: 48 (Colombian Psychodidae checklist). obandoi Wagner & Joost, 1994: 79. M. Type locality: Colombia, Huila, San Agustín, Río Magdalena. Type material: HT M (ULMG). Distr.: Colombia (Huila). Refs.: Quate & Brown 2004: 79 (rev., key); Bejarano 2006a: 48 (Colombian Psychodidae checklist). Genus Australopericoma Vaillant Australopericoma Vaillant, 1975: 172. Type species, Pericoma caudata Satchell, 1955 (by synonymy with the type species, Pericoma wirthi Quate, 1955). Comm.: With 15 species described to date, this New World genus is found distributed from the United States of America to Argentina (Quate & Brown 2004, Bravo 2007a), and is represented by a single species in Colombia. Quate & Brown (2004) revised the genus and presented a key for the identification of species. roessleri (Wagner & Joost), 1994: 77 (Arisemus). M. Type locality: Colombia, Huila, San Agustín, Río Magdalena. Type material: HT M, 4 PT (3 M and 1 F) (ULMG). Distr.: Colombia (Huila). Refs.: Quate & Brown 2004: 89 (rev., key); Bejarano 2006a: 48 (Colombian Psychodidae checklist). Genus Balbagathis Quate Balbagathis Quate, 1996: 31. Type species, Balbagathis sylvatica Quate, 1996 (orig. des.). Comm.: This Neotropical genus comprises 11 described species, which are distributed from Mexico to Brazil (Bravo 2004a, Ibáñez-Bernal 2008). Originally proposed by Quate (1996), the genus was redefined by Quate & Brown (2004), who included a key for the identification of species. The taxon was recorded in Colombia by Collantes et al. (2000) without specifying the species or locality. Undetermined sp. Distr.: Colombia (Collantes et al. 2000). Refs.: Bejarano 2006a: 48 (Colombian Psychodidae checklist). Genus Clogmia Enderlein Clogmia Enderlein, 1937: 87. Type species, Psychoda albipunctata Williston, 1893 (orig. des., as Psychoda albipennis Williston, 1893, lapsus). Lepiseodina Enderlein, 1937: 93. Type species, Psychoda tristis Meigen, 1830 (orig. des.). Comm.: Clogmia has been considered a junior synonym of the genus Telmatoscopus Eaton, 1904 (Vaillant 1972), although CATALOGUE OF PSYCHODIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 209 most taxonomists currently accept its generic status. It is represented in Colombia by the following cosmopolitan species, widely adapted to anthropogenic habitats and has been identified in other countries as an agent of human myiasis (Mohammed & Smith 1976, Kamimura & Arakawa 1986, Hyun et al. 2004, Tu et al. 2007, El-Badry et al. 2014) and as a potential mechanical vector of bacterial pathogens (Faulde & Spiesberger 2013). albipunctata (Williston), 1893: 113 (Psychoda). E., L., P., M., F. Type locality: Cuba, La Havana. Type material: LT M (USNM). Distr.: Cosmopolitan. Refs.: Ibáñez-Bernal 2008: 94 (refs. to that date, Mexico records); Boumans 2009: 1 (Netherlands records); Boumans et al. 2009: 153 (biol., rev., Belgium records); Gelardi et al. 2009: 780 (causative agent of nasal obstruction); García-Solache et al. 2010: 306 (mol. biol.); Boumans 2011: 62 (Netherlands records); Rocha et al. 2011: 179 (egg morphology); Curler & Moulton 2012: 606 (mol. tax.); Espíndola et al. 2012: 494 (mol. tax.); Faulde & Spiesberger 2012: 134 (hospital infestations); Ježek et al. 2012: 122 (Czech and Slovakia records); Kvifte 2012: 34 (Afrotropical Psychodidae cat.); Oboňa & Ježek 2012a: 280 (biol.), 2012b: 388 (range expansion in Slovakia); Faulde & Spiesberger 2013: 51 (potential vector); Janssens et al. 2013: 1 (mol. biol.); Kvifte et al. 2013: 64 (Croatia records); Wagner 2013 (European Psychodidae checklist); Wagner & Weber 2013: 291 (Luxembourg record); El-Badry et al. 2014: 247 (causative agent of human urinary myiasis); Jiménez-Guri et al. 2014: 1 (dev. biol., description of the life cycle). Note: A recompilation of references on Clogmia albipunctata (Williston, 1893) can be found in Ibáñez-Bernal (2008) and Boumans et al. (2009), thus only studies published subsequently to these are cited above. meridionalis (Eaton), 1894: 194 (Pericoma). snowii (Haseman), 1907: 311 (Psychoda). albonigra (Brunetti), 1908: 374 (Psychoda). legnothisa (Speiser), 1909: 44 (Psychoda). erecta (Curran), 1926: 102 (Psychoda). nocturna (Santos Abréu), 1930: 115 (Psychoda). nigrithorax (Santos Abréu), 1930: 115 (as Psychoda nocturna var. nigrithorax). guianica (Curran), 1934: 309 (Psychoda). giesi (Vargas), 1953: 130 (Syntomolaba). haranti (Mirouse), 1958: 93 (Telmatoscopus). Genus Didicrum Enderlein Didicrum Enderlein, 1937: 105. Type species, Pericoma griseata Tonnoir, 1929 (orig. des.). Mecysmia Enderlein 1937: 108. Type species, Pericoma schoenemanni Enderlein, 1937 (orig. des.). Podolepria Enderlein, 1937: 105. Type species, Pericoma inornata Tonnoir, 1929 (orig. des.). Comm.: This Gondwanan genus is comprised of 23 species (Moya-Arévalo et al. 2012, Omad 2014), 12 of which are present in South America, including the only species listed here, which was recently described from Colombia. colombensis Moya-Arévalo, Ibáñez-Bernal & Suárez-Landa, 2012: 81. M., F. Type locality: Colombia, Cundinamarca, Parque Nacional Natural Chingaza, Bosque de Palacio. Type material: HT M, 3 PT F (ICNMHN); 3 PT (1 M and 3 F) (LACM); 6 PT (2 M and 4 F) (IEXA). Distr.: Colombia (Boyacá, Cundinamarca). Genus Feuerborniella Vaillant Feuerborniella Vaillant, 1971: 37. Type species, Psychoda obscura Tonnoir, 1919 (mon.). Comm.: Feuerborniella constitutes a small genus with a worldwide distribution which includes seven species (Ibáñez-Bernal 2004a, Cordeiro et al. 2014). Four of these have a Neotropical origin, with locality records from Mexico, Colombia and Brazil. paramuna Cordeiro in Cordeiro et al., 2014: 378. M. Type locality: Colombia, Antioquia, Sonsón, Páramos, Cerro de Las Cruces. Type material: HT M, 2 PT (M and F) (CEUA); 2 PT (M and F) (DZUP). Distr.: Colombia (Antioquia). 210 · Zootaxa 4122 (1) © 2016 Magnolia Press BEJARANO & ESTRADA Genus Lepidiella Enderlein Lepidiella Enderlein, 1937: 89. Type species, Lepidiella lanuginosa Enderlein, 1937 (orig. des.). Syntomoza Enderlein, 1937: 88. Type species, Syntomoza niveitarsis Enderlein, 1937 (orig. des.). Kupara Rapp, 1945: 310. Type species, Kupara albipeda Rapp, 1945 (mon.). Comm.: This is a neotropical genus containing 14 species, distributed from Mexico to Brazil, including the Caribbean island of Saint Lucia (Ibáñez-Bernal 2008, 2010, Bravo & Araújo 2013b). In Colombia it is represented by a single species originally described from this country. larryi Ibáñez-Bernal, 2010: 35. M., F. Type locality: Colombia, Magdalena, Parque Nacional Natural Sierra Nevada de Santa Marta, San Lorenzo. Type material: HT M, 3 PT (2 M and 1 F) (ICNMHN); 3 PT M (LACM); 3 PT M (IEXA). Distr.: Colombia (Magdalena). Genus Maruina Müller Maruina Müller, 1895: 480. Type species, Maruina pilosella Müller, 1895 (sub. des., Enderlein 1937: 110). Comm.: This New World genus includes 37 species and has a wide geographic range extending from Canada to Argentina, including the Antilles (Dowes 1972, Hogue 1973, 1990, Bravo 2005). A taxonomic revision of the genus was performed by Hogue (1973), in which he recognized two subgenera and presented keys for the morphological identification of adults and immatures. Four of the 33 species described from the Neotropics have been found in Colombia. Subgenus Aculcina Hogue Aculcina Hogue, 1973: 13. Type species, Maruina cholita Hogue, 1973 (orig. des.). chaborra Hogue, 1973: 24. L., P., F. Type locality: Colombia, Boyacá, NW of Arcabuco, Valle del Río Pomeca. Type material: HT F, 2 PT F (AMNH); PT F (LACM). Distr.: Colombia (Boyacá). Refs.: Bravo 2004c: 5 (tax.); Bejarano 2006a: 48 (Colombian Psychodidae checklist). chiringa Hogue, 1990: 186. L., P., M., F. Type locality: Colombia, Cundinamarca, Quebrada El Puerto. Type material: HT M, AT F, 14 PT (LACM); 2 PT (M and F) (ICNMHN); 2 PT (M and F) (USNM). Distr.: Colombia (Cundinamarca). Refs.: Bravo 2004c: 5 (tax.); Bejarano 2006a: 48 (Colombian Psychodidae checklist). colombicana Wagner & Joost, 1994: 83. M. Type locality: Colombia, Bogotá. Type material: HT M (ULMG); PT M (RW). Distr.: Colombia (Bogotá). Refs.: Bravo 2004c: 5 (tax.); Bejarano 2006a: 48 (Colombian Psychodidae checklist). Subgenus Maruina Hogue Maruina Hogue, 1973: 27. Type species, Maruina pilosella Müller, 1895 (orig. des.). campesina Hogue, 1973: 55. L., P., M. Type locality: Colombia, Cundinamarca, road Sibaté-La Aguadita. Type material: HT M (AMNH); 3 PT M (AMNH, LACM). Distr.: Colombia (Cundinamarca). Refs.: Bravo 2004c: 5 (tax.); Bejarano 2006a: 48 (Colombian Psychodidae checklist). Genus Paramormia Enderlein Paramormia Enderlein, 1935: 248. Type species, Pericoma fratercula Eaton, 1893 (orig. des.). Duckhousiella Vaillant, 1972: 54. Type species, Pericoma ustulata Walker, 1856 (orig. des.) Comm.: This genus, represented in the Neotropics by a single species, Paramormia fucarta (Kincaid, 1899), was recorded in Colombia by Wagner & Joost (1994) although without specifying the species. CATALOGUE OF PSYCHODIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 211 Undetermined sp. Distr.: Colombia (Boyacá, Cundinamarca (Wagner & Joost 1994)). Refs.: Collantes et al. 2000 (Colombia record); Bejarano 2006a: 49 (Colombian Psychodidae checklist). Genus Parasetomima Duckhouse Parasetomima Duckhouse, 1968: 29. Type species, Parasetomima umbella Duckhouse, 1968 (orig. des.). Comm.: This small genus, composed of two species endemic to Brazil, Parasetomima umbella Duckhouse, 1968 and Pa. ornata Bravo, 2004 (Duckhouse 1968, Bravo 2004b), was recorded in Colombia by Collantes et al. (2000) without specifying the species or locality. Undetermined sp. Distr.: Colombia (Collantes et al. 2000). Refs.: Bejarano 2006a: 48 (Colombian Psychodidae checklist). Genus Psychoda Latreille Psychoda Latreille, 1796: 152. Type species, Tipula phalaenoides Linnaeus, 1758 (mon.). Tinearia Schellenberg, 1803: pl. 40. Type species, Psychoda alternata Say, 1824 (sub. des., Coquillett, 1910: 615). Trichoptera Meigen, 1803: 261. Type species, Tipula phalaenoides Linnaeus, 1758 (sub. des., Coquillett, 1910: 616). Logima Eaton, 1904: 58. Type species, Psychoda erminea Eaton, 1904 (orig. des.). Copropsychoda Vaillant, 1971: 30. Type species, Psychoda brevicornis Tonnoir, 1940 (mon.). Psychodocha Ježek, 1984: 135. Type species, Psychoda cinerea Banks, 1894 (orig. des.). Psycha Ježek, 1984: 136. Type species, Psychoda grisescens Tonnoir, 1922 (orig. des.). Psychomora Ježek, 1984: 137. Type species, Psychoda trinodulosa Tonnoir, 1922 (orig. des.). Chodopsycha Ježek, 1984: 138. Type species, Psychoda lobata Tonnoir, 1940 (orig. des.). Psychodula Ježek, 1984: 138. Type species, Psychoda minuta Banks, 1894 (orig. des.). Ypsydocha Ježek, 1984: 140. Type species, Psychoda setigera Tonnoir, 1922 (orig. des.). Falsologima Ježek & van Harten, 1996: 73. Type species, Psychoda savaiiensis Edwards, 1928 (orig. des.). Psychana Ježek & van Harten, 2005: 207. Type species, Psychoda rujumensis Ježek & van Harten, 2005 (orig. des.). Apsycha Ježek, 2007: 146. Type species, Psychoda pusilla Tonnoir, 1922 (orig. des.). Comm.: With a wide global distribution, this constitutes one of the most species-rich genera of non- haematophagous psychodids with about 450 described species (Salmela et al. 2012), some of which have become adapted to synanthropic habitats, e.g. Psychoda alternata Say, 1824, Ps. albipennis Zetterstedt, 1850 and Ps. cinerea Banks, 1894, and associated with cases of human myiasis, i.e. Ps. alternata and Ps. albipennis (Kamimura 1967, Oğuz et al. 2012, Yones et al. 2014). The genus has been subjected to different revisions (Quate 1959, Vaillant 1972, Ježek 1984, Bravo et al. 2006), which range from the recognition of a single genus Psychoda s.l. made up of 12 subgenera (Bravo et al. 2006), to a división of the taxon in 13 genera (Ježek 1984, 2007, Ježek & van Harten 2005). Seven of the 75 species found in the Neotropics (Bravo et al. 2006, 2010a, Cordeiro & Bravo 2008, Ibáñez-Bernal 2008, Cordeiro et al. 2011, Ježek et al. 2011) have been recorded from Colombia and are listed here, without consideration of their subgeneric classification. aitkeni Quate, 1959: 218. M., F. Type locality: Trinidad and Tobago, Trinidad, Saint George, Chaguaramas. Type material: HT F (USNM). Distr.: Colombia (Collantes et al. 2000), Nicaragua (Collantes & Martínez-Ortega 1999), Panama (Duckhouse 1973), Trinidad. Refs.: Bejarano 2006a: 48 (Colombian Psychodidae checklist); Bravo et al. 2006: 11 (Latin American and Caribbean checklist). alternata Say, 1824: 358. E., L., P., M., F. Type locality: USA, Pennsylvania, Philadelphia. Type material: lost (Berdén 1952). Distr.: Cosmopolitan. Refs.: Ibáñez-Bernal 2008: 97 (refs. to that date, Mexico records); Mazurkiewicz & Kubrakiewicz 2008: 267 (dev. biol.); Boumans 2011: 58 (Netherlands records); Cordeiro et al. 2011: 6 (Brazil records, key); Ježek et al. 2011: 192 (Tinearia species checklist); Kvifte et al. 2011: 185 (Norway record); Curler & Moulton 2012: 606 (mol. tax.); Espíndola et al. 2012: 494 (mol. tax.); Kvifte 2012: 42 (Afrotropical Psychodidae cat.); Omad 2012: 262 (Argentina records); Ali et al. 2013: 11 (Iraq records); Kvifte et al. 2013: 64 (Croatian checklist); Wagner 2013 (European Psychodidae checklist); Elgueta & Ježek 2014: 79 (Chilean Psychodidae checklist); Kvifte & Boumans 2014: 12 (Norway records and DNA barcoding); Salmela et al. 2014: 40 (Finnish Diptera checklist). Note: Only references subsequent to Ibáñez-Bernal (2008) are cited above. 212 · Zootaxa 4122 (1) © 2016 Magnolia Press BEJARANO & ESTRADA tripunctata Macquart, 1838a: 81; 1838b: 85. marginepunctata von Roser, 1840: 50. septempunctata Phillipi, 1865: 631. conspicillata Hutton, 1881: 13. schizura Kincaid, 1899: 32. floridica Haseman, 1907: 316. nocturnala Haseman, 1907: 319. bengalensis Brunetti, 1908: 371. albimaculata Welch, 1912: 411. dakotensis Dyar, 1926: 108. marmorosa Santos Abréu, 1930: 123 (as Psychoda alternata var. marmorosa). floridica Johannsen, 1934: 25 (as Psychoda alternata var. floridica Haseman, 1907). alternicula Quate, 1955: 222. M., F. Type locality: USA, Florida, W. Palm Beach, Morrison Field. Type material. HT F, AT M, others PT (M and F) (USNM). Distr.: Colombia (Collantes et al. 2000), Brazil (Cordeiro et al. 2011), Israel (Wagner 1984), Mexico, Nicaragua (Collantes & Martínez-Ortega 1999), Panama, Trinidad, USA (incl. Florida Keys Islands (Wagner & Hribar 2005)) (Duckhouse 1973). Refs.: Ibáñez-Bernal 2008: 99 (refs. to that date, Mexico records); Cordeiro et al. 2011: 6 (in key to Brazilian species); Ježek et al. 2011: 192 (Tinearia species checklist). bogotensis Wagner & Joost, 1994: 82. M. Type locality: Colombia, Bogotá, Quebrada Chicó. Type material: HT M (ULMG). Distr.: Colombia (Bogotá). Refs.: Bejarano 2006a: 49 (Colombian Psychodidae checklist); Bravo et al. 2006: 12 (Latin American and Caribbean checklist). cordiforma Quate, 1996: 70. F. Type locality: Costa Rica, Guanacaste, La Pacífica. Type material: HT F (INBIO); 3 PT F (LACM, USNM). Distr.: Colombia (Collantes et al. 2000); Costa Rica, Nicaragua (Collantes & MartínezOrtega 1999). Refs.: Bejarano 2006a: 49 (Colombian Psychodidae checklist); Bravo et al. 2006: 12 (Latin American and Caribbean checklist). mimica Quate, 1996: 73. F. Type locality: Costa Rica, Limón, Puerto Viejo de Talamanca. Type material: HT F (INBIO); 8 PT F (BMNH, LACM, USNM). Colombia (Collantes et al. 2000); Costa Rica, Nicaragua (Collantes & Martínez-Ortega 1999). Refs.: Bejarano 2006a: 49 (Colombian Psychodidae checklist); Bravo et al. 2006: 12 (Latin American and Caribbean checklist). savaiiensis Edwards, 1928: 74. M., F. Type locality: Samoa, Savaii, Salailua. Type material: HT M (BMNH). Distr.: Pantropical. Refs.: Ibáñez-Bernal 2008: 104 (full refs. to that date, Mexico records); Cordeiro et al. 2011: 14 (Brazil records, key); Kvifte 2012: 40 (Afrotropical Psychodidae cat.); Sinclair 2013: 37 (Galapagos Islands checklist). rarotongensis Satchell, 1953: 183. lucia Quate, 1954: 349. Subfamily Sycoracinae Jung, 1954 Genus Sycorax Haliday Sycorax Haliday in Curtis, 1839: 745. Type species, Sycorax silacea Haliday in Curtis, 1839 (mon.). Microdixa Müller, 1927: 535. Type species, Microdixa scutigera Müller, 1927 (orig. des.). Comm.: One fossil and 18 extant species of this genus (Petrulevičius et al. 2011, Santos et al. 2011, Curler & Jacobson 2012, Santos et al. 2013a) have been recorded from the Neotropics, one of which, Sycorax wampukrum Bravo & SalazarValenzuela, 2009, feeds on the blood of harlequin frogs (Bravo & Salazar-Valenzuela 2009). Two taxonomic proposals have been put forward for these species, that of three genera proposed by Duckhouse (1972) and the monogeneric of Ježek (1999); we prefer to follow the latter in the absence of molecular phylogenetic studies which would permit its generic CATALOGUE OF PSYCHODIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 213 status to be resolved. Keys for the species of the Neotropical region are found in Bravo et al. (2010b) and Santos et al. (2013a). Five Sycorax species have been described to date in Colombia. andicola Young, 1979: 21. M. Type locality: Colombia, Cauca, Cerro Munchique. Type material: HT M (FSCA); PT M (BMNH, FSCA or INS). Distr.: Colombia (Cauca). Refs.: Ježek 1999: 84 (world checklist); Bejarano 2006a: 49 (Colombian Psychodidae checklist); Azar et al. 2007: 10 (world checklist); Bravo 2007b: 527 (in key to Neotropical species); Bravo et al. 2010b: 836 (in key to Neotropical species); Curler & Jacobson 2012: 61 (world checklist). colombiensis Young, 1979: 23. M., F. Type locality: Colombia, Valle del Cauca, Anchicayá. Type material: HT M, AT F (FSCA); 6 PT (1 M and 5 F) (BMNH, FSCA, INS). Distr.: Colombia (Valle del Cauca). Refs.: Ježek 1999: 84 (world checklist); Bejarano 2006a: 49 (Colombian Psychodidae checklist); Azar et al. 2007: 10 (world checklist); Bravo 2007b: 527 (in key to Neotropical species); Bravo et al. 2010b: 836 (in key to Neotropical species); Curler & Jacobson 2012: 61 (world checklist). fairchildi Young, 1979: 23. M., F. Type locality: Colombia, Valle del Cauca, Anchicayá. Type material: HT M, AT F (FSCA); 8 PT (6 M and 2 F) (BMNH, FSCA, INS). Distr.: Colombia (Antioquia, Valle del Cauca). Refs.: Ježek 1999: 84 (world checklist); Bejarano 2006a: 49 (Colombian Psychodidae checklist); Azar et al. 2007: 10 (world checklist); Bravo 2007b: 527 (in key to Neotropical species); Bravo et al. 2010b: 836 (in key to Neotropical species); Curler & Jacobson 2012: 61 (world checklist). trispinosa Young, 1979: 25. M., F. Type locality: Colombia, Valle del Cauca, Anchicayá. Type material: HT M, AT F (FSCA); 24 PT (15 M and 9 F) (BMNH, FSCA, INS). Distr.: Colombia (Valle del Cauca). Refs.: Ježek 1999: 84 (world checklist); Bejarano 2006a: 49 (Colombian Psychodidae checklist); Azar et al. 2007: 10 (world checklist); Bravo 2007b: 527 (in key to Neotropical species); Bravo et al. 2010b: 836 (in key to Neotropical species); Curler & Jacobson 2012: 62 (world checklist). utriensis Bejarano, Duque & Vélez, 2008: 29. M. Type locality: Colombia, Chocó, Bahía Solano, Parque Nacional Natural Ensenada de Utría. Type material: HT M (VHET). Distr.: Colombia (Chocó). Refs.: Curler & Jacobson 2012: 61 (world checklist); Santos et al. 2009b: 64 (tax., key). Subfamily Trichomyiinae Tonnoir, 1922 Genus Trichomyia Haliday Trichomyia Haliday in Curtis, 1839: 745. Type species, Trichomyia urbica Haliday in Curtis, 1839 (mon.). Diplonema Loew, 1845: 7. Type species, Diplonema buceras Loew, 1845 (mon.). Phalaenomyia Loew, 1845: 9. Type species, Diplonema buceras Loew, 1845 (sub. des., Evenhuis, 1994: 195). Termitodipteron Holmgren, 1905: 533. Type species, Termitodipteron wasmanni Holmgren, 1905 (mon.). Lepria Enderlein, 1937: 112. Type species, Lepria squamosa Enderlein, 1937 (orig. des.). Eubonetia Vargas & Díaz-Nájera, 1953: 155. Type species, Trichomyia cirrata Coquillett, 1902 (orig. des.). Comm.: This constitutes one of the most diverse genera of the fauna of Neotropical non-haematophagous psychodids, with 74 extant species (Araújo & Bravo 2013) and 8 fossil ones described (Quate 1961, 1963, Stebner & Solórzano-Kraemer 2014). Although four new subgenera have been proposed from the Neotropics (Bravo 1999, 2001, Araújo & Bravo 2013, Bravo & Araújo 2013a), most of the species described remain ungrouped, including two of the five species recorded in Colombia. Subgenus Brachiotrichomyia Bravo & Araújo Brachiotrichomyia Bravo & Araújo, 2013a: 330. Type species, Trichomyia quatei Bravo, 2001 (orig. des.). risaraldensis Bejarano, Pérez-Doria & Sierra, 2009a: 296. M. Type locality: Colombia, Rirasalda, Marsella, Jardín 214 · Zootaxa 4122 (1) © 2016 Magnolia Press BEJARANO & ESTRADA Botánico “Alejandro Humboldt”. Type material: HT M, 3 PT M (VHET). Distr.: Colombia (Risaralda). Refs.: Santos & Leite 2012: 44 (western hemisphere checklist); Bravo & Araújo 2013a: 334 (tax., key). Subgenus Opisthotrichomyia Bravo Opisthotrichomyia Bravo, 2001: 50. Type species, Trichomyia brevitarsa (Rapp, 1945) (orig. des.). andina Bejarano, Pérez-Doria & Sierra, 2010: 76. M. Type locality: Colombia, Risaralda, Marsella. Type material: HT M, 7 PT M (VHET). Distr.: Colombia (Risaralda). Refs.: Santos & Leite 2012: 42 (western hemisphere checklist). brevitarsa (Rapp), 1945: 310 (Psychoda). M., F. Type locality: Panama, Canal Zone, Barro Colorado Island. Type material: HT M (AMNH). Distr.: Colombia (Bolívar (incl. Isla Fuerte (Bejarano 2006b)), Córdoba (Bejarano 2006a), Sucre (Bejarano 2006b)), Costa Rica (Quate 1996), Mexico (Ibáñez-Bernal 2004b), Panama, Peru, Surinam, Venezuela (Alexander et al. 2001b). Refs.: Duckhouse 1974: 61 (redescription, new combination); Quate 1996: 8 (tax., key); Quate 1999: 416 (tax.); Bravo 2001: 50 (as subgenotype, key); Ibáñez-Bernal 2004b: 3 (female description); Bejarano 2006a: 49 (Colombian Psychodidae checklist); Santos & Leite 2012: 42 (western hemisphere checklist). maldonadoi (Vargas), 1953: 128 (Eubonetia). Species unplaced to subgenus colosensis Pérez-Doria, Hernández & Bejarano, 2010: 39. M. Type locality: Colombia, Sucre, Colosó, Reserva Forestal Protectora Serranía de Coraza y Montes de María. Type material: HT M, 2 PT M (CZUS). Distr.: Colombia (Sucre). Refs.: Santos & Leite 2012: 42 (western hemisphere checklist). quimbaya Bejarano, Pérez-Doria & Sierra, 2009b: 98. M. Type locality: Colombia, Rirasalda, Marsella, Jardín Botánico “Alejandro Humboldt”. Type material: HT M (VHET). Distr.: Colombia (Risaralda). Refs.: Santos & Leite 2012: 43 (western hemisphere checklist). TABLE 1. Psychodidae species recorded from Colombia. Subfamily Genus Species Author Bruchomyiinae Nemopalpus cancer Wagner & Stuckenberg, 2012 mopani De León, 1950 patriciae Alexander, 1987 phoenimimos Quate & Alexander, 2000 avellari (Costa Lima, 1932) beaupertuyi (Ortíz, 1954) galindoi (Fairchild & Hertig, 1947) hamata (Fairchild & Hertig, 1947) leopoldoi (Rodríguez, 1953) mesai Sherlock, 1962 pentacantha (Barretto, 1947) pintoi (Costa Lima, 1932) abonnenci (Floch & Chassignet, 1947) abunaensis Martins, Falcão & Silva, 1965 aclydifera (Fairchild & Hertig, 1952) Phlebotominae Brumptomyia Lutzomyia ...Continued on next page CATALOGUE OF PSYCHODIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 215 TABLE 1. (Continued) Subfamily Genus Species Author amazonensis (Root, 1934) andina Osorno, Osorno-Mesa & Morales, 1972 antioquiensis (Wolff & Galati, 2002) antunesi (Coutinho, 1939) aragaoi (Costa Lima, 1932) araracuarensis Morales & Minter, 1981 atroclavata (Knab, 1913) auraensis (Mangabeira, 1942) ayrozai (Barretto & Coutinho, 1940) barrettoi barrettoi (Mangabeira, 1942) barrettoi majuscula Young, 1979 begonae (Ortíz & Torrez, 1975) bernalei (Osorno-Mesa, Morales & Osorno, 1967) bettinii Feliciangeli, Ramírez-Pérez & Ramírez, 1988 bicolor Fairchild & Theodor, 1971 bifoliata Osorno-Mesa, Morales, Osorno & Muñoz, 1970 bispinosa (Fairchild & Hertig, 1951) bursiformis (Floch & Abonnenc, 1944) campbelli (Damasceno, Causey & Arouck, 1945) camposi (Rodríguez, 1950) caprina Osorno-Mesa, Morales & Osorno, 1972 carpenteri (Fairchild & Hertig, 1953) carrerai (Barretto, 1946) cayennensis cayennensis (Floch & Abonnenc, 1941) ceferinoi (Ortíz & Álvarez, 1963) cellulana Young, 1979 cerqueirai (Causey & Damasceno, 1945) chagasi (Costa Lima, 1941) chassigneti (Floch & Abonnenc, 1944) choti (Floch & Abonnenc, 1941) christenseni Young & Duncan, 1994 cirrita Young & Porter, 1974 claustrei Abonnenc, Léger & Fauran, 1979 columbiana (Ristorcelli & Van Ty, 1941) conviti Ramírez-Pérez, Martins & Ramírez, 1976 cuzquena Martins, Llanos & Silva, 1975 damascenoi (Mangabeira, 1941) dasymera (Fairchild & Hertig, 1961) davisi (Root, 1934) dendrophyla (Mangabeira, 1942) disiuncta Morales, Osorno & Osorno-Mesa, 1974 ...Continued on next page 216 · Zootaxa 4122 (1) © 2016 Magnolia Press BEJARANO & ESTRADA TABLE 1. (Continued) Subfamily Genus Species Author dreisbachi (Causey & Damasceno, 1945) dubitans (Sherlock, 1962) duncanae Le Pont, Martínez, Torrez-Espejo & Durjardin, 1998 dysponeta (Fairchild & Hertig, 1952) emberai Bejarano, Duque & Vélez, 2004 erwindonaldoi (Ortíz, 1978) evangelistai Martins & Frahia, 1971 evansi (Núñez-Tovar, 1924) fairtigi Martins, 1970 falcata Young, Morales & Ferro, 1994 ferroae Young & Morales, 1987 flaviscutellata (Mangabeira, 1942) furcata (Mangabeira, 1941) geniculata (Mangabeira, 1941) gomezi (Nitzulescu, 1931) gorbitzi (Blancas, 1959) hartmanni (Fairchild & Hertig, 1957) hernandezi (Ortíz, 1965) hirsuta (Mangabeira, 1942) howardi Young, 1979 ignacioi Young, 1972 infraspinosa (Mangabeira, 1941) isovespertilionis (Fairchild & Hertig, 1958) lerayi Le Pont, Martínez, Torrez-Espejo & Dujardin, 1998 lichyi (Floch & Abonnenc, 1950) limafalcaoae (Wolff & Galati, 2002) longiflocosa Osorno-Mesa, Morales, Osorno & Muñoz, 1970 longipalpis (Lutz & Neiva, 1912) longispina (Mangabeira, 1942) lutziana (Costa Lima, 1932) marinkellei Young, 1979 martinezi Young & Morales, 1987 micropyga (Mangabeira, 1942) migonei (França, 1920) monstruosa (Floch & Abonnenc, 1944) moralesi Young, 1979 nematoducta Young & Arias, 1984 nevesi (Damasceno & Arouck, 1956) nocticola Young, 1973 ...Continued on next page CATALOGUE OF PSYCHODIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 217 TABLE 1. (Continued) Subfamily Genus Species Author nuneztovari (Ortíz, 1954) oresbia (Fairchild & Hertig, 1961) osornoi (Ristorcelli & Van Ty, 1941) ovallesi (Ortíz, 1952) pabloi Barreto, Burbano & Young, 2002 pajoti Abonnenc, Léger & Fauran, 1979 panamensis (Shannon, 1926) paraensis (Costa Lima, 1941) pia (Fairchild & Hertig, 1961) pilosa (Damasceno & Causey, 1944) preclara Young & Arias, 1984 punctigeniculata (Floch & Abonnenc, 1944) quasitownsendi Osorno, Osorno-Mesa & Morales, 1972 rangeliana (Ortíz, 1952) reburra (Fairchild & Hertig, 1961) recurva Young, 1973 reducta Feliciangeli, Ramírez-Pérez & Ramírez, 1988 richardwardi Ready & Fraiha, 1981 rorotaensis (Floch & Abonnenc, 1944) rosabali (Fairchild & Hertig, 1956) ruii Arias & Young, 1982 runoides (Fairchild & Hertig, 1953) saltuosa Young, 1979 sanguinaria (Fairchild & Hertig, 1957) saulensis (Floch & Abonnenc, 1944) sauroida Osorno-Mesa, Morales & Osorno, 1972 scaffi (Damasceno & Arouck, 1956) scorzai (Ortíz, 1965) sericea (Floch & Abonnenc, 1944) serrana (Damasceno & Arouck, 1949) shannoni (Dyar, 1929) shawi Fraiha, Ward & Ready, 1981 sherlocki Martins, Silva & Falcão, 1971 sipani Fernández, Carbajal, Alexander & Need, 1994 sordellii (Shannon & Del Ponte, 1927) spinicrassa Morales, Osorno-Mesa, Osorno & Muñoz, 1969 strictivilla Young, 1979 suapiensis Le Pont, Torrez-Espejo & Dujardin, 1997 thula Young, 1979 tihuiliensis Le Pont, Torrez-Espejo & Dujardin, 1997 ...Continued on next page 218 · Zootaxa 4122 (1) © 2016 Magnolia Press BEJARANO & ESTRADA TABLE 1. (Continued) Subfamily Genus Species Author tolimensis Carrasquilla, Munstermann, Marín, Ocampo & Ferro, 2012 tortura Young & Rogers, 1984 torvida Young, Morales & Ferro, 1994 townsendi (Ortíz, 1960) trapidoi (Fairchild & Hertig, 1952) triacantha (Mangabeira, 1942) trinidadensis (Newstead, 1922) triramula (Fairchild & Hertig, 1952) tuberculata (Mangabeira, 1941) ubiquitalis (Mangabeira, 1942) umbratilis Ward & Fraiha, 1977 undulata (Fairchild & Hertig, 1950) vattierae Le Pont & Desjeux, 1992 velezi Bejarano, Vivero & Uribe, 2010 venezuelensis (Floch & Abonnenc, 1948) vespertilionis (Fairchild & Hertig, 1947) wagleyi (Causey & Damasceno, 1945) walkeri (Newstead, 1914) witoto Young & Morales, 1987 yencanensis (Ortíz, 1965) ylephiletor (Fairchild & Hertig, 1952) youngi Feliciangeli & Murillo, 1987 yucumensis (Le Pont, Caillard, Tibayrenc & Desjeux, 1986) yuilli Young & Porter, 1972 nigrosaccula Fairchild & Hertig, 1951 rotundipennis Fairchild & Hertig, 1951 guhli Wagner & Joost, 1994 martinezi Wagner & Joost, 1994 obandoi Wagner & Joost, 1994 Australopericoma roessleri (Wagner & Joost, 1994) Balbagathis sp. Clogmia albipunctata (Williston, 1893) Didicrum colombensis Moya-Arévalo, Ibáñez-Bernal & Suárez-Landa, 2012 Feuerborniella paramuna Cordeiro, 2014 Lepidiella larryi Ibáñez-Bernal, 2010 Maruina campesina Hogue, 1973 chaborra Hogue, 1973 chiringa Hogue, 1990 colombicana Wagner & Joost, 1994 Warileya Psychodinae Arisemus ...Continued on next page CATALOGUE OF PSYCHODIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 219 TABLE 1. (Continued) Subfamily Sycoracinae Trichomyiinae Genus Species Paramormia sp. Parasetomima sp. Psychoda aitkeni Quate, 1959 alternata Say, 1824 alternicula Quate, 1955 bogotensis Wagner & Joost, 1994 cordiforma Quate, 1996 mimica Quate, 1996 savaiiensis Edwards, 1928 andicola Young, 1979 colombiensis Young, 1979 fairchildi Young, 1979 trispinosa Young, 1979 utriensis Bejarano, Duque & Vélez, 2008 andina Bejarano, Pérez-Doria & Sierra, 2010 brevitarsa (Rapp, 1945) colosensis Pérez-Doria, Hernández & Bejarano, 2010 quimbaya Bejarano, Pérez-Doria & Sierra, 2009 risaraldensis Bejarano, Pérez-Doria & Sierra, 2009 Sycorax Trichomyia Author References Abonnenc, E., Arias, J.R., Léger, N. & Young, D.G. (1980) Sur Lutzomyia davisi (Root, 1934) et les espèces de morphologie comparable (Diptera, Phlebotomidae). Annales de Parasitologie Humaine et Comparée, 55 (6), 707–719. Abonnenc, E. & Léger, N. (1976) Sur une classification rationnelle des diptères Phlebotomidae. 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(1995) Entomological aspects of a leishmaniasis control project based on community participation in riverine communities of SW Colombia. Boletín de Malariología y Saneamiento Ambiental, 35 (Supplement 1), 29–40. Alexander, J.B. (1987) A new species of Nemapalpus (Diptera; Psychodidae; Bruchomyiinae) from northeastern Colombia. Florida Entomologist, 70 (3), 376–381. http://dx.doi.org/10.2307/3495071 Alexander, J.B., Takaoka, H., Eshita, Y., Gómez, E.A. & Hashiguchi, Y. (1992) New records of phlebotomine sandflies (Diptera: Psychodidae) from Ecuador. Memórias do Instituto Oswaldo Cruz, 87 (1), 123–130. http://dx.doi.org/10.1590/S0074-02761992000100019 Ali, H.B., Augul, R.S.H. & Mzhr, N.N. (2013) New record of moth fly species; Psychoda alternata Say, 1824 (Diptera: Psychodidae: Psychodinae) from Iraq. Journal of University of Anbar for Pure science, 7 (1), 11–14. Alvar, J., Vélez, I.D., Bern, C., Herrero, M., Desjeux, P., Cano, J., Jannin, J. & den Boer, M. (2012) Leishmaniasis worldwide and global estimates of its incidence. PLoS ONE, 7 (5), e35671. http://dx.doi.org/10.1371/journal.pone.0035671 Araújo, M.A. & Bravo, F. (2013) A new subgenus and species of Neotropical Trichomyia (Diptera: Psychodidae). Zoologia, 30 220 · Zootaxa 4122 (1) © 2016 Magnolia Press BEJARANO & ESTRADA (4), 458–462. http://dx.doi.org/10.1590/S1984-46702013000400014 Arias, J.R. & Young, D.G. (1982) Sandflies of the Central Amazon of Brazil. Lutzomyia (Trichophoromyia) ruii n. sp. (Diptera: Psychodidae). Revista Brasileira de Biologia, 42 (1), 249–253. Arismendi-Solano, M.J. (2005) Estudio de foco de leishmaniosis en la zona de “tierra caliente” del municipio de Sucre Santander. Trabajo de Grado. Escuela de Biología, Universidad Industrial del Santander, Bucaramanga, 48 pp. Arrivillaga, J., Pérez, C., Flores, M., Enriquez, S., Vaca, F., Medina, B. & Benítez, W. 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(Diptera, Psychodidae). Revista Brasileira de Biologia, 3 (2), 183–189. Barretto, M.P. & Duret, J.P. (1953) Uma nova espécie de flebótomo da República Argentina: Phlebotomus microcephalus n. sp. (Diptera, Psychodidae). Papéis Avulsos do Departamento de Zoologia, 11 (20), 341–344. Bejarano, E.E. (2002) Sobre la evolución de los flebotomíneos americanos (Diptera: Psychodidae): un llamado a los sistemáticos moleculares. Revista Colombiana de Entomología, 28 (2), 211–212. Bejarano, E.E. (2006a) Lista actualizada de los psicódidos (Diptera: Psychodidae) de Colombia. Folia Entomológica Mexicana, 45 (1), 47–56. Bejarano, E.E. (2006b) Subfamilia Trichomyiinae (Psychodidae) en el territorio continental e insular colombiano. Acta Biológica Colombiana, 11 (1), 37–41. Bejarano, E.E., Castro, M., Perez-Doria, A., Hernández-Oviedo, E., Vélez, A. & Vélez, I.D. (2007a) Primer informe de Lutzomyia França en el departamento de Guainía, Amazonia Colombiana, y de Brumptomyia mesai Sherlock (Diptera: Psychodidae) en el litoral Caribe Colombiano. Neotropical Entomology, 36 (6), 990–993. http://dx.doi.org/10.1590/S1519-566X2007000600027 CATALOGUE OF PSYCHODIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 221 Bejarano, E.E., Duque, P. & Vélez, I.D. (2004) Taxonomy and distribution of the series pia of the Lutzomyia verrucarum group (Diptera: Psychodidae), with a description of Lutzomyia emberai n. sp. Journal of Medical Entomology, 41 (5), 833–841. http://dx.doi.org/10.1603/0022-2585-41.5.833 Bejarano, E.E., Duque, P. & Vélez, I.D. (2006) Redescripción de la hembra de Lutzomyia vattierae (Diptera: Psychodidae, Phlebotominae) de la serranía de La Macarena, Colombia. Biomédica, 26 (4), 556–561. http://dx.doi.org/10.7705/biomedica.v26i4.325 Bejarano, E.E., Duque, P. & Vélez, I.D. 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Transactions of the Royal Society of Tropical Medicine and Hygiene, 77 (2), 280. http://dx.doi.org/10.1016/0035-9203(83)90097-4 Zuluaga, W.A., López, Y.L., Osorio, L., Salazar, L.F., González, M.C., Ríos, C.M., Wolff, M.I. & Escobar, J.P. (2012) Vigilancia de insectos de importancia en salud pública durante la construcción de los proyectos hidroeléctricos Porce II y Porce III, Antioquia, Colombia, 1990–2009. Biomédica, 32 (3), 321–331. http://dx.doi.org/10.7705/biomedica.v32i3.668 238 · Zootaxa 4122 (1) © 2016 Magnolia Press BEJARANO & ESTRADA Zootaxa 4122 (1): 239–245 http://www.mapress.com/j/zt/ Copyright © 2016 Magnolia Press Article ISSN 1175-5326 (print edition) ZOOTAXA ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.4122.1.21 http://zoobank.org/urn:lsid:zoobank.org:pub:6501CADA-5D0C-41DB-BE20-D012C3452AAA FAMILY SCATOPSIDAE DALTON DE SOUZA AMORIM Departamento de Biologia, FFCLRP, Universidade de São Paulo, Av. Bandeirantes 3900, 14.040–901 Ribeirão Preto SP, BRAZIL. E–mail: dsamorim@usp.br Abstract An updated catalogue of the family Scatopsidae in Colombia is presented. Seven species of six genera have being recorded for the country, but four of these species are still undescribed. The Ectaetiinae genus Ectaetia, the Psectrosciarinae genera Anapausis and Psectrosciara, and at least the Scatopsinae genera Scatopse, Efcookella, and Abrhexoza are expected to be present in the fauna of the family in Colombia. Key words: catalogue, Colombia, Distribution, Scatopsidae, taxonomy Introduction Scatopsids are tiny midges, usually black or blackish brown, exceptionally with areas with yellow, shiny or dull, from 0.6 to 3.0 mm long. The maxillary palpus is 1–segmented in the entire family and the antenna have 5–10 flagellomeres usually short and with a regular row of setae. The radial veins in the wing are well sclerotized, but the medial and cubital veins are typically faint. The abdominal membrane has a unique condition, the microtrichia being modified forming parallel long stripes along each segment. They do not fly well, but may run over the substrate. Scatopsid larvae live in a variety of substrates, including decaying plant or animal material, humid wood holes etc. Pupae are found envolved in the exuvia of the last larval instar, a feature shared with Canthyloscelidae. The relationships of the Scatopsoidea with the Bibionomorpha was originally based on the wing venation (Hennig 1954, 1973). Larval features (Wood & Borkent 1989) and the fusion of the meron to the thoracic pleural sclerites (Amorim 1994), however, strongly suggest a relationship with the suborder Psychodomorpha (see Amorim & Yeates 2006). Some recent analyses (e.g., Wiegmann et al. 2011) using molecular data point out to a relationship with the Bibionomorpha, but the taxon sampling of these papers are sufficiently poor to allow a misplacement of some important families in the system. The connections of Scatopsidae and Canthyloscelidae, on the other hand, have never been questioned (e.g., Hennig 1954, 1973; Wood & Borkent 1989; Amorim 1994). Recently, the family Valeseguyidae has been proposed by Amorim & Grimaldi (2006) to be the sister group of the clade composed of Canthyloscelidae + Scatopsidae. The Scatopsidae presently include 36 genera in the world that encompass almost 400 described species. The family, however, is certainly much more diverse, having received little attention in the field or in sorting. Amorim (1994) proposed a phylogenetic classification for the family based on Amorim (1982), with four taxa with subfamilial status. The Aspistinae have two Holarctic genera, the Ectaetiinae have a single genus, and the Psectrosciarinae have two tribes, each with a single genus. The Scatopsinae enclose four tribes and have the bulk of the diversity in the family. The tribes are Rhegmoclematini, Scatopsini, Colobostematini, and Swammerdamellini. Rhegmoclematini is further divided into four subtribes, Holoclematina, Diamphidicina, Rhegmoclematina, and Parascatopsina. In the Rhegmoclematini, Holoclema Amorim & Haenni is endemic to the neotropics and Diamphidicus Cook is known from Chile and Australia. Parmaferia Cook is absent in the Neotropical region, while both other genera of the tribe—Thripomorpha Enderlein and Rhegmoclema Enderlein—have Neotropical species, as well as Parascatopse Cook, and Neorhegmoclemina Cook. Austroclemina Cook is known only from Australia, while Rhegmoclemina is Accepted by S. Nihei: 10 Jun. 2015; published: 14 Jun. 2016 239 typically Holarctic, with a species known from Mexico. The Scatopsini have four genera, of which Apiloscatopse Cook, Reichertella Enderlein and Pharsoreichertella Cook are absent in the Neotropical region. Scatopse is known from an introduced, Cosmopolitan species, S. notata Linnaeus. In the Colobostematini, the genera Colobostema Enderlein, Cookella Freeman, and Holoplagia Enderlein are present in the recent fauna. The extinct genus Procolobostema Cook is known from Mexico and Dominican Republic amber fossils. Ferneiella Cook, Borneoscatopse Freeman, and Lumpuria Edwards are absent in the Neotropics. In the Swammerdamellini, the genera Pararhexosa Freeman, Swammerdamella Enderlein, Coboldia Melander, Akorhexoza Cook, Abrhexosa Freeman, Quateiella Cook and Brahemyia Amorim have species known from the Neotropical region, while Hawomersleya Cook is exclusive from Australia and Rhexosa Enderlein is known from the Nearctic and Palearctic regions. Fossil records that may belong to the stem–group of the family, with a branched R5, are known from Upper Jurassic/Lower Cretaceous (Rohdendorf 1946; Kalugina & Kovalev 1985), included in a separate extinct family, Protoscatopsidae. This is congruent with the presence of Canthyloscelidae in the Jurassic (Amorim 2000a), in such a way that this would be the same age for the sister family, the Scatopsidae. The earliest scatopsid compression fossils known are from the Lower Cretaceous, described by Kovalev (1986). There are unpublished Cretaceous amber fossils of the subfamily Ectaetiinae from different deposits (see reviews in Amorim 1998; Nel & Prokop 2004). Records and genera of Scatopsidae expected to occur in Colombia For Colombia, seven species of six genera have being recorded, though four of these species are still undescribed (Table 1). Additionally, there are genera still without records, but that are expected to occur due to their presence in close by areas. Subfamily Ectatiinae Enderlein The genus Ectaetia Enderlein is known from different parts of the Neotropical Region and is expected to occur in Colombia, even though there are no records yet. Subfamily Psectrosciarinae Cook The Neotropical species of Anapausis Enderlein, 1912 were revised by Amorim & Balbi (2006), with species known from Chile, southern Brazil, Panama and Costa Rica. The genus is most certainly present in Colombia. Psectrosciara Kieffer, 1911 is known from Mexico, Costa Rica and Belize, besides a species from Chile, and shall be found in the Colombian fauna of the family. Subfamily Scatopsinae Newman Tribe Rhegmoclematini Cook Thripomorpha Enderlein, 1905 Neorhegmoclemina Cook, 1955 Tribe Scatopsini Cook There are no records of Scatopse Geoffroy, 1762 for Colombia, but S. notata Linnaeus is a species with worldwide distribution and should be found in Colombia at some stage. Tribe Colobostematini Cook Colobostema Enderlein, 1926 Holoplagia Enderlein, 1912 240 · Zootaxa 4122 (1) © 2016 Magnolia Press AMORIM Species of the genus Efcookella Haenni, 1998 are known from South America and it would be no surprise that it comes to be found in Colombia. Tribe Swammerdamellini Cook Swammerdamella Enderlein, 1912 Coboldia Melander, 1916 Brahemyia Amorim, 2007 At least the genus Abrhexoza Freeman, 1990 should also be present in Colombia. Acronyms used for the depositories AMNH—American Museum of Natural History, Department of Entomology, New York, USA. BMNH—The Natural History Museum, Department of Entomology, London, UK. DEI—Deutsches Entomologisches Institut, Deutsche Akademie der Landwirts–wissenschaften zu Berlin, Eberswalde, Germany CAIM—Collection of Arthropods with Medical Importance (CAIM), Instituto de Diagnóstico y Referencia Epidemiológicos, Mexico, D.F., Mexico CAS—California Academy of Sciences, Department of Entomology, San Francisco, USA CUI—Cornell University, Cornell University Insect Collection, Cornell, USA MNM—Magyar Természettudományi Múzeum (Hungarian Natural History Museum), Budapest, Hungary MHNN—Muséum d’Histoire Naturelle Neuchâtel, Neuchâtel, Switzerland MNHNP—Muséum National d’Histoire Naturelle, Paris, France MZSP—Museu de Zoologia da Universidade de São Paulo, São Paulo, Brazil NMNH—National Museum of Natural History, Washington, D.C., USA PAN—Polish Academy of Science, Museum of the Institute of Zoology, Warsaw, Poland. UCMP—University of California Museum of Paleontology, Berkeley, USA UMIC—University of Minnesota Insect Collection, St. Paul, USA ZMB—Zoologisches Museum Berlin, Berlin, Germany List of abbreviations cat.—catalogue distr.—geographic distribution n. comb. —new combination orig. des.— original designation pl.— plate redescr.— redescription refs.— references rev.—revision syn.—synonymy term.—terminalia T—type HT—holotype M (m#)—Male F (f#)—Female CATALOGUE OF SCATOPSIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 241 Catalogue of Scatopsidae of Colombia Family Scatopsidae Newman, 1834 Subfamily Scatopsinae Newman, 1834 Tribe Rhegmoclematini Cook, 1955 Subtribe Rhegmoclematina Cook, 1965a Genus Thripomorpha Enderlein Thripomorpha Enderlein, 1905: 448. Type species, Thripomorpha paludicola Enderlein, 1905 (orig. des.). Refs.: Cook, 1955a (rev.). Aldrovandiella Enderlein, 1912: 278. Type species, Scatopse halterata Meigen, 1838 (orig. des.). Refs.: Haenni, 1995: 331 (syn.). blantoni (Cook), 1955b: 363, pl. IV i–k (M segment 7, M term., wing) (Rhegmoclema). Type locality: Panama, Bocas del Toro Prov., Almirante. HT M (NMNH). N. comb. Distr.: Panama, Colombia (Valle del Cauca(Rio Raposo)). Genus Neorhegmoclemina Cook Rhegmoclemina (Neorhegmoclemina) Cook, 1955: 358. Type species, Rhegmoclemina parva Cook, 1955 (orig. des.). Refs.: Cook, 1955b (rev.); Amorim, 1994 (generic status). undescribed sp. 1 Locality: Colombia, Valle del Cauca, Rio Raposo. undescribed sp. 2 Locality: Colombia, Valle del Cauca, Rio Raposo. Tribe Colobostematini Amorim, 1994 Genus Colobostema Enderlein Colobostema Enderlein, 1926: 140. Type species, Colobostema oldenbergi Enderlein, 1926 (orig. des.). Refs.: Cook, 1956c (rev.). undescribed sp. 1 Locality: Colombia, Valle del Cauca, Rio Raposo. Genus Holoplagia Enderlein Holoplagia Enderlein, 1912: 267. Type species, Scatopse transversalis Loew, 1846 (orig. des.) (= Scatopse lucifuga Loew). Refs.: Cook, 1957 (rev.); Freeman, 1990 (redefinition). guamensis (Johannsen), 1946: 187. 1a–e [wing (the wing figured is that of Coboldia fuscipes, not of H. guamensis], M term.) (Scatopse). Type locality: “Guam”. T (CUI). Distr.: Micronesia, Hawaii, Truk, Ghana, Accra, Panama, Colombia, Brazil. Refs.: Hardy, 1956: 91 (redescr., n. comb.), figs. 2a (mx. palpus), 2b (wing), 2c (M segment 7 and term.), 2d–e (F segment 7 and term.); Cook, 1957: 610 (redescr.), figs. 7C (M term.), 7E (F term.), 7K (wing); Cook, 1967 (cat.); Cook, 1989 (cat.). 242 · Zootaxa 4122 (1) © 2016 Magnolia Press AMORIM Tribe Swammerdamellini Cook, 1972 Genus Coboldia Melander Coboldia Melander, 1916: 17. Type species, C. formicarum Melander, 1916 (orig. des.) = Scatopse fuscipes Meigen, 1830. Refs.: Cook, 1974 (rev.), 1981; Meade & Cook, 1961 (biology of immatures). Rhaeboza Enderlein, 1936: 55. Type species, Scatopse fuscipes Meigen, 1830 (orig. des.). Masatierra Enderlein, 1938: 666. Type species, M. ferruginea Enderlein, 1938 (orig. des.) = Scatopse fuscipes Meigen. fuscipes (Meigen), 1830: 314 (Scatopse). Type locality: “Europe”. T (MNHNP) (Duda, 1928a: 22). Distr.: Cosmopolitan. Refs.: Hardy, 1956: 99 (as Scatopse) (redescr., illustr.); Cook, 1957: figs. 3A–B (M term.), 3D (spiracular sclerite), 3N (F sternite 7), 5D (F term.), 5F (M segment 7), 6B (wing), 6G (M tergite 8); Cook, 1963: 17, figs. 47 (F term.), 48 (M segment 7), 49 (M term.), 59 (wing); Rabello & Forattini, 1962: 303 (as Rhexoza coutinhoi) (descr., immature, biol.); Cook, 1974: 70, figs. 78 (M term.); 79 (F term.); Cook, 1967 (cat., as Scatopse); Haenni & Vaillant, 1994 (biol.). Genus Brahemyia Amorim Brahemyia Amorim, 2007: 48. Type species, Rhexoza lobata Cook, 1956 (orig. des.). Refs.: The larva attributed to the genus was illustrated in Guimarães & Amorim (2006). undescribed sp. 1 Locality: Colombia, Valle del Cauca, Rio Raposo TABLE 1. Scatopsidae species recorded from Colombia. Subfamily Genus Species Author Scatopsinae Thripomorpha blantoni Cook, 1955 Neorhegmoclemina undescribed sp. 1 Neorhegmoclemina undescribed sp. 2 Colobostema undescribed sp. 1 Holoplagia guamensis (Johannsen, 1946) Coboldia fuscipes (Meigen, 1830) Brahemyia undescribed sp. 1 References Amorim, D.S. (1982) Sistemática filogenética dos Scatopsidae. Master’s Thesis, Universidade de São Paulo, São Paulo, v + 173 pp + 74 pls. Amorim, D.S. (1994) A new suprageneric classification of the Scatopsidae (Diptera: Psychodomorpha). Iheringia, Série Zoologia, 77, 107–112. Amorim, D.S. (1998) Amber fossil Scatopsidae (Diptera: Scatopsidae). I. Considerations on Scatopsidae described fossils, Procolobostema roseni sp.n. from Dominican amber and the position of Procolobostema in the family. American Museum Novitates, 3227, 1–17. Amorim, D.S. (2000a) A new phylogeny and phylogenetic classification of the Canthyloscelidae (Diptera: Psychodomorpha). 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Moscow, Paleontologicheskogo Instituta Akademii a Nauk, 198 pp. [in Russian] Kovalev, V.G. (1986) Insects in the early Cretaceous ecosystems of Western Mongolia. Trudy Sovremenye–Mongolsky Paleogicheskii Ekspeditsiya, Academy of Sciences of the USSR, 28, 125–154. [in Russian] Meade, A.B. & Cook, E.F. (1961) Notes on the biology of Scatopse fuscipes Meigen. Entomological News, 72, 13–18. Meigen, J.W. (1830) Systematische Beschreibung der bekannten europäischen zweiflügeligen Insekten. Vol. 6. Schulz, Hamm, xi + 401 pp. Meigen, J.W. (1838) Systematische Beschreibung der bekannten europäischen zweiflügeligen Insekten. Siebenter Theil oder Supplementband. Schultz, Hamm, xii + 434 + [1] pp. Melander, A.L. (1916) The dipterous family Scatopsidae. Bulletin Washington Agricultural Experiment Station, 130, 1–21. Nel, A. & Prokop, J. (2004) New Paleogene Scatopsidae (Diptera: Nematocera) in amber from the Czech Republic and France. Acta Societatis Zooligicae Bohemicae, 68, 91–98. Newman, E. (1834) Attempted division of the British Insects into natural Orders. Entomologist´s Magazine, 2, 379–441. Rabello, E.S. & Forattini, O.P. (1962) Sobre uma nova espécie do gênero Rhexoza Enderlein (Diptera, Scatopsidae). Papéis Avulsos do Departamento de Zoologia, 15, 303–309. Rohdendorf, B. (1946) [Evolution of wing and phylogeny of nematoceran Diptera Oligoneura]. Trudy Paleontologicheskogo Instituta Akademii a Nauk, SSSR, 13 (2), 5–108. [in Russian] Wiegmann, B.M., Trautwein, M., Winkler, I., Barr, N., Kim, J.-W, Lambkin, C., Bertone, M.A., Cassel, B.K., Bayless, K.M., Heimberg, A.M., Wheeler, B.M., Peterson, K.J., Pape, T., Sinclair, B.J., Skevington, J.H., Blagoderov, V., Caravas, J., Kutty, S.N., Schmidt-Ott, U., Kampmeier, G.E., Thompson, F.C., Grimaldi, D.A., Beckenbach, A.T., Courtney, G.W., Friedrich, M., Meier, R. & Yeates, D.K. (2011) Episodic radiations in the fly tree of life. Proceedings of the National Academy of Sciences of the United States of America, 108, 5690–5695. http://dx.doi.org/10.1073/pnas.1012675108 Wood, D.M. & Borkent, A. (1989) Phylogeny and classification of the Nematocera. In: McAlpine, J.F. & D.M. Wood (Coords.), Manual of the Nearctic Diptera. Vol. 3. Research Branch Monograph 29, Agriculture Canada, Ottawa, pp. 1333–1370. CATALOGUE OF SCATOPSIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 245 Zootaxa 4122 (1): 246–248 http://www.mapress.com/j/zt/ Copyright © 2016 Magnolia Press Article ISSN 1175-5326 (print edition) ZOOTAXA ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.4122.1.22 http://zoobank.org/urn:lsid:zoobank.org:pub:9F895302-0A36-4F35-8C9C-1D5A3A7052DC FAMILY RHAGIONIDAE CHARLES MORPHY D. SANTOS1* & DANIEL D. D. CARMO2 1 Universidade Federal do ABC, Centro de Ciências Naturais e Humanas, Rua Santa Adélia, 166, Bairro Bangu, 09210-170, Santo André, SP, Brazil. E-mail: charlesmorphy@gmail.com. 2 Instituto Nacional de Pesquisas da Amazônia, Av. André Araújo, 2936, Bairro Aleixo, 69060-001, Manaus, AM, Brazil. E-mail: dandorndias@gmail.com *Corresponding author: charlesmorphy@gmail.com Abstract The family Rhagionidae is one of the oldest Brachyeran lineages. Its monophyly is still uncertain. There are four rhagionid genera distributed in Neotropical Region but only three species of Chrysopilus are found in Colombia. Key words: catalogue, Chrysopilus, Colombia, distribution, Neotropical, Rhagionidae Introduction Rhagionidae (snipe flies) is one of the families that constitute the infraorder Tabanomorpha, along with Austroleptidae, Vermileonidae, Pelecorhynchidae, Oreoleptidae, Tabanidae and Athericidae (Stuckenberg 2001, Zloty et al. 2005). The group comprises one of the most ancient lineages within Brachycera, with a fossil species from Lower Triassic (Krzeminski & Krzeminska 2003). The monophyly of Rhagionidae has been an issue of discussion (Nagatomi 1982, Stuckenberg 2001, Santos 2008, Kerr 2010). Kerr (2010) published the first cladistics analysis focusing specifically on the relationships among the rhagionid genera, based both on morphological and molecular evidences. According to him, Rhagionidae is monophyletic, but unpublished data from one of us (Santos) suggests that a group of rhagionid genera is more closely related to the clade (Oreoleptidae+(Athericidae+Tabanidae)) than to the other rhagionids. Such result corroborates the paraphyly of the family as proposed by Wiegmann et al. (2000) The biology of the Rhagionidae is poorly known. The larvae are reported as predators (Stuckenberg 2001, Kerr 2010). Adults of some Chrysopilus and Rhagio predate other insects, and at least one genus, Symphoromyia Frauenfeld, is hematophagous (Stuckenberg 2001). Species size range from small to medium (4–20 mm), and the adults rest on shrubs in rain forests (Santos 2008, Kerr 2010). Rhagionids can be differentiated by the absence of a scale on the postspiracular esclerite (Kerr 2010, Carvalho et al. 2012). In some genera, the female terminalia has an invagination in the proximal segment of the cercus, the posteroventral cercal lobe, also present in the Pelecorhynchidae (Carmo & Santos 2011, Fig. 9 A–D). Nagatomi (1982) and Kerr (2010) present diagnostic keys to the genera of Rhagionidae. There are approximately 720 extant species in the family, distributed in 22 genera (Santos 2008). Four of these are found in the Neotropics: Chrysopilus Macquart, Rhagio Fabricius, Atherimorpha White and Sierramyia Kerr (Lindner) (Santos & Amorim 2007, Santos 2006, 2008). Rhagio and Chrysopilus are the most diversified and widespread rhagionids (Nagatomi 1982, Santos 2008). Until the present moment, Chrysopilus is the only genus of Rhagionidae found in Colombia, with three reported species (James 1968, Santos 2008) (Table 1). This genus is diagnosed by gonocoxites fused with the sternite IX (Carmo & Santos 2011, Fig. 3 A,C) and by wing features such as R4+5 fork almost in a right angle (Santos & Amorim 2007). 246 Accepted by S. Nihei: 26 May 2015; published: 14 Jun. 2016 Acronyms used for the depositories NMW—National Museum Wales, Cardiff, United Kingdom. BMNH—British Museum of Natural History, London, United Kingdom. List of abbreviations cat.—catalogue descr.—description distr.—distribution ref.—reference subs. des.—subsequent designation Catalogue of Rhagionidae of Colombia Family Rhagionidae Samouelle, 1819 Genus Chrysopilus Macquart Chrysopilus Macquart, 1826: 403. Type species: Rhagio diadema (Fabricius) (subs. des. Westwood, 1840: 134). Ref.: Curran, 1931 (partial key). caducus (Wiedemann, 1828): 579. (Leptis) Type locality: “Brazil”. Distr.: Venezuela, Colombia, Brazil. Refs.: Lindner, 1924: 74 (descr.), James, 1968: 3 (cat.). elegans Schiner, 1868: 168. Type locality: “Colombia”. Distr.: Costa Rica to Peru (NMW). Ref.: James, 1968: 3 (cat.). lucifer (Walker, 1852): 164. Type locality: “Colombia”. Distr.: Colombia, Peru (BMNH). Ref.: James, 1968: 3 (cat.). TABLE 1. Rhagionidae species recorded from Colombia. Genus Species Author Chrysopilus caducus (Wiedemann, 1828) elegans Schiner, 1868 lucifer (Walker, 1852) References Carmo, D.D.D. & Santos, C.M.D. (2011) Morfologia comparada das terminálias masculina e feminina dos Rhagionidae (Diptera, Tabanomorpha) Neotropicais. Papéis Avulsos de Zoologia, 51 (30), 465–479. Carvalho, C.J.B, Rafael, J.A., Couri, M.S. & Silva, V.C. (2012) Diptera Linnaeus, 1758. In: Rafael, J.A., Mello, G.A.R., de Carvalho, C.J.B., Casari, S.A. & Constantino, R. (Eds.), Insetos do Brasil. Editora Holos, Ribeirão Preto, pp. 702–743. Kerr, P.K. (2010) Phylogeny and classification of Rhagionidae, with implications for Tabanomorpha (Diptera: Brachycera). Zootaxa, 2592, 1–133 Krzeminski, W. & Krzeminska, E. (2003) Triassic Diptera: descriptions, revisions and phylogenetic relations. Acta Zoologica Cracoviensia, 46 (Supplement), 153–184. CATALOGUE OF RHAGIONIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 247 James, M.T. (1968) Family Rhagionidae (Leptidae). In: Papavero, N. (Ed.), A catalogue of the Diptera of the Americas South of the United States. Museu de Zoologia da Universidade de São Paulo, São Paulo, pp. 1–12. Lindner, E. (1924) Dipterologische Studien. II. Rhagionidae neotropical. Konowia, 3, 65–75. Nagatomi, A. (1982) The genera of Rhagionidae (Diptera). Journal of Natural History, 16, 31–70. http://dx.doi.org/10.1080/00222938200770041 Santos, C.M.D. (2006) Description of two new species of Neorhagio (Diptera, Tabanomorpha, Rhagionidae), and remarks of a controversial female character. Zootaxa, 1174, 49–62. Santos, C.M.D. (2008) Geographical distribution of Tabanomorpha (Diptera, Brachycera): Athericidae, Austroleptidae, Oreoleptidae, Rhagionidae, and Vermileonidae. Entomobrasilis, 1 (3), 43–50. http://dx.doi.org/10.12741/ebrasilis.v1i3.25 Santos, C.M.D. & Amorim, D.S. (2007) Chrysopilus (Diptera: Rhagionidae) from Brazil: redescription of Chrysopilus fascipennis Bromley and description of eleven new species. Zootaxa, 1510, 1–33. Schiner, J.R. (1858) Diptera. In: Wullerstorf-Urbair, B. von (Ed.), Reise der österreichischen Fregatte Novara um die Erde in den Jahren 1857, 1858, 1859 unter den Befehlen des Commodore B. von Wüllerstorf-Urbair. Zoologischer Theil. Vol. 2. Abt. I. K. Gerold’s Son, Wien, 388 pp. Stuckenberg, B.R. (2001) Prunning the tree: a critical review of classifications of the Homeodacytla (Diptera, Brachycera), with new perspectives and an alternative classification. Studia Dipterologica, 8, 3–42. Wiedemann, C.R.W. (1828) Aussereuropäische zweiflügelige Insekten. Vol. 1. Schultz, Hamm, xxxii + 608 pp. Wiegmann, B.M., Tsaur, S.-C., Webb, D.W., Yeates, D.K. & Cassel, B.K. (2000) Monophyly and relationships of the Tabanomorpha (Diptera: Brachycera) based on 28S ribosomal gene sequences. Annals of the Entomological Society of America, 93, 1031–1038. http://dx.doi.org/10.1603/0013-8746(2000)093[1031:MAROTT]2.0.CO;2 Zloty, J. & Sinclair, B.J. (2005) Discovered in our backyard: a new genus and species of a new family from the Rocky Mountains of North America (Diptera, Tabanomorpha). Systematic Entomology, 30 (2), 248–266. http://dx.doi.org/10.1111/j.1365-3113.2005.00270.x 248 · Zootaxa 4122 (1) © 2016 Magnolia Press SANTOS & CARMO Zootaxa 4122 (1): 249–301 http://www.mapress.com/j/zt/ Copyright © 2016 Magnolia Press Article ISSN 1175-5326 (print edition) ZOOTAXA ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.4122.1.23 http://zoobank.org/urn:lsid:zoobank.org:pub:68C0653C-9D55-4E98-9CEF-A95318600585 FAMILY TABANIDAE MARTA WOLFF E1*. & DANIEL RAFAEL MIRANDA-ESQUIVEL2 1 Grupo de Entomologia, Instituto de Biología, Universidad de Antioquia, Colombia. E-mail: martha.wolff@udea.edu.co Laboratorio de Sistematica y Biogeografica, Escuela de Biología, Universidad Industrial de Santander Bucaramanga Colombia. E-mail: dmiranda@uis.edu.co *Corresponding author: martha.wolff@udea.edu.co 2 Abstract This catalogue presents 256 species distributed in 32 genera, six tribes and three subfamilies recorded to Colombia. For each species we present the available geographical information and most important references. Key words: catalogue, Colombia, Tabanidae, distribution Introduction Tabanidae are a large Tabanomorpha family, (Diptera, Brachycera) with a worldwide distribution and approximately 4350 described species. For the neotropics there are aproximately 1500 species. Adults can be recognized by their stout bodies, measuring between 3–30 mm length; a big head occupied mainly by the compound eyes, separated by front in females and males. The eyes may present transversal stripes with yellow and green coloration patterns. Antennae are composed by scape, pedicel and flagellum, this latter consists of one large basal segment and ends with a series of cancelled segments. Proboscis are strong and thick, adapted to penetrate tissues. Females are hematophagous while males are phytophagous; however, in many species, male and female feed exclusively on nectar. Adults are found from open areas to forests and have diurnal behavior, but some few species are crepuscular or nocturnal. Larvae are predators of diverse animals (arthropods, mollusks and annelids). This family has been well studied in part, because some species could be vectors of human and animals diseases. Studies on taxonomy as well as medical aspects of the family were particularly intensive in Colombia between the 1950s and the end of the century. One of the major studies was made by Bequaert & Renjifo (1946), who covered a significant area of the national territory, they listed 129 species with their respective localities. Also relevant are the works made by Lee, Fairchild & Barretto (1969) in the Valle del Cauca state and the monograph of Wilkerson (1979), about the horseflies of the Western Colombian, with emphasis in the departments of Chocó, Valle del Cauca and Cauca; in which includes some records to Antioquia, supplied by Porter. According to the information recorded in Colombia there are 256 species (Table 1). However, it is expected that number to be much higher, since there are many unexplored areas, such as the mountain ranges of the Andes, the Caribbean coast or the amazonian forests. This catalog and the classification with synonyms adopted is based on Henriques & Rafael (1995) revision of genus Acanthocera, Coscaron & Papavero (2009) catalogue of neotropical Diptera, Krolow & Henriques (2010) revision of genus Chlorotabanus, Turcatel et al (2010) revision of genus Stibasoma, Krolow et al (2015) revision of genus Pityocera and includes information about specific localities on each department. The departments are underline and specific localities are in brackets. At the end we present a table with all species, rank and author. Accepted by S. Nihei: 10 Apr. 2015; published: 14 Jun. 2016 249 Acronyms used for the depositories AMNH—American Museum of Natural History, New York, U.S.A. BMNH—The Natural History Museum, London, England. CAS—Department of Entomology, California Academy of Sciences, San Francisco, U.S.A. FSCA—Florida State Collection of Arthropods, Gainesville, U.S.A. IEAUT—Istituto di Entomologia Agraria della Università di Torino, Turin IOC—Instituto Oswaldo Cruz, Rio de Janeiro, Brazil. MCZ—Museum of Compartative Zoology, Harvard University, Cambridge, England. MNHNP—Muséum National d´Histoire Naturelle, Paris, France. MLUH—Wissenschaftlich Zoologie, Bereits Biowissenschaften, Martin Luther Universität, Halle am den Saale MNHUB—Nuseum für Naturkunde der Humboldt-Universität, Berlin. MZUM—Museum of Zoology, University of Michigan, Ann Arbor. U.S.A. NMW—Naturhistorisches Museum, Wien. NRS—Naturhistoriska Riksmuseet, Stockholm USNM—National Museum of Natural History, Smithsonian Institution, Washington, D. C. ZMKU—Zoologiske Museum, Copenhagen ZSBS—Zoologische Staatssammlung des Bayerischen Staates, Munich. Gainesville: Florida State Collection of Arthropods, Division of Plant Industry, 1911 34th Street S. W., P. O. Box 147100, Gainesville, Florida 32614-7100. U. S. A. List of abbreviations aut.—automatic cat.—catalogue col.—collected des.—designation distr.—geographic distribution i.e.—exemple km.—kilometers m.—meters mon.—monotypy nom. nov.—nomen novum (new name) orig. des.—by original designation pp.—pages preocc.—preoccupied refs.—references rev.—revision sp., spp.—species ssp.—subspecies subg.—subgenus subs.—subsequent var.—variety P.N.N. Parque Nacional Natural HT—Holotype NT—Neotype LT—Lectotype ST—Syntype PT—Paratype F (f#)—Female M (m#)—Male 250 · Zootaxa 4122 (1) © 2016 Magnolia Press E & MIRANDA-ESQUIVEL Catalogue of Tabanidae of Colombia Family Tabanidae Latreille, 1802 Subfamily Chrysopsinae Blanchard, 1840 Tribe Chrysopsini Blanchard, 1840 Genus Chrysops Meigen Chrysops Meigen, 1800: 23. Type species, Tabanus caecutiens Linnaeus, 1758 (subs. mon., Meigen, 1803: 267). Suppressed by I.C.Z.N., 1963: 339. Chrysops Meigen, 1803: 267. Type species, Tabanus caecutiens Linnaeus, 1758 (mon.). Chrysops, subg. Heterochrysops Kröber, 1920: 50. Type species, Chrysops flavipes Meigen, 1804 (subs. des., Bequaert, 1924: 31). Refs.: Fairchild, 1967a: 331. Chrisops Fraga G, 1936: 248, error. Refs.: Kröber, 1925-1926 (key, figs.); Philip, 1955a (key); Coscaron & Papavero, 2009: 8 (cat.). auroguttatus Kröber, 1930a: 71. Type locality: “Colombia, Trinidad”. HT F (BMNH). Distr.: Colombia (Cauca (Guapi), Chocó (La Teresita), Valle del Cauca (Rio Raposo)), Mexico to Colombia, Trinidad. Refs.: Kröber, 1934: 225 (cat.; as auroguttata); Bequaert & Renjifo-Salcedo, 1946: 58 (as auroguttata); Lee et al., 1969: 444 (as auroguttata); Fairchild, 1971: 30 (cat.); Moucha, 1976: 55 (cat.); Wilkerson, 1979: 182; Fairchild & Burger, 1994: 55 (cat.); Coscaron & Papavero, 2009: 9 (cat.). incisa Macquart of Bequaert, 1931: 535 and of Fairchild, 1942a: 3, misident. bulbicornis Lutz, 1911a: 79. Type locality: Brazil, NW. São Paulo. HT F (IOC). Distr.: Colombia (Chocó, Antioquia), Bolivia, Brazil, Paraguay, Peru, ?Trinidad. Refs.: Fairchild & Burger, 1994: 56 (cat.); Coscaron & Papavero, 2009: 9 (cat.). calogaster Schiner, 1868: 103. Type locality: “South America”. HT F (NMW). Distr.: Colombia (Amazonas (Leticia), Boyacá (Muzo), Cauca, Chocó (La Teresita; El Valle; Utria), Cundinamarca (Villeta), Meta (Restrepo)), Brazil, Costa Rica to Colombia, ?Peru. Refs.: Bequaert & Renjifo-Salcedo, 1946: 58; Fairchild, 1971: 30 (cat.); Moucha, 1976: 56 (cat.); Wilkerson, 1979: 184; Fairchild & Burger, 1994: 56 (cat.), Coscaron & Papavero, 2009: 9 (cat.). chiriquensis Fairchild, 1939: 259. Type locality: Panama, Chiriquí, Boquete, 3000-4000m. HT F (MCZ). Distr.: Colombia (Antioquia (Caucasia), Magdalena (Cincinnati, Santa Marta)), Guatemala, Mexico, Panama. Refs.: Bequaert & Renjifo-Salcedo, 1946: 58; Fairchild, 1971: 31 (cat.); Moucha, 1976: 57 (cat.); Wilkerson, 1979: 186; Fairchild & Burger, 1994: 56 (cat.); Parra-Henao et al., 2008: 182; Coscaron & Papavero, 2009: 9 (cat.). incisus Macquart, 1846: 173 (1846: 44). Type locality: “New Grenada” (Colombia). ST F (BMNH). Distr.: Bolivia, Brazil, Colombia, Guyana, Peru, Surinam. Refs.: Kröber, 1934: 227 (cat.; as incisa); Bequaert & RenjifoSalcedo, 1946: 58 (as incisa); Fairchild, 1971: 32 (cat.); Moucha, 1976: 61 (cat.); Fairchild & Burger, 1994: 57 (cat.); Coscaron & Papavero, 2009: 11 (cat.). fulviceps Walker, 1854: 285. brasiliensis Ricardo, 1901: 314. aurofasciata Kröber, 1926: 354 tenuifasciata Kröber, 1929a: 525. aureofasciatus Fairchild, 1967a: 335 (Kröber’s HT), error. laetus Fabricius, 1805: 112. Type locality: “South America. Smidt” [i.e., Guyana, between rivers Demerara and Essequibo; cf. Papavero, 1971: 21]. HT F (ZMKU). Distr.: Colombia (Amazonas (El Encanto), Antioquia (near Medellín; Porcecito, Rio Porce), Caquetá (Florencia), Chocó (Andagoya; Utría), Meta (Villavicencio; El Caibe; CATALOGUE OF TABANIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 251 Restrepo), Putumayo (Puerto Limón), Valle del Cauca (Sonso; Hacienda El Tablazo, Tuluá), Vaupés)), ?Argentina, Brazil, ?Paraguay, Surinam. Refs.: Kröber, 1934: 227 (cat. as laeta); Bequaert & Renjifo-Salcedo, 1946: 57 (as laeta); Fairchild & Burger, 1994: 58 (cat.); Coscaron & Papavero, 2009: 11 (cat.). tenuistria Kröber, 1926: 341(as laetus var.) leucospilus Wiedemann, 1828: 202. Type locality: “Brazil”. HT F (MNHUB). Distr.: Colombia (Antioquia (Providencia), Boyacá (Muzo), Meta (Restrepo, El Caibe; Villavicencio; Bosque Ocoa), Vaupés (between San José del Guaviare and Calamar)), Panama to Brazil, Paraguay. Refs.: Kröber, 1934: 228 (cat.); Bequaert & RenjifoSalcedo, 1946: 57 (as leucospila); Fairchild, 1971: 32 (cat.); Moucha, 1976: 62 (cat.); Wilkerson, 1979: 188; Fairchild & Burger, 1994: 58 (cat.); Coscaron & Papavero, 2009: 12 (cat.). guttuta Wiedemann, 1828: 203. leucospillus Krolow et al., 2007: 254 (list). melaenus Hine, 1925: 17. Type locality: Panama, Chiriquí, Progreso. HT F (MZUM). Distr.: Colombia (Chocó (La Teresita), Santander (Jasmín), Valle del Cauca (Bajo Calima, 10km E. Buenaventura)), ?Nicaragua, Costa Rica to Venezuela. Refs.: Bequaert & Renjifo-Salcedo, 1946: 58 (as melaena); Fairchild, 1971: 32 (cat.); Moucha, 1976: 63 (cat.); Wilkerson, 1979: 189; Fairchild & Burger, 1994: 58 (cat.) Coscaron & Papavero, 2009: 12 (cat.). incisus Macquart of Kröber, 1926: 344 (misident.). mexicanus Kröber, 1926: 248 (as calogaster var.). Type locality: Mexico, Colima, Volcán Colima. HT F (ZSBS). Distr.: Colombia (Antioquia (Providencia), Cauca, Chocó, Valle del Cauca (Rio Raposo)), México to Colombia. Refs.: Lee et al., 1969: 444 (as mexicana); Fairchild, 1971: 32 (cat.); Moucha, 1976: 63 (cat.); Wilkerson, 1979: 191; Fairchild & Burger, 1994: 58 (cat.); Coscaron & Papavero, 2009: 12 (cat). nexosus Fairchild (unpublished 1978) in Wilkerson, 1979: 192. Type locality: Panama, Veraguas, Santa Fé. HT F (FSCA). Distr.: W. Colombia (Valle del Cauca (Rio Sabaletas)), Costa Rica, Panama. Refs.: Wilkerson, 1979: 192; Fairchild & Burger, 1994: 58 (cat.); Coscaron & Papavero, 2009: 13 (cat.). mexicana Kröber of Fairchild, 1942a: 8, 1947: 565, part, misident. renjifoi Bequaert, 1946a: 7. Type locality: Colombia, Valle de Cauca, Rio Yurumangui, Veneral. HT F (MCZ). Distr.: Colombia (Valle de Cauca (Veneral, Rio Yurumari)). Refs.: Bequaert & Renjifo-Salcedo, 1946: 58; Fairchild, 1971: 34 (cat.); Moucha, 1976: 67 (cat.); Wilkerson, 1979: 194; Fairchild & Burger, 1994: 60 (cat.); Coscaron & Papavero, 2009: 14 (cat.). reticulatus Wilkerson, 1979: 195. Type locality: Panama, San Blás, Rio Mandinga. HT F, PT F 2 (FSCA). Distr.: Colombia (Valle del Cauca (Rio Raposo)), Panama. Refs.: Wilkerson, 1979: 195; Fairchild & Burger, 1994: 60 (cat.); Coscaron & Papavero, 2009: 14 (cat.). soror Kröber, 1925: 245. Type locality: “Venezuela”. HT F (NMW). Distr.: Colombia (Antioquia, Boyacá (Muzo), Valle del Cauca (Rio Raposo)), Belize, Costa Rica, Guatemala, Honduras, Panama, Venezuela. Refs.: Bequaert & Renjifo-Salcedo, 1946: 58; Lee et al., 1969: 445; Fairchild, 1971: 34 (cat.); Moucha, 1976: 68 (cat.); Wilkerson, 1979: 197; Fairchild & Burger, 1994: 60 (cat.); Coscaron & Papavero, 2009: 14 (cat.). varians Wiedemann, 1828: 208. Type locality: “Brazil”. LT F (NMW). Distr.: Colombia (Amazonas (El Encanto), Antioquia (near Medellìn, Porcecito, Rio Porce), Caldas, Caquetá (Florencia), Cauca (Guapi), Chocó (Andagoya, Utría), Meta (El Caibe, Restrepo, Villavicencio), Putumayo (Puerto Limón), Santander, Valle del Cauca (Bajo Calima, Sonso, Tuluá)), Argentina, Brazil, Ecuador, Guyana, Panama, Paraguay, Peru, Trinidad, Venezuela. Refs.: Lee et al., 1969: 445; Fairchild, 1971: 34 (cat.); Moucha, 1976: 69 (cat.); Wilkerson, 1979: 198; Fairchild & Burger, 1994: 60 (cat.); Coscaron & Papavero, 2009: 15 (cat); Pisciotti & Miranda, 2011: 71 (report). tardus Wiedemann, 1828: 577. sublaeta Philip, 1955b: 72 ( as laeta ssp.). laetus, authors, not Fabricius, misident. nigroviolaceus, authors, not Kröber, misident., part. 252 · Zootaxa 4122 (1) © 2016 Magnolia Press E & MIRANDA-ESQUIVEL variegatus (De Geer), 1776: 230 (Tabanus). Type locality: “Surinam”. HT F (NRS). Distr.: Colombia (Antioquia (near Medellìn; Porcecito, Rio Porce; Caucasia), Cauca (near Caloto, hacienda San Julian, Guapi), Chocó (La Teresita), Magdalena (Tucurinca), Meta (El Caibe, Villavicencio), Putumayo (Puerto Limón, Mocoa), Santander (Hacienda Florencia 20km W. San Vicente, Puerto Olaya), Valle del Cauca (near Cali, Municipio La Candelaria, Palmira Hacienda La Carmelita, Rio Frío), Vaupés (between San José del Guaviare and Calamar), West Indies, southern Mexico to Argentina. Refs.: Kröber, 1934: 224 (cat.; as variegata); Bequaert & Renjifo-Salcedo, 1946: 57 (as variegata); Lee et al., 1969: 445 (as variegata); Fairchild, 1971: 35 (cat.); Moucha, 1976: 69 (cat.); Wilkerson, 1979: 199; Fairchild & Burger, 1994: 61 (cat.); Parra-Henao et al., 2008: 182 (as variegata); Coscaron & Papavero, 2009: 15 (cat.). costatus Fabricius, 1794: 373 (Tabanus). molestus Guérin-Méneville, 1835: 542 (1844: 542) (preocc. Wiedemann, 1828). vulneratus Rondani, 1848: 104. subfascipennis Macquart, 1855: 55 (1855: 35). amazonius Rondani, 1863: 81. crucians Jaennicke, 1867: 312 (preocc. Wiedemann, 1828). peruviensis Kröber, 1925: 235 (as costatus var.). weberi Bequaert, 1946a: 9. Type locality: Guyana, Orenoque. HT F (MCZ). Distr.: E. Colombia (Vaupés (Mitú)), Brazil, French Guiana, Guyana, Peru, Venezuela. Refs.: Bequaert & Renjifo-Salcedo, 1946: 58; Fairchild, 1971: 35 (cat.); Fairchild & Burger, 1994: 61 (cat.); Coscaron & Papavero, 2009: 16 (cat.). Tribe Rhinomyzini Enderlein, 1922 Genus Betrequia Oldroyd Betrequia Oldroyd, 1970: 250. Type species, Betrequia ocellata Oldroyd, 1970 (orig. des.). ocellata Oldroyd, 1970: 252. Type locality: Brazil, Ceará, Paracuru [3°23’50"S, 39°2’23"W]. HT F (BMNH). Distr.: Colombia (without defined locality), Brazil. Refs.: Fairchild & Burger, 1994: 53 (cat.); Coscaron & Papavero, 2009: 18 (cat.). Subfamily Pangoniinae Rondani, 1856 Tribe Pangoniini Rondani, 1856 Genus Esenbeckia Rondani Esenbeckia Rondani, 1863: 83, 95 (as Esenbekia p. 83) (1864: 83). Type species, Silvius vulpes Wiedemann, 1828 (orig. des.). Esenbackia Surcouf, 1909: 257, error. Dyspangonia Lutz, 1905: 22, 27 (no. 11). Type species, Pangonia fuscipennis Wiedemann, 1828 (subs. des., Bequaert, 1924: 28). Refs.: Fairchild, 1961a: 239, Coscaron & Papavero, 2009: 21 (cat.). Subgenus Esenbeckia Rondani, 1863 balteata Wilkerson, 1979: 129. Type locality: Colombia, Valle, Rio Bravo, Playa Rica. HT F, l F PT (FSCA). Distr.: Colombia (Valle del Cauca). Refs.: Wilkerson, & Fairchild, 1983: 528; Fairchild & Burger, 1994: 20 (cat.); Coscaron & Papavero, 2009: 22 (cat.). bella Philip, 1961: 111. Type locality: Colombia, Magdalena, Don Diego, 100m. HT F (AMNH). Distr.: Colombia (Magdalena). Refs.: Fairchild, 1971: 6 (cat.); Moucha, 1976: 17 (cat.); Wilkerson & Fairchild, 1983: 530; Fairchild & Burger, 1994: 20 (cat.); Coscaron & Papavero, 2009: 22 (cat.). CATALOGUE OF TABANIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 253 cisandeana Wilkerson, & Fairchild, 1983: 531. Type locality: Colombia, Putumayo, Mocoa, 650m. HT F, PT F (FSCA). Distr.: Colombia (Putumayo), Chile, Peru. Refs.: Fairchild & Burger, 1994: 20 (cat.); Coscaron & Papavero, 2009: 22 (cat.). diaphana (Schiner), 1868: 99 (Pangonia). Type locality: “Colombia”. HT F (NMW). Distr.: Colombia (Meta (Restrepo), Putumayo (Puerto Limón; Urcusique; San Pedro)), Venezuela. Refs.: Kröber, 1934: 240 (cat.); Bequaert & Renjifo-Salcedo, 1946: 59; Fairchild, 1971: 6 (cat.); Moucha, 1976: 18 (cat.); Wilkerson, 1979: 21; 1978: 21; Fairchild & Burger, 1994: 21 (cat.); Coscaron & Papavero, 2009: 22 (cat.). gracilis Kröber, 1931a: 255. Type locality: Peru, Loreto, Pebas. HT F (MLUH). Distr.: Argentina, Colombia, Peru. Refs.: Kröber, 1934: 241 (cat.); Fairchild, 1971: 7 (cat.); Moucha, 1976: 18 (cat.); Fairchild & Burger, 1994: 22 (cat.); Coscaron & Papavero, 2009: 24 (cat.). insignis Kröber, 1931a: 255. Type locality: “Brazil”. HT ? (BMNH). Distr.: Colombia (Caquetá), Brazil. Refs.: Kröber, 1934: 242 (cat.); Fairchild, 1971: 8 (cat.); Moucha, 1976: 19 (cat.); Fairchild & Burger, 1994: 22 (cat.); Coscaron & Papavero, 2009: 24 (cat.). minuscula Wilkerson, 1979: 131. Type locality: Colombia, Valle, Rio Frío, La Carmelita. HT F, PT F (FSCA). Distr.: Colombia (Valle del Cauca (Rio Frío, La Carmelita). Refs.: Wilkerson & Fairchild, 1983: 545; Fairchild & Burger, 1994: 22 (cat.); Coscaron & Papavero, 2009: 24 (cat.). osornoi Fairchild, 1942d: 196 (as illota ssp.). Type locality: Panama, Panama Prov., Rio Chagres, Moja Polo. HT F (MCZ). Distr.: Colombia (Amazonas (Leticia), Cundinamarca, Meta (Restrepo), Santander (Rionegro)), Costa Rica to Colombia, Brazil, Guyana, Surinam, Trinidad, Venezuela. Refs.: Bequaert & Renjifo-Salcedo, 1946: 60 (as illota var.); Fairchild, 1971: 7 (cat.; as illota var.); Wilkerson, 1979: 130 (as E (E) illota osornoi); Fairchild & Burger, 1994: 23 (cat.); Coscaron & Papavero, 2009: 25 (cat.). guianense Fairchild, 1942d: 196 (as illota ssp.). meridionae Fairchild, 1942d: 196 (as illota ssp.). prasiniventris (Macquart), 1846: 157 (1846: 29) (Pangonia). Type locality: “Colombia”. Distr.: Colombia (Cundinamarca (Bogotá), Magdalena (Rio Frío; Aguadulce, San Lorenzo)), Guatemala to Ecuador, Brazil, Trinidad. Refs.: Kröber, 1934: 242 (cat.); Bequaert & Renjifo-Salcedo, 1946: 59; Fairchild, 1971: 8 (cat.); Moucha, 1976: 19 (cat.); Wilkerson, 1979: 133; Fairchild & Burger, 1994: 23(cat.); Coscarón & Papavero, 2009: 25 (cat.). semiviridis Ricardo, 1900a: 181 (Pangonia) . prasiventris Hunter, 1901: 134 (Pangonia). subvaria (Walker), 1848: 150 (Tabanus). Type locality: “Venezuela”. HT F (BMNH). Distr.: Colombia, Venezuela. Refs.: Kröber, 1934: 243 (cat.); Bequaert & Renjifo-Salcedo, 1946: 59; Fairchild, 1971: 9 (cat.); Moucha, 1976: 19 (cat.); Fairchild & Burger, 1994: 23 (cat.); Coscarón & Papavero, 2009: 26 (cat.). testaceiventris (Macquart), 1848: 169 (1848: 9) (Pangonia). Type locality: Ecuador, Quito. ST F (BMNH). Distr.: Colombia (Cundinamarca (Tapia Las Cruces, Bogotá), Meta (Villavicencio), Valle del Cauca (Peñas Blancas, Lago Calima)), ?Chile, Ecuador, Venezuela. Refs.: Bequaert & Renjifo-Salcedo, 1946: 60; Fairchild, 1971: 9 (cat.); Moucha, 1976: 19 (cat.); Wilkerson, 1979: 134; Fairchild & Burger, 1994: 23 (cat.); Coscarón & Papavero, 2009: 26 (cat.). umbra Walker, 1850a: 19 (Pangonia). sexmaculata Enderlein, 1925: 288. lindigi Kröber, 1934: 243 (cat.), nomen nudum. tigrina Wilkerson, 1979: 136. Type locality: Colombia, Valle, Peñas Blancas. HT F, PT F (FSCA). Distr.: Colombia (Valle del Cauca Peñas Blancas)), Ecuador. Refs.: Wilkerson & Fairchild, 1983: 555; Fairchild & Burger, 1994: 24 (cat.); Coscarón & Papavero, 2009: 26 (cat.). 254 · Zootaxa 4122 (1) © 2016 Magnolia Press E & MIRANDA-ESQUIVEL tinctipennis Kröber, 1931a: 256. Type locality: Colombia, Chocó, Condoto. HT F (BMNH). Distr.: Colombia (Chocó (Condoto), Valle del Cauca (Rio Raposo)). Refs.: Kröber, 1934: 243 (cat.); Bequaert & Renjifo-Salcedo, 1946: 59; Lee et al., 1969: 443; Fairchild, 1971: 9 (cat.); Moucha, 1976: 20 (cat.); Wilkerson, 1979: 138; Wilkerson & Fairchild, 1983: 555; Fairchild & Burger, 1994: 24 (cat.); Coscaron & Papavero, 2009: 26 (cat.). translucens (Macquart), 1846: 154 (1846: 26) (Pangonia). Type locality: “Brazil”. Distr.: Colombia (Antioquia (Providencia, La Tirana, 25km W., 22km S. Zaragoza), Boyacá (Muzo), Chocó (Rio Nimiquía), Putumayo (Mocoa, Puerto Asís), Valle del Cauca (Rio Raposo, Rio Sabaletas, Bajo Anchicayá, Candelaria)), México to Peru, N. Brazil. Refs.: Bequaert & Renjifo-Salcedo, 1946: 59; Barretto, 1957: 76; Lee et al., 1969: 443; Fairchild, 1971: 9 (cat.); Moucha, 1976: 20 (cat.); Wilkerson, 1979: 139; Wilkerson & Fairchild, 1983: 555; Fairchild & Burger, 1994: 24 (cat.); Coscaron & Papavero, 2009: 26 (cat.). fusiformis Walker, 1850a: 19 (Pangonia). fusciformis Hine, 1925: 7 (Pangonia), misspelling. Subgenus Proboscoides Philip Proboscoides Philip, 1943: 111 (as genus). Type species, Proboscoides fairchildi Philip, 1943 (orig. des.). Refs.: Coscaron & Papavero, 2009: 27 (cat.). ecuadorensis ssp. chagresensis Fairchild, 1942d: 192 (as sp.). Type locality: Panama, Colón, New San Juan on Rio Chagres. HT F (MCZ). Distr.: Colombia (Magdalena (Rio Frío)), Panama. Refs.: Bequaert & Renjifo-Salcedo, 1946: 59; Fairchild, 1971: 10 (cat.); Moucha, 1976: 20 (cat.; as syn. of ecuadorensis Lutz & Castro, 1935); Wilkerson, 1979: 141; Fairchild & Burger, 1994: 25 (cat.); Coscaron & Papavero, 2009: 27 (cat.). Tribe Scionini Enderlein, 1922 Genus Fidena Walker, 1850 Subgenus Fidena Walker, 1850 aureopygia Kröber, 1931b: 24. Type locality: Colombia, Chocó, Opogodó HT F (?), PT (BMNH). Distr.: Colombia (Chocó (Opogodó), Valle del Cauca (Ají, Rio Naya)), Ecuador. Refs.: Kröber, 1934: 245 (cat.); Bequaert & Renjifo-Salcedo, 1946: 61; Fairchild, 1971: 18 (cat.); Moucha, 1976: 34 (cat.); Wilkerson, 1979: 144; Fairchild & Burger, 1994: 40 (cat.); Coscaron & Papavero, 2009: 36 (cat.). auribarba (Enderlein), 1925: 276 (Melpia). Type locality: Colombia, Cordillera. HT F (MNHUB). Distr.: Colombia (Chocó (Condoto, Rios San Juan-Baudò), Valle del Cauca (41km E., 20km N. Buenaventura, Lago Calima, Bajo Anchicayá, Rio Raposo, Rio Sabaletas)), Ecuador, Panama. Refs.: Kröber, 1934: 245 (cat.); Bequaert & Renjifo-Salcedo, 1946: 61; Fairchild, 1971: 18 (cat.); Moucha, 1976: 34 (cat.); Lee et al., 1969: 444; Wilkerson, 1979: 145; Fairchild & Burger, 1994: 40 (cat.); Coscaron & Papavero, 2009: 36 (cat.). albibarba Enderlein, 1925: 276 (Melpia; as auribarba var.; not Enderlein, 1925: 293). columbiensis Kröber, 1930b: 177 (Melpia). eriomeroides (Lutz), 1909a: 649, (Epipsila). Type locality: Brazil, São Paulo, Serra da Cantareira, Cachoerinha. LT F (IOC). Distr.: Colombia (Antioquia (Anorí (La Tirana), San Roque (Providencia), 22km S. Zaragoza), Meta (Villavicencio, Restrepo), Nariño (Pasto), Valle del Cauca (Bajo Anchicayá)), Costa Rica to Peru, Brazil. Refs.: Fairchild, 1971: 19 (cat.); Moucha, 1976: 35 (cat.); Wilkerson, 1979: 147; Fairchild & Burger, 1994: 42 (cat.); Coscaron & Papavero, 2009: 38 (cat.). gracilis Kröber, 1930b: 181 (Melpia). nigricornis Kröber, 1931b: 19 (Epipsila). nitida Philip, 1958: 70. CATALOGUE OF TABANIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 255 flavipennis Kröber, 1931b: 24. Type locality: Venezuela, Guarino. HT M (MNHUB). Distr.: Colombia (Antioquia, Chocó (La Teresita), Valle del Cauca (near Rio Dagua, Municipio Buenaventura)), Mexico to Venezuela. Refs.: Lee et al., 1969: 444; Fairchild, 1971: 20 (cat.); Moucha, 1976: 35 (cat.); Wilkerson, 1979: 148; Fairchild & Burger, 1994: 42 (cat.); Coscaron & Papavero, 2009: 38 (cat.). isthmiae Fairchild, 1941b: 642. niger Ricardo of Hine, 1925: 11 (Costa Rica), missident. flavipennis ssp. vallensis Wilkerson, 1979: 149. Type locality: Colombia, Valle, lower Anchicayá. HT F (FSCA). Distr.: Colombia (Valle del Cauca (Bajo Anchicayá, Rio Sabaletas, Bajo Calima)). Refs.: Fairchild & Burger, 1994: 42 (cat.); Coscaron & Papavero, 2009: 38 (cat.). laterina (Rondani), 1850: 370 (Pangonia). Type locality: Ecuador, Rio Napo. NT F (FSCA). Distr.: Colombia (Putumayo [no sex stated] lost), Ecuador. Refs.: Fairchild & Burger, 1994: 43 (cat.); Coscaron & Papavero, 2009: 39 (cat.). ochrapogon Wilkerson, 1979: 150. Type locality: Colombia, Putumayo, Puerto Pepino, 500m. HT F (FSCA). Distr.: Colombia (Cauca (Cerro Munchique), Caquetá (Florencia), Putumayo (Puerto Pepino)), Argentina, Bolivia, Ecuador, Peru. Refs.: Fairchild & Burger, 1994: 45 (cat.); Coscaron & Papavero, 2009: 41 (cat.). pubescens (Lutz), 1909a: 644 (Erephopsis). Type locality: Brazil, São Paulo, Jacutinga. LT F (IOC). Distr.: Colombia (Putumayo (Puerto Asis)), Brazil. Refs.: Fairchild & Burger, 1994: 46 (cat.); Bequaert & RenjifoSalcedo, 1946; Coscaron & Papavero, 2009: 42 (cat.). rhinophora (Bellardi), 1859: 46, (Pangonia). Type locality: “Mexico”. HT F (IEAUT). Distr.: Colombia (Boyacá (Muzo), Cauca (Cerro Minchique), Chocó, Magdalena, Meta (Villavicencio), Valle del Cauca (Lago Calima, Playa Rica, Peñas Blancas)), Mexico to Venezuela, Peru. Refs.: Kröber, 1934: 250 (cat.); Bequaert & Renjifo-Salcedo, 1946; Fairchild, 1971: 23 (cat.); Moucha, 1976: 37 (cat.); Wilkerson, 1979: 152; Fairchild & Burger, 1994: 46 (cat.); Coscaron & Papavero, 2009: 42 (cat.). pyrausta Osten Sacken, 1886: 43 (Pangonia). auricauda Enderlein, 1925: 276 (Melpia). schildi (Hine), 1925: 11 (Erephopsis). Type locality: Costa Rica, La Suiza de Turrialba. HT F (USNM). Distr.: Colombia (Antioquia (Providencia)), Brazil, Costa Rica to Colombia, French Guiana. Refs.: Kröber, 1934: 250 (cat.); Fairchild, 1971: 23 (cat.); Moucha, 1976: 38 (cat.); Wilkerson, 1979: 153; Fairchild & Burger, 1994: 46 (cat.); Coscaron & Papavero, 2009: 43 (cat.). nigricans Kröber, 1930b: 181 (Melpia; preocc. Lutz, 1909). nigricorpus Kröber, 1934: 291 (cat.; footnote; nom. nov. for nigricans Kröber, 1930). sulfurea Wilkerson, 1979: 155. Type locality: Colombia, Valle, 8km below dam at lago Calima, 1250m. HT F (FSCA). Distr.: Colombia (Cauca (La Cumbre), Caquetá (Florencia), Magdalena (NW. Sierra Nevada de Santa Marta, Vista Nieve, Cuchilla San Lorenzo), Valle del Cauca (Lago Calima)). Refs.: Fairchild & Burger, 1994: 47 (cat.); Coscaron & Papavero, 2009: 43 (cat.). Genus Pityocera Giglio-Tos, 1896 Subgenus Elaphella Bezzi Elaphella Bezzi, 1913: 56 (nom. nov. for Dicrania Macquart 1834). Type species, Pangonia cervus Wiedemann, 1828 (aut.). Refs.: Coscaron & Papavero, 2009: 45 (cat.). Allodicrania Enderlein, 1913: 253 (nom. nov. for Dicrania Macquart, 1834). Stichocera Hine, 1920a: 185 (nom. nov. for Dicrania Macquart, 1834). Stichochera Surcouf, 1921: 99, error. 256 · Zootaxa 4122 (1) © 2016 Magnolia Press E & MIRANDA-ESQUIVEL cervus (Wiedemann), 1828: 194 (Pangonia). Type locality: Brazil, Pará. HT F (MNHUB). Distr.: Colombia (Meta (Villavicencio, Bosque Oca; Lago El Dorado, Rio Vaupés)), Bolivia, N. Brazil, Guyana, Peru, Surinam. Refs.: Bequaert & Renjifo-Salcedo, 1946: 60 (Elaphella); Fairchild, 1971: 28 (cat.); Moucha, 1976: 41 (cat.); Wilkerson, 1979: 178; Fairchild & Burger, 1994: 52 (cat.); Coscaron & Papavero, 2009: 45 (cat.); Krolow et al., 2015: 304 (rev.). comprehensa Walker, 1850a: 11 (Pangonia). ramicornis Ricardo, 1900a: 100, nomen nudum. Subgenus Pityocera Giglio-Tos festai Giglio-Tos, 1896: 3, (as festae, error). Type locality: Panama, Darien, Rio Lara, Laguna Pita. HT F (IEAUT). Distr.: Colombia (Chocó (La Teresita), Cauca (Guapi)), Ecuador, Panama. Refs.: Bequaert & Renjifo-Salcedo, 1946: 60; Fairchild, 1971: 27 (cat.); Moucha, 1976: 41 (cat.); Wilkerson, 1979: 179 (as festae); Fairchild & Burger, 1994: 51 (cat.); Coscaron & Papavero, 2009: 46 (cat.); Krolow et al., 2015: 308 (rev.). festae, error (named after Enrico Festa (cf. Papavero, 1973: 352-353)). Genus Scione Walker Pangonia, subg. Scione Walker, 1850: 10. Type species, Pangonia incompleta Macquart, 1846 (mon.). Refs.: Coscaron & Papavero, 2009: 51 (cat.). Diclisa Schiner, 1867: 305. Type species, Pangonia incompleta Macquart, 1846 (orig. des.). Rhinotriclista Enderlein, 1922: 338 (in key). Type species, Diclisa maculipennis Schiner, 1868 (orig. des.). albifasciata (Macquart), 1846: 156 (1846: 28) (Pangonia). Type locality: “Colombia” (as Nouvelle Grenade). ST F (BMNH). Distr.: Colombia (Cauca (Cerro de Munchique), Chocó (La Unión), Huila (Finca Meremberg, near Leticia), ?Magdalena (Vista Nieve, W. San Lorenzo, NW. Sierra Nevada de Santamarta), ?Meta (Villavicencio), ?Putumayo)), Ecuador. Refs.: Kröber, 1934: 232 (as albofasciata); Bequaert & Renjifo-Salcedo, 1946: 62; Fairchild, 1971: 25 (cat.); Moucha, 1976: 39 (cat.); Wilkerson, 1979: 159; Fairchild & Burger, 1994: 49 (cat.); Coscaron & Papavero, 2009: 51 (cat.). albofasciata Kröber, 1934: 232 (cat.) error. aurulans (Wiedemann), 1830: 620 (Pangonia). Type locality: Mexico, Oaxaca. LT F (MNHUB). Distr.: Colombia (Magdalena (Vista Nieve, San Lorenzo; Cerro Quemado)), Mexico to Costa Rica. Refs.: Kröber, 1934: 233 (cat.); Bequaert & Renjifo-Salcedo, 1946: 61; Fairchild & Burger, 1994: 49 (cat.); Coscaron & Papavero, 2009: 51 (cat.). rostrifera Bellardi, 1859: 47 (Pangonia). misera Osten Sacken, 1886: 47 (Diclisa). aurulans ssp. lurida Enderlein, 1925: 301. Type locality: “Colombia”. LT F (MNHUB). Distr.: Colombia, Venezuela. Refs.: Fairchild, 1971: 25 (cat.); Moucha, 1976: 39 (cat.); Fairchild & Burger, 1994: 49 (cat.); Coscaron & Papavero, 2009: 52 (cat.). brevibeccus Wilkerson, 1979: 161. Type locality: Colombia, Valle, lago Calima, 8km below dam. HT F, 6 F PT (FSCA). Distr.: Colombia (Valle del Cauca (Lago Calima)). Refs.: Fairchild & Burger, 1994: 49 (cat.); Coscaron & Papavero, 2009: 52 (cat.). cuprea Wilkerson, 1979: 163 (as cupreus). Type locality: Colombia, Cauca, Cerro de Munchique, 2450m. HT F (FSCA). Distr.: Colombia (Cauca (Cerro de Munchique, 2450m)). Refs.: Fairchild & Burger, 1994: 49 (cat.; as cupreus); Coscaron & Papavero, 2009: 52 (cat.). distincta (Schiner), 1868: 102 (Diclisa). Type locality: “South America” (Venezuela). HT F (NMW). Distr.: Colombia (Cundinamarca (Medina, upper Rio Guacavía), Putumayo (upper Rio Putumayo)), Venezuela. Refs.: Bequaert & Renjifo-Salcedo, 1946: 62; Fairchild, 1971: 26 (cat.); Moucha, 1976: 40 (cat.); Fairchild & Burger, 1994: 49 (cat.); Coscaron & Papavero, 2009: 52 (cat.). CATALOGUE OF TABANIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 257 equivexans Wilkerson, 1979: 164. Type locality: Colombia, Valle del Cauca, Peñas Blancas, 1750m, 10km W. of Cali. HT F ca. 200 F PT (FSCA). Distr.: Colombia (Cauca (Cerro de Munchique, La Peña del Perro, Cerro Torres), Valle del Cauca (Bajo Calima; Buenaventura; Peñas Blancas; W. Queremal)), Ecuador. Refs.: Fairchild & Burger, 1994: 50 (cat.); Coscaron & Papavero, 2009: 52 (cat.). flavescens (Enderlein), 1930: 65 (Rhinotriclista). Type locality: ‘Ecuador’. HT F (MNHUB). Distr.: Colombia (Cauca (Cerro de Munchique, El Asomadero), Valle del Cauca (Lago Calima, Rio Bravo, Peñas Blancas)), Ecuador. Refs.: Kröber, 1934: 233 (cat.); Wilkerson, 1979: 166; Fairchild & Burger, 1994: 50 (cat.); Coscaron & Papavero, 2009: 52 (cat.). fulvosericea (Kröber), 1931b: 26, (Fidena). Type locality: Panama, Darien (as Colombia). HT F (BMNH). Distr.: Costa Rica to Colombia (W. of Andes). Refs.: Kröber, 1934: 247 (cat.); Bequaert & Renjifo-Salcedo, 1946: 61; Fairchild, 1971: 27 (cat.; as syn. of rufescens Ricardo, 1900); Moucha, 1976: 40 (cat.; as syn. of rufescens Ricardo, 1900); Wilkerson, 1979: 173 (as syn. of rufescens Ricardo, 1900); Fairchild & Burger, 1994: 50 (cat.); Coscaron & Papavero, 2009: 53 (cat.). aureopygia Fairchild, 1942d: 184. incompleta (Macquart), 1846: 155 (1845: 27), (Pangonia). Type locality: “Colombia”. ST F [Lille], ? F (MNHNP). Distr.: Colombia. Refs.: Kröber, 1934: 234 (cat.); Bequaert & Renjifo-Salcedo, 1946: 64; Fairchild, 1971: 26 (cat.); Moucha, 1976: 40 (cat.); Fairchild & Burger, 1994: 50 (cat.); Coscaron & Papavero, 2009: 53 (cat.). maculipennis (Schiner), 1868: 102 (Diclisa). Type locality: “South America” (Venezuela). ST F (NMW). Distr.: Colombia (Cauca (Munchique, La Peña del Perro, Cerro Torres), Chocó (La Unión), ?Cundinamarca (Bogotá), Meta (Villavicencio, Restrepo), Valle del Cauca (Lago Calima; Rio Bravo-Playa-Rica; Peñas Blancas)), Costa Rica to Venezuela, Ecuador. Refs.: Kröber, 1934: 234 (cat.); Bequaert & Renjifo-Salcedo, 1946: 62; Fairchild, 1971: 26 (cat.); Moucha, 1976: 40 (cat.); Wilkerson, 1979: 168; Fairchild & Burger, 1994: 50 (cat.); Coscaron & Papavero, 2009: 53 (cat.). aurea (Macquart), 1847: 29 (1847: 13). minor (Macquart), 1847: 29 (1847: 13) (Pangonia). Type locality: “?America”. HT F (BMNH). Distr.: Colombia (Cundinamarca (Bosque Calderón, Bogotá; Monserrate; Arrayán; Bogotá), Meta, Putumayo (upper Rio Putumayo)), Venezuela. Refs.: Bequaert & Renjifo-Salcedo, 1946: 63; Fairchild, 1971: 27 (cat.); Moucha, 1976: 40 (cat.); Fairchild & Burger, 1994: 50 (cat.); Coscaron & Papavero, 2009: 53 (cat.). incompleta Macquart of Schiner, 1868: 101 (Diclisa), missident. minuta Szilády, 1926: 28. Type locality: Colombia, Sierra San Lorenzo. Distr.: Colombia (Magdalena (Sierra San Lorenzo)). Refs.: Kröber, 1934: 234 (cat.); Bequaert & Renjifo-Salcedo, 1946: 64; Fairchild, 1971: 27 (cat.); Moucha, 1976: 40 (cat.); Fairchild & Burger, 1994: 51 (cat.); Coscaron & Papavero, 2009: 54 (cat.). nigripes (Kröber), 1930k: 152, (Rhinotriclista). Type locality: “New Grenada” (Colombia). Distr.: unknown. HT F (MLUH). Refs.: Kröber, 1934: 234 (cat.); Bequaert & Renjifo-Salcedo, 1946: 63; Fairchild, 1971: 27 (cat.); Moucha, 1976: 40 (cat.); Fairchild & Burger, 1994: 51 (cat.); Coscaron & Papavero, 2009: 54 (cat.). obscurefemorata Kröber, 1930a: 172. Type locality: “Colombia, Peru”. ST F (MNHUB). Distr.: Colombia (Antioquia (Sonsón), Cauca (Cerro de Munchique)), Ecuador, Peru. Refs.: Kröber, 1934: 234 (cat.); Bequaert & Renjifo-Salcedo, 1946: 63; Fairchild, 1971: 27 (cat.); Moucha, 1976: 40 (cat.); Wilkerson, 1979: 170; Fairchild & Burger, 1994: 51 (cat.); Coscaron & Papavero, 2009: 54 (cat.). punctata Szilády, 1926: 27, 29. Type locality: Colombia, Sierra San Lorenzo. Distr.: Colombia (Magdalena (Sierra San Lorenzo; Vista Nieve, San Lorenzo Mountains)), ?Ecuador. Refs.: Kröber, 1934: 234 (cat.); Bequaert & Renjifo-Salcedo, 1946: 63; Fairchild, 1971: 27 (cat.); Moucha, 1976: 40 (cat.); Fairchild & Burger, 1994: 51 (cat.); Coscaron & Papavero, 2009: 54 (cat.). 258 · Zootaxa 4122 (1) © 2016 Magnolia Press E & MIRANDA-ESQUIVEL punstata Fairchild & León, 1986: 105. rhinothrix Wilkerson, 1979: 172. Type locality: Colombia, Valle, Peñas Blancas, 1750m, 10km W. of Cali. HT F, 2F PT (FSCA). Distr.: Colombia (Valle del Cauca (Peñas Blancas)). Refs.: Fairchild & Burger, 1994: 51 (cat.); Coscaron & Papavero, 2009: 54 (cat.). rufescens (Ricardo), 1900b: 294 (Erephrosis). Type locality: Peru, Callanga. PT F (BMNH). Distr.: Colombia (?Chocò or Antioquia), Costa Rica to Bolivia. Refs.: Kröber, 1934: 234 (cat.); Bequaert & Renjifo-Salcedo, 1946: 63; Fairchild, 1971: 27 (cat.); Moucha, 1976: 40 (cat.); Wilkerson, 1979: 173; Fairchild & Burger, 1994: 51 (cat.); Coscaron & Papavero, 2009: 54 (cat.). generosa Enderlein, 1925: 302 (Rhinotriclista; Enderlein’s type) rufipes (Kröber), 1930k: 147 (Rhinotriclista). Type locality: “Venezuela”. Distr.: Colombia (Meta (Restrepo)), Venezuela. Refs.: Bequaert & Renjifo-Salcedo, 1946: 63; Fairchild, 1971: 27 (cat.); Moucha, 1976: 40 (cat.); Fairchild & Burger, 1994: 51 (cat.); Coscaron & Papavero, 2009: 54 (cat.). serrata Wilkerson, 1979: 174 (as serratus, error). Type locality: Colombia, Valle, “Rio Bravo-Playa Rica”, 15km below dam at lago Calima. HT F (FSCA). Distr.: Colombia (Valle del Cauca (Rio Bravo-Playa Rica; Bajo Anchicayá)). Refs.: Fairchild & Burger, 1994: 51 (cat.; as serratus); Coscaron & Papavero, 2009: 54 (cat.). youngi Wilkerson, 1979: 176. Type locality: Colombia, Valle, Peñas Blancas, 10km W. of Cali. HT F, PT F (FSCA). Distr.: Colombia (Valle del Cauca). Refs.: Fairchild & Burger, 1994: 51 (cat.); Coscaron & Papavero, 2009: 54 (cat.). Subfamily Tabaninae Latreille, 1802 Tribe Diachlorini Lutz, 1909 Genus Acanthocera Macquart, 1834 Subgenus Acanthocera Macquart, 1834 marginalis Walker, 1854: 268. Type locality: Brazil, Pará. HT F (BMNH). Distr.: , Brazil, Colombia, Ecuador, Guyana, Peru, Surinam, Trinidad. Refs.: Fairchild, 1971: 59 (cat.); Moucha, 1976: 196 (cat.); Fairchild & Burger, 1994: 93 (cat.); Henriques & Rafael, 1995: 420 (rev.); Coscaron & Papavero, 2009: 56 (cat.). formosa Kröber, 1930a: 79 Subgenus Nothocanthocera Fairchild Dichelacera, subg. Nothocanthocera Fairchild, 1969 209. Type species, Acanthocera tenuicornis Lutz, 1915 (orig. des.). Refs.: Wilkerson, 1981 (key). Henriques & Rafael, 1995: 421 (rev.). albomarginata (Kröber), 1930a: 76. 17-18 (Spheciogaster). Type locality: Colombia, Chocó, Condoto. HT F (BMNH). Distr.: Colombia (Chocó (La Teresita; Condoto)). Refs.: Kröber, 1934: 253 (cat.; Spheciogaster albomarginatus); Bequaert & Reinjifo-Salcedo 1946: 83 (Acanthocera); Fairchild, 1971: 65 (cat.); Moucha, 1976: 199 (cat.); Wilkerson, 1979: 260; Fairchild & Burger, 1994: 100 (cat.); Henriques & Rafael, 1995: 422 (rev.); Coscaron & Papavero, 2009: 87 (cat.). trigonifera (Schiner), 1868: 95 (Acanthocera). Type locality: “South America” (Venezuela). ST F (NMW). Distr.: Colombia (Cundinamarca (Bogotá, reported by Krober, 1928), Magdalena (Santa Marta, F.L. Gallego), Rio Magdalena, Pehlke col)), Venezuela. The occurrence of this species near Bogota needs to be confirmed (Bequaert & Reinjifo-Salcedo, 1946). Refs.: Bequaert & Reinjifo-Salcedo, 1946: 83 (Acanthocera); Fairchild, 1971: 66 CATALOGUE OF TABANIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 259 (cat.); Moucha, 1976: 199 (cat.); Fairchild & Burger, 1994: 101 (cat.); Henriques & Rafael, 1995: 427 (rev.); Coscaron & Papavero, 2009: 88 (cat.). Genus Bolbodimyia Bigot Bolbodimyia Bigot, 1892b: 162. Type species, Bolbodimyia bicolor Bigot, 1892 (mon.). Papavero, 2009: 59 (cat.). Snowielius Hine, 1904: 230. Type species, Snowielius atratus Hine 1904 (mon.). Refs.: Coscaron & bicolor Bigot, 1892b: 162. Type locality: Venezuela, Mérida. HT F (BMNH). Distr.: Colombia (Boyacá (Muzo), Chocó (between Rio San Juan and Rio Baudó), Meta (Restrepo; Villavicencio; Retiro near Villavicencio), Valle del Cauca (Bajo Anchicayá, 41km and 21km N. Buenaventura; Cali; Rio Sabaletas; Peñas Blancas)), Bolivia, Guyana, Venezuela. Refs.: Bequaert & Renjifo-Salcedo, 1946: 64; Fairchild, 1971: 54 (cat.); Moucha, 1976: 192 (cat.); Wilkerson, 1979: 229; Fairchild & Burger, 1994: 86 (cat.); Coscaron & Papavero, 2009: 60 (cat.). celeroides Stone, 1954: 251. Type locality: Colombia, Meta, Restrepo, 500m. HT F (MCZ). Distr.: Colombia (Meta, Valle del Cauca (Lago Calima, Peñas Blancas)), Ecuador, Peru. Refs.: Fairchild, 1971: 54 (cat.); Moucha, 1976: 192 (cat.); Wilkerson, 1979: 231; Fairchild & Burger, 1994: 86 (cat.); Coscaron & Papavero, 2009: 60 (cat.). erythrocephala (Bigot), 1892a: 668 (Atylotus). Type locality: “Panama”. HT F (BMNH). Distr.: Colombia (Valle del Cauca (41km and 20km N. Buenaventura; Rio Sabaletas)), Costa Rica, Panama. Refs.: Fairchild, 1971: 54 (cat.); Moucha, 1976: 192 (cat.); Wilkerson, 1979: 231; Fairchild & Burger, 1994: 86 (cat.); Coscaron & Papavero, 2009: 60 (cat.). galindoi Fairchild, 1964a: 172. Type locality: Panama, Darien, Tacarcuna Yellow Fever Station, 665m. HT F (MCZ). Distr.: Colombia (Antioquia (Rio Anorí Valley)), Costa Rica to Colombia. Refs.: Wilkerson, 1979: 232; Fairchild & Burger, 1994: 86 (cat.); Coscaron & Papavero, 2009: 60 (cat.). nigra Stone, 1934: 190. Type locality: Ecuador, Guayaquil. HT F (USNM). Distr.: Colombia (Meta (Villavicencio), Valle del Cauca (Peñas Blancas; 41km E. and 20km N. Buenaventura; Lago Calima)), Ecuador, Peru. Refs.: Kröber, 1934: 258 (cat.); Fairchild, 1971: 54 (cat.); Moucha, 1976: 192 (cat.); Wilkerson, 1979: 234; Fairchild & Burger, 1994: 86 (cat.); Coscaron & Papavero, 2009: 60 (cat.). philipi Stone, 1954: 253. Type locality: Guatemala, Acatenango. HT F (USNM). Distr.: Colombia (Antioquia, Boyacá), Costa Rica, El Salvador, Guatemala, Panama. Refs.: Wilkerson, 1979: 235; Fairchild & Burger, 1994: 86 (cat.); Coscaron & Papavero, 2009: 60 (cat.). Genus Catachlorops Lutz, 1913 Subgenus Amphichlorops Lutz, 1913 Amphichlorops Lutz, 1909b: 29, nomun nudum. Amphichlorops Lutz, 1913a: 486, 487 (as genus, defined in key) (1914: 166, 167 (sep., pp. 6, 7). Type species, Tabanus flavus Wiedemann, 1828 (subs. des., Bequaert 1924: 27). Refs.: Fairchild, 1951a: 120, 1961: 237. Coscaron & Papavero, 2009: 61 (cat.). Stypochela Enderlein, 1922: 345 (in key). bogotanus (Enderlein), 1925: 386 (Stypochela). Type locality: Colombia, Bogotá, Cartago and Las Pavas. LT F (MNHUB). Distr.: Colombia (Valle del Cauca (Cartago; Las Pavas) the locality Bogotá is probably erroneus). Refs.: Kröber, 1934: 264 (cat.; Stypochela as bogotana); Fairchild, 1971: 69 (cat.; Stypochela); Moucha, 1976: 204 (cat.); Bequaert & Renjifo-Salcedo, 1946: 71 (Dasychela (Stypochela)); Fairchild & Burger, 1994: 104 (cat.); Coscaron & Papavero, 2009: 61 (cat.). 260 · Zootaxa 4122 (1) © 2016 Magnolia Press E & MIRANDA-ESQUIVEL bogotana Enderlein, 1922: 345 (Stypochela), nomen nudum. vespertinus (Bequaert & Renjifo-Salcedo), 1946: 67 (Dichelacera (Psalidia); nom. nov. for elongatus Macquart, 1846) (as vespertina). Type locality: “Nouvelle Grenade”. HT F, PT F 1 (LACM). Distr.: Colombia (Chocó (Rio Nimiquía), Meta (Villavicencio; Restrepo), Valle del Cauca (Lago Calima; La Carmelita; Rio Frío), Ecuador, Peru, Bolivia. Refs.: Fairchild, 1971: 69 (cat.); Moucha, 1976: 204 (cat.); Wilkerson, 1979: 268; Fairchild & Burger, 1994: 104 (cat.); Coscaron & Papavero, 2009: 61 (cat.). elongatus Macquart, 1846: 166 (1846: 38), (Tabanus; preocc. Wiedemann, 1828). Subgenus Catachlorops Lutz, 1913 alphus Wilkerson, 1979: 272. Type locality: Colombia, Valle, 3.6km W. Queremal, 1130m. HT F (FSCA). Distr.: Colombia (Valle del Cauca (W. Queremal)). Refs.: Fairchild & Burger, 1994: 101 (cat.); Coscaron & Papavero, 2009: 61 (cat.). fumipennis Kröber, 1931c: 347. Type locality: Brazil, Mato Grosso. HT F (NMW). Distr.: Brazil, Colombia. Refs.: Fairchild, 1971: 67 (cat.); Moucha, 1976: 201 (cat.); Fairchild & Burger, 1994: 102 (cat.); Coscaron & Papavero, 2009: 62 (cat.). nigripalpis (Macquart), 1846: 168 (1846: 40) (Tabanus). Type- locality: ‘Nouvelle Grenade’ (Colombia). HT F (BMNH). Distr.: Colombia (Magdalena (Santa Marta), Santander)), Venezuela. Refs.: Kröber, 1934: 273 (cat.), Bequaert & Renjifo-Salcedo, 1946: 67 (Dichelacera (?Catachlorops)); Fairchild, 1971: 68 (cat.); Moucha, 1976: 201 (cat.); Fairchild & Burger, 1994: 103 (cat.); Coscaron & Papavero, 2009: 63 (cat.). calopterus Schiner, 1868: 91 (Tabanus). rufescens (Fabricius), 1805: 100 (Tabanus). Type locality: “South America. Smidt” (i.e., Guyana, between rivers Demerara and Essequibo; cf. Papavero, 1971: 21). HT F (ZMKU). Distr.: Colombia (Magdalena (Vista Nieve, Sierra Nevada de Santa Marta)), Brazil, French Guiana, Guyana. Refs.: Bequaert & Renjifo-Salcedo, 1946: 67 (Dichelacera (Catachlorops)); Fairchild & Burger, 1994: 103 (cat.); Coscaron & Papavero, 2009: 63 (cat.). ruficeps Kertész, 1900: 68 (Tabanus), error. scutellatus Kröber, 1931c: 348. Subgenus Psalidia Enderlein, 1922 fulmineus (Hine), 1920a: 186 (Tabanus). Type locality: Panama, Canal Zone, Gatún. Distr.: Colombia (Antioquia (Providencia), Boyacá (Muzo), Cauca (Rio Guapi), Chocó (La Teresita, upper Curiche, Rio Curiche), Valle del Cauca (Rio Raposo; Bajo Calima, 10km E. Buenaventura)), Ecuador, Honduras to Panama. Refs.: Bequaert & Renjifo-Salcedo, 1946: 68 (Dichelacera (Psalidia)) (as fulminea); Lee et al., 1969: 448 (as var. ocellata); Wilkerson, 1979: 270; Fairchild & Burger, 1994: 107 (cat.); Coscaron & Papavero, 2009: 65 (cat.). furcata Bigot, 1892a: 631 (Bellardia; preocc. Wiedemann, 1828). festivus Hine 1920a: 187 (Tabanus; preocc. Wiedemann, 1828). ocellata Enderlein 1925: 393 (Psalidia). medemi Philip, 1969b: 149. Type locality: Colombia, Amazonas, lower Rio Caquetá, La Pedrera. HT F (CAS), PT F 2 (FSCA), PT F 2 (SMNS). Distr.: Colombia (Amazonas), Brazil. Refs.: Fairchild & Burger, 1994: 107 (cat.); Coscaron & Papavero, 2009: 65 (cat.). Subgenus Psarochlorops Fairchild, 1969 Catachlorops, subg. Psarochlorops Fairchild, 1969: 211. Type species, Dichelacera testacea Macquart, 1846 (orig. des.). auripilis (Philip), 1960a: 86 (Dichelacera (Catachlorops)). Type locality: Colombia, Nariño, 18mi. SW. of Mocoa, CATALOGUE OF TABANIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 261 1910m. HT F (CAS). Distr.: Colombia (Nariño). Refs.: Fairchild, 1971: 70 (cat.); Moucha, 1976: 203 (cat.); Fairchild & Burger, 1994: 106 (cat.); Coscaron & Papavero, 2009: 66 (cat.). difficilis (Kröber), 1931g: 93 (Dichelacera). Type locality: Brazil, Amazonas, Tefé, and Guyana, Demerara. ST F (IZPAN). Distr.: Brazil, Colombia, Ecuador, Guyana. Refs.: Fairchild, 1971: 70 (cat.); Moucha, 1976: 203 (cat.); Fairchild & Burger, 1994: 106 (cat.); Coscaron & Papavero, 2009: 66 (cat.). quadrimaculatus (Macquart), 1846: 167 (1846: 39) (Tabanus). Type locality: ‘Nouvelle Grenade’ (Colombia). HT F (BMNH). Distr.: Colombia, Venezuela. Refs.: Kröber, 1934: 271 (cat. Amphichlorops); Bequaert & RenjifoSalcedo, 1946: 67 (Dichelacera (Catachlorops)); Fairchild, 1971: 70 (cat.); Moucha, 1976: 203 (cat.); Fairchild & Burger, 1994: 106 (cat.); Coscaron & Papavero, 2009: 66 (cat.). poecilopterus Schiner, 1868: 91 (Tabanus). siculus Wilkerson, 1979: 274. Type locality: Colombia, Valle, lago Calima, 8km below dam, 1250m. HT F (FSCA). Distr.: Colombia (Valle del Cauca (Lago calima; 3.6km W. Queremal; Rio Dagua)). Refs.: Fairchild & Burger, 1994: 106 (cat.); Coscaron & Papavero, 2009: 66 (cat.). testaceus (Macquart), 1846: 157 (1846: 29), (Dichelacera). Type locality: “Colombia”. HT F (BMNH). Distr.: Colombia (Meta (Restrepo; Villavicencio), Vaupés (between San José del Guaviare and Calamar)), Brazil, Guyana, Venezuela. Refs.: Kröber, 1934: 267; Bequaert & Renjifo-Salcedo, 1946: 67 (Dichelacera (Catachlorops)) (as testacea); Fairchild, 1971: 70 (cat.); Moucha, 1976: 203 (cat.); Fairchild & Burger, 1994: 106 (cat.); Coscaron & Papavero, 2009: 66 (cat.). repanda Walker, 1848: 190 (Dichelacera). Genus Chlorotabanus Lutz Chlorotabanus Lutz, 1913a: 487 (sep., p. 6) (1914: 167 (sep., p. 7)). Type species, Tabanus mexicanus Linnaeus, 1758 (mon.). Refs.: Fairchild, 1951a: 121, 1961: 238. Coscaron & Papavero, 2009: 67 (cat.); Krolow & Henriques, 2010: 2 (rev.). fairchildi Wilkerson, 1979: 237. 30A-C. Type locality: Colombia, Valle del Cauca, Rio Sabaletas, 4km E. of Sabaletas. HT F, PT F 4 (FSCA). Distr.: Colombia (Cauca (Guapi), Chocó (Teresita, quebrada docordo between Cucurrupi & Noanama; Rio San Juan), Valle del Cauca (Rio Sabaletas; Bajo Calima; La Bocana; Buenaventura, Rio Dagua)). Refs.: Fairchild & Burger, 1994: 87 (cat.); Coscaron & Papavero, 2009: 67 (cat); Krolow & Henriques, 2010: 9 (rev.). inanis (Fabricius), 1787: 356 (Tabanus). Type locality: “Cayenne. Rohr” [cf. Papavero, 1971: 20]. NT F (ZMKU). Distr.: Colombia (Amazonas (Leticia), Antioquia (Puerto Berrio), Cauca (Quebrada Huanqui), Meta (Villavicencio, Carimagua; Caibe; Restrepo), Putumayo (Mocoa; Rio Putumayo between Itiquilla and Puerto Arana; La Tagua; santa Rosa)), S. Mexico S. to S. Brazil. Refs.: Kröber, 1934: 296 (cat.; Tabanus (Chlorotabanus)); Fairchild, 1971: 55 (cat.); Moucha, 1976: 193 (cat.); Bequaert & Renjifo-Salcedo, 1946: 73; Wilkerson, 1979: 239; Fairchild & Burger, 1994: 87 (cat.); Coscaron & Papavero, 2009: 67 (cat.); Krolow & Henriques, 2010: 14 (rev.). inanis Kröber, 1930: 15–16 (key); 1934: 296 (Tabanus (Chlorotabanus)). inanis Philip & Fairchild, 1956: 316–317 (Chlorotabanus (Chlorotabanus)). inanis Coscarón & Papavero, 2009b: 3 (Chlorotabans) (manual), error. ochroleucus Meigen, 1804: 172 (Tabanus). sulphureus Palisot de Beauvois, 1819: 222 (Tabanus). sulphureus Macquart, 1847: 35 (Tabanus; preocc. Palisot de Brauvois, 1819). inconspicuus Walker, 1848: 171 (Tabanus). viridiflavus Walker, 1850b: 1xvi (Tabanus) Material examined. COLOMBIA: Santander: Cimitarra, Puerto Olaya, Central Termocentro-Isagen, 110m, 1 male, 12.vi.2012, Castaño-Rivera leg., trap light (CEUA). 262 · Zootaxa 4122 (1) © 2016 Magnolia Press E & MIRANDA-ESQUIVEL leucochlorus Fairchild, 1961c: 439, fig. 2. Type-locality: Brazil, Amapá, Serra do Navio. HT F (MZUSP). Distr.: Colombia (Amazonas (30 km aval de La Chorrera, Rio Igaraparana), N. Brasil, French Guiana, Guyana, Surinam, E. Peru, Venezuela, Refs.: Krolow & Henriques, 2010: 18 (ver.). leuconotus Krolow & Henriques, 2010: 21, figs. 8a–i. Type locality: Brazil, Amazonas, Manaus. HT F (INPA), PT 35 F (BMNH, CZMA, INPA, MNHN, MPEG). Distr.: Colombia (Putumayo (Amazonas, Puerto Nariño), Guyana, French Guiana, Brazil, Peru. Ref.: Henriques et al., 2012: 279. mexicanus (Linnaeus), 1758: 602 (Tabanus). Type locality: “South America”. HT F (NRS). Distr.: Colombia (Antioquia (Murindó; Providencia), Cauca (Rio Micay; Guapi), Chocó (along the Atrato River; Arquía; Andagoya), Valle del Cauca (Buenaventura; Rio Raposo; Rio Sabaletas)), S. Mexico to Ecuador (Esmeraldas), N. Brazil, Trinidad. Refs.: Kröber, 1934: 297 (cat.; Tabanus (Chlorotabanus)); Bequaert & Renjifo-Salcedo, 1946: 73; Lee et al., 1969: 450; Fairchild, 1971: 55 (cat.); Moucha, 1976: 193 (cat.); Wilkerson, 1979: 240; Fairchild & Burger, 1994: 87 (cat.); Coscaron & Papavero, 2009: 68 (cat.); Krolow & Henriques, 2010: 24 (rev.). mexicanus, Kröber, 1930: 15, 18 (key and diagnosis) (Tabanus); 1934: 297 (catalog); Bequaert, 1940: 449 (citation); 1944: 17 (citation). mexicanus Philip & Fairchild, 1956: 315–316 (Chlorotabanus (Chlorotabanus). olivaceus De Geer, 1776: 229, pl. 30, fig. 6 (Tabanus). punctatus Fabricius, 1787: 355 (Tabanus). tetrapunctus Thunberg, 1827a: 57 (Tabanus). tetrapunctatus Philip, 1967: 1235 (Tabanus), error. Genus Cryptotylus Lutz Cryptotylus Lutz, 1913a: 487 (sep., p. 6) (1914a: 166 (sep. p. 6)) (as Chryptotylus, error). Type species, Tabanus unicolor Wiedemann, 1828 (mon.). Refs.: Coscaron & Papavero, 2009: 68 (cat.). Ommalia Enderlein, 1923: 545. Type-species, Ommalia viridis Enderlein, 1925 (sub. des., Enderlein, 1925a: 388) = Cryptotylus unicolor (Wiedemann, 1828). chloroticus (Philip & Fairchild), 1956: 320, (Chlorotabanus (Cryptotylus). Type locality: Panama, Panama Prov., Moja Pollo. HT F, PT F (MCZ), PT F 7, M 2 (FSCA), other PT (USNM, MCZ, NMW, BMNH, DECU, and CAS). Distr.: Colombia (Meta (Villavicencio)), Panama, Trinidad. Refs.: Fairchild, 1971: 77 (cat.); Moucha, 1976: 206 (cat.); Wilkerson, 1979: 314; Fairchild & Burger, 1994: 115 (cat.); Coscaron & Papavero, 2009: 68 (cat.). limonus Townsend of Fairchild 1940a: 720, Bequaert 1944: 17, Stone 1944: 132 and Bequaert & Renjifo Salcedo, 1946: 73, misident. Refs.: Coscaron & Papavero, 2009: 68 (cat.). unicolor (Wiedemann), 1828: 141 (Tabanus). Type locality: “Brazil”. Distr.: Colombia (Antioquia (Caucasia), Chocó, Meta (Hacienda Yacuana near Villavicencio), Magdalena River), Panama to Brazil, Argentina, Paraguay. Refs.: Bequaert & Renjifo-Salcedo, 1946: 73; Fairchild, 1971: 78 (cat.); Moucha, 1976: 206 (cat.); Wilkerson, 1979: 315; Fairchild & Burger, 1994: 115 (cat.); Parra-Henao et al., 2008: 182; Coscaron & Papavero, 2009: 69 (cat.). ferrugineus Thunberg 1789: 91 (Tabanus; preocc. Ström, 1768). castaneus Macquart 1834: 198 (Tabanus). ochraceus Macquart 1838a: 153 (1838: 149) (Tabanus). sulphureus Macquart 1847: 35 (1847: 19) (Tabanus). princeps Brèthes 1910: 478 (Tabanus). viridis Enderlein 1925: 389 (Ommalia). Genus Dasybasis Macquart Dasybasis Macquart, 1847: 41 (1847: 25). Type species, Dasybasis appendiculata Macquart 1847 (mon.). Pseudoselasoma CATALOGUE OF TABANIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 263 Brèthes, 1910: 475. Type species, Pseudoselasoma opacum Brèthes, 1910 (subs. des., Enderlein, 1922: 346 (in key)). Refs.: Trojan, 1994: 161; Coscaron & Papavero, 2009: 69 (cat.). Archiplatius Enderlein, 1922: 348 (in key). Stypommia Enderlein, 1922: 346 (in key). Coscaronia Trojan, 1994: 126. n. syn by Coscaron & Papavero, 2009: 69 (cat.). montium (Surcouf), 1919: 229, (Tabanus). Type locality: Ecuador, Casitagua. HT F (MNHNP). Distr.: Colombia (Cauca (Parque de Puracé; Lago San Rafael; Cerro de Munchique), Cundinamarca (Monserrate near Bogotá), Valle del Cauca (above Tenerife in Paramo)), Bolivia, Ecuador, Venezuela. Refs.: Bequaert & Renjifo-Salcedo, 1946: 76 (Agelanius colombianus); Wilkerson, 1979: 203; Fairchild & Burger, 1994: 66 (cat.); Coscaron & Papavero, 2009: 73 (cat.). columbianus Enderlein, 1925: 374 (Archiplatius). colombianus Fairchild & León, 1986: 115, error. schineri (Kröber), 1931h: 87. 18-19 (Tabanus (Poecilosoma)). Type locality: “Venezuela”. HT F (NMW). Distr.: Colombia (Cundinamarca; Huila (20km W. Leticia, 2600m)), Venezuela, Ecuador. Refs.: Fairchild, 1971: 41 (cat.); Moucha, 1976: 222 (cat.); Wilkerson, 1979: 205; Fairchild & Burger, 1994: 68 (cat.); Coscaron & Papavero, 2009: 75 (cat.). REVISAR HUILA histrio Wiedemann, of Schiner 1868: 87 (Tabanus), misident. osornoi Bequaert, 1946b: 86 (Agelanius). Genus Dasychela Enderlein, 1922 Subgenus Dasychela Enderlein, 1922 limbativena (Enderlein), 1925: 386 (Dicladocera). Type locality: Colombia, Bogotá, Guayabal. HT F (MNHUB). Distr.: Colombia (Cundinamarca (Guayabal)). Refs.: Kröber, 1934: 268 (cat.; Dicladocera (Dicladocera)); Bequaert & Renjifo-Salcedo, 1946: 70; Fairchild, 1971: 72 (cat.); Moucha, 1976: 207 (cat.); Fairchild & Burger, 1994: 108 (cat.); Coscaron & Papavero, 2009: 77 (cat.). limbativena Enderlein 1922: 345 (Dicladocera), nomen nudum. ocellus (Walker), 1848: 143 (Pangonia). Type locality: Ecuador, Quito. HT F (BMNH). Distr.: Colombia (Meta (Villavicencio), Valle del Cauca (Lago Calima)), Ecuador, Peru. Refs.: Fairchild, 1971: 72 (cat.); Moucha, 1976: 207 (cat.); Wilkerson, 1979: 276; Fairchild & Burger, 1994: 108 (cat.); Coscaron & Papavero, 2009: 77 (cat.). Genus Diachlorus Osten Sacken, 1876 Diabasis Macquart, 1834: 207 (preocc. Hoffmannsegg, 1819). Type species, Tabanus bicinctus Fabricius, 1805 (subs. des., Coquillett, 1910: 532). Refs.: Coscaron & Papavero, 2009: 78 (cat.). Diachlorus Osten Sacken, 1876: 475 (nom. nov. for Diabasis Macquart, 1834). Type species, Tabanus bicinctus Fabricius, 1805 (aut.). anduzei Stone, 1944: 132. Type locality: Venezuela, Bolívar, [Rio?] Surukum. HT F (USNM), PT F 1 (FSCA). Distr.: Colombia, Ecuador, Venezuela. Refs.: Fairchild & Burger, 1994: 82 (cat); Coscaron & Papavero, 2009: 78 (cat.). curvipes (Fabricius), 1805: 107 (Haematopota). Type locality: “South America. Smidt” (i.e., Guyana, between rivers Demerara and Essequibo; cf. Papavero, 1971: 21). HT F (ZMKU). Distr.: Colombia (Antioquia (Providencia), Boyacá (Muzo), Cauca (Guapi), Chocó, Valle del Cauca (Veneral, Rio Yurumanguí), ?Vaupes (Rio Apoporis, La Isla)), Bolivia, Brazil, Costa Rica, French Guiana, Panama to Surinam, Peru, Trinidad. Refs.: Kröber, 1934: 256 (cat.); Bequaert & Reinjifo-Salcedo, 1946: 83; Fairchild, 1971: 52 (cat.); Moucha, 1976: 209 (cat.); Wilkerson, 1979: 225; Fairchild & Burger, 1994: 83 (cat); Coscaron & Papavero, 2009: 79 (cat.). 264 · Zootaxa 4122 (1) © 2016 Magnolia Press E & MIRANDA-ESQUIVEL ataenia Macquart, 1838a: 156 (1838: 152) (Diabasis). varipes Walker, 1854: 289 (Chrysops; preocc. Rondani, 1848). terminalis Macquart, 1855: 56 (1855: 36) (Chrysops; preocc. Walker, 1848). fuscistigma Lutz, 1913b: 148, pl. 12. Type locality: Brazil, Amazonas, Rio Madeira. HT F (IOC). Distr.: Bolivia, Brazil, Colombia, Ecuador, Peru, Surinam. Refs.: Fairchild & Burger, 1994: 83 (cat.); Coscaron & Papavero, 2009: 80 (cat.). ochraceus Macquart of Kröber, 1928: 116, misident. angustifrons Kröber, 1930a: 74. jobbinsi Fairchild, 1942e: 296. Type locality: Panama, Darien, El Real. HT F (MCZ). Distr.: Colombia (Cauca (Guapi), Chocó (La Teresita), Valle del Cauca (Buenaventura; Rio Raposo; Ladrilleros, 30km N. Buenaventura)), Brazil, Costa Rica to Ecuador. Refs.: Lee et al., 1969: 446; Fairchild, 1971: 53 (cat.); Moucha, 1976: 209 (cat.); Wilkerson, 1979: 226; Fairchild & Burger, 1994: 84 (cat.); Coscaron & Papavero, 2009: 80 (cat.). leticia Wilkerson & Fairchild, 1982: 641. Type locality: Colombia, Amazonas, 17km N. of Leticia. HT F, PT l F (FSCA). Distr.: Colombia (Amazonas), Brazil, Peru. Refs.: Fairchild & Burger, 1994: 84 (cat.); Coscaron & Papavero, 2009: 80 (cat.). pechumani Fairchild, 1972: 223. Type locality: Peru, Cuzco, Quincemil, 2450m. HT F (DECU), PT F l (FSCA). Distr.: Brazil, Colombia, Peru. Refs.: Fairchild & Burger, 1994: 84 (cat.); Coscaron & Papavero, 2009: 81 (cat.). xynus Fairchild, 1972: 226. Type locality: Peru, Cuzco, Quincemil, 2450m. HT F (DECU), PT F 4 (FSCA). Distr.: Brazil, Colombia, Peru, Surinam. Refs.: Fairchild & Burger, 1994: 84 (cat.); Coscaron & Papavero, 2009: 81 (cat.). Genus Dichelacera Macquart, 1838 Subgenus Desmatochelacera Fairchild Dichelacera subg. Desmatochelacera Fairchild, 1969: 210. Type species, Dichelacera transposita Walker, 1854 (orig. des.). albitibialis Burger, 1999: 54. Type locality: Peru, Loreto, Boquerón, 550m. HT F (LACM), PT F 19 (FSCA, LACM). Distr.: Colombia (Rio Putumayo). Refs.: Coscaron & Papavero, 2009: 82 (cat.). transposita Walker, 1854: 151 (Dichelacera). Type locality: “West Coast of America” (Ecuador). HT F (BMNH). Distr.: Colombia (Antioquia (Rio Anori valley; Providencia), Cauca (Rio Micay), Chocó (between Rio San Juan and Ro Baudó), Valle del Cauca (Rio Raposo; Puerto Merizalde on the Rio Naya; Rio Sabaletas; 41km E. and 20km N. Buenaventura)), Costa Rica, Ecuador, Guyana, Panama. Refs.: Kröber, 1934: 273 (cat.; Catachlorops), Bequaert & Renjifo-Salcedo, 1946: 67 (Dichelacera (Catachlorops)); Fairchild, 1971: 65 (cat.); Moucha, 1976: 200 (cat.); Lee et al., 1969: 448 (Catachlorops); Wilkerson, 1979: 263; Fairchild & Burger, 1994: 100 (cat.); Coscaron & Papavero, 2009: 82 (cat.). Subgenus Dichelacera Macquart Dichelacera, subgenus Dichelacera Macquart, 1838. Type species, Tabanus cervicornis Fabricius, 1805. Refs.: Coscaron & Papavero, 2009: 82 (cat.). burgeri, Coscarón & Papavero, 2009: 82, nom. nov. for Dichelacera (Dichelacera) aurata Burger, 1994. Type locality: Colombia, Valle, Rio Zabaletas, 45km E. of Zabaletas. Distr.: Colombia (Antioquia, Cauca, Valley (probably Valle del Cauca). Refs.: Coscaron & Papavero, 2009: 82 (cat.). aurata Burger, 1994: 56. chocoensis Fairchild & Philip, 1960: 26. Type locality: Colombia, Chocó, Rio Nimiquia, Genené. HT F (MCZ), PT CATALOGUE OF TABANIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 265 M 1 (CAS), PT F 7 (FSCA). Distr. Colombia (Antioquia, Chocó (Rio Nimiquia), Cauca (Rio Micay), Valle del Cauca (Buenaventura; Rio Anchicayá; Rio Raposo)), Ecuador. Refs.: Lee et al., 1969: 447; Fairchild, 1971: 61 (cat.); Moucha, 1976: 197 (cat.); Wilkerson, 1979: 249; Fairchild & Burger, 1994: 95 (cat.); Coscaron & Papavero, 2009: 83 (cat.). damicornis (Fabricius), 1805: 101 (Tabanus). Type locality: “South America. Smidt” [i.e., Guyana, between rivers Demerara and Essequibo; cf. Papavero, 1971:21]. HT F (ZMKU). Distr.: ?Colombia, Venezuela to Brazil. Refs.: Fairchild, 1971: 61 (cat.); Moucha, 1976: 197 (cat.); Fairchild & Burger, 1994: 95 (cat.); Coscaron & Papavero, 2009: 83 (cat.). damicornis Hine, 1920b: 317, error. capricornis Kröber, 1931d: 414. fasciata Walker, 1850: 68. Type locality: “North America”. HT F (BMNH). Distr.: Colombia (Cauca (Guapi), Chocó (between Rio San Juan and Rio Baudó; Rio Nimiquía; Genené; Utría; Alto Curichi; Curichi) Valle del Cauca (Buenaventura; Rio Raposo; Rio Sabaletas; Bajo Calima; Rio Bravo-Playa Rica; 12km E. Buenaventura; 41km E. and 20km N. Buenaventura; Ladrilleros)), Nicaragua to Ecuador. Refs.: Lee et al., 1969: 447; Fairchild, 1971: 61 (cat.); Moucha, 1976: 197 (cat.); Wilkerson, 1979: 251; Fairchild & Burger, 1994: 95 (cat.); Coscaron & Papavero, 2009: 83 (cat.). cervicornis Fabricius of Osten Sacken, 1886: 58, missident. analis Hine, 1920b: 316. costaricensis Kröber, 1931d: 411. hartmanni Fairchild & Philip, 1960: 37. Type locality: Panama, Chiriquí, Santa Clara. HT F (MCZ), PT F l (FSCA). Distr.: Colombia (Santander), Panama. Refs.: Fairchild, 1971: 62 (cat.); Moucha, 1976: 198 (cat.); Fairchild & Burger, 1994: 96 (cat.); Coscaron & Papavero, 2009: 84 (cat.). marginata Macquart, 1847: 30 (1847: 14). Type locality: French Guiana, Cayenne. HT F (BMNH). Distr.: Colombia (Antioquia, Boyacá (Muzo), Chocó (Rio Curiche; La Teresita; Lloró; Rio Cabi near Quibdó; Andagoya), Magdalena (Rio Don Diego), Meta (Restrepo; Villavicencio), Santander (Jazmín), Valle del Cauca (Quebrada San Joaquin near Buenaventura)), Nicaragua to N. Brazil and Peru. Refs.: Kröber, 1934: 266 (cat.); Bequaert & Renjifo-Salcedo, 1946: 66; Fairchild, 1971: 63 (cat.); Moucha, 1976: 198 (cat.); Wilkerson, 1979: 253; Fairchild & Burger, 1994: 97 (cat.); Coscaron & Papavero, 2009: 84 (cat.). hinnulus Walker, 1850b: cxxii. emarginata Hine, 1917: 293, misspell. melanosoma Hine, 1920b: 316. Type locality: Costa Rica, San Mateo, Higuito. HT F (USNM). Distr.: Colombia (Chocó (La Teresita)), Costa Rica to N. Colombia. Refs.: Fairchild, 1971: 63 (cat.); Moucha, 1976: 198 (cat.); Wilkerson, 1979: 254; Fairchild & Burger, 1994: 97 (cat.); Coscaron & Papavero, 2009: 85 (cat.). regina Fairchild, 1940b: 699. Type locality: Panama, Coclé, El Valle, 2500ft. HT F (MCZ). Distr.: Colombia (Antioquia, Cauca (Rio Micay), Chocó (upper Curiche; Rio Curiche, between Rio San Juan and Rio Baudó; Itsmina; Rio Nimiquía), Meta (Restrepo), Valle del Cauca (Rio Raposo; bajo Anchicayá, Lago Calima; Rio BravoPlaya Rica; 20km below dam Lago Calima; 41km E. and 20km N. Buenaventura; Puerto Merizalde on Rio Naya; Veneral on Rio Yurumanguí)), Honduras to Ecuador. Refs.: Bequaert & Renjifo-Salcedo, 1946: 66; Lee et al., 1969: 448; Fairchild, 1971: 64 (cat.); Moucha, 1976: 198 (cat.); Wilkerson, 1979: 256; Fairchild & Burger, 1994: 98 (cat.); Coscaron & Papavero, 2009: 86 (cat.). grandis Ricardo of Hine, 1917: 293, misident. submarginata Lutz, 1915: 86. Type locality: “Venezuela”. LT F (IOC). Distr.: Colombia (Antioquia (Anori River valley), Cundinamarca (Yacopí), Meta (Ocoa, Villavicencio), Putumayo (La Tagua), Santander (Gualilo, Landázuri)), Bolivia, Costa Rica to Venezuela, E. Peru. Refs.: Kröber, 1934: 267 (cat.); Bequaert & RenjifoSalcedo, 1946: 66; Fairchild, 1971: 64 (cat.); Moucha, 1976: 199 (cat.); Wilkerson, 1979: 258; Fairchild & Burger, 1994: 98 (cat.); Coscaron & Papavero, 2009: 86 (cat.). tricallosa Kröber, 1931d: 414. 266 · Zootaxa 4122 (1) © 2016 Magnolia Press E & MIRANDA-ESQUIVEL villavoensis Fairchild & Philip, 1960: 74. Type locality: Colombia, Meta, Villavicencio (as Viliavicencio, error). HT F (MCZ), PT F 2 (FSCA). Distr.: Colombia (Santander, Meta), Brazil, Ecuador, Peru. Refs.: Fairchild, 1971: 65 (cat.); Moucha, 1976: 199 (cat.); Fairchild & Burger, 1994: 99 (cat.); Coscaron & Papavero, 2009: 87 (cat.). submarginata Lutz of Bequaert & Renjifo-Salcedo, 1946: 66, misident. Subgenus Idiochelacera Fairchild Dichelacera, subg. Idiochelacera Fairchild, 1969: 210. Type species, Dichelacera subcallosa Fairchild & Philip, 1960 (orig. des.). subcallosa Fairchild & Philip, 1960: 83. Type locality: Colombia, Santander, Rio Suárez, 900m. HT F, PT M 1 (AMNH). Distr.: Colombia (Antioquia (Providencia), Chocó (between Rio San Juan and Rio Baudó), Santander, Valle del Cauca (Bajo Anchicayá)), Costa Rica, Ecuador, Panama, Peru. Refs.: Fairchild, 1971: 65 (cat.); Moucha, 1976: 200 (cat.); Wilkerson, 1979: 261; Fairchild & Burger, 1994: 99 (cat.); Coscaron & Papavero, 2009: 87 (cat.). Subgenus Orthostyloceras Lutz, 1933 Orthostylus Lutz & Neiva, 1914: 74 (sep., p. 8) (preocc. Beck, 1837). Type species, Orthostylus ambiguus Lutz & Neiva, 1914 (mon.). Orthostyloceras Lutz in Borgmeier, 1933: 298 (as genus; nom. nov. for Orthostylus Lutz, & Neiva, 1914). Type species, Orthostylus ambiguus Lutz & Neiva, 1914 (aut.). Refs.: Coscaron & Papavero, 2009: 88 (cat.). aurata Wilkerson, 1979: 265. Type locality: Colombia, Cauca, Cerro de Munchique, El Tambo, 2400m. HT F (FSCA). Distr.: Colombia (Cauca (Cerro de Munchique)). Refs.: Fairchild & Burger, 1994: 100 (cat.); Coscaron & Papavero, 2009: 88 (cat.). Genus Dicladocera Lutz, 1913 Dicladocera Lutz, 1909b: 29, 1911b: 34, nomen nudum. Dicladocera Lutz, 1913b: 486, 487 (sep., pp. 6, 7) (1914a: 166 (sep., p. 6) (as Dichladocera). Type species, Tabanus guttipennis Wiedemann, 1828 (subs. des., Enderlein, 1922: 346 (in key)). Refs.: Fairchild, 1951a: 121, 1961a: 238; . Coscaron & Papavero, 2009: 89 (cat.); Henriques et al., 2012: 278. Dasychela Enderlein of Bequaert & Renjifo-Salcedo, 1946: 68, misident. (part). argentomacula Wilkerson, 1979: 280. Type locality: Colombia, Valle, Peñas Blancas, 10km W. of Cali. HT F, PT F 5 (FSCA). Distr.: Colombia (Valle del Cauca (Peñas Blancas; 10km W. Cali; Anchicayá; Lago Calima; Tres Esquinas; Habana; Buga)), Bolivia, Ecuador, Peru. Refs.: Fairchild & Burger, 1994: 109 (cat.); Coscaron & Papavero, 2009: 89 (cat.). argenteomaculata Fairchild & Burger, 1994: 109 (cat.), error. basirufa (Walker), 1850b: 32 (Tabanus). Type locality: “Colombia”. HT F (BMNH). Distr.: Colombia (CaucaHuila (Páramo de las Papas sources of Rio Magdalena), Cundinamarca (near Guasca), Nariño (near Pasto)), Ecuador. Refs.: Kröber, 1934: 268 (cat.); Bequaert & Renjifo-Salcedo, 1946: 71 (Dasychela); Fairchild, 1971: 73 (cat.); Moucha, 1976: 227 (cat.); Wilkerson, 1979: 283; Fairchild & Burger, 1994: 109 (cat.); Coscaron & Papavero, 2009: 89 (cat.). steinheili Enderlein, 1925: 384. pallidetibia Kröber, 1940. beaveri Wilkerson, 1979: 284. Type locality: Colombia, Cauca, Cerro de Munchique, 2450m. HT F (FSCA). Distr.: Colombia (Cauca (Cerro de Munchique)). Refs.: Fairchild & Burger, 1994: 109 (cat.); Coscaron & Papavero, 2009: 89 (cat.). calimaensis Wilkerson, 1979: 286. Type locality: Colombia, Valle, lago Calima, 8km below dam, 1250m. HT F, CATALOGUE OF TABANIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 267 PT 5 F (FSCA). Distr.: Colombia (Valle del Cauca (Lago Calima)). Refs.: Fairchild & Burger, 1994: 109 (cat.); Coscaron & Papavero, 2009: 89 (cat.). clara (Schiner), 1868: 88 (Tabanus). Type locality: “Venezuela”. ST F (NMW). Distr.: Colombia (Valle del Cauca (Peñas Blancas; Lago Calima)), Ecuador, Venezuela, Peru. Refs.: Kröber, 1934: 308 (cat.; Tabanus (Tabanus), as clarus); Wilkerson, 1979: 287 (as clarus, error); Fairchild & Burger, 1994: 110 (cat.); Coscaron & Papavero, 2009: 89 (cat.). umbratipenne Fairchild 1958: 522. dalessandroi Wilkerson, 1979: 289. Type locality: Colombia, Valle, lower Anchicayá, 400m. HT F (FSCA). Distr.: Colombia (Valle del Cauca (Bajo Anchicayá)). Refs.: Fairchild & Burger, 1994: 110 (cat.); Coscaron & Papavero, 2009: 89 (cat.). distomacula Wilkerson, 1979: 290. Type locality: Colombia, Cauca, Cerro de Munchique, 2450m. HT F (FSCA). Distr.: Colombia (Cauca (Cerro de Munchique)), Ecuador. Refs.: Fairchild & Burger, 1994: 110 (cat.); Coscaron & Papavero, 2009: 89 (cat.). hirsuta Wilkerson, 1979: 291. Type locality: Colombia, Cauca, Parque de Puracé, lago San Rafael, 3500m. HT F, PT F 2 (FSCA). Distr.: Colombia (Cauca (Parque de Puracé, Lago San Rafael)), Ecuador. Refs.: Fairchild & Burger, 1994: 110 (cat.); Coscaron & Papavero, 2009: 90 (cat.). leei Fairchild in Wilkerson, 1979: 293. Type locality: Colombia, Cauca, Páramo de Puracé, Sierra de Puracé, vic. Cali, 3500m. HT F, PT F 2, M 1 (FSCA). Distr.: Colombia (Cauca (Parque de Puracé, Lago San Rafael)). Refs.: Fairchild & Burger, 1994: 110 (cat.); Coscaron & Papavero, 2009: 90 (cat.). macula (Macquart), 1846: 171 (1846: 43) (Tabanus). Type locality: unknown (?Colombia). HT F (BMNH). Distr.: Colombia (Cundinamarca (Bogotá, Arrayán, Fusagasugá, Páramo), Huila (20km W. Leticia), Magdalena (NW. Sierra Nevada de Santa Marta; San Lorenzo Mts), Valle del Cauca (La Cumbre)), Bolivia, Ecuador, Peru, Venezuela. Refs.: Bequaert & Renjifo-Salcedo, 1946: 70 (Dasychela); Fairchild, 1971: 73 (cat.); Moucha, 1976: 227 (cat.); Wilkerson, 1979: 295; Fairchild & Burger, 1994: 110 (cat.); Coscaron & Papavero, 2009: 90 (cat.). auribarbis Macquart, 1848: 12 (Tabanus). argyrophorus Schiner, 1868: 90 (Tabanus). maculata Pechuman, 1944: 55, error. minos (Schiner), 1868: 88 (Tabanus). Type locality: “South America” (Venezuela). LT F (NMW). Distr.: Colombia (Cauca (Silvia), Santander (Páramo del Almorzadero)), Bolivia, ?Brazil, Ecuador, Venezuela. Refs.: Kröber, 1934: 269 (cat.; Dicladocera (Dasyrhamphis)); Bequaert & Renjifo-Salcedo, 1946: 75; Wilkerson, 1979: 296; Fairchild & Burger, 1994: 111 (cat.); Coscaron & Papavero, 2009: 90 (cat.). nigrocoerulea (Rondani), 1850: 191 (Tabanus). Type locality: “Venezuela”. ST F (IEAUT, Naples). Distr.: Colombia (Valle del Cauca (Peñas Blancas, Lago Calima)), ?Brazil, Ecuador, Panama, ?Peru, Venezuela. Refs.: Fairchild, 1971: 74 (cat.); Moucha, 1976: 228 (cat.) Wilkerson, 1979: 297; Fairchild & Burger, 1994: 111 (cat.); Coscaron & Papavero, 2009: 90 (cat.). Lutzi Surcouf, 1921: 54 (Stigmatophthalmus). acheronitens Kröber, 1931d: 403. pruinosa Wilkerson, 1979: 298. Type locality: Colombia, Valle, Peñas Blancas. HT F, PT F 2 (FSCA). Distr.: Colombia (Cauca (Cerro de Munchique), Valle del Cauca (Peñas Blancas, Lago Calima)), Ecuador. Refs.: Fairchild & Burger, 1994: 111 (cat.); Coscaron & Papavero, 2009: 91 (cat.). riveti (Surcouf), 1919: 226 (Tabanus). Type locality: Ecuador, between Guanasilla and San Nicolás. ST F (MNHNP). Distr.: Colombia (Valle del Cauca (Bajo Anchicayá, Rio Sabaletas)), Ecuador. Refs.: Wilkerson, 1979: 300; Fairchild & Burger, 1994: 111 (cat.); Coscaron & Papavero, 2009: 91 (cat.). 268 · Zootaxa 4122 (1) © 2016 Magnolia Press E & MIRANDA-ESQUIVEL submacula (Walker), 1850b: 30 (Tabanus). Type locality: ‘Colombia’. HT F (BMNH). Distr.: Colombia (Boyacá (near Muzo, Maripí), ?Magdalena, ?Valle)), Ecuador. Refs.: Fairchild, 1971: 75 (cat.); Moucha, 1976: 228 (cat.); Wilkerson, 1979: 302; Fairchild & Burger, 1994: 111 (cat.); Coscaron & Papavero, 2009: 91 (cat.). Genus Hemichrysops Kröber Hemichrysops Kröber, 1930c: 237. Type species, Hemichrysops fascipennis Kröber, 1930 (mon.). Refs.: Coscaron & Papavero, 2009: 92 (cat.). fascipennis Kröber, 1930c: 237. Type locality: “Colombia”. HT F (BMNH). Distr.: Colombia (Valle del Cauca (Rio Raposo)), Costa Rica, Panama. Refs.: Kröber, 1934: 231 (cat.); Bequaert & Renjifo-Salcedo, 1946: 59; Lee et al., 1969: 451; Fairchild, 1971: 53 (cat.); Moucha, 1976: 226 (cat.); Wilkerson, 1979: 203; Fairchild & Burger, 1994: 85 (cat); Coscaron & Papavero, 2009: 92 (cat.). vecordis Fairchild 1956: 16 (Tabanus; nom. nov. for fascipennis Kröber, 1930; when in Tabanus, fascipennis Kröber, 1930 was preocc., Macquart, 1845, by Coscaron & Papavero, 2009: 93 (cat.)). Genus Himantostylus Lutz Himantostylus Lutz, 1913a: 183 (sep., p. 44). Type species, Himantostylus intermedius Lutz, 1913. Refs.: Fairchild, 1966: 211. Coscaron & Papavero, 2009: 93 (cat.). intermedius Lutz, 1913b: 183 (sep., p. 44), Type locality: Peru, Puno, Yahuarmayo. HT M (IOC). Distr.: Colombia (Antioquia, Valle del Cauca (Rio Raposo; Rio Sabaletas)), Panama to Bolivia. Refs.: Lee et al., 1969: 446; Fairchild, 1971: 51 (cat.); Moucha, 1976: 208 (cat.); Wilkerson, 1979: 223; Fairchild & Burger, 1994: 81 (cat); Coscaron & Papavero 2009: 93 (cat.). Genus Lepiselaga Macquart, 1838 Subgenus Conoposelaga Barretto Conoposelaga Barretto, 1949b: 87 (as genus). Type species, Lepiselaga aberrans Lutz, 1913 (orig. des.). Refs.: Coscaron & Papavero, 2009: 94 (cat.). colombiana Fairchild, 1966: 215. Type locality: Colombia, Meta, Barbascal. HT F (MCZ), PT F l (FSCA). Distr.: Colombia (Meta). Refs.: Fairchild, 1971: 85 (cat.); Moucha, 1976: 207 (cat.); Fairchild & Burger, 1994: 125 (cat.); Coscaron & Papavero, 2009: 94 (cat.). Subgenus Lepiselaga Macquart, 1838 crassipes (Fabricius), 1805: 108 (Haematopota). Type locality: “South America. Smidt” (i.e., Guyana, between rivers Demerara and Essequibo; cf. Papavero, 1971: 21). HT F (ZMKU). Distr.: Colombia (Amazonas (Santa Sofia Island), Antioquia (Caucasia), Caldas (Dorada, Magdalena River), Cauca (Rio Micay), Chocó (Atrato River, Sautatá; ), Cordoba (Monteria), Meta (Hacienda Yacuana near Villavicencio), Nariño (Rio Patía), Santander (Puerto Olaya; Barrancabermeja), Valle del Cauca (Buenaventura, Rio Raposo; Candelaria; Puerto Merizalde, Rio Naya; Cali)), Mexico to N. Argentina, Cuba, Jamaica, Hispaniola, Puerto Rico. Refs.: Kröber, 1934: 253 (cat.); Bequaert & Renjifo-Salcedo, 1946: 64; Lee et al., 1969: 446; Fairchild, 1971: 84 (cat.); Moucha, 1976: 207 (cat.); Wilkerson, 1979: 351; Fairchild & Burger, 1994: 125 (cat.); Parra-Henao et al., 2008: 183; Coscaron & Papavero, 2009: 94 (cat.). lepidotus Wiedemann, 1828: 193 (Tabanus). recta Loew, 1869: 46 (Lepidoselaga). CATALOGUE OF TABANIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 269 major Szilády, 1926: 21. fenestrata Hack, 1953: 346 (as crassipes var.). Genus Leucotabanus Lutz Leucotabanus Lutz, 1913a: 487 (1914: 167 (sep., p. 7)). Type species, Tabanus leucaspis Wiedemann, 1828 (subs. mon., Lutz & Neiva, 1914:71 (sep., p. 5)) = Leucotabanus exaestuans (Linnaeus, 1758). Refs.: Fairchild, 1951a: 123; 1961: 240, Coscaron & Papavero, 2009: 95 (cat.). canithorax Fairchild, 1941a: 634, (nom. nov. for albicans Macquart of Macquart 1845, misident). Type locality: “Colombia”. HT F (MCZ). Distr.: Colombia (Magdalena (Sevilla)), Mexico, Belice, Panama, Guyana, Trinidad. Refs.: Bequaert & Renjifo-Salcedo, 1946: 74; Fairchild, 1971: 83 (cat.); Moucha, 1976: 230 (cat.); Wilkerson, 1979: 347; Fairchild & Burger, 1994: 123 (cat.); Coscaron & Papavero, 2009: 95 (cat.). albicans Macquart of Macquart, 1846: 37, misident. exaestuans (Linnaeus), 1758: 601 (Tabanus). Type locality: “America”. HT F (NRS). Distr.: Colombia (Antioquia, Boyacá, Chocó (La Teresita), Meta (Villavicencio), Putumayo (Puerto Asis), Santander, Valle del Cauca (Rio Sabaletas; Bajo Anchicayá), Vaupés), Mexico to Bolivia, Argentina, Trinidad. Refs.: Fairchild, 1971: 83 (cat.); Moucha, 1976: 230 (cat.); Wilkerson, 1979: 348; Fairchild & Burger, 1994: 123 (cat.); Coscaron & Papavero, 2009: 95 (cat.). leucaspis Wiedemann, 1828: 179 (Tabanus). albicans Macquart, 1846: 165 (1846: 37) (Tabanus; preocc. Macquart 1834). albiscutellatus Macquart, 1850: 338 (1850: 34) (Tabanus). cingulifer Walker, 1857: 123 (Tabanus). melanopterus Brèthes, 1910: 479 (Tabanus). exoestuans Barretto, 1957: 88, error. flavinotum (Kröber), 1934: 309 (cat.; Tabanus (Tabanus); nom. nov. for nigriflavus Kröber, 1931. Type locality: Brazil, Amazonas. Distr.: Colombia (Antioquia (Providencia), Boyacá (Muzo), Chocó), Costa Rica to Colombia, Brazil, Peru, Venezuela. Refs.: Bequaert & Renjifo-Salcedo, 1946: 74; Fairchild, 1971: 83 (cat.); Moucha, 1976: 230 (cat.); Wilkerson, 1979: 350; Fairchild & Burger, 1994: 123 (cat.); Coscaron & Papavero, 2009: 96 (cat.). nigriflavus Kröber, 1931e: 292 (Tabanus; preocc. Kröber, 1930). janinae Fairchild, 1970: 845. Type locality: French Guiana, Forêt d’Acarouany. HT F (MNHNP), PT F 2 (FSCA). Distr.: Brazil, Colombia, French Guiana, Surinam. Refs.: Fairchild & Burger, 1994: 123 (cat.); Coscaron & Papavero, 2009: 96 (cat.). pauculus Fairchild, 1951b: 456. Type locality: Brazil, Pará HT F (MCZ). Distr.: Colombia, Venezuela, Brazil. Refs.: Fairchild, 1971: 83 (cat.); Moucha, 1976: 231 (cat.); Fairchild & Burger, 1994: 124 (cat.); Coscaron & Papavero, 2009: 96 (cat.). Genus Phaeotabanus Lutz Phaeotabanus Lutz, 1913a: 487 (sep., p. 6) (1914a: 168 (sep., p. 8). Type species, Tabanus litigiosus Walker, 1850 (subs. des., Bequaert, 1924: 29). Refs.: Fairchild, 1951a: 124, 1961a: 243. Coscaron & Papavero, 2009: 98 (cat.). cajennensis (Fabricius), 1787: 355 (Tabanus). Type locality: French Guiana, “Cayenne. Rohr” (cf. Papavero, 1971: 20). LT F (ZMKU). Distr.: Colombia (Meta (region of Rio Negro, S. of Villavicencio), Tolima (between Mediación and Ibagué)), S. to Brazil, Bolivia, Trinidad to Colombia and French Guiana. Refs.: Kröber, 1934: 307 (cat.; Tabanus (Tabanus)); Bequaert & Renjifo-Salcedo, 1946: 76 (Stenotabanus); Fairchild, 1971: 56 (cat.); Moucha, 1976: 194 (cat.); Fairchild & Burger, 1994: 88 (cat.); Coscaron & Papavero, 2009: 98 (cat.). 270 · Zootaxa 4122 (1) © 2016 Magnolia Press E & MIRANDA-ESQUIVEL caiennensis Kertész 1908: 232 (cat.; Tabanus); Lutz, 1928: 61, error. caiennensus Lutz, 1928: caption of fig. 8 (Pl. IX). decoloratus Kröber, 1931f: 301 (Tabanus (Tabanus)). innotescens (Walker), 1854: 327 (Tabanus). Type locality: Brazil, Pará, Rio Tapajós. HT F (BMNH). Distr.: Colombia. Refs.: Fairchild & Burger, 1994: 88 (cat.); Coscaron & Papavero, 2009: 99 (cat.). hesperus Lutz, 1912: 8, fig. 3 (Tabanus). aurora Macquart of Bequaert, 1926: 233 and Kröber, 1929e: 246 (female only), misident. pallidipalpis Stone, 1944: 131 (Cryptotylus). nigriflavus (Kröber), 1930d: 284, (Tabanus). Type locality: “French Guiana”. Distr.: Colombia. Refs.: Fairchild & Burger, 1994: 89 (cat.); Coscaron & Papavero, 2009: 100 (cat.). obscurehirtus Kröber, 1930d: 284 (Tabanus (Phaeotabanus); as aphanopterus var.; preocc. Ricardo, 1908). obscuripilis Kröber, 1934: 304 (cat.; Tabanus (Phaeotabanus), as aphanopterus var., nom nov. for obscurehirtus, Kröber, 1930). obscuripilus Philip, 1960a: 81 (as aphanopterus var.), error. obscurepilis Moucha, 1976: 195 (cat.), error. obscurepilus Fairchild & Burger, 1994: 89, error. phaeopterus Fairchild, 1964b: 176. Type locality: Panama, Darien, Rio Tapalices, 200m. HT F, PT M 1 (MCZ), PT F 2 (FSCA). Distr.: Colombia (Antioquia, Chocó (between Rio San Juan and Rio Baudó), Valle del Cauca (Bajo Anchicayá; Rio Sabaletas, 41Km E. and 20km N. of Buenaventura)), Ecuador, Panama, Peru. Refs.: Wilkerson, 1979: 241; Fairchild & Burger, 1994: 89 (cat.); Coscaron & Papavero, 2009: 100 (cat.). serenus (Kröber), 1931f: 304 (Tabanus (Tabanus)). Type locality: “Venezuela”. HT F (NMW). Distr.: Colombia (Antioquia (Providencia), Boyacá, Meta (Restrepo)), Ecuador, Panama, Venezuela. Refs.: Fairchild, 1971: 82 (cat.; Stypommisa as serena); Moucha, 1976: 232 (cat.; Stypommisa); Wilkerson, 1979: 342 (Stypommisa) (as serena); Fairchild & Burger, 1994: 90 (cat.); Coscaron & Papavero, 2009:100 (cat.). vapidus Fairchild, 1964a: 172 (Stenotabanus). Genus Philipotabanus Fairchild, 1943 Subgenus Melasmatabanus Fairchild Philipotabanus, subg. Melasmatabanus Fairchild, 1964a: 179. Type species, Tabanus fascipennis Macquart, 1846 (orig. des.). criton (Kröber), 1934: 304 (cat.; Tabanus (Phaeotabanus), nom. nov. for columbianus Kröber, 1931). Type locality: Colombia, Fusagasugá. Distr.: Colombia (Antioquia (Providencia), Cundinamarca (Fusagasuga), Chocó (Rio Curiche), Santander (San Vicente, Rio Suarez), Valle del Cauca (Rio Raposo)), Brazil, Ecuador, Panama. Refs.: Kröber, 1934: 304 (cat.); Bequaert & Reinjifo-Salcedo 1946: 82 (Tabanus (?Philipotabanus); Fairchild, 1971: 80 (cat.); Moucha, 1976: 230 (cat.); Wilkerson, 1979: 318; Fairchild & Burger, 1994: 118 (cat.); Coscaron & Papavero, 2009: 100 (cat.). columbianus Kröber, 1931g: 90 (Phaeotabanus; preocc. Enderlein, 1925). keenani Fairchild, 1947: 574 (Tabanus (Philipotabanus)). fascipennis (Macquart), 1846: 163 (1846: 35), (Tabanus). Type locality: “Nouvelle Grenade” (Colombia). LT F (BMNH). Distr.: Colombia (Antioquia, Boyacá (Muzo), Valle del Cauca (Lower Anchicayá, 41km E. and 20km Buenaventura)), Panama, Venezuela. Refs.: Bequaert & Renjifo-Salcedo, 1946: 82 (Tabanus (Philipotabanus)); Fairchild, 1971: 80 (cat.); Moucha, 1976: 230 (cat.); Wilkerson, 1979: 319; Fairchild & Burger, 1994: 118 (cat.); Coscaron & Papavero, 2009: 101 (cat.). CATALOGUE OF TABANIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 271 nigripennis Wilkerson, 1979: 321. Type locality: Colombia, Valle, lago Calima. HT F, PT F 2 (FSCA). Distr.: Colombia (Chocó, Valle del Cauca), Ecuador E. of Andes. Refs.: Fairchild & Burger, 1994: 118 (cat.); Coscaron & Papavero, 2009: 101 (cat.). Subgenus Mimotabanus Fairchild Philipotabanus, subg. Mimotabanus Fairchild, 1964a: 179. Type species, Tabanus (Philipotabanus) inauratus Fairchild (orig. des.). Refs.: Coscaron & Papavero, 2009: 101(cat.). annectans Fairchild, 1975: 691. Type locality: Colombia, Antioquia, Rio Anori valley, Providencia. HT F, PT F 4 (FSCA). Distr.: Colombia (Antioquia (Providencia)). Refs.: Fairchild & Burger, 1994: 117 (cat.); Coscaron & Papavero, 2009: 101 (cat.). fucosus Fairchild, 1958: 526. Type locality: Colombia, Meta, Villavicencio (El Retiro). HT F (MCZ), PT F l (FSCA). Distr.: Colombia (Meta), Peru. Refs.: Fairchild, 1971: 79 (cat.); Moucha, 1976: 229 (cat.); Fairchild & Burger, 1994: 117 (cat.); Coscaron & Papavero, 2009: 101 (cat.). opimus Fairchild, 1975: 691. Type locality: Colombia, Cundinamarca, Monterredondo. HT F (DECU), PT F 8, M 1 (FSCA). Distr.: Colombia (Cundinamarca), Ecuador to Peru. Refs.: Fairchild & Burger, 1994: 118 (cat.); Coscaron & Papavero, 2009: 101 (cat.). phalaropygus Fairchild, 1964a: 177. Type locality: Panama, Darien, Rio Tuira Yellow Fever Station. HT F (MCZ), PT F l (FSCA). Distr.: Colombia (Valle del Cauca (Rio Pance near Buenos Aires, Cali)), Panama, Peru. Refs.: Wilkerson, 1979: 322; Fairchild & Burger, 1994: 118 (cat.); Coscaron & Papavero, 2009: 101 (cat.). plenus (Hine), 1907: 225 (Tabanus). Type locality: Guatemala, Izabal. ST F (AMNH, OSU). Distr.: Guatemala to Colombia. Refs.: Fairchild, 1971: 80 (cat.); Moucha, 1976: 229 (cat.); Fairchild & Burger, 1994: 118 (cat.); Coscaron & Papavero, 2009: 101 (cat.). porteri Fairchild, 1975: 692. Type locality: Colombia, Antioquia, Rio Anori valley, La Tirana. HT F (MCZ), PT F 3 (FSCA), PT F l (BMNH). Distr.: Colombia (Antioquia (Rio Anori valley, La Tirana), Valle del Cauca (41km E. and 21km N. Buenaventura, Rio Raposo, Rio Bravo-Playa Rica, 15km below dam Lago Calima, Rio Sabaletas)). Refs.: Wilkerson, 1979: 323; Fairchild & Burger, 1994: 118 (cat.); Coscaron & Papavero, 2009: 101 (cat.). tanypterus Wilkerson, 1979: 325. Type locality: Colombia, Valle, near Santa María, 200-2500m. HT F (FSCA). Distr.: Colombia (Valle del Cauca). Refs.: Fairchild & Burger, 1994: 118 (cat.); Coscaron & Papavero, 2009: 101 (cat.). vulpinus Fairchild, 1975: 690. Type locality: Colombia, Buenaventura, Rio Raposo. HT F (FSCA). Distr.: Colombia (Valle del Cauca (Rio Raposo, Bajo Anchicayá)). Refs.: Wilkerson, 1979: 326; Fairchild & Burger, 1994: 118 (cat.); Coscaron & Papavero, 2009: 101 (cat.). Subgenus Philipotabanus Fairchild, 1943 chrysothrix (Fairchild), 1943: 459 (Tabanus (Philipotabanus)). Type locality: Panama, Canal Zone, Madden Lake, Rio Pequení. Distr.: Colombia (Chocó), Panama. Refs.: Fairchild, 1971: 78 (cat.); Fairchild & Burger, 1994: 116 (cat.); Coscaron & Papavero, 2009: 102 (cat.). grassator (Fairchild), 1953: 278, (Tabanus (Philipotabanus)). Type locality: Panama, Bocas del Toro, Almirante. HT F (MCZ). Distr.: Colombia (Valle del Cauca), Panama. Refs.: Fairchild & Burger, 1994: 116 (cat.); Coscaron & Papavero, 2009: 102 (cat.). magnificus (Kröber), 1934: 299 (cat.; Tabanus (Phaeotabanus), nom. nov. for formosus Kröber, 1930). Type 272 · Zootaxa 4122 (1) © 2016 Magnolia Press E & MIRANDA-ESQUIVEL locality: Ecuador, Pucay, Santo Domingo; Colombia; Costa Rica. Distr.: Colombia (Antioquia, Cauca (Rio Micay), Chocó (between Rio San Juan and Rio Baudó, La Teresita, Curiche, Rio Nimiquia, Itsmina Andagoya, Rio Cabí near Quibdó, Condoto, Utria), Meta (Rio Guayuriba), Santander (Alto Rio), Valle del Cauca (Buenaventura, Rio Anchicayá, Rio Raposo, Rio Sabaletas, Bajo Anchicayá, Bajo Calima, Rio Bravo-Playa Rica, 15Km below dam Lago Calima, 41km E. and 20km N. Buenaventura, 3.6km W. Queremal, Cali to Buenaventura road, Veneral on Rio Yurumanguí, Puerto Merizalde on Rio Naya, Quebrada San Joaquín near Buenaventura)), Honduras to Venezuela and Ecuador. Refs.: Kröber, 1934: 305 (cat.; Tabanus (Phaeotabanus)); Bequaert & Renjifo-Salcedo, 1946: 82 (Tabanus (Philipotabanus); Lee et al., 1969: 451; Fairchild, 1971: 79 (cat.); Moucha, 1976: 229 (cat.); Wilkerson, 1979: 327; Fairchild & Burger, 1994: 116 (cat.); Coscaron & Papavero, 2009: 102 (cat.). formosus Kröber, 1930d: 299 (Tabanus (Phaeotabanus); preocc. Walker, 1848). nigrinubilus (Fairchild)1953: 279 (Tabanus (Philipotabanus)). Type locality: Panama, Bocas del Toro, Almirante. HT F (MCZ), PT F 2 (FSCA). Distr.: Colombia (Antioquia, Cauca, Chocó (upper Curiche river), Valle del Cauca (Rio Raposo, 41km E. and 20km N. Buenaventura, Rio Sabaletas, Bajo Calima)), Costa Rica, Ecuador, Honduras, Panama. Refs.: Lee et al., 1969: 452; Fairchild, 1971: 79 (cat.); Moucha, 1976: 229 (cat.); Wilkerson, 1979: 329; Fairchild & Burger, 1994: 117 (cat.); Coscaron & Papavero, 2009: 102 (cat.). pterographicus (Fairchild), 1943: 459, (Tabanus (Philipotabanus)). Type locality: Panama, Chagres river region, Moja Pollo. HT F (MCZ).Distr.: Colombia (Chocó), Costa Rica, Ecuador, Panama. Refs.: Fairchild, 1971: 79 (cat.); Moucha, 1976: 229 (cat.); Wilkerson, 1979: 331; Fairchild & Burger, 1994: 117 (cat.); Coscaron & Papavero, 2009: 103 (cat.). tenuifasciatus (Kröber), 1930d: 296, (Tabanus (Phaeotabanus)). Type locality: Peru, Pichís, Puerto Yessup. HT F (DFS). Distr.: Brazil, Colombia, Ecuador, Peru. Refs.: Fairchild, 1971: 79 (cat.); Moucha, 1976: 229 (cat.); Fairchild & Burger, 1994: 117 (cat.); Coscaron & Papavero, 2009: 103 (cat.). Genus Pseudacanthocera Lutz & Neiva, 1914 Pseudacanthocera Lutz, 1913a: 487 (sep., p. 6) (1914a: 167 (sep., p. 7)), nomen nudum. Pseudacanthocera Lutz & Neiva, 1914: 71 (sep., p. 5). Type species, Silvius marginatus Macquart, 1838: 1, error = Silvius sylveirii Macquart, 1838 (mon.). Refs. Coscaron & Papavero, 2009: 103 (cat.) Pseudoselasoma Brèthes of Enderlein, 1925: 350 (part), misident. Pseudocanthocera Fairchild, 1951a: 124, error. brevicornis (Enderlein), 1925: 351 (Pseudoselasoma). Type locality: Paraguay, San Bernardino. LT F, PLT M (MNHUB). Distr.: Argentina, Bolivia, Brazil, Colombia, Paraguay. Refs.: Kröber, 1934: 260 (cat.); Fairchild, 1971: 84 (cat.; as brevicorne); Moucha, 1976: 227 (cat.; as brevicorne); Fairchild & Burger, 1994: 124 (cat.; as brevicorne); Coscaron & Papavero, 2009: 103 (cat.). fraterna Kröber, 1930a: 80. lenkoi Bernardi 1975: 75. Genus Selasoma Macquart Selasoma Macquart, 1838a: 187 (1838: 303). Type species, Tabanus tibialis Fabricius, 1805 (orig. des.). Refs.: Coscaron & Papavero, 2009: 104 (cat.). tibiale (Fabricius), 1805: 102 (Tabanus). Type locality: “South America. Smidt”. HT F (ZMKU). Distr.: Colombia (Antioquia (Providencia), Meta (Villavicencio)), Mexico to N. Argentina, Trinidad. Refs.: Bequaert & RenjifoSalcedo, 1946: 64; Fairchild, 1971: 55 (cat.); Moucha, 1976: 226 (cat.); Wilkerson, 1979: 236; Fairchild & Burger, 1994: 86 (cat.); Coscaron & Papavero, 2009: 104 (cat.). chalybeus Perty, 1833: 183 (Hadrus). cyaneum Walker, 1848: 208 (Hadrus). CATALOGUE OF TABANIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 273 Genus Spilotabanus Fairchild Spilotabanus Fairchild, 1969: 218. Type species, Tabanus multiguttatus Kröber, 1930 (orig. des.). multiguttatus (Kröber), 1930d: 285, (Tabanus (Phaeotabanus)). Type locality: Colombia, Bogotá. HT F destroyed (Col. Kröber). Distr.: Colombia (Cauca (Cerro de Munchique), Cundinamarca, Magdalena (Sierra San Lorenzo), Putumayo (on Amazon side of Andes along road leading to Mocoa)), Ecuador, Peru. Refs.: Kröber, 1934: 305 (cat.; Tabanus (Phaeotabanus)); Bequaert & Renjifo-Salcedo, 1946: 82 (Tabanus (Philipotabanus); Fairchild, 1971: 57 (cat.); Moucha, 1976: 230 (cat.); Wilkerson, 1979: 243; Fairchild & Burger, 1994: 90 (cat.); Coscaron & Papavero, 2009: 105 (cat.). triaurius Wilkerson, 1979: 245. Genus Stenotabanus Lutz, 1913 Subgenus Brachytabanus Fairchild Stenotabanus, subg. Brachytabanus Fairchild, 1942e: 300. Type species, Stenotabanus longipennis Kröber, 1929 (orig. des.). Refs.: Coscaron & Papavero, 2009: 107 (cat.). longipennis Kröber, 1929b: 125. Type locality: “Venezuela”. HT F (NMW). Distr.: Colombia (Boyacá (Muzo), Meta (Villavicencio), Santander (Bocas del Rosario, Rio Magdalena and an unknow locatin on the upper Magdalena River), Putumayo (Urcusique), Valle del Cauca (Rio Raposo)), Costa Rica, Panama, Venezuela. Refs.: Bequaert & Renjifo-Salcedo, 1946: 75 (Brachytabanus); Lee et al., 1969: 450; Fairchild, 1971: 50 (cat.); Wilkerson, 1979: 219; Fairchild & Burger, 1994: 79 (cat.); Coscaron & Papavero, 2009: 107 (cat.). Subgenus Stenotabanus Lutz, 1913 brunneus Wilkerson, 1979: 208. Type locality: Valle del Cauca, Tres Esquinas, Habana, Buga. HT F, PT F 1. (Gainesville). Distr.: Colombia (Valle del Cauca). Refs.: Fairchild & Burger, 1994: 73 (cat.). chrysonotus Wilkerson, 1979: 209. Type locality: Colombia, Valle, Rio Sabaletas, 4km E. of Sabaletas. HT F (FSCA). Distr.: Colombia (Valle del Cauca (Rio Sabaletas)). Refs.: Fairchild & Burger, 1994: 73 (cat.); Coscaron & Papavero, 2009: 109 (cat.). detersus (Walker), 1850b: 31 (Tabanus). Type locality: “Colombia”. HT F (BMNH). Distr.: Colombia (Valle del Cauca (Peñas Blancas)), Ecuador. Refs.: Kröber, 1934: 303 (cat. Tabanus (Macrocormus)); Fairchild, 1971: 45 (cat.); Moucha, 1976: 210 (cat.); Bequaert & Renjifo-Salcedo, 1946: 76; Wilkerson, 1979: 211; Fairchild & Burger, 1994: 73 (cat.); Coscaron & Papavero, 2009: 109 (cat.). incipiens (Walker), 1860: 275 (Tabanus). Type locality: Brazil, Amazon. HT F (headless, BMNH). Distr.: Colombia (Antioquia, Meta, Vaupes (La Isla, Rio Apoporis) and unknown location on upper Magdalena River)), Argentina, Brazil, Guatemala to Bolivia. Refs.: Fairchild, 1971: 46 (cat.); Moucha, 1976: 211 (cat.); Wilkerson, 1979: 212; Fairchild & Burger, 1994: 74 (cat.); Coscaron & Papavero, 2009: 110 (cat.). sexannulata Enderlein, 1922: 348 (Styposelaga), nomen nudum. sexannulata Enderlein, 1925: 342 (Styposelaga). luteolineatus Wilkerson, 1979: 214. Type locality: Colombia, Cauca, Guapi. HT F (FSCA). Distr.: Colombia (Cauca). Refs.: Fairchild & Burger, 1994: 74 (cat.); Coscaron & Papavero, 2009: 110 (cat.). maculifrons (Hine), 1907: 222 (Tabanus). Type locality: Guatemala, Puerto Barrios. ST F (OSUC). Distr.: Colombia (Meta (Restrepo; Rio Caney near Restrepo; Villavicencio)), Costa Rica, Honduras, Panama, Trinidad, Venezuela. Refs.: Bequaert & Renjifo-Salcedo, 1946: 78; Fairchild, 1971: 46 (cat.; as syn. of incipiens Walker, 1860); Moucha, 1976: 211 (cat.; as syn. of incipiens Walker, 1860); Coscaron & Papavero, 2009: 110 (cat.). 274 · Zootaxa 4122 (1) © 2016 Magnolia Press E & MIRANDA-ESQUIVEL nigriculus Wilkerson, 1979: 215. Type locality: Colombia, Valle, Rio Sabaletas, 4km E. of Sabaletas. HT F (FSCA). Distr.: Colombia (Valle del Cauca). Refs.: Fairchild & Burger, 1994: 75 (cat.); Coscaron & Papavero, 2009: 110 (cat.). obscurus Kröber, 1929b: 124. Type locality: “Venezuela”. HT F (NMW). Distr.: Colombia (Antioquia, ?Choco, Putumayo (Urcusique), Valle del Cauca (Lago Calima)), Argentina, Brazil, Bolivia, Costa Rica and Panama to Colombia, Ecuador, Peru, Venezuela. Refs.: Fairchild, 1971: 47 (cat.); Moucha, 1976: 211 (cat.); Wilkerson, 1979: 217; Fairchild & Burger, 1994: 75 (cat.); Coscaron & Papavero, 2009: 110 (cat.). constabulorum Fairchild, 1942e: 304. flavofemoratus Kröber, 1929b: 124 (as obscurus var.). sordidatus Fairchild, 1958: 525. Type locality: Panama, San Blás, Rio Tangandi, Camp 2. HT F (MCZ), PT F 1 (FSCA). Distr.: Colombia (Antioquia), Costa Rica, Panama. Refs.: Wilkerson, 1979: 218; Fairchild & Burger, 1994: 76 (cat.); Coscaron & Papavero, 2009: 111 (cat.). wilkersoni Chainey et al., 1999: 121. Type locality: Colombia, Valle, Anchicay, km 80 El Caudal. HT F, PT F 1 (FSCA). Distr.: Colombia (Valle del Cauca). Refs.: Coscaron & Papavero, 2009: 112 (cat.). Subgenus Wilkersonia Fairchild & Burger, 1994 Stenotabanus, subg. Stilbops Wilkerson, 1979: 221. Type species, Stenotabanus (Stilbops) roxannae Wilkerson, 1979 (orig. des.). Stenotabanus, subg. Wilkersonia Fairchild & Burger, 1994: 80 (nom. nov. for Stilbops Wilkerson, 1979). Type species, Stenotabanus (Stilbops) roxannae Wilkerson, 1979 (aut.). Refs.: Coscaron & Papavero, 2009: 112 (cat.). roxannae Wilkerson, 1979: 221 (Stenotabanus (Stilbops)). Type locality: Colombia, Chocó, between Rio San Juan and Rio Baudó. HT F (FSCA). Distr.: Colombia (Chocó). Refs.: Fairchild & Burger, 1994: 80 (cat); Coscaron & Papavero, 2009: 112 (cat.). Genus Rhabdotylus Lutz Rhabdotylus Lutz, 1909b: 29 (1911: 34), nomen nudum. Rhabdotylus Lutz, 1913a: 487 (sep., p. 6) (1914: 166 (sep., p. 6)). Type species, Tabanus planiventris Wiedemann, 1828 (subs. des., Bequaert, 1924: 30). Refs.: Coscaron & Papavero, 2009: 112 (cat.); Turcatel et al., 2010: 2 (rev.); Henriques et al., 2012: 278. Gymnochela Enderlein, 1925: 388. Type species, Tabanus planiventris Wiedemann, 1828 (orig. des.). venenatum (Osten Sacken), 1886: 54 (Tabanus). Type locality: Panama, Volcán de Chiriquí. LT F (BMNH). Distr.: Colombia (Valle del Cauca (Lago Calima, Peñas Blancas)), Guatemala to Ecuador. Refs.: Fairchild, 1971: 77 (cat.; as venenata); Wilkerson, 1979: 312 (as venenata); Fairchild & Burger, 1994: 114 (cat.; as venenata); Coscaron & Papavero, 2009: 113 (cat.). venatum Moucha, 1976: 205 (cat.), error. Genus Stibasoma Schiner, 1867 Subgenus Stibasoma Schiner, 1867 apicimacula Fairchild, 1940b: 686. Type locality: Panama, Coclé, El Valle, 2500 ft. HT F (MCZ). Distr.: Colombia (Valle del Cauca (Rio Raposo, Rio Sabaletas; Anchicaya)), Costa Rica, Panama. Refs.: Lee et al., 1969: 449; Fairchild, 1971: 75 (cat.); Moucha, 1976: 204 (cat.); Wilkerson, 1979: 305; Fairchild & Burger, 1994: 112 (cat.); Coscaron & Papavero, 2009: 113 (cat.); Turcatel et al., 2010: 4 (rev.). chionostigma (Osten Sacken), 1886: 54 (Tabanus). Type locality: Guatemala, Vera Paz, Panama. HT F (BMNH). CATALOGUE OF TABANIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 275 Distr.: Colombia (Cauca (Guapi), Valle del Cauca (Buenaventura)), Mexico to Colombia. Refs.: Bequaert & Renjifo-Salcedo, 1946: 64; Fairchild, 1971: 75 (cat.); Moucha, 1976: 204 (cat.); Wilkerson, 1979: 306; Fairchild & Burger, 1994: 112 (cat.); Coscaron & Papavero, 2009: 113 (cat.): Turcatel et al., 2010: 13 (rev.). pachycephalum Bigot, 1892a: 636. flavistigma Hine, 1912: 516. bifenestrata Philip, 1966: 526. festivum (Wiedemann), 1828:135 (Tabanus). Type locality: “Brazil”. HT F (MNHUB). Distr.: Argentina, Brazil, Colombia (Putumayo (Villagarzón, as Vila Garzon, error), French Guiana, Trinidad and Tobago, Venezuela. Refs. Turcatel et al., 2010: 17 (rev.). mallophoroides Walker, 1857: 123 (Tabanus). mallophoroides Knab, 1913: 410 (Stibasoma). var. mallophoroides var. Fairchild, 1971: 75 dyridophorum Knab, 1913: 410. ssp dyridophorum, Fairchild, 1971: 75. flaviventris (Macquart), 1848: 171 (1848: 11) (Tabanus). Type locality: Brazil, Amazonas, Rio Negro. HT F (BMNH). Distr.: Colombia (Nariño (Barbacoas); Valle del Cauca (Rio Raposo)), Mexico to N. Brazil. Refs.: Lee et al., 1969: 449 (as flaviventre); Fairchild, 1971: 76 (cat.; as flaviventre); Moucha, 1976: 205 (cat.; as flaviventre); Wilkerson, 1979: 308 (as flaviventre); Fairchild & Burger, 1994: 113 (cat.; as flaviventre); Coscaron & Papavero, 2009: 114 (cat.); Turcatel et al., 2010: 20 (rev.). dives Walker, 1848: 166 (Tabanus). dives Knab, 193: 408 (Stibasoma) euglossa Lutz, 1915: 117. flaviventre Macq., which it undoubtedly is”). stilbium Fairchild, 1953: 264. sulfurotaenium Fairchild, 1971: 76 (cat.), misspell. sulfurotaeniatum Kröber, 1931i: 338. ssp. pullum Fairchild & Aitken, 1960: 5 [as pulla,]. sulfureotaeniatum Moucha, 1976: 205 (cat.), error fulvohirtum (Wiedemann), 1828: 155 (Tabanus). Type locality: “Brazil”. LT F (NMW). Distr.: Colombia (Amazonas (La Chorrera); (Boyacá (Guaguaquí); Chocó (La Teresita), Meta (Villavicencio); Putumayo (Mocoa); Valle del Cauca (Rio Raposo)), Bolivia, Brazil, Costa Rica to Trinidad, Ecuador, Guyana, ?Paraguay. Refs.: Kröber, 1934: 263 (cat.); Bequaert & Renjifo-Salcedo, 1946: 64; Lee et al., 1969: 449; Fairchild, 1971: 76 (cat.); Moucha, 1976: 205 (cat.); Wilkerson, 1979: 310; Fairchild & Burger, 1994: 113 (cat.); Coscaron & Papavero, 2009: 114 (cat.); Turcatel et al., 2010: 23 (rev.). compactus Walker, 1854: 222 (Tabanus). dives Walker of Lutz, 1915: 108. fulvihirtum Pechuman, 1942: 54, error. panamense Curran, 1934: 152 (as theotaenia ssp.). Type locality: Panama, Canal Zone, Barro Colorado Island. HT F (AMNH). Distr.: Colombia (Chocó (Curiche)), Honduras to Ecuador, ?Venezuela. Refs.: Fairchild, 1971: 76 (cat.; as panamensis); Moucha, 1976: 205 (cat.; as panamensis); Wilkerson, 1979: 311 (as panamensis); Fairchild & Burger, 1994: 114 (cat.; as panamensis); Coscaron & Papavero, 2009: 115 (cat.). Genus Stypommisa Enderlein Stictotabanus Lutz & Neiva, 1914: 72, 73 (sep., pp. 6, 7), nomen nudum. Refs.: Fairchild, 1951a: 125, 1961a: 244; Coscaron & Papavero, 2009: 115 (cat.). Stypommisa Enderlein, 1923: 545. Type species, Stypommisa punctipennis Enderlein, 1953 (sub. des. Enderlein, 1925: 369) = Stypommisa scythropa (Schiner, 1868). Refs.: Fairchild & Wilkerson, 1986 (rev.); Coscaron & Papavero, 2009: 115 (cat.). 276 · Zootaxa 4122 (1) © 2016 Magnolia Press E & MIRANDA-ESQUIVEL Therioplectes Zeller of Enderlein, 1925: 357, part, misident. Stypommia Enderlein of Kröber, 1929c: 52, 1930a: 248, misident. (part). Sziladya Enderlein of Kröber, 1929c: 118, misident. Sciladynus Kröber, 1930i: 256, error (misident. of Sziladynus Enderlein). Enderleiniella Kröber, 1932: 153 (preocc. Becker 1912). Type species, Therioplecles punctulipennis Enderlein, 1925 (orig. des.) = Stypommisa rubrithorax (Macquart, 1838). Dudaelia Strand, 1932: 142 (nom. nov. for Enderleiniella Kröber, 1932). Type species, Therioplectes punctulipennis Enderlein 1925 (aut.) = Stypommisa rubrithorax (Macquart, 1838). anoriensis Fairchild & Wilkerson, 1986: 11. Type locality: Colombia, Antioquia, Rio Anori valley, La Tirana, 570m. HT F, PT F 16 (FSCA). Distr.: Colombia (Antioquia (Rio Anorí), (Chocó (Rio San Juán-Baudó), Valle del Cauca (Rio Raposo; Buenaventura; Bajo Anchicayá; Rio Bravo Playa Rica; below dam at Lago Calima, Ladrilleros, 30km N. Buenaventura; Rio Sabaletas, 4km E. Sabaletas; Lago Calima, 8km below dam), Antioquia (Rio Anori valley, La Tirana), Cauca)), Ecuador, Peru. Refs.: Fairchild & Burger, 1994: 119 (cat.); Coscaron & Papavero, 2009: 116 (cat.). bipuncta Fairchild in Wilkerson, 1979: 334. Type locality: Colombia, Valle, Rio Frío, La Carmelita. HT F, PT F 3 (FSCA). Distr.: Colombia (Valle del Cauca (Rio Frío, La Carmelita)). Refs.: Fairchild & Wilkerson, 1986: 16; Fairchild & Burger, 1994: 119 (cat.); Coscaron & Papavero, 2009: 116 (cat.). brunnea (Wilkerson), 1979: 208, (Stenotabanus). Type locality: Colombia, Valle, Tres Esquinas, Habana, Buga. HT F, PT F l (FSCA). Distr.: Colombia (Valle del Cauca). Refs.: Fairchild & Burger, 1994: 73 (cat.; Stenotabanus (Stenotabanus)); Coscaron & Papavero, 2009: 116 (cat.). captiroptera (Kröber), 1930: 10, (Tabanus). Type locality: “Venezuela”. HT F (NMW). Distr.: Colombia (Antioquia (Zaragoza, Rio Anorí valley La Tirana), Cauca (Guapi), Chocó (La Teresita; Curiche River, between Rio San Juan and Rio Baudó), Putumayo (Puerto Asis), Valle del Cauca (25km E. Buenaventura, Rio Sabaletas, 4km E. Sabaletas, Bajo Anchicayá, Bajo Calima, 41km E., 20km N. Buenaventura)), Brazil, ?Paraguay, Mexico to Guyana. Refs.: Fairchild, 1971: 81 (cat.); Moucha, 1976: 231 (cat.); Wilkerson, 1979: 335; Fairchild & Wilkerson, 1986: 18; Fairchild & Burger, 1994: 120 (cat.); Coscaron & Papavero, 2009: 116 (cat.). xenium Fairchild, 1947: 568 (Stenotabanus (Stenotabaus)). flavescens Köber of Barretto, 1957: 82 (from Guyana), misident. flavescens (Kröber), 1930i: 249 (Stypommia). Type locality: “Peru”. NT F (ZSBS). Distr.: Colombia (Valle del Cauca (Rio Raposo), Putumayo (Mocoa)), Colombia to Bolivia. Refs.: Lee et al., 1969: 451; Fairchild, 1971: 81 (cat.); Moucha, 1976: 231 (cat.); Fairchild & Wilkerson, 1986: 20; Fairchild & Burger, 1994: 120 (cat.); Coscaron & Papavero, 2009: 116 (cat.). abdominalis Philip, 1960a: 88 (Stypommia). glandicolor (Lutz), 1912: 5 (Tabanus). Type locality: Brazil, Mato Grosso. LT F (IOC). Distr.: Colombia (Meta (Villavicencio)), Costa Rica, Surinam, French Guiana, Peru, Bolivia, Brazil. Refs.: Fairchild & Wilkerson, 1986: 24; Fairchild & Burger, 1994: 120 (cat.); Coscaron & Papavero, 2009: 117 (cat.). hypographa ssp. neofurva Philip, 1969a: 123, (Stoypomisa [sic]). Type locality: Bolivia, Cochabamba, Chapare, Villa Tunari, 500m. HT F (IML). Distr.: Colombia (Putumayo (Mocoa)), Panama to Bolivia. Refs.: Fairchild & Wilkerson, 1986: 25; Fairchild & Burger, 1994: 120 (cat.); Coscaro & Papavero, 2009: 117 (cat.). jaculatrix (Fairchild), 1942e: 309 (Stenotabanus (Stenotabanus)). Type locality: Panama, Canal Zone, Bohio. HT F (MCZ). Distr.: Colombia (Antioquia (Providencia), Chocó (La Teresita; Curiche River)), Guyana, Panama. Refs.: Wilkerson, 1979: 337; Fairchild & Wilkerson, 1986: 27 (as jaculator); Fairchild & Burger, 1994: 120 (cat.; as jaculator); Coscaron & Papavero, 2009: 117 (cat.). maruccii (Fairchild), 1947: 569, (Stenotabanus (Stenotabanus)). Type locality: Panama, San Blás, Pito. HT F CATALOGUE OF TABANIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 277 (MCZ). Distr.: Colombia (Antioquia, Chocó (Curiche River, between Rio San Juan and Rio Baudó), Valle del Cauca (Rio Raposo, 25km E. Buenaventura, Rio Sabaletas, Bajo Anchicayá, Bajo Calima, 10km E. Buenaventura, 41km E. 20km N. Buenaventura)), Nicaragua to Ecuador. Refs.: Lee et al., 1969: 450; Fairchild, 1971: 82 (cat.); Moucha, 1976: 232 (cat.); Wilkerson, 1979: 339; Fairchild & Wilkerson, 1986: 30; Fairchild & Burger, 1994: 121 (cat.); Coscaron & Papavero, 2009: 117 (cat.). frondicolus Fairchild, 1951b: 453 (Stenotabanus) pequeniensis (Fairchild), 1942e: 308, (Stenotabanus (Stenotabanus)). Type locality: Panama, Canal Zone, Rio Pequení, Police Station. HT F (MCZ), PT F l (FSCA). Distr.: Colombia (Antioquia (Rio Anori, SW. Zaragoza; Rio Anori valley, Providencia; La Tirana, Providencia), Cauca (Guapi), Chocó (La Teresita; Curiche; between Rio San Juan and Rio Baudó), Meta (Restrepo), Putumayo (Mocoa; Puerto Asis), Valle del Cauca (Bajo Anchicayá; Lago Calima; Rio Sabaletas; 25km E. Buenaventura; 41km E. 20km N. Buenaventura; Anchicayá dam)), Costa Rica to Bolivia. Refs.: Bequaert & Renjifo-Salcedo, 1946: 76 (Stenotabanus); Fairchild, 1971: 82 (cat.); Moucha, 1976: 232 (cat.); Wilkerson, 1979: 340; Fairchild & Wilkerson, 1986: 35; Fairchild & Burger, 1994: 121 (cat.); Coscaron & Papavero, 2009: 117 (cat.). undet. sp. Wilkerson, 1979: 345. Distr.: Colombia (Antioquia (Rio Anori valley), Chocó (between Rio San Juan and Rio Baudó), Valle del Cauca (8km below dam Lago Calima; Plata Rica; Ladrilleros; Rio Sabaletas, 41km E. 20km N. Buenaventura; Bajo Anchicayá; Buenaventura). Refs.: Coscaron & Papavero, 2009: 118 (cat.). Tribe Tabanini Latreille, 1802 Genus Phorcotabanus Fairchild Stenotabanus, subg. Phorcotabanus Fairchild, 1961b: 441. Type species, Tabanus cinereus Wiedemann, 1821 (orig. des.). Refs.: Coscaron & Papavero, 2009: 119 (cat.). cinereus (Wiedemann), 1821: 84 (Tabanus). Type locality: “South America”. LT F (NMW). Distr.: Colombia (Meta (Villavicencio), Argentina, Bolivia, Brazil, Ecuador, Peru. Refs.: Kröber, 1934: 307 (cat.; Tabanus (Tabanus)); Bequaert & Renjifo-Salcedo, 1946: 75 (Phaeotabanus (Aegialomyia)); Fairchild, 1971: 50 (cat.; Stenotabanus (Phorcotabanus)); Moucha, 1976: 213 (cat.; Stenotabanus (Phorcotabanus)) and 216 (cat.; Dasybasis); Fairchild & Burger, 1994: 128 (cat.); Coscaron & Papavero, 2009: 119 (cat.). Genus Poeciloderas Lutz Poecilosoma Lutz, 1909b: 29 (1913: 487 (1914: 167 (sep., p. 7)) (preocc. Hübner, 1819). Type species, Tabanus quadripunctatus Fabricius, 1805 (subs. des., Bequaert, 1924: 30). Refs.: Coscaron & Papavero, 2009: 120 (cat.). Poeciloderas Lutz, 1921: 15. Type species, Tabanus quadripunctatus Fabricius, 1805 (mon.). Refs.: Coscaron & Papavero, 2009: 120 (cat.). Poecilochlamys Lutz, 1922: 147 (no sp. included). Dasyphyrta Enderlein, 1922: 344 (in key). Hybopelma Enderlein, 1922: 351 (in key). Hypopelma, misspell. Poecilochlamys Lutz in Lutz & Núñez Tovar, 1928: (1955: 61). Hypopelma Kröber, 1929d: 178, error. allusiosis Wilkerson, 1979: 354. Type locality: Colombia, Valle, Peñas Blancas, 1750m, 10km W. of Cali. HT F, PT F 163 (FSCA). Distr.: Colombia (Valle del Cauca (Peñas Blancas; Candelaria; Lago Calima; 3.6km W. Queremal), Cauca, Huila (Finca Merenberg)). Refs.: Coscaron & Papavero, 2009: 120 (cat.). allusiosus Fairchild & Burger, 1994: 126 (cat.), error. quadripunctatus (Fabricius), 1805: 99 (Tabanus; as 4punctatus). Type locality: “South America. Smidt” (i.e., Guyana, between rivers Demerara and Essequibo; cf. Papavero 1971a:21). HT F (ZMKU). Distr.: Colombia 278 · Zootaxa 4122 (1) © 2016 Magnolia Press E & MIRANDA-ESQUIVEL (Amazonas (Leticia), Antioquia (Medellín), Cauca (Guapi), Meta (Villavicencio; Restrepo), Putumayo (Mocoa; Puerto Asis), Valle del Cauca (Peñas Blancas; Cali; Buenaventura; Lago Calima; Tres Esquinas; Habana; Buga; Buenaventura)), Mexico to Argentina. Refs.: Bequaert & Renjifo-Salcedo, 1946: 74 (Hybomitra) (as quadripunctata); Fairchild, 1971: 86 (cat.); Wilkerson, 1979: 356; Fairchild & Burger, 1994: 127 (cat.); Coscaron & Papavero, 2009: 121 (cat.). elegans Thunberg, 1827b: 61 (Tabanus). punctipennis Macquart, 1838a: 185 (1838: 301) (Tabanus). maculipennis Macquart, 1846: 162 (1846: 34) (Tabanus; preocc. Wiedemann, 1828). amabilis Walker, 1848: 154 (Tabanus). nigropunctatus Bellardi, 1859: 67 (Tabanus). habilis Brèthes, 1910: 483 (Tabanus). dasyphyrtina Enderlein, 1925: 356. platyventris Kröber, 1931h: 88 (Tabanus (Poecilosoma)). amabilinus Philip, 1960a: 89 (as quadripunctatus var.). amabilis authors, not Walker. Genus Tabanus Linnaeus Tabanus Linnaeus, 1758: 601. Type species, Tabanus bovinus Linnaeus, 1758 (subs. des., Latreille, 1810: 443). Refs.: Coscaron & Papavero, 2009: 122 (cat.). Bellardia Rondani, 1863: 81 (preocc. Robineau-Desvoidy, 1863). Type species, Tabanus albonotatus Bellardi, 1859 (orig. des.) = Tabanus oculus Walker, 1848. Macrocormus Lutz, 1909b: 29, nomen nudum. Neotabanus Lutz, 1909b: 29, nomen nudum. Macrocormus Lutz, 1911b: 35, nomen nudum. Chelotabanus Lutz, 1913a: 487 (sep., p. 5) (1914: 166 (sep., p. 6)). Macrocormus Lutz, 1913a: 487 (1914: 167 (sep., p. 7)). Neotabanus Lutz, 1913a: 487 (1914a: 167 (sep., p. 7)) (preocc. Ricardo, 1911). Odontotabanus Lutz in Lutz, et al., 1918: 166. Brachypsalidia Enderlein, 1922: 344 (in key). Chelommia Enderlein, 1922: 345 (in key). Hybostraba Enderlein, 1923: 545. Straba Enderlein, 1923: 545. Lophotabanus Szilády, 1926: 25. Bellaria Strand, 1928: 48 (nom. nov. for Bellardia Rondani 1863). Taeniotabanus Kröber, 1930j: 140, nomen nudum. Taeniotabanus Kröber, 1931h: 68. Astigmatophthalmus Kröber, 1931i: 297. Alliomma Borgmeier, 1934: 222 (footnote). albocirculus Hine, 1907: 227. Type locality: Costa Rica, Cartago, Tucurrique. HT F (USNM). Distr.: Colombia (Antioquia (Caucasia), Cauca (Guapi), Caquetá (P.N.N Chiribiquete; Rio Mesay; R. Cuñaré-Amú), Chocó (La Teresita; Curiche River; between Rio San Juan and Rio Baudó; Andagoya; Rio Sucio; Rio Sucio Sautatá), Cundinamarca (Yocaima), Valle del Cauca (Bajo Calima, Rio Raposo, Rio Sabaletas)), Nicaragua, Costa Rica, Panama, Guyana, Ecuador. Refs.: Lee et al., 1969: 452; Fairchild, 1971: 88 (cat.); Moucha, 1976: 87 (cat.); Wilkerson, 1979: 363; Fairchild & Burger, 1994: 130 (cat.); Coscaron & Papavero, 2009: 123 (cat.); Parra-Henao et al., 2008: 182; Pisciotti & Miranda, 2011: 4 (report). albocirculatus Kröber, 1929c: 122, error. obscuriventris Kröber, 1930e: 6, (preocc. Kröber, 1929). obscurigaster Kröber, 1934: 303 (cat.; Tabanus (Macrocormus). amazonensis (Barretto), 1949a: 41 (Chelommia). Type locality: Brazil, Amazonas, Manaus, banks of lower rio Negro. HT F (MZUSP). Distr.: Colombia, Guyana, Brazil. Refs.: Fairchild, 1971: 99 (cat.; as syn. of pellucidus Fabricius 1805); Moucha, 1976: 88 (cat.; as syn. of pellucidus Fabricius 1805); Fairchild & Burger, 1994: 130 (cat.); Coscaron & Papavero, 2009: 123 (cat.). CATALOGUE OF TABANIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 279 albopruinosus Kröber, 1930a: 85 (Tabans (Lophotabanus); preocc. Szilády, 1923) angustifrons Macquart, 1848: 172 (1848: 12). Type locality: French Guiana, Cayenne. HT F (BMNH). Distr.: Colombia (? Vaupés), Venezuela, French Guiana, Peru, Brazil. Refs.: Bequaert & Renjifo-Salcedo, 1946: 80 (Tabanus (Tabanus) alboater Walker); Fairchild, 1971: 98 (cat.; as syn. of pellucidus Fabricius 1805); Moucha, 1976: 88 (cat. ; as syn. of pellucidus Fabricius 1805); Coscaron & Papavero, 2009: 123 (cat.). alboater Walker, 1850b: lxvi. atricornis Bigot, 1892a: 683 (preocc. Meigen, 1838). aniptus Fairchild, 1976: 242. Type locality: W. Colombia, Buenaventura, Rio Raposo. HT F (MCZ), PT F 6 (FSCA). Distr.: Colombia (Cauca (Guapi), Chocó (Curiche River), Valle del Cauca (Rio Raposo; Buenaventura; Guayabal; Rio Cajambre; Ladrilleros; Rio Anchicayá)), Panama, Ecuador. Refs.: Wilkerson, 1979: 365; Fairchild & Burger, 1994: 131 (cat.); Coscaron & Papavero, 2009: 124 (cat.). trivittatus Fabricius of Lee et al., 1969: 455, misident. antarcticus Linnaeus, 1758: 602. Type locality: “America calidiore” (Surinam). Distr.: Colombia (Chocó (Andagoya), Meta (Restrepo; Villavicencio)), Brazil, Surinam to Peru, Trinidad, Venezuela. Refs.: Kröber, 1934: 306 (cat.; Tabanus (Tabanus)); Bequaert & Renjifo-Salcedo, 1946: 81 (Tabanus (Lophotabanus) xipe Kröber); Fairchild, 1971: 89 (cat.); Moucha, 1976: 88 (cat.); Fairchild & Burger, 1994: 131 (cat.); Coscaron & Papavero, 2009: 124 (cat.). surinamensis Kröber, 1929c: 123 (preocc. Macquart 1838). xipe Kröber, 1934: 296 (cat.; Tabanus (Lophotabanus), nom. nov for surinamensis Kröber, 1929 by Coscaron & Papavero, 2009: 124 (cat.)). bigoti Bellardi 1859: 58 (nom. nov. for apicalis Macquart 1847). Type locality: “Mexico”. Distr.: Mexico to Colombia, Venezuela. Refs.: Kröber, 1934: 270 (cat.; Chelommia); Fairchild, 1971: 89 (cat.); Moucha, 1976: 94 (cat.); Fairchild & Burger, 1994: 132 (cat.); Coscaron & Papavero, 2009: 125 (cat.). apicalis Macquart, 1847: 36 (1847: 20) (preocc. Wiedemann 1828). brevihamus Enderlein, 1925: 390 (Ommallia). appretiatus Kröber, 1934: 306 (cat.; Tabanus (Tabanus), nom. nov. for apicalis Macquart, 1847). callosus Macquart, 1848: 171 (1848: 11). Type locality: Brazil, Amazonas, rio Negro. HT F (BMNH). Distr.: Colombia (Amazonas, Vaupés), Peru, Guyana, Brazil. Refs.: Fairchild, 1971: 103 (cat.; as syn. of trivittatus Fabricius, 1805); Moucha, 1976: 97, 148 (cat.; as syn. of trivittatus Fabricius, 1805); Fairchild & Burger, 1994: 132 (cat.); Coscaron & Papavero, 2009: 125 (cat.). cicur Fairchild, 1942c: 160 (Tabanus (Neotabanus)). Type locality: Colombia, Meta, Restrepo. HT F (MCZ), PT F l (FSCA). Distr.: Colombia (Meta (La Unión 12km E. Restrepo)), Ecuador, Guyana, Bolivia, Brazil. Refs.: Bequaert & Renjifo-Salcedo, 1946: 79 (Tabanus (Neotabanus)); Fairchild, 1971: 92 (cat. ; as syn. of dorsiger Wiedemann, 1821); Moucha, 1976: 98, 104 (cat.; as syn. of dorsiger Wiedemann, 1821); Fairchild & Burger, 1994: 133 (cat.); Coscaron & Papavero, 2009: 126 (cat.). claripennis (Bigot), 1892a: 675 (Atylotus). Type locality: “Australia” (in error). HT F (BMNH). Distr.: Colombia (Antioquia (Caucasia), Cauca (Finca Corredor, 20km S. Cali), ?Cordoba (Monteria), Magdalena (Santa Marta), Meta (Barbascal), Valle del Cauca (Candelaria, Finca San Luis; Pance; Peñas Blancas; Cali)), West Indies, Costa Rica to Paraguay, Argentina, Brazil and Chile. Refs.: Bequaert & Renjifo-Salcedo, 1946: 78 (Tabanus (Neotabanus) hookeri Knab) Fairchild, 1971: 90 (cat.); Moucha, 1976: 99 (cat.); Wilkerson, 1979: 367; Fairchild & Burger, 1994: 133 (cat.); Parra-Henao et al., 2008: 182; Coscaron & Papavero, 2009: 126 (cat.). filiolus Williston of Johnson, 1919: 428 (Mexico, Jamaica), misident. ameghinoi Brèthes, 1910: 478. hookeri Knab, 1915: 48. ovalipalpus Enderlein, 1925: 357 (Hybostraba). pygmaeus Enderlein, 1925: 375 (Archiplatius; preocc. Williston, 1887). 280 · Zootaxa 4122 (1) © 2016 Magnolia Press E & MIRANDA-ESQUIVEL flavifrons Szilády, 1926: 22. frontosus Kröber, 1931f: 279 (Tabanus (Agelanius)). ?venezuelensis Kröber, 1931f: 303 (Tabanus (Tabanus)). pulchella Kröber, 1931h: 73, fig. 5 (Tabanus (Hybostraba); preocc. Loew, 1858). pulcher Kröber, 1934: 297 (cat.; Tabanus (Hybostraba), nom. nov. for pulchella Kröber, 1931). pusio Kröber, 1934: 301 (cat.; Tabanus (Agelanius), nom. nov. for pygmaeus Enderlein, 1925 by Coscaron & Papavero, 2009: 127 (cat.)). colombensis Macquart, 1846: 165 (1846: 37). Type locality: “Colombia”. HT F (BMNH). Distr.: Colombia (Meta (Villavicencio; Restrepo), Santander (Puerto Olaya), Vaupés (between Rio San José del Guaviare and Calamar)), U. S. A. (S. Texas) to Trinidad, Brazil, Ecuador, Venezuela. Refs.: Bequaert & Renjifo-Salcedo, 1946: 79 (Tabanus (Neotabanus) amplifrons Kröber); Fairchild, 1971: 90 (cat.); Moucha, 1976: 100 (cat.); Fairchild & Burger, 1994: 134 (cat.); Coscaron & Papavero, 2009: 127 (cat.). truquii Bellardi, 1859: 64. fur Williston, 1901: 261 (preocc. Williston, 1887). amplifrons Kröber, 1933: 354 (Tabanus (Neotabanus)). trilineatus Latreille of Bequaert, 1940b: 450, 452, misident. commixtus Walker, 1860: 273. Type locality: “Mexico”. HT F (BMNH). Distr.: Colombia (Chocó (La Teresita; Curiche River)), S. Mexico to Venezuela, Hispaniola, Trinidad, Martinique. Refs.: Wilkerson, 1979: 369; Fairchild & Burger, 1994: 134 (cat.); Coscaron & Papavero, 2009: 127 (cat.). compactus Walker of Aldrich, 1905: 204, misident. maya Bequaert, 1931: 546. truquii Bellardi of Fairchild, 1971: 103 (cat.), misident crassicornis Wiedemann, 1821: 71. Type locality: “?America”. HT F (ZMKU). Distr.: Brazil, Colombia, French Guiana, Surinam, Venezuela. Refs.: Fairchild, 1971: 98 (cat.; as syn. of pellucidus Fabricius, 1805); Moucha, 1976: 101 (cat.; as syn. of pellucidus Fabricius, 1805); Fairchild & Burger, 1994: 135 (cat.); Coscaron & Papavero, 2009: 128 (cat.). discus Wiedemann, 1828: 123. Type locality: “Brazil”. HT F (MNHUB). Distr.: Colombia (Vaupés (Mitú)), Brazil, Ecuador, French Guiana, Guyana, Surinam, Trinidad, ?Venezuela. Refs.: Bequaert & Renjifo-Salcedo, 1946: 80 (Tabanus (Tabanus)); Barretto, 1957: 85 (Chelotabanus); Fairchild & Burger, 1994: 135 (cat.); Coscaron & Papavero, 2009: 128 (cat.). infuscatipennis Surcouf, 1919: 230, nomen nudum. cervinus Kröber, 1929e: 254. communis Kröber, 1930a: 83 (Tabanus (Lophotabanus)). ?melanocnemis Barretto, 1957: 83 (Chelommia). duckei Fairchild, 1984: 18. Type locality: Venezuela, Amazonas, Mt. Marahuaca, N. slope, Benitez camp. HT F, PT F 5 (FSCA). Distr.: Colombia, Venezuela. Refs.: Fairchild & Burger, 1994: 136 (cat.); Coscaron & Papavero, 2009: 129 (cat.). eldridgei Fairchild, 1973: 327. Type locality: Colombia, Chocó, Curiche river. HT F, PT F 10 (FSCA), PT F 2 (UNHD). Distr.: Colombia (Cauca (Guapi), Chocó (Curiche River), Valle del Cauca (Rio Raposo)), W. coast of South America from N. Colombia to Ecuador. Refs.: Wilkerson, 1979: 371; Fairchild & Burger, 1994: 136 (cat.); Coscaron & Papavero, 2009: 129 (cat.). glaucus Wiedemann, 1819: 42. Type locality: “Brazil”. LT F (NMW). Distr.: E. Bolivia, Brazil, E. Colombia, Peru, Venezuela. Refs.: Fairchild, 1971: 94 (cat.); Moucha, 1976: 111 (cat.); Fairchild & Burger, 1994: 138 (cat.); Coscaron & Papavero, 2009: 131 (cat.). cinerarius Wiedemann, 1828: 121. CATALOGUE OF TABANIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 281 tenens Walker, 1850b: lxv (preocc. Walker, 1850). confligens Walker, 1854: 326 (nom. nov. for tenens Walker, 1850). grandis Kröber, 1931h: 68, fig. l (Tabanus (Straba); preocc. Szilády, 1923). guapiensis Wilkerson, 1979: 373. Type locality: Colombia, Cauca, Guapi HT F, PT F 9 (FSCA). Distr.: Colombia (Cauca (Guapi), Nariño, Valle del Cauca (Ladrilleros; Rio Raposo), Chocó (Curiche river)). Refs.: Fairchild & Burger, 1994: 138 (cat.); Coscaron & Papavero, 2009: 131 (cat.). hirtitibia Walker, 1850: 33. Type locality: “Colombia”. HT F (BMNH). Distr.: Colombia (Caquetá (Florencia), Valle del Cauca (Lago Calima; 41km E. 20km N. Buenaventura; Rio Bravo-Playa Rica; Pance; Peñas Blancas)), Venezuela, Ecuador, Bolivia, Peru. Refs.: Kröber, 1934: 310 (cat.; Tabanus (Tabanus)); Bequaert & RenjifoSalcedo, 1946: 80 (Tabanus (?Tabanus)); Fairchild, 1971: 95 (cat.); Moucha, 1976: 113 (cat.); Wilkerson, 1979: 375; Fairchild & Burger, 1994: 139 (cat.); Coscaron & Papavero, 2009: 132 (cat.). cinnamomeus Schiner, 1868: 90 (preocc. Doleschall 1858). fibulata Enderlein, 1922: 345 (Chelommia), nomen nudum. bogotana Enderlein, 1925: 349 (Dasyommia). fibulata Enderlein, 1925: 382 (Chelommia). hirtibia Fairchild & León, 1986: 117 (Tabanus, (Taeniotabanus), error. howdeni (Philip), 1978: 121, (Hybomitra). Type locality: Mexico, Durango, 105km W. of El Salto, 2700m. HT F (CNC). Distr.: Colombia. Refs.: Fairchild & Burger, 1994: 139 (cat.); Coscaron & Papavero, 2009: 132 (cat.). importunus Wiedemann, 1828: 127. Type locality: “Brazil”. Distr.: Colombia (Magdalena (Aracataca), Meta (Restrepo; El Caibe), Cordoba (Monteria), Tolima (Venadillo)), Bolivia to Brazil, Guyana, Panama, Paraguay, Peru, Trinidad. Refs.: Bequaert & Renjifo-Salcedo, 1946: 80 (Tabanus (Tabanus)); Fairchild, 1971: 95 (cat.); Moucha, 1976: 114 (cat.); Wilkerson, 1979: 376; Fairchild & Burger, 1994: 139 (cat.); Coscaron & Papavero, 2009: 132 (cat.). monogramma Wiedemann, 1828: 150. valterii Macquart, 1838b: 184 (1839: 300). lividus Walker, 1848: 162. albidicollis Macquart, 1850: 336 (1850: 32). semisordidus Walker, 1854: 208. albicollis Hunter, 1901: 139, error. lutzi Kröber, 1934: 296 (cat.; Tabanus (Lophotabanus), nom. nov. for macquarti Kröber, 1930). Type locality: “Venezuela”. Distr.: Colombia, Venezuela. Refs.: Fairchild, 1971: 96 (cat.); Moucha, 1976: 120 (cat.); Fairchild & Burger, 1994: 141 (cat.); Coscaron & Papavero, 2009: 134 (cat.). macquarti Kröber, 1930a: 83 (Tabanus (Lophotabanus); preocc. Schiner 1868)). macquarti Schiner, 1868: 89. Type locality: “South America” (Venezuela). ST F (NMW). Distr.: Colombia (Antioquia, Boyacá (Muzo), Cauca (Rio Micay), Meta (Villavicencio, San José; Restrepo), Putumayo, Santander (Rio Suárez), Valle del Cauca (Rio Raposo), Vaupés), Guatemala, Costa Rica to Peru and Surinam. Refs.: Bequaert & Renjifo-Salcedo, 1946: 72 (Alliomma); Fairchild, 1967b: 248 (Schiner’s ST); Lee et al., 1969: 453; Fairchild, 1971: 96 (cat.); Moucha, 1976: 121 (cat.); Wilkerson, 1979: 377; Fairchild & Burger, 1994: 141 (cat.); Coscaron & Papavero, 2009: 134 (cat.). validus Hine, 1920a: 188 (preocc. Wiedemann, 1828). indescriptus Kröber, 1931j: 50, nomen nudum. fulvohirta Kröber, 1931j: 51 (Gymnochela; as Bigot,i var.; preocc. Wiedemann 1828). ratus Kröber, 1934: 315 (cat.; Tabanus (Tabanus), nom. nov. for validus Hine, 1920 by Coscaron & Papavero, 2009: 134 (cat.)). maculineuris Macquart, 1855: 51 (1855: 31). Type locality: “South America”. HT F (BMNH). Distr.: unknown. 282 · Zootaxa 4122 (1) © 2016 Magnolia Press E & MIRANDA-ESQUIVEL Refs.: Kröber, 1934: 276 (Stypommisa as maculinervis); Fairchild, 1971: 96 (cat.); Moucha, 1976: 121 (cat.); Coscaron & Papavero, 2009: 134 (cat.). maculinervis Hunter, 1901: 141 (cat.). nebulosus De Geer, 1776: 227. Type locality: “Surinam”. HT F (NRS). Distr.: Colombia (Antioquia (Caucasia), Atlántico, Magdalena, Meta, Valle del Cauca)), Argentina, Belize, ?Barbados to Brazil, Trinidad. Refs.: Kröber, 1934: 295 (Tabanus (Lophotabanus) ferrifer Walker); Bequaert & Renjifo-Salcedo, 1946: 80 (Tabanus (Tabanus) ferrifer Walker); Philip, 1952: 312 (De Geer’s HT); Fairchild, 1971: 97 (cat.); Moucha, 1976: 125 (cat.); Fairchild & Burger, 1994: 142 (cat.); Parra-Henao et al., 2008: 182; Coscaron & Papavero, 2009: 135 (cat.). ?surinamensis Macquart, 1838a: 136 (1838: 132). incertus Macquart, 1838a:151 (1838: 147). ferrifer Walker, 1850a: 30. interpunctus Lutz, 1918: 167, nomen nudum. erythraeus Bigot, of Brèthes, 1921: 68, misident druyvesteijni Szilády, 1926: 26. ornativentris Kröber, 1929c: 182 (Hybostraba, in the syn. of druyvesteijni Szilády, 1926) transversalifuscatus Kröber, 1930i: 263 (as synonym of druyvesteijni Szilády, 1926). nubeculipennis Kröber, 1931e: 298. nematocallus Fairchild, 1984: 29. Type locality: Brazil, Amazonas, Manaus (Reserva Adolpho Ducke). HT F (INPA), PT F 14 (FSCA). Distr.: Bolivia, Brazil, Colombia, Guyana, Peru, Venezuela. Refs.: Fairchild & Burger, 1994: 142 (cat.); Coscaron & Papavero, 2009: 135 (cat.). nereus Fairchild, 1943: 446. Type locality: Panama, Canal Zone, Balboa, Thatcher Ferry. HT F (MCZ). Distr.: Colombia (Cauca (Guapi), Chocó (Curiche river)), Ecuador, Pacifíc coast of Panama. Refs.: Wilkerson, 1979: 379; Fairchild & Burger, 1994: 142 (cat.); Coscaron & Papavero, 2009: 135 (cat.). occidentalis Linnaeus, 1758: 602. Type locality: “America meridionalis”. NT F (FSCA). Distr.: Colombia (Amazonas (Caucaya; Tarapacá; Leticia, El Puerto), Antioquia (near Porcecito Rio Porce; El Dos, Turbo; Micuro Rio León; Puerto Berrio), Atlántico (Puerto Colombia; Calamar; Barranquilla; Sabanilla), Boyacá (Muzo), Caquetá (P.N.N. Chiribiquete; Rio Mesay; Rio Sararamano; Rio Cuñaré; Cuñaré-Amú), Chocó (Andagoya; Sautatá; Rio Atrato; Rio Sucio, Tilupa; Rio Sucio, Sautata), Magdalena (Sevilla; Rio Frío; Sierra Nevada de Santa Marta, Guachaca 500m), Meta (Restrepo; Villavicencio), Putumayo (La Tagua; Puerto Limón; San Pedro), Santander (Bocas del Rosario; Puerto Wilches; Barranca; Barrancabermeja), Valle del Cauca (Hacienda La Brisa, Ginebra; Ginebra; Hacienda El Tablazo, Tuluá; Tres Esquinas, Habana Buga; Guayabal, Rio Cajambre; Hacienda La Carmelita, Rio Frío; Hormiguero; Cali; Farallones de Cali), Vaupes (between San José del Guaviare and Calamar)), Mexico to Argentina, Trinidad. HT F lost, Refs.: Kröber, 1934: 294 (Tabanus (Neotabanus)), Bequaert & RenjifoSalcedo, 1946: 77 (Tabanus (Neotabanus) lineola var. carneus Bellardi and T (N) lineola var. plangens Walker); Fairchild, 1971: 108 (cat.); Lee et al., 1969: 453; Fairchild & Burger, 1994: 143 (cat.); Coscaron & Papavero, 2009: 136 (cat.); Pisciotti & Miranda 2011: 10 (report). triceps Thunberg, 1827a: 59. dorsiger Wiedemann, 1821: 83. modestus Wiedemann, 1828: 146. importunus Macquar, 1847: 34 (1847: 18) (preocc. Wiedemann 1828). consequa Walker, 1850b: cxxi. plangens Walker, 1854: 199. dorsovittatus Macquart, 1855: 50 (1855: 30). carneus Bellardi, 1859: 62. appendiculatus Hine, 1906: 22. ochrophilus Lutz, 1914: 46, 49 (sep., pp. 7, 10) (Neotabanus). oculus Walker, 1848: 157. Type locality: “British Honduras” (Belize). ST F (BMNH). Distr.: Colombia (Santander CATALOGUE OF TABANIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 283 (Jazmín, between Puerto Wilches and Bucaramanga)), Mexico to Panama. Refs.: Bequaert & Renjifo-Salcedo, 1946: 81 (Tabanus (Lophotabanus)); Fairchild, 1971: 98 (cat.); Moucha, 1976: 128 (cat.); Fairchild & Burger, 1994: 144 (cat.); Coscaron & Papavero, 2009: 137 (cat.). bipartitus Walker, 1848: 158. albonotatus Bellardi, 1859: 56. olivaceiventris Macquart, 1847: 34 (1847: 18). Type locality: Brazil, Pará. Distr.: Colombia (Antioquia (Caucasia), Cauca (Puerto Tejada), Magdalena (Rio Frío), Valle del Cauca (Tenerife; Palmira; Yumbo)), Panama to Brazil, Trinidad. Refs.: Bequaert & Renjifo-Salcedo, 1946: 80 (Tabanus (Tabanus)); Fairchild, 1971: 98 (cat.); Moucha, 1976: 128 (cat.); Wilkerson, 1979: 380; Fairchild & Burger, 1994: 144 (cat.); Coscaron & Papavero, 2009: 138 (cat.). punctum Rondani, 1848:105. imponens Walker, 1857: 122. pulverulentus Bigot, 1892a: 665 (Atylotus). olivaceiventris Lutz, 1911b: 35, nomen nudum. viridescens Kröber, 1931e: 296. coriarius Kröber, 1934: 296 (cat.; as syn. of imponens Walker, 1857). pellucidus Fabricius, 1805: 97. Type locality: “South America. Smidt” (i.e., Guyana, between rivers Demerara and Essequibo; cf. Papavero, 1971: 21). HT F (ZMKU). Distr.: Brazil, Colombia, Ecuador, Guyana, Peru, Surinam, Venezuela. Refs.: Kröber, 1934: 313 (cat.); Fairchild, 1971: 98 (cat.); Moucha, 1976: 130 (cat.); Fairchild & Burger, 1994: 145 (cat.); Coscaron & Papavero, 2009: 138 (cat.). albibarbis Wiedemann, 1824: 21. ?flavifacies Macquart, 1846: 164 (1846: 36). rufoniger Walker, 1850a: 66. senior Walker, 1850a: 67. melanocnemis Barretto, 1957: 83 (Chelommia). crassicornis Wiedemann of Fairchild, 1967c: 96. pellucidis Contreras-Liochtenberg, 2002: 198, error. perplexus Walker, 1850a: 32. Type locality: “Colombia”. HT F (BMNH). Distr.: Colombia, Ecuador. Refs.: Kröber, 1934: 269 (cat; Dicladocera (Dicladocera) as perplexa); Bequaert & Renjifo-Salcedo, 1946: 70 (Dasychela) (as perplexa); Fairchild, 1971: 99 (cat.); Moucha, 1976: 131 (cat.); Fairchild & Burger, 1994: 145 (cat.); Coscaron & Papavero, 2009: 139 (cat.). peruvianus Macquart, 1848: 173 (1848: 13). Type locality: Ecuador, Quito. HT F (BMNH). Distr.: ?Brazil, Colombia, Ecuador, Venezuela. Refs.: Kröber, 1934: 270 (as peruviana); Bequaert & Renjifo-Salcedo, 1946: 68 (Dichelacera (?Psalidia) peruviana); Fairchild, 1971: 99 (cat.); Moucha, 1976: 131 (cat.); Fairchild & Burger, 1994: 145 (cat.); Coscaron & Papavero, 2009: 139 (cat.). piceiventris Rondani 1848:106. Type locality: “Brazil” (Pará, Belém; cf. Papavero, 1973: 342-343, material collected by Ghiliani). HT F (Naples). Distr.: Bolivia, Brazil, Colombia, Ecuador, French Guiana, Guyana, Peru, Surinam, Trinidad, Venezuela. Refs.: Coscaron & Papavero, 2009: 139 (cat.). basivitta Walker, 1850b: lxviii. viduus Walker, 1850b: lxviii. marginenevris Macquart, 1855: 49 (1855: 29) (Tabanus). bitinctus Walker, 1857: 123. marginenrvis Hunter, 1901: 142 (cat.; Tabanus); error. polyphemus Fairchild, 1958: 527. Type locality: Panama, Panama, Cerro Jefe, La Victoria. HT F (MCZ), PT F 12 (FSCA). Distr.: Mexico (Veracruz) to Colombia. Refs.: Fairchild, 1971: 99 (cat.); Moucha, 1976: 132 (cat.); Fairchild & Burger, 1994: 146 (cat.); Coscaron & Papavero, 2009: 139 (cat.). fumomarginatus Hine of Fairchild, 1951b: 455, misident. 284 · Zootaxa 4122 (1) © 2016 Magnolia Press E & MIRANDA-ESQUIVEL praeteritus Fairchild, 1947: 570. Type locality: Panama, Panama, Chepo. HT F, PT F 1 (MCZ), PT F 1, M 1 (USNM), PT M 1 (AMNH), PT F 2 (FSCA). Distr.: Colombia (Cauca (Guapi), Valle del Cauca (Rio Raposo)), Costa Rica to Colombia. Refs.: Wilkerson, 1979: 382; Fairchild & Burger, 1994: 146 (cat.); Coscaron & Papavero, 2009: 140 (cat.). primitivus Walker, 1848: 177. Type locality: “South America”. HT F (BMNH). Distr.: unknown. Refs.: Kröber, 1934: 303 (cat.; Tabanus (Macrocormus)); Fairchild, 1971: 99 (cat.); Moucha, 1976: 132 (cat.); Fairchild & Burger, 1994: 146 (cat.); Coscaron & Papavero, 2009: 140 (cat.). pseudoculus Fairchild 1942b: 13 (Tabanus (Bellardia)). Type locality: Panama, Panama, Rio Chagres, Moja Pollo. HT F, PT F 3, M 1 (MCZ). Distr.: Colombia (Chocó (La Teresita; Curiche River), Valle del Cauca (Bajo Calima)), Trinidad, Guatemala to Colombia, Venezuela and Ecuador. Refs.: Bequaert & Renjifo-Salcedo, 1946: 81 (Tabanus (Lophotabanus)); Fairchild, 1971: 100 (cat.); Moucha, 1976: 133 (cat.); Wilkerson, 1979: 383; Fairchild & Burger, 1994: 147 (cat.); Coscaron & Papavero, 2009: 140 (cat.). pungens Wiedemann, 1828: 175. Type locality: Uruguay, Montevideo. LT F (NMW). Distr.: Colombia (Amazonas (Santa Sofia Island, Rio Amazonas), Antioquia, Cauca (Finca corredor 40km S. Cali; Guapi), Cordoba (Monteria), Meta (Villavicencio; Restrepo), Tolima (Venadillo), Valle del Cauca (Buenaventura; Rio Raposo; Candelaria; Bugalagrande)), U.S.A. (Texas), Neotropics (except West Indies and Chile), Trinidad. Refs.: Bequaert & RenjifoSalcedo, 1946: 79 (Tabanus (Neotabanus) angustivitta Kröber); Lee et al., 1969: 454; Fairchild, 1971: 100 (cat.); Moucha, 1976: 134 (cat.); Wilkerson, 1979: 384; Fairchild & Burger, 1994: 147 (cat.); Coscaron & Papavero, 2009: 140 (cat.). desertus Walker, 1850b: lxix. nuntius Walker, 1854: 207. univittatus Macquart, 1855: 50 (1855: 30). sallei Bellardi, 1859: 61. propinquus Bellardi, 1859: 65 (preocc. Macquart, 1855). discifer Bigot, 1892a: 684 (preocc. Walker, 1850). angustivitta Kröber, 1929e: 250 (as dorsiger var.) (1933a: 352 (Tabanus (Neotabanus); again as n. var.). ruficornis Kröber, 1931f: 287 (Tabanus (Agelanius); preocc. Fabricius 1775). rufus Kröber, 1931f: 287 (Agelanius; preocc. Scopoli 1763). erythrocerus Kröber, 1934: 300 (cat.; Tabanus (Agelanius), nom. nov. for ruficornis Kröber, 1931, by Coscaron & Papavero, 2009: 141). ruficolor Kröber, 1934: 302 (cat.; Tabanus (Agelanius), nom. nov. for rufus Kröber, 1931, by Coscaron & Papavero, 2009: 141). restrepoensis Fairchild, 1942c: 163. Type locality: Colombia, Meta, Restrepo. HT F (MCZ). Distr.: Colombia (Meta (Restrepo)), Argentina, Bolivia, Ecuador, Guyana, Peru. Refs.: Bequaert & Renjifo-Salcedo, 1946: 78 (Tabanus (Neotabanus)); Fairchild, 1971: 101 (cat.); Moucha, 1976: 135 (cat.); Fairchild & Burger, 1994: 148 (cat.); Coscaron & Papavero, 2009: 141 (cat.). rixator Fairchild, 1943: 448. Type locality: Panama, Panama, Paitilla Point. HT F (MCZ). Distr.: Colombia (Valle del Cauca (Buenaventura, Rio Raposo (Pacific coast)), Ecuador, Panama. Refs.: Lee et al., 1969: 454; Fairchild, 1971: 101 (cat.); Moucha, 1976: 135 (cat.); Fairchild & Burger, 1994: 148 (cat.); Coscaron & Papavero, 2009: 141 (cat.). rubiginipennis Macquart, 1846: 167 (1846: 39). Type locality: “Colombia, temperate region”. HT F (BMNH). Distr.: Colombia (Cauca (Cerro de Munchique), Meta (Restrepo)), Ecuador. Refs.: Kröber, 1934: 269 (cat.; Dicladocera (Dasyrhamphis); Bequaert & Renjifo-Salcedo, 1946: 75 (Hybomitra); Fairchild, 1971: 74 (cat.; Dicladocera); Moucha, 1976: 228 (cat.; Dicladocera); Wilkerson, 1979: 301 (Dicladocera); Fairchild & Burger, 1994: 148 (cat.); Coscaron & Papavero, 2009: 141 (cat.). adustus Walker, 1850a: 34. rubigipennis Moucha, 1976: 136 (cat.). CATALOGUE OF TABANIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 285 rubripes Macquart, 1838a: 138 (1838: 134). Type locality: French Guiana, Cayenne (? in error, type labelled “Brazil. Sylveira” [= Rio de Janeiro; cf. Papavero, 1971: 144-147]). HT F (MNHNP). Distr.: Colombia (Antioquia (Rio Anori valley), Meta (Villavicencio; Restrepo)), Panama to Paraguay. Refs.: Fairchild, 1971: 101 (cat.); Moucha, 1976: 136 (cat.); Wilkerson, 1979: 386; Fairchild & Burger, 1994: 148 (cat.); Coscaron & Papavero, 2009: 142 (cat.). ?derivatus Walker, 1848: 151. pruinosus Kröber, 1929c: 125. pruinicorpus Kröber, 1934: 296 (cat.; Tabanus (Lophotabanus), nom. nov. for pruinosus Kröber, 1929). lophus Philip 1960a: 92 (Tabanus (Lophotabanus)). secundus Walker, 1848: 180. Type locality: unknown. HT F (BMNH). Distr.: Colombia (Caquetá (P.N.N. Chiribiquete, Cuñaré-Amú; Rio Sararamano; Rio Mesay, Rio Cuñaré), Chocó (Rio Sucio, Cocarica), Cauca (Rio Raposo), Meta (Villavicencio)), Guatemala to Peru, Surinam and Paraguay. Refs.: Bequaert & Renjifo-Salcedo, 1946: 78 (Tabanus (Neotabanus) lineola var. stenocephalus Hine); Fairchild, 1971: 92 (cat.; as syn. of dorsiger Wiedemann, 1821); Moucha, 1976: 103, 138 (cat.; as syn. of dorsiger Wiedemann, 1821); Fairchild & Burger, 1994: 149 (cat.); Coscaron & Papavero, 2009: 142 (cat.); Pisciotti & Miranda 2011: 9 (as stenocephalus) (report). stenocephalus Hine 1906: 27. sextriangulus Gorayeb & Rafael, 1984: 49. Type locality: Brazil, Amazonas, Manaus (Campus da Universidade Federal do Amazonas). HT F (MPEG), PT F 24 (FSCA). Distr.: Colombia, Peru, Brazil. Refs.: Fairchild & Burger, 1994: 149 (cat.); Coscaron & Papavero, 2009: 142 (cat.). simplex Walker, 1850a: 34. Type locality: “Colombia”. Distr.: Colombia (Caquetá (Florencia; La Vega; P.N.N. Chiribiquete), Cundinamarca, Huila (Finca Merenberg near Leticia), Valle del Cauca (Peñas Blancas; 10km W Cali)). HT F (BMNH). Refs.: Bequaert & Renjifo-Salcedo, 1946: 82 (Tabanus (?subgenus); Fairchild, 1971: 74 (cat.; Dicladocera); Moucha, 1976: 228 (cat.; Dicladocera); Wilkerson, 1979: 343 (Stypommisa); Fairchild & Wilkerson, 1986: 2; Fairchild & Burger, 1994: 149 (cat.); Coscaron & Papavero, 2009: 142 (cat.); Pisciotti & Miranda 2011: 11 (report). rufiventris Macquart, 1846: 166 (1846: 38) (preocc. Fabricius, 1805). indiorum Bequaert & Renjifo-Salcedo, 1946: 75 (Hybomitra; nom. nov. for rufiventris Macquart, 1846). sorbillans Wiedemann, 1828: 140. Type locality: “Brazil”. HT F (MNHUB). Distr.: Colombia (Boyacá, Casanare, Meta (Restrepo)), Venezuela, Trinidad, to Paraguay, Brazil and Argentina. Refs.: Bequaert & Renjifo-Salcedo, 1946: 82 (Tabanus (Macrocormus)); Fairchild, 1971: 101 (cat.); Moucha, 1976: 141 (cat.); Fairchild & Burger, 1994: 149 (cat.); Coscaron & Papavero, 2009: 143 (cat.). ?fuscus Thunberg, 1827a: 58 (preocc. Wiedemann, 1819). terminus Walker, 1848: 160. ?pseudosorbillans Lutz, 1909b: 29, nomen nudum. sylvestris Brèthes, 1910: 482. flavostriatus Kröber, 1929d: 188 (preocc. Schuurmans Stekhoven 1926). rubrofemorata Kröber, 1931h: 400 (Bellardia). flavolineatus Philip, 1960b: 31 (nom. nov. for flavostriatus Kröber, 1930). undet. sp. Wilkerson, 1979: 393 (Colombia, Boyacá, Muzo, 900m; Chocó, La Teresita; Antioquia: 25Km W. of Zaragoza). Ref.: Coscaron & Papavero, 2009: 143 (cat.). surifer Fairchild, 1964a: 183. Type locality: Panama, Panama, Cerro Campana. HT F (MCZ), PT F 77, M 2 (FSCA). Distr.: Colombia (Antioquia, Boyacá (Muzo), Cauca (Rio Micay; Guapi), Chocó (Rio Nimiquía; La Teresita; Curiche; between Rio San Juan and Rio Baudó), Meta, Valle del Cauca (Bajo Anchicayá; Lago Calima; Bajo Calima; Rio Bravo-Playa Rica; Rio Sabaletas; Ladrilleros; Rio Raposo)), Costa Rica, Ecuador, Panama, Peru. Refs.: Lee et al., 1969: 454; Fairchild, 1971: 102 (cat.; as thimeana ssp.); Moucha, 1976: 145 (cat.; as syn. of thiemeanus Enderlein, 1925); Wilkerson, 1979: 386; Fairchild & Burger, 1994: 150 (cat.); Coscaron & Papavero, 2009: 144 (cat.). bigoti var. A; Fairchild 1943: 442. 286 · Zootaxa 4122 (1) © 2016 Magnolia Press E & MIRANDA-ESQUIVEL thiemeanus (Enderlein), 1925: 389 (Ommalia). Type locality: “Colombia, Cordillera, tierra caliente”. HT F (MNHUB). Distr.: Colombia (Chocó (between Rio San Juan and Rio Baudó), Valle del Cauca (Lago Calima; Rio Bravo-Playa Rica; Rio Sabaletas; 41km E. Buenaventura; Bajo Anchicayá; 3.6km W. Queremal; Rio Raposo; Cali)), Ecuador. Refs.: Kröber, 1934: 270 (cat.; as thiemeana); Bequaert & Renjifo-Salcedo, 1946: 73 (Alliomma) (as thiemeana); Fairchild, 1971: 102 (cat.; as thimeana, error); Moucha, 1976: 146 (cat.; as thimeana, error); Wilkerson, 1979: 389 (as thiemeana, error); Coscaron & Papavero, 2009: 144 (cat.). interrupta Enderlein, 1925: 390 (Ommallia). trivittatus Fabricius, 1805: 104 (as 3-vittatus). Type locality: “South America. Smidt” [i.e., Guyana, between rivers Demerara and Essequibo; cf. Papavero, 1971: 21]. LT F (ZMKU). Distr.: Colombia (Antioquia (El Dos, Turbo), Caquetá (P.N.N. Chiribiquete, Rio Cuñaré-Amú) Meta (Villavicencio), Putumayo (Mocoa; San Pedro) Valle del Cauca (Rio Anchicayà; Rio Raposo; Rio Frío)), Brazil, ?Costa Rica, Guyana, ?Panama, , Surinam. Refs.: Bequaert & Renjfo-Salcedo, 1946: 78 (Tabanus (Neotabanus) fumatipennis Kröber, ) Lee et al., 1969: 455; Fairchild, 1971: 103 (cat.); Moucha, 1976: 148 (cat.); Fairchild & Burger, 1994: 151 (cat.); Coscaron & Papavero, 2009: 145 (cat.); Pisciotti & Miranda, 2011: 13 (report). fumatipennis Kröber, 1933: 341 (Tabanus (Neotabanus)). brunniventris Kröber, 1933: 343 (Tabanus (Neotabanus); as callosus var.; preocc. Schuurmans Stekhoven, 1926). bruniventris Barretto, 1957: 86 (Taeniotabanus), error. unipunctatus (Bigot), 1892a: 663 (Atylotus). Type locality: “South America”. Distr.: Colombia (Chocó (Alto Curiche), Valle del Cauca (Rio Raposo)), Mexico to Colombia. Refs.: Fairchild, 1971: 103 (cat.); Moucha, 1976: 149 (cat.); Wilkerson, 1979: 390; Fairchild & Burger, 1994: 151 (cat.); Coscaron & Papavero, 2009: 145 (cat.). fumomarginatus Hine of Kröber, 1929c: 131 (Tabanus (Lophotabanus)), misident. piraticus Fairchild, 1942b: 15. unistriatus Hine, 1906: 28. Type locality: Costa Rica, San Carlos. HT F (BMNH). Distr.: Colombia (Caquetá (P.N.N. Chiribiquete, Rio Mesay; Rio Sararamano; Puerto Abeja; Rio Cuñaré; Rio Cuñaré-Amú), Cauca (Rio Micay; Guapi), Chocó (between Rio San Juan and Rio Baudó), Valle del Cauca (Rio Raposo; 25Km E. Buenaventura; 41km E., 20km N. Buenaventura; Bajo Anchicayá; Rio Sabaletas)), Guatemala to Ecuador, ?Guyana, ?Peru. Refs.: Lee et al., 1969: 455; Fairchild, 1971: 104 (cat.); Moucha, 1976: 149 (cat.); Wilkerson, 1979: 391; Fairchild & Burger, 1994: 152 (cat.) Coscaron & Papavero, 2009: 146 (cat.); Pisciotti & Miranda 2011: 14 (report). vittiger ssp. guatemalanus Hine, 1906: 24 (as sp.). Type locality: Guatemala, San José. ST F M (OSU). Distr.: U. S. A. (Florida), Bahamas, West Indies (Cuba, Cayman islands, Jamaica, Puerto Rico), SE. Mexico to Surinam, French Guiana and Brazil. Refs.: Fairchild, 1971: 102 (cat.; as subsimilis ssp.); Moucha, 1976: 144 (cat.; as subsimilis ssp.); Fairchild & Burger, 1994: 152 (cat.); Coscaron & Papavero, 2009: 146 (cat.). bellardii Szilády, 1926: 23. truquii Bellardi of Bequaert, 1940: 352 (M only), misident. appendiculatus Hine of Bequaert, 1940a: 451, 453 (M only), misident. carneus Bellardi of Bequaert, 1940a: 451, 453 (F only), misident. caymanicus Fairchild, 1942c: 180 (as vittiger ssp.). angustivitta Kröber of Bequaert & Renjifo-Salcedo, 1946: 79, misident. wilkersoni Fairchild, 1983: 31, (nom. nov. for despectus Fairchild, 1942). Type locality: Brazil, Amazonas, rio Branco, Vista Alegre. Distr.: Brazil, E. Colombia, French Guiana, Peru. Refs.: Fairchild & Burger, 1994: 152 (cat.); Coscaron & Papavero, 2009: 146 (cat.). despectus Fairchild, 1942c: 161 (Tabanus (Neotabanus)); preocc. Kröber, 1930). xuthopogon Fairchild, 1984: 36. Type locality: Peru, Loreto, Iquitos. HT F, PT F 4 (FSCA). Distr.: Brazil, E. Colombia, E. Ecuador, E. Peru. Refs.: Fairchild & Burger, 1994: 153 (cat.); Coscaron & Papavero, 2009: 146 (cat.). CATALOGUE OF TABANIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 287 Acknowledgments To Universidad de Antioquia, Proyecto Sostenibilidad 2014-2015 (MW). TABLE 1. Tabanidae species recorded from Colombia. Subfamily Tribe Genus Chrysopsinae Chrysopsini Chrysops Pangoninae Rhinomyzini Betrechia Pangoniini Esenbeckia Scionini Fidena Subgenus Species author auroguttatus Kröber, 1930 bulbicornis Lutz, 1911 calogaster Schiner, 1868 chiriquensis Fairchild, 1939 incisus Macquart, 1846 laetus Fabricius, 1805 leucospilus Wiedemann, 1828 melaenus Hine,1925 mexicanus Kröber, 1926 nexosus Wilkerson, 1979 renjifoi Bequaert, 1946 reticulatus Wilkerson, 1979 soror Kröber, 1925 varians Widemann, 1828 variegatus (De-Geer, 1776) weberi Bequaert, 1946 ocellata Oldroyd, 1970 balteata Wilkerson, 1979 bella Philip, 1961 cisandeana Wilkerson, & Fairchild, 1983 diaphana (Schiner, 1868) gracilis Kröber, 1931 insignis Kröber, 1931 minuscula Wilkerson, 1979 osornoi Fairchild, 1942 prasiniventris (Macquart, 1846) subvaria (Walker, 1848) testaceiventris (Macquart, 1848) tigrina Wilkerson, 1979 tinctipennis Kröber, 1931 translucens (Macquart, 1846) Proboscoides ecuadorensis ssp. chagresensis Fairchild, 1942 Fidena aureopygia Kröber, 1931 auribarba (Enderlein, 1925) eriomeroides (Lutz, 1909) flavipennis Krober, 1931 Esenbeckia ...Continued on next page 288 · Zootaxa 4122 (1) © 2016 Magnolia Press E & MIRANDA-ESQUIVEL TABLE 1. (Continued) Subfamily Tribe Genus Pityocera Subgenus Species author flavipennis ssp. vallensis Wilkerson, 1979 laterina (Rondani, 1850) ochrapogon Wilkerson, 1979 pubescens (Lutz, 1909) rhinophora (Bellardi, 1859) schildi (Hine, 1925) sulfurea Wilkerson, 1979 Elaphella cervus (Widemann, 1828) Pityocera festai Giglio-Tos, 1896 albifasciata (Macquart, 1846) aurulans (Wiedemann, 1830) Scione aurulans ssp. lurida Enderlein, 1925 Tabaninae Diachlorini Acanthocera brevibeccus Wilkerson, 1979 cuprea Wilkerson, 1979 distincta (Schiner, 1868) equivexans Wilkerson, 1979 flavescens (Enderlein, 1930) fulvosericea (Krober, 1931) incompleta (Macquart, 1846) maculipennis (Schiner, 1868) minor (Macquart, 1947) minuta Szilády, 1926 nigripes (Kröber, 1930) obscurefemorata Kröber, 1930 punctata Szilády, 1926 rhinothrix Wilkerson, 1979 rufescens (Ricardo, 1900) rufipes (Kröber, 1930) serrata Wilkerson, 1979 youngi Wilkerson, 1979 Acanthocera marginalis Walker, 1854 Nothocanthocera albomarginata (Kröber, 1930) trigonifera (Schiner, 1868) Bolbodimyia Catachlorops Amphichlorops bicolor Bigot, 1892 celeroides Stone, 1954 erythrocephala (Bigot, 1892) galindoi Fairchild, 1964 nigra Stone, 1934 philipi Stone, 1954 bogotanus (Enderlein, 1925) vespetinus (Bequaert & RenjifoSalcedo, 1946) ...Continued on next page CATALOGUE OF TABANIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 289 TABLE 1. (Continued) Subfamily Tribe Genus Subgenus Species author Catachlorops alphus Wilkerson, 1979 fumipennis Kröber, 1931 nigripalpis (Macquart, 1846) rufescens (Fabricius, 1805) fulmineus (Hine, 1920) medemi Philip, 1969 auripis (Philip, 1960) difficilis (Kröber, 1931) quadrimaculatus (Macquart, 1846) siculus Wilkerson, 1979 testaceus (Macquart, 1846) fairchildi Wilkerson, 1979 inianis (Fabricius, 1787) leucochlorus Fairchild, 1961 leuconotus Krolow & Henriques, 2010 mexicanus (Linnaeus, 1758) chloroticus (Philip & Fairchild, 1956) unicolor (Wiedemann, 1818) montium (Surcouf, 1919) schineri (Kröber, 1931) limbativena (Enderlein, 1925) ocellus (Walker, 1848) anduzei Stone, 1944 curvipes (Fabricius, 1805) fuscistigma Lutz, 1931 jobbinsi Fairchild, 1942 leticia Wilkerson, & Fairchild, 1982 pechumani Fairchild, 1972 xynus Fairchild, 1972 albitibialis Burger, 1999 transposita Walker,, 1854 burgeri Coscaron & Papavero, 2009 chocoensis Fairchild & Philip, 1960 damicornis (Fabricius, 1805) fasciata Walker, 1850 hartmanni Fairchild & Philip, 1960 marginata Macquart, 1847 melanosoma Hine, 1920 Psalidia Psarochlorops Chlorotabanus Chryptolylus Dasybasis Dasychela Dasychela Diachlorus Dichelacera Desmatochelacera Dichelacera ...Continued on next page 290 · Zootaxa 4122 (1) © 2016 Magnolia Press E & MIRANDA-ESQUIVEL TABLE 1. (Continued) Subfamily Tribe Genus Subgenus Species author regina Fairchild, 1940 submarginata Lutz, 1915 villavoensis Fairchild & Philip, 1960 Idiochelacera subcallosa Fairchild & Philip, 1960 Orthostyloceras aurata Wilkerson, 1979 argentomacula Wilkerson, 1979 basirufa (Walker, 1850) beaveri Wilkerson, 1979 calimaensis Wilkerson, 1979 clara (Schiner, 1868) dalessandroi Wilkerson,1979 distomacula Wilkerson, 1979 hirsuta Wilkerson, 1979 leei Wilkerson, 1979 macula (Macquart, 1846) minos (Schiner, 1868) nigrocoerulea (Rondani, 1850) pruinosa Wilkerson, 1979 riveti (Surcouf, 1919) submacula (Walker, 1850) Hemichrysops fascipennis Kröber, 1930 Himantostylus intermedius Lutz, 1913 Conoposelaga colombiana Fairchild, 1966 Lepiselaga crassipes (Fabricius, 1805) Dicladocera Lepiselaga Leucotabanus Phaeotabanus Philipotabanus Melasmatabanus Mimotabanus canithorax Fairchild, 1941 exaestuans (Linnaeus, 1758) flavinotum (Kröber, 1934) janinae Fairchild, 1970 pauculus Fairchild, 1951 cajennensis (Fabricius, 1787) innotescens (Walker, 1854) nigriflavus (Kröber, 1930) phaeopterus Fairchild, 1964 serenus (Kröber, 1931) criton (Kröber, 1934) fascipennis (Macquart, 1846) nigripennis Wilkerson, 1979 annectans Fairchild, 1975 fucosus Fairchild, 1958 opimus Fairchild, 1975 phalaropygus Fairchild, 1964 ...Continued on next page CATALOGUE OF TABANIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 291 TABLE 1. (Continued) Subfamily Tribe Genus Subgenus Species author plenus (Hine, 1907) porteri Fairchild, 1975 tanypterus Wilkerson, 1979 vulpinus Fairchild, 1975 chrysothrix (Fairchild, 1943) grassator (Fairchild, 1953) magnificus (Kröber, 1934) nigrinubilus (Fairchild, 1953) pterographicus (Fairchild, 1943) tenuifasciatus (Kröber, 1930) Pseudacanthocera brevicornis (Enderlein, 1925) Selasoma tibiale (Fabricius, 1805) Spilotabanus multiguttatus (Kröber, 1930) Brachytabanus longipennis Kröber, 1929 Stenotabanus brunneus Wilkerson, 1979 chrysonotus Wilkerson,1979 detersus (Walker, 1850) incipiens (Walker, 1860) luteolineatus Wilkerson, 1979 maculifrons (Hine, 1907) nigriculus Wilkerson, 1979 obscurus Kröber, 192 sordidatus Fairchild, 1958 wilkersoni Chainey, Hall & Gorayeb, 1999 roxannae Wilkerson, 1979 venenatum (Osten Sacken, 1886) apicimacula Fairchild, 1940 chionostigma (Osten Sacken, 1886) festivum (Wiedemann, 1828) flaviventris (Macquart, 1848) fulvohirtum (Wiedemann, 1828) panamense Curran, 1934 anoriensis Fairchild & Wilkerson, 1986 bipuncta Fairchild & Wilkerson, 1979 brunnea (Wilkerson, 1979) captiroptera (Kröber, 1930) flavescens (Kröber, 1930) glandicolor (Lutz, 1912) hypographa ssp. neofurva Philip, 1969 Philopotabanus Stenotabanus Wilkersonia Rhabdotylus Stibasoma Stypommisa Stibasoma ...Continued on next page 292 · Zootaxa 4122 (1) © 2016 Magnolia Press E & MIRANDA-ESQUIVEL TABLE 1. (Continued) Subfamily Tribe Tabanini Genus Species author jaculatrix (Fairchild, 1942) maruccii (Fairchild, 1947) pequeniensis (Fairchild, 1942) undeter.sp. Wilkerson, 1979 Phorcotabanus cinereus (Wiedemann, 1821) Poeciloderas allusiosis Wilkerson, 1979 quadripunctatus (Fabricius, 1805) albocirculus Hine,1907 amazonensis (Barretto, 1949) angustifrons Macquart, 1848 aniptus Fairchild, 1976 antarcticus Linnaeus, 1758 bigoti Bellardi, 1859 callosus Macquart, 1848 cicur Fairchild, 1942 claripennis (Bigot, 1892) colombensis Macquart, 1846 commixtus Walker, 1860 crassicornis Wiedemann, 1821 discus Wiedemann, 1828 duckei Fairchild, 1984 eldridgei Fairchild, 1973 glaucus Wiedemann, 1819 guapiensis Wilkerson, 1979 hirtitibia Walker, 185 howdeni (Philip, 1978) importunus Wiedemann, 1828 lutzi Kröber, 1934 macquarti Schiner, 1868 maculineuris Macquart, 1855 nebulosus De Greer, 1776 nematocallus Fairchild, 1984 nereus Faichild, 1943 occidentalis Linnaeus, 1758 oculus Walker, 1848 olivaceiventris Macquart, 1847 pellucidus Fabricius, 1805 perplexus Walker, 1850 peruvianus Macquart, 1848 piceiventris Rondani, 1848 polyphemus Rondani, 1848 praeteritus Fairchil, 1947 Tabanus Subgenus ...Continued on next page CATALOGUE OF TABANIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 293 TABLE 1. (Continued) Subfamily Tribe Genus Subgenus Species author primitivus Walker,, 1848 pseudoculus Fairchild, 1942 pungens Wiedemann, 1828 restrepoensis Fairchild, 1942 rixator Fairchild, 1943 rubiginipennis Macquart, 1838 rubripes Macquart, 1838 secundus Walker, 1848 sextriangulus Gorayeb & Rafael, 1984 simplex Walker, 1850 sorbillans Wiedemann, 1828 undet. sp. Wilkerson, 1979 surifer Fairchild, 1964 thiemeanus (Enderlein, 1925) trivittatus Fabricius, 1805 unipunctatus (Bigot, 1892) unistriatus Hine, 1906 vittiger ssp. guatemalanus Hine, 1906 wilkersoni Fairchild, 1983 xuthopogon Fairchild, 1984 References Aldrich, J.M. (1905) A catalogue of North American Diptera. Smithsonian Institution Miscellaneous Collection, 46 (2), 1–680. [Series: Publication (Smithsonian Institution); 1444] Barretto, M.P. (1949a) Estudos sobre tabânidas brasileiros. VII. Gênero “Chelommia” Enderlain, 1922, com as descrições de três novas espécies (Diptera, Tabanidae). Revista Brasileira de Biologia, 9 (1), 39–48. Barretto, M.P. (1949b) Estudos sobre tabânidas brasileiros. X. Conoposelaga, nov. gên. (Diptera, Tabanidae). Anais da Faculdade de Medicina da Universidade de São Paulo, 24, 87–93. Barretto, M.P. (1957) Tabânidas da Guiana Inglêsa, com as descrições de seis novas espécies da fauna guianoamazônica (Diptera, Tabanidae). Revista Brasileira de Malariologia e Doenças Tropicais, 8 (l), 75–89. [“1956”] Bellardi, L. (1859) Saggio di Ditterologia Messicana. Vol. 1. Dalla Stamperia Reale, Torino, 264 pp. [pp. 46–76] Bequaert, J. (1924) Notes upon Surcouf´s treatment of the Tabanidae in the Genera Insectorum and upon Enderlein’s proposed new classification of this family. Psyche, 31, 24–40. http://dx.doi.org/10.1155/1924/84890 Bequaert, J. (1926) Medical report of the Hamilton Rice Seventh Expedition to the Amazon, in conjunction with the Department of Tropical Medicine of Harvard University, 1924–1925. Part II. Medical and economic entomology. Tabanidae. Contributions from the Harvard Institute for Tropical Biology and Medicine, 4, 155–257. Bequaert, J. (1931) Tabanidae of the Peninsula of Yucatan, Mexico, with descriptions of new species. Journal of The New York Entomological Society, 39 (4), 533–553. Bequaert, J. (1940a) The Tabanidae of the Antilles. Revista de Entomologia, 11 (1–2), 253–369. Bequaert, J. (1940b) Tabanidae of the Island of Trinidad, B.W. I. Bulletin of Entomological Research, 30 (4), 447–453. http://dx.doi.org/10.1017/S0007485300040621 Bequaert, J. (1944) Further studies of the Tabanidae of Trinidad, B. W. I. Psyche, 51, 12–21. http://dx.doi.org/10.1155/1944/71329 Bequaert, J. (1946a) Descriptions of three new Neotropical species of Chrysops (Diptera, Tabanidae). Psyche, 53 (1–2), 6–12. http://dx.doi.org/10.1155/1946/54672 Bequaert, J.C. (1946b) Appendix A new Agelanius from Colombia. Psyche, 4, 86–88. 294 · Zootaxa 4122 (1) © 2016 Magnolia Press E & MIRANDA-ESQUIVEL Bequaert, J. & Renjifo-Salcedo, S. (1946) The Tabanidae of Colombia (Diptera). Psyche, 53 (3–4), 52–86. http://dx.doi.org/10.1155/1946/40437 Bernardi, N. (1975) Notas sobre tabanídeos americanos I. Sobre o gênero Pseudacanthocera Lutz, com descrição de uma espécie nova. Revista Brasileira de Entomologia, 19 (2), 73–77. Bezzi, M. (1913) Einige alte und neue Namen bei den Dipteren. Societas Entomologica, 28 (13), 56. Bigot, J.M.F. (1892a) Descriptions de diptères nouveaux ou peu connus. Tabanidi. Mémoires de la Société Zoologique de France, 5, 602–692. Bigot, J.M.F. (1892b) Nova genera Dipterorum. Wiener entomologische Zeitung, 11, 161–162. Blanchard, E. (1840) Histoire naturelle des insectes. Orthoptères, névroptères, hémiptères, hyménoptères, lepidoptères et diptères. Duménil, P. (Ed.), Paris, 672 pp. http://dx.doi.org/10.5962/bhl.title.59226 Borgmeier, T. (1933) A proposito da nomenclatura dos Tabanidae da região neotropica. Revista de Entomologia, 3 (3), 286–303. Borgmeier, T. (1934) [Footnote to Kröber’s catalogue of Tabanidae]. Revista de Entomologia, 4, 222. Brèthes, J. (1910) Dípteros nuevos ó poco conocidos de Sud-América. Anales del Museo Nacional de Historia Natural de Buenos Aires, 20, 469–484. Burger, J.F. (1999) Notes on and descriptions of new and little-known species of Neotropical Tabanidae (Diptera). Memoirs on Entomology, International, 14, 51–74. Chainey, J.E., Hall, M.J.R. & Gorayeb, I.S. (1999) Stenotabanus Lutz (Diptera: Tabanidae): an identification guide to the South American species. Memoirs on Entomology, International, 14, 75–134. Contreras-Lichtenberg, R. (2002) On the history of the Diptera collection at the Museum of Natural History in Vienna. Annales Zoologici, 52 (2), 195–200. Coquillett, D.W. (1910) The Type species of the North American genera of Diptera. Proccedings of the United State National Museum, 37, 499–647. Coscarón, S. (1979) Notas sobre tabánidos argentinos. El género Chrysops Meigen. 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Mitteilugen der Deutsche Entomologischen Gesellschaft, 1923 (5), 544–545. Enderlein, G. (1925) Studien an blutsaugenden Insekten. I. Grundlagen eines neuen Systems der Tabaniden. Mitteilungen aus dem Zoologischen Museum in Berlin, 11 (2), 255–409. Enderlein, G. (1930) Dipterologische Studien XX. Tabanidae. Deutsche Entomologische Zeitschrift, 1930, 65–66. Fabricius, J.C. (1775) Systema entomologiae, sistens insectorvm classes, ordines, genera, species adiectis synonymis, loci descriptionibvs, observationibvs. Flensburgi et Lispiae [= Flensburg and Leipzig] Kortii, 832 pp. http://dx.doi.org/10.5962/bhl.title.36510 Fabricius, J.C. (1787) Mantissa insectorum sistens species nuper detectas, Tome 1. Hafniae, apud Proft et Storch, Copenhagen, 383 pp. Fabricius, J.C. (1794) Entomologiae Systematicae Emendata et Aucta. Secundum. Classes, Ordines, Genera, Species. Volume 4. Hafniae [=Copenhagen], 472 pp. Fabricius, J.C. (1805) Systema antliatorum secundum ordines, genera, species adiectis synonymis, locis, observationibus, descriptionibus. C. Reichard, Brunsvigae [= Brunswick], xiv + 15–372 + [1 (errata)] + 30 pp. Fairchild, G.B. (1939) Two new species of Chrysops (Diptera: Tabanidae) from Panama. Proceedings of the Entomological Society of Washington, 41 (9), 257–260. Fairchild, G.B. (1940a) Notes on Tabanidae (Diptera) from Panama. I. The genera Chlorotabanus and Cryptotylus. Revista de Entomologia, 11 (3), 713–723. Fairchild, G.B. (1940b) Notes on Tabanidae (Diptera) from Panama. II. The genus Dichelacera Macquart and related genera. Annals of the Entomological Society of America, 33 (4), 683–700. http://dx.doi.org/10.1093/aesa/33.4.683 Fairchild, G.B. (1941a) Notes on Tabanidae (Diptera) from Panama. IV. The genus Leucotabanus Ad. Lutz. Annals of the Entomological Society of America, 34 (3), 629–638. http://dx.doi.org/10.1093/aesa/34.3.629 CATALOGUE OF TABANIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 295 Fairchild, G.B. (1941b) Notes on Tabanidae (Diptera) from Panama. VI. The genus Fidena Walker. Annals of the Entomological Society of America, 34 (3), 639–646, 6 figs. Fairchild, G.B. (1942a) Notes on Tabanidae (Diptera) from Panama. III. The genus Chrysops Meigen. Proceedings of the Entomological Society of Washington, 44 (1), 1–8. Fairchild, G.B. (1942b) Notes on Tabanidae (Diptera) from Panama. V. The genus Tabanus, subgenus Bellardia Rondani. Psyche, 49 (1–2), 8–17. http://dx.doi.org/10.1155/1942/42581 Fairchild, G.B. (1942c) Notes on Tabanidae (Diptera) from Panama. VII. The subgenus Neotabanus Ad. Lutz. Annals of the Entomological Society of America, 35 (2), 153–182. http://dx.doi.org/10.1093/aesa/35.2.153 Fairchild, G.B. (1942d) Notes on Tabanidae (Diptera) from Panama. VIII. The genera Pityocera, Scione and Esenbeckia. Annals of the Entomological Society of America, 35 (2), 183–199. http://dx.doi.org/10.1093/aesa/35.2.183 Fairchild, G.B. (1942e) Notes on Tabanidae (Diptera) from Panama. IX. The genera Stenotabanus Lutz., Lepiselaga Macquart and related genera. Annals of the Entomological Society of America, 35 (3), 289–309. http://dx.doi.org/10.1093/aesa/35.3.289 Fairchild, G.B. (1943) Notes on Tabanidae (Diptera) from Panama. X. The genus Tabanus Linneus, and resumé of the Tabanidae of Panama. Annals of the Entomological Society of America, 35 (4), 441–474. [1942] Fairchild, G.B. (1947) Additional notes on the Tabanidae of Panama. Annals of the Entomological Society of America, 39 (4), 564–575. [1946] Fairchild, G.B. (1951a) The generic names for Tabanidae proposed by Adolfo Lutz. Psyche, 57 (4), 117–127. [1950] Fairchild, G.B. (1951b) Descriptions and notes on Neotropical Tabanidae. Annals of the Entomological Society of America, 44 (3), 441–462. http://dx.doi.org/10.1093/aesa/44.3.441 Fairchild, G.B. (1953) Notes on Neotropical Tabanidae (Diptera) with descriptions of new species. Annals of the Entomological Society of America, 46 (2), 259–280. http://dx.doi.org/10.1093/aesa/46.2.259 Fairchild, G.B. (1956) Synonymical notes on Neotropical flies of the family Tabanidae (Diptera). Smithsonian Miscellaneous Collections, 131 (3), 1–38. Fairchild, G.B. (1958) Notes on Neotropical Tabanidae. II. Descriptions of new species and new records for Panama. Annals of the Entomological Society of America, 51 (6), 517–530. http://dx.doi.org/10.1093/aesa/51.6.517 Fairchild, G.B. (1961a) The Adolfo Lutz, collection of Tabanidae (Diptera). I. The described genera and species, condition of the collection, and selection of lectotypes. Memórias do Instituto Oswaldo Cruz, 59 (2), 185–249. http://dx.doi.org/10.1590/S0074-02761961000200006 Fairchild, G.B. (1961b) The Adolfo Lutz, collection of Tabanidae (Diptera). II. Status of the names published without description. Memórias do Instituto Oswaldo Cruz, 59 (3), 278–295. http://dx.doi.org/10.1590/S0074-02761961000300003 Fairchild, G.B. (1964a) Notes on Neotropical Tabanidae (Diptera). IV. Further new species and new records for Panama. Journal of Medical Entomology, 1 (2), 169–185. http://dx.doi.org/10.1093/jmedent/1.2.169 Fairchild, G.B. (1964b) A new genus and species of Neotropical horsefly (Diptera: Tabanidae). Psyche, 70 (4), 193–196, 3 figs. Fairchild, G.B. (1966) Notes on Neotropical Tabanidae (Diptera). VI. A new species of Lepiselaga Macquart with remarks on related genera. Psyche, 72 (3), 210–217. Fairchild, G.B. (1967a) Notes on Neotropical Tabanidae (Diptera). IX. The species described by Otto Kröber. Studia Entomologica, 9 (1–4), 329–379. Fairchild, G.B. (1967b) Notes on Neotropical Tabanidae (Diptera). X. The species described by Schiner, J.R. and others. 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Walker, F. (1848) List of the specimens of dipterous insects in the collection of the British Museum. Vol. 1. London, 229 pp. Walker, F. (1850a) Diptera. Part I. In: Saunders, W.W. (Eds.), Insecta Saundersiana, or characters of undescribed insects in the collection of W.W. Saunders,Esq.,f.r.s., f.l.s., &c. Vol. l. VanVoorst, London, pp. 1–76. Walker, F. (1850b) Characters of undescribed Diptera in the British Museum. Newman’s Zoologist, 8 (Appendix), [Art. l]: lxv–lxxii, [Art. 5] xcv–xcix, [Art. 15] cxxi–cxxii. Walker, F. (1854–1855) List of the specimens of dipterous insects in the collection of the British Museum. 5 (Supplement 1) & 6 (Supplement 2), London, 506 pp. [pp. 1–330 (1854), pp. 331–506 (1855)] Walker, F. (1857–1858) Characters of undescribed Diptera in the collection of W.W. Saunders. Transactions of the Entomological Society of London, New Series, Series 2, 4, 119–158 (1857); 190–235 (1858). Walker, F. (1860) Characters of undescribed Diptera in the collection of W.W. Saunders. Transactions of the Entomological Society of London, New Series, 5, 268–296. Wiedemann, C.R.W. (1819) Brasilianische Zweiflügler. (Wiedemann’s). Zoological Magazine, l (3), 40–56. Wiedemann, C.R.W. (1821) Diptera exotica. Pars 1. Tabulis aeneis duabus. 2nd Edition. (No publisher), Kiliae [= Kiel], xix + 224 pp., 2 pls. http://dx.doi.org/10.5962/bhl.title.14404 Wiedemann, C.R.W. (1824) Munus rectoris in Academia Christiana Albertina aditurus analecta entomologica ex Museo Regio Havniens: maxime congesta profert iconibusque illustrat. Kiliae e regio typographeo scholarum, Kiel, 60 pp. http://dx.doi.org/10.5962/bhl.title.77322 Wiedemann, C.R.W. (1828) Aussereuropäische Zweiflügelige Insekten. Vol 1. Schulzischen, Hamm, xxxii + 608 pp. Wiedemann, C.R.W. (1830) Aussereuropäische zweiflügelige Insekten. 2. Schulzischen, Hamm, xii + 684 pp. Williston, S.W. (1895) Exotic Tabanidae. Kansas University Quarterly, 3, 184–194. Wilkerson, R.C. (1979) Horse flies (Diptera: Tabanidae) of the Colombian departments of Chocó, Valle and Cauca. Cespedesia, 8 (31–35), 87–433. Wilkerson, R.C. (1981) Two new species of Dichelacera (Nothocanthocera) Fairchild, with a key to the species of the subgenus (Diptera: Tabanidae). Proceedings of the Entomological Society of Washington, 83 (1), 64–71. Wilkerson, R.C. & Fairchild, G.B. (1982) Five new species of Diachlorus (Diptera: Tabanidae) from South America, with a revised key to species and new locality records. Proceedings of the Entomological Society of Washington, 84 (3), 636–650. Wilkerson, R.C. & Fairchild, G.B. (1983) A review of the South American species of Esenbeckia, subgenus Esenbeckia (Diptera: Tabanidae). Journal of Natural History, 17, 519–567. http://dx.doi.org/10.1080/00222938300770451 CATALOGUE OF TABANIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 301 Zootaxa 4122 (1): 302–305 http://www.mapress.com/j/zt/ Copyright © 2016 Magnolia Press Article ISSN 1175-5326 (print edition) ZOOTAXA ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.4122.1.24 http://zoobank.org/urn:lsid:zoobank.org:pub:4D2563CA-13BA-4374-A124-455A73F28806 FAMILY XYLOPHAGIDAE DIEGO AGUILAR FACHIN Departamento de Biologia, Faculdade de Filosofia, Ciências e Letras de Ribeirão Preto, Universidade de São Paulo, Av. Bandeirantes, 3900, 14040–901, Ribeirão Preto, SP, Brazil. E–mail: diegoafachin@gmail.com Abstract This family is recorded for the first time to Colombia, with only one undescribed species of genus Rachicerus. Key words: Catalogue, Colombia, diversity, Rachicerus, Xylophagidae Introduction Xylophagidae Fallén, 1810 are a small family of lower Brachycera infraorder Xylophagomorpha (Woodley 1989, Woodley et al. 2009), with a widespread distribution in the world, except in the Afrotropical Region. These flies have body length ranging from 2 to 25 mm and are easily recognized by the antennal flagellum composed of 7–8 or more than 15 flagellomeres, tibial spurs 1–2–2, pulvilliform empodium, the unmodified wing venation—M3 can be open or closed at wing margin—and by the lack of a scale-like elevation behind the metathoracic spiracle (Woodley 2009). Additionally, Rachicerus Walker, 1854, the largest genus, is recognized by the pectinate antennal flagellum with more than eight flagellomeres. Immature stages of xylophagid species are known mainly for Coenomyia Latreille, 1797, Rachicerus and Xylophagus, Meigen, 1803, where they are frequently found under bark of fallen tree (Xylophagus) or associated with decaying wood (Rachicerus), as predators of other insect larvae (James 1981, Woodley 2009). In the neotropics, the immature stages are still poorly known with only the description of pupa of Heterostomus curvipalpis Bigot, 1857 (Coscarón et al. 2013). The adults usually are found on vegetation, particularly near water or fallen trees (Carrera 1945, James 1981, Nagatomi 1982). There are 133 described species of Xylophagidae in the world, in nine genera (Woodley 2011), with most of them into two genera, Rachicerus (69 species) and Xylophagus (24 species). The richest regions are the Palearctic and the Oriental Region, with 46 and 62 described species, respectively (Woodley 2011). Only eleven species in three genera have been described to the Neotropical Region (Woodley 2011), of which six species were described to Brazil—all of them by Carrera (1940, 1945)—and other five species described for different countries (Table 1). Xylophagus vetustus Walker, 1854 from Brazil was stated by Woodley (2011) as a Therevidae species. Additionally, one undescribed species of Rachicerus is recorded to Colombia, as stated in this work (see below). The catalog records only one undescribed species of Rachicerus (Table 2). The family Xylophagidae, as well as the genus Rachicerus are recorded for the first time to Colombia. The recorded species and the classification adopted were based mainly the on information of the world catalogue of the family (Woodley 2011). Additional literature and material belonging to the Instituto de Investigación de Recursos Biológicos Alexander von Humboldt (IAvH) also were consulted. The geopolitical departments are underline and specific localities are in brackets. 302 Accepted by S. Nihei: 10 Apr. 2015; published: 14 Jun. 2016 TABLE 1. Checklist of Neotropical species of Xylophagidae, with known gender and geographic data. Species Gender Geographical distribution Male Female x x Chile Rachicerus bellus Osten Sacken - x Panama Rachicerus lanei Carrera x - Brazil Rachicerus lopesi Carrera - x Brazil Rachicerus marcusi Carrera - x Brazil Rachicerus nigripalpus Loew - x Mexico Rachicerus oliverioi Carrera - x Brazil Rachicerus picticornis Kertész x - Costa Rica Rachicerus shannoni Carrera - x Brazil Rachicerus varipes Loew x x Cuba Xylophagus vetustus Walker - x Brazil Heterostomus curvipalpis Bigot Acronyms used for the depositories IAvH—Instituto de Investigación de Recursos Biológicos Alexander von Humboldt List of abbreviations des.—designation distr.—geographic distribution emend.—emended m.—meters mon.—monotypy, monotype syn. nov.—new synonym orig. des.—by original designation sp.—species subs. des.—subsequent designation syn.—synonym M (m#)—male Catalogue of Xylophagidae of Colombia Family Xylophagidae Fallén, 1810 Genus Rachicerus Walker, 1854 Rachicerus Walker, 1848: 124. Nomen nudum. Attributed to Haliday. Rachicerus Walker, 1854: 103. Type species, Rachicerus fulvicollis Walker, 1854 (mon.). Rhyphomorpha Walker, 1861: 275. Type species, Rhyphomorpha bilinea Walker, 1861 (mon.). Antidoxion Snellen van Vollenhoven, 1863: 6. Type species, Antidoxion fulvicorne Snellen van Vollenhoven, 1863 (mon.). Refs.—Osten Sacken, 1881: 409 (syn.); Woodley, 2011: 476 (cat.). Rhachicerus Loew, 1872: 114, emend. Ref.—Woodley, 2011: 476 (cat.). Antidoxia Marschall, 1873: 321, error. Ref.—Woodley, 2011: 476 (cat.). Rhacicerus Malloch, 1917: 346, error. Ref.—Woodley, 2011: 476 (cat.). Rhachicerella Enderlein, 1921: 167. Type species, Rachicerus honestus Osten Sacken, 1877 (orig. des.). Refs.—Nagatomi, 1970: 418 (syn.); Woodley, 2011: 476 (cat.). CATALOGUE OF XYLOPHAGIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 303 Rachicerella Carrera, 1945: 120, error. Ref.—Woodley, 2011: 476 (cat.). Gymnorhachicerus Frey, 1954: 7. Type species, Gymnorhachicerus pilosus Frey, 1954, (orig. des.). Ref.—Woodley, 2011: 468 (cat., syn.). undescribed sp. Distr.—Colombia (Chocó (PNN Los Katios, Centro Administrativo Sautatá, Afuera del Bosque)). Material examined. COLOMBIA: Chocó: PNN Los Katios, Centro Administrativo Sautatá, Afuera del Bosque, 07º51’N, 77º08’W, 30 m, 1 M (m#), 29.v–13.vi.2003, P. Lopéz Leg. M.3757, Malaise 4 [IAvH]. Acknowlegdments This study was supported by FAPESP Grants 2011/14472-6, 2013/02824-0 and 2014/05793-1. TABLE 2. Xylophagidae species recorded to Colombia. Genus Species Author Rachicerus sp. Walker, 1854 References Carrera, M. (1940) Novas espécies do gênero Rachicerus Walker, 1854 (Dipt.). Arquivos de Zoologia do Estado de São Paulo, São Paulo, 1, 387–394. Carrera, M. (1945) Contribuição ao conhecimento do gênero Rachicerus Walker, 1854 (Diptera, Rachiceridae Handlirsch, 1908). Papéis Avulsos do Departamento de Zoologia, 5, 119–134. [São Paulo] Coscarón, S., Coscarón, M.D.C. & Gil-Azevedo, L.H. (2013) On the enigmatic Heterostomus curvipalpis Bigot, 1857, with a description of the pupa (Diptera, Brachycera). Zootaxa, 3616 (3), 268–276. http://dx.doi.org/10.11646/zootaxa.3616.3.4 Enderlein, G. (1921) Über die phyletisch älteren Stratiomyiiden subfamilien (Xylophaginae, Chiromyzinae, Solvinae, Beridinae, und Coenomyiinae). Mitteilungen aus dem Zoolo-gischen Museum in Berlin, 10 (1), 151–214. Fallén, C.F. (1810) Specimen entomologicum novam Diptera disponendi methodum exhibens. Berlingianis, Lundae [Lund], 26 pp. Frey, R. (1954) Studien über ostasiatische Dipteren. III. Rhachiceridae, Rhagionidae, Hilarimorphidae. Notulae Entomologicae, 34 (1), 1–25. James, M.T. (1981) Xylophagidae [Chapter] 34. In: McAlpine, J.F., Peterson, B.V., Shewell, G.E., Teskey, H.J., Vockeroth, J.R. & Wood, D.M. (Eds.), Manual of Neartic Diptera. Vol. 1. Research Branch Monograph 27, Agriculture Canada, Ottawa, i–vi, pp. 489–492. Loew, H. (1872) Diptera Americae septentrionalis indigena. Centuria decima. Berliner Entomologische Zeitschrift, 16 (1), 49–124. http://dx.doi.org/10.1002/mmnd.18720160110 Malloch, J.R. (1917) A preliminary classification of Diptera, exclusive of Pupipara, based upon larval and pupal characters, with keys to imagines in certain families. Part I. Bulletin of the Illinois State Laboratory of Natural History, 12 (3), i–vi, 161–409. http://dx.doi.org/10.5962/bhl.title.9383 Marschall, A.F. (1873) Nomenclator zoologicus continens nomina systematica generum animalium tam viventium quam fossilium, secundum ordinem alphabeticum disposita sub auspiciis et sumptibus C. R. Societatis ZoologicoBotanicae. Caroli Ueberreuter (M. Salzer), Vindobonae [=Vienna]. I–IV, [2], 482 pp. Nagatomi, A. (1970) Rachiceridae (Diptera) from the Oriental and Palaearctic Regions. Pacific Insects, 12 (2), 417–466. Nagatomi, A. (1982) Notes on Rachiceridae (Diptera). Memoirs of the Kagoshima University Research Center for the South Pacific, 3 (1), 39–65. [Kagoshima] Osten Sacken, C.R. (1877) Western Diptera: Descriptions of new genera and species of Diptera from the region west of the Mississippi and especially from California. Bulletin of the United States Geological and Geographical Survey of the Territories, 3 (2), 189–354. http://dx.doi.org/10.5962/bhl.title.57939 304 · Zootaxa 4122 (1) © 2016 Magnolia Press FACHIN Osten Sacken, C.R. (1881) Enumeration of the Diptera of the Malay Archipelago collected by Prof. Odoardo Beccari, M.r L. M. d’Albertis and others. Annali del Museo Civico di Storia Naturale di Genova, 16, 393–492. Snellen van Vollenhoven, S.C. (1863) Over eene merkwaardige soort van tweevleugelig insect, Antidoxion fulvicorne. Verslagen en Mededeelingen der Koninklijke Akademie van Weten-schappen. Afdeeling Natuurkunde, 15, 1–7. Walker, F. (1848) List of the specimens of dipterous insects in the collection of the British Museum. Vol. 4. Part I. British Museum, London, 229 pp. Walker, F. (1854) List of the specimens of dipterous insects in the collection of the British Museum. Vol. 6. Part V. Supplement I. British Museum, London, 1330 pp. Walker, F. (1861) Catalogue of the dipterous insects collected in Batchian, Kaisaa and Makian, and at Tidon in Celebes, by Mr. A. R. Wallace, with descriptions of new species. Journal of the Proceedings of the Linnean Society, 5 (20), 270–303. http://dx.doi.org/10.1111/j.1096-3642.1861.tb02237.x Woodley, N.E. (1989) Phylogeny and classification of the “orthorrhaphous” Brachycera. In: McAlpine, J.F., Peterson, B.V., Shewell, G.E., Teskey, H.J., Vockeroth, J.R. & Wood, D.M. (Eds.), Manual of Neartic Diptera. Vol. 3. Research Branch Monograph 32, Agriculture Canada, Ottawa, pp. 1371–1395. Woodley, N.E. (2009) Xylophagidae [Chapter] 35. In: Brown, B.V., Borkent, A., Cumming, J.M., Wood, D.M., Woodley, N.E. & Zumbado, M.A. (Eds.), Manual of Central American Diptera. Vol. 1. NCR Research Press, Boca Raton, Ottawa, pp. 509–512. Woodley, N.E. (2011) A World Catalog of the Xylophagidae (Insecta: Diptera). In: Thompson, F.C., Brake, I. & Lonsdale, O. (Eds.), Contributions to the Biosystematic Database of World Diptera. Myia, 12, pp. 522–568. Woodley, N.E., Borkent, A. & Wheeler, T.A. (2009) Phylogeny of Diptera [Chapter 5]. In: Brown, B.V., Borkent, A., Cumming, J.M., Wood, D.M., Woodley, N.E. & Zumbado, M.A. (Eds.), Manual of Central American Diptera. Vol. 1. NCR Research Press, Boca Raton, Ottawa, pp. 79–94. CATALOGUE OF XYLOPHAGIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 305 Zootaxa 4122 (1): 306–311 http://www.mapress.com/j/zt/ Copyright © 2016 Magnolia Press Article ISSN 1175-5326 (print edition) ZOOTAXA ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.4122.1.25 http://zoobank.org/urn:lsid:zoobank.org:pub:45AE96AB-57C9-4415-90D0-F07975721237 FAMILY PANTOPHTHALMIDAE MARTA WOLFF1, YARDANY RAMOS-PASTRANA1, 2 & FERNANDO VALLEJO3 1 Grupo de Entomologia, Instituto de Biología, Universidad de Antioquia, Medellín-Colombia, Centro de Investigación de la biodiversidad Andino-Amazónica –IMBIANAN- Programa de Biología, Universidad de La Amazonia, Florencia, Caquetá, 3 Facultad de Ciencias Agripecuarias, Universidad de Caldas, Manizales-Colombia. E-mail: martha.wolff@udea.edu.co; ya.ramos@udla.edu.co; luis.vallejo_e@ucaldas.edu.co *Corresponding author: martha.wolff@udea.edu.co 2 Abstract This catalogue features 11 species distributed in 2 genera recorded to Colombia. Three species of Pantophthalmus are recorded for the first time to the country. Key words: catalogue, Colombia, distribution, Pantophthalmidae Introduction Pantophthalmidae are a family of Diptera exclusive to the Neotropical Region, with 20 species and two genera: Opetiops with one species from South America, and Pantophthalmus with 19 species occurring from southern Mexico to northern Argentina (Val 1976, 1992). Eight species are recorded from Colombia (Papavero 2009). Adults may be recognized as stout flies, body size between 20–55 mm in length; big head occupied largely by the eyes, holoptic in males and dichoptic in females, with three ocelli; face with conic or round beak; antennae with two short basal joints and the third 8-segmented; short proboscis and palpi with two segments; thorax usually with brown or black longitudinal stripes; wing with small calypter, short prefurca; veins R4 and R5 divergent enclosing wing tip, fourth posterior cell closed, anal cell closed; legs slender with or without apical spurs in posterior femora. It is not clear if the adults feed or not (Val 1992), the eggs are laid on a wide diversity of trees (Papavero 1967), and the larvae live and excavate horizontal galleries in living and dead trees (Woodley 2009). This catalogue and the classification adopted are based on the Systematics and Evolution of Pantophthalmidae by Val (1976) and the Catalogue of Neotropical Pantophthalmidae by Papavero (2009), the holotypes sex was taking of Val 1976. Three species are recorded for the first time to Colombia, Pantophthalmus chuni, P. comptus and P. engeli. For each species, distributional data, when available, include departments (underline) and specific localities (in brackets). Acronyms used for the depositories BMNH—National Museum of Natural History, London, UK BZM—Berliner Zoologische Museum, Humboldt-Universität, Berlin, Germany CEUA—Colección Entomológica Universidad de Antioquia, Colombia IZPAN—Instytut Zoologiczny Polskiej Akademii Nauk, Warsaw, Poland LEUC—Laboratorio de Entomología, Universidad de Caldas, Manizales, Colombia. NMW—Naturhistorisches Museum Wien, Vienna, Austria SMTD—Staatliches Museum für Naturkunde, Dresden, Germany 306 Accepted by S. Nihei: 13 Apr. 2015; published: 14 Jun. 2016 UAM-E—Colección de Entomología, Museo de Historia Natural, Universidad de La Amazonia, Colombia ZSBS—Zoologische Sammlung des Bayerischen Staates, Munich, Germany List of abbreviations cat.—catalogue n. comb.—new combination leg.—legitur (collector) des.—designation distr.—geographic distribution m.—meters mon.—monotypy nom. nov.—nomen novum (new name) orig. des.—by original designation pp.—pages refs.—references sp., spp.—species LT—lectotype HT—Holotype M (m#)—male F (f#)—female Catalogue of Pantophthalmidae of Colombia Family Pantophthalmidae Bigot, 1882 Genus Opetiops Enderlein Opetiops Enderlein, 1921: 231. Type species, Pantophthalmus alienus Hermann, 1916 (orig. des.). alienus (Hermann), 1916: 43, (Pantophthalmus). Type locality: Brazil, Santa Catarina, Corupá (as Kolonie Hansa(Humboldt). HT F (ZSBS). Distr.: Colombia (Boyacá (Muzo), Santander), Brazil, Panama, Peru. Refs.: Papavero, 1967: 3 (cat.); Val, 1976: 69; Amat, 2005: 96 (list); Papavero, 2009: 2 (cat.). Genus Pantophthalmus Thunberg Pantophthalmus Thunberg, 1819: viii. Type species, Pantophthalmus tabaninus Thunberg, 1819 (mon.). Refs.: Papavero, 1967: 1; Val, 1976: 70; Papavero, 2009: 2 (cat.). Pantophtalmus Thunberg, 1819: vii (in title), error. Rhaphiorhynchus Wiedemann, 1821: 59. Acanthomera Wiedemann, 1821: 60. Raphiorhynchus Lepeletier & Serville, 1825: 544, error. Raphiorhynchus Loew, 1862: 19, error. Megalomyia Bigot, 1880: v. Megalemyia Bigot, 1881: 455, error or emend. Raphiorhyncus Bigot, 1881: 458, error. Accanthomera Bigot in Scudder, 1882: 3, error. Raphiorrhynchus Enderlein, 1914: 582, error. Atopomyia Austen, 1923: 596. Lycops Enderlein, 1931: 361 (in key), 367 Meraca Enderlein, 1934: 181. bellardii (Bigot) in Bellardi, 1862: 213 (Acanthomera; nom. nov. for picta Wiedemann of Bellardi, 1859). Type CATALOGUE OF PANTOPHTHALMIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 307 locality: “Mexico”. HT F (BMNH). Distr.: Colombia (?Buena Vista, Chocó (Nuquí), Valle del Cauca (Rio Dagua)), Bolivia, Brazil, Costa Rica, Ecuador, El Salvador, Guatemala, Mexico, Nicaragua, Panama, Peru, Venezuela. Refs.: Austen, 1923: 588; Papavero, 1967: 2 (cat.); Val, 1976: 72; Papavero, 2002: 1(cat.); Amat, 2005: 96 (list.); Papavero, 2009: 2 (cat.). picta Wiedemann of Bellardi, 1859: 76 (Acanthomera), misident. championi Osten Sacken, 1886: 67 (Acanthomera). sp. Osten Sacken, 1912: 108 (Acanthomera). helleri Enderlein, 1912: 108, 110. championi Osten Sacken of Enderlein, 1921: 231, misident. [cf. Enderlein, 1931: 374]. versicolor Austen, 1923: 572. latifrons Enderlein, 1931: 374. Material examined. COLOMBIA: Chocó: Nuquí, Morromico, 10m, 1 female, 24 ix.1999, M. Wolff leg., net house (CEUA). chuni (Enderlein), 1912: 102, 103, (Acanthomera). Type locality: Peru, Junín, Chanchamayo. LT M (IZPAN) (des. by Val, 1976: 75). Distr.: Colombia (Caquetá (Florencia, Aguas Negras), Putumayo (Mocoa, San José del Pepino), Argentina, Bolivia, Brazil, Ecuador, French Guiana, Peru. Refs.: Val, 1976: 74; Papavero, 2009: 3 (cat.). New record to Colombia. leuckarti Enderlein, 1912: 107, 113. helleriana Enderlein, 1914: 578 (in key), 581 (Acanthomera). Material examined. COLOMBIA: Caquetá: Florencia, vereda Aguas Negras, 450m, 2 females, 23 v.2012, Y. Ramos-Pastrana Leg., net in forest (UAM-E). Putumayo: Mocoa, vereda San José del Pepino, vía Mocoa, Villa Garzón “fin del mundo”, 560m, 1 female, 26 iii.2015, M. Angulo-González leg., net in forest (UAM-E). comptus Enderlein, 1912: 107, 117. Type locality: “South America”. HT F (SMTD). Distr.: Colombia (Caldas municipio de La Merced), Caquetá (Florencia, Sebastropol)), Belize, Brazil, French Guiana, Guyana, Peru, Surinam. Refs.: Val, 1976: 75; Papavero, 2009: 3 (cat.). New record to Colombia. conspicuus Austen, 1923: 581. sigma Enderlein, 1931: 372 (Opetiops). Material examined. COLOMBIA: Caldas: municipio de La Merced, 1819m, 1 male, 18 viii.2009, TrujilloToro leg., net in grass (LEUC). Caqueta: Florencia, vereda Sebastropol, 380m, 1 female, 23 v.2011, Y. RamosPastrana leg., net in forest (UAM-E). engeli (Enderlein), 1931: 362 (Acanthomera). Type locality: Panama, Chiriquei. HT M ZSBS. Distr.: Colombia (Caldas), Panama. Refs.: Val, 1976: 76; Papavero, 2009: 3 (cat.). New record to Colombia. Material examined. COLOMBIA: Caldas: Ecoparque los Alcázares, 2150m, 1 male, 15 x.2010, Montoya & Ramírez leg., net in Guadua angustifolia (LEUC); Manizales, 2000m, 1 male, 27 vii.2010, L. Patiño leg., net in grass (LEUC). frauenfeldi (Schiner), 1868: 78 (Acanthomera). Type locality: “Colombia”. Type ?NMW not found by Val (1976: 78). Distr.: Colombia (Antioquia (Medellín, San Antonio de Prado), Boyacá (Muzo), Caldas (municipio de Aranzazu; vereda Lusitania; Municipio de Villamaría; Manizales; Reserva Rio Blanco), Caquetá (Florencia; Aguas Negras) Cundinamarca (Bogotá), Santander (Suarez river)), Bolivia, Ecuador, Peru, Venezuela. Refs.: Kertész, 1908: 294 (cat.; n. comb.); Austen, 1923: 565; Papavero, 1967: 3 (cat.; Rhaphiorhynchus); Val, 1976: 77; Amat, 2005: 96 (list.); Papavero, 2009: 3 (cat.). Material examined. COLOMBIA: Antioquia: Medellín, San Antonio de Prado, 2100m, 1 female, 23 ix.2007, Castaño Leg., net forest (CEUA). Caldas: vereda Lusitania, Villamaría, 2050m, 1 female, 20 iv. 2008, Castro-Hoyos leg., net (LEUC); idem, municipio de Aranzazu, 1900m, 1 female, iii.2006, Arango-Salazar leg., net in Cospositae (LEUC); Manizales, Reserva Rio Blanco, 1900m, 1 female, 3 iv.2005, M. Wolff leg., bats network in forest (CEUA 19167); Manizales, Granja Tesorito, 2480m; 1 male, 18 iv.2008, F. Vallejo leg., net in grass (LEUC). Caquetá: Florencia, vereda Aguas Negras, 280m, 1 female, 23 v.2012, Y. Ramos-Pastrana leg., net in forest; idem, barrio Ventilador, 280m, 1 female, 21 iv.2011, Y. Ramos-Pastrana leg., net (UAM-E). 308 · Zootaxa 4122 (1) © 2016 Magnolia Press WOLFF ET AL. kerteszianus (Enderlein), 1914: 578 (in key and text) (Acanthomera; as kertésziana). Type locality: Peru, Juanjui. LT F (BZM) (des. by Val, 1976: 79). Distr.: Colombia (Without specific locality), Bolivia, Brazil, Panama, Peru. Refs.: Papavero, 1967: 3 (cat.; Rhaphiorhynchus); Val, 1976: 78; Papavero, 2002: 2 (cat.); Amat, 2005: 96 (list.); Papavero, 2009: 3 (cat.). conspicabilis Austen, 1923: 577. planiventris (Wiedemann), 1821: 60 (Rhaphiorhynchus). Type locality: “Brazil”. T not found (Val, 1976: 82). Distr.: Colombia (Cundinamarca (Bogotá), Santander (Barrancabermeja; Suarez river)), Bolivia, Brazil, Costa Rica, Ecuador, French Guiana, Guatemala, Guyana, Mexico, Nicaragua, Panama, Peru, Surinam, Trinidad, Venezuela. Refs.: Kertész, 1908: 296 (cat.; Rhaphiorhynchus); Papavero, 1967: 3 (cat.; Rhaphiorhynchus); Val, 1976: 81 (comb. n); Papavero, 2002: 1 (cat.); Amat, 2005: 95, 96 (list.); Papavero, 2009: 4 (cat.). crassipalpis Macquart, 1847: 27 (Acanthomera). bigoti Bellardi, 1862: 15 (Acanthomera). rothschildi (Austen), 1909: 129, (Rhaphiorhynchus). Type locality: Bolivia, Santa Cruz, Buenavista. HT M (BMNH). Distr.: Colombia (Meta (Huape river Orinoco region)), Bolivia, Brazil, Ecuador. Refs.: Papavero, 1967: 4 (Rhaphiorhynchus); Val, 1976: 84; Papavero, 2002: 1 (cat.); Amat, 2005: 96 (list.); Papavero, 2009: 5 (cat.). ocellata Enderlein, 1931: 370 (Atopomyia; as rotschildi var.). tabaninus Thunberg, 1819: vii, [Pantophtalmus]. Type locality: Lesser Antilles, Saint Barthélemy (Forsström) (cf. Papavero, 1971: 103). Type lost (cf. Val, 1976: 88). Distr.: Colombia (Amazonas (Parque Natural Amacayacu); Antioquia (Caucasia; San Luis), Caquetá (Florencia), Chocó (Riosucio; Cacarica; Quibdó), Meta (Villavicencio), Putumayo (Valle del Guamés)), Bolivia Brazil, French Guiana, Guatemala, Guyana, Lesser Antilles, Panama, Peru, Surinam, Trinidad, Venezuela. Refs.: Kertész, 1908: 296 (cat.); Papavero, 1967: 2 (cat.); Val, 1976: 87; Papavero, 2002: 2 (cat.); Amat, 2005: 95, 96 (list.); Papavero, 2009: 5 (cat.). seticornis Wiedemann, 1828: 108, pl. 2, fig. 1 (Acanthomera). heydenii Wiedemann, 1828: 555 (Acanthomera). immanis Wiedemann, 1830: 623 (Acanthomera). picta Wiedemann of d’Orbigny in Cuvier, 1849: pl. 172bis, fig. 1 (Acanthomera), misident. rubriventris Bigot, 1880: v (Acanthomera). setiformis Bigot, 1881: 454, 455 (Megalemyia), error. tabanius Kertész, 1908: 296, error. gigas Enderlein, 1912: 109, 110. fastuosus Knab, 1914: 27. gigans Bezzi, 1912: 287, error. reticornis Enderlein, 1921: 231, error. Material examined. COLOMBIA: Antioquia: Caucasia, 80m, 1 male, iv.1987, A. Arango Leg., net (CEUA); idem, San Luis, 80m, 1 female, ix.2000, M. Castaño Leg., net forest (CEUA). Caquetá, Florencia, 450m, 1 female, 20 viii.2007, Y. Ramos-Pastrana leg., net. vittatus (Wiedemann), 1828: 109, (Acanthomera). Type-locality: “Brazil”. LT M (BZM) (des. by Val, 1976: 91). Distr.: Colombia (Meta (Villavicencio)), Bolivia, Brazil, French Guiana, Peru, Surinam, Trinidad, Venezuela . Refs.: Papavero, 1967: 2 (cat.); Val, 1976: 89; Papavero, 2002: 1 (cat.); Amat, 2005: 96 (list); Papavero, 2009: 6 (cat). servillei Guérin-Méneville, 1835: 543, pl. 98, fig. 2 (Acanthomera). fulvida Bigot, 1880: v (Acanthomera). CATALOGUE OF PANTOPHTHALMIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 309 TABLE 1. Pantophthalmidae species recorded from Colombia Genus Species Author Opetiops alienus (Hermann, 1916) Pantophthalmus bellardi (Bigot, 1862) chuni Enderlein, 1912 comptus Enderlein, 1912 engeli (Enderlein, 1931) frauenfeldi (Schiner, 1868) kerteszianus (Enderlein, 1914) planiventris (Wiedemann, 1821) rothschildi (Austen, 1909) tabaninus Thunberg, 1819 vittatus (Wiedemann, 1828) References Amat, E.C. (2005) New records of timber flies (Diptera: Pantophthalmidae) from Colombia. Entomotropica, 20, 95–96. Austen, E.E. (1909–1910) Two remarkable new species of Diptera. Novitates Zoologicae. Tring, 16, 129–131. [1909 & 1910] Austen, E.E. (1923) A revision of the family Pantophthalmidae (Diptera) with descriptions of new species and a new genus. Proceeding of the Zoological Society of London, 551–598. Bellardi, L. (1862) Saggio di Ditterologia Messicana. Appendice, Stamperia Reale, Torino [=Turin], 28 pp. Bellardi, L. (1859) Saggio di Ditterologia Messicana. Vol. 1. Dalla Stamperia Reale, Torino, 1–80. Bezzi, M. (1912) Os gigantes entre as moscas inimigas da sylvicultura brasileira. Chácaras e Quintais, 5, 1–4. [São Paulo] Bigot, J.M.F. (1880) Description d’un genre nouveaux et de nouvelles espèces d’Acanthomeridae Wiedemann [sic]. Annales de la Société Entomologique de France, Series 5, 10 (Bull), v–vi. Bigot, J.M.F. (1881) Diptères nouveaux ou peu connus. 18e partie. XXVIII. Acanthomeridae (Wiedemann, Dipt. exot. [sic]).]. Annales de la Société Entomologique de France, Series 6, 1, 453–460. Cuvier, G.L.C.F.D. (1849) Le régne animal distribué d’aprés son organisation. Disciples ed., Paris, Masson. Insectes 2, 1–443. Enderlein, G. (1912) Dipterologische Studien. I. Die Dipterenfamilie Pantophthalmidae. Zoologischer Anzeiger Leipzig, 41, 97–118. Enderlein, G. (1914) Dipterologische Studien. XIII. Weitere Beiträge zur Kenntniss der Pantophthalmiden. Zoologischer Anzeiger Leipzig, 44, 577–586. Enderlein, G. (1921) Dipterologische Studien. XI. Zoologischer Anzeiger Leipzig, 52, 227–231. Enderlein, G. (1931) Klassifikation der Pantophthalmiden. Sitz Berlinische Gesllschaft Naturforschender Freunde, Berlin, 1930, 361–376. Enderlein, G. (1934) Dipterologica. II. Sitz Berlinische Gesllschaft Naturforschender Freunde, Berlin, 181–190. Guérin-Méneville, M.F.E. (1835) Iconographie du règne animal de George Cuvier. Baillière, Paris, 3, 543–544. Hermann, F. (1916) Ein neuer Pantophthalmus nebst kritischen Bemerkungen über die Systematik der Pantophthalmiden (Dipt.). Deutsche Entomologische Zeitschrift, 43–49. http://dx.doi.org/10.1002/mmnd.48019160109 Kertész, K. (1908) Catalogus dipterorum hucusque descriptorum. Vol. 3. Lipsiae & Budapestini [=Leipzig and Budapest], 367 pp. http://dx.doi.org/10.5962/bhl.title.5141 Knab, F. (1914) A new Pantophthalmus (Diptera, Pantophthalmidae). Insecutor Inscitiae Mentruus, 2, 27–29. Lepeletier, A.L.M. & Serville, J.G.A. (1825) In: Latreille, P.A. (Ed.), Encyclopédie Méthodique Histoire Naturelle. Entomologie, ou Histoire Naturelle des Crustacés, des Arachnides et des Insectes. Tome dixieme. Agasse, Paris, pp. 1–344 Loew, H. (1862) Monographs of the Díptera of North America. Part I. Smithsonian Miscellaneous Collections, 6, 1–221. Macquart, J. (1847) Diptères exotiques nouveaux ou peu connus. Suite de 2e. supplément. Mémoires de la Societé Royale des Sciences, de l’Agriculture et des Arts de Lille, 1846, 21–120. Osten Sacken, C.R. (1886) Diptera. In: Godman, F.D. & Salvin, O. (Eds.), Biologia Centrali-Americana. Zoologia. Insecta. Diptera. Vol 1. Taylor & Francis, London, 378 pp. Papavero, N. (1967) Family Pantophthalmidae. In: Papavero, N. (Ed.), A catalogue of the Diptera of the Americas south of the United States. Museu de Zoologia, Universidade de São Paulo, São Paulo, pp. 8. Papavero, N. (1971) Essays on the history of Neotropical Dipterology, with special reference to collectors (1750–1905). Vol. 1. 310 · Zootaxa 4122 (1) © 2016 Magnolia Press WOLFF ET AL. Museu de Zoologia, Universidade de São Paulo, São Paulo, vii + 216 pp. Papavero, N. (2002) Insecta – Díptera – Pantophthalmidae. Fauna da Amazônia Brasileira, 11, 1–4. Papavero, N. (2009) Catalogue of Neotropical Diptera. Pantophthalmidae. Neotropical Diptera, 19, 1–11. [Ribeirão Preto] Schiner, J.R. (1868) Diptera. In: von Wüllerstorf-Urbair, B. (Ed.), Reise der österreichischen Fregatte Novara um die Erde in den Jahren 1857, 1858, 1859 unter den Befehlen des Commodore B. Von Wüllerstorf-Urbair. Zoologischer Theil. Zweiter Band. 1. Abtheilung B. K. Gerold’s Sohn, Wien [= Vienna], pp. i–vi + 1–388. Thunberg, C.P. (1819) Beskrifning och teckning på en fórut okånd westindisk fluga, Pantopthalmus tabaninus. Götteborgs k. Vettensk. och Vitterhets Samhället. Handl. 3, vii–x, pl. 1. Val, F.C. (1976) Systematics and evolution of the Pantophthalmidae (Diptera, Brachycera). Arquivos de Zoologia, 27, 51–164. http://dx.doi.org/10.11606/issn.2176-7793.v27i2p51-164 Val, F.C. (1992) Pantophthalmidae of Central America and Panama (Diptera). In: Quintero, D. & Aiello, A. (Eds.), Insects of Panama and Mesoamerica. Select Studies, Oxford University Press, Oxford, pp. 600–610. [692 pp.] Wiedemann, C.R.W. (1821) Diptera exotica (Pars. I). In academia Kiliensi, Kiel, 244 pp. Wiedemann, C.R.W. (1828) Aussereuropäische Zweiflügelige Insekten. Vol 1. In der Schulzischen, Hamm, xxxii + 608 pp. Wiedemann, C.R.W. (1830) Aussereuropäische zweiflügelige Insekten. Vol. 2. Hamm, xii + 684 pp. Woodley, N.E. (2009) Pantophthalmidae. In: B. Brown; Borkent, A, Cumming, J.M, Wood, D.M, Woodley, N.E. & Zumbado, M.A. (Eds.), Manual of Central American Diptera. Vol 1. NRC Research Press, Ottawa, pp. 513–515. CATALOGUE OF PANTOPHTHALMIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 311 Zootaxa 4122 (1): 312–341 http://www.mapress.com/j/zt/ Copyright © 2016 Magnolia Press Article ISSN 1175-5326 (print edition) ZOOTAXA ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.4122.1.26 http://zoobank.org/urn:lsid:zoobank.org:pub:43593E4D-8E4F-4BC2-9C0B-D822DACE7EAA FAMILY STRATIOMYIDAE DIEGO AGUILAR FACHIN1* & CRISTIANE VIEIRA DE ASSIS-PUJOL2 1 Departamento de Biologia, Faculdade de Filosofia, Ciências e Letras de Ribeirão Preto, Universidade de São Paulo, Av. Bandeirantes 3900, Ribeirão Preto, SP–14.040-901, Brazil. 2 Laboratório de Zoologia, Universidade Católica de Brasília, QS 7 lote 1, Bl. M/ s. 331, Taguatinga, DF-72.030-170, Brazil. *Corresponding author, E–mail: diegoafachin@gmail.com. Abstract The family Stratiomyidae has more than 2,800 described species, of which 1001 species belongs to the Neotropics. This catalog for Colombia presents 87 species distributed in 32 genera, and ten subfamilies. Merosargus gracilis and the genus Microchrysa, with a single species M. bicolor are recorded for the first time to Colombia. The fauna is very expressive but still poorly known, representing nearly one tenth of the Neotropical diversity of the family in numbers of species, and one fifth of generic diversity. Key words: catalogue, Colombia, distribution, diversity, Stratiomyidae Introduction Stratiomyidae are a family of lower Brachycera, with small to fairly big flies and body length varying from 2 to 34 mm. They are recognized mainly by the wing venation, with their radial veins concentrated in the anterior part of the wing and a small discal cell from which medial veins radiate (Woodley 2009). The family is placed within the infraorder Stratiomyomorpha, as sister-group of the Xylomyidae (Woodley 1989; Woodley et al. 2009). There are more than 2,800 described species of Stratiomyidae in the world, included in 377 genera (Woodley 2001, 2011). From these total, 1001 species in 162 genera occur in the Neotropical region. The family has often been divided into 12 subfamilies: Antissinae, Beridinae, Chiromyzinae, Chrysochlorininae, Clitellarinae, Hermetiinae, Nemotelinae, Pachygastrinae, Parhadrestiinae, Raphiocerinae, Sarginae e Stratiomyinae (Woodley 2001). Brammer & Dohler (2010) published the most recent phylogenetic analysis of the family, improving the understanding of the relationships in the group. In their hypothesis, the subfamilies Antissinae, Stratiomyinae, Sarginae and Clitellariinae, in the way they have traditionally being defined, may correspond to non-monophyletic groups. The catalog records 87 described species, included in 32 genera, and ten subfamilies to Colombia. A summary table of the data given here is presented in Table 1. Furthermore, Merosargus gracilis Schiner, 1868b and the genus Microchrysa Loew, 1855a, with a single species M. bicolor (Wiedemann), 1830 are recorded for the first time to Colombia. The examined material of these species belongs to the National Museum of Natural History, Washington, D.C, United States (USNM) and Canadian National Collection, Arachnids and Nematodes, Ottawa, Ontario, Canada (CNC), respectively. The records of the species and the classification adopted here were based mainly on information of the family catalogues to Neotropical region (James 1973) and to the world (Woodley 2001, 2011). Additional literature and material were also consulted. The departments of Colombia are underline and specific localities are in brackets. The term “allotype”, “paratype” and syntype were used in this catalogue only for the specimens that were designated as such in the original description. 312 Accepted by S. Nihei: 10 Apr. 2015; published: 14 Jun. 2016 Acronyms used for the depositories AMNH—American Museum of Natural History, New York, New York, United States; BMNH—Department of Entomology, Natural History Museum, London, England; CAS—California Academy of Sciences, San Francisco, California, United States; CNC—Canadian National Collection of Insects, Ottawa, Ontario, Canada; HNHM—Hungarian Natural History Museum, Budapest, Hungary; INBIO—Instituto Nacional de Biodiversidad, Santo Domingo de Heredia, Costa Rica; KU—Entomological Laboratory, Kyushu University, Fukuoka, Japan; LACM—Natural History Museum of Los Angeles County, Los Angeles, California, United States; LSL—Linnean Society of London, London, England; MCZ—Museum of Comparative Zoology, Cambridge, Massachusetts, United States; MHNLI—Musée d’Histoire Naturelle et d’Ethnographie, Lille, France; MIZT—Museo Regionale di Storia Naturale, Torino, Italy; MNHN—Muséum National d’Histoire Naturelle, Paris, France; NMW—Naturhistorisches Museum Wien, Vienna, Austria; NRS—Naturhistoriska Riksmuseet, Stockholm, Sweden; PAN—Museum of the Institute of Zoology, Polish Academy of Sciences, Warsaw, Poland; RNH—Nationaal Natuurhistorisch Museum, Leiden, Netherlands; SEMC—Snow Entomological Museum, Department of Entomology, University of Kansas, Lawrence, Kansas, United States; SMF—Forschungsinstitut und Naturmuseum Senckenberg, Frankfurt/Main, Germany; SMNS—Staatliches Museum für Naturkunde Stuttgart, Stuttgart, Germany; UNK—Unknown depositary of type material; USNM—National Museum of Natural History, Washington, D.C., United States; USU—Department of Biology, Utah State University, Logan, Utah, United States; UZMC—Natural History Museum of Denmark, Copenhagen, Denmark; WSU—Department of Entomology, Washington State University, Pullman, Washington, United States; ZIL—Museum of Zoology, Lund, Sweden; ZMHU—Museum für Naturkunde, Institut für Systematische Zoologie, Berlin, Germany; ZSBS—Zoologische Staatssammlung München, Munich, Germany. List of abbreviations cat.—catalogue des.—designation distr.—geographic distribution emend.—emended m.—meters mon.—monotypy nom. nov.—new name orig. des.—by original designation preocc.—preoccupied ssp.—subspecies subs. des.—subsequent designation syn.—synonym AT—allotype HT—holotype LT—lectotype PT—paratype ST—syntype CATALOGUE OF STRATIOMYIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 313 M (m#)—male F (f#)—female Catalogue of Stratiomyidae of Colombia Family Stratiomyidae Latreille, 1802 Subfamily Beridinae Westwood, 1838 Genus Arcuavena Woodley Arcuavena Woodley, 1995: 58. Type species, Hoplacantha nigerrima Lindner, 1949a (orig. des.). singularis (Macquart), 1846: 185 (Nemotelus). Type locality: “Colombia”. Distr.—Colombia, Honduras, Venezuela. HT F [BMNH]. Ref.—Woodley, 1995: 62 (rev. of the subfamily, desc. of new species, key to genera). Genus Heteracanthia Macquart Heteracanthia Macquart, 1850: 347. Type species, Heteracanthia ruficornis Macquart, 1850 (orig. des.). Heleracanthia Macquart, 1850: plate 5, error. Heteracantha Williston, 1896: 46, error. ruficornis Macquart, 1850: 348. Type locality: Venezuela, Caracas. Distr.—Bolivia, Brazil (Pará, São Paulo), Colombia, Costa Rica, Ecuador, El Salvador, French Guiana, Guatemala, Honduras, Mexico (Chiapas, Quintana Roo, Tabasco, Veracruz), Panama, Peru, Suriname, Trinidad, Venezuela. HT M [MNHN]. Refs.—Lindner, 1933c: 200; McFadden, 1972a: 122; Woodley, 2001: 70 (cat.). Clitellaria scutellaris Walker, 1854: 313. Type locality: Brazil, Santarém. HT sex? [BMNH], destroyed? Ref.—Woodley, 2001: 70 (cat., syn.). Genus Oplachantha Rondani Oplachantha Rondani, 1863: 87. Type species, Beris mexicana Bellardi, 1859 (orig. des.). Oplacantha Bigot, 1879a: 185, error. Hoplacantha Brauer, 1882: 88, error. Hoplacantha Scudder, 1882: 166, emend. of Oplachantha Rondani. Oplacantha Scudder, 1882: 236, emend. of Oplachantha Rondani. Olachantha Hunter, 1900: 122, error. albihirta James, 1977: 312. Type locality: Colombia, Meta, 3 miles W of Villavicencio, 920 m. Distr.—Colombia, (Meta (3 miles W of Villavicencio, 920 m)). HT M [CAS]. Refs.—James, 1977: 312 (desc. of new species, key to species); Woodley, 1995: 133 (rev. of the subfamily, desc. of new species, key to genera). albitarsis (Macquart), 1846: 175 (Beris). Type locality: “Venezuela”. Distr.—Colombia, Venezuela. LT M [BMNH]. Refs.—Woodley, 1995: 133, 148 (sub. des.; rev. of the subfamily, desc. of new species, key to genera). atricalx James, 1977: 311. Type locality: Peru, Cuzco Department, Quincemil, 700 m. Distr.—Colombia (Meta (3 miles W of Villavicencio, 920 m)), Peru. HT M [CNC]; 1 PT M [CAC = CAS]. Refs.—James, 1977: 311 (desc. of new species, key to species); Woodley, 1995: 133 (rev. of the subfamily, desc. of new species, key to genera). atriceps James, 1977: 314. Type locality: Peru, Cuzco Department, Quincemil, 700 m. Distr.—Colombia, (Meta (3 miles W of Villavicencio, 920 m)), Peru. HT F [CNC]; PT 1 F [UNK]. Refs.—James, 1977: 315 (desc. of new species, key to species); Woodley, 1995: 133 (rev. of the subfamily, desc. of new species, key to genera). 314 · Zootaxa 4122 (1) © 2016 Magnolia Press FACHIN & ASSIS-PUJOL caerulescens (Schiner), 1868b: 71 (Beris). Type locality: “Colombia”. Distr.—Colombia. HT F [NMW], destroyed? Ref.—Woodley, 2001: 73 (cat.). Beris coerulescens Kertész, 1908b: 124, error. Ref.—Woodley, 2001: 73 (cat.). formosa Enderlein, 1921: 207. Type locality: Colombia, “Cordilleren, Terra caliente”. Distr.—Colombia, Costa Rica, Panama. HT M [ZMHU]. Ref.—Woodley, 1995: 133 (rev. of the subfamily, desc. of new species, key to genera). pallida James, 1977: 312. Type locality: Colombia, Nariño, 16 miles E of Santiago, 2730 m. Distr.—Colombia (Nariño). HT F [CAS]. Ref.—Woodley, 1995: 134 (rev. of the subfamily, desc. of new species, key to genera). pulchella (Williston), 1888: 245 (Beris). Type locality: Brazil, Mato Grosso, Chapada. Distr.—Bolivia, Brazil (Mato Grosso), Colombia (Meta (3 miles W of Villavicencio, 920 m)), Ecuador, Peru (Cuzco, Vilcanota, Tingo Maria). LT M [AMNH]. Refs.—James, 1977: 309 (desc. of new species, key to species); Woodley, 1995: 134, 151 (sub. des.; rev. of the subfamily, desc. of new species, key to genera). Hoplacantha iubata Enderlein, 1921: 208. Type locality: Peru, Vilcanota. ST F [ZMHU]; ST M [HNHM], destroyed. Ref.—Woodley, 2001: 74 (cat.). viriata Enderlein, 1921: 206. Type locality: Peru, Cuzco Department, Callanga. Distr.—Colombia (Meta (3 miles W of Villavicencio, 920 m)), Peru. ST 1 M, 1 F [HNHM], destroyed. Refs.—James, 1977: 311 (desc. of new species, key to species); Woodley, 1995: 134 (rev. of the subfamily, desc. of new species, key to genera). Subfamily Chiromyzinae Brauer, 1880 Genus Nonacris Walker Nonacris Walker, 1850: 7. Type species, Nonacris transequa Walker, 1850 (mon.). Nanacris Nagatomi, 1991: 23, error. longicornis Enderlein, 1921: 164. Type locality: “Colombia”. Distr.—Colombia. HT F [ZMHU]. Ref.—Pujol-Luz et al., 2001: 1. nigriventris Enderlein, 1921: 165. Type localities: Colombia and Ecuador, Santa Inéz (there is no lectotype designation). Distr.—Colombia, Ecuador (Santa Inéz). ST 2 M, 1 F [ZMHU]. Refs.—Enderlein, 1921: 163, Figs 1– 2 (desc. of new species); Pujol-Luz et al., 2001: 1. scutellaris Enderlein, 1921: 164. Type locality: Colombia, Cundinamarca, Bogotá, Tapias la Cruzes. Distr.— Colombia Colombia (Cundinamarca (Bogotá, Tapias la Cruzes). ST 2 M, 1 F [ZMHU]. Ref.—Pujol-Luz et al., 2001: 1. transequa Walker, 1850: 7. Type locality: “South America”. Distr.—Colombia (“Cordillieren, Terra fria”), Venezuela. HT M [BMNH]. Ref.—Enderlein, 1921: 163–164. Chiromyza transsequa Kertész, 1908b: 144, error. Ref.—Woodley, 2001: 51 (cat.). Subfamily Clitellariinae Brauer, 1882 Genus Acropeltates Kertész Acropeltates Kertész, 1923: 118. Type species, Acropeltates fasciata Kertész, 1923 (orig. des.). Monacanthella James, 1936: 49. Type species, Monacanthella gracilis James, 1936 = Acropeltates kerteszi Szilády, 1929 (orig. des.). CATALOGUE OF STRATIOMYIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 315 diversicornis Kertész, 1923: 121. Type locality: “Colombia”. Distr.—Colombia. HT M [HNHM], destroyed. Ref.—Woodley, 2001: 149 (cat.). fasciata Kertész, 1923: 120, Fig. 16. Type locality: Peru, Urubamba River Meshagua and Ukayali River, Unini. Distr.—Colombia, Peru. ST 2 M, 2 F [HNHM], destroyed; ST 2 M, 2 F [MHU]. Ref.—Woodley, 2001: 149 (cat.). Genus Chordonota Gerstaecker Chordonota Gerstaecker, 1857: 311. Type species, Cyphomyia inermis Wiedemann, 1830 (mon.). leiophthalma Williston, 1896: 302. Type locality: St. Vincent. Distr.—Colombia (Boyacá (Muzo), Meta), Grenada, St. Vicent. ST M [BMNH], 1 M [UNK]. Refs.—James, 1940: 122 (desc. of new species, key to species); Woodley, 2001: 155 (cat.). Genus Cyphomyia Wiedemann Cyphomyia Wiedemann, 1819: 54. Type species, Stratiomys cyanea Fabricius, 1794 (sub. des. of Duponchel in d'Orbigny, 1844: 548). Ref.—Evenhuis & Thompson, 1990: 234. Cyphomia Rondani, 1863: 85, error. Ciphomia Rondani, 1863: 86, error. Cyphomya Rondani, 1863: 94, error. Rondania Jaennicke, 1867: 324 (preocc. by Robineau-Desvoidy, 1850; Bigot, 1854). Type species, Rondania obscura Jaennicke, 1867 (mon.). Neorondania Osten Sacken, 1878: 50 (nom. nov. for Rondania Jaennicke, 1867). Type species, Rondania obscura Jaennicke, 1867. Gyneuryparia Enderlein, 1914b: 604. Type species, Cyphomyia pilosissima Gerstaecker, 1857 (orig. des.). Ciphomyia James, 1980: 260, error. Cephomyia Mead, 1992: 5, error. albitarsis (Fabricius), 1805: 80 (Stratiomys). Type locality: “America meridionali”. Distr.—Bolivia, Brazil (Pará, Roraima), Colombia (Valle del Cauca), Costa Rica (San Mateo, Higuito), Ecuador, Guyana, Honduras, Mexico (Chiapas, Nayarit, San Luis Potosí, Veracruz, Yucatán), Panama (Caño Saddle, Gatun Lake; Porto Belo; Tabogal), Paraguay, Peru, Trinidad, Venezuela (Valera). ST 1 F [UZMC]. Refs.—Giglio-Tos, 1893: 9; James, 1939d: 599 (desc. of new species); James, 1940: 140 (desc. of new species, key to species); Rafael, 1991: 331 (checklist). Cyphomyia fenestrata Macquart, 1846: 176. Type locality: Mexico, Yucatán, Merida. ST M/F [UNK]. Ref.— Woodley, 2001: 158 (cat.). altifrons James, 1939d: 603. Type locality: Colombia, Cundinamarca, Bogotá. Distr.—Colombia (Cundinamarca (Bogotá)), Brazil, Peru, Bolivia? HT M [USNM]. Ref.—Woodley, 2001: 158 (cat.). androgyna Osten Sacken, 1886b: 34. Type locality: Panama, Volcan de Chiriqui, Bugaba. Distr.—Colombia, Costa Rica, Ecuador, Panama (Chiriqui). ST 1 M, 4 F [BMNH]. Ref.—Woodley, 2001: 158 (cat.). Cyphomyia chiriquina Lindner, 1935: 409. Type locality: Panama (Chiriqui). HT F [PAN]. Ref.—Woodley, 2001: 158 (cat.). banksi James, 1937: 149. Type locality: Panama, Canal Zone, Barro Colorado Island. Distr.—Colombia, Costa Rica, El Salvador, Mexico (Veracruz), Panama (Canal Zone, Barro Colorado Island). HT F [MCZ]. Ref.— Woodley, 2001: 159 (cat.). chrysodota Perty, 1833a: plate 36, Fig. 13. Type locality: Brazil, “montibus Provinciae Minarum”. Distr.—Belize, Brazil, Colombia (Boyacá (Muzo)), Costa Rica, Ecuador, Guyana, Mexico (Chiapas), Panama. Description given in Perty, 1833b: 184. ST M [ZSBS]. Refs.—James, 1940: 142 (desc. of new species, key to species); Woodley, 2001: 159 (cat.). 316 · Zootaxa 4122 (1) © 2016 Magnolia Press FACHIN & ASSIS-PUJOL cyanea (Fabricius), 1794: 265 (Stratiomys). Type locality: French Guiana. Distr.—Bolivia, Brazil (Pará), Colombia (Cundinamarca (Bogotá)), Costa Rica, Ecuador, French Guiana, Guyana, Panama, Paraguay, Peru, Venezuela. ST F [MNHN], lost. Refs.—Coquebert, 1804: plate XXIII, Fig. 4; Lindner, 1929: 262; James 1939d: 604 (desc. of new species); James, 1940: 143 (desc. of new species, key to species). Cyphomyia caerulea Macquart, 1834 (nom. nov. for Stratiomys cyanea Fabricius, 1794): explication des planches 4: Planche VI, Fig. 3. Ref.—Woodley, 2001: 159 (cat.). Cyphomyia coerulea James, 1953: 313, error. Ref.—Woodley, 2001: 159 (cat.). dispar Schiner, 1868b: 54. Type locality: “Colombia”. Distr.—Colombia (Boyacá (Muzo)). ST 13 M, 25 F [NMW]. Refs.—Bigot, 1876: 487; James, 1940: 136 (desc. of new species, key to species). ecuadoriensis Enderlein, 1914b: 601. Type locality: Ecuador, Archidona. Distr.—Colombia, Ecuador (Archidona), Peru. ST 1 M, 1 F [PAN]. Refs.—James, 1940: 143 (desc. of new species, key to species); James, 1973: 26.26 (cat.); Woodley, 2001: 160 (cat.). Cyphomyia ecuadorensis James, 1973: 26.26, error. Ref.—Woodley, 2001: 160 (cat.). erectispinis Lindner, 1935: 408. Type locality: Colombia, Hacienda Pehlke. Distr.—Colombia (Chocó (Andagoya), Caldas (Victoria, Hacienda Pehlke)). HT F [PAN]. Refs.—James, 1940: 143 (desc. of new species, key to species); Woodley, 2001: 160 (cat.). fassli Lindner, 1949a: 862. Type locality: Colombia, Boyacá, Muzo, 400–800 m. Distr.—Colombia (Boyacá (Muzo)). HT F [BMNH]. Ref.—Woodley, 2001: 160 (cat.). Cyphomyia fassli Lindner, 1949b: 786. Nomen nudum. Ref.—Woodley, 2001: 160 (cat.). formosa James, 1940: 138–139; Fig 2: 133. Type locality: Colombia, Boyacá, Muzo, 900 m. Distr.—Colombia (Boyacá (Muzo)), Brazil (Amazonas)? HT F [MCZ]. Refs.—James, 1940: 138–139 (desc. of new species, key to species); Woodley, 2001: 160 (cat.). geniculata Gerstaecker, 1857: 290. Type locality: “Colombia”. Distr.—Colombia. HT F [ZMHU]. Ref.—Woodley, 2001: 160 (cat.). planifrons James, 1939d: 602. Type locality: Colombia, Cundinamarca, Bogotá. Distr.—Colombia (Cundinamarca (Bogotá)), Peru, Brazil. HT F [USNM]. Ref.—Woodley, 2001: 162 (cat.). schwarzi James, 1940: 135. Type locality: Colombia, Western Cordillera, Cali District, 6500 feet. Distr.— Colombia (Valle del Cauca (Western Cordillera, Cali District, 6500 feet)). HT F [AMNH]. Refs.—James, 1940: 135 (desc. of new species, key to species); Woodley, 2001: 162 (cat.). unicolor (Walker), 1854: 312 (Clitellaria). Type locality: Brazil, Amazonas, Tefé (as Ega). Distr.—Brazil (Amazonas, Pará), Colombia, Peru. ST 1 M, 2 F [BMNH]; ST 1 sex? [BMNH], lost. Refs.—James, 1940: 140 (desc. of new species, key to species); James, 1973: 26.28 (cat.) Woodley, 2001: 163 (cat.). Cyphomyia cyanocephala Gerstaecker, 1857: 274. Type locality: “South America”. HT F [ZMHU]. Ref.— Woodley, 2001: 163 (cat.). varipes Gerstaecker, 1857: 283. Type localities: Mexico, Costa Rica, Venezuela (there is no lectotype designation). Distr.—Colombia (Boyacá (Muzo)), Costa Rica, Guatemala, Mexico (Chiapas), Nicaragua, Panama, Venezuela. ST 3 F (ZMHU). Refs.—Osten Sacken, 1886b: 34 (desc. of new species); James 1940: 137 (desc. of new species, key to species); James, 1973: 26.28 (cat.) violacea Macquart, 1855: 60. Type locality: Brazil, Pará. Distr.—Brazil (Pará), Colombia, Ecuador. ST M, 8 F [BMNH]. Refs.—Lindner, 1929: 264; James, 1940: 144 (desc. of new species, key to species). wiedemanni Gerstaecker, 1857: 302. Type locality: Brazil, Pará. Distr.—Argentina, Bolivia, Brazil (Amazonas, CATALOGUE OF STRATIOMYIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 317 Goiás, Mato Grosso do Sul, Minas Gerais, Pará, Roraima), Colombia (Boyacá (Muzo)), Costa Rica, Ecuador, French Guiana, Guyana, Mexico, Nicaragua, Panama, Paraguay, Peru (Iquitos (Roqué)), Suriname, Tobago, Trinidad, Venezuela. ST M, F [NMW]; ST 1 M, 1 F [ZIL]; ST 2 M, 3 F [ZMHU]; ST sex? [ZMHU]; Refs.— Lindner, 1933a: 203; Lindner, 1933c: 329; James, 1939d: 599–600; James, 1940: 147 (desc. of new species, key to species); James, 1973: 26.28 (cat.); Rafael, 1991: 331 (checklist). Cyphomyia flavispinis Macquart, 1855: 60. Type locality: “Amérique méridionale”. ST 1 M, 1 F [BMNH]. Ref.—Woodley, 2001: 163 (cat.). Cyphomyia albispinis Costa, 1866: 39. Nomen nudum. Ref.—Woodley, 2001: 163 (cat.). Cyphomyia cyanispis Bigot, 1876: 86. Nomen nudum. Ref.—Woodley, 2001: 163 (cat.). Cyphomyia cyanispinis Bigot, 1876: 487. Type locality: Brazil (“Amazonia”). HT M [BMNH]. Refs.— Woodley, 2001: 163 (cat.); Woodley, 2011: 410 (cat.). Cyphomyia wiedemanni ssp. peruana Lindner, 1933c: 329. Type locality: Peru, Iquitos, Roqué. HT F [SMF]. Ref.—Woodley, 2001: 163 (cat.). Cyphomyia wiedemanni ssp. burmeisteri Lindner, 1949a: 786. Nomen nudum. Ref.—Woodley, 2001: 163 (cat.). Cyphomyia wiedemanni ssp. burmeisteri Lindner, 1949b: 863. Type locality: Brazil, “Amazon”. ST 2 M [BMNH]. Ref.—Woodley, 2001: 164 (cat.). Cyphomyia viedemanni James, 1953: 311, error. Ref.—Woodley, 2001: 164 (cat.). Cyphomyia cyanispis Woodley, 2001: 163, error. Ref.—Woodley, 2011: 401 (cat.). Genus Euryneura Schiner Euryneura Schiner, 1868a: 308. Type species, Stratiomys fascipennis Fabricius, 1805 (orig. des.). propinqua Schiner, 1868b: 57. Type locality: Colombia. Distr.—Neotropical: Colombia, Mexico (Veracruz), Venezuela. Neartic: USA (Arizona). HT M [NMW]. Refs.—Giglio-Tos, 1893: 15; Cole, 1969: 158. Genus Labocerina Enderlein Labocerina Enderlein, 1914b: 579, 615. Type species, Stratiomys atrata Fabricius, 1805 (orig. des.). Labacerino Enderlein, 1914b: 615, error. Labocerino James, 1940: 124, error. atrata (Fabricius), 1805: 83 (Stratiomys). Type locality: “America meridionali”. Distr.—Colombia (Boyacá (Muzo)), Guyana, Mexico (Veracruz), Panama, Peru, Venezuela. ST M [UZMC]. Ref.—James, 1940: 125 (desc. of new species, key to species). Chordonota carbonaria Bellardi, 1862: 11. Type locality: Mexico. HT M [MIZT], destroyed. Ref.—Woodley, 2001: 168 (cat.). Genus Quichuamyia Brammer Quichuamyia Brammer, 2005: 2. Type species, Quichuamyia aplanatantennae Brammer, 2005 (orig. des.). aplanatantennae Brammer, 2005: 3, Figs 1–13. Type locality: Ecuador, Napo, Reservation Ethnica, Waorani, 1 km S of Onkone Gare. Distr.—Colombia (Amazonas (22 km NW Leticia)), Costa Rica (Heredia (La Selva 3 km S. Porto Viejo)), Cartago (Turrialba, tres Equis, P.N. Barbilla, 2 km S. de Estácion Barbilla)); Ecuador (Napo (Reservation Ethnica, Waorani, 1 km S of Onkone Gare)), (Misahualli near Tena), (Limacocha), (Coca, Napo R.)); Panama (Panama Province (Cerro Campana)); Peru (Monson Valley, Tingo Maria), (Quincemil Cuzco)). HT M [USNM]; AT F [USNM]; PT: 4 M, 3 F [USNM]; 3 F [USU]; 5 F [CNC]; 1 F [LACM]; 1 F [INBIO]; 1 F [CAS]. Ref.—Brammer, 2005: 9 (desc. of new species, key to species). 318 · Zootaxa 4122 (1) © 2016 Magnolia Press FACHIN & ASSIS-PUJOL Subfamily Chrysochlorininae Woodley, 2001 Genus Chrysochlorina James Chrysochlorina James, 1939a: 33. Type species, Sargus vespertilio Fabricius, 1805 (orig. des.). femoralis (Curran), 1929: 4 (Chrysochlora). Type locality: Colombia, Magdalena, Rio Frio. Distr.—Colombia (Magdalena (Rio Frio)), Mexico. HT M [AMNH]. Refs.—Curran, 1934: 315 (desc. of new species, key to species); James, 1939a (desc. of new species, key to species): 34; Iide, 1966: 112 (desc. of new species, key to species). frosti James, 1939a: 35. Type locality: Panama, Canal Zone, Barro Colorado Island. Distr.—Colombia (Boyacá (Muzo)), Panama (Canal Zone (Barro Colorado Island)). HT M [AMNH]. Refs.—James, 1939a (desc. of new species, key to species): 36; Iide, 1966: 112 (desc. of new species, key to species). haterius (Walker), 1849: 516 (Sargus). Type locality: “Venezuela”. Distr.—Colombia, Venezuela. ST 1 M, 1 sex? (stated F) [BMNH]. Ref.—Woodley, 2001: 176 (cat., syn.). Chrysochlora plana (Walker), 1851: 82. Type locality: “Colombia”. HT M (stated F) [BMNH]. Ref.—Smith & Chainey, 1989: 142 (syn.). varia (Curran), 1929: 3 (Chrysochlora). Type locality: Guyana, Bartica. Distr.—Bolivia, Brazil (Amazonas, Mato Grosso do Sul, Roraima), Colombia, Guyana (Bartica), Honduras, Panama, Suriname. HT M [AMNH]. Refs.— James, 1939a: 34 (desc. of new species, key to species); Iide, 1966: 112 (desc. of new species, key to species); Rafael, 1991: 331 (checklist). Subfamily Hermetiinae Loew, 1862 Genus Hermetia Latreille Hermetia Latreille, 1804: 192. Type species, Musca illucens Linnaeus, 1758 (mon.). Thorasena Macquart, 1838: 177. Type species, Hermetia pectoralis Wiedemann, 1824 (orig. des.). Stenothorax Agassiz, 1846: 369 (preocc. by Harris 1826; Harris 1827; Harris 1842), emend. of Thorasena Macquart, 1838. Massicyta Walker, 1856: 8. Type species, Massicyta bicolor Walker, 1856 (mon.). Massycyta Loew, 1860: 75, error. Masicyta Marschall, 1873: 337, error. Acrodesmia Enderlein, 1914c: 3. Type species, Acrodesmia luederwaldti Enderlein, 1914c = Hermetia albitarsis Fabricius, 1805 (orig. des.). Scammatocera Enderlein, 1914c: 5. Type species, Scammatocera virescens Enderlein, 1914c (orig. des.). Ref.—James, 1975: 31 (cat., syn.). illucens (Linnaeus), 1758: 589 (Musca). Type locality: “South America”. Distr.—Nearctic: USA (Alabama, Arizona, Arkansas, Califomia, Connecticut, Delaware, District of Columbia, Florida, Georgia, Indiana, Iowa, Kansas, Louisiana, Maryland, Massachusetts, Mississippi, Missouri, New Hampshire, New Jersey, North Carolina, Pennsylvania, South Carolina, Texas, Virginia) Neotropical: Argentina, Belize, Brazil (Amazonas, Bahia, Distrito Federal, Espírito Santo, Goiás, Mato Grosso, Mato Grosso do Sul, Minas Gerais, Pará, Paraná, Pernambuco, Rio de Janeiro, Rio Grande do Norte, Rio Grande do Sul, Roraima, Santa Catarina, São Paulo), British Virgin Islands, Chile, Colombia, Costa Rica, Ecuador, Dominica, Dominican Republic, El Salvador, Grenada, Guatemala, Guyana, Haiti, Honduras, Jamaica, Mexico (Chiapas, Chihuahua, Jalisco, Nuevo León, Oaxaca, Quintana Roo, San Luis Potosí, Sinaloa, Tamaulipas, Veracruz, Yucatán), Panama, Paraguay, Peru, Puerto Rico, Suriname, Trinidad, Venezuela, Uruguay. Palaearctic: Albania, Canary Islands, Croatia, France, Italy, Japan, Malta, Spain, Switzerland, Yugoslavia. Afrotropical: Cameroun, Congo, Ghana, Ivory Coast, Kenya, Madagascar, Mali, Namibia, South Africa, Tanzania, Zaire, Zambia. Oriental: India, Indonesia (lava, Sulawesi), Japan (Ryukyu Islands), Malaysia, Nepal, Philippines, Sri Lanka, Taiwan, Thailand, Vietnam. Australian: Australia (Queensland, Westem Australia), Belau, Bonin Islands, French Polynesia (Austral Islands, Society Islands), Guam, Hawaiian CATALOGUE OF STRATIOMYIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 319 Islands, Indonesia (Irian Jaya), Kiribati (Gilbert Island), Marshall Islands, Micronesia, New Caledonia, New Zealand, Northem Marianas, Papua New Guinea (Bougainville Island, Papua New Guinea), Solomon Islands, Vanuatu, Westem Samoa. ST sex? [NRS]. Refs.—Williston, 1888: 245; Lindner, 1928b: 239; Lindner, 1929: 260; Lindner, 1933c: 330; Carrera & Lane, 1945: 127 (checklist); James, 1949: 3; Iide & Mileti, 1976: 924–933 (redescription); Rafael, 1991: 331 (checklist); Woodley, 2001: 182 (cat.). Musca leucopa Linnaeus, 1767: 983. Type locality: “America”. ST 1 sex? [LSL]. Ref.—Woodley, 2001: 182 (cat.). Hermetia rufiventris Fabricius, 1805: 63. Type locality: “America meridionali”. ST 2 F [UZMC]. Ref.— Woodley, 2001: 182 (cat.). Hermetia pellucens Macquart, 1834: explication des planches 4: Planche V, Fig. 11, error. Ref.—Woodley, 2001: 182 (cat.). Hermetia nigrifacies Bigot, 1879a: 200. Type locality: “Mexico”. LT M [BMNH]. Refs.—McFadden, 1972b: 257; Woodley, 2001: 182 (cat.). Hermetia mucens Riley & Howard, 1889: 353, error. Ref.—Woodley, 2001: 182 (cat.). Hermetia illucens var. nigritibia Enderlein, 1914c: 9. Type locality: Brazil, Santa Catarina. HT sex? [PAN]. Ref.—Woodley, 2001: 182 (cat.). Hermetia illuscens Copello, 1926: 23, error. Ref.—Woodley, 2001: 182 (cat.). rufitarsis Macquart, 1846: 177. Type locality: “Colombia”. Distr.—Colombia. ST 1 F [MNHN]; ? [MHNLI]. Ref.—Woodley, 2001: 184 (cat.). tincta (Walker), 1851: 82 (Chrysochlora). Type locality: “Colombia”. Distr.—Colombia. HT F [BMNH]. Ref.— Woodley, 2001: 185 (cat., syn). Genus Patagiomyia Lindner Patagiomyia Lindner, 1933c: 331. Type species, Patagiomyia cyphomyioides Lindner, 1933b (orig. des.). cyphomyioides Lindner, 1933c: 331. Type locality: Peru, Iquitos, Mishujacu. Distr.—Bolivia, Colombia, Ecuador, Peru (Iquitos (Mishujacu). HT M [SMF]. Ref.—Woodley, 2001: 185 (cat.). Subfamily Nemotelinae Kertész, 1912 Genus Nemotelus Geoffroy Nemotelus Geoffroy, 1762: 450, 542. Type species, Musca pantherina Linnaeus, 1758 (sub. des. of I.C.Z.N. 1957a: 85). Nemotela Latreille, 1797: 164, error. Nemotilus Turton, 1801: 655, error. Nemoletus Say, 1823: 29, error. Nematotelus Agassiz, 1846: 248, emend. of Nemotelus Geoffroy. Nemoteles Siebke, 1863: 150, error. Akronia Hine, 1901: 113. Type species, Akronia frontosa Hine, 1901 (orig. des.). Ref.—Woodley, 2001: 310 (cat., syn.). Alisonia Williston, 1908: 171, error. Acronia Enderlein, 1914c: 2, error. Geitonomyia Kertész, 1923: 122. Type species, Geitonomyia transsylvanica Kertész, 1923 (orig. des.). Ref.—Woodley, 2001: 310 (cat., syn.). Epideicticus Kertész, 1923: 126. Type species, Nemotelus haemorrhous Loew, 1857 (orig. des.). Ref.—James, 1980: 262 (cat., syn.). Geitonomya Pleske, 1925b: 106, error. Galeodioptriger Lindner, 1933b: 3. Nomen nudum. Galeodioptriger Pleske in Lindner, 1937b: 108. Nomen nudum. Galeodioptriger Rozkošný, 1977: 10. Type species, Nemotelus bipunctatus Loew, 1846 (orig. des.). Described as subgenus of Nemotelus. 320 · Zootaxa 4122 (1) © 2016 Magnolia Press FACHIN & ASSIS-PUJOL Geitenomyia Berezovsky & Nartshuk, 1993: 97, error. Cluninemotelus Mason, 1997b: 49. Type species, Nemotelus clunipes Lindner, 1960 (orig. des.). Ref.—Woodley, 2001: 310 (cat., syn.). Paranemotelus Mason, 1997b: 106 Type species, Nemotelus exsul James, 1960 (orig. des.). Described as subgenus of Nemotelus. Temonelus Mason, 1997b: 110. Type species, Nemotelus grootaerti Mason, 1997b (orig. des.). Described as subgenus of Nemotelus. Subgenus Nemotelus Geoffroy lanatus James, 1974a: 6. Type locality: Colombia, Nariño, 10 miles E of Pasto Nariño, 3000 m. Distr.—Colombia (Nariño (10 miles E of Pasto Nariño, 3000 m)). HT F [CAS]. Ref.—Woodley, 2001: 317 (cat.). Nemotelus laniger James, 1974a: 2, error. Ref.—Woodley, 2001: 317 (cat.). Subgenus Camptopelta Williston Camptopelta Williston, 1917: 23. Type species, Camptopelta aldrichi Williston, 1917 (mon.). Nematotelus Séguy, 1926: 30. Type species, Nemotelus nigrinus Fallén, 1817 (mon.). Attributed to Osten Sacken. Nemotelinus Enderlein, 1936: 79. Type species, Nemotelus nigrinus Fallén, 1817 (mon.). Described as genus. Melanonemotelus Pleske in Lindner, 1937b: 107. Nomen nudum. Melanonemotelus Szilády, 1941: 100. Type species, Nemotelus nigrinus Fallén, 1817 (mon.). nudus Kertész, 1914: 525, Fig. 54. Type locality: Colombia, Barranquilla. Distr.—Brazil, Colombia (Atlantico (Barranquilla). HT F [HNHM], destroyed. Ref.—Woodley, 2001: 327 (cat.). Subfamily Pachygastrinae Loew, 1856 Genus Artemita Walker Artemita Walker, 1854: 61. Type species, Clitellaria amenides Walker, 1849 (sub. des., of Brauer, 1882: 86). Described as subgenus of Clitellaria Meigen, 1803. Artemida Loew, 1860: 75, error. Hypocerina Bigot, 1879a: 207. Nomen nudum. Psegmomma Enderlein, 1914a: 304. Type species, Acanthina argentea Osten Sacken, 1886b (orig. des). argentea (Osten Sacken), 1886b: 41 (Acanthina). Type localities: Costa Rica, Caché and Colombia (there is no lectotype designation). Distr.—Colombia (Chocó (Teresita)), Costa Rica, Ecuador (Napo (Coca, Napo River)), Guatemala, Mexico, Panama (Canal Zone (Barro Colorado Island), (San Blas Province)), Peru (Cuzco (Quincemil), Madre de Dios (Avispas)). ST 1 M (BMNH); ST M (ZMHU). Refs.—Kertész, 1914: 490 (desc. of fauna); James, 1971: 65 (desc. of new species, key to species); James, 1973: 26.50 (cat.); James et al., 1980: 10 (rev. of middle America fauna, desc. of new species, key to species). Psegmomma aureostriatum Enderlein, 1914a: 304. Type locality: Costa Rica. ST 18 F [PAN]. Ref.—Woodley, 2001: 83 (cat.). aurata (Macquart), 1846: 179 (Acanthina). Type locality: “Colombia”. Distr.—Colombia, Costa Rica, Ecuador (Napo (Coca, Napo River)), Panama (Darién (Cerro Pirre)), Peru (Madre de Dios (Avispas)), Venezuela. HT F [BMNH]. Refs.—Kertész, 1914: 489–490, Figs 28, 30 (desc. of fauna); James, 1971: 65 (desc. of new species, key to species); James et al. 1980: 10 (rev. of middle America fauna, desc. of new species, key to species). Acanthina aurata Bigot, 1879a: 207. Type locality: “Colombia”. HT F [BMNH]. Preocc., primary homonym of Acanthina aurata Macquart, 1846. hieroglyphica (Wiedemann), 1830: 50 (Acanthina). Type locality: “Brazil”. Distr.—Bolivia, Brazil (Amazonas, Pará, Roraima, Rio de Janeiro, Santa Catarina), Colombia, Costa Rica (Bataan, Batagu), El Salvador, Ecuador (Coca, Napo River), Guatemala (San Marcos (Finca La Paz, near La Reforma, 3.500 ft)), Guyana (Bartica), CATALOGUE OF STRATIOMYIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 321 Honduras, Mexico, Panama ((Bocas del Toro), Canal Zone (Barro Colorado Island)), Paraguay, Peru (20 mi W of Pucalpa), Suriname, Trinidad. ST F [ZMHU]. Refs.—Kertész, 1914: 484–485, Figs 29, 31 (desc. of fauna); Lindner, 1928b: 244; James, 1939d: 606; Lindner 1951a: 260 (desc. of fauna); James, 1971: 68 (desc. of new species, key to species); James et al. 1980: 11–12 (rev. of middle America fauna, desc. of new species, key to species); Rafael, 1991: 331 (checklist); Ururahy-Rodrigues & Pujol-Luz, 2007: 342–343, Figs 13–17 (redesc. of species). Acanthina ornata Macquart, 1846: 179. Type locality: Colombia. HT F [MNHN]. Ref.—Woodley, 2001: 84 (cat.). Clitellaria halala Walker, 1849: 523. Type locality: Honduras. HT F [BMNH]. Ref.—Woodley, 2001: 84 (cat.). Acanthina marmorata Gerstaecker, 1857: 337. Type locality: Brazil. ST 2 F [ZMHU]. Ref.—Woodley, 2001: 84 (cat.). Genus Caenacantha Wulp Caenacantha Wulp, 1885: 58. Type species, Caenacantha bipartita Wulp, 1855 (mon.). Spaniomyia Kertész, 1914: 462. Type species, Artemita pulchripennis Brauer, 1882 (orig. des.). Ref.—Woodley, 2011: 379 (cat., syn.). Pachycyanomyia McFadden, 1972b: 262. Nomen nudum. bipartita Wulp, 1885: 59. Type locality: Colombia, Cundinamarca, Bogotá. Distr.—Colombia (Cundinamarca (Bogotá)). HT F [RNH]. Ref.—Woodley, 2001: 90 (cat.). Genus Chalcidomorphina Enderlein Chalcidomorphina Enderlein, 1914a: 298. Type species, Chalcidomorphina aurata Enderlein, 1914a (orig. des.). Calcidomorphina Lindner, 1949a: 787, error. Chalcomorphina James, 1967a: 20, error. aurata Enderlein, 1914a: 299. Type locality: Colombia, Hacienda Pehlke. Distr.—Brazil (Rio de Janeiro, Roraima, Santa Catarina), Colombia (Caldas (Victoria, Hacienda Pehlke)), Costa Rica, Ecuador, Guyana (Bartica), Mexico (Chiapas (Palenque), (16 km S Malpasso), Veracruz (16 km W Tetzonapa), (Lake Catemaco)), Panama (Canal Zone (Barro Colorado Island), (Madden Lake Camp, Chagres)), Peru (Madre de Dios (Avispas), (Tingo Maria)), Venezuela. HT F [PAN]. Refs.—Lindner, 1951b: 260, 249, Fig. 1 (desc. of fauna); James, 1974b: 29 (desc. of fauna of the South America); James et al. 1980: 17–18 (rev. of middle America fauna, desc. of new species, key to species); Rafael, 1991: 332 (checklist); Pujol-Luz & Xerez, 1999: 298, Figs 1–9 (desc. of larva). Chalcidomorphina crewi Johnson, 1920: 115. Type locality: Guyana (Bartica). HT F [MCZ]. Ref.—Woodley, 2001: 91 (cat., syn.). Genus Ligyromyia Kertész Ligyromyia Kertész, 1916: 200. Type species, Ligyromyia columbiana Kertész, 1916 (orig. des.). columbiana Kertész, 1916: 201, 202, Fig. 42. Type locality: Colombia, Sierra San Lorenzo. Distr.—Argentina (Villa Padre Monti, Burruyacu), Brazil (São Paulo), Colombia (Magdalena (Sierra San Lorenzo), Peru, Trinidad. HT M [HNHM], destroyed. Refs.—Lindner, 1933a: 200; James, 1967b: 100 (desc. of fauna); Woodley, 2001: 115 (cat.). Ligyromyia colombiana James, 1967b: 100, error. Ref.—Woodley, 2001: 115 (cat.). 322 · Zootaxa 4122 (1) © 2016 Magnolia Press FACHIN & ASSIS-PUJOL Genus Panacris Gerstaecker Panacris Gerstaecker, 1857: 346. Type species, Panacris lucida Gerstaecker, 1857 (mon.). Comm.: Spyridopa Gerstaecker, 1857 is treated as synonym by James et al. (1980: 4) and Woodley (2011: 400), however, the genus is considered valid here. The latter author of this work examined the type of Spyridopa, Spyridopa tarsalis Gerstaecker, 1857, (Pujol-Luz & Assis-Pujol, 2002: 2). Tijucameru Pujol-Luz & Galinkin, 2004 also treated as synonym by Woodley (2011: 400) is considered valid here. This genus erected for Panacris maxima Kertész, 1908a was based in the genitalia differences, as well as the other morphological differences (Pujol-Luz & Galinkin, 2004: 35–36), when compared to the other type-species of Panacris and Spyridopa. Therefore, we recognize Spyridopa and Tijucameru as distinct genera. protrudens James in James, McFadden & Woodley, 1980: 6. Type locality: Panama, Canal Zone, Barro Colorado Island. Distr.—Colombia (Chocó (Teresita)), Costa Rica, Panama (Canal Zone (Barro Colorado Island)), Peru (Madre de Dios (Avispas), (Cuzco (Quincemil)). HT F [WSU]. Ref.—James et al., 1980: 6 (rev. of middle America fauna, desc. of new species, key to species). Genus Vittiger Kertész Vittiger Kertész, 1909: 395. Type species, Vittiger schnusei Kertész, 1909 (mon.). schnusei Kertész, 1909: 396. Type locality: Peru, “mouth of Pachitea River”. Distr.—Brazil (Rio de Janeiro, Santa Catarina), Colombia, Peru. ST 1 M, 1 F [UNK]. Refs. –Lindner, 1969: 4; Xerez & Pujol-Luz, 2001: 338, Figs 1–6 (desc of larva); Woodley, 2009: 547 (manual of fauna). Subfamily Raphiocerinae Schiner, 1868b Genus Lysozus Enderlein Lysozus Enderlein, 1914b: 579. Type species, Lysozus columbianus Enderlein, 1914b (orig. des.). Lyzozus Frey, 1934: 304, error. columbianus Enderlein, 1914b: 580. Type locality: Colombia, Hacienda Pehlke. Distr.—Colombia (Caldas (Victoria, Hacienda Pehlke)). HT sex? [PAN]. Ref.—James, 1943: 371 (desc. of new species, key to species). Genus Raphiocera Macquart Raphiocera Macquart, 1834: 253. Type species, Sargus armatus Wiedemann, 1830 (orig. des.). Hoplistes Macquart, 1834: 253. Type species, Sargus hoplistes Wiedemann, 1830 by absolute tautonymy (as synonym of Hoplistes pomaceus Macquart, 1834). Ref.—Pimentel & Pujol-Luz, 2001: 11 (rev. of subfamily, desc. of new species, key to species; syn.). Syn. by Pimentel & Pujol-Luz, 2001: 11. Basentidema Macquart, 1838: 197. Type species, Basentidema syrphoides Macquart, 1838 = Sargus hortulanus Wiedemann, 1830 (mon.) Ref.—Pimentel & Pujol-Luz, 2001: 11 (rev. of subfamily, desc. of new species, key to species; syn.). Syn. by Pimentel & Pujol-Luz, 2001: 11. Rhaphiocera Macquart, 1846: 182, error. Rhaphiocera Agassiz, 1846: 321, emend. of Raphiocera Macquart, 1834. Rhamphiocera Scudder, 1882: 292, emend. of Raphiocera Macquart, 1834. Rhapiocera Wulp, 1879: 9, error. mipartita (Macquart), 1846: 182 (Rhaphiocera). Type locality: Colombia. Distr.—Colombia. ST M [UNK]. Refs.—James, 1943: 379 (desc. of new species, key to species); Pimentel & Pujol-Luz, 2001: 28 (species placed as incertae sedis, stated as a probable synonym of Neoraphiocera pipopiuna Pimentel & Pujol-Luz, 2001; rev. of subfamily, desc. of new species, key to species); Woodley, 2001: 238 (cat.). Rhaphiocera bipartita Walker, 1854: 79, error. Ref.—Woodley, 2001: 238 (cat.). CATALOGUE OF STRATIOMYIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 323 ornata (Macquart), 1846: 183 (Rhaphiocera). Type locality: Colombia. Distr.—Bolivia, Colombia, Ecuador. ST 3 sex? [BMNH]. Refs.—James, 1943: 378 (desc. of new species, key to species); Pimentel & Pujol-Luz, 2001: 28 (species placed as incertae sedis; rev. of subfamily, desc. of new species, key to species). Subfamily Sarginae Walker, 1834 Genus Himantigera James Himantigera James in James & McFadden, 1982: 19. Type species, Himantigera silvestris McFadden, 1982 (orig. des.). dichroa (Schiner), 1868b: 62 (Chrysonotus dichrous). Type locality: Colombia. Distr.—Colombia. HT M [NMW]. Ref.—Woodley, 2001: 191 (cat.). Genus Merosargus Loew Merosargus Loew, 1855a: 144. Type species, Sargus obscurus Wiedemann, 1830 (orig. des.). Compsosoma Brauer, 1882: 79. Type species, Compsosoma chalconota Brauer, 1882 (orig. des.). Syn. by Fachin & Amorim, 2015: 24. Aloipha Enderlein, 1914b: 584. Type species, Aloipha cyaneoscutellata Enderlein, 1914b (orig. des.). Coenosargus Enderlein, 1914b: 587. Type species, Coenosargus azureus Enderlein, 1914b (orig. des.). Aloiphina Lindner, 1949a: 810. Type species, Aloiphina stigmatica Lindner, 1949a (mon.). Merusargus James & McFadden, 1971: 11, error. australis James in James & McFadden, 1971: 36 (Merosargus aurivena ssp. australis). Type locality: Colombia, Meta Department, Restrepo, 500 m. Distr.—Colombia (Meta (Restrepo, 500 m)), Peru, Venezuela. HT M [WSU]; AT 1 F (CNC). Refs.—James & McFadden, 1971: 36 (rev. of genus, desc of new species, key to species); Woodley, 2001: 194, 338 (cat., sub. des.). cingulatus Schiner, 1868b: 62. Type locality: “South America”. Distr.—Brazil (Minas Gerais, Paraná, Santa Catarina, São Paulo), Colombia, Costa Rica, Ecuador, EI Salvador, Guatemala, Honduras, Mexico (Campeche, Chiapas, Oaxaca, Tabasco, Veracruz, Yucatán), Nicaragua, Panama, Trinidad, Venezuela. ST 1 M, 5 F [NMW]. Refs.—Carrera & Lane, 1945: 128 (checklist); Woodley, 2001: 195 (cat.); Fontenelle et. al., 2012; Fachin & Amorim, 2015: 108 (taxonomic revision). complicatus James in James & McFadden, 1971: 43. Type locality: Colombia, Meta, 3 miles W of Villavicencio, 920 m. Distr.—Colombia (Meta (3 miles W of Villavicencio, 920 m)), Venezuela. HT M [CAS]; AT 1 F [WSU]; PT 1 F [WSU]. Ref.—James & McFadden, 1971: 43 (rev. of genus, desc of new species, key to species). frontatus Schiner, 1868b: 63. Type locality: Colombia. Distr.—Colombia. HT M [NMW]. Ref.—Woodley, 2001: 197 (cat.). gorgona (Lindner), 1949a: 814 (Acrochaeta). Type locality: Colombia, Gorgona Island. Distr.—Colombia (Gorgona Island). HT M [BMNH]. Ref.—Woodley, 2001: 197 (cat.). gowdeyi Curran, 1928: 33. Type locality: Colombia, Rio Frio. Distr.—Brazil (Minas Gerais), Colombia (Antioquia (Aljibes, Providencia, c.33 km. SW Zaragoza; Rio Frio), Vaupés (Rio Guayabero, Charco San José)), Costa Rica, Ecuador, Guatemala, Panama, Peru, Trinidad. HT M [BMNH]. Refs.—Woodley, 2001: 197 (cat.); Fontenelle et. al., 2012. Material examined. COLOMBIA: Antioquia: Aljibes, Providencia, c.33 km. Sv. Zaragoza, 1.xii. 1970, Richard W. Pinger, 1 F [USNM]. COLOMBIA: Vaupes: Rio Guayabero, Charco San José, 28.i.69, R.E. Dietz, 1 M [USNM]. 324 · Zootaxa 4122 (1) © 2016 Magnolia Press FACHIN & ASSIS-PUJOL gracillis Williston, 1888: 249. Type locality: Brazil, Mato Grosso, Chapada. Distr.—Brazil (Mato Grosso, Minas Gerais, Roraima, Santa Catarina, São Paulo), Colombia (Rio Raposo), Ecuador (Pastaza (Napo River), (Napo (Coca, Napo River)), Peru (Madre de Dios (Avispas), Cuzco (Quincemil), (Monson Valley, Tingo Maria)), Venezuela (Caracas). HT F [AMNH]. Ref.—James & McFadden, 1971: 27 (rev. of genus, desc of new species, key to species); Rafael, 1991: 331 (checklist); Iide & Pujol-Luz, 1999: 1–7 (redescription); Fontenelle et al., 2012. Material examined. COLOMBIA: Rio Raposo, v.1964, V.H. Lee. 1 M [USNM]. incanus James & McFadden, 1971: 11 (Merosargus lyricus ssp. incanus). Type locality: Peru, Cuzco, Quincemil, 700 m. Distr.—Colombia (Meta (3 mi. W Villavicencio), Costa Rica (Turrialba (La Suiza)), Peru (Cuzco (Quincemil, 700 m), (Madre de Dios (Avispas), (Monson Valley, Tingo Maria)). HT M [CNC]. Refs.—James & McFadden, 1971: 11 (rev. of genus, desc. of new species, key to species); Woodley, 2001: 194, 338 (cat., sub. des.). Merusargus lyricus ssp. incanus James & McFadden, 1971: 11, error. Merusargus lyricus ssp. incanus Woodley, 2001: 197, error. notatus (Lindner), 1949a: 807 (Coenosargus). Type locality: Colombia, Gorgona Island. Distr.—Colombia (Gorgona Island). HT sex? [BMNH]. Ref.—Woodley, 2001: 199 (cat.). quadrifasciatus Lindner, 1941: 178. Type locality: Peru, Silvia, 520 m. Distr.—Colombia, Ecuador, Peru. ST 1 M, 1 F [ZMUH], destroyed; 1 F [SMNS]. Ref.—Woodley, 2001: 200 (cat.). stamineus (Fabricius), 1805: 253 (Scaeva staminea). Type locality: “America meridionali”. Distr.—Belize, Brazil (Minas Gerais, Pará), Colombia, Costa Rica, Ecuador, El Salvador, Guatemala, Honduras, Mexico (Campeche, Chiapas, Hidalgo, Morelos, Nayarit, Oaxaca, Puebla, San Luis Potosí, Tabasco, Veracruz, Yucatán), Panama, Peru. HT M [UZMC]. Refs.—Lindner, 1929: 258; Lindner, 1933a: 201; Woodley, 2001: 200 (cat.). Sargus stramineus Osten Sacken, 1878: 45, error. Ref.—Woodley, 2001: 200 (cat.). Genus Microchrysa Loew Chrysomyia Macquart, 1834: 262. Type species, Musca polita Linnaeus, 1758 (sub. des. of Westwood, 1840: 130). Suppressed by I.C.Z.N. (1987: 148). Microchrysa Loew, 1855a: 146. Type species, Musca polita Linnaeus, 1758 (orig. des.). Clorisoma Rondani, 1856: 168. Type species, Sargus pallipes Meigen, 1830 = Sargus flavicornis Meigen, 1822 (orig. des.). Chrysomya Rondani, 1856: 168, error. Crhysomyia Bigot, 1857: 523, error. Chlorisoma Costa, 1857: 444, error. Chlorosia Rondani, 1861: 11 (nom. nov. for Clorisoma Rondani, 1856). Type species, Sargus pallipes Meigen, 1830 = Sargus flavicornis Meigen, 1822. Myochrysa Rondani, 1861: 11 (nom. nov. for Chrysomyia Macquart, 1834). Type species, Musca polita Linnaeus, 1758. Myochrysa Rondani, 1863: 28 (nom. nov. for Chrysomyia Macquart, 1834). Type species, Musca polita Linnaeus, 1758. Preocc. by Myochrysa Rondani, 1861. Microchryza Bigot, 1879a: 186, error. Chlorosoma Scudder, 1882: 71, emend. of Clorisoma Rondani. Myiochrysa Scudder, 1882: 218, emend. of Myochrysa Rondani. Myochrisa Bigot, 1887a: 29, error. Myiochrysa Aldrich, 1905: 180, error. Psaronius Enderlein, 1914b: 590. Type species, Psaronius viridis Enderlein, 1914b (orig. des.). Preocc. by Enderlein, 1912. Psaroninus Enderlein, 1937: 431 (nom. nov. for Psaronius Enderlein, 1914b). Type species, Psaronius viridis Enderlein, 1914b. Michrochrysa Ôuchi, 1940: 283, error. Mychrochrysa Csiby & Tóth, 1981: 200, error. Chrymichrosa Mason, 1997a: 31. Type species, Microchrysa elmari Lindner, 1960 (orig. des.). Described as subgenus of Microchrysa. bicolor (Wiedemann), 1830: 41 (Sargus bicolor). Type locality: “Crab Island” near Puerto Rico. Distr.—Argentina, Belize, Brazil (Minas Gerais, Pernambuco, Rio de Janeiro, Rio Grande do Norte, Roraima, Santa Catarina), Colombia (Cundinamarca (Monterredondo)), Costa Rica, Dominica, Dominican Republic, El Salvador, CATALOGUE OF STRATIOMYIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 325 Guatemala, Honduras, Jamaica, Mexico (Campeche, Chiapas, Colima, Guerrero, Morelos, Nayarit, Oaxaca, Sinaloa, Veracruz, Yucatán), Nicaragua, Panama, Puerto Rico, Uruguay. LT M [NMW]. Refs.—Lindner, 1929: 260; Lindner, 1933a: 201; James, 1949: 3; Lindner, 1951b: 249; McFadden, 1972a: 124 (lectotype designation); Rafael, 1991: 331 (checklist); Woodley, 2001: 203 (cat.). Chrysomyia rufiventris Macquart, 1834: 264. Type locality: “Guyane”. ST M [MNHN], lost. Microchrysa bruesi Johnson, 1919: 426. Type locality: Jamaica, Kingston, Liguanea Plain. HT M [MCZ]. Microchrysa abdominalis James, 1936: 51. Type locality: Panama, Canal Zone, Ancon. HT M [MCZ]. Microchrysa bicolor var. maculata Lindner, 1949a: 819. Type locality: Brazil, Santa Catarina, Nova Teutonia. ST 1 M, 1 F [BMNH], 2 M, F [UNK]. Material examined. COLOMBIA: Cundinamarca: Monterredondo, 16.xi.1960, J. Foerster, 1 M (CNC). Genus Ptecticus Loew Ptecticus Loew, 1855a: 142. Type species, Sargus testaceus Fabricius, 1805 (orig. des.). Ref.: Loew, 1855b: IX. Plecticus Loew, 1855a: 147, error. Plectiscus Loew, 1855a: plate, error. Pedicella Bigot, 1856: 63, 85. Type species, Sargus petiolatus Macquart, 1838 (sub. des. of Nartshuk & Rozkošný, 1984: 295). Ptecicus Loew, 1860: VII, error. Macrosargus Bigot, 1879a: 187 (nom. nov. for Pedicella Bigot, 1856). Type species, Sargus petiolatus Macquart, 1838. Pletiscus Scudder, 1884: 253, error. Gongrozus Enderlein, 1914b: 585. Type species, Gongrozus nodivena Enderlein, 1914b (orig. des.). Ref.: James, 1975: 18 (cat., syn.). Nacrosurgus Ôuchi, 1940: 281, error. Pteticus Lindner, 1965: 225, error. Petecticus Daniels, 1979: 578, error. Pteoticus Shi, 1992: 1116, error. conopsoides Schiner, 1868b: 66. Type locality: Colombia. Distr.—Colombia. ST M, F [NMW]. Ref.—Woodley, 2001: 211 (cat.). nigrifrons Enderlein, 1914b: 584 (Ptecticus testaceus var. nigrifrons). Type locality: Mexico, Chiapas. Distr.— Belize, Brazil (Santa Catarina), Colombia (Boyacá (Muzo, 900 m)), Costa Rica, El Salvador, Mexico (Chiapas, Jalisco, Morelos, Nayarit, Nuevo León, Oaxaca, San Luis Potosí, Tabasco, Tamaulipas, Veracruz), Panama, Peru. ST 4 M, 11 F [PAN]. Refs.—James, 1973: 26.17 (cat.); Woodley, 2001: 214 (cat.). Ptecticus nubilipennis James, 1938: 202. Type locality: Colombia, Boyacá, Muzo, 900 m. HT M [MCZ]. Ref.—Woodley, 2001: 214 (cat., syn.). tenuis Schiner, 1868b: 66. Type locality: Colombia. Distr.—Colombia. HT F [NMW]. Ref.—Woodley, 2001: 218 (cat.). terminalis James, 1938: 202. Type locality: Colombia, Boyaca, Muzo, 900 m. Distr.—Colombia (Boyacá (Muzo, 900 m)). HT M [MCZ]. Ref.—Leal & Oliveira, 1979: 77 (rev. of brasilian species). Genus Sargus Fabricius Sargus Fabricius, 1798: 549. Type species, Musca cupraria Linnaeus, 1758 (sub. des. of Latreille, 1810: 442). Chrysonotus Loew, 1855a: 146 (preocc. by Swainson, 1837). Type species, Musca bipunctata Scopoli, 1763 (orig. des.). Chrysochroma Williston, 1896: 47 (nom. nov. for Chrysonotus Loew, 1855a). Type species, Musca bipunctata Scopoli, 1763. Chrysonotomyia Hunter, 1900: 124 (nom. nov. for Chrysonotus Loew, 1855a). Type species, Musca bipunctata Scopoli, 1763. Geosargus Bezzi, 1907: 53 (nom. nov. for Sargus Fabricius, 1798). Type species, Musca cupraria Linnaeus, 1758. Surgus Ôuchi, 1938: 55, error. 326 · Zootaxa 4122 (1) © 2016 Magnolia Press FACHIN & ASSIS-PUJOL Pedicellina James, 1952: 225 (preocc. by Sars, 1835). Type species, Sargus notatus Wiedemann, 1830 = Sargus fasciatus Fabricius, 1805 (orig. des.). Himantoloba McFadden, 1970: 274. Type species, Chrysonotus flavopilosus Bigot, 1879b (orig. des.). Eumenogastrina Enderlein, 1914b: 589. Type species, Eumenogastrina angusta Enderlein, 1914b (orig. des.). Ref.—Woodley, 2011: 389 (cat., syn.). isthmi James in James & McFadden, 1982: 14. Type locality: Panama, Canal Zone, Barro Colorado Island. Distr.—Colombia (Chocó (Curiche River)), Costa Rica (Cartago), Panama (Canal Zone, Barro Colorado Island)). HT F [WSU]; PT 4 F [WSU]. Ref.—James & McFadden, 1982: 14 (rev. of subfamily on middle America, desc. of new species, key to species). thoracicus Macquart, 1834: 261. Type locality: “Amérique méridionale”. Distr.—Brazil (Mato Grosso, Paraná, Rio de Janeiro, Rio Grande do Norte, Santa Catarina, São Paulo), Colombia, Costa Rica, Guatemala, Honduras, Mexico (Campeche, Chiapas, Michoacán, Morelos, Nuevo León, Oaxaca, Puebla, Tamaulipas, Veracruz, Yucatán), Panama. ST F [UNK]. Refs.—Williston, 1888: 248–249; Lindner, 1933a: 201 (as Merosargus thoracicus); Lindner, 1933c: 328 (as Merosargus thoracicus); James 1941: 304 (key to species); James, 1949: 3; Viana & Xerez, 2003: 80 (description of puparium); Fachin & Amorim, 2015: 110 (taxonomic revision). Sargus concinnus Osten Sacken, 1886a: 23. Type locality: Guatemala, San Gerónimo. LT F [BMNH]. Refs.— McFadden, 1972b: 261; Woodley, 2001: 229 (cat.). stenus James in James & McFadden, 1982: 16. Type locality: Panama, Canal Zone, Barro Co1orado Island. Distr.—Colombia (Meta (5 km w Villavicencio)), Costa Rica (Heredia), Panama (Canal Zone (Barro Co1orado Island)). HT F [WSU]; PT 2 M, 2 F [KU]; 1 M [CAS]. Ref.—James & McFadden, 1982: 16 (rev. of subfamily on middle America, desc. of new species, key to species). Subfamily Stratiomyinae Latreille, 1802 Genus Hedriodiscus Enderlein Hedriodiscus Enderlein, 1914b: 608. Type species, Odontomyia brevifacies Macquart, 1855 (orig. des.). Hedriodiscina Enderlein, 1914b: 609. Type species, Stratiomys chloraspis Wiedemann, 1830 (sub. des. of James, 1953: 318). Hederodiscus Zack, 1984: 27, error. leucogaster (James), 1933: 69 (Odontomyia). Type locality: United States, Texas, Galveston. Distr.—Nearctic: United States (Califomia, Texas); Neotropical: Colombia, Mexico. HT M [SEMC]. Ref.—Woodley, 2001: 264 (cat.). quadrilineatus (Macquart), 1834: 248 (Odontomyia quadrilineata). Type locality: “Amérique méridionale”. Distr.—Argentina, Brazil, Colombia, Paraguay. ST F [UNK]. Ref.—Woodley, 2001: 264 (cat.). Odontomyia histrio Walker, 1851: 77. Type locality: Brazil. ST 1 F [BMNH]. Refs.—Lagrange, 1993: 70 (syn.); Woodley, 2001: 264. trivittatus (Say), 1829: 160 (Stratiomys trivittata). Type locality: Mexico. Distr.—Nearctic: United States (Alabama, Arizona, Califomia, Florida, Minnesota, Texas, Wisconsin); Neotropical: Colombia, Dominican Republic, El Salvador, Guatemala, Mexico (Chiapas, Colima, Guanajuato, Guerrero, Jalisco, Mexico, Michoacán, Morelos, Nayarit, Nuevo León, San Luis Potosí, Sinaloa, Sonora, Tabasco, Veracruz). ST F [UNK]. Ref.— Woodley, 2001: 265 (cat.). Odonthomyia tritaeniata Bellardi, 1859: 38. Type locality: Mexico. LT F [MIZT]. Ref.—Woodley, 2001: 265 (cat.). Odontomyia tritoeniata Giglio-Tos, 1893: 70, error. Ref.—Woodley, 2001: 265 (cat.). CATALOGUE OF STRATIOMYIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 327 Genus Hoplitimyia James Hoplitimyia James, 1934: 443. Type species, Stratiomys constans Loew, 1872 (orig. des.). Himantomyia James, 1939c: 39. Type species, Hoplitimyia clavata James, 1939b (orig. des.). Described as subgenus of Hoplitimyia. panamensis James in James & McFadden, 1979: 11. Type locality: Panama, Darién Province, Rio Indio. Distr.— Colombia (Valle del Cauca (64 km E of Buenaventura, 570 m), Ecuador, Panama (Darién Province (Rio Indio)). HT F [WSU]. Ref.—James & McFadden, 1979: 11 (rev. of subfamily on middle America, desc. of new species, key to species). Genus Odontomyia Meigen Eulalia Meigen, 1800: 21. Type species, Musca hydroleon Linnaeus, 1758 (sub. des. of Coquillett, 1910: 541). Suppressed by I.C.Z.N., 1963: 339. Odontomyia Meigen, 1803: 265. Type species, Musca hydroleon Linnaeus, 1758. (sub. des. of Westwood, 1840: 130). Odontomyja Fabricius, 1805: 79, error. Odontomya Latreille, 1809: 274, error. Odontomga Oken, 1815: plate XIV, error. Odonthomya Rondani, 1856: 170, error. Opseogymnus Costa, 1857: 443. Type species, Opseogymnus flavosignatus Costa, 1857 = Stratiomys limbata Wiedemann in Meigen, 1822 (orig. des.). Odonthomyia Bellardi, 1859: 32 error. Odontomeija Walker, 1860 error. Euceromys Bigot, 1877a: 88. Nomen nudum. Euceromys Bigot, 1877b: 102. Type species, Stratiomys nexura Walker, 1858 (mon.). Euceratomyia Rye, 1879: 191, emend. of Euceromys Bigot, 1877b. Cyrtopus Bigot, 1883: 132. Type species, Cyrtopus fastuosus Bigot, 1883 (mon.). Ref.: Woodley 2011: 459 (cat., syn.). Euceratomys Bigot, 1887b: CCV, error. Euceromyia Wulp, 1896: 52, error. Euceromyia Kertész, 1908b: 85, emend. of Euceromys Bigot, 1877b. Trichacrostylia Enderlein, 1914b: 607. Type species, Stratiomys angulata Panzer, 1798 (orig. des.). Neuraphanisis Enderlein, 1914b: 608. Type species, Stratiomys tigrina Fabricius, 1775 (orig. des.). Catatasina Enderlein, 1914b: 608. Type species, Stratiomys argentata Fabricius, 1794 (orig. des.). Scapanocnema Enderlein, 1914b: 610. Type species, Scapanocnema spathulipes Enderlein, 1914b (orig. des.). Ref.—Woodley, 2011: 459 (cat., syn.). Odomtomyia Matsumura, 1915: 44, error. Stratiomyiopsis Pleske, 1922: 332. Type species, Stratiomys ornata Meigen, 1822 (orig. des.). Described as subgenus of Eulalia. Hadracantha Pleske, 1922: 333. Type species, Stratiomys flavissima Rossi, 1790 (orig. des.). Described as subgenus of Eulalia. Clitellariopsis Pleske, 1922: 333. Type species, Stratiomys limbata Wiedemann in Meigen, 1822 (orig. des.). Described as subgenus of Eulalia. Achylomyia Pleske, 1922: 334. Type species, Musca microleon Linnaeus, 1758 (orig. des.). Described as subgenus of Eulalia. Zoniomyia Pleske, 1922: 334. Type species, Odontomyia pictifrons Loew, 1854 (orig. des.). Described as subgenus of Eulalia. Sratiomyiopsis Pleske, 1925a: 26, error. Odontomyiina Enderlein, 1930: 70. Type species, Stratiomys virgo Wiedemann, 1830 (orig. des.). Eularia Ôuchi, 1940: 275, error. Clytellariopsis Szilády, 1941: 92, error. Orthogonicera Lindner, 1951a: 187. Type species, Odontomyia hirayamae Matsumura, 1916 (mon.). Stratiodonta Lindner, 1972: 14. Type species, Odontomyia poecilopoda Bezzi, 1906 (mon.). Described as subgenus of Odontomyia. pachycephala Schiner, 1868b: 60. Type locality: Colombia. Distr.—Colombia. HT F [NMW]. Ref.—Woodley, 2001: 281 (cat.). Genus Stratiomys Geoffroy Stratiomys Geoffroy, 1762: 449, 475. Type species, Linnaeus, 1758 (sub. des. of I.C.Z.N., 1957b: 123). 328 · Zootaxa 4122 (1) © 2016 Magnolia Press FACHIN & ASSIS-PUJOL Hirtea Scopoli, 1763: 367. Type species, Hirtea longicornis Scopoli, 1763 (mon.). Stratiomiis Scopoli, 1777: 408, error. Ssratiomys Panzer, 1792: 14, error. Stratiomyx Rossi, 1794: 60, error. Stratiomis Walckenaer, 1802: 377, error. Stratyomis Schellenberg, 1803: 34, error. Stratyomys Griffith & Pidgeon, 1832: 700, error. Stratiomyia Macquart, 1838: 179, error. Hoplomyia Zeller, 1842: 828 [misprinted “882”] (nom. nov. for Stratiomys Geoffroy, 1762). Type species, Musca chamaeleon Linnaeus, 1758. Stratiomyia Agassiz, 1846: 354, emend. of Stratiomys Geoffroy, 1762. Hoptomyia Loew, 1846: 467, error. Thyreodontha Rondani, 1863: 77. Type species, Stratiomys strigata Fabricius, 1781 = Hirtea longicornis Scopoli, 1763 (orig. des.). Strutiomys Walker, 1864: 203, error. Stratiomya Schiner, 1868b: 51, error. Thyreodonta Marschall, 1873: 348, error. Stratiotomyia Rye, 1879: 188, emend. of Stratiomys Geoffroy, 1762. Stratomyia Bigot, 1887b: CCV, error. Stratiomia Hunter, 1900: 130, error. Thyreodonta Bezzi, 1903: 16, emend. of Thyreodontha Rondani, 1863. Oreomyia Pleske, 1922: 326. Type species, Stratiomys concinna Meigen, 1822 (sub. des. of Lindner, 1937a: 73). Holopyrrhoceromyia Pleske, 1922: 326. Type species, Stratiomys pyrrhocera Loew, 1869 = Stratiomys ruficornis Zetterstedt, 1842 (orig. des.). Described as subgenus of Oreomyia. Hemipyrrhoceromyia Pleske, 1922: 326. Type species, Stratiomys concinna Meigen, 1822 (orig. des.). Described as subgenus of Oreomyia. Laternigera Pleske, 1922: 327. Type species, Stratiomys furcata Fabricius, 1794 = Musca singularius M. Harris, 1778 (orig. des.). Described as subgenus of Stratiomyia. Eustratiomyia Pleske, 1922: 327. Type species, Musca chamaeleon Linnaeus, 1758 (orig. des.). Described as subgenus of Stratiomyia. Amauromyia Pleske, 1922: 328. Type species, Stratiomys lugubris Loew, 1871 (orig. des.). Described as subgenus of Stratiomyia. Poecilothorax Pleske, 1922: 329. Type species, Stratiomys nobilis Loew, 1871 (orig. des.). Described as subgenus of Stratiomyia. Diademomyia Pleske, 1922: 329. Type species, Stratiomys cenisia Meigen, 1822 (orig. des.). Described as subgenus of Stratiomyia. Metastratiomyia Pleske, 1922: 330. Type species, Stratiomys equestris Meigen, 1835 (orig. des.). Described as subgenus of Stratiomyia. Electrolophidion Pleske, 1922: 331. Type species, Stratiomys zarudnyi Pleske, 1899 (orig. des.). Described as subgenus of Stratiomyia. Parastratiomyia Pleske, 1922: 331. Type species, Stratiomys ventralis Loew, 1847 (orig. des.). Described as subgenus of Stratiomyia. Stereoxantha Pleske, 1922: 332. Type species, Stratiomyia sinensis Pleske, 1901 (orig. des.) Described as subgenus of Stratiomyia. Stiatiomyia Pleske, 1922: 332, error. Stratioyms Ôuchi, 1940: 272, error. Stratiomyis Ôuchi, 1940: 272, error. Strariomyia Shi, 1992: 1116, error. Comm.: We do not agree at all with the opinion of Woodley (2001: 346, footnote 126) on the synonymy of Stratiomicrodon Pujol-Luz & Papavero, 1998 with Stratiomys Geoffroy, 1762. Stratiomicrodon erected for Stratiomys convexa Wulp, 1881 has important diagnostic features when compared with other species of Stratiomys, especially by its four flagellomeres of the antenna and the scutellar spines very close on the scutellum, exclusive conditions of the genus. Therefore, we recognize the genus Stratiomicrodon as valid, with a single species Stratiomicrodon convexus (Wulp, 1881). constricta Walker, 1860: 269. Type locality: Mexico. Distr.—Colombia, El Salvador, Guatemala, Jamaica, Mexico (Chiapas, Tamaulipas, Veracruz), Nicaragua, Panama, Venezuela. HT M [BMNH]. Ref.—Woodley, 2001: 298 (cat.). Stratiomys pinguis Walker, 1860: 270. Type locality: Mexico. HT F [BMNH]. Ref.—Woodley, 2001: 298 (cat.). CATALOGUE OF STRATIOMYIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 329 Genus Zuerchermyia Woodley Zuercheria Lindner, 1928a: 102 (preocc. by Reitter, 1908). Type species, Zuercheria malachitis Lindner, 1928a (mon.). Zuerchermyia Woodley, 2001: 27, 306 (nom. nov. for Zuercheria Lindner, 1928a). Type species, Zuercheria malachitis Lindner, 1928a. pustulosa (James), 1938: 201 (Zuercheria). Type locality: Colombia, Magdalena Aracataca. Distr.—Colombia (Magdalena (Aracataca)). HT M [MCZ]. Ref.—James, 1938: 201–202 (desc. of new species). Acknowlegdments This study was supported by FAPESP Grants 2011/14472-6, 2013/02824-0 and 2014/05793-1. TABLE 1. Stratiomyidae species recorded to Colombia. Subfamily Genus Beridinae Chiromyzinae Clitellariinae Species Author Arcuavena singularis (Macquart, 1846) Heteracanthia ruficornis Macquart, 1850 Oplachantha albihirta James, 1977 albitarsis (Macquart, 1846) atricalx James, 1977 atriceps James, 1977 caerulescens (Schiner, 1868b) formosa Enderlein, 1921 pallida James, 1977 pulchella (Williston, 1888) viriata Enderlein, 1921 longicornis Enderlein, 1921 nigriventris Enderlein, 1921 scutellaris Enderlein, 1921 transequa Walker, 1850 diversicornis Kertész, 1923 fasciata Kertész, 1923 Chordonota leiophthalma Williston, 1896 Cyphomyia albitarsis (Fabricius, 1805) altifrons James, 1939d androgyna Osten Sacken, 1886b banksi James, 1937 chrysodota Perty, 1833ab cyanea (Fabricius, 1794) dispar Schiner, 1868b ecuadorensis Enderlein, 1914b erectispinis Lindner, 1935 fassli Lindner, 1949a formosa James, 1940 Nonacris Acropeltates Subgenus ...Continued on next page 330 · Zootaxa 4122 (1) © 2016 Magnolia Press FACHIN & ASSIS-PUJOL TABLE 1. (Continued) Subfamily Chrysochlorininae Hermetiinae Genus Subgenus Species Author geniculata Gerstaecker, 1857 planifrons James, 1939d schwarzi James, 1940 unicolor (Walker, 1854) varipes Gerstaecker, 1857 violacea Macquart, 1855 wiedemanni Gerstaecker, 1857 Euryneura propinqua Schiner, 1868b Labocerina atrata (Fabricius, 1805) Quichuamyia aplanatantennae Brammer, 2005 Chrysochlorina femoralis (Curran, 1929) frosti James, 1939a haterius (Walker, 1849) varia (Curran, 1929) illucens (Linnaeus, 1758) rufitarsis Macquart, 1846 tincta (Walker, 1851) cyphomyioides Lindner, 1933c Nemotelus lanatus (James, 1974a) Camptopelta nudus Kertész, 1914 argentea (Osten Sacken, 1886b) aurata (Macquart, 1846) hieroglyphica (Wiedemann, 1830) Caenacantha bipartita Wulp, 1885 Chalcidomorphina aurata Enderlein, 1914a Ligyromyia columbiana Kertész, 1916 Panacris protudens James in James, McFadden & Woodley, 1980 Vittiger schnusei Kertész, 1909 Lysozus columbianus Enderlein, 1914b Raphiocera mipartita (Macquart, 1846) ornata (Macquart, 1846) Himantigera dichroa (Schinner, 1868b) Merosargus australis James in James & McFadden, 1971 cingulatus Schiner, 1868b complicatus James in James & McFadden, 1971 frontatus Schiner, 1868b gorgona (Lindner, 1949a) gowdeyi Curran, 1928 Hermetia Patagiomyia Nemotelinae Pachygastrinae Raphiocerinae Sarginae Nemotelus Artemita ...Continued on next page CATALOGUE OF STRATIOMYIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 331 TABLE 1. (Continued) Subfamily Genus Species Author gracilis Williston, 1888 incanus James & McFadden, 1971 notatus (Lindner, 1949a) quadrifasciatus Lindner, 1941 stamineus (Fabricius, 1805) Microchrysa bicolor (Wiedemann, 1830) Ptecticus conopsoides Schiner, 1868b nigrifrons Enderlein, 1914b tenuis Schiner, 1868b terminalis James, 1938 isthmi James in James & McFadden, 1982 thoracicus Macquart, 1834 stenus James in James & McFadden, 1982 leucogaster (James, 1933) quadrilineatus (Macquart, 1834) trivittatus (Say, 1829) Hoplitimyia panamensis James in James & McFadden, 1979 Odontomyia pachycephala Schiner, 1868b Stratiomys constricta Walker, 1860 Zuerchermyia pustulosa (James, 1938) Sargus Stratiomyinae Hedriodiscus Subgenus References Agassiz, L. 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I–XVI, 440 pp. CATALOGUE OF STRATIOMYIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 341 Zootaxa 4122 (1): 342–345 http://www.mapress.com/j/zt/ Copyright © 2016 Magnolia Press Article ISSN 1175-5326 (print edition) ZOOTAXA ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.4122.1.27 http://zoobank.org/urn:lsid:zoobank.org:pub:9AC4EC48-CC2A-4D1B-A976-86FBEB87065A FAMILY XYLOMYIDAE DIEGO AGUILAR FACHIN1* & CRISTIANE VIERA DE ASSIS-PUJOL2 1 Departamento de Biologia, Faculdade de Filosofia, Ciências e Letras de Ribeirão Preto, Universidade de São Paulo, Av. Bandeirantes 3900, 14040–901, Ribeirão Preto, SP, Brazil. 2 Universidade Católica de Brasília, Laboratório de Zoologia, QS 7 lote 1, Bl. M/ s. 331, Taguatinga, 72030-170, DF, Brazil. * Corresponding author, E–mail: diegoafachin@gmail.com Abstract This family is poor known in Colombia with only two known species of two genera, Solva tuberculata and Arthropeina colombiana. Key words: Arthropeina, Catalogue, Colombia, diversity, Solva, Xylomyidae Introduction Xylomyidae Verral, 1901 are a family of lower Brachycera infraorder Stratiomyomorpha, sister-group of the Stratiomyidae (Woodley 1989, Woodley et al. 2009). These flies have body length ranging from 4 to 14 mm, and color quite varied. They are easily recognized by the presence of spurs on middle and hind tibia, by the conical antenna—with most of flagellomeres uniform in shape and color—, by the elongated discal cell, and by cell m3 closed before the wing margin. Quite few is known about the biology of these flies in the Neotropical region, but often the adults are associated with wooded areas and the immatures occur under bark of fallen trees (Webb 1984, Woodley 2009, 2011). Currently, there are 143 described species of Xylomyidae in the world, in four genera, with most of these species recorded to the Paleartic and Oriental regions, 46 and 50 described species, respectively (Woodley 2011, and more recent papers). Only 16 species in three genera have been recorded to the Neotropical Region: Arthropeina Lindner, 1949 (six species), Solva Walker 1859 (seven species) and Xylomya Rondani, 1861 (three species) (Table 1). Despite of the more recent work revising the genus Arthropeina (Fachin & Amorim, 2014), the family as a whole still is poorly known in the Neotropical Region. Papavero & Artigas (1991) were the first to discuss the genera relationships in Xylomyidae under a phylogenetic perspective, using mainly characters of female genitalia. Recently, Woodley (2011) proposed a new phylogenetic hypothesis for the Xylomyidae genera, making a reanalysis of some characters already available in the literature. In his proposal, Solva + Arthropeina and Coenomyiodes + Xylomya appear as sister-groups. The catalog records two species from Colombia: Solva tuberculata Webb, 1984 and Arthropeina colombiana Fachin & Amorim, 2014 (see below). The recorded species and the classification adopted were based mainly on information of the catalogue of Neotropical region (James 1975) and the world catalogue of the family (Woodley 2011). Additional literature also was consulted. The geopolitical departments are underline and specific localities are in brackets. The terms “allotype” and “paratype” were used in this catalogue only for the specimens that were designated as such in the original description. A summary table of the data given here is presented in Table 2. 342 Accepted by S. Nihei: 26 May 2015; published: 14 Jun. 2016 TABLE 1. Checklist of Neotropical species of Xylomyidae with known gender and geographic data. Species Gender Geographical distribution Male Female Arthropeina colombiana Fachin & Amorim - x Colombia Arthropeina diadelothorax Fachin & Amorim - x Brazil Arthropeina fulva Lindner x x Brazil Arthropeina lindneri Fachin & Amorim - x Guyana, Brazil Arthropeina melanochroma Fachin & Amorim x x Ecuador, Peru, Brazil Arthropeina pseudofulva Fachin & Amorim - x Brazil, Bolivia Solva brasiliana (Lindner) x - Brazil Solva decora Webb x x United States, Mexico, Costa Rica Solva flavicoxis (Enderlein) - x Mexico, Brazil, Paraguay Solva flavomaculata (Leonard) x - Mexico Solva pallipes (Loew) x x Canada, United States, Mexico Solva truncativena (Enderlein) x x Costa Rica, Panama Solva tuberculata Webb x x Mexico, Guatemala, Belize, Costa Rica, Colombia, Bolivia Xylomya elongata (Osten Sacken) x - Guatemala Xylomya fulviventris (Bigot) - x Mexico Xylomya prista Enderlein x - Mexico Acronyms used for the depositories AMNH—American Museum of Natural History, New York, New York, USA; CAS—California Academy of Sciences, San Francisco, California, USA; IAvH—Instituto de Investigación de Recursos Biológicos Alexander von Humboldt, Villa de Leyva, Colombia; CUID—Department of Entomology, Cornell University, Ithaca, New York, USA; FSCA—Museum of Entomology, Gainesville, Florida, United States; INHS—Illinois Natural History Survey, Champaign, Illinois, USA; KSU—Department of Entomology, Kansas State University, Manhattan, Kansas, USA; MSUEL—Department of Entomology, Michigan State University, East Lansing, Michigan, USA; UCD—Bohart Museum of Entomology, Davis, California, USA; USNM—National Museum of Natural History, Washington, D.C., USA; USU—Department of Biology, Utah State University, Logan, Utah, USA. List of abbreviations cat.—catalogue distr.—geographic distribution mon.—monotypy orig. des.—by original designation subs. des.—subsequent designation AT—allotype HT—holotype PT—paratype F (f#)—female M (m#)—male CATALOGUE OF XYLOMYIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 343 Catalogue of Xylomyidae of Colombia Family Xylomyidae Verral, 1901 Genus Arthropeina Lindner, 1949 Arthropeina Lindner, 1949: 789. Type species, Arthropeina fulva Lindner, 1949 (mon.). colombiana Fachin & Amorim, 2014: 244. Type locality: Colombia (Valle del Cauca (PNN [Parque Nacional Natural] Farallones de Cali, Cgto. La Meseta)). Distr.—Colombia (Valle del Cauca). Ref.—Fachin & Amorim, 2014: 239, Figs 21–26 (female terminalia); 247, Fig. 61 (thorax, dorsal view); 250, Figs 68–69 (wing and palpus); 251, Fig. 79 (dorsal habitus); 252, Fig. 85 (lateral habitus); 254, Fig. 92 (geographic distribution). HT F (f#) [IAvH]. Genus Solva Walker, 1859 Solva Walker, 1859: 98. Type species, Solva inamoena Walker, 1859 (mon.). Subulonia Enderlein, 1913: 545. Type species, Subulonia truncativena Enderlein, 1913 (orig. des.). Prista Enderlein, 1913: 546. Type species, Subula vittata Doleschall, 1859 (orig. des.). Ceratosolva de Meijere, 1914: 21. Type species, Ceratosolva cylindricornis de Meijere, 1914 (mon.). Pararthropeas Brunetti, 1920: 108. Type species, Pararthropeas thereviformis Brunetti, 1920 (orig. des.). Hanauia Enderlein, 1920: 281. Type species, Xylophagus marginatus Meigen 1820, (sub. des., Enderlein, 1921: 170). Hanavia Brunetti, 1923: 165, error. Formosolva James, 1939: 32. Type species, Solva concavifrons James, 1939 (orig. des.). Described as subgenus of Solva. Salva Ôuchi, 1943: 486, error. Phloophila Hull, 1945: 263. Type species, Subula pallipes Loew, 1863 (orig. des.). Described as subgenus of Solva. tuberculata Webb, 1984: 288. Type locality: Mexico (San Luis Potosí (15 miles N. Tamazunchale). Distr.—Nearctic: Mexico (Morelos, San Luis Potosí). Neotropical: Belize (Cayo), Bolivia (Beni), Colombia (Nariño), Costa Rica (Cartago, La Suiza, San Mateo), Guatemala, Mexico (Chiapas, San Luis Potosí, Veracruz), Peru. Refs.—Webb, 1984: 288–290 (description of the species); Woodley, 2011: 440 (cat.); CAS database. HT M (m#) [CAS]; AT 1 F (f#) [USU]; PT 28? [AMNH, CAS, CUI, FSCA, INHS, KSU, MSUEL, UCD, USNM, USU]. Acknowlegdments This study was supported by FAPESP Grants 2011/14472-6, 2013/02824-0 and 2014/05793-1. TABLE 2. Xylomyidae species recorded to Colombia. Genus Species Author Solva tuberculata Webb, 1984 Arthropeina colombiana Fachin & Amorim, 2014 References Brunetti, E. (1920) Diptera Brachycera. In: Shipley, A.E. (Ed.), The Fauna of British India, including Ceylon and Myanmar. Vol. 1. Taylor and Francis, London, pp. i–x + 1–401. Brunetti, E. (1923) Second revision of the Oriental Stratiomyidae. Record of the Indian Museum, 25 (1), 45–180. Doleschall, C.L. (1859) Derde bijdrage tot de kennis der dipteren fauna van Nederlandsch Indië. Natuurkundig Tijdschrift voor Nederlandsch-Indie, Djakarta, 17, 73–128. Enderlein, G. (1913) Dipterologische Studien. V. Zur Kenntnis der Familie Xylophagidae. Zoologischer Anzeiger, 42 (12), 533–552. 344 · Zootaxa 4122 (1) © 2016 Magnolia Press FACHIN & ASSIS-PUJOL Enderlein, G. (1920) 20. Ord. Diptera, Fliegen, Zweiflügler. In: Brohmer, P. (Ed.), Fauna von Deutschland. Ein Bestimmungsbuch unserer heimischen Tierwelt. Quelle & Meyer, Leipzig, pp. I–VIII + 265–315. Enderlein, G. (1921) Über die phyletisch älteren Stratiomyiidensubfamilien (Xylophaginae, Chiromyzinae, Solvinae, Beridinae und Coenomyiinae). Mitteilungen aus dem Zoologischen Museum in Berlin, 10 (1), 151–214. Fachin, D.A. & Amorim, D.S. (2014) Taxonomic revision of the Neotropical genus Arthropeina Lindner, 1949 (Diptera: Xylomyidae). Zootaxa, 3827 (2), 231–257. http://dx.doi.org/10.11646/zootaxa.3827.2.6 Hull, F.M. (1945) Notes upon flies of the genus Solva Walker. Entomological News, 55, 263–265. James, M.T. (1939) New Formosan Stratiomyidae in the collection of the Deutsches Entomologisches Institut. Arbeiten über Morphologische und Taxonomische Entomologie, Berlin-Dahlem, 6 (1), 31–37. James, M.T. (1975) Family Xylomyidae [Chapter] 25. In: Stone, A., Sabrosky, C.W., Wirth, W.W., Foote, R.H. & Coulson, J.R. (Eds.), A catalog of the Diptera of America north of Mexico. Agricultural Research Service, United States Department of Agriculture, Washington, D.C., pp. I–IV + 299–319. Lindner, E. (1949) Neotropische Stratiomyiiden des Britischen Museums in London. - Theil I. Annals and Magazine of Natural History, Series 12, 1 (11), 782–821. Loew, H. (1863) Diptera Americae septentrionalis indigena. Centuria tertia. Berliner. Entomologische Zeitschrift, 7 (1–2), 1–55. http://dx.doi.org/10.1002/mmnd.18630070104 Meigen, J.W. (1820) Systematische Beschreibung der bekannten europäischen zweiflügeligen Insekten. Zweiter Theil. Friedrich Wilhelm Forstmann, Aachen. I–X, 1–363. Meijere, J.C.H. de. (1914) Studien über südostasiatische Dipteren. VIII. Tijdschrift voor Entomologie, 56 (Supplement), 1–99. Ôuchi, Y. (1943) Contributiones ad Cognitionem Insectorum Asiae Orientalis 13. Notes on some dipterous insects from Japan and Manchoukuo. Shanghai Sizenkagaku Kenkyusyo Iho, 13 (6), 483–492. [in Japanese with English summary] Papavero, N. & Artigas, J.N. (1991) Phylogeny of the American genera of Solvidae (Xylomyidae) (Diptera), with illustrations of the female spermatheca. Gayana Zoologia, 55 (2), 101–113. Verrall, G.H. (1901) A list of British Diptera. University Press, Cambridg, 47 pp. http://dx.doi.org/10.5962/bhl.title.20297 Walker, F. (1859) Catalogue of the dipterous insects collected at Makessar in Celebes, by Mr. A.R. Wallace, with descriptions of new species [part]. Journal of the Proceedings of the Linnean Society, Zoology, 4 (15), 97–144. Webb, D.W. (1984) A revision of the Nearctic species of the family Solvidae (Insecta: Diptera). Transactions of the American Entomological Society, 110 (3), 245–293. Woodley, N.E. (1989) Phylogeny and classification of the “orthorrhaphous” Brachycera. In: McAlpine, J.F., Peterson, B.V., Shewell, G.E., Teskey, H.J., Vockeroth, J.R. & Wood, D.M. (Eds.), Manual of Neartic Diptera. Vol. 3. Research Branch Monograph 32, Agriculture Canada, Ottawa, pp. 1371–1395. Woodley, N.E. (2009) Xylomyidae [Chapter] 37. In: Brown B.V., Borkent, A., Cumming, J.M., Wood, D.M., Woodley, N.E. & Zumbado, M.A. (Eds.), Manual of Central American Diptera. Vol. 1. NCR Research Press, Boca Raton, Ottawa, pp. 517–519. Woodley, N.E. (2011) A World Catalog of the Xylomyidae (Insecta: Diptera). In: Thompson, F.C., Brake, I. & Lonsdale, O. (Eds.), Contributions to the Biosystematic Database of World Diptera. Myia, 12, pp. 417–453. Woodley, N.E., Borkent, A. & Wheeler T.A. (2009) Phylogeny of Diptera [Chapter 5]. In: Brown, B.V., Borkent, A., Cumming, J.M., Wood, D.M., Woodley, N.E. & Zumbado, M.A. (Eds.), Manual of Central American Diptera. Vol. 1. NCR Research Press, Boca Raton, Ottawa, pp. 79–94. CATALOGUE OF XYLOMYIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 345 Zootaxa 4122 (1): 346–349 http://www.mapress.com/j/zt/ Copyright © 2016 Magnolia Press Article ISSN 1175-5326 (print edition) ZOOTAXA ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.4122.1.28 http://zoobank.org/urn:lsid:zoobank.org:pub:57599AD6-9906-443B-B4D6-92B38F585F98 FAMILY ACROCERIDAE JÉSSICA PAULA GILLUNG Bohart Museum of Entomology, 1124 Academic Surge, University of California Davis, CA, USA, 95616. Email: jpg.bio@gmail.com Abstract A catalog of the Acroceridae from Colombia is provided. Five species and five genera are recorded for the country, including one undescribed Ocnaea species. Key words: catalogue, Colombia, Distribution, Neotropical, Spider flies, spider parasitoid Introduction Spider-flies are very diverse in shape, size and color, although all species have a greatly enlarged calypter and broad eyes that encompass almost the entire head. Some species are balloon-shaped with greatly swollen abdomens, while others may be slender bodied and extremely hunchbacked. Body size is particularly variable, ranging from 2 to 20 mm. Coloration may be metallic, varying from blue, purple, green, red, gold, orange, to yellow, while some species may be black or brown (Schlinger 1981, Nartshuk 1997). Some species mimic wasps, bees, and even beetles (e.g., Leucopsina, Mesophysa, Pterodontia) (Cole 1919). In general, species of Acrocerinae are small, have a minute head, scarce pilosity covering the body, and a swollen abdomen. Panopinae species, on the other hand, are usually bigger and covered by dense pilosity, and may have a slender or very swollen abdomen. Species of Philopotinae are very conspicuous flies with a hunchback body shape, in which the postpronotal lobes are greatly developed and fused medially forming a collar behind the head, and with a great variety of sizes and colors. In most acrocerid species, the eyes are holoptic in both sexes and may be pilose or bare. The antennae may be inserted at the upper portion of the head, in the middle of the frons, or at the lower part of the head. Proboscis length is extremely variable, ranging from longer than body length to smaller than the antennae; in some species, however, the mouthparts are lacking or are vestigial, not forming a proboscis. The wings may be hyaline or maculated, and the presence or absence of wing markings may be a dimorphic character in some species (e.g., Philopota). The wing venation is remarkably variable; some genera have a relatively complete set of wing veins, in which costa, R1, R2+3, R4, R5, M1, M2, M3, CuA1, CuA2 and A1 are present, while others have a varying degree of wing losses. In Acroceridae, cell r4+5 is generally bisected by crossvein 2r-m, and the presence of this additional crossvein may be a synapomorphy of the group. The costal vein can be complete, ending at the anal lobe, or it can be incomplete and end at the wing tip. In some genera, M3 is fused to CuA1 before wing margin, leading to the formation of a cell m3. The adults feed on nectar and may be efficient pollinators with variable mouthpart length, but some species do not feed. Sexual dimorphism takes place in many species, but in some species the sexes are quite indistinguishable. Immatures are internal parasitoids of spiders with a hypermetamorphic life cycle comprised of four larval instars. The first instar is a small, free-living planidium that either actively search for the spider host or sit and wait for it to pass by (Schlinger 1987, Cady et al. 1993). Acroceridae are worldwide distributed and comprise approximately 530 species in 55 genera (Schlinger et al. 2013). Neotropical fauna comprises all three subfamilies and is represented by 19 genera and approximately 100 species. There are five species and five genera recorded for Colombia, but the actual diversity is presumably much greater. The family is traditionally divided into three subfamilies (Acrocerinae, Panopinae and Philopotinae), but the monophyly of Acrocerinae has recently been questioned (Winterton et al. 2007). 346 Accepted by S. Nihei: 10 Apr. 2015; published: 14 Jun. 2016 Material from several collections in Brazil and in the USA was examined, including Evert I. Schlinger's collection, currently housed in the California Academy of Sciences (CAS). Within these material, only one specimen from Colombia was found, a new species of the genus Ocnaea. Thus, the list of Colombian species is based mainly on data from the literature. Three catalogs were consulted: Hunter (1901), Kertész (1909) and Arnaud (1979), and records obtained from the Diptera Site (Pape & Thompson 2011) were also included. Four described and one undescribed species are recorded for Colombia. The undescribed species belongs to the genus Ocnaea and was not included in the catalog. A list of the Colombian species is presented in Table 1. Acronyms used for the depositories CAS—California Academy of Sciences, San Francisco, CA, USA. OUMNH—Hope Entomological Collections, University Museum, Oxford, UK. List of abbreviations cat.—catalog diag.—diagnosis distr.—geographic distribution F (f#)—Female HT—holotype M (m#)—Male refs.—references orig. des.— by original designation Catalogue of Acroceridae of Colombia Family Acroceridae Leach, 1815 Subfamily Acrocerinae Schiner, 1868 Genus Ogcodes Latreille Ogcodes Latreille, 1797: 154. Type species, Musca gibbosa Linnaeus, 1758 (by subsequent monotypy (Latreille, 1802: 432)). colombiensis Schlinger, 1960: 311. Type locality: Colombia, Cundinamarca, 1 mile west of Villeta. HT M (CAS). Distr.: Colombia (Cundinamarca). Refs.: Arnaud, 1979: 204 (location of holotype). Subfamily Panopinae, Schiner, 1868 Genus Pterodontia Gray Pterodontia Gray, 1832: 779. Type species, P. flavipes Gray (by monotypy). analis Macquart, 1846: 226. Type locality: Colombia, "Region temperee". HT M (OUMNH, Coll. Bigot). Distr.: Colombia. Refs.: Walker, 1855: 347 (diag.); Hunter, 1901: 150 (cat.); Howard, 1902: 134 (note); Kertész, 1909: 6 (cat.); Carrera, 1946: 81 (distinction from P. flavonigra); Sabrosky, 1948: 392 (notes). dimidiata Westwood, 1876: 513. Type locality: “Colombia”. HT sex unspecified (OUMNH). Distr.: Colombia. CATALOGUE OF ACROCERIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 347 Refs.: Hunter, 1901: 151 (cat.); Kertész, 1909: 6 (cat.); Carrera, 1946: 81 (distinction from P. flavonigra); Sabrosky, 1948: 392 (note). Genus Pteropexus Macquart Pteropexus Macquart, 1846: 225. Type species, P. bicolor Macquart (orig. des.). bicolor Macquart 1846: 226. Type locality: “Colombia”. HT sex unspecified (OUMNH). Distr.: Colombia. Refs.: Walker, 1855: 335 (diag., synonymical notes); Wulp, 1882: 88, pl. 10, fig. 1 (drawing of wing); Kertész, 1909: 11 (cat.); Verrall, 1909: 451, fig. 259C (drawing of wing). TABLE 1. List of the Acroceridae recorded from Colombia. Subfamily Genus Species Author Acrocerinae Ogcodes colombiensis Schlinger, 1960 Panopinae Ocnaea sp. undescribed Pterodontia analis Macquart, 1846 dimidiata Westwood, 1876 bicolor Macquart, 1846 Pteropexus References Arnaud, P.H. Jr. (1979) A catalog of the types of Diptera in the collection of the California Academy of Sciences. Myia, 1, 1–505. Cady, A., Leech, R., Sorkin, L., Stratton, G. & Caldwell, M. (1993) Acrocerid (Insecta: Diptera) life histories, behaviours, host spiders (Arachnida: Araneida), and distribution records. The Canadian Entomologist, 125, 931–944. http://dx.doi.org/10.4039/Ent125931-5 Carrera, M. (1946) Moscas parasitas de aranhas. Chácaras e Quintais, 73, 174–178. Cole, F.R. (1919) The dipterous family Cyrtidae in North America. Transactions of the American Entomological Society, 45, 1–79. Gray, G.R. (1832) The class Insecta arranged by the Baron Cuvier, with supplementary additions to each order by Edward Griffith and Edward Pidgeon, and notices of new genera and species by George Gray, Esq, Volume second. In: Griffith, E. & Pidgeon, E. (Eds.), The animal kingdom arranged in conformity with its organisation by the Baron Cuvier with supplementary additions to each order. Whittaker, Treacher & Co., London, pp. 1–793. Howard, L.O. (1902) The Insect Book. Doubleday Page & Co., New York, 429 pp. Hunter, W.D. (1901) A catalogue of the Diptera of South America, Part II. Transactions of the American Entomological Society, 27, 121–155. Kertész, K. (1909) Catalogus Dipterorum hucusque descriptorum. Vol. 4. Oncodidae, Nemestrinidae, Mydaidae, Apioceridae, Asilidae. Museum Nationale Hungaricum, Budapest, 349 pp. Latreille, P.A. (1797) Précis des caractères génériques des insectes, disposés dans un ordre naturel. Prévôt, Paris, 201 pp. Latreille, P.A. (1802) Histoire naturelle, générale et particulière, des crustacés et des insectes. Tome troisième. Families naturelles des genres. Dufart, Paris, 467 pp. Macquart, J. (1846) Diptères exotiques nouveaux ou peu connus. Supplement. Mémoires de la Société des Sciences, de l'Agriculture et des Arts de Lille, 1844, 133–364. Westwood, J.O. (1876) Notae Dipterologicae. Descriptions of new genera and species of the family Acroceridae. Transactions of the Entomological Society of London, 3, 507–518. Nartshuk, E.P. (1997) Family Acroceridae. In: Papp, L. & Darvas, B. (Eds.), Manual of Palaearctic Diptera. Vol. 2. Science Herald, Budapest, pp. 469–485. Pape, T. & Thompson, F.C. (2011) Systema Dipterorum. Version 1.0. Available from: http://www.diptera.org/ (accessed 14 July 2013) 348 · Zootaxa 4122 (1) © 2016 Magnolia Press GILLUNG Sabrosky, C.W. (1948) A further contribution to the classification of the North American spider parasites of the Acroceratidae (Diptera). The American Midland Naturalist, 39, 382–430. http://dx.doi.org/10.2307/2421592 Schlinger, E.I. (1960) A revision of the genus Ogcodes Latreille, with particular reference to species of the Western Hemisphere. Proceedings of the United States National Museum, 111, 227–336. http://dx.doi.org/10.5479/si.00963801.111-3429.227 Schlinger, E.I. (1981) Acroceridae. In: Brown, B.V., Borkent, A., Cumming, J.M., Wood, D.M., Woodley, N.E., & Zumbado, M.A. (Eds.), Manual of Nearctic Diptera. Vol. 1. NRC Research Press, Ottawa, pp. 575–584. Schlinger, E.I. (1987) The biology of Acroceridae (Diptera): True endoparasitoids of spiders. In: Nentwig, W. (Ed.), Ecophysiology of spiders. Springer Verlag, Berlin, pp. 319–327. http://dx.doi.org/10.1007/978-3-642-71552-5_24 Schlinger, E.I., Gillung, J.P. & Borkent, C.J. (2013) New spider flies from the Neotropical Region (Diptera, Acroceridae) with a key to New World genera. ZooKeys, 270, 59–93. http://dx.doi.org/10.3897/zookeys.270.4476 Verrall, G.H. (1909) British flies, Vol. V, Stratiomyidae and succeeding families of the Diptera Brachycera of Great Britain. Gurney & Jackson, London, 780 pp. Walker, F. (1855) List of the specimens of dipterous insects in the collection of the British Museum. Part VI. Supplement II. British Museum, London, pp. 331–506. Winterton, S.L., Wiegmann, B.M. & Schlinger, E.I. (2007) Phylogeny and Bayesian divergence time estimations of small-headed flies (Diptera: Acroceridae) using multiple molecular markers. Molecular Phylogenetics and Evolution, 43, 808–832. http://dx.doi.org/10.1016/j.ympev.2006.08.015 Wulp, F.M. van der (1882) Americanishe Diptera, No. 1. Tijdschrift voor Entomologie, 25, 77–136. CATALOGUE OF ACROCERIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 349 Zootaxa 4122 (1): 350–352 http://www.mapress.com/j/zt/ Copyright © 2016 Magnolia Press Article ISSN 1175-5326 (print edition) ZOOTAXA ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.4122.1.29 http://zoobank.org/urn:lsid:zoobank.org:pub:74C56D01-388E-4864-85E6-75C5A0278A89 FAMILY NEMESTRINIDAE CECÍLIA KOSMANN1,* & CARLOS JOSÉ EINICKER LAMAS2 1 Depto. de Ciências Biológicas, Faculdade de Ciências e Letras de Assis - UNESP, Av. Dom Antonio, 2100, Assis, SP,19806-900, Brazil. 2 Museu de Zoologia da Universidade de São Paulo. Av. Nazaré 482, 04263-000, São Paulo, SP, Brazil. E-mail: ceciliakosmann@gmail.com; einicker@usp.br. *Corresponding author: ceciliakosmann@gmail.com Abstract Nemestrinidae (Diptera, Nemestrinoidea) are a relatively rare group of flies, and the knowledge on the Colombian fauna is still incipient. According to the available literature, only one species is recorded to this country, Hyrmophlaeba magdalena Bernardi, 1977. Key words: catalogue, Colombia, distribution, Nemestrinidae, tangle-veined flies Introduction The Nemestrinidae, also known as tangle-veined flies, comprise a group of brachycerous flies with a worldwide distribution (Bernardi 1973). The family has around 250 described species, distributed in 23 genera (Woodley 2009). In South America, five genera and 60 species are recorded (Bernardi 1973, 1977). The genera are geographically segregated and where they occur, they are dominant (Bernardi 1973). Besides the extant species, around 30 fossil species have been described, mainly from the Upper Jurassic and Cretaceous (Ansorge & Mostovski 2000). The Nemestrinidae are medium-sized flies (4–16 mm), and their body is usually covered with dense pilosity (Woodley 2009). The head is wide, sometimes wider than the thorax (Bernardi 1973), with large eyes, bare to densely pilose, ranging from clearly holoptic to widely dicoptic in males; mouthparts varying from vestigial to very long (Bernardi 1973; Woodley 2009). The thorax is unmodified, with long and slender legs (Woodley 2009). The wings are usually longer than the body, ranging from hyaline to opaque, with a complex and characteristic venation, which includes a so-called diagonal vein formed by many fusions of veins of the radial and medial areas (Bernardi 1973; Woodley 2009). Their unique wing venation pattern is considered as an autapormophy of the group (Yeates 1994). Adults frequently visit flowers of many different groups, feeding upon the nectar, and are known to live in open areas (Bernardi 1973). Larvae are endoparasites of grasshopper and beetles (Evenhuis 1994), and all species for which the biology is known have hypermetamorphosis (Woodley 2009). This catalogue and the classification adopted follow Papavero & Bernardi (2009), including information on type. Acronyms used for the depositories AMNH—American Museum of Natural History, New York, USA 350 Accepted by S. Nihei: 26 May 2015; published: 14 Jun. 2016 List of abbreviations cat.—catalogue dist.—geographic distribution orig. des.—by original designation refs.—references rev.—revision spp.—species subg.—subgenus syn.—synonym HT—Holotype F (f#)—female Catalogue of Nemestrinidae of Colombia Family Nemestrinidae Griffith & Pidgeon, 1832 Subfamily Hirmoneurinae Loew, 1860 Tribe Hyrmophlaebini Bernardi, 1976 Genus Hyrmophlaeba Rondani Hermoneura Philippi, 1865: 655. Incorrect orig. spelling. Hirmoneura, subg. Hyrmophlaeba Rondani, 1863: 51. Type species, Hirmoneura brevirostris Macquart, 1846 (orig. des.). Refs.: Bernardi, 1976: 32 (generic status), 1977: 248 (key to spp.). Hirmoneura, subg. Austrohirmoneura Bequaert, 1932: 15. Type species, Hermoneura bellula Philippi, 1865 (orig. des.). Refs.: Bernardi, 1976: 32 (syn.). Hirmophlaeba, Hirmophloeba, Hyrmophloeba, error. Refs.: Bernardi, 1977: 248-249 (key to non-Chilean spp.). magdalena Bernardi, 1977: 250. Type locality: Colombia, Magdalena, Rio Frio. HT F (AMNH). Distr.: Colombia (Magdalena). Refs.: Bernardi, 1977: 250 (rev.); Papavero & Bernardi, 2009: 7 (cat.). Material examined. COLOMBIA: Magdalena: Rio Frio, HT female, 19.xii.1926, G. Salt leg. (AMNH). TABLE 1. Nemestrinidae species recorded from Colombia. Subfamily Tribe Genus Species Author Hirmoneurinae Hyrmophlaebini Hyrmophlaeba magdalena Bernardi, 1977 References Ansorge, J. & Mostovski, M. (2000) Redescription of Prohirmoneura jurassica Handlirsch, 1906 (Diptera, Nemestrinidae) from the Lower Tithonian lithographic limestone of Eichstätt (Bavaria). Neues Jahrbuch für Geologie und Paläontologie Monastshefte, 4, 235–243. Bequaert, J. (1932) The Nemestrinidae (Diptera) in the V. v. Röder collection. Zoologische Anzeiger, 100, 13–33. Bernardi, N. (1973) The genera of the family Nemestrinidae (Diptera: Brachycera). Arquivos de Zoologia, 24, 211–318. http://dx.doi.org/10.11606/issn.2176-7793.v24i4p211-318 Bernardi, N. (1976) Classificação da subfamília Hirmoneurinae (Diptera, Nemestrinidae). Papéis Avulsos de Zoologia, São Paulo, 30, 25–33. Bernardi, N. (1977) A tribo Hyrmophlaebini (Diptera, Nemestridae, Hirmoneurinae), com treze espécies novas da América do Sul. Papéis avulsos de Zoologia, São Paulo, 30, 239–259. CATALOGUE OF NEMESTRINIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 351 Evenhuis, N.L. (1994) Family Nemestrinidae. In: Evenhuis, N.L. (Ed.), Fossil flies of the world (Insecta: Diptera). Backhuys, Leiden, pp. 313–315. Griffith, E. & Pidgeon, E. (1832) Part 34, in their The class Insecta arranged by the Baron Cuvier, with supplementary additions to each order by Edward Griffith, F. L. S., A. S. & Edward Pidgeon, Esq. and notices of new genera and species by George Gray, Esq. Volume the second, in Griffith, E., et al., The animal kingdom arranged in conformity with its organization by the Baron Cuvier with supplementary additions to each order. Vol. 15. Whittake, Treacher & Co., London, pp. 577–796. Loew, H. (1860) Die Dipteren-Fauna Südafrika’s. Erste Abtheilung. Abhandlungen des Naturwissenschaftlichen Vereins für Sachsen und Thüringen in Halle (1858–1861), 2, 57–402, pls. 1–2. [Also publ. sep., xi + 330 pp., 2 pls., Berlin, 1860] Macquart, J. (1846) Diptères exotiques nouveaux ou peu connus. [ler.] Supplément. Memoires de la Societe des Sciences de I' Agriculture et des Arts de Lille, (1845) 1844, 133–364, pls. 1–20. [Also separately published as Supplément [I], pp. 5–238, 20 pls, Paris, 1846] Papavero, N. & Bernardi, N. (2009) Catalogue of Neotropical Diptera, Nemestrinidae. Neotropical Diptera, 7, 1–16. Philippi, R.A. (1865) AufzählungderchilenischenDipteren. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien, 15, 595–782, pls. 1–7. Rondani, C. (1856) Dipterologiae italicae prodromus. Vol. I. Genera italica., A. Stoschi, Parmae [=Parma], 226 + [2] pp. http://dx.doi.org/10.5962/bhl.title.8160 Rondani, C. (1863) Diptera exotica revista et annotata. Novis nonnullis descriptis. E. Soliani, Modena, 99pp. [Also sep. publ., under the title: Dipterorum species et genera aliqua exotica, in Archivio Per La Zoologia, L'Anatomia E La Fisiologia, Modena, (1863) 3: 1–99, 1 pl., 1864] Woodley, N.E. (2009) Nemestrinidae. In: Brown, B.V., Borkent, A, Cumming, J.M., Wood, D.M., Woodley, N.E. & Zumbado, M.A. (Eds.), Manual of Central American Diptera. Vol. 1. NRC Research Press, Ottawa, pp. 557–560. Yeates, D.K. (1994) The cladistics and classification of the Bombyliidae (Diptera: Asiloidea). Bulletin of the American Museum of Natural History, 219, 1–191. 352 · Zootaxa 4122 (1) © 2016 Magnolia Press KOSMANN & LAMAS Zootaxa 4122 (1): 353–371 http://www.mapress.com/j/zt/ Copyright © 2016 Magnolia Press Article ISSN 1175-5326 (print edition) ZOOTAXA ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.4122.1.30 http://zoobank.org/urn:lsid:zoobank.org:pub:B30752DE-A5CB-42F5-8299-B4F25CD0CB84 FAMILY ASILIDAE MARTA WOLFF1 & CARLOS JOSÉ EINICKER LAMAS2 1 Grupo de Entomologia, Instituto de Biologia, Universidad de Antioquia. Colombia. E-mail: martha.wolff@udea.edu.com 2 Museu de Zoologia da Universidade de São Paulo. Av. Nazaré, 481, Ipiranga, 04263–000, São Paulo, SP, Brazil. E-mail: einicker@usp.br *Corresponding author: einicker@usp.br Abstract Asilidae is one of the largest Diptera families with more than 7,000 recognized species worldwide. All their species are predators on arthropods, mainly insects. This catalogue presents 71 species distributed in 26 genera, ten tribes or generic groups and four subfamilies. For each species we present the available geographical information and relevant references. Key words: Asilidae, Robber Flies, catalogue, Colombia, distribution, Biodiversity Introduction Asilidae are a distinct part of the global insect fauna with slightly more than 7,000 species distributed over all biogeographic regions, making Asilidae one of the most diverse taxa within Diptera (Dikow 2004). All species of Asilidae are predators: they feed on other insects, or occasionally other kinds of arthropods, such as arachnids. Predation is well known for the adult robber fly, and a large number of studies have investigated this behavior. Asilid larvae are likewise predaceous, based on available knowledge, which is far more limited than that for the adult (Fisher 2009). Colombia is listed as one of the world’s “megadiverse” countries, hosting close to 14% of the planet’s biodiversity. Colombia possesses a rich complexity of ecological, climatic, biological and ecosystem components, but only 72 species and 26 genera of robber flies are known from Colombia (Table 1), which represents less than 5% of the Neotropical fauna of this family. These numbers show how the Colombian fauna is still poorly known and emphasize the necessity of extensive sampling projects to be developed in this megadiverse country. This catalogue is primarily based upon the Neotropical catalogue of Papavero (2009). Acronyms used for the depositories AMNH—American Museum of Natural History, New York, USA. BMNH—The Natural History Museum, London, UK. CAS—California Academy of Sciences, San Francisco, California, USA. CNC—Canadian National Collection, Ottawa, Ontario, Canada. DZUC—Departamento de Zoología, Universidad de Concepción, Concepción, Chile. FMNH—The Field Museum of Natural History, Chicago, Illinois, USA. IZPAN—Instytut Zoologiczny Polskiej Akademii Nauk, Warsaw, Poland. KIEL—Zoologisches Museum, Christian-Albrechts Universität, Kiel, Germany. MACN—Museo Argentino de Ciencias Naturales, Buenos Aires, Argentina. MNHN—Museum National d’Histoire Naturelle, Paris, France. MNVL—Museum d’Histoire Naturelle de Ville de Lille, Lille, France. Accepted by S. Nihei: 26 May 2015; published: 14 Jun. 2016 353 MZUN—Museu dell’Istituto di Zoologia dell’Università di Napoli, Naples, Italy. NMW—Naturhistorisches Museum, Vienna, Austria. SMFD—Forschungsinstitut und Museum Senckenberg, Frankfurt, a.M., Germany. TORO—Instituto e Museo di Zoologia, Università di Torino, Turin, Italy. UDEC—Universidad de Concepción, Concepción, Chile. UMO—University Museum, University of Oxford, Oxford, UK. UNKN—Type depository unknown or not traced in this study. USNM—National Museum of Natural History, Washington, D.C., USA. ZMHB—Museum für Naturkunde, Humboldt-Universität, Berlin, Germany. ZMUC—Zoologisk Museum, University of Copenhagen, Copenhagen, Denmark. List of abbreviations aut.—automatic cat.—catalogue des.—designation distr.—geographic distribution emend.—emendations m.—meters misident.—misidentifications mon.—monotypy n. comb.—new combination nom. nov.—nomen novum (new name) orig. des.—by original designation pp.—pages preocc.—preoccupied refs.—references rev.—revision spp.—species sub.—subsequent subg.—subgenus HT—Holotype NT—Neotype LT—Lectotype ST—Syntype T—Type (unspecified) F (f#)—Female M (m#)—Male Catalogue of Asilidae of Colombia Family Asilidae Latreille, 1802 Subfamily Asilinae Latreille, 1802 Efferia-group Genus Albibarbefferia Artigas & Papavero Albibarbefferia Artigas & Papavero, 1997a: 68. Type species, Erax abibarbis Macquart, 1838 (orig. des.). Ref.: Papavero, 2009: 14 (cat.). 354 · Zootaxa 4122 (1) © 2016 Magnolia Press WOLFF & LAMAS Erax, barbatus-group of Hine, 1919: 108. Efferia, albibarbis-group of Wilcox, 1966: 104. ?heteroptera (Macquart), 1846: 211 (1846: 83), pl. 8, fig. 6 (Erax). Type locality: “Colombia” (as “Nouvelle Grenade”, cold region) HT f# (UMO). Refs.: Martin & Papavero, 1970: 65 (cat.; Efferia, as heteropterus, error); Artigas & Papavero, 1997a: 68 (n. comb.); Papavero, 2009: 15 (cat.). Genus Diplosynapsis Enderlein Diplosynapsis Enderlein, 1914a: 259. Type species, Diplosynapsis argentifascia Enderlein, 1914 (orig. des.). Ref.: Papavero, 2009: 17 (cat.). halterata Enderlein, 1914a: 260. Type locality: Colombia, Hacienda Pehlke, 900 m [cf. Papavero, 1973a: 299]. HT m# (IZPAN). Distr.: Colombia (Caldas (Victoria, hacienda Pehlke)). Refs.: Hull, 1962 (2): 479; Artigas & Papavero, 1997a: 70; Papavero, 2009: 17 (cat.). Genus Nerax Hull Nerax Hull, 1962 (2): 476. Type species, Asilus aestuans Linnaeus, 1758 (orig. des.). Ref.: Papavero, 2009: 9 (cat.). Erax Scopoli of authors (part), misident. Efferia Coquillett of authors, misident. Erax, aestuans-group of Hine, 1919: 120. Efferia, aestuans-group of Wilcox, 1966: 94. pictipennis (Schiner), 1868: 182 (Erax). Type locality: “South America” (Colombia). HT m# (NMW). Distr.: Colombia ?. Refs.: Hull, 1962 (2): 478 (n. comb.); Martin & Papavero, 1970: 67 (cat.); Artigas & Papavero, 1997a: 72; Papavero, 2009: 21 (cat.). Genus Triorla Parks Triorla Parks, 1968: 175. Type species, Asilus interruptus Macquart, 1834 (orig. des.). Refs.: Rueda, 2004: 88 (cat.); Papavero, 2009: 23 (cat.). interrupta (Macquart), 1834: 310 (Asilus). Type locality: U.S.A., Georgia. T (lost). Distr.: Colombia, Costa Rica, Guatemala, Honduras, Mexico, Nicaragua Panama, U.S.A. Refs.: Hull, 1962 (2): 478 (Nerax); Artigas & Papavero, 1997a: 73; Papavero, 2009: 24 (cat.). maculatus Macquart, 1838: 111 (1839: 227), pl. 9, figs. 6a-b (Erax), part. lateralis Macquart, 1838: 116 (1839: 232) (Erax). concolor Walker, 1851: 130 (Erax). perrumpens Walker, 1860: 283 (Asilus). villosus Bellardi, 1861: 149 (49) (Erax). perrumpeus Parks, 1968: 176; Papavero & Ibáñez-Bernal, 2001: 96 (history of Mexican dipterology), error. spinosa Tomasovic, 2002: 58, fig. 17. Type locality: “Colombia”. HT m# (MNVL). Ref.: Papavero, 2009: 24 (cat.). striola (Fabricius), 1805: 172 (Dasypogon). Type locality: “South America”. ST sex? (ZMUC) [cf. Zimsen, 1964: 471]. Distr.: Colombia, Guyana, Panama, Surinam south to Paraguay, Venezuela. Refs.: Williston, 1891: 87 (cat.); Hull, 1962 (2): 479 (Nerax); Martin & Papavero, 1970: 69 (cat.); Artigas & Papavero, 1997a: 90; Rueda, 2004: 88 (Cat.); Papavero, 2009: 24 (cat.). maculatus Macquart, 1838: 116 (1839: 232), pl. 9, figs. 6a-b (Erax), part. sicyon Walker, 1849: 399 (Asilus). CATALOGUE OF ASILIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 355 stimicon Walker, 1851: 129 (Erax). dilectus Walker, 1855: 632 (Erax). zetterstedti Jaennicke, 1867: 362 (Erax). Eicherax-group Genus Eicherax Bigot, Eristicus Loew, 1848: 396 (preocc. Wesmael, 1844). Type species, Erax nigripes Bellardi, 1861 (sub. des., Hull, 1962: 476) = Eicherax bellardii (Schiner, 1868). Eicherax Bigot, 1857: 548 (in key). Type species, Erax simplex Macquart, 1848 (orig. des.). Ref.: Papavero, 2009: 25 (cat.). Neoeristicus Osten Sacken, 1878: 81 (cat.; nom. nov. for Eristicus Loew, 1848). costatus (Schiner), 1868: 181 (Erax). Type locality: Colombia. ST sex? (NMW). Distr.: Colombia, Venezuela. Refs.: Williston, 1891: 85 (cat.; Erax); Hull, 1962 (2): 478 (Nerax); Martin & Papavero, 1970: 70 (cat.). fulvithorax (Macquart), 1838: 113 (1839: 229) (Erax). Type locality: Brazil, Pará, Belém, and French Guiana. ST sex? (MNHN). Distr.: Brazil, Colombia, French Guiana, Peru. Refs.: Hull, 1962 (2): 478 (Nerax); Lamas, 1972: 313; Artigas & Papavero, 1995a: 37; Papavero, 2009: 25 (cat.). parphorus Walker, 1851: 127 (Erax). Lecania-group Genus Ctenodontina Enderlein Ctenodontina Enderlein, 1914a: 260. Type species, Ctenodontina pectinatipes Enderlein, 1914 (orig. des.). Ref.: Martin & Papavero, 1970: 70 (syn.); Papavero, 2009: 30 (cat.). Catostola Hull, 1958c: 320. Type species, Catostola carrerai Hull, 1958 (orig. des.) = Ctenodontina maya Carrera & d´Andretta, 1953. pectinatipes Enderlein, 1914a: 261, figs. 6-7. Type locality: Colombia, Caldas, Victoria (Hacienda Pehlke), 900 m [cf. Papavero, 1973a: 299]. HT m# (IZPAN). Distr.: Colombia (Caldas). Refs.: Hull, 1962 (2): 480, figs. 2508, 2524; Martin & Papavero, 1970: 71 (cat.); Artigas & Papavero, 1995b: 36; Papavero, 2009: 30 (cat.). Mallophora-group Promachina Lehr, 1969: 532, 549, nomen nudum. Refs.: Papavero, 2009: 33 (cat.). Genus Mallophora Macquart, 1838 Mallophora Macquart, 1834: 300. Type species, Asilus bomboides Wiedemann, 1821 (sub. des., Coquillett, 1910: 565). Ref.: Artigas & Angulo, 1980 (rev.); Papavero, 2009: 33 (cat.). argentipes Macquart, 1838: 38 (1839: 204). Type locality: “Guyana”. HT m# (MNHN). Distr.: Argentina, Brazil, Colombia, French Guiana, Guyana, Panama, Paraguay, Surinam, Trinidad-Tobago, Venezuela. Refs.: Hull, 1962 (2): 468; Artigas & Papavero, 1997b: 101; Artigas & Hengst, 1999: 134 (key); Papavero, 2009: 36 (cat.). atra Macquart, 1834: 302. Type locality: “Brazil”. NeoType locality: Brazil, Santa Catarina, Nova Teutônia. T (lost). NT m# (DZUC) [des. by Artigas & Angulo, 1980: 34]. Distr.: USA (Minesota, Florida) to Argentina. Colombia. Refs.: Hull, 1962 (2): 468; Artigas & Papavero, 1997b: 101; Artigas & Hengst, 1999: 134 (key), Papavero, 2009: 36 (cat.). 356 · Zootaxa 4122 (1) © 2016 Magnolia Press WOLFF & LAMAS freycineti Macquart, 1838: 85 (1839: 201). ventralis Macquart, 1838: 91 (1839: 207). albifrons Walker, 1850a: xcviii. testaceipes Macquart, 1850: 380 (1850: 76). xylocopiides Walker, 1851: 111. xyocopides Walker, 1855: 579; error. testaceitarsis Macquart, 1854: 74 (1854: 54) . coeruleiventris Thomson, 1869: 467. nigra Williston, 1885: 58. lynchi Gemignani, 1930: 141. flavicollis Curran 1941: 272 (in key), nomen nudum. nitidula Curran, 1941: 272 (in key), nomen nudum. vorax Curran 1941: 272 (in key), 278. lucida Curran, 1941: 272 (in key), 278. curiosa Curran, 1941: 272 (in key), 279. jemima Curran, 1941: 272 (in key), 279. jemina Martin & Papavero, 1970: 74 (cat.), error. calida (Fabricius), 1787: 358 (Asilus). Type locality: Bolivia, Cordillera, San Antonio. T (KIEL) (only indeterminable remains) [cf. Zimsen, 1964: 468]. NT m# (DZUC). Distr. Mexico south America (except Chile), Colombia (Santander). Refs.: Hull, 1962 (2): 468; Artigas & Papavero, 1997b: 102; Artigas & Hengst, 1999: 134 (key), Papavero, 2009: 38 (cat.). nigritarsis Fabricius of Macquart, 1838: 86, misident. albicincta Bromley, 1929: 47. callida Curran, 1934a: 5, error abana Curran, 1934a: 5 (in key), 7. clavitarsis Curran, 1941: 271 (in key), 282. cingulata Artigas & Angulo, 1980: 53, figs. 2, 122, 136, 176. Type locality: Peru, Pucallpa, Rio Ucayali, 200 m. HT f# (UDEC). Distr.: Colombia, Panama, Peru. Refs.: Artigas & Papavero, 1997b: 102; Papavero, 2009: 38 (cat.). inca Curran, 1941: 270 (in key), 273. Type locality: Peru, Rio Santiago. HT f# (AMNH). Distr.: Brazil, Colombia, Paraguay, Peru. Refs.: Hull, 1962 (2): 469; Artigas & Papavero, 1997b: 102; Rueda, 2004: 86 (cat.); Papavero, 2009: 39 (cat.). minos (Wiedemann), 1824: 478 (Asilus). Type locality: Uruguay, Montevideo. NT m# (UDEC). Distr.: Belize, Brazil, Colombia, Ecuador, El Salvador, Guatemala, Mexico, Panama, Peru, Trinidad-Tobago, Uruguay, Venezuela. Refs.: Hull, 1962 (2): 469; Martin & Papavero, 1970: 75 (cat.); Artigas & Papavero, 1997b: 103; Papavero, 2009: 40 (cat.). craverii Bellardi, 1861: 122 (22). pulchella Curran, 1925: 260. schwarzi Curran, 1934a: 4. tolteca Curran, 1941: 272. pipiens Curran, 1941: 272. nigrifemorata Macquart, 1838: 90 (1839: 206). Type locality: “?Brazil”. HT f# (MNHN). Distr.: Argentina, Brazil, Panama, Colombia, Ecuador, Guyana, Surinam. Refs.: Hull, 1962 (2): 469 (as ?syn. of nigritarsis Fabricius, 1805); Artigas & Papavero, 1997b: 103; Artigas & Hengst, 1999: 135 (key); Papavero, 2009: 40 (cat.). tibialis Macquart, 1846: 77, pl. 8, fig. 3 (preocc. Macquart, 1838). cruralis Rondani, 1863: 46 (nom. nov. for tibialis Macquart, 1846). ada Curran, 1941: 171 (in key), 184. geijskesi Curran, 1942: 58. fairchildi Curran, 1942: 58. CATALOGUE OF ASILIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 357 nigritarsis (Fabricius), 1805: 168 (Dasypogon). Type locality: “South America (Smidt)” [i. e., Guyana, between rivers Demerara and Essequibo; cf. Papavero, 1971a: 21]. T (ZMUC) [cf. Zimsen, 1964: 470. [NT from Bolivia, Chapare, 400m DZUC, des. by Artigas & Angulo, 1980: 85, invalid]. Distr.: Argentina, Bolivia, Brazil, Colombia, Cuba, Ecuador, French Guiana, Guyana, Panama, Paraguay, Peru, Surinam, Trinidad-Tobago, Venezuela. Refs.: Hull, 1962 (2): 469; Artigas & Papavero, 1997b: 103, Artigas & Hengst, 1999: 135 (key); Papavero, 2009: 40 (cat.). cinerascens Walker, 1855: 581. socculata Rondani, 1848: 93. bruneri Bromley, 1929: 287. fusca Bromley, 1934: 350. pluto (Wiedemann), 1828: 477 (Asilus). Type locality: Argentina, Sierra Córdoba. NT m# (UDEC). Distr.: Argentina, Brazil, Colombia, Uruguay. Refs.: Hull, 1962 (2): 469; Artigas & Papavero, 1997b: 103; Artigas & Hengst, 1999: 135 (key); Rueda, 2004: 86 (cat.); Papavero, 2009: 41 (cat.). vegeta Lynch Arribálzaga, 1882: 140. copelloi Gemignani, 1930: 138. rufiventris Macquart, 1838: 91 (1839: 207). Type locality: unknown. HT f# (MNHN). Distr.: Argentina, Bolivia, Brazil, Colombia, French Guiana, Paraguay. Refs.: Hull, 1962 (2): 469; Artigas & Papavero, 1997b: 104; Artigas & Hengst, 1999: 135 (key); Rueda, 2004: 87 (cat.); Papavero, 2009: 42 (cat.). ?nigriventris Jaennicke, 1867: 361. tibialis Macquart, 1838: 85 (1939: 201). Type locality: Brazil, Pará, surroundings of Belém; French Guiana, “dans les grands bois aux bords de l’Oyapock”, and Cayenne. ST sex? (MNHN). Distr.: Argentina, Bolivia, Brazil, Colombia, French Guiana, Guyana, Peru, Surinam, Venezuela. Refs.: Hull, 1962 (2): 469; Artigas & Papavero, 1997b: 104; Artigas & Hengst, 1999: 135 (key); Rueda, 2004: 87 (cat.); Papavero, 2009: 43 (cat). fascipennis Macquart, 1850: 379 (1850: 75). fasciata Walker, 1851: 112. Genus Promachus Loew Trupanea Macquart, 1838: 91 (1839: 209) (preocc. Guettard, 1762). Type species, Asilus maculatus Fabricius, 1775 (orig. des.). Promachus Loew, 1848: 390. Type species, Asilus maculatus Fabricius, 1775 (sub. des., Coquillett, 1910: 595). Ref.: Papavero, 2009: 45 (cat.). fuscipennis (Macquart), 1846: 209 (1846: 81), pl. 8, fig. 4 (Trupanea). Type locality: “Colombia, warm region”. ST m# (UMO), sex? (MNHN). Distr.: Colombia, Venezuela. Refs.: Martin & Papavero, 1970: 80 (cat.); Artigas & Papavero,1997c: 105; Papavero, 2009: 45 (cat.). Ommatius-group Ommatiini Hardy, 1927: 384, 1927b: 388, 390. Type-genus, Ommatius Wiedemann, 1821. Ommatiinae Ricardo, 1929: 120. Ommatini Hull, 1962 (2): 433. Genus Ommatius Wiedemann Ommatius Wiedemann, 1821: 213. Type species, Asilus marginellus Fabricius, 1781 (sub. des., Coquillett, 1910: 579). Refs.: Scarbrough, 2008: 2-4 (key to species-groups); Papavero, 2009: 47 (cat.). Emphysomera Schiner of authors, misdent. 358 · Zootaxa 4122 (1) © 2016 Magnolia Press WOLFF & LAMAS ampliatus Scarbrough, 2002: 138 (in key), 140, figs. 1, 18-26. Type locality: Colombia, Cauca, 20 km W of Silvia (as Causa). HT m# (CNC). Distr.: Colombia (Cauca), Antioquia (Medellín)). Refs.: Papavero, 2009: 48 (cat.); Fisher, 2009: 624. angulosus Scarbrough, 2002: 139 (in key), 145, figs. 2, 27-35. Type locality: Colombia, Valle del Cauca, 6 miles W of Cali. HT m# (CAS). Distr.: Colombia (Valle del Cauca), Ecuador. Refs.: Papavero, 2009: 48 (cat.). dolabriformis Scarbrough, 2002: 139 (in key), 166, figs. 7, 78-83. Type locality: Colombia, San Sebastián de Rábago, Sierra Nevada de Santa María, 2000 m. HT m# (FMNH). Distr.: Colombia (Cesar (San Sebastian de Rábago)). Refs.: Papavero, 2009: 49 (cat.). rufipes Macquart, 1846: 216 (1846: 88) (preocc. Macquart, 1838). Type locality: “Colombia”. T (lost). Distr.: Colombia. Refs.: Williston, 1891: 90 (cat.); Hull, 1962 (2): 435; Martin & Papavero, 1970: 60 (cat.); Papavero, 2009: 51 (cat.). scopifer Schiner, 1868: 194. Type locality: “Colombia”. HT m# (NMW). Distr.: Colombia. Refs.: Williston, 1891: 90 (cat.); Hull, 1962 (2): 435; Martin & Papavero, 1970: 60 (cat.); Papavero, 2009: 52 (cat.). uncatus Scarbrough, 1993: 731 (in key to males), 733 (in key to females), 750, figs. 132-142. Type locality: Peru, Huánuco, San Domingo. HT m# (USNM) Distr.: Colombia (Amazonas (Rio Tacana), Brazil, Peru. Refs.: Scarbrough, 2007: 472; Papavero, 2009: 52 (cat.). Genus Taurhynchus Artigas & Papavero Taurhynchus Artigas & Papavero, 1995c: 148. Type species, Asilus xanthopterus Wiedemann, 1828 (orig. des.). Ref.: Papavero, 2009: 55 (cat.). aurolineatus (Macquart), 1846: 214 (1846: 86), pl. 8, fig. 9 (Proctacanthus). Type locality: “Colombia” (as “Nouvelle Grenade”). ST m# (lost) (UMO). Distr.: Colombia. Refs.: Hull, 1962 (2): 488 (Proctacanthus); Martin & Papavero, 1970: 77 (cat.); Papavero, 2009: 55 (cat.). bromleyi (Curran), 1931: 22 (Proctacanthus). Type locality: Colombia, Magdalena, Rio Frio, HT m# (AMNH). Distr.: Colombia (Magdalena (Rio Frio)), Venezuela. Refs.: Hull, 1962 (2): 488 (Proctacanthus); Papavero, 2009: 55 (cat.). flavipennis (Macquart), 1846: 213 (1846: 85), pl. 8, fig. 11 (Proctacanthus). Type locality: “Colombia”. T (lost). Refs.: Williston, 1891: 88 (cat.); Hull, 1962(2): 488 (Proctacanthus); Papavero, 2009: 55 (cat.). mystaceus (Macquart), 1846: 214 (1846: 86) (Proctacanthus). Type-localkity: “Colombia”. HT f# (lost) (UMO). Refs.: Hull, 1962 (2): 488 (Proctacanthus); Papavero,2009: 56 (cat.). salti (Curran), 1934a: 9 (in key), 11 (Proctacanthus). Type locality: Colombia, Magdalena, Rio Frío. HT m# (AMNH). Distr.: Colombia (Magdalena (Rio Frio)). Refs.: Hull, 1962 (2): 488 (Proctacanthus); Papavero, 2009: 56 (cat.). Proctacanthus-group Genus Eccritosia Schiner Eccritosia Schiner, 1866: 674. Type species, Asilus barbatus Fabricius, 1787 (orig. des.). Refs.: Lamas, 1973 (rev.); Papavero 2009: 53 (cat.). Ecoritosia Bromley, 1951: 25, error. CATALOGUE OF ASILIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 359 barbata (Fabricius), 1787: 358 (Asilus). Type locality: “French Guiana, Cayenne (Rohr)” [cf. Papavero, 1971a: 20]. T sex? (KIEL) [cf. Zimsen, 1964: 468]. Distr.: Argentina, Bolivia, Brazil, Colombia, Costa Rica, Guyana, Honduras, Mexico, Panama, Paraguay, Peru, Venezuela, U.S.A. Refs.: Artigas & Hengst, 1999: 135 (key); Rueda, 2004: 88 (cat.); Papavero 2009: 53 (cat.). amphinome Walker, 1849: 387 (Asilus), part. tricolor Walker, 1850: xcviii (Mallophora). Tribe Isopogonini Hardy, 1948 Genus Aphamartania Schiner Aphamartania Schiner, 1866: 671. Type species, Aphamartania frauenfeldi Schiner, 1866 (orig. des.). Ref.: Papavero, 2009: 53 (cat.). flavipennis (Macquart), 1846: 192 (1846: 64), pl. 7, fig. 4 (Dasypogon). Type locality: “Colombia” (as “Nouvelle Grenade”). HT f# (UMO). Distr.: Colombia. Refs.: Papavero, 1971b: 21 (n. comb.). Tribe Lastaurini Papavero, 1973b Genus Blepharepium Rondani Blepharepium Rondani, 1848: 89. Type species, Blepharepium luridum Rondani (mon.). Refs.: Papavero & Bernardi, 1973: 168 (key to spp.); Papavero, 2009: 66 (cat). Senobasis Macquart of aurthors, part, misident. Planetolestes Lynch Arribálzaga, 1879: 147. Type species, Laphria coarctata Perty, 1833 (orig. des.). One species known as fossil: Blepharepium borealis (James, 1939: 45; as Senobasis) (Oligocene; U. S. A., Colorado). annulatum (Bigot), 1857a: 331, pl. 20, figs. 3, 3a (Senobasis). Type locality: “Cuba”. HT m# (partially destroyed) (MNHN). Distr.: Colombia Antioquia (Medellín near Veracruz (as Mexico)), Cesar (Zapatosa (as Sapatosa region Chiriguaná)), Aruba, Belize, Bonaire, Costa Rica, Cuba, El Salvador, Guatemala, Honduras, Mexico, Panama, St. Eustatius Trinidad, Venezuela. Refs.: Hull, 1962 (1): 233; Papavero & Bernardi, 1973: 172, 174, 175, figs. 4-6, map 3; Papavero, 2009: 66 (cat). secabilis Walker, 1860: 276 (Dasypogon). cajennense (Fabricius), 1787: 360 (Asilus). Type locality: Guyana, Cuyuni-Mazaruni, Bartica, Kartabo. NT m# (MZUSP) [des. by Papavero & Bernardi, 1973: 178, 179]. Distr.: Colombia (Putumayo (La chorrera (as lacorezza)), (as Puntamayo), Bolivia, Brazil, Guyana, Peru, Surinam, Trinidad, Venezuela. Refs.: Hull, 1962 (1): 233; Lamas, 1972: 307 (cat.); Papavero & Bernardi, 1973: 181, figs. 1-17, map 1; Papavero, 2009: 66 (cat). cayennensis Gmelin, 1792: 162 (Asilus), error. inserens Walker, 1851: 91. auricincta Schiner, 1867: 371 (Senobasis). inca Curran, 1942: 54. vorax Curran, 1942: 53. Genus Diogmites Loew Diogmites Loew, 1866: 21 (Cent. 7, no. 36). Type species, Diogmites platypterus Loew, 1866 (sub. des., Coquillett, 1910: 533). Ref.: Papavero, 2009: 68 (cat.). Deromyia Philippi of authors, misident. amethystinus Carrera, 1953: 178, fig. 2. Type locality: Colombia, Meta, Restrepo, 500m. T sex?, Bomley Coll. 360 · Zootaxa 4122 (1) © 2016 Magnolia Press WOLFF & LAMAS (USNM?). Distr.: Colombia (Meta (Restrepo)). Refs.: Hull, 1962 (1): 232; Martin & Papavero, 1970: 24 (cat.); Papavero, 2009: 69 (cat.). castaneus (Macquart), 1838: 35 (1839: 151) (Dasypogon). Type locality: Brazil, Pará (i. e., Pará, Belém). HT f# (MNHN). Distr.: Bolivia, Brazil, Colombia, Peru. Refs.: Hull, 1962 (1): 232; Martin & Papavero, 1970: 25 (cat.); Lamas, 1972: 306 (cat.); Papavero, 2009: 69 (cat.). fulvus Bigot, 1878: 426 (Dasypogon). fulvis Bromley, 1946: 106 (cat.; Senobasis), error. winthemi (Wiedemann), 1821: 223 (Dasypogon). Type locality: “South America”. HT f# (NMW). Distr.: Brazil, Colombia. Refs.: Hull, 1962 (1): 232; Martin & Papavero, 1970: 27 (cat.); Papavero, 2009: 72 (cat). Genus Lastaurus Loew Dasypogon, subg. Lastaurus Loew, 1851: 11. Type species, Lastaurus anthracinus Loew, 1851 (orig. des.) = lugubris (Macquart, 1846). Refs.: Papavero, 2009: 73 (cat.). Morimna Walker, 1851: 104. Type species, Morimna mallophorides Walker, 1851 (mon.) = Lastaurus fallax (Macquart, 1838). Lastauropsis Carrera, 1949: 107. Type species, Lastauropsis villosus Carrera, 1949 (orig. des.). Refs.: Artigas & Papavero, 1988: 210 (syn.). fallax (Macquart), 1846: 191 (1846: 63), pl. 7, fig. 5 (Dasypogon). Type locality: “Colombia” (as “Nouvelle Grenade”). HT f# (UMO). Distr.: ?Argentina, ?Bolivia, Colombia, Ecuador, Peru, Venezuela. Refs.: Williston, 1891: 73 (cat.); Hull, 1962 (1): 239; Martin & Papavero, 1970: 30 (cat.); Lamas, 1972: 307 (cat.); Papavero, 2009: 73 (cat). bombimorpha Rondani, 1850: 368 (Dasipogon). mutabilis Loew, 1851: 12. mallophorides Walker, 1851: 104, pl. 6, figs. 2, 22 (Morimna). flavipellitus Enderlein, 1914b: 173 (as fallax var.). mallophoroides Kertész, 1909: 144. fenestratus Bigot, 1878: 424. Type locality: “Colombia” (as “Nouvelle Grenade”). HT f# (UMO). Distr.: Colombia. Refs.: Williston, 1891: 73 (cat.); Hull, 1962 (1): 239; Martin & Papavero, 1970: 30 (cat.); Papavero, 2009: 73 (cat.). lugubris (Macquart), 1846: 192 (1846: 64) (Dasypogon). Type locality: “Colombia” (as Nouvelle Grenade). T (lost). Distr.: ?Argentina, Colombia, Mexico, Venezuela. Refs.: Hull, 1962 (1): 239; Martin & Papavero, 1970: 30 (cat.); Artigas & Hengst, 1999: 137 (key); Papavero, 2009: 73 (cat.). anthracinus Loew, 1851: 12 (Dasypogon). Genus Senobasis Macquart Senobasis Macquart, 1838: 52 (1839: 168). Type species, Senobasis analis Macquart, 1838 (sub. des., Bromley, 1934: 332). Refs.: Papavero, 1975: 265 (key to spp.); Papavero, 2009: 78 (cat.). Stenobasis Agassiz, 1846: 338, emend. Astylium Rondani, 1850: 185. Type species, Astylium dorsiger Rondani, 1850 (orig. des.). Lochites Schiner, 1866: 655 (key, 671 (12868: 163; second erection of genus) (preocc. Cabanius & Heine, 1860). Type species, Laphria ornata Wiedemann, 1819 (orig. des.). Astylum Kertész, 1909: 83, emend. Stenobasis Kertész, 1909: 123, emend. Lochitomyia Brèthes, 1925: 105 (nom. nov. for Lochites Schiner, 1866). Type species, Laphria ornata Wiedemann, 1819 (aut.). aedon (Walker), 1849: 343 (Dasypogon). Type locality: “Venezuela”. T sex? (BMNH) Distr.: Colombia, CATALOGUE OF ASILIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 361 Venezuela. Refs.: Hull, 1962 (1): 106 (Stichopogon); Papavero, 1975: 269, 271 (key, cat.); Papavero, 2009: 79 (cat.). staurophora Schiner, 1868: 164 (Lochites). analis Macquart, 1838: 53 (1839: 169), pl. 5, figs. 1, 1a, 1b. Type locality: French Guiana, Mana river (as “Brésil, Lamana”, in error). HT m# (MNHN). Distr.: Colombia (Amazonas (Rio Tacana) (as Rio Tocana)), Brazil, French Guiana, Guyana, Peru. Refs.: Hull, 1962 (1): 422, figs. 675, 1276, 1286, 2107, 2131, 2165; Papavero, 1975: 271, figs. 28, 131-133; Papavero, 2009: 79 (cat.). tibialis Curran, 1934b: 332. corsair Bromley, 1951: 12. Type locality: Panama, Rio Indio. HT m# (USNM). Distr.: Colombia (Chocó (Pacific coast)), Costa Rica, Panama. Refs.: Hull, 1962 (1): 422; Papavero, 1975: 277, figs. 29, 45, 83, 149-151; Papavero, 2009: 79 (cat.). mendax Curran, 1934b: 333. Type locality: Guyana, Cuyuni-Mazaruni, Bartica, Kartabo. HT m# (AMNH). Distr.: Colombia (Amazonas (upper Rio Loreto, Yuca), Brazil, Ecuador, French Guiana, Guyana, Peru, Venezuela. Refs.: Hull, 1962 (1): 422; Papavero, 1975: 284, figs. 161-163; Papavero, 2009: 79 (cat). Tribe Atomosiini Lynch Arribálzaga, 1882 Genus Atomosia Macquart Atomosia Macquart, 1838: 73 (1839: 189). Type species, Atomosia incisuralis Macquart, 1838 (sub. des., Coquillett, 1010: 512) = puella (Wiedemann, 1828). Refs.: Papavero, 2009: 85 (cat.). Paratractia Hull, 1962 (1): 384. Type species, Laphria dasypus Wiedemann, 1828 (orig. des,). nigroaenea Walker, 1851: 154. Type locality: Colombia, Bogotá. HT f# (BMNH). Distr.: Colombia (Bogotá). Refs.: Martin & Papavero, 1970: 47 (cat.); Papavero, 2009: 87 (cat). Genus Cerotainia Schiner Cerotainia Schiner, 1866: 662 (key), 673 (1868: 170, second erection of genus). Type species, Laphria xanthoptera Wiedemann, 1828 (orig. des.). Refs.: Papavero, 2009: 90 (cat.). Ceratotaenia Lynch Arribálzaga, 1880: 52, emend. Cyphotomyia Williston, 1889: 257. Type species, Cyphotomyia lynchii Williston, 1889 (orig. des.). Protichisma Hermann, 1912: 35. Type species, Protichisma longimanus Hermann, 1912 (orig. des.). aurata Schiner, 1868: 171. Type locality: “Colombia”. HT m# (NMW). Refs.: Hull, 1962 (1): 395; Martin & Papavero, 1970: 51 (cat.); Papavero, 2009: 90 (cat.). Genus Dissmeryngodes Hermann Dissmeryngodes Hermann, 1912: 75. Type species, Laphria antica Wiedemann, 1828 (orig. des.). Refs.: Papavero, 2009: 92 (cat.). Centrolaphria Enderlein, 1914a: 241. Type species, Centrolaphria columbiana Enderlein, 1914 (orig. des.) = Dissmerynodes nigripes (Macquart, 1838). Distr.: Colombia (Caldas (Victoria, Hacienda Pehlke). Catonomyia Hull, 1962 (1): 380. Type species, Catonomyia spiculata Hull, 1962 (orig. des.). nigripes (Macquart), 1838: 74 (1839: 190) (Atomosia). Type locality: “Brazil”. HT f# (MNHN). Distr.: Brazil, Colombia, Peru. Refs.: Hull, 1962 (1): 379, fig. 651 (Atractia); Papavero, 2009: 92 (cat). dispar Walker, 1855: 570 (Atomosia). columbiana Enderlein, 1914a: 242 (Centrolaphria). 362 · Zootaxa 4122 (1) © 2016 Magnolia Press WOLFF & LAMAS Tribe Dasytrichini Lynch Arribálzaga Dasythricaria Lynch Arribálzaga, 1882: 245. Type genus, Dasythrix Loew, 1851. Refs.: Dikow, 2003: 465. Refs.: Papavero, 2009: 96 (cat.). Erecmoneminae Hermann, 1912: 21, part. Dasythrichini Artigas et al., 1988: 212, 237, as new, error. Genus Cryptomerynx Enderlein Cryptomerynx Enderlein, 1914a: 254. Type species, Cryptomerynx laphriicornis Enderlein, 1914 (orig. des.). Refs.: Papavero, 2009: 96 (cat.). Alipiolaphria Carrera, 1951: 116. Type species, Alipiolaphria mirandai Carrera, 1951 (orig. des.). laphriicornis Enderlein, 1914a: 254, figs. 1-2. Type locality: “Colombia, [Pehlke]”. HT m# (IZPAN). Distr.: Colombia (Caldas (Victoria, Hacienda Pehlke)). Refs.: Hull, 1962 (1): 344, figs. 2515, 2522; Papavero, 2009: 96 (cat). Tribe Laphriini Macquart, 1838 Genus Laphria Meigen Lapria Meigen, 1800: 25. Type species, Asilus gibbosus Linnaeus, 1758 (sub. des., Coquillett, 1910: 557). Suppressed by I. C. Z. N., 1963: 359. Refs.: Papavero, 2009: 99 (cat.). Laphria Meigen, 1803: 270. Type species, Asilus gibbosus Linnaeus, 1758 (sub. des.; Latreille, 1810: 443). Bombomima Enderlein, 1914a: 258. Type species, Laphria fulvithorax Fabricius, 1805 (orig. des.) = thoracica (Fabricius, 1805). melanogaster Wiedemann, 1821: 236. Type locality: “Mexico and Savannah”. Distr.: ?Colombia, Mexico, U.S.A. T (UNKN). Refs.: Hull, 1962 (1): 325 [Laphria (Bombomima)]; Papavero, 2009: 99 (cat). Unrecognized species of Laphriinae rubescens Bigot, 1878: 225 (Laphria). Type locality: “Colombia”. HT m# (UNKN). Refs.: Hull, 1962 (1): 323 (Laphria); Martin & Papavero, 1970: 38 (cat.); Papavero, 2009: 102 (cat). venezuelensis Macquart, 1846: 202 (1846: 74), pl. 7, fig. 15 (Laphria). Type locality: “Venezuela”. ST sex? (UMO). Distr.: Colombia, Venezuela. Refs.: Williston, 1891: 80 (cat.; Laphria); Hull, 1962 (1): 323 (Laphria); Martin & Papavero, 1970: 38 (cat.; Laphria). violacea Macquart, 1846: 202 (1846: 74) (Laphria). Type locality: “Colombia”. T (lost). Refs.: Williston, 1891: 80 (cat.); Hull, 1962 (1): 323 (Laphria); Martin & Papavero, 1970: 38 (cat.); Papavero, 2009: 102 (cat). Subfamily Laphystiinae Hendel, 1936 Genus Triclioscelis Roeder Triclioscelis Roeder, 1900: 337. Type species, Triclioscelis burmeisteri Roeder, 1900 (sub. des., Hermann, 1920: 183). Ref.: Curran, 1934a: 2 (key to spp.), Papavero, 2009: 105 (cat.). CATALOGUE OF ASILIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 363 salti (Curran), 1931: 11 (in key), 18 (Laphystia). Type locality: Colombia, Magdalena, Ciénaga. HT f# (AMNH). Distr.: Colombia (Magdalena). Refs.: Curran, 1934a: 2; Hull, 1962 (1): 76 (Laphystia); Martin & Papavero, 1970: 10 (cat.); Papavero, 2009: 106 (cat). Unrecognized species of Laphystiinae columbina Schiner, 1868: 175 (Laphystia). Type locality: “Colombia”. T (NMW). Refs.: Williston, 1891: 81 (cat.); Hull, 1962 (1): 76 (Laphystia); Martin & Papavero, 1970: 9 (cat.; Laphystia); Artigas et al., 1997: 53; Papavero, 2009: 106 (cat). Subfamily Leptogastrinae Schiner, 1862b Genus Leptogaster Meigen Leptogaster Meigen, 1803: 269. Type species, Asilus tipuloides Linnaeus, 1758 (mon.; misident.) = Leptogaster cylindrical (De Geer, 1776). Refs.: Papavero, 2009. 108 (cat.). Gonypes Latreille, 1805: 309. Type species, Asilus tipuloides Linnaeus, 1758 (mon.; misident.) = Leptogaster cylindrica (De Geer, 1776). One species known as fossil: Leptogaster prior Melander (1947: 44) (Oligocene; U. S. A., Colorado). nubeculosa Bigot, 1878: 455. Type locality: “Colombia”. HT m#? (UMO). Distr.: Colombia, Refs.: Williston, 1891: 68 (cat.); Kertész, 1909: 55 (cat.); Hull, 1962 (1): 299 (cat.); Papavero, 2009: 109 (cat). Genus Leptopteromyia Williston Leptopteromyia Williston, 1907: 1. Type species, Leptopteromyia gracilis Williston, 1907 (sub. mon., Williston, 1908: 195, figs. 35, 77). Refs.: Papavero 2009: 109 (cat.). colombiae Martin, 1971: 266 (in key), 269. Type locality: Colombia, Cundinamarca, 11 miles W of “Congueza”, 1340m. HT m# (CAS). Distr.: Colombia (Cundinamarca (Caquezá ) (as Conqueza error)). Refs.: Papavero, 2009: 110 (cat). Subfamily Trigonomiminae Enderlein, 1914b Genus Holcocephala Jaennicke Discocephala Macquart, 1838: 50 (1839: 166) (preocc. Jaennicke, 1932). Type species, Discocephala rufiventris Macquart, 1838 (sub. des., Coquillett, 1910: 534) = Holcocephala abdominalis (Say, 1823). Refs.: Papavero, 2009: 135 (cat.). Holcocephala Jaennicke, 1867: 359 (nom. nov. for Discocephala Macquart, 1838). Type species, Discocephala rufiventris Macquart, 1838 (aut.) = Holcocephala abdominalis (Say, 1823). Helcocephala Loew, 1874: 377, error. Holocephala Williston, 1891: 72 (cat.), error. Arthriticopus Enderlein, 1914b: 161. Type species, Arthriticopus nodosipes Enderlein, 1914 (orig. des.) Rhipidocephala Hermann, 1926: 174. Type species, Discocephala analis Macquart, 1846 (orig. des.). analis (Macquart), 1846: 197 (1846: 69) (Discocephala). Type locality: “Colombia”. T (lost). Distr.: Colombia. Refs.: Hull, 1962 (1): 65 (Rhipidocephala); Martin & Papavero, 1970: 6 (cat.); Papavero, 2009: 135 (cat.). ?apicalis Walker, 1854: 497 (Discocephala). nodosipes (Enderlein), 1914b: 161, fig. 4 (Arthriticopus). Type locality: Colombia: Cundinamarca, Fusagasugá, and Caldas, Victoria (Hacienda Pehlke), 900 m [cf. Papavero, 1973a: 299]; ST sex? (IZPAN). Distr.: Colombia (Caldas (Victoria), Cundinamarca (Fusagasugá)). Refs.: Hull, 1962 (1): 64 [Holcocephala (Arthriticopus)]; Martin & Papavero, 1970: 7 (cat.); Papavero, 2009: 136 (cat). 364 · Zootaxa 4122 (1) © 2016 Magnolia Press WOLFF & LAMAS Unrecognized and unplaced species of Asilidae albipilosus Macquart, 1846: 220 (1846: 92) (Asilus). Type locality: “Brazil” and Colombia, Bogotá. T (lost). Refs.: Williston, 1891: 90 (cat.; Asilus); Hull, 1962 (2): 545 (Asilus); Martin & Papavero, 1970: 88 (cat.); Papavero, 2009: 138 (cat.). caeruleiventris Macquart, 1846: 221 (1846: 93) (Asilus). Type locality: “Colombia”. T (lost). Refs.: Hull, 1962 (2): 545 (Asilus); Martin & Papavero, 1970: 88 (cat.; Asilus); Papavero, 2009: 138 (cat.). cinereum Bigot, 1878: 432 (Bathypogon). Type locality: “Colombia”. HT f# (UMO). Refs.: Hull, 1962 (1): 150 (Bathypogon); Martin & Papavero, 1970: 18 (cat.); Papavero, 2009: 139 (cat.). colombiae Macquart, 1838: 148 (1848: 264) (Asilus). Type locality: “Colombia”. HT f# (MNHN). Refs.: Hull, 1962 (2): 545 (Asilus); Martin & Papavero, 1970: 88 (cat.); Papavero, 2009: 139 (cat.). columbiae Williston, 1891: 90 (cat.; Asilus), error. flavipennis Macquart, 1846: 192 (1846: 64), pl. 7, fig. 4 (Dasypogon; preocc. Wiedemann, 1828). Type locality: “Colombia” (as Nouvelle Grenade). HT f# (UNKN). Refs.: Williston, 1891: 76 (cat.); Hull, 1962 (1): 228 (Dasypogon); Martin & Papavero, 1970: 89 (cat.); Papavero, 2009: 139 (cat.). gamaxus Walker, 1851: 142 (Asilus). Type locality: “Colombia”. HT m# (UNKN). Refs.: Williston, 1891: 90 (cat.); Martin & Papavero, 1970: 89 (cat.); Papavero, 2009:139 (cat.) . gammaxus Hull, 1962 (2): 545 (Asilus), error. lebasii Macquart, 1838: 148 (1848: 264) (Asilus). Type locality: “Colombia”. T (lost). Refs.: Williston, 1891: 90 (cat.); Hull, 1962 (2): 545 (Asilus); Martin & Papavero, 1970: 89 (cat.); Papavero, 2009: 139 (cat). lebasii Macquart, 1838: 38 (1839: 154) (Dasypogon). Type locality: “Colombia”. T (lost). Refs.: Hull, 1962 (1): 228 (Dasypogon); Martin & Papavero, 1970: 89 (cat.); Papavero, 2009: 139 (cat). tatius Walker, 1851: 148 (Asilus). Type locality: “Colombia”. T (lost). Refs.: Williston, 1891: 91 (cat.); Hull, 1962 (2): 545 (Asilus); Martin & Papavero, 1970: 91 (cat.); Papavero, 2009: 140 (cat). TABLE 1. Asilidae species recorded from Colombia. Subfamily Tribe or Generic group Genus Species Author Asilinae Efferia-group Albibarbefferia ?heterocera (Macquart, 1846) Diplosinapsis halterata Enderlein, 1914 Nerax commiles (Walker, 1851) pictipennis (Schiner, 1868) interrupta (Macquart, 1834) spinosa Tomasovic, 2002 stirola (Fabricius, 1805) costatus (Schiner, 1868) fulvithorax (Macquart, 1838) pectinatipes Enderlein, 1914 Triorla Eicherax-group Eicherax Lecania-group Ctenodontina Mallophora-group Mallophora argentipes Macquart, 1838 atra Macquart, 1834 calida (Fabricius, 1787) ...Continued on next page CATALOGUE OF ASILIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 365 TABLE 1. (Continued) Subfamily Tribe or Generic group Ommatius-group Genus Species Author cingulata Artigas & Angulo, 1980 inca Curran, 1941 minos (Wiedemann, 1824) nigrifemorata Macquart, 1838 nigritarsis (Fabricius, 1805) pluto (Wiedemann, 1828) rufiventris Macquart, 1838 tibialis Macquart, 1838 Promachus fuscipennis (Macquart, 1846) Ommatius ampliatus Scarbrough, 2002 angulosus Scarbrough, 2002 dolabriformis Scarbrough, 2002 rufipes Macquart, 1846 scopifer Schiner, 1868 uncatus Scarbrough, 1993 aurolineatus (Macquart, 1846) bromleyi (Curran, 1931) flavipennis (Macquart, 1846) mystaceus (Macquart, 1846) barbata (Fabricius, 1787) salti (Curran, 1934) Taurhynchus Proctacanthus-group Eccritosia Isopogonini Aphamartania flavipennis (Macquart, 1846) Lastaurini Blepharepium annulatum (Bigot, 1857) cajennense (Fabricius, 1787) amethystinus Carrera, 1953 castaneus (Macquart, 1838) wintemi (Wiedemann, 1821) fallax (Macquart, 1846) fenestratus Bigot, 1878 lugubris (Macquart, 1846) aedon (Walker, 1849) analis Macquart, 1838 corsair Bromley, 1951 mendax Curran, 1934 Atomosia nigroaenea Walker, 1851 Cerotainia aurata Schiner, 1868 Dissmeryngodes nigripes (Macquart, 1838) Cryptomeryns laphriicornis Enderlein, 1914 Laphria melanogaster Wiedemann, 1821 rubescens Bigot, 1878 violacea Macquart, 1846 venezuelensis Macquart, 1846 Diogmites Lastaurus Senobasis Atomosiini Dasytrichini Unrecognized species of Laphriidae ...Continued on next page 366 · Zootaxa 4122 (1) © 2016 Magnolia Press WOLFF & LAMAS TABLE 1. (Continued) Subfamily Tribe or Generic group Laphystiinae Genus Species Author Triclioscelis salti (Curran, 1931) columbina Schiner, 1868 Leptogaster nubeculosa Bigot, 1878 Leptopteromyia colombiae Martin, 1971 Holcocephala analis (Macquart, 1846) nodosipes (Enderlein, 1914) albipilosus Macquart, 1846 caeruleiventris Macquart, 1846 cinereum Bigot, 1878 colombiae Macquart, 1838 flavipennis Macquart, 1846 gamaxus Walker, 1851 lebasii Macquart, 1838 lebasii Macquart, 1838 tatius Walker, 1851 Unrecognized species of Laphystiinae Leptogastrinae Trigonomiminae Unrecognized and unplaced species of Asilidae References Agassiz, L. (1846) Nomenclator zoologicus, continens nomina systematica generum animalium tam viventium quam fossilium, secundum ordinem alphabeticum disposita, adjectis auctoribus, libris, in quibus reperiuntur, anno editionis, etymologia et familiis, ad quas pertinent, in singulis classibus. 1842-1846 [1847]. Jent et Gassmann, Soloduri [=Solothurn], 393 pp Artigas, J.N. & Angulo, A.O. (1980) Revisión del gênero Mallophora Macquart por sistemática alfa y numérica (Diptera–Asilidae). Gayana, Zoologia, 43, 1–182. Artigas, C.J.N. & Hengst, M. (1999) Clave ilustrada para los géneros de asílidos argentinos. Revista Chilena de Historia Natural, 72 (1), 107–150. Artigas, J.N. & Papavero, N. (1995a) The American genera of Asilidae (Diptera): Keys for identification with an atlas of female spermathecae and other morphological details. IX.8. Subfamily Asilinae Leach, Eicherax -group, with a catalogue of the Neotropical species. Boletin de la Sociedad de Biologia de Concepcion, 66, 35–42. Artigas, J.N. & Papavero, N. (1995b) The American genera of Asilidae (Diptera): Keys for identification with an atlas of female spermathecae and other morphological details. IX.4. Subfamily Asilinae Leach, Glaphyropyga- group, with the proposal of two new genera and a catalogue of the Neotropical species. Boletin de la Sociedad de Biologia de Concepcion, 66, 11–33. Artigas, J.N. & Papavero, N. (1995c) The American genera of Asilidae (Diptera): Keys for identification with an atlas of female spermathecae and other morphological details. IX.7. Subfamily Asilinae Leach, Proctacanthus group, with the proposal of a new genus and a catalogue of the Neotropical species. Gayana Zoologia, 59 (2), 145–160. Artigas, J.N. & Papavero, N. (1997a) The American genera of Asilidae (Diptera): Keys for identification with an atlas of female spermathecae and other morphological details. IX.1. Subfamily Asilinae Leach (including Apocleinae Lehr). Key to generic groups. Arquivos de Zoologia, 34 (2), 57–63. http://dx.doi.org/10.11606/issn.2176-7793.v34i1-4p57-63 Artigas, J.N. & Papavero, N. (1997b) The American genera of Asilidae (Diptera): Keys for identification with an atlas of female spermathecae and other morphological details. IX.2. 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Hathaway, New Haven, 405 pp. [James, T (Ed.)] http://dx.doi.org/10.5962/bhl.title.20989 Wilcox, J. (1966) Efferia Coquillett in America north of Mexico (Diptera: Asilidae). Proceedings of the California Academy of Sciences, 39 (2), 85–234. Zimsen, E. (1964) The type material of I. C. Fabricius. Munksgaard, Copenhagen, 656 pp. CATALOGUE OF ASILIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 371 Zootaxa 4122 (1): 372–381 http://www.mapress.com/j/zt/ Copyright © 2016 Magnolia Press Article ISSN 1175-5326 (print edition) ZOOTAXA ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.4122.1.31 http://zoobank.org/urn:lsid:zoobank.org:pub:4ECD03DF-7333-473D-92C5-F4EDCEFC8A6A FAMILY BOMBYLIIDAE CARLOS JOSÉ EINICKER LAMAS1* & NEAL L. EVENHUIS2 1 Museu de Zoologia da Universidade de São Paulo, Avenida Nazaré, 481, Ipiranga, 04263-000, São Paulo, SP, Brazil. einicker@usp.br 2 J. Linsley Gressitt Center for Research in Entomology, Bishop Museum, 1525 Bernice Street, Honolulu, Hawaii 96817, USA. neale@bishopmuseum.org *Corresponding author: einicker@usp.br Abstract Bombyliidae is one of the largest Diptera families with more than 4,500 recognized species worldwide. Their species vary from robust to thin, and may be small to large (2-20mm) and looks like bees or wasps. They also present great variation in color. Adults can often be seen either resting and sunning themselves on trails, rocks or twigs or feeding on flowering plants as they are nectar feeders. All reared bee flies are predators or parasitoids of arthropods. The Colombian fauna of bombyliids comprises at the moment 22 species, and 12 genera, of which, six are endemic species. Nonetheless, this number may be much higher, as Colombia is a megadiverse country and there are not many specimens of this family deposited in collections all over the world. Key words: Catalogue, Bee Flies, Biodiversity, Colombia Introduction Bee flies, as the members of the family Bombyliidae are commonly called, comprise a diverse and speciose assemblage of brachycerous flies. They are one of the largest families of Diptera, with more than 4,500 species found on all continents except Antarctica and also many oceanic islands (Evenhuis & Greathead 1999). They have a remarkable range in size, color and shapes, being powerful and extremely agile in flight. Individuals can often be seen either resting in the open on trails or on rocks or twigs sunning themselves, or feeding on a variety of flowering plants. Adults are pollinators of many flower plants as a consequence of being nectar feeders (males) and obligate nectar and pollen feeders (females) (Evenhuis & Greathead 1999). Immatures are mainly parasitoids on the immatures of other holometabolous insects, but some species are known to prey on the egg pods of orthopterans (Yeates & Greathead 1997). As a result of being parasitoids, many species are important natural enemies of major pests, but others develop in nests of solitary wasps and bees and are occasionally considered to be pests when they kill the larvae of bees which are natural pollinators for crops as alfalfa in the United States (Evenhuis & Greathead 1999). Colombia is listed as one of the world’s “megadiverse” countries, hosting close to 14% of the planet’s biodiversity. Colombia possesses a rich complexity of ecological, climatic, biological and ecosystem components, but only 22 species and 12 genera of bee flies are known from Colombia (Table 1) which represents less than 2.5% of the Neotropical fauna of this family. These numbers show how the Colombian fauna is still poorly known and emphasize the necessity of extensive sampling projects to be developed in this megadiverse country. Acronyms used for the depositories AMNH—American Museum of Natural History, New York, USA. BMNH—The Natural History Museum, London, UK. 372 Accepted by S. Nihei: 26 May 2015; published: 14 Jun. 2016 CAS—California Academy of Sciences, San Francisco, California, USA. MNHN—Museum National d’Histoire Naturelle, Paris, France. MNVL—Museum d’Histoire Naturelle de Ville de Lille, Lille, France. MRSN—Museo Regionale Scienze Naturale, Torino, Italy. MZUN—Museu dell’Istituto di Zoologia dell’Università di Napoli, Naples, Italy. NMW—Naturhistorisches Museum, Vienna, Austria. SMFD—Forschungsinstitut und Museum Senckenberg, Frankfurt, a.M., Germany. USNM—National Museum of Natural History, Washington, D.C., USA. UZIU—Zoological Museum, Uppsala University, Uppsala, Sweden. ZMHB—Museum für Naturkunde, Humboldt-Universität, Berlin, Germany. ZMUC—Zoologisk Museum, University of Copenhagen, Copenhagen, Denmark. List of abbreviations aut.—automatic cat.—catalogue col.—collector des.—designation distr.—geographic distribution i.e.—example m.—meters mon.—monotypy nom. nov.—nomen novum (new name) orig. des.—by original designation pp.—pages preocc.—preoccupied refs.—references rev.—revision sp., spp.—species ssp.—subspecies subs.—subsequent subg.—subgenus syn.—synonymy HT—Holotype NT—Neotype LT—Lectotype ST—Syntype T—Type (unspecified) NA—Not applicable M (m#)—Male F (f#)—Female Catalogue of Bombyliidae of Colombia Family Bombyliidae Latreille, 1802 Subfamily Toxophorinae Schiner, 1868 Tribe Systropodini Brauer, 1880 CATALOGUE OF BOMBYLIIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 373 Genus Systropus Wiedemann, 1820 Systropus Wiedemann, 1820: 18. Type species: Systropus macilentus Wiedemann, 1820, by orig. des. (on plate). Céphène Latreille, 1825: 496 (unjustified new replacement name for Systropus Wiedemann, 1820). [Unavailable; vernacular name without nomenclatural status. Cephenus Berthold, 1827: 506 (unnecessary new replacement name for Systropus Wiedemann, 1820 [as “Systrophus”]). Type species: Systropus macilentus Wiedemann, 1820, aut. Cephenes Latreille, 1829: 505. Type species: Systropus macilentus Wiedemann, 1820, by subs. des. (Evenhuis, 1991: 25). [Unavailable; name proposed in syn. with Systropus Wiedemann and not made available before 1961.] Systropus Jensen, 1832: 335. Type species: Systropus macilentus Jensen, 1832 [preocc. = Systropus macilentus Wiedemann, 1820], by mon.. [Preocc. Wiedemann, 1820.] Xystrophus Agassiz, 1846: 393 (unnecessary emendation of Systropus Wiedemann, 1820). Type species: Systropus macilentus Wiedemann, 1820, aut. Cephenius Enderlein, 1926: 70. Type species: Systropus studyi Enderlein, 1926, by orig. des. Coptopelma Enderlein, 1926: 70. Type species: Coptopelma schineri Enderlein, 1926 [misidentification, = Systropus sanguineus Bezzi, 1921], by orig. des.. Pioperna Enderlein, 1926: 70. Type species: Cephenus femoratus Karsch, 1880, by orig. des. Symballa Enderlein, 1926: 70, 92. Type species: Systropus leptogaster Loew, 1860 [misidentification = Systropus holaspis, Speiser, 1914], by orig. des. Subgenus Systropus Wiedemann, 1820 columbianus (Karsch), 1880: 657 (Cephenus). Type locality: “Colombia”. HT ♂ [ZMHB]. Distr.: Neotropical: Brazil (Pará), Colombia, Ecuador, Panama, Peru. Refs.: Painter & Painter, 1963: 306; 1974: 73. Tribe Toxophorini Schiner, 1868 Genus Toxophora Meigen, 1803 Toxophora Meigen, 1803: 270. Type species: Toxophora maculata Meigen, 1804 [preocc. = Asilus fasciculatus Villers, 1789], by subs. mon. (Meigen, 1804: 273). Eniconevra Macquart, 1840: 110 [1841: 388]. Type species: Eniconevra fuscipennis Macquart, 1840, by mon. Heniconevra Agassiz, 1846: 138, 178 (unjustified emendation of Eniconevra Macquart), 1840. Type species: Eniconevra fuscipennis Macquart, 1840, aut. Heniconeura Bezzi, 1903: 189 (unjustified emendation of Eniconevra Macquart, 1840). Type species: Eniconevra fuscipennis Macquart, 1840, aut. Toxomyia Hull, 1973: 232 (as subgenus of Toxophora Meigen). Type species: Toxophora maxima Coquillett, 1886, by orig. des. Enicocera, error for Eniconevra. cuprea (Fabricius), 1787: 366 (Bombylius). Type locality: “Cajennae” [= French Guiana]. ST 1 (sex unknown) [ZMUC]. Distr.: Nearctic: Mexico (Morelos, Nayarit). Neotropical: Brazil (Goiás, Mato Grosso, Pará), Colombia, French Guiana, Panama, Suriname. Refs.: d’Andretta & Carrera, 1950: 354; Painter & Painter, 1974: 87; Cunha et al. 2011: 52. Subfamily Bombyliinae Latreille, 1802 Tribe Bombyliini Hull, 1973 Genus Heterostylum Macquart, 1848 Heterostylum Macquart, 1848: 195 (35). Type species: Heterostylum flavum Macquart, 1848 [= Tanyglossa hirsuta Thunberg, 1827], by orig. des. Comastes Osten Sacken, 1877: 256. Type species: Comastes robustus Osten Sacken, 1877, by mon. [Preocc. by Jan, 1863.] Unduloptomyia Evenhuis, 1978: 247 (new replacement name for Comastes Osten Sacken, 1877). Type species: Comastes robustus, Osten Sacken, 1877, aut. Heterostilum, incorrect original spelling of Heterostylum [Macquart, 1848: 231(71)]. 374 · Zootaxa 4122 (1) © 2016 Magnolia Press LAMAS & EVENHUIS hirsuta Thunberg, 1827: 67 (Tanyglossa). Type locality: “Brasilia” [= Brazil or Uruguay]. HT ♀ [UZIU]. Distr.: Neotropical: Argentina, Brazil, Colombia, Paraguay, Venezuela. Refs.: Painter & Painter, 1974: 24; Cunha et al. 2007: 18. flavum Macquart, 1848: 196(36) (Heterostylum). Type locality: “Brésil”. HT ♀ [BMNH]. Distr.: Neotropical: Argentina, Brazil, Colombia, Paraguay, Venezuela. Refs.: Painter & Painter, 1974: 24; Cunha et al. 2007: 18. xanthobasis Curran, 1929: 6 (Heterostylum). Type locality: Colombia. HT ♂ [AMNH]. Distr.: Neotropical: Argentina, Brazil, Colombia, Paraguay, Venezuela. Refs.: Painter & Painter, 1974: 24; Cunha et al. 2007: 18. Unplaced species of Bombyliinae Latreille, 1802 hyalipennis Macquart, 1846: 245(117) (Bombylius). Type locality: “Colombie” [= Colombia, Ecuador, or Venezuela]. T ♂ (lost?) [MRSN] (originally in Brême Collection)]. Distr.: Neotropical: Colombia or Ecuador or Venezuela. Refs.: Evenhuis & Greathead, 1999: 204. Subfamily Ecliminae Hall, 1969 Genus Lepidophora Westwood, 1835 Lepidophora Westwood, 1835: 447. Type species: Ploas aegeriiformis Gray, 1832 [= Toxophora lepidocera Wiedemann, 1828], by mon. secutor Walker, 1857: 145. Type locality: “Valley of the Amazon” [= Brazil (Amazonas or Pará)] HT ♂ [BMNH]. Distr.: Neotropical: Brazil (Acre, Amazonas, Roraima), Colombia, Guyana, Peru. Refs.: Rodrigues & Lamas, 2013: 30. Subfamily Lomatiinae Schiner, 1868 Genus Macrocondyla Rondani, 1863 Macrocondyla Rondani, 1863: 54, 55 [1864: 54, 55]. Type species: Macrocondyla pictinervis Rondani, 1863, by orig. des. Ulosometa Hull, 1973: 359. Type species: Ulosometa falcata Hull, 1973 [= Exoprosopa flavinevris Macquart, 1846], by orig. des. Ulosoma, incorrect original spelling of Ulosometa (Hull, 1973: 62). Subgenus Macrocondyla Rondani, 1863 flavinevris Macquart, 1846: 237(109) (Exoprosopa). Type locality: “Colombie” [= Colombia, Ecuador, or Venezuela]. LT ♀ [des. by Painter & Painter (1974: 102) [MNHN]]. Distr.: Neotropical: Argentina, Brazil (Minas Gerais), Colombia. Refs.: Painter & Painter, 1974: 101; Lamas & Couri, 1996: 314. falcata Hull, 1973: 360 (Ulosometa). Type locality: Argentina. HT ♂ [USNM]. flavinervis, error for flavinevris. Subfamily Anthracinae Latreille, 1804 Tribe Anthracini Latreille, 1804 Genus Anthrax Scopoli, 1763 CATALOGUE OF BOMBYLIIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 375 Anthrax Scopoli, 1763: 358. Type species: Musca morio Linnaeus, 1758 [misidentification, = Musca anthrax Schrank, 1781], by mon. Leucamoeba Sack, 1909: 520. Type species: Bibio aethiops Fabricius, 1781, by orig. des. Chalcamoeba Sack, 1909: 522. Type species: Anthrax virgo Egger, 1859, by orig. des. angustipennis Macquart, 1840: 64 [1841: 342]. Type locality: French Guiana. HT ♂ [MNHN]. Distr.: Neotropical: Bolivia, Brazil (Acre, Amazonas, Maranhão, Pará, Rio De Janeiro, Roraima), Colombia, French Guiana, Guyana, Panama, Peru, Trinidad, Venezuela. Refs.: Marston, 1970: 65. binotata Macquart, 1846: 241(113). Type locality: “Nouvelle-Grenade” [= Colombia, Ecuador, or Venezuela]. ST 2 ♀ [BMNH]. [Preocc. by Wiedemann, 1820.]. Distr.: Neotropical: Bolivia, Brazil (Acre, Amazonas, Maranhão, Pará, Rio De Janeiro, Roraima), Colombia, French Guiana, Guyana, Panama, Peru, Trinidad, Venezuela. Refs.: Marston, 1970: 65. cathetodaithmos Marston, 1970: 49. Type locality: Mexico, Colima. HT ♀ [USNM]. Distr.: Nearctic: Mexico (Colima). Neotropical: Brazil (Amazonas, Pará), Colombia, Costa Rica, Panama, Peru. Refs.: Marston, 1970: 49. gideon Fabricius, 1805: 124. Type locality: “America meridionali”. ST 2 ♀ [ZMUC]; ST 1 ♀ [SMFD]. Distr.: Neotropical: Brazil (Paraná, Rio de Janeiro, Rio Grande do Sul, Santa Catarina), Colombia, Costa Rica, French Guiana, Guyana, Jamaica, Mexico (Veracruz-Llave), Paraguay, Peru, Puerto Rico, Suriname, Venezuela. Refs.: Marston, 1970: 85. acroleuca Wiedemann, 1828: 312. Type locality: “Südamerika” [= Brazil (from type specimen label data)]. ST 5 ♂, 1♀ [NMW]; ST 1 ♀ in [ZMHB]. Distr.: Neotropical: Brazil (Paraná, Rio de Janeiro, Rio Grande do Sul, Santa Catarina), Colombia, Costa Rica, French Guiana, Guyana, Jamaica, Mexico (Veracruz-Llave), Paraguay, Peru, Puerto Rico, Suriname, Venezuela. Refs.: Marston, 1970: 85. propinqua (Schiner), 1868: 123 (Argyromoeba). Type locality: “Südamerika” [= Venezuela (see Marston, 1970: 88)]. HT ♀ [NMW]. Distr.: Neotropical: Brazil (Paraná, Rio de Janeiro, Rio Grande do Sul, Santa Catarina), Colombia, Costa Rica, French Guiana, Guyana, Jamaica, Mexico (Veracruz-Llave), Paraguay, Peru, Puerto Rico, Suriname, Venezuela. Refs.: Marston, 1970: 88. luctuosa Macquart, 1840: 70 [1841: 348]. Type locality: “Patrie inconnue” [= French Guiana (see Marston, 1970: 49)]. HT ♀ [MNHN]. Distr.: Neotropical: Brazil (Acre), Colombia, Costa Rica, French Guiana, Mexico (Chiapas, Oaxaca), Panama, Venezuela. Refs.: Marston, 1970: 47. midas Fabricius, 1805: 124. Type locality: “America meridionali”. HT 1 ♀ [ZMUC]. Distr.: Neotropical: Brazil (Acre, Amazonas, Goiás, Mato Grosso, Minas Gerais, Rio Grande do Norte, São Paulo), Colombia, Costa Rica, Guyana, Mexico (Chiapas, Yucatán), Venezuela. Refs.: Marston, 1970: 53. guianica Curran, 1934: 362. Type locality: “British Guiana” [= Guyana]. HT ♂ [AMNH]. Distr.: Neotropical: Brazil (Acre, Amazonas, Goiás, Mato Grosso, Minas Gerais, Rio Grande do Norte, São Paulo), Colombia, Costa Rica, Guyana, Mexico (Chiapas, Yucatán), Venezuela. Refs.: Marston, 1970: 53. mexicanus Cole, 1957: 202. Type locality: Mexico, Chiapas. HT ♂ [CAS]. Distr.: Neotropical: Brazil (Acre, Amazonas, Goiás, Mato Grosso, Minas Gerais, Rio Grande do Norte, São Paulo), Colombia, Costa Rica, Guyana, Mexico (Chiapas, Yucatán), Venezuela. Refs.: Marston, 1970: 53. Tribe Exoprosopini Becker, 1912 Genus Exoprosopa Macquart, 1840 Exoprosopa Macquart, 1840: 35 [1841: 313]. Type species: Anthrax pandora Fabricius, 1805, by subsequent designation (Coquillett, 1910: 544) [des. conserved by Opinion 1632 (I.C.Z.N., 1991: 82).] Trinaria Mulsant, 1852a: 20 [1852b: 180]. Type species: Anthrax interrupta Mulsant, 1852 [preocc. = Anthrax rutila Pallas & Wiedemann, 1818], by subs. des.. (Coquillett, 1910: 617). Argyrospila Rondani, 1856: 162, 202. Type species: Anthrax jacchus Fabricius, 1805, by orig. des. (as “jaccus”). 376 · Zootaxa 4122 (1) © 2016 Magnolia Press LAMAS & EVENHUIS Exoptata Coquillett, 1887: 13 (as subgenus of Exoprosopa Macquart). Type species: Exoprosopa divisa, Coquillett, 1887, by mon. Argyrospyla, incorrect original spelling of Argyrospila (Rondani, 1856: 162). antica (Walker), 1852: 183 (Anthrax). Type locality: Colombia. HT ♀ [BMNH]. Distr.: Neotropical: Colombia. Refs.: Painter & Painter, 1974: 123. argentifasciata Macquart, 1846: 237(109). Type locality: “Colombie” [= Colombia, Ecuador, or Venezuela]. ST 1 ♂, 6 ♀ [BMNH]. Distr.: Nearctic: Mexico (Guerrero, Jalisco, Michoácan de Ocampo, Morelos, Puebla, San Luis Potosí, Veracruz-Llave). Neotropical: Colombia, Costa Rica, Mexico (Chiapas, Oaxaca, Yucatán), Panama. Refs.: Painter & Painter, 1974: 124. limbipennis Macquart, 1846: 238(110). Type locality: Mexico, Yucatán. ST 2 ♀# [MNHN]; ST 2 ♂ [BMNH]. Distr.: Nearctic: Mexico (Guerrero, Jalisco, Michoácan de Ocampo, Morelos, Puebla, San Luis Potosí, VeracruzLlave). Neotropical: Colombia, Costa Rica, Mexico (Chiapas, Oaxaca, Yucatán), Panama. Refs.: Painter & Painter, 1974: 124. panamensis Curran, 1930: 9. Type locality: Panama. HT ♂ [AMNH]. Distr.: Neotropical: Brazil (Mato Grosso), Colombia, Panama. Genus Hyperalonia Rondani Hyperalonia Rondani, 1863: 58 [1864: 58]. Type species: Anthrax erythrocephala Fabricius, 1805, by subs. des. (Dallas, 1866: 649). surinamensis Rondani, 1863: 58 [1864: 58]. Type locality: Suriname. HT ♂ [MZUN]. Distr.: Neotropical: Brazil (Amazonas, Bahia, Mato Grosso, Pará, São Paulo), Colombia, Ecuador, Guyana, Peru, Suriname. Refs.: Couri & Lamas, 1994: 121. Genus Ligyra Newman Ligyra Newman, 1841: 220. Type species: Anthrax bombyliformis Macleay, 1826, by mon. Velocia Coquillett, 1886: 158. Type species: Anthrax cerberus Fabricius, 1794, by orig. des. [Preocc. by Robineau-Desvoidy, 1863.] Paranthrax Paramonov, 1931: 57(57). Type species: Paranthrax africanus Paramonov, 1931 [= Hyperalonia niveifrons Bezzi, 1914], by mon. [Preocc. by Paranthrax Bigot, 1876.] Paranthracina Paramonov, 1933: 56 [1934: 27; 1936: 31] (replacement name for Paranthrax Paramonov, 1931). Type species: Paranthrax africanus Paramonov, 1931 [= Hyperalonia niveifrons Bezzi, 1914], aut. Lygira, error for Ligyra. Vulpia, error for Velocia. latreillii (Wiedemann), 1830: 633 (Anthrax). Type locality: “Oaxara im Mexikanischen” [= Mexico (Oaxaca)]. HT ♂ [ZMHB]. Distr.:Nearctic: Mexico (México). Neotropical: Argentina, Bolivia, Brazil, Colombia, Costa Rica, Guatemala, Honduras, Jamaica, Mexico (Chiapas, Oaxaca, Tabasco, Yucatán), Panama, Paraguay, Peru. Tribe Villini Hull, 1973 Genus Chrysanthrax Osten Sacken Chrysanthrax Osten Sacken, 1886b: 113 (as subgenus of Anthrax Scopoli), 121 (as “group” of Anthrax Scopoli). Type species: Anthrax fulvohirta Wiedemann, 1821 [= Anthrax cypris Meigen, 1820], by mon. [See Evenhuis (1991: 27) for clarification of type fixation of this genus.] CATALOGUE OF BOMBYLIIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 377 dichotoma (Schiner), 1868: 119 (Exoprosopa). Type locality: “Südamerika” [= Venezuela]. LT ♂ [des. by Painter & Painter (1974: 192) [NMW]]. Distr.: Neotropical: Colombia, Venezuela. Refs.: Painter & Painter, 1974: 192. bipartita (Bigot), 1892: 346 (Exoprosopa). Type locality: “Colombie”. HT ♀ [BMNH]. Distr.: Neotropical: Colombia, Venezuela. Refs.: Painter & Painter, 1974: 192. procedens (Walker), 1852: 185 (Anthrax). Type locality: “Columbia” [= Colombia]. HT ♀ [BMNH]. Distr.: Neotropical: Colombia. Refs.: Painter & Painter, 1974: 264. Genus Hemipenthes Loew Hemipenthes Loew, 1869: 28 [1872: 142]. Type species: Musca morio Linnaeus, 1758, by subs. des. (Coquillett, 1894: 90). Isopenthes Osten Sacken, 1886a: 80, 96. Type species: Isopenthes jaennickeana Osten Sacken, 1886, by subs. des. (Coquillett, 1910: 556). ignea (Macquart), 1846: 241(113) (Anthrax). Type locality: “Colombie” [= Colombia, Ecuador, or Venezuela]. LT ♀ [des. by Painter & Painter (1974: 233) [BMNH]. Distr.: Neotropical: Colombia, Guyana, Panama. Refs.: Painter & Painter, 1974: 233. Genus Villa Lioy Villa Lioy, 1864: 732. Type species: Anthrax concinna Meigen, 1820 [= Anthrax abbadon Fabricius, 1794], by subs. des. (Coquillett, 1910: 619). Hyalanthrax Osten Sacken, 1886b: 112 [1887: 134] (as subgenus of Anthrax Scopoli). Type species: Anthrax faustina Osten Sacken, 1887, by subs. des. (Coquillett, 1910: 553). Aspiloptera Künckel d’Herculais, 1905: 145. Type species: Anthrax flava Meigen, 1820 [= Musca hottentotta Linnaeus, 1758], by mon. Protepacmus Cockerell, 1916: 94. Type species: Protepacmus setosus Cockerell, 1916, by mon. colombiana Evenhuis & Greathead, 1999: 492 (new replacement name for Anthrax vicina Macquart, 1846). Distr.: Neotropical: Colombia. vicina Macquart, 1846: 240(112) (Anthrax). Type locality: “Colombie” [= Colombia, Ecuador, or Venezuela]. T ♀ (lost?) [MNHN]. [Preocc. by Macquart, 1840.] n. syn. Distr.: Neotropical: Colombia. Refs.: Painter & Painter, 1974: 285. diminutiva (Macquart), 1846: 241(113) (Anthrax). Type locality: “Colombie” [= Colombia, Ecuador, or Venezuela]. HT ♀ [MNHN]. Distr.:Neotropical: Colombia. Refs.: Painter & Painter, 1974: 198. UNPLACED SPECIES OF VILLINI HULL gradata Macquart, 1847a: 53 [1847b: 69] (Anthrax). Type locality: “Colombie” [= Colombia, Ecuador, or Venezuela]. HT ?♂ [MNVL]. Distr.:Neotropical: “Colombie” [= Colombia, Ecuador or Venezuela]. Refs.: Painter & Painter, 1974: 227. latifimbria Walker, 1852: 175 (Anthrax). Type locality: “Columbia”[= Colombia]. HT (sex unknown) [(lost) [BMNH ] (J.E. Chainey, pers. comm.)]. Distr.: Neotropical: Colombia. 378 · Zootaxa 4122 (1) © 2016 Magnolia Press LAMAS & EVENHUIS TABLE 1. Bombyliidae species recorded from Colombia. Subfamily Tribe Genus Species Author Toxophorinae Systropodini Systropus columbianus (Karsch, 1880) Toxophorini Toxophora cuprea (Fabricius, 1787) Bombyliini Heterostylum hirsutum (Thunberg, 1827) Unplaced species of Bombyliinae “Bombylius” hyalipennis Macquart, 1846 Lepidophora secutor Walker, 1857 Bombyliinae Ecliminae Lomatiinae Lomatiini Macrocondyla (Macrocondyla) flavinevris (Macquart, 1846) Anthracinae Anthracini Anthrax angustipennis Macquart, 1840 Anthrax cathetodaithmos Marston, 1970 Anthrax gideon Fabricius, 1805 Anthrax luctuosus Macquart, 1840 Anthrax midas Fabricius, 1805 Exoprosopa antica (Walker, 1852) Exoprosopa argentifasciata Macquart, 1846 Exoprosopa panamensis Curran, 1930 Hyperalonia surinamensis Rondani, 1863 Ligyra latreillii (Wiedemann, 1830) Chrysanthrax dichotomus (Schiner, 1868) Chrysanthrax procedens (Walker, 1852) Hemipenthes ignea (Macquart, 1846) Villa colombiana (Evenhuis & Greathead, 1999) Villa diminutiva (Macquart, 1846) “Anthrax” gradatus Macquart, 1847 “Anthrax” latifimbrius Walker, 1852 Exoprosopini Villini Unplaced species of Villini References Agassiz, J.L.R. 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Nazaré, 481, Ipiranga, 04263–000, São Paulo, SP, Brazil * Corresponding author. E-mail: juliacalhau@gmail.com Abstract Mydidae (Diptera, Asiloidea) are a relatively rare group of flies, and the knowledge on the Colombian mydids is very poor. According to available literature, only two species are registered to this country, Protomydas coerulescens (Olivier, 1811) and P. rubidapex (Wiedemann, 1830). Key words: catalogue, Colombia, Mydidae, Protomydas Introduction Mydidae (Diptera, Asiloidea) are a relatively rare group of flies, currently comprising 11 subfamilies, 66 genera and about 460 valid species (Dikow 2010). This family includes the largest flies in the world: Gauromydas heros, endemic to Brazil, can reach 70 mm in body length (Autuori 1952). Some species are mimics of Hymenoptera, especially Pompilidae (Zikán 1942; Cooper 1981; Meyer et al. 1984; Nelson 1986). Adult mydids can be identified by the following features: head usually with sunken vertex; face medially gibbous (gibbosity), with setulae forming a mystax; medial ocellus regular, but lateral ocelli often highly modified, linear and adjacent to a central callus (except in Megascelinae and Rhaphiomidinae); antenna with elongate postpedicel, often with basal stalk and apical thickened club; palpus one-segmented, frequently vestigial; hind femur sometimes thickened, with ventral tuberculate spiniform setae; hind tibia sometimes with ventral carina, often prolonged on apical spine; wing with apical branches of M peculiarly curved anteriorly, parallel to distal portion of R5 and posterior wing margin; Tergite 2 with a pair of callosities (bullae) on posterior margin (absent in Megascelinae and Rhaphiomididae); male hypopygium lacking lateral ejaculatory apodeme and gonostylus. The majority of the species occur in warmer climates (Dikow 2010), and are commonly associated to open vegetation, although some species of Mydinae are found in tropical rain forests and wet habitats (Wilcox & Papavero 1975). The adults are essentially flower feeders (Drury 1773; Zikán 1942, 1944; Rogers & Mattoni 1993; Williams 1995), but females of some species seem not feed at all as adults (Zikán 1942). Most of what it is known about larval biology of Mydidae is concerned to Mydinae and its association with nests of “sauva ants” (Formicidae, Attinae) (Zikán 1942, 1944; Autuori 1952, 1971). The mydid larvae live in the “garbage pan” [“panela de lixo”] of the ant nest, where residues of decaying plants acumulate and seem to attract several Coleoptera, especially some Dynastinae (Melolonthidae) (Eidmann 1937). Zikán (1942, 1944) believed that the mydid larvae prey upon those dynastine larvae. The knowledge on the Colombian Mydidae is very poor, and only two species are registered to this country (Papavero 2009). In the last five years, while working on Mydidae material deposited in many collections around the world, the present authors found no specimens from Colombia. For this reason, this catalog is only based on literature, and only two species are registered to the country (Table 1). 382 Accepted by S. Nihei: 26 May 2015; published: 14 Jun. 2016 Acronyms used for the depositories BZM—Berliner Zoologischer Museum, Humboldt-Universität, Berlin, Germany MZUSP—Museu de Zoologia da Universidade de São Paulo, São Paulo, Brazil OXF—Hope Department of Entomology, Oxford University, Oxford, England UUZM—Museum of Evolution, Uppsala University, Uppsala, Sweden List of abbreviations comb.—combination cat.—catalogue des.—designation distr.—geographic distribution hist. Mexican dipterol.—Contributions to a history of Mexican Dipterology Man. Neot. Dip.—Manual of Neotropical of Diptera misident.—misidentification orig. des.—by original designation pp.—pages refs.—references syn.—synonym HT—Holotype NT—Neotype ST—Syntype F (f#)—Female M (m#)—Male Catalogue of Mydidae of Colombia Family Mydidae Latreille, 1809 Subfamily Mydinae Tribe Mydini Subtribe Protomydina Genus Protomydas Wilcox, Papavero & Pimentel Protomydas Wilcox, Papavero & Pimentel, 1989: 13. Type species, Mydas coerulescens Olivier, 1811 (orig. des.). coerulescens (Olivier), 1811: 83 (Mydas). Type locality: “South America”. Neotype-locality: Brazil, Rio de Janeiro, Itatiaia. HT lost. NT M (MZUSP) [des. by Wilcox, Papavero & Pimentel, 1989: 14]. Distr.: Colombia, Brazil (Amazonas, Mato Grosso, Minas Gerais, Pará, Rio de Janeiro, São Paulo). Refs.: Wiedemann, 1831: 33 (Mydas); Westwood, 1841: 50 (Midas); Gertaecker, 1868: 95 (Mydas; syn. of Mydas giganteus Thunberg, 1818); Becher, 1882: 146, (Mydas); Hunter, 1901: 154 (cat.; Mydas); Kertész, 1909: 37 (cat.; Mydas); d´Andretta, 1951: 10, (Mydas); Papavero & Wilcox, 1968: 7 (cat.; Mydas); Wilcox, Papavero & Pimentel, 1989: 14 (comb.; syn. of Mydas argyrostomus Gerstaecker, 1868); Artigas & Papavero, 1990: 41-42 (spermathecae); Papavero & Artigas, 1990: 125 (cat.); Papavero et al., 2002: 3 (cat.); Papavero, 2009: 12 (cat.); Papavero & Artigas, 2009: 37, (Man. Neot. Dip.). giganteus Thunberg, 1818: 246, (Mydas, as gigantea). Type locality: “Brazil”. ST (UUZM). Distr.: Brazil CATALOGUE OF MYDIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 383 (Minas Gerais). Refs.: Wiedemann, 1821: 116 (Midas, as gigantea), 1824: 60 (Mydas, as gigantea), 1828: 239 (Mydas); Latreille, 1829: 172, (Mydas); Wiedemann, 1831: 36, 38, (Mydas); Macquart, 1834: 274 (Mydas); Westwood, 1841: 50 (Midas); Macquart, 1838: 126 (Mydas); Walker, 1848: 228 (Mydas); Rondani, 1850: 184 (Mydas); Walker, 1854: 363 (Mydas). argyrostomus Gerstaecker, 1868: 94 (Mydas). Type locality: “Colombia”. HT M (BZM). Distr.: Colombia, Brazil (Pará: Óbidos). Refs.: Hansen, 1883: 13–21 (Mydas); Hunter, 1901: 153 (cat.; Mydas); Kertész, 1909: 35 (cat.; Mydas); D´Andretta, 1951: 28, 38, 117, 140, 153. rubidapex (Wiedemann), 1830: 626 (Midas). Type locality: “Mexico”. HT M (BZM). Distr.: Brazil (Amazonas, Pará), Colombia (Boyacá (Muzo)), Mexico, Venezuela. Refs.: Wiedemann, 1831: 36, 40, (Mydas); Westwood, 1841: 51 (Midas); Walker, 1854: 361 (Mydas); Bellardi, 1861: 5 (Mydas); Gerstaecker, 1868: 96 (Mydas); Schiner, 1868: 153 (Mydas); Osten Sacken, 1874: 175 (Mydas); Brauer, 1885: 387, (Mydas); Osten Sacken, 1886: 70 (Mydas); Williston, 1898: 55 (in key and text; Mydas); Hunter, 1901: 154 (cat.; Mydas); Williston et al., 1900: 267 (Mydas); Aldrich, 1905: 252 (cat.; Mydas); Kertész, 1909: 40 (cat.; Mydas); d´Andretta, 1951: 21; Papavero & Wilcox, 1968: 7 (cat.); Wilcox, Papavero & Pimentel, 1989: 16 (comb.; syn. of Mydas dives Westwood, 1841; syn. of Mydas heros Perty of D´Andretta, 1951: 20 (part), misident.); Papavero & Artigas, 1990: 125 (cat.); Papavero, 1996: 632 (cat.); Papavero & Ibáñez Bernal, 2001: 75 (hist. Mexican dipterol.); Papavero et al., 2002: 3 (cat.); Papavero, 2009: 12 (cat.); Papavero & Artigas, 2009: 11, 15 (Man. Neot. Dip.). dives Westwood, 1841: 50, pl. 13,(Midas). Type locality: “Unknown”. HT F (OXF). Distr.: Brazil (Pará), Colombia (Boyacá (Muzo)). Refs.: Walker, 1848: 228 (Mydas), 1854: 363 (Mydas); Gerstaecker, 1868: 94 (Mydas); Osten Sacken, 1886: 70 (Mydas); Williston, 1898: 55 (in key and text; Mydas); Hunter, 1901: 154 (cat.; Mydas); Aldrich, 1905: 154 (cat.; Mydas); Kertész, 1909: 37 (cat.; Mydas); Curran, 1934: 326 (Mydas); d´Andretta, 1951: 37, 60, 120, 149 (Mydas); Papavero & Ibáñez Bernal, 2001: 87 (hist. Mexican dipterol.). heros Perty of D´Andretta, 1951: 20 (Mydas; part, specimens from Brazil, Amazonas: Borba-Guajará, and Colombia, Muzo), misident. TABLE 1. Mydidae species recorded from Colombia. Subfamily Tribe Subtribe Genus Mydinae Mydini Protomydina Protomydas species author coerulescens (Olivier, 1811) rubidapex (Wiedemann, 1830) References Agassiz, L. (1846) Nomenclator zoologici index universalis. Jent & Gassmann, Soloduri, viii + 393 pp. Aldrich, J.M. (1905) A catalogue of North American Diptera. 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Boletim do Ministério da Agricultura, 33, 43–55. 386 · Zootaxa 4122 (1) © 2016 Magnolia Press CALHAU & LAMAS Zootaxa 4122 (1): 387–392 http://www.mapress.com/j/zt/ Copyright © 2016 Magnolia Press Article ISSN 1175-5326 (print edition) ZOOTAXA ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.4122.1.33 http://zoobank.org/urn:lsid:zoobank.org:pub:6358CE14-E49C-4111-AEEE-E7F785E69FE7 FAMILY THEREVIDAE PAULA FERNANDA MOTTA RODRIGUES1,2,3* & JULIA CALHAU1 1 Museu de Zoologia da Universidade de São Paulo. Avenida Nazaré, 481 Ipiranga, 04263-000. São Paulo-SP, Brazil. Programa de Pós-Graduação em Ciências Biológicas (Zoologia) do Instituto de Biociências, Universidade de São Paulo. 3 Conselho Nacional de Desenvolvimento Científico e Tecnológico (CNPq), fellow. * Corresponding author: paulafm_rodrigues@yahoo.com.br 2 Abstract This catalogue presents the Colombian fauna of stiletto flies (Diptera: Therevidae). According to available literature, only four species, distributed in three genera, are registered to this country, Ataenogera brevicornis (Bromley, 1934), Brachylinga punctifrons (Kröber, 1914), B. xanthoperna Irwin and Webb, 1991 and Chromolepida pruinosa (Coquillett, 1904). Key words: catalogue, Colombia, distribution, Therevidae Introduction Stiletto flies (Diptera: Asiloidea: Therevidae) are a moderately sized group of lower brachyceran flies that along with Scenopinidae, Apsilocephalidae and Evocoidae, comprise the therevoid clade (Yeates et al. 2003, 2006, Woodley et al. 2009). Therevidae is divided into four subfamilies and two tribes: Agapophytinae, Phycinae, Therevinae (Cyclotelini and Therevini (non-cycloteline Therevinae) (Gaimari & Irwin 2000)) and Xestomyzinae, and occurs on all continents except Antarctica (Webb et al. 2013). Worldwide, there are 1.123 described species in 119 genera (Thompson 2008), and this number represents only 56.15% of the estimated fauna of Therevidae (Gaimari & Webb 2009). The Neotropical fauna comprises 147 species (Amorim et al. 2002). In the New World, adult stiletto flies can be identified by the following features: cuticle and prominent macrosetae pale yellow to black, entirely or partially covered with pubescence; males generally holoptic (except in some genera of Phycinae), females dichoptic; wing with R4 sinuous, divergent from R5, r4 enclosing apex of wing, bm truncate apically, d elongate from which arise veins M1, M2, and M3, m-cu present with base of m3 truncate, r-m attached to basal half of d, cup closed near wing margin; abdomen with eight pregenital segments, often with silver or bronze pubescence dorsally in males; tergite 8 in males usually broad and rectangular, with posterior margin broadly emarginate; epandrium rectangular to quadrate, wider than long in most species; cerci one-segmented, usually free basally; hypandrium often reduced, strap-like, occasionally absent; gonocoxites bulbous, separated medially in most species (except in Cyclotelini, all of which have gonocoxites medially fused), gonostyli apically; aedeagus usually narrow, tapered posteriorly with an internal ejaculatory apodeme; sternite 8 of the female terminalia conspicuous; acanthophorite macrosetae strongly reduced in Phycinae and Xestomyzinae, present and distinct posterodorsally in Therevinae and Agapophytinae; female Phycinae has three spermathecae but lack a spermathecal sac, while all other therevids have a spermathecal sac in addition to spermathecae, with two spermathecae in Therevinae and Xestomyzinae, and three in Agapophytinae (although the Australian genus Bonjeania Irwin and Lyneborg has only one) (Webb et al. 2013). Therevids are relatively rare and active during daytime, when they feed mostly on water, plant and insect secretions or even on insect excretions (Irwin & Lyneborg 1981b). They usually occur in dry areas (Majer 1997), but they can also be found in mountains and tropical humid regions (Gaimari & Webb 2009). Larvae feed on other insects, mainly beetles, butterflies, moths and other flies that live in leaf litter, tree holes or sandy terrains (Irwin & Accepted by S. Nihei: 10 Apr. 2015; published: 14 Jun. 2016 387 Webb 1991). A few species may even be cannibals and others are phytophagous (Majer 1997). Therevids are infrequently collected and sampled, because the predator larvae live inside dry and fragile substrates, and for this reason adults are found in the same areas. Thevids are better collected and sampled using malaise traps for their behavior could be easily predicted: usually they are flying in arid environments searching for water or displacing themselves along trails (Gaimari & Webb 2009). Males of some species tend to set on beams of light in trails, usually waiting for females to mate (Irwin & Lyneborg, 1981b). Some species may even possess a very specific substrate for repose, from sand spots, rocks, herbs, leaves to trunks (Gaimari & Webb 2009). The therivid fauna of the Neotropical region is poorly studied and undercollected, although many new taxa are recognized. This catalogue of Colombian Therevidae and the classification adopted is based on the catalogue by Webb et al. (2013), the first catalogue of New World Therevidae. According to available literature, only four species, distributed in three genera, are registered to Colombia (Table 1). Acronyms used for the depositories AMNH—American Museum of Natural History, Division of Invertebrate Zoology, New York, New York, USA. INBio—Instituto Nacional de Biodiversidad, Sección Artrópodos, Santo Domingo de Heredia, Costa Rica. INPA—Instituto Nacional de Pesquisas da Amazônia, Coleção de Invertebrados, Amazonas, Manaus, Brazil. USNM—National Museum of Natural History, Department of Entomology, Washington, D.C., USA. List of abbreviations abd.—abdomen cat.—catalogue comb.—combination desc.—description desig.—designation distr.—geographic distribution F (f#)—Female figs.—figures genit.—genitalia HT—holotype M (m#)—Male max. palp.—maxillary palpus mon.—monotypy nom.—nomenclature NT—neotype orig.—original phylog.—phylogeny refs.—references sp.—species syn.—synonym trans.—translation Catalogue of Therevidae of Colombia Family Therevidae Newman, 1834 Subfamily Phycinae Lyneborg, 1976 388 · Zootaxa 4122 (1) © 2016 Magnolia Press RODRIGUES & CALHAU Genus Ataenogera Kröber Ataenogera Kröber, 1914: 31. Type species, Ataenogera abdominalis Kröber, 1914 (mon.). Leptocera Kröber, 1928b: 117. Type species, Leptocera gracilis Kröber, 1928b (mon.). Preoccupied by Olivier (1813: 489). Ziehenia Kröber, 1929a: 434. Replacement name for Leptocera Kröber (1928b: 117) nec Olivier (1813: 489). Epileptocera Richards, 1929: 171. Unjustified replacement name for Leptocera Kröber (1928b: 117) nec Olivier (1813: 489). Ziehenimyia. Misspelling of Ziehenia. Kröber (1929b: 172 desc.), Hauser and Webb (2007: 43 nom.). brevicornis (Bromley), 1934: 361 (Henicomyia). Type locality: Guyana, Bartica District [=Cuyuni-Mazaruni], Kartabo. HT F (AMNH). Distr.: Nearctic. Mexico (Jalisco, Michoacán, Morelos, Puebla). Neotropical. Brazil (Amazonas, Mato Grosso do Sul, Pará, Rondônia, Santa Catarina), Colombia (Magdalena), Costa Rica (Alajuela, Guanacaste, Puntarenas), El Salvador (La Libertad), Guatemala (Zacapa), Guyana (Cuyuni-Mazaruni), Nicaragua (Rivas), Panama (Darién, Panamá (Canal Zone)), Peru (San Martin), Trinidad, Uruguay, Venezuela (Aragua, Barinas, Guárico). Refs.: Webb & Irwin, 1988: 39 (nom., syn. desig. of A. abdominalis Kröber; Henicomyia); Hauser & Webb, 2007: 42 (revised status), 44 (key), 52 (nom., desc., comb. change), 54 (distr.), Figs. 20 male frons, 21 female frons, 22–27 male genit., 28 female abd., 29 female genit., 57 (distr. map); Gaimari & Webb, 2009: 644 (distr.); Webb et al., 2013: 16 (cat.). Subfamily Therevinae Newman, 1834 Tribe Cyclotelini Gaimari and Irwin, 2000 Genus Brachylinga Irwin and Lyneborg Brachylinga Irwin and Lyneborg 1981a: 232. Type species, Psilocephala baccata Coquillett, 1893 (orig. des.). punctifrons (Kröber), 1914: 58 (Psilocephala). Type locality: Guatemala, Departamento Esquintla, Monterrico. NT M (MEI 108667) (INBC). Distr.: Nearctic. Mexico (Jalisco, Nayarit, Puebla). Neotropical. Colombia (Bolivar), Costa Rica (Guanacaste, Puntarenas), El Salvador (Ahuachapán, La Libertad), Guatemala (Esquintla, Guatemala, Quetzaltenango), Mexico (Chiapas, Colima, Guerrero, Oaxaca, Sinaloa), Nicaragua (Masaya), Panama (Chiriqui, Panamá (Canal Zone)). Refs.: Kröber, 1914: 38 (male key; Psilocephala), 40 (female key; Psilocephala); Kröber, 1928a: 7 (distr.; Psilocephala), 8 (male key; Psilocephala), 11 (female key; Psilocephala), 16 (desc.; Psilocephala), Figs. 11 female frons, ant., 11a male ant.; (Psilocephala); Metz et al., 2003: 248 (NT desig., comb. change), 250 (trans. orig. desc.); Webb & Metz, 2006: 14 (male key), 17 (female key), 123 (nom.), 124 (desc.), Figs. 20, 43, 66, 89, 112, 135, 158, 181 male genit., 202 female genit., 215 distr. map; Webb et al., 2013: 27 (cat.). xanthoperna Irwin and Webb, 1991: 87 (checklist), 91 (distr.), Figs. 11 female frons, 12 female ant., 13 female max. palp., 14 female wing, 15–16 female genit.. Type locality: Brazil, Roraima, Rio Uraricoera, Ilha de Maracá. HT F (MEI 006913) (INPA). Distr.: Neotropical. Brazil (Roraima), Colombia (Magdalena), Venezuela (Aragua, Guárico, Monagas, Nueva Esparta, Zulia). Refs.: Webb & Metz, 2006: 15 (male key), 17 (female key), 143 (desc.), Figs. 24, 47, 70, 92, 116, 139, 162, 185 male genit., 206 female genit., 213 distr. map; Webb et al., 2013: 27 (cat.). Tribe Therevini (non-Cyclotelini) Newman, 1834 Genus Chromolepida Cole Chromolepida Cole, 1923: 23. Type species, Psilocephala pruinosa Coquillett, 1904 (orig. des.). Chrmomolepida, misspelling of Chromolepida. pruinosa (Coquillett), 1904: 91 (Psilocephala). Type locality: Nicaragua, Granada. HT M (MEI 009218) [USNM CATALOGUE OF THEREVIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 389 (Type No. 7795)]. Distr.: Nearctic. Mexico (Puebla). Neotropical. Colombia (Magdalena), Costa Rica (Alajuela, Guanacaste, San José), Guatemala, Guyana, Honduras, Nicaragua (Granada), Venezuela (Aragua, Guárico). Refs.: Kröber, 1911: 499 (male key; Psilocephala), 522 (desc.; Psilocephala); Kröber, 1913: 33 (sp. list; Psilocephala), 37 (male key; Psilocephala); Kröber, 1928a: 7 (distr.; Psilocephala), 9 (male key; Psilocephala); Kertész, 1909: 164 (cat.; Psilocephala); Cole, 1923: 23 (key), 24 (desc., comb. change); Irwin & Lyneborg, 1981a: 260 (sp. list); Webb & Irwin, 1995: 197 (phylog.), 204 (key), 220 (nom., desc.), 221 (distr.), Figs. 1 phylog., 42 ant., 43 max. palp., 44 wing, 45–51 male genit., 52 female head, 53–55 female genit., 58 distr. map; Webb et al., 2013: 34 (cat.). TABLE 1. Therevidae species recorded from Colombia. 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(Eds.), Manual of Palaeractic Diptera. Vol. 2. Science Herald, Budapest, Hungary, pp. 519–529. Marschall, A.F. de (1873) Nomenclator Zoologicus continens nomina systematica generum animalium tam viventium quam fossilium, secundum ordinem alphabeticum disposita. C. Ueberreuter, Vindobonae [= Wien], 482 pp. Metz, M.A., Webb, D.W. & Irwin, M.E. (2003) A review of the genus Psilocephala Zetterstedt (Diptera: Therevidae), with the description of four new genera. Studia Dipterologica, 10 (1), 227–266. Newman, E. (1834) Attempted division of British insects into natural orders. Entomological Magazine, 2, 379–431. Olivier, A.G. (1813) Premièr mèmoire sur quelques insectes qui attaquent les céréales. Mémoires de la Société d'Agriculture, Departement de la Seine, 16, 477–495. Ôuchi, Y. (1943) Diptera Sinica. Thervidae 1. On three new stilleto flies from East China. Shanghai Sizenkagu Kenkyūsyo Ihō, 13, 477–482. Perty, J.A.M. (1841) Allgemeine Naturgeschichte, als philosophische und Humanitäts-Wissenschaft für Naturforscher, Philosophen und das höher gebildete Publikum. Vol. 3. C. Fischer, Bern, xii + 467–1119 pp. Richards, O.W. (1929) Systematic notes on the Borboridae (Diptera), with descriptions of new species of Leptocera (Limosina). Entomologist's Monthly Magazine, 65 (3), 171–176. Schoch, G. (1890) Fauna insectorum Helvetiae, Diptera. Bolli & Böcherer, Schaffhausen, 27 pp. Thompson, F.C. (2008) The Diptera site. The biosystematic database of world Diptera. Nomenclator status statistics. Version 10.5. Available from: http://www.sel.barc.usda.gov/diptera/names/status/bdwdstat.htm/ (accessed 5 August 2013) Walker, F. (1874) Notes on Diptera and lists of species. First series. E.W. Janson, London, [1] + 37 pp. Webb, D.W. & Irwin, M.E. (1988) The genera Ataenogera and Phycus in the New World. Proceedings of the Entomological Society of Washington, 91 (1), 35–50. [1989] Webb, D.W. & Irwin, M.E. (1995) The New World genus Chromolepida Cole (Diptera: Therevidae: Therevinae). Proceedings of the Entomological Society of Washington, 97, 197–224. Webb, D.W. & Metz, M.A. (2006) A revision of the New World genera Brachylinga Irwin and Lyneborg and Lysilinga Irwin and Lyneborg (Diptera: Therevidae: Thereviilae) with the descripction of a new genus, Elcaribe Webb. Zootaxa, 1288, 1–241. Webb, D.W., Gaimari, S.D., Hauser, M., Holston, K.C., Metz, M.A., Irwin, M.E., Kampmeier, G.E. & Algmin, K. (2013) An annotated catalogue of the New World Therevidae (Insecta: Diptera: Asiloidea). Zootaxa, 3600 (1), 1–105. http://dx.doi.org/10.11646/zootaxa.3600.1.1 Woodley, N.E., Borkent, A. & Wheeler, T.A. (2009) Phylogeny of the Diptera. In: Brown, B.V., Borkent, A., Cumming, J.M., Wood, D.M., Woodley, N.E. & Zumbado, M. (Eds.), Manual of Central American Diptera. Vol. 1. National Research Council Research Press, Ottawa, pp. 79–94. Woodworth, C.W. (1913) Guide to California Insects. The Law Press, Berkeley, 360 pp. http://dx.doi.org/10.5962/bhl.title.49919 Yeates, D.K., Irwin, M.E. & Wiegmann, B.M. (2003) Ocoidae, a new family of asiloid flies (Diptera: Brachycera: Asiloidea), based on Ocoa chilensis gen. and sp.n. from Chile, South America. Systematic Entomology, 28, 417–431. CATALOGUE OF THEREVIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 391 http://dx.doi.org/10.1046/j.1365-3113.2003.00224.x Yeates, D.K., Irwin, M.E. & Wiegmann, B.M. (2006) Evocoidae (Diptera : Asiloidea), a new family name for Ocoidae, based on Evocoa, a new replacement name for the Chilean genus Ocoa Yeates, Irwin, and Wiegmann. Systematic Entomology, 31 (2), 373. http://dx.doi.org/10.1111/j.1365-3113.2006.00332.x 392 · Zootaxa 4122 (1) © 2016 Magnolia Press RODRIGUES & CALHAU Zootaxa 4122 (1): 393–396 http://www.mapress.com/j/zt/ Copyright © 2016 Magnolia Press Article ISSN 1175-5326 (print edition) ZOOTAXA ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.4122.1.34 http://zoobank.org/urn:lsid:zoobank.org:pub:9DDDB807-5D14-417B-BD83-CC6577C81B30 FAMILY EMPIDIDAE JOSÉ ALBERTINO RAFAEL1* & JOSENIR TEIXEIRA CÂMARA2 1 Instituto Nacional de Pesquisas da Amazônia, INPA, Coordenação de Biodiversidade, Caixa Postal 2223, 69080–971, Manaus, Amazonas, Brazil. E-mail: jarafael@inpa.gov.br 2 Instituto Nacional de Pesquisas da Amazônia, INPA, Programa de Pós–Graduação em Entomologia, Caixa Postal 2223, 69080–971, Manaus, Amazonas, Brazil. E-mail: josenircamara@gmail.com * Corresponding author: jarafael@inpa.gov.br Abstract Empididae is one of the biggest families of Diptera with around 3,000 known species. The family is poorly known in Colombia, with only six species reported and this work provides information on their distribution. No endemic genera or species have been recorded to date from Colombia. Key words: catalogue, Colombia, dance-flies, distribution Introduction The Empididae are small flies, from 1–12 mm in length, usually dark in color, sometimes yellow and rarely with metallic green or blue reflections. Adults are mostly predators, but some species feed on pollen and nectar (Steyskal & Knutson 1981). They are found resting in vegetation or flying in search of possible prey. Males of some species of Empidinae offer a gift to the female before copulation. Flower-feeding of some species of Empis is indicative of their importance as pollinators for some species of plants. Adults are usually predators of smaller insects, including Diptera (also other empidids), Hemiptera, Thysanoptera, Lepidoptera, Trichoptera, Hymenoptera, Neuroptera, Ephemeroptera, Plecoptera, Coleoptera, Collembola and Acari (Tuomikoski 1952, Smith 1969, Cumming & Cooper 1993). Adults can be captured using flight interception traps like Malaise trap, emergence trap, yellow pan traps and by sweeping with entomological nets on the vegetation. Adults should not be placed in alcohol or other liquids that remove bristles and coloration patterns necessary for their identification, except when you want to perform molecular analyses. Large adults should be mounted on pin numbers 1 or 2 as soon as they die, or placed in cotton layers in case they cannot be mounted immediately. Larvae generally live in moist soil, decaying roots or water bodies and are predators of others small organisms (Cumming & Cooper 1993). Some species with aquatic larvae are predators of larvae of blackflies (Hamada 1993). Empidids are cosmopolitan and are represented by 3,000 species in the World, however, it is possible that there are approximately 10,000 undescribed species. About 700 species are known in the Neotropical Realm in about 70 genera (Yang et al. 2007). Large gaps in the taxonomic knowledge of the Empididae remain for most regions of the World (Sinclair & Cumming 2006). The knowledge of the Neotropical fauna is restricted to few countries and virtually all genera need to be reviewed in a broader context for this region; the most well-known fauna belonging to Argentina, Brazil and Chile. The Colombian fauna is especially poorly known. Considering it is a highly diverse country any survey will result certainly in many new species and many new records increasing significantly the number of previously recorded species. Nothing is known of phenology, biology or behavior for Colombian empidids. Empididae belong to the superfamily Empidoidea considered a monophyletic lineage and a sister-group of Accepted by S. Nihei: 10 Apr. 2015; published: 14 Jun. 2016 393 Cyclorrapha (Sinclair & Cumming 2006). Recently Sinclair & Cumming (2006), based on cladistic morphological characters, justified its division into several families: Empididae, Hybotidae, Atelestidae, Brachystomatidae and Dolichopodidae. This chapter deals only with Empididae sensu Sinclair & Cumming (2006) comprising the following subfamilies/tribes: Empidinae (Empidini and Hilarini), Heremodromiinae (Chelopodini and Hemerodromiini) and Clinocerinae, whose position in the family is uncertain. Bearing in mind the great variety of environments that the Colombian territory offers, we believe that the actual number of species in the country is a lot larger than that recorded to date. Hence it is to be expected that collection in different regions of the country as well as the development of taxonomic expertise to identify the material will result in notable advances in knowledge of taxonomy, biology and distribution of the family in Colombia. Acronyms used for the depositories CAS—Department of Entomology, California Academy of Sciences, San Francisco, USA. CEUA—Colección Entomológica de la Universidad de Antioquia, Medellin, Colombia INPA—Instituto Nacional de Pesquisas da Amazônia, Manaus, Brazil. NHM—Natural History Museum, Vienna, Austria. List of abbreviations cat.—catalogue des.—designation distr.—distribution HT—holotype mon.—monotypy p, ; pp.—page(s) refs.—references rev.—revision subs. des.—subsequent designation F (f#)—Female M (m#)—Male Catalogue of Empididae of Colombia Family Empididae Subfamily Empidinae Schiner, 1862 Tribe Empidini Schiner, 1862 Genus Lamprempis Wheeler & Melander Lamprempis Wheeler & Melander, 1901: 366. Type species, Empis chichimeca Wheeler & Melander, 1901 (des. Coquillett 1903: 252). columbi Schiner, 1868: 208 (Empis). Type locality: Colombia. HT F (probably NHM). Refs.: Smith, 1967 (cat.); Yang et al., 2007 (cat.). 394 · Zootaxa 4122 (1) © 2016 Magnolia Press RAFAEL & CÂMARA Genus Macrostomus Wiedemann Macrostomus Wiedemann, 1817: 59. Type species, Hybos ferrugineus Fabricius (mon.). Refs.: Smith, 1967: 29 (cat.). alpinus Rafael & Cumming, 2006: 50. Type locality: Colombia, Cauca, Popayan. HT M (CAS). Distr.: Colombia, Ecuador. Refs.: Rafael & Cumming, 2006; Yang et al., 2007 (cat.). occidentalis Rafael & Cumming, 2006: 55. Type locality: Colombia, Cauca, Buenaventura. HT M (CAS). Distr.: Colombia. Refs.: Rafael & Cumming, 2006; Yang et al., 2007 (cat.). Genus Opeatocerata Melander Opeatocerata Melander 1928: 135. Type species: Empis rubida Wheeler & Melander, 1901. agudeloi Câmara & Rafael, 2014: 507. Type locality: Letícia, Colombia. HT M (CEUA). Distr.: Colombia. Refs.: Câmara & Rafael, 2014 (rev.). tanimboca Câmara & Rafael, 2014: 541. Type locality: Letícia, Colombia. HT M (CEUA). Distr.: Colombia; Brazil. Refs.: Câmara & Rafael, 2014 (rev.). Tribe Hilarini Collin, 1961 Genus Hilarempis Bezzi Hilarempis Bezzi, 1905: 443. Type species, Hilarempis nudifacies Bezzi, 1905 (des. Melander, 1928). vanellus Schiner, 1868: 206 (Hilara). Type locality: Colombia. HT M (probably NHM). Distr.: Colombia. Refs.: Smith, 1967 (cat.); Yang et al., 2007 (cat.). Acknowledgements We thank Dr. Adrian Plant for helping with English; the Conselho Nacional de Desenvolvimento Científico e Tecnológico (CNPq) for the award of a fellowship to J.A.R. (grant 300.305/2007–9), for the Ph.D. scholarship to J.T.C. (proc. 551.991/2011–9) and for financial resources (grant 401.243/2012–5). To Fundação de Amparo à Pesquisa do Estado do Amazonas (FAPEAM) by financial support (edital numbers 021/2011 and 020/2013). To Dra Marta Wolff for the loan of CEUA specimens. TABLE 1. Empididae species recorded from Colombia. Subfamily Tribe Genus Species Autor Empidinae Empidini Lamprempis columbi (Schiner, 1868) Macrostomus alpinus Rafael & Cumming, 2006 Macrostomus occidentalis Rafael & Cumming, 2006 Opeatocerata agudeloi Câmara & Rafael, 2014 Opeatocerata tanimboca Câmara & Rafael, 2014 Hilarempis vanellus (Schiner, 1868) Hilarini CATALOGUE OF EMPIDIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 395 References Bezzi, M. (1905) Empididae neotropicae Musei Nationalis Hungarici. Annales Historia Naturalis Musei Nationalis Hungarici, 33, 424–460. Câmara, J.T. & Rafael, J.A. (2014) Revision of Opeatocerata Melander, 1928 (Diptera: Empididae: Empidinae). Zootaxa 3846: 502–546. Coquillett, D.W. (1903) The genera of the dipterous family Empididae, with notes and new species. Proceedings of the Entomological Society of Washington, 5, 245–272. Cumming, J.M. & Cooper, B.E. (1993) Techniques for obtaining adult-associated immature stages of predacious tachydromiine flies (Diptera: Empidoidea), with implications for rearing and biocontrol. Entomological News, 104, 93–101. Hamada, N. (1993) Association between Hemerodromia sp. (Diptera, Empididae) and Simulium perflavum (Diptera, Simuliidae) in Central Amazonia, Brazil. Memórias do Instituto Oswaldo Cruz, 88 (1), 169–170. http://dx.doi.org/10.1590/S0074-02761993000100029 Melander, A.L. (1928) Diptera, Fam. Empididae. In: Wytsman, O. (Ed.), Genera Insectorum. Fasc. 185. Louis DesmetVerteneuil, Bruxelles, 434 pp. Rafael, J.A & Cumming, J.M. (2006) New species of Macrostomus Wiedemann (Diptera, Empididae) from Colombia, Ecuador and Peru. Zootaxa, 1136, 49–64. Schiner, J.R. (1868) Diptera. In: Wüllerstorf-Urbair, B. von (Ed.), Reise der Österreichischen Fregatte Novara um die Erde in den Jahren 1857, 1858, 1859, unter den Befehlen des Commodore B. Von Wuelerstorf-Urbair. Zoologischer Teil 2, Abt. I (B). K. Gerold’s Son, Wien, pp. 1–388. http://dx.doi.org/10.5962/bhl.title.7913 Sinclair, B.J. & Cumming, J.M. (2006) The morphology, higher-level phylogeny and classification of the Empidoidea (Diptera). Zootaxa, 1180, 1–172. Smith, K.G.V. (1967) 39. Family Empididae, In: Papavero, N. (Ed.), A Catalogue of Diptera of Americas South of the United States. Departamento de Zoologia da Secretaria da Agricultura do Estado de São Paulo, São Paulo, pp. 67. Smith, K.G.V. (1969) The Empididae of southern Africa (Diptera). Annals of Natal Museum, 19, 1–342. Steyskal, G.C. & Knutson, L.V. (1981) Empididae. In: McAlpine, J.F., Peterson, B.V., Shewell, G.E., Teskey, H.J., Vockeroth, J.R. & Wood, D.M. (Eds.), Manual of Nearctic Diptera. Vol. 1. Research Branch Monograph 27, Agriculture Canada, Ottawa, pp. 607–624. Tuomikoski, R. (1952) Über die Nahrung der Empididen-Imagines (Dipt.) in Finnland. Annales Entomologicae Fennicae, 18 (4), 170–181. Wheeler, W.M & Melander, A.L. (1901) Empididae In: Godman, F.D. & Salvin, O. (Eds.), Biologia Centrali Americana. Zoologia-Insecta-Diptera. Vol. 1. Supplement. Taylor & Francis, London, pp. 366–376. Wiedemann, C.R.W. (1817) Ueber einige neue Fliegen-Gattungen. Zoologischen Magazin, 1, 57–60. Yang, D., Zhang, K., Yao, G. & Zhang, J. (2007) World Catalog of Empididae (Insecta: Diptera). China Agricultural University Press, Beijing, vi + 599 pp. 396 · Zootaxa 4122 (1) © 2016 Magnolia Press RAFAEL & CÂMARA Zootaxa 4122 (1): 397–403 http://www.mapress.com/j/zt/ Copyright © 2016 Magnolia Press Article ISSN 1175-5326 (print edition) ZOOTAXA ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.4122.1.35 http://zoobank.org/urn:lsid:zoobank.org:pub:C6C0A4C6-A104-41A6-AF51-3756ECE2754E FAMILY HYBOTIDAE ROSALY ALE-ROCHA1 & RAFAEL AUGUSTO PINHEIRO DE FREITAS-SILVA2 1 Coordenação de Biodiversidade, Instituto Nacional de Pesquisas da Amazônia, Av.André Araújo, 2936, Petrópolis, CEP 69067-375, Manaus, Amazonas, Brazil. E-mail: alerocha@inpa.gov.br 2 Fellowship CNPq; Graduate Program in Entomology; E-mail: rfreitas.silva@yahoo.com.br *Corresponding author: alerocha@inpa.gov.br Abstract Hybotidae is a diverse family of the order Diptera, suborder Brachycera, superfamily Empidoidea. Hybotid flies are generally yellow to black, are morphologically diverse, and are distributed worldwide. The monophyly of the family is based on: palpifer and fore tibial gland present, laterotergite bare and R4+5 unbranched. Hybotids are mainly predators and are usually found on the vegetation, logs and other surfaces, or in flight during displacement and forage. This catalogue, based on the study of specimens and available literature records, summarizes and updates the information on the Colombian fauna of Hybotidae, which includes 19 species distributed in six genera. Key words: Colombia, distribution, Hybotidae, Neotropical Region Introduction Hybotidae are cosmopolitan, with approximately 1,700 species described in 66 genera. Of these, 187 species in 25 genera have been documented for South America (Yang et al. 2007). Hybotid flies occur virtually in all terrestrial biotopes and in several environmental conditions (Chvála 1975). Adults are small to medium in size (1.0-7.5mm), black to yellow, never metallic, and are morphologically diverse. Larvae have the terminal segment rounded with a protuberance below the posterior spiracles; locomotor projections on the first seven abdominal segments, and cuticular spines organized in rows. The pupae are very diverse and usually have short respiratory horns or pedicellate spiracles on thorax and abdomen, but the prothoracic spiracles are much shorter than those found in Dolichopodidae (Dyte 1967). Even though hybotids are primarily predators, there are some records of species that feed on pollen and nectar (Chvála 1976, Melander 1906, Downes & Smith 1969). Hybotid flies are normally found on leaves, trunks and other surfaces, or in flight during displacement and foraging. Ocydromiinae flies swarm as part of their mating and courtship behavior, but in Hybotinae and Tachydromiinae flies the predatory and reproductive activities have been displaced to solid surfaces such as leaves, the ground and tree trunks (Chvála 1976). The biology of the immatures is virtually unknown, but there is a suspicion that they are predators (Cumming & Cooper 1993) and develop on the ground, on moss, in tree holes (Chvála 1975), litter or rotting wood (Steyskal & Knutson 1981). The most common methods of collecting hybotids are traps that intercept them during flight, for instance Malaise and Suspended traps, colored pan traps (primarily the yellow ones), and swapping with entomological nets. The first phylogenetic proposal for the classification of Hybotidae was advanced by Chvála (1983), who included in it three subfamilies traditionally placed in Empididae s. lat. (namely Hybotinae, Ocydromiinae and Tachydromiinae). Presently, the family includes the subfamilies Hybotinae, Ocydromiinae, Oedaleinae, Tachydromiinae and Trichininae, which are clustered based on the following synapomorphies: presence of palpifer (special sclerite of the mouthparts joining the stipes with the palpi), fore tibial gland present, gonocoxal apodeme restricted to anterolateral margin of hypandrium, laterotergite bare and vein R4+5 unbranched (Sinclair & Cumming 2006). Accepted by S. Nihei: 10 Apr. 2015; published: 14 Jun. 2016 397 Species from three hybotid subfamilies occur in the neotropics: Hybotinae, Ocydromiinae and Tachydromiinae. The first one, Hybotinae, include 543 species in 17 genera, being that 83 species in eight genera occur in South America, with 17 species in Colombia (Ale-Rocha 2007, 2008; Ale-Rocha & Vieira 2008; Yang et al. 2007). The second, Ocydromiinae, include 84 species in 15 genera worldwide, with 31 species in eight genera in South America, and one species registered for Colombia (Sinclair & Cumming 2007; Yang et al. 2007). Finally, Tachydromiinae, the largest of the three, include 1,201 species in 27 genera (Shamshev & Grootaert, 2012) occurring in all biogeographical regions (except the Antarctic); in South America, there are 73 species in nine genera, but only one species is known to occur in Colombia (Table 1). Acronyms used for the depositories CAS—California Academy of Sciences, San Francisco, U.S.A CEUA—Colección Entomológica, Universidad de Antioquia, Medellín, Colombia IBUNAM—Instituto de Biología de la Universidad Nacional Autónoma de México INPA—Instituto Nacional de Pesquisas da Amazônia, Manaus, Brazil NHM—Natural History Museum, London, England SNSD—Senckenberg Naturhistorische Sammlungen Dresden List of abbreviations cat.—catalogue cit.—citation diagn.—diagnosis distr.—geographic distribution mon.—by monotypy orig. des.—by original designation refs.—references subs. des.—by subsequent designation pp.—pages preocc.—preoccupied rev.—revision redesc.—redescription part.—partim HT—Holotype LT—Lectotype F (f#)—Female M (m#)—Male. Catalogue of Hybotidae of Colombia Family Hybotidae Meigen, 1820 Subfamily Hybotinae Meigen, 1820 Genus Cerathybos Bezzi Cerathybos Bezzi, 1909: 303. Type species, Cerathybos schnusei Bezzi, 1909 by mon. Refs.: Smith, 1967: 14 (cat.); Rafael & Ale-Rocha, 1995: 519 (rev. Bezzi’s types); Yang et al., 2007: 281 (cat.). 398 · Zootaxa 4122 (1) © 2016 Magnolia Press ALE-ROCHA & FREITAS-SILVA bezzii Ale-Rocha, 2002: 294. Type locality: Ecuador, Napo, Muyuna. HT M [NHM]. Distr.: Bolivia, Colombia (Putumayo (Mocoa)), Equador. Refs.: Yang et al., 2007: 282 (cat.). schnusei Bezzi, 1909: 303. Type locality: Unini, Ucayale River, Peru. HT F [SNSD]. Distr.: Brasil, Colombia (Putumayo (Mocoa)), Peru. Refs.: Ale-Rocha & Rafael 1995: 189 (key); Ale-Rocha, 2002: 297 (distr.); Yang et al., 2007: 282 (cat.). Genus Euhybus Coquillett Euhybus Coquillett, 1895: 437. Type species, Hybos purpureus Walker, 1849 by subs. des. of Coquillett, 1903. Refs.: Smith, 1967: 11 (cat.); Rafael & Ale-Rocha, 1995: 521 (rev. Bezzi’s types); Yang et al., 2007: 282 (cat.). amazonicus Ale-Rocha, 2002: 304. Type locality: Equador, Morona, Santiago, Sierra Tucutu. HT M NHM]. Distr.: Brazil, Colombia (Putumayo (Villa Garzon)), Ecuador. Refs.: Ale-Rocha & Vieira, 2008: 114 (distr.); Yang et al., 2007: 283 (cat.). eurypterus (Bezzi), 1909: 306 (Hybos). Type locality: Peru, Pichis River, Puerto Bermúdez, Mouth of the river Pachitea. LT M [SNSD]. Distr.: Bolivia, Brazil, Colombia (Caquetá), Ecuador, Peru, Trindad. Refs.: Smith, 1967: 12 (cat.); Rafael & Ale-Rocha, 1995: 521 (rev. Bezzi’s types); Ale-Rocha, 2002: 311 (dist.); Ale-Rocha & Vieira, 2008: 115 (dist.); Yang et al., 2007: 284 (cat.). niger Ale-Rocha & Rafael, 2004: 363. Type locality: Ecuador, Napo. HT M [NHM]. Distr.: Colombia (Puntumayo, Caqueta), Ecuador. Refs.: Yang et al., 2007: 284 (cat.). pilosus (Schiner), 1868: 202 (Hybos). Type locality: Colombia. HT M [location?]. Distr.: Bolivia, Colombia. Refs.: Smith, 1967: 13 (cat.); Yang et al., 2007: 285 (cat.). setulosus Ale-Rocha, 2002: 316. Type locality: Colombia (Puntumayo (Mocoa)). HT M [NHM]. Distr.: Colombia (Puntumayo). Refs.: Yang et al., 2007: 285 (cat.). stigmaticuss (Schiner), 1868: 202 (Hybos). Type locality: Colombia. HT M [location?]. Distr.: Colombia. Refs.: Yang et al., 2007: 286 (cat.). Genus Lactistomyia Melander Lactistomyia Melander, 1902: 250. Type species, Lactistomyia insolita Melander, 1902 by mon. Refs.: Melander, 1928: 35 (key); Coquillett, 1903: 251 (as subgenus of Hybos); Bezzi, 1909: 311 (discussion); Smith, 1967: 14 (cat.); Yang et al., 2007: 284 (cat.); Ale-Rocha, 2008: 1 (rev.). hyalina Bezzi, 1909: 312. Type-Locality: Peru, Pasco, Pichis, Puerto Bermudez. HT F [SNSD]. Distr.: Colombia (Narino (Barbacoas)), Peru. Refs.: Collin, 1933: 28 (cit.); Curran, 1931: 6 (key); Smith, 1967:14 (cat.); Rafael & Ale-Rocha, 1995: 532 (rev. Bezzi’s types); Ale-Rocha, 2008: 4, 5 (key, distr.). Genus Neohybos Ale-Rocha & Carvalho Neohybos Ale-Rocha & Carvalho, 2003: 3. Type species, Hybos luridus Bezzi, 1909 by orig. des. Refs.: Ale-Rocha & Rafael, 2004: 497 (distr.); Yang et al., 2007: 300 (cat.) Hybos Meigen, 1803: 269 (part). Euhybus Coquillett, 1895: 437 (part). CATALOGUE OF HYBOTIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 399 brunnescens Ale-Rocha, 2007: 33, 37. Type locality: Colombia, Huila, Cueva de los Guacharos. HT M [NHM]. Distr.: Colombia (Huila (Cueva de Los Guacharos)). colombiensis Ale-Rocha, 2007: 34, 42. Type locality: Colombia, Putumayo, Villa Garzon. HT M [NHM]. Distr.: Colombia (Putumayo). cooperi Ale-Rocha, 2007: 34, 42. Type locality: Colombia, Huila, Culva of los Guacharis. HT M [NHM]. Distr.: Colombia (Huila). elegans Ale-Rocha, 2007: 35, 43. Type locality: Ecuador, Napo, Tena. HT M [NHM]. Distr.: Colombia (Valle del Cauca (Buena Ventura), Nariño (Barbacoas)), Ecuador. hallexus (Smith), 1963: 32, 156 (Euhybus). Type locality: Guiana, Kaieteur. HT M [NHM]. Distr.: Brazil, Colombia, Guiana, Peru. Refs.: Smith, 1967: 12 (cat.); Ale-Rocha & Carvalho, 2003: 8 (rev.); Ale-Rocha, 2007: 34 (key); Yang et al., 2007: 300 (cat.). rossi Ale-Rocha, 2007: 35, 48. Type locality: Colombia (Villaviceneio (3 m. w.)). HT M [CAS]. Distr.: Colombia (Meta). schlingeri Ale-Rocha, 2007: 34, 48 (key, desc.). Type locality: Colombia, Tolima, 14 mi. W. Fresno. HT M [CAS]. Distr.: Colombia (Tolima). tenuis Ale-Rocha, 2007: 35, 51. Type locality: Ecuador, Napo, Tena. HT M [NHM]. Distr.: Colombia (Tolima), Ecuador. Subfamily Ocydromiinae Schiner, 1862 Genus Chvalaea Papp & Földvári Chvalaea Papp & Földvári, 2002: 354. Type species, Leptopeza rugosiventris Strobl, 1910 by orig. des. Refs.: Ale-Rocha, 2006: 17 (neotropical species); Yang et al., 2007: 329 (cat.). amazonica Ale-Rocha, 2006: 19. Type locality: Brazil, Amazonas, Santa Izabel do Rio Negro. HT M[INPA]. Distr.: Brazil, Colombia (Antioquia), Peru. Refs.: Yang et al. 2007: 329 (cat.). Examined material. COLOMBIA: Antioquia: Medellin, Santa Elena, Parque Piedras Blancas, 1 male, 03 vii.2013 (CEUA). Subfamily Tachydromiinae Schiner, 1862 Genus Elaphropeza Macquart Elaphropeza Macquart, 1827: 86. Type species, Tachydromia ephippiata Fallén, 1815 by mon. Refs.: Melander, 1928: 305 (cat., part); Smith, 1962: 202 (key); 1963: 153; 1967: 4 (cat.); Yang et al., 2007: 370 (cat.); Freitas-Silva & Ale-Rocha, 2009: 33 (key). Ctenodrapetis Bezzi, 1904: 351 (as subgenus of Drapetis). flavida (Williston), 1896: 308 (Drapetis). Type locality: West Indies, Saint Vincent, Windward side. Distr.: Saint Vincent, Brazil, Colombia (Meta, Valle del Cauca), Cuba, Ecuador, Haiti, Jamaica, Mexico, Peru, Puerto Rico, U.S.A. Refs.: Melander, 1928: 314 (cat.); Smith, 1967: 39.4 (cat.); Yang et al., 2007: 373 (cat.); Freitas-Silva & Ale-Rocha, 2009: 36 (rev.). bergonzoi Raffone, 2000: 3. Refs.: Freitas-Silva & Ale-Rocha, 2009: 39. 400 · Zootaxa 4122 (1) © 2016 Magnolia Press ALE-ROCHA & FREITAS-SILVA Remarks Three additional species, which are probably undescribed, have been collected from Colombia. They are housed in CEUA and belong to Hybos Meigen (previously only recorded from Central America and southern Brazil), Hoplopeza Bezzi (from Chile) and Syneches Walker (see table). This is the first record of Hybos and Hoplopeza for Colombia. Acknowledgements We sincerely thank Marta Wolff (Universidad de Antioquia, Colombia) for loaning the material examined. We also thank Coordenação de Aperfeiçoamento de Pessoal de Nível Superior (CAPES) and Fundação de Amparo à Pesquisa do Estado do Amazonas (FAPEAM) for the support to this work by the project “Incremento do conhecimento da diversidade, formação e fixação de especialistas em Sistemática de Diptera (Insecta) na Amazônia” and Conselho Nacional de Desenvolvimento Científico e Tecnológico (CNPq) by the project “Sistemática morfológica de Hybotidae (Diptera, Empidoidea) com ênfase nas espécies neotropicais de Ocydromiinae e Hybotinae”. TABLE 1. Hybotidae species recorded from Colombia. Subfamily Tribe Genus species Hybotinae Author Meigen, 1820 Hybotini Meigen, 1820 Cerathybos Bezzi, 1909 bezzii Ale-Rocha, 2002 schnusei Bezzi, 1909 Euhybus Coquillett, 1895 amazonicus Ale-Rocha, 2002 eurypterus (Bezzi, 1909) niger Ale-Rocha & Rafael, 2004 pilosus (Schinner, 1868) setulosus Ale-Rocha, 2002 stigmaticuss (Schinner, 1868) Hybos Meigen, 1803 sp. Lactistomyia Melander, 1902 hyalina Neohybos Syneches Bezzi, 1909 Ale-Rocha & Carvalho, 2003 brunnescens Ale-Rocha, 2007 colombiensis Ale-Rocha, 2007 cooperi Ale-Rocha, 2007 elegans Ale-Rocha, 2007 hallexus (Smith, 1963) rossi Ale-Rocha, 2007 schlingeri Ale-Rocha, 2007 tenuis Ale-Rocha, 2007 Walker, 1852 ...Continued on next page CATALOGUE OF HYBOTIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 401 TABLE 1. (Continued) Subfamily Tribe Genus species Author sp. Ocydromiinae Schiner, 1862 Chvalaea Papp & Földvári, 2002 amazonica Hoplopeza Ale-Rocha, 2006 Bezzi, 1909 sp Tachydromiinae Meigen, 1822 Elaphropeza Macquart, 1827 flavida (Williston, 1896) References Ale-Rocha, R. (2002) Revisão do gênero Euhybus Coquillett (Diptera, Empididae, Hybotinae) da Região Neotropical. Grupo dimidiatus. Acta Amazonica, 32 (2), 299–324. http://dx.doi.org/10.1590/1809-43922002322324 Ale-Rocha, R. 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Transactions of the Entomological Society of London, 44 (3), 253–449. http://dx.doi.org/10.1111/j.1365-2311.1896.tb00965.x Yang, D., Zhang, K., Yao, G. & Zhang, J. (2007) World catalog of Empididae (Insecta: Diptera). Chine Agricultural University Press, Pequim, 599 pp. CATALOGUE OF HYBOTIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 403 Zootaxa 4122 (1): 404–413 http://www.mapress.com/j/zt/ Copyright © 2016 Magnolia Press Article ISSN 1175-5326 (print edition) ZOOTAXA ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.4122.1.36 http://zoobank.org/urn:lsid:zoobank.org:pub:1EB8F6DE-DC7B-4C15-9D71-91A7F91C201E FAMILY DOLICHOPODIDAE RENATO SOARES CAPELLARI Instituto Federal de Educação, Ciência e Tecnologia do Triângulo Mineiro, Campus Uberlândia. Caixa Postal 1020. Uberlândia, Minas Gerais, Brasil, 38400-970. E-mail: rscapellari@gmail.com Abstract The Dolichopodidae recorded from Colombia are listed based on literature data only and currently comprise 30 species in 14 genera (six subfamilies). Information on type material and depository institutions is given. Key words: Catalogue, Colombia, Diptera, Dolichopodidae Introduction Dolichopodidae are a large and diverse lineage of Empidoidea (along with Atelestidae, Brachystomatidae, Hybotidae and Empididae), with over 7,600 described species in some 250 genera from all zoogeographical regions (Grichanov 2014, Yang et al. 2006). Around 1,200 dolichopodid species and 75 genera are known from Neotropics, a small fraction of the world fauna which is obviously a subestimative of the real number for that Region. Adults of the family are predators, usually feeding on small soft-bodied arthropods and annelids (see Ulrich 2004 for a review) and found virtually everywhere, but tending to be more numerous in moist habitats, like riparian vegetation and humid forests. Adult dolichopodids can be usually recognized by their slender and elongate legs, metallic body color and reduced wing venation. Males often display distinctive modifications involving chaetotaxy, sculptures and enlargement of body parts (e.g., palpus, antenna, legs and wings) which are assumed to play a role during male-female recognition during courtship. The Colombian dolichopodid fauna is very poorly studied, and most of the 30 recorded species (14 genera; summarized in Table 1) are reported only by their original description and known just from the type locality. The species list provided below is based on literature data only, and the species whose distributional ranges overlap Colombia, like Plagioneurus univittatus Loew, Chrysotus spectabilis (Loew), and Chrysotus mundus (Loew), are omitted if not formally recorded from there. The classification adopted here follows that of Ulrich (1981), with few additions and modifications. Acronyms used for the depositories AMNH—American Museum of Natural History, New York, USA BMNH—The Natural History Museum, London, England CAS—Department of Entomology, California Academy of Sciences, San Francisco, USA CNC—Canadian National Collection of Insects, Ottawa, Canada HNHM—Hungarian Natural History Museum, Budapest, Hungary INPA—Instituto Nacional de Pesquisas da Amazônia, Manaus, Brazil MNHNP—Muséum National d´Histoire Naturelle, Paris, France NMW—Naturhistorisches Museum, Wien, Austria MPCB—Marc Pollet Collection, Belgium 404 Accepted by S. Nihei: 10 Apr. 2015; published: 14 Jun. 2016 MZSP—Museu de Zoologia da Universidade de São Paulo, São Paulo, Brazil SNSD—Senckenberg Naturhistorische Sammlung Dresden, Dresden, Germany USNM—National Museum of Natural History, Smithsonian Institution, Washington, D.C., USA ZMHB—Museum für Naturkunde der Humboldt-Universität, Berlin, Germany List of abbreviations aut.—automatic des.—designation mon.— by monotypy orig. des.— by original designation preocc.—preoccupied ref(s).—reference(s) rev.—revision subs.—subsequent HT—holotype PT—paratype ST—syntype F (f#)—Female M (m#)—Male Catalogue of Dolichopodidae of Colombia Family Dolichopodidae Latreille, 1809 Subfamily Achalcinae Grootaert & Meuffels, 1997 Genus Achalcus Loew Achalcus Loew, 1857a: 30. Type species, Porphyrops flavicollis Meigen, 1824: 56, subs. des. by Robinson (1970: 22). Ref.: Pollet, 2005 (rev.). cyanocephalus Pollet 2005: 41 (fig. 6). Type locality: Colombia, Cauca, 27 mi. S. of Popayán. HT M (CAS). Distr.: Colombia (Cauca). Ref.: Yang et al., 2006: 24 (cat.). Subfamily Diaphorinae Schiner, 1864 Genus Achradocera Becker Achradocera Becker 1922: 207. Type species, Achradocera femoralis Becker, subs. des. by Robinson (1970: 31). Refs.: Parent, 1933a (key); Bickel, 2000 (review). femoralis Becker, 1922: 208. Type locality: Trinidad, Port of Spain; Colombia, Baranquilla; & Paraguay, San Bernardino. ST 3 M (HNHM, destroyed). Distr.: Trinidad, Colombia (Atlántico (Barraquilla)). Paraguay. Refs.: Robinson, 1970: 32 (cat., as subgenus of Chrysotus); Yang et al., 2006: 31 (cat.). Genus Chrysotus Meigen Chrysotus Meigen, 1824: 40. Type species, Musca nigripes Fabricius, subs. des. by Westwood (1840: 134). Refs.: Parent, 1928; Van Duzee, 1931b (keys); Capellari & Amorim, 2012 (taxonomic notes). CATALOGUE OF DOLICHOPODIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 405 exceptus Becker, 1922: 190. Type locality: Colombia, Sierra San Lorenzo. HT M (HNHM, destroyed). Distr.: Colombia (Magdalena (Sierra San Lorenzo)). Refs.: Robinson, 1970: 29 (cat.); Yang et al., 2006: 94 (cat.). Genus Lyroneurus Loew Lyroneurus Loew, 1857b: 38. Type species, Lyroneurus coerulescens Loew, subs. des. by Robinson (1970: 35). Refs.: Parent, 1930a (key); Capellari & Amorim, 2010 (taxonomic notes). fratellus Becker, 1922: 183. Type locality: Colombia, Sierra San Lorenzo & Argentina, Lules. ST 2 M (HNHM, destroyed). Distr.: Colombia (Magdalena (Sierra San Lorenzo)), Argentina. Refs.: Robinson, 1970: 35 (cat., as subgenus of Diaphorus); Yang et al., 2006: 72 (cat., as Diaphorus). luteoviridis Parent, 1935: 95. Type locality: Colombia, Cordillera, near Rio Yurumanguí. HT M (MNHNP). Distr.: Colombia (Valle del Cauca), Brazil. Refs.: Robinson, 1970: 35 (cat., as subgenus of Diaphorus); Yang et al., 2006: 76 (cat., as Diaphorus). occultus Becker, 1922: 185. Type locality: Colombia, Sierra San Lorenzo. ST 2 M (HNHM, destroyed). Distr.: Colombia (Magdalena (Sierra San Lorenzo)). Refs.: Robinson, 1970: 36 (cat., as subgenus of Diaphorus); Yang et al., 2006: 78 (cat., as Diaphorus). Genus Symbolia Becker Symbolia Becker, 1922: 210. Type species, Symbolia ochracea Becker (mon.). Ref.: Robinson, 1966 (rev.). Cyrtosymbolia Parent, 1931: 10. Type species, Cyrtosymbolia sinuosa Parent (mon.). lugubris Parent, 1934a: 266 (pl. 68, figs. 15–17). Type locality: Colombia, Gorgona Is. HT M (BMNH). Distr.: Colombia (Gorgona Is.). Refs.: Robinson, 1970: 37 (cat.); Yang et al., 2006: 87 (cat.). Subfamily Dolichopodinae Latreille, 1809 Genus Cheiromyia Dyte Cheirocerus Parent, 1930b: 13. Type species, Cheirocerus palmaticornis Parent (mon.). Preoc. by Cheirocerus Eigenmann. Cheiromyia Dyte, 1980: 223. Type species, Cheirocerus palmaticornis Parent, aut. Replacement name for Cheirocerus Parent, 1930b. Refs.: Capellari & Amorim, 2009 (taxonomic notes); Brooks et al., 2010 (rev.). brevitarsis Brooks, 2010 in Brooks et al., 2010: 48 (figs. 2D, 4, 8A): 48. Type locality: French Guiana, Sinnamary. HT M (CNC), PT 24 (BMNH, CNC, INPA, MPCB, MZSP, USNM). Distr.: Colombia (Meta), Guyana, French Guiana, Brazil. Ref.: Capellari & Amorim, 2009: 59 (taxonomic notes). Genus Paraclius Loew Paraclius Loew, 1864: 97. Type species, Pelastoneurus arcuatus Loew, subs. des. by Coquillett (1910: 583). Leptocorypha Aldrich, 1896: 315. Type species: Leptocorypha pavo Aldrich (mon.). Ref.: Van Duzee, 1930 (key). difficilis Becker, 1922: 47. Type locality: Trinidad, Port of Spain & Colombia, “S. Maria” [sic? =Santa Marta?]. ST 6M, 1F (HNHM, destroyed). Distr.: Trinidad, Colombia. Refs.: Robinson, 1970: 48 (cat.); Yang et al., 2006: 184 (cat.). 406 · Zootaxa 4122 (1) © 2016 Magnolia Press CAPELLARI mixtus Parent, 1934a: 269 (pl. 69, figs. 24–26). Type locality: Colombia, Gorgona Is. HT M, PT F (BMNH). Distr.: Colombia (Gorgona Is.). Refs.: Robinson, 1970: 48 (cat.); Yang et al., 2006: 187 (cat.). Genus Pelastoneurus Loew Paracleius Bigot, 1859: 215, 227. Type species, Dolichopus heteronevrus Macquart (mon.). Senior syn. of Pelastoneurus Loew, 1861. Suppressed by the I.C.Z.N (2004) (see Brooks et al., 2002). Pelastoneurus Loew, 1861: 36. Type species, Pelastoneurus vagans Loew, subs. des. by Coquillett (1910: 586). Ref.: no suitable keys or revisions are available for Neotropical species. Metapelastoneurus Aldrich, 1894: 152. Type species, Metapelastoneurus kansensis Aldrich (mon.). Sarcionus Aldrich, 1901: 341. Type species, Pelastoneurus lineatus Aldrich (orig. des.) Phylarchus Aldrich, 1901: 342. Type species, Phylarchus tripartitus Aldrich (mon.). Preoc. by Phylarchus Simon. Paraclius Kertész, 1909: 230. Type species, Dolichopus heteronevrus Macquart, aut. Unjustified emendation of Paracleius Bigot, 1859. Proarchus Aldrich, 1910: 100. Type species, Phylarchus tripartitus Aldrich, aut. New name for Phylarchus Aldrich, 1901. nigritibia Parent, 1934a: 271 (pl. 70: figs. 30, 31). Type locality: Colombia, Villavicencio, Quatiquia River. HT M (BMNH). Distr.: Colombia (Meta (Villavicencio)). Refs.: Robinson, 1970: 51 (cat.); Yang et al., 2006: 196 (cat.). umbripictus Becker, 1922: 73. Type locality: USA, Carolina & Colombia, Barranquilla, Bogotá. ST 2F (ZMHB). Distr.: USA, Mexico, Belize, Colombia (Atlántico (Barranquilla), Cundinamarca (Bogotá)). Refs.: Robinson, 1970: 51 (cat.); Yang et al., 2006: 197 (cat.). Genus Tachytrechus Haliday Ammobates Stannius, 1831a: 33. Type species, Ammobates notatus Stannius, subs. des. by Rondani, 1856: 143. Preocc. by Ammobates Latreille. Tachytrechus Stannius, 1831c: 261 (nomen nudum, see Brooks 2005). Tachytrechus Haliday, 1851: 173. Type species, Ammobates notatus Stannius, automatic. Replacement name for Ammobates Stannius, 1831a. Ref.: no suitable keys or revisions are available for Neotropical species. Stannia Rondani, 1857: 14. Type species: Ammobates notatus Stannius, automatic. Unnecessary new name for Ammobates Stannius, 1831a. Gongophora Philippi, 1875: 86. Type species, Gongophora medinae Philippi (mon.). Polymedon Osten Sacken, 1877: 317. Type species, Polymedon flabellifer Osten Sacken (mon.). Macellocerus Mik, 1878: 5. Type species, Tachytrechus moechus Loew (orig. des.). Psilischium Becker, 1922: 93. Type species, Psilischium laevigatum Becker (mon). Gonioneurum Becker 1922: 98. Type species, Gonioneurum varum Becker (mon.). Syntomoneurum Becker, 1922: 123. Type species, Syntomoneurum alatum Becker (mon.). analis (Parent, 1954): 226 (figs. 7–10) (Syntomoneurum); Brooks (2005). Type locality: Colombia, Boyacá, Muzo. HT M (MNHNP). Distr.: Colombia (Boyacá (Muzo)). Refs.: Robinson, 1970: 58 (cat., as Syntomoneurum in subfamily Hydrophorinae); Brooks & Wheeler, 2002: 317, figs. 5–7 (redescription); Yang et al., 2006: 208 (cat.); Brooks & Cumming, 2008: 5, fig. 2 (as Tachytrechus, diagnosis). giganteus (Brooks, 2002) in Brooks & Wheeler, 2002: 321 (figs. 12–22) (Syntomoneurum); Brooks (2005). Type locality: Colombia, Antioquia, 10 km E of Medellin. HT M, PT M and PT 2F (USNM). Distr.: Colombia (Antioquia). Refs.: Yang et al., 2006: 210 (cat.); Brooks & Cumming, 2008: 14, fig. 6 (as Tachytrechus, diagnosis). potens (Parent, 1934a): 271 (pl. 70: fig. 29) (Paraclius); Robinson (1970). Type locality: Colombia, Gorgona Is. HT M (BMNH). Distr.: Colombia (Gorgona Is.). Refs.: Robinson, 1970: 55 (cat.); Yang et al., 2006: 215 (cat.). varus (Becker, 1922): 99 (figs. 37, 38) (Gonioneurum); Brooks (2005). Type locality: Colombia, Sierra San Lorenzo. ST 2 M (HNHM, destroyed). Distr.: Colombia (Magdalena (Sierra San Lorenzo)). Refs.: Robinson, 1970: 46 (cat., as Gonioneurum); Yang et al., 2006: 217 (cat.). CATALOGUE OF DOLICHOPODIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 407 Subfamily Medeterinae Lioy, 1863-64 Genus Medetera Fischer von Haldheim Medetera Fischer von Waldheim, 1819: 7. Type species, Medetera carnivora Fischer von Waldheim [=Musca diadema Linnaeus] (mon.). Refs.: Van Duzee, 1929, 1931a (keys). Taechobates Haliday, 1832: 356. Type species, Hydrophorus jaculus Fallén, subs. des. by Coquillett (1910: 611). Orthobates Wahlberg, 1844: 109. Type species, Hydrophorus jaculus Fallén, subs. des. by Coquillett (1910: 581). Anorthus Loew, 1850: 117. Type species, Hydrophorus jaculus Fallén (mon.). Oligochaetus Mik, 1878: 7. Type species, Medeterus plumbellus Meigen (orig. des.) Saccopheronta Becker, 1914: 125. Type species, Saccopheronta nudipes Becker (mon.). Elongomedetera Hollis, 1964: 26. Type species, Elongomedetera thoracica Hollis (orig. des.). Asioligochaetus Negrobov, 1966: 877 (as subgenus). Type species, Oligochaetus vlasovi Stackelberg (orig. des.). Lorea Negrobov, 1966: 878 (as subgenus). Type species, Medetera spiniger Stackelberg (orig. des.). pedestris Becker, 1922: 137 (fig. 58). Type locality: Peru, Ucayali, Unini, mouth of River Pachitea. ST 6 (SNSD) (Kallweit & Negrobov 1994). Distr.: Peru, Surinam, Colombia. Refs.: Parent, 1935: 96 (distr.); Robinson, 1970: 19 (cat.); Yang et al., 2006: 298 (cat., as Saccopheronta). Subfamily Sciapodinae Bigot, 1917 Genus Condylostylus Bigot Condylostylus Bigot, 1859: 215. Type species, Psilopus bituberculatus Macquart (orig. des.). Ref.: Parent, 1929 (key). Dasypsilopus Bigot, 1859: 215. Type species, Psilopus pilipes Macquart (orig. des.) Eurostomerus Bigot, 1859: 215. Type species, Psilopus coerulus Macquart, nomen nudum (=Eurostomerus coerulus Bigot, validated by generic description). Oedipsilopus Bigot, 1859: 224. Type species, Psilopus posticatus Wiedemann (orig. des.). Tylochaetus Bigot, 1888: xxiv. Type species: Psilopus bituberculatus Macquart (orig. des.). Laxina Curran, 1934: 230. Type species, Dolichopus patibulatus Say (orig. des.) biseta Becker, 1922: 353. Type locality: “Colombia” (Cordillera) ST 2M (ZMHB). Distr.: Colombia. Refs.: Robinson, 1970: 4 (cat.); Yang et al., 2006: 397 (cat.). cinctiventris Van Duzee, 1934: 371 (fig. 7). Type locality: Colombia, Sierra San Lorenzo, Vista Nieve. HT M (AMNH). Distr.: Colombia (Magdalena (Sierra San Lorenzo)). Refs.: Robinson, 1970: 5 (cat.); Yang et al., 2006: 398 (cat.). facetus Parent, 1933b: 169 (figs. 10–12). Type locality: Colombia, Gorgona Is. HT M (BMNH). Distr. Colombia (Gorgona Is.). Refs.: Robinson, 1970: 6 (cat.); Yang et al., 2006: 401 (cat.). gorgonensis Parent, 1933b: 175 (fig. 21). Type locality: Colombia, Gorgona Is. HT M (BMNH). Distr.: Colombia (Gorgona Is.). Refs.: Robinson, 1970: 7 (cat.); Yang et al., 2006: 403 (cat.). impatiens Becker, 1922: 310. Type locality: "Venezuela". ST 4M and 1F (NMW). Distr.: Colombia, Venezuela. Refs.: Robinson, 1970: 8 (cat.); Yang et al., 2006: 303 (cat.) loriferus (Parent, 1934b): 116 (figs. 9, 10) (Chrysosoma); Bickel (1994). Type locality: Colombia, Bogotá. HT M (MNHNP). Distr.: Colombia (Cundinamarca (Bogotá)). Refs.: Robinson, 1970: 2 (cat., as Chrysosoma); Yang et al., 2006: 405 (cat.). mirus Parent, 1931: 4 (pl. 1: figs. 8, 9). Type locality: Peru, Chanchamayo. HT M (SNSD). Distr.: Peru, Colombia. Refs.: Parent, 1935: 96 (distr.); Robinson, 1970: 9 (cat.); Yang et al., 2006: 406 (cat.). 408 · Zootaxa 4122 (1) © 2016 Magnolia Press CAPELLARI nubeculus Becker, 1922 (figs. 123, 124). Type locality: Colombia, Aracataca. ST M and 6F (HNHM, destroyed). Distr.: Colombia (Magdalena (Aracataca)). Refs.: Robinson, 1970: 9 (cat.); Yang et al., 2006: 407 (cat.). productipes Parent, 1934b: 121 (figs. 17, 18). Type locality: Colombia, Bogotá. HT M (MNHNP). Distr. Colombia (Cundinamarca (Bogotá)). Refs.: Robinson, 1970: 10 (cat.); Yang et al., 2006: 409 (cat.). squamifer Becker, 1922: 340 (figs. 134, 135). Type locality: Peru, Rio Urubamba & Colombia, Aracataca. ST M (SMTD); ST 2 M (HNHM, destroyed). Distr.: Peru, Colombia (Magdalena (Aracataca)). Refs.: Robinson, 1970: 12 (cat.); Kallweit & Negrobov, 1994 (type cat.); Yang et al., 2006: 411 (cat.). Genus Mesorhaga Schiner Mesorhaga Schiner, 1868: 217. Type species, Mesorhaga tristis Schiner (mon.). Refs.: Naglis, 2000; Bickel, 2007 (keys). Aptorthus Aldrich, 1893: 48. Type species, Aptorthus albiciliatus Aldrich, subs. des. by Coquillett (1910: 509). tristis Schiner, 1868: 217. Type locality: “Colombia”. ST 2 M (NMW). Distr.: Colombia. Refs.: Robinson, 1970: 14 (cat.); Yang et al., 2006: 429 (cat.). Subfamily Sympycninae Aldrich, 1905 Genus Pseudosympycnus Robinson Pseudosympycnus Robinson, 1967: 901. Type species, Sympycnus palpiger Van Duzee (orig. des.). Ref.: Robinson, 1967 (rev.). singularis (Parent, 1934a): 259 (pl. 67: figs. 6–9) (Sympycnus); Robinson (1970). Type locality: Colombia, Gorgona Is. ST 3 (BMNH). Distr.: Colombia (Gorgona Is.). Refs.: Robinson, 1970: 38 (cat.); Yang et al., 2006: 448 (cat.). Genus Stolidosoma Becker Stolidosoma Becker, 1922: 213. Type species, Stolidosoma permutans Becker, subs. des. by Robinson (1967: 893). Ref.: Robinson, 1967 (rev.). permutans Becker, 1922: 217 (figs. 88, 89). Type locality: Colombia, Sierra San Lorenzo. ST 3M and 3F (HNHM, destroyed). Distr.: Mexico, Panama, Colombia (Magdalena (Sierra San Lorenzo)), Guyana. Refs.: Robinson, 1967: 895 (rev.), 1970: 39 (cat.); Yang et al., 2006: 449 (cat.). Acknowledgments The preparation of this paper benefited from FAPESP grants 2008/58224–3 and 2013/01392-0. TABLE 1. Species of Dolichopodidae recorded from Colombia. Subfamily Genus Species Author Achalcinae Achalcus cyanocephalus Pollet, 2005 Diaphorinae Achradocera femoralis Becker, 1922 ...Continued on next page CATALOGUE OF DOLICHOPODIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 409 TABLE 1. (Continued) Subfamily Dolichopodinae Genus Species Author Chrysotus exceptus Becker, 1922 Lyroneurus fratellus Becker, 1922 luteoviridis Parent, 1935 occultus Becker, 1922 Symbolia lugubris Parent, 1934 Cheiromyia brevitarsis Brooks, 2010 Paraclius difficilis Becker, 1922 mixtus Parent, 1934 nigritibia Parent, 1934 umbripictus Becker, 1922 analis (Parent, 1954) giganteus (Brooks, 2002) potens (Parent, 1934) varus (Becker, 1922) Pelastoneurus Tachytrechus Medeterinae Medetera pedestris Becker, 1922 Sciapodinae Mesorhaga tristis Schiner, 1868 Condylostylus biseta Becker, 1922 cinctiventris Van Duzee, 1934 facetus Parent, 1933 gorgonensis Parent, 1933 mirus Parent, 1931 impatiens Becker, 1922 loriferus (Parent, 1934) nubeculus Becker, 1922 productipes Parent, 1934 squamifer Becker, 1922 Pseudosympycnus singularis (Parent, 1934) Stolidosoma permutans Becker, 1922 Sympycninae References Aldrich, J.M. (1893) New genera and species of Psilopinae. The Kansas University Quarterly, 2, 47–50. 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Öfversigt af Kongliga Vetenskaps-Akademiens Förhandlingar, 1, 106–110. 412 · Zootaxa 4122 (1) © 2016 Magnolia Press CAPELLARI Westwood, J.O. (1840) Order XIII. Diptera Aristotle (Antliata Fabricius. Halteriptera Clairv.). In: An introduction to the modern classification of insects. Synopsis of the genera of British insects. Longman, Orme, Brown, Green and Longmans, London, 158 pp. Yang, D., Zhu, Y.J., Wang, M. & Zhang, L.L. (2006) World Catalog of Dolichopodidae (Insecta: Diptera). China Agricultural University Press, Beijing, 704 pp. CATALOGUE OF DOLICHOPODIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 413 Zootaxa 4122 (1): 414–451 http://www.mapress.com/j/zt/ Copyright © 2016 Magnolia Press Article ISSN 1175-5326 (print edition) ZOOTAXA ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.4122.1.37 http://zoobank.org/urn:lsid:zoobank.org:pub:18CFF0A1-DF97-4D45-8C4F-8DDB1D669BB6 FAMILY PHORIDAE DANILO CÉSAR AMENT1* & BRIAN V. BROWN2 1 Universidade de São Paulo, Faculdade de Filosofia, Ciências e Letras de Ribeirão Preto, Departamento de Biologia, Av. Bandeirantes 3900, 14040–901, Ribeirão Preto, SP, Brasil. E-mail: danament@gmail.com 2 Natural History Museum of Los Angeles County, 900 Exposition Boulevard, Los Angeles, CA, 90007, USA. Email: bbrown@nhm.org *Corresponding author: danament@gmail.com. Abstract The Phoridae recorded from Colombia are catalogued, totaling 226 species distributed in 19 genera and three subfamilies. For each species, distribution, hierarchic position and cited references are presented. Key words: catalogue, Colombia, diversity, distribution, Phoridae Introduction Phoridae are probably one of the largest families of Diptera, numbering now only 4,000 described species, but estimated to probably include closer to 40–50,000 species. Because of their small size and astounding diversity, phorids are much less studied than almost any other group of flies. It is unknown how many species inhabit Colombia, but Brown (1998) estimated over 2,500 species from the much smaller landmass of Costa Rica. Many genera of Neotropical Region phorids have not yet been recorded from Colombia, but are known to occur there. A diagnosis of phorids is small (0.5–4 mm) cyclorrhaphan flies, ptilinal suture absent. Frons usually with distinctive setation, with one ventral and one dorsal pair of interfrontal setae, one ventral and one dorsal pair of fronto-orbital setae, one pair of inner and one pair of outer vertical setae, and one pair of postocellar setae; often one or more pairs of supra-antennal setae also present. Some or all frontal setae absent in some species. Mouthparts and palpus varied. Antenna has pedicel deeply inserted into first flagellomere; shape of first flagellomere varied, from small and rounded to greatly enlarged and flattened; Arista usually present and apical to dorsal. Thorax shortened, scutum arched ("humpbacked"), scutellum usually with one small anterior pair and one larger posterior pair of setae, but sometimes with many pairs of setae. Katepisternum produced into mesepimeron. Wing present, reduced, or absent. Wing venation highly characteristic, with thickened costal and radial veins, and thin, linear posterior veins without any crossveins linking them. Halter present, rarely absent. Hind femur of most phorids flattened, broad. At least hind tarsomere with one longitudinal setal palisade. Most phorid flies are parasitoids, many of which attack ants, bees, and other social insects; others are mycophagous, herbivorous, or predaceous. Some are scavengers, including the best-known species of the family. Larval life history was reviewed by Disney (1994) and Brown (2010). The life history of over 90% of the species is unknown. Phorids are greatly undersampled in Colombia, as in other parts of the Neotropical Region. The largest collections were amassed by the Colombia Arthropod Project, led by Michael Sharkey and Brian Brown in the 1990s. Malaise trap samples collected during this project are housed at the Instituto Alexander von Humboldt in Villa de Leyva. Other significant collections are those of Lee and Barreto (1969) at Rio Raposo (housed in the Smithsonian Institution, Washington DC), and Malaise trap samples taken at Amacayacu National Park by Bert Klein, which are housed in the University of Colombia, Bogotá entomology collection. This catalogue deals with the names of species of Phoridae recorded from Colombia and their synonyms. The records were obtained mostly with the help of the catalogue of the Phoridae of the world and its supplement 414 Accepted by S. Nihei: 17 Apr. 2015; published: 14 Jun. 2016 (Borgmeier 1968, 1971), searches on literature and the online version of the Zoological Records (http:// apps.webofknowledge.com). The 226 species recognized for Colombia are organized by alphabetical order within their higher categories: subgenus (when one exists), genus and subfamily. The subfamily classification follows Brown et al. (2015). At the end, a table is presented with a summary of the species, its higher categories, author and year of publication (Table 1). Acronyms used for the depositories CBFC—Colección Boliviana de Fauna, La Paz, Bolivia. CUMZ—University Museum of Zoology Insect Collection, Cambridge, United Kingdom. FMNH—Field Museum of Natural History, Chicago, USA. IAvH—Instituto Alexander von Humboldt, Villa de Leyva, Colombia. INBC—Instituto Nacional de Bioversidad, Heredia, Costa Rica. INPA—Instituto Nacional de Pesquisas da Amazônia, Manaus, Brazil. LACM—Entomology Section, Natural History Museum of Los Angeles County, Los Angeles, USA. MNHB—Museum der Naturkunde für Humboldt Universität zu Berlin, Germany. MNHN—Muséum National d'Histoire Naturelle, Paris, France. MPEG—Departamento de Entomologia, Museu Paraense Emilio Goeldi, Belém, Brazil. MUSM—Museo de Historia Natural, Universidad Nacional Mayor de San Marcos, Lima, Peru. MZSP—Museu de Zoolgia, Universidade de São Paulo, São Paulo, Brazil. SMTD—Staatsliches Museum für Tierkunde, Dresden, Germany. UNCB—Museo de Historia Natural, Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Santa Fé de Bogotá, Colombia. USNM—United States National Museum, Smithsonian Institution, Washington, DC, USA. ZMFK—Zoologisches Forschungsmuseum Alexander Koenig, Bonn, Germany. ZMUA—Zoologisch Museum, University of Amsterdam, Holland. List of abbreviations (and latin words used) biol.—Biology cat.—catalog cgto—corregimiento (little village) des.—designation distr.—distribution emend.—emendation gp.—group mon.—monotypy nec—and not (of ), nor (of ) nom. nudum—nomen nudum orig. des.—by original designation p. p.—in part passim—here and there preocc.—preoccupied refs.—references s. str.—sensu stricto spp.—species subg.—subgenus subs.—subsequent syn.—synonym AF—Afrotropical CATALOGUE OF PHORIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 415 HT—Holotype LT—Lectotype NT—Neotropical ST—Syntype F (f#)—Female M (m#)—Male Catalogue of Phoridae of Colombia Family Phoridae Latreille, 1796 Subfamily Chonocephalinae Brown et al. 2015 Genus Chonocephalus Wandolleck Chonocephalus Wandolleck, 1898: 428. Type species, Chonocephalus dorsalis Wandolleck, 1898 (mon.). Refs.: Becker, 1901: 86; Brues, 1906: 12; 1915b: 103 (key); 1915a: 144; Schmitz, 1927b: 26; 1928b: 87; 1929a: 168; Borgmeier, 1935b: 257; 1963a: 247 (key); 1967a: 201; 1968: 80 (cat.); 1971a: 179 (cat.); Paulian, 1958: 10; Beyer, 1964: 296; Disney, 1986: 77; 1994: 270; 2005: 393 (AF); 2008: 1 (NT). Heterophora Santos Abreu, 1921: 81 (nec Borgmeier). Refs.: Schmitz, 1928b: 87 (syn.). depressus Meijere, 1912: 151. Type locality: Indonesia, Sumatra, Medan de Bussy. LT F (ZMUA). Distr.: Bolivia (La Paz), Colombia (Valle del Cauca (Rio Raposo)), Hawaii (Lanikai Oahu), also recorded from the Oriental and Palaearctic regions. Refs.: Rao, 1961: 163; Borgmeier, 1968: 80 (cat.); Disney, 1991a: 208; 2002: 14 (LT des.); 2005: 14; 2008: 13. similis: Collin, 1912: 105 (part) nec Brues, 1905: 554. Refs.: Disney, 2002: 14 (misidentification) ecitophilus Borgmeier & Schmitz, 1923: 145. Refs.: Disney, 2002: 14 (syn.). japonicus Schmitz, 1941a: 82. Refs.: Disney, 2002: 15 (syn.). dimakae Paulian, 1958: 12. Refs.: Disney, 2005: 14 (syn.). pallidulus Beyer, 1964: 298. Refs.: Disney, 2002: 15 (syn.). simiolus Beyer, 1964: 300. Refs.: Disney, 2002: 15 (syn.). dominicanus Borgmeier, 1967a: 206. Type locality: Dominica, Clarke Hall. HT M (USNM). Distr.: Brazil (Amazonia), Colombia (Valle del Cauca (Rio Raposo), Cauca)), Costa Rica (Puerto Viejo), Dominica (Clarke Hall), Ecuador (Quevedo Pichilinguo, Galapagos Islands), Panama (Gamboa), Trinidad (Cumaca). Refs.: Borgmeier, 1968: 81 (cat.); Disney, 2008: 16. similis: Collin, 1912 (part): nec Brues, 1905: 554. punctifascia nec Borgmeier, 1935b: 257. wirthorum: Disney, 1980: 363. Refs.: Disney, 2002: 18 (syn.). globipygus Borgmeier, 1967a: 204. Type locality: Panama, Barro Colorado Island. HT M (USNM). Distr.: Colombia (Valle del Cauca (Rio Raposo)), Costa Rica (Limon), Ecuador (Rio Palenque), Panama (Barro Colorado Island, Gamboa). Refs.: Borgmeier, 1968: 81 (cat.); Disney, 2008: 21. wirthorum Disney, 1980: 363 (1 male; misidentification) heymonsi Stobbe, 1913: 131. Type locality: Surinam, Paramaribo. ST 5F (MNHB). Distr.: Argentina (Salta), Belize (Punta Gorda), Brazil (Amazonia), Colombia (Valle del Cauca (Rio Raposo)), Cuba (Guantanamo Bay Naval Base), Dominica, Ecuador (Napo, Galapagos Islands), Nicaragua (8 mi. S. La. Trinidad), Panama (Canal Zone), Puerto Rico, Surinam (Paramaribo), Trinidad (St Augustine) USA (Hawaii). Refs.: Borgmeier, 1968: 81 (cat.); Disney, 2002: 23; 2005: 24; 2008: 23. similis: Collin, 1912: 105, part (one female only) nec Brues, 1905 (misidentification) jamaicensis Brues, 1915b: 102. Refs.: Disney, 1981: 207 (syn.). 416 · Zootaxa 4122 (1) © 2016 Magnolia Press AMENT & BROWN punctifascia Borgmeier, 1935b: 257. Refs.: Prado, 1976: 587 (syn. C. jamaicensis). vadoni Paulian, 1958: 11. Refs.: Prado, 1976: 587 (syn. C. jamaicensis). brisbanensis Beyer, 1960: 85. Refs.: Disney, 2002: 23 (syn.). digitalis: Borgmeier, 1967a: 207 (part) (misidentification). leei Disney, 2008: 26. Type locality: Colombia, Rio Raposo. HT M (USNM). Distr.: Colombia (Valle del Cauca (Rio Raposo)). raposoensis Disney, 2008: 34. Type locality: Costa Rica, Puntarenas, Las Cruces Biological Station. HT M (CUMZ). Distr.: Colombia (Valle del Cauca (Rio Raposo)), Costa Rica (Puntarenas). vitiodepressus Disney, 2008: 38. Type locality: Costa Rica, Puntarenas, Las Cruces Biological Station. HT M (CUMZ). Distr.: Colombia (Chocó, Valle del Cauca), Costa Rica (Puntarenas), Mexico (Chiapas). Subfamily Metopininae Rondani, 1856 (emend.) Genus Apocephalus Coquillett Apocephalus Coquillett 1901: 501. Type species, Apocephalus pergandei Coquillett, 1901 (orig. des.). Refs.: Brues, 1904: 372; 1906: 8; 1915a: 110; 1924: 43 (key); Malloch, 1912: 442 (key); Schmitz, 1927a: 98; 1929a: 135; Borgmeier, 1925: 185 (key); 1937: 207 (subg.); 1958: 314, 316 (key); 1961: 38 (key); 1963a: 163 (key); 1968: 68 (cat.); 1971a: 183 (cat.); Beyer, 1965: 189; Brown, 1992: 81; 1993b: 218 (subg. Mesophora); 1994a: 1; 1997: 3 (attophilus gp.); 2000a: 5 (miricauda gp.); 2002: 6 (pergandei gp.); 2012: 20 (mucronatus gp.); Disney, 1994: 267. Pseudoplastophora Schmitz, 1915a: 327. Refs.: Borgmeier, 1968: 68 (syn.; cat.). Pleurophorina Borgmeier, 1969a: 66. Refs.: Brown, 1997: 3 (syn.). Zyziphora Peterson and Robinson, 1976: 119. Refs.: Brown, 1992: 81 (syn.). Anaclinusa Borgmeier, 1969a: 63. Refs.: Brown, 2000a: 5 (syn.). Borgmeieria Prado, 1976: 582. Refs.: Brown, 1997 (syn.). Macrocerides Borgmeier, 1927: 31 (p. p.). Refs.: Brown, 2011: 698. Subgenus Apocephalus s. str. Borgmeier, 1937 Apocephalus s. str. Borgmeier, 1937: 209. Type species, Apocephalus pergandei Coquillett, 1901 (orig. des.). asymmetricus Brown, 1997: 41. Type locality: Colombia, Rio Raposo. HT F (USNM). Distr.: Colombia (Valle del Cauca (Rio Raposo)), Ecuador (Sucumbios). barbarus Brown, 2002: 27. Type locality: Costa Rica, Heredia, La Selva Biological Station. HT F (INBC). Distr.: Colombia (Magdalena), Costa Rica (Guanacaste, Heredia). brochus Brown, 2000a: 20. Type locality: Colombia, Amazonas, 22 km NW Leticia. HT F (LACM). Distr.: Brazil (Pará), Colombia (Amazonas). catholicus Brown, 2000a: 48. Type locality: Costa Rica, Limon, 7 km SW Bribri. HT F (LACM). Distr.: Brazil (Amazonas), Colombia (Amazonas, Valle del Cauca), Costa Rica (Alajuela, Guanacaste, Heredia, Limon, Puntarenas), Ecuador (Esmeraldas, Napo, Pichincha), Panama (Darien). crassilatus Brown, 2000a: 12. Type locality: Costa Rica, Heredia, La Selva Biological Station. HT F (LACM). Distr.: Brazil (Minas Gerais), Colombia (Amazonas), Costa Rica (Alajuela, Guanacaste, Heredia), Panama (Canal Zone). densepilosus Borgmeier, 1971b: 110. Type locality: Brazil, Pará, Utinga. HT F (MZSP). Distr.: Brazil (Pará, Roraima), Colombia (Amazonas), Ecuador (Napo, Sucumbios), Peru (Madre de Dios). Refs.: Brown, 2000a: 36. CATALOGUE OF PHORIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 417 dinoponerae Brown, 2000a: 51. Type locality: Colombia, Amazonas, 22 km NW Leticia. HT F (UNCB). Distr.: Colombia (Amazonas). facettalis Borgmeier, 1961: 52. Type locality: Brazil, Rio de Janeiro, Taquara. HT F (MZSP). Distr.: Brazil (Amazonas, Rio de Janeiro), Colombia (Amazonas), Costa Rica (Guanacaste, Heredia, Limon, Puntarenas, San José), Ecuador (Guyas, Sucumbios), Panama (Canal Zone), Peru (Madre de Dios). Refs.: Borgmeier, 1968: 70 (cat.); Brown, 2002: 22. fernandezi Brown, 2002: 17. Type locality: Colombia, Amazonas, 22 km NW Leticia. HT F (UNCB). Distr.: Colombia (Amazonas). flexiseta Brown, 2002: 31. Type locality: Costa Rica, San José, Ciudad Colon. HT F (LACM). Distr.: Brazil (Amazonas), Colombia (Caquetá), Costa Rica (Heredia, Limon, Puntarenas, San José), Ecuador (Sucumbios), Panama (Canal Zone, San Blas). kungae Brown, 2000a: 27. Type locality: Colombia, Amazonas, Amacayacu National Park. HT F (LACM). Distr.: Colombia (Amazonas (Amacayacu, National Natural Park)). longimanus Brown, 2012: 20. Type locality: Colombia, Bolívar, SFF Colorados. HT F (IAVH). Distr.: Colombia (Bolívar). lopesi (Borgmeier), 1969a: 64 (Anaclinusa). Type locality: Brazil, Pará Belém. HT F (MZSP). Distr.: Brazil (Pará), Colombia (Amazonas), Costa Rica (Limón, Puntarenas), Guyana (Berbice). Refs.: Bogmeier, 1971b: 5; Brown, 2000a: 33. melinus Brown, 2000a: 9. Type locality: Ecuador, Napo, Yasuni Biological Research Station. HT F (LACM). Distr.: Colombia (Amazonas), Ecuador (Napo). neivai (Borgmeier), 1928: 167 (Macrocerides). Type locality: Brazil, Santa Catarina, Bom Retiro. HT F (MZSP). Distr.: Argentina (Santa Fe), Brazil (Santa Catarina), Colombia (Amazonas). Refs.: Borgmeier, 1931: 217 (Macrocerides); 1958: 341 (Macrocerides); 1968: 72 (cat., Macrocerides); 1969: 73; Brown, 1997: 11; Brown et al., 2010: 171. paraponerae Borgmeier, 1958: 324. Type locality: Panama, Barro Colorado Island. HT F (University of Kansas). Distr.: Brazil (Amazonas, Rio de Janeiro, Rondônia), Colombia (Amazonas, Rio Raposo), Costa Rica (Guanacaste, Heredia, Puntarenas), Ecuador (Napo, Pichincha, Sucumbios), Guyana (Berbice), Panama (Barro Colorado Island, Canal Zone, Darien, San Blas), Peru (Madre de Dios), Venezuela (Amazonas). Refs.: Borgmeier, 1968: 72 (cat.); Brown, 2000a: 6. reburrus Brown, 2002: 14. Type locality: Venezuela, Zulia. HT F (LACM). Distr.: Colombia (Magdalena), Venezuela (Zulia). roeschardae Brown, 2000a: 10. Type locality: Colombia, Amazonas, Amacayacu PNN. HT F (UNCB). Distr.: Brazil (Rio de Janeiro), Colombia (Amazonas (Amacayacu, National Natural Park)), Ecuador (Napo), Peru (Madre de Dios). scaurus Corona & Brown, 2004: 691. Type locality: Costa Rica, Heredia, La Selva Biological Station. HT F (INBC). Distr.: Colombia (Magdalena), Costa Rica (Heredia). setitarsus Brown, 1997: 44. Type locality: Costa Rica, San José, Braulio Carrillo National Park. HT F (LACM). Distr.: Brazil (Amazonas), Colombia (Valle del Cauca (Rio Raposo)), Costa Rica (Guanacaste, Limón, San José). Refs.: Brown et al., 2010: 178; Guillade & Folgarait, 2011: 34. 418 · Zootaxa 4122 (1) © 2016 Magnolia Press AMENT & BROWN spinilatus Brown, 1997: 50. Type locality: Costa Rica, Heredia, La Selva Biological Station. HT F (INBC). Distr.: Colombia (Valle del Cauca), Costa Rica (Heredia). tanyurus Brown, 2000a: 50. Type locality: Ecuador, Sucumbios, Sacha Lodge. HT F (LACM). Distr.: Colombia (Valle del Cauca), Ecuador (Sucumbios). tenuitarsus Brown, 1997: 13. Type locality: Costa Rica, Heredia, La Selva Biological Station. HT F (LACM). Distr.: Colombia (Valle del Cauca), Costa Rica (Alajuela, Heredia). torulus Brown, 2000a: 17. Type locality: Ecuador, Esmeraldas, Bilsa Biological Station. HT F (LACM). Distr.: Colombia (Valle del Cauca), Ecuador (Esmeraldas). trifidus Brown, 2000a: 49. Type locality: Peru, Madre de Dios, Zona Reserva Manu, Pakitza. HT F (MUSM). Distr.: Brazil (Pará, Roraima), Colombia (Amazonas), Ecuador (Napo, Sucumbios), Guyana (Berbice), Peru (Madre de Dios). Subgenus Mesophora Borgmeier, 1937 Apocephalus subg. Mesophora Borgmeier, 1937: 209. Type species, Apocephalus mortifer Borgmeier, 1937 (orig. des.). antennatus Malloch, 1913: 273. Type locality: USA, Maryland, Plummers Island. HT F (USNM). Distr.: Canada (Ontario, Quebec), Colombia, USA (Connecticut Delaware, Florida, Maryland, Michigan, Missouri, New York, North Carolina, Tennessee, Virginia, West Virginia). Refs.: Borgmeier, 1963a: 166; 1968: 74 (cat.); Brown, 1993b: 222; 1994b: 145. Genus Cryptophora Borgmeier Cryptophora Borgmeier, 1935c: 490. Type species, Cryptophora coeca Borgmeier, 1935c (mon.). Refs.: Borgmeier, 1968: 86 (cat.). colombiae Seevers, 1941: 186. Type locality: Colombia, Villavicencio. HT F (FMNH). Distr.: Colombia (Meta (Villavicencio). Refs.: Borgmeier, 1968: 86 (cat.). Genus Eibesfeldtphora Disney Eibesfeldtphora Disney et al., 2009: 601. Type species Apocephalus curvinervis Malloch, 1914 (orig. des.). Refs.: Brown et al., 2012: 34. Neodohrniphora Malloch, 1914 (p. p). Neodohrniphora subg. Eibesfeldtphora Disney, 1996: 1378. Refs.: Brown, 2001: 394 Neodohrniphora subg. Wallerphora Disney, 1996: 1378. Refs.: Brown, 2001: 394 (syn.). arcuata Brown, 2001: 395. Type locality: Colombia, Amazonas, Amacayacu PNN. HT F (UNCB). Distr.: Bolivia (Santa Cruz), Colombia (Amazonas (Amacayacu, National Natural Park)), Costa Rica (Puntarenas), Ecuador (Napo), Guyana (Berbice), Panama (San Blas), Peru (Madre de Dios). Neodohrniphora sp. ('small species'): Brown, 1999: 97. attae Disney, 1996: 1381. Type locality: Trinidad, Simla Valley. HT F (ZMFK). Distr.: Brazil (Roraima), Colombia (Cauca), Costa Rica (Guanacaste, Heredia, Limon, Puntarenas, San José), Guyana (Berbice), Panama (San Blas), Peru (Madre de Dios), Trinidad (Simla Valley). Refs.: Brown, 2001: 395. Apocephalus sp.: Beyer, ln Eibl-Eibesfeldt & Eibl-Eibesfeldt, 1967: 280. Neodohrniphora sp. 1: Disney, 1994: 80. CATALOGUE OF PHORIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 419 declinata Borgmeier, 1925: 214. Type locality: Brazil, Rio de Janeiro, Petrópolis. HT F (MZSP). Distr.: Brazil (Minas Gerais, Rio de Janeiro, São Paulo), Colombia (Amazonas), Guyana (Berbice). Refs.: Borgmeier, 1928: 120; 1929: 496; 1968: 228 (cat., Neodohrniphora); Prado, 1976: 593; Brown, 2001: 399. curvinervis: Borgmeier, 1962a: 479 (p. p.). Refs.: Borgmeier, 1968: 228 (p. p., cat.). dissita Brown, 2001: 401. Type locality: Colombia, Amazonas, Amacayacu PNN. HT F (UNCB). Distr.: Bolivia (Santa Cruz), Colombia (Amazonas (Amacayacu, National Natural Park)), Ecuador (Napo). Neodorhniphora sp. ('large species'): Brown, 1999: 97. leei Brown, 2001: 404. Type locality: Colombia, Rio Raposo. HT F (USNM). Distr.: Colombia (Valle del Cauca (Rio Raposo)). pala Brown, 2001: 404. Type locality: Colombia, Amazonas, Amacayacu PNN. HT F (UNCB). Distr.: Colombia (Amazonas (Amacayacu, National Natural Park)). prolixa Brown, 2001: 404. Type locality: Ecuador, Napo, Tena. HT F (LACM). Distr.: Colombia (Amazonas), Ecuador (Napo, Sucumbios), Panama (San Blas). Genus Homalophora Borgmeier Homalophora Borgmeier, 1923b: 849. Type species, Homalophora reichenspergeri Borgmeier, 1923b (mon.). Refs.: Borgmeier, 1923c: 177; 1958: 401 (key); 1960: 341; 1968: 101 (cat.); Schmitz, 1927b: 27; 1928b: 70; 1929a: 167; Brues, 1932b: 141; Disney, 1994: 276. Neoplatyphora Borgmeier, 1923d: 1224 (nom. nudum); 1958: 401. guatiquiae Seevers, 1941: 188. Type locality: Colombia, Villavicencio. HT F (FMNH). Distr.: Colombia (Meta (Villavicencio)). Refs.: Borgmeier, 1968: 101 (cat.). metae Seevers, 1941: 189. Type locality: Colombia, Villavicencio. HT F (FMNH). Distr.: Colombia (Meta (Villavicencio)). Refs.: Borgmeier, 1968: 101 (cat.). Genus Idiophora Borgmeier Idiophora Borgmeier, 1960: 365. Type species, Ecitomyia termitoxena Seevers, 1941 (mon.). Refs.: Borgmeier, 1968: 102 (cat.); Disney, 1994: 276. termitoxena (Seevers), 1941: 186 (Ecitomyia). Type locality: Colombia, Villavicencio. HT F (FMNH). Distr.: Colombia (Meta (Villavicencio)). Refs.: Borgmeier, 1960: 366; 1968: 102 (cat.). Genus Megaselia Rondani Megaselia Rondani, 1856: 137. Type species, Megaselia crassineura Rondani, 1856 = Phora costalis Roser, 1840 (orig. des.). Refs.: Borgmeier, 1925: 137 (Aphiochaeta); 1961: 71; 1962a: 291 (key, NT spp.); 1964: 258 (key subg. Aphiochaeta, USA); 1966: 2 (key Megaselia s. str., USA); 1967b: 200 (key oriental Aphiochaeta); 1967c: 83 (key oriental Megaselia s. str.); 1968: 104 (cat.); 1971: 8 (key); Schmitz, 1927a: 111; 1929a: 140; 1956: 387; Malloch, 1935: 336 (key, Samoa); Brues, 1950: 59; Beyer, 1958: 38 (keys); 1959b: 66; 1959c: 3 (key, Africa); 1960: 23 (key, Australia); 1964: 273 (key, Hawaii); 1966: 190; 1967: 335 (key, Micronesia); Ferrar, 1987: 1; Disney, 1989: 1139; Brown, 1992: 73. Aphiochaeta Brues, 1904: 337. Refs. Brues, 1906: 9; 1915a: 111; Malloch, 1910: 91; 1912: 445 (key); Schmitz passim before 1926; Lundbeck, 1922: 201; Borgmeier, 1925: 137; Schmitz: 1927a: 111 (subg.). Plastophora Brues, 1905: 551. Megaselida Becker et al., 1907: 175 (emend.; unjustified). Pericyclocera Schmitz, 1927b: 24. Refs.: Disney, 1994: 279 (syn.). Paraphiochaeta Malloch, 1914: 27. Refs.: Schmitz, 1929a: 141. 420 · Zootaxa 4122 (1) © 2016 Magnolia Press AMENT & BROWN graminicola (Borgmeier), 1969a: 102 (Pericyclocera). Type locality: Brazil, Pará, Belém. HT F (USNM). Distr.: Brazil (Pará), Colombia (Leticia). Refs.: Borgmeier, 1971a: 206 (Pericyclocera). neocorynurae Gonzalez et al., 2002: 74. Type locality: Colombia, Boyacá, SFF Iguaque. HT M (UNCB). Distr.: Colombia (Boyacá (Iguaque, Santuario de Fauna y Flora). parianae (Borgmeier), 1969a: 105 (Pericyclocera). Type locality: Brazil, Pará, Belém. HT F (USNM). Distr.: Brazil (Pará), Colombia (Leticia), Panama (Colón). Refs.: Borgmeier, 1971a: 206 (Pericyclocera). perdita (Malloch), 1912: 459 (Aphiochaeta). Type locality: USA, Plummers Island. HT F (USNM). Distr.: Brazil, Colombia, Ecuador, Mexico, USA. Refs.: Brues, 1915a: 126 (Aphiocaeta); Brues, 1950: 68; Borgmeier, 1962a: 293; 1964: 262; 1968: 199 (cat.). picta (Lehman), 1822: 43 (Phora) Type locality: Europe (type lost). Distr.: Europe, North and South America, Philippines, Japan. Refs.: Mik, 1896: 112 (Phora); Aldrich, 1896: 435 (Phora); Becker, 1901: 52 (Phora); Wood, 1908: 169 (Phora); Brues, 1904: 361 (Aphiochaeta); 1915a: 126 (Aphiochaeta); Strobl, 1910: 121 (Aphiochaeta); Malloch, 1912: 451 (Aphiochaeta); Schmitz, 1918: 136 (Aphiochaeta); 1929a: 20, 23, 187; 1941b: 34 (distr.); Lundbeck, 1922: 233 (Aphiochaeta); Brues, 1950: 70; Borgmeier, 1962a: 305 (key); 1966: 10; 1967c: 84 (key); 1968: 146 (cat.). ininterrupta Zetterstedt, 1838: 797 (Trineura). Refs.: Zetterstedt, 1848: 2878; 1860: 6478; Schiner, 1864: 337 (Phora); Coquillett, 1895: 104. subquadrifasciata Zetterstedt, 1852: 4826 (Trineura). Refs.: Schmitz, 1929a: 20; 1953a: 136 (syn.). atlantica Brues, 1904: 362 (Aphiochaeta). Refs.: Brues, 1950: 73; Borgmeier, 1961: 110; 1962a: 298 (syn.). picta var. fasciiventris Brues, 1919: 432 (Paraphiochaeta) (nec Enderlein). Refs.: Borgmeier, 1962a: 299 (syn.). calianensis Brues, 1936: 419. Refs.: Borgmeier, 1962a: 483 (syn.). equivocata Brues, 1936: 420. Refs.: Borgmeier, 1967c: 97 (syn.). infuscata Bridarolli, 1940: 487. Refs.: Borgmeier, 1962a: 296 (syn.). rufipes (Meigen), 1804: 313 (Trineura). Type locality: Germany, Aachen. HT F (probably in the MNHN). Distr.: Cosmopolitan. Refs.: Fabricius, 1805: 336 (Trineura); Zetterstedt, 1838: 795 (Trineura); 1848: 2857 (Trineura); 1852: 4823 (Trineura); Meigen, 1830: 216 (Phora); Macquart, 1835: 627 (Phora); Schiner, 1864: 340 (Phora); Strobl, 1892: 202 (Phora); Becker, 1901: 59 (Phora); Coquillett, 1904: 51 (Phora); Wood, 1909: 194, 243 (Phora); Keilin, 1911: 61 (Phora); Brues, 1904: 368 (Aphiochaeta); 1915a: 130 (Aphiochaeta); Bezzi, 1911: 60 (Aphiochaeta); Malloch, 1912: 494 (Aphiochaeta); Silva Figueroa, 1916: 9 (Aphiochaeta); Lundbeck, 1922: 375 (Aphiochaeta); Borgmeier, 1925: 167 (Aphiochaeta); 1966: 72; 1968: 152 (cat.); Schmitz, 1929a: passim; 1931: 48; 1938a: 31; 1941b: 34 (distr.); Taylor, 1944: 120; Hennig, 1952: 144; Hesjedal, 1984: 113. annulata Meigen, 1804: 314 (Trineura). Refs.: Meigen, 1830: 216 (Trineura); Macquart, 1835: 627 (Trineura); Schmitz, 1929a: 15 (syn.). pallipes Latreille, 1809: 395 (Phora). Refs.: Dufour, 1840: 54 (Phora); Schmitz, 1929a: 3 (syn.). vulgaris Fallén, 1823: 6 (Trineura). Refs.: Schmitz, 1929a: 4 (syn.). semiflava Hartig, 1837 (Phora). Refs.: Engel, 1916: 57 (Phora); Schmitz, 1929a: 20 (syn.). heracleellae Bouché, 1834: 102 (Phora). Refs.: Strobl, 1892: 202 (Phora); Schmitz, 1929a: 7 (syn.). sphingicides Bouché, 1834 (Phora). Refs.: Schmitz, 1929a: 7 (syn.). coleopterorum Brischke, 1869: 4 (Phora). Refs.: Schmitz, 1929a: 25 (syn.). omnivora Hudson, 1892 (Phora). Refs.: Schmitz, 1929a: 199 (Phora). juanfernandezica Enderlein, 1938: 661 (Lioyella). Refs.: Borgmeier, 1967d: 404 (syn.). scalaris (Loew), 1866: 53 (Phora). Type locality: Cuba. LT F (MCZC). Distr.: Cosmopolitan. Refs.: Brues, 1904: 364; 1915a: 132; 1928a: 160 (biol.); 1950: 70; Malloch, 1912: 467 (Aphiochaeta); Schmitz, 1929a: 29; 1956: 394; Fernandez, 1949: 93 (biol.); Bohart, 1947: 406; Bohart & Gressit, 1951 passim; Beyer, 1964: 287; 1967: 347; Ondraschek, 1953: 347 (biol.); Hennig, 1952: 143; Haider, 1956: 191; Borgmeier, 1962a: 307 (key); 1966: 3 (key); CATALOGUE OF PHORIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 421 1967b: 84, 89 (key), 101; 1968: 153 (cat.); Mainx, 1964: 415; El-Miniawi & Moustafa, 1965: 81; McCrae, 1967: 241; Robinson, 1975: 384; Tumrasvin et al., 1977: 74 (biol.); Prawirodisastro & Benjamin, 1979: 317; Benner & Ostermeyer, 1980: 103; Dias Macieira et al., 1983: 223; Garris, 1983: 688; Curtis et al., 1987: 265; 1989: 47; Carareto & Mourao, 1988: 995; Singh & Rana, 1989: 228; Traut & Willhoeft, 1990: 407; Traut et al., 1990: 77; Willhoeft & Traut, 1990: 237; Benner, 1991: 351; Curtis & Benner, 1991: 85; Wolf et al., 1993: 272; Andreotti et al., 2003: 46; Koller et al., 2003: 1; Solis et al., 2005: 339 (biol.); Sukontason et al., 2005: 113; Brown & Oliver, 2007: 85; Rasmussen & Noor, 2009: 1; Sorensen et al., 2009: 73; Miranda-Miranda, 2011: 344; Vanin et al., 2013: 209. xanthina Speiser, 1908: 148 (Aphiochaeta). Refs.: Schmitz, 1929a: 29 (syn.). fissa Becker, 1908: 193 (Aphiochaeta). Refs.: Schmitz, 1929a: 29 (syn.). conjuncta Becker, 1908: 193 (Aphiochaeta). Refs.: Schmitz, 1929a: 29 (syn.). banksi Brues, 1909: 6 (Aphiochaeta). Refs. Brues, 1936: 380; Borgmeier, 1961: 110 (syn.). circumsetosa Speiser, 1911: 348 (Aphiochaeta). Refs.: Schmitz, 1929a: 29 (syn.). ferruginea Brunetti, 1912: 507 (Aphiochaeta). Refs.: Schmitz, 1929a: 29 (syn.). repicta Schmitz, 1915b: 108 (Aphiochaeta). Refs.: Schmitz, 1929a: 29 (syn.). plusiivorax Enderlein, 1929: 104 (Lioyella). Refs.: Beyer, 1967: 347 (syn.). Genus Melaloncha Brues Melaloncha Brues, 1904: 374. Type species, Melaloncha pulchella Brues, 1904 (subs. des.). Refs.: Brues, 1906: 8; 1912: 136; 1915a: 109; Malloch, 1912: 500; Schmitz, 1927b: 19, 26; 1927a: 147; 1929a: 144; Borgmeier, 1925: 223; 1934: 167; 1938: 44; 1959: 167; 1968: 219 (cat.); 1971b: 125; Brown, 2004a: 386 (cingulata-gp.); 2004b: 1 (Udamochiras subg.); 2005: 241 (furcata-gp.); 2006: 1; Gonzalez & Brown, 2004: 1; Brown & Kung, 2006: 1 (ungulata-gp.); Kung, 2008: 491; Brown & Smith, 2010: 649; Lucia et al., 2013: 72 (biol.); Lutz & Brown, 2013: 238 (biol.). Melanoloncha (emend.) Becker et al., 1907: 176. (unjustified emendation) Udamochiras Enderlein, 1912b: 42; 1924: 281. Refs.: Brues 1912: 136 (syn.). Subgenus Melaloncha s. str. Brown, 2004b Melaloncha s. str. Brown, 2004b: 10. Type species, Melaloncha pulchella Brues, (orig. des.). acicula Brown, 2006: 44. Type locality: Colombia, Amazonas, PNN Amacayacu. HT F (UNCB). Distr.: Brazil (Amazonas), Colombia (Amazonas (Amacayacu, National Natural Park)), Ecuador (Napo). acoma Brown & Kung, 2006: 4. Type locality: Costa Rica (Heredia, La Selva Biological Station. HT F (LACM). Distr.: Argentina (Misiones), Bolivia (Beni, La Paz), Colombia (Chocó, Magdalena), Costa Rica (Heredia, Puntarenas, San José), Mexico (Vera Cruz), Peru (Madre de Dios, San Martin, San Antonio de Cumbaza). caligula Brown & Kung, 2006: 12. Type locality: Colombia, Amazonas, Amacayacu PNN. HT F (UNCB). Distr.: Colombia (Amazonas (Amacayacu, National Natural Park)). candida Brown & Kung, 2006: 6. Type locality: Brazil, Pará, Caiçara. HT F (MPEG). Distr.: Brazil (Amazonas, Pará), Colombia (Amazonas). clandestina Brown, 2005: 255. Type locality: Colombia, Amazonas, Amacayacu PNN. HT F (UNCB). Distr.: Colombia (Amazonas (Amacayacu, National Natural Park)). debilis Brown & Kung, 2006: 11. Type locality: Costa Rica (Heredia (La Selva Biological Station)). HT F (INBC). Distr.: Colombia (Chocó), Costa Rica (Heredia). diastata Brown & Kung, 2006: 9. Type locality: Colombia, Amazonas, Amacayacu PNN. HT F (UNCB). Distr.: Colombia (Amazonas (National Natural Park)). 422 · Zootaxa 4122 (1) © 2016 Magnolia Press AMENT & BROWN digitalis Borgmeier, 1959: 176. Type locality: Brazil, Santa Catarina, Nova Teutônia. HT M (MZSP). Distr.: Brazil (Rio de Janeiro, Rondônia, Santa Catarina), Colombia (Amazonas, Caquetá), Costa Rica (Alajuela, Guanacaste, Heredia, Puntarenas, San José), Mexico (Tamaulipas), Panama (Fort Gulick), Peru (Madre de Dios, San Martin). Refs.: Borgmeier, 1968: 220 (cat.); Brown, 2006: 17. hirsuta Brown, 2006: 54. Type locality: Colombia, Cundinamarca, Arbelaez. HT F (UNCB). Distr.: Colombia (Cundinamarca). hyalinipennis Borgmeier, 1933: 95. Type locality: Argentina, San José de Chiquitos. HT M (Staatliches Museum für Naturkunde, Stuttgart). Distr.: Argentina (San José de Chiquitos), Brazil (Amazonas, Bahia, Mato Grosso, São Paulo), Colombia (Amazonas), Ecuador (Napo), Peru (Madre de Dios, San Martin). Refs.: Borgmeier, 1968: 220 (cat.); Brown, 2006: 56. juxta Brown & Kung, 2006: 10. Type locality: Colombia, Chocó, PNN Utría. HT F (UNCB). Distr.: Colombia (Chocó (Utria, National, Natural Park)), Costa Rica (Puntarenas (5.5 km SW Rincon)). kungae Brown, 2005: 246. Type locality: Ecuador, Napo, Yasuni National Park. HT F (LACM). Distr.: Bolivia (La Paz), Brazil (Amazonas, Rondonia), Colombia (Amazonas, Caquetá, Vichada), Ecuador (Napo), Peru (Madre de Dios, San Martin). lamellata Borgmeier, 1934: 169. Type locality: Bolivia, Mapiri, Sarampioni. HT F (SMTD). Distr.: Bolivia (Mapiri), Colombia (Magdalena). Refs.: Borgmeier, 1968: 220 (cat.); Brown, 2004a: 388. maculata Borgmeier, 1934: 176. Type locality: Costa Rica, Higuito, San Mateo. HT F (USNM). Distr.: Bolivia (La Paz), Brazil (Bahia), Colombia (Cauca), Costa Rica (Alajuela, Heredia, Higuito, Limón, San José), Nicaragua (Rio San Juan). Refs.: Borgmeier, 1968: 221 (cat.); Brown, 2006: 58. nannocauda Brown, 2006: 59. Type locality: Brazil, Amazonas, Reserva Campina. HT F (INPA). Distr.: Brazil (Amazonas), Colombia (Amazonas (Reserva Campina)). punctifrons Borgmeier, 1934: 171. Type locality: Bolivia, Mapiri, Lorenzapata. HT F (Dresden). Distr.: Bolivia (Mapiri), Brazil (Pará), Colombia (Amazonas, Caquetá, Valle del Cauca), Costa Rica (Alajuela, Guanacaste, Heredia), Panama (Darien). Refs.: Borgmeier, 1968: 221 (cat.); Borgmeier & Prado, 1975: 54; Brown, 2006: 16. rodeoensis Brown, 2006: 33. Type locality: Costa Rica, San José, Zona Protectora El Rodeo. HT F (LACM). Distr.: Colombia (Amazonas), Costa Rica (San José). striatula Borgmeier, 1934: 176. Type locality: Costa Rica, Higuito, San Mateo. HT F (USNM). Distr.: Brazil (Amazonas), Colombia (Chocó), Costa Rica (Alajuela, Cartago, Guanacaste, Heredia, Higuito, Puntarenas), Panama (San Blas) Peru (Madre de Dios, San Martin). Refs.: Borgmeier, 1968: 222 (cat.); Brown, 2006: 63. stylata (Schiner), 1868: 224 (Phora). Type locality: Venezuela. HT F (Naturhistorisches Museum Wien). Distr.: Colombia, Venezuela. Refs.: Brues, 1904: 375; Schmitz, 1927a: 176; 1928b: 12; 1929a: 147; Borgmeier, 1934: 170; 1959: 179; 1968: 222 (cat.); Brown, 2006: 42. tuparroensis Brown & Kung, 2006: 14. Type locality: Colombia, PNN El Tuparro. HT F (UNCB). Distr.: Colombia (Vichada (El Tuparro, National Natural Park)). Subgenus Udamochiras Brown, 2004b Melaloncha subg. Udamochiras Brown, 2004b: 11. Type species: U. colossia Enderlein, 1924: 281 (orig. des.). Udamochiras Enderlein, 1912b: 42. Type species: U. colossia Enderlein, 1924: 281 (orig. des.). CATALOGUE OF PHORIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 423 ancistra Gonzalez & Brown, 2004: 8. Type locality: Colombia, Boyacá, SFF Iguaque, Cabaña Chaina. HT F (UNCB). Distr.: Colombia (Boyacá (Iguaque, Santuario de Fauna y Flora)). flavilata Brown, 2004b: 14. Type locality: Colombia, Amazonas, Amacayacu PNN HT F (UNCB). Distr.: Colombia (Amazonas (Amacayacu, National Natural Park)). Genus Melittophora Brues Melittophora Brues, 1928b: 134. Type species, Melittophora salti Brues, 1928b (mon.). Refs.: Schmitz, 1929a: 200; Borgmeier, 1968: 279 (cat.); Brown, 1993a: 121. salti Brues, 1928b: 136. Type locality: Colombia, Magdalena, Rio Frio. HT F (not found). Distr.: Colombia (Magdalena (Rio Frio)). Refs.: Borgmeier, 1968: 279 (cat.). Genus Pseudacteon Coquillett Pseudacteon Coquillett, 1907: 208. Type species, Pseudacteon crawfordi Coquillett, 1907 (orig. des.). Refs.: Wood, 1908: 168; Schmitz, 1916: 30; 1918: 124; 1924: 139; 1929a: 138; 1938b: 171; Wasmann, 1918: 317; Lundbeck, 1922: 425; Steyskal, 1944: 1 (key); Borgmeier, 1923e: 239 (biol.); 1925: 235 (key NT); 1962b: 27; 1963a: 198; 1968: 246 (cat.); 1971: 210 (cat.); Disney 1991b: 283; Brown, 1992: 84; Plowes et al., 2009: 937. Plastophora (p. p.): Brues, 1907b: 430; 1915a: 137; Malloch, 1912: 500; Schmitz, 1914: 528, 557. andinus Plowes et al., 2009: 941. Type locality: Peru, Madre de Dios, Manu NP. HT F (LACM). Distr.: Bolivia (Beni), Colombia (Amazonas), Ecuador (Napo, Sucumbios), Peru (Madre de Dios). antiguensis (Malloch), 1912: 502 (Plastophora). Type locality: Antigua. HT M (USNM). Distr.: Antigua, Barbados (Holetown), Colombia (Vichada (El Tuparro, National Natural Park)), Dominica (Clark Hall), Guyana (Berbice), Suriname (Brokopondo), Trinidad (Arima Vly). Refs.: Borgmeier, 1963a: 200; 1968: 246 (cat.); Borgmeier & Prado, 1975: 76; Disney, 1991b: 287; Plowes et al., 2009: 941. caudalis Borgmeier, 1963a: 200. Refs.: Disney, 1991b: 287, 332 (misidentification). grandis Borgmeier, 1963a: 200. (misidentification). arcuatus Borgmeier, 1969a: 42. Type locality: Dominica, Clarke Hall. HT F (USNM). Distr.: Colombia (Magdalena, Vichada), Costa Rica (Heredia), Dominica (Clarke Hall), Ecuador (Sucumbios), Nicaragua (12 km N Matagalpa), Suriname (Brokopondo). Refs.: Borgmeier, 1968: 246 (cat.); Feener and Brown, 1992: 80; Plowes et al., 2009: 954. browni Disney, 1991b: 291. Type locality: Costa Rica, Heredia, La Selva Biological Station. HT F (CUMZ). Distr.: Colombia (Cauca); Costa Rica (Alajuela, Guanacaste, Heredia, Limon, Puntarenas, San Jose), Guatemala (Sacatepéquez, Suchitepéquez), Honduras (Catacamas), Nicaragua (Rio San Juan), Grenada (Volcan Mombacho), Panama (Darien, Veraguas). Refs.: Feener and Brown, 1992: 80; Plowes et al., 2009: 942. deltoides Plowes et al., 2009: 952. Type locality: Suriname, Brokopondo, Brownsberg Nature Park. HT F (LACM). Distr.: Colombia (Magdalena, Vichada), Ecuador (Napo, Sucumbios), Guyana (Berbice), Suriname (Brokopondo). grandis Greene, 1941: 183 Type locality: Jamaica, Negril Point, Station 583. HT F (USNM). Distr.: Bahamas (San Salvador Island), British Virgin Islands (Guana Island), Colombia (Bolívar, Magdalena), Dominican Rep. (La Cumbre), Jamaica (Milk River Bank, Negril Point, Runaway Bay), Puerto Rico (Salinas), Venezuela (Zuila). Refs.: Disney, 1991b: 296; Plowes et al., 2009: 944. antiguensis: Borgmeier, 1963a: 200 (misidentification); 1968: 246 (cat., misidentification). 424 · Zootaxa 4122 (1) © 2016 Magnolia Press AMENT & BROWN hippeus Plowes et al., 2009: 949. Type locality: USA, Texas, San Roque Riv.. HT F (LACM). Distr.: Colombia (Meta), Costa Rica (Guanacaste, Limon, Puntarenas, San José), Ecuador (Napo, Sucumbios), Guatemala (Sacatepéquez), Honduras (Catacamas), Trinidad (Arima Valley), USA (Texas). crawfordi: Feener 1987: 148 (misidentification); Morrison et al., 1999: 198 (misidentification) quinni Plowes et al., 2009: 953. Type locality: Costa Rica, Guanacaste, Santa Rosa NP. HT F (LACM). Distr.: Colombia (Magdalena, Vichada), Costa Rica (Guanacaste, Heredia), Mexico (Jalisco). Genus Pseudohypocera Malloch Pseudohypocera Malloch, 1912: 439. Type species, Pseudohypocera clypeata Malloch = kerteszi (Enderlein), 1912b (mon.). Refs.: Borgmeier, 1925: 183; 1968: 249 (cat.); 1971: 210 (cat.); Schmitz, 1927a: 97; 1929a: 134; Disney, 1994: 288. kerteszi (Enderlein), 1912b: 48 (Aphiochaeta). Type locality: Costa Rica. HT F (destroyed). Distr.: Brazil (Bahia, Pernambuco, Rio de Janeiro, São Paulo), Colombia (Sucre (Sincé)), Costa Rica, Mexico (Jalisco), Paraguay (Pastoreo). Refs.: Brues, 1915a: 122 (Aphiochaeta); Borgmeier, 1935a: 438; 1968: 249 (cat.); 1971: 210 (cat.); Prado, 1976: 598. clypeata Malloch, 1912: 439. Refs.: Brues, 1915a: 110. nigrofascipes Borgmeier & Schmitz, 1923: 132. Refs.: Borgmeier, 1925: 183; 1930: 92; Brues, 1928b: 137; Pickel, 1928: 67 (biol.). minor Schmitz, 1927a: 97. Refs.: Borgmeier, 1930: 97 (syn.). Pseudohypocera sp.: Nogueira-Neto, 1970: 297. Genus Syntermophora Seevers Syntermophora Seevers, 1941: 183. Type species, Syntermophora microphthalma Seevers, 1941 (mon.). Refs.: Borgmeier, 1968: 259 (cat.); Disney, 1994: 291. microphthalma Seevers, 1941: 184. Type locality: Colombia (Villavicencio). HT F (FMNH). Distr.: Colombia (Meta (Villavicencio). Refs.: Borgmeier, 1968: 259 (cat.). Genus Tayrona Brown & Kung Tayrona Brown & Kung, 2004. Type species, Tayrona nitifrons Brown & Kung, 2004 (mon.). nitifrons Brown & Kung, 2004: 6. Type locality: Colombia, Magdalena, PNN Tayrona, Zaino. HT M (UNCB). Distr.: Colombia (Magdalena (Tayrona, National Natural Park)). Subfamily Phorinae Rondani, 1856 Genus Chaetocnemistoptera Borgmeier Chaetocnemistoptera Borgmeier, 1923f: 51. Type species, Hypocera semifurcata Borgmeier, 1923b (mon.). Refs.: Schmitz, 1927a: 32; 1929a: 94; Brown, 1992a: 52; Ament, 2014: 301. Chaetopleurophora (p. p.): Borgmeier, 1961: 4; 1968: 14 (cat.), 1969a: 36. palpalis (Borgmeier), 1961: 8 (Chaetopleurophora). Type locality: Brazil, Santa Catarina, Nova Teutônia. HT M (MZSP). Distr.: Brazil (Rio de Janeiro, Rondônia, Santa Catarina), Colombia (Amazonas, Chocó, Magdalena), Costa Rica (Puntarenas), Ecuador (Napo, Sucumbios), Panama (Canal Zone). Refs.: Borgmeier, 1968: 13 (Chaetopleurophora, cat.); Brown, 1992a: 52; Ament, 2014: 308. CATALOGUE OF PHORIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 425 lobata Ament, 2014: 311. Type locality: Bolivia, La Paz, 40 km N Caranavi. HT M (CBFC). Distr.: Brazil (Amazonas), Bolivia (La Paz), Colombia (Amazonas, Vaupés, Vichada), Ecuador (Napo, Sucumbios), French Guiana (Regina), Peru (Madre de Dios). magdalena Ament, 2014: 312. Type locality: Colombia, Magdalena, PNN Tayrona. HT M (IAVH). Distr.: Colombia (Bolivar, Magdalena (Tayrona, Nacional, Natural Park)), Costa Rica (Puntarenas), Panama (Darien Cruce de Mono). phaeonota Ament, 2014: 314. Type locality: Colombia, Huila, PNN Cueva de los Guácharos. HT M (LACM). Distr.: Colombia (Huila (Cueva de Los Guácharos, National Natural Park)), Ecuador (Sucumbios). pityropyga Ament, 2014: 315. Type locality: Argentina, Misiones, Reserva Vida Silvestre Urugua-í. HT M (LACM). Distr.: Argentina (Misiones), Brazil (Paraná, Santa Catarina), Colombia (Amazonas), Ecuador (Sucumbios). trichopoda Ament, 2014: 317. Type locality: Colombia, Chocó, PNN Utria, Boroboro. HT M (IAVH). Distr.: Colombia (Amazonas, Chocó (Utria, Boroboro, National Natural Park), Cundinamarca, Huila, Valle del Cauca), Costa Rica (Puntarenas), Ecuador (Esmeraldas), Panama (Parque Nacional Campana), Venezuela (Aragua Rancho Grande Stn). Genus Coniceromyia Borgmeier Coniceromyia Borgmeier, 1923a: 338. Type species, Coniceromyia epicantha Borgmeier, 1923a (mon.). Refs.: Borgmeier, 1925: 123; 1950: 283; 1963b: 450 (key); 1968: 19 (cat.); 1971: 179 (cat.); Brown, 1992: 53; Disney, 1994: 271; Kung & Brown, 2000: 1; Ament & Amorim, 2010: 36. browni Kung, 2009: 50. Type locality: Colombia, Amazonas, PNN Amacayacu, Matamata. HT M (IAVH). Distr.: Colombia (Amazonas (Amacayacu, National Natural Park)). caliga Kung & Brown, 2001: 267. Type locality: Colombia, Nariño, RN La Planada, Vía Hondon. HT M (UNCB). Distr.: Colombia (Nariño (La Planada, Reserva Natural)). reina Kung & Brown, 2001: 268. Type locality: Colombia, Boyacá, SFF Iguaque. HT M (UNCB). Distr.: Colombia (Boyacá (Iguaque, National Natural Park), Cundinamarca). sakaii Kung, 2009: 53. Type locality: Colombia, Vichada, PNN El Tuparro. HT M (IAvH). Distr.: Colombia (Vichada (El Tuparro, National Natural Park)). valdesi Kung, 2009: 54. Type locality: Colombia, Putumayo, PNN La Paya. HT M (IAvH). Distr.: Colombia (Putumayo (La Paya, National Natural Park)). Genus Cyrtophorina Borgmeier & Prado Cyrtophorina Borgmeier & Prado, 1975: 85. Type species, Cyrtophorina deinocerca Borgmeier & Prado, 1975 (orig. des.). Refs.: Brown, 1993a: 119; 2007: 118; Disney, 1994: 272. deinocerca Borgmeier & Prado, 1975: 85. Type locality: Brazil, Rondônia, Vilhena. HT M (MZSP). Distr.: Brazil (Pará, Rondônia), Colombia (Amazonas), Ecuador (Napo), Pastaza. Refs.: Brown, 1993a: 120; 2007: 120. gorgonensis Brown, 2007: 126. Type locality: Colombia, Cauca, Isla Gorgona. HT M (UNCB). Distr.: Colombia (Cauca (Isla Gorgona). 426 · Zootaxa 4122 (1) © 2016 Magnolia Press AMENT & BROWN Genus Dohrniphora Dahl Dohrniphora Dahl, 1898: 188. Type species, Dohrniphora dohrni Dahl, 1898 (mon.). Refs.: Dahl, 1899: 65; Becker, 1901: 83; Schmitz, 1918: 107; 1920: 97; , Borgmeier, 1925: 88 (key); 1960: 258 (key); 1961: 10; 1963a: 63 (key); 1966: 134; 1968: 29 (cat.); 1969a: 53; 1971a: 10 (cat.); Beyer, 1959a: 4 (Diplonevra, key); Borgmeier & Prado, 1975: 11; Prado, 1976: 566; Disney, 1990: 339 (key); 1994: 274; 2003: 1 (AF); Brown, 1992: 60; 2008: 204; Kung & Brown, 2005: 55; 2006: 1931; Brown & Kung, 2007: 157; 2010: 1; Hash et al., 2011: 13; 2013: 401 (phylogeny). Diploneura subg. Dohrniphora: Schmitz, 1929a: 107; 1949: 204, 241. Pronomiorpha Enderlein, 1912a: 116. Crepidopachys Enderlein, 1912b: 16. acanthiventris Brown & Kung, 2010: 34. Type locality: French Guiana, L’Oyapock, St. Georges de l’Oyapock. HT M (LACM). Distr.: Colombia (Amazonas), French Guiana (L’Oyapock). acarinata Brown & Kung, 2010: 61. Type locality: Colombia, Amazonas, Amacayacu NP. HT M (IAVH). Distr.: Colombia, Amazonas (Amacayacu National Park)). acuta Brown & Kung, 2010: 82. Type locality: Colombia, Valle del Cauca, PNN Farallones de Cali. HT M (IAvH). Distr.: Colombia (Risaralda, Valle del Cauca (Farallones de Cali, National Natural Park)). alvarengai Prado, 1976: 567. Type locality: Brazil, Mato Grosso. HT M (MZSP). Distr.: Brazil (Amazonas, Mato Grosso, Rondônia, Roraima), Colombia (Amazonas, Caquetá, Putumayo), Costa Rica (Alajuela, Guanacaste, Puntarenas, San José), Ecuador (Napo), French Guiana (Regina), Panama (Chiriquí), Peru (Madre de Dios, Cuzco). Refs.: Brown & Kung 2007: 161. amacayavia Brown & Kung, 2010: 64. Type locality: Colombia, Amazonas, Amacayacu NP. HT M (IAvH). Distr.: Brazil (Amazonas), Colombia (Amazonas (Amacayacu, National Natural Park)), Costa Rica (San José). anchicayensis Brown & Kung, 2007: 162. Type locality: Colombia, Valle del Cauca, PNN Farallones de Cali. HT M (UNCB). Distr.: Colombia (Valle del Cauca (Farallones de Cali, National Natural Park)). Refs.: Brown & Kung, 2010: 55. anteroventralis Borgmeier, 1960: 264. Type locality: Brazil, Santa Catarina, Nova Teutônia. HT M (MZSP). Distr.: Argentina (Misiones), Brazil (Amazonas, Pará, Paraná, Rio de Janeiro, Santa Catarina), Colombia (Amazonas, Bolívar, Caquetá, Cauca, Chocó, Magdalena, Putumayo, Vaupés, Vichada), Costa Rica (Alajuela, Guanacaste, Heredia, Limón, Puntarenas), Ecuador (Chico, Manabi, Napo, Pichincha), French Guiana (Regina), Honduras (Olancho), Nicaragua (Rio San Juan), Panama (Canal Zone, Darien), Peru (Madre de Dios, Pakitza), Trinidad (Arima Valley, Morne Blue), Venezuela (Zulia). Refs.: Borgmeier, 1968: 30 (cat.); Borgmeier & Prado, 1975: 14; Disney, 1983: 455; Brown & Kung, 2007: 163. anterosetalis Borgmeier & Prado, 1975: 14. Refs.: Brown & Kung, 2007: 163 (syn.). trudiae Disney, 1983: 455. Refs.: Brown & Kung, 2007: 163 (syn.). aprosmicta Brown & Kung, 2010: 35. Type locality: Colombia, Amazonas, Amacayacu NP. HT M (IAvH). Distr.: Colombia (Amazonas (Amacayacu, National Natural Park)). atrilinea Brown & Kung, 2010: 56. Type locality: Colombia, Amazonas, Amacayacu NP. HT M (IAvH). Distr.: Bolivia (La Paz), Colombia (Amazonas (Amacayacu, National Natural Park)), Ecuador (Napo), Peru (Madre de Dios). basilacuna Brown & Kung, 2010: 10. Type locality: Colombia, Caquetá, PNN Chiribiquete. HT M (IAvH). Distr.: Colombia (Caquetá (Chiribiquete National Natural Park)). berezovskiyi Brown & Kung, 2010: 11. Type locality: Colombia, Amazonas, Amacayacu NP. HT M (IAvH). CATALOGUE OF PHORIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 427 Distr.: Brazil (Amazonas, Rondônia), Bolivia (La Paz), Colombia (Amazonas (Amacayacu, National Natural Park), Caquetá, HuilaHuila, Vaupés)), French Guiana (Regina). bicavatura Brown & Kung, 2010: 108. Type locality: Costa Rica, Heredia, La Selva Biological Station. HT M (LACM). Distr.: Brazil (Rio de Janeiro), Colombia (Amazonas, Putumayo), Costa Rica (Guanacaste, Heredia, Puntarenas), French Guiana (Regina), Panama (Canal Zone). bicostula Kung & Brown, 2005: 57. Type locality: Colombia, Amazonas, PNN Amacayacu. HT M (UNCB). Distr.: Brazil (Amazonas, Rondonia, Roraima), Colombia (Amazonas (Amacayacu, National Natural Park)), Peru (Cuzco). Refs.: Brown, 2008: 205. biseriata Borgmeier, 1960: 284. Type locality: Brazil, Rio de Janeiro, Petrópolis. HT M (MZSP). Distr.: Argentina (Misiones), Brazil (Amazonas, Rio de Janeiro), Colombia (Amazonas), Ecuador (Pichincha). Refs.: Borgmeier, 1968: 30 (cat.); Borgmeier & Prado, 1975: 24; Brown & Kung, 2010: 85. brevisimplex Brown & Kung, 2010: 41. Type locality: Colombia, Amazonas, Amacayacu NP. HT M (IAvH). Distr.: Colombia (Amazonas (Amacayacu, National Natural Park), Caquetá)). broadheadi Disney, 1983: 453. Type locality: Panama, Canal Zone, Pipeline Road. HT M (CUMZ). Distr.: Colombia (Amazonas, Magdalena), Panama (Canal Zone). Refs.: Brown & Kung, 2007: 189. bugenitalis Brown & Kung, 2010: 28. Type locality: Colombia, Amazonas, Amacayacu NP. HT M (IAvH). Distr.: Colombia (Amazonas (Amacayacu, National Natural Park)). buscki Malloch, 1912: 432 (Dohrniphora venusta, var. buscki). Type locality: Panama, (La) Cabima. HT M (USNM). Distr.: Bolivia (La Paz), Brazil (Amazonas), Colombia (Amazonas, Chocó, Putumayo, Valle del Cauca, Vichada), Costa Rica (Guanacaste, Heredia, Limón, Puntarenas), French Guiana (Regina), Panama (Canal Zone, (La) Cabima), Peru (Madre de Dios). Refs.: Borgmeier, 1961: 110; 1968: 31 (cat.); Borgmeier & Prado, 1975: 16; Brown & Kung, 2010: 22. laminata Borgmeier, 1960: 274. Refs.: Borgmeier, 1961: 110 (syn.). caguanensis Brown & Kung, 2010: 74. Type locality: Colombia, Caquetá, San Vicente de Caguán. HT M (IAvH). Distr.: Colombia (Caquetá (San Vicente del Caguan)). circularis Brown & Kung, 2007: 190. Type locality: Colombia, Amazonas, PNN Amacayacu. HT M (UNCB). Distr.: Colombia (Amazonas (Amacayacu, National Natural Park)). circumflexa Borgmeier, 1960: 285. Type locality: Brazil, Santa Catarina, Nova Teutônia. HT M (MZSP). Distr.: Brazil (Santa Catarina), Colombia (Amazonas, Cauca), Costa Rica (Cartago, Heredia, Puntarenas, San José), Panama (Canal Zone). Refs.: Borgmeier, 1968: 31 (cat.); Brown & Kung, 2010: 66. clariloba Brown & Kung 2007: 168. Type locality: Colombia, Amazonas, PNN Amacayacu. HT M (UNCB). Distr.: Colombia (Amazonas (Amacayacu, National Natural Park)), Peru (Cuzco). cocaensis Brown & Kung, 2007: 190. Type locality: Colombia, Amazonas, PNN Amacayacu. HT M (UNCB). Distr.: Brazil (Rondônia), Colombia (Amazonas (Amacayacu, National Natural Park)), Ecuador (Napo). cognata Borgmeier, 1960: 280. Type locality: Brazil, Santa Catarina, Nova Teutônia. HT M (MZSP). Distr.: Brazil (Amazonas, Roraima, Santa Catarina), Colombia (Amazonas, Chocó, Huila, Meta, Putumayo, Valle del Cauca, Vichada), Costa Rica (Heredia, Limón), French Guiana (l’Oyapock, Regina), Panama (Canal Zone). Refs.: Borgmeier, 1968: 31 (cat.); Brown & Kung, 2010: 103. ventralis Borgmeier & Prado, 1975: 26. Refs.: Brown & Kung, 2007: 186; Brown & Kung, 2010: 103 (syn.). 428 · Zootaxa 4122 (1) © 2016 Magnolia Press AMENT & BROWN consimilis Brown & Kung 2007: 168. Type locality: Costa Rica, Heredia, Chilamate. HT M (LACM). Distr.: Colombia (Valle del Cauca), Costa Rica (Cartago, Guanacaste, Heredia, Puntarenas), Peru (Cuzco). conspicua Brown & Kung, 2010: 58. Type locality: Ecuador, Pichincha 17 km E Santo Domingo. HT M (LACM). Distr.: Colombia (Chocó, Magdalena, Valle del Cauca), Costa Rica (Heredia), Ecuador (Pichincha), Panama (Colon). mississippiensis: Brown & Kung, 2007: 177 (misidentification). continua Brown & Kung, 2010: 13. Type locality: Costa Rica, Heredia, La Selva Biological Station. HT M (LACM). Distr.: Bolivia (La Paz), Brazil (Amazonas), Colombia (Amazonas, Caquetá, Meta, Putumayo, Valle del Cauca, Vaupés), Costa Rica (Heredia), French Guiana (Regina), Peru (Madre de Dios). cootei Brown & Kung 2007: 169. Type locality: Colombia, Amazonas, Leticia, Cgto. La Pedrera. HT M (UNCB). Distr.: Brazil (Amazonas), Colombia (Amazonas (Leticia, Corregimiento La Pedrera), Vaupés), Costa Rica (Puntarenas). cornuta (Bigot), 1857: 348 (French edition: 1857: 827) (Phora). Type locality: Cuba. ST 1M, 1 F (Paris Museum). Distr.: Cosmopolitan (some NT records: Argentina (Buenos Aires, La Rioja, Mendoza), Colombia (Cauca), Costa Rica (San José), Dominica (Anse Bouleau), Dominican Republic (Pedernales), Jamaica (Runaway Bay), Puerto Rico (Rio Grande). Refs.: Schmitz, 1926: 44 (Diploneura); 1927a: 49 (Diploneura); 1929b: 22 (Diploneura); 1929a: 112 (Diploneura); 1951: 241 (Diploneura); Santos Abreu & Schmitz, 1934: 65 (Diploneura); Brues, 1936: 369 (Diploneura); Frey, 1936: 70 (Diploneura); de Mejeire, 1938: 230 (Diploneura); Bohart, 1947: 407 (Diploneura); Hennig, 1952: 151 (Diploneura); Beyer, 1959a: 4 (Diploneura); 1964: 272 (Diploneura); Borgmeier, 1960: 277; 1963a: 65; 1968: 31 (cat.); Kaneko et al., 1961: 241 (Diploneura); Kloter et al., 1977: 775; Barnes, 1990: 474 (biol.); Kung & Brown, 2006: 1932; Feng & Liu, 2012: 1528. navigans Frauenfeld, 1867: 154 (Phora). Refs.: Schmitz, 1929a: 24 (Phora). cleghorni Bigot, 1890: 191 (Phora). Refs.: Collin, 1912: 106 (Phora). chlorogastra Becker, 1901: 32 (Phora). mordax Brues, 1911: 531 (Phora). venusta: Malloch, 1912: 432. bequaerti Schmitz, 1914: 105. flaviventris Silva Figueroa, 1916: 15 (Apocephalus). Refs.: Borgmeier, 1957: 24 (syn.; Apocephalus). divaricata var. basalis Santos Abreu, 1921: 11. divaricata var. obscura Santos Abreu, 1921: 11. fulva Santos Abreu, 1921: 12. Refs.: Bridarolli, 1939: 5 (Diplonevra); Schmitz, 1951: 245; Borgmeier, 1968: 32 (syn.; cat.). opposita Borgmeier, 1925: 107. Refs.: Borgmeier, 1960: 277. crockeri Van Duzee, 1933: 70. Refs.: Curran, 1934: 152 (syn.); Borgmeier, 1960: 277 (syn.). willowsi Van Duzee, 1933: 70; Borgmeier, 1960: 277 cavifemur Borgmeier, 1969b: 1 (p. p., male only), misidentification cunarensis Brown & Kung, 2010: 114. Type locality: Colombia, Caquetá, PNN Chiribiquete. HT M (IAVH). Distr.: Colombia (Caquetá (Chiribiquete, National Natural Park). curticerca Brown & Kung, 2007: 191. Type locality: Colombia, Amazonas, PNN Amacayacu. HT M (UNCB). Distr.: Colombia (Amazonas (Amacayacu ,National Natural Park)). decrescens Brown & Kung, 2007: 169. Type locality: Peru, Madre de Dios, Tambopata Research Center. HT M (MUSM). Distr.: Brazil (Amazonas), Colombia (Amazonas, Vaupés), Ecuador (Napo), Peru (Madre de Dios). didyma Brown & Kung, 2007: 192. Type locality: Ecuador, Sucumbios, Sacha Lodge. HT M (LACM). Distr.: Colombia (Amazonas), Ecuador (Sucumbios). CATALOGUE OF PHORIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 429 digitata Brown & Kung, 2007: 192. Type locality: Colombia, Amazonas, PNN Amacayacu. HT M (UNCB). Distr.: Colombia (Amazonas (Amacayacu, National Natural Park)). dispar (Enderlein), 1912b: 29 (Phora). Type locality: Brazil, Santa Catarina, Hammonia. LT M (Polish Academy of Sciences). Distr.: Argentina, Bolivia, Brazil, Colombia, Costa Rica, Dominica, Dominican Republic, Ecuador, French Guiana, Jamaica, Montserrat, Panama, Peru, Puerto Rico. Refs.: Schmitz, 1923: 55; Borgmeier, 1935a: 431 (Diploneura, LT design.); 1960: 276; 1968: 32 (cat.); 1969b: 2; Kung & Brown, 2006: 1934. dispah: Borgmeier, 1925: 109 (lapsus). divaricata (Aldrich), 1896: 437 (Phora). Type locality: Saint Vincent. HT M (USNM). Distr.: Argentina, Brazil, Colombia, Costa Rica, Dominica (Cabrit Swamp, Layou), Dominican Republic (Hato Mayor, La Estrelleta, Pedernales), French Guiana, Jamaica (Hardwar Gap, Runaway Bay), Nevis (Newcastle), Panama, Puerto Rico (El Yunque, Maricao, Rio Grande), Saint Kitts (Greenhill, Wingfield Mtn.), Saint Lucia (Anse La Raye), Saint Vincent, USA. Refs.: Borgmeier, 1961: 111; 1968: 33 (cat.); Barnes, 1991: 307; Kung & Brown, 2006: 1936; Brown & Kung, 2007: 171; 2010: 91. venusta: Brues, 1915a: 95 (misidentification). cornuta: Schmitz, 1929: 22, 25, 32 (misidentification). obscuriventris Borgmeier, 1925: 105. Refs.: Disney & Kistner, 1997: 24 (syn.). diffusa Borgmeier, 1960: 281. Refs.: Borgmeier, 1961: 111 (syn.). recurvata Borgmeier, 1960: 272. Refs.: Brown & Kung, 2010: 91 (syn.). perplexa: Borgmeier, 1963a (p. p., some males, misidentification). cavifemur Borgmeier, 1969b: 1 (p. p., female only). Refs.: Kung & Brown, 2006: 1936 (syn.). caverna Disney, 1995: 421. Refs.: Brown & Kung, 2010: 91 (syn.). ecitophila Borgmeier, 1960: 271. Type locality: Brazil, Goiás, Campinas. HT F (MZSP). Distr.: Argentina (Misiones), Bolivia (La Paz), Brazil (Amazonas, Minas Gerais, Rio de Janeiro), Colombia (Amazonas, Bolívar, Chocó, Magdalena, Meta, Nariño, Valle del Cauca, Vaupés), Costa Rica (Guanacaste, Heredia, Limón, Puntarenas, San José), Ecuador (Napo, Pichincha), French Guiana (Regina), Mexico (Jalisco), Panama (Canal Zone), Peru (Madre de Dios), Venezuela (Aragua). Refs.: Borgmeier, 1968: 33 (cat.); Borgmeier & Prado, 1975: 36; Brown & Kung, 2010: 23. geminata Borgmeier, 1960: 283. Refs.: Borgmeier, 1968: 33 (syn.). emmesta Brown & Kung, 2007. Type locality: Colombia, Amazonas, PNN Amacayacu. HT M (UNCB). Distr.: Colombia (Amazonas (Amacayacu, National Natural Park)). femoralis Borgmeier, 1960: 265. Type locality: Panama, Darien, Patino Point. HT M (USNM). Distr.: Colombia (Magdalena), Panama (Canal Zone, Darien). Refs.: Borgmeier, 1968: 33 (cat.); Brown & Kung, 2007: 193. fernandezi Brown & Kung, 2010: 107. Type locality: Colombia, Amazonas, Amacayacu NP. HT M (IAVH). Distr.: Colombia (Amazonas (Amacayacu, National Natural Park)). flammidorsa Brown & Kung, 2010: 30. Type locality: Peru, Madre de Dios, Pakitza. HT M (MUSM). Distr.: Colombia (Caquetá), Peru (Madre de Dios). flexicarina Brown & Kung, 2010: 85. Type locality: Colombia, Amazonas, Amacayacu NP. HT M (IAVH). Distr.: Colombia (Amazonas (Amacayacu, National Natural Park), Putumayo)), Argentina (Misiones). foveolata Borgmeier, 1960: 278. Type locality: Brazil, Rio de Janeiro, Petrópolis. HT M (MZSP). Distr.: Brazil (Amazonas, Rio de Janeiro), Colombia (Amazonas), Ecuador (Napo). Refs.: Borgmeier, 1968: 33 (cat.); Borgmeier & Prado, 1975: 29; Brown & Kung, 2010: 77. fuliginata Brown & Kung, 2010: 43. Type locality: El Salvador, La Libertad, Finca la Giralda. HT M (LACM). 430 · Zootaxa 4122 (1) © 2016 Magnolia Press AMENT & BROWN Distr.: Colombia, Costa Rica (Cartago, Heredia, Limón, Puntarenas), El Salvador (La Libertad), Mexico (Oaxaca, Veracruz). castaneicoxa: Borgmeier, 1963a: 66 (p. p., specimens not from Brazil, misidentification). fulleri Brown & Kung, 2010: 37. Type locality: Colombia, Caquetá, PNN Chiribiquete. HT M (IAvH). Distr.: Brazil (Amazonas), Colombia (Caquetá (Chiribiquete, National Natural Park)). fundula Brown & Kung, 2010: 37. Type locality: Colombia, Amazonas, Amacayacu NP. HT M (IAVH). Distr.: Colombia (Amazonas (Amacayacu, National Natural Park)), Ecuador (Napo). fuscicoxa Borgmeier, 1923a: 327. Type locality: Brazil, Rio de Janeiro, Petrópolis. HT M (MZSP). Distr.: Argentina (Buenos Aires), Brazil (Rio de Janeiro, Santa Catarina), Colombia (Bolívar), Ecuador (Napo), Peru (Cuzco). Refs.: Borgmeier, 1968: 33 (cat.); Borgmeier & Prado, 1975: 24; Brown & Kung, 2010: 110. gigantea (Enderlein), 1912b: 18 (Phora). Type locality: Brazil, Santa Catarina, (Lüderwaldt). HT M (Polish Academy of Sciences, Warsaw, Poland). Distr.: Bolivia (La Paz (40 km N Caranavi, Chulumani)), Brazil (Rio de Janeiro, Santa Catarina, São Paulo), Colombia (Chocó, Huila, Nariño, Norte de Santander, Risaralda, Valle del Cauca), Costa Rica (Alajuela, Cartago, Guanacaste, Limón, Puntarenas, San José), Ecuador (Napo, Pichincha), Honduras (Olancho), Nicaragua (Matagalpa), Panama (Darien, San Blas), Peru (Madre de Dios, San Martin), Venezuela (Aragua). Refs.: Schmitz, 1923: 54; Borgmeier, 1925: 94; 1960: 261; 1968: 33 (cat.); Enderlein, 1924: 273 (Diploneura); Schmitz 1929: 32, 108 (Diploneura); 1949: 204 (Diploneura); Brown & Kung, 2007: 173. rubriventris Borgmeier, 1923b: 580. Refs.: Borgmeier, 1925: 95; 1960: 261 (syn.). gravis Borgmeier & Prado, 1975: 32. Type locality: Brazil (Pará (Belém)). HT M (MZSP). Distr.: Brazil (Pará), Colombia (Amazonas). Refs.: Brown & Kung, 2010: 48. hamartia Brown & Kung, 2007: 194. Type locality: Colombia, Amazonas, Leticia. HT M (CNCI). Distr.: Colombia (Amazonas (Leticia), Peru (Upper Rio Pachitea). pyricornis: Brues, 1944: 151 (Diploneura (Dohrniphora)); Borgmeier, 1961: 13 (p. p., paratype only). heliconiae Brown & Kung, 2010: 68. Type locality: Costa Rica, Alajuela, Albergue de Heliconia. HT M (LACM). Distr.: Costa Rica (Alajuela, Cartago, Heredia), Colombia (Chocó), Mexico (Jalisco). hexaseta Brown & Kung, 2010: 17. Type locality: Colombia, Magdalena, PNN Tayrona. HT M (IAVH). Distr.: Colombia (Cauca, Magdalena (Tayrona, National Natural Park)), Mexico (Jalisco). ignobilis Borgmeier, 1960: 279. Type locality: Brazil, Rio de Janeiro, Petrópolis. HT M (MZSP). Distr.: Brazil (Minas Gerais, Rio de Janeiro), Colombia (Amazonas, Vichada), French Guiana (L’Oyapock), West Indies (Grenada). Refs.: Borgmeier, 1968: 34 (cat.); Brown & Kung, 2010: 68. incomitata Brown & Kung, 2007: 194. Type locality: Colombia, Amazonas, PNN Amacayacu. HT M (UNCB). Distr.: Colombia (Amazonas (Amacayacu, National Natural Park)). inornata Brown & Kung, 2007: 195. Type locality: Colombia, Amazonas, PNN Amacayacu. HT M (UNCB). Distr.: Colombia (Amazonas (Amacayacu, National Natural Park), Caquetá, Putumayo). knabi Malloch, 1912: 431. Type locality: Panama, Cabima. HT M (USNM). Distr.: Brazil (Rondonia), Colombia (Amazonas), Costa Rica (Heredia, Limón, Puntarenas, San José), Guyana (Berbice), Panama (Cabima, Canal Zone). Refs.: Borgmeier, 1968: 34 (cat.); Borgmeier & Prado, 1975: 17; Brown & Kung, 2010: 86. lacunosa Brown & Kung, 2007: 195. Type locality: Panama, Canal Zone, Barro Colorado Is. HT M (LACM). Distr.: Colombia (Amazonas, Caquetá, Chocó, Magdalena), Guyana (Berbice), Panama (Canal Zone, Darien). CATALOGUE OF PHORIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 431 lamellifera Borgmeier, 1961: 16. Type locality: Peru, Iquitos. HT M (USNM). Distr.: Colombia (Amazonas), Peru (Iquitos). Refs.: Borgmeier, 1968: 34 (cat.); Borgmeier & Prado, 1975: 24; Brown & Kung, 2010: 79. lobata Borgmeier, 1960: 269. Type locality: Suriname, Mazaruni River, Geijskes. HT M (MZSP). Distr.: Brazil (Belem), Colombia (Amazonas), Suriname (Mazaruni River). Refs.: Borgmeier, 1968: 34 (cat.); Brown & Kung, 2007: 196. longinoi Brown & Kung, 2010: 92. Type locality: Costa Rica, Heredia, La Selva Biological Station. HT M (LACM). Distr.: Brazil (Amazonas), Colombia (Amazonas), Costa Rica (Heredia). longisetosa Brown & Kung, 2010: 74. Type locality: Brazil, Amazonas, Manaus. HT M (INPA). Distr.: Brazil (Amazonas), Colombia (Caquetá). lugens Borgmeier, 1960: 286. Type locality: Brazil, Santa Catarina, Nova Teutônia. HT M (MZSP). Distr.: Argentina (Misiones), Brazil (Santa Catarina), Colombia (Amazonas), Costa Rica (Puntarenas, San José). Refs.: Borgmeier, 1968: 34 (cat.); Brown & Kung, 2010: 93. luteicincta Borgmeier, 1960: 282. Type locality: Brazil, Rio de Janeiro. HT M (only a hind leg glued to a card mount, MZSP). Distr.: Brazil (Amazonas, Santa Catarina, Rio de Janeiro), Colombia (Amazonas, Vaupés), Ecuador (Napo), Peru (Madre de Dios). Refs.: Borgmeier, 1968: 34 (cat.); Brown & Kung, 2010: 86. matamatae Brown & Kung, 2010: 48. Type locality: Colombia (Amazonas (Amacayacu NP)). HT M (IAVH). Distr.: Brazil (Rondônia), Colombia (Amazonas). membranea Brown & Kung, 2007: 177. Type locality: Costa Rica, San José, Zurquí de Moravia. HT M (LACM). Distr.: Bolivia (La Paz), Colombia (Nariño, Risaralda), Costa Rica (San José), Venezuela (Lara). mesofemoralis Brown & Kung, 2007: 196. Type locality: Panama, Darien, Cruce de Mono. HT M (LACM). Distr.: Colombia (Chocó, Magdalena), Panama (Canal Zone, Darien). minerva Borgmeier, 1961: 17. Type locality: Panama, Canal Zone. HT M (USNM). Distr.: Colombia (Chocó), Costa Rica (Heredia, Puntarenas), Panama (Canal Zone). Refs.: Borgmeier, 1968: 35 (cat.); Brown & Kung, 2010: 94. minervoidea Brown & Kung, 2010: 117. Type locality: Colombia, Vaupés, Estación Biologica Mosiro-Itajura. HT M (IAvH). Distr.: Colombia (Amazonas, Vaupés). minilobata Brown & Kung, 2010: 117. Type locality: Colombia, Amazonas, Amacayacu NP. HT M (IAVH). Distr.: Colombia (Amazonas (Amacayacu, National Natural Park)), Ecuador (Napo). mississippiensis Khalaf, 1971: 274. Type locality: USA, Gainesville (Mississippi). HT M (USNM). Distr.: Colombia, Costa Rica, USA. Refs.: Brown & Kung, 2007: 177. mixtura Brown & Kung, 2010: 99. Type locality: Colombia, Amazonas, 22 km NW Leticia. HT M (IAvH). Distr.: Brazil (Amazonas, Rondonia), Colombia (Amazonas, Vichada), Ecuador (Napo), French Guiana (l’Oyapock). shannoni: Brown & Kung, 2007: 182 (misidentification). nigricarina Brown & Kung, 2010: 56. Type locality: Costa Rica, Heredia, La Selva Biological Station. HT M (LACM). Distr.: Costa Rica (Alajuela, Cartago, Guanacaste, Heredia, Limón, Puntarenas, San José), Colombia (Chocó, Magdalena, Valle del Cauca), Mexico (Sinaloa), Nicaragua (Zelaya), Panama (Canal Zone). notoplicata Brown & Kung, 2010: 120. Type locality: Colombia, Amazonas, Amacayacu NP. HT M (IAvH). Distr.: Brazil (Amazonas), Colombia (Amazonas (Amacayacu, National Natural Park)). 432 · Zootaxa 4122 (1) © 2016 Magnolia Press AMENT & BROWN oricilla Kung and Brown, 2005: 60. Type locality: Honduras, Olancho, Catacamas. HT M (LACM). Distr.: Colombia (Amazonas, Bolívar, Chocó, Magdalena, Valle del Cauca), Costa Rica (Alajuela, Guanacaste, Heredia, San José), Honduras, Mexico (Chiapas), Panama (Canal Zone, Darien). Refs.: Brown, 2008: 205. orthogramma Brown & Kung, 2010: 25. Type locality: Colombia, Valle del Cauca, PNN Farallones de Cali. HT M (IAvH). Distr.: Colombia (Chocó, Valle del Cauca (Farallones de Cali, National Natural Park)). ospinae Brown & Kung, 2010: 70. Type locality: Colombia, Caquetá, PNN Chiribiquete. HT M (IAVH). Distr.: Colombia (Caquetá (Chiribiquete, National Natural Park)). ovibarba Brown & Kung, 2007: 197. Type locality: Ecuador, Napo, Yasuni NP. HT M (LACM). Distr.: Colombia (Vichada), Ecuador (Napo). palenquensis Brown & Kung, 2010: 110. Type locality: Ecuador, Pichincha, 47 km S of Santo Domingo. HT M (LACM). Distr.: Colombia (Chocó, Magdalena), Ecuador (Pichincha). pallidens Borgmeier & Prado, 1975: 22. Type locality: Costa Rica, Higuito, San Mateo. HT M (MZSP). Distr.: Colombia (Meta), Costa Rica (Heredia, Higuito, Limón, Puntarenas). Refs.: Brown & Kung, 2010: 87. paraguayana Brues, 1907a: 400 (Phora). Type locality: Paraguay, Asunción. ST 2F (Hungarian National Museum, probably destroyed during Hungarian Revolt). Distr.: Argentina (Misiones), Bolivia (S. Inicua R. Alto Beni), Brazil (Amapá, Amazonas, Espirito Santo, Maranhão, Mato Grosso, Minas Gerais, Rio de Janeiro, Rondônia, Roraima, São Paulo), Colombia (Amazonas, Bolívar, Chocó, Vichada), Costa Rica (Alajuela, Cartago, Guanacaste, Heredia, Limón, Puntarenas, San José), Ecuador (Napo, Pastaza), French Guiana (Regina), Guyana (Berbice), Panama (Canal Zone), Paraguay (Asunción, Villarica), Peru (Cuzco, Madre de Dios), Venezuela (Aragua). Refs.: Brues, 1915a: 96; Schmitz, 1929a: 108 (Diplonevra); Borgmeier, 1960: 269; 1968: 36 (cat.); Brown & Kung, 2007: 178. conspicua Borgmeier, 1923b: 628. Refs.: Borgmeier, 1925: 99; 1960: 269 (syn.); Schmitz, 1929a: 108 (Diplonevra); Brues, 1932a: 134 (Diploneura). anterodorsalis (Schmitz), 1928a: 28 (Diploneura). Refs.: Schmitz, 1929a: 108 (Diploneura); Borgmeier, 1960: 269 (syn.). paralobata Brown & Kung 2007: 198. Type locality: Ecuador, Sucumbios, Sacha Lodge. HT M (LACM). Distr.: Colombia (Amazonas, Magdalena), Ecuador (Pastaza, Sucumbios). parvicarinata Brown & Kung, 2010: 49. Type locality: Colombia, Amazonas, PNN Amacayacu. HT M (IAvH). Distr.: Colombia (Amazonas (Amacayacu, National Natural Park)). parvicava Brown & Kung, 2010: 79. Type locality: Colombia, Amazonas, Amacayacu NP. HT M (IAvH). Distr.: Colombia (Amazonas (Amacayacu, National Natural Park)). parvipala Brown & Kung, 2010: 111. Type locality: Costa Rica, Puntarenas, 24 km W of Pan American Hwy. HT M (LACM). Distr.: Colombia (Amazonas, Chocó), Costa Rica (Heredia, Limón, Puntarenas). phyzela Brown & Kung, 2010: 44. Type locality: Colombia, Amazonas, Amacayacu NP. HT M (IAvH). Distr.: Colombia (Amazonas (Amacayacu, National Natural Park)), Trinidad (Arima Valley). porrasae Brown & Kung, 2010: 51. Type locality: Costa Rica, Heredia, La Selva Biological Station. HT M (LACM). Distr.: Colombia (Chocó, Magdalena, Valle del Cauca, Vichada), Costa Rica (Heredia, Puntarenas, San José), Ecuador (Pichincha), Panama (Canal Zone). preacuta Brown & Kung, 2010: 107. Type locality: Colombia, Valle del Cauca, PNN Farallones de Cali. HT M (IAVH). Distr.: Colombia (Valle del Cauca Farallones de Cali, National Natural Park)). CATALOGUE OF PHORIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 433 procera Borgmeier, 1960: 273. Type locality: Brazil, Santa Catarina, Blumenau. HT M (MZSP). Distr.: Brazil (Roraima, Santa Catarina), Colombia (Amazonas, Cauca, Valle del Cauca), Costa Rica (Heredia, Puntarenas), Ecuador (Napo), French Guiana (Regina), Guyana (Berbice), Panama (Canal Zone). Refs.: Borgmeier, 1968: 36 (cat.); Borgmeier & Prado, 1975: 28; Brown & Kung, 2010: 25. pyricornis Brues, 1944: 151. Type locality: Peru, Upper Rio Pachitea. HT M (Cornell University). Distr.: Brazil (Amazonas), Colombia (Amazonas), Peru. Refs.: Borgmeier, 1968: 37 (cat.); Brown & Kung, 2007: 200. rachelae Disney, 1983: 454. Type locality: Panama, Canal Zone, Pipeline Road. HT M (CUMZ). Distr.: Colombia (Amazonas), Ecuador (Napo), Panama (Canal Zone). Refs.: Brown & Kung, 2007: 200. sanmartinensis Brown & Kung, 2010: 80. Type locality: Colombia, Amazonas, Amacayacu NP. HT M (IAVH). Distr.: Brazil (Alagoas, Amazonas), Colombia (Amazonas (Amacayacu, National Natural Park)). sarmientoi Brown & Kung, 2007: 182. Type locality: Colombia, Valle del Cauca, PNN Farallones de Cali. HT M (UNCB). Distr.: Colombia (Amazonas, Valle del Cauca (Farallones de Cali, National Natural Park)). sarriae Brown & Kung, 2010: 60. Type locality: Colombia, Valle del Cauca, PNN Farallones de Cali. HT M (IAVH). Distr.: Colombia (Valle del Cauca (Farallones de Cali, National Natural Park))), Costa Rica (Guanacaste, San José), Mexico (Chiapas). segregata Brown & Kung, 2010: 101. Type locality: Costa Rica, Heredia, La Selva Biological Station. HT M (LACM). Distr.: Colombia (Chocó), Costa Rica (Heredia, Limón, Puntarenas). shannoni Borgmeier, 1961: 14. Type locality: Peru, Iquitos. HT M (USNM). Distr.: Colombia (Amazonas, Meta), Peru. Refs.: Borgmeier, 1968: 37 (cat.); Brown & Kung, 2007: 182; 2010: 101. sharkeyi Brown & Kung, 2007: 201. Type locality: Colombia, Amazonas, PNN Amacayacu. HT M (UNCB). Distr.: Colombia (Amazonas (Amacayacu, National Natural Park)). simplex Borgmeier & Prado, 1975: 13. Type locality: Trinidad. HT M (MZSP). Distr.: Colombia (Vichada), Trinidad. Refs.: Brown & Kung, 2010: 46. sinuicarina Brown & Kung, 2010: 52. Type locality: Colombia, Caquetá, San Vicente de Caguán. HT M (IAVH). Distr.: Colombia (Amazonas, Caquetá (San Vicente de Caguán)), Ecuador (Napo). sinuicoxa Brown & Kung, 2010: 14. Type locality: Colombia, Amazonas, Amacayacu NP. HT M (IAVH). Distr.: Bolivia (Alto Beni), Brazil (Amazonas), Colombia (Amazonas (Amacayacu, National Natural Park)), Caquetá, Putumayo, Vichada), Ecuador (Napo), Peru (Madre de Dios). sinuosa Borgmeier, 1960: 266. Type locality: Panama, Canal Zone, Fort Clayton. HT M (USNM). Distr.: Colombia (Amazonas, Magdalena), Panama (Canal Zone, Darien). Refs.: Borgmeier, 1968: 37 (cat.); Brown & Kung, 2007: 201. smithi Brown & Kung, 2007: 184. Type locality: Peru, Madre de Dios, Tambopata Research Center. HT M (MUSM). Distr.: Brazil (Amazonas), Colombia (Amazonas), French Guiana (Regina), Peru (Madre de Dios). Refs.: Brown & Kung, 2010: 39. sulcatula Borgmeier, 1960: 283. Type locality: Brazil, Amapá, Serra do Navio. HT M (MZSP). Distr.: Brazil (Amapá, Pernambuco), Colombia (Amazonas, Chocó), Costa Rica (Heredia, Puntarenas), French Guiana (Regina), Panama (Alhajuelo, Canal Zone, Mojinga, Sabana). Refs.: Borgmeier, 1968: 38 (cat.); Brown & Kung, 2007: 184; 2010: 20. 434 · Zootaxa 4122 (1) © 2016 Magnolia Press AMENT & BROWN taura Brown & Kung, 2010: 15. Type locality: Costa Rica, Heredia, La Selva Biological Station. HT M (LACM). Distr.: Colombia (Cauca, Chocó, Valle del Cauca), Costa Rica (Guanacaste, Heredia, Puntarenas, San José), Ecuador (Carchi), Panama (Canal Zone). tayronensis Brown & Kung, 2010: 89. Type locality: Colombia, Magdalena, PNN Tayrona. HT M (IAVH). Distr.: Colombia (Bolívar, Magdalena (Tayrona, National Natural Park)). utriensis Brown & Kung, 2007: 185. Type locality: Colombia, Chocó, PNN Utría. HT M (UNCB). Distr.: Brazil (Amazonas), Colombia (Chocó (Utría, National Natural Park)), Costa Rica (Heredia). Refs.: Brown & Kung, 2010: 73. wangae Brown & Kung, 2007: 186. Type locality: Ecuador, Napo, Yasuni NP. HT M (LACM). Distr.: Colombia (Amazonas), Ecuador (Napo). wilkinsoni Brown & Kung, 2007: 187. Type locality: Ecuador, Napo, Yasuni NP. HT M (LACM). Distr.: Colombia (Amazonas, Caquetá, Vaupés), Ecuador (Napo). xiei Brown & Kung, 2007: 187. Type locality: Ecuador, Napo, Yasuni NP. HT M (LACM). Distr.: Brazil (Amazonas), Colombia (Amazonas), Ecuador (Napo), Panama (Canal Zone), Peru (Madre de Dios). Genus Neopleurophora Brown Neopleurophora Brown, 1992: 51. Type species, Phora scutellata Brues, 1904 (orig. des.). Refs.: Ament & Amorim, 2013: 1. acrochaetopyga Ament & Amorim, 2013: 19. Type locality: Colombia, Chocó, PNN Utria Send. Cocalito. HT M (IAvH). Distr.: Colombia (Chocó (Utria, National Natural Park), Valle del Cauca, Magdalena), Costa Rica (Heredia, San José), Nicaragua (Rio San Juan Refugio Bartola). balbii Ament & Amorim, 2013: 42. Type locality: Panama, Canal Zone, Barro Colorado Is. HT M (LACM). Distr.: Colombia (Vichada), Costa Rica (San José, Puntarenas, Limon), Ecuador (Sucumbios), Panama (Canal Zone). cauca Ament & Amorim, 2013: 43. Type locality: Colombia, Cauca, PNN Gorgona. HT M (IAvH). Distr.: Colombia (Cauca (Gorgona Is, National Natural Park)). ceratopyga Ament & Amorim, 2013: 53. Type locality: Colombia, Meta, PNN Macarena)). HT M (IAvH). Distr.: Brazil (Pará), Colombia (Meta (Macarena, National Natural Park), Vichada, Vaupés)), French Guiana (Regina). chocoensis Ament & Amorim, 2013: 54. Type locality: Colombia, Chocó, PNN Utria Boroboro. HT M (IAvH). Distr.: Colombia (Chocó (Utria, National Natural Park)). dichaeta Ament & Amorim, 2013: 22. Type locality: Colombia, Huila, PNN Cueva de los Guácharos. HT M (IAVH). Distr.: Colombia (Huila (Cueva de Los Guacaros, National Natural Park)). dicrodrilus Ament & Amorim, 2013: 60. Type locality: Colombia, Risaralda, SFF Otún Quimbaya. HT M (IAVH). Distr.: Colombia (Risaralda (Otun Quimbaya, Santuario de Fauna y Flora)). diffusa Ament & Amorim, 2013: 56. Type locality: Colombia, Putumayo, PNN La Paya. HT M (IAVH). Distr.: Brazil (Amazonas), Colombia (Amazonas, Putumayo (La Paya, National Natural Park), Vaupés), Ecuador (Sucumbios, Napo), French Guiana (Regina). dolichopyga Ament & Amorim, 2013: 29. Type locality: Colombia, Cauca, PNN Gorgona. HT M (IAVH). Distr.: CATALOGUE OF PHORIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 435 Brazil (Rondônia), Colombia (Cauca (Gorgona Is, National Natural Park), Chocó); Costa Rica (Puntarenas), French Guiana (Regina). kleini Ament & Amorim, 2013: 12. Type locality: Colombia, Putumayo, PNN La Paya. HT M (IAVH). Distr.: Brazil (Amazonas), Colombia (Amazonas, Caquetá, Putumayo (La Paya, National Natural Park), Vaupés), Ecuador (Sucumbios). marquesi Ament & Amorim, 2013: 36. Type locality: Ecuador, Napo, Yasuni NP. HT M (LACM). Distr.: Brazil (Pará), Colombia (Chocó, Amazonas), Costa Rica (Guanacaste), Ecuador (Napo, Pichincha, Sucumbios). microssoma Ament & Amorim, 2013: 14. Type locality: Colombia, Amazonas, PNN Amacayacu. HT M (IAVH). Distr.: Brazil (Amazonas), Colombia (Amazonas (Amacayacu, National Natural Park), Putumayo), Ecuador (Napo, Zamora-Chinchipe), French Guiana (Regina). odontopyga Ament & Amorim, 2013: 57. Type locality: Ecuador, Napo, Yasuni NP. HT M (LACM). Distr.: Colombia (Amazonas, Valle del Cauca), Costa Rica (Puntarenas), Ecuador (Napo), French Guiana (Regina), Panama (Canal Zone). platypyga Ament & Amorim, 2013: 63. Type locality: Colombia, Amazonas, PNN Amacayacu. HT M (IAVH). Distr.: Colombia (Amazonas (Amacayacu, National Natural Park), Vaupés). polychaetopoda Ament & Amorim, 2013: 66. Type locality: Colombia, Bolívar, SFF Colorados. HT M (IAVH). Distr.: Colombia (Bolívar (Colorados, Santuario de Fauna y Flora), Chocó), Costa Rica (Puntarenas), Panama (Darien Cruce de Mono). prionotopyga Ament & Amorim, 2013: 17. Type locality: Costa Rica, Limón, 16km W Guapiles. HT M (LACM). Distr.: Colombia (Valle del Cauca), Costa Rica (Limón), Nicaragua (Zelaya). ptychodrilus Ament & Amorim, 2013: 37. Type locality: Ecuador, Sucumbios, Sacha Lodge. HT M (LACM). Distr.: Colombia (Amazonas), Ecuador (Sucumbios). scleropyga Ament & Amorim, 2013: 8. Type locality: Colombia, Putumayo, PNN La Paya. HT M (IAVH). Distr.: Colombia (Putumayo (La Paya, National Natural Park), Vaupés). tanytarsus Ament & Amorim, 2013: 23. Type locality: Colombia, Risaralda, SFF Otún Quimbaya. HT M (IAVH). Distr.: Colombia (Risaralda (Otun Quimbaya, Santuario de Fauna y Flora) Valle del Cauca), Costa Rica (Guanacaste, Heredia). Genus Phora Latreille Phora Latreille, 1796: 125 (no species); 1802: 464 (des. of genotype). Type species, Musca aterrima Fabricius, 1794 (preocc. by M. aterrima Villers, 1789 = Trineura atra Meigen, 1804) (subs. mon. Latreille, 1802). Refs.: Latreille, 1805: 394; Becker, 1901: 6 (p. p.); Brues, 1915a: 107; 1950: 44; Lundbeck, 1922: 186; Schmitz, 1918: 117, 119; 1920: 110; Schmitz & Wirth, 1954: 114; Borgmeier, 1963a: 99 (key); 1968: 42 (cat.); 1971: 182 (cat.); Beyer, 1965: 21; Brown, 1992: 47; Disney, 1994: 286. Trineura Meigen, 1803: 276. Type species, Musca aterrima Fabricius, 1794 (subs. des. Brues, 1906). Refs.: Meigen, 1804: 312; Fallén, 1823: 7; Zetterstedt, 1848: 2890; Schiner, 1864: 347; Becker, 1901: 78; Brues, 1904: 339; 1906: 7; Schmitz, 1940: 102; 1953b: 309. Philodendria Rondani, 1856: 136. Type species, Tephritis aterrima Fabricius (orig. des.; =Musca aterrima Fabricius, 1794). americana Schmitz & Wirth, 1954: 124. Type locality: Mexico, Cuenavarca. HT M (USNM). Distr.: Colombia (Cundinamarca (Funza, La Picota)), Costa Rica (Cartago, Heredia), Ecuador (14 km NE Ambato), Mexico (Cuenavarca, D.F., San Jacinto, Jalapa), USA (Texas, Arizona, California, New Mexico, New York). Refs.: Borgmeier, 1963a: 111; 1968: 42 (cat.); Brown, 2000b: 978. 436 · Zootaxa 4122 (1) © 2016 Magnolia Press AMENT & BROWN TABLE 1. Phoridae species recorded from Colombia. Subfamily Genus Chonocephalinae Chonocephalus Metopininae Apocephalus Subgenus Apocephalus Mesophora Cryptophora Eibesfeldtphora Homalophora Species Author depressus Meijere, 1912 dominicanus Borgmeier, 1967 globipygus Borgmeier, 1967 heymonsi Stobbe, 1913 leei Disney, 2008 raposoensis Disney, 2008 vitiodepressus Disney, 2008 asymmetricus Brown, 1997 barbarus Brown, 2002 brochus Brown, 2000 catholicus Brown, 2000 crassilatus Brown, 2000 densepilosus Borgmeier, 1971 dinoponerae Brown, 2000 facettalis Borgmeier, 1961 fernandezi Brown, 2002 flexiseta Brown, 2002 kungae Brown, 2000 longimanus Brown, 2012 lopesi (Borgmeier, 1969) melinus Brown, 2000 neivai Borgmeier, 1931 paraponerae Borgmeier, 1958 reburrus Brown, 2002 roeschardae Brown, 2000 scaurus Corona & Brown, 2004 setitarsus Brown, 1997 spinilatus Brown, 1997 tanyurus Brown, 2000 tenuitarsus Brown, 1997 torulus Brown, 2000 trifidus Brown, 2000 antennatus Malloch, 1913 colombiae Seevers, 1941 arcuata Brown, 2001 attae Disney, 1996 declinata Borgmeier, 1925 dissita Brown, 2001 leei Brown, 2001 pala Brown, 2001 prolixa Brown, 2001 guatiquiae Seevers, 1941 ...Continued on next page CATALOGUE OF PHORIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 437 TABLE 1. (Continued) Subfamily Genus Subgenus Idiophora Megaselia Species Author metae Seevers, 1941 termitoxena (Seevers, 1941) graminicola (Borgmeier, 1969) neocorynurae Gonzalez et al., 2002 parianae (Borgmeier, 1969) perdita (Malloch, 1912) picta (Lehman, 1822) rufipes (Meigen, 1804) scalaris (Loew, 1866) acicula Brown, 2006 acoma Brown & Kung, 2006 caligula Brown & Kung, 2006 candida Brown & Kung, 2006 clandestina Brown, 2005 debilis Brown & Kung, 2006 diastata Brown & Kung, 2006 digitalis Borgmeier, 1959 hirsuta Brown, 2006 hyalinipennis Borgmeier, 1933 juxta Brown & Kung, 2006 kungae Brown, 2005 lamellata Borgmeier, 1934 maculata Borgmeier, 1934 nannocauda Brown, 2006 punctifrons Borgmeier, 1934 rodeoensis Brown, 2006 striatula Borgmeier, 1934 stylata (Schiner, 1868) tuparroensis Brown & Kung, 2006 ancistra Gonzalez & Brown, 2004 flavilata Brown, 2004 Melittophora salti Brues, 1928 Pseudacteon andinus Plowes et al., 2009 antiguensis (Malloch, 1912) arcuatus Borgmeier, 1969 browni Disney, 1991 deltoides Plowes et al., 2009 grandis Greene, 1941 hippeus Plowes et al., 2009 quinni Plowes et al., 2009 Pseudohypocera kerteszi (Enderlein, 1912) Syntermophora microphthalma Seevers, 1941 Melaloncha Melaloncha Udamochiras ...Continued on next page 438 · Zootaxa 4122 (1) © 2016 Magnolia Press AMENT & BROWN TABLE 1. (Continued) Subfamily Phorinae Genus Species Author Tayrona nitifrons Brown & Kung, 2004 Chaetocnemistoptera palpalis (Borgmeier, 1961) lobata Ament, 2014 magdalena Ament, 2014 phaeonota Ament, 2014 pityropyga Ament, 2014 trichopoda Ament, 2014 browni Kung, 2009 caliga Kung & Brown, 2000 reina Kung & Brown, 2000 sakaii Kung, 2009 valdesi Kung, 2009 deinocerca Borgmeier & Prado, 1975 gorgonensis Brown, 2006 acanthiventris Brown & Kung, 2010 acarinata Brown & Kung, 2010 acuta Brown & Kung, 2010 alvarengai Prado, 1976 amacayavia Brown & Kung, 2010 anchicayensis Brown & Kung, 2007 anteroventralis Borgmeier, 1960 aprosmicta Brown & Kung, 2010 atrilinea Brown & Kung, 2010 basilacuna Brown & Kung, 2010 berezovskiyi Brown & Kung, 2010 bicavatura Brown & Kung, 2010 bicostula Kung and Brown, 2005 biseriata Borgmeier, 1960 brevisimplex Brown & Kung, 2010 broadheadi Disney, 1983 bugenitalis Brown & Kung, 2010 buscki Malloch, 1912 caguanensis Brown & Kung, 2010 circularis Brown & Kung, 2007 circumflexa Borgmeier, 1960 clariloba Brown & Kung 2007 cocaensis Brown & Kung, 2007 cognata Borgmeier, 1960 consimilis Brown & Kung 2007 conspicua Brown & Kung, 2010 continua Brown & Kung, 2010 cootei Brown & Kung 2007 Coniceromyia Cyrtophorina Dohrniphora Subgenus ...Continued on next page CATALOGUE OF PHORIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 439 TABLE 1. (Continued) Subfamily Genus Subgenus Species Author cornuta (Bigot, 1857) cunarensis Brown & Kung, 2010 curticerca Brown & Kung, 2007 decrescens Brown & Kung, 2007 didyma Brown & Kung, 2007 digitata Brown & Kung, 2007 dispar (Enderlein, 1912) divaricata (Aldrich, 1896) ecitophila Borgmeier, 1960 emmesta Brown & Kung, 2007 femoralis Borgmeier, 1960 fernandezi Brown & Kung, 2010 flammidorsa Brown & Kung, 2010 flexicarina Brown & Kung, 2010 foveolata Borgmeier, 1960: 278 fuliginata Brown & Kung, 2010 fulleri Brown & Kung, 2010 fundula Brown & Kung, 2010 fuscicoxa Borgmeier, 1923 gigantea (Enderlein, 1912) gravis Borgmeier & Prado, 1975 hamartia Brown & Kung, 2007 heliconiae Brown & Kung, 2010 hexaseta Brown & Kung, 2010 ignobilis Borgmeier, 1960 incomitata Brown & Kung, 2007 inornata Brown & Kung, 2007 knabi Malloch, 1912 lacunosa Brown & Kung, 2007 lamellifera Borgmeier, 1961 lobata Borgmeier, 1960 longinoi Brown & Kung, 2010 longisetosa Brown & Kung, 2010 lugens Borgmeier, 1960 luteicincta Borgmeier, 1960 matamatae Brown & Kung, 2010 membranea Brown & Kung, 2007 mesofemoralis Brown & Kung, 2007 minerva Borgmeier, 1961 minervoidea Brown & Kung, 2010 minilobata Brown & Kung, 2010 mississippiensis Khalaf, 1971 ...Continued on next page 440 · Zootaxa 4122 (1) © 2016 Magnolia Press AMENT & BROWN TABLE 1. (Continued) Subfamily Genus Neopleurophora Subgenus Species Author mixtura Brown & Kung, 2010 nigricarina Brown & Kung, 2010 notoplicata Brown & Kung, 2010 oricilla Kung and Brown, 2005 orthogramma Brown & Kung, 2010 ospinae Brown & Kung, 2010 ovibarba Brown & Kung, 2007 palenquensis Brown & Kung, 2010 pallidens Borgmeier & Prado, 1975 paraguayana Brues, 1907 paralobata Brown & Kung 2007 parvicarinata Brown & Kung, 2010 parvicava Brown & Kung, 2010 parvipala Brown & Kung, 2010 phyzela Brown & Kung, 2010 porrasae Brown & Kung, 2010 preacuta Brown & Kung, 2010 procera Borgmeier, 1960 pyricornis Brues, 1944 rachelae Disney, 1983 sanmartinensis Brown & Kung, 2010 sarmientoi Brown & Kung, 2007 sarriae Brown & Kung, 2010 segregata Brown & Kung, 2010 shannoni Borgmeier, 1961 sharkeyi Brown & Kung, 2007 simplex Borgmeier & Prado sinuicarina Brown & Kung, 2010 sinuicoxa Brown & Kung, 2010 sinuosa Borgmeier, 1960 smithi Brown & Kung, 2007 sulcatula Borgmeier, 1960 taura Brown & Kung, 2010 tayronensis Brown & Kung, 2010 utriensis Brown & Kung, 2007 wangae Brown & Kung, 2007 wilkinsoni Brown & Kung, 2007 xiei Brown & Kung, 2007 acrochaetopyga Ament & Amorim, 2013 balbii Ament & Amorim, 2013 cauca Ament & Amorim, 2013 ceratopyga Ament & Amorim, 2013 ...Continued on next page CATALOGUE OF PHORIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 441 TABLE 1. (Continued) Subfamily Genus Phora Subgenus Species Author chocoensis Ament & Amorim, 2013 dichaeta Ament & Amorim, 2013 dicrodrilus Ament & Amorim, 2013 diffusa Ament & Amorim, 2013 dolichopyga Ament & Amorim, 2013 kleini Ament & Amorim, 2013 marquesi Ament & Amorim, 2013 microssoma Ament & Amorim, 2013 odontopyga Ament & Amorim, 2013 platypyga Ament & Amorim, 2013 polychaetopoda Ament & Amorim, 2013 prionotopyga Ament & Amorim, 2013 ptychodrilus Ament & Amorim, 2013 scleropyga Ament & Amorim, 2013 tanytarsus Ament & Amorim, 2013 americana Schmitz & Wirth, 1954 References Aldrich, J.M. (1896) Dolichopodidae; Phoridae. Transactions of the Royal Entomological Society of London, 1896, 253–446. Ament, D.C. (2014) Taxonomic revision of the genus Chaetocnemistoptera Borgmeier (Diptera: Phoridae), with the description of five new species. Zootaxa, 3753 (4), 301–322. http://dx.doi.org/10.11646/zootaxa.3753.4.1 Ament, D.C. & Amorim, D.S. 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(1990) Molecular differentiation of the homomorphic sex chromosomes in Megaselia scalaris (Diptera) detected by random DNA probes. Chromosoma, 99, 237–242. http://dx.doi.org/10.1007/BF01731698 Wolf, K.W., Jeppesen, P. & Mitchell, A. (1993) Spermatid Nucleus of Megaselia scalaris Loew (Insecta, Diptera, Phoridae): A Study Using Anti-Histone Antibodies, Scanning Electron Microscopy, and a Centromere-Specific Oligonucleotide. Molecular Reproduction and Development, 35, 272–276. http://dx.doi.org/10.1002/mrd.1080350309 Wood, J.H. (1908) On the British species of Phora (Part II). Entomologist's Monthly Magazine, 44, 164–174, 215–218, 253–254. Wood, J.H. (1909) On the British species of Phora. Part II, cont. Entomologist's Monthly Magazine, 45, 24–29, 59–63, 113–120, 143–149 191–195, 240–244. Zetterstedt, J.W. (1838/40) Insecta Lapponiae. Leipzig, VI + 140 pp. [Diptera pp. 477–868] Zetterstedt, J.W. (1848) Diptera Scandinaviae disposita et descripta. Vol. 7. ex Officina Lundbergiana, sumtibus auctoris, Lundae, 354 pp. [pp. 2581–2934] Zetterstedt, J.W. (1852) Diptera Scandinaviae disposita et descripta. Vol. 11. ex Officina Lundbergiana, sumtibus auctoris, Lundae, 456 pp. [pp. V–VII + 4091–4546] Zetterstedt, J.W. (1860) Diptera Scandinaviae disposita et descripta. Vol. 14. ex Officina Lundbergiana, sumtibus auctoris, Lundae, 419 pp. [pp. 6191–6609] CATALOGUE OF PHORIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 451 Zootaxa 4122 (1): 452–456 http://www.mapress.com/j/zt/ Copyright © 2016 Magnolia Press Article ISSN 1175-5326 (print edition) ZOOTAXA ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.4122.1.38 http://zoobank.org/urn:lsid:zoobank.org:pub:EB6C55D3-5780-4912-A351-D80E84B7511C FAMILY PIPUNCULIDAE JOSÉ ALBERTINO RAFAEL1* & HUGO CESAR RODRIGUEZ2 1 Instituto Nacional de Pesquisas da Amazônia (INPA), Coordenação de Biodiversidade, Av. André Araújo 2936, CP 478, 69011–970, Manaus, Brazil; E–mail: jarafael@inpa.gov.br 2 Laboratorio de Enemigos Naturales de Plagas Agrícolas del NOA, PROIMI–Biotecnología, Av. Belgrano y Pje. Caseros (CP 4000), San Miguel de Tucumán, Tucumán, Argentina. E–mail: huguitorod@hotmail.com * Corresponding author: jarafael@inpa.gov.br Abstract Pipunculidae presently have around 1,400 known species, but it is poorly known in Colombia, with only ten reported species in all territory. The genera Eudorylas Aczél and Tomosvaryella Aczél are the most abundant. This work provides information on the species distribution. No endemic genera have been recorded to date from Colombia. Key words: big-headed fly, catalogue, Colombia, distribution Introduction Pipunculids, also known as big-headed flies, are small flies measuring 2.0–11.5 mm long. They are closely related to Syrphidae but can be differentiated by the large compound eyes that occupy most of the globose or semiglobose head; by the lack of spurious vein; and by the open R4+5 cell (Rafael & Skevignton 2010). The pterostigma is a character present in most genera. Globally 1,401 species have been described to date in 28 genera or subgenera, and it is likely that more than 2,700 species exist (De Meyer 1996, De Meyer & Skevington 2000). In South America it is known so far 203 species (Rodríguez & Rafael 2012). They are classified in 18 genera divided into three subfamilies: Chalarinae, Nephrocerinae y Pipunculinae (Rafael & De Meyer 1992, Skevington & Yeates 2000, Rafael & Skevington 2010). The taxonomy of this family is well known. The species identification typically relies on characters of the terminalia and in most cases only males can be identified. The biology of the group remains poorly know because few studies have examined life cycles and host relationships. Most studies in Latin America have their foundation in Hardy (1948), who studied specimens that were collected in various Latin America countries and mostly deposited in Unites States museums (Rodríguez & Rafael 2012). The only study on the fauna of Pipunculidae in Colombia is the work of Hardy (1963), describing four new species and recording in that occasion the first known host family in the Neotropical Region. Most species that have been studied are exclusively endoparasitoids of Auchenorrhyncha (Hemiptera), records include Cercopidae, Cixiidae, Flatidae, Fulgoridae and Membracidae, but in the Neotropical region there are only records of parasitism for Cicadellidae and Delphacidae (Virla & Rafael 1996). The most effective method for studying pipunculid biology involves capturing potential hosts by sweep-collecting in croplands and forest areas; this allows not only knowing but offer information about the place where each species lives. Pipunculidae are widely distributed in Latin America, occurring from northern Mexico to the Tierra del Fuego Island of Chile and are found in a variety of environments (Rodríguez & Rafael 2012). Bearing in mind these facts and the great variety of environments that the Colombian territory offers, we believe that the amount of actual species in the country is a lot larger than the recorded one to date, that is why the collects in different regions as well as the formation of specialists to identify the material will increase notably the knowledge of biology and distribution of the family in Colombia. Nowadays there are 10 species recorded of pipunculids (Table 1), most of 452 Accepted by S. Nihei: 26 May 2015; published: 14 Jun. 2016 them on the west region of the country, in the Pacific and Andean ecoregions. The main depositaries of the Pipunculidae collections are the collections of the Instituto de Investigación de Recursos Biológicos Alexander von Humboldt of Bogotá which held 190 specimens, though there are few samples identified and information is not digitalized; and the Museo Entomológico Francisco Luis Gallego of Medellín has five species of the genus Dorylomorpha, collected in the state of Nariño. Acronyms used for the depositories INPA—Instituto Nacional de Pesquisas da Amazônia, Manaus, Brazil. MCZ—Museum of Comparative Zoology–Harvard University, Cambridge, Massachusetts, USA. MNRJ—Museu Nacional do Rio de Janeiro, Rio de Janeiro, Brazil. USNM—United States National Museum, Smithsonian Institution, Washington, D. C, USA. List of abbreviations cat.—catalogue distr.—distribution HT—Holotype orig. des.—by original designation pp.—pages PT—paratype refs.—reference(s) rev.—revision Catalogue of Pipunculidae of Colombia Family Pipunculidae Subfamily Pipunculinae Latreille, 1802 Genus Amazunculus Rafael Amazunculus Rafael, 1986: 16. Type species: Dorilas (Eudorylas) platypodus Hardy, 1950 (orig. des.). cordigaster Galinkin & Rafael, 2008: 519. Type locality: Colombia, Amacayacu Mata-mata. HT ♂ (INPA). Distr.: Colombia (Cauca; Amazonas), Brazil. Refs.: Galinkin & Rafael, 2008; Rodriguez & Rafael, 2012 (cat.). Host unknown. Genus Dorylomorpha Aczél Dorylomorpha Aczél, 1939: 22 (Tomosvaryella subgenus). Type species: Pipunculus rufipes Meigen, 1824 (orig. des.). reveloi Hardy, 1963: 259. Type locality: Colombia, Boyaca, Bonza. HT ♂ (USNM). Distr.: Colombia (Cundinamarca). Refs.: Albrecht, 1990 (rev.); Rodriguez & Rafael, 2012 (cat.). Host unknown. Genus Eudorylas Aczél Eudorylas Aczél, 1940: 151. Type species,Cephalops opacus Fallén, 1816 (orig. des.). CATALOGUE OF PIPUNCULIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 453 Metadorylas Rafael, 1987: 35. Type species, Pipunculus schreiteri Shannon, 1927. Synonymy by Skevington & Yeates (2001). absonditus (Hardy, 1954): 14. Type locality: Brazil, Rio de Janeiro. Refs. Hardy, 1963. HT ♀ (MNRJ). Distr.: Colombia (Nariño), Brazil. Refs.: Hardy, 1963 (Colombian species). Host: Cicadulina pastusae Ruppel & De Long (Cicadellidae). dumicolus (Hardy, 1963): 265. Type locality: Colombia, Pedregal. HT ♂ (USNM). Distr.: Colombia (Nariño). Refs.: Rafael, 1993 (Mexican and American Central species); Rodríguez & Rafael, 2012 (cat.). Host unknown. spinosus (Hardy, 1948): 4. Type locality: Costa Rica, Hamburg Farm. HT ♀ (MCZ). Distr.: Colombia (Cundinamarca), Argentina, Brazil. Refs.: Rafael, 1990 (revision of Metadorylas); Rodríguez & Rafael, 2012 (cat.). Host: Cicadulina pastusae Ruppel & De Long and Dalbulus maidis (Cicadellidae). subopacus subopacus (Loew, 1866): 176. Type locality: USA, Washington. HT ♀ (MCZ). Distr.: Colombia (Nariño), Mexico, Peru, Argentina. Refs.: Rafael, 1990; Virla et al., 2009 (hosts); Rodríguez & Rafael, 2012 (cat.). Host: Cicadulina pastusae Ruppel & De Long, Scaphytopius acutus (Say), Dalbulus maidis (DeLong & Wolcott) and Dalbulus elimatus (Ball) (Cicadellidae). Genus Microcephalops De Meyer Microcephalops De Meyer, 1989: 120. Type species, Pipunculus banksi Aczél, 1940 (orig. des.). williamsi (Hardy, 1954): 54. Type locality: Brazil, Rio de Janeiro. HT ♂ (USNM). Distr.: Colombia, Brazil, Peru. Refs.: Rafael, 1996 (Dominican Republic species); Rodríguez & Rafael, 2012 (cat.). Host unknown. Genus Tomosvaryella Aczél Tomosvaryella Aczél,1939: 22. Type species, Pipunculus sylvaticus Meigen, 1824 (orig. des.). prostata Hardy, 1963: 261. Type locality: Colombia, Cundinamarca. HT ♂ (USNM). Distr.: Colombia (Cundinamarca). Refs.: Rodríguez & Rafael, 2012 (cat.). Host unknown. scopulata Hardy, 1963: 262. Type locality: Colombia, Palmira. HT ♂ (USNM). Distr.: Colombia (Nariño, Tolima, Valle del Cauca), Argentina, Cuba, Dominica, Mexico, Nicaragua. Refs.: Rodríguez & Rafael, 2012 (cat.). Host unknown. subvirescens (Loew, 1872): 87. Type locality: USA, Texas. Ref. Rafael & Ale-Rocha 1997. HT ♂ (MCZ). Distr.: Colombia, Bahamas, Brazil, Belize, Chile, Dominica, Jamaica, Puerto Rico. Refs: Virla et al., 2009; Rodríguez & Rafael, 2012 (cat.). Host: Circulifer tenellus Baker (Cicadellidae). Acknowledgements To Conselho Nacional de Desenvolvimento Científico e Tecnológico (CNPq) for the award of a fellowship to J.A.R. (grant 300.305/2007–9). To CAPES for the Ph.D. scholarship to H.C.R. (proc. 551.991/2011–9). To Fundação de Amparo à Pesquisa do Estado do Amazonas (FAPEAM) by financial support (edital numbers 021/ 2011 and 020/2013). Dr. Sergio Orduz and Jhon Alveiro Quiroz Gamboa coordinators of Museo Entomológico Francisco Luis Gallego, Medellin, Colombia and Claudia Alejandra María Uribe coordinator of Colecciones Biológicas del Instituto de Investigación de Recursos Biológicos Alexander von Humboldt, Villa de Leyva, Colombia, for information on the specimens deposited in their institutions. 454 · Zootaxa 4122 (1) © 2016 Magnolia Press RAFAEL & RODRIGUEZ TABLE 1. Pipunculidae species recorded from Colombia. Subfamily Tribe Genus Pipunculinae Eudorylini Amazunculus Tomosvaryellini Dorylomorpha Eudorylini Eudorylas Subgenus Pipunculina species author cordigaster Galinkin & Rafael, 2008 reveloi Hardy, 1963 absonditus (Hardy, 1954) dumicolus (Hardy, 1963) spinosus (Hardy, 1948) subopacus (Loew, 1866) Microcephalopsini Microcephalops williamsi (Hardy, 1954) Tomosvaryellini Tomosvaryella prostata Hardy, 1963 scopulata Hardy, 1963 subvirescens (Loew, 1872) References Albrecht, A. (1990) Revision, phylogeny and classification of the genus Dorylomorpha (Diptera, Pipunculidae). Acta Zoologica Fennica, 188, 1–240. Aczél, M. (1939) Das System der Familie Dorylaidae. Dorylaiden–Studien I. Zoologischer Anzeiger, 125, 15–23. Aczél, M. (1940) Vorarbeiten zu einer Monographie der Dorylaiden (Dipt.). Dorylaiden–Studien V. Zoologischer Anzeiger, 132, 149–169. De Meyer, M. (1989) Systematics of the Neartic species of the genus Cephalops Fallén (Diptera, Pipunculidae). Journal of African Zoology, 104, 181–189. De Meyer, M. (1996) World catalogue of Pipunculidae (Diptera). Institut Royal des Sciences Naturelles de Belgique, Documents de Travail, 86, 1–127. De Meyer, M. & Skevington, J.H. (2000) First addition to the world catalogue of Pipunculidae. Bulletin de L' Institut Royal des Sciences Naturelles de Belgique. Entomologie, 70, 5–11. Fallén, C.F. (1816) Syrphici Sveciae. [Part]. Lundae, Berling, 22 pp. Galinkin, J. & Rafael, J.A. (2008) Revisão do gênero neotropical Amazunculus Rafael (Diptera: Pipunculidae) com descrição de três novas espécies e chave de identificação. Acta Amazonica, 38 (3), 517–524. http://dx.doi.org/10.1590/S0044-59672008000300017 Hardy, D.E. (1948) Neotropical Dorilaidae (Pipunculidae) Studies, Part 1 (Diptera). Psyche, 55, 1–15. http://dx.doi.org/10.1155/1948/25050 Hardy, D.E. (1950) Neotropical Dorilaidae Studies. Part 2 (Pipunculidae, Diptera). Revista de Entomologia, 21, 433–448. Hardy, D.E. (1954) Neotropical Dorilaidae Studies. Part III. Brazilian species and a key to the know species of Dorilas sens. lat. Boletim do Museu Nacional do Rio de Janeiro, 123, 1–60. Hardy, D.E. (1963) Studies in Pipunculidae (Diptera) of Colombia. Hawaiian Entomological Society, 18, 259–266. Loew, H. (1866) Diptera Americae septentrionalis indigena. Centuria decima. Berliner Entomogische Zeitschrift, 9, 127–186. http://dx.doi.org/10.1002/mmnd.18650090303 Loew, H. (1872) Diptera Americae septentrionalis indigena. Centuria decima. Berliner Entomogische Zeitschrift, 16, 49–124. http://dx.doi.org/10.1002/mmnd.18720160110 Meigen, J.W. (1824) Systematische Beschreibung der bekannten europaischen zweiflugeligen Insekten. Vierter Theil. Schulz–Wunderman, Hamm, xii + 428 pp. Rafael, J.A. (1986) Amazunculus, a new genus of pipunculid from the Amazon basin (Diptera: Pipunculidae). Amazoniana, X (1), 15–19. Rafael, J.A. (1987) Two new genera of Pipunculidae (Diptera) from the New World: Metadorylas gen. n. and Elmohardyia gen. n., with new synonyms, designation of lectotypes and revalidation of a species. Revista Brasileira de Entomologia, 31 (1), 35–39. Rafael, J.A. (1990) As espécies neotropicais de Metadorylas Rafael (Diptera: Pipunculidae). Boletim do Museu Paraense Emilio Goeldi, Serie Zoologia, 6 (2), 127–164. Rafael, J.A. (1993) Espécies de Eudorylas Aczél (Diptera, Pipunculidae) do México e América Central. Revista Brasileira de Entomologia, 37 (4), 751–762. Rafael, J.A. (1996) Pipunculidae (Insecta: Diptera) of the Dominican Republic: New records and description of new species. Annals of Carnegie Museum, 65 (4), 363–381. Rafael, J.A. & De Meyer, M. (1992) Generic classification of the family Pipunculidae (Diptera): a cladistic analysis. Journal of Natural History, 26, 637–658. CATALOGUE OF PIPUNCULIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 455 http://dx.doi.org/10.1080/00222939200770391 Rafael, J.A. & Ale–Rocha, R. (1997) Chilean Pipunculidae (Diptera). Key to genera, new records, description of a new species of Eudorylas Aczél and a catalog of species. Revista Chilena de Entomología, 24, 31–36. Rafael, J.A. & Skevington, J.H. (2010) Pipunculidae (big-headed flies). In: Brown B.V; Borkent A; Cumming J.M; Wood D.M; Woodley N.E and Zumbado M.A (Eds.), Manual of Central American Diptera. Vol. 2. NRC Research Press, Ottawa, Ontario, Canada, pp. 793–803. Rodríguez, H.C. & Rafael, J.A. (2012) Pipunculidae (Diptera) of Latin America and the Caribbean: A Catalog of Species with Notes on Biology and Pipunculid–Host Associations Pipunculidae. Lambert Academic Publishing, Germany, 50 pp. Shannon, R.C. (1927) Some new Diptera from Argentina. Revista de la Sociedad Entomologica Argentina, 1 (4), 31–42. Skevington, J.H. & Yeates, D.K. (2000) Phylogeny of the Syrphoidea (Diptera) Inferred from mtDNA Sequences and Morphology with Particular Reference to Classification of the Pipunculidae (Diptera). Molecular Phylogenetics and Evolution, 16 (2), 212–224. http://dx.doi.org/10.1006/mpev.2000.0787 Skevington, J.H. & Yeates, D.K. (2001) Phylogenetic classification of Eudorylini (Diptera: Pipunculidae). Systematic Entomology, 26, 421–452. http://dx.doi.org/10.1046/j.0307-6970.2001.00160.x Virla, E.G. & Rafael, J.A. (1996) Datos bionomicos preliminares y descripcion de la hembra de Cephalops penepauculus (Hardy) (Diptera, Pipunculidae) um parasitoide de Delphacidae (Homoptera, Auchenorrhyncha) en Argentina. CIRPON, Revista de Investigaciones, X (1–4), 33–36. Virla, E.G., Moya-Raygoza, G. & Rafael, J.A. (2009) First record of Eudorylas schreiteri (Shanon) (Diptera: Pipunculidae) as a parasitoid of the corn leafhopper (Hemiptera: Cicadellidae) in Argentina, with a table of pipunculid-host associations in the Neotropical region. Neotropical Entomology, 38 (1), 152–154. http://dx.doi.org/10.1590/S1519-566X2009000100019 456 · Zootaxa 4122 (1) © 2016 Magnolia Press RAFAEL & RODRIGUEZ Zootaxa 4122 (1): 457–537 http://www.mapress.com/j/zt/ Copyright © 2016 Magnolia Press Article ISSN 1175-5326 (print edition) ZOOTAXA ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.4122.1.39 http://zoobank.org/urn:lsid:zoobank.org:pub:F95F1B6A-A1BB-46EE-95E3-0B8C5B16ADA0 FAMILY SYRPHIDAE AUGUSTO L. MONTOYA Instituto de Biología, Grupo de Entomología Universidad de Antioquia, Medellín Colombia. E–mail: guto.spider@gmail.com, aleon.montoya@udea.edu.co Abstract This paper is an updated Catalogue of Syrphidae from Colombia, including 337 species belonging to 3 subfamilies, 11 tribes, 57 genera and 16 subgenera. Each species is listed under its current accepted taxon, including the original reference, type locality, distribution by state/province in Colombia, in addition to a complete bibliographic reference. Species in the genera Calostigma, Claraplumula, Eristalinus, Fazia, Hermesomyia, Hybobathus, Mimocalla, Menidon, Monoceromyia, Orphnabaccha, Pelecinobaccha, Peradon, Rhinoprosopa, Sphiximorpha and Styxia are listed for the first time from Colombia. The advance in the knowledge of the Colombian Syrphidae is briefly discussed. Key words: Catalogue, Colombia, Syrphidae, Flower Flies, Geographical Distribution, Colombian Biodiversity, updated checklist Introduction Syrphidae are one of the most specious Diptera families in the Neotropical region with many undescribed species waiting to be discovered and described (Thompson et al. 1976; Thompson 1981, 1999a; Thompson et al. 2010). The family is known as hover flies because exhibit high maneuverability in flight, remaining hovering in a static position. The flower flies are pollinators, feeding on pollen and nectar, an energetic resource for flight and mating events (Rotheray & Gilbert 2011). Immature states display a wide range of lifestyles and adaptations. Syrphinae are predators of colonial insects such as Hemiptera with beneficial effects as biocontrol agents (Rojo et al. 2003). In Syrphinae, some species are predators of aquatic insect in bromeliads (Rotheray et al. 2000), while others are secondarily leaf–miners (Nishida et al. 2003), stem–borers (Zuijen & Nishida 2010) or pollen–feeders (Reemer & Rotheray 2009). Eristalinae feed on decaying organic matter, with some species feeding on animal dung, water reservoirs in Phytotelmata, while others species are saprophagous feeding on exuding plant sap (Gilbert et al. 1994). Microdontinae are predators of immature states of ants (Reemer & Ståhls 2013). The study of flower flies in Colombia started with Macquart (1842–1850), who described four new species. Until 1892, the number of species was low, with only 25 species described by six authors. Between 1925 and 1951, many species were described (50 species), mainly supported by the works of Hull (1925–1951) and Curran (1925–1941). In 1976, Thompson listed 96 species from Colombia. In posterior studies, new species were described, and numerous new synonymies, new combinations, new records and various nomenclatural changes were proposed, increasing the number of Colombian species to 200 (Hippa & Thompson 1983; Thompson 1994; 1999a, 2006; Gutiérrez et al. 2005; Restrepo-Ortiz & Carrejo 2009). The present work provides an updated catalogue of the Colombian Syrphidae including the examination of material in collections in addition to a literature review (Montoya et al. 2012; Ricarte et al. 2012; Mengual, 2014; Miranda et al. 2014). The catalogue contains 337 species and 57 genera. Species in the genera Calostigma, Claraplumula, Eristalinus, Fazia, Hermesomyia, Hybobathus, Mimocalla, Menidon, Monoceromyia, Orphnabaccha, Pelecinobaccha, Peradon, Rhinoprosopa, Sphiximorpha and Styxia are listed for the first time from Colombia (Table 1). Valid names were taken from the Nomenclator of the BioSystematic Database of World Diptera (Thompson 2013). Abbreviations follow the proposed standard of Thompson (1999b). Accepted by S. Nihei: 20 Apr. 2015; published: 14 Jun. 2016 457 Acronyms used for the depositories AMNH—American Museum of Natural History, New York, USA. ANSP—The Academy of Natural Sciences, Philadelphia, PA, USA. BMNH—The Natural History Museum, London, UK. CAS—Department of Entomology, California Academy of Sciences, SF, USA. CEUA—Colección de Entomología Universidad de Antioquia, Medellín, Colombia. CMP—Carnegie Museum of Natural History, Section of Insect and Spiders, Pittsburgh, PA, USA. CNC—Canadian National Collection of Insects, Ottawa, ON, Canada. CUI—CUIC—Cornell University Insect Collection, Ithaca, NY, USA. DEI—Deutsches Entomologisches Institut, Deutschen Akademie der Landwirtswissenschaften zu Berlin, Germany. ICN—Instituto de Ciencias Naturales de la Universidad Nacional de Colombia Sede Bogotá, Bogotá, Colombia. INBio—Instituto Nacional de Biodiversidad, Santo Domingo de Heredia, Costa Rica. IZUSN—Instituto di Zoologia, Universita degli Studia di Napoli, Portici, Italy. LSL—Linnean Society, Burlington House, Piccadilly, London, UK. MACN—Museo Argentino de Ciencias Naturales ‘Bernardino Rivadivia’, Buenos Aires, Argentina. MCZ—Museum of Compartative Zoology, Harvard University, Cambridge, MA, USA. MEFLG—Museo Entomológico Francisco Luis Gallego, Universidad Nacional de Colombia, Medellín, Colombia. MEPB—Museo Entomológico Piedras Blancas, Universidad de Antioquia, Medellín, Colombia. ML—Rijksmuseum van Natuurlijke Histerie, Leiden, Netherlands. MLUH—Martin-Luther-Universität, Wissenschaftsbereich Zoologie, Halle (Saale), Germany. MNHNP—Muséum National d´Histoire Naturelle, Paris, France. MNHNS—Mueso Nacional de Historia Natural, Santiago, Chile. MZUSP—Museu de Zoologia, Universidade de Sao Paulo, Sao Paulo, Brazil. NMW—Naturhistorisches Museum Wien, Vien, Austria OHSU—Ohio State University, Columbus, USA. QZAZ—Departamento de Biología, Pontífica Universidad Católica del Ecuador, Quito, Ecuador. SMF—Forschungsinstitut und Naturmuseum Senckenberg, Hessen, Germany. SMT—Department of Entomology Collection, Staatliches Museum für Tierkunde, Dresden, Forschungsstelle, Dresden, Germany. UChS—Museo Entomologico, Universidad de Chile, Santiago, Chile. UK—Snow Entomological Museum, University of Kansas, Lawrence, USA. UMO—Hope Entomological Collections, University Museum, Park Road, Oxford, UK. UNAB—Museo Entomológico de la Facultad de Agronomía, Universidad Nacional de Colombia, Bogotá, Colombia. USNM—National Museum of Natural History, Smithsonian Institution, Washington, D.C, USA. UZMC—University of Copenhagen, Zoological Museums, Department of Entomology, Copenhagen, Denmark. ZMHU—Museum für Naturkunde der Humboldt Universitat zu Berlin, Bereich Zoologisches Museum, Berlin, Germany. List of abbreviations aut.—automatic cat.—catalogue col.—Collected desc.—description des.—designation distr.—geographic distribution emend.—emendation i.e.—example 458 · Zootaxa 4122 (1) © 2016 Magnolia Press MONTOYA m.—meters mon.—monotypy nom. nov.—nomen novum (new name) orig. des.—by original designation pp.—pages preocc.—preoccupied refs.—references rev.—revision sp.—species ssp.—subspecies subs.—subsequent subg.—subgenus Vsr.—Van Sommeren—Rydon trap HT = Holotype NT = Neotype LT = Lectotype ST = Syntype T = Type (unspecified) NA = Not applicable F (f#)= Female M (m#)= Male Catalogue of Syrphidae of Colombia Family Syrphidae Latreille, 1897 Superfamily Syrphoidea Coquillett, 1901 Subfamily Syrphinae Latreille, 1802 Tribe Bacchini Williston Genus Argentinomyia Lynch Arribálzaga Argentinomyia Lynch Arribalzaga, 1891:199 (40). Type species, testaceipes Lynch Arribalzaga (mon.). Refs.: Fluke, 1945:1–30 (key and rev.); Fluke, 1957b:261–279 (study of males genitalia); Thompson et al. 1976:46 (cat.), Thompson, 1981:105 (desc. notes); Thompson, 1999a:325 (key to Argentinomyia spp.), 327 (key to A. octomaculata), 338 (status notes); Marinoni et al. 2007:149, 150 (desc, key, illustrations); Thompson & Skevington, 2014 (key and review of melanostomine genera groups). Rhysops Williston, 1907:2. Type species, Melanostoma rugosonasus Williston (Coquillett, 1910:601, as rugonasus Williston). Proposed for “Melanostoma scitulum, rugonasus, melanocerus, etc.”. Braziliana Curran, 1925b:252. Type species, Melanostoma longicornis Williston (orig. des.) = longicornis Walker. Allograptina Enderlein, 1938:226. Type species, Allograptina octomaculata Enderlein. altissima (Fluke), 1945:20 (Melanostoma). Type locality: Ecuador, Imbabura, Cuicocha. HT M (AMNH). Distr.: Colombia (Antioquia (Andes, Belmira, Medellín), Cundinamarca (Bogotá), Tolima (Murillo)), Ecuador. Refs.: Fluke, 1945:20, figs. 48 & 49 (abdomen) 1957b:276, figs. 95–96 (M genitalia); Thompson et al. 1976:42 (cat.). Material examined. COLOMBIA: Antioquia: Andes, Cerros Caramanta, Paramo Laguna de Santarita, 5.550249°, -75.988733°, 3610m, 1male, 1female, 3-5.iii.2009, Leg. M. Wolff, Net (CEUA); Belmira, Paramo de Belmira, 6.707065°, -75.714136°, 3057m, 1male, 2females, 5.iii.2009, Leg. A.L. Montoya, Net (CEUA); Medellín, Corregimiento de Santa Elena, Vereda Piedras Blancas, en Robledal, 6.291588°, -75.493345°, 2200m, 1male, 26.ix.2009, Leg. C. Bota, Net (CEUA); Cundinamarca: Bogotá, 4.592901°, -74.081040°, 2700m, 1male, 25.iii.1937, Leg. Osorno (in the collection of F.M. Hull); Bogotá, 4.592901°, -74.081040°, 2700m, 1male, CATALOGUE OF SYRPHIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 459 25.iii.1937, Leg B. Guerra (USNM); Tolima: Murillo, Robledal, 4.870242N, 75.177614W, 3000m, 1male, 6.ix.2012, Leg. C. Bota, Net (CEUA). bolivariensis (Fluke), 1945:19 (Melanostoma). Type locality: Ecuador, Bolivar, Hacienda Talahua. HT M (AMNH). Distr.: Colombia (Antioquia (Belmira)), Ecuador. Refs.: Fluke, 1945:19, figs. 29 (head), 45 (abdomen) 1957b:276, (M genitalia); Thompson et al. 1976:42 (cat.). Material examined. COLOMBIA: Antioquia: Belmira, Paramo de Belmira, 6.707065°, -75.714136°, 3057m, 1male, 5.iii.2009, Leg. A.L. Montoya, Net (CEUA). browni (Fluke), 1945:18 (Melanostoma). Type locality: Ecuador, Bolivar, Hacienda Talahua. HT M (AMNH). Distr.: Colombia (Antioquia (Belmira)), Ecuador. Refs.: Fluke, 1945:18, figs. 27 (antenna), 28 (head), 44 (abdomen) 1957b:276, figs. 107–108 (M genitalia); Thompson et al. 1976:42 (cat.). Material examined. COLOMBIA: Antioquia: Belmira, Paramo de Belmira, 6.707065°, -75.714136°, 3057m, 1male, 5.iii.2009, Leg. A.L. Montoya, Net (CEUA). columbiana (Enderlein), 1938:202 (Braziliana). Type locality: Colombia, "Cordillera, tierra templada". HT M/F (ZMHU). Distr.: Colombia (Magdalena). Refs.: Thompson et al. 1976:42 (cat.). colombianum (Enderlein), 1938:198 (Melanostoma). crenulata (Williston), 1891:12 (Melanostoma). Type locality: Mexico: Guerrero, Xucumanatlan, 7000ft & Omilteme, 8000ft. ST M H (BMNH). Distr.: Colombia (Antioquia (Betania)). Refs.: Williston, 1891:12, pi. 1:5 (habitus), 5a–b (head); Fluke, 1945: 5, fig. 3 (head). Material examined. COLOMBIA: Antioquia: Betania, Vereda La Oculta, 5.736143°, -76.028237°, 17002100m, 2male, 8.viii.1990, Forest, Leg O. Ortega, Net (Coll. O. Ortega). festiva Fluke, 1945:10. Type locality: Ecuador, Tungurahua, Baños. HT M (AMNH). Distr.: Colombia (Antioquia (La Ceja, Rionegro), Cundinamarca (Silvania)), Ecuador. Refs.: Fluke, 1945:10, figs. 13 (head), 42 (abdomen); Thompson et al. 1976:43 (cat.). Material examined. COLOMBIA: Antioquia: La Ceja, Alto de La Union, 6.016132°, -75.370182°, 2500m, 1male, 10.iv.2008, Leg. A.L. Montoya, Net (CEUA); Rionegro, Cabeceras, Finca La escondida, 6.140929°, 75.411164°, 2100m, 17.v.2013, Leg. D. Caraballo, Manual (CEUA); Cundinamarca: Silvania, Vereda Azafranal, 4.342730°, -74.404917°, 1500m, 1female, 6.xii.2009, Leg. N. Flórez, Net in Coffea arabica (UNAB). longicornis (Walker), 1837:343 (Pipiza). Type locality: "South America". HT F (BMNH). Distr.: Colombia (Valle del Cauca (Venta Queremal and Buenaventura)), Brazil, Costa Rica, Ecuador, Guatemala, Peru. Refs.: Fluke, 1945:3, fig. 1 (head) 1957b:276, figs. 4, 14 133 (M genitalia); Thompson et al. 1976:43 (cat.); Borges & Pamplona, 2003:164, figs. 2 (antenna), 3 (abdomen). Material examined. COLOMBIA: Valle del Cauca: Venta Queremal and Buenaventura, 1000–1300m, 1female, Leg. H.F. Schwarz (AMNH). luculenta (Fluke), 1945:17 (Melanostoma). Type locality: Ecuador, Tungurahua, Baños. HT M (AMNH). Distr.: Colombia (Antioquia (Guarne, La Ceja), Cundinamarca (Bogotá, Mosquera)), Ecuador. Refs.: Fluke, 1945:17, figs. 26 (head), 46 (abdomen); Thompson et al. 1976:43 (cat.). Material examined. COLOMBIA: Antioquia: Guarne, Vereda Piedras Blancas, 6.291588°, -75.493345°, 2200m, 1male, 1female, 2.x.2008, Leg. A.L. Montoya (CEUA); La Ceja, Alto de La Union, 6.016132°, 75.370182°, 2500m, 2males, 10-xi.2008, Leg. A.L. Montoya, Net (CEUA); Cundinamarca: Bogotá, Alto del Cable, 4.629103°, -74.049218°, 3100m, 30.vi.1978, Leg. M.L.J.M (ICN); Bogotá, Jardínes Barrio Palermo, 4.666300°, -74.068344°, 2600m, 6.viii.2007, Leg. C.A. Fernández & A. Fernández (ICN); Mosquera, UNAL Marengo lote 4, 4.680641°, -74.216488°, 2550m, 1female, 27.iv.2011, Leg. S. Guaqueta Net in Feijoba, Acca sellowiana (O.Berg) Burret (Myrtaceae) (UNAB). neotropica Curran, 1937:4 (Melanostoma). Type locality: Brazil: Sao Paulo. HT M (AMNH). Distr.: Colombia 460 · Zootaxa 4122 (1) © 2016 Magnolia Press MONTOYA (Antioquia, (Anori), Tolima), Argentina, Brazil. Refs.: Fluke, 1945:8, figs. 9 (head), 39 & 40 (abdomen); Fluke, 1958:278, figs. 117 & 118 (M genitalia). Material examined. COLOMBIA: Antioquia: Anori, Vereda El Roble, Reserva Natural Arrierito Antioqueño, 6.984806°, -75.111667°, 1600–1700m, 1male, 25–27.v.2012, Leg. Entomology Group, Net (CEUA). nigrans (Fluke), 1945:8 (Rhysops). Type locality: Brazil: Santa Catarina. Nova Teutonia. HT F (AMNH). Distr.: Colombia (Boyacá (Buenavista)). Refs.: Fluke, 1945, fig. 8 (head); Fluke, 1958:279, figs. Ill & 112 (M genitalia). Material examined. COLOMBIA: Boyacá: Buenavista, Vereda Patiño, 5.650832°, -74.286905°, 871m, 1female, 8.IV.2004, Leg. W. Avila, Net in ‘Granadilla’ Passiflora edulis Sims, 1818 (Passifloraceae) (UNAB). opaca (Fluke), 1945:10 (Rhysops). Type locality: Ecuador: Chimborazo, Urbina, Cerro Chimborazo, 3650m. HT M (AMNH). Distr.: Colombia (Antioquia (Belmira)), Ecuador. Refs.: Fluke,1945: 26, fig 15 (head profile of M); 1958: 279, figs. 115 & 116 (male genitalia). Material examined. COLOMBIA: Antioquia: Belmira, Paramo de Belmira, 6.707065°, -75.714136°, 3057m, 1male, 5.iii.2009, Leg. A.L. Montoya, Net. (CEUA). peruviana (Shannon), 1927:7 (Braziliana). Type locality: Peru, Rio Charape. HT M (USNM). Distr.: Colombia (Cauca (Morales)), Peru. Refs.: Shannon, 1927:7; Thompson et al. 1976:44 (cat.). Material examined. COLOMBIA: Cauca: Morales, El Pomorroso, Finca el Recuerdo, 2.701350°, 76.811080°, 1600m, 12.ix.2013, Leg. D. Caraballo, Manual. rex Fluke, 1945:21 (Melanostoma). Type locality: Ecuador: Hacienda Talahua, Bolivar. HT M (AMNH). Distr.: Colombia (Cundinamarca (La Palma)). Refs.: Thompson et al. 1976:44 (cat.). Material examined. COLOMBIA: Cundinamarca: La Palma, Murca, 5.329289°, -74.385364°, 1530m, 1male, ix.2010, Leg. J. Calderon, Net (UNAB). thiemei (Enderlein), 1938:201 (Braziliana). Type locality: Colombia. "Cordillera, tierra templada". HT M/F (ZMHU). Distr.: Colombia (Magdalena). Refs.: Thompson et al. 1976:44 (cat.). tropica (Curran), 1937:3 (Melanostoma). Type locality: Brazil, Sao Paulo, Campos do Jordao. HT M (AMNH). Distr.: Argentina, Brazil, Colombia (Antioquia (Andes, Girardota, Guarne)), Ecuador. Refs.: Curran, 1937: 3 (descr.); Thompson et al. 1976:44 (cat.). Material examined. COLOMBIA: Antioquia: Andes, Paramo de Santa Rita, 5.550249°, -75.988733°, 3600m, 8.xii.2010, 1female, Leg. M. Wolff (CEUA); Girardota, Vereda El Palmar, Secundary forest, 6.343502° 75.435102°, 2200m, 3female, 22-27.vii.2009, Leg. N. Uribe & G. Valencia Vsr. Viscera (CEUA); Guarne, Vereda Piedras Blancas, Parque Ecologico Piedras Blancas, cerca de quebrada, 6.291588°, -75.493345°, 2200m, 1female, 5.ii.2009, Leg. A.L. Montoya, Net (CEUA). Genus Leucopodella Hull Leucopodella Hull, 1948:94. Type species, Baccha lanei Curran (orig. des.). Refs.: Thompson et al. 1976:46 (cat.); Thompson, 1981:103 (key to the West Indies spp.) 183 (key to Neotropical spp.); Marinoni et al. 2007:149, 150 (desc, key, illustrations); Thompson & Skevington, 2014 (key and review of melanostomine genera groups). Xestoprosopa Hull, 1949c:94. Type species, Baccha delicatula Hull (orig. des.). delicatula (Hull), 1943:78 (Baccha). Type locality: Ecuador, Tungurahua, Baños. HT M (CNC). Distr.: Colombia (Antioquia (Medellin)), Costa Rica. Refs.: Hull, 1943:78, fig. 72 (abdomen); Thompson et al. 1976:46 (cat.); Thompson, 1999a:325 (key). Material examined. COLOMBIA: Antioquia: Medellín, 6.267404°, -75.585789°, 1570m, 1male, 23.x.2009, Leg. A.L. Montoya (CEUA). gracilis (Williston), 1891:34 (Baccha). Type locality: Mexico, Vera Cruz, Orizaba; Guerrero, Omiltemi. HT M/F CATALOGUE OF SYRPHIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 461 (BMNH). Distr.: Colombia (Antioquia (Caldas, Jardin, Medellín), Cundinamarca (Arbeláez), Huila (San Agustín), Quindio (Salento)), Ecuador, Mexico. Refs.: Curran, 1926:102; 1928:35; Wolcott, 1936:348 1848:463; Hull, 1943c:49, fig. 73 (abdomen); 1949a:94, 103, fig. 73, (abdomen) 186 (abdomen), 209 (wing); Thompson et al. 1976:46 (cat.); Thompson, 1981:103 (key and desc.) 184 (species key). Material examined. COLOMBIA: Antioquia: Caldas, Alto La Romera, 6.090000°, -75.638000°, 1560m, 1male, Leg. A.L. Montoya, Net (CEUA); Jardin, 5.583337°, -75.833332°, 2000m, 1male, 7.xi.2014, Leg. A.L. Montoya, Net (CEAU); Medellín, San Antonio de Prado, Vereda Yarumalito, 6.186412°, -75.658630°, 2300m, 1female, 7–8.xi.2008, Leg. C. Bota & N. Uribe, Net (CEUA); Cundinamarca: Arbeláez, 4.270965°, -74.419444°, 1417m, 1male, 23.iv.1994, Leg. N. León (UNAB); Huila: San Agustín, 1.927474°, -76.297857°, 1700m, 1male, 22.vii.2008, Leg. A.L. Montoya, Net (CEUA); Quindio: Salento, Vereda Canaan, 4.613714°, -75.581637°, 1800m, 1male, Leg. A.L. Montoya, Net (CEUA). Genus Platycheirus Lepeletier & Serville Cheilosia Panzer, 1809:14. Type species, Syrphus rosarum Fabricius, 1787 (mon.). To prevent confusion the ICZN should be asked to suppress Cheilosia Panzer to preserve existing usage of Cheilosia Meigen, 1822, and to place Platycheirus on the Official List of Generic Names. Platycheirus Lepeletier & Serville, 1828:513. Type species, Syrphus scutatus Meigen, 1822 (subs. des, Westwood, 1840:137). Refs.: Fluke, 1943 (key); Andersson, 1970:236–240 (Taxonomic notes); Vockeroth, 1990:660 (revisio of Neartical spp.); Thompson, 1972a:77–84 (desc. notes). Thompson, 1999a:325 (key); Marinoni et al. 2007:149, 150 (desc, key, illustrations); Thompson & Skevington, 2014 (key and review of melanostomine genera groups). Platychirus Agassiz, 1846:295, emend. Pyrophaena Schiner, 1860:21 3. Type species, Syrphus rosarum Fabricius, 1787 (orig. des.). Pachysphyria Enderlein, 1938:196. Type species, Scaeva ambigua Fallen, 1817 (orig. des.). Carposcalis Enderlein, 1938:199. Type species, Syrphus stegnus Say, 1829 (orig. des.). Subgenus Carposcalis Enderlein chalconota (Philippi), 1865:747 (Syrphus). Type locality: Chile, Santiago. HT ? (MNHNS). Distr.: Colombia (Cundinamarca), Argentina, Chile, Ecuador. Refs.: Fluke, 1945:15 (desc.), figs. 22 (head), 23 (antenna), 38 (abdomen); 1958:273, figs. 51–52 (M genitalia); Thompson et al. 1976:40 (cat.). ecuadoriensis (Fluke), 1945:16 (Melanostoma). Type locality: Ecuador, Imbabura. HT M (AMNH). Distr.: Colombia (Antioquia (Urrao), Cundinamarca (Bogotá, Facatativá)), Ecuador. Refs.: Fluke, 1945:16, figs. 25 (head), 59 (front tibia ofmales); Fluke, 1958:273, figs. 59 & 60 (M genitalia); Hull, 1949b:289, fig. 10c (head); Thompson et al. 1976:40 (cat.). Material examined. COLOMBIA: Antioquia: Urrao, Paramo del Sol, 6.416558°, -76.107506°, 3500m, 1female, 15.xi.2009, Leg. C. Bota (CEUA); Cundinamarca: Bogotá, 4.662568°, -74.080007°, 2600m, 10.v.1968, Leg. Wilches P. (ICN); Bogotá, Jardínes Barrio Palermo, 4.677811°, -74.101080°, 2650m, 20.vii.2007, Leg. C.A. Fernández & A. Fernández (ICN); Bogotá, Predios Universidad Nacional I.C.N., 4.638194°, -74.084046°, 2567m, 18.vii.1972, Leg. M. Arrieta (ICN); Bogotá, 7.ii.1978, Leg. C. Hernandez (ICN); Bogotá, Botanic Garden, 4.668332°, -74.100229°, 2551m, 20.ix.2009, Leg. M. Zamora (ICN); Facatativá, 4.814185°, -74.354995° 2600m, 1.i.2008, Leg. C.A. Fernández & A. Fernández (ICN); La Mesa, Lago Pedro Palo, 4.629429°, -74.469603°, 1200m, 22.xi.1970, Leg. E. Soler (ICN); La Calera, Vereda El Hato, Finca El Atardecer, 4.727495°, -73.907958°, 3000m, 1female, 6-10.i.2010, E. Ricaurte, Manua (CEUA); Risaralda: Pereira, Parque Natural Nacional los Nevados, 4.872737°, -75.426886°, 3000m, 4female, 6.v.2009, Leg. A. Bustamante (CEUA). fenestrata (Macquart), 1842:163 (103) (Syrphus). Type locality: "Chile". HT F (MNHNP). Distr.: Colombia (Antioquia (Belmira, Bello, La Ceja, Urrao), Cundinamarca (Bogotá, Chingaza, Subachoque), Brazil, Chile, Mexico. Refs.: Macquart, 1842:163 (103), tab. 17, fig. 6 (habitus), 6a (head); Macquart, 1850:458 (154), tab. 14, fig. 13 (head); Fluke, 1945:15 (desc. notes), figs. 21 (head), 57 (male front tibia); Fluke, 1958:273, figs. 56 & 64 (M genitalia); Thompson et al. 1976:40 (cat.). Material examined. COLOMBIA: Antioquia: Belmira, Paramo de Belmira, 6.707065°, -75.714136°, 462 · Zootaxa 4122 (1) © 2016 Magnolia Press MONTOYA 3057m, 1female, 5.iii.2009, Leg. A.L. Montoya (CEUA); Bello, San Felix, Paramo de Las Baldias, 6.332653°, 75.646599°, 3050m, 1male, 17.ii.2013, Leg. A.L. Montoya, Net (CEUA); La Ceja, Alto la Union, Forest, 6.016132°, -75.370182°, 2500m, 2male, 10.ix.2008, Leg. A.L. Montoya, Net (CEUA); Urrao, Paramo del Sol, 6.416558°, -76.107506°, 3500m, 8female, 3male, 8.ix.2009, Leg. C. Bota (CEUA); Cundinamarca: Bogotá, Alto del cable, 4.629103°, -74.049218°, 3100m, 30.vi.1978, Leg. M.L.M.J (ICN); Bogotá, 1.v.1968, Leg. Aguirre (ICN); Bogotá, Jardín de casa, 4.668332°, -74.100229°, 2600m, 4.iv.1976, Leg S. Zuluaga. (ICN); Bogotá, Botanic Garden, 4.668332°, -74.100229°, 2600m, 18.x.1989, Leg. I. Arévalo (ICN); 25.vii.2009, Leg. M. Zamora (ICN); Chingaza, Estacion Monte Redondo, 4.481898°, -73.721867°, 3036m, 1male, 1female, 19.ix.2002, Leg. E. Martinez, decaying Pig (CEUA); Subachoque, Paramo de El Tablazo, 5.013671°, -74.184844°, 3400m, 7.vi.2008, Leg. C.A. Fernández & A. Fernández (ICN). hesperius (Vockeroth), 1990:696. Type locality: Alabhama, CA, USA. HT M (USNM). Distr.: Colombia (Antioquia (Andes), Cundinamarca (Chingaza), Mexico. Refs.: Vockeroth, 1990:696 (cat. and desc.). Material examined. COLOMBIA: Antioquia: Andes, Paramo de Santa Rita, 5.550249°, -75.988733°, 3600m, 8.xii.2010, 1male, Leg. A.L. Montoya (CEUA); Cundinamarca: Chingaza, Estacion Monte Redondo, 4.481898°, -73.721867°, 3036m, 3female, 19.ix.2002, Leg. E. Martinez, decaying Pig (CEUA). Genus Talahua Fluke Melanostoma, subg, Talahua Fluke, 1945:22. Type species, fervidum Fluke (orig. des.). Refs.: Fluke, 1945:23 (key); Fluke, 1958:279, fig 123 (M genitalia); Thompson et al. 1976:45 (cat.); Thompson, 1999a:325 (key), 338; Thompson & Skevington, 2014 (key and review of melanostomine genera groups). fervida (Fluke), 1945:23 (Melanostoma). Type locality: Ecuador, Bolivar, Hacienda Talahua. HT M (AMNH). Distr.: Colombia (Antioquia, Boyacá, Quindio), Ecuador. Refs.: Fluke, 1945:23 (desc, key), figs. 34 (head), 35 (M genitalia), 53 (abdomen); Fluke, 1958:279, fig. 123 (M genitalia); Thompson et al. 1976:45 (cat.); Thompson, 1999a:325–338 (key and taxonomic notes). Material examined. COLOMBIA: Antioquia: San Jose de la Montaña, Vereda el Congo sector la Laguna, Potrero, 6.804244°, -75.689613°, 3100–3200m, 1female, 10–14.ix.2011, Leg. L. Rios, Net (CEUA). Genus Tuberculanostoma Fluke Tuberculanostoma Fluke, 1943:425. Type species, antennatum Fluke (orig. des.). Ref. Fluke, 1943 (key); Thompson & Skevington, 2014 (key and review of melanostomine genera groups). antennatum Fluke, 1943:426. Type locality: Ecuador, Bolivar, Hacienda Talahua. HT M (AMNH). Distr.: Colombia (Antioquia (Liborina, Urrao), Cundinamarca (Chingaza), Risaralda (Pereira)), Ecuador, Peru, Venezuela. Refs.: Fluke, 1943:426, figs. 1–2 (head), 3 (antenna); Thompson et al. 1976:45 (cat.). Material examined. COLOMBIA: Antioquia: Liborina, Vereda El Playon, Finca La Pava, 6.665323°, 75.719095°, 3000-3197m, 1female, 5-8.xi.2011, Leg. J.D. Castaño & C. Bota, Net (CEUA); Urrao, Paramo del Sol, 6.416558°, -76.107506°, 3500m, 8female, 5male, 8.ix.2009, Leg. C. Bota (CEUA); Cundinamarca: Chingaza, Estacion Monte Redondo, 4.481898°, -73.721867°, 3036m, 3female, 19.ix.2002, Leg. E. Martinez, decaying Pig (CEUA); Risaralda: Pereira, Parque Natural Nacional los Nevados, 4.872737°, -75.426886°, 3000m, 3000m, 1female, 6.v.2009, Leg. A. Bustamante (CEUA). pectinis Fluke, 1943:430. Type locality: Ecuador, Bolivar, Hacienda Talahua. HT M (AMNH). Distr.: Colombia (Antioquia (Urrao), Cundinamarca), Ecuador. Refs.: Fluke, 1943:430, figs. 6 (head), 9 (profemur); Thompson et al. 1976:45 (cat.). Material examined. COLOMBIA: Antioquia: Urrao, Paramo del Sol, 6°1'43.24"N, 76°7'3.29"W, 3200m, 1female, 8.ix.2009, Leg. C. Bota (CEUA). CATALOGUE OF SYRPHIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 463 Genus Xanthandrus Verral Xanthandrus Verrall, 1901:316. Type species, Musca comptus Harris (Coquillett, 1910:620). Refs.: Refs.: Fluke, 1937 (key); Thompson, 1981:105 (taxonomic notes); Thompson, 1999a:325 (key); Borges & Pamplona, 2003:155–167 (rev. and desc.); Marinoni et al. 2007a:149, 150 (desc, key, illustrations); Thompson & Skevington, 2014 (key and review of melanostomine genera groups). Subgenus Xanthandrus Verral bucephalus (Wiedemann), 1830:126 (Syrphus). Type locality: "Brazil". HT M (ZMHU). Distr.: Colombia (Antioquia (La Pintada), Sucre (Chengue), Brazil, Peru. Refs.: Verral, 1901:317; Fluke, 1936:62, 63 (key and citation); Fluke, 1937:6 (key); Fluke, 1958:279, figs. 119–120 (M genitalia); Thompson et al. 1976:44 (cat.); Borges & Pamplona, 2003:158 (desc. and illustrations); Reemer, 2010:194 (desc, notes), fig. 18 (habitus,males). Material examined. COLOMBIA: Antioquia: La Pintada, 5.736521°, -75.632087°, 650m, 1male, 26.v.2007, Leg.N. Uribe, Vsr. Fish (CEUA); Sucre: Chengue, 9.614540°, -75.398641°, 53m, Leg. C. Elster (ICN). mellinoides (Macquart), 1846:265 (137) (Syrphus). Type locality: "Colombia". HT M (UMO). Distr.: Colombia, Mexico, Venezuela. Refs.: Thompson et al. 1976:45 (cat.); Borges & Pamplona, 2003:160 (desc. and illustrations). euceratus Bigot, 1884:82 (Melanostoma?) (description). plaumanni Fluke, 1937:8. Type locality: Brazil, Santa Catarina, Nova Teutonia. HT F (AMNH). Distr.: Colombia (Boyacá, Valle del Cauca), Brazil. Refs.: Fluke, 1937:8, fig. 4 (abdomen); Thompson et al. 1976:45 (cat.); Borges & Pamplona, 2003:164 (desc. and illustrations). Tribe Syrphini Genus Allograpta Osten Sacken Allograpta Osten Sacken, 1875:49. Type species, Scaeva obliqua Say (mon.). Refs.: Curran, 1928:36 (key to Puerto Rican spp.); Vockeroth, 1969:126–1130 (desc.); Vockeroth, 1973 (syn.); Thompson, 1981:23 (key); Thompson, 1999a:327, 328 (key); Thompson et al. 2000:54 (Key to the Costarican spp.); Marinoni et al. 2007a:147, 150 (desc, key, illustrations); Mengual et al. 2009:12 (key and phylogenetic rev.); Thompson 2012 (nomenclatural change, syns.). Neoscaeva Frey, 1946:170. Type species, Syrphus aeruginosifrons Schiner (orig. des.). Microsphaerophoria Frey, 1946:168. Type species, plaumanni Frey (orig. des.). Miogramma Frey, 1946:165. Type species, Syrphus javanus Wiedemann (orig. des.). Helenomyia Bankowska, 1962:311. Type species, Syrphus javanus Wiedemann (orig. des.). Paraxanthogramma Tao & Chiu, 1971:74. Nomen nudum based on Syrphus javanus Wiedemann as nakamurae Matsumura. annulipes (Macquart), 1850:464 (160) (Paragus). Type locality: Brazil, Bahia. HT M (MNHNP). Distr.: Colombia (Cundinamarca (Bogotá), Risaralda (Santa Rosa de Cabal)), Bolivia, Brazil, Ecuador, Peru. Refs.: Macquart, 1850:464 (160), pi. 14, fig. 18 (head); Fluke, 1942:16, fig. 34 (abdomen); Thompson et al. 1976:33 (cat.); Reemer, 2010:169 (desc, and notes), fig. 8 (habitus). Material examined. COLOMBIA: Cundinamarca: Bogotá, 4.596100°, -74.084879°, 2599m, Net (ICN); Risaralda: Santa Rosa de Cabal, Termales de Santa Rosa, 4.837708°, -75.548600°, 2000m, 1male, 10.vi.2013, Leg. A.L. Montoya, Net (CEUA). aperta Fluke, 1942:19. Type locality: Guyona, Kaieteur, Savannah. HT M (AMNH). Distr.: Colombia (Cundinamarca (Bogotá)), Guyana, Suriname. Refs.: Fluke, 1942:19, fig. 48 (abdomen); Thompson et al. 1976:34 (cat.); Reemer, 2010:171 (desc, and notes), fig. 9 (habitus). Material examined. COLOMBIA: Cundinamarca: Bogotá, 4.596100°, -74.084879°, 2599m, 7males, 5females, Net (ICN). 464 · Zootaxa 4122 (1) © 2016 Magnolia Press MONTOYA bilineella Enderlein, 1938:219. Type locality: Argentina, Prov. Buenos Aires. HT M (ZMHU). Distr.: Colombia, Argentina. Refs.: Thompson et al. 1976:34 (cat.). exotica (Wiedemann), 1830:136 (Syrphus). Type locality: "Brazil". HT ? (NMW). Distr.: Colombia (Antioquia (Medellín), Boyacá (Aquitania, Sogamoso), Cundinamarca (Bogotá, Cajicá, Chipaqué, La Vega, La Esperanza, Machetá, Mosquera, Sibaté, Soacha, Suba, Subachoque, Usme, Vianí), Boyacá, Meta (Acacías)), Argentina, Brazil, Costa Rica. Refs.: Fluke, 1942:24, fig. 47 (abdomen); 1950a:129, figs. 136 & 138 (M genitalia); Thompson et al. 1976:34 (cat.); Reemer, 2010:171 (desc, and notes), fig. 10 (habitus). bilineata Enderlein, 1938a:220. Material examined. COLOMBIA: Antioquia: Medellín, Universidad de Antioquia, 6.266375°, 75.568989°, 1465m, 1male, Leg. A.L. Montoya (CEUA); Boyacá: Aquitania, Laguna de la Tota, 5.583541°, 72.900514°, 3050m, 16.x.1999 (UNAB); Sogamoso, 5.722996°, -72.923443°, 2549m, 06.iv.2004, Leg. M. Sánchez, Net (UNAB); Cundinamarca: Fontibón, Parque Centenario, cerca de caño, 4.685171°, -74.151528°, 2549m, 31.viii.2003, Leg. E. Ortíz, Net (UNAB); Bogotá, Botanic Garden, 4.668332°, -74.100229°, 2551m 18.viii.2009, Leg. M. Zamora (ICN); Bogotá, UNal I.C.N, 4.638194°, -74.084046°, 2567m, 24.xii.1971, Leg. Fernández–A., J.L. Fernández–C (ICN); Cajicá, Vereda El Canelo, 4.920434°, -74.027161°, 2564m, 04.xi.2000, Leg. A. Morales., H. Monroy (UNAB); Chipaqué, 4.450001°, -74.050000°, 2479m 18.iv.2004, Leg. Z. Muñoz, Net (UNAB); La Esperanza, 4.269725°, -74.431312°, 2599m, 09.v.2004, Leg. A. Sepúlveda, Net (UNAB); La Vega, 4.999916°, -74.350009°, 1215m, 12.xi.1995, Leg. Gallego (UNAB); Machetá, 5.080217°, -73.607475°, 2100m, Km. 21, 2110m, 23.xi.2003, Leg. A. Rodríguez (UNAB); Mosquera, Finca Marengo, 4.680641°, -74.216488°, 2554m, 15.xi.2003, Leg. D. Gracía. J. Díaz, Net (UNAB); Sibaté, 4.483597°, -74.266475°, 2180m, 01.v.2000, Leg. N. Santana, M. Mazorra & J. Espinosa (UNAB); Soacha, Vereda Las Piedras, 4.574535°, -74.228384°, 2566m, 06.xi.1999 (UNAB); Suba, 4.757031°, -74.088591°, 2599m 18.xii.1988, Leg. Ospina (UNAB); Subachoque, Finca Potosi, 4.932714°, -74.183370°, 2663m, 06.ix.2003, Leg. L. Bermúdez (UNAB); Usme, 4.522906°, -74.106378°, 2790m, 08.xi.2004, Leg. A. Reyes & G. Villeta, Net (UNAB); Vianí, 4.883333°, -74.566667°, 1498m, 25.v.2004, Leg. G. Velandía & J. Romero, Net (UNAB); Meta: Acacías, 3.987806°, -73.759747°, 523m, 29.xi.1997, Leg. Aranguren (UNAB). falcata Fluke, 1942:16. Type locality: Brazil, Santa Catarina, Nova Teutonia. HT M (AMNH). Distr.: Colombia (Antioquia (Medellín)), Costa Rica, Ecuador, Peru, Venezuela. Refs.: Fluke, 1942:16, fig. 36 (abdomen); Thompson et al. 1976:35 (cat.). Material examined. COLOMBIA: Antioquia: Medellín, San Antonio de Prado, Vereda Yarumalito, 6.186412°, -75.658630°, 2300m, 1male, 7–8.xi.2008, Leg. C. Bota & N. Uribe, Net (CEUA). hastata Fluke, 1942:17. Type locality: Brazil, Sao Paulo. HT M (AMNH). Distr.: Colombia (Valle del Cauca), Brazil, Peru. Refs.: Fluke, 1942:17, fig. 37 (abdomen); Thompson et al. 1976:35 (cat.). neotropica Curran, 1936b:14. Type locality: Colombia, Popayan. HT M (AMNH). Distr.: Colombia (Antioquia, Boyacá (Gachetá, Ramiriqui), Cauca (Popayán), Cundinamarca (Bogotá, Cota, Facatativá, Machetá, Pulí, Ubaque), Magdalena, Tolima (Ambalema), Valle del Cauca (Buga, Palmira)), Argentina, Brazil, Costa Rica, Ecuador, Hispaniola, Jamaica. Refs.: Fluke, 1942:24, fig. 49 (abdomen); 1950a:141, fig. 41 (M genitalia); Vockeroth, 1969:130 (citation); Thompson et al. 1976:36 (cat.). Material examined. COLOMBIA: Boyacá: Gachetá, Vereda Salinas, 4.820836°, -73.636574°, 1850m, 07.iv.2004, Leg. O. Lanchero, Net (UNAB); Ramiriqui, Vereda Escobal, 5.399818°, -73.333428°, 2500m, 18.xi.2003, Leg. D. Fonseca (UNAB); Cundinamarca: Bogotá, Parque La Florida, 4.732002°, -74.146014°, 2600m, 10.xi.2001, Leg. E. Peláez, Net (UNAB); Cota, Finca Peñas Blancas, 4.812890°, -74.106085°, 2547m, 13.i.2005, D. Carrillo Leg., Malaise trap (UNAB); Facatativá, Finca Luisa María, 4.814821°, -74.346605°, 2600m 18.ii.2005, Leg. C. Jara, Net (UNAB); Machetá, El Caucho, 5.080217°, -73.607475°, 2100m, 12.x.2003, Leg. F. Jiménez, Net (UNAB); Pulí, Vereda El Manantial, 4.687593°, -74.705016°, 1200m, 12.iii.1998, Leg. C. Jiménez (UNAB); Ubaque, Vereda Romero Alto, Finca La Esperanza, 4.484418°, -73.934194°, 2320m, 12.iv.2008, Leg. J.C. Almaciga, Net (UNAB); Tolima: Ambalema, Vereda Pajonales, Hacienda Pajonales, 4.783333°, -74.766667°, 850m, 09.iv.2001, Leg. J. Esguerra (UNAB); Valle del Cauca: Buga, 3.900000°, -76.300000°, 1067m, 10.v.2004, CATALOGUE OF SYRPHIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 465 Leg. W. Olaya & P. Villa, Manual (UNAB); Palmira, Unipalma, 3.587073°, -76.248936°, 1059m, 25.xi.2002, Leg. A. Farias & O. Mateus, Net (UNAB). obliqua (Say), 1823:89 (Scaeva). Type locality: "United States". HT ? Lost. Distr.: Colombia (Antioquia (Medellín), Cundinamarca (Anolaima), Risaralda, Tolima (Espinal)), Argentina, Costa Rica, Jamaica. Refs.: Osten Sacken, 1878:126 (syn.); Fluke, 1942:24, fig. 46 (abdomen); Vockeroth, 1969:165, fig. 85 (M genitalia); Thompson et al. 1976:36 (cat.); Thompson,1981:25, fig. 20 (thorax, lateral view), 28 (wing). Material examined. COLOMBIA: Antioquia: Medellín, 6.267404°, -75.585789°, 1570m, Universidad de Antioquia, 1male, 1female, 11-xi.2008, Leg. A.L. Montoya, Net (CEUA); Cundinamarca: Anolaima, La Esmeralda, 4.766197°, -74.467662°, 1656m, 21.ix.2003, Leg. C. Matiz, Net (UNAB); Tolima: Espinal, 4.162687°, -74.874972°, 322m, 16.viii.1974, Leg. G. Daza (UNAB). quadricincta Enderlein, 1938:214. Type locality: Colombia, Bogota. HT F (ZMHU). Distr.: Colombia (Cundinamarca). Refs.: Thompson et al. 1976:37 (cat.). tectiforma Fluke, 1942:18, fig, 45 (abdomen). Type locality: Ecuador, Imbabura Cuicocha, 3200 ft. HT M (AMNH). Distr.: Colombia (Antioquia (Bello, Betulia, Urrao)), Ecuador. Refs.: Fluke, 1942:18, fig, 45 (abdomen); Fluke, 1950a: l29, fig. 145 (M genitalia); Thompson et al. 1976:38 (cat.). Material examined. COLOMBIA: Antioquia: Bello, San Felix, Paramo de Las Baldias, 6.332653°, 75.646599°, 3050m, 1female, 17.ii.2013, Leg. A.L. Montoya, Net (CEUA); Betulia, Vereda el Indio, La Mina, Alto de San José, Paramo, 5.771994°, -76.086152°, 3284-3300m, 1male, 14-18.xii.2013, Leg. C. Henao & J. Cogollo (CEUA); Urrao, Paramo del Sol, 6.416558°, -76.107506°, 3500m, 8.ix.2009, Leg. A. Franco, Malaise trap (MEFLG). teligera Fluke, 1942:18. Type locality: Brazil, Santa Catarina, Nove Teutonia. HT F (AMNH). Distr.: Colombia (Antioquia (Jardín), Cundinamarca), Brazil, Costa Rica, Ecuador. Refs.: Fluke, 1942:18, fig. 44 (abdomen); Thompson et al. 1976:38 (cat.). Material examined. COLOMBIA: Jardín, 5.578169°, -75.829912°, 1972m, 1female 18.v.2006, Leg. J. Cardona (CEUA). Genus Calostigma Shannon, 1927 Calostigma Shannon, 1927:8. Type species, Calostigma elnora Shannon, 1927 (orig. des.). Refs.: Curran, 1930. Callostigma Curran, 1930c:8. Misspelling. Callistigma Hull, 1944b:41. Misspelling. elnora species group ornatipes (Curran), 1927:3 (Baccha). Type locality: Puerto Rico, Cayey. HT M (AMNH). Distr.: Colombia (Antioquia (San Roque)), Puerto Rico. Refs.: Hull, 1949c:232, fig. 194 (abdomen), 260, fig. 306 (wing); Thompson et al. 1976:24 (cat.); Thompson, 1981:34, fig 39 (head, lateral view), 48 (leg andmales genitalia complex), 65 (wing). Material examined. COLOMBIA: Antioquia: San Roque, 6.503175°, -74.926885°, 1800m, 1female, 8.ii.2009, Leg. A.L. Montoya (CEUA). Genus Claraplumula Shannon Claraplumula Shannon, 1927:8. Type species, latifacies Shannon (orig. des.). Refs.: Thompson, 2012 (generic status). latifacies (Shannon), 1927:8 (Claraplumula). Type locality: Peru, Tincochaca, 7000 ft. HT M (USNM). Distr.: 466 · Zootaxa 4122 (1) © 2016 Magnolia Press MONTOYA Colombia (Antioquia (Copacabana, Jardin)), Ecuador, Peru. Refs.: Fluke, 1942:22, figs. 4 (head), 5 (abdomen); 1950a:127, figs. 127 & 125 (M genitalia); Vockeroth, 1969:165, fig. 86 (M genitalia); Thompson et al. 1976:36 (cat.). Material examined. COLOMBIA: Antioquia: Copacabana, Vereda El Cabuyal, 6.345579°, -75.503714°, 1450m, 1male, Leg. Velez & Marin, Vsr. Fish (MEPB); Jardin, Vereda Florestar-El Cañon, Finca Los Robles 5.583337°, -75.833332°, 2000m, 1female, 28-abr-2001, Leg. G. Etnomology, Net (CEUA). Genus Dasysyrphus Enderlein Dasysyrphus Enderlein, 1938:208. Type species, Scaeva albostriata Fallen (orig. des.). Refs.: Vockeroth, 1969:73 (description, nomenclatural notes); Thompson et al. 1976:10 (cat.), Thompson, 1999a:326 (key); Locke & Skevington, 2013: 1–80 (Rev. of Neartic species). Conosyrphus Matsumura 1918: 11 (name preocc. by Conosyrphus Frey, 1915). Type species Conosyrphus okunii Matsumura, 1918 (orig. des.) Syrphella Goffe, 1944: 129. Type species, Scaeva tricincta Fallén, 1817 Dendrosyrphus Dusek & Laska, 1967: 365. Type species, Syrphus lunulatus Meigen, 1822 lotus (Williston), 1887b:75 (Syrphus). Type locality: U.S.A, Arizona. HT F (USNM). Distr.: Colombia (Antioquia (Betulia), Valle del Cauca), Ecuador, Mexico. Refs.: Fluke, 1942:3; Thompson et al. 1976:10 (cat.); Mengual et al. 2008:7 (distrib. record). Material examined. COLOMBIA: Antioquia: Betulia, Vereda el Indio, La Mina, Alto de San José, Paramo, 5.771994°, -76.086152°, 3284-3300m, 3male, 14-18.xii.2014, Leg. C. Henao & J. Cogollo (CEUA). Genus Eupeodes Osten Sacken Eupeodes Osten Sacken, 1877:328. Type species, Eupeodes volucris Osten Sacken (mon). Refs.: Thompson et al. 1976:9 (cat.); Thompson, 1999a:328 (key). Posthonia Enderlein, 1938:203. Type species, longipenis Enderlein (orig. des.). volucris Osten Sacken, 1877:329. Type locality: U.S.A, California, Los Angeles, Marin County, Yosemite; Nevada; Utah; Colorado, Denver, Clear Creek, Spanish Peaks. HT M/F (MCZ). Distr.: Colombia (Cundinamarca), Mexico. Refs.: Vockeroth, 1969:150, fig. 34 (M genitalia); Thompson et al. 1976:9 (cat.). longipenis Enderlein, 1938:203 (Posthonia). Genus Fazia Shannon Fazia Shannon, 1927:25. Type species, bullaephora Shannon (orig. des.) = decemmaculata (Rondani). Refs.: Thompson, 2012 (generic status). Chasmia Enderlein, 1938a:213 (name preocc. by Enderlein, 1922). Type species, hians Enderlein (orig. des.). Metepistrophe Hull, 1949b:293 (Proposed as a subg. of Epistrophe). Type species, Epistrophe altissima Fluke (orig. des.). Misidentified Type species, Originally named type was Epistrophe remigis Fluke, but Fluke (1951) noted that the specimen used by Hull was mislabelled by him and was actually altissima Fluke. Hence, under Article 70.3 of the International Code of Zoological Nomenclature (ICZN 1999), Mengual et al. 2009 fixed the Type species, of Metepistrophe Hull to the taxonomical species, altissima Fluke. Metallograpta Hull, 1949b:293 (Proposed as a subg. of Epistrophe). Type species, Allograpta colombia Curran (orig. des.). alta Curran, 1936b:15. Type locality: Colombia, Popayan. HT M (AMNH). Distr.: Colombia (Cauca, Cundinamarca), Ecuador. Refs.: Fluke, 1942:23, fig. 31 (abdomen); Fluke, 1950a:129, fig. 134 (M genitalia); Thompson et al. 1976:33 (cat.). flavomaculata Hull, 1937a:169. colombia Curran, 1925c:349. Type locality: Colombia, Sierra Quemada. HT F (CUI). Distr.: Colombia CATALOGUE OF SYRPHIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 467 (Magdalena), Brazil, Costa Rica, Ecuador. Refs.: Fluke, 1942:12 (desc.), figs. 25 (head), 26 (abdomen); 1950a:129, fig. 135 (M genitalia); Thompson et al. 1976:34 (cat.). fasciata Curran, 1932:4. Type locality: "Peru". HT F (AMNH). Distr.: Colombia (Magdalena), Ecuador, Peru. Refs.: Fluke, 1942:23, figs. 28 (head), 29 (abdomen); 1950a:129, fig. 137 (M genitalia); Thompson et al. 1976:35 (cat.). bisinterrupta Enderlein, 1938:210 (Fazia). bisinterrpta Thompson et al. 1976:34. Misspelling. fascifrons (Macquart), 1846:265 (Syrphus). Type locality: "Nouvelle Grenade". HT M (UMO). Distr.: Colombia (Magdalena, Valle del Cauca). Refs.: Macquart, 1846:265 (137), pi. 11, fig. 11 (wing); Thompson et al. 1976:35 (cat.). imitator (Curran), 1925c:351 (Epistrophe). Type locality: Colombia, Vista Nieve, San Lorenzo Mt. HT M (CUI). Distr.: Colombia (Antioquia (Alejandria, Bello, Caldas, Jardin), Caldas, Cundinamarca, Magdalena (Santa Marta), Quindio (Salento)), Bolivia, Costa Rica. Refs.: Fluke, 1942:23, figs. 20 (head), 21 (abdomen); Thompson et al. 1976:36 (cat.). Material examined. COLOMBIA: Antioquia: Alejandria, 6.401714°, -75.058838°, 1200m, 1male, 8.xi.07, Leg. A.L. Montoya, Manual (CEUA); Bello, San Felix, Paramo de Las Baldias, 6.332653°, -75.646599°, 3050m, 2female, 17.ii.2013, Leg. A.L. Montoya, Net (CEUA); Caldas, Alto La Romera, 6.090000°, -75.638000°, 1560m, 1male, 2females, Leg. A.L. Montoya, Net (CEUA); Jardin, Alto del Indio, Finca Pipinta, 2females, 14-18.xii.2013, Leg. S. Ruiz (CEUA); Quindio: Salento, Vereda Canaan, 4.613714°, -75.581637°, 1800m, 1male, Leg. A.L. Montoya, Net (CEUA). nasigera (Enderlein), 1938:212 (Fazia). Type locality: "Colombia". HT M/F (ZMHU). Distr.: Colombia. Refs.: Thompson et al. 1976:36 (cat.). roburoris (Fluke), 1942:11 (Epistrophe). Type locality: Ecuador, Bolivar, Hacienda Talahua. HT M (AMNH). Distr.: Argentina, Colombia (Antioquia (Belmira)), Ecuador. Refs.: Fluke, 1942:11, figs. 18 (abdomen) 18 (head); Thompson et al. 1976:37 (cat.). Material examined. COLOMBIA: Antioquia: Belmira, Morro Cabildo Verede, 6.6579, -75.672, 3200m, 1female, 30.ix.2011, Leg. J.D. Castaño & C. Bota, Malaise trap (CEUA). rostrata (Bigot), 1884:102 (Sphaerophoria). Type locality: Mexico. HT F (UMO). Distr.: Colombia (Valle del Cauca), Costa Rica, Mexico. Refs.: Thompson et al. 1976:37 (cat.). similis Curran, 1925c:350. Type locality: Colombia, Vista Nieve. HT F (CUI). Distr.: Colombia (Magdalena), Argentina, Brazil, Costa Rica. Refs.: Fluke, 1942:23, fig. 27 (abdomen); 1950a:129, fig. 144 (M genitalia); Thompson et al. 1976:37 (cat.). strigifacies (Enderlein), 1938:211 (Fazia). Type locality: "Brazil". HT M/F (ZMHU). Distr.: Colombia (Antioquia (Bello), Valle del Cauca), Brazil. Refs.: Thompson et al. 1976:37 (cat.). Material examined. COLOMBIA: Antioquia: Bello, San Felix, Paramo de Las Baldias, 6.332653°, 75.646599°, 3050m, 1female, 17.ii.2013, Leg. A.L. Montoya, Net (CEUA). Genus Hermesomyia Vockeroth Hermesomyia Vockeroth, 1969:121. Type species, Hermesomyia bacchiformis Vockeroth, 1969 (orig. des.) = Baccha wulpiana Lynch Arribalzaga, 1891. wulpiana (Lynch), 1891:250 (1892c :47) (Baccha, nom. nov. for tricinctus Wulp). Type locality: Argentina, Prov. 468 · Zootaxa 4122 (1) © 2016 Magnolia Press MONTOYA Tucuman. HT M (BMNH). Distr.: Colombia (Antioquia (Guarne)), Argentina, Brazil, Costa Rica, Ecuador. Refs.: Wulp, 1888:376, pi. 10, fig. 8 (habitus); Kertész, 1910:165 (cat.); Hull, 1943b:50 (cat.); Hull, 1949:126 (key ref., abdomen pattern); Fluke, 1956:259 (cat.), 267 (cat.); Vockeroth, 1969:122, figs. 13 18; Thompson et al. 1976:25 (Syn.); Rotheray et al. 2000:390 (desc, and syn.), fig. 8–12 (illustrations of larval structures). Material examined. COLOMBIA: Antioquia: Guarne, Vereda Piedras Blancas, 6.291588°, -75.493345°, 2660m, 1female, Leg. Entomology Group, rearing from Bromeliad (MEPB). Genus Hybobathus Enderlein Hybobathus Enderlein, 1938:233. Type species, Hybobathus quadrilineatus Enderlein, 1938 (orig. des.). Hypobathus Fluke, 1956:217. Misspelling. Callisyrphus Frey, 1946:154. Type species, Syrphus rubricosus Wiedemann, 1830 (orig. des.). Calliscaeva Frey, 1946:171. Incorrect original spelling of Callisyrphus (rev. by Fluke, 1956:199). Aulacibaccha Hull, 1949a:96 Type species, Baccha titan Hull, 1947a (orig. des.) = Baccha arx Fluke, 1936. (Proposed as a subg. of Baccha). arx species group arx (Fluke), 1936:60 (Baccha). Type locality: Brazil, "Cois Parma Cos". HT F (AMNH). Distr.: Colombia ((Alejandria, Fredonia, Medellín, San Roque), Quindio (Filandia) Risaralda (Pereira)), Brazil, Costa Rica. Refs.: 1936:60, fig. 2 (abdomen), 3 (wing); Hull, 1949c:248, figs. 259–261 (abdomens); 274, fig. 361 (wing); Thompson et al. 1976:13 (cat.). Material examined. COLOMBIA: Antioquia: Alejandria, 6.401714°, -75.058838°, 1200m, 1male, 8.xi.07, Leg. A.L. Montoya, Vsr. Fish (CEUA); Fredonia, 5.916573°, -75.683347°, 1010m, 1male, Leg. Entomology Group, Net (CEUA); Medellín, Universidad Nacional de Colombia, 6.267404°, -75.585789°, 1570m, 2males, 10.i.2009, Leg. A.L. Montoya, larvae preying nymphs of Antianthe expansa Germar 1835 (Membracidae) (CEUA); San Roque, San Jose del Nus, 6.503175°, -74.926885°, 830m, 1male, 5.i.2007, Leg. N. Uribe, Vsr. Liver (CEUA); Quindio: Filandia, Hacienda Bengala, 4.685291°, -75.617533°, 2000m, 1male, 1female, 10.ix.2008, Leg. A.L. Montoya, Net (CEUA); Risaralda: Pereira, Wildlife Sanctuary Otún Quimbaya, 4.728880°, 75.577850°, 1890m, 1female, v.2001, Leg. H. Vargas, Vsr. Fish (CEUA). flavipennis (Wiedemann), 1830:123 (Syrphus). Type locality: "Brazil". HT M/F (ZMHU). Distr.: Colombia (Antioquia (San Roque)), Brazil, Ecuador, Peru. Refs.: Hull, 1949c:204, fig. 65 (abdomen); Thompson et al. 1976:18 (cat.); Reemer, 2010:179 (desc.), fig 62–63 (habitus, males & female). Material examined. COLOMBIA: Antioquia: San Roque, San Jose del Nus, 6.503175°, -74.926885°, 1800m, 5.i.2007, Leg. N. Uribe (CEUA). lineatus (Macquart), 1846:267 (139) (Baccha). Type locality: Mexico, Yucatan. HT F (MNHNP). Distr.: Colombia (Antioquia (Nariño)), Bahamas, Jamaica. Refs.: Macquart, 1846:267 (139), tab. 20, fig. 5 (habitus); Hull, 1949c:236, fig. 206–207 (abdomen); Thompson, 1981:54; Thompson et al. 1976:21 (cat.). Material examined. COLOMBIA: Antioquia: Nariño, 5.631484°, -75.245454°, 2400m, 1male, 24.vii.2010, Leg. V. Pérez (CEUA). lividus (Schiner), 1868:343 (Baccha). Type locality: "South America". HT M/F (NMW). Distr.: Colombia (Antioquia (San Roque), Sucre (Chengue)), Brazil, Guyana, Honduras, Mexico, Panama, Peru, Trinidad. Refs.: Hull, 1949c:196, fig. 26 (abdomen), 281, fig. 124 (abdomen), 268, fig. 336 (wing); Thompson et al. 1976:21 (cat.). Material examined. COLOMBIA: Antioquia: San Roque, San Jose del Nus, 6.503175°, -74.926885°, 1800m, 5.i.2007, Leg. N. Uribe (CEUA); Sucre: Chengue, 9.614517°, -75.398629°, 53m, Leg. C. Elster (ICN). norina (Curran), 1941:267 (Baccha). Type locality: Brazil, Santa Catarina, Nova Teutonia. HT M (AMNH). Distr.: Colombia (Valle del Cauca), Argentina, Bolivia, Brazil. Refs.: Hull, 1949c:196, fig. 23 (abdomen), 218, fig. 131 (abdomen), 270, fig. 347 (wing); Thompson et al. 1976:23 (cat.); Reemer, 2010:182 (desc.), fig 61 (habitus). CATALOGUE OF SYRPHIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 469 persimilis (Curran), 1930a:2 (Baccha). Type locality: Brazil, Mato Grosso, Chapada. HT F (AMNH). Distr.: Colombia (Quindio (Filandia), Risaralda (Pereira)), Brazil. Refs.: Hull, 1949c:210, fig. 93 (abdomen). Material examined. COLOMBIA: Quindio: Filandia, Hacienda Bengala, 4.685291°, -75.617533°, 2000m, 1male, 1female, 10.ix.2008, Leg. A.L. Montoya, Net (CEUA); Risaralda: Pereira, Wildlife Sanctuary Otún Quimbaya, 4.728880°, -75.577850°, 1890m, 1male, 1female, 13-14-iv.11, Leg. N. Uribe, Vsr. Chiken (CEUA). quadrilineatus (Enderlein), 1938:233 (Hybobathus). Type locality: Colombia, "Cordillera, tierra templada". HT F (ZMHU). Distr.: Colombia. Refs.: Thompson et al. 1976:54 (cat.). Genus Mimocalla Hull Mimocalla Hull, 1943a: 46. Type species, Baccha capitata Loew, 1863 (orig, des.). (Proposed as a subg. of Baccha). Refs.: Zumbado & Thompson, 2000 (rev. and desc.) Therantha Hull, 1943a:47. Type species, Baccha atypica Curran, 1930 (orig, des.) = Syrphus conjunctus Wiedemann, 1830. (Proposed as a subg. of Baccha). capitatus species group gigantea (Schiner), 1868:340 (Baccha). Type locality: "South America". HT M (NMW). Distr.: Colombia (Antioquia (Girardota, La Ceja, Medellín), Boyacá (Arcabuco, Tunja), Cundinamarca (Bogotá), Meta, Risaralda (Pereira)), Costa Rica, Venezuela. Refs.: Thompson et al. 1976:19 (cat.); Zumbado & Thompson, 2000:776 (key), 781, fig 8–9 (M genitalia complex), 785 (desc.). Material examined. COLOMBIA: Antioquia: Girardota, Vereda El Palmar, 6.343502° -75.435102°, 2200m, 1male, Leg. N. Uribe, Vsr. Fish (CEUA); La Ceja, Alto la Union, Forest, 6.016132°, -75.370182°, 2500m, 1male, 10.ix.2008, Leg. A.L. Montoya, Net (CEUA); Medellín, Vereda Piedras Blancas, 6.291588°, -75.493345°, 2300m, 3male, Leg. A. Velez, Vsr. Fish (CEUA); Boyacá: Arcabuco, Pomeca, 5.750000°, -73.433333°, 2650m, 1.iv.1997, Leg. F. Escobar (ICN); Tunja, Santuario Iguaque Quebrada La Laguna, 6.044559°, -73.154046°, 2340m, 3.iii.1990, Leg. H. Hurtado (ICN); Cundinamarca: Bogotá, Universidad Nacional de Colombia, 4.638194°, -74.084046°, 2567m, 2600m, iv.2011, Leg. E. Vergara, R. Simbaqueba (UNAB); Risaralda: Pereira, Wildlife Sanctuary Otún Quimbaya, 4.728880°, -75.577850°, 1890m, 1female, 13-14.iv.11, Leg. N. Uribe, Vsr. Chiken (CEUA). tristani Zumbado, 2000:788. Type locality: Costa Rica. Heredia, Santo Domingo, INBio. HT M (INBio). Distr.: Colombia (Antioquia (Caldas, Guarne), Meta (Villavicencio)), Costa Rica, Mexico. Refs.: Zumbado & Thompson, 2000:774, fig. 1 (habitus ofmales), 778, fig. 6–7 (M genitalia complex). Material examined. COLOMBIA: Antioquia: Caldas, 6.090000°, -75.638000°, 1560m, 28.xi.2007 (MEFLG); Guarne, Vereda Piedras Blancas, 6.291588°, -75.493345°, 2660m, 1female, Leg. Entomology Group (MEPB); Meta: Villavicencio, Forest Bavaria, 4.150000°, -73.633333°, 467m, 8.x.2005 (ICN). willistoni Thompson (nom. nov. for dolosa Williston). Type locality: Peru, Chanchamayo. HT M (BMNH). Distr.: Colombia (Antioquia (Caldas), Risaralda (Pereira)), Costa Rica, Mexico. Refs.: Williston, 1891:37, pi. 1:7 (habitus), 7a–b (head) (name preocc. by Walker, 1857 (Baccha)); Hull, 1949c:234, fig. 202 (abdomen); Thompson et al. 1976:30 (cat.); Zumbado & Thompson, 2000:784, fig. 18–19 (M genitalia complex), 790 (desc.). Material examined. COLOMBIA: Antioquia: Caldas, 6.090000°, -75.638000°, 1560m, 28.xi.2007 (MEFLG). Risaralda: Pereira, Wildlife Sanctuary Otún Quimbaya, 4.728880°, -75.577850°, 1890m, 1female, 1314-iv.11, Leg. N. Uribe, Vsr. Chiken (CEUA). Genus Ocyptamus Macquart Ocyptamus Macquart, 1834:554. Type species, fascipennis Macquart (Coquillett, 1910:577) = fuscipennis Say. Refs.: Curran, 1928:34 (key to 8 Puerto Rican spp.); 1941:257–265 (key); Hull, 1949c:100–135 (key); Telford, 1973 (key to 19 Puerto Rican spp.); Thompson, 1981:35 (desc. Notes, key to West Indies spp, species groups); 470 · Zootaxa 4122 (1) © 2016 Magnolia Press MONTOYA Thompson, 1999a:326–327 (key); Rotheray et al. 2000:385–398 (acquatic larvae, desc.); Marinoni et al. 2007:147, 150 (desc, key, illustrations); Reemer, 2010:173–187 (desc, key and syn.). Baccha. authors. Cryptamus Stahl, 1883: 97. Misspelling. Baccha (Ocyptamus). Hull, 1949a. Baccha funebris species group. Hull, 1949a. Ocyptamus cylindricus species group. Thompson, 1981. cylindricus species group anthiphates (Walker), 1849:589 (Syrphus). Type locality: "Jamaica". HT M (BMNH). Distr.: Colombia (Antioquia (Alejandria), Boyacá (Togüí), Caldas, Cundinamarca, Risaralda, Santander (Cimitarra), Tolima), Costa Rica, Cuba, Hispaniola, Jamaica, Puerto Rico. Refs.: Thompson, 1981:46; Thompson et al. 1976:12 (cat.); Thompson, 1981:46 (desc, syn.), fig. 1 (habitus). Material examined. COLOMBIA: Alejandria, 6.401714°, -75.058838°, 1200m, 1female, 8.xi.2007, Leg. A.L. Montoya, Vsr. Fish (CEUA); Boyacá: Togüí, 1830m, 1.ii.1978, Leg. I. de Arévalo (ICN); Santander: Cimitarra, 6.310723°, -73.944432°, 263, 1female, 14.vi.2005, Leg. M. Castaño & C. Rivera (CEUA). dimidiatus (Fabricius), 1781:434 (Syrphus). Type locality: "Americae meridionalis insulis". HT M (?). Distr.: Colombia (Antioquia, Caldas (Samaná, Norcasia), Cundinamarca (Bogotá, Cáqueza), Meta (Guayabetal, Via Apiay Villavo), Santander (Piedecuesta), Sucre (San Onofre), Tolima, Valle del Cauca), Brazil, Costa Rica, Cuba, Hispaniola, Jamaica, Less Antillas, Mexico, Puerto Rico. Refs.: Hull, 1949c:240, fig. 232 (abdomen); Thompson et al. 1976:16 (cat.); Thompson, 1981:34, fig 34 (dorsal view of posterolateral portion of thorax), 38 (occiputs lateral view), 47 (M genitalia complex); Reemer, 2010:178 (desc.), fig 29–30 (habitus, males, female), 101 (larva). Material examined. COLOMBIA: Caldas: Samaná, Norcasia, 5.538340°, -74.969231°, 980m, 6.x.1992, Leg. Molano A. Camero (ICN); Cundinamarca: Bogotá, km91 carretera Bogotá–Villavo Pipiralito, 4.139059°, 73.780962°, 1000m, 14.vi.1980, Leg. Hernandez C. Bohorquez (ICN); Cáqueza, Vereda Ubatoque, 4.405278°, 73.950000°, 1743m, 14.viii.1989, Leg. L.P. Moreno (ICN); Meta: Guayabetal, 4.215273°, -73.814434°, 1100m, 18.x.1976, Leg. Chavarro Gerardo (ICN); Via Apiay Villavo, 4.105925°, -73.597319°, 380m, Leg B. González. (ICN); Santander: Piedecuesta, 6.987834°, -73.047684°, 1100m, 12.x.1978, Leg. L. Caballero (ICN); Sucre: San Onofre, 9.646080°, -75.581129°, 5m, 11.x.2003, 1female, Leg. Entomology Group (CEUA). funebris Macquart, 1834:554. Type locality: "Brazil". HT (MNHNP) (lost?). Distr.: Colombia (Antioquia), Brazil, Costa Rica. Refs.: Hull, 1949c:276, figs. 160 & 162 (abdomens); 278, fig. 370 (wing); Thompson et al. 1976:18 (cat.). gastrostactus (Wiedemann), 1830:123 (Syrphus). Type locality: "Brazil". HT M (AMNH). Distr.: Colombia (Antioquia (Alejandria), Caldas, Cundinamarca, Meta, Risaralda), Brazil, Costa Rica, Venezuela. Refs.: Hull, 1943f:65, fig. 2 (alula); 1949c:226, figs. 163–164 (abdomens); 258, figs. 299–300 (wings); Thompson et al. 1976:19 (cat.). Material examined. COLOMBIA: Antioquia: Alejandria, 6.401714°, -75.058838°, 1200m, 1male, 8.xi.07, Leg. A.L. Montoya, Vsr. Fish (CEUA). lepidus species group cultrinus (Curran), 1939b:7 (Baccha). Type locality: Panama, Canal Zone, Barro Colorado Island. HT M (AMNH). Distr.: Colombia (Antioquia (Puerto Berrio)), Panama. Refs.: Hull, 1949c:210, fig. 92 (abdomen); 242, figs 236 & 241 (abdomen); Thompson et al. 1976:16 (cat.). Material examined. COLOMBIA: Antioquia: Puerto Berrio, 6.489819°, -74.401992°, 114m, 1male, 24.ix.2004, Leg. Entomology Group (CEUA). hyalipennis (Curran), 1930e:15 (Callostigma). Type locality: Honduras, Tegucigalpa. HT F (AMNH). Distr.: Colombia (Antioquia (San Roque), Nariño (Barbacoas)), Argentina, Brazil, Honduras. Refs.: Hull, 1949c:230, fig. 183 (abdomen), 260, fig. 311 (wing); Thompson et al. 1976:20 (cat.). CATALOGUE OF SYRPHIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 471 Material examined. COLOMBIA: Antioquia: San Roque, Corregimiento de Sna Jose del Nus, 6.503175°, 74.926885°, 1800m, 1male, 1.v.2012, Leg. N. Uribe, Vsr. Liver (CEUA); Nariño: Barbacoas, Corregimiento Altaquer, Reserva Natural Privada Río Ñambí, 1.651397°, -78.163467°, 93m, 1male, 1370m (ICN). luctuosus (Bigot), 1883:334 (Baccha). Type locality: "Mexico". HT F (BMNH). Distr.: Colombia (Antioquia (Yarumal)), Brazil, Costa Rica, Mexico, Puerto Rico. Refs.: Hull, 1949c:242, fig. 239 (abdomen); Fluke, 1956:252; Thompson et al. 1976:21 (cat.), Rotheray et al. 2000:386 (desc, and syn.), fig. 1–7 (illustrations of larval structures); Reemer, 2010:182 (desc.), fig 48–49 (habitus,males, female). luctuosus Montoya et al. 2012 (Ocyptamus) (cat.). Material examined. COLOMBIA: Antioquia: Yarumal, 6.961666°, -75.426785°, 2500m, Leg. L. Barrientos (CEUA). verona (Curran), 1941:272 (Baccha). Type locality: Colombia, Vista Nieve. HT F (AMNH). Distr.: Colombia (Magdalena). Refs.: Hull, 1949c:240, 254; figs. 225 (abdomen), 288 (wing); Thompson et al. 1976:29 (cat.). vierecki (Curran), 1930a:6 (Baccha). Type locality: Panama, Canal Zone, Barro Colorado Island. HT M (AMNH). Distr.: Colombia, Panama. Refs.: Hull, 1949c:236, fig. 213 (abdomen); Thompson et al. 1976:29 (cat.). melanorrhina species group sativa (Curran), 1941:275 (Baccha). Type locality: Brazil, Santa Catarina, Nova Teutonia. HT M (AMNH). Distr.: Colombia, Brazil. Refs.: Hull, 1949e:248 figs. 250, 264 (abdomen), 283 (wing); Thompson et al. 1976:27 (cat.). arsinoe (Hull), 1949c:250 (Baccha). stenogaster species group stenogaster (Williston), 1888:266 (Baccha). Type locality: Brazil, Mato Grosso, Chapada. HT M (AMNH). Distr.: Colombia (Antioquia (La Pintada), Huila (San Agustin)), Argentina, Brazil, Mexico, Puerto Rico. Refs.: Hull, 1949e:232, 266, 272; figs. 189 & 190 (abdomen), 331 & 357 (wing); Thompson et al. 1976:27 (cat.). Material examined. COLOMBIA: Antioquia: La Pintada, 1male, 16.ii.2007, Leg. N. Uribe, Vsr. Fish (CEUA); Huila: San Agustín, 1male, 22.vii.2008, Leg. A.L. Montoya (CEUA). titania species group filissima (Hull), 1943f:79 (Baccha). Type locality: Colombia, Muzo. HT F (Hull Collection Unknown). Distr.: Colombia (Boyacá). Refs.: Hull, 1949e:200, fig. 42 (abdomen); Thompson et al. 1976:18 (cat.). Genus Orphnabaccha Hull Orphnabaccha Hull, 1949a:93. Type species, Baccha coerulea Williston, 1891 (orig. des.). Mercurymyia Fluke, 1950:140 Type species, Epistrophe caldus Walker, 1852 (orig. des.). (Proposed as a subg. of Stenosyrphus). Synonymy Thompson et al. 1976:11. Refs.: Vockeroth, 1969:118 (Description, taxonomic notes). caldus species group ampla (Fluke), 1942:6 (Epistrophe). Type locality: Brazil, Santa Catarina, Nova Teutonia. HT F (AMNH). Distr.: Colombia (Valle del Cauca), Argentina, Brazil, Panama. Refs.: Fluke, 1942:6, fig. 7 (abdomen); Fluke, 1950a:125, fig. 98 (M genitalia); Vockeroth, 1969:164, fig. 76 (M genitalia); Thompson et al. 1976:12 (cat.). calda (Walker), 1852:232 (Syrphus). Type locality: "Brazil". HT M (BMNH). Distr.: Colombia (Valle del Cauca), Argentina, Brazil, Colombia (Valle del Cauca), Paraguay. Refs.: Fluke, 1937 :4, fig. 6 (abdomen); Vockeroth, 1969:163, fig. 75 (M genitalia); Thompson et al. 1976:14 (cat.). 472 · Zootaxa 4122 (1) © 2016 Magnolia Press MONTOYA coerulea (Williston), 1891:38 (Baccha). Type locality: Mexico, Guerrero; Omiltemi, Amula, Xucumanatlan, Sierra de las Aguas Escondidas. HT M/F (BMNH). Distr.: Colombia (Cauca), Costa Rica. Refs.: Hull, 1949c:224, fig. 159 (abdomen), 272, fig. 355 (wing); Vockeroth, 1969:163, fig. 74 (M genitalia); Thompson et al. 1976:14 (cat.). delimbata (Enderlein), 1938:224 (Allograpta). Type locality: Colombia, Bogota. HT M (ZMHU). Distr.: Colombia (Cundinamarca). Refs.: Thompson et al. 1976:34 (cat.). pteronis (Fluke), 1942:5 (Epistrophe). Type locality: Ecuador, Tungurahua, Volcan Tungurahua, Minza Chica. HT M (AMNH). Distr.: Colombia (Antioquia (Belmira)), Ecuador. Refs.: Fluke, 1950a:125, fig. 108 (M genitalia); Vockeroth, 1969:164, fig. 79 (superior lobe); Thompson et al. 1976:26 (cat.). Material examined. COLOMBIA: Antioquia: Belmira, Paramo de Belmira, 6.707065°, -75.714136°, 3057m, 1male, 5.iii.2009, Leg. A.L. Montoya, Net (CEUA). trabis (Fluke), 1942:5 (Epistrophe). Type locality: Ecuador, Tungurahua, Volcan Tungurahua, Runtun. HT M (AMNH). Distr.: Colombia (Valle del Cauca), Argentina, Ecuador. Refs.: Fluke, 1942:5, fig. 6 (abdomen); 1950a:125, fig. 109 (M genitalia); Vockeroth, 1969:165, fig. 81 (superior lobe); Thompson et al. 1976:28 (cat.). Genus Pelecinobaccha Shannon Pelecinobaccha Shannon, 1927:10. Type species, Baccha peruviana Shannon, 1927 (orig. des.). (Proposed as a subg. of Baccha). adspersa species group adspersa (Fabricius), 1805:200 (Baccha). Type locality: "America meridionali”. HT M (UZMC). Distr.: Colombia (Antioquia (Alejandria, Maceo), Chocó, Boyacá, Meta), Costa Rica, Ecuador, Guyana, Mexico, Peru. Refs.: Hull, 1943f:65, fig. 2 (alula); 1949c:138 (desc.) 188, fig. 33 (abdomen); Thompson et al. 1976:12 (cat.); Reemer, 2010:175 (desc, and notes), figs. 22–23 (habitus, males and female). Material examined. COLOMBIA: Antioquia: Alejandria, Vereda San Jose, 6.371942°, -75.109771°, 1879m, 10.xii.2007, Leg. A.L. Montoya (CEUA); Maceo, Vereda Las Brisas, 6.540547°, -74.770327°, 840m, 18.iii.2006, Leg. C. Vélez (CEUA). oviphora (Hull), 1943f:73 (Baccha). Type locality: Colombia, Muzo. HT F (CNC). Distr.: Colombia (Boyacá). Refs.: Hull, 1949c:232, fig. 191 (abdomen); Thompson et al. 1976:24 (cat.). ovipositoria (Hull), 1943f:76 (Baccha). Type locality: Colombia. HT F (CNC). Distr.: Colombia (Meta). Refs.: Hull, 1949c:230, fig. 178 (abdomen); Thompson et al. 1976:24 (cat.). squamagula Miranda, 2014:70. Type locality: Brazil, Santa Catarina, Nova Teutônia. HT F (CNC). Distr.: Colombia (Cundinamarca (Anapoima)), Brazil, Bolivia. Refs.: Miranda et al. 2014: 70 (fig. 14). brevipennis species group cyclops (Hull), 1947a:406 (Baccha). Type locality: Colombia, Muzo. HT M (Hull Collection Unknown). Distr.: Colombia (Boyacá). Refs.: Hull, 1949c:234, fig. 196 (abdomen); Thompson et al. 1976:16 (cat.); Miranda et al. 2014: 39. gracilitas Miranda, 2014:43. Type locality: Amazonas, Manaus, Boa Vista. HT M (CNC). Distr.: Brazil, Colombia (Boyacá (Muzo)). Refs.: Miranda et al. 2014: 43 (fig. 5). CATALOGUE OF SYRPHIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 473 peruviana species group alucard Miranda, 2014:21. Type locality: Costa Rica, Guanacaste, Eco Museo, Río Aguas Claras, 420m. HT M (INBio). Distr.: Colombia (Antioquia (Girardota)), Costa Rica. Refs.: Miranda et al. 2014: 23 (figs. 15e–i). Material examined. COLOMBIA: Antioquia: Girardota, Vereda Manga Arriba, Secundary forest, 6.343502° -75.435102°, 2200m, 22-26.viii.2009, 1female, Leg. G. Valencia & N. Uribe, Vsr. Viscera (CEUA). andrettae Miranda, 2014:23. Type locality: Ecuador, Napo, Jatun Sacha Biol. Res., 6 km E Misahuali, 450m. HT F (QZAZ). Distr.: Colombia (Amazonas (Letitia)). Refs.: Miranda et al. 2014: 23 (figs. 18a–g). avispas Miranda, 2014:25. Type locality: Peru, Madre de Dios, Avispas, 400m. HT F (CNC). Distr.: Bolivia, Brazil, Colombia (Antioquia (Alejandria)), Ecuador, Peru. Refs.: Miranda et al. 2014:119 (fig. 19). Material examined. COLOMBIA: Antioquia: Alejandria, Vereda San Jose, 6.371942°, -75.109771°, 1879m, Forest, 1male, 1female, 10.viii.2007, Leg. A.L. Montoya, Vsr. Fish (CEUA). clarapex (Wiedemann), 1830:684 (Syrphus). Type locality: Brazil. HT F (NMW). Distr.: Colombia ((Magdalena (Cerro Patron, Rio Frio), Boyacá (Muzo)), Bolivia, Brazil, Colombia, Costa Rica, Ecuador, Guatemala, Peru, Venezuela. Refs.: Hull, 1949c:214, fig. 109 (abdomen); 271, fig. 346 (wing); Thompson et al. 1976:14 (cat.); Miranda et al. 2014: 31 (fig. 22). colombiana Curran, 1941:282 (Baccha). colombianus Thompson et al. 1976:14 (Ocyptamus) (cat.). dracula (Hull), 1943c:52 (Baccha). Type locality: Honduras. HT M (CNC). Distr.: Colombia (Antioquia (La Pintada)), Costa Rica, Guatemala, Honduras, Mexico, Peru. Refs.: Hull, 1949a:152 (redesc.), 232 (fig. 205, abdomen); Miranda et al. 2014: 39, 127 (fig. 27). Material examined. COLOMBIA: Antioquia: La Pintada, 5.736521°, -75.632087°, 650m, 1male, 16.ii.2007, Leg. N. Uribe, Vsr. Fish (CEUA). hiantha (Hull), 1943a:74 (Baccha). Type locality: Brazil, Santa Catarina, Nova Teutônia. HT F (AMNH). Distr.: Brazil, Colombia (Boyacá (Muzo)), Brazil, Costa Rica, Peru. Refs.: Miranda et al. 2014: 44 (fig. 29). ida (Curran), 1941:279 (Baccha). Type locality: Brasil, Santa Catarina, Nova Teutônia. HT M (AMNH). Distr.: Argentina, Brazil, Colombia (Quindio (Filandia)), Ecuador, Paraguay. Refs.: Hull, 1949a:156 (redesc.), 159 (fig. 107, M abdomen), 194 (fig. 12, F abdomen; fig. 18 M abdomen), 262 (fig. 319, M wing), 274 (fig. 358, F wing); Thompson et al. 1976: 20 (cat. cit.); Miranda et al. 2014:131 (fig. 31). Material examined. COLOMBIA: Quindio: Filandia, Hacienda Bengala, 4.685291°, -75.617533°, 2000m, 3male, 10.ix.2008, Leg. A.L. Montoya (CEUA). peruviana Shannon, 1927:10 (Baccha). Type locality: Peru, Chanchamayo. HT F (USNM). Distr.: Colombia (Amazonas (Leticia)), Peru. Refs.: Hull, 1949a: 207 (fig. 69, Fabdomen; fig. 70, M abdomen); Thompson et al. 1976: 25 (cat. cit.); Miranda et al. 2014: 63, 108 (fig. 34i-j). Material examined. COLOMBIA: Amazonas: Leticia, Puerto Nariño Amazonas, - -3.774094°, 70.381153°, 98m, 10.vii.2012, 1female, Leg. C. Bota, Net (CEUA) pilipes (Schiner), 1868:342 (Baccha). Type locality: "South America". HT M/F (NMW). Distr.: Colombia (Antioquia (La Pintada), Valle del Cauca, Vaupés (Nbra Koyo)), Brazil. Refs.: Hull, 1949c:210, fig. 94 (abdomen), 272, fig. 350 (wing); Thompson et al. 1976:25 (cat.). nigrocilia var. hirtipes Hull, 1943e:215 (Baccha). (missing FCT). nigrocilia var. inclusa Hull, 1943e:215 (Baccha). (missing FCT). Material examined. COLOMBIA: Antioquia: La Pintada, 5.736521°, -75.632087°, 650m, 1male, 16.ii.2007, Leg. N. Uribe, Vsr. Fish (CEUA); Vaupés: Nbra Koyo, 0.983621°’, -70.864236°, 180, 1male, Leg. J. Gonzalez (CEUA). 474 · Zootaxa 4122 (1) © 2016 Magnolia Press MONTOYA pucallpa Miranda, 2014:68. Type locality: Peru, Ucayali, Pucallpa. HT M (CNC). Distr.: Colombia (Antioquia (Puerto Berrio)), Peru. Refs.: Miranda et al. 2014: 68, 136 (figs. 36h–l). Material examined. COLOMBIA: Antioquia: Puerto Berrio, Alto de las Aguilas, Hacienda Manaos, 6.500865°, -74.546317°, 404m, 1male, 5-6.vi.2013, Leg. M. Wolff & INPA, Net (CEUA). telescopica Curran, 1930:74. Type locality: Colombia. HT F (AMNH). Distr.: Colombia, Panama. Refs.: Hull, 1943a:44 (fig. 1, alula); Hull, 1949a:205 (fig. 60, abdomen), 207 (fig. 71, abdomen), 276 (fig. 369, wing) 1843j:94, fig. 71 (abdomen) 1849a:169 (redesc.), 224 (fig. 154, abdomen); Thompson et al. 1976:28 (cat.). stipa Hull, 1949a:110 (Baccha). transatlantica (Schiner), 1868:343 (Baccha). Type locality: "Brazil & Colombia". HT M/F (NMW). Distr.: Colombia, Brazil, Peru. Refs.: Hull, 1949c:216, 284; figs. 119 & 120 (abdomen), 392 & 393 (wing); Thompson et al. 1976:28 (cat.). susio species group pandora (Hull), 1942i:102 (Baccha). Type locality: Panama, Code prov, El Valle. HT M (MCZ). Distr.: Colombia (Antioquia (La Pintada, San Roque), Caldas (Supia)), Costa Rica, Panama. Refs.: Hull, 1949c:198, fig. 38 (abdomen); Thompson et al. 1976:24 (cat.). Material examined. COLOMBIA: Antioquia: La Pintada, 5.736521°, -75.632087°, 650m, 2male, 8female, 16.ii.07 Leg. N. Uribe, Vsr. Fish (CEUA); San Roque, 6.503175°, -74.926885°, 1800m, 3male, 3female, 8.ii.2009, Leg. A.L. Montoya (CEUA); Caldas: Supia, Vereda Piedras, Quebrada Piedras, 5.557611°, -75.577439°, 600m, 1male, 2.iii.2009, Leg. A.L. Montoya, Net (CEUA). Genus Pseudodoros Becker Pseudodoros Becker, 1903:92. Type species, nigricollis Becker (mon.). Baccha, subg. Dioprosopa Hull, 1949c:99 (also 1949b:296). Type species, Syrphus clavatus Fabricius (mon.). Refs.: Thompson, 1981:33 (desc. and notes); Thompson, 1999a:327 (key); Kassebeer, 2000:80 (desc, key); Marinoni et al. 2007:148, 150 (desc, key, illustrations). Subgenus Dioprosopa Hull Dioprosopa Hull 1949: 99. Type species, Syrphus clavatus (Fabricius), 1794 (mon.) clavatus (Fabricius), 1794:298 (Syrphus). Type locality: "Americae meridionalis Insulis". HT M/F (UZMC). Distr.: Colombia (Antioquia (Alejandria, Copacabana, La Pintada, Santafé de Antioquia), Arauca, Boyacá (Villa de Leyva, Cáqueza), Caldas (Pensilvania), Cundinamarca, Meta, Putumayo, Quindío (Filandia), Santander (Bucaramanga, Piedecuesta, San Gil y Guane), Sucre (San Onofre), Tolima, Valle del Cauca (Farallones)), Argentina, Bahamas, Brazil, Costa Rica, Cuba, Ecuador, Hispaniola, Jamaica, Less Antillas, Puerto Rico. Refs.: Weidemann, 1830:94 (desc. based on type); Townsend, 1895b:37 (desc. notes); Williston, 1896:347 (syn.; distr. notes); Wolcott, 1923:7, fig. 7 (larva), 8 (pupa), 9 (adult) (life hist.) 1826:218 (prey recs); Hull, 1943c:48, fig. 7 (abdomen) 1849e:192, fig. 7 (abdomen) 1849d:288, fig. 9a (head); Thompson et al. 1976:10 (cat.); Thompson, 1981:33, (desc. notes); Kassebeer, 2000:80, fig 1f (female abdomen), 2g–h (M genitalia complex, dist. notes); Reemer, 2010:171 (desc, and notes), fig. 11, 12 (habitus). Material examined. COLOMBIA: Antioquia: Alejandria, Vereda San Jose, 6.371942°, -75.109771°, 1879m, Forest, 1male, 1female, 10.viii.2007, Leg. A.L. Montoya, Vsr. Fish (CEUA); Copacabana, Barrio Machado, 6.345579°, -75.503714°, 1450m, 12.ix.2007, Leg. M. Giraldo, Manual (CEUA); La Pintada, 5.736521°, -75.632087°, 650m, 1female, 26.vi.2007, Leg. N. Uribe, Vsr. Fish (CEUA); Santafé de Antioquia, 6.512784°, 75.825832°, 460m, 25.v.2007, Leg. A.L. Montoya (CEUA); Boyacá: Villa de Leyva, 5.638191°, -73.535870°, 2200m, 24.x.1997, Leg. F. Fernández (ICN); Caldas: Pensilvania, Vereda La Rioja, 5.384070°, -75.160990°, 1956m, 1female, 7.xi.20013, Leg. C. Bota & C. Flórez, Net (CEUA); Cundinamarca: Cáqueza, Vereda Ubatoque, CATALOGUE OF SYRPHIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 475 4.405278°, -73.950000°, 1743m, 21.viii.1989, Leg. L.P. Moreno (ICN); Guayabetal, 4.215287°, -73.814548°, 1120m, Leg. G.B. Gonzalez (ICN); Quindio: Filandia, 4.685291°, -75.617533°, 1700m, 1male, 13.xi.2009, Leg. A.L. Montoya (CEUA); Santander: Bucaramanga, Cañon del Chicamocha, 6.770764°, -73.008988°, 1400m, 22.xii.2010, Leg. A.L. Montoya (CEUA); Piedecuesta, 6.987834°, -73.047684°, 1110m, 12.x.1978, Leg. J. Caballero (ICN); San Gil y Guane, 6.551825°, -73.134334°, 1300m, 5.iv.2009, Leg. Fernández–Alonso & J.L. Fernández–Castillo (ICN); Sucre: San Onofre, 9.646080°, -75.581129°, 5m, 1male, 11.x.2004, GEUA Leg., Vsr. Banana (CEUA); Valle del Cauca: Farallones de Cali, 3.367176°, -76.687470°, 3400m, 2.v.1993, Leg. C. Andrade (ICN). Genus Relictanum Miranda Relictanum Miranda, 2014:85. Type species, Baccha crassa Walker, 1852. Ref.: Miranda, 2014. Baccha tristis species group, in part. Hull, 1949a. crassum (Walker), 1852:222 (Baccha). Type locality: Brazil. HT M (BMNH). Distr.: Brazil, Colombia (Amazonas (Leticia)), Ecuador, Guyana, Panama, Peru, Suriname, Trinidad. Refs.: Miranda et al. 2014: 90 (fig. 44). nero Curran, 1939:8. Type locality: Guyana, Kaieteur. HT M (BMNH). Distr.: Colombia (Boyacá (Muzo)), Bolivia, Brazil, French Guiana, Mexico, Peru. Refs.: Hull, 1949c:159 (desc.), 224, fig. 151 (abdomen) 182 (redesc.), 224 (fig. 153, abdomen), 258 (fig. 298, wing); Thompson et al. 1976:20 (cat.); Miranda et al. 2014: 94 (fig. 47). infanta Hull, 1943f:74 (Baccha). nigrocilia (Hull), 1943g:215 (Baccha). Type locality: Brazil, Sao Paulo, Ilha Seca. HT F (AMNH). Distr.: Colombia, Brazil. Refs.: Hull, 1949c:162 (desc.), 244, fig. 156, 248 (abdomen), Hull, 1949c:220, fig. 139 (abdomen); Thompson et al. 1976:23 (cat.). hirtipes (Hull), 1943g:215 (Baccha). (missing FCT). inclusa (Hull), 1943g:2l5 (Baccha). (missing FCT). schwarzi (Curran), 1939b:9 (Baccha). Type locality: Colombia, Western Cordillera, Cali HT M (AMNH). Distr.: Colombia (Valle del Cauca (Cali)), Costa Rica, Peru. Refs.: Hull, 1949c:214, 278; figs. 111 (abdomen), 375 (wing); Thompson et al. 1976:27 (cat.); Miranda et al. 2014. Genus Rhinoprosopa Hull Oligorhina Hull, 1937a: 30 (name preocc. by Fairmaire & Germain 1863). Type species, aenea Hull (orig. des.). Rhinoprosopa Hull, 1942:23 (nom. nov. for Oligorhina Hull). Refs.: Thompson, 2012 (generic status); Mengual, 2014 (generic rev.) flavophylla (Hull), 1943d:319 (Rhinoprosopa). Type locality: Peru, Perene Valley. El Campamiento. HT F (CUI). Distr.: Colombia (Antioquia (Amalfi, Belmira, Caldas, Ciudad Bolivar, Envigado, Jardin), Caldas (Anserma), Quindío (Salento), Risaralda (Pueblo Rico)), Costa Rica, Peru. Refs.: Hull, 1949b:269, fig. 6b (head); 1949c:222, fig. 146 (abdomen); Thompson et al. 1976:35 (cat.); Mengual, 2014:10 Figs. 30 (M, head, lateral view), 31–32 (M, habitus, dorsal and frontal view), 33 (M, head, frontolateral view). Material examined. COLOMBIA: Antioquia: Amalfi, Vereda Guayabito, Finca Costar Rica, 1840m, 1male, 19-feb-2005, Leg. Velez & Diaz (CEUA); Belmira, Paramo de Santa Inés (El Morro), 6.778957°, -75.699390°, 3200m, 1male, Abril 24 2011, Leg. J. Cardona & J.D. Cardona, Net (CEUA); Belmira, Morro Cabildo Verede, 6.6579, -75.672, 3200m, 30.ix.2011, Leg. J.D. Castaño & C. Bota, Malaise (CEUA); Caldas, Alto La Romera, 6.090000°, -75.638000°, 1560m, 2male, Leg. E. Alarcón (CEUA); Ciudad Bolivar, Vereda La Mina, 5.772073°, 76.032002°, 2400m, 1male, Leg. E. Garces, Net (CEUA); Envigado, El Salado, Jardin en Borde dde Quebrada, 8 mar 2009, Leg. C. Bota, Net (CEUA); Jardin, Vereda Florestar-El Cañon, Finca Los Robles 5.583337°, - 476 · Zootaxa 4122 (1) © 2016 Magnolia Press MONTOYA 75.833332°, 2000m, Net, Rastrojo, 28-abr-2001, Leg. G. Etnomology (CEAU); Caldas: Anserma, 5.249026°, 75.795074°, 1700m, 1male, 1female, 15.xi.2008, Leg. A.L. Montoya (CEUA); Quindio: Salento, Vereda Canaan, 4.613714°, -75.581637°, 1800m, 1male, Leg. A.L. Montoya, Net (CEUA); Risaralda: Pueblo Rico, Vereda Montebello, Rio Claro, Forest, 5.248847°, -76.060004°, 1350m, 1male, 15-14.iv.2014, Leg. C. Bota, M. Loaiza, C. Flórez, Vsr. Copro (CEUA). lucifer (Hull), 1943g:216 (Rhinoprosopa). Type locality: Ecuador, Pinas, 1600 ft. HT M (AMNH). Distr.: Colombia (Sucre (San Onofre), Tolima, Valle del Cauca (San Antonio)), Costa Rica, Ecuador. Refs.: Hull, 1949c:222, fig. 141 (abdomen); Thompson et al. 1976:36 (cat.). Material examined. COLOMBIA: Sucre: San Onofre, 9.646080°, -75.581129°, 5m, 4males, 1male, Leg. A. Velez & L. Rios, Vsr. Fruit (CEUA). nasuta (Bigot), 1884:103 (Sphaerophoria). Type locality: “Mexico”. HT M (UMO). Distr.: Colombia (Valle del Cauca (Palmira)), Costa Rica, Ecuador, El Salvador, Guatemala, Mexico, Venezuela. Refs.: Thompson et al. 1976:36 (cat.); Mengual, 2014: 15, Figs. 45–46 (F, habitus, lateral and dorsal view), 47–48 (F, head, frontal and frontolateral view), 49–50 (M, habitus, lateral and dorsal view), 51–52 (M, head, frontal and frontolateral view). sycorax (Hull), 1947b:239 (Rhinoprosopa). Type locality: Costa Rica, La Suiza. HT F (USNM). Distr.: Colombia, (Antioquia (Bello), Valle del Cauca), Costa Rica, Venezuela. Refs.: Thompson et al. 1976:37 (cat.); Mengual, 2014: 18, Figs. 53–54 (F, habitus, dorsal and lateral view), 55–56 (M, lateral and dorsal view), 57–58 (F, head, frontal and frontolateral view), 59–60 (M, head, frontal and frontolateral view). Material examined. COLOMBIA: Antioquia: Bello, San Felix, Paramo de Las Baldias, 6.332653°, 75.646599°, 3050m, 1female, 17.ii.2013, Leg. A.L. Montoya, Net (CEUA). Genus Salpingogaster Schiner Amathia Walker, 1852: 223 (name preocc. by Lamouroux 1812; Roux, 1828; Duponchel, 1829; Rathke, 1837). Type species, costalis Walker (mon.). Salpingogaster Schiner, 1868:344. Type species, pygophora Schiner (orig. des.). Refs.: Sack, 1920 (rev. key); Curran, 1941 (key); Knutson, 1971 (biol.); Thompson, 1981:63 (key); Thompson, 1999a:326 (key); Marinoni et al. 2007:148, 150 (desc, key, illustrations). Flexineura Bezzi, 1908:294. Nomen nudum. abdominalis Sack, 1920:242. Type locality: "Bolivia". HT M (SMT). Distr.: Colombia (Quindío (Salento)), Bolivia. Refs.: Sack, 1920:242, fig. D (tip of abdomen); Thompson et al. 1976:31 (cat.). Material examined. COLOMBIA: Quindio: Salento, 4.613714°, -75.581637°, 1800m, 3males, 28.iv.2009, Leg. A.L. Montoya (CEUA). cornuta Hull, 1944a:168. Type locality: Venezuela". HT F (NMW). Distr.: Colombia (Antioquia (Andes), Caldas, Caqueta (Chiribiquete), Cauca, Cundinamarca (La Mesa)), Venezuela. Refs.: Thompson et al. 1976:31 (cat.). Material examined. COLOMBIA: Antioquia: Andes, Vereda Quebrada arriba, Forest pescado, 5.600370°, 75.965892°, 2100-2500m, 6male, 26-29-ii-1997, Leg. O. Ortega (Coll. O. Ortega); Caquetá: Chiribiquete, Río Cuñeré, 0.746043°, -72.738590°, 100m, 8.viii.1994, Leg. A. Amarillo (ICN); Cundinamarca: La Mesa, Laguna Pedro Palo, 4.629429°, -74.469603°, 2031m, 16.iii.1997, Leg. V. González (UNAB). limbipennis Williston, 1891:29. Type locality: Mexico, Guerrero, Amula, 6000 ft. HT F (BMNH). Distr.: Colombia (Antioquia (Andes)), Mexico. Refs.: Sack, 1920:256 (desc.); Thompson et al. 1976:32 (cat.). Material examined. COLOMBIA: Antioquia: Andes, Paramo de Santa Rita, 5.550249°, -75.988733°, 3600m, 8.xii.2010, 1male, Leg. A.L. Montoya (CEUA). nigra Schiner, 1868:344. Type locality: "South America". HT M/F (NMW). Distr.: Colombia (Antioquia (San Roque), Boyacá, Caldas, Cauca (Popayán), Cundinamarca (San Pedro de Jagua, Tibirita, Villeta), Huila, Meta CATALOGUE OF SYRPHIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 477 (Reserva La Macarena Caño Curia, San Martín, Villavicencio), Nariño (Barbacoas), Quindío, Valle del Cauca (Restrepo)), Brazil, Costa Rica, Guatemala, Hispaniola, Jamaica, Mexico, Panama, Peru, Puerto Rico, Venezuela. Refs.: Schiner, 1868:344, pi. 4: fig. 4 (habitus), 4a (wing); Sack, 1920:254 (desc.), figs, a (wing), t, u, & v (abdomen); Thompson, 1981:67, fig 81 (abdomen); Thompson et al. 1976:32 (cat.); Reemer, 2010:190 (desc.), fig 17 (habitus males). Material examined. COLOMBIA: Antioquia: San Roque, Vereda La Mora, Alto El Cuatro, 6.503175°, 74.926885°, 1800m, x.2008, Leg. H. Paredes (UNAB); Cauca: Popayán, Vereda Rejoya Finca Helechaux, 2.454604°, -76.608737°, 1025m, 21.iv.1994, Leg. C. Sarmiento (ICN); Cundinamarca: San Pedro de Jagua, 4.647699°, -73.325182°, 775m, 8.ix.1988, Leg. D.N. Padilla (ICN); Tibirita, 5.054444°, -73.508056°, 1880m, 22.xi.2003, Leg. O. Munar, Net (UNAB); Villeta, 5.008319°, -74.473554°, 992m, 21.xi.2003, Leg. A. Molano, Net (UNAB); Meta: Reserva La Macarena Caño Curia, 2.072107°, -73.955920°, 380m, 24.ix.1989, Leg. W. Cubillos (ICN); San Martín, Finca El Caduceo, 3.694490°, -73.693630°, 340m, 15.v.2006, Leg. G. Aves (ICN); Villavicencio, Vereda La Vanguardia, Pozo Azul, 4.150000°, -73.633333°, 467m, 16.iv.2005 (ICN); Nariño: Barbacoas, Corregimiento Altaquer. Reserva Natural Privada Río Ñambí, 1.651397°, -78.163467°, 93m, 13.vii.1995, Leg. C. Sarmiento (ICN); Valle del Cauca: Restrepo, Camino Hacienda El Piral, 3.811560°, 76.515178°, 1000m, 6.ii.1984, Leg. P. Saray (ICN). panamana Hull, 1944c:33. Type locality: Panama, Bella Vista. HT M (MCZ). Distr.: Colombia (Antioquia (Urrao)), Panama. Refs.: Thompson et al. 1976:32 (cat.). Material examined. COLOMBIA: Antioquia: Urrao, 6.416558°, -76.107506°, 3500m, 6.i.1991, Leg. A. Madrigal (MEFLG). pygophora Schiner, 1868:344. Type locality: "South America". HT M (NMW). Distr.: Colombia (Antioquia (Gomez Plata), Cundinamarca (Sasaima)), Brazil, Costa Rica, Cuba, Mexico, Panama. Refs.: Sack, 1920:239 (desc.), fig. b (tip of males abdomen); Hull, 1949b:264, fig. 3e (head); Thompson et al. 1976:32 (cat.); Thompson, 1981:64, fig 67a, b (wing andmales genitalia complex). Material examined. COLOMBIA: Antioquia: Gomez Plata, 6.683154°, -75.216575°, 1800m, 3males, 1female, 29.x.1998, Leg. Entomology Group (CEUA); Cundinamarca: Sasaima, 4.966667°, -74.433333°, 1221m, 5.ii.2005, Leg.V. Castiblanco, Manual (UNAB). vera Hull, 1944a:169. Type locality: Bolivia, Rio Songo. HT M (NMW). Distr.: Colombia (Antioquia (Caldas)), Bolivia. Refs.: Thompson et al. 1976:32 (cat.). Material examined. COLOMBIA: Antioquia: Caldas, 6.090000°, -75.638000°, 1560m, 1male, 10.i.1973, Leg. R. Vélez (MEFLG). Genus Scaeva Fabricius Scaeva Fabricius, 1805:248. Type species, Musca pyrastri Linnaeus, 1758 (Subs. des.; Curtis, 1834:509). Catabomba Osten Sacken, 1877:329 Type species, Musca pyrastri Linnaeus, 1758 (mono.). Lasiopthicus Rondani, 1845:459. Type species, Musca pyrastri Linnaeus, 1758 (orig. des.). Refs.: Shannon & Aubertin, 1933 (key); Thompson, 1999a:325, 327 (key). occidentalis Shannon, 1927:29. Type locality: Chile, Santiago. HT M (USNM). Distr.: Colombia (Antioquia (Medellín)), Argentina, Chile. Refs.: Vockeroth, 1969:150, fig. 36 (M genitalia); Thompson et al. 1976:9 (cat.). Material examined. COLOMBIA: Antioquia: Medellín, 6.267404°, -75.585789°, 1570m, 1female, iv.1945, Leg. F.L. Gallego (MEFLG). Genus Styxia Hull Styxia Hull, 1943a: 46. Type species, Styxia eblis Hull, 1943 (orig. des.). (Proposed as a subg. of Baccha). 478 · Zootaxa 4122 (1) © 2016 Magnolia Press MONTOYA eblis (Hull), 1943f (abdomen) (Styxia). Type locality: Ecuador, Bolivar, Hacienda Talahua. HT M (AMNH). Distr.: Colombia (Antioquia (Betulia)), Ecuador. Refs.: Hull, 1949c:208, fig. 85 (abdomen). Material examined. COLOMBIA: Antioquia: Betulia, Vereda el Indio, La Mina, Alto de San Jose, Paramo, 5.771994°, -76.086152°, 3284-3300m, 1male, 14-18.xii.2013, Leg. C. Henao & J. Cogollo (CEUA). Genus Syrphus Fabricius Syrphus Fabricius, 1775:762. Type species, Musca ribesii Linnaeus (see Wirth et al. 1965:558) to preserve established usage. Refs.: Fluke, 1942:2–3 (key, rev. Neotropical spp.); Vockeroth, 1969:55–57 (descript); Thompson, 1981:23 (desc. note); Thompson, 1999a:326 (key); Thompson et al. 2000:7 (key and descriptions); Marinoni et al. 2007:148, 150 (desc, key, illustrations). laceyorum Thompson, 2000:6. Type locality: Costa Rica, San José, Parque Nacional Braulio Carillo, Estacion Barva. HT M (INBio). Distr.: Colombia (Caldas (Riosucio), Cundinamarca), Costa Rica. Refs.: Thompson et al. 2000: 3, fig. 1b (habitus), 5, fig. 3a–c (M genitalia), 6 (desc.); Gutiérrez et al. 2005. Material examined. COLOMBIA: Caldas: Riosucio, Camino hacia el cerro Ingurumá, 5.421334°, 75.719539°, 2200m, 1female, 7.i.2013, Leg. A.L. Montoya (CEUA). octomaculatus Walker, 1837:344. Type locality: "Chile". HT F (BMNH). Distr.: Colombia (Antioquia (Andes)), Argentina, Costa Rica, Mexico. Refs.: Macquart, 1842:150 (90), pi. 13, fig. 8 (head), Macquart, 1850:457 (153), pi. 14, fig. 12 (head); Fluke, 1942:3, Fig. 1 (abdomen); Thompson et al. 1976:8 (cat.); Thompson et al. 2000:8 (key). Material examined. COLOMBIA: Antioquia: Andes, Paramo de Santa Rita, 5.550249°, -75.988733°, 20002600m, 1male, 8.xii.2010, Leg. A.L. Montoya (CEUA). phaeostigma Wiedemann, 1883:130. Type locality: "Brazil". HT M (NMW). Distr.: Argentina, Brazil, Colombia (Santander (California)). Refs.: Fluke, 1942:2 (redesc.), fig. 3 (abdomen); Fluke, 1950a:117, fig. 15 (male genitalia). Material examined. COLOMBIA: Santander: California, Vereda California, 7.377812°, -72.938002°, 2800–3000m, 1male, 12–19.ix.2012, Leg. A. Vélez & J.D. Sánchez (CEUA). shorae Fluke, 1950b:440 (nom. nov. for willistoni Fluke). Type locality: Mexico, Guerrero, Omiltemi, Costa Rica, Volcan de Irazu, Rio Sucio. HT F (BMNH). Distr.: Colombia (Antioquia (Bello, Belmira, Caldas, Guarne, La Ceja, San Jose de la Montaña, San Pedro), Boyacá (Buenavista, Villa de Leyva) Cundinamarca (Bogotá, Chocóntá, Facatativá, Machetá, Silvania), Quindío (Filandia), Risaralda (Pereira) Santander (California, Charalá, Zapatoca), Valle del Cauca (Sevilla)), Argentina, Costa Rica, Chile. Refs.: Fluke, 1942:22, fig. 2 (abdomen); Fluke, 1950a:117, fig. 14 (M genitalia); Thompson et al. 1976:8 (cat.); Thompson et al. 2000:8 (key). Material examined. COLOMBIA: Antioquia: Bello, San Felix, Paramo de Las Baldias, 6.332653°, 75.646599°, 3050m, 1male, 17.ii.2013, Leg. A.L. Montoya, Net (CEUA); Belmira, Paramo de Belmira, 6.707065°, -75.714136°, 3057m, 3males, 1female, 5.iii.2009, Leg. A.L. Montoya, Net (CEUA); Caldas, Alto de San Miguel, 6.090000°, -75.638000°, 1560m, 1male, 27.vii.2009, Leg. N. Uribe, Net (CEUA); Guarne, Vereda Piedras Blancas, 6.291588°, -75.493345°, 2660m, 2females, Leg. Entomology Group (CEUA); La Ceja, Alto la Union, Forest, 6.016132°, -75.370182°, 2500m, 10.ix.2008, 1female, Leg. A.L. Montoya, Net (CEUA); San Jose de la Montaña, Vereda el Congo sector la Laguna, Potrero, 6.804244°, -75.689613°, 3100–3200m, 1female, 10–14.ix.2011, Leg. L. Rios, Net (CEUA); San Pedro, Vereda Cerezales, 6.483237°, -75.630955°, 2800m, 1female, Leg. V. Pérez Net in a decomposed pig (CEUA); Boyacá: Buenavista, Vereda Patiño, Las Lomas, 5.650832°, -74.286905°, 1001m, 04.iv.2004, Leg. W. Ávila, Net (UNAB); Villa de Leyva, Bella Vista Km130 Carretera Villavo, 4.139059°, 73.780962°, 1500m, Leg. M. Alvarez (ICN); Cundinamarca: Bogotá, Botanic Garden, 4.668332°, -74.100229°, 2551m, 20.ix.2009, Leg. M. Zamora (ICN); 23.xii.2006, Leg. Fernández–A., J.L. Fernández–C (ICN); Bogotá, Jardínes Barrio Palermo, 4.677811°, -74.101080°, 2650m, 1.iii.2008, Leg. C.A. Fernández & A. Fernández (ICN); Chocontá, Reserva La Macarena Caño Curia, 5.152037°, -73.623517°, 2800m, 14.vii.2009, Leg. C.A. Fernández & A. Fernández (ICN); Facatativá, vía Madrid–Facatativá, The Elitte Flowers, El Corzo, Finca Margaritas, 4.814185°, -74.354995° 2600m, 13.vii.2005, Leg. A. Mendoza (UNAB); Machetá, El Caucho, 5.080217°, - CATALOGUE OF SYRPHIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 479 73.607475°, 2100m, 12.x.2003, Leg. F. Jiménez, Net (UNAB); Silvania, El Vergel, 4.342730°, -74.404917°, 1470m, 13.xi.2003 (UNAB); Quindio: Filandia, Hacienda Bengala, 4.685291°, -75.617533°, 2000m, 1male, 2females, 10.ix.2008, Leg. A.L. Montoya (CEUA); Risaralda: Pererira, PNR Ucumari, La Pastora, 4.708555°, 75.487224°, 2500m, 1.x.2001, Leg. G. Bernal, A. Carlos (ICN); Pererira, La Pastora, 4.708555°, -75.487224°, 2500m, 1female, 1.x.2001, Leg. C. Bota & J. Cardona, Net (CEUA); Santander: Charalá, 6.304190°, 73.173909°, 1859m, 27.xi.1979, Leg. G.I.I. de Arévalo (ICN); California, Vereda California, 7.377812°, 72.938002°, 2800–3000m, 1male, 12–19.ix.2012, Leg. A. Vélez & J.D. Sánchez (CEUA); Zapatoca, Betulia, Cuenca del Rio El Reno, 6.904303, -73.293379, 1700-2000m, 9-11.viii.20013, Leg. C. Bota & C. Florez, Net (CEUA); Valle del Cauca: Sevilla, Bella Vista K.130 Carretera Villavo, 4.268890°, -75.936100°, 2000m, 2.iv.1974, Leg. J. Aguirre (ICN). Tribe Pipizini Genus Trichopsomyia Williston Trichopsomyia Williston, 1888:259. Type species, polita Williston (Hull, 1949b: 330). Refs.: Fluke, 1937 (key); Thompson, 1972a:99 (desc.); Thompson, 1981:116 (desc. and syn.; Thompson, 1999a:332 (key). Halictomyia Shannon, 1927:13. Type species, boliviensis Shannon (orig. des.). Parapenium Collin, 1952:85. Type species, Pipiza carbonaria Meigen (orig. des.). urania Hull, 1949a: 73. Type locality: Peru, Pucallpa. HT M (Hull). Distr. Colombia Colombia (Boyacá (Tunja)), Peru. Refs.: Thompson et al. 1976:68 (cat.); Thompson 2006. Material examined. COLOMBIA: Boyacá: Tunja, Santuario Iguaque Carrizal, 6.044559°, -73.154046°, 2340m, 8.i.1992, Leg. C. Sarmiento (ICN). Tribe Toxomerini Genus Toxomerus Macquart Toxomerus Macquart, 1855:112 (1855:92). Type species, notatus Macquart (orig. des.) = geminatus (Say). Refs.: Hull, 1941:45–49 (key); Thompson, 1981:66 (key); Thompson, 1999a:326 (key); Metz & Thompson, 2001:225–256 (rev. of larger spp.); Thompson & Thompson, 2006:317–331 (rev. of Chilean spp); Marinoni et al. 2007:149, 150 (desc, key, illustrations); Borges & Couri, 2009:1–72 (rev. of Brazilian spp.); Reemer, 2010:190–194 (rev. of Surinam spp.); Mengual, 2011:1–26 (rev. of Black–tie dress spp.). Mesogramma Loew, 1866a:157 (1872b:31) (name preocc. by Stephens, 1850). Type species, parvula Loew (Williston, 1887:98). Mesograpta Loew, 1872:114 (1872b:290) (nom. nov. for Mesogramma Loew). Type species, parvula Loew (aut.). Mitrosphen Enderlein, 1938:227. Type species, Syrphus tibicen Wiedemann (orig. des.). Antiops Enderlein, 1938:228. Type species, Mesograpta ? saphiridiceps Bigot (orig. des.). anthrax (Schiner), 1868:350 (Mesogramma). Type locality: "South America". HT M/F (NMW). Distr.: Colombia (Magdalena (Aracataca)), Costa Rica, Ecuador, Guyana, Peru. Refs.: Thompson et al. 1976:48 (cat.); Mengual, 2011:6 (key), 8 (diagn. and syn.), (thorax, lateral view, M genitalia). vitrecens (Hull), 1930:142 (Mesogramma). apegiensis (Harbach), 1974:31 (Mesogramma). Type locality: Brazil, Belen de Para. HT M (CNC). Distr.: Colombia, Brazil. Refs.: Borges & Couri, 2009:11 (desc.), (F genitalia); Reemer & Rotheray, 2009:939–949 (larval morphology and food preferences); Reemer, 2010:190 (desc, syn., habitus M and F). interruptus (Enderlein), 1938:227 (name preocc. by Philippi, 1865 (Mitrosphen)). aquilinus Sack, 1941:100. Type locality: Peru, Pichis, Puerto Yessup. HT M (SMT). Distr.: Colombia ((Magdalena, Quindio (Quimbaya), Risaralda, Sucre (San Onofre)), Bolivia, Brazil, Ecuador, Mexico, Panama, 480 · Zootaxa 4122 (1) © 2016 Magnolia Press MONTOYA Peru, Venezuela. Refs.: Sack, 1941:100 (1951:96), (abdomen); Hull, 1941:35, (abdomen); Thompson et al. 1976:48 (cat.); Metz & Thompson, 2001:231 (redesc.), (abdomen, wing, M genitalia). Material examined. COLOMBIA: Quindio: Quimbaya, Vereda Kerman, Km 7, 4.614947°, -75.807598°, 1339m, 1female, 1.vii.2003, Leg. H. Vargas (CEUA); Sucre: San Onofre, 9.646080°, -75.581129°, 5m, 4males, 1male, Leg. A. Velez & L. Rios, Vsr. Fruit (CEUA). brevifacies (Hull), 1941:20 (Mesogramma). Type locality: Ecuador, Baños, Runtum Trail. HT M (AMNH). Distr.: Colombia (Cauca), Ecuador. Refs.: Thompson et al. 1976:49 (cat.). centaureus (Hull), 1951a:18 (Mesogramma). Type locality: Colombia, Cali, West Cordillera. HT M (AMNH). Distr.: Colombia (Valle del Cauca). Refs.: Thompson et al. 1976:49 (cat.). dispar (Fabricius), 1794:309 (Syrphus). Type locality: “America meridionalis” restricted to Virgin Islands, St. Croix (Thompson, 1981:86). HT M/F (UZMC). Distr.: Colombia (Antioquia (Anorí, Caldas, Ciudad Bolivar, Caucasia, Cocorná, Guarne, La Pintada, Nariño, Puerto Berrio, San Roque, Santafé de Antioquia), Caldas (Norcacia, Pensilvania), Cundinamarca (Bogotá), Meta, Quindío (Filandia, Salento), Risaralda (Pereira), Tolima, Valle del Cauca), Brazil, Costa Rica, Cuba, El Salvador, Guatemala, Hispaniola, Honduras, Jamaica, Less Antillas, Mexico, Puerto Rico, Suriname, Venezuela. Refs.: Thompson et al. 1976:39 (cat.); Thompson, 1981:25, (thorax, lateral view, abdomen, a palem, b darkm, c very dar F, wing), (M genitalia complex), 119 (F genitalia); Thompson & Thompson, 2006:321 (desc.), (abdomen, M genitalia); Borges & Couri, 2009:29 (desc.), (thorax, lateral view, abdomen, dorsal view); Reemer, 2010:191 (desc, syn.), (habitus M and F); Mengual, 2011:7 (key), 10 (diagn. and syn.), (abdomen morphotypes), (wing, M genitalia). Material examined. COLOMBIA: Antioquia: Anori, Vereda El Roble, Reserva Natural Arrierito Antioqueño, 6.984806°, -75.111667°, 1600–1700m, 1male, 25.v.2012, Leg. Entomology Group, Vsr. Fish (CEUA); Caldas, Alto La Romera, 6.090000°, -75.638000°, 1560m, 1male, Leg. A.L. Montoya (CEUA); Ciudad Bolivar, Corregimiento Farallones, Vereda La Arboleda, Quebrada La Arboleda, 1male, 2-7.vii.2012, Leg. E.C. Bota & C. Flórez, Net (CEUA); Caucasia, Hacienda La Candelaria, 7.983333°, -75.200000°, 80m, 1.ix.2006; Leg. Entomology Group, Net, (CEUA); Cocorná, Vereda El Viao, Secundary forest, 6.050187°, -75.166752°, 1950m, 1male, Leg. L. Rios, Net (CEUA); Guarne, Vereda Piedras Blancas, 6.291588°, -75.493345°, 2660m, 1male, Leg. Entomology Group (CEUA); La Pintada, Camping Los Farallones, 5.743018°, -75.604756°, 660m, 1female, Leg. J.D. Sanchez, Vsr. Copro (CEUA); Nariño, Vereda Puente Linda, 5.631484°, -75.245454°, 2400m, 2males, Leg. L. Rios, Net (CEUA); Puerto Berrio, Vereda Cristalina, RUNA, 6.489819°, -74.401992°, 114m, 1male, 25-ix.2006, Leg. Entomology Group, Net (CEUA); San Roque, Corregimiento San Jose del Nus, 6.503175°, -74.926885°, 1800m, 1male, 5.i.2007, Leg. N. Uribe, Net (CEUA); Santafé de Antioquia, 6.512784°, -75.825832°, 460m, 1male, Leg. A.L. Montoya, Net (CEUA); Caldas: Norcacia, Sector El Tigre, Forest, 5.560100°, -74.893349°, 700m, 1female, 5.xii.2009, Leg. A.L. Montoya, Net (CEUA); Pensilvania, Vereda La Rioja, 5.384070°, -75.160990°, 1956m, 1male, 7.xi.2013, Leg. C. Bota & C. Flórez, Net (CEUA); Cundinamarca: Bogotá, Suba, San Jose, 4.757031°, -74.088591°, 2555m, 6.ii.1974, Leg. J. Peréa (ICN); Quindio: Filandia, Hacienda Bengala, 4.685291°, -75.617533°, 2000m, 9male, 10.ix.2008, Leg. A.L. Montoya (CEUA); Salento, Vereda Canaan, 4.613714°, 75.581637°, 1800m, 4males, Leg. A.L. Montoya, Net (CEUA); Risaralda: Pereira, Vereda Mundo Nuevo, Finca la Acuarela, 1550m, 1female, 18.x.2006, Leg. D. Grisales, Cerdo 1 Manual (CEUA); Pereira, PNR Ucumari, La Pastora, 4.708555°, -75.487224°, 2500m, 2males, 7.iv.2008, Leg. A. Bustamante, Net (CEUA). duplicatus (Wiedemann), 1830:142. Type locality: Uruguay, Montevideo. HT M/F (NMW). Distr.: Colombia ((Antioquia (San Pedro de los Milagros)), Argentina, Brazil, Mexico, Puerto Rico, Uruguay. Refs.: Thompson et al. 1976:50 (cat.); Thompson & Thompson, 2006:322 (desc.), (abdomen, M genitalia); Borges & Couri, 2009:31 (desc. and syn.), (wing, F genitalia). Material examined. COLOMBIA: Antioquia: San Pedro de los Milagros, 1male, 20.vi.2001, Leg. V. Pérez (CEUA). flaviplurus (Hall), 1927:239 (Mesogramma). Type locality: Guatemala, Puerto Barrios. HT F (USNM). Distr.: Colombia (Valle del Cauca), Brazil, Costa Rica, Guatemala, Trinidad. Refs.: Curran, 1930:1 (key); 1934:398 (key); CATALOGUE OF SYRPHIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 481 Hall, 1927:239, (abdomen); Thompson et al. 1976:50 (cat.); Borges & Couri, 2009:34 (desc.), (thorax, lateral view, wing, abdomen, M genitalia, F genitalia); Mengual, 2011:5, 6 (key), 13 (diagn. and syn.), (abdomen, thorax, lateral view). floralis (Fabricius), 1798:563 (Syrphus). Type locality: "Cajenne". HT M/F (UZMC). Distr.: Colombia (Antioquia (Caucasia, La Pintada), Boyacá (Pauna), Cundinamarca (Cachipay, Medina), Meta (Acacías)), Bahamas, Brazil, Chile, Costa Rica, Cuba, El Salvador, Guatemala, Hispaniola, Jamaica, Less Antillas, Mexico, Nicaragua, Puerto Rico, Venezuela. Refs.: Hull, 1941:35, (abdomen); Thompson et al. 1976:50 (cat.); Thompson, 1981:68, (abdomen F, wing, M genitalia complex, F genitalia); Thompson & Thompson, 2006:323 (desc.), (abdomen, M genitalia); Borges & Couri, 2009:35 (desc, syn.); Reemer, 2010:191 (desc., syn.), (habitus F and M). Material examined. COLOMBIA: Antioquia: Caucasia, Hacienda La Candelaria, 7.983333°, -75.200000°, 80m, 1male, 1.ix.2006; Leg. Entomology Group, Net, (CEUA); La Pintada, Camping Los Farallones, 5.743018°, 75.604756°, 660m, 1female, Leg. J.D. Sanchez, Vsr. Viscera (CEUA); Boyacá: Pauna, Manote bajo La Rochela, 1280m, 26.iv.1979, Leg. I. de Arevalo (ICN); Cundinamarca: Cachipay, 10.v.1990, Leg. F. Torres (ICN); Medina, Riberas Río Gasaguan pequeño, 4.508319°, -73.361189°, 440m, 16.x.1989, Leg. M. García (ICN); Meta: Acacías, Vereda San Jose, 3.987806°, -73.759747°, 660m, 4.xii.1985, Leg. C. López (ICN). Det F.C. Thompson, 2009. hieroglyphicus (Schiner), 1868:348 (Mesogramma). Type locality: "South America". HT M (NMW). Distr.: Colombia (Cauca), Ecuador. Refs.: Hull, 1941:31, (abdomen); Thompson et al. 1976:51 (cat.); Mengual, 2011:5 (key), 16 (diagn. and syn.), (abdomen, head, lateral view). idalius (Hull), 1951a:11 (Mesograma). Type locality: Ecuador, Oriente, Puyo. HT M (AMNH). Distr.: Colombia (Antioquia (Itagui), Valle del Cauca), Brazil, Ecuador. Refs.: Thompson et al. 1976:51 (cat.); Borges & Couri, 2009:36 (desc.), (Head, lateral view, M genitalia, F genitalia). Material examined. COLOMBIA: Antioquia: Itagui, 1750m, 1male, 10.x.2004, Leg. E. Alarcon (CEUA). insignis (Schiner), 1868:347 (Mesogramma). Type locality: "South America". HT F (NMW). Distr.: Colombia Antioquia (Alejandria, Caldas, Guarne, Jardin, La Ceja, Medellín, Nariño, San Vicente), Boyacá, Caldas (Manizales), Quindío (Filandia), Risaralda (Pereira), Valle del Cauca), Bolivia, Ecuador, Venezuela. Refs.: Schiner, 1868:347, (habitus, head); Thompson et al. 1976:51 (cat.); Metz & Thompson, 2001:234 (redesc.), (abdomen, thorax), (wing, M genitalia). elongata Hull, 1941:46 (Mesogramma). Material examined. COLOMBIA: Antioquia: Alejandria, Vereda El Carbón, 6.403715°, -75.064663°, 1445m, 1male, Leg. Entomology Group, Net (CEUA); Caldas, Alto La Romera, 6.090000°, -75.638000°, 1560m, 1female, Leg. E. Alarcón (CEUA); Guarne, Vereda Piedras Blancas, 6.291588°, -75.493345°, 2660m, 1female, Leg. Entomology Group (MEPB); Jardin, Vereda La Herrera, Cuchilla Jardin Tamesis, 5.568645°, -75.737518°, 2750m, 1female, 19.v.2006, Leg. Velez, Perez & Vargas, Net (CEUA); La Ceja, Alto la Union, Forest, 6.016132°, 75.370182°, 2500m, 10.ix.2008, 1male, Leg. A.L. Montoya, Net (CEUA); Medellín, San Antonio de Prado, Vereda Yarumalito, 6.186412°, -75.658630°, 2300m, 3males, 7–8.xi.2008, Leg. C. Bota & N. Uribe, Manual (CEUA); Nariño, Vereda Puente Linda, 5.631484°, -75.245454°, 2400m, 1female, Leg. L. Rios, Net (CEUA); San Vicente, Vereda Chaparral, 3.750000°, -75.583333°, 1354m, 1female, Leg. M. Wolff & A. Bustamante, Net (CEUA); Caldas: Manizales, Vereda las Palomas, Reserva Rio Blanco, 5.111962°, -75.462904°, 2550m, 2males, Leg. A. Velez, Net (CEUA); Quindio: Filandia, Hacienda Bengala, 4.685291°, -75.617533°, 2000m, 3males, 10.ix.2008, Leg. A.L. Montoya (CEUA); Risaralda: Pereira, Wildlife Sanctuary Otún Quimbaya, 4.728880°, -75.577850°, 1890m, 1female, vi.2010, Leg. C. Bota, Net (CEUA). lacrymosus (Bigot), 1884:107 (Mesograpta?). Type locality: "Brazil; Mexico". HT M/F (UMO). Distr.: Colombia (Antioquia (Alejandria, La Pintada, Maceo, Medellín, Nariño, San Roque, San Vicente), Boyacá, Caldas (Norcacia)), Brazil, Costa Rica, Mexico. Refs.: Hull, 1941:31, (abdomen); Thompson et al. 1976:51 (cat.); Borges & Couri, 2009:38, (wing, M genitalia, F genitalia); Reemer, 2010:192 (desc, syn.), (habitus, M and F). Material examined. COLOMBIA: Antioquia: Alejandria, Vereda San José, 6.403715°, -75.064663°, 1445m, 1male, 1female, Leg. Entomology Group, Net (CEUA); La Pintada, 5.736521°, -75.632087°, 650m, 482 · Zootaxa 4122 (1) © 2016 Magnolia Press MONTOYA 1female, 4.v.2007, Leg. M.C. Vélez, Vsr. Viscera (CEUA); Maceo, Reserva Cañón del Río Alicante, 6.540547°, 74.770327°, 840m, 1female, Leg. S. Perez, Net (CEUA); Medellín, Barrio Belén, 6.216630°, -75.599930°, 1500m, 1female, Leg. A.L. Montoya, Net (CEUA); Nariño, Vereda Puente Linda, 5.631484°, -75.245454°, 2400m, 2females, Leg. L. Rios, Net (CEUA); San Roque, Corregimiento San Jose del Nus, 6.503175°, -74.926885°, 1800m, 2females, 5.i.2007, Leg. A.L. Montoya, Net (CEUA); San Vicente, Vereda El Chaparal 1800m, 2females, Leg. A.L. Montoya, Net (CEUA); Caldas: Norcacia, 5.560100°, -74.893349°, 700m, 3females, 5.xii.2007, Leg. A.L. Montoya (CEUA). laenas (Walker), 1852:241 (Syrphus). Type locality: "Brazil". HT M (BMNH). Distr.: Brazil, Colombia. Refs.: Hull, 1943e:41, fig. 79 (abdomen) ; Mengual 2011: 18 (desc.), 12, fig. 12 (abdomen, M, dorsal), 25 (thorax, lateral). nitidiventris Curran, 1930b:9 (Mesogramma). mutuus (Say), 1829:164 (Syrphus). Type locality: "Mexico". HT M/F (?). Distr.: Colombia, Costa Rica, Guatemala, Mexico). Refs.: Hull, 1943e:43, (abdomen); Thompson et al. 1976:52 (cat.). nigripunctus (Enderlein), 1938:231 (Mesogramma). Type locality: Colombia, Cordillera. HT M (ZMHU). Distr.: Colombia. Refs.: Thompson et al. 1976:52 (cat.). nitidus (Schiner), 1868:349 (Mesograma). Type locality: "South America". ST M (NMW). Distr.: Colombia, Panama. Refs.: Thompson et al. 1976:53 (cat.); Mengual 2011:17, fig. 24 (Thorax), 18 (desc.). norma (Curran), 1930b:10 (Mesogramma). Type locality, Panama, Canal Zone, Corozal. HT M (AMNH). Distr.: Colombia (Risaralda), Brazil, Costa Rica, Ecuador, Mexico, Panama, Paraguay, Venezuela. Refs.: Hull, 1943e:29, (abdomen); Thompson et al. 1976:53 (cat.); Borges & Couri, 2009:43 (desc.), (abdomen, F genitalia). mulio (Hull), 1941c:311 (Mesogramma). Thompson et al. 1976:52 (dist. record). Material examined. COLOMBIA: Quindio: Quimbaya, Vereda Kerman, Km 7, 4.614947°, -75.807598°, 1339m, 1female, 1.vii.2003, Leg. H. Vargas (CEUA); Risaralda: Pereira, Wildlife Sanctuary Otún Quimbaya, 4.728880°, -75.577850°, 1890m, 1female, vi.2010, Leg. C. Bota, Vsr. Fish (CEUA). ochraceus (Hull), 1942f:19 (Mesogramma). Type locality: Colombia, Bogota. HT F (MCZ). Distr.: Colombia (Cundinamarca). Refs.: Thompson et al. 1976:53 (cat.). ophiolinea (Hull), 1943f:68 (Baccha). Type locality: Colombia, Villavicencio. HT F (CNC). Distr.: Colombia (Meta). Refs.: Hull, 1943f:68, (abdomen); Hull, 1949c:202, (abdomen); Thompson et al. 1976:53 (cat.). pallipes (Bigot), 1884:106 (Mesograpta?). Type locality: "Mexico". HT F (UMO). Distr.: Colombia (Antioquia (Bello, Jardín, Medellín, San Pedro de Los Milagros, Sonson), Boyacá, Caldas (Manizales)), Costa Rica, Mexico. Refs.: Hull, 1943e:47, (abdomen); Thompson et al. 1976:55 (cat.). Material examined. COLOMBIA: Antioquia: Bello, San Felix, Paramo de Las Baldias, 6.332653°, 75.646599°, 3050m, 2males, 17.ii.2013, Leg. A.L. Montoya, Net (CEUA); Jardin, Vereda La Herrera, Cuchilla Jardin Tamesis, 5.568645°, -75.737518°, 2750m, 1male, 3–4.xi.2013, Leg. Velez, Perez, Vargas, Net (CEUA); Medellín, 6.267404°, -75.585789°, 1570m, 1.v.2010, Leg. N. Uribe (CEUA); San Pedro de Los Milagros, 2male, 1male, 20.vi.2001, Leg. A. Ocampo (CEUA); Sonson, Cerro Las Palomas, 5.537061°, -75.273872°, 3000–3200m, 2males, Leg. C. Bota & L. Ríos, Net (CEUA); Caldas: Manizales, Vereda las Palomas, Reserva Rio Blanco, 5.111962°, -75.462904°, 2550m, 1male, Leg. A. Velez, Manual (CEUA). paraduplicatus Borges, 2009:44. Type locality: Brazil, São Paulo: Salesópolis. HT M (CNC). Distr.: Colombia (Antioquia (Medellín)), Brazil. Refs.: Borges & Couri, 2009:44 (org. des.), (Thorax, lateral vew, wing, hind leg, abdomen, M genitalia complex, F genitalia). Material examined. COLOMBIA: Antioquia: Medellín, 6.267404°, -75.585789°, 1570m, v.2010, Leg. Entomology Group (CEUA). pictus (Macquart), 1842:159 (Syrphus). Type locality: "Guyane". HT F (MNHNP). Distr.: Colombia (Antioquia CATALOGUE OF SYRPHIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 483 (Belmira)), Greater Antilles, Lesser Antilles, Mexico south to Argentina. Refs.: Macquart), 1842:159 (99) pi. 16 fig. 11 (habitus) Hull, 19431:30, fig. 13 (abdomen). Material examined. COLOMBIA: Antioquia: Belmira, Paramo de Belmira, 6.707065°, -75.714136°, 3057m, 1male, 1female, 5.iii.2009, Leg. A.L. Montoya, Net (CEUA). politus (Say), 1823:88 (Scaeva). Type locality: "United States ". HT M (USNM). Distr.: Colombia (Antioquia (Alejandria, La Ceja), Cesar, Cundinamarca (Apulo, Mosquera, La Mesa, Villeta), Magdalena (Santa Marta), Casanare (Paz de Ariporo), Meta (Acacías, Villavicencio), Risaralda (Pereira, Santuario), Tolima (Falán), Valle del Cauca (Buga, Cerrito, Palmira), Argentina, Bahamas, Brazil, Costa Rica, Cuba, Ecuador, El Salvador, Guatemala, Hispaniola, Honduras, Jamaica, Less Antillas, Mexico, Peru, Puerto Rico, Venezuela. Refs.: Macquart, 1842:97 (132) abdomen); Hull, 1941:29, (abdomen); Thompson et al. 1976:53 (cat.); Thompson, 1981:81, (M genitalia complex, key); Metz & Thompson, 2001:228, 240 (key & redesc.), (abdomen, thorax, wing, larva), Reemer & Rotheray, 2009:939–949 (larval morphology and food preferences); Borges & Couri, 2009:47 (descript, syn.), (F genitalia); Reemer, 2010:193 (desc, syn.), (habitus, F). Material examined. COLOMBIA: Antioquia: Alejandria, Vereda San José, 6.403715°, -75.064663°, 1445m, 1male, Leg. Entomology Group, Net (CEUA); La Ceja, Alto la Union, Forest, 6.016132°, -75.370182°, 2500m, 1male, 10.ix.2008, Leg. A.L. Montoya, Net (CEUA); Cundinamarca: Apulo, 421m, ix.1999, Leg. L. Millán (UNAB); Mosquera, 4.680641°, -74.216488°, 2550m, 30.ix.2002, Leg. R. Chizaba (UNAB); La Mesa, 4.629469°, -74.466633°, 1298m, 12.ix.2003, Leg. R. Parra (UNAB); Villeta, Buenos Aires, Utica, 5.008319°, 74.473554°, 992m, 29.x.1977, Leg. I. de Arévalo (ICN); Magdalena: PNN Tayrona Neguanje, 11.276050°, 74.195519°, 121m, 1.viii.1996, Leg. F. Escobar (ICN); Casanare: Paz de Ariporo, 670m, 28.iii.2002, Leg.J. Peña & M. Guzmán, Net (UNAB); Meta: Acacías, 3.987806°, -73.759747°, 502m, 23.v.2002, Leg. J. Cárdenas (UNAB); Villavicencio, 4.150000°, -73.633333°, 467m, 27.v.2004, Leg.J. Escalante & M. Romero, Manual (UNAB); Risaralda: Pereira, Ins. Pol. Combia, 2137m, 27.v.1999, Leg. Cely, Forero & Rojas (UNAB); Santuario, Vereda La Esmeralda, Natural National Park Tatama, 5.105551°, -76.008496°, 2000m, 18-30.x.2013, Leg. C. Bota & C. Flórez, Net (CEUA); Tolima: Falán, Vereda San Antonio, Finca Villa Liz, 983m 18.viii.2003, Leg. H. Ávila (UNAB); Valle del Cauca: Buga, Semillas del Valle, 3.900000°, -76.300000°, 969m, 14.x.2003, Leg. E. Ortíz, Net (UNAB); Cerrito, Semillas del Valle, 987m, 14.x.2003, Leg. P. Rodríguez, Net (UNAB); Palmira, 3.587073°, 76.248936°, 1059m, 20.v.2002, Leg. J. Martínez, Manual (UNAB). polygraphicus (Hull), 1940a:249 (Mesogramma). Type locality: Brazil, Sao Paulo, Bateia. HT F (CNC). Distr.: Colombia (Antioquia (San Vicente), Cundinamarca, Meta, Quindio (Filandia), Santander (Charalá)), Brazil, Peru. Refs.: Hull, 1941:31, (abdomen); Thompson et al. 1976:54 (cat.); Borges & Couri, 2009:48 (desc. and syn.), (abdomen, M genitalia, F genitalia). Material examined. COLOMBIA: Antioquia: San Vicente, Vereda El Chaparal 1800m, 1male, Leg. A.L. Montoya, Net (CEUA); Meta: km91 carretera Bogotá–Villavo Pipiralito, 4.139059°, -73.780962°, 1000m, 14.vi.1980, Leg. C. Hernandez (ICN); Quindio: Filandia, Hacienda Bengala, 4.685291°, -75.617533°, 2000m, 1male, 1female, 10.ix.2008, Leg. A.L. Montoya, Net (CEUA); Santander: Charalá, Cañaverales, 6.304190°, 73.173909°, 1800m, 28.xi.1978, Leg. I. de Arévalo (ICN). procrastinatus Metz, 2001:242. Type locality: Brazil, Santa Catarina, Nova Teutonia. HT M (USNM). Distr.: Colombia (Quindio (Quimbaya)) Argentina, Brazil, Panama. Refs.: Metz & Thompson, 2001:242, (abdomen, thorax, wing, M genitalia); Borges & Pamplona, 2001:49, (F genitalia). Material examined. COLOMBIA: Quindio: Quimbaya, Vereda Kerman, Km 7, 4.614947°, -75.807598°, 1339m, 1male, Leg. H. Vargas, Vsr. Fish (CEUA). pulchellus (Macquart), 1846:266 (Syrphus). Type locality: Guadeloupe, San Domingo. HT F (UMO). Distr.: Colombia (Antioquia (La Pintada, San Roque)), Bahamas, Brazil, Costa Rica, Cuba, El Salvador, Hispaniola, Honduras, Jamaica, Mexico, Panama, Puerto Rico, Venezuela. Refs.: Macquart, 1846:266, (abdomen); Hull, 1941:34, (abdomen); Thompson, 1981:67, (abdomen F), wing, M genitalia complex); Borges & Couri, 2009:51, figs. 143–145 (F genitalia); Reemer, 2010:193 (desc, syn.), (habitus M and F). Material examined. COLOMBIA: Antioquia: La Pintada, 5.736521°, -75.632087°, 650m, 1female, 484 · Zootaxa 4122 (1) © 2016 Magnolia Press MONTOYA 4.v.2007, Leg. M.C. Vélez, Vsr. Fish (CEUA); San Roque, Corregimiento San Jose del Nus, 6.503175°, 74.926885°, 1800m, 2males, 5.i.2007, Leg. A.L. Montoya, Net (CEUA). saphiridiceps (Bigot), 1884:105 (Mesograpta?). Type locality: "Mexico". HT M (UMO). Distr.: Colombia (Meta (Restrepo)), Bolivia, Costa Rica, Ecuador, Peru, Venezuela. Refs.: Thompson et al. 1976:54 (cat.); Metz & Thompson, 2001:244 (redesc.), (abdomen, wing, M genitalia). flavus Hull, 1941:432 (Mesogramma) sylvaticus (Hull), 1943f:34 (Mesogramma). Type locality: Ecuador, Baños. HT M (AMNH). Distr.: Colombia, Brazil, Ecuador. Refs.: Hull, 19431:41, (abdomen); Thompson et al. 1976:55 (cat.); Borges & Couri, 2009:55, (head lateral view, abdomen, F genitalia). vierecki (Curran), 1930b:13 (Mesogramma). Type locality: Colombia, Sero Quenado. HT F (AMNH). Distr.: Colombia (Caqueta). Refs.: Hull, 1941:37, (abdomen); Thompson et al. 1976:56 (cat.). virgulatus (Macquart), 1850:456 (Syrphus). Type locality: "Patrie inconnue, probablement le Bresil". HT M (UMO). Distr.: Colombia (Antioquia (Caldas, La Pintada, San Roque), Huila (Obando), Quindio (Filandia)), Brazil, Costa Rica, Guatemala, Mexico, Nicaragua, Panama, Venezuela. Refs.: Macquart, 1850:456 (152), (abdomen); Thompson et al. 1976:56 (cat.); Borges & Couri, 2009:58 (desc, and syn.), (abdomen, M genitalia, F genitalia); Reemer, 2010:194 (desc, syn,), (habitus M and F). Material examined. COLOMBIA: Antioquia: Caldas, Alto La Romera, 6.090000°, -75.638000°, 1560m, 3males, Leg. A.L. Montoya (CEUA); Caldas, Alto de San Miguel, 6.090000°, -75.638000°, 1560m, 27.vii.2009, Leg. J. Cardona, N. Franz, G. Mazo, Net (CEUA); La Pintada, 5.736521°, -75.632087°, 650m, 2females, 26.v.2007, Leg. N. Uribe, Vsr. Fish (CEUA); San Roque, Corregimiento San Jose del Nus, 6.503175°, 74.926885°, 1800m, 1female, 5.i.2007, Leg. N. Uribe, VSR. Fish (CEUA); Huila: Obando, 1.926307°, 76.300314°, 1850m, 2males, 1female, Leg. A.L. Montoya, Net (CEUA); Quindio: Filandia, Hacienda Bengala, 4.685291°, -75.617533°, 2000m, 1male, 1female, 10.ix.2008, Leg. A.L. Montoya (CEUA). watsoni (Curran), 1930b:5 (Mesogramma). Type locality: Haiti, Aux Cayes. HT M (UMO). Distr.: Colombia (Antioquia, Cundinamarca), Bolivia, Brazil, Costa Rica, Cuba, El Salvador, Hispaniola, Guatemala, Jamaica, Less Antillas, Mexico, Peru, Puerto Rico, Venezuela. Refs.: Curran, 1930b:5, (ventral view of M genitalia); Hull, 1941:33, (abdomen); Thompson et al. 1976:56 (cat.); Thompson, 1981:25, (thorax, lateral view), (M genitalia complex); Borges & Couri, 2009:68, (F genitalia); Reemer, 2010:194 (desc, syn.), (habitus, M, F). Subfamily Microdontinae Rondani, 1845 Genus Menidon Reemer Menidon Reemer, 2013:39. Type species, Microdon falcatus Williston, 1887:9. falcatus Williston, 1887:9. Type locality: Mexico, Isthmus of Tehuantepec. HT F (USNM). Distr.: Colombia (Antioquia (La Pintada)), Bolivia, Brazil, Costa Rica, El Salvador, Honduras, Mexico. Refs.: Thompson et al. 1976:64 (cat.); Thompson, 2007:283–291 (redesc, LT design. and syn.), figs. 1 (male, lateral view), 2 (female, lateral view), 3–4 (head, frontoblique view), 5–6 (thorax, dorsal view), 7 (abdomen, dorsal view), 8 (scutellum, dorsal view), 10 (M genitalia); Reemer & Ståhls, 2013:12–13, 39 (key, diagnosis and taxonomic notes), 133 (status notes), figs 151 (habitus dorsal), 152 (habitus lateral), 153 (head frontal). Material examined. COLOMBIA: Antioquia: La Pintada, Camping Los Farallones, 5.743018°, 75.604756°, 660m, Leg. J.D. Sanchez, Manual (CEUA). CATALOGUE OF SYRPHIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 485 Genus Microdon Meigen Microdon Meigen, 1803:275. Type species, Musca mutabilis Linnaeus (mon.). Refs.: Curran, 1941:248–250 (key to Neotropical spp.); Thompson, 1969:74–85 (key and notes); Thompson, 1981:112 (key and illustrations of West Indies spp.); Thompson, 1999a:332 (key); Cheng & Thompson, 2010:25, 40 (key, diagnosis and notes); Reemer & Ståhls, 2013:16, 21 (key, diagnosis and taxonomic notes), figs 27–34 (habitus dorsal, wing,males genitalia, head lateral). Aphritis Latreille, 1804:193. Type species, auropupescens Latreille, 1805 (mon.). Colacis Gistel, 1848: x. (nom. nov. for Microdon Meigen). Parmula Heyden, 1825:589. Type species, cocciformis Heyden, 1825:589 (mon.). Scutelligera Spix, 1824:148. Type species, ammerlandia Spix, 1824:124 (mon.). Scutigerella Haas, 1924:148. Misspelling of Spix 1824. Subgenus Chymophila Macquart Chymophila Macquart, 1834:485. Type species, splendens Macquart, 1834:486 (mon.). Refs.: Cheng & Thompson (2008); Reemer & Ståhls, 2013. Chimophila Osten Sacken, 1875:46. Misspelling. Eumicrodon Curran, 1925a:50. Type species, Microdon fulgens Wiedemann, 1830:82 (orig. des.). cyaneiventris (Macquart), 1846:249 (Aphritis). Type locality: "Colombia". HT F (MCZ). Distr.: Colombia. Refs.: Macquart, 1846:249 (121), pi. 10, fig. 14 (wing); Thompson et al. 1976:64 (cat.); Reemer & Ståhls, 2013:136 (status notes). marceli Curran, 1936b:7. Type locality: Brazil, Mato Grosso, Chapada. HT M (AMNH). Distr.: Colombia (Antioquia (Gomez Plata), Bolívar (Zambrano)), Brazil. Refs.: Thompson et al. 1976:66 (cat.); Reemer & Ståhls, 2013:136 (status notes). Material examined. COLOMBIA: Antioquia: Gómez Plata, 6°41'0.03"N, 75°13'0.00"W (MEFLG); Bolívar: Zambrano, Hacienda Monterrey, 9°44'59.88"N, 74°49'59.91"W, 70m, 2.i.1997, Leg. Ulloa G. Fernández (ICN). Genus Mixogaster Macquart Mixogaster Macquart, 1842:74 (14). Type species, conopsoides Macquart (orig. des.). Refs.: Hull, 1954 (rev. Neotropical spp.), Carrera & Lane, 1958 (key); Thompson, 1969:74–85 (key and notes); Thompson, 1999a:331 (key); Cheng & Thompson, 2010:23, 30 (key, diagnosis and notes). Myxogaster Kertesz, 1910:351. Incorrect original spelling of Mixogaster Macquart. Myxogaster Shiraki, 1930:8. Incorrect original spelling of Mixogaster Macquart. Myxogasteroides Shiraki, 1930:9. Type species, nigripennis Sack (orig. des.). Subjective syn. of Paramicrodon (Hull, 1949:317). conopsoides Macquart, 1842: 74 (14). Type locality: Brazil, Rio de Janeiro. HT F (MNHNP). Distr.: Colombia (Antioquia (Barbosa, Cisneros), Caquetá (San Vicente del Caguán)), Brazil. Refs.: Macquart, 1842:74 (14), tab. 3, fig. 1 (habitus), la (head); Thompson et al. 1976:58 (cat.); Reemer & Ståhls, 2013:142 (status notes). Material examined. COLOMBIA: Antioquia: Barbosa, Forest, 6.439000°, -75.333000°, 1330m, 1male, 8.xii.1997, Leg. L. Lopez, Net (CEUA); Cisneros, Montaña al lado del Rio Azul, Primary Forest, 6.538319°, 75.088603°, 1038m, 1female, 4.i.2009, Leg. A. Bustamante, Net (CEUA); Caquetá: San Vicente del Caguán, Vereda Las Vegas, 2.116999°, -74.766711°, 335m, 11.ix.1995, Leg. C. Sarmiento (ICN). pithecofascia Hull, 1944d:512. Type locality: Colombia, Restrepo. HT F (CNC). Distr.: Colombia (Meta). Refs.: Hull, 1954:23 (desc.) figs. 3 & 15 (head), 32 (abdomen); Thompson et al. 1976:58 (cat.); Reemer & Ståhls, 2013:143 (status notes). 486 · Zootaxa 4122 (1) © 2016 Magnolia Press MONTOYA Genus Peradon Reemer Peradon Reemer, 2013:61. Type species, Mulio bidens Fabricius, 1805. bidens species group angustus (Macquart), 1846:250 (122) (Aphritis). Type locality: "Cayenne". HT M (MNHNP). Distr.: Colombia, Guyana. Refs.: Thompson et al. 1976:63 (cat.); Reemer & Ståhls, 2013:145 (status notes). trivittatus species group diaphanus Sack, 1921:146. Type locality: Paraguay, St. Trinidad. HT M (DEI). Distr.: Colombia (Meta (Villavicencio)), Brazil, Paraguay. Refs.: Sack, 1921:146, figs. 21 (habitus), 22 (head); Thompson et al. 1976:64 (cat.); Reemer & Ståhls, 2013:146 (status notes). Material examined. COLOMBIA: Meta: Villavicencio, 4.150000°, -73.633333°, 467m, 1male, Leg. ICN group (ICN). trivittatus Curran, 1925c:344. Type locality: Guyana, Kartabo. HT M (AMNH). Distr.: Colombia (Meta (Villavicencio)), Brazil, Guyana. Refs.: Thompson et al. 1976:67 (cat.); Reemer & Ståhls, 2013:146 (status notes). Material examined. COLOMBIA: Meta: Villavicencio, Forest Bavaria, 4°9'0.00"N, 73°37'60.00"W, 580m, 7.iii.1996, Leg. C. Sarmiento (ICN). Subfamily Eristalinae Newman, 1834 Tribe Brachyopini Genus Lepidomyia Loew Lepidomyia Loew, 1864a:69 (1864b:231). Type species, calopiis Loew (mon.). Refs.: Hull, 1946b (rev, key); Thompson, 1968:272 (key and notes); Thompson, 1972b:116–117 (description); Thompson et al. 1976:88 (cat.); Thompson, 1981:141 (taxonomic notes); Thompson, 1999a:330 (key). Lepidostola Mik, 1886:278 (unjustified nom. nov. for Lepidomyia Loew). Lepromyia Williston, 1887:31 (unjustified nom. nov. for Lepidomyia Loew). colombia Curran, 1929:495 (Lepidostola). Type locality: Colombia, Rio Frio, Magdalena. HT F (AMNH). Distr.: Colombia (Magdalena). Refs.: Hull, 1946b:10–11; Thompson et al. 1976:88 (cat.). Genus Orthonevra Macquart Orthonevra Macquart, 1829:188 (40). Type species, Chrysogaster elegans Meigen (mon.). Refs.: Sedman, 1964 (rev, Keys); Thompson, 1972b: 119–121 (description); Thompson et al. 1976:90–91 (cat.); Thompson, 1981:141 (taxonomic notes); Thompson, 1999a:332. Orthoneura, emend. Cryptineura Bigot, 1859:308. Type species, hieroglyphca Bigot (mon.)= nitida Wiedemann. Chrysogaster authors (part). argentina Brèthes, 1922:141 (Chrysogaster). Type locality: Argentina, Buenos Aires, Sierra de la Ventana. HT M (MACN). Distr.: Colombia (Antioquia (Medellin)), Argentina. Refs.: Thompson et al. 1976:91 (cat.). Material examined. COLOMBIA: Antioquia: Medellín, 6.267404°, -75.585789°, 1570m, 1male, Leg. N. Uribe, Net (CEUA). shannoni (Curran), 1925a:119 (Chrysogaster). Type locality: Costa Rica, San Mateo, Huigito. HT M (USNM). CATALOGUE OF SYRPHIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 487 Distr.: Colombia (Antioquia (Girardota, Envigado), Cundinamarca (Sopo)), Costa Rica. Refs.: Thompson et al. 1976:91 (cat.). Material examined. COLOMBIA: Antioquia: Girardota, Vereda La Honda, 6.343502° -75.435102°, 2200m, 1male, Leg. N. Uribe, VSR. Fruit (CEUA); Envigado, Vereda El Rellano, 6.162140°, -75.563731°, 1800m, 1male (CEUA); Medellín, 6.267404°, -75.585789°, 1570m, 1male, 1female, v.2010, Leg. N. Uribe (CEUA); Cundinamarca: Sopo, 4.916667°, -73.933333°, 2900m, 21.vii.2013, Leg. D. Guzman. Tribe Ceriodini Genus Ceriana Rafinesque Ceria Fabricius, 1794 (4): 277 (preocc. By Scopoli 1863). Type species, Ceria clavicornis Fabricius, 1794 (= Musca conopsoides Linnaeus, 1758) by subs. des. Latreille, 1810:443. Cina Fabricius, 1798:557. Unavailable name cited in generic synonymy. Ceriana Rafinesque, 1815:131 (nom. nov. for Ceria Fabricius). Type species, Ceria clavicornis Fabricius, 1794 = Musca conopsoides Linnaeus, 1758. Refs.: Curran, 1941:243–244 (key to New world spp.); Vockeroth, 1971:282 (notes, syn. and generic status); Thompson, 1972b:129–132 (desc.); Thompson, 1999a:328 (key). Tenthredomyia Shannon, 1925(13):50. Type species, Ceria abbreviata Loew, 1864 (orig. des.). Styloceria Enderlein, 1936:6(1):127. Type species, Musca conopsoides Linnaeus, 1758, by mon. cacica (Walker), 1860:288 (Ceria). Type locality: "Mexico". HT F (BMNH). Distr.: Colombia (Antioquia (Gomez Plata)), Costa Rica, Mexico. Refs.: Thompson et al. 1976:94 (cat.). Material examined. COLOMBIA: Antioquia: Gómez Plata, Hacienda Normandia, 6.683154°, -75.216575°, 1800m, 1male, 29.x.1998, Leg. J.C. Agudelo, Net (MEFLG). Genus Monoceromyia Shannon Monoceromyia Shannon, 1922d:41. Type species, Ceria tricolor Loew (mon.). Refs.: Curran, 1941:243–244 (key to New world spp.); Vockeroth, 1971:282 (notes, syn. and generic status); Thompson, 1972b:129–132 (desc.); Thompson, 1999a:328 (key). vittipes (Curran), 1941:246 (Cerioides). Type locality: Peru, Puerto Bermiidez. HT M (AMNH). Distr. Colombia (Meta (San Juan de Arama)), Peru Refs.: Thompson et al. 1976:94 (cat.). Material examined. COLOMBIA: Meta: San Juan de Arama, 3.350009°, -73.876655°, 420m (ICN). Genus Polybiomyia Shannon Polybiomyia Shannon, 1925:56. Type species, schwarzi Shannon (orig des.). Refs.: Shannon, 1925 (key), Curran, 1941:243–244 (key to New world spp.); Vockeroth, 1971:282 (notes, syn. and generic status); Thompson, 1999a:328 (key). bergrothi (Williston), 1892:77 (Ceria). Type locality: Mexico, Vera Cruz, Atoyac. HT M (AMNH). Distr.: Colombia (Antioquia (San Roque), Cundinamarca), Costa Rica, Mexico. Refs.: Thompson et al. 1976:95 (cat.). Material examined. COLOMBIA: Antioquia: San Roque, Corregimiento San Jose del Nus, 6.503175°, 74.926885°, 1800m, 1male, 5.i.2007, Leg. N. Uribe, Vsr. Liver (CEUA). odontomera (Curran), 1941:244 (Cerioides). Type locality: Brazil, Mato Grosso, Chapada. HT M (AMNH). Distr.: Colombia (Bolívar (Zambrano), Cundinamarca (Tocaima)), Brazil. Refs.: Thompson et al. 1976:95 (cat.). Material examined. COLOMBIA: Bolívar: Zambrano, Hacienda Monterrey, 9.749954°, -74.833324°, 30m, 29.i.1994, Leg. Fernández F. Ulloa (ICN); Cundinamarca: Tocaima, 4.471774°, -74.637839°, 400m, 1male, 13.vi.1984, Leg. W. Jaramillo (UNAB). 488 · Zootaxa 4122 (1) © 2016 Magnolia Press MONTOYA Genus Sphiximorpha Rondani Sphiximorpha Rondani, 1850:212. Type species, Ceria subsessilis Illiger (orig. des.). Refs.: Curran, 1941 (key); Vockeroth, 1971:282 (notes and generic status); Thompson, 1999a:328 (key). Sphyxiviorpha, emend. Cerioides Rondani, 1850:211 (originally cited in synonymy; validated by Becker et al. 1907:156). Type species, Ceria subsessilis Illiger, 1807 (aut.). Ceriodes, Incorrect original spelling. Cerioides, subg. Polistoceria Hull, 1949b: 380. Type species, kerteszi Shannon (orig. des.). roederii (Williston), 1888:289 (Ceria). Type locality: Brazil, Mato Grosso, Chapada. HT M (AMNH). Distr.: Colombia (Caquetá (San Vicente del Caguán), Meta (San Juan de Arama)), Peru, Brazil. Refs.: Thompson et al. 1976:94 (cat.). Material examined. COLOMBIA: Caquetá: San Vicente del Caguán, Vereda Las Vegas, 2.116999°, 74.766711°, 355m, 12.ix.1995, Leg. C. Sarmiento (ICN); Meta: San Juan de Arama, 3.350009°, -73.876655°, 380m, 13.x.1991, Leg. W. C (ICN). rubrobrunnea (Hull), 1944:403 (Cerioides). Type locality: Colombia, Muzo. HT F (NMW). Distr.: Colombia (Boyacá). Refs.: Thompson et al. 1976:94 (cat.). wulpii (Williston), 1888:290 (Ceria). Type locality: Brazil, Mato Grosso, Chapada; Rio de Janeiro. HT M/F (AMNH). Distr.: Colombia (Antioquia (La Pintada, Porce, San Roque), Boyacá (Puerto Colombia), Caldas (Norcacia), Caquetá (San Vicente del Caguán)), Brazil. Refs.: Sack, 1921:149, fig. 25 (puparium); Thompson et al. 1976:94 (cat.). Material examined. COLOMBIA: Antioquia: La Pintada, 5.736521°, -75.632087°, 650m, 2females, 26.v.2007, Leg. N. Uribe, Vsr. Fish (CEUA); Porce, 6.794288°, -75.121970°, 1500m, 1male, ix.1997, A. Smith, Trampa pescado (MEFLG); San Roque, Forest El Topacio, 6.503175°, -74.926885°, 1800m, 1male, Leg. A.L. Montoya, Vsr. Fish (CEUA); Boyacá: Puerto Colombia, Puerto Romero, 6.000040°, -74.416702°, 170m, 25.iv.2001 (ICN); Caldas: Norcacia, Sector El Tigre, Forest, 5.560100°, -74.893349°, 750m, 1male, 22.ii.2010, Leg. L. Rios & D. Gaurisas, Vsr. Fish (CEUA); Caquetá: San Vicente del Caguán, Vereda Las Vegas, 2.116999°, 74.766711°, 355m, Leg. C. Sarmiento (ICN). Tribe Eristalini Genus Dolichogyna Macquart Dolichogyna Macquart, 1842:125 (65). Type species, fasciata Macquart (orig. des.) = chiliensis (Guerin–Meneville). Refs.: Fluke, 1951a (rev, key); Thompson, 1972b:134–135 (description); Thompson, 1999a:329 (key); Thompson, 2000:377 (key). rostrata (Macquart), 1846:259 (131) (Helophilus). Type locality: "Nouvelle–Grenade". HT M (BMNH). Distr.: Colombia. Refs.: Thompson et al. 1976:98 (cat.). Genus Eristalinus Rondani Eristalinus Rondani, 1845:453. Type species, Musca sepulchralis Linnaeus (Subs. mon, Rondani 1857:38). Refs.: Thompson, 1999a:329 (key); 340 (notes on taxa); Thompson, 2003:3, 10 (key, syn. and taxonomic notes). Lathyrophthalmus Mik, 1897:114. Type species, Conops aeneus Scopoli (orig. des.). Lathyrophthalmus, subgenus of Metalloeristalis Kanervo, 1938:43. Type species, Conops aeneus Scopoli (orig. des.). Oreristalis Séguy, 1851:16. Nomen nudum. CATALOGUE OF SYRPHIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 489 Subgenus Eristalodes Mik Eristalodes Mik 1897: 114. Type species, Eristalis taeniopus (Wiedemann), 1818:42. calops (Bigot), 1880:227 (Eristalinus). Type locality: Colombia. HT F (BMNH). Distr.: Colombia. Refs.: Thompson et al. 1976:104 (cat.). fulvitarsis (Rondani), 1863:6 (Eristalinus). Type locality: Colombia. HT M (BMNH). Distr.: Colombia. Refs.: Thompson et al. 1976:105 (cat.). rufitarsis Macquart, 1855:109. (Eristalis; preocc. by Macquart, 1842). Genus Eristalis Latreille Tubifera Meigen, 1800:34. Type species, Musca tenax Linnaeus (Coquillett, 1910:618). Suppressed by I.C.Z.N 1863:339. Elophilus Meigen, 1803:274. Type species, Musca tenax Linnaeus (Latreille, 1810:443). Suppressed by I.C.Z.N 1893:256. Eristalis Latreille, 1804:194. Type species, Musca tenax Linnaeus (Subs. des. by Curtis, 1832:432). Refs.: Thompson et al. 1976:101; Thompson, 1972b:139–140 (desc.); Thompson, 1981:145 (taxonomic notes); Thompson, 1997:209–237 (key, desc. notes and syn.); Thompson, 1999a:330 (key); Thompson, 2003:4, 11 (key, syn. and taxonomic notes). Helophilus Leach, 1817:159, emend. of Elophilus. Cristalis Wiedemann, 1828:10. Misspelling. Eristaloides Rondani, 1845:453. Type species, Musca tenax Linnaeus (Coquillett, 1910:540). Eristalomyia Rondani, 1857:38. Type species, Musca tenax Linnaeus (orig. des.). Eriops Lioy, 1864:743 (Preocc. Klug, 1808). Type species, Musca tenax Linnaeus (Subs. des. by Goffe, 1946:29). Eristalomyia Scudder, 1882:127, emend. of Eristalomya. Subgenus Eoseristalis Kanervo Eoseristalis Karnervo, 1938:40. Type species, Eristalis cerealis Fabricius, 1805:232. bogotensis Macquart, 1842:112 (52). Type locality: Colombia, Santa Fe de Bogota. HT F (MNHNP). Distr.: Colombia (Antioquia (La Ceja, San Pedro, Santa Elena, Yarumal), Cundinamarca, Meta (Puerto Gaitán), Nariño (Pasto, Vereda Gualmatan)), Ecuador, Peru, Venezuela. Refs.: Hull, 1925b:308 (desc.), pi. 11:7 (abdomen), 2:12 (head), 15 (mesonotum); Thompson et al. 1976:101 (cat.); Thompson, 1997:221 (desc. and notes), fig. 13 (M genitalia). assimilis Macquart, 1846:257 (129) (Eristalis). Material examined. COLOMBIA: Antioquia: La Ceja, Alto la Union, Forest, 6.016132°, -75.370182°, 2500m, 10.ix.2008, 1male, Leg. A.L. Montoya, Net (CEUA); San Pedro, Vereda Cerezales, 6.483237°, 75.630955°, 2800m, 1male, Leg. V. Pérez Net in a decomposed pig (CEUA); Santa Elena, 6.212280°, -75.497798°, 2600m, 1male, x.1963, Leg. R. Velez (MEFLG); Turbo, 8.119779°, -76.694502°, 50m, 1male, viii.1956, Leg. F. Gallego (MEFLG); Yarumal, 6.961666°, -75.426785°, 2500m, iv.1982, Leg. J.A. Quiroz, En luz (MEFLG); Meta: Puerto Gaitán, 4.310873°, -72.073758°, 160m, 23.iv.1993, Leg. C. González (ICN); Nariño: Pasto, 1.207852°, 77.277216°, 2800m, 1male, ii.1958, Leg. F. Gallego (MEFLG); Vereda Gualmatan, En arvences, 0.921479°, 77.563786°, 2900m, 1female, 12.iv.2006, Leg. N. Uribe, Net (CEUA). Det F.C. Thompson, 2009. Subgenus Eristalis Latreille tenax (Linnaeus), 1758:591 (Musca). Type locality: "Europe". HT M/F (LSL). Distr.: Colombia (Amazonas (Leticia), Antioquia (Bello, La Ceja, San Pedro de Los Milagros), Boyacá (Aquitania, Duitama, Laguna Pialagua, Páez, Pajarito, Sogamoso), Cundinamarca (Anolaima, Bogotá, Choachí, Facatativá, Guatavita, La Vega, Tocaima), Huila (San Agustín), Magdalena, Meta (Bella Vista, Villavicencio), Nariño (Vereda Sandona), Quindío (Salento), Risaralda (Pereira), Tolima (Melgar, Murillo)), Widespread. Refs.: Williston, 1887:160 (desc.), pi. 7: fig. 7 (wing), 7a (head), 7b (antenna), 7c (hind leg); Thompson et al. 1976:101 (cat.); Thompson, 1981:146 (syn. and notes); Thompson, 1997:214 (desc, and notes), fig 4 (habitus, F, dorsal view), 5 (larva). 490 · Zootaxa 4122 (1) © 2016 Magnolia Press MONTOYA ?columbica Macquart, 1855:108 (88) (Eristalis). Material examined. COLOMBIA: Amazonas: Leticia, Km 7, Tarapaca Ciudad Titama, -4.191131°, 69.941422°, 78m, 2.viii.2007, Leg. Students of Animal Systematic (ICN); Antioquia: Bello, San Félix, 6.333736°, -75.646006°, 2424–3150m, 2males, Leg. C. Bota & N. Uribe, Net (CEUA); La Ceja, Alto la Union, Forest, 6.016132°, -75.370182°, 2500m, 10.ix.2008, 1male, Leg. A.L. Montoya, Net (CEUA); San Pedro de Los Milagros, 6.475640°, -75.616844°, 1male, 20.vi.2001, Leg. V. Pérez (CEUA); Boyacá: Aquitania, Laguna de Tota, 5.583541°, -72.900514°, 3050m, 28.v.1988, Leg. M. García (ICN); Duitama, Paramo de la Rosa, 5.874538°, 73.051187°, 3405m, 28.xi.1978, Leg. I. Arevalo (ICN); Páez, 5.083333°, -73.050000°, 1200m, 20.xi.1973, Leg. E. Cruz (ICN); Pajarito, Comijoque, 5.416672°, -72.666681°, 2000m, 9.iii.1980, Leg. C. Gamboa (ICN); Sogamoso, El Tomate, 5.722996°, -72.923443°, 2549m, 06.iv.2004, Leg. M. Sánchez, Net (UNAB); Cundinamarca: Anolaima, 4.766197°, -74.467662°, 1656m, 13.x.1974, Leg. J. Reyes (UNAB); Bogotá, 4.606156°, -74.077404°, 2600m, 10.xi.1980, Leg. A. Escallón (UNAB); Bogotá, UNal I.C.N, 4.638194°, -74.084046°, 2567m, 7males, 5females, Leg. Fernández–A. & J.L. Fernández–C (ICN); Bogotá, Alto del cable, 4.629103°, -74.049218°, 3100m, 7.vii.1976, Leg. M.L.J.M (ICN); Bogotá, 7.ii.1978, Leg. C. Hernandez (ICN); Choachí, Vereda Resguardo alto, Finca Villa Nueva, 4.529210°, -73.924817° 1827m, 03.v.2008, Leg. E. Herrera, Net (UNAB); Facatativá, Vía Facatativá a Cerro El Hato, 4.814185°, -74.354995°, 2700m, 16.ix.2007, Leg. Fernández–A. & J.L. Fernández–C (ICN); Facatativá, Barrio Tibguesusa, 4.814185°, -74.354995° 2600m, 20.viii.2009, Leg. R. Garzón (ICN); Guaduas, 5.069440°, -74.598052°, 1000m, Leg. M.C. Isaza (ICN); Guatavita, 4.934444°, -73.834444°, 2600m, 16.vi.1974, Leg. L.G. Martínez (ICN); La Calera, 4.727495°, -73.907958°, 2800m, 30.iv.1978, Leg. M.L.J.M (ICN); La Vega, 4.999916°, -74.350009°, 1250m, 8.vi.1972, Leg. G. Cortés (ICN); Tocaima, 4.471774°, 74.637839°, 400m, 9.iii.1973, Leg. Bastid. N. (ICN); Villeta, 5.008319°, -74.473554°, 992m, 30.x.1976, Leg. N. Pinzón (ICN); Huila: San Agustín, 1.927474°, -76.297857°, 1700m, 10 M Liraila, 21.xii.1972, Leg. M.A. Mazorra (ICN); Magdalena: Cienaga, 10.935439°, -74.246833°, 15m, 30.iii.1972, Leg. P.A. Jiménez (ICN); Meta: Bella Vista K.130 Carretera Villavo, 4.139059°, -73.780962°, 1252m, 10.ix.1976, Leg. M. Rodríguez (ICN); Villavicencio, 4.150000°, -73.633333°, 467m, 11.xi.1976, Leg. L. Orozco (ICN); Nariño: Sandona, 1.283333°, 77.466667°, 1970m, 1female, Leg. N. Uribe, Manual (CEUA); Quindio: Salento, Vereda Canaan, 4.613714°, 75.581637°, 1800m, 1female, Leg. A.L. Montoya, Net (CEUA); Risaralda: Pereira, Finca Calamar, 4.885069°, 75.529611°, 2137m, 28.v.1999a, Leg. C. Forero (UNAB); Tolima: Melgar, 4.203889°, -74.642778°, 323m, 2.x.1976, Leg. J. Martínez (ICN); Murillo, Robledal, 4.870242N, 75.177614W, 3000m, 1female, 6.ix.2012, Leg. C. Bota, Net (CEUA). Genus Habromyia Williston Habromyia Williston, 1888:284. Type species, coeruleithorax Williston (mon.). Refs.: Thompson, 1972b:136–137 (des.); Thompson, 1975:292 (status notes); Thompson, 1999a:329 (key); 340 (key to spp. and notes); Thompson, 2000:376 (key). Edwardsietta Hull, 1941i: 437. Lycopale Hull, 1944f: 129. Type species, Merornacrus vittata Hull (orig. des.) = lizeri (Brèthes). Criorthrix Hull, 1949b: 391. Type species, Habromyia rectilinea Hull (orig. des.). magnifica (Bigot), 1880:85 (Plagiocera). Type locality: Colombia. HT M (OXF). Distr.: Colombia (Santander (Carmen de Chucuri)). Refs.: Hull, 1949b: 390, fig. 24h (head); Thompson et al. 1976:101 (cat.); Thompson, 1999a:329 (key). Material examined. COLOMBIA: Santander: Carmen de Chucuri, La Bodega, 6.660277°, -73.384553°, 2300–2400m, 1male, 16.viii.2001, Leg. P. Duque & A. Vélez, Vsr. Fish (CEUA). Genus Lejops Rondani Lejops Rondani, 1857: 33. Type species, Mallota vittata Meigen. Refs.: Curran, 1939a (key); Thompson, 2000:377 (key). Asemosyrphus Bigot, 1882:128. Type species, oculiferus Bigot (Aldrich, 1933:168) = mexicanus (Macquart). Dimorphomyia Bigot, 1885b:173. Type species, calliphoroides Bigot (mon.) = mexicanus (Macquart) (see Aldrich, 1933:168). CATALOGUE OF SYRPHIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 491 Subgenus Asemosyrphus Bigot mexicanus (Macquart), 1842:124 (64) (Helophilus). Type locality: "Mexico". HT M/F (UMO). Distr.: Colombia (Antioquia (Bello, Medellín, San Pedro), Boyacá (Paipa), Caldas (Anserma), Cundinamarca (Anolaima, Bogotá, Cota, Cunuca, Choachí, Cogua, Facatativá, Fontibón, Granada, Guayabal de Síquima, Tocaima, La Vega, Madrid, Mosquera, Sopó, Subachoque, Silvania, Tabio), Quindio (Armenia, Finlandia), Santander (Barbosa), Tolima (Guamo, Líbano)), Mexico. Refs.: Macquart, 1842:124 pi. 11, fig. 2 (habitus), 2a (head); Curran, 1939a:2 (key); Thompson et al. 1976:97 (cat.); Gutierrez et al. 2005. Material examined. COLOMBIA: Antioquia: Bello, San Felix, Paramo de Las Baldias, 6.332653°, 75.646599°, 3050m, 5males, 4females, 17.ii.2013, Leg. A.L. Montoya, Net (CEUA); Medellín, San Antonio de Prado, Vereda Yarumalito, 6.186412°, -75.658630°, 2300m, 1female, 7–8.xi.2008, Leg. C. Bota & N. Uribe, Manual (CEUA); San Pedro, Vereda Cerezales, 6.483237°, -75.630955°, 2800m, 3females, 30.vi. 2001, Leg. V. Pérez, Net in a decomposed pig (CEUA); Caldas: Anserma, Vereda San Pedro, 5.250776°, -75.795434°, 1669m, 1male, 7females, 10.x.2004, Leg. A.L. Montoya (CEUA); Cundinamarca: Anolaima, Balsillas, 4.766197°, 74.467662°, 1656m 18.xi.2003, Leg. R. Matíz, Net (UNAB); Bogotá, Botanic Garden, 4.668332°, -74.100229°, 2551m, 7.vii.2009, Leg. M. Zamora (ICN); Bogotá, UNal I.C.N, 4.638194°, -74.084046°, 2567m, 12.iii.2005, Leg. J.L. Fernández (ICN); Choachí, Vereda Resguardo alto, Finca Villa Nueva, 4.529210°, -73.924817° 1827m, 22.iii.2008, Leg. E. Herrera, Net (UNAB); Cogua, Vereda El Altico, Finca El Oasis, 5.066667°, -73.983333°, 2631m, 07.xi.2004, Leg. K. Cantor, Net (UNAB); Cota, Jerusalen, Finca La Esmeralda, 4.812890°, -74.106085°, 2547m, 07.ii.2005, Leg. D. Carrillo, Malaise trap (UNAB); Facatativá, Vía Facatativá a El Rosal Cerro El Hato, 4.814821°, -74.346605°, 2650m, 9.iii.2008, Leg. C.A. Fernández & A. Fernández (ICN); Granada, Vereda San Raimundo, Finca Los Laureles, 4.521013°, -74.344264°, 2100m, 16.v.2001 (UNAB); Guayabal de Síquima, 4.883144°, -74.466607°, 1630m, 06.iv.2004, Leg. P. Rubio. Z. Castro, Net (UNAB); La Vega, 4.999916°, 74.350009°, 1250m, Km 4, 1215m, 02.iv.1999, Leg. P. Monroy (UNAB); Madrid, 4.737827°, -74.272839°, 2254m, 07.xi.2003, Leg. F. Moreno, Manual (UNAB); Mosquera, Finca Marengo, 4.680641°, -74.216488°, 2546m, 07.v.2004, Leg. B. Maldonado, Net (UNAB); Sopó, Parque Sopó, 4.913452°, -73.939245°, 2650m, 22.vii.2007, Leg. C.A. Fernández & A. Fernández (ICN); Subachoque, Finca Potosi, 4.932714°, -74.183370°, 2663m, 07.xi.2003, Leg. L. Bermúdez (UNAB); Silvania, 4.342730°, -74.404917°, 1600m, 03.x.1996, Leg. S. Fuentes (UNAB); Tabio, Vereda Poveda, 4.920148°, -74.094387° 2589m, 16.v.2000, Leg. H. Sandoval (UNAB); Tocaima, 4.471774°, -74.637839°, 2547m, 03.ii.2005, Leg. D. Carrillo, Malaise trap (UNAB); Boyacá: Paipa, Jardínes, 5.790345°, -73.124558°, 2500m, 20.vii.2006, Leg. Fernández–A. & J.L. Fernández–C (ICN); Quindio: Armenia, Sena, 4.533852°, -75.693470°, 1500m, 18.x.2001, Leg. A. Sarmiento, Net (UNAB); Finlandia, 4.682608°, -75.634175°, 2200m, 2females, 30.ii.2009, Leg. A.L. Montoya, Net (CEUA); Santander: Barbosa, Cite, 5.932922°, -73.621119°, 1800m, 28.viii.2004, Leg. D. Mejía, Net (UNAB); Tolima: Guamo, Vereda Pringamosa, 4.022141°, -74.967567°, 300m, 12.iv.2001, Leg. D. Ávila. A., Sánchez & R. Mora (UNAB); Líbano, 4.920330°, -75.071350°, 1800m, 09.iv.2004, Leg. C. Valbuena, Net (UNAB). Genus Lycastrirhyncha Bigot Lycastrirhyncha Bigot, 1859:307. Type species, nitens Bigot (mon.). Refs.: Doesburg, 1962:31 (key); Thompson, 1972b:145–146 (description); Thompson et al. 1976:102 (cat.); Thompson, 1999a:329 (key); Thompson, 2003:4, 13 (key, syn. and taxonomic notes). Licastrirhyncha Williston, 1908:256. Misspelling. Lycastrirrhyncha Curran, 1934:264. Misspelling. nitens Bigot, 1859:307. Type locality: "Amazonia". HT F (UMO). Distr.: Colombia (Córdoba (Tres Palmas), Chocó (Riosucio)), Brazil, Costa Rica, Ecuador, Guatemala, Guyana, Mexico, Suriname. Refs.: Hull, 1949b:393, fig. 25b (habitus), 25m (head); Thompson et al. 1976:102 (cat.); Montoya et al. 2012 (new record). Material examined. COLOMBIA: Córdoba: Tres Palmas, in Laguna, 8.489913°, -75.949317°, 30m, 1male, 3.i.1975, Leg. R. Vélez (MEFLG); Chocó: Riosucio, Santatá, 7.440560°, -77.118900°, 7m, 1male, 27.vii.1978, Leg. H. Echeverri (ICN). 492 · Zootaxa 4122 (1) © 2016 Magnolia Press MONTOYA Genus Mallota Meigen Mallota Meigen, 1822:377. Type species, Syrphus fuciformis Fabricius (Rondani, 1845:452). Refs.: Curran, 1940:12 (key); Fluke, 1957a:127 (cat.); Thompson, 1972b:138–139 (description); Thompson et al. 1976:99 (cat.); Thompson, 1999a:329 (key); Thompson, 2000:375 (key); Thompson & Zumbado, 2002:89–107 (rev. of Mesoamerican Mallota spp.). Imatisma Macquart, 1842:127 (67). Type species, Eristalis posticatus Fabricius (orig. des.). Subgenus Mallota Meigen aberrans Shannon, 1927:17. Type locality: Peru, Paltaybamba. HT F (USNM). Distr.: Colombia (Huila), Costa Rica, Peru. Refs.: Curran, 1940:12 (key); Thompson et al. 1976:99 (cat.); Thompson & Zumbado, 2002:91 (redesc.), 93, fig 2a–c (M genitalia complex). analis (Macquart), 1846:260 (138) (Imatisma). Type locality: "Nova Grenada". HT F (BMNH). Distr.: Colombia. Refs.: Macquart, 1846:260 (138). pi. 11, fig. 8 (habitus); Thompson et al. 1976:99 (cat.). apis Thompson, 2002:95. Type locality: Costa Rica, San José, Estación Santa Elena, Las Nubes. HT M (INBio). Distr.: Colombia (Antioquia (Betania, Guarne, Rionegro)), Costa Rica, Bolivia, Peru. Refs.: Thompson & Zumbado, 2002:7, fig. 4a–c (M genitalia complex). Material examined. COLOMBIA: Antioquia: Betania, Vereda La Oculta, Betania, Vereda La Oculta, 5.736143°, -76.028237°, 1700-2100m, 1female, 8.viii.1990, Forest, Leg. O. Ortega, Net (Coll. O. Ortega); Guarne, Vereda Piedras Blancas, 6.291588°, -75.493345°, 2660m, Leg. Entomology Group (MEPB); Rionegro, Vereda El Carmín, 6.140929°, -75.411164°, 2100m, 1male, Leg. Entomology Group, Net (CEUA). colombii (Macquart), 1850:445. Type locality: Colombia. HT F (MNHNP). Distr.: Colombia, Peru, Venzuela. Refs.: Macquart, 1850:445 (141), tab. 13, fig. 3 (habitus), 3a (head), 3b (antenna), 3c (hind leg); Thompson et al. 1976:100 (cat.). inversa Shannon, 1927:16. Type locality: Peru, Chanchamayo. HT F (USNM). Distr.: Colombia (Antioquia (Betania, Girardota, Guarne, Rionegro, Sonson), Boyacá (Duitama), Cundinamarca (Usme), Risaralda (Parque Tatamá), Santander (Verda California)), Peru. Refs.: Thompson et al. 1976:100 (cat.). Material examined. COLOMBIA: Antioquia: Betania, Vereda La Oculta, 5.736143°, -76.028237°, 17002100m, 1male, Forest, 8-viii-1990, Leg. O. Ortega, Net (Coll. O. Ortega); Girardota, Vereda El Palmar, 6.343502° -75.435102°, 2200m, 1male, Leg. N. Uribe, Vsr. Fish (CEUA); Guarne, Vereda Piedras Blancas, 6.291588°, 75.493345°, 2200m, 3males, 2female, Leg. Entomology Group (MEPB). Other specimens with the same label information, except for: Vereda Juan Cojo, 1male, Leg. C. Bota, Net (CEUA); Rionegro, Vereda El Carmin, 6.140929°, -75.411164°, 2100m, 5.xi.2005, Leg. N. Franco, Net (CEUA); Sonson, Cerro Las Palomas, 5.537061°, -75.273872°, 3000–3200m, 2males, Leg. C. Bota & L. Ríos, Net (CEUA); Boyacá: Duitama, 5.825391°, 73.040268°, 2530m, 22.vii.2004, Leg. N. Rosas, Manual (UNAB); Cundinamarca: Usme, Parque Entrenubes Cerro Gavilán, 4.522906°, -74.106378°, 2745m, 10.v.2003, Leg. Gasca H. Benavides (ICN); Risaralda: Parque Tatamá, Rio San Rafael, 5.086394°, -76.036662°, 2430m, 8.i.1993, Leg. A. Amarillo (ICN); Santander: California, Vereda California, 7.377812°, -72.938002°, 2800–3000m, 3females, 12–19.ix.2012, Leg. A. Vélez & J.D. Sánchez (CEUA). mystacia Fluke, 1939:372. Type locality: Panama, Chiriqui, El Volcan. HT M (AMNH). Distr. Panama, Colombia (Antioquia (Caldas)). Refs.: Fluke, 1939:372, fig. 12 (head). Material examined. COLOMBIA: Antioquia: Caldas, Vereda Primavera, 6.090000°, -75.638000°, 1560m, 1female, 17.ix.2014, Leg. S. Gutierrez, Manual (Coll. O. Ortega). nigra Shannon, 1927:16. Type locality: Ecuador, Santa Inez. HT F (BMNH). Distr.: Colombia (Caldas (Riosucio)), Ecuador. Refs.: Thompson et al. 1976:100 (cat.). Material examined. COLOMBIA: Caldas: Riosucio, 1male, 2females, 13.vii.2008, C. Gutierrez (Catalina Gutierrez, pers. comm.). CATALOGUE OF SYRPHIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 493 rubricunda Curran, 1940:13. Type locality: Ecuador, Tungurahua, Mt. Tungurahua. HT M (AMNH). Distr.: Colombia (Antioquia (Guarne, Sonson), Cundinamarca (Chingaza, Choachí), Santander (Vereda California)), Ecuador. Refs.: Thompson et al. 1976:100 (cat.). Material examined. COLOMBIA: Antioquia: Guarne, Vereda Piedras Blancas (MEPB); Sonson, Paramo de Sonson, 5.537061°, -75.273872°, 3000m, 1male, 1female, 20.vi.2010, Leg. L. Ríos (CEUA); Cundinamarca: Chingaza, Estacion Monte Redondo, 4.481898°, -73.721867°, 3036m, 5females, 26.x.2002, Leg. E. Martinez, Net–Pig (CEUA); Choachí, Paramo Cruz Verde, 4.544337°, -73.968663°, 3600m, 16.xi.1991, Leg.C. Sarmiento (ICN); Santander: California, Vereda California, 7.377812°, -72.938002°, 2800–3000m, 3males, 12–19.ix.2012, Leg. A. Vélez & J.D. Sánchez (CEUA). salti Curran, 1929:499. Type locality: Colombia, Santa Marta, Vista Nieve. HT F (AMNH). Distr.: Colombia (Magdalena). Refs.: Thompson et al. 1976:100 (cat.). Genus Meromacrus Rondani Plagiocera Macquart, 1842:119 (preocc. Klug 1834). Type species, Milesia cruciger Wiedemann (orig. des.) = acutus Fabricius. Preocc. by Klug, 1834 (Hymenoptera). Meromacrus Rondani, 1848:70. Type species, Meromacrus ghilianii Rondani (mon.). Refs.: Thompson, 1972b:144–145 (description); Thompson, 1999:330 (key); Thompson, 1981:155 (descript, key); Blatch et al. 2003:13–36 (rev. of Mesoamerican Meromacrus spp.); Thompson, 2003:6, 12 (key, syn. and taxonomic notes). Plagiacera Stahl, 1883:97. Misspelling. Pteroptila Loew, 1866:165. Type species, decorus Loew (subs. des. of Coquillett, 1910:598). Promilesia Lynch Arribalzaga, 1892:241. Type species, nectarinoides Lynch Arribalzaga (mon.). Metameromacrus Hull, 1942:1. No Type species, designated. Nomen nudum. Thalamopales Hull, 1949:401. Type species, Helophilus scitus Walker (orig. des.). anna Curran, 1936:4. Type locality: Panama, Canal Zone, Fort Randolph. HT M (AMNH). Distr.: Colombia (Antioquia (San Luis)), Brazil, Costa Rica, Guyana, Panama, Peru. Refs.: Hull, 1942:2 (key ref.); Fluke, 1957a:147 (cat.); Doesburg, 1962:26 (note); Thompson et al. 1976:111 (cat.); Blatch et al. 2003:15 (redesc.) fig. 2 (male abdomen), 7 (M genitalia). Material examined. COLOMBIA: Antioquia: San Luis, 6.041873°, -74.993729°, 1445m, 1male, 1female, Leg. S. López, Manual (CEUA). basiger (Walker), 1860:290 (Eristalis). Type locality: "Amazon Region". HT F (BMNH). Distr.: Colombia (Antioquia (Puerto Berrio, Puerto Nare, San Roque)). Refs.: Sack, 1920:270 (desc.), figs, d (head), f (mesonotum); Thompson et al. 1976:111 (cat.). Material examined. COLOMBIA: Antioquia: Puerto Berrio, Alto de las Aguilas, Hacienda Manaos, 6.500865°, -74.546317°, 404m, 1male, 5.vii.2013, Leg. Entomology Group, Vsr. Fish (CEUA); Puerto Nare, Vereda La Suiza, 6.190087°, -74.587306°, 220m, 1female, Leg. M. Castaño & M.C. Vélez, Pitfall (CEUA); San Roque, Forest El Topacio, 6.503175°, -74.926885°, 1800m, 1male, Leg. A.L. Montoya, Vsr. Fish (CEUA). circumdatus (Bigot), 1875:471 (Sphixea). Type locality: "Colombia". HT F (UMO). Distr.: Colombia, Peru. Refs.: Sack, 1920:263 (description), fig. b (mesonotum). Thompson et al. 1976:112 (cat.). fucatus Hull, 1930:143. Type locality: Surinam, Paramaribo. HT F (ANSP). Distr.: Colombia (Antioquia (Jardín)), Brazil, Suriname. Refs.: Hull, 1942e: ll, fig. 11 (mesonotum); Thompson et al. 1976:112 (cat.). Material examined. COLOMBIA: Antioquia: Jardín, 5.578169°, -75.829912°, 1972m, 1male, 20.v.2011, Leg. J. Cardona (CEUA). laconicus (Walker), 1852:227 (Milesia). Type locality: "Brazil". HT F (BMNH). Distr.: Colombia (Antioquia (Medellín), Magdalena (Santa Marta)), Costa Rica, Ecudar, Panama, Guyana, Venezuela. Refs.: Williston, 1886:324 (citation); Kertész, 1910:330 (cat.); Sack, 1920:268 (mesonotum, descr, key ref.), Sack, 1921:144 (Larva, descr. larva); Hine, 1924:22 (key, descriptive notes); Curran, 1936:2 (taxonomic notes); Hull, 1942:3 (key 494 · Zootaxa 4122 (1) © 2016 Magnolia Press MONTOYA ref.); Fluke, 1957a:154 (species incertis sedis); Thompson et al. 1976:112 (cat.); Thompson, 1981:163 (key ref., taxonomic status); Blatch et al. 2003:24 (redesc.), figs. 6 (abdomen), 10 (M genitalia). Material examined. COLOMBIA: Antioquia: Medellín, 6.267404°, -75.585789°, 1570m, 1male, 1female, 10.i.2004, Leg. S. Lopez (CEUA); Magdalena: Santa Marta, Corregimiento de Taganga, 11.248643°, 74.211974°, 11m, 1male, vi.2004, Leg. A. Ruiz, Manual (CEUA). obscurus Hine, 1924:21. Type locality: Guatemala, Los Amates. HT M (OHSU). Distr.: Colombia (Antioquia (San Roque)), Guatemala, Costa Rica. Refs.: Hull, 1942:2 (as obscura, key ref.); Fluke, 1957a:150 (cat.); Thompson et al. 1976:112 (cat.); Blatch et al. 2003:31 (redesc.), figs. 3 (thorax), 13 (M genitalia). Material examined. COLOMBIA: Antioquia: San Roque, Forest El Topacio, 6.503175°, -74.926885°, 1800m, 1male, Leg. A.L. Montoya, Vsr. Fish (CEUA). scitus (Walker), 1857:152 (Helophilus). Type locality: "Valley of Amazon". HT F (BMNH). Distr.: Colombia (Amazonas (Quebrada Mata Mata–Igapo), Quindio (Finlandia)), Brazil. Refs.: Hull, 1949b:390, fig. 241 (head); Thompson et al. 1976:113 (cat.). Material examined. COLOMBIA: Amazonas: Quebrada Matamata–Igapo, -3.435850°, -70.143596°, 160m, 1.i.1996, Leg. L. Lomanco (ICN); Quindio: Finlandia, 4.685291°, -75.617533°, 2200m, 1male, 30.ii.2009, Leg. A.L. Montoya, Net (CEUA). unicolor Wulp, 1883:131 (Eristalis). Type locality: "Guadeloupe". HT F (ML). Distr. Colombia. Refs.: Thompson et al. 1976:113 (cat.). Genus Palpada Macquart Palpada Macquart, 1834:512. Type species, Palpada scutellata Macquart (mon.) = scutellaris (Fabricius). Refs.: Curran, 1928:42 (key to Puertorican spp.) 1830c:3–6, 134:407–411 (key to World spp. as part of Eristalis); Bean, 1949 (M genitalia); Hull, 1949b:390, fig. 241 (head); Thompson, 1972:142–144 (desc.); Thompson et al. 1976:113 (cat.); Thompson, 1997:209–237 (rev, key and notes); Morales & Marinoni, 2009:301–347 (Cladistic analysis and taxonomic rev. of scutellaris group); Thompson, 2003:4, 13 (key, syn. and taxonomic notes); Mengual & Thompson, 2008 (species group and rev. of ruficeps subgroup). Doliosyrphus Bigot, 1882:120. Type species, Doliosyrphus scutellatus Bigot (subs. des. by Williston, 1887:178) = scutellaris (Fabricius). Dolyosyrphus, Bigot, 1883:228. Misspelling. agrorum species group (= Gymnopalpada Vockeroth) agrorum (Fabricius), 1787:335 (Syrphus). Type locality: "America". HT M/F (UZMC). Distr.: Argentina, Brazil, Colombia (Antioquia (Caldas, Medellin, Sopetran), Bolivar (Pinillos), Cordoba (Tres Palmas)), Honduras, Paraguay Panama, Peru, Venezuela, U.S.A. (Louisiana, Florida), West Indies (Cuba, Puerto Rico). Ref.: Hull, 1925a:38 (description), pi. 1:3 (abdomen), 2:14 (mesonotum); Bean, 1949:143, fig. 4 (M genitalia). Material examined. COLOMBIA: Antioquia: Caldas, 6.080608°, -75.644307°,1900m, ix.1946, Leg. F. Gallego (MEFLG); Medellin, Cerro El volador, Rastrojo bajo, 6.268165°, -75.581523°, 1547m, 3males, i.2008, Leg. A. Carpio (MEFLG); Sopetran, Bosque, ix.1971, Leg. R. Velez (MEFLG); Bolivar: Pinillos, En Maleza, 8.920310°, -74.461586°,14m, viii.1946, Leg. F. Gallego (MEFLG); Cordoba: Tres Palmas, Bosque, 8.488256°, 75.949109°, 30m, 5males, 2females, ix.1972, Leg. R. Velez (MEFLG). albifrons (Wiedemann), 1830:189 (Eristalis). Type locality: "Brazil". HT M (NMW). Distr.: Colombia (Bolivar (Mucura), Magdalena (La Rinconada), Nariño (La Rinconada), Santander (Charalá, Cimitarra), Sucre (San Onofre)), Bahamas, Brazil, Costa Rica, Cuba, Ecuador, Guatemala, Mexico, Puerto Rico, Jamaica, Hispaniola. Refs.: Hull, 1925a:35 (desc.), pi. 1:7 (abdomen); Bean, 1949:144, fig. 3 (M genitalia); Thompson et al. 1976:107 (cat.); Thompson, 1981:151 (synomyms and notes), figs. 177 (male abdomen, dorsal) 180 (head,males, dorsal view). Material examined. COLOMBIA: Bolivar: Mucura, Isla de San Bernardo, 9.781757°, -75.872394°, 6m, CATALOGUE OF SYRPHIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 495 2males, 1female, 6.i.2003, Leg. M. Wolff & P. Duque, Net (CEUA); Magdalena: La Rinconada, 9.231884°, 74.492215°, 17m, Leg. C. Elster (ICN); Nariño: La Rinconada, 1.507414°, -77.112540°, 2300m, Leg. C. Elster (ICN); Santander: Charalá, Virolin, 6.304190°, -73.173909°, 1800m, 1.iii.1981 (ICN); Cimitarra, Corregimiento Puerto Olaya, Central Termocentro–Isagen, Humedal Norte, 6.473122°, -74.395383°, 110m, 15.vi.2005, 1female, Leg. M. Cataño & C. Rivera, Light trap (CEUA); Sucre: San Onofre, 9.646080°, -75.581129°, 5m, 4males, 1female, Leg. C. Velásquez (CEUA). atrimana (Loew), 1866a:167 (also 1872b:41) (Eristalis). Type locality: "Cuba". HT F (MCZ). Distr.: Colombia (Antioquia (Caucasia, San Luis, San Roque, Maceo, Puerto Berrio), Caldas (Norcacia), Chocó (Rio Truandó), Huila (Obando), Santander (Cimitarra)), Bahamas, Brazil, Costa Rica, Cuba, Ecuador, Guatemala, Jamaica, Hispaniola, Mexico, Puerto Rico. Refs.: Williston, 1887:173 (desc.); Curran, 1930c:16 (desc.); Thompson et al. 1976:103 (cat.); Thompson, 1981:151 (synomyms and notes). Material examined. COLOMBIA: Antioquia: Caucasia, Barrio El Triangulo, 7.983333°, -75.200000°, 50m, 1female, 18.v.2010, Leg. O. Sierra, Net (CEUA); Maceo, Reserva Cañón del Río Alicante, 6.540547°, 74.770327°, 840m, 1female, Leg. J. Vélez, Manual (CEUA); San Luis, Cañon de Rio Claro, Forest, 5.894165°, 74.856963°, 350m, 1female, 26.xi.2009, Leg. I. Ceballos, Vsr. Fish (CEUA); San Roque, Corregimiento San Jose del Nus, 6.503175°, -74.926885°, 1800m, 1male, 8.ii.2009, Leg. A.L. Montoya, Net (CEUA); Puerto Berrio, 6.489819°, -74.401992°, 114m, 1male, Leg. A.L. Montoya, Manual (CEUA); Caldas: Norcacia, 5.560100°, 74.893349°, 750m, 1female, Leg. Norberto, Manual (CEUA); Chocó: Rio Truandó, 7.149891°, -77.199761°, 100m, 10.i.1957 (ICN); Huila: Obando, 1.926307°, -76.300314°, 1850m, 1female, Leg. A.L. Montoya, Net (CEUA); Santander: Cimitarra, Corregimiento Puerto Olaya, 6.473122°, -74.395383°, 110m, 1male, 1female, Leg. G. Zapata, Net (CEUA). cosmia (Schiner), 1868:362 (Eristalis). Type locality: "Venezuela". HT M/F (NMW). Distr.: Colombia (Antioquia (Caldas, Guarne, Medellín)), Venezuela. Refs.: Thompson et al. 1976:104 (cat.). Material examined. COLOMBIA: Antioquia: Caldas, Alto La Romera, 6.090000°, -75.638000°, 1560m, 1male, Leg. E. Alarcón (CEUA); Guarne, Vereda Piedras Blancas, Parque Piedras Blancas, 6.291588°, 75.493345°, 2300m, 1female, 11.vi.2004, Leg. A. Velez (CEUA); Medellín, 6.267404°, -75.585789°, 1570m, 1female, Leg. E. Alarcón, Net (CEUA). Det F.C. Thompson, 2009. elegans (Blanchard), 1852:406 (Eristalis). Type locality: Chile, Coquimbo. HT M/F (MNHNP). Distr.: Colombia (Antioquia (La Ceja, San Pedro)). Refs.: Hull, 1925a:26 (desc.), pi. 2:13 (abdomen), 16 (mesonotum); Thompson et al. 1976:105 (cat.). Material examined. COLOMBIA: Antioquia: La Ceja, Alto la Union, Forest, 6.016132°, -75.370182°, 2500m, 1male, 1female, Leg. A. Bustamante, Net (CEUA); San Pedro, Vereda Cerezales, 6.483237°, -75.630955°, 2800m, 1male, 30.vi. 2001, Leg. V. Pérez, Net in a decomposed pig (CEUA). expicta (Walker), 1860:290 (Eristalis). Type locality: "Mexico". HT M (BMNH). Distr.: Colombia (Antioquia (La Ceja, Puerto Berrio, San Roque)), Costa Rica, Mexico. Refs.: Thompson et al. 1976:105 (cat.). Material examined. COLOMBIA: Antioquia: La Ceja, Alto la Union, Forest, 6.016132°, -75.370182°, 2500m, 1male, Leg. A.L. Montoya, Net (CEUA); Puerto Berrio, 6.489819°, -74.401992°, 114m, 1male, Leg. A. Bustamante, Net (CEUA); San Roque, Corregimiento San Jose del Nus, 6.503175°, -74.926885°, 1800m, 1male, Leg. A.L. Montoya, Net (CEUA). florea (Hull), 1925a:39 (Eristalis). Type locality: Guyana, Arakaba & Bartica, Kartabo. HT M (OHSU). Distr.: Colombia (Antioquia (Anorí, San Roque), Caqueta (Vereda Sebastopolo)), Costa Rica, Guyana, Mexico, Panama, Trinidad. Refs.: Hull, 1925a:39, pi. 1:3 (abdomen); Thompson et al. 1976:105 (cat.). Material examined. COLOMBIA: Antioquia: Anorí, 7.073631°, -75.146379°, 1546m, 1female, 1female, Leg. Bustamante, Net (CEUA); San Roque, Vereda El Iris, 6.503175°, -74.926885°, 1800m, 1female, 10.iv.2014, Leg. Entomology Group, Pitfall copro (CEUA); Caqueta: Florencia, Vereda Sebastopolo, Finca La Magoloa, 1.598268°, -75.543651°, 305m, 2females, 25.xii.2004, Leg. Entomology Group, Net–Pig (CEUA). Det F.C. Thompson, 2009. 496 · Zootaxa 4122 (1) © 2016 Magnolia Press MONTOYA minutalis (Williston), 1891:64 (Eristalis). Type locality: Mexico, Tabasco, Teapa. HT M/F (BMNH). Distr.: Colombia, Mexico. Refs.: Williston, 1891:64, pi. 2:6 (habitus), 6a (head); Hull 1925a:42 (desc.); Thompson et al. 1976:107 (cat.). monticola (Roder), 1892:9 (Eristalis; nom. nov. for montana Roder). Type locality: Colombia. ST F (MLUH). Distr.: Colombia (Boyacá). Refs.: Roder, 1892:9, fig. 1 (habitus); Thompson et al. 1976:107 (cat.). montana Roder, 1886:263 (Eristalis; preocc. Williston, 1882). ochracea (Williston), 1888:279 (Eristalis). Type locality: Brazil, Mato Grosso, Chapada. HT M (AMNH). Distr.: Colombia, Brazil, Costa Rica, Mexico, Guyana. Refs.: Thompson et al. 1976:107 (cat.). pusilla (Macquart), 1842:114 (54) (Eristalis). Type locality: French Guiana, Mana River. HT M (MNHNP). Distr.: Colombia, Argentina, Costa Rica. Refs.: Hull, 1925a:36 (description), pi. 1:2; Bean, 1949:144, fig. 2 (M genitalia) (abdomen). 2:17 (mesonotum); Thompson et al. 1976:108 (cat.); Thompson, 1981:152 (synomyms and notes). rufiventris (Macquart), 1846:257 (129) (Eristalis). Type locality: "Colombia". HT M/F (UMO). Distr.: Colombia (Antioquia (Maceo, Puerto Berrio, San Luis), Costa Rica. Refs.: Hull, 1925a:33 (desc.), pi. 1:4 (abdomen); Thompson et al. 1976:108 (cat.). Material examined. COLOMBIA: Antioquia: Maceo, Reserva Cañón del Río Alicante, 6.540547°, 74.770327°, 840m, 1female, Leg. A.L. Montoya, Net (CEUA); Puerto Berrio, 6.489819°, -74.401992°, 114m, 1female, Leg. A.L. Montoya, Net (CEUA); San Luis, Rio Claro, 5.894165°, -74.856963°, 600m, 1male, Leg. Entomology Group, Net (CEUA). taenia (Wiedemann), 1830:174 (Eristalis). Type locality: Uruguay, Montevideo. HT M (AMNH). Distr.: Colombia (Antioquia (La Ceja, San Roque), Caldas (Anserma), Quindio (Filandia)), Brazil, Venezuela. Refs.: Hull, 1925a:38 (desc.); Curran, 1930c:13 (descriptive notes); Thompson et al. 1976:109 (cat.). Material examined. COLOMBIA: Antioquia: La Ceja, Alto la Union, Forest, 6.016132°, -75.370182°, 2500m, 2males, 2females, 5.xii.2007, Leg. A.L. Montoya (CEUA); San Roque, Corregimiento San Jose del Nus, 6.503175°, -74.926885°, 1800m, 1male, 10.ix.2008, Leg. A.L. Montoya (CEUA); Caldas: Anserma, Vereda San Pedro, 5.250776°, -75.795434°, 1669m, 1male, 10.x.2004, Leg. E. Alarcón (CEUA); Quindio: Filandia, Hacienda Bengala, 4.685291°, -75.617533°, 2000m, 1male, 10.ix.2008, Leg. A.L. Montoya (CEUA). thalia (Hull), 1942a:86 (Eristalis). Type locality: Colombia, Dept. Cundinamarca, Bogota. HT M (MCZ). Distr.: Colombia (Cundinamarca). Refs.: Thompson et al. 1976:110 (cat.). vierecki (Curran), 1930c:10 (Eristalis). Type locality: Colombia, Vista Nieve. HT M (AMNH). Distr.: Colombia (Bolivar), Costa Rica, Bolivia. Refs.: Thompson et al. 1976:110 (cat.). scutellaris species group (Palpada sensu stricto)) conica (Fabricius), 1805:190 (Milesia). Type locality: "America meridional". HT M/F (UZMC). Distr.: Colombia (Amazonas, Antioquia (Alejandria, Guarne), Chocó (Quibdó, Riosucio), Cundinamarca (Medina), La Guajira (El Jagüey), Meta (Puerto López, Villavicencio), Tolima (Chaparral, Melgar)), Bolivia, Brazil, Costa Rica, Ecuador, Guatemala, Guyana, Panama, Peru. Refs.: Hull, 1925b:290 (desc.), pi. 1:6 (abdomen); Thompson et al. 1976:104 (cat.); Morales & Marinoni, 2009:318 (desc, syn.), figs 20–21, 45–46 (head, frontal and lateral), 61–62 (Thorax, dorsal), 83, 91, 107 (hind leg), 131 (wing), 142, 157 (abdomen), 163 186 189, 220 (M genitalia), 242, 251, 263 (F, M genitalia). Material examined. COLOMBIA: Amazonas: 1.iv.1975, Leg. J. Idrobo (ICN); Antioquia: Alejandria, Vereda El Carbón, 6.403715°, -75.064663°, 1445m, 1male, Leg. Entomology Group, Net (CEUA); Guarne, Vereda La Peña, 6.286535°, -75.423037°, 2800m, 1male, Leg. V. Pérez, Net in decaying pig (CEUA); Chocó: Quibdó, 5.693715°, -76.654892°, 37m, 21.iii.1982, Leg. J. M. Idrobo (ICN); Riosucio, Tilupo, 7.440560°, -77.118900°, 7m, 6.iv.1978, Leg. H. Echeverri (ICN); Riosucio, Santatá, 7.440560°, -77.118900°, 7m, 27.iv.1978, Leg. H. Echeverri CATALOGUE OF SYRPHIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 497 (ICN); Cundinamarca: Medina, Vereda Choapal, 4.508319°, -73.361189°, 440m, 26.vii.1986, Leg. Students of Animal Systematic (ICN); La Guajira: El Jagüey, 10.626761°, -71.705081°, 71m, 1.vi.1981, Leg. J. Rincón (ICN); Meta: Puerto López, Vereda Bajo Menegua, 4.066137°, -73.048348°, 181m, 18.iv.1984, Leg. E.A. (ICN); Villavicencio, Vereda La Vanguardia, Pozo Azul, 4.150000°, -73.633333°, 467m, 16.iv.2005 (ICN); Tolima: Chaparral, 3.750000°, -75.583333°, 1354m, 90.xii.1972, Leg. L. Quiñones (ICN); Melgar, 4.203889°, 74.642778°, 323m, 24.iv.1976, Leg. M.R Leon (ICN). erratica (Curran), 1930c:19 (Eristalis). Type locality: Panama, Canal Zone, Barro Colorado Island. HT M (AMNH). Distr.: Colombia (Antioquia, Magdalena, Valle del Cauca, Vichada), Bolivia, Brazil, Costa Rica, Ecucador, Guyana, Panama, Peru, Panama, Trinidad, Venzuela. Refs.: Thompson et al. 1976:105 (cat.); Morales & Marinoni, 2009:325 (desc, syn.), figs 11–12 (habitus), 49 (head, lateral), 65 (Thorax, dorsal), 84, 92 (hind leg), 133 (wing), 144, 159 (abdomen), 165 188, 201, 222 (M genitalia), 244, 253, 262 (F M genitalia). scutellaris (Fabricius), 1805:190 (Milesia). Type locality: "America meridionali". HT M/F (UZMC). Distr.: Colombia (Antioquia (Andes, Carepa, Fredonia, Ituango, La Pintada, San Roque, Santa Bárbara), Caldas (Riosucio), Meta (Vista hermosa), Brazil, Bolivia, Costa Rica, Ecuador, Guyana, Mexico, Peru, Panama, Suriname, Trinidad, Venezuela. Refs.: Macquart, 1834:513, pi. 11, fig. 19; Williston, 1887:178 pi. 8:8 (head); Hull, 1925b:291 (desc.), pi. 2:16 18 (head); Doesburg, 1962:23, fig. 1 (habitus), la (head), lb (hind leg), 2, 4 (puparium), 3 (habitus); Thompson et al. 1976:109 (cat.); Morales & Marinoni, 2009:343 (desc, syn.), figs. 37–39 (head, dorsal, lateral), 73 (thorax, dorsal), 78 (scutellum, lateral), 89, 95, 118 (hind leg), 138 (wing), 150, 153 (abdomen) 180 184, 207, 228 (M genitalia), 248, 257, 262 (F M genitalia). Material examined. COLOMBIA: Antioquia: Andes, Corregimiento de Santa Rita, 5.550249°, 75.988733°, 3600m, 2males, 5.xii.2009, Leg. J.D. Sanchez & A.L. Montoya, Net (CEUA); Carepa, Estacion Tulenapa, Corpoica, 7.759545°, -76.654668°, 48m, 2males, 8.ii.2009, Leg.Duque, Net (CEUA); Fredonia, 5.916573°, -75.683347°, 1010m, 1male, Leg. Entomology Group, Net (CEUA); Ituango, 7.166667°, -75.750000°, 1600m, 1male, 15.ii.1989, Leg. J. Garces, Net. (CEUA); La Pintada, Hacienda Montenegro, 5.736521°, 75.632087°, 650m, 1male, 1female, Leg. Velez & Grisales, Net (CEUA); San Roque, Corregimiento San Jose del Nus, 6.503175°, -74.926885°, 1800m, 1male, Leg. M. Castaño, Net (CEUA); Santa Bárbara, 5.882086°, 75.575273°, 1846m, 14.vi.1993, Leg. G. Jiménez (UNAB); Caldas: Riosucio, Cerro Ingurumá, 5.421334°, 75.719539°, 2200m, 1female, 7.i.2013, Leg. A.L. Montoya (CEUA); Meta: Vista Hermosa, Caño Blanco, 2.744073°, -73.748008°, 250m, 25.iii.1997, Leg. C. Bojacá (UNAB). vinetorum species group (= Trichopalpada Vockeroth). aemula (Williston), 1891:64 (Eristalis). Type locality: Mexico, Guerrero: Rio Papagaio, 1200 ft.; Chilpancingo, 4600 ft.; Rincon, 2800 ft.; Venta de Zopilot 2800 ft.; Guatemala, San Geronimo; Panama: Bugaba. HT M (BMNH). Distr.: Colombia (Antioquia (San Roque), Meta), Costa Rica, Ecuador, Mexico, Venezuela. Refs.: Williston, 1891:64, pi. 2:5 (habitus); Hull, 1925b:288 (desc.), pi. 1:1 (abdomen), 2:18 (hind leg); Thompson et al. 1976:111 (cat.). Material examined. COLOMBIA: Antioquia: San Roque, Corregimiento San Jose del Nus, 6.503175°, 74.926885°, 1800m, 1female, Leg. M. Castaño, Net (CEUA). amazon (Curran), 1930c:23 (Eristalis). Type locality: Brazil, Amazonas, Rio Caiari–Uaupes. HT F (AMNH). Distr.: Colombia (Antioquia (Puerto Berrio), Caldas (Norcacia)), Brazil, Venezuela. Refs.: Thompson et al. 1976:103 (cat.). Material examined. COLOMBIA: Antioquia: Puerto Berrio, Barrio Lamalesna, 6.489819°, -74.401992°, 114m, 1male, Leg. Pérez & Vélez, Net (CEUA); Puerto Berrio, 6.489819°, -74.401992°, 114m, 1female, Leg. M. Castaño, Net (CEUA); Caldas: Norcacia, 5.560100°, -74.893349°, 900m, 1male, Leg. Pérez & Vélez, Net (CEUA). flavoscutellata (Hull), 1937a:173 (Eristalis). Type locality: Costa Rica, Hiquito, San Mateo. HT F (USNM). Distr.: Colombia (Valle del Cauca), Costa Rica. Refs.: Hull, 1937a:173, figs. 1 (tip of abdomen), 7 (hind leg); Thompson et al. 1976:105 (cat.). 498 · Zootaxa 4122 (1) © 2016 Magnolia Press MONTOYA furcata (Wiedemann), 1819:51 (Eristalis). Type locality: "Brazil". HT M/F (NMW). Distr.: Colombia (Antioquia (Amaga, Caldas, Itagui, La Ceja), Boyacá (Buenavista, Gachetá, Paipa), Caldas (Anserma, Samaná), Cundinamarca (Bogotá, Fusagasugá, Guayabal de Síquima), Huila (Obando), Magdalena (Santa Marta), Meta (Villavo), Quindío, Risaralda, Santander (Charalá, Piedecuesta), Tolima (Espinal, Ibagué), Valle del Cauca (Darien, Restrepo, Villa Carmelo)), Argentina, Brazil. Refs.: Macquart, 1842:100 (40), pi. 3, fig. 6 (head); Hull, 1925a:29 (desc.), pi. 2:11 (abdomen), 20 (mesonotum); Thompson et al. 1976:106 (cat.). Material examined. COLOMBIA: Antioquia: Amaga, 6.050000°, 1 -75.700000°, 546m, 1female, Leg. A.L. Montoya, Net (CEUA); Caldas, Alto La Romera, 6.090000°, -75.638000°, 1560m, 1male, Leg. A.L. Montoya, Net (CEUA); Itagüí, Vereda La María, 6.189489°, -75.627675°, 2546m, 1female, Leg. A.L. Montoya, Net (CEUA); La Ceja, Alto la Union, Forest, 6.016132°, -75.370182°, 2500m, 1female, Leg. A.L. Montoya, Net (CEUA); Boyacá: Buenavista, Vereda Miraflores, 5.650832°, -74.286905°, 800m, 03.xi.2003, Leg. D. Melo, Manual (UNAB); Gachetá, Vereda Salinas, 4.820836°, -73.636574°, 1850m, 07.iv.2004, Leg. O. Lanchero (UNAB); Paipa, 5.790345°, -73.124558°, 2517m, 03.viii.1999, Leg. Cely, Forero & Rojas (UNAB); Caldas: Anserma, Vereda San Pedro, 5.250776°, -75.795434°, 1669m, 1male, Leg. A.L. Montoya, Net (CEUA); Samaná, Corregimiento de Norcasia, La Laguna, 5.538340°, -74.969231°, 980m, 9.xi.2003, Leg. A. Rodríguez (UNAB); Cundinamarca: Bogotá, 4.598411°, -74.076182°, 2600m, 20.vi.1976, Leg. A. Bernal (ICN); Tena, 4.624207°, -74.239034°, 1340m, 27.iv.1991, Leg. A. Castillo (ICN); Fusagasugá, 4.338328°, -74.363968°, 1850m, 12.iv.2000, Leg. Estévez (UNAB); Guayabal de Síquima, Vereda El Resguardo, 4.883333°, -74.466667°, 1630m, 2005, Leg. F. Cruz, Net (UNAB); Huila: Obando, Estrecho del Magdalena, 1.926307°, -76.300314°, 1800m, 1male, Leg. A.L. Montoya, Net (CEUA); Magdalena: Santa Marta, Guachaca, 11.248643°, -74.211974°, 11m, 4.xi.1976, Leg. J. Blanco (ICN); Meta: Villavo, 4.139059°, -73.780962°, 1500m, 03.viii.1979, Leg. J.L. Pinto (ICN); Santander: Charalá, Rio Virolin, 6.304190°, -73.173909°, 1800m, 1840m, 8.xii.1978, Leg. I. Arevalo (ICN); Piedecuesta, 6.987834°, 73.047684°, 1100m, 12.x.1978, Leg. J. Caballero (ICN); Tolima: Espinal, 4.162687°, -74.874972°, 322m, 24.xi.2003, Leg.F. Moreno, Manual (UNAB); Ibagué, 4.440663°, -75.244141°, 1285m, 05.i.1999, Leg. Cely, Forero & Rojas (UNAB); Valle del Cauca: Darién camino Rio Azul, Rio Bravo, 3.935850°, -76.675300°, 1300m, 8.ii.1984, Leg. C. Valdés (ICN); Restrepo, Camp. Rio Azul, 3.882081°, -76.573971°, 1500m, 7.iii.1984, Leg. D. Acosta (ICN). geniculata (Fabricius), 1805:237 (Eristalis). Type locality: "America meridionali". HT M (UZMC). Distr.: Colombia (Sucre (San Onofre)), Brazil. Refs.: Stephens, 1846:29, pi. 47:3 (habitus); Thompson et al. 1976:106 (cat.). Material examined. COLOMBIA: Sucre: San Onofre, 9.646080°, -75.581129°, 5m, 2males, 1female, Leg. Entomology Group, Net (CEUA). Det F.C. Thompson, 2009. maculata (Sack), 1941:112 (Eristalis). Type locality: Peru, Pachitea River. HT M/F (SMT). Distr.: Colombia (Chocó (Atrata Valley, Boca de Arquia)), Ecuador, Peru. Refs.: Thompson et al. 1976:106 (cat.). mexicana (Macquart), 1847:75 (59) (Eristalis). Type locality: "Mexico". HT M/F (BMNH). Distr.: Colombia (Antioquia (Cocorná, Caldas, El Peñol, Guarne, Itagui, Medellín, San Roque), Sucre, Valle del Cauca), Ecuador, Peru. Refs.: Thompson et al. 1976:107 (cat.). Material examined. COLOMBIA: Antioquia: Cocorná, 6.050187°, -75.166752°, 830m, 2female, Leg. Entomology Group, Net (CEUA); Caldas, 6.090000°, -75.638000°, 1560m, 1male, Leg. Entomology Group, Net (CEUA); El Peñol, 6.218496°, -75.243396°, 1420m, 1female, Leg. A.L. Montoya, Net (CEUA); Guarne, Vereda La Peña, 6.286535°, -75.423037°, 2800m, 1female, Leg. A.L. Montoya, Net (CEUA); Itagüí, Vereda La María, 6.189489°, -75.627675°, 1800, 1female, Leg. C. Bota, Net (CEUA); Medellín, Corregimiento Santa Elena, 6.212280°, -75.497798°, 1830m, 1male, Leg. Entomology Group, Net (CEUA); San Roque, Corregimiento San Jose del Nus, 6.503175°, -74.926885°, 1800m, 1male, Leg. A.L. Montoya, Net (CEUA). Det F.C. Thompson, 2009. mirabilis (Hull), 1925b:305 (Eristalis). Type locality: Colombia, Vista Nieve. HT F (AMNH). Distr.: Colombia (Antioquia (Caldas, El Carmen de Viboral, Girardota, Itagui, Medellín, Montebello), Cundinamarca (Chía, Fusagasugá, La Mesa), Huila (Gigante), Magdalena (Santa Marta), Meta (Puerto Gaitán, Restrepo), Risaralda (Santa Rosa de Cabal)), Paraguay. Refs.: Hull, 1925b:305, pi. 2:11 (mesonotum); Thompson et al. 1976:107 (cat.). CATALOGUE OF SYRPHIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 499 Material examined. COLOMBIA: Antioquia: Caldas, 6.090000°, -75.638000°, 1560m, 1male, Leg. A.L. Montoya, Net (CEUA); El Carmen de Viboral, Vereda El Carmen, 6.083333°, -75.333333°, 2100m, Leg. M. Zuluaga, Manual (CEUA); Itagüí, Vereda La María, 6.189489°, -75.627675°, 2546m, 1male, Leg. A.L. Montoya, Net (CEUA); Girardota, Vereda Juan Cojo, 6.343502° -75.435102°, 2200m, 1female, Leg. C. Bota, Net (CEUA); Medellín, Corregimiento San Antonio de Prado, Vereda Potrerito, 6.186412°, -75.658630°, 2300m, 1female, Leg. Bustamante, Net (CEUA); Medellín, 6.267404°, -75.585789°, 1570m, 16.ii.1981, Leg. A. Reyes (ICN); Montebello, Vereda El Guabial, 5.916667°, -75.500000°, 1250m, 5.v.2010, Leg. BACL (CEUA); Cundinamarca: Chía, Cerro La valvanera, 4.861565°, -74.032534°, 2582m, 21.iii.2004, Leg. O. Lugo, Net (UNAB); La Mesa, Laguna Pedro Palo, 4.629429°, -74.469603°, 1200m, 27.ii.1997, Leg. L. Castillo (UNAB); Fusagasugá, 4.338328°, -74.363968°, 1771m, 15.vii.1971, Leg. G. Baquero (ICN); La Mesa, 4.629469°, -74.466633°, 1000m, 10.v.1972, Leg. J. Carretero (ICN); Útica, 5.xi.1965 (ICN); Huila: Gigante, 2.386942°, -75.546096°, 800m, 16.xi.1971, Leg. A. Sobrino (ICN); Meta: Puerto Gaitán, 4.310873°, -72.073758°, 160m, 1.v.1993, Leg. C. González (ICN); Restrepo, 4.243144°, -73.573992°, 560m, 11.ix.1974, Leg. A. Bernal (ICN); Risaralda: Santa Rosa de Cabal, Termales de Santa Rosa, 4.837708°, -75.548600°, 2000m, 10.vi.2013, Leg. A.L. Montoya, Net (CEUA). vinetorum (Fabricius), 1798:562 (Syrphus; as vinctorum, emended by Fabricius 1799:48). Type locality: "America Insulis". HT M/F (UZMC). Distr.: Colombia (Amazonas (Mata Mata), Antioquia (Alejandria, Maceo, San Roque), Boyacá (Berbeo), Caqueta, Cundinamarca (La Mesa, Madrid, Villeta), Guainía (Puerto Inirida El Salario), Huila (Gigante), Meta (Bella vista, Guayabetal, San Juan de Arama), Santander (Bucaramanga, Cimitarra), Sucre (San Onofre), Valle del Cauca (Restrepo)), Argentina, Costa Rica, Cuba, Ecuadro, Guatemala, Hispaniola, Jamaica, Less Antilles, Puerto Rico. Refs.: De Geer 1776:145, pi. 29, fig. 1 (habitus); Williston, 1887:171 (desc.), pi. 7:8a (hind leg); Hull, 1925a:40 (description), pi. 1:9 (abdomen); Bean, 1949:146, fig. 8 (M genitalia); Thompson et al. 1976:110 (cat.); Thompson, 1981:153 (syn. and notes), figs. 170 (hind leg of male) 182 (wing); Thompson, 1990:15–16 (distributional notes). Material examined. COLOMBIA: Antioquia: Alejandria, Maceo, San Roque. Santander: Cimitarra, Corregimiento Puerto Olaya, Central termocentro–Isagen, Humedal Norte, 6.473122°, -74.395383°, 110m, 15.vi.2005, 1female, Leg. M. Cataño & C. Rivera, Light trap (CEUA); Sucre: San Onofre, Reserva Natural Sanguare, 9.646080°, -75.581129°, 5m, 3males, 2females, 27.v.2009, Leg. A.L. Montoya, Net (CEUA). ruficeps subgroup prietorum Mengual, 2008:32. Type locality: Colombia, Valle del Cauca, Cali, Cerro San Antonio. HT M (USNM). Distr.: Colombia (Valle del Cauca). Refs.: Mengual & Thompson, 2008:32 (orig. des.), pi 4, figs. 2 (habitus, dorsal), 5 (habitus, lateral), 6 (wing), 7 (M genitalia). ruficeps (Macquart), 1842:111 (51) (Eristalis). Type locality: Colombia, Santa Fe de Bogota. HT M (MNHNP). Distr.: Colombia (Antioquia (La Ceja, Sonson, San Jose de la Montaña), Boyacá (Miraflores, Tunja, Villa de Leyva), Caldas (Manizales), Cauca (Guayas), Cundinamarca (Bogotá, Guatavita, La Mesa, Quetame, San miguel), Meta (Puerto Gaitán), Tolima (Ibagué)), Ecuador, Mexico, Venezuela. Refs.: Hull, 1925b:307 (desc.), pi. 2:10 (head); Thompson et al. 1976:108 (cat.); Mengual & Thompson, 2008:34 (key and notes), fig. 1 (habitus, dorsal). Material examined. COLOMBIA: Antioquia: La Ceja, Alto la Union, Forest, 6.016132°, -75.370182°, 2500m, 1female, 10.ix.2008, Leg. A.L. Montoya, Net (CEUA); Sonson, Cerro Las Palomas, 5.537061°, 75.273872°, 3000m, 1male, 1female, 19.ix.2009, Leg. L. Rios, Net (CEUA); San Jose de la Montaña, Vereda el Congo sector la Laguna, Potrero, 6.804244°, -75.689613°, 3100–3200m, 7females, 10–14.ix.2011, Leg. L. Rios, Net (CEUA); Boyacá: Miraflores, 5.196389°,-73.144444°, 1820m, 05.vi.1989, Leg. F. Ramírez (UNAB); Tunja, Santuario Iguaque Quebrada La Laguna, 6.044559°, -73.154046°, 2340m, 5.viii.1990, Leg. A. Amarillo (ICN); Villa de Leyva, 5.638191°, -73.535870°, 2200m, 1.xi.2001, Leg. M. Alvarez (ICN); Caldas: Manizales, 5.072373°, -75.506634°, 2216m, Leg. F. Rodríguez (UNAB); Cauca: Guayas, 2.673707°, -76.959274°, 1446m, 8.xi.1976, Leg. R.M. Gómez (ICN); Cundinamarca: Bogotá, 4.626403°, -74.04548°, 2600m, 5.x.1968, Leg. P. Wilches (ICN); Bogotá, Alto del cable, 4.629103°, -74.049218°, 3100m, 5females, 7.vii.1978, Leg. MLJM (ICN); Guatavita, 4.934444°, -73.834444°, 2900m, 30.x.1999, Leg. L. Bérmudez (UNAB); La Mesa, 4.629469°, 74.466633°, 1298m, 15.v.1994, Leg. G. Rincón (UNAB); Quetame, Corregimiento El Calvario, 4.333333°, - 500 · Zootaxa 4122 (1) © 2016 Magnolia Press MONTOYA 73.850000°, 1880m, 11.vi.1979, Leg. C. Vergara (ICN); San miguel, 5.550000°, -73.750000°, 2750m, 30.iv.1941 (ICN); Meta: Puerto Gaitán, 4.310873°, -72.073758°, 160m, 1.v.1977, Leg. C. González (ICN); Tolima: Ibagué, 4.440663°, -75.244141°, 1300m, 1.ii.1970, Leg. T. De los Reyes (ICN). suprarufa Thompson, 1999a:344. Type locality: Ecuador, S. Otavalo. HT M (MZUSP). Distr.: Colombia (Antioquia (Urrao)), Ecuador. Refs.: Mengual & Thompson, 2008:34, figs. 3 (Key and habitus, dorsal). Material examined. COLOMBIA: Antioquia: Urrao, Paramo de Frontino, 6.416558°, -76.107506°, 3500m, 1female, 6.i.1991, Leg. G. Abril (MEFLG). Genus Quichuana Knab Quichuana Knab, 1813:13. Type species, sylvicola Knab (orig. des.). Refs.: Hull, 1946a (rev, key); Thompson, 1972:135–136 (desc.); Thompson et al. 1976:98 (cat.); Thompson, 1981:143 (key and taxonomic notes); Thompson, 1999a:329 (key); Thompson, 2000:375 (key); Ricarte et al. 2012:74 (taxonomic rev. and keys). Quihuana, Hull, 1943h:510. Misspelling. angustiventris (Macquart), 1855:110 (Merodon). Type locality: ‘Valley of the Amazon’. HT M (BNHM). Distr.: Colombia (Antioquia (San Luis), Chocó, Caldas, Cundinamarca), Brazil, Costa Rica, Peru Trinidad, Venezuela, and Surinam. Refs.: Macquart, 1855:110 (90), pi. 5, fig. 2 (head); Thompson et al. 1976:98 (cat.); Ricarte et al. 2012:74 (desc., key), figs. 5–6 (male genitlia), 73 (habitus, males). Material examined. COLOMBIA: Antioquia: San Luis, Rio Claro, 5.894165°, -74.856963°, 350m, 1male Leg. Entomology Group, Net (CEUA). brevicera Hull, 1946a:7. Type locality: Colombia, Atrata Valley, Boca de Arquia. HT M (BMNH). Distr.: Colombia (Chocó). Refs.: Thompson et al. 1976:98 (cat.); Ricarte et al. 2012:83 (description, status notes and key). inca var. brevicera Hull, 1946b. calathea Shannon, 1925b:111. Type locality: Panama, Porto Bello. HT M/F (USNM). Distr.: Colombia (Antioquia (Nariño, San Luis, San Roque), Chocó, Risaralda, Quindio (Quimbaya), Risaralda (Pereira), Valle del Cauca (Caicedonia, Restrepo), Costa Rica, Cuba, Panama. Refs.: Hull, 1946a:16, fig. 13 (head) (desc. note); Thompson et al. 1976:98 (cat.); Ricarte et al. 2012:85 (desc., key), figs. 74 (male head, dorsal view). Material examined. COLOMBIA: Antioquia: Nariño, Vereda Puente Linda, 5.631484°, -75.245454°, 2400m, 2females, Leg. L. Rios, Net (CEUA); San Luis, Rio Claro, 5.894165°, -74.856963°, 350m, 1male, 2females, Leg. A. Vélez, Vsr. Fish (CEUA); San Roque, Corregimiento San Jose del Nus, 6.503175°, -74.926885°, 1800m, 1female, Leg. N. Uribe, Vsr. Liver (CEUA); Quindio: Quimbaya, Vereda Kerman, Km 7, 4.614947°, 75.807598°, 1339m, 9males, 3females, Leg. H. Vargas, Vsr. Fish (CEUA); Risaralda: Pereira, Wildlife Sanctuary Otún Quimbaya, 4.728880°, -75.577850°, 1890m, 1female, vi.2010, Leg. C. Bota, Vsr. Fish (CEUA); Valle del Cauca: Caicedonia, 4.340810°, -75.832995°, 1400m, 16.x.2001, Leg. M. Gullen (UNAB); Restrepo, Camino Rio Bravo, Campo Alegre, 3.811560°, -76.515178°, 1000m, 4.ii.1984, Leg. O. Cepeda (ICN). dolorosa Hull, 1946:6. Type locality: Bolivia-Mapiri. HT M (AMNH). Distr.: Bolivia, Colombia (Risaralda (Belen de Umbria)). Refs.: Thompson et al. 1976:98 (cat.); Ricarte et al. 2012:91 (desc., key). Material examined. COLOMBIA: Risaralda: Belen de Umbria, Vereda La Isla, 5.137655°, -75.830656°, 1000m, 1male, 8.ii.2015, Leg. A.L. Montoya, rearing from Bromeliad (CEUA). fasciata (Sack), 1941:115 (also 1951:111) (Myiatropa). Type locality: Peru, Paso de Lares. HT M/F (SMT). Distr.: Colombia (Boyacá), Costa Rica, Panama, Peru. Refs.: Thompson et al. 1976:98 (cat.); Ricarte et al. 2012:92 (desc., key), figs. 11 (antenna, F laterarl view), 77 (F LT, habitus, dorsal view). picadoi Knab, 1913:14. Type locality: "Costa Rica". HT F (USNM). Distr.: Costa Rica, Colombia (Antioquia (Hispania, Medellín)), Surinam. Refs.: Hull, 1946a: 16, figs. 12 & 25 (head), 17 (antenna); Thompson et al. 1976:99 (cat.); Ricarte et al. 2012:106 (desc., key), figs 75-76 (M head, anterior, lateral view). CATALOGUE OF SYRPHIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 501 Material examined. COLOMBIA: Antioquia: Hispania, Finca La fé, 5.799893°, -75.899989°, 922m, 2females, 20.v.2008, Leg. J.D. Suaza, reared from a water tank of Guadua (CEUA); Medellín, 6.267404°, 75.585789°, 1570m, Universidad de Antioquia, 2male, 1female, 11-xi.2008, Leg. A.L. Montoya, reared from Xanthosoma sp. (CEUA). quixotea Hull, 1946a:14, fig. 6 (head). Type locality: "Colombia". HT M/F (CNC). Distr.: Colombia (Boyacá (Muzo), Santander (Olaya)), Costa Rica. Refs.: Thompson et al. 1976:99 (cat.); Ricarte et al. 2012:109 (description, key). Tribe Merodontini Genus Eumerus Meigen Eumerus Meigen, 1822:202. Type species, Syrphus tricolor Fabricius (Curtis, 1839:749). Genus–group name conserved by ICZN (1993, Opinion 1747). Refs.: Latta & Cole, 1933 (rev, key); Thompson et al. 1976:92 (cat.); Thompson, 1999a:332; Thompson, 2008:3 (key), fig. 15 (wing). funeralis (Meigen), 1822:208. Type locality: Germany & Austria. HT F (MNHNP). Distr.: Colombia (Magdalena). Refs.: Thompson et al. 1976:92 (cat.) Tribe Milesiini Genus Cacoceria Hull Cacomyia Hull, 1930:147 (preocc. Coquillett, 1906). Type species, cressoni Hull (orig. des.). Cacoceria Hull, 1936a:227 (nom. nov. for cacomyia Hull). Refs.: Thompson, 1968:272 (key and notes); Thompson, 1972:179–180 (desc, synonms); Thompson, 1999a:333 (key). willistoni Hull, 1949d:225. Type locality: Peru, Chanchamayo. HT M (CNC). Distr.: Colombia (Antioquia (El Retiro, Jardin), Risaralda (Pereira)), Argentina, Peru. Refs.: Thompson et al. 1976:91 (cat.). Material examined. COLOMBIA: Antioquia: El Retiro, 6.054442°, -75.501817°, 2150m, Trampa de Emergencia, 1female, Leg. M. Wolff (CEUA); Jardin, 5.584248°, -75.832689°, 2200m, 1female, ii.1980, Leg. A. Molina P. (MEFLG); Risaralda: Pereira, Parque Natural Nacional los Nevados, 4.872737°, -75.426886°, 2900m, 1female, Leg. Bustamante, Net (CEUA). Genus Chalcosyrphus Curran Chalcosyrphus Curran, 1925:122 (Proposed as a subg. of Chalcomyia). Type species, atra Curran (orig. des.) = Chalcomyia despressa Shannon. Refs.: Curran, 1941 (key); Thompson, 1972:157–158 (desc, synonms); Thompson, 1975:292 (status notes); Thompson et al. 1976:114 (cat.); Thompson, 1999a:336 (key). Subg. Xylotomina Shannon, 1926:7. Type species, Xylota vecors Osten Sacken (orgi. des.). As first revisor, Thompson (1975) select Xylotomina as senior to Xylotodes. Planes Rondani, 1863:9 (preocc. Bowditch, 1825; Saussure, 1862). Type species, Xylota vagans Wiedemann (mon.). Xylotodes Shannon, 1926:7. Type species, Brachypalpus inarmatus Hunter (orgi. des.). Neplas Porter, 1927b:96 (nom. nov. for Planes Rondani). Type species, Xylota vagans Wiedmann (aut.). Cheiroxylota Hull, 1949:361. Type species, Xylota dimidiate Brunetti (orgi. des.). americana (Schiner), 1868:367. Type locality: South America. HT M/F (NMW). Distr.: Colombia. Refs.: Thompson et al. 1976:115 (cat.). 502 · Zootaxa 4122 (1) © 2016 Magnolia Press MONTOYA vagabondans (Hull), 1941c:62 (Planes). Type locality: Colombia, Restrepo. HT M (CNC). Distr.: Colombia (Meta). Refs.: Thompson et al. 1976:115 (cat.). Genus Macrometopia Philippi Macrometopia Philippi, 1865:740. Type species, atra Philippi (mon.). Refs.: Hull, 1949b:365, fig. 221 (head); Thompson, 1972:152–153 (description); Thompson, 1999a:333 (key), 347 (species key). maculipennis Thompson, 1999a:348. Type locality: Colombia, West Cordillera, Monte Socorro. HT F (BMNH). Distr.: Colombia (Valle del Cauca), Peru. Refs.: Thompson, 1999a:347 (key), 348 (orig. des.), pi 378, fig 29 (wing). Genus Meropidia Hippa & Thompson Meropidia Hippa & Thompson, 1983:111. Type species, neurostigma (mon.). Refs.: Hippa & Thompson, 1982:111–115; Thompson, 1999a:335 (key); Morales et al. 2013. flavens Hippa & Ståhls, 2013: 60. Type locality: Bogota. HT F (NMNH). Distr.: Colombia (Antioquia (Andes), Cundinamarca). Refs.: Morales et al. 2013: 60–62, Fig. 9–15 (head, anterior and lateral view, habitus dorsa and lateral). Material examined. COLOMBIA: Antioquia: Andes, Vereda Quebrada arriba, Forest pescado, 5.600370°, 75.965892°, 2100-2500m, 1female, 26-29-ii-1997, Leg. O. Ortega (Coll. O. Ortega). neurostigma Hippa, 1982:111. Type locality: Bolivia, Cochabamba, Incacha. HT M (CNC). Distr.: Colombia (Antioquia (Andes)), Bolivia. Refs.: Hippa, 1982:111 (orig. des.), pi 112, fig. 1–4 (M genitalia complex), 5 (thoraxic pleuron, lateral view), 6 (hind leg, lateral view), 7 (wing), 8 (face,males, frontal view), 9 (facial profile, female), 10 (head,males, lateral view), 115, fig 11 (abdomen), 12 (thorax). Material examined. COLOMBIA: Antioquia: Andes, Paramo de Santa Rita, 5.550249°, -75.988733°, 3600m, 1male, 2females, 8.xii.2010, Leg. A.L. Montoya, Net (CEUA). nigropilosa Thompson, 1983:113. Type locality: Colombia, West Cordillera, Monte Socorro. HT F (BMNH). Distr.: Colombia (Valle del Cauca). Refs.: Hippa, 1982:113 (orig. des.). Genus Spilomyia Meigen Spilomyia Meigen, 1803:273. Type species, Musca diophthalma Linnaeus (Coquillett, 1910: 617). Ref.: Curran, 1951 (key). gratiosa Wulp, 1888: 372. Type locality: Argentina, Tucuman Prov. HT F (lost?). Distr. Brazil (Sao Paulo), Colombia (Risaralda (Pererira)), Argentina. Refs.: Thompson, 1996. Material examined. COLOMBIA: Risaralda: Pererira, PNR Ucumari, La Florida, 4.708555°, -75.487224°, 2600m, vii.1990, Leg. L. Sneider (ICN). Genus Sterphus Philippi Sterphus Philippi, 1865:737. Type species, autumnalis Philippi (orig. des.) = coeruleus Rondani. Refs.: Kertész, 1910 (cat.); Shannon, 1926:45 (cat.); Shannon & Aubertin, 1933:155 (desc. notes, distr. data); Hull, 1949:369 (desc.); Thompson, 1972:153–154 (desc, synonms); Thompson, 1973:1185–240 (phylogenetic rev, keys and taxonomic notes); Thompson, 1999a:333–334 (key). Crepidomyia Shannon, 1926a:47. Type species, tricrepis Shannon (orig. des.) = jamaicensis Gmelin. Tatuomyia Shannon, 1926:48. Type species, batesi Shannon, 1926 (orig. des.). CATALOGUE OF SYRPHIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 503 Senoceria Hull, 1930:144. Type species, spinifemorata Hull, 1930 (orig. des.) = Xylota coarctata Wiedemann, 1830. Stherphus, Fluke, 1957a:106. Misspelling. Subgenus Ceriogaster Williston Ceriogaster Williston 1888: 285. Type species, Ceriogaster foscithorax Williston, 1888: 286 chlorpygus species group chloropyga (Schiner), 1868:366 (Xylota). Type locality: "Columbien" (=Venezuela). HT M/F (NMW). Distr.: Colombia (Antioquia (Andes), Santander (Vereda California)), Ecuador, Venezuela. Refs.: Fluke, 1957a:157 (cat.); Thompson, 1973:212 (redesc.); Thompson et al. 1976:117 (cat.). Material examined. COLOMBIA: Antioquia: Andes, Vereda Quebrada arriba, Forest pescado, 5.600370°, 75.965892°, 2100-2500m, 1female, 26-29-ii-1997, Leg. O. Ortega (Coll. O. Ortega); Santander: California, Vereda California, 7.377812°, -72.938002°, 2800–3000m, 1male, 12–19.ix.2012, Leg. A. Vélez & J.D. Sánchez (CEUA). coarctatus species group coarctatus (Wiedemann), 1830:100 (Xylota). Type locality: "Brazil". HT F (SMF). Distr.: Colombia (Antioquia (Maceo, Puerto Berrio), Brazil, Costa Rica, Panama. Refs.: Hull, 1930:146, figs. 2 (mesonotum), 5 (head), 6 (hind leg); Fluke, 1957a:105 (cat.); Thompson, 1973:187 (head, lateral view) 181 (key) 185, fig. 14 (wing of typic population), 15 (wing of Panama Population), 20 (abdomen), pi 219 (redesc.), pi 230, fig. 33 (M genitalia); Thompson et al. 1976:117 (cat.); Zumbado & Thompson, 1997:79, 86 (notes, and key). Material examined. COLOMBIA: Antioquia: Maceo, 6.540547°, -74.770327°, 840m, 1male, Leg. M. Wolff, Vsr. Fish (CEUA); Puerto Berrio, Alto de las Aguilas, Hacienda Manaos, 6.500865°, -74.546317°, 404m, 1male, 5.vii.2013, Leg. M. Wolff, Vsr. Fish (CEUA). transversa (Walker), 1857:153 (Syritta). Type locality: "Valley of Amazon". Distr.: HT M (BMNH). Distr.: Colombia (Sucre (San Onofre)), Valley of Amazon. Refs.: Thompson et al. 1976:116 (cat.). Material examined. COLOMBIA: Sucre: San Onofre, 9.646080°, -75.581129°, 5m, 1female, 11.x.2003, Leg. C. Rivera & P. Duque, Net (CEUA). cybele species group andicus Hippa, 1978:136. Type locality: Bolivia, Coroico, 1200m. HT F (BMNH). Distr.: Bolivia, Colombia (Santander (Vereda California)). Refs.: Hippa, 1978:136 (cat.). Material examined. COLOMBIA: Santander: California, Vereda California, 7.377812°, -72.938002°, 2800–3000m, 2male, 1 female, 12–19.ix.2012, Leg. A. Vélez & J.D. Sánchez (CEUA). aureus Hippa, 1978:137. Type locality: Colombia. Monte Socorro, 3500 m. HT F (BMNH). Distr.: Colombia (Guarne), Valle del Cauca). Refs.: Hippa, 1978:137 (cat.). Material examined. COLOMBIA: Antioquia: Guarne, Vereda Piedras Blancas, Parque Piedras Blancas, 6.291588°, -75.493345°, 2200m, 2300m, 2females, 29.x.2004, Leg. A. Velez, Manual (CEUA). cybele (Hull), 1951c:184 (Crepidomyia). Type locality: Peru, Chanchamayo. HT F (CNC). Distr.: Colombia (Cundinamarca), Peru. Refs.: Fluke, 1957a:106 (cat.); Thompson, 1973:216 (redesc.); Thompson et al. 1976:117 (cat.); Hippa & Thompson, 1994:486 (distributional record), fig 6 (hind leg), pi 488, 14 (antenna, female). cydippe Hippa, 1994:189. Type locality: Colombia, West Cordillera, Monte Socorro, 3800m. HT M/F (BMNH). Distr.: Colombia (Antioquia (Betania), Valle del Cauca). Refs.: Hippa & Thompson, 1994:486, fig. 4 (hind leg), 487, fig. 7 (M genitalia), 488, fig. 11 (antenna) 18 (aedeagus), 489 (orig. desc.), fig. 22 (abdomen). Material examined. COLOMBIA: Antioquia: Betania, Vereda La Oculta, 5.736143°, -76.028237°, 17002100m, 2male, Forest, 8-viii-1990 Leg. O. Ortega, Net (Coll. O. Ortega). 504 · Zootaxa 4122 (1) © 2016 Magnolia Press MONTOYA fassli Hippa, 1978:138 Type locality: Colombia, West Cordillera, Monte Socorro, 3500 m. HT F (BMNH). Distr.: Colombia (Antioquia (Andes, Sonson), Valle del Cauca)). Refs.: Hippa, 1978:137 (cat.). Material examined. COLOMBIA: Antioquia: Andes, Vereda Quebrada arriba, Forest pescado, 5.600370°, 75.965892°, 2100-2500m, 1female, 26-29-ii-1997, Leg. O. Ortega (Coll. O. Ortega); Sonson, Cerro Las Palomas, 5.537061°, -75.273872°, 3045m, 1female, Leg. A. Cardona & D. Alvarez, Pitfall (CEUA). icarum Hippa, 1978 Type locality: Colombia, West Cordillera, Monte Socorro, 3800m. HT F (BMNH). Distr.: Colombia (Santander (Vereda California), Valle del Cauca). Refs.: Hippa, 1978:140 (cat.). Material examined. COLOMBIA: Santander: California, Vereda California, 7.377812°, -72.938002°, 2800–3000m, 1male, 12–19.ix.2012, Leg. A. Vélez & J.D. Sánchez (CEUA). janzeni Thompson, 1994:491. Type locality: Costa Rica, Puntarenas Prov., Monteverde. HT M (INBio). Distr.: Costa Rica, Colombia (Antioquia (Medellín)), Panama. Refs.: Thompson, 1994:491, fig 5 (Hind femur, lateral view.), 9 (M genitalia, lateral view.), 13 (F Antenna), 18 (M Aedeagus), 20, 24 (F, M abdomen dorsal view). Material examined. COLOMBIA: Antioquia: Medellín, Vereda Piedras Blancas, 6.291588°, -75.493345°, 2200m, 1female, 19.vi.2012, Leg. G. Entomological, Net (MEPB). nitidicollis Hippa, 1978:140. Type locality: Colombia, West Cordillera, Monte Socorro, 3800m. HT F (BMNH). Distr.: Colombia (Valle del Cauca). Refs.: Hippa, 1978:140 (cat.). ochripes Hippa, 1978:142. Type locality: Colombia, West Cordillera, Monte Socorro, 3800m. HT F (BMNH). Distr.: Colombia (Valle del Cauca). Refs.: Hippa, 1978:142 (cat.). pilifer Hippa, 1978:140. Type locality: Colombia, West Cordillera, Monte Socorro, 3800m. HT F (BMNH). Distr.: Colombia (Valle del Cauca). Refs.: Hippa, 1978:20 (cat.). sapphirifer Hippa, 1978:142. Type locality: Colombia, West Cordillera, Monte Socorro, 3800m. HT M (BMNH). Distr.: Colombia (Antioquia (Caldas, Guarne), Valle del Cauca). Refs.: Hippa, 1978:142 (cat.). Material examined. COLOMBIA: Antioquia: Caldas, 6.090000°, -75.638000°, 1560m, 1female, ix.1973, Leg. R. Velez, En maleza (MEFLG); Guarne, Vereda El Placer, Robledal, 6.291562°,-75.497416°, 2300m, 1male, 1–5.iii.2007, Leg. A. Velez, Vsr. Fish (CEUA). fascithorax species group fascithorax Kertész, 1910:86 (as foscithorax; emend. by Kertész, 1910:308; Curran, 1930f:10). Type locality: Brazil, Mato Grosso: Corumba & Rio de Janeiro. HT M/F (AMNH). Distr.: Colombia (Antioquia), Brazil. Refs.: Thompson et al. 1976:115 (cat.). plagiatus species group jamaicensis (Gmelin), 1790:427 (Musca). Type locality: Jamaica. HT M/F (?). Distr.: Colombia (Antioquia (Alejandria, Caldas, Girardota, Medellín)), Costa Rica, Cuba, Bahamas, Bolivia, Jamaica, Less Antillas, Peru, Puerto Rico. Refs.: Thompson, 1973:196, fig. 18 (abdomen), 224, fig. 31 (M genitalia), pi 227 (desc, synonymy); Thompson et al. 1976:39 (cat.); Thompson, 1981:169, fig 187 (male head, lateral view) 183 (syn. and desc.); Zumbado & Thompson, 1997:82, 86 (notes and key). Material examined. COLOMBIA: Antioquia: Alejandria, Vereda El Carbón, 6.403715°, -75.064663°, 1800m, 2males, Leg. A.L. Montoya, Net (CEUA); Caldas, Alto La Romera, 6.090000°, -75.638000°, 1560m, 2females, Leg. A.L. Montoya, Net (CEUA); Girardota, Vereda La Honda, 6.343502° -75.435102°, 2200m, 400m, 1male, Leg. N. Uribe, Vsr. Fish (CEUA); Medellín, Vereda Piedras Blancas, en Robledal, 6.291588°, -75.493345°, 2200m, 1female, 26.ix.2009, Leg. C. Bota, Net (CEUA). CATALOGUE OF SYRPHIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 505 plagiatus Wiedemann, 1830:98 (Xylota). Type locality: Brazil. HT F (NMW). Distr.: Colombia (Antioquia (Maceo)), Brazil. Refs.: Kertész, 1910:304 (cat.); Shannon, 1926:48 (desc. note, distr, rec.); Curran, 1934:404 (notes on type, distr. rec); Fluke, 1957a:106 (cat.); Thompson, 1973:220, fig. 13 (wing). Material examined. COLOMBIA: Antioquia: Maceo, Reserva Cañón del Rio Alicante, 6.540547°, 74.770327°, 840m, 1female, Leg. S. Perez, Net (CEUA). Tribe Rhingini Genus Rhingia Scopoli Rhingia Scopoli, 1763:358. Type species, Conops rostrata Linnaeus (mon.). Refs.: Thompson, 1972:102 (description); Thompson, 1999a:333 (key). longirostris Fluke, 1943:430. Type locality: Ecuador, Bolivar, Hacienda Talahua. HT M (AMNH). Distr.: Colombia (Antioquia (Jardín, La Ceja, Liborina, Medellín, San Jose de la Montaña), Nariño (Barbacoas), Risaralda (Pereira)), Ecuador. Refs.: Fluke, 1943:430, figs. 10 (wing tip), 11 (head); Thompson et al. 1976:69 (cat.). Material examined. COLOMBIA: Antioquia: Jardín, Reserva Natural Cuchilla Jardín Támesis, 5.531351°, -75.821373°, 2700–2800m, Leg. Vélez, Pérez & Vargas, Pitfall (CEUA); Jardín, 5.531351°, -75.821373°, 2400m, 1female, Leg. A.L. Montoya, Net (CEUA); La Ceja, Alto de la Union, 6.016132°, -75.370182°, 2500m, 1male, Leg. Entomology Group, Net (CEUA); Liborina, Vereda El Playon, Finca La Pava, 6.665323°, -75.719095°, 30003197m, 1male, 5-8.xi.2011, Leg. J.D. Castaño & C. Bota, Net (CEUA); Medellín, San Antonio de Prado Vereda Yarumalito, 6.186412°, -75.658630°, 2300m, 1male, Leg. Entomology Group, Net in decaying pig P3 (CEUA); San Jose de la Montaña, Vereda el Congo sector la Laguna, Potrero, 6.804244°, -75.689613°, 3100–3200m, 2females, 10–14.ix.2011, Leg. L. Rios, Net (CEUA); Nariño: Barbacoas, Via Sandona, Quebrada Chacaguaico, 1.651397°, -78.163467°, 93m, 1female, Leg. N. Uribe, Manual (CEUA); Risaralda: Pereira, Wildlife Sanctuary Otún Quimbaya, 4.728880°, -75.577850°, 1890m, 1female, vi.2010, Leg. C. Bota, Vsr. Fish (CEUA). nigra Macquart, 1846:261. Type locality: "Colombia". HT M (UMO). Distr.: Colombia (Antioquia (Caldas, Medellín, San Jose de la Montaña), Cundinamarca (Bogotá), Huila (San Agustín), Risaralda (Pereira)), Costa Rica, Ecuador, Mexico. Refs.: Macquart, 1846:261 (133), tab. 11:10 (head); Fluke, 1943:427, fig. 12 (head); Thompson et al. 1976:69 (cat.). Material examined. COLOMBIA: Antioquia: Caldas, 6.090000°, -75.638000°, 1560m, 05.ii.1998, Leg. R. Landinez, Net (UNAB); Jardin, Vereda La Herrera, 5.568645°, -75.737518°, 2050m, 1female, 4.vi.2011, Leg. C.M Arenas, Net (CEUA); Medellín, Barrio Belén, 6.216630°, -75.599930°, 1500m, 1female, Leg. A.L. Montoya, Net (CEUA); San Jose de la Montaña, Vereda el Congo, 6.804244°, -75.689613°, 3100–3200m, 1female, 10–14.ix.2011, Leg. L. Rios, Malaise trap (CEUA); Cundinamarca: Bogotá, Carretera Bogotá La Mesa. Sitio Sabrinsky, 4.629469°, -74.466633°, 1000m, 20.iv.1984, Leg. M. García (ICN); Huila: San Agustín, 1.927474°, 76.297857°, 1700m, 1female, 18.v.2006, Leg. A. L. Montoya (CEUA); Risaralda: Pereira, PNR Ucumari, La Pastora, 4.708555°, -75.487224°, 2500m, 6.vii.1990, Leg. C.S. Suárez (ICN). Tribe Volucellini Genus Copestylum Macquart Copestylum Macquart, 1846:252 (124). Type species, flaviventris Macquart (mon.) = marginatum Say. Refs.: Curran, 1930d (keys), 1935 (key to marginatum group), 1939a (key), l947b (key to vesicularium group), 1953 (keys to abdominal and esuriens groups); Fluke, 1953 (key to scutellatum group); Thompson, 1972:108 (desc. and relationships), 111 (key); Thompson, 1999a:330, 332 (key); Rotheray et al. 2007 (key and desc.); Restrepo & Carrejo, 2009:141–150 (distrib. records). Phalacromya Rondani, 1848:67. Type species, submetallica Rondani (Coquillett, 1910:587). Phalacromyia, emend. 506 · Zootaxa 4122 (1) © 2016 Magnolia Press MONTOYA Glaurotricha Thomson, 1869:493. Type species, muscaria Thomson (mon.). Haurotricha Thomson, 1869:493. Incorrect original spelling of Glaurotricha Thomson (1st reviser, Bigot, 1883:250). Atemnocera Bigot, 1882:114. Type species, Volucella scutellata Macquart (mon.). Apophysophora Williston, 1888:276. Type species, scutellata Williston (Curran, 1930d:5) = trituberculatum Thompson. Megametopon Giglio Tos, 1891:5. Type species, Ophromyia nasica Williston (Williston, 1892:79 & Giglio Tos, 1892e:44). Ophromyia Williston, 1891:55. Type species, nasica Williston (mon.). Camerania Giglio Tos, 1892a:3. Type species, macrocephada Giglio Tos (as Temnocera megacephala? Loew (orig. des.). Viereckomyia Curran, 1925b:246. Type species, Temnocera gibbera Schiner (orig. des.). Volucella, subg. Volosyrpha Shannon, 1928:575. Type species, tibialis Shannon (orig. des.) = rufitarse Thompson. Volucellosia Curran, 1930d:5. Type specie Volucella fornax Townsend (orig. des.). Subgenus Copestylum Macquart flaviventre Macquart, 1846:253 (125). Type locality: "Colombia, Venezuela". HT M/F (UMO). Distr.: Colombia, Costa Rica, Mexico, Venezuela. Refs.: Macquart, 1846:253 (125), pi. 10, figs. 16 (habitus), 16a (head); Thompson et al. 1976:75 (cat.). Subgenus Lepidopsis Curran viride (Williston), 1888:275 (Volucella). Type locality: Brazil, Mato Grosso, Chapada. HT M F (AMNH). Distr.: Colombia (Amazonas, Antioquia (Puerto Berrio), Meta), Brazil, Costa Rica, Peru. Refs.: Thompson et al. 1976:87 (cat.); Restrepo & Carrejo, 2009:149 (cat.). Material examined. COLOMBIA: Antioquia: Puerto Berrio, 6.489819°, -74.401992°, 114m, 1male, 24.ix.2005, Leg. Entomology Group (MEPB). Subgenus Megametopon (= Ophromyia) Giglio Tos amethystinum (Bigot), 1875:479 (Volucella, as amethistina; emend.: Williston, 1891:52). Type locality: "Mexico". HT M (UMO). Distr.: Colombia (Cauca, Nariño, Risaralda), Brazil, Costa Rica, Mexico. Refs.: Thompson et al. 1976:71 (cat.); Restrepo & Carrejo, 2009:146 (cat.). Subgenus Phalacromya Rondani apicale (Loew), 1866a:151 (1872b:25) (Volucella). Type locality: "Cuba". HT M (MCZ). Distr.: Colombia (Magdalena (Santa Marta)), Costa Rica, Cuba, Jamaica, Less Antillas, Puerto Rico. Refs.: Thompson et al. 1976:71 (cat.). Material examined. COLOMBIA: Magdalena: Santa Marta, PNN Tayrona Neguanje, 11.276050°, 74.195519°, 121m, 1.viii.1996, Leg. F. Escobar (ICN). aster (Curran), 1939b:3 (Volucella). Type locality: Peru, Piches and Perene Valleys. HT F (AMNH). Distr.: Colombia (Antioquia (Alejandria, Bolivar, Guarne, Medellín, San Vicente), Boyacá, Caldas (Pensilvania), Caqueta, Huila, Magdalena, Nariño, Risaralda (Pereira), Santander (Guavatá)), Peru. Refs.: Thompson et al. 1976:71 (cat.); Restrepo & Carrejo, 2009:146 (cat.). Material examined. COLOMBIA: Antioquia: Alejandria, Vereda El Carbón, 6.403715°, -75.064663°, 1400m, Leg. A.L. Montoya, Vsr. Fruit (CEUA); Bolivar, Farallones de Citara, Vereda La Mina, Rio Buen Aventón, 5.772073°, -76.032002°, 2400m, 27.iii.20011, Leg. D. Ramirez, Net (CEUA); Gurane, Vereda Piedras Blancas, 6.291588°, -75.493345°, 2300m, 1female, Leg. A. Velez, Net (MEPB); Medellín, Corregimiento San Antonio de Prado, Vereda Potrerito, 6.186412°, -75.658630°, 2300m, 1female, Leg. M. Castaño, Net (CEUA); San Vicente, Vereda El Chaparal, 6.257308°, -75.357379°, 1800m, 2females, Leg. A.L. Montoya, Net (CEUA); Caldas: Pensilvania, Vereda La Rioja, 5.384070°, -75.160990°, 1956m, Leg. C. Bota & C. Flórez, Net (CEUA); Risaralda: Pereira, Reserva Ucumari, El Cedral, 4.708555°, -75.487224°, 2500m, 1female, Leg. A. Bustamante, Net (CEUA); Santander: Guavatá, 5.950028°, -73.700151°, 1870m, 14.vi.1999, Leg. R. Gómez (UNAB). CATALOGUE OF SYRPHIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 507 aureum (Fluke), 1951b:13 (Volucella). Type locality: Colombia, Cundinamarca, Zipacoa. HT F (AMNH). Distr.: Colombia (Boyacá (Villa de Leyva). Refs.: Fluke, 1951b:13, figs. 6 (head), 27 (scutellum); Thompson et al. 1976:71 (cat.). Material examined. COLOMBIA: Boyacá: Villa de Leyva, 5.638191°, -73.535870°, 2200m 18.xi.1996, Leg. M. Galvis (ICN). brunnigaster (Hull), 1943k:41 (Volucella). Type locality: Colombia, Meta. HT M (UZMC). Distr.: Colombia (Boyacá, Cundinamarca, Huila, Magdalena, Meta, Risaralda (Pereira), Quindio (Filandia)). Refs.: Thompson et al. 1976:72 (cat.); Restrepo & Carrejo, 2009:146 (cat.). Material examined. COLOMBIA: Risaralda: Pereira, Wildlife Sanctuary Otún Quimbaya, 4.728880°, 75.577850°, 1890m, 1female, Leg. Entomology Group, Vsr. Fish (CEUA); Quindio: Filandia, Granja Experimental Bengala, Forest, 4.685291°, -75.617533°, 2000m, 13–16.xi.2009, Leg. Entomology Group, Vsr. fish (CEUA). caesariatum (Williston), 1891:49 (Volucella). Type locality: Mexico, Orizaba. HT F (BMNH). Distr.: Colombia (Magdalena, Putumayo), Mexico. Refs.: Thompson et al. 1976:73 (cat.); Restrepo & Carrejo, 2009:146 (cat.). Material examined. COLOMBIA: Antioquia: Copacabana, Vereda El Cabuyal, 6.345579°, -75.503714°, 1650m, 1male, 1female, Leg. Grisales & Rivera, Manual (CEUA). chalybescens (Wiedemann), 1830:204 (Volucella). Type locality: "Brazil". HT M (NMW). Distr.: Colombia (Amazonas, Antioquia (Alejandria, Apartado, El Bagre, La Pintada, San Luis, San Roque), Quindío (Quimbaya), Bolivar, Boyacá, Cundinamarca, Meta, Tolima, Putumayo, Quindío, Vaupés, Vichada), Brazil, Cuba, Panama, Trinidad. Refs.: Curran, 1926a:59 (desc.); Thompson et al. 1976:73 (cat.); Restrepo & Carrejo, 2009:146 (cat.). Material examined. COLOMBIA: Antioquia: Alejandria, Vereda San Jose, 6.371942°, -75.109771°, 1879m, 1female, Leg. A.L. Montoya, Vsr. Fish (CEUA); Apartado, Via Carepa, Parque de Los Encuentros, 7.759545°, -76.654668°, 30m, Rastrojo, 27-.xi.2004, Leg. Velez & Grisales, Vsr. Fish (CEUA); El Bagre, Vereda 505 Parcelas, interior de Bosque, 7.594170°, -74.811900°, 150m, 2males, Leg. L. Perez & M. Quiroz, Net (CEUA); La Pintada, 5.736521°, -75.632087°, 650m, 1female, Leg. L.A. Vergara, Vsr. Viscera (CEUA); San Luis, Cañon de Rio Claro, Forest, 5.894165°, -74.856963°, 350m, 1male, 1female, 26.xi.2009, Leg. I. Ceballos, Vsr. Fish (CEUA); San Roque, 6.503175°, -74.926885°, 1800m, 1male, Leg. N. Uribe, Vsr. Viscera (CEUA); Quindio: Quimbaya, Vereda Kerman, Km 7, 4.614947°, -75.807598°, 1339m, 1male, Leg. H. Vargas, Vsr. Fish (CEUA). cinctiventre (Curran), 1930d:16 (Volucella). Type locality: Panama, Canal Zone, Barro Colorado Island. HT F (AMNH). Distr.: Colombia (Antioquia (Guarne, San Roque), Amazonas (Caño Lorena), Huila), Caldas (Manizales)), Panama, Trinidad. Refs.: Thompson et al. 1976:73, 82 (cat.); Restrepo & Carrejo, 2009:146, 148 (cat.); Ricarte et al.2015:42 (desc., syn.). Material examined. COLOMBIA: Antioquia: Gurane, Vereda Piedras Blancas, 6.291588°, -75.493345°, 2200m, 1female, Leg. A. Velez, Vsr. Fish (CEUA); San Roque, Corregimiento San Jose del Nus, 6.503175°, 74.926885°, 1800m, 1male, 2females, Leg. G. Enotomoloy, Vsr. Fish (CEUA); Amazonas: Caño Lorena, 3.712698°, -70.333443°, 150m, Leg. F. Fernandez (ICN); Caldas: Manizales, Vereda las Palomas, Reserva Rio Blanco, 5.111962°, -75.462904°, 2550m, 1male, Leg. A. Velez, Net (CEUA). circumdatum (Walker), 1857:154 (Temnocera). Type locality. Brazil (as "Valley of Amazon"). HT F (BMNH). Distr.: Colombia (Antioquia (San Roque), Meta, Putumayo, Risaralda, Norte de Santander, Vichada), Brazil, Costa Rica. Refs.: Fluke, 1957a:156 (species incertae sedis); Kertesz, 1910:195 (citation); Sack, 1921:137 (Bolivia, biology, immatures, fig. 10 (puparium), fig. 1c (anterior spiracular process); Curran, 1926:52 (key ref.) 1830:7 (key ref.) 1834:380 (key ref.) 1839:2 (key ref); Fluke, 1957a:76 (cat.); Thompson et al. 1976:73 (cat.); Thompson & Marinoni 2003:217–223 (redesc. and notes) figs. 1 (adult, lateral view), 2 (head, lateral view), 3 (thorax, dorsal), 4 (abdomen), 5 (M genitalia complex); Restrepo & Carrejo, 2009:146 (cat.). Material examined. COLOMBIA: Antioquia: San Roque, 6.503175°, -74.926885°, 1800m, 1male, Leg. N. Uribe, Vsr. Viscera (CEUA). Det F.C. Thompson, 2009. 508 · Zootaxa 4122 (1) © 2016 Magnolia Press MONTOYA circumscriptum (Curran), 1939b:13 (Volucella). Type locality: Brazil, Sao Paulo. HT M/F (AMNH). Distr.: Colombia (Vaupés), Brazil. Refs.: Thompson et al. 1976:73 (cat.); Restrepo & Carrejo, 2009:146 (cat.). contumax (Curran), 1939b:4 (Volucella). Type locality: "Brazil". HT M (AMNH). Distr.: Colombia (Nariño, Risaralda), Brazil. Refs.: Thompson et al. 1976:74 (cat.); Restrepo & Carrejo, 2009:146 (cat.). correctum (Curran), 1927c:87 (Volucella). Type locality: Costa Rica, San Jose. HT M (UK). Distr.: Colombia (Antioquia (Caucasia, La Pintada, Maceo, San Luis, San Roque), Caldas, Meta, Quindio (Quimbaya), Santander (Cimitarra)), Costa Rica, Panama, Tinidad. Refs.: Thompson et al. 1976:74 (cat.); Restrepo & Carrejo, 2009:146 (cat.). Material examined. COLOMBIA: Antioquia: Caucasia, Barrio El Camello, 7.983333°, -75.200000°, 50m, 1male, iv.2010, Leg. Entomology Group (CEUA); La Pintada, Hacienda Montenegro, 5.736521°, -75.632087°, 650m, 1female, Leg. N. Uribe, Vsr. Fish (CEUA); Maceo, Reserva Cañón del Rio Alicante, 6.540547°, 74.770327°, 840m, 1female, Leg. M. Wolff, Vsr. Fish (CEUA); San Luis, Reserva Natural Cañon del Rio Claro, 5.894165°, -74.856963°, 350m, 1female, Leg. A. Velez, Vsr. Fish (CEUA); San Roque, Corregimiento San Jose del Nus, 6.503175°, -74.926885°, 1800m, 1male, 1.v.2007, Leg. Entomology Group, Vsr. Viscera (CEUA); Quindio: Quimbaya, Vereda Kerman, Km 7, 4.614947°, -75.807598°, 1339m, 9males, 3females, Leg. H. Vargas, Vsr. Fish (CEUA); Santander: Cimitarra, Corregimiento Puerto Olaya, 6.473122°, -74.395383°, 110m, 2males, 1male, 14.vi.2005, Leg. M. Castañ & C. Rivera, Black Light trap (CEUA). corumbense (Curran), 1930d:18 (Volucella). Type locality: Brazil, Mato Grosso, Corumba. HT F (AMNH). Distr.: Colombia (Antioquia (Medellín), Bolivar, Cauca, Magdalena, Norte de Santander), Brazil. Refs.: Thompson et al. 1976:74 (cat.); Restrepo & Carrejo, 2009:146 (cat.). Material examined. COLOMBIA: Antioquia: Medellín, Vereda Piedras Blancas, 6.291588°, -75.493345°, 2350m, 22.vii.2011, Leg. Entomology Group (MEPB). delila (Hull), 1950c:230 (Volucella). Type locality: Peru, Chanchamayo. HT M (CNC). Distr.: Colombia (Amazonas, Huila), Peru. Refs.: Thompson et al. 1976:74 (cat.); Restrepo & Carrejo, 2009:148 (cat.). dispar (Macquart), 1846:251 (Volucella). Type locality: "New Grenada". HT M/F (UMO). Dist. Colombia (Antioquia), Guatemala, Panama, Venezuela. Refs.: Macquart, 1846:251 (123), pi. 11, figs. 2 (wing), 2a (head); Thompson et al. 1976:75 (cat.). florella (Hull), 1944f:49 (Volucella). Type locality: Guyana, Potaro River, Tumatumari. HT F (CUI). Distr.: Colombia (Caldas (Norcacia), Huila), Guyana. Refs.: Thompson et al. 1976:76 (cat.); Restrepo & Carrejo, 2009:147 (cat.). Material examined. COLOMBIA: Caldas: Norcacia, 5.560100°, -74.893349°, 700m, 1female, Leg. A.L. Montoya, Net (CEUA). flukei (Curran), 1936b: 10 (Volucella). Type locality: Brazil, Parana. HT F (AMNH). Distr.: Brazil, Colombia (Antioquia (Alejandria, Ciudad Bolivar, Copacabana), Risaralda (Pereira)). Thompson et al. 1976:75 (cat.). Material examined. COLOMBIA: Antioquia: Alejandria, 6.401714°, -75.058838°, 1200m, 1female, 8.xi.07, Leg. A.L. Montoya, Vsr. Fish (CEUA); Ciudad Bolivar, Carretera La Sucia, 1200m, 22.v.2011, Leg. E. Corrrea (CEUA); Copacabana, Vereda El Cabuyal, 6.345579°, -75.503714°, 1450m, 1female, Leg. Grisales & Rivera, Manual (CEUA); Risaralda: Pereira, Wildlife Sanctuary Otún Quimbaya, 4.728880°, -75.577850°, 1890m, 3males, 3females, v.2001, Leg. H. Vargas, Vsr. Fish (CEUA). fornax (Townsend), 1895a:613 (Volucella). Type locality: Mexico, Baja California, El Taste. HT M (CAS). Distr. Distr.: Colombia (Cundinamarca), Mexico. Refs.: Thompson et al. 1976:76 (cat.); Restrepo & Carrejo, 2009:147 (cat.). CATALOGUE OF SYRPHIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 509 fraudulentum (Williston), 1891:48 (Volucella). Type locality: Mexico, Guerrero, Venta de Zopilote; North Yucatan, Temax. HT M/F (BMNH). Distr.: Colombia (Antioquia (Carepa, Maceo), Huila), Mexico. Refs.: Williston, 1891:48, pi. 1:13 (habitus), 13a (head), 13b (antenna); Thompson et al. 1976:76 (cat.); Restrepo & Carrejo, 2009:147 (cat.). Material examined. COLOMBIA: Antioquia: Carepa, Hacienda Tulenapa Universidad de Antioquia, 7.759545°, -76.654668°, 47m, 8.ii.2009, Leg. Entomology Group (CEUA); Maceo, Reserva Cañon del rio Alicante, Native forest, 6.540547°, -74.770327°, 840m, 1male, 12.v.2000, Leg. S. Perez, Net (CEUA). frauenfeldi (Schiner), 1868:358 (Temnocera). Type locality: "Chile". HT M/F (NMW). Distr.: Colombia (Antioquia (Puerto Berrio, Rionegro, San Luis, San Roque), Amazonas, Caldas (Norcacia), Caqueta, Meta, Nariño, Putumayo, Vaupés), Argentina, Chile. Refs.: Shannon & Aubertin, 1933:170, fig. 35a (head); Fluke, 1915b:19 (desc.), fig. 32 (scutellum); Thompson et al. 1976:76 (cat.); Restrepo & Carrejo, 2009:147 (cat.). Material examined. COLOMBIA: Antioquia: Puerto Berrio, Vereda Cristalina, RUNA, 6.489819°, 74.401992°, 114m, 1male, 25-ix.2006, Leg. Entomology Group, Net (CEUA); Rionegro, Corregimiento de San Antonio de Pereira, Parque las Veraneras, 6.134884°, -75.393827°, 2100m, 1male, Leg. C. Perez, Net (CEUA); San Luis, Rio Claro, 5.894165°, -74.856963°, 350m, 1male, 6.ix.2007, Leg. Entomology Group, Vsr. Fish (CEUA); San Roque, Corregimiento San Jose del Nus, 6.503175°, -74.926885°, 1800m, 3males, 1.v.2007, Leg. Entomology Group, Vsr. Fish (CEUA); Caldas: Norcacia, Sector El Tigre, Forest, 5.560100°, -74.893349°, 700m, 1male, 22.ii.2010, Leg. L. Rios & D. Gaurisas, Vsr. Fish (CEUA). fulvicorne (Bigot), 1883:84 (Volucella). Type locality: "Panama". HT M/F (UMO). Distr.: Colombia (Antioquia (San Roque)), Costa Rica, Panama. Refs.: Thompson et al. 1976:76 (cat.). Material examined. COLOMBIA: Antioquia: San Roque, Vereda El Iris, 6.503175°, -74.926885°, 1800m, 1female, 10.iv.2014, Entomology Group, Pitfall copro (CEUA). hambletoni (Fluke), 1951b:16 (Volucella). Type locality: Peru, Cariete. HT M (AMNH). Distr.: Colombia (Antioquia (Anori, Maceo, San Luis), Caldas (Anserma), Magdalena Santander (Betulia)), Ecuador, Peru. Refs.: Fluke, 1951b:16, figs. 16 (head), 41 (antenna), 57 (M genitalia); Thompson et al. 1976:77 (cat.); Restrepo & Carrejo, 2009:147 (cat.). Material examined. COLOMBIA: Antioquia: Anori, Vereda El Roble, Reserva Natural Arrierito Antioqueño, 6.984806°, -75.111667°, 1600–1700m, 1female, 25.v.2012, Leg. Entomology Group, Vsr. Fish (CEUA); Maceo, Reserva Cañon del rio Alicante, Native forest, 6.540547°, -74.770327°, 840m, 1male, 23.i.2009, Leg. M. Wolff, Vsr. Fish (CEUA); San Luis, Rio Claro, 5.894165°, -74.856963°, 350m, 1male, 21.xi.2009, Leg. Entomology Group, Vsr. Fish (CEUA); Caldas: Anserma, Vereda Palo Blanco, 5.231550°, -75.776772°, 1800m, 2males, Leg. A.L. Montoya, Manual on spider net (CEUA); Santander: Betulia, Vereda Palma de Oro, Hacienda La Antigua, 6.890176°, -73.271165°, 1700m, x.2001, Leg. A. Velez & P. Duque, Vsr. Fish (CEUA). hydrofenestra (Hull), 1943n:513 (Volucella). Type locality: Colombia, Muzo. HT F (CNC). Distr.: Colombia (Boyacá, Cundinamarca). Refs.: Thompson et al. 1976:77 (cat.). imitans (Curran), 1926a:63 (Volucella). Type locality: Mexico. HT M (AMNH). Distr.: Colombia (Antioquia (Santafe de Antioquia), Magdalena (Santa Marta)), Costa Rica, Mexico. Refs.: Thompson et al. 1976:77 (cat.); Restrepo & Carrejo, 2009:147 (cat.). Material examined. COLOMBIA: Antioquia: Santafé de Antioquia, 6.512784°, -75.825832°, 460m, 1female, Leg. A.L. Montoya, Net (CEUA); Magdalena: Santa Marta, PNN Tayrona Neguanje, 11.276050°, 74.195519°, 121m, Leg. F. Escobar (ICN). inconsistens (Curran), 1939b:3 (Volucella). Type locality: Brazil, Mt. Roraima. HT F (AMNH). Distr.: Colombia (Boyacá), Brazil, Costa Rica. Refs.: Thompson et al. 1976:77 (cat.); Restrepo & Carrejo, 2009:147 (cat.). infractum Thompson, 1981:130. Type locality: Dominica, Freshwater Lake, 2500 ft. HT M (USNM). Distr.: Colombia (Amazonas, Antioquia (Amalfi, Anori, San Luis), Chocó, Caldas, Huila, Meta, Nariño, Risaralda 510 · Zootaxa 4122 (1) © 2016 Magnolia Press MONTOYA (Pereira), Vaupés, Valle del Cauca), Less Antillas. Refs.: Thompson, 1981:130 (orig. desc.), fig. 143 (face, lateral view), 160 (M genitalia); Restrepo & Carrejo, 2009:147 (cat.). Material examined. COLOMBIA: Antioquia: Amalfi, 6.916667°, -75.083331°, 1700m, Leg. Entomology Group, Vsr. Fish (CEUA); Anori, Vereda El Roble, Reserva Natural Arrierito Antioqueño, 6.984806°, -75.111667°, 1600–1700m, 1female, 25–27.v.2012, Leg. Entomology Group, Net (CEUA); San Luis, Rio Claro, 5.894165°, 74.856963°, 350m, 3males, Leg. Entomology Group, Vsr. Fish (CEUA); Risaralda: Pereira, Wildlife Sanctuary Otún Quimbaya, 4.728880°, -75.577850°, 1890m, 5female, Leg. Entomology Group, Vsr. Fish (CEUA). lanei (Curran), 1936b:11 (Volucella). Type locality: Brazil, Sao Paulo, Juquia. HT M (AMNH). Distr.: Colombia (Nariño, Vaupés), Brazil. Refs.: Thompson et al. 1976:77 (cat.); Restrepo & Carrejo, 2009:147 (cat.). latevitatum (Curran), 1939b:16 (Volucella). Type locality: Colombia, between Queremal and Buenaventura. HT M (AMNH). Distr.: Colombia (Valle del Cauca), Bolivia, Peru. Refs.: Thompson et al. 1976:77 (cat.). Material examined. COLOMBIA: Santander: Carmen del Chucuri, Vereda Cascajal, 6.647307°, 73.501338°, 700m, 1female, Leg. A. Vélez & P. Duque, Vsr. Fish (CEUA). longirostre (Macquart), 1846:252 (Volucella). Type locality: ¨Patria Inconnue¨. HT F (UMO). Distr.: Colombia (Boyacá), Chile. Refs.: Restrepo & Carrejo, 2009:147 (cat.). macquarti (Curran), 1926a:60 (Volucella). Type locality: Honduras, Prieta. HT F (AMNH). Distr.: Colombia (Amazonas, Caqueta (Florencia), Huila, Putumayo, Risaralda, Vaupés), Brazil, Guyana, Hispaniola, Mexico, Trinidad. Refs.: Thompson et al. 1976:78 (cat.); Restrepo & Carrejo, 2009:147 (cat.). Material examined. COLOMBIA: Caquetá: Florencia, Vereda Sebastopol, Finca La Magola, 1.598268°, 75.543651°, 305m, 25.xii.2004, Leg. Entomology Group, Net on decomposed Pig (CEUA). meretricias (Williston), 1888:272 (Volucella). Type locality: Brazil, Mato Grosso, Chapada; Rio de Janeiro. HT M/F (AMNH). Distr.: Colombia (Antioquia (Puerto Berrio, La Pintada, San Luis, San Roque)), Brazil. Refs.: Thompson et al. 1976:79 (cat.). Material examined. COLOMBIA: Antioquia: Puerto Berrio, Alto de las Aguilas, Hacienda Manaos, 6.500865°, -74.546317°, 404m, 1female, 5.vii.2013, Leg. Entomology Group, Net (CEUA); La Pintada, Hacienda Montenegro, 5.736521°, -75.632087°, 650m, 1male, Leg. N. Uribe, Vsr. Fish (CEUA); San Luis, Rio Claro, 5.894165°, -74.856963°, 350m, 3males, Leg. Entomology Group, Vsr. Fish (CEUA); San Roque, Corregimiento San Jose del Nus, 6.503175°, -74.926885°, 1800m, 1male, 1female, Leg. N. Uribe, Vsr. liver (CEUA). mocanum (Curran), 1936b:11 (Volucella). Type locality: Guatemala, Moca Guatalon. HT F (AMNH). Distr.: Colombia (Cundinamarca, Huila, Risaralda), Guyana, Peru, Venezuela. Refs.: Thompson et al. 1976:79 (cat.); Restrepo & Carrejo, 2009:148 (cat.). muscarium (Thomson), 1869:493 (Hanrotricha). Type locality: Argentina, Buenos Aires. HT M (AMNH). Distr.: Colombia, Argentina. Refs.: Thompson et al. 1976:79 (cat.). nigripes (Bigot), 1857:296 (Phalacromyia). Type locality: "Chile". HT M (UMO). Distr.: Colombia (Huila (San Agustín)). Refs.: Bigot, 1857a:296, pi. 7:9 (head); Thompson et al. 1976:80 (cat.). Material examined. COLOMBIA: Huila: San Agustín, 1.927474°, -76.297857°, 1700m, 1male, 22.vii.2008, Leg. A.L. Montoya, Manual on spider net (CEUA). obscurior (Curran), 1939b:5 (Volucella). Type locality: Brazil, Santa Catarina, Nova Teutonia. HT M (AMNH). Distr.: Colombia (Antioquia (Macea, San Luis, San Roque), Chocó (Acandí), Quinidio (Filandia), Risaralda (Pereira, Santuario)), Argentina, Brazil. Refs.: Thompson et al. 1976:80 (cat.). Material examined. COLOMBIA: Antioquia: Maceo, Vereda San Pedro, finca San Pedro, 6.540547°, 74.770327°, 840m, Forest, 1male, 17.ii.2012, Leg. Entomology Group, Vsr. Fish (CEAU); San Luis, Rio Claro, 5.894165°, -74.856963°, 350m, 1female, Leg. Entomology Group, Vsr. Fruit (CEUA); San Roque, 6.503175°, - CATALOGUE OF SYRPHIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 511 74.926885°, 1800m, 1female, Leg. N. Uribe, Vsr. Viscera (CEUA); Chocó: Acandí, Punta Napu, Forest near the Sea, 8.417380°, -77.148863°, 0-50m, i.2010; Leg. Bustamante, Net (CEUA); Quindio: Filandia, Hacienda Bengala, 4.685291°, -75.617533°, 2000m, 1female, 10.ix.2008, Leg. A.L. Montoya (CEUA); Risaralda: Pereira, Wildlife Sanctuary Otún Quimbaya, 4.728880°, -75.577850°, 1890m, 4males, 1female, Leg. Entomology Group, Vsr. Fish (CEUA); Santuario, Vereda La Esmeralda, Natural National Park Tatama, 5.105551°, -76.008496°, 2000m, 18-30.x.2013, Leg. C. Bota & C. Flórez, Net (CEUA). opinator (Williston), 1891:51 (Volucella). Type locality: Mexico, Vera Cruz, Atoyac. HT F (BMNH). Distr.: Colombia (Magdalena, Risaralda), Costa Rica, Mexico. Refs.: Williston, 1891:51, pi. 1:14 (habitus), 14a (head), 14b (antenna); Thompson et al. 1976:80 (cat.); Restrepo & Carrejo, 2009:148 (cat.). pallens (Wiedemann), 1830:204 (Volucella). Type locality: "Brazil". HT F (NMW). Distr.: Colombia (Antioquia (Caucasia, La Pintada, Puerto Berrio), Arauca (Fortul), Bolivar (Mompox), Cundinamarca (La Mesa), Sucre (San Onofre), Tolima (Espinal)), Argentina, Brazil, Puerto Rico. Refs.: Thompson et al. 1976:81 (cat.). Material examined. COLOMBIA: Antioquia: Caucasia, 7.983333°, -75.200000°, 80m, Leg. Entomology Group, Light trap (CEUA); La Pintada, Hacienda Montenegro, 5.736521°, -75.632087°, 650m, 1male, Leg. N. Uribe, Vsr. Fish (CEUA); Puerto Berrio, 6.489819°, -74.401992°, 114m, 1male, 1female, 25.xi.2004, Leg. Vélez & Grisales (CEUA); Arauca: Fortul, 6.792220°, -71.775600°, 250m, 23.iv.2002, Leg. N. Ulloa (UNAB); Bolivar: Mompox, La Sebastiana, 9.163064°, -74.546328°, 10m, 1female, 27.ii.2000, Leg. M. Wolff, Net (CEUA); Cundinamarca: La Mesa, Zona urbana, 4.629469°, -74.466633°, 1298m, 12.ix.2003, Leg. A. Forero, Net (UNAB); Sucre: San Onofre, 9.646080°, -75.581129°, 5m, 1female, Leg. A.L. Montoya, Net (CEUA); Tolima: Espinal, 4.162687°, -74.874972°, 322m 18.v.1995, Leg. A. Perilla (UNAB). Det F.C. Thompson, 2009. parina (Fluke), 1951b:25 (Volucella). Type locality: Chile, Arica, Parinacota. HT M (UChS). Distr.: Colombia (Antioquia, Chocó, Magdalena). Refs.: Fluke, 1951b:25, figs. 14 (head), 61 (M genitalia); Thompson et al. 1976:81 (cat.); Restrepo & Carrejo, 2009:148 (cat.). persimile (Williston), 1888:273 (Volucella). Type locality: Brazil, Destares & Mato Grosso, Chapada; Rio de Janeiro. HT M/F (AMNH). Distr.: Colombia (Antioquia (Alejandria, San Luis, San Roque), Caldas (Irra), Caqueta)), Brazil. Refs.: Thompson et al. 1976:81 (cat.). Material examined. COLOMBIA: Antioquia: Alejandria, Vereda San Jose, 6.371942°, -75.109771°, 1879m, 1female, Leg. A.L. Montoya, Vsr. Fish (CEUA); San Luis, 6.042529°, -74.993289°, 1160m, 1female, Leg. A. Vélez, Vsr. Fish (CEUA); San Roque, 6.503175°, -74.926885°, 1800m, 1female, Leg. N. Uribe, Vsr. Viscera (CEUA); Caldas: Irra, 5.271445°, -75.665865°, 745m, 1male, Leg. J.D. Suaza, Manual (CEUA). pica (Schiner), 1868:355 (Phalocromyia). Type locality: Colombia. HT M/F (NMW). Distr.: Colombia (Antioquia (Andes, Medellín)), Brazil, Costa Rica, Mexico, Venezuela. Refs.: Thompson et al. 1976:81 (cat.). Material examined. COLOMBIA: Antioquia: Andes, Vereda La Mesenia, 5.604827°, -75.954463°, 2100m, 1male, Leg. L. Gomez, 13.ii.11, Net (CEUA); Medellín, Vereda Piedras Blancas, 6.291588°, -75.493345°, 2200m, Leg. G. Entomoloy (MEPB). pictum (Wiedemann), 1830:201 (Volucella). Type locality: "Brazil". HT M/F (NMW). Distr.: Colombia (Amazonas, Putumayo, Vaupés), Argentina, Brazil, Costa Rica, Cuba, Mexico, Panama. Refs.: Curran, 1925a:206, pi. 10:145 (M genitalia); Doesburg, 1966:96, fig. 66 (habitus); Thompson et al. 1976:82 (cat.); Restrepo & Carrejo, 2009:148 (cat.). posticum (Say), 1829:166 (Volucella). Type locality: "Mexico". Distr.: U.S.A. (California to Colorado), south to Mexico. HT M (ANSP). Distr.: Colombia (Antioquia (Itagui, Fredonia, Medellín, Santa fe de Antioquia), Cundinamarca (Fusa)), Mexico. Thompson et al. 1976:82 (cat.). Material examined. COLOMBIA: Antioquia: Itagüí, Vereda La María, 6.189489°, -75.627675°, 2546m, 1female, Leg. L. Gómez, Net (CEUA); Fredonia, Vereda Jonas, 5.916573°, -75.683347°, 1010m, Leg. E. Flórez, Net (CEUA); Medellín, 6.267404°, -75.585789°, 1570m, 1445, 1female, Leg. N. Herrera, Net (CEUA); Santafé de 512 · Zootaxa 4122 (1) © 2016 Magnolia Press MONTOYA Antioquia, 6.512784°, -75.825832°, 500m, 1male, Leg. A.L. Montoya, Net (CEUA); Cundinamarca: Fusagasuga, 4.338328°, -74.363968°, 1771m, 15.xi.1969, Leg. O. Torres (ICN); Tena, 4.624207°, -74.239034°, 1340m, 27.v.1991, Leg. A. Castillo (ICN). prasinus (Schiner), 1868:354 (Phalacromyia). Type locality: "South America". HT M/F (NMW). Distr.: Colombia (Boyacá, Cundinamarca, Magdalena, Risaralda), South America. Refs.: Thompson et al. 1976:82 (cat.); Restrepo & Carrejo, 2009:148 (cat.). pseudopallens Thompson, 1981:131. Type locality: Jamaica, Perkins. HT M (CMP). Distr.: Colombia (Amazonas, Putumayo, Vaupés), Jamaica. Refs.: Thompson, 1981:131 (orig. desc.), fig 153, 156 (M genitalia complex), pi 133 (key); Restrepo & Carrejo, 2009:148 (cat.). punctiferum (Bigot), 1875:475 (Volucella). Type–locality: "Amazonia". HT M (BMNH). Distr.: Colombia (Amazonas, Bolívar (Zambrano), Risaralda), Guatemala. Refs.: Thompson et al. 1976:82 (cat.). punctigena (Hull), 1937a:165 (Volucella). Type locality: Guatemala, Siquinola. HT F (USNM). Distr.: Colombia (Bolívar (Zambrano), Boyacá), Mexico. Refs.: Hull, 1937a:165, fig. 6 (abdomen); Thompson et al. 1976:82 (cat.); Restrepo & Carrejo, 2009:148 (cat.). Material examined. COLOMBIA: Bolívar: Zambrano, Hacienda Monterrey, 9.749954°, -74.833324°, 30m, 1.viii.1996, Leg. F. Escobar (ICN). purpuriferum (Bigot), 1875:475 (Volucella). Type–locality: "Amazonia". HT M (BMNH). Distr.: Colombia (Risaralda), Mexico. Refs.: Thompson et al. 1976:82 (cat.); Restrepo & Carrejo, 2009:148 (cat.). roraima (Curran), 1939b:3 (Volucella). Type locality: Brazil, Mt. Roraima. HT M (AMNH). Distr.: Colombia (Huila), Brazil. Refs.: Thompson et al. 1976:83 (cat.); Restrepo & Carrejo, 2009:148 (cat.). selectum (Curran), 1939b:15 (Volucella). Type locality: Brazil, Santa Catarina, Nova Teutonia. HT M (AMNH). Distr.: Colombia (Antioquia (Alejandria, Santafe de Antioquia), Risaralda (Pereira)), Brazil. Refs.: Thompson et al. 1976:83 (cat.). Material examined. COLOMBIA: Antioquia: Alejandria, Vereda El Carbón, 6.403715°, -75.064663°, 1800m, 2males, Leg. A.L. Montoya, rearing in laboratory (CEUA); Santafé de Antioquia, 6.512784°, -75.825832°, 460m, 1female, Leg. A.L. Montoya, Net (CEUA); Risaralda: Pereira, Wildlife Sanctuary Otún Quimbaya, 4.728880°, -75.577850°, 1890m, 4males, 1female, Leg. Entomology Group, Vsr. Fish (CEUA). sexmaculatum (Palisot de Beauvois), 1819: :224 (Syrphus). HT M/F (Lost). Type locality: Dominican Republic, Santo Domingo. Dist. Colombia (Antioquia (Carepa, Girardota)), Southern U.S.A. Refs.: Palisot de Beauvois, 1819: :224, pi. 3:8 (habitus). Material examined. COLOMBIA: Antioquia: Carepa, Hacienda Tulenapa Universidad de Antioquia, 7.759545°, -76.654668°, 48m, Vii.2013, Leg. Entomology Group (CEUA); Girardota, 6.343502° -75.435102°, 2200m, xii.1981, Leg. E. Arcila (MEFLG). spinithorax (Lynch Arribalzaga), 1892b:190 (1892c:141) (Temnocera). Type locality: Argentina, Chaco, Colonia Azara. HT F (MACN). Distr.: Colombia (Caqueta, Putumayo, Vaupés, Vichada), Argentina, Brazil, Panama. Refs.: Thompson et al. 1976:84 (cat.); Restrepo & Carrejo, 2009:148 (cat.). Material examined. COLOMBIA: Caldas: Irra, 5.271445°, -75.665865°, 745m, 1female, Leg. J.D. Suaza (CEUA). subcoeruleum (Rondani), 1863:3 (Phalacromyla). Type locality: "America meridionalis" = (Colombia?). HT M (IZUSN). Distr.: Colombia. Refs.: Thompson et al. 1976:84 (cat.). CATALOGUE OF SYRPHIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 513 tumicephalum (Hull), 1943c:35 (Volucella). Type locality: "Peru". HT M (BMNH). Distr.: Colombia (Antioquia (Alejandria, San Roque), Caldas (Irra), Magdalena (Santa Marta)), Peru. Refs.: Thompson et al. 1976:85 (cat.). Material examined. COLOMBIA: Antioquia: Alejandria, Vereda San Jose, 6.371942°, -75.109771°, 1879m, 1female, Leg. A.L. Montoya, Vsr. Fish (CEUA); San Roque, San Jose del Nus, 6.503175°, -74.926885°, 1800m, 5.i.2007, Leg. N. Uribe (CEUA); Caldas: Irra, 5.271445°, -75.665865°, 745m, 1male, Leg. J.D. Suaza, reared from Bromeliad (CEUA); Magdalena: Santa Marta, PNN Tayrona, 11.276050°, -74.195519°, 121m, 8.vi.1974, Leg. A. Ibagos (ICN). tympanitis (Fabricius), 1805:226 (Syrphus). Type locality: "America meridional". HT M/F (UZMC). Distr.: Colombia (Amazonas, Antioquia (Carmen de Viboral, Medellín, Puerto Berrio, San Luis), Cauca, Chocó (Riosucio), Meta (Villavicencio), Santander (Cimitarra), Sucre (San Marcos), Valle del Cauca (Santiago de Cali)), Costa Rica, Guatemala, Mexico, Panama. Refs.: Thompson et al. 1976:85 (cat.); Restrepo & Carrejo, 2009:148 (cat.). Material examined. COLOMBIA: Antioquia: Carmen de Viboral, Vereda El Carmen, 2100m, Leg. M. Zuluaga, Manual (CEUA); Medellín, 6.267404°, -75.585789°, 1570m, 1male, Leg. N. Herrera, Net (CEUA); Puerto Berrio, Alto de las Aguilas, Hacienda Manaos, 6.500865°, -74.546317°, 404m, 1male, 5.vii.2013, Leg. Entomology Group, Light trap (CEUA); San Luis, Vereda La Palma, Bosque fragmentado, 6.037362°, 75.004162°, 1000m, 13.viii.2013, Leg. Entomology Group, Net (CEUA); Chocó: Riosucio, Tilupo, 7.440560°, 77.118900°, 7m, 10.iv.1978, Leg. H. Echeverri (ICN); Meta: Villavicencio, Forest Bavaria, 4.150000°, 73.633333°, 467m, 9.iv.2005, Leg. Students of Animal Systematic (ICN); Santander: Cimitarra, Corregimiento Puerto Olaya, 6.473122°, -74.395383°, 110m, 2males, 1 female, 14.vi.2005, Leg. M. Castañ & C. Rivera (CEUA); Sucre: San Marcos, 8.660492°, -75.134688°, 27m, 25.vi.2003, Leg. J. Martínez, Net (UNAB); Valle del Cauca: Santiago de Cali, Farallones de Cali, 3.367176°, -76.687470°, 3400m, 5.v.1993, Leg. C. Andrade (ICN). ulrica (Hull), 1950a:68 (Volucella). Type locality: Peru, Chanchamayo. HT F (CNC). Distr.: Colombia (Amazonas, Antioquia, Huila, Meta, Risaralda (Pereira), Vaupés, Valle del Cauca), Peru. Refs.: Thompson et al. 1976:86 (cat.); Restrepo & Carrejo, 2009:148 (cat.). Material examined. COLOMBIA: Risaralda: Pereira, Wildlife Sanctuary Otún Quimbaya, 4.728880°, 75.577850°, 1890m, 1male, Leg. Entomology Group, Net (CEUA). vagum (Wiedemann), 1830:205 (Volucella). Type locality: "Brazil". HT M (NMW). Distr.: Colombia (Amazonas (Caño Lorena), Antioquia (Amalfi, Alejandria, Bello, Caicedo, Copacabana, Gomez Plata, Jericó, La Pintada, Maceo, Medellín, San Jeronimo, Puerto Berrio, San Luis, San Roque, Tamesis), Atlántico (Luruaco), Boyacá (Pauna), Caldas, Chocó, Caqueta, Cundinamarca (La Mesa, La Palma, Mesitas del Colegio, Puerto Salgar, Topaipi, Villeta, Viotá), Huila, Meta (Bella Vista, Villavicencio), Putumayo, Risaralda (Pereira), Santander, Sucre, Tolima (Espinal, Ibagué)) Vaupés, Valle del Cauca), Brazil, Costa Rica, Guyana, Less Antillas, Peru. Refs.: Thompson et al. 1976:86 (cat.); Restrepo & Carrejo, 2009:149 (cat.). Material examined. COLOMBIA: Amazonas: Caño Lorena, -3.712698°, -70.333443°, 150m, 1.viii.1994, Leg. F. Fernandez (ICN); Antioquia: Amalfi, 1050m, 1male, Leg. P. Duque, Vsr. Fish (CEUA); Alejandria, Vereda San Jose, 6.371942°, -75.109771°, 1879m, 1female, Leg. A.L. Montoya, Vsr. Fruit (CEUA); Bello, 6.340400°, 75.562271°, 1500m, 1female, Leg. J. Cardona, Net (CEUA); Caicedo, 6.403933°, -75.975283°, 2200m, 1female, Leg. L. Urrea, Net (CEUA); Copacabana, Vereda El Cabuyal, 6.345579°, -75.503714°, 1450m, 1female, Leg. Grisales & Rivera, Manual (CEUA); Gómez Plata, Vereda La Clara, 6.683154°, -75.216575°, 1800m, 1female, Leg. Entomology Group, Net (CEUA); Jericó, Cauca Viejo, 5.753990°, -75.809029°, 400m, 1male, Leg. Entomology Group, Vsr. (CEUA); La Pintada, Hacienda Montenegro, 5.736521°, -75.632087°, 650m, 1female, Leg. N. Uribe, Vsr. Onion (CEUA); Maceo, 6.540547°, -74.770327°, 840m, 1female, Leg. M. Wolff, Vsr. Fish (CEUA); Medellín, 6.267404°, -75.585789°, 1570m, 1female, Leg. A.L. Montoya, Manual (CEUA); San Jeronimo, 6.432764°, -75.714291°, 1225m, Leg. A. Vélez, Vsr. Fish (CEUA); Puerto Berrio, 6.489819°, 74.401992°, 114m, 1female, Leg. Bustamante, Net (CEUA); Támesis, 5.666667°, -75.716667°, 500–800m, 1female, Leg. J. Posada, Light trap (CEUA); San Luis, Rio Claro, 5.894165°, -74.856963°, 350m, 1male, Leg. Entomology Group, Vsr. Fruit (CEUA); San Roque, Corregimiento San Jose del Nus, 6.503175°, -74.926885°, 1800m, 1male, 1female, Leg. N. Uribe, Vsr. Liver (CEUA); Atlántico: Luruaco, Vitelma, 10.647233°, - 514 · Zootaxa 4122 (1) © 2016 Magnolia Press MONTOYA 75.082286°, 30m, 09.v.1969, Leg. C. Bernal (UNAB); Boyacá: Pauna, Manote bajo La Rochela, 5.666667°, 73.983333°, 1280m, 25.iv.1979 (ICN); Cundinamarca: La Mesa, El Palmar, 4.629469°, -74.466633°, 1298m, 01.iv.2004, Leg. M. Calderón (UNAB); Granada, 4.518974°, -74.351273°, 450m, 23.xi.2003, Leg. R. Parra (UNAB); La Palma, 5.332998°, -74.383346°, 1462m, 11.xi.1995, Leg. F. Rodríguez (UNAB); Mesitas del Colegio, Vereda Santa Marta, Finca Los Guayabos, 4.567024°, -74.433012°, 1600m, 08.ix.2001, Leg. J. Velasco, Net (UNAB); Puerto Salgar, Vereda Los Campos, 5.499381°, -74.583475°, 828m 18.iv.2004, Leg. H. Gamba. E. Rueda, Net (UNAB); Topaipi, 5.372367°, -74.366351°, 1323m 18.iii.1996, Leg. L. Ospina, Light trap (UNAB); Villeta, 5.008319°, -74.473554°, 992m, 20.iii.1999, O. Higuera. Leg. R. Quevedo (UNAB); Viotá, Javitá, Finca La Esperanza, 4.433434°, -74.516643°, 567m, 03.xi.2002, Leg. V. Cortés, Net (UNAB); Meta: Bella Vista Carretera Villavo, 4.139059°, -73.780962°, 1120m, 1.ix.2010, Leg. L. Orozco (ICN); Villavicencio, Forest Bavaria, 4.150000°, -73.633333°, 467m, 29.iv.2004, Leg. Students of Animal Systematic (ICN); Risaralda: Pereira, Wildlife Sanctuary Otún Quimbaya, 4.728880°, -75.577850°, 1890m, 4female, Leg. Entomology Group, Vsr. Fish (CEUA); Santander: Carmen del Chucuri, Vereda Cascajal, 6.647307°, -73.501338°, 690m, 1male, Leg. A. Vélez & P. Duque, Vsr. Fish (CEUA); Tolima: Espinal, 4.162687°, -74.874972°, 322m, 24.xi.2003, Leg. F. Moreno, Manual (UNAB); Ibagué, 4.440663°, -75.244141°, 1300m, 13.vi.1970, Leg. T de los Reyes (ICN). verdigaster (Hull), 1943k:41 (Volucella). Type locality: Colombia, Bogota. HT M (USNM). Distr.: Colombia (Cundinamarca). Refs.: Thompson et al. 1976:86 (cat.). vierecki (Curran), 1925b:250 (Volucella). Type locality: Colombia, Vista Nieve. HT F (AMNH). Distr.: Colombia (Amazonas, Magdalena, Cauca, Huila, Nariño). Refs.: Thompson et al. 1976:86 (cat.); Restrepo & Carrejo, 2009:149 (cat.). villarica (Fluke), 1951b:23 (Volucella). Type locality: Paraguay, Villarica. HT F (AMNH). Distr.: Colombia (Cundinamarca), Brazil, Paraguay. Refs.: Fluke, 1951b:23, figs. 12 (head), 62 (M genitalia); Thompson et al. 1976:86 (cat.); Restrepo & Carrejo, 2009:149 (cat.). virescens (Williston), 1891:42 (Phalacromyia). Type locality: Guatemala, El Tumbador. HT F (BMNH). Distr.: Colombia (Amazonas, Antioquia (Medellín, San Vicente), Boyacá (Buenavista, Sogamoso), Cauca, Caldas (Manizales), Cundinamarca (Caquezá), Huila, Magdalena, Meta, Nariño, Norte de Santander, Quindío, Risaralda (Pereira), Santander (Carmen del Chucuri), Valle del Cauca), Costa Rica, Guatemala. Refs.: Thompson et al. 1976:86 (cat.); Restrepo & Carrejo, 2009:149 (cat.). Material examined. COLOMBIA: Antioquia: Medellín, 6.267404°, -75.585789°, 1570m, Leg. M. Castaño, Manual (CEUA); San Vicente, Vereda Chaparral, 3.750000°, -75.583333°, 1354m, 1female, Leg. M. Wolff & A. Bustamante, Net (CEUA); Boyacá: Buenavista, Vereda Patiño, Las Lomas, 5.650832°, -74.286905°, 1889m, 15.iv.2004, Leg. W. Ávila, Net (UNAB); Sogamoso, El Cortijo, 5.722996°, -72.923443°, 2549m, 10.iv.2004, Leg. M. Sánchez, Net (UNAB); Caldas: Manizales, 5.072373°, -75.506634°, 2216m, 1 Female Leg. Entomology Group, Vsr. Fish (CEUA); Cundinamarca: Caquezá, Los Guayabos, 4.405452°, -73.949893°, 21.ix.2003, Leg. Rodríguez (UNAB); Risaralda: Pereira, Wildlife Sanctuary Otún Quimbaya, 4.728880°, -75.577850°, 1890m, 1female, Leg. Entomology Group, Vsr. Fish (CEUA); Santander: Carmen del Chucuri, Vereda Cascajal, 6.647307°, -73.501338°, 690m, 1female, Leg. A. Vélez & P. Duque, Vsr. Fish (CEUA). viride (Williston), 1888:275 (Volucella). Type locality: Brazil, Mato Grosso, Chapada. HT M/F (AMNH). Distr.: Colombia (Amazonas, Meta), Brazil, Costa Rica, Peru. Refs.: Thompson et al. 1976:87 (cat.); Restrepo & Carrejo, 2009:149 (cat.). wulpi (Goot), 1964:214 (Volucella; nom. nov. for testaceum Wulp). Type locality: "Venezuela, Curagao". HT M/F (UNKNOWN ML?). Distr.: Colombia (Antioquia (V. Medellín), Guajira (Uribia) Magdalena), Venezuela. Refs.: Thompson et al. 1976:87 (cat.); Restrepo & Carrejo, 2009:149 (cat.). Material examined. COLOMBIA: Antioquia: V. Medellín, 6.267404°, -75.585789°, 1570m, 1male, i.1947, Leg. F. Gallego (MEFLG); Guajira: Uribia, Santana, 12.0833330, -72.0499080, 18m, 25.i.2013, Leg. C. Valverde, Vsr. Fruit (CEUA). CATALOGUE OF SYRPHIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 515 Subgenus Viereckomyia Curran gibberum (Schiner), 1868:358 (Temnocera). Type locality: "Colombia". HT F (NMW). Distr.: Colombia (Antioquia (Caldas, Ciudad Bolivar, Guarne)), Costa Rica. Peru. Refs.: Curran, 1925b:246 (desc.males); Thompson et al. 1976:76 (cat.). Material examined. COLOMBIA: Antioquia: Caldas, 6.090000°, -75.638000°, 1560m, 1female, Leg. Norberto, Manual (CEUA); Ciudad Bolivar, Vereda La Mina, 5.772073°, -76.032002°, 2400m, 1male, Leg. E. Garces, Net (CEUA); Gurane, Vereda Piedras Blancas, 6.291588°, -75.493345°, 2300m, 1female, Leg. A. Velez, Manual (CEUA). rospigliosii (Brèthes), 1920:31 (also 1937:2) (Volucella). Type locality: Peru, Chanchamayo region. HT M/F (MACN). Distr.: Colombia (Caldas (Anserma)), Peru. Refs.: Thompson et al. 1976:83 (cat.). Material examined. COLOMBIA: Caldas: Anserma, Vereda Palo Blanco, 5.231550°,-75.776772°, 1800m, 1male, Leg. A.L. Montoya, Manual on spider net (CEUA). Subgenus Volosyrpha Shannon hirtipes (Macquart), 1850:434 (130) (Volucella). Type locality: Bolivia, Yungas. HT M (MNHNP). Distr.: Colombia, Argentina, Bolivia, Brazil. Refs.: Macquart, 1850:434 (130), pi. 12, figs. 13 (scutellum), 13a (hind leg); Thompson et al. 1976:77 (cat.). rufitarse Thompson, 1976:83 (nom. nov. for tibiale Shannon). HT F (MACN). Distr.: Colombia (Antioquia (Caldas, Gómez Plata, Puerto Nare), Huila)), Costa Rica. Refs.: Thompson et al. 1976:83 (cat.); Restrepo & Carrejo, 2009:148 (cat.). Material examined. COLOMBIA: Antioquia: Caldas, Cariaño, Confines, Secundary forest, 6.090000°, 75.638000°, 1560m, 1male, i.2002, Leg. E.E. Martinez, Vsr. (MEFLG); Gómez Plata, Hacienda Normandia, 6.683154°, -75.216575°, 1800m, 1female, 29.x.1998, Leg. J.C. Agudelo, Net (MEFLG); Puerto Nare, 6.190087°, 74.587306°, 200m, 1female, M. Castaño & C. Vélez Leg., Pitfall copro (CEUA). Genus Ornidia Lepeletier & Serville Ornidia Lepeletier & Serville, in Latreille et al. 1828:786. Type species, Syrphus obesus Fabricius (orig. des.). Refs.: Curran, 1930d (key); Val 1872 (key, biometry, evolution,males genitalia); Thompson, 1972:106–107 (desc. and relationships); Thompson, 1991 (rev); Whittington & Rotheray, 1997:365–372 (Afrotropical distr. and desc. of larva); Thompson, 1999a:330 (key), Morales & Köhler, 2004:93–102 (key, rev. and floral preferences); Rotheray et al. 2005 (key and rev. of larvae); Carvalho–Filho & Esposito, 2009:59–64 (key, rev. and males genitalia comparison). Volucella, subg. Ornidia: Hull, 1949:348 (description) aemula (Willison), 1888:272 (Volucella). Type locality: Bolivia, Santa Cruz, Piedra Blanca. (4 km West of Corumba, Mato Grosso, Brazil). HT F (AMNH). Distr.: Colombia (Sucre), Bolivia, Brazil. Refs.: Val, 1972 (key, biometry, evolution, M genitalia); Thompson et al. 1976:69 (cat.); Thompson, 1991:251 (redesc.), fig. 1 (facial profile), 5 (wing), 8, 9, 13 (M genitalia complex). Material examined. COLOMBIA: Sucre: San Onofre, 9.646080°, -75.581129°, 5m, 11.x.2003, Leg. Entomology Group (CEUA). major Curran, 1930d:2. Type locality: "Brazil". HT M (AMNH). Distr.: Colombia (Amazonas (Caño Lorena), Antioquia (Alejandria, Ebéjico, Envigado, Medellín), Caldas (Manizales), Risaralda (Pereira), Santander (La Belleza), Valle del Cauca (Restrepo)), Brazil, Costa Rica, Ecuador, Panama, Peru. Refs.: Thompson et al. 1976:69 (cat.); Thompson, 1981:120 (desc. distributional notes); Thompson, 1991:255 (redesc.), fig. 1 (facial profile), 11, 15 (M genitalia complex); Morales & Köhler, 2004:93–102 (key, rev. and floral preferences); Rotheray et al. 2005:423 (larval key and desc.), fig 1, 2, 5, 9, 12, 15 (larval structures). 516 · Zootaxa 4122 (1) © 2016 Magnolia Press MONTOYA Material examined. COLOMBIA: Amazonas: Caño Lorena, -3.712698°, -70.333443°, 150m, 1.viii.1994 (ICN); Antioquia: Alejandria, Vereda San Jose, 6.371942°, -75.109771°, 1879m, 1male, Leg. A.L. Montoya, Net (CEUA); Ebéjico, 6.316833°, -75.781921°, 1480m, 1male, Leg. M. Castaño, Manual (CEUA); Envigado, 6.162140°, -75.563731°, 1800m, 1male, 1female, Leg. Castaño & Rivera Leg., Manual (CEUA); Medellín, Cola del Zorro, 6.036796°,-75.556175°, 2600m, 1male, Leg. M. Wolff (CEUA); Caldas: Manizales, Vereda El Palomar, 5.121763°, -75.644841°, 900m, 1male, Leg. Pérez & Vélez, Net (CEUA); Risaralda: Pereira, 4.795782°, 75.675455°, 1480m, 1male, Leg. M. Castaño, Manual (CEUA); Santander: La Belleza, Vereda La Playa, 5.866667°, -73.983333°, 2000m, 04.i.1997, Leg. Medina (UNAB); Valle del Cauca: Restrepo, Vereda Rio bravo, 3.825209°, -76.539032°, 1460m, 10.ii.1984, Leg. de I. Arévalo (ICN). obesa (Fabricius), 1775:763 (Syrphus). Type locality: "America Insulis"= Virgin Islands. HT M/F (UZMC). Distr.: Colombia (Amazonas (Leticia, Puerto Buenaventura Rio Igara–Paraná), Antioquia (Alejandria, Caldas, Cisneros, San Roque), Atlántico (Barranquilla), Boyacá (Aguaclara, Buenavista, Macanal, Miraflores, Muzo, Otanche, Santa María), Caldas (Anserma, La Dorada, Manizales, Pácora), Casanare (Aguazul, Villanueba, Yopal), Cauca (Miraflor), Cesar (Valledupar), Chocó (Coqui, Riosucio), Cundinamarca (Albán, Anapoima, Anolaima, Arbeláez, Bogotá, Caquezá, Chía, Chinauta, Choachí, Cogua, El Colegio, Fusagasugá, Girardot, Guaduas, Guayabetal, La Mesa, La Palma, Nilo, Pandi, San Antonio del Tequendama, Suba, Terra, Rueda), Magdalena (La Picua, Santa Marta), Meta (Acacías, Buenavista, Cumaral, La Macarena, Puerto Gaitán, Puerto Lleras, Puerto López, San Juan de Arama, Villavicencio), Nariño (Tumaco), Norte de Santander (Ocaña), Putumayo (Mocoa), Quindío (Córdoba), Risaralda (Pereira, La Pastora), Santander (Barbosa, Guepsa, La Belleza, Piedecuesta, Cimitarra), Tolima (Carmen de Apicala, Chaparral, Melgar, Ibagué), Valle del Cauca (La Unión, Palmira, Restrepo, Santiago de Cali), Vaupes (Nboa Koyo)), Widespread. Refs.: Val, 1972 (key, biometry, evolution, M genitalia); Thompson et al. 1976:69 (cat.); Thompson, 1981:120 (redesc. distributional notes); Thompson, 1991:256 (desc.), fig. 3 (facial profile), 4 (notopleuron and adjacent structures, dorsal view), 6 (wing), 10 and 14 (M genitalia complex); Whittington & Rotheray, 1997:365–372 (Afrotropical distr, illustration and desc. of larva); Rotheray et al. 2005:423 (larval key and desc.), fig. 3, 61 9, 12 (larval structures); Carvalho–Filho & Esposito, 2009:59–64 (key and rev), 61, fig. 6–7, 10, 13 (M genitalia complex). Material examined. COLOMBIA: Amazonas: Leticia, Rio Guayurita Ocoa, -4.205429°, -69.932808°, 73m, 2.v.2002, Leg. Students of Animal Systematics (ICN); Leticia, El Puerto, -4.205429°, -69.932808°, 73m, 18.xi.1970, Leg. M.A.Méndez (ICN); Puerto Buenaventura Rio Igara–Paraná, -2.277969°, -71.793312°, 130m, 1.iv.1975, Leg. I.C. Sáenz (ICN); Antioquia: Alejandria, Vereda San Jose, 6.371942°, -75.109771°, 1879m, 1 Female Leg. A.L. Montoya, Manual (CEUA); Caldas, Alto de San Miguel, 6.090000°, -75.638000°, 1560m, 05.ii.1998, Leg. R. Landinez, Net (UNAB); Cisneros, 6.538319°, -75.088603°, 1038m, 08.xi.2003, Leg. R. Parra (UNAB); San Roque, Corregimiento San Jose del Nus, 6.503175°, -74.926885°, 1800m, Leg. A.L. Montoya, Vsr. Fish (CEUA); Atlántico: Barranquilla, Bella Vista K.130 Carretera Villavo, 11.001465°, -74.795511°, 33m, 20.x.1970, Leg. G. Zambrano (ICN); Boyacá: Aguaclara, Hacienda Don Antonio, 4.874329°, -72.865297°, 580m, 15.iii.1970, Leg. P. A. Figueroa (ICN); Buenavista, Vereda Santo Domingo, 5.650832°, -74.286905°, 1482m, 03.xi.2002, Manual (UNAB); Macanal, 4.980023°, -73.311745°, 1850m, 20.viii.1973, Leg. M. Zambrano (UNAB); Miraflores, Vereda El Cajón, 5.196389°,-73.144444°, 1820m, 24.iii.1997, Leg. M. Mora (UNAB); Muzo, Peña negra, 5.533119°, -74.099392°, 850m, 14.v.1972, Leg. J. Corredor (UNAB); Páez, km91 carretera Bogotá–Villavo Pipiralito, 5.083333°, -73.050000°, 1200m, 20.ii.1973, Leg. E. Cruz (ICN); Otanche, 5.749824°, 74.249682°, 1050m, 11.v.1990, Leg. G. Villalba (UNAB); Santa María, Sendero Ecológico, 4.857735°, 73.266876°, 1200m, 20.iv.1997, Leg. G. Amat (ICN); Caldas: Anserma, Vereda San Pedro, 5.250776°, 75.795434°, 1669m, 1female, Leg. A.L. Montoya, Net (CEUA); La Dorada, 5.449480°, -74.667940°, 188m, 16.iv.1971, Leg. A. Gutiérrez (UNAB); Manizales, 5.072373°, -75.506634°, 2216m, 27.iv.2005, Leg. N. Luque. D. Jamaica, Net (UNAB); Pácora, 5.529230°, -75.459550°, 1815m, 15.iii.1999, Leg. J. Arbelaez. F. Betancourt (UNAB); Casanare: Aguazul, Vereda La Rosa, 5.175734°, -72.557936°, 290m, 27.x.1996, Leg. C. Gómez (UNAB); Villanueva, 4.628254°, -72.915737°, 400m, 22.vi.1996, Leg. J. Vargas (UNAB); Yopal, La Nieta, Finca Maracaibo, 5.306471°, -72.434005°, 350m, 18.ix.2003m, Leg. J. López, Pitfall (UNAB); Cauca: Miraflores, 2.588550°, -76.862722°, 1800m, 29.iii.2002, Leg. R. Hernández (UNAB); Cesar: Valledupar, 10.461934°, 73.253460°, 182m, 05.iv.1972, Leg. N. Esguerra (UNAB); Chocó: Coqui, 5.607387°, -77.328848°, 179m, Leg. CATALOGUE OF SYRPHIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 517 Entomology Group, Net (CEUA); Riosucio, Santatá, 7.440560°, -77.118900°, 7m, 4.iii.1978, Leg. H. Echeverri (ICN); Cundinamarca: Albán, 4.878333°, -74.438333°, 2251m, 09.xii.1995, Leg. Benincore (UNAB); Anapoima, 4.551687°, -74.535691°, 710m, 13.ix.2003, Leg. J. Cárdenas, Manual (UNAB); Arbeláez, 4.270965°, 74.419444°, 1417m, 09.vi.1996, Leg. A. Bejarano (UNAB); Bogotá, UNal I.C.N, 4.638194°, -74.084046°, 2567m, 20.ii.1971, Leg. J. Granada (ICN); Cáqueza, km91 carretera Bogotá–Villavo Pipiralito, 4.405278°, -73.950000°, 1743m, 6.viii.1989, Leg. L.P. Moreno (ICN); Chía, 4.861565°, -74.032534°, 2582m, 29.ix.2003, Leg. A. Rodríguez (UNAB); Chinauta, 4.308640°, -74.444130°, 1620m, 23.v.1999, Leg. Cely, Forero & Rojas (UNAB); Choachí, 4.529210°, -73.924817° 1827m, 18.v.1996, G. Ascencio. Leg. C. Alvarez (UNAB); Cogua, 5.066667°, 73.983333°, 2631m, 20.iii.1971, Leg. D'Silvestri (UNAB); El Colegio, 4.567024°, -74.433012°, 983m, 22.ii.1992, Leg. M. Daza (UNAB); Fusagasugá, 4.338328°, -74.363968°, 1771m, 1.xii.1969, Leg. D. Castro (ICN); Girardot, 4.300725°, -74.805450°, 281m, ii.1984, Leg. E. Vargas (UNAB); Guaduas, 5.069440°, -74.598052°, 1001m, 06.v.1999, Leg. J. Arbelaez & F. Betancourt (UNAB); Guayabetal, Villavista, 4.215287°, -73.814548°, 1000m, 5.vi.1970, Leg. R. Alvarez (ICN); La Mesa, San Javier, 4.629469°, -74.466633°, 1298m, 04.vi.1995, Leg. L. Palacios (UNAB); La Palma, 5.332998°, -74.383346°, 1462m, 20.v.1994, Leg. D. Angel, Net (UNAB); Nilo, 10 M Liraila, 4.306975°, -74.599994°, 420m, 1.vii.1972, Leg. B.N. Páez (ICN); Pandi, Vereda El Chorro, 4.187360°, 74.487134°, 1024m, 24.ix.2002, Leg. M. Duarte Fuentes (ICN); San Antonio del Tequendama, Laguna Pialagua, 4.616661°, -74.350002°, 1550m, Leg. G. Hernández (ICN); Suba, San Jose, 4.757031°, -74.088591°, 2555m, Leg. C. Rodríguez Medina (ICN); Terra, 4.636041°, -74.365618°, 1340m, 27.iv.1991, Leg. A. Castillo (ICN); Magdalena: La Picua, Parque Tayrona, 11.276050°, -74.195519°, 121m, 5.iii.1978, Leg. H. Echeverri (ICN); Santa Marta, Cimarca Guayabetal, 11.248643°, -74.211974°, 11m, 22.iv.1976, Leg. A. Acosta (ICN); Meta: Acacías, 3.987806°, -73.759747°, 522m 18.viii.1976, Leg. S. Rodríguez (UNAB); Buenavista, 5.650832°, 74.286905°, 1482m, Leg. G.J. Moreno (ICN); Cumaral, R.N. Kaliawirinae, 4.250000°, -73.486389°, 550m, 7.xi.2005, Leg. N. Ocampo (ICN); Cumaral, Bella Vista K.130 Carretera Villavo, 4.250000°, -73.486389°, 420m, 8.x.2009, Leg. R. Garon (ICN); Cumaral, Vereda San José, 4.250000°, -73.486389°, 480m, 28.iii.1993, Leg. C. Nelson (UNAB); La Macarena, 2.072107°, -73.955920°, 390m, 28.iii.1997, Leg. E. Bastidas (UNAB); Puerto Lleras, 3.268224°, -73.377700°, 250m, 27.v.1978, Leg. J. Uribe (UNAB); Puerto Gaitán, 4.310873°, -72.073758°, 160m, 1.xi.1977, Leg. P.A. Jiménez (ICN); Puerto López, 4.066137°, -73.048348°, 181m, 6.vi.1970, Leg. F. Sarmiento (ICN); San Juan de Arama, 3.350009°, -73.876655°, 450m, 25.iii.1997, Leg. Rodain (UNAB); Villavicencio, 4.150000°, -73.633333°, 467m, 14.vi.1972, Leg. C. Garzón (UNAB); Nariño: Tumaco, 1.808369°, -78.763991°, 2m, 15.i.1972, Leg. M. Corredor (UNAB); Norte de Santander: Ocaña, Vereda Agua de la virgen, 8.232143°, -73.350083°, 1202m, 25.i.1997, Leg. J. Santiago (UNAB); Putumayo: Mocoa, 1.150000°, 76.647500°, 695m, 31.iii.1996, Leg. L. Becerra (UNAB); Quindio: Córdoba, Vía Armenia, Km. 22, 4.393050°, 75.687778°, 1800m, 18.x.2003, Leg. R. Peña (UNAB); Risaralda: Pereira, Finca Calamar, 4.820499°, 75.633310°, 1820m, 28.v.1999, Leg. Cely, Forero & Rojas (UNAB); Pereira, PNR Ucumari, La Pastora, 4.708555°, -75.487224°, 2500m, 1.xi.1977, Leg. P.A. Jiménez (ICN); Santander: Barbosa, 5.932922°, 73.621119°, 1800m, 26.viii.2004, Leg. D. Mejía, Net (UNAB); Cimitarra, Bella Vista K.130 Carretera Villavo, 6.310723°, -73.944432°, 263, 5.x.1969, Leg E.Barrera. (ICN); Guepsa, Vereda San Isidro, 6.038430°, 73.581924°, 1483m, 28.xi.1997, Leg. S. Olarte (UNAB); La Belleza, Vereda La Playa, 5.866667°, -73.983333°, 2000m, 03.i.1997, Leg. Medina (UNAB); Piedecuesta, Ins. Pol. Mesa de Ruitoque, 6.987834°, -73.047684°, 1010m, 03.iv.1994, Leg. G. Jaimes (UNAB); Tolima: Carmen de Apicala, 4.150000°, -74.733333°, 380m, 22.v.1995, Leg. H. López (UNAB); Chaparral, Granja Sixto Iriarte, Barrio Beltrán, 3.750000°, -75.583333°, 1354m, 27.viii.2003, Leg.A. Díaz, Net (UNAB); Melgar, 4.203889°, -74.642778°, 323m, 01.vi.1997, Leg. J. Paez (UNAB); Honda, 5.190185°, -74.742401°, 255m, 29.xi.1982, Leg. A. Sobrino (ICN); Ibagué, Villavista, 4.440663°, -75.244141°, 1300m, 18.iv.1974, Leg. Rivera & Alcira (ICN); Mariquita, Vereda La Guardia, 5.250000°, -74.916667°, 960m, 23.ix.1978, Leg. G. Murillejo (ICN); Valle del Cauca: La Unión, 4.529890°, 76.108690°, 975m, 03.x.2003, Leg. R. Parra (UNAB); Obando, 4.569255°, -75.969261°, 939m, 04.xi.2003, Leg. A. Díaz, Net (UNAB); Palmira, Corpoica, 3.587073°, -76.248936°, 1059m, 15.x.2003, Leg. P. Rodríguez, Manual (UNAB); Restrepo, Vereda Rio Bravo, 3.811560°, -76.515178°, 1000m, 8.ii.1984, Leg. de I. Arévalo (ICN); Santiago de Cali, 3.378086°, -76.491055°, 1500, 1.v.1970, Leg. L. Angel (ICN); Sevilla, Buenavista, 4.268890°, 75.936100°, 2000m, 4.ii.1974, Leg. J. Aguirre (ICN); Vaupés: Nboa Koyo, 0.983621°’, -70.864236°, 180m, 1female, Leg. J. Gonzales, Net (CEUA). 518 · Zootaxa 4122 (1) © 2016 Magnolia Press MONTOYA whiteheadi Thompson, 1991:257. Type locality: Pamana, Cerro Campana. HT M (CAS). Distr.: Colombia (Antioquia), Costa Rica, Panama. Refs.: Thompson, 1991:257 (orig. desc.), fig. 12, 16 (M genitalia complex) 18 (habitus); Rotheray et al. 2005:424 (larval key and desc.), fig 4, 8, 11, 14 (larval structures). Acknowledgments I am grateful to Marta Wolff for inviting me to participate in the realization of the Catalogue of Syrphidae from Colombia. I would like to acknowledge the curators and directors of the visited collections for permission to study material in their care: CEUA, M. Wolff; ICN, C. E. Sarmiento; MEPB, A. Vélez; MEFLG, S. Orduz, J. Quiroz and UNAB, F. Serna, E. Vergara. I am grateful to F. Christian Thompson (Entomology Dept, Smithsonian Institution) for verification and identification of some syrphid specimens and for his support during the development of this project. I also thank N. Carrejo, C. Ruiz, and C. Restrepo of the MEUV for their help in the confirmation and identification of part of the material. I am also grateful to the Samuel Wendell Williston Diptera Research Foundation and the organizing Committee for financial assistance to attend the 6th International Symposium on the Syrphidae in Glasgow, (UK) and the 8th International Congress of Dipterology in Potsdam, Germany. Thanks to the Project “Diversidad de insectos de Colombia”, led by the Instituto de Investigación en Recursos Biológicos Alexander von Humboldt (IAvH). I thank the CODI (Comite para el Desarrollo de Investigaciones Universidad de Antioquia) Act number 20067, and to SOCOLEN for the “Beca de incentivo a la investigación Entomología en la Universidades Colombianas” (TG 03–2009). I also thank everyone who collects this invaluable material, especially to the members of the Entomology Group University of Antioquia (GEUA). Many thanks to the editorial board, especially to Carolina Henao Sepúlveda for useful comments that substantially enriched the final version of this manuscript. TABLE 1. Syrphidae species recorded from Colombia. Subfamily Genus Syrphinae Allograpta Argentinomyia Subgenus Species Author annulipes (Macquart, 1850) aperta Fluke, 1942 bilineella Enderlein, 1938 exotica (Wiedemann, 1830) falcata Fluke, 1942 hastata Fluke, 1942 neotropica Curran, 1936 obliqua (Say, 1823) quadricincta Enderlein, 1938 tectiforma Fluke, 1942 teligera Fluke, 1942 altissima (Fluke, 1945) bolivariensis (Fluke, 1945) browni (Fluke, 1945) columbiana (Enderlein, 1938) crenulata (Williston, 1891) festiva Fluke, 1945 longicornis (Walker, 1837) luculenta (Fluke, 1945) neotropica Curran, 1937 nigrans (Fluke, 1945) ...Continued on next page CATALOGUE OF SYRPHIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 519 TABLE 1. (Continued) Subfamily Genus Subgenus Species Author opaca (Fluke, 1945) peruviana (Shannon, 1927) rex Fluke, 1945 thiemei (Enderlein, 1938) tropica (Curran, 1937) Calostigma ornatipes (Curran, 1927) Claraplumula latifacies (Shannon, 1927) Dasysyrphus lotus (Williston, 1887) Eupeodes volucris Osten Sacken, 1877 Fazia alta Curran, 1936 colombia Curran, 1925 fasciata Curran, 1932 fascifrons (Macquart, 1846) imitator (Curran, 1925) nasigera Enderlein, 1938 roburoris (Fluke, 1942) rostrata (Bigot, 1884) similis Curran, 1925 strigifacies (Enderlein, 1938) Hermesomyia wulpiana Lynch Arribalzaga, 1891 Hybobathus arx (Fluke, 1936) flavipennis (Wiedemann, 1830) lineatus (Macquart, 1846) lividus (Schiner, 1868) norina (Curran, 1941) persimilis (Curran, 1930) quadrilineatus (Enderlein, 1938) delicatula (Hull, 1943) gracilis (Williston, 1891) gigantea (Schiner, 1868) tristani Zumbado, 2000 willistoni Thompson, 1976 anthiphates (Walker, 1849) cultrinus (Curran, 1939) dimidiatus (Fabricius, 1781) filissima (Hull, 1943) funebris Macquart, 1834 gastrostactus (Wiedemann, 1830) hyalipennis (Curran, 1930) luctuosus (Bigot, 1883) sativa (Curran, 1941) Leucopodella Mimocalla Ocyptamus ...Continued on next page 520 · Zootaxa 4122 (1) © 2016 Magnolia Press MONTOYA TABLE 1. (Continued) Subfamily Genus Subgenus Orphnabaccha Pelecinobaccha Platycheirus Pseudodoros Relictanum Rhinoprosopa Carposcalis Dioprosopa Species Author stenogaster (Williston, 1888) verona (Curran, 1941) vierecki (Curran, 1930) ampla (Fluke, 1942) calda (Walker, 1852) coerulea (Williston, 1891) delimbata (Enderlein, 1938) pteronis (Fluke, 1942) trabis (Fluke, 1942) adspersa (Fabricius, 1805) alucard Miranda, 2014 andrettae Miranda, 2014 avispas Miranda, 2014 cyclops (Hull, 1947) clarapex (Wiedemann, 1830) dracula (Hull, 1943) gracilitas Miranda, 2014 hiantha (Hull, 1943) ida (Curran, 1941) oviphora (Hull, 1943) ovipositoria (Hull, 1943) pandora (Hull, 1942) peruviana Shannon, 1927 pilipes (Schiner, 1868) pucallpa Miranda, 2014 squamagula Miranda, 2014 telescopica Curran, 1930 transatlantica (Schiner, 1868) chalconota (Philippi, 1865) ecuadoriensis (Fluke, 1945) fenestrata (Macquart, 1842) hesperius (Vockeroth, 1990) clavatus (Fabricius, 1794) crassum (Walker, 1852) nero Curran, 1939 nigrocilia (Hull, 1943) schwarzi (Curran, 1939) flavophylla (Hull, 1943) lucifer (Hull, 1943) nasuta (Bigot, 1884) sycorax (Hull, 1947) ...Continued on next page CATALOGUE OF SYRPHIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 521 TABLE 1. (Continued) Subfamily Genus Subgenus Species Author abdominalis Sack, 1920 cornuta Hull, 1944 limbipennis Williston, 1891 nigra Schiner, 1868 panamana Hull, 1944 pygophora Schiner, 1868 vera Hull, 1944 Scaeva occidentalis Shannon, 1927 Styxia eblis Hull, 1943 Syrphus laceyorum Thompson, 2000 octomaculatus Walker, 1837 phaeostigma Wiedemann, 1883 shorae Fluke, 1950 Talahua fervida (Fluke, 1945) Toxomerus anthrax (Schiner, 1868) apegiensis (Harbach, 1974) aquilinus Sack, 1941 brevifacies (Hull, 1941) centaureus (Hull, 1951) dispar (Fabricius, 1794) duplicatus (Wiedemann, 1830) flaviplurus (Hall, 1927) floralis (Fabricius, 1798) hieroglyphicus (Schiner, 1868) idalius (Hull, 1951) insignis (Schiner, 1868) lacrymosus (Bigot, 1884) laenas (Walker, 1852) mutuus (Say, 1829) nigripunctus (Enderlein, 1938) nitidus (Schiner, 1868) norma (Curran, 1930) ochraceus (Hull, 1942) ophiolinea (Hull, 1943) pallipes (Bigot, 1884) paraduplicatus Borges, 2009 pictus (Macquart, 1842) politus (Say, 1823) polygraphicus (Hull, 1940) procrastinatus Metz, 2001 pulchellus (Macquart, 1846) Salpingogaster ...Continued on next page 522 · Zootaxa 4122 (1) © 2016 Magnolia Press MONTOYA TABLE 1. (Continued) Subfamily Genus Species Author saphiridiceps (Bigot, 1884) sylvaticus (Hull, 1943) vierecki (Curran, 1930) virgulatus (Macquart, 1850) watsoni (Curran, 1930) Trichopsomyia urania Hull, 1949 Tuberculanostoma antennatum Fluke, 1943 pectinis Fluke, 1943 bucephalus (Wiedemann, 1830) mellinoides (Macquart, 1846) plaumanni Fluke, 1937 falcatus Williston, 1887 cyaneiventris (Macquart, 1846) marceli Curran, 1936 conopsoides Macquart, 1842 pithecofascia Hull, 1944 angustus (Macquart, 1846) diaphanus Sack, 1921 trivittatus Curran, 1925 Cacoceria willistoni Hull, 1949 Ceriana cacica (Walker, 1860) Chalcosyrphus americana (Schiner, 1868) vagabondans (Hull, 1941) Copestylum flaviventre Macquart, 1846 Lepidopsis viride (Williston, 1888) Megametopon amethystinum (Bigot, 1875) Phalacromya apicale (Loew, 1866) aster (Curran, 1939) aureum (Fluke, 1951) brunnigaster (Hull, 1943) caesariatum (Williston, 1891) chalybescens (Wiedemann, 1830) cinctiventre (Curran, 1930) circumdatum (Walker, 1857) circumscriptum (Curran, 1939) contumax (Curran, 1939) correctum (Curran, 1927) corumbense (Curran, 1930) delila (Hull, 1950) dispar (Macquart, 1846) florella (Hull, 1944) Xanthandrus Microdontinae Subgenus Xanthandrus Menidon Microdon Chymophila Mixogaster Peradon Eristalinae Copestylum ...Continued on next page CATALOGUE OF SYRPHIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 523 TABLE 1. (Continued) Subfamily Genus Subgenus Species Author flukei (Curran, 1936) fornax (Townsend, 1895) fraudulentum (Williston, 1891) frauenfeldi (Schiner, 1868) fulvicorne (Bigot, 1883) hambletoni (Fluke, 1951) hydrofenestra (Hull, 1943) imitans (Curran, 1926) inconsistens (Curran, 1939) infractum Thompson, 1981 lanei (Curran, 1936) latevitatum (Curran, 1939) longirostre (Macquart, 1846) macquarti (Curran, 1926) meretricias (Williston, 1888) mocanum (Curran, 1936) muscarium (Thomson, 1869) nigripes (Bigot, 1857) obscurior (Curran, 1939) opinator (Williston, 1891) pallens (Wiedemann, 1830) parina (Fluke, 1951) persimile (Williston, 1888) pica (Schiner, 1868) pictum (Wiedemann, 1830) posticum (Say, 1829) prasinus (Schiner, 1868) pseudopallens Thompson, 1981 punctiferum (Bigot, 1875) punctigena (Hull, 1937) purpuriferum (Bigot, 1875) roraima (Curran, 1939) selectum (Curran, 1939) sexmaculatum (Palisot de Beauvois, 1819) spinithorax (Lynch Arribalzaga, 1892) subcoeruleum (Rondani, 1863) tumicephalum (Hull, 1943) tympanitis (Fabricius, 1805) ulrica (Hull, 1950) vagum (Wiedemann, 1830) verdigaster (Hull, 1943) ...Continued on next page 524 · Zootaxa 4122 (1) © 2016 Magnolia Press MONTOYA TABLE 1. (Continued) Subfamily Genus Subgenus Species Author vierecki (Curran, 1925) villarica (Fluke, 1951) virescens (Williston, 1891) viride (Williston, 1888) wulpi (Goot, 1864) gibberum (Schiner, 1868) rospigliosii (Brèthes, 1820) hirtipes (Macquart, 1850) rufitarse Thompson, 1976 rostrata (Macquart, 1846) calops (Bigot, 1880) fulvitarsis (Rondani, 1863) Eoseristalis bogotensis Macquart, 1842 Eristalis tenax (Linnaeus, 1858) Eumerus funeralis (Meigen, 1822) Habromyia magnifica (Bigot, 1880) mexicanus (Macquart, 1842) Lepidomyia colombia Curran, 1929 Lycastrirhyncha nitens Bigot, 1859 Macrometopia maculipennis Thompson, 1999 aberrans Shannon, 1927 analis (Macquart, 1846) apis Thompson, 2002 colombii (Macquart, 1850) inversa Shannon, 1927 mystacia Fluke, 1939 nigra Shannon, 1927 rubicunda Curran, 1940 salti Curran, 1929 anna Curran, 1936 basiger (Walker, 1860) circumdatus (Bigot, 1875) fucatus Hull, 1930 laconicus (Walker, 1852) obscurus Hine, 1824 scitus (Walker, 1857) unicolor Wulp, 1883 flavens Hippa & Ståhls, 2013 neurostigma Hippa, 1982 nigropilosa Thompson, 1983 vittipes Curran, 1941 Viereckomyia Volosyrpha Dolichogyna Eristalinus Eristalis Lejops Mallota Meromacrus Meropidia Monoceromyia Eristalodes Asemosyrphus Mallota ...Continued on next page CATALOGUE OF SYRPHIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 525 TABLE 1. (Continued) Subfamily Genus Subgenus Ornidia Orthonevra Palpada Polybiomyia Quichuana Species Author aemula (Willison, 1888) major Curran, 1930 obesa (Fabricius, 1775) whiteheadi Thompson, 1991 argentina Brèthes, 1822 shannoni (Curran, 1925) aemula (Williston, 1891) agrorum (Fabricius, 1787) albifrons (Wiedemann, 1830) amazon (Curran, 1930) atrimana (Loew, 1866) conica (Fabricius, 1805) cosmia (Schiner, 1868) elegans (Blanchard, 1852) erratica (Curran, 1930) expicta (Walker, 1860) flavoscutellata (Hull, 1937) florea (Hull, 1925) furcata (Wiedemann, 1819) geniculata (Fabricius, 1805) macula (Sack, 1841) mexicana (Macquart, 1847) minutalis (Williston, 1891) mirabilis (Hull, 1925) monticola (Roder, 1892) ochracea (Williston, 1888) prietorum Mengual, 2008 pusilla (Macquart, 1842) ruficeps (Macquart, 1842) rufiventris (Macquart, 1846) scutellaris (Fabricius, 1805) suprarufa Thompson, 1999a taenia (Wiedemann, 1830) thalia (Hull, 1942) vierecki (Curran, 1930) vinetorum (Fabricius, 1798 bergrothi (Williston, 1892) odontomera (Curran, 1941) angustiventris (Macquart, 1855) brevicera Hull, 1946 calathea Shannon, 1925 ...Continued on next page 526 · Zootaxa 4122 (1) © 2016 Magnolia Press MONTOYA TABLE 1. (Continued) Subfamily Genus Subgenus Rhingia Sphiximorpha Spilomyia Sterphus Ceriogaster Species Author dolorosa Hull, 1946 fasciata (Sack, 1841) picadoi Knab, 1913 quixotea Hull, 1946 longirostris Fluke, 1943 nigra Macquart, 1846 roederii Williston, 1888 rubrobrunnea (Hull, 1944) wulpii (Williston, 1888) gratiosa Wulp, 1888 andicus Hippa, 1978 aureus Hippa, 1978 chloropyga (Schiner, 1868) coarctatus (Wiedemann, 1830) cybele (Hull, 1951) cydippe Hippa, 1994 fascithorax Kertesz, 1910 fassli Hippa, 1978 icarum Hippa, 1978 jamaicensis (Gmelin, 1890) janzeni Thompson, 1994 nitidicollis Hippa, 1978 ochripes Hippa, 1978 plagiatus Wiedemann, 1830 pilifer Hippa, 1978 sapphirifer Hippa, 1978 transversa (Walker, 1857) References Agassiz, L. 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Tijdschrift voor entomologie, 31, 359–376. Zuijen, van M.P. & Nishida, K. (2010) Life history and immature stages of Allograpta zumbadoi Thompson, a phytophagous flower fly (Diptera: Syrphidae: Syrphinae). Studia dipterologica, 7, 37–51. Zumbado, M.A. & Thompson, F.C. (1997) Nuevas especies de Sterphus (Diptera, Syrphidae) de Costa Rica con notas sobre especies presentes en Costa Rica. The Southwestern Entomologist, 22, 79–90. CATALOGUE OF SYRPHIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 537 Zootaxa 4122 (1): 538–548 http://www.mapress.com/j/zt/ Copyright © 2016 Magnolia Press Article ISSN 1175-5326 (print edition) ZOOTAXA ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.4122.1.40 http://zoobank.org/urn:lsid:zoobank.org:pub:1CB61FE5-0F37-4C11-9CB2-EFCF0038B414 FAMILY MICROPEZIDAE STEPHEN MARSHALL1, ANDERSON CARDONA2 & MARTHA WOLFF2 1 School of Environmental Sciences, University of Guelph, Guelph, Ontario, Canada N1G 2W1. E-mail: samarsha@uoguelph.ca. Grupo de Entomología, Instituto de Biología, Universidad de Antioquia. Colombia. Emails: martha.wolff@udea.edu.co, andybond8@gmail.com. *Corresponding author: samarsha@uoguelph.ca 2 Abstract Sixty species and 14 genera of Micropezidae are recorded from Colombia, including four species of Micropezinae and 56 species of Taeniapterinae. The following species are newly recorded from the country: Grallipeza affinis, G. ecuadoriensis, G. obscura, G. nigrinotata, G. pseudosimplex, Ptilosphen comis, P. conveniens, P. enderleni, P. facetus and P. mimicus. Key words: catalogue, Colombia, Micropezidae, distribution Introduction The family Micropezidae Loew occurs in all zoogeographic regions, but is particularly diverse in the Neotropical Region. The total of 51 species recorded for Colombia by Steyskal (1968) is a great underestimate because most neotropical species of Micropezidae remain undescribed, and most described species are effectively unidentifiable except by direct comparison with the type specimens. Fourteen genera and 60 species are listed here, but several genera remain difficult to define, and others are clearly paraphyletic as treated by Steyskal (1968). The entire New World fauna is currently under revision by the first author and associated graduate students, and the number of undescribed species (manuscript names) currently greatly exceeds the number of published names. We expect that at least 200 species of Micropezidae occur in Colombia. At the family level, adult micropezids are recognizable as distinctively long-legged, slender flies of 4-20 mm (usually 6-12 mm) with veins R4+5 and M strongly convergent at the wing tip, katepisternal bristles present (usually in a vertical row), arista dorsobasal, ocellar bristle absent, and vibrissa absent. Adults of some genera are attracted to decomposing matter and feces, but most species for which biological data are available develop in dead wood, roots or stems. Colombian species are recorded from sea level to 3000m. Most species for which the type locality is Colombia were described by Enderlein (1922), on the basis of specimens added to the collection in Berlin by Otto Thieme. Thieme, a professor at the Sophien Real Gymnasium in Berlin until 1907, made significant collections in South America but little is known of the details of his trips, and his micropezid specimens are simply labeled "cordillera tierra caliente" or “cordillera tierra templada”. Type depositories are here given only if we have examined type material, which is the case for the majority of species. A single sex is listed for syntype series if a specimen has been selected as lectotype but not yet designated. The classification adopted here is based on the catalogue by Skeyskal (1968) and the micropezid chapter in the Manual of Central American Diptera chapter by Marshall (2010). It also incorporates unpublished information from the first author's unpublished research on the New World Taeniapterinae, which includes revisions of Grallipeza, Ptilosphen, and Mesoconius (now including Zelatractodes). New country records and corrected distributions are given for those genera only. New synonymies explained and justified in unpublished papers are indicated here with the noncommittal "probably a synonym of". The division of Taeniaptera sensu Steyskal into Paragrallomyia, Grallomyia, and Taeniaptera is based on Marshall (2010) and molecular phylogenetic data in Jackson et al. (2015). The inclusion of Zelatractodes in Mesoconius is based on Marshall (2015). 538 Accepted by S. Nihei: 25 May 2015; published: 14 Jun. 2016 We underline Colombian Department names and parenthetically note specific localities. At the end, we provide a table that includes all species found in Colombia (Table 1). Acronyms for type depositories ANSP—Academy of Natural Sciences of Philadelphia, Pennsylvania, USA. BMNH—British Museum Natural History, United Kingdom, London. CUIC—Cornell University Insect Collection, Ithaca, New York, USA. NHMW—Naturhistorisches Museum Wien, Vienna, Austria. SMTD—Staatliches Museum für Tierkunde, Dresden, Germany. UMO—Oxford University Museum of Natural History, United Kingdom. USNM —National Museum of Natural History, Washington D.C., USA. ZMHB—Museum für Naturkunde der Humboldt-Universität, Berlin, Germany. ZMUC—Zoological Museum University of Copenhagen, Denmark. ZMUH—Finnish Museum of Natural History, Helsinki, Finland. List of abbreviations auct.—auctorum (authors) cat.—catalogue distr.—distribution refs.—references PNN—Parque Nacional Natural F (f#)—Female M (m#)—Male Catalogue of Micropezidae of Colombia Family Micropezidae Loew, 1862 Subfamily Micropezinae Enderlein, 1922 Genus Micropeza Meigen, 1803 Tylos Meigen, 1800: 31. Type species, Musca corrigiolata Linaeus (Coquillett, 1910: 618). Suppressed by I.C.Z.N., 1955: 267. Tylus, emend. Micropeza Meigen, 1803: 276. Type species, Musca corrigiolata Linnaeus (mon.). Neriocephalus Enderlein, 1922: 160. Type species, Micropeza appendiculata Schiner (orig. des.). Metopobrachia Enderlein, 1922: 161. Type species, Micropeza obscura Bigot (orig. des.). Cliopeza Enderlein, 1922: 162. Type species, Calobata pectoralis Wiedemann (orig. des.). Neotylus Hendel, 1932: 121. Type species, Micropeza brasiliensis Schiner (orig. des.). Neotypus , error. Protylos Aczél, 1949: 238 (as subgenus). Type species, Micropeza stigmatica Wulp (orig. des.). Phantasma Robineau-Desvoidy, 1830: 739. Type species, Musca filiformis Fabricius (= corrigiolata L.) (orig. des.). annulata Schiner, 1868: 249. Type locality: "South America” (Venezuela). Distr.: Colombia, Venezuela. Refs.: Enderlein, 1922: 162; Steyskal, 1968: 2 (cat.). bogotana (Enderlein), 1922: 162 (Cliopeza). Type locality: Colombia, Bogotá. HT M (ZMHB). Distr.: Colombia (Bogotá D.C). Ref.: Hennig, 1936: 206 (rev.); Steyskal, 1968: 3 (cat.). CATALOGUE OF MICROPEZIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 539 brasiliensis Schiner, 1868: 249. Type locality: Brazil and Colombia. Distr.: Brazil, Colombia, Panama, Venezuela. Refs.: Hendel, 1932:17; Steyskal, 1968: 3 (cat.). columbiana (Enderlein), 1922: l61 (Metopobrachia). Type locality: Colombia, Bogotá and Ibagué. HT F (ZMHB). Distr.: Colombia (Cundinamarca (Bogotá); Tolima (Ibagué)). Refs.: Hennig, 1936: 206 (rev.); Steyskal, 1968: 3 (cat.). Subfamily Taeniapterinae Cresson, 1930 Genus Calosphen Hennig, 1934 Calosphen Hennig, 1934: 310. Type species, Tanypoda strophium Enderlein (orig. des.). Ref.: Steyskal, 1968: 5 (cat.). pleuralis (Cresson), 1930: 321 (Rainieria). Type locality: Colombia, San Lorenzo Mountains, Vista Nieve. HT F? (ANSP), PT F (CNM). Distr.: Colombia (Magdalena (Vista Nieve, San Lorenzo). Refs.: Hennig, 1934: 311 (rev.); Steyskal, 1968: 5 (cat.); Roback, 1969: 553. strophium (Enderlein), 1922: 203 (Tanypoda). Type locality: Colombia, Cordillera, "tierra templada". HT F (ZMHB). Distr.: Colombia (Cordillera, “tierra templada”), Venezuela. Refs.: Hennig, 1934: 311 (rev.); Steyskal, 1968: 5 (cat.). Genus Cliobata Enderlein, 1923 Cliobata Enderlein, 1923: 540. Type species, Catobata guttipennis Wiedemann (orig. des.). Ref.: Steyskal, 1968: 5 (cat.). biguttata (Enderlein), 1922: 205 (Tanypoda). Type locality: Colombia, "tierra templada". HT F (ZMHB). Distr.: Colombia (Cordillera, “tierra templada”). Ref.: Steyskal, 1968: 5 (cat.). diadema (Wiedemann), 1830: 533 (Calobata). Type locality: "Brazil". HT F (NHMW). Distr.: Colombia (Boyacá) Muzo), Brazil. Refs.: Enderlein, 1922: 212; Hennig, 1935: 41; Steyskal, 1968: 5 (cat.). nigrifilum (Walker), 1852:393 (Calobata). Type locality: "South America" HT F (BMNH). Genus Grallipeza Rondani, 1850 Grallipeza Rondani, 1850: 180. Type species, Calobata unimaculata Macquart (orig. des.). Refs.: Enderlein, 1922: 189; Steyskal, 1968: 6 (cat.). Systellapha Enderlein, 1922: 189. Type species, ornatithorax Enderlein (orig. des.). affinis Hennig, 1934: 308 Type locality: Bolivia, Mapiri, San Carlos HT F (SMTD) . Distr.: Colombia (Vichada (PNN EI Tuparro, Cerro Tomas); Meta (PNN Sierra de la Macarena, Cano Cuja); Caquetá (PNN Serrania de Chiribiquete Cunare-Anu); Putumayo (PNN La Paya, Cabaña La Paya)), Bolivia, Brazil, Ecuador. Refs.: Steyskal, 1868: 6 (cat.); Hennig, 1934: 308 (rev.). New record to Colombia. ecuadoriensis (Enderlein), 1922: 193 (Systellapha). Type locality: Ecuador. Guayaquil. HT F (ZMHB). Distr.: Colombia (Amazonas (PNN Amacayacu Malambata), Brazil, Guyana, Ecuador, Peru, Trinidad, Venezuela. Refs.: Steyskal, 1868: 7 (cat.); Hennig, 1934: 306 (rev.). New record to Colombia. obscura Hennig, 1934: 300. Type locality: Bolivia, Mapiri 21.03, S. Carlos. HT F (SMTD). Distr.: Colombia (Putumayo (PNN La Paya Cabana Viviano, Varzea)), Bolivia, Brazil, Ecuador, Guyana, Peru. Ref.: Steyskal, 1868: 7 (cat.). New record to Colombia. 540 · Zootaxa 4122 (1) © 2016 Magnolia Press MARSHALL ET AL. nigrinotata Hennig, 1934: 304. Type locality: Bolivia, Mapiri. HT F (SMTD). Distr.: Colombia (Amazonas (PNN Amacayacu, Matamata); Magdalena (PNN Tayrona, Zaino); Bolivar (SFF Los Colorados, Alto el Mirador); Vichada (PNN El Tuparro, Centro Administrativo; Bosque Sabana), Bolivia, Brazil, Trinidad. Ref.: Steyskal, 1868: 7 (cat.). New record to Colombia. pseudosimplex Hennig, 1934: 309. Type locality: Bolivia, Mapiri. HT F (SMTD) Distr: Colombia (Amazonas (PNN Amacayacu, San Martin; Matamata), Peru, Bolivia, Brazil. Refs.: Steyskal, 1868: 8 (cat.); Hennig, 1934: 309 (rev.). New record to Colombia. suavis (Enderlein), 1922: 194 (Systellapha). Type locality: Brazil, Pará, Ilia Marajó. Soure. HT F (ZMHB) Distr.: Colombia (Magdalena (Magdalena Valley, El Banco)), Brazil. Refs.: Hennig, 1934: 306 (rev.); Steyskal, 1968: 8 (cat.). Probable synonym of ecuadoriensis unifasciata (Fabricius), 1805: 262 (Calobata). Type locality : "South America". LT F (ZMUC). Distr.: Colombia (Cordillera “tierra caliente”), Guyana, Panama. Refs.: Hennig, 1934: 302 (rev.); Steyskal, 1968: 8 (cat.). columbiana Enderlein, 1922: 193 (Systellapha). female Ht headless ZMHB: Colombia, Cordillera, "tierra caliente". Refs.: Steyskal, 1968: 8 (cat.). unimaculata (Macquart), 1846: 343 (sep. p. 2151, pl. 19, fig. 4 (Calobata). Type locality: "Colombia". HT F (UMO). Distr.: Colombia, Peru, Brazil. Refs.: Hennig, 1934: 300; Steyskal, 1968: 8 (cat.). Probable synonym of unifasciata. Genus Mesoconius Enderlein, 1922 Mesoconius Enderlein 1922: 176. Type species, infestus Enderlein (orig. des.) Zelatractodes Enderlein 1922: 169. Type species, filipes Enderlein (orig. des.). Aristobata Frey, 1927: 69. Type species, melini Frey (orig. des). albimanus (Enderlein), 1922: 212 (Scipopus). Type locality: Peru. Cuzco, Callanga, Pinipini Valley, 1500m, 1900, O. Garlepp. Enderlein indicates in his description that there is a single type male, but the Enderlein collection in MNBG includesonly two "types", both good female specimens examined by the first author in 2002. Distr.:Colombia (Villa Elvira; Cordillera “tierra caliente”), Bolivia, Peru. Refs.: Frey, 1927: 73; Steyskal, 1968: 9 (cat.). auristrigatus Enderlein, 1922: 178. Type locality: Colombia. HT F (ZMHB). Distr.: Colombia. Ref.: Steyskal, 1968: 9 (cat.). enderleini Frey, 1927: 72. Type locality: Colombia. Aguatal. HT F (ZMUH). Distr.: Colombia. Ref.: Steyskal, 1968: 9 (cat.). flaviceps Enderlein, 1922:178. Type locality: Colombia. HT M (MNBG). Distr.: Colombia. Refs.: Steyskal, 1968: 9 (cat.). garleppi (Enderlein), 1922: 201 (Tanypoda). Type locality: Peru. HT M (ZMHB). Distr.: Colombia, Ecuador, Bolivia. Ref.: Steyskal, 1968: 20 (cat.). rufiventris (Enderlein), 1922: 204 (Tanypoda). Type locality: Colombia. HT F (ZMHB). Distr.: Colombia. Ref.: Steyskal, 1968: 20 (cat.). ujhelyianus Enderlein, 1922: 177. Type locality: Colombia, Sierra San Lorenzo. HT M (ZMHB). Distr.: Colombia (Magdalena). Ref.: Steyskal, 1968: 9 (cat.). versicolorus (Enderlein), 1922: 203 (Tanypoda versicolora). Type locality: Colombia. HT F (ZMHB). Distr.: Colombia. Ref.: Steyskal, 1968: 20 (cat.). CATALOGUE OF MICROPEZIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 541 Genus Metasphen Frey Metasphen Frey, 1927: 71. Type species, Metasphen vigilans Frey (orig. des.). gilvimana (Cresson) 1926: 269. Type locality: Panama, Alhajuela. HT M (USNM). Distr.: Colombia (Amazonas (PNN Amacayacu, San Martín), Valle del Cauca (PNN Farallones de Cali, Alto Anchicayá)), Costa Rica, Ecuador, Honduras, Panama. Refs.: Steyskal, 1968: 14 (cat.); Marshall, 2004: 531. obscura (Hennig), 1935: 65. (Rainieria (Globomyia)). Type locality: Peru, Meshagua, Urubamba River. HT F (SMTD). Distr.: Colombia (Amazonas (PNN Amacayacu, Mocagua, Matamata)), Ecuador, Peru. Refs.: Steyskal, 1968: 14 (cat.); Marshall, 2004: 534. vigilans Frey 1927:71. Type locality: Brazil, Amazonas, Taracuá, Rio Vaupéz, and Manaus. LT F (MZHF) (des. by Marshall, 2004: 533). Distr.: Colombia (Amazonas (PNN Amacayacu, Matamata), Brazil, Ecuador. Refs.: Steyskal, 1968: 10 (cat.); Marshall, 2004: 533. Genus Plocoscelus Enderlein, 1922 Plocoscelus Enderlein, 1922: 226. Type species, Calobata arthritica Wiedemann (orig. des.). Ref.: Steyskal, 1968: 10 (cat.). Rhoecius Enderlein, 1922:227. Type species, valgus Enderlein (orig. des.) = niger (Schiner). arthriticus (Wiedemann), 1830: 546 (Calobata). Type locality: Brazil HT M (ZMHB). Distr.: Mexico to Panama, Surinam, Venezuela, Colombia (Isla Gorgona; Cordillera “tierra caliente”), ?Peru. Refs.: Enderlein, 1922: 227; Steyskal, 1968: 10 (cat.). myrmex Schiner, 1868:254 (Cardiacephala). Type locality: “South America”. Distr.: Colombia (Magdalena (Aracataca, Hacienda Victoria, Sierra San Lorenzo). brevipennis (Walker), 1852: 389 (Calobata). Type locality: "Brazil". Distr.: Bolivia, Brazil, Colombia (Cordillera “tierra caliente”), Ecuador, Peru, Surinam, Venezuela. Refs.: Enderlein, 1922: 227; Steyskal, 1968: 10 (cat.); Albuquerque, 1989: 22. harenosus Cresson, 1930: 333 (as Ploscelus, lapsus). Type locality: Colombia, Puerto Colombia, Atlántico. HT M (ANSP). Distr.: Colombia (Atlántico (Puerto Colombia)), Mexico to Panama, Guyana. Ref.: Steyskal, 1968: 10 (cat.). niger (Schiner), 1868: 255 (Cardiacephala). Type locality : "South America". Distr.: Mexico to Panama, Bolivia, Brazil, Colombia, Guyana, Peru, Venezuela. Refs.: Steyskal, 1968:10 (cat.). valgus Enderlein, 1922: 228 (Rhoecius). Type locality: Bolivia, Sara; Colombia. Genus Poecilotylus Hennig, 1934 Subgenus Poecilotylus Hennig, 1934 obliqua (Fabricius), 1805: 261 (Calobata). Type locality: "South America". HT F (ZMUC). Distr.: Guatemala to Panama, Bolivia, Brazil, Colombia (Cordillera, “tierra caliente”), Guianas, Peru. Venezuela Refs.: Hennig, 1934: 102; Steyskal, 1968:12 (cat.). Genus Ptilosphen Enderlein, 1922 Ptilosphen Enderlein, 1922: 222. Type species, Calobata insignis Wiedemann (orig. des.). Ref.: Steyskal, 1968: 13 (cat.). 542 · Zootaxa 4122 (1) © 2016 Magnolia Press MARSHALL ET AL. comis Cresson, 1930: 347. Type locality: Guyana, Kamakusa. HT M (ANSP). Distr.: Colombia (Amazonas (Leticia)), Bolivia, Brazil, Ecuador, Guyana, Peru. Ref.: Steyskal, 1968: 13 (cat.). New record to Colombia. subgentilis Hendel, 1936: 71. Type locality: Brazil, Pernanbuco, Recife; Pará, Belém. conveniens (Wulp) 1897: 373 (Calobata). Type locality: Panama, Bugaba. ST F (BMNH) Distr.: Colombia (Meta (23km N.W. Villavicencio, quebrada Susumuco)), Bolivia, Brazil, Costa Rica, Peru, Venezuela. Ref.: Steyskal, 1968: 13 (cat.). New record to Colombia. cyaneiventris (Macquart), 1846: 343 (sep. p. 215), pl. 19, fig. 5 (Calobata). Type locality: "Colombia". Distr.: Colombia (Magdalena (Aracataca); Bogotá (probably error); Cordillera “tierra caliente”; Hac. Pehlke; ?Cartagena)), Mexico to Panama, Venezuela, Brazil. Refs.: Enderlein, 1922: 224; Cresson, 1930: 346; Hennig, 1934: 316; Steyskal, 1968: 13 (cat.). enderleini Cresson 1930: 346. Type locality: Peru, Puerto Bermúdez, Río Pichís. HT M (CUIC). Distr.: Brazil, Colombia, Ecuador, Peru. Ref.: Steyskal, 1968: 13 (cat.). New record to Colombia. facetus Enderlein 1922: 225 (Ptilosphen). Type locality: Ecuador, Balzapamba. HT M (ZMHB) Distr.: Colombia (Valle del Cauca (Buenaventura, Anchicaya), Ecuador. Ref.: Steyskal, 1968: 13 (cat.). New record to Colombia. gentilis Cresson, 1930: 348. Type locality: Costa Rica, San Carlos. HT M (USNM) Distr.: Colombia (Isla Gorgona), Costa Rica, Ecuador, Panama. Refs.: Hennig, 1934: 318 (rev.); Steyskal, 1968: 13 (cat.). insignis (Wiedemann), 1830: 533 (Calobata). Type locality: “Brazil”. ST M (ZMHB) Distr.: Colombia (Cordillera “tierra caliente”), Trinidad, Guyana, Brazil, Peru. Refs.: Enderlein, 1922: 224; Steyskal, 1968: 13 (cat.). mimicus Cresson 1930: 348. Type locality: Peru, Pichís and Perene Valleys. HT F (USNM) Distr.: Colombia (Cueva de Los Guácharos, Cueva del Indio), Costa Rica, Ecuador. Ref.: Steyskal, 1968: 13 (cat.). New record to Colombia. rufifrons Enderlein, 1922: 224. Type locality: Colombia, Cordillera, "tierra caliente". ST F (ZMHB) Distr.: Colombia (Cordillera “tierra caliente”), Ecuador, Panama. Refs.: Hennig, 1934: 37; Hennig, 1937: 523; Steyskal, 1968: 13 (cat.). Probable synonym of C. cyaneiventris. cinctiventris Enderlein, 1922:225. Type locality: Colombia, Cordillera, "tierra caliente". Bolivia, Brazil, Colombia, Costa Rica, Peru, Venezuela. Ptilosphen tetrastigma auct., not Schiner (misidentifications) Genus Rainieria Rondani, 1843 Rainieria Rondani, 1843: 40. Type species, Calobata calceata Fallen (orig. des.). Tanipoda Rondani, 1856: 116. Type species, Calobata calceata Fallen (orig. des.). Tanypoda, emend. Comm.: No Rainieria were listed from Colombia in Steyskal 1968, but the taxon currently treated as the subgenus Rainieriella Aczel certainly occurs in Colombia. Genus Scipopus Enderlein, 1922 Subgenus Phaeopterina Frey, 1927 Phaeopterina Frey, 1927: 73 (as genus). Type species, heteropus Frey (orig, des.). Phaeopterinus, emend. Steyskal, 1968: 15 (cat.). CATALOGUE OF MICROPEZIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 543 sexguttatus Enderlein, 1922: 209. Type locality: Venezuela; Colombia. ST F (ZMHB) Distr.: Colombia, Venezuela. Ref.: Steyskal, 1968: 15 (cat.). Subgenus Scipopus Enderlein, 1922 Scipopus Enderlein, 1922: 208. Type species, Calobata erythrocephala Fabricius (orig. des.). belzebul (Schiner), 1868: 251 (Calobata). Type locality: “South America”. HT F (NHMW) Distr.: Colombia (Boyacá (Muzo)), Panama south to Brazil and Bolivia. Refs.: Enderlein 1922: 210; Hennig, 1934: 324 (rev.); Steyskal, 1968: 15 (cat.). diversus (Schiner), 1868: 250 (Calobata). Type locality : "South America”. HT F (NHMW) Distr.: Colombia (Cordillera, tierra templada), Guyana, Venezuela, Brazil, Peru, Bolivia, Paraguay. Refs.: Enderlein 1922: 211; Hennig, 1934: 326 (rev.); Steyskal, 1968: 16 (cat.). erythrocephalus (Fabricius), 1805: 260 (Calobata). Type locality: “South America”. HT damaged, M or F? (ZMUC) Distr.: Colombia (Bogotá D.C), Panama south to Brazil and Bolivia. Refs.: Enderlein 1922: 210; Steyskal, 1968: 16 (cat.). frit Cresson, 1926: 270. Type locality: Honduras. Sangrelaya. HT M (ANSP). Distr.: Colombia (Isla Gorgona), Mexico to Panama, Venezuela. Refs.: Hennig, 1934: 325 (rev.); Steyskal, 1968: 16 (cat.). gorgonae Hennig, l934: 329. Type locality: Colombia, Isla Gorgona. HT M (BMNH) Distr.: Colombia (Isla Gorgona). Ref.: Hennig, 1934: 329; Steyskal, 1968: 16 (cat.). limbativertex Enderlein, 1922: 211. Type locality: Colombia, Cordillera, "tierra caliente", and Muzo; Brazil, Mato Grosso; Mexico, Chiapas, Soconusco. ST M (ZMHB) (manuscript lectotype M Colombia) Distr.: Mexico to Panama, Colombia (Boyacá (Muzo); Hac. Pehlke; Cordillera “tierra caliente”)), Brazil, Bolivia. Refs.: Hennig, 1934: 328 (rev.); Steyskal, 1968: 16 (cat.). nigripennis (Hendel), 1922 (April) : 231 (Tanypoda). Type locality: Ecuador, Balzapamba. Distr.: Colombia (Cordillera, “tierra caliente”), Ecuador. Refs.: Hennig, 1934: 328 (rev.); Steyskal, 1968: 16 (cat.). Genus Grallomyia Rondani, 1850 Grallomya Rondani, 1850: 180. Type species, Calobata tarsata Wiedemann (orig. des.). Grallomyia, emend. Steyskal, 1968: 16 (cat.). tarsata (Wiedemann), 1830: 541 (Calobata). Type locality: Type series from Brazil and Surinam; the one specimen labeled "type" in the BMNH has no locality data, this specimen is provisionally treated as the holotype. HT M (BMNH) Distr.: Colombia (Cundinamarca (Bogotá D.C); río Magdalena), Mexico to Panama, Trinidad south to Brazil and Bolivia. Refs.: Enderlein, 1922: 218; Hennig, 1934: 93 (rev.); Steyskal, 1968: 19 (cat.). Genus Paragrallomyia Hendel, 1933 Grallomyia, subg. Paragrallomyia Hendel, 1933: 63. Type species, Musca annulata Fabricius (orig, des.). Grallomyia, subg. Neograllomyia, Hendel, 1933:65. Type species, longivitta Hendel (orig. des,) = vittipennis (Coquillett). aeripennis (Enderlein), 1922:214 (Grallomyia). Type locality: Peru, Cuzco, Callanga, Pinipini Valley. HT F (ZMHB) Distr.: Colombia (La Cumbre, probably belong to Valle del Cauca department), Bolivia, Peru. Refs.: Cresson, 1930: 340; Hennig, 1934: 91; Steyskal, 1968: 17 (cat.). 544 · Zootaxa 4122 (1) © 2016 Magnolia Press MARSHALL ET AL. angulata (Loew), 1866: 47 (Cent. 7, no. 87) (Calobata). Type locality : *'New Granada" = Colombia. (a Colombian specimen in ZMHB labelled "angulata det Loew" is probably the type). Distr.: Honduras to Panama, Colombia (Isla Gorgona; Magdalena (Aracataca), Nariño (Tumaco), Hac. Pehlke)), Trinidad, south to Argentina (Tucumán). Refs.: Enderlein, 1922: 216; Cresson, 1930: 351; Hennig, 1934: 80 (rev.); Steyskal, 1968: 17 (cat.). annulata (Fabricius), 1787: 348 (Musca). Type locality: Cayenne=French Guiana. (ZMHB) Distr.: Colombia, French Guiana and Surinam south to Uruguay and Argentina. Refs.: Enderlein, 1922: 215; Steyskal, 1968: 17 (cat.). gorgonae Hennig, 1934 : 90. Type locality : Colombia, Isla Gorgona. HT F (BMNH) Distr.: Colombia (Isla Gorgona). Ref.: Steyskal, 1968: 17 (cat.). lineata (Enderlein), 1922: 217 (Grallomyia). Type locality: “Colombia”. Distr.: HT M (ZMHB) Distr.: Colombia. Refs.: Hennig, 1934: 83 (rev.); Steyskal, 1968: 18 (cat.). platycnema (Loew), 1866: 47 (Calobata). Type locality: New Granada (=Colombia). Distr.: Colombia (Nariño (Tumaco)), Mexico to Panama, West Indies, Ecuador. Refs.: Hennig, 1934: 90 (rev.); Steyskal, 1968: 19 (cat.). thiemei (Enderlein), 1922: 217 (Grallomyia). Type locality: Colombia, Cordilleras “tierra caliente”. HT F (ZMHB) Distr.: Colombia. Refs.: Hennig, 1934: 85 (rev.); Steyskal, 1968: 19 (cat.). Genus Taeniaptera Macquart, 1835 Taeniaptera Macquart, 1835: 491. Type species, trivittata Macquart (mon.). Grallopoda Rondani, 1850:178, 180. Type species, Calobata albimana Macquart (orig. des.) = trivittata Macquart. lasciva (Fabricius), 1798: 564 (Musca). Type locality: Cayenne=French Guiana. HT destroyed, pin only (ZMUC) Distr.: Colombia (Cundinamarca (Fusagasuga)), Southernmost USA, Mexico, West Indies, South to Paraguay and northern Argentina. Refs.: Enderlein, 1922: 212; Hennig, 1934: 87 (rev.); Steyskal, 1968: 18 (cat.). iunulata Wulp, 1897: 372, pl 9. fig. 26 (Calobata). Type locality: Mexico, Mazatlán, Presidio; Frontera and Teapa; North Yucatán; Tabasco; Medellín, near Veracruz (error, this locality is from Colombia (Antioquia department)). vulpes Cresson, 1926: 273. Type locality: Panama, Trinidad Río. HT M (USNM) Distr.: Panama, Colombia (río Magdalena). Refs.: Hennig, 1934: 85 (rev.); Steyskal, 1968: 19 (cat.). TABLE 1. Micropezidae species recorded from Colombia. Subfamily Genus Micropezinae Micropeza Taeniapteriinae Calosphen Cliobata Grallipeza Subgenus Species Author annulata Schiner, 1888 bogotana (Enderlein), 1922 brasiliensis Schiner, 1868 columbiana (Enderlein), 1922 pleuralis (Cresson), 1930 strophium (Enderlein), 1922 biguttata (Enderlein), 1922 diadema (Wiedemann), 1830 affinis Hennig, 1934 ...Continued on next page CATALOGUE OF MICROPEZIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 545 TABLE 1. (Continued) Subfamily Genus Subgenus Species Author ecuadoriensis (Enderlein, 1922) obscura Hennig, 1934 nigrinotata Hennig, 1934 pseudosimplex Hennig, 1934 suavis (Enderlein, 1922) unifasciata (Fabricius, 1805) unimaculata (Macquart, 1846) Grallomyia tarsata (Wiedemann, 1830) Mesoconius albimanus (Enderlein, 1922) auristrigatus Enderlein, 1922 enderleini Frey, 1927 flaviceps Enderlein, 1922 ujhelyianus Enderlein, 1922 gilvimana (Cresson, 1926) vigilans Frey, 1927 obscura (Hennig, 1935) aeripennis (Enderlein, 1922) angulata (Loew, 1866) annulata (Fabricius, 1787) gorgonae Hennig, 1934 lineata (Enderlein, 1922) platycnema (Enderlein, 1922) thiemei (Enderlein, 1922) arthriticus (Wiedemann, 1830) brevipennis (Walker, 1852) harenosus Cresson, 1930 niger (Schiner, 1868) obliqua (Fabricius, 1805) comis Cresson, 1930 conveniens (Wulp, 1897) cyaneiventris (Macquart, 1846) facetus Enderlein, 1922 gentilis Cresson, 1930 insignis (Wiedemann, 1830) mimicus Cresson, 1930 rufifrons Enderlein, 1922 tetrastigma (Schiner, 1868) Phaeopterina sexguttatus Enderlein, 1922 Scipopus belzebul (Schines, 1868) Metasphen Paragrallomyia Plocoscelus Poecilotylus Poecilotylus Ptilosphen Scipopus ...Continued on next page 546 · Zootaxa 4122 (1) © 2016 Magnolia Press MARSHALL ET AL. TABLE 1. (Continued) Subfamily Genus Taeniaptera Zelatractodes Subgenus Species Author diversus (Schiner, 1868) erythrocephalus (Fabricius, 1805) frit Cresson, 1926 gorgonae Hennig, 1934 limbativertex Enderlein, 1922 nigripennis (Hendel, 1922) lasciva (Fabricius, 1798) vulpes Cresson, 1926 garleppi (Enderlein, 1922) rufiventris (Enderlein, 1922) versicolorus (Enderlein, 1922) References Aczel, M.L. (1949) Catálogo de la familia de las “Tylidae” (Calobatidae + Micropezidae + Neriidae, Diptera) en la región Neotropical. Acta Zoologica Lilloana, 8, 309–374. Albuquerque, L.P. (1989) Cardiacephalina subtr. N. de Taeniapterini (Diptera, Micropezidae, Taenipaterinae). Memórias do Instituto Oswaldo Cruz, 84, 19–26. http://dx.doi.org/10.1590/S0074-02761989000800009 Coquillet, D.W. (1910) The type species of the North American genera of Diptera. Proceedings of the Unites States National Museum, 37, 499–647. http://dx.doi.org/10.5479/si.00963801.37-1719.499 Cresson, E.T. (1926) Descriptions of new genera and species of Diptera (Ephydridae and Micropezidae). Transactions of the American Entomological Society, 58, 249–274. Cresson, E.T. (1930) Notes on and descriptions of some neotropical Neriidae and Micropezidae (Diptera). Transactions of the American Entomological Society, 56, 307–362. Enderlein, G. (1922) Klasification der Micropeziden. Archiv für Naturgeschichte, Series A, 88 (85), 140–229. Enderlein, G. (1923) Neue Beiträge zur Kenntnis der Micropeziden (Dipt.). Deutsche Entomologische Zeitschrift, 5, 540–543. Fabricius, J.C. (1787) Mantissa insectorum sistens species nuper detectarum. Vol. 2. Impensis C.G. Proft, Hafniae [= Copenhagen], 382 pp Fabricius, J.C. (1798) Supplementum Entomologia systematica. Proft et Storch, Hafniae [= Copenhagen], 572 pp. Fabricius, J.C. (1805) Systema antiatorum secundum ordines, genera, species adiectis synonymis, locis, observationibus, descriptionibus. C. Reichard, Brunsvigae, 373 pp. Frey, R. (1927) Zur Systematik der Diptera Haplostomata. III. Fam. Micropezidae. Notulae Entomologicae, 7, 65–76. Hendel, F. (1922) Einige neue Micropeziden. Deutsche Entomologische Zeitschrift, 231–235. Hendel, F. (1932) Die Ausbeute der Deutschen Chaco-Expedition 1925/26. - Diptera. (Fortsetzung). XXV–XXVI. Sciomyzidae, Lauxaniidae, Tanypezidae, Lonchaeidae, Tylidae, Drosophilidae, Milichiidae. Konowia, 11, 98–145. Hendel, F. (1933) Ueber einige Typen Wiedemann's und Schiner's von acalyptraten Musciden aus Südamerika, nebst einige verwandten Arten (Dipt.). Revista de Entomologia, 5, 58–83. Hennig, W. (1934) Revision der Tyliden (Dipt., Acalypt.). Stettiner Entomologische Zeitung, 95, 65–198, 294–330. Hennig, W. (1935) Revision der Tyliden (Dipt.,Acalypt.). I. Teil: Die Taeniapterinae Amerikas. Stettiner Entomologische Zeitung, 96, 27–67. Hennig W. (1936) Revision der Tyliden (Dipt., Acalypt.). II. Teil: Die auβeramerikanischen Taeniapterinae, die Trepidariinae und Tylinae. Allgemeines über die Tyliiden. Konowia, 15, 129–144, 201–239. Hennig, W. (1937) Tylidae (Micropezidae, Diptera), aus der Sammlung des British Museum of Natural History. Nachträge zur “Revision der Tyliden-II”. Annals and Magazine of Natural History Series, Series 10, 19, 521–523. http://dx.doi.org/10.1080/00222933708655295 International Commission on Zoological Nomenclature (ICZN) (1955) Opinion 369. Suppression under the plenary powers of the generic names Tylos Meigen, 1800. In: Hemming, F. (Ed.), Opinions and Declatarations rend. Internatioanl Coomission on Zoological Nomenclature, 11, 265–300. Jackson, M.D., Marshall, S.A. &. Skevington, J.H. (2015) Molecular phylogeny of the Taeniapterini (Diptera:Micropezidae) using nuclear and mitochondrial DNA, with a reclassification of the genus Taeniaptera Macquart. Insect Systematics & CATALOGUE OF MICROPEZIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 547 Evolution. [in press] Loew, H. (1866) Diptera Americae septentrionalis indigena. Centuria septima. Berlin Entomologische Zeitschrift, 10, l–54. http://dx.doi.org/10.1002/mmnd.18660100103 Macquart, J. (1835) Historie Naturelle des Insects. Dipteres. Tome deuxieme. Librairie Encyclopédique de Roret, Paris, 703 pp. Macquart, J. (1846) Diptères exotiques nouveaux ou peu connus, 1er. Mémories de la Société Royale des Sciences, de l´ Agriculture et des Arts de Lille, 1844, 133–364, pls. 1–20. [1845) Marshall, S.A. (2004) A review of the genus Metasphen Frey including Rainieria (Globomyia) Hennig new synonym. Studia Dipterologica, 11, 529–536. Marshall, S.A. (2010) Micropezidae Chapter 55. In: Brown, B.V., Borkent, A., Cumming, J.M., Wood, D.M., Woodley, N.E. & Zumbado, M.A. (Eds.), Manual of Central American Diptera. Vol 2. NRC Research Press, Ottawa, pp. 805–813. Marshall, S.A. (2015) Mesoconius Enderlein (Diptera, Micropezidae, Taeniapterinae) of Central America. Zootaxa, 3914 (5), 525–540. http://dx.doi.org/10.11646/zootaxa.3914.5.2 Meigen, J.W. (1800) Nouvelle classification des mouches a deux ailes (Diptera L.) d´apres un plan tout nouveau. J.J. Fuchs, Paris, 40 pp. Meigen, J.W. (1803) Versuch einer neuen Gattungseintheilung der europäischen zweiflügeligen Insekten. Magazin für Insektenkunde, 2, 259–281. Roback, S.S. (1969) The Genera, Subgenera and Species Described by E. T. Cresson, Jr. 1906–1949. Transactions of the American Entomological Society, 95 (4), 517–569. Rondani, C. (1843) Quattro specie di insetti proposti come tipi di generi nouvi. Memoria sesta peer servir ala ditterologia italiana. (Accad. Sci. Ist. Bologna) Nouvi Annali dele Scienze Naturali, 10, 32–46. Rondani, C. (1850) Osservazioni sopra alquante specie di Esapodi Ditteri del Museo Torinese. Nuovi Annali delle Scienze Naturali, Series 3, 2, 165–197, pls. 4. [Bologna] Rondani, C. (1856) Diterologiae Italicae Prodromus. Vol. I. Genera Italica ordinis dipterorum ordinatim disposita et distincta et in famílias et stirpes aggregata. A. Stocchi, Parmae [=Parma], 226 + [2] pp. Schmidt, E. (1906) Beitrag zur Kenntnis der Membraciden. Neue Genera und Spezies in der Sammlung des Stettiner Museums. Stettiner Entomologische Zeitung, 67, 359–373. Schiner, J.R. (1868) Diptera. In: Reise der österreichischen Fregatte Novara um die Erde in den Jahren 1857, 1858, 1859 unter den Befehlen des Commodore B. von Wüllerstorf-Urbair. Vol. 2. Part I. Section B. K. Gerold's Sohn, Wien [=Vienna], pp. i–vi + 1–388, pls. 1–4. Steyskal, G.C. (1968) 48. Family Micropezidae. In: Papavero, N. (Ed.), A Catalogue of the Diptera of the Americas South of the United States. Departamento de Zoología, Secretaria da Agricultura, São Paulo, pp. 1–33. Walker, F. (1852) Diptera. Part IV. In: Saunders, W.W (Ed.), Insecta Saundersiana, London, pp. 253–414. Wiedemann, C.R.W. (1830) Aussereuropäische zweiflügelige Insekten. Vol. 2. n der Schulzischen Buchhandlung, Hamm, xii + 684 pp. Wulp, F.M Van Der. (1897) Group Calobatinae. In: Godman, F.D. & Salvin, O. (Eds.), Biologia Centrali-Americana, Zoology, Insecta, Diptera. Vol. 2. Published for the editors by R. H. Porter, London, pp. 363–376. 548 · Zootaxa 4122 (1) © 2016 Magnolia Press MARSHALL ET AL. Zootaxa 4122 (1): 549–554 http://www.mapress.com/j/zt/ Copyright © 2016 Magnolia Press Article ISSN 1175-5326 (print edition) ZOOTAXA ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.4122.1.41 http://zoobank.org/urn:lsid:zoobank.org:pub:3FCD28CE-54B9-4D3C-8D7B-8CBA8B2FFA9B FAMILY NERIIDAE TATIANA SEPÚLVEDA* & CLAUDIO J. B. DE CARVALHO Departamento de Zoologia, Universidade Federal do Paraná, Postal Box 19020, 81531–980, Curitiba, Paraná, Brazil. E-mail: tatiana.sepulveda.villa@gmail.com; cjbcarva@ufpr.br * Corresponding author: tatiana.sepulveda.villa@gmail.com Abstract Here we provide a catalog of the Neriidae from Colombia, with all six genera and 17 species in alphabetical order. With each genus we provide references and with each species we include the original name, type-locality, acronym of the institution where the type is deposited and the known geographical distribution. In the case of samples only from Colombia, we provide additional details for locality, relevant synonymy and additional references. Key words: Catalog, Colombia, Diptera, distribution, Nerioidea Introduction The Neriidae comprise a small group of acalyptrate flies (116 species in 17 genera, Pape et al. 2011) with rather long legs, porrect antennae with dorso-apical or apical arista and pedicel with a triangular inner process. Neriidae feed on sap, rotten fruit and other decaying matter (Aczél 1961; Mangan & Baldwin 1986; Buck 2010) and are found in all biogeographic regions, tending to predominate in the tropics (Hennig 1937; Steyskal 1968). The Neriidae were first considered a subfamily of Micropezidae in its first revision by Enderlein (1922). In the second general revision, Hennig (1937) listed 34 species in nine genera for the Neotropical region. Neriidae were considered to be within the Tylidae (today, Micropezidae) in the first Neotropical catalog by Aczél (1949). Later, Neriidae was restored as a family in the first complete revision of the American representatives, in which 39 species in 11 genera were recognized (Aczél 1951). The family is split into the subfamilies Telostylinae and Neriinae. Only the Neriinae occur in the Americas and are identified by the expanded and medially divided upper face and lunule, which forms a prominently exposed antennal base (Steyskal 1987; Buck 2010). Steyskal (1968) listed eight species in three genera in Colombia in his catalog of the species reviewed by Aczél (1961). Recently, another four new species have been described from South America (Sepúlveda et al. 2013a, 2014). Today, nine genera are known in the New World (Buck 2010), of which six (and a total of 17 species) have been recorded in Colombia and are included in this catalog, in which we follow the classification proposed by Aczél (1961) and Buck (2010). We list the genera and species in alphabetical order and each species is followed by information including authorship, original publication, original genus name parenthetically, type locality, type-material information (including the acronym of the institution housing the material in square brackets), general distribution (countries) except for the United States, Mexico, Brazil and Argentina, when we also provide state or provincial citations parenthetically. For each Colombian species, we also include the specific locality and department parenthetically, followed by detailed references and the relevant synonymy. At the end of the list of species, a table includes all species known from Colombia, including rank, author(s) and year of publication. Accepted by S. Nihei: 16 Apr. 2015; published: 14 Jun. 2016 549 Acronyms for the depositories CEUA—Colección Entomológica Universidad deAntioquia, Medellin, Colombia. DEBU—University of Guelph Insect Collection, Guelph, Canada. IAvH—Instituto de Investigación de Recursos Biológicos Alexander von Humboldt, Villa de Leyva, Colombia. MZUSP—Museu de Zoologia da Universidade de São Paulo, São Paulo, Brazil. NMW—Naturhistorisches Museum Wien, Wien, Austria. OUM—University Museum of Natural History, Oxford, United Kingdom. SMT—Staatliches Museum für Tierkunde, Dresden, Germany. ZMHB—Museum für Naturkunde der Humboldt-Universität, Berlin, Germany. USNM—National Museum of Natural History, Smithsonian Institution, Washington, D. C. ZMHB—Museum für Naturkunde der Humboldt-Universität, Berlin, Germany. ZMUC—University of Copenhagen, Zoological Museum, Copenhagen, Denmark. List of abbreviations cat.—catalogue distr.—distribution fig./figs—figure/figures HT—Holotype LT—Lectotype misident.—misidentification mon.—by monotypy n. comb.—new combination n. syn.—new synonym orig. des.—by original designation PNN—National Natural Park pp.—pages PT—Paratype redesc.—redescription Refs.—references RN—Natural Reserve SFF—Flora and Fauna Sanctuary SN—Syntype subs. des.—by subsequent designation M (m#)—Male F (f#)—Female Catalogue of Neriidae of Colombia Family Neriidae Westwood, 1840 Subfamily Neriinae Enderlein, 1922 Genus Cerantichir Enderlein, 1922 Cerantichir Enderlein, 1922: 155. Type species, Nerius flavifrons Bigot, 1886 (orig. des.; misident.) = enderleini Hennig, 1937. peruana (Hennig), 1937: 261 (Odontoloxozus). Type locality: Peru, Urubamba river, Umuhuankiali. HT M (SMT). Distr.: Costa Rica to Bolivian and Midwest Region of Brazil, Colombia (Putumayo (PNN La Paya)). Refs.: 550 · Zootaxa 4122 (1) © 2016 Magnolia Press SEPÚLVEDA & DE CARVALHO Sepúlveda et al. 2013b: 126 (key; figs. 1B, 1D, 2B, 2D), 127 (figs. 3B, 3C, 3E, 3I, 3J, 3K), 129 (redesc.; figs. 5A, 5B), 130 (distr.). Genus Eoneria Aczél, 1951 Eoneria Aczél, 1951: 570. Type species, Eoneria blanchardi Aczél, 1951 (orig. des.). aczeli Sepúlveda & Carvalho, 2013a: 252. Type locality: Colombia, Magdalena, PNN Tayrona, Neguanje. HT M (IAvH). Distr.: Colombia (Magdalena (PNN Tayrona, Neguanje), Atlántico (Barranquilla, Uniatlántico)). Genus Glyphidops Enderlein, 1922 Glyphidops Enderlein, 1922: 150. Type species, Nerius filosus Fabricius, 1805 (orig. des.). Subgenus Glyphidops Enderlein, 1922 bullatus (Enderlein), 1922: 151 (Chaetomeristes). Type locality: Ecuador, Napo Province, Archidonia. HT M (ZMHB). Distr.: Colombia (Risaralda (SFF Otún Quimbaya), Huila (PNN Cueva de Los Guácharos), Nariño (RN La Planada)), Ecuador, Peru. Refs.: Sepúlveda et al., 2014: 142 (key), 142 (redesc.), 144 (figs. 1A-G), 169 (distr.). coracinus Sepúlveda, Wolff & Carvalho, 2014: 145. Type locality: Colombia, Risaralda, SFF Otún Quimbaya. HT M (IAvH). Distr.: Colombia (Risaralda (SFF Otún Quimbaya), Putumayo (PNN La Paya, Loma Baja), Amazonas (PNN Amacayacu, San Martin)), Peru, Bolivia. filosus (Fabricius), 1805: 262 (Nerius). Type locality: “America Meridional”. HT sex? (ZMUC). Distr.: Costa Rica, Guatemala, Honduras, Panamá, Trinidad, Surinam, British Guyana, Brazil (Amazonas, Roraima, Pará, Acre, Alagoas, Bahia, Minas Gerais, Espírito Santo, Rio de Janeiro, Mato Grosso, São Paulo), Colombia (Antioquia (Santa Bárbara; La Pintada; Jericó, Cauca Viejo), Meta (PNN La Macarena, Caño Curía), Vaupés (Estación Biólogica Mosiro-Itajura, Caparú)), Ecuador, Peru, Bolivia. Refs.: Sepúlveda et al., 2014: 142 (key), 148 (redesc.), 149 (figs. 4A-E), 169 (distr.). ruselatus Sepúlveda, Wolff & Carvalho, 2014: 152. Type locality: Colombia, Antioquia, La Ceja. HT M (CEUA). Distr.: Colombia (Antioquia (La Ceja; Medellín; Andes; Carepa), Norte de Santander (Cucutilla), Santander (Carmen de Chucurrí; Pie de Cuesta), Risaralda (SFF Otún Quimbaya), Quindío (Finlandia), Huila (PNN Cueva de Los Guacharos), Caldas (Anselma), Valle del Cauca (PNN Farallones de Cali)), Ecuador. steyskali Sepúlveda, Wolff & Carvalho, 2014: 154. Type locality: Ecuador, Puerto Orellana. HT M (USNM). Distr.: Colombia (Caquetá (Sucre), Huila (PNN Cueva de los Guacharos), Meta (PNN Sumapaz), Vaupés (Estación Biológica Moshiro-Itajura (Caparú)), Putumayo (PNN La Paya), Amazonas (PNN Amacayacu, Matamata)), Ecuador, Brazil (Amazonas), Peru. Subgenus Oncopsia Enderlein, 1922 Dictyonerius Enderlein, 1922: 159. Type species: Nerius pluricellatus Schiner, 1868: 248 (orig. des.). Refs.: Aczél 1961: 329. carrerai Aczél, 1961: 316. Type locality: Brazil, São Paulo, São Vicente. HT M (MZUSP). Distr.: Colombia (Antioquia (Girardota), Amazonas (PNN Amacayacu, Matamata)), Brazil (São Paulo), Guyana. Refs.: Sepúlveda et al. 2014: 142 (key), 159 (redesc.), 160 (figs. 9A–D), 171 (distr.). durus (Cresson), 1926: 260 (Nerius). Type locality: Costa Rica, Higuito, San Mateo. HT F (USNM). Distr.: Costa Rica, Panama, Colombia (Antioquia (Nariño, Vereda Puente Linda; San Luis, RN Cañón del Rio Claro), Chocó (PNN Los Katíos), Risaralda (Otún Quimbaya, El Molinillo), Huila (PNN Cueva de los Guácharos, Cabaña CATALOGUE OF NERIIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 551 Cedros), Cauca (PNN Gorgona, Antigua Laguna), Valle del Cauca (PNN Farallones de Cali, Anchicaya; Ginebra, Loma Gorda), Meta (PNN Sumapaz, Jardín Botánico), Putumayo (PNN La Paya, Loma Alta), Vaupés (Estación Biológica Mosiro Itajura, Caparú), Amazonas (PNN Amacayacu, Matamata; PNN Amacayacu, San Martín)), Venezuela, Brazil (Amazonas, Pará, São Paulo, Rio de Janeiro, Paraná), Ecuador, Peru, Bolivia, Argentina (Tucumán). Refs.: Sepúlveda et al., 2014: 142 (key), 160 (redesc.), 161 (figs. 10A–E, 17F–I), 171 (distr.). flavifrons (Bigot), 1886: 372 (Nerius). Type locality: Mexico. HT F (OUM). Distr.: USA (Arizona), Mexico (Baja California sur, Tabasco), Honduras, Nicaragua, El Salvador, Costa Rica, Panama, Colombia (Magdalena (Sierra Leona, Hacienda Cincinnati), Antioquia (Carepa, Tunelapa; La Pintada; Maceo, RN Cañón del rio Alicante; San Luis, Rio Samaná Norte; Nariño, Vereda Puente Linda; Santa Bárbara), Caldas (Norcasia), Chocó (Nuquí, Corregimiento Jobi), Meta (PNN Sumapaz, Jardín Botánico), Putumayo (PNN La Paya, Loma Alta), Amazonas (PNN Amacayacu, Matamata; PNN Amacayacu, San Martín)), Trinidad and Tobago, British Guyana, Brazil (Amazonas, Pará, Acre), Ecuador, Bolivia. Refs.: Hennig 1937: 252 (key, listed, n. com.); Aczél 1949: 378 (cat.); Steyskal 1968: 3 (cat.); Sepúlveda et al. 2014: 142 (key), 163 (redesc.), 164 (figs. 11A–E, 17J–L)), 171 (distr.). pluricellatus (Schiner), 1868: 248 (Nerius). Type locality: “South America” (Venezuela). ST F (NMW). Distr.: Costa Rica, Panama, Colombia (Antioquia (Carepa, Tunelapa; La Pintada; Rio Claro; Zaragoza, Providencia; Jericó, Cauca Viejo), Chocó (Jobi; PNN Los Katíos), Meta (Villavicencio, Quatquia)), Venezuela, Argentina (Mendoza). Refs.: Hendel 1933: 59 (redescr.); Hennig 1937: 262 (redescr.); Aczél 1961: 315 (key), 329 (redesc., n. comb.), 335 (fig. 59); Steyskal 1968: 3 (cat.); Sepúlveda et al., 2014: 142 (key), 167 (redesc.), 168 (figs. 13A–E,), 171 (distr.). Genus Longina Wiedemann, 1830 Longina Wiedemann, 1830: 553. Type species: Longina abdominalis Wiedemann, 1830 (mon.). anguliceps Buck & Marshall, 2004: 25. Type locality: Ecuador, Provincia Napo, RN Sierra Azul. HT F (DEBU). Distr.: Colombia (Norte de Santander (Cucutilla), Santander (Pie de Cuesta), Risaralda (SFF Otún Quimbaya), Boyacá (SFF Iguaqué), Quindío (Finlandia), Cauca (PNN Munchique, Sector La Romelia), Huila (PNN Cueva de los Guácharos)), Ecuador. semialba Buck & Marshall, 2004: 28. Type locality: Ecuador, Provincia Napo, Cosanga. HT F (DEBU). Distr.: Colombia (Risaralda (SFF Otún Quimbaya), Quindío (SFF Otún Quimbaya), Huila (PNN Cueva de Los Guácharos), Nariño (RN La Planada)), Ecuador. Genus Loxozus Enderlein, 1922 Loxozus Enderlein, 1922: 156. Type species: Loxozus clavicornis Enderlein (orig. des.). clavicornis Enderlein, 1922: 156. Type locality: Colombia, “cordilleras, tierra templada”. HT F (ZMHB). Distr.: Colombia. Genus Nerius Fabricius, 1805 Nerius Fabricius, 1805: 264. Type species: Nerius pilifer Fabricius, 1805: 264 (post. des. Coquillett 1910: 575). pilifer Fabricius, 1805: 264. Type locality: “America meridional”. HT M (ZMUC). Distr.: Mexico (Tabasco), Haiti, Costa Rica, Panama, Colombia (La Guajira (Rioacha, Corregimiento El Abra), Atlántico (Barranquilla, Uniatlántico), Magdalena (Santa Marta), Bolivar (Corregimiento El Arenal), Sucre (San Onofre, RN Sanguaré), Antioquia (Maceo, RN Cañón del Rio Alicante; La Pintada; Jericó, Cauca Viejo; Nariño, Vereda Puente Linda), 552 · Zootaxa 4122 (1) © 2016 Magnolia Press SEPÚLVEDA & DE CARVALHO Santander (Campamento, Santa Isabel), Caldas (Norcasia, Sector El Tigre), Vichada (PNN El Tuparro, Ceroo Tomás), Putumayo (PNN La Paya, Quebrada Blanca)), Venezuela, British Guyana, Surinam, Brazil (Amazonas, Pará, Acre, Tocantins, Rôndonia, Bahia, Mato Grosso, Goiás, Minas Gerais, Mato Grosso do Sul, São Paulo), Ecuador, Peru, Bolivia, Paraguay, Argentina (Tucumán). Refs.: Macquart, 1835: 493 (as brunneus, desc.); Hennig 1937: 246 (remarks); Aczél 1949: 376 (cat.); Aczél 1961: 333 (key), 344 (remarks); Steyskal 1968: 4 (cat.); Carvalho-Filho & Esposito 2008: 60 (distr., key). brunneus Macquart, 1835: 493; Hennig 1937: 246. plurivittatus Bigot, 1886: 372. Type locality: Mexico. HT M (OUM). Distr.: Mexico (Veracruz, Córdoba), Panama, Dominican Republic, Trinidad, British Guyana, Venezuela, Colombia (Caldas (Norcasia), Antioquia (Maceo, RN Cañón del Rio Alicante; San Roque, Corregimiento San José del Nus; La Pintada; Nariño, Vereda Puente Linda; Carepa, Tunelapa), Cundinamarca (PNN Sumapaz), Meta (PNN Sumapaz, Jardín Botánico; Villavicencio, Rio Oca), Valle del Cauca (PNN Farallones de Cali, Anchicaya), Putumayo (PNN La Paya), Vaupés (Estación Biológica Mosiro-Itajura, Caparú), Amazonas (PNN Amacayacu, Matamata)), Ecuador, Peru, Bolivia, Brazil (Amazonas, Pará, Acre). Refs.: Aczél 1961: 262 (listed), 336 (redesc.), 339 (distr.); Steyskal 1968: 5 (cat.); Carvalho-Filho & Esposito 2008: 60 (listed, distr.). purpusianus (Enderlein), 1922: 157 (Brachantichir). Type locality: Mexico, Xiapas, Soconusco. LT M (ZMHB). Distr.: Mexico (Chiapas), Costa Rica, Colombia (Caldas (Tolima, Hacienda Pehlke)). Refs.: Hennig 1937: 247 (redesc.); Aczél 1949: 376 (cat.); Steyskal 1968: 5 (cat.). Acknowledgments The authors would like to thank to Conselho Nacional de Desenvolvimento Científico e Tecnológico (CNPq) for the research grant (TAS proc. 130370/2011-8; CJBC proc. 304713/2011–2). Thanks to James J. Roper for review of the English. TABLE 1. Species of Neriidae recorded for Colombia. Genus Species Author Cerantichir peruana (Hennig, 1937) Eoneria aczeli Sepúlveda & Carvalho, 2013 bullatus (Enderlein, 1922) coracinus Sepúlveda, Wolff & Carvalho, 2014 filosus (Fabricius, 1805) ruselatus Sepúlveda, Wolff & Carvalho, 2014 steyskali Sepúlveda, Wolff & Carvalho, 2014 carrerai Aczél, 1961 durus (Cresson, 1926) flavifrons (Bigot, 1886) pluricellatus (Schiner, 1868) anguliceps Buck & Marshall, 2004 semialba Buck & Marshall, 2004 Loxozus clavicornis Enderlein, 1922 Nerius pilifer Fabricius, 1805 plurivittatus Bigot, 1886 purpusianus (Enderlein, 1922) Glyphidops Glyphidops Subgenus Glyphidops Oncopsia Longina CATALOGUE OF NERIIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 553 References Aczél, M.L. (1949) Catálogo de la familia de las Tylidae (Calobatidae + Micropezidae + Neriidae, Diptera) en la Region Neotropical. Acta Zoológica Lilloana, 8, 309–389. Aczél, M.L. (1951) Morfología externa y División Sistemática de las “Tanypezidiformes” con sinopsis de las especies argentinas de “Tylidae” (“Micropezidae”) y “Neriidae” (Dipt.). Acta Zoologica Lilloana, 11, 483–589. Aczél, M.L. (1961) A revision of American Neriidae (Diptera, Acalyptratae). Studia Entomologica, 4, 257–346. Bigot, J.M.F. (1886) Diptères nouveaux ou peu connus. 29e partie (suite) (1). XXXVII. §2e. Essai D´une classification synoptique du groupe Tanypezidi (mihi) et description de genres et d'espèces inédits. Annales de la Société Entomologique de France, 6, 372–373. Buck, M. (2010) Neriidae. In: Brown, B.V., Borkent, A., Cumming, J.M., Wood, D.M., Woodley, N.E. & Zumbado, M.A. (Eds.), Manual of Central American Diptera. Vol. 2. NRC Research Press, Ottawa, pp. 815–819. Buck, M. & Marshall, S.A. (2004) A review of the genus Longina Wiedemann, with descriptions of two new species (Diptera, Neriidae). Studia Dipterologica, 11, 23–32. Carvalho-Filho, F.S. & Esposito, M.C. (2008) Neriidae (Diptera: Schizophora) of the Brazilian Amazon: new records of genera and species, and key to species. Neotropical Entomology, 37, 58–62. http://dx.doi.org/10.1590/S1519-566X2008000100008 Coquillett, D.W. (1910) The type-species of the North American genera of Diptera. Proceedings of the United States National Museum, 6, 166–192. http://dx.doi.org/10.5962/bhl.title.57938 Cresson, E.T. Jr. (1926) Descriptions of new genera and species of Diptera (Ephydridae and Micropezidae). Transactions of the American Entomological Society, 52, 249–274. Enderlein, G. (1922) Klassifikation der Micropeziden. Archiv für Naturgeschicthe, (A) 88, 140–229. Fabricius, J.C. (1805) Systema antliatorum secundum ordines, genera, species, adiectis synonyms, locis, observationibus, descriptionibus. Reichard, Brunsvigae (=Brunswick), pp. 264–265. Hendel, F. (1933) Ueber einige Typen Wiedemann´s und Schiner´s von acalyptraten Musciden aus Südamerika, nebst eiginen verwandten Arten. Revista de Entomologia, 3, 50–83. Hennig, W. (1937) Üebersichtüeber die Arten der Neriiden und üeber die Zoogeographiedieser Acalyptraten-Gruppe. Stettiner Entomologische Zeitung, 98, 240–280. Macquart, P.J.M. (1835) Histoire naturelle des insectes. Diptères. Tome deuxième. Ouvrage accompagné de planches. Roret, Paris, 703 pp. Mangan, R.L. & Baldwin, D. (1986) A new cryptic species of Odontoloxozus (Neriidae: Diptera) from the Cape Region of Baja California Sur (Mexico). Proceedings of the Entomological Society of Washington, 88, 110–121. Pape, T., Blagoderov, V. & Mostovski, B.M. (2011) Order Diptera Linnaeus, 1758. In: Zhang, Z.-Q. (Ed.), Animal biodiversity: An outline of higher-level classification and survey of taxonomic richness. Zootaxa, 3148, 222–229. Sepúlveda, T., Wolff, M.I. & Carvalho, C.J.B. (2013a) Revision of the Neotropical genus Eoneria Aczél (Diptera: Neriidae) with description of a new species from Colombia. Zootaxa, 3636 (2), 245–256. http://dx.doi.org/10.11646/zootaxa.3636.2.2 Sepúlveda, T., Wolff, M.I. & Carvalho, C.J.B. (2013b) Revision of the neotropical genus Cerantichir (Diptera: Neriidae) with new records and a key to species. Revista Colombiana de Entomología, 39 (1), 125–131. Sepúlveda, T., Wolff, M.I. & Carvalho, C.J.B. (2014) Revision of the New World genus Glyphidops Enderlein (Diptera: Neriidae). Zootaxa, 3785 (2), 139–174. Schiner, I.R. (1868) Diptera. In: Wullerstorf-Urbair, B. von (Ed.), Reise der österreichischen Fregatte Novara um die Erde in den Jahren 1857, 1858, 1859 unter den Befehlen des Commodore B. von Wüllerstorf-Urbair. Zoologischer Theil, Vol. 2 (Abt. 1). B. K. Gerald's Sohn, Wien, pp. 1–388. [vi + 388 pp., 4 pls.] Steyskal, G.C. (1968) 49. Family Neriidae. In: Papavero, N. (Ed.), A Catalogue of Diptera of the Americas South of the United States. Departamento de Zoologia, Secretaria da Agricultura, São Paulo, 7 pp. Steyskal, G.C. (1987) Neriidae. In: McAlpine, J.F., Peterson, B.V., Shewell, G.E., Teskey, H.J., Vockroth, J.R. & Wood, D.M. (Eds.), Manual of Neartic Diptera. Vol. 2. Research Branch Monograph 28, Agriculture Canada, Ottawa, Ontario, pp. 769–771. Westwood, J.O. (1840) An Introduction to the Modern Classification of Insects; founded on the Natural Habits and Corresponding Organisation of the different Families. Volume II. Longman, Orme, Brown, Green, and Longmans, London, xi + 587 pp. Wiedemann, C.R.W. (1830) Aussereuropische zweiflgelige Insekten Als Fortsetzung des Meigenschen Werkes. Erster Theil. Schulz, Hamm, xii + 684 pp., 5 pls. 554 · Zootaxa 4122 (1) © 2016 Magnolia Press SEPÚLVEDA & DE CARVALHO Zootaxa 4122 (1): 555–557 http://www.mapress.com/j/zt/ Copyright © 2016 Magnolia Press Article ISSN 1175-5326 (print edition) ZOOTAXA ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.4122.1.42 http://zoobank.org/urn:lsid:zoobank.org:pub:A525554E-4435-4938-92B1-4F8F177F6A08 FAMILY TANYPEZIDAE MARTA WOLFF*1 & DIANA GRISALES2 1 Grupo de Entomologia, Instituto de Biologia, Universidad de Antioquia. Colombia. Laboratório de Biodiversidade e Biogeografia de Diptera, Departmento de Zoologia, UFPR, Caixa postal 19020, Curitiba, PR., Brazil, 81531-980. E-mail: martha.wolff@udea.edu.co; ochoa310@gmail.com * Corresponding author: martha.wolff@udea.edu.co 2 Abstract The Tanypezidae of Colombia are catalogued. Nine species in one genus are presented and geographical information are provided for all species, when available. Key words: catalogue, Diptera, distribution, Colombia Introduction Tanypezidae is a small family of Acalyptratae Diptera. These flies are medium sized with body slender, long legs and veins R4+5 and M converging apically. The biology and larval stages are practically unknown (Lonsdale 2014). Only 28 species are known in two genera, Tanypeza Fallén (2 species, restricted to the Holarctic region) and Neotanypeza Hendel (26 species, restricted to the New World) (Lonsdale 2014). In Colombia only nine species are reported in this genus but it is expected to discover many new species and new occurrences. This catalogue is based on the Neotropical Catalogue of Tanypezidae (Steyskal 1967) and on the World Catalogue of the family (Lonsdale 2014). We underline Colombian department names and parenthetically note specific localities. At the end, we provide a table that includes all species found in Colombia (Table 1). Acronyms used for the depositories CBFC—Colección Boliviana de Fauna, La Paz, Bolivia. CAS—California Academy of Sciences, San Francisco, USA. DEBU—University of Guelph Insect Collection, Guelph, Ontario, Canada. MRSN—Museo Regionale de Scienze Naturali, Turin, Italy. NMW—Naturhistorisches Museum Wien, Vienna, Austria. USNM—National Museum of Natural History, Washington, D.C., USA. ZMHB—Museum für Naturkunde der Humboldt-Universität, Berlin, Germany. List of abbreviations cat.—catalogue des.—designation distr.—geographic distribution m.—meters Accepted by S. Nihei: 26 May 2015; published: 14 Jun. 2016 555 orig. des.—by original designation pp.—pages redescr.—redescription refs.—references rev.—revision stat. reinst.—status reinstated subs.—subsequent T—type LT—lectotype F (f#)—Female M (m#)—Male Catalogue of Tanypezidae of Colombia Family Tanypezidae Bigot, 1880 Genus Neotanypeza Hendel Neotanypeza Hendel, 1903: 203 (as subgenus of Tanypeza). Type species, Tanypeza elegans Wiedemann (subs. des., Enderlein, 1913: 225). Refs.: Steyskal, 1967: 1 (cat.); 1987: 774; Lonsdale, 2013: 11 (rev., phylogeny); 2014: 414 (cat.). Scipopeza Enderlein, 1913: 228. Type species, Scipopeza grandis Enderlein (orig. des.). Polphopeza Enderlein, 1913: 226. Type species, Polphopeza quadrisetosa Enderlein (orig. des.). Tripolphopeza Enderlein, 1936: 41. Type species, Polphopeza elegantina Enderlein (orig. des.). Tritanypeza Enderlein, 1936: 42. Type species, Tritanypeza cubitofusca Enderlein (orig. des.). alopecia Lonsdale, 2013: 16. Type locality: Ecuador, Napo, Res. Ethnica Waorani, 1 km S Onkone Gare Camp, 00°3910”S, 76°26’W. HT M (USNM). Distr.: Colombia (Cundinamarca, Finca Bellavista, Sasaima), Bolivia, Brazil, Ecuador, Paraguay, Peru. Refs.: Lonsdale, 2014: 415 (cat.). callitarsis (Rondani), 1850: 181. Type locality: Venezuela, Junin Prov., La Merced, Chanchamayo, 800m. HT M (MRSN) (redescr. Lonsdale, 2013: 19). Distr.: Colombia (Cundinamarca, Finca Bellavista, Sasaima), Argentina, Bolivia, Brazil, Costa Rica, Ecuador, Guatemala, Mexico, Paraguay, Peru, Venezuela. Refs.: Lonsdale, 2013: 19; 2014: 415 (cat.). nigripalpis Hennig, 1936: 36. cubitofusca Enderlein, 1936: 44 (Tritanipeza). claripennis (Schiner), 1868: 247 (stat. reinst. by Lonsdale, 2013: 20). Type locality: Venezuela. LT M (NMW) (des. Lonsdale 2013: 20). Distr.: Argentina, Brazil, British Honduras, Colombia, Costa Rica, Dominica, Ecuador, El Salvador, Guatemala, Mexico, Panama, Peru, St. Vincent, Venezuela. Refs.: Lonsdale, 2013: 20; 2014: 415 (cat.). dallasi (Shannon), 1927: 41. Type locality: Argentina, Tucuman, quebrada de Lules. LT M (USNM) (des. Lonsdale 2013: 21). Distr.: Argentina, Bolivia, Colombia, Costa Rica, Dominica, El Salvador, Guatemala, Mexico, Panama. Refs.: Lonsdale, 2013: 21; 2014: 415 (cat.). flavicalx (Enderlein), 1936: 46. Type locality: Colombia, Cordillere, Tierra Caliente. HT F (ZMHB). Distr.: Bolivia, Brazil, Colombia, Ecuador, Panama, St. Lucia, Trinidad, Venezuela. Refs.: Steyskal, 1967: 2; Lonsdale, 2013: 25; 2014: 416 (cat.). montana (Enderlein), 1936: 40 (Polphopeza). Type locality: Colombia. Cordilleren, Terra Fria. LT F (ZMHB) (redes. Lonsdale 2013: 27). Distr.: Colombia (Meta, 3 m W of Villavicencio, 920 m), Argentina, Bolivia, Brazil, Costa Rica, Ecuador, Panama, Peru, Venezuela. Refs.: Steyskal, 1967: 2; Lonsdale, 2013: 27; 2014: 416 (cat.). 556 · Zootaxa 4122 (1) © 2016 Magnolia Press WOLFF & GRISALES posthos Lonsdale, 2013: 31. Type locality: Bolivia, La Paz, Coroico, Cerro Uchumachi, 2550 m, 16°12′43′′S, 67°42′49′′W, cloud forest. HT M (CBFC). Distr.: Bolivia, Colombia (Antioquia, Queb. La Iguana, 17 km NW Medellin), Ecuador, Peru, Venezuela. Refs.: Lonsdale, 2013: 31; 2014: 417 (cat.). symmetros Lonsdale, 2013: 34. Type locality: Colombia, Las Hermosas National Park, Valle Paramo, 2700 m. HT M (CAS). Distr.: Colombia (Valle del Cauca, Páramo Las Hermosas). Refs.: Lonsdale, 2014: 418 (cat.). vexilla Lonsdale, 2013: 34. Type locality: Ecuador, Napo, Sierra Azul Res., 14 km W of Cosanga, 2200 m, 0°40′55′′S, 77°58′09′′W, HT M (DEBU). Distr.: Ecuador, Colombia (Boyacá, SFF Iguaque, 5°25′12′′N, 73°27′24′′W, 2850 m), Peru, Venezuela. Refs.: Lonsdale, 2013: 31; 2014: 418 (cat.). Acknowledgements To Estrategia de Sostenibilidad 2015-2015 University of Antioquia for the grants to MW and to the Conselho Nacional de Desenvolvimento Científico e Tecnológico for their postdoctoral support to DG. TABLE 1. Tanypezidae species recorded to Colombia. Genus Species Author Neotanypeza alopecia Lonsdale, 2013 callitarsis (Rondani, 1850) claripennis (Schiner, 1868) dallasi (Shannon, 1927) flavicalx (Enderlein, 1936) montana (Enderlein, 1936) posthos Lonsdale, 2013 symmetros Lonsdale, 2013 vexilla Lonsdale, 2013 References Bigot, J.M.F. (1880) Dipteres nouveaux ou peu connus. 12e partie. XVIII. Genres Plagiocera (Macq.), Formosia (Guerin) et Rutilia (Rob.-Desv.). XIX. Diopsidae (Macq., J. Bigot). Annales de la Société Entomologique de France, 5, 85–89, 90–94. Enderlein, G. (1913) Dipterologische Studien II. Zur Kenntnis der Tanypezinen. Zoologischer Anzeiger, 42, 224–229. Enderlein, G. (1936) Klassifikation der Tanypeziden. Deutsche Entomologische Zeitschrift, 1936, 39–47. Hendel, F. (1903) Ueber die systematische Stellung von Tanypeza Fall. (Dipt.). Weiner Entomologische Zeitung, 22, 201–205. Hennig, W. (1936) Beiträge zur Kenntnis des Kopulationsapparates und der Systematik der Tanypeziden (Dipt., Acalypt.). Deutsche Entomologische Zeitschrift, 1936, 27–38. Lonsdale, O. (2013) Review of the families Tanypezidae and Strongylophthalmyiidae, with a revision of Neotanypeza Hendel (Diptera: Schizophora). Smithsonian Contributions to Zoology, 641, 1–60. http://dx.doi.org/10.5479/si.19436696.641.1 Lonsdale, O. (2014) World Catalogue of the family Tanypezidae (Diptera: Schizophora). Zootaxa, 3857 (3), 412–422. http://dx.doi.org/10.11646/zootaxa.3857.3.4 Rondani, C. (1850) Osservazione sopra alquante specie di esapodi ditteri del Museo Torinese. Nuovi annali delle scienze naturali, Bologna, Series 2, 165–197. Schiner, I.R. (1868) Diptera. In: von Wullerstorf-Urbair, B. (Ed.), Reise der österreichischen Fregatte Novara. Zoology, 2 (1), pp. i–vi+ 1–388, pls. 1–4. [B.K. Gerold’s Sohn, Wien] http://dx.doi.org/10.5962/bhl.title.7913 Steyskal, G.C. (1967) 52. Family Tanypezidae. In: A catalogue of the Diptera of the Americas South of the United States. Museu de Zoologia. Universidad de São Paulo, São Paulo, 4 pp. CATALOGUE OF TANYPEZIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 557 Zootaxa 4122 (1): 558–560 http://www.mapress.com/j/zt/ Copyright © 2016 Magnolia Press Article ISSN 1175-5326 (print edition) ZOOTAXA ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.4122.1.43 http://zoobank.org/urn:lsid:zoobank.org:pub:3A6AC4C8-D630-435F-A80B-4A78DFCAC4D7 FAMILY SYRINGOGASTRIDAE ALESSANDRA RUNG Insect Biosystematist Plant Pest Diagnostics Branch, CDFA, 3294 Meadowview Road, Sacramento, CA 95832-1448, U.S.A. E-mail: arung@cdfa.ca.gov Abstract This catalog features the three species of Syringogastridae that have been reported for Colombia. It is entirely based on literature data and contains no new records. Key words: catalogue, Colombia, Distribution, Syringogaster , Syringogastridae Introduction The Syringogastridae occur in the Neotropical region with exception of the Antilles, from which they are known only through the fossil record (Dominican amber) (Marshall et al. 2009). They are distributed from Tamazunchale, San Luis Potosí, Mexico to Eldorado, Misiones, Argentina (Marshall et al. 2009). The family is monotypic, with a total of 21 extant species, three of which occur in Colombia (Table 1). Almost nothing is known about the biology of Syringogaster. Syringogastrids walking on leaves resemble ants (especially Pseudomyrmex Lund) that often occur on the same leaf surfaces (Papavero 1964). This resemblance includes movement, size and shape, making syringogastrids difficult to distinguish from ants until they take flight. Marshall et al. (2009) and Marshall & Buck (2010) summarized what is known about the biology of syringogastrids. Syringogaster adults are characterized by the following combination of characters (Marshall & Buck 2009): Small (body length 4–6 mm), ant-looking flies; head spherical, separated from mesothorax by long, collar-like prothorax; setae reduced; head setae limited to outer verticals and (usually) ocellar bristles; thorax only with postalar and one pair of scutellar bristles; pedicel with dorsal seam, arista bipectinate. Wing partially infuscated or banded, constricted basally, anal portion of wing reduced, alula apparently absent; Costal vein without breaks; subcostal vein incomplete, more or less coalescing with vein R1 Costal vein; cell cup complete. Forelegs short and separated from midlegs by wide distance; hind femur swollen and with anteroventral and posteroventral row of stout, short, spine-like setulae. Scutum with distinct supra-alar carina, metathorax with large, cylindrical posterior extension, postmetacoxal bridge high. Abdomen petiolate, with first three or four segments fused. Male with fully developed and symmetrical tergite 6. Synsternite 7+8 forming complete ring separate from epandrium; surstyli and cerci present. Hypandrium with prominent, setose, ventrolateral lobes; hypandrial bridge usually developed; distiphallus large and complex, usually curled up on left side of postabdomen. Female lacking oviscape; apical segments forming short telescoping ovipositor. The larvae are not known. Feijen (1989) and Meyer & Hilger (2000) gave information on egg morphology. Species of Syringogaster are somewhat similar to some other ant mimics, especially small Micropezidae in the genera Metasphen Frey and Globopeza Marshall (Marshall & Buck 2010). Syringogastrids are easily distinguished from Psilidae (the family in which they were originally described by Cresson 1912), and most other Neotropical flies, by the long collar-like pronotum, the hind femur swollen with ventral row of spines, the reduced head chaetotaxy, the basally narrowed wing, and the ant-like body shape (Marshall & Buck 2009). 558 Accepted by S. Nihei: 10 Apr. 2015; published: 14 Jun. 2016 Acronyms used for the depositories ANSP —Academy of Natural Sciences, Philadelphia, Pennsylvania, U.S.A. DEBU—University of Guelph Insect Collection, Guelph, Ontario, Canada. List of abbreviations cat. —catalog distr.—geographic distribution orig. des.—by original designation refs.—references rev.—revision HT—Holotype F (f#)—Female M (m#)—Male Catalogue of Syringogastridae of Colombia Family Syringogastridae Cresson, 1912 Genus Syringogaster Cresson Syringogaster Cresson, 1912: 392 (as Psilidae). Type species, Syringogaster rufa Cresson (orig. des.). Refs.: Prado, 1975: 1 (cat.); Prado and Papavero, 2002: 1 (checklist, Amazon); Marshall et al. 2009: 1–80 (key, rev.); Marshall and Buck 2010: 843–846 (Manual of Central American Diptera). brunnea Cresson, 1912: 394. Type locality: Costa Rica. Peralta Sta. HT F (ANSP). Distr.: Colombia (Valle del Cauca (Río Raposo)), Costa Rica, Ecuador, Guatemala, Panama. Refs.: Curran, 1934: 373; Hennig, 1958: 542, fig. 127c; Marshall et al., 2009: 16, 28–29 (key, rev., figs. 17–20); Marshall and Buck 2010:843 fig. 61. 1, 845 (Manual of Central American Diptera, discussion). brunneina Marshall & Buck, 2009 in Marshall et al., 2009: 30. Refs.: Marshall and Buck 2010: 844, 845, figs. 61.2–3 (Manual of Central American Diptera). Type locality: Trinidad. Arima, Windblow Ridge, Simla House. HT M (DEBU). Distr.: Colombia (Amazonas (Leticia), Caldas, Valle del Cauca (Buenaventura), Córdoba (Montería)), Costa Rica, Guatemala, Nicaragua, Panama, Suriname, Trinidad and Tobago, Venezuela. brunnea auctt., nec Cresson, 1912: Prado, 1969: 51.1 (in part). palenque Marshall & Buck, 2009 in Marshall et al., 2009: 53. Type locality: Ecuador. Pichincha, 47 km s. Sto. Domingo, Río Palenque Biological Station, 250 m. HT F (DEBU). Distr.: Colombia (Córdoba (Montería)), Ecuador. TABLE 1. Syringogastridae species recorded from Colombia. Genus species author Syringogaster brunnea Cresson, 1912 brunneina Marshall & Buck, 2009 palenque Marshall & Buck, 2009 CATALOGUE OF SYRINGOGASTRIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 559 References Cresson, E.T. (1912) Descriptions of several new Neotropical acalyptrate Diptera. Entomological News, 23, 389–396. Curran, C.H. (1934) The families and genera of North American Diptera. Ballou Press, New York, 512 pp. http://dx.doi.org/10.5962/bhl.title.6825 Feijen, H.R. (1989) Diopsidae. In: Griffiths, G.C.D. (Ed.), Flies of the Nearctic Region, 9 (12), pp. 1–122. [E. Schweizerbart'sche Verlagsbuchhandlung, Stuttgart] Hennig, W. (1958) Die Familien der Diptera Schizophora und ihre phylogenetischen Verwandtschaftsbeziehungen. Beiträge zur Entomologie, 8 (5–6), 505–688. Marshall, S.A., Buck, M., Skevington, J.H. & Grimaldi, D. (2009) A revision of the family Syringogastridae (Diptera: Diopsoidea). Zootaxa, 1996, 1–80. Marshall, S.A. & Buck, M. (2010) 61. Syringogastridae (Syringogastrid flies). In: Brown, B.V., Borkent, A., Cumming, J.M., Wood, D.M., Woodley, N.E. & Zumbado, M.A. (Eds.), Manual of Central American Diptera. Vol. II. NRC Research Press, Ottawa, pp. 843–846. Meier, R. & Hilger, S. (2000) On the egg morphology and phylogenetic relationships of Diopsidae (Diptera: Schizophora). Journal of Zoological Systematics and Evolutionary Research, 38, 1–36. http://dx.doi.org/10.1046/j.1439-0469.2000.381128.x Papavero, N. (1964) Notes on the myrmecomimicry of Syringogaster rufa Cresson (Diptera, Acalyptratae, Megamerinidae). Papéis Avulsos de Zoologia, 16 (12), 110–113. [São Paulo] Prado, A.P. (1969) Syringogastridae, uma nova família de dípteros Acalyptratae, com descrição de seis espécies nova do gênero Syringogaster Cresson. Studia Entomologica, 12 (14), 1–32. Prado, A.P. (1975) 51. Family Syringogastridae. In: Papavero, N. (Ed.), A Catalogue of the Diptera of the Americas South of the United States. Departamento de Zoologia, Secretaria de Agricultura, São Paulo, pp. 1. Prado, A.P. & Papavero, N. (2002) Insecta - Diptera - Syringogastridae. Fauna da Amazônia Brasileira, 4, 1. [Belém] 560 · Zootaxa 4122 (1) © 2016 Magnolia Press RUNG Zootaxa 4122 (1): 561–565 http://www.mapress.com/j/zt/ Copyright © 2016 Magnolia Press Article ISSN 1175-5326 (print edition) ZOOTAXA ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.4122.1.44 http://zoobank.org/urn:lsid:zoobank.org:pub:6B48CF8C-EF50-47CB-A95D-52DEDB688279 FAMILY CONOPIDAE LEONARDO SILVESTRE GOMES ROCHA1,2* & CÁTIA ANTUNES DE MELLO-PATIU1,3 1 Laboratório de Biodiversidade e Sistemática de Diptera, Departamento de Entomologia, Museu Nacional, Universidade Federal do Rio de Janeiro, Rio de Janeiro, Brazil. 2 Instituto Federal de Educação, Ciência e Tecnologia do Rio de Janeiro, Campus Nilópolis, Rio de Janeiro, Brazil. E-mail: leonardo.rocha@ifrj.edu.br 3 CNPq research fellow. E-mail: camello@acd.ufrj.br *Corresponding author: leonardo.rocha@ifrj.edu.br Abstract A catalogue of the Conopidae known from Colombia, with 16 species distributed in four genera, is presented. The richest group is Stylogaster Macquart. Geographic distributions are given for each species. Key words: biodiversity, catalogue, Conopidae, distribution, Neotropical Introduction Conopidae Latreille, 1802, or thick-headed flies, are a family of parasitoid Diptera, found all over the world, except in Polar Regions and Pacific islands. About 800 extant species in 56 genera are recognized (Gibson & Skevington 2013, Pape & Thompson 2013), and three fossils are known, from Baltic and Dominican amber (dating minimally from ca. 47 mya) (Hennig 1966, Gibson & Skevington 2013, Rocha et al. 2015). Conopids can be recognized by the head thicker than the slender body, dorsal arista or apical stylus, elongated antennae and prementum, cell r4+5 closed, vein R4+5 deflected forwards, and specialized female genitalia (elongated and tube-shaped, formed by the segments 6, 7, 8 and 9, in stylogastrines or formed by flat plates on sternite 5 in the others). Adults of Conopidae are nectarivorous, feeding mainly on the flowers of Apiaceae, Asteraceae and Lamiaceae (Lopes 1937, Smith & Peterson 1987). Larvae are parasites mainly in hymenopterans, except the stylogastrines, which develop into orthopteroids and at least oviposit on calyptrate Diptera (Couri & Pont 2006, Couri & Barros 2010). Gibson et al. (2012) and Gibson & Skevington (2013) recognized five subfamilies in the family classification: Conopinae, Dalmanniinae, Myopinae, Stylogastrinae and Zodioninae, but only Conopinae, Stylogastrinae and Zodioninae are recorded for Colombia. The family is poorly known in Colombia and, to date, only 16 species in four genera are recorded in the literature. Data are based on the catalogue of Papavero (1971), the monograph of Camras and Parrillo (1985), and recent literature. The sex and depository collection of the types are given, but [?] is used when this information is unknown. Recorded departments are underlined, specific localities are in brackets, and “Colombia” is used when specifications of departments or localities are not available. Most of the records of Conopidae from Colombia are unspecific about the department or cities, making difficult the identification of sampled regions. The result of this catalog reflects the few studies on Conopidae in this country and the conopid fauna shall be actually much richer than the known records, since there are many potentially rich habitats in all over Colombian territory. Sixteen species of Conopidae are recorded from Colombia. Stylogaster Macquart is the richest group, with 10 species. Physoconops Szilády with 4 species, and Physocephala Schiner and Zodion Latreille, with one species each, complete the list (Table 1). Accepted by S. Nihei: 10 Apr. 2015; published: 14 Jun. 2016 561 Acronyms used for the depositories AMNH—American Museum of Natural History, New York. BMNH—The Natural History Museum, London. CAS—California Academy of Sciences, San Francisco. FMNH—Field Museum of Natural History, Chicago. NMW—Naturhistorisches Museum Wien, Vien. SMT—Staatliches Museum fur Tierkunde, Dresden. UKaL— University of Kansas, State Biological Survey of Kansas Invertebrate Collection, Lawrence. USNM—National Museum of Natural History, Smithsonian Institution, Washington. ZMUC—Zoological Museum, University of Copenhagen, Copenhagen. List of abbreviations cat.—catalogue des.—designation distr.—by distribution mon.—monotypy orig. des.—by original designation refs.—references rev.—revision ST—Syntype HT—Holotype F (f#)—Female M (m#)—Male ?—unknown Catalogue of Conopidae of Colombia Family Conopidae Latreille, 1802 Genus Physocephala Schiner Physocephala Schiner, 1861: 137. Type species, Conops rufipes Fabricius, 1781 (orig. des.). inhabilis (Walker), 1849: 672 (Conops). Type locality: Unknown. HT F (BMNH). Distr.: Mexico to Argentina, “Colombia”. Refs.: Camras, 1957: 215 (rev.); Papavero, 1971: 7 (cat.). Genus Physoconops Szilády Physoconops Szilády, 1926: 588. Type species, Conops brachyrrynchus Macquart, 1843 (orig. des.) = Conops obscuripennis Williston, 1882. Subgenus Aconops Kröber Aconops Kröber, 1919: 142. Type species, Conops antennatus Malloch, 1933. costatus (Fabricius), 1805: 175, (Conops). Type locality: “America Meridionali”. HT sex? (ZMUC). Distr.:“Colombia”, Surinam, Brazil. Refs.: Camras, 1955: 174 (rev.); Papavero, 1971: 3 (cat.). 562 · Zootaxa 4122 (1) © 2016 Magnolia Press ROCHA & DE MELLO-PATIU Subgenus Pachyconops Camras Pachyconops Camras, 1955: 161. Type species, Conops bulbirostris Loew, 1853. brachyrrhynchus (Macquart), 1843: 172, (Conops). Type locality: “Amerique Septentrionale”. HT (?). Distr.: United States, Mexico, “Colombia”. Refs.: Camras, 1955: 163 (rev.); Papavero, 1971: 4 (cat.). gracilis (Williston), 1885: 377, (Conops). Type locality: USA, Arizona. HT M (UKaL). Distr.: United States, Mexico, “Colombia”. Refs.: Camras, 1955: 163 (rev.); Papavero, 1971: 4 (cat.). guyanicus (Curran), 1934: 417, (Conops). Type locality: Guyana, Bartica, Kartabo. HT F (AMNH). Distr.: “Colombia”, Guyana, Venezuela, Peru, Brazil. Refs.: Camras, 1955 (rev.): 167; Papavero, 1971: 4 (cat.). Genus Stylogaster Macquart Stylogaster Macquart, 1835: 38. Type species, Conops stylata Fabricius, 1805 (mon.). abdominalis Kröber, 1914: 352. Type locality: Bolivia, La Paz, Mapirí, Sarampiuní, 700m. HT F (SMT). Distr.: Colombia (Meta (Villavicencio)), Ecuador, Bolivia. Refs.: Aldrich, 1930: 26; Papavero, 1971: 13 (cat.); Camras & Parrillo, 1985: 120 (rev.). breviventris Aldrich, 1930: 19. Type locality: Guyana, Bartica, Kartabo. HT F (AMNH). Distr.: “Colombia”, Ecuador, Peru, Guyana; Brazil. Refs.: Papavero, 1971: 13 (cat.); Camras & Parrillo, 1985: 117 (rev.). decorata Aldrich, 1930: 16. Type locality: Costa Rica, San Mateo: Higuito & La Suiza de Turrialba. HT M (USNM). Distr.: Mexico, Costa Rica, Panama, “Colombia”, Ecuador, Peru, Brazil. Refs.: Papavero, 1971: 14 (cat.); Camras & Parrillo, 1985: 121 (rev.). geijskesi Curran, 1942: 65. Type locality: Surinam, Serabreck. HT M (AMNH). Distr.: Colombia (Valle del Cauca (San José de Anchicaya)), Surinam. Refs.: Papavero, 1971: 14 (cat.); Camras & Parrillo, 1985: 124 (rev.). indistincta Aldrich, 1930: 24. Type locality: Panama, Canal Zone. Cano Saddle, Gatún Lake. HT M (USNM). Distr.: Guatemala, Costa Rica, Panama, “Colombia”, Peru, Brazil. Refs.: Papavero, 1971: 14 (cat.); Camras & Parrillo, 1985: 120 (rev.). minuta Townsend, 1897: 27. Type locality: Mexico, Veracruz, Rio Nautla. ST M F (BMNH). Distr.: Mexico, Guatemala, Honduras, Panama, “Colombia”, Ecuador, Venezuela, Peru. Refs.: Aldrich, 1930: 13; Papavero, 1971: 14 (cat.); Camras & Parrillo, 1985: 118 (rev.). paradecorata Camras & Parrillo, 1985: 121. Type locality: Brazil, Santa Catarina, Nova Teutônia, 300-500m. HT M (FMNH). Distr.: Costa Rica, Colombia (Valle del Cauca (Buenaventura, San José de Anchicaya)), Ecuador, Peru, Brazil. rectinervis Aldrich, 1930: 23. Type locality: Guyana, Bartica, Kartabo. HT M (AMNH). Distr.: Costa Rica, “Colombia”, Ecuador, Peru, Guyana; Brazil. Refs.: Papavero, 1971: 15 (cat.); Camras & Parrillo, 1985: 117 (rev.). schlingeri Camras & Parrillo, 1985: 113. Type locality: Colombia, Valle, 6miles W Calli. HT M (CAS). Distr.: Colombia (Valle del Cauca (Cali, San José de Anchicaya)). stylata (Fabricius), 1805: 177, (Conops). Type locality: “America meridionale”. HT F (ZMUC). Distr.: “Colombia”, Ecuador, Brazil, Paraguay, Argentina. Refs.: Aldrich, 1930: 7; Papavero, 1971: 15 (cat.); Camras & Parrillo, 1985: 115 (rev.); Rocha & Mello-Patiu, 2009: 559 (rev.). CATALOGUE OF CONOPIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 563 Genus Zodion Latreille Zodion Latreille, 1797: 162. Type species, Myopa cinerea Fabricius, 1794. pictum Schiner, 1868: 370. Type locality: “Colombien” (= Colombia, Ecuador or Venezuela). HT F (NMW). Distr.: Mexico; Costa Rica, “Colombia”, Venezuela, Brazil, Bolivia. Refs.: Kröber, 1927: 130; Camras, 1953: 397 (rev.); Pearson & Camras, 1978: 205 (rev.); Papavero, 1971: 12 (cat.). TABLE 1. Species of Conopidae recorded for Colombia. Genus Species Author Physocephala inhabilis (Walker, 1849) Physoconops costatus (Fabricius, 1805) brachyrrhynchus (Macquart, 1843) gracilis (Willinston, 1885) guyanicus (Curran, 1934) abdominalis Kröber, 1914 breviventris Aldrich, 1930 decorata Aldrich, 1930 geijskesi Curran, 1942 indistincta Aldrich, 1930 minuta Townsend, 1897 paradecorata Camras & Parrillo, 1985 rectinervis Aldrich, 1930 schlingeri Camras & Parrillo, 1985 stylata (Fabricius, 1805) pictum Schiner, 1868 Stylogaster Zodion References Aldrich, J.M. (1930) American Two-Winged Flies of the Genus Stylogaster Macquart. Proceedings of the U.S. National Museum, 78, 1–27. http://dx.doi.org/10.5479/si.00963801.78-2852.1 Camras, S. & Parrillo, P.P. (1985) Review of New World Stylogaster (Diptera: Conopidae). Annals of the Entomological Society of America, 78, 111–126. http://dx.doi.org/10.1093/aesa/78.1.111 Camras, S. (1953) Notes on Neotropical Zodion and Parazodion (Diptera: Conopidae). Journal of Natural History, Series 12, 65 (6), 395–399. Camras, S. (1955) A review of the New World flies of the genus Conops and allies (Diptera: Conopidae). Proceedings of the United States National Museum, 105, 155–187. http://dx.doi.org/10.5479/si.00963801.3355.155 Camras, S. (1957) A review of New World Physocephala (Diptera: Conopidae). Annals of the Entomological Society of America, 50 (3), 213–218. http://dx.doi.org/10.1093/aesa/50.3.213 Couri, M.S. & Pont, A.C. (2006) Eggs of Stylogaster Macquart (Diptera: Conopidae) on Madagascan muscids (Diptera: Muscidae). Proceedings of the California Academy of Sciences, 57 (16), 473–478. 564 · Zootaxa 4122 (1) © 2016 Magnolia Press ROCHA & DE MELLO-PATIU Couri, M.S. & Barros, G.P.S. (2010) Diptera hosts of Stylogaster Macquart (Diptera, Conopidae) from Madagascar and South Africa. Revista Brasileira de Entomologia, 54, 361–366. http://dx.doi.org/10.1590/S0085-56262010000300003 Curran, C.H. (1934) The Diptera of Kartabo, Bartica District, British Guiana, with descriptions of new species from other British Guiana localities. Bulletin of the American Museum of Natural History, 66, 287–532. Curran, C.H. (1942) American Diptera. Bulletin of the American Museum of Natural History, 80, 51–84. Fabricius, J.C. (1781) Species Insectoruon Exibentes Eorum Differentias Specificas, Synonyma Auctorum, Loca Natalia, Metamorphosin, Adiectis Observationibus, Descriptionibus. Tom II. Impensis Carol. Ernest Bohnii, Hamburg, 552 pp. Fabricius, J.C. (1794) Entomologia Systematica emendate et aucta. Vol. 4. 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CATALOGUE OF CONOPIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 565 Zootaxa 4122 (1): 566–575 http://www.mapress.com/j/zt/ Copyright © 2016 Magnolia Press Article ISSN 1175-5326 (print edition) ZOOTAXA ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.4122.1.45 http://zoobank.org/urn:lsid:zoobank.org:pub:E7B06831-4715-46F5-8B86-8565AB78580D FAMILY LONCHAEIDAE JULIANA MORGADO FERNANDES & MÁRCIA SOUTO COURI* Museu Nacional, Universidade Federal do Rio de Janeiro. E-mail: jumorgado89@gmail.com, courimarcia@gmail.com *Corresponding author: courimarcia@gmail.com Abstract This family was poorly known in the Neotropical region, mainly in Colombia. However, recent studies increased its knowledge with new distribution records. This catalogue presents 44 species distributed in three genera. Key words: catalogue, Colombia, distribution, Lonchaeidae Introduction Lonchaeidae Rondani, 1856 species are moderately small sized Diptera (body length 3.0-6.0 mm), stout, relatively setose, shiny black to metallic blue or green, with wings usually hyaline and abdomen broad and flat. Frons narrower in male than in female. Ocellar, inner and outer vertical setae strong. Vibrissa absent, but one or more subvibrissal vibrissa-like setae sometimes present. Face broad, usually without strong carina. Eye large, pilose or bare. Thorax with one postpronotal seta, two notopleurals, one presutural and one postsutural supralar, two postsutural dorsocentral, two prescutelar acrostichal, often one weak intra-alar, and two postalars. Legs mostly dark, tarsi sometimes partially or entirely yellow. Wing with anal lobe and alula well developed, subcosta complete and with costagial and humeral weakenings and subcostal break. Halter dark brown (Norrbom 2010). The larvae of most Lonchaeidae are invaders in diseased or injured plants, a few species of Dasiops Rondani are pests of Passiflora (Passifloraceae), and some species of Neosilba McAlpine and Lonchaea Fallén breed in damaged fruit, vegetables or cacti. In Colombia, specimens were obtained from fruits and attacking Passiflora (Norrbom & McAlpine 1997). Lonchaeidae belong to the superfamily Tephritoidea and may be the most basal clade of the superfamily (McAlpine 1989, Korneyev 1999). The family has a worldwide distribution, with 523 species in nine genera in the world, approximately 100 species in four genera in the Neotropical region (Norrbom 2010), and 44 species in three genera recorded from Colombia (Table 1). Acronyms used for the depositories AMNH—American Museum of Natural History, New York, USA BMNH—The National Museum of Natural History, London, England CAS—California Academy of Science, USA CNC—Canadian National Collection of Insects, Ontario, Canada IZAM—Instituto de Zoología Agrícola, Universidad Central de Venezuela, Maracay, Venezuela MEUNPRG—Museo de la Universidad Nacional Pedro Ruiz Gallo, Lambayeque, Peru MENT-UT—Museo de Entomología de la Universidsd del Tolima, Tolima, Colombia MZUSP—Museu de Zoologia, Universidade de São Paulo, São Paulo, Brazil USNM—National Museum of Natural History, Washington D.C., USA ZMUC—Zoological Museum, University of Copenhagen, Copenhagen, Denmark 566 Accepted by S. Nihei: 10 Apr. 2015; published: 14 Jun. 2016 List of abbreviations cat.—catalogue col.—collector des.—designation distr.—geographic distribution mon.—monotypy n. comb.—new combination orig. des.—original designation pp.—pages refs.—references rev.—revision HT—Holotype NT—Neotype LT—Lectotype ST—Syntype T—Type (unspecified) NA—Not applicable F (f#)—Female M (m#)—Male Catalogue of Lonchaeidae of Colombia Family Lonchaeidae Rondani, 1856 Subfamily Dasiopinae Morge, 1963 Tribe Dasiopini Morge, 1963 Genus Dasiops Rondani Dasiops Rondani, 1856: 120. Type species, Lonchaea latifrons Meigen, 1826, des. I.C.Z.N., 1963: 114. Dasiopa Rondani; Schiner, 1868 (misspelling) Dasyops Rondani; Coquillet, 1910 (misspelling) Dasiops, error for Dasyops brevicornis (Williston), 1896: 379 (Lonchaea). Type locality: St. Vincent, West Indies. HT F (BMNH). Distr.: Saint Vincent Islands, Jamaica and Colombia. Refs.: Norrbom & McAlpine, 1997: 196; Hernandez, 2011: 30; Castro, 2012: 22;Castro et al., 2012: 6682; Wychuys et al., 2012: 96; Uchôa, 2012: 285. Dasiops brevicornis, Steyskal, 1980: 169. (n. comb.); Castro et al., 2012: 6682 (first record for Colombia). caustonae Norrbom & McAlpine, 1997: 197. Type locality: In Passiflora mollissima, Venezuela, Muchuchies. HT F (IZAM). Distr.: Venezuela, Bolivia, Ecuador and Colombia. Refs.: Causton & Peña, 2002: 169; Umaña, 2005: 59; Hernandez, 2011: 31; Castro et al., 2012: 6689, Castro, 2012: 19-20; Uchôa, 2012: 285. chotanus Korytkowski, 1971: 93. Type locality: Peru, Cajamarca, Pasamayo, Chota,. HT F (UNPRG). Distr.: Peru and Colombia. Refs.: Causton & Peña, 2002: 169; Hernandez, 2011: 33; Castro et al., 2012: 6687 (first record for Colombia). clarionus McAlpine, 1964a: 682. Type locality: Mexico, Baja California,Clarion Isld. HT M (CAS). Distr.: Mexico, Peru and Colombia. Refs.: Korytkowski & Ojeda, 1971: 96; Hernadez, 2011: 34; Castro et al., 2012: 6689 (first record for Colombia). CATALOGUE OF LONCHAEIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 567 curubae Steykal, 1980: 166. Type locality: Colombia, Cundinamarca, San Bernardo. HT F (USNM). Distr.: Colombia (Cundinamarca (San Bernardo)). Refs.: Norrbom & McAlpine, 1997: 199; Causton & Peña, 2002: 270; Umanã, 2005: 59; Hernandez, 2011: 36; Castro, 2012: 23; Wychuys et al., 2012: 92; Uchôa, 2012: 285; Carrero et al., 2013: 438. dentatus Norrbom & McAlpine, 1997: 200. Type locality: Peru, Junin. HT F (USNM). Distr.: Peru and Colombia. Refs.: Causton & Peña, 2002: 270; Hernandez, 2011: 38; Uchôa, 2012: 285, Carrero et al., 2013: 438. frieseni Norrbom & McAlpine, 1997: 200 Type locality: Brazil, Rio de Janeiro, Teresopolis. HT F (CNC). Distr.: Brazil and Colombia. Refs.: Aguiar-Menezes et al., 2004: 115; Hernandez, 2011: 42; Wychuys et al., 2012: 96; Uchôa, 2012: 285. gracilis Norrbom & McAlpine, 1997: 202. Type locality: Colombia, Coachí. HT F (USNM). Distr.: Colombia, Panamá, Venezuela and Costa Rica. Refs.: Causton & Peña, 2002: 270; Umanã, 2005: 60; Hernandez, 2011: 43; Carrero et al., 2013: 438; Castro, 2012: 17; Wychuys et al., 2012: 96; Uchôa, 2012: 285. inedulis Steyskal, 1980: 167. Type locality: Panamá, Chiriquí. HT F (USNM). Distr.: Colombia, Panamá and Peru. Refs.: Norrbom & McAlpine, 1997: 203; Causton & Peña, 2002: 270; Uchôa-Fernandes et al., 2002; AguiarMenezes et al., 2004: 115; Umanã, 2005: 60; Hernandez, 2011: 47; Wychuys et al., 2012: 96; Uchôa, 2012: 285; Carrero, 2013: 19-63; Carrero et al., 2013: 437-447; Felix, 2013: 6. longulus Norrbom & McAlpine, 1997: 205. Type locality: Brazil, São Paulo, Santa Isabel. HT F (MZUSP). Distr.: Brazil and Colombia. Refs.: Aguiar-Menezes et al., 2004: 114; Uchôa, 2012: 286; Felix, 2013: 6. luzestelae Hernandez, 2011: 75 Type locality: Colombia, Valle del Cauca. HT F (MEUNPRG). Distr.: Colombia (Valle del Cauca) and Panamá. Refs.: Castro, 2012: 26. passifloris McAlpine, 1964a: 691. Type locality: USA, Florida. HT F (USNM). Distr.: USA and Colombia. Refs.: Causton & Peña, 2002: 270; Umanã, 2005: 59; Hernandez, 2011: 50; Castro, 2012: 23; Wychuys et al., 2012: 96; Uchôa, 2012; 286. peruanus Hennig, 1948: 348. Type locality: Peru, Huánuco, Boca Pachieta. HT F. Distr.: Peru and Colombia. Ref.: Hernandez, 2011: 52. Dasiops peruana (Hennig), McAlpine, 1964a: 689. Dasiops peruanus (Hennig), Korytkowski & Ojeda, 1971: 96. plaumanni McAlpine, 1964a: 662. Type locality: Brazil, Nova Teutonia,. HT F (CNC). Distr.: Brazil and Colombia. Refs.: Hernandez, 2011: 54; Castro, 2012: 25; Castro et al., 2012: 6684. plumeus McAlpine, 1964a: 676. Type locality: Colombia, Rio Sambingo, 32 millas N. La Union Narino. HT F (CAS). Distr.: Colombia. Refs.: Korytkowski & Ojeda, 1971; Hernandez, 2011: 57. rugifrons Hennig, 1948: 349. Type locality: Peru, Cuzco, Rosalina. HT F (?) . Distr.: Peru, Venezuela and Colombia. Refs.: Korytkowski & Ojeda 1971: 95; Norrbom & McAlpine, 1997: 206; Causton & Peña, 2002:169; Umaña, 2005: 60; Castro, 2012: 24; Castro et al., 2012: 6684; Uchôa, 2012: 286. rugulosos Norrbom & McAlpine, 1997: 207. Type locality: Mexico. HT F (USNM). Distr.: Trinidad and Tobago, and Colombia. Refs.: Hernandez, 2011: 62; Uchôa, 2012: 286. saltans (Townsend), 1913: 265 (Acucula). Type locality: Peru, Santa Ana, Rio Rimac Valley. HT (?). Distr.: Colombia and Peru. Refs.: Causton & Peña, 2002: 270; Umaña, 2005: 65; Delgado & Kondo, 2010: 1-10; Henandez, 2011: 64. 568 · Zootaxa 4122 (1) © 2016 Magnolia Press FERNANDES & COURI yepezi Norrbom & McAlpine, 1997: 207. Type locality: Colombia, Antioquia. HT F (USNM). Distr.: Colombia (Antioquia) and Brazil. Refs.: Causton & Peña, 2002: 270; Umanã, 2005: 60; Hernandez, 2011: 66; Castro, 2012: 18; Wychuys et al., 2012: 96; Uchôa, 2012: 286; Carrero et al., 2013: 438 Subfamily Lonchaeinae Rondani, 1856 Tribe Lonchaeini Rondani, 1856 Genus Lonchaea Fallén Lonchaea Fallén, 1820: Type species: Musca chorea Fabricius, 1782, des. Westwood, 1840: 150. Lonchoea, error for Lonchaea. aculeata Bezzi, 1910: 24. Type locality: Brazil. HT (sex?). Distr.: Peru and Colombia. Refs.: Korytkowski & Ojeda 1971: 107; Luna, 1988: 31; Wychuys et al., 2012: 96. cristula McAlpine, 1964b: 744. Type locality: Mexico, Teotihuacan. HT M (CNC). Distr.: Texas, Mexico, Ecuador, Peru and Colombia. Refs: Korytkowski & Ojeda, 1971: 101; Luna, 1988: 29; Wychuys et al., 2012: 96. curiosa McAlpine, 1964b: 749. Type locality: Colombia, Funza. HT M (CNC). Distr.: Mexico, Colombia (Cundinamarca (Funza)), Ecuador and Peru. Refs.: Korytkowski & Ojeda, 1971: 101; Luna, 1988: 30; Wychuys et al., 2012: 96. echinapinna McAlpine, 1964b: 754. Type locality: Colombia, Palmira, Tolima. HT M (CNC). Distr.: Colombia (Tolima, Cauca (Popayán), Valle del Cauca (Palmira, w. Cali)). Ref.: Luna, 1988: 32. longicornis Williston, 1896: 378. Type locality: St. Vincent, West Indies. HT M (BMNH). Distr.: USA, Antillas, Colombia and Peru. Refs.: Korytkowski & Ojeda 1971: 107, Luna, 1988: 31. Genus Neosilba McAlpine Neosilba McAlpine, 1962:138-144. Type species: Lonchaea batesi Curran, 1932. amphora Galeano-Olay & Canal, 2012: 368. Type locality: “Colombia”, Santander, Provincia De Vélez, Chipata. HT M (MEN-UT). Distr.: Colombia. angusta Galeano-Olaya & Canal, 2012: 374. Type locality: “Colombia”, Tolima. HT M (MEN-UT). Distr.: Colombia (Tolima). batesi (Curran), 1932: 10 (Lonchaea). Type locality: Guatemala, Escuintla. HT M (AMNH). Distr.: Guatemala, Mexico, Colombia, Panamá, Florida (USA), Peru, Costa Rica and El Salvador. Refs.: McAlpine & Steyskal, 1982: 110; Korytkowsky & Ojeda. 1971: 113; Ahlmark & Steck, 1997: 116; Wychuys et al., 2012: 96; Carrero et al., 2012: 438; Uchôa, 2012: 286. Silba batesi, Korytkowsky & Ojeda. 1971: 113 (n. comb.) Neosilba batesi, McAlpine, 1962: 138-140 (n. comb.); McAlpine & Steyskal, 1982: 110. bifida Strikis & Prado, 2005: 2-3. Type locality: Brazil, São Paulo, São Bento do Sapucai. HT F (MZUSP). Distr.: Brazil and Colombia (Tolima (Ibagué)). Ref.: Caires et al., 2009: 274; Strikis & Prado, 2009, 50; Galeano-Olaya & Canal, 2012: 385 (first record for Colombia); Uchôa, 2012: 286; Uchôa et al., 2012: 633. certa (Walker), 1850: 364 (Anthomyia). Type locality: “Brazil”. HT M (BMNH). Distr.: Brazil, Puerto Rico, CATALOGUE OF LONCHAEIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 569 Mexico, Colombia, Costa Rica and El Salvador. Refs.: McAlpine & Steyskal, 1982: 112; Urrutia & Korytkowsky, 2004: 60; Freitas, 2008: 55; Gattelli et al., 2008: 237; Rodrigues, 2008: 19; Souza et al., 2008: 520; Caires et al., 2009: 274; Souza-Filho et al., 2009: 33; Strikis & Prado, 2009: 50; Uchôa & Nicácio, 2010: 731; Montes et al., 2012: 4; Uchôa, 2012: 287; Uchôa et al., 2012: 633. Lonchaea certa, Stein 1901: 192 (n. comb.) Silba certa, Gaud & Martorell 1973: 249 (n. comb.) Neosilba certa, McAlpine & Steyskal 1982: 112 (n. comb.) chinchinaia Urrutia & Korytkowsky, 2004: 61. Type locality: Colombia, Norte de Santander, Blonay. HT M (MZUSP). Distr.: Ecuador, Colombia (Norte de Santander (Chinácota, Blonay) and Peru. colombiana Urrutia & Korytkowsky, 2004: 63. Type locality: Colombia, Cundinamarca. HT M (MZUSP). Distr.: Colombia (Cundinamarca) and Panamá. concava Galeano-Olaya & Canal, 2012: 371. Type locality: Colombia, Tolima, Espinal. HT M (MEN-UT). Distr.: Colombia (Tolima (Espinal). convexa Galeano-Olaya & Canal, 2012: 370. Type locality: Colombia, Tolima, Ibagué. HT M (MEN-UT). Distr.: Colombia (Tolima (Ibagué). cryptocerta Urrutia & Korytkowsky, 2004: 67. Type locality: Colombia, Cali, La Elvira. HT M (MZUSP). Distr.: Colombia (Valle del Cauca (Cali, La Elvira)) and Brazil. distospinosa Galeano-Olaya & Canal, 2012: 377. Type locality: Colombia, Tolima, Cajamarca. HT M (MEN-UT). Distr.: Colombia (Tolima (Cajamarca)). glaberrima (Wiedemann), 1830: 475 (Lonchaea). Type locality: “West Indies”. HT F (ZMUC). Distr.: India, Trinidad, Brazil, Colombia, Peru, Ecuador, Panama, Costa Rica, Honduras, Guatemala, Mexico and El Salvador. Refs.: McAlpine & Steyskal, 1982: 114; Urrutia & Korytkowsky, 2004: 72; Bittencourt et al., 2006: 282; Rodrigues, 2008: 19; Souza- Filho et al., 2009: 33; Strikis & Prado, 2009: 50; Delgado & Kondo, 2010: 96; Uchôa & Nicácio, 2010: 731; Camargos et al., 2011: 615-616; Uchôa, 2012: 288; Felix, 2013: 6. Carpolonchaea pendula Hennig, 1948: 360 (n. comb.) Silba pendula, Korytkowski & Ojeda, 1971: 112 (n. comb.) Neosilba glaberrima, McAlpine & Steyskal 1982: 114 (n. comb.) major (Malloch), 1920: 246 (Lonchaea). Type locality: Mexico, Chiapas,. HT M (BMNH). Distr.: Peru, Ecuador, Colombia, Panama, El Salvador and Mexico. Refs.: McAlpine & Steyskal, 1982: 117; Urrutia & Korytkowsky, 2004: 82; Uchôa, 2012: 289. Carpolonchaea magna, Hennig, 1948: 24 (n. comb) Neosilba major, McAlpine, 1962: 140 (n. comb); McAlpine & Steyskal, 1982: 117. Silba magna, Korytkowski & Ojeda, 1971: 111 (n. comb) orbata Galeano-Olaya & Canal, 2012: 372. Type locality: Colombia, Tolima, Ibagué. HT M (MEN-UT). Distr.: Colombia. parva (Hennig), 1948: 359 (Carpolonchaea). Type locality: Colombia, Tolima, Libano. HT M (MEN-UT). Distr.: Costa Rica, Mexico, Peru, Brazil, Honduras and Colombia (Tolima (Líbano)). Refs.: McAlpine & Steyskal, 1982: 121; Urrutia & Korytkowsky, 2004: 93; Bittencourt et al., 2006: 282; Rodrigues, 2008: 19; Galeano-Olaya & Canal, 2012: 385 (first record for Colombia); Uchôa, 2012: 290; Felix, 2013: 6. Silba parva, Korytkowski & Ojeda, 1971: 111. (n. comb) Neosilba parva, McAlpine, 1962: 140-143 (n. comb); McAlpine & Steyskal, 1982:121. Lonchaea pendula Bezzi, 1919: 249, pro parte. 570 · Zootaxa 4122 (1) © 2016 Magnolia Press FERNANDES & COURI pendula (Bezzi), 1919: 249 (Lonchaea). Type locality: Brazil, Campinas. Distr.: Brazil, Mexico, Peru and Colombia. LT M (USNM). Refs.: McAlpine & Steyskal, 1982: 123; Urrutia & Korytkowsky, 2004: 95; Bittencourt et al., 2006: 282; Freitas, 2008: 52; Rodrigues, 2008: 19; Souza et al., 2008: 520; Caires et al., 2009: 274; Strikis & Prado, 2009: 50; Uchôa & Nicácio, 2010: 731; Camargos et al., 2011: 615-66; Uchôa Montes et al., 2012: 290; Felix, 2013: 6. perezi (Romero & Ruppel), 1973: 165 (Silba). Type locality: Puerto Rico. Distr.: Brazil, Guayana, Colombia, Cuba, Honduras, Jamaica, Puerto Rico, Republic Dominican, USA, Caribe, Mexico, Costa Rica and El Salvador. HT M (USNM). Refs.: McAlpine & Steyskal, 1982: 126; Urrutia & Korytkowsky, 2004: 97; Uchôa, 2012: 291. Neosilba perezi, Waddill & Weems, 1978: 1-3 (n. comb.) Lonchaea pendula Bezzi, 1919, parte. piracea Galeano-Olaya & Canal, 2012: 383. Type locality: Colombia, Tolima, Ibagué. HT M (MEN-UT). Distr.: Colombia (Tolima (Ibagué)). plana Galeano-Olaya & Canal, 2012: 365. Type locality: Colombia, Tolima, Ibagué. HT M (MENT-UT). Distr.: Colombia. (Tolima (Ibagué)). pseudopendula (Korytkowski & Ojeda), 1971: 113. (Silba). Type locality: Peru, Torgopampa. HT M (MEUNPRG). Distr.: Colombia, Peru, Costa Rica and Panama. Ref.: Steyskal, 1978:72; Urrutia & Korytkowsky, 2004: 99. Neosilba pseudopendula, McAlpine & Steyskal, 1982: 127. (n. comb.) spiculata Galeano-Olaya & Canal, 2012: 374. Type locality: Colombia, Tolima, Cajamarca. HT M (MEN-UT). Distr.: Colombia (Tolima (Cajamarca)). tolimensis Galeano-Olaya & Canal, 2012: 366. Type locality: Colombia, Tolima, Cajamarca. HT M (MEN-UT). Distr.: Colombia. (Tolima (Cajamarca)). zadolicha McAlpine & Steyskal, 1982: 127. Type locality: Columbia [Colombia], Cundinamarca?, Viota. HT M (USNM). Distr.: Brazil, Peru, Colombia and Panamá. Refs.: Aguiar-Menezes et al., 2004: 115; Santos et al., 2004: 653; Urrutia & Korytkowsky, 2004: 105; Silva et al., 2006: 669; Gattelli et al., 2008: 237; Freitas, 2008: 47; Rodrigues, 2008: 19; Souza et al., 2008: 520; Caires et al., 2009: 274; Souza-Filho et al., 2009: 33; Strikis & Prado, 2009: 50; Delgado & Kondo, 2010: 96; Uchôa & Nicácio, 2010: 731; Camargos et al., 2011: 615-616; Montes et al., 2012: 4; Nunes et al., 2012: 8; Wychuys et al., 2012: 96; Uchôa, 2012: 291; Uchôa et al., 2012: 633; Felix, 2013: 6. TABLE 1. Lonchaeidae species recorded from Colombia Subfamily Dasiopinae Tribe Genus Species Author Dasiops brevicornis (Williston, 1896) caustonae Norrbom & McAlpine, 1997 chotanus Korytkowski,1971 clarionus McAlpine, 1964 curubae Steykal, 1980 dentatus Norrbom & McAlpine, 1997 frieseni Norrbom & McAlpine, 1997 gracilis Norrbom & McAlpine, 1997 inedulis Steyskal, 1980 ...Continued on next page CATALOGUE OF LONCHAEIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 571 TABLE 1. (Continued) Subfamily Lonchaeinae Tribe Lonchaeini Genus Lonchaea Neosilba Species Author longulus Norrbom & McAlpine, 1997 luzestelae Hernandez, 2011 passifloris McAlpine, 1964 peruanus (Hennig, 1948) plaumanni McAlpine, 1964 plumeus McAlpine, 1946 rugifrons Hennig, 1948 rugulosus Norrbom & McAlpine, 1997 saltans (Towsend, 1913) yepezi Norrbom & McAlpine,1997 aculeata Bezzi, 1910 cristula McAlpine, 1964b curiosa McAlpine, 1964b: 744 echinapinna McAlpine, 1964a longicornis Williston, 1896 amphora Galeano-Olaya & Canal, 2012 angusta Galeano-Olaya & Canal, 2012 batesi (Curran, 1932) bifida Strikis & Prado, 2005 concava Galeano-Olaya & Canal, 2012 convexa Galeano-Olaya & Canal, 2012 cryptocerta Urrutia & Korytkowsky, 2004 distospinosa Galeano-Olaya & Canal, 2012 glaberrima (Wiedemann, 1830) major (Malloch, 1920) orbata Galeano-Olaya & Canal, 2012 parva (Hennig, 1948) pendula (Bezzi, 1919) perezi (Romero & Ruppel, 1973) piracea Galeano-Olaya & Canal, 2012 plana Galeano-Olaya & Canal, 2012 pseudopendula (Korytkowski & Ojeda, 1971) spiculata (Galeano-Olaya & Canal, 2012 tolimensis Galeano-Olaya & Canal, 2012 zadolicha McAlpine & Steyskal, 1982 References Aguiar-Menezes, E.N., Nascimento, R.B. & Menezes, E.B. 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In: Wüllerstorf-Urbair, B. von (Ed.), Reise der österreichischen Fregatte Novara um die Erde in den Jahren 1857, 1858, 1859 unter den Befehlen des Commodore B. Von Wüllerstorf-Urbair. Zoologischer Theil. Zweiter Band. 1. Abtheilung. B.K. Gerold’s Son, Wien [= Vienna], 388 pp. Silva, F.F., Meirelles, R.N., Redaelli, L.R. & Dal Soglio, F.K. (2006) Diversity of Flies (Diptera: Tephritidae and Lonchaeidae) in Organic Citrus Orchards in the Vale do Rio Caí, Rio Grande do Sul, Southern Brazil. Neotropical Entomology, 35 (5), 666–670. http://dx.doi.org/10.1590/S1519-566X2006000500015 Stein, P. (1901) Die Walkerischen aussereuropäisen Anthomyiden in der Sammlungder British Museum zu London (Dipt.). Zeitschrift für systematische Hymenoptera und Diptera, 1, 185–221. Steyskal G.C. (1980) Two Winged flies of the genus Dasiops (Diptera: Lonchaeidae) attacking flowers or fruit of species of Passiflora (passion fruit, granadilla, curuba, etc). 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Brazilian Journal of Biology, 69 (1), 31–40. http://dx.doi.org/10.1590/s1519-69842009000100004 Townsend, C.H.T. (1913) A jumping-maggot which lives in cactos blooms ((Acucula saltans Gen. Et sp. Nova). Canadian Entomologist, 45, 262–265. http://dx.doi.org/10.4039/Ent45262-8 Uchôa, M.A., Oliveira, I., Molina, R.M.S. & Zucchi, R.A. (2002) Species diversity of frugivorous flies (Diptera: Tephritoidea) from hosts in the cerrado of the State of Mato Grosso do Sul, Brazil. Neotropical Entomology, 31 (4), 515–524. http://dx.doi.org/10.1590/S1519-566X2002000400002 Uchôa, M.A. & Nicácio, J. (2010) New Records of Neotropical Fruit Flies (Tephritidae), Lance Flies (Lonchaeidae) (Diptera: Tephritoidea), and Their Host Plants in the South Pantanal and Adjacent Areas, Brazil. Annals of the Entomological Society of America, 103 (5), 723–733. http://dx.doi.org/10.1603/AN09179 Uchôa, M.A. (2012) Fruit Flies (Diptera: Tephritoidea): Biology, Host Plants, Natural Enemies, and the Implications to Their Natural Control, Integrated Pest Management and Pest Control - Current and Future Tactics. In: Soloneski, S. & Larramendy, M.L. (Eds.), Integrated pest management and pest control: current and future tactics. Rijeka: InTech, 2012. Cap.12, p.271-300. Available from: http://cdn.intechopen.com/pdfs/29609/ InTechFruit_flies_diptera_Tephritoidea_biology_host_plants_natural_enemies_and_the_implications_to_their_natural_co ntrol.pdf (accessed 22 September 2015) Uchôa, M.A., Caires, C.S., Nicácio, J.N. & Duarte, M. (2012) Frugivory of Neosilba Species (Diptera: Lonchaeidae) and Thepytus echelta (Lepidoptera: Lycaenidae) on Psittacanthus (Santalales: Loranthaceae) in Ecotonal Cerrado-South Pantanal, Brazil. Florida Entomologist, 95 (3), 630–640. http://dx.doi.org/10.1653/024.095.0314 Umaña, M.V. (2005) Moscas de la fruta del gênero Dasiops (Diptera: Lonchaeidae) associados a la curuba y recomendaciones generales para su manejo agroecológico em la vereda Canón, município de Sutamarchán-Boyacá. Revista Colombiana de 574 · Zootaxa 4122 (1) © 2016 Magnolia Press FERNANDES & COURI Entomología, 31 (1), 59–65. Urrutia, F.M. (2004) Revisión Taxonómica Parcial del Género Neosilba McAlpine (Diptera: Lonchaeidae) em la Región Neotropical. Tesis de doctorado, Programa CentroAmericano de Maestría en Entomología, Universidad do Panamá, Panamá, 146 pp. Waddill, V.H., Weems, J.R. (1978) The cassava shoot fly, Neosilba perezi (Romero and Ruppel) (Diptera: Lonchaeidae). Entomology Circular, 187, 1–2. [local: Florida Dept. Agric.] Walker, F. (1850) Insecta Saundersiana. In: Saunders, W.W. (Ed.), Insecta Saundersiana. Van Voorts, London, pp. 76. Westwood, J.O. 1840. Order XIII. Diptera Aristotle (Antilata Fabricius. Halteriptera Clairv.). In: An Introduction to the Modern Classification of Insects; Founded on the Natural Habits and Corresponding Organisation of the Different Families. Synopsis of the genera of British Insects. Orme, Brown, Green and Longmans, London, pp. 125–154. Wiedemann, C.R.W. (1830) Aussereuropäische zweiflügelige Insecten. Vol. 2. Schulz, Hamm, 648 pp., 5 pls. Williston, S.W. (1896) On the Diptera of St Vincent (West Indies). Transactions of the Entomological Society of London, 1896, 253–446. http://dx.doi.org/10.1111/j.1365-2311.1896.tb00965.x CATALOGUE OF LONCHAEIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 575 Zootaxa 4122 (1): 576–578 http://www.mapress.com/j/zt/ Copyright © 2016 Magnolia Press Article ISSN 1175-5326 (print edition) ZOOTAXA ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.4122.1.46 http://zoobank.org/urn:lsid:zoobank.org:pub:DD7EDC4B-269F-4D13-9294-440CEB811177 FAMILY PIOPHILIDAE MARTA WOLFF1*, SANDRA PÉREZ1 & DIANA GRISALES2 1 Grupo de Entomologia, Instituto de Biologia, Universidad de Antioquia, Medellín-Colombia. Laboratório de Biodiversidade e Biogeografia de Diptera, Departamento de Zoologia, UFPR, Caixa postal 19020, Curitiba, PR., Brazil, 81531-980. E-mail: martha.wolff@udea.edu.co; sandra.perez@udea.edu.co; ochoa310@gmail.com * Corresponding author: martha.wolff@udea.edu.co 2 Abstract Piophilidae is a little family poorly known in Colombia, with only Piophila casei (L.) and Stearibia nigriceps Meigen reported so far. This catalogue expands the distribution of these species to other localities in the country. Key words: catalogue, Colombia, Diptera, distribution Introduction Piophilidae are known as skipper flies. They are similar to ants in appearance, measuring 3–8 mm long and shining black to dull brown and yellow flies with wings frequently glassy. They could be identified by the head wide as the thorax, with ocelli present. Frons setulose, with zero to two lateroclinate orbital setae. Postocellar setae divergent, one to two vibrissae present Arista usually short. Thorax with zero to two postpronotal. Wing rounded, longer than abdomen, with anal and alula developed, entirely hyaline in Neotropical species. Larvae are typical muscomorph maggots, long, slender, cylindrical with spiculose intersegmental rings (Ozerov & Norrbom 2010). The natural history of the species of Piophilidae is poorly known for the Neotropical region, except for the introduced species They are scavengers as larvae, growing in animal or vegetable decomposing matter; larvae of many species live in animal carrion (Ozerov & Norrbom 2010), hidden between the joints and bones. Piophilidae are cosmopolitan flies and several species are synanthropic and widespread (Ozerov & Norrbom 2010). In the world occur 82 species in 19 genera, and 10 species in the Neotropical region (de Carvalho et al. 2012). In Colombia only two species are reported, Piophila casei (Linnaeus) and Stearibia nigriceps Meigen, both collected in forensic entomology studies. In this catalogue we underline Colombian Department names and parenthetically note specific localities. At the end, we provide a table that includes the species known in Colombia (Table 1). Acronyms used for the depositories CEUA—Colección Entomológica de la Universidad de Antioquia, Medellín, Colombia. LSL—Linnean Society, Burlington House, Piccadilly, London WIV OLQ, England. List of abbreviations cat.—catalogue distr.—distribution m.— meters 576 Accepted by S. Nihei: 26 May 2015; published: 14 Jun. 2016 mon.—monotypy refs.—references T—type Catalogue of Piophilidae of Colombia Family Piophilidae Macquart, 1835 Genus Piophila Fallén Piophila Fallén, 1810: 20. Type species, Musca casei (Linnaeus) 1758: 597 (mon.). Refs.: Pape & Thompson, 2013 (cat. online) casei Linnaeus, 1758: 597 (Musca). Type locality: “Europe”. T (unspecified sex) (LSL). Distr.: Colombia (Cundinamarca (Páramo Chingaza 3030m), Antioquia (Medellín)), Argentina, Bolivia, Brazil, Costa Rica, Cuba, Chile, Ecuador, Jamaica, Ilhas Juan Fernandez, Mexico, Paraguay, Venezuela), Cosmopolitan. Refs.: Steyskal, 1967:1; Wolff et al., 2001:55. Material examined. COLOMBIA: Antioquia: Mpio. [Municipio] Medellín, 1450m, 30 females, v-vi.1999, A. Uribe-Toro leg., net-pig (CEUA); ídem: 26 females, ix-2002, S. Pérez leg., net-pig (CEUA); Boyaca: Tunja, 2690m, 1 male, 8 females, vi-2002, Yuseff & Daza leg., net-pig (CEUA). Genus Stearibia Lioy Stearibia Lioy, 1864: 1105. Type species, Piophila foveolata Meigen (mon.) = nigriceps Meigen, 1826: 397. Refs.: Pape & Thompson, 2013 (cat. online) nigriceps Meigen, 1826: 397. Type locality: “Europe”. T adult (sex unspecifed) (museum unespecifed). Distr.: Colombia (Antioquia (Medellín 1450m), Boyacá (Tunja 2690m), Risaralda (Pereira)), Argentina, Chile, Ecuador, Peru, and Holarctic Region. Refs.: Steyskal, 1967: 1; Pérez et al., 2005 (as foveolata): 3; Martinez et al., 2007: 183; Ozerov & Norrbom, 2010: 869; Grisales et al., 2010: 640. Material examined. COLOMBIA: Antioquia: Mpio. [Municipio] Medellín, 1450m, 38 females, vi-1999, A. Uribe-Toro leg., (CEUA); ídem: 4 females, ix-2002, S. Pérez leg., net-pig (CEUA). TABLE 1. Piophilidae species recorded from Colombia. Genus Species Author Piophila casei (Linnaeus, 1758) Stearibia nigriceps Meigen, 1826 References de Carvalho, C.J.B., Rafael, J.A., Couri, M.S. & Silva, V.C. (2012) Diptera. In: Rafael J.A., Mello, G.A.R., de Carvalho, C.J.B., Casari, S.A. & Constantino, R. (Eds.), Insetos do Brasil. Holos Editora, São Paulo, pp. 701–744. Fallén, C.F. (1810) Specimen Entomologicum novam Diptera disponendi methodum exhibens. Berlingianus, Lundae [= Lund], 26 pp. Grisales, D., Ruiz, M. & Villegas, S. (2010) Insects associated with exposed decomposing bodies in the Colombian Andean Coffee Region. Revista Brasileira de Entomologia, 54 (4), 637–644. http://dx.doi.org/10.1590/S0085-56262010000400016 Linnaeus, C. (1758) Systema naturae per regna tria naturae. Vol. 1. 10th Edition. Holmiae (= Stockolm), 824 pp. Lioy, P. (1864) I dittero distribuiti secondo um nuovo método di classificazione naturale (cont.). Atti dell' Realle Istituto Veneto di Scienze, Lettere. Arti, Series 3, 9, 499–518, 569–604, 719–771, 879–910, 989–1027, 1087–1126, 1311–1352 (cont.). CATALOGUE OF PIOPHILIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 577 Macquart, P.J.M. (1835) Histoire Naturelle des insects, Dipteres. Tome deuxieme. Roret, Paris, 710 pp. Martínez, E., Duque, P. & Wolff, M. (2007) Succession pattern of carrion-feeding insects in Paramo, Colombia. Forensic Science International, 166, 182–189. http://dx.doi.org/10.1016/j.forsciint.2006.05.027 Meigen, J.W. (1826) Systematische Beschreibung der bekannten europäischen zweiflügeligen Insekten. Vol. 5. Fortsmann, Aachen, 412 pp. Ozerov, A. & Norrbom, A. (2010) Piophilidae. In: Brown B.V., Borkent A., Cumming J.M., Wood, D.M., Woodley, N.E. & Zumbado, M.A. (Eds.), Manual of Central American Diptera. Vol. 2. NR C-CNRC, Ottawa, pp. 865–869. Pape, T. & Thompson, F.C. (Eds.) (2013) Systema Dipterorum, version 1.5. http://www.diptera.org/ (accessed April 20 2015) Pérez, S., Duque, P. & Wolff, M. (2005) Successional Behavior and Occurrence Matrix of Carrion-Associated Arthropods in the Urban Area of Medellín, Colombia. Journal of Forensic Science, 50 (2), 1–7. http://dx.doi.org/10.1520/JFS2004046 Steyskals, G.C. (1967) 68. Family Piophilidae. In: Papavero, N. (Ed.), A Catalogue of the Diptera of the Americas South of the United States. Departamento de Zoologia, Secretaria da Agricultura, São Paulo, 33 pp. Wolff, M., Uribe, A., Ortiz, A. & Duque, P. (2001) A preliminar study of forensic entomology in Medellín, Colombia. Forensic Science International, 120, 53–59. http://dx.doi.org/10.1016/S0379-0738(01)00422-4 578 · Zootaxa 4122 (1) © 2016 Magnolia Press WOLFF ET AL. Zootaxa 4122 (1): 579–581 http://www.mapress.com/j/zt/ Copyright © 2016 Magnolia Press Article ISSN 1175-5326 (print edition) ZOOTAXA ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.4122.1.47 http://zoobank.org/urn:lsid:zoobank.org:pub:F8B8C529-6ED9-4D54-AFD5-B8A76CC54EE2 FAMILY PLATYSTOMATIDAE LISIANE DILLI WENDT Instituto Nacional de Pesquisas da Amazônia (INPA), Coordenação de Pesquisas em Biodiversidade (CBio), Av. André Araújo, 2936, Petrópolis, CEP 69011–970, Caixa–Postal 478, Manaus, Amazonas, Brasil. Fellowship CAPES/FAPEAM. E-mail: lisidilli@yahoo.com.br Abstract Platystomatidae (Signal Flies) are one of the largest families of Tephritoidea, with about 1200 species and four subfamilies, worldwide distributed. However, Platystomatidae are not well represented in the New World, and in the Neotropical Region only four genera and 26 species, belonging to Platystomatinae, are recorded. The family is a group understudied in Colombia and only one species is recorded to the country. Key words: catalogue, Colombia, distribution, Neotropical Region, Platystomatinae Introduction Platystomatidae are recognized by small to mediam sized flies (body length 2.5–20.0mm) and body ranging from slender to very robust, generally with metallic color. Head with setae reduced in number, one or two orbital setae and frontal seta absent; vibrissa absent to rudimentary. Thorax with prosternum generally broad, bare or ciliated; postmetacoxal bridge complete or incomplete; katepisternal and presutural dorsocentral setae absent; one anepisternal seta; one or none postsutural dorsocentral setae; one to three scutellar setae. Legs moderately developed, without conspicuous setae. Wing usually patterned with dark spots or bands; humeral break present and subcostal break absent; vein Sc complete, curving forward to end in costal vein; vein R1 setulose; cell cup never with a pointed extension. Abdomen with sternite 1 present and developed. Female with segment 6 reduced or absent. Male with abdominal sternite 6 reduced or absent; lateral and medial surstily developed; one or two prensisetae; distiphallus long with developed pre-apical glans (absent in some groups, e.g. some genera of Trapherinae from Australia); terminal filaments absent (e.g. Naupoda Osten-Sacken; Scholastes Loew) or present (e.g. Lamprogaster Macquart; Loxoneuroides Hendel). Platystomatidae are one of the largest families of Tephritoidea and contains some of the most morphologically bizarre forms in this superfamily. Over 1,200 species have been described in 119 world genera and four subfamilies (McAlpine 2001). With worldwide distribution, Platystomatidae are not well represented in the New World and in the Neotropical region only Platystomatinae, with four genera and 26 species, are recorded (Steyskal 1968). The family is a group understudied in Colombia and only one species is recorded to the country (Table 1), a figure certainly requiring revision. Acronyms used for the depositories BMNH—Natural History Museum, London, England OUMNH—Oxford University Museum of Natural History, Oxford, England ZMUC—Zoological Museum University of Copenhagen, Copenhagen, Denmark Accepted by S. Nihei: 10 Apr. 2015; published: 14 Jun. 2016 579 List of abbreviations cat.—catalogue cit.—citation descr.—description disc.—discussion distr.—geographic distribution e.g.—for example figs.—figures mon.—by monotypy redescr.—redescription refs.—references spp.—species syn.—synonym, synonymy HT—Holotype F (f#)—Female M (m#)—Male Catalogue of Platystomatidae of Colombia Family Platystomatidae Schiner, 1862 Subfamily Platystomatinae Schiner, 1862 Genus Senopterina Macquart Senopterina Macquart, 1835: 453. Type species, Dacus brevipes (Fabricius) (mon.). Bricinnia Walker, 1861: 324. Type species, Bricinia flexivitta Walker (mon.). Bricinniela Giglio-Tos, 1893: 13; 1895: 45. Type species, Bricinniela cyanea Giglio-Tos (mon.). Stenopterina, misspelling. brevipes (Fabricius), 1805: 272 (Dacus). Type locality: “South America”. HT sex? (ZMUC). Distr.: Mexico, Trinidad, Guyana, Brazil, Colombia, Peru, Bolivia, Paraguay. Refs.: Wiedemann, 1830: 513 (diag. as Dacus brevipes); Macquart, 1835: 454 (diag.); Walker, 1849: 1071 (list); D’Orbigny, 1861: 552 (cit., disc.); Hendel, 1914a: 103–105 (key, redescr.); 1914b: 61 (cit., syn. list); Curran, 1934: 424 (spp. list from Kartabo); Shewell, 1962: 194 (disc., figs. 1–3, 13); Meier et al., 2005: website (collection inventory, HT); Whittington, 2010: 904 (fig. 6, head). splendens (Macquart), 1846: 337 (plate 18, fig. 8, 8a, head, wing) (Herina). HT F (OUMNH). Type locality: “Colombia”. Syn. by Hendel, 1914a: 104. angusta (Walker), 1852: 374 (Ortalis). HT M (BMNH, according to Sistema Dipterorum site). Type locality: "Brazil". Syn. by Hendel, 1914a: 104. TABLE 1. Platystomatidae species recorded to Colombia Subfamily Genus Specific epithet Author Platystomatinae Senopterina brevipes (Fabricius, 1805) References Curran, C.H. (1934) The Diptera of Kartabo, Bartica District, British Guiana. Bulletin of the American Museum of Natural History, 66, 287–532. 580 · Zootaxa 4122 (1) © 2016 Magnolia Press WENDT D’Orbigny, M.C. (1861) Dictionnaire universel d’histoire naturelle résumant et complétant tous les faits presents par les incyclopédies les anciens dictionnaires scientifiques. Chez les editeurs MM. Renard, Martinet et cie, Paris, 649 pp. Fabricius, J.C. (1805) Systema antliatorum secundum ordines, genera, species adiectis synonymis, locis, observationibus, descriptionibus. Carolum Reichard, Brunsvigae (= Brunswick), 373 + 30 pp. Giglio-Tos, E. (1893) Diagnosi di nuovi generi e di nuove specie di Ditteri. IX. Bolletino dei Musei di Zoologia ed Anatomia Comparata della Regia Universita di Torino, 8, 1–14. Giglio-Tos, E. (1895) Ditteri del Messico, Parte IV. Memorie dela Reale Accademia delle Scienze di Torino, 45, 1–73. Hendel, F. (1914a) Die Arten der Platystominen. Abhandlungen der Kaiserlich-Königlichen Zoologisch-Botanischen Gesellschaft in Wien, 8 (1), 1–410. Hendel, F. (1914b) Diptera, Fam. Muscaridae, Subfam. Platystominae. In: Wytsman, P. (Ed.), Genera Insectorum, Fasc. 157, pp. 1–179. [Bruxelles] Macquart, J. (1835) Histoire naturelle des insectes. Diptères, 2, 1–703. Macquart, J. (1846) Diptères exotiques nouveaux ou peu connus. Mémoires de la Société royale des sciences, de l'agriculture et des arts de Lille, 1844, 133–364. http://dx.doi.org/10.3853/j.0067-1975.53.2001.1327 McAlpine, D.K. (2001) Review of the Australasian genera of signal flies (Diptera: Platystomatidae). Records of the Australian Museum, 53, 113–199. http://dx.doi.org/10.3853/j.0067-1975.63.2011.1585 Meier, R., Buch, W., Buhl, P.N., Klass, K.D., Michelsen, V. & Petersen, J.F.T. (2005) Collection inventory of the Diptera collection of the Zoological Museum, University of Copenhagen (ZMUC). Available from: http://www.zmuc.dk/EntoWeb/ collections-databaser/Diptera/dipterasites.htm (accessed 31 May 2013) Steyskal, G.C. (1968) 55. Family Platystomatidae. In: Papavero, N. (Ed.), A catalogue of the Diptera of the Americas South of the United States. Departamento de Zoologia, Secretaria da Agricultura, São Paulo, 4 pp. Shewell, G.E. (1962) A new Canadian species of Stenopterina Macq. with notes on the species allied to brevipes (Fab.) (Diptera; Otitidae). The Canadian Entomologist, 94, 194–200. http://dx.doi.org/10.4039/Ent94194-2 Walker, F. (1849) List of the specimens of dipterous insects in the collection of the British Museum. Part IV. Printed by order of the Trustees [British Museum], London, pp. 689–1172. Walker, F. (1852) Diptera. Part IV. In: Saunders, W.W. (Ed.), Insecta Saundersiana: Or characters of undescribed insects in the collection of William Wilson Saunders. Vol. 1. London, pp. 253–414. Walker, F. (1861) Characters of undescribed insects in the collection of W.W. Saunders. The Transactions of the Entomological Society of London, 5, 268–334. Whittington, A.E. (2010) Platystomatidae (Signal flies). In: Brown, B.V., Borkent, A., Cumming, J.M., Wood, D.M., Woodley, N. & Zumbado, M.A. (Eds.), Manual of Central American Diptera. Vol. 2. NRC Research Press, Ottawa, pp. 903–907. CATALOGUE OF PLATYSTOMATIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 581 Zootaxa 4122 (1): 582–584 http://www.mapress.com/j/zt/ Copyright © 2016 Magnolia Press Article ISSN 1175-5326 (print edition) ZOOTAXA ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.4122.1.48 http://zoobank.org/urn:lsid:zoobank.org:pub:5C0FDE91-E459-43C5-BE75-A5BD53EEFD53 FAMILY PYRGOTIDAE RAMON LUCIANO MELLO1*& CARLOS JOSÉ EINICKER LAMAS2 1 Universidade Federal de Mato Grosso do Sul, CCBS (Zoologia), Caixa Postal 549, Cidade Universitária, 79070–900, Campo Grande, MS, Brazil. E-mail: ramon.mello@ufms.br 2 Museu de Zoologia da Universidade de São Paulo, 04218-970, São Paulo, SP, Brazil. E-mail: einicker@usp.br *Corresponding author: ramon.mello@ufms.br. Abstract Pyrgotidae is a family of endoparasitics flies of beetles with worldwide distribution. The Neotropical fauna is composed by 59 valid species names disposed in 13 genera. The occurrence of Pyrgota longipes Hendel is the first record of the family in Colombia. Key words: Catalogue, Diptera, Tephritoidea, Pyrgotidae, Colombia, Neotropical Region Introduction Pyrgotidae is a small family of tephritoid flies, whose immatures develop as internal parasitoids into the abdomen of beetles of the family Scarabaeidae (Coleoptera, Scarabaeoidea) (Forbes 1908, Aczél 1956a, Paramonov 1958, Steyskal 1987, Hernandez-Ortiz 2010). Recently, Mello et al. (2010) discussed the possible parasitoid association between the pyrgotid species Carrerapyrgota bernardii Mello, Lamas & Rafael with Pelidnota sordida Germar (Scarabaeidae, Rutelinae). The family has a worldwide distribution with the majority of the species concentrated in the tropical areas (Korneyev 2006). According to Mello & Lamas (in prep.), the Neotropical fauna is composed by 59 valid species names disposed in 13 genera. This catalogue presents for the first time the occurrence of Pyrgotidae in Colombia (Table 1). Specimens of Pyrgota longipes Hendel from Antioquia and Cundinamarca are registered. Acronyms used for the depositories CEUA—Colección Entomológica de la Universidad de Antioquia, Antioquia, Colombia. MEFLG—Museo Entomológico Francisco Luis Gallego, Medellín, Colombia. NHMW—Naturhistorisches Museum Wien. Vienna, Austria. UNAB—Museo Entomológico facultad de Agronomía, Universidad Nacional de Agronomía, Bogotá, Colombia. List of abbreviations cat.—catalogue col.—collector distr.—distribution mon.—monotypy occ.—occurrence reds.—redescription 582 Accepted by S. Nihei: 10 Apr. 2015; published: 14 Jun. 2016 refs.—references tax.—taxonomic M (m#)—Male F (f#)—Female Catalogue of Pyrgotidae of Colombia Family Pyrgotidae Schiner, 1868 Pyrgotinae Schiner, 1868: 229. Type genus, Pyrgota Wiedemann, 1830. Subfamily Pyrgotinae Schiner, 1868 Tribe Pyrgotini Schiner, 1868 Genus Pyrgota Wiedemann, 1830 Pyrgota Wiedemann, 1830: 580. Type species, Pyrgota undata Wiedemann, 1830 (mon.). Refs.: Gerstäcker 1860: 186 (tax. notes); Loew 1873: 72 (reds.); Hendel 1908: 7 (tax. notes); Hendel 1914: 78 (key); Malloch 1929: 258 (tax. notes); Hendel 1934: 143 (key); Enderlein 1942: 101 (key); Aczél 1956a: 162, 168–171, 173 (tax. notes), 181 (checklist); 1956b: 13, 17 (key), 17 (tax. notes); Steyskal 1965: 657 (cat.); Steyskal 1967: 4 (cat.); Steyskal 1978: 151 (key); Steyskal 1987: 814 (key); Hernández-Ortiz 2010: 960 (key), 960 (distr.). Oxycephala Macquart, 1843: 354. Type species, Oxycephala fuscipennis Macquart, 1843 (mon.). longipes Hendel, 1908: 145. Type locality: Brazil, Rio Grande do Sul. ST 1M, 1F (NHMW). Distr.: Brazil, Colombia. Refs.: Bezzi 1929: 2 (checklist); Hennig 1952: 609 (new occ.); Aczél 1956a: 166–169, 171, 174 (tax. notes), 181 (checklist); 1956b: 21 (reds.); Steyskal 1967: 4 (cat.). Examined material: COLOMBIA: Antioquia, Santo Domingo Porce, iii/1997, R. Véllez col., 1 M and 3 F (MEFLG); San Roque, San Jose del Nus. 880 m. Bosque, 13/iiv/2011, A. Clavijo col., 1 F (CEUA). Cundinamarca, La Palma, ix/2009, J. Guerrero col., 1 M (UNAB); x/2009, 1 M (UNAB); Tibacuy, La Vuelta, 20/ix/2009, R. Simbaqueba col., 1 M (UNAB); Fusagasuga, 11/vi/1996, D. Forero col., 1 F (UNAB). Acknowledgments The authors are very grateful to the curators Erika Navarro (UNAB) and John Gamboa (MEFLG) for the loan of material and to Dr. Marta Wolff (CEUA) by sending information about the collection; and to the Conselho Nacional de Desenvolvimento Científico e Tecnológico (CNPq) for the PhD fellowship of the first author (process nº. 141656/2007–70). TABLE 1. Pyrgotidae species recorded from Colombia. Tribe Genus Species Author Pyrgotini Pyrgota longipes Hendel, 1908 References Aczél, M. (1956a) Revisíon parcial de las Pyrgotidae neotropicales y Antárticas con synopsis de los géneros y especies (Diptera, Acalyptratae). Revista Brasileira de Entomologia, 4, 161–184. Aczél, M. (1956b) Revisíon parcial de las Pyrgotidae neotropicales y Antárticas con synopsis de los géneros y especies (Diptera, Acalyptratae). Revista Brasileira de Entomologia, 5, 1–70. Bezzi, M. (1929) Australian Pyrgotidae. Proceedings of the Linnean Society of New South Wales, 54, 1–20. Enderlein, G. (1942) Klassifikation der Pyrgotiden. Sitzungsberichte der Gesellschaft Naturforschender Freunde, 2, 98–134. CATALOGUE OF PYRGOTIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 583 Forbes, S.A. (1908) On the life history, habits, and economic relations of the white-grubs and may-beetles (Lachnosterna). Bulletin of the Illinois Agricultural Experimental Station, 116, 447–480. Gerstäcker, F. (1860) Beschreibung einiger aus gezeichneten neuen Dipteren aus der Familie Muscariae. Stettiner Entomologische Zeitung, 21, 163–202. Hendel, F. (1908) Diptera, Familien Muscaridae, subfamilien Pyrgotinae. Genera Insectorum, 79, 1–33. Hendel, F. (1914) Neue Beiträge zur Kenntnis der Pyrgotinen. Archiv für Naturgeschichte, 79, 77–117. Hendel. F. (1934) Übersicht über die Gattungen der Pyrgotiden, nebst Beschreibung neuer Gattungen und Arten. Encyclopédie Entomologique, Diptera, 7, 141–156. Hennig, W. (1952) Die Larvenformen der Dipteren. Vol. 3. Akademi Verlag, Berlin, 628 pp. Hernández-Ortiz, V. (2010) Pyrgotidae (Pyrgotid flies). In: Brown, B.V., Borkent, A., Cumming, J.M., Wood, D.M., Woodley, N.E., & Zumbado, M.A. (Eds.), Manual of Central American Diptera. Vol. 2. National Research Council Press, Ottawa, pp. 955–961. Korneyev, V. (2006) A revision of Afrotropical species of the Eupyrgota (Diptera, Pyrgotidae): the spinifemur group and latipennis subgroup of species. Vestnik Zoologii, 40, 3–25. Loew, H. (1873) Monographs of the Diptera of North America part III. Smithsonian Miscellaneous Collections, 256, 1–351. Macquart, J. (1843) Dipteres exotiques nouveaux ou peu conns. Supplément I Memoires de La Société Royale des Sciences, de l’Agriculture et des Arts, de lille, 1844, 133–364. Malloch, J.R. (1929) Exotic Muscaridae (Diptera); XXIV. Family Pyrgotidae. Annals and Magazine of Natural History, 3, 240–261. Mello, R.L., Lamas, C.J.E. & Rafael, J.A. (2010) Revision of the Neotropical genus Carrerapyrgota Aczél (Diptera, Pyrgotidae) with the description of two new species. Zootaxa, 2515, 45–64. Paramonov, S.J. (1958) A review of Australian Pyrgotidae (Diptera). Australian Journal of Entomology, 6 (1) , 89–138. http://dx.doi.org/10.1071/zo9580089 Schiner, J.R. (1868) Diptera. In: von Wüllerstorf-Urbair, B. (Ed.), Reise der österreichischen Fregatte Novara um die Erde in den Jahren 1857, 1858, 1859 unter den Befehlen des Commodore B. von Wüllerstorf-Urbair. Zoologischer Theil. Vol. 2. Abt. I. K. Gerold’s Son, Wien, pp. 1–388. Steyskal, G.C. (1965) Family Pyrgotidae. In: Stone, A., Sabrosky, C.W., Wirth, W.W., Foote, R.H. & Coulson, T.R. (Eds.), A catalog of the Diptera of America north of Mexico. USDA, Agricultural Research Service, Agriculture Handbooks 276, pp. 657–658. Steyskal, G.C. (1967) Family Pyrgotidae. In: Papavero, N. (Ed.), A catalogue of the Diptera of the America South of the United States. Museu de Zoologia, São Paulo, 56, pp. 1–8. Steyskal, G.C. (1978) Synopsis of the North American Pyrgotidae (Diptera). Proceedings of the Entomological Society of Washington, 80 (2), 149–155. Steyskal, G.C. (1987) Pyrgotidae. In: McAlpine, J.F., Peterson, B.V., Shewell, G.E., Teskey, H.J., Vockeroth, J.R., Wood, D.M. (Eds.), Manual of Nearctic Diptera. Vol. 2. Research Branch Monograph 28, Agriculture Canada, Ottawa, pp. 813–816. Wiedemann, C.R.W. (1830) Aussereuropäische zweifluegeliche Insekten. Vol. 2. Schulzischen Buchhandlung, Hamm, xii, 684 pp. 584 · Zootaxa 4122 (1) © 2016 Magnolia Press MELLO & LAMAS Zootaxa 4122 (1): 585–595 http://www.mapress.com/j/zt/ Copyright © 2016 Magnolia Press Article ISSN 1175-5326 (print edition) ZOOTAXA ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.4122.1.49 http://zoobank.org/urn:lsid:zoobank.org:pub:D67463D4-37F8-47CA-9C98-8261E5C0D863 FAMILY RICHARDIIDAE LISIANE DILLI WENDT1* & ROSALY ALE-ROCHA1, 2 1 Instituto Nacional de Pesquisas da Amazônia (INPA), Coordenação de Pesquisas em Biodiversidade (CBio), Av. André Araújo, 2936, Petrópolis, CEP 69011–970, Caixa–Postal 478, Manaus, Amazonas, Brasil. Fellowship CAPES/FAPEAM; E-mail: lisidilli@yahoo.com.br; 2 Fellowship PQ/CNPq; E-mail: alerocha@inpa.gov.br * Corresponding author: lisidilli@yahoo.com.br Abstract Richardiidae are a family of “acalyptrate” Diptera represented by ca. 180 species distributed in the New World, mostly in the Neotropical region. The species that occur in Colombia have received little attention from taxonomists, and the great majority of them are known only from their type localities. Currently, 14 genera and 23 species are known to occur in the country. Key words: catalogue, Colombia, distribution, Neotropical Region, Richardiinae Introduction The following characteristics are peculiar to the Richardiid flies: small to medium-sized (body length 3–15mm), body robust or slender. Head with one orbital seta and frontal seta absent. Thorax with prosternum very reduced, bare or ciliated; katepisternum developed, higher than anepisternum; postmetacoxal bridge complete or incomplete; presutural dorsocentral seta absent; one anepisternal seta; one or two post-sutural dorsocentral setae; one or two scutellar setae. Wing generally with apical spot, median band and/or other marks; subcostal break present; humeral break present in Epiplateinae and absent in Richardiinae; R1 vein swollen at apex; cup closed apically by a short sinuous or slightly convex vein. Legs long, slender to robust, usually banded; spine-like setae present on ventral surface of at least hind femur in Richardiinae (except Setellia Robineau-Desvoidy), absent in all femora in Epiplateinae. Abdomen with strong preapical setae laterally on syntergite 1+2; sternite 1 very reduced or absent. Female with two spermathecae. Male with abdominal tergites 6 and 7 very reduced or vestigial; lateral surstylus partial or entirely free from epandrium; one (in Richardiinae) or two (in Epiplateinae) prensisetae; distiphallus long and generally setulose. Very little is known about the biology of immature and adult richardiids. A few records on the breeding behavior of these flies suggest that the larval diet is composed of rotting vegetable matter, rotting fruit (Steyskal 1958, Martinez et al. 2000), decaying trunks (Steyskal 1973), and flowers (Steyskal 1973, 1987). Shannon, Suspended and Malaise are optimal traps for collecting the adult stage. Furthermore, there are records of adult richardiids collected in association with animal carcasses (e.g. Carvalho et al. 2000, Carvalho & Linhares 2001, Moretti et al. 2008). The family Richardiidae is classified in the Tephritoidea and only occurs in the New World, mostly in the Neotropical region. The family is currently represented by about 200 species in 32 genera and is divided into two subfamilies, Richardiinae and Epiplateinae (Hancock 2010). The Richardiinae is conspicuously more diverse, encompassing more than 90% of the species in the family (Steyskal 1968). The species that occur in Colombia have received little attention from taxonomists, and the great majority of them are known only from their type localities. Currently, 14 genera and 23 species are known to occur in the country (Table 1). The present catalogue provides a list of species from Colombia based on available literature and observed material. For genus name the following is provided: the original reference, type species, current valid name (when Accepted by S. Nihei: 10 Apr. 2015; published: 14 Jun. 2016 585 appropriate), designation, and synonyms listed chronologically. For each species name, the following information is given: the original reference with the original generic assignment when it differs from the present assignment, the type locality quoted in full, the nature and location of the type series, the geographical distribution in a sequence that runs from north to south and west to east, chronologically listed synonyms, all known available and unavailable published names, and bibliography of all literature cited. Acronyms used for the depositories ANSP—Academy of Natural Sciences, Pennsylvania, Philadelphia, USA BMNH—Natural History Museum, London, England CE-TDEA—Colección Entomológica Tecnológico de Antioquia, Institución Universitária, Medellín, Colombia CEUA—Colección Entomológica, Universidad de Antioquia, Medellín, Colombia DEIC—Deutsches Entomologisches Institut, Müncheberg, Germany HNHM—Hungarian Natural History Museum, Budapest, Hungarian MCZ—Museum of Comparative Zoology, Harvard University, Cambridge, Massachusetts, USA MEFLG—Museo Entomológico Francisco Luís Gallego, Medellín, Colombia MIZT—Museo Zoologico di Torino, Università di Torino, Torino, Italy MLPA —Universidad Nacional de La Plata, Museo de la Plata, La Plata, Argentina MNHN—Muséum National d'Histoire Naturelle, Paris, France MRSN—Museo Regionale di Scienze Naturali, Torino, Italy MZPW—Polish Academy of Science, Museum of the Institute of Zoology, Warsaw, Poland NMHW—Naturhistorisches Museum Wien, Wien, Austria OUMNH—Oxford University Museum of natural History, Oxford, England USNM—National Museum of Natural History, Smithsonian Institution, Washington, D.C., USA ZMHB—Museum für Naturkunde der Humbolt-Universität zu Berlin, Berlin, Germany ZMUC—Zoological Museum University of Copenhagen, Copenhagen, Denmark List of abbreviations cat.—catalogue cit.—citation col.—collector comb.—combination des.—designation descr.—description diag.— diagnosis disc.—discussion distr.—geographic distribution e.g.—for example n., nov.—new mon.—monotypy orig. des.—by original designation pp.—pages redescr.—redescription refs.—references rev.—revision sp., spp.—species subs.—subsequent syn.—synonym, synonymy HT—Holotype 586 · Zootaxa 4122 (1) © 2016 Magnolia Press WENDT & ALE-ROCHA PT—Paratype ST—Syntype T—Type (unspecified) F (f#)—Female M (m#)—Male Catalogue of Richardiidae of Colombia Family Richardiidae Snow, 1896 Subfamily Epiplateinae Steyskal, 1987 Genus Automola Loew Automola Loew, 1873: 118; Type species, Ortalis atomaria Wiedemann, 1830: 461; subs. des. (Coquillett, 1910: 512). Wulpia Bigot, 1886: 371. Type species, Wulpia caloptera Bigot (mon.). Syn. by Hendel, 1911a: 184. atomaria (Wiedemann), 1830: 461 (Ortalis). Type locality: “Brazil”. ST M (NHMW). Distr.: Mexico, Costa Rica, Colombia (Caldas (Norcasia)) (new record), Guyana, Brazil, Peru, Bolivia, Paraguay, Argentina. Refs.: Loew, 1873: 11, 118 (disc. as Ortalis trifasciata; cit.); Wulp, 1898: 390 (material list, disc.; plate 10, fig. 22, wing); Aldrich, 1905: 588 (cat.); Coquillett, 1910: 512 (subs. des.); Hendel, 1911a: 184–187 (key, redescr.); Curran, 1934a: 293, 425 (spp. list, material list); 1934b, 1965: 276 (fig. 34, wing); Hennig, 1937: 22 (material list); Aczél, 1950: 8 (cat.); Steyskal, 1958: 303 (key, disc., material list, distr.); 1968: 2 (cat.); Wendt & Ale-Rocha, 2012: 49 (cit.); 2013: 1091–1093 (cit., comments, distr., distr. map). trifasciata (Wiedemann), 1830: 461 (Ortalis). Type locality: “Brazil” ST F (NHMW). Syn. by Hendel, 1911a: 185. Refs.: Loew, 1868: 324 (cit. as Ortalis); 1873: 11 (disc., as Ortalis trifasciata), 118 (cit.); Wulp, 1898: 390 (cit.); Hendel, 1911a: 185 (syn.); Aczél, 1950: 8 (cat.); Steyskal, 1968: 2 (cat.). mexicanum (Giglio-Tos), 1893: 12 (Platystoma); 1895: 46 (fig. 24, wing). HT M (MIZT or MRSN ?). Type locality: Mexico, Tuxpango. Syn. by Hendel, 1911a: 185. Refs.: Hendel, 1911a: 185 (syn.); Aczél, 1950: 8 (cat.); Steyskal, 1968: 2 (cat.). leopardina Blanchard, 1938: 92 (figs. 6, 9). Type locality: Argentina, Misiones, Porto Bemberg. HT F (MLPA). Syn. by Steyskal, 1958: 303. Refs.: Aczél, 1950: 8 (cat.); Steyskal, 1958: 303 (syn.); 1968: 2 (cat.). Material examined. COLOMBIA: Caldas: Norcasia, Sector El Tigre, 05º33’09.8”N 74º52’23.1”W, 700m, Forest, 1 female, 20.ii.2010, D.Y. Gaurisas and L. Rios col., Van Sommeren-Rydon trap (CEUA). Subfamily Richardiinae Snow Genus Batrachophthalmum Hendel Batrachophthalmum Hendel, 1911a: 394. Type species, Batrachophthalmum rufiventri Hendel (by mon.). quimbaya Carvalho, Wolff & Wendt, 2011: 44. Type locality: Colombia, Risaralda, Pereira, Otún Quimbaya. HT M, PT M (CEUA). Distr.: Colombia (Risaralda (Pereira)). Refs.: Wendt & Ale-Rocha, 2014: 306–310 (female desc., figs. 1A–F, 2A–K, 3A–I, female habitus, head, thorax, terminalia, sternites, spermathecae; male terminalia and sternites). Material examined. COLOMBIA: Risaralda: Pereira, Santuario de Flora y Fauna Otún-Quimbaya, HT and PT 1 male, v. 2001, Grupo de Entomologia col. (CEUA) 12796; idem, 1890m, 1 female, 07.v.2001, Bosque Nativo, Van Sommeren-Rydon trap baited with fish (CEUA) 16283. CATALOGUE OF RICHARDIIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 587 Genus Coilometopia Macquart Coilometopia Macquart, 1847: 107. Type species, Coilometopia ferruginea Macquart (by mon.) = Scatophaga trimaculata Fabricius, 1805. costalis Hendel, 1911a: 256. HT F (NMHW). Type locality: “Venezuela”. Distr.: Colombia (Santander (Piedecusta Sevilla)) (new record), Venezuela, Brazil, Bolivia. Refs.: Hennig, 1937: 32 (material list); Aczél, 1950: 22 (cat.); Steyskal, 1968: 3 (cat.). Material examined. COLOMBIA: Santander: Piedecuesta Sevilla, Vereda Cristales, Rasgon Farm, Forest, 2200m, 1 female, 25.viii.2001, Duque & Vélez cols., Van Sommeren-Rydon Trap (CEUA). longicornis Hendel, 1911a: 255. Type locality: Peru, Vilcanota. ST M (HNHM). Distr: Colombia (Quindio (Quimbaya)) (new record); Brazil, Peru, Bolivia. Refs.: Hennig, 1937: 32 (material list); Aczél, 1950: 23 (cat.); Steyskal, 1968: 3 (cat.). Material examined. COLOMBIA: Quindío: Quimbaya, Vereda Kerman, Km 7, Panaca, 1 male, 01.vii.2003, H. Vargas col. (CEUA). trimaculata (Fabricius), 1805: 208 (Scatophaga). Type locality: “South America”. HT sex? (ZMUC). Distr.: Costa Rica, Panama, Trinidad, Guyana, Colombia (Antioquia (La Pintada, Maceo, Caucásia), Sucre (San Onofre), Meta (San Martin)) (new records), Venezuela, Brazil, Ecuador, Peru, Bolivia. Refs.: Loew, 1873: 13, 68, 188 (disc. as Coilometopia), 26 (cit. as Trypeta), 68 (cit. as Scatophaga); Coquillett, 1910: 526 (cit. as Scatophaga); Hendel, 1911a: 254–255 (key, taxonomy); 1911b: 24 (spp. list, figs. 40–42, head, wing); 1936: 80 (material list); Enderlein 1912: 113 (material list); Curran, 1934a: 293, 423 (spp. list, material list, human excrement); Hennig, 1937: 32 (material list); Aczél, 1950: 23 (cat.); Steyskal, 1968: 3 (cat.); Meier et al., 2005: website (collection inventory, HT); Hancock, 2010: 876, 878 (fig. 19, head, record to Costa Rica). flavus (Fabricius), 1805: 274 (Dacus). Type locality: “South America”. HT sex? (ZMUC). Syn. by Hendel 1911a: 154. Refs.: Loew, 1873: 68 (disc. as Dacus); Hendel, 1911a: 254 (syn.); 1911b: 24 (spp. list); Steyskal, 1968: 3 (cat.); Meier et al., 2005: website (collection inventory, HT). maculipennis (Macquart), 1846: 339 (plate 18, fig. 12, habitus dorsal) (Odontomera). Type locality: “Colombia”. HT F (OUMNH, (Fairmaire collection)). Syn. by Loew, 1873: 68. Refs.: Schiner, 1868: 256, 258 (disc. as Odontomera); Loew, 1873: 68 (disc. as Odontomera); Hendel, 1911a: 254 (cit. as Odontomera); 1911b: 24 (spp. list); Aczél, 1950: 23 (cat. as Odontomera); Steyskal, 1968: 3 (cat.). ferruginea Macquart, 1847: 107 (plate 5, figs. 14a–c, head, antenna). Type locality: “Brazil”. HT M (OUMNH, (Bigot Collection)). Syn. by Loew, 1873: 188. Refs.: Loew, 1873: 68, 188 (disc., syn.); Hendel, 1911b: 24 (spp. list); Aczél, 1950: 23 (cat.); Steyskal, 1968: 3 (cat.). Material examined. COLOMBIA: Antioquia: Maceo, Reserva Cañon del rio Alicante, Forest, 400m, 2 males, 1 female, 25.i.2009, M. Wolff col. (CEUA); La Pintada, Montenegro Farm, Comfenalco, 5º43’24”N 75º37’15”W, Bosque, 850m, 1 female , 20.iv.2007, N. Uribe col. (CEUA); Caucasia, Sector Las Malvinas, La Cacaotera Farm, 07º58’06.05”N 75º11’54.52”, 3 males, 20.iv.2010, Entomology Group col., Van Sommeren-Rydon trap baited with fish (CEUA); Sucre: San Onofre, Sanguaré Reserve, Palmavinal, 09º42’36.80”N 75º41’00.38”W, 15m, 2 males, 2 females, 26–29.v.2009, Entomology Group col., Van Sommeren-Rydon trap baited with fish (CEUA); Meta: San Martin, Vereda La Novilla, 03º31’37”N 73º24’16”W, 249m, 7 males, 5 females, 06.vi.2013, Perez and Gomez-Pinerez col., Van Sommeren-Rydon trap baited with fish and offal (CE-TDEA). Genus Euolena Loew Euolena Loew, 1873: 183. Type species, Michogaster egregius Gerstaecker (by orig. des.). egregia (Gerstaecker), 1860: 179 (Michogaster). Type locality: “Colombia”. ST M, F (ZMHB). Distr.: Colombia, Venezuela, Ecuador. Refs.: Schiner, 1868: 255 (cit. as Michogaster); Hendel 1911a: 390 (diag.); 1911b: 42 (spp. list, figs. 64, 65, habitus, head); Enderlein, 1912: 109 (material list); Hennig, 1938a: 118 (material list); Aczél, 1950: 40 (cat.); Steyskal, 1968: 4 (cat.); Hancock, 2010: 875, 878 (fig. 15, habitus, cit.). 588 · Zootaxa 4122 (1) © 2016 Magnolia Press WENDT & ALE-ROCHA ornatus Schiner, 1868: 255 (Michogaster). Type locality: “South America”. ST M, F (NMHW). Syn. by Hendel, 1911b: 42. Refs.: Hendel, 1911b: 42 (syn.); Steyskal, 1968: 4 (cat.). Material examined. “Columbia”, syntypes 1 male, 4 females, Moritz col. (ZMHB). Genus Hemixantha Loew Hemixantha Loew, 1873: 190. Type species, Hemixantha spinipes Loew (by mon.). fascinventris Enderlein, 1912: 112. Type locality: Colombia. HT F (MZPW). Distr.: Colombia, Brazil. Refs.: Lopes, 1936: 476 (disc.); Hennig, 1937: 308 (key, cit.); Aczél, 1950: 25 (cat.); Steyskal, 1968: 4 (cat.). fasciventris Hendel, 1936: 80. Type locality: Brazil, Pará, Taperinha. HT M (MNHW). Syn. by Aczél 1950: 25. Refs.: Aczél, 1950: 25 (cat.); Steyskal, 1968: 4 (cat.). Note: Both this and the aforementioned species were described as valid species. Aczél (1950) and Steyskal (1968) treated them as synonymous, however the examination of types is necessary to clarification on status of both species. Genus Melanoloma Loew Melanoloma Loew, 1873: 192. Type species, Trypeta cyanogaster Wiedemann (by orig. des.). Melanolema Enderlein, 1912: 112 (misspelling). Melanolomina Curran, 1934b: 281. Type species, Odontomera varians Schiner (by orig. des.). decrepita Hendel, 1911a: 367. Type locality: “Venezuela”. HT F (NMHW). Distr.: Colombia, Venezuela, Peru, Bolivia. Refs.: Hendel, 1911b: 31 (spp. list); Enderlein, 1912: 112 (material list); Curran, 1934b, 1965: 283 (figs. 98, 107, wing, head); Hennig, 1938a: 112 (material list); Aczél, 1950: 31 (cat.); Steyskal, 1968: 6 (cat.). viatrix Hendel, 1911a: 269. Type locality: Bolivia, Songo. HT F (HNHM). Distr.: Colombia (Antioquia (Vegachi), (Maceo)) (new record), Venezuela, Bolivia. Refs.: Hennig, 1938a: 112 (material list); Aczél, 1950: 30 (cat.); Steyskal, 1968: 6 (cat.); Arévalo & Osório, 1995: 1–8 (biol.); Martínez et al., 2000: 135–140 (biol., pest, figs. 1–3, adult, larvae, pupa); Montilla et al., 2007: 107–112 (biol.); 2008: 403–407 (distr. in Venezuela); Moreno et al., 2009: 98–100 (biol., figs. 1A–L, pupae and puparium); Hancock, 2010: 878 (cit.). Material examined. COLOMBIA: Antioquia: Vegachi, 14 males, 6 females, ii.1992, Yepes col., on pineapple fruit (MEFLG), idem, 3 males, 6 females, iii.1992, idem , idem (MEFLG); Maceo, 1 male, 3 females, vii.1992, idem (MEFLG). Genus Odontomera Macquart Odontomera Macquart, 1843: 372. Type species, Odontomera ferruginea Macquart (by mon.). Cyrtometopa Loew, 1873: 179. Type species, Odontomera ferruginea Macquart (by orig. des.). canonigra Enderlein, 1912: 110. Type locality: “Colombia”. HT F (MZPW). Distr.: Colombia. Refs.: Hendel, 1936: 79 (disc.); Hennig, 1938a: 114 (material list); Aczél, 1950: 33 (cat.); Steyskal, 1968: 7 (cat.). nitens (Schiner), 1868: 258 (Michogaster). Type locality: “Colombia”. ST M (NMHW). Distr.: Mexico, Costa Rica, Colombia, Venezuela, Bolivia. Refs.: Hendel, 1911a: 370, 372 (key, redescr.); 1911b: 34 (spp. list, figs. 54–56, head, wing); Curran, 1934b, 1965: 282–283 (figs. 80, 97, head, wing); Hennig, 1938a: 114 (material list); Aczél, 1950: 35 (cat.); Steyskal, 1958: 304, 310 (cit., male terminalia descr., figs. 6, 7, male terminalia); 1968: 7 (cat.); Korneyev, 1999: 22 (phylogeny of Tephritoidea, ingroup); Becerril-Morales & Macías-Ordóñez, 2009: 245–262 (behavior); Hancock, 2010: 878 (cit.). cinctella (Wulp), 1898: 389 (figs. 20, 20a, habitus, head) (Cyrtometopia). Type locality: Mexico, Tabasco, Teapa, São Lorenzo near Cordova. ST M, F (USNM). Syn. by Hendel, 1911b: 34. Refs.: Wulp, 1898: 388 (key, CATALOGUE OF RICHARDIIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 589 Cyrtometopia); Aldrich, 1905: 599 (cat. as Cyrtometopia); Hendel, 1911a: 372 (cit. as Cyrtometopia), 1911b: 34 (syn.); Aczél, 1950: 35 (cat.); Steyskal, 1968: 7 (cat.). Genus Poecilomyia Hendel Poecilomyia Hendel, 1911a: 183, 256. Type species, Poecilomyia longicornis Hendel (by mon.). longicornis Hendel, 1911a: 256. Type locality: “Venezuela” and Panama, Chiriqui. ST M, F (NHMW). Distr.: Panama, Colombia, Venezuela, Peru, Bolivia. Refs.: Hendel, 1911b: 24 (spp. list, figs. 43–45, head, wing); Curran, 1934b, 1965: 278, 282 (figs. 49, 93, wing, head); Hennig, 1937: 33 (material list); Aczél, 1950: 24 (cat.); Steyskal, 1968: 8 (cat.); Hancock, 2010: 876, 878 (fig. 18, head, cit.). longicornis var. colombiana Enderlein, 1912: 106. Type locality: Colombia, Fusagasuga. HT M (MZPW). Distr.: Colombia. Refs.: Hennig, 1937: 33 (cit.); Aczél, 1950: 24 (cat.); Steyskal, 1968: 8 (cat.). Genus Richardia Robineau-Desvoidy Richardia Robineau-Desvoidy, 1830: 728. Type species, Richardia saltatoria Robineau-Desvoidy (by mon.). Merodina Macquart, 1835: 378. Type species, Dacus podagrica Fabricius (1805) (by mon.). Gnathoplasma Enderlein, 1912: 99. Type species, Gnathoplasma infestans Enderlein (by orig. des.). Scelidodontia Enderlein, 1913: 553. Type species, Scelidodontia hendeliana Enderlein (by orig. des.). Polphopsis Enderlein, 1927: 127. Type species, Richardia telescopica Gerstaecker (by orig. des.). Zetekomyia Curran, 1934b: 275. Type species, Zetekomyia banski Curran, nomen nudum (by orig. des.) = Richardia telescopica Gerstaecker. Richarula Williston, 1908: 279 (misspelling). annulata (Macquart), 1835: 434 (Herina). Type locality: “Colombia”. HT M (MNHN). Distr.: Panama, Colombia, Venezuela, Guyana Brazil. Refs.: Macquart, 1843: 362 (cit. as Herina); Rondani, 1848: 82 (disc. as Herina); Gerstaecker, 1860: 163 (cat.); Schiner, 1868: 259 (disc., cit. as Herina.); Hendel, 1911a: 208, 242 (key, redescr.); 1911b: 21 (spp. list); Speiser, 1911: 246 (disc.); Hennig, 1937: 28 (cit.); Aczél, 1950: 16 (cat.); Steyskal, 1968: 9 (cat.). eurycephala Gerstaecker, 1860: 165. Type locality: “South America”. HT M (ZMHB). Syn. by Hendel, 1911b: 21. Refs.: Schiner, 1868: 259 (disc.); Hendel, 1911b: 21 (syn.); Aczél, 1950: 16 (cat.); Steyskal, 1968: 9 (cat.). flavipes Schiner, 1868: 260. Type locality: “South America”. HT M (NHMW). Distr.: Colombia (Antioquia (Puerto Berrio), Quindío (Quimbaya)) (new records), Guyana, Venezuela. Refs.: Hendel, 1911a: 210 (key), 1911b: 21 (cit.; plate 2, fig. 32, hind leg); Curran, 1934a: 293, 421, 423 (spp. list, key, material list, human excrement); Hennig, 1937: 30 (cit.); Aczél, 1950: 17 (cat.); Steyskal, 1968: 9 (cat.); Alencar et al., 2013: 205, 207 (cit., key to Brazilian Amazon spp., fig. 19, hind leg). Material examined. COLOMBIA: Antioquia: Puerto Berrio, Vereda Cristalina, Reserva RUNA, Bosque, 1 male, 24.ix.2005, Entomology Group col., Pitfall trap baited with fruit (CEUA); Quindío: Quimbaya, 1200m, 1 male, 25.vi.2003, H. Vargas col., Jama Solar (CEUA). hendeliana (Enderlein), 1913: 553 (fig. hind femora) (Scelidodontia). Type locality: Colombia, Hacienda Pehlke. HT M (MZPW). Distr.: Colombia. Refs.: Hennig, 1937: 30 (cit.); Aczél, 1950: 17 (cat.); Steyskal, 1968: 9 (cat.). infestans (Enderlein), 1912: 99 (Gnathoplasma). Type locality: Colombia, Hacienda Pehlke. HT M (MZPW). Distr.: Panama, Colombia. Refs.: Hennig, 1937: 31 (cit. as Gnathoplasma); Aczél, 1950: 22 (cat. as Gnathoplasma); Steyskal, 1968: 9 (cat., comb. nov.); Alencar et al., 2013: 202 (cit.). Wendt & Ale-Rocha, 2015: 158, 160 (key, diag., disc., figs. 3, 20–23, habitus, head). Material examined. COLOMBIA: Pehlke's Farm, HT male, iv-vi.1908, E. Pehlke col. (MAPW); Cordillera, Tierra Caliente, 1 male, without date, Thieme col., (ZMHB). 590 · Zootaxa 4122 (1) © 2016 Magnolia Press WENDT & ALE-ROCHA podagrica (Fabricius), 1805: 272 (Dacus). Type locality: “South America”. HT sex? (ZMUC). Distr.: Mexico, Costa Rica, Trinidad, Colombia (Antioquia (Gómez Plata), (San Jeronimo), Quindío, (Quimbaya)) (new records), Guyana, Brazil, Peru, Bolivia. Refs.: Wiedemann, 1830: 445 (as Cordylura podagrica, diag., plate 10, figs. 9a, b, c, habitus, palpus, antenna); Macquart, 1843: 362 (disc.); Rondani, 1848: 82 (disc.); Gerstaecker, 1860: 163 (cat.); Schiner, 1868: 259 (disc.); Wulp, 1898: 385 (key, diag.); Aldrich, 1905: 599 (cat.); Hendel, 1911a: 209, 210 (key, redescr.); 1911b: 21 (spp. list, figs. 30, 31, habitus, head); Speiser, 1911: 247 (disc.); Enderlein, 1912: 101 (fig. 3, fore leg); Cole, 1927: 444 (disc. about genitalia, fig. 251, male genitalia); Curran, 1934a: 293, 421 (spp. list, key, material list); Hennig, 1937: 25 (material list); Aczél, 1950: 19 (cat.); Steyskal, 1958: 310 (male terminalia descr., figs, 19–21, male terminalia); 1968: 9 (cat.); Korneyev, 1999: 22 (phylogeny of Tephritoidea, ingroup); Meier et al., 2005: website (collection inventory, HT). Material examined. COLOMBIA: Antioquia: Gómez Plata, Vereda La Clara, 1000m, 1 male, 09.vii.2003, Entomology Group col., Jama (CEUA); San Jeronimo, Parque Los Tamarindos, Sendero Ecológico, Rastrojo, 925m, 1 female, 29.xi.2005, A. Velez cols., Van Sommeren-Rydon trap baited with fish (CEUA); idem, La Pintada, Montenegro Farm, Comfenalco 5º43’24”N 75º37’15”W, 859m, Forest, 1 male, 1 female, 20.iv.2007, N. Uribe col. (CEUA); Quindío: Quimbaya, Vereda Kerman, Km7, Panaca, 1 male, 01.vii.2003, H. Vargas col. (CEUA). stylops Hennig, 1938b: 8 (figs. 1, 2, head, wing). Type locality: Colombia, Muzo. HT M (DEIC). Distr.: Colombia (Antioquia (San Luis, Maceo, Sant Carmen de Chucuri, Sant Pie de Cuesta)) (new record). Refs.: Aczél, 1950: 21 (cat.); Steyskal, 1968: 10 (cat.); Carvalho et al., 2011: 41 (key); Alencar et al., 2013: 202 (cit.). Material examined: COLOMBIA: Alto de los Andes, 2200m, 1 male, 1 female, 8.x.1908, (BMNH); Aguatal, 1 male, without date and collector (BMNH); Antioquia: San Luis, Proyecto Porvenir 1, Quebrada, 400m, 1 male, 22–26.viii.2011, S. Angel and C. Bota cols. (CEUA); Maceo,Vereda San Pedro, 06º27’N 74º47’W, 890–1100m, Forest, 1 male, 17–19.ii.2012, Entomology Group col., Van Sommeren-Rydon trap baited with fish (CEUA); Santa Carmen de Chucuri, Vereda Cascajales, Forest, 1 male, 19–21.xi.2000, A. Vélez and P. Duque cols., Van Sommeren-Rydon trap baited with fish (CEUA); Pie de Cuesta, Vereda Cristalina, Forest, 2300–2400m, 1 male, 16.viii.2001, P. Duque and A. Vélez cols.,Van Sommeren-Rydon trap baited with fish (CEUA). telescopica Gerstaecker, 1860: 167 (plate 2, 2a, habitus, head); Type locality: Brazil, Amazon River. HT M (ZMHB). Distr.: Nicaragua, Costa Rica, Panama, Colombia (Valle del Cauca (Buenaventura)) (new record), Brazil. Refs.: Schiner, 1868: 260 (disc.); Hendel, 1911a: 209 (key); Hennig, 1937: 30 (cit. as Polyphopsis); Aczél, 1950: 21 (cat.); Steyskal, 1968: 10 (cat.); Grimaldi & Fenster, 1989: 2 (table hipercephalic Diptera, figs. 1a–c, female head, male habitus, male head); Zumbado, 2006: 151, 152, 255 (figs. male and female, habitus); Carvalho et al., 2011: 42–43 (cit., key); Alencar et al., 2013: 202, 206 (cit., key to Brazilian Amazon spp.). huberi (Cresson), 1923: 259 (Plagiocephalus). Type locality: Nicaragua, Eden. HT M (ANSP). Refs.: Aczél, 1950: 21 (cat.); Steyskal, 1968: 10 (cat.). banski (Curran), 1934b, 1965: 281 (Zetekomyia) (figs. 42, 105, wing, head) (without descriptive statement). Syn. by Steyskal, 1968. Refs.: Aczél, 1950: 32 (cat.); Steyskal, 1968: 10 (cat., syn.). Material examined. COLOMBIA: Valle del Cauca: Buenaventura, Rio Yurumanguí, Guandal, 2 males, 31.i.1998, F. Riascos col. (MEFLG). undulata Hennig, 1937: 27. Type locality: Colombia, Cesar, Chiriguana District, Lake Sapatoza Region. HT M (BMNH). Distr.: Colombia (Cesar (Chiriguana)). Refs.: Aczél, 1950: 20 (cat.); Steyskal, 1968: 10 (cat.). viridiventris Wulp, 1898: 386. Type locality: Mexico, Tabasco, Teapa. ST M, F (USNM); ST M (MCZ). Distr.: Mexico, Costa Rica, Surinam, Colombia, Guyana, Brazil, Peru, Bolivia. Refs.: Aldrich, 1905: 599 (cat.); Hendel, 1911a: 209, 212 (key, redescr.); 1911b: 21 (spp. list, fig.35, wing); Hennig, 1937: 27 (cit.); Aczél, 1950: 20 (cat.); Steyskal, 1968: 10 (cat.). Genus Sepsisoma Johnson Sepsisoma Johnson, 1900: 327. Type species, Michogaster geniculatus Schiner (by orig. des.). Parasespsisoma Hendel, 1936: 80. Type species, Sepsisoma flavescens Johnson (by mon.). CATALOGUE OF RICHARDIIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 591 geniculatum (Schiner), 1868: 257 (Michogaster). Type locality: “South America”. ST M (NMHW). Distr.: Colombia, Venezuela, Brazil. Refs.: Hendel, 1911a: 379, 381 (key, redescr.); 1911b: 17 (spp. list, figs. 58, 59, head); 1936: 80 (material list, note); Lindner, 1930: 60 (disc.); Aczél, 1950: 38 (cat.); Steyskal, 1968: 11 (cat.). Remarks Some additional species from Colombia were found in a material recently examined, nevertheless it was not possible to identify them until a specific level (see Table 1) and probably these species are new. These species, included in the Table 1 below, belong to four genera for the first time recorded to the country, and they are the follows: Beebeomyia Curran (with three unidentified species); Megalothoraca Hendel (with one species); Setellia Robineau-Desvoidy (with at least three species) and Epiplatea Loew (with at least one species). Acknowledgements We sincerely thank Dr. K. Goodger (BMNH) for loaning material; and Dr. Marta Wolff (CEUA), Eduardo C. Amat Garcia (CE-TDEA) and John Alexander Pulgarín (MEFLG) for welcoming us at the their respective collection and for loaning material. We thank Fundação de Amparo à Pesquisa do Estado do Amazonas (FAPEAM) and Coordenação de Aperfeiçoamento de Pessoal de Nível Superior (CAPES) for supporting the Project “Incremento do conhecimento da diversidade, formação e fixação de especialistas em Sistemática de Diptera (Insecta) na Amazônia”. We are grateful to CAPES, FAPEAM and Conselho Nacional de Desenvolvimento Científico e Tecnológico (CNPq) for the scholarships provided. TABLE 1. Richardiidae species recorded from Colombia. Subfamily Genus Species Author Epiplateinae Automola atomaria (Wiedemann, 1830) Epiplatea sp.* Richardiinae Batrachophthalmum quimbaya Beebeomyia spp.* Carvalho, Wolff & Wendt, 2011 Coilometopia costalis Hendel, 1911 Coilometopia longicornis Hendel, 1911 Coilometopia trimaculata (Fabricius, 1805) Euolena egregia (Gerstaecker, 1860) Hemixantha fascinventris Enderlein, 1912 Megalothoraca sp.* Melanoloma decrepitum Hendel, 1911 Melanoloma viatrix Hendel, 1911 Odontomera canonigra Enderlein, 1912 Odontomera nitens (Schiner, 1868) Poecilomyia longicornis Hendel, 1911 Poecilomyia longicornis var. colombiana Enderlein, 1912 Richardia annulata (Macquart, 1835) Richardia flavipes Schiner, 1868 Richardia hendeliana Enderlein, 1913 Richardia infestans (Enderlein, 1912) Richardia podagrica (Fabricius, 1805) Richardia stylops Hennig, 1938b Richardia telescopica Gerstaecker, 1860 Richardia undulata Hennig, 1937 ...Continued on next page 592 · Zootaxa 4122 (1) © 2016 Magnolia Press WENDT & ALE-ROCHA TABLE 1. (Continued) Subfamily Genus Species Author Richardia viridiventris Wulp, 1898 Sepsisoma geniculatum (Schiner, 1868) Setellia spp.* (*See comments above in the Remarks) References Aczél, M. (1950) Catalogo de la sufamilia americana “Richardiinae”(Dipt. Acalypt. Otitidae). Acta Zoologica Lilloana, 9, 5–47. Aldrich, J.M. (1905) A catalogue of North American Diptera (or two-winged flies). Smithsonian Miscellaneous Collections, 46, 1–680. Alencar, L.B. de, Wendt, L.D. & Ale-Rocha, A. (2013) Richardia Robineau-Desvoidy (Diptera, Tephritoidea, Richardiidae) from Central Amazon, Brazil. Revista Brasileira de Entomologia, 57, 202–208. http://dx.doi.org/10.1590/S0085-56262013005000015 Arévalo, P.E. & Osório, O.M. (1995) Consideraciones generales sobre Melanoloma viatrix Hendel, uma nueva plaga de la piña. Revista Colombiana de Entomologia, 21, 1–8. Becerril-Morales, F. & Macías-Ordóñez, R. (2009) Territorial contests within and between two species of flies (Diptera: Richardiidae) in the wild. 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Giglio-Tos, E. (1895) Ditteri del Messico. Part 4, 74 pp. Torino. Grimaldi, D. & Fenster, G. (1989) Evolution of extreme sexual dimorphisms: structural and behavioral convergence among broad-headed male Drosophilidae (Diptera). Americam Museum Novitates, 2939, 1–15. CATALOGUE OF RICHARDIIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 593 Hancock, E.G. (2010) Richardiidae (Richardiid flies). In: Brown, B.V., Borkent, A., Cumming, J.M., Wood, D.M., Woodley, N.E. & Zumbado, M.A. (Eds.), Manual of Central American Diptera. Vol. 2. NRC Research Press, Ottawa, pp. 871–879. Hendel, F. (1911a) Die arten der dipteren-subfamilie Richardiinae. Deutsche Entomologische Zeitschrift, 1911, 181–212, 239–270, 367–396. Hendel, F. (1911b) Diptera. Fam. Muscaridae, Subfam. Richardiinae. Genera Insectorum, 113, 1–56. Hendel, F. (1936) Ergebnisse einer zoologischen Sammelreise nach Brasilien, insbesondere in das Amazonasgebiet, ausgeführt Von Dr. H. Zerny. 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(1868) Die amerikanische Ulidiina. Berliner Entomologische Zeitschrift, 11, 283–326. http://dx.doi.org/10.1002/mmnd.18670110308 Loew, H. (1873) Monographs of the Diptera of North America. Part III. Smithsonian Miscellaneous Collection, 256, 1–358. Lopes, H. de S. (1936) Sobre três espécies novas de Richardiidae do Brasil. Revista de Entomologia, 6, 475–480. Macquart, J. (1835) Histoire naturelle des insectes. Diptères, 2, 1–703. Macquart, J. (1843) Diptères exotiques nouveaux ou peu connus. Mémoires de la Société royale des sciences, de l'agriculture et des arts de Lille, 1842, 162–460. Macquart, J. (1846) Diptères exotiques nouveaux ou peu connus. Mémoires de la Société royale des sciences, de l'agriculture et des arts de Lille, 1844, 133–364. Macquart, J. (1847) Diptères exotiques nouveaux ou peu connus. Mémoires de la Société royale des sciences, de l'agriculture et des arts de Lille, 1846, 21–120. Martinez, N.B., Rosales, C.R. & Godoy, F. 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In: Wüllersdorff-Urbair, B. von (Ed.), Reise der Osterreichischen Fregatte Novara um die Erde in den Jahren 1857, 1858, 1859 unter den Befehlen des Commodore B. von Wüllerstorf-Urbair. Zoologischer Theil., Vol. 2, Abt. IB. K. Gerold’s Son,, 388 pp. Speiser, P. (1911) Zur kenntnis aussereuropäischer Dipteren. Jahrücher des Nassauischen Vereins für Naturkunde Wiesbaden, 64, 237–261. Steyskal, G.C. (1958) Notes on the Richardiidae, with a review of the species known to occur in the United States (Diptera, Acalyptratae). Annals of the Entomological Society of America, 51, 302–310. http://dx.doi.org/10.1093/aesa/51.3.302 Steyskal, G.C. (1968) Famly Richardiidae. In: Papavero, N. (Ed.) A catalogue of the Diptera of the Americas South of the United States. Departamento de Zoologia, Secretaria da Agricultura, São Paulo, 20 pp. Steyskal, G.C. (1973) The genera of the family Thyreophoridae and the species of the genus Omomyia, with one new species (Diptera). 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(2015) Antlered richardiid flies: new species of Richardia (Tephritoidea:Richardiidae) with antler-like genal processes. Entomological Science, 18, 153–166. http://dx.doi.org/10.1111/ens.12122 Wiedemann, C.R.W. (1830) Aussereuropäische zweiflügelige Insekten. Part II. in der Schulzischen Buchhandlung, Hamm, XII + 684 + XI pp. Williston, S.W. (1908) Manual of North American Diptera. 3rd Edition. James T. Hathaway, New Haven, 405 pp. http://dx.doi.org/10.5962/bhl.title.20989 Wulp, F.M. van der (1898) Group Ortalinae. In: Godman, F.D. & Salvin, O. (Eds.), Biologia Centralli-Americana. ZoologiaInsecta-Diptera. Vol. 2. Taylor & Francis, London, pp. 377–400. Zumbado, M.A. (2006) Dípteros de Costa Rica Y la America Tropical. Instituto Nacional de Biodiversidad, Santo Domingo de Heredia, 267 pp. CATALOGUE OF RICHARDIIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 595 Zootaxa 4122 (1): 596–621 http://www.mapress.com/j/zt/ Copyright © 2016 Magnolia Press Article ISSN 1175-5326 (print edition) ZOOTAXA ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.4122.1.50 http://zoobank.org/urn:lsid:zoobank.org:pub:D9CF6C6C-6295-4DCD-B247-A1AB25B24036 FAMILY TEPHRITIDAE MARCOANDRE SAVARIS1*, LUCIANE MARINONI1 & ALLEN L. NORRBOM2 1 Universidade Federal do Paraná, Departamento de Zoologia, Centro Politécnico, Jardim das Américas, Caixa Postal 19020, CEP 81531-980, Curitiba, Paraná, Brazil. E-mails: plaumannimyia@gmail.com and lmarinoni@ufpr.br 2 Systematic Entomology Laboratory, USDA, ARS, c/o Smithsonian Institution, P.O. Box 37012, MRC 168, Washington, DC 200137012, USA. E-mail: allen.norrbom@ars.usda.gov * Corresponding author: plaumannimyia@gmail.com Abstract The present Catalogue includes 90 species and 23 genera of Tephritidae that have been recorded in Colombia. Four subfamilies (Blepharoneurinae, Dacinae, Trypetinae and Tephritinae), and eight tribes (Acrotaeniini, Carpomyini, Dacini, Eutretini, Myopitini, Noeetini, Tephritini, and Toxotrypanini) are represented in the Colombian territory. The species Toxotrypana littoralis and Tomoplagia pleuralis are new records for the country. Trypeta conferta Walker, 1853 is here formally transferred to the genus Neomyopites. Key words: catalogue, Colombia, distribution, taxonomy, Tephritidae Introduction The Tephritidae are a family of acalyptrate flies, commonly known as fruit flies. They comprise more than 4,800 species in approximately 500 genera of worldwide distribution (Norrbom et al. 1999a, Norrbom 2010, and unpubl. data). Species of this family have brightly colored and/or patterned bodies and wings but are not metalic (except Tachinisca cyaneiventris which has a purplish abdomen, and several species from Madagascar). The size of the adults varies from 2 to 35 mm in length and they can be recognized mainly by the shape of the subcosta (bent sharply forward subapically and usually weaker or foldlike beyond the bend) and by the presence of inclinate frontal setae. In most species the basal cubital cell has a distal acute projection (Foote et al. 1993, Norrbom 2010). Fruit flies are distributed in all biogeographic regions except in extreme desert and polar areas, where their hosts are scarce or absent (Foote et al. 1993; Thompson 1999). Within the Diptera, Tephritidae is the family of greatest agricultural relevance. Most species are phytophagous and many pest species have been studied extensively, due to damage they cause in plants of economic interest (Foote 1967, Aluja 1994, Aluja & Norrbom 1999, Norrbom et al. 1999a, Aluja & Mangan 2008). Many species use fruits as a substrate for their larval development causing severe economic impact and quarantine restrictions imposed by many countries (Foote 1967, Aluja & Mangan 2008). The immatures of more than half of the species in the New World, however, develop in flowers, seeds, stems or roots of plant species of the family Asteraceae (Foote 1967, Foote et al. 1993, Prado et al. 2002, Norrbom & Prado 2006, Norrbom 2010). Although many species are considered pests, some species of Tephritinae are used in weed biological control (Thompson 1999, Rieder et al. 2001, Norrbom 2010). There are also some representatives of the subfamily Tachiniscinae that parasitize larvae of Lepidoptera, family Saturniidae (Norrbom 2010). In the Neotropical Region the number of taxa described is about 916 species in 71 genera (Norrbom 2010, and unpubl. data). Norrbom et al. (1999b) listed 52 species in 20 genera specifically from Colombia, although others were implied to occur there (e.g., as occurring from “Costa Rica to Peru”). Since then, additional species were described or recorded for that country (Hernández-Ortiz 2002, Norrbom & Caraballo 2003, Norrbom et al. 2005, Norrbom 2006, Martínez-Alava 2007, Norrbom & Condon 2010, Canal 2010, Norrbom & Korytkowski 2012). 596 Accepted by S. Nihei: 20 Apr. 2015; published: 14 Jun. 2016 Herein we list 90 species in 23 genera known from Colombia, the higher taxa listed phylogenetically, the species in alphabetical order (Table 1). We believe that as a result of new studies and surveys the number of species of fruit flies in the country will undoubtedly be much higher. Acronyms used for the depositories AMNH—American Museum of Natural History, New York, USA BMNH—The Natural History Museum, London, United Kingdom CPARJ—Centro de Pesquisas Agropecuarias Centro-Sul, EMBRAPA, Rio de Janeiro, Brazil FSCA—Florida State Collection of Arthropods, Gainesville, USA IML—Fundacion e Instituto Miguel Lillo, Universidad Nacional de Tucumán, Tucumán, Argentina IMZ—Museo ed Istituto di Zoologia Sistematica, Universita di Torino, Torino, Italy INbio—Instituto de Biodiversidad, Santo Domingo de Heredia, Costa Rica IOC—Instituto Oswaldo Cruz, Rio de Janeiro, Brazil IPV—Instituto de Patologia Vegetal, Buenos Aires, Argentina IZAM—Universidad Central de Venezuela, Maracay, Venezuela MACN—Museo Argentino de Ciencias Naturales Bernardino Rivadavia, Buenos Aires, Argentina MCZ—Museum of Comparative Zoology, Cambridge, USA MCSNM—Museo Civico di Storia Naturale, Milan, Italy MEN-UT—Museo del Laboratorio de Entomología de la Universidad del Tolima, Ibagué, Colombia MLUH—Wissenschaftsbereich Zoologie, Martin-Luther-Universitata Halle, Germany MMS—MacLeay Museum, University of Sydney, Australia MNHNP—Museum National d’Histoire Naturelle, Paris, France MNM—Magyar Termeszettudomanyi Muzeum Allattara (Hungarian Natural History Museum), Budapest, Hungary MUSENUV—Museo de Entomología de la Universidad del Valle, Cali, Colombia MZUSP—Museu de Zoologia, Universidade de São Paulo, São Paulo, Brazil NHMW—Naturhistorisches Museum Wien, Austria NRS—Naturhistoriska Riksmuseet, Sektionen fur entomologi, Stockholm, Sweden PAN—Polish Academy of Science, Museum of the Institute of Zoology, Warsaw, Poland SANC—South African National Collection of Insects, South Africa SMT—Staatliches Museum fur Tierkunde, Dresden, Germany SMN—Staatliches Museum fur Naturkunde, Stuttgart, Germany UMO—University Museum of Natural History, Oxford, United Kingdom UPRG—Universidad Nacional “Pedro Ruiz Gallo”, Lambayeque, Peru USNM—National Museum of Natural History, Washington DC, USA UZMC—Natural History Museum of Denmark, Copenhagen, Denmark ZMHU—Museum fur Naturkunde, Humboldt Universitat, Berlin, Germany List of abbreviations A— adult cat.—catalogue comm.—comments desc.—description distr.—geographical distribution emend.—emendation ft.—feet incosp.—incorrect original spelling m—meters CATALOGUE OF TEPHRITIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 597 missp.—misspelling mon.—monotypy new comb.—new combination nom. nov.—nomen novum orig. des.—original designation orig. desc.—original description preocc.—preoccupied redesc.—redescription refs.—references rev.—revision subs. des.—subsequent designation syn.—synonymy T—type HT—holotype LT—lectotype NT—neotype ST—syntype F (f#)—female M (m#)—male Catalogue of Tephritidae of Colombia Family Tephritidae Newman, 1834 Subfamily Blepharoneurinae Korneyev, 1994 Genus Blepharoneura Loew Blepharoneura Loew, 1873: 272. Type species, Trypeta poecilogastra Loew (orig. des.). Refs.: Wulp, 1899: 402, 411 (keys to 7 spp.); Hendel, 1914b: 82 (in key); Hendel, 1914c: 5 (in key), 20 (key to 15 spp.); Bates, 1933: 48 (comm.); Hering, 1941: 123 (in key), 132 (key to 3 spp.); Aczél, 1950a: 195 (cat.); Aczél, 1953: 104 (in key); Foote, 1967: 17 (cat.); Foote, 1980: 7 (in key), 20 (rev.); Norrbom et al., 1999b: 105 (cat.); Norrbom & Condon, 2010: 8 (rev. femoralis group). furcifer Hendel, 1914c: 23. Type locality: Peru, Pachitea-Münd. [mouth of Río Pachitea], 150 m. LT F (NHMW). Distr.: Bolivia, Brazil, Colombia, Ecuador, Guyana, Peru, and Venezuela. Refs.: Hering, 1935: 226 (in key); Hering, 1941: 133 (in key); Aczél, 1950a 196 (cat.); Foote, 1967: 18 (cat.); Hardy, 1968: 110 (type data); Norrbom et al., 1999b: 106 (cat.); Norrbom & Condon, 2010: 67 (redesc.). quadristriata Wulp, 1899: 413. Type locality: Mexico, Tabasco, Teapa. HT M (BMNH). Distr.: Colombia, Costa Rica, Ecuador, Guatemala, and Mexico. Refs.: Hendel, 1914c: 21 (in key, cat.); Aczél, 1950a: 197 (cat.); Hering, 1941: 134 (in key); Foote, 1965: 241 (type data); Foote, 1967: 18 (cat.); Norrbom et al., 1999b (cat.); Norrbom & Condon, 2010: 96 (redesc.). Subfamily Dacinae Loew, 1861 [as Dacina] Tribe Dacini Genus Ceratitis Macleay Ceratitis MacLeay, 1829: 482. Type species, citriperda MacLeay (mon.) (= capitata Wiedemann). Refs.: Bezzi, 1909: 273 (comm.), 278 (key to 18 spp.); Hendel, 1914b: 76 (in key); Hendel, 1914c: 4 (in key); Bezzi, 1924a: 98 (key to 2 spp.); 598 · Zootaxa 4122 (1) © 2016 Magnolia Press SAVARIS ET AL. Aczél, 1950a: 190 (cat.); Foote, 1967: 20 (cat.); Foote, 1980: 8 (in key), 22 (rev.); Yepes & Vélez, 1989: 74 (distr.); Norrbom et al., 1999b: 118 (cat.). capitata Wiedemann, 1824: 55 (Tephritis). Type locality: India orient., mare indico. T M (UZMC). Distr.: Tropical Africa. Introduced to Madagascar, Mauritius, Réunion, Seychelles, North Africa, southern Europe, Middle East, Neotropics, western Australia, Hawaii. Refs.: Wiedemann, 1830: 496 (type data); Bezzi, 1909: 276 (comm.); Hendel, 1914c: 10 (comm.); Aczél, 1950a: 191 (cat.); Foote, 1967: 20 (cat.); Yepes & Vélez, 1989: 76, 83 (distr.); Portilla et al., 1994: 262 (distr.); Gonzáles et al., 1988: 333 (distr.); Núñez Bueno et al., 2004: 5 (comm.); Uchôa, 2012: 279 (distr.). Comm.: An introduced exotic pest, commonly known as the “Mediterranean fruit fly”. citriperda MacLeay, 1829: 482. Type locality: Azores, Sao Miguel. ST M/F (MMS). (syn.). hispanica Breme, 1842: 188. Type locality: Spain, Malaga. ST M/F (IMZ). (syn.). asparagi Bezzi, 1924b: 480 (Pardalaspis). Type locality: South Africa, Cape, East London. HT F (SANC). (syn.). Subfamily Trypetinae Loew, 1861 Tribe Carpomyini Norrbom, 1989 [as Carpomyina] Genus Cryptodacus Hendel Cryptodacus Hendel, 1914b: 84. Type species, obliquus Hendel (orig. des.). (preocc. Gundlach, 1861; suspension of ICZN rules required to validate usage, in the interest of stability we maintain the usage of this name). Refs.: Aczél, 1950a: 192 (cat.); Aczél, 1954: 72 (in key); Foote, 1967: 21 (cat.); Foote, 1980: 8 (in key), 24 (rev.); Norrbom, 1994: 38 (rev. of 8 spp.); Norrbom et al., 1999b: 128 (cat.). Cryptodacus Hendel, 1914c: 12. Type species, obliquus Hendel (orig. des.). (preocc. Gundlach, 1861, Hendel, 1914b: 84). Lezca Foote, 1978: 27. Type species, tau Foote (orig. des.). Norrbom 1994: 38 (syn.). Muhabbetiella Koçak & Kemal, 2009: 11. Nom. nov. for Cryptodacus Hendel, 1914b: 84. Type species, Cryptodacus obliquus Hendel (automatic). ornatus Norrbom, 1994: 41. Type locality: Colombia, “Cordill., terra caliente”. HT F (ZMHU). Distr.: Brazil, Colombia, and Ecuador. Refs.: Norrbom et al., 1999b: 128 (cat.); Norrbom & Korytkowski, 2008: 32 (comm.), 42 (in key). Genus Rhagoletis Loew Rhagoletis Loew, 1862: 44. Type species, Musca cerasi Linnaeus (mon.). Refs.: Hendel, 1927: 74 (key to 4 spp.); Hering, 1941: 124 (in key), 141 (key to 3 spp.); Aczél, 1954: 76 (key to 10 spp.); Hering, 1958: 1 (key to 9 spp.); Bush, 1966: 454 (rev. of 22 spp.); Kandybina, 1977: 129 (key to larvae of 15 spp.); Foote, 1981: 9 (rev. of 21 spp.); Foote et al., 1993: 328 (key to 25 spp.). [pomonella Walsh, 1867: 343 (Trypeta). Type locality: USA, Illinois. LT F (MCZ). Lectotype designated by Bush, 1966: 457. Distr.: Canada, Mexico, and United States; record from Colombia doubtful, needs confirmation. Refs.: Loew, 1873: 265 (redesc.); Garcés & Gallego, 1947: 472 (comm.); Bush, 1966: (rev.); Foote, 1981: 5 (distr.), 7 (host), 10 (in key), 52 (comm.); Norrbom et al., 1999b: 201 (cat.); Weems & Fasulo, 2006: 1 (distr.). Comm.: The occurence of Rhagoletis pomonella in Colombia is based on the report of Garcés & Gallego (1947) who mention this fly in apples. However, the identification was not confirmed and this species has not been reported from Colombia subsequently, thus its occurance in the country is doubtful.] Genus Zonosemata Benjamin Zonosemata Benjamin, 1934: 17. Type species, Trypeta electa Say (orig. des.). Refs.: Bush, 1965: 307 (rev. of 5 spp.); Foote, CATALOGUE OF TEPHRITIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 599 1967: 66 (cat.); Steyskal, 1975a: 231 (key to larvae of 2 spp.); Foote, 1980: 8 (in key), 51 (rev.); Hernández-Ortiz, 1989: 207 (key to 7 spp.); Norrbom, 1990: 53 (comm. on 5 spp.); Norrbom, 2002b: 614 (key). scutellata Hendel, 1936: 73 (Cryptodacus). Type locality: Brazil, Natal. HT M (NHMW). Distr.: Brazil, Colombia, and Venezuela. Refs.: Steyskal, 1974: 234 (desc.); Norrbom, 1990: 53 (comm.); Norrbom et al., 1999b: 249 (cat.); Norrbom, 2002b: 615 (in key), 620 (redesc.). ica Steyskal, 1974: 234. Type locality: Colombia, Norte de Santander, Cucuta. HT F (USNM). Norrbom, 1990: 53 (syn.). Tribe Toxotrypanini Hancock, 1986 [as Toxotrypaninae] Genus Anastrepha Schiner Anastrepha Schiner, 1868: 263. Type species, Dacus serpentinus Wiedemann, 1830 (orig. des.). Refs.: Bezzi, 1909: 280 (comm.); Hendel, 1914a: 66, 1914b: 84 (in key), 1914c: 5 (in key), 13 (comm.); Greene, 1934: 130 (rev.); Lima, 1934b: 487 (rev. of 62 spp.); Stone, 1939b: 282 ((Pseudodacus) rev. of 4 spp. of daciformis group); Stone, 1939a: 340 ((Lucumaphila) rev. of 10 spp.); Hering, 1941: 123 (in key), 136 (key to 14 spp. Peru); Stone, 1942a: 1 (rev. of 126 spp.); Stone, 1942b: 298 (rev. of 7 spp.); Aczél, 1950a: 198 (cat.); Blanchard, 1961: 281 (monograph of 38 spp., Argentina); Foote, 1967: 6 (cat.); Steyskal, 1977b: 3 (key to 141 spp.); Zucchi, 1978: 1 (monograph of 77 spp., Brazil); Foote, 1980: 7 (in key), 19 (rev.); Caraballo, 1981: 1 (monograph of 37 spp., Venezuela); Norrbom, 1985: 1 (rev. of cryptostrepha, daciformis, robusta & schausi species groups); Norrbom & Kim, 1988: 164 (rev. of schausi group); Yepes & Vélez, 1989: 74 (distr.); Steck et al., 1990: 342 (key to larvae of 13 spp.); Norrbom, 1991: 101 (rev. of grandis species group); Hernández-Ortiz, 1992: 1 (monograph of 28 spp., Mexico); Foote et al., 1993: 85 (key to 19 spp.); Norrbom et al., 1999b: 76 (cat.); Norrbom, 2002a: 390 (rev. of serpentina species group); Norrbom et al., 2003: 77 (rev. hastata species group); Norrbom & Korytkowski, 2009: 1 (rev. of robusta species group). Acrotoxa Loew, 1873: 227. Type species, Dacus fraterculus Wiedemann (subs. des. Bezzi, 1909: 280). Pseudodacus Hendel, 1914a: 66. Type species, Anastrepha daciformis Bezzi (orig. des.). Phobema Aldrich, 1925: 7. Type species, atrox Aldrich (orig. des.). Lucumaphila Stone, 1939a: 340. Type species, sagittata Stone (orig. des.). Pseudodacus Hendel, 1914b: 97. Type species, Anastrepha daciformis Bezzi (orig. des.) (preocc. Hendel, 1914a: 66). Pseudodacus Hendel, 1914c: 13. Type species, Anastrepha daciformis Bezzi (orig. des.) (preocc. Hendel, 1914a: 66). acuminata Canal, 2010: 33. Type locality: Colombia, Tolima, Ibagué, Santafé de los Guaduales, 1.107 m. HT F (MEN-UT). Distr.: Colombia. Refs.: Castañeda et al., 2010: 267 (distr.). alveata Stone, 1942a: 72. Type locality: Panama, La Campana. HT F (USNM). Distr.: Argentina, Colombia, Guatemala, Mexico, Panama, and Venezuela. Refs.: Aczél, 1950a: 199 (cat.); Foote, 1967: 6 (cat.); Norrbom et al., 1999b: 76 (cat.); Martínez-Alava, 2007: 37 (distr.); Canal, 2010: 42 (distr.); Castañeda et al., 2010: 267 (distr.). anomoiae Norrbom, 2002a: 398. Type locality: Colombia, Boyacá, Guaguaqui. HT F (USNM). Distr.: Colombia and Panama. Refs.: Canal, 2010: 43 (distr.). antunesi Lima, 1938: 63. Type locality: Brazil, Bahia, Piraja. HT F (IOC). Distr.: Brazil, Colombia, Guatemala, Panama, Peru, Trinidad, and Venezuela. Refs.: Stone, 1942a: 73 (redesc.); Aczél, 1950a: 199 (cat.); Foote, 1967: 7 (cat.); Korytkowski & Peña, 1968: 67 (redesc.); Norrbom et al., 1999b: 77 (cat.); Martínez-Alava, 2007: 38 (distr.); Canal, 2010: 42 (distr.). bahiensis Lima, 1937b: 60. Type locality: Brazil, Bahia. HT F (IOC). Distr.: Belize, Brazil, Colombia, Costa Rica, Ecuador, Guatemala, Guyana, Honduras, Mexico, Nicaragua, Panama, Peru, Surinam, Trinidad, and Venezuela. Refs.: Stone, 1942a: 107 (redesc.); Aczél, 1950a: 200 (cat.); Foote, 1967: 7 (cat.); Korytkowski & Peña, 1968: 68 (redesc.); Carrejo & González, 1994: 89 (distr.); Norrbom et al., 1999b: 77 (cat.); Martínez & Serna, 2005: 109 (distr.); Canal, 2010: 42 (distr.); Uchôa, 2012: 276 (distr.). [binodosa Stone, 1942a: 57. Type locality: Brazil, Pará. HT F (BMNH). Distr.: Brazil. Refs.: Aczél, 1950a: 201 600 · Zootaxa 4122 (1) © 2016 Magnolia Press SAVARIS ET AL. (cat.); Foote, 1967: 8 (cat.); Norrbom et al., 1999b: 77 (cat.); Norrbom & Korytkowski, 2009: 13 (redesc.). The record of this species from Colombia reported by Carrejo & González, 1994: 87 was based on a misidentification of an undetermined species of the dentata group (Norrbom & Korytkowski 2009), and there are no other records of it from Colombia.] carreroi Canal, 2010: 34. Type locality: Colombia, Tolima, Ibagué, Totumo. HT F (MEN-UT). Distr.: Colombia. Refs.: Castañeda et al., 2010: 267 (distr.). caudata Stone, 1942a: 85. Type locality: Brazil, São Paulo. HT F (BMNH). (The type locality is probably Brazil, Amazonas, São Paulo de Olivença). Distr.: Brazil and Colombia. Refs.: Aczél, 1950a: 203 (cat.); Foote, 1967: 8 (cat.); Steyskal, 1977b: 23 (in key); Arévalo et al., 1997: 274 (distr.); Norrbom et al., 1999b: 77 (cat.); Norrbom & Caraballo, 2003: 34 (in key), 37 (redesc.); Canal, 2010: 43 (distr.). coronilli Carrejo & González, 1993: 48. Type locality: Colombia, Valle del Cauca, Anchicaya, Buenaventura. HT F (MUSENUV). Distr.: Brazil, Colombia, Costa Rica, Guatemala, Mexico, Panama, and Surinam. Refs.: Carrejo & González, 1994: 90 (distr.); Norrbom et al., 1999b: 78 (cat.); Carrejo & González, 1999: 115 (distr.); Silva & Ronchi-Teles, 2000: 203 (distr.); Aluja et al., 2003: 1381 (distr.); Canal, 2010: 42 (distr.). curitis Stone, 1942a: 32. Type locality: Brazil, Pará. HT F (BMNH). Distr.: Brazil, Colombia, and Peru. Refs.: Aczél, 1950a: 206 (cat.); Foote, 1967: 9 (cat.); Yepes & Vélez, 1989: 88 (distr.); Norrbom, 1997: 148 (redesc.); Norrbom et al., 1999b: 78 (cat.); Canal, 2010: 42 (distr.). debilis Stone, 1942a: 47. Type locality: Panama, La Campana. HT F (USNM). Distr.: Colombia, Ecuador, and Panama. Refs.: Aczél, 1950a: 206 (cat.); Foote, 1967: 9 (cat.); Carrejo & González, 1994: 88 (distr.); Norrbom et al., 1999b: 78 (cat.); Canal, 2010: 42 (distr.). dissimilis Stone, 1942a: 40. Type locality: Haiti, Near Plaisance. HT F (USNM). Distr.: Argentina, Brazil, Colombia, Guyana, Haiti, Hispaniola, Peru, and Trinidad. Refs.: Aczél, 1950a: 207 (cat.); Foote, 1967: 9 (cat.); Korytkowski & Peña, 1968: 54 (redesc.); Carrejo & González, 1994: 90 (distr.); Norrbom et al., 1999b: 78 (cat.); Canal, 2010: 42 (distr.). correntina Blanchard, 1961: 339. Type locality: Argentina, Corrientes, Bella Vista. HT F (IPV). Steyskal, 1977b: 33 (syn.). distincta Greene, 1934: 149. Type locality: Peru, Lambayeque, Chiclaya, Hacienda Ouefe. HT F (USNM). Distr.: Brazil, British Guiana, Colombia, Costa Rica, Ecuador, Guatemala, Mexico, Panama, Peru, Trinidad, and United States. Refs.: Hering, 1941: 137 (in key); Stone, 1942a: 106 (redesc.); Aczél, 1950a: 207 (cat.); Foote, 1967: 9 (cat.); Korytkowski & Peña, 1968: 67 (redesc.); Yepes & Vélez, 1989: 83 (distr.); Carrejo & González, 1994: 89 (distr.); Gonzáles et al., 1988: 334 (distr.); Norrbom et al., 1999b: 78 (cat.); Carrejo & González, 1999: 115 (distr.); Martínez & Serna, 2005: 109 (distr.); Canal, 2010: 42 (distr.); Castañeda et al., 2010: 267 (distr.). silvai Lima, 1934b: 545. Type locality: Brazil, Rio de Janeiro; Minas Gerais, Viçosa & São Paulo. ST M/F (IOC). Stone, 1942a: 106 (syn.). doryphoros Stone, 1942b: 299. Type locality: Panama, El Cermeño. HT F (USNM). Distr.: Colombia, Costa Rica, and Panama. Refs.: Aczél, 1950a: 208 (cat.); Foote, 1967: 10 (cat.); Carrejo & González, 1994: 87 (distr.); Norrbom et al., 1999b: 78 (cat.); Canal, 2010: 43 (distr.). flavipennis Greene, 1934: 160. Type locality: Brazil, Pará, Boa Vista, Rio Tapajós. HT F (USNM). Distr.: Argentina, Brazil, Colombia, Panama, and Venezuela. Refs.: Stone, 1942a: 89 (redesc.); Aczél, 1950a: 209 (cat.); Foote, 1967: 10 (cat.); Carrejo & González, 1994: 88 (distr.); Norrbom et al., 1999b: 78 (cat.); Canal, 2010: 43 (distr.). grioti Blanchard, 1941: 60. Nomen nudum. See Blanchard, 1961: 311. CATALOGUE OF TEPHRITIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 601 fraterculus Wiedemann, 1830: 524 (Dacus). Type locality: Brazil, São Paulo. LT M (NHMW). Lectotype designation by inference of holotype by Greene, 1934: 165 (also see Loew, 1873: 223, 336, and Stone, 1942a: 78). Distr.: Argentina, Belize, Bolivia, Brazil, Colombia, Costa Rica, Cuba, Ecuador, Guatemala, Guyana, Honduras, Mexico, Nicaragua, Panama, Paraguay, Peru, Porto Rico, Republica Dominicana, Surinam, Trinidad and Tobago, United States, Uruguay, and Venezuela. Refs.: Loew, 1873: 222 (redesc.); Brèthes, 1908: 300 (cat.); Bezzi, 1909: 286 (in key); Hendel, 1914c: 14 (in key), 15 (cat.), 18 (redesc.); Fischer, 1932: 308 (distr.); Greene, 1934: 136 (in key), 164 (comm.); Lima, 1934b: 546 (redesc.); Hendel, 1936: 73 (distr.); Hering, 1941: 137 (in key), 139 (comm.); Stone, 1942a: 78 (redesc.); Aczél, 1950a: 210 (cat.); Foote, 1967: 10 (cat.); Hardy, 1968: 145 (type data); Korytkowski & Peña, 1968: 60 (redesc.); Zucchi, 1981: 289 (new syn.); Yepes & Vélez, 1989: 82 (distr.); Carrejo & González, 1994: 90 (distr.); Gonzáles et al., 1988: 334 (distr.); Norrbom et al., 1999b: 78 (cat.); Portilla et al., 1994: 262 (distr.); Patiño, 2002: 25 (distr.); Núñez Bueno et al., 2004: 7 (distr.); Martínez & Serna, 2005: 109 (distr.); Canal, 2010: 42 (distr.); Castañeda et al., 2010: 267 (distr.); Hernández-Ortiz et al., 2012: 306 (distr.); Sarmiento et al., 2012: 28 (distr.); Uchôa, 2012: 276 (distr.); Weems, 2012: 2 (distr.); Wyckhuys et al., 2012: 94 (distr.). mellea Walker, 1837: 358. (Tephritis). Type locality: Brazil, St. Paul’s [São Paulo]. LT F (BMNH). (syn.). unicolor Loew, 1862: 70. (Trypeta). Type locality: New Grenada [Colombia or Venezuela]. LT M (MCZ). Loew 1873: 222 (syn.). frutalis Weyenbergh, 1874: 165. (Anthomyia). Type locality: Argentina, Cordoba. T A destroyed?. Type data (Horn & Kahle, 1937: 301). (syn.). fraterculus var. soluta Bezzi, 1909: 284. Type locality: Brazil, São Paulo. ST M/F (MCSNM?). Stone, 1942: 78 (syn.). peruviana Townsend, 1913: 345. Type locality: Peru, Lima, Chosica, 2800 ft. HT F (USNM). Stone, 1942: 78 (syn.). braziliensis Greene,1934: 154. Type locality: Brazil, Minas Gerais, Viçosa. HT F (USNM). Stone, 1942: 78 (syn.). costarukmanii Capoor, 1954: 214. Type locality: Brazil, Minas Gerais, Itajuba. HT F (IOC). Zucchi, 1981: 289 (syn.). scholae Capoor, 1955: 27. Type locality: Brazil, Bahia, Agua Preta. LT F (IOC). Zucchi, 1981: 289 (syn.). pseudofraterculus Capoor, 1955: 28. Type locality: Brazil, Rio de Janeiro, Itatiaia. LT F (IOC). Zucchi, 1981: 289 (syn.). lambayecae Korytkowski & Peña, 1968: 63. Type locality: Peru, Lambayeque, Chiclayo, Chumbenique, 225 m. HT M (UPRG). (syn.). fraterculus var. intensa Blanchard, 1941: 60. Nomen nudum. fraterculus var. retracta Hayward, 1941: 95. Nomen nudum. fraterculus “variacion” intermissa Blanchard, 1961: 319. Nomen nudum. fraterculus “variacion” intensa Blanchard, 1961: 319. Nomen nudum. fraterculus “variacion” retracta Blanchard, 1961: 319. Nomen nudum. fraterculus “variacion” ethaleiforme Blanchard, 1961: 320. Nomen nudum. fraterculus “variacion” tipica Blanchard, 1961: 319. Nomen nudum. fraterculus “variacion” acidusaeformis Blanchard, 1961: 320. Nomen nudum. fraterculus “variacion” subtipica Blanchard, 1961: 320. Nomen nudum. costa-rukmanii Capoor, 1954: 214. (incosp. costarukmanii). costa-bukmanii Capoor, 1954: 214. (incosp. costarukmanii). pseudo-fraterculus Capoor, 1955: 28. (incosp. pseudofraterculus). grandicula Norrbom, 1991: 118. Type locality: Colombia, Amazonas, Rio Putumayo, Puerto Arica. HT F (FSCA). Distr.: Brazil and Colombia. Refs.: Norrbom et al., 1999b: 79 (cat.); Canal, 2010: 42 (distr.). grandis Macquart, 1846: 340 (Tephritis). Type locality: Nouvelle-Grenade [Colombia or Venezuela]. LT F (UMO). Distr.: Argentina, Bolivia, Brazil, Colombia, Panama, Paraguay, Peru, and Venezuela. Refs.: Bezzi, 1909: 286 (in key); Hendel, 1914c: 14 (in key), 15 (cat.); Fischer, 1932: 303 (type data), 304 (redesc.), 308 (distr.); Greene, 1934: 132 (in key), 145 (comm.); Lima, 1934b: 496 (redesc.); Hering, 1941: 137 (in key), 139 (comm.); Stone, 1942a: 22 602 · Zootaxa 4122 (1) © 2016 Magnolia Press SAVARIS ET AL. (redesc.); Aczél, 1950a: 212 (cat.); Foote, 1967: 11 (cat.); Korytkowski & Peña, 1968: 43 (redesc.); Norrbom, 1991: 104 (in key), 118 (redesc.); Carrejo & González, 1994: 87 (distr.); Norrbom et al., 1999b: 79 (cat.); Patiño, 2002: 22 (distr.); Martínez & Serna, 2005: 109 (distr.); Canal, 2010: 42 (distr.); Castañeda et al., 2010: 267 (distr.); Uchôa, 2012: 276 (distr.); Wyckhuys et al., 2012: 94 (distr.). latifasciata Hering, 1935: 227. Brazil, Santa Catarina. HT F (PAN). Lima, 1937: 63 (syn.). schineri Hendel, 1914a: 69. Bolivia, La Paz, Coroica [Coroico]. HT M (MNM). Fischer, 1932: 303 (syn.). schineri Hendel, 1914c: 19. (preocc. Hendel, 1914a: 69). hendeliana Lima, 1934b: 528. Type locality: Brazil, Amazonas, Rio Negro, São Gabriel. LT M (IOC). Distr.: Brazil and Colombia. Refs.: Stone, 1942a: 85 (redesc.); Aczél, 1950a: 214 (cat.); Foote, 1967: 12 (cat.); Steyskal, 1977b: 33 (in list); Zucchi 1981: 291 (syn. with Anastrepha longicauda); Zucchi et al., 1996: 261 (LT designation); Norrbom et al., 1999a: 80 (in cat. as syn.); Norrbom et al., 1999b: 338 (cat. as syn.); Norrbom & Caraballo, 2003: 34 (in key), 38 (new status); Canal, 2010: 43 (distr.). isolata Norrbom & Korytkowski, 2009: 30. Type locality: Ecuador, Napo, Reserva Etnica Waorani, Onkone Gare Camp. 220 m. HT F (USNM). Distr.: Brazil, Colombia, and Ecuador. Refs.: Canal, 2010: 39 (distr.). lanceola Stone, 1942a: 44. Type locality: Panama, El Cermeño. HT F (USNM). Distr.: Colombia, Costa Rica, and Panama. Refs.: Aczél, 1950a: 215 (cat.); Foote, 1967: 12 (cat.); Korytkowski & Peña, 1968: 59 (redesc.); Norrbom et al., 1999b: 80 (cat.); Canal, 2010: 40 (distr.); Castañeda et al., 2010: 267 (distr.). leptozona Hendel, 1914a: 69. Type locality: Bolivia. T A (SMT). Distr.: Bolivia, Brazil, Colombia, Guatemala, Guyana, Mexico, and Trinidad. Refs.: Hendel, 1914c: 15 (in key); Fischer, 1932: 308 (distr.); Greene, 1934: 133 (in key), 153 (comm.); Lima, 1934b: 515 (redesc.); Stone, 1942a: 39 (redesc.); Aczél, 1950a: 216 (cat.); Foote, 1967: 12 (cat.); Carrejo & González, 1994: 89 (distr.); Norrbom et al., 1999b: 80 (cat.); Carrejo & González, 1999: 115 (distr.); Canal, 2010: 42 (distr.). leptozona Hendel, 1914c: 19. Type locality: Bolivia, La Paz, Mapiri, San Antonio, 1.000 m. HT M (SMT). (preocc. Hendel 1914a: 69). limae Stone, 1942a: 67. Type locality: Panama, Panama City. HT F (USNM). Distr.: Colombia, Costa Rica, Guatemala, Mexico, Panama, and United States. Refs.: Aczél, 1950a: 216 (cat.); Foote, 1967: 12 (cat.); Yepes & Vélez, 1989: 83 (distr.); Gonzáles et al., 1988: 334 (distr.); Norrbom et al., 1999b: 80 (cat.); Martínez & Serna, 2005: 109 (distr.); Canal, 2010: 42 (distr.); Castañeda et al., 2010: 267 (distr.). loewi Stone, 1942a: 49. Type locality: Panama, La Campana. HT F (USNM). Distr.: Colombia, Costa Rica, Panama, and Venezuela. Refs.: Aczél, 1950a: 216 (cat.); Foote, 1967: 12 (cat.); Norrbom et al., 1999b: 80 (cat.); Norrbom & Korytkowski, 2012: 159 (redesc.). magna Norrbom, 1997: 149. Type locality: Venezuela, Amazonas, San Pedro de Cataniapo, 100 m. HT F (IZAM). Distr.: Colombia and Venezuela. Refs.: Canal, 2010: 42 (distr.). manihoti Lima, 1934b: 543. Type locality: Brazil, Minas Gerais, Viçosa. ST F/M (IOC). Distr.: Brazil, Colombia, Costa Rica, Ecuador, Panama, Peru, and Venezuela. Refs.: Stone, 1942a: 64 (redesc.); Aczél, 1950a: 218 (cat.); Foote, 1967: 13 (cat.); Korytkowski & Peña, 1968: 55 (redesc.); Carrejo & González, 1994: 89 (distr.); Gonzáles et al., 1988: 334 (distr.); Norrbom et al., 1999b: 80 (cat.); Martínez & Serna, 2005: 109 (distr.); Canal, 2010: 42 (distr.); Castañeda et al., 2010: 267 (distr.). manihoti Stone, 1942a: 64. (emend. manhioti Lima). manhioti Lima, 1934b: 543. (incosp. manihoti Lima). manizaliensis Norrbom & Korytkowski, 2005: 67. Type locality: Colombia, Caldas, Manizales, 2.150 m. HT F (USNM). Distr.: Colombia. Refs.: Canal, 2010: 42 (distr.); Castañeda et al., 2010: 267 (distr.). ludens: Núñez Bueno, 1981: 178 (misidentification). CATALOGUE OF TEPHRITIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 603 matertela Zucchi, 1979a: 37. Type locality: Brazil, Bahia, Cruz das Almas, Chapadinha. HT F (MZUSP). Distr.: Brazil and Colombia. Refs.: Arévalo et al. 1997: 274 (distr.); Norrbom et al., 1999b : 80 (cat.); Canal, 2010: 42 (distr.). montei Lima, 1934b: 541. Type locality: Brazil, Minas Gerais, Belo Horizonte; Rio de Janeiro, Guaratiba. ST M/F (IOC). Distr.: Argentina, Bolivia, Brazil, Colombia, Costa Rica, Ecuador, Mexico, Panama, Paraguay, and Venezuela. Refs.: Stone, 1942a: 59 (redesc.); Aczél, 1950a: 219 (cat.); Foote, 1967: 13 (cat.); Norrbom et al., 1999b: 80 (cat.); Canal, 2010: 40 (distr.); Castañeda et al., 2010: 267 (distr.). procurvata Blanchard, 1961: 308. Argentina, Misiones, Zaiman, Estacion Experimental de INTA. HT F (IPV). (syn.). mucronota Stone, 1942a: 57. Type locality: Panama, El Cermeño. HT F (USNM). Distr.: Brazil, Colombia, Ecuador, Panama, and Peru. Refs.: Aczél, 1950a: 220 (cat.); Foote, 1967: 13 (cat.); Steyskal, 1977a: 77 (desc.); Núñez-Bueno, 1981: 175, 178 (distr. and host); Yepes & Vélez, 1989: 81 (distr. and host); Molineros et al., 1992: 16, 35 (host); Carrejo & González, 1994: 88 (distr.); Norrbom et al., 1999b: 81 (cat.); Carrejo & González, 1999: 115 (distr.); Patiño, 2002: 24 (distr.); Martínez & Serna, 2005: 109 (distr.); Bomfim et al., 2007: 220 (distr.); Zucchi, 2007: 84 (host); Canal, 2010: 42 (distr.); Castañeda et al., 2010: 267 (distr.); Norrbom & Korytkowski, 2011: 5 (distr.). nunezae Steyskal, 1977a: 77. Type locality: Colombia, Cundinamarca, Cachipay. HT F (USNM). Norrbom & Korytkowski, 2011: 5 (syn.). obliqua Macquart, 1835: 464 (Tephritis). Type locality: Cuba. LT F (MNHNP). Lectotype designation by inference of holotype by Steyskal, 1975b: 357. Distr.: Antilles, Argentina, Belize, Bolivia, Brazil, Colombia, Costa Rica, Cuba, Ecuador, Guatemala, Guyana, Honduras, Mexico, Nicaragua, Panama, Paraguay, Peru, Porto Rico, Republica Dominicana, Surinam, and Venezuela. Refs.: Bezzi, 1909: 285 (in key); Hendel, 1914c: 13 (in key), 15 (cat.), 16 (redesc.); Fischer, 1932: 309 (distr.); Greene, 1934: 135 (in key), 163 (comm.); Stone, 1942a: 75 (redesc.); Aczél, 1950a: 220 (cat.); Foote, 1967: 14 (cat.); Yepes & Vélez, 1989: 81 (distr.); Carrejo & González, 1994: 90 (distr.); Gonzáles et al., 1988: 334 (distr.); Norrbom et al., 1999b: 81 (cat.); Carrejo & González, 1999: 115 (distr.); Patiño, 2002: 20 (distr.); Martínez & Serna, 2005: 109 (distr.); Canal, 2010: 42 (distr.); Castañeda et al., 2010: 267 (distr.); Sarmiento et al., 2012: 28 (distr.); Uchôa, 2012: 277 (distr.); Wyckhuys et al., 2012: 94 (distr.). fraterculus var. mombinpraeoptans Seín, 1933: 187. Type locality: Puerto Rico, Rio Piedras. HT F (USNM). Steyskal 1975: 358 (syn.). fraterculus var. ligata Lima, 1934b: 552. Type locality: Brazil, Rio de Janeiro. ST M/F (CPARJ?). Stone 1942: 68 (syn.). trinidadensis Greene, 1934: 161. Type locality: Trinidad, Tabaquite. HT F (USNM). Stone 1942: 68 (syn.). ornata Aldrich, 1925: 6. Type locality: Ecuador, Oriente, Baños. HT F (USNM). Distr.: Colombia and Ecuador. Refs.: Fischer, 1932: 309 (distr.); Greene, 1934: 131 (in key), 143 (comm.); Lima, 1934b: 495 (comm.); Stone, 1942a: 25 (redesc.); Aczél, 1950a: 221 (cat.); Foote, 1967: 14 (cat.); Yepes & Vélez, 1989: 82 (distr.); Norrbom et al., 1999b: 81 (cat.); Norrbom, 2002a: 391 (in key), 412 (redesc.); Martínez & Serna, 2005: 109 (distr.); Canal, 2010: 43 (distr.); Castañeda et al., 2010: 267 (distr.). palae Stone, 1942a: 58. Type locality: Panama, El Cermeño. HT F (USNM). Distr.: Colombia and Panama. Refs.: Aczél, 1950a: 222 (cat.); Foote, 1967: 14 (cat.); Norrbom et al., 1999b: 81 (cat.); Martínez-Alava, 2007: 39 (distr.); Canal, 2010: 43 (distr.); Castañeda et al., 2010: 267 (distr.). pallidipennis Greene, 1934: 166. Type locality: Colombia, Antioquia, Medellin. HT F (USNM). Distr.: Colombia, Panama, and Venezuela. Refs.: Stone, 1942a: 32 (redesc.); Aczél, 1950a: 222 (cat.); Foote, 1967: 14 (cat.); Yepes & Vélez, 1989: 83 (distr.); Carrejo & González, 1994: 87 (distr.); Norrbom, 1997: 153 (redesc.); Norrbom et al., 1999b: 81 (cat.); Martínez & Serna, 2005: 109 (distr.); Canal, 2010: 42 (distr.); Castañeda et al., 2010: 267 (distr.). panamensis Greene, 1934: 150. Type locality: Panama, Canal Zone, Barro Colorado Island. HT F (USNM). Distr.: 604 · Zootaxa 4122 (1) © 2016 Magnolia Press SAVARIS ET AL. Colombia, Costa Rica, and Panama. Refs.: Stone, 1942a: 59 (redesc.); Aczél, 1950a: 222 (cat.); Foote, 1967: 14 (cat.); Gonzáles et al., 1988: 333 (distr.); Norrbom et al., 1999b: 81 (cat.); Canal, 2010: 41 (distr.); Castañeda et al., 2010: 267 (distr.). parallela Wiedemann, 1830: 515 (Dacus). Type locality: Brazil. LT F (NHMW). Distr.: Brazil, Colombia, Guyana, Mexico, and Paraguay. Refs.: Bezzi, 1909: 285 (in key); Hendel, 1914c: 13 (in key), 15 (cat.); Bezzi, 1919: 374 (comm.); Fischer, 1932: 309 (distr.); Greene, 1934: 137 (in key), 168 (comm.); Lima, 1934b: 517 (comm.); Lima, 1937a: 34 (host); Lima, 1937b: 61 (comm.); Stone, 1942a: 53 (redesc.); Aczél, 1950a: 222 (cat.); Foote, 1967: 14 (cat.); Hardy, 1968: 145 (type data); Zucchi, 1979b: 263 (comm.); Norrbom, 1987: 243 (comm.); Carrejo & González, 1994: 88 (distr.); Norrbom et al., 1999b: 81 (cat.); Canal, 2010: 42 (distr.). parishi Stone, 1942b: 302. Type locality: Guyana, Bartica. HT F (USNM). Distr.: Brazil, Colombia, Costa Rica, Guyana, and Venezuela. Refs.: Aczél, 1950a: 223 (cat.); Foote, 1967: 14 (cat.); Gonzáles et al., 1988: 334 (distr.); Norrbom et al., 1999b: 81 (cat.); Martínez-Alava, 2007: 40 (distr.); Jesus et al., 2008: 135 (distr.); Canal, 2010: 43 (distr.). perdita Stone, 1942a: 76. Type locality: Brazil. HT F (USNM). Distr.: Brazil, Colombia, Panama, and Peru. Refs.: Aczél, 1950a: 223 (cat.); Foote, 1967: 14 (cat.); Korytkowski & Peña, 1968: 67 (redesc.); Norrbom et al., 1999b: 81 (cat.); Canal, 2010: 39 (distr.); Castañeda et al., 2010: 267 (distr.). pickeli Lima, 1934b: 542. Type locality: Brazil, Pernambuco, Tapera; Rio de Janeiro, Guaratiba. ST M/F (IOC). Distr.: Argentina, Brazil, Colombia, Costa Rica, Guyana, Panama, Trinidad and Tobago, and Venezuela. Refs.: Stone, 1942a: 63 (redesc.); Aczél, 1950a: 224 (cat.); Foote, 1967: 14 (cat.); Korytkowski & Peña, 1968: 59 (redesc.); Carrejo & González, 1994: 89 (distr.); Norrbom et al., 1999b: 81 (cat.); Carrejo & González, 1999: 115 (distr.); Patiño, 2002: 23 (distr.); Canal, 2010: 42 (distr.); Castañeda et al., 2010: 267 (distr.). quararibeae Lima, 1937a: 35. Type locality: Brazil, Bahia. HT F (IOC). Distr.: Brazil and Colombia. Refs.: Stone, 1942a: 44 (redesc.); Aczél, 1950a: 225 (cat.); Foote, 1967: 15 (cat.); Carrejo & González, 1994: 88 (distr.); Norrbom et al., 1999b: 82 (cat.); Canal, 2010: 42 (distr.). rheediae Stone, 1942a: 65. Type locality: Panama, Canal Zone, Barro Colorado Island. HT F (USNM). Distr.: Brazil, Colombia, Ecuador, Panama, Trinidad, and Venezuela. Refs.: Aczél, 1950a: 226 (cat.); Foote, 1967: 15 (cat.); Korytkowski & Peña, 1968: 55 (redesc.); Yepes & Vélez, 1989: 83 (distr.); Norrbom et al., 1999b: 82 (cat.); Martínez & Serna, 2005: 109 (distr.); Canal, 2010: 43 (distr.); Castañeda et al., 2010: 267 (distr.). serpentina Wiedemann, 1830: 521 (Dacus). Type locality: Brazil. T F (NHMW). Distr.: Argentina, Belize, Bolivia, Brazil, Costa Rica, Colombia, Ecuador, Guatemala, Guyana, Honduras, Mexico, Nicaragua, Panama, Peru, Surinam, Trinidad, United States, and Venezuela. Refs.: Bezzi, 1909: 286 (in key); Hendel, 1914c: 14 (in key), 16 (cat.); Fischer, 1932: 309 (distr.); Greene, 1934: 131 (in key), 142 (comm.); Lima, 1934b: 494 (redesc.); Hering, 1935: 227 (in key); Hendel, 1936: 72 (distr.); Hering, 1941: 136 (in key), 140 (comm.); Stone, 1942a: 27 (redesc.); Aczél, 1950a: 227 (cat.); Foote, 1967: 16 (cat.); Korytkowski & Peña, 1968: 49 (redesc.); Yepes & Vélez, 1989: 82 (distr.); Carrejo & González, 1994: 87 (distr.); Gonzáles et al., 1988: 334 (distr.); Norrbom et al., 1999b: 82 (cat.); Carrejo & González, 1999: 115 (distr.); Norrbom, 2002a: 393 (in key), 420 (redesc.); Martínez & Serna, 2005: 109 (distr.); Canal, 2010: 43 (distr.); Castañeda et al., 2010: 267 (distr.); Uchôa, 2012: 277 (distr.). shannoni Stone, 1942a: 23. Type locality: Peru, Loreto, Chimbote. HT M (USNM). Distr.: Brazil, Colombia, Peru, and Venezuela. Refs.: Aczél, 1950a: 228 (cat.); Foote, 1967: 16 (cat.); Korytkowski & Peña, 1968: 59 (redesc.); Norrbom, 1991: 104 (in key), 121 (redesc.); Norrbom et al., 1999b: 82 (cat.); Martínez-Alava, 2007: 40 (distr.); Deus et al., 2009: 727 (distr.); Canal, 2010: 42 (distr.). sinuosa Canal, 2010: 38. Type locality: Colombia, Tolima. Ibagué, El Placer, 1.448 m. HT F (MEN-UT). Distr.: Colombia. Refs.: Castañeda et al., 2010: 267 (distr.). CATALOGUE OF TEPHRITIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 605 sororcula Zucchi, 1979a: 39. Type locality: Brazil, São Paulo, Ribeirão Preto. HT F (MZUSP). Distr.: Brazil, Colombia, Ecuador, and Paraguay. Refs.: Norrbom et al., 1999b: 82 (cat.); Canal, 2010: 40 (distr.); Castañeda et al., 2010: 267 (distr.); Uchôa, 2012: 277 (distr.). striata Schiner, 1868: 264. Type locality: South America (Venezuela). LT F (NHMW). Distr.: Bolivia, Brazil, British Guiana, Colombia, Costa Rica, Ecuador, Honduras, Mexico, Panama, Peru, Trinidad, Tobago, Surinam, and Venezuela. Refs.: Bezzi, 1909: 285 (in key); Hendel, 1914c: 15 (in key), 16 (cat.), 19 (redesc.); Fischer, 1932: 309 (distr.); Greene, 1934: 132 (in key), 145 (comm.); Lima, 1934b: 504 (redesc.); Hering, 1941: 137 (in key), 140 (comm.); Stone, 1942a: 29 (redesc.); Aczél, 1950a: 229 (cat.); Foote, 1967: 16 (cat.); Hardy, 1968: 136 (type data); Korytkowski & Peña, 1968: 50 (redesc.); Zucchi, 1979b: 265 (comm.); Yepes & Vélez, 1989: 82 (distr.); Carrejo & González, 1994: 88 (distr.); Gonzáles et al., 1988: 334 (distr.); Norrbom et al., 1999b: 82 (cat.); Carrejo & González, 1999: 115 (distr.); Norrbom, 2002a: 391 (in key), 424 (redesc.); Patiño, 2002: 21 (distr.); Núñez Bueno et al., 2004: 7 (distr.); Martínez & Serna, 2005: 109 (distr.); Canal, 2010: 43 (distr.); Castañeda et al., 2010: 267 (distr.); Sarmiento et al., 2012: 28 (distr.); Uchôa, 2012: 278 (distr.); Wyckhuys et al., 2012: 94 (distr.). cancellaria Fabricius, 1805: 328 (Dictya). Type locality: America meridionali [Guyana]. T A (UZMC). Norrbom et al., 1999b: 82 (syn.). sylvicola Knab, 1915: 146. Type locality: Trinidad. ST F/M (USNM). Distr.: Colombia, Trinidad, and Venezuela. Refs.: Fischer, 1932: 309 (distr.); Greene, 1934: 135 (in key), 159 (comm.); Lima, 1934b: 537 (comm.); Stone, 1942a: 102 (redesc.); Foote, 1967: 16 (cat.); Norrbom et al., 1999b: 83 (cat.). tumida Stone, 1942a: 98. Type locality: Panama, La Campana. HT F (USNM). Distr.: Colombia, Costa Rica, Ecuador, Mexico, and Panama. Refs.: Foote, 1967: 17 (cat.); Carrejo & González, 1994: 90 (distr.); Norrbom et al., 1999b: 83 (cat.); Canal, 2010: 43 (distr.). velezi Norrbom, 1997: 156. Type locality: Colombia, Antioquia, Santo Domingo. HT F (USNM). Distr.: Colombia. Refs.: Canal, 2010: 42 (distr.). woodi Norrbom & Korytkowski, 2012: 163. Type locality: Costa Rica, Limón, Veragua Rainforest, 400-440 m. HT F (INBio). Distr.: Colombia, Costa Rica, and Panama. Genus Hexachaeta Loew Hexachaeta Loew, 1873: 219. Type species, Trypeta eximia Wiedemann (subs. des. Coquillett, 1910: 552). Refs.: Wulp, 1899: 402 (key to 3 spp.); Hendel, 1914b: 82 (in key); Hendel, 1914c: 5 (in key), 23 (comm.); Bates, 1933: 50 (comm.); Hering, 1941: 134 (key to 5 spp.); Aczél, 1950a: 192 (cat.); Lima, 1953b: 557 (rev. of eximia group); Lima, 1953a: 153 (rev. of amabilis group); Lima, 1954a: 277 (rev. of socialis group); Foote, 1967: 26 (cat.); Foote, 1980: 7 (in key), 31 (rev.); Foote et al., 1993: 206 (key to 3 spp.); Norrbom et al., 1999b: 158 (cat.); Hernández-Ortiz, 2006: 13 (taxonomy). Neohexachaeta Lima, 1953b: 566. Type species, guatemalensis Lima (orig. des.). Foote, 1980: 31 (syn.). amabilis Loew, 1873: 219 (Trypeta). Type locality: Mexico. ST M (MLUH). Distr.: Argentina, Belize, Bolivia, Brazil, Colombia, Costa Rica, Ecuador, Guatemala, Honduras, Mexico, Nicaragua, Panama, Paraguay, and Peru. Refs.: Hendel, 1914c: 23 (in key, cat.), 25 (redesc.); Hendel, 1936: 73 (distr.); Hering, 1941: 134 (in key), 135 (comm.); Aczél, 1950a: 193 (cat.); Lima & Leite, 1952: 297 (in key), 299 (redesc.); Lima, 1953a: 153 (rev.); Foote, 1967: 26 (cat.); Norrbom et al., 1999b: 158 (cat.); Hernández-Ortiz, 2002: 130 (distr.). amabilis var. oculata Hendel, 1914c: 23. Type locality: Peru, Urubamba, R., Rosalina. ST M (SMT, NHMW). (syn.). amabilis f. decolorata Lindner, 1928: 26. Type locality: Bolivia, Santa Cruz, Rio Tucabaca [Tucavaca]. HT M (SMN). (syn.). colombiana Lima, 1953b: 560. Type locality: Colombia, Condinamarca. HT M (USNM). Distr.: Colombia and Costa Rica. Refs.: Foote, 1967: 26 (cat.); Norrbom et al., 1999b: 158 (cat.); Hernández-Ortiz, 1999: 633 (redesc.). 606 · Zootaxa 4122 (1) © 2016 Magnolia Press SAVARIS ET AL. Genus Toxotrypana Gerstãcker Toxotrypana Gerstaecker, 1860: 191. Type species, curvicauda Gerstaecker (mon.). Refs.: Röder, 1891: 31 (comm.); Hendel, 1914b: 74 (in key); Hendel, 1914c: 4 (in key); Benjamin, 1934: 10 (redesc.); Aczél, 1950a: 180 (cat.); Blanchard, 1960: 228 (rev. of 6 spp.); Foote, 1967: 48 (cat.); Foote, 1980: 4 (in key), 48 (rev.); Yepes & Vélez, 1989: 74 (distr.); Norrbom et al., 1999b: 228 (cat.). Mikimyia Bigot, 1884: xxix. Type species, furcifera Bigot (mon.). (= curvicauda Gerstaecker). curvicauda Gerstaecker, 1860: 194. Type locality: Virgin Is., Insula St. Jean (St. John). T F (ZMHU). Distr.: Bahamas, Belize, Colombia, Costa Rica, Cuba, El Salvador, Guatemala, Honduras, Mexico, Nicaragua, Panama, United States, and Venezuela. Refs.: Röder, 1891: 31 (comm.); Hendel, 1914c: 10 (comm.); Benjamin, 1934: 11 (redesc.); Aczél, 1950a: 180 (cat.); Foote, 1967: 48 (cat.); Heppner, 1986: 1 (distr.); Norrbom et al., 1999b: 228 (cat.); Selman et al., 2012: 1 (distr.). furcifera Bigot, 1884: xxix. (Mikimyia). Brazil [probably erroneous]. HT M (UMO). (syn.). fairbatesi Munro, 1984: 160. Cuba, Soledad, Santa Clara. HT M (SANC). (syn.). littoralis Blanchard, 1960: 36. Type locality: Argentina, Corrientes. HT F (MACN). Distr.: Guatemala to Venezuela, following Andes to Argentina. Refs.: Foote, 1967: 48 (cat.); Norrbom et al., 1999b: 228 (cat.). Comm.: Toxotrypana littoralis is is newly registered from Colombia, based on 5 specimens (3 males and 2 females) in the National Museum of Natural History, Washington, DC. (USNM). These specimens originated from Cundinamarca, La Mesa, at an altitude of 575 m., collected in April 1989, by G. Sanchez, and were identified by Allen L. Norrbom (USNMENT00052674, USNMENT00214170, USNMENT00214171, USNMENT00214172, USNMENT00214173). Incertae Sedis Trypetinae Genus Pseudophorellia Lima Pseudophorellia Lima, 1934a: 139. Type species, maculata Lima (orig. des.). Refs.: Aczél, 1950a: 249 (cat.); Foote, 1967: 39 (cat.); Foote, 1980: 8 (in key), 44 (rev.); Norrbom et al., 1999b: 196 (cat.); Norrbom, 2006: 64 (rev.). anypsilon Norrbom, 2006: 79. Type locality: Panama, Parque Nacional Chagres, Altos de Pacora. HT F (USNM). Distr.: Colombia, Costa Rica, and Panama. distincta Norrbom, 2006: 93. Type locality: Colombia, Cundinamarca, Carpanta Reserve, 45.8 km E of Bogotá. HT M (USNM). Distr.: Colombia. flavicauda Norrbom, 2006: 98. Type locality: Colombia, Cauca, Río Micay Lopez, 90 m. HT F (FSCA). Distr.: Colombia. tica Norrbom, 2006: 119. Type locality: Costa Rica, Guanacaste, Parque Nacional Guanacaste, 9 km S Santa Cecilia, Estación Pitilla, 700 m. HT F (INBio). Distr.: Colombia, Costa Rica, French Guiana, and Panama. Subfamily Tephritinae Newman, 1934 [as Tephritites] Tribe Acrotaeniini Foote, Blanc & Norrbom, 1993 Genus Baryplegma Wulp Baryplegma Wulp, 1899: 416. Type species, gilva Wulp (mon.). Refs.: Hendel, 1914b: 95 (in key); Hendel, 1914c: 8 (in key), 58 (Pseudacrotaenia key to 5 spp.), 62 (comm.); Hering, 1941: 150 (Pseudacrotaenia key to 2 spp.); Aczél, 1950a: 274 (cat.); Foote, 1967: 17 (cat.); Foote, 1980: 15 (in key), 20 (rev.); Norrbom et al., 1999b: 104 (cat.). Pseudacrotaenia Hendel, 1914b: 98. Type species, Carphotricha vespillo Schiner (orig. des.). CATALOGUE OF TEPHRITIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 607 pertusa Bates, 1934: 14 (Pseudacrotaenia). Type locality: Costa Rica, San Mateo, Higuito. HT M (USNM). Distr.: Belize, Colombia, Costa Rica, Guatemala, Mexico, Nicaragua, Panama, and Venezuela. Refs.: Aczél, 1950a: 270 (cat.); Foote, 1967: 37 (cat.); Norrbom et al., 1999b: 105 (cat.). Genus Neotaracia Foote Neotaracia Foote, 1978: 31. Type species, Acrotaenia imox Bates (orig. des.). Refs.: Foote, 1979: 175 (rev. of 3 spp.); Foote, 1980: 13 (in key), 37 (rev.); Norrbom et al., 1999b: 172 (cat.). imox Bates, 1934: 11 (Acrotaenia). Type locality: Costa Rica, San Mateo, Higuito. HT F (USNM). Distr.: Belize, Colombia, Costa Rica, Ecuador, El Salvador, Guatemala, Honduras, Mexico, Nicaragua, Panama, Peru, Trinidad, and Venezuela. Refs.: Aczél, 1950a: 268 (cat.); Foote, 1967: 5 (cat.); Foote, 1978: 31(redesc.); Foote, 1979: 175 (in key, redesc.); Norrbom et al., 1999b: 172 (cat.); Norrbom et al., 2013: 21 (distr.). Genus Tetreuaresta Hendel Tetreuaresta Hendel, 1928: 368. Type species, Trypeta obscuriventris Loew (orig. des.). Refs.: Bates, 1933: 53 (comm.); Bates, 1934: 15 (comm.); Hering, 1941: 154 (key to 5 spp.); Aczél, 1950a: 276 (cat.); Foote, 1967: 43 (cat.); Steyskal, 1972b: 404 (key to 15 spp.); Foote, 1980: 13, 15 (in key), 47 (rev.); Norrbom et al., 1999b: 224 (cat.). heringi Norrbom, in Norrbom et al. 1999b: 224. Nom. nov. rufula Hering. Distr.: Colombia. Refs.: Hering, 1942b: 286 (orig. desc.); Aczél, 1950a: 278 (cat.); Foote, 1967: 44 (cat.); Norrbom et al., 1999b: 224 (cat.). rufula Hering, 1942b: 286. Preocc. by Wulp, 1899. Type locality: Colombia, “Cordillere, tierra caliente”. HT M (ZMHU). obscuriventris Loew, 1873: 313 (Trypeta). Type locality: Brazil. T F (NHMW). Distr.: Brazil, Colombia, Greater Antilles; introduced Hawaii, Fiji and Tonga. Refs.: Hendel, 1914c: 73 (redesc.); Hendel, 1928: 368 (comm.); Hering, 1941: 154 (in key), 155 (comm.); Aczél, 1950a: 277 (cat.); Foote, 1967: 44 (cat.); Norrbom et al., 1999b: 224 (cat.). columbiana Enderlein, 1911: 431 (Euaresta). Type locality: Colombia, Hacienda, Pehlke. HT M (PAN). (syn.). catharinensis Enderlein, 1911: 431 (Euaresta). Type locality: Brazil, Santa Catharina [Santa Catarina]. LT F (PAN). (syn.). Genus Tomoplagia Coquillett Tomoplagia Coquillett, 1910: 591. Nom. nov. for Plagiotoma Loew. Refs.: Coquillett, 1899: 268 (in key); Hendel, 1914b: 88 (in key); Hendel, 1914c: 6 (in key), 33 (comm.); Benjamin, 1934: 32 (redesc.); Hering, 1941: 124 (in key), 144 (key to 9 spp.); Aczél, 1950a: 241 (cat.); Aczél, 1955a: 323 (redesc.), 329 (key to 43 spp.); Aczél, 1955b: 155 (key to 45 spp.); Foote, 1967: 45 (cat.); Foote, 1980: 8 (in key), 48 (rev.); Foote et al., 1993: 411 (key to 2 spp.); Prado & Lewinsohn, 1994: 669 (taxonomy of 9 spp.); Norrbom et al., 1999b: 226 (cat.). Plagiotoma Loew, 1873: 252. Type species, Trypeta obliqua Say (subs. des. Coquillett, 1910: 591). (preocc. Dujardin, 1841). arsinoe Hering, 1942b: 285. Type locality: Colombia, “Cordillere, tierra templada”. HT M (ZMHU). Distr.: Colombia. Refs.: Aczél, 1950a: 241 (cat.); Aczél, 1955a: 333 (in key); Aczél, 1955b: 163 (in key); Foote, 1967: 45 (cat.); Norrbom et al., 1999b: 226 (cat.). arsinoë Hering, 1942b: 285. (incosp. arsinoe Hering). pleuralis Hendel, 1914c: 39. Type locality: Peru, Callanga, Río Charape. HT M (MNM). Distr.: Argentina, Ecuador, and Peru. Refs.: Hering, 1941: 144 (in key), 145 (comm.); Aczél, 1950a: 245 (cat.); Aczél, 1955a: 332 (in key), 381 (redesc.); Aczél, 1955b: 161 (in key); Foote, 1967: 47 (cat.); Norrbom et al., 1999b: 227 (cat.). Comm.: 608 · Zootaxa 4122 (1) © 2016 Magnolia Press SAVARIS ET AL. Tomoplagia pleuralis is a new record for Colombia, based on one male specimen collected in 1914 (unspecified locality) by H.A. Parish. The specimen was identified by Martín Aczél and deposited at the National Museum of Natural History, Washington, DC (USNM) (USNMENT00051053). Tribe Eutretini Munro, 1952 Genus Dictyotrypeta Hendel Dictyotrypeta Hendel, 1914b: 93. Type species, syssema Hendel (orig. des.). Refs.: Hering, 1941: 124 (in key), 148 (comm.); Aczél, 1950a: 259 (cat.); Foote, 1967: 21 (cat.); Foote 1980: 12 (in key), 25 (rev.); Norrbom et al., 1999b: 139 (cat.). Dictyotrypeta Hendel, 1914c: 49. Type species, syssema Hendel (orig. des.). (preocc. Hendel, 1914b: 93). syssema Hendel, 1914b: 93. Type locality: Peru. T A (SMT). Distr.: Colombia, Costa Rica, Ecuador, and Peru. Refs.: Hering, 1941: 149 (comm.); Aczél, 1950a: 259 (cat.); Foote, 1967: 21 (cat.); Norrbom et al., 1999b: 139 (cat.). syssema Hendel, 1914c: 50. Type locality: Peru, Mamara; Cuzco, 3.600 m. ST M/F (SMT). (preocc. Hendel 1914b: 93). Genus Eutreta Loew Eutreta Loew, 1873: 276. Type species, Trypeta sparsa Wiedemann (subs. des. Coquillett, 1910: 543). Refs.: Coquillett, 1899: 268 (in key); Hendel, 1914b: 93 (in key); Hendel, 1914c: 8 (in key), 54 (comm.); Hering, 1941: 149 (comm.); Aczél, 1950a: 264 (cat.); Foote, 1967: 24 (cat.); Stoltzfus, 1977: 369 (rev. of 30 spp.); Foote 1980: 12 (in key), 29 (rev.); Foote et al., 1993: 180 (key to subgenera & 16 spp.); Norrbom et al., 1999b: 151 (cat.). Phasmatocephala Hering, 1937: 297. Type species, Eutreta rhinophora Hering (orig. des.). Proposed as a subgenus. Icaria Schiner, 1868: 276. Type species, Trypeta sparsa Wiedemann (orig. des.) (preocc. Saussure, 1853). aczeli Lima, 1954b: 175. Type locality: Brazil. HT F (IOC). Distr.: Brazil and Colombia. Refs.: Foote, 1967: 24 (cat.); Norrbom et al., 1999b: 151 (cat.). obliqua Stoltzfus, 1977: 386. Type locality: Colombia, Rio Tocata (Jacota ?). HT M (BMNH). Distr.: Colombia and Ecuador. Refs.: Norrbom et al., 1999b: 152 (cat.). rhinophora Hering, 1937: 297. Type locality: Costa Rica, San José. NT M (PAN). Neotype designated by Stoltzfus, 1977: 387. Distr.: Belize, Colombia, Costa Rica, El Salvador, Guatemala, Honduras, Mexico, Nicaragua, Panama, and Venezuela. Refs.: Aczél, 1950a: 266 (cat.); Foote, 1967: 25 (cat.); Norrbom et al., 1999b: 152 (cat.). Tribe Myopitini Bezzi, 1910 [as Myiopitininae] Genus Neomyopites Freidberg & Norrbom Neomyopites Freidberg & Norrbom, 1999: 607. Type species, Euribia aerea Hering (orig. des.). aereus Hering, 1942c: 479 (Euribia). Type locality: Costa Rica, Cartago, La Suiza de Turrialba. HT M (MNM). Distr.: Colombia, Costa Rica, and Guatemala. Refs.: Aczél, 1950a: 183 (cat.); Foote, 1967: 54 (cat.); Norrbom et al., 1999b: 244 (cat.); Freidberg & Norrbom, 1999: 584 (new comb.). agnatus Hering, 1942a: 1 (Euribia). Type locality: Colombia. “Cordillere, terra fria”. HT F (ZMHU). Distr.: Colombia. Refs.: Aczél, 1950a: 183 (cat.); Foote, 1967: 54 (cat.); Norrbom et al., 1999b: 244 (cat.); Freidberg & Norrbom, 1999: 584 (new comb.). CATALOGUE OF TEPHRITIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 609 columbianus Hering, 1942a: 2 (Euribia). Type locality: Colombia. HT F (ZMHU). Distr.: Colombia. Refs.: Aczél, 1950a: 183 (cat.); Foote, 1967: 54 (cat.); Norrbom et al., 1999b: 244 (cat.); Freidberg & Norrbom, 1999: 584 (new comb.). confertus Walker, 1853: 379 (Trypeta). Type locality: Colombia, Bolivar, Cartagena. LT F (BMNH). Lectotype designation by inference of holotype by Foote, 1964: 319. Distr.: Colombia. Refs.: Foote, 1967: 54 (cat.); Norrbom et al., 1999b: 244 (cat.); Freidberg & Norrbom, 1999: 586 (list). Comm.: Freidberg & Norrbom (1999) treated this as an unplaced species of Myopitini, that “probably belongs in Neomyopites)”. It is here formally transferred (new comb.). cordilleranus Steyskal, 1979: 45 (Urophora). Type locality: Colombia, Cundinamarca, Guasca. HT M (USNM). Distr.: Colombia and Ecuador. Refs.: Norrbom et al., 1999b: 244 (cat.); Freidberg & Norrbom, 1999: 584 (new comb.). euryparius Steyskal, 1979: 49 (Urophora). Type locality: Colombia, Cundinamarca, Paramo de Cruz Verde, just E of Bogota, 3.150 m. HT M (AMNH). Distr.: Colombia. Refs.: Norrbom et al., 1999b: 244 (cat.); Freidberg & Norrbom, 1999: 584 (new comb.). melanops Steyskal, 1979: 51 (Urophora). Type locality: Venezuela, Cano del Tigre. HT M (USNM). Distr.: Colombia and Venezuela. Refs.: Norrbom et al., 1999b: 245 (cat.); Freidberg & Norrbom, 1999: 584 (new comb.). Genus Rhynencina Johnson Rhynencina Johnson, 1922: 24. Type species, longirostris Johnson (orig. des.). Refs.: Norrbom et al., 1999b: 203 (cat.); Steck et al., 2003: 543 (comm.). Aleomyia Phillips, 1923: 123. Type species, alpha Phillips (orig. des.) (= longirostris Johnson). dysphanes Steyskal, 1979: 47 (Urophora). Type locality: Ecuador, Pichincha, Cotocollao, 10000 ft. HT M (USNM). Distr.: Colombia and Ecuador. Refs.: Norrbom et al., 1999b: 203 (cat., comb.); Freidberg & Norrbom, 1999: 584; Steck et al., 2003: 542 (distr.). emphanes Steyskal, 1979: 48 (Urophora). Type locality: Colombia, Cundinamarca, Guasca, 35 km. NE Bogota, 1.800 m. HT F (AMNH). Distr.: Colombia. Refs.: Norrbom et al., 1999b: 203 (cat., comb.); Freidberg & Norrbom, 1999: 584 (host). Tribe Noeetini Norrbom & Korneyev, 1999 Genus Ensina Robineau-Desvoidy Ensina Robineau-Desvoidy, 1830: 751. Type species, scorzonerae Robineau-Desvoidy (subs. des. Foote & Freidberg, 1981: 30) (= sonchi Linnaeus). Refs.: Hendel, 1914b: 96 (in key); Hendel, 1914c: 9 (in key), 65 (comm.); Malloch, 1933: 273 (comm.); Hering, 1941: 152 (key to 3 spp.); Aczél, 1950a: 274 (cat.); Foote, 1980: 15 (in key), 27 (rev.); Norrbom et al., 1999b: 144 (cat.). Protensina Hendel, 1914b: 95. Type species, longiceps Hendel (orig. des.). Protensina Hendel, 1914c: 64. Type species, longiceps Hendel (orig. des.). (preocc. Hendel, 1914b: 95). hyalipennis Hennig, 1940: 13 (Protensina). Type locality: Peru, Cuzco, Cuzco, 3.500 m. HT M (SMT). Distr.: Bolivia, Colombia, Chile, Ecuador, Peru, and Venezuela. Refs.: Hering, 1941: 152 (in key), 153 (comm.); Aczél, 1950a: 275 (cat.); Steyskal, 1970: 158 (comm.); Norrbom et al., 1999b: 144 (cat.). 610 · Zootaxa 4122 (1) © 2016 Magnolia Press SAVARIS ET AL. Tribe Tephritini Newman, 1834 Genus Dyseuaresta Hendel Dyseuaresta Hendel, 1928: 368. Type species, Euaresta adelphica Hendel (orig. des.). Refs.: Benjamin, 1934: 51 (redesc.); Bates, 1934: 15 (comm.); Hering, 1941: 156 (comm.); Aczél, 1950a: 279 (cat.); Foote, 1967: 22 (cat.); Foote, 1980: 11 (in key), 26 (rev.); Norrbom et al., 1999b: 142 (cat.). mexicana Wiedemann, 1830: 511 (Trypeta). Type locality: Mexico. LT M (ZMHU). Lectotype designation by inference of holotype by Loew, 1873: 319. Distr.: Belize, Colombia, Costa Rica, Cuba, Guatemala, El Salvador, Honduras, Mexico, Nicaragua, Panama, Puerto Rico, United States, Venezuela, and West Indies. Refs.: Loew, 1873: 317 (redesc.); Benjamin, 1934: 51 (redesc.); Aczél, 1950a: 280 (cat.); Foote, 1967: 22 (cat.); Norrbom et al., 1999b: 142 (cat.); Norrbom et al., 2010: 22 (host, distr.). melanogastra Loew, 1862: 90 (Trypeta). Type locality: Cuba. ST M/F (MCZ). (syn.). plesia Curran, 1928: 71 (Euaresta). Type locality: Puerto Rico, Coamo, Springs. HT F (AMNH). (syn.). Genus Euaresta Loew Euaresta Loew, 1873: 296. Type species, Trypeta festiva Loew (subs. des. Coquillett, 1910: 540). Refs.: Coquillett, 1899: 268 (in key); Hendel, 1914b: 96 (in key); Hendel, 1914c: 6 (in key), 70 (comm.); Benjamin, 1934: 49 (redesc.); Aczél, 1950a: 281 (cat.); Aczél, 1952a: 154 (rev. of 4 spp.); Foote, 1967: 23 (cat.); Foote, 1980: 15 (in key), 27 (rev.); Foote et al., 1993: 143 (key to 8 spp.); Norrbom, 1993: 207 (key to 8 spp.); Norrbom et al., 1999b: 144 (cat.). Camaromyia Hendel, 1914b: 95. Type species, Trypeta bullans Wiedemann (orig. des.). Setigeresta Benjamin, 1934: 50. Type species, Trypeta aequalis Loew (orig. des.). Camaromyia Hendel, 1914c: 63. Type species, Trypeta bullans Wiedemann (orig. des.) (preocc. Hendel, 1914b: 95). reticulata Hendel, 1914c: 81 (Trypanea). Type locality: Peru, Lares Valley 2.000 m.; Callabamba 3.000 m.; Arequipa, Arequipa; Cuzco, Sicuani & Cuzco 3.300 m.; Junin, Tarma 3.000 m. ST M/F (SMT, MNM). Distr.: Bolivia, Colombia, Ecuador, and Peru. Refs.: Hering, 1941: 158 (comm.); Aczél, 1950a: 284 (cat.); Foote, 1967: 35 (cat.); Steyskal, 1972a: 130 (distr.); Norrbom, 1993: 205 (redesc.), 208 (in key); Norrbom et al., 1999b: 145 (cat.); Norrbom et al., 2013: 21 (distr.). apicata Becker, 1919: 195 (Tephritis). Type locality: Ecuador, Riobamba, Tulcan & Troya. ST M/F (MNHNP). Norrbom 1993: 205 (syn.). toba Lindner, 1928: 29 (Camaromyia). Type locality: Argentina, Formosa, Misión Tacaaglé. LT M (SMN). Lectotype designated by Norrbom, 1993: 200. Distr.: Argentina, Brazil, Colombia, Chile, El Salvador, Peru, Uruguay, and Venezuela. Refs.: Malloch, 1933: 274 (in key, comm.); Aczél, 1950a: 296 (cat.); Aczél, 1952a: 165 (distr.); Foote, 1967: 24 (cat.); Norrbom, 1993: 200 (redesc.), 208 (in key); Norrbom et al., 1999b: 145 (cat.). Genus Homoeothrix Hering Homoeothrix Hering, 1944: 7. Type species, Euribia lindigi Hendel (orig. des.). Refs.: Hering, 1947: 7 (emended); Aczél, 1950a: 284 (cat.); Foote, 1967: 28 (cat.); Foote, 1980: 15 (in key), 32 (rev.); Norrbom et al., 1999b: 160 (cat.); Norrbom et al., 2010: 24 (distr.). Homoeothrix Hering, 1947: 7. (emend. Homoeothrlx Hering). Homoeothrlx Hering, 1944: 7. (incosp. Homoeothrix Hering). aberrans Schiner, 1868: 273 (Oxyphora). Type locality: Colombia. LT M (NHMW). Lectotype designated by Hardy, 1968: 138. Distr.: Colombia and Venezuela. Refs.: Aczél, 1950a: 285 (cat.); Foote, 1967: 32 (cat.); Hardy, 1968: 138 (type data); Norrbom et al., 1999b: 172 (cat.); Norrbom et al., 2010: 24 (new comb.). CATALOGUE OF TEPHRITIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 611 Genus Xanthaciura Hendel Xanthaciura Hendel, 1914b: 86. Type species, Trypeta chrysura Thomson (orig. des.). Refs.: Hendel, 1914c: 7 (in key), 45 (comm.); Bates, 1933: 55 (comm.); Malloch, 1933: 266 (key to 7 spp.); Benjamin, 1934: 43 (redesc.); Hering, 1941: 147 (key to 4 spp.); Aczél, 1950a: 253 (cat.); Aczél, 1950b: 117 (key to 12 spp.); Aczél, 1952b: 253 (key to 15 spp.); Foote, 1967: 54 (cat.); Foote, 1980: 15 (in key), 50 (rev.); Foote et al., 1993: 482 (key to 4 spp.); Norrbom et al., 1999b: 246 (cat.). Tetraciura Hendel, 1914b: 90. Type species, quadrisetosa Hendel (orig. des.). Eucosmoptera Phillips, 1923: 131. Type species, Aciura testraspina Phillips (subs. des. Bates, 1933: 55). Proposed as a subgenus. Chrysaciura Aczél, 1953: 188. Type species, bipuncta Aczél (orig. des.). Tetraciura Hendel, 1914c: 48. Type species, quadrisetosa Hendel (orig. des.) (preocc. Hendel, 1914b: 90). Xanthaciura Hendel, 1914c: 45. Type species, Trypeta chrysura Thomson (orig. des.) (preocc. Hendel, 1914b: 86). chrysura Thomson, 1869: 580 (Trypeta). Type locality: Brazil, Rio de Janeiro. ST M/F (NRS). Distr.: Argentina, Belize, Bolivia, Brazil, Costa Rica, Colombia, Ecuador, Guatemala, Guyana, Honduras, Mexico, Nicaragua, Panama, Peru, Surinam, Trinidad, United States, and Venezuela. Refs.: Hendel, 1914c: 46 (in key, cat.), 47 (redesc.); Hering, 1941: 147 (in key, comm.); Aczél, 1950a: 254 (cat.); Foote, 1967: 55 (cat.); Norrbom et al., 1999b: 246 (cat.). erosa Enderlein, 1911: 458 (Aciura). Type locality: Brazil, Santa Catharina [Santa Catarina]; Colombia. ST F (PAN). (syn.). contracta Hering, 1937: 298. Type locality: Costa Rica, 8 km. W of San Jose, Farm La Caja. ST M/F (BMNH). (syn.). mallochi Aczél, 1950b: 125. Type locality: Argentina, Chaco, Colonia Benitez. HT M (IML). Distr.: Argentina, Brazil, Costa Rica, Colombia, Ecuador, Panama, and Peru. Refs.: Foote, 1967: 55 (cat.); Norrbom et al., 1999b: 247 (cat.). unipuncta Malloch, 1933: 268. Type locality: Paraguay, Encarnación. HT M (BMNH). Distr.: Argentina, Brazil, Costa Rica, Colombia, Ecuador, El Salvador, Guatemala, Honduras, Nicaragua, Panama, Paraguay, and Trinidad. Refs.: Aczél, 1950a: 256 (cat.); Foote, 1967: 56 (cat.); Norrbom et al., 1999b: 247 (cat.); Prado et al., 2002: 1024 (distr.). Family Tephritidae Incertae Sedis Tephritis unifasciata Macquart, 1835: 465. Type locality: Colombia. T M (MNHNP). Unrecognized, ST apparently lost. Distr.: Colombia. Refs.: Norrbom et al., 1999b: 250 (cat.). TABLE 1. Tephritidae species recorded from Colombia. Subfamily Tribe Blepharoneurinae Genus Species Author/year Blepharoneura furcifer Hendel, 1914 quadristriata Wulp, 1899 Dacinae Dacini Ceratitis capitata (Wiedemann, 1824) Trypetinae Carpomyini Cryptodacus ornatus Norrbom, 1994 Rhagoletis pomonella* (Walsh, 1867) Zonosemata scutellata (Hendel, 1936) Anastrepha acuminata Canal, 2010 alveata Stone, 1942 anomoiae Norrbom, 2002 Toxotrypanini ...Continued on next page 612 · Zootaxa 4122 (1) © 2016 Magnolia Press SAVARIS ET AL. TABLE 1. (Continued) Subfamily Tribe Genus Species Author/year antunesi Lima, 1938 bahiensis binodosa* Lima, 1937 Stone, 1942 carreroi Canal, 2010 caudata Stone, 1942 coronilli Carrejo & González, 1993 curitis Stone, 1942 debilis Stone, 1942 dissimilis Stone, 1942 distincta Greene, 1934 doryphoros Stone, 1942 flavipennis Greene, 1934 fraterculus (Wiedemann, 1830) grandicula Norrbom, 1991 grandis (Macquart, 1846) hendeliana Lima, 1934 isolata Norrbom & Korytkowski, 2009 lanceola Stone, 1942 leptozona Hendel, 1914 limae Stone, 1942 loewi Stone, 1942 magna Norrbom, 1997 manihoti Lima, 1934 manizaliensis Norrbom & Korytkowski, 2005 matertela Zucchi, 1979 montei Lima, 1934 mucronota Stone, 1942 obliqua (Macquart, 1835) ornata Aldrich, 1925 palae Stone, 1942 pallidipennis Greene, 1934 panamensis Greene, 1934 parallela (Wiedemann, 1830) parishi Stone, 1942 perdita Stone, 1942 pickeli Lima, 1934 quararibeae Lima, 1937 rheediae Stone, 1942a serpentina (Wiedemann, 1830) ...Continued on next page CATALOGUE OF TEPHRITIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 613 TABLE 1. (Continued) Subfamily Tribe Genus Species Author/year shannoni Stone, 1942 sinuosa Canal, 2010 sororcula Zucchi, 1979 striata Schiner, 1868 sylvicola Knab, 1915 tumida Stone, 1942 velezi Norrbom, 1997 woodi Norrbom & Korytkowski, 2012 amabilis (Loew, 1873) colombiana Lima, 1953 curvicauda Gerstaecker, 1860 littoralis Blanchard, 1960 anypsilon Norrbom, 2006 distincta Norrbom, 2006 flavicauda Norrbom, 2006 tica Norrbom, 2006 Baryplegma pertusa (Bates, 1934) Neotaracia imox (Bates, 1934) Tetreuaresta heringi Hering, 1942 obscuriventris (Loew, 1873) arsinoe Hering, 1942 pleuralis Hendel, 1914 Dictyotrypeta syssema Hendel, 1914 Eutreta aczeli Lima, 1954 obliqua Stoltzfus, 1977 rhinophora Hering, 1937 aereus (Hering, 1942) agnatus (Hering, 1942) columbianus (Hering, 1942) confertus (Walker, 1853) cordilleranus (Steyskal, 1979) euryparius (Steyskal, 1979) melanops (Steyskal, 1979) dysphanes (Steyskal, 1979) emphanes (Steyskal, 1979) Hexachaeta Toxotrypana Incertae Sedis Trypetinae Tephritinae Pseudophorellia Acrotaeniini Tomoplagia Eutretini Myopitini Neomyopites Rhynencina Noeetini Ensina hyalipennis (Hennig, 1940) Tephritini Dyseuaresta mexicana (Wiedemann, 1830) Euaresta reticulata (Hendel, 1914) toba (Lindner, 1928) aberrans (Schiner, 1868) Homoeothrix ...Continued on next page 614 · Zootaxa 4122 (1) © 2016 Magnolia Press SAVARIS ET AL. TABLE 1. (Continued) Subfamily Tribe Tephritidae Incertae Sedis Genus Species Author/year Xanthaciura chrysura (Thomson, 1869) mallochi Aczél, 1950 unipuncta Malloch, 1933 unifasciata Macquart, 1835 Tephritis * Occurrence in the country is doubtful. Acknowledgements We thank the editors, Marta Wolff and Claudio J. B. Carvalho, for inviting us to complete the fruit fly component of this catalog. We also thank Silvio S. Nihei for reviewing the manuscript. USDA is an equal opportunity provider and employer. We thank CNPq for financial support for this study (Process number 200277/2015-4). References Aczél, M.L. (1950a) Catalogo de la familia ‘Trypetidae’ (Dipt. Acalypt.) de la región Neotropical. Acta Zoologica Lilloana, 7, 177–328. Aczél, M.L. (1950b) A revision of the genus ‘Xanthaciura‘ Hendel (Trypetidae. Dipt.) based on Argentine species. Acta Zoologica Lilloana, 8, 111–146. Aczél, M.L. (1952a) El genero Euaresta Loew (= Camaromyia Hendel) en la región Neotropical. 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CATALOGUE OF TEPHRITIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 621 Zootaxa 4122 (1): 622–634 http://www.mapress.com/j/zt/ Copyright © 2016 Magnolia Press Article ISSN 1175-5326 (print edition) ZOOTAXA ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.4122.1.51 http://zoobank.org/urn:lsid:zoobank.org:pub:93FAE1F9-AF42-48B1-A010-6CB6C85CABA2 FAMILY LAUXANIIDAE VERA CRISTINA SILVA UNESP - Univ Estadual Paulista, Faculdade de Ciências Agrárias e Veterinárias, Departamento de Morfologia e Fisiologia Animal; Via de Acesso Prof. Paulo Donato Castellane, s/n; 14.806-900 Jaboticabal, SP, Brazil. E-mail: vcsilva@fcav.unesp.br Abstract An updated Catalogue of the Lauxaniidae of Colombia is presented. This acalyptratae family is poorly known in Colombia, with only 36 described species in 33 genera. This paper expands the distribution of 24 species to Colombia. At total, 63 species are reported here for Colombia. Key words: catalogue, Colombia, Diptera, distribution, Neotropical region Introduction Lauxaniidae is one of the largest Diptera Schizophora families, very common in tropical areas. Adults may be small to moderately large (2-11 mm), varied in color patterns, often with marks, spots, bands, or reticulating patterns (Shewell 1987, Gaimari & Silva 2010b). Lauxaniidae larvae frequently are recorded as saprophages, found in a diverse array of decaying plant matter (Miller 1977). A few species have been reported to be phytophagous, living in root collars, stems and leaves of clover (Tripholium; Oxalidaceae), in the ovaries of violets (Viola; Violaceae), and in the leaf–like phylloclades of prickly pear (Opuntia; Cactaceae) (Miller 1977) and in the flowerheads of Asteraceae (Silva & Mello 2008). Some adults graze fungal hyphae and spores on leaves, using special labial structures, like prongs and scoops, making them one of the few higher flies able to feed on solid particles (Mello & Silva 2007; Marshall 2012). The family comprises about 1,550 valid species (Kim 1994). In the Neotropical region there are nearly 400 described species in 53 genera (Gaimari & Silva 2010b). The monophyly of the family is well supported by several autapomorphies (McAlpine, 1989). It is included in the Lauxanioidea, together with Chamaemyiidae and Celyphidae; recently the enigmatic family Eurychoromyiidae was revised to subfamily status (Gaimari & Silva, 2010a; Gaimari, 2011). Stuckenberg (1971) proposed the subfamily Homoneurinae for a group of genera based on the presence of costal setulae extending to the apex of wing vein R4+5. Shewell (1977) proposed the subfamily Lauxaniinae for the remainder of the family, with costal setulae ending just beyond the apex of vein R2+3 and a single midtibial spur. The conditions found in Lauxaniinae are clearly plesiomorphic, and are likely part of the groundplan for the entire Muscomorpha (McAlpine, 1989), and both Stuckenberg (1971) and Shewell (1977) acknowledged the likely paraphyly for the group. Lauxaniids are uncommon in the described fossil record. Six species are known primarily from the Eocene/ Oligocene Baltic amber, but undescribed species are also known from Oligocene Dominican amber (Evenhuis, 1997). Considering that a conspectus of the Neotropical Laxaniidae is in progress by this author and S. D. Gaimari, including a catalogue to all species in the region, the data presented here represents the actual taxonomic knowledge for the family at the Neotropical Region and should be noted that some of the taxa here will have their taxonomic status changed after the publication of the conspectus. Notwithstanding this, we provided a list that is intended to help all people interested in the Colombian Diptera. The Colombian fauna of lauxaniids comprises at the moment 36 described species representing only a small fraction of the known fauna, with at least 63 species recognized (Table 1). The data provided in this catalogue, 622 Accepted by S. Nihei: 10 Apr. 2015; published: 14 Jun. 2016 along with data gathered from literature, represent the main result of the study of specimens provided by collection efforts sponsored by the Humboldt Institute and NSF to M. Sharkey (grant number DEB-0205982). Many undescribed species are discovered and they are indicated here, also in few genera an undetermined species is reported because we believe it is important to know that some genus is represented in the country, although by an unnamed species. All genus- and species-group names are listed alphabetically. Valid names are in bold type at the first mention. Synonyms when mentioned subsequently are italicized. Each published name including synonyms is given in a chronological sequence according to the scheme: the name of the species, author(s), year of description, page, genus in the original description (if different from the valid one), type locality (or localities in the case of syntypes; updated name into brackets), information about the primary types and their deposition (and, if a lectotype, a reference to the author and date of the designation), known geographic distribution. At the end we present a table with all species found in Colombia, including rank and author (Table 1). Acronyms used for the depositories AMNH—American Museum of Natural History, New York, USA CASC—California Academy of Sciences, San Francisco, USA IAHB—Instituto de Investigación de Recursos Biológicos Alexander von Humboldt, Bogotá, Colombia MCZ—Museum of Comparative Zoology, Harvard University, Cambridge, USA MNHN—Muséum National d’Histoire Naturelle, Paris, France MZUSP—Museu de Zoologia, Universidade de São Paulo, São Paulo, Brazil NHM—The Natural History Museum, London, U.K. NMW—Naturhistorisches Museum, Vienna, Austria NRM—Naturhistoriska Riksmuseum, Stockholm, Sweden OUM—Oxford University Museum of Natural History, Oxford, U.K. USNM—National Museum of Natural History, Washington, USA ZMB—Museum für Naturkunde der Humboldt-Universität, Berlin, Germany List of abbreviations cf.—confer comm.—comments descr.—description diag.—diagnosis distrib.—geographic distribution fig. / figs—figure / figures lec. des.—lectotype designation mon.—monotypy misident.—misidentification n. comb—new combination nom. nov.—nomen novum orig. des.—by original designation preocc.—preoccupied redesc.—redescription ref. / refs.—reference /references rev.—revision subg.—subgenus subs. des.—by subsequent designation syn.—synonym HT—Holotype CATALOGUE OF LAUXANIIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 623 LT—Lectotype ST—Syntype F (f#)—Female M (m#)—Male Catalogue of Lauxaniidae of Colombia Family Lauxaniidae Macquart, 1835 Subfamily Eurychoromyiinae Hendel, 1910 Genus Euryhendelimyia Gaimari & Silva Euryhendelimyia Gaimari & Silva, 2010a: 14. Type species, E. schlingeri Gaimari & Silva (orig. des.). schlingeri Gaimari & Silva, 2010a: 14. Type locality: Colombia, Nariño, 25 mi SW Mocoa, 2080 m [Putumayo]. HT M (CASC). Distrib.—Colombia (Putumayo, according to Gaimari & Silva, 2010a). Genus Exalla Gaimari Exalla Gaimari, 2011: 595. Type species, E. shewelli Gaimari (orig. des.). macalpinei Gaimari, 2011: 603. Type locality: Colombia, Nariño, R. N. La Planada, Via Hondón. HT M (IAHB). Distrib.—Colombia (Nariño). shewelli Gaimari, 2011: 607. Type locality: Colombia, Nariño, R. N. La Planada, Via Hondón. HT M (IAHB). Distrib.—Colombia (Nariño). Subfamily Lauxaniinae Shewell, 1977 Genus Agriphoneura Hendel Agriphoneura Hendel, 1925: 137. Type species, A. fumipennis Hendel (mon.). undescribed sp. 1—Locality: Colombia (Risaralda). Genus Allogriphoneura Hendel Allogriphoneura Hendel, 1925: 130. Type species, A. nigromaculata Hendel (orig. des.) = A. porcaria (Fabricius). Ref.— Hendel, 1936: 86 (key). porcaria (Fabricius, 1805: 204; Scatophaga). Type locality: "America meridionalis" (Guyana). Type ZMUC. Distrib.—Colombia (Putumayo), Guyana, Brazil, Peru, Bolivia. Ref.—Hendel, 1933: 76. nigromaculata Hendel, 1925: 130. Ref.—Hendel, 1933: 76 (syn.). Type locality: Peru, Pichis, Porto Yessup. ST (NHMW). 624 · Zootaxa 4122 (1) © 2016 Magnolia Press SILVA Genus Allominettia Hendel Allominettia Hendel, 1925: 126. Type species, A. maculatifrons Hendel (orig. des.) = xanthiceps Williston. Chaetominettia Malloch, 1926: 9. Type species, C. spinitibia Malloch (orig. des.). Refs.—Malloch, 1933: 359; Curran, 1934b: 320, fig. 31 (head); Stuckenberg, 1971: 511 (syn.) Tibiominettia Hendel, 1932: 103. Type species, T. setitibia Hendel (orig. des.). Ref.—Hendel 1936: 84; Stuckenberg, 1971: 511 (syn.). corollae (Fabricius, 1805: 331; Dictya). Type locality: "America meridionalis" (Brazil, cf. Wiedemann, 1830: 452). Type ? (probable ZMUC). Ref.—Hendel, 1925: 141 (descr., distrib.). Distrib.—Honduras, Costa Rica, Colombia (Vaupés), Guyana, Peru, Brazil, Bolivia, Paraguay. apta (Walker, 1861: 321; Sapromyza). Type locality: Mexico. HT ? F (BMNH). Ref.—Giglio-Tos 1895: 49 (syn. of Sapromyza latelimbata). latelimbata (Macquart, 1855: 140, pl. 6, fig. 18 (wing); Sapromyza). Type locality: Brazil, Pernambuco. ST M (OXUM). Refs.: Giglio-Tos, 1895: 49 (redesc.); Malloch, 1926: 10; Curran, 1934b: 316, fig. 3 (wing), 445 (descr.). Distrib.—Mexico, Trinidad, Panama to southern Brazil and Paraguay. spinitibia Malloch, 1926: 10. Type locality: Costa Rica, San Mateo, Higuito. HT F (USNM). Distrib.—Costa Rica, Colombia (Meta), Guyana. xanthiceps (Williston, 1897: 9; Sapromyza). Type locality: Brazil, "Piedra Blanco". ST (AMNH). Distrib.— Colombia (Magdalena, Meta), Ecuador, Peru, Brazil. undescribed sp. 1—Locality: Colombia (Putumayo). Genus Camptoprosopella Hendel Camptoprosopella Hendel, 1907: 223. Type species, C. melanoptera Hendel (subs. des. Hendel, 1908: 53) = Pachycerina dolorosa (Williston). Refs.—Hendel, 1908: 53; 1936: 81 (key); Malloch, 1926: 6 (key), 1933: 356; Curran, 1928: 82, 1934b: 316, fig. 7 (head), fig. 20 (wing); Shewell, 1939 (rev.). diversa Curran, 1926: 13. Type locality: Puerto Rico, Coamo Springs. HT M (AMNH). Distrib. - United States (Florida), Jamaica, Haiti, Dominican Republic, Puerto Rico, Colombia (Magdalena, Bolívar). Ref.—Shewell, 1939: 138, figs. 25–26 (male terminalia). flavipalpis Malloch, 1926: 7. Type locality: Costa Rica, San Mateo. HT F (USNM). Distrib.—Costa Rica, Colombia (Cundinamarca). Ref.—Shewell, 1939: 139. hera Shewell, 1939: 140, figs. 27–28 (male terminalia). Type locality: Guatemala, Ayutla. HT M (USNM). Distrib.—Guatemala, Colombia (Bolivar). xanthoptera Hendel, 1907: 224. Type locality: Peru, Callanga. HT F (not localized). Distrib.—Mexico, Nicaragua, Costa Rica, Colombia, Ecuador, Venezuela, Peru, Bolivia. Ref.—Shewell, 1939: 138. Genus Chaetocoelia Giglio-Tos Chaetocoelia Giglio-Tos, 1893: 11. Type species, C. palans Giglio-Tos (mon.). Refs.—Hendel, 1907: 229 (key), 1908: 52; Melander, 1913: 77; Curran, 1942: 76 (key to species, except C. tripunctata Malloch). undescribed sp. 1—Locality: Colombia (Tolima). CATALOGUE OF LAUXANIIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 625 Genus Deceia Malloch Deceia Malloch, 1923: 49. Type species, Sapromyza crevecoeuri Coquillett (orig. des.). undescribed sp. 1—Locality: Colombia (Vaupés; Risaralda). Genus Deutominettia Hendel Deutominettia Hendel, 1925: 125. Type species, D. pulchrifrons Hendel (orig. des.). Ref.—Malloch, 1928: 2 (key to species). undescribed sp. 1—Locality: Colombia (Tolima; Risaralda; Vaupés; Norte de Santander). Genus Dryosapromyza Hendel Dryosapromyza Hendel, 1933: 68. Type species, Sapromyza gigas Schiner (orig. des.). undescribed sp. 1—Locality: Colombia (Magdalena). Genus Floriminettia Broadhead Floriminettia Broadhead, 1989: 188 (key), 189 (diag.). Type species, F. coronata Broadhead (orig. des.). coronata Broadhead, 1989: 189, fig. 13E (aedeagus, hypandrium), F (surstyli). Type locality: Panama, Barro Colorado I. HT M (BMNH). Distrib.—Panama, Colombia (Magdalena). Genus Gibbolauxania Papp & Silva Gibbolauxania Papp & Silva, 1995: 191. Type species, G. elegans Papp & Silva (orig. des.). undescribed sp. 1—Locality: Colombia (Amazonas; Putumayo). Genus Griphoneura Schiner Griphoneura Schiner, 1868: 281. Type species, Lauxania imbuta Wiedemann (orig. des.). Refs.—Hendel, 1908: 51; 1925: 134 (key to Neotropical species); Malloch, 1925 (rev., key); 1928: 18; Curran, 1934b: 317, fig. 13 (wing), 318, fig. 24 (head); 1938: 4 (key to North American species). ferruginea Schiner, 1868: 281. Type locality: "South America" (Venezuela). HT M (NHMW). Distrib.—Costa Rica, Colombia. Genus Homoeominettia Broadhead Homoeominettia Broadhead, 1989: 189 (key), 190 (diag.). Type species, Minettia assimilis Malloch (orig. des.). approximata (Malloch, 1928: 3; Deutominettia). Type locality: Panama, Río Trinidad. HT M (USNM). Distrib.— Panama, Colombia (Magdalena), Venezuela, Trinidad. assimilis (Malloch, 1926: 17; Minettia). Type locality: Panama, Canal Zone, Gatún Lake, Cano Saddle. HT M (USNM). Distrib.—Panama, Colombia (Bolívar, Magdalena), Trinidad, Venezuela, Brazil. 626 · Zootaxa 4122 (1) © 2016 Magnolia Press SILVA geniseta (Malloch, 1926: 16; Minettia). Type locality: Costa Rica, San Mateo, Higuito. HT F (USNM). Distrib.— United States (Texas), Mexico, Guatemala, Costa Rica, Panama, Colombia (Magdalena). woldae Broadhead, 1989: 190, fig. 5A (apex of male abdomen). Type locality: Panama, Fortuna. HT M (BMNH). Distrib.—Panama, Colombia (Magdalena). Genus Hypagoga Hendel Hypagoga Hendel, 1907: 233. Type species, Heteromyza apicalis Schiner (orig. des.). Refs.—Hendel, 1908: 57; Hennig, 1968: 3. undescribed sp. 1—Locality: Colombia (Risaralda; Cauca). Genus Marmarodeceia Shewell Marmarodeceia Shewell, 1986: 538. Type species, Pseudogriphoneura (Neodeceia) marmorata Malloch (orig. des.). marmorata (Malloch, 1926: 22; Pseudogriphoneura (Neodeceia)). Type locality: Costa Rica, San Mateo, Higuito. HT F (USNM). Distrib.—Costa Rica, Panama, West Indies, Colombia (Cauca), Venezuela, Ecuador, Peru, Brazil, Bolivia, Paraguay. Genus Minettia Robineau-Desvoidy Minettia Robineau-Desvoidy, 1830: 646. Type species, Minettia nemorosa Robineau-Desvoidy (subs. des. Westwood, 1840: 150) = fasciata (Fallén, 1826). Refs.—Curran, 1928: 84 (key to species from Puerto Rico); 1931: 20 (revised key); Malloch, 1928: 10 (key); 1933: 359; Hendel, 1932: 103. undetermined sp. 1—Locality: Colombia (Magdalena) Genus Neogriphoneura Malloch Neogriphoneura Malloch, in Malloch & McAtee, 1924: 11. Type species, Sapromyza sordida Wiedemann (orig. des.). Refs.— Curran, 1934b: 316, fig. 6 (wing), 320, fig. 35 (head); 1942: 75 (key); Mello & Silva, 2008a (rev., key). Rhabdolauxania Hendel, 1925: 128. Type species, R. schnusei Hendel (orig. des.). Refs.—Malloch, 1928: 8; Mello & Silva, 2008a: 26 (syn.). striga Curran, 1942: 75. Type locality: Panama, Canal Zone, Barro Colorado I. HT M (AMNH). Distrib.—Costa Rica, Panama, Colombia (Bolivar), Peru, Brazil, Bolivia. Ref.—Mello & Silva, 2007: 398–399 (labellum, figs. 1– 4). tertia Curran, 1942: 76. Type locality: Panama, Patilla Point. HT M (AMNH). Distrib.—Panama, Colombia (Bolivar; Magdalena), Brazil, Paraguay. Ref.—Mello & Silva, 2007: 400 (labellum, figs. 5–8). timida Curran, 1942: 75. Type locality: Panama, Patilla Point. HT M (AMNH). Distrib.—El Salvador, Panama, Colombia (Meta; Chocó; Magdalena). undescribed sp. 1—Locality: Colombia (Magdalena) CATALOGUE OF LAUXANIIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 627 Genus Neominettia Hendel Neominettia Hendel, 1925: 140. Type species, Musca contigua Fabricius (orig. des.). Comm.: This is not Neominettia Malloch; see Scutominettia. contigua (Fabricius, 1794: 347; Musca). Type locality: "Brazil". Type ? (probable ZMUC). Distrib.—Mexico to Brazil and Bolivia. Ref.—Curran, 1934b: 318, fig. 27 (wing). undescribed sp. 1—Locality: Colombia (Cauca) Genus Oncodometopus Shewell Oncodometopus Shewell, 1986: 539. Type species, Sapromyza umbrosa Loew (orig. des.). undescribed sp. 1—Locality: Colombia (Meta). undescribed sp. 2—Locality: Colombia (Meta). undescribed sp. 3—Locality: Colombia (Meta; Cundinamarca). undescribed sp. 4—Locality: Colombia (Cauca; Risaralda; Vaupés). Genus Pachyopella Shewell Pachyopella Shewell, 1986: 540. Type species, Pachycerina ornata Melander (orig. des.). flavida (Wiedemann, 1824: 57; Lauxania). Type locality: South America. HT M (ZMUC). Distrib.—United States (Florida) to Argentina (Missiones), Colombia (Magdalena). Refs.—Malloch, 1933: 356; Shewell, 1986: 541 (n. comb.). undescribed sp. 1—Locality: Colombia (Putumayo) Genus Paraphysoclypeus Papp & Silva Paraphysoclypeus Papp & Silva, 1995: 197. Type species, P. nigropleura Papp & Silva (orig. des.). undescribed sp. 1—Locality: Colombia (Tolima) Genus Physegenua Macquart Physegenua Macquart, 1848: 220 (1848: 60). Type species, P. vittata Macquart (orig. des.). Refs.—Hendel, 1908: 15; 1925: 115 (incomplete key to Neotropical species); Melander, 1913: 77; Curran, 1934b: 317, fig. 16 (head); 1942: 67 (incomplete key to Neotropical species). Physogenia, error. Hendel 1908: 10, 15; 1910: 120; 1925: 105,115; 1932: 109; 1933: 65; Malloch & McAtee, 1924: 8, 10. Physogenna, error. Johnson, 1919: 443. Physogenua, error. Shewell, 1986: 541. banksi Curran, 1942: 67. Type locality: Panama, Canal Zone, Frijoles. HT M (AMNH). Distrib.—Panama, Colombia (Tolima). 628 · Zootaxa 4122 (1) © 2016 Magnolia Press SILVA vittifrons Hendel, 1925: 116. Type locality: Bolivia, Mapiri, San Carlos, 800 m. HT F (SMTD). Distrib.— Colombia (Bolívar), Ecuador, Peru, Bolivia, Paraguay. Genus Physoclypeus Hendel Physoclypeus Hendel, 1907: 226. Type species, Chlorops flavus Wiedemann (orig. des.). Refs.—Hendel, 1908: 19; 1925: 120 (key); Malloch, 1933: 355; Mello & Silva, 2008b (rev.). annulatus Hendel, 1925: 121. Type locality: Bolivia, Mapiri, 2000 m. LT M (NHMW). Distrib.—Colombia (Risaralda), Bolivia. Refs.—Mello & Silva, 2008b (lec. des.). coquilletti (Hendel, 1908: 30; Lauxania (as non. nov. for Lauxania lutea Coquillett)). Type locality: United States, Florida, Lake Worth and Biscayne Bay. ST ? (USNM). Distrib.—United States, Puerto Rico, Cuba, Colombia (Magdalena, Nariño, Huila, Boyacá). Ref.—Curran, 1928: 83; Mello & Silva, 2008b: 291–293 (redesc.). flavus (Wiedemann, 1830: 595; Chlorops). Type locality: Uruguay, Montevideo. HT F (NHMW). Distrib.— Colombia (Boyacá), Brazil, Bolivia, Uruguay, Argentina. Refs.—Hendel, 1908: 20; 1925: 120; 1936: 82; Malloch, 1933: 356. lineatus (Williston, 1896: 385; Sapromyza). Type locality: St. Vincent (West Indies). LT M (BMNH). Distrib.—St. Vincent, Colombia (Tolima, Boyacá, Huila, Risaralda, Cundinamarca, Putumayo, Vaupés). Refs.—Mello & Silva, 2007: 400–401 (labellum, figs. 9–11); Mello & Silva, 2008b: 299–300 (redesc., lec. des.). risaraldensis Mello & Silva, 2008b: 302, figs. 8A‒J. Type locality: Colombia, Risaralda, SFF Otún Quimbaya, Robledal. HT M (IAHB). Distrib.—Colombia (Risaralda, Magdalena, Nariño, Huila, Boyacá), Ecuador. vitattus Mello & Silva, 2008b: 309, figs. 11A-I. Type locality: Brazil, Santa Catarina, Seara. HT M (MZUSP). Distrib.—Venezuela, Colombia (Risaralda), Brazil. undescribed sp. 1—Locality: Colombia (Risaralda; Huila; Norte de Santander). Genus Poecilominettia Hendel Poecilominettia Hendel, 1932: 103. Type species, Sapromyza picticornis Coquillett (orig. des.). Refs.—Hendel, 1932: (key to species); Broadhead, 1989 (key to Panamanian species). undetermined spp.—Locality: Colombia (Tolima; Meta; Bolivar; Cauca; Risaralda; Huila; Putumayo; Vaupés; Magdalena; Chocó; Norte de Santander) Genus Pseudogriphoneura Hendel Pseudogriphoneura Hendel, 1907: 226. Type species, P. cinerella Hendel (orig. des.). Refs.—Hendel, 1908: 50; 1925: 136 (partial key); Malloch, 1926: 19 (key); Curran, 1934a: 445 (key); 1934b: 316, fig. 5 (wing), 320, fig. 38 (head); 1942: 68 (partial key). pallipes Malloch, 1926: 20. Type locality: Costa Rica, San Mateo, Higuito. HT F (USNM). Distrib.—Costa Rica, Colombia (Magdalena). undetermined spp.—Locality: Colombia (Boyacá; Cundinamarca; Magdalena; Vaupés) CATALOGUE OF LAUXANIIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 629 Genus Sapromyza Fallén Sapromyza Fallén, 1810: 18. Type species, “Tephritis flava” Fabricius (Musca flava Linnaeus, 1758, misident.) (mon.; misident.) = obsoleta Fallén, 1820. Refs.—Blanchard, 1852: 444; Coquillett, 1898: 277; Hendel, 1908: 32; Becker, 1919: 185; Malloch, 1933: 369 (key). lebasii Macquart, 1844: 191. Type locality: "Colombia". HT M ? (MNHN). Distrib.—Colombia. puella Williston, 1896: 381. Type locality: St. Vincent (West Indies). ST M F (BMNH), M (AMNH). Distrib.—St. Vincent, Colombia (Tolima, Amazonas, Cauca, Putumayo, Vaupés, Magdalena). triseriata Coquillett, 1904: 95. Type locality: Nicaragua, Chinandega. HT ? (USNM). Distrib.—Nicaragua, Costa Rica, Colombia. Ref.—Malloch, 1926: 17. undescribed sp. 1—Locality: Colombia (Boyacá; Cundinamarca) Genus Sciosapromyza Hendel Sciosapromyza Hendel, 1933: 70. Type species, Dryomyza advena Wiedemann (mon.). undescribed sp. 1—Locality: Colombia (Magdalena) Genus Scutominettia Hendel Scutominettia Hendel, 1932: 109. Type species, S. mallochi Hendel (mon.). Neominettia Malloch, 1926: 9 (preocc. Hendel, 1925, with same nominal type species). Type species, Musca contigua Fabricius (orig. des., misident.) = mallochi Hendel, 1932. Refs.—Malloch, 1928: 1 (recognyzed Neominettia Hendel as senior primary homonym); Hendel, 1932: 109. mallochi Hendel, 1932: 109 (as nom. nov. for contigua, sensu Malloch, 1926, nec Fabricius). Type localities: Mexico, Vulcano, Colima and Costa Rica, San Mateo, Higuito. ST M, F (USNM). Distrib.—Mexico, Costa Rica, Colombia (Boyacá, Meta), Paraguay. undescribed sp. 1—Locality: Colombia (Boyacá; Risaralda; Norte de Santander) Genus Siphonophysa Hendel Siphonophysa Hendel, 1907: 230. Type species, S. pectinata Hendel (mon.). Ref.—Hendel, 1908: 29 (as subg. of Lauxania). undescribed sp. 1—Locality: Colombia (Boyacá; Vaupés). Genus Trivialia Malloch Deceia, subg. Trivialia Malloch, 1923: 53. Type species, Deceia fuscocapitata Malloch (orig. des.). Ref.—Shewell, 1965: 705 (new status). undescribed sp. 1—Locality: Colombia (Putumayo; Cauca; Vaupés; Magdalena). undescribed sp. 2—Locality: Colombia (Vaupés) 630 · Zootaxa 4122 (1) © 2016 Magnolia Press SILVA Genus Xenochaetina Malloch Xenochaetina Malloch, 1923: 49. Type species, Lauxania muscaria Loew (orig. des.) = flavipennis (Fabricius). Refs.— Hendel, 1932: 108 (key); Curran, 1934b: 320, fig. 30 (wing), fig. 34 (head). undetermined spp.—Locality: Colombia (Meta; Vaupés; Nariño). TABLE 1. Lauxaniidae species recorded from Colombia. (Taxa marked with * represent a new record for the country.) Subfamily Genus Species Author Eurychoromyiinae Euryhendelimyia schlingeri Gaimari & Silva, 2010a Exalla macalpinei Gaimari, 2011 shewelli Gaimari, 2011 Lauxaniinae Agriphoneura * undescribed sp. 1 Allogriphoneura * porcaria * (Fabricius, 1805) Allominettia corollae (Fabricius, 1805) spinitibia * Malloch, 1926 xanthiceps * (Williston, 1897) undescribed sp. 1 Camptoprosopella * diversa * Curran, 1926 flavipalpis * Malloch, 1926 hera * Shewell, 1939 xanthoptera * Hendel, 1907 Chaetocoelia * undescribed sp. 1 Deceia * undescribed sp. 1 Deutominettia * undescribed sp. 1 Dryosapromyza * undescribed sp. 1 Floriminettia * coronata * Gibbolauxania * undescribed sp. 1 Griphoneura * ferruginea Schiner, 1868 Homoeominettia * approximata * (Malloch, 1928) assimilis * (Malloch, 1926) geniseta * (Malloch, 1926) woldae * Broadhead, 1989 Broadhead, 1989 Hypagoga * undescribed sp. 1 Marmarodeceia marmorata Minettia * undetermined sp. 1 Neogriphoneura striga Curran, 1942 tertia * Curran, 1942 timida Curran, 1942 (Malloch, 1926) undescribed sp. 1 Neominettia contigua (Fabricius, 1794) undescribed sp. 1 Oncodometopus * undescribed sp. 1 undescribed sp. 2 ...Continued on next page CATALOGUE OF LAUXANIIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 631 TABLE 1. (Continued) Subfamily Genus Species Author undescribed sp. 3 undescribed sp. 4 Pachyopella * flavida * (Wiedemann, 1824) undescribed sp. 1 Paraphysoclypeus * undescribed sp. 1 Physegenua * banksi * Curran, 1942 vittifrons * Hendel, 1925 annulatus * Hendel, 1925 coquilletti * (Hendel, 1908) flavus (Wiedemann, 1830) lineatus * (Williston, 1896) risaraldensis Mello & Silva, 2008b vitattus Mello & Silva, 2008b Physoclypeus undescribed sp. 1 Poecilominettia * undetermined spp. Pseudogriphoneura * pallipes * Malloch, 1926 undetermined spp. Sapromyza lebasii Macquart, 1844 puella * Williston, 1896 triseriata Coquillett, 1904a undescribed sp. 1 Sciosapromyza * undescribed sp. 1 Scutominettia * mallochi * Hendel, 1932 undescribed sp. 1 Siphonophysa * undescribed sp. 1 Trivialia * undescribed sp. 1 undescribed sp. 2 Xenochaetina undetermined spp. References Becker, T. (1919) Diptères. Brachycères. Mission Arc Méridien Amérique du Sud, Paris, 10, 163–215. Blanchard, E. (1852) Orden IX. Dipteros. 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Transactions of the Entomological Society of London, 1896, 253–446. http://dx.doi.org/10.1111/j.1365-2311.1896.tb00965.x Williston, S.W. (1897) Diptera Braziliana, part 4. Kansas University Quarterly, Series A, 6, 1–12. 634 · Zootaxa 4122 (1) © 2016 Magnolia Press SILVA Zootaxa 4122 (1): 635–641 http://www.mapress.com/j/zt/ Copyright © 2016 Magnolia Press Article ISSN 1175-5326 (print edition) ZOOTAXA ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.4122.1.52 http://zoobank.org/urn:lsid:zoobank.org:pub:DA2C180D-B6A6-4492-9A7A-0E69D2AFB7BB FAMILY ROPALOMERIDAE ROSALY ALE-ROCHA Coordenação de Biodiversidade, Instituto Nacional de Pesquisas da Amazônia, Av.André Araújo, 2936, Petrópolis, CEP 69067-375, Manaus, Amazonas, Brazil. Fellowship PQ/CNPq. E-mail: alerocha@inpa.gov.br Abstract Ropalomeridae is a small family with most species distributed in the Neotropical Region, from Mexico to Argentina, and only one Nearctic species. In Colombia, eight species distributed in four genera have been found. This catalogue, based on the study of specimens and available literature records, summarizes and updates the information on the Colombian fauna. Key words: catalogue, Colombia, distribution, Neotropical Region, Ropalomeridae Introduction Flies in the family Ropalomeridae are distributed mainly in the tropical and subtropical regions of the Americas. The family is characterized by the following combination of characters: adults robust, medium to large (6-12 mm), brown to dark brown; vertex deep, concave, eyes protruding and face with median carina or central tubercle; palpi flattened and widened; posterior thoracic spiracle with one or more setae; legs robust, especially hind femur, significantly widened, with strong ventral setae, hind tibia usually laterally compressed, curved, ornamented with long bristles or hairs dorsally; wings hyaline, lightly infuscate or strongly spotted. The morphology of the immatures is poorly understood. Lopes (1932) provided descriptions of the egg and larva of Ropalomera stictica Wiedemann. The biology of Ropalomeridae is poorly known. Adults feed on liquids that exude from plants and are collected using traps baited with rotting fruit, hydrolyzed protein, and 10% cane molasses. Ropalomeridae are included in the suborder Brachycera, infraorder Muscomorpha, superfamily Sciomyzoidea. Sepsidae are considered the sister-group of this family, and both share the middle portion of face strongly sclerotized and prominent, one or more strong subvibrissal setae, posterior thoracic spiracle with one or more developed bristles, vein R1 bare, vein A2 absent and two spermathecae (McAlpine 1989). On the other hand, nothing is known about the relationships within Ropalomeridae. The family was reviewed by Lindner (1930a), Prado (1963, 1966) and cataloged by Steyskal (1967), who also published a chapter on the fauna of the Nearctic region (Steyskal 1987). Mexican species were revised by RamírezGarcía & Hernández-Ortiz (1994), Willistoniella Mik, 1895 and Apophorynchus Williston, 1895 by Marques & Ale-Rocha (2004 and 2005, respectively), and Kroeberia Lindner, 1930 by Marques-Costa & Ale-Rocha (2005). Currently, there are 31 known species in 9 genera in the Neotropics (Ibáñez-Bernal & Hernández-Ortiz 2012: Kirst & Ale-Rocha 2012). They are distributed from the southern United States to northern Argentina, with a single species recorded from the Nearctic region, Rhytidops floridensis (Aldrich 1932). Fossils have not been found. Eight species of Ropalomeridae have been recorded from Colombia, in addition to two unrecognized species. The present catalogue provides a list of species from Colombia based on available literature and observed material. A table with all species, rank and author is presented at the end of this paper. The information included in the Catalogue comprises only the species that occur in Colombia, including synonymies. Accepted by S. Nihei: 10 Apr. 2015; published: 14 Jun. 2016 635 Acronyms used for the depositories CEUA—Colección Entomológica, Universidad de Antioquia, Medellín, Colombia IBUNAM—Instituto de Biología de la Universidad Nacional Autónoma de México, México INPA—Instituto Nacional de Pesquisas da Amazônia, Manaus, Brazil IOC—Instituto Oswaldo Cruz, Rio de Janeiro, Brazil NHM—Natural History Museum, London, England NHMW—Naturhistorisches Museum Wien, Austria ZMUC—Zoological Museum University of Copenhagen, Copenhagen, Denmark List of abbreviations cat.—catalogue cit.—citation diagn.—diagnosis distr.—geographic distribution refs.—references mon.—by monotypy orig. des.—by original designation pp.—pages preocc.—preoccupied redesc.—redescription rev.—revision HT—Holotype F (f#)—Female M (m#)—Male Catalogue of Ropalomeridae of Colombia Family Ropalomeridae Schiner, 1868 Genus Apophorhynchus Williston, 1895 Apophorhynchus Williston, 1895a: 184. Type species, Apophorhynchus flavidus Williston, 1895a, orig. des. Refs.: Williston, 1896: 280–281 (key); Lindner, 1930a: 125–126 (diagn., key); Malloch, 1941: 50 (diagn.); Steyskal, 1967: 1 (cat.); Prado & Papavero, 2002: 1 (cat.). amazonensis Prado, 1966: 248, 251–252. Type locality: Brazil, Amazonas, São Gabriel da Cachoeira, Amazonas. HT F (IOC 13210). Distr.: Brazil, Colombia, Peru. Refs.: Prado, 1967: 73–75 (male descr., distr.); Steyskal, 1967: 1 (cat.); Prado & Papavero, 2002: 1 (cat.); 2009: 2 (cat.); Marques-Costa & Ale-Rocha, 2005: 515 (rev.). Material examined. COLOMBIA: Macaqual, 1 male, 8.iii.2011, T. VSR fish 9 (CEUA). Genus Kroeberia Lindner, 1930 Kroeberia Lindner, 1930a: 125, 127. Type species, Kroeberia fuliginosa Lindner, 1930a: 127, orig. des. Refs.: Steyskal, 1967: 2 (cat.); Marques & Ale-Rocha, 2004: 315 (rev.); Prado & Papavero, 2009: 2 (cat.). fuliginosa Lindner, 1930a: 127. Type locality: Brazil, Rio Grande do Sul, Santa Cruz. HT M (NHMW). Distr.: Brazil, Colombia (Caldas, Atlántico), Mexico, Panama,Venezuela. Refs.: Aldrich, 1932: 9 (redesc.); Fischer, 1932: 450 (diag.); Prado, 1966: 253 (redesc., distr.); Steyskal, 1967: 1 (cat.); Baez, 1985: 79 (distr.); Ramírez García & 636 · Zootaxa 4122 (1) © 2016 Magnolia Press ALE-ROCHA Hernández Ortiz, 1994: 76 (redesc., distr., key); Prado & Papavero, 2002: 1 (cat.); 2009: 2 (cat.); Marques & AleRocha, 2004: 316 (rev.). Material examined. COLOMBIA: Atlantico: La Villa del Rosario, 1male, 16.iii.2013 (CEUA). Genus Ropalomera Wiedeman, 1824 Ropalomera Wiedemann, 1824: 17. Type species, Dictya clavipes Fabricius, 1805 (sub. des., Macquart, 1843: 359 [202]). Refs.: Wiedemann, 1828: 570 (redesc.); Steyskal, 1967: 2 (cat.); Prado & Papavero, 2002: 1 (cat.); 2009: 3 (cat.); Kirst & Ale-Rocha, 2012: 1 (rev.). Rhopalomera Agassiz, 1846: 4 (59), emended (injustified). femorata (Fabricius), 1805: 326 (Dictya). Type locality: Suriname [as “Sudamerica”]. HT M (ZMUC). Distr.: Colombia (Antioquia, Atlántico, Bolivar, Caldas, Caquetá, Cauca, Huila, Magdalena, Santa Marta, Sucre), Mexico, Guatemala, Nicaragua, Costa Rica, Venezuela, Brazil, Ecuador, Bolivia, Paraguay, Argentina). Refs.: Wiedemann, 1828: 571; Williston, 1895a: 184–185 (key) (Rhopalomera); Lindner, 1930a: 130 (redescr.); 1930b: 282 (Rhopalomera); Frey, 1959: 47 (key) (Rhopalomera); Prado, 1966: 215, 223, 225 (key, rev.) (Rhopalomera); 1967: 72; Steyskal, 1967: 2 (cat.); Baez, 1985: 78; Ibañes-Bernal et al., 1992: 91, 94–97 (key, redescr.); Ramírez-García & Hernández-Ortiz, 1994: 65–66, 83 (redescription, key); Prado, & Papavero, 2002: 1 (cat.); 2009: 3 (cat.); Kirst & Ale-Rocha, 2012: 8 (rev.). xanthops Williston, 1895b: 213 (Rhopalomera); Ramírez-García & Hernández-Ortiz, 1994: 65 (lectotype designation and synonymy of R. femorata). Material examined. COLOMBIA: Antioquia: Jericó, Parcelación Cauca, Viejo Litera Quebrada los Cruces, Bosque intervenido, 614m, 2 females; 1–5.vii.2009, M. Wolff (CEUA); Cerca a Quebrada de Las Cruces, Litera rocosa, 5°52ʹ21.59ʺN, 75°45ʹ 30.83ʺW, 614m, 2 females; 8.vii.2010 (CEUA 50171, 50172); 1 female, 5.vii.2009 (CEUA); Atlántico: Barranquilla, Uni-Atlantico, Campus Universitario, 11°01ʹ07.58ʺN, 74°52ʹ28.19ʺW, 48m, 7.vii.2010, C. Valverde & E. Perdomo, T. VSR (van someren trap) (CEUA); 6 males, 5.viii.2010 (CEUA); 3 males, 6.vii.2010 (CEUA); Bolívar: Cartagena, Corr. El Arenal, Finca E Peligro, 10°26ʹ56ʺN, 5°16ʹ20.53ʺW, 5males, 1 female, 19. viii.2010, C. Valverde (CEUA); Caldas: Norcasia, 05º33'09.8"N, 74º52'23.1'O, 1male, 10.ii.2010, DY. Gaurisas & L. Rios (CEUA); 1 male, 20.ii.2010, (CEUA); Magdalena: Santa Marta Unimagdalena, Parche bosque seco (dry forest), 11°13ʹ18.36ʺN, 74°11ʹ10.89ʺW, 20m, 2 males, 24.vi. 2010, C. Valverde y E. Perdomo, T. VSR (van someren trap): Fruta (CEUA); 6 females, 26.vi.2010 (CEUA); 2males, 26.vi.2010 (CEUA); Santa Marta: Colegio INEM, Granja Potrero, 11°13ʹ42.45ʺN, 74°10ʹ 13.17ʺW, 4 males, 27.viii.2010, C. Valverde & E. Perdomo, T. VSR Fruta (van someren trap with decomposition fruit), (CEUA); 4 males, 7.viii.2010 (CEUA); Sucre: San Onofre, Corregimiento Berrugas, Rva (Reserva) Sanguaré, 9°42ʹ21.6ʺN, 75°40ʹ45.6ʺW, 8m, 2 males, 7females, 29.x.2009, T. VRS Fruta (van someren trap with decomposition fruit), (CEUA); 2 males, 3 females, 30.x.2009; 3 males, 31.x.2009 (CEUA); Silvopastoril, 9°42ʹ21.6ʺN, 75°40ʹ45.6ʺW, 1 male (CEUA); Regeneracion, 1male, 2 females, 12.ii.2010, T. VRS Fruta (van someren trap with decomposition fruit), (CEUA); Regeneracion, 3 females, 14.ii.2010, T. VRS: Fruta (CEUA); Regeneracion, 2 males, 13.ii.2010, T. VRS Fruta (van someren trap with decomposition fruit), (CEUA); San Onofre, Rva (Reserva) Sanguare, T. VSR Pez (van someren trap with decomposition fish), Potrero, 2 males, 4 females 25–30.v.2009 (CEUA). latiforceps Ramírez-García & Hernández-Ortiz, 1994: 69. Type locality: Mexico, Veracruz, Estación de Biología “Los Tuxtlas”. HT M (IBUNAM). Distr.: Mexico, Panama, Costa Rica, Colombia (Antioquia). Refs.: IbáñezBernal & Hernández-Ortiz, 2012: 563 (distr.). tibialis Walker, 1852: 375. Type locality: “Brazil”. HT M (NHM). Distr. Brazil, Bolivia, Colombia (Huila, Caquetá). Refs.: Walker, 1857: 224; Williston, 1895a: 184 (key) (Rhopalomera); Prado, 1967: 72 (cit.); Steyskal, 1967: 3 (cat.); Prado & Papavero, 2002: 1 (cat.); 2009: 4 (cat.); Kirst & Ale-Rocha, 2012: 21 (rev.). walkeri Prado, 1966: 215, 243, 245 (key, description) (Rhopalomera). Materialexamined. COLOMBIA: Huila: PNN Cueva de Los Guacharos, Caquetá, PNN Chiribiqueterio, Sararemano, Bosque Cerro, 2males, 2females, 2000, Malaise (CEUA). CATALOGUE OF ROPALOMERIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 637 Genus Willistoniella Mik, 1895 Rhopalomyia Williston, 1895: 184 (preocc. name by Rübsaamen, 1892). Type species, Ropalomera pleuropunctata Wiedemann, 1824, by mon. Willistoniella Mik, 1895: 136 (nom. nov. for Rhopalomyia Williston). Type species, Ropalomera pleuropunctata Wiedemann, 1824, by automatic designation. Refs.: Williston, 1896: 280–281 (key); Lindner, 1930a: 125–129 (diagn., key, redescr.); Steyskal, 1987: 85 (key); Prado & Papavero, 2002: 2 (cat.); 2009: 4 (cat.); Marques & Ale-Rocha, 2005: 210 (rev.). pleuropunctata (Wiedemann), 1824: 18 (Ropalomera). Type locality: “Amer. Mer.”. HT F (NHMW). Distr.: Mexico, Belize, Guatemala, Honduras, Curaçao, Costa Rica, Panama, Trinidad, Colombia (Antioquia, Chocó, Sucre), Ecuador, Peru, Venezuela, Surinam, Guyana, Brazil, Bolivia, Paraguay, Argentina. Refs.: Wiedemann, 1828: 572 (redescr.); Giglio-Tos, 1894: 43 (Rhopalomera); Williston, 1895: 184–186 (Rhopalomyia); Mik, 1895: 136 (as type-species of the new name); Aldrich, 1905: 600 (Rhopalomera); 1932: 11 (distr.); Frey, 1921: 101–103 (redesc.) (Rhopalomera); Lindner, 1930a: 128–129 (redesc., distr.); 1930b:282 (cit.); Hendel, 1923: 203 (cit., distr.); 1933: 214 (cit., distr.); Fischer, 1932: 449 (description of the puparium); Steyskal, 1957: 71 (male terminalia); 1987: 943 (male terminalia, error = W. ulyssesi Marques & Ale-Rocha); Hennig, 1958: 581–590; Frey, 1959: 47 (cit., distr.); Prado, 1963: 466–469 (redesc., error = W. ulyssesi Marques & Ale-Rocha); 1966: 247 (cit., distr.); 1967: 73 (cit.); Baez, 1985: 80 (cit., distr.); Ramírez-García & Hernández-Ortiz, 1994: 72–75, 83 (diagn., distr., key, error = W. ulyssesi Marques & Ale-Rocha); Prado & Papavero, 2002: 2 (cat.); 2009: 5 (cat.); Marques & Ale-Rocha, 2005: 213 (rev.); Ibáñez-Bernal & Hernández-Ortiz, 2012: 565 (distr.). vittifrons Rondani, 1848: 81 (Rhopalomera). substituta Walker, 1858: 225 (Ropalomera). Material examined. COLOMBIA: Antioquia: Jericó, Parcelación Cauca, Viejo Litera Quebrada las Cruces, Litera rocosa, 5° 52ʹ 21.59ʺN, 75° 45ʹ 30.83ʺW, 614m, 4 males, 8.vii.2010, M. Wolff & L. Rios, T.VSR Pez (van someren trap with decomposition fish), (CEUA); 3 males, 5.vii.2009, Wolff, T. VSR Fruta (van someren trap with decomposition fruit), (CEUA); Bosque Intervenido, 614m, 1 male, 1–5.vii.2009, VSR: Fruta (CEUA); Rio Claro, Camino La Mulata, 1 male, 9.xi.2007, VSR: Banano (CEUA); Chocó: Niqui, Corr. Jobi a 50m playa, 3 females, 6.viii.2006, P. Duque & M. Wolff (CEUA); Sucre: San Onofre, Rva Sanguare, Potrero, 2 males, 25–30.v.2009, T. VSR Pez (van someren trap with decomposition fish), (CEUA); San Onofre, 9º42'21.6"N, 75º40'45.6"W, 2 males, 13.ii.2010 (CEUA). spatulata Marques & Ale-Rocha, 2005: 210. Type locality: Brazil, Amazonas, Beruri. HT M (INPA). Distr.: Honduras, Panamá, Venezuela, Colômbia (Boyacá, Meta), Equador, Brazil. Refs.: Henriques et al., 2007: 60 (distr.); Prado & Papavero, 2009: 4 (cat.). ulyssesi Marques & Ale-Rocha, 2005: 221. Type locality: Brazil, Amazonas, Manaus, R[eserva Adolpho] Ducke. HT M (INPA). Distr.: Mexico, Belize, Guatemala, Honduras, Costa Rica, Panama, Trinidad, Colombia (Antioquia, Cauca, Chocó, Meta, Sucre),Venezuela, Ecuador, Peru, Bolivia, Paraguay, Brazil. Refs.: Henriques et al., 2007: 60 (distr.); Prado & Papavero, 2009: 4 (cat.). Material examined. COLOMBIA: Antioquia: San Luis, Rio Claro, Rva (Reserva) Natural Cañon del Rio Claro, Bosque Ripario, 05°54ʹ2.98ʺN, 74 51 25.04ʺO, 350m, 6 males, 1 female, 1.xi.2009, I. Ceballos, T.VSR Pez (van someren trap with decomposition fish), (CEUA); La Pintada, Hda (Hacienda) Montenegro-Comfenalco, Bosque, 05°43ʹ 24ʺN, 75°37ʹ15ʺW, 850 m, 2 males, 15.iii.2007 (CEUA); Hotel-Comfenalco (La Pintada), 05°43ʹ24ʺN, 75°37ʹ15ʺW, 770m, 2 males, vi.2007, Al. L. Montoya (CEUA); Jericó, Parcelación Cauca Viejo, 2 males,1–5.vii.2005 (CEUA); Jericó, Parcelación Cauca Viejo cerca a quebrada Las Cruces, Litera rocosas, 05°52ʹ21.89ʺN, 75°45ʹ30.83ʺW, 614m, 2 females, 8.vii.2010, M. Wolff (CEUA); 2 females, vii.5.2009, T. VRS Pez (van someren trap with decomposition fish), (CEUA); Choco: Nuqui, Jobi Playa, 50m, 3 females, 1 male, 6.viii.2006, M. Wolff & P. Duque, T. VSR Pez (van someren trap with decomposition fish ), (CEUA); Jobi, Guacharito, Patio playa, 40m, 1 male, 1 female, 1–3.viii.2008, Wolff, T. VRS Fruta (van someren trap with decomposition fruit), (CEUA); Sucre: San Onofre, Berrugas, Rva (Reserva). Sanguaré, 9°42ʹ21.6ʺN, 75°40ʹ45.6ʺW, 8m, 1 female, 30.x.2009, T. VRS Pez (van someren trap with decomposition fish), (CEUA); 1 male, 25–30.v.2009, TVRS Fruta (van someren trap with decomposition fruit), (CEUA); Regeneración, 2 male, 1 female, 12–13.ii.2010 (CEUA). 638 · Zootaxa 4122 (1) © 2016 Magnolia Press ALE-ROCHA Unrecognized species registered to Colombia flaviceps Macquart, 1846: 226, pl. 18, fig. 6 (Ropalomera). Type locality: “Nouvelle Grenade” [=Colombia]. HT M (?). Refs.: Prado, 1966: 246 (may belong in Apophorhynchus); Prado & Papavero, 2009: 13 (cat.). varipes Walker, 1862: 375 (Ropalomera). Type locality: “Colombia”. HT (NHM?). Distr.: Colombia. Refs.: Williston, 1895a: 184 (key); Prado, 1966: 246 (treated it as unrecognizable from description); 1967: 71 (synonymy with R. femorata); Steyskal, 1967: 4 (could not locate the type at the Natural History Museum); Prado & Papavero, 2009: 13 (cat.). Acknowledgements I sincerely thank Marta Wolff (Universidad de Antioquia, Colombia) for loaning the material examined. We also thank Coordenação de Aperfeiçoamento de Pessoal de Nível Superior (CAPES) and Fundação de Amparo à Pesquisa do Estado do Amazonas (FAPEAM) for the support to this work through the project “Incremento do conhecimento da diversidade, formação e fixação de especialistas em Sistemática de Diptera (Insecta) na Amazônia” and Conselho Nacional de Desenvolvimento Científico e Tecnológico (CNPq) by the project “Sistemática morfológica de Hybotidae (Diptera, Empidoidea) com ênfase nas espécies neotropicais de Ocydromiinae e Hybotinae”. TABLE 1. Ropalomeridae species recorded from Colombia. Genus Species Author Apophorhynchus amazonensis Prado, 1966 Kroeberia fuliginosa Lindner, 1930 Ropalomera femorata (Fabricius, 1805) latiforceps Ramírez-García & Hernández-Ortiz 1994 tibialis Walker, 1852 pleuropunctata (Wiedemann, 1824) spatulata Marques & Ale-Rocha, 2005 ulyssesi Marques & Ale-Rocha 2005 flaviceps Macquart, 1846 varipes Walker, 1862 Willistoniella Unrecognized species References Agassiz, L. (1846) Nomenclatoris zoologici index universalis, continens nomina systematica classium, ordinum, familiarum et generum animalium omnium, tam viventium quam fossilium, secundum ordinem alphabeticum unicum disposita, adjectis homonymiis plantarum, nec non variis adnotationibus et emendationibus. Fasicle 12 (Indicem Universalem). Jent et Gassman, Soloduri, 393 pp. Aldrich, J.M. (1905) A Catalogue of North American Diptera. Smithsonian Institution Miscellaneous Collections, 46, 1–680. Aldrich, J.M. 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Hathaway, New Haven, 167 pp. CATALOGUE OF ROPALOMERIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 641 Zootaxa 4122 (1): 642–647 http://www.mapress.com/j/zt/ Copyright © 2016 Magnolia Press Article ISSN 1175-5326 (print edition) ZOOTAXA ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.4122.1.53 http://zoobank.org/urn:lsid:zoobank.org:pub:9CCC53AF-2CEF-4D76-BC10-5B17C6525F15 FAMILY SCIOMYZIDAE LUCIANE MARINONI1* & WILLIAM L. MURPHY2 1 Departamento de Zoologia, Setor de Ciências Biológicas, Universidade Federal do Paraná, Cx. P. 19020, CEP 80.531-980, Curitiba, Paraná, Brasil. E-mail: lmarinoni@msn.com. 2 Research Collaborator, Smithsonian Institution, 7835 Tufton Street, Fishers, Indiana 46038, U.S.A. E-mail: billmurphy8@sbcglobal.net *Corresponding author: lmarinoni@msn.com Abstract The Sciomyzidae are a family of acalyptrate flies of worldwide distribution, with 543 extant species and 14 described subspecies in 63 genera. Although 274 species in 37 genera are found in the Western Hemisphere, the sciomyzid fauna of Central and South America remains relatively unknown, comprising 103 species in 25 genera, with only seven species in five genera having been recorded from Colombia: Dictya bergi Valley, Perilimnia albifacies Becker, Pherbellia guttata (Coquillett), Sepedomerus bipuncticeps (Malloch), S. macropus (Walker), Sepedonea guianica (Steyskal), and S. isthmi (Steyskal). Key words: Acalyptratae, catalog, catalogue, Colombia, distribution, Neotropical, Neotropics, snail-killing flies, taxonomy Introduction The Sciomyzidae are a family of acalyptrate flies of worldwide distribution, with 543 extant species (Murphy 2014) and 14 described subspecies in 63 genera and 13 fossil species in five genera (Vala et al. 2012, 2013). Traditionally the family has been considered to comprise two subfamilies, Salticellinae and Sciomyzinae (Marinoni & Mathis 2000, Barker et al. 2004). More recently, DNA analyses by Tóthová et al. (2012) showed it to comprise three subfamilies: Phaeomyiinae, Salticellinae, and Sciomyzinae. Sciomyzidae is one of the best known families of Diptera. The species are considered to be “true malacophages” and are classified into 15 different behavioral groups (Knutson & Vala 2011). The biology of 241 species in 41 genera is known, with only three species not being associated with molluscs (Vala et al. 2013). Sciomyzids that prey on non-operculate aquatic snails hold promise to control species of gastropods that act as intermediate hosts of diseases such as fascioliasis and schistosomiasis (Berg & Knutson 1978, Barker et al. 2004). Although 274 species in 37 genera are found in the Western Hemisphere (Murphy 2014), the sciomyzid fauna of Central and South America remains relatively unknown, comprising 103 species in 25 genera, with only seven species in five genera having been recorded from Colombia (Table 1). This low number results from lack of information about specimens in collections and lack of collecting in Colombia. Many species known from adjacent countries (Panama, Venezuela, and Ecuador) likely occur in Colombia but have yet to be collected. Acronyms used for the depositories BMNH—Natural History Museum, London, UK CU—Cornell University, Ithaca, New York, USA MNHNP—Muséum national d’Histoire naturelle, Paris, France USNM—U.S. National Museum of Natural History, Washington, DC, USA 642 Accepted by S. Nihei: 26 May 2015; published: 14 Jun. 2016 List of abbreviations biol.—biology biol. control—biological control cat.—catalogue distr.—distribution imm. stages—immature stages kar.—karyotype key imm.—key to immature stages mon.—monotypy nom. nud.—nomen nudum orig. des.—original designation preocc.—preoccupied redesc.—redescription refs.—references syn.—synonymy F (f#)—Female M (m#)—Male The references (“Refs.”) for each taxon include only the most important publications, i.e. those providing revisions, identification keys, detailed descriptions, type notes, catalogues, etc. Catalogue of Sciomyzidae of Colombia “Sciomyzides” Fallén, 1820 Family “Sciomyzidae” Macquart, 1846 Subfamily Sciomyzinae Schiner, 1862 Tribe Sciomyzini Cresson, 1920 Genus Pherbellia Robineau-Desvoidy Pherbellia Robineau-Desvoidy 1830: 695. Type species, Pherbellia vernalis Robineau-Desvoidy 1830: 696 (mon.)—Musca punctata Fabricius 1794: 347 (preocc. Poda 1761: 116—Sciomyza schoenherri Fallén 1826: 13). Refs.: Knutson et al. 1976: 4 (cat.); Marinoni & Mathis 2000: 163-191 (cladistic analysis of the family), figs. 11, 76, 169, 220, 221 (antenna, male terminalia); Barker et al. 2004; Knutson & Vala 2011; Vala et al. 2013. Melina Robineau-Desvoidy 1830: 695 (preocc. Retzius 1788: 22). Type species Melina riparia Robineau-Desvoidy 1830: 695 (mon.)—Sciomyza dubia Fallén. Chetocera Robineau-Desvoidy 1830: 697 (Coquillett 1910: 567) Graphomyzina Macquart 1835: 558. Type species elegans Macquart 1835: 559 (orig. des.)—Sciomyza limbata Meigen 1830: 17. Notopherbellia Malloch 1933: 311. Type species chiloensis Malloch 1933: 311 (mon.). Sciomyza of authors, not Fallén 1820. guttata (Coquillett) 1901: 615 (Sciomyza) [Cresson 1920: 48 (as Melina, subgenus Graphomyzina) Steyskal 1965: 686 (Pherbellia), but not stated as new combination]. Type locality: U.S.A., Texas. F (USNM). Distr.: U.S.A. (Texas: Austin, Cameron Co.); Mexico (Chiapas, Distrito Federal Tamaulipas); Bahamas; Jamaica (St. Ann); Colombia (Valle del Cauca [Palmira]); Peru (Loreto: San Antonio). Refs.: Knutson et al., 1976: 4 (cat.); Vala et al., 2013: 85. CATALOGUE OF SCIOMYZIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 643 Tribe Tetanocerini Steyskal, 1965 Genus Dictya Meigen Dictya Meigen, 1803: 277. Type species, Musca umbrarum Linnaeus, 1758: 599 (by designation of Cresson, 1920: 70) (Traditional usage pending submission of proposal to I.C.Z.N. for fixation of type species). Refs.: Becker, 1919: 189; Knutson et al., 1976: 6 (cat.); Marinoni & Mathis, 2000: 163-191 (cladistic analysis of the family), figs. 20, 84, 174, 239241 (antenna, male terminalia); Barker et al., 2004; Knutson & Vala, 2011; Vala et al., 2013. Monochaetophora Hendel 1900: 355. Type species, Musca umbrarum Linnaeus, 1758: 599 (mon.). bergi Valley, 1977 in Valley & Berg, 1977: 3. Type locality: Costa Rica: San José Prov., San Antonio–Desamparados. M F (CU). Distr.: Costa Rica (Cartago, San José,); Colombia (Valle del Cauca). Refs.: Knutson et al., 1976: 6 (cat.); Vala et al., 2013: 97. Genus Perilimnia Becker Perilimnia Becker. 1919: 189. Type species, albifacies Becker. 1919: 189 (mon.). Refs.: Knutson et al., 1976: 8 (cat.); Marinoni & Mathis, 2000: 163-191 (cladistic analysis of the family), figs. 18, 82, 129, 172, 233-235 (antenna, male terminalia); Barker et al., 2004; Knutson & Vala, 2011; Vala et al., 2013. Notosciomyza Malloch, 1933: 313. Type species, dubia Malloch (orig. des.)—albifacies Becker, 1919: 189. albifacies Becker, 1919: 189. Type locality: Ecuador, Cuencas. M F (MNHNP). Distr.: Colombia (Cundinamarca, Nariño, Putumayo); Ecuador (Azuay, Cotopaxi, El Oro, Napo, Pichincha); Peru (Cusco, Lima); Bolivia (La Paz); Brazil (Rio Grande do Sul); Uruguay (Montevideo); Argentina (Tucumán, Jujuy, Corrientes, Cordoba, Mendoza, Buenos Aires, Rio Negro, Chubut, Santa Cruz, La Rioja); Chile (Valparaiso, Malleco, Atacama, Coquimbo, Aconcagua, Santiago, Maule, Concepción, Osorno, Llanquihu, Aysén, Magallanes). Refs.: Steyskal, 1953: 75; 1963: 123; Boyes et al., 1969: 338 (kar.); Kaczynski et al.,, 1969: figs. 1, 2, 6, 11–19, 29–34, 41–48 (male genitalia, biol., imm. stages, distr.); Knutson et al., 1976: 8 (cat.); Vala et al., 2013: 110. Genus Sepedomerus Steyskal Sepedomerus Steyskal, 1973: 144. Type species, Sepedon macropus Walker, 1849 (orig. des.). Refs.: Knutson et al., 1976: 9 (cat.); Marinoni & Mathis, 2000: 163-191 (cladistic analysis of the family), figs. 50, 113, 159, 202, 320-321 (antenna, male terminalia); Barker et al., 2004; Knutson & Vala, 2011; Vala et al., 2013. bipuncticeps (Malloch), 1933: 299, fig. 55a (Sepedon). Type locality: Argentina, Buenos Aires, La Plata City. F (BMNH). Distr.: Panama (Panama); Colombia (Valle del Cauca (El Morgan, Cali)); Brazil (Acre, Goiás, Minas Gerais, Paraná, Rio Grande do Sul); Paraguay (Villarrica, Guaira); Uruguay (Montevideo, Colón); Argentina (Distrito Federal, Chaco, Buenos Aires, La Plata, Catamarca). Refs.: Knutson et al., 1976: 9 (cat.); Vala et al., 2013; 113. macropus (Walker), 1849: 1078, fig. 556 (Sepedon). Type locality: Jamaica. M (BMNH). Distr.: U.S.A. (Texas); Cuba; Jamaica; Puerto Rico; Tobago; Trinidad; Mexico; Guatemala; Honduras; Nicaragua; Costa Rica; Panama; Colombia; Ecuador; Peru. Refs.: Malloch, 1914: 323 (syn.); Malloch, 1933: 300, fig. 55b (Sepedon); Steyskal, 1951: 276, pl. 5, fig. 24 (male hind femur and tibia); Chock et al., 1961 (biol. control); Neff & Berg, 1966 (biol., imm. stages, distr.); Boyes et al., 1969: 344 (kar.); Knutson et al., 1976: 10 (cat.); Vala et al., 2013: 113. Genus Sepedonea Steyskal Sepedonea Steyskal, 1973: 145. Type species, Sepedon lindneri Hendel, 1932: 99 (orig. des.); Refs.: Knutson et al., 1976: 10 (cat.); Marinoni & Mathis, 2000: 163-191 (cladistic analysis of the family), figs. 51, 114, 160, 203, 322-324 (antenna, male 644 · Zootaxa 4122 (1) © 2016 Magnolia Press MARINONI & MURPHY terminalia); Barker et al., 2004; Marinoni & Mathis, 2006 (cladistic analysis of the species in the genus); Knutson & Vala, 2011; Vala et al., 2013. guianica (Steyskal), 1951: 295, pl. 4, fig. 18 (Sepedon). Type locality: “Guyane” [French Guiana], Maroni, Comoro Is. [=Comoro Is., Maroni R.]. M (BMNH). Distr.: Colombia (Valle del Cauca); Venezuela (Cojedes); French Guiana (Maroni R.); Surinam (Paramaribo); Brazil (Distrito Federal, Espirito Santo, Goiás, Minas Gerais, Paraíba, Paraná, São Paulo); Argentina (Tucumán). Refs.: Steyskal 1973:145 (list); Knutson et al. 1976: 11 (cat.); Knutson & Valley 1978: 198 (review); Freidberg et al. 1991: 10 (key adult), 11 (key imm.), 16 (redesc.); figs 7, 4045, 97, 110, 113, 116, 126, 135, 138, 141; Marinoni & Mathis 2006: figs. 16, 29, 40; Vala et al. 2013: 121. isthmi (Steyskal) 1951: 291, pl. 4, fig. 20 (Sepedon). Type locality: Panama, Canal Zone, Corozal. M (USNM). Distr.: Panama (Canal Zone, Corozal, Juan Mina); Trinidad (Caroni River, Port-of-Spain); Venezuela (Aragua, Cojedes, Carabobo); Colombia (Valle); Bolivia (Rurrenabaque); Brazil (Amazonas, Goiás, São Paulo). Refs.: Steyskal, 1973: 145 (list); Knutson et al., 1976: 11 (cat.); Knutson & Valley, 1978: 197 (biol. and imm. stages); Freidberg et al., 1991: 9 (key adult), 11 (key imm.), 21 (redesc.), figs. 6, 9, 14-16, 52-57; Marinoni & Mathis, 2006: figs. 18, 31, 40. vau Mello & Bredt 1978a: 212 [nom. nud.] [Freidberg et al. 1991: 16 as Mello & Bredt 1978(b): 1459]. TABLE 1. Sciomyzidae species recorded from Colombia. Subfamily Tribe Genus Species Author Sciomyzinae Sciomyzinae Pherbellia guttata (Coquillett, 1901) Tetanocerini Dictya bergi Valley, 1977 Perilimnia albifacies Becker, 1919 Sepedomerus bipuncticeps (Malloch, 1933) macropus (Walker, 1849) guianica (Steyskal, 1951) isthmi (Steyskal, 1951) Sepedonea References Barker, G.M., Knutson, L., Vala, J.-C., Coupland, J.B. & Barnes, J.K. (2014) Overview of the biology of marsh flies (Diptera: Sciomyzidae), with special reference to predators and parasitoids of terrestrial gastropods. In: Barker, G.M. 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(1830) Systematische Beschreibung der bekannten europäischen zweiflügeligen Insekten. Vol. 6. H. W. Schultz, Hamm, xi + 401 pp., 12 pls. [pls. 55–66] Mello, D.A. & Bredt, A. (1978a) Distribuição geográfica de Sciomyzidae (Diptera: Insecta) no distrito federal e em algumas regiões de outros estados do Brasil. Ciência e Cultura, 30 (2), 212–215. Mello, D.A. & Bredt, A. (1978b) Estudos populacionais de cinco espécies de Sciomyzidae (Diptera-Insecta) no norte de Formosa, Goiás. Ciência e Cultura, 30 (12), 1459–1464. Murphy, W.L. (2014) Dictya behrmani, a new species of snail-killing fly (Diptera: Sciomyzidae) from Indiana. Proceedings of the Entomological Society of Washington, 11 (1), 91–97. Neff, S.E. & Berg, C.O. (1966) Biology and immature stages of malacophagous Diptera of the genus Sepedon (Sciomyzidae). Virginia Agricultural Experiment Station Bulletin, 566, 1–113. Poda, N. 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British Museum, London, pp. 689–1172. http://dx.doi.org/10.5962/bhl.title.20821 CATALOGUE OF SCIOMYZIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 647 Zootaxa 4122 (1): 648–652 http://www.mapress.com/j/zt/ Copyright © 2016 Magnolia Press Article ISSN 1175-5326 (print edition) ZOOTAXA ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.4122.1.54 http://zoobank.org/urn:lsid:zoobank.org:pub:3BE593E1-1D54-4E92-BB32-BFEE505D7A2E FAMILY SEPSIDAE VERA CRISTINA SILVA UNESP - Univ Estadual Paulista, Faculdade de Ciências Agrárias e Veterinárias, Departamento de Morfologia e Fisiologia Animal; Via de Acesso Prof. Paulo Donato Castellane, s/n; 14.806-900 Jaboticabal, SP, Brazil. E-mail: vcsilva@fcav.unesp.br Abstract An updated Catalogue of the Sepsidae of Colombia is presented. This small acalyptratae family is poorly known in Colombia, with only ten reported species in four genera. Key words: catalogue, Colombia, Diptera, distribution, Neotropical region Introduction The Sepsidae are a small family of the acalyptrate Diptera. Most members have an antlike appearance, caused by a basal constriction on the abdomen; they are small or medium in size (body length 2–7 mm), with wing hyaline, but sometimes with a dark spot near tip. The larvae are closely associated with various decaying organic materials, but most frequently on mammalian feces. The family is well represented in all biogeographic regions, with 318 valid species included in 37 valid genera (Ozerov 2005). Steyskal (1968) published the first catalogue of the Neotropical Sepsidae, surpassed by the World Catalogue by Ozerov (2005). A small paper about the family is available online in Spanish (Silva 1997) and the “Manual of Central American Diptera” chapter on the Sepsidae (Silva 2010) has a key to all genera known to occur in the Neotropics and a synopsis of the fauna of Central America. The Neotropical region presently includes 45 described valid species in 10 genera (Silva, submitted). In Colombia, only ten species in four genera are reported. Probably due to their habit, the species found in Colombia have a wide distributional range. Only a phylogeographic study will reveal if this is true or just an error caused by our fault in the recognition of the individuals at the species level in this morphological and ecological uniform fly family. All genus- and species-group names are listed alphabetically. Valid names are in bold type at the first mention. Synonyms when mentioned subsequently are italicized. Each published name including synonyms is given in a chronological sequence according to the scheme: the name of the species, author(s), year of description, page, genus in the original description (if different from the valid one), type-locality (or localities in the case of syntypes; updated name into brackets), information about the primary types and their deposition (and, if a lectotype, a reference to the author and date of the designation), known geographic distribution. At the end we present a table with all species found in Colombia, including rank and author (Table 1). Acronyms used for the depositories AMNH—American Museum of Natural History, New York, USA MCZ—Museum of Comparative Zoology, Harvard University, Cambridge, USA MNHN—Muséum National d’Histoire Naturelle, Paris, France MZUSP—Museu de Zoologia, Universidade de São Paulo, São Paulo, Brazil NHM—The Natural History Museum, London, U.K. NMW—Naturhistorisches Museum, Vienna, Austria 648 Accepted by S. Nihei: 10 Apr. 2015; published: 14 Jun. 2016 NRM—Naturhistoriska Riksmuseum, Stockholm, Sweden OUM—Oxford University Museum of Natural History, Oxford, U.K. USNM—National Museum of Natural History, Washington, USA ZMB—Museum für Naturkunde der Humboldt-Universität, Berlin, Germany List of abbreviations cat.—catalogue comm.—comments des.—designation distr.—geographic distribution orig. des.—by original designation phy.—phylogeny preocc.—preoccupied refs.—references rev.—revision subg.—subgenus syn.—synonym var.—variety HT—Holotype LT—Lectotype F (f#)—Female M (m#)—Male Catalogue of Sepsidae of Colombia Family Sepsidae Walker, 1833 Genus Archisepsis Silva Archisepsis Silva, 1993: 128. Type species, Sepsis armata Schiner, 1868 (orig. des.). Phalacrosepsis Ozerov, 1994: 77. Type species, Sepsis armata Schiner, 1868 (orig. des.). armata (Schiner, 1868: 261; Sepsis). Type locality: "aus Brasilien" [Brazil]. HT M (NMW). Distr.: Mexico, Cuba, Jamaica, Haiti, Dominican Republic, USA (Puerto Rico), Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Venezuela, Guyana, Ecuador, Peru, Brazil, Bolivia, Paraguay, Argentina, Uruguay. Refs.: Duda, 1926b (syn.); Silva, 1993 (rev.); Ozerov, 1994 (rev.); Zhao et al., 2013 (phy.). hoplicnema (Melander & Spuler, 1917: 17; Sepsis). Type locality: Haiti. HT M (USNM). discolor (Bigot, 1857: 346; Sepsis). Type locality: Cuba. HT F (MNHN). Distr.: Mexico, Cuba, Jamaica, Haiti, Dominican Republic, USA (Puerto Rico), Dominica, Grenada, Guatemala, El Salvador, Belize, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Venezuela, Guyana, Suriname, Ecuador, Peru, Brazil, Bolivia, Paraguay, Argentina; Palaearctic region [introduction]. Refs.: Ozerov, 1994 (rev.); Ozerov, 2005 (cat.); Zhao et al., 2013 (phy.). scabra (Loew, 1861: 42; Sepsis). Type locality: Cuba. HT F (MCZ). Distr.: Mexico to Paraguay and Argentina. Refs.: Silva, 1993 (rev.); Su et al., 2008 (phy.). haemorrhoidalis (Schiner, 1868: 261; Sepsis). Type locality: "aus Brasilien" [Brazil]. HT M (NMW). Distr.: Brazil. Refs.: Steyskal, 1968 (syn.). albicoxa (Thomson, 1869: 588; Sepsis). Type locality: “Brazil“. LT M (NRM). Distr.: Brazil. Refs.: Ozerov, 1992b (LT des.); Ozerov, 1994 (syn.). fuscus (Bigot, 1886: 387; ?Enicopus). Type locality: “Mexico“. HT F (OUM]. Distr.: Mexico. Refs.: Ozerov, 1999 (syn.). CATALOGUE OF SEPSIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 649 mequignoni (Séguy, 1936: 17; Sepsis). Type locality: Azores Is, Sãn Miguel, Sete Cidades. HT M (MNHN). Distr.: Azores. Refs.: Ozerov, 1993b (syn.). ecalcarata (Thomson, 1869: 588; Sepsis). Type locality: "California" [USA]. LT F (NRM). Distr.: Mexico, Cuba, Jamaica, Haiti, Dominican Republic, USA (Puerto Rico), Dominica, France (Martinique), Saint Lucia, Saint Vincent and the Grenadines, Grenada, Guatemala, Belize, El Salvador, Honduras, Nicaragua, Costa Rica, Panama, Colombia, Venezuela, Guyana, Ecuador, Peru, Brazil, Bolivia, Paraguay, Argentina; Nearctic region. Refs.: Ozerov, 1992b (LT des.); Ozerov, 1994 (rev.); Zhao et al., 2013 (phy.). cubensis (Bigot, 1886: 390; Nemopoda). Type locality: Cuba. LT M (OUM). Ref.: Ozerov, 1992b (LT des.); Ozerov, 1994 (syn.). xanthostoma (Bigot, 1886: 391; Nemopoda). Type locality: "Buenos Ayres" [Argentina, Buenos Aires]. LT F (OUM). Distr.: Argentina. Refs.: Ozerov, 1992b: 86 (LT des.); Ozerov, 1994 (syn.). insularis (Williston, 1896: 431; Sepsis). Type locality: St. Vincent [Saint Vincent and the Grenadines]. LT M (NHM). Distr.: St. Vincent, Ecuador, Brazil, Bolivia, Paraguay, Argentina. Refs.: Ozerov, 1993a (LT des.); Silva, 1993 (rev.); Ozerov, 1994 (syn.). bilobata (Becker, 1920: 206; Sepsis). Type-locality: Ecuador, Cuenca, 2532 m. HT M (MNHN). Distr.: Ecuador. Refs.: Ozerov, 1994 (syn.). excavata (Duda, 1926b: 87; Sepsis). Substitute name for Sepsis incisa Becker. Distr.: Belize, Guatemala, Nicaragua, Costa Rica, Panama, Colombia, Venezuela, Guyana, Ecuador, Peru, Brazil. Refs.: Silva, 1993 (rev.); Su et al., 2008 (phy.); Zhao et al., 2013 (phy.). incisa (Becker, 1920: 206; Sepsis). Junior primary homonym, preocc. by Sepsis incisa Strobl, 1894. Type locality: Ecuador Cuenca, 2532 m; LT F (MNHN). Refs.: Ozerov, 1992b: 86 (LT des.). pusio (Schiner, 1868: 262; Sepsis). Type locality: “Venezuela”. LT M (NMW). Distr.: Mexico, Cuba, Jamaica, Haiti, Dominican Republic, USA (Puerto Rico), Antigua and Barbuda, UK (Montserrat), Dominica, Saint Lucia, Saint Vincent and the Grenadines, Grenada, Guatemala, Honduras, El Salvador, Nicaragua, Costa Rica, Panama, Colombia, Venezuela, Guyana, Ecuador, Peru, Brazil, Bolivia, Paraguay, Argentina, Uruguay; Nearctic region. Refs.: Silva, 1993: 128 (rev.; LT des.); Ozerov, 1994 (rev.); Meier, 1995 (phy.); Su et al., 2008 (phy.); Zhao et al., 2013 (phy.). umbrifer (Schiner, 1868: 262; Sepsis). Type locality: “Venezuela”. LT M (NMW). Distr.: Colombia, Venezuela, Ecuador, Peru, Argentina. Refs.: Ozerov, 1992b: 85 (LT des.); Silva, 1993 (syn. with A. scabra); Ozerov, 1994 (rev.). Genus Lateosepsis Ozerov Lateosepsis Ozerov, 2004: 410. Type species, Sepsis laticornis Duda, 1926 (orig. des.). laticornis (Duda, 1926a: 45; 86–87, Pl. VIII, fig. 74; Sepsis). Type locality: Columbia [Colombia], Ujhelyi [sic], Sierra S. Lorenzo. LT M (ZMB). Distr.: Colombia (Magdalena), Venezuela, Ecuador, Bolivia, Brazil, Paraguay, Argentina. Refs.: Ozerov, 1992b: 86 (LT des.); Ozerov, 1994 (rev.); Silva, 1993 (rev.); Ozerov, 1997: 481–482. Comm.: The name Ujhelyi refered as the type locality by Duda is a Hungarian collector, József Ujhelyi, as recorded by Szilády (1926, apud Bequaert & Renjifo-Salcedo 1946) and Sándor (2013). Genus Microsepsis Silva Microsepsis Silva, 1993: 138. Type species, Sepsis armillata Melander & Spuler, 1917 (orig. des.). Ajdasepsis Ozerov, 1994: 56. Type species, Sepsis armillata Melander & Spuler, 1917 (orig. des.). furcata (Melander & Spuler, 1917: 19; Sepsis). Type locality: Jamaica. LT M (USNM). Distr.: Mexico, Cuba, 650 · Zootaxa 4122 (1) © 2016 Magnolia Press SILVA Jamaica, Dominican Republic, USA (Puerto Rico), Dominica, Saint Vincent and the Grenadines, Grenada, Guatemala, Belize, Honduras, El Salvador, Nicaragua, Costa Rica, Panama, Colombia, Venezuela, Guyana, Ecuador, Peru, Bolivia, Brazil, Paraguay, Argentina; Nearctic region. Refs.: Ozerov, 1993a: 75 (LT des.); Silva, 1993 (rev.); Ozerov, 1994 (rev.); Su et al., 2008 (phy.) ; Zhao et al., 2013 (phy.). mitis (Curran, 1927: 1; Sepsis). Type locality: Colombia, Vista Nieve, San Lorenzo Mts. HT M (AMNH). Distr.: Mexico, Dominican Republic, Guatemala, Belize, Honduras, Nicaragua, Costa Rica, Panama, Colombia (Magdalena), Venezuela, Ecuador, Bolivia, Brazil, Argentina. Refs.: Silva, 1993 (rev.); Ozerov, 1994 (rev.); Meier, 1995 (phy.); Zhao et al., 2013 (phy.). mystrion Silva, 1993: 143. Type locality: Brazil, Rio de Janeiro, Rio de Janeiro, Grajaú. HT M (MZUSP). Distr.: Brazil. Refs.: Ozerov, 2005: 23 (syn.). Genus Palaeosepsis Duda, 1926 Palaeosepsis Duda, 1926a: 43 [as a subg. of Sepsis]. Type species, Sepsis dentata Becker, 1919 (subsequent des. by Hendel, 1936: 62). maculata (Duda, 1926a: 46 [as var. of dentata]; Sepsis). Type locality: "Brasilien" [Brazil]. LT M (ZMB). Distr.: Colombia, Venezuela, Bolivia, Brazil. Refs.: Ozerov, 1992b: 86 (LT des.). TABLE 1. Sepsidae species recorded from Colombia. Subfamily Genus Species Author Sepsinae Archisepsis armata (Schiner, 1868) discolor (Bigot, 1857) ecalcarata (Thomson, 1869) excavata (Duda, 1926b) pusio (Schiner, 1868) umbrifer (Schiner, 1868) Lateosepsis laticornis (Duda, 1926a) Microsepsis furcata (Melander & Spuler, 1917) mitis (Curran, 1927) maculata (Duda, 1926a) Palaeosepsis References Becker, T. (1920) Diptères, Brachycéres. In: Mission du Service Géographique de l’Armée pour la mésure d’un Arc de Méridien Equatorial en Amérique du Sud sous le contrôle scientifique de l’Académie des Sciences, 1899-1906. Vol. X. Entomologie.–Botanie. Fascicule 2. –Opiliones. –Diptères–Myriapodes. Gauthiers-Villars, Paris, pp. 163–215. [1919] Bequaert, J.C. & S. Renjifo-Salcedo (1946) The Tabanidae of Colombia (Diptera). Psyche, 53 (3–4), 52–88. Bigot, J.M.F. (1857) Dipteros. In: Sagra, R. de la (Ed.), Historia fisica politica y natural de la Isla de Cuba. Secunde parte. Historia natural. Tomo VII. A. Bertrand, Paris, pp. 328– 349, pl. 20. [1856] Bigot, J.M.F. (1886) Diptères nouveaux ou peu connus. 29e partie (suite) XXXVII 2e. Essai d’une classification synoptique du groupe des Tanypezidi (mihi) et description de genres et d’espèces inèdits. Annales de la Société Entomologique de France, 6, 369–392. Curran, C.H. (1927) Four new American Diptera. American Museum Novitates, 275, 1–4. Duda, O. (1926a) Monographie der Sepsiden (Dipt.). I. Annalen des Naturhistorischen Museums in Wien, 39, 1–153. [1925] Duda, O. (1926b) Monographie der Sepsiden. (Dipt.). II. Annalen des Naturhistorischen Museums in Wien, 40, 1–110. Hendel, F. (1936) Ergebnisse einer zoologischen Sammelreise nach Brasilien, insbesondere in das Amazonasgebiet, ausgefürt von Dr. H. Zerny. Annalen des Naturhistorischen Museums in Wien, 47, 61–106. Loew, H. (1861) Diptera aliquot in insula Cuba collecta. Wiener Entomologische Monatschrift, 5, 33–43. CATALOGUE OF SEPSIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 651 Meier, R. (1995) Cladistic analysis of the Sepsidae (Cyclorrhapha: Diptera) based on a comparative scanning electron microscopic study of larvae. Systematic Entomology, 20, 99–128. http://dx.doi.org/10.1111/j.1365-3113.1995.tb00086.x Melander, A.L. & Spuler, A. (1917) The Dipterous families Sepsidae and Piophilidae. Bulletin State College of Washington Agricultural Experiment Station, Pullman, Washington, 143, 1–103. Ozerov, A.L. (1992a) K taksonomii dvukrylykh semeystva Sepsidae (Diptera) [On the taxonomy of flies of the family Sepsidae (Diptera)]. Byulleten’ Moskovskogo obshchestva ispytateley prirody, Otd. Biol., 97 (4), 44– 47. [in Russian] Ozerov, A.L. (1992b) New data on the neotropical sepsids (Diptera Sepsidae). Russian Entomological Journal, 1 (1), 81–86. Ozerov, A.L. (1993a) The results of the examination of the syntypes of Sepsis insularis Williston,1896 and Sepsis furcata Melander et Spuler, 1917 (Diptera Sepsidae). Russian Entomological Journal, 1 (2), 75. [1992] Ozerov, A.L. (1993b) Four new synonyms in the family Sepsidae (Diptera). Revue d’Entomologie de l’URSS, 72 (1), 248–249. Ozerov, A.L. (1994) A review of the sepsid flies of Palaeosepsini, trib.n. (Diptera Sepsidae). Russian Entomological Journal, 2 (5–6), 55–103. [1993] Ozerov, A.L. (1997) Some types of Sepsidae in the Berlin and Eberswalde Museums (Diptera). Beiträge zur Entomologie, 47 (2), 477–487. Ozerov, A.L. (1999) A review of the genus Themira Robineau-Desvoidy, 1830 (Diptera, Sepsidae) of the World, with a revision of the North American species. Russian Entomological Journal, 7 (3-4), 169–208. [1998] Ozerov, A.L. (2004) New genera and species of Sepsidae (Diptera). Russian Entomological Journal, 12 (4), 399–420. [2003] Ozerov AL. (2005) World Catalogue of the family Sepsidae (Insecta: Diptera). Zoologicheskie issledovania, 8, 1–74. Sándor, T. (2013) Magyarország fürkészlégy faunája (Diptera: Tachinidae). e-Acta Naturalia Pannonica, 5, Supplementum 1, 5–314. Available from: http://epa.oszk.hu/01900/01957/00009/pdf/EPA01957_e-acta_5_2013_suppl_311.pdf. (accessed 27 February 2014) Schiner, I.R. (1868) Diptera. In: Wüllerstorf-Urbair, B. von (Ed.), Reise der Österreichischen Fregatte Novara um die Erde in den Jahren 1857, 1858, 1859, unter den Befehlen des Commodore B. von Wüllerstorf-Urbair. Zoologischer Theil. Zweiter band. 1. Abtheilung. B. K. Gerond’s Sohn, Wien, pp. I–VI + 1–388. http://dx.doi.org/10.5962/bhl.title.1597 Séguy, E. (1936) Voyage de MM. L. Chopard et A. Méquignon aux Açores (août-septembre 1930). X. Diptères. Annales de la Société Entomologique de France, 105, 11–26. Silva, V.C. (1993) Revisão da família Sepsidae na região Neotropical. III. Os gêneros Palaeosepsis Duda, 1926, Archisepsis gen.n. e Microsepsis gen.n.; chave para os gêneros Neotropicais (Diptera, Schizophora). Iheringia, Série Zoologia, Porto Alegre, 75, 117–170. Silva, V.C. (1997) Sepsidae. In: Solís, A. (Ed.), Las famílias de insetos de Costa Rica. InBio. Available from: http:// www.inbio.ac.cr/papers/insectoscr/Texto639.html (accessed 11 February 2014) Silva, V.C. (2010) Chap. 77, Sepsidae (black scavengers flies). In: Brown, B., Wood, M. & Zumbado, M. (Eds.), Manual of Central American Diptera. Vol. 2. National Research Council Press, Ottawa, Ontario, Canada, pp. 1031–1039. Silva, V.C. (2015) Manual of the Neotropical Sepsidae (Diptera: Schizophora). Neotropical Diptera. [submitted] Steyskal, G.C. (1968) 63. Family Sepsidae. In: Papavero, N. (Ed.), A Catalogue of the Diptera of the American South of the United States. Departamento de Zoologia, Secretaria da Agricultura, São Paulo, 6 pp. Su, K.F.-Y, Kutty, S.N. & Meier, R. (2008) Morphology versus molecules: The phylogenetic relationships of Sepsidae (Diptera: Cyclorrhapha) based on morphology and DNA sequence data from ten genes. Cladistics, 24, 902–916. http://dx.doi.org/10.1111/j.1096-0031.2008.00222.x Szilády, Z. (1926) New and Old World horseflies. Biologica Hungarica, 1 (7), 1–30. Thomson, C.G. (1869) 6. Diptera. Species nova descripsit. In: Kongliga Svenska fregatten Eugenies resa omkring Jordan under befäl af C.A. Virgin, aren 1851-1853. Vetenskapliga iakttagelser pa H. Maj:t konung Oscar den förstes befallning utgifna af K. Svenska Vetenskaps-Akademien. Vol. 2 (Zoologi), [section] 1, (Insecta). P.A. Norstedt & Söner, Stockholm, pp. 443–614. Walker, F. (1833) Observations on the British species of Sepsidae. The Entomological Magazine (London), 1, 244–256. Williston, W. (1896) On the Diptera of St. Vincent (West Indies). The Transactions of the Entomological Society of London, 1896, 253–446, pls. 8–14. http://dx.doi.org/10.1111/j.1365-2311.1896.tb00965.x Zhao, L., Annie, A.S.H., Amrita, S., Yi, S.K.F. & Meier, R. (2013) Does better taxon sampling help? A new phylogenetic hypothesis for Sepsidae (Diptera: Cyclorrhapha) based on 50 new taxa and the same old mitochondrial and nuclear markers. Molecular Phylogenetics and Evolution, 69, 153–169. http://dx.doi.org/10.1016/j.ympev.2013.05.011 652 · Zootaxa 4122 (1) © 2016 Magnolia Press SILVA Zootaxa 4122 (1): 653–658 http://www.mapress.com/j/zt/ Copyright © 2016 Magnolia Press Article ISSN 1175-5326 (print edition) ZOOTAXA ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.4122.1.55 http://zoobank.org/urn:lsid:zoobank.org:pub:1840AD78-E136-4440-AB94-3725DD14C5F6 FAMILY CLUSIIDAE MARTA WOLFF1* & DIANA GRISALES2 1 Grupo de Entomologia, Instituto de Biologia, Universidad de Antioquia. Colombia. Laboratório de Biodiversidade e Biogeografia de Diptera, Departmento de Zoologia, UFPR, Caixa postal 19020, Curitiba, PR., Brazil, 81531-980. E-mail: martha.wolff@udea.edu.co; ochoa310@gmail.com * Corresponding author: martha.wolff@udea.edu.co 2 Abstract Here we catalogued 23 species in two genera of Clusiidae from Colombia. For each species the available geographical information is presented. Key words: catalogue, Colombia, Diptera, distribution Introduction Clusiidae is a Brachycera “Acalyptratae” family with worldwide distribution. Adults measure between 3–6 mm length; color yellow or dark brown and wing partially infuscate or hyaline; head round with three ocelli and ocellar bristles present; vibrissa developed; antenna porrect, arista dorsoapical and pedicel with triangular extension on margins; costal vein without breaks; and abdomen slender. This catalog is based on the studies of Lonsdale & Marshall (2006a,b; 2007a,b; 2012). For Colombia, there are registered two subfamilies, 23 species of Clusiidae in two genera (Table 1). The Colombian departments are underlined and parenthetically notes specify localities. At the end, we provide a table that includes all species known in the country (Table 1). Acronyms used for the depositories BMNH—The Natural History Museum, London, England. CNC—Canadian National Collection of Insects, Arachnids & Nematodes, Ottawa, Canada. DEBU—University of Guelph Insect Collection, Guelph, Ontario, Canada. HNHM—Zoological Department of the Hungarian Natural History Museum, Budapest, Hungary. INBio—Instituto Nacional de Biodiversidad, Santo Domingo de Heredia, Costa Rica. QCAZ—Universidad Católica del Ecuador, Quito, Ecuador. USNM—National Museum of Natural History, Smithsonian Institution, Washington, D. C. USA. List of abbreviations chang. comb.—changed combination comb. nov.—new combination by des.—designation by distr.—geographic distribution m.—meters Accepted by S. Nihei: 26 May 2015; published: 14 Jun. 2016 653 mon.—monotypy orig. des.—by original designation pp.—pages redesc.—redescription refs.—references syn.—Synonym HT—Holotype LT—Lectotype F (f#)—Female M (m#)—Male Catalogue of Clusiidae of Colombia Family Clusiidae Handel, 1916 Subfamily Clusiodinae Frey, 1960 Genus Craspedochaeta Czerny Craspedochaeta Czerny, 1903: 108. Type species, Craspedochaeta transversa Czerny, 1903 (mon.). Ref.: Lonsdale & Marshall, 2006b (rev.). Czernyola Bezzi, 1907: 52. Type species, Czernyola transversa (Czerny, 1903) (chang. comb.). Tonnoiria Malloch, 1929: 97. Type species, Tonnoiria palliseta Malloch, 1929. (orig. des.) Chaetoczernyola Sóos, 1962: 135. Type species, Chaetoczernyola novaeguineae Soós, 1962. (orig. des.). concinna species group concinna (Williston), 1896: 387 (Heteroneura). Type locality: St. Vincent and the Grenadines. St Vincent “Leeward side”. LT M (BMNH). Distr.: Argentina, Belize, Bolivia, Brazil, Colombia, Costa Rica, Ecuador, Grenada, Guatemala, Guyana, Honduras, Jamaica, Mexico, Panama, Peru, St. Vincent, Trinidad, Venezuela, United States. Ref.: Lonsdale & Marshall, 2006b: 52 (rev. and redesc.) atra Kertész, 1903: 570. atra, Melander & Argo, 1924: 25. (Czernyola). minuta (Soós), 1962: 148 (Czernyola). albohalteria (Soós), 1962: 145–146 (Czernyola). transversa species group pleuralis (Williston), 1896: 387 (Heteroneura) (comb. nov. Lonsdale & Marshall, 2006b: 44). Type locality: St. Vincent. HT M (BMNH). Distr.: Argentina, Bolivia, Brazil, Colombia, Costa Rica, Cuba, Dominican Republic, Ecuador, El Salvador, Grenada, Mexico, Panama, Peru, Puerto Rico, St. Vincent, Trinidad, Venezuela. Ref.: Lonsdale & Marshall, 2006b: 44. Genus Heteromeringia Czerny Heteromeringia Czerny, 1903: 72. Type species, Heteromeringia nigrimana (Loew, 1864) (mon.). Refs.: Lonsdale & Marshall, 2007b (rev.). czernyi species group czernyi Kertész, 1903: 568. Type locality: Peru, Callanga. HT F (HNHM). Distr.: Argentina, Belize, Bolivia, 654 · Zootaxa 4122 (1) © 2016 Magnolia Press WOLFF & GRISALES Brazil, Colombia, Costa Rica, Cuba, Dominica, Dominican Republic, Ecuador, Guatemala, Guyana, Honduras, Jamaica, Mexico, Panama, Peru, Puerto Rico, St. Lucia, Tobago, Trinidad, United States (FL), Venezuela. Ref.: Lonsdale & Marshall, 2007a: 54 (rev.) subfasciata (Curran), 1939: 2 (Sobarocephala). Subfamilia Sobarocephalinae Lonsdale & Marshall, 2006 Genus Sobarocephala Czerny Sobarocephala Czerny, 1903: 85. Type species, Sobarocephala ruebsaameni Czerny (mon.). Ref.: Soós, 1968: 4 (cat.); Sabrosky & Steyskal, 1974: 371; Lonsdale & Marshall, 2007b: 65 (rev.). Meriza Kertész, 1903: 571. Type species, Sobarocephala dorsata Czerny, 1903 (orig. des.). Syn. Melander & Argo (1924). Monorrhexa Kertész, 1903: 572. Type species, pictipennis Kertész (mon.). latipennis species group araguamyia Lonsdale & Marshall, 2012: 40. Type locality: Colombia, Vaupés, río Apoporis, Caparu Biol. Station. HT M (DEBU). Distr.: Colombia (Vaupés (Caparu)), Bolivia, Brazil, Costa Rica, Panama, Venezuela. liturata Melander & Argo, 1924: 43. Type locality: Costa Rica, La Suiza de Turrialba. HT F (USNM). Distr.: Colombia (Valle del Cauca (río Raposo), Amazonas (Leticia)), Bolivia, Brazil, Costa Rica, Ecuador, Guatemala, Guyana, Jamaica, Panama, Peru, Surinam, Trinidad. Ref.: Lonsdale & Marshall, 2012: 45 (rev., redescription). imitans Curran 1934: 443. Lonsdale & Marshall, 2012: 45 (syn.) diversipes Curran 1939: 2. Lonsdale & Marshall, 2012: 45 (syn.) ruebsaameni species-group cacumen Lonsdale & Marshall, 2012: 53. Type locality: Peru, Madre de Dios, Manu, Erika (nr-Salvacion), 500m. HT M (USNM). Distr.: Colombia (Antioquia (Quebrada La Espadera, 7km E. Medellín) as “Quep. Espareda”, error)), Belize, Costa Rica, Honduras, Mexico, Panama, Peru. Ref: Lonsdale & Marshall, 2012: 53 (rev.). isla species group isla Curran, 1939: 1. Type locality: Panama, Canal Zone, Barro Colorado Island. HT M (AMNH). Distr.: Colombia (Amazonas (Leticia) (as “Amazones Letitia”), Argentina, Bolivia, Brazil, Costa Rica, Ecuador, Guatemala, Guyana, Honduras, Mexico, Panama, Peru, Venezuela. Ref.: Lonsdale & Marshall, 2012: 74 (rev.). minthomyia Lonsdale & Marshall, 2012: 79. Type locality: Mexico, Chis., 7000’, 20 min N Bochil. HT M (CNC). Distr.: Colombia (Cundinamarca (30km SW. Bogotá, Tequendama Falls)), Costa Rica, Mexico. Ref.: Lonsdale & Marshall, 2012: 79 (rev.). interrupta species group (excluding the pectinaria subgroup) dichotomos Lonsdale & Marshall, 2012: 87. Type locality: Costa Rica, Puntarenas, Pen Osa, 5km N Puerto Jiménez, 10m (as Puerto Jiminez, 10m). HT M (INBio). Distr.: Colombia (Amazonas (Leticia), Tolima (Armero)), Argentina, Brazil, Costa Rica, Dominica, Ecuador, El Salvador, Honduras, Mexico, Panama, St. Vincent, Surinam, Trinidad, Venezuela. Lonsdale & Marshall, 2012: 87 (rev.). CATALOGUE OF CLUSIIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 655 pubens Lonsdale & Marshall, 2012: 94. Type locality: Costa Rica, Alajuela, Upala, Dos Rios, Estación San Gerrardo, 600m. HT M (INBio). Distr.: Colombia (Valle del Cauca (río Raposo)), Costa Rica. Ref.: Lonsdale & Marshall, 2012: 94 (rev.). interrupta species group pectinaria subgroup angulata Lonsdale & Marshall, 2012: 100. Type locality: Costa Rica, Puntarenas, Península de Osa, 5km N Puerto Jiménez, 10m (as Pen Osa, 5km N of Puerto Jiminez, 10m). HT M (INBio). Distr.: Colombia (Tolima (Armero)), Bolivia, Brazil, Costa Rica, Ecuador, Mexico, Peru, Trinidad, Venezuela. Ref.: Lonsdale & Marshall, 2012: 100 (rev.). conciliata Lonsdale & Marshall, 2012: 112. Type locality: Costa Rica, Limon, Bribri, 4Km NE, 50m. HT M (INBio): Distr.: Colombia (Valle del Cauca (río Raposo)), Argentina, Brazil, Costa Rica, Ecuador, Grenada, Paraguay, Peru, Trinidad, Venezuela. Ref.: Lonsdale & Marshall, 2012: 112 (rev.). epitriangulina Lonsdale & Marshall, 2012: 117 (Sobarocephala). Type locality: Costa Rica, Puntarenas: R.F. Golfo Dulce, 24km W Piedras Blancas, 200m. HT M (INBio). Distr.: Colombia (Tolima (Armero)), Argentina, Costa Rica, Guatemala, Mexico, Panama. Ref.: Lonsdale & Marshall, 2012: 117 (rev.). periglenes Lonsdale & Marshall, 2012: 136. Type locality: Colombia, Valle del Cauca, río Raposo. HT M (USNM). Distr.: Colombia (Valle del Cauca (río Raposo)). Ref.: Lonsdale & Marshall, 2012: 136 (rev.). spatulata Lonsdale & Marshall, 2012: 142. Type locality: Peru, Madre de Dios: Manu, Rio Manu, 250m, Pakitza, 12°07’S, 70°58’W. HT M (USNM). Distr.: Colombia ((Meta (3mi W Villavicencio, 920m)), Bolivia, Brazil, Costa Rica, Ecuador, Guyana, Peru, Venezuela. Ref.: Lonsdale & Marshall, 2012: 142 (rev.). subtriangulina Lonsdale & Marshall, 2012: 145. Type locality: Ecuador, Napo, Tena. HT M (QCAZ). Distr.: Colombia (Amazonas (Leticia) (as Letica), Bolivia, Costa Rica, Ecuador, Panama, Peru, Venezuela. Ref.: Lonsdale & Marshall, 2012: 145 (rev.). vestigialis Lonsdale & Marshall, 2012: 151. Type locality: Costa Rica, Puntarenas, Est. Cabuya, Rio Cabo Blanco, 50m. HT M (INBio). Distr.: Colombia (Meta (3mi W Villavicencio, 920m)), Brazil, Costa Rica, Guyana, Ecuador, Mexico, Panama, Peru, Venezuela. Ref.: Lonsdale & Marshall, 2012: 151 (rev.). flaviseta species group (S. plumata clade) columbiensis Soós 1962: 387. Type locality: Colombia, Ujhelyi. HT M (HNHM). Distr.: Colombia (Magdalena (Aracataca, Sierra San Lorenzo)). Ref: Lonsdale & Marshall, 2012: 184 (rev.). parallela Lonsdale & Marshall, 2012: 203. Type locality: Colombia, Tolima, Armero (as “Armoro”). HT M (USNM). Distr.: Colombia (Tolima (Armero)), Bolivia. Ref: Lonsdale & Marshall, 2012: 203 (rev.) plumata Melander & Argo, 1924: 46. Type locality: Panama, Canal Zone, Alajuela. HT F (USNM). Distr.: Colombia (Valle del Cauca (Buenaventura)), Bahamas, Bolivia, Costa Rica, Grand Cayman, Panama. Ref.: Lonsdale & Marshall, 2012: 208 (rev. and redesc). quadrimaculata Soós, 1963: 394. Type locality: United States, Florida, Dade Co., Evereglades National Park. HT 656 · Zootaxa 4122 (1) © 2016 Magnolia Press WOLFF & GRISALES F (USNM). Distr.: Colombia, Bahamas, Belize, Costa Rica, Mexico, Nicaragua, Panama, southeastern United States. Refs.: Sabrosky & Steyskal 1974: 374; Lonsdale & Marshall 2007b: 95 (rev, redesc); Lonsdale & Marshall, 2012: 209 (rev.). nitida Soós 1963: 393. Sabrosky & Steyskal, 1974 (syn.). Acknowledgments Thanks to Dr. Owen Lonsdale for send his papers to MW. To Estrategia de Sostenibilidad 2015-2015 University of Antioquia for the grants to MW and to the Conselho Nacional de Desenvolvimento Científico e Tecnológico for their postdoctoral support to DG. TABLE 1. Clusiidae species recorded from Colombia Subfamily Genus Species Author Clusidinae Craspedochaeta concinna (Williston, 1896) pleuralis (Williston, 1896) Heteromeringia czernyi species group Sobarocephalinae Sobarocephala czernyi Kertesz, 1903 columbiensis Soós, 1962 quadrimaculata Soós, 1963 araguamyia Lonsdale & Marshall, 2012 liturata Melander & Argo, 1924 cacumen Lonsdale & Marshall, 2012 isla Curran, 1939 minthomyia Lonsdale & Marshall, 2012 latipennis species group ruebsaameni species group isla species group interrupta species group (excluiding S. pectinaria subgroup) dichotomos Lonsdale & Marshall, 2012 pubens Lonsdale & Marshall, 2012 interrupta species group pectinaria subgroup angulata Lonsdale & Marshall, 2012 conciliata Lonsdale & Marshall, 2012 epitriangulina Lonsdale & Marshall, 2012 periglenes Lonsdale & Marshall, 2012 spatulata Lonsdale & Marshall, 2012 subtriangulina Lonsdale & Marshall, 2012 vistiagilis Lonsdale & Marshall, 2012 flaviseta species group (S. plumata clade) CATALOGUE OF CLUSIIDAE OF COLOMBIA columbiensis Soós, 1962 parallela Lonsdale & Marshall, 2012 plumata Melander & Argo, 1924 quadrimaculata Soós, 1963 Zootaxa 4122 (1) © 2016 Magnolia Press · 657 References Bezzi, M. (1907) Nomenklatorisches über Dipteren. Weiner Entomologische Zeitung , 26, 51–56. Curran, C.H. (1934) The Diptera of Kartago, Bartica District, British Guiana, with descriptions of new species from other British Guiana localities. Bulletin of the American Museum of Natural History, 66, 287–532. Curran, C.H. (1939) Four new species of Sobarocephala (Diptera: Clusiidae). American Museum Novitates, 1029, 1–2. Czerny, P.L. (1903) Revision der Heteroneuriden. Weiner Entomologische Zeitung, 22 (3), 61–108. Kertész, K. (1903) Beiträge zur kenntniss der Heteroneuriden. Annales Musei Nationalis Hungarici, 1, 566–573. Lonsdale, O. & Marshall, S.A. (2006a) Redefinition of the Clusiinae and Clusiodinae, description of the new subfamily Sobarocephalinae, revision of the genus Chaetoclusia and a description of Procerosoma gen. n. (Diptera: Clusiidae). European Journal of Entomology, 103, 163–182. Lonsdale, O. & Marshall, S.A. (2006b) Revision of the New World Craspedochaeta Czerny. Zootaxa, 1291, 1–101. Lonsdale, O. & Marshall, S.A. (2007a) Revision of the New World Heteromeringia (Diptera: Clusiidae: Clusiodinae). Beiträge zur Entomologie, 57 (1), 37–80. Lonsdale, O. & Marshall, S.A. (2007b) Revision of the North American Sobarocephala (Diptera: Clusiidae, Sobarocephalinae). Journal of the Entomological Society of Ontario, 138, 65–106. Lonsdale, O. & Marshall, S.A. (2012) Sobarocephala (Diptera: Clusiidae: Sobarocesphalinae) Subgeneric classification and Revision of the New World species. Zootaxa, 3370, 1–307. Malloch, J.R. (1929) Exotic Muscaridae (Diptera)-XXVI. Annals of the Magazine of Natural History, 10 (4), 97–120. http://dx.doi.org/10.1080/00222932908673032 Melander, A.L. & Argo, N.G. (1924) Revision of the two-winged flies of the family Clusiidae. Proceedings of the United States National Museum, 64 (11), 1–54. http://dx.doi.org/10.5479/si.00963801.64-2501.1 Sabrosky, C.W. & Steyskal, G.C. (1974) The genus Sobarocephala (Diptera: Clusiidae) in America North of Mexico. Annals of the Entomological Society of America, 67 (3), 371–385. http://dx.doi.org/10.1093/aesa/67.3.371 Soós, A. (1962) New genera and species of Clusiidae (Diptera, Acalyptrata). Acta Zoologica Academiae Scientiarum Hungaricae, 8 (1–2), 135–149. Soós, A. (1963) Identification key to the species of the “plumata-group” of the genus Sobarocephala Czerny (Diptera: Clusiidae). Acta Zoologica Academiae Scientiarum Hungaricae, 9 (3–4), 391–396. Soós, A. (1968) 84. Family Clusiidae. In: Papavero, N. (Ed.), A catalogue of the Diptera of the Americas south of the United States. Museu de Zoologia, Universidade de São Paulo, São Paulo, pp. 13. Williston, S.W. (1896) On the Diptera of St. Vincent (West Indies). Transactions of the Entomological Society of London, 1896, 253–446. http://dx.doi.org/10.1111/j.1365-2311.1896.tb00965.x 658 · Zootaxa 4122 (1) © 2016 Magnolia Press WOLFF & GRISALES Zootaxa 4122 (1): 659–676 http://www.mapress.com/j/zt/ Copyright © 2016 Magnolia Press Article ISSN 1175-5326 (print edition) ZOOTAXA ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.4122.1.56 http://zoobank.org/urn:lsid:zoobank.org:pub:7EE22515-9C7B-4EA7-B377-73D9343CF3C9 FAMILY AGROMYZIDAE VIVIANE RODRIGUES DE SOUSA* & MÁRCIA SOUTO COURI Museu Nacional, Universidade Federal do Rio de Janeiro. * Corresponding author: sousavrodrigues@gmail.com Abstract Agromyzidae are a family with many species described throughout the world. The family is known as leaf-miner flies, for the phytophagous habits of their larvae, feeding within plant tissues. This catalogue presents 74 species recorded from Colombia, distributed in 12 genera and two subfamilies. For each species we present the available geographical distribution in the country. Key words: Agromyzidae, catalogue, Colombia, distribution, leaf-miners Introduction Agromyzidae are small to median sized flies; most of the species ranging from 2–3 mm, with a maximum size of 6.5 mm. The body color is yellow, black, brown, gray, green metallic and some species show a coppery coloration. Antennae with flagelomere short and round, sometimes elongated, widened, with a projection or spine. Frontoorbital and ocellar setae are strong, postocellars divergent, fronto-orbitals in one or more rows, usually reclinate, but sometimes proclinate. Scutum with two to five dorsocentral setae, acrostichals present or absent. Costal break present; Sc vein incomplete distally or reaching C as a linear fold, independently of R1 or fusing with R1 before reaching C; CuA2 and A1 present, forming a small cell cup; A1 not reaching wing margin (Boucher 2010). Boucher (2010) summarized the relationship of Agromyzidae with other Diptera families, which remains unclear. According to Hennig (1958), Agromyzidae is more closely related to Odiniidae, or possibly to Clusiidae, while Spencer (1969) suggested proximity between Odiniidae and Agromyzidae, based on genital similarities. Odiniidae and Clusiidae were pointed out as sister groups of Agromyzidae, based on a cladistic analysis and a study of the male postabdomen, respectively by Dempewolf (2001) and Griffiths (1972). McAlpine (1989) included Agromyzidae, together with Odiniidae and Fergusoninidae, in the superfamy Agromyzoidea. The family is generally divided into two subfamilies: Agromyzinae and Phytomyzinae, differentiated by the shape of subcostal vein and larval cephalopharyngeal skeleton. The larvae of Agromyzidae species are phytophagous and attack various plant parts such as leaf, root, stem, seed and flower. Most species are leafminers, and cause great economic losses. The family has a large worldwide distribution, with 2,800 species, 469 of them, distributed in 16 genera, present in the Neotropical region (Boucher 2010; Martinez & Etienne 2002). Martinez & Etienne (2002) removed the records of Liriomyza bryoniae (Kaltenbach) and Calycomyza artemisiae (Kaltenbach) from the neotropics, as they were incorrect records. This catalogue presents 74 species recorded from Colombia, distributed in 12 genera and two subfamilies (Table 1). For each species we present the available geographical distribution in the country. Acronyms used for the depositories BC—Blanchard’s collection. BMNH—The Natural History Museum [formerly British Museum (Natural History)], London, United Kingdom. Accepted by S. Nihei: 16 Apr. 2015; published: 14 Jun. 2016 659 CAS—California Academy of Sciences, San Francisco, California, USA. ICNDIPAg—Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Santafé de Bogotá. USNM—National Museum of Natural History [formerly, United States National Museum], Washington D.C., USA. MCZ—Museum of Comparative Zoology, Harvard University Cambridge, Massachusetts, USA. SC—Spencer’s collection. This collection is deposited in BMNH. List of abbreviations cat.—catalogue distr.—geographic distribution mon.—monotypy n. comb.—new combination nom. nov.—nomen novum n. syn.—new synonym orig. des.—original designation pp.—pages preocc.—preoccupied refs.—references rev.—revision subg .—subgenus subs. des.—subsequent designation HT—Holotype NT—Neotype F (f#)—Female M (m#)—Male Catalogue of Agromyzidae of Colombia Family Agromyzidae Fallén, 1823 Subfamily Agromyzinae Hendel, 1931 Genus Calycomyza Hendel Calycomyza Hendel, 1931: 65 (as subgenus of Dizygomyza). Type species, Agromyza artemisiae Kaltenbach, 1856 (orig. des.). Refs.: Frick, 1957a: 59-70 (as subgenus), Nowakowski, 1962: 97 (as genus). colombiana Sasakawa, 1992b: 13. Type locality: Colombia, Buenaventura. HT M (CAS). Distr.: Colombia (Valle del Cauca (Buenaventura)). Refs.: Martinez & Etienne, 2002: 29 (list). ipomaeae Frost, 1931: 74 (Agromyza). Type locality: Puerto Rico. HT M (USNM). Distr.: Colombia (Cundinamarca (Finca Bella Vista, nr. Sasaima)), Barbade, Brazil (Santos), Canada, Chile, Cuba, Guadalupe, Grand-Cayman, Jamaica, La Dominique, Nicaragua, Peru, Puerto Rico, Dominican Republic, Saint-Martin, U.S.A (Florida), West Indies, Venezuela. Refs.: Spencer, 1963: 345; Spencer, 1967: 83.9 (cat.); Spencer, 1969: 150; Spencer, 1973a: 44; Spencer & Stegmaier, 1973: 76; Spencer, 1984: 26 (Colombia, cit., key); Spencer & Steyskal, 1986: 150 (key); Sasakawa, 1992b: 15 (Colombia); Spencer et al., 1992: 288; Martinez et al., 1993: 177; Martinez & Etienne, 2002: 29 (list); Korytkowski, 2014: 26. Phytobia (Calycomyza) ipomaeae, Frick, 1952: 395; 1957: 203; Spencer, 1963: 345 Calycomyza ipomaeae, Spencer, 1969: 150. 660 · Zootaxa 4122 (1) © 2016 Magnolia Press DE SOUSA & COURI lantanae Frick, 1956: 287 (Calycomyza as subgenus of Phytobia). Type locality: “Trinidad”. HT M (USNM). Distr.: Colombia (Valle del Cauca (Cali, Rio Cali)), Argentina, Bahamas, Barbade, Brazil, Galapagos, Grenade, Guadalupe, Jamaica, Mexico, Peru, Puerto Rico, San Martin, Tobago, Trinidad, Venezuela, U.S.A. (Florida and Texas), Afrotropical and Australian regions. Refs.: Spencer, 1963: 346 (male terminalia); Spencer, 1967: 83.9 (cat.); Spencer, 1973a: 45; Valladares, 1981: 226; Spencer, 1984: 26. (Colombia, cit., key); Spencer & Steyskal, 1986: 142 (key); Valladares, 1992: 193 (key); Martinez & Etienne, 2002: 29 (list); Sasakawa, 2007: 194. Calycomyza lantanae, Spencer & Stegmaier, 1973:78. meridiana Hendel, 1923: 123 (Agromyza, n. name for A. meridionalis). Type locality: “Costa Rica”. HT F (USNM). Distr.: Colombia (Cundinamarca (10 mil. W of Villeta, 1,660 m)), Brazil, British Guiana, Costa Rica, El Salvador, West Indies, Mexico, Peru, Venezuela. Refs.: Spencer, 1963: 348 (male terminalia); Spencer, 1967: 83.9 (cat.); Spencer, 1973a: 46; Spencer, 1973b: 153; Spencer & Stegmaier, 1973: 185; Sasakawa, 1992b: 15 (as Calycomyza); Sasakawa, 1992c: 823; Sasakawa, 1992d: 25; Sasakawa, 2005: 158; Martinez & Etienne, 2002: 30 (list); Korytkowski, 2014: 28. meridionalis Malloch, 1914: 89 (Agromyza; preocc. Strobl, 1900). Phytobia (Calycomyza) meridiana, Frick, 1952: 395; Spencer, 1963: 348. Calycomyza meridiana, Sasakawa, 1992b: 15. richardsi Spencer, 1963: 348 (Calycomyza as subgenus of Phytobia). Type locality: British Guiana, Mazaruni. HT M (SC). Distr.: Colombia (Valle del Cauca (Cali, 6 mil. W of Cali Valle, 1,630 m)), Guadalupe, Guyana. Refs.: Spencer, 1967: 83.9 (cat.); Spencer et al., 1992: 293; Sasakawa, 1992b: 16 (as Calycomyza); Martinez & Etienne, 2002: 30 (list). Calycomyza richardsi, Sasakawa, 1992b: 16. steviae Spencer, 1973a: 48. Type locality: Venezuela, Merida, (roadside 10 km above Timotes). HT M (SC). Distr.: Colombia (Tequedama (Bogotá to Girardot)), Venezuela. Refs.: Spencer, 1984: 26 (Colombia, key); Martinez & Etienne, 2002: 30 (list). verbesinae Spencer, 1963: 351 (Calycomyza as subgenus of Phytobia). Type locality: Brazil, São Paulo, Agua Funda. HT M (SC). Distr.: Colombia (Caldas (5 mil. N of Anserma, 1,750 m), Cundinamarca (10 mil. W of Villeta 1,660 m; Finca Bella Vista), Meta (3 mil. W of Villavicencio, 920 m), Valle del Cauca (W of Cali 1,630 m)). Brazil. Refs.: Sasakawa, 1992b: 16 (as Calycomyza, new record Colombia); Martinez & Etienne, 2002: 30 (list). Genus Japanagromyza Sasakawa Japanagromyza Sasakawa, 1958: 138. Type species, Agromyza duchesneaei Sasakawa, 1954 (orig. des.). Refs.: Martinez & Etienne, 2002: 33 (list). Geratomyza Spencer (in Spencer & Stegmaier, 1973: 140). Type species: Geratomyza maculata Spencer (orig. des.) ambigua Sasakawa, 1992b: 5. Type locality: Colombia, La Luz, Candelaria, Cundinamarca. HT F (CAS). Distr.: Colombia (Cesar (La Luz (3 km N of Costilla), Valle del Cauca (Candelaria, Finca San Luis, 1,000 m.), Cundinamarca (Finca Bella Vista, nr. Sasaima)). Refs.: Martinez & Etienne, 2002: 33 (list). clausa Sasakawa, 1992b: 6. Type locality: Colombia, Candelaria. HT M (CAS). Distr.: Colombia (Valle del Cauca (Candelaria, Finca San Luis 1,000 m)). Refs.: Martinez & Etienne, 2002: 33 (list). desmodivora Spencer, 1966a.: 4. Type locality: USA, Florida, Miami. HT M (USNM). Distr.: Colombia (Bogotá (Road to Girardot), mine in Spencer, 1963 and adults in Spencer 1966a), Ecuador, Guadalupe, Peru, Venezuela, U.S.A (Florida). Refs.: Spencer, 1973c: 77; Martinez & Etienne, 2002: 33 (list); Etienne & Martinez, 2003: 254; Korytkowski, 2014: 6. phaseoli Spencer, 1983: 49. Type locality: Venezuela, Aragua, Rancho Grande. HT M (BMNH). Distr.: Colombia CATALOGUE OF AGROMYZIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 661 (? Spencer, 1984), Argentina, Costa Rica, Peru, Venezuela. Refs.: Spencer, 1984: 9; Spencer et al., 1992: 272; Martinez & Etienne, 2002: 34 (list). Genus Melanagromyza Hendel Melangromyza Hendel, 1920: 114, 126. Type species, Agromyza aeneoventris Fallén, 1823: 1 (orig. des.). Refs.: Spencer, 1966a: 3-60. Limnoagromyza Malloch, 1920: 147. Type species: diantherae Malloch. (orig. des.). caucensis Steyskal, 1972: 265. Type locality: Colombia, Pradera, Valle del Cauca. HT M (USNM). Distr.: Colombia, (Valle del Cauca (Pradera)). Refs.: Spencer, 1984: 6; Martinez & Etienne, 2002: 38 (list). colombiensis Spencer, 1963: 309. HT M (USNM). Type locality: Colombia, Bogotá. Distr.: Colombia (Bogotá). Refs.: Spencer, 1967: 83.3 (cat.); Martinez & Etienne, 2002: 38 (list). duplicata Sasakawa, 1992b: 1. Type locality: Colombia, Cundinamarca, Candelaria. HT M (CAS). Distr.: Colombia (Valle del Cauca (6 mil. W of Cali 1,630 m; Candelaria, Valle Finca San Luis 1,000 m, nr. [near] Candelaria, Narino (15 mil. S of La Union, 2,150 m), 11 mil N. of Popayan Cuaca, 1,830 m, Cundinamarca (10 mil. W of Villeta, 1, 660 m; Finca Bella Vista, nr. [near] Sasaima)). Refs.: Martinez & Etienne, 2002: 38 (list). lini Spencer, 1963: 315. Type locality: Peru, Barranca. HT M (USNM). Distr.: Colombia (Cundinamarca (Santafé de Bogotá, Jardín Botánico)), Argentina, Chile, Peru. Refs.: Spencer, 1967: 83.4 (cat.); Spencer, 1973c: 112; Sanabria de Arévalo, 1994: 65 (key); Sasakawa, 1992c: 352; Sasakawa, 1992d: 6 (n. syn.); Sasakawa, 1994: 19; Martinez & Etienne, 2002: 39 (list); Korytkowski, 2014: 14. Melanagromyza aguilerai Spencer, 1982: 37. mallochi Hendel, 1923: 145. (Agromyza; nom. nov. for eupatoriae Malloch, 1915: 107). Type locality: “Puerto Rico”. HT M (USNM). Distr.: Colombia (Valle del Cauca (Candelaria, Finca San Luis)), Cuba, Puerto Rico. Refs.: Spencer, 1963: 316; Spencer, 1967: 83.4 (cat.); Spencer & Stegmaier, 1973: 158; Sasakawa, 1992b: 3; Martinez & Etienne, 2002: 39 (list). Agromyza mallochi Hendel, 1923: 145 (nom. nov. for eupatoriae Malloch, preocc. by eupatotii Kaltenbach, 1874). Melanagromyza mallochi, Frick, 1952: 379. minima Malloch, 1913: 328. (Agromyza). Type locality: “Trinidad”. HT M (USNM). Distr.: Colombia (Valle del Cauca (6 mil. W Cali 1,630 m, Finca San Luis, nr. [near] Candelaria.)), British Guiana, Costa Rica, El Salvador, Guatemala, Jamaica, Mexico, Panama, Peru, Trinidad, U.S.A., Venezuela. Refs.: Spencer, 1963: 317; Spencer, 1967: 83.4 (cat.); Spencer, 1973b: 151; Spencer, 1983: 43 (key); Sasakawa, 1992b: 4 (first record from Colombia); Sasakawa, 1992d: 8; Spencer et al., 1992: 258; Martinez et al., 1993: 169; Martinez & Etienne, 2002: 39 (list); Sasakawa, 2005: 152; Korytkowski, 2014: 15. Melanagromyza minima, Shewell, 1953: 468; Frick, 1957b: 200, 1959: 364; Spencer, 1963: 317; Spencer, 1966a: 12 (n. syn.), HT M from Jamaica in author’s collection; Spencer & Stegmaier, 1973: 44. Melanagromyza longicaudalis Spencer, 1963: 316. neotropica Spencer, 1963: 319. Type locality: Mexico, Cuernavaca. HT M (USNM). Distr.: Colombia (Bogotá), Argentina, Brazil (Campinas), Chile, Costa Rica, Mexico, El Salvador, Ecuador, Peru, Venezuela. Refs.: Spencer, 1967: 83.4 (cat.); Spencer, 1973a: 22; Spencer, 1973b: 151; Spencer, 1982: 10 (key); Spencer, 1983: 42 (key); Spencer, 1984: 6; Sasakawa, 1992d: 8; Sasakawa, 1992c: 815; Sanabria de Arévalo, 1994: 65; Martinez & Etienne, 2002: 39 (list); Sasakawa, 2005: 152; Korytkowski, 2014: 15. phaseolivora Spencer, 1973c: 50. Type locality: Ecuador, Pichincha, Sta. Catalina Experiment Station. HT M (USNM). Distr.: Colombia, Ecuador, Peru. Refs.: Spencer, 1984: 6; Sasakawa, 1992d: 8; Martinez & Etienne, 2002: 39 (list). 662 · Zootaxa 4122 (1) © 2016 Magnolia Press DE SOUSA & COURI setifera Spencer, 1963: 322. Type locality: British Guiana, Mazaruni. HT M (BMNH). Distr.: Colombia (Antioquia (Medellín (Intercontinental hotel)), Guyana, Peru; Refs.: Spencer, 1984: 8 (Colombia); Martinez & Etienne, 2002: 40 (list); Korytkowski, 2014: 16. socolena Sanabria de Arévalo & Zenner de Polanía, 1994: 229. Type locality: Colombia, Anolaima, Cundinamarca. HT M (ICNDIPAg). Distr.: Colombia (Cundinamarca (Anolaima, Vereda Mesitas de Callero,Finca de La Capilla)). Refs.: Martinez & Etienne, 2002: 40 (list). spilanthis Spencer, 1984: 8. Type locality: Colombia, Bogotá. HT M (SC). Distr.: Colombia (Cundinamarca (Madrid, Jardines de los Andes; Santafé de Bogotá, Jardin Botanico; Ciudad Universitaria)). Refs.: Sanabria de Arévalo, 1994: 66 (key); Martinez & Etienne, 2002: 40 (list). tomaterae Steyskal, 1972: 266. Type locality: Colombia, Valle del Cauca. HT M (USNM). Distr.: Colombia (Antioquia (Medellín), Valle del Cauca)), Ecuador, Venezuela. Refs.: Spencer, 1973a: 24 (key); Martinez & Etienne, 2002: 40 (list). turbida Sasakawa, 1992d: 815. Type locality: Ecuador, Pichincha. HT M (CAS). Distr.: Colombia (Cundinamarca (nr. Sasaima, Finca Bella Vista)), Ecuador, Guatemala. Refs.: Martinez & Etienne, 2002: 40 (list); Sasakawa, 2005: 165. wedeliae Spencer (in Spencer & Stegmaier, 1973: 54). Type locality: U. S. A., Florida, Miami. HT M (USNM). Distr.: Colombia (Bogotá), U.S.A (Florida). Refs.: Spencer, 1984: 9 (Colombia, cit., key); Martinez & Etienne, 2002: 40 (list). Genus Phytobia Lioy Phytobia Lioy, 1864: 1313. Type species, Agromyza errans Meigen, 1830 (mon.). Refs.: Nowakowiski, 1962: 97; Spencer, 1966b: 142-143 (key to species), 1969: 101 Dizygomyza, subg. Dedromyza Hendel, 1931: 22. Type species, Agromyza carbonaria Zetterstedt, 1848 (orig. des.). Liomyzina Enderlein, 1936: 180. Nomen nudum (genus-group name proposed after 1930 without designation of type species from two included species; Evenhuis, 2008: 18). Shizukoa Sasakawa, 1963: 38. Type species: Shizukoa seticopia Sasakawa, 1963, orig. des. (syn. established by Spencer, 1965). setitibialis Sasakawa, 1992b: 8. Type locality: Colombia, Amazonas, Leticia. HT M (CAS). Distr.: Colombia (Amazonas (Leticia). Refs.: Martinez & Etienne, 2002: 43 (list). Subfamily Phytomyzinae Hendel, 1931 Genus Cerodontha Rondani Odontocera Macquart, 1835: 614. Type species, Chlorops denticornis Panzer, 1806 (orig. des.) (preocc. Serville, 1833). Cerodontha Rondani 1861: 10 (replacement name for Odontocera Macquart). Refs.: Nowakowski, 1962: 100, 1967: 633-661; Spencer, 1969: 109. Micromma Philippi, 1865: 777. Type species, ?Dolichopus flavifrons Phillippi. Refs.: Spencer, 1967: 83.6 (cat., n. syn.). Subgenus Butomomyza Nowakowski Butomomyza Nowakowski, 1967: 633. Type of subgenus, Agromyza angulata Loew, 1869. lunulata Sasakawa, 1992b: 11. Type locality: Colombia, Cundinamarca. HT M (CAS). Distr.: Colombia (Cundinamarca (Finca Bella Vista, nr. Sasaima)). Refs.: Martinez & Etienne, 2002: 31 (list). CATALOGUE OF AGROMYZIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 663 Subgenus Cerodontha Rondani Odontocera Macquart, 1835: 614 (as genus). Type species, Chlorops denticornis Panzer, 1806 (orig. des.). (Junior homonym, preocc. Odontocera, Serville, 1833) Cerodontha Rondani, 1861: 10. (replacement name for Odontocera Macquart). colombiensis Spencer, 1984: 11. Type locality: Colombia, Tabio. HT M (SC). Distr.: Colombia (Cundinamarca (Tabio, 35 km N of Bogotá; Mosquera, Subachoque, Zorro; Bogotá, Hotel Bogotá Plaza; La Calera, road side Bogotá)). Refs.: Sanabria de Arévalo, 1994: 73 (key); Martinez & Etienne, 2002: 31 (list). dorsalis Loew, 1863: 54. (Odontocera). Type locality: U.S.A., District of Columbia. HT M (MCZ). Distr.: Colombia (Cundinamarca (Sabana North of Bogotá, Funza, Madrid, Mosquera, Suba, Tabio, Subachoque, La Calera, Sapó)), Argentina, Bolivia, Brazil, Canada, Chile, Costa Rica, Cuba, Ecuador, Galapagos, Guadalupe, Guatemala, La Dominique, Palaearctic region, Paraguai, Peru, Puerto Rico, San Martin, U.S.A. (Florida), Venezuela. Refs.: Spencer, 1963: 331; Spencer, 1967: 83.6 (cat.); Spencer, 1969: 143; Spencer, 1973c: 293, Spencer, 1983: 53; Spencer, 1984: 11; Spencer et al., 1992: 274; Sasakawa, 1992c: 819; Sasakawa, 1992d: 27; Sanabria de Arévalo, 1994: 73 (key); Martinez & Etienne, 2002: 40 (list); Sasakawa, 2005: 160, Sasakawa, 2007: 195. Cerodontha dorsalis, Melander, 1913: 249; Frick, 1959: 396. Cerodontha (Cerodontha) dorsalis, Spencer, 1969: 143; Spencer & Stegmaier, 1973:70. Subgenus Dizigomyza Hendel Dizigomyza Hendel, 1920: 114, 130 (as genus). Type species, Agromyza morosa Meigen, 1830 (orig. des.). Dizygomyza (Dizygomyza), Hendel, 1931:83. Phytobia (Dizygomyza), Frick, 1952: 396; 1959: 383. Cerodontha (Dizygomyza), Nowakowski, 1962: 102. andensis Spencer, 1963: 334 (Phytobia). Type locality: Colombia, Tequendama. HT F (SC). Distr.: Colombia (Cundinamarca (Tequendama)). Refs.: Martinez & Etienne, 2002: 40 (list). Cerodontha (Dizigomyza) andensis, Spencer, 1966b: 142 (n. comb.), 1967: 83.6 (cat.). Subgenus Poemyza Hendel, 1931 Dizygomyza (Poemyza) Hendel, 1931: 35. Type of subgenus, Agromyza pygmaea Meigen, 1830. Phytobia (Poemyza), Frick, 1952: 391, 1959: 379. Refs.: Spencer, 1969: 127. Cerodontha (Poemya), Nowakowski, 1962: 102, 1967: 645. Refs.: Spencer, 1969: 127. nigra Spencer, 1984: 11 (Cerodontha). Type locality: Colombia, Bogotá. HT M (AC). Distr.: Colombia (Bogotá). Refs.: Martinez & Etienne, 2002: 32 (list). Cerodontha (Poemyza) nigra, Zlobin, 2000: 64 (n. comb.); Martinez & Etienne, 2002: 32 (list). Genus Chromatomyia Hardy Chromatomyia Hardy, 1849: 385. Type species, Phytomyza periclymeni Meijere, 1924 (as P. obscurella Fallén, 1823). syngenesiae Hardy, 1849: 385. (Phytomyza) Type locality: unknown. NT M (BMNH). Distr.: Colombia (Cundinamarca (Facatativá, Vda. Mana Blanca; Mosquera, Flores Targa, Finca Candilejas; Santafé de Bogotá, Jardín Botánico; Sabana, Montserrate)), Canada, U.S.A., California (Massachusetts, Oregon, Washington), Australia, Canada, Europe, Japan, New Zealand (introduced and widespread in temperate areas). Refs.: Spencer, 1973c (as Phytomyza); Spencer, 1984: 32 (Colombia); Spencer & Steyskal, 1986: 192 (key); Martinez & Etienne, 2002: 32. Agromyza atricornis Meigen, 1830 (pro parte) Ref.: Martinez & Etienne, 2002: 32 (list). Agromyza chrysanthemi Kowartz (in Lintner, 1891) Ref.: Martinez & Etienne, 2002: 32 (list). 664 · Zootaxa 4122 (1) © 2016 Magnolia Press DE SOUSA & COURI Genus Liriomyza Mik Liriomyza Mik, 1894: 289. Type species, Liriomyza urophorina Mik, 1894 (mon.). Agrophila Lioy, 1864 (3): 9. Type species, Agromyza exilis Meigen, Refs.: Martinez & Etienne, 2002: 34 (list). Antineura Melander, 1913: 219. Type species, Antineura togata (mon.). Refs.: Martinez & Etienne, 2002: 34 (list). Haplomyza Hendel, 1914: 73, nom. nov. for Antineura Melander, 1913: 219 [not Osten Sacken, 1881 a,b]. Refs.: Martinez & Etienne, 2002: 34 (list). Praspedomyza Hendel, 1931: 77 (subgenus Dizygomyza). Type species, Dizygomyza approximata. Refs.: Martinez & Etienne, 2002: 34 (list). Triticomyza Blanchard, 1938: 356. Type species, Triticomyza cruciata. Refs.: Martinez & Etienne, 2002: 34 (list). baccharidis Spencer, 1963: 354. Type locality: Colombia, Tequendama. HT M (SC). Distr.: Colombia (Cundinamarca (Tequendema, nr. Bogotá)), Chile, U.S.A., Venezuela. Refs.: Spencer, 1967: 83.9 (cat.); Spencer, 1973a: 54; Spencer, 1984: 15 (key); Spencer & Steyskal, 1986: 227; Sanabria de Arévalo, 1994: 76 (key); Sasakawa, 1994: 30; Martinez & Etienne, 2002: 34 (list). bogotensis Sanabria de Arévalo, 1993a: 266. Type locality: Colombia, Cundinamarca Colombia. HT M (ICNDIPAg). Distr.: Colombia (Cundinamarca (Santafé de Bogotá, 2,560 m alt.)). Refs.: Sanabria de Arévalo, 1994: 77 (key); Martinez & Etienne, 2002: 34 (list). brazilliensis Frost, 1939: 97 (Agromyza). Type locality: Brazil, São Paulo, Campinas. HT M (USNM). Distr.: Colombia (Nariño (Pasto (La Unión)), Argentina, Brazil, Ecuador (Baños), Perú, Venezuela. Refs.: Mendes, 1940, 78: 207-220; Spencer, 1963: 357 (Colombia), 1967: 83.10 (cat.), Spencer, 1973a: 55; Spencer, 1973c: 118; Spencer, 1984: 15 (key); Valladares, 1984: 20 (key); Sasakawa, 1992c: 820; Martinez & Etienne, 2002: 35 (list). Liriomyza braziliensis, Frick, 1952: 402; Spencer, 1963: 357. Agromyza ecuadorensis Frost, 1939, 50: 99. (n. syn.). Liriomyza ecuadorensis, Frick, 1952: 403. chiensis Spencer, 1984: 15. Type locality: Colombia, Chía. HT M (SC). Distr.: Colombia (Cundinamarca (Chía, n. of Bogotá)). Refs.: Sanabria de Arévalo, 1994: 79 (key); Martinez & Etienne, 2002: 35 (list). cirriformis Sanabria de Arévalo, 1993a: 268. Type locality: Colombia, Cundinamarca. HT M (ICNDIPAg). Distr.: Colombia (Cundinamarca (Santafé de Bogotá 2,560 m alt.)). Refs.: Sanabria de Arévalo, 1994: 78; Martinez & Etienne, 2002: 35 (list). colombiella Spencer, 1984: 15. Type locality: Colombia, Funza, Madrid, Tabio. HT M. (SC). Distr.: Colombia (Cundinamarca (Funza, n. of Bogotá; Madrid, n. of Bogotá; Tabio, n. of Bogotá; La Calera, Guasca, se. of Bogotá)). Refs.: Sanabria de Arévalo, 1994: 79 (key); Martinez & Etienne, 2002: 35 (list). commelinae Frost, 1931: 72. (Agromyza). Type locality: U.S.A., West Indies, St. Vicent. HT M (USNM). Distr.: Colombia (Cauca (Piendamó)), Argentina, Brazil, Barbade, Costa Rica, Cuba, Guadalupe, Jamaica, Peru, San Martin, Venezuela, U.S.A. (Florida). Refs.: Spencer, 1963: 361; Spencer, 1967: 83.10 (cat.), Spencer, 1973a: 56, Spencer & Stegmaier, 1973: 101 (key); Spencer, 1983: 55; Spencer, 1984: 15; Valladares, 1984: 24 (key); Spencer & Steyskal, 1986: 286; Spencer et al., 1992: 278; Martinez & Etienne, 2002: 35 (list); Korytkowski, 2014: 43. Liriomyza commelinae, Frick, 1952: 402, 1959: 403; Silva & Oliveira, 1952: 293; Spencer, 1963: 361; Spencer & Stegmaier, 1973: 101. Liriomyza bahamondesi Blanchard, 1954: 28. dianthicola Venturi, 1949: 161 (Pseudonapomyza). Type locality: Europe. HT ? (?). Distr.: Colombia (Bogotá), Palaearctic region. Refs.: Spencer, 1984: 16 (key); Martinez & Etienne, 2002: 35 (list). Phytomyza jannonei Séguy, 1950. Refs.: Martinez & Etienne, 2002: 35 (list). Liriomyza dianthicola, Von Tschirnhaus, 1981: 319 (n. comb.). Refs.: Martinez & Etienne, 2002: 35 (list). CATALOGUE OF AGROMYZIDAE OF COLOMBIA Zootaxa 4122 (1) © 2016 Magnolia Press · 665 herrerai Spencer, 1984: 18. Type locatily: Colombia, Madrid. HT M (SC). Distr.: Colombia (Cundinamarca (Madrid, N of Bogotá)). Refs.: Martinez & Etienne, 2002: 35 (list). hordei Spencer, 1984: 18. Type locality: Colombia, Obonuco. HT M (SC). Distr.: Colombia (Nariño (Obunuco, SW of Pasto)), Ecuador. Refs.: Sasakawa, 1992c: 820; Martinez & Etienne, 2002: 35 (list). huidobrensis Blanchard, 1926: 10. Type locality: Argentina, Buenos Aires. HT F (BC). Distr.: Colombia (Antioquia (La Ceja; Medellín), Cauca (Piendamó), Cundinamarca (Sabana, Subachoque; Madrid, Funza, Chía, Bogotá, Mosquera, Fusagasugá, Zorro)), Argentina, Brazil, Chile, Costa Rica, Ecuador, Guadalupe, Guatemala, Guyana, Martinica, Peru, Republica Dominicana, Uruguai, Venezuela, U.S.A., Afrotropical, Palaearctic and Australian regions. Refs.: Spencer, 1967: 83.10 (cat.); Spencer, 1973a: 57; Spencer 1983: 55; Spencer, 1984: 18 (Colombia), Valladares, 1984: 26 (key); Spencer & Steyskal, 1986: 8; Esposito et al., 1992: 247 (redescription); Spencer et al., 1992: 279; Martinez et al., 1993: 173; Sanabria de Arévalo, 1994: 80; Martinez & Etienne, 2002: 35 (list); Korytkowski:, 2014: 45. Liriomyza cucumifoliae Blanchard, 1938: 356-359 Refs.: Martinez & Etienne, 2002: 35 (list). Liriomyza decora Blanchard, 1954: 50 pp. Refs.: Martinez & Etienne, 2002: 35 (list). Liriomyza dianthi Frick, 1958: 1-5. Refs.: Martinez & Etienne, 2002: 35 (list). Liriomyza longei Frick, 1951: 81-88. Refs.: Martinez & Etienne, 2002: 35 (list). madridensis Spencer, 1984: 19. Type locality: Colombia, Madrid. HT M (SC). Distr.: Colombia (Cundinamarca (Madrid, N of Bogotá)). Refs.: Martinez & Etienne, 2002: 36 (list). marginalis Malloch, 1913: 283 (described as Agromyza melampyga var. marginalis). Type locality: Columbia, South Carolina. HT M (USNM). Distr.: Colombia (Cundinamarca (Finca Bella Vista, nr. Sasaima), Meta (10 mil. W of Villeta 1,660 m), Valle del Cauca (Buenaventura, 3 mil. W of Villavicencio, 920 m)), Bahamas, Brazil, Chile, Costa Rica, Cuba, El Salvador, Galápagos, Guadalupe, Jamaica, La Dominic, Nicaragua, Peru, Dominican Republic, Saint-Martin, U. S. A., West Indies, Venezuela. Refs.: Spencer, 1973a: 58; Spencer, 1973c: 260; Spencer & Stegmaier, 1973: 102 (key); Spencer, 1983: 58; Spencer, 1984: 20; Spencer et al., 1992: 279; Sasakawa, 1992b: 17; Sasakawa, 1992d: 20; Martinez & et al., 1993: 174; Martinez & Etienne, 2002: 36 (list); Sasakawa, 2005: 160; Sasakawa, 2007: 195. Liriomyza marginalis, Frick, 1952: 404, 1959: 407; Spencer, 1963: 370. Refs.: Spencer & Stegmaier, 1973: 102. mariaecamilae Sanabria de Arévalo, 1993a: 270. Type locality: Colombia, Cundinamarca. HT M (ICNDIPAg). Distr.: Colombia (Cundinamarca (Santafé de Bogotá, 2,560 m alt., Jardín Botânico)). Refs.: Sanabria de Arévalo, 1994: 83 (key).Martinez & Etienne, 2002: 36 (list). menthavora Sanabria de Arévalo, 1993a: 272. Type locality: Colombia, Cundinamarca. HT M (ICNDIPAg). Distr.: Colombia (Cundinamarca (Santafé de Bogotá, 2,560 m alt.)). Refs.: Sanabria de Arévalo, 1994: 83 (key); Martinez & Etienne, 2002: 36 (list). microglossae Spencer, 1963: 363. Type locality: Brazil, São Paulo. HT M (SC). Distr.: Colombia, Argentina, Brazil (São Paulo, Nova Teutonia), Costa Rica. Refs.: Spencer, 1967: 83.11 (cat.); Spencer, 1983: 58, Valladares, 1984: 28 (key); Martinez & Etienne, 2002: 36 (list); mosquerensis Sanabria de Arévalo, 1993a: 275. Type locality: Colombia, Cundinamarca. HT M (ICNDIPAg). Distr.: Colombia (Cundinamarca (Mosquera, 2,546 m alt., Lagunade la Herrera)). Refs.: Sanabria de Arévalo, 1994: 84 (key); Martinez & Etienne, 2002: 36 (list). nigra Spencer, 1984: 21. Type locality: Colombia, Guasca. HT M (SC). Distr.: Colombia (Cundinamarca (Guasca, SE of Bogotá; La Calera, SE of Bogotá; Suba (N of Bogotá)), Ecuador. Refs.: Sasakawa, 1992c: 820; Sanabria de Arévalo, 1994: 84 (key); Matinez & Etienne, 2002: 36 (list). 666 · Zootaxa 4122 (1) © 2016 Magnolia Press DE SOUSA & COURI quadrata Malloch, 1934: 471. (Agromyza) Type locality: Chile, Concepcion. HT M (BMNH). Distr.: Colombia (Bogotá, (Monserrate)), Argentina, Chile, Ecuador, Peru, Venezuela, Refs.: Spencer, 1963: 365; Spencer, 1967: 83.11 (cat.), Spencer, 1973a: 59; Spencer, 1973c: 176; Spencer, 1984: 21; Sasakawa, 1992b: 21; Sasakawa, 1992d: 17; Sasakawa, 1992c: 821; Sasakawa, 1994: 34; Sanabria de Arévalo, 1994: 85 (key); Martinez & Etienne, 2002: 36; Korytkowski, 2014: 51. Liriomyza subandina Blanchard, 1954: 34 Liriomyza quadrata, Frick, 1952: 405 (n. comb.) robustae Spencer, 1984: 22. Type locality: Colombia, Cuaca. HT M (SC). Distr.: Colombia (Cauca (Piendamó)). Refs.: Martinez & Etienne, 2002: 36 (list). sabanae Spencer, 1984: 30 (Phytoliriomyza). Type locality: Colombia, Mosquera. HT M (SC). Distr.: Colombia (Cundinamarca (Mosquera, NW of Sabana de Bogotá)). Refs.: Martinez & Etienne, 2002: 36 (list). Liriomyza sabanae, Martinez & Etienne, 2002: 36 (n. comb.) sabaziae Spencer, 1963: 366. Type locality: Venezuela, Caracas. HT M (SC). Distr.: Colombia (Cundinamarca (Bogotá, Sabana), Antioquia (La Ceja) Cauca (Piendamó)), Costa Rica, Ecuador, Peru, U.S.A., Venezuela, Refs.: Spencer, 1967: 83.11 (cat.), Spencer, 1973a: 60; Spencer, 1983: 58; Spencer, 1984: 23; Spencer & Steyskal, 1986: 228; Sasakawa, 1992c: 821 Sanabria de Arévalo, 1994: 86 (key); Martinez & Etienne, 2002: 36 (list); Korytkowski, 2014: 53. santafecina Sanabria de Arévalo, 1993a: 276. Type locality: Colombia, Cundinamarca. HT M (ICNDIPAg). Distr.: Colombia (Cundinamarca (Santafé de Bogotá, 2,640 m alt.)). Refs.: Sanabria de Arévalo, 1994: 86 (key); Martinez & Etienne, 2002: 36 (list). sativae Blanchard, 1938: 354. Type locality: Argentina, La pampa. HT F (BC). Distr.: Colombia (Cundinamarca (10 mil. W of Villeta 1,660 m)), Argentina, Bahamas, Brazil, Canada, Chile, Costa Rica, Cuba, Guadalupe, Haiti, Jamaica, Martinique, Mexico, Nicaragua, Peru, Puerto Rico, Republica Dominicana, Trinidad, U.S.A. (Floride), Venezuela, Afrotropical, Australian, Palaearctic and Oriental regions (introduced). Refs.: Spencer, 1963: 372; Spencer, 1967: 83.11 (cat.); Spencer, 1973a: 61; Spencer, 1983: 59; Valladares, 1984: 31(key); Sasakawa, 1992b: 17; Spencer et al., 1992: 280; Martinez & et al., 1993: 174; Martinez & Etienne, 2002: 36 (list); Korytkowski, 2014: 54. Lemurimyza lycopersicae Pla & La Cruz, 1981. Refs.: Martinez & Etienne, 2002: 36 (list). Liriomyza canomarginis Frick, 1952: 395. Refs.: Martinez & Etienne, 2002: 36 (list). Liriomyza guytona Freeman, 1958: 344-345. Refs.: Martinez & Etienne, 2002: 36 (list). Liriomyza minutiseta Frick, 1952: 395. Refs.: Martinez & Etienne, 2002: 36 (list). Liriomyza munda Frick, 1957a: 59-70. Refs.: Martinez & Etienne, 2002: 37 (list). Liriomyza propepusilla Frost, 1954: 73. Refs.: Martinez & Etienne, 2002: 37 (list). Liriomyza pullata Frick, 1952: 339-452. Refs.: Martinez & Etienne, 2002: 37 (list). Liriomyza verbenicola Hering, 1951: 420pp. Ref.: Martinez & Etienne, 2002: 37 (list). solanita Spencer, 1963: 369. Type locality: Colombia, Bogotá. HT M (SC). Distr.: Colombia (B

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