zyxwvutsrqp zyxw zy Type studies in the Polyporaceae 11. Species described by J. F. C. Montagne, either alone or with other authors Leif Ryvarden Ryvarden, L. 1982. Type studies in the Polyporaceae 11. Species described by J. F. C. Montagne, either alone or with other authors. - Nord. J. Bot. 2: 75-84. Copenhagen. ISSN 0107-055X. zyxwvut zyxwv Of the 93 polypores described by Montagne, either alone or with other mycologists, 32 are accepted, 53 are synonyms, 3 names are invalid, 2 species are of unknown affinities, 2 types have not been found while 1 name is rejected. The following three combinations are proposed: Phylloporia capucina (Mont.) Ryv. comb. nov., Coltricia fragilissima (Mont.) Ryv. comb. nov., and Gloeophyllum mexicanurn (Mont.) Ryv. comb. nov. L . Ryvarden, Botanical Laboratory, Univ. of Oslo, P.O. Box 1045, Blindern, N-Oslo 3, Norway. cations in which he described polypores, is given in the references. Except for the name, which is obvious from the The French botanist J. F. C. Montagne described a number of fungi, either alone, or with C. G. D. Nees,- heading, the text of the label is cited in inverted commas M. J. Berkeley and M. C. Durieu as co-authors. Of to ease the tracing of the examined lectotype. Often these, 93 were polypores, and are treated here. Mon- there are several specimens with the same name on the tagne's herbarium is kept separate in the Paris Museum same sheet, and this may later lead to difficulties, as (P), and this certainly accounts for the fact that only 2 of holotypes, lectotypes etc. are not marked in the Pans his types could not be located. He exchanged specimens Herbarium. If the type locality is not evident from the with many of his contemporaries such as M. J. BerkeIey label, it is given in brackets. When the name of the type was found to be a and E. Fries and thus, isotypes may also be found in Kew (K), Stockholm (S) and Uppsala (UPS). Further- taxonomic synonym of an earlier described species, this more, N. T. Patouillard took samples from the Mon- is marked with =. When the name of the type was found tagne collection, probably for comparison, and these to be a nomenclatoric synonym of another species, this samples are today in the Farlow Herbarium, Harvard is marked with =. Further, there is a reference to a (FH). In the following, no designation of herbarium recent description, or, if no modem description seems means that the type is in the Pans Herbarium. It is also to exist, a detailed description is given. A fact that has caused some confusion from time to indicated whether isotypes have been found in other time is that a number of species that Montagne deherbaria. The species are arranged alphabetically according to scribed in La Sagra (1842) were marked with nov. sp. specific epithet. The generic names are abbreviated as However, all these species had been validly published follows: D = Daedalea, F = Favolus, G = Gloeoporus, earlier, although some of them as late as February 1842. H = Hexagonia, L = Lenzites, P = Polyporus and T = This date was earlier than that of the publication of La Trametes. After the name there is a reference to where Sagra's book, as already pointed out by Montagne it was published. A complete list of Montagne's publi- himself (1842a: 119). Introduction zyxwvutsr zyxw 8 NORDIC JOURNAL OF BOTANY NORD.1. BOT. 2: 75-84, MYCOL 009 75 zyxwvutsrq zyxwvut zyxwvutsrqp zyxwvutsrq P. actinoboius Mont. P. anisoporus Delast. & Mont. in Mont. 1854: 129 “Crypt. Guyan. (French Guiana) no. 347. Leprieur no. 856”. = Rigidoporus ulmarius (Fr.) Imaz. 1952: 119. 1845b: 357 “Laudum (France), M. Delastre”. = Polyporus arcularius Fr. 1821: 342. For a discussion and a description of this species, see Jahn (1980: 162). P. actinopilus Mont. 1857: 369 “Brasil, Weddel”. = Cerrena sclerodepsis (Berk.) Ryv. 1976: 204. P. auberianus Mont. P. aculeatus Mont. 1840: 205 “In sylvino, Cayenne (French Guiana), in ligno putris. Leprieur”. = Hexagonia tenuis (Hook.)Fries 1838: 1842a: 127 “Hist. nat. Cuba. cum icone. Surinam. Miquel no. 1127”. = Rigidoporus microporus (Fr.) Overeem 1924: 1. 498. P. auriformis Mont. T. aethalodes Mont. 1854: 128 (isotype in K) “Crypt. Guyan. (French Guiana) no. 359. Leprieur coll. no. 966”. = Rigidoporus microporus (Fr.) Overeem 1924: 1. zyxwvutsr zyxwvu 1857: 370 “Int. Brasil, coll. Weddell”. aethalodes (Mont.) Ryv. 1974: 276. = Coriolopsis P. blanchettianus Berk. & Mont. Fruitbody dimidiate, pileate, semicircular and appla- 1849: 238 “Bahia, Blanchet”. nate, 4-8 cm long, up to 6 crn wide and 1 cm thick, coriaceous to tough. Pileus azonate, velutinate to to- This is an accepted species in Polyporus s. str. For a mentose, dark umber to sienna brown. Pore surface description, see Reid (1976: 194) or Ryvarden & umber to deep rusty brown, pores angular, thinwalled Johansen (1980: 487). 2-3 per mm, a few wider and somewhat split radially, tubes up to 8 mm deep, concolorous with the pore surface. Context deep umber brown, 1.5 cm thick at the base. Hyphal system trimitic, generative hyphae P. brachyporus Mont. 1854: 131 “Crypt. Gyuan. (French Guiana) no. 371. Leprieur hyaline, thinwalled and with clamps at the septa, 2-3 p wide, skeletal hyphae dominating in the fruitbody, coll. no. 959”. = Echinochaete bruchyporus (Mont.) Ryv. 3-6 p wide, pale golden to rusty brown, solid to very 1978: 101. thickwalled, binding hyphae rare, tortuous, 3 4 . 5 pm For a description, see Ryvarden & Johansen (1980: wide. Catahymenium present as skeletal hyphae pro- 317). jecting into the hymenium as rounded to acute hyphal endings, smooth to finely encrusted, reminding of cystidia. Spores cylindrical, hyaline, non-amyloid, 6-7 x L. brasiliensis Mod. 2-3 pn, only seen floating. Habitat: On dead wood. 1857: 366 “Brasilia, coll. Weddel”. = Lenzites elegans (Fr.) Pat. 1900: 89. The species is undoubtedly related to C. polyzona (Pers.) Ryv., but is distinguished by a much darker bpsinus brown colour, an azonate and soft pileus and microstopically by the distinct catahymenium with its cystidia- 1842a: 126 (isotype in K) “Cuba, coll. R.de la Sagra”. like hyphal endings. It may be that fresh specimens are more ligthcoloured than described for the type. Spe&- This is an accepted Species in COriOlOpSh, for a descripmens of many trimitic species have a tendency to darken tion, see RYarden lk JOhansen (1980: 286). after prolonged storing in the herbarium. P. camwloDorus Mont. --‘ 76 “Zrypt. Guyan. (French Guiana) B38”. = Cvclomyces fuscus Fr. zyxwvutsrqp zyxwvutsrq zyxwvutsr zyxwvutsrq P. captiosus Mont. 1847: 170 “From Bonae spei. (Cape of Good Hope, South Africa). Drege no. 33”. = Coriolopsis floccosu (Jungh.) Ryv. 1972a: 230. As indicated by Reid (1975: 86) Montagne’s specimen is a bit untypical as binding hyphae are absent. However, this type of hyphae develops rather late, and the present type is a small and young specimen. P. capucinus Mont. 1857: 369 (isotype in FH) “Brazil”. Mont.) Ryv. (see below). That H. cruenta was the same as T. scabrosa was a possibility pointed out also by Montagne himself (op. cit.). P. cruentatus Mont. 1854: 129 “Guyana. Leprieur no. 1065”. = Caloporus spissw (Schw. in Fr.) Ryv. 1973: 9. zyxwvuts 3 Phylloporiu cupucinu This is a Phylloporia species and probably what Murrill (1904a: 141) described as P. parasitica, the type of which was found resupinate on a living leaf. Microscopically there is no difference between the types of P. capucinus and P. parasitica and I believe that the latter represents a tiny specimen of the former. All species in Phylloporia are known for their ability to parasitize living plants and the fruitbodies are mostly found around small stems and branches of living bushes and trees. P. capucinus is related to P . chrysita, and macroscopically the two species are very similar. Microscopically they are separated by the spores, which are larger and more ellipsoid in P. capucinus than in P. chrysista which has more or less globose spores. For a complete treatment of Phylloporia, see Ryvarden & Johansen (1980: 230-235). The following combination is proposed: For a detailed description, see Lowe (1966: 37). P. crypthacanthus Mont. 1857: 369 “Brasilia, Weddel”. = Phellinus contiguus (Fr.) Pat. 1900: 97. P. cubensis Mont. 1837: 364 “Cuba, B. M. la Sagra”. (Mont.) Pat. & Lagerh. 1891: 161. = Trumetes cubensis For a description, see Overholts (1953: 144). F. cucullatus Mont. 1842: 125 (isotype in K) “Cuba, M. Maurca”. Pseudofuvolus cucullatus (Mont.) Pat. 1900: 81. = For a description, see Ryvarden & Johansen (1980: 5 14). Phylloporia capucina (Mont.) Ryv. comb. nov. P. cycliscus Mont. Basionym: Polyporus cupucinus Mont., Ann. Sci. Nat. SCr. 4. Vol. 5: 369, 1857. 1850: 363 “Chile, Bertero”. = P. guianensis Mont. 1840: 201. F. ciliaris Mont. P. dictyopus Moat. 1843: 364 (isotype in K ) “Madagascar, Dedit. Gaudot”. arcularius Fr. I82 1: 342. = P, 1835: 345 (isotype in K) “Juan Fernandez”. This species belongs in Polyporus s. str. For a description, see Ryvarden & Johansen (1980: 491). P. concrescens Mont. 1835: 350 (isotypes in K, FH) “Ins. Juan Fernandez. Leg. Bertero”. = Rigidoporous microporus (Fr.) Overeem 1924: 1. D. dregeana Mont. G . conchoides Mont. 1842a: 126 “Cuba, Sagra”. = Gloeoporus thelephoroides (Hook.) Cunningh. 1965: 111. 1847: 171 (isotype in S) “No locality on the label, only the name written in Montagne’s hand. The collection came from Port Natal, South Africa. = Cerrenu muyenii (Kl.) Hansen (1960: 129) as indicated already by Reid (1975: 31). H. cruenta Mont. P. extenuatus DR. & Mont. in Mont. 1845a: 208 “Va vao, (Tonga, Pacific Ocean) Solefir”. = Trumetes scubrosu (Pers.) Cunn. (1965: 162). 1856: 166 “La Calle (Algerie)”. = Coriolopsis gulficu (Fr.) Ryv. 1972a: 230. 77 zy zyxwvutsrq zyxwvutsrqp zyxwvut zyxwvutsrqp P. fernandesianus Mont. P. guianensis Mont. 1835: 350 (isotype in K) “Polyporus dictyopus Mont. nunc P. fernandesi Mont. Is. Fernandez collecti Berterii”. Unknown hand. “Vix bona species, probabiliter primordia Polyporus concrescens Mont., cajus habet structuram”. In Bresadola’s hand. = Rigidoporus microporus (Fr.) Overeem 1924: 1. 1840: 201 (isotype in K) “Guiana. Leprieur”. P. flabellurn Mont. 1842a: 126 (isotype in K) “Crypt. Cuba c. icone, Pol. Membranaceus Fr., com. de la Sagra”. = Trametes membranaceus (Fr.) Kreisel 1971: 83. This is a neotropical species in Polyporus s. str. For a description, see Fidalgo (1967: 863) and Ryvarden & Johansen (1980: 496). T. guyoniana Mont. 1857: 333 “Algerie. M. Guyoy”. = Daedalea quercina Fr. 1821: 333. The type is badly eaten by insects and is sterile. The hyphal system, the consistency and colour of the context and the original description clearly point towards Ganoderma colosswn (Fr.) Baker 1918: 425. A badly developed and partly poroid specimen. Bresadola (1920: 69) believed it to be Hexagonia nitida f. trametoidea. Today only a tiny slice is left of the type, and definitive decision on its identity is difficult to ascertain. However, the pores are about 1 mm wide and somewhat sinuous, and the pileus is finely pubescent at the margin, which indicates a poroid form of D. quercina more than H. nitida, which is smooth and glabrous on the pileus. P. fragiliiimus Mont. P. heteroporus Mont. P. flavescens Mont. zyxwvutsrq zyxw 1857: 368 “P. officinah Afzelius pOsteatOS0, COlOSSO etc. affine. Brasilia Campos. Coll. Weddell.”. I 1854: 130 “Guyana in carbonarius. Leprieur”. fragilissima (Mont.) Ryv. (see below). = Coltricia 1841: 273 “Galgea (near Mauritius) M. de Duc”. = Coriolopsis asper (Jungh.) Teng 1964: 759. This species was later described by Wakefield from burnt wood in Nigeria as Polyporus pyrophilus Wakef. P. holophaeus Mont. 1916: 71. It belongs in Colrricia and the name of Mon- 1843: 361 (isotype in K) ,,No. 88, nov. sp, Mount Marsan, tagne has Priority. For full synonymy and a description, Laudon (France), in pieneci ad terram”. = Phaeolus schweinitsee Ryvarden & Johansen (1980: 111). The following zii (Fr.) Pat. 1900: 86. combination is proposed: The synonymy was noted already by Lloyd (1910: 68). Coltricia fragillissima (Mont.) Ryv. comb. nov. Basionym: Polyporus fragilissimus Mont., Ann. Sci. Nat. S6r. 4. Vol. 1: 130, 1854. - Syn. P. pyrophilus Wakef., Kew Bull. 1916: 71. P. galegensis Mont. 1841: 274 “Galega prope Mauritius”. This is a Ganoderma species, but unfortunately the type is sterile. However, the hyphal elements and the colour and the structure of the pileus point towards Ganoderma amboinense (Fr.) Pat. 1887: 171 as a possible species. For a detailed description of this species, see Steyaert (1972: 88). P. hornodermus Mont. 1857: 368 (lectotype in BPI) “Type, c. Bresadola”. = Perenniporia martius (Berk.) Ryv. 1972b: 143. P. iodinus Mont. 1841: 108 (isotype in K) “Ligno putres in Sylvis (French Guiana). Leprieur no. 532”. This is an accepted species in Cyclomyces and should be cited as C. iodinus (Mont.) Pat. 1900: 98. It is only known from the neotropics. P. inamaensis Mont. = Phel- F. giganteus Mont. 1842b: 22 “Nellgherris (India) coll. M. Perrottett”. linus inamaensis (Mont.) Ryv. 1972a: 234. 1854: 135 “Crypt. Guyan. (French Guinana) no. 391 bis. Leprieur coll. no. 958”. = Favolus brasilensis (Fries) Fries 1828: 44, as indicated already by Bresadola (1916: 230). For a description, see Ryvarden KL Johansen (1980: 175). 78 zyxwvutsrqp zyxwvuts zyxwvutsrqp zyxwvutsrq P. laetemarginatus DR. 8s Mont. in Mont. 1856: 163 “Alger”. = Oxyporus laetemarginafus (DR. & Mont.) Donk 1966: 342. For a description, see Lowe (1966: 17) or Ryvarden & Johansen (1980: 452). P. leiodermus Mont. 1854: 134 (Cisotypes in S, FH) “Guyana (French Guiana). Leprieur coll. 855). = Trametes modesta (Fr.) Ryv. 1972a: 236. For a description, see Ryvarden & Johansen (1980: 444). p. melanopus Mont. 1854: 136 “Guyana (French Guiana). Leprieur no. 942”. = . Polyporus guyanensis .Mont. 1840: 201. p, menadianus Mont. 1843: 362 “P. pergamenus Fr. P. Friesii affine. M. Menand. New York (USA)”. = Trichaprurn biformis (Fr.) Ryv. of which both the species mentioned on the label are synonyms. P. leprieurii Mont. 1840: 203 (isotype in K ) “2 Cent. Cum. icone. Guiana (French Guiana). Leprieur”. L. mexicana Mont. This species belongs in Polyporus s. str. For a description, see Ryvarden & Johansen (1980: 501) or Fidalgo & Fidalgo (1967: 859). 1843: 360 “Daedalea striafa Sw. affinis an diversa L. mexicanus Mont. Mexique, Prov. Oaxaca itin Andreu”. E Gloeophyllum mexicanum (Mont.) Ryv. (see below). P. licnoides Mont. This is an accepted species in Gloeophyllum and has previously been known as Gloeophyllum berkleyii (Sacc.) Murr. of which there is a detailed description in Fidalgo & Fidalgo (1966: 871-874). Montagne’s name has priority over that of Saccardo, and the following combination is proposed: zyxwvut zyxw 1840: 204 “ad ramos. (French Guiana). Leprieur no. 685”. = Phellinus gilvus (Schw.) Pat. 1900: 97. P. leucophaeus Mont. Gloeophyllum mexicanum (Mont.) Ryv. comb. nov. 1856: 157 “Ohio, Sullivant”. = Ganoderma apphnorum (Wallr.) Pat. 1889: 67. Basionym: Lenzites mexican@ Mont., Ann. Sci. Nat. S&. 2. Vol. 20: 360, 1843. - Syn. Daedalea rhabarbarina Berk. & Cooke in Cooke 1878: 130, (nom. illeg.) non D. rhabarbarina Mont. 1840: 205. - D . berkleyii Sacc. 1883: 381. Gloeophyllum berkleyii (Sacc.) Murr. 1905 : 370. T. marchionica Mont. 1845a: 204 (isotype in UPS) “Voy. Solifer. Nahiva (Pacific Ocean). M. Hombron”. = Trarnetes feei (Fr.) Pat. 1900: 92. p. micromegas Mont. zyx zy 1842a: 128 (isotype in K ) “Cuba, ad Ramon de la Sagra”. = Rigidoporus microporus (Fr.) Overeem 1924: 1. I. maximus Mont. 1837: 364 “Polyporo labyrinthico proximo. Cuba. Com. R. de la Sagra”. = Cerrena maxima (Mont.) Hansen 1960: 129. p, ,,,iquefi Mont. 1845b: 357 “In trunco arbori, Surinam. CI. Miquel”. 3 For a description, see Ryvarden & Johansen (1980: 517). P. megaloporus Mont. 1854: 124 “Leprieur no. 958 (French Guiana)”. Echinochaete brachyporus (Mont.) Ryv. (see above). The name is illegitimate being a homonym of P . megaloporus Pers. 1825: 88. P. monochrous Mont. 1841: 109 “Guyana (French Guiana). Leprieur no. 596”. = Tramefes modesrus (Fr.) Ryv. 1972a: 236. Synonymy already indicated by Bresadola (1916: 226). P. melanoporus Mont. 1842b: 127 “Cuba”. Murr. 1904a: 425. = Nigroformes melanoporus (Mont.) 79 zyxwvutsrq zyxwvutsrqpon zyxwvuts zyxwvutsr zyxwv F. moluccensis Mont. 1843: 365 “Ins. Ternate (Indonesia)” = Favolus spathulatus (Jungh.) LCv. 1844: 203. P. multiformis Mont. 1854: 125 (isotype in K) “Crypt. Guian. no. 844. Coll. Leprieur 939”. = Colrricia spathulafa (Hook.)Murr. 1908: 93. P. multiplicatus Mont. 1854: 128 (isotype in K ) “Guiana no. 857”. multiplicata (Mont.) Pat. 1889: 74. shiny, probably up to 10 cm long. Pore surface pale brown, pores thinwalled, angular, 2-3 per mm, tubes concolorous, up to 13 mm deep. Context very thin, brown 1-2 mm thick. Hyphal system trimitic, generative hyphae 3-4 prn wide, hyaline and with clamps, skeletal hyphae hyaline to pale yellow, swelling in KOH and then up to 12 pm wide, binding hyphae probably present, but the strongly branched segments seen may also represent the upper part of arboriform skeletal hyphae. Spores truncate, pale yellow and finely verruculose, 28-32 X 18-21 prn. The spores are probably the largest known among poroid fungi. = Ganoderrna A detailed description with synonyms can be found in Steyaert (1980: 150). 1856: 171 “Columbus (Ohio, USA) Sullivant”. = Polyporus mori Fr. 1821: 344. Often called Favolus alveolaris in American literature. The name is not mentioned in Overholts (1953). P. nilgheriensis Mont. 1842b: 22 (isotype in K ) “Nellgherris (India)”. nilgheriensis (Mont.) Cunn. 1965: 226. F. ohiensis Mont. = Phellinus For a description, see Ryvarden & Johansen (1980: 187). P. omalopilus Mont. 1842a: 128 “Crypt. Guyan. (French Guiana) no. 36. Leprieur coll. 539”. = Phellinus gilvus (Schw.) Pat. 1900: 97. P. nanus DR. & Mont. in Mont. 1856: 153 “Mascara (Algerie)”. = P. arcularius Fr. 1821: 342. P. peguanus Mont. 1834: 148. Type not found. H. nitida DR. & Mont. in Mont. 1856: 170 “In albarum ad trunc. Quercus. Julian (Algerie)”. P. phaeoporus Mont. This is an accepted species in Hexagonia. For a description, see Ryvarden (1976: 205). The species is restricted to the Quercus-zone around the Mediterranean. Donk (1969: 666) made it type species of Apoxona. 1854: 133 “Crypt. Guyan. (French Guiana) no. 370. Leprieur coll. 920”. = Coriolopsis aethalodes (Mont.) Ryv. 1974: 276. P. ochrolaccatus Mont. The type is small and probably immature. Its surface is almost glabrous, but otherwise it has the colour, the pores and the hyphal system of C. aethafades. Especially distinct is the catahymenium with swollen ends of protruding skeletal hyphae. The type is sterile. 1842a: 241 (isotypes in K, FH) “Manille (Philipine Id.) Cuming no. 1979”. = Ganoderma ochrolaccatus (Mont.) Pat. 1889: 68. This species is highly characteristic in Ganoderma because of its very large spores. Fruitbody laterally stipitate, pileus ungulate with almost vertical margin and a horizontal top, more or less circular, but slightly depressed around the attachment of the stipe, about 3 cm in diameter, 15 rnm thick, surface laccate, shiny, glabrous, distinctly sulcate along the vertical margin, wrinkled on the top (probably smooth when fresh). Yellow to ochraceous. Stipe only present as fragments today, about 5-6 mm in diameter, concolorous with the pileus or slightly darker, laccate and 80 P. phaeocanthus Berk. & Mont. in Mont. 1856: 154 “Columbus (Ohio, USA) Sullivant”. Fruitbody stipitate, pileus about 2 cm in diameter, apparently shrunken during drying, dark brown, glabrous, wrinkled. A thin agglutinated crust is present. Stipe 1.5 mm in diameter, 2-2.5 crn long, dark brown, longitudinally wrinkled. Pore surface reddish brown, pores round to angular, small, 8-1 1 per mm, tubes and context 1-3 mm thick, dark brown and dense in structure. Hyphal system monomitic, generative hyphae thinwalled, simple septate and frequently branched, 2 4 . 5 pm in diameter. Cystidia not seen. Basidia clavate with 4 zyxwv zyxwvutsrqp zyxwvu zyxwvut zyxwvutsr zyxwvutsrq zyx sterigmata, up to 35 pm long and 5-8 pm wide. Spores oblong ellipsoid, smooth, slightly thickwalled and pale golden brown (in 2.5% KOH), non-amyloid. 9-11.5 X 4.5 p.m. P. persoonii Mont. 1834: 147 “En il. Cl. Belanger Indis”. = Trametes scabrosa This is a species unknown to me. It is not mentioned in Overholts (1953). The structure and consistency of the fruitbody seem to indicate that this is no true polypore, but I do not know any poroid agaric corresponding to the description from the United States. Mycologists collecting in Ohio should try to find it again. D. rhabarbarina Mont. (Mont.) Cunningh. 1111: 111. 1840: 205 “Ad ramos vivat abnorm.. . (illegible) in sylvis Cayenne & Guyane (French Guiana), Leprieur no. 141”. = Seprobasidium rhabarbarinum (Mont.) Bres. 1916: 240. This is an accepted species in the heterobasidiomycete genus Septobasidium. For a description, see Couch (1938: 122). P. pyrrhoporus DR. & Mont. in Mont. 1856: 162 (isotype in FH), nom. rejec. “ P . vaporarius affine. Oran (Algerie). Surface inferieure d’un fragment de bois tombe dans un lieu humide. Mars 1842”. The type collection is mixed. Most of the fragments are of a monomitic species with clamped generative hyphae and abundant ellipsoid, hyaline, smooth, thinwalled spores 4-5 X 2.5-3 pm. Its pores are irregular, thinwalled and angular, 3-4 per mm, and the porewalls are dense and very brittle. No chlamydospores could be seen. This part of the collection may be Tyromyces rennyi (Berk. & Br.) Ryv. 1974: 281, which has the same spores. However, this species often has chlamydospores, but there are identical specimens without chlamydospores. Lowe (1966: 68) gives P. pyrrhoporus as a synonym of T. aneirinus, which however, usually has somewhat larger spores than those cited above. However, in the type collection of P. pyrrhoporus there are also fragments of a more small-pored resupinate species. This has a dimitic structure with skeletal hyphae. Generative hyphae were not found in these fragments. Only few spores were seen, they were of the type mentioned above and probably represent a contamination. In the original description it is stated: “Pori quoad magnitudinem et fornam valde variabilis, rotundi scilicet, angulati, laceratione dissepirnentorum elongati aut flexuosi”. Thus, it is apparent that both elements have been involved and the name has to be rejected (cf. International Code of Botanical Nomenclature, Art. 70). P. polygammus Mont. 1837: 365 (isotype in K, FH) “Ins. Cuba. Coll. Sagreana”. = Hexagonia tenuis (Hook.) Fr. 1838: 498. P. psilodermus Berk. & Mont. 1849’ 239 “Bahia (Brazil) (Blanchet) Hb. Bory”. = Trametes marianna (Pers.) Ryv. 1972a: 236. 6 Nord. J. Bot. 2(1) 1982 R. rhizomorphus Mont. 1840: 202 (isotype in K, FH) “Ad terram in sylvis Guyane et Cayenne (French Guiana), Leprieur”. This is a Polyporus s. str. It is characterized by having an entangled mass of black, cylindrical rhizomorphs from which the fruitbodies arise. Macroscopically and microscopically it is very similar to P. blanchertianus Berk. & Mont. Sharing the same type of widened binding hyphae with rather short sidebranches. Fertile specimens are desirable to check whether there are any differences in spore shspe and size. It may be that P. rhizomorphus only represents a form of P. blanchettianus. H. rhombipora Mont. 1857: 370 “Campos Prov. Minas Geras (Brazil). Coll. E. Weddell”. = Favolus brmiliensis (Fr.) Fr. 1828: 44. P. rhytiphloeus Mont. 1857: 369 “Brasilia, Weddell”. = Phellinus rhytiphloeus (Mont.) Ryv. in Ryv. & Johan. 1980: 206. For a description, see Ryvarden & Johansen (op. cit.). P. sagraeanus Mont. 1842a: 127 “Cuba” (also written on the specimen). = Tramefesfeei (Fr.) Ryv. 1972a: 236. As already indicated by Fidalgo (1968: 203), Montagne himself confused this species with Trametes cupreo-rosa (Berk.) Lloyd from South America. Leprieur coll, nos 861 and 865 in the Montagne collection are this species. The two species are sharing the same pink to reddish brown colour, but T. cupreo-rosea has larger pores and a more fibrillose and zonate pileus. 81 zyxwvutsrqp zyxwvutsrq zyxwvut zyxwvuts P. tabacinus Mont. P. senex Nees & Mont. Pheflinus senex 1835: 349 (isotype in K) “In ramos jectum, Is. Fernandez, Bertero”. sz Cycfomyces tabacinus (Mont.) Pat. 1900: 98. For a description, see Ryvarden & Johansen (1980: 2 15). For a description, see Ryvarden & Johansen (1980: 122). P. sordulentus Mont. P. tephromelas Mont. 1850: 357 “Chile”. = Crifola sordulenta (Mont.) Sing. 1962: 300. 1840: 203 “Ramis putri, ad terram, Cayenne (French Guiana)”. = Polyporus leprieurii Mont. 1840: 203. This is a stipitate species that belongs in Grifolu and there is a good description in Singer (1969: 382). P. tephroporus Mont. P. splitgerberi Mont. 1845b: 358 “P. cinerascens Schw., P. surinamensb Mont. non Miq. Surinam, Splitgerber no. 1285”. = Loweporus tephroporus (Mont.) Ryv. in Ryv. & Johan. 1980: 416. 1836: 70 “Juan de Fernandez, Bertero”. (Nees & Mont.) Imaz. 1952: 115. 1841: 109 “Emdio pores, Surinam”. (Mont.) Ryv. 1974: 274. E = Znonotus splitgerberi For a description, see Ryvarden & Johansen (1980: 416) or Wright (1976: 226). Fruitbody pileate, spathulate to dimidiate and applanate, up to 4 cm long and 3 cm wide, 1-5 mm thick, brittle when dry. Pileus golden brown to cinnamon, azonate, finely tomentose in radial lines or striae. Pore surface golden brown, pores angular and somewhat dentate, 2-4 per mm. Tubes concolorous, up to 3 mm deep. Context cinnamon and very thin. Hyphal system monomitic, generative hyphae simple septate, hyaline to golden brown, thin- to thick-walled, 2-5 pm wide. Setae none. Spores ellipsoid, smooth, pale yellow to rusty brown, 4-5 X 3 4 5 pm. Probably widespread in tropical America. Rome11 (1901: 26) described it as Phueoporus ferrugineus from Brazil and Fries (1851: 79) from Mexico, Holmquist (1972: 79) reports several collections from Venezuela. F. tewulatus Mont. 1843: 365 (isotype in K) “Crypt. Guyan. (French Guiana) no. 393. Leprieur coll. no. 599”. = Fuvolus brasilensis (Fr.) Fr. 1828: 44. zyxwvutsr zyx F. trachypus Berk. & Mont. in Mont. 1856: 154 “Colombus (Ohio, USA) Sullivant”. = P. badius ( S . F. Gray) Schw. P. tricholoma Mont. 1837: 365 (isotype in K) “Crypt. Guyan. (French Guiana) no. 336. Leprieur coll. no. 949”. P. suberis DR. & Mont. in Mont. 1856: 162 Tori daedaleoide ad Quercus suberi Cort.” (La Calle, Algerie). = Pheflinus contiguus (Fr.) Pat. 1900: 97. This is an accepted species in Polyporus. For a description, see Ryvarden & johansen (l980: 505). P. trichomallus Berk. & Mont. P. suUivantii Mont. 1842a: 243 (isotype in K) “Ohio, Am. Berkley no. 1 ex coll. Sullivant”. = Trametes pubescens (Fr.) Pil. 1939: 268. P. surinamensis Mont. - 1849: 238 “Crypt. Guyan. (French Guiana) no. 975. Leprieur coll. no. 962”. = Trichaptum perroftettii (Ltv.) Ryv. 1972a: 237. The name was based on the same collection even if Leveille (1844: 195) confused the data. The species is the type of Trichaptum Mun. 1843: 363. = P. tephroporus Mont. 1845b: 358. Nom. illeg. non P. surinamens6 Miq. 1839. Montagne became aware of this and renamed his species P . tephroporus (see below). 82 P. tristiculus Mont. 1854: 129 (isotype in K) “Guyana” zyxwvuts zyxwvutsrqp zyxwvu zyxwvutsr zyxwvuts zyxwvu The type is of unknown affinities, but belongs in Polyporus s. str. Fruitbody dimidiate with contracted base, about 2 x 2 cm and 2 - 4 m m thick. Pileus blackish, smooth, glabrous and azon’ate. Pore surface brown, pores 1 m m deep, round to angular, (3)4-5 per mm, tubes concolorous with pore surface, context dark brown, dense 1-2 m m thick. Hyphal systems dimitic, generative hyphae 2-3 pm wide and wi.th clamps, dendroid binding hyphae dominating in the fruitbody, solid, 2-5 pm wide, not conspicuously widened in the upper part. No spores and cystidia seen. The medium-sized pores remind one about P. guinnensk Mont., but this is usually a distinctly stipitate species and with a pale ochraceous pileus. It is probable that the type represents an atypical specimen of a more widespread species. 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