PHYTOLOGIA BALCANICA 12 (2): 169–180, Sofia, 2006
169
Check-list of the liverworts and hornworts of
Southeast Europe
Marko Sabovljević1,2 & Rayna Natcheva3
1
Department of Plant Ecology and Phytogeography, Institute of Botany and Botanical
Garden, Faculty of Biology, University of Belgrade, Takovska 43, 11000 Belgrade, Serbia
and Montenegro, e-mail: marko@bfbot.bg.ac.yu
2 AG Bryologie, Nees Institute für Pflanzen Biodiversität, RFW Universität Bonn,
Meckenheimer Allee 170, D-53115 Bonn, e-mail: marko@ uni-bonn.de
3 Institute of Botany, Bulgarian Academy of Sciences, Akad. G. Bonchev St., bl. 23, 1113
Sofia, Bulgaria, e-mail: renimoss@bio.bas.bg
Received: January 19, 2006 ▷ Accepted: February 25, 2006
Abstract.
The article presents the first compilation list of liverworts and hornworts from Southeast Europe. The region
of Southeast Europe includes up to date 272 species (5 hornworts and 267 liverworts), or ca. 59.9 % of the
hepatic flora of Europe. The countries with the richest hepatic flora within the region are Romania and
Bulgaria. The level of knowledge on liverworts and hornworts occurring in each of the countries of Southeast
Europe is different and some regions stand out as extremely poorly known. The number of hepatic species is
expected to increase with further investigations within Southeast Europe.
Key words:
by-country distribution, check-list, hornworts, liverworts, Southeast Europe
Introduction
Biogeographically, Southeast Europe (designated
hereafter as SE Europe) is a highly interesting area and
one of the regions with the richest flora in Europe. The
areas of Serbia and the mountainous parts of Bulgaria
are two of the six biodiversity centres in Europe
(IUCN 1987). Such high biodiversity is determined
by the various climatic and edaphic conditions in the
area and its geographic location. SE Europe exhibits almost all European climatic types, with many local variants and microclimates. The relief is extremely
varied, ranging from coastal (sea level) zones and inland steppe areas to mountain ranges above 2000 m
in altitude. The main mountain systems in SE Europe
are the Rilo-Rhodopean massive, the Carpathians,
the Balkan Range, the Dinaric Alps, and the ScardoPinds. Geology is complex: from old volcanic granite
and metamorphic rocks to Triassic and Jurassic lime-
stone, with many transitional types. The drainage network is well but unevenly developed in various parts.
SE Europe contains all major European vegetation
types, ranging from Mediterranean deciduous forests
and steppe areas, to deciduous nemoral and coniferous boreal forests, and high mountain alpine and arctic-alpine ecosystems. Furthermore, SE Europe is situated at the crossroad of biogeographic influences from
North Africa, West Asia, Central and North Europe.
All these specificities of the region of SE Europe had
set preconditions for a rich bryophyte flora. It is estimated that the number of species will reach ca. 1500,
with a total of about 2000 taxa, including intraspecifc
taxa (Sabovljević & al. 2001). However, the bryophyte
flora of SE Europe still remains very poorly known,
as compared to other regions of Europe (Sabovljević
2004). The reasons are complex but we shall mention
only a few. Historically there have been few resident
bryologists, a tendency that is even more evident to-
170
Sabovljević, M & Natcheva, R. • Liverwort check-list of Southeast Europe
day. Coupled with the large number of relatively small
in size states, this has encumbered further the bryo-floristic studies. Data on the bryophyte flora of SE Europe
are scattered in various small publications. Few of the
SE European countries have recent national check-lists
of liverworts and hornworts, e.g. Greece (Düll 1995),
Serbia and Montenegro (Sabovljević 2000), Croatia
(Sabovljević 2003), Albania (Colacino & Sabovljević
in press), Bulgaria (Ganeva & Natcheva 2003), and
Macedonia (Cekova 2005). Considering that in groups
of good dispersers like bryophytes a regional approach
is most appropriate with respect to both biodiversity
studies and conservation, a comprehensive list of the
species occurring in the entire region is greatly needed. Therefore, we present the first part of a check-list
of the bryophytes of SE Europe, dealing with the liverworts and hornworts in the region.
Methods
Study area
In the present paper SE Europe is regarded in a political sense, as comprising the states of Albania, Bulgaria,
Bosnia-Herzegovina, Croatia, Greece, Macedonia
(FYR), Romania, Serbia and Montenegro, Slovenia,
and the European part of Turkey (the Turkish Thrace)
(Fig. 1). Thus, as a geographical unit, it covers a greater area than the Balkan Peninsula, which is defined as
the land south of rivers Krka, Sava and the Danube.
Species listing
The published records were intensively searched
through for liverworts and hornworts in SE Europe.
Some recent collections have also been checked. The list
presents by-country distribution of the liverwort and
hornwort species in SE Europe, allowing a quick reference to the general distribution of each species as well
as to the total species diversity of each country. No extensive list of references will be presented in this study
due to space limitation, but the main references which
include most earlier records and some recent references with newly made records, not cited elswhere, are provided (Müller 1951-1958; Ade & Kope 1955; Pavletić
1955; Pavletić & Grom 1958; Pavletić & Zabijakin 1960;
Pócs 1960; Grom 1963, 1969; Duda 1965; Váňa & Duda
1965; Popović 1966; Boros 1968; Grolle 1970; Erdeši
1971; Bischler & Jovet-Ast 1973; Hébrard 1975; Petrov
1975; Pavletić & Pulević 1980; Düll 1983, 1995; Grgić
1983, 1989; Petrov & Ganeva 1996; Kürschner & Parolly
1997; Sabovljević 1998, 1999, 2000, 2003; Blockeel & al.
1999, 2002, 2004; Düll & al. 1999; Jakab 2000; Stefanut
2000, 2003, 2004; Papp & Sabovljević 2001, 2002, 2003;
Veljić & al. 2001; Sabovljević & Sérgio 2002; Balint &
Orban 2003; Dragićević & al. 2003; Ganeva & Natcheva
2003; Lüth 2003; Sabovljević & Cvetić 2003; Stefanut &
Stefanut 2003; Papp & al. 2004; Cekova 2005; Papp &
Erzberger 2005; Colacino & Sabovljević in press).
Nomenclature.
The nomenclature follows Grolle &
Long (2000) and Söderström & al.
(2002). A list of synonyms found
in earlier sources is provided, with
reference to the currently accepted
names (Appendix 1).
Fig. 1. The SE Europe and its position in Europe. Country abbreviations:
A – Austria; AL – Albania; BG – Bulgaria;
BH – Bosnia & Herzegovina; CG – Montenegro; GR – Greece; H – Hungary;
HR – Croatia; I – Italy; MD – Moldova,
MK – Macedonia (FYR); RO – Romania;
SK – Slovakia; SLO – Slovenia; SR – Serbia;
TR – Turkey; UA – Ukraine.
171
Phytol. Balcan. 12(2) • Sofia • 2006
Results and discussion
The present check-list includes a total of 272 species reported in SE Europe (Table 1). Of these, five are hornworts and 267 are liverworts. Thus the flora of liverworts and hornworts of SE Europe comprises 59.9 %
of the total hepatic flora of Europe and Macaronesia,
or 63.3 % if only continental Europe is considered.
The number of species falls within the range of hepatic diversity found in the other two major South
European peninsulas, the Iberian with 264, and the
Apennine with 271 species (Sabovljević 2004), and is
slightly lower than in Scandinavia (ca. 290, Damsholt
2002). The Balkan Peninsula is among the regions in
greatest need of investigation, and it is anticipated that
a high number of liverwort species will be found there
(Söderström & al. 1998). Considering that generally
there is less information on the bryophyte flora of SE
Europe as compared to other parts of Europe, it could
be expected that the number of species will further increase with some more thorough field investigations.
On a national level, the countries with richest floras
are Romania, followed by Bulgaria, Slovenia, Greece
and Croatia. The great difference in the species num-
bers among the various countries (ranging from 27 up
to 197) only partly reflects the real difference in species diversity. Forty-one species are known in only one
of the 11 SE European countries, while 31 are recorded in only two of them. This also indicates a difference in the state of knowledge of their liverwort and
hornwort floras. Particularly underestimated is the
number of species e.g. in Montenegro, Macedonia and
Albania. A higher number of local experts and a better cooperation between the bryologists from the SE
European countries are needed to improve our knowledge on national and regional level.
Thirty six species occurring within the region of SE
Europe are of conservation interest for Europe (ECCB
1995). For good dispersers such as bryophytes, a regional approach to conservation is the most appropriate one with respect to actual protection and resource
efficiency. In this context, more detailed knowledge
on the distribution of species within each country of
SE Europe is required, in order to evaluate the actual
threats and to set targets for liverwort and hornwort
conservation on regional and national level. A detailed check-list is the first step towards the achievement of these goals.
Table 1. Check-list of the liverworts and hornworts of SE Europe. Abbreviations and symbols: AL – Albania; BG – Bulgaria;
BH – Bosnia & Herzegovina; CG – Montenegro; GR – Greece incl. Crete; HR – Croatia; MK – Macedonia (FYR); RO – Romania;
SB – Serbia; SLO – Slovenia; TR – Turkey (European part only); ? – presence uncertain; (●) – record without precise locality;
* – species in the European Red List (ECCB 1995).
Species
AL
Anthocerotae
Anthoceros agrestis Paton
A. punctatus L.
Phaeoceros bulbiculosus (Brot.) Prosk.
Ph. carolinianus (Michx.) Prosk.
Ph. laevis (L.) Prosk.
●
●
Hepaticae
Anstrepta ocardensis (Hook.) Schiffn.
Anastrophyllum hellerianum (Nees ex Lindenb.) R.M. Schust.
A. michauxii (F. Weber) H. Buch
A. minutum (Schreb.) R.M. Schust.
A. saxicola (Schrad.) R.M. Schust.
Aneura pinguis (L.) Dumort.
Anthelia julacea (L.) Dumort.
A. juratzkana (Limpr.) Trevis.
Apometzgeria pubescens (Schrank) Kuwah.
Arnellia fennica (Gottsche) Lindb.*
Asterella gracilis (F. Weber) Underw.
A. lindenbergiana (Corda ex Nees) Arnell
A. saccata (Wahlenb.) A. Evans*
Athalamia hyalina (Sommerf.) S. Hatt.
A. spathysii (Lindenb.) S. Hatt.*
Barbilophozia attenuata (Mart.) Loeske
B. barbata (Schmidel ex Schreb.) Loeske
B. floerkei (F. Weber & D. Mohr) Loeske
●
●
●
●
●
BG
BH
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●
CG
GR
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MK
RO
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HR
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?
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SR
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SLO
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●
TR
172
Sabovljević, M & Natcheva, R. • Liverwort check-list of Southeast Europe
Table 1. Continuation
Species
B. hatcheri (A. Evans) Loeske
B. kunzeana (Huebener) Müll.Frib.
B. lycopodioides (Wallr.) Loeske
B. quadriloba (Lindb.) Loeske
Bazzania flaccida (Dumort.) Grolle*
B. tricrenata (Wahlenb.) Lindb.
B. trilobata (L.) Gray
Blasia pusilla L.
Blepharostoma trichophyllum (L.) Dumort.
Bucegia romanica Radian*
Calypogeia arguta Nees & Mont.
C. azurea Stotler & Crotz
C. fissa (L.) Raddi
C. integristipula Steph.
C. muelleriana (Schiffn.) Müll.Frib.
C. neesiana (C.Massal. & Carestia) Müll.Frib.
C. sphagnicola (Arnell & J.Perss.) Warnst. & Loeske
C. suecica (Arnell & J. Perss.) Müll.Frib.
Cephalozia ambigua C. Massal.
C. bicuspidata (L.) Dumort.
C. catenulata (Huebener) Lindb.
C. connivens (Dicks.) Lindb.
C. lacinulata J.B. Jack ex Spruce*
C. leucantha Spruce
C. loitlesbergeri Schiffn.
C. lunulifolia (Dumort.) Dumort.
C. pleniceps (Austin) Lindb.
Cephaloziella baumgartneri Schiffn.
C. calyculata (Durieu & Mont.) Müll.Frib.*
C. dentata (Raddi) Steph.
C. divaricata (Sm.) Schiffn.
C. elachista (J.B. Jack ex Gottsche & Rabenh.) Schiffn. *
C. elegans (Heeg) Schiffn.*
C. hampeana (Nees) Schiffn.
C. integerrima (Lindb.) Warnst.
C. massalongi (Spruce) Müll.Frib.*
C. rubella (Nees) Warnst.
C. stellulifera (Taylor ex Spruce) Schiffn.
C.turnerii (Hook.) Müll.Frib.
Chiloscyphus pallescens (Ehrh. ex Hoffm.) Dumort.
Ch. polyanthos (L.) Corda
Cladopodiella fluitans (Nees) H. Buch
Cololejeunea calcarea (Lib.) Schiffn.
C. minutissima (Sm.) Schiffn.
C. rossettiana (C. Massal.) Schiffn.
Conocephalum conicum (L.) Dumort.
C. salebrosum Szweykowski, Buczkowska & Odrzykoski
Corsinia coriandrina (Spreng.) Lindb.
Diplophyllum albicans (L.) Dumort.
D. obtusifolium (Hook.) Dumort.
D. taxifolium (Wahlenb.) Dumort.
Eremonotus myriocarpus (Carrington) Pearson
Fossombronia angulosa (Dicks.) Raddi
F. caespitiformis De Not. ex Rabenh.
F. echinata Macvicar*
F. foveolata Lindb.
F. husnotii Corb.
F. pusilla (L.) Nees
F. wondraczekii (Corda) Lindb.
Frullania dilatata (L.) Dumort.
F. fragilifolia (Taylor) Gottsche & al.*
F. inflata Gottsche*
F. jackii Gottsche
F. parvistipula Steph.*
F. riparia Hampe ex Lehm.
AL
BG
BH
CG
GR
HR
MK
RO
SR
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(●)
●
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SLO
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(●)
●
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●
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●
●
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●
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●
(●)
●
●
(●)
TR
●
●
●
●
●
●
●
●
●
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●
●
●
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●
●
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●
173
Phytol. Balcan. 12(2) • Sofia • 2006
Table 1. Continuation
Species
AL
BG
BH
CG
GR
HR
MK
RO
SR
SLO
F. tamarisci (L.) Dumort.
F. tenerifae (F. Weber) Nees
Geocalyx graveolens (Schrad.) Nees
Gongylanthus ericetorum (Raddi) Nees
Gymnocolea inflata (Huds.) Dumort.
Gymnomitrion apiculatum (Schiffn.) Müll.Frib.
G.concinnatum (Lightf.) Corda
G. corallioides Nees
Haplomitrium hookeri (Sm.) Nees*
Harpanthus flotovianus (Nees) Nees
H. scutatus (F.Weber et D. Mohr) Spruce
Jamsoniella autumnalis (DC.) Steph.
Jungermannia atrovirens Dumort.
J. borealis Damsh. & Váña
J. caespiticia Lindenb.
J. confertissima Nees
J. exertifolia Steph.
J. gracillima Sm.
J. handelii (Schiffn.) Amakawa*
J. hyalina Lyell
J. leiantha Grolle
J. obovata Nees
J. polaris Lindb.
J. pumila With.
J. sphaerocarpa Hook.
J. subelliptica (Lindb. ex Kaal.) Levier
J. subulata A. Evans*
Kurzia pauciflora (Dicks.) Grolle
Leiocolea badensis (Gottsche) Jörg.
L. bantriensis (Hook.) Jörg.
L. collaris (Nees) Schljakov
L. gillmanii (Austin) A. Evans
L. heterocolpos (Thed. ex Hartm.) H. Buch
L. turbinata (Raddi) H. Buch
Lejeunea cavifolia (Ehrh.) Lindb.
Lepidozia cupressina (Sw.) Lindenb.
L. reptans (L.) Dumort.
Lophocolea bidentata (L.) Dumort.
L. heterophylla (Schrad.) Dumort.
L. minor Nees
Lophozia ascendens (Warnst.) R.M. Schust.*
L. bicrenata (Schmidel ex Hoffm.) Dumort.
L. decolorans (Limpr.) Steph.*
L. elongata Steph.*
L. excisa (Dicks.) Dumort.
L. incisa (Schrad.) Dumort.
L. latifolia R.M.Schust.*
L. longidens (Lindb.) Macoun
L. longiflora (Nees) Schiffn.
L. obtusa (Lindb.) A. Evans
L. opacifolia Culm. ex Meyl.
L. polaris (R.M. Schust.) R.M. Schust. & Damsh.
L. sudetica (Nees ex Huebener) Grolle
L. ventricosa (Dicks.) Dumort.
L. wenzelii (Nees) Steph.
Lunularia cruciata (L.) Lindb.
Mannia androgyna (L.) A. Evans
M. fragrans (Balbis) Frye & L. Clark
M. pilosa (Hornem.) Frye & L. Clark
M. triandra (Scop.) Grolle*
Marchantia paleacea Bertol.*
M. polymorpha L.
Marchesinia mackaii (Hook.) Gray*
Marsupella adusta (Nees emend. Limpr.) Spruce*
M. alpina (Gottsche ex Husn.) Bernet
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5 • Phytol. Balcan. 12(2) • 2006
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?
?
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174
Sabovljević, M & Natcheva, R. • Liverwort check-list of Southeast Europe
Table 1. Continuation
Species
M. boeckii (Austin) Kaal.
M. brevissima (Dumort.) Grolle
M. commutata (Limpr.) Bernet
M. emarginata (Ehrh.) Dumort.
M. funckii (F.Weber & D. Mohr). Dumort.
M. ramosa Müll.Frib.
M. sparsifolia (Lindb.) Dumort.
M. sphacelata (Gieseke ex Lindenb.) Dumort.
M. sprucei (Limpr.) Bernet
Metzgeria conjugata Lindb.
M. fruticulosa (Dicks.) A. Evans
M. furcata (L.) Dumort
M. simplex Lorb. ex Müll.Frib.*
M. temperata Kuwah.
Microlejeunea ulicina (Taylor.) A. Evans
Moerkia blyttii (Moerch) Brockm.
M. hibernica (Hook.) Gottsche
Mylia anomala (Hook.) Gray
M. taylorii (Hook.) Gray
Nardia compressa (Hook.) Gray
N. geoscyphus (De Not.) Lindb.
N. insecta Lindb.
N. scalaris Gray
Nowelia curvifolia (Dicks.) Mitt.
Odontoschisma denudatum (Mart.) Dumort.
O. sphagni (Dicks.) Dumort.
Oxymitra incrassata (Brot.) Sérgio & Sim-Sim
Pallavicinia lyellii (Hook.) Carruth.*
Pedinophyllum interruptum (Nees) Kaal.
Pellia endiviifolia (Dicks.) Dumort.
P. epiphylla (L.) Corda
P. neesiana (Gottsche) Limpr.
Peltolepis quadrata (Saut.) Müll.Frib.
Petalophyllum ralfsii (Wils.) Nees & Gottsche*
Plagiochasma rupestre (J. R. Forst. & G. Forst) Steph.
Plagiochila asplenioides (L. emend. Taylor) Dumort.
P. porelloides (Torrey ex Nees) Lindenb.
Pleurocladula albescens (Hook.) Grolle
Porella arboris-vitae (With.) Grolle
P. baueri (Schiffn.) C.E.O. Jensen*
P. cordeana (Huebener) Moore
P. obtusata (Taylor) Trevis.
P. pinnata L.
P. platyphylla (L.) Pfeiff.
Preissia quadrata (Scop.) Nees
Ptilidium cilliare (L.) Hampe
P. pulcherimum (Weber) Vain.
Radula complanata (L.) Dumort.
R. lindenbergiana Gottsche ex C. Hartm.
Reboulia hemisphaerica (L.) Raddi
Riccardia chamedryfolia (With.) Grolle
R. incurvata Lindb.
R. latifrons (Lindb.) Lindb.
R. multifida (L.) Gray.
R. palmata (Hedw.) Carruth.
Riccia atromarginata Levier
R. beyrichiana Hampe ex Lehm.
R. bicarinata Lindb.
R. bifurca Hoffm.
R. canaliculata Hoffm.
R. cavernosa Hoffm.
R. ciliata Hoffm.
R. ciliifera Link ex Lindenb.
R. crinita Taylor
R. crozalsii Levier
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Phytol. Balcan. 12(2) • Sofia • 2006
Table 1. Continuation
Species
R. crustata Trab.*
R. crystalina L. emend. Raddi
R. fluitans L.
R. frostii Austin*
R. glauca L.
R. gougetiana Durieu & Mont.
R. lamellosa Raddi
R. macrocarpa Levier
R. michelii Raddi
R. nigrella DC.
R. papillosa Moris
R. perennis Steph.
R. sommieri Levier*
R. sorocarpa Bisch.
R. subbifurca Warnst. ex Corz.
R. trabutiana Steph.*
R. warnstorfii Limpr. ex Warnst.
Ricciocarpos natans (L.) Corda
Riella notarisii (Mont.) Mont.*
Sauteria alpina (Nees) Nees
Scapania aequiloba (Schwägr.) Dumort.
S. apiculata Spruce
S. aspera Bernet & M. Bernet
S. calcicola (Arnell & J. Press.) Ingham
S. compacta (A. Roth) Dumort.
S. crassiretis Bryhn
S. curta (Mart.) Dumort.
S. cuspiduligera (Nees) Müll.Frib.
S. gracilis Lindb.
S. helvetica Gottsche*
S. irrigua (Nees) Nees
S. lingulata H. Buch
S. mucronata H. Buch
S. nemorea (L.) Grolle
S. paludicola Loeske & Müll.Frib.
S. paludosa (Müll.Frib.) Müll.Frib.
S. scandica (Arnell & H.Buch) Macvicar
S. subalpina (Nees ex Lindenb.) Dumort.
S. uliginosa (Sw. ex Lindenb.) Dumort.
S. umbrosa (Schrad.) Dumort.
S. undulata (L.) Dumort.
S. verrucosa Heeg*
Southbya nigrella (De Not.) Henriq.
S. tophacea (Spruce) Spruce
Sphaerocarpos michelii Bellardi
S. texanus Austin
Targionia hypophylla L.
T. lorbeeriana Müll.Frib.
Trichocolea tomentella (Ehrh.) Dumort.
Tritomaria exsecta (Schmidel) Loeske
T. exsectiformis (Breidl.) Loeske
T. polita (Nees) Jörg.
T. quinquedentata (Huds.) H.Buch
T. scitula (Taylor) Jörg.
Total SE Europe = 272
Proportion from the total of SE European hepatic flora
Proportion from the total of European hepatic flora
(SE Europe 59.9%)
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43.5
167
61.6
27
9.9
25.9
36.8
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91
35.1
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64.4
105
38.6
88
31.9
144
52.9
157
57.9
69
25.3
20.8
38.5
22.9
19.0
31.6
34.6
15.2
44.9
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75.4
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176
Sabovljević, M & Natcheva, R. • Liverwort check-list of Southeast Europe
Appendix 1.
Major synonyms found in the bryological literature concerning SE Europe,
with reference to the currently accepted names (in bold).
Alicularia compressa (Hook.) Nees = Nardia compressa
Alicularia geoscypha De Not. = Nardia geoscyphus
Alicularia scalaris (Schrader) Corda = Nardia scalaris
Anastrophyllum myriocarpum (Carrington) Schust. ex Váňa = Eremonotus myriocarpus
Aneura latifrons Lindb. = Riccardia latifrons.
Aneura multifida (L.) Dumort. = Riccardia multifida
Aneura palmata (Hedw.) Dumort. = Riccardia palmata
Aneura sinuata (Dicks.) Dumort. = Riccardia chamedryfolia
Anthoceros dichotomus Raddi = Phaeoceros bulbiculosus
Anthoceros husnotii Steph. = Anthoceros punctatus
Anthoceros laevis L. = Phaeoceros laevis
Aplosia lanceolata (Schrad.) Dum. = Jungermannia leiantha
Barbilophozia gracilis (Schleich. ex Steph.) = Barbilophozia attenuata
Barbilophozia hatcheri (A. Evans) Loeske = Lophozia hatcheri
Bazzania denudata auct. = Bazzania flaccida
Calypogeia trichomanis auct. = Calypogeia azurea
Cephalozeilla bifidioides Doiun = Cephaloziella rubella
Cephalozeilla latzeliana Schiffn. = Cephaloziella divaricata
Cephalozia affinis Lindb. ex Steph. = Cephalozia lunulifolia
Cephalozia fluitans (Nees) Spruce = Cladopodiella fluitans
Cephalozia lammerisiana (Huebener) F. Lees = Cephalozia bicuspidata
Cephalozia media Lindb. = Cephalozia lunulifolia
Cephaloziella limprichti Warnst. = Cephaloziella stellulifera
Cephaloziella miriantha (Lindb.) Schiffn. = Cephaloziella rubella
Cephaloziella papillosa (Douin) Schiffn. = Cephalozeilla divaricata
Cephaloziella starkei (Funck.) Schiffn. = Cephaloziella divaricata
Chiloscyphus fragilis (A. Roth) Schiffn. = Chiloscyphus pallescens
Chiloscyphus polyanthos var. pallescens (Ehrh. ex Hoffm.) C. Hartm. = Chiloscyphus pallescens
Chiloscyphus rivularis (Schrad.) Hazsl.= Chiloscyphus polyanthos
Clevea hyalina (Sommerf.) Lindb. = Athalamia hyalina
Corsinia marchantioides Raddi = Corsinia coriandrina
Crossocalyx hallerianus (Nees ex Lindenb.) Meyl. = Anastrophyllum hallerianum
Dichyton calyculatum (Durieu & Mont.) Schiffn. = Cephaloziella calyculata
Eucalix hyalinus (Lyell.) Breidl. = Jungermannia hyalina.
Eucalix obovatus (Nees) Breidl. = Jungermannia obovata
Eucalyx subellipticus (Lindb. ex Kaal.) Breidl. = Jungermannia subelliptica
Fegatella conica Corda ex Opiz = Conocephalum conicum
Fimbriaria lindenbergiana Corda ex Nees = Asterella lindenbergiana
Fimbriaria pilosa Taylor = Asterella gracilis
Fossombronia loitlesbergi Schiffn. = Fossombronia wondraczekii
Fossombronia verrucosa Lindb.= Fossombronia caespitiformis
Frullania illyrica Grolle = Frullania inflata
177
Phytol. Balcan. 12(2) • Sofia • 2006
Continuation Appendix 1.
Frullania nervosa Mont. = Frullania tamarisci
Grimaldia dichotoma Raddi = Mannia androgyna
Grimaldia fragrans (Balbis) Corda = Mannia fragrans
Grimaldia pilosa (Hornem.) Lindb. = Mannia pilosa
Grimaldia rupestris (Nees) Lindb. = Mannia triandra
Haplozia atrovirens (Schleich.) Dumort. = Jungermannia atrovirens
Haplozia crenulata (Mitt.) Müll.Frib. = Jungrmannia gracilima
Haplozia pumila (With.) Dumort. = Jungermannia pumila
Haplozia riparia (Taylor) Dumort. = Jungermannia atrovirens
Haplozia schiffneri Loitl. = Jungermannia polaris
Haplozia sphaerocarpa (Hook.) Dumort. = Jungermannia sphaerocarpa
Isopaches decolorans (Limpr.) Buch = Lophozia decolorans
Jungermannia atrovirens Dumort. = Jungermannia lanceolata
Jungermannia crenulata Sm. = Jungermannia gracillima
Jungermannia eucordifolia Schljak. = Jungermannia exsertifolia ssp. cordifolia
Jungermannia karlmuelleri Grolle = Jungermannia pumila
Jungermannia lanceolata auct. = Jungermannia leiantha
Jungermannia pendletonii (Pearson) A. Evans = Jungermannia exsertifolia subsp. cordifolia
Leiocolea mülleri (Nees ex Lindb.) Dumort. = Leiocolea collaris
Leiocolea obtusa (Lindb.) H. Buch = Lophozia obtusa
Lejeunea ulicina (Taylor) Gottsche & al. = Microlejeunea ulicina
Lepidozia setacea (Web.) Mitten = Kurzia pauciflora
Leptoscyphus anomalus (Hook.) Lindb. = Mylia anomala
Leptoscyphus taylori (Hook.) Mitt. = Mylia taylorii
Lophocolea alata Mitt. ex Larter = Lophocolea bidentata
Lophocolea cuspidata (Nees) Limpr. = Lophocolea bidentata
Lophozia kiaeri Jörg. = Lophozia excisa
Lophozia porphyroleuca auct. = Lophozia longiflora
Lophozia alpestris (Schleich.) Evans = Lophozia polaris
Lophozia attenuata (Mart.) Dumort. = Barbilophozia attenuata
Lophozia bantriensis (Hook.) Steph. = Leiocolea bantriensis
Lophozia barbata (Scmidel ex Schreb.) Dumort. = Barbilophozia barbata
Lophozia collaris (Nees) Dumort. = Leiocolea collaris
Lophozia confertifolia Schiffn. = Lophozia wenzelii
Lophozia ehrhartiana (F. Weber) Inoue & Steere = Lophozia ventricosa
Lophozia floerkei (F. Weber &D. Mohr) Loeske = Barbilophozia floerkei
Lophozia guttulata (Lindb.) Evans = Lophozia longiflora
Lophozia heterocolpos (Thed. ex Hartm.) Howe = Leiocolea heterocolpos
Lophozia lycopodioides (Wallr.) Cogn. = Barbilophozia lycopodioides
Lophozia mülleri (Nees) Dumort. = Leiocolea collaris
Lophozia quadriloba (Lindb.) Evans = Barbilophozia quadriloba
Lophozia quinquedentata (Huds.) Cogniaux = Tritomaria quinquedentata
Lophozia silvicola H. Buch = Lophozia ventricosa
Lophozia turbinata (Raddi) Steph. = Leiocolea turbinata
Madotheca baueri Schiffn. = Porella baueri
Madotheca cordeana (Huebener) Dumort. = Porella cordeana
Madotheca lavigata (Schrad.) Dumort. = Porella arboris-vitae
178
Sabovljević, M & Natcheva, R. • Liverwort check-list of Southeast Europe
Continuation Appendix 1.
Madotheca platyphylla (L.) Dumort. = Porella platyphylla
Madotheca platyphylloidea (Schwein.) Dumort. = Porella platyphylla
Mannia dichotoma Udar &V. Chandra = Mannia androgyna
Mannia rupestris (Nees) Frye & L. Clark = Mannia triandra
Marsupella aquatica (Lindenb.) Schiffn. = Marsupella emarginata
Marsupella pygmaea (Limpr.) Steph. = Marsupella funckii
Marsupella varians (Lindb.) Müll.Frib. = Marsupella brevissima
Marsupella hungariaca Boros & Vajda = Marsupella funckii
Marsupella ustulata Spruce = Marsupella sprucei
Metzgeria pubescens (Schrank) Raddi = Apometzgeria pubescens
Neesiella pilosa (Horn.) Schiffn. = Asterella gracilis
Neesiella rupestris (Nees) Schifn. = Mannia triandra
Orthocaulis floerkei (Web. & Mohr.) Buch. = Barbilophozia floerkei
Oxymitra paleacea Bosch. ex Lindenb. = Oxymitra incrassata
Pellia borealis Lorb. = Pellia epiphylla
Pellia fabbroniana auct. = Pellia endiviifolia
Peltolepis grandis Lindb. = Peltolepis quadrata
Plagiochila major (Nees) S.W. Arnell = Plagiochia asplenioides
Plectocolea hyalina (Lyell) Mitt. = Jungermannia hyalina
Plectocolea obovata (Nees) Mitt. = Jungermannia obovata
Porella laevigata (Schrad.) Lindb. var. thuja Nees = Porella obtusata
Porella platyphylloidea (Schwein.) Lindb. = Porella platyphylla
Preissia commutata Nees = Preissia quadrata
Riccardia sinuata (Hook.) Trev. = Riccardia chamaedryfolia
Riccardia pinguis (L.) S. Gray = Aneura pinguis
Riccia bischoffi Hüb. = Riccia ciliifera
Riccia commutata J.B. Jack ex Levier = Riccia warnstorfii
Riccia henriquesii Lev. = Riccia bicarinata
Riccia intumescens (Bisch.) Underw. = Riccia crinita
Riccia latzeli Schiffn. = Riccia ciliifera
Riccia levieri Schiffn. = Riccia beyrichiana
Riccia minima L. = Riccia sorocarpa
Riccia raddiana J.B. Jack & Levier = Riccia sorocarpa
Riccia trichocarpa M. Howe = Riccia crinita
Scapania dentata Dumort. = Scapania undulata
Scapania obscura (H. Arn. & C. Jens.) Schiffn. = Scapania subalpina
Solenostoma atrovirens (Dumort.) Müll.Frib. = Jungermannia atrovirens
Solenostoma caespiticium (Lindenb.) Steph. = Jungermannia caespiticia
Solenostoma cordifolium (Dumort.) Steph. = Jungermannia exsertifolia ssp. cordifolia
Solenostoma crenulatum Mitt. = Jungermannia gracillima
Solenostoma gracillimum (Sm.) R.M. Schust. = Jungermannia gracillima
Solenostoma oblongifolium (Müll.Frib.) Müll.Frib. = Jungermannia pumila
Solenostoma polare (Lindb.) R.M. Schust. = Jungermannia polaris
Solenostoma pumilum (With.) Müll.Frib. = Jungermannia pumila
Solenostoma schiffneri (Loitl.) Müll.Frib. = Jungermannia polaris
Solenostoma sphaerocarpoideum (De Not.) Paton & E.F. Warb. = Jungermannia atrovirens
Solenostoma sphaerocarpum (Hook.) Stef. = Jungermannia sphaerocarpa
179
Phytol. Balcan. 12(2) • Sofia • 2006
Continuation Appendix 1.
Solenostoma sphaerocarpum (Hook.) Steph. = Jungermannia sphaerocarpa
Solenostoma triste (Nees) Müll.Frib. = Jungermannia atrovirens
Southbya stilicidiorum (Raddi) Lindb. = Southbya tophacea
Sphaerocarpos terrestris Sm. = Sphaerocarpos michelii
Sphenobolus exsectus (Schmid.) Steph. = Tritomaria exsecta
Sphenobolus hallerianus (Nees) Steph. = Anastrophyllum hallerianum
Sphenobolus politus (Nees) Steph. = Tritomaria polita
Sphenolobus hallerianus (Nees ex Lindenb.) Steph. = Anastrophyllum hallerianum
Sphenolobus michauxii (Wes.) Steph. = Anastrophyllum michauxii
Sphenolobus minutus (Schreb.) Berggr. = Anastrophyllum minutum
Teeselina pyrmidata Dumort. = Oxymitra incrassata
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Marko Sabovljevic