New or interesting lichens and lichenicolous fungi from the Vadstena area, Östergötland, Sweden ARNE THELL, VAGN ALSTRUP, ULF ARUP, MIKA BENDIKSBY, PAWEL CZARNOTA, TASSILO FEUERER, REIDAR HAUGAN, INGVAR KÄRNEFELT, JON T. KLEPSLAND, MARTIN KUKWA, ANNINA LAUNIS, ANA M. MILLANES, JURGA MOTIEJŪNAITĖ, ANDERS NORDIN, MARIA PRIETO, JUHA PYKÄLÄ, MARK R. D. SEAWARD, EINAR TIMDAL, ANDREI TSURYKAU, ORVO VITIKAINEN and MARTIN WESTBERG Thell, A., Alstrup, V., Arup, U., Bendiksby, M., Czarnota, P., Feuerer, T., Haugan, R., Kärnefelt, I., Klepsland, J. T., Kukwa, M., Launis, A., Millanes, A. M., Motiejūnaitė, J., Nordin, A., Prieto, M., Pykälä, J., Seaward, M. R. D., Timdal, E., Tsurykau, A., Vitikainen, O. & Westberg, M. 2014. New or interesting lichens and lichenicolous fungi from the Vadstena area, Östergötland, Sweden. Graphis Scripta 26(1–2): 15– 33. Tartu. ISSN 0901-7593. Results from the excursion of the 20th biennial symposium in Vadstena 11–15 August 2013 are presented. Eight localities in and around Vadstena were visited. Forty-three species are reported as new to the province Östergötland [Ostrogothia], three of which are new to the Nordic countries: Lecanora compallens, Polycoccum kerneri and Tremella caloplacae, five are new to Sweden: Cornutispora ciliata, Pyrenula chlorospila, Thelidium cf. rimulosum, Verrucaria ochrostoma and V. polystictoides, and the following are new to Östergötland: Bagliettoa baldensis, B. calciseda, Briancoppinsia cytospora, Calogaya arnoldii ssp. obliterata, Clypeococcum hypocenomycis, Cornutispora lichenicola, Endococcus exerrans, Fuscidea arboricola, Illosporium carneum, Lepraria elobata, L. jackii, L. vouauxii, Leptochidium albociliatum, Lichenoconium lecanorae, Lichenochora weillii, Marchandiobasidium aurantiacum, Micarea byssacea, Monodictys anaptychiae, M. epilepraria, Parmelia serrana, Phaeosporobolus alpinus, Placopyrenium canellum, Porpidia soredizodes, Pyrenidium actinellum, Rinodina turfacea, Stereocaulon rivulorum, Syzygospora physciacearum, Thelenella muscorum, Tremella phaeophysciae, T. ramalinae, Trichonectria rubefaciens, Verrucaria dolosa, V. inaspecta, V. infumata and V. memnonia. Arne Thell, Botanical Museum, Lund University, Box 117, SE-221 00 Lund, Sweden. E-mail: arne.thell@biol.lu.se Vagn Alstrup, Dalkær 9.2.tv, DK-2605 Brøndby, Denmark. E-mail: vagnalstrup@gmail.com Ulf Arup, Botanical Museum, Lund University, Box 117, SE-221 00 Lund, Sweden. E-mail: ulf.arup@biol.lu.se Mika Bendiksby, NTNU University Museum, Norwegian University of Science and Technology, NO-7491 Trondheim, Norway. E-mail: mika.bendiksby@ntnu.no Pawel Czarnota, Department of Plant Taxonomy and Nature Conservation, University of Gdańsk, Wita Stwosza 59, PL-80-308 Gdansk, Poland. E-mail: pawczarnota@poczta.onet.pl Tassilo Feuerer, Biocentre Klein-Flottbek, Ohnhorststrasse 18, D-22609 Hamburg, Germany. E-mail: feuerer@botanik.uni-hamburg.de Reidar Haugan, Natural History Museum, University of Oslo, P.O. Box 1172 Blindern, NO-0318 Oslo, Norway. E-mail: reidar.haugan2@getmail.no Ingvar Kärnefelt, Botanical Museum, Lund University, Box 117, SE-221 00 Lund, Sweden. E-mail: ingvar.karnefelt@biol.lu.se Jon T. Klepsland, BioFokus, Gaustadalleen 21, NO-0349 Oslo, Norway. E-mail: jon@biofokus.no GRAPHIS SCRIPTA 26 (2014) Martin Kukwa, Department of Plant Taxonomy and Nature Conservation, University of Gdańsk, Wita Stwosza 59, PL-80-308 Gdansk, Poland. E-mail: dokmak@ug.edu.pl Annina Launis, Finnish Museum of Natural History, P.O. Box 7, FI-00014 Helsinki, Finland. E-mail: annina.launis@helsinki.fi Jurga Motiejūnaitė, Nature Research Centre, Žaliųjų ežerų 49, LT-08406 Vilnius, Lithuania. E-mail: jurga.motiejunaite@botanika.lt Ana M. Millanes, Universidad Rey Juan Carlos, Departamento de Biología y Geología, E-28933 Móstoles, Madrid, Spain. E-mail: ana.millanes@urjc.es Anders Nordin, Museum of Evolution, Uppsala University, Norbyvägen 18, SE-75236 Uppsala, Sweden. Email: anders.nordin@em.uu.se Maria Prieto, Deptartment of Botany, Swedish Museum of Natural History, Box 50007, SE-104 05 Stockholm, Sweden. E-mail: maria.prieto@nrm.se Juha Pykälä, Finnish Environment Institute, P.O. Box 140, FI-00251 Helsinki, Finland. E-mail: juha.pykala@ymparisto.fi Mark R. D. Seaward, Department of Archaeological, Geographical & Environmental Sciences, University of Bradford, Bradford BD7 1DP, United Kingdom. E-mail: m.r.d.seaward@bradford.ac.uk Einar Timdal, Natural History Museum, University of Oslo, P.O. Box 1172 Blindern, NO-0318 Oslo, Norway. E-mail: einar.timdal@nhm.uio.no Andrei Tsurykau, F. Skorina Gomel State University, Sovetskaja Street 104, BL-246019 Gomel, Belarus. Email: tsurykau@gmail.com Orvo Vitikainen, Finnish Museum of Natural History, P.O. Box 7, FI-00014 Helsinki, Finland. E-mail: orvo.vitikainen@helsinki.fi Martin Westberg, Department of Botany, Swedish Museum of Natural History, Box 50007, SE-104 05 Stockholm, Sweden. E-mail: martin.westberg@nrm.se Introduction Nordic Lichen Society meetings always result in new knowledge of the lichen flora of the visited area. The previous excursion in 2011 resulted in 30 new species for Lithuania (Motiejūnaitė et al. 2012). The 20th NLF-meeting in Vadstena 11–15 August 2013 gathered 35 participants, representing 13 countries, supplied with special skills in various fields of lichenology, Thus, although Östergötland [Ostrogothia] was already reasonably well investigated lichenologically, there was a great potential to significantly increase the number of taxa known from the province. A complete list of 958 taxa, based on a continuously updated on-line version of Santesson’s list of Fennoscandian lichens (Santesson et al. 2004, Nordin et al. 2014), compiled by Anders Nordin, was distributed among the participants with a realistic goal to reach thousand. The distribution of the lichens observed were compared with Santesson’s list and the on-line Danish lichen checklist based on the printed version by Søchting & Alstrup (2002). The nomenclature follows Santesson’s list, http://www.evolutionsmuseet.uu.se /databaser/santesson.html. Collectors and herbaria Collectors: VA=Vagn Alstrup, UA=Ulf Arup, MB=Mika Bendiksby, PCz=Pawel Czarnota, TF=Tassilo Feuerer, RH=Reidar Haugan, IK=Ingvar Kärnefelt, JTK=Jon T. Klepsland, MK=Martin Kukwa, AM=Ana M. Millanes, AL=Annina Launis, JM=Jurga Motiejūnaitė, AN=Anders Nordin, MP=Maria Prieto, JP=Juha Pykälä, ATh=Arne Thell, ET=Einar Timdal, ATs=Andrei Tsurykau, OV=Orvo Vitikainen, MW=Martin Westberg. Herbaria: BILAS=the herbarium of Institute of Botany, Nature Research Centre, Vilnius, C=The Natural 16 GRAPHIS SCRIPTA 26 (2014) History Museum, Copenhagen, GSU=Belarussian Polesye Scientific Herbarium of F. Skorina Gomel State University, H=Finnish Museum of Natural History, Helsinki, HBG=Herbarium, Hamburg University, LD=Botanical Museum of Lund University, O=The Natural History Museum, University of Oslo, UGDA=Herbarium, University of Gdansk, S=Swedish Museum of Natural History, Stockholm. The localities and their lichen diversity The surroundings of Vadstena are characterized by lowland agricultural areas, the mountain Omberg in the south and a long shore of Lake Vättern. Eight localities were selected, several of them previously not visited by lichenologists; these are situated in the western part of the province Östergötland in central, southern Sweden, close to the eastern shore of Lake Vättern (Fig. 1). Localities 1–2 were visited on 12 August, 3–7 on 13 August, and locality 8 on 14 August. Figure 1. The position of the localities studied. 1. Alvastra Abbey, 2. Hästholmen, 3. Nässja ancient stone circle, 4. Herrestad church, 5. Rök runestone, 6–7. Omberg: 6. Hjässatorget (the parking area) and 7. Hjässan, the top of Omberg, 8. Vadstena town. Locality 1. Alvastra Abbey Västra Tollstad parish. 58°17'48"N, 14°39'36"E. Alvastra Abbey was a Cistercian monastery founded by monks from France in 1143. It was dissolved in 1530 and appropriated by the Crown at the time of the reformation. Beautiful ruins of calcareous stonework of the church and other 17 GRAPHIS SCRIPTA 26 (2014) buildings remain, offering a large variety of substrates for lichens. Large trees of ash (Fraxinus excelsior), and oak (Quercus robur), near the ruins are of interest, as well as the southern slopes of Omberg, dominated by beech (Fagus sylvatica), a few 100 m northwards. New to Östergötland Bagliettoa baldensis – on calcareous stone wall, MP (S-F251633). This species was previously known from six Swedish provinces (Berglund & Andersson 2008). Bagliettoa calciseda – ruins of the medieval church, on calcareous stone wall, loose piece on the ground, JP 47007 (H) The species is known from several provinces in Sweden, Norway and Finland (Santesson et al. 2004). After this first record for the province, it was, in addition, collected the day after, at locality 3, on the stone fence surrounding Nässja churchyard (Table 1). Calogaya arnoldii ssp. obliterata – on ruin walls of lime or on mortar, UA (LD). Lepraria vouauxii – on mortar, VA SE1305 (C). The species is also known from several provinces in Fennoscandia (Arup & Ekman 1991, Tønsberg 1992). Monodictys anapthychiae – the ruins of the abbey, on Anaptychia ciliaris (thallus) growing on Fraxinus excelsior, MK 12267 (UGDA). This lichenicolous fungus has been collected only once earlier in the Nordic area, in the churchyard of Håtuna in Uppland, Sweden, in 1990, also on Fraxinus (Wedin 1993). Porpidia soredizodes – open area by the road, on tile [stictic acid by TLC], MK 12271 (UGDA). This lichen species was known from scattered localities in Sweden (Muhr 1991). Tremella phaeophysciae – on thalli of Phaeophyscia orbicularis growing on trunk of old Malus domestica, JM 10373 (BILAS; duplicate at S (S-F253103)). This lichenicolous fungus growing on P. orbicularis was previously known from a few provinces (Diederich 1996). Additional species collected. Athallia pyracea – abbey ruins, on Sambucus nigra, MK 12264 (UGDA); Calogaya arnoldii ssp. obliterata – on ruin walls of lime and on mortar, UA L13235 (LD); Calogaya pusilla – on ruin walls of lime and on mortar, UA [field note]; Lecanora argentata – beech forest, on Fagus sylvatica, MK 12259 (UGDA); Lecidella elaeochroma – MB, RH & ET (O-L184467); Lecidella scabra – open area by the road, on tile [atranorin, thuringone, arthothelin by TLC], MK 12272 (UGDA), RH 11900 (OL190330); Lecidella stigmatea – beech forest, on stone, MK 12258 (UGDA); Melanelixia fuliginosa – on acid rock, VA SE1306 (C); Lichenoconium aff. xanthoriae – open area, by the road, on Xanthoria parietina, mainly on thallus, growing on bark of Sambucus nigra, MK 12273 (UGDA); Peltigera canina – on soil among grasses, JM 10401 (BILAS), abbey ruins, on bryophytes over calcareous wall, MK 12261 [det. OV] (UGDA); Peltigera horizontalis – on shaded moss-covered stone, JM 10373 (BILAS), ATs (GSU); Phaeophyscia endophoenicea – on trunk of old Malus domestica in shaded and rather damp situation, JM 10402 (BILAS); Physcia adscendens – on mortar, VA SE1310 (C); Physconia enteroxantha – on granite stone, ATs (GSU); Polycauliona cf. phlogina – abbey ruins, on Sambucus nigra, MK 12265 (UGDA); Pyrenula nitida – beech forest, on Fagus sylvatica, MK 12260 (UGDA); Ramalina farinacea – at the tourist hotel, on roadside Betula [DNA 4233], TF (HBG-015004); Ramalina fastigiata – abbey ruins, on Sambucus nigra, MK 12269 (UGDA), at the tourist hotel, on roadside Betula [DNA 4228], TF (HBG-015005); Ramalina fraxinea – abbey ruins, on Sambucus nigra, MK 12268 (UGDA); Rinodina colobina – abbey ruins, on Sambucus nigra, MK 12262 (UGDA). Red-listed in Sweden as endangered (EN); Verrucaria caerulea – on calcareous stone, MP (S-F251634); Verrucaria pinguicula – ruins of the medieval church, on calcareous stone [thallus pale brown, continuous to cracked-areolate, c. 0.15–0.2 mm thick, areoles 0.2–0.4 mm, thalli surrounded by dark prothalline line. Perithecia 0.12–0.15 mm in diam. in surface view, 3/4–1immersed in thallus; involucrellum to exciple base level, c. 30–50 µm thick, appressed to exciple; exciple 0.12–0.15 mm in diam., pale brown to brown; periphyses c. 13–25 × 2–2.5 µm; spores 12–16 × 5–6 µm. JP 18 GRAPHIS SCRIPTA 26 (2014) 47010 (H). Verrucaria pinguicula was reported from northern Finland by Pykälä (2010b). The Swedish specimen matches rather well with the syntype of V. pinguicula in H-NYL (calcem privassicam prope Muggendorf Arnold). The small type specimen (H-NYL 3096) has tiny perithecia and thin involucrellum (c. 30–40 µm thick), however, periphyses seen (c. 10–15 × 2.5 µm) were shorter than in the Swedish specimen. In the recent literature V. pinguicula has been circumscribed to include specimens with larger perithecia, rather thick involucrellum (Orange 2013), or large variation in involucrellum thickness between different specimens (Breuss & Berger 2010); Verrucaria viridula – ruins of the medieval church, on calcareous stone wall, JP 47003, 47005 (H); Vouauxiella lichenicola – on Lecanora subfusca s. l. [det. JM], ATs (GSU). Locality 2. Hästholmen Östergötland province. Västra Tollstad parish. 58°16'43"N, 14°38'05"E. Siliceous rocks on the shore of Lake Vättern, and dry pine and birch forest. New to the Nordic countries Lecanora compallens van Herk & Aptroot – siliceous rocks on the lake shore, small forest, on Populus tremula, mixed with Lecidella elaeochroma [usnic acid and zeorin by TLC], MK 12294 (UGDA). A relatively newly described species known from northwestern, central and eastern Europe, i.e. Belgium, Germany, Great Britain, Lithuania, Luxembourg, the Netherlands, Poland and Ukraine (van Herk & Aptroot 1999). Polycoccum kerneri J. Steiner – siliceous rocks on the lake shore, on thallus of Lecidea fuscoatra s. str. growing on rock [ascomata 250 µm, asci 8-spored, hamathecium branched and anastomosing, ascospores distichously arranged, 1-septate (spores more or less equal in size), brown, minutelly verruculose, 13 × 8 µm, 15 × 8 µm, 13 × 9 µm], MK 12286 (UGDA). The species has a wide distribution in Europe, Canary Islands and North America (Hafellner et al. 2002). New to Östergötland Briancoppinsia cytospora – on Evernia prunastri on trunks of Alnus glutinosa and Sorbus sp., JM 10398 (BILAS). A new coelomycetous genus was proposed for this species by Diederich et al. (2011). It is widespread in Europe and USA, but was unknown in the Nordic area until now. The species was also collected in Vadstena, locality 8, close to the entrance of Vadstena Folkhögskola. Leptochidium albociliatum – siliceous rocks on the lake shore, on saxicolous bryophytes, MK 12296 (UGDA). The species has a scattered distribution in Fennoscandia, described by Jørgensen (2007). Lichenoconium lecanorae – on apothecia of Xanthoparmelia pulla, JM 10404 (BILAS). The distribution in Fennoscandia of this lichenicolous fungus was mapped by Alstrup & Ahti (2007) and Ihlen & Wedin (2008). An additional sample was collected at locality 7 (Table 1). It was earlier known from several Swedish provinces, fewer in Norway and Finland. Marchandiobasidium aurantiacum Diederich & Schultheis [the teleomorph of Marchandiomyces aurantiacus (Diederich et al. 2003, Diederich & Lawrey 2007)] – on thalli of Physcia tenella growing on trunks and branches of Sorbus sp., JM 10409 (BILAS). Earlier known from only two Swedish provinces just north of Östergötland. The species was also collected at localities 3 and 8 (Table 1). This lichenicolous fungus was reported from southern Europe by Etayo & Diederich (1996). Prior to these new localities, the species had been found only in the Swedish provinces Södermanland and Uppland in Fennoscandia. It is probably overlooked. Placopyrenium canellum – siliceous rock, MP (S-F251639). Previously known only from the 19 GRAPHIS SCRIPTA 26 (2014) islands Gotland and Öland in the Baltic Sea. Pyrenidium actinellum – siliceous rocks on the lake shore, on Leptochidium albociliatum [near ostiolum aeruginose, spores 3-septate, brown, end cells paler, 20–21× 8–9 µm], MK 12297 (UGDA). The species was earlier known from scattered occurrences in Fennoscandia (Ihlen & Wedin 2008, Puolasma et al. 2012). Östergötland is the sixth province in Sweden for this species. Rinodina turfacea var. turfacea – scarce on moss, northern slope, JP 47026 (H). This discovery in Östergötland is the southernmost locality in Sweden. It is a northern species in Sweden, being previously known from the province Värmland and northwards (Mayrhofer & Moberg 2002). Stereocaulon rivulorum – siliceous rock [atranorin and lobaric acid by TLC, det. M. Oset], MK 12277 (UGDA). This is a northern species in Fennoscandia. This is the southernmost record of this species in Sweden, earlier known from the Province Dalarna and northwards, thus, causing a gap in the distribution range. Thelenella muscorum – calciferous siliceous rock outcrop on the shore of lake Vättern, JP 47024 (H). Its distribution is poorly known with scattered occurrences in Fennoscandia, being know also from three additional provinces, Bohuslän, Dalarna and Uppland, according to GBIF, http://www.gbif.org/species. Additional species collected. Acarospora badiofusca – on siliceous rock on the shore, UA 13241 (LD); Agonimia tristicula – rocks on the lake shore, on bryophytes over rocks, MK 12289 (UGDA), associated with Catapyrenium daedaleum, MP (S-F251638); Arthonia atra – on acid rock, VA SE1308, SE1309 (C); Athallia cerinella – on twigs of Fraxinus excelsior near the shore, UA 13245 (LD); Athallia vitellinula – on perpendicular rock on the shore, UA 13237 (LD); Bryobilimbia hypnorum – siliceous rocks on the lake shore, on bryophytes [apothecia marginate, exciple and hypothecium red brown, hymenium almost hyaline, hymenium with dark granules turning green in K, ascospores simple, 10 × 6–7 µm], MK 12298 (UGDA), MK 12299 (UGDA); Catapyrenium daedaleum, MP (S-F251638); Cetraria aculeata – on rocks [DNA 4231], TF (HBG-015010); Clypeococcum hypocenomycis – on Hypocenomyce scalaris, VA SE1302 (C); Endocarpon adsurgens – on moss on northern slope, JP 47025 (H), MP (S-F251738); Endococcus exerrans – on Rhizocarpon lecanorinum, VA SE1303 (C); Catillaria chalybeia – MB, RH & ET (O-L184476); Diploschistes muscorum – siliceous rocks on the lake shore, on Cladonia squamules and bryophytes growing on rock, MK 12282 (UGDA), on Cladonia squamules growing on rock, MK 12292 (UGDA); Henrica melaspora – on flat rock on northern slope, JP 47023 (H); Lasallia pustulata – on siliceous rock on the lake shore, MK 12279 (UGDA), ATs (GSU); Lecanora sulphurea – MB, RH & ET (O-L184472); Lecidea fuscoatra – MB, RH & ET (O-L184473); Lecidea nylanderi – siliceous rocks on the lake shore, small forest, on Pinus sylvestris, MK 12293 (UGDA), ATs (GSU); Leptogium gelatinosum – on siliceous rocks at the lake shore, ATs (GSU); Lichenoconium usneae – on thalli of Ramalina fraxinea and Evernia prunastri growing on various deciduous trees, JM 10386 (BILAS); Lichenostigma rugosum – siliceous rocks at the lake shore, on Diploschistes muscorum, MK 12287 (UGDA); Melanelixia fuliginosa – on siliceous rock [DNA 4232], TF (HBG-015011); Miriquidica deusta – MB, RH & ET (O-L184474); Miriquidica leucophaea – on rocks [miriquidic acid by TLC], RH 11909 (O-L190339); Ochrolechia parella – on siliceous rock on the lake shore, MK 12284 (UGDA); Parmelia saxatilis – on siliceous rock on the lake shore, on rock, MK 12279 (UGDA); Peltigera hymenina – on soil among grasses [det. JM], ATs (GSU); Peltula euploca – MP (SF251635); Phaeophyscia sciastra – on rocks overgrowing mosses, JM 10406 (BILAS); Placidium lachneum – MP (S-F251636); Placopyrenium fuscellum – on siliceous rocks, associated with Placopyrenium canellum, MP (S-251639); Polychidium muscicola – on rocks among mosses, JM 10407 (BILAS), siliceous rocks on the lake shore, on bryophytes growing over rocks, MK 12288 (UGDA); Porocyphus coccodes – siliceous rocks on the lake shore, MP (S-251637); Protoparmeliopsis macrocyclos – MB, RH & ET (O- 20 GRAPHIS SCRIPTA 26 (2014) L184471); Protoparmeliopsis muralis – on rocks [DNA 4237], TF (HBG-015008); Ramalina fastigiata – on rocks [DNA 4228], TF (HBG-015013); Rhizocarpon geographicum – on rocks [spores almost black, welldeveloped, epihymenium brown, K+ very faintly reddish, [DNA 4239], TF (HBG-015006); Rhizocarpon polycarpum – on rocks [DNA 4243], TF (HBG-015007); Rimularia furvella – MB, RH & ET (O-L184479); Rinodina teichophila – siliceous rock on the shore, UA 13247 (LD); Rufoplaca arenaria – on rocks [DNA 4238], TF (HBG-015011); Rufoplaca subpallida – flat siliceous rock above the shore, exposed, UA 13243 (LD); Schaereria fuscocinerea – on rocks [gyrophoric acid by TLC], RH 11910 (O-L190340); Thelopsis melathelia – on moss on north facing wall, JP 47015 (H); Xanthoparmelia delisei – on siliceous rock on the lake shore [glomelliferic, glomellic, perlatolic and gyrophoric acids by TLC], MK 12295 (UGDA); Xanthoparmelia loxodes – siliceous rocks on the lake shore, [glomelliferic, glomellic and perlatolic acids by TLC], MK 12290 (UGDA); Xanthoparmelia stenophylla – siliceous rocks on the lake shore [usnic and salazinic acids by TLC], MK 12276 (UGDA); Xanthoria aureola – on rocks [DNA 4229], TF (HBG015009). Locality 3. Nässja ancient stone circle Nässja parish. 58°27'52"N, 14°48'46"E. An historically interesting area with an ancient stone circle (c. 2000 years old and composed of 24 stones), an avenue, and 12th century church with churchyard. New to the Nordic countries Tremella caloplacae (Zahlbr.) Diederich – on Xanthoria parietina, on calcareous rocks forming the churchyard wall, AM (S-F253105), on X. parietina on Fraxinus inside the churchyard, AM (S-F253109); on Caloplaca decipiens, on calcareous rocks forming the wall next to the churchyard entrance, AM [det. AM and UA] (S-F253110). It is reported from France (NavarroRosinés & Hladun 1996) and the British Isles (Coppins in Lambley 2004: 86). New to Östergötland Bagliettoa calciseda – on top of SW-facing calcareous stone wall surrounding churchyard, JP 47034 (H). Earlier known from scattered occurrences in Fennoscandia. Collected also at locality 1 (Table 1). Cornutispora lichenicola – on Parmelia saxatilis growing on rock [main conidial axes 5–8 µm long, c. up to 2 µm wide, divergent arms not swollen or bulbous at bases], MK 12311 (UGDA). Including the present collection, it is now known from eight provinces of Sweden and two in Norway (Santesson et al. 2004, Alstrup et al. 2008). Marchandiobasidium aurantiacum Diederich & Schultheis – on Physcia tenella (thallus) growing on Fraxinus excelsior, MK 12304 (UGDA). This species was also collected at localities 2, 3 and 8 (Table 1). Tremella ramalinae – On Ramalina fraxinea on Fraxinus, inside the churchyard, AM (SF253111), MW (S-F253170), on R. fraxinea on Fraxinus, next to the stone circle, MK (SF253173). Diederich (1996) mapped T. ramalinae from Mexico and Sweden. Pippola and Kotiranta (2009) reported it new to Fennoscandia with two localities in Åland, southwesternmost Finland. This is the second locality for Sweden, since it had been collected once in the Swedish island Öland in the Baltic Sea. It was collected again on the following day in Vadstena (Table 1). Verrucaria dolosa – on calcareous stone fence surrounding churchyard, JP 47029 (H). It is very sparsely recorded in Sweden and Finland, but not in Norway (Pykälä 2007). Verrucaria inaspecta – on calcareous stone wall surrounding churchyard, N-facing, on pebbles, 21 GRAPHIS SCRIPTA 26 (2014) JP 47028 (H). Earlier reported from the provinces Närke and Västergötland in Sweden and Varsinais-Suomi in Finland (Pykälä & Breuss 2011). Verrucaria infumata – on calcareous stone wall surrounding churchyard, N-facing [together with V. muralis], JP 47027 (H), on calcareous stone wall surrounding churchyard, N-facing, on pebbles, JP 47034 (H). Reported earlier from the province Varsinais-Suomi in southwesternmost Finland (Pykälä 2008). It is also known from the three Swedish provinces Halland, Västergötland and Södermanland, according to Santesson’s list. Additional species collected. Calicium viride – on Quercus robur, ATs (GSU); Calogaya arnoldii ssp. obliterata – UA 13248 (LD); Cliostomum griffithii – on Fraxinus excelsior [det. JM], ATs (GSU); Lecanora orosthea – MB, RH & ET (O-L184480), on rock, MK 12310 (UGDA); Lichenoconium usneae – in the Vouauxiomyces ramalinae-specimen, JM 10399 (BILAS); Lichenomphalia umbellifera – on damp, well decayed wood, JM 10385 (BILAS); Muellerella hospitans – on Bacidia rubella (apothecia) growing on Fraxinus excelsior [det. JM], ATs (GSU); Parmelia sulcata – on Fraxinus excelsior [lobes short, pruinose, rhizines squarrose], MK 12306 (UGDA); Phoma cf. ficuzzae Brackel – on Ramalina fraxinea (thallus and apothecia) growing on Fraxinus excelsior [pycnidia brown in upper part, paler in the lower half, c. 125 µm in diam., conidia 4–5 × 2–3 µm], MK 12309 (UGDA). Recently reported from Lithuania (Motiejūnaitė et al. 2011). If correctly determined, this is the second record in Northern Europe, but since the determination is tentative, we prefer not to list this taxon as new to the Nordic countries. Ramalina baltica – on Fraxinus excelsior, MK 12308 (UGDA); Ramalina fraxinea – on branch of Fraxinus excelsior fallen from the canopy, MK 12307 (UGDA); Ramalina fastigiata – on branch of Fraxinus excelsior fallen from the canopy, MK 12307 (UGDA, in the specimen of R. fraxinea); Ramalina pollinaria – on Fraxinus excelsior at the church [DNA 4226], TF (HBG-015015); Rinodina bischoffii – on calcareous stone in the churchyard wall, VA SE1312 (C); Rinodina exigua – on Quercus robur, ATs (GSU); Vouauxiomyces ramalinae – on thalli of Ramalina fraxinea growing on various deciduous trees, JM 10399 (BILAS); Xanthoriicola physciae – on Xanthoria parietina (apothecia) growing on Fraxinus excelsior, MK 12305 (UGDA). Locality 4. Herrestad church Herrestad parish. 58°23'24"N, 14°48'39"E. Herrestad church is one of the best preserved medieval churches of Sweden. The whole environment, the church built of calcareous stones, gravestones and trees in the churchyard, is very lichen-rich, although no new taxa for the province were observed. Species collected. Calogaya pusilla – UA 13253 (LD); Pyrenodesmia variabilis – UA 13252 (LD); Verrucaria nigrescens – on calcareous rocks, MP (S-F251737); Xanthomendoza poeltii – on bark, VA SE1314 (C). Locality 5. Rök runestone Rök parish. 58°17'42"N, 14°46'32"E. The Rök Runestone, discovered in the 19th century in one of the walls of the local church, is one of the most famous runestones. The epiphytic lichen flora around the parking area and in the churchyard is quite rich, but naturally, there were limited possibilities to collect. New to Östergötland Syzygospora physciacearum – on Physcia aipolia on Syringa, AM (S-F253098). This lichenicolous basidiomycete has a wide world distribution. In the Nordic countries it has been recorded from the southern part of Fennoscandia (Puolasma et al. 2008). An additional specimen was collected in Vadstena in the following day (Table 1). 22 GRAPHIS SCRIPTA 26 (2014) Locality 6. Omberg: Hjässatorget (the parking area) Västra Tollstad parish. 58°18'37"N, 14°38'45"E. Omberg is a 10 km long and 3 km wide mountain in the western part of the province of Östergötland along the eastern side of Lake Vättern. The mountain has been formed over 20 Ice Ages and reaches its highest point of 263 m a.s.l at Hjässan. It is mainly composed of siliceous rock, but sandstone and calcareous rocks appear in some places. The area is characterized by old beeches (Fagus sylvatica), spruces (Picea abies), and planted and naturalised douglas firs (Pseudotsuga menziesii). New to Sweden Cornutispora ciliata Kalb – on Pertusaria amara (soralia) growing on Fagus sylvatica in the forest [main conidial axes 5–7.5 µm long, 2–2.5 µm wide, divergent arms with prominent bulbous bases and very narrow apices], MK 12313 (UGDA). A single locality from Troms in Norway was known in the Nordic area before the present report (Alstrup et al. 2008). New to Östergötland Fuscidea arboricola – on Pseudotsuga menziesii in the forest [fumarprotocetraric acid by TLC; soralia P+ red], MK 13321 (UGDA). This species is known from numerous provinces in Fennoscandia (Tønsberg 1992). Monodictys epilepraria – on Lepraria cf. incana growing on Fagus sylvatica in the forest, MK 12312 (UGDA), on Lepraria cf. incana growing on Pseudotsuga menziesii in the forest, MK 12323 (UGDA). Collected in Osby in Skåne province in 2003 by MK, which was the first record for Fennoscandia (Kukwa & Diederich 2005, Ihlen & Wedin 2008). Parmelia serrana – on the base of Fagus sylvatica at the parking area (Fig. 2), 25 Apr 2013, AT, IK & AN (LD-1637783), AN (UPS-622039). It is also known from the island Rømø in southwestermost Denmark, southwestern Finland and several provinces in southern Sweden, western and central Europe, and the Canary Islands (Thell et al. 2011, Alstrup et al. 2013). Phaeosporobolus alpinus – on Ochrolechia microstictoides (thallus) growing on Betula pendula in the forest [conidia 11–14 × 9–12 µm, consisting of 5–11 cells, cells smooth, 4–5 µm in diam.], MK 12318 (UGDA). The species has a scattered distribution in Fennoscandia (Alstrup & Hawksworth 1990, Santesson et al. 2004). Additional species collected. Biatora efflorescens – on Pseudotsuga menziesii in the forest, MK 12321a (UGDA); Biatoropsis usnearum – on Usnea sp. on the ground, probably originally on Fagus [leg. and det. MW], AM 843 (S); Lecidea nylanderi – on Pseudotsuga menziesii in the forest, MK 12322 (UGDA); Lepraria elobata – with Biatora efflorescens on Fagus sylvatica in the forest [P+ red], MK 12320 (UGDA); Lepraria jackii – on Betula pendula in the forest [atranorin and jackinic acid by TLC], MK12319 (UGDA); Lecanora argentata – on Fagus sylvatica in the forest [amphithecium with large crystals, epithecium without granules], MK 12315 (UGDA); Lichenoconium erodens – on Pertusaria cf. leioplaca, growing on Fagus sylvatica in the forest [pycnidia 50–60 µm in diam., conidia minutely verruculose, brown, 2.5–3 µm in diam.], MK 12314 (UGDA); Loxospora elatina – on a trunk of Pseudotsuga menziesii in the forest, ATs (GSU). This commonly sterile species is characterized by a thin, effuse, whitish grey thallus with convex, irregularly arranged soralia, testing bright yellow with K. Apothecia are very rare and all our specimens were sterile. The species usually grows on shaded acid tree bark, generally in damp or old woodland (Smith et al. 2009). Ochrolechia microstictoides – on Betula pendula in the forest [variolaric acid, lichesterinic acid, microstictoides unknowns by TLC; soralia C+ yellow], MK 12316 (UGDA); Phlyctis argena – on Prunus padus in the forest [thallus K+ yellow turning red, cum apothecia, but without ascospores], MK 12317 (UGDA); Scoliciosporum pruinosum – on Fagus sylvatica near the parking area, UA 13254 (LD). 23 GRAPHIS SCRIPTA 26 (2014) Figure 2. Parmelia serrana – new to Östergötland. Note the position of the isidia, along margins or clustered on ridges. Photo: Einar Timdal Locality 7. Omberg: Hjässan (the viewpoint) Västra Tollstad parish. 58°18'24"N, 14°38'55"E, 263 m a.s.l. Hjässan is at the top of Omberg and offers a marvellous view over the area. It is rich for both saxicolous and corticolous lichens (Fig. 3). New to Östergötland Illosporium carneum – the top of the mountain on Peltigera hymenina growing on saxicolous bryophytes, MK 12328 (UGDA). The species is reasonably widespread in Fennoscandia (Puolasma et al. 2012). Micarea byssacea – on bark of Quercus robur close to the top, P. Czarnota s. n. (H). A recently separated species from Micarea micrococca complex closely related to M. prasina s. str., the type species of the genus. Similarly to M. micrococca s. str. it produces methoxymicareic acid but differs in the Sedifolia grey’ pigment confined to the gel matrix in the epihymenium, hymenium and pycnidial walls, as well as in its larger, adnate and usually lead coloured apothecia (Czarnota & Guzow-Krzemińska 2010), while in M. micrococca s. str. the apothecial section is colourless throughout and ascomata always pale, without a dark pigmentation. Lichenoconium lecanorae – on thalli of Hypogymnia physodes, growing on Pseudotsuga menziesii, together with a specimen of Phoma sp. [conidia subglobose to ovoid hyaline, 4–5 × 3–4 24 GRAPHIS SCRIPTA 26 (2014) µm, with a single large guttule and subglobose, 4–5 µm diam. conidiogenous cells], JM 10418 (BILAS). This speies somewhat resembles Phoma fuliginosa, P. physciicola and P. Caloplaca, but the size and appearance of conidia and conidigeneous cells well as the host differ (Table 1). Figure 3. Some of the participants at Hjässan, Omberg. From the left, standing: Einar Timdal, Tassilo Feuerer, Jon T. Klepsland, Kristiina Mark, Martin Kukwa, Jurga Motiejūnaitė, Staffan Wall, Ingrida Prigodina-Lukošienė, Andrei Tsurykau, Piotr Grochowski, David Galloway, Lars Arvidsson, Hans-Christian Gjerlaug, Orvo Vitikainen, Mark Seaward, Juha Pykälä and Jan Ingar Båtvik. Sitting: Annina Launis, Paweł Czarnota, Mika Bendiksby, Cecilia Hultberg, David Galloway, Martti Rajamäki, Ingvar Kärnefelt, Arne Thell, Maria Prieto, Håkan Lättman and Ulf Larsson. Photo: Jan Ingar Båtvik. Additional species collected. Buellia griseovirens – on Fraxinus excelsior, MK 12327 (UGDA); Cryptothele permiscens – on rock, MW F251739 (S); Lecanora intumescens – on Sorbus aucuparia half way to the top, UA 13255 (LD); Lichenostigma rugosum – on thallus of Diploschistes scruposus, leg JTK [det. JM] 10395 (BILAS); Miriquidica deusta – MB, RH & ET (O-L184484); RH 11895 (O-L190325); Peltigera hymenina – on terricolous mosses, MK 12324 (UGDA); Pertusaria amara – on Fraxinus excelsior, MK 12326 (UGDA); Pertusaria flavida – MB, RH & ET (O-L184487); Protoparmeliopsis macrocyclos – MB, RH & ET (OL184486) (Fig. 4); Rhizocarpon geographicum – [DNA 4230], TF (HBG-015016); Rimularia furvella – RH 11896 (O-L190326). 25 GRAPHIS SCRIPTA 26 (2014) Figure 4. Protoparmeliopsis macrocyclos at Hjässan. Photo: Einar Timdal Locality 8. Vadstena Vadstena parish. 58°27'03"N, 14°53'35"E. The historical town centre of Vadstena, situated at 102 m a. s. l., is amazingly lichen-rich and a large number of different substrates and habitats are available. The participants were encouraged to perform a thorough investigation of the monastery churchyard, where Erik Acharius is buried, for a separate publication (Kärnefelt et al. 2014). New to the Nordic countries Lecanora cf. compallens Herk & Aptroot – on Ulmus cf. montana [usnic acid and zeorin by TLC], MK 12334 (UGDA). See locality 2 (Table 1). New to Sweden Thelidium cf. rimosulum – on concrete block [thallus rimose, medium greenish brown, in some parts with few soredia; perithecia (0.1–)0.15–0.2 mm in diam. in surface view, 3/4 (–1)immersed in thallus; ostiole pale, tiny; involucrellum to exciple base level, c. 30 µm thick, appressed to exciple; exciple c. 0.18 mm in diam., pale; spores (0–)1-septate, 13–16 × 6–7 µm, often constricted at septum], JP 47039 (H). The specimen is tiny, with few perithecia and mixed with Verrucaria spp. Thelidium rimosulum is characterized by the frequent occurrence of isidia and/or soredia. It was described from Poland (Ceynowa-Gieldon 2007) and later reported from Finland (Pykälä 2010a). The Swedish specimen has a similar spore size to the Polish specimens, but the involucrellum is thinner and the exciple pale. 26 GRAPHIS SCRIPTA 26 (2014) Verrucaria ochrostoma – the monastery area, the churchyard, brick wall, on mortar [thallus areolate, grey, c. 0.3–0.5 mm thick, areoles 0.3–1.2 mm; perithecia (0.1–)0.15–0.2 mm in diam. in surface view, immersed in thallus; involucrellum lacking; exciple 0.15–0.22 mm in diam., dark, apex thickened up to 50 µm thick; periphyses c. 25–30 × 1.5 µm; spores (15–)16– 22 × (7–)8–10(–11) µm], JP 47044 (H). Verrucaria ochrostoma was reported new to Fennoscandia from southern Finland by Pykälä (2007). Verrucaria glaucovirens and V. polystictoides have smaller spores, 15–20(–22) × 6–9 µm, and usually a brown thallus. Verrucaria polystictoides – the monastery area, the churchyard, brick wall, on mortar [together with V. muralis], JP 47043 (H). This species had been collected only once in Fennoscandia from East Savonia in Finland (Santesson et al. 2004). New to Östergötland Briancoppinsia cytospora – on thalli of Hypogymnia physodes on trunks of Alnus glutinosa and Sorbus sp., JM 10398 (BILAS). See locality 2 (Table 1). Lepraria vouauxii – on Ulmus cf. montana [with Bacidia rubella; pannaric acid 6-methyl ester by TLC], MK 12336 (UGDA). The species was also collected at locality 1 (Table 1). Lichenochora weillii – on Physconia enteroxantha (thallus) growing on Ulmus cf. montana, MK 12332 (UGDA). This lichenicolous fungus was previously known only from the provinces Skåne and Uppland in Sweden and southwesternmost Finland (Santesson et al. 2004, Puolasma et al. 2008). It is not found in Norway. Marchandiobasidium aurantiacum Diederich & Schultheis – on thalli of Physcia tenella growing on trunks and branches of Sorbus sp. [in the specimen of Syzygospora physciacearum], JM 10372 (BILAS). This lichenicolous fungus species was also found at localities 2 and 3. Syzygospora physciacearum – on thalli of Physcia tenella growing on trunk of Sorbus sp., JM 10372 (BILAS), on thallus of Physcia cf. tenella growing on Ulmus cf. montana, MK 12336 (UGDA), the monastery churchyard, on Physcia tenella on Syringa, AM (S-F253099). An additional sample was collected the day before at locality 5 (Table 1). Tremella ramalinae – on thallus of Ramalina fraxinea growing on Ulmus cf. montana, MK 12340 (UGDA). Collected at locality 3 the day before (Table 1). Trichonectria rubefaciens – the churchyard area, on thalli of Parmelia sulcata growing on Fraxinus excelsior; Vadstena folkhögskola area, on thalli of P. sulcata growing on Ulmus sp. This pathogenic lichenicolous fungus occurs particularly on Parmelia. Occasionally it has been found on Ramalina (Etayo 1998). This species is reported from several European countries, but the reports were rather sparse until recently. However, the new localities presented here, and by Kuznetsova et al. (2012) from northwestern Russia, from Lithuania (Motiejūnaitė et al. 2012), Estonia (Suija 2013) and Belarus (Tsurykau et al. 2013). Verrucaria aff. dolosa – on concrete block, JP47040 (H). Found also at locality 3. Verrucaria memnonia – the monastery churchyard, on concrete block [in the Thelidium cf. rimosulum sample, together with Verrucaria muralis and Acarospora glaucocarpa], JP 47039 (H). The species is sparsely recorded in Fennoscandia, and in Sweden was previously known only from the provinces Bohuslän and Västergötland. 27 GRAPHIS SCRIPTA 26 (2014) Table 1. New species for the Nordic countries, Sweden and Östergötland. Taxon Locality Distribution Voucher and herbarium Bagliettoa baldensis (A. Massal.) Vĕzda 1 New to Ög MP (S-F251633) Bagliettoa calciseda (DC.) Gueidan & Cl. Roux 1; 3 New to Ög JP 47007 (H); JP 47034 (H) Briancoppinsia cytospora (Vouaux) Diederich, Ertz, Lawrey & van den Boom 2; 8 New to Ög JM 10408; 10398 (BILAS) Calogaya arnoldii (Wedd.) Arup, Frödén & Søchting ssp. obliterata (Pers.) Gaya 1 New to Ög UA (LD) Clypeococcum hypocenomycis D. Hawksw. 2 New to Ög VA SE1302 (C) Cornutispora ciliata Kalb 6 New to Sweden MK 12313 (UGDA) Cornutispora lichenicola D. Hawksw. & B. Sutton 3 New to Ög MK 12311 (UGDA) Endococcus exerrans Nyl. 2 New to Ög VA SE1303 (C) Fuscidea arboricola Coppins & Tønsberg 6 New to Ög MK 12321 (UGDA) Illosporium carneum Fr. 7 New to Ög JM s.n. (GSU); MK 12328 (UGDA) Lecanora compallens van Herk & Aptroot 2, (8) New to the Nordic countries MK 12294 (UGDA); (MK 12333 (UGDA)) Lepraria elobata Tønsberg 6 New to Ög MK 12320 (UGDA) Lepraria jackii Tønsberg 6 New to Ög MK 12319 (UGDA) Lepraria vouauxii (Hue) R. C. Harris 1, 8 New to Ög VA SE1305 (C); MK 12336 (UGDA) Leptochidium albociliatum (Desm.) M. Choisy 2 New to Ög MK 12296 (UGDA) Lichenoconium lecanorae (Jaap) D. Hawksw. 2; 7 New to Ög JM 10404; 10418 (BILAS) Lichenochora weillii (Werner) Hafellner & R. Sant. 8 New to Ög MK 12332 (UGDA) Marchandiobasidium aurantiacum Diederich & Schultheis 2; 3; 8 New to Ög 10409 JM; 10372 [in the Syzygospora physciacearum sample] (BILAS); MK 12304 (UGDA) Micarea byssacea (Th. Fr.) Czarnota, Guzow-Krzeminska & Coppins 7 New to Ög PCz s. n. (H) Monodictys anaptychiae (Lindau) D. Hawksw. 1 New to Ög MK 12268 (UGDA) 28 GRAPHIS SCRIPTA 26 (2014) Monodictys epilepraria Kukwa & Diederich 6 New to Ög MK 12312; 12323 (UGDA) Parmelia serrana A. Crespo, M. C. Molina & D. Hawksw. 6 New to Ög ATh, (LD-1637783) Phaeosporobolus alpinus R. Sant., Alstrup & D. Hawksw. 6 New to Ög MK 12318 (UGDA) Placopyrenium canellum (Nyl.) Gueidan & Cl. Roux 2 New to Ög MP (S-F251639) Polycoccum kerneri J. Steiner 2 New to the Nordic countries MK 12286 (UGDA) Porpidia soredizodes (Lamy ex Nyl.) Hertel 1 New to Ög MK 12271 (UGDA) Pyrenidium actinellum Nyl. 2 New to Ög MK 12297 (UGDA) Rinodina turfacea (Wahlenb.) Körb. var. turfacea 2 New to Ög JP 47026 (H) Stereocaulon rivulorum H. Magn. 2 New to Ög MK 12277 (UGDA) Syzygospora physciacearum Diederich 5, 8 New to Ög AM (S-F253098; S-F253099); JM 10372 (BILAS); MK 12337 (UGDA) Thelenella muscorum (Fr.) Vain. 2 New to Ög JP 47024 (H) Thelidium cf. rimosulum Ceyn.Giełd. 8 New to Sweden JP 47039 (H) Tremella caloplacae (Zahlbr.) Diederich 3 New to the Nordic countries AM (S-F253105; S-F253109; SF253110) Tremella phaeophysciae Diederich & M. S. Christ. 1 New to Ög JM 10373 (BILAS); duplicate (SF253103) Tremella ramalinae Diederich 3, 8 New to Ög AM (S-F253111); MW (S-F253170); MK (S-F253173); MK 12340 (UGDA) Trichonectria rubefaciens (Ellis & Everh.) Diederich & Schroers 8 New to Ög – seen and determined by JM, MK & ATs Verrucaria dolosa Hepp 3 (8) New to Ög JP 47029 (H); 47040 (H) Verrucaria inaspecta Servít 3 New to Ög JP 47028 (H) Verrucaria infumata Nyl. 3 New to Ög JP 47027, 47031 (H) Verrucaria memnonia (Körb.) Arnold 8 New to Ög JP 47039 (H) [in the Thelidium cf. rimulosum sample] Verrucaria ochrostoma (Borrer ex Leight.) Trevis. 8 New to Sweden JP 47044 (H) Verrucaria polystictoides Vain. 8 New to Sweden JP 47043 (H) 29 GRAPHIS SCRIPTA 26 (2014) Additional species collected. Acarospora glaucocarpa [in the Thelidium cf. rimosulum sample, together with Verrucaria memnonia and V. muralis], JP 47039 (H); Calicium salicinum – on Ulmus cf. montana [ascospores 10–12 × 4.5–5 µm], MK 12341 (UGDA); Candelaria pacifica – the castle area, on trunks of Ulmus sp., with Polycauliona candelaria, MK 12343 (UGDA), on Tilia at the red brick (old church) tower in the town centre, 15 Oct 2009, TF (HBG-015002). The species was reported from Östergötland by Westberg and Arup (2011); Candelariella efflorescens – on Ulmus cf. montana [confirmed by MW], MK 12329 (UGDA); Chaenotheca phaeocephala – MB, RH, AL & ET (O-L187265); Chaenotheca trichialis – the monastery churchyard, brick wall, on mortar, JP 47050 (H); Chrysothrix candelaris – on Ulmus, 15 Oct 2009, TF (HBG-015001); Diplotomma alboatrum – the monastery churchyard, brick wall, on mortar, JP 47045 (H); Hypocenomyce scalaris – MB, RH, AL & ET (O-L184485); Lathagrium fuscovirens – MB, RH, AL & ET (O-L187266); Lecanora albescens – the monastery churchyard, brick wall, on mortar, together with Verrucaria savonensis [?], JP 47043 (H), together with Diplotomma alboatrum, JP 47045 (H); Lecanora chlarotera – at the monastery church, growing on Fraxinus excelsior, MK 12257 (UGDA, specimen of Vouauxiella lichenicola); Lepraria incana – on Ulmus cf. montana [divaricatic acid, zeorin by TLC], MK 12335 (UGDA); Ochrolechia subviridis – at the monastery church, on Fraxinus excelsior, MK 12256 (UGDA); Ochrolechia turneri – on Ulmus cf. montana, MK 12333, 12342 (UGDA); Pachyphiale fagicola – on Ulmus cf. montana together with Caloplaca cf. obscurella, MK 12338 (UGDA); Pertusaria albescens – MB, RH, AL & ET (O-L187268); Pertusaria coccodes – on Ulmus cf. montana, MK 12331 (UGDA); Phoma sp. – on Evernia prunastri (thallus) growing on Ulmus cf. montana [Conidia narrowly ellipsoidal, 6–8 × 1.5– 2 µm, conidiogenous cell 5 × 1.5 µm], MK 12339 (UGDA); Physconia grisea – on Ulmus cf. montana together with Melanelixia subargentifera, MK 12330 (UGDA); Physconia perisidiosa – on Ulmus cf. montana together with Melanelixia subargentifera, MK 12344 (UGDA); Protoparmeliopsis muralis – the monastery churchyard, on Ulmus sp., ATs (GSU); Ramalina fraxinea – Vadstena folkhögskola, on Tilia [DNA 4225], TF (HBG-015003); Thelidium cf. rimosulum – the monastery churchyard, on concrete block [together with Verrucaria memnonia, V. muralis and Acarospora glaucocarpa], JP 47039 (H); Thelomma ocellatum – on lignum, VA 1313 (C); Verrucaria muralis – the monastery churchyard, on concrete block [in the Thelidium cf. rimosulum sample, together with V. memnonia and Acarospora glaucocarpa], JP 47039 (H); Verrucaria viridula – the monastery churchyard, brick wall, on mortar, JP 47049 (H); Vouauxiella lichenicola – at the monastery church, on Lecanora chlarotera growing on Fraxinus excelsior, MK 12257 (UGDA); Xanthomendoza fulva – on Aesculus hippocastanum at the castle [DNA 4227], on Ulmus sp. at the castle, TF (HBG-015017), ATs (GSU). Acknowledgments Our sincere thanks to Göran Söderström and Axel Unnerbäck who arranged a historical guided tour, invited us to a banquet in their home (Acharius’s house) and for encouragement in general, to Anita Gustavsson (Vadstena Folkhögskola), to Göran Åström (in charge of the churchyards in Vadstena), and to Vadstena Buss, the local bus company. Magdalena Oset (University of Gdańsk) is thanked for the determination of Stereocaulon rivulorum. We are greatly indebted to our sponsors: Letterstedt Foundation [ Letterstedtska Föreningen], Stockholm, Ove Almborn and Elly Olsson Foundations administrated by Lund University, and the Biological Museum, Lund University. 30 GRAPHIS SCRIPTA 26 (2014) References Alstrup, V. 2004. New records in distribution of lichens and lichenicolous fungi. Graphis Scripta 16: 46–57. Alstrup, V. & Ahti, T. 2007. New reports of lichenicolous fungi, mainly from Finland and Russia. Karstenia 47: 1–4. 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