Flora and vegetation of the southern Carnarvon Basin, Western Australia

Allan Burbidge

Lists of the vascular flora of the Yelverton and Witchcliffe State Forests and the Bramley and Forest Grove National Parks from the Margaret River Plateau of south-west Western Australia are provided for the first time. A combined list of 731 taxa (87 weeds) was recorded from these areas. A total of 520 vascular plant taxa (490 native and 30 weeds) have been recorded from Yelverton Conservation Reserves, 448 vascular plant taxa (388 native and 60 weeds) from Bramley National Park, 351 vascular plant taxa (315 native and 36 weeds) from Witchcliffe State Forest and 363 vascular plant taxa (307 native and 56 weeds) from Forest Grove National Park. No native species appear to be endemic to the Plateau. No declared rare species were found, although 23 priority taxa were recorded from these reserves. Numerous disjunct and geographically significant populations are known from these parks.

The vegetation of Narran Lake Nature Reserve (4527 ha) in the central north of New South Wales approximately 30 km west of Cumborah (29°43', 147°29') in the Walgett Shire on the North Western Plains and the Darling River Plains Bioregion is described. Seven communities are defined based on flexible UPGMA analysis of cover-abundance scores of all vascular plant taxa. These communities are mapped based on ground truthing, air photo interpretation and geological substrate. All communities are simple in structure being primarily woodlands, shrublands and herbfields. Communities are: 1) Mixed Low Woodlands, 2) Mulga Low Woodlands, 3) Triodia Hummock Grasslands, 4) Chenopod Low Open Shrublands, 5) Ephemeral Herbfields, 6) Riparian Open Forests, and 7) Lignum Shrubby Thickets. A total of 325 taxa were recorded including two species listed under the NSW Threatened Species Conservation Act (1995), Lepidium monoplocoides and Goodenia macbarroni. An additional 11 species are considered to be at their geographic limit or disjunct in their distribution; 11% are exotic in origin.

Naree and Yantabulla stations (31,990 ha) are found 60 km southeast of Hungerford and 112 km northwest of Bourke, New South Wales (lat. 29° 55'S; long. 150°37'N). The properties occur on the Cuttaburra Creek within the Mulga Lands Bioregion. We describe the vegetation assemblages found on these properties within three hierarchical levels (Group, Alliance & Association). Vegetation levels are defined based on flexible UPGMA analysis of cover-abundance scores of all vascular plant taxa. These vegetation units are mapped based on extensive ground truthing, SPOT5 imagery interpretation and substrate.

DOI: 10.18195/issn.0313-122x.61.2000.077-154 Records the Western Australian Musellm Supplement No. 61: 77-154 (2000). or Flora and vegetation of the southern Carnarvon Basin, Western Australia G.J. Keighery, N. Gibson, M.N. Lyons and Allan H. Burbidge Department of Conservation and Land Management, PO Box 51, Wanneroo, Western Australia 6065, Australia Abstract This paper reports the first detailed study of the vascular flora of the southern Carnarvon Basin, an area of c. 75000 km 2 • A total flora of 2133 taxa of vascular plants was listed for the area. There are eight major conservation reserves which have 1559 taxa present in them. Most of the 574 unreserved taxa are wetland taxa, taxa of tropical affinities or those only present on the Acacia shrublands of the central basin. Vegetation patterning at a regional scale showed the major floristic boundary in the south west of the study area, which in turn reflected the major climatic gradients of the area. The other major influence on vegetation patterning was soil type. INTRODUCTION Despite Shark Bay being the site of very early visitation and study by several European expeditions (Beard, 1990; Keighery, 1990; George, 1999) the area was until recently still poorly known botanically. Beard (1975, 1976a) prepared structural vegetation maps for the whole area at a 1: 1 000000 scale and Payne et al. (1987) have undertaken land system maps (rangeland mapping) for the whole area at a 1: 250000 scale. More localised areas have been mapped and published for the Kalbarri-Shark Bay area by Beard (1976b, 1976c) at a scale of 1: 250000. Limited floristic surveys have been undertaken in the north of the study area around Lake MacLeod by Tyler (1988), Rocky Pool near Carnarvon by Kenneally (1978) and Cape Range by Keighery and Gibson (1993). Other localised studies have involved the Shark Bay World Heritage Area by Trudgen and Keighery (1995), Bernier and Dorre Islands by Royce (1962), Dirk Hartog Island by Burbidge and George (1978), Kennedy Range by Newbey and Newbey (1992) and Toolonga Nature Reserve by Burbidge et al. (1980). The present work is part of a multi-disciplinary study to sample the biodiversity of a 75000 km 2 region of Western Australia (the southern Carnarvon Basin) and identify biophysical factors related to the observed patterns of species distribution (Burbidge et al., 2000). The aim of this paper is to describe the flora and vegetation of the major landforms sampled as biodiversity sites and to discuss the composition, biogeographic patterns and conservation status of the vascular plants of the study area. Related papers in this volume deal with the wetIand flora (Gibson, Keighery and Lyons, 2000) and the flora of the temperate - arid change-over zone (Gibson, Burbidge, Keighery and Lyons, 2000). METHODS Study area The study area covered by the flora survey extended from 23°30'S to 28°00'S and from Dirk Hartog Island and Shark Bay in the west to 115°30'E (Figure 1). The southwest of the study area encompasses the change over from the South West Botanical Province to the Eremaean Province and hence is an area of major vegetation change. Almost the entire study area falls within the Carnarvon and Irwin botanical districts of Beard (1990). These districts largely correspond to two IBRA regions Geraldton Sandplain and Carnarvon (Thackway and Cresswell, 1995). Field sampling and analysis Sixty-three primary sites were sampled across the study area, stratified to cover the geographic range of the major surface types of the Basin. Sites were selected such that three to five different surfacetypes could be sampled in close proximity to facilitate fauna sampling (Figure 1). Sites 30 m x 30 m were established at each site and all vascular plants occurring in the sites were recorded. Sites were visited twice, in August 1994 and August 1995. Some additional recordings were made when the sites were visited at other times in conjunction with fauna sampling. Sites were classified according to similarities in species composition (presence / absence data) using the Czekanowski coefficient and 'un weighted pair-group mean average' fusion method (UPGMA, Sneath and G.]. Keighery, N. Gibson, M.N. Lyons, A.H. Burbidge 78 /' I.' 0 ~~'~k )j 't)j'J ( ~._-~I' 1130 14 ) / 't:--< 1150 . ~ ('- 24<>- '-~ f6 Lake (\ CU A3 Mac~ J 5 11r: 4 j.M&5 1 I 5 4 3 ~2 BO MR I la Kennedy Range National Park Camarvon Bemier Island Nature Reserve 25 0 Dorre Island Nature Reserve Shark Bay World Heritage Area Pimbee 5 4 3 2 1a"1 WO 25 50 Kilometres Existing reserves Timber Reserve CALM Leasehold la Study sites World Heritage Area Boundlll)' 1140 Figure 1 Location of 63 primary sampling sites and major conservation reserves in the study area, Flora and vegetation 79 Soka!, 1973), while species were classified using the 'two step' method of Austin and Belbin (1982) and the UPGMA fusion method. Only taxa identified to species level were included in the analysis (>95'J;, records). Annuals and geophytes were included in the analysis because all sites but one (NE2) were considered to have had good rains prior to at least one of the sampling periods. A bulked soil sample (consisting of 24 regularly spaced subsamples across the site) of the top 10 cm was collected from each site, and soil mechanical analysis and macro nutrient analyses were undertaken on these samples [see McArthur (1991) for details]. Eleven climatic parameters for each site were estimated from the climate modelling program ANUCLIM (McMahon et al., 1995). Statistical relationships between site groups were tested using Kruskal - Wallis non-parametric analysis of variance and Mann Whitney U-tests (Siege!, 1956). In addition to the 63 primary sites, flora lists were available from 75 extra sites in the southern part of the study area (Gibson, Burbidge, Keighery and Lyons, 2000) and from the wetland survey (Gibson, Keighery and Lyons, 2000). Additional sampling efforts in six mainland reserves were also undertaken to compile comprehensive species lists for the eight major reserves (Figure 1) in the study area. These lists also include previously published surveys and collections held in the Western Australian Herbarium (PERTH). The total known vascular flora of the study area (Appendix 1) was compiled from the current and previous surveys, and from collections held in PERTH. Nomenclature generally follows Green (1985) and current usage at PERTH. Over 2200 voucher specimens have been lodged in PERTH. RESULTS The results are given in two sections. The first deals with the overall vascular flora recorded for the study area, its composition and conservation. It provides a floristic context for the interpretation of the second part, which deals with quantitative patterns in the floristic composition of the biodiversity sites and how this patterning relates to the biophysical attributes of the study area. Flora Composition A total of 2133 taxa of vascular plants from 122 families were recorded from the study area (Appendix 1). These comprised 9 ferns or fern allies, 2 gymnosperms and 2122 flowering plants (of which 88 were naturalised aliens). The largest families were the Myrtaceae (231 taxa, 10.8% of the flora), Asteraceae (181 taxa, 8.5%), Poaceae (168 taxa, 7.9'Yo), Chenopodiaceae (106, 5.0%), Papilionaceae (105, 4.9'?{J), Mimosaceae (95, 4.5%), Proteaceae (93, 4.4%), Goodeniaceae (71, 3.3%), Cyperaceae (61, 2.9%), Orchidaceae (57, 2.7%), Amaranthaceae (50, 2.3°/.,), Myoporaceae (50, 2.3%) and Malvaceae (48, 2.3%). Together these 13 families comprise 51.7% of the total flora. Of the remaining 109 families, only 12 contained at least 20 species: Euphorbiaceae (36 taxa), Brassicaceae (32), Solanaceae (32), Anthericaceae (28), Zygophyllaceae (26), Apiaceae (22), Epacridaceae (22), Sterculiaceae (22), Boraginaceae (21), Chloanthaceae (21), Haemodoraceae (20) and Portulacaceae (20). These 12 families combined contributed 14.2% of the total flora. Several significant collections were made during the survey, including the re-collection of Rumex crystallinus (the first record since 1877). Three apparently new taxa were first collected during the survey: Paracaleana lyonsii ms (Orchidaceae), Calandrinia sp. Coolcalalaya (GK and NG 698) (Portulacaceae) and Eremophila glabra subsp. Zuytdorp (GK and NG 518) (Myoporaceae). The broad composition of the flora reflects the major climatic influences affecting the study area. There is a southern coastal winter rainfall zone and an inland and northern arid zone. The whole study area is still dominated by winter rainfall (Wyrwoll, Courtney and Sandercock, 2000). Immediately north of the study area, summer rainfall begins to dominate. The majority of the study area is arid and forms part of the Eremaean Botanical Province (Beard, 1975, 1976a). Thi5 pattern is also clearly seen in the results for the vegetation analysis (see below). This strong desert influence is clearly demonstrated in the composition of the flora, with arid zone taxa dominating in six of the speciose families (Amaranthaceae, Asteraceae, Goodeniaceae, Malvaceae, Myoporaceae and Poaceae) and in nearly all the smaller families (Euphorbiaceae, Brassicaceae, Solanaceae, Zygophyllaceae, Apiaceae, Sterculiaceae, Boraginaceae, Chloanthaceae, and Portulacaceae). These desert influences are from predominantly temperate or winter rainfall semi-arid or arid areas, rather than a tropical desert flora. This can be illustrated using the Asteraceae, the largest family recorded from the study area. The family has 492 native taxa present in Western Australia, of which 181 were recorded from the study area. Species richness in the family is greatest in the Avon-Wheatbelt and Coolgardie IBRA regions (areas of Mediterranean and arid-Mediterranean climate) of Western Australia (Figure 2). The family, though spread widely through the arid zone (providing the spectacular wildflower displays of the Acacia shrublands) and tropics, is species rich in temperate Western Australia. The Asteraceae of the study area are a reflection of the above pattern, being comprised mainly of G.]. Keighery, N. Gibson, M.N. Lyons, A.H. Burbidge 80 ericoid Myrtaceae, especially members of the genera Scholtzia, Baeckea and Pileanthus. There is also a considerable number of eucalypt and orchid taxa, especially on heavy soils in the Kalbarri area. The Haemodoraceae, Epacridaceae (Keighery, 1996) and the genus Acacia (Hopper and Maslin, 1978) are also species diverse on the Kalbarri sandplains. Weeds Figure 2 Species richness patterns of the native Asteraceae of Western Australia. annuals (52 taxa) that are widespread in the winter rainfall areas of the semi-arid and arid zone. There are also 20 taxa of the temperate zone that end their ranges in the Basin (not necessarily in the study area, e.g. Brachycome iberidifolia, Chthonocephalus pseudevax and Cratystylis subspinescens) and 13 tropical taxa that are at their southern limits (e.g. Blumea tenella, Launaea sarmentosa, Pluchea rubellifiora and Streptoglossa bubakii). There is also a component of 12 taxa that are either endemics or species centred on the Basin (e.g. Chthonocephalus oldfieldianus, Haptotrichion colwillii, Rhodanthe cremea and Rhodanthe oppositifolia subsp. ornata). Many of these are recently described or still unnamed. The pattern shown in the Asteraceae is repeated in the Poaceae (Keighery, 1984), Chenopodiaceae (Wilson, 1984), Amaranthaceae and Goodeniaceae (Carolin et al., 1992). The desert elements of the flora are largely of southern affinities rather than arid tropical, as would be found further north. The other major influence is that the study area contains a portion of the northern heathlands of the South-West Botanical Province, a large section of desert to the north and east, and the interzone between them (Beard, 1990). The northern heathlands are diverse in members of the Proteaceae, especially Grevillea (Gibson et al. 1997), and the In total, 88 species of naturalised vascular plant species (weeds) were recorded for the area. These weeds are nearly all annuals from the families Poaceae (41 taxa), Asteraceae (19), Brassicaceae (12) and Caryophyllaceae (10). This is a low number compared to other areas of Western Australia (Keighery, 1995), where some 1032 taxa are recorded as naturalised (ca. 9% of the total flora). But it is comparable to Cape Range, where 30 weeds were recorded by Keighery and Gibson (1993) and the Kimberley where 184 taxa have been recorded (Hussey et al., 1997). The largely intact nature of the native vegetation of the study area lowers the number of weeds. It is likely that further collecting around the townsites of Kalbarri and Carnarvon will increase the number of naturalised taxa, but it is considered that the major environmental weeds have been documented. Certain weeds have already had a severe impact on parts of the study area. Foremost are riverine sites, especially along the Gascoyne, Wooramel and Irwin Rivers where the fringing understorey vegetation is often dominated by Buffel Grass (Cenchrus ciliaris), Mosman River Grass (Cenchrus setigerus) and Castor Oil Plants (Ricinus communis). Numerous herbaceous weeds such as Cape Weed (Arctotheca calendula) dominate the Murchison River foreshore in Kalbarri National Park, probably due to past grazing. The other major areas of weed invasion are under the Acacia shrublands of Zuytdorp and Cooloomia National Park where Brassica tournefortii has replaced Parietaria debilis, after sheep, rabbit and now goat grazing pressure. There are similar, but more localised problems in the Shark Bay World Heritage Area. Several other weeds are potentially serious. Patterson's Curse (Echium plantagineum) is spreading along road verges through the southern arid zone, including North West Coastal Highway, and has the potential to replace the mass displays of everlastings in these areas. Alford et al. (2000) have reported Boxthorn (Lycium ferocissimum) on islands in the Shark Bay World Heritage area. This species is already a major pest on other offshore islands. Athel Pine (Tamarix aphylla), a serious weed of the Finke River system in the Northern Territory, is spreading along the Gascoyne River at Camarvon. I 81 Flora and vegetation Site Groups bbl 002 nal bb3 cul pel 1-I I ho4 hoS 1Ild1 1Ild2 001 003 wo3 woS wo4 1Ild4 wo1 wo2 na2 pt!4 I - I cu6 1- I J-I -_.1 I -I---I 2c 3.. I JI I 2b I I I J I I I I 2.. 1 I I 1 I I I I1 I I 3b II I 3c 3d 1 1--1-1--1-- I 1--1I I I-I I I 1--1 LI I I I-I --1- I I I 1 I I 1 I I I I-I I 1 1 I 4.. I I 4b II 1 I I I I I I I 1-1I I I I I I I I -- kel mr3 ke2 5 1I 6 I I I 1- ne2 n..3 na4 naS zu1 zu2 zu4 zuS zu3 I I I I 1 I I I I I I I I-I 1 I 1I I 1-1--1I bbS ne1 ne3 ne4 neS II 1--1--1 IIldS gj1 ke3 1<,,4 gj2 gj3 gj4 gjS mr1 keS mr2 mrS mr4 I 1I-I I 11 I 1I I 1I I I I 1- IIld3 cu2 cu3 cu4 cuS ,,11 e12 pe2 peS pe3 I I I bb2 bb4 1 1- 1I --I I I I I 0.30 I 0.48 I 0.67 I 7 I I I I I-I I I I I I I 0.86 I 1.04 8 9.. 1--1 9b I 1. 23 Dis·similarity Figure 3 Dendrogram of the nine major site groups defined by the classification of the vascular flora. I l-. 82 G.]. Keighery, N. Gibson, M.N. Lyons, A.H. Burbidge Table 1 Number of taxa recorded for the eight major reserves of the study area. Number of taxa Reserve Kalbarri National Park Toolonga Nature Reserve Zuytdorp National Park Cooloomia Nature Reserve Francois Peran National Park Bemier Island Nature Reserve Dorre Island Nature Reserve Kennedy Range National Park 1071 200 314 226 279 59 143 314 Reservation status of species There are eight major conservation reserves in this area (Figure 1). A major aspect of the present study was to attempt to produce an indication of the reservation status of individual plant taxa in the study area. Species lists for all eight major reserves were compiled. The flora of the numerous small island nature reserves in the Shark Bay World Heritage Area have been compiled separately (Alford et al., 2000; A.A. Burbidge pers. comm.). Total numbers of taxa recorded for the eight major reserves are shown in Table 1 with full lists in Appendix 1. These parks and reserves contained a total of 1559 taxa leaving 574 taxa (37% of the total) that were not recorded for any conservation area (Appendix 1). These unreserved taxa were composed of six distinctive groups. They were: • Marine angiosperms (Cymodocea, Halophila and Posidonia). These taxa are largely covered by marine reserves proposed for the W orId Heritage Area. • Wetland taxa of tropical or arid affinities (e.g. Peplidium species, Potamogeton species, Schoenoplectus species, Cyperus difformis). These species occur in the northern sector of the study area, around Lake MacLeod, major rivers and in springs near the Kennedy Range. • Terrestrial taxa of tropical affinities, especially those occurring on clay or alluvial soils that penetrate the study area in the north, usually along the major rivers or drainage lines (e.g. Crinum flaccidum) or on coastal limestones (Grevillea variifolia). • • Red sand taxa confined to the Acacia shrublands of the central region of the study area. Species confined to freshwater claypans (e.g. Lythrum sp. Towrana (Cranfield 2183) and Rumex crystallinus) • Endemics of the World Heritage Area. There are 45 endemics of the Shark Bay World Heritage Area that are largely found on Tamala and Nanga Stations. Although these are not yet in a conservation reserve, current proposals _ regarding the property (Anonymous, 1997) will see nearly all of these placed in an expanded Zuytdorp Nature Reserve. Rare and Priority Flora: There are 11 species of declared rare flora and 144 taxa of priority flora recorded in the study area (Table 2; CALM, 1998). Of these priority taxa 33 are PI, 62 are P2, 39 are P3 and 10 are P4. (See Appendix 1 for definitions of conservation codes). Of the declared rare species, eight are confined to riverine cliffs or heavy soils in the Kalbarri area (Beyeria lepidopetala, Caladenia bryceana subsp. cracens, Caladenia wanosa, Drakaea concolor, Eremophila microtheca, Hypocalymma longifolium, Lechenaultia chlorantha and Leucopogon marginatus). New records and range ends Numerous new records and range extensions were noted during the survey. One notable example was the recording of Eremophila occidens ms at Nanga, this species having been previously regarded as endemic to Cape Range. Taxa endemic to all or part of the study area and taxa at their northern or southern limits in the study area are annotated in Appendix 1. Most notable is the listing of 223 taxa at their northern limit in Kalbarri National Park, being over 20% of the Park's known flora. This is a comparable figure to another conservation area with numerous range ends, the Stirling Range. Another 229 taxa end their range within the World Heritage area to the north of Kalbarri (Trudgen and Keighery, 1995). This is further evidence that the area is a major transition zone for the vascular flora of Western Australia. Vegetation patterning A total of 626 taxa were recorded from the 63 sites (over 25% of the known flora of the study area). Of these 626 taxa, 218 (35%) were recorded in only one site and were excluded because preliminary analysis of the total dataset indicated that these 'singletons' added little information. The final dataset included 408 taxa from 63 sites. Species richness varied from nine to 66 taxa per site (with singletons excluded), with individual taxa occurring in between two and 40 sites. In this analysis site groups are partitioned at the nine group level (Figure 3) and species assemblages at the 20 group level (Table 3). Some site groups showed distinct pattern below this level and these were designated as subgroups. In all, 15 sites groups and subgroups were defined. The major floristic boundary in the southwest of the study area is clearly evident in the analysis with site groups 1 to 6 representing typical Carnarvon vegetation types and site groups 8 and 9 -~-~~~---------------------------------- F lo raandv eg e ta t ion 83 J1 3 " 2 4 ° t B e r n i e rI 2 5 ° DO r r e l l 27 ' '< F: C l 1 1 50 F igu r e4 D i s t r ibu t iono fth en in em a jo rs i t eg roup sa c ro s sth es t u d ya a~ o o 84 G.]. Keighery, N. Gibson, M.N. Lyons, A.H. Burbidge Table 2 Rare and Priority Flora recorded for the study area. [ms indicates a manuscript name currently used by Western Australian Herbarium (PERTH); see Appendix 1 for definitions of conservation codes]. Conservation Code Taxon Beyeria lepidopetala Caladenia bryceana subsp. cracens ms Caladenia hoffmanii ms Caladenia wanosa Drakaea concolor rns Drakonorchis barbarella ms Eremophila microtheca Eucalyptus beardiana Hypocalymma longifolium Lechenaultia chlorantha Leucopogon marginatus R R R R R R R R R R R Beyeria gardneri Chamelaucium oenanthum rns Chthonocephalus oldfteldianus Chthonocephalus spathulatus Eremophila cuneata rns Eremophila lanata rns Eremophila physocalyx ms Labichea eremaea Lepidium scandens Levenhookia octomaculata Macarthuria georgeana Malleostemon sp. Hardabutt Rapids (D.Bellairs 1 1 1 1 1 1 1 1 1 1 1 Millotia depauperata Persoonia brachystylis Persoonia papillosa Pityrodia axillaris Ptilotus beardii Ptilotus stipitatlls Ptilotus stirlingii var. pumilus Rhodanthe sp. Overlander (P.5.5hort 2096) Schoenia ftlifolia subsp. arenicola Scholtzia cordata ms Scholtzia sp. Eurardy O.5.Beard 6886) Scholtzia sp. Folly Hill (M.E.Trudgen 12097) Scholtzia sp. Murchison River (AS.George 7098) Sclerolaena stylosa Tetragonia coronata Thryptomene ninghanensis rns Thryptomene sp. Carrarang (M.E.Trudgen 7420) Thryptomene sp. Steep Point (M.E.Trudgen 7421) Thryptomene sp. Tamala (M.E.Trudgen 7384) Verticordia lepidophylla var. quantula Vittadinia cervicularis var. oldfteldii 1 1 1 1 1 1 1 1654A) Abutilon sp. Hamelin (AM.Ashby 2196) Abutilon sp. Quobba (H.Demarz 3858) Acacia gelasina Acacia leptospermoides subsp. obovata Acacia stereophylla var. cylindrata Acacia subrigida Acanthocarplls parviflorus Angianthus microcephalus Anthotroche myoporoides Baeckea subcuneata Bergia auriculata Beyeria cygnorum Calytrix drummondii Calytrix formosa Calytrix harvestiana 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 2 2 2 2 2 2 2 2 2 2 2 2 2 2 2 Taxon Conservation Code Calytrix paucicostata Calytrix purpllrea Chamelallcium marchantii Chthonocephalus muellerianlls Chthonocephalus tomentellus Cryptandra glabriflora Cryptandra scoparia var. microcephala Dampiera krauseana Eremaea acutifolia Eremophila glabra subsp. psammophora ms Eremophila occidens ms Eucalyptus diminuta ms Frankenia confusa Grevillea annulifera Grevillea costata Grevillea rogersoniana Grevillea stenomera Grevillea triloba Hemiandra sp. Kalbarri (D.Bellairs 1505) Malleostemon sp. Cooloomia (S.D.Hopper 1353) Malleostemon sp. Kalbarri (L.ACraven 7083) Malleostemon sp. Moonyoonooka (RJ.Cranfield 2947) Melaleuca huegelii subsp. pristicensis Microcorys tenuifolia Millotia jacksonii Murchisonia fragrans Olearia occidentissima Pityrodia glutinosa Platysace sp. Kalbarri (D.& B.Bellairs 1383) Platysace sp. Kennedy (P.G.Wilson 8450) Ptilotus alexandri Rhodanthe oppositifolia subsp. ornata Rumex crystallinus Scaevola chrysopogon Schoenus sp. Kalbarri (K.RNewbey 9352) Scholtzia sp. East Yuna (AC.Burns 6) Scholtzia sp. Galena (W.E.Blacka1l4728) Scholtzia sp. Ross Graham Lookout (S.Maley 6) Scholtzia sp. Z-Bend (Bellairs-Kalflora 912a) Sondottia glabrata Thryptomene sp. Eagle Gorge (AG.Gunness 2360) Thryptomene sp. Eurardy (D.& B.Bellairs) Thryptomene sp. Junga Dam (AG.Gunness 2383A) Thryptomene stenophylla Thysanotus kalbarriensis ms Verticordia dasystylis subsp. kalbarriensis Verticordia galeata Acacia didyma Acacia drepanophylla Acacia isoneura subsp. nimia ms Acacia plautella rns Acacia sclerosperma subsp. glaucescens Anthocercis intricata Anthotroche walcottii Arnocrinum dnlmmondii Calocephalus aervoides Calycopeplus marginatus Centrolepis cephaloformis subsp. murrayi Chamelaucium conostigmum ms Comesperma acerosum 2 2 2 2 2 2 2 2 2 2 2 2 2 2 2 2 2 2 2 2 2 2 2 2 2 2 2 2 2 2 2 2 2 2 2 2 2 2 2 2 2 2 2 2 2 2 2 3 3 3 3 3 3 3 3 3 3 3 3 3 Flora and vegetation Table 2 85 (cont.) Taxon Conservation Code Cryptandra nudiflora Dicrastylis linearifolia Eucalyptus foecunda subsp. Coolimba 3 3 Geleznowia verrucosa Goodenia sericostacfzya Grevillea candical1s Grevillea leptopoda Grevillea stel10stacfzya Gymnantfzera cllnningfzamii Hemigel1ia pimelifolia Lasiopetalum oldfieldii subsp. oldfieldii Lasiopetalum oppositifolium Lepidobolus densus ms Macartfzuria intricata Mirbelia sp. Kalbarri (MoO.Crisp 6261) Petropfzile biternata Pfzysopsis cfzrysopfzylla Pfzlegmatospermum drllmmondii 3 (M.I.H.Brooker 9556) 3 3 3 3 3 3 3 3 3 3 3 3 3 3 3 representing typical Irwin (Geraldton Sandplains) vegetation types (Figures 3 and 4). Site group 7 is represented by only a single site and its affinities are not clear. Site groups 1 to 4 were the most widespread across the study area. Site group 1 comprised the sites with a major saline influence. Species assemblage 0 is highly faithful (i.e. largely restricted) to this site group with low to moderate levels of constancy (referring to the degree of occurrence of a species assemblage in the site group). Species in this assemblage include Halosarcia indica and H. halocnemoides (samphire species), Frankenia aff. pauciflora (a species generally restricted to saline calcareous substrates) and a series of annual taxa which are restricted to this site group. This group encompasses a range of saline plots ranging from Halosarcia and Sclerostegia shrublands to an Atriplex shrubland upslope from a saline flat. Sites occurred in coastal areas from Nanga to Lake MacLeod (Figure 4) and species richness averaged 23.7 taxa/ plot. Acacia shrublands are one of the most widespread and typical communities of the Carnarvon Bioregion. Site groups 2 and 4 represent the Acacia shrublands on red sands. No single Acacia species dominated sites in site group 2 but Acacia sclerosperma and A. linophylla were the most common; sites in site group 4 were generally dominated by Acacia aneura (Mulga). Site group 2 was a very widespread group in the centre of the study area, stretching from Nanga to Boolathana and east to Meedo (Figure 4). It had the highest average species richness of 46.6 taxa / plot. While most sites in site group 2 were dominated by Acacia spp., one site along the Wooramel River Taxon Conservation Code Scaevola oldfieldli ScllOltzia sp. Ajana (T.A.Halliday 137) Stenantfzemum divaricatum Verticordia cooloomia Verticordia densiflora var. roseostella Verticordia dicfzroma var. dicfzroma Verticordia dicfzroma var. syntoma Villarsia congestiflora 3 3 3 Calotfzamnus pacfzystacfzyus Diuris recurva Goodenia neogoodenia JackBonia velutina Lepidium puberulllm Lepidosperma rupestre Plectracfzne bromoides Verticordia capillaris Verticordia polytricfza Wllrmbea murcfzisoniana 4 4 4 4 4 4 4 4 4 4 3 3 3 3 3 (MD3) had an overstorey of Eucalyptus victrix while the understorey was typical of the Acacia shrublands. Another site (B01) was dominated by Atriplex, it differed from sites in site group 1 in being rich in species belonging to assemblage K (especially in terms of herbs), an assemblage largely lacking in the site group 1. This site was classified most similar to B03 (a typical Acacia shrubland). Species assemblage A was moderately faithful to both site group 2 and site group 4, while species assemblages 0, E, and J differentiated site group 2 from other site groups. Species assemblage G was largely confined to site groups 1, 2a and 2b, species assemblage I was largely restricted to site group 2a and site group 6 and species assemblage L was most faithful to site group 2c and site group 5. The different subgroups within site group 2 reflected differences in Acacia species found on these sites. Acacia tetragonophylla was the most widespread Acacia occurring in all three subgroups, A. linophylla was largely restricted to subgroups 2a and 2b as was A. victoriae while A. grasbyi was restricted to site group 2b and A. coolgardiensis subsp. effusa was largely restricted to subgroup 2c and site group 5 (Table 3). Weedy annuals such as *Hypochaeris glabra, *Bromus arenarius, *Cenchrus ciliata, *Sonchus oleraceus and *Asphodelus fistulosa are more common in subgroups 2a and 2b compared to 2e. The coastal Acacia and Melaleuca shrublands of site group 3 are largely defined by species assemblages Q and R which are highly faithful to site group 3 but with generally only low to moderate consistency. Site group 3 as a whole has a coastal distribution extending from Edel Land to Cuvier; average species richness is high at 45.6 taxa Table 3 Sorted two way table showing the major site groups and species assemblages. Sites appear as columns, species as rows. 2a 1 2b 2c 3a Site Groups 3b 3c 3d 4a QC 0' 4b 5 6 8 7 9a 9b bbnbcplbbmbbmmlbbwwwlmmwwnplbclccccleelppplgkkggggmlkmmmlnnnnlkmklnlnnnlzzlzzz boabuelbbdooddlooooolddooaelbuluuuul11leeeljeejjjjrlerrrleeeelerelelaaaluuluuu 121311124345121133541451224156123451121253113423451152541134511321213451121453 1 1 1 1 1 1 1 1 1 1 1 11 1 1 Species Assemblage A Abutilon otocarpum Phyllanthus maderaspatensis Bulbostylis barbata Calandrinia sp. (GJK/NG 1495) Cassia helmsii Eremophila subfloccosa Maireana planifolia Asphodelus fistulosus Chenopodium cristatum Chthonocephalus spathulatus Rhodanthe charsleyae Stenopetalum pedicellare Acacia linophylla Spartothamnella teucriiflora Myriocephalus guerinae Acacia victoriae Tragus australianus Sclerolaena densiflora Boerhavia gardneri Hakea preissii Corchorus walcottii Eragrostis dielsii Ptilotus villosiflorus Triraphis mollis Eriachne aristidea Acacia aneura Eriachne pulchella Ptilotus helipteroides Tribulus astrocarpus Tripogon loliiformis Paspalidium basicladum Goodenia havilandii Enneapogon caerulescens Haloragis trigonocarpa ptilotus aervoides Stenopetalum sphaerocarpum Cleome viscosa Lepidium oxytrichum Eriachne helmsii ------+-------+-----+------+--+----+--+---+--------+----+----+---+-+---+--+--**** ***** *** *** ** * * * * * * * ** * * * * * ** ** * * * * ** ** *** ***** ***** * 1* * * ** * * ** * ** * ** 1 *** * *** ** * ** * * ** I ** * ** ** * * * ***** I * ** * * * * 1* ** *** * * ** * * * 1 1 * 1 1 1 1 1 1* *1 1 1 1 1 1 1 * 1 1 1 1 * 1 1* * 1 1 * 1 *1 * 1 1 1 1 1 1 1 1 1 * ***** 1 1 1 1 1 1 1 1 1 1 * * * * 1 * * * * 1 * * * 1 ** * * * 1 * * 1 ** * 1 * * 1 ** * * 1 ** ***** * 1 *** ** 1 * * ** 1 * ** **** 1 * * * *** *** ** * 1 ** 1 * **** * * * 1 1 * *** * **** ** 1 **** 1 1 *** * 1 1 * *** * * ****** * ** 1 1 * * **** ** 1 * **** *** 1 ** * *1* * * 1 * * ***** 1 1 *1* *** 1 *1* 1 ** 1 1 *1* * * * 1 1 * * * 1* 1 1 1 * *1 1 1 1 * *1 *1 1 1 * *1 * * ** * * * * *** * **** * ** *** * ****** ** * ** * *** * * * *** I 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 I 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 I 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 C'l ..;-. ~ III ~. ::r III ~ :z C'l ;: Cl> 0 .? :: Z r-' '< 0 ::l !:!' > ~ ~ ...l:0- c: (JQ III , . Indigofera monophylla Eriachne dominii Sida kingii Species Assemblage B Eriachne avenacea Waitzia acuminata Menkea villosula Thyridolepis mitchelliana Eremophila setacea ms Plectrachne rigidissima sida rohlenae Hibiscus coatesii Species Assemblage C Acacia xiphophylla ptilotus exaltatus Cassia sturtii ptilotus polakii Streptoglossa liatroides Panicum effusum Plectrachne schinzii Sauropus crassifolius Sida corrugata Atriplex lindleyi subsp. inflata Marsdenia viridiflora Sida aff. intricata (GJK/NG 1134) Yakirra australiensis 1 1 1 1 I * I ---- -+-------+--- I 1 I 1 1 1 1 I I 1 1 1 I * * I 1 * ---+--+----+--+---+------ -+-- 'TI *I **1 * * I* ,.,0l'ol l'ol -+----+----+---+-+- -+--+--- ::l Q.. 0( * ** - * * * * ** * * * ** ** * ---+-- * * * * * -+-- * * ** * * * ----+-- * -----+----+----+---+-+---+--+-- * +--+- -+-- ----+ -+- -+---+-+---+--+-- *** *** * ** * ----+- -+--+---+-- ** ** * - ** * *** ** * * * *** ** ** ----+-----+------+ * * * - ** ** ------+-------+-----+------+--+- +----+----+---+-+---+--+- * ** * ::l * * * ---- * * * ** ** * ;,o ~ ** * * -+------+--+----+--+---+-- ----+-------+-- ()q *** * * ~ *** * * Species Assemblage D Acacia grasbyi Sarcostemma viminale Cassia sp. Zygophyllum ovatum Atriplex semilunaris Sisymbrium iri Thysanotus speckii Species Assemblage E Brachyscome ciliaris Podolepis capillaris Tetragonia cristata Enteropogon acicularis Eremophila latrobei Maireana carnosa Eremophila clarkei Hibiscus burtonii Eremophila leucophylla Ptilotus grandiflorus Gyrostemon ramulosus 1 1* I* * ** * * * * ** * * * ** --+- * ---+ -+ ---+--+---+----- -+-- * -+----+---+-+---+ 00 -+--- '-J Table 3 (cont.) 1 2a 2b 2c Site Groups 3b 3c 3d 4a 3& 5 4b 6 7 8 9a 9b bbnbcplbbmbbmmlbbwwwlmmwwnplbclccccleelppplgkkggggmlkmmmlnnnnlkmkInlnnnlzzlzzz boabuelbbdooddlooooolddooaelbuluuuulllleeeljeejjjjrlerrrleeeelerelelaaaluuluuu 121311124345121133541451224156123451121253113423451152541134511321213451 12 1453 1 1 1 1 1 1 1 1 1 1 1 11 1 1 Species Assemblage F Acanthocarpus verticillatus Glycine canescens Gymnema granitica ms Angianthus tomentosus Lawrencia spicata Maireana georgei Zygophyllum kochii Chorizema racemosum Eriachne benthamii Calandrinia eremaea Calandrinia liniflora Lepidium linifolium Species Assemblage G Amaranthus mitchellii Commicarpus australis Portulaca oleracea Trianthema triquetra Maireana tomentosa Atriplex vesicaria Swainsona pterostylis Calotis multicaulis Cephalipterum drummondii Sclerolaena recurvicuspis Senecio glossanthus Maireana trichoptera Pogonolepis muelleriana Convolvulus erubescens Heliotropium undulatum Hypochaeris glabra Lotus cruentus Rhodanthe psammophila Rhodanthe chlorocephala Tribulus forrestii Dactyloctenium radulans Eragrostis basedowii Tribulus terrestris Lawrencia densiflora Wahlenbergia tumidifructa ------+-------+-----+------+--+----+--+---+--------+----+----+---+-+---+--+--- 1 1* 1** 1 * * * 1 1 ** * 1 * 1 1 1 1 1 1 1 1 1* 1 1* 1 1* 1 1 * *1 1* 1 1* 1 1* 1 1* 1 1* 1 1** 1 1* 1 1* 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1* 1 1 1 1 1 1 1 1 1 1 1 1* 1 1 1 1 1 1 1 1 1 1 1 1 1 1 ------+-------+-----+------+--+----+--+---+- 1 1 1 1*** **1* 1 1 ** **1* 1 1 * * 1* 1 1 * ** 1 1 * 1 * 1* 1 **** 1 * 1* * 1 1* * 1** 1 ** 1 ** **1** **1 *** ** 1 1***** 1 * 1 **1* ** 1 ** * 1 * * 1 * I 1 1 *1 ** * * 1 * 1 1 1 1 *** 1** 1* ** * 1** 1 1**** 1* * 1 ** 1** * 1* 1 * 1** * 1 1 1 * 1* 1 * 1 * 1** 1* 1 1* 1 1 1* 1 1 1* 1 1 1* 1 1 * 1* 1 1 1 1 1 *1 *1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1* 1 1 1 *1 1 1** 1 1 1 1 1 1 1 1 1 1 1 1* 1 1 1 1 *1 1 1 1 1 1 1 1 1 1 *1 1 1 1 1 1 1 1 1 *1 *1 1* 1 1 1 11 1 11 1 11 1 11 1 1I 1 I 1 1 11 1 11 1 *1 1 1 11 1 11 1 11 1 11 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 I 1 1 1 * 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 I 1 1 1 1 1 1 I 1 1 1 1 -----+----+----+---+-+---+--+--- 1 1 1 1 1 1 1 1* 1 1 1 1 1 1 1 * 1 1 *1 1 *1 1 1 *1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1* 1 1 1 1 * 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 * 1 1 1* 1 1 1* 1 1 1 *1 1 1 1 1 *1 *1 1 1 1 1 *1 I 1 1 1 1 1 * 1 * 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 I 1 1 1 1 1 1 1 1 1 I 1 1 * 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 I 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 I 1 1 1 1 1 ------+-------+-----+------+--+----+--+---+--------+----+----+---+-+---+--+--- Species Assemblage H Calandrinia ptychosperma Eucalyptus victrix Indigofera colutea Emex australis Sisymbrium erysimoides Lotus australis Species Assemblage I Acacia murrayana Pityrodia paniculata Adriana tomentosa Trichodesma zeylanicum Anthobolus foveolatus ms Indigofera occidentalis Acacia ramulosa Alectryon oleifolius Corynotheca pungens Psammagrostis wiseana Pityrodia loxocarpa Tephrosia gardneri (GJK/NG 1074) ms Eremophila crenulata Tribulus cistoides Rhodanthe citrina Goodenia tenuiloba Rhagodia drummondii Species Assemblage J Calotis hispidula Solanum cleistogamum Exocarpos sparteus Lobelia winfridae Leichardtia australis Rhagodia eremaea Senecio gregorii Jasminum calcarium Species Assemblage K Abutilon oxycarpum Euphorbia australis Heterodendrum oleifolium Cassia chatelainiana Lobelia heterophylla Acacia sclerosperma Cenchrus ciliaris Acacia tetragonophylla Enchylaena tomentosa Euphorbia boophthona Paractaenum novae-hollandiae Chenopodium gaudichaudianum * 1 1 1 1 1 1* 1 * * * * * 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 * 1 * 1 1 * 1 1 1 1 ** 1 * 1 * 1 * 1 **1 **1 ***1 * 1 * 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 * * 1 1 * 1 1 ** 1 ** 1 ** 1 1 * 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 *1 1 1 1 1 * 1 *1 1 1 1 * 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1* 1 * 1* 1* 1 * 1 1 1 * 1 * 1** 1 * 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1* 1 1 1 1 1 1 1 1 1 1 1 1 1 1** 1 1 1 1 1 1* 1 * 1* 1 1 1 1 1 1 1 1 1 1 1 1 1 ..., ...0' l'I l'I ::l Q. <: ~ ()Q ~ ------+-------+-----+------+--+----+--+---+--------+----+----+---+-+---+--+--- ** *** * * * * * 1 * 1 1 1 1 1 * * 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 *1 1 1 1 1 1 1 * 1 1 1 1 1 1 1 1 1 1 1 1 * * * 1 1 1 1 1 1 1 1 1 1 1 1 *1 1 1 1 1 1 1 ! £ o' :l * ------+-------+-----+------+--+----+--+---+--------+----+----+---+-+---+--+--- 1 1 1 1 1 1 1 1 1 1 1 1 1 1 * 1 1 1 1 1 1 1 1 1 *1 1 * 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 * 1 *1 *1 1 1 1 1* 1 1 1 1 1 1 1 1 1 1 1 * ------+-------+-----+------+--+----+--+---+--------+----+----+---+-+---+--+--- * * 1 1 1 1 1 1 1** 1 1 1* * *1** * 1 *1 1 * * 1 1 * ****1 *1 1 1 ** 1 1 1 1 1 *** * 1*** *1 1* 1**** * 1** * 1 1 * 1 ******1*****1 * ***1 ** 1*******1*****1 1 * 1*******1** * 1* 1* 1****** 1**** 1* ** 1** 1 * ****1*****1* * *1 * * * * ** * ** *** ** **** * * **** ** * 1 1 1 1 ** 1 1 1 1 1 1 * 1 * 1 1 *1 1 1 1 * 1 *1 ** 1 1 * 1 1 *1 1 1 1 ** **1 1** 1* * ***1 * *1 *1 1***1 ****** 1 1 1 1***1 ** **** 1* **1 *1***1 1 * * *1 *1* * 1 1 1 ******1 * 1 '"'" Tab l e3 ( con t . ) 1 2a 2b 2 c 3 a S i t eG roup s 3b 3 c 3d ~a ~ 6 7 8 9a 9b 5 b b n b c p l b bm b bmm l b bwww lmmww n p l b c l c c c c l e e l p p p l g k k g g g gm l kmmm l n n n n l km k l n l n n n l z z l z z z b o a b u e l b b d o o d d l o o o o o l d d o o a e l b u l u u u u l l l l e e e l j e e j j j j r l e r r r l e e e e l e r e l e l a a a l u u l u u u 453 121311124345121133541451224156123451121253113423451152541134511321213451121 1 1 1 11 11 1 1 1 1 11 1 1 T e t r a g o n i ad i p t e r a A r i s t i d ah o l a t h e r a P t i l o t u sp o l y s t a c h y u s E u p h o r b i ad r umm o n d i i S o l a n um l a s i o p h y l l um M u r c h i s o n i a v o l u b i l i s E r a g r o s t i sl a n i p e s B r a c h y s c om ec h e i l o c a r p a T r i g l o c h i nc a l c i t r a p um B r a c h y s c om ei b e r i d i f o l i a C a l a n d r i n i ap o l y a n d r a E r o d i um c y g n o r um P im e l e am i c r o c e p h a l a C r a s s u l ac o l o r a t a p t i l o t u sg a u d i c h a u d i i p t i l o t u so b o v a t u s S i d ac a l y x h y r n e n i a R h a g o d i al a t i f o l i a S t y l o b a s i ums p a t h u l a t um B r a c h y s c om el a t i s q u am e a S c l e r o l a e n ad i a c a n t h a B r a s s i c at o u r n e f o r t i i T h r e l k e l d i ad i f f u s a p t i l o t u sd i v a r i c a t u s R h a g o d i ap r e i s s i i N i c o t i a n ao c c i d e n t a l i s R o s t r a r i ap um i l a S o n c h u so l e r a c e u s S c a e v o l as p i n e s c e n s S c a e v o l at om e n t o s a Z y g o p h y l l um f r u t i c u l o s um C u s c u t a ep i th y rnum L e p i d i umr o t u n d um S a l s o l ak a l i D y s p h a n i ar h a d i n o s t a c h y a G o o d e n i ab e r a r d i a n a G n e p h o s i sa r a c h n o i d e a P a r i e t a r i ad e b i l i s Sp e c i e sA s s emb l a g eL A c a c i ac o o l g a r d i e n s i ss u b s p .e f f u s a + + + + + + + + + + + + + + + * * 1 * * * *1 * * * * ** * * * * I ** * I 11 * * *1 I I * * * *I ** * ** * * * * *I ** * * * * * * ** * * *I * * * *II I I I * * * * * * *I* * * * * * * * * * I * * * * ****I ** II * * *I I * I** * *I * * * * * *1 * ** * ** * ** ** * *I * * II I I 1 ** * * I* * * * ** * *1 * *** * * * * * * * ** * * *1 * * *1 1* I I * 1 * ** * *1 ** **1 * * ** 1 * *1 1 *1 * 1* * **1 * * * * * * * * * *1 * 11 1 1** * * * 1 * * 1 11 1 *** *1 * * ** 1 * ** 1 1 11 1 * * **1* * * 1 ** I * *I * * * ** * * * 1 * * ** * * * * * * * II I * * *1 * ** * * *I * * * * ** * * * * ** * * *I * ** * * * * * ** * * II * * *I * *1 ** * ** * * * * ***** * * * *I ** *** * * * * * * II I I* * * * * * ** * ** * * ** * * * * ** * II I * * ** * * * * * ** *** * * ** * * * * 11 1 * * * * ** * * * * ** * * * * * * ** * * * 11 1 * * * * * * * * * ** * * * * * ** * * * * * * * * * * ** * * * 11 1 * * ** **** * * * * * * * * ** * * * * * 11 * 1 ** * * * * * * * * * * 11* 1 * * * * * * * * * * ** 11 * * *1 * ** * * * * * * * ** * 11 1 * * ** * ** * * ** * ** 11 1 * * * ** * * * ** * 1 1* *1 * * * ** * ** * ** * * * * * * 11 *1 * * * * * * * 11 * 1 * * * * * * * **** ** ** * 11 1 * * * **** ** * * ** * * * * * * * * * 11 1 ** * * ** * * * * * * * ** * * * * * * * * 11 1 * * * * * ** * * ** 11 1 * ** * ** * * * * * * * * * 1 11 1 *** * * * * ** * * * * * * * * * 1* *11 1 ** * * * * * * 1 11 1 * * * * * * * * * * * 1 11 1 * * * * * * * * * ** 1 11 1 * * * * * * * * * * 1 11 1 * * * * * * * ** * * * * * * * 1 11 1 * * * * ** * * * * * * 1 11 1 * * * ** * * * * * ** * * * * * * * * * * * * 1 11 1 1 1 1* I I I* I* 1* 1 1* 1 1* 1* 1 1 1 1 1 1* 1 1 1 1 1 1 1 1 1 1 1 I I * 1 11 1 I + + + + + + + + + + + + + + + - 1 1 1 1 1 1 *1 * * * *1 1 1* 1 1 11 1 1 1* * *1 11 1 1 Schoenia cassiniana Lawrencella davenportii Uldinia ceratocarpa wai tzia ni tida Chthonocephalus tomentellus Eremophila maitlandii Porana sericea Rhyncharrhena linearis Podolepis canescens Actinobole uliginosum Solanum orbiculatum Chthonocephalus pseudevax Marsdenia australis Dianella revoluta Stipa elegantissima Species Assemblage M Acanthocarpus aff. robustus (Hopper 1367) Thysanotus patersonii Mirbelia ramulosa Melaleuca sp. (Beard 6768) Plectrachne bromoides Acanthocarpus robustus Loxocarya aspera ms Species Assemblage N Acacia latipes Lechenaultia linarioides Acacia subrigida Lasiopetalum oppositifolium Melaleuca aff. quadrifidus Eremophila occidens ms Wurmbea cernua Pityrodia verbascina Acacia longispinea Tricoryne aff. corynothecoides (GJK/NG 1274) Eucalyptus eudesmioides Lamarchea hakeifolia var. brevifolia Stenanthemum complicatum Baeckea sp. (A.S.George 11346) Brachychiton gregorii Grevillea stenobotrya Verticordia forrestii Allocasuarina acutivalvis Grevillea eriostachya Acacia rostellifera Acacia spathulifolia Calothamnus borealis Halgania cyanea Alyxia buxifolia 1* ** * * * * * * * * I * * ** *** I * 1 *1 **** * ** **** *** * * * * * * I I * * * I I * * * ** * * **** * ** *** *** * * * * * I I I I 1* ** I I * 1* ** I I* I I I I I 1* I I 1** I I I* I I * * * * * * ------+-------+-----+------+--+----+--+---+--------+- I I I I I I I I * * ** I I I I I * * * * * I *** *** * ** * * 1 * * * * * * ** * **** **** --+- -+-- -+--+- +-+- - *** * * ** * * ------+-------+-----+------+--+----+--+---+--------+----+-- -+---+ +---+--+--- I I I 1 I I I I I I I I I I I I I 1 I -----+-- I * 1 ------+- ** ** I I I I I I I I I I I I I I I I I * * * * I ** * * * I 1 ** * * * * * * -+--- * * --+--+--- I I I I I * I I I I * * * * * * * * * * * +--+---+--------+----+----+---+-+---+ * * -+--- Tab l e3 (can t . ) 2 a 1 2 c 2b 3a S i t eG roup s 3b 3 c 3d ~a 5 ~ 6 7 8 9a 9b bbnb cp lbbmbbmm lbbwww lmmwwnp lb c l c c c c l e e lppp lgkkggggm lk rnm rn lnnnn lkmk ln lnnn l z z l z z z b o a b u e l b b d o o d d l o o o o o l d d o o a e l b u l u u u u l l l l e e e l j e e j j j j r l e r r r l e e e e l e r e l e l a a a l u u l u u u 121453 121311124345121133541451224156123451121253113423451152541134511 321213451 1 1 1 11 11 1 1 1 1 11 1 1 Sp e c i e sA s s emb l a g e0 C a r p o b r o t u ss p . (G JK /NG1 7 0 7 ) R h o d a n t h es t r i c t a F r a n k e n i aa f f .p a u c i f l o r a T r i g l o c h i nc e n t r o c a r p um E r a g r o s t i sp e r g r a c i l i s G u n n i o p s i ss e p t i f r a g a L aw r e n c i av i r i d i g r i s e a P o d o l e p i sg a r d n e r i Z y g o p h y l l um c om p r e s s um H a l o s a r c i a i n d i c a H a l o s a r c i ah a l o c n em o i d e s Sw a i n s o n ak i n g i i S o n d o t t i ag l a b r a t a D i d ym a n t h u sr o e i S c l e r o l a e n ae u r o t i o i d e s Sp e c i e sA s s emb l a g eP A c a c i ar o y c e i E u c a l y p t u sm a n n e n s i s C a l a n d r i n i ac o r r i g i o l o i d e s B u r s a r i ao c c i d e n t a l i s T h y s a n o t u sm a n g l e s i a n u s C a l a n d r i n i al e hm a n n i i L e u c o c h r y s um f i t z g i b b o n i i Wu rmb ea d e n s i f l o r a C a l l i t r i sg l a u c o p h y l l a G i l b e r t a t e n u i f o l i a L y s i a n ac a s u a r i n a e T r a c h ym e n eo r n at a G r e v i l l e ap a r a d o x a P l a n t a g oa f f .h i s p i d u l a C e n t r o l e p i sd rummond iana Com e sp e rma i n t e g e r r im um G n e p h o s i s e r i o c e p h a l a P o r a n t h e r am i c r o p h y l l a G o o d e n i a o c c i d e n t a l i s T r a c h ym e n ec y a n o p e t a l a E r em o p h i l ao l d f i e l d i i p t i l o t u ss t i r l i n g i i + + + + + + + + + + + + + + + - 1 ** 1 ** 1 * *1 * *1 ** *1 * *1 * *1 ** *1 ** 1 ** 1 * * *1 * *1 * **1 * * * * 1 * ** 1 1 1 1 1 1 1 I 1 1 1 1 1 1 1 1 1 * 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 11 11 11 11 11 11 11 11 11 11 1 *1 11 I 1 11 11 11 11 11 11 11 11 11 11 11 11 11 11 11 11 11 11 11 1 1 1 1 1 1 I 1 1 1 1 1 1 1 1 1 1 1 1 I 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 I 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 I 1 1 1 1 1 1 1 1 1 1 1 1 11 11 11 11 I1 11 11 11 11 11 11 11 11 11 11 11 11 11 11 11 11 11 11 11 11 11 11 11 11 11 11 11 + + + + + + + + + + + + + + _ . , + - 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 * 1 1 * 1 1 ** 1 1 * 1 1 * * 1 * 1* * * 1 1 * * * * * 1 *1 * 1 1 1 1 1 1 1 1* 1 1 1 1 * 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1* 1 1 1 1 1 1 *1 1 1* 1 11 11 11 11 11 1*1 11 11 11 11 11 11 11 11 11 11 11 11 11 11 11 11 11 1 1 1 1 1 * * * *1 1 1* * *1 1 1* * *1 *1 1 1* 1 1 * * * *1 * * *1 1 1 1 1* * *1 1 1 * * * *1 **1 1 1* 1 1* * *1 1 1**1 1 1* * *1 1 1* 1 1 1* *1 1 1 * *1 1 1 * *1 1 1 * **1 1 1 * * 1 1* 1 * * * *1 1*1 ** *1 1 1 * 1 1 1 * 1 11 1 1 11 1 1 11* 1 1 11 1 1 * 11 * *1 1 11 1 1 * 1*1 1 1 11 1 1 1*1 1 1 11 1 1 11 1 1 11 1 1 11 1 1 1 *1 1 1 11 1 1 11 1 1 11 1 1 11 1 1 11 1 1 11 1 1 11 1 1 11 1 1 11 1 1 Brunonia australis Monachather paradoxus Velleia hispida ptilotus drummondii Synaptantha tillaeacea Gnephosis tenuissima Species Assemblage Q Acacia chartacea Diplopeltis intermedia Goodenia triodiophila Scaevola thesioides Acacia coriacea subsp. coriacea Haloragis gossei Triodia pungens Daviesia hakeoides Brachysema macrocarpum Scaevola sericophylla Triodia basedowii Dampiera incana Hibiscus sturtii Indigofera brevi dens Zygophyllum eremaeum Stipa ni tida Olearia dampieri subsp. dampieri ms Stackhousia muricata Species Assemblage R Acacia ligulata Trichanthodium skirrophorum Beyeria cinerea Eragrostis barrelieri Erodium cicutarium Triodia plurinervata Urospermum picroides Eremophila glabra Wurmbea inframediana Lepidium puberulum Olearia aff. axillaris (GJK/NG 1025) Poa drummondiana Triglochin trichophorum Podotheca gnaphalioides Exocarpos aphyllus Melaleuca cardiophylla Podotheca angustifolia Thryptomene baeckeacea Stipa crini ta Waitzia suaveolens Olearia axillaris Trachymene elachocarpa * 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 -+---1 1 1 * 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 * 1 1* *1 * 1 *1 1 ***1 1* *1 1* *1 ****1****1 1 * **1 **1 **1 **1 *** 1 ***1 ****1 * * 1 * **1* ***1 * * ****1 * **1 * * 1 ****1 ** * * * * *1 * 1 *1 * 1 -+--+----+--+---+- 1 ------+----+----+- -+-+---+--+- 1 * 1* *1* **1* *1 *1 1 ** * -+--+----+--+---+--------+----+----+---+-+---+--+--- -+- 1 ** *1 1 1 1 * * 1 1 1 1 1 * * 1 1 * 1 * 1 *1** * 1** 1** **1* 1 1 1 1 ** 1 1 1 1 1 ** 1 * 1 ** 1 1 * * * ** 1 ** 1 ** 1 * ** **1***1 **1* *1 *** **1***1 *** **1 * ** **1 * ** * * *1 1 1 1 * * 1 * I 1 1 * Table 3 (cant.) 2a 1 2b 2c 3a Site Groups 3b 3c 3d 4a 4b 5 6 8 7 9a 9b bbnbcplbbmbbmmlbbwwwlmmwwnplbclccccleelppplgkkggggmlkmmmlnnnnlkmklnlnnnlzzlzzz boabuelbbdooddlooooolddooaelbuluuuulllleeeljeejjjjrlerrrleeeelerelelaaaluuluuu 121311124345121133541451224156123451121253113423451152541134511321213451121453 1 1 1 1 1 1 1 1 1 1 1 11 1 1 Atriplex cinerea scaevola crassifolia Frankenia pauciflora Rhodanthe humboldtiana Zygophyllum ammophilum Atriplex paludosa subsp. moquiniana Senecio lautus Cyperus bulbosus Millotia myosotidifolia Bromus arenarius Daucus glochidiatus Cryptandra leuophracta Danthonia caespitosa Solanum oldfieldii Dysphania plantaginella Species Assemblage S Acacia blakelyi Actinostrobus arenarius Allocasuarina campestris Calytrix strigosa Leucopogon cucullatus Lyperanthus nigricans Tetraria microcarpa Thryptomene denticulata Acacia cavealis fiS Drosera stolonifera subsp. stolonifera Boronia coerulescens Brachysema aphyllum Monotaxis lurida Petrophile semifurcata Scaevola canescens Calothamnus blepharospermus Eremaea ebracteata Stylidium macrocarpum Stylidium repens Leucopogon cordifolius Lysinema ciliatum Neurachne alopecuroidea Petrophile brevi folia ------+-------+-----+------+--+----+--+---+--------+----+----+---+-+---+--+--- * * * * * * * * 1 1 1 1 1 * 1 1* 1* 1 1* * 1 * 1 1 1 1 1 1 1 1 1 1 1* 1 1 1 I 1 I 1 1 1 1 1 1 1 1 I I I 1 1 I I 1 1 1 1 1 1 1 1 1 1 1 1 I 1 I 1 I I I I 1 1 I 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 *1 1 1* 1** 1 1 1 1** 1 1** 1 1 *1 1 *1 1**1 * 1**1 1* 1* 1 1 1**1 *1 1 1 1 1 *1 1 1 1 *1 * *1* 1 I *1 * I 1 * I 1** 1 1 1 1 * 1 1 I 1* * 1 1 1 1 1* * 1 1 1 * I 1 *1 1* 1 1 1 1 1 1 1 1 1 I I 1 I 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 1 I I 1 I I 1 1 I I *1 I 1 1 1 1 1 1 1 1 1 1 I I I 1 1 I I 1 1 1 1 I I 1 I I 1 1 1 1 1 I 1 1 I 1 I 1 I I I 1 1 1 1 I I I I I 1 1 I I I 1 1 1 I 1 1 1 1 1 I I I 1 I 1 1 1 1 1 1 I 1 1 1 1 1 * ------+-------+-----+------+--+----+--+---+--------+----+----+---+-+---+--+--- 1 1 1 1 1 I I I 1 1 1 I 1 I 1 1 1 1 1 1 1 1 1 1 1 1 1 I 1 1 1 1 1 1 1 1 I 1 I I I I 1 1 1 1 1 1 1 1 I 1 1 1 1 I I 1 1 I 1 1 1 I 1 1 1 1 1 I 1 1 I I I 1** I 1** I 1** 1 1** I 1** I 1** I 1* * 1 1** 1 * 1* 1 * I* I *I * 1 *I * I *I * 1 *I * I *1 * 1** I** 1** 1** 1** 1** 1** I** 1** I* 1** I * 1** 1 * 1** 1 * S c h o e n u sc l a n d e s t i n u s M e l a l e u c a c o n o t h am n o i d e s M e l a l e u c a s c a b r a M e som e la ena p r e i s s i i O p e r c u l a r i a s p e rm a c o c e a Cono sp e rmum m i c r o f l o r um m s L e p i d o b o l u sp r e i s s i a n u s P h ym a t o c a r p u sp o r p h y r o c e p h a l u s D r o s e r a s t o l o n i f e r as u b s p .h um i l i s M a l l e o s t em o n s pC o o l o om i a (H o p p e r1353 ) M e som e la ena p s e u d o s t y g i a H i b b e r t i a r a c em o s a H i b b e r t i a s u b v a g i n a t a M e l a l e u c a a c e r o s a C o n o s t y l i ss t y l i d i o i d e s H i b b e r t i ap u n g e n s M e l a l e u c a a f f .l e i o p y x i s(G JK /NG1 7 0 8 ) E c d e i o c o l e am o n o s t a c h y a H i b b e r t i a c o n s p i c u a P e r s o o n i aa c i c u l a r i s B a n k s i aa s h b y i B a n k s i as c e p t r um G e l e z n ow i a v e r r u c o s a C a l y t r i xb r e v if o l i a G r e v i l l e a a n n u l i f e r a P l e c t r a c h n ed a n t h o n i o i d e s S c h o l t z i as p .F o l l yH i l l (ME T r u d g e n1 2 0 9 7 ) C a s s y t h aa u r e a Sp e c i e sA s s emb l a g eT Amph ipogon t u r b i n a t u s B o r o n i ap u r d i e a n a C a s s y t h ar a c em o s a C o n o s t y l i sa c u l e a t a o r e a l i s D r y a n d r ab o r e a l i ss u b s p .b Gompho lob ium t om e n t o s um G r e v i l l e ap r e i s s i im s p i c a t a H i b b e r t i a s p i c a t as u b s p .s P a t e r s o n i ao c c i d e n t a l i s P im e l e al e u c a n t h a P i t y r o d i ao l d f i e l d i i ~ e l o n g a t um * * * * * * * * * * * * * * * * * * * * * * * * * * ** * ** * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * * ** * * * * * * + + + -+ + - + + + + + + + + + + I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I I * * * * * * * * * * * * * * * * * * * * * * * * ~ 0 ~ ~ ~ ~ ~ ~ ~ ~ [ o ' ~ ~ 96 Figure 5 G.}. Keighery, N. Gibson, M.N. Lyons, A.H. Burbidge Differences in predicted climatic parameters between the nine site groups defined by the vascular flora. (L lowest value, U = highest value, I = first quartile, M = mean, D = median). Mean annual temperature (OC): (Kruskal-Wallis 21.52 20.80 49.9, df = 8, 22.25 p < 0.0001) 22.98 23.70 +-----------------+-----------------+-----------------+-----------------+------- L1=========DM==========3---U L---------1========MD=======3---U L-------1================M=D==============* L1M* 1 2 3 4 5 6 7 8 * L-1=* *=D=M===3U * 9 *M3U Diurnal temperature range (OC): (Kruskal-Wallis 12.00 12.88 37.5, df 13.75 = 8, p < 0.0001) 14.63 +-----------------+-----------------+----------------- 15.50 +-----------------+ 1 *========D====M==============3--------------U 2 L----------1===================M===============D==U 3 3 L---1=====D==M========3--U 4 L-----1=========M==D======* 5 L1*=* 6 L1===MD=* 7 * 8 L1=*3U 9 *=M3-U Annual temperature range (OC): (Kruskal-Wal1is 38.3, df = 8, p < 0.0001) 25.50 27.10 23.90 28.70 30.30 +-----------------+-----------------+-----------------+-----------------+ 1 *=========D===M=============3-------------U 2 L----------1===================M=============D===U 3 3 L----1===D====M========3--U 4 L-------1=========M=====D===* 5 L1MD* 6 L---1========M====DU 3 7 8 L1=MD* 9 L-1=MD* Precipitation wettest quarter (mm): (Kruskal-Wallis 30.2, df = 8, p 0.0002) 44.00 74.00 29.00 59.00 89.00 +-----------------+-----------------+-----------------+-----------------+ 1 L-1===============D==M==================3------U 2 L-----------1===========D=M=============3-------U 3 L------1===================D===M========================3-U 4 *==========D===M=============3----U 5 L-1=MDU3 6 1 LD====M========3--U 7 8 9 L1=MD* *DM3U * = Flora and vegetation Figure 5 97 (cont.) Precipitation coldest quarter (mm): 69.00 101. 0 +-----------------+- 1 2 3 4 L1=*=3U 5 6 51.8, df 133.0 8, p < 0.0001) 197.0 165.0 -+-----------------+-----------------+ *===D===M=======3------U *=====D==M=======3------U 1LD===========M=============3----------U *DM=3-U **3U 7 (Kruska1-Wa11is *=DM==3U 8 9 Annual precipitation (mm); (Kruskal-Wa11is 204.0 236.3 = 43.5, 268.5 L1==MD=* df 8, P < 0.0001) 300.8 333.0 +-----------------+-----------------+-----------------+-----------------+ 1 L-l=====*=====3---U 2 *=D======M=======3--------U 3 L-------1========D=M==========3 4 L-l=====M===D==* *DM=3U 5 6 Ll=MD=* 7 * 8 Ll==*==3U 9 / plot. The four subgroups show strong geographical and edaphic separation with subgroup 3a occurring on white dunes at Bush Bay and Cuvier, subgroup 3b occurring on red sands at Cuvier, subgroup 3c occurring on white sands at Edel Land and subgroup 3d occurring on red sands on Peron Peninsular. An separate analysis of sites in site group 3 confirmed highly faithful and constant species assemblages for subgroup 3b (i.e. Triodia basedowii, Triodia pungens, Indigofera brevidens, Eragrostis lanata, Acacia rostellifera); 3c (i.e. Acanthocarpus robustus, Lepidium puberulum, Olearia sp. Edel, Poa drummondiana, Podolepis gnaphalioides, Triglochin tridlOphorum); and 3d (i.e. Acanthocarpus aff. robustus, Solanum orbiculare, Eragrostis dielsii, Dianella revoluta, Austrostipa elegantissima) while subgroup 3a lacked species assemblages typical of the other subgroups. Site group 4 consists of the inland Acacia shrublands of the Mardathuna - Kennedy Range footslopes - Gascoyne Junction area (Figure 4). Most sites are dominated by A. aneura, two have a eucalypt overstorey while one site (GP) is dominated by Senna artemisioides ssp helmsii. Average species richness is still high (39.6 taxa/ plot) but lower than site groups 2 and 3. Species assemblage A is typical of this site type with a much higher constancy than site type 2 which shares this --------U Ll===DM===* assemblage. Assemblages Band C are largely faithful to site group 4 and differentiate between two subgroups, 4a which tends to have shallower stony soils with significant calcrete in the upper layer and 4b on deeper sandy soils (Wyrwoll, Stoneman, Elliott and Sandercock, 2000). Beard (1990) states that Mulga does not normally occur in the Camarvon region but this community type was widespread in the north east quadrant of the study area. Site group 5 consists of eucalypt - Callitris or eucalypt - Acacia woodlands on red sand on the Toolonga plateau and site group 6 are sites of the red dunes on the top of the Kennedy Range (KEl, KE2) and tall dunes at Mardathuna (MR3). Both site group 5 and 6 differ from the other Camarvon groups (site groups 1 to 4) in lacking most species from species assemblage K, which are generally ubiquitous (although site group 4 is under represented in some species). Species present from this assemblage differ between site groups 5 and 6 (Table 3). Species assemblage P is typical (ie showing both high degrees of constancy and fidelity) of site group 5 and species assemblage L is shared with other Camarvon groups. Site group 6 is largely defined by the lack of assemblages that delimit the other Carnarvon groups, average species richness is low for site group 6 (21.7 / plot) G.]. Keighery, N. Gibson, M.N. Lyons, A.H. Burbidge 98 while the red sands of the Toolonga plateau (site group 5) are much richer with an average species richness of 38.3 species/plot and similar to site groups 2 to 4. The affinities of site group 7, which consists of a single site NE2, are not clear. This site is a Banksia ashbyi-B. sceptrum woodland over heath on yellowish - red sands near the top of a dune. It was an unusual site in that no annuals were recorded on the plot in either year despite a significant annual flora being recorded at NEl (only some 3 km to the NW) in 1996. It is likely that this site is transitional between the limestone heaths of site group 9 and the eucalypt woodlands of site group 5. With the complete lack of annual taxa and only 12 perennial taxa recorded from this plot, its strongest relationship in the present classification is with the species poor site group 6. The remaining two site groups are typical of the vegetation types of the Irwin (Geraldton Sandplains) district. Site group 8 are eucalypt woodlands and tree heaths on red dunes immediately south of Shark Bay while site group 9 are limestone heaths or woodlands over limestone heaths between Kalbarri and Shark Bay (Figure 4). Species assemblage N is most typical of site group 8, largely comprising taxa of sandy habitats with a southwestern distribution. Average species richness for site group 8 was 30.0 taxa/plot. Species assemblages S and T are both highly faithful to site group 9. Both these assemblages are largely southwestern taxa common on calcareous sands. This site group can be divided into two subgroups - 9a occurring on massive limestones and 9b occurring on deeper sands over Table 4 limestone. Species in species assemblage Tare southwestern taxa generally typical of massive limestones. Average species richness for site group 9 was 39.4 taxa/plot. Environmental correlates There is a strong correlation between the site groups and most of the major climatic gradients of the study area. The temperature and rainfall parameters derived from ANUCLIM show a very high degree of inter-correlation and it was not possible from the available data to determine to which aspect of these parameters the vascular flora is responding. Most parameters showed significant differences between site groups (Figure 5). Patterns in mean annual temperature reflected geographical spread of site groups with the highest mean temperatures predicted for site groups 4 and 6 which are found in the north east of the study area and the lowest for site group 9 in the south west of the study area (Figures 4 and 5). The Widespread site groups (1, 2 and 3) had the widest range of predicted mean temperatures. The inland site groups had the highest diurnal temperature range and the highest annual temperature range (Figure 5). Most of the study area had highest rainfall in winter, with the exception of sites from the Kennedy Range (site groups 6 and 4 in part) which showed significant rainfall during the warmest quarter (Figure 5). Annual precipitation was considerably higher in the area between Kalbarri and Shark Bay (site group 9) than in any other part of the study area, with the Acacia shrublands (site Mean values of soil parameters of nine groups derived by classifying sites according to their vascular flora. Values were derived from a bulked soil sample taken from 0-10 cm layer. (S.D. indicates level of significance in difference between the means based on Kruskal - Wallis non parametric one way analysis of variance" = P< 0.05, .... = P<O.Ol, ......=P<O.OOl) EC (mS/m) Cl (%) pH (HP) Sand (%) Silt (%) Clay (%) C (%) Total N (%) Total P (ppm) P (HCO) (ppm) K (HC03) (ppm) Ca (HC03) (ppm) CEC (me%) ex Ca (me%) ex Na (me%) ex K (me%) ex Mg (me%) 1 2 3 4 651.0 0.84 8.68 84.8 6.3 8.9 0.44 0.04 185.0 15.0 345.0 15.8 4.3 4.08 5.46 0.59 2.49 3.7 0.0 7.48 92.7 1.5 5.8 0.23 0.02 129.6 8.3 135.8 0.6 2.7 1.36 0.06 0.28 0.44 9.9 0.0 9.15 93.7 1.5 4.7 0.45 0.04 222.5 10.5 57.5 25.3 3.2 2.32 0.09 0.11 0.32 12.7 0.01 6.88 90.2 3.0 6.8 0.20 0.02 135.4 6.3 167.3 0.2 3.0 1.31 0.16 0.34 0.82 Site Groups 5 6 7 8 9 S.D. 1.3 0.0 5.53 91.5 1.0 7.5 0.36 0.02 65.0 1.8 20.0 0.0 1.0 0.33 0.02 0.04 0.10 1.0 0.0 6.50 95.5 0.7 3.8 0.13 0.01 60.7 2.0 39.3 0.0 1.3 0.57 0.01 0.08 0.15 1.0 0.0 5.80 94.0 0.5 5.5 0.32 0.01 51.0 1.0 14.0 0.0 1.0 0.28 0.02 0.03 0.07 8.0 0.0 8.50 95.7 0.5 3.8 0.41 0.02 35.3 2.0 29.0 1.0 3.7 2.14 0.04 0.04 0.26 1.6 0.0 6.56 95.4 0.6 4.0 0.45 0.02 34.0 1.2 23.4 0.0 2.0 1.24 0.02 0.05 0.21 ...... ...... .... ...... ...... .. ...... ...... ...... ...... ...... ...... .... ...... ...... ...... ...... Flora and vegetation groups 2 and 4) having the lowest average rainfall (Figure 5). The nine site groups differed significantly in terms of soil chemistry and soil mechanical analysis. Soils from site group 1 were the most markedly different soils with very high levels of electrical conductivity, Cl, K, exchangeable Na, and exchangeable Mg (Table 4). Soils of site group 1 were generally loamy sands or gypsum (the two saltbush dominated plots that occurred on sand were the exceptions) while plots in the other site groups all occurred on sands except in site group 4 where two plots occurred on loamy sand. Most of the soils were acid except for the saline site group (group 1; mean pH 8.7), the red sands south of Shark Bay (site group 8) with average pH of 8.5 and the coastal Acacia and Melaleuca shrublands (site group 3) which had a mean pH of 9.1. The calcareous nature of these coastal sands can be seen from the high values for Ca and / or high exchangeable Ca. The remaining soils ranged from almost neutral for the Acacia shrublands (site group 2) to strongly acidic for the red sands of the eucalypt woodlands of the Toolonga plateau (site group 5 - average pH of 5.5). The sand dunes on the top of the Kennedy Range and at Mardathuna (site group 6) were the poorest in terms of organic C and N while site groups 1 to 4 were the most fertile in terms of total P, available P, available K, exchangeable K and exchangeable Mg. The soils of site groups 5 and 6 were more similar to the Irwin (Geraldton Sandplains) vegetation types (site groups 8 and 9) than other Carn:uvon vegetation types. This strong regional patteming in soil types was generally not seen within the geographically segregated subgroups of site group 3 (only exchangeable Ca showed significant differences between the subgroups). The low number of replicates within each subgroup may be influencing these results. Significant differences were found for some soil properties of subgroups of site groups 2 and 4; these subgroups show less well defined but still significant geographical separation (latitude was significantly different between subgroup means at P<0.05 for both groups) (Tables 5 and 6). Separation of subgroups in site groups 2 and 4 was also strongly correlated to depth of soil profile (described in Wyrwoll, Stoneman, Elliott and Sandercock, 2000). In site group 2, subgroups a and c occurred on deep sands or gravels while plots in subgroup b had soil profiles either truncated on limestone or calcrete or had soils with significant calcrete or shell deposits in the profiles. Similarly more than half the plots in subgroup 4a showed truncated profiles at depths of 30 - 90 cm underlain by massive calcrete or rock. All plots in subgroup 4b were on deep sands (> 100 cm) (Wyrwoll, Stoneman, Elliott and Sandercock, 2000). Site group 9 has a much smaller geographical spread than site 99 Table 5 Difference in soil chemistry and soil mechanical analysis between the subgroups in site group 2. Values were derived from a bulked soil sample taken from 0-10 cm layer. (S.D. indicates level of significance in difference between the means based on Kruskal - Wallis non parametric one way analysis of variance ns not significant, * = P< 0.05, ** P<O.Ol, ***=P<O.OOl) Subgroups 2b 2a EC (mS/m) 2.9 Cl (%) 0.0 7.49 pH (HP) Sand (%) 94.0 Silt (%) 1.0 5.0 Clay (%) C(%) 0.20 Total N (%) 0.02 145.7 Total P (ppm) 9.7 P (HCG}) (ppm) K (HCG,) (ppm) 145.1 0.3 Ca (HCG 3) (ppm) CEC (me%) 2.7 ex Ca (me%) 1.44 ex Na (me%) 0.04 ex K (me%) 0.30 ex Mg (me%) 0.61 Table 6 5.6 0.0 8.26 91.0 2.7 6.3 0.20 0.02 138.0 10.8 184.0 1.2 3.6 1.77 0.09 0.40 0.47 2c SD 3.0 0.0 6.83 92.5 ns ns ns ns ns ns ns ns 1.1 6.4 0.30 0.02 103.8 4. 7 848 0.3 1.8 0.92 0.06 0.17 0.22 ** ns ns ns * * Difference in soil chemistry and soil mechanical analysis between the subgroups in site group 4. Values were derived from a bulked soil sample taken from 0-10 cm layer. (S.D. indicates level of significance in difference between the means based on Kruskal - Wallis non parametric one way analysis of variance ns = not significant, * = P< 0.05, ** = P<O.Ol, ***=P<O.OOl) Subgroups 4a EC (mS/m) Cl(%) pH (HP) Sand (%) Silt (%) Clay (%) C ('Yo) Total N (%) Total P (ppm) P (1ICG,) (pp m) K (HCG,) (ppm) Ca (HCG,) (pp m) CEC (me')!o) ex Ca (meo.<,) ex Na (me%) ex K (me%) ex Mg (me%) 18.3 0.0 7.18 89.1 3.8 7.1 0.20 0.02 141.9 7.5 2113 0.3 39 1.61 0.24 0.41 1.09 4b 13 0.0 6.28 92.3 1.6 6.1 020 002 122.5 38 79.5 0.0 13 0.73 0.02 0.18 0.27 SD * ns ns ns ns ns ns ** ns ** G.]. Keighery, N. Gibson, M.N. Lyons, A.H. Burbidge 100 groups 2, 3 or 4 and while the subgroups showed no significance difference in soil chemistry or soil mechanical analysis, subgroup 9a occurred on shallow soil over limestone or in soils with significant limestone outcropping while subgroup 9b occurred on deeper sands (Wyrwoll, Stoneman, Elliott and Sandercock, 2000). DISCUSSION Flora Although the flora of the southern Carnarvon Basin is still imperfectly recorded, the current list of 2133 taxa is a marked improvement on the previous list for the entire Carnarvon Basin (an area of 91 046 km2) of 1348 taxa (Keighery and Gibson, 1993). This improvement occurred at all levels; for example Kalbarri National Park had 206 taxa recorded by Bums (1984), 717 taxa for the area by Bellairs and Bellairs (1992) and we recorded 1071 taxa from within the boundaries of the National Park. The study area has a predominantly arid flora of southern affinities with an overlay of temperate and tropical taxa at their range ends. The area is, however, species rich because it encompasses the change-over zone from the temperate to the arid zone. Species level conservation of the vascular flora shows that the reserve network in the area is heavily biased towards the southern and coastal margin of the zone. The wetland [40% of the 265 wetland taxa listed by Gibson, Keighery and Lyons (2000) are not known from reserves in the Basin] and red sandplain floras are particularly lacking from the reserve network. Vegetation patterning In a regional survey of this type the degree of discrimination within and between site groups is limited by the number of plots established. In our Table 7 survey, saltbush communities were only sampled at Bush Bay (BB1) and Boolathana (B01). In both cases these sites classified most similar to a vegetation unit found nearby based on shared saline species in one case (species assemblage 0 - BB1) and a suite of annuals (species assemblage G - B01) in the other. More sampling of this vegetation unit may have increased resolution of the current analysis. Due to the limited number of sites it was also difficult to separate floristic variation due to regional environmental gradients from that associated with local edaphic factors. This is complicated in the present analysis by the correlation in regional climatic variation with distribution of major soil types. Within some site groups for which it was possible to distinguish subgroups, it was found that geographical segregation (and hence correlation with regional environmental gradients) was still apparent; nonetheless correlations with edaphic factors were clearly demonstrated within at least three site groups. The climate of this region is unpredictable from year to year in terms of both precipitation and timing of rainfall events. During the two years of the study, at least one good season was experienced at all plots except NE2. For this reason the annual flora was included in the current analysis. However little is known of the patterns of accumulation of annuals at sites over longer time frames or the influence of time of year of rainfall events on the annual accumulation curves. Work of Zahran (1986) and Loneragan et al. (1991) provide some evidence of longer term climatic response by the herbaceous flora. Long term regular sampling at permanently located plots is needed quantify these processes in terms of changes in cover, changes in species occurrence and patterns of species accumulation through time. Annuals form a very significant proportion of the total flora of the region and therefore could be expected to be important in Percentage of taxa in each lifeform class by site group. Only taxa present in two or more sites are included. Percentage is based on total flora list recorded for each site group. Life form 6 7 8 9 42 8 58 1 18 8 17 0 14 0 16 0 4 4 55 4 0 8 58 17 2 5 70 9 8 12 60 3 49 3 12 1 56 3 98 5 2 3 4 Annual herbs / geophytes Annual graminoides 60 5 44 5 40 3 Perennial herbs Graminoides Shrub Trees 9 6 19 0 7 5 37 2 Total number of species Total number of quadrats 78 6 227 18 5 Site Group 5 1 42 0 35 1 4 4 27 5 158 153 12 73 4 10 11 10 10 Flora and vegetation 101 defining the major cornpositional gradients across the region. The analysis of the 63 primary sites shows an essentially similar result to the pattern discerned from our subjective appraisal of the biogeographic affiliations of the vascular flora. The vegetation patterning is closely correlated to major climatic gradients, as well as the distribution of different soil types across the study area. At a local scale edaphic factors were correlated with vegetation patterning but for the more geographically widespread groups it was not possible to separate effects of regional climatic differences and regional soil differences from local edaphic factors. The analysis showed the southern part of the Table 8 study area to be the most floristically complex with seven of the nine site groups (78'X,) being recorded from the 17 southern most sites (28%), reflecting the major change-over zone between the southwestern flora and the flora of more arid areas. This difference between the floras of the Geraldton Sandplain [Beard's (1990) Irwin district] and the Carnarvon can also be seen in an analysis of the proportional representation in life form class between the major classificatory groups (Table 7). Species rich shrub lands are typical of the southwestern flora and site groups 8 and 9 show the expected high percentage of shrubs compared to annual herbs. Most of the Carnarvon groups (site groups 1 to 5) show the opposite trend with Comparison of nine major floristic site types identified in the present study with the land system and Pasture Types defined by Payne et al. (1987). Divisions in the table reflect Payne et al. (1987) higher vegetation - soil type groupings. Pasture Type/ Land System 1 2 3 Site Groups 4 5 Acacia and eucalypt short grass forb Nerren Heath Nanga Zuytdorp Acacia sandplain Brown Lyons Peron Sandplain Yalbalgo Yaringa Coastal dune shrub Coast Edel Lyell Hard spinifex Kennedy Soft spinfex Cardabia 3 5 Not mapped 1 3 3 1 2 1 2 4 2 2 1 I 1 7 8 9 3 3 2 4 4 Acacia mixed shrub Mary Toolonga Acacia short grass forb Windalia Stony short grass forb Billy Mixed saltbush and bluebush Delta Sable Wrarroora Samphire Birrida McLeod 6 2 1 2 5 102 moderate to low percentage of shrubs and high percentage of herbs. Site group 6, the species poor dune vegetation of the top of the Kennedy Range and Mardathuna, do not follow this trend with a much higher than expected representation of shrubs. Further sampling following good rains would be needed to confirm this finding. Site group 7 consisted of only a single site (NE2) from which only 12 perennial taxa were recorded over the two major sampling periods. At site NE1 only 3 km distant large numbers of annuals were recorded in 1996 as with the other Nerren Nerren sites located some distance to the east. The soil chemistry, soil mechanical analysis and soil profile data (Wyrwoll, Stoneman, Elliott and Sandercock, 2000) were very similar between all Nerren Nerren sites. It is not clear if NE2 represents a local vegetation type that is naturally poor in annual species or had not received the same degree of rainfall as nearby sites. The floristic classification shows broad general agreement with the biogeographical boundaries of Beard (1990) with the tree heath on red sandplain south of Shark Bay and the coastal limestone woodlands and heaths (site groups 8 and 9) falling within the Irwin district and the saline group, the Acacia shrublands, the coastal Acacia and Melaleuca shrublands, the Mulga shrublands, and the red sand dune vegetation groups of the Toolonga Plateau and the Kennedy Range falling in the Camarvon district. The present classification does not support the inclusion of Edel Land as part of the Irwin district (renamed Geraldton Sandplain) as recently suggested by Thackway and Cresswell (1995). Our analysis clearly shows the relationships between Edel Land and Peron Peninsular with the rest of the coastal belt of the Camarvon district in site group 3. The Edel Land sites are separated as a subgroup, with this subgroup being most similar to the Peron subgroup. Our present data are not detailed enough to assess if Beard's phytogeographic boundaries are optimally placed (but see Gibson, Burbidge, Keighery and Lyons, 2000). We found no strong correlation between vegetation patteming and the broadly mapped surficial geology (Hocking, 1985). All sites occurred on Pliocene to Holocene aged alluvial or colluvial deposits or aeolian sands except the lower slopes of the Kennedy Range (KE3-5 which occurred on early Permian to early Cretaceous sediments) and one site on lower slopes of the Pell Range (GJ1 which occurred on Permian sediments). The only consistent land system mapping that has been undertaken across the study area is that produced by the Department of Agriculture rangelands survey group (Payne et al., 1987). This classification was based on 669 inventory sites sampled for perennial taxa and soil parameters in conjunction with aerial photograph interpretation. G.]. Keighery, N. Gibson, M.N. Lyons, A.H. Burbidge The resulting mapping recognised 89 land systems in 17 vegetation alliances termed Pasture Types. Emphasis was given to perennial taxa important to stock production in the definition of Pasture Types (Payne et al., 1987). These Pasture Types were classified into five broad groupings based on affinity of species present (Southwestern vs. Eremaean) and broad soil type. Our plots covered 23 of these land systems in 11 Pasture Types in four of the broad vegetation - soil groupings (Table 8). At the land system level there was only moderate correlation between the 23 land systems and the nine major floristic site groups. The nine site groups were represented by between one and seven land systems. Better correlation was evident between the floristic site groups and the major Pasture Types, with the nine site groups represented by between one and four Pasture Types. The correlation did not improve at the broadest vegetation - soil groupings (Table 8). This lack of correlation between the two studies relates to different aims and methodology used. The rangelands study used perennial taxa to help define mapped land units, while the present study used analysis of all taxa to define major floristic units across the study area. A reanalysis of the 63 sites using only perennial taxa could be expected to be more closely correlated with the rangeland classification. ACKNOWLEDGEMENTS Funding was provided by the Commonwealth through the National Reserves System Co-operative Program of the Australian Nature Conservation Agency (now Environment Australia), together with State funds provided by the Western Australian Department of Conservation and Land Management. Two referees provided valuable comment on an earlier draft of this paper. REFERENCES Alford J.J., Keighery, G.J. and Trudgen, M.E. (2000). Floras of the 29 islands of the Freycinet Estuary, Shark Bay World Heritage Area, Western Australia. CALMScience (in press). Anonymous (1997). Shark Bay Regional Strategy. Western Australian Planning Commission, Perth. Austin, M.P. and Belbin, L. (1982). A new approach to the species classification problems in floristic analysis. Australian Journal of Ecology 7: 75-89. Beard, J5. (1975). The Vegetation of the Pilbara Area, 1; 1,000,000 map and explanatory notes. University of Western Australia Press, Nedlands. Beard, J.5. (l976a). The Vegetation of the Murchison area, 1; 1,000,000 map and explanatory notes. University of Western Australia Press, Nedlands. Beard, J5. (1976b). The Vegetation of the Ajana Area, Western Australia. Vegmap Publications, Perth. Flora and vegetation Beard, J5. (1976c). The Vegetation of the Shark Bay and Edel Areas, Western Australia. Vegmap Publications, Perth. Beard, J5. (1990). Plant Life of Western Australia. Kangaroo Press, Kenthurst. Bellairs, D.R. and Bellairs, B. (1992). Checklist of Plants for Kalbarri and the Murclllson Sandplalll, (with supplement). Kalflora, Kalbarri. Burbidge, A.A, Fuller, P.J. and Cashin, K. (1980). The wildlife of the proposed Toolonga Nature Reserve, Shark Bay Shire, Western Australia. Dept. Fisheries and Wildlife Report 39. Burbidge, A.A and George, AS. (1978). The Flora and fauna of Dirk Hartog Island, Western Australia. Journal of the Royal Society of Western Australia 60: 7190. Burbidge, AI-I., McKenzie, N.L. and Harvey, MS. (2000). A biogeographic survey of the southern Camarvon Basin, Western Australia: background and methods. Records of the Western Australian Museum Supplement No. 61: 1-12. Bums, AC. (1984). List of Plants: Kalbarri National Park. National Parks Authority, Perth. CALM (1998). Declared Rare and Priority flora list - 21/10/ 1998. Unpublished Report, CALM, Perth. Carolin, RC., Rajput, M.J.M. and Morrison, D. (1992). Goodeniaceae. In AS. George (ed) Flora of Australia, Volume 35. Bnllloniaceae, Goodeniaceae, pp. 4-334. Bureau of Flora and Fauna and Australian Government Publishing Service, Canberra. George, AS (1999). William Dampier in New Holland: Australia's First Natural Historian. Bloomings Books, Hawthorn, Victoria. Gibson, N., Keighery, G.J. and Keighery, B.J. (1997). Contributions of N.H. Speck to the biogeography of Proteaceae in Western Australia. Journal of the Royal Society of Western Australia 80: 73-77. Gibson, N., Burbidge, AH., Keighery, G.J. and Lyons, M.N. (2000). The temperate to arid transition of the Irwin - Carnarvon phytogeographic boundary, Western Australia. Records of the Western Australian Museum Supplement No. 61: 155-173. Gibson, N. Keighery, G.J. and Lyons, M. (2000). The flora and vegetation of the seasonal and perennial wetlands of the southern Carnarvon Basin, Western Australia. Records of the Western Australian MusCllm Supplement No. 61: 175-199. Green, J.W. (1985). Census of the Vascular Plants of Western Australia. Department of Agriculture, Perth. Hocking, RM. (1985). Geology of the Carnarvon Basin. Geological Survey of Western Australia Bulletin No. 133. Hopper, S.D. and Maslin, B.R (1978). Phytogeography of Acacia in Western Australia. Australian Journal of Botany 26: 63-78. Hussey, B.M.J., Keighery, G.J, Cousens, RD, Dodd, J. and Uoyd, S.G. (1997). Western Weeds: a Guide to the Weeds of Western Australia. Plant Protection Society of Western Australia, Victoria Park, W.A Keighery, GJ. (1984). Phytogeography of the Monocotyledons of Western Australia. Kings Park Research Notes 8: 39-67. Keighery, G.J. (1990). Vegetation and flora of Shark Bay, Western Australia. In P.F. Berry, S.D. Bradshaw and 103 BR. Wilson (eds), Research in Shark Bay, pp. 61-88. Western Australian Museum, Perth. Keighery, G.J. (1995). How many weeds? An annotated list of the naturalised vascular plants of Western Australia. In G. Burke (ed), Invasive Weeds and Regenerating Ecosystems in Western Australia: Proceedings cif' the Conference held at Murdoch University, July 1994. Institute for Science and Technology Policy, Murdoch University, Perth. Keighery, G.J. (1996). Phytogeography, biology and conservation of Western Australian Epacridaceae. Annals of Botany 77: 347-355. Keighery, G.J. and Gibson, N. (1993). Biogeography and Composition of the flora of Cape Range peninsula, Western Australia. Records of the Western Australian Museum 45: 51-85. Kenneally, K.F. (1978). Notes on the vegetation and flora of Rocky Pool, Gascoyne River. Western Australian Herbarium Research Notes 1: 29-39. Loneragan, W.A., Holm, A.McR, Watson, I.W. (1991) Assessing long-term changes in native rangeland vegetation using functional species groups and multivariate analysis. Proceedings IVth International Rangeland Congress. April 22-26, 1991, Montpellier. McArthur, W.M. (1991). Reference Soils of South-western Australia. Department of Agriculture, Perth. McMahon, J.P., Hutchinson, M.F., Nix, H.A. and Ord, K.D. (1995). ANUCLIM Users Guide, Version 1. Centre for Resource and Environmental Studies, Australian National University, Canberra, Australia. Newbey, K.R. and Newbey, B.J. (1992). Notes on the Vegetation, Flora and Birds of Parts of the Kennedy Range, Gascoyne District, Western Australia. Unpublished report to CALM, Perth. Payne, A.L., Curry, P.J. and Spencer, G.F. (1987). An inventory and condition survey of rangelands in the Carnarvon Basin, Western Australia. Department of Agriculture Western Australia Technical Bulletin No. 73. Royce, RD. (1962). Botany .In A.J. Fraser (ed), The results of an expedition to Bernier and Dorre Islands, Shark Bay, Western Australia in July 1959. Fauna Bulletin of Fisheries Department, Western Australia 2: 31-53. Siege!, S. (1956). Non Parametric Statistics for Behavioural Sciences. McGraw-Hill, New York. Sneath, P.H.A. and Sokal, R.R. (1973). Numerical Taxonomy: The Principles and Practice of Numerical Classification. Freeman, San Francisco. Thackway, R. and Cresswell, I.D. (1995). (eds) An interim biogeographical regionalisation for Australia: a framework for establishing the natIOnal system of reserves, version 4.0. Australian Nature Conservation Agency, Canberra. Trudgen, M.E. and Keighery, G.J. (1995). Flora of the Shark Bay World Heritage Area and Environs. Report prepared by Western Australian Department of Conservation and Land Management for the Australian Heritage Commission. Tyler, J.P. (1988). Vegetation surveys near Lake MacLeod. KlIIgia 1: 49-74. Wilson, P.G. (1984). Chenopodiaceae. In A.5. George (ed) Flora of Australia, Volume 4. Phytoloccaceae to Chenopodlcaeae, pp. 81-317. Bureau of Flora and Fauna 104 and Australian Government Publishing Service, Canberra. Wyrwoll, K.-H., Courtney, J., and Sandercock, P. (2000) The climatic environment of the Carnarvon Basin, Western Australia. Records of the Western Australian Museum Supplement No. 61: 13-27. Wyrwoll, K.-H., Stoneman, T., Elliott, G. and Sandercock, P. (2000) The geo-ecological setting of the Carnarvon Basin, Western Australia: geology, geomorphology and soils of selected sites. Records of the Western Australian Museum Supplement No. 61: 29-75. G.}. Keighery, N. Gibson, M.N. Lyons, A.H. Burbidge Zahran, B.H.A. (1986) Forage composition and production of annual and biennial species on contrasting soils in rangeland grazed by sheep near Carnarvon, Western Australia. MSc (Agriculture) Thesis, University of Western Australia. Manuscript received 4 February 2000; accepted 14 April 2000. Flora and vegetation 105 APPENDIX 1 Vascular plants recorded for study area, their CALM conservation codes and occurrence in major conservation reserves. 3. Explanation of codes: R. 1. Declared Rare Flora - Extant taxa (= Threatened Flora Endangered + Vulnerable) Taxa which have been adequately searched for, and are deemed to be in the wild either rare, in danger of extinction, or otherwise in need of special protection, and have been gazetted as such, following approval by the Minister for the Environment, after recommendation by the State's Endangered Flora Consultative Committee. Priority 1 - Poorly Known Taxa Taxa which are known from one or a few (generally < 5) populations which are under threat, either due to small population size, or being on lands under immediate threat, eg. road verges, urban areas, farmland, active mineral leases, etc., or the plants are under threat from disease, grazing by feral animals etc. May include taxa with threatened populations on protected lands. Such taxa are under consideration for declaration as 'rare flora', but are in urgent need for further survey. 2. Priority Two - Poorly Known Taxa Taxa which are currently known from one or a few (generally < 5) populations, at least some of which are believed to be under immediate threat (ie. not currently endangered). Such taxa are under consideration for declaration as 'rare flora', but are in urgent need of further survey. Poorly Known Taxa Taxa which are known from several populations, at least some of which are not believed to be under immediate threat (ie. not currently endangered). Such taxa are under consideration for declaration as 'rare flora', but are in need of further survey. Conservation code [as defined by the Western Australian Department of Conservation and land Management (1998)]. Priority Three 4. Priority Four - Poorly Known Taxa Taxa which are considered to have been adequately surveyed and which, whilst being rare (in Australia), are not currently threatened by identifiable factors. These taxa require monitoring every 5 -10 years. Distribution Distributions refer to named places, pastoral leases, and conservation reserves (see below and Figure 1). Range limits refer to populations occurring at the limit of their known geographic range at or within the named place, pastoral lease or conservation reserve. VCl WHA Vacant Crown land Shark Bay World Heritage Area Conservation Reserve Codes BINR DINR CNR FPNP KNP KRNP TNR ZNP Bemier Island Nature Reserve Dorre Island Nature Reserve Cooloomia Nature Reserve Francois Peron National Park Kalbarri National Park Kennedy Range National Park Toolonga Nature Reserve Zuytdorp National Park Familyrraxon Conservation Code Distribution Acanthaceae Dipteracanthus australasicus subsp. australasicus Harnieria kempeana Hypoestes floribunda Adiantaceae Anogramma leptophylla Cheilanthes austrotenuijo/ia Chei/anthes lasiophylla Cheilanthes sieberi Agavaceae * Agave americana Aizoaceae * Carpobrotus aequilaterus Carpobrotus sp. (GJK & NG 1707) Carpobrotus virescens Disphyma crassifo/ium subsp. clavellarum Gunniopsis rodwayi Gunniopsis septifraga * Mesembryanthemum crystallinum Sesuvium portulacastrum Tetragonia coronaw Tetragonia cristata * Tetragonia decumbens Tetragonia diptera Tetragonia implexicoma Tetragonia tetragonoides Trialllhema oxycalyptra var. oxycalyptra Trialllhema pi/osa Trianthema triquetra Trianthema turgidifo/ia Zaleya galericulara Amaranthaceae Alternanthera nodi/lora * Alternanthera pungens Amaranthus clementii Amaranthus mitchellii Al1Ulrllllthus pallidi/lorus Gomphrena canescens Gomphrena cunninghamii Hemichroa diandra Pti/otus aervoides Pti/otus aff. chamaecladus Pti/otus aff. grandi/lorus 5 limit FPNP Endemic WHA 5 limit Nerren Nerren 5tn. N limitZNP KNP TNR Conservation Reserve ZNP CNR FPNP BINR DlNR KRNP Ptilotus Ptilotus Ptilotus Ptilotus Ptilotus Ptilotus Ptilotus Ptilotus Ptilotus Ptilotus Ptilotus Ptilotus Plilows Ptilotus Ptilotus Ptilotus Ptilotus Ptilotus Ptilotus Ptilotus Ptilotus Ptilotus Ptilotus Ptilotus Ptilotus Ptilotus Ptilotus Ptilotus Ptilotus Ptilotus Ptilotus Ptilotus Ptilotus al,'xandri arthrolasius astrolasills atnp!icifil!ius heardli culostachvus var calostuchyus caPitlllus ciwmaecladus divaricatus vaL diVilncatus divariclllus vaT. rubescens drummondli var drummondii drummondii var drummondii drurnmondii var muwr erwtrichus cxaltatus var exaltatus Rardneri RlUidiciwudli vaT. Raudichaudii Raudichaudii var. parViflorus JilWliiiflorus var RYimdiflorus Rraruliflorus vaT. lepidus helipteroi/ies humilis subsp. humilis latifiJiius macrocephalus flwnRlesii murrayi ohovlllus var. lancifiJiius obOVIlIUS VaT. obovatus petiolatus polystachyus forma polyslachyus roei rotundifi!!ius schwartm var ReorRei Ptilotus Ptilotus Ptilotus Ptilotus Ptilotus spathulatus StipilatuS stirlinRh vaT. pumilus stirlinRii var. stir!ingh villosif/orus 2 Endemic DHI Endemic WHA (Edel Land - Tamala Stn.) N limitZNP N limit FPNP N limit FPNP N limit WHA Endemic Peran Peninsular Endemic WHA Plilo[us sc!n-vartzii var sc}nvartzii Endemic WHA Amaryllidaceae Crinum flaceidum Anthericaceae Amocrinum drummondii Borya scirpoidea Borya spiwerocephala Caesia alfimlii ms Chamaeseilla ,·orymhoS([ Corynotheca mierantha vaT. acunthoclada S limit Lake MaCleod 3 N limit ZNP N limit KNP N limit KNP N limit Carrang Stn. Familyffaxon Corynotheca micrantha var. micrantha Corynotheca pungens Dichopogon preissii Dichopogon tyleri Laxmannia sessilif/ora subsp. sessi!iflora Murchisonia fragrans Murchisonia volubilis Sowerbaea laxiflora Thysanotus arenarius Thysanotus brittanii ms Thysanotus kalbarriensis ms Thysanotus manglesianus Thysanotus patersonii Thysanotus ramulosus Thysanotus sparteus Thysanotus speckii Thysanotus teretifo!ius Thysanotus thyrsoideus Tricoryne aff. corynothecoides (GJK & NG 1274) Tricoryne arenicola ms Tricoryne corynothecoides Tricoryne elatior Apiaceae Actinotus leucocephalus Apium annuum Apium prostratum * Cyclospermum leptophyllum Daucus glochidiatus Hydrocotyle pi!ifera var. glabrata Hydrocotyle rugulosa Hydrocotyle scutellifera Neosciadium glochidiatum Platysace commutata Platysace juncea Platysace sp. Kalbarri (0.& B.Bellairs 1383) Platysace sp. Kennedy (P.G.Wilson 8450) Platysace xerophila Trachymene aff. oleracea (GJK & NG 719) Trachymene coerulea var. leucopetala Trachymene cyanopetala Trachymene elachocurpa Trachymene glaucifolia Trachymene ornata Trachymene pi!osa Uldinia ceratocarpa Conservation Code 2 2 Distribution S limil Tamala SIn. N limil KNP Endemic to Sludy Area N limitCNR N range limit KNPrrNR N limit Tamala SIn. Endemic KNP N limit Nanga Stn. N limit Carrang SIn. Nlimit KNP N limit Tamala Stn. NlimitKNP N limit FPNP 2 2 N limit KNP Endemic KNP Endemic KRNP N limitKNP Confined KRNP NlimitKNP N limit Cobum Stn. N limitWHA N limit FPNP KNP TNR ZNP Conservation Reserve CNR FPNP BINR DINR KRNP ...o <» Apo cyn a c e a e Nl im i tN ang aS tn A lyxw h l ln /o lw *CU l /w ra f / thus roseus A r e c a c e a e *PhOe l l l ,1 doc ty l I /em A s c l ep i ad a c e a e C 'y t lo t lch l lm a f f/ lon lJUnd l lm(KRN1 1847 ) CV t la t lCh l l1 l lf l i l r lh l lnd l lm Gymnan fhe ra cU f ln ingham i i G .Ymn ema g r tm i r iu l r n s Sl im i tf rom(C ap eR ang e ) Nl im i tFPNP lWa r sd en iaau s t ra l i s A1a rsden ia g ran t t fco /a M l I rsde l lw v lnd t / lom Rh , " c lw r rhe "a I I l lea r ls Sa r co s t emnw v lm ina l es ub spau s t ru l e A sphod e l a c e a e *AS f ihodc l l ls ( IS l l l los l Is B l I l ln l le , 'emdJ i l rbo f ( / A sp l en i a t e a e P l eu ro so ru s ~I A s t e r a c e a e Nl im i tDHUFPNP Sl im i tH am e l in S tn , Ac t tn ( ) !J ( ) !e CO f1de rvd . l tum Ac t i f l ( ) / ) (J le d ru rnnumd iana Ac t lnobo lc I Ihg i lws l Im Sl im i tH am e l in S tn , Ang ia f l thu sa c roh va ! inu s Ang ian thus ( 'unn ing lw fn f i Ang la l l lhus m lc r I !cep fw l l ls 2 AnU iu !1 [hus m i fne i Ang ian thu sp re lSSU l f lUS Angwn lhus pusWus Angw l l lh l ls s p ,(A ,S ,G eo rg e11362 ) Angw l l lhus s p ,(A ,S ,G eo rg e11471 ) Ang ian !hu s {o rnen losus *Are lo lhcca ca lend l l lo A rC lo lheca pOp l l lU / i l Io As te rsuhu lo l l lS le rmdes AS le r ldeu as Be l l lda g ram /nco End em i cWHA End em i cWHA Nl im i tDH I Nl im i tKNP *B lde l ls h lp lmu l Io B len f l { } " ' {J0 ra d ru rnn I {Jnd i i B rachvscome che l l l ! I 'o rp " B rachys i ' ( )me ( " ; f fa r is B ra ch ) ' s ( 'o ln ec i l io co rpa B rachyscomc fw l lJph t la B ra ch v s co rn e ib e r id i (o /w Sl im i tZNP Farnilyrraxon Conservation Code Brachyscome latisquamea Brachyscome lineariloba Brachyscome perpusilla Calocephalus aervoides Calocephalus francisii Calocephalus knappii Calocephalus multiflorus Calocephalus sp. Pilbara (MET 11454) Calocephalus sp. Wiltenoom (A.S.George 1082) Caloti.1 hispidula Caloti.1 multicaulis * Centaurea melitensis Celllipeda cunninghamii Centipeda minima Centipeda thespidioides Cephalipterum drummondii Ceratogyne obionoides Chondropyxis halophila Chrysocephalum apiculatum ChrysoceplUllum semicalvum Chthonocephlllus muellerianus Chthonocephalus oldfieldianus ChtlulIlocephalus pseudevax Chthonocephalus spathulatus Chthonocepmllus tomelllellus * ColUla bipinnata ColUla coronopifolia Cotula cotuloides ColUla sp. Kalbarri (GJK & NG 2044) Cratystylis subspinescens Decazesia hecatocephnla Erymophyllum compactum Erymophyllum ramosum Erymophyllum ramosum subsp. involucratum Erymophyllum ramosum subsp. ramosum Erymophyllum tenellum Gilberta tenuif{!lia Gilruthia osbornei * Gnaphalium polycaulon Gnephosis acicularis Gnephosis angianthoides Gnephosis arachnoidea Gnephosis brevifolia Gnephosis eriocephala Gnephosis gynotricha Gnephosis macrocephala Gnephosis sp. Billabong (Nordenstam/Anderberg 203) Distribution 3 2 I Endemic Carnarvon to Shark Bay I 2 N limit KNP N limitDHI N limit Coburn Stn. S limit D1NR Endemic study area KNP TNR Conservation Reserve CNR FPNP BINR ZNP DINR KRNP ........ o Gnep/wsis lenuissima Haplolriclllon COIWJllli Haplolrichion conicum Hedypnois rha[{w.iio/ilides * Helianlhus Wlnuus Heliplerum cmspcdlOides Hyalosperma co[ula Hyalospemw demlssum Hvalosperma [{IUllflOSum subsp [{IUllllOSum Hyal()spernw glutin{}slurl subsp venus!um • H\lwclwens [{Iabm Isoelopsi.\' gmmlfll!olw * I..,,(JCfUCl1 serrio/a Lagem/era hue[{elll L(Hvrencella d(rvenportii Endemic southern Carnarvon Basin N limit Tamala Stn N limit Hamelin Stn. N limit Tamala Stn. Lawrence/la rrJsca LeuuJcht)'sum fitzgihb(mii Leucophyla broWflll Millolia depauper(J{a Mlllotia jaCkWlIll Millolia myosolidtjillw 2 Ivfinuria integerrinw * Mlfluna leplophvlla lvlyriocepfwlu.\' gascoyncnsis Ins Myrioceplwlu.\' guenllae Afynocephalus nudus Myriocephalus pyw,weus Myrioccphalus sp. (Demarz 12152) MyrlOccp/wlus sp. (Nordenstam &Anderberg 188) Olearia adenolasia Olearia affaxillaris (GJKJNG 1025) ()fearia axillaris Olearia dampien subsp. dampieri ms Olearia dampieri subsp. eremicola ms Olearia humitis Olearia imbricula Olearia mue/leri O/earia occidentissinw Olearia plme/eoides Olearia p/ucheacca Oleana revoluw Olearia rudis Olearia sWarlii Ostcospennurn co[cndu!aceum P/uchea rubelll!7ora P/uchea s'luarmsa Pluchea lelmnlhera Podolepis aunculaw Podolepis wnescells Podolepi.\' wpillans Endemic Carnarvon Basin Endemic Carnarvon Basin Endemic WHA Endemic WHA 2 Endemic WHA S limit Wooramel R. S limit Wooramel R. N limit Giralia Familyrraxon * * * * Podolepis gardneri Podolepis lessonii Podolepis microcephala Podotheca angustifolia Podotheca gnaphalioides Podotheca pollackii Podotheca pygmaea Pogonolepis muelleriana Pogonolepis stricta Pseudognaphalium luteoalbum Pterocaulon sphacelatum Pterodweta paniculata Reichardia tingitana Rhodanthe charsleyae Rhodanthe chloroceplwla subsp. splendida Rhodanthe citrina Rhodanthe condensata Rhodanthe corymbosa Rhodanthe cremea Rhodanthe humboldtiana Rhodanthe laevis Rhodanthe "umglesii Rhodanthe margarethae Rhodanthe maryonii Rhodanthe oppositifolia subsp. oppositifolia Rhodanthe oppositifolia subsp. ornata Rhodanthe polycephala Rhodanthe psammophila Rhodanthe rubella Rhodanthe sp. Overlander (P.S.Short 2096) Rhodanthe spicata Rhodanthe sterilescens Rhodanthe stricta Schoenia ayersii Sclwenia cassiniana Schoenia jilifolia subsp. arenicola Senecio glossanthus Senecio gregorii Senecio lautus subsp. dissectifolius Senecio lautus subsp. maritimus Sonchus oleraceus Sonchus tenerrimus Sondottia glabrata Streptoglossa cylindriceps Streptoglossa liatroides Streptoglossa tenuiflora Trichanthodium scarlettianum Conservation Code Distribution S limit Tamala Stn. N limit DHI N limit Woodleigh Stn. N limit KRNP Endemic WHA 2 NlimitDHI Endemic WHA Endemic WHA 2 Endemic WHA KNP TNR ZNP Conservation Reserve CNR FPNP BINR DINR KRNP .... .... N ...- - T r i e h a n t l l l } ( h l l l / ls k i r ropho rum '* Urosperrnum p i c ro id e s *U rs in ia (J f / lhemo ides *V e l ! c r c o p l l ) ' 1 o nd ca lba r l lm V i t tud in ia c e r v i cu la r i sv a r ce r v i cu la r i s V i f f l l d i l 1 1 l l CC /T l e l l i a n sv a r .o ld / i e ld i i V i t t ad in i ae r e l 1 1 a e a W a l t z i au c u r n i n a ! uv a r .u cununa la ~ a a cum ina tava ra lb i can s WW IZU l co r vmbo s l l W l lm i l l /und " Ww t zw podo l cp lS ~ a . Nl im i tDH I Nl im i tK a l b a r r ia r e a Nl im i tH am e l i nS t n . NI i l 1 1 i tZNP suav ( 'o !ens Av i c cnn i a c c a c .Av i c enn iama t lna Bo r ag in a ( ' c a c C y n o g h J S s um au s t ra l e *Ech lwn p la l l l l lg i l / cum H l l lg l l l /w a fT .I l rg y roph y l !a Ha lga l /w a fT Iwo ra l t s H l l l g a l / i aa l /aga l !o id e sv a r .wwg l l l !o id e s I11S H a { g a l / i aa l /ag l l l !o id e sv a rp r e iSS lwwm s H a { g a l / i aa rg v roph v l !a H a { g a l / l ab e ! J rww H a ! t ( u n i a C} ' l l IU ' ( J H I l { g a l /w c ya l / cav a r{ I l l i s e p a l l lm s H a { gw l i a gu s la t v e l / i tv a r .compa c tu sI11S Ha {gwua { , f fo ra l t s enu / {o ra H a { g a n l as Nl im i tKNP Nl im i tT am a l aS t n . Sl im i tFPNP Ha /gon ia V l SCO S ( i He l io [ rop iu l I l cu rassaV lCU f r I H e {w t ro l ' ium h e l e ra l / thum H e l io t rop ium pan i cu {a lUm H e l to t ro l ' ium andu {a lum Om p {w { , J { a p l ' u { ac o nC I lH l P { l lg iobo th r y sp {u r i s epa {u s T r i chod e smaz e y {w l l cumv a r .z e y {an i cum tJ a ck son D i s jun c tf r omM B r a s s i c a c c a c B ra s s i cajun c ca B ra s s i ca {ou rn e fo r t i i C a kU e ma r i t ima C " r o I /O pU Sd id ym / l s D ' l ' {o ta x i s mu ra l i s D i p { o t a x i st enu t f ld ia H ym e l / o { o ! J / l Sp ro cumb en s L e l ' ld ium! J i p l i c a t / lm L e l ' id iumj l J l io s / lm Nl im i tD INR Familyffaxon * * * * * Lepidium linitiJlium Lepidium lyratogynum Lepidium oxytrichum Lepidium phlebopetalum Lepidium platypetalum Lepidium pseudoruderale Lepidium puberulum Lepidium rotundum Lepidium scandens Menkea australis Menkea villosula Phlegmatospermum drummondii Raphanus raphanistrum Raphanus sativus Sisymbrium erysimoides Sisymbrium irio Sisymbrium orientale Stenopetalum anfractum Stenopetalum filifolium Stenopetalum linea re Stenopetalum pedicel/are Stenopetalum robustum Stenopetalum sphaerocarpum Caesalpiniaceae Labichea cassioides Labichea eremaea Labichea lanceolata subsp. lanceolata Labichea teretijiJlia subsp. teretifolia Petalostylis cassioides Petalostylis labicheoides Senna artemisioides subsp. filitiJlia Senna artemisioides subsp. helmsii Senna artemisioides subsp. x sturtii Senna glutinosa subsp. charlesiana Senna glutinosa subsp. chatelainiana Senna glutinosa subsp. x luerssenii Senna notabilis Senna pleurocarpa var. pleurocarpa Senna sp. Austin (AStrid 20210) Senna sp. BilIabong (J.D.Alonzo 721) Senna sp. Meekatharra (E.Bailey 1-26) Senna sp. Tamala (M.ETrudgen 7334A) Callitrichaceae * Callitriche stagna lis Conservation Code Distribution 4 N limit D1NR 3 Range extension from Wongan Hills N limit Cobum Stn. NlimitDHI N limit Cobum Stn. Endemic WHA KNP TNR ZNP Conservation Reserve CNR FPNP BINR DINR KRNP C amp anu l a c c a c W a h l c n b e r l i lU cap en s l s ~ co rnmun tS imn l e l l l l l W a h l e n b e r l i lU l Wah l enb e r l iw l im c i l t s Wah l cnb e r l i l l l mu l l luw !L , Nl im i tDH I \Vdh lenbe rg i ( lp re l s s l l Wah l enb e r l i /a I I I mI I J ! f r u c ia C app a r a c e a e Cappan sl a s l I l n l l lU Cappd r is Sp l t / f lS t lv a r num fnu !a r lc l C l eom e un l l l em ( ' { com ev i s co sa C a ryophy l l a c e a c Ce ras f iu rn g /o rnCU l1um C ' o r r i g i o l uI i t o r a ! J s s lm l i s G vp soph i la au P e t ro rhag iu v e lu t i lw Po l y ca rpa ea ("or.ym!}()S(J Pu l ) ' l 'a rpu l lt e tmph v l lum Sag inuop e ra /a a r RaWca S l I en el ia /h eav ..' ) ' i /enc! l { )C fu rna SpC l J<u laa r v en s i s Sp e rRu /anad iand ra Sp e r l iu /a r iarub ra S t c l / a r lUf i / i f i l rm / s S l c l l l 1 r i am ed ia C a su a r in a c e a e A l /uC ( J suamwa cu l lm / v / ssub sp a eu l l va ! \ , / s A l lo ca sua r / r l ( l camp e s tn s A I / ( !C ( J s / l I 1 r im l cam / I 'u /a la A l /oC l / sua r i l la d i e l s ia rw A l /oU l sua r ina h e lm s i / A l /oC l / sua r in l l hu e l i c / iww A l / oC l ! s u a r i rw hum ih s A l l o c ( l s u a r i f 1 l 1l ehnwnn iana im i tZNP Nl Nl im i tN e r r en N e r r e nS t n , Nl im i tDH I Nl im i tT am a l a ~ Casua r in t1 { fhesa C e l a s t r a c e a e PSa l lUn ( ) t f / ( } . va Psan lnUJ l lUJya ~ eph edn J id e s C en t ro l ep id a c c a e Aph e l ia nu luns Cen f ro lep is a r is fu I (J C en t ro ! ep i sc ep lw lo lo rm i : . " sub sp .rnu r ra y i Nl im i tKNP Nl im i tKNP Nl im i tKNP Nl im i tK a lb an iA r e a Familyffaxon Centrolepis Centrolepis Centrolepis Centrolepis Cemrolepis Conservation Code drummondiana eremica glabra humillima polygyna Chenopodiaceae Atriplex amnicola Atriplex bunburyana Atriplex cinerea Atriplex codonocarpa Atriplex IlOlocarpa Atriplex isatidea Atriplex lindleyi subsp. in/lata Atriplex nummularia subsp. spathulata Atriplex paludosa subsp. baudinii Atriplex paludosa subsp. moquiniana Atriplex semilunaris Atriplex spongiosa Atriplex vesicaria subsp. incompta Atriplex vesicaria subsp. variabilis * Chenopodium ambrosioides Chenopodium auricomum Chenopodium cristatum Chenopodium curvispicatum Chenopodium desertorum Chenopodium gaudichaudianum Chenopodium melanocarpum forma leucocarpum Chenopodium melanocarpum forma melanocarpum * Chenopodium murale Chenopodium pumilio Didymanthus roei Dissocarpus paradoxus Dysplumia glandulosa Dysplumia glomuli{era subsp. eremaea Dysplumia kalpari Dysph(mia plamaginella Dysphimia platycarpa Dysphania rhadinostachya subsp. in/lata Dysplumia rhadinostachya subsp. rhadinostachya Dysph(mia sphaerosperma Einadia nutans Enehylaena tomentosa var. tomentosa Eremophea aggregata Eriochiton sc!erolaenoides Halosarcia aurieulata Halosarcia doleijormis Distribution N limit KNP NlimitDHI N limit FPNP S limit Tamala Stn. KNP TNR Conservation Reserve ZNP CNR FPNP BINR DINR KRNP /w/ocnemoides subsp. /llI/ocnemoides ha/ocnemoides subsp. lenuls ""ilca subsp. hldens Indu(i subsp /eiostachya /eploc/ada pe/llIw per/iWmililla subsp. per/iWnu/ala prulflosa pteryg()spernw subsp den/feu/ora Halosarcia pteryguspenna suhsp pter.vgospernw lv/aireat/a (illU!(:'fW Mwreana aphylla /vlaircaf/a appressa Ha/omrcia Ha/osarcia Ha/omnia Ha/osarcia Ha/osarcw Ha/osw'C/a Ha/osarcla Ha!osarcia Ha/osarcia S limit Nanga Stn. S limit Nanga Stn N limit Nanga Stn :\4aircatla afkin.\'iarw Nlalreana A-1airca!la cunl0Sd CO!li-'exa Mairca!la gcorget Maircana lfltegra !rfaircat1a Afillreww lv!airca!1a Malreww AJuireana lanosa /o/n/low me/a!locoma OP!Wslll/olla pen[0J.,;(J!1{1 MiIlreana p/am/o/w Malreana p/alvcar!'a A1aircollo P()!)'Ptcr)\t;iil Maircana sc!eroplera Mlllreana sll/)f[ala Ataircat1l1 thesioides S limit Tamala Stn. Mairea!la tomentos'u lVfaireana tric!l(Jptera Afmreana trlpfera Alaireoflo rillosa Neohassia asfroc'arpa Ostco(arpum acropterurn var. acrn!Jterum S limit BINR Rha/iodia hdccald Rha/iodw crassl/o/w Rhag(hiia drurnm(}fuhi Rhagodia erenwea R/w/iodl([ /all/o/w subsp /all/o/I([ R/w/iodw /all/olld subsp recto R/wgodlil preissit subsp. obo\'([[(J Sa/so/a ka/I 5'arcocorniu bllu'kitlfW Sarcllcllmla qllll/que/lout Sclem/ael/a blcusl'is Sc!errJ!ocfw o}stota Sc/emlaefla cUI/ea la Sc/em/aerw derw/lout Sclemlaena desertlclI/a Disjunct from South Australia Familyrraxon Conservation Code Distribution Sclerolaena diacantha Sclerolaena eriacantha Sclerolaena eurotioides Sclerolaena j(Jrrestiana Sclerolaena limbata Sclerolaena medicaginoides Sclerolaena obliquicuspis Sclerolaena recurvicuspis Sclerolaena stylosa Sclerolaena tridens Sclerolaena uniflora Sclerostegia disarticulata Suaeda australis Tecticornia arborea Tecticornia verrucosa Threlkeldia diffusa Chloanthaceae Cyanostegia angustifolia Dicrastylis fulva Dicrastylis linearijiJlia Dicrastylis micrantha Dicrastylis sp. Peron Peninsula (M.E.Trudgen 7373) Dicrastylis sp. Shark Bay (J.J.Alford 1318) Lachnostachys eriobotrya Lachnostachys ferruginea Physopsis chrysophy/la Pityrodia atriplicina Pityrodia axillaris Pityrodia cuneata Pityrodia glabra Pityrodia glutinosa Pityrodia hemigenioides Pityrodia loxocarpa Pityrodia oldfieldii Pityrodia paniculata Pityrodia teckiana Pityrodia verbascina Spartothamne/la teucriif/ora 3 3 2 N limit Nerren Nerren Stn. N limit Meadow Stn. N limit Nanga Stn. Endemic WHA Endemic WHA & S to Murchison R. NlimitDHI N limit Peron Peninsular / DHI Endemic WHA Endemic Nerren Nerren Stn. area N limit Eurardy Stn. N limit Tamala Stn. S limit Cobum Stn. Clusiaceae Hypericum gramineum Hypericum japonicum Colchicaceae Burchardia multiflora Burchilrdia rosea Endemic Kalbarri Area KNP TNR Conservation Reserve ZNP CNR FPNP BINR DINR KRNP .... .... QC Wunnb ea c e rnua Nl im i tKNP lVu rmh ead e l lS / f lom ~ , J p " , ' r r n , l " c \Vu rmh ead i lo { ( i f (1 lVu rmh eam f ram edw l / i I lVu rmb eamm /wu l l l l H /u rmh ca mun ' h i s u n i d n a \Vu rmh cao t lo ra fa lVu rmb ea ) lVu rmb easp (KRN11582 Wu rmb ea Comm e l in a c e a e ~ ' ph i f yd ro id e s Nl im i tCNR 4 Nl im i tD INR am a l aS tn . Nl im i tT Nl im i tKNP Convo lvu l a c e a e Bonam ia C ' ( J ! 1 \ ' ( ) ! v u ! u s CO l lw lm lu s E vn [ vu lu sl l l s ino id e s Ipomo eaC(}S{( i [U Ipomo eam u e l l e n Ip (uw ;eapeS -C { lp rae s ub sp ,b ras i l iens is Po l ym e r i l ls p .(R JC r an f i e ld 2563 ) Po ranas e r i c ea \ \ / i lson ioh a c k lwL / s ' e i lV i / SO i l i l lhU f lu " s C r a s su l a e c a c C ra s su [a co lo ru [a acununu fu ( , ' ru s su laco [o ra fa v a r co {o raw ( . ' ra s su ladcc 'u rnb cn s d e c 'umb en s Cm s su l l l *Crassu la f lOWn . \ , ' C r l l s s u ! 1 Ip ed / ( e l lo s i I C r o S SU ! i Ip edu l / cu ! l In s Sl im i tFPNP Nl im i tC a r r angS tn . Nl im i tS a lu t a t ionI s i . Nl im i tFPNP Cu cu rb i t a c e a e A 1 u k i a nu . lde r l lspa fana Sl im i tWoo r am e l R Cunon i a c e a c Aphw l l lp e l l l !w l /, ! e l /w l ld eum Nl im i tDH I Cup r e s s a c c a c AC l i rw s t rohu sa re fwnu s Ca l l i t r i sg !au coph. . , . { fa Cu s cu t a c c a c Cu s cu la epuh vmum *C 'u s cuw p /an i j lu ra Cu s cu ta r i c to r iana Nl im i tCNR Familyrraxon Conservation Code Distribution Cymodoceaceae Amphibo/is un/urcticu Amphibolis griffithii Cymodoceu unguslala Cymodoceu serrulula Hulodule uninervis Syringodium isoeti/iJlium Cyperaceae Buumeu ucutu Buwneu urticulala Baumea junceu Bulbostylis burbuf(l Cuustis dioica Chrysitrix distigmu!osu Cyperus aff. cunninglwmii (GJK & NG 592) Cyperus ulterni/lorus Cyperus bifax Cyperus blakeanus Cyperus bulbosus Cyperus castaneus Cyperus difformis Cyperus gymnocuulos * Cyperus humulosus Cyperus ixiocarpus Cyperus rigidel/us Cyperus squurrosus Cyperus vuginutus Eleocharis acuta Eleocharis geniculuta Eleoc!wris pul/ens Guhniu lunigeru Gahniu trifidu Isolepis cernuu Isolepis congruu Isolepis cyperoides Isolepis murginata lsolepis nodosa Isolepis setiformis Lepidosperma angustatum Lepidosperma rupestre Lepidospermu scabrum Lepidosperma tenue Lipocarpha microcephala Mesomeluena preissii Mesomelaena pseudostygia N limit KNP N limit KNP N limit KNP N limit CNR S limit Gascoyne R. N IimitDHI N limit KNP N limit DHI N limit Carrang Stn. 4 N limit ZNP Endemic KNP N limit KNP N limit KNP N limit KNP N limit Carrang Stn. KNP TNR ZNP Conservation Reserve CNR FPNP BINR DINR KRNP .... N o ------------------------------------------------Sc!1{!efl()pleCIUs dis.wclwfll!1us S limit Coolcalalaya Stn. (from Hamersley Rg.) ScllOefl(lpleCfUS lalen/lorus Sch(Jcf1(Jplectus li{(lra/is Sc!1oenople,}us suhulafus N N N N 5'clwcnus andrewsif 5'clwcnus drmeria Sclwcnus clandeslinus Sc!1oenus curYI/oltus .Schocnus e!ega!ls KNP KNP ZNP KNP N limit KNP N limit CNR N limit Murchison House Stn. Sclwenus glolll/er Sc!1ocnus Sc/wel/us Sc/weflus Schoeflus limit limit limit limit glwu!J/lon/s gn/lil1lWluS !1cxal/drus !1unults Se/wcl/us lalirafls N limit CNR N limit KNP ,S('h{)enus plel(},\-tefn(ineUS N limit KNP N limit Tamala Stn. N limit Carrang Stn. Endemic Kalbarri area Endemic KNP to TNR N limit KNP N limit CNR Schoenus !/(lflUS Sclwenus (}dontocarpus Sc!1oel/us Sc!1oeflus Sclwel/us Sc!1oCI/US sp sp sp sp (Keighery & Alford 1966) G Broad Sheath (KLWilson 2633) Kalbarn (K.R.Newhey 9352) Murchison (KLWilson 2647) 2 ,S'chocnus VtJrii('cflae Tt/raria microcarpa Dasypogonaceae AWfllhocarpus afT. mhuslus (Hopper 1367) AWfllhocarpus !1ufluhs Acafll/wcarpus parVl/h!rus Acarlt!l()("arpus prelssll Acunlfwcarpus UJhusfuS Acafllh""alpus \'crllcillalus (,'a/n>{usia ('\,(ltlC[J Calccra,na gwndt/!ow Chl1nll1c.xeros serra Lmllllfldw /lIlslJlis IAurwndra nUlritinw Xero!irio!l di"'aricala Dilleniaceae Hlhbertia acemsa HiJlbertia uJ/upit'ua Hibbertia Hihhertia Hihherlia Hibberlw desfIlophylla exasperala glofllcm.w huegelti Hihhertia hypericoides Hlhherlla POfCflfilll/!ow Hlbberflll racefllosa HII!berfia sp. Red Bluff (JRWhecler 2376) 2 N limit FPNP S limit Quobba Stn, Endemie to KNP S limit ZNP S limit Tamala Stn. N limit KNP N limit KNP N limit KNP N limit KNP N limit Carrang Stn, N limit N hmit N limit N limit N limit N limit N limit N limit N limit N limit ZNP Nanga Stn KNP KNP KNP KNP KNP KNP Tamala Stn. CNR ..... ..... N Familyrraxon Conservation Code Hibbertia spicata subsp. spicata Hibbertia stellaris Hibbertia subvaginata N limit KNP N IimitCNR Dioscoreaceae Dioscorea hasti/i,!ia Droseraceae Drosera Drosera Drosera Drosera Drosera Drosera Drosera Drosera Drosera Drosera Drosera Drosera N limit Salutation Isl. bu/bosa coo/amon ms ericksoniae g/andu/igera indica macrantha subsp. nwcrantha menziesii subsp. thysanosepa/a neesii subsp. borea!is radicans ramellosa sw/onitera subsp. humilis st%nitera subsp. prostrata Elatinaceae Bergia auricu/ata Bergia perennis subsp. obtusijo!ia E/atine grati%ides Distribution N limit N limit N limit N limit KNP KNP KNP KNP N limit KNP N limitZNP N limitZNP N limit KNP N limit KNP N limit Nanga Stn. S limit KNP-Nanga Stn. 2 Emblingiaceae Emb!ingia ca/ceo/if1ora Epacridaceae Astr%ma g/aucescens Astr%ma microdonta Astr%ma serrati/i'/ium Astr%ma xerophyllum Brachy/oma sp. Kalbarri (A.H. Burbidge 4504) Conostephium preissii Leucopogon aff cordito!ius Leucopogon aff. strongy/ophyllus Leucopogon allitlii Leucopogon cOfwstephioides Leucopogon cordito!ius Leucopogon crassi/lorus Leucopogon hamu/osus Leucopogon hispidus Leucopogon leptanthus Leucopogon marginatus Leucopogon parvijlorus N limit N limit N limit N limit KNP KNP KNP KNP N limit CNR R N limitKNP N IimitKNP N limit Carrang Stn. N limit KNP N limit KNP N limit KNP N limit Kalbarri Area Presumed extinct Ajana area KNP TNR Conservation Reserve CNR FPNP BINR ZNP DINR KRNP .... N N N limit KNP Leucopogon plafllf(,tius Leucopogon polymorphus Leucopoglm slrlmgrlophrl!us N limit KNP N limit KNP Leu('(J[}()gOf1 !aNunlJlt>/lSlS Lysincnw citialum Euphorbiaceae Adriaf1i1 lomen[osa Beyeria Clnerea Beyeria N limit BINR (')'l1nes("cns Beyeria cygn()rUnl Beyena gardnen Beyeria lepidopelala BeVNw simitis Calycopcplus nwrginatus Eupllllrhia afT alsll1fj/ora (ASW 1(547) Eupllllrhia alsifllj/ora Euphorhia alolo Euphorhia auslralis subsp auslratis Euplwrhia hoophlhona Euphorhw coghlanil Eupllllrhia drummondii * Euphorhia 11f rta Euphorhia kimherlcyenns Euphorhia myrtlJldes EupllOrhia slwrkoensis Euphorhia tannctl.\'IS subsp eremophila Monolaxis graciliS Mono/iuis grmullj/om t'l1ono/axis lurida Phyllanthus calycinus Phyllanthus juernmhrii Phyllanthus laeunel!us Phyllanlhus maderaspatcnsis Phyllanlhus maillandianus Phyllanlhus sp, (ME Trudgen 7563) Poranlhera miemphylla Pseudl1nthus intrico{us Ins Ricinocmpos glaueus Ricinocarp()s murico(Us * Ricinus communis Sauropus emssif(JllUs Stachyslemon sp, (S, Van Leeuwen 210) Frankeniaceae Fmnkenia Frankenia Frankenia Frankenia Frankenia 4 R 3 S limit DINR S limit DHI N limit ZNP N limit Tamala Stn S limit DHI N limit Nanga Stn, Endemic WHA N limit ZNP S limit KNP amhita one rca conlusa lax/flora magflljica 2 N limit KNP I<'amilyrraxon Conservation Code Distribution Frankenia pauciflora Frankenia setosa Gentianaceae * Centaurium erythraea Centaurium spicatum * Centaurium tenuiflorum Geraniaceae Erodium angustilobum * Erodium aureum * Erodium cicutarium Erodium crinitum Erodium cygnorum subsp. cygnorum Pelargonium liUorale Goodeniaceae Brunonia australis Dampiera altissima Dampiera haematotricha subsp. dura Dampiera incana var. fuscescens Dampiera incana var. incana Dampiera krauseana Dampiera lindleyi Dampiera oligophylla Dampiera spicigera Dampiera stenostachya Dampiera wellsiana Goodenia aff. havilandii (GJK & NG 1505) Goodenia berardiana Goodenia caerulea Goodenia corynocarpa Goodenia drummondii Goodenia hassallii Goodenia havi!andii Goodenia maideniana Goodenia micrallllw Goodenia microptera Goodenia mimuloides Goodenia neogoodenia Goodenia occidentalis Goodenia ochracea Goodenia pinnatifida Goodenia pulchella Goodenia salmoniana Goodenia sericostachya Goodenia tenui!oba N limitZNP N limit Meadow Stn. 2 N limit KNP N limit Meadow Stn. N limit Murchison House Stn. N limit Tamala Stn. N limit KNP Nlimit KNP S limit Murchison R. N limit KNP 4 N IimitKRNP SlimitDHI 3 NlimitKNP Type only Gascoyne R. N limit Cobum Stn. KNP TNR ZNP Conservation Reserve CNR FPNP BINR DINR KRNP Goodenia Iriodioplllla G(}(Jdenia xunlh()spennu Lecherwullio chlomnlha Leehenouilio /lorilJlmdo Lechenoulllo I{(rsula Lechelloultw lil/oT/mdes Lechenaultia macriiflr!za Lechenoultio suhcwlwso Penla{Jri/o!l carey; Scacvu!fl R acacioides N limit KNP Scoel'olo anchusI/lllw ScaeV(J!a cancscens ScaeV(J/u ('hrys()!)(Jgon Sca('\'o[a crassif(J!iu .)'('a('vola CUtltlillf.{hamii Scaevolo SUlevola Scoel'olo Scal'volo SUlel'olu tlielsll glal/dub/I'm glohub/I'm humilloflil kal/ophyl/a Endemic Kalbarri Area N limit Eurardy Stn, N limit CNR N limit DHllFPNP N limit Meadow Stn, S limit Carrang Stn N limit Coburn Stn, :> N limit WHA N limit Coral Bay S limit Tamala Stn N limit S of WHA N limit KNP N limit KNP Endemic KNP Scaew){a !U!J('('()/o!a Scul'l'olu nilidu Scuevolu old/ieldil Type only Murchison R. ScaCl'o{a oxyc!O!la SUlevolo porViflllw subsp pilhome Scuevolu phle!Jopclolo Scaevo!tl {Jorocarya SUlevolu pulchelto N limit KNP N limit VCL S of Tamala Stn S limit near Carnarvon Scaevo[a repens var ereclll Ins Scaet'o[a resllucca SUlCl'olu sericophsl/u S'caevo/a spinesct'ns SUlevolo thesioldl's subsp, thesimdes Scacvola [omentusa Seal'l'olu Virgutu Symphyohusis muuople,'tm Vel/eia C)-L!lO[')()lamica Vel/cia discopfwm Vel/cia glabmw Velteia hispida Verreau:ria reirHt'ardtii S limit Nerren Nerren Stn N limit Wooramel R N limit KNP N limit KNP N limit KNP Gyrostcmonaccac CodmwUlrpus ,'Ofinl/'lllus Gyrostemo!l facenuger G}'rostenwn fartlu!osus Gyrostemofl subnudus Tenonia cyaffll/lon; N limit Coburn Stn N limit Murchison House Stn N limit KNP Familyrraxon Haemodoraceae Anigownthos humilis subsp. humilis Anigowntlws kalbarriensis Anigownthos manglesii subsp. quadrans Conostylis aculeata subsp. echinissima Conosty!is aculeata subsp. rhipidion Conosty!is aculeata subsp. septentrionora Conosty!is androstemma Conosty!is aurea Conosty!is candicans subsp. flavifo!ia Conosty!is prolifera Conostylis resinosa Conosty!is robusta Cmwsty!is stylidioides Haemodorum brevisepalum Haemodorum discolor Haemodorum paniculatum Haemodorum simulans Haemodorum spicatum Tribonanthes australis Tribonanthes violacea Haloragaceae Glischrocaryon aureum var. angustifolium G!ischrocaryon aureum var. aureum G!ischrocaryon flavescens Gonocarpus conferti/iJ!ius Haloragis gossei Haloragis trigonocarpa Myriophyllum decussatum Hypoxidaceae Hypoxis glabella var. leptantha Hypoxis occidenta!is var. occidenta!is Iridaceae Orthrosantlws laxus var. laxus Patersonia drummondii subsp. borea!is ms Patersonia graminea Pater.wnia occidenta!is Isoetaceae !soetes drummondii !soetes muellcri Juncaceae Juncus aridicola Conservation Code Distribution N limit KNP N limit Murchison House Stn. N limit Nanga Stn. Endemic to Nerren Nerren Area N limit KNP Endemic KNP to Tamala Stn. N limit KNP N limit Nanga Stn. N limit KNP N limit KNP N limit KNP NlimitDHl N limitKNP N limit KNP N limit KNP N limit CNR N IimitKNP N IimitKNP Nlimit KNP N limit Tamala Stn. N limit KNP N limitKNP N limit KNP N limit KNP N limit KNP KNP TNR Conservation Reserve ZNP CNR FPNP BINR DINR KRNP * Juneus bufrJllws Juneus kmuss;l Juneus p{wufu{ws Juncaginaceae Tng{uclllu TriR,I(}chin Trig!(}('hin Tng{uc!lln Trigl(jchin aff m{Cllmpa (GJK 1(430) ("a!('ilrapuflI centro('urpurn 111lnullSSlmUJIl mU(T(matum Trig{uchlll InclwphuruJll Lamiaceae Herniandra Hemialldm Hemiandm Hem;alldm Hemlgoua Hell/fReniu Hellugellw pungens sp (Beard 7(59) sp (Trudgen sn) sp Kalbarri (D Bellairs 15(5) 2 ,lip{alllhem mo('t(ult!ra {"JIle{t/u{1II N limit KNP Endemic WHA Endemic WHA Endemic KNP N limit KNP N limit Tamala Sin. 3 Hnnigenfu scohra IYlicrocorys !oganil1cca Mif'rfN'()rys !t'fluU()liu * Proslall/hem allnflum ProslOll/hem eampln'{!ll Prostantheru '.,>'ilkituna Salt'ia rerhenaca Weslrillgw aft' dampleri (GJK & NG 798) Wes/r;llgw damp/{'ri 2 N limit DlNR VVeslringia rigid(J Lauraceae Cassylha Cassylha Cassylha Cassyl!w Cassy/!w Cassyl!w Cassy/ha Cassyl!>a Cass.vtha (Jurea var (lutea <wrea var. {urla filtf(mnls flava g{ahella nudt/lum pUllllfrJrJll;s meemusa forma pilusa facernosa fonna racemusa Lentibulariaccae Ulricu!aria lellclla Lobeliaceae !so[onw hypucralerU(Jrmis Lube{wa{ala Lube/w glhhusa Lube!w helerophv!la N limit Shark Bay N limit FPNP N limit FPNP N limit KNP N limit KNP N limit KNP N limit KNP .... N "I Farnilyrraxon * Conservation Code Lobelia rhytidosperma Lobelia winfridae Monopsis debilis Loganiaceae Logania afL litoralis (AH Burbidge 4406) Logania biloba Logania flaviflora Logania litoralis Logania nuda Logania spermacocea Logania vaginalis Phyllangium paradoxum Loranthaceae Amyema benthllmii Amyema .fitzgeraldii Amyema gibberula var. gibberula Amyema hilliana Amyema linophylla Amyema maidenii subsp. maidenii Amyema melaleucae Amyema miquelii Amyema miraculosa subsp. miraculosa Amyema preissii Amyema sanguinea var. sanguinea Lysiana casuarinae Lysiana exocarpi Lysiana murrayi Lysiana subfalcata Nuytsia floribunda Lythraceae Ammannia baccifera Ammannia multi/lora Lythrum sp. Towrana (RJ Cranfield 2183) Rotala diandra Malvaceae Abutilon Aburilon Abutilon Abutilon Abutilon Abutilon Abutilon Abutilon afL oxycarpum (Morat 8145) amplum cryptopetalum exonemum geranioides leucopetalum otocarpum oxycarpum Distribution N limit KNP N limit VCL W of Nerren Nerren Stn. S limit KNP N limit NW Cape S limit Wooramel R. N limit ZNP S limit Carey Downs Stn. (from Barlee Range) S limit Carey Downs Stn. (from Rudall R.) S limit DINR S limit Tamala Stn. S limit Tamala Stn. KNP TNR ZNP Conservation Reserve CNR FPNP BINR DINR KRNP .... N 00 Abu t i !on sp .a l l .o x y ea rpum(Mo r a t8145 ) Abu l l !on sp .H am e l in (A .M .A shby2196 ) Abu t i !on sp .Quobb a(HD em a r z3858 ) A ! yoR yn e cun e l ! i l /1 l l lS A ! yoR yn e iw k e l / idw A ! yog rn ehu eR e l i lv a rR !o s su !a r ia e ! i l l tu sm s A ! yoR rn eh u eR e ! 1 Iv a r hueRe l t ii l lS A ( voK yn ehu eg e l i lv a r~ a a r n s A l yoR yn ep inonwnava rl epw ch !am r s A l yog yn cp i n r J ! 1 i a ! l a va r .p i t /on /ono Go s s yp iu rn au s t ru { e H ib i s cu sb ra ch y ch !a enu s H ib I s cu sb ra ch yS lphon lu s H ib i s cu sb u r tO f l l 1 H ib i s cu s coa t e s i i H ib i s cu sd rummond i l H ih i s cu s ga rdn e r i i I n s H ib i s cu sl ep to c ladu s H ib i s cu ss tu r t i lv a rR fW ld l ! lon l s H i b i SCU Ss tu r t i lv a ctn l f l ca tu s *Lavlltcracreriea Lavu le rap l e b e i l 1v a r . {o rnen losa LaH ' ren ( · ja d en sU l ( ) ra Law r en c iaR ! ( ) ! ' n e r a t a LLm ' f en c iasp Mu l e in S t a t Ion(S e l l e r317 ) 2 2 End em i cC a rn a rvon B a s in Nl im i tC a r r angS tn Nl im i tT am a l aS tn . l Nl im i tDH Lo \ -v renc ia sp i ( 'u [ ( / l . -<Hv renc ia\ ' i r id ig r i s ca *J f a !V i Ipa r v l / lo ra *lY la lvas f ruma rne r iuu lUm S idaa fLl f l t r i ca l i l(GJK& NG 1134 ) S idaca ! y xh ym enw S idaco r ruRaw Su laj ibu ! l ! i ' ra S idaj i l , ! i i f f I l lS S idain t r i ea ta S idak i nR I I la t y ea ! y x S ldap S ldaroh l ena e S idaspC am a rvon(PSS h o r t2492 ) S ldas p .P ind a r(AM l t eh e l l 3585 ) S ldas p .un i s exu a l(N .H .Sp e ek574 ) Sl im i tD INR M a r s i l c a c c a c l v fam l eaanRu s t l / i J !w Ma r s i l ea d rummond l i Ma lS i l ea e xa ra ta t v fa lS i / eah i r suw M eny an th a c c a c V i l /a lSw conR e s l l ! lo ra 3 Nl im i tKNP Farnilyrraxon Mirnosaceae Acacia aciphylla Acacia acuaria Acacia acuminata Acacia amblyophylla Acacia ampliceps Acacia anastema Acacia ancistrocarpa Acacia andrewsii Acacia aneara Acacia ashbyae Acacia aalacophylla Acacia bidentata Acacia bivenosa Acacia blakelyi Acacia cavealis ms Acacia chartacea Acacia citrinoviridis Acacia cochlearis Acacia colletioides Acacia comans Acacia coolgardiensis subsp. coolgardiensis Acacia coolgardiensis subsp. effusa Acacia coriacea subsp. coriacea Acacia coriacea subsp. pendens Acacia caspidifolia Acacia cathbertsonii subsp. cuthbertsonii Acacia didyma Acacia dielsii Acacia drepanophylla Acacia ericijiJlia Acacia erinacea Acacia jilrnesiana Acacia galeata Acacia gelasina Acacia grasbyi Acacia gregorii Acacia idiomorp}w Acacia inaeqailatera Acacia isoneura subsp. nimia ms Acacia kempeana Acacia lasiocarpa var. lasiocarpa Acacia latipes subsp. latipes ms Acacia leptospermoides subsp. leptospermoides Acacia leptospermoides subsp. obovala Acacia ligulata Acacia ligustrina Conservation Code Distribution N limit KNP N limitTNR Endemic WHA N limit ZNP N limit KNP N limit KNP N limit Tarnala Stll. N limit Murchison House Stn. N limit KNP S limit Wooramel R. 3 Endemic Shark Bay & DHI 3 Endemic Carbla Stn. area N limit Murchison House Stn. N limit Nerren Nerren Stn. 2 3 N limit Nerren Nerren Stn. N limitZNP 2 NlimitDHI N limitZNP KNP TNR ZNP Conservation Reserve CNR FPNP BINR DINR KRNP .... w o Acacia !ineo/ata subsp. lineo!a/([ Amcia linophylla N limit Eurardy Stn. Acacia ltmgispinea Acacia microhofrya AC(J("ia micrt}('ulyx A("(1("10 tn()rri,Y(}nii Acacia multisfJic(J[(l Acacia Amcia AmCla Acacia Awcia mUrUI)'{UUJ N limitCNR N limit KNP N limit KNP neuml'hrlla subsp erugata o/dlie!dll oxyclada l'a/ustns Acacia fJdraneura Awcia p!autellu 3 ms Acacia pruiw)('arpa Acacia puncricu!a!a Awcia Awcia Awcia A('acia N limit KNP illS l'ynl(l!w quadrimargll1ea quadnsu!cata romu/usa N limit ZNP N limit KNP Acacia resliacca Awcia rlwdoph!oUl Awcia mstellt!('ra N lirmt Hamelin Stn Acacia ro}'cei Acacia sa!Ju/osa Acacia saliRlla illS N limit KNP N limit KNP Acacia sa:xotilis AC(J{'ia s('irpU()!itl Acacia sc!erospernw subsp r.:/ouces('ens Awcw sclemspemw subsp sclemsperma Acacia sessilis 3 Acacia slRfWfu Awclil Amcia Amcw Acacia Acaclil Acaclil sp. (KRN 11816) spathulil()lia sphenaphvlla ms stereophylla var. cylindrata stereophylla var stereophvlla suIJrigida 2 N limit Yaringa Stn. N limit KNP N limit Eurardy Stn. N N N N limitCNR limit CNR limit Nerren Nerren Stn. limit Tamala Stn. Af'licia subtessan)g(JrJo Acacia synchronicia AcaCIa tetragonol'hyl!a Acacia Acacia Acacia Acacia Acacia * ulicifW N limit KNP vlctorwe Hianyu \i'ISeOI1(1 xanthina Acacia xiplwphvlla Neptunia dimorp!wntha ProSilpls palllda N lirmt Tamala Stn. Familyrraxon Molluginaceae Glinus IOfoides Macarthuria ausfralis Macarfhuria georgeana Macarfhuria infricata Conservation Code I 3 Moraceae Ficus platypoda var. lachnocaulos Ficas platypoda var. minor Myoporaceae Eremophila clarkei Eremophila compacta subsp. compacta Eremophila crenulata ms Eremophila cuneata ms Eremophila cuneifolia Eremophila decipiens subsp. decipiens ms Eremophila deserfi Eremophila exilijiJlia Eremophila flaccida ms Eremophila jorresfii Eremophilajraseri subsp fraseri ms Eremophila jraseri subsp. parva ms Eremophila jreelingii Eremophila gibsonii Eremophila glabra Eremophila glabra subsp. Zuytdorp (GJK & NG 518) Eremophila glabra subsp. albicans Eremophila glabra subsp. carnosa ms Eremophila glabra subsp. psammophora ms Eremophila glabra subsp. tomentosa ms Eremophila glufinosa Eremophila granitica Eremophila lanata ms Eremophila latrobei subsp. lafrobei ms Eremophila leucophyl/a Eremophila longijiJlia Eremophila mackinlayi subsp. mackinlayi ms Eremophila mackinlayi subsp. spathulafa ms Eremophila maculata subsp. brevifolia ms Eremophila mait/andii Eremophila mackinlayi subsp. mackinlayi ms Eremophila microfheca Eremophila occidens ms Eremophila o/dfieldii subsp. oldfieldii Eremophila oppositifo!ia Eremophila pantonii Distribution N limit TamaJa Stn. N limit Nanga Stn. S limit DINR Endemic WHA 2 R 2 S limit DINR S limit Nanga Stn. N limit Wooramel R. KNP TNR ZNP Conservation Reserve CNR FPNP BINR DINR KRNP E r emo l 'h ! lal 'h y l lo l 'oda m s E r emo l 'h i lal 'h y so ca l y xm s E r emo l ' l l I la I ' la lY ca l y x E r emo l ' l l I la I ' l e ro cu c l 'a E r emo l ' l l I la s e r ru lma E r emo l 'h t las e la c ea m s E r emo l 'h ! las l ' l end e l l s r n s E r em0 l 'h i las l 'u r !a m s E r emo l 'h i la s l r l i l lg y lo l 'h y l la E r emo l ' l l I la sub f /o c eo sa E r emo l 'h i laYOU I l ; ; ! t sub sp YOW I ; ; I Ir n s Nl im i tM e adow S tn . Nl im i tH am c l in S tn . End em i cWHA A1 } ' ( } ! l (J rUn lu ( 'um i f1a [um l l t ! } ' (J ! } (J ru rni n su la r e lV (WJ [J (J ru rNn l ( ) !11anum My r t a c c a c Ba e c k , 'a" Ia c ka l l i t m s Ba ( ' c k eu { lU l lRu re th l lc I n s Ba e c k eam e ; ;a f /o« J m s Ba e c k ea l ' en ta ; ;onan lha Ban k ea robu s la Ba e c k ea sp (M .ET rudg en7180 ) Ba e c k easp Mu r ch i son R iv e r(SD .Hopp c r1428 ) Ba e c k ea spMmg en cw (M .E .T rudg en12 (29 ) Ba e c k easp N ang a (AS .G eo rg e11346 ) Ba e c k ea su l JCun ea ta l ' ! en B eau j f l r t !a dam a ~ e l egun s ~ Nl im i tCobu rnS tn . Nl im i tT am a l aS tn . Nl im i tCNR 2 squu r ro su C ' ( l l l is tem ( ) f l p lu ;en i ( "eus Ca lo l lwmn l1 sb l e l 'ha ro s l ' e rm l1 s Ca lo lhamnu s bo r ea l t s Ca lo lham l l l1 s( 'h r y sa l l lh e ru s Ca lo l lwmn l1 s/ f l l 7 r !O S l 1 Ssub sp .fo rmo su s Ca lo lhamn l1 s; ; Iab c r m s C a ! l J l h am n l 1 s1 1 I im a l l J l ' h y l l u s Ca lo lhamnu s Iw lba r r ! en s ! s Ca lo l lwmnu so ld j / c ld i ! Ca ! l J l lwmnu s {h l ch y s la ch yu s Ca lo l lwmnu s quadn f /du s Nl im i tD INR Nl im i tKNP Nl im i tKNP Nl im i tT am a l aS tn . End em i cWHA Nl im i tN ang aS tn . 4 ( 'ab ) thamnu ssongu lneus Ca l y l r i xh r en jo l l l1 Ca l y l r i xd e l ' r e s sa e so la ta Ca l y l r ! xd Ca l y l r ! xd rummond t l Ca l y l r i xe ea l y ca la Ca l y l r i x/ f J r l lw sa Ca l y l r i xf ra s en Ca l y l r i x; ; lu l " /O sa Ca l y l r i x; ;mC l l t s Nl im i tWHA S tn . Nl im i tT am a l aS tn . Nl im i tEu r a rdyS tn , Nl im i tKNP Nl im i tN ang aS tn . Nl im i tT am a l aS tn 2 2 Nl im i tKNP Nl im i tKNP Nl im i tKNP Nl im i tKNP Familyrraxon Conservation Code Calytrix harvestiana 2 Calytrix leschenaultii Calytrix oldfieldii Calytrix paucicostata 2 Calytrix purpurea 2 Calytrix sapphirina Calytrix strigosa Calytrix truncatifo/ia Chamelaucium brevifo/ium 3 Chamelaucium conostigmum ms Chamelaucium marchantii 2 Chamelaucium micranthum Chamelaucium oenanthum ms Chamelaucium paucif!orum subsp. pauci!lorum ms Chamelaucium sp. Coolcalalaya (A.H.Burbidge 4233) Chamelaucium uncinatum Chamelaucium vinosum ms Conothamnus trinervis Darwinia capitel/ata Darwinia oldfieldii Darwinia paucif!ora Darwinia virescens Eremaea acuti!{)/ia 2 Eremaea dendroidea Eremaea ebracteata Eremaea ebracteata var. ebracteata Eucalyptus aff. prominens (MIH Brooker 4569) Eucalyptus aff. socia/is R Eucalyptus beardiana Eucalyptus camaldulensis var. obtuSll Eucalyptus decipiens Eucalyptus dichromophloia 2 Eucalyptus diminuta ms Eucalyptus do/ichocera ms Eucalyptus drummondii Eucalyptus erythrocorys Eucalyptus eudesmioides subsp. eudesmioides Eucalyptus eudesmioides subsp. pallida ms Eucalyptus eudesmioides subsp. sel/achiana ms Eucalyptus j1ocktoniae Eucalyptus foecunda subsp. Coolimba (M.I.H.Brooker 9556)3 Eucalyptus fruticosa Eucalyptus gillinsii Eucalyptus hypochlamydea subsp. hypochlamydea ms Eucalyptus jucunda Eucalyptus kochii Eucalyptus lenziana Distribution N limit CNR Endemic Kalbarri Area Endemic Kalbarri Area Nlimit KNP NlimitDHI N limit KNP Endemic Kalbarri Area N limitZNP N limit KNP N limit Murchison House Stn. N limit Tamala Stn. N limit KNP Endemic Kalbarri Area N limit KNP N limit KNP N limit Nanga Stn. S limit Tamala Stn. N limit ZNP N limit Towarana Stn. Endemic WHA N limit KNP S limit KNP N limit Nanga Stn. N limit CNRlCoburn Stn. KNP TNR ZNP Conservation Reserve CNR FPNP BINR DINR KRNP Eucalyptus Eucalyptus Eucalyptus Eucalsptus Eucalyptus Eucalyptus Eucalyptus Eucalyptus Eucalyptus Eucalyptus Eucalvptus Eucalyptus Eucalyptus Eucalsptus Eucalyptus leptopoda loxophleha subsp. supralaevis mallllensls subsp. vespertllla ohtusljlora old/ieldu orllrio pmllllllellS psn/ormis rif.(idula N Iimil Nerren Nerren Sin. N limit BINR N Iimil KNP N IimitWHA Endemic WHA mln'l sheathiww srna/lcalvx suhallf.(usto sllbsp. pusilla trallsomtilU'lltalis vicrnx Horna!oca(v:r afT (lurc(J HomaloclJ!Yx aureus Hspocalymma IOIlf.(ljiilll1m Lamarchea hakeljolw var. hreVl/olw Lamarchea IwkCl/iilia vac hakeljiilia Malleostemoll hurstluluset R N Iimil KNP N limil KNP N limit Nerren Nerren Sin. Endemic WHA N limit Coburn Sin. :\}ullcosfertlof1 minil....,aensis Malleostemoll pedlllu'ulatus Malleostemoll peltif.(er Mal!costemon rose us Malle'ostemoll sp Nerren Nerren (Payne 360) Malleostemoll sp Meadow Station (A.H.Burbidge sile 69) Malleostemoll sp Rabbit Proof Fence (A.H.Burbidge sile 59) . Malleostemoll sp Cooloomia (S.D.Hopper 1353) 2 Malleostemoll sp. Hardabull Rapids (DBellairs 1654A) 1 Malleostemoll sp. Kalbarri (LACraven 7083) 2 Malleostellwll sp Moonyoonooka (R.J.Cranfield 2947) 2 Malleostemoll tuherculatus Afelaleuca accm.w Melaleuca adnato Melaleuca afL leiopyxis (GJK & NG 17(8) ,Helaleuca afL eremaea (GJK & NG 1332) Melaleuca aff Iwlosericca Melaleuca all. nesophila (AHB 4520 ) Melaleuca afT. old/ieldii/nesophylla (Beard 6768) Melaleuca afL old/leldii (C. Gardner 13412) Melaleuca calotlullrlIwides Melaleuca cardiophylla Afelaleuca cilwsa Melaleuca COil crI' to Me/aleuca CO!lo{!wmnoides Melaleuca cordata Melaleuca depressa Melaleuca eleutemswchya Melaleuca ererrwea N limit Carrang Sin. N limit Coburn Sin. Endemic Nerren Nerren Sin. Endemic WHA N limit KNP N limit Tamala Stn. Endemic WHA Endemic WHA Endemic Kalbarri - Mllrchison House Stn. N limit KNP Endemic Kalbarri Area N limit KNP Familyffaxon Melaleuca julgens Melaleuca glomerata Melaleuca holosericea Melaleuca huegelii subsp. prislicensis Melaleuca lanceolata subsp. occidenlalis Melaleuca lasiandra Melaleuca lalerijlora subsp. aculifolia ms Melaleuca lalerilia Melaleuca laxif/ora Melaleuca leiopyxis Melaleuca leucadendra Melaleuca linophyl/a Melaleuca longislaminea subsp. longistaminea ms Melaleuca megacephala Melaleuca nematophyl/a Melaleuca oldfieldii Melaleuca psammophila Melaleuca radula Melaleuca rhaphiophyl/a Melaleuca scabra sen. lat. Melaleuca sp. (Trudgen sn) Melaleuca sublrigona Melaleuca Irichophyl/a Melaleuca uncinata Melaleuca undulala Melaleuca urceolaris Melaleuca viminea subsp. viminea Micromyrlus racemosa Micromyrtus sulphurea Phymatocarpus porphyrocephalus Pileanlhus aff. filifolius (Bellairs 1684) Pileanthus aff. peduncularis (Hopper 1385) Pileanthus auranlicus ms Pileanlhus bel/us ms Pileanlhus limacis Pileanlhus peduncularis Pileanthus seplenlrionalis (ML 2492) Pileanlhus vernicosus Schollzia cordala ms Scho/tzia leplanllw Schollzia sp. Cookalalaya (A.H.Burbidge sn) . Schollzia sp. Ajana (T.A.Halliday 137) Schollzia sp. East Yuna (A.C.Bums 6) Schollzia sp. Eurardy (J.S.Beard 6886) Schollzia sp. Folly Hill (M.E.Trudgen 12097) Schollzia sp. Galena (W.E.Blackall 4728) Schollzia sp. Kalbarri (N.Hoyle 623) Conservation Code 2 Distribution NlimitDHI Endemic WHA NlimitDHI N limit KNP N limit Nanga Stn. N limit KNP N limit KNP Endemic WHA N limit Carrang Stn. Endemic Murchison House Stn. N limit ZNP S limit Hamelin Stn. N limit Cobum Stn. 3 2 I I 2 Endemic Kalbarri Area KNP TNR ZNP Conservation Reserve CNR FPNP BINR DlNR KRNP S,!loll:w Sc!loll;w Sc!loll;w Sc!loll;w Sc!loll;w Sc!lo!r;w Sclwllz/{/ Sclwlwa sp Murchlson River (A.S.George 7098) NCH1hampton (AStrid 2(714) sp Red Bluff (A Gunness 2373) sp Ross Graham lookout (S.Maley 6) sp Shark Bay (MET 7429) Z-Bend (Belhurs-Kalnora 912a) ,pawlala "ben/lora 2 N limit Nanga Stn. 2 Sc!zo/t::)(J umhelli!£'Tu T!lrs!'loIl1Cnc aspera ssp glabra MS Thr.\'pffflUe!lC hacI'kC(J('Cll Thryptomene Thn'!,IOll1ene Thryptomenc Thrypfomene Thnploll1enc N limit Cohurn Stn dc('ussart[ denllclllala mw'ronu/a!o ninglwnensis "rolt/era Ins N limit CNR N limit Cohurn Stn. Thryp[()fuene racef1lulosa ThnpwlI1"ne sp Kalharri (A Gmness 2378) ThrsplOll1ene sp Wandana (AJI.Burhidge site I.') ThrsplOlI1ene sp Carrarang(M.ETrudgen 7420) Thnploll1"ne sp Eagle Gorge(A.G.Gunness 23(0) ThrsplOll1ene sp Eurardy(D.& BBellairs) ThrvplOlI1cnc sp Junga Dam(AGGunness 2383A) Thnploll1ene sp Steep Pomt(M.ETrudgen 7421) Thnploll1ene sp Tamala(METrudgen 7384) ThrvpwlI1ene slenophsl/a Thrsploll1ene slrongvlophslla Thryploll1ene slrongslop!lsl/a suhsp Tamala (METrudgen Verllcordia ((!flll/ans Verticordul chr.\'santha VCftl('(Jrdiu t'/,f-YS'{)S'lu('hys var. ('!zrysf)s[(J('h.vs Verticordia c()o!oomia Verlicordw dasyslylls suhsp. kallwrnensls Verlicordia dell.H/lora var roseoslel/a Verllcordia dens/flora var. slel/aligera \ferricurdio. dichroma vaT dichroma Verticordia dichro!lla vac Srtlfonw VertiuJrdia cri(}ceplwla Verticordia ethe!iatla var eth(J!iana Verticordiu cthe!iaf/u var j(JntlOSa Verlicordw /orresllI Verlicordia galeala Verticordia 1lIlertons VerI/cordia lepldophvlla var lepu}ophyl/a Verlicordia lepidophyl/a var ,!uamula Verl/cordw irlonadelpha var. call1lric!la Verlicordia nobilis Verticordia oculata Verticordia penflIRcUl Verli"ordia pluJlu}op!lyl/a I 2 2 2 2 7384) 4 Endemic WHA Endemic KNP to ZNP Endemic WHA N IimitCNR Endemic WHA 3 2 3 Endemic WHA 3 :3 N limit CohurnlMeadow Stn. N limit CNR N limit VCl S of Tamala Stn 2 N limit Kalharri Area Familyrraxon Verticordia picta Verticordia polytriclUl Verticordia spicata subsp. spicata Conservation Code 4 Distribution KNP TNR Conservation Reserve ZNP CNR FPNP BINR DINR KRNP N limitCNR Najadaceae Najas marina Nyctaginaceae Boerhavia burbid[?eana Boerhavia coccinea Boerhavia [?ardneri Boerhavia repanda Boerhnvia schombur[?kiana Commicarpus australis S limit Carnarvon Olacaceae Olax aurantia Oleaceae Jasminum calcarium Jasminum sp. (B. Turner I) S limit KNP S limit Carrang Sin. Ophioglossaceae Ophio[?lossum [?ramineum Ophio[?lossum lusitanicum Orchidaceae Caladenia bical/iata Caladenia bryceana subsp. cracens ms Caladenia discoidell Caladenia drummondii Caladenia .flava subsp. maculatll ms Caladenia hirtll subsp. rosea ms Caladenia hoffmanii ms Call1denia incensa ms Call1denill latifolia Caladenia lon[?icauda subsp. borealis ms Call1denill pachychilll ms ClIladenia reptans subsp. impensa ms Caladenia reptans subsp. reptllns ms ClIladenia roei Caladenia varians ms Call1denia vllrians subsp. nobi/is ms Caladenia vllrians subsp. varians ms Caladenia wanosa Cyanicula amplexans ms Cyaniculll dej{mnis ms e") -:~ /'l) R R N limit KNP ciQ" ::r /'l) N limit KNP ~ N limit KNP N limit KNP N limitKNP Endemic KNP N limit KNP ?- e") ;: '"0 .? ~ ~ r-' '< 0 :::l !!' > R N limit KNP ;:z: N limit Kalbarri Area :: ... <:r Q.; O'.l Oll /'l) C yun i cu /o g e rn rna ta I n s ( ) ' r toS !Y l i shu eg e l i l Dw r i sa fT I iU l f lo ra rAPB rown 32 ) Dw n s !an f lo ra D iu r i s ~ c D iun , D ra ko ca CO ! i c o ! o r m s D ra kono r ch lS ha rba r e l !a m s E i \ lh ran lh e ru bn l f10n l s E !Y lh ran lh e ru ema rg lnaw Eno c l l l !u sd l law lu ssub sp d l !a la lu sm s Eno c l l l !u sd l la la lu ssub sp undu !a lu sm s L "po r e / /af lmhna la M i c ro l l sm edw sub sp .m edw Pu raCU !C iU IU I YO l l s i i I n s Pa racu /cana t e rm ina l i s m s P /a soph y l !umco /n c% m s P ra soph y / !um c ypho ch l !um P ra soph y / !um e /o lu l I I P ra soph y l !um g lgon l eum P ra soph y l !um g ru c l l e P ra soph y l lum nng en s P ra soph y l lum sa rg en t l l P l e ro s t v l i sa l l/wno(G JK& NG 1867 ) sp e ra P l e ro s t v l i sa P l e ro s l r l i s /m /bow P t e ro s t y l i s nano P l e ro s l y l i sp l c /a P t e ro s t v l t s vangU lUC (J . P t e ro s t y ! i ss cuh ra P y ro r ch i sn ig r i can s Sp l cu /a ea n /ww T h e /Y I I I I / r a an l enm f e ra Th e / ym l l ra campanu !a la Th e / ym i l raj l e xuo sa Th e / ym l l ra sa rg en l l l Th e ! ym l l ra sp i ra l i s 4 R R Nl im i tKNP Nl im i tK a lb am A r e a Nl im i tKNP Nl im i tZNP Nl im i tKNP Nl im i tKNP Nl im i tKNP Nl im i tT am a l aS t n Nl im i tKNP End em i cTNR N e r r en N e r r en S t n . Nl im i tKNP Nl im i tKNP Nl im i tKNP Nl im i tKNP Nl im i tKNP Nl im i tT am a l aS tn . Nl im i tKNP Nl im i tKNP Nl im i tKNP Nl im i tKNP Nl im i tKNP O rob an ch a c c a c O robon ch e {n ino rv a r .o u s t ra l iano Ox a l id a c c a c O xa l i sp e r ennan s * Oxa l i sp e scap ra e P ap av c r a c c a c *,Aruenwneochro!euca P ap i l ion a e e a c Ao lu sph y l i c ( } {d e s Nl im i tZNP Familyrraxon Bossiaea eriocarpa Bossiaea spinescens Bossiaea walkeri Brachysema aphyllum Brachysema macrocarpum Chorizema ericiji/!ium Chorizema racemosum Clianthus formosus Crotalaria cunninghamii Cullen cinereum Cullen leucanthum ms Cullen patens Daviesia divaricata Daviesia hakeoides subsp. hakeoides Daviesia in/ricata Daviesia nudiflora subsp. nudi/lora Daviesia peduncula/a Daviesia podophylla Daviesia ramosissima Gastrolobium bennellsianum Gastrolobium oxylobioides Glycine canescens Glycine c!andestina Glycine tabacina Glycyrrhiza acanthocarpa Gompholobium sp. Kalbarri (M.D. Crisp 6300) Gompholobium Iomenlosum Indigofera australis Indigofera boviperda Indiw!fera brevidens Indigofera colutea Indigofera georgei Indigofera !inifolia Indig(ifera linnaei Indigofera monophylla Indigo/era occidenta!is ms Indigo/era sp. (KRN 11615) Isotropis cuneifolia Jacksonia aff. spinoS(l (RJ. Cranfield 2576) Jacksonia angulata Jacksonia arenicola ms Jacksonia arida ms Jacksonia calcicola ms Jacksonia cupulifera Jacksonia folio.1'il Jacksonia h<lkeoides Jacksonia rigida ms Conservation Code Distribution N IimitZNP N limit KNP N limit ZNP N limit Murchison House Sw. N limit KNP N limit Murchison House Stn. N limit KNP N limit KNP Endemic Kalbarri Area N limit Tamala Stn. N limit Nanga Stn. Endemic Carnarvon Area N IimitTNR N limit Nerren Nerren Stn. N limit KNP N limit KNP NlimitKNP KNP TNR Conservation Reserve ZNP CNR FPNP BINR DlNR KRNP ]ackson;u sternberf.:iana 4 ]acksonill l'elutinu Kennedi{[ pros/ra/a Le{'rosclllo Il{)h\'llwn ms Lep{()senw dOr'ics/{)[des Lep/osenw nwcrocarpum InS Lep/oxema /omenloSfl ms {J)IUS Iluslraln * N limit KNP N limit Hamelin Stn N limit WHA N limit Eurardy Stn. N limit Kalbani Area Endemic NW Cape to Shark Bay N limit CNR Lotus crUCfl[US Lupinus cos'oHinil Mediellgo {'oIVlllor{'l", Afedicago rrullcalulll Me!ilolus illdieus Mirbe!ill aff ralllLllosll (GJK & NG Mlrbel", aff \'flllilllllls (AHB sn) 2(59) Mlrbelw depressll N limit TNR Mlrbelw ralllLllosll Mlrbe!ill sp (Blackall 556) N limitWHA Mlrbe!ill Mirbelw Mirbe!ill Mlrbe!ill sp Zuytdorp (GJK & NG 1688) sp Denham (WEBlackall 556) sp Kalbarn (M.D.Crisp 6261) sp/fillsa AfJrbe!i1l /nelwcalvx MLlelleranthus IntollOlalus Nellleia relleulilla Ox"lolnLlIll !inellre 1'L1lrawea dllsvphvlla Rhynehoslll !lUnilllll ,)'esbania cannahina SplweroloblLllII graCile SplwerololJiulIl maeralllhLlIII Slvloslllllhes hLl!lubs S"-i'ai!ls()fUl heas/evalla Sii'uinso!w cu/cico/a 5\.vainso!lo ('Oflescens 5'wains(}na ('()mp/una/a 5\vains(ma {'(1rnuta Sit'ainsot/a {'callosa S\.,'ainsofw eleg(lf1[()ides S\vainsot/a !lal'icariuata SH'ainso!la t::racilis S\vait/so!w kingii S\vainsona lecaf/a S'wain,"lrJ!la !()fl;.;icarinato .S\vainsofla pedunculata .S\vainS0l111 [JtenJs(vlis 5\vainso!l(J rostella/a Tem{'lelof/11l bilobll TelllplelOf/11l relLlsa Te{'hroslll bldwlllll 3 Endemic Kalbani Area N limit Murchison House Stn. N limit Murchison House Stn N limit ZNP N limit KNP N limit KNP N limit KNP N limit KNP S limit KNP Endemic WHA N limit KNP N limit FPNP Conservation Code Familyffaxon * Tephrosia flammea Tephrosia gardneri Tephrosia rosea Tephrosia sp. (K.R. Newbey 11636) Trifolium campestre Trigonella suavissima Viminaria juncea Phormiaceae Dianella revoluta Stypandra glauca Pittosporaceae Billardiera bicolor var. lineata Billardiera erubescefls Billardiera ringens Bursaria occidefltalis Cheiranthera filif{}lia Cheirallthera preissiana var. preissiana Piuosporum phylliraeoides var. phylliraeoides Plantaginaceae Plalltago * Plalltago Plantago Plantago Plantago Distribution N limit KNP N limit KNP N limit Tamala Stn. N limit Nanga Stn. aff. hispidula (A.H.Burbidge 4517) coronopus debilis drummondii sp. Mt. Magnet (A.S. George 6793) Plumbaginaceae Muellerolinum salicomiaceum Poaceae * Aira caryophyllea Amphipogofl caricinus Amphipogon strictu.I· Amphipogon turbinatus Aristida aflthoxallthoides Aristida cOfltorta Aristida holathera var. holathem Astrebla elymoides Astrebla pectinata Austrostipa compressa Austrostipa crinita Austrostipa elegantissima Austrostipa macalpinei Austrostipa nitida Austrostipa tenaifolia N limit TNR N limitZNP KNP TNR Conservation Reserve ZNP CNR FPNP BINR D1NR KRNP Auslroslq){l Inchophrllll * * * * AWNrosripa fLIcker! Avellmill nllche!" A vena barba!a A t'cna Iowa A vena stcrili,\ Bothri(fChloa eH'artuuw Brachyo(>hne * Bri::a ma.yima pnJsfra{(1 '" Bri;a minor * *' * Bro!llus oreflarlUS Bromus diandrus BrofHus hordeaceus Bro!nIL) japonicus Bromus madritensis Bromus riRidus * Cenchrus ciftans * ('enchrlls echinarus '" C'enchrus setigerus Ch!oris !obala Ch/oris peUi,tala Ch!oris {'umifw Ch !on\' virgll!a Chr\'Sopogon Jallox Chrssopogon {'aU/(/us Chryso{'ogon sp, Zuytc!orp (GlK & NG 9(6) Csmbopogon wn!Jlgllus Endemic to ZNP C'ynlbopogr)JJ !NJlnbyr'inus C'ymbo{Jogon ohtecfuS ('ynodofl dacty!o!l '" [)act.v!ocleniurn australe Daus!ouenium radu!ans f)anfh()nia caf!spi!osa [)anlhonia rucemosa Dichanlhium senCtum suhsp, hum/tius Dig/tar/a brown/i Digi[orla ctefwnlha * * * * * Dip!achne /US((I Dq,!achne mueUen Dlp!achne parn/lora Ehr!wrta bren/il!/(! Ehrhllrlll ca!YC/tllI Ehrhllrtu !ongl/lora Eleus/ne ind/co Enneap()g()!l cacrules{'ens Enneapogon po!vphsllus Enteropogofl acicularis Eragrostis austra!asica Eragrostis barrelieri Eragrostis hasedrHvii N limit FPNP Farnilyrraxon • • • • • • • Eragrostis brown;; Eragrostis cumingii Eragrostis curvula Eragrostis dielsii Eragrostis elongata Eragrostis eriopoda Eragrostis falcuta Eragrostis lanipes Eragrostis leptocurpa Eragrostis pergracilis Eragrostis setifolia Eragrostis tenellula Eragrostis xerophila Eriachne aristidea Eriachne avenacea Eriachne beflthllmii Eriachne flaccida Eriachne gardneri Eriachne helmsii Eriachne lanata Eriachne mucronata Eriachne obtusa Eriachne ovata Eriachne pulchella subsp. dominii Eriachne pulchella subsp. pulchella Eriochloa australiensis Eriochloa procera Eriochloa pseudoacrotricha Eulalia aurea Eulalia fulvu Hordeum geniculatum Hordeum leporinum Iseilema eremaeum Iseilema membranaceum Iseilema vaginiflorum Lamarckia aurea Leptochloa digitata Lolium perenne Lolium rigidum Lophoch/oa pumila Monachather paradoxus Neurachne alopecuroidea Panicum australiense Panicum decompositum Panicum eUusum Panicum laevinode Paractaenum novae-hollandiae Conservation Code Distribution KNP TNR ZNP Conservation Reserve CNR FPNP BINR .... DlNR KRNP t Pa ran eu ra chn e mu e / / e r i * Pa rapho ! i s in cu r vu Pospo l id l l lm IWS lc lod l lm Pu s !w / id ium ( , I ( 'm en t i i Pu spa / id ium (D f l s tn ( ' lWn POSPO l ld l l l1 l l gmC l le POSpo l ld l l lm f l lh t f lo rum Pospo l id l l l l l l re / lex l Im *Pen ta s ch i s t i sa i ro id e s P e ro l i sran i *Phu la r i st l z ino r P log inse l l lm re fmc l l I l l l P l en ra c !m e hnu l { ( J id e s P l e c l rm 1 1 1 1 c da l l t lum i ( ) id e s P lec l rach l le d le ls l l P l e ( ' t ru ( 'hn ed rU t l ln l ( }nd i i P leu rae lme me i l ' l l le l P l eu ra c /m er ig id i s s i !1 la P / ( ' ( ' f ru ( 'hn e P (N l annua Poa d ran zmond iuna *Po{ypogO t l mO f l sp eh en s i s P sammag ro s t i sH ' i s ea fw *Ros l ra r ia c r i s la la :;: Ro s l ra r ia pum i lu *S ' ch i smu sa ra }n cu s Seh is l l l l ls bo r lm l l ls Se lO r t i ld le ls l l S e ta r iasu rg en s *S ' e ta r ia v e r t i c i / l t J lo ~ ' p {U t l z ( J s um Sp i l l l f l 'x IO l lg l to l l l lS S ! } ( J r ( J h ( J / u s( l ( , t in ( } ( ' /adu s Spombo l l ls b loke l Spombo l l ls rom l l Spombo l l ls I l l l l rh l ' l I l l S 'po roho /u sl ' t r ) { l f l lC l /S Th em eda [ r ia t /d ra Thy r tdo lep is l l l11ehe l l l i l f / { / T ragu sau s l ra l ionu s T r iod ia a f f! J a s e d oH , ' i i T r iod ia a ~ ' T r iod ia e p aC [U l rn fan s T r iod ia t T r tod io Im l lgem T r iod ia p lu r in e r va lu T r iod ia pung en s T r iod ia s ca r io su T r iod ia l r i t i co id e s Tnpogo l l lo l i l t /mms 4 Nl im i tT am a l aS t n Nl im i tFPNP Nl im i tN ang aS t n Nl im i tN ang aS t n , Familyffaxon Conservation Code Distribution 3 N limit KNP N limit KNP N limit ZNP Triraphis mollis * Trisetaria cristata * Urochloa gilesii Urochloa piligera Vulpia myuros Yakirra australiensis Polygalaceae Comesperma acerosum Comesperma calymega Comesperma integerrimum Comesperma polygaloides Comesperma scoparium Polygala isingii Polygonaceae * Emex australis Muehlenbeckia adpressa Muehlenbeckia cunninghamii Rumex crystallinus * Rumex vesicarius Portulacaceae Calandrinia aff. pumila (Andeberg 206) Calandrinia calyptrata Calandrinia corrigioloides Calandrinia disperma Calandrinia eremaea Calandrinia granulifera Calandrinia lehmannii Calandrinia linif10ra Calandrinia papil/ata Calandrinia polyandra Calandrinia polypetala Calandrinia primulijlora Calandrinia ptychosperma Calandrinia pumila Calandrinia remota Calandrinia sp. Coolcalalaya (GJK & NG 698) Calandrinia sp. Shark Bay (J.J. Alford 1376) Calandrinia sp. (GJK & NG 1495) Calandrinia stagnensis Portulaca oleracea Posidoniaceae Posidonia angustijiJIia Posidonia australis N limit Tamala Stn. N limitZNP 2 Endemic Carnarvon Basin N limit FPNP Endemic WHA KNP TNR Conservation Reserve ZNP CNR FPNP BINR D1NR KRNP ,..------------------------------------------------------ - - - - - Posidonia ('onacea Posidonia SffllUJsa Potamogetonaceae P()!alll(}gelr)!) !J('("!lfl(l!US Po{unu}gewn tnCartfla!US Ruppia mcg(u"orpa RUflflw po!rcw/,a Ruppia ruhcrosa Primulaccae * Arwgullis {lfTCnsis var an'CllSlS * Anagollls (UTCflSis va[ ('ocrulca Sa!1/()!us JUN('CUS Samolus rcpcns var. jlorihundus Samo!us repens var. paun'j"o!ius Samolus sp Shark Bay (METrudgen 7410) N limit FPNP S limit Hamelin Stn Proteaceae Adcnwlf!los a(Wlf!l{)p!lrllus Endemic to WHA N limit KNP Adc!lanth(}s ('ygl/()rum Banksw BUflksia Banksia Banksia Banksia Banksia ashh\'l oftenualu lepfoflhrlla var. melllfiCil lindlewlf/a menZlesii N limit ZNP N limit KNP N limit Tamala Stn, N limit KNP N limit Tamala Stn, N limit Nanga Stn, N limit ZNP N limit KNP Endemic Kalbarri Area N limit Nanga Stn N limit Tamala Stn, Eodemic Murchison House Stn, to Zuytdorp pri(JfUJ{CS Banksia sccpfrum Banksia vicforia(' ('oflospermum aceroswn subsp hir.\'ulUm ('ufjospernuun horca{e subsp. !Jurea!e ('{)f1(JSpennufll micnJjl(Jrurn CO!lospennum stocclwdis subsp_ sloechadis Dryandm afl. ashh)'1 (GlK & NG 1571) Dryandm horealis subsp, borealls Dryandm jmsen Dryandra s('ssllis Grevillea afl. sfenohofrya (MET 7554) Grevillea wlfluli{era Grevillea argrl'Op!lylla Grevillea hltimms subsp, bJ/lmnis Grevillea bmchrsfac!lra Grevillca Cilndelabl'Oujes Grevillea calldlcwlS Grevillca ('(Jrnmutoto Grevillea coSlofo Grel'illell dU!VlIlo/wfrYIl subsp, dU!VlIlo!Jofrya Gret'illca die/siand Grerillca erios[(1chya Grevi/lc(l gtJrd(}nitJtltJ 2 2 S limit Tamala Stn. N limit WHA N limit Tamala Stn N limit KNP N limit ZNP N limit FPNP N limit KNP N limit Nerren Nerren Stn Endemic lower Murchison R, N limit Nerren NelTcn Stn, N limit CNR ----- Familyffaxon Grevillea hakeoides subsp. stellophylla Grevillea illtricalll Grn'illea leptopoda Grevillea leucoclada Grevillea leucopteris Grevillea levis Grevillea obliquistigma subsp. obliquistigma Grevillea paradoxa Grevillea petrophiloides Grevillea pillaster Grevillea preissii Grevillea pterosperma Grevillea rogersonialla Grevillea sp. Shark Bay (MET 7553) Grevillea stellobotrya Grevillea stellomera Grevillea stellostachya Grevillea trachytheca Grevillea tri/oba Grevillea variiji,lia subsp. bUllderra ms Grevillea vestita subsp. isopogoides Hakea arida Hakea bucculelllll Hakea cundollealla Hakea circumalalll Hakea coriacea Hakea coSlll1ll Hakea illcrassata Hakea kippistialla Hakea lissocurpha Hakea orthorrhyllclw Hakea preissii Hakea pycllolleura Hakea recurva Hakea stenocarpa Hakea stellophylla Hakea suberea Hakea trij'urcalll !sopogon divergells Persoonia acicularis Persoollia angustif/ora Persoollia biglalldulosa Persoonia bowgada Persoollia brachystylis Persoollia hexagolla Persoonia papillosa Persoollia rufij10ra Conservation Code 3 Distribution NlimitDHI N limit KNP N limit KNP N IimitTNR N IimitZNP N limit KNP N limit KNP N limit KNP N limit ZNP N limitZNP 2 Endemic Shark Bay - Tamala Stn. Endemic to WHA 2 3 Endemic KNP to Tamala N limit Cobum Stn. Endemic Kalbarri Area N limit KNP S limit Lake McLeod N limit KNP 2 N limit CNRffNR N limit KNP N limit Tamala Stn. N limit KNP N limit Murchison House Stn. N limit KNP N limitZNP N limit Nanga Stn. N limit CNR N limit Nanga Stn. N limit CNR N limit KNP N limit KNP KNP TNR ZNP Conservation Reserve CNR FPNP BINR DINR KRNP N limit Nanga Stn N limit Nanga Stn N limit KNP N limit KNP N limit Tamala Stn. N limit KNP N limit Murchison House Stn N limit KNP N limit ZNP N limit KNP N limit KNP N limit KNP N limit KNP N limit Eurardy StnlCNR N IimitCNR PerSlJOf1la saundersuuw Persoonw sp. (Craven 112) Persoonia \'trtc!a PelrophIie Inlenwlll Pelropll/!e hrerifo!1I1 Pelro!,hIic (o/!lfera PetrtJ[Jhile nUUI(istm)lYo PelropllI!e S((lhnUsiu!a Petr(Jphile \'cI/litur("(it(l Perrophile \"cminwfa Pelrophl!e shull!eH(Jrlh,wlil SlIrlingw !allfolia Svnaphea rnlllTa 5'yl/aphell spinulosa subsp !Jor('u!is Xv!ome!um IIngllsllfo!wm Psilotaceae S limit KNP PSI!olum nlldum Ranuncu!aceae C!emalls !mcllnfo!w Ranuni'ulus pumlllo Runul/('ulus H'sslhilorus N limIt DHI Restionaceae N limit N limit N limit N limit N limit N limit Desnloc{adus US/h'r Ins Desmocludus hi/flrmis Ins l::cdeim'o!eu nUJ!l(JSluchru Lepidob(J!us c!wcl(Jicplw!lIs Lepidoho!lIs densus ms Lepidohollls prelsslIlnlls Lept()curpus lirisltllus Lo),."OCorYli [lorthenicu IllS LYf.{inia hur!Jata DHI KNP Tumala Stn CNR DHI KNP N limit KNP N limit KNP Rharnnaceae Cnpwndra arbullllora var horeahs Cryptandru cOIU/ala Cnplandra gllllmflora Cryptandru IcwJphruc!u Cry/Jlandru mUfi!u (',)ptandra mvriantha Cnpwndra nl/dlf!ora ('I)ptandra nulans Cryplandra pllngens Cr)ptandra scopariu var. ,nicrocephu!£l ('ryptundra s/Jyridi(Jides Srenunrhenlurn ("(Jfnplicotum Stef/anthernum dll'uricUlum Stenanthemum tnfncatum 2 N lunit DINR N limit DHI 2 3 N limit DHI S limit DHI N limit KNP . .... 'J:) --------------------------------------------------------------------------- Familyrraxon Conservation Code Stenanthemum notiale Stenanthemum pomaderroides N IimitZNP N limit KNP Rubiaceae Canthium attenuatum Canthium latifo/ium Canthium /ineare Oldenlandia galioides Opercularia aff. spermacocea (A.H.Burbidge 4502) Opercularia spermacocea Opercularia vaginata Synaptantha tillaeacea Vat. tillaeacea Rutaceae Boronia coerulescens subsp. spinescens Boronia crenulata vat. gracilis Boronia cymosa Boronia purdieana Boronia ramosa subsp. anethifolia Diplolaena geraldtonensis ms Diplolaena grandiflora Diplolaena mollis ms Eriostemon brucei Eriostemon deserti Eriostemon sericeu.l' Eriostemon sp. Kalbarri (GJK & NG 2034) Eriostemon tomentellus GeleZflowia verrucosa Phebalium tuberculosum Distribution N limit CNR N N N N N IimitZNP limit Carrang Stn. limit Hamelin Stn. limit Tamala Stn. limit KNP NlimitDHI 3 NlimitDHI Santalaceae Anthobolus jiJVeolalus Chorelrum prilzelii Exowrpos aphyllus Exocarpo.l' sparteus Leplomeria preissiana Leplomeria spinosa Santalum acuminalum Santalum lanceolatum Santalum spicalum Santalum spicalum subsp. novo Endemic Study atea Sapindaceae Aleclryon oleifo/ius subsp. oleifolius Diplopellis eriocarpa Diplopellis huegelii Vat. subinlegra Diplopellis intermedia Vat. incana N limit Nanga Stn. S limit KNP N limit FPNP N limit KNP N limit Cobum Stn. KNP TNR ZNP Conservation Reserve CNR FPNP BINR DINR KRNP D ip /op e / l iS i l l l e rm edwv a ri l l l e rm edw D ip /op e /u sp c u o / u n s Dodo l lu co umb / yoph s ' l /u Dodo l lu co op l e ra Dodo l lu f 'o bu(So r i i /o l io im i tZNP Nl Nl im i tDH I Dodo ! laca ( 'acs [J l {osa [ )odo f l l l ca Nl im i tFPNP Nl im i tKNP ' ~ Dodo l lo eu /u r r co id e s Dodo l lo co lobu /o lo Dodo ! l l lCa p c t io !a r i s Dodo fW ( 'O p l f l l /O /W Dodmw eu ! ' IU r l l I ICu e /o /w Nl im i tFPNP [ )odona caV lscosa s ub sp l1ngus t lsS l tJW Dodo l lu cu " ISCOSUs ub sp ,spu lu l i l lu S c rophu l a r i a c e a e *D i s ch i sma a r ena r ium E /u cho lo l lw I l I Im i l G lo s so s l igmu d iu l ld rum G /o s so s l igmu d rummo l ld i i M imu lu sg raC i l i s M imu lu sr ep e l l s P ( 'p l id lum u i rho ch e t lum P ep l id ium mu e l / en P ep l id lUm s p ,(NT Bu rb idg e& A K a i n 8152 ) S r emodw/ lo ru l e l l ( ( i S t emod iag ro s sa 5 ' t emod lO \ ' lSCO sa S rngu cu rn / lo ra So l an a c e a e An t lu l c e r c i sg en i s l l l id e s A l l r /w c e rC l sI l iC l j ( l lws ub sp ,ca ldu r l l l la An tho c erC l sl l i e r /o l i "s ub sp i l iC l / i l l iu An tho c e r c i sm ln ca la A l I l l l l l c e r c i sI i { {o r eu A I I I ! ro c e r c i ss p ,Sh a rkB a y(T ,E ,H ,Ap l in3335 ) An tho l ro ch er t l ) 'o J1o ro id e s A I l I ! ro r ro ch eH 'U /CO { { I I C yp ! ru l l rh e ra ra c emo su *Durura /eichlwrdlil Dubo i s iu IwpH 'ood l l L y c iumuu s t ra . ! e *Lyc iumf e ro c i s s imum N i co f iana ( , l lv ico !a *N ico / ia f /a ~ N i co r i i l f lU o c c id en tu / ISs ub sp h e sp en s N i co l iu l lu o c c id c l lw l i ss ub sp ob l iquu N ico t {a f la r o (und i lo { ia N i co [ iana s imu /a t1 s im i tCNR Nl Nl im i tKNP im i tWHA Nl 2 3 End em i cC ap eCuv i e r T am a l aS t n , Nl im i tM e adow S t n Familyrraxon * * Solanum Solanum Solanum Solanum Solanum Solanum Solanum Solanum Solanum Solanum Solanum Solanum Solanum Conservation Code americanum cleislogamum diversiflorum esuriale hesperium horridum lasiophyllum nigrum oldfieldii orbiculatum subsp. orbiculalum phlomoides slurtianum symonii N limit S of WHA Stackhousiaceae Slackhousia dielsii Stackhousia megaloplera Stackhousia mOf!ogyna Stackhousia muricala Slackhousia pubescens Stackhousia viminea Triplerococcus brunonis Sterculiaceae Brachychiton gregorii Commersonia crispa Commersonia gaudichaudii Commersonia melanopelala Commersonia microphylla Guichenolia all micranlha (Wittwer 18(9) Guichenolia ledifo/ia Guichenolia macrantha Guichenolia micrantha Hannafordia quadrivalvis Keraudrenia hermanniifolia Keraudrenia inlegrifolia Lasiopetalum anguslifo/ium Lasiopetalum oldfieldii ulSiopellllum opposilifo/ium Ru/ingia craurophylia Rulingia cygnortlm Ru/ingia densiflora Rulingia kempeana Ru/ingia IUleij]ora Rulingia malvifolia var. borealis Wallheria indica Distribution N limit just S of WHA N limit KNP N limit KNP N limit Tamala Stn. NlimitDHI 3 3 N limit DlNR NlimitDHI KNP TNR Conservation Reserve ZNP CNR FPNP BINR D1NR KRNP S ty l id ia c ea e L e l ' e l lhoo kw !C l ' ta l l t /w toma ( 'U la la L e v enh (wk ia (Jc S t y l id iu l I lb ru rw l lw l lu l I l sub sp .h rWW l l l l l l l l l l l l S t v l id lU l I l ca / ca ra tu l I l S t v ! l d lU l I lc ra s s t f i J l lU l I l S t v l id lU I I ld l c /w to l l lU I I l S t y l id iu l I l, l t sp e rmu l I l S t y l id iu l I ln om e S t v l id ium ! ep toph v l lu l I l S t y l id ium ! l I l lha tum ~ fWUToca rpum S t y l id lU l I lp t l l / e rU I I l S t y l id iumpU / J I / i e ru l I l ep e l l s S t y l id lU I I lr S t y l id iu l I lsp K a lb am (AC a r r145 ) S t y l i d iu rn ~ ~.' Su r iana c ea e S t y !oba s l l lmau s t ra ! e S t y !oba s iw l l spa th l l !a tw l l Nl im i tKNP Nl im i tKNP Nl im i tKNP Nl im i tT am a l aS tn . Nl im i tKNP Nl im i tN e r r en N e r r en S tn . lCNR Nl im i tKNP Nl im i tKNP Nl im i tCNR Nl im i tT am a l aS tn Nl im i tKNP Nl im i tKNP Nl im i tT am a l aS tn . Nl im i tMu r ch i son Hou s eS tn Tama r i ca c ea e *Tamanxaphy l la Th e lyp t e r ida c ea e C y c lo so l 'U sI l l t e r r l lp t l l s Thym e la ea c ea e P i l l l e ! ca w l l l l lo c /w r i s P im e ! ca a l l / iU s t i / i l l ia lonhw l I ja P i l l l e ! eaj P im e ! ea / i i ! / i ia l la P im e ! ea ! e l lC l . l I I t h a P im e ! ea l I l i c ro c e l 'ha !asub sp m i c ro c ep /w !a P i l l l e / eas e s s i l l s P im e / ea t r i c /w s ta ch ya T i l ia c ea e Co r c lw l 'U sC t ! IZ ! l l 'hun / i i l i l l s Co r c lw ru se !a cho ca r l 'u s Co r cho r l l s sp .C am a rvon(A .M .A shby2321 ) Co r c /w ru s wa ! co l l " T r iw l l ( e t /asp .T K imb e r l ey F lo r a T r emand ra c ea e T e t ra th e ca h i r su ta Typha c ea e Typ lw d ( )m ing en s i s Nl im i tKNP Nl im i tDH I Nl im i tT am a l aS tn . am a l aS tn Nl im i tT Farnilyffaxon Conservation Code Distribution Urticaceae Parietaria cardiostegia Parietaria debilis • Urtica urens Violaceae Hybanthus calycinus Hybanthus .floribundus subsp. .floribundus N limit Murchison House Stn. N IimitZNP Vitaceae Clematicissus angustissima N limitZNP Xanthorrhoeaceae Xanthorrhoea aff drummondii Xanthorrhoea preissii N limit KNP Zannichelliaceae Lepilaena preissii Zygophyllaceae Nitraria billardierei Tribulus astrocarpus Tribulus cistoides Tribulus .!(Jrrestii Tribulus macrocarpus Tribulus occidentalis Tribulus platypterus Tribulus sp. (KRN 11569) Tribulus terrestris Zygophylium aff. simile (RD Royce 5997) Zygophylium ammophilum Zygophylium apicularum Zygophylium aurantiacum Zygophylium biliardierei Zygophylium compressum Zygophylium eremaeum Zygophylium fruticulosum Zygophylium glaucum Zygophylium iodocarpum Zygophylium kochii Zygophylium lobularum Zygophylium ovatum Zygophylium retivalve Zygophylium simile Zygophylium sp. Karratha (J .S.Beard 3508) Zygophylium tesquorum S limit DH! S limit Harnelin Stn. NlimitDHl NlimitDHl S limit Carrang Stn. KNP TNR ZNP Conservation Reserve CNR FPNP BINR DINR KRNP

RELATED PAPERS

RELATED TOPICS

Log In

Flora and vegetation of the southern Carnarvon Basin, Western Australia

Flora and vegetation of the southern Carnarvon Basin, Western Australia

Related Papers

RELATED PAPERS

RELATED TOPICS