http://www.collembola.org/taxa/neaninae.htm - Last updated on 2024.04.26 by Frans Janssens
Checklist of the Collembola: Neanurinae

Subfamilia Neanurinae Börner C, 1901:33 sensu Cassagnau, 1989 key  key

Genus Number
of species
Adbiloba6
Afrobella9
Albanura2
Americanura24
Archilobella0
Archimorulodes0
Archineanura0
Australonura22
Balkanura3
Bilobella15
Blasconura12
Blasconurella5
Caledonimeria1
Caledonura1
Calvinura2
Camerounura1
Cansilianura1
Cassagnaua1
Catalanura3
Caucasanura2
Chaetobella9
Chirolavia2
Christobella1
Coecoloba7
Coreanura1
Crossodonthina18
Cryptonura12
Deuterobella4
Deutonura64
Digitanura1
Echinanura2
Ectonura19
Edoughnura2
Elgonura1
Endonura56
Galanura1
Ghirkanura1
Gnatholonche25
Graniloba2
Hazaranura1
Hemilobella8
Himalmeria (Himalmeria)3
Himalmeria (Yetimeria)17
Himalmeria20
Honduranura1
Hyperlobella1
Imparitubercula1
Inameria2
Intricatonura1
Itanura1
Kalanura4
Lathriopyga11
Lobella18
Lobellina20
Metanura7
Monobella10
Morulodes4
Nahuanura5
Neanura11
Neanuratobeassignedtopropergenus31
Neanurella1
Nepalanura1
Nepalimeria6
Nilgirella5
Oregonanura1
Paleonura53
Palmanura23
Paralobella14
Paramanura2
Paranura42
Paravietnura3
Parectonura1
Parvatinura5
Penelopella2
Persanura2
Phradmon6
Phylliomeria4
Pronura58
Propeanura13
Protanura12
Protolobella2
Pseudadbiloba1
Pseudobiloba1
Pseudoxenylla1
Pumilinura7
Rambutanura5
Riozura1
Sensillanura9
Siamanura15
Singalimeria1
Speleonura1
Sphaeronura3
Stenomeria1
Sulobella1
Synameria3
Tabasconura1
Tamulmeria2
Tetraloba1
Thaianura1
Thaumanura5
Travura3
Vietnura1
Vitronura24
Womersleya3
Xylanura1
Yosialina4
Yuukianura10
Zelandanura1
Neanurinae858
 Genera with number of species
Remark by ICZN. 2007:543: Placed on the Official List of Names in Zoology. Opinion 435.

Neanurinae from Singapore
2007 © Anker, A.
Neanurinae from Croatia
2008.05.25 © Keresztes, G.
Neanurinae from China
2012.10.18 © Deart, Y.

  Syn.: Neanuroidea) Neanirinae(sic) Shveenkova, YB, 2017:76 ISS
  Syn.: Neanuridae Nenurinae(sic) Smolis, A et Kuznetsova, N, 2018:53 ISS
Neanurinae from Singapore
2020.06.15 © Luna, J.




Genus Pseudoxenylla Christiansen, KA et Pike, E, 2002:175, -

Species fovealis (Christiansen, KA & Pike, E, 2002:176, -) Christiansen, KA & Pike, E, 2002:176, -, t.t.


Genus Echinanura Carpenter, 1935

Species elegans Carpenter, 1935

Species pacifica (Carpenter, 1934) Yosii, R, 1967:268


Genus Pseudadbiloba Massoud, 1963

Species africana Massoud, 1963


Genus Pseudobiloba Stach, 1951

  Syn.: Pseudachorutinae Achorutes Bonet, F, 1930:113
  Syn.: Pseudachorutinae Neanura Bonet, F, 1930:113


Species tenebrarum (Absolon, K, 1901:575) Stach, 1951, t.t.


Genus Caledonimeria Delamare Deboutteville, C et Massoud, Z, 1962:331

Species mirabilis Delamare Deboutteville, C et Massoud, Z, 1962:332, t.t.

Caledonimeria mirabilis from New Caledonia
2006 Thibaud, J.-M. © MNHN.
Caledonimeria mirabilis from New Caledonia
2019.03.09 © Brouste, D.
Caledonimeria mirabilis from New Caledonia
2019.03.15 © Brouste, D.
Caledonimeria mirabilis from New Caledonia
2019.11.13 © Brouste, D.
Caledonimeria mirabilis from New Caledonia
2019.11.13 © Brouste, D.

Tribus Neanurini Börner, 1901 sensu Smolis A & Pasnik G, 2020:10 key
            Syn.: Achorutinae Achorutini Börner, C, 1906:160
             Syn.: Neanurinae Crossodonthini(sic) Massoud, Z, 1967:342 partim ISS
            Syn.: Neanurinae Protanurini Massoud, Z, 1967:284 partim
             Syn.: Neanurinae Bilobellienne(sic) Cassagnau, P, 1983:12 ISS
             Syn.: Neanurinae Sensillanurienne(sic) Cassagnau, P, 1983:10 ISS
             Syn.: Neanurinae Sensillanurine(sic) Palacios-Vargas & Najt, 1986 ISS
             Syn.: Neanurinae Sensillanurine(sic) Palacios-Vargas, JG, 1996:822 ISS
             Syn.: Neanurinae Sensillanurina(sic) Palacios-Vargas, JG et Ríos, G, 1996:442 ISS
            Syn.: Neanurinae Sensillanurini Smolis, A et Pasnik, G, 2020:11
            Syn.: Neanurinae Paranurini Deharveng, L, 2004:424
            Syn.: Neanurinae Paranurini Smolis, A et Pasnik, G, 2020:11



             Syn.: Neanurinae Néanurienne(sic) Cassagnau, P, 1983:6 ISS
Neanurinae lignée néanurienne
Cephalic tubercles
After Deharveng L, 1983 Fig.13B
Neanurinae lignée néanurienne
Cephalic tubercles
After Deharveng L, 1983 Fig.13C
Neanurinae lignée néanurienne
Cephalic tubercles
After Deharveng L, 1983 Fig.13D
Neanurini from New Zealand
2006 Minor, M. & Robertson, A. © SoilBugs
Neanurini from Malaysia
2008.08.22 © Chin, H.-S.
Neanurini from Indonesia
2013.06.18 © Koh, J.
Neanurini from New Zealand
2014.mm.dd © Murray, A.
Neanurini from Tasmania
2014.04.04 © Murray, A.
Neanurini from New Zealand
2017.08.06 © Maurice
Neanurini from New Zealand
Pointed mouthcone
2018.09.10 © Schulz, K.
Neanurini from New Zealand
2018.09.10 © Schulz, K.
Neanurini from France
Neanura muscorum, Bilobella aurantiaca, & B. braunerae
2019.01.14 © Lebeaux, P.
Neanurini from Taiwan
2021.01.26 © Cheng, H.-J.
Neanurini from Taiwan
2021.02.02 © Cheng, H.-J.
Neanurini from Taiwan
2021.06.16 © Cheng, H.-J.
Neanurini from Taiwan
2021.06.16 © Cheng, H.-J.
Neanurini from Australia
2021.07.06 © Hobern, D.
Neanurini from Australia
2021.07.06 © Hobern, D.
Neanurini from Taiwan
2021.08.05 © Cheng, H.-J.
Neanurini from Taiwan
2021.08.12 © Cheng, H.-J.
Neanurini from Taiwan
Pointed mouthcone
Linea ventralis
2021.08.12 © Cheng, H.-J.
Neanurini ♀ juv. from Taiwan
Ventral aspect
Abd.6 under abd.5
2021.12.29 © Cheng, H.-J.
Neanurini ♀ juv. from Taiwan
Ventral aspect
Ventral abd.2-4 reduced
2021.12.29 © Cheng, H.-J.
Neanurini from China
Ocelli red
2023.06.17 © Wang, G.
Neanurini from China
Ocelli black
2023.06.18 © Wang, G.
Neanurini from New Zealand
2023.07.22 © Jewell, T.
Neanurini from New Zealand
2023.07.22 © Jewell, T.
Neanurini from Australia
2023.07.22 © Watts, Br.
Neanurini from Australia
2023.07.22 © Watts, Br.
Neanurini juv.? from New Zealand
2023.08.04 © Romera, P.
Neanurini juv. from New Zealand
Cephalic tubercles
2023.08.04 © Romera, P.
Neanurini juv. from New Zealand
Unguiculus absent
2023.08.04 © Romera, P.




Genus Archineanura Smolis, A et Pasnik, G, 2020:9


Genus Edoughnura Deharveng, L, Hamra-Kroua, S et Bedos, A, 2007:58

Species sp. Bendjaballah, M, Zoughailech, A, Brahim-Bounab, H, Hamra-Kroua, S, Bedos, A et Deharveng, L, 2018:398

            Syn.: Edoughnura sp. Zoughailech, A, 2017:90 u


Edoughnura sp. from Algeria
After Zoughailech A, 2017 Fig.45
Edoughnura sp. from Algeria
After Bendjaballah M & al, 2018 Fig.7

Species rara Deharveng, L, Hamra-Kroua, S et Bedos, A, 2007:58, t.t.

Edoughnura rara from Algeria
After Deharveng, L & Hamra-Kroua, S, 2008 ©


Genus Pumilinura Cassagnau, 1979 key

Species albanica Peja & Deharveng, 1995

Species croatica Smolis, A et Skarzynski, D, 2004:9

Species gneiweri (da Gama, 1963) Barra, 1993:76

Species hidalgoi Cassagnau, P et Palacios-Vargas, JG, 1983:3

             Syn.: Pumilinura hildagoi(sic) Barra, J-A, 1993:77 ISS

Species loksai (Dunger, 1974) Barra, J-A, 1993:77

Species travei Cassagnau, P et Peja, N, 1979:220, t.t.

Pumilinura travei
Dorsal tubercles
After Deharveng L, 1983 Fig.23C

Species vickae Barra, 1993:74


Genus Protanura Börner, C, 1906:167 key

  Syn.: Protanurini Lobella Massoud, Z, 1967:287 partim
  Syn.: Protanurini Neanura Massoud, Z, 1967:287 partim
  Syn.: Protanurini Yuukianura Massoud, Z, 1967:287
  Syn.: Lobellini Yuukianura Deharveng, L et Greenslade, P, 1992:737
  Syn.: Protanurini Yosialina Massoud, Z, 1967:287


Species aleo (Christiansen, K et Bellinger, P, 1992:72)

Species burti Fernando, 1957

Species capitata Folsom, JW, 1932:56

Species carpenteri Mukerji, 1932

             Syn.: Protanura carpenter(sic) Mandal, GP et Hazra, AK, 2009:2 ISS

Species citronella (Carpenter, GH, 1904:303) Bellinger, PF & Christiansen, KA, 1974:32

Neanura citronella from Hawaii
After Carpenter GH, 1904 Pl.IX Fig.20

Species deharvengi Thibaud & Massoud, 1980

Species lutea Cassagnau, P et Peja, N, 1979:219

Species mediterranea Stach, J, 1967:399

Protanura mediterranea from Malta
After Stach J, 1967 Pl.XXIX Fig.1

            Syn.: Protanura pseudomuscorum Pfliegler, WP et Janssens, F, 2016:204



Protanura mediterranea
Dorsal tubercles
After Deharveng L, 1983 Fig.23H

Species monticellii Caroli, 1912

            Syn.: Protanura monticellii Caroli, E, 1910:321 nn


Species papillata Cassagnau, P et Delamare Deboutteville, C, 1955:374

Species pseudomuscorum (Börner, 1903 tt) Caroli, 1912

            Syn.: Protanura mediterranea Dallai, R, 1973:500
            Syn.: Protanura monticellii Dallai, R, 1973:501
            Syn.: Protanura papillata Dallai, R, 1973:501


Protanura pseudomuscorum from Algeria
After Boldsystems.org, 2006 ©
Remark by Deharveng, L., Hamra-Kroua, S. & Bedos, A. 2007:58: This species and P. monticellii Caroli, 1912 requires redescription from topotypic material.

Protanura cf. pseudomuscorum from Algeria
After Zoughailech A, 2017 Fig.55
Protanura cf. pseudomuscorum from Algeria
After Zoughailech A, 2017 Fig.56
Protanura pseudomuscorum from Algeria
After Bendjaballah M & al, 2018 Fig.8

Species quadrioculata (Börner, 1901) Börner, C, 1906:167


Genus Lathriopyga Caroli, 1912 key

  Syn.: Neanura Lathriopyga Caroli, E, 1910:321 u
  Syn.: Neanuridae Beanura(sic) Lawrence, PN, 1970:151 lapsus ISS


Lathriopyga
Cephalic tubercles
After Deharveng L, 1983 Fig.20H
Lathriopyga
Dorsal tubercles
After Deharveng L, 1983 Fig.23I

  Syn.: Neanurinae Lathiopyga(sic) Palacios-Vargas, JG et Najt, J, 1986:24 ISS
Lathriopyga longiseta from the UK
Abd.6 hidden underneath abdomen
2009.04.08 © Crone, S.H.
Lathriopyga longiseta from the UK
Tertiary cuticular granules
2013.01.07 © Murray, A.
Lathriopyga longiseta from the UK
"Missing" 6th abdominal segment
2015.01.28 © Murray, A.



Species belisaria Cassagnau, P et Peja, N, 1979:211

Species bulgarica Smolis, A, Skarzynski, D et Pomorski, RJ, 2004:158

            Syn.: Lathriopyga hellenica Pomorski & Skarzynski, 1999


Species delmasi Massoud, Z, 1963:73

Species hellenica Ellis, WN, 1974:112

Species krizevacensis Barra, J-A, 1993:72

Species longiseta (Caroli, 1912 tt) Stach, J, 1951:71

            Syn.: Neanura longiseta Caroli, E, 1910:321 nn
            Syn.: Achorutes (Lathriopyga) longisetus var. flava Caroli, 1912
            Syn.: Neanura longiseta f. flava Caroli, E, 1910:321 nn
Remark by Busmachiu, G., Deharveng, L. & Weiner, W.M. 2010:57: Lathriopyga longiseta flava Caroli, 1912 is a species inquirenda because of an inadequate original description. It could be an albino form of L. longiseta or a good species.


            Syn.: Neanura hystrix Bagnall, RS, 1940:170


             Syn.: Achorutes longiseta(sic) Tarsia, 1936 ISS
Neanura longiseta from Madeira
Dorsal chaetotaxy
Modified after da Gama MM, 1964 Fig.31


             Syn.: Achorutes longisetis(sic) Massoud, Z, 1967:322 ISS
Neanura longiseta from the UK
After Lawrence PN, 1970 Fig.1
Neanura longiseta from the UK
After Lawrence PN, 1970 Fig.15
Beanura(sic) longiseta from the UK
After Lawrence PN, 1970 Fig.2
Lathriopyga longiseta from Greece
Cephalic and prothoracic dorsal chaetotaxy
After Ellis WN, 1976 Fig.30
Lathriopyga longiseta
After Deharveng L, 1983 Fig.1D
Lathriopyga longiseta from the UK
After Hopkin, S.P., 1997 ©
Lathriopyga longiseta from the UK
After Hopkin, S.P., 2004 ©
Lathriopyga longiseta from the UK
After Hopkin, S.P., 2004 ©
Lathriopyga longiseta from the UK
After Hopkin SP, 2007 Fig.7d
Lathriopyga longiseta from the UK
2007.03.04 © Cornwall, N.J.
Lathriopyga longiseta from the UK
2009.04.08 © Crone, S.H.
Lathriopyga longiseta from the UK
2013.01.04 © Murray, A.
Lathriopyga longiseta from the UK
Pointed buccal cone
2013.01.04 © Murray, A.
Lathriopyga longiseta from the UK
Prothoracic dorso-internal-tubercles divergent
2013.01.07 © Murray, A.
Lathriopyga longiseta from the UK
Cephalic posterior dorso-internal tubercles fused with dorso-external (**)
2015.01.24 © Murray, A.
Lathriopyga longiseta from the UK
Cephalic anterior tubercles fused but with granulated fields present
2015.01.24 © Murray, A.
Lathriopyga longiseta from the UK
Abd.6 (partly) hidden under abd.5
2015.01.25 © Murray, A.
Lathriopyga longiseta from the UK
Abdominal macrosetae long and curved
2015.01.25 © Murray, A.
Lathriopyga longiseta from the UK
Dorso-internal tubercles on abd.4 fused
2015.01.28 © Murray, A.
Lathriopyga longiseta juv. from the UK
2018.04.16 © Robson, S.
Lathriopyga longiseta from the UK
2018.06.10 © Robson, S.
Lathriopyga longiseta from Poland
2023.08.21 © Huskens, M.-L.
Leg.: Reyners, D.



Species microchaeta Cassagnau, P et Peja, N, 1979:212

Species montana Deharveng, L, 1985:213

Lathriopyga montana from France
After Deharveng, L., 1985 Fig.1A

Species nistru Busmachiu, G, Deharveng, L et Weiner, WM, 2010:54

Lathriopyga nistru from Moldova
After Busmachiu G & al, 2010 Fig.1
            Syn.: Lathriopyga bulgarica Busmachiu & Deharveng, 2008



Species primigenia Cassagnau, P et Peja, N, 1979:213

Species willemi Deharveng, L, 1985:216

Lathriopyga willemi from Greece
After Deharveng, L., 1985 Fig.2A


Genus Balkanura Cassagnau, 1979 key

Species cainadasi Cassagnau, P et Peja, N, 1979:207

Balkanura cainadasi
Dorsal tubercles
After Deharveng L, 1983 Fig.23J
             Syn.: Balkanura caindasi(sic) Smolis, A, 2003:313 ISS


Species jugoslawica (Palissa & Zivadinovic, 1974) Cassagnau, 1979, t.t.

Balkanura jugoslawica
Dorsal tubercles
After Deharveng L, 1983 Fig.23K
             Syn.: Balkanura jugoslavica(sic) Smolis, A, 2003:313 ISS


Species wiktori Smolis, A, 2003:315


Genus Monobella Cassagnau, 1979 key

Monobella
Cephalic tubercles
After Deharveng L, 1983 Fig.20I
  Syn.: Neanuridae Mnobella(sic) Shaw, P, 2017:172 ISS


Species calva Bedos, A et Deharveng, L, 1998:292

Species cassagnaui Deharveng, L, 1981:98

Monobella cassagnaui
Cephalic tubercles
After Deharveng L, 1983 Fig.20J
Monobella cassagnaui
Dorsal tubercles
After Deharveng L, 1983 Fig.23N

Species dubia Deharveng, 1986

Species edaphica Bedos, A et Deharveng, L, 1998:294

Monobella edaphica from France
After Bedos A & Deharveng L, 1998 Fig.4B

Species gersi (Deharveng, 1979) Deharveng, 1986

Species grassei (Denis, 1923) Cassagnau, 1979, t.t.

            Syn.: Achorutes Grassei(sic) Denis, J-R, 1933:229 IOS
             Syn.: Neanura graccei(sic) Bagnall, RS, 1940:163 ISS
             Syn.: Bilobella Grassei(sic) Travé, J, Gadea, E et Delamare Deboutteville, C, 1954:209 ISS
Monobella grassei
After Deharveng L, 1983 Fig.1E
Monobella grassei
Dorsal tubercles
After Deharveng L, 1983 Fig.23M
Monobella grassei from the UK
After Hopkin, S.P., 2004 ©
Monobella grassei from the UK
After Hopkin, S.P., 2004 ©
Monobella grassei from the UK
After Hopkin, S.P., 2004 ©
Monobella grassei from the UK
2007 © Cornwall, N.J.
Monobella grassei from the UK
2008.04.30 © Farrell, S.
Monobella grassei from the UK
Pointed mouthcone
2009.02.20 © Michaelson, J.
Monobella grassei from the UK
2011.12.09 © Nurcombe, E.
Monobella grassei from the UK
Ocular tubercle with 2 separate ocelli
2012.01.24 © Jones, A.
Monobella grassei from the UK
Pointed mouthcone
2012.01.24 © Jones, A.
Monobella grassei from the UK
Pointed mouthcone
2012.03.22 © Nurcombe, E.
Monobella grassei from the UK
Notice curled-up phoretic Nematoda
2012.10.20 © Murray, A.
Monobella grassei from the UK
6th abdominal segment hidden under 5th
2012.10.20 © Murray, A.
Monobella grassei from France
2012.11.02 © Quintin, C.
Monobella grassei from France
Two distinctly separated ocelli
2012.11.05 © Quintin, C.
Monobella grassei from the UK
Macrosetae on 1st abdominal segment about the same length as the segment
2013.02.04 © Nurcombe, E.
Monobella grassei from the UK
Head
2013.02.04 © Nurcombe, E.
Monobella grassei from the UK
Antenna with curved blunt sensilla
2013.02.04 © Nurcombe, E.
Monobella grassei from the UK
Two distinctly separated ocelli
2014.01.16 © Phillips, E.
Monobella grassei from the UK
Pointed mouthcone
2014.01.16 © Phillips, E.
Monobella grassei from the UK
Two distinctly separated ocelli
2015.12.13 © Horton, M.
Monobella grassei from the UK
Pointed mouthcone
2015.12.13 © Horton, M.
Monobella grassei from Holland
2016.01.15 © Krediet, A.
Monobella grassei from France
Abd.6 hidden under abd.5
2016.12.03 © Garcelon, P.
Monobella grassei from France
Distinctly pointed mouthcone
2017.01.13 © Garcelon, P.
Monobella grassei from Belgium
2018.01.03 © Krediet, A.
Monobella grassei from France
Cephalic tubercles
2023.03.12 © Clerc, O.
Monobella grassei from France
With phoretic Nematoda
curled-up around dorsal macroseta
2024.01.21 © Garcelon, P.

  Subspecies banyulensis Deharveng, L, 1981:100

  Subspecies distincta Deharveng, 1979

Monobella distincta
Dorsal tubercles
After Deharveng L, 1983 Fig.23L

  Subspecies minor Bedos, A et Deharveng, L, 1998:297

  Subspecies prodistincta Deharveng, 1986





Species jau Bedos, A et Deharveng, L, 1998:293

Monobella jau from France
After Bedos A & Deharveng L, 1998 Fig.4A

Species pyrenaica (Deharveng, 1979) Deharveng, 1986

Monobella pyrenaica from France
After Boldsystems.org, 2006 ©

Species rostrata Deharveng, 1986

Species rousseti Bedos, A et Deharveng, L, 1998:294

Monobella rousseti from France
After Bedos A & Deharveng L, 1998 Fig.5A


Genus Endonura Cassagnau, 1979 key

Endonura
Cephalic tubercles
After Deharveng L, 1983 Fig.20C
Endonura
Cephalic tubercles
After Deharveng L, 1983 Fig.20D
Endonura
Cephalic tubercles
After Deharveng L, 1983 Fig.20E
Endonura
Dorsal tubercles
After Deharveng L, 1983 Fig.23D
Endonura
Dorsal tubercles
After Deharveng L, 1983 Fig.23E

Species sp.1 Janssens F sensu Arbea JI, Brahim-Bounab, H & Hamra-Kroua S, 2013

            Syn.: Endonura sp.undescribed Arbea, JI, Brahim-Bounab, H et Hamra-Kroua, S, 2013:178


Species sp.2 Janssens F sensu Yahyapour E, Shayanmehr M, Vafaei-Shoushtari R & Arbea JI, 2022

            Syn.: Endonura sp.undescribed Yahyapour, E, Shayanmehr, M, Vafaei-Shoushtari, R et Arbea, JI, 2022:266


Species agnieszkae Smolis, A et Skarzynski, D, 2020:108

            Syn.: Endonura agnieskae(sic) Smolis, A et Skarzynski, D, 2020:108 IOS
Remark by Janssens, F. 2020.11.14: According to the etymology (p.108) : "The new species is dedicated to Agnieszka, wife of the first author." However, the new name refers to Agnieska, not to Agnieszka. This mismatch is clear evidence of an inadvertent error that must be corrected (Art.32.5.1). The name 'agnieszkae' is in line with the etymology.

            Syn.: Endonura agnieskae(sic) Smolis, A et Skarzynski, D, 2020:105 IOS
            Syn.: Endonura agnieskae(sic) Smolis, A et Skarzynski, D, 2020:109 IOS
            Syn.: Endonura agnieskae(sic) Smolis, A et Skarzynski, D, 2020:110 IOS
            Syn.: Endonura agnieskae(sic) Smolis, A et Skarzynski, D, 2020:111 IOS
            Syn.: Endonura agnieskae(sic) Yahyapour, E, Shayanmehr, M, Vafaei-Shoushtari, R et Arbea, JI, 2022:253 IOS
            Syn.: Endonura agnieskae(sic) Yahyapour, E, Shayanmehr, M, Vafaei-Shoushtari, R et Arbea, JI, 2022:272 IOS

Species aibgai Smolis & Kuznetsova, 2017

Species alavensis Pozo & Simón, 1981

Species alticola (Stach, J, 1951:36) Smolis, 2017

Species annae Smolis, A et Skarzynski, D, 2020:111

Species arbasensis Deharveng, 1979

Species asiatica Smolis, Deharveng & Kaprus, 2011

Species baculifer Deharveng, 1979

Endonura baculifer
After Deharveng L, 1983 Fig.1B
Endonura baculifer
Cephalic clypeal and antennofrontal tubercles
After Deharveng L, 1983 Fig.14A

Species caeca (Gisin, 1963) Smolis, A, Kahrarian, M, Piwnik, A, & Skarzynski, D, 2016:66

             Syn.: Neanura coeca(sic) Thibaud, J-M et Massoud, Z, 1973:164 ISS

Species cantabrica Deharveng, 1979

Endonura cantabrica from Romania
After Boldsystems.org, 2006 ©

Species carpatica Smolis, 2006

Species centaurea (Cassagnau, P & Peja, N, 1979:216) Pomorski, RJ & Skarzynski, D, 2000:118

Species ceratolabralis Smolis, A, Kahrarian, M, Piwnik, A, et Skarzynski, D, 2016:59

Species colorata (da Gama, 1964)

            Syn.: Neanura stachi Gisin, 1952 partim


Species cretensis (Ellis, WN, 1976:277)

Endonura cretensis from Israel
After Smolis A & Kaprus' IJ, 2009 Fig.1

Species cryptopyga Smolis & Kuznetsova, 2017

Species dalensi (Deharveng, 1979) Dallai, 1980

Endonura dalensi from Albania
After Traser G & Kontschán J, 2004 Fig.2

  Subspecies sardoa Dallai, R, [1984]:431

             Syn.: Endonura dalensi ssp. sardosa(sic) Traser, G et Kontschán, J, 2004:74 ISS

Species deharvengi (Cassagnau, P & Peja, N, 1979:217) Smolis, A, Kahrarian, M, Piwnik, A, & Skarzynski, D, 2016:67

Species dentifera Smolis, Skarzynski, Pomorski & Kaprus, 2007

Species dichaeta Smolis, A, Kahrarian, M, Piwnik, A, et Skarzynski, D, 2016:55

Species diminutichaeta Smolis & Kuznetsova, 2017

Species dobrolyubovae Smolis & Kuznetsova, 2017

Species dudichi (Loksa, I, 1967:289) Smolis, A, 2008:?

Neanura dudichi from Hungary
After Loksa I, 1967 Abb.1

Species gladiirostra Smolis, A et Kaprus, IJ, 2003:327

             Syn.: Endonura gladirostra(sic) Smolis, A et Kaprus, IJ, 2003:327 ISS
             Syn.: Endonura gladirostris(sic) Fanciulli, PP et Dallai, R, 2008:23 ISS

Species gladiolifer (Cassagnau, P, 1954:29) Simón, JC, Bach, C & Gaju, M, 1987:298

Deutonura gladiolifer
Cephalic clypeal and antennofrontal tubercles
After Deharveng L, 1983 Fig.14I

Species gracilirostris Smolis, Skarzynski, Pomorski & Kaprus, 2007

Species granulata (Cassagnau, P & Delamare Deboutteville, C, 1955:376) Pomorski & Skarzynski, 2000

Species ichnusae Dallai, R, [1984]:435

Species immaculata Deharveng, 1980

Species incolorata (Stach, J, 1951:56) Smolis, A, 2008:?

Species kremenitsai Smolis & Kuznetsova, 2017

Species levantica Smolis, Deharveng & Kaprus, 2011

Species longirostris Smolis, A, Shayanmehr, M, Kuznetsova, N et Lafooraki, EY, 2017:138

Species ludovicae (Denis, J-R, 1947:3)

            Syn.: Achorutes ludovicæ(sic) Denis, J-R, 1947:3 IOS
Achorutes ludovicæ(sic) from Corsica
After Denis J-R, 1947 Fig.2



Species lusatica (Dunger, 1966) Smolis, A, 2008:?

Species occidentalis Deharveng, 1979

Species ossetica Smolis & Kuznetsova, 2017

Species paracantabrica Smolis & Kuznetsova, 2017

Species paracentaurea Smolis, A, Shayanmehr, M, Kuznetsova, N et Lafooraki, EY, 2017:142

Species pejai Deharveng, 1980

Species persica Smolis, A, Kahrarian, M, Piwnik, A, et Skarzynski, D, 2016:62

             Syn.: Endonura Persica(sic) Kahrarian, M, 2019:34 ISS
             Syn.: Endonura Persica(sic) Kahrarian, M, 2019:43 ISS

Species poinsotae Deharveng, 1980

Species quadriseta (Cassagnau, P & Peja, N, 1979:218) Fanciulli, PP & Dallai, R, 2008:23

Species reticulata (Axelson, WM, 1905:790) Babenko, 2002

            Syn.: Protanura quadrioculata Linnaniemi, 1912 =?
            Syn.: Achorutes tetrophthalmus Stach, 1929 =?
            Syn.: Neanura (Endonura) tundricola Fjellberg, 1985


Species saleri Fanciulli, PP et Dallai, R, 2008:19

Species schwendingeri Smolis, A et Skarzynski, D, 2020:115

Species tartaginensis (Deharveng, 1980) Dallai, 1983

             Syn.: Endonura tartaginenis(sic) Smolis, A, Kahrarian, M, Piwnik, A, et Skarzynski, D, 2016:58 ISS

Species tatricola (Stach, J, 1951:53) Gruia, 1975

            Syn.: Neanurella szeptyckii Weiner, 1973


Species taurica (Stach, J, 1951:34) Kaprus, IJ, Pomorski, RJ, Skarzynski, D & Potapov, MB, 2005:893

Species tetrophthalma (Stach, J, 1930:282) Cassagnau, 1979, t.t.

            Syn.: Biloba tetrophthalma f. principalis Stach, J, 1951:52


             Syn.: Achorutes tetrophtalmus(sic) Smolis, A et Kaprus, IJ, 2009:1 ISS
             Syn.: Endonura tetrophtalma(sic) Smolis, A, Kahrarian, M, Piwnik, A, et Skarzynski, D, 2016:67 ISS

Species transcaucasica (Stach, J, 1951:34) Fanciulli, PP & Dallai, R, 2008:22

Endonura transcaucasica
After Boldsystems.org, 2006 ©

Species turkmenica Smolis, A, Shayanmehr, M, Kuznetsova, N et Lafooraki, EY, 2017:145

Species tyrrhenica Dallai, R, [1984]:437

Species urotuberculata Pomorski, RJ et Skarzynski, D, 2000:118

Endonura urotuberculata from Bulgaria
After Pomorski RJ & Skarzynski D, 2000 Fig.1A


Genus Cryptonura Cassagnau, 1979 key

Species sp.1 Janssens F sensu Yahyapour E, Shayanmehr M, Vafaei-Shoushtari R & Arbea JI, 2022

            Syn.: Cryptonura sp.undescribed Yahyapour, E, Shayanmehr, M, Vafaei-Shoushtari, R et Arbea, JI, 2022:265


Species abmalensis Arbea JI, Yahyapour E et Shayanmehr M in Yahyapour, E, Shayanmehr, M, Vafaei-Shoushtari, R et Arbea, JI, 2022:258

Cryptonura abmalensis from Iran
After Yahyapour E & al, 2022 Fig.16-17

Species anthrenoidea (Ellis, WN, 1976:278)

Species dirfysensis (Cassagnau, P & Peja, N, 1979:214) Smolis, A, 2002:325

Species dohezarensis Arbea JI, Yahyapour E et Shayanmehr M in Yahyapour, E, Shayanmehr, M, Vafaei-Shoushtari, R et Arbea, JI, 2022:254

Cryptonura dohezarensis from Iran
After Yahyapour E & al, 2022 Fig.1-2

Species ellisi Smolis, A, Skarzynski, D et Gwiazdowicz, DJ, 2023:345

            Syn.: Protonura sp. Gwiazdowicz & al, 2022
            Syn.: Protonura sp. Smolis, A, Skarzynski, D et Gwiazdowicz, DJ, 2023:333


Species franzi (Stach, J, 1951:70) Cassagnau, 1979, t.t.

            Syn.: Lathriopyga Franzi(sic) Stach, J, 1951:70,tt IOS

Species jubilaria Smolis, A, 2002:326

Species kuehnelti (Gisin, 1954) Deharveng, L, 1983:12

             Syn.: Cryptonura kühnelti(sic) Smolis, A, Falahati, A et Skarzynski, D, 2012:58 ISS

Species maxima Smolis, A, Falahati, A et Skarzynski, D, 2012:55

Species persica Smolis, A, Falahati, A et Skarzynski, D, 2012:52

Species sariensis Arbea JI, Yahyapour E et Shayanmehr M in Yahyapour, E, Shayanmehr, M, Vafaei-Shoushtari, R et Arbea, JI, 2022:261

Cryptonura sariensis from Iran
After Yahyapour E & al, 2022 Fig.26
Cryptonura sariensis from Iran
After Yahyapour E & al, 2022 Fig.27


Genus Neanurella Cassagnau, 1968

Species microphthalma Cassagnau, 1968, t.t.


Genus Deutonura Cassagnau, 1979 key

  Syn.: Achorutes phlægreus(sic) Denis, 1938
  Syn.: Achorutes phlegraeus f. implena(sic) Stach J in Kseneman, M, 1938:467 nn IOS
  Syn.: Achorutes phlegraeus f. implena(sic) Deharveng, L, Ait Mouloud, S et Bedos, A, 2015:470 IOS
  Syn.: Deutonura? implena Deharveng, L, Ait Mouloud, S et Bedos, A, 2015:470
  Syn.: Achorutes sp. Denis, J-R, 1947:5
  Syn.: Neanuridae Neonura(sic) Bogojevic, 1984 ISS


Deutonura
Cephalic tubercles
After Deharveng L, 1983 Fig.20F
Deutonura
Cephalic tubercles
After Deharveng L, 1983 Fig.20G
Deutonura
Dorsal tubercles
After Deharveng L, 1983 Fig.23E
Deutonura sp. from France
2020.02.06 © Dejoux, P.
Deutonura sp. from France
Abd.6 underneath abd.5
2020.02.06 © Dejoux, P.



Species sp.1 Janssens F sensu Thibaud & Massoud, 1980

             Syn.: Deutonura conjoncta(sic) Thibaud & Massoud, 1980 ISS
            Syn.: Deutonura conjuncta? Deharveng, L, Zoughailech, A, Hamra-Kroua, S et Porco, D, 2015:281

Species sp.2 Janssens F sensu Deharveng, Bedos & Duran, 2015

            Syn.: Deutonura sp.undescribed Deharveng, L, Bedos, A et Duran, V, 2015:184


Species abietis (Yosii, 1969)

Species adriani Deharveng, L, Ait Mouloud, S et Bedos, A, 2015:465

Deutonura adriani from Algeria
After Deharveng L & al, 2015 Fig.1A

            Syn.: Deutonura sp. Ait Mouloud & al, 2007



Species albella (Stach, 1921) Deharveng, L, 1983:37

Species alduidensis Arbea JI, 1987 nn

Remark by Arbea, J.I. 2007.05.07:in litt.: Unavailable manuscript name. Nomen nudum.

Species anophthalma (Massoud & Thibaud, 1968)

Species arbeai Lucianez & Simon, 1996

Species atlantica Deharveng, 1981

Deutonura atlantica from France
After Boldsystems.org, 2006 ©

Species balsainensis (Simón, 1978)

Species benzi Traser, G, Thibaud, J-M et Najt, J, 1993:221

Species betica (Deharveng, 1979) Dallai, R, [1984]:444

            Syn.: Neanura (Lathriopyga) stachi Dallai, 1969
            Syn.: Lathriopyga stachi Dallai, 1976


Species binatuber (Lee, B-H, 1974:578)

Species breviseta Smolis, A et Skarzynski, D, 2020:118

Species caerulescens Deharveng, 1982

             Syn.: Deutonura coerulescens(sic) Deharveng, L, 1983:56 ISS

Species caprai Dallai, R, [1984]:456

Species carinthiaca Deharveng, 1982

Species centralis (da Gama, 1964) Deharveng, L, 1983:34

Deutonura centralis
Cephalic clypeal and antennofrontal tubercles
After Deharveng L, 1983 Fig.14J

Species coiffaiti Deharveng, 1979

Species conjuncta (Stach, 1926) Cassagnau, P, 1983:21

            Syn.: Achorutes monoculatus Denis, 1931
            Syn.: Achorutes comatus Börner in Franz & Beier, 1948 nn


             Syn.: Achorutes conjonctus(sic) Denis, 1935 ISS
Lathriopyga conjuncta from Poland
After Stach J, 1951 Pl.1 Fig.2


             Syn.: Lathriopyga conjucta(sic) Nosek, 1957a ISS
             Syn.: Lathriopyga conjucta(sic) Nosek, 1957b ISS
             Syn.: Neanura (Lathriopyga) conjucta(sic) Cassagnau, 1966 ISS
             Syn.: Neanura conjunctata(sic) Ambroz & Nosek, 1967 ISS
             Syn.: Neanura conjucta(sic) Nosek & al, 1978 ISS
             Syn.: Neanura conjucta(sic) Falca, 1993 ISS
             Syn.: Neanura conjucta(sic) Farkas, J, 1995:46 ISS
Deutonura conjuncta from Italy
After Paolucci, P., 2006 ©
Deutonura conjuncta from Hungary
2012.05.10 © Lásló, D.
Deutonura conjuncta from Slovenia
2017.03.04 © Pisec, N.
Deutonura conjuncta from Slovenia
2017.03.04 © Pisec, N.
Deutonura conjuncta from Slovenia
Abd.6 under abd.5
2017.03.06 © Pisec, N.
Deutonura conjuncta from Slovenia
Abd.6 (*) under abd.5
2017.03.07 © Pisec, N.
Deutonura conjuncta from Slovenia
2017.03.10 © Pisec, N.
Deutonura conjuncta from France
2017.12.11 © Tiky, S.
Deutonura conjuncta from France
2017.12.11 © Tiky, S.
Deutonura conjuncta from France
2017.12.11 © Tiky, S.
Deutonura conjuncta from France
2018.01.09 © Garcelon, P.
Deutonura conjuncta from France
2018.01.09 © Garcelon, P.
Deutonura conjuncta from France
Pointed mouthcone
2018.02.08 © Garcelon, P.



Species corsica (Denis, J-R, 1947:2) Deharveng, L, Ait Mouloud, S & Bedos, A, 2015:470

Achorutes corsicus from Corsica
After Denis J-R, 1947 Fig.1

            Syn.: Deutonura insularis Deharveng, 1982
            Syn.: Deutonura insularis Deharveng, L, Ait Mouloud, S et Bedos, A, 2015:470



Species czarnohorensis Deharveng, 1982

             Syn.: Deutonura czarnohensis(sic) Deharveng, L, 1983:34 ISS
Deutonura czarnohensis(sic)
Cephalic clypeal and antennofrontal tubercles
After Deharveng L, 1983 Fig.14D



Species decolorata (da Gama & Gisin in 0381:391) Deharveng, L, 1983:34

Deutonura decolorata
Cephalic clypeal and antennofrontal tubercles
After Deharveng L, 1983 Fig.14L

Species deficiens Deharveng, 1979

Deutonura deficiens from France
After Boldsystems.org, 2006 ©
Deutonura deficiens from France
After Boldsystems.org, 2006 ©
Deutonura deficiens from France
After Boldsystems.org, 2006 ©
Deutonura deficiens from France
After Boldsystems.org, 2006 ©

  Subspecies meridionalis Deharveng, 1979

  Subspecies sylvatica Deharveng, 1982

            Syn.: Neanura plena da Gama, 1964
            Syn.: Deutonura plena ssp. plena Arbea & Jordana, 1985
            Syn.: Deutonura plena ssp. plena var. trioculata Pozo & Simón, 1982


Species deharvengi Arbea, JI et Jordana, R, 1991:100

Species dextra (Gisin, 1954)

Species ezomontana (Yosii, R, 1972:82)

Species fodinarum (Yosii, R, 1956:45)

Neanura fodinarum from Japan
After Yosii R, 1956 Pl.XXII Fig.134

Species frigida (Yosii, 1969) Deharveng & Weiner, 1984

Deutonura frigida from the USA
2022.12.16 © Parker, H.S.
Deutonura frigida from the USA
Cephalic Di and De tubercles fused
2022.12.16 © Parker, H.S.
Deutonura frigida from the USA
Abd.4 both Di tubercles fused
2022.12.16 © Parker, H.S.
Deutonura frigida from the USA
2022.12.16 © Parker, H.S.
Deutonura frigida from the USA
Dorsal cephalic tubercles
2022.12.16 © Parker, H.S.
Deutonura frigida from Canada
Cephalic Di and De tubercles fused
2023.12.20 © Barbin, T.
Deutonura frigida from Canada
Abd.4 both Di tubercles fused
2023.12.20 © Barbin, T.

Species gibbosa Porco, Bedos & Deharveng, 2010

Species gisini Deharveng, 1982

Deutonura gisini
Cephalic clypeal and antennofrontal tubercles
After Deharveng L, 1983 Fig.14H

Species granatuberis (Lee, B-H, 1974:577) Deharveng & Weiner, 1984

Neanura granatuberis from Korea
After Lee B-H, 1974 Fig.2F

Species gurreae Lucianez & Simon, 1996

Species ibicensis (Ellis, WN, 1974:128)

Species igilica Dallai, R, [1984]:451

Species ilvatica Dallai, R, [1984]:449

Species inopinata Deharveng, 1979

Species iranica Smolis, A et Skarzynski, D, 2020:125

Species jeromoltoi Deharveng, L, Bedos, A et Duran, V, 2015:184

Deutonura jeromoltoi from France
After Deharveng L, Bedos A & Duran V, 2015 Fig.8A

             Syn.: Deutonura jeremoltoi(sic) Smolis, A et Pasnik, G, 2020:6 ISS
             Syn.: Deutonura jeremoltoi(sic) Smolis, A et Pasnik, G, 2020:7 ISS
             Syn.: Deutonura jeremoltoi(sic) Smolis, A et Pasnik, G, 2020:8 ISS


Species leei Deharveng, L et Weiner, WM, 1984:19

Species luberonensis Deharveng, 1982

Deutonura luberonensis
Cephalic clypeal and antennofrontal tubercles
After Deharveng L, 1983 Fig.14F

Species mirabilis Deharveng, 1987

Species monticola (Cassagnau, P, 1954:33) Deharveng, L, 1983:8

Deutonura monticola from France
After Boldsystems.org, 2006 ©
Deutonura monticola from France
After D'Haese C in Thibaud J-M, 2017 Fig.3
Deutonura monticola from France
2018.06.19 © van Duinen, J.
Deutonura monticola from France
2018.06.19 © van Duinen, J.
Deutonura monticola from France
2018.09.23 © Garcelon, P.
Deutonura monticola from France
2018.09.25 © van Duinen, J.
Deutonura monticola from France
Ocular tubercle with 2 ocelli
2018.09.25 © van Duinen, J.
Deutonura monticola from France
2018.09.27 © van Duinen, J.
Deutonura monticola from France
2018.09.28 © Wieringa, P.
Deutonura monticola from France
Abdomen with 5 dorsally visible segments
2019.07.29 © Tiky, S.
Deutonura monticola from France
Pointed mouthcone
2019.07.29 © Tiky, S.
Deutonura monticola from France
2019.07.31 © Tiky, S.
Deutonura monticola from France
Abd.6 hidden under abd.5
2019.07.31 © Tiky, S.
Deutonura monticola from France
Leg3 with trochanter (t), femur (f), tibiotarsus (tt) and unguis (u)
2019.07.31 © Tiky, S.
Deutonura monticola from France
Unguiculus absent (*)
2019.07.31 © Tiky, S.
Deutonura monticola from France
2019.11.30 © Joachim, J.
Deutonura monticola from France
Dorso-internal tubercles of th.1 with 2 setae (*)
2019.11.30 © Joachim, J.
Deutonura monticola from France
Dorso-lateral tubercles of abd.1 with 2 setae (*)
2019.11.30 © Joachim, J.
Deutonura monticola from France
2020.02.13 © Garcelon, P.
Deutonura monticola from France
Dorsal tubercles elongated
2020.02.13 © Garcelon, P.
Deutonura monticola from France
Cephalic ocular tubercle with 2 ocelli
2020.02.13 © Garcelon, P.
Deutonura monticola from France
Cephalic tubercles
2020.02.13 © Garcelon, P.
Deutonura monticola from France
Th.1 dorso-internal tubercles with 2 setae (*)
2020.02.13 © Garcelon, P.
Deutonura monticola from France
Abd.5 with 4 tubercles; dorso-internal fused
Abd.6 underneath abd.5
2020.02.13 © Garcelon, P.
Deutonura monticola ♀ from France
Abd.5+6 ventral aspect
2020.02.13 © Garcelon, P.
Deutonura monticola ♀ from France
Gonopore with anterior and posterior lip
2020.02.13 © Garcelon, P.
Deutonura monticola ♀ from France
Left + right ventral anal valves (*)
Bilobed dorsal anal valve (**)
2020.02.13 © Garcelon, P.
Deutonura monticola from France
Collophore
2020.02.15 © Garcelon, P.
Deutonura monticola from France
Labium distinctly pointed
Linea ventralis (lv)
2020.02.15 © Garcelon, P.
Deutonura monticola from France
Foot-complex
Unguiculus absent (*)
2020.02.15 © Garcelon, P.
Deutonura monticola from France
2022.02.12 © Joachim, J.
Deutonura monticola from France
Th.1 di:de:dl tubercles
Number of setae = 2:2:1
2022.02.12 © Joachim, J.
Deutonura monticola from France
Th.2 with 2 distinct oblique folds
2024.04.25 © Massol, E.
Deutonura monticola from France
Abd.5 dorso-internal tubercles fused
2024.04.25 © Massol, E.

Species oglasicola Dallai, R, [1984]:453

Species persica Smolis, A, Shayanmehr, M et Yoosefi-Lafooraki, E, 2018:9

            Syn.: Deutonura decolorata Cox, 1982 =?
             Syn.: Neanura decolarata(sic) Falahati HosseinAbad, A, Mehr, MS et Khyroodin, A, 2013:259 ISS
            Syn.: Deutonura decolorata Shayanmehr, M, Yahyapour E, Kahrarian, M et Lafooraki EY, 2013:76 =?
            Syn.: Deutonura decolorata Smolis, A, Kahrarian, M, Piwnik, A, et Skarzynski, D, 2016:69 =?
            Syn.: Deutonura (conjuncta-group) decolorata Smolis, A, Shayanmehr, M et Yoosefi-Lafooraki, E, 2018:13 =?


Species phlegraea (Caroli, 1912) Cassagnau, 1979, t.t.

            Syn.: Neanura phlegraea Caroli, E, 1910:321 nn


             Syn.: Achorutes phlaegreus(sic) Denis, J-R, 1933:228 ISS
             Syn.: Achorutes phlægreus(sic) Denis, J-R, 1947:3 ISS
             Syn.: Achorutes Phlægreus(sic) Denis, J-R, 1947:3 ISS
             Syn.: Achorutes phlægreus(sic) Denis, J-R, 1947:5 ISS
             Syn.: Lathriopyga plegraea(sic) Stach, J, 1951:Pl.1 Fig.3 ISS
Lathriopyga plegraea(sic) from Poland
After Stach J, 1951 Pl.1 Fig.3
Deutonura phlegraea
Cephalic chaetotaxy
After Deharveng L, 1983 Fig.12C
Deutonura phlegraea
Cephalic clypeal and antennofrontal tubercles
After Deharveng L, 1983 Fig.14C

             Syn.: Deutonura phlegrea(sic) Smolis, A et Pasnik, G, 2020:6 ISS
             Syn.: Deutonura phlegrea(sic) Smolis, A et Pasnik, G, 2020:7 ISS
             Syn.: Deutonura phlegrea(sic) Smolis, A et Pasnik, G, 2020:8 ISS


Species piceae (Yosii, 1969)

Species plena (Stach, J, 1951:68) Szeptycki, A & Weiner, WM in Razowski J, 1990:20

            Syn.: Neanura phlegraea Caroli, 1912 partim


Species portucalensis (da Gama, 1964) Deharveng, 1982

Species provincialis Deharveng, 1979

Deutonura provincialis from France
After Boldsystems.org, 2006 ©
Deutonura provincialis from France
After Boldsystems.org, 2006 ©

Species quinquesetosa Deharveng, 1982

            Syn.: Neanura (Deutonura) setosa Deharveng, 1979 nec Canby, 1926
            Syn.: Neanura (Deutonura) setosa Deharveng, L, Ait Mouloud, S et Bedos, A, 2015:470


Species schulzi Smolis, A, Skarzynski, D et Gwiazdowicz, DJ, 2023:340

            Syn.: Deutonura sp. Gwiazdowicz & al, 2022
            Syn.: Deutonura sp. Smolis, A, Skarzynski, D et Gwiazdowicz, DJ, 2023:333


Species selgae Deharveng, 1979

Species sengleti Smolis, A et Skarzynski, D, 2020:122

Species similis Deharveng, 1979

Deutonura similis
Dorsal tubercles
After Deharveng L, 1983 Fig.23F

Species sinistra (Denis, 1935) Dallai & Malatesta in Calderini, 1995:6

Species stachi (Gisin, 1952) Deharveng, L, 1983:34

            Syn.: Lathriopyga phlegraea Stach, 1951, nec Caroli, 1912
            Syn.: Achorutes phlegraeus f. albella Stach, 1921


Deutonura stachi
Cephalic clypeal and antennofrontal tubercles
After Deharveng L, 1983 Fig.14B
Deutonura stachi from Romania
After Dányi, L. & Traser G, 2008 Fig.1,2

Species sylviae Dallai, R, [1984]:446

Species urbionensis Deharveng, 1979

Species vallespirensis Deharveng, 1982

Species weinerae Deharveng, 1982

Deutonura weinerae
Cephalic clypeal and antennofrontal tubercles
After Deharveng L, 1983 Fig.14G

Species yoshiiana Deharveng, L et Weiner, WM, 1984:21

Species zana Deharveng, L, Zoughailech, A, Hamra-Kroua, S et Porco, D, 2015:282

            Syn.: Deutonura sp. Deharveng, L, Hamra-Kroua, S et Bedos, A, 2007:57


Deutonura zana from Algeria
After Deharveng L et al, 2015 Fig.1
Deutonura zana from Algeria
After Deharveng L et al, 2015 Fig.2
Deutonura zana from Algeria
After Zoughailech A, 2017 Fig.40
Deutonura zana from Algeria
After Bendjaballah M & al, 2018 Fig.6
Deutonura zana juv. from Algeria
After Bendjaballah M & al, 2018 Fig.6

Species zelkovabelicea Smolis, A, Skarzynski, D et Gwiazdowicz, DJ, 2023:342

            Syn.: Endonura zelkovae(sic) Smolis, A, Skarzynski, D et Gwiazdowicz, DJ, 2023:342 IOS
            Syn.: Endonura sp. Gwiazdowicz & al, 2022
            Syn.: Endonura sp. Smolis, A, Skarzynski, D et Gwiazdowicz, DJ, 2023:333


Remark by Janssens, F. 2023.05.09: According to the etymology : "Named after the endemic tree Zelkova abelicea where new species was found." However, "zolkovae" refers to the genus "Zelkova", not to the tree "Zelkova abelicea". This meaning does not match the etymology. This mismatch is clear evidence of an inadvertent error that must be corrected (Art.32.5.1). 'zelkovabelicea' refers to 'the Endonura named after the tree Zelkova abelicea' in line with the etymology.



Genus Albanura Deharveng, 1982

Albanura
Dorsal tubercles
After Deharveng L, 1983 Fig.23I

Species nana (Cassagnau, P & Peja, N, 1979:215) Deharveng, 1982

Species secunda Smolis, A et Kuznetsova, N, 2018:3

Albanura secunda from the Caucasus
After Smolis A & Kuznetsova N, 2018 Fig.1+9


Genus Catalanura Deharveng, 1982

Catalanura
Cephalic tubercles
After Deharveng L, 1983 Fig.20H
Catalanura
Dorsal tubercles
After Deharveng L, 1983 Fig.23E

Species catalana (Deharveng, 1979) Cassagnau, P, 1983:15, t.t.

Catalanura catalana
Dorsal chaetotaxy
Modified after Deharveng L, 1983 Fig.21B

Species florae Arbea, JI et Jordana, R, 1991:100

Species najtae (Deharveng, 1979)


Genus Metanura Yosii, 1954

Species cassagnaui Deharveng, L et Weiner, WM, 1984:24

Species iriomotensis (Yosii, R, 1965:15)

Species kitayamana Yoshii, 1995

Species nagatonis Yoshii, 1995

Species okinawana (Yosii, R, 1965:13)

Species sanctisebastiani Yosii, 1954, t.t.

            Syn.: Metanura Sancti-Sebastiani(sic) Yosii, 1954,tt IOS
             Syn.: Metanura sebastiani(sic) Yosii, R, 1956:44 ISS
Neanura (Metanura) sebastiani(sic) from Japan
After Yosii R, 1956 Pl.XXIV Fig.141

             Syn.: Neanura sanctisebastini(sic) Yosii, R, 1965:14 ISS
Metanura sanctisebastiani
Dorsal tubercles
After Deharveng L, 1983 Fig.23G





Species yamashironis Yoshii, 1995


Genus Neanuratobeassignedtopropergenus Janssens F, 2017

Species aethiopica (Tarsia in Curia, 1939)

Species albipes (Nicolet, H, 1847:389), i.s.

Remark by Ellis, W.N. & Bellinger, P.F. 1973:28: Nicolet, 1847 described new species from Chile. These are also described by Nicolet in Gay, 1851[lapsus for Nicolet in Gay, 1849]. But in the first paper Nicolet cites as author of the species Nicol. in Gay. Evidently the publications appeared in a reversed order.

Species alpestris (Latzel, R, 1917:236)

Species atra (Nicolet, H, 1847:389), i.s.

Remark by Ellis, W.N. & Bellinger, P.F. 1973:28: Nicolet, 1847 described new species from Chile. These are also described by Nicolet in Gay, 1851[lapsus for Nicolet in Gay, 1849]. But in the first paper Nicolet cites as author of the species Nicol. in Gay. Evidently the publications appeared in a reversed order.

Species azrouensis (Thibaud & Massoud, 1980)

Species bakeri (Handschin, 1926), s.i.

             Syn.: Achorutes Bakeri(sic) Denis, J-R, 1933:231 ISS

Species casabianca (Fernando, 1959), i.s.

Species caucasica (Stach, 1951:35)

Species chilensis (Nicolet, H, 1847:389), i.s.

Remark by Ellis, W.N. & Bellinger, P.F. 1973:28: Nicolet, 1847 described new species from Chile. These are also described by Nicolet in Gay, 1851[lapsus for Nicolet in Gay, 1849]. But in the first paper Nicolet cites as author of the species Nicol. in Gay. Evidently the publications appeared in a reversed order.

Anoura chilensis from Chile
After Nicolet H in Gay C, 1854 Lam.1 Fig.3.
             Syn.: Anoura Chilensis(sic) Lubbock, J, 1873:200 ISS


Species coronifera (Axelson, WM, 1905:789)

Species eburnea (Gisin, 1963)

Species florida (Latzel, R, 1917:236)

Species folsomi (Stach, 1951:26), n.n.

Species growae (Christiansen, K et Bellinger, P, 1980:377)

Neanura growae from the USA
2023.03.31 © Wang, G.
Neanura growae from the USA
Eye with 2 widely separated ocelli
2023.03.31 © Wang, G.
Neanura growae ♂ from the USA
Abd.6 weakly bilobed
2024.01.27 © Wang, G.
Neanura growae ♂ from the USA
Abd.6 weakly bilobed
2024.01.27 © Wang, G.
Neanura growae ♂ from the USA
Abd.5 partly hidden under abd.6
2024.01.27 © Wang, G.
Neanura growae ♂ from the USA
Eye with 2 ocelli
2024.01.27 © Wang, G.
Neanura growae ♂ from the USA
Male genital orifice
2024.01.27 © Wang, G.
Neanura growae ♂ from the USA
First foot
2024.01.27 © Wang, G.

Species hirta (Joseph, 1881)

Species infernalis (Joseph, 1881)

Species japonica (Börner, C, 1909:107)

Species kentingensis (Lee, B-H et Kim, J-T, 1990:237)

Neanura kentlingensis(sic) from Taiwan
After Lee B-H & Park K-H, 1990 Fig.2A
             Syn.: Neanura kentlingensis(sic) Lee, B-H et Kim, J-T, 1990:238 ISS


Species laricis (Martynova, 1977)

Species loricata (Börner in Schille, 1912), n.n.

Species magna (MacGillivray, AD, 1893:128)

Neanura magna from the USA
2007.12.08 © Gross, J.
Neanura magna from the USA
2007.12.08 © Gross, J.
Neanura magna from the USA
2007.12.08 © Gross, J.
Neanura magna from the USA
2007.12.08 © Gross, J.
Neanura magna from the USA
2014.08.12 © Bertone, M.
Neanura magna from the USA
Th.2+3 dorsolateral tubercles distinctly projected
2014.08.12 © Bertone, M.
Neanura magna from the USA
Abd.4 with dorso-lateral tubercle behind dorso-external tubercle
2014.08.12 © Bertone, M.
Neanura magna from the USA
Body setae blunt and serrate
2014.08.12 © Bertone, M.
Neanura magna from the USA
Tubercles on abd.4 in triangular constellation
2016.11.28 © Dobbs, Br.
Neanura magna ♂ from the USA
Bilobed abd.6 hidden under abd.5
2016.11.28 © Dobbs, Br.
Neanura magna from the USA
Cephalic tubercles: (1)+2+4+6
2016.11.28 © Dobbs, Br.
Neanura magna from the USA
Abdominal dorsal tubercles: 6+6+6+6+4
2016.12.07 © Dobbs, Br.
Neanura magna ♀ from the USA
Rudiment of furca
2016.12.07 © Dobbs, Br.
Neanura magna juv. from the USA
2018.01.28 © Johnson, S.T.
Neanura magna from the USA
2018.04.22 © Johnson, S.T.
Neanura magna from the USA
2018.04.22 © Johnson, S.T.
Neanura magna from the USA
2022.02.06 © Hays, T.
Neanura magna from the USA
Abd.4 with 3+3 tubercles
Abd.5 with 2+2 tubercles
2022.02.06 © Hays, T.
Neanura magna from the USA
Cephalic tubercles: 1+2+4+6
2022.02.06 © Hays, T.
Neanura magna from the USA
Ant.4 with dense ventral file
2022.02.06 © Hays, T.
Neanura magna from the USA
Ocular tubercle with ocelli
2022.02.06 © Hays, T.
Neanura magna from the USA
Abd.6 hidden under abd.5
2022.02.06 © Hays, T.

Species mirabilis (Massoud, 1963), i.s.

Species multituberculata (Coleman, 1941), s.i.

Species pallida (Willem, V, 1902:10)

            Syn.: Biclavella patagonica Wahlgren, 1907


Species patagonica (Wahlgren, 1901)

            Syn.: Biclavella pallida Börner, 1906

Species pini (Becker, 1948)

Species pseudornata (Scott, 1961)

Species sexoculata (Carpenter, GH, 1916:29), i.s.

Species takoensis (Kinoshita, 1916)

Neanura takoensis from China
2006.03.14 © Botting, J.
Neanura takoensis from China
After www.yellowman.cn., 2007 ©
Neanura takoensis from China
2011.02.11 © Botting, J.
Neanura takoensis from China
2011.02.11 © Botting, J.

Species tumulosa (Li, 1991)

Species weigneri (Stach, 1922)

            Syn.: Achorutes Weigneri(sic) Stach, 1922 IOS
            Syn.: Achorutes Weigneri(sic) Denis, J-R, 1933:228 IOS
            Syn.: Achorutes Weigneri(sic) Denis, J-R, 1947:3 IOS


Genus Neanura MacGillivray, AD, 1893:314 key

  Syn.: ... Achorutes Templeton, 1835,nr ICZN
  Syn.: ... Achoreutes(sic) Koch, 1840 nr ISS
  Syn.: ... Achoreutes(sic) Templeton, 1843 nr ISS
  Syn.: Podurellen(sic) Achonetes(sic) Elditt, HL, 1854:39 ISS
  Syn.: Podurellen(sic) Achonetes(sic) Elditt, HL, 1854:40 ISS
  Syn.: Podurides(sic) Achonetes(sic) Elditt, HL, 1854:42 ISS
  Syn.: ... Acharutes(sic) Harvey, 1893 ISS
  Syn.: Poduridae Blax Koch, 1840 =?
  Syn.: ... Anoura(sic) Gervais, 1843 nec Gray, 1838 IOS
  Syn.: ... Anura Agassiz, 1846
  Syn.: Thysanoures(sic) Anoura(sic) Lucas, H in Desmarest, E, 1846:110 IOS
  Syn.: Lipurelas Anoura(sic) Nicolet, H in Gay, C, 1849:95 IOS
  Syn.: Podura Anoura(sic) Elditt, HL, 1854:44 IOS
  Syn.: ... Anoura(sic) Joseph ISS
  Syn.: Lipuridæ(sic) Anoura(sic) Lubbock, J, 1862:600 partim ISS
  Syn.: Thysanura Anoura(sic) MacGillivray, AD, 1893:128 partim ISS
  Syn.: Aphoruridæ(sic) Anoura(sic) MacGillivray, AD, 1893:314 ISS
  Syn.: Thysanura Anoura(sic) Cockerell, TDA, 1894:116 ISS
  Syn.: Thysanura Anoura(sic) Harvey, FL, 1896:422 ISS
  Syn.: Achorutidae Anoura(sic) Carpenter, GH, 1904:302 ISS
  Syn.: Collembola Anoura(sic) Bonet, F, 1930:113 ISS
  Syn.: ... Rathumoutes Templeton, 1844 nr
  Syn.: Podurides(sic) Hypogastrura Elditt, HL, 1854:42
  Syn.: ... Aphoromma MacGillivray
  Syn.: Thysanura Aphoromma Harvey, FL, 1896:423
  Syn.: Achorutidae Achorutini(sic) Börner, C, 1901:13 partim IOS
  Syn.: ... Biclavella Willem, 1902
  Syn.: ... Biloba Stach, 1949,nr Opinion 435, 1956
  Syn.: ... Imparitubercula Stach, 1951
  Syn.: Bilobidae Kodiakia Bödvarsson, H, 1960:43


  Syn.: Neanurini Neauura(sic) Yosii, R, 1965:13 ISS
Neanura
Cephalic tubercles
After Deharveng L, 1983 Fig.20B
Remark by ICZN. 2007:543: Placed on the Official List of Names in Zoology. Opinion 435.

Neanura sp. from Taiwan
2022.01.12 © Cheng, H.-J.





Species alba von Törne, 1957

            Syn.: Neanura parva Dunger, 1975 partim
Remark by Dányi, L. & Traser, G. 2008:16: The data of Neanura parva (Stach) reported by Dunger (1975) included Neanura alba as well.




Species deharvengi Smolis, A, Shayanmehr, M et Yoosefi-Lafooraki, E, 2018:5

Species judithae Smolis, A et Deharveng, L, 2017:39

Neanura judithae from Poland
After Smolis A & Deharveng L, 2017 Fig.1A

Species minuta Gisin, 1963

Neanura minuta from Turkey
After Özata MA, Sevgili H & Kaprus IJ, 2017 Fig.7

Species moldavica Busmachiu, G et Deharveng, L, 2008:?

Species muscorum (Templeton, R in Templeton, R & Westwood, JO, 1836:97) MacGillivray, AD, 1893:314, t.t., B

            Syn.: Achorutes Muscorum(sic) Templeton, R in Templeton, R et Westwood, JO, 1836:97,tt IOS
Achorutes Muscorum(sic) from Ireland
After Templeton, R, 1836 Pl.XII Fig.6


            Syn.: Blax ater Koch, 1840
            Syn.: Achorutes tuberculatus Nicolet, H, 1842:51
Achorutes tuberculatus from Switzerland
After Nicolet, H., 1842 Pl.5 Fig.1.
Anoure tuberculée from Europe
Anoura tuberculata
After Gervais P in Walckenaer CA, 1844 Pl.50 Fig.1

             Syn.: Achorutes Tuberculatus(sic) Bourlet, C, 1845:67 ISS
             Syn.: Achonetes(sic) taberculatus (sic) Elditt, HL, 1854:40 ISS
            Syn.: Anoura(sic) crassicornis Müller J, 1859
            Syn.: Anoura(sic) nigra Wankel, 1860
            Syn.: Anura gibbosa Packard, AS, 1873:27
Anoura(sic) gibbosa from the USA
Somewhat too broad from pressure
After Emerton JH in Harvey FL, 1896 Fig.1
Anoura(sic) gibbosa from the USA
Ventral aspect
After Emerton JH in Harvey FL, 1896 Fig.2
Anoura(sic) gibbosa from the USA
After Emerton JH in Harvey FL, 1896 Fig.3

            Syn.: Anoura(sic) 6-tuberculata(sic) Harvey, FL, 1896:423 IOS
            Syn.: Neanura muscorum var. purpurascens Axelson, WM, 1905:790
            Syn.: Neanura muscorum ssp. alpestris Latzel, 1917
            Syn.: Neanura betulae Selga, 1959
            Syn.: Neanura persimilis Smolis, A et Deharveng, L, 2017:46 =?



Achorutes muscorum from France
After Lucas M, 1840 Pl.1 Fig.4
Anoura muscorum from the UK
After Lubbock, J., 1873 Pl.48 Fig.1.
Neanura muscorum from New Zealand
After Salmon, J.T., 1941 Pl.46 Fig.104.
             Syn.: Achorutes Muscorum(sic) Bourlet, C, 1845:160 ISS
Neanura muscorum
Cephalic clypeal and antennofrontal tubercles
After Deharveng L, 1983 Fig.14E
Neanura muscorum
Dorsal chaetotaxy
Modified after Deharveng L, 1983 Fig.21A
Neanura muscorum
Dorsal tubercles
After Deharveng L, 1983 Fig.23B
Remark by Deharveng, L. & Bedos, A. 2000:211: The Vietnamese record possibly corresponds to Vietnura caerulea, new species, which is the most common Neanurini of Vietnam. Neanura muscorum, widespread in temperate zones, has never been met in the numerous samples we collected in the region, nor in any country of Southeast Asia.

Neanura muscorum from the UK
After Hopkin, S.P., 2002 ©
Neanura muscorum from Sweden
2005 © Hall, K.
Neanura muscorum
After Boldsystems.org, 2006 ©
Neanura muscorum from Sweden
2006 © Hall, K.
Neanura muscorum from Norway
2006.07.29 © Fauskanger, I.
Neanura muscorum from the UK
2007 © Barton, T.
Neanura muscorum from the USA
2007 © McClarin, J.
Neanura muscorum from France
2007.03.15 © Lebeaux, P.
Neanura muscorum from France, juvenile
2007.03.15 © Lebeaux, P.
Neanura muscorum from France
2007.03.15 © Lebeaux, P.
Neanura muscorum from France
2007.03.15 © Lebeaux, P.
Neanura muscorum from the UK
2007.04.01 © Cornwall, N.J.
Neanura muscorum from the UK
2008.01.06 © Campbell, A.
Neanura muscorum from the Netherlands
2008.01.15 © van Erkelens, J.A.
Neanura muscorum from France
2008.01.19 © Hugues, C.
Neanura muscorum from the UK
2008.04.18 © Farrell, S.
Neanura muscorum from the UK
2008.04.18 © Farrell, S.
Neanura muscorum from Algeria
After Hamra-Kroua, S., 2009.04.02 ©
Det. Deharveng, L.
Neanura muscorum with egg deposition
After Thibaud J-M & D'Haese C, 2010 Fig.50
2009 © D'Haese, C.
Neanura muscorum from Belgium
2011.09.09 © Henderickx, H.
Neanura muscorum juv. from the USA
2012.02.25 © Murray, T.
Neanura muscorum from the UK
With phoretic Nematoda
2012.02.29 © Nurcombe, E.
Neanura muscorum from Denmark
2012.03.21 © Bildstrup, O.
Neanura muscorum from the USA
With parasitic mite
2012.05.04 © Murray, T.
Neanura muscorum from the UK
Ocular tubercles not united with frontal tubercle
2012.06.27 © Murray, A.
Neanura muscorum from the UK
Ocular tubercle with 2+1 ocelli
2012.09.07 © Murray, A.
Neanura muscorum from the UK
Flower-like cuticular ornamentation on tubercles
2012.12.11 © Murray, A.
Neanura muscorum from the UK
Sharply pointed mouth cone
2012.12.11 © Murray, A.
Neanura muscorum from the UK
Bilobed 6th abdominal segment
2013.01.06 © Nurcombe, E.
Neanura muscorum from France
Ocular tubercle with 2 anterior ocelli and 1 posterior ocellus
2013.03.24 © Sandra.
Neanura muscorum from France
Cephalic tuberculae and pointed mouthcone
2013.03.24 © Sandra.
Neanura muscorum from the UK
Tuberculate hydrophobic cuticula
2013.12.04 © Murray, A.
Neanura muscorum from the UK
2+2 anterior ocelli
Cephalic tubercles not fused
2014.08.22 © Murray, A.
Neanura muscorum from the USA
2014.11.09 © Silsbee, D.
Neanura muscorum from the UK
Bilobed 6th abdominal segment
2015.01.28 © Murray, A.
Neanura muscorum from the UK
Moulting with uplifted abdomen
2016.01.08 © Jeffery, J.
Neanura muscorum from Canada
2016.04.29 © de Repentigny, L.-G.
Neanura muscorum from Belgium
Ocular tubercle with 2 anterior + 1 posterior ocellus
2017.12.30 © Huskens, M.-L.
Neanura muscorum from Belgium
2017.12.30 © Huskens, M.-L.
Neanura muscorum from Belgium
2018.01.10 © Huskens, M.-L.
Neanura muscorum from Belgium
Mouthcone pointed; 2 anterior ocelli
2018.01.10 © Huskens, M.-L.
Neanura muscorum from Belgium
Mouthcone pointed
Ventral aspect
2018.04.20 © Huskens, M.-L.
Neanura muscorum juv. from France
2018.10.15 © Garcelon, P.
Neanura muscorum from Holland
Ant.3 and 4 dorsally fused
2019.02.03 © Kamsteeg, G.
Neanura muscorum from Holland
Ant.3 and 4 ventrally not fused
2019.02.03 © Kamsteeg, G.
Neanura muscorum from Holland
Ocular tubercle with 2 anterior ocelli and 1 posterior ocellus
2019.02.03 © Kamsteeg, G.
Neanura muscorum ♀ from Holland
Female genital orifice
2019.02.03 © Kamsteeg, G.
Neanura muscorum ♀ from Holland
Furca absent
2019.02.03 © Kamsteeg, G.
Neanura muscorum from Belgium
Grooming with 6 legs while balancing on collophore
2019.05.01 © Huskens, M.-L.
Neanura muscorum from Belgium
Interocular macrosetae
2019.09.14 © Huskens, M.-L.
Neanura muscorum from Belgium
Pointed mouthcone (*)
2019.10.31 © Huskens, M.-L.
Neanura muscorum juv. from Belgium
2019.11.04 © Huskens, M.-L.
Neanura muscorum juv. from Belgium
Eye with 2 anterior ocelli + 1 posterior ocellus
2019.11.04 © Huskens, M.-L.
Neanura muscorum juv. from Belgium
2020.08.19 © Huskens, M.-L.
Neanura muscorum ♀ from Belgium
Furca vestigial but teratologically large
2020.11.10 © Huskens, M.-L.
Neanura muscorum ♀ from Belgium
Anus with 2 hidden ventral anal valves
and bilobed dorsal anal valve
2020.11.10 © Huskens, M.-L.
Neanura muscorum ♀ from Belgium
Abd.5 with elongate transversal female genital organ
2020.11.10 © Huskens, M.-L.
Neanura muscorum from Czechia
Forward directed pointed mouthcone
2020.11.18 © Kubát, L.
Neanura muscorum from France
Cephalic tubercles
2021.03.06 © Vol, F.
Neanura muscorum from France
Abdominal tubercles
2021.03.06 © Vol, F.
Neanura muscorum from Belgium
Forward directed pointed mouthcone
2021.03.09 © Huskens, M.-L.
Neanura muscorum ♂ from Belgium
Vestigial furca
2023.01.05 © Huskens, M.-L.
Neanura muscorum ♂ from Belgium
Abd.5 with roundish male genital organ
2023.01.05 © Huskens, M.-L.
Neanura muscorum from Belgium
Eye with 2 anterior + 1 posterior ocelli
Ocelli in obtuse angle pointing down
2023.01.06 © Huskens, M.-L.
Neanura muscorum from Belgium
Walking with "rotating" legs
2023.05.18 © Huskens, M.-L.





Species pallida Deharveng, 1979

Species parva (Stach, J, 1951:48) Gisin, 1960:99

Species persica Arbea JI, Yahyapour E et Shayanmehr M in Yahyapour, E, Shayanmehr, M, Vafaei-Shoushtari, R et Arbea, JI, 2022:266

Neanura persica from Iran
After Yahyapour E & al, 2022 Fig.37-38

Species persimilis Mills, HB, 1934:30

Neanura persimilis from the USA
Tubercles only on head and posterior body segments
2015.01.01 © Fleming, K.
Neanura persimilis from the USA
2015.09.15 © Wechsler, D.
Neanura persimilis from the USA
2022.01.24 © Dankowicz, Z.
Neanura persimilis from the USA
Abdominal tubercles
Abd.4 with 3 small dorso-external tubercles
2022.01.24 © Dankowicz, Z.
Neanura persimilis from the USA
Cephalic tubercles
2022.01.24 © Dankowicz, Z.
Neanura persimilis from the USA
Abd.5 dorso-external tubercle with 2 macrosetae
Abd.6 tubercle with 3 macrosetae
2022.01.24 © Dankowicz, Z.
Neanura persimilis from the USA
Eye ocelli 2+1 in almost straight line
2022.01.24 © Dankowicz, Z.
Neanura persimilis from the USA (in culture)
Hatchling
Feeding from plasmodium of slime mould
2023.09.01 © Gruver, Be.
Neanura persimilis from the USA (in culture)
Freshly deposited egg
2023.09.01 © Gruver, Be.
Neanura persimilis from the USA
+ 1 day days old juveniles
2023.10.22 © Gruver, Be.
Neanura persimilis juv. from the USA
7 days old
2023.10.29 © Gruver, Be.
Neanura persimilis from the USA
3 ocelli in obtuse angle pointing up
2023.12.04 © Gruver, Be.
Neanura persimilis from the USA
3 ocelli in obtuse angle pointing up
2023.12.04 © Gruver, Be.
Neanura persimilis from the USA
3 ocelli in obtuse angle pointing up
2023.12.04 © Gruver, Be.
Neanura persimilis from the USA
Light colour form
2023.12.07 © Gruver, Be.
Neanura persimilis from the USA
Abdominal dorsal tubercles
2023.12.07 © Gruver, Be.
Neanura persimilis from the USA
Fluorescency under blacklight of 365 nm UV
2023.12.12 © Gruver, Be.
Neanura persimilis from the USA
Reflex bleeding
Juvenile reacts to released alarm pheromone
2023.12.20 © Gruver, Be.
Neanura persimilis from the USA
3 ocelli in obtuse angle pointing up
Ocular tubercle with distinctly large macroseta
2024.01.13 © Coogler, J.
Neanura persimilis from the USA
Posterior abdominal tubercles
2024.01.13 © Coogler, J.
Neanura persimilis from the USA
Ready to moult
Antennae redrawn in old cuticula
2024.04.11 © Webb, D.

Species pseudoparva Rusek, 1963


Genus Thaumanura Börner, 1932

  Syn.: Bilobidae Thamanura(sic) Stach, J, 1951:Pl.1 Fig.4 ISS

Species carolii (Stach, 1920) Börner, 1932, t.t.

            Syn.: Achorutes Carolii(sic) Stach, 1920,tt IOS
Thamanura(sic) carolii from Poland
After Stach J, 1951 Pl.1 Fig.4

             Syn.: Thaumanura caroli(sic) Dallai, R, 1969:377 ISS
Thaumanura carolii from Romania
After Boldsystems.org, 2006 ©
Thaumanura carolii from Czechia
2007.07.05 © Krásenský, P.
Thaumanura carolii from Slovenia
2013.06.09 © Miran, C.
Thaumanura carolii from Slovenia
Posterior abdominal segments
2013.06.09 © Miran, C.
Thaumanura carolii from Slovenia
Ventral aspect
2013.06.09 © Miran, C.

             Syn.: Thaumanura caroli(sic) Fiera, C, 2014:229 ISS
             Syn.: Thaumanura caroli(sic) Fiera, C, 2014:230 ISS
             Syn.: Thaumanura caroli(sic) Fiera, C, 2014:231 ISS
             Syn.: Thaumanura caroli(sic) Fiera, C, 2014:232 ISS
             Syn.: Thaumanura caroli(sic) Fiera, C, 2014:233 ISS




Species dallaii Cassagnau, P et Peja, N, 1979:221

Species echinata (Kos, 1940)

            Syn.: Achorutes bornensis Schött, 1925 =?
            Syn.: Achorutes japonicus Börner, 1909 =?
            Syn.: Achorutes pterothrix Börner, 1909 =?


Species oniscoides (Latzel, R, 1917:237) Stach, J, 1951:65, s.i.

             Syn.: Achorutes Carolii(sic) Stach, 1919 =? ISS
            Syn.: Achorutes Carolii(sic) Stach, 1929
             Syn.: Achorutes Carolii(sic) Denis, J-R, 1933:230 ISS


             Syn.: Thaumanura oniscoïdes(sic) Massoud, Z, 1967:329 ISS
Thaumanura oniscoides from Hungary
2012.05.10 © Lásló, D.



Species ruffoi Dallai, R, 1969:377

Thaumanura ruffoi
After Deharveng L, 1983 Fig.1C
Thaumanura ruffoi from Italy
2011.04.09 © Piredda, C.


Genus Kalanura Smolis, A, 2007:?

Species babenkoi Smolis, A, 2007:?

Species compacta Smolis, A, 2007:?

Species spectabilis Smolis, A, 2007:?

Species stebaevae Smolis, A, 2007:?


Genus Vietnura Deharveng, L et Bedos, A, 2000:211

Species caerulea Deharveng, L et Bedos, A, 2000:212, t.t.

Vietnura caerulea from Vietnam
After Deharveng, P. & Bedos, A., 2000 Fig.1


Genus Paravietnura Smolis, A et Kuznetsova, N, 2018:44

Species insolita Smolis, A et Kuznetsova, N, 2018:49

Paravietnura insolita from the Caucasus
After Smolis A & Kuznzetsova N, 2018 Fig.9+12

Species notabilis Smolis, A et Kuznetsova, N, 2018:45, t.t.

Paravietnura notabilis from the Caucasus
After Smolis A & Kuznzetsova N, 2018 Fig.1

Species rostrata Smolis, A et Skarzynski, D, 2020:129


Genus Intricatonura Smolis, A et Bernard, EC, 2017:727, t.t.

Species fjellbergi Smolis, A et Bernard, EC, 2017:728

Intricatonura fjellbergi from the USA
After Smolis A & Bernard EC, 2017 Fig.1+11


Genus Cansilianura Dallai, R et Fanciulli, PP, 1983:235

Species malatestai Dallai, R et Fanciulli, PP, 1983:237, t.t.

Cansilianura malatestai from Italy
After Boldsystems.org, 2006 ©


Genus Caucasanura Kuznetsova & Potapov, 1988

Species besucheti Deharveng, L, 1989:817

Species stebaevae Kuznetsova & Potapov, 1988

             Syn.: Caucasanura stebayevae(sic) Deharveng, 1989 ISS
             Syn.: Caucasanura stebayevae(sic) Smolis, A et Kuznetsova, N, 2018:10 ISS


Genus Ghirkanura Kuznetsova & Potapov, 1988

  Syn.: Neanuridae (Neanurinae) Girkanura(sic) Hopkin, SP, 1997:205 ISS

Species chernovae Kuznetsova & Potapov, 1988


Genus Imparitubercula Stach, 1951

Species villosa (Kos, 1940) Stach, J, 1951:11

             Syn.: Imparitubercula villosia(sic) Cassagnau, P, 1983:21,si ISS



Genus Tetraloba Lee, B-H, 1983:37

Species seolagensis Lee, B-H, 1983:38, t.t.

Tetraloba seolagensis from Korea
After Lee B-H, 1983 Fig.1ABCD


Genus Xylanura Smolis, 2011

Species oregonensis Smolis, 2011


Genus Persanura Mehrafroz Mayvan, M, Shayanmehr, M, Smolis, A et Skarzynski, D, 2015:553

Species hyrcanica Mehrafroz Mayvan, M, Shayanmehr, M, Smolis, A et Skarzynski, D, 2015:553, t.t.

Species lencarana Smolis, A et Kuznetsova, N, 2018:6

Persanura lencarana from Azerbaijan
After Smolis A & Kuznetsova N, 2018 Fig.14


Genus Christobella Fjellberg, 1985

  Syn.: ... Cristobella Skidmore, 1995 ISS

Species ornata (Folsom, JW, 1902:89) Fjellberg, 1985

Neanura ornata from Alaska
After Folsom JW, 1902 Pl.IV Fig.2


Genus Nahuanura Palacios-Vargas, JG et Najt, J, 1986:18 key

Species ce Palacios-Vargas, JG et Najt, J, 1986:19, t.t.

Species ome Palacios-Vargas, JG et Simón-Benito JC, 2007:170

Species suenoi (Yosii, R, 1955:382) Palacios-Vargas, JG & Simón-Benito JC, 2007:175

            Syn.: Paranura s-uénoi(sic) Yosii, R, 1955:382 IOS
            Syn.: Paranura s-uénoi(sic) Yosii, R, 1955:379 IOS
            Syn.: Paranura s-uénoi(sic) Yosii, R, 1955:383 IOS
Paranura s-uénoi from Tokara Island
After Yosii R, 1955 Abb.2A

             Syn.: Paranura s-uenoi(sic) Yosii, R, 1965:10 ISS
             Syn.: Paranura s-uenoi(sic) Massoud, Z, 1967:238 ISS
             Syn.: Paranura s-uenoi(sic) Yosii, R, 1977:146 ISS
             Syn.: Paranura s-uenoi(sic) Christiansen & Bellinger, 1980 ISS
             Syn.: Paranura s-uenoi(sic) Cassagnau, 1982 ISS
             Syn.: Paranura s-uenoi(sic) Deharveng, L, 1989:104 ISS
Remark by Nicolson, D. 2007.mm.dd:in litt.: Based on art.NN the hyphen in the original spelling should be removed.


             Syn.: Nahuanura? s-uenoi(sic) Palacios-Vargas, JG et Simón-Benito JC, 2007:175 ISS
             Syn.: Paranura? s-uenoi(sic) Simón-Benito, JC et Palacios-Vargas, JG, 2008:683 ISS




Species trioculata Palacios-Vargas, JG et Simón-Benito JC, 2007:172

Species yei Palacios-Vargas, JG et Simón-Benito JC, 2007:172


Genus Paranura Axelson, 1902 key

  Syn.: ... Börneria(sic) Axelson, 1902 nec Willem, 1902


Paranura sp. from the USA
Body tubercles reduced
2024.02.09 © Brockman, T.
Paranura sp. from the USA
Abd.6 not bilobed
2024.02.09 © Brockman, T.
Paranura sp. from the USA
Eye with 2 spaced ocelli
2024.02.09 © Brockman, T.

Species sp.1 Janssens F sensu Christiansen & Reddell, 1986

            Syn.: Paranura caeca Reddell & Mitchell, 1971
            Syn.: Paranura caeca Christiansen & Reddell, 1986
            Syn.: Paranura caeca Palacios-Vargas, JG et Deharveng, L, 1987:105
            Syn.: Paranura caeca Mari Mutt, JA et Bellinger, PF, 1990:48
            Syn.: Sensillanura? caeca Palacios-Vargas, 1997
            Syn.: Paranura caeca Palacios-Vargas, 1997
            Syn.: Paranura sp.n. near anops Christiansen & Reddell, 1986


Species alpicola Kasai, H et Sawahata, T, 2023:?

Species anops Christiansen, K et Bellinger, P, 1980:346

Species bisetosa Deharveng, L, 1989:113

             Syn.: Paranura Bisetosa(sic) Simón-Benito, JC et Palacios-Vargas, JG, 2008:687 ISS

Species chiangdaoensis Deharveng, L, 1989:105

             Syn.: Paranura Chiangdaoensis(sic) Simón-Benito, JC et Palacios-Vargas, JG, 2008:687 ISS

Species coenobita Cassagnau, 1991

             Syn.: Paranura cenobita(sic) Simón-Benito, JC et Palacios-Vargas, JG, 2008:685 ISS
             Syn.: Paranura coenotiba(sic) Simón-Benito, JC et Palacios-Vargas, JG, 2008:685 ISS

Species colombiana Palacios-Vargas, JG et Peñaranda-Parada, MR, 2005:308

Species colorata Mills, HB, 1934:28


            Syn.: Paranura sexpunctata Neartic records


             Syn.: Paranura 6-punctata Mills, HB, 1934:128 ISS
             Syn.: Paranura Colorata(sic) Simón-Benito, JC et Palacios-Vargas, JG, 2008:687 ISS
Paranura colorata from the USA
2016.12.12 © Dobbs, Br.
Paranura colorata from the USA
Pointed mouthcone
2016.12.29 © Dobbs, Br.
Paranura colorata subadult from the USA
2023.01.01 © Wang, G.
Paranura colorata juv. from the USA
2023.01.01 © Wang, G.
Paranura colorata juv. from the USA
2023.01.01 © Wang, G.
?Paranura colorata hatchling? from the USA
2023.01.01 © Wang, G.



Species convallis Kasai, H et Sawahata, T, 2023:?

Species dalgeri Deharveng, L, 1989:111

Species garoensis Cassagnau, 1991

Species globulifer Deharveng, L, 1989:118

Species godavarica Cassagnau, 1991

Species ieti (Yosii, 1966) Deharveng, L, 1989:104

            Syn.: Paranura najtae Deharveng, L et Weiner, WM, 1984:11


Propeanura ieti from Korea
After Lee B-H, 1981 Fig.1A

             Syn.: Paranura Ieti(sic) Simón-Benito, JC et Palacios-Vargas, JG, 2008:687 ISS


Species impedita Palacios-Vargas, JG et Deharveng, L, 1987:111

Species johorensis (Yosii, 1976) Deharveng, L, 1989:104

Species jorgei Palacios-Vargas, JG et Deharveng, L, 1987:106

             Syn.: Paranura Jorgei(sic) Simón-Benito, JC et Palacios-Vargas, JG, 2008:687 ISS

Species koryoi Deharveng, L et Weiner, WM, 1984:7

             Syn.: Paranura Koryoi(sic) Simón-Benito, JC et Palacios-Vargas, JG, 2008:687 ISS

Species leclerci Deharveng, L, 1989:115

             Syn.: Paranura Leclerci(sic) Simón-Benito, JC et Palacios-Vargas, JG, 2008:687 ISS

Species longisensillata Palacios-Vargas, JG et Deharveng, L, 1987:111

             Syn.: Paranura Longisensillata(sic) Simón-Benito, JC et Palacios-Vargas, JG, 2008:687 ISS

Species magdalenae Simón-Benito, JC et Palacios-Vargas, JG, 2008:685

             Syn.: Paranura Magdalenae(sic) Simón-Benito, JC et Palacios-Vargas, JG, 2008:687 ISS

Species meo Deharveng, L, 1989:118

Species mjohjangensis Deharveng, L et Weiner, WM, 1984:9

             Syn.: Paranura Mjohjangensis(sic) Simón-Benito, JC et Palacios-Vargas, JG, 2008:687 ISS

Species modesta Deharveng, L, 1989:105

Species nakamurai Kasai, H et Sawahata, T, 2023:?

Species nalo Christiansen, K et Bellinger, P, 1992:68

Species nudifera Yoshii, R et Suhardjno, YR, 1992:8

Species quadrilobata Hammer, 1953

Species quadripunctata Yosii, 1966

Species reticulata Smolis & Deharveng, 2015

Species rooensis Simón-Benito, JC et Palacios-Vargas, JG, 2008:683

             Syn.: Paranura Rooensis(sic) Simón-Benito, JC et Palacios-Vargas, JG, 2008:687 ISS

Species rosea Lee, B-H et Kim, J-T, 1984:20

Species sarukhani Palacios-Vargas, JG et Deharveng, L, 1987:106

             Syn.: Paranura Sarukhani(sic) Simón-Benito, JC et Palacios-Vargas, JG, 2008:687 ISS

Species sexpunctata Axelson, 1902, t.t.

            Syn.: Börneria(sic) clavisetis Axelson, 1902
             Syn.: Paranura sexpunctata var. claviseta(sic) Womersley, H, 1935:208 ISS
            Syn.: Paranura clavisetis var. quadrimaculata Linnaniemi, 1907
            Syn.: Paranura sexpunctata ssp. colorata Mills, 1934


Paranura sexpunctata
After Deharveng L, 1983 Fig.1A
Paranura sexpunctata
After Deharveng L, 1983 Fig.12A

             Syn.: Paranura Sexpunctata(sic) Simón-Benito, JC et Palacios-Vargas, JG, 2008:687 ISS


Species sitchensis Fjellberg, 1985

Species squamosa Cassagnau, 1991

Species tamul Cassagnau, P, 1988:25

Species tapatia Palacios-Vargas, JG et Peñaranda-Parada, MR, 2005:311

Species tibiotarsalis Deharveng, L, 1989:110

Species timorensis Yoshii, R et Suhardjono, YR, 1992:76

Paranura timorensis from Indonesia
After Yoshii R & Suhardjono YR, 1992 Fig.1F

Species tridentata Lee, B-H et Kim, J-T, 1984:19

Species tsushimaensis Kasai, H et Sawahata, T, 2023:?


Genus Oregonanura Smolis, 2008

Species cascadensis Smolis, 2008 tt



Genus Sensillanura Deharveng, 1981 key  key

  Syn.: Pseudachorutini Paranura Folsom, JW, 1916:506
  Syn.: ... Achorutes Handschin, 1928 partim
  Syn.: ... Neanura da Gama, 1963 partim
  Syn.: Neanurinae Neanura(lapsus) Christiansen, K et Bellinger, P, 1998:425
  Syn.: ... Lathriopyga Massoud, 1967 partim


Species sp.1 Palacios-Vargas, JG, 1990:19

Species austriaca (da Gama, 1963 tt) Cassagnau, P, 1983:11, t.t.

Sensillanura austriaca from Algeria
After Zoughailech A, 2017 Fig.50

Species barberi (Handschin, 1928 tt) Cassagnau & Palacios-Vargas, 1983

            Syn.: Achorutes Barberi(sic) Handschin, 1928 tt IOS
            Syn.: Neanura quadrioculata Guthrie, JE, 1903:101 nec Börner, 1901
            Syn.: Biloba pseudoquadrioculata Stach, J, 1951:31



            Syn.: Achorutes Barberi(sic) Denis, J-R, 1933:230 IOS
Sensillanura barberi from the USA
2008.11.10 © Justis, S.
Sensillanura barberi from the USA
2008.11.10 © Justis, S.
Sensillanura barberi from the USA
2009.03.15 © Justis, S.
Sensillanura barberi from the USA
2009.03.15 © Justis, S.
Sensillanura barberi from the USA
2009.03.15 © Justis, S.
Sensillanura barberi from the USA
2012.12.18 © Montgomery, G.
Sensillanura barberi from the USA
After Galván AA & Palacios-Vargas JG, 2017 Fig.1+2
Sensillanura ?barberi? from the USA
Body setae acuminate
2018.01.15 © Abbott, J.C.
Sensillanura ?barberi? from the USA
2018.01.15 © Abbott, J.C.
Sensillanura ?barberi? from the USA
2019.02.07 © Bairstow, A.
Sensillanura barberi from the USA
2020.03.18 © Alloway, L.
Sensillanura barberi from the USA
2020.03.18 © Alloway, L.
Sensillanura barberi from the USA
Abd.6 hidden underneath abd.5
Abd.5 dorso-internal tubercles not fused
2020.03.18 © Alloway, L.
Sensillanura barberi juv. from the USA
Posterior ocellus not visible
2020.03.18 © Alloway, L.
Sensillanura barberi from the USA
2023.03.31 © Shahan, T.
Sensillanura barberi from the USA
2023.03.31 © Shahan, T.
Sensillanura barberi from the USA
Abd.6 hidden under abd.5
2023.03.31 © Shahan, T.
Sensillanura barberi from the USA
Pointed mouthcone
2023.03.31 © Shahan, T.
Sensillanura barberi from the USA
2024.01.05 © Kadupul.
Sensillanura barberi from the USA
Abd.6 hidden under abd.5
2024.01.05 © Kadupul.
Sensillanura barberi from the USA
2024.04.10 © Webb, D.
Sensillanura barberi from the USA
2024.04.18 © Hendrix, S.
Sensillanura barberi from the USA
2024.04.18 © Hendrix, S.
Sensillanura barberi from the USA
2024.04.18 © Hendrix, S.
Sensillanura barberi juv. from the USA
2024.04.21 © Coogler, J.



Species bullsa (Wray, 1953) Cassagnau, 1982

            Syn.: Sensillanura barberi var.? bullsa Christiansen & Bellinger, [1992:376]
            Syn.: Neanura(lapsus) bullsa Christiansen, K et Bellinger, P, 1998:425
             Syn.: Sensillanura bulsa(sic) Palacios-Vargas, JG et Catalán, E, 2010:162 ISS
Sensillanura bullsa from the USA
Eyes absent
After Galván AA & Palacios-Vargas JG, 2017 Fig.9+10



Species caeca (Folsom, JW, 1916:507) Deharveng, 1981

            Syn.: Paranura caeca Folsom, JW, 1916:507
             Syn.: Paranura coeca(sic) Yosii, R, 1955:382 ISS
             Syn.: Paranura coeca(sic) Yosii, R, 1965:11 ISS
             Syn.: Sensillanura coeca(sic) Cassagnau & Palacios-Vargas, 1983 ISS
             Syn.: Sensillanura coeca(sic) Cassagnau, 1986 ISS
Sensillanura caeca from Canada
Eyes absent
Modified after Christiansen K & Bellinger P, 1998 Fig.265A
Sensillanura caeca from the USA
2017.11.25 © Dobbs, Br.
Sensillanura caeca from the USA
2017.11.25 © Dobbs, Br.
Sensillanura caeca from the USA
Pointed mouthcone
2017.11.25 © Dobbs, Br.
Sensillanura caeca from the USA
2020.04.10 © Emile.
Sensillanura caeca from the USA
2023.03.04 © Wang, G.
Sensillanura caeca from the USA
Mouthcone pointed
2023.03.04 © Wang, G.
Sensillanura caeca from the USA
Ventral anal valves hidden
Dorsal anal valve bilobed
2023.03.04 © Wang, G.
Sensillanura caeca from the USA
Unguiculus absent
2023.03.04 © Wang, G.
?Sensillanura caeca juv.? from the USA
2023.12.23 © Wang, G.
Sensillanura caeca from the USA
2024.02.12 © Wang, G.
Sensillanura caeca from the USA
2024.02.12 © Wang, G.
Sensillanura caeca from the USA
2024.02.12 © Wang, G.
Sensillanura caeca from the USA
'Playing dead'
2024.02.12 © Wang, G.
Sensillanura caeca from the USA
Maxilla head needle-like
2024.02.12 © Wang, G.
Sensillanura caeca from the USA
Mandibula apically tridentate
2024.02.12 © Wang, G.
Sensillanura caeca from the USA
Abd.6 dorsal anal valve bilobed
Terminal tubercles with 4 setae
2024.02.12 © Wang, G.



Species caeca2 Galván, AA et Palacios-Vargas, JG, 2017:232

            Syn.: Sensillanura caeca2 Galván AA et Palacios-Vargas JG, 2017:232 nec (Folsom, 1916:507)
Sensillanura caeca from the USA
Eyes present with 2+2 ocelli
After Galván AA & Palacios-Vargas JG, 2017 Fig.11+12
Sensillanura ?caeca2? juv. from the USA
2+2 ocelli
2024.04.15 © Webb, D.
Sensillanura ?caeca2? juv. from the USA
Abd.6 hidden under abd.5
2024.04.15 © Webb, D.



Species illina (Christiansen, K & Bellinger, P, 1980:378) Cassagnau & Palacios-Vargas, 1983

Species millsi (Christiansen, K & Bellinger, P, 1980:349) Cassagnau & Palacios-Vargas, 1983

Species santizaragozai Palacios-Vargas, JG et Catalán, E, 2010:161


Genus Honduranura Palacios-Vargas, JG, 2017:3

Species centraliamericana Palacios-Vargas, JG, 2017:4, t.t.

Honduranura centraliamericana from Honduras
After Palacios-Vargas JG, 2017 Fig.1


Genus Tabasconura Palacios-Vargas, JG et Catalán, E, 2015:131

Species tapijulapana Palacios-Vargas, JG et Catalán, E, 2015:137, t.t.

            Syn.: Tabasconura tapajulapana(sic) Palacios-Vargas, JG et Catalán, E, 2015:137 IOS
             Syn.: Tabasconura tapiculapana(sic) Palacios-Vargas, JG et Catalán, E, 2015:137 ISS





Genus Americanura Cassagnau, P, 1983:11 key

  Syn.: ... Mexicanura Palacios-Vargas, 1983 nn
  Syn.: Sensillanurini Palmanura Palacios-Vargas, JG, García-Barros, E et Simón-Benito, JC, 2009:1 =?


Remark by Ellis, W.N. & Bellinger, P.F. 1984:2: Both the generic name and that of the type-species are attributed to a paper in print by Cassagnau & Palacios-Vargas (1983). As the published information contains a nomenclaturally valid diagnosis of the new genus (and by implication the type-species as well), the publication of Cassagnau (1983) takes precedence.

  Syn.: Neanurinae Americaneura(sic) Peck, SB, 2001:106 ISS
  Syn.: Collembola Americaneura(sic) Peck, SB et Jacquemart, J, 2011:4 ISS
  Syn.: Sensillanurini Americaura(sic) Palacios-Vargas, JG, Mejía-Recamier, BE et Vázquez, MM, 2017:312 ISS
  Syn.: Sensillanurini Americaura(sic) Palacios-Vargas, JG, Mejía-Recamier, BE et Vázquez, MM, 2017:314 ISS
Americanura sp. from Mexico
Predated by ant
After Murray A in Palacios-Vargas JG & Cruz-Leal JI, 2020 Fig.7



Species banksi (Denis, J-R, 1933:226) Cassagnau & Palacios-Vargas, 1983

            Syn.: Achorutes Banksi(sic) Denis, J-R, 1933:226 IOS
Achorutes Banksi(sic) from Costa Rica
After Denis J-R, 1933 Fig.10

            Syn.: Achorutes Banksi(sic) Denis, J-R, 1933:227 IOS
            Syn.: Achorutes Banksi(sic) Denis, J-R, 1933:230 IOS
            Syn.: Achorutes Banksi(sic) Denis, J-R, 1933:315 IOS
            Syn.: Achorutes Banksi(sic) Denis, J-R, 1933:318 IOS


Species bara (Christiansen, K & Bellinger, P, 1980:373) Palacios-Vargas, JG & Najt, J, 1986:24

Americanura bara? from the USA
2017.03.26 © Lee, C.

Species basseti Palacios-Vargas, JG, Simón Benito, JC et Paniagua Nucamendi, J, 2009:439

Americanura basseti from Panama
After Palacios-Vargas & al, 2009 Fig.14-15

Species belicensis Palacios-Vargas, JG, Mejía-Recamier, BE et Vázquez, MM, 2017:310

Americanura belicensis from Belize
Modified after Palacios-Vargas JG & al, 2017 Fig.1-2

Species castagnorum Palacios-Vargas, JG, Simón Benito, JC et Paniagua Nucamendi, J, 2009:439

Americanura castagnorum from Panama
After Palacios-Vargas & al, 2009 Fig.12-13

Species castillorum Palacios-Vargas & Najt, 1986:15

Species denisi Palacios-Vargas, JG, Mejía-Recamier, BE et Vázquez, MM, 2017:312

Americanura denisi from Belize
Modified after Palacios-Vargas JG & al, 2017 Fig.4-5

Species guatemalteca Cassagnau & Palacios-Vargas, 1983:2

Species imitator Cassagnau & Palacios-Vargas, 1983:2

Species interrogator Cassagnau, P et Palacios-Vargas, JG, 1983:7

Species izabalana Palacios-Vargas, JG et Ríos, G, 1996:440

Species iztac Palacios-Vargas, JG, Simón Benito, JC et Paniagua Nucamendi, J, 2009:437

Americanura iztac from Mexico
After Palacios-Vargas & al, 2009 Fig.10-11

Species janssensi Palacios-Vargas, JG, Simón Benito, JC et Paniagua Nucamendi, J, 2009:435

Americanura janssensi from Mexico
After Palacios-Vargas & al, 2009 Fig.8-9

Species macgregori Cassagnau & Palacios-Vargas, 1983:2

Species medellini Palacios-Vargas, JG et Ríos, G, 1996:437

Species mexicana Cassagnau, P, 1983:11, t.t.

Species najtae Arango, A, Villagomez, F et Palacios-Vargas, JG, 2017:52

Americanura najtae from Mexico
After Arango A, Villagomez F & Palacios-Vargas JG, 2017 Fig.1AC

Species nova (Christiansen, K & Reddell, JR, 1986:158) Palacios-Vargas, JG & Najt, J, 1986:23

Species palaciosi Paniagua Nucamendi, J, 2012:88

Species prima Cassagnau, P et Palacios-Vargas, JG, 1983:6

Species sardinasensis Palacios-Vargas, JG, Simón Benito, JC et Paniagua Nucamendi, J, 2009:433

Americanura sardinasensis from Mexico
After Palacios-Vargas & al, 2009 Fig.1-2

Species setafoliacea Cassagnau & Palacios-Vargas, 1983:2

             Syn.: Americanura setaefoliacea(sic) Palacios-Vargas, 1996 ISS
             Syn.: Americanura setaefoliacea(sic) Palacios-Vargas & Ríos, 1996 ISS

Species sotanophila Cassagnau & Palacios-Vargas, 1983:2

Species unguimitl Palacios-Vargas & Najt, 1986:11

             Syn.: Americanura unguimitli(sic) Palacios-Vargas, JG in Sánchez-Ramos, G, Reyes-Castillo, P et Dirzo, R, 2005:360 ISS



Genus Palmanura Cassagnau, P, 1983:11 key  key

Species sp. Arango, A, Palacios-Vargas, JG, Vázquez, M et Marín, EP, 2018:148

Species axelretanai Palacios-Vargas JG et Arango A in Arango, A, Palacios-Vargas, JG, Vázquez, M et Marín, EP, 2018:139

Palmanura axelretanai from Costa Rica
After Arango A et al, 2018 Fig.23+25

             Syn.: Palmanura axelreatanai(sic) Arango, A, Palacios-Vargas, JG, Vázquez, M et Marín, EP, 2018:129 ISS


Species colotlipa Palacios-Vargas, JG, 1996:820

Species contrerasi Palacios-Vargas JG et Arango A in Arango, A, Palacios-Vargas, JG, Vázquez, M et Marín, EP, 2018:134

Palmanura contrerasi from Mexico
After Arango A et al, 2018 Fig.13+16
Palmanura contrerasi from Mexico
After Arango A et al, 2018 Fig.18

Species curta Palacios-Vargas, JG, 1996:812

Species goyai Palacios-Vargas, JG et Simón-Benito, JC, 2009:268

Palmanura goyai from Mexico
After Palacios-Vargas JG & Simón-Benito JC, 2009 Fig.6,9
            Syn.: Palmanura sp.2 Palacios-Vargas, JG, García-Barros, E et Simón-Benito, JC, 2009:2



Species guilleni Palacios-Vargas, JG et Mejía-Madrid, HH, 2011:73

Species hieronimi Palacios-Vargas, JG et Simón-Benito, JC, 2009:268

            Syn.: Palmanura hieronimus(sic) Palacios-Vargas, JG et Simón-Benito, JC, 2009:268 IOS
Palmanura hieronimus(sic) from Mexico
After Palacios-Vargas JG & Simón-Benito JC, 2009 Fig.12
            Syn.: Palmanura sp.3 Palacios-Vargas, JG, García-Barros, E et Simón-Benito, JC, 2009:2



             Syn.: Palmanura hieronimus(sic) Palacios-Vargas, JG, García-Barros, E et Simón-Benito, JC, 2009:10 ISS
             Syn.: Palmanura hieronimus(sic) Palacios-Vargas, JG et Mejía-Madrid, HH, 2011:74 ISS



Species klompeni Palacios-Vargas JG et Vázquez M in Arango, A, Palacios-Vargas, JG, Vázquez, M et Marín, EP, 2018:129

Palmanura klompeni from Belize
After Arango A et al, 2018 Fig.3+4
Palmanura klompeni from Belize
After Arango A et al, 2018 Fig.6

Species lacandona Palacios-Vargas, JG, 1996:805

            Syn.: Palmanura lacandonicus(sic) Palacios-Vargas, JG, 1996:805 IOS
            Syn.: Palmanura sp. Palacios-Vargas, JG et Ríos-Saís, GA, 1995:266


             Syn.: Palmanura lancadona(sic) Palacios-Vargas, JG et Simón-Benito, JC, 2009:269 ISS



Species lindquisti Palacios-Vargas, JG, 1996:807

Species longa Palacios-Vargas, JG, 1996:821

Species matildae Palacios-Vargas, JG, 1996:806

             Syn.: Palmanura matildeae(sic) Palacios-Vargas, JG, 1996:806 ISS

Species mirabilis Cassagnau, P, 1983:11, t.t.

            Syn.: ... sp. Lavelle, Maury & Serrano, 1981


Species moralesi Palacios-Vargas JG et Arango A in Arango, A, Palacios-Vargas, JG, Vázquez, M et Marín, EP, 2018:132

Palmanura moralesi from Mexico
After Arango A et al, 2018 Fig.9+10

             Syn.: Palmanura morelasi(sic) Arango, A, Palacios-Vargas, JG, Vázquez, M et Marín, EP, 2018:146 ISS
             Syn.: Palmanura morelasi(sic) Arango, A, Palacios-Vargas, JG, Vázquez, M et Marín, EP, 2018:152 ISS


Species normae Palacios-Vargas, JG, 1996:818

Palmanura normae from Mexico
After Arango A et al, 2018 Fig.27+28

Species pascuali Palacios-Vargas, JG, 1996:814

Species primigenia Palacios-Vargas, JG, 1986:107

Species quetzalae Palacios-Vargas, JG, 1984:4

Species saham Vázquez, MM et Palacios-Vargas, JG, 2005:75

Species schusteri Palacios-Vargas, JG, 1996:809

Species sernai Palacios-Vargas, JG et Simón-Benito, JC, 2009:266

            Syn.: Palmanura sernaius(sic) Palacios-Vargas, JG et Simón-Benito, JC, 2009:266 IOS
Palmanura sernaius(sic) from Colombia
After Palacios-Vargas JG & Simón-Benito JC, 2009 Fig.4-5
            Syn.: Palmanura sp.1 Palacios-Vargas, JG, García-Barros, E et Simón-Benito, JC, 2009:2
            Syn.: Palmanura sernaius(sic) Palacios-Vargas, JG, García-Barros, E et Simón-Benito, JC, 2009:10



             Syn.: Palmanura sernaius(sic) Palacios-Vargas, JG et Mejía-Madrid, HH, 2011:74 ISS
Palmanura sernai from Colombia
After Arango A et al, 2018 Fig.30+31


             Syn.: Palmanura sernaius(sic) Arango, A, Palacios-Vargas, JG, Vázquez, M et Marín, EP, 2018:128 ISS




Species wilsoni Palacios-Vargas, JG, 1996:816

             Syn.: Palmanura wilson(sic) Palacios-Vargas, JG et Simón-Benito, JC, 2009:270 ISS



Genus Afrobella Cassagnau, P, 1983:19

Species delamarei Cassagnau, 1996

Species guineensis (Murphy, 1965) Cassagnau, P, 1983:16, t.t.

            Syn.: Neanura (Bilobella) guineense(sic) Murphy, 1965,tt IOS
            Syn.: Afrobella guineense(sic) Cassagnau, P, 1983:16 IOS
            Syn.: Bilobella guineense(sic) Thibaud, J-M, 2013:236 IOS

Species kirivatensis Cassagnau, 1996

Species mamillata Cassagnau, 1996

Species pauliani (Massoud, 1963) Cassagnau, P, 1983:21

Species scoparia Cassagnau, 1996

Species simplex (Massoud, 1963) Cassagnau, P, 1983:21

Species thibaudi Cassagnau, 1996

Species wittei Cassagnau, 1996


Genus Bilobella Caroli, E, 1912:367 key

  Syn.: Neanurini Neanura Caroli, E, 1910:321 partim


Bilobella carpatica from Ukraine
Prothoracic dorsal internal tubercles absent (*)
After Smolis A & Kaprus IJ, 2008 Fig.1

Species sp.1 Burkhardt U, Janssens F & Krediet A, 2019

Bilobella sp. from Germany
2018.03.10 © Krediet, A.

Species albanica Cassagnau, P et Peja, N, 1979:208

Species aurantiaca (Caroli, E, 1912:367) Stach, J, 1951:88, t.t.

Achorutes aurantiacus
After Caroli E, 1912 Fig.5
            Syn.: Neanura aurantiaca Caroli, E, 1910:321 nn
            Syn.: Bilobella burgalensis Simon & Pozo, 1987
            Syn.: Neanura (Bilobella) pyrenaea Selga, 1959



Remark by Maynard, E.A. 1951:70: Although the arrangement of the tubercles does not seem to agree with the meager description of this European species at my disposal, I am tentatively identifying my specimens as N. aurantiaca.

             Syn.: Bilobella auratiaca(sic) Stach, J, 1951:88 ISS
             Syn.: Neanura aurantica(sic) Rusek, J in Dlabola, J, 1977:13 ISS
Remark by Christiansen, K.A. &Bellinger, P.F. 1998:411: The occurence of this species in the Nearctic region is dubious. According to Maynard's description of his "aurantiaca", the head has 10 tubercles instead of the, 18(sic) [must be 8] described by Caroli; if this is correct, his species, at least, is probably distinct.

Bilobella aurantiaca
2005 © Domene, X.
Bilobella aurantiaca from Italy
2008.12.22 © Gregg, S.
Bilobella aurantiaca from France
2012.09.30 © Lebeaux, P.
Bilobella aurantiaca from Spain
After Arbea JI, 2013 Fig.7
Bilobella aurantiaca from Spain
2013.02.01 © Montoro, A.
Bilobella aurantiaca from Portugal
2014.04.13 © Wergen., B
Bilobella aurantiaca from Greece
With atypically deformed posterior body segments
2017.10.10 © Dimitrakopoulos, C.
Bilobella aurantiaca from Greece
2017.11.20 © Dimitrakopoulos, C.
Bilobella aurantiaca from France
2017.12.11 © Tiky, S.
Bilobella aurantiaca? juv. from the USA
Moulting
2017.12.27 © Abbott, J.
Bilobella aurantiaca? juv. from the USA
Moulting
2017.12.27 © Abbott, J.
Bilobella aurantiaca juv. from France
2018.04.09 © Tiky, S.
Bilobella aurantiaca from France
2018.11.10 © Tiky, S.
Bilobella aurantiaca from France
Mouthcone distinctly pointed
2019.10.27 © Dejoux, P.
Bilobella aurantiaca from France
2019.10.31 © Garcelon, P.

             Syn.: Bilobella auriantaca(sic) Cucini, C, Fanciulli, PP, Frati, F, Convey, P, Nardi, F et Carapelli, A, 2021:12 ISS
Bilobella aurantiaca from France
2024.03.11 © Séb, R.



Species braunerae Deharveng, L, 1981:96

Bilobella braunerae from Luxembourg
After Stomp N & Weiner WM, 2005 Fig.7
Bilobella aurantiaca(sic) from Holland
Legit Berg, M.
2007.03.13 © Heijerman, T.
Bilobella braunerae from France
2008.03.02 © Lebeaux, P.
 
Bilobella braunerae from France
2008.03.02 © Lebeaux, P.
Bilobella braunerae from France
2008.03.02 © Lebeaux, P.
Bilobella braunerae from Holland
After Berg M, 2009 Fig.1 © Heijerman T
Bilobella braunerae from Germany
2010.04.05 © Marquard, F.
Bilobella braunerae from Germany
2010.04.05 © Marquard, F.
Bilobella braunerae from Germany
2010.04.23 © Marquard, F.
Bilobella braunerae from Germany
2010.05.25 © Marquard, F.
Bilobella braunerae from Germany
2014.08.16 © Beck, K.
Bilobella braunerae from Germany
2014.08.16 © Beck, K.
Bilobella braunerae from Germany
2014.08.16 © Kluck, A.
Bilobella braunerae from Germany
1+3+4 cephalic tubercles
2014.08.21 © Kluck, A.
Bilobella braunerae from Germany
Ocular tubercle with 2 ocelli
2014.08.21 © Kluck, A.
Bilobella braunerae from Germany
Flower shaped texture of dorsal tubercles
2014.08.21 © Kluck, A.
Bilobella braunerae from Germany
2014.10.10 © Kluck, A.
Bilobella braunerae from Germany
Pointed mouthcone
2014.10.10 © Kluck, A.
Bilobella braunerae from Germany
Ocular tubercle with 2 ocelli
2014.10.21 © Kluck, A.
Bilobella braunerae from Germany
2014.10.21 © Kluck, A.
Bilobella braunerae from the UK
Leg.: Boardman, P. Det.: Shaw, P.
2015.01.28 © Shaw, P.
Bilobella braunerae from the UK
2015.01.dd © Boardman, P.
Bilobella braunerae from Germany
Sharply pointed mouth-cone
2015.08.08 © Kluck, A.
Bilobella braunerae juv. from Germany
2015.08.08 © Kluck, A.
Bilobella braunerae from Germany
2015.08.31 © Beck, K.
Bilobella braunerae from Belgium
Abd.6 partly hidden under abd.5
2017.03.08 © Huskens, M.-L.
Bilobella braunerae from Belgium
2 ocelli without pigment
2017.03.08 © Huskens, M.-L.
Bilobella braunerae from Belgium
Sharply pointed mouthcone
2017.03.10 © Huskens, M.-L.
Bilobella braunerae from Slovenia
2017.03.10 © Pisec, N.
Bilobella braunerae from Germany
2018.03.10 © Krediet, A.
Bilobella braunerae from France
Dorso-internal tubercles on th.1 absent
2020.02.06 © Dejoux, P.
Bilobella braunerae from France
2020.02.06 © Dejoux, P.

Species carpatica Smolis, A et Kaprus, IJ, 2008:510

Species coiffaiti Cassagnau, 1968

Species digitata Cassagnau, 1967

Bilobella digitata
After Deharveng L, 1983 Fig.1F
Bilobella digitata
After Potapov M in Babenko A, 1988
Bilobella digitata from Greece
2017.12.30 © Dimitrakopoulos, C.
Bilobella digitata from Greece
Abd.4 tubercles distinctly digitally elongated
2017.12.30 © Dimitrakopoulos, C.
Bilobella digitata from Greece
2017.12.30 © Dimitrakopoulos, C.
Bilobella digitata from Croatia
2019.03.dd © Lukic, M.
Bilobella digitata from Croatia
2019.03.dd © Lukic, M.

Species excolorata (Loksa, I et Rubio, I, 1966:142)

Neanura excolorata from Hungary
After Loksa I & Rubio I, 1966 Abb.17

Species ligurica Deharveng, L, 1981:97

Species mahunkai Dányi, L, 2010:29

Bilobella mahunkai from Montenegro
After Dányi, L., 2010 Fig.2
Bilobella mahunkai from Montenegro
After Dányi, L., 2010 Fig.2

Species massoudi Cassagnau, 1968

             Syn.: Bilobella massoudis(sic) Frati, F et Dell'Ampio, E, 2000:345 ISS

Species matsakisi Cassagnau, 1967

Bilobella matsakisi from Albania
After Traser G & Kontschán J, 2004 Fig.7

Species proxima Cassagnau, P et Peja, N, 1979:209

Species subaurantiaca Cassagnau, P et Peja, N, 1979:210

Bilobella subaurantiaca? from Czechia
2007.07.05 © Krásenský, P.
Bilobella subaurantiaca? from Czechia
2007.07.05 © Krásenský, P.
Bilobella subaurantiaca? from Slovenia
2017.03.04 © Pisec, N.
Bilobella subaurantiaca? from Slovenia
2017.03.10 © Pisec, N.

Species zekoi Cassagnau, P et Peja, N, 1979:210


Genus Chaetobella Cassagnau, P, 1983:19

Species buddha Cassagnau, 1993

Species eurygaster Cassagnau, 1993

Species multiseta Cassagnau, 1993

Species numatai (Yosii, 1966) Cassagnau, P, 1983:16, t.t.

Species rhodophila Cassagnau, 1993

Species sigillata Cassagnau, 1993

Species sylvatica Cassagnau, 1993

Species tonglica Cassagnau, 1993

Species verticicola Cassagnau, 1993


Genus Travura Cassagnau, P et Deharveng, L, 1980:2

Species betschi Cassagnau, P et Deharveng, L, 1980:6

Species divergens Cassagnau, P et Deharveng, L, 1980:3, t.t.

Species mikenensis Cassagnau, 2000


Genus Adbiloba Stach, 1951

Species intermedia Cassagnau, 2000

Species leleupi Cassagnau, 2000

Species plurichaetosa Cassagnau, P et Peja, N, 1979:206

Species prodoni Cassagnau, 1993

Species sikkimensis Yosii, 1966

Neanura sikkimensis from India
After Mandal GP & Hazra AK, 2009 Fig.8a

Species sokolowi (Philiptschenko, 1926) Stach, 1951, t.t.

             Syn.: Achorutes Sokolovi(sic) Denis, J-R, 1933:229 ISS


Genus Thaianura Yosii, 1961

  Syn.: Neanurinae Thaïanura(sic) Deharveng, L, 1983:6 ISS
  Syn.: Neanurinae Thaïanura(sic) Deharveng, L, 1983:22 ISS
  Syn.: Neanurinae Thaïanura(sic) Deharveng, L, 1983:24 sp ISS
  Syn.: Neanurinae Thaïanura(sic) Deharveng, L, 1983:25 ISS
  Syn.: Neanurinae Thaïanura(sic) Deharveng, L, 1983:27 ISS

Species umesaoi Yosii, 1961, t.t.


Genus Womersleya Denis, 1948

Species formosana Lee, B-H et Kim, J-T, 1990:239

Womersleya formosana from Taiwan
After Lee B-H & Park K-H, 1990 Fig.3A

Species marhia Baijal, HN, 1958:351

Womerselya(lapsus) marhia from the Himalaya
After Baijal HN, 1958 Fig.2A.

Species vicina (Denis, 1934) Denis, 1948, t.t.

Tribus Morulodini Cassagnau P, 1983:5
            Syn.: ... Protanurini Massoud, Z, 1967:284


             Syn.: Neanurinae Murolodini(sic) Arango, A, Palacios-Vargas, JG, Vázquez, M et Marín, EP, 2018:151 ISS


Genus Archimorulodes Smolis, A et Pasnik, G, 2020:9


Genus Morulodes Cassagnau, P, 1955:160

  Syn.: ... Neanura MacGillivray, 1893 partim
  Syn.: ... Kodiakia Bodvarsson, 1960


Morulodes sp. juv. from the USA
2023.12.26 © Wang, G.
Morulodes sp. juv. from the USA
Abd.6 under abd.5
2023.12.26 © Wang, G.
Morulodes sp. juv. from the USA
Maxilla with many teeth
2023.12.26 © Wang, G.
Morulodes sp. juv. from the USA
Mandibula with 7 teeth
2023.12.26 © Wang, G.
Morulodes sp. from the USA
Macrosetae unilaterally serrate
2023.12.26 © Wang, G.
Morulodes sp. juv. from the USA
Midfoot
Unguis without tooth
2023.12.26 © Wang, G.

Species ambiguus (Christiansen, K & Bellinger, P, 1980:368) Cassagnau, P, 1983:6

Neanura (Lobella) ambigua from the USA
Modified after Christiansen K & Bellinger P, 1980 Fig.276A
Morulodes ambiguus from Canada
2017.12.05 © Barbin, T.
Morulodes ambiguus from Canada
2017.12.05 © Barbin, T.
Morulodes ambiguus from Canada
Eyepatch with 3+1 ocelli
2017.12.05 © Barbin, T.
Morulodes ambiguus from Canada
Dorso-internal tubercles on abd.4 fused
Abd.6 hidden under abd.5
2017.12.05 © Barbin, T.
Morulodes ambiguus from the USA
2017.12.06 © Dobbs, Br.
Morulodes ambiguus from the USA
Dorso-internal tubercles on abd.4 fused
Abd.6 hidden under abd.5
2017.12.06 © Dobbs, Br.
Morulodes ambiguus from the USA
Head with 1+3+4 dorsal tubercles
2017.12.06 © Dobbs, Br.
Morulodes ambiguus from the USA
Broad labrum with long setae
2017.12.06 © Dobbs, Br.
Morulodes ambiguus from the USA
2022.12.30 © G, S.
Morulodes ambiguus from the USA
Eye with 3 anterior ocelli
2022.12.30 © G, S.
Morulodes ambiguus from the USA
Abd.6 hidden under abd.5
2022.12.30 © G, S.
Morulodes ambiguus from the USA
Abd.4 with fused dorso-internal tubercles
2022.12.30 © G, S.

Species rishirianus Tanaka & Ischisawa, 2002

Species serratus (Folsom, 1916 tt) Massoud, Z, 1967:287

            Syn.: Morulodes millsi Cassagnau, P, 1955:160
            Syn.: Kodiakia aleuta Bödvarsson, H, 1960:46
            Syn.: Kodiakia prima Bödvarsson, H, 1960:43


Neanura (Morulodes) serrata from the USA
Modified after Christiansen K & Bellinger P, 1998 Fig.288A
Morulodes serratus from the USA
Setae very heavily serrate
2016.11.26 © Dobbs, Br.
Morulodes serratus from the USA
Bilobed 6th abdominal segment hidden under 5th
2016.11.26 © Dobbs, Br.
Morulodes serratus from Canada
Ocular tubercle with 3 anterior ocelli + 1 posterior ocellus
2019.09.15 © Barbin, T.
Morulodes serratus from the USA
2021.01.15 © Degenhardt, J.
Morulodes serratus from the USA
Ocular tubercle with 4 ocelli
3 anterior + 1 posterior
2021.01.15 © Degenhardt, J.
Morulodes serratus from the USA
Distinctly serrate macrosetae
2021.01.15 © Degenhardt, J.
Morulodes serratus from the USA
Tubercles elongate
Darker than background colour
2021.01.15 © Degenhardt, J.
Morulodes serratus from the USA
Abd.4 dorso-external tubercles elongate
2023.01.01 © Wang, G.
Morulodes serratus from the USA
Cephalic tubercles
2023.01.01 © Wang, G.
Morulodes serratus from the USA
3rd unguis
2023.01.01 © Wang, G.

Species setosus (Canby, 1926) Deharveng, 1982

Neanura (Morulodes) setosa from the USA
Modified after Christiansen K & Bellinger P, 1998 Fig.289A
Morulodes setosus juv. from Canada
2013.04.13 © Hochhalter, R.
Morulodes setosus juv. from Canada
Eyepatch with 3 anterior and 1 posterior ocelli
2013.04.13 © Hochhalter, R.
Morulodes setosus juv. from Canada
Bilobed 6th abdominal segment hidden under 5th
2013.04.13 © Hochhalter, R.
Morulodes setosus ♂ from the USA
Abd.4 internal tubercles not fused
Abd.6 hidden under abd.5
2022.11.26 © Wang, G.
Morulodes setosus ♂ from the USA
Dorsal macrosetae unilaterally serrate
2022.11.26 © Wang, G.
Morulodes setosus ♂ from the USA
Eye with 3 + 1 ocelli
2022.11.26 © Wang, G.
Morulodes setosus ♂ from the USA
Mouthcone
2022.11.26 © Wang, G.
Morulodes setosus ♂ from the USA
Remnant furcula
2022.11.26 © Wang, G.
Morulodes setosus ♂ from the USA
Male genital papilla
2022.11.26 © Wang, G.
Morulodes setosus ♂ from the USA
Foot with reduced unguiculus (*)
2022.11.27 © Wang, G.
Morulodes setosus ♂ from the USA
Unguis with distinct inner tooth
2022.11.29 © Wang, G.
Morulodes setosus from the USA
Freshly deposited egg cluster
2023.01.28 © Wang, G.
Morulodes setosus from the USA
Egg cluster
Eyes of embryo's visible
2023.02.12 © Wang, G.
Morulodes setosus from the USA
Hatchlings
2023.02.17 © Wang, G.
Morulodes setosus from the USA
Juvenile
2023.02.17 © Wang, G.
Morulodes setosus from the USA
2023.03.18 © Wang, G.
Morulodes setosus from the USA
Unguis with distinct inner tooth
2023.03.18 © Wang, G.
Morulodes setosus juv. from the USA
2023.03.18 © Wang, G.

Tribus Lobellini Yosii R, 1965:13 sensu Cassagnau P, 1983:8
            Syn.: ... Achorutini Handschin, E in Dahl, F, 1929:39 partim
            Syn.: Neanurinae Crossodonthini(sic) Massoud, Z, 1967:342 partim IOS
             Syn.: ... Crossodonthini(sic) Deharveng, L, 2004:424 partim ISS
            Syn.: ... Neanurini Deharveng, L, 2004:424 partim
            Syn.: ... Protanurini Deharveng, L, 2004:424 partim



Neanurinae lignée lobellienne
Cephalic tubercles
After Deharveng L, 1983 Fig.13A




Genus Archilobella Smolis, A et Pasnik, G, 2020:9


Genus Coreanura Deharveng, L et Weiner, WM, 1984:27

Species filiae Deharveng, L et Weiner, WM, 1984:27, t.t.


Genus Propeanura Yosii, R, 1956:46

Species garuda Yoshii, 1976

Species hygrophila (Cassagnau, P et Deharveng, L, 1981:158)

Species hypostoma (Denis, 1929)

Species kymgangensis Deharveng, L et Weiner, WM, 1984:44

Species labukae Yoshii, 1981

Species lapidicola Cassagnau, P et Deharveng, L, 1981:155

Species leei Deharveng, L et Weiner, WM, 1984:46

Species megalops Deharveng, L et Weiner, WM, 1984:48

Species nomurai (Yosii, R, 1956:43, i.s.) Deharveng & Weiner, 1984

Yuukianura nomurai from Japan
After Yosii R, 1956 Pl.XXII Fig.131

Species parvituberculata (Yosii, R, 1965:21) Cassagnau, P, 1983:22

Bilobella parvituberculata from Japan
After Yosii R, 1965 Fig.15A
             Syn.: Neanura parvitubercula(sic) Massoud, Z, 1967:305 ISS


Species pterothrix (Börner, C, 1909:107) Yosii, 1959, t.t.

             Syn.: Neanura pterothryx(sic) Yosii, 1954 ISS
             Syn.: Neanura (Propeanura) pterothryx(sic) Yosii, R, 1956:47,tt ISS
             Syn.: Neanura pterothryx(sic) Massoud, 1967 ISS
             Syn.: Propeanura pterothryx(sic) Cassagnau, P, 1983:15,tt ISS

Species soemarwotoi Yosii, 1976

Species zehntneri (Handschin, 1920)

             Syn.: Achorutes Zehntneri(sic) Denis, J-R, 1933:231 ISS


Genus Coecoloba Yosii, R, 1956:33

Species hidana Yosii, R, 1956:36

Coecoloba hidana from Japan
After Yosii R, 1956 Pl.XIX Fig.115
             Syn.: Coecoloba hidani(sic) Massoud, 1967 ISS


Species kurasawana Yosii, R, 1956:35

Coecoloba kurasawana from Japan
After Yosii R, 1956 Pl.XVII Fig.96

Species lobella (Yosii, R, 1954:65) Yosii, R, 1956:34, t.t.

Coecoloba lobella from Japan
After Yosii R, 1956 Pl.XVII Fig.95

Species odai Yosii, R, 1956:36

Coecoloba odai from Japan
After Yosii R, 1956 Pl.XIX Fig.116

Species onychiuriformis (Yosii, 1976) Deharveng, L, 1983:59

Species plumleyi Deharveng, L, 1983:61

Species wakasana Yosii, R, 1956:37

Coecoloba wakasana from Japan
After Yosii R, 1956 Pl.XX Fig.117


Genus Lobella Börner, C, 1906:168 key

  Syn.: ... Biocularia Stach, J, 1951:6 partim
  Syn.: Lobella Telobella Cassagnau, P, 1983:18
  Syn.: Lobellini Telobella Ohira, A, Nakamori, T, Matsumoto, S, Bessho-Uehara, M, Kato, T et Oba, Y, 2023:67


Lobella sp. from Japan
2008 © Tsukiji, T.
Lobella sp. from Japan
2008 © Tsukiji, T.
Lobella sp. from Japan
2008 © Tsukiji, T.
Lobella sp. from Indonesia
2013.10.26 © Bertner, P.
Lobella sp. from Taiwan
2021.03.19 © Huate.
Lobella sp. from Taiwan
2021.03.19 © Huate.
Lobella sp. from Taiwan
Ventral aspect
2021.03.19 © Huate.
Lobella sp. from Taiwan
Anus
Dorso-ventral aspect
2021.03.19 © Huate.
Lobella? sp. from Taiwan
2021.09.28 © Cheng, H.-J.
Lobella sp. from Thailand
From culture in the USA
2022.11.28 © Pavy, R.
Lobella sp. from Thailand
From culture in the USA
2022.11.28 © Pavy, R.
Lobella sp. from Thailand
From culture in the USA
2022.11.28 © Pavy, R.
Lobella sp. from Thailand
Eye with 2 + 1 ocelli
From culture in the USA
2022.11.28 © Pavy, R.
Lobella sp. from Thailand
Pointed mouthcone
From culture in the USA
2022.11.28 © Pavy, R.
Lobella sp. from Thailand
Eye with 2+1 ocelli
From culture in the USA
2022.12.10 © Pavy, R.
Lobella sp. from Thailand
Eye with 2 anterior ocelli + 1 posterior ocellus
From culture in the USA
2022.12.10 © Pavy, R.
Lobella sp. from culture in the USA
Feeding from yeast
With everted distinctly long mouthparts
2023.12.13 © Gruver, Be.

Species fortis (Oudemans, JT in Weber, M, 1890:91) Yayuk, RS (sic), 1989:7

            Syn.: Lobella? fortis Ohira, A, Nakamori, T, Matsumoto, S, Bessho-Uehara, M, Kato, T et Oba, Y, 2023:68

Species palmeri (Wray, 1967) Palacios-Vargas & Najt, 1986:25

             Syn.: Lobella pulmei(sic) Kasai, H, Tanaka, S et Sawahata, T, 2023:517 lapsus ISS


Lobella palmeri from the USA
After Bernard, E.C., 2006 ©
Lobella palmeri from the USA
2015.05.04 © Anderson, J.
Lobella palmeri from the USA
2015.05.04 © Anderson, J.
Lobella palmeri from the USA
2019.06.08 © Montgomery, G.
Lobella palmeri from the USA
2019.06.08 © Montgomery, G.

            Syn.: Lobella? palmeri Ohira, A, Nakamori, T, Matsumoto, S, Bessho-Uehara, M, Kato, T et Oba, Y, 2023:68

Species-group sauteri


Species bicincta Cassagnau, P et Deharveng, L, 1984:3

Species bicolor Cassagnau, P et Deharveng, L, 1984:4

Species kemiri (Suhardjono, YR & Deharveng, L, 2001:?) Ohira, A, Nakamori, T, Matsumoto, S, Bessho-Uehara, M, Kato, T & Oba, Y, 2023:68

Species monocincta Cassagnau, P et Deharveng, L, 1984:4

Species punctata Cassagnau, P et Deharveng, L, 1984:7

Species sandakanensis Yoshii, 1981

             Syn.: Propeanura sandakensis(sic) Yoshii, R et Suhardjno, YR, 1992:4 ISS
             Syn.: Lobella (Propeanura) sandakaensis(sic) Ohira, A, Nakamori, T, Matsumoto, S, Bessho-Uehara, M, Kato, T et Oba, Y, 2023:68 ISS

Species sauteri (Börner, C, 1906:168) Denis, J-R, 1933:231, t.t.

            Syn.: Neanura takaoensis Kinoshita, 1932
            Syn.: Neanuridae Lobella Oba, Y, Branham, MA et Fukatsu, T, 2011:773 sp
Lobella sp. from Japan
Probably self-luminous
After Oba Y & al, 2011 Fig.7E
Lobella sp. from Japan
Emits a continuous weak green light from abdominal tubercles by stimulation
After Oba Y & al, 2011 Fig.7F

            Syn.: Lobellini Lobella Ohira, A, Nakamori, T, Matsumoto, S, Bessho-Uehara, M, Kato, T et Oba, Y, 2023:76 sp



             Syn.: Neanura (Lobella) Sauteri(sic) Carpenter, GH, 1916:30 ISS
             Syn.: Lobella Sauteri(sic) Denis, J-R, 1933:231 ISS
Lobella sauteri from Japan
After Ohira A & al, 2023 Fig.1
Lobella sauteri from Japan
Modified after Ohira A & al, 2023 Fig.5
Lobella sauteri from Japan
After Ohira A & al, 2023 Fig.24 top left
Lobella sauteri from Japan
Bioluminescence in dim light after stimulation
After Ohira A & al, 2023 Fig.24 top middle
Lobella sauteri from Japan
Bioluminescence in dark after stimulation
After Ohira A & al, 2023 Fig.24 top right
Lobella sauteri from Japan
Defence bleeding after stimulation
After Ohira A & al, 2023 Fig.26



Species setapauca Gapud, 1970

            Syn.: Lobella reducta Gapud, 1968


Species wayang (Yosii, 1976) Cassagnau, P, 1983:15

Species yambaru (Tanaka & Hasegawa, 2010) Ohira A & Nakamori T in Ohira, A, Nakamori, T, Matsumoto, S, Bessho-Uehara, M, Kato, T & Oba, Y, 2023:76


Species-group monstrum

Species decipiens Yosii, R, 1965:20

Lobella decipiens from Japan
Modified after Yosii R, 1965 Fig.14A

             Syn.: Lobella (monstrum-group) decipens(sic) Ohira, A, Nakamori, T, Matsumoto, S, Bessho-Uehara, M, Kato, T et Oba, Y, 2023:82 ISS


Species mizunasiana Yosii, 1956

             Syn.: Lobella (Lobellina) mizunashiana(sic) Yosii, R, 1977:149 ISS
             Syn.: Lobella (Lobella) mizunashiana(sic) Yosii, R, 1977:149 ISS
             Syn.: Lobella mizunaziana(sic) Deharveng, L, 1983:23 ISS

Species monstrum Ohira A et Nakomori T in Ohira, A, Nakamori, T, Matsumoto, S, Bessho-Uehara, M, Kato, T et Oba, Y, 2023:76

Lobella monstrum from Japan
After Ohira A & al, 2023 Fig.15

            Syn.: Lobella sauteri Yoshii, 1994
            Syn.: Lobella sauteri Ohira, A, Nakamori, T, Matsumoto, S, Bessho-Uehara, M, Kato, T et Oba, Y, 2023:76



Species nana Lee, B-H et Kim, J-T, 1990:242

Lobella nana from Taiwan
After Lee B-H & Park K-H, 1990 Fig.5A

Species sokamensis Deharveng, L et Weiner, WM, 1984:31

Species stachi Yosii, R, 1956:39

             Syn.: Lobella stachii(sic) Lee, B-H, 1982:220 ISS


Genus Protolobella Cassagnau, P, 1983:18

Species assamensis Yosii, 1966, t.t.

Lobella (Lobella) assamensis from India
After Mandal GP & Hazra AK, 2009 Fig.10a+b

Species maxillaris (Yosii, 1966) Ohira A & Nakamori T in Ohira, A, Nakamori, T, Matsumoto, S, Bessho-Uehara, M, Kato, T & Oba, Y, 2023:69


Genus Lobellina Yosii, 1956 key

Species sp. Yosii, R, 1977:149

            Syn.: Lobella sauteri Yosii, 1956 nec Börner


Species chosonica Deharveng, L et Weiner, WM, 1984:35

Species fusa Jiang, J-G, Wang, W-B, Xia, H, 2018:4

Lobellina fusa from China
Modified after Jiang J-G, Wang W-B & Xia H, 2018 Fig.1

Species gedehensis (Yosii, 1976), i.s.

Species gladius Hu, Jiang, Jiang & Jiang, 2019

Species hunanensis Wang, Y-R, Wang, W-B et Jiang, J-G, 2016:191

Lobellina hunanensis from China
After Wang Y-R, Wang W-B & Jiang J-G, 2016 Fig.15

Species ionescui (Massoud, Z & Gruia, M, 1973:333) Cassagnau, P, 1983:10

Lobella ionescui from Cuba
After Massoud Z & Gruia M, 1973 Fig.5G

Species ipohensis (Yosii, 1976) Deharveng, L & Weiner, WM, 1984:34

Species kitazawai (Yosii, 1969) Deharveng, L & Weiner, WM, 1984:34

Species minuta (Lee, B-H, 1980:252) Deharveng, L & Weiner, WM, 1984:34

Lobella minuta from Korea
After Lee B-H, 1980 Fig.1A

Species montana Deharveng, L et Weiner, WM, 1984:37

Species musangensis (Yosii, 1976) Deharveng, L & Weiner, WM, 1984:34

Species nanjingensis Ma, Z-C et Chen, J-X, 2008:62

Species paraminuta Deharveng, L et Weiner, WM, 1984:39

Species perfusionides (Stach, J, 1965:346) Deharveng, L & Weiner, WM, 1984:34

             Syn.: Lobella perfusoïdes(sic) Massoud, Z, 1967:333 ISS

Species pomorskii Smolis, 2017

Species proxima Deharveng, L et Weiner, WM, 1984:41

Species roseola (Yosii, 1954) Cassagnau, P, 1983:15, t.t.

Species weinerae Smolis, 2017

Species yinae Jiang, J-G, Huang, C-W et Luan, Y-X, 2018:15


Genus Cassagnaua Özdikmen, H, 2009:293

  Syn.: Lobellienne Pectinura Cassagnau, P, 1983:18 nec Forbes, 1843


Species hongkongensis (Yosii, 1976) Özdikmen, H, 2009:294, t.t.

             Syn.: Womersleya honkongensis(sic) Liu, Y-Q, Hou, D-B et Li, Z-C, 1998:126 ISS


Genus Crossodonthina Yosii, 1954 key

  Syn.: Neanurinae Crossondonthina(sic) Lee, B-H, 1980:254 ISS

Species acuminata Jiang & Wang, 2021

Species alatoserrata Yosii, R, 1965:18

Crossodonthina alatoserrata from Taiwan
Modified after Yosii R, 1965 Fig.13A

Species altamontana Yoshii, 1981

             Syn.: Crossodenthina altamontana Yayuk, 1989 ISS

Species appendiculata Yosii, R, 1956:41

Species bidentata Luo & Chen, 2009

             Syn.: Crossodonthina bidenata(sic) Kasai, H, Tanaka, S et Sawahata, T, 2023:516 ISS

Species choui Jiang & Zhang, 2012

Species clavata Jiang & Wang, 2021

Species elegans Kasai, H, Tanaka, S et Sawahata, T, 2023:510

Crossodonthina elegans from Japan
After Kasai H & al, 2023 Fig.1
Crossodonthina elegans from Japan
Dorsal chaetotaxy
After Kasai H & al, 2023 Fig.2

Species formosana Yosii, R, 1965:17

Crossodonthina formosana from Taiwan
After Yosii R, 1965 Fig.12A

Species hainana Xiong, Chen & Yin, 2005

Species koreana Yosii, R et Lee, C-E, 1963:1

Crosodonthina koreana from Korea
After Yosii R & Lee C-E, 1963 Fig.1A

            Syn.: Crossodonthina delamarei Lee, B-H, 1973:439
Crossodonthina delamarei from Korea
After Lee B-H, 1973 Fig.1A




            Syn.: Crossondonthina(sic) koreana Lee, B-H, 1980:254

Species langshanensis Hu, Jiang, Jiang & Jiang, 2019

Species laterisensillata Ohira, A, Kataoka, M, Tanooka, A et Nakamori, T, 2022:20

Crossodonthina laterisensillata from Japan
After Ohira A & al, 2022 Fig.2A

Species montana Lee, B-H et Kim, J-T, 1990:240

Crossodonthina montana from Taiwan
After Lee B-H & Park K-H, 1990 Fig.4A

Species nipponica Yosii, 1954 tt

            Syn.: Crossodonthina japonica(sic) Yosii, 1959 lapsus
            Syn.: Crossodonthina japonica(sic) Yosii, R et Lee, C-E, 1963:2 lapsus
            Syn.: Crossodonthina appendiculata Yoshii, 1995


Crossodonthina nipponica from Japan
After Yosii R, 1956 Pl.XX Fig.118
Crossodonthina nipponica from Japan
Modified after Ohira A & Nakamori T, 2023 Fig.1B

Species radiata (Salmon, JT, 1941:308) Massoud, Z, 1967:343

Neanura radiata from New Zealand
After Salmon J.T., 1941 Pl.47 Fig.117

Species tiantongshana Xiong, Chen & Yin, 2005

Species tridentiens Yue, Q-Y et Yin, W-Y, 1999:222


Genus Sphaeronura Cassagnau, P, 1983:19

Species ameliae Smolis & Greenslade, 2020

Species bornensis (Schött, 1925) Deharveng, L & Bedos, A, 2000:210

Remark by Deharveng, L. & Bedos, A. 2000:210: It is likely that the Borneo and the Vietnam forms belong to different species.

Species chaotica (Yosii, 1976) Cassagnau, P, 1983:15, t.t.


Genus Riozura Cassagnau, P, 1983:18

Species yoshibai (Yosii, 1963) Cassagnau, P, 1983:15, t.t.


Genus Yuukianura Yosii, R, 1955:383

  Syn.: Lobellini Yukianura(sic) Smolis, A, Skazynski, D, Kahrarian, M et Kaprus, IJ, 2016:299 ISS
?Yuukianura? sp. from Japan
2018.11.02 © Hara, Y.
?Yuukianura? sp. from Japan
Ant.3+4 fused; 2 ocelli
2018.11.02 © Hara, Y.
?Yuukianura? sp. from Japan
2018.11.02 © Hara, Y.



Species aphoruroides (Yosii, R, 1953:69) Yosii, R, 1955:384, t.t.

Protanura aphoruroides from Japan
After Yosii R, 1953 Fig.2.7
Yuukianura aphoruroides from Japan
After Yosii R, 1956 Pl.XXI Fig.119
Lobella (Yuukianura) aphoruroides from China
After Stach J, 1964 Plate III Fig.1
Yuukianura aphoruroides from the UK
In cultures of earthworms
After Hopkin SP, 2007 Fig.7b
Yuukianura aphoruroides from the USA (introduced)
Dorsal tubercles poorly developed
2022.02.17 © Pavy, R.
Yuukianura aphoruroides from the USA (introduced)
Dorsal tubercles poorly developed
Abd.6 visible in dorsal aspect
2022.02.17 © Pavy, R.
Yuukianura aphoruroides from the USA (introduced)
Mouthcone pointed but truncated
2022.02.17 © Pavy, R.
Yuukianura aphoruroides from the USA (introduced)
Mouthcone pointed but truncated
2022.02.17 © Pavy, R.
Yuukianura aphoruroides from the USA (introduced)
Dorsal tubercles poorly developed
2022.02.18 © Pavy, R.
Yuukianura aphoruroides from the USA (introduced)
Tubercles poorly developed
2022.09.26 © Pavy, R.
Yuukianura aphoruroides from the USA (introduced)
Abd.6 bilobed
2022.09.26 © Pavy, R.
Yuukianura aphoruroides from the USA (introduced)
Ocular tubercle with 2+1 ocelli (*)
2022.09.26 © Pavy, R.
Yuukianura aphoruroides from the USA (introduced)
2022.10.17 © Pavy, R.
Yuukianura aphoruroides from the USA (introduced)
2022.10.17 © Pavy, R.
Yuukianura aphoruroides from the USA (introduced)
2022.10.17 © Pavy, R.
Yuukianura aphoruroides from the Canada (introduced)
In culture
2022.11.20 © Chung, W.K.H.
Yuukianura aphoruroides from the Canada (introduced)
Eye with 3 unpigmented ocelli
2022.11.20 © Chung, W.K.H.
Yuukianura aphoruroides from the USA (introduced)
Collophore apex opens to stuck collophore vesicles against glass of terrarium
2022.11.30 © Pavy, R.
Yuukianura aphoruroides from the USA (introduced)
Cleaning mouthcone using 6 legs
Collophore stucked against glass of terrarium
2022.11.30 © Pavy, R.
Yuukianura aphoruroides from Taiwan
Dorso-internal tubercles absent
In commercial culture
2023.04.06 © Cheng, H.-J.
Yuukianura aphoruroides from Taiwan
Lateral aspect
In commercial culture
2023.04.06 © Cheng, H.-J.
Yuukianura aphoruroides from Taiwan
Ventral aspect
In commercial culture
2023.04.06 © Cheng, H.-J.
Yuukianura aphoruroides from Taiwan
Unguis with inner tooth
In commercial culture
2023.04.06 © Cheng, H.-J.
Yuukianura aphoruroides from Taiwan
Maxilla head dented
In commercial culture
2023.04.06 © Cheng, H.-J.
Yuukianura aphoruroides from Taiwan
Mandibula dented
In commercial culture
2023.04.06 © Cheng, H.-J.
Yuukianura aphoruroides from Taiwan
Abd.5 with 2+2 dorsal tubercles
In commercial culture
2023.04.06 © Cheng, H.-J.
Yuukianura aphoruroides from the UK
2023.11.05 © Geckoman, R.
Yuukianura aphoruroides from the UK
2023.11.05 © Geckoman, R.

Species deharvengi Smolis, 2017

Species halophila Yosii, R, 1955:384

Yuukianura halophila from Tokara Island
After Yosii R, 1955 Abb.3A

Species hawaiiensis (Bellinger, PF & Christiansen, KA, 1974:31) Deharveng, L, Palacios-Vargas, JG & Bedos, A, 2017:58

Species judithae Deharveng, L, Palacios-Vargas, JG et Bedos, A, 2017:58

Yuukianura judithae from Vanuata
After Deharveng L, Palacios-Vargas JG & Bedos A, 2017 Fig.1B

Species pacifica (Yosii, 1971) Deharveng, L, Palacios-Vargas, JG & Bedos, A, 2017:58

Species rosea (Kim, J-T & Lee, B-H, 2000:180) Deharveng, L, Palacios-Vargas, JG & Bedos, A, 2017:58

Species szeptyckii Deharveng, L et Weiner, WM, 1984:50

Species tongana (Yosii, R, 1964:9) Deharveng, L, Palacios-Vargas, JG & Bedos, A, 2017:57

Species yasudai (Yosii, 1966:479) Deharveng, L, Palacios-Vargas, JG & Bedos, A, 2017:57


Genus Deuterobella Yoshii, R et Suhardjno, YR, 1992:5

Species imadatei (Yosii, 1976), t.t.

Species murphyi (Yosii, 1976) Deharveng, L & Bedos, A, 2000:210

Remark by Deharveng, L. & Bedos, A. 2000:210: It is likely that the Malaya and the Vietnam forms belong in fact to different species.

Species tawauensis (Yoshii, 1981)

Species ternatensis (Yoshii, R et Suhardjno, YR, 1992:6)


Genus Yosialina Salmon, 1964

Species cavicola (Yosii, 1956) Ohira A & Nakamori T in Ohira, A, Nakamori, T, Matsumoto, S, Bessho-Uehara, M, Kato, T & Oba, Y, 2023:69

Lobellina? cavicola from Japan
Abd.1-4 dorso-internal tubercles absent
2014.08.23 © Hara, Y.
Lobellina? cavicola from Japan
Ocular tubercle with 2 anterior +1 posterior ocelli
2014.08.23 © Hara, Y.

Species malabarica (Yosii, 1966) Ohira A & Nakamori T in Ohira, A, Nakamori, T, Matsumoto, S, Bessho-Uehara, M, Kato, T & Oba, Y, 2023:69

Species similis (Yosii, R, 1954:66) Salmon, 1964

Species uozumii (Yosii, 1956) Ohira A & Nakamori T in Ohira, A, Nakamori, T, Matsumoto, S, Bessho-Uehara, M, Kato, T & Oba, Y, 2023:69


Genus Hemilobella Deharveng, L et Greenslade, P, 1992:729

  Syn.: Neanuridae Lobelia(sic) Ohira, A, Nakamori, T, Matsumoto, S, Bessho-Uehara, M, Kato, T et Oba, Y, 2023:88 ISS


Species sp.1 Janssens F & Pearson S, 2018

Hemilobella sp.nov. from Australia
2016.10.31 © Steve & Alison.
Hemilobella sp.nov. from Australia
Ocelli 3+3
2016.10.31 © Steve & Alison.
Hemilobella sp.nov. from Australia
Pointed mouthcone
2016.10.31 © Steve & Alison.
Hemilobella sp.nov. from Australia
Ocelli 3+3
2018.02.28 © Steve & Alison.

Species daeana (Lee, B-H, 1982:219) Ohira A & Nakamori T in Ohira, A, Nakamori, T, Matsumoto, S, Bessho-Uehara, M, Kato, T & Oba, Y, 2023:68

Lobella daeana from Korea
After Lee B-H, 1982 Fig.1A

             Syn.: Lobella daenana(sic) Lee, B-H, 1982:222 ISS
            Syn.: Lobelia(sic) daeana Ohira, A, Nakamori, T, Matsumoto, S, Bessho-Uehara, M, Kato, T et Oba, Y, 2023:88


Species ferrumequinum Deharveng, Agolin & D'Haese, 2008

Species matildae Smolis & Greenslade, 2020

Hemilobella matildae from Australia
2020.07.04 © Steve & Alison.
Hemilobella matildae from Australia
2020.07.04 © Steve & Alison.
Hemilobella matildae from Australia
Pointed mouthcone
2020.07.04 © Steve & Alison.

Species newmani (Womersley, 1933) Deharveng, L & Greenslade, P, 1992:734

Species perakensis (Yosii, 1976) Deharveng, L & Greenslade, P, 1992:736

Species pluvia Deharveng, L et Greenslade, P, 1992:732

Species rounsevelli Deharveng, L et Greenslade, P, 1992:730, t.t.


Genus Hyperlobella Cassagnau, P, 1988:24

Species kraepelini (Börner, C, 1906:169) Cassagnau, P, 1988:24, t.t.

            Syn.: Neanura dubiosa Ritter, 1910
            Syn.: Neanura dubiosa Mandal, GP et Hazra, AK, 2009:13
            Syn.: Neanura pudibunda Imms, 1912
            Syn.: Neanura pudibunda Mandal, GP et Hazra, AK, 2009:13
            Syn.: Protanura spinifera Carpenter, 1917
            Syn.: Protanura spinifera Mandal, GP et Hazra, AK, 2009:13
            Syn.: Protanura sp. Goto, 1955 =?
            Syn.: Lobella cassagnaui Prabhoo, 1971


Lobella kraepelini from Singapore
After Yosii R, 1959 Fig.6A
Lobella kraepelini from India
After Mandal GP & Hazra AK, 2009 Fig.9a


Genus Paralobella Cassagnau, P et Deharveng, L, 1984:9 key  key  key

Paralobella sp. from China
Abd.5 integument with conical cuticular tubercles
2023.07.05 © Wang, G.
Paralobella sp. from China
Th.2 tubercle with stellate formation of cuticular tubercles
2023.07.05 © Wang, G.

Species apsala (Yosii, 1976) Jiang & al, 2012

Species breviseta Luo, Y-Z et Palacios-Vargas, JG, 2016:344

Paralobella breviseta from China
Modified after Luo Y-Z & Palacios-Vargas JG, 2016 Fig.1

Species erawan (Yosii, 1976) Jiang & al, 2012

Species khaochongensis (Yosii, 1976) Bedos, 1994

             Syn.: Lobella? (Lobellina) Khaochonaensis(sic) Deharveng, 1987 ISS

Species kinabaluensis (Yoshii, 1981) Jiang & al, 2012

Species kuchierabu Kasai, H, Hayasaka, D et Sawahata, T, 2022:?

Species orousseti Cassagnau, P et Deharveng, L, 1984:9, t.t.

             Syn.: Paralobella orousetti(sic) Hopkin, SP, 2002:1 ISS
             Syn.: Paralobella orousetii(sic) Hopkin, SP, 2002:1 ISS
Paralobella orousseti
After Deharveng, L., 2002 ©



Species palustris Jiang, Luan & Yin, 2012

Species paraperfusa (Gapud, 1968) Jiang & al, 2012

Species penangensis (Yosii, 1976) Jiang & al, 2012

Species perfusa (Denis, 1934) Deharveng, L & Bedos, A, 2000:210

Lobella perfusa from Taiwan
After Lee B-H & Park K-H, 1990 Fig.6A

Species sabahna (Yosii, 1981) Jiang & al, 2012

Species selangorica (Yosii, 1976) Jiang & al, 2012

Species tianmuna Jiang, J-G, Wang, W-B, Xia, H, 2018:8

Paralobella tianmanu from China
Modified after Jiang J-G, Wang W-B & Xia H, 2018 Fig.8-9


Genus Sulobella Deharveng & Suhardjono, 2000

Species yoshii Deharveng & Suhardjono, 2000, t.t.

Tribus Paleonurini Cassagnau, 1989 key

Genus Itanura Queiroz, GC et Deharveng, L, 2015:140

Itanura sp.
Schematic of tubercles on head and abd.4-6
After Queiroz GC et Deharveng L, 2015 Fig.16
  Syn.: Paleonurini Itanaura(sic) Palacios-Vargas, JG et Soto-Adames, FN, 2017:388 ISS


Species brasiliensis (Arlé, R, 1960:164) Queiroz, GC & Deharveng, L, 2015:142

Itanura brasiliensis from Brazil
From type locality
After Queiroz GC et Deharveng L, 2015 Fig.17
Itanura brasiliensis from Brazil
From Rio de Janeiro: Itatiatia
After Queiroz GC et Deharveng L, 2015 Fig.22
Itanura brasiliensis from Brazil
From Rio de Janeiro: Serra dos Órgãos
After Queiroz GC et Deharveng L, 2015 Fig.28
             Syn.: Itanura brasilensis(sic) Arango, A, Palacios-Vargas, JG, Vázquez, M et Marín, EP, 2018:150 ISS



Genus Graniloba Cassagnau, 2000

Species imitator Cassagnau, 2000, t.t.

Species verrucosa Cassagnau, 2000


Genus Parectonura Deharveng in Tillier, 1988

Species ciliata Deharveng in Tillier, 1988:46, t.t.


Genus Caledonura Deharveng in Tillier, 1988

Species tillierae Deharveng in Tillier, 1988:49


Genus Penelopella Cassagnau, 1986

  Syn.: Paleonurini Penellopella(sic) Smolis, A, 2000:412 ISS

Species pacifica Cassagnau, 1986

             Syn.: Penelopella pacyfica(sic) Weiner, WM, Najt, J et Pasnik, G, 2012:558 ISS

Species pohli (Barra, J-A, 2006:63) Weiner, WM, Najt, J & Pasnik, G, 2012:556

Penelopella pohli from Socotra Island
After Weiner WM, Najt J & Pasnik G, 2012 Fig.2A


Genus Siamanura Deharveng, L, 1987:2

Species bisetosa Deharveng, L, 1987:18

Species breviseta Deharveng, L, 1987:15

Species burkhardti Deharveng, L, 1987:24

Species clavata Deharveng, L, 1987:15

Species gouzei Deharveng, L, 1987:20

Species hanuman (Yoshii, 1976)

Species inthanonensis Deharveng, L, 1987:6

Species leksawasdii Deharveng, L, 1987:11

Species lisu Deharveng, L, 1987:20

Species loebli Deharveng, L, 1987:27

            Syn.: Siamanura löbli(sic) Deharveng, L, 1987:27 IOS

Species maffrigali Deharveng, L, 1987:14

Species media Deharveng, L, 1987:6

Species phyllioseta Deharveng, L, 1987:23

Species primadinae Suhardjono, YR et Deharveng, L, 1992:61

             Syn.: Siamanura primadina(sic) Suhardjono, YR et Deharveng, L, 1992:61 ISS

Species trisetosa Deharveng, L, 1987:9


Genus Chirolavia Deharveng, L, 1991:54

Species gabaudei Deharveng, L, 1991:55

Species murphyi Deharveng, L, 1991:54, t.t.


Genus Pronura Delamare Deboutteville, C, 1953:823

Species sp.1 Janssens F sensu Schneider & Deharveng, 2017

            Syn.: Pronura sp.new Schneider, C et Deharveng, L, 2017:2

Species sp.2 Janssens F sensu Ospina-Sánchez, 2019

            Syn.: Pronura sp.nov. Ospina-Sánchez, CM, Soto-Adames, FN et González, G, 2020:8

Species aciculifer Cassagnau, 1996

Species aequalis Cassagnau, 1991

Species alata Cassagnau, 1996

Species amazonica Cassagnau & Oliveira, 1990

Species anjavicola Cassagnau, 1996

Species annapurnensis Cassagnau, 1991

Species australica (Yosii, 1966) Greenslade, P & Deharveng, L, 1991:852

            Syn.: Neanura cirrata Yosii, 1966 =?
            Syn.: Achorutes rosaceus Womersley, 1933 =:


Species bidoup Deharveng, L et Smolis, A, 2002:2

Species bipartita Cassagnau, 1991

Species bouakensis Cassagnau, 1996

Species caeca Cassagnau, 1996

Species camerounensis Cassagnau, 1996

Species capitata Cassagnau, 1991

Species celisi Cassagnau, 1996

Species clavisetis (Yoshii, 1981)

Species curviseta Cassagnau, 1996

Species dandicola Cassagnau, 1991

Species diplochaeta Cassagnau, 1991

Species dorsolateralis Deharveng, L et Bedos, A, 1993:191

Species edaphica Cassagnau, 1991

Species elegans Cassagnau, 1996

Species erythrina Cassagnau, 1991

Species fluctuans Cassagnau, 1996

             Syn.: Pronura fluctans(sic) Smolis, A, 2000:412 ISS

Species gaucheri Palacios-Vargas, JG, Deharveng, L et D'Haese, CA, 2011:198

Pronura gaucheri from French Guyana
After Palacios-Vargas JG et al, 2011 Fig.1
2009 © D'Haese, C.

Species gosainkundi Cassagnau, 1991

Species indiana Salmon, 1969

Species kilimanjarica Delamare Deboutteville, C, 1953:823, t.t.

Pronura kilimanjarica from Tanzania
Modified after Delamare Deboutteville C, 1953 Fig.4b
Pronura kilimanjarica from Tanzania
Modified after Delamare Deboutteville C, 1953 Fig.4a

             Syn.: Pronura kilimandjarica(sic) Massoud, Z, 1967:336 ISS
             Syn.: Pronura kilimandjarica(sic) Cassagnau, P, 1983:16 ISS
             Syn.: Pronura kilimandjarica(sic) Palacios-Vargas, JG, Deharveng, L et D'Haese, CA, 2011:198 ISS


Species kyandolirensis Cassagnau, 1996

Species laminata Cassagnau, 1996

Species lemurica Cassagnau, 1996

Species longiseta Cassagnau, 1991

Species madagascariensis Cassagnau, 1996

Species major Cassagnau, 1996

Species mignardi Massoud, 1963

Species nana Cassagnau, 1996

Species nepalica Cassagnau, 1991

Species ornata Deharveng, L et Bedos, A, 1993:190

Species paraguayana Palacios-Vargas, JG, Deharveng, L et D'Haese, CA, 2011:202

Species parvati Cassagnau, 1991

Species plumosa Cassagnau, 1991

Species pomorskii Smolis & Deharveng, 2006

Species prima Cassagnau, 1996

Species prodoni Cassagnau, 1991

Species pumilio Cassagnau, 1996

Species regularis Cassagnau, 1996

Species reticulata Cassagnau, 1991

Species ruwenzoriensis Cassagnau, 1996

Species seticola Cassagnau, 1991

Species setimigrans Cassagnau, 1996

Species shimicola Cassagnau, 1991

Species sphaeroculata Cassagnau, 1996

Species straturata Cassagnau, 1996

Species travei Cassagnau, 1991

Species trisetosa Cassagnau, 1996

Species trisulensis Cassagnau, 1991

Species yunquensis Ospina-Sánchez, CM, Palacios-Vargas, JG et González, G, 2021:582

Pronura yunquensis from Puerto Rico
After Ospina-Sánchez CM, Palacios-Vargas JG & González G, 2021 Fig.1
             Syn.: Pronura yunquesis(sic) Ospina-Sánchez, CM, Palacios-Vargas, JG et González, G, 2021:588 ISS



Genus Galanura Smolis, A, 2000:412

Species agnieszkae Smolis, A, 2000:414, t.t.

            Syn.: Galanura agnieskae(sic) Smolis, A, 2000:411 IOS
            Syn.: Galanura agnieskae(sic) Smolis, A, 2000:412 IOS
            Syn.: Galanura agnieskae(sic) Smolis, A, 2000:414 IOS
            Syn.: Galanura agnieskae(sic) Smolis, A, 2000:415 IOS
            Syn.: Galanura agnieskae(sic) Smolis & Kadej, 2014 IOS
            Syn.: Galanura agnieskae(sic) Smolis, A et Pasnik, G, 2020:9 IOS
Remark by Janssens, F. 2020.11.14: According to the etymology (p.415) : "The new species is dedicated to my wife Agnieszka." However, first the new name refers to Agnieska, not to Agnieszka. And second the author used the name 'agnieszka' in the figs 1-14. This mismatch is clear evidence of an inadvertent error that must be corrected (Art.32.5.1). The name 'agnieszkae' is in line with the etymology.



Genus Paleonura Cassagnau, P, 1982:6 key

Species sp.1 Najt & Thibaud, 1987

Species sp.2 Díaz Azpiazu, M et Palacios-Vargas, JG, 1999:93

Species aberdarensis Cassagnau, 1996

Species africana Cassagnau, 1996

Species angustior (Rusek, 1971) Cassagnau, P, 1982:6

Species anophthalma Cassagnau, 1991

Species anosyennica Cassagnau, 1996

Species badaga Cassagnau, P, 1988:28

Species barbata Cassagnau, P, 1988:30

Species bilinskii Pasnik & Weiner, 2013

Species borincana Palacios-Vargas, JG et Soto-Adames, FN, 2017:389

             Syn.: Paleonura boricana(sic) Palacios-Vargas, JG et Soto-Adames, FN, 2017:392 ISS
             Syn.: Paleonura boricana(sic) Arango, A, Palacios-Vargas, JG, Vázquez, M et Marín, EP, 2018:150 ISS

Species carayoni Massoud & Thibaud, 1987

Species cassagnaui Weiner, WM et Najt, J, 1998:229

Species coalescens Cassagnau, 1996

Species colimana Palacios-Vargas, JG et Gómez-Anaya, JA, 1995:96

            Syn.: Deutonura sp. Palacios-Vargas, JG et Gómez-Anaya, JA, 1995:101


Species daniae Palacios-Vargas, JG et Díaz, M, 1992:158

Species decorata Cassagnau, P, 1988:31

Species deharvengi Cassagnau, 1991

Species dejeani Cassagnau, 1996

Species dilatata Deharveng, L et Bedos, A, 1993:187

Species epiphytica Smolis, A et Deharveng, L, 2003:320

Species formosana (Yosii, R, 1965:9) Deharveng, L, 1989:104

Paranura formosana from Taiwan
After Yosii R, 1965 Fig.6A

Species ganesh Cassagnau, 1991

Species guadalcanarae (Yosii, 1960) Deharveng & Greenslade P, 1992:738

Species guae (Yoshii, 1976) Cassagnau, P, 1982:6

Species gurung Cassagnau, 1991

Species ili (Christiansen, K & Bellinger, P, 1992:88) Palacios-Vargas, JG & Soto-Adames, FN, 2017:393

Species indrabahadouri Cassagnau, 1991

Species insularum (Carpenter, 1935) Cassagnau, P, 1982:6

Species irregularis Cassagnau, 1991

Species khumbica (Cassagnau, 1971) Cassagnau, P, 1982:6

Species lamjungensis Cassagnau, 1991

Species lanna Deharveng, L et Bedos, A, 1993:184

Species loebli Cassagnau, P, 1988:27

Species lonavlana (Yosii, 1966) Cassagnau, P, 1982:6

Species louisi Thibaud & Najt, 1993

Species macronychia Cassagnau, P, 1988:26

Species miniseta Massoud & Thibaud, 1987

             Syn.: Paleonura minisetae(sic) Palacios-Vargas & Díaz, 1992 ISS
             Syn.: Paleonura minisetae(sic) Palacios-Vargas & Gómez-Anaya, 1995 ISS

Species monochaeta Deharveng, L et Bedos, A, 1993:185

Species monophthalma (Yosii, 1966) Cassagnau, P, 1982:6

Species nuda Cassagnau & Oliveira, 1990

Species paurochaetosa Cassagnau, 1991

Species pescadoria Palacios-Vargas, JG et Gómez-Anaya, JA, 1995:95

            Syn.: Paleonura pescadorius(sic) Palacios-Vargas, JG et Gómez-Anaya, JA, 1995:95 IOS
            Syn.: Paleonura pescadorius(sic) Palacios-Vargas, JG et Gómez-Anaya, JA, 1995:8 nn IOS
            Syn.: Paleonura pescadorius(sic) Palacios-Vargas & Gómez-Anaya, 1995 IOS
            Syn.: Paleonura pescadorius(sic) Mari Mutt, JA et Bellinger, PF, 1996:168 IOS
            Syn.: Paleonura pescadorius(sic) Palacios-Vargas, JG, 2000:28 IOS
            Syn.: Paleonura? pescadorius(sic) Palacios-Vargas, JG et Soto-Adames, FN, 2017:388 IOS
            Syn.: Pronura? pescadorius(sic) Palacios-Vargas, JG et Soto-Adames, FN, 2017:388 IOS
            Syn.: Paleonura pescadorius(sic) Ospina-Sánchez, CM, Palacios-Vargas, JG et González, G, 2021:588 IOS

Species petebellingeri Palacios-Vargas, JG et Simón-Benito, JC, 2007:320

Species plumosa Cassagnau, 1996

Species reducta (Yosii, 1966) Cassagnau, P, 1982:6

Species rosacea (Schött, H, 1917:7) Deharveng, L & Greenslade, P, 1992:738

Achorutes rosaceus from Australia
Terminal body tubercles
After Schött H, 1917 Fig.7
Achorutes rosaceus from Australia
Ocular tubercle with 2 unpigmented ocelli
After Schött H, 1917 Fig.8
             Syn.: Achorutes roseus(sic) Sarawak lapsus ISS



Species rosacea2 (Womersley, 1933) Deharveng, L & Greenslade, P, 1992:738

Achorutes rosaceus from Australia
Apex of abdomen
After Womersley H, 1939 Fig.48A
Achorutes rosaceus from Australia
Ocelli
After Womersley H, 1939 Fig.48B
Paleonura nr rosacea from Australia
2016.08.11 © Ball, D.
Paleonura nr rosacea from Australia
2016.08.11 © Ball, D.
Remark by Greenslade, P. 2018.02.22: Womersley's colour description was not based on Malanda specimens but only on those he picked up in southern parts of the continent so are quite likely to be different species.
Paleonura nr rosacea from Australia
1+2+3+6 cephalic tubercles
2018.02.26 © Steve & Alison.
Paleonura nr rosacea from Australia
1+2+3+6 cephalic tubercles
2018.02.28 © Steve & Alison.
Paleonura nr rosacea from Australia
2020.08.17 © Hyde, G.

Species saproxylica Smolis & Kadej, 2014

Species setikholensis Cassagnau, 1991

Species siva (Yosii, 1966) Cassagnau, P, 1982:6

Species spectabilis Cassagnau, P, 1982:7, t.t.

Species trisetosa Cassagnau, 1991


Genus Speleonura Palacios-Vargas, JG et Simón-Benito, JC, 2007:318

Species kenchristianseni Palacios-Vargas, JG et Simón-Benito, JC, 2007:318, t.t.


Genus Ectonura Cassagnau, 1980

Ectonura sp.
Schematic of tubercles on head and abd.4-6
After Queiroz GC et Deharveng L, 2015 Fig.15

Species sp.1 Greenslade & Deharveng, 1990

Species sp.2 da Rocha Neves, AC, de Mendonça, MC et Queiroz, GC, 2019:6

Species araucariae (Yosii, 1960) Deharveng & Bedos, 2002

Species barrai Janion, C, Bedos, A et Deharveng, L, 2011:39

Species coatesi Barra, J-A, 1994:186

Remark by Janion, C., Bedos, A. & Deharveng, L. 2011:34: The only differential character separating unambiguously E. coatesi from E. oribiensis is the morphology of modified chaetae Ag in the male.

Species judithae Deharveng & Bedos, 2002

Species koumac Deharveng & Bedos, 2002

Species lata Deharveng, Weiner & Najt, 1997

Species longirostris Deharveng, Weiner & Najt, 1997

Species minima Deharveng, Weiner & Najt, 1997

Species monochaeta Janion, C, Bedos, A et Deharveng, L, 2011:34

Species natalensis (Womersley, H, 1934:451) Cassagnau, 1980, t.t.

            Syn.: Neanura montana Paclt, 1959


Species oribiensis (Coates, 1968) Cassagnau, 1980

Species paralata Deharveng, Weiner & Najt, 1997

Species proxima Deharveng, Weiner & Najt, 1997

Species pulchra Deharveng, Weiner & Najt, 1997

Species sensillata Deharveng & Bedos, 2002

Species snowdeni Queiroz, GC et Deharveng, L, 2015:136

Ectonura snowdeni from Brazil
After Queiroz GC et Deharveng L, 2015 Fig.2

Species sylvatica Deharveng, Weiner & Najt, 1997


Genus Vitronura Yosii, 1969 key

  Syn.: Podurellen(sic) Achorutes Elditt, HL, 1854:41 partim
  Syn.: Neanurinae Matanura(sic) Yosii, R, 1977:149 ISS
  Syn.: Neanuridae Witronura(sic) Liu, Y-Q, Hou, D-B et Li, Z-C, 1998:126 ISS


Vitronura sp.
Schematic of tubercles on head and abd.4-6
After Queiroz GC et Deharveng L, 2015 Fig.12
Vitronura sp. from China
3D reproduction by Bu, Y.
2018.08.06 © Palacios-Vargas, J.G.
Vitronura sp. from China
3D reproduction by Bu, Y.
2018.08.06 © Palacios-Vargas, J.G.
Vitronura sp. from China
3D reproduction by Bu, Y.
2018.08.06 © Palacios-Vargas, J.G.
Vitronura sp. from Mexico
After Murray A in Palacios-Vargas JG & Cruz-Leal JI, 2020 Fig.2

Species sp.1 Janssens F sensu Yahyapour E, Shayanmehr M, Vafaei-Shoushtari R & Arbea JI, 2022

            Syn.: Vitronura sp.undescribed Yahyapour, E, Shayanmehr, M, Vafaei-Shoushtari, R et Arbea, JI, 2022:265


Species acuta Deharveng, L et Weiner, WM, 1984:52

Species ciliata Wang, Y-R, Wang, W-B et Jiang, J-G, 2016:187

Vitronura ciliata from China
After Wang Y-R, Wang W-B & Jiang J-G, 2016 Fig.8

Species dentata Deharveng, L et Weiner, WM, 1984:54

Species giselae (Gisin, 1950) Cassagnau, 1980

            Syn.: Anoura rosea Gervais, 1842
Anoura rosea from France
After Nicolet H, 1847 Pl.6 fig.16

             Syn.: Anura Rosea(sic) Kolenati, FA, 1858:131 ISS
Vitronura rosea from Switzerland
2008.03.17 © Haentjens, R.
Remark by Janssens, F. 2015.10.14: Given Yoshii (1995) synonymised rosea Gervais with giselae Gisin and mandarina Yosii, according to ICZN rules of priority, rosea Gervais, 1842 takes priority on giselae Gisin, 1950 and mandarina Yosii, 1954. However, this is not yet reflected in the current taxonomic hierarchy due to taxonomic complications with the type species mandarina.





Vitronura giselae
Cephalic chaetotaxy
After Deharveng L, 1983 Fig.2B
Remark by Deharveng, L. & Bedos, A. 2000:210: Identification of the Vietnamese material has to be confirmed.

Vitronura giselae from the USA
2008.11.08 © Justis, S.
Vitronura giselae from the USA
2008.11.08 © Justis, S.
Vitronura giselae from the USA
2008.11.08 © Justis, S.
Vitronura giselae from the USA
2008.11.27 © Justis, S.
Vitronura giselae from the USA
2009.02.20 © Houston, S.
Vitronura giselae from the USA
2009.10.25 © Bergen, L.
Vitronura giselae from the USA
2009.10.25 © Bergen, L.
Vitronura giselae from the USA
2010.06.18 © Montgomery, G.
Vitronura giselae from the USA
2011.12.24 © Fleming, K.
Vitronura giselae from the USA
2012.01.19 © Kouri, J.
Vitronura giselae from the USA
2012.03.26 © Bertone, M.
Vitronura giselae from the USA
2015.03.31 © Montgomery, G.
Vitronura giselae from the USA
Dorsal head tubercles: 1+2+3+6
2015.07.29 © Wechsler, D.
Vitronura giselae from Malaysia
2015.12.07 © Krásenský, P.
Vitronura giselae from France
Dorsal head tubercles: 1+2+3+6
2016.02.08 © Garcelon, P.
Vitronura giselae from France
Dorsal head tubercles: 1+2+3+6
2016.02.11 © Garcelon, P.
Vitronura giselae? from Australia
2018.02.06 © Steve & Alison.
Vitronura giselae from France
Dorsal head tubercles: 1+2+3+6
2018.02.07 © Garcelon, P.
Vitronura giselae from France
Pointed mouthcone
2018.02.07 © Garcelon, P.
Vitronura giselae from France
Abdominal tubercles
2018.02.07 © Garcelon, P.
Vitronura giselae from the USA
Cephalic tubercles : 1+2+3+6
2022.01.12 © Charny, V.
Vitronura giselae from the USA
2022.03.10 © Coogler, J.
Vitronura giselae from the USA
Unguiculus absent
2022.03.10 © Coogler, J.
Vitronura giselae from the USA
Cephalic dorsal tubercles : 1+2+3+6
Prothoracic dorsal inner tubercles absent
2022.04.08 © Dankowicz, Z.
Vitronura giselae from Japan
After Ohira A & al, 2023 Fig.24 bottom left
Vitronura giselae from Japan
Bioluminescence in dim light after stimulation
After Ohira A & al, 2023 Fig.24 bottom middle
Vitronura giselae from Japan
Bioluminescence in dark after stimulation
After Ohira A & al, 2023 Fig.24 bottom right
Vitronura giselae hatchling from the USA
2023.06.21 © Gruver, Be.
Vitronura giselae hatchling from the USA
With eggs
2023.06.21 © Gruver, Be.
Vitronura giselae from the USA
Feeding on plasmodium of Physarum polycephalum
2023.11.25 © Gruver, Be.
Vitronura giselae juv. from the USA
2024.01.13 © Coogler, J.

  Subspecies citrina Latzel, 1922

             Syn.: Achorutes roseus-citrinus(sic) Handschin, E in Dahl, F, 1929:42,si ISS

Species gressitti Cassagnau, P et Deharveng, L, 1981:161

Species joanna (Coates, 1968) Greenslade, P, 1997-:www0312

Species kunigamiensis Tanaka & Hasegawa, 2010

Species latior (Rusek, J, 1967:189)

Species luzonica Yosii, 1976

Species macgillivrayi (Denis, J-R, 1933:225) Cassagnau & Palacios-Vargas, 1983

            Syn.: Achorutes Mac Gillivrayi(sic) Denis, J-R, 1933:225 IOS
Achorutes Mac Gillivrayi(sic) from Costa Rica
After Denis J-R, 1933 Fig.7


            Syn.: Biloba costaricae Stach, J, 1951:29
Remark by Palacios-Vargas, J.G. 2010.08.09: I was in Costa Rica collecting Neanurids and found some Vitronura specimens. I realized that what was described by Denis (1933) as Achorutes macgillivrayi and that described by Stach (1951) as Biloba costaricae, are the same species.

            Syn.: Vitronura giselae Cassagnau & Palacios-Vargas, 1983 =?




            Syn.: Achorutes Mac Gillivrayi(sic) Denis, J-R, 1933:230 IOS
             Syn.: Achorutes Macgillivrayi(sic) Denis, J-R, 1933:315 ISS
             Syn.: Achorutes Macgillivrayi(sic) Denis, J-R, 1933:318 ISS



Species mandarina (Yosii, 1954) Yosii, 1976, t.t.

            Syn.: Metanura pygmaea Yosii, R, 1956:48 =?
            Syn.: Vitronura rosea Yoshii, 1995


Neanura (Propeanura) mandarina from Japan
After Yosii R, 1956 Pl.XXIV Fig.139

Species mascula Smolis & Deharveng, 2006

Species namhaeiensis (Lee, B-H, 1974:582) Cassagnau & Deharveng, 1981

Neanura namhaeiensis from Korea
After Lee B-H, 1974 Fig.7C

Species paraacuta Wang, Y-R, Wang, W-B et Jiang, J-G, 2016:184

Vitronura paraacuta from China
After Wang Y-R, Wang W-B & Jiang J-G, 2016 Fig.1

Species pygmaea (Yosii, 1954) Yosii, 1969

Species qingchengensis Jiang & Yin, 2011

Species quartadecima Gao, Y, Bu, Y et Palacios-Vargas, JG, 2012:1147

Vitronura quartadecima from China
After Gao Y, Bu Y & Palacios-Vargas JG, 2012 Fig.15

Species semilunaris (Schött, 1925) Ohira, A, Nakamori, T, Matsumoto, S, Bessho-Uehara, M, Kato, T & Oba, Y, 2023:68

            Syn.: Vitronura? semilunaris Ohira, A, Nakamori, T, Matsumoto, S, Bessho-Uehara, M, Kato, T et Oba, Y, 2023:68

Species setaebarbulata Gao, Y, Bu, Y et Palacios-Vargas, JG, 2012:1143

Vitronura setaebarbulata from China
After Gao Y, Bu Y & Palacios-Vargas JG, 2012 Fig.2

Species shaanxiensis Jiang & Yin, 2011

Species singaporiensis (Yosii, R, 1959:14) Yosii, 1976

Bilobella singaporiensis from Singapore
After Yosii R, 1959 Fig.8A

Species sinica Yosii, 1976

Species tubercula Lee, B-H et Kim, J-T, 1990:244

Vitronura tubercula from Taiwan
After Lee B-H & Park K-H, 1990 Fig.7A


Genus Blasconura Cassagnau, P, 1983:19 key

  Syn.: ... Biocularia Stach, 1951 partim


Blasconura sp.
Schematic of tubercles on head and abd.4-6
After Queiroz GC et Deharveng L, 2015 Fig.11
Blasconura sp. from Japan
After Kasai H & al, 2022 cover

Species anamalensis Cassagnau, P, 1988:37

Species batai Bedos & Deharveng, 2000

Species ceylonica Cassagnau, P, 1988:35

Species cordis Kasai, H, Hayasaka, D et Sawahata, T, 2022:?

Species hirtella (Börner, C, 1906:170) Cassagnau, P, 1983:16

            Syn.: Achorutes cirratus Handschin, E, 1926:453 =?
            Syn.: Neanura hirtella var. schötti(sic) Womersley, 1935


Lobella (Propeanura) hirtella
from Singapore
After Yosii R, 1959 Fig.7A

Remark by Deharveng, L. & Bedos, A. 2000:210: Blasconura hirtella may correspond to a complex of closely related species. The identification of the Vietnamese specimens has to be checked.

Blasconura hirtella from Singapore
2008.03.29 © Ng, M.
Blasconura hirtella from Singapore
2008.07.21 © Ng, M.
Blasconura hirtella from Singapore
2008.07.21 © Ng, M.
Blasconura hirtella from Singapore
2011.06.11 © Koh, J.
Blasconura hirtella from Singapore
2011.07.16 © Koh, J.
Blasconura hirtella from Singapore
2012.10.27 © Bay, N.
Blasconura hirtella from Singapore
2015.11.27 © Bay, N.
Blasconura hirtella from Singapore
2015.11.27 © Bay, N.

  Subspecies novaezealandiae Salmon, JT, 1941:307

            Syn.: Neanura hirtella ssp. novae-zealandiae(sic) Salmon, JT, 1941:307 IOS
             Syn.: Neanura hirtella var. nova-zelandiae(sic) Massoud, Z, 1967:318,si ISS
Blasconura hirtella ssp. novaezealandiae juv. from New Zealand
2014.02.09 © Murray, A.
Blasconura hirtella ssp. novaezealandiae from New Zealand
2014.02.13 © Murray, A.
Blasconura hirtella ssp. novaezealandiae from New Zealand
2014.02.13 © Murray, A.
Blasconura hirtella ssp. novaezealandiae from New Zealand
2014.02.13 © Murray, A.





Species palniensis Cassagnau, P, 1988:36

Species prabhooi Cassagnau, P, 1988:34

Species separata (Denis, 1934) Cassagnau, 1988

Remark by Cassagnau, 1988 in Deharveng, L. & Bedos, A. 2000:210: The attribution to the genus Blasconura is uncertain.

Species sholica Cassagnau, P, 1988:33

Species toda Cassagnau, P, 1988:32

Species triangulum Kasai, H, Hayasaka, D et Sawahata, T, 2022:?

Species vivianae Palacios-Vargas & Díaz, 1992:162


Genus Australonura Cassagnau in Dallai, 1980

Australonura from New Zealand
2006 Minor, M. & Robertson, A. © SoilBugs
Det. Deharveng, L., 2008.12.22
Australonura sp. from New Zealand
2014.03.02 © Murray, A.
Australonura sp. from New Zealand
2014.03.02 © Murray, A.
Australonura sp.
Schematic of tubercles on head and abd.4-6
After Queiroz GC et Deharveng L, 2015 Fig.13
Australonura sp. from Tasmania
2016.02.02 © Tony.
Australonura sp. from Tasmania
Head tubercles
2016.02.02 © Tony.
Australonura sp. from Tasmania
Head tubercles
2016.09.16 © Tony.
Australonura sp. from Tasmania
2016.09.16 © Tony.
Australonura sp. from Tasmania
Ocular tubercle with 2 ocelli
2016.09.16 © Tony.
Australonura? from Singapore
Dorsal cephalic tubercles : 3+3+1
2021.01.23 © Yeo, C.
Leg.: Luna, J.

Species sp.1 Palacios-Vargas, JG et Deharveng, L, 2014:42

Species ahimsa Greenslade P & Deharveng, 1990:572

Species beata Greenslade P & Deharveng, 1990:574

Species cirrata (Schött, H, 1917:8) Greenslade P & Deharveng, 1990:575

            Syn.: Achorutes hirtellus ssp. noname Denis, 1933
            Syn.: Achorutes hirtellus Womersley, 1933


             Syn.: Neanura hirtellus(sic) var. cirratus(sic) Womersley, H, 1942:24 ISS

Species friasica (Cassagnau & Oliveira, 1990) Palacios-Vargas, JG & Deharveng, L, 2014:40

            Syn.: Neanura limnophila Cassagnau & Rapoport, 1962 partim


Species gili Queiroz & Deharveng, 2014

Species grandes Greenslade P & Deharveng, 1990:576

Species grossi (Yosii, 1966) Cassagnau, P, 1983:16, t.t.

Species lawrencei Deharveng & Greenslade P, 1990:598

Species limnophila (Cassagnau & Rapoport, 1962) Palacios-Vargas, JG & Deharveng, L, 2014:35

Australonura limnophila from Argentina
After Palacios-Vargas JG & Deharveng L, 2014 Fig.2

Species meridionalis (Stach, 1951, s.i.) King, Greenslade P & Hutchinson, 1985

             Syn.: Australonura meridonalis(sic) Greenslade P & Deharveng, 1990 ISS

Species neotropica Queiroz & Deharveng, 2014

Species norfolkensis Greenslade P & Deharveng, 1990:580

Species novaecaledoniae (Yosii, R, 1960:10) Deharveng, L in Tillier, S, 1988:46

            Syn.: Neanura novae-caledoniae(sic) Yosii, R, 1960:10 IOS
             Syn.: Neanura novacaledoniae(sic) Massoud, Z, 1967:307 ISS



Species paraguayensis Palacios-Vargas, JG et Deharveng, L, 2014:42

            Syn.: Paleonura sp. Palacios-Vargas & Mejía Recamier, 2009


Species popamanasiu Deharveng & Greenslade P, 1990:602

Species quarta Greenslade P & Deharveng, 1990:582

Species redita Greenslade P & Deharveng, 1990:584

Species rennellensis Deharveng & Greenslade P, 1990:602

Species sanguisugarum Greenslade P & Deharveng, 1990:584

Species scoparia Greenslade, P et Deharveng, L, 1990:586

Species wellingtonia (Womersley, H, 1936:477) Deharveng, L, 1983:22

Achorutes hirtellus var. wellingtonia from Tasmania
After Womersley H, 1936 Fig.2a


Genus Zelandanura Deharveng, L et Wise, KAJ, 1987:143

  Syn.: Paleonurini Zealandanura(sic) Greenslade, P et Deharveng, L, 1991:853 ISS
  Syn.: ... Zealandanura(sic) Early & Gilberg, 1993 ISS

Species bituberculata Deharveng, L et Wise, KAJ, 1987:144, t.t.

            Syn.: Neanura radiata Gressit & al, 1964
            Syn.: Neanura radiata Gressit, 1964 partim
            Syn.: Neanura radiata Wise, 1964
            Syn.: Crossodonthina radiata Deharveng & Wise, 1987



Genus Phradmon Greenslade P & Deharveng, 1991

Phradmon sp.
Schematic of tubercles on head and abd.4-6
After Queiroz GC et Deharveng L, 2015 Fig.14

Species australasiae (Womersley, H, 1935:208) Greenslade, P & Deharveng, L, 1991:841, t.t.

Paranura australasiae from Australia
After Womersley H, 1939 Fig.44AD
             Syn.: Paranura australasia(sic) Salmon, JT, 1964:283 ISS


Species floccosus Greenslade P & Deharveng, 1991:844

Species maralali Greenslade P & Deharveng, 1991:845

Species schotti (Womersley, H, 1935:210) Greenslade, P & Deharveng, L, 1991:847

             Syn.: Neanura hirtella ssp. schoetti(sic) Salmon, 1941 ISS
             Syn.: Paranura australasia(sic) ssp. schoetti(sic) Salmon, JT, 1964:283 nn ISS
             Syn.: Paranura hirtella ssp. schoetti(sic) Yosii, 1966 ISS
             Syn.: Australonura schoetti(sic) Greenslade, P et Deharveng, L, 1990:591 ISS
             Syn.: Phradmon schoetti(sic) Greenslade, P et Deharveng, L, 1991:847 ISS
             Syn.: Phradmon schoetii(sic) Greenslade, P et Deharveng, L, 1991:837 ISS
             Syn.: Phradmon schoetti(sic) Greenslade, P in Houston, WWK, 1994:36 ISS

Species tasmaniae Greenslade P & Deharveng, 1991:849

Species trisetosus Greenslade P & Deharveng, 1991:851


Genus Blasconurella Deharveng, L et Bedos, A, 1992:300 key

Species arcuata Deharveng, L et Bedos, A, 1992:307, t.t.

Species intermedia Deharveng, L et Bedos, A, 1992:305

Species lobata Deharveng, L et Bedos, A, 1992:306

Species palmata Deharveng, L et Bedos, A, 1992:309

Species quinquesetosa Deharveng, L et Bedos, A, 1992:302


Genus Paramanura Cassagnau, 1986

Species najtae Cassagnau, 1986

Species problematica Cassagnau, 1991


Genus Hazaranura Cassagnau, 1991

  Syn.: Paleonurini Hazanura(sic) Smolis, A, 2000:412 ISS

Species elongata Cassagnau, 1991


Genus Nepalanura Yosii, 1966

Species paranuroides Yosii, 1966

             Syn.: Nepalanura (Nepalanura) paranurodes(sic) Mandal, GP et Hazra, AK, 2009:2 ISS



Genus Parvatinura Cassagnau, P, 1982:7

Species colcheni Cassagnau, P, 1984:9

Species dobremezi Cassagnau, P, 1982:8, t.t.

Species loebli Cassagnau, P, 1984:8

Species numica Cassagnau, P, 1984:8

Species taounjensis Cassagnau, P, 1984:8


Genus Inameria Cassagnau, P, 1983:20

Species corallina (Imms, 1912) Cassagnau, P, 1983:16, t.t.

            Syn.: Lobella (Propeanura) subcorallina Prabhoo, 1971


Species kenyensis Cassagnau, P, 1984:9


Genus Nilgirella Cassagnau, P, 1983:20

Species amboinensis Yoshii, R et Suhardjno, YR, 1992:3, i.s.

Species indica (Handschin, E, 1929:235) Cassagnau, P, 1983:16, t.t.

Achorutes indicus from South-India
After Handschin E, 1929 Fig.7

Species longiseta Cassagnau, P, 1988:44

Species palniensis Cassagnau, P, 1988:43

Species piljainae Cassagnau, P, 1988:42


Genus Phylliomeria Delamare Deboutteville, 1948

Species africana Delamare Deboutteville, 1948, t.t.

Species bebourensis Cassagnau, P, 1984:27

Species ruwenzorensis Cassagnau, P, 1984:26

Species simplex Cassagnau, P, 1984:26


Genus Gnatholonche Börner, C, 1906:168

  Syn.: Achorutes Gnatolonche(sic)) Schött, H, 1917:8 ISS
  Syn.: Paleonurini Gnatolonche(sic) Cassagnau, P, 1991:223 ISS

Species angularis (Salmon, JT, 1944:140) Stach, J, 1951:93

Gnatholonche angularis from New Zealand
2009 © Te Papa Museum
Gnatholonche angularis from New Zealand
2014.mm.dd © Murray, A.
Gnatholonche angularis from New Zealand
2014.02.17 © Murray, A.
Gnatholonche angularis from New Zealand
2014.02.17 © Murray, A.
Gnatholonche angularis from New Zealand
2014.02.17 © Murray, A.

Species annapurnensis Cassagnau, P, 1984:10

Species anomala (Yosii, 1966) Cassagnau, P, 1984:4, s.i.

Species coineaui Cassagnau, P, 1984:11

Species dentata Cassagnau, 1993

Species dobatensis Cassagnau, P, 1984:10

Species durrieui Cassagnau, P, 1984:9

Species elongata Cassagnau, P, 1984:13

Species ensifer Cassagnau, 1993

Species impariseta Cassagnau, 1993

Species intermedia (Imms, 1912) Stach, J, 1951:94

Species jesti Cassagnau, P, 1984:14

Species lamjurica Cassagnau, 1993

Species limbu Cassagnau, P, 1984:13

Species lipaspis (Börner, C, 1906:170) Handschin, E, 1926:453, t.t.

Gnatholonche lipaspis from Java
After Handschin E, 1926 Fig.4a

Species mahabaratensis Cassagnau, P, 1984:10

Species marginata Cassagnau, P, 1984:12

Species montana (Handschin, 1929) Stach, 1951:94

Species polychaetosa Cassagnau, P, 1984:14

Species prodoni Cassagnau, P, 1984:12

Species reducta Cassagnau, P, 1984:11

Species sarkimani Cassagnau, 1993

Species sensilla Salmon, 1948

Species setensis Cassagnau, 1993

Species sherpa Cassagnau, 1993


Genus Synameria Cassagnau, P, 1983:20

Species intermedia Cassagnau, P, 1990:22

Species miranda (Yosii, 1966) Cassagnau, P, 1983:16, t.t.

Species orientalis Cassagnau, P, 1990:20

Synameria orientalis from Nepal
After Cassagnau P in Thibaud J-M & D'Haese C, 2010 Fig.32


Genus Nepalimeria Cassagnau, P, 1984:5

Species coccinea Cassagnau, P, 1984:26

Species dal Cassagnau, P, 1984:24, t.t.

Species ganash Cassagnau, 1993

Species heterochaeta Cassagnau, P, 1984:25

Species khorensis Cassagnau, P, 1984:25

Species lepchana (Yosii, 1966) Cassagnau, P, 1984:4, s.i.


Genus Elgonura Cassagnau, P, 1984:5

Species erinacea Cassagnau, P, 1984:28, t.t.


Genus Singalimeria Cassagnau, P, 1984:5

Species pachyderma Cassagnau, P, 1984:24, t.t.


Genus Himalmeria Cassagnau, P, 1984:4

Subgenus Himalmeria Cassagnau, P, 1984:4 ss
Species gurung Cassagnau, P, 1984:15, t.t.

Species karmapa Cassagnau, P, 1984:16

Species ornata Cassagnau, P, 1984:16

Subgenus Yetimeria Cassagnau, P, 1984:4
Species armata Cassagnau, P, 1984:18

Species arunensis Cassagnau, P, 1984:21

Species cervicornis Cassagnau, P, 1984:21

Species cuneata Cassagnau, 1993

Species digitata Cassagnau, P, 1984:17

Species elegans Cassagnau, P, 1984:23

Species flabellifera Cassagnau, P, 1984:20

Species foliata Cassagnau, 1993

Species himalayana (Yosii, 1966) Cassagnau, P, 1984:4, s.i.

Species jaljalensis Cassagnau, P, 1984:19

Species lama Cassagnau, P, 1984:18, t.t.

Species lanata Cassagnau, P, 1984:19

Species musifera Cassagnau, P, 1984:20

Species rostrata Cassagnau, P, 1984:19

Species sikkimensis Cassagnau, P, 1984:18

Species smetanai Cassagnau, P, 1984:22

Species spatulata Cassagnau, P, 1984:20


Genus Calvinura Cassagnau, P, 1988:39

Species besucheti Cassagnau, P, 1988:39, t.t.

Species reducta Cassagnau, P, 1988:40


Genus Camerounura Cassagnau, P, 1991:221

Species delamarei Cassagnau, P, 1991:222, t.t.


Genus Digitanura Deharveng, L, 1987:145

Species quadrilobata Deharveng, L, 1987:146, t.t.

Digitanura quadrilobata from Thailand
2010.08.15 © Damrongsiri, S.


Genus Rambutanura Deharveng, 1988

Rambutanura sp. from Malaysia
2015.12.07 © Krásenský, P.
Rambutanura sp. from China
2023.06.24 © Wang, G.
Det.: Yu, D.-Y.
Rambutanura sp. from China
2023.06.24 © Wang, G.
Det.: Yu, D.-Y.

Species carcharia Smolis, A, 2007:?

Species dawydoffi (Denis, 1934) Deharveng, 1988

             Syn.: Rambutanura dawidoffi(sic) Nguyen, ATT, Thuy, KN et Arbea, JI, 2022:152 ISS

Species hunanensis Jiang & Dong, 2018

Species malayana (Yosii, 1976)

Species yoshiiana Deharveng, 1988


Genus Stenomeria Cassagnau, P, 1990:29

Species deharvengi Cassagnau, P, 1990:29, t.t.


Genus Tamulmeria Cassagnau, P, 1988:46

Species callipygos Cassagnau, P, 1988:46, t.t.

Species keralensis Cassagnau, P, 1988:47


References