041.0000
Family
Polypodiaceae J. & C.Presl; J. & C. Presl, Delic. Prag. 159 (1822)
67 genera, ca. 1700 spp. Phylogeny: Wei & Zhang 2022
Species listed in database: 1718; hybrids: 47
= Platyceriaceae Ching; Acta Phytotax. Sin. 16(3): 18 (1978)
= Drynariaceae Ching; Acta Phytotax. Sin. 16: 1-37 (1978)
= Grammitidaceae Newman; Hist. Brit. Ferns 7 (1840) (Grammitaceae auct., missp.)
= Gymnogrammitidaceae Ching; Acta Phytotax. Sin. 11: 12 (1966)
= Loxogrammaceae Ching ex Pic.Serm.; Webbia 29: 11 (1975)
= Platyceriaceae Ching; Sunyatsenia 5 (1940)
= Pleurosoriopsidaceae Sa.Kurata & Ikebe ex Ching; Acta Phytotax. Sin. 16(4): 17 (1978)
041.9025
Genus
Tomophyllum (E.Fourn.) Parris; Gard. Bull. Singapore 58(2): 245 (2007)
Distribution: Trop. Asia to China, Australia, W-Pacific. Checklist revised for this database by Parris (2020)
Species listed in database: 37
species :
Tomophyllum beleense
(Copel.) Parris; Gard. Bull. Singapore 58(2): 246 (2007)
Distribution: New Guinea (Irian Jaya, Papua New Guinea)
42 NWG
Synonyms: hidden
= Ctenopteris beleensis Copel.; Univ. Calif. Publ. Bot. 18: 225 (1942)
= Grammitis beleensis (Copel.) Christenh.; Global Fl. 1(4): 41 (2018)
species :
Tomophyllum bipinnatifidum
(Baker) Parris; Gard. Bull. Singapore 58(2): 246 (2007)
Distribution: Sulawesi; Moluccas (Seram); New Guinea (Papua New Guinea); Bismarck Arch. (New Ireland)
42 MOL NWG SUL 43 BIS
Synonyms: hidden
= Ctenopteris bipinnatifida (Baker) Copel.; Univ. Calif. Publ. Bot. 18: 225 (1942)
= Grammitis bipinnatifida (Baker) Christenh.; Global Fl. 1(4): 41 (2018)
= Polypodium bipinnatifidum Baker; J. Bot. 28: 109 (1890)
species :
Tomophyllum bipinnatum
(Copel.) Parris; Gard. Bull. Singapore 58(2): 246 (2007)
Distribution: New Guinea (Irian Jaya)
42 NWG
Synonyms: hidden
= Ctenopteris bipinnata Copel.; Univ. Calif. Publ. Bot. 18: 225 (1942)
= Grammitis calliope Christenh.; Global Fl. 1(4): 42 (2018), nom. nov.
species :
Tomophyllum brachyphlebium
(Baker) Parris; Gard. Bull. Singapore 58(2): 246 (2007)
Distribution: Sumatra
42 SUM
Synonyms: hidden
= Grammitis brachyphlebia (Baker) Christenh.; Global Fl. 1(4): 41 (2018)
= Polypodium brachyphlebium Baker; Ann. Bot. (Oxford) 8: 129 (1894)
species :
Tomophyllum brooksiae
(Alderw.) Parris; Gard. Bull. Singapore 58(2): 246 (2007)
Distribution: Sumatra; Borneo (Sarawak)
42 BOR SUM
Synonyms: hidden
= Ctenopteris brooksiae (Alderw.) Parris; Fern Gaz. 12(3): 180 (1981)
= Grammitis brooksiae (Alderw.) Christenh.; Global Fl. 1(4): 42 (2018)
= Polypodium brooksiae Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 23: 19, t. 2, f. 2 (1916)
species :
Tomophyllum callophyllum
(C.H.Wright) Parris; Fl. Penins. Malaysia Ser. 1, 1: 196 (2010)
Distribution: peninsular Malaysia (Perak, Penang, Pahang); New Guinea (Papua New Guinea)
42 MLY NWG
Synonyms: hidden
= Grammitis callophylla (C.H.Wright) Christenh.; Global Fl. 1(4): 42 (2018)
= Grammitis capillata (Brause) Christenh.; Global Fl. 1(4): 42 (2018)
= Polypodium callophyllum C.H.Wright; Bull. Misc. Inform. Kew 1909: 362 (1909)
= Polypodium capillatum Brause; Bot. Jahrb. Syst. 49: 39 (1912)
= Tomophyllum capillatum (Brause) Parris; Gard. Bull. Singapore 58(2): 247 (2007)
species :
Tomophyllum congregatifolium
(Alderw.) Parris; Gard. Bull. Singapore 58(2): 247 (2007)
Distribution: New Guinea (Irian Jaya)
42 NWG
Synonyms: hidden
= Ctenopteris congregatifolia (Alderw.) Copel.; Philipp. J. Sci. 81: 112 (1953)
= Grammitis congregatifolia (Alderw.) Christenh.; Global Fl. 1(4): 42 (2018)
= Polypodium congregatifolium Alderw.; Nova Guinea 14: 47 (1924)
species :
Tomophyllum cumingii
Parris; sp. nov. in prep.
Distribution: Philippines (Luzon)
42 PHI
Remarks: name mentioned in Ebihara et al. (2012): Rare and Threatened Pteridophytes of Asia 1.
species :
Tomophyllum daymanense
Parris; sp. nov. in prep.
Distribution: New Guinea (Papua New Guinea)
42 NWG
Remarks: name mentioned in Ebihara et al. (2012): Rare and Threatened Pteridophytes of Asia 1.
species :
Tomophyllum dissitum
Parris; sp. nov. in prep.
Distribution: Sumatra
42 SUM
Remarks: name mentioned in Ebihara et al. (2012): Rare and Threatened Pteridophytes of Asia 1.
species :
Tomophyllum donianum
(Spreng.) Fraser-Jenk. & Parris; Taxon. Revis. Indian Subcontinental Pteridophytes 75 (2008)
Distribution: China (Anhui, Guizhou, Hunan, Sichuan, Yunnan); Tibet; Taiwan; Nepal; Bhutan; N-India (Uttarakhand, Sikkim, Darjeeling, Meghalaya, Arunachal Pradesh, Himachal Pradesh); Myanmar (Kachin, Chin, etc.)
36 CHI TIB 38 TAI 40 BHU IND NEP 41 MYA
Remarks: has been treated as T. subfalcatum in the Indian subcontinent fide Fraser-Jenkins 2008
Synonyms: hidden
= Grammitis doniana (Spreng.) Christenh.; Global Fl. 1(4): 43 (2018)
= Grammitis sinica (Christ) Christenh.; Global Fl. 1(4): 49 (2018)
= Polypodium convolutum Baker; Bull. Misc. Inform. Kew 1906: 12 (1906)
= Polypodium donianum Spreng.; Syst. Veg. (ed. 16): 4(1): 54 (1827), nom. nov. for Polypodium tenellum Don
= Polypodium sinicum Christ; Bull. Herb. Boissier 7: 3 (1899)
= Polypodium subfalcatum var. sinicum (Christ) C.Chr.; Index Filic., Suppl. Tert.: 147 (1934)
= Polypodium tenellum D.Don; Prodr. Fl. Nepal.: 2 (1825), nom. illeg., non G. Forst.
= Polypodium tenuissimum Hayata; Icon. Pl. Formosan. 4: 254, f. 178 (Sept.) (1914) (non Copel. (Febr. 1914))
= Tomophyllum sinicum (Christ) Parris; Gard. Bull. Singapore 58(2): 248 (2007)
species :
Tomophyllum duriusculum
(Christ) Parris; Fern Gaz. 19(6): 209 (2013)
Distribution: Sulawesi
42 SUL
Remarks: known only from the type
Synonyms: hidden
= Grammitis duriuscula (Christ) Christenh.; Global Fl. 1(4): 43 (2018)
= Polypodium duriusculum Christ; Verh. Naturf. Ges. Basel 11: 213 (1895)
species :
Tomophyllum epaleatum
(Parris) Parris; Gard. Bull. Singapore 58(2): 247 (2007)
Distribution: Sri Lanka; Java
40 SRL 42 JAW
Synonyms: hidden
= Ctenopteris epaleata Parris; Fern Gaz. 16(5): 239 (2001 publ. 2002)
= Grammitis epaleata (Parris) Christenh.; Global Fl. 1(4): 43 (2018)
species :
Tomophyllum foersteri
(Rosenst.) Parris; Fern Gaz. 19(6): 210 (2013)
Distribution: New Guinea (Papua New Guinea, Irian Jaya)
42 NWG
Synonyms: hidden
= Grammitis foersteri (Rosenst.) Christenh.; Global Fl. 1(4): 44 (2018)
= Polypodium bipinnatifidum var. foersteri Rosenst.; Repert. Spec. Nov. Regni Veg. Beih. 12: 179 (1913)
species :
Tomophyllum hayatae
(Masam.) T.C.Hsu; Taiwania 64(4): 379 (2019)
Distribution: Taiwan
38 TAI
Synonyms: hidden
= Ctenopteris hayatae (Masam.) H.Itô; J. Jap. Bot. 11: 90 (1935)
= Polypodium hayatae Masam.; J. Soc. Trop. Agric. Formosa 2: 31 (1930)
species :
Tomophyllum hymenophylloides
(Parris) Parris; Gard. Bull. Singapore 58(2): 247 (2007)
Distribution: New Guinea (Irian Jaya: Mt. Jaya)
42 NWG
Synonyms: hidden
= Ctenopteris hymenophylloides Parris; Kew Bull. 59(2): 221 (2004)
= Grammitis hymenophylloides (Parris) Christenh.; Global Fl. 1(4): 44 (2018)
species :
Tomophyllum inconspicuum
(Blume) Parris; Gard. Bull. Singapore 58(2): 247 (2007)
Distribution: peninsular Malaysia (Pahang); Borneo (Mt. Kinabalu, Sabah); Philippines; Java; Palau
42 BOR JAW MLY PHI 62 PLU
Synonyms: hidden
= Ctenopteris inconspicua (Blume) Copel.; Philipp. J. Sci. 81: 103 (1953)
= Grammitis inconspicua (Blume) Christenh.; Global Fl. 1(4): 45 (2018)
= Polypodium gedeense Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 16: 33 (1914)
= Polypodium inconspicuum Blume; Enum. Pl. Javae 2: 130 (1828)
= Polypodium parvulum Blume; Fl. Javae, Fil. 187, t. 37 (1828) (non Bory (1810))
species :
Tomophyllum lividum
(Mett.) Parris; Fern Gaz. 19(6): 210 (2013)
Distribution: Java
42 JAW
Synonyms: hidden
= Grammitis livida (Mett.) Christenh.; Global Fl. 1(4): 45 (2018)
= Polypodium lividum Mett.; Ann. Mus. Bot. Lugduno-Batavi 2: 221 (1866)
species :
Tomophyllum macrum
(Copel.) Parris; Gard. Bull. Singapore 58(2): 247 (2007)
Distribution: Palawan; Philippines (Mindanao)
42 PHI PLW
Synonyms: hidden
= Ctenopteris macra (Copel.) Copel.; Fern Fl. Philip. 3: 529 (1960)
= Grammitis macra (Copel.) Christenh.; Global Fl. 1(4): 46 (2018)
= Polypodium macrum Copel.; Perkins, Fragm. Fl. Philip. 189 (1905)
species :
Tomophyllum millefolium
(Blume) Parris; Gard. Bull. Singapore 58(2): 247 (2007)
Distribution: Sumatra; Borneo (Mt. Kinabalu, Sabah); Sulawesi; Philippines; Java; Moluccas (Seram); New Guinea (Irian Jaya, Papua New Guinea)
42 BOR JAW MOL NWG PHI SUL SUM
Synonyms: hidden
= Ctenopteris millefolia (Blume) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 219 (1947)
= Grammitis millefolia (Blume) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 240 (1941)
= Polypodium alderwereltii Rosenst.; Index Filic., Suppl. (1906-1912): 58 (1913)
= Polypodium koningsbergeri Rosenst.; Nova Guinea 8: 726 (1912), nom. illeg., non Alderw. (1908)
= Polypodium millefolium Blume; Enum. Pl. Javae 2: 134 (1828)
species :
Tomophyllum minutum
(Blume) Parris; Gard. Bull. Singapore 58(2): 247 (2007)
Distribution: Borneo (Mt. Kinabalu, Sabah); Philippines (Luzon); Java
42 BOR JAW PHI
Synonyms: hidden
= Acrostichum serratum Poir.; Lam., Encycl., Suppl. 1: 121 (1810), non Baker (1874)
= Ctenopteris minuta (Blume) Holttum; Rev. Fl. Mal. 2: 228 (1955)
= Ctenopteris smithiana Kunze; Zoll., Syst. Verz. 37 (1854)
= Grammitis minuta (Blume) Christenh.; Global Fl. 1(4): 46 (2018)
= Polypodium minutum Blume; Enum. Pl. Javae 2: 130 (1828)
= Schizaea serrata (Poir.) Fée; Mém. Foug. 2 (Hist. Acrostich.): 112 (1845)
species :
Tomophyllum perplexum
(Parris) Parris; Gard. Bull. Singapore 58(2): 248 (2007)
Distribution: S-India (Tamil Nadu); Sri Lanka
40 IND SRL
Conservation Status: IUCN CR for India (Fraser-Jenkins 2012)
Synonyms: hidden
= Ctenopteris perplexa Parris; Fern Gaz. 16(4): 201 (2001)
= Grammitis perplexa (Parris) Christenh.; Global Fl. 1(4): 47 (2018)
species :
Tomophyllum plicatum
Parris; sp. nov. in prep.
Distribution: Sulawesi (Gunung Tambusisi)
42 SUL
Remarks: name mentioned in Ebihara et al. (2012): Rare and Threatened Pteridophytes of Asia 1.
species :
Tomophyllum polytrichum
(Copel.) Parris; Gard. Bull. Singapore 58(2): 248 (2007)
Distribution: New Guinea (Irian Jaya, Papua New Guinea)
42 NWG
Synonyms: hidden
= Ctenopteris polytricha Copel.; Univ. Calif. Publ. Bot. 18: 225 (1942)
= Grammitis polytricha (Copel.) Christenh.; Global Fl. 1(4): 47 (2018)
species :
Tomophyllum pulchrum
Parris; Fern Gaz. 21(7): 405 (2022)
Distribution: Solomon Isl. (Guadalcanal)
43 SOL
species :
Tomophyllum repandulum
(Kunze ex Mett.) Parris; Gard. Bull. Singapore 58(2): 248 (2007)
Distribution: Sri Lanka; Vietnam; Borneo (Mt. Kinabalu, Sabah, Sarawak); Sulawesi; Philippines; Moluccas (Seram); New Guinea (Irian Jaya, Papua New Guinea); Solomon Isl. (Guadalcanal)
40 SRL 41 VIE 42 BOR MOL NWG PHI SUL 43 SOL
Synonyms: hidden
= Ctenopteris repandula (Kunze ex Mett.) C.Chr. & Tardieu; Lecomte, Fl. Gener. Indo-Chine 7, 2: 533 (1941)
= Ctenopteris repandula Kunze; Zoll., Syst. Verz. 37 (1854)
= Grammitis repandula (Mett.) Christenh.; Global Fl. 1(4): 48 (2018)
= Polypodium repandulum Kunze ex Mett.; Abh. Senckenberg. Naturf. Ges. 2: 50, n. 54 (1857)
= Polypodium subfalcatum Bedd.; Handb. Ferns Brit. India, t. 189 (1866)
species :
Tomophyllum sakaguchianum
(Koidz.) Parris; Gard. Bull. Singapore 58(2): 248 (2007)
Distribution: Japan
38 JAP
Conservation Status: EN in Japan. IUCN R
Synonyms: hidden
= Ctenopteris sakaguchiana (Koidz.) H.Itô; J. Jap. Bot. 11: 90 (1935)
= Grammitis sakaguchiana (Koidz.) Sugim. ex Sa.Kurata; Hokuriku J. Bot. 6: 9 (1957)
= Polypodium sakaguchianum Koidz.; Acta Phytotax. Geobot. 1: 29 (1932)
species :
Tomophyllum secundum
(Ridl.) Parris; Fern Gaz. 19(6): 210 (2013)
Distribution: New Guinea (Irian Jaya, Papua New Guinea)
42 NWG
Synonyms: hidden
= Grammitis secunda (Ridl.) Christenh.; Global Fl. 1(4): 48 (2018)
= Polypodium secundum Ridl.; Trans. Linn. Soc. London, Bot. 9: 262 (1916)
species :
Tomophyllum sesquipinnatum
(Copel.) Parris; Fern Gaz. 19(6): 210 (2013)
Distribution: New Guinea (Irian Jaya)
42 NWG
Synonyms: hidden
= Ctenopteris sesquipinnata Copel.; Univ. Calif. Publ. Bot. 18: 225 (1942)
= Grammitis sesquipinnata (Copel.) Christenh.; Global Fl. 1(4): 48 (2018)
species :
Tomophyllum subfalcatum
(Blume) Parris; Gard. Bull. Singapore 58(2): 248 (2007)
Distribution: Thailand; Borneo; Java; Lesser Sunda Isl. (Flores); Moluccas
41 THA 42 BOR JAW LSI MOL
Synonyms: hidden
= Ctenopteris subfalcata (Blume) Kunze; Bot. Zeitung (Berlin) 120 (1848)
= Grammitis subfalcata (Blume) Ching; Bull. Fan Mem. Inst. Biol. Bot. 10: 16 (1940)
= Polypodium filipendulifolium Fée; Mém. Foug. 5 (Gen. Filic.): 240 (1852)
= Polypodium parvulum Bedd.; Ferns South. India, t. 166 (1863)
= Polypodium subfalcatum Blume; Enum. Pl. Javae 2: 130 (1828)
= Polypodium subfalcatum var. glabrum Bedd.; Ferns Brit. India: t. 189B (1866)
= Polypodium trichophyllum Baker; Bull. Misc. Inform. Kew 1906: 13 (1906)
species :
Tomophyllum subminutum
(Alderw.) Parris; Gard. Bull. Singapore 58(2): 249 (2007)
Distribution: peninsular Malaysia (Perak, Pahang, Kedah); Sumatra; Borneo (Sarawak, Mt. Kinabalu, Sabah); Java
42 BOR JAW MLY SUM
Synonyms: hidden
= Ctenopteris subminuta (Alderw.) Holttum; Rev. Fl. Mal. 2: 228, f. 127 (1955)
= Grammitis subminuta (Alderw.) Christenh.; Global Fl. 1(4): 49 (2018)
= Polypodium subminutum Alderw.; Malayan Ferns 598 (1909)
species :
Tomophyllum subrepandulum
(Christ) Parris; Gard. Bull. Singapore 58(2): 249 (2007)
Distribution: Borneo; New Guinea (Irian Jaya, Papua New Guinea); Solomon Isl.
42 BOR NWG 43 SOL
Synonyms: hidden
= Grammitis subrepandula (Christ) Christenh.; Global Fl. 1(4): 49 (2018)
= Polypodium subrepandulum Christ; Ann. Jard. Buit. II(5): 119 (1905)
species :
Tomophyllum subsecundodissectum
(Zoll.) Parris; Gard. Bull. Singapore 58(2): 249 (2007)
Distribution: Borneo (Mt. Kinabalu, Sabah); Philippines; Java; Moluccas (Seram); New Guinea (Irian Jaya, Papua New Guinea); Bismarck Arch. (New Ireland); New Caledonia
42 BOR JAW MOL NWG PHI 43 BIS 60 NWC
Synonyms: hidden
= Ctenopteris subsecundodissecta (Zoll.) Copel.; Philipp. J. Sci. 81: 114 (1953)
= Grammitis subsecundodissecta (Zoll.) Christenh.; Global Fl. 1(4): 49 (2018)
= Polypodium gedeanum Racib.; Pteridoph. Buitenzorg 96 (1898)
= Polypodium subsecundodissectum Zoll.; Syst. Verz. 38, 48 (1854)
species :
Tomophyllum terpsichoroides
Parris; sp. nov. in prep.
Distribution: New Guinea (Papua New Guinea: Mt. Saruwaged)
42 NWG
Remarks: name mentioned in Ebihara et al. (2012): Rare and Threatened Pteridophytes of Asia 1.
species :
Tomophyllum themelioides
Parris; sp. nov. in prep.
Distribution: Moluccas (Seram: Gunung Binaiya)
42 MOL
Remarks: name mentioned in Ebihara et al. (2012): Rare and Threatened Pteridophytes of Asia 1.
species :
Tomophyllum walleri
(Maiden & Betche) Parris; Gard. Bull. Singapore 58(2): 249 (2007)
Distribution: Australia (NE-Queensland)
50 AUS
Conservation Status: IUCN V
Synonyms: hidden
= Ctenopteris walleri (Maiden & Betche) S.B.Andrews; Austrobaileya 1(1): 12 (1977)
= Grammitis walleri (Maiden & Betche) Christenh.; Global Fl. 1(4): 50 (2018)
= Polypodium walleri Maiden & Betche; Proc. Linn. Soc. New South Wales 35. 799 (1910 publ. 1911)
species :
Tomophyllum warburgii
Parris; sp. nov. in prep.
Distribution: Java (Gunung Wilis)
42 JAW
Remarks: name mentioned in Ebihara et al. (2012): Rare and Threatened Pteridophytes of Asia 1.
