World Species of Biting Midges (Diptera: Ceratopogonidae)

Last updated: Feb. 20, 2013 

- If you find any errors or have any suggestions for the improvement of this catalog please contact Art 

Borkent at artborkent@telus.net 

World Species of Biting Midges (Diptera: Ceratopogonidae) 

by 

ART BORKENT 

Research Associate of the Royal British Columbia Museum, 

American Museum of Natural History, and Instituto Nacional de Biodiversidad 

691-8th Ave. SE, Salmon Arm, British Columbia, V1E 2C2, Canada. 

email: artborkent@telus.net

CONTENTS 

LOCATION OF TYPE MATERIAL ............................................................................................................................. 6 

SUBFAMILY LEBANOCULICOIDINAE BORKENT ................................................................................................. 11 

LEBANOCULICOIDES Szadziewski ............................................................................................. 11 

SUBFAMILY LEPTOCONOPINAE NOÈ ................................................................................................................. 11 

ARCHIAUSTROCONOPS Szadziewski ........................................................................................ 11 

AUSTROCONOPS Wirth and Lee ................................................................................................. 11 

FOSSILEPTOCONOPS Szadziewski ............................................................................................ 11 

JORDANOCONOPS Szadziewski ................................................................................................. 11 

LEPTOCONOPS Skuse ................................................................................................................ 12 

MINYOHELEA Borkent .................................................................................................................. 16 

SUBFAMILY FORCIPOMYIINAE LENZ .................................................................................................................. 16 

ATRICHOPOGON Kieffer .............................................................................................................. 16 

ATRICULICOIDES Remm ............................................................................................................. 27 

FORCIPOMYIA Meigen ................................................................................................................. 28 

ATOPOMYIA Yu and Liu .................................................................................................. 28 

BASSOFORCIPOMYIA Debenham ................................................................................. 28 

BLANTONIA Wirth and Dow ............................................................................................. 28 

CALOFORCIPOMYIA Saunders ...................................................................................... 29 

COLLESSOHELEA Debenham ........................................................................................ 29 

DYCEA Debenham ........................................................................................................... 30 

EUPROJOANNISIA Brèthes ............................................................................................ 30 

FORCIPOHELEA Clastrier and Delécolle ........................................................................ 32 

FORCIPOMYIA Meigen .................................................................................................... 33 

GAMPSOHELEA Yu and Liu ............................................................................................ 41 

HERAKLEOHELEA Debenham ........................................................................................ 41 

IXODEHELEA Yu and Liu ................................................................................................ 41 

JAPYGAHELEA Yu and Liu ............................................................................................. 41 

KATTANGOMYIA Debenham .......................................................................................... 41 

LASIOHELEA Kieffer ........................................................................................................ 41 

LEPIDOHELEA Kieffer ..................................................................................................... 45 

METAFORCIPOMYIA Saunders ...................................................................................... 47 

MICROHELEA Kieffer ...................................................................................................... 48 

NICOTHOHELEA Liu, Tang and Hao ............................................................................... 50 

OREINOHELEA Yu and Liu ............................................................................................. 51 

PANHELEA Remm ........................................................................................................... 51 

PEDILOHELEA de Meillon and Wirth ............................................................................... 51 

PHYTOHELEA Remm ...................................................................................................... 51 

PTEROBOSCA Macfie ..................................................................................................... 52 

RHINOHELEA de Meillon and Wirth ................................................................................ 53 

RHYNCHOFORCIPOMYIA Wirth and Dow ..................................................................... 53 

SALIOHELEA Wirth and Ratanaworabhan ...................................................................... 53 

SCHINEROMYIA Debenham ........................................................................................... 53 

SCHIZOFORCIPOMYIA Chan and LeRoux ..................................................................... 54 

SYNTHYRIDOMYIA Saunders ......................................................................................... 54 

THYRIDOMYIA Saunders ................................................................................................ 55 

TRICHOHELEA Goetghebuer .......................................................................................... 56 

TRITHICOMYIA Yu and Song .......................................................................................... 58 

TYPHONOMYIA Debenham ............................................................................................ 58 

WARMKEA Saunders ....................................................................................................... 58

SUBFAMILY DASYHELEINAE LENZ ..................................................................................................................... 58 

DASYHELEA Kieffer ...................................................................................................................... 59 

SUBFAMILY CERATOPOGONINAE NEWMAN ..................................................................................................... 72 

TRIBE CULICOIDINI KIEFFER .................................................................................................................. 72 

CULICOIDES Latreille ................................................................................................................... 72 

PARADASYHELEA Macfie .......................................................................................................... 105 

WASHINGTONHELEA Wirth and Grogan ................................................................................... 105 

TRIBE CERATOPOGONINI ..................................................................................................................... 105 

ADELOHELEA Borkent ............................................................................................................... 105 

AFROHELEA Wirth ..................................................................................................................... 106 

AGILIHELEA Yu .......................................................................................................................... 106 

ALLOHELEA Kieffer .................................................................................................................... 106 

ALLUAUDOMYIA Kieffer ............................................................................................................. 107 

ANKYLOHELEA de Meillon and Wirth ........................................................................................ 111 

ATYPHOHELEA Borkent ............................................................................................................. 111 

AUSTROHELEA Wirth and Grogan ............................................................................................ 112 

BAEODASYMYIA Clastrier and Raccurt ..................................................................................... 112 

BAEOHELEA Wirth and Blanton ................................................................................................. 112 

BAHIAHELEA Wirth ..................................................................................................................... 112 

BOREOHELEA Clastrier and Delécolle ....................................................................................... 112 

BORKENTHELEA Spinelli and Grogan ....................................................................................... 112 

BOTHAHELEA Grogan and Wirth ............................................................................................... 113 

BOTHAMIA Meiswinkel ............................................................................................................... 113 

BRACHYCRETACEA Szadziewski ............................................................................................. 113 

BRACHYPOGON Kieffer ............................................................................................................. 113 

CACAOHELEA Wirth and Grogan ............................................................................................... 117 

CALCARHELEA Wirth and Grogan ............................................................................................. 118 

CAMPTOPTEROHELEA Wirth and Hubert ................................................................................. 118 

CAPEHELEA de Meillon and Wirth ............................................................................................. 118 

CERATOCULICOIDES Wirth and Ratanaworabhan ................................................................... 118 

CERATOHELEA Wirth and Grogan ............................................................................................ 118 

CERATOPALPOMYIA Szadziewski ............................................................................................ 118 

CERATOPOGON Meigen ........................................................................................................... 119 

CHAIROPOGON Yu .................................................................................................................... 120 

CONGOHELEA Wirth and Grogan .............................................................................................. 120 

DIAPHANOBEZZIA Ingram and Macfie ...................................................................................... 120 

DOWNESHELEA Wirth and Grogan ........................................................................................... 121 

ECHINOHELEA Macfie ............................................................................................................... 121 

EOHELEA Petrunkevitch ............................................................................................................. 122 

FANTHAMIA de Meillon .............................................................................................................. 122 

FITTKAUHELEA Wirth and Blanton ............................................................................................ 123 

FOSSIHELEA Szadziewski ......................................................................................................... 123 

GEDANOHELEA Szadziewski .................................................................................................... 123 

HELEAGERON Borkent .............................................................................................................. 123 

HETEROCERATOPOGON Wirth and Grogan ............................................................................ 123 

HETEROHELEA Clastrier ........................................................................................................... 123 

HYPSIMYIA Yu ............................................................................................................................ 124 

ISTHMOHELEA Ingram and Macfie ............................................................................................ 124 

KOLENOHELEA de Meillon and Wirth ........................................................................................ 124 

LEPTOHELEA Wirth and Blanton ............................................................................................... 124 

LUCIAMYIA de Meillon ................................................................................................................ 124 

MACRUROHELEA Ingram and Macfie ....................................................................................... 124 

METACANTHOHELEA Wirth and Grogan .................................................................................. 125

MANTOHELEA Szadziewski ....................................................................................................... 125 

MEUNIEROHELEA Szadziewski ................................................................................................. 125 

MONOHELEA Kieffer .................................................................................................................. 125 

NANNOHELEA Grogan and Wirth .............................................................................................. 127 

NEOHELEA Clastrier ................................................................................................................... 127 

NEUROBEZZIA Wirth and Ratanaworabhan .............................................................................. 127 

NOTIOHELEA Grogan and Wirth ................................................................................................ 128 

NOTOCERATOPOGON de Meillon and Downes ....................................................................... 128 

OXYRIA Yu .................................................................................................................................. 128 

PALAEOBRACHYPOGON Borkent ............................................................................................ 128 

PARABEZZIA Malloch ................................................................................................................. 128 

PARALLUAUDOMYIA Clastrier ................................................................................................... 129 

PARASTILOBEZZIA Wirth and Blanton ...................................................................................... 129 

PERONEHELEA Borkent ............................................................................................................ 129 

PSEUDOSTILOBEZZIA Wirth and Ratanaworabhan .................................................................. 129 

RHYNCHOHELEA Wirth and Blanton ......................................................................................... 130 

SCHIZOHELEA Kieffer ................................................................................................................ 130 

SCHIZONYXHELEA Clastrier ..................................................................................................... 130 

SERROMYIA Meigen .................................................................................................................. 130 

SINHALOHELEA Grogan and Borkent ........................................................................................ 132 

SPINELLIHELEA Borkent, Grogan and Picado ........................................................................... 132 

STILOBEZZIA Kieffer .................................................................................................................. 132 

STILOCULICOIDES Wirth and Grogan ....................................................................................... 140 

WIRTHOHELEA Szadziewski ..................................................................................................... 140 

TRIBE HETEROMYIINI WIRTH ............................................................................................................... 140 

CLINOHELEA Kieffer .................................................................................................................. 140 

HETEROMYIA Say ...................................................................................................................... 141 

METAHELEA Edwards ................................................................................................................ 142 

NEMOROMYIA Liu ...................................................................................................................... 142 

NEUROHELEA Kieffer ................................................................................................................ 142 

PELLUCIDOMYIA Macfie ............................................................................................................ 142 

PHYSOHELEA Grogan and Wirth ............................................................................................... 142 

TETRABEZZIA Kieffer ................................................................................................................. 143 

TRIBE SPHAEROMIINI NEWMAN .......................................................................................................... 143 

ALLOIMYIA Yu and Liu ............................................................................................................... 143 

AUSTROSPHAEROMIAS Spinelli ............................................................................................... 143 

CALYPTOPOGON Kieffer ........................................................................................................... 143 

CHELOHELEA Giles and Wirth ................................................................................................... 143 

CRISPOMYIA Debenham ........................................................................................................... 143 

DIBEZZIA Kieffer ......................................................................................................................... 144 

GROGANHELEA Spinelli and Dippolito ...................................................................................... 144 

GUIHELEA Yu and Qian ............................................................................................................. 144 

HEBETULA Wirth and Debenham ............................................................................................... 144 

HOMOHELEA Kieffer .................................................................................................................. 145 

INDOBEZZIA Das Gupta and Saha ............................................................................................ 145 

JENKINSHELEA Macfie .............................................................................................................. 145 

JOHANNSENOMYIA Malloch ..................................................................................................... 146 

LANATOMYIA Debenham ........................................................................................................... 146 

LANEHELEA Wirth and Blanton .................................................................................................. 147 

LEEHELEA Debenham ............................................................................................................... 147 

MACKERRASOMYIA Debenham ................................................................................................ 147 

MACROPEZA Meigen ................................................................................................................. 147 

MALLOCHOHELEA Wirth ........................................................................................................... 148 

NEOBEZZIA Wirth and Ratanaworabhan ................................................................................... 149

NEOSPHAEROMIAS Das Gupta and Wirth ................................................................................ 149 

NILOBEZZIA Kieffer .................................................................................................................... 150 

NIPHANOHELEA Grogan and Wirth ........................................................................................... 151 

PROBEZZIA Kieffer ..................................................................................................................... 152 

SPHAEROHELEA Spinelli and Felippe-Bauer ............................................................................ 152 

SPHAEROMIAS Curtis ................................................................................................................ 153 

WANNOHELEA Yu ...................................................................................................................... 154 

XENOHELEA Kieffer ................................................................................................................... 154 

TRIBE PALPOMYIINI Enderlein ............................................................................................................... 154 

AMEROHELEA Grogan and Wirth .............................................................................................. 154 

BEZZIA Kieffer ............................................................................................................................. 154 

CLASTRIEROMYIA Spinelli and Grogan .................................................................................... 162 

PACHYHELEA Wirth ................................................................................................................... 162 

PALPOMYIA Meigen ................................................................................................................... 162 

PHAENOBEZZIA Haeselbarth .................................................................................................... 169 

TRIBE STENOXENINI Coquillett .............................................................................................................. 170 

PARYPHOCONUS Enderlein ...................................................................................................... 170 

STENOXENUS Coquillett ............................................................................................................ 171 

Taxa Unplaced in Ceratopogonidae ...................................................................................................................... 172 

ALAUTUNMYIA Borkent .............................................................................................................. 172 

ARCHICULICOIDES Szadziewski ............................................................................................... 172 

PROTOCULICOIDES Boesel ...................................................................................................... 172 

SINOPOGONITES Hong ............................................................................................................. 172 

Unplaced Genus of Forcipomyiinae ...................................................................................................................... 172 

Unplaced Genera of Ceratopogoninae .................................................................................................................. 172 

Unplaced Genus of Heteromyiini + Sphaeromiini + Palpomyiini + Stenoxenini .................................................... 172 

Unplaced Species of Forcipomyiinae .................................................................................................................... 172 

Unplaced Species of Ceratopogoninae ................................................................................................................. 173 

Unplaced species of Heteromyiini + Sphaeromiini + Palpomyiini + Stenoxenini ................................................... 173 

Unplaced Species of Ceratopogonidae ................................................................................................................. 173 

Species in Other Families Originally Considered to Be Ceratopogonidae ............................................................ 173 

TABLE 1 ................................................................................................................................................................ 233 

TABLE 2 ................................................................................................................................................................ 234

COMMENTARY ON PLACEMENT OF SOME TAXA 

Wang et al. (2012) described the genus Sinicohelea and indicated that it was a member of the 

Ceratopogoninae but unplaced to tribe. The description indicates it is a member of the 

Sphaeromiini and it is very similar to Johannsenomyia, with which it shares the derived feature of a strong row of 

bristles at the apex of male tergite 9. I have therefore placed the genus in the Sphaeromiini below. 

LOCATION OF TYPE MATERIAL 

Some publications that provide information on the location of type material of Ceratopogonidae are listed 

in table 1. The location where ceratopogonid workers have deposited most or all of their type material is listed in 

table 2. The list is not complete; some authors who have described only one or a few species and whose type 

material could not be located or confirmed were not included. However, the list provides a basis for the study and 

loan of most type material. Acronyms for museums are those proposed by Arnett and Samuelson (1986) or follow 

their method for forming acronyms: 

ANIC - Australian National Insect Collection, Division of Entomology, CSIRO, P.O. Box 1700, Canberra City, 

A.C.T. 2601, Australia. 

ANSP - Department of Entomology, Academy of Natural Sciences, 19th and the Parkway, Philadelphia, 

Pennsylvania, 19103, USA. 

BIAR - Biologicheskii Institut, Akademiya NAUK Russia, Sibirskoe Otdelenie, Frunze str. 11, 630091 

Novosibirsk, Russia. 

BMNH - Department of Entomology, Natural History Museum, London, SW7 5BD, United Kingdom. 

BPBM - Dept of Entomology, Bernice P. Bishop Museum, 1525 Bernice Street, P.O. Box 19000A, Honolulu, 

Hawaii, 96817 0916, USA. 

CASC - Department of Entomology, California Academy of Sciences, Golden Gate Park, San Francisco, 

California, 94118, USA. 

CNCI - Canadian National Collection of Insects, Eastern Cereal and Oilseed Research Centre, Agriculture 

Canada, K.W. Neatby Building, Ottawa, Ontario, K1A 0C6, Canada. 

CUIC - Department of Entomology, Comstock Hall, Cornell University, Ithaca, New York, 14853-0999, USA. 

DEFS - Departamento de Epidemiologia, Faculdade de Saude Publica, Universidade de Sao Paulo, Av. Dr.

Arnaldo, 715, 01.255, Sao Paulo, SP, Brazil. 

DEIC - Deutsches Entomologisches Institut, Eberswalder Straße 90, Muencheberg 15374, Germany. 

DPSC - Department of Parasitology, Second Military Medical College, Shanghai, China. 

ELBU - Entomology Laboratory, Department of Zoology, University of Burdwan, Burdwan 713 104, WB, India. 

ELKU - Entomological Laboratory, Faculty of Agriculture, Kyushu University, Fukuoka, Japan. 

EPRL - Entomological Pioneering Research Laboratory, University of Puerto Rico, Mayaguez, Puerto Rico, 

00708. 

FIOC - Instituto Oswaldo Cruz, Departamento de Entomologia, Caixa Postal 926, 20010 Rio de Janeiro, Rio de 

Janeiro, Brasil. 

FMMF - Lab. Ecol. Med. Pathol. Parasitologie, Faculté de Médecine, Rue Auguste-Broussonet, F-34000, 

Montpellier, France. 

FMNH - Insect Collection, Field Museum of Natural History, Roosevelt Road and Lake Shore Drive, Chicago, 

Illinois, 60605, USA. 

HNHM - Zoological Dept., Hungarian Natural History Museum, Baross utca 13, H-1088, Budapest, Hungary. 

IDRE - Laboratorio de Entomología, Instituto de Diagnóstico y Referencia Epidemiológicos, SSA. Carpio 470, Col. 

Santo Tomás, 2do. Piso, México D.F. 11340, Mexico (formerly, ISET=Instituto de Enfermedades 

Tropicales). 

IEME - Institute for Evolution, Morphology, and Ecology of Animals, Leninsky pr. 33, Moscow 117071, Russia. 

IMBC - Department of Vector and Control, Institute of Microbiology and Epidemiology, 23 (A) Qilizhuang Road, 

Fengtai District, Beijing 100071, People's Republic of China. 

IMGP - Institut und Museum für Geologie und Palaontologie der Georg-August-Universitat, Göttingen, Germany. 

INBC - Instituto Nacional de Biodiversidad, 400 Mts Oeste del Cementerio, A.P. 22-3100 Santo Domingo de 

Heredia, Costa Rica. 

INHS - Illinois Natural History Survey Insect Collection, 607 E. Peabody Drive, Champaign, Illinois, 61820, USA. 

INMA - Departamento de Arthropodologia Sanitaria, Instituto Nacional de Microbiologia, "Dr. Carlos G. Malbran," 

Avda. Velez Sarfield 563, 1281 Capital Federal, Argentina. 

IPCG - Institut Pasteur de la Guyane, Cayenne, Guyana. 

IPHT - Institute of Public Health Research, Teheran, Iran.

IPSF - Institut de Parasitologie, Faculté de Médecine, 3, Rue Koeberlé, F 67000, Strasbourg, France. 

ISNB - Collections Nationales Belges d'Insectes et d'Arachnides, Institut Royal des Sciences Naturelles de 

Belgique, 29, rue Vautier, B1040, Brussels, Belgium. 

IZBE - Institute of Zoology and Botany, Vanemuise str. 21, Tartu, 202400, Estonia. 

IZAS - Institute of Zoology, Academia Sinica, 7 Zhongguancun Lu, Haidian, 100080, Beijing, Republic of China. 

LACM - Los Angeles County Museum of Natural History, 900 Exposition Blvd., Los Angeles, California, 90007, 

USA. 

LEMQ - Lyman Entomological Museum and Research Laboratory, Macdonald College, 21111 Lakeshore, Ste.- 

Anne-de-Bellevue, Quebec, H9X 1C0. 

MABC - M. Baez, Departamento de Zoologia, Facultad de Biologia, Universidad de La Laguna, 38206 La 

Laguna, Tenerife Islas Canarias, Spain. 

MACN - Division Entomologia, Museo Argentino de Ciencias Naturales, Av. Angel Gallardo 470 (C.P. 220, Suc. 

5), 1405 Buenos Aires, Argentina. 

MCZC - Entomology Department, Museum of Comparative Zoology, Harvard University, 26 Oxford St., 

Cambridge, Massachusetts, 02138, USA. 

MEUC - Museo Entomologico, Facultad de Agronomia, Universidad de Chile, Casilla 1004, Santiago, Chile. 

MLPA - Museo de la Plata, Division Entomologia, Universidad de La Palata, Paseo del Bosque, 1900 La Plata, 

Argentina. 

MNHN - Museum National d'Histoire Naturelle, 45, rue Buffon, Paris, 75005, France. 

MRAC - Section d'Entomologie, Musée Royal de l'Afrique Centrale, Leuvensesleenweg 13, B-1980, Tervuren, 

Belgium. 

MZLU - Department of Zoology, Zoological Museum, Helgonavägen 3, S-223 62 Lund, Sweden. 

MZRF - Museo Civico de Storia Naturale "Giacomo Doria", Via Brigita Liguria 9, I-16121, Genoa, Italy. 

MZSF - Musée Zoologique, Université de Strasbourg, 29, blvd. de la Victoire, F-67000, Strasbourg, France. 

MZSP - Museu de Zoologia da Universidade de Sao Paulo, Av. Nazare, 481, C.P. 7172, 01.051, Sao Paulo, SP, 

Brazil. 

MZWP - Museum of the Earth, Al. Na Skarpie 20/26, 00-488, Warsaw, Poland. 

8

NHMW - Naturhistorisches Museum Wien, Postfach 417, Burgring 7, 1040 Wien, Austria. 

NHRS - Sektionen for entomologi, Naturhistoriska Riksmuseet, S-104 05, Stockholm, Sweden. 

NIPM - Department of Medical Entomology, National Institute of Preventive Medicine, 161 Kunyang Street, 

Nankang, Taipei, Taiwan, 11513, Republic of China. 

NMPC - Department of Entomology, National Museum of Natural History, 148 00, Praha 4, Kunratice 1, Czech 

Republic. 

NMSA - Natal Museum, 237 Loop Street, Pietermaritzburg 3201, Natal, South Africa. 

NSMT - Entomological Collections, National Science Museum (Natural History), Hyakunin-cho 3-23-1, Shinjuku-ku, Tokyo 160, 

NYSM - New York State Museum, Biological Survey, 2132 Cultural Education Center, Albany, New York, 12230, 

USA. 

NZAC - New Zealand Arthropod Collection, Entomology Division, DSIR, Private Bag, Auckland 1001, New 

Zealand. 

NZSI - National Zoological Collection, Zoological Survey of India, M-Block, New Alipur, Calcutta 700 053, India. 

ORST - Centre ORSTOM de Montpellier, Laboratoire de Taxonomie des Vecteurs 911, Avenue Agropolis, B.P. 

5045, F-34032, Montpellier Cedex, France 

PEHC - P. Havelka, Staatliche Vogelschutzwarte, Baden --- Württemberg, Hermann Schneider Allee 47, D-7500 

Karlsruhe --- Rappenwört, Germany. 

PIAM - Palaeontological Institute, The Russian Academy of Sciences, Profsoyuznaya, 113, Moscow 117321, 

Russia. 

PMNH - Peabody Museum of Natural History, Yale University, New Haven, Connecticut, 06511, USA. 

ROME - Department of Entomology, Royal Ontario Museum, Toronto, Ontario, M5S 2C6, Canada. 

RYSC - R. Szadziewski, Department of Invertebrate Zoology, University of Gdansk, Pilsudskiego 46, 81-378 

Gdynia, Poland. 

SAMR - South African Institute for Medical Research, P.O. Box 1038, Johannesburg, 2000, South Africa. 

SEMC - Snow Entomological Museum, University of Kansas, Lawrence, Kansas, 66044, USA. 

SMFD - Forschungsinstitut und Naturmuseum Senckenberg, Entomologische Section 1, Senckenberganlage 25, 

6000 Frankfurt 1, Germany. 

9

SMSH - Stovall Museum of Science and History, University of 

Oklahoma, Norman, Oklahoma, 73069, USA. 

SZRL - Systematic Zoology Museum, University of Latvia, Kromvalda bulv. 4, Riga, LV 10, Latvia. 

USNM - United States National Entomological Collection, Dept. of Entomology, U.S. National Museum of Natural 

History, Washington, DC 20560, USA. 

UZMH - Zoological Museum, Finnish Museum of Natural History, University of Helsinki, Pohjoinen Rautatiekatu 

13, SF-00100, Helsinki, Finland. 

VRIO - Veterinary Research Institute, Onderstepoort 0110, Republic of South Africa. 

ZMAN - Instituut voor Taxonomische Zoologie, Universiteit van Amsterdam, Plantage Middenlaan 64, 

Amsterdam, The Netherlands. 

ZMAS - Zoological Institute, Academy of Sciences, Universitetskaya nab., 1, St. Petersburg, 199034, Russia. 

ZMHB - Museum für Naturkunde der Humboldt Universität zu Berlin, Invalidenstrasse 43, Berlin 1040, Germany. 

ZMUC - Zoologisk Museum, Universitetsparken 15, DK 2100, Copenhagen, Denmark. 

ZSMC - Zoologische Staatssammlung, Münchhausenstrasse 21, D-8000 München 60, Germany. 

10

SUBFAMILY LEBANOCULICOIDINAE BORKENT, 2000a:361 

Genus LEBANOCULICOIDES Szadziewski 

LEBANOCULICOIDES Szadziewski, 1996: 30. Type species: Lebanoculicoides mesozoicus Szadziewski, by original 

designation. 

encantabris Pérez-de la Fuente, Delclòs, Peñalver and Arillo, 2011: 752. Spain. Lower Cretaceous. 

mesozoicus Szadziewski, 1996: 30. Lebanon. Lower Cretaceous. 

SUBFAMILY LEPTOCONOPINAE NOÈ, 1907: 143 

Genus ARCHIAUSTROCONOPS Szadziewski 

ARCHIAUSTROCONOPS Szadziewski, 1996: 34. Type species: Archiaustroconops ceratoformis Szadziewski, by original 

designation. 

alavensis Szadziewski and Arillo, 1998: 294. Spain. Lower Cretaceous. 

bocaparvus Borkent, 2000a: 374. Lebanon. Lower Cretaceous. 

borkenti Pérez-de la Fuente, Delclòs, Peñalver and Arillo, 2011: 755. Spain. Lower Cretaceous. 

ceratoformis Szadziewski, 1996: 34. Lebanon. Lower Cretaceous. 

cretaceous (Szadziewski), 1996: 36 (Austroconops). Lebanon. Lower Cretaceous. 

gracilis Szadziewski and Poinar, 2005: 355. Myanmar. Lower Cretaceous. 

hamus Borkent, 2000a: 373. Lebanon. Lower Cretaceous. 

kotejai Szadziewski and Poinar, 2005: 356. Myanmar. Lower Cretaceous. 

szadziewskii Borkent, 2000a: 372. Lebanon. Lower Cretaceous. 

Genus AUSTROCONOPS Wirth and Lee 

AUSTROCONOPS Wirth and Lee, 1958: 337. Type species: Austroconops mcmillani Wirth and Lee, by original designation. 

annettae Borkent in Borkent and Craig, 2004: 44. Australia (Western Australia). 

asiaticus Szadziewski, 2004:115. Myanmar. Lower Cretaceous. 

borkenti Szadziewski and Schlüter, 1992: 78. France. Upper Cretaceous. 

fossilis Szadziewski, 1996: 38. Lebanon. Lower Cretaceous. 

gladius Borkent, 2000a: 378. Lebanon. Lower Cretaceous. 

gondwanicus Szadziewski, 1996: 38. Lebanon. Lower Cretaceous. 

mcmillani Wirth and Lee, 1958: 337. Australia (Western Australia). 

megaspinus Borkent, 2000a: 381. Lebanon. Lower Cretaceous. 

sibiricus Szadziewski, 1996: 40. Russia. Upper Cretaceous. 

Genus FOSSILEPTOCONOPS Szadziewski 

FOSSILEPTOCONOPS Szadziewski, 1996: 46. Type species: Fossileptoconops lebanicus Szadziewski, by original 

designation. 

lebanicus Szadziewski, 1996: 47. Lebanon. Lower Cretaceous. 

Genus JORDANOCONOPS Szadziewski 

JORDANOCONOPS (as Jordanocops, misspelling) Szadziewski, 2000a: 252. Type species: Jordanconops weitschati 

Szadziewski, by original designation. 

11

weitschati Szadziewski, 2000a: 252. Jordan. Lower Cretaceous. 

Genus LEPTOCONOPS Skuse 

LEPTOCONOPS Skuse, 1889: 288. Type species: Leptoconops stygius Skuse, by monotypy. 

MYCTEROTYPUS Noè, 1905: 114. Type species: Mycterotypus bezzii Noè, designation by Carter, 1921: 3. 

PROTERSESTHES Kieffer, 1921h: 107. Type species: Tersesthes brasiliensis Lutz, by original designation. 

TERSESTHES Townsend, 1893: 370. Type species: Tersesthes torrens Townsend, by original designation. 

SCHIZOCONOPS Kieffer, 1918a: 135. Type species: Schizoconops indicus Kieffer, by monotypy. 

Subgenus BRACHYCONOPS Wirth and Atchley 

BRACHYCONOPS Wirth and Atchley, 1973: 15 (as subgenus of Leptoconops). Type species: Leptoconops californiensis 

Wirth and Atchley, by original designation. 

californiensis Wirth and Atchley, 1973: 16. USA (California). 

patagoniensis Ronderos, 1990b: 423. Argentina. 

Subgenus HOLOCONOPS Kieffer 

HOLOCONOPS Kieffer, 1918a: 135. Type species: Leptoconops kerteszi Kieffer, by original designation. 

MICROCONOPS Kieffer, 1921h: 108. Type species: Microconops vexans Kieffer, by original designation. 

acer Clastrier, 1973: 911. France. 

algeriensis Clastrier, 1975b: 41. Algeria. 

altuneshanensis Yu and Shao, in Yu, 1988: 128. China. 

americanus Carter, 1921: 22 (as variety of kerteszi). USA (Utah). 

amplifemoralis Chanthawanich and Delfinado, 1967: 298. Nepal. 

andersoni Clastrier and Wirth, 1978: 35. USA (California). 

arnaudi Clastrier and Wirth, 1978: 29. USA (California). 

ascius Yu and Hui, in Yu, 1988: 131. China. 

asilomar Clastrier and Wirth, 1978: 22. USA (California). 

atchleyi Clastrier and Wirth, 1978: 32. USA (California). 

auster Clastrier, 1981a: 91. South Africa. 

bassoi Ronderos and Spinelli, 1993: 305. Chile. 

beidaiheensis Liu and Yu, in Yu et al., 2005: 53. China. 

belkini Wirth and Atchley, 1973: 37. USA (California). 

bequaerti (Kieffer), 1925a: 405 (Holoconops; as becquaerti). Honduras. 

hondurensis Hoffman, 1926b: 135. Honduras. 

binangulus Yu, 1989b: 97. China. 

binisiculus Yu and Liu, in Yu, 1988: 130. China. 

borealis Gutsevich, 1945: 124. Russia. 

popovii Dzhafarov, 1961a: 72. Azerbaijan. 

brevistylus Mazumdar, Saha and Chaudhuri, 2010: 50. India. 

catawbae (Boesel), 1948: 69 (Holoconops). USA (Ohio). 

chenfui Yu and Xiang, in Yu, 1988: 130. China. 

conulus Yu and Liu, in Liu et al., 1990: 218 (in Liu et al., 1990: 13). China. 

dicheres Liu and Yu, in Yu et al., 2005: 62. China. 

dissimilis Clastrier, 1975a: 605. Chad. 

doyeni Spinelli and Ronderos, 1993: 115. Mexico. 

dunhuangensis Liu and Yu, in Yu et al., 2005: 63. China. 

exspectator Clastrier, 1975b: 39. Algeria. 

foleyi Clastrier, 1975b: 33. Algeria. 

12

fortipalpus Mazumdar, Saha and Chaudhuri, 2010: 51. India. 

foulki Clastrier and Wirth, 1978: 38. USA (California). 

gallicus Clastrier, 1973: 906. France. 

helobius Ma and Yu, in Ma et al., 1990: 49. China. 

interruptus (Enderlein), 1908: 460 (Mycterotypus). Namibia. 

jizhangi Liu and Yu, in Yu et al., 2005: 69. China. 

kerteszi Kieffer, 1908: 576. Egypt. 

knowltoni Clastrier and Wirth, 1978: 26. USA (California). 

lacteipennis Kieffer, 1918a: 32. Tunisia. 

laosensis Clastrier, 1974b: 71. Laos. 

latibulorum Gutsevich, 1973: 259. Kazakhstan. 

laurae (Weiss), 1912: 24 (Mycterotypus). Tunisia. 

mediterraneus (Kieffer), 1921i: 264 (Holoconops). Algeria. 

linleyi Wirth and Atchley, 1973: 49. USA (Florida). 

macfiei Clastrier, 1975b: 28. Egypt. 

mellori Clastrier and Boorman, 1987: 293. Bahrain. 

menglaensis Liu and Yu, in Yu et al., 2005: 71. China. 

monotheca Liu andYu, in Yu et al., 2005: 73. China. 

montanus Konurbajev, 1965: 134. Kyrgyzstan. 

nachitschevanicus Dzhafarov, 1961c: 78. Azerbaijan. 

nasiformus Liu and Yu, in Yu et al., 2005: 74. China. 

nevilli Clastrier, 1981a: 88. South Africa. 

ningxiaensis Liu and Yu, in Yu et al., 2005: 75. China. 

noterophilous Yu and Liu, in Liu et al., 1990: 15. China. 

pavlovskyi Dzhafarov, 1961a: 74. Azerbaijan. 

peneti (Langeron), 1913: 282 (Mycterotypus, as variety of laurae). Tunisia. 

pugnax Clastrier, 1973: 908. France. 

qinghaiensis Liu, Zhang and Gong, 2004: 49. China. 

reesi Clastrier and Wirth, 1978: 42. USA (Utah). 

riparins Yu, in Liu et al., 1990: 17 (also as riparius). China. 

rufiventris (Kieffer), 1923a: 658 (Microconops). Algeria. 

shangweni Xu and Yu, 1989: 276. China. 

sublettei Clastrier and Wirth, 1978: 36. USA (Texas). 

tarimensis Yu, 1982: 204 (1985: 406). China. 

tenebrostigmatus Mazumdar, Saha and Chaudhuri, 2010: 53. India. 

tetratheca Liu and Yu, in Yu et al., 2005: 83. China. 

tibetensis Lee, 1978: 115. China. 

transversalis (Kieffer), 1921h: 112 (Holoconops). Tunisia. 

triquetrus Liu and Yu, in Liu et al., 1990: 21. China. 

tuotuohea Liu and Gong, 2003: 549. China. 

umbellifer Clastrier, 1981a: 93. South Africa. 

utriculus Liu and Yu, in Yu et al., 2005: 89. China. 

vargasi Clastrier and Wirth, 1978: 34. Mexico. 

vexans (Kieffer), 1921h: 110 (Microconops). Algeria. 

whitseli Clastrier and Wirth, 1978: 20. USA (California). 

xuthosceles Chanthawanich and Delfinado, 1967: 299. Thailand. 

yixini Liu, 1997: 40. China. 

yunhsienensis Yu, 1963: 450. China. 

yunnanensis Lee, 1978: 116. China. 

Subgenus LEPTOCONOPS Skuse 

bezzii (Noè), 1905: 114 (Mycterotypus). Italy. 

muganicus Dzhafarov, 1962d: 241 (as subspecies of bezzii; name in Dzhafarov, 1961a: 67 a nomen nudum). 

13

Azerbaijan. 

bidentatus Gutsevich, 1960: 127. Russia. 

boreus Kalugina, 1991: 78. Russia. Upper Cretaceous. 

brasiliensis (Lutz), 1913: 66 (Tersesthes). Brazil. 

bullsbrookensis Smee, 1966: 1008. Australia (Western Australia). 

bundyensis Smee, 1966: 1007. Australia (New South Wales). 

burmiticus Szadziewski, 2004:116. Myanmar. Lower Cretaceous. 

camelorum (Kieffer), 1921h: 111 (Tersesthes). Algeria. 

capensis de Meillon and Hardy, 1953: 24. South Africa. 

carteri Hoffman, 1926b: 133. USA (California). 

casali Cavalieri and Chiossone, 1966: 45. Argentina. 

caucasicus Gutsevich, 1953: 235. Azerbaijan. 

chilensis Forattini, 1958: 38. Chile. 

chinensis Sun, 1968: 2. China. 

clava Borkent, 1997c: 4. Hungary. Upper Cretaceous. 

copiosus Borkent, 1996: 3. USA (New Jersey). Upper Cretaceous. 

curvachelus Borkent, 1996: 7. USA (New Jersey). Upper Cretaceous. 

daugeroni Choufani, Azar and Nel in Choufani et al., 2011: 286. France. Lower Cretaceous. 

demeilloni Clastrier and Nevill, 1984: 245. South Africa. 

endialis Smee, 1966: 1010. Australia (Northern Territory). 

flaviventris Kieffer, 1918a: 34. Tunisia. 

freeborni Wirth, 1952a: 115. USA (California). 

fretus Yu and Zhan, in Wang et al., 1990: 74. China. 

golanensis Clastrier, 1981c: 31. Israel. 

grandis Carter, 1921: 12. Australia (Western Australia). 

halophilus Smee, 1966: 1010. Australia (Western Australia). 

harrisoni de Meillon and Hardy, 1953: 22. South Africa. 

hongkongensis Yu, 2006: 36. China. 

hyalinipennis Kieffer, 1918a: 33. Tunisia. 

indicus (Kieffer), 1918a: 135 (Schizoconops). India. 

irritans (Noè), 1905: 118 (Mycterotypus). Italy. 

longipalpis (Kieffer), 1923a: 657 (Microconops). Algeria. 

inopinatus Huttel and Huttel, 1952b: 45. France. 

kinmenensis Lien, Lin, Weng and Chin, 1996: 32. Taiwan. 

lisbonnei Harant and Galan, 1944: 170. France. 

longicauda Yu, 1997: 56. China. 

longicornis Carter, 1921: 11. Australia (Western Australia). 

lucidus Gutsevich, 1964: 192. Kazakhstan. 

mackerrasae Smee, 1966: 1006. Australia (Queensland). 

melanderi Wirth and Atchley, 1973: 27. USA (Washington). 

mesopotamiensis (Patton), 1920: 245 (Tersesthes). Iraq. 

minutus Gutsevich, 1973: 249. Turkmenistan. 

mohavensis Wirth and Atchley, 1973: 29. USA (California). 

montigenus Clastrier, 1981b: 123. Israel. 

myanmaricus Szadziewski, 2004:117. Myanmar. Lower Cretaceous. 

nicolayi de Meillon, 1937b: 330. Democratic Republic of the Congo. 

nigripes Dzhafarov, 1961a: 68. Azerbaijan. 

nipponensis Tokunaga, 1937a: 262. Japan. 

oshimaensis Takaoka and Hayashi, 1977: 385. Japan. 

nivalis Smee, 1966: 1012. Australia (New South Wales). 

noei Clastrier and Coluzzi, 1973: 59. Italy. 

nosopheris Poinar, 2008: 468. Myanmar. Early Cretaceous. 

obscurus Smee, 1966: 1014. Australia (New South Wales). 

panamensis Ronderos and Spinelli, 1993: 308. Panama. 

14

parvichelus Chanthawanich and Delfinado, 1967: 297. Malaysia. 

petrocchiae Shannon and Del Ponte, 1927: 734. Argentina. 

primaevus Borkent, 1995: 30. Canada (Alberta). Upper Cretaceous. 

rhodesiensis Carter, 1921: 14. Zambia. 

ricardoi Ronderos and Spinelli, 1992: 43. Argentina. 

riverinaensis Smee, 1966: 1009. Australia (Australian Capital Territory). 

rossi Szadziewski, 2004:118. Myanmar. Lower Cretaceous. 

rovnensis Sontag and Szadziewski, 2011: 780. Ukraine. Eocene. 

siamensis Carter, 1921: 20. Thailand. 

leptorhynchus Chanthawanich and Delfinado, 1967: 295. Malaysia. 

maai Tokunaga, 1963a: 214. Indonesia. 


smeei Wirth and Atchley, 1973: 22. New name for longipalpis. 

longipalpis Smee, 1966: 1007 (preoccupied by Leptoconops longipalpis (Kieffer), 1923a). Australia (Western 

Australia). 

stygius Skuse, 1889: 288. Australia (New South Wales). 

subrossicus Szadziewski and Poinar, 2005: 359. Myanmar. Lower Cretaceous. 

succineus Szadziewski, 1988: 233. Denmark. Eocene. 

taiwanensis Lien, Lin and Weng, 1998a: 39. Taiwan. 

torrens (Townsend), 1893: 371 (Tersesthes). USA (New Mexico). 

venezuelensis Ortiz, 1952a: 165. Venezuela. 

wehaiensis Yu and Xue, in Yu, 1988: 132. China. 

woodhilli Lee, 1948a: 334. Australia (Northern Territory). 

zherikhini Szadziewski and Arillo, 2003:273. Spain. Lower Cretaceous. 

Subgenus MEGACONOPS Wirth and Atchley 

MEGACONOPS Wirth and Atchley, 1973: 18 (as subgenus of Leptoconops). Type species: Leptoconops floridensis Wirth, 

by original designation. 

floridensis Wirth, 1951c: 282. USA (Florida). 

Subgenus PALAEOCONOPS Borkent 

PALAEOCONOPS Borkent, 2001: 3 (as subgenus of Leptoconops). Type species: Leptoconops amplificatus Borkent, by 

original designation. 

amplificatus Borkent, 2001: 3. Lebanon. Lower Cretaceous. 

antiquus Borkent, 2001: 6. Lebanon. Lower Cretaceous. 

Subgenus PROLEPTOCONOPS Clastrier 

PROLEPTOCONOPS Clastrier, 1974a: 231 (as subgenus of Leptoconops). Type species: Leptoconops hutsoni Clastrier, by 


aviarum Gutsevich, 1973: 245. Tajikistan. 

bahreinensis Clastrier and Boorman, 1987: 297. Bahrain. 

bailangensis Liu and Yu, in Yu et al., 2005: 109. China. 

dixi de Meillon, 1936: 141. Namibia. 

fuscipennis Clastrier, Rioux and Descous, 1961: 93. Chad. 

hutsoni Clastrier, 1974a: 232. Algeria. 

turkmenicus Molotova, 1967: 627. Turkmenistan. 

werneri Wirth and Atchley, 1973: 34. USA (California). 

15

Subgenus STYLOCONOPS Kieffer 

STYLOCONOPS Kieffer, 1921h: 107. Type species: Leptoconops albiventris de Meijere, by original designation. 

ACANTHOCONOPS Carter, 1921: 24. Type species: Acanthoconops spinosifrons Carter, by original designation. 

albiventris de Meijere, 1915: 98. Indonesia. 

spinosipes Kieffer, 1917a: 190. Papua New Guinea. 

hamariensis Herzi and Sabatini, 1983: 68. Somalia. 

mooloolabaensis (Smee), 1966: 1021 (Styloconops). Australia (Queensland). 

myersi (Tonnoir), 1924: 443 (Acanthoconops). New Zealand. 

pseudosetosifrons (Smee), 1966: 1023 (Styloconops). Australia (Northern Territory). 

setosifrons (Smee), 1966: 1022 (Styloconops). Australia (Queensland). 

spinosifrons (Carter), 1921: 24 (Acanthoconops). Tanzania. 

australiensis (Lee), 1948a: 337 (Styloconops). Australia (New South Wales). 

luteigaster (Senior-White), 1929: 1 (Tersesthes). India. 

yalongensis Yu and Wang, in Yu, 1988: 132. China. 

Genus MINYOHELEA Borkent 

MINYOHELEA Borkent, 1995: 68. Type species: Minyohelea pumilis Borkent, by original designation. 

LEBANOCONOPS Szadziewski, 1996: 40. Type species: Lebanoconops lebanicus Szadziewski, by original designation. 

bacula Borkent, 2000a: 386. Lebanon. Lower Cretaceous. 

casca Borkent, 1997c: 2. Austria. Lower Cretaceous. 

falcata Borkent, 2000a: 385. Lebanon. Lower Cretaceous. 

lebanicus (Szadziewski), 1996: 42 (Lebanoconops). Lebanon. Lower Cretaceous. 

minutus (Szadziewski), 1996: 43 (Lebanoconops). Lebanon. Lower Cretaceous. 

pumilis Borkent, 1995: 69. Canada (Alberta). Upper Cretaceous. 

schleei Szadziewski, 1996: 45. Lebanon. Lower Cretaceous. 

wirthi (Szadziewski), 1996: 44 (Lebanoconops). Lebanon. Lower Cretaceous. 

SUBFAMILY FORCIPOMYIINAE LENZ, 1934: 96 

Genus ATRICHOPOGON Kieffer 

ATRICHOPOGON Kieffer, 1906a: 53 (as subgenus of Ceratopogon). Type species: Ceratopogon exilis Coquillett (= 

Ceratopogon levis Coquillett), designation by Coquillett, 1910: 512. 

KEMPIA Kieffer, 1913d: 162, 179 (as subgenus of Dasyhelea). Type species: Dasyhelea calcuttensis Kieffer, designation by 

Wirth, 1952a: 117. 

GYMNOHELEA Kieffer, 1921h: 115. Type species: Kempia longiserrus Kieffer, designation by Wirth, 1973: 348. 

DOLICHOHELEA Edwards, 1929b: 8. Type species: Dolichohelea polita Edwards, by monotypy. 

LOPHOMYIDIUM Cordero, 1929: 94. Type species: Lophomyidium uruguayense Cordero, by original designation. 

PSILOKEMPIA Enderlein, 1936: 49. Type species: Kempia appendiculata Goetghebuer, by monotypy. 

MELOEHELEA Wirth, 1956b: 16 (as subgenus of Atrichopogon). Type species: Atrichopogon meloesugans Kieffer (= 

Atrichopogon winnertzi Goetghebuer), by original designation. 

PSAMMOPOGON Remm, 1979b: 57 (as subgenus of Atrichopogon). Type species: Atrichopogon trifasciata Kieffer (= 

Atrichopogon flavolineatus Strobl), by original designation. 

ROSTROPOGON Remm, 1979b: 57 (as subgenus of Atrichopogon). Type species: Ceratopogon rostratus Winnertz, by 


IMPENSUKEMPIA Yu, in Liu et al., 2001a: 47 (as subgenus of Atrichopogon). Type species: Atrichopogon impensus, by 


BESSAMYIA Yu, Liu, Liu, Liu, Hao, Yan and Zhao, 2006: 355 (as subgenus of Atrichopogon). Type species: Atrichopogon 

bessa Yu and Yan, by original designation. 

16

abrasus Edwards, 1928: 52. Western Samoa. 

abyssiniae Kieffer, 1918a: 44. Ethiopia. 

acanthocolpus Ingram and Macfie, 1922: 260. Nigeria. 

acosmetus Ingram and Macfie, 1922: 263. Ghana. 

adamsoni Macfie, 1937a: 4. Trinidad and Tobago. 

aereum Yu and Yan, in Yu et al., 2005: 357. China. 

aethiops (Goetghebuer), 1920: 33 (Kempia). Belgium. 

africanus Ingram and Macfie, 1921: 334. Ghana. 

akizukii Tokunaga, 1940b: 274 (as akisukii). Russia. 

alainus Remm, 1980: 110. Kyrgyzstan. 

albinensis Ingram and Macfie, 1931a: 228. Argentina. 

albiscapulus Kieffer, 1918a: 42. Algeria. 

bilineatus Kieffer, 1922g: 498. Algeria. 

alticola (Kieffer), 1911b: 323 (Forcipomyia). India. 

altivolans Macfie, 1949: 113. Mexico. 

alveolatus Nielsen, 1951: 26. Denmark. 

anemotis Kieffer, 1913e: 7. Kenya. 

annulifemoratus Tokunaga, 1959: 190. Papua New Guinea. 

appendiculatus (Goetghebuer), 1920: 37 (Kempia). Belgium. 

armativentris (Kieffer), 1923a: 673 (Kempia). Algeria. 

adjacens (Kieffer), 1924b: 401 (Kempia, as variety of appendiculatus). France. 

aquilonarius Yu and Yan, in Yu et al., 2005: 379. China. 

arabicus Szadziewski, Gwizdalska-Kentzer and Gilka, 2011: 637. United Arab Emirates. 

archboldi Wirth, 1994a: 27. USA (Florida). 

arciforceps Tokunaga, 1941a: 110. Micronesia. 

arcticus (Coquillett), 1900: 396 (Ceratopogon). USA (Alaska). 

argus Kieffer, 1916a: 86. Taiwan. 

argutus Liu, Yan and Liu, 1996: 23. China. 

aridus Spinelli and Marino, in Spinelli et al., 2006: 304. Argentina. 

armaticaudalis Tokunaga and Murachi, 1959: 131. Micronesia. 

armatilabrum Clastrier, 1987b: 211. New Caledonia (France). 

assuetus Macfie, 1934c: 185 (1934d: 212). Malaysia. 

aterrimus Kieffer, 1913d: 178. India. 

atratus Kieffer, 1917a: 184. Australia (New South Wales). 

atribarbus Kieffer, 1922c: 232. Germany. 

atricollis (Goetghebuer), 1935d: 163 (Kempia). Democratic Republic of the Congo. 

atriscapulus Kieffer, 1918a: 45. Tunisia. 

atromaculatus Goetghebuer, 1933b: 111. Russia. 

atroscutellatus Edwards, 1928: 53. Western Samoa. 

atroxipes Bose and Das Gupta, in Bose et al., 2003: 261. India. 

attentus Johannsen, 1932: 416. Indonesia. 

asuturus Borkent and Picado, 2004: 29. Costa Rica. 

auricoma Kieffer, 1917b: 300. Colombia. 

australis Clastrier, 1987b: 200. New Caledonia (France). 

badiensis Clastrier, 1959b: 354. Senegal. 

bai Remm, 1980: 114. Turkmenistan. 

bakeri Macfie, 1939a: 88. Uganda. 

balseiroi Spinelli, 1982: 206. Argentina. 

bangqiensis Yan, Zhang and Yu, 1995: 100. China. 

barbatus Borkent and Picado, 2004: 23. Costa Rica. 

bargaensis Remm, 1972: 70. Russia. 

baripenis Yu and Yan, in Yu et al., 2005: 365. China. 

beccus Borkent and Picado, 2004: 31. Costa Rica. 

17

ellicosus Debenham, 1973: 74. Papua New Guinea. 

bessa Yu and Yan, in Yu et al., 2005: 355. China. 

biangulus Yan, Zhang and Yu, 1995: 96. China. 

bicolor Wirth and Ratanaworabhan, 1993: 328. Malaysia. 

bicuspis Borkent and Picado, 2004: 9. Costa Rica. 

bidaculus Yu and Yan, in Liu, 2001a: 124. China. 

bifasciatus Kieffer, 1917a: 181. Papua New Guinea. 

bifidus Ewen, in Ewen and Saunders, 1958: 694. Brazil. 

binipenis Yu and Yan, in Liu, 2001a: 53. China. 

biroi Kieffer, 1917a: 182. Papua New Guinea. 

bisetosus (Goetghebuer), 1933e: 137 (Kempia). Democratic Republic of the Congo. 

boharti Tokunaga, 1962a: 159. Japan. 

borkenti Wirth, 1994a: 25. Canada (British Columbia). 

brasiliensis Macfie, 1939c: 184. Brazil. 

brenthus Chen, Liu and Yu, 2012: 35. China. 

brevicellula Kieffer, 1921b: 8. Cameroon. 

brevicercus Yan and Yu, 2000: 402 (in Liu, 2001a: 21). China. 

brevicornis Tokunaga, in Tokunaga and Murachi, 1959: 138. Belau (USA). 

brevifurca Goetghebuer, 1948b: 10. Democratic Republic of the Congo. 

brevipalpalis Remm, 1993: 198. Russia. 

nanopalpis Borkent, in Borkent and Wirth, 1997:26. Unnecessary new name for brevipalpalis (as brevipalpis). 

brevipalpis Macfie, 1944b: 298. Trinidad and Tobago. 

brevipenis Yu and Yan, in Liu, 2001a: 67. China. 

breviserrus (Kieffer), 1924b: 400 (Kempia). France. 

brevistilus Kieffer, 1913d: 178. India. 

briani de Meillon, 1961: 45. Madagascar. 

brunnescens Statz, 1944: 141. Germany. Oligocene. 

brunnicellula Clastrier, 1959b: 366. Senegal. 

brunnipes (Meigen), 1804: 29 (Ceratopogon). Europe. 

longinervis Kieffer, 1919a: 28 (as variety of orbicularis). Romania, Slovak Republic. 

semiensis Remm, 1972: 68. Russia. 

bullus Remm, 1980: 114. Turkmenistan. 

calcuttensis (Kieffer), 1913d: 181 (Dasyhelea). India. 

callipotami Macfie, 1924: 65. Egypt. 

capistratus Yu and Yan, in Liu, 2001a: 126. China. 

caribbeanus Ewen, in Ewen and Saunders, 1958: 683. Trinidad and Tobago. 

carinatus Wirth and Ratanaworabhan, 1992: 271. Malaysia. 

carnatus Borkent and Picado, 2004: 13. Costa Rica. 

carpinteroi Marino and Spinelli, 2004c: 157. Argentina. 

casali Cavalieri and Chiossone, 1973: 153. Argentina. 

catinus Yan and Yu, in Liu, 2001a: 69. China. 

cavus Felippe-Bauer, in Felippe-Bauer et al., 2012:45. Brazil. 

celibatus Ingram and Macfie, 1923: 48. South Africa. 

celsus Yu and Yan, in Yu et al., 2005: 368. China. 

characopodus (Speiser), 1910: 749 (Helea). Tanzania. 

chazeaui Clastrier, 1987b: 199. New Caledonia (France). 

chilensis Ingram and Macfie, 1931a: 175. Argentina. 

chrysosphaerotus Ingram and Macfie, 1921: 337. Ghana. 

chuanxiensis Yu and Yan, in Liu, 2001a: 71. China. 

circatheca Bose and Das Gupta, in Bose et al., 2003: 262. India. 

clarusaliger Bose and Das Gupta, in Bose et al., 2003: 264. India. 

clastrieri Spinelli and Marino, 2007: 204. Argentina. 

clavator Debenham, 1973: 73. Papua New Guinea. 

clavifuscus Tokunaga, 1940b: 275. Japan. 

18

colossus Borkent and Picado, 2004: 10. Costa Rica. 

columbianus Kieffer, 1917b: 303. Colombia. 

comatus Bose and Das Gupta, in Bose et al., 2003: 266. India. 

comechingon Spinelli and Marino, in Spinelli et al., 2006: 306. Argentina. 

conglomeratus Kieffer, 1921b: 8. Cameroon. 

conspicuus Clastrier, Rioux and Descous, 1961: 54. Chad. 

coracinus Kieffer, 1917a: 184. Australia (New South Wales). 

corpulentus Ewen, in Ewen and Saunders, 1958: 681. Canada (British Columbia). 

costalis Macfie, 1939c: 179. Brazil. 

costaricae Macfie, 1953: 98. Costa Rica. 

costatus Yu and Yan, in Liu, 2001a: 28. China. 

cretensis Kieffer, 1919a: 28 (as variety of orbicularis). Greece. 

crinitus Ewen, in Ewen and Saunders, 1958: 717. Canada (British Columbia). 

crispantis Liu, Yan and Liu, 1996: 24. China. 

cristatus Hao and Yu, 1998:133. China. 

cryptogamus Macfie, 1939c: 182. Brazil. 

curtipalpis (Kieffer), 1923b: 136 (Kempia). Indonesia. 

dactilus Felippe-Bauer, in Felippe-Bauer et al., 2012:40. Brazil. 

daleyae Giles and Wirth, 1984: 210. Viet Nam. 

dehiscentis Yu and Yan, in Liu, 2001a: 77. China. 

dekeyseri Clastrier, 1959b: 357. Senegal. 

delpontei Cavalieri and Chiossone, 1972: 121. Argentina. 

densiplumus Kieffer, 1918a: 41. Tunisia. 

depilis Macfie, 1939c: 178. Brazil. 

deyrupi Wirth, 1994a: 26. USA (Florida). 

diandrous Yu and Yan, 2010: 200. China. 

didymothecae Macfie, 1953: 99. Costa Rica. 

dilutus Johannsen, 1932: 417. Indonesia. 

discors Macfie, 1934c: 189. Malaysia. 

distictus Kieffer, 1918a: 43. South Africa. 

dominicanus Szadziewski and Grogan, 1998b: 264. Dominican Republic. Miocene. 

domizii Spinelli, 1982: 201. Argentina. 

dorsalis Tokunaga, 1940b: 273. Japan. 

downesi Wirth, 1980: 129. USA (West Virginia). 

dubius Nielsen, 1951: 25. Denmark. 

echinatus Yu and Yan, in Yu et al., 2005: 360. China. 

echinodes Macfie, 1939c: 194. Brazil. 

edentatus Remm, 1972: 67. Russia. 

edwardsi Macfie, 1934c: 184 (1934d: 208). Malaysia. 

elektrophaeus Ingram and Macfie, 1921: 335. Ghana. 

emeiensis Deng, Liao and Yu, in Deng et al., 2011: 115. China. 

endemicus Spinelli and Marino, in Spinelli et al., 2006: 307. Argentina. 

eocenicus Szadziewski, 1988: 223. Poland. Eocene. 

epicautae Wirth, 1956b: 21. USA (Arizona). 

epixanthopygus Yu and Yan, in Yu et al., 2005: 369. China. 

eucnemus Macfie, 1939c: 185. Brazil. 

exiletergitus Yu and Yan, in Yu et al., 2005: 393. China. 

eximiunguis Bose and Das Gupta, in Bose et al., 2003: 267. India. 

ezoensis Tokunaga, 1940b: 268. Japan. 

falcatus Boesel, 1973: 214. USA (Michigan). 

falcis de Meillon and Wirth, 1981c: 571. South Africa. 

farri Wirth, 1956b: 22. USA (Massachusetts). 

femoralis Tokunaga, 1940b: 264. Japan. 

fenestriscutum Tokunaga and Murachi, 1959: 125. Micronesia. 

19

ferenudus Bose and Das Gupta, in Bose et al., 2003: 269. India. 

fiebrigi Kieffer, 1917b: 302. Paraguay. 

fimbriatus Macfie, 1939c: 194 (as variety of gordoni). Brazil. 

fitzroyi Macfie, 1932c: 34. New Zealand. 

flabellis Bose and Das Gupta, in Bose et al., 2003: 269. India. 

flavenicruris Tokunaga and Murachi, 1959: 135. Micronesia. 

flavens Tokunaga, 1940b: 266. Japan. 

flaveolus Zilahi-Sebess, 1936a: 201, 204. Poland. 

flavicaudae Macfie, 1939c: 181. Brazil. 

pilosior Macfie, 1939c: 182 (as variety of flavicaudae). Brazil. 

flaviceps Kieffer, 1912a: 28. Taiwan. 

mendax Kieffer, 1922b: 156 (as variety of flaviceps). Taiwan. 

flavidus Kieffer, 1921g: 562. Philippines. 

flavipalpis Kieffer, 1913d: 175. India. 

flavipes Lutz, 1914: 90. Brazil. 

flaviplumus Remm, 1993: 197. Russia. 

flaviscapus Remm, 1971: 194. Russia. 

flaviscutellum Tokunaga, 1940b: 277. Japan. 

flavitarsatus (Becker), 1903: 74 (Ceratopogon). Egypt. 

flavitergum Tokunaga, 1959: 187. Papua New Guinea. 

flavolineatus (Strobl), 1880: 52 (Ceratopogon). Austria. 

trifasciatus Kieffer, 1918a: 90. Turkey, Greece, Hungary, Slovak Republic, Romania. 

flavus Ewen, in Ewen and Saunders, 1958: 697. Canada (Saskatchewan). 

flumineus Macfie, 1935a: 52. Brazil. 

forcipatus (Winnertz), 1852: 30 (Ceratopogon). Germany. 

silesiacus Kieffer, 1919a: 30. Poland. 

hamifer (Goetghebuer), 1920: 38 (Kempia). Belgium. 

fulvipes Remm, 1972: 71. Turkmenistan. 

fulviscutellaris Tokunaga, 1959: 192. Papua New Guinea. 

fulvus Macfie, 1934c: 186. Malaysia. 

fusciscutellum Wirth and Ratanaworabhan, 1993: 324. Malaysia. 

fusculus (Coquillett), 1901a: 605 (Ceratopogon). USA (New Jersey). 

polydactylus Nielsen, 1951: 27. Denmark. 

fuscus (Meigen), 1804: 28 (Ceratopogon). Europe. 

fuscipes (Zetterstedt), 1850: 3644 (Ceratopogon). Denmark. 

fossicola Kieffer, 1922c: 234. Germany. 

fusinervis (Malloch), 1915a: 308 (Ceratopogon). USA (Illinois). 

gamboai Borkent and Picado, 2004: 26. Costa Rica. 

geminus Boesel, 1973: 211. USA (Ohio). 

ghashmi Boorman and Harten, 2002: 434. Yemen. 

gillipaludosus Remm, 1993: 198. Armenia. 

gilvus (Coquillett), 1905: 62 (Ceratopogon). USA (Florida). 

glaber Macfie, 1935a: 50. Brazil. 

glabricollis (Waltl), 1837 : 279 (Ceratopogon). Germany. 

globosus Kieffer, 1918a: 45. Ethiopia. 

globulifer Macfie, 1939c: 191. Brazil. 

glukhovae Huerta, 2008: 73. Mexico. 

gordoni Macfie, 1938: 164. Trinidad and Tobago. 

gracilis Hao and Yu, 1998:134 (in Liu, 2001a: 80). China. 

granditergitus Borkent and Picado, 2004: 27. Costa Rica. 

granditibialis Borkent and Picado, 2004: 20. Costa Rica. 

grandis Bose and Das Gupta, in Bose et al., 2003: 269. India. 

gressitti Tokunaga, in Tokunaga and Murachi, 1959: 137. Micronesia. 

greyi Macfie, 1932c: 35. New Zealand. 

20

griseolus (Zetterstedt), 1855: 4865 (Ceratopogon). Sweden. 

majusculus Remm, 1961b: 920. Estonia. 

grogani Huerta and Dzul, 2012: 24. Mexico. 

guianensis Macfie, 1940b: 184. Guyana. 

guttatus Yan and Yu, 2000: 403 (in Liu, 2001a: 24). China. 

haemorrhoidalis Kieffer, 1921g: 560. Taiwan. 

haesitans (Kieffer), 1922c: 234 (Gymnohelea). Germany. 

harpagonum Macfie, 1934c: 184 (1934d: 208). Malaysia. 

harrisi Macfie, 1938: 163. Trinidad and Tobago. 

helles de Meillon and Wirth, 1983a: 350. South Africa. 

hesperius Ingram and Macfie, 1922: 266. Ghana. 

hexastichus Nielsen, 1951: 27. Denmark. 

hilaris Bose and Das Gupta, in Bose et al., 2003: 271. India. 

hirsutipennis Ingram and Macfie, 1923: 49. South Africa. 

helion de Meillon, 1936: 177. South Africa. 

hirtidorsum Remm, 1961b: 922. Estonia. 

hispaniae Havelka, 1979: 58. Spain. 

hobsoni Macfie, 1932c: 32. New Zealand. 

homoius Ingram and Macfie, 1921: 338. Ghana. 

alfierii Kieffer, 1925e: 249. Egypt. 

homofacies Spinelli, in SpinelliBose, 1989: 734. Argentina. 

horni Kieffer, 1925a: 408. Sri Lanka. 

horos Yan and Yu, in Yu et al., 2005: 428. China. 

hortensis Sahuquillo and Gil Collado, 1982a: 305 (as subspecies of minutus). Spain. 

hukengicus Yu, Huang and Yang, in Huang et al., 2009:365. China. 

humicola Ewen, in Ewen and Saunders, 1958: 679. Canada (Saskatchewan). 

hystricoides Debenham, 1973: 71. Papua New Guinea. 

ilonae Gosseries, 1989: 2. New name for pilosipennis. 

pilosipennis Tokunaga, 1940b: 276 (preoccupied by Forcipomyia pilosipennis (Kieffer), 1919a). Japan. 

impensus Yu and Yan, in Liu, 2001a: 50. China. 

inacayali Spinelli and Marino, in Spinelli et al., 2006: 309. Argentina. 

inconspicuus Ewen, in Ewen and Saunders, 1958: 706. Canada (Saskatchewan). 

incultus Ewen, in Ewen and Saunders, 1958: 699. Costa Rica. 

indianus (Kieffer), 1910: 185 (Ceratopogon). India. 

rivicola (Kieffer), 1911b: 324 (Forcipomyia). India. 

infamis Bose and Das Gupta, in Bose et al., 2003: 273. India. 

infuscus Goetghebuer, 1929: 232. Belgium. 

insignipalpis Macfie, 1940a: 73. Brazil. 

insigniunguis Clastrier, 1987b: 198. New Caledonia (France). 

insigniventris Macfie, 1935a: 51. Brazil. 

insolens Bose and Das Gupta, in Bose et al., 2003: 274. India. 

insolitipes Bose and Das Gupta, in Bose et al., 2003: 273. India. 

insularis Kieffer, 1921g: 561. Taiwan. 

citrinipes Kieffer, 1922b: 154. Taiwan. 

intertextus Yu and Yan, in Yu et al., 2005: 443. China. 

isis Kieffer, 1925e: 252. Egypt. 

isolatus Liu, Yan and Liu, 1996: 24. China. 

jacobsoni (de Meijere), 1907: 212 (Ceratopogon). Indonesia. 

flavellus Kieffer, 1913d: 177. India. 

immaculatus Kieffer, 1917a: 181. Papua New Guinea. 

cavernarum Edwards, 1924: 107. India. 

rarus Johannsen, 1946: 188. Guam (USA). 

jamnbacki Wirth, 1994a: 24. USA (New York). 

japonicus Tokunaga, 1940b: 272. Japan. 

21

jejunus Macfie, 1934c: 184 (1934d: 211). Malaysia. 

jianfengensis Liu and Yan, in Liu, 2001a: 86. China. 

jubacaudalis Yu and Yan, in Yu et al., 2005: 445. China. 

kagiensis Tokunaga, 1940b: 270. Taiwan. 

kangnani Yan, Zhang and Yu, 1995: 99. China. 

kelainosoma Ingram and Macfie, 1922: 261. Ghana. 

kribiensis Kieffer, 1921b: 10. Cameroon. 

kyotoensis Tokunaga, 1940b: 275. Japan. 

lacajae Macfie, 1953: 100. Costa Rica. 

lacustris Clastrier, 1987b: 203. New Caledonia (France). 

ladislavi Tóthová, in Tóthová et al., 2009. Canada. 

lamellamarsipos Yu and Yan, in Yu et al., 2005: 345. China. 

lampronotus (Kieffer), 1911c: 335 (Ceratopogon). Seychelles. 

largipenis Yan, Zhang and Yu, 1995: 98. China. 

lassus Yan, Zhang and Yu, 1995: 97. China. 

latipygus Vaillant, 1957: 270. Algeria. 

lazoensis Remm, 1971: 197. Russia. 

levis (Coquillett), 1901a: 604 (Ceratopogon). USA (Maryland). 

exilis (Coquillett), 1902a: 86 (Ceratopogon). USA (District of Columbia). 

lindneri Wirth, 1964: 2. Tanzania. 

lituratus (Williston), 1896: 281 (Ceratopogon). St. Vincent. 

lobatus Borkent and Picado, 2004: 14. Costa Rica. 

longicalcar Remm, 1961b: 925. Estonia. 

longicalcaris Tokunaga, 1959: 182. Papua New Guinea. 

longicornis Ewen, in Ewen and Saunders, 1958: 700. Costa Rica. 

longicostus Clastrier, 1959b: 360. Senegal. 

longipalpis (Kieffer), 1923b: 135 (Kempia). Indonesia. 

longiserrus (Kieffer), 1921a: 53 (Kempia). Poland. 

longitergitus Yan and Yu, in Liu et al., 2001a: 26. China. 

lucorum (Meigen), 1818: 72 (Ceratopogon). Europe. 

sylvaticus (Winnertz), 1852: 29 (Ceratopogon). Germany. 

setosipennis (Kieffer), 1911d: 3 (Forcipomyia). Germany. 

bidentatus Kieffer, 1924b: 399 (as variety of winnertzi). France. 

seminitidus Goetghebuer, 1947: 229. Belgium. 

ludingensis Yu and Yan, in Liu et al., 2001a: 90. China. 

luteicollis (Becker), 1903: 74 (Ceratopogon). Egypt. 

flavoscutellatus (Becker), 1908: 74 (Ceratopogon, preoccupied by Dasyhelea flavoscutellata (Zetterstedt), 1850). 

Canary Islands (Spain). 

aegyptius Kieffer, 1925e: 250. Egypt. 

phrixus de Meillon, 1943: 105. South Africa. 

sanani Boorman and Harten, 2002: 434. Yemen. 

luteipes Goetghebuer, 1933e: 140. Democratic Republic of the Congo. 

lutescens Clastrier, Rioux and Descous, 1961: 59. Chad. 

lyratus Yu and Yan, in Yu et al., 2005: 448. China. 

macrodentatum Marino, Tóthová and Spinelli, 2011: 62. Argentina. 

maculatus (Lundström), 1910: 35 (Ceratopogon, as variety of minutus). Finland. 

avastensis Remm, 1959: 688. Estonia. 

hamulatus Remm, 1971: 197. Russia. 

maculipennis Clastrier, 1968: 89. French Guiana. 

maculosus Ewen, in Ewen and Saunders, 1958: 689. Canada (Saskatchewan). 

magnus Borkent and Picado, 2004: 16. Costa Rica. 

marginipilus Tokunaga, 1940b: 267. Japan. 

maritimus Tokunaga, 1940b: 270. Japan. 

mastersi (Skuse), 1889: 297 (Ceratopogon). Australia (New South Wales). 

22

matilei Clastrier, 1987b: 204. New Caledonia (France). 

medicrinis Yu and Yan, in Yu et al., 2005: 449. China. 

megalothecus Remm, 1971: 197. Russia. 

melancholicus (Meigen), 1838': 18 (Ceratopogon). Europe. 

melanimus Ingram and Macfie, 1923: 50. South Africa. 

melanoticus Macfie, 1934c: 187 (as variety of subfuscus). Malaysia. 

melinois (Speiser), 1910: 751 (Helea, as melinoessa). Tanzania. 

meloesugans Kieffer, 1922g: 495. Algeria. 

mendozae Ingram and Macfie, 1931a: 229. Argentina. 

mexicanus Huerta, 2001: 373. Mexico. 

minimus Kieffer, 1916a: 87. Taiwan. 

minutalatus Bose and Das Gupta, in Bose et al., 2003: 275. India. 

minutus (Meigen), 1830: 263 (Ceratopogon). Europe. 

fortiserrus Kieffer, 1924b: 398. France. 

turficola Kieffer, 1925b: 149. Estonia. 

parviforceps Tokunaga, 1940b: 267. Russia. 

miripalpis Kieffer, 1924b: 397. France. 

modestus Bose and Das Gupta, in Bose et al., 2003: 275. India. 

monomorphicus Marino, Tóthová and Spinelli, 2011: 65. Argentina. 

monticola Tokunaga, 1940b: 271. Taiwan. 

montigenum Yu and Yan, in Liu et al., 2001a: 92. China. 

montivagus (Kieffer), 1911b: 322 (Forcipomyia). India. 

montium Storå, 1936: 34. Canary Islands (Spain). 

muelleri (Müller), 1905: 224 (Ceratopogon). Poland. 

muelleri (Kieffer, 1906b: 336) (Ceratopogon, preoccupied by Atrichopogon muelleri (Müller), 1905). Germany. 

cornutus Nielsen, 1951: 25. Denmark. 

globularis Mayer, 1934d: 214 (as variety of trifasciatus). Sweden. 

multidens Bose and Das Gupta, in Bose et al., 2003: 277. India. 

multiplex Bose and Das Gupta, in Bose et al., 2003: 279. India. 

multispinosa Bose and Das Gupta, in Bose et al., 2003: 280. India. 

myrmedon Kieffer, 1922b: 155. Taiwan. 

quadrispinosus Remm, 1993: 196. Russia. 

pectinacaudalis Yu and Yan, in Yu et al., 2005: 457. China. 

nanus Macfie, 1940b: 184. Guyana. 

natalensis Ingram and Macfie, 1923: 52. South Africa. 

natans Kieffer, 1922g: 496. Algeria. 

nebulosus Macfie, 1939c: 192. Brazil. 

nemestrinus (Santos Abreu), 1918: 257 (Helea). Canary Islands (Spain). 

albidipes (Santos Abreu), 1918: 259 (Helea, as variety of nemestrina). Canary Islands (Spain). 

distinctus (Santos Abreu), 1918: 260 (Helea, as variety of nemestrina). Canary Islands (Spain). 

flavicans (Santos Abreu), 1918: 259 (Helea, as variety of nemestrina). Canary Islands (Spain). 

flavihalteratus (Santos Abreu), 1918: 259 (Helea, as variety of nemestrina). Canary Islands (Spain). 

fulviventris (Santos Abreu), 1918: 260 (Helea, as variety of nemestrina). Canary Islands (Spain). 

neocaledoniensis Clastrier, 1987b: 205. New Caledonia (France). 

nielamuensis Yu and Yan, in Liu et al., 2001a: 130. China. 

nigeriae Clastrier and Wirth, 1961a: 200. Nigeria. 

nigribasalis Tokunaga, 1959: 180. Papua New Guinea. 

nigripes Macfie, 1934c: 188. Malaysia. 

nigritellus Yu and Yan, in Yu et al., 2005: 406. China. 

nigrithoracius Tokunaga, 1959: 188. Papua New Guinea. 

nigrithorax Clastrier, 1987b: 207. New Caledonia (France). 

nigrofuscus Remm, 1980: 115. Kyrgyzstan. 

nigromicans Goetghebuer, 1935d: 162. Democratic Republic of the Congo. 

nilicola Kieffer, 1921b: 9. Sudan. 

23

niloticus Kieffer, 1921b: 9 (as variety of conglomeratus). Sudan. 

nilssoni de Meillon and Wirth, 1989a: 88. Madagascar. 

novaeteutoniae Macfie, 1939c: 189. Brazil. 

novaguinensis Tokunaga, 1959: 202. Indonesia. 

nubeculosus Macfie, 1949: 113. Mexico. 

nudus Zilahi-Sebess, 1940: 38. Hungary. 

obesus Goetghebuer, 1948b: 11. Democratic Republic of the Congo. 

obfuscatus Ingram and Macfie, 1931a: 173. Argentina. 

obnubilus Ingram and Macfie, 1931a: 175. Argentina. 

assimilis Ingram and Macfie, 1931a: 176. Argentina. 

obscuripes Macfie, 1933a: 77. French Polynesia (France). 

obscurus Ewen, in Ewen and Saunders, 1958: 685. Puerto Rico (USA). 

occidentalis Wirth, 1952a: 120. USA (California). 

ochrosoma (Ingram and Macfie), 1921: 340 (Kempia). Ghana. 

ocumare (Ortiz), 1952b: 254 (Monohelea). Venezuela. 

oedemerarum Storå, 1939: 16. Finland. 

okinawensis Tokunaga, 1962a: 165. Japan. 

ollicula Yan and Yu, in Liu et al., 2001a: 41. China. 

orbicularis Kieffer, 1919a: 28. Romania. 

orbitus Yu and Yan, in Yu et al., 2005: 370. China. 

origenus Kieffer, 1913d: 176. India. 

oriphilus (Kieffer), 1911b: 324 (Forcipomyia). India. 

ornatipennis Clastrier, 1987a: 271. French Guiana. 

ornatithorax Clastrier, 1987b: 208. New Caledonia (France). 

ornativentris Clastrier, 1959b: 363. Senegal. 

osiris Kieffer, 1925e: 250. Egypt. 

oviformis Liu, Yan and Liu, 1996: 24. China. 

pachito Huerta and Dzul, 2012: 22. Mexico. 

pachycnemus Macfie, 1953: 101. Costa Rica. 

pacificus Clastrier, 1987b: 210. New Caledonia (France). 

pallidicillus Yu and Zou, 1988: 85. China. 

pallidipedis Tokunaga, 1959: 194. Papua New Guinea. 

pallidipes Kieffer, 1917b: 301. Paraguay. 

palmatus Tokunaga, 1962a: 160. Japan. 

sentus Yu and Qi, in Liu et al., 1990: 219 (also as snetus and sentns). China. 

palpalis Macfie, 1939c: 196. Brazil. 

palus Bose and Das Gupta, in Bose et al., 2003: 282. India. 

parroti Clastrier, 1956: 510. Algeria. 

parvulus Kieffer, 1919a: 25. Ukraine. 

pastinaca Yu, in Yu et al., 2005: 362. China. 

paulus Remm, 1961b: 928. New name for nanus. 

nanus Remm, 1959: 686 (preoccupied by Atrichopogon nanus Macfie, 1940b). Estonia. 

pavidus (Winnertz), 1852: 33 (Ceratopogon). Germany. 

pollinivorus Downes, 1955: 442. Great Britain. 

pecteniventris Borkent, in Borkent and Wirth, 1997:27. New name for armativentris. 

armativentris (Kieffer), 1923b: 137 (Kempia, preoccupied by Atrichopogon armativentris (Kieffer), 1923a). Indonesia. 

pectinatus Macfie, 1939c: 195. Brazil. 

peculiaris Bose and Das Gupta, in Bose et al., 2003: 284. India. 

pedipalens Yan and Yu, 1999: 100. China. 

penicillatus Delécolle and Rieb, 1994: 274. Guadeloupe. 

peregrinus (Johannsen), 1908: 266 (Ceratopogon). USA (New York). 

perfuscus Ingram and Macfie, 1921: 337. Ghana. 

perplexus (Kieffer), 1913d: 182 (Dasyhelea). India. 

peruvianus Kieffer, 1917b: 302. Peru. 

24

petrosus Remm, 1980: 111. Kyrgyzstan. 

phusunensis Remm, 1971: 192. Russia. 

piceiventris Kieffer, 1917b: 301. Paraguay. 

picipes Goetghebuer, 1935d: 163. Democratic Republic of the Congo. 

pictipennis Clastrier, 1979: 30. French Guiana. 

pileolus Yu and Yan, in Liu et al., 2001a: 38. China. 

planetus Yu and Liu, 1995: 47. China. 

planusunguis Bose and Das Gupta, in Bose et al., 2003: 285. India. 

poguei Clastrier, 1987b: 201. New Caledonia (France). 

politus (Edwards), 1929b: 9 (Dolichohelea). Philippines. 

postremus (Santos Abreu), 1918: 269 (Helea). Canary Islands (Spain). 

pruinosus Kieffer, 1921g: 560. Taiwan. 

psilopterus Kieffer, 1919a: 26. Hungry, Croatia, Greece. 

pterygospinous Yu and Yan, in Liu et al., 2001a: 138. China. 

pudicus Johannsen, 1932: 418. Indonesia. 

pullatus Macfie, 1933b: 95. French Polynesia (France). 

pusillus (Kieffer), 1921b: 6 (Kempia). Cameroon. 

quadrisetosus Goetghebuer, 1933e: 141. Democratic Republic of the Congo. 

quartibrunneus Borkent and Picado, 2004: 30. Costa Rica. 

quasicomatus Bose and Das Gupta, in Bose et al., 2003: 285. India. 

quateriharpagonum Tokunaga, 1959: 198. Indonesia. 

qudratepenis Yu and Yan, in Yu et al., 2005: 411. China. 

raripilipennis Tokunaga and Murachi, 1959: 123. Belau (USA). 

redactus Borkent and Picado, 2004: 32. Costa Rica. 

remigatus Ewen, in Ewen and Saunders, 1958: 687. Brazil. 

rhiphidus Yu and Yan, in Yu et al., 2005: 459. China. 

rhynchops (Schiner), 1868: 26 (Ceratopogon). Australia (New South Wales). 

rictus Yu and Yan, in Yu et al., 2005: 338. China. 

rivalis Yu and Yan, in Liu et al., 2001a: 99. China. 

rostratus (Winnertz), 1852: 31 (Ceratopogon). Poland. 

transversalis Kieffer, 1918a: 91. Turkey, Hungary, Slovak Republic, Romania, Italy. 

ventralis Kieffer, 1918a: 91. Turkey, Hungary, Ukraine. 

homopterus Kieffer, 1919a: 29 (as variety of transversalis). Hungary, Turkey. 

coracellus Kieffer, 1919a: 30 (as variety of ventralis). Hungary, Poland, Ukraine, Greece. 

nigriventris Kieffer, 1919a: 30 (as variety of ventralis). Slovak Republic. 

putredinis Kieffer, 1922c: 233. Germany. 

rotundus Liu, Yan and Liu, 1996: 25. China. 

ruber Kieffer, 1916a: 86. Taiwan. 

formosanus Kieffer, 1918a: 89. Taiwan. 

rubidus Macfie, 1933b: 97. French Polynesia (France). 

rufescens Kieffer, 1921g: 563. Taiwan. 

rufiventris Kieffer, 1917a: 184 (as variety of atratus). Australia (New South Wales). 

rusticus Macfie, 1939c: 190. Brazil. 

ryukyuensis Tokunaga, 1962a: 164. Japan. 

sachalinensis Remm, 1993: 198. Russia. 

salisburiensis de Meillon, 1959a: 337. Zimbabwe. 

sallami Boorman and Harten, 2002: 433. Yemen. 

sanctaeclarae Macfie, 1949: 114. Mexico. 

sanctilaurentii Kieffer, 1917b: 303. Paraguay. 

saundersi Ewen, in Ewen and Saunders, 1958: 691. Puerto Rico (USA). 

schizonyx Giles and Wirth, 1982b: 825. Sri Lanka. 

scutatis Yu and Yan, in Liu et al., 2001a: 56. China. 

scutellaris Goetghebuer, 1933e: 140. Democratic Republic of the Congo. 

sebessi Remm, 1981: 32. New name for biroi. 

25

iroi Zilahi-Sebess, 1941: 83 (preoccupied by Atrichopogon biroi Kieffer, 1917a). Tunisia. 

semipilosus (Kieffer), 1912c: 5 (Forcipomyia). Sri Lanka. 

sequax (Williston), 1896: 282 (Ceratopogon). St. Vincent. 

serrulatus Kieffer, 1924b: 399. France. 

sessilis Kieffer, 1917a: 183. Papua New Guinea. 

setosicubitus Clastrier, 1987b: 210. New Caledonia (France). 

setosilateralis Borkent and Picado, 2004: 33. Costa Rica. 

setosus Kieffer, 1919c: 193. New name for setosipennis. 

setosipennis Kieffer, 1913d: 178 (preoccupied by Atrichopogon setosipennis (Kieffer), 1911d). India. 

seudoobfuscatus Spinelli, 1982: 208. Argentina. 

shortlandi Macfie, 1932c: 33. New Zealand. 

sichotensis Remm, 1971: 195. Russia. 

similis Spinelli and Marino, in Spinelli et al., 2006: 316. Argentina. 

simplex Bose and Das Gupta, in Bose et al., 2003: 287. India. 

simplicifurcatus Tokunaga, 1959: 201. Indonesia. 

singularis (Kieffer), 1921a: 52 (Kempia). Poland. 

sinuosus Wirth and Ratanaworabhan, 1992: 272. Malaysia. 

snyderi Tokunaga, in Tokunaga and Murachi, 1959: 117. Japan. 

solivagus Yu and Yan, in Yu et al., 2005: 373. China. 

sordidus Bose and Das Gupta, in Bose et al., 2003: 288. India. 

spadix Bose and Das Gupta, in Bose et al., 2003: 289. India. 

spartos Yan and Yu, in Liu et al., 2001a: 45. China. 

speculiger Nielsen, 1951: 25. Denmark. 

sphagnalis (Kieffer), 1925a: 408 (1927: 59) (Kempia). Estonia. 

spinicaudalis Tokunaga, in Tokunaga and Murachi, 1959: 129. Belau (USA). 

spiniventris Tokunaga, 1940b: 273. Japan. 

spinosus Borkent and Picado, 2004: 12. Costa Rica. 

spurius Kieffer, 1913d: 177. India. 

stannusi Ingram and Macfie, 1924c: 181. Ghana. 

subcomatus Bose and Das Gupta, in Bose et al., 2003: 289. India. 

subfuscus Macfie, 1934c: 187. Malaysia. 

sublimatus Kieffer, 1913d: 176. India. 

subtenuiatus Yu and Yan, in Liu et al., 2001a: 142. China. 

suburbanus Liu, Yan and Liu, 1996: 25. China. 

sulfuratus Debenham, 1989: 229. New name for flaveolum. 

flaveolum Tokunaga, 1959: 185 (preoccupied by Atrichopogon flaveolus Zilahi-Sebess, 1936a). Papua New Guinea. 

sumatrae Macfie, 1934c: 185 (1934d: 212). Malaysia. 

taeniatus Macfie, 1939c: 186. Brazil. 

taizi Boorman and Harten, 2002: 433. Yemen. 

wirthi Delécolle and Braverman, 1997: 101 (emended from wirthorum) (preoccupied by Atrichopogon wirthi Chan 

and Linley, 1988). Israel. 

shaubensis Boorman and Harten, 2002: 434. Yemen. 

talarum Spinelli, 1982: 204. Argentina. 

tapantiensis Borkent and Picado, 2004: 34. Costa Rica. 

tatricus Remm, 1981: 32. New name for montivagus. 

montivagus Kieffer, 1919a: 25 (preoccupied by Atrichopogon montivagus (Kieffer), 1911b). Slovak Republic. 

tegmentalis Liu, Yan and Liu, 1996: 25. China. 

tenuiatus Tokunaga, 1959: 200. Papua New Guinea. 

tenuidentis Liu, Yan and Liu, 1996: 26. China. 

tenuipalpis Liu, Yan and Liu, 1996: 26. China. 

tenuistylus Bose and Das Gupta, in Bose et al., 2003: 289. India. 

tetramischus Yu and Liu, 1995: 48. China. 

thersites (Williston), 1896: 280 (Ceratopogon). St. Vincent. 

thienemanni Kieffer, 1921a: 54. Germany. 

26

nitens Kieffer, 1921a: 54 (as variety of thienemanni). Germany. 

tirzae Borkent and Picado, 2004: 28. Costa Rica. 

titanus Boesel, 1973: 213. USA (Ohio). 

transiens (Walker), 1848: 25 (Ceratopogon). Canada. 

transversus Wirth, 1952a: 124. USA (California). 

trichopus (Thomson), 1868: 444 (Ceratopogon). China. 

trichotomma (Kieffer), 1911c: 332 (Ceratopogon). Seychelles. 

tricleaves Liu, Yan and Liu, 1996: 27. China. 

tridentistylus Tokunaga, 1959: 184. Papua New Guinea. 

trinidadensis Macfie, 1937a: 5. Trinidad and Tobago. 

tritomus Kieffer, 1919a: 24. Hungary. 

tropicus Kieffer, 1913e: 8. Kenya. 

tuberculatus Ewen, in Ewen and Saunders, 1958: 709. Trinidad and Tobago. 

turneri Ingram and Macfie, 1923: 53. South Africa. 

tutatus Yu and Yan, in Yu et al., 2005: 419. China. 

udus Yu and Yan, in Yu et al., 2005: 420. China. 

umbratilis Macfie, 1935a: 51. Brazil. 

umbrosus Macfie, 1933b: 97. French Polynesia (France). 

unguis Tokunaga, 1962a: 162. Japan. 

unilineatus Remm, 1967: 10. Azerbaijan. 

urbicola (Kieffer), 1911b: 325 (Forcipomyia). India. 

uruguayensis (Cordero), 1929: 95 (Lophomyidium). Uruguay. 

utricularis Macfie, 1953: 102. Costa Rica. 

varius Yu and Yan, in Liu et al., 2001a: 58. China. 

vastus Bose and Das Gupta, in Bose et al., 2003: 293. India. 

vepres Debenham, 1973: 73. Australia (Queensland). 

verax Bose and Das Gupta, in Bose et al., 2003: 295. India. 

vesiculosus Macfie, 1934c: 184 (1934d: 209). Malaysia. 

vestitipennis Kieffer, 1917a: 183. Papua New Guinea. 

vicinus (Santos Abreu), 1918: 262 (Helea). Canary Islands (Spain). 

victoriae de Meillon, 1942a: 96. Zimbabwe. 

vittatus Tokunaga, 1959: 196. Papua New Guinea. 

volaticus Yan and Yu, in Yu et al., 2005: 431. China. 

wallisensis Clastrier and Delécolle, 1996: 291. Wallis and Futuna Islands (France). 

warmkei Wirth, 1956c: 243. Puerto Rico (USA). 

websteri (Coquillett), 1901a: 603 (Ceratopogon). USA (Louisiana). 

winnertzi Goetghebuer, 1922: 51. Belgium. 

torgnyensis Goetghebuer, 1949: 2. Belgium. 

wirthi Chan and Linley, 1988: 189. USA (Florida). 

woodfordi Macfie, 1938: 162. Trinidad and Tobago. 

wuyishanicus Yu and Huang, in Huang et al., 2009:366. China. 

xanthoaspidium Carter, Ingram and Macfie, 1921a: 322. Ghana. 

xanthophilus (Kieffer), 1911b: 326 (Forcipomyia). India. 

xanthopus Kieffer, 1913d: 175. India. 

xanthopygus Tokunaga, 1962a: 161. Japan. 

xylochus Yu and Yan, in Yu et al., 2005: 422. China. 

yamabukiensis Clastrier, 1987c: 426. Democratic Republic of the Congo. 

yolancae Borkent and Picado, 2004: 24. Costa Rica. 

yongxinensis Yan and Yu, in Liu et al., 2001a: 117. China. 

yoshimurai Tokunaga, 1940b: 272. Japan. 

zhangmuensis Yu and Yan, in Liu et al., 2001a: 109. China. 

Genus ATRICULICOIDES Remm 

27

ATRICULICOIDES Remm, 1976b: 108. Type species: Atriculicoides macrophthalmus Remm, by original designation. 

cenomanensis Szadziewski and Schlüter, 1992: 74. France. Upper Cretaceous. 

dasyheleis Szadziewski, 1996: 55. Russia. Upper Cretaceous. 

globosus (Boesel), 1937: 47 (Lasiohelea). Canada (Manitoba). Upper Cretaceous. 

creteus (Boesel), 1937: 46 (Lasiohelea). Canada (Manitoba). Upper Cretaceous. 

incompletus Szadziewski and Schlüter, 1992: 76. France. Upper Cretaceous. 

macrophthalmus Remm, 1976b: 108. Russia. Upper Cretaceous. 

squamiciliatus Remm, 1976b: 110. Russia. Upper Cretaceous. 


swinhoei (Cockerell), 1919: 243 (Johannsenomyia). Myanmar. Lower Cretaceous. 

szadziewskii Pérez-de la Fuente, Delclòs, Peñalver and Arillo, 2011: 758. Spain. Lower Cretaceous. 

taimyricus Szadziewski, 1996: 56. Russia. Upper Cretaceous. 

Genus FORCIPOMYIA Meigen 

FORCIPOMYIA Meigen, 1818: 73,75. Type species: Tipula bipunctata Linnaeus, designation by Westwood, 1840: 126. 

Generic name first published in synonymy with Ceratopogon but available under ICZN Code Article 11(e). 

LABIDOMYIA Stephens, 1829a: 52 (1829b: 239) (unnecessary new name for Forcipomyia Meigen, as Forcipomyia 

Megerle). Type species: Tipula bipunctata Linnaeus, designation by Westwood, 1840: 126. 

TETRAPHORA Philippi, 1865: 630. Type species: Tetraphora fusca Philippi, by monotypy. 

PROHELEA Kieffer, 1911b: 319 (as subgenus of Forcipomyia). Type species: Ceratopogon decipiens Kieffer, designation 

by Brunetti, 1920: 48. 

Subgenus ATOPOMYIA Yu and Liu 

ATOPOMYIA Yu and Liu, 2000a:48 (as subgenus of Forcipomyia). Type species: Forcipomyia atopia Yu and Liu, by original 

designation. 

atopia Yu and Liu, 2000a:49. China. 

idaeus Yu and Liu, 2000a:50. China. 

Subgenus BALIOHELEA Yu and Liu 

BALIOHELEA Yu and Liu, 2006: 476 (as subgenus of Forcipomyia). Type species: Forcipomyia martinus Yu and Liu, by 


martina Yu and Liu, in Yu et al., 2005: 476. China. 

Subgenus BASSOFORCIPOMYIA Debenham 

BASSOFORCIPOMYIA Debenham, 1987b: 172 (as subgenus of Forcipomyia). Type species: Forcipomyia centurio 

Debenham, by original designation. 

centurio Debenham, 1987b: 173. Australia (Tasmania). 

inca Debenham, 1987b: 175. Australia (New South Wales). 

Subgenus BLANTONIA Wirth and Dow 

BLANTONIA Wirth and Dow, 1971: 289 (as subgenus of Forcipomyia). Type species: Forcipomyia caribbea Wirth and Dow, 


28

caribbea Wirth and Dow, 1971: 291. Jamaica. 

testudo Debenham, 1987a: 64. Australia (Queensland). 

Subgenus CALOFORCIPOMYIA Saunders 

CALOFORCIPOMYIA Saunders, 1957: 680 (as subgenus of Forcipomyia). Type species: Forcipomyia caerulea Saunders, 


annulipes Tokunaga, 1940c: 72. Japan. 

apicalis Goetghebuer, 1935d: 148. Democratic Republic of the Congo. 

trimaculata Goetghebuer, 1935d: 160. Democratic Republic of the Congo. 

rutshuruensis Goetghebuer, 1948b: 7. Democratic Republic of the Congo. 

auripes Ingram and Macfie, 1924b: 560. Ghana. 

caerulea Saunders, 1957: 681. Brazil. 

campanula de Meillon and Downes, 1986: 142. South Africa. 

coloratus Liu and Yu, in Liu et al., 2001b: 19. China. 

confragosus Liu and Yu, in Liu et al., 2001b: 21. China. 

copanensis Utmar and Wirth, 1976: 129. Honduras. 

eukosma Macfie, 1939c: 148. Brazil. 

furcifera Macfie, 1940e: 920. Brazil. 

gibba Debenham, 1987b: 180. Australia (Queensland). 

glauca Macfie, 1934b: 144. Great Britain. 

monilis Goetghebuer, 1934c: 287. Russia. 

diversipes Goetghebuer, 1936: 319. Belgium. 

splendida Wirth, 1951d: 315. USA (Virginia). 

hatoensis Utmar and Wirth, 1976: 131. Panama. 

hermosa Utmar and Wirth, 1976: 117. Brazil. 

illimis Liu and Yu, in Liu et al., 2001b: 23. China. 

lemuria de Meillon, 1961: 43. Zimbabwe. 

longipalpis Saha, Dasgupta, Gangopadhyay and Mukherjee, 2009: 8. India. 

maculipes (Goetghebuer), 1933e: 135 (Lasiohelea). Democratic Republic of the Congo. 

neocaledoniensis Clastrier and Delécolle, 1991: 183. New Caledonia (France). 

nigrescens Macfie, 1939c: 156. Brazil. 

paraglauca Saha, Dasgupta, Gangopadhyay and Mukherjee, 2009: 9. India. 

pennaticauda Debenham, 1987b: 183. Papua New Guinea. 

quasiglauca Saha, Dasgupta, Gangopadhyay and Mukherjee, 2009: 9. India. 

quokkae Debenham, 1987b: 179. Australia (Western Australia). 

remigera Utmar and Wirth, 1976: 121. Colombia. 

sabalitensis Utmar and Wirth, 1976: 127. Costa Rica. 

scitula Goetghebuer, 1935d: 159. Democratic Republic of the Congo. 

semiglauca Saha, Dasgupta, Gangopadhyay and Mukherjee, 2009: 10. India. 

squamianulipes Tokunaga and Murachi, 1959: 194. Belau (USA). 

subglauca Saha, Dasgupta, Gangopadhyay and Mukherjee, 2009: 10. India. 

swezeyi Edwards, 1928: 50. American Samoa (USA). 

takahashii Tokunaga, 1940c: 88. Taiwan. 

varicolor Saunders, 1957: 683. Brazil. 

viridis Clastrier and Delécolle, 1991: 179. New Caledonia (France). 

Subgenus COLLESSOHELEA Debenham 

COLLESSOHELEA Debenham, 1987a: 60 (as subgenus of Forcipomyia). Type species: Forcipomyia yungurara 

Debenham, by original designation. 

manzhuangensis Liu and Yu, in Liu et al., 2001b: 26. China. 

29

yungurara Debenham, 1987a: 61. Australia (Queensland). 

Subgenus DYCEA Debenham 

DYCEA Debenham, 1987d: 666 (as subgenus of Forcipomyia). Type species: Forcipomyia vespa Debenham, by original 

designation. 

capax Debenham, 1987d: 670. Australia (New South Wales). 

circinate Liu and Yu, in Liu et al., 2001b: 28. China. 

edgari Tokunaga and Murachi, 1959: 154 Micronesia. 

hamoni de Meillon, 1959a: 329. Burkina Faso. 

hikosanensis Tokunaga, 1940c: 81. Japan. 

improbiserra Debenham, 1987d: 667. Papua New Guinea. 

madeira Clastrier, 1991b: 261. Madeira (Portugal). 

pachyparamera Clastrier and Delécolle, 1991: 187. New Caledonia (France). 

transversalis Clastrier and Delécolle, 1991: 190. New Caledonia (France). 

vespa Debenham, 1987d: 668. Australia (New South Wales). 

Subgenus EUPROJOANNISIA Brèthes 

EUPROJOANNISIA Brèthes, 1914: 155. Type species: Euprojoannisia platensis Brèthes, by original designation. 

EUFORCIPOMYIA Malloch, 1915b: 312. Type species: Euforcipomyia hirtipennis Malloch (= Ceratopogon palustris Meigen), 


CRYPTOSCENA Enderlein, 1936: 51. Type species: Ceratopogon palustris Meigen, by monotypy. 

PROFORCIPOMYIA Saunders, 1957: 662 (as subgenus of Forcipomyia). Type species: Forcipomyia wirthi Saunders, by 


aequalis (Skuse), 1889: 294 (Ceratopogon). Australia (New South Wales). 

aenigma de Meillon and Downes, 1986: 146. South Africa. 

alacris (Winnertz), 1852: 25 (Ceratopogon). Germany. 

amiantos Yu and Song, 2008: 794. China. 

antipodum (Hudson), 1892: 45 (Ceratopogon). New Zealand. 

novaezelandiae Kieffer, 1922a: 145. New Zealand. 

appendicular Liu, Yan and Liu, 1996: 18. China. 

arcis de Meillon and Downes, 1986: 143. South Africa. 

astyla Tokunaga, 1940c: 92. Japan. 

asymmetrica Remm, 1980: 118. Kazakhstan. 

atomaria Tokunaga, 1940c: 93. Japan. 

avocadonis de Meillon and Wirth, 1979b: 188. South Africa. 

bahelea Liu and Yu, in Liu et al., 2001b: 39. China. 

balteatus Liu and Yu, in Liu et al., 2001b: 40. China. 

bellerophon Debenham and Wirth, 1984: 869. Australia (Queensland). 

berendti Szadziewski, 1988: 209. Baltic region. Eocene. 

bituberculifera Tokunaga, in Tokunaga and Murachi, 1959: 188. Belau (USA). 

blantoni Soria and Bystrak, 1975: 3. Brazil. 

borealis Remm, 1966: 54. Lithuania. 

breviforceps Tokunaga, 1940c: 98. Japan. 

bromeliae Saunders, 1957: 665. Brazil. 

bureschi (Zilahi-Sebess), 1934: 153 (Lasiohelea). Bulgaria. 

calamistrata Debenham and Wirth, 1984: 862. Australia (Queensland). 

calcarata (Coquillett), 1905: 64 (Ceratopogon). Mexico. 

canadensis Bystrak and Wirth, 1978: 19. Canada (Saskatchewan). 

centrosus Liu and Yu, in Liu et al., 2001b: 44. China. 

chongmingensis Liu and Yu, in Liu et al., 2001b: 46. China. 

30

claudus Liu and Yu, in Liu et al., 2001b: 48. China. 

deformis Liu and Yu, in Liu et al., 2001b: 50. China. 

deucalionis Debenham and Wirth, 1984: 875. Australia (New South Wales). 

dolichopodida Chan and Linley, 1989: 253. USA (Florida). 

dowi Bystrak and Wirth, 1978: 21. USA (Florida). 

dycei Debenham and Wirth, 1984: 871. Australia (New South Wales). 

esteparia Marino and Spinelli, 2001b: 12. Argentina. 

exemliforma Saha, Dasgupta, Gangopadhyay and Mukherjee, 2009: 12. India. 

falcifera Saunders, 1959: 39. Trinidad and Tobago. 

fermetitillans Saha, Dasgupta, Gangopadhyay and Mukherjee, 2009: 12. India. 

flavencruris Saha, Dasgupta, Gangopadhyay and Mukherjee, 2009: 13. India. 

formosana Kieffer, 1916a: 83. Taiwan. 

fuscicalcarata Bystrak and Wirth, 1978: 23. USA (Florida). 

fuscimana (Kieffer), 1921g: 559 (Ceratopogon). Taiwan. 

pennielongata Chan and LeRoux, 1971b: 733. Singapore. 

galeata Debenham and Wirth, 1984: 878. Australia (Queensland). 

grandiclara Saha, Dasgupta, Gangopadhyay and Mukherjee, 2009: 14. India. 

hardyi Wirth and Howarth, 1982: 130. USA (Hawaii). 

henningseni Szadziewski, 1988: 211. Denmark. Eocene. 

iaculum Debenham and Wirth, 1984: 883. Papua New Guinea. 

ignobilis Liu and Yu, in Liu et al., 2001b: 51. China. 

indoctacruris Saha, Dasgupta, Gangopadhyay and Mukherjee, 2009: 14. India. 

kaneohe Wirth and Howarth, 1982: 141. USA (Hawaii). 

karnyi Edwards, 1923a: 183. Indonesia. 

kyotoensis Tokunaga, 1940c: 99. Japan. 

largus Liu and Yu, in Liu et al., 2001b: 53. China. 

lingnanenis Liu and Yu, in Liu, Yan and Yu, 1999:309. China. 

lokki Remm, 1993: 199. Russia. 

longicalcar Kieffer, 1912c: 4. Sri Lanka. 

longispina Saunders, 1957: 669. Brazil. 

lota (Williston), 1896: 282 (Ceratopogon). St. Vincent. 

marsalae (Vattier and Adam), 1966a: 305 (Lasiohelea). Gabon. 

mesasiatica Remm, 1980: 118. Tajikistan. 

minor Liu, Yan and Liu, 1996: 18. China. 

miocaenica Szadziewski, 1993: 647. Germany. Eocene. 

mortuifolii Saunders, 1959: 35. Trinidad and Tobago. 

mucronis Liu and Yu, in Liu et al., 2001b: 59. China. 

nitens (Santos Abreu), 1918: 260 (Helea). Canary Islands (Spain). 

soranus de Meillon, 1943: 103. South Africa. 

navaiae Bystrak and Wirth, 1978: 31. USA (Florida). 

palasunitheca Saha, Dasgupta, Gangopadhyay and Mukherjee, 2009: 15. India. 

palikuensis Hardy, 1960: 175. USA (Hawaii). 

palusitheca Saha, Dasgupta, Gangopadhyay and Mukherjee, 2009: 16. India. 

palustris (Meigen), 1804: 28 (Ceratopogon). Germany. 

hirtipennis (Malloch), 1915b: 313 (Euforcipomyia). USA (Illinois). 

turfacea Kieffer, 1925b: 148. Estonia. 

pechumani Bystrak and Wirth, 1978: 34. USA (New York). 

perflavida Remm, 1971: 186. Russia. 

phlebotomoides Bangerter, 1933: 252. Switzerland. 

subsaltans Remm, 1961a: 189. Estonia. 

pholeter Wirth and Howarth, 1982: 139. USA (Hawaii). 

picturatus Liu and Yu, in Liu et al., 2001b: 65. China. 

piriformis (Meunier), 1904a: 228 (1904b: 239) (Ceratopogon). Baltic region. Eocene. 

pipiens Liu, Yan and Liu, 1996: 18. China. 

31

platensis (Brèthes), 1914: 156 (Euprojoannisia). Argentina. 

propenavaiae Saha, Dasgupta, Gangopadhyay and Mukherjee, 2009: 16. India. 

pseudocalcarata Saha, Dasgupta, Gangopadhyay and Mukherjee, 2009: 17. India. 

psilonota (Kieffer), 1911c: 337 (Ceratopogon). Seychelles. 

aplonota (Kieffer), 1911c: 337 (Ceratopogon). Seychelles. 

seychelleana (Kieffer), 1911c: 338 (Ceratopogon). Seychelles. 

fulvithorax (Kieffer), 1911c: 338 (Ceratopogon, as variety of seychelleana). Seychelles. 

indecora Kieffer, 1914b: 269. South Africa. 

litoralis Santos Abreu, 1918: 277. Canary Islands (Spain). 

ingrami Carter, 1919: 290. Ghana. 

egypti Macfie, 1924: 61. Egypt. 

hathor Kieffer, 1925e: 247. Egypt. 

conogensis Goetghebuer, 1933e: 132 (also as congolensis). Democratic Republic of the Congo. 

flavipilosella Goetghebuer, 1933e: 130 (also as flavopilosella, p. 135). Democratic Republic of the Congo. 

lulengaensis Goetghebuer, 1935d: 155. Democratic Republic of the Congo. 

superata Goetghebuer, 1935d: 160. Democratic Republic of the Congo. 

griseipluma Goetghebuer, 1935d: 154. Democratic Republic of the Congo. 

griseolella Goetghebuer, 1948b: 7. Democratic Republic of the Congo. 

quasiingrami Macfie, 1939c: 164. Brazil. 

ratis Liu and Yu, in Liu et al., 2001b: 71. China. 

sagittaria Debenham and Wirth, 1984: 880. Australia (Queensland). 

saltans (Winnertz), 1852: 27 (Ceratopogon). Germany. 

saltivaga (Skuse), 1889: 295 (Ceratopogon). Australia (New South Wales). 

sauteri Kieffer, 1912a: 27. Taiwan. 

clara Chan and LeRoux, 1971b: 729. Singapore. 

scapularis Goetghebuer, 1933c: 353. Belgium. 

separatim Liu, Yan and Liu, 1996: 21. China. 

setigera Saunders, 1959: 38. Trinidad and Tobago. 

setosicrus (Kieffer), 1906c: 357 (Ceratopogon). Argentina. 

simulans Johannsen, 1932: 411. Indonesia. 

sonora Wirth, 1952a: 145 (as variety of calcarata). USA (California). 

spatulifera Saunders, 1957: 667. Brazil. 

subcalcarata Saha, Dasgupta, Gangopadhyay and Mukherjee, 2009: 17. India. 

subingrami Saha, Dasgupta, Gangopadhyay and Mukherjee, 2009: 18. India. 

subnitida (Skuse), 1889: 299 (Ceratopogon). Australia (New South Wales). 

subunitheca Saha, Dasgupta, Gangopadhyay and Mukherjee, 2009: 19. India. 

titillans (Winnertz), 1852: 27 (Ceratopogon). Germany. 

divaricata (Winnertz), 1852: 25 (Ceratopogon). Germany. 

maricarmenae Sahuquillo and Gil Collado, 1982a: 303. Spain. 

tonicus Liu and Yu, in Liu et al., 2001b: 74. China. 

tsacasi Clastrier, 1983c: 38. Seychelles. 

tuberculosa Szadziewski, 1993: 648. Germany. Eocene. 

unica Bystrak and Wirth, 1978: 44. USA (Florida). 

vandiemeni Debenham and Wirth, 1984: 864. Australia (Tasmania). 

vernocheti (Clastrier), 1959c: 439 (Lasiohelea). Réunion (France). 

wansoni de Meillon, 1939a: 9. Democratic Republic of the Congo. 

wirthi Saunders, 1957: 663. USA (California). 

wirthiana Szadziewski, 1983a: 374. Algeria. 

yapensis Tokunaga and Murachi, 1959: 185. Micronesia. 

yoshimurai Tokunaga, 1940c: 91. Japan. 

Subgenus FORCIPOHELEA Clastrier and Delécolle 

FORCIPOHELEA Clastrier and Delécolle, 1991: 195 (as subgenus of Forcipomyia). Type species: Forcipomyia tillierorum 

32

Clastrier and Delécolle, by original designation. 

tillierorum Clastrier and Delécolle, 1991: 197. New Caledonia (France). 

Subgenus FORCIPOMYIA Meigen 

abyssiniae (Kieffer), 1918a: 40 (Ceratopogon). Ethiopia. 

adjecta Tokunaga, 1959: 284. Indonesia. 

akizukii Tokunaga, 1940c: 71. Russia. 

alamatae Macfie, 1937b: 73. Ethiopia. 

alatauensis Remm, 1980: 123. Kazakhstan. 

albiradialis Tokunaga, 1940c: 75. Japan. 

albonotata (Kieffer), 1910: 185 (Ceratopogon). India. 

albopunctata (Skuse), 1889: 293 (Ceratopogon). Australia (New South Wales). 

albosignata (Kieffer), 1910: 184 (Ceratopogon). Myanmar. 

albostyla Remm, 1979b: 58. Estonia. 

allocera Rieth, 1915: 438. New name for heterocera. 

heterocera Kieffer, 1914a: 234 (preoccupied by Forcipomyia heterocera Kieffer, 1913d). Germany. 

allocera Kieffer, in Thienemann and Kieffer, 1916: 491 (preoccupied by Forcipomyia allocera Rieth, 1915). New 

name for heterocera. 

altaica Remm, 1972: 62. Russia. 

annandalei Edwards, 1932a: 177. India. 

antrijovis (Kieffer), 1919a: 15 (Ceratopogon). Greece. 

antrosus Liu and Yu, in Liu et al., 2001b: 85. China. 

appendicular Liu, Yan and Liu, 1996: 18. China. 

aquatica Kieffer, 1922c: 230. Germany. 

arcigera Kieffer, 1922b: 150. Taiwan. 

argenteola Macfie, 1939c: 146. Brazil. 

armandi Harant, Huttel and Huttel, 1952: 13. Italy. 

armativentris Clastrier, 1960c: 518. Congo. 

armendarizi Sahuquillo and Gil Collado, 1982b: 744. Spain 

ashantii Ingram and Macfie, 1924b: 589. Ghana. 

rufula Goetghebuer, 1935d: 159. Democratic Republic of the Congo. 

asticta Kieffer, 1913d: 172 (as variety of albosignata). India. 

atricella Debenham, 1987c: 304. Australia (New South Wales). 

aurea Malloch, 1915a: 318. USA (Illinois). 

auronitens (Kieffer), 1910: 183 (Ceratopogon). Myanmar. 

bessa Liu and Yu, in Liu et al., 2001b: 86. China. 

biannulata Ingram and Macfie, 1924b: 557. Ghana, Nigeria, Malawi. 

quatuorguttata Goetghebuer, 1935d: 158 (also misspelled as quadriguttata). Democratic Republic of the Congo. 

bicolorata Goetghebuer, 1935d: 150. Democratic Republic of the Congo. 

marginella Goetghebuer, 1935d: 156. Democratic Republic of the Congo. 

nigricosta Goetghebuer, 1935d: 158. Democratic Republic of the Congo. 

pallidula Goetghebuer, 1948b: 6. Democratic Republic of the Congo. 

abonnenci Clastrier, 1959b: 340. Senegal. 

bikanni Chan and LeRoux, 1971b: 735. Singapore. 

binigrimaculata Tokunaga, 1940c: 68. Japan. 

bipunctata (Linnaeus), 1767: 978 (Tipula). Europe. 

trichoptera (Meigen), 1804: 31 (Ceratopogon). Germany. 

ambigua (Meigen), 1804: 32 (Ceratopogon). Europe. 

geniculata (Guérin-Méneville), 1833: 164 (Ceratopogon). France. 

palmensis Santos Abreu, 1918: 285. Canary Islands (Spain). 

squamaticrus (Kieffer), 1919a: 9 (Ceratopogon). Belgium, Serbia-Montenegro, Croatia. 

tenuisquama Kieffer, 1924b: 393. France. 

33

laguncula Kieffer, 1925d: 27. Hungry. 

flavipubens Goetghebuer, 1927b: 93. Belgium. 

bipunctatapropinqua Chan and LeRoux, 1971b: 740. Canada (Quebec). 

bitensis Kieffer, 1924b: 394. France. 

brachypetiolata Vimmer, 1928: 58. Israel. 

brachytoma Kieffer, 1922b: 149. Taiwan. 

braueri (Wasmann), 1893: 277 (Ceratopogon). Austria, Switzerland. 

brevipalpis Liu and Yu, in Liu et al., 2001b: 92. China. 

brevipalpis Saha, Dasgupta, Gangopadhyay and Mukherjee, 2009: 20. India. [needs replacement name] 

brevipedicillata (Kieffer), 1901b: 217 (Ceratopogon). France. 

brevipennis (Macquart), 1826: 179 (Ceratopogon). France. 

lateralis (Bouché), 1834: 23 (Ceratopogon). Europe. 

specularis (Coquillett), 1901a: 601. USA (Pennsylvania). 

coprophila Kieffer, 1914a: 233. Germany. 

disticta (Kieffer), 1919a: 16 (Ceratopogon). Hungry, Slovak Republic, Croatia, Romania. 

hirtidorsum (Kieffer), 1919a: 18 (Ceratopogon). Ukraine. 

nigrimana (Kieffer), 1919a: 19 (Ceratopogon). Lithuania. 

bifilis Kieffer, 1922c: 231. Germany. 

subnigra Tokunaga, 1940c: 88. Japan. 

bystraki Grogan and Wirth, 1975c: 466. USA (Virginia). 

calatheae Wirth, 1982a: 573. Dominica. 

caliginosella Wirth, 1974: 5. New name for caliginosa. 

caliginosa Ingram and Macfie, 1931a: 167 (preoccupied by Forcipomyia caliginosa (Ingram and Macfie), 1924a). 

Argentina. 

castanea (Walker), 1848: 26 (Ceratopogon). Sierra Leone. 

risbeci Séguy, 1946: 9. Senegal. 

incomptifeminiba (Austen), 1912: 107 (Ceratopogon). Ghana. 

catarinensis Marino and Spinelli, 2002: 309. Brazil. 

cattleyarum Harant and Galan, 1942b: 56. France. 

cavatus Liu and Yan in Liu, Yan and Yu, 1999:311 (Liu and Yu, in Liu et al., 2001b: 92). China. 

charon Debenham, 1987c: 276. Australia (Western Australia). 

chilensis (Philippi), 1865: 601 (Ceratopogon). Chile. 

chrysothrix (Kieffer), 1921g: 558 (Ceratopogon). Taiwan, Philippines. 

ciliata (Winnertz), 1852: 21 (Ceratopogon). Germany. 

brunnipes (Perris), 1847: 556 (Ceratopogon, preoccupied by Atrichopogon brunnipes (Meigen), 1804). France. 

perrisi (Kieffer), 1901a: 147 (Ceratopogon). New name for brunnipes. 

boleti (Kieffer), 1901a: 157 (Ceratopogon). France. 

turfosa Kieffer, 1925b: 146. Estonia. 

canicularis Goetghebuer, 1948a: 36. Belgium. 

zlatensis Damian-Georgescu, 1972: 18. Romania. 

ciliola Liu and Yu, in Liu et al., 2001b: 96. China. 

cilipes (Coquillett), 1900: 397 (Ceratopogon). USA (Alaska). 

cirrhosa Clastrier, Rioux and Descous, 1961: 50. Chad. 

claggi Tokunaga, in Tokunaga and Murachi, 1959: 150. Guam (USA). 

coarctata (Kieffer), 1901a: 161 (Ceratopogon). Romania. 

collyricus Liu, Yan and Liu, 1996: 19. China. 

concolor Malloch, 1915a: 319 (as variety of pergandei). USA (Illinois). 

confluens Kieffer, 1913d: 170. India. 

conigera Kieffer, 1913d: 168. India. 

contraria Debenham, 1987c: 307. Australia (New South Wales). 

conturbatus Liu and Yu, in Liu et al., 2001b: 100. China. 

cooki Macfie, 1932c: 28. New Zealand. 

corinneae Gosseries, 1989: 3. New name for vexans. 

vexans (de Meijere), 1909: 198 (Ceratopogon, preoccupied by Culicoides vexans (Staeger), 1839). Indonesia. 

34

corticis Kieffer, 1911d: 2. Germany. 

costata (Zetterstedt), 1838: 815 (Ceratopogon). Sweden. 

picea (Winnertz), 1852: 21 (Ceratopogon). Germany. 

latipalpis (Kieffer), 1901a: 159 (Ceratopogon). France. 

meinerti Kieffer, 1915c: 281. Denmark. 

corticicola (Kieffer), 1919a: 14 (Ceratopogon). Germany. 

turficola Kieffer, 1925b: 147. Russia. 

crassipalpis Debenham, 1987c: 303. Australia (New South Wales). 

crassipes (Winnertz), 1852: 22 (Ceratopogon). Germany. 

creesi de Meillon, Meiswinkel and Wirth, 1982: 123. South Africa. 

cubicularis Kieffer, 1911b: 320. India. 

curtus Liu and Yu, in Liu et al., 2001b: 103. China. 

dacica Damian-Georgescu, 1972: 17. Romania. 

danica Kieffer, 1915c: 280. Denmark. 

daxingensis Liu and Yu, in Liu et al., 2001b: 105. China. 

decipiens (Kieffer), 1910: 182 (Ceratopogon). India. 

declivis Debenham, 1987c: 295. Australia (New South Wales). 

desertensis Wirth and Hubert, 1960a: 639. USA (California). 

despecta Kieffer, 1922b: 152. Taiwan. 

desurvillei Macfie, 1932c: 28. New Zealand. 

dichromata Remm, in Remm and Zhogolev, 1968: 826. Ukraine. 

dirina Liu and Yu, in Liu et al., 2001b: 106. China. 

dissimilis Clastrier and Delécolle, 1991: 200. New Caledonia (France). 

distapalpis Liu and Yu, in Liu et al., 2001b: 108. China. 

dividus Liu and Yu, in Liu et al., 2001b: 110. China. 

donatoi Marino and Spinelli, 2008:790. Paraguay. 

edwardsiana Wirth, 1974: 5. New name for edwardsi. 

edwardsi Ingram and Macfie, 1931a: 160 (preoccupied by Forcipomyia edwardsi (Saunders), 1925). Argentina. 

elegans de Meillon and Wirth, 1987a: 36. South Africa. 

elegantula Malloch, 1915b: 311. USA (Illinois). 

elongata (Kieffer), 1901a: 160 (Ceratopogon). Romania. 

ensifera Macfie, 1934c: 182. Malaysia. 

eobreviflagellata Szadziewski, 1988: 220. Poland. Eocene. 

eocostata Szadziewski, 1988: 202. Poland. Eocene. 

episcopa Debenham, 1987c: 291. Australia (New South Wales). 

euzierei Harant and Galan, 1943: 122. France. 

excellans Johannsen, 1932: 413. Indonesia. 

fangchengensis Liu and Yu, in Liu et al., 2001b: 111. China. 

fasciata Tokunaga, 1940c: 78. Japan. 

fascilatipes Saha, Dasgupta, Gangopadhyay and Mukherjee, 2009: 21. India. 

fascicauda Tokunaga, 1940c: 62 (1940e: 167). Micronesia. 

fascicornis Tokunaga, 1962a: 171. Japan. 

fengjiensis Yu, Chen and He, in Yu et al., 2007: 487. China. 

fidelis Krivosheina, 1968: 586. Russia. 

filicauda Remm, 1993: 200. Russia. 

fimbriata (Coquillett), 1901a: 601 (Ceratopogon). USA (District of Columbia). 

flava (Williston), 1896: 280 (Ceratopogon). St. Vincent. 

flaviceps Kieffer, 1913d: 173 (as variety of macrothrix). India. 

flavicincta Santos Abreu, 1918: 282. Canary Islands (Spain). 

flavicoxis Goetghebuer, 1935d: 152. Democratic Republic of the Congo. 

wittei Goetghebuer, 1948b: 5. Democratic Republic of the Congo. 

flavipectoralis Tokunaga, 1959: 280. Papua New Guinea. 

flavomaculata Vimmer, 1928: 58. Israel. 

flavominima Remm, 1971: 184. Russia. 

35

formicaria (Kieffer), in Kieffer and Thienemann, 1908: 2 (Ceratopogon). France. 

freyi Storå, 1936: 32. Canary Islands (Spain). 

frigidus Liu and Yu, in Liu et al., 2001b: 114. China. 

fulvescens Santos Abreu, 1918: 284. Canary Islands (Spain). 

fulvipes Saha, Dasgupta, Gangopadhyay and Mukherjee, 2009: 22. India. 

fulvotibialis Saha, Dasgupta, Gangopadhyay and Mukherjee, 2009: 22. India. 

funebris (Meigen), 1830: 262 (Ceratopogon). Europe. 

furfura de Meillon and Wirth, 1989b: 205. Zimbabwe. 

fusca (Philippi), 1865: 630 (Tetraphora). Chile. 

patagonica Ingram and Macfie, 1931a: 165. Argentina. 

fusciforceps (Kieffer), 1918a: 38 (Ceratopogon). Guinea. 

fusciparamera Szadziewski and Grogan, 1998b: 266. Dominican Republic. Miocene. 

gedanicola Szadziewski, 1988: 200. Poland. Eocene. 

geniflava Tokunaga, 1959: 278. Papua New Guinea. 

genualis (Loew), 1866: 128 (Ceratopogon). Cuba. 

propinqua (Williston), 1896: 279 (Ceratopogon). St. Vincent. 

raleighi Macfie, 1938: 160. Trinidad and Tobago. 

globofuscitheca Saha, Dasgupta, Gangopadhyay and Mukherjee, 2009: 23. India. 

gloriose Liu and Yu, in Liu et al., 2001b: 117. China. 

grimaldii Szadziewski and Grogan, 1998b: 268. Dominican Republic. Miocene. 

guamensis Tokunaga and Murachi, 1959: 166. Guam (USA). 

guangdongensis Liu and Yu, in Liu et al., 2001b: 119. China. 

guilleaumei Goetghebuer, 1935b: 413. Belgium. 

guttata de Meillon and Wirth, 1989b: 204. Zimbabwe. 

haranti (Huttel and Huttel), 1952a: 31 (Monohelea). France. 

harpegonata Wirth and Soria, 1975: 19. Puerto Rico (USA). 

heilongjiangensis Liu and Yu, in Liu et al., 2001b: 121. China. 

herbaceus Liu and Yu, in Liu et al., 2001b: 123. China. 

heterocera Kieffer, 1913d: 173. India. 

himalayae Kieffer, 1911b: 320. India. 

hirsuta Ingram and Macfie, 1924b: 577. Ghana. 

hirtula (Zetterstedt), 1838: 815 (Chironomus). Sweden. 

simulata Walley, 1932: 165. Canada (Ontario). 

paradoxa Krivosheina, 1968: 588. Russia. 

holofuscicruris Saha, Dasgupta, Gangopadhyay and Mukherjee, 2009: 24. India. 

hurdi Wirth, 1952a: 143. USA (California). 

hutteli Arnold and Jarry, 1956: 136. France. 

hybrida Remm, 1980: 122. Tajikistan. 

hydratus Liu, Yan and Liu, 1996: 19. China. 

hygrophila Kieffer, 1925b: 147. Estonia. 

sphagnicola Kieffer, 1925b: 148. Estonia. 

occidentalis Wirth, 1952a: 141. USA (California). 

imaculata Vimmer, 1928: 58. Israel. 

imparitheca Saha, Dasgupta, Gangopadhyay and Mukherjee, 2009: 24. India. 

ingenua Macfie, 1934c: 180. Malaysia. 

intrudens Liu, Yan and Liu, 1996: 19. China. 

ishikariensis Yamashita, Kitamura and Nakamura, 1957: 91. Japan. 

ishizuchiensis (Udaka), 1959: 18 (Lasiohelea). Japan. 

kaltenbachi (Winnertz), 1852: 19 (Ceratopogon). Germany. 

kilemae Kieffer, 1913e: 4. Kenya. 

kisantuensis Goetghebuer, 1933e: 134. Democratic Republic of the Congo. 

kribiensis (Kieffer), 1921b: 6 (Ceratopogon). Cameroon. 

krzeminskii Szadziewski, 1988: 217. Poland. Eocene. 

kulickae Szadziewski, 1988: 219. Poland. Eocene. 

36

laboulbeni (Perris), 1870: 138 (Ceratopogon). France. 

lagonigera Kieffer, 1922b: 151. Taiwan. 

lanceolata Macfie, 1934c: 178 (1934d: 205). Malaysia. 

laxus Liu and Yu, in Liu et al., 2001b: 129. China. 

lepida (Winnertz), 1852: 23 (Ceratopogon). Germany. 

lepidopa (Kieffer), 1917a: 179 (Ceratopogon). Papua New Guinea. 

leptolepis Krivosheina and Remm, 1974: 116. Russia. 

letabana de Meillon, Meiswinkel and Wirth, 1982: 125. South Africa. 

leucochaeta (Kieffer), 1921b: 5 (Ceratopogon). Cameroon. 

lignicola (Brunetti), 1912: 445 (Ceratopogon). India. 

lochmocola Zou and Yu, 1991: 57. China. 

longiconus Liu and Yu, in Liu et al., 2001b: 132. China. 

longiseta Yu and Liu, 1999: 246 (Liu and Yu, in Liu et al., 2001b:134). China. 

longisetosa Krivosheina and Remm, 1974: 118. Russia. 

helvetica Delécolle and Schiegg, 1999b: 389. Switzerland. 

lugubris (Zetterstedt), 1855: 4863 (Ceratopogon). Sweden. 

lurida Saha, Dasgupta, Gangopadhyay and Mukherjee, 2009: 25. India. 

luteofulvous Liu and Yu, in Liu et al., 2001b: 136. China. 

lyneborgi Szadziewski, 1988: 196. Denmark. Eocene. 

macronyx Goetghebuer, 1933e: 131. Democratic Republic of the Congo. 

longiventris Goetghebuer, 1935d: 155. Democratic Republic of the Congo. 

stanleyi Macfie, 1939a: 86. Uganda. 

macrorhyncha (Kieffer), 1910: 183 (Ceratopogon). India. 

macrothrix Kieffer, 1911b: 325. India. 

macswaini Wirth, 1952a: 130. USA (California). 

maculipennis Tokunaga, 1940c: 64. Japan. 

magnasacculus Liu and Yu, in Liu et al., 2001b: 138. China. 

magnipunctata Tokunaga, 1940c: 90. Japan. 

manchuriensis Tokunaga, 1941b: 90. China. 

marini Spinelli and Dippolito, 1995: 155. Venezuela. 

maura (Kieffer), 1918a: 37 (Ceratopogon). Mozambique. 

mcmillani Clastrier and Wirth, 1961a: 191. Nigeria. 

melanchroa Ingram and Macfie, 1924b: 591. Nigeria. 

minitheca Marino and Spinelli, 2001a: 111. Chile. 

minor Liu, Yan and Liu, 1996: 18. China. 

mira Johannsen, 1932: 410. Indonesia. 

miricornis Kieffer, 1916a: 81. Taiwan. 

monticola Kieffer, 1913d: 171. India. 

monticolonia Tokunaga, 1966b: 284. Afghanistan. 

multipicta Ingram and Macfie, 1931a: 157. Argentina. 

murphyi Clastrier and Wirth, 1961a: 194. Gambia. 

muzoni Marino and Spinelli, 2004a: 150. Argentina. 

myrmecophila (Egger), 1863: 1109 (Ceratopogon). Austria. 

nigeriensis Ingram and Macfie, 1924b: 592. Nigeria. 

nigra (Winnertz), 1852: 17 (Ceratopogon). Germany. 

sphagnorum Kieffer, 1925b: 149. Estonia. 

nigrans Remm, 1962b: 188. Estonia. 

nigricoxis Goetghebuer, 1935d: 158. Democratic Republic of the Congo. 

elongata Goetghebuer, 1935d: 152 (preoccupied by Forcipomyia elongata (Kieffer), 1901a). Democratic Republic of 

Congo. 

thripsiformis Goetghebuer, 1935d: 161. Democratic Republic of the Congo. 

lugardi Macfie, 1939a: 85. Uganda. 

kibatiensis Goetghebuer, 1948b: 5. Democratic Republic of the Congo. 

elongatula Goetghebuer, 1948b: 6. New name for elongata. 

37

atripes Goetghebuer, 1948b: 8. Democratic Republic of the Congo. 

africana Goetghebuer, 1948b: 8. Democratic Republic of the Congo. 

nigrotibialis Ingram and Macfie, 1924b: 571. Ghana. 

nilicola (Kieffer), 1925e: 253 (Apelma). Egypt. 

niligena (Kieffer), 1921b: 4 (Ceratopogon). Sudan. 

nilotica (Kieffer), 1921b: 3 (Lepidohelea). Sudan. 

noctivaga Kieffer, 1912c: 3. Sri Lanka. 

notata Macfie, 1939a: 84. Uganda. 

nuncupata Macfie, 1949: 110. Mexico. 

obscura (Walker), 1848: 26 (Ceratopogon). Canada. 

ochracea Vimmer, 1928: 58. Israel. 

orientalis Kieffer, 1923b: 135. Indonesia. 

ornaticrus Kieffer, 1912c: 1. Sri Lanka. 

ornatipennis Macfie, 1939c: 151. Brazil. 

ornatipes (Kieffer), 1918a: 88 (Ceratopogon). Sri Lanka. 

paenealbiradialis Saha, Dasgupta, Gangopadhyay and Mukherjee, 2009: 26. India. 

palpipotens Saha, Dasgupta, Gangopadhyay and Mukherjee, 2009: 26. India. 

pallida (Winnertz), 1852: 15 (Ceratopogon). Germany. 

pallidipes Santos Abreu, 1918: 276 (as variety of bipunctata). Canary Islands (Spain). 

rustica (Kieffer), 1919a: 19 (Ceratopogon). Hungary, Croatia. 

cataneii Kieffer, 1923a: 662. Algeria. 

pallidistyla Krivosheina and Remm, 1974: 121. Russia. 

palmarum Debenham, 1987c: 299. Papua New Guinea. 

parda Debenham, 1987c: 295. Australia (New South Wales). 

parva (Walker), 1848: 26 (Ceratopogon). Canada. 

parvicellula Ingram and Macfie, 1931b: 195. New Zealand. 

peradeniyae Dessart, 1963: 186. New name for calcarata. 

calcarata Kieffer, 1912c: 3 (preoccupied by Forcipomyia calcarata (Coquillett), 1905). Sri Lanka. 

calcarhelea Wirth, 1973: 352. New name for calcarata. 

pergandei (Coquillett), 1901a: 602 (Ceratopogon). USA (District of Columbia). 

persa Liu and Yu, in Liu et al., 2001b: 139. China. 

picheyrei Harant and Galan, 1942a: 135. Algeria. 

pictiscutaris Tokunaga, 1959: 270. Papua New Guinea. 

pictoni Macfie, 1938: 161. Trinidad and Tobago. 

pilosa (Coquillett), 1902a: 87 (Ceratopogon). USA (District of Columbia). 

pinamarensis Spinelli, 1983a: 121. Argentina. 

pingxiangensis Liu and Yu, in Liu et al., 2001b: 141 (also as pinxiangensis). China. 

pinicola Bystrak and Messersmith, 1980: 109. USA (Maryland). 

piroskyi Cavalieri, 1961b: 172. Argentina. 

poulaineae Ingram and Macfie, 1931a: 227. Argentina. 

praealtus Liu and Yu, in Liu et al., 2001b: 143. China. 

praecincta Santos Abreu, 1918: 279. Canary Islands (Spain). 

pretoriana (Kieffer), 1918a: 39 (Ceratopogon). South Africa. 

pseudomicrohelea Szadziewski, 1988: 205. Baltic region. Eocene. 

pseudonigra Delécolle and Schiegg, 1999b: 381. Switzerland. 

pseudobipunctata Saha, Dasgupta, Gangopadhyay and Mukherjee, 2009: 27. India. 

psychasta Kieffer, 1913d: 170. India. 

puhtuensis Remm, 1979b: 58. Estonia. 

pulchrithorax Edwards, in Saunders, 1924: 209. Great Britain. 

pulla de Meillon and Wirth, 1981b: 527. South Africa. 

pulluspadicruris Saha, Dasgupta, Gangopadhyay and Mukherjee, 2009: 27. India. 

punctatipennis Kieffer, 1924b: 394 (as variety of braueri). France. 

punctipes Edwards, 1928: 49. Western Samoa. 

punctumalbum (Kieffer), 1917a: 178 (Ceratopogon). Papua New Guinea. 

38

quadriflava Tokunaga, 1959: 272. Papua New Guinea. 

quasialbiradialis Saha, Dasgupta, Gangopadhyay and Mukherjee, 2009: 28. India. 

quatei Wirth, 1952a: 142. USA (California). 

quechua Marino and Spinelli, 2002: 312. Argentina. 

qufuensis Liu and Yu, in Liu et al., 2001b: 146. China. 

radicicola Edwards, in Saunders, 1924: 208. Great Britain. 

padi Remm, 1979b: 59. Estonia. 

radiifera (Kieffer), 1918a: 35 (Ceratopogon). South Africa. 

recussus Liu and Yu, in Liu et al., 2001b: 148. China. 

regula (Winnertz), 1852: 16 (Ceratopogon). Germany. 

metatarsis Tokunaga, 1940c: 69. Japan. 

longimaculata Tokunaga, 1940c: 85. Japan. 

repandus Liu and Yu, in Liu et al., 2001b: 149. China. 

resinicola (Kieffer), 1901a: 159 (Ceratopogon). France. 

rioplatensis Marino and Spinelli, 2002: 314. Argentina. 

rufescens (Kieffer), 1918a: 40 (Ceratopogon). Tunisia. 

ruralis Liu and Yu, in Liu et al., 2001b: 151. China. 

sahariensis Kieffer, 1923a: 663. Algeria. 

armaticrus Kieffer, 1923a: 661. Algeria. 

tuzeti Huttel and Huttel, 1952c: 178. France. 

acanthophora Remm, in Havelka, 1976a: 227. Kyrgyzstan. 

onusta Remm, 1980: 122. Kazakhstan. 

salmi (de Meijere), 1909: 197 (Ceratopogon). Indonesia. 

sanguinolenta Kieffer, 1925a: 407. Germany. 

securis Chan and LeRoux, 1971b: 751. Singapore. 

semirustica Remm, in Remm and Zhogolev, 1968: 828. Ukraine. 

sensillata Clastrier, 1983c: 44. Seychelles. 

sexannulata Clastrier, 1983c: 45. Seychelles. 

sexvittata Wirth, 1956c: 248. Costa Rica. 

sihlwaldensis Delécolle and Schiegg, 1999b: 385. Switzerland. 

skusei Debenham, 1987c: 279. Australia (New South Wales). 

somuncurensis Marino and Spinelli, 2001a: 116. Argentina. 

spatuligera Macfie, 1949: 111. Mexico. 

sphagnophila Kieffer, 1925a: 406. Russia. 

solonensis Wirth, 1951d: 315. USA (Virginia). 

oreophila Remm, 1972: 63 (as oreophilia). Russia. 

spiculata Sinha, Mazumdar, and Chaudhuri, 2003: 75. India. 

spinuliforceps Tokunaga, 1959: 275. Papua New Guinea. 

squamigera Kieffer, in Thienemann and Kieffer, 1916: 491. Sweden. 

apricans (Kieffer), 1919a: 19 (Ceratopogon). Slovak Republic. 

squamipes (Coquillett), 1902a: 88 (Ceratopogon). USA (New Mexico). 

brumalis (Long), 1902: 3 (Ceratopogon). USA (Texas). 

squamitibia Lutz, 1914: 88. Brazil. 

soriai Wirth, 1991b: 168. Brazil. 

squamitibialis Tokunaga, 1959: 269. Papua New Guinea. 

stenammatis (Long), 1902: 10 (Ceratopogon). USA (Connecticut). 

striaticornis (Kieffer), 1918a: 36 (Ceratopogon). Algeria. 

nigerrima Goetghebuer, 1933e: 131. Democratic Republic of the Congo. 

iphias de Meillon, 1936: 169. South Africa. 

suberis Clastrier, 1956: 496. Algeria. 

flavirustica Remm, in Remm and Zhogolev, 1968: 827. Ukraine. 

subfrigidus Liu and Yu, in Liu et al., 2001b: 154. China. 

subgedanicola Szadziewski, 1993: 645. Germany. Eocene. 

subpallida Tokunaga, 1962a: 181. Japan. 

39

subruralis Liu and Yu, in Liu et al., 2001b: 156. China. 

subtilis Johannsen, 1932: 412. Indonesia. 

subulipenis Sinha, Mazumdar, and Chaudhuri, 2003: 77. India. 

sudanensis Macfie, 1947b: 70. Sudan. 

supplex Debenham, 1987c: 297. Australia (Queensland). 

surculus Liu and Yu, in Liu et al., 2001b: 157. China. 

swezeyana Tokunaga and Murachi, 1959: 145. Guam (USA). 

swezeyanaadfinis Chan and LeRoux, 1971b: 747. Singapore. 

tangae Kieffer, 1913e: 6. Tanzania. 

tapleyi Ingram and Macfie, 1931b: 196. New Zealand. 

taragui Marino, Spinelli, and Cazorla, 2002: 17. Argentina. 

guarani Marino and Spinelli, 1999: 448 (preoccupied by Forcipomyia guarani Ronderos and Spinelli, 1999). 

tauffliebi Clastrier, 1960c: 515. Congo. 

tavetae Kieffer, 1913e: 6. Kenya. 

tenuis (Winnertz), 1852: 25 (Ceratopogon). Germany. 

tenuisquamipes Wirth, 1952b: 89. Chile. 

tetraclada Kieffer, 1912c: 2. Sri Lanka. 

tetrasticta (Kieffer), 1919a: 11 (Ceratopogon). Hungary. 

texana (Long), 1902: 10 (Ceratopogon). USA (Texas). 

theobromae Kieffer, 1912c: 3. Sri Lanka. 

thienemanni Kieffer, 1912b: 103. France. 

tienshanica Remm, 1980: 120. Kazakhstan. 

tigripes Ingram and Macfie, 1924b: 580. Ghana. 

tortor Debenham, 1987c: 286. Australia (New South Wales). 

tortula Liu and Yu, in Liu et al., 2001b: 160. China. 

townesi Wirth, 1952a: 137. USA (California). 

townsendi Knab, 1915: 111. Peru. 

trigonata Johannsen, 1932: 412. Indonesia. 

trilineata Goetghebuer, 1934a: 90. Russia, Austria. 

trinotata Kieffer, 1913d: 169. India. 

tristicta Kieffer, 1913d: 169. India. 

turanorustica Remm, 1980: 118. Uzbekistan. 

turbinata (Meunier), 1904a: 227 (1904b: 238) (Ceratopogon). Baltic region. Eocene. 

uncula (Meunier), 1904a: 227 (1904b: 238) (Ceratopogon). Baltic region. Eocene. 

unculiformis Szadziewski, 1993: 646. Germany. Eocene. 

unimaculata Sinha, Mazumdar, and Chaudhuri, 2003: 78. India. 

uramaensis Spinelli and Dippolito, 1995: 156. Venezuela. 

urbana Goetghebuer, 1937: 273. Belgium. 

urnigera Kieffer, 1925e: 247. Egypt. 

utae Knab, 1915: 109. Peru. 

variicrus Kieffer, 1913d: 172 (as variety of albosignata). India. 

venetiana (Kieffer), 1919a: 12 (Ceratopogon). Italy. 

ventralis Borkent, in Borkent and Wirth, 1997:38. New name for abdominalis. 

abdominalis Tokunaga, 1940c: 87 (preoccupied by Forcipomyia abdominalis (Santos Abreu), 1918). Japan. 

vesicula de Meillon and Wirth, 1983a: 350. South Africa. 

vexillaria Debenham, 1987c: 291. Australia (New South Wales). 

villiersi Clastrier, 1972a: 98. Congo. 

vittata Tokunaga, 1940c: 71. Japan. 

wahgi Debenham, 1987c: 285. Papua New Guinea. 

waldemari Szadziewski, 1983a: 370. Algeria. 

waldenia de Meillon, 1940: 464. South Africa. 

wheeleri (Long), 1902: 12 (Ceratopogon). USA (Texas). 

wygodzinskyi Cavalieri, 1961a: 17. Argentina. 

delpontei Cavalieri, 1961b: 169. Argentina. 

40

yamana Marino and Spinelli, 2001a: 118. Chile. 

yamauchii Tokunaga, 1940c: 84. Japan. 

youngi Wirth, 1982a: 579. Panama. 

zhangmuensis Liu and Yu, in Liu et al., 2001b: 162. China. 

zhongshanensis Liu and Yu, in Liu et al., 2001b: 164. China. 

zonogaster Ingram and Macfie, 1931a: 168. Chile. 

Subgenus GAMPSOHELEA Yu and Liu 

GAMPSOHELEA Yu and Liu, 2006: 584 (as subgenus of Forcipomyia). Type species: Forcipomyia dentapenis Yu and Liu, 


dentapenis Yu and Liu, in Yu et al., 2005: 584. China. 

Subgenus HERAKLEOHELEA Debenham 

HERAKLEOHELEA Debenham, 1987a: 97 (as subgenus of Forcipomyia). Type species: Forcipomyia grallator Debenham, 


grallator Debenham, 1987a: 97. Australia (New South Wales). 

Subgenus IXODEHELEA Yu and Liu 

IXODEHELEA Yu and Liu, 2006: 586 (as subgenus of Forcipomyia). Type species: Forcipomyia ambiguous Yu and Liu, by 


ambiguoa Yu and Liu, in Yu et al., 2005: 586. China. 

Subgenus JAPYGAHELEA Yu and Liu 

JAPYGAHELEA Yu and Liu, 2006: 588 (as subgenus of Forcipomyia). Type species: Forcipomyia pinnatus Yu and Liu, by 


pinnata Yu and Liu, in Yu et al., 2005: 588. China. 

Subgenus KATTANGOMYIA Debenham 

KATTANGOMYIA Debenham, 1987a: 98 (as subgenus of Forcipomyia). Type species: Forcipomyia taura Debenham, by 


orbis Debenham, 1987a: 102. Australia (New South Wales). 

taura Debenham, 1987a: 101. Australia (New South Wales). 

Subgenus LASIOHELEA Kieffer 

CENTRORHYNCHUS Lutz, 1913: 62 (preoccupied by Centrorhynchus Steven or Fisher Waldheim, 1829). Type species: 

Centrorhynchus stylifer Lutz, by original designation. 

LASIOHELEA Kieffer, 1921h: 115. Type species: Atrichopogon pilosipennis Kieffer (= Ceratopogon velox Winnertz), 


PARAPTEROBOSCA Harant, Huttel and Huttel, 1951: 468. Type species: Parapterobosca anthropophila Harant, Huttel and 

Huttel, by original designation. 

DACNOFORCIPOMYIA Chan and Saunders, 1965: 527 (as subgenus of Forcipomyia). Type species: Forcipomyia 

anabaenae Chan and Saunders, by original designation. 

41

abditan Yu, in Yu et al., 2005: 694 (Lasiohelea). China. 

agas (Rondani), 1875: 462 (Ceratopogon). Malaysia. 

aiyurae Debenham, 1983: 29. Papua New Guinea. 

alia Yu and Yan, 2007: 45 (Lasiohelea). Malaysia. 

americana Szadziewski and Grogan, 1998b: 270. Dominican Republic. Miocene. 

anabaenae Chan and Saunders, 1965: 528. Singapore. 

aeschrodenta (Yu and Liu), 1981: 61 (Lasiohelea). China. 

angyria (Yu and Wirth), 1997: 21 (Lasiohelea). Malaysia. 

anitae Huerta and Ibanez-Bernal, 1996: 350. Mexico. 

ancoriformis Tokunaga, in Tokunaga and Murachi, 1959: 226. Micronesia. 

anthropophila (Harant, Huttel and Huttel), 1951: 468 (Parapterobosca). Ivory Coast. 

araea (Yu and Wirth), 1997: 23 (Lasiohelea). Malaysia. 

attenuata Saunders, 1964: 469. Costa Rica. 

baltea Boorman, 1990: 128. Oman. 

bambusa (Liu and Yu), 1996: 50 (Lasiohelea). China. 

bernicla (Yu and Wirth), 1997: 24 (Lasiohelea). Malaysia. 

bidenta (Yu and Wirth), 1997: 26 (Lasiohelea). Malaysia. 

bifidipenis (Yu and Wirth), 1997: 27 (Lasiohelea). Malaysia. 

binifoliaceus (Yu and Wirth), 1997: 29 (Lasiohelea). Malaysia. 

bispica (Yu and Wirth), 1997: 30 (Lasiohelea). Malaysia. 

bladapida Yu and Yan, 2007: 46 (Lasiohelea). Malaysia. 

bonasa (Yu and Wirth), 1997: 31 (Lasiohelea). Malaysia. 

boophila Lien, 1991: 85. Taiwan. 

borneoensis (Yu and Wirth), 1997: 32 (Lasiohelea). Malaysia. 

breviprobosca Yu, in Yu et al., 2005: 748 (Lasiohelea). China. 

brevisicae Debenham, 1983: 26. Australia (New South Wales). 

brevitarsata (Ingram and Macfie), 1924a: 385 (Lasiohelea). Ghana. 

cacaophila Ronderos and Spinelli, 1999: 152. Venezuela. 

caelomacula Liu, Yan and Liu, 1996: 10 (Lasiohelea). China. 

caesariata Debenham, 1983: 8. Australia (Queensland). 

caledonica Clastrier and Delécolle, 1993: 132. New Caledonia (France). 

carolinensis (Tokunaga), 1940a: 208 (Lasiohelea). Micronesia. 

chiengmai (Yu and Wirth), 1997: 35 (Lasiohelea). Thailand. 

citowitschi Debenham, 1983: 17. Australia (Queensland). 

collessi (Yu and Wirth), 1997: 36 (Lasiohelea). Malaysia. 

collicola Lien, 1989: 43. Taiwan. 

colum Debenham, 1983: 7. Papua New Guinea. 

cornuta Saunders, 1964: 464. Costa Rica. 

crocea Debenham, 1983: 15. Papua New Guinea. 

cubitalis (Kieffer), 1918a: 88 (Ceratopogon). Singapore. 

cuculli Debenham, 1983: 34. Australia (Queensland). 

cultella (Yu and Xiang), in Yu, 1988: 127 (Lasiohelea, as Lasoihelea). China. 

cymodocea Yu and Nie, in Nie et al., 2003: 240 (Lasiohelea). China (on ship from Davao, Philippines). 

dandongensis (Ding and Yu), 1990: 53 (Lasiohelea). China. 

danxianensis (Yu and Liu), 1982a: 25 (Yu and Liu, 1982b: 306; Yu, 1982: 203) (Lasiohelea). China. 

diaoluoensis (Yu and Liu), 1982a: 26 (Yu and Liu, 1982b: 306; Yu, 1982: 203) (Lasiohelea). China. 

dirus (Liu, Yan and Liu), 1996: 11 (Lasiohelea). China. 

divergena (Yu and Wen), in Yu and Liu, 1982a: 28 (Lasiohelea). China. 

emeishana (Yu and Liu), 1982a: 31 (Yu and Liu, 1982b: 300) (Lasiohelea). China. 

eminenta Yu, in Yu et al., 2005: 761 (Lasiohelea). China. 

fairfaxensis Wirth, 1951d: 317. USA (Virginia). 

fenestrarum Debenham, 1983: 25. Australia (New South Wales). 

fengyani Yu, in Yu et al., 2005: 710 (Lasiohelea). China. 

ficula (Yu and Wirth), 1997: 41 (Lasiohelea). Malaysia. 

42

flaveola Clastrier and Delécolle, 1993: 136. New Caledonia (France). 

flavescens Saunders, 1964: 471. Philippines. 

forficula (Yu and Wirth), 1997: 44 (Lasiohelea). Malaysia. 

gracilidenta Yu, in Yu et al., 2005: 762 (Lasiohelea). China. 

gramencola Lien, 1991: 90. Taiwan. 

gripha Borkent, in Borkent and Wirth, 1997:39. New name for paradoxa. 

paradoxa (Yu and Liu), 1990: 16 (Lasiohelea, preoccupied by Forcipomyia paradoxa Krivosheina, 1968). China. 

guangxiensis (Lee), 1975: 435 (Lasiohelea). China. 

guarani Ronderos and Spinelli, 1999: 153. Paraguay. 

habros Chen, Liu and Yu, 2012: 34 (Lasiohelea). China. 

hainana (Liu, Yan and Liu), 1996: 12 (Lasiohelea). China. 

hicksae Debenham, 1983: 30. Australia (Queensland). 

homaliae Lien, 1991: 90. Taiwan. 

hortensis (Yu and Liu), 1981: 62 (Lasiohelea). China. 

hualuensis Lien, 1989: 49. Taiwan. 

humilavolita (Yu and Liu), 1982a: 34 (Yu and Liu, 1982b: 302) (Lasiohelea). China. 

hygroecia Yu, Liang and Chen, in Chen et al., 2007: 171 (Lasiohelea). China. 

incomposita Debenham, 1983: 20. Papua New Guinea. 

infans Debenham, 1983: 28. Australia (Queensland). 

insigniforceps Macfie, 1939c: 165. Brazil. 

interceda Yu, in Yu et al., 2005: 717 (Lasiohelea). China. 

intermedia Saunders, 1964: 468. Costa Rica. 

jinileei Yu, in Yu et al., 2005: 718 (Lasiohelea). China. 

kamilaroi Debenham, 1983: 29. Australia (New South Wales). 

koba Yu, in Yu et al., 2005: 719 (Lasiohelea). China. 

labidentis (Yu and Liu), 1982a: 37 (Yu and Zhang, 1982: 187) (Lasiohelea). China. 

lanyuensis Lien, 1989: 54. Taiwan. 

latifolia (Yu and Wirth), 1997: 48 (Lasiohelea). Malaysia. 

latiunguis Clastrier, 1983c: 47. Seychelles. 

lefanui Carter, 1916: 131. Ghana. 

atratula Goetghebuer, 1935d: 149. Democratic Republic of the Congo. 

atrosetosa Goetghebuer, 1935d: 149. Democratic Republic of the Congo. 

squamipes (Ingram and Macfie), 1924a: 385 (Lasiohelea, as variety of lefanui, preoccupied by Forcipomyia 

squamipes (Coquillett), 1902a). Probably Tanzania. 

lepta (Yu and Wirth), 1997: 51 (Lasiohelea). Malaysia. 

longicornis (Tokunaga), 1940d: 103 (Lasiohelea). Japan. 

longineura Saunders, 1964: 470. Philippines. 

longipalpula (Yu and Wirth), 1997: 54 (Lasiohelea). Malaysia. 

lui (Liu, Yan and Liu), 1996: 14 (Lasiohelea). China. 

lushana (Yu and Liu), 1982a: 38 (Yu and Wang, 1982: 19; Yu, 1982: 203) (Lasiohelea). China. 

luzona (Yu and Wirth), 1997: 56 (Lasiohelea). Philippines. 

magnispinosa Clastrier and Delécolle, 1993: 137. New Caledonia (France). 

mahensis (Kieffer), 1911c: 339 (Ceratopogon). Seychelles. 

manasi Maheshwari, 2003: 72. India. 

matautuensis Clastrier and Delécolle, 1996: 304. Wallis and Futuna Islands (France). 

matsumurai Kitaoka, 1994: 3. Japan. 

megadentis Lien, 1989: 57. Taiwan. 

mengi (Yu and Liu), in Liu and Yu, 1996: 49 (Lasiohelea). China. 

ministra Debenham, 1983: 32. Australia (Northern Territory). 

mixta (Yu and Liu), 1982a: 40 (Yu and Liu, 1982b: 301) (Lasiohelea). China. 

montana de Meillon and Downes, 1986: 147. South Africa. 

mopsa de Meillon and Hardy, 1954: 71. South Africa. 

moriokensis Kitaoka, 1994: 1. Japan. 

multidentata Ronderos and Spinelli, 1999: 153. Brazil. 

43

multidentis Yu, in Yu et al., 2005: 727 (Lasiohelea). China. 

multisensora Yu, in Yu et al., 2005: 773 (Lasiohelea). China. 

multispina (He and Yu), 1990: 63 (Lasiohelea). China. 

nanjingensis (Yu and Liu), 1982a: 41 (Yu and Wang, 1982: 20) (Lasiohelea). China. 

nasuensis Kitaoka, 1994: 3. Japan. 

nema Debenham, 1983: 19. Australia (Queensland). 

nemerosa Yu, in Yu et al., 2005: 778 (Lasiohelea). China. 

neotokunagai Szadziewski and Borkent, 2003: 255. New name for tokunaga. 

tokunaga (Yu and Wirth), 1997: 77 (Lasiohelea)(preoccupied by Forcipomyia tokunagai (Oka and Asahina), 1948). 

Malaysia. 

neowirthi Szadziewski and Borkent, 2003: 255. New name for wirthi. 

wirthi (Yu and Wirth), 1997: 80 (preoccupied by Forcipomyia wirthi Saunders, 1957). Malaysia. 

nepala (Yu), 2000a: 12 (Lasiohelea, misspelled as nepaia). Nepal. 

nhulunbuyensis Debenham, 1983: 32. Australia (Northern Territory). 

nigeriae (Ingram and Macfie), 1924a: 382 (Lasiohelea). Nigeria. 

dewulfi (Goetghebuer), 1933e: 136 (Lasiohelea). Democratic Republic of the Congo. 

nipponica (Tokunaga), 1940d: 102 (Lasiohelea). Japan. 

oreita (Liu and Yu), 1996: 51 (Lasiohelea). China. 

ostiola (Yu and Wirth), 1997: 60 (Lasiohelea). Philippines. 

oxypenis (Yu and Wirth), 1997: 62 (Lasiohelea). Malaysia. 

oxyria (Yu and Liu), in Yu et al., 1985: 72 (Lasiohelea). China. 

paenedentula Debenham, 1983: 16. Australia (Queensland). 

parenthesis Debenham, 1983: 24. Australia (Queensland). 

paucidentis Lien, 1991: 98. Taiwan. 

parvitas (Liu and Yu), 1996: 49 (Lasiohelea). China. 

pensiledentia (Yu and Wirth), 1997: 63 (Lasiohelea). Malaysia. 

perae Debenham, 1983: 23. Solomon Islands. 

perangusta (Goetghebuer), 1935d: 168 (Dasyhelea). Democratic Republic of the Congo. 

peregrinator Debenham, 1983: 12. Australia (Queensland). 

phototropia (Yu and Liu), 1982a: 45 (Yu and Zhang, 1982: 189) (Lasiohelea). China. 

plumosa Debenham, 1983: 9. Papua New Guinea. 

propria Chan and LeRoux, 1970: 272. Malaysia. 

pungobovis (Yu and Liu), 1982a: 48 (Lasiohelea). China. 

putea Debenham, 1983: 14. Australia (Queensland). 

quadrangula Yu and Yan, 2007: 45 (Lasiohelea). Malaysia. 

quasicornuta Saunders, 1964: 467. Costa Rica. 

quinquedentis (Yu and Zhou), in Yu, 1988: 128 (Lasiohelea). China. 

rastraria Debenham, 1983: 10. Australia (Northern Territory). 

relicta (Yu and Wen), in Yu and Liu, 1982a: 50 (Lasiohelea). China. 

rhamphis (Yu and Wirth), 1997: 67 (Lasiohelea). Malaysia. 

richardlanei Clastrier, 1983c: 50. Seychelles. 

ripa (Yu and Liu), 2000b:125 (Lasiohelea). China. 

russelli Debenham, 1983: 11. Australia (Queensland). 

saltensis (Cavalieri), 1962: 360 (Lasiohelea). Argentina. 

saxicola Lien, 1991: 98. Taiwan. 

serridentata Debenham, 1983: 18. Solomon Islands. 

sibirica (Buyanova), 1962: 43 (Lasiohelea). Russia. 

sirycta (Yu and Liu), 2000b:126 (Lasiohelea). China. 

spinipenis Tokunaga, in Tokunaga and Murachi, 1959: 227. Belau (USA). 

squamitarsata (Clastrier), 1959b: 351 (Lasiohelea). Senegal. 

stellaris (Yu and Wirth), 1997: 68 (Lasiohelea). Malaysia. 

stimulans (de Meijere), 1909: 197 (Ceratopogon). Indonesia. 

stylifer (Lutz), 1913: 63 (Centrorhynchus). Brazil. 

succinea Szadziewski, 1988: 194. Poland. Eocene. 

44

taipei Lien, 1991: 102. Taiwan. 

taiwana (Shiraki), 1913: 291 (Ceratopogon). Taiwan. 

notialis (Yu and Liu), 1982a: 43 (Lasiohelea). China. 

tambunana (Yu and Wirth), 1997: 73 (Lasiohelea). Malaysia. 

taoyuanensis Lien, 1989: 72. Taiwan. 

tawauensis (Yu and Wirth), 1997: 74 (Lasiohelea). Malaysia. 

tenuidentis (Yu and Wirth), 1997: 75 (Lasiohelea). Malaysia. 

thabinana de Meillon, 1959b: 6. South Africa. 

thiensis Clastrier and Delécolle, 1993: 139. New Caledonia (France). 

thyesta Yu, Chen and He, in Chen et al., 2007: 172 (Lasiohelea). China. 

tibetana Yu, in Yu et al., 2005: 738 (Lasiohelea). China. 

townsvillensis (Taylor), 1918: 169 (Culicoides). Australia (Queensland). 

turgepeda (Yu and Liu), 1982a: 52 (Yu and Liu, 1982b: 304) (Lasiohelea). China. 

uncusidentis (Liu, Yan and Liu), 1996: 16 (Lasiohelea). China. 

uncusipenis (Yu and Liu), 1982a: 52 (Yu and Zhang, 1982: 187) (Lasiohelea). China. 

uncuspromissa Chan and LeRoux, 1970: 276. Singapore. 

ussurica Remm, 1971: 190. Russia. 

velox (Winnertz), 1852: 28 (Ceratopogon). Germany. 

halterata (Winnertz), 1852: 28 (Ceratopogon). Germany. 

nitens (Kieffer), 1919a: 20 (Ceratopogon, preoccupied by Forcipomyia nitens (Santos Abreu), 1918). Hungary. 

pilosipennis (Kieffer), 1919a: 23 (Atrichopogon). Hungary, Slovak Republic. 

hungarica (Kieffer), 1921d: 298 (Ceratopogon). New name for nitens. 

decrescens Kieffer, 1924b: 391. France. 

silesiae (Kieffer), 1925d: 49 (Lasiohelea). Hungary. 

montschadskyi (Dzhafarov), 1962c: 196 (Lasiohelea). Azerbaijan. 

virgula (Yu and Wen), in Yu et al., 1985: 73 (Lasiohelea). China. 

wanjungensis Lien, 1989: 72. Taiwan. 

whitcombei Boorman, 1990: 126. Oman. 

woodruffi Szadziewski and Grogan, 1998b: 272. Dominican Republic. Miocene. 

wulai Lien, 1991: 109. Taiwan. 

wuyiensis (Shen and Yu), 1990: 66 (Lasiohelea). China. 

yui (Liu, Yan and Liu), 1996: 13 (Lasiohelea). China. 

zhenbaodaoensis (Yu and Liu), 1987: 83 (Lasiohelea). China. 

zonaphalla (Yu and Liu), 1982a: 53 (Yu and Liu, 1982b: 305) (Lasiohelea). China. 

Subgenus LEPIDOHELEA Kieffer 

LEPIDOHELEA Kieffer, 1917b: 364. Type species: Ceratopogon chrysolophus Kieffer, by original designation. 

abercrombyi Macfie, 1938: 161. Trinidad and Tobago. 

acinacium Debenham, 1987c: 329. Australia (New South Wales). 

acinacis Wirth and Spinelli, 1993b: 615. USA (Maryland). 

adversari Debenham, 1987c: 321. Australia (Queensland). 

anguliforceps Tokunaga, 1940c: 94. Japan. 

annulatipes Macfie, 1939c: 154. Brazil. 

antilleana Szadziewski and Grogan, 1998b: 274. Dominican Republic. Miocene. 

articulatus Ma and Yan, in Ma et al., 2000:15. China. 

bahiensis Wirth and Spinelli, 1993a: 113. Brazil. 

basendjiorum Dessart, 1962: 140. Democratic Republic of the Congo. 

basifemoralis Wirth and Spinelli, 1993a: 113. Jamaica. 

beckae Wirth, 1976b: 82. USA (Florida). 

bicolor Lutz, 1914: 89. Brazil. 

discoloripes Macfie, 1939c: 159. Brazil. 

bifida Wirth and Spinelli, 1993a: 115. Jamaica. 

45

ifurcifera Tokunaga, 1959: 281. Papua New Guinea. 


bullata Debenham, 1987c: 315. Australia (Capital Territory). 

cacaoi Dessart, 1963: 186. New name for theobromae. 

theobromae Dessart, 1961: 367 (preoccupied by Forcipomyia theobromae Kieffer, 1912c). Democratic Republic of 

Congo. 

calotricha Kieffer, 1911b: 322. India. 

cerbera Debenham, 1987c: 335. Australia (New South Wales). 

christiansoni Wirth and Hubert, 1960a: 640. USA (California). 

chrysolopha (Kieffer), 1911c: 333 (Ceratopogon). Seychelles. 

grata Goetghebuer, 1935d: 153 (preoccupied by Forcipomyia grata Macfie, 1934c). Democratic Republic of the 

Congo. 

guttatella Goetghebuer, 1935d: 146 (also as guttatula). Democratic Republic of the Congo. 

annulator Goetghebuer, 1948b: 6. New name for grata. 

chrysosuccinea Szadziewski and Grogan, 1998b: 274. Dominican Republic. Miocene. 

cochisei Wirth and Spinelli, 1993b: 618. USA (Arizona). 

convexipenis Wirth and Spinelli, 1993a: 116. Colombia. 

davidi Debenham, 1987c: 330. Australia (New South Wales). 

demeter Debenham, 1987c: 319. Australia (Victoria). 

domibicolor Szadziewski and Grogan, 1998b: 276. Dominican Republic. Miocene. 

dubia Macfie, 1939c: 162. Brazil. 

dubiamima Wirth and Spinelli, 1993b: 620. USA (Maryland). 

eadsi Wirth and Spinelli, 1993b: 622. USA (Texas). 

edmistoni Wirth and Spinelli, 1993b: 624. USA (Maryland). 

euthystyla Wirth and Spinelli, 1993a: 117. Colombia. 

flavifemoris Macfie, 1940d: 24. Guyana. 

grata Macfie, 1934c: 178 (1934d: 203). Malaysia. 

gravesi Wirth and Spinelli, 1993a: 119. USA (North Carolina). 

hastata de Meillon and Wirth, 1981c: 565. South Africa. 

herediae Wirth and Spinelli, 1993a: 119. Costa Rica. 

hobbsi Wirth and Spinelli, 1993a: 121. Dominica. 

ikinae de Meillon, Meiswinkel and Wirth, 1982: 127. South Africa. 

kuanoskeles Macfie, 1939c: 150. Brazil. 

lacrimatorii Macfie, 1939c: 161. Brazil. 

lagoenae Debenham, 1987c: 327. Australia (New South Wales). 

lepidosuccinea Szadziewski and Grogan, 1998b: 278. Dominican Republic. Miocene. 

luteigenua Wirth and Spinelli, 1992: 353. Costa Rica. 

maculatus Liu, Yan and Liu, 1996: 20. China. 

maculosicrura Tokunaga, 1959: 265. Papua New Guinea. 

manis Debenham, 1987c: 344. Papua New Guinea. 

matilei Clastrier and Delécolle, 1991: 203. New Caledonia (France). 

medipala Debenham, 1987c: 320. Australia (New South Wales). 

paenulata Debenham, 1987c: 334. Australia (Western Australia). 

palliscuta Tokunaga, 1959: 259. Papua New Guinea. 

pampoikila Ingram and Macfie, 1924b: 570. Ghana. 

parvicrater Debenham, 1987c: 336. Australia (Capital Territory). 

pectinis Liu and Yu, in Liu et al., 2001b: 172. China. 

pictisquamipennis Tokunaga, 1959: 260. Papua New Guinea. 

pricei Wirth and Spinelli, 1993b: 627. USA (Texas). 

pulcherrima Santos Abreu, 1918: 272. Canary Islands (Spain). 

formosae (Kieffer), 1922b: 153 (Lepidohelea). Taiwan. 

lepidota Ingram and Macfie, 1924b: 566. Ghana. 

ornatipes (Kieffer), 1921b: 1 (Lepidohelea, preoccupied by Forcipomyia ornatipes (Kieffer), 1918a). Cameroon. 

variegata Goetghebuer, 1933e: 133. Democratic Republic of the Congo. 

46

marsafae Ghonaim, Ibrahim and Ali, 2001: 42. Egypt. 

qionghainensis Liu and Yu, in Liu et al., 2001b: 177. China. 

quinqueremis Debenham, 1987c: 341. Papua New Guinea. 

randensis de Meillon, 1931: 335. South Africa. 

randensoides Dessart, 1961: 364. Democratic Republic of the Congo. 

roseae de Meillon, Meiswinkel and Wirth, 1982: 129. South Africa. 

scurra Debenham, 1987c: 340. Australia (Queensland). 

semihamata Debenham, 1987c: 339. Australia (New South Wales). 

seminole Wirth, 1976b: 81. USA (Florida). 

semota Debenham, 1987c: 328. Australia (Queensland). 

solutus Liu and Yu, in Liu et al., 2001b: 179. China. 

squamipennis Ingram and Macfie, 1924b: 562. Ghana. 

squamithorax Clastrier, 1972b: 170. French Guiana. 

statirae de Meillon, 1936: 165. South Africa. 

tibialis Remm, 1961a: 188. Estonia. 

tortuosa Debenham, 1987c: 322. Australia (New South Wales). 

unifascicornis Tokunaga, 1959: 263. Papua New Guinea. 

unitheca Tokunaga, in Tokunaga and Murachi, 1959: 183. Belau (USA). 

usingeri Wirth and Spinelli, 1993b: 627. USA (California). 

varipennis Wirth and Williams, 1957: 8. Bermuda (Great Britain). 

venusta Ingram and Macfie, 1924b: 569. Ghana. 

weemsi Wirth and Spinelli, 1993a: 123. USA (Florida). 

werneri Wirth and Spinelli, 1993b: 632. Mexico. 

winderi Wirth, 1991b: 171. Brazil. 

xichangensis Liu and Yu, in Liu et al., 2001b: 181. China. 

Subgenus METAFORCIPOMYIA Saunders 

METAFORCIPOMYIA Saunders, 1957: 685 (as subgenus of Forcipomyia). Type species: Forcipomyia cerifera Saunders, 


albipluma Spinelli, Marino and Borkent, 2012: 27. Costa Rica. 

anniae Spinelli, Marino and Borkent, 2012: 11. Costa Rica. 

atenasensis Spinelli, Marino and Borkent, 2012: 13. Costa Rica. 

campana Debenham, 1987b: 193. Australia (New South Wales). 

cerifera Saunders, 1957: 685. Brazil. 

colona Debenham, 1987b: 191. Australia (Capital Territory). 

crepidinis Debenham, 1987b: 196. Australia (Western Australia). 

darwini Marino and Spinelli, 2003: 24. Chile. 

defatigatus Liu and Yu, in Liu et al., 2001b: 183. China. 

fehrerorum Grogan and Sigrist, 2007: 531. USA (Maryland). 

furculae Debenham, 1987b: 190. Papua New Guinea. 

galliarii Marino and Spinelli, 1999: 5. Argentina. 

germinata Spinelli, Marino and Borkent, 2012: 29. Costa Rica. 

grandiseta Spinelli, Marino and Borkent, 2012: 16. Costa Rica. 

heroni Spinelli, Marino and Borkent, 2012: 31. Costa Rica. 

hesiones de Meillon, 1936: 167. South Africa. 

longiflagellata Spinelli, Marino and Borkent, 2012: 18. Costa Rica. 

macroseta Spinelli, Marino and Borkent, 2012: 20. Costa Rica. 

maculosa Ingram and Macfie, 1931a: 159. Argentina. 

mapuche Marino and Spinelli, 2003: 26. Argentina. 

morenoi Marino and Spinelli, 2003: 26. Argentina. 

novaguineae Tokunaga, 1959: 289. Papua New Guinea. 

osaensis Spinelli, Marino and Borkent, 2012: 15. Costa Rica. 

47

pluvialis Malloch, 1923: 5. USA (Maryland). 

pseudocerifera Spinelli, Marino and Borkent, 2012: 21. Costa Rica. 

rivalis Spinelli, Marino and Borkent, 2012: 7. Costa Rica. 

ronderosae Spinelli, Marino and Borkent, 2012: 33. Costa Rica. 

rupicola Debenham, 1987b: 194. Australia (New South Wales). 

rursa Spinelli, Marino and Borkent, 2012: 24. Costa Rica. 

stewarti de Meillon and Downes, 1986: 149. South Africa. 

stigmatipennis Tokunaga, 1959: 268. Papua New Guinea. 

tomaculorum Debenham, 1987b: 188. Australia (New South Wales). 

totus Liu and Yu, in Liu et al., 2001b: 185. China. 

triclotae Clastrier and Delécolle, 1991: 209. New Caledonia (France). 

truncata Spinelli, Marino and Borkent, 2012: 9. Costa Rica. 

williamsi Marino and Spinelli, 1999: 5. Argentina. 

Subgenus MICROHELEA Kieffer 

MICROHELEA Kieffer, 1917b: 364. Type species: Atrichopogon microtomus Kieffer, designation by Kieffer, 1921b: 7, with 

explanatory notation on erroneous previous type designation). 

PHASMIDOHELEA Mayer, 1937b: 233. Type species: Phasmidohelea crudelis Mayer (= Forcipomyia mayeri Forattini and 

Lane), by original designation. 

abbaekiefferi Debenham, 1987d: 656. Papua New Guinea. 

alleni Clastrier and Wirth, 1995: 117. Costa Rica. 

almus Liu and Yu, in Liu et al., 2001b: 190. China. 

amazonica Wirth, 1971b: 241. Brazil. 

ambientis Liu and Yu, in Liu et al., 2001b: 192. China. 

amieuensis Clastrier and Delécolle, 1993: 149. New Caledonia (France). 

angustipalpis Saha, Dasgupta, Gangopadhyay and Mukherjee, 2009: 30. India. 

antioquiae Clastrier and Wirth, 1995: 142. Colombia. 

arnhemi Debenham, 1987d: 652. Australia (Northern Territory). 

asquamata Clastrier and Delécolle, 1993: 142. New Caledonia (France). 

australiensis (Kieffer), 1917a: 178 (Ceratopogon). Papua New Guinea. 

beatulus Liu and Yu, in Liu et al., 2001b: 193. China. 

belemensis Clastrier and Wirth, 1995: 140. Brazil. 

bispinula Liu and Yu, 1997: 26. China. 

brasiliana Clastrier and Wirth, 1995: 116. Peru. 

breelandi Clastrier and Wirth, 1995: 114. Panama. 

brevilabellata Clastrier and Wirth, 1995: 136. French Guiana. 

broadheadi Clastrier and Wirth, 1995: 129. Panama. 

callithorax (Kieffer), 1911c: 335 (Ceratopogon, as variety of lasionota). Seychelles. 

castneri Clastrier and Wirth, 1995: 111. Peru. 

catarina Clastrier and Wirth, 1995: 131. Brazil. 

coheni Clastrier and Wirth, 1995: 132. Ecuador. 

colombiana Clastrier and Wirth, 1995: 102. Colombia. 

cristata Clastrier and Wirth, 1995: 135. Brazil. 

cylindripalpis Clastrier and Delécolle, 1993: 147. New Caledonia (France). 

desutterae Clastrier and Wirth, 1995: 100. French Guiana. 

donskoffi Clastrier and Wirth, 1995: 125. French Guiana. 

dunklei Clastrier and Wirth, 1995: 130. Peru. 

eriophora (Williston), 1896: 279 (Ceratopogon). St. Vincent. 

esakiana Tokunaga, 1940a: 206. Micronesia. 

felippebauerae Clastrier and Wirth, 1995: 122. Peru. 

flavitibialis Tokunaga and Murachi, 1959: 170. Micronesia. 

48

foliacceus Liu and Yu, in Liu et al., 2001b: 199. China. 

folipenis Liu and Yu, 1997: 24. China. 

franklini Clastrier and Wirth, 1995: 141. Costa Rica. 

fuliginata Clastrier, 1983c: 54. Seychelles. 

fuliginosa (Meigen), 1818: 86 (Ceratopogon). Germany. 

villosa (Zetterstedt), 1850: 3645 (Ceratopogon). Sweden. 

crudelis (Karsch), 1886: 18 (Ceratopogon). Germany. 

hirtipes (de Meijere), 1907: 209 (Ceratopogon). Indonesia. 

brevimana Lundström, 1910: 32. Finland. 

inornatipennis (Austen), 1912: 107 (Ceratopogon). Nigeria. 

erucicida Knab, 1914: 65. USA (Florida). 

crudelis Knab, 1914: 66 (preoccupied by Forcipomyia crudelis (Karsch), 1886). Mexico. 

coquilletti (Kieffer), 1917b: 297 (Ceratopogon). USA (New York). 

tropica (Kieffer), 1917b: 297 (Ceratopogon). Costa Rica. 

obscura Santos Abreu, 1918: 276 (as variety of bipunctata, preoccupied by Forcipomyia obscura (Walker), 1848). 

Canary Islands (Spain). 

alboclavata (Kieffer), 1919a: 12 (Ceratopogon, also as asticta). Slovak Republic. 

canaliculata (Goetghebuer), 1920: 110 (Ceratopogon). Belgium. 

nilotheres Macfie, 1924: 62. Egypt. 

ornaticrus Ingram and Macfie, 1924b: 577 (as variety of inornatipennis; preoccupied by Forcipomyia ornaticrus 

Kieffer, 1912c). Ghana. 

atripennis Goetghebuer, 1935d: 149. Democratic Republic of the Congo. 

auripila Goetghebuer, 1935d: 150. Democratic Republic of the Congo. 

curtimana Goetghebuer, 1935d: 151. Democratic Republic of the Congo. 

grisescens Goetghebuer, 1935d: 154. Democratic Republic of the Congo. 

vicina Goetghebuer, 1935d: 161. Democratic Republic of the Congo. 

longiradialis Tokunaga, 1940c: 66. Japan. 

takagii Tokunaga, 1941b: 90. China. 

wansoni (Harant and Baur), 1946: 141 (Lasiohelea, preoccupied by Forcipomyia wansoni de Meillon, 1939a). 

Democratic Republic of the Congo. 

brookmani Wirth, 1952a: 140. USA (California). 

santosi Remm, 1981: 32. New name for obscura. 

galapagensis (Coquillett), 1901b: 372 (Ceratopogon). Galápagos Islands (Ecuador). 

galbiventris Borkent, in Borkent and Wirth, 1997:43. New name for flaviventris. 

flaviventris (Kieffer), 1917a: 179 (Ceratopogon, preoccupied by Palpomyia flaviventris (Czerny and Strobl), 1909). 

Papua New Guinea. 

globularis Edwards, 1928: 48. Western Samoa. 

gracilalitheca Saha, Dasgupta, Gangopadhyay and Mukherjee, 2009: 31. India. 

grandcolasi Clastrier and Wirth, 1995: 112. French Guiana. 

gressitti Tokunaga and Murachi, 1959: 152. Belau (USA). 

guyana Clastrier and Wirth, 1995: 124. French Guiana. 

inflatipalpis Clastrier and Delécolle, 1993: 153. New Caledonia (France). 

insignipalpis Macfie, 1949: 109. Mexico. 

insulae Clastrier and Delécolle, 1993: 151. New Caledonia (France). 

intonsa Chan and LeRoux, 1971b: 743. Singapore. 

iquitosensis Clastrier and Wirth, 1995: 139. Peru. 

ixodoides (Fiebrig-Gertz), 1928: 290 (Ceratopogon). Paraguay. 

kawensis Clastrier and Wirth, 1995: 120. French Guiana. 

lasionota (Kieffer), 1911c: 334 (Ceratopgon). Seychelles. 

lignaria Debenham, 1987d: 655. Australia (Queensland). 

longurius Liu and Yu, 1997: 19. China. 

luteisquamosa Wirth, 1972: 575. Brazil. 

mayeri Forattini and Lane, 1955: 4. New name for crudelis. 

crudelis (Mayer), 1937b: 233 (Phasmidohelea, preoccupied by Forcipomyia crudelis (Karsch), 1886). Costa Rica. 

49

menzeli Clastrier and Wirth, 1995: 106. Ecuador. 

microtoma (Kieffer), 1917b: 299. Brazil. 

minisquamosa Wirth, 1972: 570. Belize. 

moorei Clastrier and Wirth, 1995: 133. Brazil. 

neotropica Clastrier and Wirth, 1995: 126. Peru. 

nibleyi Tokunaga, 1962a: 178. Japan. 

nigrimaxillata Clastrier and Wirth, 1995: 118. Peru. 

notothena Liu and Yu, 1997: 22. China. 

novaeteutoniae Clastrier and Wirth, 1995: 107. Brazil. 

nudifascipes Saha, Dasgupta, Gangopadhyay and Mukherjee, 2009: 32. India. 

obesa Costa Lima, 1928: 85. Brazil. 

ocrearum Debenham, 1987d: 664. Papua New Guinea. 

ornata Tokunaga, 1940c: 61 (1940e: 166). Belau (USA). 

ovatitheca Saha, Dasgupta, Gangopadhyay and Mukherjee, 2009: 33. India. 

paenefuliginosa Saha, Dasgupta, Gangopadhyay and Mukherjee, 2009: 33. India. 

penultimata Wirth, 1972: 573. Brazil. 

perpusillus Liu and Yu, in Liu et al., 2001b: 209. China. 

peruviana Clastrier and Wirth, 1995: 123. Peru. 

phototropisma Liu and Yu, in Liu et al., 2001b: 211. China. 

pinjiensis Liu and Yu, in Liu et al., 2001b: 213. China. 

plaumanni Clastrier and Wirth, 1995: 127. Brazil. 

proximornata Debenham, 1987d: 657. Papua New Guinea. 

qiongzhongensis Liu, Yan and Liu, in Yu et al., 2005: 624. New name for flava. 

flava Liu, Yan and Liu, 1996: 20 (preoccupied by Forcipomyia flava (Williston)). China. 

raposoensis Clastrier and Wirth, 1995: 144. Colombia. 

rettenmeyerorum Clastrier and Wirth, 1995: 104. Panama. 

serrulifimbria Tokunaga, 1959: 283. Papua New Guinea. 

similis Liu and Yu, 1997: 20. China. 

similitheca Saha, Dasgupta, Gangopadhyay and Mukherjee, 2009: 34. India. 

squamosa Lutz, 1914: 87. Brazil. 

stelechos Debenham, 1987d: 649. Australia (Queensland). 

subspadicifascia Tokunaga and Murachi, 1959: 167. Micronesia. 

tegula Liu and Yu, 1997: 27. China. 

tenuifascicruris Saha, Dasgupta, Gangopadhyay and Mukherjee, 2009: 35. India. 

tettigonaris Wirth and Castner, 1990: 159. Peru. 

thomasi Clastrier and Wirth, 1995: 115. Brazil. 

tiwaka Clastrier and Delécolle, 1993: 144. New Caledonia (France). 

tuthilli Tokunaga, in Tokunaga and Murachi, 1959: 174. Marshall Islands. 

valleensis Clastrier and Wirth, 1995: 145. Colombia. 

vernoni Clastrier and Wirth, 1995: 138. Colombia. 

wagneri (Séguy), 1941: 85 (Phasmidohelea). Brazil. 

walschaertsi Gosseries, 1989: 3. New name for monilis. 

monilis Tokunaga, 1962a: 179 (preoccupied by Forcipomyia monilis (Goetghebuer), 1934c). Japan. 

weihaiensis Xue and Yu, 1998: 317. China. 

willisi Debenham, 1987d: 661. Australia (New South Wales). 

willistoni Wirth, 1971b: 242. Puerto Rico (USA). 

wuxiensis Liu and Yu, in Liu et al., 2001b: 222. China. 

Subgenus NICOTHOHELEA Liu, Tang and Hao 

NICOTHOHELEA Liu, Tang and Hao, 2002: 230 (as Nicothoehelea Yu and Liu, 2006: 633) (as subgenus of Forcipomyia). 

Type species: Forcipomyia delenificus Liu, Tang and Hao, by original designation. 

delenifica Liu, Tang and Hao, 2002: 231 (Yu and Liu, in Yu et al., 2005: 633). China. 

50

Subgenus OREINOHELEA Yu and Liu 

OREINOHELEA Yu and Liu, 2006: 635 (as subgenus of Forcipomyia). Type species: Forcipomyia lenis Liu and Yu, by 


lenis Liu and Yu, in Yu et al., 2005: 635. China. 

Subgenus PANHELEA Remm 

PANHELEA Remm, 1980: 117 (as subgenus of Forcipomyia). Type species: Forcipomyia pontica Remm (= Ceratopogon 

brevicubitus Goetghebuer), by original designation. 

aristolochiae (Rondani), 1860: 134 (Ceratopogon). Italy. 

brevicubita (Goetghebuer), 1920: 27 (Ceratopogon). Belgium. 

pontica Remm, in Remm and Zhogolev, 1968: 831. Ukraine. 

hissarica Remm, 1980: 117. Tajikistan. 

Subgenus PEDILOHELEA de Meillon and Wirth 

PEDILOHELEA de Meillon and Wirth, 1980: 9 (as subgenus of Forcipomyia). Type species: Forcipomyia clastrieri Dessart, 


aitkeni de Meillon and Wirth, 1980: 12. Brazil. 

archboldi de Meillon and Wirth, 1980: 13. Dominica. 

australis Clastrier and Delécolle, 1991: 212. New Caledonia (France). 

bilancea Liu and Yu, in Yu et al., 2005: 637. China. 

brinchangensis de Meillon and Wirth, 1980: 14. Malaysia. 

brincki de Meillon, 1959a: 328. South Africa. 

clastrieri Dessart, 1963: 183. New name for brevitarsata. 

brevitarsata (Clastrier), 1959b: 345 (Lepidohelea, preoccupied by Forcipomyia brevitarsata (Ingram and 

Macfie), 1924). Senegal. 

draconis de Meillon and Wirth, 1980: 17. South Africa. 

eremita Szadziewski, Gwizdalska-Kentzer and Gilka, 2011: 638. United Arab Emirates. 

eshowensis de Meillon, 1937b: 361. South Africa. 

forcipis de Meillon and Wirth, 1980: 18. Malaysia. 

hailaerensis Liu and Yu, in Yu et al., 2005: 639. China. 

haroldi de Meillon and Wirth, 1989b: 208. South Africa. 

proavia Debenham, 1987d: 642. Australia (Capital Territory). 

raposoi de Meillon and Wirth, 1980: 21. Colombia. 

spangleri de Meillon and Wirth, 1980: 21. Guatemala. 

spilmani de Meillon and Wirth, 1980: 22. Dominica. 

tasmani Macfie, 1932c: 27. New Zealand. 

Subgenus PHYTOHELEA Remm 

PHYTOHELEA Remm, 1971: 189 (as subgenus of Forcipomyia). Type species: Ceratopogon bromelicola Lutz, by original 

designation. 

alocasiae Tokunaga, 1961b: 120. Papua New Guinea. 

antiguensis Saunders, 1957: 700. Antigua. 

bacoti (Ingram and Macfie), 1923: 55 (Apelma). Sierra Leone. 

natalia (de Meillon), 1936: 158 (Lasiohelea). South Africa. 

belkini de Meillon and Wirth, 1979e: 198. New Zealand. 

51

evis (Johannsen), 1927c: 205 (Apelma). USA (Hawaii). 

bromelicola (Lutz), 1914: 84 (Ceratopogon). Brazil. 

caribbeana Saunders, 1957: 696. Trinidad and Tobago. 

comis Johannsen, 1932: 409. Indonesia. 

dominicana de Meillon and Wirth, 1979e: 195. Dominica. 

edwardsi (Saunders), 1925: 260 (Apelma). Brazil. 

eophytoheleana Szadziewski, 1988: 215. Baltic region. Eocene. 

fijiensis (Macfie), 1945: 1 (Apelma). Fiji. 

forfices Debenham, 1987a: 114. Australia (Queensland). 

grandis Chan and LeRoux, 1971b: 758. Singapore. 

hamaticauda Tokunaga, in Tokunaga and Murachi, 1959: 212. Belau (USA). 

jocosa Saunders, 1957: 701. Trinidad and Tobago. 

mahamutius Liu and Yu, in Liu et al., 2001b: 222. New name for jocosus. 

jocosus Yu and Maha, 1998: 205 (preoccupied by Forcipomyia jocosa Saunders, 1957). China. 

keilini (Saunders), 1925: 265 (Apelma). Brazil. 

magna (Saunders), 1925: 266 (Apelma). Brazil. 

marksae Tokunaga, 1961b: 117. Papua New Guinea. 

musae Clastrier and Delécolle, 1994: 51. French Guiana. 

nicopina Chan and LeRoux, 1971b: 754. Singapore. 

oligarthra Saunders, 1957: 698. Puerto Rico (USA). 

onustagalea Yu, 2000b: 163. New name for onustus. 

onustus Yu and Liu, 1999: 247 (preoccupied by Forcipomyia onusta Remm, 1980). China. 

sabroskyi Tokunaga, in Tokunaga and Murachi, 1959: 211. Belau (USA). 

tzaneenensis de Meillon and Wirth, 1979e: 184. South Africa. 

Subgenus PTEROBOSCA Macfie 

PTEROBOSCA Macfie, 1932a: 266. Unavailable name; proposed after 1930 without type species designation. 

PTEROBOSCA Macfie, 1940f: 16. Type species: Ceratopogon aeschnosuga de Meijere, by original designation. 

adhesipes (Macfie), 1932a: 270 (Pterobosca). Indonesia. 

aerobates (Macfie), 1936a: 64 (Pterobosca). India. 

aeschnosuga (de Meijere), 1923: 137 (Ceratopogon). Indonesia. 

ariel (Macfie), 1932a: 275 (Pterobosca). Indonesia. 

asahinai Tokunaga, 1962a: 188. Japan. 

chrysopipennis Wirth, 1966: 29. New name for chrysopae. 

chrysopae Tokunaga, in Tokunaga and Murachi, 1959: 235 (preoccupied by Forcipomyia chrysopae Mayer, 1934b). 

Belau (USA). 

crinume (Tokunaga), 1932: 1 (Dasyhelea). Japan. 

esakii (Tokunaga), 1940a: 210 (Pterobosca). Mariana Islands (USA). 

esakia Tokunaga, in Tokunaga and Murachi, 1959: 234. New name for esakii. 

farri Wirth, 1966: 29. Jamaica. 

feminae (Tokunaga), 1940c: 100 (1940e: 168) (Pterobosca). Belau (USA). 

fidens (Macfie), 1936b: 228 (Pterobosca). China. 

fusicornis (Coquillett), 1905: 63 (Ceratopogon). USA (Florida). 

fur (Johnson), 1913: 444 (Ceratopogon). Bermuda (Great Britain). 

macfiei (Costa Lima), 1937b: 616 (Pterobosca). Brazil. 

floridana (Johannsen), 1950: 143 (Pterobosca). USA (Florida). 

hutsoni Wirth and Ratanaworabhan, 1976: 242. Aldabra (Seychelles). 

incubans (Macfie), 1937c: 111 (Pterobosca). Belize. 

lairdi (Wirth), 1956a: 363 (Pterobosca). Solomon Islands. 

latipes (Macfie), 1936b: 227 (Pterobosca). China. 

mollipes (Macfie), 1932a: 272 (Pterobosca). Liberia. 

neodebenhamae Szadziewski and Borkent, 2003: 256. New name for debenhamae. 

52

debenhamae Cranston, in Orr and Cranston, 1997: 1850 (preoccupied by Forcipomyia debenhamae Clastrier and 

Delécolle, 1991). Brunei. 

odonatiphila (Macfie), 1932a: 274 (Pterobosca). Indonesia. 

ogatai Tokunaga, 1961c: 3. New Caledonia (France). 

paludis (Macfie), 1936a: 63 (Pterobosca). Great Britain. 

pinheyi Clastrier and Legrand, 1984: 173. Mauritius. 

tokunagai (Oka and Asahina), 1948: 107 (Pterobosca). Japan. 

Subgenus RHINOHELEA de Meillon and Wirth 

RHINOHELEA de Meillon and Wirth, 1979d: 881 (as subgenus of Forcipomyia). Type species: Forcipomyia briani de 

Meillon and Wirth, by original designation. 

briani de Meillon and Wirth, 1979d: 882. South Africa. 

khoisana de Meillon and Wirth, 1979d: 885. South Africa. 

Subgenus RHYNCHOFORCIPOMYIA Wirth and Dow 

RHYNCHOFORCIPOMYIA Wirth and Dow, 1972: 863 (as subgenus of Forcipomyia). Type species: Forcipomyia 

messersmithi Wirth and Dow, by original designation. 

brachyrhyncha Wirth and Dow, 1972: 867. Colombia. 

cylindrica Wirth and Dow, 1972: 865. El Salvador. 

dorsalis Wirth and Dow, 1972: 870. Mexico. 

flavisimplexa Saha, Dasgupta, Gangopadhyay and Mukherjee, 2009: 35. India. 

guamai Wirth and Dow, 1972: 869. Brazil. 

messersmithi Wirth and Dow, 1972: 864. El Salvador. 

puracensis Wirth and Dow, 1972: 867. Colombia. 

zeteki Wirth and Dow, 1972: 870. Panama. 

Subgenus SALIOHELEA Wirth and Ratanaworabhan 

SALIOHELEA Wirth and Ratanaworabhan, 1978: 494 (as subgenus of Forcipomyia). Type species: Forcipomyia leei Wirth 

and Ratanaworabhan, by original designation. 

boudinoti Clastrier and Delécolle, 1991: 216. New Caledonia (France). 

brevicosta (Clastrier), 1960c: 520 (Lasiohelea). Congo. 

calleida Yu, in Yu et al., 2005: 648. New name for claris (as clarus). 

claris Yu, 2001: 159 (preoccupied by Forcipomyia clara Chan and LeRoux, 1971b). China. 

cicuta Yu, in Yu and Song, 2008: 795. New name for ciliata. 

ciliata Liu and Yu, in Yu et al., 2005: 646 (preoccupied by Forcipomyia ciliata (Winnertz), 1852). China. 

deminuta Tokunaga and Murachi, 1959: 219. Micronesia. 

digita Spinelli, Marino and Borkent, 2012: 42. Costa Rica. 

infrensi Liu and Yu, in Yu et al., 2005: 649. China. 

isolata Liu and Yu, in Yu et al., 2005: 650. China. 

leei Wirth and Ratanaworabhan, 1978: 498. Colombia. 

perrara Liu and Yu, in Yu et al., 2005: 651. China. 

rima Yu and Xue, in Yu et al., 2005: 652. China. 

stami Wirth and Ratanaworabhan, 1978: 502. Democratic Republic of the Congo. 

Subgenus SCHINEROMYIA Debenham 

SCHINEROMYIA Debenham, 1987a: 67 (as subgenus of Forcipomyia). Type species: Forcipomyia gandangara Debenham, 


53

gandangara Debenham, 1987a: 67. Australia (New South Wales). 

Subgenus SCHIZOFORCIPOMYIA Chan and LeRoux 

SCHIZOFORCIPOMYIA Chan and LeRoux, 1971a: 271 (as subgenus of Forcipomyia). Type species: Forcipomyia 

petersoni Chan and LeRoux (= Forcipomyia borbonica Clastrier), by original designation. 

anna de Meillon, 1959a: 331. South Africa. 

borbonica Clastrier, 1959c: 436. Réunion (France). 

penniornata Tokunaga and Murachi, 1959: 163. Guam (USA). 

fuscimaculata Hardy, 1960: 170. USA (Hawaii). 

stabilis Sen and Das Gupta, 1968: 95. India. 

petersoni Chan and LeRoux, 1971a: 272. Singapore. 

chazeaui Clastrier and Delécolle, 1991: 222. New Caledonia (France). 

cinctipes (Coquillett), 1905: 64 (Ceratopogon). USA (Florida). 

clavula Debenham, 1987d: 639. Australia (New South Wales). 

harpa Spinelli and Borkent, 2004b: 3 (male, female; Costa Rica). Distr.- Costa Rica. 

lecordeurorum de Meillon, Meiswinkel and Wirth, 1982: 132. South Africa. 

lydiae Clastrier and Delécolle, 1991: 219. New Caledonia (France). 

monoceros Debenham, 1987d: 634. Australia (New South Wales). 

rudebecki de Meillon, 1959a: 330. South Africa. 

tinia Krivosheina, 1968: 583. Russia. 

warreni de Meillon and Wirth, 1981b: 528. South Africa. 

yirrkala Debenham, 1987d: 636. Australia (Northern Territory). 

Subgenus SYNTHYRIDOMYIA Saunders 

SYNTHYRIDOMYIA Saunders, 1957: 688 (as subgenus of Forcipomyia). Type species: Lasiohelea acidicola Tokunaga, by 


acidicola (Tokunaga), 1937b: 455 (Lasiohelea). Japan. 

minuta Goetghebuer, 1947: 228 (preoccupied by Forcipomyia minuta (Tokunaga), 1940d). Belgium. 

colemani Wirth, 1952a: 146. USA (California). 

ansericolli Debenham, 1987a: 59. Australia (Northern Territory). 

bucera Debenham, 1987a: 55. Australia (Queensland). 

caestuum Debenham, 1987a: 53. Australia (Queensland). 

calchaqui Spinelli and Marino, 1997: 188. Argentina. 

contigoa Liu and Yu, in Yu et al., 2005: 655. China. 

corsoni Macfie, 1926: 355. Tanzania. 

exigua Ingram and Macfie, 1924b: 587. Ghana. 

flexoductitheca Saha, Dasgupta, Gangopadhyay and Mukherjee, 2009: 37. India. 

floridensis Dow and Wirth, 1972: 195. USA (Florida). 

hengqinensis Yu, Sun and Li, in Sun et al., 2009: 105. China. 

hylecoeta Yu and Xu, 2000: 449. China. 

knockensis Goetghebuer, 1938: 375. Belgium. 

bequaerti Goetghebuer, 1942: 1. Belgium. 

abludens Remm, in Remm and Zhogolev, 1968: 830. Ukraine. 

margaritae Szadziewski, 1983a: 377. Algeria. 

mendica Liu and Yu, in Yu et al., 2005: 659. China. 

murina (Winnertz), 1852: 26 (Ceratopogon). Europe. 

abdominalis (Santos Abreu), 1918: 265 (Helea, as variety of murina). Canary Islands (Spain). 

aurosparsum (Kieffer), 1919a: 65 (Apelma). Hungary. 

sulfurea Kieffer, 1923a: 664. Algeria. 

54

hirtipalpis Kieffer, 1924b: 392. France. 

sate Kieffer, 1925e: 245. Egypt. 

longitarsis Tokunaga, 1940c: 92 (preoccupied by Forcipomyia longitarsis (Malloch), 1915b). Taiwan. 

moascari Macfie, 1943a: 147. Egypt. 

attonsa Goetghebuer, 1950: 1. Belgium. 

tokunagai Wirth, 1973: 356 (preoccupied by Forcipomyia tokunagai (Oka and Asahina), 1948. New name for 

longitarsis. 

submurina Remm, 1980: 115 (as sibmurina, as subspecies of murina). Russia. 

nanshengwei Yu, Liang and Su, in Yu et al., 2007: 487. China. 

operimenti Debenham, 1987a: 58. Australia (Queensland). 

oryx Debenham, 1987a: 54. Australia (Northern Territory). 

sanctaeclarae Wirth, 1952b: 90. Chile. 

soibelzoni Marino and Spinelli, 2001c: 14. Argentina. 

tenuiforceps Macfie, 1939c: 167. Brazil. 

tertiaricola Szadziewski and Grogan, 1998b: 281. Dominican Republic. Miocene. 

tympanista Debenham, 1987a: 56. Australia (Queensland). 

unituberculata Tokunaga, in Tokunaga and Murachi, 1959: 197. Belau (USA). 

xiangshanensis Yu, Liu and Chen, in Liu et al., 2009: 522. China. 

yuani Yu, Su and Chen, in Yu et al., 2007: 487. China. 

Subgenus THYRIDOMYIA Saunders 

THYRIDOMYIA Saunders, 1925: 268. Type species: Thyridomyia palustris Saunders (= Ceratopogon monilicornis 

Coquillett), by original designation. 

angelicae Tokunaga, 1940c: 97. Japan. 

biskraensis Kieffer, 1923a: 665 (as variety of seneveti). Algeria. 

sergenti Clastrier, 1956: 502. Algeria. 

imeretica Remm, 1967: 5. Georgia. 

blascoi Delécolle and Rieb, 1993: 112. Spain. 

carolinea Tokunaga, in Tokunaga and Murachi, 1959: 214. Belau (USA). 

cineornata Saha, Dasgupta, Gangopadhyay and Mukherjee, 2009: 38. India. 

colombiae Wirth, 1970: 435. Colombia. 

concinna Liu and Yu, in Yu et al., 2005: 663. China. 

falcinella (Kieffer), 1911c: 338 (Ceratopogon). Seychelles. 

fererugosa Saha, Dasgupta, Gangopadhyay and Mukherjee, 2009: 38. India. 

frutetorum (Winnertz), 1852: 29 (Ceratopogon). Germany. 

seneveti Kieffer, 1922g: 500. Algeria. 

aethiopiae Ingram and Macfie, 1924b: 582. Ghana. 

dasyptera Goetghebuer, 1934b: 214 (Dasyhelea). Israel. 

japonica (Tokunaga), 1937a: 267 (Lasiohelea). Japan. 

aspinosa Saunders, 1957: 692. Canada (Saskatchewan). 

frutetosuccinea Szadziewski and Grogan, 1998b: 280. Dominican Republic. Miocene. 

gokwe de Meillon and Wirth, 1989b: 209. Zimbabwe. 

gossympina Chan and LeRoux, 1970: 286. Singapore. 

hamata Tokunaga, in Tokunaga and Murachi, 1959: 207. Belau (USA). 

hoplia Liu and Yu, in Yu et al., 2005: 666. China. 

inconspicuosa (Ingram and Macfie), 1924a: 391 (Lasiohelea). Ghana. 

ishigakii Tokunaga, 1962a: 182. Japan. 

jamaicensis Wirth, 1970: 436. Jamaica. 

jipajapae Wirth, 1970: 437. Panama. 

johannseni Thomsen, 1935: 286. USA (New York). 

kii (Tokunaga), 1940d: 107 (Lasiohelea). Japan. 

kitasirakawae Tokunaga, 1940c: 94. Japan, Taiwan. 

55

litoraurea (Ingram and Macfie), 1924a: 389 (Lasiohelea). Ghana. 

minutissima Remm, 1961a: 192. Estonia. 

masaakii Debenham, 1989: 235. New name for esakii. 

esakii (Tokunaga), 1940e: 169 (1940d: 101) (Lasiohelea, preoccupied by Forcipomyia esakii (Tokunaga), 1940a). 

Belau (USA). 

mercuratus Liu, Yan and Liu, 1996: 21 (as mercuratas). China. 

minima (Tokunaga), 1940d: 104 (Lasiohelea). Japan. 

minuta (Tokunaga), 1940d: 106 (Lasiohelea). Japan. 

monilicornis (Coquillett), 1905: 63 (Ceratopogon). Canada (British Columbia). 

hirta (Lundström), 1910: 34 (Ceratopogon). Finland. 

microcera (Kieffer), 1919a: 54 (Dasyhelea). Slovak Republic. 

palustris (Saunders), 1925: 269 (Thyridomyia, preoccupied by Forcipomyia palustris (Meigen), 1804). Great Britain. 

curticornis Goetghebuer, 1933c: 354. Belgium. 

curticornis Goetghebuer, 1949: 1 (preoccupied by Forcipomyia curticornis Goetghebuer, 1933c). Belgium. 

kabashae de Meillon, 1959a: 333. Democratic Republic of the Congo. 

nana (Macfie), 1939c: 171 (Lasiohelea). Brazil. 

neomonticola Borkent, in Borkent and Wirth, 1997:48. New name for monticola. 

monticola (Tokunaga), 1937a: 270 (Lasiohelea, preoccupied by Forcipomyia monticola Kieffer, 1913d). Japan. 

nodosa Saunders, 1959: 43. Costa Rica. 

nudocula (Tokunaga), 1937a: 269 (Lasiohelea). Japan. 

oxyria Yu, Ke and Li, in Sun et al., 2009: 105. China. [preoccupied and needs new name if Lasiohelea oxyria Yu and Liu, 

considered to be a species of Forcipomyia] 

riojana Spinelli and Marino, 1997: 188. Argentina. 

rugosa Chan and LeRoux, 1970: 280. Canada (Quebec). 

murinoides Remm, 1971: 188. Estonia. 

sinuosa Dow and Wirth, 1972: 189. Mexico. 

skiaphila (Clastrier), 1960c: 522 (Lasiohelea). Congo. 

tenuichela Dow and Wirth, 1972: 191. USA (California). 

tugelensis de Meillon, 1959a: 333. South Africa. 

univesicula Macfie, 1939c: 170. Brazil. 

ursuli Remm, 1972: 66. Russia. 

vertexcava Chan and LeRoux, 1970: 282. Singapore. 

watshami de Meillon and Wirth, 1989b: 210. Zimbabwe. 

xiphoidea Liu and Yu, in Yu et al., 2005: 675. China. 

yaana Liu and Yu, in Yu et al., 2005: 676. China. 

Subgenus TRICHOHELEA Goetghebuer 

APELMA Kieffer, 1919a: 64 (preoccupied by Billberg, 1820). Type species: Apelma auronitens Kieffer (= Trichohelea 

tonnoiri Goetghebuer), designation by Macfie, 1940f: 16. 

TRICHOHELEA Goetghebuer, 1920: 39. Type species: Trichohelea tonnoiri Goetghebuer, by original designation. 

NEOFORCIPOMYIA Tokunaga, in Tokunaga and Murachi, 1959: 200 (as subgenus of Forcipomyia). Type species: 

Ceratopogon pectinunguis de Meijere, by original designation. 

aeronautica Macfie, 1935b: 265. Guyana. 

aliena Debenham, 1987a: 83. Australia (New South Wales). 

araneivora Clastrier and Legrand, 1991: 155. Guinea. 

austrina Macfie, 1932c: 29. New Zealand. 

basiflava Tokunaga, in Tokunaga and Murachi, 1959: 202. Micronesia. 

baueri Wirth, 1956a: 361. USA (Arizona). 

bifidicola Szadziewski, 1993: 649. Germany. Eocene. 

blanda Yu and Liu, in Yu et al., 2005: 6678. China. 

caliginosa (Ingram and Macfie), 1924a: 386 (Lasiohelea). Ghana. 

castania Liu and Yu, in Yu et al., 2005: 680. China. 

56

chaetoptera Remm, 1962b: 188. Estonia. 

chirurga Debenham, 1987a: 87. Australia (New South Wales). 

cliens Debenham, 1987a: 76. Papua New Guinea. 

collinsi Lane, 1977: 308. Ethiopia. 

consortis de Meillon and Wirth, 1989b: 212. South Africa. 

crinita Saunders, 1964: 474. Canada (Saskatchewan). 

danaisi (Floch and Abonnenc), 1949: 2 (1950a: 72) (Lasiohelea). Venezuela. 

debenhamae Clastrier and Delécolle, 1991: 228. New name for arcis. 

arcis Debenham, 1987a: 91 (preoccupied by Forcipomyia arcis de Meillon and Downes, 1986). Australia (Northern 

Territory). 

eotrichoheleana Szadziewski, 1988: 213. Baltic region. Eocene. 

eques (Johannsen), 1908: 266 (Ceratopogon). USA (New York). 

chrysopae Mayer, 1934b: 259. Germany. 

chrysopivora (Tokunaga), 1939b: 371 (Lasiohelea). Japan. 

equitans (Edwards), 1933c: 251 (Lasiohelea). Malaysia. 

ferrea Debenham, 1987a: 76. Australia (Queensland). 

geometrica (Clastrier), 1959c: 437 (Lasiohelea). Réunion (France). 

goniognatha Wirth and Messersmith, 1971: 20. Mexico. 

imparidentes Debenham, 1987a: 77. Fiji. 

insignicornis (Macfie), 1947a: 29 (Lasiohelea). Uganda. 

insignis (Skuse), 1889: 298 (Ceratopogon). Australia (New South Wales). 

sydneyensis (Skuse), 1889: 302 (Ceratopogon). Australia (New South Wales). 

intrepida Macfie, 1936b: 228. Peru. 

kuscheli Sublette and Wirth, 1980: 330. New Zealand. 

leptognatha Wirth and Messersmith, 1971: 21. USA (Kansas). 

limnetis Ingram and Macfie, 1931a: 169. Argentina. 

shannoni (Ingram and Macfie), 1931a: 171 (Lasiohelea). Argentina. 

lunata Debenham, 1987a: 90. Australia (New South Wales). 

lushanensis Yu and Liu, in Yu et al., 2005: 681. China. 

macheti Clastrier and Legrand, 1990: 168. French Guiana. 

mcateei Wirth, 1956a: 359. USA (Maryland). 

saundersi Chan, in Chan and LeRoux, 1965: 87. Canada (Quebec). 

mexicana Wirth, 1956a: 361. Mexico. 

monoplectron Lane, 1977: 306. Tanzania. 

okadai (Tokunaga), 1939b: 370 (Lasiohelea). Russia. 

opilionivora (Lane), 1947c: 159 (Lasiohelea). Brazil. 

pacifica (Macfie), 1933a: 75 (Lasiohelea). French Polynesia (France). 

papuensis Lane and Cotman, 1986: 617. Papua New Guinea. 

pectinunguis (de Meijere), 1923: 138 (Ceratopogon). Indonesia. 

pennambula (Macfie), 1932a: 279 (Lasiohelea). Indonesia. 

quateriungula Tokunaga, 1959: 293. Indonesia. 

roubaudi Clastrier and Delécolle, 1997: 379. French Guiana. 

samoensis (Edwards), 1928: 51 (Lasiohelea). Western Samoa. 

sayhuequei Marino and Spinelli, 2004b: 2257. Argentina. 

scorpio Debenham, 1987a: 88. Australia (New South Wales). 

sector Debenham, 1987: 83. Australia (New South Wales). 

simulataris Liu and Yu, in Yu et al., 2005: 684. China. 

subauronitens Tokunaga, 1940c: 96. Japan. 

succinicola Szadziewski, 1993: 650. Germany. Eocene. 

tehuelche Marino and Spinelli, 2004b: 2260. Argentina. 

tipulivora Macfie, 1936b: 229. Indonesia. 

tonnoiri (Goetghebuer), 1920: 39 (Trichohelea). Belgium. 

auronitens (Kieffer), 1919a: 65 (Apelma, preoccupied by Forcipomyia auronitens (Kieffer), 1910). Slovak Republic. 

papilionivora Edwards, 1923b: 23. Great Britain. 

57

umbellicola Remm, 1971: 189. Russia. 

trinidadensis Saunders, 1964: 476. Trinidad and Tobago. 

tripeda Liu and Yu, in Yu et al., 2005: 685. China. 

tsutsumii Tokunaga, 1960b: 76. Japan. 

tumula Debenham, 1987a: 92. Australia (New South Wales). 

veroensis Wirth and Messersmith, 1971: 24. USA (Florida). 

wudangensis Yu and Liu, in Yu et al., 2005: 687. China. 

Subgenus TRITHICOMYIA Yu and Song 

TRITHICOMYIA Yu and Song, 2008: 793 (as subgenus of Forcipomyia). Type species: Trithicomyia huainanensis Yu and 

Song, by original designation. 

huainanensis Yu and Song, 2008: 793. China. 

Subgenus TYPHONOMYIA Debenham 

TYPHONOMYIA Debenham, 1987a: 93 (as subgenus of Forcipomyia). Type species: Forcipomyia anachoreta Debenham, 


anachoreta Debenham, 1987a: 94. Australia (New South Wales). 

Subgenus WARMKEA Saunders 

WARMKEA Saunders, 1957: 671 (as subgenus of Forcipomyia). Type species: Forcipomyia bicolor Saunders (= 

Forcipomyia lesliei Wirth), by original designation. 

aeria Saunders, 1957: 675. Puerto Rico (USA). 

albiacies Debenham, 1987b: 170. Australia (Queensland). 

brunnea de Meillon and Downes, 1986: 151. South Africa. 

distinctiscuta Tokunaga, 1962a: 173. Japan. 

fishi Wirth and Soria, 1980: 145. USA (Florida). 

galindoi Wirth and Soria, 1980: 146. Panama. 

kaufmannae Wirth and Derron, 1976: 230. Sao Tomé and Principe. 

lesliei Wirth, 1974: 12. New name for bicolor. 

bicolor Saunders, 1957: 672 (preoccupied by Forcipomyia bicolor Lutz, 1914). Puerto Rico (USA). 

louriei (Macfie), 1935a: 49 (Lasiohelea). Brazil. 

narthekophora Macfie, 1939c: 166. Brazil. 

malayae Saunders, 1957: 678. Malaysia. 

samyda Yu and Liu, in Yu et al., 2005: 690. China. 

spinosa Saunders, 1957: 676. Puerto Rico (USA). 

terrestris Saunders, 1964: 479. Trinidad and Tobago. 

tuberculata Saunders, 1957: 677. Trinidad and Tobago. 

Species of FORCIPOMYIA unplaced to subgenus 

longitarsis (Malloch), 1915b: 314 (Euforcipomyia). USA (Illinois). 

nabilis Yu, in Yu and Song, 2008: 795. New name for notata. 

notata Yu and Liu, in Yu et al., 2005: 691 (preoccupied by Forcipomyia notata Macfie, 1939a). China. 

reburra Borkent, in Borkent and Wirth, 1997:50. New name for villosa. 

villosa (Macfie), 1934c: 183 (Lasiohelea, preoccupied by Forcipomyia villosa (Zetterstedt), 1850). Malaysia. 

SUBFAMILY DASYHELEINAE LENZ, 1934: 96 

58

Genus DASYHELEA Kieffer 

DASYHELEA Kieffer, 1911d: 5. Type species: Dasyhelea halophila Kieffer, by monotypy. 

PROKEMPIA Kieffer, 1913d: 163, 179 (as subgenus of Dasyhelea). Type species: Dasyhelea ornaticornis Kieffer, 

designation by Wirth, 1973: 358. 

PSEUDOCULICOIDES Malloch, 1915a: 309. Type species: Ceratopogon mutabilis Coquillett, by original designation. 

TETRAHELEA Kieffer, 1925a: 423. Type species: Culicoides insignicornis Kieffer, by original designation. 

DICRYPTOSCENA Enderlein, 1936: 51. Type species: Dasyhelea inclusa Kieffer, by original designation. 

SEBESSIA Remm, 1979b: 55 (as subgenus of Dasyhelea). Type species: Dasyhelea flavopyga Zilahi-Sebess, by original 

designation. 

BORKENTIMYIA Yu, Liu, Liu, Liu, Hao, Yan and Zhao, 2006: 321 (as subgenus of Dasyhelea). Type species: Dasyhelea 

forsteri Grogan and Wirth, by original designation. 

abdita Yu, in Yu et al., 2005: 122. China. 

abdominalis (Santos Abreu), 1918: 303 (Culicoides). Canary Islands (Spain). 

abhazica Remm, 1967: 21. Georgia. 


abreui (Kieffer), 1921e: 7 (Culicoides). New name for flaviventris. 

flaviventris (Santos Abreu), 1918: 292 (Culicoides, as variety of versicolor, preoccupied by Dasyhelea flaviventris 

(Goetghebuer), 1910). Canary Islands (Spain). 

abronica Yu, 2001: 160. China. 

abyenensis Boorman and Harten, 2002: 440. Yemen. 

actita Yu, in Yu et al., 2005: 125. China. 

acuminata Kieffer, 1919a: 60. Hungary. 

polita Edwards, 1921: 124. Great Britain. 

verticillata Kieffer, 1925d: 63. France, Hungary. 

littoralis Goetghebuer, 1934c: 289. Belgium. 

adami Vattier, 1964: 1160. Congo. 

adjaniae Gosseries, 1989: 3. New name for acuminata. 

acuminata Goetghebuer, 1935d: 165 (preoccupied by Dasyhelea acuminata Kieffer, 1919a). Democratic Republic of 

Congo. 

goetghebueri Wirth, 1980: 159, in Wirth et al., 1980 (preoccupied by Dasyhelea goetghebueri Kieffer, 1919a). New 

name for acuminata. 

adjecta Mazumdar and Chaudhuri, 2009a: 192. India. 

aegealitis Spinelli and Wirth, 1984d: 589. Jamaica. 

aeratipennis (Skuse), 1889: 303 (Ceratopogon). Australia (New South Wales). 

affinis (Johannsen), 1932: 422 (Tetraphora, as variety of tersa). Indonesia. 

afghanica Navai, 1994: 359. Afghanistan. 

africana Clastrier, 1959b: 379. Senegal. 

agrila Yu, in Yu et al., 2005: 309. China. 

aithalodes Remm, 1971: 200. Russia. 

albidipes (Santos Abreu), 1918: 291 (Culicoides, as variety of versicolor). Canary Islands (Spain). 

intermedia (Santos Abreu), 1918: 310 (Culicoides). Canary Islands (Spain). 

guanchense Clastrier, 1966: 702. Canary Islands (Spain). 

albiscapula Clastrier, 1959b: 377. Senegal. 

albohalterata (Santos Abreu), 1918: 299 (Culicoides, as variety of sericatus). Canary Islands (Spain). 

albopruinosa Goetghebuer, 1933e: 143. Democratic Republic of the Congo. 

alboscutellata Clastrier, 1959c: 424. Réunion (France). 

alboverrucosa Remm, 1967: 17. Azerbaijan. 

algarum Kieffer, 1924a: 21. Germany. 

alonensis (Strobl), 1906: 396 (Ceratopogon). Spain. 

alticola Liu and Zhang, in Liu, Zhang and Luo, 2002: 257. China. 

alula Yu, in Yu et al., 2005: 259. China. 

ambita Yu, in Yu et al., 2005: 177. China. 

59

ampelis Yu, in Yu et al., 2005: 126. China. 

ampullariae Macfie, 1934c: 279. Malaysia. 

ancora (Coquillett), 1902a: 87 (Ceratopogon). USA (Florida). 

andensis Ingram and Macfie, 1931a: 180. Argentina. 

antilleana Szadziewski and Grogan, 1998b: 257. Dominican Republic. Miocene. 

antiqua Palmer, 1957: 266 (as subspecies of australis). USA (California). Miocene. 

apiculata Yu, Chen and Yan, in Yu et al., 2005: 688. China. 

arciforceps Tokunaga, 1940d: 129. Japan. 

clavifulva Tokunaga, 1940d: 135. Japan. 

arenivaga Macfie, 1943a: 151 (as variety of inconspicuosa). Egypt. 

arenosa Kieffer, 1925e: 255. Algeria. 

argentinensis Kieffer, 1925c: 92. Argentina. 

assimilis (Johannsen), 1932: 425 (Tetraphora). Indonesia. 

atlantis Wirth and Williams, 1957: 11. Bermuda (Great Britain). 

atra de Meillon and Wirth, 1989b: 213. Zimbabwe. 

atrata Wirth, 1952a: 164. USA (California). 

atrifemorata Clastrier, 1961b: 264. Senegal. 

atrolaevis Goetghebuer, 1933e: 144. Democratic Republic of the Congo. 

atronotata Macfie, 1938: 158. Solomon Islands. 

atrostriata Goetghebuer, 1933e: 145. Democratic Republic of the Congo. 

aucklandensis Sublette and Wirth, 1980: 331. New Zealand. 

auli Remm, 1962a: 123. Estonia. 

aurantiaca (Kieffer), 1911c: 342 (Culicoides). Seychelles. 

mahensis (Kieffer), 1911c: 342 (Culicoides, as variety of aurantiaca). Seychelles. 

aurensis Remm, 1981: 32. New name for biskraensis. 

biskraensis Goetghebuer, 1939: 59 (preoccupied by Dasyhelea biskraensis Kieffer, 1923a). Algeria. 

aurosa Yu, in Yu et al., 2005: 129. China. 

australis Wirth, 1952b: 92. Chile. 

awadi Boorman and Harten, 2002: 440. Yemen. 

azteca Huerta and Grogan, 2006: 893. Mexico. 

bactriana Remm, 1980: 104. Uzbekistan. 

baculata Remm, 1967: 15. Georgia. 

bahamensis (Johnson), 1908: 71 (Ceratopogon). Bahamas. 

bajensis Wirth, 1978: 192. Mexico. 

baltica Remm, 1966: 58. Estonia. 

bambusaoris Yu, in Yu et al., 2005: 130. China. 

bamianica Navai, 1994: 363. Afghanistan. 

barbaros Yu, in Yu et al., 2005: 277. China. 

batouensis Yu, in Yu et al., 2005: 275. China. 

begueti Kieffer, 1922g: 508. Algeria. 

astyla Kieffer, 1922g: 510. Algeria. 

hirtipes Kieffer, 1922g: 510 (as variety of begueti, preoccupied by Dasyhelea hirtipes (Kieffer), 1917b). Algeria. 

bensoni Edwards, 1933a: 91. Great Britain. 

vernalis Remm, 1979b: 56. Estonia. 

bermudae Wirth and Williams, 1957: 11. Bermuda (Great Britain). 

biannulata Clastrier, Rioux and Descous, 1961: 68. Chad. 

bicolorea Goetghebuer, 1934d: 191. Democratic Republic of the Congo. 

bicornis Remm, 1993: 195. Russia. 

bicrenata Kieffer, 1923a: 668. Algeria. 

bifida Zilahi-Sebess, 1936b: 44 (as variety of fasciigera). Hungary. 

excellentis Borkent, 1997a: 91. USA (Hawaii). 

bifurcata Kieffer, 1923a: 669. Algeria. 

bilineata Goetghebuer, 1920: 45. Belgium. 

insignipalpis Kieffer, 1925d: 62 (as variety of versicolor). Europe. 

60

saxicola (Edwards), 1929a: 426 (Tetraphora). Great Britain. 

geleiana Zilahi-Sebess, 1930a: 11 (1931: 321). Hungary. 

montana Zilahi-Sebess, 1940: 48 (as variety of dufouri). Hungary. 

lithotelmatica Strenzke, in Thienemann, 1950: 178. Austria. 

tecticola Remmert, 1953: 334. Germany. 

dieuzeidei (Vaillant), 1957: 265 (Culicoides). France. 

karelica Glukhova and Brodskaya, 1997: 443. Russia. 

bilobata Kieffer, 1915b: 65. France. 

luteiventris Goetghebuer, 1934a: 90. Austria. 

spiralis Remm, 1966: 60. Lithuania. 

bipunctata Wirth and Messersmith, 1977: 304. Seychelles. 

biseriata Wirth and Beaver, 1979: 49. Malaysia. 

biskraensis Kieffer, 1923a: 672 (as variety of begueti). Algeria. 

bistriata Kieffer, 1915a: 472. Germany. 

biunguis Kieffer, 1925a: 409. Russia. 

bolei Ingram and Macfie, 1923: 54. Ghana. 

boothi Ingram and Macfie, 1921: 331. Nigeria. 

borbonica Clastrier, 1959c: 419. Réunion (France). 

borgmeieri Wirth and Waugh, 1976: 225. Trinidad and Tobago. 

borkenti Yu, in Yu et al., 2005: 321. China. 

brachibaios Yu and Liu, in Yu et al., 2005: 226. China. 

brevicornis Waugh and Wirth, 1976: 245. USA (New York). 

brevicosta Waugh and Wirth, 1976: 238. USA (Virginia). 

brevicula Yu, in Yu et al., 2005: 133. China. 

breviforceps Tokunaga, 1962a: 198. Japan. 

brevimana (Kieffer), 1910: 187 (Culicoides). India. 


browneae Pierce, 1966: 87. USA (California). Miocene. 

bullocki Tokunaga, 1958: 75. South Korea. 

cacaoi Wirth and Waugh, 1976: 231. Trinidad and Tobago. 

cactorum Wirth and Hubert, 1960a: 642. Mexico. 

caelata Yu, in Yu et al., 2005: 137. China. 

caerulea Yu and Hao, in Yu et al., 2005: 138. China. 

caesia Remm, 1993: 193. Russia. 

lugensis Brodskaya, 1995: 9. Russia. 

calvescens Macfie, 1938: 157. USA (Hawaii). 

calycata Remm, 1972: 74. Russia. 

canariensis (Santos Abreu), 1918: 293 (Culicoides, as variety of versicolor). Canary Islands (Spain). 

brunnea (Santos Abreu), 1918: 295 (Culicoides, as variety of versicolor). Canary Islands (Spain). 

fenestralis (Santos Abreu), 1918: 293 (Culicoides, as variety of versicolor). Canary Islands (Spain). 

ornatigaster (Santos Abreu), 1918: 294 (Culicoides, as variety of versicolor). Canary Islands (Spain). 

hiemalis (Santos Abreu), 1918: 314 (Culicoides). Canary Islands (Spain). 

albidigaster (Santos Abreu), 1918: 316 (Culicoides, as variety of hiemalis). Canary Islands (Spain). 

inmaculata (Santos Abreu), 1918: 315 (Culicoides, as variety of hiemalis). Canary Islands (Spain). 

quinquetaeniata Clastrier, 1966: 699. Canary Islands (Spain). 

caribbeana Spinelli and Wirth, 1984d: 596. Jamaica. 

carolinensis Tokunaga, 1941a: 112. Micronesia. 

cellulana de Meillon and Wirth, 1983a: 353. South Africa. 

centridorsalis Tokunaga, 1963c: 39. Japan. 

chambiensis Goetghebuer, 1935d: 166. Democratic Republic of the Congo. 

chani Wirth and Linley, 1990: 274. USA (Florida). 

chimaira Yu and Wang, in Yu et al., 2005: 278. China. 

cincta (Coquillett), 1901a: 605 (Ceratopogon). USA (Florida). 

albopicta Ingram and Macfie, 1931a: 187. Argentina. 

61

penthesileae Macfie, 1935a: 55. Brazil. 

circellata Yu, in Yu et al., 2005: 180. China. 

cirrata Yu, in Yu et al., 2005: 181. China. 

claviculifera Tokunaga, 1962a: 202. Japan. 

coarctata Kieffer, 1914a: 231. Switzerland. 

cogani Wirth, 1990: 238. Aldabra (Seychelles). 

columbiana (Kieffer), 1917b: 304 (Culicoides). Colombia. 

columna de Meillon and Wirth, 1989b: 213. Zimbabwe. 

communis Kieffer, 1918a: 55. Algeria. 

compta Yu and Hao, in Yu et al., 2005: 139. China. 

confinis (Johannsen), 1932: 426 (Tetraphora). Indonesia. 

connexa Yu, in Yu et al., 2005: 280. China. 

consuta Yu and Zhao, in Yu et al., 2005: 182. China. 

contexta Yu, in Yu et al., 2005: 183. China. 

contigua (Johannsen), 1932: 423 (Tetraphora). Indonesia. 

corinneae Gosseries, 1991: 42. New name for scutellata. 

scutellata (Meigen), 1830: 262 (Ceratopogon, preoccupied by Ceratopogon scutellatus Say, 1829). Europe. 

choneta Yu and Zou, in Yu et al., 2005: 279. China. 

cornuta Remm, 1972: 77. Russia. 

correntina Ronderos and Díaz, in Ronderos et al., 2004: 194. Argentina. 

coruscantis Yu and Zhang, in Yu et al., 2005: 141. China. 

crassipilosa Tokunaga, 1956: 114. Japan. 

crassiseta Borkent and Forster, 1986: 1285. Canada (Ontario). 

crassivena Mazumdar and Chaudhuri, 2009a: 193. India. 

cremnica Yu and Ding, in Yu et al., 2005: 299. China. 

croceoscutellata Goetghebuer, 1933e: 143. Democratic Republic of the Congo. 

cuneata Remm, 1979b: 57. Russia. 

curticoma Kieffer, 1923b: 138. Indonesia. 

curtiradialis Tokunaga, 1940d: 130. Japan, Taiwan. 

curtivitta Tokunaga, 1940d: 124. Japan. 

curta Yu and Yan, in Yu et al., 2005: 264. China. 

dampfi Kieffer, 1925b: 150. Estonia. 

estonica Kieffer, 1925a: 409. Estonia. 

turfacea Kieffer, 1925b: 151. Estonia. 

dara Pierce, 1966: 88. USA (California). Miocene. 

davidi Clastrier, 1983c: 27. Seychelles. 

decempunctata (Skuse), 1889: 301 (Ceratopogon). Australia (New South Wales). 

decoratissima (Strobl), 1910: 264 (Ceratopogon). Austria. 

deemingi Boorman and Harten, 2002: 441. Oman 

dehalperti Macfie, 1937b: 76. Ethiopia. 

deiras Yu, in Yu et al., 2005: 184. China. 

deleasma Yu, in Yu et al., 2005: 282. China. 

dellapei Díaz and Spinelli, in Díaz et al., 2010: 2828. Argentina. 

denticulata Yu, in Yu et al., 2005: 238. China. 

dentiforceps Tokunaga, 1940d: 121. Japan. 

desertorum Szadziewski, Gwizdalska-Kentzer and Gilka, 2011: 641. United Arab Emirates. 

dicrae Yu, in Yu et al., 2005: 142. China. 

digna Borkent, 1997a: 92. USA (Hawaii). 

dilatata Yu, in Yu et al., 2005: 143. China. 

dimota Remm, 1972: 79. Russia. 

dioxyria Hao and Yu, 2001: 12. China. 

diplosis Kieffer, 1914a: 232. Germany. 

distalis Kieffer, 1918a: 94. Turkey. 

divergens de Meillon, 1959b: 22. South Africa. 

62

diversiloba Yu, in Yu et al., 2005: 144. China. 

dixa Yu, Sun and Gong, in Sun et al., 2010:247. China. 

dominicana Szadziewski and Grogan, 1998b: 258. Dominican Republic. Miocene. 

doratos Yu, in Yu et al., 2005: 310. China. 

dujanga Yu, in Yu et al., 2005: 146. China. 

dunhuangensis Yu and Liu, in Yu et al., 2005: 185. China. 

dupliforceps Tokunaga, in Tokunaga and Murachi, 1959: 264. Kiribati. 

dybasi Tokunaga and Murachi, 1959: 291. Belau (USA). 

echinatus Liu, Yan and Liu, 1996: 50. China. 

echma Yu, in Yu et al., 2005: 300. China. 

effusa Liu and Yu, 2001: 277. China. 

egypti Macfie, 1943a: 149 (as variety of inconspicuosa). Egypt. 

enygros Yu, in Yu et al., 2005: 147. China. 

eodicryptoscenica Szadziewski, 1988: 228. Poland. Eocene. 

erema Yu and Huang, in Yu et al., 2005: 284. China. 

eremita Remm and Nazarmukhamedov, 1969: 56. Tajikistan. 

erici Havelka, 1978b: 62. Austria. 

ermeri Remm, 1967: 17. Georgia. 

erythrogaster (Santos Abreu), 1918: 307 (Culicoides). Canary Islands (Spain). 

esakii Tokunaga, 1940a: 212. Micronesia. 

europaea Remm, 1962a: 122 (as subspecies of bifurcata). Estonia. 

excelsa Yu and Deng, in Yu et al., 2005: 148. China. 

eximia (Santos Abreu), 1918: 312 (Culicoides). Canary Islands (Spain). 

falcata de Meillon and Downes, 1986: 152. South Africa. 

falculata Remm, in Remm and Zhogolev, 1968: 833. Ukraine. 

fallax Bose and Das Gupta, in Bose et al., 2002: 376. India. 

fasciigera Kieffer, 1925b: 151. Estonia. 

furcata Zilahi-Sebess, 1940: 50. Hungary. 

nigra Zilahi-Sebess, 1940: 50 (as variety of furcata). Hungary. 

fenerivensis de Meillon, 1961: 48. Madagascar. 

ferreyrai Díaz and Spinelli, in Díaz et al., 2011: 33. Argentina. 

ferruginosa Remm and Nazarmukhamedov, 1969: 54. Turkmenistan. 

festiva Wirth, 1952a: 161. USA (California). 

filibranchia (Lutz), 1914: 85 (Ceratopogon). Brazil. 

filiductus Díaz and Spinell, in Díaz et al., 2009:153. Argentina. 

flava Carter, Ingram and Macfie, 1921b: 196. Ghana. 

flava Yu, in Yu et al., 2005: 229. China. [preoccupied and needs new name] 

flaveola Clastrier, Rioux and Descous, 1961: 73. Chad. 

flavescens Tokunaga and Murachi, 1959: 250. Marshall Islands. 

flavibasalis Tokunaga, 1940e: 173 (1940d: 118). Micronesia. 

flavicauda Macfie, 1939c: 201. Brazil. 

flavicaudalis Tokunaga and Murachi, 1959: 248. Micronesia. 

flaviformis Carter, Ingram and Macfie, 1921b: 201. Ghana. 

flavifrons (Guérin-Méneville), 1833: 165 (Ceratopogon). France. 

obscura (Winnertz), 1852: 45 (Ceratopogon, preoccupied by Forcipomyia obscura (Walker), 1848). Germany. 

versicolor (Winnertz), 1852: 45 (Ceratopogon). Germany. 

dufouri (Laboulbène), 1869: 163 (Ceratopogon). France. 

hippocastani (Mik), 1888: 185 (Ceratopogon). Austria. 

rufithorax Strobl, 1910: 261 (Ceratopogon, as variety of versicolor). Croatia. 

brevitibialis Goetghebuer, 1919: 72. Belgium. 

goetghebueri Kieffer, 1919a: 53. Belgium. 

sensualis Kieffer, 1919a: 55. Croatia, Greece. 

lignicola Kieffer, 1919a: 57. Czech Republic. 

paludicola Kieffer, 1925b: 152. Estonia. 

63

oppressa Thomsen, 1935: 285. USA (New York). 

septuosa Borkent, in Borkent and Wirth, 1997:58. New name for obscura (Winnertz). 

flavimana (Santos Abreu), 1918: 292 (Culicoides, as variety of versicolor). Canary Islands (Spain). 

flavipicta Ingram and Macfie, 1922: 249. Ghana. 

flaviscapula Kieffer, 1918a: 93. Turkey. 

flaviventris (Goetghebuer), 1910: 96 (Culicoides). Belgium. 

flavohumeralis Goetghebuer, 1935d: 166. Democratic Republic of the Congo. 

flavopyga Zilahi-Sebess, 1940: 49. Hungary. 

flavoscapularis Goetghebuer, 1935d: 167. Democratic Republic of the Congo. 

flavoscutellata (Zetterstedt), 1850: 3648 (Ceratopogon). Norway. 

egens (Winnertz), 1852: 43 (Ceratopogon). Germany. 

heracleae Kieffer, 1919a: 59. Greece. 

halobia (Kieffer), 1924a: 12 (Prokempia (preoccupied by Dasyhelea halobia Kieffer, 1924a). Germany. 

floricola Kieffer, 1922g: 510 (as variety of begueti). Algeria. 

foliacea Yu, in Yu et al., 2005: 149. China. 

fontana de Meillon and Wirth, 1981b: 533. South Africa. 

forcepina Ma and Yu, in Yu et al., 2005: 243. China. 

forficata de Meillon and Wirth, 1989b: 215. Senegal. 

formosae (Kieffer), 1912a: 29 (Culicoides). Taiwan. 

formosana Kieffer, 1921g: 565. Taiwan. 

sauteri Tokunaga, 1940d: 127. New name for formosana. 

fornicata Yu and Liu, in Yu et al., 2005: 150. China. 

forsteri Grogan and Wirth, 1981c: 97. Solomon Islands. 

fossata Yu, in Yu et al., 2005: 152. China. 

fosteri Clastrier, 1983c: 28. Seychelles. 

franzella Goetghebuer, 1950: 3. Austria. 

unbedarfti Havelka, 1978a: 177. Germany. 

fratercula (Santos Abreu), 1918: 305 (Culicoides). Canary Islands (Spain). 

fueguina Díaz and Spinelli, in Díaz et al., 2010: 2832. Argentina. 

fulcillata Yu, in Yu et al., 2005: 153. China. 

fulvicauda Macfie, 1933b: 101. French Polynesia (France). 

fulvosa Remm, 1967: 15. Azerbaijan. 

fumala Tokunaga, 1940e: 176 (1940d: 117). Belau (USA). 

fungivora Krivosheina and Remm, 1974: 123. Russia. 

furcillifera Tokunaga, in Tokunaga and Murachi, 1959: 275. Micronesia. 

furcula de Meillon and Wirth, 1989b: 216. South Africa. 

furva Remm, 1967: 21. Russia. 

fusca Carter, Ingram and Macfie, 1921b: 204. Ghana. 

fusca Yu, in Yu et al., 2005: 154. China. [preoccupied and needs new name] 

fusciformis Carter, Ingram and Macfie, 1921b: 209. Ghana. 

fuscipleuris Carter, Ingram and Macfie, 1921b: 200. Ghana. 

fusciscutellata Carter, Ingram and Macfie, 1921b: 187. Ghana. 

fuscocincta Remm, 1967: 12. Azerbaijan. 

galbiscutellata Borkent 1997b: 7. New name for luteoscutellata. 

luteoscutellata Carter, Ingram and Macfie, 1921b: 191 (preoccupied by Dasyhelea luteoscutellata (Santos Abreu), 

1918). Ghana. 

galea Yu and Ma, in Yu et al., 2005: 287. China. 

gargola Díaz and Spinelli, in Díaz et al., 2010: 2834. Argentina. 

gedanica Szadziewski, 1988: 226. Poland. Eocene. 

gemma Yu, in Yu et al., 2005: 190. China. 

georgei Huttel and Huttel, 1954: 39. France. 

gigantosalpinx de Meillon, 1937b: 371. South Africa. 

gluchovae Brodskaya, 1996: 193 (as glukhovae). Kyrgyzstan. 

gnopheros Yu, in Yu et al., 2005: 156. China. 

64

gongylophoda Yu and Huang, 2006: 48. China. 

grata (Johannsen), 1932: 426 (Tetraphora). Indonesia. 

gressitti Tokunaga, in Tokunaga and Murachi, 1959: 251. Belau (USA). 

grisea (Coquillett), 1901a: 602 (Ceratopogon). USA (District of Columbia). 

subcaerulea Thomsen, 1935: 284. USA (New York). 

griseithorax Goetghebuer, 1933e: 145. Democratic Republic of the Congo. 

griseola Wirth, 1978: 195. Mexico. 

grogani Díaz and Spinelli, in Díaz et al., 2011:35. Chile. 

guadeloupensis Delécolle and Rieb, 1994: 272. Guadeloupe. 

halobia Kieffer, 1924a: 20. Germany. 

halophila Kieffer, 1911d: 5 (1913c: 255). Croatia. 

hama Yu, in Yu et al., 2005: 244. China. 

hamardabani Remm, 1972: 75. Russia. 

hamata de Meillon and Wirth, 1989b: 217. Zimbabwe. 

hanoiensis Yu, 2003a. Vietnam. 

hawaiiensis Macfie, 1934a: 133. USA (Hawaii). 

hei Yu, Sun and Gong, in Sun et al., 2010:246. China. 

heliophila Macfie, 1943a: 150 (as variety of inconspicuosa). Egypt. 

hesperos Yu and Yan, in Yu et al., 2005: 158. China. 

heurui Kieffer, 1923b: 139. Indonesia. 

heyuxiani Yu, Yuan and Zeng, in Yu et al., 2005: 687. China. 

hihifoi Clastrier and Delécolle, 1996: 311. Wallis and Futuna Islands (France). 

hippolytae Macfie, 1935a: 54. Brazil. 

hirsuta Yu, in Yu et al., 2005: 159. China. 

hirtipes (Kieffer), 1917b: 305 (Culicoides). Peru. 

hispaniolae Szadziewski and Grogan, 1998b: 260. Dominican Republic. Miocene. 

hitchcocki Wirth, 1976a: 381. Tonga. 

holosericea (Meigen), 1804: 27 (Ceratopogon). Europe. 

homocera Kieffer, 1919a: 56. Croatia. 

hondurensis Spinelli and Wirth, 1984d: 600. Belize. 

horrida Yu, in Yu et al., 2005: 210. China. 

huasteca Huerta and Ibanez-Bernal, 1999: 499. Mexico. 

humilis Macfie, 1938: 159. Solomon Islands. 

hutsoni Wirth, 1990: 240. Aldabra (Seychelles). 

imperfecta de Meillon and Wirth, 1989b: 217. Zimbabwe. 

incisurata Remm, 1962a: 114. Estonia. 

inconspicuosa Carter, Ingram and Macfie, 1921b: 191. Ghana. 

indecora (Kieffer), 1912a: 29 (Culicoides). Taiwan. 

infula Yu, in Yu et al., 2005: 191. China. 

insignicornis (Kieffer), 1913e: 9 (Culicoides). Kenya. 

insons Mayer, 1934c: 177. Indonesia. 

insperata Yu and Yan, in Yu et al., 2005: 192. China. 

insularis Tokunaga, 1940a: 214. Micronesia. 

insulicola Tokunaga and Murachi, 1959: 307. Marshall Islands. 

internata Yu, in Yu et al., 2005: 323. China. 

intonsa Debenham, 1987e: 347. Cocos (Keeling) Islands (Australia). 

ismailiae Macfie, 1943a: 152. Egypt. 

jamaicensis Spinelli and Wirth, 1984d: 602. Jamaica. 

jingboi Yu, in Yu et al., 2005: 288. China. 

johannseni (Malloch), 1915a: 311 (Pseudoculicoides). USA (California). 

joycei de Meillon, 1936: 193. South Africa. 

jucunda Macfie, 1932c: 38. New Zealand. 

judithae Pierce, 1966: 88. USA (California). Miocene. 

kanakoffi Pierce, 1966: 89. USA (California). Miocene. 

65

kibunensis Tokunaga, 1940d: 134. Japan. 

kitaokai Tokunaga, 1963c: 42. Japan. 

kodoriensis Remm, 1993: 195. Russia. 

korfoensis Remm, 1993: 193. Russia. 

kruppi Boorman and Harten, 2002: 444. Oman. 

kunmingensis Zhao and Yu, 1997: 6. China. 

kurensis Remm, 1967: 14. Azerbaijan. 

mayeri Spataru and Damian-Georgescu, 1970: 422. Romania. 

kyotoensis Tokunaga, 1940d: 132. Japan. 

kyrenica Remm, 1972: 75. Russia. 

labinoda Mazumdar and Chaudhuri, 2009a: 195. India. 

labourdonnaisi Clastrier, 1959c: 422. Réunion (France). 

lacustris Ingram and Macfie, 1931a: 190. Argentina. 

laeta (Johannsen), 1932: 427 (Tetraphora). Indonesia. 

lakistos Yu and Liu, in Yu et al., 2005: 230. China. 

larundae de Meillon, 1936: 195. South Africa. 

latiforceps Clastrier, 1983c: 29. Seychelles. 

ledi Remm, 1993: 190. Russia. 

lediformis Remm, 1993: 191. Russia. 

leporis de Meillon and Wirth, 1981c: 572. South Africa. 

leptobranchia Waugh and Wirth, 1976: 236. USA (Maryland). 

leptoclada Remm, 1967: 19. Azerbaijan. 

lieni Yu, in Yu et al., 2005: 311. China. 

lineata (Kieffer), 1917a: 186 (Culicoides). Papua New Guinea. 

linlingae Yu and Huang, 2006: 48. China. 

longicauda Yu, in Yu et al., 2005: 212. China. 

longicoma Kieffer, 1923b: 139. Indonesia. 

longicornis Kieffer, 1913d: 180. India. 

longipenis Yu, in Yu et al., 2005: 291. China. 

longituba Kieffer, 1922g: 511. Algeria. 

longquana Yu and Liu, in Yu et al., 2005: 312. China. 

longuria Yu, in Yu et al., 2005: 292. China. 

lucida Remm, in Remm and Zhogolev, 1968: 831. Georgia. 

ludingensis Zhang and Yu, 1996: 202. China. 

lutea Remm, in Remm and Zhogolev, 1968: 834. Ukraine. 

luteicauda Tokunaga, 1962a: 192. Japan. 

luteipalpis (Santos Abreu), 1918: 292 (Culicoides, as variety of versicolor). Canary Islands (Spain). 

luteocincta Kieffer, 1925e: 254. Egypt. 

luteogrisea Wirth and Williams, 1957: 10. Bermuda (Great Britain). 

luteola Goetghebuer, 1934d: 191. Democratic Republic of the Congo. 

luteoscutellata (Santos Abreu), 1918: 300 (Culicoides, as variety of sericatus). Canary Islands (Spain). 

macrostoma (Kieffer), 1910: 188 (Culicoides). India. 

maculata Macfie, 1943b: 119. Bahamas. 

magna Mazumdar and Chaudhuri, 2009a: 197. India. 

major (Malloch), 1915a: 311 (Pseudoculicoides). USA (Illinois). 

ampla Borkent, in Borkent and Wirth, 1997:51. Unnecessary new name for major. 

mayor (Strobl), 1906: 397 (Ceratopogon, as variety of sericata). Spain. 

aperta Goetghebuer and Timon David, 1937: 415. France. 

wuelkeri Mayer, 1959: 96. Spain. 

malleola Remm, 1962a: 119. Estonia. 

manassi Remm, 1980: 106. Kyrgyzstan. 

mangshi Yu and Yan, 2010: 201. China. 

maricola Remm, 1993: 195. Russia. 

maritima Tokunaga, 1940d: 126. Japan. 

66

matubae de Meillon, 1937b: 375. South Africa. 

mcmillani Clastrier and Wirth, 1961b: 333. Gambia. 

mediomunda Minaya, 1978: 79. Peru. 

meloae Díaz and Spinelli, in Díaz et al., 2011:37. Argentina. 

messersmithi Waugh and Wirth, 1976: 241. USA (Virginia). 

microsporea Hao and Yu, 2001: 13. China. 

minima Kieffer, 1913d: 181. India. 

minuscula (Skuse), 1889: 299 (Ceratopogon). Australia (New South Wales). 

minuticola Szadziewski and Grogan, 1998b: 262. Dominican Republic. Miocene. 

minutissima Goetghebuer, 1933e: 142. Democratic Republic of the Congo. 


miotheca Yu and Deng, in Yu et al., 2005: 245. China. 

mocambicana de Meillon, 1942c: 9. Mozambique. 

modesta (Winnertz), 1852: 43 (Ceratopogon). Germany. 

aestiva (Winnertz), 1852: 42 (Ceratopogon). Germany. 

longipalpis Kieffer, 1913b: 37. Germany. 

inclusa Kieffer, 1918b: 188. Czech Republic. 

strobli Kieffer, 1919a: 63. Spain. 

pratensis Goetghebuer, 1920: 44. Belgium. 

bihamata Kieffer, 1923a: 667. Algeria. 

moascari Macfie, 1943a: 153. Egypt. 

densipilosa Tokunaga, 1963c: 41. Japan. 

monilicornis Kieffer, 1923a: 671. Algeria. 

monosticta (Ingram and Macfie), 1923: 60 (Thysanognathus). Tanzania. 

monticola Ingram and Macfie, 1931a: 188. Chile. 

montivaga (Kieffer), 1910: 188 (Culicoides). India. 

montivegeta Yu, 2000b: 163. China. New name for montana. 

montana Yu and Ma, 1998: 274 (preoccupied by Dasyhelea montana Zilahi-Sebess, 1940). China. 

morrisoni Grogan and Wieners 2006: 468. Bahamas. 

morula Yu and Xue, in Yu et al., 2005: 247. China. 

multifascia Tokunaga and Murachi, 1959: 320. Micronesia. 

muradovi Remm, 1980: 108. Turkmenistan. 

mutabilis (Coquillett), 1901a: 602 (Ceratopogon). USA (District of Columbia). 

myrmedon Kieffer, 1917a: 188. Papua New Guinea. 

nachitai Yu and Liu, in Yu et al., 2005: 248. China. 

nandina Zhao and Yu, 1997: 6. China. 

navai Xue and Yu, 2002: 39. China. 

navaiae Waugh and Wirth, 1976: 240. USA (Michigan). 

necrophila Spinelli and Rodriguez, 1999: 59. Argentina. 

neglecta (Winnertz), 1852: 46 (Ceratopogon). Germany. 

neobifurcata Wirth, 1976c: 15. New name for bifurcata. 

bifurcata Wirth, 1952a: 161 (preoccupied by Dasyhelea bifurcata Kieffer, 1923a). USA (California). 


nepenthicola Wirth and Beaver, 1979: 47. Malaysia. 

nigella (Skuse), 1889: 300 (Ceratopogon). Australia (New South Wales). 

nigeriae Ingram and Macfie, 1921: 329. Nigeria. 

nigricans Carter, Ingram and Macfie, 1921b: 194. Ghana. 

nigrina Clastrier, Rioux and Descous, 1961: 71. Chad. 

nigripygma Tokunaga, in Tokunaga and Murachi, 1959: 310. Micronesia. 

nigristigmata Tokunaga and Murachi, 1959: 246. Marshall Islands. 

nigritula Clastrier, Rioux and Descous, 1961: 66. Chad. 

nigrofusca Carter, Ingram and Macfie, 1921b: 207. Ghana. 

nigroris Tokunaga and Murachi, 1959: 301. Micronesia. 

nilssoni Szadziewski, 2000b: 478. Canary Islands (Spain). 

67

nitida Yu and Liu, in Yu et al., 2005: 313. China. 

nitidula (Kieffer), 1910: 189 (Culicoides). Myanmar, India. 

noctuabunda Remm, 1980: 109. Tajikistan. 

norvegica Szadziewski and Hagan, 2000: 462. Norway. 

notata Goetghebuer, 1920: 47. Belgium. 

semistriata Goetghebuer, 1921: 176. Belgium. 

sziladyi Zilahi-Sebess, 1936b: 42. Hungary. 

nudipennis Kieffer, 1921b: 11. Cameroon. 

nyasae Ingram and Macfie, 1925: 286. Malawi. 

obscurella Goetghebuer, 1935d: 167. Democratic Republic of the Congo. 

occasa Zhang and Yu, 1996: 201. China. 


omoxantha Ingram and Macfie, 1922: 251. Nigeria. 

ona Díaz and Spinelli, in Díaz et al., 2010: 2836. Argentina. 

opaca (Kieffer), 1910: 188 (Culicoides). India. 

oreocincta Remm, 1980: 103. Kyrgyzstan. 

oribates Macfie, 1932c: 40. New Zealand. 

ornata Yu, in Yu et al., 2005: 160. China. 

ornaticornis Kieffer, 1913d: 180. India. 

pabloi Díaz and Spinelli, in Díaz et al., 2011:37. Chile. 

pacifica Macfie, 1933b: 99. French Polynesia (France). 

pallida Macfie, 1933b: 100 (as variety of pacifica). French Polynesia (France). 

pailemanensis Díaz and Spinelli, in Díaz et al., 2011:40. Argentina. 

paivai (Kieffer), 1910: 193 (Culicoides). India. 

palauensis Tokunaga, 1940e: 177 (1940d: 117). Belau (USA). 

pallens Wirth, 1952a: 152. USA (California). 

pallescentis Yu and Liu, in Yu et al., 2005: 215. China. 

pallidicola Yu, in Yu et al., 2005: 293. China. 

pallidihalter Carter, Ingram and Macfie, 1921b: 184. Ghana. 

pallidiscutellata Yu, in Yu et al., 2005: 163. China. 

pallidiventris (Goetghebuer), 1931: 211 (Tetraphora). Germany. 

olivacea Remm, 1962a: 117. Estonia. 

pallivittae Tokunaga, in Tokunaga and Murachi, 1959: 269. Micronesia. 

palloris Tokunaga and Murachi, 1959: 298. Mariana Islands (USA). 

paracincta Wirth, 1969: 576. Galápagos Islands (Ecuador). 

paracuminata Remm, 1980: 104. Tajikistan. 

paragrata Remm, 1972: 72. Russia. 

parahybae Macfie, 1940a: 74. Brazil. 

parallela Remm, 1962a: 125. Estonia. 

parcepilosa Kieffer, 1923a: 670. Algeria. 

parva Borkent, in Borkent and Wirth, 1997:57. New name for minuscula. 

minuscula Zilahi-Sebess, 1940: 51 (preoccupied by Dasyhelea minuscula (Skuse), 1889). New name for minima. 

minima Zilahi-Sebess, 1936b: 41 (preoccupied by Dasyhelea minima Kieffer, 1913d). Hungary. 

parvifulva Tokunaga, 1940d: 139. Taiwan. 

parvinigra Tokunaga, 1940d: 133. Taiwan. 

parvistylata Tokunaga and Murachi, 1959: 259. Micronesia. 


chilensis Ingram and Macfie, 1931a: 186. Chile. 

pauca Yu and Zhan, in Yu et al., 2005: 314. China. 

paulistana Forattini and Rabello, 1957: 245. Brazil. 

peculiopa Navai, 1994: 377. Afghanistan. 

peliliouensis Tokunaga, 1940e: 177 (1940d: 117). Belau (USA). 

pelletieri Clastrier, 1959c: 426. Réunion (France). 

penicillata Yu and Liu, in Yu et al., 2005: 216. China. 

68

pentalineata Wirth and Hubert, 1960a: 644. Mexico. 

perfida (Johannsen), 1932: 424 (Tetraphora). Indonesia. 

perrara Remm, 1980: 108. Turkmenistan. 

philotherma Macfie, 1953: 103. Costa Rica. 

picata Macfie, 1932b: 494. Malawi. 

pictiventris (Kieffer), 1911b: 327 (Culicoides). India. 

placenta Yu and Wang, in Yu et al., 2005: 315. China. 

platychaeta Hardy, 1960: 185. USA (Hawaii). 

pollex Borkent and Forster, 1986: 1286. Bahamas. 

pollinosa Wirth, 1952a: 156. USA (California). 

pritchardi Wirth, 1952a: 163. USA (California). 

pseudocincta Waugh and Wirth, 1976: 225 (as pseudocinta). USA (New York). 

pseudoincisurata Waugh and Wirth, 1976: 233. USA (Georgia). 

pulchripes (Santos Abreu), 1918: 308 (Culicoides). Canary Islands (Spain). 

tenerifensis Clastrier, 1966: 694. Canary Islands (Spain). 

pumila Macfie, 1939c: 202. Brazil. 

punctatipennis Kieffer, 1921b: 11. Cameroon. 

punctiformis Goetghebuer, 1935d: 168. Democratic Republic of the Congo. 

punctiventris Goetghebuer, 1940: 71. Germany. 

sericatoides Zilahi-Sebess, 1940: 53. Hungary. 

siccicola Remm, in Remm and Zhogolev, 1968: 832. Ukraine. 

punctosa Remm, 1993: 190. Russia. 

pusi Yu and Yan, in Yan and Yu, 2008: 481. China. 

pusilla (Lutz), 1913: 65 (Centrorhynchus). Brazil. 

pygmaea (Williston), 1896: 278 (Ceratopogon). St. Vincent. 

pyrsonota Macfie, 1953: 104. Costa Rica. 

qinghaiensis Liu and Shi, in Liu, et al., 2005: 32. China. 

quadrilobata (Kieffer), 1910: 189 (Culicoides). India. 

quarternihamata Tokunaga and Murachi, 1959: 295. Micronesia. 

quassata Yu, in Yu et al., 2005: 249. China. 

radialis Edwards, 1928: 53. Western Samoa. 

raoheensis Zou and Yu, in Yu et al., 2005: 218. China. 

raripilosa Tokunaga, 1940e: 172 (1940d: 117). Belau (USA). 

ravida Yu, in Yu et al., 2005: 199. China. 

retorta Ingram and Macfie, 1921: 333. Sierra Leone. 

revoluta Yu and Wen, in Yu et al., 2005: 200. China. 

reynoldsi Ingram and Macfie, 1931a: 185. Chile. 

ricardoi Díaz and Spinelli, in Díaz et al., 2011:41. Argentina. 

robustiforceps Tokunaga, 1962a: 187. Japan. 

rostrosericea Remm, 1993: 191. Russia. 

rugula Yu and Zhang, in Yu et al., 2005: 165. China. 

russa Macfie, 1933a: 79. French Polynesia (France). 

ryckmani Wirth and Hubert, 1960a: 646. USA (California). 

ryukyuensis Tokunaga, 1962a: 196. Japan. 


saetula Yu, in Yu et al., 2005: 318. China. 

sagittifera Tokunaga and Murachi, 1959: 292. Mariana Islands (USA). 

salinaria de Meillon and Wirth, 1981b: 530. South Africa. 

salta de Meillon and Downes, 1986: 155. South Africa. 

saltensis Spinelli and Wirth, 1984d: 606. Mexico. 

sanctaemariae Wirth, 1952a: 162. USA (California). 

santaemarthae (Kieffer), 1917b: 306 (Culicoides). Colombia. 

saprophila Kieffer, 1925a: 410. Germany. 

scapularis (Kieffer), 1910: 189 (Culicoides). India. 

69

schizothrixi Lee and Wirth, in Lee et al., 1989: 453. Singapore. 

schumanni Navai, 1994: 379. Afghanistan. 

scissurae Macfie, 1937a: 15. Trinidad and Tobago. 

koenigi Delécolle and Rieb, 1994: 267. Guadeloupe. 

scotti (Kieffer), 1911c: 341 (Culicoides). Seychelles. 

scutellaris Kieffer, 1918a: 93. Turkey. 

sella Yu, in Yu et al., 2005: 319. China. 

seminigra Navai, 1994: 381. Afghanistan. 

sericata (Winnertz), 1852: 34 (Ceratopogon). Germany. 

serristernum Remm, 1967: 18. Georgia. 

setigera (Kieffer), 1910: 190 (Culicoides). India. 

setoensis Tokunaga, 1940d: 137. Japan. 

sexlineata Goetghebuer, 1935d: 169. Democratic Republic of the Congo. 

seychellensis (Kieffer), 1911c: 341 (Culicoides). Seychelles. 

shannoni Ingram and Macfie, 1931a: 183. Chile. 

shaoshana Yu, in Yu et al., 2005: 304. China. 

siculla Yu and Zhao, in Yu et al., 2005: 201. China. 

signata Goetghebuer, 1935d: 169. Democratic Republic of the Congo. 

signicornecula Mazumdar and Chaudhuri, 2009a: 197. India. 

similaris Remm, 1972: 77. Russia. 

similinigrina Navai, 1994: 384. Afghanistan. 

similis Carter, Ingram and Macfie, 1921b: 189. Ghana. 

simillima (Johannsen), 1932: 423 (Tetraphora). Indonesia. 

simpliradialis Tokunaga, 1940d: 127. Japan. 

simulator Goetghebuer, 1935d: 170. Democratic Republic of the Congo. 

sinclairi Borkent, 1991: 110. Galápagos Islands (Ecuador). 

skierskae Szadziewski, 1985a: 91. Algeria. 

sonorensis Wirth, 1978: 197. Mexico. 

soriai Wirth and Waugh, 1976: 229. Brazil. 

soutini de Meillon and Wirth, 1987a: 38. South Africa. 

spathicerca Wirth, 1969: 577. Galápagos Islands (Ecuador). 

speciosa Clastrier, 1983c: 34. Seychelles. 

spiniforma Waugh and Wirth, 1976: 243. USA (New York). 

spinula Yu and Kong, in Yu et al., 2005: 296. China. 

stackelbergi Remm, 1993: 194. Russia. 

stanislavi Szadziewski, 1988: 230. Poland. Eocene. 

stellata Remm, in Remm and Zhogolev, 1968: 831. Ukraine. 

stemlerae Waugh and Wirth, 1976: 233. USA (Maryland). 

stenoceras Palmer, 1957: 271. USA (California). Miocene. 

sternalis Remm, 1980: 110. Tajikistan. 

storai Borkent, 1997b: 14. New name for canariensis. 

canariensis Storå, 1936: 36 (as variety of flavoscutellata, preoccupied by Dasyhelea canariensis (Santos Abreu), 

1918). Canary Islands (Spain). 

strigosa Kieffer, 1922g: 514. Algeria. 

striipennis Tokunaga, in Tokunaga and Murachi, 1959: 312. Micronesia. 

suarezi Spinelli and Ronderos, 1987: 11. Argentina. 

subaequalis Kieffer, 1915a: 472. Germany. 

subcommunis Yu, in Yu et al., 2005: 269. China. 

subehinata Yu, Zeng, and Yuan, in Yu et al., 2005: 687. China. 

subflava Yu and Hao, in Yu et al., 2005: 219. China. 

subgrata Tokunaga, 1961a: 180. Indonesia. 

sublettei Wirth, 1987: 73. USA (Texas). 

subperfida Tokunaga, 1940e: 174 (1940d: 117). Belau (USA). 

subscutellata Tokunaga, 1940e: 175 (1940d: 118). Belau (USA). 

70

subtilis Yu and Zhang, in Yu et al., 2005: 170. China. 

sudis Yu and Qi, in Yu et al., 2005: 250. China. 

suntari Remm, 1993: 195. Russia. 

surrecta Yu and Shi, in Yu et al., 2005: 252. China. 

symmetria Tokunaga, 1962a: 191. Japan. 

szegedensis Zilahi-Sebess, 1940: 53. Hungary. 

taibei Yu, in Yu et al., 2005: 233. China. 

taiwana Tokunaga, 1940d: 119. Taiwan. 

taiwaniensis Borkent, in Borkent and Wirth, 1997: 58. New name for formosana. 

formosana Kieffer, 1922b: 157 (preoccupied by Dasyhelea formosana Kieffer, 1921g). Taiwan. 

tamsi Wirth and Messersmith, 1977: 305. Seychelles. 

tarsalis Kieffer, 1919a: 62. Hungary. 

taurica Remm, in Remm and Zhogolev, 1968: 834. Ukraine. 

tenebrosa (Coquillett), 1905: 64 (Ceratopogon). USA (California). 

tenerrima Goetghebuer, 1935d: 170. Democratic Republic of the Congo. 

tenuipalpis Yu, in Yu et al., 2005: 320. China. 

ternipenis Yu and Yan, in Yu et al., 2005: 305. China. 

tersa (Johannsen), 1932: 421 (Tetraphora). Indonesia. 

tessicola Remm, 1972: 74. Russia. 

thalestris Macfie, 1935a: 55. Brazil. 

theobromatis Wirth and Derron, 1976: 232. Sao Tomé and Principe. 

thienemanni Spataru and Damian-Georgescu, 1970: 425 (as tienemanni). Romania. 

thompsoni de Meillon, 1936: 202. South Africa. 

thomsenae Wirth, 1952a: 160. USA (California). 

tianshana Yu and Ma, 1998: 273. China. 

tibestiensis Clastrier, Rioux and Descous, 1961: 61. Chad. 

tigalatensis (Santos Abreu), 1918: 303 (Culicoides, as variety of scutellatus). Canary Islands (Spain). 

townesi Tokunaga and Murachi, 1959: 267. Marshall Islands. 

transvaalensis de Meillon, 1959a: 335. South Africa. 

traverae Thomsen, 1935: 285. USA (New York). 

triaina Yu, in Yu et al., 2005: 255. China. 

trifasciata Kieffer, 1918a: 92. Turkey. 

trigonata Mayer, 1934c: 181. Indonesia. 

tristyla Wirth, 1952a: 165. USA (California). 

tropica Clastrier, 1959b: 369. Senegal. 

truncata Tokunaga and Murachi, 1959: 297. Belau (USA). 

tshatkalensis Remm, 1980: 106. Kyrgyzstan. 

tuberculata Remm, 1980: 109. Tajikistan. 

tugelae de Meillon, 1936: 197. South Africa. 

turanicola Remm and Nazarmukhamedov, 1969: 56. Kazakhstan. 

serrata Navai, 1994: 383. Afghanistan. 


grenieri Clastrier, 1966: 703. Canary Islands (Spain). 

malibui Yu, 2008:165. France. 

tymicola Remm, 1993: 194. Russia. 

uncinata Deng and Yu, in Yu et al., 2005: 256. China. 

undosternum Remm, 1972: 77. Russia. 

unguistyla Remm, 1972: 79. Russia. 

unicolor Remm, 1962a: 119. Estonia. 

upsilon de Meillon and Wirth, 1987a: 39. South Africa. 

varicornis (Santos Abreu), 1918: 288 (Culicoides). Canary Islands (Spain). 

velutina Remm, 1993: 193. Russia. 

verdieri Huttel and Huttel, 1951: 103. France. 

vidua Yu, in Yu et al., 2005: 171. China. 

71

villosipes (Kieffer), 1917b: 305 (Culicoides). Paraguay. 

viridans Forattini and Rabello, 1957: 247. Brazil. 

vittata Tokunaga, 1940d: 127. Japan. 

vittula Tokunaga, in Tokunaga and Murachi, 1959: 271. Guam (USA). 

vittulae Tokunaga, 1940d: 120. Japan. 

waldiae Macfie, 1937b: 77. Ethiopia. 

wanlina Yu and Li, in Yu et al., 2005: 325. China. 

williamsi Wirth and Waugh, 1976: 228. Trinidad and Tobago. 

winderi Wirth and Waugh, 1976: 233. Brazil. 

wirthi Clastrier, 1974c: 135. New name for insignicornis. 

insignicornis Clastrier, Rioux and Descous, 1961: 76 (preoccupied by Dasyhelea insignicornis (Kieffer), 1913e). 

Chad. 

wirthicola Szadziewski, 1985a: 94. Pakistan. 

wushi Ysmar and Yu, 2003: 52 (Yu and Ma, in Yu et al., 2005: 202). China. 

xichangensis Yu, in Yu et al., 2005: 235. China. 

yoshimurai Tokunaga, 1940d: 124. Japan. 

zavreli Kieffer, 1918b: 188. Czech Republic. 

zonalis Yu and Yang, in Yu et al., 2005: 298. China. 

SUBFAMILY CERATOPOGONINAE NEWMAN, 1834: 388 

TRIBE CULICOIDINI KIEFFER, 1911d: 1, 1911b: 319 

Genus CULICOIDES Latreille 

CULICOIDES Latreille, 1809: 251. Type species: Culicoides punctatus Latreille (= Ceratopogon punctatus Meigen), by 

monotypy. 

PADROSIA Rafinesque, 1815: 130 (unnecessary new name for Culicoides Latreille). Type species: Culicoides 

punctatus Latreille (= Ceratopogon punctatus Meigen), automatic. 

OECACTA Poey, 1853: 238. Type species: Oecacta furens Poey, by monotypy. 

PSYCHOPHAENA Philippi, 1865: 628. Type species: Psychophaena pictipennis Philippi (= Culicoides venezuelensis Ortiz 

and Mirsa), by monotypy. 

HAEMATOMYIDIUM Goeldi, 1905: 137. Type species: Haematomyidium paraense Goeldi, by original designation. 

COTOCRIPUS Brèthes, 1912: 451. Type species: Cotocripus caridei Brèthes, by monotypy. 

OXYHELEA Kieffer, 1921b: 14 (as subgenus of Culicoides). Type species: Culicoides dentatus Kieffer, by monotypy. 

DIPLOSELLA Kieffer, 1921h: 113 (as subgenus of Culicoides). Type species: Culicoides sergenti Kieffer, by monotypy. 

HAEMOPHORUCTUS Macfie, 1925: 349. Type species: Haemophoructus maculipennis Macfie, by monotypy. 

PROSAPELMA Kieffer, 1925a: 417. Type species: Prosapelma cinerea Kieffer, by original designation. 

SYNHELEA Kieffer, 1925a: 423. Type species: Culicoides tropicalis Kieffer, designation by Wirth et al., 1980: 160. 

HOFFMANIA Fox, 1948: 21 (as subgenus of Culicoides). Type species: Culicoides inamollae Fox and Hoffman (= 

Culicoides insignis Lutz), by original designation. 

BELTRANMYIA Vargas, 1953a: 34 (as subgenus of Culicoides). Type species: Culicoides crepuscularis Malloch, by original 

designation. 

SELFIA Khalaf, 1954: 38 (as subgenus of Culicoides). Type species: Culicoides hieroglyphicus Malloch, by original 

designation. 

MONOCULICOIDES Khalaf, 1954: 39 (as subgenus of Culicoides). Type species: Ceratopogon nubeculosus Meigen, by 


MACFIELLA Fox, 1955: 217 (as subgenus of Culicoides). Type species: Ceratopogon phlebotomus Williston, by original 

designation. 

AVARITIA Fox, 1955: 218 (as subgenus of Culicoides). Type species: Ceratopogon obsoletus Meigen, by original 

designation. 

TRITHECOIDES Wirth and Hubert, 1959: 2 (as subgenus of Culicoides). Type species: Culicoides flaviscutatus Wirth and 

Hubert, by original designation. 

ANILOMYIA Vargas, 1960: 37 (as subgenus of Culicoides). Type species: Culicoides covagarciai Ortiz, by original 

72

designation. 

DRYMODESMYIA Vargas, 1960: 40 (as subgenus of Culicoides). Type species: Culicoides copiosus Root and Hoffman, by 


DIPHAOMYIA Vargas, 1960: 40 (as subgenus of Culicoides). Type species: Culicoides baueri Hoffman, by original 

designation. 

GLAPHIROMYIA Vargas, 1960: 41 (as subgenus of Culicoides). Type species: Culicoides scopus Root and Hoffman, by 


MATAEMYIA Vargas, 1960: 43 (as subgenus of Culicoides). Type species: Culicoides mojingaensis Wirth and Blanton, by 


MEIJEREHELEA Wirth and Hubert, 1961: 23 (as subgenus of Culicoides). Type species: Ceratopogon guttifer de Meijere, 


PONTOCULICOIDES Remm, in Remm and Zhogolev, 1968: 840 (as subgenus of Culicoides). Type species: Culicoides 

tauricus Gutsevich, by original designation. 

CALLOTIA Vargas and Kremer, 1972: 242 (as subgenus of Culicoides). Type species: Culicoides saevus Kieffer, by original 

designation. 

WIRTHOMYIA Vargas, 1973: 112 (as subgenus of Culicoides). Type species: Culicoides segnis Campbell and Pelham- 

Clinton, by original designation. 

SENSICULICOIDES Shevchenko, 1977: 133 (as subgenus of Culicoides). Type species: Ceratopogon pictipennis Staeger, 


REMMIA Glukhova, 1977: 116 (as subgenus of Culicoides). Type species: Ceratopogon schultzei Enderlein, by original 

designation. 

SILVATICULICOIDES Glukhova, 1977: 117 (as subgenus of Culicoides). Type species: Ceratopogon fascipennis Staeger, 


NEOCULICOIDES Boorman and Lane, 1979: 327 (preoccupied by Neoculicoides Pierce, 1966). Type species: 

Neoculicoides taylori Boorman and Lane, by original designation. 

NULLICELLA Lee, 1982: 165 (as subgenus of Culicoides). Type species: Culicoides lasaensis Lee, (by original 

designation). 

SINOCOIDES Chu, 1983: 26 (as subgenus of Culicoides). Type species: Culicoides hamiensis Chu, Qian and Ma, by 


JILINOCOIDES Chu, 1983: 28 (as subgenus of Culicoides). Type species: Culicoides dunhuaensis Chu, by original 

designation. 

STIGMOCULICOIDES Isaev, 1988: 15 (as subgenus of Culicoides). Type species: Culicoides stigma (Meigen), by original 

designation. 

AMOSSOVIA Glukhova, 1989: 226 (as subgenus of Culicoides). Type species: Culicoides dendrophilus Amosova, by 


SILVICOLA Mirzaeva and Isaev, 1990: 98 (as subgenus of Culicoides). Type species: Culicoides grisescens Edwards, by 


TOKUNAGAHELEA Dyce and Meiswinkel, 1995: 131 (as subgenus of Culicoides). Type species: Culicoides mikros Dyce 

and Meiswinkel, by original designation. 

FASTUS Liu, in Yu et al., 2005: 1196 (as subgenus of Culicoides). Type species: Culicoides alpigenus Yu and Liu, by original 

designation. 

MARKSOMYIA Bellis and Dyce, 2011: 36 (as subgenus of Culicoides). Type species: Culicoides marksi Lee and Reye, by 


abbreviatipennis Statz, 1944: 142. Germany. Oligocene. 

abchazicus Dzhafarov, 1964: 263. Georgia. 

absitus Liu and Yu, 1990: 20. China. 

acanthostomus Wirth and Hubert, 1989: 92. Malaysia. 

accraensis Carter, Ingram and Macfie, 1920: 241. Ghana. 

achrayi Kettle and Lawson, 1955: 37. Great Britain. 

acotylus Lutz, 1913: 69. Brazil. 

panamericanus Fox, 1947: 90. Mexico. 

actoni Smith, 1929: 255 (1932: 180). India. 

okumensis Arnaud, 1956a: 119. Japan. 

73

imperceptus Das Gupta, 1962a: 538. India. 

adamskii Wirth, 1990: 242. Aldabra (Seychelles). 

adersi Ingram and Macfie, 1923: 56. Kenya and Tanzania. 

africanus Clastrier, 1959a: 186. Senegal. 

agamus Borkent, 1995: 63. Canada (Manitoba). Upper Cretaceous. 

agas Wirth and Hubert, 1989: 438. Indonesia. 

aitkeni Wirth and Blanton, 1968a: 214. Trinidad and Tobago. 

alachua Jamnback and Wirth, 1963: 187. USA (Florida). 

alahialinus Barbosa, 1952: 11 (1953: 12). Ecuador. 

alaskensis Wirth, 1951a: 84. USA (Alaska). 

alatavicus Gutsevich and Smatov, in Smatov and Isimbekov, 1971: 61. New name for fuscus. 

fuscus Gutsevich and Smatov, 1966: 71 (preoccupied by Culicoides fuscus Goetghebuer, 1952). Kazakhstan. 

alazanicus Dzhafarov, 1961b: 75. Azerbaijan. 

musilator Kremer and Callot, 1961: 693. France. 

albibasis Wirth and Hubert, 1959: 31. Malaysia. 

albicans (Winnertz), 1852: 41 (Ceratopogon). Germany. 

albifascia Tokunaga, 1937a: 319. Taiwan. 

albipennis Kieffer, 1919a: 32. France. 

albomaculus Root and Hoffman, 1937: 164. Mexico. 

albopunctatus Clastrier, 1960a: 84. Congo. 

albosparsus Kieffer, 1918a: 52. Ethiopia. 

albovenosus Khamala and Kettle, 1971: 53. Uganda. 

neoangolensis Kremer, 1972: 83. Angola. 

albuquerquei Wirth and Blanton, 1973: 424. Brazil. 

alexanderi Wirth and Hubert, 1962: 190. USA (Massachusetts). 

pseudopiliferus Wirth and Hubert, 1962: 189. USA (Maryland). 

alexandrae Dzhafarov, 1962a: 211. Azerbaijan. 

algeriensis Clastrier, 1957: 426. Algeria. 

alishanensis Chen, 1983: 80 (1988: 151). China. 

allantothecus Wirth and Hubert, 1989: 121. Malaysia. 

almeidae Cambournac, 1970b: 251. Portugal. 

almirantei Wirth and Blanton, 1959: 454. Panama. 

alpicola (Strobl), 1910: 263 (Ceratopogon, as variety of oculatus). Austria. 

alpigenus Yu and Liu, in Yu et al., 2005: 1197. China. 

altaicus Remm, 1972: 80. Russia. 

alticola Kieffer, 1913e: 11. Tanzania. 

alvarezi Ortiz, 1957: 161. Venezuela. 

amaniensis Khamala and Kettle, 1971: 25. Tanzania. 

ambericus Szadziewski and Grogan, 1998a: 40. Dominican Republic. Miocene. 

ameliae Browne, 1980: 543. Colombia. 

amossovae Remm, 1971: 205. Russia. 

analis Santos Abreu, 1918: 297. Canary Islands (Spain). 

andicola Wirth and Lee, 1967: 5. Colombia. 

andinus Wirth and Lee, 1967: 17. Colombia. 

andrewsi Causey, 1938: 410. Thailand. 

angkaensis Kitaoka, Takaoka and Choochote, 2005: 285. Thailand. 

angolensis Caeiro, 1961: 289. Angola. 

annandalei Majumdar and Das Gupta, in Gangopadhyay and Das Gupta, 2000: 125. India. 

annettae Spinelli and Borkent, 2004a: 365. Costa Rica. 

annosus Borkent, 1995: 65. Canada (Manitoba). Upper Cretaceous. 

annuliductus Wirth, in Vitale et al., 1981: 150. Panama. 

anophelis Edwards, 1922: 161. Malaysia and India. 

antefurcatus Wirth and Blanton, 1959: 315. Panama. 

antennalis Lee and Reye, 1953: 386. Australia (New South Wales). 

74

anthropophygas Yu annd Liu, in Yu et al., 2005: 892. China. 

antilleanus Szadziewski and Grogan, 1998a: 40. Dominican Republic. Miocene. 

antioquiensis Spinelli, Santamaría, Cabrera, Ronderos and Suárez, 2009: 82. Colombia. 

antunesi Forattini, 1954b: 315. Brazil. 

aomoriensis Kitaoka, 1991: 289. Japan. 

arabiensis Boorman, 1989: 173. Oman. 

aragaoi Tavares and Luna Dias, 1980: 393. Brazil. 

arakawai (Arakawa), 1910: 411 (Ceratopogon, as arakanae). Japan. 

sugimotonis Shiraki, 1913: 286. Taiwan. 

alboguttatus Kieffer, 1921g: 563. Taiwan. 

shimai (Sasaki), 1928: 687 (Ceratopogon). Japan. 

daleki Smith and Swaminath, 1932: 185. India. 

micropunctatus Tokunaga, 1951: 105. Indonesia. 

niigataensis Takahashi, 1958: 115 (as variety of arakawae). Japan. 

arboreus Gutsevich, 1952: 90. Russia. 

arboricola Root and Hoffman, 1937: 166. USA (Maryland). 

archboldi Wirth and Blanton, 1970b: 39. Dominica. 

ardentissimus Tokunaga, 1940a: 216. Micronesia. 

ardleyi Tokunaga, 1962b: 474. Papua New Guinea. 

arenarius Edwards, 1922: 164. Somalia. 

arenicola Howarth, 1985: 79. Laos. 

aricola Kieffer, 1922f: 391. Austria. 

arizonensis Wirth and Hubert, 1960a: 655. USA (Arizona). 

arnaudi Hubert and Wirth, 1961: 238. Japan. 

arubae Fox and Hoffman, 1944: 109. Netherlands Antilles (Netherlands). 

asiaticus Gutsevich and Smatov, 1966: 75. Kazakhstan. 

atchleyi Wirth and Blanton, 1969a: 559. USA (Alaska). 

atelis Wirth, 1982b: 249. Panama. 

aterinervis Tokunaga, 1937a: 312. Japan. 

atratus Statz, 1944: 147. Germany. Oligocene. 

atripalpis Wirth and Blanton, 1973: 325. Brazil. 

atripennis Shevchenko, 1972: 77. Ukraine. 

aureus Ortiz, 1951c: 585. Venezuela. 

miyamotoi Wirth and Blanton, 1953b: 231. Panama. 

austeni Carter, Ingram and Macfie, 1920: 261. Ghana. 

austerus Statz, 1944: 144. Germany. Oligocene. 

australiensis Kieffer, 1917a: 188. Papua New Guinea. 

austropalpalis Lee and Reye, 1955: 240. New name for palpalis. 

palpalis Lee and Reye, 1953: 380 (preoccupied by Culicoides palpalis Macfie, 1948). Australia (Queensland). 

austroparaensis Spinelli, in Spinelli et al., 2005: 141. Argentina. 

autumnalis Sen and Das Gupta, 1959a: 628. India. 

avilaensis Oritz and Mirsa, 1951: 593. Venezuela. 

azerbajdzhanicus Dzhafarov, 1962a: 211. Azerbaijan. 

azureus Wirth and Blanton, 1959: 377. Panama. 

badooshensis Khalaf, 1961: 460. Iraq. 

bahiensis Barbosa, 1947: 11. Brazil. 

bahrainensis Boorman, 1989: 177. Bahrain. 

baisasi Wirth and Hubert, 1959: 12. Philippines. 

bajensis Wirth and Moraes, 1979: 291. Mexico. 

bakeri Vargas, 1954: 27. Mexico. 

balsapambensis Ortiz and León, 1955: 569 (as variety of pifanoi). Ecuador. 

balticus Szadziewski, 1988: 41. Poland. Eocene. 

bambusicola Lutz, 1913: 62. Brazil. 

bancrofti Lee and Reye, 1953: 387. Australia (New South Wales). 

75

aniwa Felippe-Bauer, in Felippe-Bauer, et al., 2009: 852. Brazil. 

barbosai Wirth and Blanton, 1956a: 161. Panama. 

barnetti Wirth and Hubert, 1959: 32. Malaysia. 

barrosmachadoi Callot, Kremer and Molet, 1967: 39. Angola. 

barthi Tavares and Souza, 1978: 619. Brazil. 

bassetorum Callot, Kremer and Molet, 1973: 378. Lesotho. 

batesi Wirth and Blanton, 1973: 426. Brazil. 

sanmartini Wirth and Barreto, 1978: 553. Colombia. 

baueri Hoffman, 1925: 297. USA (Maryland). 

bayano Wirth, in Vitale et al., 1981: 152. Panama. 

beaveri Wirth and Barreto, 1978: 557. Colombia. 

beckae Wirth and Blanton, 1967: 213. USA (Florida). 

bedfordi Ingram and Macfie, 1923: 57. South Africa. 

begueti Clastrier, 1957: 432. Algeria. 

belemensis Wirth and Blanton, 1973: 427. Brazil. 

belgicus Kieffer, 1919a: 43. Belgium. 

belkini Wirth and Arnaud, 1969: 509. French Polynesia (France). 

benarrochi Ortiz and Mirsa, 1952b: 126 (as benarrochei). Venezuela. 

bergi Cochrane, 1973: 311. USA (New York). 

bermudensis Williams, 1956: 298. Bermuda (Great Britain). 

bernardae Itoua and Cornet, 1986: 244. Congo. 

besscus Liu and Yu, 1990: 15. China. 

beybienkoi Dzhafarov, 1962b: 183. Azerbaijan. 

biarcuatus Vimmer, 1932: 137. Israel. 

bickleyi Wirth and Hubert, 1962: 188. USA (Maryland). 

bicornus Liu and Yu, in Yu et al., 2005: 1288. China. 

bipalus Yu, 1991: 52. New name for biclavatus. 

biclavatus Deng and Yu, in Zhang et al., 1990: 27 (preoccupied by Culicoides biclavatus Kieffer, 1924a). China. 

bicolor Statz, 1944: 146. Germany. Oligocene. 

bicultellus Yu and Liu, 1990: 3. China. 

biestroi Spinelli and Ronderos, 1991: 86. Argentina. 

bifidus Borkent, 1996: 19. USA (New Jersey). Upper Cretaceous. 

bigeminus Wirth and Hubert, 1989: 336. Malaysia. 

biguttatus (Coquillett), 1901a: 604 (Ceratopogon). USA (District of Columbia). 

bilobatus Kieffer, 1912c: 6. Sri Lanka. 

bimaculatus Floch and Abonnenc, 1942b: 3. French Guiana. 

biscapus Kieffer, 1925a: 418. Vietnam. 

bisignatus Kieffer, 1921b: 14. Cameroon. 

bisolis Kremer and Brunhes, 1973: 287. Madagascar. 

blantoni Vargas and Wirth, 1955: 33. Mexico. 

bodemensis Clastrier and Delécolle, 1996: 302. Indonesia. 

bodenheimeri Vimmer, 1932: 138. Israel. 

bolitinos Meiswinkel, 1989: 30. South Africa. 

boliviensis Spinelli and Wirth, 1984a: 172. Bolivia. 

boophagus Macfie, 1937c: 116. Malaysia. 

boormani Giles and Wirth, 1985: 365. Malaysia. 

borinqueni Fox and Hoffman, 1944: 110. Puerto Rico (USA). 

bottimeri Wirth, 1955: 356. USA (Texas). 

multidentatus Atchley and Wirth, 1975: 1421. USA (California). 

bougainvillae Tokunaga, 1962b: 492. Papua New Guinea. 

boydi Wirth and Mullens, 1992: 1006. USA (California). 

brachcordylus Liu and Zhao, in Yu et al., 2005: 1290. China. 

brasilianum Forattini, 1956a: 81. Brazil. 

bredini Wirth and Blanton, 1970b: 41. Dominica. 

76

evipalpis Delfinado, 1961: 654. Philippines. 

brevipenis Mai and Yu, in Hao et al., 1990: 42. China. 

brevitarsis Kieffer, 1917a: 187. Australia. 

robertsi Lee and Reye, 1953: 386. Australia (Queensland). 

radicitus Delfinado, 1961: 657. Philippines. 

superfulvus Das Gupta, 1962b: 253. India. 

bricenoi Ortiz, 1951a: 445. Venezuela. 

brinchangensis Wirth and Hubert, 1989: 190. Malaysia. 

brodzinskyi Szadziewski and Grogan, 1998a: 40. Dominican Republic. Miocene. 

bromophilus Kieffer, 1922f: 391. Austria. 


brosseti Vattier and Adam, 1966a: 297. Gabon. 

brownei Spinelli, in Spinelli et al., 1993: 24. Colombia. 

brucei Austen, 1909: 282. Uganda. 

pseudopulicaris Goetghebuer, 1935d: 173. Democratic Republic of the Congo. 

hirtius de Meillon and Lavoipierre, 1944: 58. South Africa. 

brunnicans Edwards, in Edwards et al., 1939: 43. Great Britain. 

bubalus Delfinado, 1961: 658. Philippines. 

buettikeri Boorman, 1989: 179. Oman. 

buhetoensis Takahasi, 1941: 81. China. 

bullus Borkent, 1995: 61. Canada (Alberta). Upper Cretaceous. 

bundyensis Lee and Reye, 1955: 239. Australia (New South Wales). 

bunrooensis Lee and Reye, 1955: 238. Australia (Queensland). 

burylovae Glukhova and Khabirov, 1977: 52 (as burylovi). Tajikistan. 

butleri Wirth and Hubert, 1960a: 650. USA (Arizona). 

bwambanus de Meillon, 1952: 173. Uganda. 

byersi Atchley, 1967: 983. USA (New Mexico). 

cacticola Wirth and Hubert, 1960a: 653. USA (California). 

calcaratus Wirth and Hubert, 1989: 157. Malaysia. 

caldasi Browne, 1980: 535. Colombia. 

calexicanus Wirth and Rowley, 1971: 156. USA (California). 

californiensis Wirth and Blanton, 1967: 215. USA (California). 

caliginosus Goetghebuer, 1952: 2. Belgium. 

calloti Kremer, Delécolle, Bailly-Choumara and Chaker, 1979: 195. Morocco. 

cambodiensis Chu, 1986: 253. Cambodia. 

cameronensis Kitaoka, 1983: 94. Malaysia. 

cameroni Campbell and Pelham-Clinton, 1960: 222. Great Britain. 

camicasi Cornet and Chateau, 1971: 160. Senegal. 

camposi Ortiz and León, 1955: 569. Ecuador. 

fairchildi Wirth and Blanton, 1955a: 102. Panama. 

canadensis (Boesel), 1937: 47 (Atrichopogon). Canada (Manitoba). Upper Cretaceous. 

cancer Hogue and Wirth, 1968: 2. Costa Rica. 

cancrisocius Macfie, 1946: 15. Fiji. 

capillosus Borkent, in Borkent and Wirth, 1997:64. New name for hirtus. 

hirtus Xue and Yu, in Zhang et al., 1990: 30 (preoccupied by Culicoides hirtius de Meillon and Lavoipierre, 1944). 

China. 

capricorniae Dyce and Wirth, 1997: 586. Australia (Queensland). 

caprilesi Fox, 1952: 364. Venezuela. 

kintzi Wirth and Blanton, 1953a: 72. Panama. 

caridei (Brèthes), 1912: 451 (Cotocripus). Argentina. 

setifer (Lutz), 1913: 64 (Centrorhynchus). Brazil. 

carpenteri Wirth and Blanton, 1953a: 72. Panama. 

carpophilus Wirth and Hubert, 1989: 196. Malaysia. 

carri Pierce, 1966: 91. USA (California). Miocene. 

77

carsiomelas Wirth and Blanton, 1955a: 100. Panama. 

carvalhoi Wirth and Blanton, 1973: 429. Brazil. 

casei Grogan and Szadziewski, 1988: 809. USA (New Jersey). Upper Cretaceous. 

cassideus Zhang and Yu, in Zhang et al., 1990: 27. China. 

castillae Fox, 1946a: 251. Honduras. 

gibsoni Wirth, 1952d: 246. Guatemala. 

flochabonnenci Ortiz and Mirsa, 1952a: 267. Venezuela. 

cataneii Clastrier, 1957: 438. Algeria. 

catharinae Kremer, Delécolle and Braverman, 1991: 151. Egypt. 

caucaensis Wirth and Lee, 1967: 19. Colombia. 

caucoliberensis Callot, Kremer, Rioux and Descous, 1967: 827. France. 

causeyi Majumdar and Das Gupta, in Gangopadhyay and Das Gupta, 2000: 127. India. 

cavaticus Wirth and Jones, 1956: 166. USA (California). 

ceranowiczi Szadziewski, 1988: 45. Poland. Eocene. 

certus Das Gupta, 1962a: 537. India. 

ceylanicus Kieffer, 1912c: 5. Sri Lanka. 

chacoensis Spinelli and Wirth, 1984a: 174. Argentina. 

chagyabensis Lee, 1982: 169. China. 

changbaiensis Qu and Ye, 1995:95. China. 

charadraeus Arnaud, 1956a: 97. Japan. 

charruus Spinelli and Martinez, 1992: 176. Uruguay. 

chateaui Cornet, 1970: 357. Senegal. 

chaverrii Spinelli and Borkent, 2004a: 373. Costa Rica. 

chazeaui Clastrier and Delécolle, 1996: 300. Wallis and Futuna Islands (France). 

cheahi Kitaoka, 1983: 92. Malaysia. 

chengduensis Zhou and Lee, 1984a: 222. China. 

cheni Kitaoka and Tanaka, 1985: 43. Taiwan. 

chewaclae Glick and Mullen, 1983: 378. USA (Alabama). 

choochotei Kitaoka and Takaoka, in Kitaoka et al, 2005: 287. Thailand. 

chiopterus (Meigen), 1830: 263 (Ceratopogon). Europe. 

amoenus (Winnertz), 1852: 35 (Ceratopogon). Germany. 

similis (Goetghebuer), 1927a: 203 (Monohelea, preoccupied by Culicoides similis Carter, Ingram and Macfie, 1920). 

Belgium. 

dobyi Callot and Kremer, 1969a: 610. France. 

chitinosus Gutsevich and Smatov, 1966: 67. Kazakhstan. 

chrysonotus Wirth and Blanton, 1956b: 226. Panama. 

cilipes Kieffer, 1921d: 275. Latvia. 

cinereus (Kieffer), 1925a: 417 (Prosapelma). Slovak Republic. 

circumbasalis Tokunaga, 1959: 232. Indonesia. 

praesignis Delfinado, 1961: 652. Philippines. 

circumscriptus Kieffer, 1918a: 49. Tunisia. 

nadayanus Kieffer, 1918a: 95. Turkey. 

edwardsi Goetghebuer, 1921: 177. Belgium. 

algarum Kieffer, 1924a: 18. Germany. 

salicola Kieffer, 1924b: 405. Norway. 

pictidorsum Kieffer, 1924b: 406 (as variety of salicola). Norway. 

albonotatus Vimmer, 1932: 133 (preoccupied by Culicoides albonotatus Kieffer, 1918a). Israel. 

albosignatus Vimmer, 1932: 135. Israel. 

polymaculatus Vimmer, 1932: 135. Israel. 

pulcher Zilahi-Sebess, 1934: 155 (misspelled as pulscher). Bulgaria. 

kirovabadicus Dzhafarov 1964: 228. Azerbaijan. 

matsuensis Lien, Weng and Lin, 1996: 20 (valid subspecies of circumscriptus). Taiwan. 

meridionalis Xue, Liu and Yu, 2003: 112 (valid subspecies of circumscriptus). China. 

citroneus Carter, Ingram and Macfie, 1920: 259. Ghana. 

78

claggi Tokunaga, in Tokunaga and Murachi, 1959: 338. Bonin Islands. 

clarkei Carter, Ingram and Macfie, 1920: 246. Ghana. 

clastrieri Callot, Kremer and Deduit, 1962: 156. France. 

clavipalpis Mukerji, 1931: 1052. India. 

candidus Sen and Das Gupta, 1959a: 620. India. 

cleaves Liu, 1995: 9. China. 

clintoni Boorman, 1984: 164. Great Britain. 

clivus Yu and Liu, 1990: 3. China. 

coarctatus Clastrier and Wirth, 1961b: 312. Nigeria. 

cochisensis Wirth and Blanton, 1967: 216. USA (Arizona). 

cockerellii (Coquillett), 1901a: 603 (Ceratopogon). USA (Colorado). 

combinothecus Yu and Liu, in Yu et al., 1986: 211. China. 

commatis Wirth and Blanton, 1959: 321. Panama. 

comparis Liu and Yu, in Yu et al., 2005: 917. China. 

comosioculatus Tokunaga, 1956: 121. Japan. 

chaetophthalmus Amosova, 1957: 237. Russia. 

caucasicus Sergejev, 1959: 203 (as subspecies of chaetophthalmus). Russia. 

setosus Gutsevich, 1960: 106. Ukraine. 

conaensis Liu and Yu, 1990: 19. China. 

concatervans Liu and Yu, in Yu et al., 2005: 826. China. 

confusus Carter, Ingram and Macfie, 1920: 250. Ghana. 

congolensis Clastrier, 1960a: 98. Congo. 

continualis Qu and Liu, 1982: 102. China. 

contubernalis Ortiz and León, 1955: 574 (as variety of rozeboomi). Ecuador. 

copiosus Root and Hoffman, 1937: 171. Mexico. 

coracinus Borkent, in Borkent and Wirth, 1997:65. New name for niger. 

niger Tokunaga, 1941b: 98 (as nigrus, as variety of arcuatus, preoccupied by Culicoides niger Root and Hoffman, 

1937). China. 

cordiformis Kieffer, 1927: 61. Estonia. 

cordiger Macfie, 1934c: 193. Malaysia. 

magnesianus Lee and Reye, 1953: 385. Australia (Queensland). 

corneti Kremer, 1972: 92. Angola. 

corniculus Liu and Qu, 1981: 419. China. 

apiculatus Yu and Zhang, in Yu, 1988: 134. China. 

cornutus de Meillon, 1937b: 332. South Africa. 

coronalis Lee and Reye, 1955: 234. Australia (Queensland). 

corsicus Kremer, Leberre and Beaucournu-Saguez, 1971: 655. France. 

corsoni Ingram and Macfie, 1921: 324. Ghana. 

corti Causey, 1938: 411. Thailand. 

costalis Tokunaga, 1963b: 132. Papua New Guinea. 

coutinhoi Barretto, 1944: 96. Brazil. 

covagarciai Ortiz, 1950b: 457. Venezuela. 

beebei Fox, 1952: 366. Venezuela. 

crassipilosus Tokunaga, 1937a: 276. Japan. 

balius Arnaud, 1956a: 96. Japan. 

crassus Tokunaga, 1962b: 508. Papua New Guinea. 

crepuscularis Malloch, 1915a: 303. USA (Illinois). 

crescentis Wirth and Blanton, 1959: 317. Panama. 

crucifer Clastrier, 1968: 85. French Guiana. 

cuiabai Wirth, 1982b: 250. Brazil. 

culiciphagus Wirth and Hubert, 1959: 11. Solomon Islands. 

cummingi Spinelli and Borkent, 2004a: 379. Costa Rica. 

cunctans (Winnertz), 1852: 42 (Ceratopogon). Germany. 

cuniculus Lee and Reye, 1953: 389. Australia (Queensland). 

79

cylindricornis Wirth and Blanton, 1973: 430. Brazil. 

cylindripalpis Wirth and Hubert, 1989: 98. Malaysia. 

daedaloides Wirth and Blanton, 1959: 330. Panama. 

daedalus Macfie, 1948: 83. Mexico. 

dalessandroi Wirth and Barreto, 1978: 556. Colombia. 

damnosus Delfinado, 1961: 642. Philippines. 

dampfi Root and Hoffman, 1937: 169. Mexico. 

darlingtonae Wirth and Blanton, 1971a: 39. Trinidad and Tobago. 

dasyheleiformis Szadziewski, 1988: 36. Baltic region. Eocene. 

dasyophrus Macfie, 1940d: 27. Guyana. 

dasyops Clastrier, 1958b: 217. Senegal. 

davidi Spinelli, in Spinelli et al., 1993: 30. Colombia. 

daviesi Wirth and Blanton, 1968b: 251. Guyana. 

davisi Wirth and Rowley, 1971: 157. USA (Washington). 

deanei Felippe-Bauer and Wirth, 1987: 416. Brazil. 

debilipalpis Lutz, 1913: 60. Brazil. 

khalafi Beck, 1957: 104. USA (Florida). 

ichesi Ronderos and Spinelli, 1995b: 77. Argentina. 

decor (Williston), 1896: 281 (Ceratopogon). St. Vincent. 

definitus Sen and Das Gupta, 1959a: 622. India. 

defoliarti Atchley and Wirth, 1979: 527. USA (Arizona). 

dekeyseri Clastrier, 1958b: 214. Senegal. 

delfinadoae Wirth and Hubert, 1989: 386. Malaysia. 

deltus Edwards, in Edwards et al., 1939: 48. Great Britain. 

lupicaris Downes and Kettle, 1952: 76. Great Britain. 

dendrophilus Amosova, 1957: 240. Russia. 

reesi Bullock and Akiyama, 1959: 24. Japan. 

denisae Clastrier, 1971: 290. French Guiana. 

denisoni Boorman, 1988: 152. Greece. 

denningi Foote and Pratt, 1954: 20. Canada (Saskatchewan). 

dentatus Kieffer, 1921b: 13. Cameroon. 

denticulatus Wirth and Hubert, 1962: 193. USA (Wisconsin). 

dentiformis McDonald and Lu, 1972: 403. Taiwan. 

derisor Callot and Kremer, 1965: 330. France. 

desertorum Gutsevich, 1959: 675. Turkmenistan and Uzbekistan. 

pictus Khalaf, 1961: 455. Iraq. 

brevifrontis Smatov and Isimbekov, 1971: 62. Kazakhstan. 

oxianus Smatov, in Smatov and Kravets, 1976: 282. Kazakhstan. 

desytoculus Liu and Zhao, 1998: 10. China. 

dewulfi Goetghebuer, 1936: 320. Belgium. 

pseudochiopterus Downes and Kettle, 1952: 67. Great Britain. 

diabolicus Hoffman, 1925: 294. Panama. 

diamouanganai Itoua and Cornet, 1986: 237. Congo. 

dicrourus Wirth and Blanton, 1955b: 123. Panama. 

diffusus Spinelli, in Spinelli et al., 1993: 34. Brazil. 

digitalis Remm, 1973: 178. Mongolia. 

dikhros Tokunaga, 1962b: 507. Papua New Guinea. 

dingriensis Yu and Liu, in Yu et al., 2005: 1056. China. 

discrepans Ortiz and Mirsa, 1951: 595. Venezuela. 

dispar Clastrier, 1959a: 175. Senegal. 

dispersus Gutsevich and Smatov, 1966: 76. Kazakhstan. 

distinctipalpis Majumdar and Das Gupta, in Gangopadhyay and Das Gupta, 2000: 128. India. 

distinctipennis Austen, 1912: 101. Nigeria. 

multiguttatus (Goetghebuer), 1935d: 156 (Forcipomyia). Democratic Republic of the Congo. 

80

distinctus Sen and Das Gupta, 1959a: 618. India. 

diversus Felippe-Bauer, in Felippe-Bauer, et al., 2003: 1052. Peru. 

divisus Wirth and Hubert, 1989: 198. Malaysia. 

doeringae Atchley, 1967: 1014. USA (New Mexico). 

dominicanus Wirth and Blanton, 1970d: 146. Dominica. 

donggangensis Liu, Zhou and Wu, 2007:211. China. 

downesi Wirth and Hubert, 1962: 186. Canada (Ontario). 

dryadeus Wirth and Hubert, 1972: 41. Malaysia. 

duartei Tavares and Luna Dias, 1980: 395. Brazil. 

dubiosum Dyce and Wirth, 1997: 591. Australia (Queensland). 

dubitatus Kremer, Rebholtz-Hirtzel and Delécolle, 1976: 233. Angola. 

dubius Arnaud, 1956a: 100. Japan. 

duddingstoni Kettle and Lawson, 1955: 42. Great Britain. 

dukinensis Mirzaeva, 1985: 98. Russia. 

dumdumi Sen and Das Gupta, 1959a: 628. India. 

dungunensis Wirth and Hubert, 1989: 125. Malaysia. 

dunhuaensis Chu, 1983: 28. China. 

dunni Wirth and Blanton, 1959: 328. Panama. 

duodenarius Kieffer, 1922b: 157. Taiwan. 

dureti Ronderos and Spinelli, 1995a: 59. Paraguay. 

dutoiti de Meillon, 1943: 100. South Africa. 

vagus Cornet and Chateau, 1971: 156. Senegal. 

dycei Lee and Reye, 1953: 390. Australia (New South Wales). 

dzhafarovi Remm, 1967: 25. Azerbaijan. 

dzhafarovi Callot, Kremer, Molet and Bach, 1968: 95. France. 

eadsi Wirth and Blanton, 1971a: 37. USA (Texas). 

edeni Wirth and Blanton, 1974a: 23. USA (Florida). 

efferus Fox, 1952: 365. Peru. 

effusus Delfinado, 1961: 658. Philippines. 

elbeli Wirth and Hubert, 1959: 27. Malaysia. 

eldridgei Wirth and Barreto, 1978: 561. Colombia. 

elemae Pappas and Pappas, 1989: 228. USA (Nebraska). 

elizabethae Dyce and Wirth, 1997: 584. Papua New Guinea. 

elongatulus Statz, 1944: 143. Germany. Oligocene. 

elongatus Chu and Liu, 1978: 83. China. 

elutus Macfie, 1948: 75. Mexico. 

enderleini Cornet and Brunhes, 1994: 157. Senegal. 

engubandei de Meillon, 1937b: 337. South Africa. 

eoselficus Szadziewski, 1988: 43. Poland. Eocene. 

equatoriensis Barbosa, 1952: 13 (1953: 14) (as variety of debilipalpis). Ecuador. 

erairai Kono and Takahasi, 1940: 70. Japan. 

eremicus Yu and Liu, in Yu et al., 2005: 1296. China. 

erikae Atchley and Wirth, 1979: 532. USA (New Mexico). 

eriodendroni Carter, Ingram and Macfie, 1920: 250. Ghana. 

erkaensis Yu and Yang, in Yu, 1988: 135. China. 

esakii Esaki, 1939: 234. Micronesia. 


espinolai Felippe-Bauer and Lourenco-de-Oliveira, 1987: 149. Brazil. 

estevezae Ronderos and Spinelli, 1994: 47. Argentina. 

eublepharus Macfie, 1948: 86. Guyana. 

transferrans Ortiz, 1953a: 801. Venezuela. 

evansi Wirth and Blanton, 1959: 342. Panama. 

eupurus Dyce and Wirth, 1997: 582. Indonesia. 

excavatus Khamala and Kettle, 1971: 24. Uganda. 

81

expallens Remm, 1973: 176. Mongolia. 

exspectator Clastrier, 1959a: 177. Senegal. 

fadzili Kitaoka, 1983: 97. Malaysia. 

faghihi Navai, 1971: 199. Iran. 

fagineus Edwards, in Edwards et al., 1939: 147. Great Britain. 

farri Wirth and Blanton, 1970d: 148. Jamaica. 

fascipennis (Staeger), 1839: 594 (Ceratopogon). Denmark. 

distictus Kieffer, in Thienemann and Kieffer, 1916: 492. Sweden. 

albonotatus Kieffer, 1918a: 94. Turkey. 


felippebauerae Spinelli, in Spinelli et al., 2007: 660. Brazil. 

fernandezi Ortiz, 1954: 223. Venezuela. 

fernandoi Tavares and Sousa, 1979: 611. Brazil. 

ferreyrai Ronderos and Spinelli, 1995a: 61. Argentina. 

festivipennis Kieffer, 1914a: 235. Germany. 

odibilis Austen, 1921: 114. Israel. 

winnertzi Edwards, 1926a: 406. Germany. 

fieldi Wirth and Blanton, 1956c: 50. Honduras. 

filamentis Liu, Yan and Liu, 1996: 34. China. 

filariferus Hoffman, 1939: 172. Mexico. 

filicinus Gornostaeva and Gachegova, 1972: 522. Russia. 

filiductus Wirth, in Vitale et al., 1981: 155. Panama. 

filipalpis Remm, 1976b: 111. Russia. Upper Cretaceous. 

firuzae Dzhafarov, 1958: 245. Azerbaijan. 

fittkaui Wirth and Blanton, 1973: 432. Brazil. 

flavescens Macfie, 1937c: 114 (as variety of anophelis). Malaysia. 

subflavescens Wirth and Hubert, 1959: 14. Indonesia. 

flavidorsalis Tokunaga, 1959: 245. Papua New Guinea. 

flavidorsum Shevchenko and Lisetsky, 1969: 1413. Russia. 

flavimaculinotalis Tokunaga, 1940e: 180 (1940d: 142). Micronesia. 

flavipes Vimmer, 1932: 136. Israel. 

flavipulicaris Dzhafarov, 1964: 238. Azerbaijan. 

flavipunctatus Kitaoka, 1975: 199. Japan. 

flavirostris Vimmer, 1932: 137. Israel. 

flaviscriptus Tokunaga, 1959: 252. Papua New Guinea. 

flaviscutatus Wirth and Hubert, 1959: 34. Indonesia. 

flaviscutellaris Wirth and Hubert, 1989: 132. Malaysia. 

flavitibialis Kitaoka and Tanaka, 1985: 42. Taiwan. 

flavivenulus Costa Lima, 1937a: 418. Brazil. 

flavus Gornostaeva, 1980: 81 (as subspecies of grisescens). Russia. 

flavisomum Mirzaeva, 1984b: 67. Russia. 

anadyriensis Mirzaeva, 1984b: 70. Russia. 

flinti Wirth, 1982b: 251. Argentina. 

florenciae Messersmith, 1972: 167. Colombia. 

floridensis Beck, 1951: 135. USA (Florida). 

flukei Jones, 1956: 30. USA (Wisconsin). 

flumineus Macfie, 1937c: 116. Malaysia. 

fluvaitilis Xiang and Yu, in Ma et al., 1990: 47. China. 

fluvialis Macfie, 1940d: 25. Guyana. 

fluviatilis (Lutz), 1914: 82 (Johannseniella). Brazil. 

scorzai Ortiz, 1956: 93. Venezuela. 

foleyi Kieffer, 1922g: 503. Algeria. 

footei Wirth and Jones, 1956: 162. USA (Virginia). 

forattinii Ortiz, 1961: 211. Venezuela. 

82

fordae Lee, 1988: 53 (as fordae Wirth and Hubert). China. 

fordae Wirth and Hubert, 1989: 133 (preoccupied by Culicoides fordae Lee, 1988). Malaysia. 

fossicola Kieffer, 1922c: 236. Germany. 

fossilis Pierce, 1966: 91. USA (California). Miocene. 

foxi Ortiz, 1950c: 461. Puerto Rico (USA). 

fragmentum Tokunaga, 1962b: 507. Papua New Guinea. 

franclemonti Cochrane, 1974: 128. USA (New York). 

franklini Spinelli, in Spinelli et al., 1993: 45. Panama. 

freeborni Wirth and Blanton, 1969b: 217. USA (California). 

freitasi Wirth and Blanton, 1973: 434. Brazil. 

fretensis Wang and Yu, in Wang et al., 1990: 73. China. 

frohnei Wirth and Blanton, 1969b: 219. USA (Alaska). 

fukienensis Chen and Tsai, 1962: 395. China. 

fukudai Wada, 1990: 56. Japan. 

fulbrighti Lee and Reye, 1963: 361. Australia (New South Wales). 

fulvithorax (Austen), 1912: 105 (Johannseniella). Kenya. 

citrinus Kieffer, 1921b: 15. Cameroon. 

ruficollis Goetghebuer, 1935d: 174. Democratic Republic of the Congo. 

fulvus Sen and Das Gupta, 1959a: 628. India. 

furcillatus Callot, Kremer and Paradis, 1962: 771. France. 

furens (Poey), 1853: 238 (Oecacta). Cuba. 

maculithorax (Williston), 1896: 277 (Ceratopogon). St. Vincent. 

dovei Hall, 1932: 88. USA (Georgia). 

birabeni Cavalieri, 1966: 59. Venezuela. 

furensoides Williams, 1955: 271. USA (Michigan). 

dickei Jones, 1956: 28. USA (Wisconsin). 

fuscus Goetghebuer, 1952: 1. Belgium. 

fusipalpis Wirth and Blanton, 1973: 435. Brazil. 

gabaldoni Ortiz, 1954: 221. Venezuela. 

gabrieli Spinelli, Santamaría, Cabrera, Ronderos and Suárez, 2009: 82. Colombia. 

galindoi Wirth and Blanton, 1953a: 73. Panama. 

galliardi Callot, Kremer and Molet, 1973: 381. Lesotho. 

gambiae Clastrier and Wirth, 1961b: 308. Gambia. 

gaponus Yu, 1982: 202. China. 

garciai Wirth and Hubert, 1989: 304. Malaysia. 

gedanensis Szadziewski, 1988: 46. Poland. Eocene. 

gejgelensis Dzhafarov, 1964: 282. Azerbaijan. 

gemellus Macfie, 1934c: 192. Malaysia. 

geminus Macfie, 1937d: 472. Malaysia. 

bifasciatus Tokunaga, 1951: 104. Indonesia. 

gentilis Macfie, 1934c: 191. Malaysia. 

gentiloides Kitaoka and Tanaka, 1985: 41. Taiwan. 

geocheloneoides Dyce and Meiswinkel, 1995: 140. Indonesia. 

germanus Macfie, 1940d: 27. Guyana. 

gewertzi Causey, 1938: 409. Thailand. 

giganteus Khamala and Kettle, 1971: 35. Kenya. 

gigas Root and Hoffman, 1937: 172. Canada (Saskatchewan). 

ginesi Ortiz, 1951c: 586. Venezuela. 

glabellus Wirth and Blanton, 1956c: 47. Panama. 

glabrior Macfie, 1940d: 27 (as variety of debilipalpis). Guyana. 

grahambelli Forattini, 1956b: 35. Panama. 

glabripennis Goetghebuer, 1935d: 171 (1948b: 12). New name for nudipennis. 

nudipennis Goetghebuer, 1933e: 146 (preoccupied by Culicoides nudipennis Kieffer, 1922g). Democratic Republic 

of Congo. 

83

septemmaculatus Goetghebuer, 1935d: 175. Democratic Republic of the Congo. 

spinifer Khamala and Kettle, 1971: 51. Kenya. 

gladysae Kettle, Elson and Dyce, 1976: 173. Australia (Queensland). 

gluchovae Mirzaeva, 1974: 27. Russia. 

wushenensis Lee (as Li), 1974: 353. China. 

glushchenkoae Glukhova, 1989: 200. Russia 

goeldii Wirth and Blanton, 1973: 437. Brazil. 

gorgasi Wirth and Blanton, 1953b: 232. Panama. 

gornostaevae Mirzaeva, 1984a: 371. Russia. 

gouldi Wirth and Hubert, 1989: 139. Malaysia. 

gracilior Statz, 1944: 143. Germany. Oligocene. 

gracilipes Vaillant, 1954: 227. France. 

grahamii Austen, 1909: 280. Ghana. 

habereri Becker, 1909: 289. Cameroon. 

hostilissimus (Pittaluga), 1911a: 29 (1911b: 69; 1912: 597) (Oecacta). Guinea. 

kivuensis Goetghebuer, 1935d: 173. Democratic Republic of the Congo. 

trichopis de Meillon, 1937b: 328. Democratic Republic of the Congo. 

grandensis Grogan and Phillips, 2008: 197. USA (Utah). 

grandibocus Borkent, 1996: 19. USA (New Jersey). Upper Cretaceous. 

gregsoni Wirth and Blanton, 1969b: 222. Canada (British Columbia). 

grenieri Vattier and Adam, 1966b: 723. Congo. 

griffithi Wirth and Hubert, 1989: 307. Thailand. 

griseidorsum Kieffer, 1918a: 46. Algeria. 

grisescens Edwards, in Edwards et al., 1939: 146. Great Britain. 

remmi Damian-Georgescu, 1972: 16. Romania. 

arschanicus Mirzaeva, 1984b: 69. Russia. 

guadeloupensis Floch and Abonnenc, 1950b: 2. Guadeloupe. 

guamai Wirth and Blanton, 1973: 438. Brazil. 

guangxiensis Liu and Hao, 2003: 139. China. 

guarani Ronderos and Spinelli, 1994: 48. Argentina. 

guerrai Wirth and Blanton, 1971a: 41. Trinidad and Tobago. 

guineensis Kieffer, 1918a: 54. Guinea. 

gulbenkiani Caeiro, 1959: 155. South Africa. 

gutsevichi Sen and Das Gupta, 1958b: 163 (1959b: 65). New name for orientalis. 

orientalis Gutsevich, 1952: 92 (preoccupied by Culicoides orientalis Macfie, 1932b). Russia. 

bisulcatus Gutsevich, 1959: 681. New name for orientalis. 

guttatus (Coquillett), 1904c: 35 (Ceratopogon). Brazil. 

guttifer (de Meijere), 1907: 209 (Ceratopogon). Indonesia. 

guttipennis (Coquillett), 1901a: 603 (Ceratopogon). USA (Ohio). 

guyanensis Floch and Abonnenc, 1942a: 4. French Guiana. 

recifensis Barbosa, 1943: 263. Brazil. 

stubalensis Fox, 1946a: 254. Trinidad and Tobago. 

gymnopterus Edwards, 1926b: 247. Malaysia. 

haematopotus Malloch, 1915a: 302. USA (Illinois). 

haerbalingensis Liu and Wang, in Liu et al., 2010: 63. China. 

hainanensis Lee, 1975: 433. China. 

haitiensis Delécolle, Raccurt and Rebholtz, 1986: 108. Haiti. 

halonostictus Wirth and Hubert, 1989: 431. Thailand. 

hamiensis Chu, Qian and Ma, 1982: 109. China. 

hanae Braverman, Delécolle and Kremer, 1983: 678. Egypt. 

haranti Rioux, Descous and Pech, 1959: 432. France. 

hasegawai Kanasugi and Kitaoka, 2001:228. Japan. 

hawsi Wirth and Rowley, 1971: 159. USA (Washington). 

hayakawai Kitaoka, 1984: 301. Japan. 

84

hayesi Matta, 1967: 75. Honduras. 

hegneri Causey, 1938: 402. Thailand. 

heliconiae Fox and Hoffman, 1944: 108. Venezuela. 

rozeboomi Barbosa, 1947: 26. Trinidad and Tobago. 

heliophilus Edwards, 1921: 124. Great Britain. 

latifrontis Shakirzjanova, 1962: 256. Kazakhstan. 

kobachidzei Dzhafarov, 1964: 348. Georgia. 

helveticus Callot, Kremer and Deduit, 1962: 164. Switzerland. 

hengduanshanensis Lee, 1984: 88. China. 

henryi Lee and Reye, 1963: 360. Australia (Queensland). 

herero (Enderlein), 1908: 460 (Ceratopogon). Namibia. 

hermani Spinelli and Borkent, 2004a: 363. Panama. 

heteroclitus Kremer and Callot, in Callot and Kremer, 1965: 333 (Callot and Kremer, 1966: 941). France. 

hewitti Causey, 1938: 413. Thailand. 

hieroglyphicus Malloch, 1915a: 297. USA (Arizona). 

hildae Cornet and Nevill, 1979: 179. South Africa. 

himalayae Kieffer, 1911b: 326. India. 

hinmani Khalaf, 1952b: 353. USA (Oklahoma). 

hinnoi Howarth, 1985: 33. Laos. 

hirsutus Khamala and Kettle, 1971: 59. Uganda. 

hirtipennis Delfinado, 1961: 662. Philippines. 

hirtulus (Coquillett), 1900: 396 (Ceratopogon). USA (Alaska). 

hispanicolus Szadziewski and Grogan, 1998a: 41. Dominican Republic. Miocene. 

histrio Johannsen, 1946: 190 (as variety of guttifer). Guam (USA). 

mackayensis Lee and Reye, 1953: 383. Australia (Queensland). 

hitchcocki Spinelli and Wirth, 1984a: 176. Peru. 

hoffmani Fox, 1946a: 251. Trinidad and Tobago. 

hoffmanioides Wirth and Hubert, 1989: 173. Malaysia. 

hoguei Wirth and Moraes, 1979: 293. USA (California). 

hokkaidoensis Kitaoka, 1984: 303. Japan. 

holcus Lee, 1980: 85. China. 

hollandiensis Tokunaga, 1959: 226. Indonesia. 

hollensis (Melander and Brues), 1903: 13 (Ceratopogon). USA (Massachusetts). 

canithorax Hoffman, 1925: 284. USA (Georgia). 

homochrous Remm, in Remm and Zhogolev, 1968: 835. Ukraine. 

homotomus Kieffer, 1922b: 158. Taiwan. 

osakensis Iwata, 1935: 7. Japan. 

denmeadi Causey, 1938: 403. Thailand. 

hondurensis Spinelli and Borkent, 2004a: 369. Honduras. 

hornsbyensis Lee and Reye, 1963: 361. Australia (New South Wales). 

horridus Yu and Deng, in Yu, 1988: 135. China. 

hortensis Khamala and Kettle, 1971: 29. Uganda. 

horticola Lutz, 1913: 61. Brazil. 

bachmanni Spinelli, in Spinelli et al., 2005: 146. Argentina. 

huambensis Caeiro, 1961: 179 (also as huambenis). Angola. 

huayingensis Zhou and Lee, 1984b: 293. China. 

huaynacapaci Felippe-Bauer, in Felippe-Bauer et al., 2008b: 308. Peru. 

huberti Howarth, 1985: 35. Laos. 

huffi Causey, 1938: 406. Thailand. 

hui Wirth and Hubert, 1961: 16. Taiwan. 

hulinensis Liu and Yu, 1996: 135 (Liu et al., 1999: 341). China. 

humeralis Okada, 1941: 20. Japan. 

huochengensis Ma and Yu, in Ma et al., 1990: 47. China. 

husseyi Wirth and Blanton, 1971c: 76. USA (Florida). 

85

hyalinus Tokunaga, 1962b: 472. Papua New Guinea. 

hylas Macfie, 1940d: 26. Guyana. 

ibericus Dzhafarov, 1963: 54 (1964: 355). Azerbaijan. 

ibriensis Boorman, 1989: 181. Oman. 

ignacioi Forattini, 1957: 215. Brazil. 

saintjusti Tavares and Ruiz, 1980: 27. Brazil. 

iliensis Gutsevich and Smatov, 1966: 68. Kazakhstan. 

imicola Kieffer, 1913e: 11. Kenya. 

pallidipennis Carter, Ingram and Macfie, 1920: 265. Ghana. 

iraqensis Khalaf, 1957b: 343. Iraq. 

minutus Sen and Das Gupta, 1959a: 622. India. 

pseudoturgidus Das Gupta, 1962a: 537. India. 

imitator Ortiz, 1953b: 808. Venezuela. 

immaculatus Lee and Reye, 1953: 375. Australia (Queensland). 

impunctatus Goetghebuer, 1920: 55. Belgium. 

minor Tokunaga, 1941b: 97 (as variety of impunctatus). China. 

impusilloides Spinelli and Wirth, 1984a: 178. Brazil. 

incertus Yu and Zhang, in Yu, 1988: 136. China. 

indianus Macfie, 1932b: 488. India. 

cylindratus Kitaoka, 1980: 16. Japan. 

inermis Spinelli, Santamaría, Cabrera, Ronderos and Suárez, 2009: 84. Colombia. 

inexploratus Sen and Das Gupta, 1959a: 628. India. 

inflatipalpalis Tokunaga, 1963b: 125. Papua New Guinea. 

infulatus Delfinado, 1961: 644. Philippines. 

innoxius Sen and Das Gupta, 1959a: 626. India. 

inornatipennis Carter, Ingram and Macfie, 1920: 227. Ghana. 

inornatithorax Das Gupta, 1963: 36. India. 

insignipennis Macfie, 1937d: 469. Malaysia. 

insignis Lutz, 1913: 51. Brazil. 

inamollae Fox and Hoffman, 1944: 110. Puerto Rico (USA). 

painteri Fox, 1946a: 257. Honduras. 

insinuatus Ortiz and León, 1955: 577. Ecuador. 

insolatus Wirth and Hubert, 1960a: 654. Mexico. 

insolens Choudhuri and Das Gupta, in Choudhuri et al., 1986: 54. India. 

insulanus Macfie, 1933a: 77. French Polynesia (France). 

insularis Kitaoka, 1980: 18. Japan. 

interrogatus Lee and Reye, 1963: 358. Australia (New South Wales). 

inthanonensis Kitaoka, Takaoka and Choochote, 2005: 288. Thailand. 

inyoensis Wirth and Blanton, 1969a: 565. USA (California). 

iphthimus Zhou and Lee, 1984b: 295. China. 

iranicus Navai, 1971: 202. Iran. 

iriartei Fox, 1952: 368. Venezuela. 

vargasi Wirth and Blanton, 1953a: 74. Panama. 

iriomotensis Kitaoka, 1975: 198. Japan. 

irwini Spinelli and Wirth, 1984a: 180. Chile. 

isechnoensis Glick, 1990: 108. Kenya. 

isioloensis Cornet, Nevill and Walker, 1974: 240. Kenya. 

isoregalis Majumdar and Das Gupta, in Majumdar et al., 1997: 28. India. 

jacksoni Atchley, 1970: 258. USA (New Mexico). 

jacobsoni Macfie, 1934d: 215. Indonesia. 

buckleyi Macfie, 1937c: 117. Malaysia. 

kitaokai Tokunaga, 1955: 6. Japan. 

unisetiferus Tokunaga, 1959: 236. Papua New Guinea. 

jamaicensis Edwards, 1922: 165 (as variety of loughnani). Jamaica. 

86

jamesi Fox, 1946b: 244. USA (Montana). 

jamnbacki Wirth and Hubert, 1962: 192. USA (New York). 

japonicus Arnaud, 1956a: 106. Japan. 

litoreus Amosova, 1957: 241. Russia. 

javae Tokunaga, 1951: 103. Indonesia. 

javanicus Salm, 1917b: 139. Indonesia. 

jefferyi Kitaoka, 1983: 96. Malaysia. 

jianfenglingensis Liu, 1995: 9. China. 

jimmiensis Tokunaga, 1959: 229. Papua New Guinea. 

jiulongensis Liu and Wang, in Liu, et al., 2011: 170. China. 

jouberti Huttel, Huttel and Verdier, 1953: 98. Gabon. 

jonesi Wirth and Hubert, 1960a: 650. USA (Texas). 

jucundus Statz, 1944: 147. Germany. Oligocene. 

judaeae Macfie, 1933c: 79. Israel. 

juddi Cochrane, 1973: 316. USA (New York). 

jumineri Callot and Kremer, 1969b: 1112. Tunisia. 

jurbergi Felippe-Bauer, in Felippe-Bauer, et al., 2005: 51. Peru. 

jurensis Callot, Kremer and Deduit, 1962: 160. France. 

jurutiensis Trindade and Felippe-Bauer, 2011a:61. Brazil. 

kadenensis McDonald, Bolinguit and Lu, 1973: 639. Japan. 

kaifongensis Yu, 1982: 202. China. 

kaimosiensis Khamala and Kettle, 1971: 58. Kenya. 

kaluginae Remm, 1976b: 110. Russia. Upper Cretaceous. 

kampa Felippe-Bauer, Veras, Castellon and Moreira, 2000: 35. Brazil. 

kamrupi Sen and Das Gupta, 1958b: 163 (1959a: 617; 1959b: 65). New name for albipennis. 

albipennis Smith and Swaminath, 1932: 184 (preoccupied by Culicoides albipennis Kieffer, 1919a). India. 

kanagai Khamala and Kettle, 1971: 49. Kenya. 

kangdingensis Yu and Liu, in Yu et al., 2005: 1204. China. 

karagiensis Smatov and Aldabergenov, 1973: 24. Kazakhstan. 

karakumensis Gutsevich and Molotova, in Gutsevich, 1973: 180. Turkmenistan. 

karenensis Glick, 1990: 162. Kenya. 

kasimi Khalaf, 1961: 464. Iraq. 

kayi Bellis and Dyce, 2011: 46. Australia (Western Australia). 

kelantanensis Wirth and Hubert, 1989: 346. Malaysia. 

kelinensis Lee, 1978: 58 (1979a: 99). China. 

kepongensis Lee, 1988: 69 (as kepongensis Wirth and Hubert). China. 

kepongensis Wirth and Hubert, 1989: 346 (preoccupied by Culicoides kepongensis Lee, 1988). Malaysia. 

kerichoensis Khamala and Kettle, 1971: 37. Kenya. 

kettlei Breidenbaugh and Mullens, 1999: 150. USA (California). 

kibatiensis Goetghebuer, 1935d: 172. Democratic Republic of the Congo. 

volatilis Goetghebuer, 1935d: 176. Democratic Republic of the Congo. 

kibunensis Tokunaga, 1937a: 298. Japan. 

cubitalis Edwards, in Edwards et al., 1939: 40. Great Britain. 

ponkikiri Kono and Takahasi, 1940: 74. Japan. 

sitinohensis Okada, 1941: 18. Japan. 

kinabaluensis Wirth and Hubert, 1989: 211. Indonesia. 

kinari Howarth, 1985: 44. Laos. 

kingi Austen, 1912: 104. Sudan. 

nilotes Kieffer, 1925e: 257. Egypt. 

kirbyi Glick and Mullen, 1983: 380. USA (Maryland). 

kirgizicus Glukhova, 1973: 111. Kyrgyzstan. 

kirinensis Lee, 1976: 47. China. 

kisangkini Howarth, 1985: 47. Laos. 

klossi Edwards, 1933c: 252. Malaysia. 

87

knowltoni Beck, 1956: 136. USA (Florida). 

kobae Cornet and Chateau, 1971: 148. Senegal. 

kohistanensis Kitaoka and Shinonaga, 1989: 29. Pakistan. 

kolymbiensis Boorman, 1988: 154. Greece. 

komiensis McDonald, Bolinguit and Lu, 1973: 641. Japan. 

komarovi Mirzaeva, 1985: 99. Russia. 

konglinensis Yu and Kong, in Xue et al., 1992: 52. China. 

kongmiaoensis Liu and Zhou, 2006: 468. China. 

koreensis Arnaud, 1956a: 109. South Korea. 

palustris Amosova, 1957: 243. Russia. 

korossoensis (Huttel and Huttel), 1952d: 472 (Monohelea). Mali. 

kotonkan Boorman and Dipeolu, 1979: 35. Nigeria. 

krameri Clastrier, 1959a: 194. Senegal, Ivory Coast and Mali. 

ciliodentatus Khamala and Kettle, 1971: 28. Uganda. 

kribiensis Kieffer, 1921b: 18. Cameroon. 

krombeini Giles, Wirth and Messersmith, 1981: 542. Sri Lanka. 

kucheensis Liu and Yu, in Yu et al., 2005: 1302. China. 

kugitangi Ataev, 1976: 79. Turkmenistan. 

kumbaensis Callot, Kremer, Mouchet and Bach, 1965: 540. Cameroon. 

kurensis Dzhafarov, in Gutsevich, 1960: 100. Azerbaijan. 

kuripako Felippe-Bauer, in Felippe-Bauer et al., 2010: 863. Brazil. 

kusaiensis Tokunaga, 1940a: 215. Micronesia. 

kuscheli Wirth and Blanton, 1978: 236. Chile. 

kyushuensis Wada, 1986: 146. Japan. 

labis Yu and Liu, in Yu et al., 2005: 1303. China. 

lacustris Ronderos, 1990a: 116. Argentina. 

lahillei (Iches), 1906: 264 (Ceratopogon). Argentina. 

lahontan Wirth and Blanton, 1969b: 223. USA (California). 

laimargus Zhou and Lee, 1984a: 221. China. 

lamborni Ingram and Macfie, 1925: 283. Malawi. 

landauae Kremer, Rebholtz-Hirtzel and Bailly-Choumara, 1975: 206. Morocco. 

lanei Ortiz, 1950a: 431. Panama. 

langeroni Kieffer, 1921i: 262. Tunisia. 

lansangensis Howarth, 1985: 58. Laos. 

lanyuensis Kitaoka and Tanaka, 1985: 39. Taiwan. 

laoensis Howarth, 1985: 37. Laos. 

laoshanensis Yu and Kang, in Yu, 1988: 137. China. 

lasaensis Lee, 1978: 63 (1979a: 100). China. 

latifrons Khamala and Kettle, 1971: 23. Uganda. 

latipennis Kieffer, 1919a: 37. Romania, Ukraine. 

laurae Pierce, 1966: 91. USA (California). Miocene. 

leanderensis Lee and Reye, 1963: 355. Australia (Queensland). 

leechi Wirth, 1977: 53. USA (California). 

leei Tokunaga, 1960b: 72. Papua New Guinea. 

leizhouensis Lai and Yu, in Lai et al., 1990: 69. China. 

lenae Glushchenko and Mirzaeva, 1970: 37. Russia. 

lengi Yu and Liu, 1990: 10. China. 

lenti Tavares and Luna Dias, 1980: 396. Brazil. 

leoni Barbosa, 1952: 17 (1953: 19). Ecuador. 

leopoldoi Ortiz, 1951c: 579. Venezuela. 

leucostictus Kieffer, 1911c: 340. Seychelles. 

praetermissus Carter, Ingram and Macfie, 1920: 240. Ghana. 

egypti Macfie, 1924: 66 (as variety of distinctipennis). Egypt. 

pharao Kieffer, 1925e: 259. Egypt. 

88

lichyi Floch and Abonnenc, 1949: 1 (1950a: 69). Venezuela. 

lieni Chen, 1979: 99. Taiwan. 

liliputanus Statz, 1944: 142. Germany. Oligocene. 

limai Barretto, 1944: 99. Brazil. 

limonensis Ortiz and León, 1955: 576. Ecuador. 

lingshuiensis Lee, 1975: 434. China. 

lini Kitaoka and Tanaka, 1985: 46. Taiwan. 

lisicarruni Moncada, Carrasqui9lla and Spinelli, 2010: 978. Colombia. 

liui Wirth and Hubert, 1961: 20. Taiwan. 

liubaensis Liu and Wu, 2005:448. China. 

liukueiensis Kitaoka and Tanaka, 1985: 47. Taiwan. 

lobatoi Felippe-Bauer, in Felippe-Bauer and Quintelas, 1994: 25. Brazil. 

loisae Jamnback, 1965: 77. USA (New York). 

longicercus Kitaoka, 1980: 19. Japan. 

longicollis Glukhova, 1971: 507. Ukraine. 

longidens Arnaud, 1956a: 111. Japan. 

paucidentatus Kitaoka, 1973: 216. Japan. 

longior Hagan and Reye, 1986: 340. Australia (Queensland). 

longipalpis Delfinado, 1961: 645. Philippines. 

longipennis Khalaf, 1957b: 348. Iraq. 

flavisimilis Dzhafarov, 1964: 291. Azerbaijan. 

longiporus Chu and Liu, 1978: 86. China. 

longiradialis Tokunaga, 1962b: 499. Papua New Guinea. 

longirostris Qu and Wang, 1994: 486. China. 

longlinensis Yu, 1982: 202. China. 

paradoxus Yu and Liu, 1990: 2. China. 

longzhouensis Hao and Yu, 1990: 40. China. 

lopesi Barretto, 1944: 102. Brazil. 

lophortygis Atchley and Wirth, 1975: 1422. USA (California). 

loughnani Edwards, 1922: 165. Jamaica. 

loxodontis Meiswinkel, 1992: 147. South Africa. 

lungchiensis Chen and Tsai, 1962: 397 (as subspecies of peregrinus). China. 

megaforticeps Kitaoka, 1973: 212. Japan. 

luglani Jones and Wirth, 1958: 89. USA (Texas). 

lui Yu and Liu, 1990: 10. China. 

lulianchengi Chen, 1983: 93 (1988: 152). China. 

lutealaris Wirth and Blanton, 1956b: 225. Panama. 

luteolus Wirth and Hubert, 1989: 100. Malaysia. 

luteovenus Root and Hoffman, 1937: 156. Mexico. 

lutzi Costa Lima, 1937a: 419. Brazil. 

lyrinotatus Wirth and Blanton, 1955b: 126. Panama. 

maai Wirth and Hubert, 1989: 312. Malaysia. 

macclurei Wirth and Hubert, 1989: 348. Malaysia. 

macfiei Causey, 1938: 411. Thailand. 

macintoshi Cornet and Nevill, 1980: 383. South Africa. 

macieli Tavares and Ruiz, 1980: 29. Brazil. 

mackerrasi Lee and Reye, 1963: 355. Australia (Northern Territory). 

macrostigma Wirth and Blanton, 1953b: 230. Panama. 

maculipennis (Macfie), 1925: 349 (Haemophoructus). Singapore. 

maculiscutellaris Tokunaga, 1959: 249. Indonesia. 

maculitibialis Lien, Weng and Lin. 1997: 171. Taiwan. 

madagascarensis de Meillon, 1961: 40. Madagascar. 

magnificus Sen and Das Gupta, 1959a: 622. India. 

magnipalpis Wirth and Blanton, 1953a: 76. Panama. 

89

magnipictus Tokunaga, 1962b: 494. Papua New Guinea. 

magnithecalis Majumdar and Das Gupta, in Gangopadhyay and Das Gupta, 2000: 124. India. 

magnus Colaco, 1946: 219, 237 (as variety of hirtius). South Africa. 

majorinus Chu, 1977: 99. China. 

malariologiensis Perruolo, 1990: 28. Venezuela. 

malayae Macfie, 1937d: 471. Malaysia. 

malevillei Kremer and Coluzzi, 1971: 415. Italy. 

mamaensis Lee, 1979a: 101. China. 

mammalicolus Szadziewski and Grogan, 1998a: 41. Dominican Republic. Miocene. 

manchuriensis Tokunaga, 1941b: 98. China. 

setiger Goetghebuer, 1938: 379 (preoccupied by Dasyhelea setigera (Kieffer), 1910). Belgium. 

goetghebueri Arnaud, 1956b: 94. Belgium. New name for setiger. 

machardyi Campbell and Pelham-Clinton, 1960: 235. Great Britain. 

ochraceimaculatus Shevchenko, 1970b: 8. Ukraine. 

ochraceipennis Shevchenko, 1970b: 9. Ukraine. 

mesostigma Remm, 1971: 201. Russia. 

vistulensis Skierska, 1973: 289. Poland. 

manhauensis Yu, 1982: 202. China. 

manikumari Wirth and Hubert, 1989: 103. Malaysia. 

marcleti Callot, Kremer and Basset, 1968: 271. Algeria. 

marginalis Lee and Reye, 1963: 354. Australia (New South Wales). 

marginatus Delfinado, 1961: 646. Philippines. 

marginus Chu, 1984: 24. New name for marginalis. 

marginalis Chu and Liu, 1978: 84 (preoccupied by Culicoides marginalis Lee and Reye, 1963). China. 

margipictus Qu and Wang, 1994: 487. China. 

marinkellei Wirth and Lee, 1967: 13. Colombia. 

marinus Yu and Zhu, in Lai et al., 1990: 70. China. 

maritimus Kieffer, 1924a: 16. Germany. 

submaritimus Dzhafarov, 1962a: 206. Azerbaijan. 

marksi Lee and Reye, 1953: 392. Australia (New South Wales). 

marmoratus (Skuse), 1889: 304 (Ceratopogon). Australia (New South Wales). 

marshi Wirth and Blanton, 1956b: 220. Panama. 

martinezi Wirth and Blanton, 1970b: 43. Trinidad and Tobago. 

martyrius Trindade and Felippe-Bauer, 2011a:62. Brazil. 

maruim Lutz, 1913: 48. Brazil. 

recifei Barbosa, 1947: 25. Brazil. 

mathisi Giles and Wirth, 1983: 36. Sri Lanka. 

matsuzawai Tokunaga, 1950: 64. Japan. 

mayeri Goetghebuer, 1935a: 3. Germany. 

mcdonaldi Wirth and Hubert, 1989: 224. New name for monticola. 

monticola McDonald and Lu, 1972: 413 (as subspecies of pulicaris). Taiwan. 

mcdowelli Delfinado, 1961: 647. Philippines. 

mckeeveri Brickle and Hagan, 1999: 39. Belize. 

mcmillani Lee and Reye, 1953: 391. Australia (New South Wales). 

megacanthus Palmer, 1957: 272. USA (California). Miocene. 

meijereri Kieffer, 1919c: 192. New name for guttipennis. 

guttipennis (de Meijere), 1906: 69 (Ceratopogon, preoccupied by Culicoides guttipennis (Coquillett), 1901a). 

Indonesia. 

melanesiae Macfie, 1939b: 368. Papua New Guinea. 

melleus (Coquillett), 1901a: 604 (Ceratopogon). USA (Florida). 

mellipes Wirth and Hubert, 1989: 179. Malaysia. 

menghaiensis Lee, 1980: 87. China. 

menglaensis Chu and Liu, 1978: 88. China. 

mesghalii Navai, 1973: 196. Iran. 

90

metagonatus Wirth and Blanton, 1956b: 221. Panama. 

micayensis Spinelli, Santamaría, Cabrera, Ronderos and Suárez, 2009: 85. Colombia. 

micheli Cornet and Chateau, 1971: 164. Senegal. 

micromaculatus Vimmer, 1932: 139. Israel. 

midorensis Arnaud, 1956a: 113. Japan. 

miharai Kinoshita, 1918: 156. Korea. 

mihunensis Chu, 1983: 29. China. 

mikros Dyce and Meiswinkel, 1995: 136. Papua New Guinea. 

milnei Austen, 1909: 283. Kenya. 

lugens Kieffer, 1918a: 51. South Africa. 

minasensis Felippe-Bauer, 1987: 147. Brazil. 

minimaporus Liu and Yu, in Yu et al., 2005: 890. China. 

minimus Wirth and Hubert, 1989: 251. Malaysia. 

minipalpis Wirth and Hubert, 1989: 351. Malaysia. 

minutissimus (Zetterstedt), 1855: 4860 (Ceratopogon). Sweden. 

nanulus Kieffer, 1919a: 38. Hungary. 

albihalter Kieffer, 1919a: 37 (1921f: 785). Germany. 

bychowskyi Dzhafarov, 1964: 344. Azerbaijan. 

tugaicus Dzhafarov, 1960a: 1183. Azerbaijan. 

minutunculus Yu and Liu, in Yu et al., 2005: 1312. China. 

mioceneus Pierce, 1966: 93. USA (California). Miocene. 

miombo Meiswinkel, 1991: 161. Malawi. 

mirsae Ortiz, 1953a: 801. Venezuela. 

mirzaevae Glukhova and Khabirov, 1977: 50 (as mirzaevi). Tajikistan. 

mississippiensis Hoffman, 1926a: 158. USA (Mississippi). 

mohave Wirth, 1952a: 187. USA (California). 

mojingaensis Wirth and Blanton, 1953b: 232. Panama. 

molestior Kieffer, 1911e: 514. New name for molestus. 

molestus Kieffer, 1910: 192 (preoccupied by Culicoides molestus (Skuse), 1889). India. 

iniquus Sen and Das Gupta, 1958b: 163 (1959a: 620; 1959b: 65). New name for molestus. 

molestus (Skuse), 1889: 305 (Ceratopogon). Australia (New South Wales). 

mollis Edwards, 1928: 55. Western Samoa. 

molotovae Glukhova and Braverman, 1999: 309. Turkmenistan. 

mongolensis Yao, 1964: 287. China. 

transcaspius Molotova, 1966: 654. Turkmenistan. 

monicae Spinelli and Borkent, 2004a: 372. Costa Rica. 

monocylichnus Yu and Liu, in Yu et al., 2005: 851. China. 

monoensis Wirth, 1952a: 193. USA (California). 

monothecalis Tokunaga, 1962b: 509. Papua New Guinea. 

montanus Shakirzjanova, 1962: 258. Kazakhstan. 

monticola Wirth and Lee, 1967: 15. Colombia. 

pichindensis Browne, 1980: 538. Colombia. 

moreensis Lee and Reye, 1955: 239. Australia (New South Wales). 

moreli Clastrier, 1959a: 189. Senegal and Ivory Coast. 

morisitai Tokunaga, 1940d: 149. China. 

mihensis Arnaud, 1956a: 115. Japan. 

nagahanai Tokunaga, 1956: 119. Japan. 

mortivallis Wirth and Blanton, 1971b: 465. USA (California). 

motoensis Lee, 1978: 75 (1979a: 102). China. 

moucheti Cornet and Kremer, 1970: 266. Chad. 

mukerjii Majumdar and Das Gupta, in Gangopadhyay and Das Gupta, 2000: 125. India. 

mulrennani Beck, 1957: 103. USA (Florida). 

multifarious Liu, Gong and Zhang in Liu, Gong, Zhang, and Shi 2003: 359. China. 

multifidous Liu and Yu, in Yu et al., 2005: 1241. China. 

91

multimaculatus Taylor, 1918: 169. Australia (Victoria). 

multipunctatus Malloch, 1915a: 296. USA (Illinois). 

murphyi Clastrier and Wirth, 1961b: 303. Nigeria. 

murrayi Wirth and Hubert, 1989: 388. Indonesia. 

murtalai Boorman and Dipeolu, 1979: 41. Nigeria. 

musajevi Dzhafarov, 1961c: 75. Azerbaijan. 

muscicola Kieffer, 1925d: 85. France. 

mykytowyczi Lee and Reye, 1963: 359. Australia (New South Wales). 

mystacinus Vimmer, 1932: 139. Israel. 

nagarzensis Lee, 1978: 77 (1979a: 98). China. 

nairobiensis Glick, 1990: 131. Kenya. 

nampui Howarth, 1985: 28. Laos. 

nanellus Wirth and Blanton, 1969a: 564. USA (California). 

nankanensis Lien, Weng and Lin, 1996: 21. Taiwan. 

nanpingensis Yu and Song, in Yu et al., 1986: 209. China. 

nanus Root and Hoffman, 1937: 165. USA (Maryland). 

narrabeenensis Lee and Reye, 1963: 357. Australia (New South Wales). 

nasuensis Kitaoka, 1984: 303. Japan. 

nattaiensis Lee and Reye, 1955: 237. Australia (New South Wales). 

navaiae Lane, 1983: 534. Saudi Arabia. 

neavei Austen, 1912: 102. Uganda. 

neghmei Vargas, 1955: 673. Mexico. 

neofagineus Wirth and Blanton, 1969b: 227. USA (Arizona). 

neomelanesiae Tokunaga, 1963b: 123. Papua New Guinea. 

neomontanus Wirth, 1976c: 15. New name for montanus. 

montanus Wirth and Blanton, 1969b: 225 (preoccupied by Culicoides montanus Shakirzjanova, 1962). USA (Utah). 

neopalpalis Tokunaga, 1962b: 483. Papua New Guinea. 

neopalpifer Chen, 1983: 21. China. 

neoparaensis Tavares and Souza, 1978: 621. Brazil. 

neopulicaris Wirth, 1955: 355. USA (Texas). 

neoregalis Majumdar and Das Gupta, in Majumdar et al., 1997: 29. India. 

neoschultzei Boorman and Meiswinkel, in Boorman, 1989: 192. Oman. 

nevilli Cornet and Brunhes, 1994: 158. Senegal. 

newsteadi Austen, 1921: 113. Israel. 

biclavatus Kieffer, 1924a: 14. Germany. 

halophilus Kieffer, 1924b: 404. Norway. 

edwardsi Goetghebuer, 1933f: 46 (as variety of pulicaris, preoccupied by Culicoides edwardsi Goetghebuer, 1921). 

Europe. 

edwardsianus Goetghebuer, 1933d: 367 (as variety of pulicaris). Great Britain. 

nibleyi Hubert and Wirth, 1961: 237. Japan. 

nielamensis Liu and Deng, 2000: 246. China. 

nigeriae Ingram and Macfie, 1921: 325. Nigeria. 

nigrifemur Spinelli, Santamaría, Cabrera, Ronderos and Suárez, 2009: 85. Colombia. 

nigrigenus Wirth and Blanton, 1956b: 222. Panama. 

nigripennis Carter, Ingram and Macfie, 1920: 253. Ghana. 

nigripes Wirth and Hubert, 1989: 353. Thailand. 

nigritus Fei and Lee, 1984b: 345. China. 

nigroannulatus Goetghebuer, 1932b: 5. Indonesia. 

nigrosignatus Kieffer, 1901a: 161. France. 

nilogenus Kieffer, 1921b: 17. Sudan. 

nilophilus Kieffer, 1921b: 20. Sudan. 

niphanae Wirth and Hubert, 1989: 316. Thailand. 

nipponensis Tokunaga, 1955: 4. Japan. 

nitens Edwards, 1933c: 252. Malaysia. 

92

nivosus de Meillon, 1937b: 341. South Africa. 

nobrei Caeiro, 1961: 241. Angola. 

nocivum (Harris), 1841: 405 (Simulium). Eastern North America. 

noshaquensis Tokunaga, 1966b: 285. Afghanistan. 

notatus Delfinado, 1961: 648. Philippines. 

papuae Tokunaga, 1962b: 481. Papua New Guinea. 

novaguineanus Tokunaga, 1959: 223. Indonesia. 

novairelandi Tokunaga, 1962b: 512. Papua New Guinea. 

novamexicanus Atchley, 1967: 1012. USA (New Mexico). 

nubeculosus (Meigen), 1830: 263 (Ceratopogon). Europe. 

puncticollis Goetghebuer, 1912: 205 (preoccupied by Culicoides puncticollis (Becker), 1903). Belgium. 

punctaticollis Goetghebuer, 1920: 56. New name for puncticollis. 

nudipalpis Delfinado, 1961: 655. Philippines. 

nudipennis Kieffer, 1922g: 507. Algeria. 

nujiangensis Liu, 1990: 59. China. 

nukabirensis Wada, 1979: 204. Japan. 

nunomemoguri Kitaoka, 1980: 18. Japan. 

nuntius Cambournac, 1970a: 249. Portugal. 

nupurius Kanasugi and Kitaoka, 2001:227. Japan. 

nyakini Howarth, 1985: 49. Laos. 

nyungnoi Howarth, 1985: 24. Laos. 

obesus Statz, 1944: 145. Germany. Oligocene. 

obnoxius Fox, 1952: 365. Venezuela. 

obscuratus Statz, 1944: 144. Germany. Oligocene. 

obscurus Tokunaga and Murachi, 1959: 347. Belau (USA). 

pungens de Meijere, 1909: 200 (preoccupied by Culicoides pungens (Kieffer), 1901a). Indonesia. 

obsoletus (Meigen), 1818: 76 (Ceratopogon). Europe. 

varius (Winnertz), 1852: 35 (Ceratopogon). Germany. 

yezoensis (Matsumura), 1911: 60 (Ceratopogon). Russia. 

obscuripes Santos Abreu, 1918: 297 (as variety of varius). Canary Islands (Spain). 

lacteinervis Kieffer, 1919a: 47. Slovak Republic, Ukraine. 

rivicola Kieffer, 1921a: 56. Germany. 

clavatus Kieffer, 1921a: 56. Germany. 

heterocerus Kieffer, 1921a: 57. Germany. 

pegobius Kieffer, 1922c: 235. Germany. 

kabyliensis Kieffer, 1922g: 505. Algeria. 

concitus Kieffer, 1922e: 71. Germany. 

intermedius Okada, 1941: 22 (as variety of obsoletus, preoccupied by Dasyhelea intermedia (Santos Abreu), 1918). 

Japan. 

sintrensis Cambournac, 1956: 591. Portugal. 

seimi Shevchenko, 1967: 173. Ukraine. 

obuncus Borkent, 1995: 64. Canada (Manitoba). Upper Cretaceous. 

occidentalis Wirth and Jones, 1957: 21 (as subspecies of variipennis). USA (California). 

ochrothorax Carter, 1919: 298. Ghana. 

octosignatus Kieffer, 1921b: 17. Democratic Republic of the Congo. 

oculatus (Strobl), 1910: 262 (Ceratopogon). Austria. 

ocumarensis Ortiz, 1950b: 455. Venezuela. 

odai Boorman, 1989: 195. Oman. 

odiatus Austen, 1921: 112. Israel. 

niger Dzhafarov, 1960a: 1183 (as variety of pallidicornis, preoccupied by Culicoides niger Root and Hoffman, 1937). 

Azerbaijan. 

indistinctus Khalaf, 1961: 461. Iraq. 

lailae Khalaf, 1961: 458. Iraq. 

kurektshaicus Dzhafarov, 1962a: 209. Azerbaijan. 

93

conicus Remm, in Remm and Zhogolev, 1968: 839. Ukraine. 

odiosus Kieffer, 1910: 192. India. 

okazawai Wada, 1990: 60. Japan. 

okinawensis Arnaud, 1956a: 118. Japan. 

oklahomensis Khalaf, 1952b: 355 (as subspecies of villosipennis). USA (Oklahoma). 

olyslageri Kremer and Nevill, 1972: 467. South Africa. 

omogensis Arnaud, 1956a: 119. Japan. 

onderstepoortensis Fiedler, 1951: 6. South Africa. 

onoi Tokunaga, 1940d: 147. China. 

opertus Liu and Yu, 1990: 15. China. 

oregonensis Wirth and Rowley, 1971: 162. USA (Oregon). 

orescius Yu and Liu, in Yu et al., 2005: 1170. China. 

orestes Wirth and Hubert, 1989: 222. Malaysia. 

orientalis Macfie, 1932b: 490. Malaysia. 

nayabazari Das Gupta, 1963: 35. India. 

orjuelai Wirth and Lee, 1967: 6. Colombia. 

ornatus Taylor, 1913: 73. Australia (Queensland). 

ostroushkoae Glukhova, 1989: 220. Ukraine. 

ousairani Khalaf, 1952b: 354. USA (Oklahoma). 

ovalis Khamala and Kettle, 1971: 88. Kenya. 

owyheensis Jones and Wirth, 1978: 57 (misspelled as owyheenis). USA (Idaho). 

oxystoma Kieffer, 1910: 193. India. 

kiefferi Patton, 1913: 336 (preoccupied by Culicoides kiefferi Goetghebuer, 1910). India. 

mesopotamiensis Patton, 1920: 246. Iraq. 

pattoni Kieffer, 1921e: 7. India. New name for kiefferi. 

housei Causey, 1938: 407. Thailand. 

punctigerus Tokunaga, 1951: 101. Indonesia. 

alatus Das Gupta and Ghosh, 1956b: 162. India. 

pabloi Browne, 1980: 541. Colombia. 

pachymerus Lutz, 1914: 83. Brazil. 

padusae Mirzaeva, 1989: 87 (1990: 90). Russia. 

pakistanensis Kitaoka and Shinonaga, 1989: 30. Pakistan. 

paksongi Howarth, 1985: 25. Laos. 

palauensis Tokunaga, in Tokunaga and Murachi, 1959: 348. Belau (USA). 

palawanensis Delfinado, 1961: 648. Philippines. 

pallidicornis Kieffer, 1919a: 46. Hungary, Romania. 

susae Kieffer, 1919a: 44. Italy. 

dileucus Kieffer, 1921a: 55. France. 

brunneiscutellatus Zilahi-Sebess, 1933: 151 (as variety of pallidicornis). Hungary. 

bruneoscutellatus Zilahi-Sebess, 1934: 155 (as variety of pallidicornis). Bulgaria. 

niger Root and Hoffman, 1937: 168. USA (Maryland). 

pallidimaculosus Tokunaga, 1959: 219. Papua New Guinea. 

pallidizonatus Tokunaga, 1963b: 135. Papua New Guinea. 

pallidothorax, Lee and Reye, 1963: 362. Australia (Northern Territory). 

pallidulus Yu, 1991: 52. New name for obscuratus. 

obscuratus Ding and Yu, 1990: 54 (preoccupied by Culicoides obscuratus Statz, 1944). China. 

pallidus Khalaf, 1957b: 338. Iraq. 

stackelbergi Dzhafarov, 1962a: 210. Azerbaijan. 

palmerae James, 1943: 151. USA (Colorado). 

palpalis Macfie, 1948: 78. Mexico. 

palpifer Das Gupta and Ghosh, 1956a: 122. India. 

palpisimilis Wirth and Hubert, 1989: 390. Malaysia. 

pamiricus Zhogolev, 1973: 185. Tajikistan. 

pampaensis Spinelli and Wirth, 1984a: 182. Argentina. 

94

pampangensis Delfinado, 1961: 650. Philippines. 

pampoikilus Macfie, 1948: 79. Mexico. 

dominicii Ortiz, 1951b: 7. Venezuela. 

panamensis Barbosa, 1947: 22. Panama. 

alambiculorum Macfie, 1948: 81. Mexico. 

pancensis Browne, 1980: 537. Colombia. 

pangkorensis Wirth and Hubert, 1989: 326. Malaysia. 

paolae Boorman, in Boorman et al., 1996: 501. Italy. 

papillatus Khamala and Kettle, 1971: 84. Kenya. 

papilliger Borkent, in Borkent and Wirth, 1997:78. New name for papillatus. 

papillatus Kitaoka and Shinonaga, 1989: 26 (preoccupied by Culicoides papillatus Khamala and Kettle, 1971). 

Pakistan. 

papuensis Tokunaga, 1962b: 513. Papua New Guinea. 

parabarnetti Wirth and Hubert, 1989: 146. Malaysia. 

parabubalus Wirth and Hubert, 1989: 224. Malaysia. 

paradisionensis Boorman, 1988: 156. Greece. 

paraensis (Goeldi), 1905: 137 (Haematomyidium). Brazil. 

undecimpunctatus Kieffer, 1917b: 307. Argentina. 

paraflavescens Wirth and Hubert, 1959: 15. Sri Lanka. 

paragarciai Dyce, 1996: 313. Papua New Guinea. 

parahumeralis Wirth and Hubert, 1989: 110. Malaysia. 

paraignacioi Spinelli, in Spinelli et al., 1993: 66. Colombia. 

paraimpunctatus Borkent, 1995: 60. New name for canadensis. 

canadensis Wirth and Blanton, 1969b: 211 (preoccupied by Culicoides canadensis (Boesel), 1937). USA 

(Minnesota). 

paraliui Das Gupta, 1962a: 538. India. 

paramalayae Wirth and Hubert, 1989: 226. Malaysia. 

paramaruim Wirth and Blanton, 1973: 443. Brazil. 

parapiliferus Wirth and Blanton, 1974b: 71. USA (New York). 

parararipalpis Das Gupta, 1963: 41. India. 

pararegalis Majumdar and Das Gupta, in Majumdar et al., 1997: 28. India. 

parascopus Wirth and Blanton, 1978: 238. Mexico. 

parauapebensis Trindade and Felippe-Bauer, 2011b:42. Brazil. 

parroti Kieffer, 1922g: 502. Algeria. 

parvimaculatus Lee and Reye, 1953: 391. Australia (New South Wales). 

parviscriptus Tokunaga, 1959: 213. Papua New Guinea. 

parvulus Khamala and Kettle, 1971: 62. Kenya. 

pastus Kitaoka, 1980: 11. Japan. 

patagoniensis Ronderos and Spinelli, 1997: 34. Argentina. 

paucienfuscatus Barbosa, 1947: 23. Brazil. 

paulipictus Tokunaga, 1977: 45. Papua New Guinea. 

pechumani Cochrane, 1974: 133. USA (New York). 

pecosensis Wirth, 1955: 358. USA (Texas). 

peculiaris Majumdar and Das Gupta, in Gangopadhyay and Das Gupta, 2000: 127. India. 

pelius Liu and Yu, 1990: 23. China. 

peliliouensis Tokunaga, in Tokunaga and Esaki, 1936: 55. Belau (USA). 

ejercitoi Delfinado, 1961: 643. Philippines. 

pellucidus Khamala and Kettle, 1971: 62. Tanzania. 

pendleburyi Wirth and Hubert, 1989: 85. Malaysia. 

pentamaculatus Smatov, in Smatov and Kravets, 1976: 283. Kazakhstan. 

perakensis Kitaoka, 1983: 95. Malaysia. 

peregrinus Kieffer, 1910: 191. India. 

judicandus Bezzi, 1916: 8 (1917: 108). Philippines. 

esmoneti Salm, 1917b: 136. Indonesia. 

95

philippinensis Kieffer, 1921g: 564. Philippines. 

assamensis Smith and Swaminath, 1932: 183 (as variety of peregrinus). India. 

quadratus Tokunaga, 1951: 108. Indonesia. 

perettii Cornet and Chateau, 1971: 145. Senegal. 

perijaensis Perruolo, 2006b: 27. Venezuela. 

perornatus Delfinado, 1961: 651. Philippines. 

multinotatae Tokunaga, 1962b: 475. Papua New Guinea. 

peruvianus Felippe-Bauer, in Felippe-Bauer, et al., 2003: 1054. Peru. 

petersi Tokunaga, 1962b: 490. Papua New Guinea. 

petronius Liu and Yu, in Yu et al., 2005: 1091. China. 

phaeonotus Wirth and Blanton, 1959: 326. Panama. 

phasmatua Liu and Yu, 1997: 120. China. 

phlebotomus (Williston), 1896: 281 (Ceratopogon). St. Vincent. 

amazonius Macfie, 1935a: 52. Brazil. 

photophilus Kieffer, 1911d: 5. Germany. 

picadoae Spinelli and Borkent, 2004a: 381. Costa Rica. 

pictellum (Rondani), 1869: 190 (Ceratopogon). Italy. 

pictilis Wirth and Hubert, 1989: 392. Malaysia. 

pictimargo Tokunaga and Shogaki, 1953: 286. Japan. 

pictipennis (Staeger), 1839: 594 (Ceratopogon). Denmark. 

arcuatus (Winnertz), 1852: 39 (Ceratopogon). Germany. 

guttularis Kieffer, 1919a: 45. Hungary. 

maculatus Zilahi-Sebess, 1936a: 202 (as variety of pictipennis, preoccupied by Culicoides maculatus (Shiraki), 

1913). Hungary. 

achkamalicus Dzhafarov, 1964: 251. Azerbaijan. 

luganicus Shevchenko, 1972: 75. Ukraine. 

picturatus Kremer and Deduit, 1961: 701. France. 

pifanoi Ortiz, 1951c: 588. Venezuela. 

tricoloratus Wirth and Blanton, 1953b: 233. Panama. 

pikongkoi Howarth, 1985: 51. Laos. 

piliferus Root and Hoffman, 1937: 163. USA (Maryland). 

pilosipennis Kieffer, 1925e: 257. Egypt. 

pilosus Wirth and Blanton, 1959: 332. Panama. 

pingtanensis Yu, 1982: 202. China. 

platiradius Tokunaga, 1963b: 128. Papua New Guinea. 

plaumanni Spinelli, in Spinelli et al., 1993: 69. Argentina. 

poikilonotus Macfie, 1948: 82. Mexico. 

cacozelus Macfie, 1948: 85. Mexico. 

hertigi Wirth and Blanton, 1953b: 229. Panama. 

polynesiae Wirth and Arnaud, 1969: 518. American Samoa (USA). 

polypori Wirth and Blanton, 1968a: 212. Panama. 

polystictus Kieffer, 1921c: 181. Paraguay. 

pongsomiensis Chu, 1986: 254. Cambodia. 

popayanensis Wirth and Lee, 1967: 10. Colombia. 

poperinghensis Goetghebuer, 1953: 127. Belgium. 

posoensis Wirth and Blanton, 1969a: 562. USA (California). 

pretoriensis Kremer and Nevill, 1972: 464. South Africa. 

prolixipalpis Wirth and Hubert, 1989: 426. Malaysia. 

propinquus Macfie, 1948: 81. Mexico. 

propriipennis Macfie, 1948: 84. Mexico. 

prussicus Szadziewski, 1988: 48. Baltic region. Eocene. 

pseudocrescentis Tavares and Luna Dias, 1980: 397. Brazil. 

pseudocordiger Wirth and Hubert, 1989: 447. Malaysia. 

pseudodiabolicus Fox, 1946a: 256. Trinidad and Tobago. 

96

pseudoheliconiae Felippe-Bauer, in Felippe-Bauer et al., 2008a: 260. Peru. 

pseudoheliophilus Callot and Kremer, 1961a: 682. France. 

albihalteratus Goetghebuer, 1935b: 414 (as variety of neglectus, preoccupied by Dasyhelea albohalteratus (Santos 

Abreu), 1918). Belgium. 

pseudolangeroni Kremer, Chaker and Delécolle, 1981: 291. Tunisia. 

pseudopallidipennis Clastrier, 1958b: 197. Senegal. 

pseudopallidus Khalaf, 1961: 466. Iraq. 

pseudopalpalis Wirth and Hubert, 1989: 394. Indonesia. 

pseudoregalis Majumdar and Das Gupta, in Majumdar et al., 1997: 30. India. 

pseudosalinarius Chu, 1981: 309. China. 

pseudostigmatus Tokunaga, 1959: 234. Indonesia. 

pulchellus Liu and Zhao, 1998: 12. China. 

pulchripennis Macfie, 1939c: 200. Brazil. 

pulicaris (Linnaeus), 1758: 603 (Culex). Europe. 

setosinervis Kieffer, 1913a: 8. Germany. 

pullatus Kieffer, 1915a: 474. Germany. 

stephensi Carter, 1916: 135. Egypt. 

cinerellus Kieffer, 1919a: 40. Ukraine. 

quinquepunctatus Goetghebuer, 1921: 177. Belgium. 

flaviplumus Kieffer, 1924a: 19. Germany. 

sawamotoi Kono and Takahasi, 1940: 75. Russia. 

pumilus (Winnertz), 1852: 46 (Ceratopogon). Germany. 

punctatus (Meigen), 1804: 29 (Ceratopogon). Europe. 

punctatus Latreille, 1809: 252 (preoccupied by Culicoides punctatus (Meigen), 1804). France. 

ocellaris Kieffer, 1921d: 276 (as variety of pulicaris). Latvia. 

kasachstanicus Shakirzjanova, 1963: 63 (as subspecies of pulicaris). Kazakhstan. 

puncticollis (Becker), 1903: 75 (Ceratopogon). Egypt. 

algecirensis (Strobl), 1900: 170 (Ceratopogon, as form of pulicaris). Spain. Name suppressed by ICZN Opinion 

1643. 

impressus Kieffer, 1918a: 47. Tunisia. 

distigma Kieffer, 1922g: 502. Algeria. 

donatieni Kieffer, 1922g: 504. Algeria. 

sciniphes Kieffer, 1925e: 261. Egypt. 

bipunctatus Vimmer, 1932: 133. Israel. 

tripunctatus Vimmer, 1932: 137. Israel. 

wenigi Vimmer, 1932: 138. Israel. 

flavitarsis Vimmer, 1932: 137. Israel. 

griseovittatus Vimmer, 1932: 133. Israel. 

luteosignatus Vimmer, 1932: 140. Israel. 

vavrai Vimmer, 1932: 140. Israel. 

punctithorax Carter, Ingram and Macfie, 1920: 235. Ghana. 

pungobovis Liu, Yan and Liu, 1996: 35. China. 

puracensis Wirth and Lee, 1967: 7. Colombia. 

puripennis Austen, 1921: 109. Israel. 

purus Lee and Reye, 1963: 352. Australia (New South Wales). 

pusilloides Wirth and Blanton, 1955a: 104. Panama. 

pusillus Lutz, 1913: 52. Brazil. 

putianensis Lee, 1988: 96. China. 

pycnostictus Ingram and Macfie, 1925: 284. Malawi. 

alexis de Meillon, 1936: 147. South Africa. 

meeserellus de Meillon, 1936: 151. South Africa. 

hysipyles de Meillon, 1936: 156. South Africa. 

pygmaeus Tokunaga, 1963b: 139. Indonesia. 

qabdoensis Lee, 1979a: 103. China. 

97

qianshanensis Fei, 1982: 105. China. 

qianweiensis Yu, 1982: 202. China. 

qingdaoensis Kong and Yu, 1990: 58. China. 

qinghaiensis Fei and Lee, 1984a: 182. China. 

qionghaiensis Yu and Liu, 1990: 4. China. 

qiongzhongensis Liu, Yan and Liu, 1996: 35. China. 

quadrisignatus Kieffer, 1921b: 16. Cameroon. 

quadrivittatus Vimmer, 1932: 136. Israel. 

quasiparaensis Clastrier, 1971: 286. French Guiana. 

quasiregalis Majumdar and Das Gupta, in Majumdar et al., 1997: 30. India. 

quatei Wirth and Hubert, 1989: 333. Malaysia. 

quaterifasciatus Tokunaga, 1959: 240. Papua New Guinea. 

queenslandae Dyce and Wirth, 1997: 589. Australia (Queensland). 

qufuensis Xue and Yu, in Xue et al., 1992: 52. China. 

quinquelineatus Goetghebuer, 1934d: 192. Democratic Republic of the Congo. 

quinquemaculatus Vimmer, 1932: 132. Israel. 

quqiaoensis Chen, in Lee, 1988: 99. China. 

rabauli Macfie, 1939b: 367. Papua New Guinea. 

angularis Lee and Reye, 1953: 384. Australia (New South Wales). 

rachoui Tavares and Souza, 1978: 622. Brazil. 

radiomaculatus Khamala and Kettle, 1971: 85. Kenya. 

rageaui Vattier and Adam, 1966b: 727. Congo. 

rangeli Ortiz and Mirsa, 1952b: 126. Venezuela. 

donajii Vargas, 1954: 28. Mexico. 

patulipalpis Wirth and Blanton, 1959: 421. Panama. 

raposoensis Wirth and Barreto, 1978: 561. Colombia. 

raripalpis Smith, 1929: 256 (1932: 181). India. 

rariradialis Das Gupta, 1963: 34. India. 

rarus Das Gupta, 1963: 40. India. 

ravus de Meillon, 1936: 151. South Africa. 

tokwensis de Meillon, 1942a: 97. Zimbabwe. 

fuscicaudae Macfie, 1947b: 75. Sudan. 

subravus Cornet and Chateau, 1971: 167. Senegal. 

reconditus Campbell and Pelham-Clinton, 1960: 227. Great Britain. 

recurvus Delfinado, 1961: 663. Philippines. 

reevesi Wirth, 1952a: 193. USA (California). 

regalis Majumdar and Das Gupta, in Majumdar et al., 1997: 31. India. 

remerki Boorman and Dipeolu, 1979: 51. Nigeria. 

remotus Kieffer, 1918a: 54. Guinea. 

reticulatus Lutz, 1913: 49. Brazil. 

rhizophorensis Khamala and Kettle, 1971: 77. Kenya. 

ribeiroi Lemblé, Messaddeq, Capela and Kremer, 1990: 267. Portugal. 

riebi Delécolle, Mathieu, and Baldet, 2005: 70. France. 

riethi Kieffer, 1914a: 237. Germany. 

cordatus Kieffer, 1921h: 114 (1921d: 275). Latvia. 

crassiforceps Kieffer, 1924a: 15. Germany. 

riggsi Khalaf, 1957a: 198 (as subspecies of piliferus). USA (Oklahoma). 

riouxi Callot and Kremer, 1961a: 679. France. 

ritzei Dzhafarov, 1964: 346. Georgia. 

robini Cornet, 1970: 352. Senegal. 

rochenus Cambournac, 1970a: 250. Portugal. 

rodriguezi Ortiz, 1968: 67. Venezuela. 

ronderosae Spinelli and Borkent, 2004a: 377. Costa Rica. 

rostratus Wirth and Blanton, 1956b: 218. Panama. 

98

ubzovi Dzhafarov, 1960a: 1181. Azerbaijan. 

rugulithecus Wirth and Hubert, 1989: 113. Malaysia. 

ruiliensis Lee, 1980: 86. China. 

ruizi Forattini, 1954a: 189. Brazil. 

rutilus Ingram and Macfie, 1921: 326 (as variety of inornatipennis). Ghana. 

rutshuruensis Goetghebuer, 1935d: 174. Democratic Republic of the Congo. 

ryckmani Wirth and Hubert, 1960a: 656. USA (California). 

saboyae Cornet, 1970: 355. Senegal. 


sacrilegus Xue and Yu, 1991: 53. China. 

saevanicus Dzhafarov, 1960b: 94. Armenia. 

saevus Kieffer, 1922g: 506. Algeria. 

drenskii Zilahi-Sebess, 1934: 154. Bulgaria. 

puncticeps Goetghebuer, 1934a: 91. Austria. 

micromaculithorax Khalaf, 1957b: 340. Iraq. 


baghdadensis Khalaf, 1957b: 341 (as subspecies of similis). Iraq. 

coluzzii Callot, Kremer and Bailly-Choumara, 1970: 710. Tunisia. 

sajanicus Mirzaeva, 1971: 35. Russia. 

salebrosus Liu and Shi in Liu, Gong, Zhang, and Shi 2003: 360. China. 

salihi Khalaf, 1952b: 351. USA (Oklahoma). 

salinarius Kieffer, 1914a: 236. Germany. 

halobius Kieffer, 1914a: 237. Germany. 

meinerti Kieffer, 1915c: 284. Denmark. 

punctatidorsum Kieffer, 1924a: 17. Germany. 

saltaensis Spinelli and Wirth, 1984a: 183. Argentina. 

saltonensis Wirth, 1952a: 173 (as subspecies of cockerellii). USA (California). 

samoensis Wirth and Arnaud, 1969: 513. American Samoa (USA). 

sanguisuga (Coquillett), 1901a: 604 (Ceratopogon). USA (Maryland). 

saninensis Tokunaga, 1956: 116. Japan. 

epactius Arnaud, 1956a: 102. Japan. 

litus Arnaud, 1956a: 111. Japan. 

santanderi Browne, 1980: 536. Colombia. 

santonicus Callot, Kremer, Rault and Bach, 1966: 514. France. 

sarawakensis Wirth and Hubert, 1959: 28. Malaysia. 

sasai Kitaoka, 1975: 192. Japan. 

saundersi Wirth and Blanton, 1969a: 557. USA (Washington). 

scanloni Wirth and Hubert, 1962: 187. USA (Virginia). 

schramae Giles, Wirth and Messersmith, 1981: 539. Sri Lanka. 

roswelli Giles and Wirth, 1983: 38. Sri Lanka. 

schultzei (Enderlein), 1908: 459 (Ceratopogon). Namibia. 

irroratus Goetghebuer, 1948b: 12. Democratic Republic of the Congo. 

scopus Root and Hoffman, 1937: 170. Mexico. 

scoticus Downes and Kettle, 1952: 65. Great Britain. 

segnis Campbell and Pelham-Clinton, 1960: 232. Great Britain. 

sejfadinei Dzhafarov, 1958: 247. Azerbaijan. 

flavidus Dzhafarov, 1959: 470. Azerbaijan. 

selangorensis Wirth and Hubert, 1989: 355. Malaysia. 

sellersi Boorman and Dipeolu, 1979: 55. Nigeria. 

semicircum Tokunaga, 1959: 208. Indonesia. 

semimaculatus Clastrier, 1958a: 55. Algeria. 

karajevi Dzhafarov, 1961c: 77. Azerbaijan. 

sensillatus Mirzaeva, 1971: 33. Russia. 

kuraiensis Remm, 1972: 82. Russia. 

99

sergenti Kieffer, 1921h: 113. Algeria. 

citrinellus Kieffer, 1923a: 674. Algeria. 

mosulensis Khalaf, 1957b: 339. Iraq. 

turkmenicus Gutsevich, 1959: 678. Turkmenistan. 

shahgudiani Navai, 1973: 201. Iran. 

shaklawensis Khalaf, 1957b: 345. Iraq. 

caspius Gutsevich, 1959: 676. Russia. 

shamaensis Yu and Deng, in Zhang et al., 1990: 31. China. 

shanghaiensis Qu, Cao and Liu, 2010: 322. China. 

shermani Causey, 1938: 404. Thailand. 

shimoniensis Khamala and Kettle, 1971: 54. Kenya. 

shortti Smith and Swaminath, 1932: 183. India. 

fortis Sen and Das Gupta, 1959a: 622. India. 

siamensis Wirth and Hubert, 1989: 359. Thailand. 

sibiricus Mirzaeva, 1964: 218. Russia. 

sierrensis Wirth and Blanton, 1969b: 232. USA (California). 

sigmoidus Lee and Reye, 1963: 359. Australia (Capital Territory). 

signatus Kieffer, 1921b: 19. Sudan. 

sikkimensis Das Gupta, 1963: 35. India. 

silvestrii Kieffer, 1918a: 50. Cameroon. 

similis Carter, Ingram and Macfie, 1920: 255. Ghana. 

simulans Vimmer, 1932: 136. Israel. 

simulator Edwards, in Edwards et al., 1939: 40. Great Britain. 

sinanoensis Tokunaga, 1937a: 331. Japan. 

obsoletiformis Amosova, 1957: 233. Russia. 

sitiens Wirth and Hubert, 1960a: 652. USA (California). 

slovacus Országh, 1969: 1. Slovak Republic. 

smeei Tokunaga, 1960b: 73. Papua New Guinea. 

snowi Wirth and Jones, 1956: 163. USA (Virginia). 

sogdianus Gutsevich, 1966: 673. Uzbekistan. 

sommermanae Wirth and Blanton, 1969b: 234. USA (Alaska). 

sonorensis Wirth and Jones, 1957: 18 (as subspecies of variipennis). USA (Arizona). 

albertensis Wirth and Jones, 1957: 17 (as subspecies of variipennis). Canada (Alberta). 

australis Wirth and Jones, 1957: 15 (as subspecies of variipennis). USA (Louisiana). 

sordidellus (Zetterstedt), 1838: 820 (Ceratopogon). Greenland (Denmark). 

sousadiasi Caeiro, 1961: 309. Angola. 

speciosus (Meunier), 1904a: 229 (1904b: 240) (Ceratopogon). Baltic region. Eocene. 

sphagnumensis Williams, 1955: 269. USA (Michigan). 

lacicola Arnaud, 1956a: 110 (as laciocola). Japan. 

carjalaensis Glukhova, 1957: 249. Russia. 

sphenostylus Remm, 1976b: 112. Russia. Upper Cretaceous. 

spiculae Howarth, 1985: 53. Laos. 

spinapenis Yu and Hao, in Hao et al., 1990: 41. China. 

spinosus Root and Hoffman, 1937: 172. USA (Maryland). 

spinoverbosus Qu and Wang, 1994: 488. China. 

spinulosus Khamala and Kettle, 1971: 87. Uganda. 

spurius Wirth and Blanton, 1959: 433. Panama. 

stagetus Lee, 1979b: 33. China. 

stanicicus Shevchenko, 1970a: 84. Ukraine. 

stellaris Yu and Liu, 1990: 6. China. 

stellifer (Coquillett), 1901a: 604 (Ceratopogon). USA (District of Columbia). 

stepicola Remm, in Remm and Zhogolev, 1968: 838. Ukraine. 

markevitshi Shevchenko, 1969: 47. Ukraine. 

aquilinus Smatov and Kravets, 1976: 284. Kazakhstan. 

100

stercorarius Khamala and Kettle, 1971: 60. Uganda. 

stigma (Meigen), 1818: 73 (Ceratopogon). Europe. 

kiefferi Goetghebuer, 1910: 96. Belgium. 

cordiformitarsis Carter, 1916: 134. Egypt. 

unimaculatus Goetghebuer, 1920: 57. New name for kiefferi. 

stigmoides Callot, Kremer and Deduit, 1962: 166. France. 

stigmalis Wirth, 1952d: 245. Guatemala. 

stigmaticus Kieffer, 1911d: 4. Germany. 

stilobezzioides Foote and Pratt, 1954: 33. USA (New York). 

stonei James, 1943: 149. USA (Colorado). 

weesei Khalaf, 1952a: 65 (1952b: 351). USA (Oklahoma). 

stupulosus Yu and Zhang, in Deng et al., 1990: 35. China. 

suarezi Rodriguez and Wirth, 1986: 313. Colombia. 

subarakawae Yu and Zou, in Hao et al., 1990: 41. China. 

subchangbaiensis Liu and Ren, in Liu et al., 2010: 64. China. 

subcircumscriptus Yu, 1982: 202. China. 

subdubius Tokunaga, 1962c: 53. Japan. 

subfagineus Delécolle and Ortega, 1998: 285. Spain. 

subfasciipennis Kieffer, 1919a: 44. Hungary. 

analis Kieffer, 1925d: 81 (as variety of subfasciipennis, preoccupied by Culicoides analis Santos Abreu, 1918). 

Belgium. 

subgedanensis Szadziewski, 1993: 608. Germany. Eocene. 

subgrisescens Dzhafarov, 1964: 249. Azerbaijan. 

subimmaculatus Lee and Reye, 1953: 375. Australia (New South Wales). 

sublatifrontis Smatov and Isimbekov, 1971: 61. Kazakhstan. 

sublettei Atchley, 1967: 997. USA (New Mexico). 

submagnesianus Tokunaga, 1962b: 484. Papua New Guinea. 

subpalpifer Lee, 1988: 104 (as subpalpifer Wirth and Hubert). China. 

subpalpifer Wirth and Hubert, 1989: 115 (preoccupied by Culicoides subpalpifer Lee, 1988). Malaysia. 

subpunctatus Liu and Yu, 1996: 137 (Liu et al., 1999: 341). China. 

subregalis Majumdar and Das Gupta, in Majumdar et al., 1997: 31. India. 

subschultzei Cornet and Brunhes, 1994: 158. South Africa. 

subsejfadinei Liu, Wang and Hao, 1999: 201. China. 

subsylvarum Remm, 1981: 32. Russia. 

succineus Remm, 1976b: 113. Russia. Upper Cretaceous. 

succivarius Szadziewski, 1988: 40. Poland. Eocene. 

sumatrae Macfie, 1934c: 190 (1934d: 215). Malaysia. 

amamiensis Tokunaga, 1937a: 325. Japan. 

kagiensis Tokunaga, 1937a: 327. Taiwan. 

ohmorii Takahashi, 1958: 113 (as subspecies of amamiensis). Japan. 

assimilis Delfinado, 1961: 660. Philippines. 

superfluthecus Yu and Li, in Yu et al., 1986: 210. China. 

suspectus Zhou and Lee, 1984b: 294. China. 

suzukii Kitaoka, 1973: 212. Japan. 

swaminathi Majumdar and Das Gupta, in Gangopadhyay and Das Gupta, 2000: 128. India. 

sylvicola Khamala and Kettle, 1971: 36. Kenya. 

tadzhikistanicus Zhogolev, 1969: 114. Tajikistan. 

tahemanensis Liu and Ma, 2001: 199. China. 

tainanus Kieffer, 1916b: 114. Taiwan. 

maculatus (Shiraki), 1913: 294 (Ceratopogon, preoccupied by Atrichopogon maculatus (Lundström), 1910). Taiwan. 

kii Tokunaga, 1937a: 284. Japan. 

sigaensis Tokunaga, 1937a: 322. Japan. 

kyotoensis Tokunaga, 1937a: 329. Japan. 

suborientalis Tokunaga, 1951: 106. Indonesia. 

101

taipingshanensis Liu and Guo, 2009:50. China. 

taiwanensis Kitaoka and Tanaka, 1985: 44. Taiwan. 

talgariensis Gutsevich and Smatov, 1966: 74. Kazakhstan. 

tamada Howarth, 1985: 39. Laos. 

tamaensis Perruolo, 2006a: 115. Venezuela. 

tamboensis Wirth and Lee, 1967: 20. Colombia. 

taonanensis Ren, Wang and Liu, 2006: 388. China. 

tarapaca Spinelli and Wirth, 1984a: 184. Chile. 

tatebeae Kitaoka, 1991: 291. Japan. 

tauffliebi Clastrier, 1960a: 96. Congo. 

tauricus Gutsevich, 1959: 677. Ukraine. 

tavaresi Felippe-Bauer and Wirth, 1988: 261. Brazil. 

tawauensis Wirth and Hubert, 1989: 161. Malaysia. 

taylori (Boorman and Lane), 1979: 327 (Neoculicoides). Nigeria. 

tayulingensis Chen, 1983: 77 (1988: 153). China. 

tbilisicus Dzhafarov, 1964: 309. Azerbaijan, Georgia. 

dendriticus Boorman, 1976: 102. Great Britain. 

tentorius Austen, 1921: 110. Israel. 

tenuifasciatus Wirth and Hubert, 1989: 243. Malaysia. 

tenuilobus Wirth and Blanton, 1959: 354. Panama. 

tenuipalpis Wirth and Hubert, 1959: 16. Taiwan. 

tenuipennis Statz, 1944: 147. Germany. Oligocene. 

tenuistylus Wirth, 1952a: 178. USA (California). 

teretipalpis Wirth and Barreto, 1978: 557. Colombia. 

tessellatus Yu, 1982: 202. China. 

testudinalis Wirth and Hubert, 1962: 191. USA (Pennsylvania). 

tetrathyris Wirth and Blanton, 1959: 409. Panama. 

thurmanae Wirth and Hubert, 1989: 360. Thailand. 

tianmushaensis Chu, 1981: 310. China. 

tiaratus Liu and Zhao, in Yu et al., 2005: 1109. China. 

tibetensis Chu, 1977: 102. China. 

tidwelli Spinelli, in Spinelli et al., 1993: 74. Colombia. 

tienhsiangensis Chen, 1983: 78 (1988: 153). China. 

tissoti Wirth and Blanton, 1966: 279. USA (Florida). 

tobaensis Tokunaga, 1937a: 317. Japan. 

todatangae Wirth and Blanton, 1973: 447. Brazil. 

tohokuensis Okada, 1941: 16. Japan. 

tokunagai Arnaud, 1956a: 132. Japan. 

tonmai Howarth, 1985: 30. Laos. 

tororoensis Khamala and Kettle, 1971: 48. Uganda. 

torreyae Wirth and Blanton, 1971c: 73. USA (Florida). 

torridus Wirth and Hubert, 1960a: 654. Mexico. 

toshiokai Kitaoka, 1975: 194. Japan. 

towadaensis Okada, 1941: 25. Japan. 

toyamaruae Arnaud, 1956a: 133. Japan. 

translucens Khamala and Kettle, 1971: 83. Tanzania. 

trapidoi Wirth and Barreto, 1978: 554. Colombia. 

travassosi Forattini, 1957: 198. Brazil. 

travisi Vargas, 1949: 233. New name for simulans. 

simulans Root and Hoffman, 1937: 167 (preoccupied by Culicoides simulans Vimmer, 1932). USA (Maryland). 

horneae Foote and Pratt, 1954: 25. USA (New York). 

triallantionis Howarth, 1985: 42. Laos. 

triangulatus Shevchenko, 1970b: 11. Ukraine. 

trifasciellus Goetghebuer, 1935d: 175. Democratic Republic of the Congo. 

102

trifidus Spinelli and Borkent, 2004a: 385. Costa Rica. 

trilineatus Fox, 1946a: 250. Virgin Islands (USA). 

trimaculatus McDonald and Lu, 1972: 415. Taiwan. 

trimaculipennis Wirth and Hubert, 1989: 245. Malaysia. 

trinidadensis Hoffman, 1925: 286. Trinidad and Tobago. 

oliveri Fox and Hoffman, 1944: 108. Haiti. 

wokei Barbosa, 1947: 28 (preoccupied by Culicoides wokei Fox, 1947). Panama. 

diminutus Barbosa, 1951: 163. New name for wokei. 

tripallidus Tokunaga, 1959: 220. Papua New Guinea. 

trisignatus Kieffer, 1921b: 21. Cameroon. 

tristanii Huttel, Huttel and Verdier, 1953: 100. Gabon. 

tristriatulus Hoffman, 1925: 294 (as variety of cockerellii). USA (California). 

tritenuifasciatus Tokunaga, 1959: 242. Papua New Guinea. 

trivittatus Vimmer, 1932: 136. Israel. 

subneglectus Vimmer, 1932: 139. Israel. 

bulbostylus Khalaf, 1961: 463. Iraq. 

trizonatus Tokunaga, 1963b: 134. Papua New Guinea. 

tropicalis Kieffer, 1913e: 10. Kenya. 

babrius de Meillon, 1943: 112. Zimbabwe. 

trouilleti Itoua and Cornet, 1986: 239. Congo. 

truncatus Borkent, 2000b: 461. USA (New Jersey). Upper Cretaceous. 

truncorum Edwards, in Edwards et al., 1939: 41. Great Britain. 

sylvarum Callot and Kremer, 1961b: 389. France. 

tsutaensis Wada, 1990: 65. Japan. 

tuamsombooni Kitaoka, Takaoka and Choochote, 2005: 290. Thailand. 

tunkinensis Mirzaeva, 1985: 97. Russia. 

turanicus Gutsevich and Smatov, in Smatov and Isimbekov, 1971: 61. New name for kasachstanicus. 

kasachstanicus Gutsevich and Smatov, 1966: 70 (preoccupied by Culicoides kasachstanicus Shakirzjanova, 1963). 

Kazakhstan. 

turgeopalpulus Liu and Yu, 1990: 23. China. 

turgidus Sen and Das Gupta, 1959a: 626. India. 

tuttifrutti Meiswinkel, Cornet and Dyce, in Meiswinkel and Linton, 2003: 42. South Africa. 

tyrrelli (Boesel), 1937: 48 (Dasyhelea). Canada (Manitoba). Upper Cretaceous. 

ukrainensis Shevchenko, 1970b: 14. Ukraine. 

uncistylus Wirth and Hubert, 1989: 449. Philippines. 

undentaris Liu, 2002: 191. China. 

unetensis Perruolo, 2001: 35. Venezuela. 

unicolor (Coquillett), 1905: 65 (Ceratopogon). USA (California). 

unicus Delfinado, 1961: 665. Philippines. 

uniradialis Wirth and Blanton, 1953a: 70. Panama. 

univittatus Vimmer, 1932: 139. Israel 

agathensis Callot, Kremer and Rioux, 1963: 121. France. 

uruguayensis Ronderos, 1990a: 117. Uruguay. 

usingeri Wirth, 1952a: 192. USA (California). 

ustinovi Shevchenko, 1962: 673. Ukraine. 

utahensis Fox, 1946b: 246. USA (Utah). 

utowana Jamnback, 1965: 109. USA (New York). 

variatus Liu, Yan and Liu, 1996: 36. China. 

variifrons Glukhova and Ivanov, in Ivanov and Glukhova, 1967: 810. Russia. 

variipennis (Coquillett), 1901a: 602 (Ceratopogon). USA (Virginia). 

venezuelensis Ortiz and Mirsa, 1950: 137. Venezuela. 

pictipennis (Philippi), 1865: 628 (Psychophaena, preoccupied by Culicoides pictipennis (Staeger), 1839). Chile. 

ortizi Fox, 1952: 366. Venezuela. 

ventralis Borkent, in Borkent and Wirth, 1997:84. New name for abdominalis. 

103

abdominalis Statz, 1944: 145 (preoccupied by Dasyhelea abdominalis (Santos Abreu), 1918). Germany. Oligocene. 

venustus Hoffman, 1925: 290. USA (Maryland). 

verbosus Tokunaga, 1937a: 303. Taiwan. 

verecundus Macfie, 1948: 76. Mexico. 

vernoni Wirth and Blanton, 1973: 448. Brazil. 

vespertinus Yu and Ma, in Ma et al., 1990: 48. China. 

vetustus Breidenbaugh and Mullens, 1999: 156. USA (California). 

vexans (Staeger), 1839: 593 (Ceratopogon). Denmark. 

pungens (Kieffer), 1901a: 163 (Palpomyia). France. 

perpungens Kieffer, 1919a: 33. New name for pungens. 

ajbassovi Shakirzjanova, 1962: 254. Kazakhstan. 

vicinus Clastrier, 1960a: 104. New name for intermedius. 

intermedius Clastrier, 1959a: 173 (preoccupied by Dasyhelea intermedia (Santos Abreu), 1918). Senegal. 

victoriae Macfie, 1941: 67. Australia (Victoria). 

magnimaculatus Lee and Reye, 1953: 388. Australia (New South Wales). 

vidourlensis Callot, Kremer, Molet and Bach, 1968: 99. France. 

villosipennis Root and Hoffman, 1937: 165. USA (Maryland). 

vitreipennis Austen, 1921: 108. Israel. 

vitshumbiensis Goetghebuer, 1935d: 176. Democratic Republic of the Congo. 

volcanensis Wirth and Blanton, 1959: 389. Panama. 

vomensis Boorman and Dipeolu, 1979: 61. Nigeria. 

wadai Kitaoka, 1980: 14. Japan. 

wakuensis McDonald, Bolinguit and Lu, 1973: 647. Japan. 

chaetocellaris Kitaoka, 1973: 214. Japan. 

walkeri Boorman, 1979: 69. Kenya. 

wallacei Wirth and Blanton, 1973: 449. Brazil. 

wandashanensis Wang and Liu, 1999: 328. China. 

wansoni Goetghebuer, 1935c: 477. Democratic Republic of the Congo. 

obscuripennis Clastrier and Wirth, 1961b: 318. Gambia. 

wardi Boorman, 1989: 209. United Arab Emirates. 

waringi Lee and Reye, 1955: 240. Australia (Western Australia). 

wenzeli Delfiando, 1961: 649. Philippines. 

werneri Wirth and Blanton, 1971b: 463. USA (Arizona). 

williamsi Spinelli, in Spinelli et al., 2005: 147. Paraguay. 

willistoni Wirth and Blanton, 1953c: 116. Panama. 

williwilli Lee and Reye, 1955: 234. Australia (New South Wales). 

wirthi Foote and Pratt, 1954: 36. USA (Montana). 

wirthomyia Vargas, 1953b: 227. Mexico. 

wisconsinensis Jones, 1956: 32. USA (Wisconsin). 

wokei Fox, 1947: 90. Panama. 

aethionotus Wirth and Blanton, 1955b: 121. Panama. 

wuyiensis Chen, 1981: 93 (1982: 129). China. 

xanifer Wirth and Blanton, 1968a: 210. Panama. 

xanthoceras Kieffer, 1917a: 186. Papua New Guinea. 

xanthogaster Kieffer, 1918a: 53. Guinea. 

xinghaiensis Yu, 1982: 202. China. 

xinjiangensis Chu, Qian and Ma, 1982: 107. China. 

xinpingensis Qu and Liu, 1982: 101. China. 

yanbianensis Liu, Wu and Wang, 2006:321. China. 

yadongensis Chu, 1984: 24. New name for spinulosus. 

spinulosus Chu, 1977: 100 (preoccupied by Culicoides spinulosus Khamala and Kettle, 1971). China. 

yaeyamaensis Kitaoka, 1975: 196. Japan. 

yankari Boorman and Dipeolu, 1979: 62. Nigeria. 

yamii Lien, Lin and Weng, 1998b:57. Taiwan. 

104

yaracuyensis Ortiz, 1959: 364 (as yaracuyanus). Venezuela. 

yasumatsui Tokunaga, 1941a: 113. Micronesia. 

lingensis Tokunaga, 1963b: 129. Papua New Guinea. 

yemenensis Boorman, 1989: 211. Yemen. 

yichunensis Yu, 1982: 202. China. 

yigongensis Liu and Deng, 2010: 578. China. 

yoosti Borkent, 2000b: 462. USA (New Jersey). Upper Cretaceous. 

yoshimurai Tokunaga, 1941a: 114. Micronesia. 

youngi Wirth and Barreto, 1978: 562. Colombia. 

yuchihensis Lien, Lin and Weng, 1998b:58. Taiwan. 

yukonensis Hoffman, 1925: 291. Canada (Yukon Territory). 

yui Liu and Hao, in Yu et al., 2005: 1117. China. 

yunanensis Chu and Liu, 1978: 81. China. 

zentae Bellis and Dyce, 2011: 44. Australia (Queensland). 

zhangmensis Deng and Yu, in Deng et al., 1990: 35. China. 

zhiyingi Yu and Liu, 1990: 8. China. 

zhogolevi Remm, in Remm and Zhogolev, 1968: 836. Ukraine. 

zhongningensis Yu, 1982: 201 (1981: 31). China. 

zhuhaiensis Yu and Hao, in Yu, 1988: 137. China. 

zikaensis Khamala and Kettle, 1971: 56. Uganda. 

zuluensis de Meillon, 1936: 145. South Africa. 

acastus de Meillon, 1947: 118. South Africa. 

zumbadoi Spinelli and Borkent, 2004a: 383. Costa Rica. 

zyzzamaculatus Yu, 1982: 202. China. 

Genus PARADASYHELEA Macfie 

PARADASYHELEA Macfie, 1940f: 17. Type species: Dasyhelea brevipalpis Ingram and Macfie, by original designation. 

albipunctata Wirth and Lee, 1959: 116. Australia (New South Wales). 

boucheti Clastrier, 1989b: 134. New Caledonia (France). 

brevipalpis (Ingram and Macfie), 1931a: 178 (Dasyhelea). Argentina. 

egregria (Macfie), 1932c: 36 (Dasyhelea). New Zealand. 

harrisoni Wirth, 1981: 384. New Zealand. 

ingrami Spinelli and Grogan, 2003: 572. Argentina. 

macfiei Spinelli and Grogan, 2003: 574. Argentina. 

minuta Wirth and Lee, 1959: 118. Australia (New South Wales). 


olympiae Wirth and Blanton, 1969c: 98. USA (Washington). 

reyei Elson-Harris and Kettle, 1985: 233. Australia (Queensland). 

Genus WASHINGTONHELEA Wirth and Grogan 

WASHINGTONHELEA Wirth and Grogan, 1988: 97. Type species: Washingtonhelea frommeri Wirth and Grogan, by 


frommeri Wirth and Grogan, 1988: 98. USA (California). 

taimyrica Szadziewski, 1996: 71. Russia. Upper Cretaceous. 

TRIBE CERATOPOGONINI 

Genus ADELOHELEA Borkent 

ADELOHELEA Borkent, 1995: 45. Type species: Adelohelea glabra Borkent, by original designation. 

105

glabra Borkent, 1995: 46. Canada (Manitoba). Upper Cretaceous. 

magyarica Borkent, 1997c: 6. Hungary. Upper Cretaceous. 

Genus AFROHELEA Wirth 

AFROHELEA Wirth, 1965b: 230. Type species: Parabezzia capensis de Meillon and Hardy), by original designation. 

capensis (de Meillon and Hardy), 1954: 67 (Parabezzia). South Africa. 

Genus AGILIHELEA Yu 

AGILIHELEA Yu, in Yu et al., 2005: 1326. Type species: Agilihelea jiania Yu, by original designation. 

jiania Yu, in Yu et al., 2005: 1326. China. 

Genus ALLOHELEA Kieffer 

ALLOHELEA Kieffer, 1917b: 364. Type species: Sphaeromias pulchripennis Kieffer, by original designation. 

afra Clastrier and Delécolle, 1990: 130. Guinea. 

africana (Clastrier), 1960b: 282 (Monohelea). Congo. 

allocota Yu and Zhang, in Yu et al., 2005: 1329. China. 

ampligonata (Ratanaworabhan and Wirth), 1972: 468 (Monohelea). Malaysia. 

annulata Yu and Yan, 2004:37(Yu, in Yu et al., 2005: 1330). China. 

arboricola Clastrier and Delécolle, 1990: 138. Guinea. 

arcuata (Ratanaworabhan and Wirth), 1972: 469 (Monohelea). Malaysia. 

basiflava (Tokunaga), 1963a: 248 (Monohelea). Papua New Guinea. 

basilobata (Ratanaworabhan and Wirth), 1972: 463 (Monohelea). Malaysia. 

bellula (Macfie), 1934c: 282 (Monohelea). Malaysia. 

borealosinica Yu and Liu, in Yu et al., 2005: 1339. China. 

bottimeri Wirth, 1991a: 502. USA (Texas). 

brinchangensis (Ratanaworabhan and Wirth), 1972: 452 (Monohelea). Malaysia. 

camptostyla (Ratanaworabhan and Wirth), 1972: 461 (Monohelea). Malaysia. 

capitata (Ratanaworabhan and Wirth), 1972: 459 (Monohelea). Malaysia. 

chelagonata (Ratanaworabhan and Wirth), 1972: 454 (Monohelea). Malaysia. 

digitata (Ratanaworabhan and Wirth), 1972: 466 (Monohelea). Malaysia. 

distortifemur Wirth, 1991a: 499. USA (Florida). 

fruticosa Yan and Yu, in Liu et al., 1996: 359. China. 

guineensis Clastrier and Delécolle, 1990: 135. Guinea. 

harpagonifera (Debenham), 1972: 9 (Monohelea). Australia (Capital Territory). 

inflativena (Tokunaga), 1962a: 214 (Monohelea). Japan. 

insularis (Tokunaga), 1941a: 114 (Monohelea). Micronesia. 

japonica (Udaka), 1959: 20 (Monohelea). Japan. 

jianfengensis Liu and Yu, in Liu et al., 1996: 358. China. 

johannseni (Wirth), 1953b: 153 (Monohelea). USA (Virginia). 

kindiae Clastrier and Delécolle, 1990: 140. Guinea. 

limosa Clastrier and Delécolle, 1990: 143. Guinea. 

litoraurea (Ingram and Macfie), 1921: 344 (Monohelea). Ghana. 

venustula (Goetghebuer), 1935d: 178 (Monohelea). Democratic Republic of the Congo. 

makonde (de Meillon and Wirth), 1983a: 363 (Monohelea). South Africa. 

maureenae (de Meillon and Wirth), 1987a: 53 (Monohelea). South Africa. 

meeseri (de Meillon), 1939a: 16 (Monohelea). South Africa. 

mimas (de Meillon), 1939a: 13 (Monohelea). South Africa. 

106

minxia Yu and Yan, 2004:39 (Yu and Yan, in Yu et al., 2005: 1333). China. 

nebulosa (Coquillett), 1901a: 606 (Ceratopogon). USA (New Jersey). 

neotropica Wirth, 1991a: 503. Jamaica. 

nigripes (Ratanaworabhan and Wirth), 1972: 460 (Monohelea). Thailand. 

pallifemorata (Remm), 1980: 91 (Monohelea). Tajikistan. 

papuae (Tokunaga), 1963a: 245 (Monohelea). Papua New Guinea. 

parafurcata (Ratanaworabhan and Wirth), 1972: 458 (Monohelea). Malaysia. 

paucimaculata Clastrier and Delécolle, 1990: 136. Guinea. 

pedicellata Wirth, 1991a: 502. USA (Florida). 

pulchripennis (Kieffer), 1911c: 344 (Sphaeromias). Seychelles. 

quatei (Ratanaworabhan and Wirth), 1972: 457 (Monohelea). Malaysia. 

solidipedalis (Tokunaga), 1963a: 247 (Monohelea). Papua New Guinea. 

subigripes Yu and Deng, in Yu et al., 2005: 1336. China. 

superlobata (Ratanaworabhan and Wirth), 1972: 462 (Monohelea). Thailand. 

sylvatica (Clastrier), 1960b: 284 (Monohelea). Congo. 

tenuilobata (Ratanaworabhan and Wirth), 1972: 455 (Monohelea). Thailand. 

tessellata (Zetterstedt), 1850: 3642 (Ceratopogon). Sweden. 

illustris (Winnertz), 1852: 53 (Ceratopogon). Germany and Great Britain. 

tokunagai (Remm), 1993: 185 (Monohelea). Russia. 

tricuspis (Debenham), 1972: 10 (Monohelea). Australia (New South Wales). 

vespertilio Szadziewski, Gwizdalska-Kentzer and Gilka, 2011: 644. United Arab Emirates. 

weemsi Wirth, 1991a: 501. USA (Florida). 

yorkensis (Debenham), 1972: 27 (Monohelea). Australia (Queensland). 

zherihhini (Remm), 1993: 185 (Monohelea). Russia. 

zhuhaiensis Yu and Hao, in Yu et al., 2005: 1337. China. 

Genus ALLUAUDOMYIA Kieffer 

ALLUAUDOMYIA Kieffer, 1913e: 12. Type species: Alluaudomyia imparunguis Kieffer, by monotypy. 

NEOCERATOPOGON Malloch, 1915b: 310. Type species: Ceratopogon bellus Coquillett, by original designation. 

PRIONOGNATHUS Carter, Ingram and Macfie, 1921a: 309 (preoccupied by Prionognathus LaFerté-Sénectère, 1851). Type 

species: Prionognathus marmoratus Carter, Ingram and Macfie, by original designation. 

THYSANOGNATHUS Ingram and Macfie, 1922: 244. New name for Prionognathus Carter, Ingram and Macfie. Type 

species: Prionognathus marmoratus Carter, Ingram and Macfie, automatic. 

ISOECACTA Garrett, 1925: 9. Type species: Isoecacta poeyi Garrett (= Ceratopogon bellus Coquillett), by original 

designation. 


abdominalis Wirth and Delfinado, 1964: 606. Malaysia. 

acicula Yu, in Yu et al., 2005: 1369. China. 

adunca Wirth and Delfinado, 1964: 609. Thailand. 

albigena Wirth and Delfinado, 1964: 618. Thailand. 

albopicta (Ingram and Macfie), 1922: 244 (Thysanognathus). Ghana. 

alpina Debenham, 1971: 146. Australia (New South Wales). 

altalocei Delécolle and Rieb, 1989: 127. France. 

amazonica Spinelli and Wirth, 1984c: 676. Brazil. 

angulata Wirth and Delfinado, 1964: 611. Malaysia. 

annulata Wirth and Delfinado, 1964: 637. Malaysia. 

annulipes Wirth and Delfinado, 1964: 637. Thailand. 

australiensis Debenham, 1971: 133 (valid subspecies of annulipes). Australia (Queensland). 

anserina de Meillon and Wirth, 1983a: 354. South Africa. 

appendiculata Debenham, 1971: 151. Australia (New South Wales). 

asperata Yu and Liu, in Yu et al., 2005: 1348. China. 

astera Tokunaga, 1963a: 222. Papua New Guinea. 

107

aterivena Tokunaga, 1940d: 154. Japan. 

bella (Coquillett), 1902a: 87 (Ceratopogon). USA (Washington, D.C.). 

poeyi (Garrett), 1925: 9 (Isoecata). Canada (British Columbia). 

bertrandi Harant and Cellier, 1949: 10. France. 

bicornis Debenham, 1971: 159. Australia (Northern Territory). 

biestroi Spinelli, 1988: 130. Argentina. 

bifasciata Tokunaga, 1963a: 220. Papua New Guinea. 

bifurcata Wirth and Delfinado, 1964: 633. Malaysia. 

bimaculata Clastrier and Wirth, 1961a: 228. Nigeria. 

bipunctata Tokunaga and Murachi, 1959: 356. Belau (USA). 

bispinula Zhang and Yu, in Zhang et al., 2004: 317 (Yu and Deng, in Yu et al., 2005: 1349). China. 

bistorta Yu and Zou, in Yu et al., 2005: 1377. China. 

boucheti Clastrier, 1985c: 194. New Caledonia (France). 

brandti Tokunaga, 1963a: 223. Papua New Guinea. 

brevicosta Clastrier, 1960b: 261. Congo. 

brevis Wirth and Delfinado, 1964: 608. Malaysia. 

candidata Yu, 1999: 220. China. 

caracalla Yu and Kong, in Yu et al., 2005: 1351. China. 

caribbeana Spinelli and Wirth, 1984c: 678. Belize. 

catarinensis Spinelli and Wirth, 1984c: 681. Brazil. 

claudia de Meillon, 1942a: 92. Zimbabwe. 

cobra de Meillon and Wirth, 1987a: 41. South Africa. 

columinis Liu, Yan and Liu, 1996: 42. China. 

congolensis de Meillon, 1939b: 12. Democratic Republic of the Congo. 

conjuncta Kieffer, 1918a: 52. Guinea. 

crumena Yu and Liu, in Yu et al., 2005: 1379. China. 

dekeyseri Clastrier, 1958c: 487. Senegal. 

delfinadoae Giles and Wirth, 1984: 211. Malaysia. 

demeilloni Clastrier and Wirth, 1961a: 223. Nigeria. 

debilipenis Sinha, Mazumdar and Chaudhuri, 2005: 116. India. 

desma Yu, in Yu et al., 2005: 1346. China. 

distispinulosa Spinelli and Wirth, 1984c: 682. Brazil. 

epsteini Giles and Wirth, 1987: 459. New Caledonia (France). 

estevezae Spinelli and Wirth, 1984c: 684. Mexico. 

exigua Clastrier, 1985c: 193. New Caledonia (France). 

falcata de Meillon and Wirth, 1983a: 356. South Africa. 

fimbriatinervis Clastrier, 1958b: 230. Senegal. 

finitima Sinha, Mazumdar and Chaudhuri, 2005: 118. India. 

fittkaui Spinelli and Wirth, 1984c: 686. Brazil. 

flexistyla Chaudhuri and Ghosh, 1980: 422. India. 

flexuosa Yu and Hao, in Yu et al., 2005: 1352. China. 

footei Wirth, 1952c: 428. USA (Florida). 

forcipulata Yu and Liu, in Yu et al., 2005: 1372. China. 

formosana Okada, 1942: 317 (as variety of maculipennis). Taiwan. 

fragilicornis Clastrier, 1958c: 489. Senegal. 

fragmentum Debenham, 1971: 163. Australia (Northern Territory). 

fregata Yu, Li and Nie, in Nie et al., 2009:162. China (on ship from Paradip, India). 

fumosipennis Debenham, 1971: 135. Australia (Northern Territory). 

fuscipennis Wirth and Delfinado, 1964: 612. Thailand. 

fuscipes Wirth and Delfinado, 1964: 606. Malaysia. 

fuscitarsis Chaudhuri, Das Gupta and Chatterjee, 1981: 147. India. 

fuscula Yu and Liu, in Yu et al., 2005: 1355. China. 

gloriosa Kieffer, 1925a: 422. Austria. 

griffithi Wirth and Delfinado, 1964: 635. Thailand. 

108

guarani Spinelli, 1988: 131. Uruguay. 

halterata Remm, 1993: 187. Russia. 

hirsutipennis Clastrier, 1960b: 267. Congo. 

huberti Wirth and Delfinado, 1964: 622. Malaysia. 

hygropetrica Vaillant, 1954: 228. France. 

immaculata Tokunaga, 1963a: 219. Indonesia. 

imparunguis Kieffer, 1913e: 12. Kenya. 

inaequalis Wirth and Delfinado, 1964: 613. Thailand. 

infuscata Wirth and Delfinado, 1964: 621. Malaysia. 

inexspectata Clastrier, 1983c: 10. Seychelles. 

insignis Chaudhuri, Das Gupta and Chaudhuri, 1972: 94. India. 

insulana Tokunaga and Murachi, 1959: 358. Micronesia. 

insulicola Tokunaga and Murachi, 1959: 361. Micronesia. 

jimmensis Tokunaga, 1963a: 229. Papua New Guinea. 

lactella Remm, 1980: 97. Kyrgyzstan. 

latipennis (Skuse), 1889: 308 (Ceratopogon). Australia (New South Wales). 

leei Spinelli and Wirth, 1984c: 687. Colombia. 

limosa Clastrier, 1961a: 432. France. 

ljatifeidae Remm, 1967: 27 (as latifeidae). Azerbaijan. 

longzhouensis Hao and Yu, 1991: 45. China. 

louisi de Meillon and Wirth, 1981b: 535. South Africa. 

lucania Li and Yu, 1997: 124. China. 

lunata de Meillon and Wirth, 1983a: 356. South Africa. 

macclurei Wirth and Delfinado, 1964: 622. Malaysia. 

maculiabdominis Yu, Nie and Li, in Nie et al., 2007: 476. Australia or India. 

maculipennis (Carter, Ingram and Macfie), 1921a: 316 (Prionognathus). Ghana. 

maculithorax (Carter, Ingram and Macfie), 1921a: 319 (Prionognathus). Ghana. 

maculosa de Meillon, 1936: 172. South Africa. 

maculosipennis Tokungaga, 1940e: 181 (1940d: 154). Micronesia. 

maculosissima Wirth and Delfinado, 1964: 624. Indonesia. 

magoebai de Meillon, Meiswinkel and Wirth, 1982: 134. South Africa. 

magna Wirth and Delfinado, 1964: 632. Thailand. 

marginalis Wirth and Delfinado, 1964: 625. Malaysia. 

marmorata (Carter, Ingram and Macfie), 1921a: 312 (Prionognathus). Ghana. 

marmorea Clastrier, 1960b: 258. Congo. 

mcmillani Clastrier and Wirth, 1961a: 236. Nigeria. 

megaparamera Williams, 1957: 327. USA (Michigan). 

meeseri (de Meillon and Hardy), 1954: 74 (Ceratopogon). Zimbabwe. 

melanesiae Clastrier, 1985c: 190. New Caledonia (France). 

melanosticta (Ingram and Macfie), 1922: 248 (Thysanognathus). Ghana. 

meridiana Clastrier, 1978: 25. France. 

monopunctata Tokunaga and Murachi, 1959: 354. Micronesia. 

mouensis Giles and Wirth, 1987: 463. New Caledonia (France). 

mynistensis Remm, 1979b: 55. Estonia. 

natalensis de Meillon, 1939b: 14. South Africa. 

needhami Thomsen, 1935: 287. USA (New York). 

pentaspila Remm and Glukhova, 1971: 304. Russia. 

neocaledoniensis Clastrier, 1985c: 188. New Caledonia (France). 

nilogenes (Kieffer), 1925e: 262 (Thysanognathus). Egypt. 

novaguineae Tokunaga, 1963a: 227. Papua New Guinea. 

nubeculosa Spinelli and Wirth, 1984c: 689. Brazil. 

ocellata Remm, 1980: 99. Tajikistan. 

onoi Tokunaga, 1972: 15. Japan. 

opacata Yu, in Yu et al., 2005: 1380. China. 

109

ornatithorax Chaudhuri, Das Gupta and Chaudhuri, 1972: 89. India. 

pacifica Clastrier, 1985c: 195. New Caledonia (France). 

papuae Tokunaga, 1963a: 217. Papua New Guinea. 

parafurcata Wirth and Delfinado, 1964: 640. Malaysia. 

paraspina Wirth, 1952c: 429. USA (Georgia). 

parva Wirth, 1952c: 431. USA (Florida). 

downesi Wirth, 1952c: 433. USA (Virginia). 

peculiaris Chaudhuri, Das Gupta and Chaudhuri, 1972: 100. India. 

personata Debenham, 1971: 133. Australia (Queensland). 

petersi Tokunaga, 1963a: 228. Papua New Guinea. 

platipyga Tokunaga, 1963a: 221. Papua New Guinea. 

plaumanni Spinelli and Wirth, 1984c: 691. Brazil. 

polyommata Macfie, 1947b: 76. Sudan. 

poguei Giles and Wirth, 1987: 461. New Caledonia (France). 

prima Clastrier, 1976: 205. French Guiana. 

pseudomaculipennis (Carter, Ingram and Macfie), 1921a: 318 (Prionognathus). Ghana. 

pseudomaculithorax Clastrier, 1958b: 225. Senegal. 

pseudomarginalis Wirth and Delfinado, 1964: 628. Philippines. 

punctiradialis Chaudhuri, Das Gupta and Chaudhuri, 1972: 97. India. 

punctivenosa Wirth and Grogan, 1988: 20. New name for punctiradialis. 

punctiradialis Spinelli and Wirth, 1984c: 692 (preoccupied by Alluaudomyia punctiradialis Chaudhuri, Das Gupta and 

Chaudhuri, 1972). Brazil. 

punctulata Wirth and Delfinado, 1964: 639. Malaysia. 

quadripunctata (Goetghebuer), 1934c: 288 (Culicoides). Russia. 

quadrispina Tokunaga, 1972: 17. Japan. 

quasivudu Stam, 1964: 195. Democratic Republic of the Congo. 

quinquenebulosa Wirth and Delfinado, 1964: 619. Malaysia. 

quinquepicina Yu and Zhyang, in Yu et al., 2005: 1361. China. 

quinquepunctata Tokunaga, 1940b: 256. Japan. 

remmi Szadziewski, 1983b: 395. Algeria. 

reyei Debenham, 1971: 157. Australia (Northern Territory). 

riparia Clastrier, 1978: 29. France. 

falcata Knoz and Ratajsky, 1987: 566 (preoccupied by Alluaudomyia falcata de Meillon and Wirth, 1983a). Czech 

Republic. 

bohemiae Boorman, 1997: 173. New name for falcata. 

rostrata Yu and Zhang, in Zhang et al., 2004: 317 (Yu and Deng, in Yu et al., 2005: 1362). China. 

rudolfi de Meillon and Downes, 1986: 171. South Africa. 

sagaensis Tokunaga, 1940b: 257. Japan. 

schnacki Spinelli, 1983b: 403. Argentina. 

senta de Meillon, 1936: 175. South Africa. 

sexpunctata Spinelli and Wirth, 1984c: 695. Colombia. 

shogakii Tokunaga, 1960a: 74. Japan. 

siebenschwabi Havelka, 1982: 59. Spain. 

signosoma Yu and Zhang, in Yu et al., 2005: 1363. China. 

similiforceps Clastrier, 1960b: 265. Congo. 

simulata Sinha, Mazumdar and Chaudhuri, 2005: 119. India. 

smeei Tokunaga, 1963a: 230. Papua New Guinea. 

sophiae Stam, 1964: 193. Democratic Republic of the Congo. 

sordidipennis Clastrier and Wirth, 1961a: 233. Gambia. 

soutini de Meillon and Wirth, 1983a: 359. South Africa. 

spinellii Wirth and Grogan, 1988: 21. New name for tripunctata. 

tripunctata Spinelli and Wirth, 1984c: 697 (preoccupied by Alluaudomyia tripunctata Chaudhuri, Das Gupta and 

Chaudhuri, 1972). Colombia. 

spinosiforceps Clastrier, 1977: 345. Guinea. 

110

spinosipes Tokunaga, 1962a: 206. Japan. 

splendentis Liu, Yan and Liu, 1996: 42. China. 

splendida (Winnertz), 1852: 47 (Ceratopogon). Europe. 

sternalis Wirth and Delfinado, 1964: 617. Thailand. 

stictipennis Wirth, 1952a: 197. USA (California). 

streptomera Remm, 1980: 99. Turkmenistan. 

striata Remm, 1993: 187. Russia. 

subadunca Chaudhuri, Das Gupta and Chaudhuri, 1972: 95. India. 

subannulata Wirth and Delfinado, 1964: 636. Malaysia. 

subflexuosa Yu, in Yu et al., 2005: 1353. China. 


tauffliebi Clastrier, 1960b: 263. Congo. 

tenuiannulata Spinelli and Wirth, 1984c: 696. Guatemala. 

tenuistylata Tokunaga, 1959: 296. Indonesia. 

thurmanorum Wirth and Delfinado, 1964: 629. Thailand. 

tiberghieni Neveu, 1978: 355. France. 

depuncta Remm, 1980: 101. Tajikistan. 

tillierorum Clastrier, 1985c: 191. New Caledonia (France). 

tokunagai Wirth and Delfinado, 1964: 633. New name for splendida. 

splendida Tokunaga, 1963a: 216 (preoccupied by Alluaudomyia splendida (Winnertz), 1852). Papua New Guinea. 

transvaalensis de Meillon, 1947: 119. South Africa. 

tripartita Okada, 1942: 316 (as variety of sagaensis). Taiwan. 

tripunctata Chaudhuri, Das Gupta and Chaudhuri, 1972: 91. India. 

tropaea Yu, in Yu et al., 2005: 1366. China. 

typica Chaudhuri, Das Gupta and Chaudhuri, 1972: 91. India. 

undecimpunctata Tokunaga, 1940b: 257. Japan. 

unguistyla Debenham, 1971: 139. Australia (New South Wales). 

ussurica Isaev, 1993: 70. Russia. 

varia Debenham, 1971: 154. Australia (Queensland). 

variegata Glick and Mullen, 1982: 540. USA (Alabama). 

verecunda Debenham, 1971: 143. Papua New Guinea. 

vicina Clastrier, 1960b: 260. Congo. 

vudu de Meillon and Hardy, 1954: 65. Cameroon. 

wansoni de Meillon, 1939b: 10. Democratic Republic of the Congo. 

wirthi Williams, 1957: 328. USA (Michigan). 

xanthocoma (Kieffer), 1913d: 182 (Culicoides). India. 

youngi Spinelli and Wirth, 1984c: 699. Colombia. 

Genus ANKYLOHELEA de Meillon and Wirth 

ANKYLOHELEA de Meillon and Wirth, 1987b: 384. Type species: Ankylohelea montana de Meillon and Wirth, by original 

designation. 

montana de Meillon and Wirth, 1987b: 384. South Africa. 

Genus ATYPHOHELEA Borkent 

ATYPHOHELEA Borkent, 1998: 138. Type species: Johannsenomyia macroneura Malloch, by original designation. 

cothurnata (Meunier), 1904a: 231 (1904b: 242) (Ceratopogon). Baltic region. Eocene. 

flagella Meunier, 1904a: 230 (1904b: 241) (Ceratopogon). Baltic region. Eocene. 

macroneura (Malloch), 1915a: 337 (Johannsenomyia). USA (Kansas). 

nigra (Wirth), 1952a: 208 (Neurohelea). USA (California). 

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Genus AUSTROHELEA Wirth and Grogan 

AUSTROHELEA Wirth and Grogan, 1988: 22. Type species: Monohelea shannoni Wirth and Blanton, by original 

designation. 

antipodalis (Ingram and Macfie), 1931b: 205 (Monohelea). New Zealand. 

brevipes (Lee), 1948c: 355 (Monohelea). Australia (Western Australia). 

campbellensis (Tokunaga), 1964b: 289 (Monohelea). New Zealand. 

ferruginea (Macfie), 1932c: 49 (Monohelea). New Zealand. 

shannoni (Wirth and Blanton), 1972b: 175 (Monohelea). Argentina. 

tasmaniensis (Lee), 1948c: 354 (Monohelea). Australia (Tasmania). 

tonnoiri (Macfie), 1932c: 46 (Monohelea). New Zealand. 

Genus BAEODASYMYIA Clastrier and Raccurt 

BAEODASYMYIA Clastrier and Raccurt, 1979b: 100. Type species: Baeodasymyia modesta Clastrier and Raccurt, by 

monotypy. 

christopheri Borkent, in Borkent and Craig, 1999: 15. Costa Rica. 

dominicana Szadziewski and Grogan, 1994: 220. Dominican Republic. Miocene. 

gustavoi Borkent, in Borkent and Craig, 1999: 20. Paraguay. 

lydiae Borkent, in Borkent and Craig, 1999: 19. Costa Rica. 

michaeli Borkent, in Borkent and Craig, 1999: 13. Costa Rica. 

modesta Clastrier and Raccurt, 1979b: 100. Haiti. 

Genus BAEOHELEA Wirth and Blanton 

BAEOHELEA Wirth and Blanton, 1970c: 95. Type species: Baeohelea nana Wirth and Blanton, by original designation. 

nana Wirth and Blanton, 1970c: 95. Dominica. 

Genus BAHIAHELEA Wirth 

BAHIAHELEA Wirth, 1992: 276. Type species: Bahiahelea brasiliensis Wirth, by original designation. 

brasiliensis Wirth, 1992: 280. Brazil. 

Genus BOREOHELEA Clastrier and Delécolle 

BOREOHELEA Clastrier and Delécolle, 1990: 146. Type species: Boreohelea afrotropica Clastrier and Delécolle, by 


afrotropica Clastrier and Delécolle, 1990: 146. Guinea. 

Genus BORKENTHELEA Spinelli and Grogan 

BORKENTHELEA Spinelli and Grogan, 1993: 321. Type species: Borkenthelea nothofagus Spinelli and Grogan, by original 

designation. 

harii Spinelli and Grogan, 2001: 148. Argentina. 

nerudai Spinelli and Grogan, 2001: 151. Chile. 

nothofagus Spinelli and Grogan, 1993: 323. Argentina. 

quatei Spinelli and Grogan, 2001: 153. Argentina. 

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Genus BOTHAHELEA Grogan and Wirth 

BOTHAHELEA Grogan and Wirth, 1983: 199. Type species: Bothahelea phelpsi Grogan and Wirth, by original designation. 

gigantostyla Grogan and Wirth, 1983: 203. Zimbabwe. 

nama de Meillon and Wirth, 1987a: 42. South Africa. 

phelpsi Grogan and Wirth, 1983: 200. Zimbabwe. 

Genus BOTHAMIA Meiswinkel 

BOTHAMIA Meiswinkel, 1987: 300. Type species: Bothamia demeilloni Meiswinkel, by original designation. 

demeilloni Meiswinkel, 1987: 300. South Africa. 

Genus BRACHYCRETACEA Szadziewski 

BRACHYCRETACEA Szadziewski, 1996: 63. Type species: Brachycretacea taimyrica (Remm), by original designation. 

taimyrica (Remm), 1976b: 115 (Baeohelea). Russia. Upper Cretaceous. 

Genus BRACHYPOGON Kieffer 

BRACHYPOGON Kieffer, 1899: 69. Type species: Ceratopogon vitiosus Winnertz, by original designation. 

Subgenus BRACHYPOGON Kieffer 

acutus Yu, in Yu et al., 2005: 1388. China. 

afifi Boorman and Harten, 2002: 452. Yemen. 

americanus Szadziewski and Grogan, 1998a: 42. Dominican Republic. Miocene. 

apunctipennis Spinelli and Grogan, 1998: 64. Brazil. 

arabicus Szadziewski, Gwizdalska-Kentzer and Gilka, 2011: 646. United Arab Emirates. 

artemis Debenham, 1991: 771. Australia (New South Wales). 

baculus Das, Mazumdar, and Chaudhuri, 2010: 456. India. 

balticus Szadziewski, 1988: 93. Baltic region. Eocene. 

beaufortanensis Delécolle and Rieb, 1992: 33. France. 

sudowicus Szadziewski, 2001: 211. Poland. 

bergensis (de Meillon and Hardy), 1953: 26 (Ceratopogon). South Africa. 

bifidus Spinelli and Grogan, 1998: 66. Dominica. 

bifurcus Debenham, 1991: 780. Australia (New South Wales). 

bimaculatus Spinelli and Grogan, 1998: 67. Colombia. 

bonaerensis Spinelli, 1990: 744. Argentina. 

boormani Grogan and de Meillon, 1993: 400. Senegal. 

brazzai (Vattier and Adam), 1966b: 736 (Ceratopogon). Congo. 

bryanae Debenham, 1991: 774. Australia (New South Wales). 

caecus Sarkar and Das Gupta, in Saha and Das Gupta, 2006:18. India. 

calchaqui Spinelli, 1990: 746. Argentina. 

calloti Grogan and de Meillon, 1993: 403. Senegal. 

canadensis Downes, 1976: 1147. Canada (Quebec). 

carpinteroi Spinelli and Marino, 2008: 120. Peru. 

chuanxicus Yu and Wen, in Yu et al., 2005: 1389. China. 

citrithecus Yu and Lai, in Yu et al., 2005: 1390. China. 

complanothecus Yu, in Yu et al., 2005: 1392. China. 

compressus Yu and Liu, in Yu et al., 2005: 1393. China. 

corius (de Meillon and Hardy), 1954: 69 (Ceratopogon). South Africa. 

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emphuxi (Vattier and Adam), 1966b: 734 (Ceratopogon). Congo. 

corneti Grogan and de Meillon, 1993: 392. Senegal. 

corniger Debenham, 1991: 777. Australia (New South Wales). 

curtus Debenham, 1991: 796. Australia (Queensland). 

debenhamae Grogan and de Meillon, 1993: 405. Senegal. 

defectivus (Strobl), 1906: 400 (Ceratopogon). Spain. 

delecollei Grogan and de Meillon, 1993: 404. Senegal. 

demeilloni Grogan and Wirth, 1993: 30. Zimbabwe. 

ecuadorensis Spinelli and Grogan, 1998: 68. Ecuador. 

emeiensis Wen and Yu, in Wen et al., 1991: 30. China. 

eocenicus Szadziewski, 1988: 91. Denmark. Eocene. 

ethelae Spinelli and Grogan, 1998: 62. Panama. 

fagicola Delécolle and Schiegg, 1999a: 31. Switzerland. 

flecteris Das, Mazumdar, and Chaudhuri, 2010: 457. India. 

fuscivenosus (Lutz), 1914: 94 (Palpomyia). Brazil. 

gearyae Debenham, 1991: 786. Australia (New South Wales). 

gedanicus Szadziewski, 1988: 95. Poland. Eocene. 

gongdongi Yu and Huang, in Yu et al., 2009:48. China. 

gravidus Debenham, 1991: 790. Australia (Queensland). 

griffithsae Debenham, 1991: 797. Australia (Queensland). 

grobleri de Meillon and Wirth, 1981c: 575. South Africa. 

hamiltoni de Meillon and Wirth, 1981c: 576. South Africa. 

halimos Yu and Li, in Li et al., 2008: 706. China. 

havelkai Grogan and de Meillon, 1993: 402. Central African Republic. 

hercules Debenham, 1991: 788. Australia (New South Wales). 

idolon Debenham, 1991: 791. Australia (Queensland). 

impar (Johannsen), 1938: 223 (Ceratopogon). Puerto Rico (USA). 

infarctipes Delécolle and Grogan, 1990: 133. Senegal. 

infrequens Saha and Das Gupta, in Saha and Das Gupta, 2006:20. India. 

insularis Spinelli and Grogan, 1998: 70. Cuba. 

insulicola (Tokunaga), in Tokunaga and Murachi, 1959: 350 (Ceratopogon). Micronesia. 

institor Debenham, 1991: 796. Australia (New South Wales). 

jorgei Spinelli and Marino, 2008: 122. Peru. 

kokocinskii Szadziewski, 1983b: 393. Algeria. 

kremeri Szadziewski and Havelka, 1984: 353. North Korea. 

krugeri de Meillon and Wirth, 1981c: 578. South Africa. 

krzeminskii Szadziewski and Havelka, 1984: 344. North Korea. 

laneae Swanson and Grogan, 2011: 533. USA (Alabama). 

latifemoris Clastrier, 1989c: 190. Guinea. 

libanius (de Meillon), 1943: 104 (Ceratopogon). South Africa. 

limushanensis Liu, Yan and Liu, 1996: 41. China. 

lobulus Yu and Zhang, in Yu et al., 2005: 1399. China. 

lorica Debenham, 1991: 793. Australia (Queensland). 

medusae Debenham, 1991: 778. Australia (New South Wales). 

maitangensis Yu, Wang and Tan, in Wang et al., 2009:181. China. 

miocaenicus Szadziewski, 1993: 610. Germany. Eocene. 

mireillae (Vattier and Adam), 1966b: 733 (Ceratopogon). Congo. 

monicae Spinelli and Grogan, 1998: 70. Colombia. 

nanchuanensis Yu, in Yu et al., 2005: 1400. China. 

neotericus Yu and Zhang, in Wen et al., 1991: 32. China 

nicolaii Debenham, 1991: 798. Papua New Guinea. 

nieves (Havelka), 1976b: 91 (Ceratopogon). Germany. 

novaguineae (Tokunaga), 1964a: 294 (Ceratopogon). Papua New Guinea. 

obesus Szadziewski, Gwizdalska-Kentzer and Gilka, 2011: 647. United Arab Emirates. 

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optimus Saha and Das Gupta, in Saha and Das Gupta, 2006:21. India. 

pakistanicus Szadziewski and Havelka, 1984: 346. Pakistan. 

paraensis Wirth and Blanton, 1970c: 99 (as parasensis). Brazil. 

peruensis Spinelli and Marino, 2008: 118. Peru. 

petersi (Tokunaga), 1964a: 293 (Ceratopogon). Papua New Guinea. 

phymos Delécolle and Grogan, 1990: 129. Senegal. 

placodus Yu, in Yu et al., 2005: 1403. China. 

pollices Debenham, 1991: 783. Australia (New South Wales). 

proprius Sarkar and Das Gupta, in Saha and Das Gupta, 2006:21. India. 

pseudocanadensis Sarkar and Das Gupta, in Saha and Das Gupta, 2006:22. India. 

pseudoparaensis Spinelli and Grogan, 1998: 64. Brazil. 

riebi Grogan and de Meillon, 1993: 397. Senegal. 

ringueleti Spinelli, 1990: 748. Argentina. 

schmitzi Spinelli and Grogan, 1998: 65. Brazil. 

senegalensis (de Meillon and Wirth), 1955: 275 (Ceratopogon). Senegal. 

sitius (de Meillon), 1959a: 345 (Ceratopogon). South Africa. 

soavei Spinelli and Marino, 2008: 128. Peru. 

sparsus Yu, in Yu et al., 2005: 1404. China. 

spatuliformis Spinelli and Grogan, 1998: 65. Brazil. 

spinosipes Clastrier, 1989c: 194. Guinea. 

spinosissimus Clastrier, 1989c: 195. Guinea. 

subiectus Debenham, 1991: 789. Australia (New South Wales). 

szadziewskii Grogan and de Meillon, 1993: 400. Senegal. 

tabernaculum Debenham, 1991: 784. Australia (New South Wales). 

telesfordi Spinelli and Grogan, 1998: 71. St. Vincent. 

tokunagai Grogan and Wirth, 1981c: 96. Solomon Islands. 

usticius Yu and Liu, in Yu et al., 2005: 1405. China. 

vitiosus (Winnertz), 1852: 49 (Ceratopogon). Germany. 

minimus (Kieffer), 1924b: 402 (Anakempia). France. 

niger (Mayer), 1934a: 291 (Trishelea). Poland. 

woodruffi Spinelli and Grogan, 1998: 72. Dominica Republic. 

wudangshanensis Yu, in Yu et al., 2005: 1408. China. 

xingyangensis Yu, in Yu et al., 2005: 1409. China. 

yarimii Boorman and Harten, 2002: 453. Oman. 

zhuhaiensis Yu and Hao, in Yu et al., 2005: 1411. China. 

Subgenus ISOHELEA Kieffer 

ISOHELEA Kieffer, 1917b: 295. Type species: Ceratopogon lacteipennis Zetterstedt (misidentified, = Psilohelea 

sociabilis Goetghebuer), by original designation. 

NILOHELEA Kieffer, 1921b: 22. Type species: Nilohelea albipennis Kieffer, by monotypy. 

TRISHELEA Kieffer, 1925b: 153. Type species: Trishelea incompleta Kieffer, by original designation. 

abdominalis (Tokunaga), 1940b: 259 (Ceratopogon). Japan. 

aethiopicus (Clastrier, Rioux and Descous), 1961: 87 (Ceratopogon). Chad. 

africanus de Meillon, 1929: 248. South Africa. 

albipennis (Kieffer), 1921b: 22 (Nilohelea). Sudan. 

alpinus (Clastrier), 1961a: 413 (Ceratopogon). Austria. 

aquilonalis (Clastrier), 1961a: 421 (Ceratopogon). Finland. 

babiogorenis Szadziewski, in Szadziewski et al., 1994: 5. Poland. 

basiflagellatus (Tokunaga), 1940b: 263 (Ceratopogon). Japan. 

beskidicus Krzywinski, in Szadziewski et al., 1994: 7. Poland. 

bialoviesicus Krzywinski, in Szadziewski et al., 1994: 9. Poland. 

bilobatus (Remm), 1993: 189 (Ceratopogon). Kazakhstan. 

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orealis (Kieffer), 1919b: 41 (1919a: 68) (Psilohelea). Norway. 

borkenti Spinelli and Cazorla, 2004: 3. Argentina. 

carpaticus Szadziewski, in Szadziewski et al., 1994: 12. Poland. 

caucasicus (Remm), 1967: 28 (Ceratopogon). Georgia. 

clastrieri Szadziewski, 1983b: 386. Algeria. 

clavatus (Clastrier), 1966: 706 (Ceratopogon). Canary Islands (Spain). 

cuacuahuitlus Huerta and Borkent, 2005: 115. Mexico. 

darvazi (Remm), 1974c: 55 (Ceratopogon). Tajikistan. 

dehiscens Debenham, 1991: 802. Australia (New South Wales). 

dilatatus Yu, in Yu et al., 2005: 1413. China. 

dominicanus Szadziewski and Grogan, 1998a: 42. Dominican Republic. Miocene. 

falcifer (Tokunaga), 1940b: 261 (Ceratopogon). Russia. 

ferganicus (Remm), 1974c: 56 (Ceratopogon). Kyrgyzstan. 

flaviventris (Kieffer), 1919a: 66 (Psilohelea). Croatia. 

croaticus (Remm), 1981: 32 (Ceratopogon). New name for flaviventris. 

glukhovae Grogan and de Meillon, 1993: 406. Senegal. 

griseipennis (Storå), 1945: 34 (Isohelea). Azores (Portugal). 

hadrosaurus Debenham, 1991: 801. Australia (New South Wales). 

hamoni (Vattier and Adam), 1966b: 730 (Ceratopogon). Congo. 

henningseni Szadziewski, 1988: 89. Denmark. Eocene. 

herati Navai, 1997: 184. Afghanistan. 

hudjakovi (Remm), 1974c: 53 (Ceratopogon). Russia. 

hugoi Spinelli and Grogan, 1994: 2. Colombia. 

huochengensis Yu and Xiang, in Yu et al., 2005: 1414. China. 

hyperboreus (Clastrier), 1961a: 415 (Ceratopogon). Norway. 

incerticruris Sarkar and Das Gupta, in Saha and Das Gupta, 2006:23. India. 

incompletus (Kieffer), 1925b: 153 (Trishelea). Poland. 

thienemanni (Mayer), 1940: 162 (Helea). Sweden. 

lapiae (Clastrier), 1961a: 416 (Ceratopogon). Finland. 

xuguitensis Cao and Chen, 1984: 298 (Culicoides). China. 

jaroslavi Szadziewski, 1983b: 388. Algeria. 

laricis (Remm), 1974c: 51 (Ceratopogon). Russia. 

longipalpis (Kieffer), 1925a: 412 (Trishelea). Russia. 

lunatus Borkent, in Borkent and Wirth, 1997:93. New name for arcuatus. 

arcuatus (Remm), 1993: 189 (Ceratopogon, preoccupied by Culicoides arcuatus (Winnertz), 1852). Russia. 

magnipalpis (Clastrier), 1961a: 409 (Ceratopogon). France. 

mapuche Spinelli, 1990: 750. Argentina. 

misionensis Spinelli, 1990: 752. Argentina. 

montanus (Tokunaga), 1940d: 152 (Ceratopogon). Japan. 

monticola (Tokunaga), 1940b: 262 (Ceratopogon). Japan. 

montivagus Yu and Wen, in Wen et al., 1991: 35. China. 

nipponensis Wirth and Grogan, 1988: 32. New name for albitarsis. 

albitarsis (Tokunaga), 1940b: 261 (Ceratopogon, preoccupied by Schizohelea albitarsis (Wiedemann), 1817). Japan. 

nitidulus (Edwards), 1921: 125 (Psilohelea). Great Britain. 

crassiforceps (Kieffer), 1925a: 411 (Trishelea). Russia. 

turfaceus (Kieffer), 1925a: 415 (Anakempia). Russia. 

finniae (Clastrier), 1961a: 417 (Ceratopogon). Finland. 

norvegicus Szadziewski and Hagan, 2000: 459. Norway. 

oreinus (Remm), 1974c: 55 (Ceratopogon). Kyrgyzstan. 

pallidipennis Spinelli and Grogan, 1994: 6. Honduras. 

pasquieri (Clastrier), 1961a: 402 (Ceratopogon). Algeria. 

pelecatus (Remm), 1974c: 54 (Ceratopogon). Kyrgyzstan. 

perpusillus (Edwards), 1921: 125 (Psilohelea). Great Britain. 

polonicus Szadziewski, 1988: 88. Poland. Eocene. 

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prominulus (Meunier), 1904a: 228 (1904b: 239) (Ceratopogon). Baltic region. Eocene. 

prominuloides Szadziewski and Grogan, 1998a: 43. Dominican Republic. Miocene. 

pruinosus (Wirth), 1952a: 201 (Helea). USA (California). 

qinghai Yu and Liu, in Yu et al., 2005: 1421. China. 

raohenicus Yu and Liu, in Yu et al., 2005: 1422. China. 

remmi Wirth and Grogan, 1988: 32. New name for falcatus. 

falcatus (Remm), 1974c: 54 (Ceratopogon, preoccupied by Stilobezzia falcata (Meunier), 1904a). Russia. 

sahariensis (Clastrier), 1961a: 422 (Ceratopogon). Algeria. 

saxatilis (Clastrier), 1961a: 411 (Ceratopogon). Algeria. 

serratus (Lewis), 1956: 46 (Helea). USA (Connecticut). 

sevanicus (Remm), 1974c: 50 (Ceratopogon). Armenia. 

silecis Szadziewski, 1990b: 485. Poland. 

sociabilis (Goetghebuer), 1920: 68 (Psilohelea). Belgium. 

sentiger Borkent, in Borkent and Wirth, 1997:94. New name for spiniger. 

spiniger (Remm), 1974c: 53 (Ceratopogon, preoccupied by Serromyia spinigera (Loew), 1850). Kyrgyzstan. 

spinosus Liu, Yan and Liu, 1996: 25. China. 

stigmalis (Coquillett), 1902a: 86 (Ceratopogon). USA (New Mexico). 

stolida Sarkar, Saha and Das Gupta, in Saha and Das Gupta, 2006:23. India. 

subnitidulus Yu and Wang, in Yu et al., 2005: 1424. China. 

surae Szadziewski, 1984a: 555. Algeria. 

susunai (Remm), 1974c: 51 (Ceratopogon). Russia. 

sylvaticus Yu and Liu, in Wen et al., 1991: 35. China. 

taivoi (Remm), 1974c: 50 (Ceratopogon). Kyrgyzstan. 

tiiveli (Remm), 1980: 103 (Ceratopogon). Tajikistan. 

turkestanicus (Remm), 1974c: 56 (Ceratopogon). Kyrgyzstan. 

ussuriensis (Glukhova), 1979: 169 (Isohelea). Russia. 

vaillanti (Mayer), 1955: 111 (Helea). Algeria. 

ajjerensis (Clastrier), 1961a: 408 (Ceratopogon). Algeria. 

vanharteni Szadziewski, Gwizdalska-Kentzer and Gilka, 2011: 647. United Arab Emirates. 

wirthi Spinelli, 1990: 753. Argentina. 

wirthorum Grogan and de Meillon, 1997: 130. Zimbabwe. 

yariensis (Tokunaga), 1940d: 151 (Ceratopogon). Japan. 

zavoicus Szadziewski, in Szadziewski et al., 1994: 29. Poland. 

Subgenus SARISSOHELEA Debenham 

SARISSOHELEA Debenham, 1991: 798 (as subgenus of Brachypogon). Type species: Brachypogon kraussi Grogan and 

Wirth, by original designation. 

alexandros Debenham, 1991: 800. Papua New Guinea. 

kraussi Grogan and Wirth, 1981c: 93. Solomon Islands. 

maai (Tokunaga), 1964a: 298 (Ceratopogon). Indonesia. 

papuensis (Tokunaga), 1964a: 296 (Ceratopogon). Papua New Guinea. 

peregrinator (Edwards), 1928: 56 (Ceratopogon). Western Samoa. 

Species of BRACHYPOGON unplaced to subgenus 

obscurus (Santos Abreu), 1918: 323 (Ceratolophus, as variety of rufigastris). Canary Islands (Spain). 

rufigastris (Santos Abreu), 1918: 321 (Ceratolophus). Canary Islands (Spain). 

singularis (Santos Abreu), 1918: 317 (Ceratolophus). Canary Islands (Spain). 

Genus CACAOHELEA Wirth and Grogan 

CACAOHELEA Wirth and Grogan, 1988: 33. Type species: Cacaohelea youngi Wirth and Grogan, by original designation. 

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adusta Borkent and Picado, 2008: 28. Costa Rica. 

curva Borkent and Picado, 2008: 28. Costa Rica. 

glukhovae Borkent and Picado, 2008: 28. Costa Rica. 

youngi Wirth and Grogan, 1988: 34. Costa Rica. 

Genus CALCARHELEA Wirth and Grogan 

CALCARHELEA Wirth and Grogan, 1988: 35. Type species: Alluaudomyia bimater de Meillon and Hardy, by original 

designation. 

bimater (de Meillon and Hardy), 1953: 25 (Alluaudomyia). South Africa. 

Genus CAMPTOPTEROHELEA Wirth and Hubert 

CAMPTOPTEROHELEA Wirth and Hubert, 1960b: 89. Type species: Camptopterohelea hoogstraali Wirth and Hubert, by 


admirabilis Das Gupta and Sarkar, 1982: 73. India. 

distincta Das Gupta and Sarkar, 1982: 70. India. 

hoogstraali Wirth and Hubert, 1960b: 90. Philippines. 

javanensis Wirth and Wada, 1979: 347. Indonesia. 

tokunagai Wirth and Wada, 1979: 349. Malaysia. 

Genus CAPEHELEA de Meillon and Wirth 

CAPEHELEA de Meillon and Wirth, 1987b: 387. Type species: Capehelea steli de Meillon and Wirth, by original 

designation. 

steli de Meillon and Wirth, 1987b: 388. South Africa. 

Genus CERATOCULICOIDES Wirth and Ratanaworabhan 

CERATOCULICOIDES Wirth and Ratanaworabhan, 1971a: 170. Type species: Helea longipennis Wirth, by original 

designation. 

aliciae Huerta and Borkent, 2005: 112. Mexico. 

blantoni Wirth and Ratanaworabhan, 1971a: 174. USA (Maryland). 

danicus Szadziewski, 1988: 102. Denmark. Eocene. 

longipennis (Wirth), 1952a: 201 (Helea). USA (California). 

moravicus Knoz, 1987: 390. Czech Republic. 

gracilipes (Remm), 1967: 27 (Ceratopogon, preoccupied by Bezzia gracilipes (Winnertz), 1852). Georgia. 

havelkai Wirth and Grogan, 1988: 40. New name for gracilipes. 

remmi Gosseries, 1989: 2. New name for gracilipes. 

tontoeguri (Havelka), 1980: 86 (Ceratopogon). Germany. 

virginianus (Wirth), 1951d: 318 (Helea). USA (Virginia). 

Genus CERATOHELEA Wirth and Grogan 

CERATOHELEA Wirth and Grogan, 1988: 40. Type species: Ceratopogon advena de Meillon, by original designation. 

advena (de Meillon), 1959a: 341 (Ceratopogon). Zimbabwe. 

Genus CERATOPALPOMYIA Szadziewski 

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CERATOPALPOMYIA Szadziewski, 1988: 171. Type species Ceratopalpomyia eocenica Szadziewski, by original 

designation. 

eocenica Szadziewski, 1988: 173. Denmark. Eocene. 

Genus CERATOPOGON Meigen 

HELEA Meigen, 1800: 18. Type species: Ceratopogon communis Meigen, designation by Coquillett, 1910: 549. Generic 

name suppressed by ICZN Opinion 678. 

CERATOPOGON Meigen, 1803: 261. Type species: Tipula barbicornis Linnaeus (misidentified, = Ceratopogon communis 

Meigen), by monotypy. 

CERAPOGON Rafinesque, 1815: 130 (unnecessary new name for Ceratopogon Meigen). Type species: 

Ceratopogon communis Meigen, automatic. 

PSILOHELEA Kieffer, 1915c: 284 (misspelled as Psilohelca), 1917: 294. Type species: Ceratopogon candidatus Winnertz 

(= Ceratopogon niveipennis Meigen), by original designation. 

ANAKEMPIA Kieffer, 1924a: 13. Type species: Dasyhelea grandiforceps Kieffer, by original designation. 

DIPLOHELEA Kieffer, 1925a: 420. Type species: Diplohelea parvula Kieffer (= Ceratopogon grandiforceps Kieffer), by 

original designation). 

abstrusus Borkent and Grogan, 1995: 40. Canada (British Columbia). 

adductus Borkent and Grogan, 1995: 53. Mongolia. 

algidus Borkent and Grogan, 1995: 80. Canada (British Columbia). 

annettae Borkent and Grogan, 1995: 87. Canada (British Columbia). 

arcanus Borkent and Grogan, 1995: 44. USA (Maryland). 

bitterfeldi Szadziewski, 1993: 613. Germany. Eocene. 

boomerangus Borkent and Grogan, 1995: 33. USA (Maryland). 

cavatus Borkent and Grogan, 1995: 67. North Korea. 

ceranowiczi Szadziewski, 1988: 77. Baltic region. Eocene. 

cinaedicus Borkent and Grogan, 1995: 54. USA (Colorado). 

cirrosus Remm, 1974c: 45. Russia. 

communis Meigen, 1804: 27. Germany. 

contextus Yu and Ma, in Yu et al., 2005: 1429. China. 

crypticus Szadziewski, 1988: 70. Baltic region. Eocene. 

culicoidithorax Hoffman, 1926a: 156. USA (New York). 

curtus Borkent and Grogan, 1995: 64. USA (Alaska). 

curvistylus Remm, 1993: 189. Russia. 

denticulatus Borkent and Grogan, 1995: 58. Great Britain. 

dsungaricus Remm, 1980: 101. Kazakhstan. 

elevatus Borkent and Grogan, 1995: 48. Austria. 

eminens Meunier, 1904a: 229 (1904b: 240). Baltic region. Eocene. 

ferulae Remm, 1974c: 43. Kyrgyzstan. 

forcipiformis Meunier, 1904a: 235 (1904b: 246). Baltic region. Eocene. 

defectus Meunier, 1904a: 229 (1904b: 240). Baltic region. Eocene. 

fumipennis Remm, 1974c: 41. Kyrgyzstan. 

gedanicus Szadziewski, 1988: 74. Baltic region. Eocene. 

gigaforceps Remm, 1973: 180. Russia. 

grandiforceps (Kieffer), 1913a: 7 (Dasyhelea). Great Britain. 

parvula (Kieffer), 1925a: 420 (Diplohelea). Great Britain. 

grogani Szadziewski, 1988: 64. Baltic region. Eocene. 

hennigi Szadziewski, 1988: 60. Baltic region. Eocene. 

inverecundus Borkent and Grogan, 1995: 46. Canada (British Columbia). 

kolensis Remm, 1993: 189. Russia. 

kotejai Szadziewski, 1993: 616. Germany. Eocene. 

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kurilensis Remm, 1974c: 41. Russia. 

lacteipennis Zetterstedt, 1838: 820. Norway. 

edentatus (Edwards), 1921: 124 (Psilohelea). Great Britain. 

sphagnicola (Kieffer), 1924b: 403 (Anakempia). Great Britain. 

conjunctus (Kieffer), 1925a: 416 (Anakempia). Great Britain. 

longitarsis (Mayer), 1940: 164 (Helea). Great Britain. 

magniforceps (Kieffer), 1925a: 413 (Trishelea). Russia. 

mallochi (Cole), in Cole and Lovett, 1921: 213 (Hartomyia). USA (Oregon). 

mammulus Borkent and Grogan, 1995: 65. Canada (Yukon Territory). 

margaritae Szadziewski, 1988: 80. Baltic region. Eocene. 

miocaenicus Szadziewski, 1993: 617. Germany. Eocene. 

multisetosus Borkent and Grogan, 1995: 76. Canada (Nova Scotia). 

naccinervis Borkent, in Borkent and Wirth, 1997:97. New name for crassinervis. 

crassinervis (Goetghebuer), 1920: 62 (Stilobezzia, preoccupied by Palpomyia crassinervis (de Meijere), 1907). 

Belgium. 

nanalobus Borkent and Grogan, 1995: 97. Baltic region. Eocene. 

niveipennis Meigen, 1818: 73. Germany. 

candidatus Winnertz, 1852: 57. Netherlands. 

pallicoleatus Borkent and Grogan, 1995: 83. USA (Oregon). 

pandus Borkent and Grogan, 1995: 53. USA (Wisconsin). 

paraeminens Borkent and Grogan, 1995: 99. Baltic region. Eocene. 

paucisetosus Remm, 1974c: 43. Estonia. 

piotrowskii Szadziewski, 1988: 75. Baltic region. Eocene. 

pisinnus Borkent and Grogan, 1995: 97. Baltic region. Eocene. 

pontis Borkent and Grogan, 1995: 56. USA (Kansas). 

pubiantennalis Borkent and Grogan, 1995: 74. Canada (Northwest Territories). 

rectus Borkent and Grogan, 1995: 84. USA (Colorado). 

relictus Borkent and Grogan, 1995: 32. USA (California). 

remmicola Szadziewski, 1988: 71. Baltic region. Eocene. 

ritzkowskii Szadziewski, 1988: 78. Baltic region. Eocene. 

romanicus Damian-Georgescu, 1972: 19. Romania. 

seculus Borkent and Grogan, 1995: 79. USA (Maryland). 

subeminens Szadziewski, 1993: 617. Germany. Eocene. 

succinicola Szadziewski, 1993: 620. Germany. Eocene. 

tertiaricus Szadziewski, 1988: 62. Baltic region. Eocene. 

unguis (Tokunaga), 1940b: 255 (Culicoides). Japan. 

willisi Borkent and Grogan, 1995: 81. USA (Maryland). 

Genus CHAIROPOGON Yu 

CHAIROPOGON Yu, in Yu et al., 2005: 1430. Type species: Chairopogon chengdeiensis Yu and Hao, by original 

designation. 

chengdeiensis Yu and Hao, in Yu et al., 2005: 1430. China. 

Genus CONGOHELEA Wirth and Grogan 

CONGOHELEA Wirth and Grogan, 1988: 46. Type species: Serromyia fuligipennis Clastrier, by original designation. 

fuligipennis (Clastrier), 1960b: 291 (Serromyia). Congo. 

Genus DIAPHANOBEZZIA Ingram and Macfie 

DIAPHANOBEZZIA Ingram and Macfie, 1931a: 223. Type species: Diaphanobezzia pellucida Ingram and Macfie, by 

120


araucaria Spinelli, 1996: 77. Argentina. 

patagonica Spinelli and Grogan, 1990: 127. Argentina. 

pellucida Ingram and Macfie, 1931a: 223. Argentina. 

spinellii Wirth and Grogan, 1988: 48. Argentina. 

Genus DOWNESHELEA Wirth and Grogan 

DOWNESHELEA Wirth and Grogan, 1988: 50. Type species: Monohelea stonei Wirth, by original designation. 

balboa (Lane and Wirth), 1964: 225 (Monohelea). Panama. 

bicornis Felippe-Bauer and Quintelas, 1993b: 185. Brazil. 

bimaculata Clastrier and Delécolle, 1990: 148. Ivory Coast. 

blantoni (Lane and Wirth), 1964: 217 (Monohelea). Panama. 

carioca (Tavares and Silva Pereira), 1978: 157 (Monohelea). Brazil. 

castroi (Tavares and Silva Pereira), 1978: 159 (Monohelea). Brazil. 

cebacoi (Lane and Wirth), 1964: 218 (Monohelea). Panama. 

charrua Felippe-Bauer and Spinelli, 1994: 161. Uruguay. 

chiapasi (Lane and Wirth), 1964: 219 (Monohelea). Nicaragua. 

chirusi (Lane and Wirth), 1964: 218 (Monohelea). Panama. 

colombiae (Lane and Wirth), 1964: 220 (Monohelea). Colombia. 

deanei Felippe-Bauer and Quintelas, in Felippe-Bauer et al., 1995: 395. Trinidad. 

fluminensis Felippe-Bauer and Quintelas, 1993a: 33. Brazil. 

fuscipennis (Lane and Wirth), 1964: 221 (Monohelea). Colombia. 

grogani Huerta, Felippe-Bauer, and Spinelli, 2012: 64. Mexico. 

guianae (Wirth), 1953b: 150 (Monohelea). Guyana. 

lanei Felippe-Bauer and Borkent in Felippe-Bauer et al., 2011: 21. Brazil. 

leei (Debenham), 1972: 16 (Monohelea). Australia (New South Wales). 

macclurei (Ratanaworabhan and Wirth), 1972: 448 (Monohelea). Malaysia. 

mcdanieli (Tokunaga), in Tokunaga and Murachi, 1959: 411 (Monohelea). Belau (USA). 

multilineata (Lutz), 1914: 93 (Palpomyia). Brazil. 

nigeriae (Ingram and Macfie), 1922: 268 (Monohelea). Nigeria. 

nigra (Tokunaga), 1963a: 243 (Monohelea). Papua New Guinea. 

notialisinica Yu and Hao, in Yu et al., 2005: 1432. China. 

nubeculosa (Macfie), 1932c: 45 (Monohelea). New Zealand 

oliveirai Felippe-Bauer, in Felippe-Bauer and Silva, 2008: 400. Brazil. 

panamensis (Lane and Wirth), 1964: 221 (Monohelea). Panama. 

quasidentica Felippe-Bauer and Quintelas, 1993a: 37. Brazil. 

scanloni (Ratanaworabhan and Wirth), 1972: 449 (Monohelea). Thailand. 

sepikensis (Debenham), 1972: 13 (Monohelea). Papua New Guinea. 

stenochora Wirth and Giles, 1990: 446. Fiji. 

stonei (Wirth), 1953b: 148 (Monohelea). USA (Louisiana). 

unimaculata (Debenham), 1972: 5 (Monohelea). Australia (New South Wales). 

whartoni (Ratanaworabhan and Wirth), 1972: 451 (Monohelea). Malaysia. 

xanthogonua (Tokunaga), 1963a: 242 (Monohelea). Papua New Guinea. 

Genus ECHINOHELEA Macfie 

ECHINOHELEA Macfie, 1940c: 187. Type species: Echinohelea ornatipennis Macfie, by original designation. 

Subgenus ECHINOHELEA Macfie 

australiensis Debenham, 1970c: 152. Australia (Capital Territory). 

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lantoni Wirth, 1994c: 232. Panama. 

flava Tokunaga, 1963a: 235. Papua New Guinea. 

harbelensis de Meillon, 1960: 409. Liberia. 

hardyi Tokunaga, 1963a: 235. Papua New Guinea. 

jamaicensis Wirth, 1994c: 234. Jamaica. 

laensis Tokunaga, 1963a: 236. Papua New Guinea. 

lanei Wirth, 1951d: 319. USA (Virginia). 

leei Wirth, 1994c: 237. Colombia. 

longirostris Debenham, 1970c: 147. Papua New Guinea. 

macfiei Lane, 1948: 228. Brazil. 

moresbyensis Debenham, 1970c: 149. Papua New Guinea. 

neotropica Wirth, 1994c: 237. Colombia. 

notipes Debenham, 1970c: 152. Australia (Queensland). 

ornata Clastrier, 1984c: 368. Guinea. 

ornatipennis Macfie, 1940c: 188. Guyana. 

pallida Debenham, 1970c: 155. Papua New Guinea. 

panamensis Wirth, 1994c: 239. Panama. 

papuensis Tokunaga, 1966a: 112. Papua New Guinea. 

pastoriana Clastrier, 1984c: 362. Guinea. 

pictipennis Tokunaga, 1963a: 231. Papua New Guinea. 

richardsi Macfie, 1940c: 189. Guyana. 

smarti Macfie, 1940c: 190. Guyana. 

vicina Clastrier, 1984c: 366. Guinea. 

voltana de Meillon, 1959b: 15. Burkina Faso. 

zonata Tokunaga, 1963a: 234. Papua New Guinea. 

Subgenus ECHINOIDESHELEA Wirth 

ECHINOIDESHELEA Wirth, 1994c: 231 (as subgenus of Echinohelea). Type species: Echinohelea aitkeni Wirth, by original 

designation. 

aitkeni Wirth, 1994c: 231. Brazil. 

Genus EOHELEA Petrunkevitch 

EOHELEA Petrunkevitch, 1957: 208. Type species: Eohelea stridulans Petrunkevitch (= Ceratopogon sinuosus Meunier), 


fossicola Szadziewski, 1993: 624. Germany. Eocene. 


grogani Szadziewski, 1988: 164. Poland. Eocene. 

miocaenea Szadziewski, 1993: 625. Germany. Eocene. 

petrunkevitchi Szadziewski, 1984b: 39 (1985b: 127). Poland. Eocene. 

sakhalinica Szadziewski, 1990a: 78. Russia. Tertiary. 

sinuosa (Meunier), 1904a: 234 (1904b: 245) (Ceratopogon). Baltic region. Eocene. 

stridulans Petrunkevitch, 1957: 209. Baltic region. Eocene. 

Genus FANTHAMIA de Meillon 

FANTHAMIA de Meillon, 1939c: 103 (as subgenus of Ceratopogon). Type species: Ceratopogon adulator de Meillon, by 


abdita de Meillon and Wirth, 1987a: 46. South Africa. 

adulator (de Meillon), 1939c: 104 (Ceratopogon). South Africa. 

122

aniculae de Meillon and Downes, 1986: 167. South Africa. 

cardinis de Meillon and Wirth, 1979c: 194. Zimbabwe. 

carina (de Meillon and Hardy), 1953: 28 (Ceratopogon). South Africa. 

draconis de Meillon and Downes, 1986: 170. South Africa. 

forsteri de Meillon and Downes, 1986: 169. South Africa. 

gigantulus de Meillon and Wirth, 1979c: 196. South Africa. 

imperfecta (Goetghebuer), 1935d: 172 (Culicoides). Congo. 

jonkeri de Meillon and Wirth, 1987a: 48. South Africa. 

mennoi de Meillon and Wirth, 1981c: 582. South Africa. 

montana de Meillon and Downes, 1986: 167. South Africa. 

ornatipennis (de Meillon), 1939c: 106 (Ceratopogon). South Africa. 

sani de Meillon and Downes, 1986: 168. South Africa. 

Genus FITTKAUHELEA Wirth and Blanton 

FITTKAUHELEA Wirth and Blanton, 1970a: 7. Type species: Fittkauhelea amazonica Wirth and Blanton, by original 

designation. 

amazonica Wirth and Blanton, 1970a: 9. Brazil. 

Genus FOSSIHELEA Szadziewski 

FOSSIHELEA Szadziewski, 1988: 104. Type species: Ceratopogon gracilitarsis Meunier, by original designation. 

gracilitarsis (Meunier), 1904a: 235 (1904b: 246) (Ceratopogon). Baltic region. Eocene. 


Genus GEDANOHELEA Szadziewski 

GEDANOHELEA Szadziewski, 1988: 166. Type species: Gedanohelea loewi Szadziewski, by original designation. 

loewi Szadziewski, 1988: 168. Poland. Eocene. 


wirthi Szadziewski, 1988: 170. Denmark. Eocene. 

Genus HELEAGERON Borkent 

HELEAGERON Borkent, 1995: 47. Type species: Heleageron arenatus Borkent, by original designation. 

arenatus Borkent, 1995: 48. Canada (Alberta). Upper Cretaceous. 

grimaldii Borkent, 1996: 15. USA (New Jersey). Upper Cretaceous. 

Genus HETEROCERATOPOGON Wirth and Grogan 

HETEROCERATOPOGON Wirth and Grogan, 1988: 55. Type species: Heteroceratopogon poguei Wirth and Grogan, by 


poguei Wirth and Grogan, 1988: 57. New Caledonia (France). 

Genus HETEROHELEA Clastrier 

HETEROHELEA Clastrier, 1985b: 272. Type species: Heterohelea minutissima Clastrier, by original designation. 

minutissima Clastrier, 1985b: 272. New Caledonia (France). 

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Genus HYPSIMYIA Yu 

HYPSIMYIA Yu, in Yu et al., 2005: 1433. Type species: Hypsimyia emeiensis Yu and Li, by original designation. 

emeiensis Yu and Li, in Yu et al., 2005: 1433. China. 

Genus ISTHMOHELEA Ingram and Macfie 

ISTHMOHELEA Ingram and Macfie, 1931a: 208. Type species: Isthmohelea disjuncta Ingram and Macfie, by original 

designation. 

disjuncta Ingram and Macfie, 1931a: 209. Chile. 

Genus KOLENOHELEA de Meillon and Wirth 

KOLENOHELEA de Meillon and Wirth, 1981a: 513. Type species: Kolenohelea dycei de Meillon and Wirth, by original 

designation. 

calcarata (Goetghebuer), 1920: 64 (Monohelea). Belgium. 

sharpi (Edwards), 1929a: 427 (Stilobezzia). Great Britain. 

fusca (Goetghebuer), 1932a: 125 (Stilobezzia). Belgium. 

dycei de Meillon and Wirth, 1981a: 517. South Africa. 

equilis de Meillon and Wirth, 1987a: 48. South Africa. 

fuscipennis Clastrier, 1984e: 369. Guinea. 

inusitata de Meillon and Wirth, 1981c: 586. South Africa. 

jarmilae de Meillon and Downes, 1986: 172. South Africa. 

kindiae Clastrier, 1984e: 366. Guinea. 

leonina de Meillon and Wirth, 1987a: 50. South Africa. 

luciae de Meillon and Wirth, 1983a: 361. South Africa. 

milleri de Meillon and Wirth, 1981a: 521. South Africa. 

mira de Meillon and Wirth, 1981c: 583. South Africa. 

mirabunda de Meillon and Wirth, 1981a: 522. South Africa. 

mirifica de Meillon and Wirth, 1981a: 519. South Africa. 

monticola de Meillon and Downes, 1986: 174. South Africa. 

ornata Clastrier, 1984e: 367. Guinea. 

pedi de Meillon and Wirth, 1987a: 50. South Africa. 

pugilator de Meillon and Wirth, 1987a: 52. South Africa. 

uysorum de Meillon, Meiswinkel and Wirth, 1982: 138. South Africa. 

Genus LEPTOHELEA Wirth and Blanton 

LEPTOHELEA Wirth and Blanton, 1970a: 12. Type species: Leptohelea icronyx Wirth and Blanton, by original designation. 

micronyx Wirth and Blanton, 1970a: 12. Colombia. 

Genus LUCIAMYIA de Meillon 

LUCIAMYIA de Meillon, 1937b: 380. Type species: Luciamyia biloba de Meillon, by original designation. 

biloba de Meillon, 1937b: 380. South Africa. 

irrita (de Meillon), 1937b: 383 (Monohelea). South Africa. 

Genus MACRUROHELEA Ingram and Macfie 

124

MACRUROHELEA Ingram and Macfie, 1931a: 203. Type species: Macrurohelea caudata Ingram and Macfie, by original 

designation. 

caudata Ingram and Macfie, 1931a: 205. Chile. 

commoni Lee, 1963: 339. Australia (New South Wales). 

dycei Grogan and Wirth, 1985: 133. Australia (New South Wales). 

fuscipennis Spinelli and Grogan, 1990: 128. Argentina. 

gentilii Spinelli and Grogan, 1984: 963. Argentina. 

irwini Grogan and Wirth, 1980b: 140. Chile. 

kuscheli Wirth, 1965c: 49. Chile. 

leei Grogan and Wirth, 1985: 131. Australia (Queensland). 

monotheca Spinelli and Grogan, 1984: 965. Argentina. 

paracaudata Grogan and Wirth, 1980b: 141. Chile. 

setosa Wirth, 1965c: 49. Chile. 

similis Spinelli and Grogan, 1990: 131. Argentina. 

thoracica Ingram and Macfie, 1931a: 206. Argentina. 

wirthi Spinelli and Grogan, 1984: 965. Argentina. 

yamana Spinelli and Grogan, 1999: 710. Argentina. 

Genus METACANTHOHELEA Wirth and Grogan 

METACANTHOHELEA Wirth and Grogan, 1988: 65. Type species: Metacanthohelea cogani Wirth and Grogan, by original 

designation. 

cogani Wirth and Grogan, 1988: 66. Aldabra (Seychelles). 

Genus MANTOHELEA Szadziewski 

MANTOHELEA Szadziewski, 1988: 144. Type species: Ceratopogon lacus Meunier, by original designation. 


laca (Meunier), 1904a: 232 (1904b: 243) (Ceratopogon). Baltic region. Eocene. 

Genus MEUNIEROHELEA Szadziewski 

MEUNIEROHELEA Szadziewski, 1988: 148. Type species: Meunierohelea nielseni Szadziewski, by original designation. 

CHIMAEROHELEA Debenham, 1988: 801. Type species: Chimaerohelea caligula Debenham, by original designation. 

caligula (Debenham), 1988: 803 (Chimaerohelea). Australia (Queensland). 

gedanicola Szadziewski, 1988: 153. Poland. Eocene. 

miocaenica (Szadziewski), 1993: 622 (Chimaerohelea). Germany. Eocene. 

nielseni Szadziewski, 1988: 150. Denmark. Eocene. 

wirthi Szadziewski, 1988: 157. Denmark. Eocene. 

Genus MONOHELEA Kieffer 

MONOHELEA Kieffer, 1917b: 294. Type species: Monohelea hieroglyphica Kieffer, by original designation. 

accipiter Debenham, 1972: 34. Australia (New South Wales). 

affinis Felippe-Bauer and Spinelli, 1991: 201. Brazil. 

aguirrei Tavares and Sousa, 1980: 97. Brazil. 

albiclavatoris Tokunaga and Murachi, 1959: 407. Micronesia. 

125

andersoni Wirth and Grogan, 1981: 30. USA (Maryland). 

andrei Vattier and Adam, 1966b: 744. Congo. 

annulata Debenham, 1972: 18. Australia (Northern Territory). 

archibaldoi Tavares and Sousa, 1980: 98. Brazil. 

avicularis Remm, 1993: 185. Azerbaijan. 

baltica Szadziewski, 1988: 130. Poland. Eocene. 

beaveri Wirth and Giles, 1990: 448. Fiji. 

bidens Debenham, 1972: 35. Australia (Capital Territory). 

bidentata Felippe-Bauer and Spinelli, 1994: 163. Argentina. 

bifurcata Wirth and Williams, 1964: 308. USA (Massachusetts). 

boucheti Clastrier, 1993b: 161. New Caledonia (France). 

boudinoti Clastrier, 1993b: 160. New Caledonia (France). 

brasiliensis Lane, 1948: 226. Brazil. 

candescens Debenham, 1972: 25. Australia (New South Wales). 

chalybeata Clastrier, 1960b: 286. Congo. 

chazeaui Clastrier, 1985d: 752. New Caledonia (France). 

clavipes Macfie, 1932c: 48. New Zealand. 

clunipes (Loew), 1850: 30 (Ceratopogon). Baltic region. Eocene. 

cognata Clastrier, 1960b: 289. Congo. 

coloisuvae Wirth and Giles, 1990: 449. Fiji. 

cunasi Lane and Wirth, 1964: 232. Panama. 

dorsotaeniata Clastrier, 1985d: 756. New Caledonia (France). 

estonica Remm, 1965: 182. Estonia. 

fairchildi Lane and Wirth, 1964: 214. Panama. 

falcata Debenham, 1972: 7. Australia (New South Wales). 

fijiensis Wirth and Giles, 1990: 451. Fiji. 

floridensis Wirth and Williams, 1964: 309. USA (Florida). 

gemina Debenham, 1972: 39. Australia (New South Wales). 

grogani Delécolle and Rieb, 1995: 34. USA (Arkansas). 

guaimiesi Lane and Wirth, 1964: 227. Panama. 

hainanena Liu, Yan and Liu, 1996: 45. China. 

hainanensis Liu and Yan, 1996: 19. China 

hieroglyphica Kieffer, 1917b: 312. Paraguay. 

hirsuta Wirth and Grogan, 1981: 36. USA (Maryland). 

hissarica Remm, 1980: 91. Tajikistan. 

hyalinipennis Clastrier, 1993b: 162. New Caledonia (France). 

impunctatipennis Clastrier, 1985d: 748. New Caledonia (France). 

inflatistyla Debenham, 1972: 30. Australia (New South Wales). 

jamnbacki Wirth and Williams, 1964: 308. USA (Michigan). 

knighti Wirth and Williams, 1964: 305. USA (Florida). 

lanei Wirth, 1953b: 142. USA (Florida). 

legrandi Clastrier, 1984a: 49. Gabon. 

leveri Wirth and Giles, 1990: 452. Fiji. 

lutea Tokunaga, 1963a: 241. Papua New Guinea. 

macfiei Wirth, 1953b: 143. USA (Louisiana). 

maculipennis (Coquillett), 1905: 64 (Ceratopogon). USA (Florida). 

magnitheca Wirth and Grogan, 1981: 42. USA (Maryland). 

maya Felippe-Bauer, Huerta and Ibáñez-Bernal, 2000: 815. Mexico. 

mayeri Ortiz, 1950d: 202. Venezuela. 

minuscula Clastrier, 1985d: 758. New Caledonia (France). 

neocaledonica Clastrier, 1993b: 158. New Caledonia (France). 

novaguinensis Tokunaga, 1959: 309. Papua New Guinea. 

obscura Wirth and Williams, 1964: 304. USA (Massachusetts). 

orientalis Ratanaworabhan and Wirth, 1972: 444. Thailand. 

126

ornata Wirth, 1953b: 144. USA (Florida). 

ornatissima Clastrier, 1985d: 750. New Caledonia (France). 

ornatithorax Clastrier and Delécolle, 1990: 153. Guinea. 

pahangensis Ratanaworabhan and Wirth, 1972: 443. Malaysia. 

palauensis Tokunaga, in Tokunaga and Murachi, 1959: 408. Belau (USA). 

pallida Clastrier and Delécolle, 1990: 151. Guinea. 

palustria Debenham, 1972: 28. Australia (Queensland). 

paranigripes Saha and Das Gupta, 1991: 6. India. 

periucunda de Meillon and Wirth, 1981c: 587. South Africa. 

phoenix Debenham, 1972: 38. Australia (New South Wales). 

polychroma Clastrier, 1958b: 252. Senegal. 

poncai Lane and Wirth, 1964: 228. Panama. 

ponticulifera Debenham, 1972: 38. Australia (Capital Territory). 

praeclara Goetghebuer, 1935d: 178. Democratic Republic of the Congo. 

pseudochelagonata Saha and Das Gupta, 1991: 8. India. 

radialis Debenham, 1972: 22. Australia (Queensland). 

roraimensis Felippe-Bauer and Spinelli, 1991: 202. Brazil. 

scirpi (Kieffer), 1901a: 157 (Palpomyia). France. 

scutofasciata Clastrier, 1985d: 754. New Caledonia (France). 

sinica Yu and Deng, in Yu et al., 2005: 1435. China. 

smeei Tokunaga, 1963a: 239. Papua New Guinea. 

subpahangensis Saha and Das Gupta, 1991: 7. India. 

texana Wirth, 1953b: 143. USA (Texas). 

tigrina (Skuse), 1889: 306 (Ceratopogon). Australia (New South Wales). 

transversalis Clastrier, 1960b: 290. Congo. 

umbrosipennis Debenham, 1972: 32. Australia (Queensland). 

unica Sinha and Das Gupta, 1992: 26. India. 

urracaisi Lane and Wirth, 1964: 231. Panama. 

uruguayensis Felippe-Bauer and Spinelli, 1998: 63. Uruguay. 

ussurica Remm, 1971: 208. Russia. 

wirthi Khalaf, 1969: 407. USA (Mississippi). 

Genus NANNOHELEA Grogan and Wirth 

NANNOHELEA Grogan and Wirth, 1980a: 374. Type species: Ceratopogon bourioni Clastrier, by original designation. 

bourioni (Clastrier), 1961a: 424 (Ceratopogon). Algeria. 

clastrieri Grogan and Wirth, 1980a: 381. Colombia. 

eocenica Szadziewski, 1988: 101. Denmark. Eocene. 

fuscipennis (Tokunaga), 1964a: 298 (Ceratopogon). Papua New Guinea. 

grogani Szadziewski, 1988: 99. Denmark. Eocene. 

tamil Grogan and Wirth, 1990: 348. Sri Lanka. 

Genus NEOHELEA Clastrier 

NEOHELEA Clastrier, 1988a: 53. Type species: Neohelea pastoriana Clastrier, by original designation. 

pastoriana Clastrier, 1988a: 55. Guinea. 

Genus NEUROBEZZIA Wirth and Ratanaworabhan 

NEUROBEZZIA Wirth and Ratanaworabhan, 1972b: 244. Type species: Bezzia granulosa Wirth, by original designation. 

MEDEOBEZZIA Yu, 1995: 287 (also as Meddeobezzia, Medeobizzia). Type species: Medeobezzia singularis Yu, by 

monotypy. 

127

granulosa (Wirth), 1952a: 240 (Bezzia). USA (California). 

singularis (Yu), 1995: 287 (Medeobezzia). China. 

tsacasi Clastrier, 1983c: 64. Seychelles. 

Genus NOTIOHELEA Grogan and Wirth 

NOTIOHELEA Grogan and Wirth, 1979c: 283. Type species: Notiohelea chilensis Grogan and Wirth, by original 

designation. 

chilensis Grogan and Wirth, 1979c: 284. Chile. 

pilosa Spinelli and Grogan, 1990: 133. Argentina. 

Genus NOTOCERATOPOGON de Meillon and Downes 

NOTOCERATOPOGON de Meillon and Downes, 1986: 157. Type species: Notoceratopogon minutus de Meillon and 

Downes, by original designation. 

alcides (de Meillon and Hardy), 1954: 74 (Ceratopogon). South Africa. 

minutus de Meillon and Downes, 1986: 161. South Africa. 

natalensis (de Meillon), 1937b: 378 (Ceratopogon). South Africa. 

vockerothi de Meillon and Downes, 1986: 162. South Africa. 

Genus OXYRIA Yu 

OXYRIA Yu, in Yu et al., 2005: 1436. Type species: Oxyria xui Yu, by original designation. 

xui Yu, in Yu et al., 2005: 1437. China. 

Genus PALAEOBRACHYPOGON Borkent 

PALAEOBRACHYPOGON Borkent, 1995: 70. Type species: Palaeobrachypogon vetus Borkent, by original designation. 

aquilonius (Boesel), 1937: 49 (Ceratopogon). Canada (Manitoba). Upper Cretaceous. 

grandiforceps Borkent, 2000b: 463. USA (New Jersey). Upper Cretaceous. 

macronyx (Remm), 1976b: 113 (Ceratopogon). Russia. Upper Cretaceous. 

remmi Borkent, 1995: 76. Canada (Alberta). Upper Cretaceous. 

vetus Borkent, 1995: 78. Canada (Alberta). Upper Cretaceous. 

Genus PARABEZZIA Malloch 

PARABEZZIA Malloch, 1915a: 358. Type species: Parabezzia petiolata Malloch, by original designation. 

alexanderi Wirth, 1965b: 219. USA (Massachusetts). 

arenosa Clastrier and Raccurt, 1979a: 172. Haiti. 

atchleyi Grogan and Wirth, 1977a: 67. USA (Texas). 

balseiroi Spinelli and Grogan, 1987: 13. Argentina. 

blantoni Wirth, 1965b: 220. Panama. 

brasiliensis Spinelli and Grogan, 1987: 34. Brazil. 

brunnea Wirth, 1965b: 220. Panama. 

bystraki Grogan and Wirth, 1977a: 70. USA (Maryland). 

caribbeana Clastrier and Raccurt, 1979a: 173. Haiti. 

cayoensis Spinelli and Grogan, 1987: 36. Belize. 

clastrieri Spinelli and Grogan, 1987: 23. El Salvador. 

128

costalis Wirth, 1965b: 221. Panama. 

downesi Wirth, 1965b: 222. Canada (Ontario). 

eupetiolata Grogan and Wirth, 1977a: 79. USA (New York). 

falcipennis Clastrier, 1960b: 293. Congo. 

insolita Vattier and Adam, 1966b: 769. Congo. 

fuscipennis Wirth, 1965b: 223. Panama. 

grogani Szadziewski and Wirth, 1983: 359. Algeria. 

haitiensis Clastrier and Raccurt, 1979a: 170. Haiti. 

hondurensis Spinelli and Grogan, 1987: 25. Honduras. 

horvathi Grogan and Wirth, 1977a: 81. USA (Florida). 

huberti Grogan and Wirth, 1977a: 75. USA (Maryland). 

inaequalis Spinelli and Grogan, 1987: 38. Brazil. 

inermis (Coquillett), 1902a: 86 (Ceratopogon). USA (Arizona). 

jamaicensis Wirth, 1965b: 224. Jamaica. 

neunguis Grogan and Wirth, 1977a: 65. USA (Virginia). 

obscura de Meillon and Wirth, 1981c: 588. South Africa. 

orientalis Giles and Wirth, 1982b: 823. Sri Lanka. 

pallida Spinelli and Grogan, 1987: 40. Mexico. 

panamensis Wirth, 1965b: 225. Panama. 

petiolata Malloch, 1915a: 359. USA (Illinois). 

pseudunguis Spinelli and Grogan, 1987: 30. Panama. 

raccurti Spinelli and Grogan, 1987: 42. El Salvador. 

spangleri Wirth, 1965b: 227. Puerto Rico (USA). 

stagni de Meillon and Wirth, 1981b: 538. South Africa. 

texensis Grogan and Wirth, 1977a: 60. USA (Texas). 

uncinata (Johannsen), 1943: 761 (Stilobezzia). USA (Alabama). 

unguis Wirth, 1965b: 228. USA (Arizona). 

williamsi Wirth, 1965b: 229. USA (Michigan). 

wirthi Clastrier and Raccurt, 1979a: 169. Haiti. 

Genus PARALLUAUDOMYIA Clastrier 

PARALLUAUDOMYIA Clastrier, 1960b: 268. Type species: Paralluaudomyia maculata Clastrier, by original designation. 

maculata Clastrier, 1960b: 269. Congo. 

Genus PARASTILOBEZZIA Wirth and Blanton 

PARASTILOBEZZIA Wirth and Blanton, 1970a: 10. Type species: Parastilobezzia leei Wirth and Blanton, by original 

designation. 

leei Wirth and Blanton, 1970a: 10. Colombia. 

Genus PERONEHELEA Borkent 

PERONEHELEA Borkent, 1995: 80. Type species: Peronehelea chrimikalydia Borkent, by original designation. 

chrimikalydia Borkent, 1995: 82. Canada (Alberta). Upper Cretaceous. 

frigidus (Remm), 1976b: 115 (Ceratopogon). Russia. Upper Cretaceous. 

sibirica Szadziewski, 1996: 69. Russia. Upper Cretaceous. 

Genus PSEUDOSTILOBEZZIA Wirth and Ratanaworabhan 

PSEUDOSTILOBEZZIA Wirth and Ratanaworabhan, 1973: 177. Type species: Pseudostilobezzia macclurei Wirth and 

129

Ratanaworabhan, by original designation. 

macclurei Wirth and Ratanaworabhan, 1973: 178. Viet Nam. 

wirthi Yu and Yan, 2003: 238. China. 

Genus RHYNCHOHELEA Wirth and Blanton 

RHYNCHOHELEA Wirth and Blanton, 1970c: 96. Type species: Rhynchohelea monilicornis Wirth and Blanton, by original 

designation. 

monilicornis Wirth and Blanton, 1970c: 98. USA (Florida). 

Genus SCHIZOHELEA Kieffer 

SCHIZOHELEA Kieffer, 1917b: 295. Type species: Ceratopogon copiosus Winnertz (= Ceratopogon leucopeza Meigen), by 

monotypy. 

KIEFFEROMYIA Mayer, 1937a: 303. Type species: Kiefferomyia gorana Mayer (= Ceratopogon leucopeza Meigen), by 


armata (Remm), 1993: 182 (Monohelea). Russia. 

incerta (Clastrier), 1963: 60 (Monohelea). Algeria. 

lampropeza (Remm), 1967: 29 (Monohelea). Georgia. 

leucopeza (Meigen), 1804: 29 (Ceratopogon). Europe. 

albitarsis (Wiedemann), 1817: 67. (Ceratopogon). Germany. 

copiosa (Winnertz), 1852: 56 (Ceratopogon). Germany. 

polita (Coquillett), 1901a: 606 (Ceratopogon). USA (Massachusetts). 

stecki (Kieffer), 1915b: 65 (Bezzia). Switzerland. 

gorana (Mayer), 1937a: 303 (Kiefferomyia). Germany. 

xanthopeza (Clastrier), 1963: 55 (Monohelea). Belgium, Norway, Finland. 

pekae (Remm), 1980: 89 (Monohelea). Kyrgyzstan. 

spathulata (Remm), 1993: 183 (Monohelea). Armenia. 

Genus SCHIZONYXHELEA Clastrier 

SCHIZONYXHELEA Clastrier, 1984b: 1. Type species: Schizonyxhelea guyana Clastrier, by original designation. 

forattinii Wirth and Grogan, 1988: 81. Brazil. 

guyana Clastrier, 1984b: 2. French Guiana. 

Genus SERROMYIA Meigen 

SERROMYIA Meigen, 1818: 83. Type species: Ceratopogon femoratus Meigen, by monotypy. 

PRIONOMYIA Stephens, 1829b: 237. Type species: Ceratopogon femoratus Meigen, designation by Westwood, 1840: 126. 

ATMOBIA Bigot, 1857: 519. Type species: Ceratopogon femoratus Meigen, designation by Evenhuis and Pont, 2004:15. 

CERATOLOPHUS Kieffer, 1899: 69, (preoccupied by Barboza de Bocage, 1873). Type species: Ceratopogon femoratus 

Meigen , by original designation. 

JOHANNSENIELLA Williston, 1907: 1. New name for Ceratolophus Kieffer. Type species: Ceratopogon femoratus Meigen, 

automatic. 

CERATOLOPHANA Strand, 1928: 48. New name for Ceratolophus Kieffer. Type species: Ceratopogon femoratus Meigen, 

automatic. 

aethiopiae Clastrier and Wirth, 1961a: 219. Gambia. 

agathae de Meillon and Wirth, 1983b: 388. South Africa. 

alphea (Heyden), 1870: 251 (Ceratopogon). Germany. Eocene. 

130

austera Statz, 1944: 150. Germany. Oligocene. 

colorata Statz, 1944: 150. Germany. Oligocene. 

spinofemorata Statz, 1944: 151. Germany. Oligocene. 

anomalicornis (Loew), 1850: 30 (Ceratopogon). Baltic region. Eocene. 

arabica Szadziewski, Gwizdalska-Kentzer and Gilka, 2011: 648. United Arab Emirates. 

atra (Meigen), 1818: 84 (Ceratopogon). Latvia. 

albitarsis Kieffer, 1919a: 71. Hungary. 

micronyx Kieffer, 1919a: 70. Hungary. 

spinosipes Kieffer, 1919a: 72. Hungary. 

nitens Goetghebuer, 1920: 73. Belgium. 

barberi Wirth, 1952a: 205. USA (California). 

bicolor Borkent, in Borkent and Bissett, 1990: 196. Germany. 

borealis Borkent, in Borkent and Bissett, 1990: 186. Canada (Alberta). 

crassifemorata Malloch, 1914a: 218. USA (Illinois). 


femorata (Meigen), 1804: 28 (Ceratopogon). Europe. 

armata (Meigen), 1818: 83 (Ceratopogon). Germany. 

foersteri (Meigen), 1838: 21 (Ceratopogon). Great Britain. 

flavipes (Gimmerthal), 1847: 144 (Ceratopogon, preoccupied by Palpomyia flavipes (Meigen), 1804). Great Britain. 

flavicrus (Kieffer), 1906a: 63 (Palpomyia). New name for flavipes. 

inermipes Kieffer, 1919a: 73. Great Britain. 

festiva Kieffer, 1911c: 346. Seychelles. 

flaviventris Remm, 1993: 182. Russia. 

hainana Yu and Yan, 2002: 168. China. 

heveli Giles and Wirth, 1982a: 442. Sri Lanka. 

ledicola Kieffer, 1925b: 156. Estonia. 

europaea Clastrier, 1963: 61. Austria. 

macronyx Goetghebuer, 1933c: 355. Belgium. 

maculipennis Giles and Wirth, 1982a: 444. Sri Lanka. 

mangrovi Delécolle and Braverman, 1987: 57. Egypt. 

meiswinkeli de Meillon and Wirth, 1983b: 390. South Africa. 

morio (Fabricius), 1775: 800 (Culex). Great Britain. 

nudipennis Kieffer, 1913a: 10. Great Britain. 

neethlingi de Meillon and Wirth, 1983b: 392. South Africa. 

nocticolor Kieffer, 1914b: 268. South Africa. 

armipes (de Meillon), 1959a: 343 (Ceratopogon, preoccupied by Palpomyia armipes (Meigen), 1838). South Africa. 

corinneae (Gosseries), 1989: 2 (Ceratopogon). New name for armipes. 

nudicolis Borkent, in Borkent and Bissett, 1990: 188. USA (Maine). 

pacifica Remm, 1990: 4 (Remm, in Borkent and Bissett, 1990: 201). Russia. 

pendleburyi Macfie, 1934c: 280. Malaysia. 

polonica Szadziewski, 1988: 135. Poland. Eocene. 

punctata Giles and Wirth, 1982a: 446. Sri Lanka. 

reyei Debenham, 1970d: 161. Australia (Northern Territory). 

rossi de Meillon and Wirth, 1983b: 396. Madagascar. 

rufitarsis (Meigen), 1818: 83 (Ceratopogon). Europe. 

gelida Kieffer, 1925a: 429. Latvia. 

bispinosa Goetghebuer, 1936: 321. Belgium. 

dipetala Remm, 1965: 182. Estonia. 

ryszardi Borkent, in Borkent and Bissett, 1990: 207. Poland. Eocene. 

sierrensis Borkent, in Borkent and Bissett, 1990: 190. USA (California). 

silvatica de Meillon and Downes, 1986: 175. South Africa. 

sinuosa Borkent, in Borkent and Bissett, 1990: 207. Poland. Eocene. 

spinigera (Loew), 1850: 30 (Ceratopogon). Baltic region. Eocene. 

elongata (Meunier), 1904a: 231 (1904b: 242) (Ceratopogon, preoccupied by Forcipomyia elongata (Kieffer), 1901a). 

131

Baltic region. Eocene. 

stuckenbergi de Meillon and Wirth, 1983b: 398. South Africa. 

subinermis Kieffer, 1919a: 73 (as variety of spinosipes). Hungary. 


tecta Borkent, in Borkent and Bissett, 1990: 205. Austria. 

vockerothi Borkent, in Borkent and Bissett, 1990: 191. Canada (Manitoba). 

zuluensis de Meillon and Wirth, 1981c: 589. South Africa. 

Genus SINHALOHELEA Grogan and Borkent 

SINHALOHELEA Grogan and Borkent, 1992: 314. Type species: Sinhalohelea gansi Grogan and Borkent, by original 

designation. 

dayongi Yu, 2003b:201. China. 

gansi Grogan and Borkent, 1992: 315. Sri Lanka. 

pingyii Yu, in Yu et al., 2005: 1443. China. 

Genus SPINELLIHELEA Borkent, Grogan and Picado 

SPINELLIHELEA Borkent, Grogan and Picado, 2008: 623. Type species: Spinellihelea brevicosta Borkent, Grogan and 

Picado, by original designation. 

brevicosta Borkent, Grogan and Picado, 2008: 627. Costa Rica. 

Genus STILOBEZZIA Kieffer 

STILOBEZZIA Kieffer, 1911a: 118. Type species: Stilobezzia festiva Kieffer, by original designation. 

HARTOMYIA Malloch, 1915a: 339. Type species: Ceratopogon pictus Coquillett (= Stilobezzia coquilletti Kieffer), by original 

designation. 

Subgenus ACANTHOHELEA Kieffer 

ACANTHOHELEA Kieffer, 1917a: 198. Type species: Acanthohelea pruinosa Kieffer, by monotypy. 

PALEOTETRAGONEURA Meunier, 1920a: 897 (1920b: 1221). Type species: Tetragoneura veterana Meunier, by 

monotypy. 

NEOSTILOBEZZIA Goetghebuer, 1934a: 53 (as subgenus of Stilobezzia). Unavailable name; proposed after 1930 without 

type speciesdesignation. 

NEOSTILOBEZZIA Wirth, 1953a: 63. Type species: Ceratopogon ochraceus Winnertz, by original designation. 

acrotrichis Tokunaga, 1959: 301. Papua New Guinea. 

afra Clastrier, 1991a: 298. Guinea. 

afrotropica Clastrier, 1991a: 302. Guinea. 

alba Tokunaga, 1940d: 155. Japan. 

albiabdominalis Tokunaga and Murachi, 1959: 368. Micronesia. 

amaniensis de Meillon, in Wirth et al., 1980: 168. New name for monticola. 

monticola de Meillon, 1960: 403 (preoccupied by Stilobezzia monticola Tokunaga, 1940b). Tanzania. 

amazonica Clastrier, 1991a: 306. French Guiana. 

amplistyla Clastrier, 1989a: 13. Guinea. 

anomalapennis Yu, in Yu et al., 2005: 1449. China. 

antipodalis Ingram and Macfie, 1931b: 203. New Zealand. 

armatibiae Tokunaga, 1963a: 260. Papua New Guinea. 

artistyla Das Gupta and Wirth, 1968: 41. Thailand. 

atrichopogon Lane and Forattini, 1956: 208. Panama. 

badia Macfie, 1932c: 41. New Zealand. 

132

iangulata Tokunaga, 1963a: 274. Papua New Guinea. 

bicinctipes Ingram and Macfie, 1931a: 195. Chile. 

bifurcata Tokunaga, 1959: 307. Papua New Guinea. 

borkenti Cazorla and Spinelli, 2012: 74. Argentina. 


brevistyla Clastrier, 1989a: 12. Guinea. 

calcaris Tokunaga and Murachi, 1959: 382. Micronesia. 

carayoni Clastrier, 1986a: 284. Guinea. 

centripictura Tokunaga, 1963a: 252. Indonesia. 

cereola Clastrier, 1963: 51. Macedonia. 

challieri Vattier and Adam, 1966b: 741. Congo. 

clavicula Tokunaga, 1963a: 270. Papua New Guinea. 

chlorogastrula Yu and Yuan, 2007:76. China. 

constans Das Gupta and Wirth, 1968: 42. Philippines. 

corneti Clastrier, 1991a: 303. Burkina Faso. 

crassiforceps Tokunaga, 1963a: 266. Papua New Guinea. 

crassivenosa Das Gupta and Wirth, 1968: 44. Pakistan. 

curvistyla Cazorla and Spinelli, 2007: 182. Argentina. 

differens de Meillon, 1960: 406. Liberia. 

dispartheca Das Gupta, Chaudhuri and Sanyal, 1971: 440. India. 

distinctigenitalis Das Gupta, Chaudhuri and Sanyal, 1971: 438. India. 

djalonensis Clastrier, 1988c: 127. Guinea. 

dominicana Szadziewski and Grogan, 1998a: 45. Dominican Republic. Miocene. 

donskoffi Clastrier, 1988a: 60. Democratic Republic of the Congo. 

douryi Clastrier, 1963: 42. Algeria. 

downesi Cazorla and Spinelli, 2012: 77. Chile. 

edwardsi Ingram and Macfie, 1931a: 198. Argentina. 

eliptaminensis Tokunaga, 1963a: 265. Papua New Guinea. 

falcata (Meunier), 1904a: 233 (1904b: 244) (Ceratopogon). Baltic region. Eocene. 

spinosa (Meunier), 1904a: 234 (1904b: 245) (Ceratopogon). Baltic region. Eocene. 

fasciscutata Das Gupta and Wirth, 1968: 46. Malaysia. 

fitzroyensis Lee, 1948c: 350. Australia (New South Wales). 

flaccisacca Yu and Zhang, in Yu et al., 2005: 1452. China. 

flava Chaudhuri, Das Gupta and Sinharay, 1974: 49. India. 

flavida de Meillon and Wirth, 1987a: 56. South Africa. 

fortistyla Das Gupta and Wirth, 1968: 48. Malaysia. 

fulva de Meillon and Downes, 1986: 176. South Africa. 

fulvacea Remm, 1980: 95. Tajikistan. 

fulviscuta Tokunaga and Murachi, 1959: 399. Micronesia. 

furcellata Remm, 1980: 93. Tajikistan. 

furcipes de Meillon, 1960: 404. Liberia. 

furva Ingram and Macfie, 1931a: 200. Argentina. 

fuscidorsum Kieffer, 1921a: 58. Germany. 

fuscigenua Tokunaga and Murachi, 1959: 373. Belau (USA). 

fusciscutellata Tokunaga and Murachi, 1959: 376. Micronesia. 

fuscula Wirth, 1952a: 204. USA (California). 

fusistylata Tokunaga and Murachi, 1959: 392. Micronesia. 

gallica Clastrier, 1991a: 305. France. 

genitalis Lee, 1948c: 349. Australia (Tasmania). 

gigantiforceps Clastrier, 1989a: 6. Guinea. 

gracilenta Yu and Zou, in Yu et al., 2005: 1454. China. 

gracilis (Haliday), 1833: 152 (Ceratopogon). Great Britain. 

dorsalis (Zetterstedt), 1850: 3644 (Ceratopogon). Sweden. 

gressitti Tokunaga and Murachi, 1959: 372. Micronesia. 

133

grogani Cazorla and Spinelli, 2012: 87. Chile. 

guarani Cazorla and Spinelli, 2010: 46. Argentina. 

guianae (Macfie), 1940d: 28 (Acanthohelea). Guyana. 

hirsuta Ingram and Macfie, 1931a: 201. Argentina. 

illustris Das Gupta, Chaudhuri and Sanyal, 1971: 441. India. 

impartheca Das Gupta, Chaudhuri and Sanyal, 1971: 443. India. 

ingrami Cazorla and Spinelli, 2012: 90. Argentina. 

insigniforceps Clastrier, 1989a: 9. Guinea. 

insolita Das Gupta and Wirth, 1968: 49. Malaysia. 

kindiae Clastrier, 1988c: 121. Guinea. 

kunashiri Remm, 1993: 186. Russia. 

kutscheri Szadziewski, 1993: 637. Germany. Eocene. 

lanceloti Macfie, 1937b: 78. Ethiopia. 

latiforceps Tokunaga, in Tokunaga and Murachi, 1959: 386. Belau (USA). 

setigera Tokunaga, in Tokunaga and Murachi, 1959: 387 (valid subspecies of latiforceps). Belau (USA). 

latistyla Clastrier, 1989a: 10. Guinea. 

latiunguis Clastrier, 1985e: 460. Guinea. 

lengi Yu, in Yu et al., 2005: 1456. China. 

limai Wirth and Derron, 1976: 234. Sao Tomé and Principe. 

longicornis Goetghebuer, 1934d: 193. Democratic Republic of the Congo. 

longisacca Yu and Deng, in Yu et al., 2005: 1457. China. 

longistyla Tokunaga, 1941a: 117. Micronesia. 

lutacea Edwards, 1926a: 412. Great Britain. 

lutea (Malloch), 1918a: 18 (Hartomyia). USA (Illinois). 

mallochi Hoffman, 1924: 283. USA (New York). 

luteola de Meillon, 1940: 460. South Africa. 


macclurei Das Gupta and Wirth, 1968: 51. Malaysia. 

magnitheca Das Gupta and Wirth, 1968: 52. Indonesia. 

manaosensis Lane and Forattini, 1958: 205. Brazil. 

menglaensis Yu and Huang, in Yu et al., 2005: 1459. China. 

miripes Das Gupta and Wirth, 1968: 54. Malaysia. 

monopicta Cazorla and Spinelli, 2010: 48. Argentina. 

monticola Tokunaga, 1940b: 279. Japan. 

natalensis de Meillon, 1939a: 20. South Africa. 

nebulosa Tokunaga, 1963a: 252. Papua New Guinea. 

nigerrima Ingram and Macfie, 1931a: 196. Argentina. 

nigriapicalis Tokunaga, 1963a: 275. Papua New Guinea. 

niveus Liu, Yan and Liu, 1996: 44. China. 

obesa Das Gupta and Wirth, 1968: 56. Indonesia. 

obesigenitalis Das Gupta and Wirth, 1968: 58. Malaysia. 

ochracea (Winnertz), 1852: 48 (Ceratopogon). Germany. 

rufithorax Kieffer, 1919a: 84. Slovak Republic. 

scutellata Goetghebuer, 1920: 111. Belgium. 

ohakunei Ingram and Macfie, 1931b: 202. New Zealand. 


orientis de Meillon and Wirth, 1981b: 541. South Africa. 

ornata Lane and Forattini, 1958: 206. Panama. 

ornaticrus Ingram and Macfie, 1931a: 194. Chile. 

ornatithorax Clastrier, 1988c: 128. Guinea. 

ovatitheca Das Gupta, Chaudhuri and Sanyal, 1971: 445. India. 

pallidicornis Tokunaga and Murachi, 1959: 366. Belau (USA). 

papillata Remm, 1980: 93. Tajikistan. 


134

paranaense Cazorla and Spinelli, 2010: 50. Argentina. 

parvaeungulae Das Gupta and Wirth, 1968: 61. Malaysia. 

parvitheca Das Gupta and Wirth, 1968: 59. Malaysia. 


pessoni Neveu, 1977: 230. France. 

postcervix Tokunaga, 1959: 303. Indonesia. 

pruefferi Szadziewski, 1992a: 83. Algeria. 

pruinosa (Kieffer), 1917a: 198 (Acanthohelea). Australia (New South Wales). 

pseudoparvitheca Das Gupta, Chaudhuri and Sanyal, 1971: 447. India. 

rava Ingram and Macfie, 1931a: 203. Chile. 

reflexa Tokunaga, 1963a: 269. Papua New Guinea. 

robusta Das Gupta and Wirth, 1968: 62. Malaysia. 

royi Das Gupta, Chaudhuri and Sanyal, 1971: 447. India. 


aureola Clastrier, 1963: 49. Algeria. 

samoana Edwards, 1928: 57. Western Samoa. 

sanctibernardini Kieffer, 1917b: 308. Paraguay. 

saxonica Szadziewski, 1993: 638. Germany. Eocene. 

scutica Chaudhuri, Das Gupta and Sinharay, 1974: 47. India. 

seguyi Clastrier, 1990: 399. Guinea. 

semiartistyla Sinha, Das Gupta, and Chaudhuri, 2003a: 530. India. 

setigeripes Tokunaga, 1963a: 268. Papua New Guinea. 

setigeriscutellata Tokunaga and Murachi, 1959: 394. Micronesia. 

similisegmenta Tokunaga, 1959: 306. Papua New Guinea. 

simulator de Meillon, 1960: 408. Liberia. 

singularis Clastrier, 1985e: 464. Guinea. 

soror Johannsen, 1932: 432. Indonesia. 

spadicicoxalis Tokunaga and Murachi, 1959: 396. Micronesia. 

spadicitibialis Tokunaga and Murachi, 1959: 370. Micronesia. 

speculae Macfie, 1934c: 284. Malaysia. 

spinifemorata Tokunaga, 1963a: 262. Indonesia. 

stonei Wirth, 1953a: 66. USA (Virginia). 

subalba Das Gupta and Wirth, 1968: 64. Malaysia. 

subnebulosa Das Gupta and Wirth, 1968: 38. Thailand. 

subsoror Tokunaga, 1941a: 116. Micronesia. 

subumbrosa Das Gupta, Chaudhuri and Sanyal, 1971: 449. India. 

succinea Ingram and Macfie, 1931a: 200. Argentina. 

szadziewskii Chaudhuri and Das Gupta, 1991: 182. New name for falcata. 

falcata Chaudhuri, Das Gupta and Sinharay, 1974: 48 (preoccupied by Stilobezzia falcata (Meunier), 1904a). India. 

tasmaniensis Lee, 1948c: 347. Australia (Tasmania). 

tenuiforceps Tokunaga and Murachi, 1959: 384. Belau (USA). 

thibaulti Neveu, 1977: 226. France. 

thyridofera Tokunaga, 1959: 304. Indonesia. 

tibialis Lane and Forattini, 1956: 209. Panama. 

tomensis Wirth and Derron, 1976: 236. Sao Tomé and Principe. 

tonnoiri Macfie, 1932c: 43. New Zealand. 

trilineata de Meillon and Wirth, 1983a: 365. South Africa. 

trimaculata de Meillon and Wirth, 1987a: 57. South Africa. 

truncata Tokunaga, in Tokunaga and Murachi, 1959: 389. Micronesia. 

umbrosa Das Gupta, Chaudhuri and Sanyal, 1971: 452. India. 

unicellula Clastrier, 1985e: 463. Guinea. 

unifascidorsalis Tokunaga, in Tokunaga and Murachi, 1959: 379. Belau (USA). 

vandeli Vattier and Adam, 1966b: 738. Congo. 

varia Ingram and Macfie, 1931a: 191. Chile. 

135

venefica Das Gupta and Wirth, 1968: 39. Thailand. 

veterana (Meunier), 1920a: 897 (1920b: 1221) (Tetragoneura). Germany. Oligocene. 

goetghebueri Statz, 1944: 148. Germany. Oligocene. 

vittula Tokunaga, 1963a: 260. Papua New Guinea. 

wanlinensis Yu and Li, in Yu et al., 2005: 1461. China. 

wirthicola Szadziewski and Grogan, 1998a: 45. New name for succinea. 

succinea Szadziewski, 1993: 639 (preoccupied by Stilobezzia succinea Ingram and Macfie, 1931a). Germany. 

Eocene. 

xanthogaster Das Gupta and Wirth, 1968: 65. Malaysia. 

xerophila Cazorla and Spinelli, 2010: 52. Argentina. 

Subgenus DEBENHAMIA Wirth and Grogan 

DEBENHAMIA Wirth and Grogan, 1988: 89 (as subgenus of Stilobezzia). Type species: Stilobezzia collessi Wirth and 

Grogan, by original designation. 

collessi Wirth and Grogan, 1988: 91. Australia (New South Wales). 

dycei Wirth and Grogan, 1988: 93. Australia (Western Australia). 

Subgenus EUKRAIOHELEA Ingram and Macfie 

EUKRAIOHELEA Ingram and Macfie, 1921: 347. Type species: Eukraiohelea africana Ingram and Macfie, designation by 

Macfie, 1940f: 22. 

africana (Ingram and Macfie), 1921: 347 (Eukraiohelea). Ghana. 

amnigena (Macfie), 1935a: 56 (Eukraiohelea). Brazil. 

brevicostalis Das Gupta and Wirth, 1968: 28. Malaysia. 

clarifemorata Das Gupta and Wirth, 1968: 18. Malaysia. 

dorsofasciata (Lutz), 1914: 96 (Palpomyia). Brazil. 

elegantula (Johannsen), 1907: 109 (Bezzia). USA (Kansas). 

subsessilis Kieffer, 1917b: 311. Paraguay. 

maculitibia Lane and Forattini, 1956: 207. Panama. 

foyi (Ingram and Macfie), 1922: 270 (Eukraiohelea). Nigeria. 

fuscipes Das Gupta and Wirth, 1968: 20. Thailand. 

fusciterga Das Gupta and Wirth, 1968: 22. Thailand. 

minima Kieffer, 1918a: 100. India. 

minuta Das Gupta and Wirth, 1968: 23. Thailand. 

punctifemorata Das Gupta and Wirth, 1968: 25. Malaysia. 

versicolor (Ingram and Macfie), 1921: 351 (Eukraiohelea). Ghana. 

Subgenus STILOBEZZIA Kieffer 

addita Clastrier, 1986b: 380. Democratic Republic of the Congo. 

albicoxa Lane and Forattini, 1956: 210. Panama. 

albocincta Kieffer, 1917b: 309. Paraguay. 

americana Kieffer, 1917b: 310. Paraguay. 

angustipennis Clastrier, 1988c: 125. Guinea. 

ani Yu, in Yu et al., 2005: 1464. China. 

antennalis (Coquillett), 1901a: 606 (Ceratopogon). USA (District of Columbia). 

antilleana Szadziewski and Grogan, 1998a: 46. Dominican Republic. Miocene. 

areolaris (Kieffer), 1911c: 345 (Sphaeromias). Seychelles. 

atronitens Goetghebuer, 1933e: 148. Democratic Republic of the Congo. 

baojia Liu and Shi, 2002: 187. China. 

136

asizonata Tokunaga, 1963a: 254. Indonesia. 

bata de Meillon and Hardy, 1954: 62. Cameroon. 

beckae Wirth, 1953a: 69. USA (Florida). 

bessa Yu and Zhang, in Yu et al., 2005: 1465. China. 

bicolor Lane, 1947a: 208. Brazil. 

bimacula (Kieffer), 1910: 201 (Palpomyia). India. 

bimaculata Lane and Forattini, 1956: 211. Panama. 

bispinosa Kieffer, 1917b: 310. Paraguay. 

bizonata Tokunaga, 1963a: 256. Papua New Guinea. 

blaesospira Yu and Deng, in Yu et al., 2005: 1467. China. 

blantoni Lane and Forattini, 1956: 211. Panama. 

boharti Tokunaga, 1962a: 211. Japan. 

brevicornis Clastrier, 1988c: 130. Guinea. 

browni Wirth and Giles, 1990: 454. Fiji. 

bulla Thomsen, 1935: 289. USA (New York). 

caribe Lane and Forattini, 1958: 208. Panama. 

castanea Macfie, 1934c: 284 (1934d: 220). Malaysia. 

chaconi Macfie, 1938: 166. Trinidad and Tobago. 

chasteli Clastrier, 1967: 115. Cambodia. 

punctivenosa Das Gupta and Wirth, 1968: 106. Thailand. 

chlorosa Clastrier, 1986b: 364. New name for viridis. 

viridis Goetghebuer, 1935d: 179 (preoccupied by Stilobezzia viridis (Coquillett), 1901a). Democratic Republic of 

Congo. 

claripennis Clastrier, 1958b: 243. Senegal. 

claripes Das Gupta and Wirth, 1968: 128. South Korea. 

tokunagai Das Gupta and Wirth, 1968: 131. Japan. 

maihensis Remm, 1971: 209. Russia. 

clavella Yu, in Yu et al., 2005: 1468. China. 

congestiterga Das Gupta and Wirth, 1968: 124. Malaysia. 

coquilletti Kieffer, 1917b: 308. New name for picta. 

picta (Coquillett), 1905: 60 (Ceratopogon, preoccupied by Sphaeromias pictus (Meigen), 1818). USA (Virginia). 

coracina Kieffer, 1917b: 311. Paraguay. 

crassipes Kieffer, 1918a: 101. India. 

crassistyla Das Gupta and Wirth, 1968: 125. Philippines. 

crossi de Meillon and Wirth, 1981c: 591. South Africa. 

debilipes Das Gupta and Wirth, 1968: 134. Malaysia. 

decora Kieffer, 1916a: 89. Taiwan. 

diminuta Lane and Forattini, 1958: 209. Panama. 

distinctifasciata Das Gupta and Wirth, 1968: 135. Thailand. 

diversa (Coquillett), 1901a: 607 (Ceratopogon). USA (New Jersey). 

dorsosignata Sinha, Das Gupta, and Chaudhuri, 2003a: 529. India. 

dryadum Macfie, 1940b: 186. Guyana. 

dubitans Lane, Forattini and Rabello, 1955: 85. Brazil. 

dureti Lane and Forattini, 1958: 210. Brazil. 

ensistyla Chaudhuri, Das Gupta and Chatterjee, 1981: 149. India. 

erectiseta Liu, Yan and Liu, 1996: 43. China. 

esmeralda Lane and Forattini, 1958: 211. Panama. 

eximitarsis Das Gupta and Wirth, 1968: 115. Malaysia. 

femoralis Lane and Forattini, 1956: 212. Panama. 

festiva Kieffer, 1911a: 118. India. 

scutellaris Kieffer, 1912c: 7 (as variety of festiva). Sri Lanka. 

fiebrigi Kieffer, 1917b: 309. Paraguay. 

filapenis Yu and Liu, 1991: 50. China. 

flavipectoralis Remm, 1993: 187. Russia. 

137

flavirostris (Winnertz), 1852: 52 (Ceratopogon). Germany. 

albicornis Kieffer, 1919a: 84. Austria. 

flavirostroides (Strobl), 1880: 64 (Ceratopogon). Austria. 

lateralis (Waltl), 1837 : 279 (Ceratopogon, preoccupied by Forcipomyia lateralis (Bouché), 1834). Germany. 

flavizonata Tokunaga, 1963a: 263. Indonesia. 

fortipes Das Gupta and Wirth, 1968: 72. Malaysia. 

gambiae Clastrier and Wirth, 1961a: 216. Gambia. 

glauca Macfie, 1939c: 204. Brazil. 

fluminensis Lane, 1947a: 210. Brazil. 

grandis Lane and Forattini, 1958: 213. Panama. 

harurii Boorman and Harten, 2002: 454. Yemen. 

hirta Borkent, in Borkent and Wirth, 1997:111. New name for pruinosa. 

pruinosa Wirth, 1952a: 203 (preoccupied by Stilobezzia pruinosa (Kieffer), 1917a). USA (California). 

hirtaterga Yu, 1989a: 475. China. 

hollandia Tokunaga, 1959: 299. Indonesia. 

huberti Das Gupta and Wirth, 1968: 85. Malaysia. 

immodentis Liu, Yan and Liu, 1996: 44. China. 

imparungulae Das Gupta and Wirth, 1968: 136. Malaysia. 

inermipes Kieffer, 1912c: 8. Sri Lanka. 

lineata Kieffer, 1913d: 185. India. 

biroi Kieffer, 1918a: 102. Singapore. 

aberrans Johannsen, 1932: 431. Indonesia. 

esakiana Tokunaga, 1940e: 183 (1940d: 155). Micronesia. 

inusitata (Johannsen), 1946: 190 (Eukraiohelea). Guam (USA). 

inkisiensis Clastrier, 1986b: 379. Democratic Republic of the Congo. 

intermedia de Meillon, 1939a: 18. South Africa. 

isthmostheca Das Gupta and Wirth, 1968: 116. Thailand. 

jinggangshana Yu, Liu and Ma, in Liu et al., 2010: 572. China. 

kiefferi Lane, 1947a: 205. Brazil. 

punctipes Wirth, 1953a: 79. USA (Florida). 

kisantuensis Clastrier, 1986b: 369. Democratic Republic of the Congo. 

lasioterga Das Gupta and Wirth, 1968: 126. Malaysia. 

leucopeza Clastrier, 1958b: 249. Senegal. 

lijiangi Yu, Zhang and Mo, in Deng et al., 2011: 116. China. 

limnophila Ingram and Macfie, 1922: 267. Ghana. 

longiforceps Clastrier, 1960b: 274. Congo. 

longihamata Tokunaga, 1963a: 258. Papua New Guinea. 

macfiei Lane, 1947a: 213. Brazil. 

maculata Lane, 1947a: 207. Brazil. 

maculipes Macfie, 1933b: 103. French Polynesia (France). 

mahensis Clastrier, 1983c: 76. Seychelles. 

maia Lane and Forattini, 1958: 204. Panama. 

merceri Cazorla and Spinelli, in Cazorla et al., 2005: 290. Peru. 

modesta Lane, 1947a: 206. Brazil. 

mutabilis Clastrier, 1986c: 63. Guinea. 

nasicae de Meillon, 1959b: 9. Congo. 

navaiae Wirth and Grogan, 1981: 86. USA (New York). 

nigroflava Lane and Forattini, 1958: 215. Costa Rica. 

nitela Yu and Yan, 2005: 168. Vietnam. 

notata (de Meijere), 1907: 210 (Ceratopogon). Indonesia. 

nudisthmostheca Das Gupta and Wirth, 1968: 118. Sri Lanka. 

nyei Clastrier, 1983c: 77. Seychelles. 

obscura Lane and Forattini, 1958: 216. Panama. 

oxiana Remm, 1980: 95. Tajikistan. 

138

pallescens Lane and Forattini, 1958: 218. Panama. 

pallidicollis Yu, in Yu et al., 2005: 1480. China. 

pallidipes Clastrier, 1983c: 78. Seychelles. 

pallidiventris (Malloch), 1915a: 344 (Hartomyia). USA (Illinois). 

palpalis Tokunaga, 1963a: 255. Indonesia. 

panamensis Lane and Forattini, 1958: 218. Panama. 

parvula Goetghebuer, 1933e: 149. Democratic Republic of the Congo. 

pastoriana Clastrier, 1986c: 66. Guinea. 

paucimaculata Clastrier, 1984d: 46. Guinea. 

paucipictipes Das Gupta and Wirth, 1968: 98. Indonesia. 

pauliani de Meillon, 1961: 57. Madagascar. 

paulistensis Lane, 1947a: 200. Brazil. 

perspicua Johannsen, 1932: 433 (as variety of notata). Indonesia. 

photophila Clastrier, 1984d: 43. Guinea. 

pictipes Kieffer, 1917a: 191. Australia (New South Wales). 

poikiloptera (Ingram and Macfie), 1922: 276 (Parabezzia). Ghana. 

propristyla Das Gupta and Wirth, 1968: 129. Malaysia. 

pseudofestiva Das Gupta and Wirth, 1968: 89. Thailand. 

pseudonotata Das Gupta and Wirth, 1968: 91. Malaysia. 

pseudopunctulata Cazorla and Ronderos, in Cazorla et al., 2012: 400. Argentina. 

punctulata Lane, 1947a: 204. Brazil. 

quatei Das Gupta and Wirth, 1968: 111. Viet Nam. 

rabelloi Lane, 1947a: 203. Brazil. 

rotunditheca Das Gupta and Wirth, 1968: 73. Malaysia. 

rufa Kieffer, 1921b: 23. Cameroon. 

rutshuruensis Clastrier, 1986b: 373. Democratic Republic of the Congo. 

seychelleana Clastrier, 1983c: 79. Seychelles. 

silvicola Macfie, 1940b: 185. Guyana. 

similans Lane and Forattini, 1956: 214. Panama. 

simplex Lane and Forattini, 1958: 222. Panama. 

spinipes Das Gupta and Wirth, 1968: 74. Philippines. 

spinitarsis Das Gupta and Wirth, 1968: 92. Malaysia. 

spiniterga Das Gupta and Wirth, 1968: 119. Malaysia. 

spirogyrae Carter, Ingram and Macfie, 1921a: 325. Ghana. 

subfestiva Das Gupta and Wirth, 1968: 94. Thailand. 

subflava Das Gupta and Wirth, 1968: 76. Malaysia. 

subviridis Macfie, 1934d: 218. Indonesia. 

esakii Tokunaga, 1940e: 183 (1940d: 155). Micronesia. 

supernotata Das Gupta and Wirth, 1968: 95. Malaysia. 

sybleae Wirth, 1953a: 82. USA (Virginia). 

tauffliebi de Meillon, 1959b: 12. Congo. 

tenebrosa Macfie, 1933b: 101. French Polynesia (France). 

tenuicolorata Das Gupta and Wirth, 1968: 123. Malaysia. 

tetragona Goetghebuer, 1934d: 193. Democratic Republic of the Congo. 

thomsenae Wirth, 1953a: 83. USA (Florida). 

scutata Lane and Forattini, 1961: 92. Panama. 

transversa Lane and Forattini, 1958: 222. Panama. 

traubi Das Gupta and Wirth, 1968: 77. Malaysia. 

travassosi Lane, 1947a: 210. Brazil. 

tropica Clastrier, 1958b: 247. Senegal. 

unifasciata Tokunaga, 1963a: 262. Indonesia. 

venezuelensis Ortiz, 1950d: 199 (as variety of glauca). Venezuela. 

virescens Kieffer, 1919a: 84. Bulgaria. 

viridis (Coquillett), 1901a: 607 (Ceratopogon). USA (New Jersey). 

139

viridiventris (Kieffer), 1910: 203 (Palpomyia). Myanmar. 

vittata Clastrier, 1960b: 276. Congo. 

vulgaris Yu, 1989a: 478. China. 

wenganga Yu, Wu and Liu, 2011:46. China. 

williamsi Cazorla and Spinelli, in Cazorla et al., 2005: 290. Peru. 

wirthi Lane and Forattini, 1956: 214. Panama. 

wudangshanensis Yu and Liu, 1991: 47. China. 

wygodzinskyi Lane, 1947a: 212. Brazil. 

zonata Tokunaga, 1963c: 43. Japan. 

Species of Stilobezzia unplaced to subgenus 

kurthi Borkent, 2000b: 464. USA (New Jersey). Upper Cretaceous. 

pikei Borkent, 2012: 763. Canada (Alberta). Upper Cretaceous. 

Genus STILOCULICOIDES Wirth and Grogan 

STILOCULICOIDES Wirth and Grogan, 1988: 95. Type species: Stilobezzia ugandae Ingram and Macfie, by original 

designation. 

africanus Clastrier, 1993a: 134. Guinea. 

congoensis Clastrier, 1993a: 139. Congo. 

guineensis Clastrier, 1993a: 141. Guinea. 

kindiae Clastrier, 1993a: 138. Guinea. 

ugandae (Ingram and Macfie), 1923: 62 (Stilobezzia). Uganda. 

vauclusensis Clastrier, 1993a: 141. France. 

Genus WIRTHOHELEA Szadziewski 

WIRTHOHELEA Szadziewski, 1988: 158. Type species: Wirthohelea trifida Szadziewski, by original designation. 

trifida Szadziewski, 1988: 159. Poland. Eocene. 

TRIBE HETEROMYIINI WIRTH, 1962: 275 

Genus CLINOHELEA Kieffer 

CLINOHELEA Kieffer, 1917b: 295. Type species: Ceratopogon variegatus Winnertz (= Ceratopogon unimaculata 

Macquart), by original designation. 

Subgenus CERATOBEZZIA Kieffer 

CERATOBEZZIA Kieffer, 1917b: 326. Type species: Ceratobezzia fallax Kieffer, by original designation. 

fallax Kieffer, 1917b: 326. Paraguay. 

barrettoi Lane and Duret, 1954: 249. Brazil. 

Subgenus CLINOHELEA 

albopennis Lane, 1944: 259. Brazil. 

argentina Lane and Duret, 1954: 248. Argentina. 

bimaculata (Loew), 1861: 311 (Ceratopogon). USA (District of Columbia). 

bisignata (Kieffer), 1910: 200 (Palpomyia). India. 

curriei (Coquillett), 1905: 62 (Ceratopogon). Canada (British Columbia). 

nebulosa (Malloch), 1915a: 322 (Palpomyia). USA (Michigan). 

140

damascenoi Lane and Duret, 1954: 250. Brazil. 

dimidiata (Adams), 1903: 27 (Ceratopogon). USA (Arizona). 

dryas Debenham, 1974: 12. Australia (Queensland). 

flagellata (Edwards), 1916: 500 (Palpomyia). Australia (North Australia). 

fuscoalata Remm, 1993: 181. Russia. 

hollandiae Tokunaga, 1966a: 111. Indonesia. 

horacioi Lane, 1944: 257. Brazil. 

hygropetrica Clastrier, 1983b: 19. Guinea. 

insperata de Meillon and Wirth, 1981c: 592. South Africa. 

lacustris Macfie, 1939a: 91. Uganda. 

longipalpis Kieffer, 1918a: 96. Singapore. 

longitheca Grogan and Wirth, 1975b: 286. USA (Florida). 

muzoni Spinelli and Duret, 1993: 46. Colombia. 

neivai Lane, 1944: 252. Brazil. 

nigeriae (Ingram and Macfie), 1923: 68 (Ceratobezzia). Nigeria. 

nigripes Macfie, 1939c: 205. Brazil. 

nubifera (Coquillett), 1905: 61 (Ceratopogon). USA (Florida). 

pachydactyla Kieffer, 1918a: 96. Singapore. 

papuensis Tokunaga, 1966a: 109. Papua New Guinea. 

podagrica Goetghebuer, 1948b: 17. Democratic Republic of the Congo. 

pseudobisignata Saha and Das Gupta, 1996: 7. India. 

pseudonubifera Grogan and Wirth, 1975b: 280. USA (Maryland). 

reperticia Yu and Zhang, 1996:294. China. 

rubriceps Kieffer, 1917b: 318. Paraguay. 

saltanensis Lane and Duret, 1954: 252. Argentina. 

tasmaniensis Lee, 1948d: 65. Australia (Tasmania). 

tenuipes Tokunaga, 1966a: 110. Indonesia. 

tenuissima (Kieffer), 1917a: 195 (Sphaeromias). Papua New Guinea. 

townsendi Lane, 1944: 256. Brazil. 

townesi Lane, 1944: 254. Brazil. 

trimaculata Clastrier, 1983b: 24. Guinea. 

unimaculata (Macquart), 1826: 178 (Ceratopogon). France. 

maculipes (Meigen), 1830: 265 (Ceratopogon). Europe. 

variegata (Winnertz), 1852: 59 (Ceratopogon). Germany. 

usheri de Meillon, 1959b: 20. Mozambique. 

usingeri Wirth, 1952a: 209. USA (California). 

wygodzinskyi Lane, 1948: 231. Brazil. 

Genus HETEROMYIA Say 

HETEROMYIA Say, 1825: plate 35. Type species: Heteromyia fasciata Say, by monotypy. 

PACHYLEPTUS Walker, 1856b: 426. Type species: Pachyleptus fasciatus Walker (= Heteromyia nigra Kieffer), by 

monotypy. 

antequerae (Lynch Arribálzaga), 1893: 227 (Pachyleptus). Argentina. 

bejaranoi Duret and Lane, 1955: 36. Argentina. 

castanea Lane, 1946a: 214. Brazil. 

chaquensis Duret and Lane, 1955: 37. Argentina. 

clavata Williston, 1900: 225. Mexico. 

rufa Kieffer, 1917b: 325. Colombia. 

caloptera Kieffer, 1919c: 192. New name for rufa. 

correntina Duret and Lane, 1955: 39. Argentina. 

dominicana Szadziewski and Grogan, 1997: 254. Dominican Republic. Miocene. 

fasciata Say, 1825: plate 35. North America. 

141

festiva (Loew), 1861: 314 (Ceratopogon). USA (Pennsylvania). 


lamprogaster Edwards, 1933b: 87. Argentina. 

nigra Kieffer, 1917b: 326. Paraguay. 

fasciatus (Walker), 1856b: 426 (Pachyleptus, preoccupied by Heteromyia fasciata Say, 1825). South America. 

orellana (Roback), 1957: 1 (Palpomyia). Peru. 

prattii Coquillett, 1902a: 88. USA (Virginia). 

wokei Wirth and Grogan, 1977: 181. Nicaragua. 

Genus METAHELEA Edwards 

METAHELEA Edwards, 1929b: 12. Type species: Metahelea metallescens Edwards, by monotypy. 

bifasciata (Kieffer), 1917a: 194 (Palpomyia). Australia (Queensland). 

metallescens Edwards, 1929b: 12. Philippines. 

serafini Szadziewski, 1998: 247. Russia. Eocene. 

Genus NEMOROMYIA Liu and Yu 

NEMOROMYIA Liu and Yu, 1991: 25. Type species: Nemoromyia nemorosa Liu and Yu, by original designation. 

nemorosa Liu and Yu, 1991: 26. China. 

Genus NEUROHELEA Kieffer 

NEUROHELEA Kieffer, 1925d: 112. Type species: Ceratopogon luteitarsis Waltl, by monotypy. 

luteitarsis (Waltl), 1837 : 279 (Ceratopogon). Germany. 

subsessilis (Goetghebuer), 1921: 181 (Clinohelea). Belgium. 

Genus PELLUCIDOMYIA Macfie 

PELLUCIDOMYIA Macfie, 1939a: 99. Type species: Pellucidomyia ugandae Macfie, by original designation. 

MACFIEHELEA Lane, 1946a: 208. Type species: Macfiehelea oliveirai Lane, by original designation. 

blantoni (Lane), 1956b: 435 (Macfiehelea). Panama. 

dycei Debenham, 1970a: 137. Australia (Queensland). 

lanei Wirth and Ratanaworabhan, 1971b: 446. Colombia. 

leei Wirth, 1960: 2. Australia (Queensland). 

oliveirai (Lane), 1946a: 209 (Macfiehelea). Brazil. 

sambulena (de Meillon), 1942c: 14 (Bezzia). Mozambique. 

ugandae Macfie, 1939a: 100. Uganda. 

wirthi (Lane), 1956b: 437 (Macfiehelea). Panama. 

Genus PHYSOHELEA Grogan and Wirth 

PHYSOHELEA Grogan and Wirth, 1979b: 53. Type species: Neurohelea oedidactyla Ingram and Macfie, by original 

designation. 

obtusa (Meunier), 1904a: 235 (1904b: 246) (Ceratopogon). Baltic region. Eocene. 

cothurnatula (Meunier), 1904a: 231 (1904b: 242) (Ceratopogon). Baltic region. Eocene. 

oedidactyla (Ingram and Macfie), 1931a: 212 (Neurohelea). Argentina. 

turgidipes (Ingram and Macfie), 1931a: 214 (Neurohelea). Chile. 

142

Genus TETRABEZZIA Kieffer 

TETRABEZZIA Kieffer, 1917b: 296. Type species: Dibezzia spinigera Kieffer, by original designation. 

africana Clastrier, 1982a: 307. Guinea. 

argentea Ingram and Macfie, 1923: 70. Nigeria. 

diazi de Meillon, 1961: 55. Madagascar. 

pictipennis (Kieffer), 1913d: 191 (Dibezzia). India. 

soeiroi (de Meillon), 1942c: 16 (Bezzia). Mozambique. 

spinigera (Kieffer), 1914c: 313 (Dibezzia). India. 

TRIBE SPHAEROMIINI NEWMAN, 1834: 388 

Genus ALLOIMYIA Yu and Liu 

ALLOIMYIA Yu, in Yu et al., 2005: 1493. Type species: Alloimyia yanjiensis Yu and Xu, by original designation. 

yanjiensis Yu and Xu, in Yu et al., 2005: 1494. China. 

Genus AUSTROSPHAEROMIAS Spinelli 

AUSTROSPHAEROMIAS Spinelli, 1997: 224. Type species: Austrosphaeromias apricans (Ingram and Macfie), by original 

designation. 

apricans (Ingram and Macfie), 1931a: 219 (Palpomyia). Argentina. 

sentior (Ingram and Macfie), 1931a: 222 (Palpomyia, as variety of apricans). Chile. 

chilensis Ingram and Macfie, 1931a: 218. Chile. 

wirthi Spinelli, 1997: 228. Argentina. 

Genus CALYPTOPOGON Kieffer 

CALYPTOPOGON Kieffer, 1910: 209. Type species: Calyptopogon albitarsis Kieffer (= Macropeza gibbosus Wiedemann), 

by monotypy. 

brevitarsis Macfie, 1939d: 8. Thailand. 

gibbosus (Wiedemann), 1824: 10 (Macropeza). Indonesia. 

albitarsis Kieffer, 1910: 209. India. 

kiefferi (Johannsen), 1927b: 423 (Macropeza). New name for albitarsis. 

similis (Johannsen), 1927b: 424 (Macropeza). Taiwan. 

javanensis (Kieffer), 1910: 210 (Macropeza). Indonesia. 

edwardsi (Macfie), 1939d: 9 (Macropeza). Sri Lanka. 

maithonensis Saha and Das Gupta, in Saha et al., 1995: 30. India. 

mirus Sarkar and Das Gupta, in Saha et al., 1995: 32. India. 

pseudoalbitarsis Sarkar and Das Gupta, in Saha et al., 1995: 31. India. 

Genus CHELOHELEA Giles and Wirth 

CHELOHELEA Giles and Wirth, 1985: 364. Type species: Chelohelea comata Giles and Wirth, by original designation. 

comata Giles and Wirth, 1985: 365. Malaysia. 

Genus CRISPOMYIA Debenham 

143

CRISPOMYIA Debenham, 1974: 48. Type species: Crispomyia receptaculum Debenham, by original designation. 

monacha Debenham, 1974: 52. Australia (Western Australia). 

receptaculum Debenham, 1974: 51. Australia (Western Australia). 

Genus DIBEZZIA Kieffer 

DIBEZZIA Kieffer, 1911a: 120. Type species: Dibezzia clavata Kieffer, by original designation. 

brevistila Kieffer, 1911a: 122. Bangladesh. 

clavata Kieffer, 1911a: 120. India. 

debenhamae Wirth and Ratanaworabhan, 1981a: 290. Malaysia. 

gideoni de Meillon and Wirth, 1981c: 595. South Africa. 

Genus GROGANHELEA Spinelli and Dippolito 

GROGANHELEA Spinelli and Dippolito, in Spinelli et al., 1995: 166. Type species: Groganhelea rondoniensis Spinelli and 

Dippolito, by original designation. 

rondoniensis Spinelli and Dippolito, in Spinelli et al., 1995: 167. Brazil. 

Genus GUIHELEA Yu and Qian 

GUIHELEA Yu and Qian, in Yu et al., 2005: 1496. Type species: Guihelea jingxiensis Yu and Qian, by original designation. 

jingxiensis Yu and Qian, in Yu et al., 2005: 1497. China. 

Genus HEBETULA Wirth and Debenham 

HEBETULA Wirth and Debenham, 1977: 282. Type species: Mixohelea lemur Debenham, by original designation. 

armiger (Debenham), 1974: 20 (Mixohelea). Australia (Queensland). 

atypipennis (Tokunaga), 1966a: 127 (Palpomyia). Indonesia. 

bequaerti (Goetghebuer), 1933e: 151 (Palpomyia). Democratic Republic of the Congo. 

bisulca (Debenham), 1974: 24 (Mixohelea). Australia (Western Australia). 

centonis (Debenham), 1974: 21 (Mixohelea). Australia (Queensland). 

gladiator (Debenham), 1974: 23 (Mixohelea). Australia (Capital Territory). 

hexacantha (Kieffer), 1911c: 343 (Sphaeromias). Seychelles. 

imparunguis (Kieffer), 1917a: 194 (Palpomyia). Papua New Guinea. 

australiensis (Kieffer), 1917b: 364 (Mixohelea). New name for imparunguis Kieffer. 

infirmior (Debenham), 1974: 28 (Mixohelea). Australia (New South Wales). 

latifrons (Debenham), 1974: 29 (Mixohelea). Australia (Queensland). 

lemur (Debenham), 1974: 32 (Mixohelea). Australia (Western Australia). 

lucta (Debenham), 1974: 31 (Mixohelea). Australia (New South Wales). 

maai (Tokunaga), 1966a: 115 (Xenohelea). Indonesia. 

matileorum Clastrier, 1985b: 268. New Caledonia (France). 

muroides (Debenham), 1974: 30 (Mixohelea). Australia (New South Wales). 

novaebritainica (Tokunaga), 1966a: 130 (Palpomyia). Papua New Guinea. 

novaecaledoniae Clastrier, 1985b: 271. New Caledonia (France). 

sarmensis (Tokunaga), 1966a: 116 (Xenohelea). Indonesia. 

sensilis (Debenham), 1974: 27 (Mixohelea). Australia (New South Wales). 

tonnoiri (Lee), 1948d: 66 (Xenohelea). Australia (Tasmania). 

uveae Clastrier and Delécolle, 1996: 308. Wallis and Futuna Islands (France). 

144

versicolor (Debenham), 1974: 35 (Mixohelea). Australia (Western Australia). 

Genus HOMOHELEA Kieffer 

HOMOHELEA Kieffer, 1917b: 364. Type species: Palpomyia abjuncta Kieffer, designation by Macfie, 1940f: 26. 

SCHIZODACTYLUS Ingram and Macfie, 1921: 353 (preoccupied by Schizodactylus Brullé, 1835). Type species: 

Schizodactylus telmatoscopus Ingram and Macfie, by monotypy. 

ANKISTRODACTYLUS Ingram and Macfie, 1922: 272. New name for Schizodactylus Ingram and Macfie. Type species: 

Schizodactylus telmatoscopus Ingram and Macfie, automatic. 

abjuncta (Kieffer), 1913d: 189 (Palpomyia). India. 

agrestia Ghosh, Majumdar, Mazumdar and Chaudhuri, 2009:92. India. 

albitudinis de Meillon and Wirth, 1981b: 543. South Africa. 

barkudensis Edwards, 1932a: 179. India. 

delanoe (de Meillon), 1942b: 113 (Palpomyia). Zimbabwe. 

diabola (de Meillon), 1961: 52 (Sphaeromias). Madagascar. 

dispartivena Ghosh, Majumdar, Mazumdar and Chaudhuri, 2009:94. India. 

diunota Ghosh, Majumdar, Mazumdar and Chaudhuri, 2009:96. India. 

iberica Delécolle, Blasco-Zumeta and Rieb, 1997: 338. Spain. 

insolita Ghosh, Majumdar, Mazumdar and Chaudhuri, 2009:98. India. 

insons (Johannsen), 1932: 436 (Sphaeromias). Indonesia. 

obscuripes (Macfie), 1934d: 225 (Sphaeromias). Indonesia. 

kraussi (Tokunaga), in Tokunaga and Murachi, 1959: 415 (Sphaeromias). Guam (USA). 

lacteal Ghosh, Majumdar, Mazumdar and Chaudhuri, 2009:100. India. 

longicosta (Goetghebuer), 1933e: 150 (Palpomyia). Democratic Republic of the Congo. 

melia de Meillon, 1943: 112. Zimbabwe. 

optima Ghosh, Majumdar, Mazumdar and Chaudhuri, 2009:102. India. 

splendida Ghosh, Majumdar, Mazumdar and Chaudhuri, 2009:104. India. 

stuckenbergi (de Meillon), 1961: 51 (Sphaeromias). Madagascar. 

telmatoscopa (Ingram and Macfie), 1921: 353 (Schizodactylus). Ghana. 

Genus INDOBEZZIA Das Gupta and Saha 

INDOBEZZIA Das Gupta and Saha, 1995: 638. Type species: Indobezzia nova Das Gupta and Saha, by original 

designation. 

nova Das Gupta and Saha, 1995: 638. India. 

recens Das Gupta and Saha, 1995: 639. India. 

Genus JENKINSHELEA Macfie 

JENKINSIA Kieffer, 1913d: 165 (preoccupied by Jenkinsia Jordan and Evermann, 1896). Type species: Jenkinsia 

setosipennis Kieffer, by original designation. 

JENKINSHELEA Macfie, 1934c: 177. New name for Jenkinsia Kieffer. Type species: Jenkinsia setosipennis Kieffer, 

automatic. 

accraensis (Ingram and Macfie), 1923: 41 (Jenkinsia). Ghana. 

albaria (Coquillett), 1895: 308 (Ceratopogon). USA (Florida). 

aequalis (Malloch), 1915a: 336 (Johannsenomyia). USA (Illinois). 

blantoni Grogan and Wirth, 1977b: 139. USA (Florida). 

corea de Meillon, 1942b: 116. Zimbabwe. 

distincta de Meillon and Wirth, 1983a: 367. South Africa. 

djalonensis Clastrier, 1983a: 261. Guinea. 

magnipennis (Johannsen), 1908: 268 (Johannseniella). USA (New York). 

145

niphanae Grogan and Wirth, 1981b: 45. Thailand. 

paliki Szadziewski, 1992b: 77. North Korea. 


parsetosipennis Saha and Das Gupta, 1996: 8. India. 

polyxenae de Meillon, 1936: 189. South Africa. 

rhodesiensis de Meillon, 1937a: 261. Zimbabwe. 

setosiforceps Grogan and Wirth, 1981b: 50. Malaysia. 

setosipennis (Kieffer), 1913d: 165 (Jenkinsia). India. 

stenoptera Remm, 1979a: 178. Turkmenistan. 

stonei Grogan and Wirth, 1977b: 135. USA (Florida). 

sudwalai de Meillon and Wirth, 1983a: 369. South Africa. 

tokunagai Grogan and Wirth, 1981b: 49. Thailand. 

trisensillata Clastrier, 1983a: 266. Guinea. 

Genus JOHANNSENOMYIA Malloch 

JOHANNSENOMYIA Malloch, 1915a: 332. Type species: Johannsenomyia halteralis Malloch (= Ceratopogon argentata 

Loew), designation by Wirth, 1952a: 211. 

DICROHELEA Kieffer, 1917b: 363. Type species: Palpomyia filicornis Kieffer, designation by Macfie, 1940f: 26. 

abdominalis (Tokunaga), 1966a: 120 (Dicrohelea). Papua New Guinea. 

albidorsata (de Meillon), 1937b: 368 (Dicrohelea). South Africa. 

angulosa de Meillon and Wirth, 1983a: 369. South Africa. 

annulicornis Malloch, 1918b: 230. USA (Illinois). 

argentata (Loew), 1861: 310 (Ceratopogon). USA (District of Columbia). 

halteralis Malloch, 1915a: 338 USA (Illinois). 

blantoni (Lane and Wirth), 1961: 81 (Dicrohelea). Panama. 

ceylanica (Kieffer), 1912c: 6 (Dibezzia). Sri Lanka. 

filicornis (Kieffer), 1910: 196 (Palpomyia). India. 

filitarsis (Kieffer), 1910: 202 (Palpomyia). India. 

grahami (Ingram and Macfie), 1923: 65 (Palpomyia). Nigeria. 

imparunguis (Becker), 1903: 72 (Ceratopogon). Egypt. 

inermicrus (Kieffer), 1913d: 186 (Sphaeromias). India. 

interrupta (Kieffer), 1910: 199 (Palpomyia). India. 

lalokiensis (Lee), 1948d: 69 (Dicrohelea). Papua New Guinea. 

lanei Wirth, 1965a: 4. Brazil. 

longistila (Kieffer), 1911a: 121 (Dibezzia). India. 

maai (Tokunaga), 1966a: 122 (Dicrohelea). Indonesia. 

nigeriae (Ingram and Macfie), 1923: 67 (Palpomyia). Nigeria. 

nigra Goetghebuer, 1933e: 149. Democratic Republic of the Congo. 

novaebrittanica (Tokunaga), 1966a: 118 (Dicrohelea). Papua New Guinea. 

novaeguineae (Tokunaga), 1966a: 119 (Dicrohelea). Indonesia. 

petersi (Tokunaga), 1966a: 151 (Probezzia). Papua New Guinea. 

pseudoargentata Saha and Das Gupta, 1996: 9. India. 

schoutedeni (Goetghebuer), 1933e: 150 (Palpomyia). Democratic Republic of the Congo. 

scriba Debenham, 1974: 39. Papua New Guinea. 

sejuncta (Kieffer), 1916a: 90 (Sphaeromias). Taiwan. 

uncinata (Kieffer), 1916a: 91 (Sphaeromias). Taiwan. 

Genus LANATOMYIA Debenham 

LANATOMYIA Debenham, 1974: 43. Type species: Lanatomyia miles Debenham, by original designation. 

dichroa (Edwards), 1933c: 253 (Dicrohelea). Malaysia. 

146

electra Debenham, 1974: 46. Australia (Northern Territory). 

miles Debenham, 1974: 44. Australia (Capital Territory). 

Genus LANEHELEA Wirth and Blanton 

LANEHELEA Wirth and Blanton, 1972a: 433. Type species: Lanehelea leei Wirth and Blanton, by original designation. 

leei Wirth and Blanton, 1972a: 434. Colombia. 

spinifemur Wirth and Blanton, 1972a: 436. Colombia. 

Genus LEEHELEA Debenham 

LEEHELEA Debenham, 1974: 54. Type species: Homohelea hollandiensis Tokunaga, by original designation. 

brevidensis Nandi, Mazumdar and Chaudhuri, 2012: 53. India. 

fuscilateralis Nandi, Mazumdar and Chaudhuri, 2012: 55. India. 

hispida Debenham, 1974: 57. Australia (New South Wales). 

hollandiensis (Tokunaga), 1966a: 123 (Homohelea). Indonesia. 

leucophaeata Debenham, 1974: 60. Australia (Queensland). 

magnidentata Nandi, Mazumdar and Chaudhuri, 2012: 57. India. 

parvicapusula Nandi, Mazumdar and Chaudhuri, 2012: 58. India. 

punctipes (Macfie), 1934c: 287 (Sphaeromias). Malaysia. 

wasselli Debenham, 1974: 58. Australia (Queensland). 

Genus MACKERRASOMYIA Debenham 

MACKERRASOMYIA Debenham, 1970b: 140. Type species: Heteromyia brevibarba Kieffer, by original designation. 

brevibarba (Kieffer), 1917a: 192 (Heteromyia). Australia (Queensland). 

caesia (Macfie), 1934c: 288 (1934d: 224) (Palpomyia). Malaysia. 

indica (Kieffer), 1913d: 183 (Heteromyia). India. 

magna (Das Gupta and Wirth), 1971: 880 (Neosphaeromias). Viet Nam. 

marginata Debenham, 1970b: 143. Australia (Queensland). 

pingxiangensis Yu, in Yu et al., 2005: 1500. China. 

wongsirii Wirth and Ratanaworabhan, 1981b: 404. Thailand. 

zumpti de Meillon and Wirth, 1979a: 182. Zimbabwe. 

Genus MACROPEZA Meigen 

MACROPEZA Meigen, 1818: 87. Type species: Macropeza albitarsis Meigen, by monotypy. 

MACROPTILUM Becker, 1903: 76. Type species: Macroptilum nudum Becker, by monotypy. 

HAASIELLA Kieffer, 1913d: 190. Type species: Haasiella semiflava Kieffer, by original designation. 

abonnenci (Clastrier), 1958c: 495 (Bezzia). Senegal. 

aethiopica (Ingram and Macfie), 1923: 45 (Macroptilum). South Africa. 

meeseri (de Meillon), 1936: 204 (Macroptilum). South Africa. 

affinis Macfie, 1939d: 10. South Africa. 

albitarsis Meigen, 1818: 87. Europe. 

valvata (Winnertz), 1852: 72 (Ceratopogon). Germany. 

bayeri (de Meillon), 1937b: 351 (Macroptilum). South Africa. 

blantoni Wirth and Ratanaworabhan, 1972d: 216. USA (Florida). 

calcipennis (Macfie), 1939d: 11 (Macroptilum). Sierra Leone. 

capensis Macfie, 1939d: 10. South Africa. 

147

fluviatilis (de Meillon), 1940: 463 (Macroptilum). South Africa. 

geari de Meillon and Wirth, 1981b: 547. South Africa. 

insignipennis (Macfie), 1939d: 11 (Macroptilum). Sierra Leone. 

longipes (Séguy), 1931b: 645 (Macroptilum, as Macroptylum). Mozambique. 

natalensis (de Meillon), 1937b: 355 (Macroptilum). South Africa. 

navasi (Séguy), 1934: 10 (Macroptilum, as Macroptylum). Spain. 

nigeriae (Ingram and Macfie), 1923: 43 (Macroptilum). Nigeria. 

nigra (Séguy), 1931b: 646 (Macroptilum, as Macroptylum). Mozambique. 

nuda (Becker), 1903: 77 (Macroptilum). Egypt. 

pallidipes (Séguy), 1931b: 646 (Macroptilum, as Macroptylum). Madagascar. 

pamunkeiana Knausenberger and Wirth, 1980: 128. USA (Virginia). 

semiflava (Kieffer), 1913d: 190 (Haasiella). India. 

senegalensis (Clastrier), 1958b: 239 (Bezzia). Senegal. 

sibayae (de Meillon), 1936: 183 (Bezzia). South Africa. 

stephensi (Ingram and Macfie), 1921: 369 (Probezzia). Ghana. 

Genus MALLOCHOHELEA Wirth 

MALLOCHOHELEA Wirth, 1962: 278. Type species: Johannsenomyia albibasis Malloch, by original designation. 

aenipes (Macfie), 1940c: 193 (Bezzia). Guyana. 

albibasis (Malloch), 1915b: 315 (Johannsenomyia). USA (Illinois). 

albiclava (Kieffer), 1921g: 566 (Dicrohelea). Taiwan. 

albihalter Wirth, 1962: 280. USA (Michigan). 

alpina (Clastrier), 1962b: 282 (Johannsenomyia). Switzerland. 

aspera Debenham, 1974: 88. Australia (Tasmania). 

atripes Wirth, 1962: 281. USA (New Jersey). 

aukurabis de Meillon and Wirth, 1983a: 371. South Africa. 

australiensis (Lee), 1948d: 67 (Johannsenomyia). Australia (Capital Territory). 

boettcheri (Edwards), 1929b: 11 (Palpomyia). Philippines. 

borneana (Macfie), 1934c: 289 (Palpomyia). Malaysia. 

caudellii (Coquillett), 1905: 63 (Ceratopogon). Canada (British Columbia). 

errinae (de Meillon), 1940: 461 (Palpomyia). South Africa. 

flavidula (Malloch), 1914a: 230 (Johannseniella). USA (Illinois). 

fluminea de Meillon and Wirth, 1981b: 550. South Africa. 

hamata de Meillon and Wirth, 1987a: 60. Madagascar. 

hansfordi de Meillon and Wirth, 1983a: 372. South Africa. 

hardyi (Tokunaga), 1966a: 139 (Nilobezzia). Papua New Guinea. 

inermis (Kieffer), 1909b: 38 (Johannseniella). Germany. 

kiefferi (Goetghebuer), 1921: 182 (Johannsenomyia). Belgium. 

kirki (Macfie), 1939a: 92 (Sphaeromias). Kenya. 

limitrofe Spinelli and Felippe-Bauer, 1990a: 87. Argentina. 

luaboensis (de Meillon), 1959b: 3 (Johannsenomyia). Mozambique. 

martae Szadziewski, 2005: 364. Russia. Eocene. 

microcera (Kieffer), 1915c: 287 (Palpomyia). Denmark. 

munda (Loew), 1864: 381 (Ceratopogon). Germany. 

breviforceps (Kieffer), 1915c: 289 (Palpomyia). Denmark. 

dentata (Kieffer), 1909b: 38 (Johannseniella). Germany. 

nemoralis (Macfie), 1940c: 190 (Johannsenomyia). Guyana. 

nigripes (Macfie), 1939c: 216 (Johannsenomyia). Brazil. 

nitida (Macquart), 1826: 178 (Ceratopogon). France. 

clavicornis (Kieffer), 1919a: 88 (Sphaeromias, as variety of nitida). Slovak Republic. 

longiforceps (Kieffer), 1919a: 88 (Sphaeromias, as variety of nitida). Denmark. 

prominens (Johannsen), 1932: 435 (Johannsenomyia). Indonesia. 

148

pullata (Wirth), 1952a: 213 (Johannsenomyia). USA (California). 

bicellii (Lane), 1961b: 449 (Nilobezzia). Brazil. 

remota (Kieffer), 1919a: 87 (Sphaeromias, as variety of nitida). Slovak Republic. 

bulgarica (Zilahi-Sebess), 1934: 157 (Sphaeromias, as variety of nitida). Bulgaria. 

dimidiata (Kieffer), 1921a: 59 (Sphaeromias). France. 

sabroskyi (Tokunaga), in Tokunaga and Murachi, 1959: 412 (Palpomyia). Belau (USA). 

sanctaeluciae (de Meillon), 1937b: 365 (Palpomyia). South Africa. 

satelles Debenham, 1974: 86. Australia (New South Wales). 

scandinaviae (Clastrier), 1962b: 286 (Johannsenomyia). Norway. 

senex Debenham, 1974: 89. Australia (Queensland). 

setigera (Loew), 1864: 380 (Ceratopogon). Germany. 

maihensis Remm, 1971: 209. Russia. 

shibuyai (Tokunaga), 1940b: 277 (Johannsenomyia). Japan. 

sidis (de Meillon), 1959b: 5 (Johannsenomyia). South Africa. 

silvicola (Goetghebuer), 1920: 75 (Johannsenomyia). Belgium. 

siricis (de Meillon), 1961: 50 (Sphaeromias). Madagascar. 

smithi (Lewis), 1956: 47 (Johannsenomyia). USA (Connecticut). 

spinipes Wirth, 1962: 282. USA (Georgia). 

stygia Debenham, 1974: 85. New name for nigra. 

nigra (Tokunaga), 1966a: 124 (Johannsenomyia, preoccupied by Johannsenomyia nigra Goetghebuer, 1933e). 

Papua New Guinea. 

sybleae (Wirth), 1952a: 212 (Johannsenomyia). USA (California). 

termophila (Spinelli), 1984: 197 (Neobezzia). Argentina. 

texensis Wirth, 1962: 283. USA (Texas). 

tianshanica Remm, 1980: 85. Kyrgyzstan. 

tumidicornis Debenham, 1974: 87. Australia (Queensland). 

turneri (Ingram and Macfie), 1923: 63 (Sphaeromias). South Africa. 

unca de Meillon and Wirth, 1987a: 61. Kenya. 

variegata Wirth, 1962: 283. USA (Michigan). 

vernalis Remm, 1965: 185. Estonia. 

yanana Yu and Wu, in Yu et al., 2005: 1501. China. 

yunnana Yu and Zou, in Yu et al., 2005: 1503. China. 

Genus NEOBEZZIA Wirth and Ratanaworabhan 

NEOBEZZIA Wirth and Ratanaworabhan, 1972a: 477. Type species: Neobezzia clavipes Wirth and Ratanaworabhan, by 


albitarsis Wirth and Ratanaworabhan, 1972a: 478. Brazil. 

amnicola (Macfie), 1940d: 30 (Bezzia). Guyana. 

blantoni Wirth and Ratanaworabhan, 1972a: 482. Panama. 

brasiliae (Lane), 1961a: 37 (Macropeza). Brazil. 

clavipes Wirth and Ratanaworabhan, 1972a: 485. Brazil. 

costaricae Wirth and Ratanaworabhan, 1972a: 486. Costa Rica. 

fittkaui Wirth and Ratanaworabhan, 1972a: 489. Brazil. 

wirthi Spinelli and Felippe-Bauer, 1990a: 88. Argentina. 

Genus NEOSPHAEROMIAS Das Gupta and Wirth 

NEOSPHAEROMIAS Das Gupta and Wirth, 1971: 875. Type species: Neosphaeromias gibbus Das Gupta and Wirth, by 


afrotropicalis Clastrier, 1983a: 268. Guinea. 

choriodes Grogan and Wirth, 1982: 198. Solomon Islands. 

149

gibbus Das Gupta and Wirth, 1971: 877. Thailand. 

niger Das Gupta and Wirth, 1971: 879. Sri Lanka. 

Genus NILOBEZZIA Kieffer 

NILOBEZZIA Kieffer, 1921b: 24. Type species: Nilobezzia armata Kieffer, by monotypy. 

CRESPINIA Kieffer, 1923b: 141. Type species: Crespinia brevipalpis Kieffer, by monotypy. 

PARROTIA Kieffer, 1923b: 140. Type species: Parrotia flaviventris Kieffer (= Nilobezzia kiefferi Wirth), by original 

designation. 

SPHAEROBEZZIA Zilahi-Sebess, 1940: 108 (as subgenus of Bezzia). Type species: Bezzia paradoxa Zilahi-Sebess (= 

Ceratopogon formosa Loew), by monotypy. 

acanthopus (de Meijere), 1907: 215 (Ceratopogon). Indonesia. 

conspicua (Johannsen), 1932: 441 (Bezzia, as variety of raphaelis). Indonesia. 

albilata Das Gupta, in Mazumdar, et al., 2009: 132. India. 

albipennis (Kieffer), 1911a: 124 (Bezzia, as variety of nigricans). India. 

allotropica (Kieffer), 1913d: 193 (Bezzia). India. 

animifascia Das Gupta, in Mazumdar, et al., 2009: 133. India. 

aranea Debenham, 1974: 72. Australia (New South Wales). 

arciscuta Das Gupta, in Mazumdar, et al., 2009: 135. India. 

arcuatipes (Kieffer), 1913d: 195 (Bezzia, as variety of hamifera). Indonesia. 

armata Kieffer, 1921b: 24. Sudan. 

albipennis Kieffer, 1921b: 25 (as variety of armata, preoccupied by Nilobezzia albipennis (Kieffer), 1911a). Sudan. 

flaviventris Kieffer, 1921b: 25 (as variety of armata, preoccupied by Nilobezzia flaviventris (Kieffer), 1910). Sudan. 

atoporna Yu and Zhang, 1997: 307. China. 

atroxifemorata Das Gupta, in Mazumdar, et al., 2009: 137. India. 

badia (Johannsen), 1932: 442 (Bezzia, as variety of flaviventris). Indonesia. 

bakeri (Kieffer), 1921g: 570 (Probezzia). Philippines. 

basispinigera Debenham, 1974: 74. Australia (Queensland). 

belligera (de Meillon), 1940: 459 (Dicrobezzia). South Africa. 

borkenti Das Gupta, in Mazumdar, et al., 2009: 139. India. 

brevicornis (Wirth), 1952a: 215 (Sphaeromias). USA (California). 

brevipalpis (Kieffer), 1923b: 141 (Crespinia). Indonesia. 

claripennis (Kieffer), 1916a: 97 (Bezzia). Taiwan. 

curticornis (Kieffer), 1917a: 196 (Bezzia). Australia (New South Wales). 

triquetrinotata Debenham, 1974: 64. Australia (New South Wales). 

curvopennis Yu, in Yu et al., 2005: 1506. China. 

diffidens (Johannsen), 1932: 441 (Bezzia, as variety of ochriventris. Indonesia. 

discritus Yu, in Yu et al., 2005: 1507. China. 

disjuncta (Kieffer), 1913d: 189 (Palpomyia). India. 

duodenalis Liu, Yan and Liu, 1996: 46. China. 

facialis (Kieffer), 1910: 206 (Bezzia). India. 

fijiensis Wirth and Giles, 1990: 456. Fiji. 

flavida Remm, 1980: 86. Tajikistan. 

flaviventris (Kieffer), 1910: 204 (Bezzia). Nepal. 

formosa (Loew), 1869a: 1 (Ceratopogon). Hungary. 

paradoxa (Zilahi-Sebess), 1940: 108 (Bezzia). Hungary. 

formosana (Kieffer), 1912a: 30 (Bezzia). Taiwan. 

fusca Kieffer, 1921b: 25 (as variety of armata). Sudan. 

fuscitarsus Debenham, 1974: 78. Australia (Queensland). 

hamifera (Kieffer), 1913d: 194 (Bezzia). India. 

henanei Clastrier, 1962a: 121. Algeria. 

hunyani de Meillon, 1943: 111. Zimbabwe. 

japana (Tokunaga), 1939a: 284 (Bezzia). Japan. 

150

kerteszi (Kieffer), 1916a: 97 (Bezzia). Taiwan. 

kiefferi Wirth, 1973: 383. New name for flaviventris. 

flaviventris (Kieffer), 1923b: 140 (Parrotia, preoccupied by Nilobezzia flaviventris (Kieffer), 1910). Indonesia. 

lacteipennis (Kieffer), 1910: 206 (Bezzia). India. 

leucothrix Remm, 1980: 88. Turkmenistan. 


magnitheca Das Gupta, in Mazumdar, et al., 2009: 141. India. 

mallochi Wirth, 1962: 284. USA (Michigan). 

manicata Clastrier, 1958c: 500. Senegal. 

minor (Wirth), 1952a: 216 (Sphaeromias). USA (California). 

neotropica (Macfie), 1940a: 78 (Bezzia). Brazil. 

nigra Sen and Das Gupta, 1958a: 69. India. 

nigricans (Kieffer), 1910: 207 (Bezzia). India. 

nigritibialis (Ingram and Macfie), 1921: 371 (Dicrobezzia). Ghana. 

ningxia Yu, in Yu et al., 2005: 1510. China. 

nilotica (Kieffer), 1925e: 263 (Parrotia). Egypt. 

nipponensis (Tokunaga), 1939a: 287 (Bezzia). Japan. 

ochriventris Edwards, 1929b: 12. Philippines. 

opaca Das Gupta, in Mazumdar, et al., 2009: 142. India. 

ordospina Das Gupta, in Mazumdar, et al., 2009: 144. India. 

paraensis (Lane), 1958: 33 (Bezzia). Brazil. 

posticata (Zetterstedt), 1850: 3658 (Ceratopogon). Norway. 

schineri (Kieffer), 1919a: 93 (Palpomyia). Austria. 

grisea (Zilahi-Sebess), 1936b: 44 (Sphaeromias). Hungary. 

raphaelis (Salm), 1917b: 135 (Ceratopogon). New name for blanchardi. 

blanchardi (Salm), 1917a: 106 (Ceratopogon, preoccupied by Ceratopogon blanchardi Iches, 1906). Indonesia. 

nigriventris (Kieffer), 1923b: 141 (Parrotia). Indonesia. 

robusta (de Meillon), 1937b: 346 (Bezzia). South Africa. 

capensis de Meillon and Hardy, 1954: 70. South Africa. 

schwarzii (Coquillett), 1901a: 605 (Ceratopogon). USA (Texas). 

setipes (Coquillett), 1905: 59 (Ceratopogon). USA (Texas). 

banksi (Gerry), 1933: 94 (Bezzia). Cuba. 

brasiliensis (Lane), 1958: 28 (Bezzia). Brazil. 

scotti (Kieffer), 1911c: 348 (Probezzia). Seychelles. 

longipennis (Kieffer), 1911c: 349 (Probezzia). Seychelles. 

semirufa (Kieffer), 1922b: 161 (Probezzia). Taiwan. 

setoensis (Tokunaga), 1939a: 286 (Bezzia). Japan. 

simplicior Debenham, 1974: 77. Australia (New South Wales). 

subtilicrinis Debenham, 1974: 80. Australia (Northern Territory). 

tenuicolor Das Gupta, in Mazumdar, et al., 2009: 146. India. 

typica Das Gupta, in Mazumdar, et al., 2009: 147. India. 

vaga (Kieffer), 1911b: 329 (Bezzia). Nepal. 

virago Debenham, 1974: 68. Australia (Northern Territory). 

whartoni Lee, 1948b: 343. Papua New Guinea. 

yasumatsui Wirth and Ratanaworabhan, 1981b: 409. Thailand. 

zibanensis Clastrier, 1962a: 123. Algeria. 

Genus NIPHANOHELEA Grogan and Wirth 

NIPHANOHELEA Grogan and Wirth, 1981d: 201. Type species: Niphanohelea bannae Grogan and Wirth, by original 

designation. 

bannae Grogan and Wirth, 1981d: 202. Thailand. 

151

Genus PROBEZZIA Kieffer 

PROBEZZIA Kieffer, 1906a: 57 (as subgenus of Bezzia). Type species: Ceratopogon venustus Meigen (= Tipula seminigra 

Panzer), designation by Coquillett, 1910: 594. 

DICROBEZZIA Kieffer, 1919a: 127. Type species: Ceratopogon venustus Meigen (= Tipula seminigra Panzer), by original 

designation. 

albitibia Wirth, 1971a: 732. USA (Virginia). 

albiventris (Loew), 1861: 311 (Ceratopogon). USA (Georgia). 

atriventris Wirth, 1951b: 31. USA (Michigan). 

baptosmixa Yu, in Yu et al., 2005: 1513. China. 

bottimeri Wirth, 1971a: 734. USA (Texas). 

concinna (Meigen), 1818: 77 (Ceratopogon). Europe. 

fairchildi Wirth, 1994b: 142. USA (Florida). 

flavonigra (Coquillett), 1905: 60 (Ceratopogon). Canada (British Columbia). 

flavipeda Yu and Zhang, in Yu et al., 2005: 1514. China. 

fuscipennis Wirth, 1971a: 735. USA (Maine). 

glicki Wirth, 1994b: 142. USA (Alabama). 

infuscata Malloch, 1915b: 316. USA (Illinois). 

jamnbacki Wirth, 1971a: 735. USA (Michigan). 

ludoviciana Wirth, 1951b: 28. USA (Louisiana). 

manshurica Remm, 1993: 180. Russia. 

meadi Wirth, 1994b: 143. USA (Florida). 

nigra Wirth, 1971a: 736. USA (Louisiana). 

pallida Malloch, 1914b: 138. USA (Illinois). 

rosewalli Wirth, 1951b: 32. USA (Louisiana). 

sabroskyi Wirth, 1951b: 31. USA (Michigan). 

seminigra (Panzer), 1798: plate 10 (Tipula). Germany. 

varipes Curtis, 1829: plate 285. Europe. 

munda (Loew), 1869a: 3 (Ceratopogon, preoccupied by Mallochohelea munda (Loew), 1864). Europe. 

venusta (Meigen), 1818: 78 (Ceratopogon). Austria. 

inflata (Winnertz), 1852: 71 (Ceratopogon). Germany. 

spinosidorsum Kieffer, 1921a: 61. Poland. 

borealis (Clastrier), 1962a: 119 (Bezzia). Finland. 

smithii (Coquillett), 1901a: 600 (Ceratopogon). USA (New Jersey). 

sugiyama (Tokunaga), 1940b: 282 (Dicrobezzia). Japan. 

unica (Johannsen), 1934: 345 (Lasiobezzia). USA (New York). 

weemsi Wirth, 1994b: 143. USA (Florida). 

williamsi Wirth, 1971a: 737. USA (Michigan). 

wirthi Spinelli and Grogan, 1997: 230. USA (Florida). 

xanthogaster (Kieffer), 1917b: 329 (Bezzia). New name for elegans. 

elegans (Coquillett), 1901a: 599 (Ceratopogon, preoccupied by Sphaeromias elegans (Winnertz), 1852). USA (New 

Jersey). 

Genus SINICOHELEA Yu, Wang and Tan 

SINICOHELEA Yu, Wang and Tan, in Wang et al., 2012: 42. Type species: Sinicohelea xuanjui, by original designation. 

xuanjui Yu, Wang and Chen, in Wang et al., 2012: 43. China. 

Genus SPHAEROHELEA Spinelli and Felippe-Bauer 

SPHAEROHELEA Spinelli and Felippe-Bauer, 1990b: 195. Type species: Sphaerohelea biestroi Spinelli and Felippe-Bauer, 

152

y original designation. 

biestroi Spinelli and Felippe-Bauer, 1990b: 197. Argentina. 

Genus SPHAEROMIAS Curtis 

SPHAEROMIAS Curtis, 1829: plate 285. Type species: Sphaeromias albomarginatus Curtis (= Ceratopogon fasciatus 

Meigen), by original designation. 

XYLOCRYPTA Kieffer, 1899: 69. Type species: Ceratopogon fasciatus Meigen, by original designation. 

bifidus Wirth and Grogan, 1979: 893. USA (Maryland). 

brevispinus (Kieffer), 1911a: 117 (Palpomyia). India. 

cinereus (Kieffer), 1910: 197 (Palpomyia). India. 

conjunctus Kieffer, 1916a: 93. Taiwan. 

connexus Kieffer, 1916a: 92. Taiwan. 

corsoni (Ingram and Macfie), 1924c: 182 (Ankistrodactylus). Ghana. 

decoratus Debenham, 1974: 54. Australia (Queensland). 

discolor (de Meijere), 1907: 214 (Ceratopogon). Indonesia. 

javanensis Macfie, 1934c: 285. Indonesia. 

distictus (Kieffer), 1911a: 115 (Palpomyia). India. 

eugenei de Meillon and Wirth, 1987a: 62. Zimbabwe. 

fasciatus (Meigen), 1804: 30 (Ceratopogon). Europe. 

cingulatus (Meigen), 1804: 30 (Ceratopogon). Europe. 

albomarginatus Curtis, 1829: plate 285. Great Britain. 

procerus (Zetterstedt), 1855: 4867 (Ceratopogon). Sweden. 

ocularis (Kieffer), 1924c: 397 (Xylocrypta). Germany. 

goetghebueri de Meijere, 1946: 9. Netherlands. 

inermipes Kieffer, 1916b: 115. Taiwan. 

inermithorax (Kieffer), 1918a: 99 (Palpomyia). Singapore. 

lineus Mazumdar and Chaudhuri, 2009b: 286. India. 

litoraureus Ingram and Macfie, 1921: 359. Ghana. 

longipennis (Loew), 1861: 313 (Ceratopogon). USA (Pennsylvania). 

meeseri de Meillon, 1942a: 95. Zimbabwe. 

ornatipennis (Goetghebuer), 1933b: 113 (Sphaeromias). Russia. 

nubeculosus (Tokunaga), 1941a: 100 (Palpomyia). China. 

ornatipes Mazumdar and Chaudhuri, 2009b: 287. India. 

par (Ingram and Macfie), 1922: 272 (Ankistrodactylus). Ghana. 

servilia de Meillon, 1943: 25. Mozambique. 

photophilus (Kieffer), 1913d: 188 (Palpomyia). India. 

pictipes (Kieffer), 1910: 199 (Palpomyia). India. 

pictus (Meigen), 1818: 80 (Ceratopogon). Germany. 

punctatus (Meigen), 1830: 264 (Ceratopogon, preoccupied by Culicoides punctatus (Meigen), 1804). Europe. 

elegans (Winnertz), 1852: 58 (Ceratopogon). Poland. 

candidatus (Loew), 1856: 23 (Ceratopogon, preoccupied by Ceratopogon candidatus Winnertz, 1852). Europe. 

miricornis (Kieffer), 1919a: 77 (Xylocrypta). Hungary. 

copiosus (Kieffer), 1925a: 423 (Xylocrypta). Czech Republic. 

sparus Borkent, in Borkent and Wirth, 1997:123. New name for punctata Meigen, 1830. 

pistiae (Ingram and Macfie), 1922: 274 (Palpomyia). Ghana. 

prominens Mazumdar and Chaudhuri, 2009b: 287. India. 

simililineus Mazumdar and Chaudhuri, 2009b: 287. India. 

spiniferus (Kieffer), 1916a: 94 (Palpomyia). Taiwan. 

stictonotus Kieffer, 1911a: 118. India. 

tetrastictus (Kieffer), 1911a: 116 (Palpomyia). India. 

theileri de Meillon and Wirth, 1981b: 552. South Africa. 

153

verbosus Mazumdar and Chaudhuri, 2009b: 289. India. 

Genus WANNOHELEA Yu 

WANNOHELEA Yu, in Yu et al., 2005: 1517. Type species: Wannohelea huoqiuensis Yu, by original designation. 

huoqiuensis Yu, in Yu et al., 2005: 1517. China. 

Genus XENOHELEA Kieffer 

XENOHELEA Kieffer, 1917b: 295. Type species: Xenohelea pruinosa Kieffer, by original designation. 

MIXOHELEA Kieffer, 1917b: 364. Type species: Palpomyia pulchripes Kieffer, designation by Wirth, 1973: 384. 

albinervis (Kieffer), 1913d: 187 (Palpomyia). India. 

ciliaticrus (Kieffer), 1921g: 568 (Mixohelea). Taiwan. 

galatea de Meillon, 1942c: 23. Mozambique. 

inaequalis (Kieffer), 1910: 200 (Palpomyia). India. 

luteinervis (de Meijere), 1907: 214 (Ceratopogon). Indonesia. 

nuansriae Wirth and Ratanaworabhan, 1981b: 411. Thailand. 

polydora Macfie, 1934d: 226. Indonesia. 

polysticta (Kieffer), 1911a: 117 (Palpomyia). India. 

pruinosa Kieffer, 1917b: 295 (1918a: 96). India. 

pulchripes (Kieffer), 1910: 198 (Palpomyia). India. 

roseiventris (Kieffer), 1911a: 114 (Palpomyia). India. 

rufiventris (Kieffer), 1910: 208 (Bezzia). India. 

spinosipes (Goetghebuer), 1948b: 13 (Palpomyia). Democratic Republic of the Congo. 

spinosus Liu, Yan and Liu, 1996: 46. China. 

TRIBE PALPOMYIINI Enderlein, 1936: 49 

Genus AMEROHELEA Grogan and Wirth 

AMEROHELEA Grogan and Wirth, 1981a: 1280. Type species: Amerohelea galindoi Grogan and Wirth, by original 

designation. 

dalcyi Grogan and Wirth, 1981a: 1289. Brazil. 

fasciata Grogan and Wirth, 1981a: 1283. Belize. 

frontispina (Dow and Turner), 1976: 138 (Bezzia). USA (Texas). 

galindoi Grogan and Wirth, 1981a: 1294. Colombia. 

nelsoni Grogan and Wirth, 1981a: 1290. Brazil. 

paranaensis Gaddi, Spinelli and Grogan in Gaddi et al., 2011: 301. Argentina. 

pseudofasciata Grogan and Wirth, 1981a: 1286. Brazil. 

ronderosi Grogan and Wirth, 1981a: 1298. Colombia. 

similis Spinelli, 1989: 27. Uruguay. 

sordidipes (Macfie), 1939c: 209 (Palpomyia). Brazil. 

spinellii Grogan and Wirth, 1981a: 1292. Colombia. 

vargasi Grogan and Wirth, 1981a: 1296. Belize. 

xerophila Gaddi, Spinelli and Grogan in Gaddi et al., 2011: 305. Argentina. 

Genus BEZZIA Kieffer 

BEZZIA Kieffer, 1899: 69. Type species: Ceratopogon ornatus Meigen, by original designation. 

PSEUDOBEZZIA Malloch, 1915a: 351. Type species: Ceratopogon expolitus Coquillett, by original designation. 

154

ALLOBEZZIA Kieffer, 1917b: 296. Type species: Ceratopogon expolitus Coquillett, by original designation. 

LASIOBEZZIA Kieffer, 1925d: 54. Type species: Bezzia pilipennis Lundström, by original designation. 

HOMOBEZZIA Macfie, 1932b: 496. Type species: Homobezzia nyasae Macfie, by monotypy. 

SIVABEZZIA Remm, 1974a: 440 (as subgenus of Bezzia). Type species: Bezzia campanai Clastrier, by original designation. 

PYGOBEZZIA Remm, 1974a: 441 (as subgenus of Bezzia). Type species: Bezzia strobli Kieffer (= Ceratopogon albicornis 

Meigen), by original designation. 

ASPINABEZZIA Dow and Turner, 1976: 122 (as subgenus of Bezzia). Type species: Ceratopogon glaber Coquillett, by 


acuta Remm, 1974b: 891. Estonia. 

adamsi Tokunaga and Murachi, 1959: 418. Micronesia. 

aegyptia Kieffer, 1925e: 264. Egypt. 

affinis (Staeger), 1839: 595 (Ceratopogon). Denmark. 

africana Ingram and Macfie, 1923: 71. South Africa. 

hopkinsi de Meillon and Hardy, 1954: 62. Cameroon. 

aitkeni Spinelli and Wirth, 1991: 5. Brazil. 

aklavikensis Wirth and Grogan, 1983: 495. Canada (Northwest Territories). 

albicornis (Meigen), 1818: 74 (Ceratopogon). Europe. 

pallidetarsata (Strobl), 1900: 171 (Ceratopogon). Spain. 

strobli Kieffer, 1919a: 122. Serbia-Montenegro. 

brevinervis Kieffer, 1919a: 122. Hungary. 

atrata (Macfie), 1944a: 126 (Homobezzia). Egypt. 

albidorsata Malloch, 1915a: 349. USA (Illinois). 

albuquerquei Lane, 1961a: 43. Brazil. 

aldanica Remm, 1974b: 893. Russia. 

algeriana Clastrier, 1962a: 89. Algeria. 

alia Liu and Yu, in Yu et al., 2005: 1525. China. 

amana de Meillon and Wirth, 1981b: 554. South Africa. 

ammossovi Remm, 1974b: 894. Russia. 

ampla Saha and Chaudhuri, in Saha et al., 2009: 2. India. 

analis Kieffer, 1913d: 196 (as variety of gracilipes). India. 

andersonorum Wirth and Grogan, 1983: 497. USA (Maryland). 

angulata Remm, 1974b: 900. Tajikistan. 

annulipes (Meigen), 1830: 264 (Ceratopogon). Europe. 

bidentata Kieffer, 1901a: 162. France. 

fossicola Kieffer, 1912b: 102. Germany. 

ploenensis Kieffer, 1921a: 62. Germany. 

media Kieffer, 1925d: 126 (preoccupied by Bezzia media (Coquillett), 1904b). Belgium. 

phragmitis Kieffer, 1925a: 428. Germany. 

digramma Kieffer, 1925a: 428. Poland. 


sicarti Clastrier, 1962a: 83. France. 

apicata Malloch, 1914c: 284. USA (Illinois). 

araucana Spinelli and Wirth, 1990: 14. Argentina. 

armatipes Kieffer, 1910: 207. India. 

atacina Clastrier, 1962a: 65. France. 

atrifemorata Clastrier, 1962a: 69. France. 

atripluma Kieffer, 1919a: 123. Hungary. 

atrovittata Remm, 1972: 87. Russia. 

australiensis Kieffer, 1917a: 197. Australia (New South Wales). 

badiifemorata Tokunaga and Murachi, 1959: 419. Micronesia. 

bargaensis Remm, 1974a: 436. Russia. 

bengalensis Kieffer, 1913d: 196 (as variety of gracilipes). India. 

biannulata Wirth, 1952a: 237. USA (California). 

bicolor (Meigen), 1804: 31 (Ceratopogon). Europe. 

155

typhae (Kieffer), 1919a: 130 (Probezzia). Czech Republic. 

brehmiana (Kieffer), 1924a: 22 (Probezzia). Austria. 

algarum (Zilahi-Sebess), 1930b: 186 (Probezzia). Hungary. 

rufithorax (Goetghebuer), 1933a: 287 (Probezzia). Germany. 

copiosa (Thomsen), 1935: 292 (Probezzia). USA (New York). 

bilineata Wirth, 1952a: 230. USA (California). 

bistorta Yu and Hao, in Yu et al., 2005: 1536. China. 

bivittata (Coquillett), 1905: 60 (Ceratopogon). USA (California). 

blandiata Remm, 1967: 32. Azerbaijan. 

blantoni Spinelli and Wirth, 1989b: 771. Honduras. 

bohemica Kieffer, 1919a: 118. Czech Republic. 

boiemica Kieffer, 1922e: 73 (as variety of taeniata). Czech Republic. 

boluensis Yu and Ma, in Yu et al., 2005: 1536. China. 

bresi Huttel and Huttel, 1951: 102. France. 

brevicornis (Kieffer), 1917b: 328 (Allobezzia). Paraguay. 

brevipennata Saha and Chaudhuri, in Saha et al., 2009: 4. India. 

brevipluma Kieffer, 1919a: 121. Hungary, Croatia. 

bromeliae Spinelli and Wirth, 1991: 9. Panama. 

bruna Yu, in Yu et al., 2005: 1526. China. 

bucina Li and Yu, 1997: 121. China. 

calceata (Walker), 1856a: 239 (Ceratopogon). Great Britain. 

trilobata Kieffer, 1922c: 237. Germany. 

calcuttensis Kieffer, 1913d: 195. India. 

campanai Clastrier, 1962a: 103. Algeria. 

capitata Wirth and Grogan, 1983: 503. Honduras. 

carioca Lane, 1958: 30. Brazil. 

catarinensis Spinelli and Wirth, 1990: 17. Brazil. 

cayoensis Spinelli and Wirth, 1991: 6. Belize. 

chelistyla Wirth and Grogan, 1983: 504. USA (Arizona). 

chilensis Spinelli and Ronderos, 2001: 752. Chile. 

chrysolopha Kieffer, 1912a: 31. Taiwan. 

chuzhouensis Yu, in Yu et al., 2005: 1556. China. 

circumdata Staeger, 1839: 596 (Ceratopogon). Denmark. 

solstitialis (Winnertz), 1852: 78 (Ceratopogon). Europe. 

hydrophila Kieffer, 1909b: 38 (1911d: 6). Germany. 

sieberti Kieffer, 1921d: 277. Latvia. 

aquatilis Goetghebuer, 1923: 105. Belgium. 

stagnalis Kieffer, 1925d: 129 (as variety of hydrophila). Germany. 

clarkei Tokunaga and Murachi, 1959: 423. Micronesia. 

clavipennis Spinelli and Wirth, 1989b: 775. Brazil. 

cockerelli Malloch, 1915a: 346. USA (Colorado). 

obscura (Malloch), 1915a: 355 (Probezzia). USA (New York). 

collessi Wirth and Ratanaworabhan, 1981b: 415. Thailand. 

conjunctivena Tokunaga, 1966a: 147. Papua New Guinea. 

coracina (Zetterstedt), 1850: 3646 (Ceratopogon). Sweden. 

albipes (Winnertz), 1852: 77 (Ceratopogon). Germany. 

fusciclava Kieffer, 1919a: 121. Hungary. 

corvina Remm, 1974b: 902. Russia. 

crista Yu, in Yu et al., 2005: 1540. China. 

curtiforceps Goetghebuer, 1929: 233 (1934a: 78). Belgium. 

cyrtonotum Remm, 1974b: 898. Tajikistan. 

datana Yu, Chen and Liang, in Yu et al., 2007: 488. China. 

decincta Edwards, 1932b: 47. Great Britain. 

demeilloni Haeselbarth, 1975: 368. Madagascar. 

156

dentata Malloch, 1914c: 284. USA (Illinois). 

dentifemur Spinelli and Wirth, 1991: 3. Colombia. 

dessarti Haeselbarth, 1980: 656. Democratic Republic of the Congo. 

dewulfi Goetghebuer, 1935d: 181. Democratic Republic of the Congo. 

dinhaiensis Yu and He, in Yu et al., 2005: 1557. China. 

diversipes Clastrier, 1958c: 490. Senegal. 

dividua Tokunaga, 1966a: 148. Indonesia. 

dorsasetula Dow and Turner, 1976: 89. USA (New York). 

downesi Dow and Turner, 1976: 93. USA (Maine). 

echinata Clastrier, 1985a: 49. New Caledonia (France). 

edwardsi de Meillon, 1938: 268. Zambia. 

elongata Zilahi-Sebess, 1940: 101. Hungary. 

eocenica Szadziewski, 1988: 187. Poland. Eocene. 

excavata Tokunaga, 1966a: 145. Papua New Guinea. 

exclamationis Kieffer, 1918a: 61. Tunisia. 

exigua Goetghebuer, 1935a: 4. Belgium. 

expedita Sinha and Dasgupta, in Sinha, et al., 2003: 24. India. 

expolita (Coquillett), 1901a: 600 (Ceratopogon). USA (New Jersey). 

johnsoni (Coquillett), 1901a: 600 (Ceratopogon). USA (New Jersey). 

fairchildi Wirth, 1983b: 303. USA (Florida). 

fangchengi Yu and Mai, in Yu et al., 2005: 1549. China. 

fascispinosa Clastrier, 1962a: 91. France. 

sexstrigata Remm, 1971: 213. Russia. 

longiradia Dow and Turner, 1976: 97. Canada (Ontario). 

fenestrata Clastrier, 1962a: 86. France. 

filiducta Spinelli and Wirth, 1991: 3. Colombia. 

flava Tokunaga, 1939a: 292. Japan. 

flavescens Kieffer, 1913d: 196 (as variety of gracilipes). India. 

flavicornis (Staeger), 1839: 599 (Ceratopogon). Denmark. 

flavipalpis (Winnertz), 1852: 80 (Ceratopogon). Germany. 

flavipluma Kieffer, 1919a: 126. Slovak Republic. 

spinifera Goetghebuer, 1920: 106. Belgium. 

spinosula Clastrier, 1962a: 100. France. 

flavicorporis de Meillon, 1939a: 22. South Africa. 

flavipennis Tokunaga, 1939a: 290. Japan. 

spinosa Tokunaga, 1940d: 161. Japan. 

flavitarsis Malloch, 1914c: 283. USA (Illinois). 

flavitibia Dow and Turner, 1976: 140. USA (New York). 

flavoscutellaris Haeselbarth, 1975: 362. Zimbabwe. 

flinti Spinelli and Wirth, 1989a: 113. Dominica. 

fluminensis Lane, 1948: 236. Brazil. 

fontana Liu, Yan and Liu, 1996: 47. China. 

fortigenitalis Sinha and Dasgupta, in Sinha, et al., 2003: 26. India. 

foyi Ingram and Macfie, 1921: 361. Nigeria. 

fuliginata Clastrier, 1962a: 116. Serbia-Montenegro. 

fusca Spinelli and Wirth, 1991: 9. Colombia. 

fuscifemoris Remm, 1971: 216. Russia. 

gibbera (Coquillett), 1905: 60 (Ceratopogon). Cuba. 

gibberella Wirth and Grogan, 1983: 508. USA (Maryland). 

glabra (Coquillett), 1902a: 85 (Ceratopogon). USA (Florida). 

glaucivena Sinha and Dasgupta, in Sinha, et al., 2003: 27. India. 

globulosa Spinelli and Wirth, 1990: 18. Puerto Rico (USA). 

goianensis Lane, 1961a: 44. Brazil. 

gracilipes (Winnertz), 1852: 72 (Ceratopogon). Germany. 

157

gressitti Tokunaga, 1966a: 149. Papua New Guinea. 

griseata Remm, 1972: 85. Russia. 

griseipes Clastrier, Rioux and Descous. 1961: 85. Chad. 

grogani Spinelli and Wirth, 1990: 19. Panama. 

hainana Liu, Yan and Liu, 1996: 48. China. 

haroldi de Meillon and Wirth, 1987a: 66. South Africa. 

hihifoi Clastrier and Delécolle, 1996: 296. Wallis and Futuna Islands (France). 

hissarica Remm, 1974b: 891. Tajikistan. 

hoggarensis Clastrier, 1962a: 81. Algeria. 

hondurensis Spinelli and Wirth, 1990: 20. Honduras. 

huochengensis Yu and Ma, in Yu et al., 2005: 1560. China. 

imbifida Dow and Turner, 1976: 38. USA (Maryland). 

indecora Kieffer, 1912c: 8. Sri Lanka. 

inflatifemora Tokunaga, 1966a: 141. Papua New Guinea. 

insolita de Meillon and Wirth, 1983a: 376. South Africa. 

insularis Kieffer, 1921g: 571. Taiwan. 

jubata Spinelli and Wirth, 1990: 20. Colombia. 

jamaicensis Spinelli and Wirth, 1989a: 115. Jamaica. 

japonica Tokunaga, 1939a: 280. Japan. 

suffusa Dow and Turner, 1976: 110. Canada (Manitoba). 

jiaonana Yu and Xue, in Yu et al., 2005: 1551. China. 

kazlauskasi Remm, 1966: 67. Lithuania. 

kelamayi Yu and Ma, in Yu et al., 2005: 1561. China. 

kempi Kieffer, 1913d: 197. India. 

kiefferiana Goetghebuer, 1934a: 80. Germany. 

kitaokai Tokunaga, 1963c: 46. Japan. 

kuhetiensis Remm, 1967: 33. Azerbaijan. 

kurensis Remm, 1967: 34. Azerbaijan. 

laciniastyla Dow and Turner, 1976: 42. USA (Florida). 

lancanga Yu, in Yu et al., 2005: 1553. China. 

latipalpis Clastrier, 1962a: 106. Tunisia. 

leei Spinelli and Wirth, 1990: 21. Colombia. 

lenkoi Lane, 1958: 31. Brazil. 

leucogaster (Zetterstedt), 1850: 3659 (Ceratopogon). Sweden. 

xanthocephala Goetghebuer, 1911: 95. Belgium. 

parvidens Kieffer, 1914a: 240. Sweden. 

picticornis Kieffer, 1914a: 241. Germany. 

danica Kieffer, 1915c: 291. Denmark. 

grisea Kieffer, 1919a: 119. Hungary. 

brachycera Kieffer, 1919a: 119. Hungary. 

belgica Kieffer, 1919a: 120. Belgium. 

hungarica Zilahi-Sebess, 1930b: 190. Hungary. 

lewvanichae Wirth and Ratanaworabhan, 1981b: 422. Thailand. 

liangpinensis Yu, in Yu et al., 2005: 1527. China. 

longiforceps Tokunaga, 1959: 312. Indonesia. 

longipennis Statz, 1944: 151. Germany. Oligocene. 

longisaeta (Spataru), 1973: 287 (Probezzia). Romania. 

lophophora Clastrier, 1988a: 58. Guinea. 

lucida de Meillon, 1939a: 24. Democratic Republic of the Congo. 

lujingi Yu, Chen and Pan, in Yu et al., 2007: 488. China. 

lutea Wirth and Ratanaworabhan, 1981b: 417. Thailand. 

luteiventris Wirth and Grogan, 1983: 510. USA (Virginia). 


maculifemorata Tokunaga and Murachi, 1959: 424. Micronesia. 

158

magnisetula Dow and Turner, 1976: 46. Canada (Manitoba). 

mallochi Wirth, 1951d: 323. USA (Virginia). 

mathisi Spinelli and Wirth, 1989a: 117. Ecuador. 

mazaruni Macfie, 1940c: 193. Guyana. 

coloradensis Wirth, 1952a: 238. USA (California). 

media (Coquillett), 1904b: 166 (Ceratopogon). USA (New Jersey). 

medusa Yu, Li and Nie, in Nie et al., 2005: 103. Indonesia or China. 

megatheca Spinelli and Wirth, 1990: 22. Colombia. 

melanesiae Clastrier, 1985a: 45. New Caledonia (France). 

melanoflava Clastrier, 1958b: 235. Senegal. 

melanoflavida Clastrier and Wirth, 1961a: 205. Nigeria. 

mellori Boorman and Harten, 2002: 456. Oman. 

mesotibialis Spinelli and Wirth, 1990: 23. Belize. 

mexicana Spinelli and Wirth, 1991: 6. Mexico. 

meyensis Vattier and Adam, 1966b: 748. Congo. 

micronyx Kieffer, 1922b: 162. Taiwan. 

crassistyla Tokunaga, 1966a: 143. Indonesia. 

minuta Remm, 1974b: 895. Kyrgyzstan. 

mohave Wirth and Grogan, 1983: 511. USA (California). 

mollis Johannsen, 1932: 442. Indonesia. 

monacantha Kieffer, 1925a: 427. Russia. 

mongolica Remm, 1972: 88. Russia. 

monotheca Sinha and Dasgupta, in Sinha, et al., 2003: 29. India. 

morvani Clastrier, 1962a: 95. Algeria. 

multiannulata (Strobl), 1906: 400 (Ceratopogon). Spain. 

gandavensis Goetghebuer, 1935a: 4. Belgium. 

strigula Clastrier, 1962a: 68. France. 

multispinosa Clastrier, 1962a: 77. Algeria. 

murina Kieffer, 1922b: 162. Taiwan. 

murphyi Clastrier and Wirth, 1961a: 207. Gambia. 

myrmedon (Kieffer), 1921g: 571 (Probezzia). Taiwan. 

narynica Remm, 1973: 183. Kyrgyzstan. 

naseri Boorman and Harten, 2002: 457. Yemen. 

neixianga Yu and Guo, in Yu et al., 2005: 1529. China. 

nicator de Meillon, 1959b: 18. South Africa. 

nigerrima Haeselbarth, 1965b: 9. South Africa. 

nigrialula Tokunaga, in Tokunaga and Murachi, 1959: 428. Belau (USA). 

nigriclava Kieffer, 1921g: 572. Taiwan. 

nigripes Wirth and Grogan, 1983: 513. USA (Utah). 

nigrita Clastrier, 1962a: 117. Norway. 

nigritibialis Spinelli and Wirth, 1991: 5. Belize. 

nigritula (Zetterstedt), 1838: 820 (Ceratopogon). Norway. 

tenebricosa Goetghebuer, 1912: 208. Belgium. 

atripes Zilahi-Sebess, 1940: 100. Hungary. 

nigrofasciata Tokunaga, in Tokunaga and Murachi, 1959: 429. Micronesia. 

nigroflava Remm, 1974b: 891. Russia. 

niokoloensis Clastrier, 1958c: 493. Senegal. 

niphatoda Yu, 2000: 162. China. 

nobilis (Winnertz), 1852: 79 (Ceratopogon). Germany. 

setulosa (Loew), 1861: 312 (Ceratopogon). USA (District of Columbia). 

barberi (Coquillett), 1901a: 601 (Ceratopogon). USA (Maryland). 

punctata Lundström, 1910: 42 (as variety of leucogaster). Finland. 

leucosticta Kieffer, 1919a: 124. Russia. 

cinerella Kieffer, 1919a: 124. Hungary. 

159

armaticrus Kieffer, 1919a: 125. Hungary. 

csikiana Kieffer, 1919a: 126. Russia. 

chrysocoma Kieffer, 1922d: 355. Poland. 

acanthodes Macfie, 1940c: 192. Guyana. 

atlantica Wirth and Williams, 1957: 13. Bermuda (Great Britain). 

nobiliformis Clastrier, 1962a: 74. France. 

nodosipes Kieffer, 1924a: 22. Austria. 

numidiana Clastrier, 1962a: 72. Algeria. 

nyasae (Macfie), 1932b: 497 (Homobezzia). Malawi. 

agathae de Meillon, 1940: 455. South Africa. 

obelisca Dow and Turner, 1976: 104. USA (New York). 

omanensis Boorman and Harten, 2002: 458. Oman. 

ornata (Meigen), 1830: 262 (Ceratopogon). Europe. 

vittiger (Zetterstedt), 1850: 3642 (Ceratopogon). Sweden. 

albosignata Kieffer, 1919a: 117. Hungary. 

ornatissima (Kieffer), 1911c: 349 (Probezzia). Seychelles. 

bipunctata (Kieffer), 1911c: 350 (Probezzia, as variety of ornatissima). Seychelles. 

pachypyga Remm, 1974a: 441. Tajikistan. 

pallidipes Clastrier and Wirth, 1961a: 209. Gambia. 

palustris Clastrier, 1962a: 94. France. 

papillistyla Sinha and Dasgupta, in Sinha, et al., 2003: 31. India. 

papuae Tokunaga, 1966a: 150. Indonesia. 

pediaureola Tokunaga and Murachi, 1959: 427. Micronesia. 

perplexa Dow and Turner, 1976: 57. USA (Florida). 

picipes Goetghebuer, 1948b: 17. Democratic Republic of the Congo. 

pilipennis Lundström, 1916b: 674. Sweden. 

pilosella Remm, 1974b: 896. Kazakhstan. 

platyura (Macfie), 1947b: 77 (Palpomyia). Sudan. 

dubia Clastrier, Rioux and Descous, 1961: 81. Chad. 

propriostyla Sinha and Dasgupta, in Sinha, et al., 2003: 33. India. 

prospicula Remm, 1974a: 435. Kyrgyzstan. 

pruinosa (Coquillett), 1905: 59 (Ceratopogon). Canada (British Columbia). 

modocensis Wirth, 1952a: 233. USA (California). 

pseudobscura Wirth, 1951d: 324. USA (Virginia). 

pseudogibbera Spinelli and Wirth, 1990: 23. Panama. 

pseudovenustula Spinelli and Wirth, 1991: 8. Brazil. 

pulchripes Kieffer, 1917b: 330. Paraguay. 

pulverea (Coquillett), 1901a: 600 (Ceratopogon). USA (New Jersey). 

punctipennis (Williston), 1896: 278 (Ceratopogon). St. Vincent. 

pygmaea Goetghebuer, 1920: 100. Belgium. 

raoheensis Yu and Liu, in Yu et al., 2005: 1562. China. 

raposoensis Spinelli and Wirth, 1991: 4. Colombia. 

rhodesiensis Haeselbarth, 1975: 360. Zimbabwe. 

rhynchostylata Remm, 1974b: 891. Russia. 

riparia Clastrier, 1985a: 51. New Caledonia (France). 

roldani Spinelli and Wirth, 1981: 187. Argentina. 

rossii Clastrier, 1962a: 55. Algeria. 

rufescens Remm, 1971: 213. Russia. 

rufifascies Goetghebuer, 1932a: 126. Germany. 

rufipes (Kieffer), 1911c: 347 (Pachyleptus). Seychelles. 

femoralis (Kieffer), 1911c: 348 (Pachyleptus, as variety of rufipes). Seychelles. 

sahariensis Clastrier, 1962a: 97. Algeria. 

saileri Wirth, 1983a: 770. USA (Alaska). 

sajana Remm, 1972: 89. Russia. 

160

quadridens Remm, 1972: 89. Russia. 

sandersoni Wirth and Grogan, 1983: 514. USA (Arizona). 

schmitzorum Dippolito and Spinelli, in Dippolito et al., 1995: 54. Brazil. 

segermanae Haeselbarth, 1975: 359. South Africa. 

separata Kieffer, 1916a: 96. Taiwan. 

serena Johannsen, 1932: 443. Indonesia. 

sergenti Kieffer, 1922g: 515. Algeria. 

setigera Spinelli and Wirth, 1990: 24. Colombia. 

setosa Remm, 1974b: 892. Russia. 

setosinotum Wirth and Grogan, 1983: 517. USA (Maryland). 

sevanica Remm, 1974a: 438. Armenia. 

sexspinosa Edwards, 1928: 57. American Samoa (USA). 

shihei Yu and Li, in Yu et al., 2005: 1543. China. 

signata (Meigen), 1804: 29 (Ceratopogon). Europe. 

heusdensis Goetghebuer, 1935b: 414. Belgium. 

sinica Hao and Yu, 2003: 200. China. 

sivashica Remm, in Remm and Zhogolev, 1968: 842. Ukraine. 

snowi Lane, 1958: 34. Guatemala. 

soikai (Harant, Huttel and Huttel), 1952: 12 (Dicrobezzia). Italy. 

sordida Wirth, 1952a: 232. USA (California). 

spathula Wirth and Grogan, 1983: 518. USA (Maryland). 

spicata Dow and Turner, 1976: 77. USA (Florida). 

spinosella Clastrier, 1983c: 21. Seychelles. 

spinositibialis Tokunaga and Murachi, 1959: 421. Micronesia. 

suavis Johannsen, 1932: 444. Indonesia. 

subfusca Macfie, 1939c: 218. Brazil. 

sulfureicruris Tokunaga and Murachi, 1959: 425. Micronesia. 

tadsignata Remm, 1974b: 896. Tajikistan. 

taeniata (Haliday), in Walker, 1856a: 238 (Ceratopogon). Great Britain. 

brehmii Kieffer, 1925d: 129. France. 

tasmaniensis Lee, 1948b: 341. Australia (Tasmania). 

ternidenta Yu, 2000: 161. China. 

tenuipennis Yu and Zhang, in Yu et al., 2005: 1564. China. 

texensis Wirth and Grogan, 1983: 519. USA (Texas). 

tirawati Wirth and Ratanaworabhan, 1981b: 420. Thailand. 

transitiva Remm, 1974a: 434. Russia. 

trichia Yu and Liu, in Yu et al., 2005: 1565. China. 

trispinosa Kieffer, 1911a: 123. India. 

tshernovskii Remm, 1993: 180. Armenia. 

tuoliensis Yu and Xiang, in Yu et al., 2005: 1544. China. 

turbipes Sinha and Dasgupta, in Sinha, et al., 2003: 35. India. 

turkmenica Glukhova, 1979: 163. Turkmenistan. 

turmidifemura Yu and Xu, in Yu et al., 2005: 1545. China. 

turrita de Meillon and Wirth, 1983a: 378. South Africa. 

twinni Wirth, 1983a: 774. Canada (Manitoba). 

umlalazia de Meillon, 1940: 457. South Africa. 

uncistyla Dow and Turner, 1976: 80. USA (New Mexico). 

unispina Dow and Turner, 1976: 114. USA (Alaska). 

ussurica Glukhova, 1979: 167. Russia. 

varia Haeselbarth, 1975: 368. Zimbabwe. 

varicolor (Coquillett), 1902b: 84 (Ceratopogon). USA (New York). 

ventanensis Spinelli, in Spinelli, et al., 2012: 62. Argentina. 

venustula (Williston), 1896: 278 (Ceratopogon). St. Vincent. 

concoloripes Macfie, 1940d: 31. Guyana. 

161

vilbastei Remm, 1971: 215. Russia. 

vitilevuensis Wirth and Giles, 1990: 458. Fiji. 

vittata Tokunaga, 1966a: 146. Indonesia. 

wanyuanensis Yu, in Yu et al., 2005: 1566. China. 

winnertziana Kieffer, 1919a: 113. New name for gracilis. 

gracilis (Winnertz), 1852: 76 (Ceratopogon, preoccupied by Stilobezzia gracilis (Haliday), 1833). Germany. 

wirthi Haeselbarth, 1965b: 10. Zimbabwe. 

woodruffi Spinelli and Wirth, 1989a: 120. Jamaica. 

xanthogaster (Kieffer), 1919a: 130 (Probezzia). Russia. 

xinyangensis Yu, in Yu et al., 2005: 1547. China. 

yaeyama (Tokunaga), 1962a: 216 (Nilobezzia). Japan. 

yasumatsui Wirth and Ratanaworabhan, 1981b: 424. Thailand. 

yigonga Yu and Yan, in Yu et al., 2005: 1533. China. 

zajantshkauskasi Remm, 1966: 68. Lithuania. 

zhangmuensis Yu and Zhang, in Yu et al., 2005: 1567. China. 

zhangxingwangi Yu and Gao, in Yu et al., 2005: 1570. China. 

zhongnina Yu and Liu, in Yu et al., 2005: 1569. China. 

zonatipes Tokunaga, 1966a: 146. Indonesia. 

Genus CLASTRIEROMYIA Spinelli and Grogan 

CLASTRIEROMYIA Spinelli and Grogan, 1985: 330. Type species: Clastrieromyia schnacki Spinelli and Grogan, by original 

designation. 

dycei Spinelli and Grogan, 1986: 456. Uruguay. 

kremeri Spinelli and Grogan, 1985: 332. Brazil. 

schnacki Spinelli and Grogan, 1985: 331. Ecuador. 

uruguayensis Spinelli and Grogan, 1986: 458. Uruguay. 

Genus PACHYHELEA Wirth 

PACHYHELEA Wirth, 1959a: 50. Type species: Ceratopogon magnus Coquillett (= Ceratopogon pachymerus Williston), by 


albidiventris (Kieffer), 1917b: 316 (Sphaeromias). Colombia. 

pachymera (Williston), 1900: 224 (Ceratopogon). Mexico. 

magna (Coquillett), 1905: 61 (Ceratopogon). USA (Texas). 

latifemoris (Ingram and Macfie), 1931a: 231 (Johannsenomyia). Argentina. 

Genus PALPOMYIA Meigen 

PALPOMYIA Meigen, 1818: 82. Type species: Ceratopogon flavipes Meigen, by monotypy. Generic name first published in 

synonymy with Ceratopogon but available under ICZN Code Article 11(e). 

APOGON Rondani, 1856: 175 (preoccupied by Apogon Lacepede, 1802). Type species: Ceratopogon hortulanus Meigen (= 

Ceratopogon flavipes Meigen), by original designation. 

ALASION Rondani, 1857: 14. New name for Apogon Rondani. Type species: Ceratopogon hortulanus Meigen (= 

Ceratopogon flavipes Meigen), automatic. 

MIOPALPOMYIA Pierce, 1966: 95. Type species: Miopalpomyia shilo Pierce, by original designation. 

NEOPALPOMYIA Pierce, 1966: 95. Type species: Neopalpomyia freyi Pierce, by original designation. 

PARAPALPOMYIA Pierce, 1966: 97. Type species: Parapalpomyia ryshkoffi Pierce, by original designation. 

GLUHHOVIA Remm, 1976a: 174 (as subgenus of Palpomyia). Type species: Palpomyia miki Goetghebuer, by original 

designation. 

abdominalis Kieffer, 1922b: 161 (as variety of tainana). Taiwan. 

162

abrupta Yu, in Yu et al., 2005: 1575. China. 

aculeata Ingram and Macfie, 1931a: 215. Chile. 

afra Goetghebuer, 1948b: 14. Democratic Republic of the Congo. 

alba Yu and Liu, in Yu et al., 2005: 1576. China. 

albiditarsis Kieffer, 1910: 203. India. 

albipennis Kieffer, 1901a: 163. France. 

aldrichi (Malloch), 1915a: 326 (Heteromyia). USA (Idaho). 

algeriana Clastrier, 1962b: 244. Algeria. 

almeidai (Lane), 1946b: 219 (Sphaeromias). Brazil. 

altispina Grogan and Wirth, 1975a: 32. USA (New York). 

amplofemoria Yu and Zhang, in Yu et al., 2005: 1577. China. 

ancorifera Tokunaga, 1966a: 129. Papua New Guinea. 

angustipennis Kieffer, 1919a: 105. Hungary. 

aquilotibialis Borkent, in Borkent and Wirth, 1997:130. New name for nigrotibialis. 

nigrotibialis Remm, 1976a: 183 (preoccupied by Palpomyia nigrotibialis Goetghebuer, 1933d). Kyrgyzstan. 

arcutibia Yu, in Yu et al., 2005: 1578. China. 

arenosa (Remm and Nasarmuchamedov), 1969: 54 (Bezzia). Uzbekistan. 

arkitensis Remm, 1976a: 183. Kyrgyzstan. 

armata Kieffer, 1921a: 60. France. 

armatipes Wirth, 1952a: 222. USA (California). 

armigera de Meillon and Downes, 1986: 177. South Africa. 

armipes (Meigen), 1838: 20 (Ceratopogon). Germany. 

erythrocephala (Staeger), 1839: 579 (Ceratopogon). Denmark. 

rufipecta (Winnertz), 1852: 62 (Ceratopogon). Poland. 

ashantii Ingram and Macfie, 1923: 64. Ghana. 

aspina Grogan and Wirth, 1979a: 37. USA (Texas). 

aterrima Goetghebuer, 1921: 183. Belgium. 

atriclava Kieffer, 1921g: 569. Taiwan. 

atricoxa Kieffer, 1919a: 110. Romania. 

balozeti Clastrier, 1962b: 260. Algeria. 

barrettoi Lane, 1947b: 442. Brazil. 

basalis (Walker), 1848: 27 (Ceratopogon). USA (New York). 

trivialis (Loew), 1861: 309 (Ceratopogon). USA (District of Columbia). 

tenuicornis (Malloch), 1915a: 328 (Heteromyia). USA (Wisconsin). 

belkini Grogan and Wirth, 1979a: 72. USA (California). 

bicolor Macfie, 1941: 67. Malawi. 

bipicta Remm, 1976a: 175. Tajikistan. 

bispinosa Kieffer, 1915c: 286. Denmark. 

kiefferi Goetghebuer, 1934a: 69. New name for bispinosa. 

boliviensis Kieffer, 1917b: 324. Bolivia. 

bottimeri Grogan and Wirth, 1979a: 75. USA (Texas). 

brachialis (Haliday), 1833: 152 (Ceratopogon). Great Britain. 

longipennis Kieffer, 1919a: 104. Slovak Republic. 

fusciclava Kieffer, 1919a: 109. Slovak Republic. 

nemorivaga Goetghebuer, 1920: 84. Belgium. 



brevicornis Edwards, 1926a: 422. Great Britain. 

brevipes Goetghebuer, 1948b: 14. Democratic Republic of the Congo. 

buettikeri Boorman and Harten, 2002: 458. Saudi Arabia. 

burmae Kieffer, 1910: 196. Myanmar. 

calcarata Edwards, 1929b: 10. Philippines. 

calderana Storå, 1936: 37. Canary Islands (Spain). 

callangana Kieffer, 1917b: 323. Peru. 

163

canadensis Grogan and Wirth, 1979a: 109. USA (California). 

cantuaris Ingram and Macfie, 1931b: 207. New Zealand. 

carioca Lane, 1960: 384. Brazil. 

carrerai (Lane), 1948: 235 (Dicrobezzia, as carreirai). Brazil. 

castanea Macfie, 1939c: 211. Brazil. 

castellana (Strobl), 1900: 170 (Ceratopogon). Spain. 

catarinensis Lane, 1960: 385. Brazil. 

chongqingi Yu, in Yu et al., 2005: 1581. China. 

citrinipes Kieffer, 1922e: 72. Czech Republic. 

columbiana Kieffer, 1917b: 323. Colombia. 

concoloripes Clastrier, 1962b: 258. Norway. 

conifera Macfie, 1939c: 214. Brazil. 

coroicoensis Wirth, 1974: 54. New name for boliviensis. 

boliviensis (Kieffer), 1917b: 331 (Jenkinsia, preoccupied by Palpomyia boliviensis Kieffer, 1917b). Bolivia. 

craponnei Clastrier, 1962b: 242. France. 

crassicrus Kieffer, 1917b: 321. Paraguay. 

crassifemur Kieffer, 1918a: 98. Sri Lanka. 

festiva Kieffer, 1918a: 99 (as variety of crassifemur). Sri Lanka. 

crassinervis (de Meijere), 1907: 213 (Ceratopogon). Indonesia. 

crassipalpis Sinha, Das Gupta, and Chaudhuri, 2003b: 75. India. 

flexidigita Sinha, Das Gupta, and Chaudhuri, 2003b: 79. India. 

novita Saha and Das Gupta, 2005: 63. India. 

cressoni (Malloch), 1915a: 327 (Heteromyia). USA (Pennsylvania). 

croatica Kieffer, 1919a: 99 (as variety of serripes). Slovak Republic, Romania, Hungary. 

csikii Remm, 1981: 31. New name for longicornis. 

longicornis Kieffer, 1919a: 107 (preoccupied by Palpomyia longicornis (Williston), 1896). Romania. 

curtata Yu and Deng, in Yu et al., 2005: 1582. China. 

decima Lee, 1948d: 58. Australia (Tasmania). 

deminutipalpis Sinha, Das Gupta, and Chaudhuri, 2003b: 79. India. 

barbara Saha and Das Gupta, 2005: 62. India. 

distincta (Haliday), 1833: 152 (Ceratopogon). Great Britain. 

ephippium (Zetterstedt), 1855: 4873 (Ceratopogon). Sweden. 

rubra Kieffer, 1919a: 101. Romania. 

flaviscutellum Kieffer, 1919a: 103. Romania. 

ruficornis Goetghebuer, 1934c: 289 (as variety of distincta). Europe. 

divisa Kieffer, 1916a: 94. Taiwan. 

downesi Grogan and Wirth, 1979a: 112. Canada (Manitoba). 

ebejeri Boorman and Harten, 2002: 458. Oman. 

ectasa Yu, in Yu et al., 2005: 1588. China. 

edwardsi Cockerell, 1921: 469. Great Britain. Oligocene. 

elgoni Macfie, 1939a: 97. Kenya. 

erikae Szadziewski, 1993: 642. Germany. Eocene. 

exotica Yu and Nie, in Nie and Yu, 2006:376. China (on ship from Tomita, Japan). 

fergana Remm, 1976a: 183. Kyrgyzstan. 

flaviceps (Johannsen), 1908: 268 (Johannseniella). USA (New York). 

opacithorax (Malloch), 1915a: 329 (Heteromyia). USA (Illinois). 

hirta (Malloch), 1915a: 330 (Heteromyia). USA (Illinois). 

flavipecta Kieffer, 1918a: 59. Tunisia. 

flavipes (Meigen), 1804: 28 (Ceratopogon). Europe. 

hortulana (Meigen), 1818: 81 (Ceratopogon). Europe. 

mutabilis Clastrier, 1962b: 246. France. 

flavitarsis (Meigen), 1838: 20 (Ceratopogon). Europe. 

flavitibialis Santos Abreu, 1918: 325 (as variety of flavipes). Canary Islands (Spain). 

flaviventris (Czerny and Strobl), 1909: 130 (Ceratopogon, as variety of flavipes). Spain. 

164

floralis (Meigen), 1804: 27 (Ceratopogon). Europe. 

freyi (Pierce), 1966: 95 (Neopalpomyia). USA (California). Miocene. 

fulva Yu and Qi, in Yu et al., 2005: 1585. China. 

fulvastra Yu, in Yu et al., 2005: 1586. China. 

fulvescens Kieffer, 1919a: 100. Romania. 

fumiptera Yu and Zhang, in Yu et al., 2005: 1587. China. 

fusca de Meillon, 1938: 266. Mozambique. 

fuscipecta Kieffer, 1918a: 60. Tunisia. 

atrimana Kieffer, 1918a: 60 (as variety of fuscipectus). Tunisia. 

fuscipeda Yu and Liu, in Yu et al., 2005: 1590. China. 

fuscitibia Yu and Deng, in Yu et al., 2005: 1591. China. 

gilva de Meillon and Wirth, 1987a: 67. South Africa. 

globulifera Remm, 1971: 211. Russia. 

grossipes Goetghebuer, 1920: 88. Belgium. 

guarani Lane, 1946b: 221. Brazil. 

guyana Clastrier, 1992: 117. French Guiana. 

hastata Grogan and Wirth, 1975a: 34. USA (New York). 

helviscutellata Borkent, in Borkent and Wirth, 1997:132. New name for flavoscutellata. 

flavoscutellata (Strobl), 1900: 170 (Ceratopogon, as variety of flavipes, preoccupied by Dasyhelea flavoscutellata 

(Zetterstedt), 1850). Spain. 

himalayae (Kieffer), 1911b: 328 (Dibezzia). India. 

hispida Borkent, in Borkent and Wirth, 1997:132. New name for multispinosa. 

multispinosa Spinelli and Grogan, 1989: 6 (preoccupied by Palpomyia multispinosa (Pierce), 1966). El Salvador. 

iberaensis Spinelli and Cazorla, 2006: 1068. Argentina. 

indistincta Remm, 1976a: 180. Russia. 

indivisa Kieffer, 1916a: 95. Taiwan. 

inermicollis Kieffer, 1917b: 323. Paraguay. 

inflatifemoralis Tokunaga, 1966a: 131. Papua New Guinea. 

infuscata Kieffer, 1919a: 104. Slovak Republic, Croatia. 

insularis Spinelli and Grogan, 1989: 3. Puerto Rico (USA). 

jamnbacki Grogan and Wirth, 1979a: 88. USA (New York). 

jantari Szadziewski, 1988: 182. Poland. Eocene. 

jimmensis Tokunaga, 1966a: 126. Papua New Guinea. 

johannseni (Lane), 1948: 233 (Dicrobezzia). Brazil. 

johnstoni Macfie, 1939a: 96. Uganda. 

jonesi Grogan and Wirth, 1975a: 17. USA (Wisconsin). 

kernensis Wirth, 1952a: 223. USA (California). 

kilembei Macfie, 1939a: 94. Uganda. 

kirgizica Glukhova, 1979: 161. Kyrgyzstan. 

korni Havelka, 1980: 89. Germany. 

kurwana de Meillon and Wirth, 1987a: 69. South Africa. 


lacorum Debenham, 1989: 250. New name for abdominalis. 

abdominalis (Tokunaga), 1966a: 106 (Heteromyia, preoccupied by Palpomyia abdominalis Kieffer, 1922b). 

Indonesia. 

lacteipennis Remm, 1976a: 183. Russia. 

lacustris Lane, Forattini and Rabello, 1955: 83. Brazil. 

laensis Tokunaga, 1966a: 134. Papua New Guinea. 

lanceolifera Tokunaga, 1966a: 133. Papua New Guinea. 

langyaensis Yu, in Yu et al., 2005: 1592. China. 

leucopogon Kieffer, 1911a: 114. India. 

limnochares (Macfie), 1940a: 76 (Sphaeromias). Brazil. 

lineata (Meigen), 1804: 30 (Ceratopogon). Europe. 

cinerea (Macquart), 1826: 177 (Ceratopogon). France. 

165

inotata (Staeger), 1839: 596 (Ceratopogon). Denmark. 

slossonae (Coquillett), 1905: 61 (Ceratopogon). USA (New Hampshire). 

algarum Kieffer, in Kieffer and Thienemann, 1908: 1. Germany. 

winnertzi Kieffer, 1914a: 238. Germany. 

illinoisensis Malloch, 1914a: 219. USA (Illinois). 

octasema Kieffer, 1914a: 239. Germany. 

spinosissima Kieffer, 1915c: 285. Denmark. 

unifasciata Kieffer, 1922e: 72. Czech Republic. 

murina Kieffer, 1925a: 426. Germany. 

obesa Goetghebuer, 1927b: 94. Belgium. 

nigrotibialis Goetghebuer, 1933d: 370 (as variety of lineata). Belgium. 

pruinescens Thomsen, 1935: 290. USA (New York). 

edwardsi de Meijere, 1946: 10 (preoccupied by Palpomyia edwardsi Cockerell, 1921). Netherlands. 

stagnalis Clastrier, 1962b: 274. France. 

linsleyi Wirth, 1952a: 224. USA (California). 

longa Remm, 1976a: 189. Georgia. 

longtana Yu, in Yu et al., 2005: 1593. China. 

lundstroemi Remm, 1981: 31. New name for bispinosa. 

bispinosa Lundström, 1916a: 15 (preoccupied by Palpomyia bispinosa Kieffer, 1915c). Finland. 

luteifemorata Edwards, 1926a: 419. Great Britain. 

luteiventris Santos Abreu, 1918: 326 (as variety of flavipes). Canary Islands (Spain). 

lutzi Lane, 1947b: 440. Brazil. 

maai Tokunaga, 1966a: 132. Papua New Guinea. 

macracantha Clastrier, 1962b: 252. France. 

magali de Meillon and Wirth, 1981b: 557. South Africa. 

magna Tokunaga, 1966a: 130. Papua New Guinea. 

magnispinosa Clastrier, 1962b: 250. France. 

mahyoubi Boorman and Harten, 2002: 458. Yemen. 

manilensis Edwards, 1929b: 10. Philippines. 

mapuche Spinelli, Grogan and Ronderos, in Spinelli, et al., 2009:50. Argentina. 

marinoi Spinelli, Grogan and Ronderos, in Spinelli, et al., 2009:53. Argentina. 

melacheira Remm, 1976a: 186. Russia. 

mellichroa Macfie, 1939c: 212. Brazil. 

micans Kieffer, 1918a: 61. Tunisia. 

microchela Kieffer, 1917b: 321. Colombia. 

montana Tokunaga, 1939a: 304. Japan. 

monticola Tokunaga, 1939a: 306. Japan. 

morenae (Strobl), 1900: 170 (Ceratopogon). Spain. 

multisaeta Yu and Xu, in Yu et al., 2005: 1594. China. 

multispinosa (Pierce), 1966: 97 (Neopalpomyia). USA (California). Miocene. 

nakali Boorman and Harten, 2002: 459. Oman. 

namwambae Macfie, 1939a: 99. Uganda. 

nana Zilahi-Sebess, 1934: 157. Bulgaria. 

nelsoni Macfie, 1932c: 50. New Zealand. 

nigricans Macfie, 1934c: 288. Malaysia. 

nigridorsum Kieffer, 1919a: 111. Slovak Republic. 

nigrina Clastrier, 1962b: 226. France. 

nigripalpis Goetghebuer, 1934a: 91. Spain. 

nigripecta Kieffer, 1919a: 108. Slovak Republic. 

nigripes (Meigen), 1830: 265 (Ceratopogon). Europe. 

pratensis (Meigen), 1830: 264 (Ceratopogon). Europe. 

bryocrypta Kieffer, 1925a: 425. France. 

nigritarsalis Remm, 1976a: 185. Kyrgyzstan. 

nigrithorax de Meillon, 1929: 248. South Africa. 

166

nigroflava Lane, 1947b: 446. Brazil. 

nigroscutellata Lane, 1947b: 444. Brazil. 

nitela Yu and Ding, in Yu et al., 2005: 1596. China. 

nitidissima Goetghebuer, 1948b: 15. Democratic Republic of the Congo. 

novaeguineae Tokunaga, 1966a: 135. Papua New Guinea. 

novaeirelandensis Tokunaga, 1966a: 135. Papua New Guinea. 

novitibialis Grogan and Wirth, 1975a: 13. USA (New York). 

obscurella Clastrier, 1962b: 256. France. 

occidentalis Grogan and Wirth, 1979a: 115. USA (Utah). 

oliffi de Meillon and Hardy, 1954: 72. South Africa. 

oliveirai Lane, 1947b: 443. Brazil. 

pacifica Remm, 1981: 31. New name for aterrima. 

aterrima Kieffer, 1922b: 159 (preoccupied by Palpomyia aterrima Goetghebuer, 1921). Taiwan. 

pallida (Tokunaga), 1966a: 107 (Heteromyia). Papua New Guinea. 

pallidipeda Yu and Liu, in Yu et al., 2005: 1597. China. 

pampana Lane, 1960: 386. Brazil. 

panacantha Clastrier, 1962b: 270. France. 

paraensis Lane, 1960: 387. Brazil. 


paulistensis Lane, 1947b: 445. Brazil. 

pendleburyi Edwards, 1933c: 254. Malaysia. 

pilea Yu, in Yu et al., 2005: 1598. China. 

plebeiella Grogan and Wirth, 1975a: 20. USA (New York). 

plebeja (Loew), 1861: 313 (Ceratopogon). USA (Pennsylvania). 

polyacantha Clastrier, 1962b: 268. France. 

portali Macfie, 1939a: 95. Uganda. 

praeusta (Loew), 1869b: 50 (Ceratopogon). Germany. 

apicalis (Strobl), 1898: 612 (Ceratopogon, as variety of flavipes, preoccupied by Ceratopogon apicalis Roser, 1840). 

Bosnia-Herzegovina. 

terminalis Kieffer, 1919a: 103. Slovak Republic. 

schineri Goetghebuer, 1920: 82 (as variety of flavipes, preoccupied by Nilobezzia schineri (Kieffer), 1919a). Belgium. 

pruinulosa Remm, 1976a: 178. Georgia. 

pseudolacustris Dippolito and Spinelli, in Dippolito et al., 1995: 55. Brazil. 

pseudorivularis Das Gupta, in Das Gupta, et al. 2009: 47. India. 

pseudorufa Grogan and Wirth, 1975a: 26. USA (Maryland). 

puberula Remm, 1976a: 196. Estonia. 

pubescens Kieffer, 1919a: 105. Slovak Republic. 

crassipes Goetghebuer, 1920: 112. Belgium. 

turfacea Kieffer, 1925b: 155. Russia. 

quadrispinosa Goetghebuer, 1920: 83. Belgium. 

quadristriata (Strobl), 1910: 263 (Ceratopogon). Austria. 

raignieri Goetghebuer, 1931: 211. Belgium. 

rastellifera Macfie, 1932c: 51. New Zealand. 

remmi Havelka, 1974: 622. Germany. 

reversa Remm, 1972: 83. Russia. 

riedeli Szadziewski, 1988: 183. Poland. Eocene. 

riouxi Huttel and Huttel, 1951: 99. France. 

rivularis Kieffer, 1911b: 328. India. 

rossica Remm, 1965: 185. Russia. 

rubiginosa Grogan and Wirth, 1975a: 22. USA (New York). 

rufa (Loew), 1861: 314 (Ceratopogon). USA (Pennsylvania). 

ruficrus Goetghebuer, 1948b: 15. Democratic Republic of the Congo. 

rufipes (Meigen), 1818: 81 (Ceratopogon). Europe. 

spinosior Kieffer, 1919a: 106. Slovak Republic, Hungary, Croatia. 

167

pseudospinipes Goetghebuer, 1933b: 112. Russia. 

plurispinosa Clastrier, 1962b: 265. France. 

rutshuruensis Goetghebuer, 1948b: 15. Democratic Republic of the Congo. 

ruwenzorii Macfie, 1939a: 98. Uganda. 

ryshkoffi (Pierce), 1966: 98 (Parapalpomyia). USA (California). Miocene. 

scabra (Coquillett), 1905: 62 (Ceratopogon). Mexico. 

scalpellifera Grogan and Wirth, 1975a: 33. USA (New York). 

schmidti Goetghebuer, 1934e: 36. Iraq. 

miki Goetghebuer, 1934a: 91. Hungary. 

semiermis Goetghebuer, 1914: 190. Belgium. 

goetghebueri Kieffer, 1919a: 98. Belgium. 

seneveti Clastrier, 1962b: 239. Algeria. 

septentrionalis Spinelli, Grogan and Ronderos, in Spinelli, et al., 2009:58. Argentina. 

serraticauda Tokunaga, 1966a: 128. Papua New Guinea. 

serripes (Meigen), 1818: 82 (Ceratopogon). Europe. 

transfuga (Staeger), 1839: 598 (Ceratopogon). Denmark. 

tarsata (Zetterstedt), 1855: 4874 (Ceratopogon). Sweden. 

ruficeps Kieffer, 1918a: 59. Tunisia. 

shilo (Pierce), 1966: 95 (Miopalpomyia). USA (California). Miocene. 

simplitheca Das Gupta, in Das Gupta, et al. 2009: 47. India. 

sinanoensis Tokunaga, 1939a: 300. Japan. 

singularis de Meillon, 1937b: 362. South Africa. 

spinifemur (Lane), 1948: 234 (Dicrobezzia). Brazil. 

spinipes (Meigen) in Panzer, 1806: plate 14 (Ceratopogon). Germany. 

fulva (Macquart), 1826: 181 (Ceratopogon). France. 

ferruginea (Meigen), 1830: 265 (Ceratopogon). Denmark. 

parviforceps Kieffer, 1925d: 98. France. 

spinosa Lutz, 1914: 93. Brazil. 

spinulosa Goetghebuer, 1935d: 180. Democratic Republic of the Congo. 

stella Tokunaga, 1966a: 136. Indonesia. 

stonei Wirth, 1951d: 322. USA (Virginia). 

subalpina Lee, 1948d: 59. Australia (Capital Territory). 

subaspera (Coquillett), 1901a: 606 (Ceratopogon). USA (New Mexico). 

maculicrus Ingram and Macfie, 1931a: 230. Argentina. 

essigi Wirth, 1952a: 225. USA (California). 

subfuscula Ingram and Macfie, 1931a: 216. Chile. 

succincta (Meigen), 1818: 85 (Ceratopogon). Portugal. 

succinea Szadziewski, 1988: 185. Denmark. Eocene. 

tainana Kieffer, 1912a: 29. Taiwan. 

tamaricis Remm, 1976a: 176. Kyrgyzstan. 

tamioi Lane, 1960: 388. Brazil. 

tanycornis Borkent, in Borkent and Wirth, 1997:134. New name for longicornis. 

longicornis (Williston), 1896: 280 (Ceratopogon, preoccupied by Ceratopogon longicornis Waltl, 1837 ). St. Vincent. 

tasmanica (Lee), 1948d: 62 (Heteromyia). Australia (Tasmania). 

tauffliebi Vattier and Adam, 1966b: 762. Congo. 

tenuicrus Kieffer, 1917b: 320. Paraguay. 

terrea (Meigen), 1818: 85 (Ceratopogon). Germany. 

oldenbergi Goetghebuer, 1932a: 127. Germany. 

thomsoni Macfie, 1939a: 96. Uganda. 

tibialis (Meigen), 1818: 82 (Ceratopogon). Austria. 

atripecta Kieffer, 1919a: 101. Serbia-Montenegro. 

laticollis Goetghebuer, 1922: 55. Belgium. 

nipponica Tokunaga, 1939a: 297. Japan. 

tinctipennis Kieffer, 1919a: 110. Romania, Slovak Republic. 

168

obscuripennis Kieffer, 1919a: 110. Croatia. 

semifumosa Goetghebuer, 1922: 58. Belgium. 

adusta Kieffer, 1925a: 426. Germany. 

nigritella Clastrier, 1962b: 255. France. 

trifasciata Wirth, 1952a: 222. USA (California). 

turanica Remm, 1976a: 176. Kazakhstan. 

tuvae Remm, 1972: 84. Russia. 

umbella Macfie, 1939c: 208. Brazil. 

urpicifemoris Macfie, 1932c: 52. New Zealand. 

vallouisensis Clastrier, 1962b: 272. France. 

variipila Kieffer, 1924b: 407. Germany. 

venetiae Harant, Huttel and Huttel, 1952: 14. Italy. 

verna de Meillon and Wirth, 1987a: 70. South Africa. 

versicolor Macfie, 1939c: 215. Brazil. 

vittata Clastrier, Rioux and Descous, 1961: 78. Chad. 

walteri Grogan and Wirth, 1979a: 62. USA (West Virginia). 

weemsi Grogan and Wirth, 1979a: 35. USA (Florida). 

wirthi Lane, Forattini and Rabello, 1955: 82. Brazil. 

wirthorum Szadziewski and Grogan, 1997: 257. Dominican Republic. Miocene. 

wittei Goetghebuer, 1948b: 14. Democratic Republic of the Congo. 

xanthothorax Remm, 1976a: 180. Tajikistan. 

xizanga Yu and Liu, in Yu et al., 2005: 1605. China. 

yamana Spinelli, Grogan and Ronderos, in Spinelli, et al., 2009:63. Argentina. 

yasumatsui Tokunaga, 1940d: 158. Japan. 

yuanqingi Yu, in Yu et al., 2005: 1607. China. 

zernyi Goetghebuer, 1934a: 92. Spain. 

ziliangi Yu, in Yu et al., 2005: 1604. China. 

zyzza Yu and Zou, in Yu et al., 2005: 1608. China. 

Genus PHAENOBEZZIA Haeselbarth 

PHAENOBEZZIA Haeselbarth, 1965a: 297. Type species: Probezzia pistiae Ingram and Macfie, by original designation. 

assimilis (Johannsen), 1932: 445 (Bezzia). Indonesia. 

ateles (Macfie), 1940a: 78 (Bezzia). Brazil. 

astyla Spinelli and Wirth, 1986: 232. Colombia. 

beni de Meillon and Wirth, 1983a: 379. South Africa. 

chonganensis Yu and Shen, 1998: 62. China. 

cinnae (de Meillon), 1936: 185 (Palpomyia). South Africa. 

conspersa (Johannsen), 1932: 446 (Bezzia). Indonesia. 

eucera (Kieffer), 1911a: 123 (Bezzia). India. 

excisa (Clastrier), 1962a: 110 (Bezzia). Algeria. 

fulvithorax (Malloch), 1915a: 354 (Probezzia). USA (Illinois). 

griseipennis (Clastrier), 1958c: 498 (Nilobezzia). Senegal. 

javana (Kieffer), 1923b: 142 (Probezzia). Indonesia. 

lineola (Kieffer), 1910: 205 (Bezzia, as variety of gracilipes). India. 

gracilipes (Kieffer), 1910: 205 (Bezzia, preoccupied by Bezzia gracilipes (Winnertz), 1852). India. 

mashonensis (Ingram and Macfie), 1923: 73 (Bezzia). Zimbabwe. 

maya Spinelli and Wirth, 1986: 234. Belize. 

mellipes Wirth and Ratanaworabhan, 1981b: 427. Thailand. 

minutistyla (Tokunaga), 1939a: 293 (Bezzia). Japan. 

nitens Liu, Yan and Liu, 1996: 48. China. 

nzuari (Vattier and Adam), 1966b: 750 (Probezzia). Congo. 

troglophila (Vattier and Adam), 1966b: 754 (Nilobezzia). Congo. 

169

opaca (Loew), 1861: 312 (Ceratopogon). USA (District of Columbia). 

incerta (Malloch), 1915a: 358 (Probezzia). USA (Illinois). 

pajoti (Vattier and Adam), 1966b: 752 (Probezzia). Congo. 

congolensis (Vattier and Adam), 1966b: 758 (Nilobezzia). Congo. 

pistiae (Ingram and Macfie), 1921: 364 (Probezzia). Ghana. 

probata (de Meillon), 1937b: 349 (Bezzia). South Africa. 

rara Saha and Das Gupta, 2001: 250. India. 

rubiginosa (Winnertz), 1852: 72 (Ceratopogon). Germany. 

glyceriae (Kieffer), 1913a: 9 (Bezzia). Germany. 

nitidiventris (Goetghebuer), 1923: 104 (Probezzia). Belgium. 

radialis (Goetghebuer), 1936: 323 (Bezzia). Belgium. 

sabroskyi Wirth and Grogan, 1982: 189. USA (Maryland). 

seychelleana (Kieffer), 1911c: 350 (Probezzia). Seychelles. 

spekei (Macfie), 1939a: 101 (Nilobezzia). Uganda. 

stuckenbergi Haeselbarth, 1965a: 308. South Africa. 

suavis Saha and Das Gupta, 2001: 251. India. 

tenuiforceps (Clastrier), 1962a: 109 (Bezzia). Algeria. 

tropica (Clastrier and Wirth), 1961a: 212 (Nilobezzia). Gambia. 

vacunae (de Meillon), 1936: 180 (Bezzia). South Africa. 

vulgaris Saha and Das Gupta, 2001: 251. India. 

wirthi Szadziewski and Grogan, 1997: 259. Dominican Republic. Miocene. 

zuluensis Haeselbarth, 1965a: 306. South Africa. 

TRIBE STENOXENINI Coquillett, 1899: 61 

Genus PARYPHOCONUS Enderlein 

PARYPHOCONUS Enderlein, 1912: 57. Type species: Paryphoconus angustipennis Enderlein, by original designation. 

aemulus Macfie, 1940b: 180. Guyana. 

amapaensis Lane, 1961b: 450. Brazil. 

angustipennis Enderlein, 1912: 57. Brazil. 

anomalicornis Kieffer, 1917b: 333. Colombia and Paraguay. 

apicalis Spinelli and Wirth, 1984b: 889. Ecuador. 

barrettoi Lane, 1946a: 203. Brazil. 

batesi Lane, 1961c: 455. Brazil. 

brunneipennis Spinelli and Wirth, 1984b: 890. Brazil. 

ecuadorensis Spinelli and Wirth, 1984b: 892. Ecuador. 

enderleini Lane, 1956a: 301. Brazil. 

fittkaui Spinelli and Wirth, 1984b: 892. Brazil. 

flavidus (Johannsen), 1943: 761 (Ceratobezzia). Guyana. 

lanei Wirth, 1959b: 236. Brazil. 

flinti Spinelli and Wirth, 1984b: 894. Mexico. 

fusciradialis Spinelli and Wirth, 1984b: 895. Brazil. 

fuscus Lane, 1946a: 206. Brazil. 

goianensis Lane, 1961c: 456. Brazil. 

grandis Macfie, 1939d: 6. Brazil. 

guianae Macfie, 1940b: 180. Guyana. 


latipennis Spinelli and Wirth, 1984b: 898. Colombia. 

leei Spinelli and Wirth, 1984b: 898. Colombia. 

macfiei Lane, 1946a: 203. Brazil. 

maya Spinelli and Wirth, 1984b: 899. Belize. 

mayeri Wirth, 1959b: 236. Brazil. 

170

misionensis Spinelli, 1998: 52. Argentina. 

neotropicalis (Lane), 1948: 229 (Macropeza). Brazil. 

nigripes Macfie, 1939d: 8. Argentina. 

nubifer Macfie, 1939d: 5. Brazil. 

oliveirai Lane, 1956a: 303. Brazil. 

paranaensis Spinelli and Wirth, 1984b: 902. Argentina. 

paulistensis Lane, 1961c: 457. Brazil. 

peruvianus Mauad and Spinelli, 2011: 487. Peru. 

sonorensis Wirth and Ratanaworabhan, 1972c: 1374. Mexico. 

steineri Spinelli and Wirth, 1984b: 905. Peru. 

subflavus Macfie, 1940d: 23. Guyana. 

travassosi Lane, 1956a: 304. Brazil. 

taragui Spinelli, 1998: 53. Argentina. 

telmatophilus (Macfie), 1940a: 77 (Bezzia). Brazil. 

terminalis (Coquillett), 1904a: 90 (Ceratopogon). Nicaragua. 

unimaculatus Macfie, 1940b: 179. Guyana. 

wirthi Lane, 1961c: 458. Brazil. 

wygodzinskyi Lane, 1946a: 208. Brazil. 

Genus STENOXENUS Coquillett 

STENOXENUS Coquillett, 1899: 61. Type species: Stenoxenus johnsoni Coquillett, by monotypy. 

aductus Dippolito and Spinelli, in Dippolito et al., 1995: 56. Brazil. 

arcuatus Wirth and Ratanaworabhan, 1972c: 1379. Panama. 

blantoni Wirth and Ratanaworabhan, 1972c: 1379. Panama. 


carrerai Lane, 1956a: 305 (as carreirai). Brazil. 

coomani Séguy, 1931a: 210. Viet Nam. 

insigninervis Macfie, 1934c: 290. Malaysia. 

dimorphus Kieffer, 1909a: 47. Peru. 

excentricus Lane, 1961c: 453. Brazil. 

fulvus Johannsen, 1927a: 70. Peru. 

johnsoni Coquillett, 1899: 61. USA (New Jersey). 

lanei Wirth and Ratanaworabhan, 1972c: 1381. Panama. 

limpidus Wirth and Ratanaworabhan, 1972c: 1383. Costa Rica. 

marginalis Wirth and Ratanaworabhan, 1972c: 1385. Panama. 

niger Lane, 1948: 230. Brazil. 

paraensis Lane, 1956a: 306. Brazil. 

pastorianus Clastrier, 1982b: 293. Guinea. 

pauliani Vattier and Adam, 1966b: 767. Congo. 

pseudofulvus Spinelli, 1998: 54. Argentina. 

setiger Macfie, 1939c: 138. Brazil. 

stenopterus Wirth and Ratanaworabhan, 1972c: 1387. Philippines. 

usutae de Meillon, 1959a: 355. South Africa. 

171

Taxa Unplaced in Ceratopogonidae 

Genus ALAUTUNMYIA Borkent 

ALAUTUNMYIA Borkent, 1996: 11. Type species: Alautunmyia elongata Borkent, by original designation. 

elongata Borkent, 1996: 13. USA (New Jersey). Upper Cretaceous. 

Genus ARCHICULICOIDES Szadziewski 

ARCHICULICOIDES Szadziewski, 1996: 29. Type species: Archiculicoides schleei Szadziewski, by original designation. 

acraorum (Borkent), 2000a: 367 (Protoculicoides). Lebanon. Lower Cretaceous. 

schleei Szadziewski, 1996: 29. Lebanon. Lower Cretaceous. 

unus (Borkent), 2000a: 367 (Protoculicoides). Lebanon. Lower Cretaceous. 

Genus PROTOCULICOIDES Boesel 

PROTOCULICOIDES Boesel, 1937: 50. Type species: Protoculicoides depressus Boesel, by original designation. 

burmiticus Szadziewski and Poinar, 2005: 352. Myanmar. Lower Cretaceous. 

ciliates Borkent, 2012: 762. Canada (Alberta). Upper Cretaceous. 

depressus Boesel, 1937: 51. Canada (Manitoba). Upper Cretaceous. 

punctus Borkent, 2000a: 368. Lebanon. Lower Cretaceous. 

skalskii Szadziewski and Arillo, 1998: 292. Spain. Lower Cretaceous. 

succineus Szadziewski, 1996: 31. Lebanon. Lower Cretaceous. 

Genus SINOPOGONITES Hong 

SINOPOGONITES Hong, 2002a: 167 [2002b: 109]. Type species: Sinopogonites eocenicus Hong, by original designation. 

eocenicus Hong, 2002a: 168 [2002b: 109]. China. Eocene. 

Unplaced Genus of Forcipomyiinae 

DIDYMOPHLEPS Weyenbergh, 1883: 108. Type species: Didymophleps hortorum Weyenbergh, by monotypy. 

Unplaced Genera of Ceratopogoninae 

NEOCULICOIDES Pierce, 1966: 93. Type species: Neoculicoides jeanneae Pierce, by original designation. 

PARACULICOIDES Pierce, 1966: 94. Type species: Paraculicoides rouseae Pierce, by original designation. 

Unplaced Genus of Heteromyiini + Sphaeromiini + Palpomyiini + Stenoxenini 

EOPALPOMYITIS Hong, Guo, and Ren, 2000: 226 (Hong, 2002: 165). Type species: Palpomyia unca Hong, by original 

designation. 

Unplaced Species of Forcipomyiinae 

Ceratopogon albipennis Meigen, 1818: 73. Germany. 

Ceratopogon anomalus Macquart, 1826: 180. France. 

Ceratopogon cantabricus Strobl, 1900: 169. Spain. 

Ceratopogon annulatus Meigen, in Waltl, 1835: 67. Germany. 

Ceratopogon cypri Kieffer, 1918a: 86. Cyprus. 

172

Ceratopogon koniae Kieffer, 1918a: 87. Turkey. 

Ceratopogon leucopterus Meigen, 1804: 28. Germany. 

Ceratopogon mediterraneus Kieffer, 1919a: 16. Greece. 

Ceratopogon pallipes Meigen, 1818: 74. Germany. 

Ceratopogon tristis Meigen, 1830: 263. Germany. 

Didymophleps hortorum Weyenbergh, 1883: 110. Argentina. 

Helea erronea Speiser, 1910: 741. France. 

Unplaced Species of Ceratopogoninae 

Ceratopogon escheri Giebel, 1856: 252. Baltic region. Eocene. 

Ceratopogon kamus de Meillon, 1959a: 340. South Africa. 

Johannsenomyia hotchkissae Pierce, 1966: 94. USA (California). Miocene. 

Monohelea nigrita Goetghebuer, 1941: 288. Germany. 

Neoculicoides jeanneae Pierce, 1966: 94. USA (California). Miocene. 

Paraculicoides rouseae Pierce, 1966: 94. USA (California). Miocene. 

Unplaced species of Heteromyiini + Sphaeromiini + Palpomyiini + Stenoxenini 

Ceratopogon apicalis Roser, 1840: 50. Europe. 

Tipula goezii Schrank, 1803: 72. Germany. 

Ceratopogon quadrimaculatus Strobl, 1906: 399. Spain. 

Ceratopogon ruficornis Macquart, 1826: 178. France. 

Ceratopogon succinotus Meigen, 1818: 85. Portugal. 

Eopalpomyitis unca (Hong), 1981: 58 (Palpomyia). China. Eocene. 

Unplaced Species of Ceratopogonidae 

Cecidomyia pennicornis Zetterstedt, 1850: 3696. Sweden. 

Ceratopogon claripennis Lynch Arribálzaga, 1893: 229. Argentina. 

Ceratopogon crassipalpis Meunier, 1912: 365. Madagascar. Holocene? 

Ceratopogon longicornis Waltl, 1837: 279. Germany. 

Ceratopogon varipes Meigen, 1838: 18. Germany. 

Ceratopogon madagascariensis Meunier, 1912: 365. Madagascar. Holocene? 

Ceratopogon scutellatus Say, 1829: 150. USA (Indiana). 

Ceratopogon tenuipes Meunier, 1912: 366. Madagascar. Holocene? 

Ceratopogon barbipes Gimmerthal, 1846: 102. Latvia. 

Ceratopogon nemorosus Meigen, 1804: 24. Germany. 

Ceratopogon pusillus Holmgren, in Holmgren and Aurivillius, 1883: 182. Russia. 

Ceratopogon univittatus Zetterstedt, 1838: 820. Sweden. 

Ceratopogon viscatus Meunier, 1912: 364. Madagascar. Holocene? 

Ceratopogon vittatus Wiedemann, 1817: 66. Germany. 

Tipula subsultans Schrank, 1803: 77. Germany. 

Species in Other Families Originally Considered to Be Ceratopogonidae 

Ceratopogon blanchardi Iches, 1906: 266. Undetermined Chironomidae. 

Ceratopogon humeralis Zetterstedt, 1838: 820. Junior synonym of Cricotopus ephippium (Zetterstedt). Chironomidae. 

Ceratopogon imperfectus Skuse, 1889: 307. A valid species of Pseudosmittia Goetghebuer. Chironomidae. 

Ceratopogon pusio Zetterstedt, 1850: 3647. Undetermined Chironomidae. 

Ceratopogon validinervis Zetterstedt, 1850: 3645. Junior synonym of Mycetobia pallipes (Meigen). Anisopodidae. 

173

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176

Brunetti, E. 1912. New Oriental Diptera. Records of the Indian Museum 7:445-513. 

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Bullock, H.R. and J. Akiyama 1959. A new biting midge from Japan and Korea (Diptera, Heleidae). Japanese Journal of 

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Buyanova, O.F. 1962. Description of a new species of blood-sucking midges 

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Byers, G.W., F. Blank, W.J. Hanson, D.F. Beneway and R.W. Fredrichson. 1962. Catalogue of the types in the Snow 

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Bystrak, P.G. and D.H. Messersmith. 1980. A new species of midge of the genus Forcipomyia Meigen (Diptera: 

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Caeiro, V.M.P. 1959. Culicoides gulbenkiani, a new species of Culicoides (Diptera Ceratopogonidae) in South Africa. 

Onderstepoort Journal of Veterinary Research 28:155-167. 

– 1961. Contribuicao para o estudo das espécies Angolanas do género Culicoides Latreille, 1809 (Diptera, 

Ceratopogonidae). Junta de Investigacoes Cientificas do Ultramar, Estudos, Ensaios, e Documentos 86:17-359, 58 

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Callot, J. and M. Kremer. 1961a. Culicoides riouxi et Culicoides pseudoheliophilus, especes nouvelles du groupe des 

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688. 

– 1961b. Scission de l'espèce Culicoides truncorum Edwards (Diptera: Ceratopogonidae) en deux espèces. Bulletin de la 

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– 1965. Sur quelques Diptères Nématocères du Var, avec la description d'espèces nouvelles de Culicoides 

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– 1966. Culicoides heteroclitus n. sp. (Diptères, Cératopogonidés). Proceedings of the 1st International Congress of 

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– 1969a. Description de Culicoides dobyi n. sp. (Diptera Ceratopogonidae). Bulletin de la Société de Pathologie Exotique 

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– 1969b. Description d'un Culicoide nouveau C. jumineri (Dipt. Cératopogonidé) trouvé en Tunisie. Ibid.:1112-1118. 

Callot, J., M. Kremer and H. Bailly-Choumara. 1970. Description de Culicoides coluzzii n. sp. (Dipt., Ceratopogonidae). 

Bulletin de la Société Zoologique de France 95:709-718. 

Callot, J., M. Kremer and M. Basset. 1968. Culicoides marcleti n. sp. et nouvelles localisations de Culicoides (Diptères, 

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Callot, J., M. Kremer and Y. Deduit. 1962. Nouvelles especes et nouvelles localisations de Culicoides (Diptera: 

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Callot, J., M. Kremer and B. Molet. 1967. Cératopogonidés (Diptères) de la région éthiopienne et particulièrement d'Angola 

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Annales de Parasitologie Humaine et Comparée 48:377-386. 

Callot, J., M. Kremer, B. Molet and A. Bach. 1968. Nouvelles espèces, nouvelles localisations de Culicoides (Dipt. Ceratop.) 

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Callot, J., M. Kremer, J. Mouchet and A. Bach. 1965. Contribution à l'étude de Cératopogonidés (Diptera) de Kumba 

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Callot, J., M. Kremer and C. Paradis. 1962. Culicoides furcillatus n. sp. et Culicoides setosus Gutzevich, Diptères 

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Callot, J., M. Kremer, B. Rault and A. Bach. 1966. Contribution à l'étude des Culicoides de l'Ouest de la France. Annales de 


Callot, J., M. Kremer and J.-A. Rioux. 1963. Sur des Culicoides (Diptera: Ceratopogonidae) dont une espèce et une variété 

nouvelles du Midi de la France. Annales de Parasitologie Humaine et Comparée 38:121-129. 

Callot, J., M. Kremer, J.-A. Rioux and S. Descous. 1967. Culicoides des Pyrénées Orientales. Description de C. 

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177

Cambournac, F.J.C. 1956. Culicoides (Nematocera, Ceratopogonidae): sua ocorrencia em Portugal - Descricao de C. 

sintrensis sp. n. Anais do Instituto de Medicina Tropical (Lisbon) 13:589-595. 

– 1970a. Lista das espécies do género Culicoides (Nematocera, Ceratopogonidae) encontrada em Portugal. Anais da 

Escola Nacional de Saude Publica e de Medicina Tropical 4:249-250. 

– 1970b. Culicoides almeidae (Nematocera, Ceratopogonidae) sp. n.; uma nova espécie encontrada em Portugal. Anais da 

Escola Nacional de Saude Publica e de Medicina Tropical 4:251-257. 

Campbell, J.A. and E.C. Pelham-Clinton. 1960. A taxonomic review of the British species Culicoides Latreille (Diptera, 

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summary]. Acta Zootaxonomica Sinica 9:298-300. 

Carter, H.F. 1916. On three new African midges. Annals of Tropical Medicine and Parasitology 10:131-138, pl. 3. 

– 1919. New West African Ceratopogoninae. Ibid. 12:289-302, pl. 8. 

– 1921. A revision of the genus Leptoconops, Skuse. Bulletin of Entomological Research 12:1-28. (June). 

Carter, H.F., A. Ingram and J.W.S. Macfie. 1920. Observations on the ceratopogonine midges of the Gold Coast with 

descriptions of new species Part II. Annals of Tropical Medicine and Parasitology 14:211-274, pls. 7-8. 

– 1921a. Observations on the ceratopogonine midges of the Gold Coast with descriptions of new species Part III. Ibid.:309- 

331. 

– 1921b. Observations on the ceratopogonine midges of the Gold Coast with descriptions of new species Part IV. Ibid. 

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Causey, O.R. 1938. Culicoides of Siam with descriptions of new species. American Journal of Hygiene 27:399-416, 8 pls. 

Cavalieri, F. 1961a. Notas sobre Ceratopogonidae (Diptera, Nematocera) I. Forcipomyia (Forcipomyia) wygodzinskyi sp. n. 

para Tierra del Fuego. Revista Sociedad de la Entomologica Argentina 23:17-19. 

– 1961b. Notas sobre Ceratopogonidae (Diptera, Nematocera) II. Sobre dos nuevas especies de Forcipomyia para Tierra 

del Fuego, Forcipomyia (F.) delpontei n. sp. y F. (F.) piroskyi. Acta Zoologica Lilloana 18:169-175 (1962). 

– 1962. Notas sobre Ceratopogonidae (Dipt. Nematocera) III. Sobre un nuevo diptero hematofago para Argentina, 

Lasiohelea saltensis n. sp. y notas sobre las especies neotropicales de Lasiohelea. Ibid.:359-365. 

– 1966. Notas sobre Ceratopogonidae (Diptera, Nematocera) V. Descripcion de una nueva especie de Venezuela: 

Culicoides (Oecacta) birabeni sp. n. Physis 26:59-63. 

Cavalieri, F. and I.F. Chiossone. 1966. Datos sobre la familia de dipteros hematofagos Leptoconopidae en la Argentina, con 

la descripcion de Leptoconops (Leptoconops) casali sp. n. y la redescripcion de L. (L.) petrocchiae Shannon y Del 

Ponte, 1927. Physis 26:43-57. 

– 1972. Notas sobre el género Atrichopogon Kieffer, 1906, de la Argentina (Diptera, Ceratopogonidae). I. Consideraciones 

generales y descripcion de Atrichpogon delpontei sp. nov. Revista de la Sociedad Entomologica Argentina 34:119- 

129. 

– 1973. Notas sobre el género Atrichopogon Kieffer, 1906, de la Argentina (Diptera, Ceratopogonidae). II. Descripcion de 

Atrichpogon casali sp. nov. Physis 32:151-159. 

Cazorla, C.G. and G.R. Spinelli. 2007. A new species of Patagonian Stilobezzia (Acanthohelea) and a redescription of S. (A.) 

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Cazorla, C.C., G.R. Spinelli and F. Díaz. 2005. Two new species of the subgenus Stilobezzia (Stilobezzia) Kieffer from 

Peruvian Amazonia (Diptera: Ceratopogonidae). Amazoniana 18:289-297. 

Cazorla, C.G., M.M. Ronderos, G.R. Spinelli, S.R.S. Torreias and R. L. Ferreira Keppler. 2012. A new species of Stilobezzia 

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Chan, K.L. and E.J. LeRoux. 1965. Description of Forcipomyia (Neoforcipomyia) sp. n. and redescription of Forcipomyia 

(Neoforcipomyia) eques (Johannsen) (Diptera: Ceratopogonidae), with an account of the digestive and reproductive 

systems. Phytoprotection 46:74-104. 

– 1970. New species of Forcipomyia (Diptera: Ceratopogonidae) described in all stages. Canadian Entomologist 102:271-- 

293. 

– 1971a. A new subgenus and species of midge (Diptera: Ceratopogonidae) from Singapore. Ibid. 103:271-276. 

– 1971b. Nine new species of Forcipomyia (Diptera: Ceratopogonidae) described in all stages. Ibid.:729-762. 

178

Chan, K.L. and J.R. Linley. 1988. Description of Atrichopogon wirthi new species (Diptera: Ceratopogonidae) from leaves of 

the water lettuce (Pistia stratiotes) in Florida. Florida Entomologist 71:186-201. 

– 1989. A new Florida species of Forcipomyia (Euprojoannisia) (Diptera: Ceratopogonidae) from leaves of the water lettuce, 

Pistia stratiotes. Ibid. 72:252-262. 

Chan, K.L. and L.G. Saunders. 1965. Forcipomyia (Dacnoforcipomyia) anabaenae, a new blood-sucking midge from 

Singapore, described in all stages (Diptera: Ceratopogonidae). Canadian Journal of Zoology 43:527-540. 

Chanthawanich, N. and M.D. Delfinado. 1967. Some species of Leptoconops of the oriental and pacific regions (Diptera, 

Ceratopogonidae). Journal of Medical Entomology 4:294-303. 

Chaudhuri, P.K. and S.K. Das Gupta. 1991. Stilobezzia szadziewskii Chaudhuri & Das Gupta, a replacement name for 

Stilobezzia falcata Chaudhuri, Das Gupta & Sinharay (preoccupied) (Diptera: Ceratopogonidae). Oriental Insects 

25:182. 

Chaudhuri, P.K. and M. Ghosh. 1980. Addition to the studies of genus Alluaudomyia Kieffer (Diptera: Ceratopogonidae) of 

India. Oriental Insects 14:421-424. 

Chaudhuri, P.K., S.K. Das Gupta and A.K. Chatterjee. 1981. Alluaudomyia fuscitarsis n. sp. and Stilobezzia ensistyla n. sp. 

from India (Diptera: Ceratopogonidae). Entomologica Scandinavica 12:147-150. 

Chaudhuri, P.K., S.K. Das Gupta and D.K. Chaudhuri. 1972. Biting midges of the genus Alluaudomyia Kieffer (Diptera: 

Ceratopogonidae) from India. Oriental Insects 6:83-110. 

Chaudhuri, P.K., S.K. Das Gupta and D.C. Sinharay. 1974. Three new species of Stilobezzia Kieffer (Diptera: 

Ceratopogonidae) from India. Oriental Insects 8:47-50. 

Chen, C.-S. 1979. Descriptions of Culicoides lieni n. sp. and C. hainanensis Lee from Taiwan (Diptera: Ceratopogonidae). 

Chinese Journal of Microbiology 12:99-104. 

– 1983. The genus Culicoides of Taiwan (Diptera: Ceratopogonidae) [in Chinese]. Ph.D. thesis, Graduate School of Plant 

Pathology and Entomology, National Taiwan University, Taiwan, 226 pp., pls. 1-4. 

– 1988. Descriptions of four new species of Culicoides (Diptera: Ceratopogonidae) from Taiwan. Chinese Journal of 

Entomology 8:151-156. 

Chen, H.-Y., Y.-Q. Liu, and Y.-X. Yu. 2012. Two new biting midges of the subfamily Forcipomyiinae Lenz (Diptera: 

Ceratopogonidae) from China. Zootaxa 3582:33-36. 

Chen, J.-L., R.-D. Chen, Z.-H. Liang, Z.-M. He and Y.-X. Yu. 2007. Preliminary report of blood-sucking midges in Hong Kong 

(Diptera: Ceratopogonidae) [in Chinese, English summary]. Acta Parasitologica et Medica Entomologica Sinica 

14:169-173. 

Chen, K.C. 1981. A new species of Culicoides from Wuyi-Mountain (Diptera: Ceratopogonidae) [in Chinese, English 

summary]. Entomotaxonomia 3:93-94. 

– 1982. A preliminary survey of blood-sucking midges (Diptera: Ceratopogonidae) in Wuyi Shan Fujian (in Chinese, English 

summary). Wuyi Science Journal 2:128-131. 

Chen, K.C. and L.L. Tsai. 1962. The bloodsucking midges (Ceratopogonidae) of Fukien [in Chinese, English summary]. Acta 

Entomologica Sinica 11:394-400. 

Choudhuri, P.P., M. Bose, S.K. Das Gupta and P.K. Choudhuri. 1986. Anophelis group of Culicoides Insects 

(Ceratopogonidae: Diptera) of India. Burdwan University Science Journal 3:52-55. 

Chu, F.-I. 1977. New species and records of Culicoides (Diptera: Ceratopogonidae) from Tibet China [in Chinese, English 

summary]. Acta Entomologica Sinica 20:99-105. 

– 1981. On the blood-sucking midges (Diptera: Ceratopogonidae) from the coastal regions of south-eastern China [in 

Chinese, English summary]. Ibid. 24:307-313. 

– 1983. Two new subgenera and two new species of Culicoides from China (Diptera: Ceratopogonidae) [in Chinese, English 


– 1984. Two new names of Chinese Culicoides (Diptera: Ceratopogonidae) [in Chinese]. Ibid. 6:24. 

– 1986. The biting midges of Cambodia, with descriptions of two new species (Diptera: Ceratopogonidae) [in Chinese, 

English summary]. Ibid. 8:251-261. 

Chu, F.-I. and S.-C. Liu. 1978. A taxonomic study of the genus Culicoides (Diptera: Ceratopogonidae) of Yunan, with 

descriptions of five new species [in Chinese, English summary]. Acta Entomologica Sinica 21:79-90. 

Chu, F.-I., J.-Q. Qian and D.-X. Ma. 1982. Records of biting midges collected from Xinjiang, China with descriptions of two 

new species (Diptera: Ceratopogonidae) [in Chinese, English summary]. Acta Entomologica Sinica 25:105-110. 

Clastrier, J. 1956. Notes sur les Cératopogonidés. I.-Quatre espèces du groupe Forcipomyia d'Algérie et de Tunisie. 

Archives de l'Institut Pasteur Algérie 34:496-512. 

– 1957. Notes sur les Cératopogonidés. II.-Quelques Culicoides à ailes tachetées. Ibid. 35:404-444. 

179

– 1958a. Notes sur les Cératopogonidés. III.-Culicoides semimaculatus n. sp. d'Algérie. Ibid. 36:55-60. 

– 1958b. Notes sur les Cératopogonidés. IV.-Cératopogonidés d'Afrique Occidentale Francaise. Ibid.:192-258. 

– 1958c. Notes sur les Cératopogonidés. V.-Cératopogonidés d'Afrique Occidentale Francaise (2). Ibid.:487-505. 

– 1959a. Notes sur les Cératopogonidés. VI.-Cératopogonidés d'Afrique Occidentale Francaise (3). Ibid. 37:167-197. 

– 1959b. Notes sur les Cératopogonidés. VII.-Cératopogonidés d'Afrique Occidentale Francaise (4). Ibid.:340-383. 

– 1959c. Notes sur les Cératopogonidés. VIII.-Cératopogonidés de L'Ile de la Réunion. Ibid.:412-446. (Sept.). 

– 1960a. Notes sur les Cératopogonidés. IX.-Cératopogonidés de la République du Congo. Ibid. 38:79-105. 

– 1960b. Notes sur les Cératopogonidés. X.-Cératopogonidés de la République du Congo (2). Ibid.:258-298. 

– 1960c. Notes sur les Cératopogonidés. XI.-Cératopogonidés de la République du Congo (3). Ibid.:510-526. 

– 1961a. Notes sur les Cératopogonidés. XV.-Ceratopogon et Alluaudomyia de la Région Paléarctique. Ibid. 39:401-437. 

– 1961b. Le Parc National du Niokolo-Koba (Deuxième fascicule). XVII. (Diptera Ceratopogonidae). Mémoires de l'Institut 

Francais d'Afrique Noire 62:257-272. 

– 1962a. Notes sur les Cératopogonidés. XVI.-Espèces du genre Bezzia Kieffer ou apparentées de la région Paléarctique. 


– 1962b. Notes sur les Cératopogonidés de la région Paléarctique. XVII. Nouveaux Palpomyia Meig. et Johannsenomyia 

Mall. Ibid.:225-288. 

– 1963. Notes sur les Cératopogonidés. XVIII.-Espèces du genre Stilobezzia Kieffer ou Apparentées de la Région 

Paléarctique. Ibid. 41:41-68. 

– 1966. Cératopogonidés des Iles Canaries (Dipt. Nematocera). Annales de la Société de Entomologique de France 2:693- 

710. 

– 1967. Note sur deux Cératopogonidés du Cambodge: Styloconops spinosifrons (Carter 1921) et Stilobezzia chasteli n. sp. 

Bulletin de la Société Entomologique de France 72:115-120. 

– 1968. Deux Ceratopogonides nouveaux de la Guyane Francaise. Archives de l'Institut Pasteur de la Guyane Francaise et 

de L'Inini 21:85-92. 

– 1971. Deux nouveaux Culicoides (Diptera, Ceratopogonidae) de la Guyane Francaise. Annales de Parasitologie 46:285- 

294. 

– 1972a. Contribution à la faune du Congo (Brazzaville). Mission A. Descarpentries et A. Villiers. Bulletin de l'Institut 

Fondamental d'Afrique Noire 34:98-103. 

– 1972b. Forcipomyia (F.) squamithorax n. sp. de la Guyane francaise. Bulletin de la Société Entomologique de France 

77:170-176. 

– 1973. Le genre Leptoconops, sous-genre Holoconops dans le midi de la France (Dipt. Ceratopogonidae). Annales de la 

Société de Entomologique de France 9:895-920. 

– 1974a. Leptoconops (Proleptoconops) hutsoni n. sg., n. sp. du Sahara Algerien Septentrional (Diptera, Ceratopogonidae). 

Parassitologia 16:231-238. 

– 1974b. Leptoconops laoensis n. sp. du sud-est asiatique. Nouvelle Revue d'Entomologie 4:71-74. 

– 1974c. Dasyhelea insignicornis (Kieffer, 1913) et Dasyhelea wirthi nov. n. (Diptera, Ceratopogonidae). Ibid.:135. 

– 1975a. Description de quelques males d'Holoconops (Dipt. Ceratopogonidae). Annales de la Société de Entomologique 

de France 11:587-607. [Received at BMNH Feb. 23, 1976]. 

– 1975b. Le genre Leptoconops, sous-genre Holoconops en Afrique du Nord (Diptera, Ceratopogonidae). Archives de 

l'Institut Pasteur Algérie 50-51 (1972-73):23-52. [Received at BMNH Oct. 20]. 

– 1976. Alluaudomyia prima n. sp. de la Guyane Francaise (Diptera, Ceratopogonidae). Nouvelle Revue d'Entomologie 

6:205-207. 

– 1977. Description d'un nouvel Alluaudomyia de la République de Guinée (Diptera, Ceratopogonidae). Ibid. 7:345-348. 

– 1978. Deux nouveaux Alluaudomyia de la faune francaise (Dipt. Ceratopogonidae). Entomologiste 34:25-31. 

– 1979. Un nouvel Atrichopogon de la Guyane Francaise (Diptera: Ceratopogonidae). Revue Francaise d'Entomologie 1:30- 

32. 

– 1981a. Description de trois nouveaux Holoconops de la République sud-africaine. Bulletin de la Société Entomologique de 

France 86:87-97. 

– 1981b. Description d'un nouveau Leptoconops d'Israel (Diptera, Ceratopogonidae). Nouvelle Revue d'Entomologie 

11:123-126. 

– 1981c. Note sur deux Leptoconops (s.str.) du procheorient: l'un connu, l'autre nouveau pour la science (Diptera, 

Ceratopogonidae). Revue Francaise d'Entomologie 3:31-33. 

– 1982a. Description d'une nouvelle espèce afrotropicale de Tetrabezzia (Diptera, Ceratopogonidae). Cahiers ORSTOM, 

Série Entomologie Médicale et Parasitologie 20:307-312. 

180

– 1982b. Stenoxenus pastorianus n. sp. de la République de Guinée (Diptera, Ceratopogonidae). Nouvelle Revue 

d'Entomologie 12:293-297. 

– 1983a. Description de trois nouvelles espèces afrotropicales de Ceratopogonidae (Dipt., Nematocera) revenant aux 

genres Jenkinshelea et Neosphaeromias. Annales de la Société de Entomologique de France 19:261-272. 

– 1983b. Description de deux nouvelles espèces afrotropicales concernant les genres Ceratobezzia et Clinohelea (Diptera, 

Ceratopogonidae). Revue Francaise d'Entomologie 5:19-26. 

– 1983c. Ceratopogonidae des Iles Seychelles (Diptera, Nematocera). Mémoires du Muséum National d'Histoire Naturelle, 

Nouvelle Série, Série A, Zoologie 126:83 pp. 

– 1984a. Description d'une nouvelle espéce afrotropicale de genre Monohelea (Diptera, Ceratopogonidae). Nouvelle Revue 

d'Entomologie (N.S.) 1:49-53. 

– 1984b. Schizonyxhelea guyana n. g., n. sp. de la Guyane Francaise (Diptera, Ceratopogonidae). Revue Francaise 


– 1984c. Révision des espèces afrotropicales du genre Echinohelea (Diptera, Ceratopogonidae) avec description de trois 

espèces nouvelles. Bulletin du Muséum National d'Histoire Naturelle, Paris 4th Série, 6:361-376. 

– 1984d. Description de deux nouveaux Stilobezzia (Diptera, Ceratopogonidae) de la région afrotropicale. Cahiers 

ORSTOM, Série Entomologie Médicale et Parasitologie 22:43-49. 

– 1984e. Description de trois nouveaux Kolenohelea (Diptera, Ceratopogonidae) de la République de Guinée. Annales de la 


– 1985a. Ceratopogonidae de Nouvelle-Calédonie III. Genre Bezzia (Diptera, Nematocera). Cahiers ORSTOM, Série 

Entomologie Médicale et Parasitologie 23:45-54. 

– 1985b. Ceratopogonidae de Nouvelle-Calédonie. I. Genres Echinohelea Macfie, Hebetula Wirth and Debenham, 

Heterohelea n. g. (Diptera, Nematocera). Nouvelle Revue d' Entomologie (N.S.) 2:267-275. 

– 1985c. Ceratopogonidae de Nouvelle-Calédonie IV. Genre Alluaudomyia (Diptera, Nematocera). Description de six 

espèces nouvelles; simplification de l'identification des femelles. Cahiers ORSTOM, Série Entomologie Médicale et 

Parasitologie 23:187-210. 

– 1985d. Ceratopogonidae de Nouvelle-Calédonie II. Genre Monohelea (Diptera, Nematocera). Annales de Parasitologie 

Humaine et Comparée 60:747-759. 

– 1985e. Description de trois nouveaux Stilobezzia de la région afrotropicale dont la nervatin alaire est modifiée, apparentés 

a S. differens de Meillon (Dipt. Ceratopogonidae). Annales de la Société de Entomologique de France 21:457-465. 

– 1986a. Stilobezzia carayoni, n. sp. de la République de Guinée (Diptera, Ceratopogonidae). Ibid. 22:284-285. 

– 1986b. Révision des Stilobezzia africains décrits par M. Goetghebuer (Diptera, Ceratopogonidae). Revue Zoologique 

Africaine 100:363-383. 

– 1986c. Description de deux nouveaux Stilobezzia de la République de Guinée (Diptera, Ceratopogonidae). Revue 

Francaise d'Entomologie 8:63-68. 

– 1987a. Atrichopogon ornatipennis, n. sp. de la Guyane francaise (Diptera, Ceratopogonidae). Entomologiste 43:271-274. 

– 1987b. Ceratopogonidae de Nouvelle-Calédonie V. Genre Atrichopogon (Diptera, Nematocera). Cahiers ORSTOM, Série 


– 1987c. Note sur deux Atrichopogon du Zaire: A. quadrisetosus Goetghebuer et A. yamabukiensis n. sp. (Diptera, 

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– 1988a. Description d'un genre nouveau et de trois espèces nouvelles de Ceratopogonidae africains (Dipt. Nematocera). 


– 1988b. Diptères Ceratopogonidae de Nouvelle-Calédonie. 6. Note sur le genre Dasyhelea. Mémoires du Muséum 

National d'Histoire Naturelle, Série A, Zoologie 142:75-82. 

– 1988c. Description de cinq nouveaux Stilobezzia de la République de Guinée (Diptera, Ceratopogonidae). Nouvelle 

Revue d'Entomologie 5:121-132. 

– 1989a. Description de cinq nouveaux Stilobezzia originaires de la République de Guinée (Diptera, Ceratopogonidae). 

Revue Francaise d'Entomologie 11:5-14. 

– 1989b. Ceratopogonidae de Nouvelle-Calédonie. VII. Genre Paradasyhelea [Diptera, Nematocera]. Ibid.:133-137. 

– 1989c. Description de trois Brachypogon nouveaux et remarquables de la République de Guinée (Dipt. Ceratopogonidae). 


– 1990. Description de Stilobezzia seguyi, n. sp. de la République de Guinée (Diptera: Ceratopogonidae). Annales de la 


– 1991a. Description de cinq nouveaux Stilobezzia apparentés à S. insolita Das Gupta & Wirth, originaires des régions 

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181

95:297-310 (1990). 

– 1991b. Description de Forcipomyia (Dycea) madeira n. sp. de l'Ille de Madère, et présence en Finlande de F. 

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– 1992. Description de Palpomyia guyana n. sp. de la Guyane francaise (Diptera, Ceratopogonidae). Revue Francaise 


– 1993a. Description de cinq espèces nouvelles de Stiloculicoides Wirth and Grogan originaires de l'Afrique tropicale et du 

département francais de Vaucluse (Diptera, Ceratopogonidae). Nouvelle Revue d'Entomologie 10:133-143. 

– 1993b. Diptera Ceratopogonidae de Nouvelle-Calédonie. 10. Genre Monohelea. Mémoires du Muséum National 

d'Histoire Naturelle, Série A, Zoologie 157:157-164. 

Clastrier, J. and J. Boorman. 1987. Description de deux nouveaux Leptoconops l.s. de Bahrein. Bulletin de la Société 

Entomologique de France 91:293-300. 

Clastrier, J. and M. Coluzzi. 1973. Leptoconops (Leptoconops) bezzii (Noé, 1905) et Leptoconops (Leptoconops) noei n. sp. 

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Clastrier, J. and J.-C. Delécolle. 1990. Description d'un nouveau genre et de nouvelles espéces africaines des genres 

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– 1991. Diptera Ceratopogonidae de Nouvelle-Calédonie. 8. Genre Forcipomyia. Mémoires du Muséum National d'Histoire 

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– 1993. Diptera Ceratopogonidae de Nouvelle-Calédonie. 9. Genre Forcipomyia, sous-genres Lasiohelea et Microhelea. 

Ibid. 157:131-156. 

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– 1991. Forcipomyia (Trichohelea) araneivora n. sp. ectoparasite d'une aragnée habitant les monts nimba en Guinée 

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Clastrier, J. and E.M. Nevill. 1984. Leptoconops (Leptoconops) demeilloni, a new species from the Cape Duneveld of South 

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Clastrier, J. and C. Raccurt. 1979a. Quatre nouveaux Parabezzia de la Republique d'Haiti (Diptera, Ceratopogonidae). 

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– 1979b. Baeodasymyia modesta n. g., n. sp, de la République d'Haiti (Diptera, Ceratopogonidae). Annales de Parasitologie 

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Clastrier, J. and W.W. Wirth. 1961a. Notes sur les Cératopogonidés. XIII. Cératopogonidés de la région Éthiopienne. 


– 1961b. Notes sur les Cératopogonidés. XIV. Cératopogonidés de la région Éthiopienne (2). Ibid.:302-337. 

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182

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– 1904a. New Diptera from Central America. Proceedings of the Entomological Society of Washington 6:90-98. 

– 1904b. New North American Diptera. Ibid.:166-192. 

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– 1970b. Australasian Ceratopogonidae (Diptera, Nematocera). Part XII: The status of the genus Heteromyia Say in the 

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– 1970c. Australasian Ceratopogonidae (Diptera, Nematocera). Part XIII: Australian and New Guinea species of 

Echinohelea Macfie. Ibid.:145-159. 

– 1970d. Australasian Ceratopogonidae (Diptera, Nematocera). Part XIV: The genus Serromyia Meigen. Ibid.:160-165. 

– 1971. Australasian Ceratopogonidae (Diptera, Nematocera). Part XV: The genus Alluaudomyia Kieffer in Australia and 

New Guinea. Ibid. 96:128-174. 

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– 1987b. The biting midge genus Forcipomyia (Diptera: Ceratopogonidae) in the Australasian Region (exclusive of New 

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184

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– 1992. Nouvelle contribution à l'étude des Cératopogonidés de Savoie. Description de Brachypogon (Brachypogon) 

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34. 

– 1999b. Contribution à l'étude des Cératopogonidés de Suisse. III. Description de trois espèces nouvelles appartenant au 

genre Forcipomyia Meigen (Diptera, Nematocera). Bulletin de la Société Entomologique de France 104:381-392. 

Delécolle, J.-C., J. Blasco-Zumeta and J.-P. Rieb. 1997. Nouvelle contribution à l'étude des Cératopogonidés d'Espagne. 

Description de Homohelea iberica n. sp., et redescription de Palpomyia miki Goetghebuer, 1934 et de Culicoides 

brevifrontis Smatov & Isimbekov, 1971 (Diptera, Nematocera). Nouvelle Revue d'Entomologie 14:337-351. 

Delécolle, J.-C., B. Mathieu and T. Baldet. 2005. Nouvelle contribution à l'étude des Culicoides de Corse. II. Mise à jour de la 

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Delécolle, J.-C., C.P. Raccurt and C. Rebholtz. 1986. Culicoides haitiensis n. sp. et Culicoides borinqueni Fox et Hoffman en 

Haiti. I. Étude morphologue (Diptera, Ceratopogonidae). Nouvelle Revue d'Entomologie 3:107-115. 

Delfinado, M.D. 1961. The Philippine biting midges of the genus Culicoides (Diptera: Ceratopogonidae). Fieldiana Zoology 

33:627-675, pls. 8-14. 

Delfinado, M.D. and D.E. Hardy. 1971. Type specimens of Philippine Diptera. Notulae Entomologicae 51:15-32. 

De Meijere, J.C.H. 1906. Diptera. Résultats de l'Expédition Scientifique Néerlandaise à la Nouvelle-Guinée en 1903 sous les 

auspices de Arthur Wichmann. Nova Guinea 5:67-99, 1 pl. 

– 1907. Studien über Südostasiatische Dipteren. I. Tijdschrift voor Entomologie 50:196-264. 

– 1909. Blutsaugende Micro-Dipteren aus Niederlandisch Ostindien. Tijdschrift voor Entomologie 52:191-204. 

– 1915. Dipteren aus Nord-Neu-Guinea gesammelt von Dr. P.N. Van Kampen und K. Gjellerup in den Jahren 1910 und 

1911. Ibid. 58:98-139. 

– 1923. Ceratopogon-Arten als Ectoparasiten anderer Insekten. Ibid. 66:137-142. 

– 1946. Zevende Supplement op de Nieuwe Naamlijst van Nederlandsche Diptera van 1898. (Eerste Supplement op mijne 

Naamlijst van 1939). Ibid. 87:1-25. 

De Meillon, B. 1929. Some Ceratopgoninae from the Transvaal. Transaction of the Entomological Society of London 77:245- 

249. 

185

– 1931. A new species of Forcipomyia (Diptera, Ceratopogonidae) from the Transvaal, with a description of its early stages. 

Ibid. 79:335-340. 

– 1936. Entomological studies. Studies on insects of medical importance in South Africa-Part III. South African 

Ceratopogonidae. Part II. Some new and unrecorded species. Publications of the South African Institute for Medical 

Research 7:141-207. 

– 1937a. A new Jenskinshelea (Dipt. Ceratopogonidae) from southern Rhodesia. Annals of the South African Museum 

32:261-263. 

– 1937b. Entomological studies. Studies on insects of medical importance in South Africa and adjacent territories (Part IV). 

2. Records and species from South Africa. Publications of the South African Institute for Medical Research 7:332- 

385. 

– 1938. Notes on African Ceratopogonidae (Diptera). Proceedings of the Royal Entomological Society of London (B) 7:266- 

270. 

– 1939a. Notes on Ceratopogonidae (Dipt. Nematocera) from southern Africa. Journal of the Entomological Society of 

Southern Africa 1:9-25. 

– 1939b. Notes on Ceratopogonidae (Diptera Nematocera) from southern Africa - II. Ibid. 2:7-17. 

– 1939c. A new sub-genus of Ceratopogon. Ruwenzori Expedition 1934-5 (British Museum of Natural History) 1(5):103-107. 

– 1940. Ceratopogonidae (Diptera, Nematocera) from southern Africa. Transaction of the Royal Entomological Society of 

London 90:455-466. 

– 1942a. New Nematocera from the Ethiopian Region. Journal of the Entomological Society of Southern Africa 5:87-98. 

– 1942b. Ceratopogonidae (Dipt. Nematocera) from southern Rhodesia. Transaction of the Rhodesia Scientific Association 

39:113-119. 

– 1942c. Simuliidae and Ceratopogonidae (Dipt. Nematocera) from the colony of Mocambique. Estacao Anti-malarica de 

Lourenco Marques, 26 pp., pls. 1-4. 

– 1943. New records and new species of Nematocera (Diptera) from the Ethiopian Region. Journal of the Entomological 

Society of Southern Africa 6:90-113. 

– 1947. New records and species of biting insects from the Ethiopian Region. II. Ibid. 10:110-124. 

– 1952. A new species of Culicoides (Diptera: Ceratopogonidae) from Uganda. Proceedings of the Royal Entomological 

Society of London (B) 21:173. 

– 1959a. Chapter XVII Diptera (Nematocera): Ceratopogonidae. in B. Hanstrom, P. Brinck, G. Rudebeck (eds.), South 

African Animal Life. Vol. 6, pp. 325-355. Stockholm: Almquist and Wiksell. 553 pp. 

– 1959b. New Ceratopogonidae (Diptera: Nematocera) from Africa. Novos Taxa Entomologicos 13:3-24. 

– 1960. New Ceratopogonidae (Diptera: Nematocera) from the Subsaharan region. Journal of the Entomological Society of 


– 1961. The Madagascan Ceratopogonidae. Revista Entomologica Mocambique 4:37-64. 

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De Meillon, B. and F. Hardy. 1953. New records and species of biting insects from the Ethiopian Region - IV. Journal of the 

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– 1954. New records and species of biting insects from the Ethiopian region. V. Ibid. 17:62-85. 

De Meillon, B. and M. Lavoipierre. 1944. New records and species of biting insects from the Ethiopian Region. Journal of the 


De Meillon, B. and W.W. Wirth. 1955. A new species of Ceratopogon from West Africa (Diptera, Ceratopogonidae). 

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– 1979a. Subsaharan Ceratopogonidae (Diptera) I. A new South African species of Mackerrasomyia Debenham. Journal of 

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– 1979b. Subsaharan Ceratopogonidae (Diptera) II. Ceratopogonidae taken on flowers of avocado, Persea americana, in 

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– 1979c. Subsaharan Ceratopogonidae (Diptera) III. New species and records of the genus Fanthamia de Meillon. Ibid.:191- 

196. 

– 1979d. Subsaharan Ceratopogonidae (Diptera) IV. Rhinohelea, a new subgenus of Forcipomyia from the south-west Cape 

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– 1979e. A taxonomic review of the subgenus Phytohelea of Forcipomyia (Diptera: Ceratopogonidae). Proceedings of the 

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– 1980. A new subgenus of Forcipomyia, with descriptions of eight new species (Diptera: Ceratopogonidae). Ibid. 82:9-24. 

186

– 1981a. Subsaharan Ceratopogonidae (Diptera) V. Kolenohelea, a new genus of African Stilobezziini. Annals of the Natal 

Museum 24:513-523. 

– 1981b. Subsaharan Ceratopogonidae (Diptera) VI. New species and records of South African biting midges collected by 

A.L. Dyce. Ibid.:525-561. 

– 1981c. Subsaharan Ceratopogonidae (Diptera) VII. The biting midges of the Kruger National Park, South Africa, exclusive 

of the genus Culicoides. Ibid.:563-601. 

– 1983a. Subsaharan Ceratopogonidae (Diptera) IX. New species and records from southern Africa. Ibid. 25:347-381. 

– 1983b. Subsaharan Ceratopogonidae (Diptera) XI. The genus Serromyia Meigen. Ibid.:383-402. 

– 1987a. Subsaharan Ceratopogonidae (Diptera) XII. New species and records, mainly from South Africa. Journal of the 


– 1987b. Subsaharan Ceratopogonidae (Diptera) XIII. Two new South African genera of the tribe Ceratopogonini. Ibid.:383- 

391. 

– 1989a. A new pollen feeding Atrichopogon midge from Madagascar, with notes on closely related subsaharan species 

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– 1989b. Subsaharan Ceratopogonidae (Diptera) XIV. New species and records of Forcipomyia and Dasyhelea, mainly from 

Zimbabwe and Transvaal, South Africa. Journal of the Entomological Society of Southern Africa 52:201-221. 

– 1991. The genera and subgenera (excluding Culicoides) of the Afrotropical Biting Midges (Diptera: Ceratopogonidae). 

Annals of the Natal Museum 32:27-147. 

De Meillon, B., R. Meiswinkel and W.W. Wirth. 1982. Subsaharan Ceratopogonidae (Diptera) VIII. Seven new species from 

the northern Transvaal. Journal of the Entomological Society of Southern Africa 45:123-143. 

Deng, C., G. Zhan and Y.-X. Yu. 1990. Descriptions of Culicoides zhangmensis sp. nov. and Culicoides stupulosus sp. nov. 

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Deng, C.Y., Z.-Y. Liao, Y.-Z. Zhang, D.-P. Mo, and Y.-X. Yu. 2011. Two new species and a new record of biting midges from 

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Dessart, P. 1961. Contribution a l'etude des Ceratopogonidae (Diptera) (II) Revision des Forcipomyia Congolais decrits par le 

Dr. Goetghebuer. Bulletin et Annales de la Société Royale d'Entomologie de Belgique 97:315-376. 

– 1962. Contribution à l'étude des Ceratopogonidae (Diptera) IV Les Forcipomyia pollinisateurs du Cacaoyer (2). Revue de 

Zoologie et de Botanique Africaines 65:139-148. 

– 1963. Contribution à l'étude des Ceratopogonidae (Diptera) VI.- Remarques sur quelques especes du genre Forcipomyia. 

Bulletin et Annales de la Société Royale d'Entomologie de Belgique 99:182-188. 

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42. 

Ding, Z. and Y.-X. Yu. 1990. Descriptions of Lasiohelea dandongensis sp. nov. and Culicoides obscuratus sp. nov. (Diptera: 

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Dow, M.I. and E.C.Turner. 1976. A revision of the Nearctic species of the genus Bezzia (Diptera: Ceratopogonidae). 

Occasional Papers from the Department of Entomology, Virginia Polytechnic Institute and State University, Research 

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Dow, M.I. and W.W. Wirth. 1972. Studies on the genus Forcipomyia, 2. The Nearctic species of the subgenera Thyridomyia 

and Synthyridomyia (Diptera: Ceratopogonidae). Annals of the Entomological Society of America 65:177-201. 

Downes, J.A. 1955. The food habits and description of Atrichopogon pollinivorus sp. n. (Diptera: Ceratopogonidae). 

Transactions of the Royal Entomological Society of London 106:439-453. 

– 1976. A new species of Brachypogon (Diptera: Ceratopogonidae), the first record of the genus in Canada. Canadian 


– 1978. The Culicoides variipennis complex: a necessary re-alignment of nomenclature (Diptera: Ceratopogonidae). Ibid. 

110:63-69. 

Downes, J.A. and D.S. Kettle. 1952. Descriptions of three species of Culicoides Latreille (Diptera: Ceratopogonidae) new to 

187

science, together with notes on and a revised key to the British species of the pulicaris and obsoletus groups. 

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Duret, J.P. and J. Lane. 1955. Novas Heteromyia da Argentina (Diptera. Ceratopogonidae). Dusenia 6:35-40. 

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Dyce, A.L. and W.W. Wirth. 1997. The Purus group, a newly recognised natural species-group in the genus Culicoides 

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– 1959. Culicoides flavidus, a new species of the genus Culicoides Kieff. (Diptera, Heleidae) from Transcaucasia [in 

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– 1960a. Fauna of bloodsucking Heleidae (Diptera) of bottomland forest in the Kura Plain, in Azerbaidjan [in Russian, 

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– 1960b. Materials for the fauna of biting flies (Diptera, Heleidae) of Armenia with description of a new species [in Russian]. 

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– 1961a. Contribution to the systematics of bloodsucking flies of the genus Leptoconops Skuse in Transcaucasia and 

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Meditsinskikh Nauk 1961 (1): 67-76. 

– 1961b. Fauna of bloodsucking Heleidae of the genus Culicoides Latr. of the Alazan Valley in Azerbaijan [in Russian]. Ibid. 

1961 (2): 73-76. 

– 1961c. Description of three species of Heleidae from Asia Minor [in Russian]. Ibid. 1961(10): 75-80. 

– 1962a. New species of bloodsucking midges (Diptera, Heleidae) from the valley of the Kura River, Transcaucasus [in 

Russian]. Entomologicheskoe Obozrenie 41:206-219. Translation in Entomological Review 41:121-131. 

– 1962b. Biting midges (Diptera, Heleidae) from the northeast part of Azerbaidzhan [in Russian]. Entomologicheskii Sbornik. 

Baku: Izdatel'stvo Akademii Nauk Azerbaidzhanskoi SSR 1:183-192. 

– 1962c. A new species of Heleidae of the genus Lasiohelea Kieff. (Diptera) from Azerbaidzhan [in Russian]. Ibid.: 193-200. 

– 1962d. Morphology of preimaginal phases of Leptoconops bezzii muganicus Dzhaf. of blood sucking sand-flies (Diptera, 

Heleidae) [in Russian, English summary]. Zoologichesky Zhurnal 41:241-246. 

– 1963. New species of biting midges (Diptera, Heleidae) from the eastern region of Georgia (USSR) [in Russian]. Doklady 

Akademii Nauk Azerbaidzhanskoi 19:53-55. 

– 1964. Blood-sucking midges (Diptera, Heleidae) of the Transcaucasus. Akademija Nauk Azerbaidzanskoi SSR, Instituta 

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Edwards, F.W. 1916. Two new Australian Diptera. Annals and Magazine of Natural History Ser. 8, 18:498-502. 

– 1921. Diptera Nematocera from Arran and Loch Etive. Scottish Naturalist 1921:59-61, 89-92, 121-125. 

– 1922. On some Malayan and other species of Culicoides, with a note on the genus Lasiohelea. Bulletin of Entomological 

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– 1923a. New and litlle-known [sic] nematocerous Diptera from Java. Treubia 3:180-183. 

– 1923b. New and old observations on Ceratopogonine midges attacking other insects. Annals of Tropical Medicine and 

Parasitology 17:19--29. 

– 1924. II. Chironomidae. Records of the Indian Museum 26:107-108. 

– 1926a. On the British biting midges (Diptera, Ceratopogonidae). Transactions of the Entomological Society of London 

74:389-426, 2 pls. 

– 1926b. Diptera Nematocera from the mountains of Borneo. Sarawak Museum Journal 3:243-278, pls. 9-10. 

– 1928. Nematocera, p. 23-102. In: Insects of Samoa and other Samoan terrestrial Arthropoda. Part VI. Diptera. Fascicle 2. 

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– 1929a. British non-biting midges (Diptera, Chironomidae). Transactions of the Royal Entomological Society 77:279-430. 

– 1929b. Philippine nematocerous Diptera II. Notulae Entomologicae 9:1-14. 

– 1932a. Some Chironomidae (Diptera) from Barkuda Island, Chilka Lake. Records of the Indian Museum 34:177-183. 

– 1932b. Notes on Highland Diptera, with descriptions of six new species. Scottish Naturalist 194:43-52. 

– 1933a. Some Perthshire Diptera. Scottish Naturalist 201:87-92. 

188

– 1933b. Die Ausbeute der deutschen Chaco-Expedition 1925/26. Diptera (Fortsetzung) XXXIX-XLI. Cecidomyiidae, 

Chironomidae, Ceratopogonidae. Konowia 12:86-88. 

– 1933c. XX. Diptera Nematocera from Mount Kinabalu. Journal of the Federated Malay States Museums 17:223-296. 

Edwards, F.W., H. Oldroyd and J. Smart. 1939. British blood-sucking flies. London: British Museum. viii + 156 pp., 45 pls. 

Egger, J. 1863. Dipterologische Beiträge. Verhandlungen der Kaiserlich-Königlichen Zoologisch-Botanischen Gesellschaft in 

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Elson-Harris, M.M. and D.S. Kettle. 1985. A new species of Paradasyhelea Macfie (Diptera: Ceratopogonidae), with 

descriptions and keys to the immature stages of Australian Paradasyhelea. Journal of the Australian Entomological 

Society 24:233-240. 

Enderlein, G. 1908. Neue Ceratopogoninen aus Südafrika. Denkschriften der medizinisch-naturwissenschaftlichen 

Gesellschaft zu Jena 13:459-461. 

– 1912. Paryphoconus, eine neue Chironomiden-gattung aus Brasilien. Stettiner entomologische Zeitung 73:57-60. 

– 1936. Ordnung: Zweiflügler, Diptera. Abt. 16, 259 pp. In P. Brohmer, P. Ehrmann and G. Ulmer (eds.), Die Tierwelt 

Mitteleuropas 6:Insekten III Teil. Leipzig. 

Esaki, T. 1939. Injurious arthropoda to man in mandated South Sea Islands of Japan (first report). in Volumen Jubilare pro 

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Ewen, A.B. and L.G. Saunders. 1958. Contributions toward a revision of the genus Atrichopogon based on characters of all 

stages (Diptera, Heleidae). Canadian Journal of Zoology 36:671-724. 

Fabricius, J.C. 1775. Systema entomologiae, sistens insectorum classes, ordines, genera, species adiectis synonymis, locis, 

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Fei, S.-H. 1982. A new species of Culicoides from Liaoning Province (Diptera: Ceratopogonidae) [in Chinese, English 

summary]. Zoological Research 3:105-106. 

Fei, S.-H. and T.-S. Lee. 1984a. A new species of Culicoides from Qinghai Province, China [in Chinese, English summary]. 

Acta Zootaxonomica Sinica 9:182-183. 

– 1984b. Description of a new species of Culicoides from China (Diptera: Ceratopogonidae) [in Chinese, English summary]. 

Acta Entomologica Sinica 27:345-347. 

Felippe-Bauer, M.L. 1987. A new Neotropical Culicoides from Minas Gerais (Diptera: Ceratopogonidae). Memorias do 

Instituto Oswaldo Cruz 82:147-148. 

Felippe-Bauer, M.L. and R. Lourenco-de-Oliveira. 1987. A new Neotropical biting midge of the Culicoides debilipalpis group 

(Diptera: Ceratopogonidae). Ibid. 82:149-150. 

Felippe-Bauer, M.L. and A.R. Quintelas. 1993a. Two new Brazilian predaceous midges of the genus Downeshelea Wirth and 

Grogan (Diptera: Ceratopogonidae). Ibid. 88:33-38. 

– 1993b. Downeshelea bicornis, a new Neotropical predaceous midge from Brazil (Diptera: Ceratopogonidae). Ibid.: 185- 

187. 

– 1994. Culicoides lobatoi, a new Brazilian biting midge of the Limai group (Diptera: Ceratopogonidae). Ibid. 89:25-27. 

Felippe-Bauer, M.L. and C.S. Silva. 2008. Downeshela (sic) oliveirai, a new neotropical predaceous midge from northern 

Brazil (Diptera: Ceratopogonidae). Iheringia, série Zoológica 98:400-403. 

Felippe-Bauer, M.L. and G.R. Spinelli. 1991. New Brazilian predaceous midges of the genus Monohelea Kieffer and 

redescription of M. guaimiesi (Diptera: Ceratopogonidae). Memorias do Instituto Oswaldo Cruz 86:201-207. 

– 1994. Two new Neotropical species of Monohelea Kieffer and Downeshelea Wirth & Grogan (Diptera: Ceratopogonidae). 

Ibid. 89:161-165. 

– 1998. A new Neotropical species of Monohelea Kieffer from Uruguay (Diptera: Ceratopogonidae). Ibid. 93:63-64. 

Felippe-Bauer, M.L. and W.W. Wirth. 1987. A new Culicoides of the Stigmalis group (Diptera: Ceratopogonidae). Ibid. 

82:415-419. 

– 1988. Culicoides tavaresi, a new species from the state of Rio de Janeiro, Brazil (Diptera: Ceratopogonidae). Ibid. 83:261- 

263. 

Felippe-Bauer, M.L., A.G. Cáceres, C. S. Silva, W. Valderrama-Bazan and A. Gonzales-Perez. 2003. Two new Culicoides of 

the paraensis species group (Diptera: Ceratopogonidae) from the Amazonian region of Peru. Ibid. 98:1051-1058. 

– 2005. A new Culicoides (Diptera : Ceratopogonidae) of the subgenus Diphaomyia from Peru. Ibid. 100:51-53. 

Felippe-Bauer, M.L., A.G. Cáceres, C. S. Silva, W. Valderrama-Bazan, A. Gonzales-Perez, and J.M. Costa. 2008a. 

Description of Culicoides pseudoheliconiae sp. n. from Peruvian Amazon and revalidation of Culicoides contubernalis 

189

Ortiz & Leon (Diptera: Ceratopogonidae). Ibid. 103:259-262. 

– 2008b. Culicoides huaynacapaci, a new species from Department of Cajamarca, Peru (Diptera, Ceratopogonidae). 

Iheringia, série Zoológica 98:308-310. 

Felippe-Bauer, M.L., C.P. Damasceno, V. Py-Daniel, and G.R. Spinelli. 2009. Culicoides baniwa sp.nov. from the Brazilian 

Amazon Region witn a synopsis of the hylas species group (Diptera: Ceratopogonidae). Memorias do Instituto 

Oswaldo Cruz 104:851-857. 

Felippe-Bauer, M.L., C.P. Damasceno, R. Lopes da Trindade and V. Py-Daniel. 2010. A new Culicoides (Diptera: 

Ceratopogonidae) of the Reticulatus species group from Brazilian Amazon Region. Ibid. 105: 863-865. 

Felippe-Bauer, M.L., T.N. da Silva, E.S. Ribeiro and A. Borkent. 2011. A new species of Downeshelea Wirth & Grogan and a 

redescription of the male of Downeshelea cebacoi (Lane & Wirth) (Diptera: Ceratopogonidae). Zootaxa 2780:20-28. 

Felippe-Bauer, M.L., T.N. da Silva, and J.R.C. Alves. 2012. Two new species of Atrichopogon Kieffer from Rio de Janeiro, 

Brazil (Diptera: Ceratopogonidae). Zootaxa 3566: 39–50. 

Felippe-Bauer, M.L., H. Huerta and S. Ibañez Bernal. 2000. A new species of predaceous midge of the genus Monohelea 

Kieffer from Mexico (Diptera: Ceratopogonidae). Ibid. 95:815-818. 

Felippe-Bauer, M.L., A.R. Quintelas and G.R. Spinelli. 1995. A new Neotropical predaceous midge, Downeshelea deani and 

redescription of Downeshelea guianae (Wirth) (Diptera: Ceratopogonidae). Ibid. 90:395-399. 

Felippe-Bauer, M.L., R.S. Veras, E.G. Castellon and N.A. Moreira. 2000. A new Culicoides from the Amazonian Region, 

Brazil (Diptera: Ceratopogonidae). Ibid. 95:35-37. 

Fiebrig-Gertz, C. 1928. Un Diptere ectoparasite sur un Phasmide: Ceratopogon ixodoides n. sp. Annales de Parasitologie 


Fiedler, O.G.H. 1951. The South African biting midges of the genus Culicoides (Ceratopogonid., Dipt.). Onderstepoort 

Journal of Veterinary Research 25(2): 3-33. 

Floch, H. and E. Abonnenc. 1942a. Cératopogonides hématophages de la Guyane Française. Publication de l'Institut 

Pasteur de la Guyane et du Territory de L'Inini 37:1-10. 

– 1942b. Cératopogonidés hématophages de la Guyane Française - II. Ibid. 49:1-5. 

– 1949. Sur les Cératopogonidés du Venezuela Description de deux especes nouvelles: Culicoides lichyi et Lasiohelea 

danaisi. Ibid. 196:1-5. 

– 1950a. Ceratopogonides nouveaux du Venezuela Culicoides lichyi et Lasiohelea danaisi n. sp. Boletin de Entomologia 

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190

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191

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– 1911. Un chironomide nouveau de Belgique. Bezzia xanthocephala nov. sp. Ibid. 11:95-97. 

– 1912. Quelques chironomides nouveaux de Belgique. Annales Biologie Lacustre 5:204-214. 

– 1914. Contribution à l'étude des Chironomides de Belgique. Ibid. 7:165-229. 

– 1919. Observations sur les larves et les nymphes de quelques chironomides de Belgique. Ibid. 9:51-78. 

– 1920. Ceratopogoninae de Belgique. Mémoires du Musée Royal d'Histoire Naturelle de Belgique 8(3): 1-116. 

– 1921. Chironomides de Belgique et spécialement de la zone des Flandres. Ibid. 8(4): 1-208. 

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– 1923. Nouveaux matériaux pour l'étude de la faune des Chironomides de Belgique. 2 e note. Ibid. 12:103-120. 

– 1927a. Observations sur quelques Ceratopogonines de Belgique (Dipt. Chironom.). Bulletin & Annales de la Société 

Entomologique de Belgique 67:202-204. 

– 1927b. Nouveaux matériaux pour l'étude de la faune des Chironomides de Belgique. 3 e note. Annales Biologie Lacustre 

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– 1929. Deux formes nouvelles de Ceratopogonidae de Belgique. Bulletin & Annales de la Société Entomologique de 

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– 1931. Ceratopogonidae et Chironomidae nouveaux d'Europe. Ibid. 71:211-218. 

– 1932a. Ceratopogonidae et Chironomidae nouveaux ou peu connus d'Europe. Ibid. 72:125-130. 

– 1932b. Ceratopogonidae et Chironomidae. Résultats Scientifiques du Voyage aux Indes Orientales Néerlandaises de LL. 

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– 1933a. Ceratopogonidae et Chironomidae nouveaux ou peu connus d'Europe, Deuxieme Note (1). Bulletin & Annales de 

la Société Entomologique de Belgique 72:287-294. 

– 1933b. Ceratopogonidae et Chironomidae de la Sibérie Orientale. Ibid. 73:111-120. 

– 1933c. Ceratopogonidae et Chironomidae nouveaux ou peu connus d'Europe (Quatrième note). Ibid.: 353-361. 

– 1933d. Catalogue des Cératopogonides de Belgique. Ibid.: 363-372. 

– 1933e. Ceratopogonidae et Chironomidae du Congo Belge. Revue de Zoologie et de Botanique Africaines 24:129-151. 

– 1933f. 13a. Heleidae (Ceratopogonidae). in E. Lindner (ed.), Die Fliegen der palaearktischen Region 3:1-48, pls. 1-6 (= 

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192

– 1934b. Ceratopogonides et Chironomides recueillis en Palestine. Bulletin & Annales de la Société Entomologique de 

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– 1934c. Ceratopogonidae et Chironomidae nouveaux ou peu connus d'Europe (Cinquieme Note). Ibid.: 287-294. (Aug.). 

– 1934d. Cératopogonides et Chironomides du Congo Belge. -2e Note. Revue de Zoologie et de Botanique Africaines 

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– 1934e. Zur Erforschung des Persischen Golfes. (Beitrag Nr. 15). Ceratopogonidae et Chironomidae. Arbeiten über 

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Série B 8:3-14. 

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– 1935c. Un Culicoides nouveau du Bas-Congo. Revue de Zoologie et de Botanique Africaines 26:477-478. 

– 1935d. Cératopogonides récoltés par le Dr. De Wulf au Congo Belge. Ibid. 27:145-181. 

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– 1937. Ceratopogonidae et Chironomidae nouveaux ou peu connus d'Europe (7e note). Ibid. 77:273-280. 

– 1938. Note sur quelques Cératopogonides de Belgique. Ibid. 78:375-389. 

– 1939. Cératopogonides et Chironomides recueillis en Algérie. Ibid. 79:59-62. 

– 1940. Chironomides de Laponie Suédoise. Ibid. 80:55-72. 

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– 1948a. Deux Diptères nouveaux de Belgique. Ibid. 84:36-39. 

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– 1950. Ceratopogonidae et Chironomidae nouveaux ou peu connus d'Europe (Quatorzieme note). Ibid. 26(47): 1-15. 

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– 1981a. A new American genus of predaceous midges related to Palpomyia and Bezzia (Diptera: Ceratopogonidae). 

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– 1981d. A new genus of predaceous midges of the tribe Sphaeromiini from Thailand (Diptera: Ceratopogonidae). Ibid.: 

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– 1982. A new species of Neosphaeromias from the Solomon Islands (Diptera: Ceratopogonidae). Ibid. 24:198-201. 

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– 1954. The speciation of the genus Culicoides (Diptera, Heleidae). Ibid. 47:34-51. 

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Kieffer, J.J. 1899. Description d'un nouveau genre et tableau des genres européens de la famille des Chironomides (Dipt.). 


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348. 

– 1906c. Description d'un genre nouveau et de quelques espèces nouvelles de diptères de l'Amérique de Sud. Ibid.: 349- 

358. 

– 1908. Description d'une espèce nouvelle de chironomides d'Égypte. Annales Historico-Naturales Musei Nationalis 

Hungarici 6:576-577. 

– 1909a. Considérations sur le genre Stenoxenus Coq. avec description d'une espèce nouvelle. Ibid. 7:46-49. 

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198

– 1910. Etude sur les chironomides des Indes Orientales, avec description de quelques nouvelles especes d'Egypte. 

Memoirs of the Indian Museum 2:181-242. 

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pls. 6-7. 

– 1911b. Les chironomides de (Tendipedidae) de l'Himalaya et d'Assam. Ibid.: 319-349. (Dec.). 

– 1911c. The Percy Sladen Trust Expedition to the Indian Ocean in 1905. Under the leadership of Mr. J. Stanley Gardiner. 

Vol. 3. No. XV. - Diptera, Chironomidae der Seychellen-inseln, aus der Sammlung von Mr. H. Scott. Transactions of 

the Linnean Society of London (2nd Ser.) 14:331-366. 

– 1911d. Nouvelles descriptions de chironomides obtenus d'éclosion. Bulletin de la Société d'Histoire Naturelle de Metz 

27:1-60. (Nov. 1). 

– 1911e. Bemerkungen zur Arbeit des Herrn Dr. Speiser über die Dipteren-Gruppe der sogenanneten Heleinae. 

Zoologische Jahrbücher. Abteilung für Systematik, Geographie und Biologie der Tiere 30:509-526. 

– 1912a. Tendipedidae (Chironomidae) (Dipt.). Supplementa Entomologica 1:27-43. 

– 1912b. Quelques nouveaux tendipédides (Dipt.) obtenus d'éclosion (2e note). Bulletin de la Société Entomologique de 

France 1912:101-103. 

– 1912c. Nouveaux Chironomides (Tendipedidae) de Ceylan. Spolia Zeylanica 8:1-24. 

– 1913a. Nouveaux chironomides (tendipédides) d'Allemagne. Bulletin de la Société d'Histoire Naturelle de Metz 28:7-35. 

– 1913b. Nouvelle contribution a la connaissance des tendpédides d'Allemagne. Ibid.: 37-44. 

– 1913c. Dasyhelea halophila n. sp., eine neue halophile Zuckmücke. Biologischen Centralblatt 33:255-258. 

– 1913d. Nouvelle etude sur les chironomides de l'Indian Museum de Calcutta. Records of the Indian Museum 9:119-197, 

pls. 11-12. 

– 1913e. Chironomidae et Cecidomyidae. in C. A. Alluaud and R. Jeannel (eds.), Voyage de Ch. Alluaud et R. Jeannel en 

Afrique orientale (1911-1912). Resultats scientifiques. Diptera (= Vol. 5): 1-43, Paris, 351 pp. 

– 1914a. Zwölf neue Culicoidinenarten. Archiv für Hydrobiologie und Planktonkunde, Supplement 2:231-241. 

– 1914b. South African Chironomidae (Diptera). Annals of the South African Museum 10:259-270. 

– 1914c. Quelques nouveaux chironomides des Indes. Records of the Indian Museum 10:313-315. 

– 1915a. Neue halophile Chironomiden. Archiv für Hydrobiologie, Suppl. 2:472-482. 

– 1915b. Neue Chironomiden aus Mitteleuropa. Brotéria, Serie Zoologica 13:65-87. 

– 1915c. Über dänische Chironomiden. Entomologiske Meddelelser 10:280-297 [280-284 published July 31; 285-297 

published Dec. 31]. 

– 1916a. Tendipedides (chironomides) de Formose. Annales Historico-Naturales Musei Nationalis Hungarici 14:81-121. 

– 1916b. Tendipedidae (Dipt.). Supplementa Entomologica 5:114-117. 

– 1917a. Chironomides d'Australie conservés au Musée National Hongrois de Budapest. Annales Historico-Naturales Musei 

Nationalis Hungarici 15:175-228. 

– 1917b. Chironomides d'Amérique conservés au Musée National Hongrois de Budapest. Ibid.: 292-364. 

– 1918a. Chironomides d'Afrique et d'Asie conservés au Museum National Hongrois de Budapest. Ibid. 16:31-136. 

– 1918b. Beschreibung neuer, auf Lazarettschiffen des östlichen Kriegsschauplatzes und bei Ignalino in Litauen von Dr. W. 

Horn gesammelter Chironomiden, mit Uebersichtstabellen einiger Gruppen von paläarktischen Arten (Dipt.). 

Entomologische Mitteilungen 7:35-53, 94-110, 163-170, 177-188. 

– 1919a. Chironomides d'Europe conservés au Musée National Hongrois de Budapest. Annales Historico-Naturales Musei 

Nationalis Hungarici 17:1-160. (Dec. 29). 

– 1919b. Chironomiden der Nördlichen Polarregion. Entomologische Mitteilungen 8:40-48 [Received at BMNH May 3], 110- 

120 [Received at BMNH Aug. 23]. 

– 1919c. Observations sur les chironomides (Dipt.) décrits par J. R. Malloch. Bulletin de la Société Entomologique de 

France 1919:191-194. 

– 1921a. Chironomides nouveaux ou peu connus de la région paléarctique. Bulletin de la Société d'Histoire Naturelle de la 

Moselle 29:51-109. 

– 1921b. Chironomides de l'Afrique Équatoriale. Annales de la Société Entomologique de France 90:1-56. 

– 1921c. Description de quelques Chironomides exotiques. Annales de la Société Scientifique de Bruxelles 40:181-186. 

– 1921d. Chironomides de Courlande. Ibid.: 275-298. (June 30). 

– 1921e. Notes synonymiques. Bulletin de la Société Entomologique de France 1921:7. 

– 1921f. Neue Chironomiden aus Mitteleuropa. Archiv für Hydrobiologie, Suppl. 2:785-808. 

– 1921g. Chironomides des Philippines et de Formose. Philippine Journal of Science 18:557-593. 

– 1921h. Sur quelques Diptères piqueurs de la tribu des Ceratopogoninae. Archives de l'Institut Pasteur de l'Afrique du Nord 

199

1:107-115. (March). 

– 1921i. Nouvelles observations sur les Diptères piqueurs de la tribu des Ceratopogoninae. Ibid.: 262-268. 

– 1922a. Notice sur quelques chironomides d'Amérique et de Nouvelle-Zélande. Annales de la Société Linnéenne de Lyon 

68:145-148 (1921). 

– 1922b. Étude sur les chironomides de Formose. Ibid.: 149-163 (1921). 

– 1922c. Nouveaux Chironomides piqueurs habitant le Sleswig-Holstein. Annales de la Société Scientifique de Bruxelles 

41:230-237. 

– 1922d. Nouveaux Chironomides à larves aquatiques. Ibid.: 355-367. 

– 1922e. Chironomides nouveaux ou peu connus de la région Paléarctique. Ibid. 42:71-128. 

– 1922f. Observations biologiques sur les Chironomides piqueurs avec descriptions de deux espèces nouvelles. Archives 

de l'Institut Pasteur de l'Afrique du Nord 2:387-392. 

– 1922g. Nouveaux Chironomides piqueurs habitant l'Algérie. Ibid.: 494-518. 

– 1923a. Ceratopogonines recueillis au Sahara Constantinois. Archives de l'Institut Pasteur Algérie 1:654-683. [Received at 

BMNH May 5, 1924]. 

– 1923b. Chironomides piqueurs de Java. Annales de la Société Scientifique de Bruxelles 43:134-143. (Dec. 12). 

– 1924a. Chironomides nouveaux ou rares de l'Europe centrale. Bulletin de la Société d'Histoire Naturelle de la Moselle 

30:10-110. 

– 1924b. Quelques nouveaux chironomides piqueurs de l'Europe centrale. Archives de l'Institut Pasteur Algérie 2:391-408. 

– 1924c. Quelques chironomides nouveaux et remarquables du nord de l'Europe. Annales de la Société Scientifique de 

Bruxelles 43:390-397. 

– 1925a. Nouveaux genres et nouvelles espèces de chironomides piqueurs. Archives de l'Institut Pasteur Algérie 3:405-430. 

– 1925b. 16. Chironomiden der Hochmoore Nordeuropas und des östlichen Mitteleuropas. Beitrage zur Kunde Estlands 

10:145-163. 

– 1925c. Chironomides de la République Argentine. Annales de la Société Scientifique de Bruxelles 44:73-92. 

– 1925d. Diptères (Nématocères piqueurs): Chironomidae Ceratopogoninae. Faune de France 11:1-139. 

– 1925e. Chironomides d'Egypte (Dipt.). Bulletin de la Société Royale Entomologique d'Egypte 1924:244-312. 

– 1927. Weitere Beiträge zur Chironomidenfauna Estlands. Tartu Ülikooli juures oleva Loodusuurijate Seltsi aruanded 

toimetanud H.G. Perlitz 33:59-70 (1926). 

Kieffer, J.J. and A. Thienemann. 1908. Neue und bekannte Chironomiden und ihre Metamorphose. Zeitschrift für 

Wissenschaftliche Insektenbiologie 4:1-10, 33-39, 78-84, 124-128, 184-190, 214-219, 256-259, 277-286. 

Kinoshita, S. 1918. Studies on a bloodsucking Culicoides in Korea [in Japanese]. Dobutsugaki Zasshi (Zoological Magazine) 

30:155-160, pl. 3. 

Kitaoka, S. 1973. Descriptions of four new species and the hitherto unknown males of four species of the Culicoides (Diptera, 

Ceratopogonidae) from Amami-Oshima, Japan. National Institute of Animal Health Quarterly 13:211-219. 

– 1975. Five new species of Culicoides (Diptera: Ceratopogonidae) of the Nansei Islands. Ibid. 15:192-200. 

– 1980. Six new species and males of some species of Japanese Culicoides (Diptera: Ceratopogonidae). Ibid. 20:11-22. 

– 1983. Five new species of the genus Culicoides (Diptera: Ceratopogonidae) from West Malaysia. Ibid. 23:92-98. 

– 1984. Three new species of Culicoides biting midges (Diptera: Ceratopogonidae) from Japan. Japanese Journal of 

Sanitary Zoology 35:301-305. 

– 1991. Two new species of biting midges of the genus Culicoides (Diptera: Ceratopogonidae) from northernmost Honshu, 

Japan. Ibid. 42:289-292. 

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201

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Lundström, C. 1910. Beiträge zur Kenntnis der Dipteren Finlands VI. Chironomidae. Acta Societatis pro Fauna et Flora 

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– 1916a. Beiträge zur Kenntnis der Dipteren Finlands X. Suppl. 4. Bibionidae, Chironomidae, Tipulidae. Ibid. 44(2): 1-26. 

– 1916b. Dipteren au Sarekgebiet. Diptera Nematocera. Naturwissenschaftliche Untersuchungen des Sarekgebirges in 

Schwedisch-Lappland 4:666-680. 

Lutz, A. 1913. Contribuicao para o estudo das Ceratopogoninas hematofagas do Brazil. Memorias do Instituto Oswaldo Cruz 

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– 1914. Contribuicao para o conhecimento das Ceratopogoninas do Brasil. Ibid. 6:81-99, pls. 8-9 

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– 1925. A new blood-sucking midge from Singapore. Ibid.: 349-351. 

– 1926. Ceratopogonidae from Dar-Es-Salaam. Ibid. 16:355-357. 

205

– 1932a. Ceratopogonidae from the wings of dragonflies. Tijdschrift voor Entomologie 75:265-283. 

– 1932b. Some new or little-known Ceratopogonidae. Annals and Magazine of Natural History Ser. 10, 9:485-499. 

– 1932c. New Zealand biting midges (Diptera, Ceratopogonidae). Annals of Tropical Medicine and Parasitology 26:23-53. 

– 1933a. Ceratopogonidae from the Society Islands. Bulletin of the Bernice P. Bishop Museum 113:75-80. 

– 1933b. Ceratopogonidae from the Marquesas Islands. Ibid. 114:93-103. 

– 1933c. A new species of Culicoides from Palestine. Annals of Tropical Medicine and Parasitology 27:79-81. 

– 1934a. Notes on Ceratopogonines (Dipt.) from Hawaii. Stylops 3:133-134. 

– 1934b. A new British Forcipomyia (Diptera, Ceratopogonidae). Entomologist's Monthly Magazine 70:144-145. (June). 

– 1934c. Report on a collection of Ceratopogonidae from Malaya. Annals of Tropical Medicine and Parasitology 28:177-194, 

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– 1934d. Fauna Sumatrensis. Bijdrage No. 75, Ceratopogonidae (Diptera). Tijdschrift voor Entomologie 77:202-231. (Nov. 

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– 1935a. Ceratopogonidae (Dipt.) from the river Amazon. Stylops 4:49-56. 

– 1935b. A new ceratopogonid (Dipt.) from British Guiana. Ibid.: 265. 

– 1936a. Two new species of Ceratopogonidae (Diptera) from the wings of dragonflies. Proceedings of the Royal 

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– 1936b. Four species of Ceratopogonidae (Diptera) from the wings of insects. Ibid.: 227-230. 

– 1937a. Ceratopogonidae from Trinidad. Annals and Magazine of Natural History Ser. 10, 20:1-18. 

– 1937b. Ceratopogonidae (Diptera) from Ethiopia and British Somaliland. Proceedings of the Royal Entomological Society 

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– 1937c. Notes on Ceratopogonidae (Diptera). Ibid.: 111-118. 

– 1937d. Three new species of Culicoides (Diptera, Ceratopogonidae) from Malaya. Annals of Tropical Medicine and 

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– 1940b. Ceratopogonidae (Diptera) from British Guiana and Trinidad. Part 1. Ibid.: 179-186. 

– 1940c. Ceratopogonidae (Diptera) from British Guiana and Trinidad. Part 2. Ibid.: 187-195. 

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– 1943b. A new species of Dasyhelea (Diptera) from the Bahama Islands. Ibid.: 119-120. 

– 1944a. A new species of Homobezzia (Diptera, Ceratopogonidae) from Egypt. Ibid. 13:125-126. 

– 1944b. Ceratopogonidae collected in Trinidad from cacao flowers. Bulletin of Entomological Research 35:297-300. 

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– 1946. A new species of Culicoides (Diptera, Ceratopogonidae) from Fiji. Ibid. 15:15-16. 

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– 1947b. Ceratopogonidae from the Anglo-Egyptian Sudan. Ibid.: 69-78. 

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– 1950b. Estudios en Culicoides IV. Revisión de las especies americanas del sub-género Hoffmania Fox 1948, con la 

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– 1950c. Estudios en Culicoides V. Informes sobre una nueva especie y lista de los machos cuyas genitalias son conocidas. 

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– 1950d. Descripcion de Stilobezzia glauca venezuelensis n. vr. y Monohelea mayeri n. sp. Memorias de la Sociedad de 

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210

(Townsend, 1893). Ibid. 17:163-171. 

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– 1953a. Nueva contribución al conocimiento de los caracteres morfológicos externos de las hembras americanas del 

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– 1952b. Sobre las especies Americanas del género "Culicoides" Latr., (Diptera, ACeratopogonidae@) con una espermateca. 

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– 1990a. Biting midges (Insecta: Diptera: Ceratopogonidae) from Sakhalin amber. Prace Muzeum Ziemi 41:77-81. 

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– 1992a. Ceratopogonidae (Diptera) from Algeria. VII. Stilobezzia and Kolenohelea. Acta Universitatis Nicolai Copernici 

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– 1992b. A new biting midge of genus Jenskinshelea from the Democratic People's Republic of Korea (Diptera, 

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– 1993. Biting midges (Diptera, Ceratopogonidae) from Miocene Saxonian amber. Acta Zoologica Cracoviensia 35:603-656. 

– 1996. Biting midges from Lower Cretaceous amber of Lebanon and Upper Cretaceous Siberian amber of Taimyr (Diptera, 

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– 2000a. Biting midges (Diptera: Ceratopogonidae) from the Lower Cretaceous amber of Jordan. Ibid. 69:251-256. 

– 2000b. A new species of Dasyhelea from the Canary Islands (Diptera: Ceratopogonidae). Ibid.: 477-481. 

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Szadziewski, R. and D.V. Hagan. 2000. Two new species of biting midges from Norway (Diptera: Ceratopogonidae).Ibid. 

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Szadziewski, R. and P. Havelka. 1984. A review of the Palaearctic biting midges of the subgenus Brachypogon (s. str.) 

220

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– 1940d. Biting midges from Japan and neighbouring countries, including Micronesian Islands, Manchuria, North China and 

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– 1974. Family Ceratopogonidae. in A Catalog of the Diptera of the Americas south of the United States, Fasc. 14:89 pp. 

– 1976a. A new species and new records of Dasyhelea from the Tonga Islands and Samoa (Diptera: Ceratopogonidae). 

Proceedings of the Hawaiian Entomological Society 22:381-383. 

– 1976b. Forcipomyia pictoni Macfie and descriptions of two new related species from Florida (Diptera: Ceratopogonidae). 

Florida Entomologist 59:77-84. 

– 1976c. New names for North American Ceratopogonidae (Diptera). Proceedings of the Entomological Society of 


– 1977. A new Culicoides biting midge from California (Diptera: Ceratopogonidae). Pan-Pacific Entomologist 53:53-55. 

– 1978. New species and records of intertidal biting midges of the genus Dasyhelea Kieffer from the Gulf of California 

(Diptera: Ceratopogonidae). Pacific Insects 18:191-198. 

– 1980. A new species and corrections in the Atrichopogon midges of the subgenus Meloehelea attacking blister beetles 


– 1981. Paradasyhelea harrisoni n. sp. from the Auckland Islands and additional records of subantarctic Ceratopogonidae. 

New Zealand Journal of Zoology 8:383-386. 

– 1982a. The cacao-pollinating midges of the Forcipomyia argenteola group (Diptera: Ceratopogonidae). Proceedings of the 


– 1982b. New species of Neotropical Culicoides (Diptera: Ceratopogonidae). Florida Entomologist 65:248-253. 

– 1983a. The North American species of the cockerelli and dorsasetula groups of the predaceous midge genus Bezzia, 

subgenus Homobezzia (Diptera: Ceratopogonidae). Proceedings of the Entomological Society of Washington 

85:762-782. 

– 1983b. The North American predaceous midges of the Bezzia bicolor group (Diptera: Ceratopogonidae). Florida 


– 1987. A new species of Dasyhelea (Diptera: Ceratopogonidae) from rock pools in the southwestern United States. Journal 

of the North American Benthological Society 6:72-76. 

– 1990. The biting midges of Aldabra Atoll, Indian Ocean (Diptera: Ceratopogonidae). Proceedings of the Entomological 


225

– 1991a. The predaceous midge genus Allohelea Kieffer in the western hemisphere (Diptera: Ceratopogonidae). Florida 


– 1991b. New and little-known species of Forcipomyia (Diptera: Ceratopogonidae) associated with cocoa pollination in 

Brazil. Proceedings of the Entomological Society of Washington 93:163-175. 

– 1992. A new genus of Ceratopogonini (Diptera: Ceratopogonidae) from Brazil. Ibid. 94:276-281. 

– 1994a. The subgenus Atrichopogon (Lophomyidium) with a revision of the Nearctic species (Diptera: Ceratopogonidae). 

Insecta Mundi 8:17-36. 

– 1994b. New species and records of predaceous midges of the genus Probezzia from Florida and Alabama (Diptera: 

Ceratopogonidae). Florida Entomologist 77:136-145. 

– 1994c. The western hemisphere species of the predaceous midge genus Echinohelea, with descriptions of six new species 

(Diptera: Ceratopogonidae). Insecta Mundi 8:227-242. 

Wirth, W.W. and P.H. Arnaud. 1969. Polynesian biting midges of the genus Culicoides (Diptera: Ceratopogonidae). Pacific 

Insects 11:507-520. 

Wirth, W.W. and W.R. Atchley. 1973. A review of the North American Leptoconops (Diptera: Ceratopogonidae). Graduate 

Studies Texas Tech University (5), 57 pp. 

Wirth, W.W. and P. Barreto. 1978. New species of Culicoides biting midges (Diptera: Ceratopogonidae) from Colombia. 

Journal of Medical Entomology 14:553-564. 

Wirth, W.W. and R.A. Beaver. 1979. The Dasyhelea biting midges living in pitchers of Nepenthes in southeast Asia (Diptera, 

Ceratopogonidae). Annales de la Société Entomologique de France 15:41-52. 

Wirth, W.W. and F.S. Blanton. 1953a. Studies in Panama Culicoides (Diptera: Heleidae): I, Descriptions of six new species. 

Journal of the Washington Academy of Sciences 43:69-77. 

– 1953b. Studies in Panama Culicoides (Diptera, Heleidae). II. Descriptions of six additional new species. Journal of 

Parasitology 39:229-236. 

– 1953c. Studies in Panama Culicoides (Diptera, Heleidae). III. A new species related to phlebotomus (Williston). 

Entomological News 64:113-120. 

– 1955a. Studies in Panama Culicoides (Diptera, Heleidae) IV. Descriptions of three new species. Bulletin of the Brooklyn 

Entomological Society 50:100-106. (October). 

– 1955b. Studies in Panama Culicoides (Diptera, Heleidae) V. Descriptions of three new species of the subgenus Oecacta 

Poey. Ibid.: 121-127. 

– 1956a. A new species of salt-marsh sand fly from Florida, the Bahamas, Panama and Ecuador: its distribution and 

taxonomic differentiation from Culicoides furens (Poey) (Diptera, Heleidae). Florida Entomologist 39:157-162. 

– 1956b. Studies in Panama Culicoides VII. The species of the pulicaris and cova-garciai groups (Diptera, Heleidae). 


– 1956c. Studies in Panama Culicoides (Diptera, Heleidae). IX. Two new species related to leoni Barbosa and reevesi Wirth. 

Bulletin of the Brooklyn Entomological Society 51:45-52. 

– 1959. Biting midges of the genus Culicoides from Panama (Diptera: Heleidae). Proceedings of the United States National 

Museum 109:237-482. 

– 1966. A new day-biting sand fly from the southeastern States (Diptera, Ceratopogonidae). Florida Entomologist 49:279- 

281. 

– 1967. The North American Culicoides of the guttipennis group (Diptera: Ceratopogonidae). Ibid. 50:207-232. 

– 1968a. A revision of the Neotropical biting midges of the hylas group of Culicoides (Diptera: Ceratopogonidae). Ibid. 

51:201-215. 

– 1968b. A new Culicoides from Guyana (Diptera: Ceratopogonidae). Ibid.: 251-252. 

– 1969a. New species and records of Culicoides from western North America. Proceedings of the Entomological Society of 


– 1969b. North America [sic] Culicoides of the pulicaris group (Diptera: Ceratopogonidae). Florida Entomologist 52:207-243. 

– 1969c. A new Nearctic species of the genus Paradasyhelea Macfie. Pan-Pacific Entomologist 45:97-100. 

– 1970a. New genera of Neotropical Ceratopogonidae (Diptera). Florida Entomologist 53:7-14. 

– 1970b. New species of Neotropical Culicoides (Diptera: Ceratopogonidae). Ibid.: 39-45. 

– 1970c. Notes on Brachypogon Kieffer (Diptera, Ceratopogonidae), a new species and two new Neotropical genera of the 

tribe Ceratopogonini. Ibid.: 93-104. 

– 1970d. A review of the Culicoides Nigrigenus group, with two new species (Diptera: Ceratopogonidae). Entomological 

News 81:141-151. 

– 1971a. New Neotropical sandflies of the Culicoides debilipalpis group (Diptera: Ceratopogonidae). Proceedings of the 

226


– 1971b. New western Culicoides of the stonei group. Journal of the Kansas Entomological Society 44:459-467. 

– 1971c. New species and synonymy of Florida Culicoides (Diptera: Ceratopogonidae). Florida Entomologist 54:73-78. 

– 1972a. Lanehelea, a new Neotropical genus of Sphaeromiini (Diptera: Ceratopogonidae). Studia Entomologica 15:433- 

438. 

– 1972b. A new Patagonian biting midge of the the genus Monohelea, Isthmohelea (Diptera: Ceratopogonidae). Florida 


– 1973. A review of the maruins or biting midges of the genus Culicoides (Diptera: Ceratopogonidae) in the Amazon Basin. 

Amazoniana 4:405-470. 

– 1974a. A new Florida sand fly closely related to Culicoides haematopotus Malloch (Diptera: Ceratopogonidae). Florida 


– 1974b. New synonymy and a correction in the Culicoides piliferus group (Diptera: Ceratopogonidae). Ibid.: 71-75. 

– 1978. Two new species of Neotropical Culicoides (Diptera: Ceratopogonidae). Pan-Pacific Entomologist 54:236-240. 

Wirth, W.W. and J.L. Castner. 1990. New Neotropical species of "Stick-tick" (Diptera: Ceratopogonidae) from katydids. 


Wirth, W.W. and M.L. Debenham. 1977. Hebetula, a new genus of the predaceous midge tribe Sphaeromiini (Diptera: 


Wirth, W.W. and M.D. Delfinado. 1964. Revision of the oriental species of Alluaudomyia Kieffer (Diptera, Ceratopogonidae). 

Pacific Insects 6:599-648. 

Wirth, W.W. and J.O. Derron. 1976. New species and records of biting midges (Diptera: Ceratopogonidae) associated with 

cacao in Sao Tomé, West Africa. Bulletin de la Société Entomologique Suisse 49:229-237. 

Wirth, W.W. and M.I. Dow. 1971. Studies on the genus Forcipomyia III. Blantonia, a new subgenus in the Trichohelea 

complex (Diptera: Ceratopogonidae). Florida Entomologist 54:289-295. 

– 1972. Studies on the genus Forcipomyia 4. Rhynchoforcipomyia, a new Neotropical subgenus in the Trichohelea complex 

(Diptera: Ceratopogonidae). Annals of the Entomological Society of America 65:862-872. 

Wirth, W.W. and F.E. Giles. 1990. New species and records of predaceous midges from Fiji (Diptera: Ceratopogonidae). 


Wirth, W.W. and W.L. Grogan. 1977. Taxonomic notes on the genus Heteromyia Say and a new species from Nicaragua 


– 1979. Natural history of Plummers Island, Maryland. XXIV. Biting midges (Diptera: Ceratopogonidae). 2. The species of the 

tribes Heteromyiini and Sphaeromiini. Proceedings of the Biological Society of Washington 91:847-903. 

– 1981. Natural History of Plummer's Island, Maryland XXV. Biting midges (Diptera: Ceratopogonidae). 3. The species of the 

tribe Stilobezziini. Bulletin of the Biological Society of Washington 5:1-102. 

– 1982. The predaceous midges of the genus Phaenobezzia in North America (Diptera: Ceratopogonidae). Memoirs of the 


– 1983. The Nearctic species of the Bezzia bivittata group (Diptera: Ceratopogonidae). Proceedings of the Biological 


– 1988. The predaceous midges of the world (Diptera: Ceratopogonidae; Tribe Ceratopogonini). Flora and Fauna Handbook 

4. E. J. Brill, xv + 160 pp. 

Wirth, W.W. and F.G. Howarth. 1982. The "Forcipomyia ingrami" complex in Hawaii (Diptera: Ceratopogonidae). 

Proceedings of the Hawaiian Entomological Society 24:127-151. 

Wirth, W.W. and A.A. Hubert. 1959. Trithecoides, a new subgenus of Culicoides (Diptera, Ceratopogonidae). Pacific Insects 

1:1-38. 

– 1960a. Ceratopogonidae (Diptera) reared from cacti, with a review of the copiosus group of Culicoides. Annals of the 

Entomological Society of America 53:639-658. 

– 1960b. Camptopterohelea a new genus of Ceratopogonidae from the Philippines (Diptera). Fieldiana, Zoology 42:89-91. 

– 1961. New species and records Taiwan Culicoides (Diptera: Ceratopogonidae). Pacific Insects 3:11-26. 

– 1962. The species of Culicoides related to piliferus Root and Hoffman in eastern North America (Diptera, 


– 1972. A new Oriental species of Culicoides breeding in tree rot cavities (Diptera: Ceratopogonidae). Journal of the 

Washington Academy of Sciences 62:41-42. 

– 1989. The Culicoides of Southeast Asia (Diptera: Ceratopogonidae). Memoirs of the American Entomological Institute 

44:i-iv, 1-508. 

Wirth, W.W. and R.H. Jones. 1956. Three new North American species of tree-hole Culicoides (Diptera, Heleidae). 

227


– 1957. The North American subspecies of Culicoides variipennis (Diptera, Heleidae). United States Department of 

Agriculture Technical Bulletin 1170, iii + 35 pp. 

Wirth, W.W. and D.J. Lee. 1958. Australasian Ceratopogonidae (Diptera, Nematocera). Part VIII: A new genus from Western 

Australia attacking man. Proceedings of the Linnean Society of New South Wales 83:337-339. 

– 1959. The genus Paradasyhelea Macfie, with descriptions of two new species from eastern Australia (Diptera: 

Ceratopogonidae). Bulletin of the Brooklyn Entomological Society 54:114-121. 

– 1967. New species of Culicoides from high altitudes in the Colombian Andes (Diptera: Ceratopogonidae). Proceedings of 

the United States National Museum 124:1-22. 

Wirth, W.W. and J.R. Linley. 1990. Description of Dasyhelea chani new species (Diptera: Ceratopogonidae) from leaves of 

the water lettuce (Pistia stratiotes) in Floirda [sic]. Florida Entomologist 73:273-279. 

Wirth, W.W. and D.H. Messersmith. 1971. Studies on the genus Forcipomyia. 1. The North American parasitic midges of the 

subgenus Trichohelea (Diptera: Ceratopogonidae). Annals of the Entomological Society of America 64:15-26. 

– 1977. Notes on the biting midges of the Seychelles (Diptera: Ceratopogonidae). Proceedings of the Entomological Society 

of Washington 79:293-309. 

Wirth, W.W. and A.P.A. de Moraes. 1979. New records and new species of biting midges from salt marshes in California and 

Mexico (Diptera: Ceratopogonidae). Pan-Pacific Entomologist 55:287-298. 

Wirth, W.W. and C. Morris. 1985. The taxonomic complex, Culicoides variipennis. In Bluetongue and related orbiviruses, pp. 

165-175. New York: Alan R. Liss, Inc., xxi + 746 pp. 

Wirth, W.W. and B.A. Mullens. 1992. Culicoides boydi (Diptera: Ceratopogonidae): a potential vector of Hemorrhagic Disease 

Viruses to Desert Bighorn Sheep in southern California. Journal of Medical Entomology 29:1006-1010. 

Wirth, W.W. and N.C. Ratanaworabhan. 1971a. Ceratoculicoides, a new genus related to Ceratopogon Meigen (Diptera: 


– 1971b. Notes on Neotropical Pellucidomyia (Diptera, Ceratopogonidae). Annals of the Entomological Society of America 

64:446-448. 

– 1972a. Neobezzia, a new Neotropical biting midge genus of the tribe Sphaeromiini (Diptera: Ceratopogonidae). Journal of 

the Kansas Entomological Society 45:476-490. 

– 1972b. A new genus of biting midge from California related to Neurohelea Kieffer (Diptera, Ceratopogonidae). Pan-Pacific 


– 1972c. A revision of the tribe Stenoxenini (Diptera: Ceratopogonidae). Annals of the Entomological Society of America 

65:1368-1388. 

– 1972d. Notes on the genus Macropeza Meigen and description of a new species from Florida (Diptera: Ceratopogonidae). 


– 1973. Pseudostilobezzia, a new genus of biting midge from Viet Nam (Diptera: Ceratopogonidae). Proceedings of the 


– 1976. A new species of parasitic midge (Forcipomyia (Pterobosca)) from Aldabra, with descriptions of its presumed larva 

and pupa and systematic notes on the subgenera of Forcipomyia (Ceratopogonidae). Systematic Entomology 1:241- 

245. 

– 1978. Studies on the genus Forcipomyia. V. Key to subgenera and description of a new subgenus related to 

Euprojoannisia Brethes (Diptera: Ceratopogonidae). Proceedings of the Entomological Society of Washington 

80:493-507. 

– 1981a. The Oriental species of the genus Dibezzia Kieffer (Diptera: Ceratopogonidae). Ibid. 83:287-295. 

– 1981b. New species and records of predaceous midges (Diptera: Ceratopogonidae) from rice paddies in Thailand. Pacific 

Insects 23:396-431. 

– 1992. Two new Oriental species of Atrichopogon related to A. jacobsoni (Diptera: Ceratopogonidae). Oriental Insects 

26:265-274. 

– 1993. The Oriental biting midges of the genus Atrichopogon related to Dolichohelea polita Edwards (Diptera: 


Wirth, W.W. and W.A. Rowley. 1971. A revision of the palmerae group of the genus Culicoides. Journal of the Kansas 


Wirth, W.W. and S.J. Soria. 1975. A new Neotropical Forcipomyia midge closely related to F. (F.) genualis (Loew) (Diptera: 

Ceratopogonidae). Revista Theobroma 5:19-27. 

– 1980. Studies on the genus Forcipomyia VI. The Neotropical species of the subgenus Warmkea (Diptera: 


228

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– 1993a. The American species of the annulatipes group of the subgenus Lepidohelea, genus Forcipomyia (Diptera: 

Ceratopogonidae). Insecta Mundi 6:109-125. (1992) 

– 1993b. The North American species of the Forcipomyia (Lepidohelea) bicolor subgroup (Diptera: Ceratopogonidae). 


Wirth, W.W. and Y. Wada. 1979. Two new species of the genus Camptopterohelea Wirth and Hubert from southeast Asia 


Wirth, W.W. and W.T. Waugh. 1976. Five new Neotropical Dasyhelea midges (Diptera: Ceratopogonidae) associated with 

culture of Cocoa. Studia Entomologica 19:223-236. 

Wirth, W.W. and R.W. Williams. 1957. The biting midges of the Bermuda Islands, with descriptions of five new species. 


– 1964. New species and records of North American Monohelea (Diptera: Ceratopogonidae). Annals of the Entomological 

Society of America 57:302-310. 

Wirth, W.W., B. de Meillon and E. Haeselbarth. 1980. 10. Family Ceratopogonidae. in R. W. Crosskey (ed.), Catalogue of 

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Xue, J. and Y.-X. Yu. 1991. A new species of Culicoides from Qufu City, Shandong Province, China (Diptera: 


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Yan, G., Y. Zhang and Y.-X. Yu. 1995. Five new species of the genus Atrichopogon from Tibet, China (Diptera: 

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– 1982. Chapter III. The identification of important biting midges of China [in Chinese]. in Manual of Important Chinese 

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– 1985. Description of Leptoconops (Holoconops) tamimensis sp. nov. (Diptera: Ceratopogonidae) [in Chinese, English 


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232

TABLE 1 

Published sources listing type material of Ceratopogonidae in a museum or geographic area (many other museums 

have websites listing their type specimens) 

Museum or Geographical Area Author 

British Museum (BMNH) Townsend and Boorman (1990); Townsend unpublished 

ms 

California Academy of Sciences (CASC) Arnaud (1979) 

Canadian National Collection (CNCI) Cooper (1991) 

University of Sao Paulo (DEFS) Forattini et al. (1971) 

Deutsches Entomologisches Institut (DEIC) Blech and Rohlfien (1987) 

Illinois Natural History Survey (INHS) Webb (1980) 

Musée Royal de l'Afrique Centrale, Belgium (MRAC) Kremer et al. (1975) 

Museo de La Palata, Argentina Marino, et al., 2002 

South African Institute for Medical Research (SAMR) Segerman (1995) 

Snow Entomological Museum (Kansas) (SEMC) Byers et al. (1962) 

Zoological Institute, St. Petersburg (ZMAS) Glukhova and Brodskaya (1995) 

Australasian Region Debenham (1979) 

Philippines Delfinado and Hardy (1971) 

233

TABLE 2 

Location of type material described by most ceratopogonid systematists. Museums listed hold most or all of 

the type specimens designated by a given author although many authors have described scattered species held by 

museums not listed here. All authors who have coauthored with Wirth or Yu have deposited all or most of their 

material in the depositories given below under these two authors and they are not listed separately. 

Author Museum 

Adams SEMC 

Amossova ZMAS 

Arnaud CASC; USNM 

Atchley USNM 

Austen BMNH 

Barbosa USNM 

Beck USNM 

Becker ZMHB 

Boesel PMNH; ROME 

Boorman BMNH 

Borkent CNCI, USNM, INBC 

Brèthes MNHN; MACN 

Browne USNM 

Bujanova ZMAS 

Callot IPSF 

Carter BMNH 

Causey USNM 

Cavalieri INMA? 

Chan LEMQ; CNCI; USNM 

Chaudhuri ELBU 

Chen NIPM 

Chu DPSC 

Clastrier MNHN 

Cochrane USNM 

Coquillett USNM 

Cornet IPSF; ORST 

Das Gupta NZSI 

Debenham ANIC 1 

Delécolle MZSF 

Delfinado USNM; FMNH 

de Meijere ZMAN 

de Meillon SAMR; NMSA; USNM 

Dessart MRAC 

Dow USNM; CNCI 

Downes CNCI 

Dzhafarov ZMAS 

Edwards BMNH 

Enderlein ZMHB 

Ewen CNCI 

Fabricius ZMUC 

Felippe-Bauer FIOC 

Floch IPCG 

Foote USNM 

Forattini MZSP 

Fox EPRL 

234

Garrett CNCI 

Giles USNM 

Gimmerthal SZRL 

Glick USNM 

Glukhova ZMAS 

Goeldi BMNH 

Goetghebuer ISNB; MRAC 

Gornostaeva ZMAS 

Grogan USNM; CNCI 

Gutsevich ZMAS 

Haeselbarth SAMR; USNM 

Hardy BPBM 

Havelka PEHC 

Heyden BMNH 

Hoffman NYSM; USNM 

Hogue LACM 

Howarth BPBM 

Hubert USNM 

Huttel FMMF 

Ingram BMNH 

Jamnback NYSM 

Johannsen CUIC 

Jones USNM 

Kalugina PIAM 

Kettle BMNH 

Khalaf SMSH; BMNH 

Khamala BMNH 

Kieffer HNHM; DEIC; BMNH; MNHN; 2 ; ZMSC (Thienemann material) 

Kitaoka NSMT 

Knab USNM 

Knausenberger USNM 

Kremer MNHN 

Krivosheina IEME 

Lane, J. DEFS; BMNH 

Lane, R. P. BMNH 

Lee, D. J. ANIC; SPUS 

Lee, T-S. IZAS 

Lewis USNM 

Lien NIPM 

Linnaeus contact BMNH; NHRS 

Loew MCZC; IMGP; ZMHB 

Lutz FIOC; BMNH (see discussion by Belkin et al., 1971) 

Macfie BMNH, DEIC 

Macquart MNHN 

Malloch INHS; SEMC 

Mayer DEIC; PEHC 

McDonald NIPM 

Meigen MNHN 

Meiswinkel VRIO 

Meunier IMGP 

Mirzaeva BIAR; ZMAS 

Molotova ZMAS 

Navai IPHT; USNM 

Neveu MNHN; personal collection 

235

Nielsen ZMUC 

Ortiz EPRL 

Pappas USNM 

Petrunkevitch ZMUC 

Philippi MEUC 

Pierce LACM 

Remm IZBE 

Roback ANSP 

Root USNM 

Rondani MZRF 

Ronderos MLPA 

Sahuquillo collection destroyed 5 

Santos Abreu MABC 

Saunders CNCI; USNM 

Séguy MNHN 

Sen NZSI 

Shannon INMA 

Shevchenko ZMAS 

Skuse ANIC 

Smatov ZMAS 

Smee SPUS 

Spinelli MLPA; USNM 

Staeger ZMUC 

Statz LACM 

Strobl ZMHB 

Storå UZMH 

Szadziewski RYSC; MZWP 

Thomsen CUIC 

Tokunaga BPBM; ELKU; USNM; CASC 

Tonnoir NZAC 

Townsend SEMC 

Vargas IDRE 

Vimmer NMPC 

Wada NSMT 

Walker BMNH 

Wiedemann NHMW; ZMUC 

Williams USNM 

Williston SEMC; BMNH 

Winnertz NHMW; SMFD; ZMHB; MZLU; some in Oxford 

Wirth (and coauthors) USNM; CASC; CNCI; BMNH; 3,4 

Yu (and coauthors) IMBC 

Zetterstedt MZLU 

1 Specimens originally recorded as deposited in SPHTM now in ANIC 

2 Many destroyed or lost 

3 Scattered specimens in many other museums. 

4 See Arnaud and Arnaud, 1997 for detailed list. 

5 Victor Sarto Monteys (pers. comm., March, 2012) 

236

World Species of Biting Midges (Diptera: Ceratopogonidae)

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