The beetles of decaying wood in Ireland - National Parks & Wildlife ...

The beetles of decaying wood in Ireland. 

A provisional annotated checklist of 

saproxylic Coleoptera. 

Irish Wildlife Manuals No. 65

The beetles of decaying wood in Ireland. A provisional 

annotated checklist of saproxylic Coleoptera. 

Keith N. A. Alexander 1 & Roy Anderson 2 

1 59 Sweetbrier Lane, Heavitree, Exeter EX1 3AQ; 2 1 Belvoirview Park, Belfast BT8 7BL, N. Ireland 

Citation: Alexander, K. N. A. & Anderson, R. (2012) The beetles of decaying wood in Ireland. A 

provisional annotated checklist of saproxylic Coleoptera. Irish Wildlife Manuals, No. 65. National 

Parks and Wildlife Service, Department of the Arts, Heritage and the Gaeltacht, Dublin, Ireland. 

Keywords: beetles; saproxylic; deadwood; timber; fungal decay; checklist 

Cover photo: The Rhinoceros Beetle, Sinodendron cylindricum © Roy Anderson 

The NPWS Project Officer for this report was: Dr Brian Nelson; brian.nelson@ahg.gov.ie 

Irish Wildlife Manuals Series Editors: F. Marnell & N. Kingston 

© National Parks and Wildlife Service 2012 

ISSN 1393 – 6670

Saproxylic beetles of Ireland 

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Contents 

Executive Summary........................................................................................................................................ 2 

Acknowledgements ........................................................................................................................................ 2 

Introduction..................................................................................................................................................... 3 

Aims and methods...................................................................................................................................... 3 

Analysis of status ........................................................................................................................................ 4 

Special features of the Irish saproxylic fauna.......................................................................................... 8 

Important sites............................................................................................................................................. 9 

The checklist of Irish saproxylic Coleoptera ............................................................................................. 16 

Structure of the checklist.......................................................................................................................... 16 

Species accounts........................................................................................................................................ 17 

References .................................................................................................................................................... 131 

Appendix 1 Recorders and list of abbreviations..................................................................................... 144 

Appendix II Taxonomic checklist of species ........................................................................................... 146 

Index of beetle species................................................................................................................................ 158


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Executive Summary 

A full list is provided of the saproxylic (wood-decay) beetles known to have been resident in Ireland 

in modern times, together with information about their ecology and distribution, with site records 

organised by vice county. 

For each of the 274 species an estimation of their current status is suggested, and species with an 

apparent association with old growth (ancient woodland and/or ancient wood pasture conditions) are 

also identified. At least 208 species appear to be native to Ireland, while 9 of these may have become 

extinct within the past 100 years. Only 25 species appear to be common and widespread, in contrast to 

the 104 species (50% of the natives) which have been assessed as ‘rare’. This suggests a serious 

imbalance in species status, reflecting the overall rarity and fragility of saproxylic habitats across 

Ireland. A list of important sites is included. 

Acknowledgements 

Brian Nelson arranged for this independent report to be published by NPWS. The authors would like 

to thank the recorders who have generously made their records available to us: Martin Cawley, Tom 

Gittings and Jervis Good. Also our thanks go to Eugenie Regan of the National Biodiversity Data 

Centre for encouragement to move towards publication.


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Introduction 

Saproxylic and other woodland invertebrates have a reputation for being very poorly represented in 

Ireland (Speight 1986, 1989b) although Alexander (2003) suggests that this merely reflects the overall 

species representation in Ireland in comparison with Britain and continental Europe – a factor of 

Ireland’s biogeographic relationships – rather than saproxylics being rarer than any other habitat 

grouping. However, what may be true is that the background fauna – the common and widespread 

species which can be found very widely across Ireland – is a relatively small range of species, while 

the more restricted species form a greater proportion of the fauna than might be expected. Species 

richness is certainly greatest in sites which are almost certainly ancient woodlands and ancient wood- 

pastures (including historic demesne parklands), and it is suggested that the species which are 

virtually exclusive to these relatively rich sites are the remains of Ireland’s old growth fauna 

(Alexander et al., 2002). Ancient tree-scapes do appear to be relatively scarce across Ireland, although 

probably not as scarce as many people assume – the historic demesnes have hardly been studied at all. 

The definition of saproxylic used for the purposes of this list is the one proposed by Alexander (2008b) 

as it emphasises fungal decay of wood and does not require trees to be senescent or dead (Speight, 

1989c): Species which are involved in or dependent on the process of fungal decay of wood, or on the 

products of that decay, and which are associated with living as well as dead trees. Conventionally two 

further groupings of organisms are included within the definition: i) sap-run associates, ie species 

dependent on fluxes of sap and its decomposition products, and ii) organisms other than fungi that 

feed directly on wood. 

This checklist is presented in order to provide a complete statement of where we are with knowledge 

of the current Irish saproxylic beetle fauna. Subfossil records have not been included. It is hoped that 

this baseline data will provide a framework for developing an Irish Red List of saproxylic beetles and 

enable the development of site quality statistics along the lines of those already in use in Britain 

(Alexander, 2004; Fowles et al., 1999). 

Aims and methods 

This checklist aims to include all resident breeding species, and so includes the long-established native 

fauna plus introductions which have proved able to maintain viable populations in Ireland. Casual 

imports are omitted where their status is clear-cut, e.g. Silvanus bidentatus in pallets associated with a 

consignment of computer parts from Japan (Good & Hume, 1991), but recent arrivals are included as 

these may become established. 

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The data is primarily derived by extraction from published literature and contact with the active 

recorders that are known to the authors. The record is almost certainly not complete however, as 

unpublished information will largely remain uncollated for the time being. 

Analysis of status 

274 species are listed of which at least 208 are native – see Table 1. Nine of these species may now be 

extinct, having not been reported since Johnson & Halbert (1902) – see below. This is clearly a rich 

fauna, despite the various opinions which have been expressed over the years – see Introduction. 

However, only 25 species can be considered common and widespread, in contrast to the 104 (50%) 

species which have been assessed as rare in Ireland. This suggests a serious imbalance in species 

status, reflecting the overall rarity and fragility of saproxylic habitats across Ireland. 

Eight additional species are thought to be doubtfully Irish as only old records have been reported and 

no supporting voucher specimens are available, while the status of a further 8 have proved impossible 

to assess on the scant information available. 35 species are established non-natives, largely 

accidentally brought in by commercial forestry activities. There is also a clear trend of recent arrivals 

making first landfall in Northern Ireland, presumably carried from Scotland on air currents – 8 species 

have first appeared in the north-eastern counties and have proceeded to spread. 

The species presumed to be extinct in Ireland are: Nossidium pilosellum (Marsham), Amphicyllis globus 

(F.), Agathidium rotundatum Gyllenhal, Euplectus punctatus Mulsant, Cypha seminulum (Erichson), 

Calambus bipustulatus (L.), Malthodes dispar (Germar), Anthocomus fasciatus (L.) and Conopalpus testaceus 

(Olivier). These represent a wide variety of saproxylic habitat types. 

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Figure 1: The distinctive larva of the Red-headed Cardinal Beetle Pyrochroa serraticornis © Keith Alexander


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Figure 2: Larvae of Sinodendron cylindricum bore in decaying heartwood of broad-leaved trees © Roy Anderson 

Figure 3: Two wood-decay fungi Ganoderma resinaceum (left) and Inonotus dryadeus (right) © Roy Anderson 

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TABLE 1 The Irish status of saproxylic beetle species grouped by family (and subfamily for selected 

groups). (For heading codes see ‘Structure of Checklist’ below.) 

Family/subfamily All Ex R Sc Un Wi Cm Est Ca Re Do Ucl 

Histeridae 1 1 

Ptiliidae 9 1 5 3 

Leiodidae 8 2 3 2 1 

Scydmaenidae 1 1 

Staphylinidae 

• Omaliinae 

• Pselaphinae 

• Phloeocharinae 

• Tachyporinae 

• Aleocharinae 

• Scaphidiinae 

• Piestinae 

• Staphylininae 

12 6 4 2 

7 1 6 

1 1 

1 1 

32 1 15 2 7 4 1 1 1 

2 1 1 

1 1 

6 4 1 1 

Lucanidae 1 1 

Clambidae 1 1 

Scirtidae 1 1 

Eucnemidae 1 1 

Elateridae 6 1 1 1 2 1 

Lycidae 1 1 

Cantharidae 10 1 1 5 1 1 1 

Bostrichidae 1 1 

Anobiidae 11 4 2 1 1 3 

Lymexylidae 1 1 

Phloiophilidae 1 1 

Trogosittidae 1 1 

Cleridae 2 1 1


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Family/subfamily All Ex R Sc Un Wi Cm Est Ca Re Do Ucl 

Dasytidae 1 1 

Malachiidae 2 1 1 

Aspidiphoridae 1 1 

Nitidulidae 19 6 1 4 2 2 2 1 1 

Monotomidae 7 1 2 1 2 1 

Laemophloeidae 1 1 

Cryptophagidae 5 3 1 1 

• Atomariinae 

2 1 1 

Erotylidae 2 1 1 

Cerylonidae 3 1 2 

Endomychidae 1 1 

Corylophidae 1 1 

Latridiidae 1 1 

Mycetophagidae 2 2 

Ciidae 12 4 2 1 1 3 1 

Tetratomidae 3 2 1 

Melandryidae 6 1 4 1 

Mordellidae 1 1 

Colydiidae 3 2 1 

Tenebrionidae 4 1 1 2 

Oedemeridae 2 1 1 

Pyrochroidae 1 1 

Salpingidae 9 5 1 1 1 1 

Scraptiidae 7 2 1 2 2 

Cerambycidae 22 9 1 2 2 2 2 2 1 1 

Anthribidae 1 1 

Curculionidae 17 4 2 1 1 5 2 2 

• Scolytinae 

19 1 2 1 2 11 1 1 

TOTALS 274 9 104 27 29 13 25 35 6 8 7 8 

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About 50 species appear to have an association with ancient woodland and ancient wood pasture 

conditions (old growth species) – nearly 25% of the natives. This emphasises the great importance of 

these habitats for saproxylic beetles, and also provides an opportunity for developing conservation 

quality statistics for site evaluation and comparison. 

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Figure 4: Caledon Deer Park, Co. Tyrone © Keith Alexander 

Special features of the Irish saproxylic fauna 

Ireland is the global epicentre for at least one species of saproxylic beetle. The Holly Weevil 

Rhopalomesites tardyi was discovered new to science and named after James Tardy a prominent Dublin 

entomologist of the early 19 th century – as Cossonus tardii – and taken by him and N.A.Vigors in July, 

1822, near Powerscourt waterfall, Co. Wicklow, under the bark of decayed hollies. Strangely, Tardy is 

reported as saying “I have in vain sought for it in places abounding as much in holly and in similar 

situations in the same country” (Nash, 1983). Today it is one of the most widespread species to be 

found, occurring in hedgerow and boundary stands of trees as well as old woodlands. Interestingly it 

has subsequently been found in Britain, although it is very localised and concentrated along the 

western seaboard. Its global range is restricted to offshore islands and extreme western coasts of


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Europe from the British Isles to the Azores, taking in coastal north-west France and southern Portugal 

(eg in driftwood at Rio Arade, Portugal, 24.vi.1994, RA). 

The presence of the red click beetle Ampedus pomonae in Ireland but not Britain is another interesting 

feature of the Irish saproxylic beetle fauna. However, it is widespread on the near Continent and its 

absence from Britain is the curiosity rather than its presence in Ireland. 

A third species is also of note, Quedius lyszkowskii, as it is only known globally from Ireland and 

Scotland. It was described new to science only recently (Lott 2010), having previously been 

overlooked within Q. aetolicus by modern workers and historically with Q. puncticollis in Irish 

collections. Its earliest known record is from 1905 and so it seems reasonable to assume that it is long- 

established native – at least until evidence to the contrary is forthcoming. 

Important sites 

As already stated, the saproxylic beetle fauna of Ireland is not well-studied and undoubtedly 

important sites remain overlooked. The situation has been made worse by the spurious notion that 

rare and threatened species are primarily associated with semi-natural vegetation and therefore the 

parklands of historic demesnes would not be expected to be of special interest. In reality many of the 

historic demesnes actually protected and maintained more features of earlier Irish landscapes than did 

the average area of countryside. Saproxylic interest is focused on trees rather than woodlands, and 

sites which retain relatively large concentrations of veteran trees over time support the richest variety 

of saproxylic beetles. Such sites tend to be the historic demesnes rather than the ancient woodlands – 

themselves relatively rare in Ireland anyway. Most historic demesnes have not been assessed for their 

saproxylic interests – the compilation of an inventory of such sites and an investigation into the fauna 

present should be a high priority for invertebrate conservation in Ireland. Recent work in Northern 

Ireland has demonstrated the great value of historic parklands for Ireland’s wood-decay invertebrates 

(Alexander et al., 2007). 

The sites are listed in order of known importance for saproxylic beetles. Only eight areas are currently 

known to support more than one species of great rarity: 

Killarney National Park area, Co. Kerry 

This is the outstanding site historically, very well studied up to the early 20 th century, notably by 

Edwin Bullock, but very neglected in recent decades. There is an urgent need for a re-assessment in 

view of the management changes that have been taking place. More than 20 notable species – 

considerably more than any other area that has been studied; the only known Irish site for the Six- 

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spotted Longhorn Beetle Anoplodera sexguttata (1859 & 1879); one of only two known sites for the Fire- 

winged Beetle Pyropterus nigroruber (discovered in 1866 & still there in 1989). The majority of the 

records stand merely as ‘Killarney’ making it impossible to allocate particular records to particular 

locations within the area. Derrycunihy and Tomies Woods are well-known to be good examples of the 

habitat, but equally the open parkland areas are also notably important for saproxylic beetles, 

especially Muckross Park. The whole area forms a mosaic of ancient woodland, ancient wood pasture 

and historic parkland, which in turn forms the largest extent of veteran tree habitat anywhere in 

Ireland. Greater attention to its saproxylic beetle importance is needed as this will greatly inform site 

management for conservation objectives. 

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Figure 5: Glena Wood, Killarney National Park, Co.. Kerry © John Cross 

Powerscourt Deer Park and area, Co. Wicklow 

A very famous site known to the early recorders; the only known Irish site for the black and red 

flower beetle Ischnomera sanguinicollis (still there in 1985); one of very few known sites for the fungus 

beetle Anisotoma orbicularis (1927) and the false darkling beetle Melandrya caraboides (1824).


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Glengarriff Woods, Co. Cork 

Only known Irish site for Mordellistena neuwaldeggiana (1924); one of only two known sites for the false 

click beetle Melasis buprestoides (1985); best Irish site for Hornet Longhorn Beetle Leptura aurulenta; one 

of very few known sites for the ambrosia beetle Hylecoetus dermestoides. Glengarriff Castle Demesne is 

of especial note for its veteran oaks; it was here that the ambrosia beetle was found and this is also a 

good area for the longhorn beetle. 

Glencar area, Co. Kerry 

Figure 6: Glengarriff, Co. Cork © Keith Alexander 

Long famous as the only site in Ireland for the red click beetle Ampedus pomonae (discovered here in 

1902 and still there, but now also known from Co. Clare); also the first area where the false click beetle 

Melasis buprestoides was found (1924); also one of only two known sites for the narrow-waisted beetle 

Lissodema denticolle (1906 & 1924); the first place that the ambrosia beetle Hylecoetus dermestoides was 

found in Ireland (1902). 

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Crom Castle Demesne, Co. Fermanagh 

Accessible for recording only since 1987 and with a steady increase in notable finds: first Irish site for 

the false darkling beetle Abdera flexuosa, the fungus beetle Agathidium confusum and the bracket fungus 

beetle Dorcatoma dresdensis; one of few modern localities for the fungus beetle Anisotoma orbicularis and 

the short-winged mould beetle Bibloporus bicolor. An important and unusually large concentration of 

old parkland, as well as having areas of more typical ancient wood pasture and ancient woodland. 

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Figure 7: Veteran trees on Inisherk, Crom Castle Estate, Co. Fermanagh © Keith Alexander 

Charleville Park and Woods, Co. Offaly 

Also only recorded in recent years; first Irish record for the longhorn beetle Grammoptera ustulata 

(2003) and one of few Irish sites for the minute fungus beetle Cis villosulus (2003). An important area 

for the oak high forest habitat as well as remnant parkland.


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Belvoir Park, Co. Down 

Figure 8: A mature tree and regeneration at Charleville, Co. Offaly © John Cross 

Has received a lot of attention in recent decades: the only Irish site for the old forest spider beetle 

Ptinus subpilosus (1992 onwards) and also with one of the Killarney rarities, the rove beetle Atheta 

pilicornis. This site contains many old parkland trees especially oak – of 130 native oaks with a girth of 

3m or more, 55 have girths in the range of 4-6m (Simon, 2005). 

St John’s Wood, Co. Roscommon 

First investigated in any detail in 2010 (Alexander 2011); notably, one of few Irish sites for the short- 

winged mould beetle Euplectus piceus and the stone beetle Stenichnus bicolor; first known site in the 

Republic for the bracket fungus beetle Dorcatoma dresdensis. This wood is said to be the largest and 

most natural woodland in the Irish midlands. Its coppice-with-standards structure means that it 

contains a large number of veteran trees, particularly oaks, as well as old hazels and alders. 

Other sites, with at least one great rarity: 

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The Great Deer Park, Glenarm, Co. Antrim – one of only two known sites for the fungus beetle 

Agathidium confusum. This site has a notably large extent of semi-natural ancient wood pasture and is 

an outstanding site in the context of Northern Ireland (Alexander et al., 2007). 

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Figure 9: The Great Deer Park, Glenarm, Co. Antrim © Keith Alexander 

Slish Wood, Co. Sligo – one of only two sites for the Fire-winged Beetle Pyropterus nigroruber. 

Remnant ancient oak woodland. 

Shane’s Castle Park, Co. Antrim – an important modern site for another Killarney rarity, the sap 

beetle Epuraea terminalis. Parkland and broad-leaved woodland. 

Castle Coole Park, Co. Fermanagh – one of the few modern sites for the rove beetle Hapalaraea 

pygmaea; a large expanse of historic parkland with many veteran oak, etc. 

Tynan Abbey Park, Co. Tyrone – one of the few modern sites for the rove beetle Hapalaraea pygmaea; a 

large expanse of historic parkland with many veteran oak, etc; more or less continuous with Caledon 

Deer Park which is another important area of historic parkland.


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It is very obvious that historic parklands and ancient wood pastures feature very strongly in this short 

list, while conventional ancient woodland is relatively poorly represented. However, the detailed 

survey of St John’s Wood (Alexander, 2011) has demonstrated that ancient woodlands may also be 

rich in saproxylic invertebrates. The overwhelming majority of ancient woodland sites listed by Perrin 

and Daly (2010) have not yet been assessed for their saproxylic beetle interest. 

Figure 10: A sap run, an important microhabitat for woodland insects © Keith Alexander 

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The checklist of Irish saproxylic Coleoptera 

Structure of the checklist 

The checklist of species considered saproxylic is given in Appendix II. The listing in this and the 

species accound section below follows the current taxonomic checklist order (Duff, 2008), with species 

name, proposed Irish status (see below), and a short account of its ecology. The records follow and are 

ordered by vice county, with locality name, grid reference (where readily available), recorders name, 

date of record, and source. The sequence is in date order, the earliest record first. Where a historic 

record is ambiguous to county then the record is placed in the most likely vice county. 

The Irish status in bold is provided as a starting point, based on existing evidence, to stimulate 

thought and discussion. This is very likely to be modified as knowledge progresses. The expressions 

used are as follows: 

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Extinct (Ex) – no records since about 1900 

Rare (R) – only a few modern records 

Scarce (Sc) – known from only about ten modern localities 

Uncommon (Un) – known from many counties, some with many records 

Widespread (Wi) – known from most counties and plenty of records; only the year of records 

are provided for the most widely recorded species 

Common & Widespread (Cm) – only VC distribution provided 

Established introduction (Est) – species believed to have been brought into Ireland 

incidentally through commerce; a few have first been found only very recently 

Casual Introduction (Ca) – an occasional record only, no established populations known 

Recent arrival (Re) – believed to have arrived naturally 

Doubtful (Do) – only a single report but no evidence of a resident population 

Unclear (Ucl) – too little information for meaningful assessment of status 

The recorder for all individual records is given, usually as an acronym to save space (see Appendix 1), 

followed by any literature references (usually bracketed). All records employed in this review (and


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many paraphrased as vice-county records for common species) have been uploaded to the Recorder 

database held by CEDar LRC, Ulster Museum and to the National Biodiversity Data Centre (NBDC), 

Waterford. 

A full list of references is appended. 

Species accounts 

FAMILY HISTERIDAE Hister Beetles 

Predatory, especially on larvae of other deadwood inhabiting insects, also on mites and springtails. 

The following species is the only Irish one found associated with wood decay, but others are 

occasionally found in this situation although more strongly associated with other habitat types. 

Abraeus perpusillus (Marsham, 1812) syn. globosus (Hoffman, J., 1803) – Rare 

In moist rotten wood of various broad-leaved trees; usually within ancient woodland, wood pasture, 

or historic parkland in Britain. A curious distribution in Ireland – in the central eastern counties, but 

perhaps favouring the relatively warmer and drier districts. 

H19 Kildare: Louisa Bridge, Royal Canal, in Polyporus squamosus on poplar, viii.1942, AWS 

(O’Mahony, 1943b). 

H21 Dublin: St. Annes Park (O2037), rot hole in beech, 19.v.1994, JAO (Owen, 1997). 

H38 Down: Hollymount NNR, under bark on beech by Ganoderma bracket, 25.iii.2012, RA. 

FAMILY PTILIIDAE Feather-winged Beetles 

Mould-feeders, living between the bark and sapwood of dead trees or in rot-holes, where conditions 

are slightly moist and mouldy. 

Nossidium pilosellum (Marsham, 1802) – Extinct 

Develops in decaying wood, particularly rotten stumps and damp, rotten, fungus-colonised timber, 

and especially elm and beech in Britain. Ireland: ‘very rare, on rotten stumps (Haliday, 1855)’ (Johnson 

& Halbert, 1902) – no details provided. 

Ptenidium laevigatum Erichson, 1845 – Rare 

Possibly not a true saproxylic. 

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H2 North Kerry: Kenmare Demesne (V99), a few specimens taken together in a damp, decaying tree 

stump, between 18 & 25 July 1913, OEJ (Janson, 1914). 

Ptinella aptera (Guérin-Méneville, 1839) – Rare old growth species 

Under bark of decaying oak and beech wood. 

H2 North Kerry: Killarney, EFB (Bullock, 1928). 

Ptinella cavelli (Broun, 1893) – Established introduction 

Under tight bark of dead broad-leaves and conifers; native to New Zealand, widespread in Britain and 

the north of Ireland where first found in 1975 (Nash, Anderson & O’Connor, 1997). 

Ptinella denticollis (Fairmaire, 1858) – Rare old growth species 

Under tight bark of dead broad-leaved timber. Only a single record reported. 

H2 North Kerry: Killarney, EFB (O’Mahony, 1929a). 

Ptinella errabunda Johnson, 1975 – Established introduction 

Under tight bark of most species of dead trees; probably native to New Zealand. Found abundantly by 

RA in decaying wood in several parkland sites west of Belfast (Nash, Anderson & O’Connor, 1997). 

H38 Down: Lagan Meadows, under bark of Scots pine, 27.xi.1992, RA. 

H39 Antrim: Newforge, R. Lagan, under bark of rotting Japanese larch with fruiting decay fungus 

Coniophora puteana, 20.x.1992, RA (Nash, Anderson & O’Connor, 1997); Ballyskeagh, Belfast, under 

beech bark, 8.xii.1992, RA. 

Ptinella limbata (Heer, 1841) – Rare old growth species 

Under bark of various dead broad-leaved trees and conifers; old forest areas. This record may possibly 

be an early observation of the tiny yellowish Antipodean species P. errabunda which RA finds 

occasionally under pine bark in Northern Ireland but which is otherwise unreported from the south. 

H21 Dublin: Tolka Valley, under bark on pine logs, JNH (Halbert, 1924). 

Ptinella taylorae Johnson, 1977 – Established introduction 

Under tight bark of dead trees; native to New Zealand; not uncommon in W. England & Ireland 

(Johnson 1977). 

H39 Antrim: Rea’sWood, Antrim, under alder bark, 21.x.2006, RA. 

Pteryx suturalis (Heer, 1841) – Rare 

Under bark and in rotten wood of dead broad-leaved trees, rarely in conifer. 

H2 North Kerry: near Killarney, EFB (Bullock, 1928) (O’Mahony, 1929a). 

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H39 Antrim: Rea’s Wood, Antrim, 17.iv.1974, AGI, det CJ. 

FAMILY LEIODIDAE 

All species of Anisotoma have an obligate association with slime fungi, with adults and larvae feeding 

on the spores. Species of Agathidium are most likely primarily associated with slime fungi but the 

evidence is less clear – not all species appear to be associated with decaying wood. The species appear 

typical of shady woodlands rather than more open situations. 

Anisotoma humeralis (Fabricius, 1792) – Uncommon old growth species 

Modern sites almost invariably ancient woodlands and ancient wood pastures. 

H1 South Kerry: Kenmare, JRH (in Halbert, 1898c) (Johnson & Halbert, 1902); Rossbehy, 1902, HSJD 

(Donisthorpe, 1902). 

H2 North Kerry: nr. Killarney, 1910-1914, EFB; Ross Castle, Killarney, 22.v.1995, PJH. 

H16 West Galway: Connemara National Park (L75), 1994, MCDS (Speight, 2004). 

H18 Offaly: Charleville Wood (N315226), v. 2003, RA (Regan & Anderson, 2004) 

H20 Wicklow: (Halbert, 1898c, in A.H. Haliday collection, Dublin Museum); Powerscourt Deerpark 

(O1912), swept along R. Dargle, 9.vi.1919, JNH (Halbert, 1922); 19.vi.1993, KNAA; Clara, iv.1929, AWS 

(O’Mahony, 1929b); Devil’s Glen, 9.vi.1934, EO (O’Mahony, 1934). 

H23 Westmeath: Lough Derravaragh, JNH (Johnson & Halbert, 1902). 

H25 Roscommon: St John’s Wood (M9956 & N0055), 6.viii. & 8.x.2010, KNAA. 

H29 Leitrim: Acres Lake (G966100), two knocked from dead Pinus, 23.iv.2011, MC. 

H30 Cavan: Cloverhill, vi.1916, GWN (Nicholson, 1917). 

H33 Fermanagh: Reilly Wood, Crom (H32), 28.v.1992, KNAA; Inisherk, Crom, FIT, 1.vii.1992- 

7.vii.1992, APF; Florencecourt, 24.viii.2010, RA. 

H36 Tyrone: Brookend NR, Lough Neagh, 8.v.2007, RA. 

H38 Down: Stormont, Belfast, 17.ix.1994, RA; Tollymore Forest Park, 27.vi.1998, RA. 

H39 Antrim: Shane’s Castle, vi.1902, CWB (Halbert, 1910b); 1.vii.2010, RA; Great Deer Park, Glenarm 

(D2911), 20.v.2006, KNAA. 

H40 Londonderry: Banagher Glen (C6704), FIT, 8.vii.2007, KNAA. 

Anisotoma orbicularis (Herbst, 1792) – Rare old growth species 

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Only known from two of Ireland’s richest areas for saproxylics. 

H20 Wicklow: Powerscourt, 15.viii.1927, AWS (O’Mahony, 1927). 

H33 Fermanagh: Inisherk, Crom, 1.vi.1992, 7.vii.1992, 5.viii.1992, 4.ix.1992; Inishfendra, Crom, 15- 

16.vii.1993, FIT samples, APF. 

Amphicyllis globus (Fabricius, 1792) – Extinct 

In decaying timber, mostly in ancient woodland areas in Britain. No modern reports. 

H11 Kilkenny: Thomastown, one swept “in a marshy place on the banks of the Nore, July, 1901", JNH 

(Johnson & Halbert, 1902). 

Agathidium confusum Brisout, 1863 – Rare old growth species 

First noted in Ireland in 1993 but regarded as an overlooked native. 

H33 Fermanagh: Inishfendra Island, Crom, FIT, 16.vi.1993 – 15.vii.1993, APF. 

H39 Antrim: Great Deer Park, Glenarm (D2911), in numbers from the twigs of a fallen large oak 

branch, 20.v.2006, KNAA (Alexander, 2008a). 

Agathidium nigripenne (Fabricius, 1792) – Scarce 

Only reported across the more northern counties; not northern in Britain. 

H15 South-east Galway: Coole Park (M433043), several swept, 22.v.2010, PJH. 

H25 Roscommon: St John’s Wood (M9956), various dates in 2010, KNAA. 

H27 West Mayo: Westport, JJW (Fowler, 1891). 

H33 Fermanagh: Inisherk Island, Crom, FIT, 1.vi.1992 – 7.vii. 1992, APF. 

H34 East Donegal: Rathmullan, JJW (Fowler, 1891) (Johnson & Halbert, 1902); Foyle district, 1900, 

CWB. 

H37 Armagh: Armagh, WFJ (Johnson & Halbert, 1902); Annagarriff NR, Peatlands Park, 29.v.2000, RA 

(Anderson, Simms & Nelson, 2000). 

H38 Down: Stormont, 7.v.1994, 13.v.1994, RA; Belvoir Forest, 23.iv.1998, RA; Hillsborough, 9.viii.2000, 

RA; Shooter’s Island, Quoile (J494470), 23.ii.2012, MDB. 

H39 Antrim: Farr’s Bay, Randalstown, 4.vii.1986, RCW (Welch, 1986); Belfast Castle, 11.vii.2004, 

9.iii.2008, RA. 

H40 Londonderry: Ness Wood (C528118), 19.v.2007, KNAA. 

Agathidium rotundatum (Gyllenhal, 1827) – Extinct 

20


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H33 Fermanagh: Belleisle (Porter, 1898) (Johnson & Halbert, 1902). 

H37 Armagh: Palace Demesne, in moss, WFJ (Johnson, 1892a); Coney Island, Lough Neagh, WFJ 


H39 Antrim: Murlough, GWC, det. JNH (Johnson & Halbert, 1902). 

Agathidium seminulum (Linnaeus, 1758) – Rare 

H2 North Kerry: near Killarney, pre-1914, EFB (Bullock, 1914). 

H33 Fermanagh: Munnilly Bridge, Old Ulster Canal, 19.vii.2011, RA. 

H39 Antrim: Cranmore Park, Belfast, 18.iv.2005, RA. 

Agathidium varians Beck, 1817 – Scarce 

H20 Dublin: The Dingle (Waterstown Park), v. 1935, JNH (1937). 

H33 Fermanagh: Inisherk, Crom, FIT, 1.vi.1992 – 7.vii.1992, APF; Castle Coole Park (H2643), FIT, viii- 

14.ix.2007, KNAA. 

H38 Down: Belvoir Forest, Belfast, 24.ii.1999, RA. 

H39 Antrim: Shane’s Castle, CWB, 1902, det. JNH (Johnson & Halbert, 1902: Suppl. List); Rea’s Wood, 

Antrim, 3.xii.1994, RA; Great Deer Park, Glenarm (D2911), FIT, vii-19.viii.2007, KNAA. 

H40 Londonderry: Drenagh Estate (C6923), FIT, viii-15.ix.2007, KNAA. 

FAMILY SCYDMAENIDAE Stone Beetles 

Predatory on mites, in moist situations; a few frequent decaying wood. 

Stenichnus bicolor (Denny, 1825) – Rare old growth species 

Under bark and in moist crumbly decaying timber of various dead trees; widespread in moister areas 

of Britain; mostly in ancient wood pastures. 

H2 North Kerry: near Killarney, pre 1914, EFB, det GCC. 

H3 West Cork: Glandore (W23), one under a stone close to the shore, JNH, det GCC (Johnson & 

Halbert, 1902). 

H25 Roscommon: St John’s Wood (N0055), FIT, v-7.vi.2010, KNAA. 

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FAMILY STAPHYLINIDAE Rove Beetles 

A small number are restricted to decaying wood, but many are associated with fungal fruiting bodies 

and the degree of association with wood-decay fungi is often unclear. 

STAPHYLINIDAE SUBFAMILY OMALIINAE 

Phyllodrepoidea crenata (Ganglbauer, 1895) – Rare old growth species 

Adults and larvae occur under bark of relatively freshly dead trees or branches, usually broad-leaved 

trees in England, pines in the relict pine forest of Scotland; they feed on fungal decay. A speciality of 

the hill country of the N and W of Britain, presumably also submontane in Ireland. Hammond (1980) 

refers to the Killarney population and the date of the first report but does not comment on whether or 

not there have been any subsequent records. 

H2 North Kerry: Killarney district, “fairly common in decaying fungi on trees in woods throughout 

the whole district, occurring frequently in the late Autumn months”, Bullock (1928); Deliphrum 

crenatum in O’Mahony (1929a). 

Coryphium angusticolle Stephens, 1834 – Rare old growth species 

Lives beneath bark of dead oak and other trees, and in red-rotten oak, mostly in wooded areas, 

possibly only ancient sites; widespread in Britain. 

H2 North Kerry: Killarney (Bullock 1914). 

H37 Armagh: Derryadd Lough, Peatlands Park, 16.iii.2002, RA. 

H38 Down: Holywood, sea coast (Haliday, 1841, 1855; in a rotten stump, Haliday ms.) (Johnson & 

Halbert, 1902); Belvoir Forest, Belfast, 12.iv.1995 & 17.iii.1998, RA. 

H40 Londonderry: Culmore, CWB (Buckle, 1900) (Johnson & Halbert, 1902). 

Acrulia inflata (Gyllenhal, 1813) – Rare old growth species 

Adults and larvae under fungoid bark on sound timber of various dead broad-leaved trees, where 

moist interface; general scavengers, feeding on insect material; adults also feed on fungal spores. In 

Britain confined to hill country of N and W; similarly submontane in Ireland. 

H2 North Kerry: Killarney, EFB (Bullock 1914), ‘the single Irish record’ (Hammond, 1980). 

H40 Londonderry: Banagher Glen, a pair among fruiting bodies of fungus Hymenoscyphus calyculus on 

oak branch, 6.xi.1999, RA. 

22


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Genus Dropephylla 

Adults and larvae are found under the bark of dead wood, although the adults of some (perhaps all) 

are attracted to blossom in early summer. Adults and larvae are cannibalistic and feed on small insects 

or other arthropods living under bark. 

Dropephylla devillei Bernhauer, 1902 syn. grandiloqua Luze, 1910 – Scarce old growth species 

Under bark of broadleaves and conifers in old forest areas in Britain; widespread but rare in Europe as 

a whole. Hammond (1980) regards the species as having spread into England recently from Scotland – 

the evidence for such mobility does not bear close scrutiny however. Most Irish records are from the 

classic saproxylic sites. 

H18 Offaly: Charleville Wood (N315226), v. 2003 (Regan & Anderson, 2004) 

H19 Kildare: Curragh Camp (H774118), Scots pine bark, 17.iii.2012, RA. 

H33 Fermanagh: Castle Coole Park, 17.viii.2006, KNAA. 

H36 Tyrone: Baronscourt Park, 15.vi. & 12.ix.2006, KNAA. 

H37 Armagh: Peatlands Country Park, 17.iii.1993, 23.x.1993, RA; Slieve Gullion Forest (J039197), 

conifer bark, 8.iii.2012, RA. 

H39 Antrim: Cave Hill, in numbers under bark of Scots pine, RA (Hammond, 1980); Great Deer Park, 

Glenarm, 13.vi., 19.viii. & 16.ix.2006, KNAA. 

Dropephylla gracilicornis (Fairmaire & Laboulbène, 1856) – Scarce 

Under bark and in rotten wood of dead branches of broad-leaved trees, especially oak. 

H2 North Kerry: Killarney (Bullock, 1930). 

H9 Clare: specimens seen (Hammond, 1980). 

H21 Dublin: Lucan Demesne, under bark, coll. JNH, det. D. Sharp (Johnson & Halbert, 1902: 

“Omalium brevicorne” probably refers to this species). 

H36 Tyrone: Baronscourt Park, 15.vi.2006, KNAA. 

H38 Down: specimens seen (Hammond, 1980); Donard Park, Newcastle, 31.xii.2011, RA. 

H39 Antrim: Lady Dixon Park, Belfast, 21.v.1980, RA; Great Deer Park, Glenarm, 19.viii.2006, KNAA. 

Dropephylla ioptera (Stephens, 1832) – Scarce 

Occasional under bark of dead branchwood; broad-leaved trees; adults visit flowers of rowan, etc; 

widespread in Britain, “probably very local” in Ireland (Hammond, 1980). 

H2 North Kerry: near Killarney, pre 1914, EFB (Bullock, 1914); Ballyseedy Wood, xi.2001, JAG. 

23


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H18 Offaly: Charleville Wood (N315226), v. 2003, RA (Regan & Anderson, 2004). 

H19 Kildare: Liffey bank at Celbridge, AWS. 

H22 Meath: Balrath, ix.1910, GWN (Nicholson, 1911). 

H25 Roscommon: Mote Park (Halbert, 1898a, as Omalium iopterum). 

H33 Fermanagh: Castle Coole Park, 17.viii.2006, KNAA; Garrison, Lough Melvin, 30.v.2007, DAL; 

Knockninny Hill, 1.vi.2007, RA; Filbert Hill, Castle Coole, 15.viii.2010, RA; Old Crom Castle 

(H364239), 24.v.2012, MDB. 

H36 Tyrone: Baronscourt Park, 24.x.2006, KNAA; Wood Lough (H759602), 24.v.2012, RA. 

H37 Armagh: Peatlands Park, 29.v.2000, RA. 

H38 Down: Hillsborough Forest, 20.vi.1995, RA; Morelands Meadow, Belvoir, 22.v.2004, RA. Donard 

Park, 31.xii.2011, RA. 

H39 Antrim: Lady Dixon Park, 21.v.1980, RA; Barnett’s Park, 19.v.1994, RA; Dungonnell dam, 

21.vi.1995, RA; Great Deer Park, Glenarm, 22.v.2007, KNAA. 

H40 Londonderry: Ness Wood, 19.v.2007, KNAA. 

Dropephylla koltzei Jászay & Hlavac, 2006 nec vilis (Erichson, 1840) – Scarce 

Under bark of dead broad-leaved trees and conifers; widespread in Britain, “probably local” in Ireland 

(Hammond, 1980). 

H22 Meath: Balrath, 26.ix.1913, GWN (Nicholson, 1914). 

H30 Cavan: Cloverhill, ix.1913, GWN (Nicholson, 1914). 

H33 Fermanagh: Trory, Lower Lough Erne (H225483), under bark on dead sallow, 23.v.1995, KNAA. 

H36 Tyrone: Ballybriest, Lough Fea, 5.vi.1994, RA. 

H37 Armagh: Moyrourkan Lough, 18.x.1980, RA; Derryadd Lough (H914604), 11.v.2012, RA. 

H38 Down: Donard Demesne, Newcastle, under bark of felled beech, vi.1923, OEJ (Janson, 1924); 

Leitrim Lodge, Mournes, 1.iii.1980, RA; Morelands Meadow, Belvoir, 15.iv.2004, RA. 

H39 Antrim: Carr’s Glen, CWB (Johnson & Halbert, 1902); Belfast Castle, 26.i.1980, RA; Barnett’s Park, 

19.v.1994, RA. 

Hapalaraea pygmaea (Paykull, 1800) – Rare old growth species 

Largely confined to areas of mature woodland, where it is found in bracket fungi, bird nests and 

squirrel dreys in the tree canopy, rotten wood, etc.; probably not an obligate wood-decay species. 

24


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H2 North Kerry: Kenmare Demesne (V98), ‘Bolitophagus pygmaeus’ in decaying tree stumps, 18- 

25.vii.1913, OEJ (Janson, 1914). 

H33 Fermanagh: Rossyvullan Hill, Castle Coole, one in Polyporus squamosus on dead standing 

sycamore, 20.vii.2010, RA. 

H37 Armagh: Tynan Abbey, three in Polyporus squamosus on elder, 26.vii.2011, RA/MDB. 

H39 Antrim: near Belfast, R. Templeton (Johnson, 1889), ‘now in collection of Belfast Nat. Hist. & 

Philosophical Society’ (Johnson & Halbert, 1902); Hammond (1980) regards this record as 

unconfirmed. 

Phloeonomus punctipennis Thomson, C.G., 1867 – Common & Widespread 

Adults and larvae under bark of various dead broad-leaved wood; a fungal feeder; under-recorded 

due to past confusion with P. pusillus; “probably widely distributed…. generally distributed” in 

Ireland H2, H4, H13, H21, H25, H30, H36, H33, H37, H38, H39 (Hammond, 1980). 

Phloeonomus pusillus (Gravenhorst, 1806) – Common & Widespread 

Adults and larvae live under bark, mostly of dead conifer wood; widespread in Britain; “apparently 

less generally distributed in Ireland than P. punctipennis” H1, H2, H15, H17, H19, H20, H34, H36, H37, 

H38, H40 (Hammond, 1980). 

Phloeostiba plana (Paykull, 1792) – Rare old growth species 

Adults attracted to fresh sap of broad-leaved trees – they feed on the sap; larvae develop under dead 

bark, feeding on sap as well as insects; mostly in areas of ancient woodland. 

H1 South Kerry: Glencar, vii.1924, OEJ (Janson & Bonaparte-Wyse, 1924). 

H2 North Kerry: Flesk and Killarney, “seen specimens” (Hammond, 1980). 

H20 Wicklow: Ballard Wood (T1495), 14.vi.1993, KNAA. 

H22 Meath: Balrath, vi.1916, GWN (Nicholson, 1917). 

H36 Tyrone: Baronscourt Park (H3682), 12.ix.2006, KNAA. 

Xylostiba monilicornis (Gyllenhal, 1810) – Rare 

Confined in Britain to ancient broad-leaved woodland and pasture woodland, mainly in the midlands 

and north. Found in gill fungi and in Piptoporus betulinus on birch, also reported from oak, beech and 

pine. Appears during winter and in early spring. Only recently found in Ireland but presumed to be 

an overlooked native. 

H37 Armagh: Slieve Gullion Forest Park (J038193), under bark on dead standing silver fir, 8.iii.2012, 

RA. 

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H38 Down: Donard Demesne, Newcastle (J373301), 1 pair under fungoid bark of veteran beech on the 

ground, 9.i.2012, RA; Leitrim Lodge (J223256), 1 under bark of dead standing Scots pine, 29.ii.2012, 

RA. 

STAPHYLINIDAE SUBFAMILY PSELAPHINAE Short-Winged Mould Beetles 

Predatory, particularly on mites; a number associated with deadwood. 

Euplectus bescidicus Reitter, 1881 – Rare 

Lives amongst insect borings and frass in old trees. 

H2 North Kerry: near Killarney, in burrows of wood-wasp Sirex in an old tree, v.1928, EFB 

(O’Mahony, 1929a). 

Euplectus infirmus Raffray, 1910 – Rare 

In rotten wood and under bark, especially willow. Ireland (Pearce, 1957). 

H6 Waterford: Waterford, JAP (Power, 1878). 

H33 Fermanagh: Castle Coole Park (H2643), from FIT in hollow horse chestnut, 14.ix.2006, KNAA. 

H38 Down: Hillsborough Estate (J243576), 1 in pitfall in parkland, 5.vi.1992, RA. 

Euplectus karstenii (Reichenbach, 1816) – Rare 

H20 Wicklow: Ashford, ARF (Johnson & Halbert, 1902). 

H21 Dublin: Dublin unlocalised, ARH det. AHH (Hogan, 1854); Dundrum, FHB det. JNH (Johnson & 


H33 Fermanagh: Castle Coole Park (H2643), from FIT in hollow horse chestnut, 14.ix.2006, KNAA. 

Euplectus mutator Fauvel, 1895 syn. fauveli Guillebeau, 1888 – Rare 

In bird nests, under dead bark and in rotten wood; oak and beech. Ireland (Pearce, 1957). 

H2 North Kerry: Bullock’s house at Flesk, Killarney, EFB (O’Mahony, 1929a). 

Euplectus piceus Motschulsky, 1835 – Rare old growth species 

Under bark of oak and beech, and in red-rotten heartwood decay in oak. 

H2 North Kerry: near Killarney, EFB (Bullock, 1914). 

H25 Roscommon: St John’s Wood (M9956), under bark on fallen oak branch, 14.iv.2010, KNAA. 

Euplectus punctatus Mulsant, 1861 – Extinct 

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In moist crumbly rotten wood; primarily a relict species of old growth forest; Ireland (Pearce, 1957). 

H2 North Kerry: ‘Kerry’, JAP (Power, 1878). 

Bibloporus bicolor (Denny, 1825) – Rare old growth species 

Under bark and in rotten wood of a wide variety of trees, broad-leaves and conifers. 

H2 North Kerry: Killarney, Bullock (1914). 

H20 Wicklow: Powerscourt, Heasler (O’Mahony, 1929a). 

H33 Fermanagh: Inishfendra, Crom, FIT, 16.vi-15.vii.1993, APF. 

STAPHYLINIDAE SUBFAMILY PHLOEOCHARINAE 

Phloeocharis subtilissima Mannerheim, 1830 – Scarce 

Amongst debris under beech bark, in moss on trees, on bracket fungi especially Daedaleopsis 

confragrosa on Salix, etc. 

H1 South Kerry: Valentia Island (V47), taken commonly by Miss M.J. Delap, in decaying Fuchsia sticks 

and in moss (Halbert 1898b) (Johnson & Halbert, 1902). 

H2 North Kerry: Kenmare Demesne (V99), in some numbers beneath the bark of dead firs, between 18 

& 25 July 1913, OEJ (Janson, 1914). 

H9 Clare: Kilrush, v.1914, EFB. 

H34 East Donegal: Milford, ix.1906, Halbert (1937). 

H37 Armagh: Peatlands Park (H904606), 23.v.2007, RA; Derryadd Lough (H911606), in Piptoporus on 

birch, 11.v.2012, RA. 

H38 Down: Rostrevor, 15.iv.1974, CAMR; Rostrevor Oakwood, 2000 (Anderson, Simms & Nelson, 

2000); Tollymore Park, 6.iv.2011, MDB; Donard Park (J372298), 9.i.2012, RA. 

H39 Antrim: Shane’s Castle, Sept 1902, C.W. Buckle (Halbert 1910); under bark of fallen firs, June 

1923, O.E. Janson (1924); Rea’s Wood, Antrim, 17.iv.1974, CAMR. 

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STAPHYLINIDAE SUBFAMILY TACHYPORINAE 

Sepedophilus littoreus (Linnaeus, 1758) – Scarce 

Mycetophagous species, feeding on fungal hyphae not fruiting bodies - rarely away from dead and 

decaying wood of some kind; occasionally found amongst leaf and other vegetable litter. Generally 

distributed in Britain, largely in ancient woodlands. Although not known prior to 1977, it seems 

probable that the species is an old woodland relict in Ireland (Anderson, 1978). 

H6 Waterford: Dunkitt Castle meadow, pitfall in riverine grassland, v.-vi, 1999, JAG. 

H19 Kildare: Pollardstown Fen, pitfall in fen, vi.1998, v.-vii.2002, JAG. 

H33 Fermanagh: near Inishteige, Upper Lough Macnean (H0242), female from roadside grass cuttings 

bordering freshly felled scrub woodland, 12.iv.1977, RA (Anderson, 1978); Inisherk, Crom, FIT, vi.- 

vii.1992, APF. 

H34 East Donegal: Dunkineely, from Polyporus squamosus on stump, 6.vi.1999, KNAA. 

H38 Down: Tollymore Forest, under piled up Douglas fir bark, 9.iv.1993, RA. 

H39 Antrim: near Portmore Lough (J119686), pair from under bark around fruiting fungus Phlebia 

merismoides on a rotting alder log in marginal old alder woodland, 24.xi.1984, RA (Anderson, 1985); 

Bucklebrook House, Upper Ballinderry, 31.iii.2011, MDB. 

H40 Londonderry: by Toneduff Bridge, R. Faughan (C5107), male in basidiomycete fungi on beech 

logs, 31.x.1977, RA (Anderson, 1978). 

STAPHYLINIDAE Subfamily ALEOCHARINAE 

Alaobia pallidicornis (Thomson, C.G., 1856) – Uncommon 

At sap, bracket fungi, and under fungoid bark in forest trees. Local and mainly south-eastern in 

Britain, but more generally distributed in Ireland. Possibly not an obligate saproxylic. 

H18 Offaly: Charleville Wood, Tullamore, in bracket fungus on horse chestnut, 24.v.2003, RCW 

(Regan & Anderson, 2003). 

H19 Kildare: Louisa Bridge, Royal Canal, in ‘Polyporus’ on poplar, viii.1942, EO (O’Mahony, 1943). 

H30 Cavan: Cloverhill, at sap of damaged oak, vi.1916, GWN (Nicholson, 1920). 

H33 Fermanagh: Rossyvullan Hill, Castle Coole, under fungoid bark on oak limb, 28.v.2010, RA. 

28


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H37 Armagh: Tynan Abbey Estate, in Polyporus squamosus on birch, 26.vii.2011, RA; Tynan Abbey, in 

Trametes on ash, 17.viii.2011, RA. 

H38 Down: Hydebank Woods, under fungoid bark of sycamore log, 25.x.2007, RA; Mount Stewart, in 

Russula under beech, 11.viii.2010, RA; Dundrum Castle (J406371), a pair in fungi on dead wood, 

14.iii.2012, MDB. 

H39 Antrim: Glenarm Nature Reserve, in Piptoporus on birch, 8.vi.2011, RA. 

Atheta basicornis (Mulsant & Rey, 1852) – Scarce 

Under bark on periodically inundated dead branches and trunks along fast-flowing rivers; feeds on 

fungal material. 

H15 South-east Galway: Lough Coy Turlough, Ardrahan, pitfall, 6.v.1996, JAG. 

H18 Offaly: Bullock Island, Shannon, under willow bark, 24.v.2003, RCW (Regan & Anderson, 2003). 

H30 Cavan: Cloverhill, under pine bark, vi.1916, GWN (Nicholson, 1920). 

H33 Fermanagh: Rossyvullan Hill, Castle Coole, under fungoid bark on oak, 28.v.2010; Munilly 

Bridge, Old Ulster Canal, under willow bark, 19.vii.2011, RA. 

H37 Armagh: Oxford Island (J047621), in fungus on willow, 25.iii.2012, MDB 

H38 Down: Murlough NNR, in Agaricus campestris, dunes, 23.ix.2006, RA. 

H39 Antrim: Portmore Lough, under bark of willow and alder, 19.vi.2009, 26.vi.2009, RA. 

Atheta pilicornis (Thomson, 1852) – Rare 

Chiefly subcortical, often with scolytids; adults also reported from wood-decay and terricolous fungi, 

moss and among dead leaves, but larval habits not known; damp woodlands. 

H2 North Kerry: near Killarney, in dead leaves, vi.1932, EFB det. Ste. Claire Deville (Bullock, 1932). 

H38 Down: Belvoir Forest (J339696), 1 male in bolete Xerocomus chrysenteron under oak, 11.ix.1997, RA; 

Dundrum Castle woods, 1 female on Polyporus squamosus on horse chestnut, 20.vii.2009, RA. 

Bessobia fungivora (Thomson, 1867) – Rare 

In rotting fungi or on fungoid wood in forests; probably not an obligate saproxylic. Historically 

known only from a record of R.E. Dillon for Clonbrock, a specimen having been determined by J.N. 

Halbert (Johnson & Halbert, 1902). Dillon has become notorious for some of the outlandish records 

claimed for Clonbrock which, but for a modern record, would have made the species doubtfully Irish. 

H16 North-east Galway: Clonbrock, RED, det. JNH (Johnson & Halbert, 1902). 

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H38 Down: Hollymount NNR (J465440), in Stereum bracket fungi on beech logs, 18.i.2011, RA 

(Anderson, 2011). 

Dadobia immersa (Erichson, 1837) – Rare 

Under bark, usually pine, but also reported from decaying alder trunks and Nothofagus. 

H1 South Kerry: Glencar (V78), several shaken from the chippings of felled trees, 8.vii.1924, OEJ 

(Janson & Bonaparte-Wyse, 1924). 

Dinaraea aequata (Erichson, 1837) – Common & Widespread 

Under bark of various trees, usually near fungoid growths. H2, H4, H5, H15, H18, H19, H20, H23, 

H33, H36, H37, H39, H40. 

Dinaraea linearis (Gravenhorst, 1802) – Rare old growth species 

Under bark. 

H2 North Kerry: Killarney, Bullock (1914) and collection specimens, EFB (Good, 1990); Ballyseedy 

Wood, xi.2001, JAG. 

H4 Mid Cork: Curraghabinny Wood, near Crosshaven, in frass on oak log, 20.vii.2000, RA. 

H37 Armagh: Derryadd Lough (H911606), in Piptoporus on birch, 11.v.2012, RA; Loughgall Country 

Park (H909513), under bark, dead alder, MDB. 

Traumoecia picipes (Thomson, 1856) – Uncommon 

In fruit bodies of wood-decay fungi such as Piptoporus betulinus, Meripilus giganteus & Hypholoma; 

overwintering in tussocks, moss, compost, etc. Identified as an indicator species of “well-developed 

woodland habitat” (Good & Butler, 1995). 

H2 North Kerry: Ballyseedy Wood, xi.2001, JAG; Muckross Woods, 2.ix.2011, RA. 

H6 Waterford: Lismore, 13.iii.2011, RA. 

H12 Wexford: Johnstown Castle, 14.viii.2009, SM. 

H20 Wicklow: Powerscourt Demesne, Sept/Oct, 1990, JAG (Good & Butler, 1995). 

H31 Louth: Bellurgan Foresty (J095099), 22.iii.2012, RA. 

H33 Fermanagh: Inisherk Island, 31.iii.1997, RA; Florencecourt, 16.iii.2011, RA. 

H36 Tyrone: Baronscourt Estate, 16.v.2006, RA. 

H37 Armagh: Tynan Abbey, 26.vii.2011, 17.viii.2011, RA; Slieve Gullion Forest (J037194), 8.iii.2012, 

RA. 

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H38 Down: Belvoir Forest, 16.iv.2004; Mount Stewart, 13.ix.2003, 17.x.2009, 13.viii.2010, RA; 

Killynether Wood, 11.viii.2007, RA; Helen’s Tower, 26.vii.2008, 10.ii.2011, RA; Rostrevor Oakwood 

NNR, 7.iii.2011, MDB. 

H39 Antrim: near Lough Neagh (O’Mahony, 1929a); Shane’s Castle, vi.1902, CWB (Halbert, 1910b); 

Barnett’s Park, 2.x.1996, RA; Belfast Castle, 11.vii.2004, RA. 

Thamiaraea hospita (Märkel, 1844) – Rare 

At exuding sap on tree trunks, especially oak. 

H21 Dublin: Lucan Demesne, in borings of the goat moth on oak, 1909, JNH (Halbert, 1910a). 

Bolitochara lucida (Gravenhorst, 1802) – Rare 

In fleshy fungus on old stumps. 

H21 Dublin: Glencullen, AWS (Stelfox, 1924) (O’Mahony, 1929a). 

H39 Antrim: Fair Head & Murlough Bay, 15.vi.1992, KNAA. 

Bolitochara obliqua Erichson, 1837 – Common & Widespread 

Under bark of dead wood of various deciduous trees, especially associated with the small bracket 

fungus Trametes versicolor; adult is a fungal feeder, larvae also feeding on phloem and dead larvae. 

Common in Britain and Ireland: H9, H18, H19, H20, H21, H22, H28, H32, H33, H34, H36, H37, H38, 

H39. 

Leptusa fumida (Erichson, 1839) – Common & Widespread 

Under bark on dead wood of various trees, also in rot holes; feed mostly on detritus and fungi, but 

will eat insect larvae and eggs. Widespread in Ireland: H1, H2, H4, H13, H18, H19, H20, H21, H30, 

H33, H36, H37, H38, H39, H40. 

Leptusa norvegica Strand, A., 1941 – Established Introduction 

Lives under bark in the runs of ambrosia beetles, etc, on conifer logs. 

H39 Antrim: Portglenone Forest, in Tomicus burrows, Scots pine, SC (Anderson & Clawson, 2004). 

H38 Down: Donard Park, under loose bark of silver fir, 31.xii.2011, RA. 

Leptusa pulchella (Mannerheim, 1830) – Status unclear 

Under dead bark on a wide variety of tree species. Mainly in ancient woodlands in Britain, especially 

where wet. ‘Leptusa analis’ in Johnson & Halbert (1902). Only reported from the northernmost 

counties, where widespread. 

H33 Fermanagh: Florencecourt, under bark, beech log, 16.iii.2011, RA. 

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H36 Tyrone: Caledon Deer Park (H7543), 16.viii.2006, KNAA. 

H38 Down: Rostrevor Oakwood, under bark of oak logs, 13.v.2000 (Anderson, Simms & Nelson, 

2000); Murlough NNR, under dying elm bark, 15.xi.2010; Hollymount Wood NNR, in Trametes 

versicolor, ash log, 18.i.2011, RA; The Drinns, Tollymore Forest, under bark, 6.iv.2011, MDB; Mallard 

plantation, Castleward, under bark of Pseudotsuga log, 21.viii.2010, RA; Shooter’s Island, Quoile 

(J494470), 1 in deer dung(!), 23.ii.2012, MDB; Tullyframe Wood (J264169), under bark on beech, 

27.iii.2012, RA. 

H39 Antrim: Murlough Bay, one under bark, ix.1901, GWC det. JNH; Great Deer Park, Glenarm 

(D2911), 20.v.2006, KNAA; Glenarm Nature Reserve, under bark of oak log, 8.vi.2011, 14.vi.2011, in 

Piptoporus on birch, both RA. 

H40 Londonderry: Altnaheglish Forest, Banagher, Sitka spruce stump, 6.viii.2005, RA. 

Leptusa ruficollis (Erichson, 1839) – Recent arrival 

Under bark on deadwood; favours thin fibrous-barked woody plants notably Tilia & frequently 

Clematis; ancient woodlands in Britain, but appears to be a recent arrival in Ireland. 

H38 Down: Mount Stewart House, 1 in rotting Russula under beech, 13.viii.2010, RA; Temple of the 

Winds, Mount Stewart, 1 under fresh bark on fallen beech bough, 5.x.2010, RA. 

Gyrophaenini are associated with fungal fruiting bodies, generally wood decay fungi but not 

exclusively so; larvae spore-feeding. 

Agaricochara latissima (Stephens, 1832) – Uncommon 

H2 North Kerry: Killarney, Bullock (1914) (O’Mahony 1929); Ballyseedy Wood, ix.2001, JAG. 

H30 Cavan: Cloverhill, 13.ix.1913, GWN (Nicholson, 1920; O’Mahony 1929). 

H36 Tyrone: Baronscourt Park, 15.vi.2006, KNAA, det. RA. 

H37 Armagh: Ardress House, 25.ii.2011, RA. 

H38 Down: Dane’s Wood, Clandeboye, 21.x.1973, CAMR; Stormont, 18.ix.2004, RA; Castle Espie, 

19.viii.2007, RA; Belvoir Forest, 22.xi.2007, RA; Murlough NNR, 15.ix.2010, RA; Hollymount NNR, 

18.i.2011, RA. 

H39 Antrim: Ballycastle, GWC (Johnson & Halbert, 1902); Barnett’s Park, 29.ix.1993, RA; Portmore 

Churchyard, 29.ix.2009, RA; Belfast Castle, 23.i.2011, RA; Glenarm Nature Reserve, 1.xi.2011, RA. 

Gyrophaena affinis Mannerheim, 1830 – Uncommon 

H16 North-east Galway: Clonbrock (M7439), in fungi on trees (Halbert, 1896; Johnson & Halbert, 

1902). 

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H20 Wicklow: Powerscourt Deerpark, 20.vi.1928, EO (O’Mahony, 1929a). 

H23 Westmeath: Shurock House, pitfall, vi.2007, P. O’Connell. 

H25 Roscommon: St John’s Wood (M9956 & N0055), Armillaria on oak roots, 1.ix.2010 & frequent in 

FITs, 2.xi.2010, KNAA. 

H32 Monaghan: Castle Leslie, Glaslough, 18.vii.2009, RA. 

H36 Tyrone: Gortin Glen Forest (H484819), 21.vii.2012, RA. 

H38 Down: Barnett’s Park, 29.ix.1993, RA; Hillsborough Forest, 28.iv.1995, RA; Tollymore Forest, 

27.vi.1998, RA; Stormont Estate, 18.ix.2004, RA; Castle Espie, 19.viii.2007, RA; Newforge, Belfast, 

23.vii.2008, RA; Helen’s Tower, Clandeboye, 26.vii.2008, RA; Killynether Wood (J475722), 29.vi.2012, 

RA. 

H39 Antrim: Shane’s Castle Demesne, Lough Neagh, “only recorded from Galway in the Irish list”, 

vi.1923, OEJ (Janson, 1924); Rea’s Wood, Antrim, 3.xii.1994, RA; Portglenone Forest, 28.viii.2004, RA; 

Glenarm Nature Reserve, 14.vi.2011, RA. 

Gyrophaena bihamata Thomson, C.G., 1867 – Rare 

H2 North Kerry: near Killarney, EFB (Bullock, 1928; O’Mahony, 1929a). 

H32 Monaghan: Castle Leslie, Glaslough, in Polyporus squamosus on ash, 18.vii.2009, RA. 

H39 Antrim: Rea’s Wood, in Clitocybe under spruce, 30.vi.1986, RCW (Welch, 1986); in Hirneola 

auricula-judae on sycamore, 3.vii.2004, RA. 

Gyrophaena fasciata (Marsham, 1802) – Common & Widespread 

H2,H6,H9,H17,H20,H24,H25,H30,H32,H33,H36,H37,H38,H39. 

Gyrophaena gentilis Erichson, 1839 – Uncommon 

H20 Wicklow: Powerscourt Demesne, 1 with Daedalea quercina on an oak stump, ix-x.1990, Good & 

Butler (1995). 

H34/40: Foyle district, CWB (Buckle, 1900; Johnson & Halbert, 1902. Suppl. List). 

H32 Monaghan: Castle Leslie, Glaslough, 18.vii.2009, RA. 

H33 Fermanagh: Lough Fadda, Blessingbourne, 27.vii.2011, RA. 

H38 Down: Belvoir Forest, 27.x.1993, 9.ix.1995, 28.ix.1996, RA; Hollymount NNR, 5.x.1996, RA; 

Shane’s Castle, 3.vii.2004, RA; Belfast Castle, 11.vii.2004, RA; Killynether Wood (J475722), in Polyporus 

squamosus on sycamore, 29.vi.2012, RA. 

33


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H39 Antrim: Ballycastle, in fungi, “obtained commonly in woods near Ballycastle by G.W. Chaster” 

(Johnson & Halbert, 1902); Shane’s Castle, vi.1902, CWB (Halbert 1910); Rea’s Wood, 12.ix.2007, RA. 

Gyrophaena manca Erichson, 1839 syn. angustata (Stephens, 1832) – Rare 

Larvae feed especially on spores of Polyporus squamosus. A Palaearctic species, widespread in Europe, 

but very local in Britain and rare in Ireland. 

H15 South-east Galway: near Aughrim (M825268), 3m & 10f in bracket fungus Polyporus squamosus on 

cut mature ash stump in a hedge in an area of esker ridge scrub, old pasture and hedgerows, JAG 

(Good, 1997). 

H37 Armagh: Tynan Abbey Estate (H715414), abundant in Polyporus squamosus on elder, 26.vii.2011, 

RA/MDB. 

H38 Down: Killynether Wood (J475722), in Polyporus squamosus on sycamore, 29.vi.2012, RA. 

Gyrophaena minima Erichson, 1837 – Rare 

H28 Sligo: Glencar, GWC, det. GCC (Johnson & Halbert, 1902). 

H38 Down: Belvoir Forest, 5 from Stereum hirsutum on a Prunus laurocerasus log, 1.ix.1997, RA. 

H39 Antrim: Ballycastle, GWC, det. JNH (Johnson & Halbert, 1902). 

Gyrophaena nana (Paykull, 1800) – Rare 

H22 Meath: Balrath, new to Irish list, ix.1910, GWN (Nicholson, 1911) (O’Mahony, 1929a) 

Gyrophaena poweri Crotch, 1867 – Rare 

H2 North Kerry: Ross Castle Lakeside, Killarney, in fungi, EFB (Bullock, 1935). 

Gyrophaena pulchella Heer, 1839 – Rare 

H2 North Kerry: Killarney, Bullock (O’Mahony, 1929a). 

H22 Meath: Balrath, new to Irish list, 26.ix.1913, GWN (Nicholson, 1914) (O’Mahony, 1929a). 

H36 Tyrone: Drum Manor, a pair in fungi, 18.x.2010, MDB det. RA. 

H38 Down: Belvoir Forest, 6 in fresh fungus Clitocybe geotropa under beech, 27.x.1993, RA (Anderson, 

Simms & Nelson, 2000). 

Gyrophaena strictula Erichson, 1839 – Rare old growth species 

Characteristically associated with the bracket fungus Daedalea quercina but occasionally reported with 

other fungi; this bracket fungus is itself very localised in Ireland and known from only seven sites in 

four vice-counties (Muskett & Malone, 1980). The beetle has been identified as an indicator species of 

34


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“well-developed woodland habitat” (Good & Butler, 1995); widely distributed and not uncommon in 

N Europe. 

H20 Wicklow: Powerscourt Demesne (O221172), 8m, 3f, in Daedalea quercina on oak stump, ix-x.1990, 

JAG (Good, 1991a; Good & Butler, 1995). 

Anomognathus cuspidatus (Erichson, 1839) – Scarce 

Under fungoid bark of dead beech, oak, etc. 

H2 North Kerry: near Killarney, 1914, EFB (Bullock, 1914). 

H15 South-east Galway: Ardfry, under bark, vi.1921, JNH (Halbert, 1937). 

H20 Wicklow: Ballyward Bridge, under bark, vi.1921, JNH (Halbert, 1937). 

H21 Dublin: Grand Canal bank, near Lucan, under bark of felled elms, larvae extremely abundant but 

adults rather scarce, May, JNH (Johnson & Halbert, 1902. Suppl. List). 

H37 Armagh: Tynan Abbey Estate, 21.vi.2011, RA. 

H38 Down: Morelands Meadow, Belvoir, 18.xi.1992, 15.iv.2004, RA; Stormont Estate, 7.v.1994, 

13.v.1994, RA; Hydebank Woods, 25.x.2007, RA; Mount Stewart, 16.viii.2010, RA; Temple of the 

Winds, 5.x.2010, RA. 

H39 Antrim: Lagan Meadows, Belfast (J334699), decaying Bjerkandera adusta bracket fungi on a 

sycamore stump, 11.iv.1993, Anderson (1995); Portmore Lough, RSPB Reserve, 19.vi.2009, RA; Moss 

Plantation, Randalstown, 16.vii.2010, RA. 

Homalota plana (Gyllenhal, 1810) – Uncommon 

Under bark on dead beech etc; adult feeds on detritus and fungi. 

H2 North Kerry: Deer Park, Killarney, one under bark, EFB (Bullock, 1935). 

H25 Roscommon: St John’s Wood (M9956), 14.iv. & 12.v.2010, KNAA. 

H37 Armagh: Annagarriff NNR (H902616), sycamore bark, 9.v.2000, RA; Peatlands Park, 2000 

(Anderson, Simms & Nelson, 2000); Tynan Abbey Estate (H755414), bark, beech, 26.vii.2011, RA. 

H38 Down: Morelands Meadow (J337697), beech logs, 18.xi.1992, RA; Belvoir Forest (J341695), bark 

Scots pine, 11.iv.1995, RA; Hydebank woods (J342677), bark, sycamore, 25.x.2007, RA; Mount stewart 

(J550699), in rotting Russula, 11.viii.2010, RA; Castle Ward (J572502), bark, ash, 19.iii.2011, RA; 

Shooter’s Island, Quoile (J494471), under sycamore bark, 23.ii.2012, RA. 

H39 Antrim: Farr’s Bay NNR (J089870), bark, Picea log, 4.vii.1986, RCW (Welch, 1986); Belfast Castle 

(J327793), fungoid bark on beech, 11.vii.2004, RA; Moss Plantation, Shane’s Castle (J094893), bark, oak, 

20.viii.2010, RA. 

35


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Cypha seminulum (Erichson, 1839) – Extinct 

In rotting wood, fungi, under bark and in moss; elm and sycamore. Ireland (Haliday collection). 

H34/40: Foyle District, CWB (Buckle, 1900; Johnson & Halbert, 1902). 

Oligota apicata (Erichson, 1837) – Rare 

Occurs in bracket fungi on trees: reported from Stereum hirsutum, Polyporus squamosus, Trametes 

versicolor; possibly preys on Cis beetle larvae; also recorded in leaf litter inside a hollow oak, under 

beech bark, etc. 

H15 South-east Galway: Coole Park (M433043), 22.v.2010, PJH. 

H38 Down: Helen’s Tower, Clandeboye, in rotting Trametes/Bjerkandera on beech logs, 10.ii.2011, RA. 

H39 Antrim: Shane’s Castle Demesne, Lough Neagh, two sifted from debris, vi.1923, OEJ (Janson, 

1924). 

Ischnoglossa prolixa (Gravenhorst, 1802,) – Rare 

Found beneath bark on freshly dead timber; widespread on the Continent. 

H15 South-east Galway: Ardfry, under bark, vi.1921, JNH (Halbert, 1937). 

H16 North-east Galway: Clonbrock (M7439), collected by R.E. Dillon, det JNH (Johnson & Halbert, 

1902). 

H19 Kildare: Curragh camp (M774118), 2 under bark of Scots pine, 17.iii.2012, RA. 

H20 Wicklow: Ballyward Bridge, under bark, vi.1921, JNH (Halbert, 1937); Powerscourt Demesne, 16 

under Pseudotsuga bark, Sept/Oct, 1990, JAG (Good & Butler, 1995). 

H21 Dublin: Dundrum, 1900, EFB (Halbert, 1900; Johnson & Halbert, 1902). 

H37 Armagh: Slieve Gullion Forest Park (J037194), 1 under bark of dead standing Scots pine, 8.iii.2012, 

RA; Loughgall (H909513), under bark of dead alder, 3.v.2012, MDB. 

H38 Down: Belvoir Forest (J342694), in Bjerkandera adusta on oak stump, 23.iv.2004, RA; Leitrim Lodge 

(J223256), under bark of dead standing Pinus contorta, 29.ii.2012, RA. 

Phloeopora testacea (Mannerheim, 1830) – Uncommon 

Predatory, beneath bark on deadwood of various broad-leaved trees. Phloeopora reptans (syn.) reported 

from Ireland by Haliday and there are Irish-taken specimens in his collection (Johnson & Halbert, 

1902). 

H2 North Kerry: near Killarney, 1914, EFB (Bullock, 1914); Kenmare Demesne, Killarney, P. reptans in 

some numbers under bark of recently felled pines, 25.vi.-30.vi.1919, OEJ (Janson, 1920). 

36


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H19 Kildare: Curragh camp (N774118), 17.iii.2012, RA. 

H20 Wicklow: Powerscourt Deerpark, 18.ii.2007, RA. 

H21 Dublin: Clontarf, date unknown, OE (O’Mahony, 1938). 

H33 Fermanagh: Florencecourt, 31.vii.2010, RA. 

H37 Armagh: Annagarriff NNR, 29.v.2000, RA. 

H38 Down: Belvoir Forest, 28.ix.1993, 12.iv.1995, RA; Redburn Country Park, 6.ix.2006, RA; Hydebank 

Woods, 25.x.2007, RA; Castle Espie, 26.viii.2009, RA. 

H39 Antrim: Barnett’s Park, 19.v.1994, RA; Belfast Castle, 11.vii.2004, RA; Ram’s Island, L. Neagh, 

4.vi.2005, RA; Shane’s Castle Estate, 1.vii.2010, RA; Moss Plantation, Randalstown, 16.vii.2010, RA. 

H40 Londonderry: Leganannon Pot, 4.iv.1999, RA. 

STAPHYLINIDAE SUBFAMILY SCAPHIDIINAE – shining fungus beetles 

Scaphisoma agaricinum (Linnaeus, 1758) – Uncommon 

Amongst decaying wood and at fungal fruit bodies. 

H14 Laois: Glendine Valley, Slieve Bloom, 1907, JNH (Halbert, 1907a). 

H15-17 Galway: Woodford, JNH (Johnson & Halbert, 1902); Derryclare Wood (L8249), in birch 

polypore, 20.v.1988, KNAA. 

H20 Wicklow: Powerscourt Deer Park (O1912), 19.vi.1993, KNAA. 

H21 Dublin: 1878, JAP (Power, 1878); Woodlands, JNH (Halbert, 1894; Johnson & Halbert, 1902). 

H23 Westmeath: Derravaragh, JNH (Johnson & Halbert, 1902). 


H33 Fermanagh: Inishfendra, Crom, FIT, 15.vii.1993-24.viii.1993, APF; Molly Mountain, 23.vi.1995, 

RA; Castle Coole Park, 2.vi.1992 & 18.v.2006, KNAA. 

H36 Tyrone: Caledon Park, 16.viii.2006, KNAA; Brookend NR, L. Neagh, 8.v.2007, RA. 

H37 Armagh: Lowry’s Lough, sweeping gorse, WFJ (Johnson, 1892a) (Johnson Halbert, 1902); 

Derryadd Lough (H914609), in old Piptoporus, 11.v.2012, MDB. 

H38 Down: Dundrum Castle (J406371), in fungi on dead wood, 14.iii.2012, MDB. 

H39 Antrim: Shane’s Castle Demesne, Lough Neagh, June 1923, OEJ (Janson, 1924). Great Deer Park, 

Glenarm, 28.vii. & 19.viii.2006, KNAA. 

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Scaphisoma boleti (Panzer, 1793) – Scarce 

H2: North Kerry: Ardagh (W9788), near Killarney, vii.1927, JNH (Halbert, 1937). 

H16 North-east Galway: Clonbrock (M7439), coll R.E. Dillon, det. JNH, (Johnson & Halbert, 1902); 

Ardfry, vi.1916, JNH (Halbert, 1937). 

H21 Dublin: Dundrum, JNH (Halbert, 1895b) (Johnson & Halbert, 1902). 

H33 Fermanagh: Summerhill Lough, xii.1905, WFJ (Johnson, 1906); Florencecourt, 21.vi.2010, RA; 

Filbert Hill, Castle Coole, 26.vi.2010, RA. 

H34 East Donegal: St. Ernan’s island (G9278), 30.vi.1908, WFJ (Johnson, 1909). 

H37 Armagh: Tynan Abbey, 1.vii.2011, RA. 

H39 Antrim: Rea’s Wood, Antrim, 30.vi.1986, RCW (Welch, 1986). 

STAPHYLINIDAE SUBFAMILY PIESTINAE 

Siagonium quadricorne Kirby, 1815 – Scarce 

Saprophagous beneath moist sappy bark on the freshly dead wood of various broad-leaved trees, 

especially elm. Old records from southern counties, where still present; recently appeared in north. 

H5 East Cork: Fota Wildlife Park, 1 adult, 1 larva under bark of Scots pine, 11.vii.2001, RA. 

H13 Carlow: Borris, under bark of moist decaying ash stump, iv.1895, JNH (Halbert, 1895e) (Johnson 

& Halbert, 1902) 

H19 Kildare: Leixlip, “seen specimens taken in August 1942” (Hammond, 1980). 

H37 Armagh: Tynan Abbey, in Polyporus squamosus on elder, 26.vii.2011, RA; under oak bark among 

lumber, 29.ix.2011, RA. 

H38 Down: Hydebank Woods, Belfast, under bark of sycamore log with Quedius lyszkowskii, 25.x.2007, 

RA; Murlough NNR, under bark of dead elm, 15.xi.2010, RA; Temple Water, Castleward, under ash 

bark, 19.iii.2011, RA; Shooter’s Island (J580493), under bark of dead standing sycamore (32 on a single 

tree), 23.ii.2012, RA. 

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Figure 11: Siagonium quadricorne © Roy Anderson 

STAPHYLINIDAE SUBFAMILY STAPHYLININAE 

Atrecus affinis (Paykull, 1789) – Common & Widespread 

Under dead bark and in rotten wood of various trees, conifers & broad-leaved trees. Widespread and 

locally common throughout Ireland. Baptolinus alternans in Johnson & Halbert (1902). 

H1,H2,H4,H5,H15,H20,H27,H29,H32,H33,H35,H36,H37,H38,H39,H40. 

Gabrius splendidulus (Gravenhorst, 1802) – Status unclear 

Under dead bark, especially of beech. The Furlong record below is from very atypical habitat and may 

be due to a misidentification. The Mount Stewart record has been verified and suggests a recent 

arrival in the north. 

H1 South Kerry: Glenbeigh (V69), sea weed, ARF (Johnson & Halbert, 1902). 

H38 Down: Mount Stewart, 19.v.1992, APF. 

Quedius brevicornis (Thomson, 1860) – Rare 

Lives within nests of birds inside tree cavities; also found in tree rot holes and fungi. 

H6 Waterford: Waterford, JAP (Power, 1878) (Johnson & Halbert, 1902). 

39


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H39 Antrim: Collin Glen (J270720), 6.ix.2002, KNAA. 

Quedius lyszkowskii Lott, 2010 – Rare 

A recently described species, previously overlooked within Q. aetolicus Kraatz by modern workers and 

historically with Q. puncticollis in Irish collections (Lott, 2010). So far known only from Scotland and 

Ireland; may be endemic to the British Isles. Of the historical Irish records of ‘Q. puncticollis’ only 

material from Killarney and Portmarnock have so far been examined. Summer and autumn records 

are typically associated with decaying wood, but one winter record is from atypical habitat and may 

have been a dispersing individual. 

H2 North Kerry: Killarney, 22.vi.1927, LHBW (BMNH) (Lott, 2010). 

H21 Dublin: Portmarnock, 7.vi.1905, collector unknown (BMNH) (Lott, 2010). 

H36 Tyrone: Baronscourt Park (H3682), under dead bark on veteran beech, 12.ix.2006, KNAA. 

H38 Down: Hydebank Woods, under fungoid sycamore bark, 25.x.2007, RA. 

H39 Antrim: Barnett’s Park, under sycamore bark, 12.xii.1979, RA; Belfast Castle, under fungoid bark 

on beech, 11.vii.2004, RA; Bucklebrook House, Upper Ballinderry, in hen litter, 6.xi.2010, MDB. 

H40 Londonderry: Roe Valley Country Park, under bark of stacked conifer logs, 16.vii.2011, RA. 

Quedius plagiatus Mannerheim, 1843 syn. laevigatus Gyllenhal, 1810 – Rare old growth species 

Lives beneath loose bark and amongst moist well-rotted timber; broad-leaved trees & conifers. 

H1 South Kerry: Lickeen Wood, Glencar (V78), one in an oak stump, July 1924, OE Janson (Janson & 

Bonaparte-Wyse 1924) 

H2 North Kerry: Rossbeigh Dunes, no date, EFB (O’Mahony, 1929a); near the base of Torc Mountain 

(V98), 2 in a much decayed fir-log, vi.1919, OEJ (Janson, 1920); Derrycunihy Wood (V9181), 

18.viii.1990, KNAA; Muckross Woods, 2.ix.2011, 4.ix.2011, RA. 

H20 Wicklow: Powerscourt Demesne, 2 under Pseudotsuga bark, Sept/Oct, 1990, Good & Butler (1995); 

Ballard Wood, Glendalough (T1495), 14.vi.1993, KNAA. 

H21 Dublin: Glendhu, Heasler (O’Mahony, 1929a). 

H30 Cavan: Cloverhill, Nicholson (O’Mahony, 1929a). 

Quedius truncicola Fairmaire & Laboulbène, 1856 syn. ventralis (Aragona, 1830) – Rare 

In wet, very rotten timber, often with dense, clay-like blackish mould, in tree holes and hollow trunks, 

generally beneath bird nests; also reported from rotten fungi and at sap. 

H21 Dublin: Portmarnock, var rufipennis, two, 21.ix.1925, E. O’Mahony, det. PMH (Nash, Anderson & 

O’Connor, 1997). 

40


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FAMILY LUCANIDAE Stag Beetles 

Sinodendron cylindricum (Linnaeus, 1758) Rhinoceros Beetle – Widespread 

Bores in decaying heartwood of large broad-leaved trees, and also pine, including stumps. Irish hosts 

include alder, birch, oak and sweet chestnut (Speight, 1985). Has been reported from floating timber 

in both freshwater and seawater. 

H1 Kerry South: remains in ash stump in large pine wood near Glenbeigh (V69), vi.1902 

(Donisthorpe, 1903) (Bouskell, 1904). 

H2 North Kerry: Muckross Demesne, June 1924, OE Janson (Janson & Bonaparte-Wyse 1924); 

Reenadinna (V9586), 28.iv.2002, MC. 

H10 North Tipperary: Ballyvillane (R8681), 24.iv.2002, MC. 

H14 Laois: Abbeyleix, 1901, JMB (Johnson & Halbert, 1902). 

H15 South-east Galway: Clarinbridge (M4119), 20.x.1996, MC; Garryland Wood, 24.ix.2007, RA. 

H20 Wicklow: Bray, GCC; Powerscourt, 1900, JSS (Johnson & Halbert, 1902); Powerscourt Deerpark 

(O1912), in decayed trees, 9.vi.1919, Halbert (1922); 15-19.vi.1993, KNAA. 

H21 Dublin: Dollymount, 1900, JSS (Johnson &Halbert 1902); Killiney Hill, pair, 23.v.1983, leg. R 

Mulcahy, det JAG (Good, 2002); Waterstown Park, 21.v.2011, RA. 

H23 Westmeath: Hare Island in Lough Ree, Halbert (1900) (Johnson &Halbert 1902); Crookedwood 

(N4662), 25.iv.1996, MC. 

H25 Roscommon: Mote Park (M86), on willow, Halbert (1898a; Johnson & Halbert, 1902); Rindoon 

Wood (N05), adults & larvae in moist rotten wood in dead trunks of alder and ash, JAG (Good, 2002); 

galleries in dead ash, 14.iv.2010, KNAA. St John’s Wood (M9956), many dates, 2010, KNAA. 

H28 Sligo: Rathcarrick (G6334), 18.ii.1993, MC; Templehouse (G6218), remains in faeces (?pine 

marten), 23.vi.1993, MC; Union Wood (G680285), 14.vi.1999, KNAA. 

H29 Leitrim: Curraun (H103006), vi.2004, TG. 

H30 Cavan: Lough Oughter, 1893, WFJ (Johnson, Halbert & Carpenter, 1894). 

H33 Fermanagh: Tempo, C Langham (Langham, 1897; Johnson &Halbert 1902); Marble Arch, 

24.vi.1988, DAL; Inishfendra, Corlatt Wood, Reilly Wood, etc, Crom Castle Estate (H32), 1992, KNAA 

& APF, 4.xi.2002 & 6.vii.2007, KNAA; Castle Coole Estate (H2643), 2.vi.1992, KNAA; many dates, 2006, 

KNAA; Filbert Hill, Castle Coole, 26.vi.2010, RA; Rossyvullan Hil, Castle Coole, 28.v.2010, RA; 

Florencecourt, 21.vi.2010, RA. 

41


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H36 Tyrone: Caledon Deer Park (H7543), many dates, 2006, KNAA. 

H37 Armagh: Tynan Abbey Estate, 21.vi.2011, 1.vii.2011, 26.vii.2011, RA. 

H39 Antrim: Portmore Lough, 8.x.2009, RA; Shane’s Castle, 1.vii.2010, RA. 

FAMILY CLAMBIDAE 

Clambus pallidulus Reitter, 1911 – Rare 

42 

Figure 12: Sinodendron cylindricum © Keith Alexander 

Has been reported in Britain from a hollow apple tree, in debris in a rotten elm stump, and from moss 

among rotten logs, implying a saproxylic association. 

H2 North Kerry: Killarney district, identified at the time as Clambus minutus, male & female, ii.1918; 

male, Flesk, iv.1939, EFB (Johnson, 1997). 

FAMILY SCIRTIDAE 

Prionocyphon serricornis (Müller, P.W.J., 1821) – Uncommon 

Develops in water-logged hollows in old trees, especially beech, and including external hollows 

amongst roots; larvae aquatic, feed on detritus from dead leaves; adults active fliers, short-lived. 

‘Appears to be common on beech trees just about everywhere here but has been completely 

overlooked (by me) until now. I have now looked in six locations and found it in five.’ (RA, Jan. 2008). 

These latter finds (also below) refer to larvae only, although adults are relatively easy to rear. 

Assessed as Data Deficient in Foster et al. (2009).


____________________________ 

Figure 13: Prionocyphon serricornis larva © Roy Anderson 

H20 Wicklow: Powerscourt Deerpark, a male, viii.1927, AWS (O’Mahony, 1927, 1928a); two larvae & 

some exuvial fragments in tree holes of mature beeches beside the new club-house of the golf course, 

5.xii.2007, GNF & JD Reynolds (Foster & Reynolds, 2008). 

H33 Fermanagh: Finlane, Florencecourt, 6.iii.2011, pools on stand of beeches, BN, GNF, MDB, RA. 

H36 Tyrone: Drum Manor (H757776), larvae on beech, 19.i.2008, RA (Anderson, 2008a). 

H37 Armagh: Slieve Gullion Forest (J039195), 8.iii.2012, MDB/RA. 

H38 Down: Minnowburn Beeches (J326686), larvae plus numerous exuviae in several pools on beech 

root bases, 10.xii.2007, RA, (Anderson, 2008a); Killynether Wood, 2.ii.2008, RA; Temple of the Winds, 

Mount Stewart, 8.iii.2008, RA; Donard Demesne (J373302), 31.xii.2011, RA. 

H39 Antrim: Barnett’s Park (J323686), one small colony on a large beech, 16.xii.2007, RA (Anderson, 

2008a); Donard Demesne, 31.xii.2011; Lady Dixon Park, Belfast, 17.i.2008, RA. 

H40 Londonderry: Springhill Estate (H866824), several colonies on mature beech, 17.xii.2007, RA 

(Anderson, 2008a). 

FAMILY EUCNEMIDAE False Click Beetles 

Melasis buprestoides (Linnaeus, 1761) – Rare old growth species 

Develops in rather hard dead timber, especially boughs, of a wide variety of broad-leaved trees; in 

standing & fallen timber. SW Ireland only, although more widespread in Britain where scarcest in the 

west. 

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H1 South Kerry: Lickeen Wood, Glencar (V78), one female on a dead portion of a standing oak, 

9.vii.1924, Janson & Bonaparte-Wyse (1924). 

H3 West Cork: Glengarriff Woods (V920565), one on bracken under an oak near Lady Bantry’s 

Lookout, 12.vii.1985, JPOC. & MAOC (O’Connor & O’Connor 1987). 

FAMILY ELATERIDAE Click Beetles 

Calambus bipustulatus (Linnaeus, 1767) – Extinct 

Larvae a predator, living in relatively soft rotten wood in stumps, trunks and hanging boughs of oak 

& other broadleaves; adult reported nectaring at blackthorn and hogweed in daylight. 

H6 Waterford: H. Tempest took it near Waterford; specimen in NMI (Johnson & Halbert, 1902). 

Denticollis linearis (Linnaeus, 1758) – Common & Widespread 

Larvae develop under bark on dead wood and in decaying heartwood; broad-leaved trees and pine; 

omnivorous, feeding on live larvae as well as dead phloem tissues, etc. Also develops on moorlands, 

where larvae are active in the upper peat and moss layers. ‘Very local’ in Ireland (Johnson & Halbert, 

1902) although more widespread than then appreciated once the larvae are recognised. H1, H2, H3, 

H9, H20, H25, H33, H34, H36, H38, H39, H40. 

Ampedus balteatus (Linnaeus, 1758) –- Doubtful 

44 

Figure 14: Denticollis linearis © Roy Anderson 

Develops in red-rotten heartwood of stumps and boughs of various trees. The Rathlin record lacks 

credibility as many other records on Hardy’s list have also been rejected. The Kerry specimens have


____________________________ 

not been seen by a competent authority apart from Hardy. It seems unlikely that such a distinctive 

species could have been overlooked for so long. 

H1/2 Kerry: J.R. Hardy, in litt. (Halbert & Johnson, 1902). 

H39 Antrim: Rathlin Island, at roots of broom, JRH (Hardy, 1897) (Halbert & Johnson 1902).] 

Ampedus pomonae (Stephens, 1830) – Rare 

Primarily known from the Glencar area of Co. Kerry, where it has been found developing in soft red- 

rotten heartwood of birch on peatland. Its recent discovery in County Clare suggests that it may be 

more widespread than was once supposed. 

H1 South Kerry: Glencar, several taken, vi.1902 (Donisthorpe, 1903); ‘Ampedus praeustus specimens’ on 

a road, HSJD (Donisthorpe, 1917); 29.vi.1934, AWS, det. MCDS, confirmed J. Chassain. (Speight, 

1989b); Caragh Lough (V7190), 1989 (Mendel, 1990; Mendel & Clarke, 1996); adult and three larvae in 

birch logs, bog and fens at south end, 18.v.1994, JAO (Owen, 1997). 

H9 Clare: Ballyogan Lough (R370912), rich fen, v.vi.2007, & Fir Lough (R630832), reedswamp by area 

cutover bog, vii.viii.2007, in Malaise traps, TG. 

Ampedus pomorum (Herbst, 1784) – Uncommon 

Larvae develop in decayed timber of oak, birch, pine & probably other trees; primarily associated with 

birches around peatlands, but also in areas of ancient wood pasture. 

H9 Clare: bog near Finlough, coll. F. Neale (Halbert, 1896; Johnson & Halbert, 1902). 

H10 North Tipperary: near Rathcabban (N0007), dry peat bog with birch trees, 9.vii.1996, JAO (Owen, 

1997). 

H13 Carlow: Borris, PF (Johnson & Halbert, 1902). 

H14 Laois: Abbeyleix, JMB (Browne, 1901; Johnson & Halbert, 1902). 

H15 South-east Galway: Portumna Bay, The Quay, Park, etc (M841032), v. 2003 (Regan & Anderson, 

2004). 

H16 North-east Galway: Clonbrock (M7439), JNH (Halbert, 1896; Johnson & Halbert, 1902). 

H18 Offaly: Seagull Bog, near Tullamore, JNH (Halbert, 1895c; Johnson & Halbert, 1902); All Saints 

Bog, 1988, MCDS (Speight, 1990). 

H19 Kildare: Prosperous Bog, 1935, EO (O’Mahony, 1935b). 

H20 Wicklow: Powerscourt “Elater sanguineus” “among peat earth, in rotten branch of oak tree, 

17.ii.1849, ARF (Haliday ms list) (Johnson & Halbert, 1902); Powerscourt Deerpark, two “Elater 

45


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praeustus” dug out of an old alder stump by River Dargle, JNH (Halbert, 1922) – the museum 

specimen at NMI is A. pomorum (Speight, 1989b). 

H22 Meath: Balrath, vi.1916, GWN (Nicholson, 1917); Carlanstown, vi.1916, GWN (Nicholson, 1917). 

H30 Cavan: Cloverhill, v.1914, GWN (Nicholson, 1915). 

H34/40 East Donegal & Londonderry: Kilderry, CWB (Buckle, 1900; Johnson & Halbert, 1902). 

H37 Armagh: Churchill, 1887, WFJ (Johnson, 1888a; Johnson & Halbert, 1902); Derryadd, Peatlands 

Park, 18.v.1992, 21.v.2007, 15.iv.2010, RA; Peatlands Park, 21.v.2007, RA; Argory Moss (H877575), 

26.iv.2012, MDB. 

H39 Antrim: Garry Bog (9328/30), two off sallow bush at edge raised bog, 7.vi.1992; Great Deer Park, 

Glenarm (D2911), in dead oak branch, 20.v. & 13.vi.2006, KNAA. 

H40 Londonderry: Iniscarn Forest, Moyola Park, 17.v.1985, RA; Kennedy’s Quarry, Macosquin, 

6.vi.1992, RA; Ballynahone More Bog, 28.v.2007, RA. 

46 

Figure 15: Ampedus pomorum © Roy Anderson 

Melanotus castanipes (Paykull, 1800) – Common & Widespread 

Larvae most frequently develop in red-rotted heartwood, but also in decaying wood generally. The 

adult flies after dark and is attracted to light. Included within M. villosus (Fourcroy, 1785) in Britain 

and Ireland until separated by Mendel (2004) who comments he has M. castanipes from the west of


____________________________ 

Ireland while M. villosus predominates in SE England. H1, H2, H13, H15, H16, H18, H20, H25, H30, 

H31, H33, H34, H36, H37, H38, H38, H39, H40. 

FAMILY LYCIDAE Net-winged Beetles 

Figure 16: Melanotus castanipes © Roy Anderson 

Pyropterus nigroruber (De Geer, 1774) – Rare old growth species 

Larvae develop in well-decayed red-rotten heartwood of various trees, especially birch and beech; 

adults attracted to blossom and often found sunning in woodland glades. Vulnerable in Ireland 

(Speight, 1990). Montane and northern in continental Europe. 

H2 North Kerry: Killarney, summer 1866 JRH (Rye, 1867); several taken by beating birch trees and by 

sweeping, July 1898, JRH & GWC (Halbert, 1898c); Ross Castle, Killarney, swept off birch, GWC 

(Johnson & Halbert, 1902); Flesk, Killarney, 6.x.1930, EFB; Muckross, ix.1953, EFB (Speight, 1990) – 

both very late dates; Muckross Peninsula (V9586), larvae collected from red-rotted beech log in 

ash/oak woods on sandstone, ii.1989, hatched 5-15.v.1989, MCDS (Speight, 1990); Cromaglan, 1907, 

AWS (Halbert, 1907a). 

H28 Sligo: Slish Wood (G7432), one beaten from vegetation along lakeshore, 16.vii.1992, MC (Cawley, 

2005). 

47


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FAMILY CANTHARIDAE Soldier Beetles 

Larvae of Malthininae are thought to develop in decaying branchwood or heartwood, although the 

larvae of some Malthodes have been found as active predators over branch and trunk surfaces of living 

trees. 

Malthinus balteatus Suffrian, 1851 – Rare, requires confirmation as a native 

Associated with poorly-drained broad-leaved woodland, particularly wooded streamsides, but also 

along base of wooded limestone escarpments. Strangely just a single report from Ireland, which may 

suggest a misidentifcation. Current recording activity suggests that it is absent from Ireland. 

H2 North Kerry: Muckross Demesne, one beaten from whitethorn near the lake shore, 29.vi.1924, OEJ 

(Janson & Bonaparte-Wyse, 1924). 

Malthinus flaveolus (Herbst, 1786) – Widespread 

Widespread in broad-leaved woodland and hedgerows. The lack of records from the southern 

counties is peculiar, but it does appear much more localised than it does in Britain. 

H20 Wicklow: Glen of the Downs (O2611), 19.vi.1993, KNAA. 

H21 Dublin: Barnacullia, vi.1892, HKGC (Cuthbert, 1892a). 

H25 Roscommon: St John’s Wood (M9956), 9.vi.2010, KNAA. 

H27 West Mayo: Belclare, 1910-1912, JNH (Johnson & Halbert, 1912). 

H33 Fermanagh: Portora, Enniskillen, 19.v.1946, UM (coll. unknown). Castle Coole Park (H2643), 

17.vi.2006, KNAA. 

H37 Armagh: Mullinure, Armagh, WFJ (Johnson, 1892a); Argory Moss (H881578), 1.vii.1992, KNAA. 

H38 Down: Greencastle Td., 31.viii.1946, UM (coll. unknown); Belvoir Forest, 5.vii.1993, RA; Shimna 

R., Newcastle, 29.vii.1995, RA; Rostrevor Oakwood NNR (J186173), two swept, 11.vii.2007, KNAA; 

Castle Ward Park (J5649), 20.viii.2006, KNAA. 

H39 Antrim: Farr’s Bay NR, Randalstown, 4.vii.1986, RCW (Welch, 1986); Newforge, Belfast, 

15.vi.1994, RA; Breen Wood NNR, 13.vii.2005, RA; Shane’s Castle, 1.vii.2010, RA. 

H40 Londonderry: The Umbra NR, Magilligan, 5.vii.2005, RA. Ness Wood ASSI (C528118), 7.vii.2007, 

KNAA. 

Malthinus seriepunctatus Kiesenwetter, 1852 – Uncommon 

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Common in the south-east of Ireland (Johnson & Halbert, 1902), and widespread but local elsewhere; 

becomes more coastal in distribution in the north, as it does in Britain. 

H3 West Cork: Glandore (W23), JNH (Johnson & Halbert, 1902). 

H4 Mid Cork: Shippool Woods, Bandon River (W5757), 23.v.1994; Rock Castle Woods, Bandon River 

(W5653), 6.vi.1994; Rineen Wood (W190338), 12.vi.1996, KNAA. 

H6 Waterford: Glenshelane Valley & Lismore, JNH (Johnson & Halbert, 1902); Ballyeelinan (X2181), 

10.viii.1990, KNAA. 

H11 Kilkenny: Thomastown, JNH (Johnson & Halbert, 1902). 

H12 Wexford: Wexford district, JNH (Johnson & Halbert, 1902). 

H20 Wicklow: Bray, 1904, WFJ; Vale of Clara, vii.1919, JNH (Halbert, 1937); Glen of the Downs, 14- 

19.vi.1993; Deputy’s Pass (T2390), 17.vi.1993; Powerscourt Deerpark (O1912), 15.19.vi.1993, KNAA. 

H31 Louth: south, JNH (Halbert, 1900b) (Johnson & Halbert, 1902). 

H33 Fermanagh: Correl Glen, 5.viii.1993, RA. 

H38 Down: Rostrevor Oakwood NNR (J186173), swept in numbers, 11.vii.2007, KNAA; Dundrum 

Coastal Path, 30.vi.2009, RA; Mount Stewart, 11.viii.2010, RA. 

H39 Antrim: Glenarm NR, 14.vi.2011, RA. 

Malthodes dispar (Germar, 1824) – Rare 

A species of wet woodlands and shaded riverbanks. 

H2 North Kerry: Torc Wood, GWC, det. JNH (Johnson & Halbert, 1902). 

H6 Waterford: Lismore (X09), 5.vii.1922, OEJ (Janson et al., 1923; O’Meara, 2008). 

Malthodes flavoguttatus Kiesenwetter – Uncommon 

Most frequent in acidic oak and birch woodlands of hill country districts. 

H1 South Kerry: Cloonee Loughs, Kenmare, JNH (Halbert, 1898c) (Johnson & Halbert, 1902); Tomies 

Wood, Lough Leane (V9188), 26.v.1994, KNAA. 

H2 North Kerry: Killarney, vii.1927, JNH (Halbert, 1937). 

H3 West Cork: Glengarriff Woods (V918570), vii.1924, JNH (Halbert, 1937); 24.v.1994 & 13.vi.1996, 

KNAA. 

H4 Cork Mid: Inchy Bridge & Ummera (W466457), 14 & 15.vi.1996, KNAA. 

H6 Waterford: Glenshelane Valley near Cappoquin, JNH (Johnson & Halbert, 1902). 

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H20 Wicklow: Clara Oak Wood, emerged from oak sticks collected by AWS, 22.iv.1934, by 4.vi.1934; 

teste P. Harwood; very local and new to Co. Wicklow (O’Mahony, 1935a); Glendalough Oakwood 

(T1296), 13.vi.1993, KNAA. 

H27 West Mayo: Croagh Patrick, 1910-1912, JNH (Johnson & Halbert, 1912). 

H28 Leitrim: Truskmore, 19.vii.2008, RA. 

H33 Fermanagh: Carrickreagh Bay, Lough Erne, 11.vi.1941, anon., UM. 

H34 East Donegal: Ardnamona Wood, Lough Eske (G9684), 10.vi.1999, KNAA. 

H35 West Donegal: Lough Gartan (C0616), 13.vi.1992, KNAA. 

H38 Down: Slieve Donard (J3529), 8.vii.1992, KNAA; Altataggart Mountain, 9.vii.2005, RA. 

H39 Antrim: Carr’s Glen, CWB det. JNH (Johnson & Halbert, 1902); Murlough Bay (D1941), 

15.vi.1992, KNAA. 

Malthodes fuscus (Waltl, 1838) – Uncommon 

Most frequent in acidic oak and birch woodlands in hill country districts. ‘Malthodes pellucidus’ in 

Johnson & Halbert (1902). 

H1 South Kerry: Kenmare, 1898, JNH (Halbert, 1898c). 

H2 North Kerry: Derrycunihy Wood, GWC det. JNH (Johnson & Halbert, 1902); Torc Woods, 

Killarney (V968850), 6.vi.1994, KNAA. 

H4 Mid Cork: Shippool Woods, Bandon River (W5757), 23.v. & Rock Castle Woods, Bandon River 

(W5653), 6.vi.1994, KNAA. 

H6 Waterford: Glenshelane Valley, JNH (Johnson & Halbert, 1902). 

H11 Kilkenny: Woodstock, JNH (Johnson & Halbert, 1902). 

H20 Wicklow: Devil’s Glen (T2499), 17.vi.1993, KNAA; Deputy’s Pass (T2390), 17.vi.1993, KNAA; 

Glendalough Oakwood (T1296), 13.vi.1993, KNAA. 

H27 Mayo West: Belclare, 1910-1912, JNH (Johnson & Halbert, 1912). 

H28 Sligo: Slish Wood (G740317), 12-17.vi.1999; Union Wood (G680285), 14.vi.1999, KNAA; Sriff 

Wood (G793341), 17.vi.1999, KNAA. 

H33 Fermanagh: Florencecourt, 21.vi.2010, RA. 

H34 East Donegal: Ballyarr Wood (C1820), 14.vi.1992, KNAA. 

H38 Down: Rostrevor Oakwood NNR (J1817), 11.vii.2007, KNAA. 

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H39 Antrim: Carr’s Glen, CWB, det. JNH (Johnson & Halbert, 1902); Clare Forest, Ballycastle, 

1.vii.1986, RCW; Breen Wood (D1233), 7.vi.1992 & 9.vii.2007, KNAA; Fair Head (D1843), 11.vi.1992, 

KNAA; Murlough Bay (D1941), 16.vi.1992, KNAA; Collin Glen, vii.1992, RA. 

H40 Londonderry: Ness Wood ASSI (C528118), 7 & 8.vii.2007, KNAA. 

Malthodes guttifer Kiesenwetter, 1852 – Uncommon 

First recorded in Ireland by Johnson & Halbert (1902) as M. mysticus (see Halbert, 1937) then later by 

O’Mahony (1935a); now known to be relatively widespread. In damp woodland situations. 

H2 North Kerry: Killarney, vii.1927, JNH (Halbert, 1937). Torc Woods (V968850), 6.vi.1994, KNAA. 

H3 West Cork: Glengarriff Castle Estate (V9455), in Malaise trap, 2005, TG det. KNAA. 

H7 South Tipperary: Clonmel, vii.1928, JNH (Halbert, 1937). 

H15 South-east Galway: Coole & Garryland (M4304), 4.vi.1994, KNAA. 

H20 Wicklow: Arklow, vii.1926, JNH (1937); Clara Oak Wood, collected by AWS on 22.iv.1934, 

emerged from oak sticks by 4.vi.1934; det. KGB (O’Mahony, 1935a). 

H23 Westmeath: Belclare, M. mysticus, JNH (Clare Island Report, part 28; O’Mahony, 1929a) re-det. as 

M. guttifer (Halbert, 1937). 

H25 Roscommon: St John’s Wood (N0056), 6.viii.2010, KNAA. 

H27 Mayo West: Belclare, recorded as M. mysticus (Halbert, 1937). 

H28 Sligo: Hazelwood, L. Gill, 20.vii.2008, RA. 

H33 Fermanagh: Brougher Mountain, 9.vi.2006, RA; Correl Glen NNR (H074546), RA; 5.vii.2007, 

KNAA; Rossyvullan Hill, Castle Coole, 28.v.2010, RA. 

H38 Down: Slieve Donard, lower slopes (J3529), 9.vii.1992, KNAA; Belvoir Park, 4.vii.1993, RA; 

Rostrevor Oakwood NNR (J186173), 11.vii.2007, KNAA. 

H39 Antrim; Shane’s Castle, 3.vi.1995, RA; Farr’s Bay NNR, 4.vii.1986, RCW (Welch, 1986); Breen 

Wood (D123336), 13.vii.2005, RA & 9.vii.2007, KNAA; Glenarm Great Deer Park (D2911), 29.vii.2006, 

KNAA; Glenarm NR, 14.vi.2011, RA. 

H40 Londonderry: Banagher Glen (C670045), 14.vii.2005, RA & 8.vii.2007, KNAA; Ness Wood ASSI 

(C528118), 8.vii.2007, KNAA; The Umbra, 1.vi.2011, RA. 

Malthodes marginatus (Latreille, 1806) – Common & Widespread 

Larvae develop in decaying wood or beneath bark on dead timber; mainly predatory on insect larvae 

including dead ones, but will also feed on decaying timber to some extent. Common in Ireland 

51


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(Johnson & Halbert, 1902): Ulster, Connaught, Leinster, Munster; H1, H2, H3, H4, H9, H15, H16, H18, 

H20, H21, H23, H24, H25, H27, H28, H33, H34, H35, H37, H38, H39, H40. 

Malthodes minimus (Linnaeus, 1758) – Doubtful 

Irish status unclear: “these records require verification, as there may have been some confusion 

with…” M. pumilus (Johnson & Halbert, 1902). Anderson et al. (1997) list M. minimus without 

comment; no authenticated Irish specimens. No modern reports. 

H2 North Kerry: Killarney, old Kenmare Road, Furlong ms. (Johnson & Halbert, 1902). 

H21 Dublin: taken by Mr Tardy (Halbert 1854) (Johnson & Halbert, 1902). 

H38 Down: near Belfast, Halbert (1885) (Johnson & Halbert, 1902).] 

Malthodes pumilus (Brébisson, 1835) – Uncommon 

Usually found by sweeping tall grasses beneath mature and older trees, in open parkland as well as 

closed canopy high forest. M. atomus (Johnson & Halbert, 1902). 

H2 North Kerry: Muckross Demesne, June 1924, OEJ (Janson & Bonaparte-Wyse, 1924). 

H9 Clare: Cratloe (Johnson & Halbert, 1902). 

H11 Kilkenny: (Johnson & Halbert, 1902). 

H12 Wexford: (Johnson & Halbert, 1902). 

H20 Wicklow: (Johnson & Halbert, 1902). 

H21 Dublin: Santry, JNH (Halbert, 1894); Lambay Island, vi-x.1906, JNH (Halbert, 1907b). 

H27 Mayo West: Westport, 1910-1912, JNH (Johnson & Halbert, 1912). 

H28 Sligo: Slish Wood (G740317), swept in numbers along wooded lakeside, 12-17.vi.1999, KNAA. 


H33 Fermanagh: Inishfendra, Crom, FIT, 1992, APF. Castle Coole Park (H2643), 17.vi.2006, KNAA. 

H34/40: Foyle District (Buckle, 1900) (Johnson & Halbert, 1902). 

H36 Tyrone: Baronscourt Park (H3682), 15.vi.2006, KNAA. 

H38 Down: (Johnson & Halbert, 1902); Ballymacormick Point, 21.v.2007, MGT. 

H39 Antrim: (Johnson & Halbert, 1902); Great Deer Park, Glenarm (D2911), one swept from shady 

woodland along river, 13.vi.2006, KNAA. 

H40 Londonderry: Drenagh Estate (C6923), 14.vi.2006, KNAA. 

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FAMILY BOSTRICHIDAE False Powder-post Beetles 

Develop in dead hard timber continuously until interior reduced to powder. 

Lyctus linearis (Goeze, 1777) – Rare 

Usually found on fresh oak palings; it develops in dead sapwood of oak, beech & ash. Subfossil 

remains have been found in Viking Dublin (Reilly, 2003) 

H1 South Kerry: Kenmare, Lyctus canaliculatus swept amongst herbage by JRH, (Halbert, 1898c; 

Johnson & Halbert, 1902). 

FAMILY ANOBIIDAE 

Hedobia imperialis (Linnaeus, 1767) – Rare 

Has been reared from dead stems of Rosa and hawthorn. 

H1 South Kerry: Kenmare, in a thorn fence at Carrigacappeen, JRH (Halbert, 1898; Johnson & Halbert, 

1902). 

H15 South-east Galway: Portuma Forest Park, 1 in Piptoporus on birch, 23.v.2003, G.M. Orledge, det. 

M. Barclay. 

Ptinus subpilosus Sturm, 1837 – Rare old growth species 

A spider beetle, living in old hollow trees and under loose flaking dead bark on tree trunks, mainly of 

oak, but also old pines in Scottish Highlands; also occasionally found as a scavenger in bird nests. 

Relict old forest species in Britain; in Ireland it may be a relict old forest species surviving in a few 

historic demesnes. 

H39 Down: Belvoir Park Forest (J336695), a female from under flaking bark of an old sycamore, 

25.xi.1992, RA; same locality, 2 males 2 females on 2.iii and 5.iii.2007, RA (Anderson, 1994); same 

locality, 10 in dry crevices with spiders webs, 7.ii.2012, MDB/RA. 

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54 

Figure 17: Ptinus subpilosus © Roy Anderson 

Grynobius planus (Fabricius, 1787) – Common & Widespread 

Develops in dead heartwood of various broad-leaved trees. Locally common in old trees: H1, H2, H3, 

H6, H9, H20, H21, H25, H26, H27, H28, H33, H35, H36, H37, H38, H39, H40. 

Figure 18: Grynobius planus © Roy Anderson 

Dryophilus pusillus (Gyllenhal, 1808) – Established introduction 

In wood and debris of pine and larch; locally common on conifers and evidently extending its range in 

Ireland (Halbert, 1937a). H6, H13, H19, H20, H21, H22, H30, H33, H37, H38, H39. 

Ochina ptinoides (Marsham, 1802) – Uncommon 

Develops in dead thick stems of ivy on trees. Ochina hederae in Donegal, Antrim, Down, Armagh, 

Galway, Meath, Dublin, Wicklow, Kilkenny, Waterford, Kerry (Johnson & Halbert, 1902).


____________________________ 

H2 North Kerry: near the base of Torc Mountain (V98), in plenty by beating holly and ivy, 1919, OEJ 

(Janson, 1920). 

H4 Mid Cork: Charleville (R5322), from Taxus in graveyard, 8.ix.2005, MC. 

H7 South Tipperary: Clonmel (S200224), from Taxus in graveyard, 7.vi.2007, MC. 

H18 Offaly: Charleville Castle Park (N3123), 16.vi.1993, KNAA. 

H21 Dublin: Clontarf, 1928. EO (O’Mahony, 1928b). 

H22 Meath: Laytown sandhills, 16.vi.1894, GHC, det. JNH (Halbert, 1895b). 

H25 Roscommon: St John’s Wood (M9956 & N0056), 6.viii.2010, KNAA. 

H33: Fermanagh: Castle Coole Park (H2643), 17.vi.2006; Inisherk, Crom Castle, 6.vii.2007, KNAA. 

H37 Armagh: Loughgall Demesne, sweeping, WFJ (Johnson, 1892a). 

H38 Down: Cairnshill, Belfast, 23.vii.1987; Newforge, Belfast, 15.vi.2004, RA; Rostrevor Oakwood 

NNR (J186173), 9.vii.2005, RA & 11.vii.2007, KNAA. 

H39 Antrim: Farr’s Bay NNR, 4.vii.1986, RCW (Welch, 1986). 

H40 Londonderry: Drenagh Estate (C6923), 14.vi.2006, KNAA. 

Xestobium rufovillosum (De Geer, 1774) Deathwatch Beetle – Established Introduction 

Bores in hard dead heartwood of several hardwood species where damp and fungal decay is present - 

in building timbers one fungus in particular Donkioporia expansa may be especially important; larval 

period anything between 1 and 13 years, usually 3-7; wings well-developed, but flight very rarely 

recorded, and unlikely to colonise buildings naturally. Flight occurs only when temperatures exceed 

17°C and flying beetles are attracted to light. Very rare in Ireland (Hickin, 1953) and only known from 

building timbers. 

H7 South Tipperary: Knocklofty, evidence of this beetle attacking timber behind plaster in a house, 

25.ii.1953, L. Brooks (Hickin, 1953). 

H38 Down: Cregagh, Belfast, 31.iii.1994, in the wooden handle of a corkscrew originally from 

Germany, “the loud ticking was, as might be imagined, mysterious”, Robert Nash, UM. 

Ernobius mollis (Linnaeus, 1758) – Established Introduction 

Develops in dead branches of softwoods, larvae consuming the bark but scoring sapwood, i.e. a 

cambium feeder. Indigenous to north Temperate regions, being common in northern Europe, 

especially in Scandinavia; widespread in Britain & Ireland; introduction in many parts of World. 

H1/2 Kerry: Furlong ms. (Johnson & Halbert, 1902). 

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H21 Dublin: Clontarf, 1928, vi.1935, EO (O’Mahony, 1928, 1935). 

H7 South Tipperary: Thomastown Plantation, 4.vii.2008, RM det. RA. 

H38 Down: Altataggart Mountain, Mournes, on Scots Pine, 9.vii.2005, RA. 

H39 Antrim: Greenmount Agricultural College, in dying Picea abies by pond, 30.vi.1986, RCW (Welch, 

1986). 

H40 Londonderry: The Umbra, swept off Pinus nigra, 5.vii.2005, RA. 

Anobium inexpectatum Lohse, 1954 – Rare, possibly overlooked 

Develops in dead ivy stems. Very similar to A. punctatum but the aedeagus and ovipositor styles differ 

– only recently distinguished. One recent Irish record - suggests an overlooked native. 

H1 South Kerry: Uragh Wood, Inchiquin Lough (V829632), 1 male fogged from oakwood, RM det. 

RA. 

Anobium punctatum (De Geer, 1774) Furniture Beetle – Widespread 

Eggs laid in a crack or groove in exposed dead sapwood of living and dead trees, larvae bore 

primarily within sapwood, and pupate just beneath outer surface. Anobium domesticum is ‘common in 

old houses: Ulster, Connaught, Leinster, Munster’ (Johnson & Halbert, 1902) – few reports of wild- 

living beetles. Subfossil remains have been found in Viking Dublin (Reilly 2003) 

H6 Waterford: in buildings at two locations (O’Meara, 2008). 

H9 Clare: Inishmaan, Aran Islands, in domestic dwelling, 28.v.1997, RA. 

H10 North Tipperary: Castlelough (R7483), loughside vegetation, 10.vii.1996, JAO (Owen, 1997). 

H33 Fermanagh: Castle Coole Park (H2643), 2.vi.1992 & 26.vii.2006, KNAA. 

H35 West Donegal: Ardara, WFJ (Johnson, 1893). 

H36 Tyrone: Baronscourt Park (H3682), 25.vii.2006; Caledon Deer Park (H7543), 26.vii. & 16.viii.2006, 

KNAA. 

H37 Armagh: Tynan Abbey Estate, under fungoid bark on beech, RA. 

H38 Down: Belfast Docks, corn stores, 1942-46, AFO (O’Farrell & Butler, 1948); Copeland Bird 

Observatory Island, 1971-1973, CAMR; Derrymore House Park, Bessbrook (J0528), 6.vii.1992, KNAA; 

Terrace Hill, Ballynahatty, swept off winter barley, 7.viii.2006, RA; Hillsborough Forest, swept off 

Douglas fir, 3.viii.2006, RA; Castle Ward Park (J5649), 30.vii. & 20.viii.2006, KNAA. 

H39 Antrim: Whitepark Bay ASSI, on walls of hostel, 27.vi.2007, MGT. 

H40 Londonderry: Drenagh Estate (C6923), 28.vii.2006, KNAA. 

56


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Ptilinus pectinicornis (Linnaeus, 1758) – Uncommon 

Bores in exposed dry heartwood of old broad-leaved trees, making small pinholes; especially in beech, 

but also in most other species. 

H1 South Kerry: Tomies Wood (V9089), 26.v.1994, KNAA. 

H7 South Tipperary: Thomastown Plantation (S214209), 4.vii.2008, RM det. RA; Shanbally Castle 

Estate (R977165), 17.iii.2012, RA. 

H14 Laois: Emo Park, vi.1938, AWS (O’Mahony, 1938). 

H21 Dublin: College Park, ARH (Hogan, 1854); Artane, in an old tree stump, G.P. Farran (Halbert, 

1900b; Johnson & Halbert, 1902); St Annes, 1924, EO (O’Mahony, 1924); Clontarf, Raheny, Sutton, 

1928, EO (O’Mahony, 1928b). 

H25 Roscommon: St John’s Wood (M9956), 6.viii.2010, KNAA. 

H33 Fermanagh: Tempo, C. Langham (Langham, 1897) (Johnson & Halbert, 1902). Castle Coole Park 

(H2643), 17.vi & 27.vii.2006, KNAA. 

H36 Tyrone: Baronscourt Park (H3682), on dead beech, various dates, 2006; Caledon Deer Park 

(H7543), vi. & vii.2006, KNAA. 

H37 Armagh: Tynan Abbey Estate, dead beech, 1.vii.2011, RA. 

H38 Down: Belvoir Forest, on beech trunk, 19.vi.1984, RA; Portulla Wood, Quoile, on beech trunk, 

20.vi.1984, RA; Derrymore House Park, Bessbrook (J0528), 6.vii.1992, KNAA; Mount Stewart Estate, 

19.v.1992, APF. 

H39 Antrim: Deerpark Hotel, Antrim, on scarred oak, 30.vi.1986; Farr’s Bay NNR, in partially dead 

beech, 4.vii.1986, RCW (Welch, 1986); Shane’s Castle, at Tilia flowers, 1.vii.2010, RA; Mount Stewart, 

galleries in beecxh, 13.viii.2010, RA. 

H40 Londonderry: Drenagh Estate (C6923), on dead beech, 14.vi.2006, KNAA. 

Dorcatoma dresdensis Herbst, 1792 – Rare old growth species 

In Ireland has only been found developing in the dead, persistent annual brackets of Inonotus dryadeus 

on veteran oak trees and I. radiatus on dead alder stems, but in Britain it favours dead hard perennial 

bracket fungi on broad-leaved trees, incl. Ganoderma and Phellinus spp. Although discovered very 

recently it is most likely an overlooked native. Alexander (2009) notes emergence holes in I. dryadeus 

almost certainly of this species at a number of historic parkland sites across Northern Ireland, but with 

confirmation from only Crom. 

57


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H6 Waterford: Lismore Castle (X049988), reared from I. radiatus on old dead alder stems by R. 

Blackwater, 14.iv.2012, RA. 

H25 Roscommon: St John’s Wood (M9956), occupied brackets of I. radiatus on old dead alder stems, 

7.x.2010, confirmed by rearing, KNAA (Alexander, 2011). 

H28 Sligo: Doorly Park (G707354), emerged 22.v.2011 from old bracket fungi on Alnus in carr 

woodland, collected 11.v.2011, MC. 

H29 Leitrim: Lough Allen (G964110), in old bracket fungus on Alnus, 28.v.2011, MC. 

H33 Fermanagh: Inisherk, Crom Castle Estate (H357248), occupied I. dryadeus bracket on old parkland 

oak, 6.vii.2007, confirmed by rearing, KNAA (Alexander, 2009), also 6.vii.2010, MDB. 

FAMILY LYMEXYLIDAE 

58 

Figure 19: Dorcatoma dresdensis © Roy Anderson 

Hylecoetus dermestoides (Linnaeus, 1761) – Rare old growth species 

Develops in freshly dead timber and stumps of hardwoods and softwoods; adult stage very short, 

only a few days, from early April to early July; adult beetle bores vertically into the heartwood and 

establishes networks of galleries, unbranched and curved, in which eggs are laid and the larvae 

develop; the bore dust is ejected creating distinctive piles of dust; the larvae feed on ambrosia fungus


____________________________ 

growing on walls of the galleries. Primarily SW in Ireland and modern records only from a single site 

protected as a SAC under European legislation. 

H1 South Kerry: Glencar Valley, taken out of birch and in flight, vi.1902, HSJD (Donisthorpe, 1902 & 

1903; Bouskell, 1904). 

H2 North Kerry: Killarney, EFB (O’Mahony, 1929a). 

H3 West Cork: Glengarriff Castle Demesne (V9455), male & female in malaise trap, 2005, coll. T. 

Gittings, det. KNAA (Alexander, 2010a). 

H20 Wicklow: Powerscourt Deerpark, AWS (O’Mahony, 1929a); taken in numbers on 30.v.1933, EO 

(O’Mahony, 1934). 

H21 Dublin: Glendhu, R. Heasler (O’Mahony, 1929a). 

FAMILY PHLOIOPHILIDAE 

Phloiophilus edwardsii Stephens, 1830 – Rare old growth species 

Develops in the fungus Peniophora quercina which fruits on the aerial dead lower boughs of various 

mature open-grown broad-leaved trees and shrubs, but especially oak; generally in old wood pasture 

and parkland situations; adults active in autumn only. 

H2 North Kerry: Killarney, by sweeping, ix, TVW (note on cover of Haliday’s ms List of Irish Insects 

(O’Mahony, 1929). 

H19 Kildare: Liffey bank at Celbridge, one on grass under an oak tree, x.1929, JNH (Halbert, 1937). 

H33 Fermanagh: Castle Coole Park (H2643), from dead branches on two mature parkland oaks in 

Front Park, 14.ix. & 30.x.2006, KNAA (Alexander, 2010a). 

H36 Tyrone: Baronscourt Park (H3682), one in FIT hung on veteran parkland oak, 2006, KNAA 

(Alexander, 2010a). 

FAMILY TROGOSSITIDAE Flat Beetles 

Thymalus limbatus (Fabricius, 1787) – Doubtful 

Larvae and adults live beneath loose bark on decaying broad-leaved wood, especially oak, and in the 

later stages of white-rot decay, when the heartwood is dry and soft. The reliability of this record seems 

questionable – it is not normally difficult to find when present and yet extensive searches have failed 

to find any. 

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H1 South Kerry: Kenmare, ‘taken in numbers amongst a mass of leaves and fungi’, JRH, vii.1898 

(Halbert, 1898c; Johnson & Halbert, 1902). 

FAMILY CLERIDAE Checkered Beetles 

Tillus elongatus (Linnaeus, 1758) – Doubtful 

A predator of other beetles developing in the dead heartwood of old broad-leaved trees, especially 

larvae of the anobiid Ptilinus pectinicornis. The larvae hunt nocturnally under bark and on the outside 

of the tree. The single Irish record has not been confirmed in recent years. 

H1 South Kerry: Kenmare, June, Furlong ms. (Johnson & Halbert, 1902). 

Thanasimus formicarius (Linnaeus, 1758) – Rare old growth species 

Larvae and adults feed on the larvae of wood-boring beetles. Rare in Ireland, where primarily 

associated with Holly Weevil Rhopalomesites tardyi (qv). 

H1 South Kerry: Bealalaw Bridge, Glencar (V78), a pair in cop. on old holly stump riddled with 

Rhopalomesites galleries, a short way downstream 7.v.2010, KNAA. 

H2 North Kerry: Innisfallen, Lough Leane (V98), in old hollies, between 18 & 25 July 1913, OE Janson 

(1914); Muckross Demesne, on the trunks of some very decrepit holly trees, June 1919, OE Janson 

(1920); Muckross Demesne, June 1924, OE Janson (Janson & Bonaparte-Wyse, 1924); Killarney, larva in 

BMNH coll. (Emden 1943); Upper Lake (V9081), deciduous woodland, 16.v.1994, Owen (1997). 

H9 Clare: Derrygeeha Lough (R150564), Malaise trap, vi.vii.2007, TG. 

H20 Wicklow: Powerscourt Deerpark, in a decayed holly, Furlong coll. (Johnson & Halbert, 1902); one 

swept from bracken & another sunning on old holly tree, 9.vi.1919, JNH (Halbert, 1922); a specimen 

on a log, EO (O’Mahony, 1929b). 

H21 Dublin: “on the seashore” by Mr Bulwer (Curtis, 1839); Grand Canal Docks, JNH (Halbert, 1900) 


H28 Sligo: Slish Wood (G740318) & mixed woodland, Dooney Rock (G722323), 3.vi.2011, MC. 

H35 Donegal: Ballyboyle, adult with Rhopalomesites in white-rotten heartwood of open-grown 

sycamore on eroded seaward side of drumlin, 18.xi.2009, KNAA (Alexander, 2010b). 

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FAMILY DASYTIDAE 

Dasytes aeratus Stephens, 1830 – Unclear 

Figure 20: Thanasimus formicarius © Keith Alexander 

Favours open woodland situations, especially on neutral to base-rich soils: adults usually found at 

blossom, especially on hawthorns; the larvae are carnivorous over and under bark on live trunks as 

well as deadwood, feeding on dead invertebrates - has been reared in numbers from decaying oak 

wood. A curious distribution in Ireland; easy to find when present in England, and so the lack of Irish 

records does appear real; potentially a casual introduction. 

H1 South Kerry: Kenmare, JNH (Halbert, 1898c; Johnson & Halbert, 1902). 

H11 Kilkenny: Brownstown Wood (S656299), canopy fogging, 10.vi.2008, RM det. RA. 

H19 Kildare: Maynooth, HKGC & GHC (Cuthbert & Carpenter, 1894; Johnson & Halbert, 1902). 

H21 Dublin: Santry, JNH (Halbert, 1893; Johnson & Halbert, 1902). 

FAMILY MALACHIIDAE Malachite Beetles 

Malachius bipustulatus (Linnaeus, 1758) – Scarce 

Larvae partly predatory in galleries of wood-borers, partly feeding on their excreta and larval skins; 

adults sun-loving and feed on pollen, also seen to attack and eat the beetle Dasytes aeratus in Britain. 

Very local in Ireland (Johnson & Halbert, 1902); mainly southern – southern and eastern in Britain. 

The Rathlin record of Hardy is highly suspect. 

61


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H1 South Kerry: Cloonee Loughs, Kenmare, JNH (Halbert, 1898) (Johnson & Halbert, 1902). 

H6 Waterford: Curraghmore, as Malachius viridis, LHBW (Bonaparte-Wyse, 1897). 

H14 Laois: Abbeyleix, 1901, abundant on hawthorn blossom, JMB (Johnson & Halbert, 1902). 

H18 Offaly: All Saints Bog (N0010), several in birch woodland, 1988, MCDS (Speight, 1990). 

H21 Dublin: in a demesne near Lucan, JNH (Halbert, 1892); Waterstown Park (The Dingle, O088357), 

abundant over a wide area with 7 or 8 in an average sweep in pasture by the R. Liffey, 21.v.2011, RA. 

H30 Cavan: Castle Saunderson (H4119), female ovipositing on poroid bracket fungus, 14.vi.2004, M. 

McMullan, det. Howard Fox. 

H39 Antrim: Rathlin, JRH (Hardy, 1897; Johnson & Halbert, 1902). 

Anthocomus fasciatus (Linnaeus, 1758) – Extinct 

62 

Figure 21: Malachius bipustulatus © Roy Anderson 

Larvae probably predatory in borings of anobiid beetles in deadwood; adults usually found by sweep- 

netting beneath trees or at umbel flowers. The single Irish record would be doubtful if it were not for 

the voucher specimen. 

H20 Wicklow: ‘Mr Furlong’s specimen is now in Dublin Museum’ (Johnson & Halbert, 1902).


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FAMILY ASPIDIPHORIDAE 

All of the known Aspidiphoridae breed exclusively in slime fungus spore bodies. 

Aspidiphorus orbiculatus (Gyllenhal, 1808) – Rare old growth species 

H3 West Cork: Glengarriff Castle Demesne, in Malaise trap samples, 2005, TG, det IM. 


H12 Wexford: Forth Hills, JNH (Johnson & Halbert, 1902). 

H20 Wicklow: Powerscourt Demesne, vi.1926, JNH (Halbert, 1937). 

FAMILY NITIDULIDAE Sap or Blossom Beetles 

A number of species are attracted to sap flows, especially during fermentation; at freshly cut stumps, 

dying trees attracting the attention of bark beetles and the ambrosia beetle Hylecoetus, as well as 

exudations caused by the wood-boring larva of the Goat Moth Cossus. Dispersing adults may also feed 

casually on other moist decaying organic matter. 

Epuraea aestiva (Linnaeus, 1758) – Common & Widespread 

Adults at flowers of trees and shrubs, e.g. rowan and gorse; larval habits unknown but assumed to be 

sap beetles. Common in Ireland: Ulster, Connaught, Leinster, Munster (Johnson & Halbert, 1902). H1, 

H2, H5, H9, H19, H20, H21, H27, H33, H37, H38, H39, H40. 

Epuraea angustula Sturm, 1844 – Rare old growth species 

Associated with the borings of Trypodendron bark beetles in freshly dead trunks and boughs, 

especially of birch, oak and beech. 

H21 Dublin: Clontarf, in fungi, 1928, OE (O’Mahony, 1928b). 

H33 Fermanagh: Portora, Enniskillen, 3 specimens, 1.vi.1941, anon., UM. 

H35 Donegal: Glentogher, “three in the borings of Trypodendron domesticum in a fallen beech tree” 

CWB (Buckle, 1900) (Johnson & Halbert, 1902). 

H39 Antrim: Deramore (J332696), 1 male under bark on wind-thrown oak bough, 6.iii.1996, RA. 

Epuraea biguttata (Thunberg, 1784) syn. unicolor (Olivier, 1790) – Uncommon 

At sappy birch and oak stumps, and bracket fungi; also may develop in compost where conditions 

suitable. 

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H2 North Kerry: Muckross, vi.1919, OEJ (Janson, 1920); Muckross, in slivers of oak lumber, yard, 

2.ix.2011, RA. 

H20 Wicklow: Powerscourt Demesne, E. obsoleta, iv.1925, JNH (Halbert, 1937). 

H22 Meath: Balrath, 26.ix.1913, GWN (Nicholson, 1914). 

H30 Cavan: Cloverhill, v.1914, in runs of Hylurgops, GWN (Nicholson, 1915). 

H32 Monaghan: Castle Leslie, 1 female in Polyporus squamosus on ash, 18.vii.2009, RA. 

H33 Fermanagh: Rossyvullan Hill, Castle Coole, in Polyporus squamosus on sycamore, 20.vii.2010, RA. 

H34 East Donegal: Kilderry, E. obsoleta by beating ivy, CWB (Buckle, 1900) (Johnson & Halbert, 1902). 

H36 Tyrone: Baronscourt Park (H3682), beneath bark on sappy fallen branch, 12.ix.2006, KNAA. 

H38 Down: Castlewellan, as E. obsoleta (Patterson, 1885) (Johnson & Halbert, 1902); Inch Abbey, swept 

off haw blossom, hedge, 22.v.1995, RA; Killynether Wood (J475722), in Polyporus squamosus on fallen 

sycamore, 29.vi.2012, MDB/RA. 

H39 Antrim: Shane’s Castle, E. obsoleta, vi.1902, JNH (Halbert, 1910b); Langford Lodge (J088749), 

under sappy bark on windthrown oak bough, 28.v.2012, RA. 

H40 Londonderry: Ness Wood ASSSI (C528118), beneath sappy bark of fallen & sawn oak trunk, 

8.vii.2007, KNAA. 

Epuraea distincta (Grimmer, 1841) – Recent arrival 

Associated with fruit bodies of the bracket fungus Daedaleopsis confragosa which decays dead stems of 

willow and birch. First noted in 2007 and now widely known across north-east. 

H36 Tyrone: Coalisland Canal, Cranebrook, 24.iv.2010, MDB det. RA. 

H37 Armagh: Derryadd Lough (H917603), 21.v.2007, RA (Anderson, 2007a); Brackagh Moss, 

14.iv.2010, MDB, det. RA. 

H38 Down: Rostrevor Oakwood NNR (J1817), one in FIT, 11.vii.2007, KNAA. 

H39 Antrim: Portmore L. RSPB Reserve, 10.vii.2009, RA. 

Epuraea longula Erichson, 1845 – Scarce 

Adults have been found at Goat Moth Cossus burrows, but are more regularly found at woodland 

flowers; also reported from rotten elm; larval habitat not known. 

H4 Mid Cork: Courtaparteen, on meadowsweet, streambank, 16.vii.2000, RA 

H16 North-east Galway: Clonbrock (M7439), RED, det. JNH (Johnson & Halbert, 1902). 


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H31 Louth: Bellurgan Forest (J095099), under sappy bark on fresh beech log, 22.iii.2012, RA. 

H33 Fermanagh: Inisherk, Crom, in Inonotus dryadeus on oak, 31.iii.1997, RA. 

H34/40: Foyle District, CWB (Buckle, 1900; Johnson & Halbert, 1902). 

H37 Armagh: Loughgall Country Park, in Polyporus squamosus on veteran ash, 26.viii.2008, RA. 

H38 Down: Tollymore Forest, under piled up Douglas fir bark, 9.x.1993, RA. 

Epuraea marseuli Reitter, 1872 syn. pusilla (Illiger, 1798) – Uncommon 

Feed on the eggs and larvae of scolytid bark beetles under sappy bark, although also considered by 

some authors to be fungivores or scavengers; colonise freshly dead wood once colonised by scolytids - 

attracted by scolytid pheromones; conifers and broadleaves. Adults also found at blossom and in tree 

fungi. Under fir bark (Johnson & Halbert, 1902). 

H2 North Kerry: Flesk, 1914, EFB (Bullock, 1914); Muckross Demesne, vi.1919, OEJ (Janson, 1920) 

H3 West Cork: Spanish Island, Roaringwater Bay, vi-vii.2010, T. Daguerre det. RA. 

H4 Mid Cork: Blarney Castle, under oak bark, 10.vii.2000, RA. 

H17 North-east Galway: Tuam, JNH (Johnson & Halbert, 1902). 

H21 Dublin: Dublin, ARH (Hogan, 1854) (Johnson & Halbert, 1902). 

H22 Meath: Balrath, vii.1912, GWN (Nicholson, 1913). 

H34/40: Foyle District, CWB (Buckle, 1900) (Johnson & Halbert, 1902). 

H35 West Donegal: Corravaddy, bark of Sitka spruce, 11.vi.1998, RA. 

H36 Tyrone: Caledon Deer Park (H7543), off oak foliage, 17.v.2006; Baronscourt Park (H3682), under 

bark on sappy fallen branch 12.ix.2006, KNAA. 

H37 Armagh: Peatlands Park, 29.v.2000, RA (Anderson, Simms & Nelson, 2000). 

H38 Down: Tollymore Forest, Douglas fir bark, 9.x.1993, RA. 

H39 Antrim: Shane’s Castle Demesne, Lough Neagh, under bark of fallen firs, vi.1923, OEJ (Janson, 

1924). 

Epuraea melanocephala (Marsham, 1802) – Common & Widespread 

Larval habits unknown; adults found at flowers of trees and shrubs, and amongst foliage; overwinter 

in leaf litter. Added to the Irish list by Irwin (1975). – but see O’Mahony 1934. Probably widespread 

but overlooked. H3, H4, H18, H21, H25, H37, H38, H39. 

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Epuraea melina Erichson, 1843 – Rare 

Adults usually found at flowers; larval habitat unknown. A curious miscellany of reports, all old, & 

not reported since 1946. 

H 5 East Cork: Cobh (Queenstown), JJW (Walker, 1895; Johnson & Halbert, 1902). 

H6 Waterford: JAP (Power, 1878; Johnson & Halbert, 1902). 

H33 Fermanagh: Portora, 17.v.1946, anon., det. T. Delaney, UM; Farnamullan, 19.v.1946, anon., det. T. 

Delaney, UM. 

H38 Down: Conlig, near Bangor, JJW (Walker, 1895; Johnson & Halbert, 1902); Greencastle, 

Carlingford Lough, 8.vi.1945, anon., det. T. Delaney. 

Epuraea pallescens (Stephens, 1835) syn. florea Erichson, 1845 – Widespread 

At flowers and tree-sap; also in fungi. ‘Probably common, but there are few records’ - H6, H12, H21, 

H31, H33, H34/35, H37, H38, H39 (Johnson & Halbert, 1902). 

H4 Mid Cork: Cobh, iv-v.1995, JJW (Walker, 1895). 


H27 West Mayo: Mullaranny, 1910-1912, JNH (Johnson & Halbert, 1912). 

H31 Louth: Carlingford, 1893, WFJ (Johnson, Halbert & Carpenter, 1894). 

H34 East Donegal: Coolmore (Rossnowlagh), 1893, WFJ (Johnson, Halbert & Carpenter, 1894). 

H36 Tyrone: Baronscourt Park (H3682), knocked from aerial dead branches on common lime, 

2.vii.2006, KNAA. 

H37 Armagh: Mullinure, Armagh, WFJ (Johnson, 1992a). 

H39 Antrim: Ballycastle, ix.1901, GWC (Chaster & Tomlin, 1902); Langford lodge (J088749), sappy 

bark of windthrown oak bough, 28.v.2012, RA. 

Epuraea rufomarginata (Stephens, 1830) – Rare 

Amongst borings of the beetle Trypodendron domesticus in a cut bough; also under dead spruce bark 

and in oak faggots; and in Daldinia fungus on ash. 

H2 North Kerry: Killarney district, Epuraea parvula “crawling about on freshly cut logs”, EFB (Bullock, 

1928). 

H31 Louth: Bellurgan Forest (J095099), under sappy bark on fresh beech log, 22.iii.2012, RA. 

H38 Down: Stormont, under bark of Acer platanoides faggots, 7.v.1994, RA. 

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H39 Antrim: Moss Plantation, Shane’s Castle, under bark of freshly thrown oak branch, 16.vii.2010, 

RA. 

Epuraea silacea (Herbst, 1783) syn. deleta Sturm, 1844 – Widespread 

At flowers, under sappy bark on dead wood, and in bracket fungi. Very rare in mid-Europe. Bullock 

(1930) is credited with the first Irish record in Anderson et al. (1997). Locally common in fungi 


H2 North Kerry: Killarney, one specimen by sweeping…within a few miles of Killarney, EFB (Bullock, 

1930). 

H3 West Cork: Trafraska, SE of Baltimore (W0525), 2.vi.1996, KNAA. 

H4 Mid Cork: Mount Desert, 19.ii.2006, MC. 

H6 Waterford: JAP (Power, 1878) E. deleta (Johnson & Halbert, 1902). 

H8 Limerick: JNH as E. deleta (Johnson & Halbert, 1902). 

H9 Clare: JNH as E. deleta (Johnson & Halbert, 1902). 

H17 North-east Galway: Clonbrock (M7439), JNH (Halbert, 1896) E. deleta (Johnson & Halbert, 1902). 

H20 Wicklow: Ballard Wood, Glendalough (T1495), 14.vi.1993; Devil’s Glen, 17.vi.1993, KNAA. 

H21 Dublin: E. silacea Hogan (1854) but queried by Haliday (Johnson & Halbert, 1902). Dublin, JNH 

(Halbert, 1900b) E. deleta (Johnson & Halbert, 1902); Clontarf, in fungi, EO (O’Mahony, 1928b). 

H22 Meath: Balrath, ix.1910, GWN (Nicholson, 1911). 

H23 Westmeath: Mullingar district, JNH as E. deleta (Johnson & Halbert, 1902). 

H25 Roscommon: Mote Park, JNH (Halbert, 1895d) E. deleta (Johnson & Halbert, 1902). 

H27 West Mayo: Westport Demesne, JNH (Johnson & Halbert, 1912). 

H33 Fermanagh: Correl Glen, 17.vii.2007, KNAA. 

H37 Armagh: Armagh City, WFJ, E. deleta (Johnson & Halbert, 1902); Tynan Abbey estate, 26.vii.2011, 

RA. 

H38 Down: Comber, CWB E. deleta (Johnson & Halbert, 1902); Murlough Dunes (J410350), 20.v.1992, 

KNAA; Drumlea, Quoile, on haw flowers, 22.v.1995, RA. 

H40 Londonderry: Drenagh Estate (C6923), 14.vi.2006; Banagher Glen (C670045), under bark, 

20.v.2007, KNAA. 

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Epuraea terminalis (Mannerheim, 1843) syn. adumbrata Mannerheim, 1852 – Rare old growth 

species 

Under bark of sappy dead oak, birch and pine timber. 

H2 North Kerry: Killarney district, Epuraea immunda, “a few taken on flowers of Umbelliferae”, 

Bullock (1928). 

H39 Antrim: Moss Plantation, Shane’s Castle (J092893), 2 males of splintered, fresh-thrown oak 

branch, 16.vii.2010, RA. 

Epuraea thoracica Tournier, 1872 – Established Introduction 

Under bark on deadwood and on resinous stumps and planks of conifers; generally with scolytids; 

mainly Scottish in Britain, but appears to have spread with conifer forestry. 

H15 South-east Galway: Pollnaknockaun Wood (M740014), v. 2003 (Regan & Anderson, 2004). 

H21 Dublin: ARH (Hogan, 1854; Johnson & Halbert, 1902). 

H34/40: Foyle District, Epuraea oblonga, CWB (Buckle, 1900; Johnson & Halbert, 1902). 

H39 Antrim: Victoria College grounds, Cranmore, 1 under bark of large conifer stump, 18.iv.2005, RA. 

Soronia grisea (Linnaeus, 1758) – Uncommon 

Develop in sap-flows from injured broadleaved trees of many species; widespread in the oak-wood 

areas of Scotland although Speight (1989a) suggests primarily associated with ash in Ireland. 

H2 North Kerry: Killarney, vii.1919, EFB; Deerpark, Killarney, 27.iv.1931, EFB (Speight, 1989a). 

H11 Kilkenny: Inistioge, 8-16.viii.1902, H.J. Turner (Turner, 1903). 

H13 Carlow: Kilcarry Br., 5.viii.1929, AWS (Speight, 1989a). 


H16 Galway: Galway, JJW (Johnson & Halbert, 1902); Knockboy (L7735), beaten from ash in mixed 

woods, 24.v.1974, MCDS (Speight, 1989a). 

H18 Offaly: Mongan Bog (N0230), in pitfall trap on raised bog, 24.viii.1983, JAG (Speight, 1989a). 

H20 Wicklow: Kilruddery, etc, on bones, ARH (Hogan, 1854) (Johnson & Halbert, 1902). 

H21 Dublin: Dundrum, FHB (Johnson & Halbert, 1902); St Anne’s, Clontarf (O2137), xii.1924, EO 

(Speight, 1989a); Phoenix Park (O0935), beaten from hawthorn & ash, 17.iv.1982, JPOC (Speight, 

1989a). 

H33 Fermanagh: Inisherk, Crom, FIT 4.ix-5.x.1992, APF. 

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H38 Down: near Belfast, JNH (Halbert, 1885); Ballintogher, Downpatrick, 1 swept in grain field, 

17.viii.2006, RA. 

H39 Antrim: Cranmore, Belfast, R. Templeton (Johnson & Halbert, 1902); Ballycastle, BT (Johnson & 

Halbert, 1902, Supplementary List); Lough Neagh (near Gawley’s gate?), 1 only, 3-4.ix.1935, EO 

(Crawford & O’Mahony, 1935; Speight, 1989a); Rea’s Wood, 1 on ash trunk during flood, 10.xii.1999, 

RA; Great Deer Park, Glenarm (D2911), three from soggy bracket of Fistulina on old oak, 16.ix.2006, 

KNAA. 

Soronia punctatissima (Illiger, 1794) – Rare 

Develop in sap-flows from injured broadleaved trees of many species; widespread in the oak-wood 

areas of Scotland but associated more with ash in Ireland (Speight, 1989a). ‘An Irish-taken specimen in 

Dublin Museum, which had been for many years in the Trinity College collection’ (Johnson & Halbert, 

1902). 

H2 North Kerry: Muckross Demesne, one beneath a boletus on an oak, June 1919, OE Janson (1920); 

Muckross, male, vii.1935, EFB; Muckross (V9585), male under loose bark over fungus-infested sap-run 

on large living ash in deciduous woods, 20.v.1988, MCDS (Speight, 1989a). 

H11 Kilkenny: Inistioge (S6337), in Goat Moth Cossus infected tree, H.J. Turner (Turner, 1903). 

H12 Wexford: Kilcarry Bridge, 5.viii.1929, AWS (Speight, 1989a). 

H30 Cavan: Cloverhill (H4115), 2 at sap on damaged oak, GWN (Nicholson, 1917). 

H33 Fermanagh: Inisherk, Crom (H357243), 1 in FIT 1.vi.-7.vii.1992, APF. 

Glischrochilus quadripunctatus (Linnaeus, 1758) – Casual Introduction? 

On conifers. Just a few isolated records, suggesting possibly introductions. 

H2 North Kerry: near Killarney, EFB (O’Mahony, 1929a). 

H19 Kildare: Athy, beside R. Barrow (S6893), 30.viii.1980, D.Murphy (Speight, 1989a). 

H36 Tyrone: Dungannon, Ips quadripustulata, Haliday ms (Johnson & Halbert, 1902). 

Glischrochilus hortensis (Geoffroy in Fourcroy, 1785) – Uncommon 

Adults usually at sappy stumps, in fungi, or amongst chippings from broad-leaved trees or conifers; 

attracted to a wide range of decaying substrates and also known as a pest of soft fruits. Adults & 

larvae have been found inside decaying Fistulina bracket and beneath sappy bark on oak stump 

(KNAA, pers obs). G. quadripunctata Herbst in O’Mahony (1929). 

H2 North Kerry: Cruagh, Killarney, 17.vii.1930, EFB (O’Mahony, 1929a; Speight, 1989a). 

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H4 Mid Cork: Rock Castle Woods (W5653), 6.vi.1994 & Shippool Woods (W5757), Bandon River, 

23.v.1994, KNAA; River Blackwater, near Mallow, 14.x.2001, S. Gregory (KNAA); Crosshaven 

(W7761), 7.v.1995, Ballyphehane, Cork City (W6869), i.vi.1998 & Mount Desert (W628720), 19.ii.2006, 

MC (Cawley, 2011a). 

H5 East Cork: near Tallow (W9993), in numbers inside raspberries and tomatoes in polytunnel; also 

reported causing similar damage from near Cobh, D. Berridge (O’Connor, 2005); Ballyannan Wood 

(W875720), Malaise trap, 2003 & Ballylanders (W949623), netted, 23.viii.2011, TG. 

H6 Waterford: near Lismore, reported causing damage to soft fruit, D. Berridge (O’Connor, 2005); 

Kilbarry Bog, 3.ix. & 9.ix.2001 (O’Meara, 2008); Rincrew (X0880), 30.ix.2004, MC (Cawley, 2011a). 

H11 Kilkenny: Kilkenny, 24.iv.1925, R.A. Phillips (Speight, 1989a). 

H12 Wexford: New Ross, 24.v.1994, PJH; woodland bordering River Barrow, Mountgarrett (S7229), 

13.iv.1998, MC (Cawley, 2011a). 

H14 Laois: Abbeyleix Wood (S4284), 16.iii.2003, MC (Cawley, 2011a). 

H19 Kildare: Sallins, 5.v.1935, AWS (Speight, 1989a); Moore Abbey Wood (N635089), 27.iv.2007 & 

banks of Grand Canal, Barrettstown (N874235), 9.viii.2008, MC (Cawley, 2011a). 

H21 Wicklow: Three Lakes, Table Mountain, pitfall 5.vi.-28.viii.1998, MN det. RA/SM. 

H22 Meath: Balrath, 26.ix.1913, vi.1916, GWN (Nicholson, 1914, 1917) 

H25 Roscommon: St John’s Wood (M9956), alighting on stack of freshly cut poles, 13.iv.2010, KNAA. 


H37 Armagh: Loughgall Country Park, 26.viii.2008, RA. 

Glischrochilus quadriguttatus (Fabricius, 1777) – Rare 

As above; feeds on fermenting sap; associated with ash in Ireland (Speight, 1989a). 

H2 North Kerry: Cahirnane, Killarney, vi.1908; Killarney, viii.1915, EFB (Speight, 1989a); Killarney, 

1928, “two specimens from under bark” (Bullock, 1928); Muckross (V9585), under loose bark on 

fungus-infested sap run on large, living ash, 10.v.1988, MCDS (Speight, 1989a). 

H21 Dublin: Glendhu, at sap, v.1928, H. Heasler (O’Mahony 1929; Speight, 1989a). 

H30 Cavan: Cloverhill, Nicholson (1915). 

H31 Louth: Bellurgan Forest (J095099), 3 under bark of fallen sycamore, 22.iii.2012, RA/MDB. 

Pityophagus ferrugineus (Linnaeus, 1758) – Established Introduction 

Under bark on dead pine; adults have also been reported from plant roots in arable land in Britain. 

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H2 North Kerry: Muckross, coll. Furlong, in Haliday ms (Johnson & Halbert, 1902). 

H17 Galway: Tuam, GHC (Carpenter, 1895) (Johnson & Halbert, 1902). 

H22 Meath: Laytown Sandhills, 1894 (DNFC, 1894) (Johnson & Halbert, 1902); Balrath (N7173), 5 

under bark of felled pine, vi.1916, GWN (Nicholson, 1917). 

H36 Tyrone: Ballyard, Trillick (H339587), on silver fir, 3.vi.1998, SC. 

FAMILY MONOTOMIDAE 

Larvae feed on larvae of other small beetles, including certain scolytid bark beetles; in damp 

conditions where there is mould or sap, especially beneath bark on freshly dead wood. 

Rhizophagus bipustulatus (Fabricius, 1793) – Casual introduction? 

Adults and larvae under dead bark of most species of broad-leaved trees; feed on fungal hyphae and 

will also eat dead scolytid bark beetle larvae. Not included (in error) in Anderson et al. (1997), but it is 

a ubiquitous species in Britain and the near Continent, and it might be expected to be either common 

in Ireland or entirely absent. A single isolated old record suggests a possible casual introduction. 

H2 North Kerry: Near Killarney, pre 1914, EFB (Bullock, 1914). 

Rhizophagus cribratus Gyllenhal, 1827 – Scarce 

Usually found around tree roots in litter, etc, especially oaks. Under bark, but not common (Johnson & 

Halbert, 1902). No modern records. 


H4 Mid Cork: Castle Cor near Mallow, TVW (Wollaston, 1847). 

H16 West Galway: Galway, Fowler (1891) (Johnson & Halbert, 1902). 

H21 Dublin: Little Dargle, JNH (Halbert, 1900b; Johnson & Halbert, 1902). 


H27 West Mayo: Achill, 1910-1912, JNH (Johnson & Halbert, 1912); Westport Demesne, 1910-1912, 

JNH (Johnson & Halbert, 1912). 


H36 Tyrone: Caledon Forest, WFJ (Johnson, 1896). 

H37 Armagh: Armagh City, WFJ (Johnson, 1891); Palace Grounds, in moss, WFJ (Johnson, 1892a); 

Loughgilly, WFJ (Johnson & Halbert, 1902). 

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Rhizophagus depressus (Fabricius, 1793) – Established Introduction 

Under bark of dead pine. 

H2 North Kerry: Kenmare Demesne, Killarney in some numbers under bark of recently felled pines, 

25.vi-30.vi.1919, OEJ (Janson, 1920); Rossbeigh dunes, vii.1902, HSJD (Donisthorpe, 1902). 

H17 North-east Galway: Tuam, GHC (Carpenter, 1895). 

H18 Offaly: Clonad Wood, JNH (Halbert, 1895c) (Johnson & Halbert, 1902). 

H21 Dublin: Clontarf, EO (O’Mahony, 1928b). 

H33 Fermanagh: Ballyard, Trillick, 3.vi.1998, SC. 

H34/40: Foyle district, CWB (Buckle, 1900; Johnson & Halbert, 1902). 

H37 Armagh: Poyntzpass, WFJ (Johnson, 1900; Johnson & Halbert, 1902); Peatlands Park, under Scots 

pine bark, abundant, 29.v.2000 RA (Anderson, Simms & Nelson, 2000). 

H39 Antrim: Shane’s Castle Demesne, Lough Neagh, under bark of fallen firs, June 1923, OEJ (Janson, 

1924); Portgenone Forest, 10.vi.2004, SC det. RA. 

Rhizophagus dispar (Paykull, 1800) – Common & Widespread 

Under bark of most dead broad-leaved trees, and in bracket fungi, also on plantation conifers. 

Common, often abundant in Ireland: H1, H2, H4,H6, H9, H16, H17, H18, H20, H21, H25, H26, H27, 

H28, H33, H34, H35, H36, H37, H38, H39, H40. 

Rhizophagus ferrugineus (Paykull, 1800) – Scarce 

Under bark on deadwood and in heart-rot; also at heartwood of sawn logs where in contact with soil. 

H2 North Kerry: Kenmare Demesne (V99), in some numbers under bark of recently felled pines, 

between June 25 & 30, 1919, O.E. Janson (1920). 

H4 Mid Cork: Castle Cor, TVW (Wollaston, 1847). 

H15-17 Galway: Tuam, under fir bark (Carpenter, 1895;Johnson & Halbert, 1902). 

H21 Dublin: Merrion Demesne, 1854 (Hogan, 1854). 


H33: Fermanagh: Inisherk (H357248), Crom Castle Estate, one in flight trap in shoreline carr 

woodland, vii.2007, KNAA. 

H36 Tyrone: Davagh Bridge, Davagh Forest, 2.vi.1998, SC. 

H34 Donegal: Foyle district, CWB (Buckle, 1900; Johnson & Halbert, 1902). 

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H38 Down: Hillsborough Forest, 28.viii.1993, RA; Audley’s Castle, 20.ii.2008, RA; Leitrim Lodge 

(J223256), 29.ii.2012, RA. 

H39 Antrim: Breen Wood NNR (D123336), 13.vii.2005, RA, 9.vii.2007, KNAA. 

Rhizophagus parallelocollis Gyllenhal, 1827 – Established Introduction 

Present as a native saproxylic in Britain, under bark on dead wood, but only known from 

synanthropic situations in Ireland. 

H21 Dublin: Finglas, on vegetable refuse, xi.1924, JNH (Halbert, 1937). 

H37 Armagh: Cathedral Grammar School, one in room, WFJ (Johnson, 1892a). 

H39 Antrim: Armoy Churchyard, in numbers on tombstones, ix.1901, GWC (Chaster & Tomlin, 1902). 

Rhizophagus perforatus Erichson, 1845 – Rare 

Under bark on dead broad-leaved trees. Adults may occasionally be taken away from larval habitat, 

including town gardens. 


H15-17 Galway: Galway, JJW (Fowler, 1891) (Johnson & Halbert, 1902). 

H21 Dublin: local, JNH (Halbert, 1900b); Clontarf, EO (O’Mahony, 1928b). 

H22 Meath: Balrath, in bracket on an ash stump, 26.ix.1913, GWN (Nicholson, 1914). 

H30 Cavan: Cabra Castle, under bark, vii.1912, GWN (Nicholson, 1913). 

H37 Armagh: Palace Demesne, in moss, WFJ (Johnson, 1892a). 

H38 Down: Rostrevor Oakwood, under bark on oak logs, 13.v.2000, RA (Anderson, Simms & Nelson, 

2000) 

H39 Antrim: Ballycastle, BT (Johnson & Halbert, 1902, Supplementary List). 

FAMILY LAEMOPHLOEIDAE 

Cryptolestes ferrugineus (Stephens, 1831) – Established Introduction 

In Britain, under bark on beech, especially where still sappy; also on oak, horse chestnut, etc; also in 

granaries. Irish records have all been from synanthropic situations. Ireland, Haliday coll. (Johnson & 


H21 Dublin: Clontarf, from a city corn store, EO (O’Mahony, 1927). 

H31 Louth: south, HKGC (Halbert, 1900b; Johnson & Halbert, 1902). 

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FAMILY CRYPTOPHAGIDAE Silken Fungus Beetles 

CRYPTOPHAGIDAE SUBFAMILY CRYPTOPHAGINAE 

Henoticus serratus (Gyllenhal, 1808) – Rare 

Under bark on mouldy decaying wood, especially birch & beech, but also oak, spruce, pine, etc; larvae 

possibly in fungal mycelium; adults attracted to Salix catkins, hawthorn blossom, sugary exudates, 

bracket fungi. Added to the Irish list in 1923, but not reported since. 

H38 Down: Donard Demesne, Newcastle, two under bark of felled beech, vi.1923, OEJ (Janson, 1924). 

Cryptophagus spp. These small beetles are usually fungal or detritus feeders. 

Cryptophagus dentatus (Herbst, 1793) – Common & Widespread 

In bracket fungi, decaying twigs & leaves; bird nests in tree branches; with mouldy materials in 

buildings; larvae will feed on dead or dying insect larvae. Not a true saproxylic but almost invariably 

present in decaying wood. Common: Ulster, Connaught, Leinster, Munster (Johnson & Halbert, 1902); 

H1, H4, H19, H21, H25, H27, H33, H36, H37, H38, H39, H40. 

Cryptophagus intermedius Bruce, 1934 – Rare 

Known from borings of the bark beetle Hylesinus varius under ash bark and at sycamore sap. 

H38 Down: Deramore, 2 in abandoned grass bale, 26.iv.1995, (Anderson, Simms & Nelson, 2000). 

Cryptophagus denticulatus Heer, 1841 – Rare 

Known in Ireland from gathered hay and under fungoid bark on trees; ecology poorly understood and 

only distinguished from amongst C. dentatus in the 1950s. 

H9 Clare: Cragroe Castle, 3 sieving hay on barn floor, 26.v.2003, RCW (Regan & Anderson, 2003). 

H38 Down: Morelands Meadow, 1 under bark on slender part of a fallen beech branch in parkland, 

18.xi.1992, RA. 

H39 Antrim: Shane’s Castle, 1 under fungoid bark of a large oak log, 1.vii.2010, RA. 

Cryptophagus ruficornis Stephens, 1830 – Recent Arrival 

Associated with the fungus Daldinia concentrica fruiting on deadwood of ash, and D. loculata on burnt 

birch. Added to the Irish list only in 1992; conceivably an overlooked native, but more likely a recent 

colonist from Scotland, especially in view of Daldinia being most frequent in Ireland in the eastern and 

south-eastern counties and the only reports of the beetle are from the north-east. 

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H37 Armagh: Loughgall Country Park, frequent in Polyporus squamosus on veteran ash, 26.viii.2008, 

RA. 

H38 Down: Belvoir Forest, in rotted Fistulina on veteran oak, 12.x.2003, RA; Hillsborough Forest, in 

pitfall under Douglas fir, 29.vi.2005, RA; Mount Stewart, Sea Plantation, in fungoid bark on beech, 

13.ix.2003, RA. 

H39 Antrim: Ballyskeagh (J280671), one under loose dry bark of a splintered, burnt ash stump, 

8.xii.1992, RA (Nash, Anderson & O’Connor, 1997); Mountsandel Wood, Coleraine, in old chaffinch 

nest in tree hole, 9.ii.2008, RA. 

H40 Londonderry: Drenagh Estate (C6923), plentiful on old sycamore in avenue with burnt out 

hollow base and Daldinia fruiting, 14.vi.2006, KNAA. 

CRYPTOPHAGIDAE SUBFAMILY ATOMARIINAE 

Adults and larvae of most species probably feed on fungal hyphae and moulds. With the present state 

of knowledge, individual species are particularly difficult to allocate to the decaying wood category 

with confidence when individuals have been found in leaf litter or flood refuse (both including 

decaying twigs), as well as more distinct decaying wood habitats. 

Atomaria pulchra Erichson, 1846 – Established Introduction? 

Has been found in all kinds of decaying conifer wood: heaps of bark shavings, burnt wood, sawn logs 

and stacked timber. 

H38 Down: Rostrevor Forest (J189184), one male in Ips typographus pheromone trap in Sitka spruce 

stand, 27.vii.2006, SC det. RA (Anderson & Clawson, 2006). 

Atomaria umbrina (Gyllenhal, 1827) – Rare 

Associated with the fruiting bodies of wood-rotting fungi, especially the gill fungi Armillaria mellea 

and Pholiota spp, fruiting from decaying wood in woodlands. Adults also taken in grass heaps. 

H1 South Kerry: Lickeen Wood, Glencar, vi.1924, OEJ (Janson & Bonaparte-Wyse, 1924). 


H21 Dublin: Finglas, x.1924, JNH (Halbert, 1924). 

H37 Armagh: Armagh, WFJ det. C. Johnson; Mullinure, Loughry’s Lough, WFJ (Johnson, 1892a). 

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FAMILY EROTYLIDAE 

Dacne bipustulata (Thunberg, 1781) – Scarce 

Adults normally frequent fruiting brackets of the softer polypore fungi on trunks of broad-leaved 

trees; has been reared from Laetiporus sulphureus, Piptoporus betulinus and Pleurotus ostreatus. 

Widespread in northern counties, but surprisingly just one report from the south. 

H19 Kildare: a quarter mile east of Landenstown House, swept from beech in a small plantation, 

9.v.1948, AWS (Stelfox, 1952). 

H33 Fermanagh: Castle Coole Park (H2643), from Bjerkandera adusta on beech, 18.v. & 27.vii.2006, 

KNAA; Crom Castle Park (H3624), at very fresh Laetiporus sulphureus on standing dead oak, 18.v.2007, 

KNAA; Rossyvullan Hill, Castle Coole, in Polyporus squamosus on sycamore, 20.vii.2010, RA. 

H36 Tyrone: Caledon Deer Park (H7543), one at Bjerkandera adusta on old sycamore, 16.vi.2006, & 

teneral on P squamosus, 16.viii.2006, KNAA. 

H37 Armagh: Loughgall Country Park, abundant in Polyporus squamosus on veteran ash, including 

several entirely pale (rufous) specimens, 26.viii.2008, RA; Tynan Abbey Estate, in Polyporus squamosus 

on elder, 26.vii.2011, RA. 

H38 Down: Hillsborough Forest, Bjerkandera on beech logs, 24.iii.2003, RA. 

H39 Antrim: Lagan Meadows, Belfast, several from decaying Bjerkandera adusta on a sycamore stump, 

9.iii.1993, RA (Anderson, 1995); Newforge Lane, Belfast, in Gymnopilus on Scots pine, 9.vii.2008, RA; 

Shane’s Castle, in Bjerkandera on beech log, 1.vii.2010, RA. 

H40 Londonderry: Springhill House (H8683), mixed woodland, 25.vi.1992, KNAA (Anderson, 1995). 

Triplax aenea (Schaller, 1783) – Doubtful 

Usually associated with the fungus Pleurotus ostreatus growing on trunks of broad-leaved trees. 

“Certainly Irish” in Haliday’s ms list; one of J Tardy’s captures, but specimens cannot be traced 

(Johnson & Halbert, 1902); not included in Anderson et al. (1997). A distinctive species and one 

unlikely to be overlooked. 

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FAMILY CERYLONIDAE 

Feed on fungal hyphae and spores. 

Cerylon fagi Brisout de Barneville, 1867 – Rare old growth species 

Lives under bark of deadwood and within decaying heartwood, especially oak, and especially in the 

later stages of decay; also found in beech and ash. Very localised in Europe. [Records for Coole Park 

and Charleville Wood in Regan & Anderson (2004) refer to C. ferrugineum]. 

H2 North Kerry: Killarney district, crawling on log, EFB (Bullock, 1928). 

H15 South-east Galway: Garryland wood, under fungoid bark on beech, 24.ix.2007, RA. 

Cerylon ferrugineum Stephens, 1830 – Scarce old growth species 

Develops beneath bark on dead broad-leaved timber in the early stages of decay; feeds on fungal 

hyphae and spores. Mostly in ancient woodland and wood pasture; widespread through northern 

and central Europe. 

H1 South Kerry: Tomies Wood (V9089), under bark on dead oak, 14.v.1988, 17.viii.1990, 

7.xi.2002KNAA; Uragh Wood (V8362), 25.v.1994, KNAA. 

H2 North Kerry: nr. Killarney, EFB (Bullock, 1914); Derrycunihy Wood (V8981 & V9080), 17.viii.1990 

& 13.x.2001; Knockrear Park, Killarney, 7.xi.2002, KNAA. 

H15 South-east Galway: Punchbowl, Gort (M4500), 24.v.1988, KNAA. 

H18 Offaly: Charleville Wood (N315226), in Laetiporus, v. 2003 (Regan & Anderson, 2004). 

H21 Dublin: The Millenium Park, Blanchardstown (O066387), emerged from bracket fungi on a 

rotting log in mixed planted woodland verge in this suburban park, 5.iii.2012, MC. 

H25 Roscommon: St John’s Wood (M9956 & N0055), 7.vi. & 6.viii.2010, KNAA. 

H28 Sligo: Union Wood (G680285), 14.vi.1999, KNAA. 

H33 Fermanagh: Knocknabrass, 26.v.1992; Corlatt Wood, 27.v.1992; Reilly Wood, 28.v.1992; Killy 

Upper, 7.vii.1992, all Crom Castle Estate, KNAA; Castle Coole Park (H2643), 2.vi.1992, 18.v. & 

17.viii.2006, KNAA; Castle Archdale, under bark of fallen oak branch, 11.v.1997, RA; Florencecourt, 

under bark on oak, 21.vi.2010, RA; Rossyvullan Hill, Castle Coole, fungoid oak bark, 26.vi.2010, RA; 

Wattle Bridge, under bark on hedgerow ash, 11.viii.2011. 

H35 West Donegal: Hollymount, Rathmullan (C279271), 22.ii.2012, MDB. 

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H36 Tyrone: Baronscourt Park (H3682), beneath bark on fallen oak branches, 16.v.2006, RA & KNAA; 

25.vii & 12.ix.2006, KNAA. 

H37 Armagh: Peatlands Park, under bark on dead birch, 23.x.1993, RA; Tynan Abbey Estate, under 

fungoid bark on oak, 21.vi.2011, RA. 

Cerylon histeroides (Fabricius, 1793) – Scarce old growth species 

Develops beneath bark on dead broad-leaved timber, generally at later stage in succession when 

cavities moist; feed on fungal hyphae and spores. 

H2 North Kerry: Killarney, JNH (Halbert, 1898c; Johnson & Halbert, 1902). 

H4 Mid Cork: Blarney Castle, 10.vii.2000, RA. 

H19 Kildare: Celbridge, ix.1936, JNH (Halbert, 1937). 

H25 Roscommon: St John’s Wood (M9956 & N0056), 5.6.viii & 7.x.2010, KNAA. 

H33 Fermanagh: Castle Coole Park (H2643), 17.vi.2006, KNAA; Knockninny ASSI, 1.vi.2007, RA; 

Rossyvullan Hill, Castle Coole, 20.vii.2010, RA. 

H36 Tyrone: Caledon Park (H7543), 17.v.2006, KNAA; Brookend ASSI, 8.v.2007, RA. 

H37 Armagh: Peatlands Park, 23.x.1993, RA (Anderson, Simms & Nelson, 2000); Annagarriff NNR, in 

Formica nest, 3.iv.2007, RA; The Argory, 29.iii.2011, RA; Tynan Abbey Estate, 21.vi.2011, RA. 

H38 Down: Minnowburn Beeches, 29.i.1994; Rostrevor Oakwood NNR, 13.v.2000, RA (Anderson, 

Simms & Nelson, 2000). 

H39 Antrim: Belvoir Park, RS (Johnson & Halbert, 1902); Deramore, 11.xi.1992, RA; Rea’s Wood, 

Antrim, 20.iv.1995, RA; Farr’s Bay NNR, 4.vii.1986, RA; Moss Plantation, Shane’s Castle, 16.vii.2010. 

H40 Londonderry: Magilligan, CWB (Johnson & Halbert, 1902); Scab Island, Lough Beg, 6.xii.1975, 

RA. 

FAMILY ENDOMYCHIDAE False Ladybirds 

Endomychus coccineus (Linnaeus, 1758) – Recent arrival 

Lives gregariously with its larvae on or around fungoid growth under bark of dead timber; especially 

in beech, but also in apple, crack willow, horse chestnut and birch. Sites tend to be low-lying marshy 

areas along floodplains, or close by. First noted in Ireland in 1976 and spreading steadily in north-east 

(RA). 

H33 Fermanagh: Rossyvullan Hill, Castle Coole, 24.viii.2010, RA. 

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H36 Tyrone: Brantry Lough (H748534), under bark of dead standing beech, 26.i.2012, MDB/RA. 

H37 Armagh: Derrycarne Road, Portadown (J008576), seen on several occasions between 1990 and 

2012 in rural garden/orchard, most recently 12.v.2012, BN; Tynan Abbey Estate, 26.vii.2011, RA. 

H38 Down: Morelands Meadow, Belvoir, 1.xi.1992, RA; Belvoir Forest, 1.ix.1997, RA; Killynether 

Wood, 2.ii.2008, RA; Newtownbreda Village, urban garden, Belfast, 30.iii.2008; Shooter’s Island 

(J494470), under bark of dead standing sycamore, 23.ii.2012, MDB/RA. 

H39 Antrim: Rea’s Wood, Antrim, 1st record - seven clinging to dry bark under a pile of fresh wind- 

felled willow boughs in native alder woodland, on juvenile fruiting bodies of Trametes, 2.x.1976, RA 

(Anderson, 1977); Rea’s Wood, 3.xii.1994, RA; Clement Wilson Park, Lagan Valley (J325695), 8.i.1992, 

RA (Anderson, 1992); Lady Dixon Park, Belfast, 28.ii.2005, RA; Belfast Castle, 12.ii,2007, RA; Toome 

Weir, Lough Neagh, 3.v.2007; Quarterland, Dundrod, rural garden, 5.xi.2011, RA. 

H40 Londonderry: Springhill House, 26.iv.2003, RA; The Umbra, 2.i.2004, RA. 

FAMILY CORYLOPHIDAE 

Figure 22: Endomychus coccineus © Roy Anderson 

Orthoperus nigrescens Stephens, 1829 syn. mundus Matthews, A., 1885 – Rare 

Under fungoid bark. 

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H19 Kildare: Louisa Bridge on Royal Canal, Leixlip, one, 11.ix.1942, AWS det. PMH (Nash, Anderson 

& O’Connor, 1997). 

FAMILY LATRIDIIDAE Brown Scavenger or Plaster Beetles 

Most feed on mould. 

Enicmus testaceus (Stephens, 1830) – Rare 

In ripe slime fungus on beech and other trunks; Reticularia lycoperdon the main host. 

H9 Clare: Lough Gash (R3967), one on silt by lough, vi.1989 (Lott & Bilton, 1991). 

H33 Fermanagh: Molly Mountain (H233282), one from Inonotus radiatus brackets on sycamore in open 

pastureland, 23.vi.1995, RA (Anderson, 1998; Anderson et al., 2000). 

H38 Down: Hollymount NNR (J459432), in grass clippings, 31.v.2011, MDB det. RA. 

FAMILY MYCETOPHAGIDAE Hairy Fungus Beetles. 

Associated with decaying bark and wood. 

Litargus connexus (Geoffroy in Fourcroy, 1785) – Recent arrival 

Develops in Pyrenomycetes (Diatrype, Hypoxylon, Daldinia). Adult beetles feed on wood-decay fungi 

more generally (Pleurotus, Fomes, Piptoporus, Polyporus) and may also be found under dead bark. Only 

recently discovered in Northern Ireland (Anderson, 1993). The fungus found on birch locally is 

Daldinia loculata, after recent revisions of the species concept in Daldinia. 

H37 Armagh: Derryadd Lough (H915604), 3 under loose bark in fire-damaged birch stumps which 

were extensively affected by a red heart rot, on cutover peat, 18.v.1992, RA (Anderson, 1993); 

Derryadd (H911606), 1 in Piptoporus on birch, 11.v.2012, MDB/RA. 

Mycetophagus multipunctatus Fabricius, 1793 – Recent arrival 

With bracket fungi on ash and other broad-leaved trees, and especially along alluvial floodplain 

situations. 

H38 Antrim: Rea’s Wood (J143583), 8 adults & 4 larvae in Polyporus on Salix fragilis, 12.iv.2007, RA 

(Anderson, 2007b); Portmore Lough, RSPB Reserve (J122688), 3 in Daedaleopsis confragosa on sallow, 

willow carr, 8.x.2009, RA/MDB; Shane’s Castle (J115885), 1 dead under fungoid bark on oak, 1.vii.2010, 

RA. 

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FAMILY CIIDAE Minute Tree Fungus Beetles 

Figure 23: Mycetophagus multipunctatus © Roy Anderson 

Develop in bracket and other fungi in and on dead and decaying timber. While the larvae appear to 

have restricted fungal associations, adults may feed on wood-decay fungi more widely. 

Octotemnus glabriculus (Gyllenhal, 1827) – Common & Widespread 

Develop chiefly in young, expanding brackets of Trametes versicolor; also in Pseudotrametes gibbosa; 

particularly characteristic of beech Fagus, but also found in association with other broad-leaved tree 

species. ‘Common where it occurs’ (Johnson & Halbert, 1902). H4, H5, H6, H15, H16, H18, H19, H21, 

H25, H27, H29, H33, H37, H39, H40. 

Orthocis alni (Gyllenhal, 1813) – Scarce 

Reputedly associated with Jew's Ear Fungus Auricularia auricula-judae, mainly on dead stems of elder, 

but also regularly knocked from aerial dead branches on open-grown oak trees; larvae develop in the 

soft fungoid sapwood of the colonised dead host tissues. The record for Phoenix Park (Halbert, 1893) 

is to be deleted (Johnson & Halbert, 1902). For many years only known from a few old records, but 

recently shown to be more widespread on veteran open-grown parkland trees in historic demesnes. 

H1 South Kerry: Glencar (V78), vii.1924, OEJ (Jansen & Wyse, 1924). 

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H2 North Kerry: Muckross Demesne, a few beneath a ‘boletus’ (i.e. a polypore) on an oak, vi.1919, 

OEJ (Janson, 1920) 

H25 Roscommon: Mote Park, JNH (Halbert, 1898a). 

H33 Fermanagh: Castle Coole Park (H260430), aerial dead branches on oak, 2.vi.1992 & 17.vi.2006, 

KNAA; Crom Castle, beaten from Ganoderma on oak, 21.viii.2004, RA 

H36 Tyrone: Baronscourt Park (H3682), beaten from aerial dead branches on parkland oak, 12.ix.2006, 

KNAA. 

H37 Armagh: Loughgall Country Park, in Polyporus squamosus on sycamore, 26.viii.2008, RA; Tynan 

Abbey Estate, in Inonotus dryadeus on oak, 1.vii.2011, RA. 

H39 Antrim: Ballycastle, Tomlin coll., NMW, Cardiff (G. Orledge pers.comm.); Barnett’s Park, Belfast, 

in Inonotus radiatus on alder with larvae, 11.i.2011, Polyporus squamosus on elder, 26.vii.2011, RA. 

Cis bidentatus (Olivier, 1790) – Common & Widespread 

Most regularly develops in the brackets of Piptoporus betulinus and Laetiporus sulphureus, but also 

reported from Pleurotus, Polyporus squamosus, and Ganoderma spp. Locally abundant (Johnson & 

Halbert, 1902). H15, H18, H19, H20, H21, H33, H36, H37, H38, H39, H40. 

Cis bilamellatus Wood, 1884 – Established Introduction 

An Australian species which has been accidentally introduced into Britain and Ireland. Develops in 

soft bracket fungi. First noted in Ireland in 2003. 

H3 West Cork: Glengarriff Woods, on birch polypore, v.2003, GMO. 

H38 Down: Belvoir Forest, 15.iv, 23.iv.2004, RA; Rostrevor Oakwood (J186173), from Stereum 

gausapatum on fallen oak bough 23.v.2007, KNAA; Helen’s Tower, Clandeboye, 10.ii.2011, RA; 

Portaferry House, 30.xi.2011, RA; Leitrim Lodge (J223256), under bark on dead Pinus contorta, 

29.ii.2012. 

H39 Antrim: Portmore Lough, RSPB Reserve, 8.x.2009, RA; Barnett’s Park, Belfast, 11.i.2011, RA. 

Cis boleti (Scopoli, 1763) – Common & Widespread 

Develop in the fully expanded fruit bodies of the fungus Trametes versicolor, although adults may also 

be found feeding at other fungi. This is the most frequent species of the family. Common: Ulster, 

Connaught, Leinster, Munster (Johnson & Halbert, 1902). H3, H4, H5, H9, H12, H13, H15, H16, H18, 

H25, H29, H33, H36, H37, H38, H39, H40. 

Cis fagi Waltl, 1839 – Rare old growth species 

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Primarily develops in the mycelium of Laetiporus sulphureus in cubical red-rotten oak heartwood; 

larvae have also reported from subcortical mycelial sheets of Armillaria sp; adults reported widely, 

feeding at a wide variety of wood-decay fungi. 

H2 North Kerry: Killarney district, Cis fuscatus “in dry fungi on rotten wood”, EFB (Bullock, 1928). 

H18 Offaly: Charleville Wood (N320227), 24. v. 2003, RCW (Regan & Anderson, 2004). 

Cis festivus (Panzer, 1793) – Uncommon 

In fungal brackets, particularly Stereum spp., on decaying timber. 


H5 East Cork: Cobh (Queenstown), under beech bark, iv-v.1895, JJW (Walker, 1895; Johnson & 


H13 Carlow: Borris, JNH (Johnson & Halbert, 1902). 

H15 South-east Galway: Coole Park (M430042), 25.v. 2003, GMO; Pollnaknockaun Wood (M740014), 

23.v. 2003, GMO; Portumna Forest, 23.v.2003, GMO (all - Regan & Anderson, 2004). 

H18 Offaly: Charleville Wood (N315226), 24.v. 2003, RA (Regan & Anderson, 2004) 

H25 Roscommon: St John’s Wood (M9956 & N0056), knocked from dead hazel pole, 6.x.2010; reared 

from old bracket fungus Inonotus radiatus collected 7.x.2010, KNAA. 

H33 Fermanagh: Inisherk, Crom, 27.vi.2000, RA; Castle Coole Park (H2643), aerial dead branch on 

oak, 17.vi.2006, KNAA. 

H36 Tyrone: Baronscourt Park (H3682), 15.vi.2006, KNAA. 

H37 Armagh: Peatlands Park, 12.viii.2000, RA. 

H38 Down: Rostrevor Oakwood, 13.v.2000, RA (Anderson, Simms & Nelson, 2000); Morelands 

Meadow, 15.iv.2004, RA; Castle Ward Park (J5649), aerial dead branches on parkland oaks, 20.viii.2006 

& 28.x.2006, KNAA; Shooter’s Island (J494470), in Stereum hirsutum on oak, 23.ii.2012, RA. 

H39 Antrim: Newforge Plots, 11.xi.1992, RA; Great Deer Park, Glenarm (D2911), 22.v. & 24.x.2007, 

KNAA; Glenarm Nature Reserve, 1.xi.2011, RA. 

H40 Londonderry: Banagher Glen (C670045), aerial dead branch with Stereum rugosum on hazel, 

25.v.2007, KNAA. 

Cis hispidus (Paykull, 1798) – Rare 

Develop in the brackets of the fungi Trametes hirsutus and T. versicolor; possibly favours open wood 

pasture situations. 

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H14 Laois: Maryborough, WES (Johnson & Halbert, 1902). 

Cis nitidus (Fabricius, 1792) – Widespread 

Develops in the brackets of Ganoderma australe in particular; the larvae have particularly heavily 

developed mandibles for chewing this exceptionally woody fungus; can also develop in Piptoporus 

betulinus. Most often found in old parklands or wood pastures. 

H2 North Kerry: near the base of Torc Mountain (V98), in a ‘boletus’ (ie a polypore), vi.1919, OEJ 

(Janson, 1920); Muckross, in birch bracket, 11.v. onwards, 1938 (Donisthorpe, 1938). 

H5 East Cork: Fota Wildlife Park, 13.iv.2011, RA. 

H9 Clare: Lough Graney (R559949), 25.v. 2003, GMO (Regan & Anderson, 2004). 

H11 Kilkenny: Thomastown, JNH (Johnson & Halbert, 1902). 

H12 Wexford: Killurin, JNH (Johnson & Halbert, 1902). 

H13 Carlow: Borris, JNH (Halbert, 1895e; Johnson & Halbert, 1902). 

H15 South-east Galway: Portumna Bay, The Quay, Park, etc (M841032), v. 2003; Derrycrag Wood 

(R738994), 23.v. 2003 (all - Regan & Anderson, 2004). 

H16 West Galway: Galway, JJW (Johnson & Halbert, 1902). 

H18 Offaly: Charleville Wood (N315226), 24.v. 2003, GMO/RCW (Regan & Anderson, 2004) 

H21 Dublin: Dublin (Hogan, 1854); Phoenix Park, JNH (Halbert, 1893) (Johnson & Halbert, 1902). 

H23 Westmeath: Athlone district, JNH (Johnson & Halbert, 1902). 

H25 Roscommon: Mote Park, vi.1897, JNH (Halbert, 1898a) (Johnson & Halbert, 1902). 

H31 Louth: Bellurgan Forest (J096098), 22.iii.2012, RA/MDB. 

H32 Cavan: Cloverhill, ix.1913, GWN (Nicholson, 1914). 

H33 Fermanagh: Castle Coole Park (H2643), G australe, 2.vi.1992, 18.v & 27.vii.2006; Crom Castle Park 

(H32), 7.vii.1992, KNAA. 

H36 Tyrone: Baronscourt Park (H3682), G australe, 25.vii.2006; Caledon Park (H7543), 17.v.2006, 

KNAA. 

H37 Armagh: Slieve Gullion Forest (J037194), 8.iii.2012, RA. 

H38 Down: near Belfast, AHH (Haliday, 1885; Johnson & Halbert, 1902); Castle Ward Park (J5649), 

9.vii.1992 & 14.vi.2006, KNAA; Mount Stewart (J5570), 27.vi.1992, KNAA; 11.viii.2010, RA; Stormont 

Estate, 20.v.1994, RA; Belvoir Forest, 16.xii.1993, 15.iv.2004, RA; Murlough NNR, 15.xi.2010, RA; 

Portaferry Estate, 30.xi.2011, RA. 

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H39 Antrim: Barnett’s Park, 16.xi.1993, RA; Collin Glen (J270720), 6.ix.2002, KNAA; Great Deer Park, 

Glenarm (D2911), iii.2006, KNAA; Moss Plantation, Shane’s Castle, 16.vii.2010, RA; Langford Lodge 

(J088749), 28.v.2012, RA. 

H40 Londonderry: Springhill Estate (H8683), 25.vi.1992, KNAA; Drenagh Estate (C6923), G australe, 

14.vi.2006, KNAA; Roe Valley Country Park, 16.vii.2011, RA. 

Cis vestitus Mellié, 1849 – Scarce 

Mainly on aerial dead branches on open-grown oak trees, generally with the fungus Peniophora 

quercina; also reported from polypore fungi. For many years only known from the Bullock record from 

Killarney, but recently shown to be widespread on veteran open-grown parkland trees in historic 

demesnes. 

H2 North Kerry: Killarney area, in fungi in woods, EFB (Bullock, 1935); Gortracuisane Oakwood, 

Killarney (V9484), aerial dead branch on oak, 13.x.2001, KNAA. 

H3 West Cork: Glengarriff Castle Demesne (V9455) & Big Meadow (V918570), Glengarriff Woods, 

6.xi.2004, KNAA. 

H33 Fermanagh: Castle Coole Park (H2643), aerial dead branches on parkland oaks, v-x.2006, KNAA; 

Crom Castle Park (H3624), aerial dead branches on parkland oaks, 6.vii.2007, KNAA. 

H36 Tyrone: Baronscourt Park (H3682), v-x.2006, Caledon Deer Park (H7543), vi. vii. & ix. 2006, 

KNAA. 

H38 Down: Castle Ward Park (J5649), aerial dead branches on parkland oaks and common lime, v- 

ix.2006; also from Laetiporus sulphureus, 17.ix.2006, KNAA. 

H39 Antrim: Great Deer Park, Glenarm (D2911), vi-xii.2006 & 22.v.2007, KNAA; Langford Lodge 

(J088749), in crevices in splintered oak bough, 28.v.2012, RA. 

H40 Londonderry: Drenagh Estate (C6923), oak and common lime, 16.vii, 26.vii, 13.xi.2006, KNAA; 

Banagher Glen (C670045), aerial dead branch with Stereum rugosum on hazel, 8.vii.2007, KNAA. 

Cis villosulus (Marsham, 1802) syn. setiger Mellié, 1849 – Rare 

Larvae develop under fungoid bark colonised by Trametes versicolor. Adults feed on hyphae of a wider 

variety of wood-decay fungi. 

H2 North Kerry: Killarney district, “in an old stump”, EFB (Bullock, 1928). 

Ennearthron cornutum (Gyllenhal, 1827) – Rare 

Larvae develop in the fruiting bodies of various bracket fungi. 

H2 North Kerry: Killarney district, “in fungi on rotten wood”, EFB (Bullock, 1928). 

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FAMILY TETRATOMIDAE 

Associated with bracket fungi. 

Hallomenus binotatus (Quensel, 1790) – Rare 

Develops in the fruiting bodies of large polypore fungi in ancient wood pastures, particularly in 

Laetiporus sulphureus. Ireland (O’Mahony, 1941). The single Irish record is curious and may actually 

suggest a casual introduction 

H21 Dublin: Clontarf, in Polyporus squamosus on a dead elm, vi.1941, EO (O’Mahony, 1941). 

Tetratoma ancora Fabricius, 1791 – Rare old growth species 

Larvae under encrusting fungal fruiting bodies on dead and decaying standing stems of hazel and 

other broadleaved trees, generally in deep shade; ancient woodlands. First reported in Ireland in 1939, 

but recently demonstrated to be more widespread than previously appreciated – 4 sites now 

identified. A reliable marker of old growth (Crowson 1987). 

H25 Roscommon: St. John’s Wood, 23.vi.2009, RM det. RA; M9956 & N0056, FITs, 6.viii. & 8.x.2010, 

KNAA. 

H38 Down: Rostrevor, one under bark of a dead Scots pine, viii.1939, CTG (Gimingham, 1942); 

Rostrevor Oakwood NNR (J186173), two knocked from aerial dead hazel poles, 11.vii.2007, KNAA. 

H40 Londonderry: Ness Wood ASSI (C5211), one in flight trap, 7.vii.2007, KNAA. Banagher Glen 

(C670045), two knocked from aerial dead hazel poles, 8.vii.2007, KNAA. 

Tetratoma fungorum Fabricius, 1790 – Recent Arrival 

This species develops successfully, and most commonly, in fruiting bodies of Piptoporus betulinus on 

birch Betula; and has been found in Britain developing in Pleurotus cornucopiae, P. ostreatus, Flammulina 

velutipes, Fistulina hepatica and Paxillus panuoides; adults have also been taken on Inonotus cuticularis, 

Bjerkandera adusta and Polyporus squamosus. The first Irish record was in 1992, but it has been found a 

number of times since, suggesting an expanding population. 

H36 Tyrone: Brantry Lough (H748534), in fallen Piptoporus under birch, 26.i.2012, MDB/RA. 

H37 Armagh: Peatlands Park, 2 + several larvae in Piptoporus betulinus on dead birch, 15.iv.2010, RA; 

Ballymoyer Wood (H974311) larvae in Piptoporus on birch, 2.ii.2012, MDB/RA. 

H38 Down: Carrowdore Castle (J584732), teneral in Hericium cirrhatum on beech stump, 23.x.2002; 

Belvoir Forest, three in decaying Bjerkandera adusta on oak stump, 23.iv.2004, RA (Anderson, 2006); 

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Murlough NNR, 2 under bark in dead elm, 15.xi.2010, RA; Shooter’s Island (J494471), in Flammulina 

velutipes on sycamore, 23.ii.2012, MDB/RA. 

H39 Antrim: Murlough Bay (D189425), off birch polypore in birch woodland, 16.vi.1992, KNAA 

(Alexander & Foster, 1995); Belfast Castle (J326794), adults & larvae in Flammulina velutipes on wych 

elm, 13.iii.2005, RA (Anderson, 2006); Belfast Castle, 8 larvae in Pleurotus cornucopiae on dead wych 

elm, 9.iii.2008, RA. 

Figure 24: Tetratoma fungorum © Roy Anderson 

FAMILY MELANDRYIDAE False Darkling Beetles 

Orchesia micans (Panzer, 1793) – Rare old growth species 

Develops in a variety of large polypore fungi: in lowland Britain especially Inonotus hispidus fruiting 

from standing live mature and older ash trees, but in the west primarily in I. radiatus on standing dead 

alder and birch stems; has also been reported from Fistulina hepatica on live oak; mainly but possibly 

not exclusively in ancient woodland and wood pasture. All Irish records come from I. radiatus 

brackets. 

H1 South Kerry: Kenmare, reared in numbers from a fungus found growing on birch, 1898, JNH 

(Halbert, 1898c; Johnson & Halbert, 1902) – presumably Inonotus radiatus. 

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H33 Fermanagh: Correl Glen NNR (H074546), adult knocked from old I. radiatus on alder snag by 

Sillees River, 5.vii.2007, KNAA; Florencecourt (H178344), several reared from I. radiatus on alder, 

16.iii.2011, MDB det. RA; Finlane, upper Florencecourt (H171325), 2 in I. radiatus on alder, hedgerow, 

20.vii.2012, RA. 

H36 Tyrone: Round Lough (H444483), frequent in I. radiatus on alder, 22.vii.2011, RA; Cullentra 

Lough (H474475), in I. radiatus on alder, MDB. 

H39 Antrim: Great Deer Park, Glenarm (D2911), adult and larvae with old I. radiatus brackets on 

alder, 20.v.2006; Glenarm Woods ASSI (D3010), reared from larvae collected 22.v.2007, parasitic wasps 

hatched as well, KNAA. 

88 

Figure 25: Orchesia micans © Roy Anderson 

Orchesia minor Walker, 1836 – Rare old growth species 

Develops in the fruiting bodies of a variety of wood-decay polypore fungi and possibly certain 

Ascomycetes, mainly on aerial dead branches; especially in permanently damp woodlands, in carr or 

gorge situations. Most often found in ancient woodland and wood pasture. Long only known in 

Ireland from Co. Kerry, but recently shown to be more widespread.


____________________________ 

H2 North Kerry: Muckross Demesne, three specimens of ‘Clinocara tetratoma’ in a fungoid growth 

under the bark of a dead holly, vi.1919, OEJ (Janson, 1920). 

H33 Fermanagh: Correl Glen NNR (H074537), one knocked from dead boughs of up-ended hazel 

stool, 17.v.2007, KNAA. 

H38 Down: Rostrevor Oakwood NNR (J186173), two knocked from aerial dead hazel poles and 

another swept from Luzula, 11.vii.2007, KNAA. 

H39 Antrim: Great Deer Park, Glenarm (D2911), 1 knocked from dead branch in lower canopy of old 

open-grown oak, 27.x.2006, & 1 from FIT on edge of hazel thicket; Glenarm Woods, ASSI (D3011), 2 

knocked from aerial dead hazel poles, 10.vii.2007, KNAA. 

Orchesia undulata Kraatz, 1853 – Uncommon 

Develops in decaying aerial dead branches of oak trees; also reported from other broad-leaved trees to 

some extent. Adults are relatively mobile and occasionally turn up in association with the dead wood 

of a wider variety of tree species outside of the period June/July, and these may have been attracted to 

fruiting wood-decay fungi for feeding (particularly in May) or be merely sheltering between periods 

of activity, especially while over-wintering. Adults are occasionally found at hawthorn and umbel 

blossom. Rare and threatened in central Europe. 

H1 South Kerry: Kenmare, one found in a “mass of fungus-grown leaves”, 1898, JRH (Halbert, 1898c; 

Johnson & Halbert, 1902); Glencar Valley, beaten off birch, vi.1902, HSJD (Donisthorpe, 1902, Bouskell, 

1904); Tomies Wood, Lough Leane (V9088), elytron beneath loose bark on fallen oak branch, 3.vi.1994, 

KNAA (Alexander, 1997b). 

H2 North Kerry: near Killarney, EFB (Bullock, 1914); Muckross Demesne, a few found in a fungoid 

growth under bark of a dead holly, June 1919, OEJ (Janson, 1920); vi.1924, OEJ (Janson & Bonaparte- 

Wyse, 1924). Open-grown oak in field by Reenadinna Wood (V9586), 4.ix.1998, KNAA. 

H4 Mid Cork: Glinny, Riverstick, under oak bark, 17.vii.2000, RA. 

H9 Clare: Mooghaun Lough (R412715), in Malaise trap, vi.2006, TG. 

H16 West Galway: Ballynahinch Woods (L7648), beneath loose bark on fallen oak branch, 3.vi.1994, 

KNAA (Alexander, 1997b). 

H20 Wicklow: Altidore Glen, one swept, vii.1929, JNH (Halbert, 1937); Powerscourt Deerpark, on 

fungi on a felled log, AWS (O’Mahony, 1929b). 

H25 Roscommon: St John’s Wood (N0055 & 0056), two knocked from aerial dead hazel stems in old 

coppices, 8.viii.2010, KNAA. 

H28 Leitrim: Killegar House, under bark of polar logs, 23.x.2010, RA. 

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H33 Fermanagh: Inisherk, Crom (H353244), one in FIT within ancient woodland, 7.vii. – 4.ix.1992, 

APF (Alexander, 1997b); Culliaghs Wood, Crom, in Stereum hirsutum on oak, 21.viii.2004, RA; Castle 

Coole Park (H2643), 17.viii. & 14.ix.2006, KNAA; Rossyvullan Hill, Castle Coole (H266429), under 

bark on beech log, 24.viii.2010, RA. 

H35 West Donegal: Glenveagh Oakwood (C010190), adult at rest on underside of rowan branch, 

14.vi.1992, KNAA (Alexander, 1997b). 

H36 Tyrone: Baronscourt Park (H3682), 2 dead beneath bark on fallen bough beneath open-grown 

parkland oak, 16.v.2006; one swept under ancient oak, 15.vi.2006, KNAA. Caledon Deer Park (H7543), 

vi., vii., & viii.2006, KNAA. 

H37 Armagh: Tynan Abbey Estate, 2 under tight beech bark, 1 under Castanea bark, 1.vii.2011, RA. 

H38 Down: Belvoir Forest, Belfast (J336696) under bark of fallen bough of veteran oak, 7.ii.2012, 

MDB/RA; Leitrim Lodge (J223256), 1 under bark of dead upright Pinus contorta, 29.ii.2012, MDB. 

H39 Antrim: Shane’s Castle, under bark of splintered oak, 1.vii.2010, RA. 

H40 Londonderry: Drenagh Estate (C6923), aerial dead lime branch with Peniophora sp, 5.ix.2006, 

KNAA. 

Abdera flexuosa (Paykull, 1799) – Rare old growth species 

Mainly develops in the small brackets of the wood-decay fungus Inonotus radiatus which decays dead 

standing stems of alder, willow and birch. Also found on I. dryadeus fruiting on oak Quercus in Co. 

Fermanagh and Phellinus pini in Aberdeenshire. The Co. Leitrim records from pine may suggest 

development in the dead persistent annual brackets of Phaeolus schweinitzii which have the same soft 

fibrous consistency as Inonotus spp. Most records are from ancient woodlands and wood pastures. 

Currently only known in Ireland from Cos Fermanagh (where widespread), Tyrone and Leitrim. 

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Figure 26: Abdera flexuosa © Roy Anderson 

H29 Leitrim: Acres Lake (G966100), one emerged 27.v.2011 from a small amount of dead Pinus, 

collected 23.iv.2011, MC; Lough Allen (G964110), in a small amount of dead Pinus sylvestris, 28.v.2011, 

MC. 

H33 Fermanagh: Inisherk, Crom (H357243), four on bracket fungus of Inonotus dryadeus on parkland 

oak, 1.vi.1992, APF (Alexander & Foster, 1995); Crom Park (H32), knocked from I. radiatus on dead 

alder stem in open by Lough Nalughoge, 18.v.2007, KNAA; Molly Mountain (H233282), two from I. 

radiatus brackets on sycamore in open pastureland, 23.vi.1995, RA (Anderson, 1998); Castle Coole 

Park, knocked from I. radiatus brackets on alders, 18.v.2006, KNAA; Correl Glen NNR (H074537), 

knocked from I. radiatus brackets on alders, 17.v.2007; one swept, 5.vii.2007, KNAA. 

H36 Tyrone: Caledon Deer Park (H7543), numerous on old I. radiatus brackets on alder, 16.vi.2006, 

KNAA; Cullentra Lough (H474475), in I. radiatus on alder, 22.vii.2011, MDB det. RA; Brantry Lough 

(H746534), reared from I. radiatus brackets on alder, collected 26.i.2012, MDB/RA; Wood Lough 

(H762603), 1, climbing upright decorticate alder pole, 24.v.2012, RA. 

H39 Antrim: Deramore, Belfast (J333697), reared from I. radiatus brackets on alder, collected 7.ii.2012, 

RA/MDB. 

Melandrya caraboides (Linnaeus, 1761) – Rare old growth species 

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Develops in relatively soft moist white-rotted heartwood of boughs, trunks and stumps; various 

broad-leaved trees, especially ash and beech; mostly associated with ancient woodlands and wood 

pastures, including linear sites such as riverside trees. Very local (Johnson & Halbert, 1902). 

H20 Wicklow: “Powerscourt Deerpark, June 24, in rotten thorn stump plentifully”, Furlong coll. 

(Johnson & Halbert, 1902); Powerscourt Deerpark, some numbers on logs, 20.vi.1928, AWS/EO 

(O’Mahony, 1929b). 

H21 Dublin: Malahide (O24), ARH (Hogan, 1854; Johnson & Halbert, 1902). 

Conopalpus testaceus (Olivier, 1790) – Extinct old growth species 

Develops in aerial dead branches, especially of oak, also hazel; adults may occasionally visit flowers, 

especially umbellifers; associated with ancient wood pastures. Single unlocalised Irish report: two 

taken by Mr Tardy (fide Haliday ms). These are now in the Dublin Museum and Trinity College 

collections and are marked as having been found in Ireland (Johnson & Halbert, 1902). 

FAMILY MORDELLIDAE Tumbling Flower Beetles 

The larvae of most genera in this family develop in galls or the stems of herbaceous plants, but a few 

specialise in decaying wood. 

Mordellistena neuwaldeggiana (Panzer, 1796) – Rare old growth species 

Has been reared in Britain from hornbeam and field maple branch wood in the early stages of decay; 

adults are attracted to blossom; most records are from relict old forest or wood pasture. 

H3 West Cork: Glengarriff, 1924, JNH (O’Mahony, 1929a). 

FAMILY COLYDIIDAE Cylindrical Bark Beetles. 

Mostly predatory. 

Pycnomerus angulatus (Broun, 1893) – Established introduction. 

Under bark and in soft heartwood of dead, mainly coniferous, wood. An import from New Zealand, 

not recorded elsewhere in the Northern Hemisphere. 

H38 Down: Mount Stewart House (J554695), 1 under bark of Pinus radiata logs in carpark, 26.x.2002, 

RA; Sea Plantation, Mount Stewart (J552695), 17 in white-rot debarked logs of Picea abies, 26.vii.2003, 

RA det. PMH; ditto, 3 under sycamore log, 12.x.2011, RA. 

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Pycnomerus fuliginosus Erichson, 1842 – Established introduction 

Under bark of dead trunks and boughs of broadleaved trees; an Australian import. First noted in 

Ireland in 1981. 

H37 Armagh: Derrymore, Newry (J058277), under bark on dead beech, 6.vii.1992, KNAA; Ballymoyer 

Wood (National Trust) (H970311), in powdery frass from Euophryum confine on upright dead Picea 

abies, 2.ii.2012, MDB/RA; Slieve Gullion Forest (J040196), under bark of ash log, 8.iii.2012, MDB/RA. 

H38 Down: Maghery south of Mourne Park, two under bark of a partly decayed alder on a gravelly 

river bank (J268151), 9.v.1981, RA (O’Connor et al., 1983); Rostrevor Oakwood (J186173), on oak logs, 

13.v.2000, RA (Anderson, Simms & Nelson, 2000); on oak, 28.vi.1992, 23.v.2007, KNAA; Derryleckagh 

Wood ASSI (J121253), in powdery rotten oak wood with Euophryum confine, 17.ii.2012, MDB/RA; 

Tullyframe, Mourne Park (J265168), in soft white-rot ash, 27.iii.2012, MDB/RA. 

Synchita humeralis (Fabricius, 1792) – Doubtful 

Feed on fungus-colonised sappy bark or timber, of birch, hawthorn, hazel, alder, and especially beech; 

has been reared from Daldinia in long-established birchwoods. Not included as Irish in Anderson et al. 

(1997). 

H38 Down: near Belfast, AHH (Haliday, 1885), ‘Monotoma juglandis’ in this reference; the specimens 

cannot be traced (Johnson & Halbert, 1902). 

FAMILY TENEBRIONIDAE Darkling Beetles 

Tenebrio molitor Linnaeus, 1758 Mealworm Beetle – Established Introduction 

In stored-products and a domestic pest; also develops in bird nests and in bat roosts where larvae are 

scavengers; occasionally found in decaying timber. In flour stores and houses (Johnson & Halbert, 

1902). 

H8 Limerick: Limerick City, EFB (Bullock, 1914). 

H21 Dublin: JNH (Halbert, 1900b) (Johnson & Halbert, 1902). 

H34 East Donegal: Buncrana, in coastal dunes, C.R. Algeo (Algeo, 1907). 

H39 Antrim: Belfast, in timber yard, CWB (Johnson & Halbert, 1902); Belfast docks, 1942-1946, 

AFO/PMB (O’Farrell & Butler, 1948). 

Palorus ratzeburgi (Wissmann, 1848) – Established Introduction 

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Associated with wood borers (Scolytidae and Bostrychidae) in its natural habitat, but predominantly 

found in stored cereal products and other commodities in Ireland, particularly in mouldy grain 

residues previously attacked by grain weevils; occasionally on imports, more frequently in flour and 

provender mills, grain stores, maltings, etc; widely distributed in Europe. Has been found in decaying 

wood and bracket fungi in Britain in recent years, perhaps responding to climate change. 

H2 North Kerry: golf-links near Ballybunion, P. melinus under bark on a paling, HKGC (Cuthbert, 

1898; Johnson & Halbert, 1902). 

H21 Dublin: Clontarf, viii.1925, EO (O’Mahony, 1926). 

H39 Antrim: Belfast docks, 1942-1946, AFO (O’Farrell & Butler, 1948). 

Nalassus laevioctostriatus (Goeze, 1777) – Common & Widespread 

Develops in decaying timber in ancient woodlands and wood pastures; also in peaty soils of heaths. 

Larvae feed indiscriminately on organic material. Adults nocturnal, feeding on algae encrusting 

timber or lichens on heaths. Mostly short-winged, but with occasional reports of flying individuals. 

Helops striatus “common on heather on high ground, and under fir bark”, Antrim, Down, Galway, 

Louth, Dublin, Wicklow, Carlow, Wexford, Waterford, Cork, Kerry (Johnson & Halbert, 1902). H1, H2, 

H3, H6, H12, H13, H14, H15, H16, H20, H21, H38, H39. 

Gonodera luperus (Herbst, 1783) – Rare 

Adults usually found by sweep-netting in calcareous woodlands; ecology not known; larvae 

presumed to be saproxylic but may not be. 

H2 North Kerry: Killarney district, “a few by brushing young trees close to Ross Castle”, EFB 

(Bullock, 1928). 

H40 Londonderry: Magilligan Sands, one Cistela luperus found in a sand-pocket during a strong wind, 

CWB (Buckle, 1900) (Johnson & Halbert, 1902). 

FAMILY OEDEMERIDAE False Blister Beetles 

This family includes a mix of stem-borers and wood-decay species. 

Nacerdes melanura (Linnaeus, 1758) Wharf Borer – Established Introduction 

In waterlogged timber, especially at waterline zone of coastal and estuarine areas, in pilings, wharves 

& timber supporting riverbanks; also in railway sleepers; larvae develop in flooded timber, mainly 

softwood but occasionally in oak which is damp or wet and in process of fungal decay. 

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H6 Waterford: Tramore Back Strand (S50), 27.vi.1922, OEJ (Janson et al., 1923); Ballyvoyle (X3894), 

sandy coastal bank, 9.vii.1998, MC det. KNAA. 

H12 Wexford: adults in large numbers in house in June; originated in nearby wooden quays? 

(O’Connor, 1981); Raven Point (T118232), in large numbers on dead pine, 11.vii.2011, RA. 

H21 Dublin: Glasnevin Botanic Gardens, on Scrophularia (Hogan, 1854); Dundrum and Phoenix Park, 

JNH (Johnson & Halbert, 1902); Dublin (O1833), abundant in house near sea, June (O’Connor, 1983); 

Portmarnock (02541), in pieces of wood in disturbed sand dune foreshore, 30.vi.2011, MC. 

H28 Sligo: Sligo Docks (G63), under a stone on disused railway track, 11.vii.1993, MC, det. KNAA. 

H34 East Donegal: Foyle district, CWB (Buckle, 1900; Johnson & Halbert, 1902). 

H38/39 Down/Antrim: Sand Quay, Belfast, CWB (Johnson & Halbert, 1902, Supplementary List). 

Ischnomera sanguinicollis (Fabricius, 1787) – Rare old growth species 

Develops in old relatively soft white-rotten wood of wych elms and other broadleaved tree species; 

adults attracted to blossom of hawthorn, field maple, sycamore, oak, lime, and guelder rose. “Irish 

examples … in the collections of the Dublin Museum and Trinity College. According to Haliday’s MS 

list this species was taken by Mr J Hardy in Ireland” (Johnson & Halbert, 1902). Other references: 

Stephens (1832); Curtis (1839). 

H20 Wicklow: Powerscourt Deerpark, one swept from beside the R. Dargle, v.1926, AWS (O’Mahony, 

1929b, & annotated copy of Johnson & Halbert, 1902 in the National Museum, Dublin, acc. Speight, 

1985); deciduous woodland along R. Dargle (O2215), in pupal cell in moist rotting wood within top of 

major split in trunk of large, old, living sycamore, MCDS (Speight, 1985). 

FAMILY PYROCHROIDAE Cardinal Beetles 

Pyrochroa serraticornis (Scopoli, 1763) Red-headed Cardinal Beetle – Rare 

Larvae (Figure 1) develop under bark on freshly dead stems of various broad-leaved trees. Rare in 

Ireland, where known from just a few southern and south-eastern counties – never found repeatedly 

at any location – so with a curious status. The larvae are very distinctive and yet the authors have 

never encountered any; it seems feasible that it is merely a casual introduction with imported timber 

consignments. 

H1 South Kerry: Caragh Lake (V79), larvae in some rotten beech wood and an adult on a fern leaf, 

1903, F. Bouskell (Johnson & Halbert, 1902; Halbert, 1908) 

H5 East Cork: Currabeha (W900 962), 11.vii.2008, TG. 

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H8 Limerick: Adare Demesne, 1863, Furlong (Haliday Ms.) (Johnson & Halbert, 1902); Mount Coote 

(R631274), one swept from bluebells under horse chestnut & beech, 15.v.2000, J Walsh det. JPOC 

(Walsh, 2001). 

H9 Clare: Rineanna, 1 swept off bracken, 21.v.2010, PJH. 

H7 South Tipperary: Knocknanuss, Horse and Jockey (S1449), several on hogweed, semi-natural 

woodland, 25.v.2011, photographed,John Fogarty. 

H12 Wexford: Oaklands Forest, New Ross (S715255), a female crawling on a herb by a small stream 

within the forest, 18.vi.1982, JPOC (O’Connor & O’Connor, 1984); Tintern Abbey, two males on nettles 

and buttercups by an old wall, 15.vi.1990, JPOC (O’Connor & Rigney, 1991); Ballycarney (S9749), 

roadside vegetation, 20.v.1994, JAO (Owen, 1997); Rosslare, on a house wall near a sycamore wood, 

27.v.2005, MC (Cawley, 2009b). 

H13 Carlow: Cloughristick Wood, Leighlinbridge (S702697), male from vegetation by river Barrow 

near the wood, and another on grass on the river bank within the wood, 19.vi.1982, JPOC (O’Connor 

& O’Connor, 1984); Altamont Gardens, by River Slaney (S8665), adult in flight, 9.vi.1991, MAOC; 

Bunclody (S887586), domestic garden, on cotoneaster under pine, 18.vi.2012, Brian Power. 

H14 Laois: Donaghmore (S264797), 4.v.2010, TG. 

96 

Figure 27: Red-headed Cardinal Beetle Pyrochroa serraticornis © Keith Alexander


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FAMILY SALPINGIDAE Narrow-waisted Bark Beetles 

Mainly live under bark on deadwood, though some in small branches and twigs, where adult and 

larva prey on other insects. 

Lissodema denticolle (Gyllenhal, 1813) syn. quadripustulata (Marsham, 1802) – Rare old growth 

species 

In dead branches of a wide variety of trees, mostly broadleaves but including pine, and generally 

where still attached to the tree. No reports since 1924. 

H1 South Kerry: Caragh Lake, ‘common on oak’, vii.1906, JNH (Halbert, 1937); Lickeen, Glencar 

(V7085), vi-vii.1924, OEJ (Janson & Bonaparte-Wyse, 1924). 

H2 North Kerry: Killarney, beaten off oak near the Upper Lake, JNH (Halbert, 1898c) (Johnson & 


Rabocerus foveolatus (Ljungh, 1824) – Rare 

In dead wood; reported from beech and pine. Recorded in Ireland by Halbert (1937a) and not reported 

since. 

H2 Kerry North: Kenmare Demesne, Killarney, one Sphaeriestes foveolatus swept, viii.1925, JNH 

(Halbert, 1937). 

Rabocerus gabrieli Gerhardt, 1901 – Rare 

In dead wood. 

H20 Wicklow: River Dargle near Powerscourt (O2015), sweeping along a streambank, EO (O’Mahony, 

1931); Maulin Mountain, Enniskerry (O1813), beaten from birch trees at 100 ft. on east side, AWS 

(O’Mahony, 1932). 

H39 Antrim: Rea’s Wood, Antrim (J1385), ix.1936, OE (O’Mahony, 1938). 

Sphaeriestes ater (Paykull, 1798) – Rare 

In burnt twigs. Status of early records unclear due to confusion in identification of the genus 

(Anderson, 2009) – ‘may prove to be widespread in eastern Ireland’. 

H2 North Kerry: Dingle (V4499), two tenerals on the shoreline, 30.vi.1971, CAMR det. RA, in UM 

(Anderson, 2009). 

H14 Laois: The Curragh, vi.1938, EO (O’Mahony, 1938). 

H21 Dublin: JAP (Power, 1878). 

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H22 Meath: Balrath, v.1914, GWN (Nicholson, 1915). 

H27 West Mayo: Croagh Patrick, JNH (Johnson & Halbert, 1912). 

H38 Down: Dundrum Bay, vi.1923, OEJ (Janson, 1924). 

Sphaeriestes castaneus (Panzer, 1796) – Established Introduction 

On conifers, in dead and dying branches. No modern reports. 

H2 North Kerry: Muckross Demesne, beating dead boughs, vi.1919, OEJ (Janson, 1920). 

H4 Mid Cork: Kanturk, TVW (Wollaston, 1847) (Johnson & Halbert, 1902). 

H16 North-east Galway: Clonbrock (M7439), JNH (Halbert, 1896) (Johnson & Halbert, 1902). 

H21 Dublin: Rush, HKGC (DNFC, 1894); Santry, JNH (Halbert, 1894; Johnson & Halbert, 1902); 

Clontarf, EO (O’Mahony, 1928b); Howth, EO (O’Mahony, 1928b); St. Annes Park, EO (O’Mahony, 

1924). 

H34/40: Foyle district, CWB (Buckle, 1900) (Johnson & Halbert, 1902). 

H37 Armagh: Little Castledillon, WFJ (mistakenly det. as S. aeratus) (Johnson & Halbert, 1902). 

Sphaeriestes reyi (Abeille, 1874) – Rare 

In burnt twigs. Salpingus aeratus in Johnson & Halbert (1902). Status of early records unclear due to 

confusion in identification of the genus (Anderson, 2009). 

H20 Wicklow: Enniskerry, ix.1931, JNH (Halbert, 1937). 

H21 Dublin: Royal canal at Blanchardstown, Tibradden, JNH (Halbert, 1894, 1895b; Johnson & 

Halbert, 1902); St Annes Park, OE (O’Mahony, 1924); Clontarf, ‘not common’, OE (O’Mahony, 1928b). 

H38 Down: Bloody Bridge, Mourne Mountains (J380272), one swept off burnt Western Gorse, 

14.v.2007, RA (Anderson, 2009b). 

H39 Antrim: Cave Hill, CWB det. GCC (Johnson & Halbert, 1902). 

Vincenzellus ruficollis (Panzer, 1794) – Uncommon 

Under bark on various broad-leaved trees, especially hawthorn & beech. Previously known as 

“Rhinosimus viridipennis”. 

H1 South Kerry: Kenmare, vii.1898, JNH (Halbert, 1898c; Johnson & Halbert, 1902). 

H21 Dublin: Santry, on moss on old tree stump, JNH (Halbert, 1894); Dundrum, under elm bark, JNH 

(Halbert, 1900b) (Johnson & Halbert, 1902); Malahide, EO (O’Mahony, 1928b); Clontarf, 1 under bark, 

EO (O’Mahony, 1935b). 

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H25 Roscommon: Mote Park, vi.1897, JNH; Mount Talbot, JNH (Halbert, 1898a; Johnson & Halbert, 

1902); St. John’s Wood, on ash, canopy fogging, 23.vi.2009, RM. 


H31 Louth: Bellurgan Forest (J096099), under bark of dead sycamore, 22.iii.2012, MDB/RA. 

H33 Fermanagh: Inisherk Island, Crom, FIT 4.ix-5.x.1992, APF. 

H34/40: Foyle district, CWB (Buckle, 1900) (Johnson & Halbert, 1902). 

H38 Down: Derrymore Park, Bessbrook (J0528), 6.vii.1992, KNAA; Newtownards, 1 swept from 

saltmarsh grassland, 24.vi.1998, RA; Castle Ward Park (J5649), 21.v.2006, KNAA. 

H39 Antrim: Newforge, Lagan Canal, under bark of Nothofagus, 20.x.1992, RA; Deramore, in Stereum 

hirsutum on beech log, 11.xi.1992, RA; Lagan Meadows, under fungoid bark of beech logs, 27.xi.1992, 

RA; Lady Dixon Park, 11 under bark with Stereum on beech logs, 11.i.2005, RA. 

Salpingus planirostris (Fabricius, 1787) – Common & Widespread 

Under bark on various broad-leaved trees in early stages of decay; normally saprophagous, but will 

also feed on insect larvae. Widespread: Derry, Antrim, Down, Armagh, Galway, Dublin, Wexford, 

Cork, Kerry (Johnson & Halbert, 1902). Common, at least in the north. H3, H5, H12, H16, H18, H21, 

H25, H27, H28, H33, H36, H38, H39, H40. 

Salpingus ruficollis (Linnaeus, 1761) – Widespread 

As above, less common (Johnson & Halbert, 1902). Mainly in the north. 

H9 Clare: Blackweir Bridge, Newmarket-on-Fergus (R389715), 26.v. 2003, RCW (Regan & Anderson, 

2004) 

H16 West Galway: Connemara National Park (L75), 1994 (Speight, 2004). 

H21 Dublin: Santry, JNH (Halbert, 1894; Johnson & Halbert, 1902); Clontarf, Howth, EO (O’Mahony, 

1928b); St. Annes Park, 29 in tree stump, EO (O’Mahony, 1936). 


H25 Roscommon: St John’s Wood (N0056), FIT, 6.viii.2010, KNAA. 

H27 West Mayo: Westport, JJW (Fowler, 1891) (Johnson & Halbert, 1902) 

H33 Fermanagh: Belleisle (Porter, 1898) (Johnson & Halbert, 1902); Inisherk, Crom, FIT 4.ix-5x.1992, 

APF. Castle Coole Park (H2643), FIT, 25.x.2006, KNAA. 

H36 Tyrone: Baronscourt Park (H3682), FITs, various dates, 2006, KNAA. 

H37 Armagh: Palace Grounds, Dean’s Hill, WFJ (Johnson, 1888b, 1892a) (Johnson & Halbert, 1902). 

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H38 Down: Tollymore Park (Patterson, 1885); Lough Neagh, Haliday ms. (Johnson & Halbert, 1902); 

Newforge, HLO det. WFJ (Johnson, 1914); Kircassock, under sycamore bark flakes, 27.xi.1976; 

Derrymore Park, Bessbrook (J0528), 6.vii.1992, KNAA; Tollymore Forest, in a small Piptoporus on 

birch, 9.x.1993, RA; Ballyskeagh, under bark of Tilia, 8.xii.1992, RA; Giant’s Ring, Edenderry, under 

sycamore flakes, 24.x.1992, RA; Castle Ward Park (J5649 ), 28.x.2006, KNAA; Dundrum Castle, under 

fungoid bark on horse chestnut, 20.xvii.2009, RA; Shooter’s Island (J494470), 23.ii.2012, RA. 

H39 Antrim: Ballycastle, BT (Johnson & Halbert, 1902); Newforge Lagan Canal, under bark of 

Nothophagus, 20.x.1992; Belfast Castle, under fungoid beech bark, 11.vii.2004, RA; Lady Dixon Park, 

Belfast, under fungoid beech bark with Stereum, 11.i.2005, RA; Great Deer Park, Glenarm (D2911 ), 

16.ix.2006, KNAA; Breen Wood (D1233), FIT, 9.vii.2007, KNAA. 

H40 Londonderry: Drenagh Estate (C6923), 19.v. & 15.ix.2006, KNAA. 

FAMILY SCRAPTIIDAE 

Most, if not all, develop in dead wood; larvae of some have been found below loose bark; feed largely 

on wood fibres and fungi, although will take animal food; adults frequent flowers. 

Anaspis fasciata (Forster, 1771) syn. humeralis (Fabricius, 1775) – Rare 

Has been reared in numbers in Britain from dead fallen branchwood of oak, hornbeam and larch; 

frequently found at hawthorn blossom, also umbelliferous flowers. 

H1 South Kerry: Kenmare, ‘A. geoffroyi’ found on privet, JRH (Halbert, 1898c). 

H33 Fermanagh: Lough Yoan, Castle Coole, 2.vi.1992, KNAA. 

Anaspis frontalis (Linnaeus, 1758) – Widespread 

Has been reared from deadwood of oak and maple in Sweden; frequently found at hawthorn blossom. 

Widespread but not common: Ulster, Connaught, Leinster, Munster (Johnson & Halbert, 1902). 

H1 South Kerry: Kenmare, vii.1898, JNH (Halbert, 1898c); Lickeen Wood (V7086), 4.v.2010, KNAA. 

H2 North Kerry: Cloghereen Wood, Muckross (V9786), cut grass in parkland, 8.vii.1996, JAO (Owen, 

1997). 

H7 South Tipperary: Mallow (W5298), roadside vegetation, 6.vii.1996, JAO (Owen, 1997). 

H9 Clare: Slieve Carron (M3303), 9.v.2010, KNAA. 

H18 Offaly: Charleville Wood (N315226), 24.v. 2003, RCW (Regan & Anderson, 2004). 

H23 Westmeath: Annagh near Athlone (N093436), 20.v.2012, Oscar Vorst. 

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H25 Roscommon: St John’s Wood (M9956 & N0056), widely, 10.vi.2010, KNAA. 

H37 Armagh: Dean’s Hill, Armagh, WFJ (Jphnson, 1892a); Peatlands Park, at haw flowers, 29.v.2000, 

RA (Anderson, Simms & Nelson, 2000); Derryinver, Lough Neagh, sweeping Filipendula, 14.viii.1996, 

RA; Derrymagown, Argory, haw flowers, 4.v.1996, RA; Loughgall Forest NR, on oak foliage, 

14.v.2005, RA. 

H39 Antrim: Greenmount Agricultural College, beating Cotoneaster, Rea’s Wood, sweeping, 30.vi.1986, 

RCW (Welch, 1986); Shane’s Castle, at Tilia flowers, 1.vii.2010, RA. 

H40 Londonderry: Banagher Glen (C670045), on flowers, 20.v.2007, KNAA. 

Anaspis garneysi Fowler, 1889 – Scarce 

Has been reared from dry wood mould of oak, beech & larch (Levey 2009). The record Dublin 

(Halbert, 1893) is to be deleted (Johnson & Halbert, 1902). Widely found in Northern Ireland by flight 

trapping during 2007. 

H2 North Kerry: Killarney area, by general sweeping, EFB (Bullock, 1932). 

H18 Offaly: Bullock Island, Shannon Callows (N026181), 24.v. 2003, RCW (Regan & Anderson, 2004). 

H33 Fermanagh: Crom Castle Park (H3624), hawthorn blossom 18.v., & Inisherk (H357248), FIT, 

23.v.2007, KNAA; Old Crom Castle (H364239), swept, 24.v.2012, K.A. Bryan. 

H37 Armagh: Loughgall Forest NR, on haw flowers, 14.v.2005, RA. 

H38 Down: Rostrevor Oakwood NNR, on sycamore foliage, 21.v.2005, RA; Rostrevor Oakwood NNR 

(J1817), 23.v., & FIT, 11.vii.2007, KNAA. 

H39 Antrim: Great Deer Park, Glenarm (D2911), 13.vi.2007; Glenarm Woods ASSI (D3010), 10.vii.2007, 

KNAA. 

H40 Londonderry: Ness Wood ASSI (C5211), 7.vii.2007; Banagher Glen (C670045), FIT, 8.vii.2007, 

KNAA. 

Anaspis lurida Stephens, 1832 – Rare 

Has been reared from dead fallen branchwood of oak, alder, aspen, elm, hazel, larch & from birch 

polypore; larvae possibly epiphytic rather than saproxylic (Levey, 2009). Adults attracted to blossom, 

including elder. 

H3 West Cork: Glengarriff Castle Demesne (V9455), in Malaise trap, 26.v.2005, coll. TG, det KNAA. 

H23 Westmeath: Whinning Wood, Portlick (N0548), 14.v.2010, KNAA. 

H39 Antrim: Cullybacky, WFJ (Johnson, 1891) ‘A. subtestacea’ (Johnson & Halbert, 1902); Rea’s Wood, 

Antrim, sweeping carr, 30.vi.1986, RCW (Welch, 1986). 

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Anaspis maculata (Geoffroy in Fourcroy, 1785) – Common & Widespread 

Develops in small girth branchwood of a wide variety of broad-leaved trees; frequent at blossom. 

Common: Ulster, Connaught, Leinster, Munster (Johnson & Halbert, 1902). H1, H2, H4, H5, H6, H9, 

H15, H18, H21, H23, H33, H34, H35, H36, H37, H38, H39, H40. 

Anaspis regimbarti Schilsky, 1895 – Common & Widespread 

Has been reared from a larva found in a decaying oak log, and from large girth oak branchwood. 

‘Anaspis ruficollis’ (syn.) common: Ulster, Connaught, Leinster, Munster (Johnson & Halbert, 1902). H1, 

H2, H3, H9, H15, H18, H33, H34, H35, H36, H37, H38, H39, H40. 

Anaspis rufilabris (Gyllenhal, 1827) – Widespread 

Has been reared from large girth oak branchwood in England. ‘Only known from Ulster’ (Johnson & 

Halbert, 1902) – this has proved mistaken. 

H1 South Kerry: Tomies Wood (V9089), 26.v.1994, KNAA. 

H3 West Cork: Lissangle, Skibereen (W1040), 4.vi.1996, KNAA. 

H9 Clare: Blackweir Bridge, Newmarket-on-Fergus (R389715), v. 2003 (Regan & Anderson 2004) 

H18 Offaly: Charleville Wood (N315226), v. 2003 (Regan & Anderson 2004). 

H23 Westmeath: Whinning Wood, Portlick (N0548), 14.v.2010, KNAA. 

H25 Roscommon: St John’s Wood (M9956 & N0056), 7.vi & 7.viii.2010, KNAA. 

H28 Sligo: Sriff Point, Lough Gill (G788339), 17.vi.1999, KNAA. 

H33 Fermanagh: Castle Coole Park (H2643), 2.vi.1992, 17.vi.2006 & 18.v.2007; Correl Glen (H074537), 

17.v.2007; Crom Castle Park (H3624), 18.v.2007, KNAA. 

H35 West Donegal: Rathmullan, CWB det. GCC (Johnson & Halbert, 1902) 

H36 Tyrone: Baronscourt Park (H3682), 15.vi. & 25.vii.2006, KNAA; Wood Lough (H762603), 

24.v.2012, RA. 

H37 Armagh: WFJ det. GCC (Johnson & Halbert, 1902); Peatlands Park (H897613), 11.vi.2012, K.A. 

Bryan. 

H38 Down: Rostrevor Oakwood (VJ1817), 23.v.2007, KNAA. 

H39 Antrim: Lagan Canal near Lisburn, coll. CWB det. GCC (Johnson & Halbert, 1902); Great Deer 

Park, Glenarm (D2911), 13.vi.2006; Glenarm Woods ASSI (D2911), 22.v. & 10.vii. 2007, KNAA; Breen 

Wood (D1233), 21.v2007, KNAA. 

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H40 Londonderry: Springhill Park (H8683), 25.vi.1992; Banagher Glen (C670045), 20.v. & 8.vii.2007; 

Ness Wood ASSI (C528118), 2.vii.2007, KNAA. 

[Anaspis thoracica (Linnaeus, 1758) 

Ireland (Bullock, 1935, according to Anderson et al., 1997, but this is an error).] 

FAMILY CERAMBYCIDAE Longhorn Beetles 

Many exotic longhorns turn up in timber yards and buildings due to importation of timber. Some 

have become established. Obvious casuals are not included. Speight (1988) provides some Irish threat 

statuses. 

Rhagium bifasciatum Fabricius, 1775 – Common & Widespread 

Develops in the cut stumps of conifers; it probably occurs wherever there are conifer plantations in 

Ireland. It is also frequent in rotten stumps of various broadleaved trees, including oak and alder; also 

found living in 2000 year old pine stumps exposed from beneath bogs (Speight, 1988). Common & 

widespread in Ireland. “Common in fir plantations” (Johnson & Halbert, 1902). H1, H2, H3, H4, H5, 

H6, H12, H14, H16, H17, H18, H20, H25, H27, H28, H29, H30, H33, H34, H36, H37, H38, H39, H40. 

Figure 28: Rhagium bifasciatum (left) and Rhagium mordax (right) © Roy Anderson 

Rhagium mordax (De Geer, 1775) – Widespread 

Larvae develop in decaying timber, preferring the cambium and outer sapwood of rotting boles or 

stumps; most often found in oak, but also in a wide range of other broad-leaved trees. ‘R. inquisitor’ 

“not uncommon on oak, beech and hawthorn, in old woods in the north and west of Ireland, but rare 

in the east” (Johnson & Halbert, 1902). Widely distributed in Ireland (Speight, 1988). The absence of 

reports from the south and east appears to hold true. 

H1 South Kerry: Tomies Wood (V9089), 1988 & 1990; Uragh Wood (V8362), 1994, KNAA. 

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H2 North Kerry: nr. Killarney, EFB (Bullock, 1914); W.F. de V. Kane (Fowler, 1891); Waterville, J.W. 

Yerbury; Muckross, ARF (Johnson & Halbert, 1902); Muckross Park, 1998, KNAA; Derrycunihy Wood 

(V8981 & 9181), 1990, KNAA; Galways River, Upper Lake, 1994, JAO; Galways Bridge (V914803), 

1994, KNAA; Killarney National Park (V9490), 22.iv.1999, MC; Muckross Woods, 2011, RA; Ladies 

View, 2011, RA. 

H3 West Cork: Glengarriff Woods (V918570), J.W. Yerbury (Johnson & Halbert, 1902); 1988 & 1996, 

KNAA; V912572, beaten off hawthorn, 19.v.2012, BN. Glengarriff Castle Demesne (V946553), 2005, TG. 

H7 South Tipperary: Shanbally Castle Estate (R977165), 17.iii.2012, RA. 

H9 Clare: Arduan Lough (R485706), v.vi.2006, & Derrygeeha Lough (R150564), v.vi.2007, Malaise 

traps, TG. 

H14 Laois: Abbeyleix, JMB (Browne, 1901) (Johnson & Halbert, 1902). 

H15 South-east Galway: Punchbowl, Gort (M4500), Caher Wood, Lough Graney (R5591), & 

Derrykeel Wood (R5199), 1988, KNAA Portumna Bay, The Quay, Park, etc (M841032), 2003 (Regan & 

Anderson, 2004); Scarriff Lough (R413935), 2007, TG; Garryland Wood, 2007, RA. 

H16 West Galway: Connemara National Park (L75), 1994 (Speight, 2004). 

H18 Offaly: Charleville Wood (N315226), 2003, RCW (Regan & Anderson, 2004). 

H20 Wicklow: Powerscourt, 1919, JNH (Halbert, 1922); Powerscourt Deer Park (O1912), 1993, KNAA; 

Powerscourt, 2007, RA. Ballard Woods, Glendalough (T1495), 1993, KNAA. 

H23 Westmeath: Hare Island, Lough Ree, JNH (Halbert, 1900a); 1977, MCDS. 

H25 Roscommon: Cloonca Wood, JNH (Halbert, 1898a) (Johnson & Halbert, 1902). 

H27 West Mayo: Crossmolina, W.F. de V Kane (Fowler, 1891) (Johnson & Halbert, 1902). Barnarinnia 

Woods, Lough Mask (M0660), 1992, KNAA. 

H28 Sligo: W.F. de V. Kane (Fowler, 1891) (Johnson & Halbert, 1902); Killerry (G7732), 23.v.1993, MC; 

Rathcarrick (G636347), 13.ii.1994, MC; Ardcree (G6422), MC; Union Wood (G680285), 1999, KNAA & 

5.v.2011, MC. 

H29 Leitrim: Glencar Woods (G7543), 1998, KNAA. 

H33 Fermanagh: Ballinamallard (Johnson & Halbert, 1902); Correl Glen, 1988, RA; Shane’s Castle, 

1988, RA; Castle Caldwell FNR (H0260), 1992, KNAA; Crom Castle Estate (H32), 1992, 2002, 2007, 

KNAA; Isle Namanfin, 1993, RA; Florencecourt, 1997 & 2010, RA; Inisherk Island, 2000, RA; Castle 

Coole Park (H2643), 2006, KNAA; Rossyvullan, Castle Coole, 2010, RA. 

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H34 East Donegal: Ardnamona Wood, Lough Eske (G9685), 1992 & 1999, KNAA; Glenveagh 

Oakwood (C010190), 1992, KNAA. 

H35 West Donegal: Rathmullan, JJW (Johnson & Halbert, 1902). 

H36 Tyrone: W.F. de V. Kane (Fowler, 1891; Johnson & Halbert, 1902). Caledon Deer Park (H7543), 

2006, KNAA. 

H37 Armagh: Tynan Abbey Estate, 2011, RA 

H38 Down: Tollymore Park, WFJ (Johnson & Halbert, 1902). 

H39 Antrim: Great Deer Park, Glenarm (D2911), 2006, KNAA; Glenarm Woods ASSI, 2007, KNAA; 

Moss Plantation, Shane’s Castle, 2010, RA; Glenarm Nature Reserve, 2011, RA. 

H40 Londonderry: Walworth Wood, CWB (Buckle, 1900; Johnson & Halbert, 1902). Ness Wood ASSI 

(C528118), 2007, KNAA. 

Grammoptera abdominalis (Stephens, 1831) syn. variegata (Germar, 1824) – Rare 

Larva feed on the decayed outer sapwood of dead aerial branches of oak, pear and sweet chestnut, 

boring deeper to pupate; preferred fungal host is Vuilleminia comedens (M. Rejzek, in lit.); adults 

attracted to blossom. 

H38 Down: near Belfast, 1885, Haliday – “an Irish specimen [of G. analis] in Haliday collection” 

(Johnson & Halbert, 1902); Murlough NNR (J410352), several on haw flowers, 17.v.2005, RA. 

Grammoptera ruficornis (Fabricius, 1781) – Common & Widespread 

Larvae in dead twigs and decaying small branches of many broad-leaved trees; adults attracted to 

blossom. Common in Ireland (Johnson & Halbert, 1902); one of Ireland’s most widely distributed and 

most frequently encountered longhorn beetles (Speight, 1988). H1, H2, H3, H4, H5, H7, H9, H12, H14, 

H15, H16, H17, H18, H20, H21, H22, H23, H25, H27, H28, H29, H30, H33, H35, H36, H37, H38, H39, 

H40. 

Grammoptera ustulata (Schaller, 1783) – Rare old growth species 

Larvae have been reared from dry dead or mouldy lichen-covered twigs of field maple and oak, also 

known from other broad-leaved trees, where they feed on the outer sapwood; adults at blossom. 

Discovered in Ireland in 2003 but assumed to be an overlooked native. 

H18 Offaly: Charleville Wood (N320227), one swept under beech, 24.v.2003, RCW (Regan & 

Anderson, 2004). 

Leptura aurulenta (Fabricius, 1793) Hornet Beetle – Rare 

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Larvae develop in the cambial layer of large sections of freshly dead wood, where in open sunny 

situations; broad-leaved trees, especially of oak. In Ireland largely confined to Counties Kerry, Cork 

and Wicklow. ‘Vulnerable’ in Speight (1988). 

H1 South Kerry: “This area was visited by a large party of naturalists in July 1898 none of whom 

reported this beetle, so perhaps it colonised from Glengarriff subsequently” (Stelfox, 1937); Slaheny 

Bridge, Kilgarvan, one at a stump in the woods, 31.vi.1935, several more, 1.vi.1935, AWS (Stelfox, 

1937); near Drumanassig Bridge (V9587), between Kenmare and Glengarriff, commonly at stumps of 

beech by the roadside, 6.vii.1935, AWS (Stelfox, 1937); Dunkerron Castle, west of Kenmare, on beech 

stumps, 7.vii.1935, AWS (Stelfox, 1937); Derreen district, southern side of Kenmare Bay, 1935, S.W. 

Kemp (Stelfox, 1937); Kenmare, vii.1945, EFB (Speight, 1988); Ardtully (V9973), 5.vii.1976, M. de 

Courcy Williams (Speight, 1988); Cloonee Loughs (V8362), on Rubus flowers in clearing in oak woods, 

22.vii.1978, MCDS (Speight, 1988). 

H2 North Kerry: Dungeel (V8195), near Killorglin, two from felled alder, 22.vii.2001, M. O’Sullivan 

(O’Sullivan, 2003); Muckross Woods (V98), 1 at umbel, 20.viii.1992, RN. 

H3 West Cork: Glengarriff (V95), “First taken …many years ago by A. Furlong and E.P. Wright; and 

recently W.F. deV. Kane captured 2 ... presented to the Dublin Museum” (Johnson & Halbert, 1902); 

Lord Bantry’s Demesne (Kenmare Demesne), W.F. de V. Kane (Fowler, 1891); common at Glengarriff 

in vii.1924, flying about tree stumps, mostly of oak and birch, and to bramble flowers quite close to the 

village, Halbert (1937a); extraordinarily abundant around Glengarriff, vii.1935 outnumbered R. 

maculata in the oakwoods about The Lodge (Stelfox, 1937); on Kenmare side of Glengarriff (V95), 1950, 

& Barley Lake (V85), 1952, H.C. Huggins (Huggins, 1953); not uncommon at Glengarriff; collected a 

series for the BMNH in 1952; at flowers of purple loosestrife and ragwort, but most taken on an oak 

stump on the side of the Kenmare Road, H.C. Huggins (Huggins, 1960); Glengarriff (V9157), on 

Oenanthe flowers in clearing in mixed woods, 22.vii.1976, MCDS (Speight, 1988); Glengarriff Castle 

Demesne (V9455), flying about stumps of oak and birch, vi.1924, JNH (Halbert, 1937); 2005, in Malaise 

traps, TG; 2012, one photographed in nature reserve, per Clare Heardman. 

H6 Waterford: Glenmore, in Blackwater valley, between Lismore and Fermoy, two, 29.vii.1928, AWS 

(Stelfox, 1937). 

H8 Wexford: Vinegar Hill, Enniscorthy (S9840), 16.vii.2012, Lynn Cheeseman, det. RA. 

H20 Wicklow: Rathdrum Forest (T18), 1924, EO (O’Mahony, 1924); Shelton Abbey, NW of Arklow, on 

tree stumps, vii.1926, JNH (Halbert, 1937); Clara Bridge, flying to Angelica, 12.vii.1926, JNH (Halbert, 

1937); The Meeting of the Waters, many dead in a beech stump; by no means rare from Rathdrum 

down the Avonmore valley to near Arklow (Stelfox, 1937); Avondale Forest Park (T1985), mixed 

woods, 23.vi.1975, MCDS (Speight, 1988). 

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Leptura quadrifasciata (Linnaeus, 1758) Four-banded Longhorn – Rare 

Develops in dead and decaying stumps, especially birch, but also a wide range of other broad-leaved 

trees, and once reared from spruce; larvae make meandering galleries deep in the sapwood; wood 

sizes down to 15cm diameter, wet or dry, but dry areas needed for pupation; adults sun-loving and fly 

on hot sultry days, attracted to blossom. NB Leptura aurulenta often mis-deteremined as this species by 

people not familiar with it. ‘Vulnerable’ in Speight (1988). 

H2 North Kerry: Galways River at Upper Lake (V9081), Killarney, 16.v.1994, JAO (Owen, 1997). 

[H3 West Cork: Glengarriff, in Lord Bantry’s Demesne, 12 taken in half hour on ragweed, by W.F. de 

V. Kane (Johnson & Halbert, 1902); Halbert (1937a) comments on his inability to find a trace of this 

species here; Stelfox (1937) queries the identification since it has not been there since]. 

H9 Clare: Castletown Lough (R388777), Cragmoher Lough (R293867) & Lough Ardnamurry 

(R381797), Malaise traps, vi.viii.2006, TG; Ballycar, Newmarket-on-Fergus (R419619), in rural garden, 

24.vi.2012, John Murphy det. RA. 

H10 North Tipperary: near Rathcabban (N0007), dry peat bog with birch trees, 9.vii.1996, JAO (Owen, 

1997). 

H15 South-east Galway: Portumna (M8303), elytron in rotten birch, mixed woods beside lake, 

15.ii.1975, MCDS (Speight, 1988). 

H18 Offaly: All Saints Bog (N0010), in birch woods, MCDS (Speight, 1988). 

H20 Wicklow: Clara Bridge, one flying to Angelica, 12.vii.1926, JNH (Halbert, 1937); Lake Park, Lough 

Dan, 1924, EO (O’Mahony, 1924); repeatedly seen in county on various dates between 12.vii. & 4.viii., 

either in the Vale of Clara or near Coolballintaggart in the Ow Valley (Stelfox, 1937); west of Kippure 

House, upper Liffey valley, female egg-laying in crevices of a stump of a beech tree, 16.viii.1939, Mrs 

Stelfox (Stelfox, 1943). 

H35 West Donegal: Ards Forest (C0834), deciduous woodland, 11.viii.1978, M. de Courcy Williams 

(Speight, 1988); Ards Forest (C0734), 12.viii.1978, RA. 

Anoplodera sexguttata (Fabricius, 1775) – Rare; old growth species 

Develops in red-rotten oak wood in shade or semi-shade, generally lying logs in moist conditions; 

adults most often seen nectaring at flowers of plants such as hogweed and water dropwort; often in 

small glades in dense forests. ‘Endangered’ (Speight, 1988). 

H2 North Kerry: two near Muckross Abbey, 1859, E. Birchall, in J.R. Hardy coll.; Dinish Island, near 

Muckross, male swept off flowers, summer 1879, JRH (Johnson & Halbert, 1902). 

Alosterna tabacicolor (De Geer, 1775) – Uncommon 

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Larvae develop in old damp rotten stumps of hazel, hornbeam, maple, pine, etc, often boring deeply 

into sapwood; adults attracted to blossom. Local in Ireland ((Johnson & Halbert, 1902); possibly a 

relict ancient woodland species (Alexander, 1997a). 

H1 South Kerry: from rowan blossom in large pine wood near Glenbeigh (V6690), vi.1902, 

(Donisthorpe, 1903) (Bouskell, 1904). 

H2 North Kerry: Killarney, viii.1912, etc, EFB (Bullock, 1914; Speight, 1988). 

H5 East Cork: Owennacurra River (W8576), open areas along wooded riverside, 4.vi.1999, KNAA. 



H14 Laois: Abbeyleix, JMB (Browne, 1901) (Johnson & Halbert, 1902); Durrow (S410790), 14.vi.2010, 

MC. 

H20 Wicklow: Delgany, WES (Johnson & Halbert, 1902); Glen of the Downs (O2610), oak woods, 

11.vii.1979, D. Dowling (Speight, 1988); (O2215), on umbel flowers in deciduous woods, 14.vi.1975, 

MCDS (Speight, 1988); Devil’s Glen, 14.vii.1986, RA. 

H21 Dublin: Glasnevin, etc, ARH (Hogan, 1854); Barnacullia, HKGC (Cuthbert, 1892a) (Johnson & 


H25 Roscommon: Mote Park, JNH (Halbert, 1898a) (Johnson & Halbert, 1902); St. John’s Wood 

(M9956), 5.vi.1999 & 7.vi.2010, KNAA. 

H28 Sligo: Slish Wood (G740317), 12 & 17.vi.1999, KNAA; 25.vi.2009, MC (Cawley, 2009a). 

H33 Fermanagh: Reilly Wood, Crom (H337253), 28.v.1992; Gole Wood, Crom (H333252), 1.vi.1992, 

KNAA (Alexander, 1997a). 

H38 Down: near Belfast, AHH (Haliday, 1885) (Johnson & Halbert, 1902). 

H39 Antrim: Great Deer Park, Glenarm (D2911), 13.vi.2006; Glenarm Woods Nature Reserve, 

22.v.2007, KNAA & 14.vi.2011, 20.vi.2012, RA. 

Rutpela maculata (Poda, 1761) – Widespread 

Develops in moist decaying wood of stumps and roots of broad-leaved trees and pine, but particularly 

birch; adults attracted to blossom. Probably generally distributed in Ireland if somewhat local in 

certain districts (Stelfox, 1937). ‘Strangalia armata’ (Johnson & Halbert, 1902). 

H1 South Kerry: Kenmare, vii.1898, JNH (Halbert, 1898c); Waterville, coll. Yerbury (Johnson & 

Halbert, 1902); Glencar, vii.1902, HSJD (Donisthorpe, 1903); Derrycunihy, vii-viii.1906, LHBW 

(Bonaparte-Wyse, 1906); Lickeen Wood, Glencar, viii.1910, LHBW (Bonaparte-Wyse, 1911). 

108


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H2 North Kerry: Killarney (Hogan, 1855); Upper Lake & Kenmare, JNH (Halbert, 1898c); V9088, oak 

woods, 20.vi.1975, MCDS; V8362, on Rubus flowers, deciduous woods, 27.vii.1976, MCDS; Camillan 

Wood , Muckross (V9486), 12.viii.1990, KNAA (Alexander, 1997a). 

H3 West Cork: Glengarriff Castle (V948555), Furlong ms & Kane (Johnson & Halbert, 1902); 2005 & 

2006, TG; Glengarriff (V9157), 6.v.1975, MCDS; near Ballyoughtera (W0930), 13.vii.1983, MCDS 

(Speight, 1988); banks of Bandon River (W550556), 24.v.1998, MC; banks of River Sullane, Macroom 

(W339732), 30.vi.2010, MC. 

H5 East Cork: Ballygrunna (X018808), 21.vi.1998, MC. 

H6 Waterford: Ballymacmague Fen, 26.vii.2009, RA; Cappagh Loughs, 27.vii.2009, RA. 

H9 Clare: St. Senan's Lough (R046542), Malaise trap, vi.vii.2007, TG. 

H11 Kilkenny: Thomastown and Woodstock, JNH (Johnson & Halbert, 1902). 

H13 Carlow: plentiful in Mr Kavanagh’s Demesne near Borris, PF (Johnson & Halbert, 1902). 

H16 Galway: Derryclare Wood (L8349), on meadowsweet flowers in oak wood, 30.vii.1975, MCDS 

(Speight, 1988). 

H18 Offaly: Charleville Wood (N3122), oak/ash woods, 5.vii.1984, MCDS (Speight, 1988). 

H20 Wicklow: Powerscourt, ARH (Hogan, 1854); Avoca, Furlong ms.; Devil’s Glen, AHH (Haliday, 

1857); R. Dargle, JNH (Halbert, 1900b); Powerscourt Demesne (O2215), 27.vi.1976, MCDS; Avondale 

Forest (T1985), 24.viii.1978, MCDS; nr. Kilmacanogue (O2513), alder/willow swamp, 27.vii.1981, 

MCDS (Speight, 1988); Devil’s Glen, 14.vii.1986, RA; Powerscourt (17.viii.1986, RA . 

H23 Westmeath: Hare Island, L. Ree, vi.1976, MCDS (Speight, 1977). 

H25 Roscommon: N0055, 20.vi.1976, MCDS (Speight, 1988). 

H28 Sligo: Rockwood, Lough Gill, 1901, DNFC (DNFC, 1901); Hazelwood (G712360, 27.vi.1995, MC; 

Killerry (G7733), 9.viii.2009, MC; Union Wood (G682273), 10.viii.2009, MC. 

H27 West Mayo: Knockmore, Pontoon, 1907, LHBW (Bonaparte-Wyse, 1907). 

H29 Leitrim: Dromahair, vi.1957, CAC (Collingwood, 1959). 

H30 Cavan: Gartanoul, 16.iv.1940, RCF (Speight, 1988). 

H33 Fermanagh: Inishdivann, L. Erne, 9.viii.1936, anon., UM. 

H38 Down: near Belfast, 1885, AHH (Johnson & Halbert, 1902); Tollymore Forest, 9.vii.1976, 

16.viii.1985, 6.viii.1986, RA. 

Stenurella melanura (Linnaeus, 1758) – Doubtful 

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Larvae have been found in thin, decayed oak branches and in broom roots; also in other trees. Irish 

example in Haliday collection, but no locality data (Johnson & Halbert, 1902). It would seem unlikely 

that a resident population of a large and colourful beetle would be overlooked for so long. 

Asemum striatum (Linnaeus, 1758) – Established introduction 

Mainly develops in pine stumps and exposed roots of freshly cut trees, but will use larch and spruce; 

has spread with softwood plantation forestry. First noted in Ireland in 1973 (Speight, 1976). 

H14 Laois: Abbeyleix (S4382), in rotten conifer log at edge of young conifer plantation on drained 

valley bog, 2.vi.1973, MCDS (Speight, 1976). 

H20 Wicklow: Glendalough State Forest (T128962), 1 in emergence trap on Sitka spruce stumps, 

vi.2006, Aoife Dillon. 

H40 Londonderry: Umbra NR (C724357), several emerging from Pinus contorta roots, 1.vi.2011, 

RA/MDB. 

110 

Figure 29: Asemum striatum © Roy Anderson 

Nathrius brevipennis (Mulsant, 1839) – Casual Introduction 

Larva develops particularly in thin woody stems, such as osier, dog-rose stems, wattle fences and 

wickerwork.


____________________________ 

H2 North Kerry: Killarney, vii.1907, EFB (Bullock, 1914; Speight, 1988).] 

Aromia moschata (Linnaeus, 1758) Musk Beetle – Rare 

Develops in willows and sallows, S. cinerea and S. alba, in wetlands and water meadows; larvae and 

pupae in dead wood of trunks and branches; has also been reported from alder, poplar and other 

broad-leaved trees; reputedly prefers young healthy growths rather than established trees and so - 

strictly - not a saproxylic; larvae can endure flooding for many weeks; adults fly in hot sunshine and 

visit blossom. “Local… headquarters… in the south-west, where it has been taken in some numbers in 

the Killarney and Glengariff districts” (Johnson & Halbert, 1902). “Common in parts of Kerry and 

West Cork, but very rare or absent from most districts in the north and east of Ireland” (Stelfox, 1959). 

Also turns up as an import. ‘Vulnerable’ in Speight (1988). 

H1 South Kerry: ‘Kenmare & Caragh Lake’, J.W. Yerbury (Champion, 1901); Kenmare, 2 on sallows, 

vii-viii.1906, LWBW (Bonaparte-Wyse, 1906). Ardea East, Tuosist (V790627), 9.viii.2011, F. Moore. 

H2 North Kerry: Killarney, E.P. Wright (Johnson & Halbert, 1902); Killarney, viii.1909, 1914, 1917, 

EFB; Cahirnane, viii.1930, EFB, NMI; Muckross, 4 on umbellifers, vii-viii.1906, LHBW (Bonaparte- 

Wyse, 1906); Muckross (V9685), on flowers of Polygonum, roadside, edge of oak woodland along lake 

shore, 19.ix.1986, MCDS (Speight, 1988). 

H3 West Cork: coll. R. Patterson; Glengarriff, Furlong coll. (Johnson & Halbert, 1902). 

H16 North-east Galway: Clonbrock (M7439), on a decaying willow, JNH (Halbert, 1898a), coll. R.E. 

Dillon (Johnson & Halbert, 1902) – it is unclear if this is a reliable report. 

H38 Down: near Belfast, AHH (Haliday, 1885) (Johnson & Halbert, 1902); Seaforde Estate (J4043), 

female reared from insect bored willow trunk material, coll. 11.viii.1988, emerged 2.ii.1989, RN (Nash, 

1989). 

Callidium violaceum (Linnaeus, 1758) Violet Longhorn – Established Introduction 

In dead birch, pine, oak, etc, but mainly conifers, larvae feeding between bark and sapwood; not on 

standing timber, but favours milled softwood. Ireland (Bullock, 1914). No modern records, presumed 

to have died out. 

H1 South Kerry: Lickeen Wood, vi.1907, F. Bouskell (Halbert, 1908); Lickeen Wood, Glencar (V78) 

(Kaufmann, 1948) [these almost certainly refer to the same record]. 

H2 North Kerry: near Killarney, EFB (Bullock, 1914); Killarney, vi.1915, EFB (Speight, 1988). 

Phymatodes testaceus (Linnaeus, 1758) – Casual Introduction? 

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Develops in and beneath the dying and dead bark of freshly fallen boughs and on the main trunk of 

living oak trees; also known from other tree species on occasion. Regularly transported outside its 

native range by the timber trade. Unknown in Ireland until the records given below. 

H37 Armagh: Tynan Abbey Estate (H760425), 3 specimens & numerous exit holes, under tight bark 

on the underside of a sundered oak branch, about 2 m above the ground, veteran parkland oak, 

1.vii.2011, MDB/RA (Bryan & Anderson, 2011); 1 specimen in a similar situation plus numerous exit 

holes at H761415, 26.vii.2011, MDB/RA. 

112 

Figure 30: Phymatodes testaceus © Roy Anderson 

Clytus arietis (Linnaeus, 1758) Wasp Beetle – Rare 

Develops in a variety of dead broad-leaved trees, and occasionally in conifers. Very rare in Ireland 

(Johnson & Halbert, 1902), where also occasionally imported with timber. O’Connor & Nash (1981) 

comment on the difficulty of discerning specimens derived from imports from any established native 

population. There is probably a native population in the woods of County Wicklow and possibly in 

South Kerry. ‘Endangered’ in Speight (1988). 

H1 South Kerry: “taken near Kenmare and another on the Glengariff Road, by Mr Hardy”, coll. JRH 

(Halbert, 1898c), but Edwin Bullock never found it further north at Killarney (O’Connor & Nash, 

1981).


____________________________ 

H20 Wicklow: Devil’s Glen, on flowers of mountain ash at head of glen, 14.vi.1935, AWS (annotation 

of a museum copy of Johnson & Halbert, 1902) (O’Connor & Nash, 1981); Deputy’s Pass (T2390), male 

on bramble leaf in pathside glade in ancient oak woodland, 17.vi.1993, KNAA (Alexander, 1997a). 

H21 Dublin: Iona Road, Drumcondra, one from a garden, vi.1914, NMI (O’Connor & Nash, 1981) 

H38 Down: Castlereagh, Belfast, one emerged from timber imported from Scandinavia, 18.iii.1980, 

T.Hedley (O’Connor & Nash, 1981) 

H39 Antrim: Cranmore, Belfast, one in coll. of Belfast Nat Hist & Phil Soc, taken by R. Templeton. 

H40 Londonderry: Magherfelt (H8990), emerged into house living space, from (?) furniture or roof 

timbers, 2.ii.1984, RA. 

Lamia textor (Linnaeus, 1758) The Weaver Beetle – Status unclear 

Associated with Salix, birch and aspen in damp woodland, the larvae developing in living healthy 

roots or boles of both young trees and moist decaying old boles; adults nocturnal, resting by day on 

trunks and roots; will nibble leaves of host plant. 

H2 Kerry North: (Kaufmann, 1948) – details not known. 

Pogonocherus hispidulus (Piller & Mitterpacher, 1783) – Rare 

Larvae under bark of recently dead branches or thin stems of broadleaves; especially hazel & apple. 

Ireland (1885, Haliday). ‘P. bidentatus’ in Johnson & Halbert (1902). May be confined to ancient 

woodlands. 

H2 North Kerry: Flesk, Killarney, vi.1957, EFB (Speight, 1988). 

H20 Wicklow: Glen of the Downs, one dead dug out of dead branch, vi.1888, JMB (Browne, 1892). 

Powerscourt, 20.vi.1937, ix.1938, EO (Speight, 1988). 

H25 Roscommon: St John’s Wood (M9956), FIT, 7.vi. & one on hazel leaf in sunny ride, 5.viii.2010, 

KNAA. 

H37 Armagh: Loughgall Manor, one found in grounds by Mrs Johnson (Johnson, 1892a). 

H38 Down: Rostrevor Oakwood (J1817), 1 in moth trap, 8.vi.1999, R. Thompson det. BN (Anderson, 

Simms & Nelson, 2000) 

H39 Antrim: Great Deer Park, Glenarm (D2911), one in FIT in old hazel stand, 6.vi.2006, KNAA. 

Pogonocherus hispidus (Linnaeus, 1758) – Scarce 

Develops in thin dead branches of a variety of broad-leaved trees; especially in old hedgerows. ‘P. 

dentatus’ rare (Johnson & Halbert, 1902). 

113


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H1 South Kerry: Caragh Lake, one swept off nettles at the Hotel end of the lake, vi.1902, 

(Donisthorpe, 1903; Bouskell, 1904); 23.vii.1928, EFB (Speight, 1988) 

H2 North Kerry: Muckross Demesne, vi.1924, OEJ (Janson & Bonaparte-Wyse, 1924); Ross, Killarney, 

26.ix.1931, EFB; Ardagh, Killarney, iii.1934, EFB; Killarney, v.1935, EFB; Flesk, Killarney, ix.1953, EFB 


H5 East Cork: Ballyannan Wood (W880719), Malaise trap, iii.iv.2004, TG. 

H6 Waterford: Curraghmore House, Portlaw, 1916, LHBW (Bonaparte-Wyse, 1916). 

H7 South Tipperary: Doon (R978062), Malaise trap, v.2004, TG. 

H9 Clare: Lough Oona (R404976), Malaise trap, v.vi.2007, TG. 

H12 Wexford: Slaney Bank near Killurin, JNH (Johnson & Halbert, 1902). 

H15 Galway Southeast: Kylebrack, coll. Quirke (Speight, 1988). 

H20 Wicklow: Clara, 23.xi.1933, AWS (Speight, 1988); Powerscourt, bred from birch, 1937, EO 

(O’Mahony, 1938). 

H21 Dublin: Dublin, JAP (Power, 1878; Johnson & Halbert, 1902). 

H29 Leitrim: Curraun (H103004), Malaise trap, v.vi.2004, TG. 

H30 Cavan: Farnham, on ivy, 3.iv.1943, RCF (Speight, 1988). 

H33 Fermanagh: Inishfendra, Crom (H3623), 28.v.1992, APF (Alexander, 1997a). 

H37 Armagh: Derrycarne Road, Portadown, beaten off apple tree in orchard, 8.ix.1998, BN. 

Leiopus nebulosus (Linnaeus, 1758) – Uncommon 

Larvae develop in freshly dead hazel branches, 2-6cm in diameter, with rather thin bark; galleries tight 

under the bark in the cambial layer; in branches with relatively low moisture; also reared from oak, 

lime and other broadleaved trees. Apparently widely distributed in Ireland, even if only seldom 

recorded (Speight, 1988). A very similar species, L. linnei, has recently been described and is known 

widely across Europe including southern Britain. 

H1 South Kerry: Lickeen Wood, Glencar, vii.1924, OEJ (Janson & Bonaparte-Wyse, 1924). 

H2 North Kerry: Killarney, ARH (Hogan, 1855) (Johnson & Halbert, 1902); Muckross Demesne, 

vi.1919, OEJ (Janson, 1920); Muckross Demesne, vi.1924, OEJ (Janson & Bonaparte-Wyse, 1924); 

Cahirnane, Killarney, iv.1938, EFB; Killarney, vi.1938, EFB; V9088, 16.viii.1978, M. de Courcy Williams 


H3 West Cork: Glengarriff Castle Demesne (V946 553), Malaise trap, vii.viii.2005, TG. 

114


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H6 Waterford: Lismore, JNH (Halbert, 1895a) (Johnson & Halbert, 1902). 

H9 Clare: Cratloe Wood, coll F. Neale (Johnson & Halbert, 1902). Cloonmore fen (R396747), 

Cragmoher Lough (R293867) & Lickaun Fen (R327823), Malaise traps, vi.vii.2006, TG. 


H14 Laois: Abbeyleix, JMB (Browne, 1901) (Johnson & Halbert, 1902). 

H16 Galway Northeast: Clonbrock (M7439), JNH (Halbert, 1896) (Johnson & Halbert, 1902). 

H18 Offaly: Clara Bog (N2429), birch/willow scrub, 4.vii.1984, MCDS; Charleville Wood (N3122), oak 

woods, 5.vii.1984, MCDS (Speight, 1988). 

H20 Wicklow: Powerscourt,3.vi.1927, EO; Glenmore, 16.vi.1939, EO; Knocksink Wood (O2117), oak 

woods, 2.viii.1978, MCDS; Lake park (O1602), swept bilberry under oak, 19.ix.1979, MCDS (Speight, 

1988). 

H21 Dublin: Rathfarnham (Hogan, 1854); Lucan Demesne, on oak, JNH (Halbert, 1900b) (Johnson & 

Halbert, 1902); Phoenix Park, 18.iv.1945, EO; Dublin, beaten from oak in suburban garden 16.vi.1987, 

MCDS (Speight, 1988). 

H23 Westmeath: Hare Island, Lough Ree (N0446), oak woods on an island, 19.vi.1976, MCDS 


H25 Roscommon: Mote Park, JNH (Halbert, 1898a). 

H33 Fermanagh: Crom Castle Park (H359246), 1 from aerial twigs of fallen oak branch, 6.vii.2007, 

KNAA. 

H36 Tyrone: Caledon Deer Park (H7543), 26.vii.2006, KNAA. 

H37 Armagh: Tandragee Castle, 7.v.1896, WFJ (Johnson, 1896); Tynan Abbey Estate, under bark on 

dead beech and beaten from Castanea, 1.vii.2011, RA. 

H38 Down: near Belfast, AHH (Haliday, 1885) (Johnson & Halbert, 1902). 

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Tetrops praeustus (Linnaeus, 1758) – Rare 

116 

Figure 31: Leiopus nebulosus © Roy Anderson 

Probably develops in dead branches; in old hedgerows, fruit trees, etc. ‘Endangered’ in Speight (1988). 

H13 Carlow: left bank of River Barrow above St. Mullins, one beaten from hawthorn, 16.v.1935, AWS 

(Stelfox, 1935). 

FAMILY ANTHRIBIDAE Fungus Weevils 

Choragus sheppardi Kirby, W., 1819 – Status unclear 

Larvae develop in decaying wood of old ivy, in hedges and woods. Only two old records, so unclear 

whether an established population. 

H21 Dublin: Templeogue, coll. G.P. Farran (Halbert, 1900b; Johnson & Halbert, 1902); Clontarf, EO 

(O’Mahony, 1928b).


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FAMILY CURCULIONIDAE Weevils 

Rhopalomesites tardyi (Curtis, 1825) Holly Weevil – Common & Widespread 

Develops in the dead heartwood of broad-leaved timber; bores in cambium and xylem, forming 

random galleries; flight holes 1.57-3.42mm in diameter. Ireland has most of the world population of 

this species; largely coastal and western in Britain; reported in Germany but status unclear. “Locally 

abundant in wooded districts throughout Ireland” (Johnson & Halbert, 1902). H1, H2, H3, H5, H12, 

H15, H16, H20, H21, H23, H27, H28, H31, H33, H34, H35, H36, H38, H39, H40. 

Pselactus spadix (Herbst, 1795) – Status unclear 

Figure 32: Rhopalomesites tardyi © Roy Anderson 

In rotten timber, coastal; old groins, driftwood, etc. Only one old Irish record. 

H21 Dublin: South Bull, in a piece of drift wood (Halbert, 1900b) (Johnson & Halbert, 1902). 

Pseudophloeophagus aeneopiceus (Boheman, 1845) – Scarce 

In damp rotten timber of various broad-leaved trees, in coastal woods and other coastal situations; 

also in driftwood; rare in buildings. Caulotrypis (Johnson & Halbert, 1902). 

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H1 South Kerry: Pointanskoh (V579984), in discarded pallet on roadside bank, 23.ix.2008, MC 

(Cawley, 2011b). 

H3 West Cork: near Rathmore (W0928), 3.iii.1983, MCDS (Morris, 1987); Sherkin Island, R.Moore 

(Morris, 1987) 

H4 or 5 Mid or East Cork: ‘found in profusion near Cork’ (Stephens, 1832). 

H21 Dublin: Portmarnock, common in old ash tree, CWB (Halbert, 1900b); Lambay Island, in elder at 

the Calico Hole, vi-x.1906, JNH (Halbert, 1907b); North Bull Island (O252834), 12 in large driftwood, 

dunes, 13.vii.2009, RA; Sutton (O252397), under a piece of wood at HWM in a small saltmarsh, 

19.vii.2011, MC. 

H28 Sligo: Ballincar (G678382), under processed wood at estuarine HWM, 13.v.2011, MC; Pollnagat 

(G383387), under a piece of wood, exposed grassy cliff top behind shingle beach, 9.vi.2011, MC. 

H38 Down: Killard NNR (J6043), in driftwood, 24.iv.2007, MGT. 

Euophryum confine (Broun, 1881) – Established introduction 

Immigrant New Zealand species, first recorded in Ireland by Hickin (1952), and the first outdoor 

records in 1955 (McClenaghan, 1987). Always found associated with timber where damp and decay 

evident; dense channelled galleries in decayed heartwood. Regularly found in cuboidal red rot of 

fungus Laetiporus sulphureus outdoors, but also in wet rot on timbers in buildings. Anderson (1984) 

summarised Irish records, while others were added by McClenaghan (1987). H1, H3, H4, H5, H6, H8, 

H18, H21, H23, H25, H28, H31, H33, H35, H36, H37, H38, H39, H40. 

Euophryum rufum (Broun, 1880) – Casual Introduction 

Possibly not a distinct species; added to the Irish list on a specimen from Exchequer Street, Dublin, 

1976 (O’Connor, 1978) and a second from Flesk, Killarney, in the Bullock collection, misdet. as 

Pentarthum huttoni (O’Connor, 1978). A secondary pest of timber in buildings. Dublin City (O’Connor 

1981, 1984). 

Pentarthrum huttoni Wollaston, 1854 – Established introduction 

Larvae and adults bore into timber which is damp and colonised by fungus; more often in 

floorboards, etc; hard- and softwoods, but softer layers of wood eaten away first, leaving harder rings 

untouched; requires sufficiently damp climate. Possibly a long-established non-native, as all old 

records are from wooden casks at ports. Irish records summarised by O’Mahony (1950). Ireland (AF 

O’Farrell & PM Butler, 1948). 

[H2 North Kerry: Killarney, Bullock, in lit. (O’Mahony, 1929a) – NB specimen of Euophryum rufum 

misdet as Pentarthum in Bullock collection (O’Connor, 1978)] 

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H20 Wicklow: Newtownmountkennedy, J. Forwood (Morris, 1993a). 

H21 Dublin: Clontarf, O’Mahony viii.1925, EO (O’Mahony, 1927) (O’Connor, 1981). 

H27 West Mayo: Glas Valley, Ballycurran, ix-x.1984, G.F.B. Pullar (Morris, 1997); infesting hospital 

with dry rot, 20.xii.1985 (O’Connor et al., 1990). 

H39 Antrim: Belfast Docks, 1942-1946, AFO (O’Farrell & Butler, 1948); Larne (D4002), in hardwood 

panelling in house, 24.xi.1985, RA. 

Macrorhyncolus littoralis (Broun, 1880) – Casual 

In driftwood. Possibly native to Australia, but well-established in New Zealand, South Africa and the 

Americas. The only other Palaearctic records appear to be two from Kent, England, in 1987 and 1988. 

H12 Wexford: Ballyteige Burrow (S938056), 40 in 2 colonies under driftwood on beach, 27.vi.2006, 

MGT (Telfer, 2007). 

Cryptorhynchus lapathi (Linnaeus, 1758) – Scarce 

Burrows in dead trunks of alder, poplar, willow and birch, sometimes even live trees. “Taken on one 

occasion in some numbers on bark of sallows” (Johnson & Halbert, 1902). 

H2 North Kerry: near Killarney, EFB (Bullock, 1914); Cloghereen at Muckross Lake, 3.vi.1919 – 

18.vi.1919, OEJ (Janson, 1920). 

H6 Waterford: Dunhill (S5304), 29.vi.1988, JPOC (Morris, 1997). 

H12 Wexford: on the R. Boura, Furlong ms. (Johnson & Halbert, 1902) 

H20 Wicklow: Boulnalea, Furlong ms; Murragh, HKGC (Johnson & Halbert, 1902); Arklow, vii.1926, a 

number in cop. on trunk of Salix, JNH (Halbert, 1937). 

H28 Sligo: Gleniff picnic site, 18.vii.2008, RA. 

H29 Leitrim: Clooncoe Lough (N1091), 29.v.1985, MCDS (Morris, 1987). 

H37 Armagh: Coney Island, Lough Neagh, WFJ (Johnson, 1895; Johnson & Halbert, 1902); Brackagh 

Moss NNR, 16.vi.1999, R. Key. 

H39 Antrim: near Gawley’s Gate, Lough Neagh, two beaten from osiers, ix.1935, WMC (Crawford & 

O’Mahony, 1935); Montiaghs Moss ASSI, 16.vi.1999, A.P. Fowles. 

Acalles misellus Boheman, 1844 syn turbatus nec Boheman, 1844 – Uncommon 

Larvae probably develop in dead twigs and small branches; found in woods, hedgerows and 

occasionally isolated field trees. More oceanic than ptinoides; may be specially associated with ivy. 

M.G. Morris (pers. comm.) considers it to be overlooked because of its cryptic habits and that it is 

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actually quite common in Ireland (Morris, 1997; Alexander, 1997c). VCs 1, 2, 19, 21, 34, 39 (Morris, 

1993a). 

H2 North Kerry: near Killarney, EFB (Bullock, 1914); near the base of Torc Mountain (V98), in plenty 

by beating holly and ivy, vi.1919, OEJ (Janson, 1920); Muckross Demesne, vi.1924, OEJ (Janson & 

Bonaparte-Wyse, 1924). 

H3 West Cork: near Inchigeelach (W2467), 12.vii.1996, MGM (Morris, 1997); Courtmacsherry Wood 

(W524428), 6.ii.2008 & Poulgorm Bridge (W210353), 27.ii.2008, MC (Cawley, 2009c) 

H4 Mid Cork: Shippool Woods, Bandon River (W5757), 23.v.1994, KNAA (Alexander, 1997c). 

H8 Limerick: Poultallin Point (R239520), 24.viii.2008, MC (Cawley, 2009c). 

H9 Clare: Ballyvaughan Harbour (M2307), 6.vi.1995; Ballyeighter Loughs (R3595), 6.vi.1995, MGM 

(Morris, 1997). 

H10 North Tipperary: near Annagh Castle (R8290), 9.vii.1996, MGM (Morris, 1997). 

H14 Laois: Grantstown Wood (S3380), 29.v.2007, MC (Cawley, 2009c). 

H15 South-east Galway: Garryland Wood, 21.v.1994, RA. 

H18 Offaly: Cedar Hill, near Shinrone (S1092), 1.vi.1995, MGM (Morris, 1997). 

H19 Kildare: Leixlip Spa (N9936), vii.1991, MGM (Morris, 1992). 

H20 Wicklow: Grange near Newcastle (O300025), 31.v.1995, MGM (Morris, 1997). 

H21 Dublin: Dundrum (Halbert, 1900b) (Johnson & Halbert, 1902); St Annes Park, 1924, EO 

(O’Mahony, 1924); Clontarf, EO (O’Mahony, 1928b); Portmarnock Strand, EO (O’Mahony, 1928b); 

Sutton (O253396), under tiles lying on ivy, 19.vii.2011, MC. 

H22 Meath: Hamwood (N987409), 20.x.2008, MC (Cawley, 2009c). 

H23 Westmeath: Cappankelly (N058446), 22.iii.2012, MC. 

H28 Sligo: Lugnadetra (G6228), 4.vi.1995, Morris (1997). 

H29 Leitrim: Milltown Wood (G874404), 16.iv.2009, MC (Cawley, 2009c). 

H33 Fermanagh: Inisherk, Crom (H353244), in tray below FIT, 8.viii. – 4.ix.1992, APF (Alexander, 

1997c); Crom Castle Park (H359246), 6.vii.2007, KNAA. 

H34/40: Foyle district, CWB (Buckle, 1900) (Johnson & Halbert, 1902) 

H38 Down: Hillsborough Forest, pitfall, 28.v.1993, RA (Alexander, 1997c); Belvoir Forest, 27.viii.1993, 

RA; Donard Demesne, 31.xii.2011, RA. 

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H39 Antrim: Ballycastle, BT (Johnson & Halbert, 1902); Craigagh Wood (D227323), one under bark on 

fallen beech branch, 17.vi.1992, KNAA (Alexander, 1997c). 

Acalles ptinoides (Marsham, 1802) – Rare old growth species 

Develops in dead twigs on trees; litter layer of lowland oak woods; ancient woodland and old 

heathland; a reliable marker of old growth (Crowson 1987). Only three modern localities known. 

H1 South Kerry: Glencar, July 1924, OEJ (Janson & Bonaparte-Wyse, 1924). 

H2 North Kerry: Muckross Woods, 1923, OEJ (Janson & Bonaparte-Wyse). 

H3 West Cork: Glandore, JNH (Johnson & Halbert, 1902). 

H12 Wexford: Forth Hills, JNH (Johnson & Halbert, 1902); Curracloe, vi.1936, JNH (Halbert, 1937); 

Kelly’s Wood, Camlin Hill (S711255), 12.vi.2008, MC (Cawley, 2009c). 


H20 Wicklow: Bray Head, JNH (Halbert, 1895b) (Johnson & Halbert, 1902). 

H21 Dublin: Dundrum, JNH (Halbert, 1900b; Johnson & Halbert, 1902). 

H23 Westmeath: Crookedwood (N464628), 16.v.2008, MC (Cawley, 2009c). 

H25 Roscommon: St John’s Wood (M9956 & N0056), leaf litter, 10.v & FIT, 6.viii.2010, KNAA. 

H31 Louth: Carlingford, WFJ (Johnson & Halbert, 1902). 

H34 Donegal: Foyle district (Buckle, 1900) (Johnson & Halbert, 1902). 

Kyklioacalles roboris (Curtis, 1835) – Rare old growth species 

Develop in small twigs of trees; confined to ancient woodland. Only one modern record. 

H1 South Kerry: Rossbehy (V6490), S. Stevens (Stevens, 1871) (Johnson & Halbert, 1902). 

H13 Carlow: Kilcarry, 1.v.1929, specimens collected by AWS (Morris, 1987; 1993a). 

H20 Wicklow: Enniskerry Glen, 1931, AWS/EO (O’Mahony, 1932). 

H25 Roscommon: St John’s Wood (M9956), off ivy, 26.vii.2010, & FIT, 24.viii.2010, KNAA. 

Magdalis armigera (Fourcroy, 1785) – Rare 

Female drills hole in dead elm branch and deposits egg inside; larvae feed on cambium; in hedgerows 

& scrubby places. 

H2 North Kerry: Killarney, three in Bullock collection, vii.1918, EFB (Morris, 1993a). 

H3 West Cork: Farrandeligeen, Castlehaven (W1730), beaten from roadside hedge with dead elm, 

7.vi.1996, KNAA (Alexander, 1997c); 

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H6 Waterford: Dromana Forest, near Cappoquin, JNH (Johnson & Halbert, 1902). 

H18 Offaly: Cedar Hill, near Shinrone (S1092), 1.vi.1995, MGM (Morris, 1997). 

H21 Dublin: Santry, on elms, JNH (Halbert, 1894, 1922) (Johnson & Halbert, 1902). Phoenix Park 

(O1235), 11.vi.1983, JPOC (Morris, 1997). 

H33 Fermanagh: Castle Coole Park (H251429), abundant on re-growth from base of dead elm, 

2.vi.1992, KNAA (Alexander, 1997c). 

Magdalis carbonaria (Linnaeus, 1758) – Rare 

Associated with birch, the larvae developing in dead twigs and branches. Considered an ancient 

woodland relict species in Britain and this may be true in Ireland as well – all known sites have high 

quality fauna. 

H20 Wicklow: Powerscourt Deerpark, two swept from bracken beneath birch, 9.vi.1919, JNH (Halbert, 

1922); Morris (1993a). Glendalough Wood (T1296), one swept from open woodland, 13.vi.1993, KNAA 

(Alexander, 1997e). 

H25 Roscommon: St John’s Wood (M9956), 9.vi.2010, KNAA. 

H39 Antrim: Glenarm Nature Reserve (D304110), 2 swept off sprouting hazel, RA. 

H40 Londonderry: Banagher Glen (C672045), 8.vii.2007, KNAA. 

122 

Figure 33: Magdalis carbonaria © Roy Anderson


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Magdalis ruficornis (Linnaeus, 1758) – Status unclear 

Removed from the Irish list by Morris (1993b). Unlocalised material in Haliday collection. 

Hylobius abietis (Linnaeus, 1758) – Established introduction 

Associated with Scots pine. “Locally common about fir plantations” (Johnson & Halbert, 1902). A 

common conifer pest (Morris, 1993a, 1997). H1, H3, H6, H7, H9, H10, H12, H13, H14, H20, H21, H25, 

H28, H31, H33, H34, H36, H37, H38, H39, H40. 

Pissodes castaneus (De Geer, 1775) – Recent Introduction 

A native of continental Europe, this feeds on the inner bark and cambium of pine species, and 

although all ages may be attacked, there is a bias towards younger trees so plantation pine may be 

particularly affected. 

H29 Leitrim: Gleniff, Annacoona (G7246), on lodgepole pine, 11.vi.1997, SC (Clawson & Anderson, 

2006). 

Pissodes pini (Linnaeus, 1758) – Recent Introduction 

Feeds on the inner bark and cambium of weakened pine trees in plantations. 

H33 Fermanagh: Ely Lodge Forest (H172522), larvae and adults at base of Scots pine, 22.iv.1997, SC 

(Clawson & Anderson, 2006). 

Figure 34: Pissodes pini © Roy Anderson 

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CURCULIONIDAE Subfamily SCOLYTINAE Bark Beetles 

Species feeding on wood (xylem) and/or phloem are usually restricted to one or a few hosts, whereas 

those which carry their own symbiotic fungi which break down the xylem (ambrosia beetles) may 

colonize a larger range of hosts. Many species have been imported in timber and some have become 

established. A number are more strictly phytophagous, their larvae feeding in the still living inner 

bark of stressed or moribund stems or branches, but these have been included in the list as they 

remain present in the dead tissues. 

Scolytus multistriatus (Marsham, 1802) Small Elm Bark Beetle – Recent Arrival 

In smaller, thinner barked, dying branches of various broad-leaved trees but especially elm. Only 

discovered in Ireland as recently as 1980. 

H8 Limerick: Adare, possible galleries, winter 1979 (O’Callaghan, 1982). 

H21 Dublin: Phoenix Park, 48 attracted to pheromone traps, summer 1980, O’Callaghan (1982). 

Scolytus rugulosus (Müller, P.W.J., 1818) Small Fruit Tree Bark Beetle – Rare 

Mainly on apple trees in orchards; probably overlooked, although possibly a new arrival. 

H33 Fermanagh: Inisherk, Crom (H357241), beaten from an old apple tree in an abandoned orchard, 

24.v.2009, APF. 

Scolytus scolytus (Fabricius, 1775) Common Elm Bark Beetle – Widespread 

Mainly in elm, also other broad-leaved trees; adults feed on bark of top living twigs after emerging 

from thicker dead bark where the larvae develop. First found in Ireland in 1943 (Quirke, 1943) 

although probably long established and widespread. 

H2 North Kerry: Bellview Wood, Kenmare Estate, Killarney, larvae, pupae and adults from an old 

elm, 1943, Quirke (1943); Flesk, Killarney, 1948, 1949, 1952, 1955, EFB (O’Connor et al., 1991). 

H3 West Cork: Lough Hyne House Grounds (W0928), 2.vi.1996; Castlehaven Castle Bay (W175300), 

7.vi.1996; KNAA. 

H4 Mid Cork: near Riverstick (W6759), viii.1980, JAG (O’Connor et al., 1991). 

H6 Waterford: wood near Cappoquin, undated, JNH (O’Meara, 2008); near Dunhill (S5305), 4.vii.1989, 

JPOC (O’Connor et al., 1991); Ballyeelinan (X2181), 10.viii.1990, KNAA. 

H8 Limerick: Curraghchase (R4149), 21.vi.1979, J. Mullin (O’Connor et al., 1991). 

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H13 Carlow: Carlow City, adults inside house with elm firewood, 8.v.1987, D. Phillips (O’Connor et 

al., 1991). 

H19 Kildare: Donadea (N8332), 8.iii.1987; Milltown (N7617), dead beetles in dead elm on canal bank, 

30.iv.1987, JPOC (O’Connor et al., 1991); Leixlip Spa (N9936), vii.1991, MGM (Morris, 1992). 

H20 Dublin: Slade of Saggart (O0324), in riverbank elm tree, 18.vii.1982, J.M.O’Connor (O’Connor et 

al., 1991). 

H28 Sligo: Markree Castle Park (G6925), 30.v.1992, KNAA. 

H33 Fermanagh: Crom Castle Park (H32), 1.vi.1992; Castle Coole Park (H2643), 2.vi.1992 & 30.iii.2006, 

KNAA. 

H38 Down: Mount Stewart Demesne (J5570), 18.viii.1992; Rowallane Estate (J409575), 22.ix.2004; 

Castle Ward Park (J5649), 2.iv.2006, KNAA. 

H39 Antrim: Collin Glen (J270720), 19.v.1992; Great Deer Park, Glenarm (D2911), galleries widely, 

2006, KNAA. 

H40 Londonderry: Springhill (H8683), 25.vi.1992, KNAA. 

Pityophthorus pubescens (Marsham, 1802) – Established introduction 

Pine and spruce; in small dead stems of less than 1cm diameter. Known from Galway, Wicklow, 

Wexford, Clare and Cork (Johnson & Halbert, 1902); also Cavan, Dublin, Kerry, Meath (O’Connor et 

al. 1991). 

Cryphalus asperatus (Gyllenhal, 1813) – Established Introduction 

Develops in dead branches of spruce but not a pest species. Known from Kerry and Wicklow 

(O’Connor et al., 1991). 

Dryocoetes autographus (Ratzeburg, 1837) – Established introduction 

In dead and dying conifers. Until recently, reported in Ireland only as imports (O’Mahony, 1935b; 

O’Connor et al., 1991). 

H29 Leitrim: Truskmore (G7446), 1 in pitfall on summit, 11.v-16.vi.2004, SM det. RA. 

H36 Tyrone; Ballyard, Trillick (H3458), on silver fir, 1.vii.1997, SC; Pomeroy Forest (H7072), on silver 

fir, 2.vi.1998, SC. 

H37 Armagh: Camlough Wood (J0524), under bark at base of Picea abies, 7.vi.2004, SC; Slieve Gullion 

Forest (J038196), bred from Picea sitchensis bark, MDB det. RA; Ballymoyer Wood (H974311), reared 

from Tsuga heterophylla wood, 2.ii.2012, MDB det. RA. 

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H38 Down: Hillsborough Forest (J2558), adults emerging from bark on Picea abies thinnings, 

11.iv.1995, RA. 

H39 Antrim: Farr’s Bay NNR (J0887), in Picea logs, 4.vii.1986, RCW (Welch, 1986); Randalstown Forest 

(J0888), on Sitka spruce, 8.ix.1993, SC. 

Dryocoetes villosus (Fabricius, 1793) – Uncommon 

Develops in relatively thick bark of freshly dead oak boughs and trunks; also known from sweet 

chestnut, beech and lime. 

H2 North Kerry: Killarney, including Cahirnane, Deerpark and Flesk, 1914-1951, EFB (O’Connor et 

al., 1991); Muckross Park (V5458), 4.ix.1998, KNAA. 

H3 West Cork: Glengarriff Woods (V918573), developing in felled old oaks, 13.vi.1996 (Alexander, 

1997d); Big Meadow (V918570), 6.xi.2004, KNAA. 

H14 Laois: Abbeyleix (S4284), Old Town Oak, 16.vi.1993, KNAA (Alexander, 1997d); 

H18 Offaly: Charleville Castle (N3123), 16.vi.1993 KNAA (Alexander, 1997d); 

H33 Fermanagh: Reilly Wood, Crom (H3624), in thick bark of live open-grown oak outside wood 

edge, 28.v.1992 (Alexander, 1997d); Inisherk Island (H353244), FIT, 1.vi.–7.vii.1992, APF (Alexander, 

1997d) & FIT, 6.vii.2007, KNAA; Castle Coole Park (H2643), 19.v & 17.vi.2006, KNAA. 

H34 East Donegal: Foyle district, CWB (O’Connor et al., 1991). 

H36 Tyrone: Caledon Deer Park (H7543), various dates, 2006, KNAA. 

H37 Armagh: Slieve Gullion Forest (J038193), on oak, 8.iii.2012, RA; 

H38 Down: Rostrevor Oakwood (J1817), 28.vi.1992 & 11.vii.2007, KNAA (Alexander, 1997d); Castle 

Ward Park (J5649), 23.viii.1998, KNAA; Tullyframe, Mourne Park (J265168), on oak bark, 27.iii.2012, 

MDB/RA. 

H39 Antrim: Great Deer Park, Glenarm (D2911), various dates, 2006, KNAA. 

H40 Londonderry: Walworth Wood, July, CWB (Johnson & Halbert, 1902). 

Orthotomicus laricis (Fabricius, 1792) – Established introduction 

Develops under bark of dead conifers, chiefly pine. First noted by O’Mahony (1934). 

H2 North Kerry: Flesk, Killarney, EFB (O’Connor et al., 1991). 

H13 Laois: Emo Park (N5406), in pine logs, iv.1934, EO (O’Mahony, 1934). 

H21 Dublin: Glenasmole, in poles used as a railing, EO (O’Mahony, 1940). 

H33 Fermanagh: Rosslea Wood (H5431), on Pinus sylvestris, 15.iii.1993, SC. 

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Pityogenes bidentatus (Herbst, 1783) – Established introduction 

Pine and spruce; in small dead thin-barked branches. Added to the Irish list by Johnson (1892a) and 

now known from Antrim, Armagh, Dublin, East Cork, North Kerry, Wicklow (O’Connor et al., 1991). 

H2 North Kerry: Cahirnane, Flesk, EFB (O’Connor et al., 1991). 

H5 East Cork: Ballyhoura (O’Connor et al., 1991). 

H20 Wicklow: Kilruddery (O’Connor et al., 1991). 

H21 Dublin: St Annes Park, in pines, 1924, EO (O’Mahony, 1924). 

H37 Armagh: Armagh, in a garden, WFJ (Johnson, 1892a). 

H38 Down: Belvoir Forest (J3469), galleries under bark of dismasted larch, 16.v.1996, RA. 

H39 Antrim: Shane’s Castle, vi.1902, JNH (Halbert, 1910b); vi.1923, OEJ (Janson, 1924). 

Trypodendron domesticum (Linnaeus, 1758) – Scarce old growth species 

An ambrosia beetle, developing in the sapwood of a wide range of freshly dead broad-leaved timber 

including oak, maple, alder, beech, birch and ash. Adults excavate deep galleries in the sappy timber 

and feed on the fruiting bodies of fungi cultivated therein. 

H2 North Kerry: near Killarney, EFB (Bullock, 1914); Muckross Demesne, in decaying wood, vi.1919, 

OEJ (Janson, 1920) 

H4 Mid Cork: Blarney, EFB (Bullock, 1914). 

H5 East Cork: Ballyannan Wood (W880719), Malaise trap, iii.iv.2004, TG. 

H6 Waterford: Lismore, under bark on beech log, 13.ii.2011, RA. 

H20 Wicklow: Avoca, 15.iv.1927, AWS; Powerscourt Deerpark, in decayed alder trunk, 9.vi.1919, JNH 


H25 Roscommon: Cloonca Wood, JNH (Halbert, 1898a; Johnson & Halbert, 1902). St John’s Wood 

(M9956), 13.iv.2010, KNAA. 

H27 West Mayo: Westport Demesne, JNH (Johnson & Halbert, 1912). 

H33 Fermanagh: Castle Coole Park (H260430), fragments in decaying oak branch, 2.vi.1992, KNAA 

(Alexander, 1997d); Inisherk, Crom (H353244), two in FIT, 1.vi. – 7.vii.1992 & another 5.viii. – 4.ix.1992, 

APF (Alexander, 1997d); Correl Glen (H074545), FIT, 5.vii.2007, KNAA. 

H35 West Donegal: Glentogher, “borings of Trypodendron domesticum in a fallen beech tree” CWB 

(Buckle, 1900; Johnson & Halbert, 1902). 

H38 Down: Morelands Meadow, under bark, beech logs, 18.xi.1992, RA. 

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H39 Antrim: Great Deer Park, Glenarm, FITs, 20.v.2006, KNAA; Carr’s Glen, under bark on hazel, 

23.iii.2010, RA; Rea’s Wood, under burnt bark of sycamore, RA; Barnett’;s Park, Belfast, under bark on 

oak branch, 25.iii.1982, RA. 

H40 Londonderry: Drenagh Demesne (C6923), 14.vi.2006, KNAA. 

Trypodendron lineatum (Olivier, 1795) Conifer Ambrosia Beetle – Established Introduction 

Dead wood of pine, spruce, larch, and fir. 

H30 Cavan: Cloverhill, specimen in pine bark, v.1914, GWN (Nicholson, 1915). 

H37 Armagh: Slieve Gullion Forest (J040196), hibernating under ash bark (!), 8.iii.2012, RA. 

H39Antrim: Farr’s Bay NNR (J089870), attacking Picea stumps, 4.vii.1986, RCW (Welch, 1986). 

Hylesinus crenatus (Fabricius, 1787) Large Ash Bark Beetle – Scarce 

Chiefly in large old dying ash, but also known in other countries from oak, walnut, etc, in rather thick 

bark of the trunk. No modern records. 

H2 North Kerry: Muckross, 1940, 1946, 1948, EFB (O’Connor et al., 1991). 

H7 South Tipperary: Tipperary town, A.C. Forbes (Forbes, 1910; O’Mahony 1929). 

H10 North Tipperary: Templemore, 25.v.1927, R.A. Phillips (O’Connor et al., 1991). 

H21 Dublin: Tolka Valley, 1912, JNH (O’Mahony, 1929a). 

H22 Meath: Athboy, A.C. Forbes (O’Mahony, 1929a). 

H33 Fermanagh: Old Crom Castle (H364239), swept near lakeshore, 24.v.2012, K.A. Bryan. 

Hylesinus varius (Fabricius, 1775) Common Ash Bark Beetle – Widespread 

In ash; in standing and fallen recently dead trunks and boughs; makes short blind hibernation 

galleries in crotches of live ash trees, where it is associated with bacterial disease ash rose canker. 

Surprisingly poorly known in Ireland, although probably overlooked. 

H2 North Kerry: Ardagh, Killarney, 9.v.1931, EFB; Flesk, Killarney, iv.1953, v.1954, EFB (O’Connor et 

al., 1991). 

H5 East Cork: Glanmire, JNH (Johnson & Halbert, 1902); Cork city (as Hylesinus fraxini) in O’Mahony 

(1924). 

H17 Galway: Muckanagh (M7951), 11.iv.2010, KNAA. 

H19 Kildare: Landenstown, 13.ix.1942, AWS (O’Connor et al., 1991). 

H20 Wicklow: Devil’s Glen, 12.ix.1926, AWS (O’Connor et al., 1991). 

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H21 Dublin: Lucan Demesne, Hylesinus fraxini, JNH (Halbert, 1893); Dundrum, GHC (Carpenter, 

1894) (Johnson & Halbert, 1902); Raheny, Clontarf (as Hylesinus fraxini) in O’Mahony (1924). 

H25 Roscommon: Mount Talbot (M8155), 16.iv.2010, KNAA. 

H29 Leitrim: Ballinamore, in firewood, ix.1987, D. Quinn (O’Connor et al., 1991). 

H33 Fermanagh: Crom Castle Park (H32), 26.v.1992, KNAA. 

H36 Tyrone: Caledon Deer Park (H7543), 17.v.2006, KNAA. 

H37 Armagh: Jerrettspass, 13.xii.1912, WFJ (Johnson, 1912); Loughgall Country Park, 15.iv.2008, RA. 

H38 Down: Belfast, 28.x.1986, Forest Service Collection (O’Connor et al., 1991); Castle Ward Park 

(J5649), 21.v.2006, KNAA. 

H39 Antrim: Moss Plantation, Shane’s Castle, 16.vii.2010, RA. 

Pteleobius vittatus (Fabricius, 1787) – Status unclear 

In recently dead thin-barked elm, and occasionally reported from ash and lime. Johnson and Halbert 

(1902) cite Ireland for Hylesinus vittatus (Haliday coll.) but O’Connor et al. (1991) were unable to 

confirm this as no material appears to have survived. Subfossil remains have been found in Viking 

Dublin (Reilly 2003) 

Hylastes ater (Paykull, 1800) – Established Introduction 

Develops in dying pine. All old records of this species are believed to be H. brunneus Erichson 


H2 North Kerry: Kenmare Demesne, plenty dug out of the stumps of recently felled pines, between 

June 25 & 30, 1919, OEJ (Janson, 1920). 

H20 Wicklow: Camcor River Bridge (N2104), 16.vi.1993, KNAA; Three Lakes Table Mountain 

(T034983), pitfall, 5.vi-28.viii.1998, Myles Nolan det. RA. 

Hylastes brunneus Erichson, 1836 – Established Introduction 

Develops in dying pine, widely across Ireland and long-established. All old Irish records for H. ater 

have proved to be this species (O’Connor et al., 1991). A serious pest of pine afforestation. H2, H3, H5, 

H6, H7, H17, H20, H21, H30, H34, H36, H37, H38, H40. 

Hylastes opacus Erichson, 1836 – Established introduction 

Develops in dead pine, but also reported from elm and ash. A serious pest of pine afforestation. 

H2 North Kerry: Muckross, EFB ( Bullock, 1914); Kenmare Demesne, plenty dug out of the stumps of 

recently felled pines, 25.vi-30.vi.1919, OEJ (Janson, 1920). 

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H5 East Cork: Ballyhoura, 25.vi.1975, Forestry Commission Collection (O’Connor et al., 1991). 

H7 South Tipperary: Killballyboy Wood (S0113), roadside, 11.v.1994, JAO (Owen, 1997). 

H12 Wexford: The Raven (T1126), swept in conifer plantation, 4.vi.1986, JPOC (O’Connor et al., 1991). 

H14 Laois: Emo Park, Portarlington, iv.1934, EO (O’Connor et al., 1991). 

H20 Wicklow: Devil’s Glen, 16.vi.1934, EO (O’Connor et al., 1991). 

H31 Louth: south, HKGC (Cuthbert, 1892b). 

Hylurgops palliatus (Gyllenhal, 1813) – Established introduction 

Develops under bark of dead conifer timber; common and widespread in Ireland (O’Connor et al., 

1991). H1, H2, H5, H14, H17, H18, H20, H21, H29,H30, H33,H34, H36, H37, H38, H39, H40. 

Tomicus piniperda (Linnaeus, 1758) Pine Shoot Beetle – Established introduction 

Dead conifers, especially Pinus; tunnels under bark longitudinally; two broods annually. Widespread 

in pine plantations in Ireland (O’Connor et al., 1991). H2, H5, H10, H12, H17, H18, H20, H21, H22, 

H36, H37, H38, H39, H40. 

130


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Appendix 1 Recorders and list of abbreviations 

The following are the abbreviations used for recorders in the species accounts 

AFO A.F. O’Farrell JAP J.A. Power 

AGI A.G. Irwin JJW J.J. Walker 

AHH A.H. Haliday JMB J.M. Browne 

APF A.P. Foster JNH J.N. Halbert 

ARF A.R. Furlong JPOC J.P. O’Connor 

ARH A.R. Hogan JRH J.R. Hardy 

AWS A.W. Stelfox KNAA K.N.A. Alexander 

BN B. Nelson LHBW L.H. Bonaparte-Wyse 

BT B. Tomlin MAOC M.A. O’Connor 

CAMR C.A.M. Reid MC M. Cawley 

CJ C. Johnson MCDS M.C.D. Speight 

CTG C.T. Gimingham MDB M.D. Bryan 

CWB C.W. Buckle MGM M.G. Morris 

DAL D.A. Lott MGT M.G. Telfer 

EFB E.F. Bullock OEJ O.E. Janson 

EO E. O’Mahony PJH P.J. Hodge 

GCC G.C. Champion RA R. Anderson 

GHC G.H. Carpenter RCF R.C. Faris 

GNF G.N. Foster RCW R.C. Welch 

GMO G.M. Orledge RM R. Martin 

GWC G.W. Chaster RN R. Nash 

GWN G.W. Nicholson SC S. Clawson 

HKGC H.K. Gore Cuthbert SM S. McCormack 

HSJD H.St.J. Donisthorpe TG T. Gittings 

IM I. McClenaghan TVW T.V. Wollaston 

JAG J.A. Good WES W.E. Sharp 

JAO J.A. Owen 

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Other abbreviations used in the text are as follows 

BMNH British Museum (Natural History) NMI National Museum of Ireland 

FIT flight interception trap UM Ulster Museum 

145


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Appendix II Taxonomic checklist of species 

HISTERIDAE Hister Beetles 

146 

Abraeus perpusillus (Marsham, 1812) 

PTILIIDAE Feather-winged Beetles 

LEIODIDAE 

Nossidium pilosellum (Marsham, 1802) 

Ptenidium laevigatum Erichson, 1845 

Ptinella aptera (Guérin-Méneville, 1839) 

Ptinella cavelli (Broun, 1893) 

Ptinella denticollis (Fairmaire, 1858) 

Ptinella errabunda Johnson, 1975 

Ptinella limbata (Heer, 1841) 

Ptinella taylorae Johnson, 1977 

Pteryx suturalis (Heer, 1841) 

Anisotoma humeralis (Fabricius, 1792) 

Anisotoma orbicularis (Herbst, 1792) 

Amphicyllis globus (Fabricius, 1792) 

Agathidium confusum Brisout, 1863 

Agathidium nigripenne (Fabricius, 1792) 

Agathidium rotundatum (Gyllenhal, 1827) 

Agathidium seminulum (Linnaeus, 1758) 

Agathidium varians Beck, 1817 

SCYDMAENIDAE Stone Beetles 

Stenichnus bicolor (Denny, 1825) 

STAPHYLINIDAE Rove Beetles 

Subfamily OMALIINAE 

Phyllodrepoidea crenata (Ganglbauer, 1895) 

Coryphium angusticolle Stephens, 1834


____________________________ 

Acrulia inflata (Gyllenhal, 1813) 

Dropephylla devillei Bernhauer, 1902 

Dropephylla gracilicornis (Fairmaire & Laboulbène, 1856) 

Dropephylla ioptera (Stephens, 1832) 

Dropephylla koltzei Jászay & Hlavac, 2006 

Hapalaraea pygmaea (Paykull, 1800) 

Phloeonomus punctipennis Thomson, C.G., 1867 

Phloeonomus pusillus (Gravenhorst, 1806) 

Phloeostiba plana (Paykull, 1792) 

Xylostiba monilicornis (Gyllenhal, 1810) 

Subfamily PSELAPHINAE Short-winged Mould Beetles 

Euplectus bescidicus Reitter, 1881 

Euplectus infirmus Raffray, 1910 

Euplectus karstenii (Reichenbach, 1816) 

Euplectus mutator Fauvel, 1895 

Euplectus piceus Motschulsky, 1835 

Euplectus punctatus Mulsant, 1861 

Bibloporus bicolor (Denny, 1825) 

Subfamily: PHLOEOCHARINAE 

Phloeocharis subtilissima Mannerheim, 1830 

Subfamily: TACHYPORINAE 

Sepedophilus littoreus (Linnaeus, 1758) 

Subfamily: ALEOCHARINAE 

Alaobia pallidicornis (Thomson, C.G., 1856) 

Atheta basicornis (Mulsant & Rey, 1852) 

Atheta pilicornis (Thomson, 1852) 

Bessobia fungivora (Thomson, 1867) 

Dadobia immersa (Erichson, 1837) 

Dinaraea aequata (Erichson, 1837) 

Dinaraea linearis (Gravenhorst, 1802) 

147


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148 

Traumoecia picipes (Thomson, 1856) 

Thamiaraea hospita (Märkel, 1844) 

Bolitochara lucida (Gravenhorst, 1802) 

Bolitochara obliqua Erichson, 1837 

Leptusa fumida (Erichson, 1839) 

Leptusa norvegica Strand, A., 1941 

Leptusa pulchella (Mannerheim, 1830) 

Leptusa ruficollis (Erichson, 1839) 

Agaricochara latissima (Stephens, 1832) 

Gyrophaena affinis Mannerheim, 1830 

Gyrophaena bihamata Thomson, C.G., 1867 

Gyrophaena fasciata (Marsham, 1802) 

Gyrophaena gentilis Erichson, 1839 

Gyrophaena manca Erichson, 1839 

Gyrophaena minima Erichson, 1837 

Gyrophaena nana (Paykull, 1800) 

Gyrophaena poweri Crotch, 1867 

Gyrophaena pulchella Heer, 1839 

Gyrophaena strictula Erichson, 1839 

Anomognathus cuspidatus (Erichson, 1839) 

Homalota plana (Gyllenhal, 1810) 

Cypha seminulum (Erichson, 1839) 

Oligota apicata (Erichson, 1837) 

Ischnoglossa prolixa (Gravenhorst, 1802) 

Phloeopora testacea (Mannerheim, 1830) 

Subfamily: SCAPHIDIINAE Shining Fungus Beetles 

Subfamily: PIESTINAE 

Scaphisoma agaricinum (Linnaeus, 1758) 

Scaphisoma boleti (Panzer, 1793) 

Siagonium quadricorne Kirby, 1815


____________________________ 

Subfamily: STAPHYLININAE 

LUCANIDAE Stag Beetles 

CLAMBIDAE 

SCIRTIDAE 

Atrecus affinis (Paykull, 1789) 

Gabrius splendidulus (Gravenhorst, 1802) 

Quedius brevicornis (Thomson, 1860) 

Quedius lyszkowskii Lott, 2010 

Quedius plagiatus Mannerheim, 1843 

Quedius truncicola Fairmaire & Laboulbène, 1856 

Sinodendron cylindricum (Linnaeus, 1758) Rhinoceros Beetle 

Clambus pallidulus Reitter, 1911 

Prionocyphon serricornis (Müller, P.W.J., 1821) 

EUCNEMIDAE False Click Beetles 

Melasis buprestoides (Linnaeus, 1761) 

ELATERIDAE Click Beetles 

Calambus bipustulatus (Linnaeus, 1767) 

Denticollis linearis (Linnaeus, 1758) 

Ampedus balteatus (Linnaeus, 1758) 

Ampedus pomonae (Stephens, 1830) 

Ampedus pomorum (Herbst, 1784) 

Melanotus castanipes (Paykull, 1800) 

LYCIDAE Net-winged Beetles 

Pyropterus nigroruber (De Geer, 1774) 

CANTHARIDAE Soldier Beetles 

Malthinus balteatus Suffrian, 1851 

Malthinus flaveolus (Herbst, 1786) 

Malthinus seriepunctatus Kiesenwetter, 1852 

Malthodes dispar (Germar, 1824) 

Malthodes flavoguttatus Kiesenwetter, 1852 

149


____________________________ 

150 

Malthodes fuscus (Waltl, 1838) 

Malthodes guttifer Kiesenwetter, 1852 

Malthodes marginatus (Latreille, 1806) 

Malthodes minimus (Linnaeus, 1758) 

Malthodes pumilus (Brébisson, 1835) 

BOSTRICHIDAE False Powder-post Beetles 

ANOBIIDAE 

LYMEXYLIDAE 

PHLOIOPHILIDAE 

Lyctus linearis (Goeze, 1777) 

Hedobia imperialis (Linnaeus, 1767) 

Ptinus subpilosus Sturm, 1837 

Grynobius planus (Fabricius, 1787) 

Dryophilus pusillus (Gyllenhal, 1808) 

Ochina ptinoides (Marsham, 1802) 

Xestobium rufovillosum (De Geer, 1774) Deathwatch Beetle 

Ernobius mollis (Linnaeus, 1758) 

Anobium inexpectatum Lohse, 1954 

Anobium punctatum (De Geer, 1774) Furniture Beetle 

Ptilinus pectinicornis (Linnaeus, 1758) 

Dorcatoma dresdensis Herbst, 1792 

Hylecoetus dermestoides (Linnaeus, 1761) 

Phloiophilus edwardsii Stephens, 1830 

TROGOSSITIDAE Flat Beetles 

Thymalus limbatus (Fabricius, 1787) 

CLERIDAE Checkered Beetles 

DASYTIDAE 

Tillus elongatus (Linnaeus, 1758) 

Thanasimus formicarius (Linnaeus, 1758) 

Dasytes aeratus Stephens, 1830


____________________________ 

MALACHIIDAE Malachite Beetles 

ASPIDIPHORIDAE 

Malachius bipustulatus (Linnaeus, 1758) 

Anthocomus fasciatus (Linnaeus, 1758) 

Aspidiphorus orbiculatus (Gyllenhal, 1808) 

NITIDULIDAE Sap or Blossom Beetles 

MONOTOMIDAE 

Epuraea aestiva (Linnaeus, 1758) 

Epuraea angustula Sturm, 1844 

Epuraea biguttata (Thunberg, 1784) 

Epuraea distincta (Grimmer, 1841) 

Epuraea longula Erichson, 1845 

Epuraea marseuli Reitter, 1872 

Epuraea melanocephala (Marsham, 1802) 

Epuraea melina Erichson, 1843 

Epuraea pallescens (Stephens, 1835) 

Epuraea rufomarginata (Stephens, 1830) 

Epuraea silacea (Herbst, 1783) 

Epuraea terminalis (Mannerheim, 1843) 

Epuraea thoracica Tournier, 1872 

Soronia grisea (Linnaeus, 1758) 

Soronia punctatissima (Illiger, 1794) 

Glischrochilus quadripunctatus (Linnaeus, 1758) 

Glischrochilus hortensis (Geoffroy in Fourcroy, 1785) 

Glischrochilus quadriguttatus (Fabricius, 1777) 

Pityophagus ferrugineus (Linnaeus, 1758) 

Rhizophagus bipustulatus (Fabricius, 1793) 

Rhizophagus cribratus Gyllenhal, 1827 

Rhizophagus depressus (Fabricius, 1793) 

Rhizophagus dispar (Paykull, 1800) 

151


____________________________ 

LAEMOPHLOEIDAE 

152 

Rhizophagus ferrugineus (Paykull, 1800) 

Rhizophagus parallelocollis Gyllenhal, 1827 

Rhizophagus perforatus Erichson, 1845 

Cryptolestes ferrugineus (Stephens, 1831) 

CRYPTOPHAGIDAE Silken Fungus Beetles 

Subfamily CRYPTOPHAGINAE 

Henoticus serratus (Gyllenhal, 1808) 

Cryptophagus dentatus (Herbst, 1793) 

Cryptophagus intermedius Bruce, 1934 

Cryptophagus denticulatus Heer, 1841 

Cryptophagus ruficornis Stephens, 1830 

Subfamily ATOMARIINAE 

EROTYLIDAE 

CERYLONIDAE 

Atomaria pulchra Erichson, 1846 

Atomaria umbrina (Gyllenhal, 1827) 

Dacne bipustulata (Thunberg, 1781) 

Triplax aenea (Schaller, 1783) 

Cerylon fagi Brisout de Barneville, 1867 

Cerylon ferrugineum Stephens, 1830 

Cerylon histeroides (Fabricius, 1793) 

ENDOMYCHIDAE False Ladybirds 

CORYLOPHIDAE 

Endomychus coccineus (Linnaeus, 1758) 

Orthoperus nigrescens Stephens, 1829 

LATRIDIIDAE Brown Scavenger or Plaster Beetles 

Enicmus testaceus (Stephens, 1830) 

MYCETOPHAGIDAE Hairy Fungus Beetles 

Litargus connexus (Geoffroy in Fourcroy, 1785)


____________________________ 

Mycetophagus multipunctatus Fabricius, 1793 

CIIDAE Minute Tree Fungus Beetles 

TETRATOMIDAE 

Octotemnus glabriculus (Gyllenhal, 1827) 

Orthocis alni (Gyllenhal, 1813) 

Cis bidentatus (Olivier, 1790) 

Cis bilamellatus Wood, 1884 

Cis boleti (Scopoli, 1763) 

Cis fagi Waltl, 1839 

Cis festivus (Panzer, 1793) 

Cis hispidus (Paykull, 1798) 

Cis nitidus (Fabricius, 1792) 

Cis vestitus Mellié, 1849 

Cis villosulus (Marsham, 1802) 

Ennearthron cornutum (Gyllenhal, 1827) 

Hallomenus binotatus (Quensel, 1790) 

Tetratoma ancora Fabricius, 1791 

Tetratoma fungorum Fabricius, 1790 

MELANDRYIDAE False Darkling Beetles 

Orchesia micans (Panzer, 1793) 

Orchesia minor Walker, 1836 

Orchesia undulata Kraatz, 1853 

Abdera flexuosa (Paykull, 1799) 

Melandrya caraboides (Linnaeus, 1761) 

Conopalpus testaceus (Olivier, 1790) 

MORDELLIDAE Tumbling Flower Beetles 

Mordellistena neuwaldeggiana (Panzer, 1796) 

COLYDIIDAE Cylindrical Bark Beetles 

Pycnomerus angulatus (Broun, 1893) 

Pycnomerus fuliginosus Erichson, 1842 

153


____________________________ 

154 

Synchita humeralis (Fabricius, 1792) 

TENEBRIONIDAE Darkling Beetles 

Tenebrio molitor Linnaeus, 1758 Mealworm Beetle 

Palorus ratzeburgi (Wissmann, 1848) 

Nalassus laevioctostriatus (Goeze, 1777) 

Gonodera luperus (Herbst, 1783) 

OEDEMERIDAE False Blister Beetles 

Nacerdes melanura (Linnaeus, 1758) Wharf Borer 

Ischnomera sanguinicollis (Fabricius, 1787) 

PYROCHROIDAE Cardinal Beetles 

Pyrochroa serraticornis (Scopoli, 1763) Red-headed Cardinal Beetle 

SALPINGIDAE Narrow-waisted Bark Beetles 

SCRAPTIIDAE 

Lissodema denticolle (Gyllenhal, 1813) 

Rabocerus foveolatus (Ljungh, 1824) 

Rabocerus gabrieli Gerhardt, 1901 

Sphaeriestes ater (Paykull, 1798) 

Sphaeriestes castaneus (Panzer, 1796) 

Sphaeriestes reyi (Abeille, 1874) 

Vincenzellus ruficollis (Panzer, 1794) 

Salpingus planirostris (Fabricius, 1787) 

Salpingus ruficollis (Linnaeus, 1761) 

Anaspis fasciata (Forster, 1771) 

Anaspis frontalis (Linnaeus, 1758) 

Anaspis garneysi Fowler, 1889 

Anaspis lurida Stephens, 1832 

Anaspis maculata (Geoffroy in Fourcroy, 1785) 

Anaspis regimbarti Schilsky, 1895 

Anaspis rufilabris (Gyllenhal, 1827) 

Anaspis thoracica (Linnaeus, 1758)


____________________________ 

CERAMBYCIDAE Longhorn Beetles 

Rhagium bifasciatum Fabricius, 1775 

Rhagium mordax (De Geer, 1775) 

Grammoptera abdominalis (Stephens, 1831) 

Grammoptera ruficornis (Fabricius, 1781) 

Grammoptera ustulata (Schaller, 1783) 

Leptura aurulenta (Fabricius, 1793) Hornet Beetle 

Leptura quadrifasciata (Linnaeus, 1758) Four-banded Longhorn 

Anoplodera sexguttata (Fabricius, 1775) 

Alosterna tabacicolor (De Geer, 1775) 

Rutpela maculata (Poda, 1761) 

Stenurella melanura (Linnaeus, 1758) 

Asemum striatum (Linnaeus, 1758) 

Nathrius brevipennis (Mulsant, 1839) 

Aromia moschata (Linnaeus, 1758) Musk Beetle 

Callidium violaceum (Linnaeus, 1758) Violet Longhorn 

Phymatodes testaceus (Linnaeus, 1758) 

Clytus arietis (Linnaeus, 1758) Wasp Beetle 

Lamia textor (Linnaeus, 1758) The Weaver Beetle 

Pogonocherus hispidulus (Piller & Mitterpacher, 1783) 

Pogonocherus hispidus (Linnaeus, 1758) 

Leiopus nebulosus (Linnaeus, 1758) 

Tetrops praeustus (Linnaeus, 1758) 

ANTHRIBIDAE Fungus Weevils 

Choragus sheppardi Kirby, W., 1819 

CURCULIONIDAE Weevils 

Rhopalomesites tardyi (Curtis, 1825) Holly Weevil 

Pselactus spadix (Herbst, 1795) 

Pseudophloeophagus aeneopiceus (Boheman, 1845) 

Euophryum confine (Broun, 1881) 

155


____________________________ 

156 

Euophryum rufum (Broun, 1880) 

Pentarthrum huttoni Wollaston, 1854 

Macrorhyncolus littoralis (Broun, 1880) 

Cryptorhynchus lapathi (Linnaeus, 1758) 

Acalles misellus Boheman, 1844 

Acalles ptinoides (Marsham, 1802) 

Kyklioacalles roboris (Curtis, 1835) 

Magdalis armigera (Fourcroy, 1785) 

Magdalis carbonaria (Linnaeus, 1758) 

Magdalis ruficornis (Linnaeus, 1758) 

Hylobius abietis (Linnaeus, 1758) 

Pissodes castaneus (De Geer, 1775) 

Pissodes pini (Linnaeus, 1758) 

Subfamily SCOLYTINAE Bark Beetles 

Scolytus multistriatus (Marsham, 1802) Small Elm Bark Beetle 

Scolytus rugulosus (Müller, P.W.J., 1818) 

Scolytus scolytus (Fabricius, 1775) Common Elm Bark Beetle 

Pityophthorus pubescens (Marsham, 1802) 

Cryphalus asperatus (Gyllenhal, 1813) 

Dryocoetes autographus (Ratzeburg, 1837) 

Dryocoetes villosus (Fabricius, 1793) 

Orthotomicus laricis (Fabricius, 1792) 

Pityogenes bidentatus (Herbst, 1783) 

Trypodendron domesticum (Linnaeus, 1758) 

Trypodendron lineatum (Olivier, 1795) Conifer Ambrosia Beetle 

Hylesinus crenatus (Fabricius, 1787) Large Ash Bark Beetle 

Hylesinus varius (Fabricius, 1775) Common Ash Bark Beetle 

Pteleobius vittatus (Fabricius, 1787) 

Hylastes ater (Paykull, 1800) 

Hylastes brunneus Erichson, 1836


____________________________ 

Hylastes opacus Erichson, 1836 

Hylurgops palliatus (Gyllenhal, 1813) 

Tomicus piniperda (Linnaeus, 1758) Pine Shoot Beetle 

157


____________________________ 

Index of beetle species 

158 

A 

Abdera flexuosa................................. 12, 90, 91, 132, 153 

Abraeus perpusillus ...............................................17, 146 

Acalles misellus....................................................119, 156 

Acalles ptinoides..................................................121, 156 

Acrulia inflata ........................................................22, 147 

Agaricochara latissima ..........................................32, 148 

Agathidium confusum ....................... 12, 14, 20, 131, 146 

Agathidium nigripenne..........................................20, 146 

Agathidium rotundatum....................................4, 20, 146 

Agathidium seminulum..........................................21, 146 

Agathidium varians ...............................................21, 146 

Alaobia pallidicornis ..............................................28, 147 

Alosterna tabacicolor...................................107, 134, 155 

Ampedus balteatus................................................44, 149 

Ampedus pomonae.............................. 9, 11, 45, 138, 149 

Ampedus pomorum .........................................45, 46, 149 

Amphicyllis globus .............................................4, 20, 146 

Anaspis fasciata...................................................100, 154 

Anaspis frontalis ..................................................100, 154 

Anaspis garneysi..................................................101, 154 

Anaspis lurida ......................................................101, 154 

Anaspis maculata ................................................102, 154 

Anaspis regimbarti...............................................102, 154 

Anaspis rufilabris .................................................102, 154 

Anaspis thoracica ................................................103, 154 

Anisotoma humeralis.............................................19, 146 

Anisotoma orbicularis................................ 10, 12, 19, 146 

Anobium inexpectatum .........................................56, 150 

Anobium punctatum..............................................56, 150 

Anomognathus cuspidatus ....................................35, 148 

Anoplodera sexguttata..................................10, 107, 155 

Anthocomus fasciatus........................................4, 62, 151 

Aromia moschata ........................................111, 139, 155 

Asemum striatum ........................................110, 142, 155 

Aspidiphorus orbiculatus .......................................63, 151 

Atheta basicornis...................................................29, 147 

Atheta pilicornis .............................................. 13, 29, 147 

Atomaria pulchra .......................................... 75, 133, 152 

Atomaria umbrina................................................. 75, 152 

Atrecus affinis........................................................ 39, 149 

B 

Bessobia fungivora........................................ 29, 133, 147 

Bibloporus bicolor ........................................... 12, 27, 147 

Bolitochara lucida ................................................. 31, 148 

Bolitochara obliqua............................................... 31, 148 

C 

Calambus bipustulatus...................................... 4, 44, 149 

Callidium violaceum ............................................ 111, 155 

Cerylon fagi ........................................................... 77, 152 

Cerylon ferrugineum.............................................. 77, 152 

Cerylon histeroides................................................ 78, 152 

Choragus sheppardi ............................................ 116, 155 

Cis bidentatus........................................................ 82, 153 

Cis bilamellatus ..................................................... 82, 153 

Cis boleti................................................................ 82, 153 

Cis fagi................................................................... 82, 153 

Cis festivus............................................................. 83, 153 

Cis hispidus............................................................ 83, 153 

Cis nitidus .............................................................. 84, 153 

Cis vestitus............................................................. 85, 153 

Cis villosulus .................................................... 12, 85, 153 

Clambus pallidulus ................................................ 42, 149 

Clytus arietis................................................ 112, 131, 155 

Conopalpus testaceus ....................................... 4, 92, 153 

Coryphium angusticolle......................................... 22, 146 

Cryphalus asperatus............................................ 125, 156 

Cryptolestes ferrugineus ....................................... 73, 152 

Cryptophagus dentatus......................................... 74, 152 

Cryptophagus denticulatus ................................... 74, 152 

Cryptophagus intermedius .................................... 74, 152 

Cryptophagus ruficornis ........................................ 74, 152 

Cryptorhynchus lapathi ....................................... 119, 156


____________________________ 

Cypha seminulum.............................................. 4, 36, 148 

D 

Dacne bipustulata ..................................76, 132, 142, 152 

Dadobia immersa.................................................. 30, 147 

Dasytes aeratus..................................................... 61, 150 

Denticollis linearis ................................................. 44, 149 

Dinaraea aequata ................................................. 30, 147 

Dinaraea linearis ................................................... 30, 147 

Dorcatoma dresdensis..................12, 13, 57, 58, 131, 150 

Dropephylla devillei............................................... 23, 147 

Dropephylla gracilicornis....................................... 23, 147 

Dropephylla ioptera .............................................. 23, 147 

Dropephylla koltzei................................................ 24, 147 

Dryocoetes autographus..................................... 125, 156 

Dryocoetes villosus.............................................. 126, 156 

Dryophilus pusillus ................................................ 54, 150 

E 

Endomychus coccineus.............................78, 79, 132, 152 

Enicmus testaceus......................................... 80, 132, 152 

Ennearthron cornutum.......................................... 85, 153 

Epuraea aestiva..................................................... 63, 151 

Epuraea angustula ................................................ 63, 151 

Epuraea biguttata................................................. 63, 151 

Epuraea distincta .......................................... 64, 132, 151 

Epuraea longula .................................................... 64, 151 

Epuraea marseuli .................................................. 65, 151 

Epuraea melanocephala ....................................... 65, 151 

Epuraea melina ..................................................... 66, 151 

Epuraea pallescens................................................ 66, 151 

Epuraea rufomarginata......................................... 66, 151 

Epuraea silacea ..................................................... 67, 151 

Epuraea terminalis.......................................... 14, 68, 151 

Epuraea thoracica ................................................. 68, 151 

Ernobius mollis ...................................................... 55, 150 

Euophryum confine ........................93, 118, 132, 138, 155 

Euophryum rufum ....................................... 118, 139, 156 

Euplectus bescidicus.............................................. 26, 147 

Euplectus infirmus................................................. 26, 147 

Euplectus karstenii ................................................ 26, 147 

Euplectus mutator................................................. 26, 147 

Euplectus piceus.............................................. 13, 26, 147 

Euplectus punctatus...........................................4, 26, 147 

G 

Gabrius splendidulus..............................................39, 149 

Glischrochilus hortensis .........................69, 134, 140, 151 

Glischrochilus quadriguttatus................................70, 151 

Glischrochilus quadripunctatus..............................69, 151 

Gonodera luperus ..................................................94, 154 

Grammoptera abdominalis..................................105, 155 

Grammoptera ruficornis ......................................105, 155 

Grammoptera ustulata..................................12, 105, 155 

Grynobius planus ...................................................54, 150 

Gyrophaena affinis ................................................32, 148 

Gyrophaena bihamata...........................................33, 148 

Gyrophaena fasciata..............................................33, 148 

Gyrophaena gentilis...............................................33, 148 

Gyrophaena manca ...............................................34, 148 

Gyrophaena minima ..............................................34, 148 

Gyrophaena nana ..................................................34, 148 

Gyrophaena poweri ...............................................34, 148 

Gyrophaena pulchella............................................34, 148 

Gyrophaena strictula .....................................34, 135, 148 

H 

Hallomenus binotatus....................................86, 141, 153 

Hapalaraea pygmaea ......................................14, 24, 147 

Hedobia imperialis.................................................53, 150 

Henoticus serratus.................................................74, 152 

Homalota plana.....................................................35, 148 

Hylastes ater........................................................129, 156 

Hylastes brunneus................................................129, 156 

Hylastes opacus ...................................................129, 157 

Hylecoetus dermestoides.................................11, 58, 150 

Hylesinus crenatus...............................................128, 156 

Hylesinus varius .............................................74, 128, 156 

Hylobius abietis....................................................123, 156 

Hylurgops palliatus ..............................................130, 157 

I 

Ischnoglossa prolixa...............................................36, 148 

Ischnomera sanguinicollis........................10, 95, 142, 154 

159


____________________________ 

160 

K 

Kyklioacalles roboris ............................................121, 156 

L 

Lamia textor ........................................................113, 155 

Leiopus nebulosus................................ 114, 116, 133, 155 

Leptura aurulenta ................................. 11, 105, 107, 155 

Leptura quadrifasciata ........................................107, 155 

Leptusa fumida......................................................31, 148 

Leptusa norvegica..........................................31, 133, 148 

Leptusa pulchella...................................................31, 148 

Leptusa ruficollis............................................32, 133, 148 

Lissodema denticolle........................................11, 97, 154 

Litargus connexus..........................................80, 132, 152 

Lyctus linearis ........................................................53, 150 

M 

Macrorhyncolus littoralis.....................................119, 156 

Magdalis armigera ..............................................121, 156 

Magdalis carbonaria....................................122, 131, 156 

Magdalis ruficornis..............................................123, 156 

Malachius bipustulatus............................ 61, 62, 136, 151 

Malthinus balteatus...............................................48, 149 

Malthinus flaveolus ...............................................48, 149 

Malthinus seriepunctatus ......................................48, 149 

Malthodes dispar...............................................4, 49, 149 

Malthodes flavoguttatus .......................................49, 149 

Malthodes fuscus...................................................50, 150 

Malthodes guttifer.........................................51, 141, 150 

Malthodes marginatus ..........................................51, 150 

Malthodes minimus...............................................52, 150 

Malthodes pumilus ................................................52, 150 

Melandrya caraboides.....................................10, 91, 153 

Melanotus castanipes.............................. 46, 47, 138, 149 

Melasis buprestoides ............................... 11, 43, 140, 149 

Mordellistena neuwaldeggiana.......................11, 92, 153 

Mycetophagus multipunctatus................ 80, 81, 132, 153 

N 

Nacerdes melanura ...............................................94, 154 

Nalassus laevioctostriatus .....................................94, 154 

Nathrius brevipennis............................................110, 155 

Nossidium pilosellum ........................................ 4, 17, 146 

O 

Ochina ptinoides ................................................... 54, 150 

Octotemnus glabriculus ........................................ 81, 153 

Oligota apicata...................................................... 36, 148 

Orchesia micans .............................................. 87, 88, 153 

Orchesia minor...................................................... 88, 153 

Orchesia undulata................................................. 89, 153 

Orthocis alni .......................................................... 81, 153 

Orthoperus nigrescens .......................................... 79, 152 

Orthotomicus laricis ............................................ 126, 156 

P 

Palorus ratzeburgi................................................. 93, 154 

Pentarthrum huttoni ........................................... 118, 156 

Phloeocharis subtilissima ...................................... 27, 147 

Phloeonomus punctipennis ................................... 25, 147 

Phloeonomus pusillus............................................ 25, 147 

Phloeopora testacea ............................................. 36, 148 

Phloeostiba plana.................................................. 25, 147 

Phloiophilus edwardsii........................................... 59, 150 

Phyllodrepoidea crenata ....................................... 22, 146 

Phymatodes testaceus ........................ 111, 112, 133, 155 

Pissodes castaneus.............................................. 123, 156 

Pissodes pini................................................ 123, 134, 156 

Pityogenes bidentatus......................................... 127, 156 

Pityophagus ferrugineus ....................................... 70, 151 

Pityophthorus pubescens .................................... 125, 156 

Pogonocherus hispidulus..................................... 113, 155 

Pogonocherus hispidus........................................ 113, 155 

Prionocyphon serricornis..........42, 43, 132, 135, 140, 149 

Pselactus spadix .................................................. 117, 155 

Pseudophloeophagus aeneopiceus ............. 117, 134, 155 

Pteleobius vittatus............................................... 129, 156 

Ptenidium laevigatum ........................................... 17, 146 

Pteryx suturalis...................................................... 18, 146 

Ptilinus pectinicornis ....................................... 57, 60, 150 

Ptinella aptera....................................................... 18, 146 

Ptinella cavelli ....................................................... 18, 146 

Ptinella denticollis ................................................. 18, 146 

Ptinella errabunda................................................. 18, 146 

Ptinella limbata..................................................... 18, 146


____________________________ 

Ptinella taylorae.................................................... 18, 146 

Ptinus subpilosus................................13, 53, 54, 132, 150 

Pycnomerus angulatus.......................................... 92, 153 

Pycnomerus fuliginosus................................. 93, 140, 153 

Pyrochroa serraticornis ........4, 95, 96, 134, 140, 143, 154 

Pyropterus nigroruber................10, 14, 47, 134, 142, 149 

Q 

Quedius brevicornis............................................... 39, 149 

Quedius lyszkowskii......................................9, 38, 40, 149 

Quedius plagiatus ................................................. 40, 149 

Quedius truncicola ................................................ 40, 149 

R 

Rabocerus foveolatus............................................ 97, 154 

Rabocerus gabrieli................................................. 97, 154 

Rhagium bifasciatum .......................................... 103, 155 

Rhagium mordax................................................. 103, 155 

Rhizophagus bipustulatus ..................................... 71, 151 

Rhizophagus cribratus........................................... 71, 151 

Rhizophagus depressus ......................................... 72, 151 

Rhizophagus dispar ............................................... 72, 151 

Rhizophagus ferrugineus....................................... 72, 152 

Rhizophagus parallelocollis ................................... 73, 152 

Rhizophagus perforatus ........................................ 73, 152 

Rhopalomesites tardyi................................8, 60, 117, 155 

Rutpela maculata................................................ 108, 155 

S 

Salpingus planirostris ............................................ 99, 154 

Salpingus ruficollis................................................. 99, 154 

Scaphisoma agaricinum ........................................ 37, 148 

Scaphisoma boleti ................................................. 38, 148 

Scolytus multistriatus.................................. 124, 139, 156 

Scolytus rugulosus............................................... 124, 156 

Scolytus scolytus ..................................................124, 156 

Sepedophilus littoreus....................................28, 132, 147 

Siagonium quadricorne............................38, 39, 133, 148 

Sinodendron cylindricum .........................1, 5, 41, 42, 149 

Soronia grisea ........................................................68, 151 

Soronia punctatissima ...........................................69, 151 

Sphaeriestes ater ...........................................97, 133, 154 

Sphaeriestes castaneus..........................................98, 154 

Sphaeriestes reyi............................................98, 133, 154 

Stenichnus bicolor............................................13, 21, 146 

Stenurella melanura ............................................109, 155 

Synchita humeralis.................................................93, 154 

T 

Tenebrio molitor ....................................................93, 154 

Tetratoma ancora..........................................86, 135, 153 

Tetratoma fungorum ...............................86, 87, 132, 153 

Tetrops praeustus ................................................116, 155 

Thamiaraea hospita...............................................31, 148 

Thanasimus formicarius...........................60, 61, 131, 150 

Thymalus limbatus.................................................59, 150 

Tillus elongatus......................................................60, 150 

Tomicus piniperda................................................130, 157 

Traumoecia picipes ................................................30, 148 

Triplax aenea .........................................................76, 152 

Trypodendron domesticum............................63, 127, 156 

Trypodendron lineatum .......................................128, 156 

V 

Vincenzellus ruficollis.............................................98, 154 

X 

Xestobium rufovillosum .........................................55, 150 

Xylostiba monilicornis............................................25, 147 

161

The beetles of decaying wood in Ireland - National Parks & Wildlife ...

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