The Genus Amorphophallus - Bjorn Malkmus Rare Seed Nursery
The Genus Amorphophallus - Bjorn Malkmus Rare Seed Nursery
The Genus Amorphophallus - Bjorn Malkmus Rare Seed Nursery
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<strong>The</strong> <strong>Genus</strong> <strong>Amorphophallus</strong> (Titan Arums)<br />
Origin, Habit and General Information<br />
<strong>The</strong> genus <strong>Amorphophallus</strong> is<br />
well known for the famous<br />
<strong>Amorphophallus</strong> titanum,<br />
commonly known as "Titan<br />
Arum". <strong>The</strong> Titan Arum holds the<br />
plant world record for an<br />
unbranched single<br />
inflorescence. <strong>The</strong> infloresence<br />
eventually may reach up to three<br />
meters and more in height.<br />
Besides this<br />
oustanding species more than 200 <strong>Amorphophallus</strong> species have been<br />
described - and each year some more new findings are published. A<br />
more or less complete list of all validly described <strong>Amorphophallus</strong><br />
species and many photos are available from the website of the<br />
International Aroid Society (http://www.aroid.org). If you are<br />
interested in this fascinating genus, think about becoming a member of<br />
the International Aroid Society! <strong>The</strong> International Aroid Society is the<br />
worldwide leading society in aroids and offers a membership at a very<br />
low price and with many benefits!<br />
A different website for those interested in <strong>Amorphophallus</strong> hybrids is:<br />
www.amorphophallus-network.org This page features some awe-inspiring new hybrids,<br />
e.g. <strong>Amorphophallus</strong> 'John Tan' - an unique and first time ever cross between<br />
<strong>Amorphophallus</strong> variabilis X <strong>Amorphophallus</strong> titanum!<br />
<strong>The</strong> majority of<br />
<strong>Amorphophallus</strong> species is<br />
native to subtropical and<br />
tropical lowlands of forest<br />
margins and open, disturbed<br />
spots in woods throughout<br />
Asia. Few species are found in<br />
Africa (e.g. <strong>Amorphophallus</strong><br />
abyssinicus, from West to<br />
East Africa), Australia<br />
(represented by a single<br />
species only, namely <strong>Amorphophallus</strong> galbra, occuring in Queensland, North Australia<br />
and Papua New Guinea), and Polynesia respectively. Few species, such as<br />
<strong>Amorphophallus</strong> paeoniifolius (Madagascar to Polynesia), serve as a food source<br />
throughout the Asian region.<br />
As in other related genera within the broad Aroid family (Araceae), such as Arisaema,<br />
Dracontium, and Typhonium, most species develop a large, subterranean round to vertical<br />
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elongated tuber or fleshy rhizome, which serves as a storage organ and may gain up to<br />
60kg and more in the tallest species, such as<br />
<strong>Amorphophallus</strong> hewitii (Sarawak, East<br />
Malaysia), <strong>Amorphophallus</strong> titanum (Sumatra,<br />
Indonesia), and <strong>Amorphophallus</strong> gigas<br />
(Sumatra, Indonesia). In most species a single,<br />
long-stemmed, umbrella-like, dissected leaf is<br />
developed during each growing cycle. This single<br />
leaf resembles almost a small tree in the above<br />
mentioned giant members within the genus. In<br />
certain species, such as <strong>Amorphophallus</strong><br />
bulbifer (widespread throughout India), a<br />
secondary leaf may develop under favorable growing conditions. <strong>Seed</strong>lings and young<br />
plants received via in-vitro propagation or by means of leaf cuttings tend to make several<br />
subsequent leaves before they will enter their true first dormancy period.<br />
On maturity, plants usually produce a single inflorescence before the leaf unfolds. Few<br />
species may develop several inflorescences per tuber (e.g. <strong>Amorphophallus</strong><br />
polyanthus) and/or make their inflorescence after laf development. Some<br />
<strong>Amorphophallus</strong> species will rest for an entire year after flowering, i.e. they will skip one<br />
growing cycle. <strong>The</strong> inflorescence is composed of a sometimes very colorful outer spathe (a<br />
pseudo petal) and an inner spadix. <strong>The</strong> spadix may be shorter or protruding the spathe,<br />
and usually has a different color. <strong>The</strong> spadix hosts quite small male and female flowers at<br />
its base. <strong>The</strong> upper part (e.g. the spadix appendix) is usually the only visible part of the<br />
spadix. On the day of opening the female flowers are receptive and the spaidx releases a<br />
sometimes very unpleasant strong scent, which is very attractive for its main pollinators,<br />
such as flies, and certain beetles. On the second day (or later) the female flowers are not<br />
receptive anymore, whereas the male flowers release their pollen. Thus one would usually<br />
need two inflorescence at more or less the same time in order to successfully transmit<br />
polen from a first inflorescence to a second flowering plant. Few species are apomictic and<br />
may set viable seeds without pollination (e.g. <strong>Amorphophallus</strong> bulbifer,<br />
<strong>Amorphophallus</strong> henryi, <strong>Amorphophallus</strong> kiusianus, and <strong>Amorphophallus</strong> muelleri).<br />
After successful pollination the female flowers develop into some colorful berries, which<br />
may contain one to rarely two or three seeds each. In most species berries are bright<br />
orange to scarlet, whereas some may be bluish-lilac (e.g. <strong>Amorphophallus</strong> kiusianus<br />
from South Japan, East China, and Taiwan) or rarely whitish-green to yellowish-green.<br />
<strong>The</strong> ovate to elliptic berries are somewhat fleshy and are mostly dispersed by birds.<br />
Cultivation<br />
In cultivation, most species are easily grown in any humus-rich, organic, very well drained<br />
and well aerated soil. <strong>The</strong> majority of <strong>Amorphophallus</strong> species can be grown at some 20°C<br />
(optimal 25°C) during summer in a semi-moist soil in a partially shaded spot with<br />
protection from direct sunlight, especially during hours at midday. Few tropical species,<br />
such as the famous <strong>Amorphophallus</strong> titanum from Sumatra, require an overall higher<br />
average temperature of some 25°C throughout their entire growing cycle and during<br />
dormancy as well. Few species (mostly the beforehand mentioned tropical species) have a<br />
prolonged growing cycle which exceeds 12 months, or they may be even evergreen (e.g.<br />
<strong>Amorphophallus</strong> coaetaneus). However most species will follow seasons and their<br />
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single leaf will yellow from late<br />
summer to early winter. At this<br />
point, plants should be kept<br />
drier at a minimum of some<br />
15°C. Tropical species require a<br />
higher temperature and very<br />
careful watering during<br />
dormancy. It is not essential,<br />
that dormant tubers are kept<br />
completely dry, but soil should<br />
not be soaked wet at any time<br />
during dormancy. As a matter of fact, a dry storage will inevitably lead to losses in most<br />
tropical species. However, most African species from seasonally dry areas (e.g.<br />
<strong>Amorphophallus</strong> andranogydroensis, <strong>Amorphophallus</strong> consimilis, and<br />
<strong>Amorphophallus</strong> dracontioides) are best kept completely dry during dormancy.<br />
Formerly, <strong>Amorphophallus</strong> species had been separated into two distinct cultivation groups<br />
with different cultivation requirements concerning soil composition, minimum temperatures<br />
and water suppply during dormancy. On grounds of some more recent and more detailed<br />
cultivation experiences, most species can be successfully grown in one and the same soil<br />
mixture (see above). Different temperatures are not necessary during dormancy either.<br />
However, several highland species will come into growth earlier, probably in mid winter<br />
already, and hence they will require artificial light. It is not necessary either to keep soil<br />
completely dry during dormancy. Several species will show less vigor and minor growth, if<br />
the tuber is kept completey dry during dormancy. In most species, a slightly moist soil will<br />
be suitable (with exception for a couple of more delicate African species). As a general<br />
rule: the moister the soil, the higher temperatures have to be during dormancy. Tropical<br />
species require a temperature optimum of not below 25°C throughout the year, whereas<br />
some 15°C to 20°C are sufficient for all other species.<br />
Most tubers stay dormant for some 3 to 7 months. Increasing temperatures during daytime<br />
stimulate new growth in most <strong>Amorphophallus</strong> species from spring to mid summer.<br />
Tropical species are less predictable concerning their growth cycle and may well come into<br />
growth at any time of the year. Thus artificial light during winter months and heat are<br />
essential factors to successfully cultivate these species. As soon as the new leaf appears<br />
from the center of the tuber, plants shall be watered carefully on a more regular basis and<br />
feeded weekly till mid summer with any common fertilizer. Roots are developped primarily<br />
from the upper part of the tuber in most species, thus the tuber shall be planted deep<br />
enough, so that the roots may develop freely. Three times the size of the tuber is a<br />
suitable depth for most species.<strong>The</strong> tuber in immature specimens may increase its weight<br />
threefold or even more in a single season, hence the growing pot has to be of an adequate<br />
size to fit the newly developping tuber.<br />
Propagation by <strong>Seed</strong>s<br />
Most <strong>Amorphophallus</strong> are extremely easily raised from preferably fresh seeds. If seeds<br />
cannot be sowed immediately, they may be stored for some weeks or months. <strong>Seed</strong>s from<br />
<strong>Amorphophallus</strong> kiusianus and <strong>Amorphophallus</strong> konjac showed still good germination<br />
rates after more than 12 months storage at 7°C in semi-moist peat moss. However it is<br />
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essential that seeds are constantly kept slightly moist (e.g. in slightly moist peat moss) and<br />
at lower temperatures of some 10°C in order to delay the germination process. Once<br />
completely desiccated or frozen, <strong>Amorphophallus</strong> seeds will have lost their viability and<br />
they will not germinate anymore. Prior to sowing the outer flesh has to be removed by<br />
thoroughly rinsing it away with water. Wear some protective hand gloves, for the flesh<br />
contains some very irritant substances. <strong>The</strong> flesh also contains some chemical which act<br />
as germination inhibitors. <strong>The</strong>se susbtances will siginificantly delay germination, if the<br />
entire berry is sowed. <strong>Seed</strong>s are potted some 1 cm deep in the same soil mix as one<br />
would use for ault tubers. Soil should be kept slightly moist at all times. <strong>The</strong> optimal<br />
temperature is some 25°C at day<br />
and some 20°C at night. One should<br />
add a couple of degrees for all<br />
tropical species. Most seedlings will<br />
occur within some 2 to 6 weeks<br />
(examples from the left: 6 weeks old<br />
<strong>Amorphophallus</strong> gomboczianus<br />
seedlings and 6 months old<br />
<strong>Amorphophallus</strong> laoticus seedlings). Some may take a couple of weeks more. <strong>Seed</strong>s of<br />
<strong>Amorphophallus</strong> kiusianus may well need some 6 months and more to germinate. Most<br />
seedlings show a prolonged growing cycle in comparison to older and mature specimens<br />
within the same species. Thus pots should kept warm in partial shade throughout the year<br />
as long as seedlings are “in green”, i.e. as long as they have a leaf. Several seedlings may<br />
make not only one, but two or more subsequent leaves before they enter dormancy for the<br />
first time. As a general rule, young dormant seedling tubers, as well as any other small<br />
offset tubers or leaf bulbils, shall not be kept completely dry during dormancy, as they may<br />
desiccate more easily.<br />
Propagation by Offsets & Leaf Bulbils<br />
Apart from seeds, most species vegetatively propagate as well. <strong>The</strong>y either propagate by<br />
stoloniferous offsets (e.g. <strong>Amorphophallus</strong> albus, <strong>Amorphophallus</strong> antsingyensis,<br />
<strong>Amorphophallus</strong> bangkokensis, <strong>Amorphophallus</strong> impressus, <strong>Amorphophallus</strong><br />
krausei, and <strong>Amorphophallus</strong> konjac), by small adjacent tubers (e.g. <strong>Amorphophallus</strong><br />
thaiensis, <strong>Amorphophallus</strong> odoratus, <strong>Amorphophallus</strong><br />
paeoniifolius, and <strong>Amorphophallus</strong> yunnanensis) or rarely<br />
by leaf bulbils (e.g. <strong>Amorphophallus</strong> beccarii,<br />
<strong>Amorphophallus</strong> bufo, <strong>Amorphophallus</strong> bulbifer [see foto<br />
with large central leaf bulbil and three developping adjecent<br />
bulbils], <strong>Amorphophallus</strong> muelleri, <strong>Amorphophallus</strong><br />
yuloensis, and <strong>Amorphophallus</strong> xiei). <strong>The</strong>se offsets and<br />
small tubers shall stay attached to the main tuber until they can<br />
be separated easily. Leaf bulbils can be easily separated from the leaf after the leaf will<br />
have withered.<br />
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Propagation by Leaf Cuttings<br />
Leaf cuttings and in-vitro cultivation are some more dedicated advanced methods of<br />
vegetative propagation. Whereas in-vitro propagation requires some rather expensive<br />
material and special techniques and experience, it is quite easy and rather inexpensive to<br />
take leaf cuttings, even for not that experienced private cultivators. It has to be noted, that<br />
not all species can be successfully propagated from leaf cuttings. For example, so far this<br />
method has failed in <strong>Amorphophallus</strong> konjac and in <strong>Amorphophallus</strong> paeoniifolius.<br />
And in those species, which can be grown from leaf cuttings, not all cuttings will eventually<br />
make a tuber. But for the rarer species it is quite worth the effort, especially for those,<br />
which do not make any offsets or leaf bulbils (e.g. <strong>Amorphophallus</strong> borneensis,<br />
<strong>Amorphophallus</strong> decus-silvae, <strong>Amorphophallus</strong> gigas, and <strong>Amorphophallus</strong><br />
titanum). From the latter four species, <strong>Amorphophallus</strong> titanum is easily propagated via<br />
leaf cuttings, provided that leaflets are mature and of an adequate size.<br />
<strong>The</strong> following material will be necessary to take leaf<br />
cuttings:<br />
- a transparent plastic box with cover, large enough to host<br />
the cuttings<br />
- small pots with a bottom hole, pereferably transparent to<br />
easily control root growth<br />
- a sharp and clean knife and some alcohol solution for<br />
desinfection<br />
- some lables and a pencil<br />
- preferably some rooting hormone, especially formulated<br />
for leaf cuttings<br />
- finely granulated, clean washed pumice or any other<br />
inorganic small grained substrate<br />
A healthy leaf shall be chosen for taking cuttings. <strong>The</strong> cutting<br />
should be preferably made in the early morning hours, as<br />
long as the leaf is turgescent. <strong>The</strong> cut should be directly<br />
made above a branching point (see dotted red line).<br />
Under optimal conditions, the leaf is some 1 to 3 months old.<br />
Younger leaves may be too imature, whereas older leaves<br />
may be programmed to enter dormancy already.<br />
As an exception to the above rules, leaf cuttings can be<br />
taken at any time in a rescue try for a rare species with signs<br />
of a rotting tuber. It may prove easier to rescue a species by<br />
taking leaf cuttings instead of trying to rescue the infested<br />
tuber.<br />
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A typical cutting is some 10cm to 20cm (or more) long,<br />
depending on species and its size of leaflets. <strong>The</strong> cutting<br />
consists of at least one to three leaflets, maybe even some<br />
more in very small species to have a sufficiently large<br />
surface for photosynthesis.<br />
<strong>The</strong> cut should be made straight with a very sharp knife and<br />
without squeezing the tissue.<br />
As a general rule, from well grown plants 1/3 of a leaf can be<br />
taken. This way, the mother plant has a still sufficiently large<br />
leaf surface for photosynthesis and to increase its own tuber<br />
in size or to maintain its size. In very large mature<br />
specimens up to 1/2 of the leaf may be taken for leaf<br />
cuttings.<br />
In the left example of <strong>Amorphophallus</strong> macrorhizus, some<br />
1/4 was taken from the total leaf surface.<br />
In between cuttings, the knife blade should be desinfected,<br />
e.g. by dipping it into alcohol. Let the knife blade dry up<br />
before taking the next cutting.<br />
Preferably use a rooting hormone as directed on the<br />
package. Some species may be successfully propagated<br />
without using any rooting hormone. However, in general the<br />
success rate will be much lower and it may take longer for<br />
the cutting to make a new basal tuber.<br />
Do not forget to write a label for each pot in order to not get<br />
lost with identifications, especially if cuttings from more than<br />
one species are going to be taken!<br />
After taking the cutting, immediately dip it into the powder,<br />
shake off any excessive powder and plant the leaf cut<br />
vertically into a small pot. Depending on pot size, one or<br />
more cuttings will fit into a single pot (in the left example, two<br />
cuttings were planted). Pumice or any other inorganic<br />
substrate will work best. Organic substrates (such as peat<br />
based soils and the like) are less recommended, for these<br />
will usually promote bacterial and fungal growth.<br />
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Pests and Diseases<br />
Carefully place the small pot in a transparent plastic box and<br />
put on the cover to raise air humidity. <strong>The</strong> box should be<br />
large and tall enough, so that the leaflet tip will not touch the<br />
cover (possible rotting). Store the plastic box in a shaded<br />
place with some indirect sunlight at some 25°C to 30°C.<br />
Control humidity every other day and if necessary, mist the<br />
cuttings.<br />
Cuttings will start to produce some callus tissue after some 4<br />
to 12 weeks and a couple of weeks later some roots will<br />
form. A first small new leaf will eventually appear from the<br />
base of the cutting and/or the cutting will make a small<br />
dormant tuber before it will wither. If the leaf cutting withers<br />
and no new leaf has been created, carefully check its base.<br />
In most cases it will have made a small dormant tuber.<br />
Cultivation for these newly formed tubers is the same as<br />
mentioned above for growing seedlings.<br />
In general, <strong>Amorphophallus</strong> species are rather trouble-free and easy-going in cultivation,<br />
provided that some basic cultivation guidelines are complied.<br />
However, dormant tubers are sometimes prone to fungal or bacterial infections and rotting,<br />
especially in very moist soils and at rather low temperatures at the same time. If an<br />
infection is detected (parts of the tuber start to get slimy or mashy), an effort of rescue may<br />
be started by generously cutting away all infected parts. <strong>The</strong> rest of the tuber shall hence<br />
be dusted with some power fungicide and kept absolutely dry for several days before<br />
being repotted in a rather dry soil. In general and as a prevention measure against fungal<br />
infections, older plants should be kept rather dry during dormancy - with the exception of<br />
several tropical species, which have to be kept slightly moist throughout the year to<br />
prevent the tuber from dessication. If dormant tubers are stored unpotted and completely<br />
dry, they are more easily infested by mealy bugs. Thus a slightly moistened soil should be<br />
maintained in all species except for African members, which prefer a dry soil during<br />
dormancy. Leaves may be infested rarely with aphis and more often with spider mites.<br />
This especially occurs in winter months and if plants are kept at rather low air humidity<br />
inside the house. Any available insecticide spray respectively systemic solution fomulated<br />
against aphids, spider mites and/or mealy bugs will work fine. Always refer to the<br />
directions of use as stated on the package. And before treating an entire collection with<br />
some new insectide, a first trial should be made on a single plant in order to check for any<br />
unwanted side-effects.<br />
Another, and much more serious pest are nematodes. <strong>The</strong>se are very small and almost<br />
transparent worms with some 2mm to 4 mm in length. All dormant tubers shall be repotted<br />
every other year and inspected for these insects. An infestation can be detected for small<br />
bumby warts on the outside of the tuber. <strong>The</strong>se warts will eventually implode and leave<br />
behind a small mashy crater. This wound may be the entry site for a secondary bacterial<br />
and/or fungal infection. Unfortunately there is no reliable nematocide available for an use<br />
in a private collection. If such an infestation is detected, the best thing would be to throw<br />
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the entire pot (including the plant and the soil) away. For the rarer and hence pricier<br />
species, you may want to try to rescue the plant. However, an infested plant has to be kept<br />
in quarantine at all times and its pot should never get into contact with the open ground or<br />
a growing bench with other pots. Thoroughly wash the tuber removing any adherent soil. It<br />
may help to soak the tuber in some undiluted hypochlorite solution (standard bleach, wear<br />
some protective hand gloves for handling!) for some three minutes in order to remove<br />
some bacteria, fungal spores and nematodes from the tuber surface. Afterwards rinse the<br />
tuber thoroughly again with water to remove any remaining solution and let it dry a couple<br />
of days. After that repot the tuber using a new pot and fresh soil. Do not try to sterilize the<br />
old pot, but throw it away under all circumstances. Tuber offsets will inevitably be infested<br />
as well and these do not serve to regain a nematode-free stock. However, seeds, leaf<br />
bulbils and leaf cuttings are usually nematode-free, and this is the second-best option to<br />
rescue an infested plant. After successful propagation the mother plant shall be discarted<br />
as soon as possible.<br />
Plant & <strong>Seed</strong> Source<br />
An extensive list of <strong>Amorphophallus</strong> and other aroid species is available here:<br />
http://www.rareplants.de/shop<br />
http://www.plantdelights.com<br />
At present we are propagating a very good<br />
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the following genera: <strong>Amorphophallus</strong>,<br />
Anchomanes, Ariopsis, Arisaema, Arum,<br />
Biarum, Colletogyne, Dracontium,<br />
Dracunculus, Gorgonidium, Helicodiceros,<br />
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Pseudohydrosme, Spathantheum,<br />
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Plant Delights <strong>Nursery</strong> is the best source of<br />
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You can buy a wide variety of aroids such as<br />
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Ears, Pinellia, Philodendron, Sauromatum<br />
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