Inventory of the flora of Big Bend National Park Brewster County ...

INVENTORY OF THE FLORA OF 

BIG BEND NATIONAL PARK 

BREWSTER COUNTY, TEXAS 

[FIRST WORKING DRAFT] 

Compiled by: 

Richard D. Worthington, Ph.D. 

Floristic Inventories of the Southwest Program 

P. O. Box 13331 

El Paso, TX 79913 

El Paso, Texas 

MAY, 2001

INVENTORY OF THE FLORA OF 

BIG BEND NATIONAL PARK 

Brewster County, Texas 

LICHENS 

One can consult Bennett and Wetmore (2005) for an introduction to the National Parks database of 

lichens reported from the North American National Parks. They report 39 references and 277 taxa from 

BBNP. They estimate that 91-99 % of the lichen species are now known. 

Buellia spuria (Schaer.) Anzi 

[Bungartz & Nash, 2004] 

[Brewster Co.: BBNP, Wetmore 19248 (MIN)] 

Buellia subaethalea de Lesd. 

[Bungartz, Elix & Nash, 20040 

Brewster Co.: BBNP, Nash 10537b (ASU)] 

Caloplaca brouardii (B. de Lesd.) Zahlbr. 

[Wetmore & Karnefelt, 1998] 

[Brewster Co.: BBNP, Boot Canyon, Wetmore 40954 (MIN)] 

Caloplaca eygyra (Tuck.) Zahlbr. 


[Brewster Co.: BBNP, Blue Creek Canyon, Wetmore 19484 (CANL; MIN)] 

Caloplaca galactophylla (Tuck.) Zahlbr. 


[Brewster Co.: BBNP, Dagger Flats, Wetmore 19295 (MIN)] 

Caloplaca microphyllina (Tuck.) Hasse 

[Wetmore, 2004] 

[Brewster Co.: BBNP, South Rim Trail to Emory Peak, Anderson & shushan S18788 (BM)] 

Caloplaca persimilis Wetmore 


[Brewster Co.: BBNP, East Rim, Wetmore 18346 (ASU, FH, LSU, MIN, WIS)] 

Caloplaca saxicola (Hoffm.) Nordin 


[Brewster Co.: BBNP, N side of Chisos Mts., Nash 15032 (ASU; LSU; MIN)] 

Caloplaca sonorae Wetmore 


[Brewster Co.: BBNP, Emory Peak, Wetmore 40628 (MIN)] 

Caloplaca texana Wetmore & Karnefelt 


[Brewster Co.: BBNP, Noot Canyon, Wetmore 18277 (ALTA; COLO; MICH; MIN); 

Casa Grande, Wetmore 40893 (MIN); Emory Peak, 6000 ft., 11 Apr 1980, Wetmore 

40629 (Type: MIN; Isotypes: ASU; CANL)] 

Catapyrenium “lachneum” 

[Thomson, 1987, mapped] 

Note: True C. lachneum does not occur in the area. All material identified under that name has 

to be reviewed following the O. Breuss monographic work. 

Catapyrenium plumbeum (de Lesd.) J. W. Thomson 

[Dermatocarpon plumbeum (de Lesd.) Zahlbr.; Endopyrenium plumbeum de Lesd.] 


Catapyrenium rufescens (Ach.) Breuss 

[Breuss & McCune, 1994]

[Brewster Co.: Wetmore 18266 (LD)] 

Catapyrenium tuckermanii (Rav. ex Mont.) J. W. Thomson 


Dermatocarpon moulinsii (Mont.) Zahlbr. 

[Steyermark & Moore, 1933] 

[Brewster Co.: Chisos Mts., Moore & Steyermark] 

Dimelaena oreina (Ach.) Norman 

[Rinodina oreina (Ach.) Wain.] 



Flavoparmelia caperata (L.) Hale 

[Bennett & Wetmore, 2005] 

Haematomma subpuniceum (Mull. Arg.) de Lesd. 

[Culberson, 1963] 

[Brewster Co.: Emory Peak, 1900-2100 m, Anderson & Shushan (DUKE)] 

Heppia lutosa (Ach.) Nyl. 


[Brewster Co.: Lost Mine Trail from Panther Gap, Wetmore 14265B; Hot Springs near Rio 

Grande Village, Wetmore 18189, 18193; Old Mine Road near Rio Grande Village, Wetmore 

18377; Pulliam Ridge, Wetmore 18203A] 

Leptogium denticulatum Tuck. 

[Sierk, 1964] 

[Brewster Co.: S rim trail to Emory Peak, 1959, Anderson & Shushan (COLO)] 

Leptogium furfuraceum (Harm.) Sierk 

[Sierk, 1964] 

[Brewster Co.: S rim trail to Emory Peak, 1959, Anderson & Shushan 18806 (ABSL, COLO, 

WIS, FH)] 

Leptogium plicicatile (Ach.) Leighton 

[Collema plicatile Ach.] 



Lichinella cribellifera (Nyl.) P. Moreno & Egea 

[Schultz, 2005] 

Lichinella iodopulchra (Croz.) P. Moreno & Egea 


Lichinella minnesotensis (Fink) Essl. In Essl. & Egan 


Lichinella nigritella (Lettau) P. Moreno & Egea 


Neocatapyrenium disparatum Breuss 

[Breuss, 2005] 

[Brewster Co.: BBNP, Chisos Mts., Boot Canyon, Wetmore 18258 (MIN, type)] 

Peltigera collina (Ach.) Schrader 

[Peltigera scutata (Dickson) Duby] 

[Thomson, 1950] 

[Records: Boot Spring, Chisos Mts., 1937, Warnock (MO)] 

Peltigera rufescens (Weiss.) Humb. 

[Peltigera canina (L.) Willd. var. rufescens (Weiss.) Mudd] 


[Records: Chisos Mts., Boot Spring, 1937, Warnock (MO)] 

Peltula euploca (Ach.) Poelt 


[Brewster Co.: South Rim Trail to Emory Peak, Anderson & Shushan S-18693 (COLO); 

Boulder Meadows, Wetmore 18492; Grapevine Hills NW of Headquarters, Wetmore 18287A, 

18293A; Green Gulch, Wetmore 18437B; Pine Canyon, Wetmore 18422] 

Peltula obscurans (Nyl.) Gyelnik

var. obscurans 


[Brewster Co.: Green Gulch, Wetmore 18445; 2 rd. mi. from hwy. on rd. to Grapevine 

Hills, Wetmore 18294A] 

var. deserticola (Zahlbr.) Wetmore 


[Brewster Co.: below Burro Mesa pouroff, Wetmore 18312B; Dagger Flats, Wetmore 18208; 

2 mi. W jct. rd. to Castolon, Wetmore 18302; Grapevine Hills NW of Headquarters, Wetmore 

18288; 2 mi. from hwy. on rd. to Grapevine Hills, Wetmore 18294B; Green Gulch, Wetmore 

18430; Hot Springs Road near Rio Grande Village, Wetmore 18192, 18187; 4 mi. W jct. main 

road and road to Basin Campground, Wetmore 18295A; Old Mine Road near Rio Grande 

Village, Wetmore 18379; 7 mi. W jct. jct. rd. to Basin Campground, Wetmore 18296; W of jct. 

Going to Santa Elena Canyon along TX 170, Wetmore 14304, 14307 

var. hassei (Zahlbr.) Wetmore 


[Brewster Co.: Lost Mine Trail from Panther Pass, Wetmore 14265A; Pulliam Ridge, Wetmore 

18203B] 

Peltula omphaliza (Nyl.) Wetmore 


[Brewster Co.: below Burro Mesa pouroff, Wetmore 18312; 4 mi. W jct. main rd. and rd. to 

Basin Campground, Wetmore 18295C] 

Peltula patellata (Bagl.) Swainscow & Krog 

[Heppia polyspora Tuck.; Peltula polyspora (Tuck.) Wetmore] 


[Brewster Co.: Green Gulch, Wetmore 18443; Old Mine Road near Rio Grande Village, Wetmore 

18378] 

Peltula placodizans (Zahlbr.) Wetmore 


[Brewster Co.: across from Basin Campground, Wetmore 14252A; Boulder Meadows, Wetmore 

18411; Grapevine Hills, Wetmore 18293B; Green Gulch, Wetmore 18436, 18437A] 

Peltula richardsii (Herre) Wetmore 


[Brewster Co.: 2 mi. W jct. rd. to Castolon, Wetmore 18298; Hot Springs near Rio Grande 

Village, Wetmore 18185; Old Mine Road near Rio Grande Village, Wetmore 18380; W of jct. 

Going to Santa Elena Canyon along TX 170, Wetmore 14305] 

Physcia aipolia (Ehrh. ex Humb.) Furnr. 

[Bennett & Wetmore, 2005] 

Physcia stellaris (L.) Nyl. 



Pseudevernia intensa (Nyl.) Hale & Culb. 

[Parmelia furfuracea (L.) Ach.] 

[Hale, 1955, mapped] 

Psora crenata (Taylor) Reinke 

[Steyermark & Moore, 1933; Timdal, 1986, mapped] 


Psora decipiens (Hedwig) Hoffm. 



Psora globifera (Ach.) A. Massal. 



Psora pseudorussellii Timdal 

[Timdal, 1986, mapped] 

Psora russellii (Tuck.) A. Schneider 

[Timdal, 1986, mapped]

Ramalina unifolia Thomson 


[Brewster Co.: BBNP, Casa Grande, Wetmore 19520 (MIN)] 

Rhizoplaca chrysoleuca (Sm.) Zopf 

[McCune, 1987, mapped] 

Rinodina intermedia Bagl. 

[Mayrhofer, Sheard, Grassler & Elix, 2001] 

[Brewster Co.: BBNP, 7000 ft., Anderson & Shushan S18791 (COLO)] 

Staurothele areolata (Ach.) Lett. 


[Brewster Co.: BBNP, Emory Peak, Anderson & Shushan 18712 (COLO)] 

Usnea amblyoclada (Mull. Arg.) Zahlbr. 

[Clerc & Herrera-Campos, 1997] 

[Brewster Co., BBNP, Wetmore 40874] 

Xanthoria parietina (L.) Th. Fr. 



AYTONIACEAE 

HEPATOPHYTA 

Plagiochasma rupestre (Forst.) Steph. 

[Schuster, 1992] 

[Brewster Co.: near Boot Spring, Chisos Mts., BBNP, RMS 81-1247] 

Plagiochasma wrightii Sulliv. 

[Steyermark & Moore, 1933; Schuster, 1992] 

[Brewster Co.: Chisos Mts., Moore & Steyermark 3033; trail to South Rim, 

Emory Peak and to Boot Spring, Chisos Mts., BBNP, Shushan B-2885, RMS 

81-1240, 81-1247] 

Reboulia hemispherica (L.) Raddi ssp. acrogyna (Schust.) Schust. 

[Asterella bolanderi (Aust.) Underw. ssp. acrogyna Schust.] 

[Schuster, 1985, 1992] 

[Brewster Co.: Chisos Mts., BBNP, on track from Boot Spring to South Rim, 

6500-7000 ft., RMS 81-245 (Type)] 

CLEVEACEAE 

Athalamia pygmaea Schust.

CODONIACEAE 

[Schuster, 1985, 1992] 

[Brewster Co.: Chisos Mts., BBNP, near Boot Spring, 24 Dec. 1981, RMS 81-1251a 

(Type)] 

Fossombronia brasiliensis Steph. 


[Brewster Co.: Chisos Mts., Moore & Steyermark 3372] 

JUBULACEAE 

Frullania riparia Hampe ex Lehm 

[Mueller & Magill, 1979] 

[Brewster Co.: BBNP, Chisos Mts., Lost Mine Peak, Magill 1241, 1248, 

1275, 1776, 1792 (SRSC)] 

Frullania squarrosa (Reinw. Et al.) Dum. 


[Brewster Co.: BBNP, Chisos Mts., Emory Peak, Magill 1874 (SRSC)] 

RICCIACEAE 

Riccia andina K. Mull. ssp. chinophora Schust. 

[Schuster, 1992, 1992] 

[Brewster Co.: Chisos Mts., BBNP, N or NW of Boot Spring on slope of 

Emory Peak, 6500 ft., 24 Dec 1981, RMS 81-1241 (Type)] 

Riccia lamellosa Raddi 

[Schuster, 1985, 1992] 

[Brewster Co., Chisos Mts., BBNP, near Boot Spring, 24 Dec 1981, Schuster (growing 

with his type of Athalamia pygmaea Schust.)] 

Note: It is not clear if this collection reported by Schuster as R. lamellosa was not later 

determined to be R. andina and described above as a new subspecies. 

Riccia macallisteri M. A. Howe 


[Records: Brewster Co.: BBNP, rock outcrop E of Old Ore Rd., 0.8 km W of Tunnel, 

Magill 828 (SRSC)] 

Riccia sorocarpa Bisch. 

[Schuster, 1992] 

[Brewster Co.: Chisos Mts., BBNP, on track from Boot Spring to South Rim, 6500- 

7000 ft., RMS 81-245 (with type coll. of Reboulia hemispherica acrogyna)] 

TARGIONIACEAE 

Note: Mueller & Magill (1979) report Ricciocarpus nutans (L.) Corda from 8 km W of 

Study Butte which indicates it is expected in the park. 

Targionia heterophylla L. 

[Mueller & Magill, 1979; Schuster, 1992] 

[Records: Brewster Co.: BBNP, Chisos Mts., Boulder Meadow, on soil at base of 

boulders, Magill 1708 (SRSC); Chisos Mts., near Boot Spring, 24 Dec 1981, Schuster]

FABRONIACEAE 

BRYOPHYTA 

Fabronia ciliaris (Brid.) Brid. var. ciliaris 

[Fabronia octoblepharis (Schleich.) Schwaegr.] 



FUNARIACEAE 

Funaria hygrometrica Hedw. 



HEDWIGIACEAE 

Hedwigia ciliata (Hedw.) P. Beauv. 

[Hedwigia albicans (Web.) Lindb.] 


[Brewster Co.: Chisos Mts., Moore & Steyermark 3174]

HYPNACEAE 

Homomallium mexicanum Card. 



LESKEACEAE 

Lindbergia brachyptera (Mitt.) Kindb. 

[Crum, 1956] 

[Brewster Co.: Chisos Mts., Whitehouse 24662, 24664; BBNP, Whitehouse 19548] 

POTTIACEAE 

Leptodontium flexifolium (With.) Hampe ex Lindb. 

[Janssens & Zander, 1980, mapped] 

Scopelophila cataractae (Mitt.) Broth. 

[Reese, 1989] 

[Brewster Co.: BBNP, Burro Mesa in Apache Canyon near its head (29 Deg 17’N, 

103 Deg 23’W), Reese 17344 (BUF; DUKE; LAF; MICH; MO; NY; US)] 

Syntrichia chisosa (Magill, Delgadillo and Stark) Zand. 

[Tortula chisosa Magill, Delgadillo and Stark] 

[Brewster Co.: Chisos Mts., canyon below Lost Mine Trail, W side Lost Mine Peak, 

1600 m, 19 Jun 1973, Magill 1341 (Type: MO)] 

Syntrichia obtusissima (C. Mull.) Zand. 

[Tortula obtusissima (C. Mull.) Mitt.] 



Tortella humilis (Hedw.) Jenn. 

[Tortella caespitosa (Schwaegr.) Limpr.] 



Weissia controversa Hedw. 

[Reese, 1988.] 

[Brewster Co.: BBNP, Chisos Mts., trail to Emory Peak, Magill 2015] 

PTYCHOMITRIACEAE 

Ptychomitrium sinense (Mitt.) Jaeg. 

[Ptychomitrium leibergii Best.] 

[Cao & Vitt, 1994, mapped] 

LYCOPODIOPHYTA 

SELAGINELLACEAE (Spikemoss Family) 

Selaginella arizonica Maxon Arizona Spikemoss 

[Tryon, 1955, mapped] 

[Brewster Co.: unnamed drainage to N of Chisos Basin Campground and

E of Oak Creek, BBNP, 29 DEG 17’N, 103 DEG 18’40”W, 5400 ft., 16 Mar 

1983, Yatskievych (UTEP)] 

Selaginella densa Rydb. var. densa 


Selaginella lepidophylla (Hook. & Greville) Spring. in Resurrection Plant 

Martius et al. 

[Sperry, 1938; Wells, 1965; Eickmeier, 1988] 

[Brewster Co.: Dead Horse Mts. (Wells, 1965); Hot Springs (Sperry, 

1938)] 

Selaginella mutica D. C. Eaton ex L. Underwood var. 

limitanea Weatherby 

[Sperry, 1938; Tryon, 1955] 

[Brewster Co.: Pulliam Canyon, Pine Canyon and S slope Pulliam Bluff 

(Sperry, 1938); ridge S of Emory Peak, Chisos Mts., 2300 m, 22 Jun 1931, 

Moore & Steyermark (GH, MO, US)] 

Selaginella x neomexicana Maxon 

Selaginella peruviana (J. Milde) Hieron. Peruvian Spikemoss 

[Sperry, 1938; Tryon, 1955, mapped] 

[Brewster Co.: Pine Canyon and Oak Creek Canyon near Window 

(Sperry, 1938)] 

Selaginella pilifera A. Braun Resurrection Plant 

[Sperry, 1938] 

[Brewster Co.: Pulliam Canyon (Sperry, 1938)] 

Selaginella rupincola L. Underwood 

Selaginella underwoodii Hieron. Underwood’s Spikemoss 

[Brewster Co.: Maple Canyon, BBNP, 29 DEG 17’40”N, 103 DEG 

17’20”W, 5900 ft., 12 Mar 1985, Yatskievych (UTEP)] 

Selaginella viridissima Weatherby Slender Spikemoss 

[Tryon, 1955; Valdespino, 1993] 

[Brewster Co.: near waterfall at upper end of trail in Pine Canyon, BBNP, 

29 DEG 16’N, 103 DEG 15’W, 5600 ft., 17 Mar 1983, Yatskievych (UTEP); 

Chisos Mts., 10 Jul 1944, Cory 44831 (GH, US)] 

Selaginells wrightii Hieron Wright’s Spikemoss 


EQUISETACEAE (Horsetail Family) 

EQUISETOPHYTA 

Equisetum x ferrissi Clute 

Note: This is a hybrid between the next two species. 

Equisetum hyemale L. ssp. affine (Engelm.) Calder & Taylor 

Equisetum laevigatum A. Braun Smooth Scouring Rush 

ASPLENIACEAE (Spleenwort Family) 

POLYPODIOPHYTA 

Asplenium resiliens Kunze Black-stemmed Spleenwort 

[Palmer, 1930; Sperry, 1998] 

[Records: Pulliam Canyon, S slope Casa Grande, Green Gulch and Boot 

Spring (Sperry, 1938); Boot Spring, 25 May 1928, Palmer 34183]

AZOLLACEAE (Azolla Family) 

Azolla mexicana C. Presl. 

DENNSTAEDTIACEAE 

Pteridium aquilinum (L.) Kuhn var. pubescens L. Underwood Western Bracken 

DRYOPTERIDACEAE (Wood Fern Family) 

Cystopteris reevesiana Lellinger Southwestern Brittle Fern 

Phanerophlebia umbonata L. Underwood 

[Sperry, 1938; Yatskievych, 1993, 1996] 

[Brewster Co.: Pulliam Canyon and slopes of Casa Grande (Sperry, 1938); Chisos Mts., 

Basin, Maple Canyon, 12 May 1985, Yatskievych & McCrary 85-05; side of Casa Grande, 

Basin in Chisos Mts., Warnock 178 (ARIZ; GH; TEX-LL)] 

Woodsia neomexicana Windham New Mexico Cliff Fern 


[Brewster Co.: Pine Canyon (Sperry, 1938)] 

Woodsia phillipsii Windham Phillip’s Cliff Fern 

[Windham, 1993] 

[Records: trail from Boot Spring to South Rim, Warnock 1105 (TEX-LL; US)] 

Woodsia plummerae Lemmon Plummer’s Cliff Fern 

Note: Steyermark and Moore (1933) report Woodsia mexicana Fee (no. 3167) from 

the Chisos Mts. but the collection is most likely now annotated as one of the above. 

MARSILEACEAE (Water-clover Family) 

Marsilea vestita Hook. & Greville 

OPHIOGLOSSACEAE (Adder’s-tongue Family) 

Ophioglossum engelmannii Prantl Limestone Adder’s-tongue 

POLYPODIACEAE (Polypody Family) 

Pleopeltis riograndensis (T. Wendt) Andrews & Windham in Rio Grande Scaly Polypody 

Windham 

[Polypodium thysanolepis A. Braun ex Klotzsch var. riograndense T. Wendt] 


[Brewster Co.: Boot Canyon (Sperry, 1938)] 

PTERIDACEAE (Maidenhair Fern Family) 

Adiantum capillus-veneris L. Venus’s-hair Fern; Southern Maidenhair 

[Adiantum modestum Underw.] 


[Records: lower Oak Creek Canyon (Sperry, 1998); Oak Canyon, 25 May 1928, Palmer 

34122] 

Argyrochosma limitanea (Maxon) Windham ssp. mexicana 

(Maxon) Windham 

[Notholaena limitanea Maxon ssp. mexicana Windham] 

[Sperry, 1938; Correll, 1949]

[Brewster Co.: Willow Creek Canyon (Sperry, 1938)] 

Argyrochosma microphylla (Mett. ex Kuhn) Windham Littleleaf Cloakfern 

[Notholaena parvifolia Tryon; Pellaea microphylla Mett. ex Kuhn] 

[Steyermark & Moore, 1933; Sperry, 1938; Wells, 1965] 

[Brewster Co.: Dead Horse Mts. (Wells, 1965); Chisos Mts. (Sperry, 1938)] 

Astrolepis cochisensis (Goodd.) Benham & Windham ssp. Cochise Scaly Cloakfern 

cochisensis 

[Notholaena cochisensis Goodd.; Notholaena sinuata 

(Lagasca ex Swartz) Kaulfuss var. cochisensis (Goodd.) 

Weatherby] 

Astrolepis integerrima (Hook.) Benham & Windham Wavyleaf Cloakfern 

[Notholaena sinuata (Lagasca ex Swartz) Kaulfuss var. 

integerrima (Hook.) Nickel] 

[Palmer, 1930] 

[Records: Laguna or Green Gulch, 25 May 1928, Palmer 34167] 

Astrolepis sinuata (Lagasca ex Swartz) Benham & Windham Bulb Lipfern 

[Notholaena sinuata (Lag. ex Swartz) Kaulfuss] 


[Records: Juniper Canyon, 23 May 1928, Palmer 34100] 

Note: Subspecies sinuata and mexicana are mapped by Benham and Windham (1993) 

as occurring in the area. 

Astrolepis windhamii Benham 

Bommeria hispida (Kuhn) Underwood Hairy Bommeria 

[Sperry, 1938; Haufler, 1979] 

[Records: Chisos Mts. (Sperry, 1938); BBNP, Brenckle 51046 (US); Chisos Mts., 

Rossbach 4887 (US)] 

Cheilanthes aemula Maxon Texas Lipfern 

Chaelanthes alabamensis (Buckley) Kunze Alabama Lipfern 

[Palmer, 1930; Sperry, 1938; Wells, 1965] 

[Records: Dead Horse Mts. (Wells, 1965); Pulliam Canyon and S slope 

of Pulliam Bluff (Sperry, 1938); unnamed drainage to N of Chisos Basin Campground 

and E of Oak Creek, BBNP, 29 DEG 17’N, 103 DEG 18’40”W, 5400 ft., 16 Mar 1983, 

Yatskievych (UTEP); Boot Spring, 25 May 1928, Palmer 34134, 34147] 

Cheilanthes bonariensis (Willdenow) Proctor Bonaire Lipfern 


[Records: Mt. Emory and S slope Pulliam Bluff (Sperry, 1938); Juniper Canyon, 

23 May 1928, Palmer 34110] 

Cheilanthes eatoni Baker in Hook. & Baker Eaton’s Lipfern 

[Cheilanthes castanea Maxon] 

[Palmer, 1930; Steyermark & Moore, 1933; Sperry, 1938; Correll, 1949; McVaugh, 1952, 

mapped; Jones & Jones, 1993] 

[Records: Pinnacle’s trail towards Boot Spring (Jones & Jones, 1993); Pine 

Canyon, lower Oak Canyon and between Boot Spring and South Rim (Sperry, 1938); 

Moore & Steyermark 3191; Juniper Canyon, 23 May 1928, Palmer 34104] 

Cheilanthes feei T. Moore Slender Lipfern 


[Records: SE of Basin Campground in unnamed ravine on N side of Casa 

Grande, BBNP, 29 DEG 16’30”N, 103 DEG 17’30”W, 6400 ft., 14 Mar 1983, 

Yatskievych (UTEP); Boot Spring, 25 May 1928, Palmer 34187] 

Cheilanthes fendleri Hook. Fendler’s Lipfern 

Cheilanthes horridula Maxon Rough Lipfern; Prickly Lipfern 

Cheilanthes kaulfussii Kunze Glandular Lipfern 


[Records: Chisos Mts. (Sperry, 1938)] 

Cheilanthes lendigera (Cavanilles) Swartz Beaded Lipfern 

[Maxon, 1939; Correll, 1949]

[Records: Crown Mtn., Chisos Mts., 23 Aug 1937, Warnock] 

Cheilanthes lindheimeri Hook. Fairy Swords 


[Records: Chisos Mts. (Sperry, 1938); Oak Canyon, 24 May 1928, Palmer 34123; 

near Boot Spring, 25 May 1928, Palmer 34166] 

Cheilanthes tomentosa Link Woolly Lipfern 


[Records: Pulliam Canyon, S slope Pulliam Bluff, Boot Canyon and 

Pine Canyon (Sperry, 1938); Oak Canyon, 25 May 1928, Palmer 34085, 34087] 

Cheilanthes villosa Davenport ex Maxon Villous Lipfern 

Cheilanthes wootonii Maxon Wooton’s Lipfern 

Cheilanthes wrightii Hook. Wright’s Lipfern 



Notholaena aliena Maxon 

[Records: Burro Mesa pouroff, 29 DEG 13’N, 103 DEG 24’W, 3550 ft., 

10 Mar 1985, Yatskievych (UTEP)] 

Notholaena aschenboroniana Klotzsch 

Notholaena copelandii C. C. Hall 

Notholaena grayi Davenport ssp. grayi Gray’s Cloakfern 

Notholaena greggii (Mett. ex Kuhn) Maxon 

[Steyermark & Moore, 1933; Correll, 1949; Windham, 1993] 

[Records: atop tunnel just W Rio Grande Village on Rd. from Panther 

Jct. to Rio Grande Village, 29 DEG 11’N, 102 DEG 58’W, 2050 ft., 14 Mar 

1985, Yatskievych (UTEP) Boquillas Canyon, Moore & Steyermark 3348 (MO)] 

Notholaena nealleyi Sexton ex J. M. Coulter 

Notholaena neglecta Maxon 

Notholaena standleyi Maxon Star Cloakfern 

[Palmer, 1930; Sperry, 1938; Seigler & Wollenweber, 1983] 

[Records: S slope of Pulliam Bluff (Sperry, 1938); Juniper Canyon, 23 May 1928, 

Palmer 34105; Oak Canyon, 24 May 1928, Palmer 34151; BBNP, Correll & Correll 

30599 (TEX-LL)] 

Pellaea atropurpurea (L.) Hook. Purple Cliffbrake 

[Palmer, 1930; Sperry, 1938; Tryon, 1957; Jones & Jones, 1993] 

[Records: Pinnacle’s trail towards Boot Springs (Jones & Jones, 1993); 

S slope of Pulliam Bluff (Sperry, 1938); Boot Spring, 25 May 1928, Palmer 34186] 

Pellaea cordifolia (Sesse & Mocino) A. R.Smith 

[Pellaea cordata (Cav.) Sm.; Pellaea sagittata (Cav.) Link var. cordata (Cav.) A. Tryon] 

[Sperry, 1938; Tryon, 1957] 

[Brewster Co.: Pinnnacle Mts. (Sperry, 1938)] 

Pellaea intermedia Mett. ex Kuhn Creeping Cliffbrake 

[Palmer, 1930; Sperry, 1938; Tryon, 1957, 1968, mapped; Jones & Jones, 1993] 


Divide, Pulliam Canyon, S slope of Casa Grande and Boot Canyon (Sperry, 1938); 

Juniper Canyon, 22 May 1928, Palmer 34078] 

Pellaea ternifolia (Cav.) Link 

[Sperry, 1938; Correll, 1949; Tryon, 1957; Windham, 1993] 

ssp. arizonica Windham 

[Records: Mt. Emory and Pulliam Canyon (Sperry, 1938); along south rim of Basin, Tryon & 

Tryon 5040 (ssp. arizonica: MEXU; UT)] 

Note: Subspecies ternifolia and arizonica are reported from the park. 

Pellaea wrightiana Hook. Wright’s Cliffbrake 

[Palmer, 1930; Perry, 1938; Tryon, 1957] 

[Records: NW slope Pulliam Bluff (Sperry, 1938); Oak Canyon, 25 May 1928, Palmer 

34131]

CUPRESSACEAE (Cypress Family) 

CONIFEROPHYTA (Conifers) 

Cupressus arizonica Greene Arizona Cypress 

[Steyermark & Moore, 1933; Sperry, 1938; Wolf, 1948; Correll, 1967; Powell, 1998; Karges 

& Zech, 2001] 

[Brewster Co.: “forested canyons in the Chisos Mts. (Correll reviewed 17 sheets); 

upper Boot Spring valley, Chisos Mts., 22 Jun 1931, Moore & Steyermark 3207 (MO, CAS); 

Boot Spring, Warnock & Hinckley 7138 (SRSC), Palmer 34177 (CAS); upper Juniper Canyon, 

15-18 Jul 1921, Ferris & Duncan 2725 (CAS)] 

Juniperus coahuilensis (Martinez) Gaussen ex R. P. Adams Roseberry 

[Juniperus erythrocarpa Cory] 

[Cory, 1936; Sperry, 1938; Correll, 1967, as J. monosperma; Zanoni, 1978; Adams & 

Zanoni, 1979; Adams & Kistler, 1991; Powell, 1998] 

[Brewster Co.: Chisos Mts. (see Correll, 1967, for a listing of records); Laguna in Chisos 

Mts., W base of Mt. Emory, 2 Dec 1933, Cory 7642 (Type of J. erythrocarpa: A; Isotype: 

MO)] 

Note: This species hybridizes with J. pinchotii in the Basin (Adams & Kistler, 1991). 

Juniperus deppeana Steudel var. deppeana Alligator Juniper 

[Sperry, 1938; Correll, 1967; Powell, 1998] 

[Brewster Co.: Chisos Mts. (numerous records listed in Correll, 1967)] 

Juniperus flaccida Schlecht. var. flaccida Drooping Juniper 

[Steyermark & Moore, 1933; Sperry, 1938; Correll, 1967; Jones & Jones, 1993; 

Adams, 1993; Powell, 1998] 

[Brewster Co.: upper slopes of the Chisos Mts. (see Correll, 1967, for numerous 

records); Panther Pass, BBNP, 26 Jun 1978, Freeman s.n. (UTEP); Chisos Mts., Moore & 

Steyermark 3330 (MO)] 

Juniperus pinchotii Sudworth Pinchot Juniper; Redberry Juniper 

[Juniperus texensis Van Melle] 

[Steyermark & Moore, 1933; Van Melle, 1952; Wells, 1965; Correll, 1967; Zanoni, 1978; 

Adams & Zanoni, 1979] 

[Brewster Co.: Chisos Mts. (a number of records are listed in Correll, 1967); BBNP, Brenchle 

51019 (Type of J. texensis: NY; Isotype: A)] 

Note: This species hybridizes with J. coahuilensis in the Basin (Adams & Kistler, 1991). 

PINACEAE (Pine Family) 

Pinus arizonica Engelm. var. stormiae Martinez 

[Powell, 1998] 

[Brewster Co.: Pine Canyon, N slope of Crown Mt., S slope Boot Canyon (Powell, 

1998)] 

Pinus cembroides Zuccarini Mexican Pinyon 

[Sperry, 1938; Correll, 1967; Jones & Jones, 1993; Powell, 1998] 

[Brewster Co.: Chisos Mts. (numerous records listed by Correll, 1967); Dead Horse 

Mts.(Powell, 1998); lower end Lost Mine Trail, BBNP, 26 Jun 1978, Freeman s.n. 

(UTEP)] 

Pinus ponderosa Dougl. ex Lawson var. scopulorum Engelm. in Rocky Mountain Ponderosa Pine 

S. Watson 


[Brewster Co.: Chisos Mts. (nine collections reviewed by Correll, 1967)] 

Note: Powell (1998) references communication from Bailey to the effect that this 

taxon does not occur in the Chisos Mts., the above collections being Pinus arizonica.

Pinus remota (Little) Bailey & Hawksworth Papershell Pinyon 


[Brewster Co.: Dead Horse Mts. and 0.5 mi. NW Sue Peak, BBNP (Powell, 1998)] 

Pseudotsuga menziesii (Mirbel) Franco var. glauca (Mayr) Franco Rocky Mountain Douglas-fir 


[Brewster Co.: Chisos Mts. (numerous records listed by Correll, 1967)] 

GNETOPHYTA (Gnetophytes) 

EPHEDRACEAE (Mormon-tea or Joint-fir Family) 

Ephedra antisyphilitica Berl. ex C. A. Mey Vine Ephedra 

[Sperry, 1938; Correll, 1967] 

[Brewster Co.: slopes of the Dead Horse Mts., 08 Jul 1937, Warnock 490] 

Note: The above record is not cited by Powell (1998). 

Ephedra nevadensis S. Wats. var. aspera (Engelm. ex S. Wats.) Boundary Ephedra; Popotillo 

L. Benson 

[Sperry, 1938; Wells, 1965; Correll, 1967] 

[Brewster Co.: (see Correll, 1967 for numerous records in BBNP)] 

Ephedra torreyana S. Wats. var. powelliorum Wendt 


[Brewster Co.: 6.9 mi. N Panther Junction and Tornillo Flats (Powell, 1998)] 

Ephedra trifurca Torr. ex S. Wats. Mexican-tea 

[Sperry, 1938; Correll, 1967] 

[Brewester Co.: 4 mi. N of Tornillo Creek, BBNP, rte. 51, 02 Apr 1959, Correll 

20654; 3 mi. SE Costolon, 29 May 1938, Cutler 1881, 1879; Hot Springs, BBNP, 

27 Mar 1948, Goodman & Waterfall 4623; near Solis Ranch and Boquillas, 20 Jul 

1937, Marsh 143; Chisos Mts., 27 Jun 1931, Moore & Steyermark 3287; W slope 

of the Dead Horse Mts., 02 Apr 1942, Sperry 2044, 2047; et al. in Correll, 1967] 

ACANTHACEAE (Acanthus Family) 

MAGNOLIOPHYTA (Flowering Plants) 

MAGNOLIOPSIDA (Dicots) 

Anisacanthus linearis (Hagen) Henrickson & Lott Dwarf Anisacanth 

[Anisacanthus insignis Gray] 

[Steyermark & Moore, 1933; Sperry, 1938; Wasshausen, 1967; Henrickson & Lott, 1982; 

Daniel, 1984; Powell, 1998] 

[Records: Chisos Mts., Painted Gap, Warnock 905 (ARIZ, SRSC, TEX-LL); 

“numerous localities in Big Bend Park and Chisos Mts., Paint Gap Hills, Oak and Willow 

Creek canyons, 4 mi. W of Government Springs, Basin, W of Lone Mt., Grapevine Hills 

(SRSC); lower Blue Creek Canyon, 1370 m, 23 Jun 1931, Moore & Steyermark 3202 (Type:

MO; Isotypes: GH; NY; UC); see Wasshausen (1967) for more records] 

Anisacanthus puberulus (Torr.) Henrickson & Lott Pinky Anisacanth 


[Brewster Co.: Chisos Mts. and Oak Creek near Cattail Falls trailhead (SRSC)] 

Carlowrightia arizonica Gray Arizona Carlowrightia 


[Brewster Co.: Chisos Mts., Blue Creek, head of Boquillas Canyon, and Hot Springs 

(SRSC); Sierra Quemada near Domenguez Spring, BBNP, Warnock & Wallmo 12304 

(TEX-LL)] 

Carlowrightia linearifolia (Torr.) Gray 

[Sperry, 1938; Wasshausen, 1967; Daniel, 1983, 1984] 

[Brewster Co.: 0.5-0.8 mi. W Chisos Basin Campground near Window Trail, Daniel 565 

(CAS, DUKE, ENCB, F, GH, MICH, MO, NY, TEX-LL, UC, US); Chisos Mts. (SRSC); 

Chisos Mts., lower Green Gulch, 30 Aug. 1936, Sperry 472 (US); Devil’s Backbone to Fisk 

Canyon, 01 Aug 1937, Warnock 1090; W side Ward Spring, BBNP, 18 Aug 1955,Warnock 

13092 (TEX-LL); near Chisos Mts., 12 Sep 1915, Young 188 (MO); Blue Creek Canyon, BBNP, 

Correll 33827 (TEX-LL, US); Chisos Mts., Nealley 436 (351) (F, US); 13 mi. SE Costolon, 

Powell & Powell 3025 (TEX-LL); 2 mi. N main road on Grapevine Spring Road, Semple 

(MO, US); Oak Creek Canyon, Chisos Mts., Sperry 462 (US); lower Oak Creek, Warnock 

1091 (GH); lower Willow Creek, Warnock 1091[sic] (TEX-LL); upper Heath Canyon, 

Warnock & Hinkley BG238 (TEX-LL)] 

Carlowrightia parvifolia Brandeg. 


[Brewster Co.,: Blue Creek Canyon, BBNP (SRSC)] 

Carlowrightia serpyllifolia Gray 

[Wasshausen, 1967; Powell, 1998; Daniel, 1983] 

[Brewster Co.: Chisos Mts. et al. (SRSC); Oak Canyon below the “Window,” Chisos 

Mts., 09 Nov. 1964, Correll & Correll 30621 (GH, TEX-LL, MO, BRIT); Cattail Canyon, 

Chisos Mts., 23 Sep 1938, Cory 30186, 30187 (A); 14 mi. E Costolon, S edge of Chisos Mts., 

12 May 1937, Cutler 1018 (MO), 05 Mar 1937, Cutler 733 (MO); et al. in Wasshausen, 1967; 

Heath Canyon, Correll 31572 (TEX-LL, US); pouroff of Burro Mesa, BBNP, Correll & 

Correll 35384 (GH, TEX-LL, MO, NCU, BRIT, UC); Cattail and Oak Canyons in Chisos 

Mts., Sperry 321 (TEX-LL); Pinnacles of Chisos Mts. at Fisk Canyon hut, Warnock 321 

(TEX-LL); Baldy Peak in Chisos Mts., Warnock 12990 (TEX-LL); Hot Springs, Warnock 

13926 (TEX-LL)] 

Dyschoriste decumbens (Gray) Kuntze Narrowleaf Dyschoriste 

[Brewster Co.: Basin at Saddle above Green Gulch, 15 Jul 1981, Freeman s.n. 

(UTEP)] 

Dyschoriste linearis (Torr. & Gray) Kuntze 


[Brewster Co.: Basin and Green Gulch (Sperry, 1938)] 

Justicia warnockii B. L. Turner Warnock Justicia 

[Wasshausen, 1967; Powell, 1998] 

[Brewster Co.: Chisos Mts. and Dead Horse Mts. (SRSC); near Mckinney Springs, 

Dead Horse Mts., 19 Jul 1952, Warnock 10786; Dead Man’s Cut, BBNP, 10 Aug 1955, 

Warnock 13063; 5 mi. from Ward Spring, Chisos Mts., 18 Aug 1955, Warnock 13007; 

Big Brushy Canyon of Dead Horse Mts., 25 Jun 1950, Warnock & Hinckley BG-137; 

above Tunnel W of Rio Grande Village, 20 Aug 1982, Van Devender s.n. (UTEP)] 

Ruellia parryi Gray Parry Ruellia 

[Steyermark & Moore, 1933; Sperry, 1938; Tharp & Barkley, 1949; Turner, 1991, mapped; 

Powell, 1998] 

[Records: 14 mi. E of Castolon, 7 Mar 1937, Cutler (GH); Oak Canyon, 24 May 1928, 

Palmer 34128 (NY); Chisos Mts., 26 May 1928, Palmer 34203 (A); 3 mi. W of Glenn 

Spring, 20 Aug 1939, Sperry 1553 (GH); Glenn Spring, 3 May 1937, Warnock (GH; TEX-LL); 

Santa Elena Canyon, Steyermark & Moore 3459 (MO)] 

Siphonoglossa pilosella (Nees) Torr.


[Brewster Co.: shrub areas N of Chisos Mts. (Sperry, 1938)] 

Stenandrium barbatum Torr. & Gray Shaggy Stenandrium 


[Brewster Co.: desert areas near Persimmon Gap (Sperry, 1938] 

ACERACEAE (Mple Family) 

Acer grandidentatum Nutt. var. grandidentatum Bigtooth Maple 

[Sperry, 1938; Jones & Jones, 1993; Powell, 1998] 

[Brewster Co.: Pinnacle’s trail towards Boot Springs (Jones & Jones, 1993); 

Boot and Pulliam Canyons (Sperry, 1938)] 

AMARANTHACEAE (Amaranthus Family) 

Alternanthera caracasana Kunth Matt Chaff-flower 

[Sperry, 1938; Reed, 1969] 

[Brewster Co.: Juniper Canyon, Chisos Mts., Jul 1935, Marsh 197; Chisos Mts., 

05 Aug 1931, Mueller s.n.] 

Amaranthus albus L. White Amaranth 

[Amaranthus pubescens (Uline & Bray) Rydb.] 

[Reed, 1969] 

[Brewster Co.: Chisos Mts., 27 Aug. 1915, Young s.n.; pinnacles, Chisos Mts., 22 Jul 

1937, Warnock 463; Chisos Pens between Todd Hill and Burro Mesa, 01 Aug 1955, 

Warnock 12764] 

Amaranthus arenicola I. M. Johnst. Torrey Amaranth 

[Amaranthus torreyi auct. non Gray] 

[Reed, 1969] 

[Brewster Co.: Terlingua Flats near Study Butte, BBNP, 22 Jul 1957, Correll & 

Johnston 18363; McKinney Springs, Dear Horse Mts., 19 Jul 1952, Warnock 10796; 

S of Ward Spring, Chisos Mts., 18 Aug 1955, Warnock 13010; Oak Creek Canyon, 

03 Aug 1955, Warnock 12811; Basin of Chisos Mts., 31 Aug. 1938, Warnock C647; 

Fresno Springs, Chisos Mts., 29 Jun 1937, Warnock 347; upper Green Gulch, 14 Sep 

1941, Sperry 1966; Baldy Peak, Basin of Chisos Mts., 06 Sep 1950, Warnock 9605; 

Onion Springs, Paint Gap Hills, 05 Aug 1955, Warnock 13083; lower Blue Canyon, 

07 Sep 1933, Wolff 4488; 1.5 mi. N Costolon, 28 Sep 1938, Park & Cory 30507; 

along river at Mariscal, 22 Jul 1937, Sperry 2070] 

Amaranthus blitoides S. Wats. Prostrate Pigweed 

[Sperry, 1938; Reed, 1969] 

[Brewster Co.: Chisos Mts., 06 Aug 1937, Marsh 182; Chisos Windmill, 19 Jun 1931, 

Tharp 8851] 

Amaranthus crassipes Schlecht. 

[Reed, 1969] 

[Brewster Co.: Chisos Mts., 20 Jul 1937, Marsh 128] 

Amaranthus fimbriatus (Torr.) Benth. ex S. Wats. Fringed Amaranth 

[Reed, 1969] 

[Brewster Co.: Onion Springs, BBNP, 05 Aug. 1955, Warnock 13083] 

Amaranthus hybridus L. Green Amaranth 

[Sperry, 1938; Reed, 1969] 

[Brewster Co.: Boot Springs, Chisos Mts., 26 Sep 1938, Parks & Cory 30374] 

Amaranthus palmeri S. Wats. Palmer Amaranth 

[Sperry, 1938; Reed, 1969] 

[Brewster Co.: Chisos Mts., 20 Jul 1937, Marsh 136; upper Green Gulch, 14 Sep 

1941, Sperry 1965; RR between Todd Hill and Baro Mesa, BBNP, 01 Aug 1955, 

Warnock 12782; Boquillas (Sperry, 1938)]

Amaranthus powellii S. Wats. ssp. powellii Powell Amaranth 

[Sperry, 1938; Reed, 1969; Costea et al., 2001] 

[Brewster Co.: Chisos Mts. State Park, Jul 1935, Marsh 89, 04 Aug 1931, Mueller 

7986 (US)] 

Amaranthus scleropoides Uline & Bray 


[Brewster Co.: Chisos Mts. (Sperry, 1938)] 

Amaranthus wrightii S. Wats. Wright Amaranth 

[Reed, 1969] 

[Brewster Co.: Boot Springs, 09 Sep 1933, Parks & Cory 7129; Boot Springs Area, 

08 Sep 1933, Wolff 4592; upper Blue Creek Canyon, 10 Sep 1933, Parks & Cory 

7300] 

Froelichia arizonica Thornb. ex Standl. Arizona Snakecotton 

[Sperry, 1938; Reed, 1969] 

[Brewster Co.: Green Gulch, Chisos Mts., 05 Aug 1946, Correll 13659; Oak 

Creek Canyon, 06 Sep 1940, Sperry 1763; Chisos Mts., 25 Jun 1937, Marsh 16] 

Froelichia floridana (Nutt.) Moq. Florida Snakecotton 

[Sperry, 1938; Reed, 1969] 

[Brewster Co.: Chisos Mts., Jul 1935, Marsh 92; Onion Springs, Paint Gap Hills, 

BBNP, 05 Aug 1955, Warnock 12972; Green Gulch, 14 Sep 1941, Sperry 1979] 

Note: Reed (1969) determined some material to be var. campestris (Small) Fern. 

Froelichia gracilis (Hook.) Moq. Slender Snakecotton 

[Froelichia braunii Standl.] 

[Reed, 1969] 

[Brewster Co.: south rim, Chisos Mts., 26 Sep 1938, Parks & Cory 30376; 1.5 mi. 

N of Costolon, 28 Sep 1938, Parks & Cory 30509] 

Froelichia interrupta (L.) Moq. Texas Snakecotton 

[Reed, 1969] 

[Brewster Co.: lower Blue Creek Canyon, Chisos Mts., 07 Sep 1933, Wolff 

4510] 

Gomphrena nitida Rothrock Pearly Globe-amaranth 

[Sperry, 1938; Reed, 1969] 

[Brewster Co.: Cattail Canyon, Chisos Mts., 23 Sep 1938, Cory s.n., 23 Sep 1938, 

Parks & Cory 30167, 30168; Divide (Sperry, 1938)] 

Gossypianthus lanuginosus (Poir.) Moq. 


[Brewster Co.: lower Willow Creek Canyon (Sperry, 1938)] 

Guilleminea densa (Willd. in Roem. et Schult.) Moq. var. Cottonflower 

aggregata Uline & Bray 

[Sperry, 1938; Reed, 1969; Henrickson, 1987] 

[Brewster Co.: Green Gulch, Chisos Mts., 04 Aug 1936, Sperry 1243 (US); more 

records in Reed (1969)] 

Iresine heterophylla Standl. Standley Bloodleaf 

[Sperry, 1938; Reed, 1969] 

[Brewster Co.: Pine Canyon, Chisos Mts., BBNP, 17 Jun 1963, Correll & Wasshausen 

27880, et al.; Blue Creek Canyon, 30 Sep 1933, Wolff 4679, et al.] 

Iresine leptoclada (Hoof. f.) Henrickson & Sandberg Bloodleaf 

[Dicrairus leptocladus Hook. f.] 

[Sperry, 1938; Reed, 1969; Henrickson & Sundberg, 1986; Powell, 1998] 

[Brewster Co.: Green Gulch in Chisos Mts., BBNP, 06 Sep 1950, Warnock 9617; et al. 

in Reed (1969); upper Juniper Spring (Sperry, 1938)] 

Tidestromia carnosa (Steyermark) I. M. Johnston Fleshy Tidestrom 

[Tidestromia lanuginosa (Nutt.) Standl. var. carnosa (Steyerm.) Cory] 

[Steyermark, 1932; Steyermark & Moore, 1933; Reed, 1969] 

[Brewster Co.: Blue Creek, 26 May 1928, Palmer 34196; Muskhog Spring, 

30 Jul 1938, Warnock C577; between Todd Hill and Burro Mesa, BBNP, 01 Aug

1955, Warnock 12802; Tornillo Creek, Aug 1883, Havard 104, 17 Apr 1967, 

correll & Rollins 23608; between Avary Canyon and Tornillo Pass, 06 Sep 1950, 

Warnock 9632; et al. in Reed (1969); gravelly flat 3 mi. E of Study Butte, 762 m, 

26 Jun 1931, Moore & Steyermark 3795 (GH – Type)] 

Tidestromia lanuginosa (Nutt.) Standl. Woolly Tidestrom 

[Sperry, 1938; Reed, 1969] 

[Brewster Co.: 2 mi. S Study Butte, BBNP, 28 Sep 1947, Warnock 7374; lower 

Blue Creek Canyon, 07 Sep 1933, Wilff 4525; mesa near Chisos Mts., 13 Sep 1916, 

Young s.n.; Burman’s Pasture, Chisos Mts., 01 Sep 1915, Young s.n.; Fresno Springs, 

Chisos Mts., 29 Jun 1937, Warnock 671; between Nugent and Glenn Springs, 29 Oct 

1939, Sperry 1511; et al. in Reed (1969); Pinnacle Mts. (Sperry, 1938)] 

Tidestromia suffruticosa (Torr.) Standl. Shrubby Tidestrom 

[Sperry, 1938; Reed, 1969; Powell, 1998] 

[Brewster Co.: Chisos Mts., 16 Jul 1937, Marsh 105; Tornillo Creek bed at Hot Springs, 

02 Dec 1937, Sperry 1598; Boquillas Canyon, 06 Aug 1937, Warnock 694; et al. in 

Reed (1969); Mesa de Anguila and Boquillas (Sperry, 1938)] 

ANACARDIACEAE (Sumac Family) 

Rhus microphylla Engelm. ex Gray Littleleaf Sumac; Desert Sumac 

[Sperry, 1938; Powell, 1998] 


Rhus trilobata Nutt. var. trilobata Fragrant Sumac; Squawbush 

[Barkley, 1937; Sperry, 1938; Powell, 1998] 

[Brewster Co.: Basin and Green Gulch (Sperry, 1938); upper Blue Creek Canyon, 

1520 m, 01 Jul 1931, Moore & Steyermark 3326 (CAS; MO); Chisos Mts., 19 Jun 1931, 

Mueller 8066 (MO)] 

Rhus virens Lindheimer ex Gray var. virens Evergreen Sumac; Tobacco 

[Sperry, 1938; Wells, 1965; Turner, 1996, mapped; Powell, 1998] 

[Brewster Co.: Basin and Green Gulch (Sperry, 1938); Sue Peak, Dead Horse Mts. 

9Wells, 1965)] 

Toxicodendron radicans (L.) O. Ktze. ssp. eximium (Greene) Gillis Poisonivy 

[Barkley, 1937; Powell, 1998] 

[Brewster Co.: Chisos Mts., Warnock 1212 (ARIZ; GH; TEX-LL); Chisos Mts., Willow 

Creek Canyon, 23 May 1928, Palmer 34076 (MO; A; NY) and Sperry 201 (US); Goode Place, 

Young s.n. (GH; TEX-LL); Oak Canyon, Cory 7010 (GH) and Mueller 8063 (F; TEX-LL); 

Oak Canyon, base of Ward Mtn., Gillis 5745 (MSC)] 

APIACEAE (Parsley Family) 

Aletes acaulis (Torr.) Coult. & Rose 


[Brewster Co.: upper slopes of Casa Grande (Sperry, 1938)] 

APOCYNACEAE (Dogbane Family] 

Apocynum androsaemifolium L. Spreading Dogbane 



Telosihonia hypoleuca (Benth.) Henr. Bottomwhite Rocktrumpet 

[Macrosiphonia hypoleuca (Benth.) Muell.-Arg.] 


[Brewster Co.: upper Chisos near the Laguna (Powell, 1998)] 

Telosiphonia macrosiphon (Torr.) Henr. Plateau Rocktrumpet 

[Macrosiphonia macrosiphon (Torr.) Heller]

[Sperry, 1938; Henrickson, 1996; Powell, 1998] 

[Brewster Co.: Pulliam Bluff, below Pine Canyon, upper Cattail Canyon and 

hill N of Chisos Mts. (Sperry, 1938); Chisos Mts., 05 Aug 1931, Muller 8123 (F; 

MICH; NY; TEX; US)] 

ARISTOLOCHCHIACEAE (Birthwort Family) 

Aristolochia wrightii Seem. Wright Dutchman’s Pipe 

[Aristolochia brevipes Benth.] 


[Brewster Co.: Pinnacle Mts., Oak Creek Canyon, Mesa de Anguila and 15 mi. S 

of Persimmon Gap (Sperry, 1938)] 

ASCLEPIADACEAE (Milkweed Family) 

Asclepias asperula (Dcne.) Wods. ssp. capricornu (Woods.) Woods. Spider Antelopehorn 

[Asclepiodora decumbens (Nutt.) Gray] 


[Brewster Co.: Basin and Panther Spring (Sperry, 1938)] 

Asclepias glaucescens Kunth 

[Asclepias elata Benth.] 

[Steyermark & Moore, 1933; Sperry, 1938] 

[Brewster Co.: Willow Creek Canyon (Sperry, 1938); head of Green Gulch, Chisos 

Mts., Moore & Steyermark 3417 (MO)] 

Asclepias nummularia Torr. Yerba de Cuervo 


[Brewster Co.: lower Green Gulch (Sperry, 1938)] 

Asclepias oenotheroides Cham. & Schlect. Hierba de Zizotes 

[Asclepias longicornu Benth.] 


[Brewster Co.: Tornillo Flats, lower Green Gulch and Croton Spring (Sperry, 

1938)] 

Asclepias subverticillata (Gray) Vail Poison Milkweed 

[Asclepias galioides auct. non Kunth] 


[Brewster Co.: Tornillo Creek (Sperry, 1938)] 

Asclepias texana Heller 

[Sperry, 1938, as A. perennis Walt.; Edwards & Wyatt, 1994; Wyatt et al., 1998] 

[Brewster Co.: Pulliam Canyon and lower Oak Creek Canyon (Sperry, 1938); Boot Spring, 

6900 ft. (GA); Boot Spring Trail, 5700 ft., Dec 1992 (GA); Laguna Meadow Trail, 6000 ft. (GA); 

Laguna Meadow, 6700 ft. (GA)] 

Cynanchum pringlei (Gray) Henrickson 

[Cynanchum barbigerum (Scheele) Shinners var. breviflorum Shinners] 

[Steyermark & Moore, 1933; Sperry, 1938; Shinners, 1964] 

[Brewster Co.: Chisos Mts., BBNP, granite peak in center of basin, 15-19 Jul 1952, 

Webster 4340 (BRIT) (Type of breviflorum); Basin, Oak Creek Canyon and lower 

Juniper Spring (Sperry, 1938); Chisos Mts., Moore & Steyermark 3365] 

Cynanchum racemosum (Jacq.) Jacq. var. unifarium (Scheele) E. Sundell 

[Rouliniella unifaria (Scheele) Vail] 


[Brewster Co.: Panther Canyon (Sperry, 1938); Chisos Mts., Moore & Steyermark 3398] 

Matelea reticulata (Engelm. ex Gray) Woods. 

[Vincetoxicum reticulatum (Engelm.) Heller; Gonolobus reticulatus 

Engelm. ex Gray] 

[Sperry, 1938]

[Brewster Co.: Pulliam Canyon and S slope Pulliam Bluff (Sperry, 1938)] 

Sarcostemma crispum Benth. Wavyleaf Twinevine 

[Funastrum crispum Benth.) Schlechter] 


[Brewster Co.: Chinese Wall (Sperry, 1938)] 

Sarcostemma cynanchoides Dcne. var. hartwegii (Vail) Shinners Arroyo Twinevine 

[Funastrum heterophyllum (Engelm.) Standl.] 


[Brewster Co.: Neville Spring and San Francisco Canyon (Sperry, 1938)] 

Sarcostemma torreyi (Gray) Woods. Soft Twinevine 

[Funastrum torreyi (Gray) Schltr.] 


[Brewster Co.: Basin, Green Gulch and lower Juniper Spring (Sperry, 1938); Basin 

at Saddle above Green Gulch, 15 Jul 1981, Freeman s.n. (UTEP); Chisos Mts., 

Moore & Steyermark 3234] 

ASTERACEAE (Sunflower Family) 

Acourtia nana (Gray) Reveal & King Desert Holly 

[Perezia nana Gray] 


[Brewster Co.: Basin (Sperry, 1938)] 

Acourtia runcinata (Lag. ex D. Don) B. L. Turner 


[Brewster Co.: Fresno Creek and Pinnacle Mts. (Sperry, 1938)] 

Acourtia wrightii (Gray) Reveal & King Brownfoot; Pink Perezia 


[Brewster Co.: Juniper Canyon and Chinese Wall (Sperry, 1938); Chisos Mts, Moore 

& Steyermark 3337] 

Ageratina herbaceum (Gray) King & H. E. Robins. 

[Sperry, 1938; Johnston, 1967] 

[Brewster Co.: Chisos Mts., 24 Aug 1931, Mueller 8177 (TEX-LL), 14 Aug 1941, 

Mueller 8182 (TEX-LL; BRIT); from Laguna to Boot Springs, 26 Aug 1937, Warnock 

1142 (TEX-LL); trail to south rim, Chisos Mts., 25 Aug 1937, Warnock 21723 (TEX-LL); 

near summit Mt. Emory, 07 Aug 1946, Correll 13700 (BRIT); Laguna of the Chisos Mts., 

10 Sep 1950, Warnock 9685 (TEX-LL; BRIT); trail into the Laguna, 10 Sep 1950, 

Warnock 9669 (TEX-LL; BRIT); Lost Mine Peak, 12 Sep 1961, Correll & Johnston 

24519 (TEX-LL)] 

Ageratina rothrockii (Gray) King & H. E. Robins. 

[Eupatorium rothrockii Gray] 


[Brewster Co.: basin-slopes of Mt. Emory, 09 Oct 1935, Sperry 542 (TEX-LL); trail to 

Chinese Wall, Chisos Mts., 01 Sep 1947, Warnock 6999 (BRIT); Lost Mine Peak, 12 

Sep 1961, Correll & Johnston 24511 (TEX-LL)] 

Ageratina wrightii (Gray) King & H. E. Robins. 

[Eupatorium wrightii Gray] 


[Brewster Co.: Chinese Wall (Sperry, 1938); Basin to Laguna (Powell, 1998)] 

Amblyolepis setigera DC. 

[Bierner, 1990] 

[Brewster Co.: Chisos Mts., Warnock 440 (TEX-LL)] 

Aphanostephus ramosissimus DC. var. humilis (Benth.) Turner & Plaind Dozedaisy 

Birdsong 

[Sperry, 1938; Turner, 1984, mapped] 

[Brewster Co.: Rio Grande near San Vicente Canyon (Sperry, 1938)]

Artemisia dracunculus L. 



Artemisia ludoviciana Nutt. Louisiana Sagebrush 

var. ludoviciana 


[Brewster Co.: Pine Canyon and Boot Spring to South Rim (Sperry, 1938)] 

var. mexicana (Willd. ex Spreng.) Keck 

[Sperry, 1938} 

[Brewster Co.: slopes of Mt. Emory (Sperry, 1938)] 

var. redolens (Gray) Keck. 



Baccharis havardii Gray Havard Baccharis 

[Sperry, 1938; Mahler & Waterfall, 1964; Powell, 1998] 

[Brewster Co.: Mt. Emory, Chisos Mts., 07 Aug 1946, Correll 13693 9BRIT); 

Chisos Mts., 14 Aug 1931, Mueller 8192 (GH; BRIT; TEX-LL); Boot Spring; above 

the Laguna; Basin; Pine Canyon; upper Cattail Canyon] 

Baccharis neglecta Britt. Roosevelt Weed; New Deal Weed 


[Brewster Co.: Rio Grande in Costolon area (SRSC)] 

Baccharis pteronioides DC. Yerba de Pasmo Baccharis 

[Baccharis ramulosa (DC.) Gray] 

[Steyermark & Moore, 1933; Sperry, 1938; Mahler & Waterfall, 1964; Powell, 1998] 

[Brewster Co.: Pulliam Bluff, Chisos Mts., 02 Jun 1937, Warnock 728 (TEX-LL); Chisos 

Mts., Moore & Steyermark 3346] 

Baccharis salicifolia (Ruiz & Pavon) Pers. Seepwillow 

[Baccharis glutinosa Pers.] 

[Sperry, 1938; Mahler & Waterfall, 1964; Powell, 1998] 

[Brewster Co.: Chisos Mts., 10 Aug 1931, Mueller 8194 (TEX-LL); along Rio 

Grande, Boquillas and Panther Creek (Sperry, 1938)] 

Baccharis salicina Torr. & Gray Willow Baccharis 


[Brewster Co.: along Rio Grande (Sperry, 1938)] 

Baccharis wrightii Gray Wright Baccharis 


[Brewster Co.: Tornillo Flats, BBNP (SRSC)] 

Bahia absinthifolia Benth. var. dealbata (Gray) Gray Hairyseed Bahia 


[Brewster Co.: hills adjacent to Chisos Mts., Hot Springs and Tornillo 

Flats (Sperry, 1938)] 

Bahia biternata Gray 


[Brewster Co.: dry unwooded canyons of Chisos Mts. (Sperry, 1938)] 

Bahia dissecta (Gray) Britt. Ragleaf Bahia 

[Sperry, 1938; Ellison, 1964] 

[Records: Dog Creek Wash (Sperry, 1938); Chisos Mts., Mueller 8239 (CAS; GH, MICH; 

TEX-LL; US] 

Bahia pedata Gray Bluntscale Bahia 

[Sperry, 1938; Ellison, 1964] 

[Records: flats N of Chisos Mts. (Sperry, 1938); Chisos Mts., 1 Aug 1931, Mueller 8228 

(TEX-LL)] 

Baileya multiradiata Harv. & Gray ex Gray Desert Baileya 

[Sperry, 1938, Turner, 1993, mapped] 

[Brewster Co.: Persimmon Gap, lower Green Gulch, Hot Springs, Boquillas 

and along Rio Grande (Sperry, 1938)]

Berlandiera lyrata Benth. Lyerleaf Greeneyes 


[Brewster Co.: Pulliam Bluff (Sperry, 1938)] 

Bidens leptocephala Sherff 


[Brewster Co.: Pulliam Canyon and lower Cattail Canyon (Sperry, 1938)] 

Brickellia brachyphylla (Gray) Gray var. terlinguensis Flyr 

[Flyr, 1968] 

[Brewster Co.: Chisos Mts., 07 Aug 1915, Young s.n. (TEX-LL) (Type)] 

Brickellia californica (Torr. & Gray) Gray California Brickellbush 


[Brewster Co.: S slope of Mt. Emory (Sperry, 1938)] 

Brickellia coulteri Gray Coulter Brickellia 

[Steyermark & Moore, 1933; Turner, 1990, mapped; Powell, 1998] 

[Brewster Co.: Crown Mtn. in Chisos Mts. and near McKinney Springs in 

Dead Horse Mts. (Powell, 1998); Boquillas Canyon region, Moore & Steyermerk 3438 

(MO)] 

Note: Turner (1991) recognized varieties and mapped the BBNP plants as var. brachiata 

(Gray) Turner. 

Brickellia cylindracea Gray & Engelm. 


[Brewster Co.: Pine Canyon, upper Blue Creek and Panther Spring (Sperry, 1938)] 

Brickellia fendleri Gray 

[Eupatorium fendleri Gray] 

[Sperry, 1938; Correll & Johnston, 1970] 

[Brewster Co.: Chisos Mts. (Sperry, 1938; Correll & Johnston, 1970)] 

Brickellia laciniata Gray Splitleaf Brickellia 



Brickellia shineri Flyr 

[Eupatorium parryi Gray] 

[Steyermark & Moore, 1933; Correll & Johnston, 1970] 

[Brewster Co.: Chisos Mts., head of Cat Tail Canyon, Moore & Steyermark 3408 (MO)] 

Brickellia veronicifolia (Kunth) Gray var. petrophila (B. L. Robins.) 

B. L. Robins. 

[Sperry, 1938; Correll & Johnston, 1970; Powell, 1998] 

[Brewster Co.: Chisos Wall area (Sperry, 1938); Lost Mine Peak, Green Gulch, 

Chinese Wall, Laguna Trail and upper Cattail Canyon (Powell, 1998)] 

Carminatia tenuiflora DC. 

[Sperry, 1938; Turner, 1988] 

[Brewster Co.: upper Green Gulch (Sperry, 1938); upper Green Gulch, Warnock 387 (GH, 

TEX-LL)] 

Carpochaete bigelovii Gray Bigelow Bristlehead 

[Steyermark & Moore, 1933; Sperry, 1938; Powell & Powell, 1978; Turner, 1987; 

Jones & Jones, 1993; Powell, 1998] 


Pine Canyon, Green Gulch, Dead Man’s Cut and near Boot Springs (Powell, 1998); 

Divide and Green Gulch (Sperry, 1938); Chisos Mts., Averett 240, n=11 (SRSC or TEX-LL); 

above Lost Mine Peak Trail, Chisos Mts., 02 Apr 1959, Correll 20692 

(TEX-LL); Chisos Mts., Moore & Steyermark 3201] 

Centaurea americana Nutt. American Basketflower 


[Brewster Co.: desert N of Chisos Mts. (Sperry, 1938)] 

Chaetopappa ericoides (Torr.) Nesom Baby White Aster 

[Aster leucelene Blake; Leucelene ericoides (Torr.) Greene] 

[Sperry, 1938; Shinners, 1946]

[Brewster Co.: Chisos Mts. (Sperry, 1938); Chisos Mts., Mueller 8196 (GH)] 

Chaptalia texana Greene 

[Chaptalia nutans (L.) Polak. var. texana (Greene) Burkart] 

[Sperry, 1938; McVaugh, 1952, mapped; Nesom, 1995, mapped] 

[Records: lower Cattail Canyon (Sperry, 1938)] 

Chloracantha spinosa (Benth.) Nesom var. spinosa Devilweed 

[Aster spinosus Benth.; Erigeron ortegae Blake; Leucosyris spinosa (Benth.) Greene] 

[Sundberg, 1991, mapped; Powell, 1998] 

[Records: Rio Grande below Hot Springs (Powell, 1998)] 

Chromolaena bigelovii (Gray) King & Robinson 

[Eupatorium bigelovii Gray] 

[Nesom & Turner, 2006] 

[Records: Chisos Mts., Warnock 9587 (SRSC); Sierra Quemada, Claro 2 Spring, Bartel 211 

(SRSC)] 

Chrysactina mexicana Gray Damianita 

[Steyermark & Moore, 1933; Sperry, 1938; Wells, 1965; Strother, 1977, mapped] 

[Records: Dead Horse Mts. (Wells, 1965); Pulliam Bluff, Basin, Tornillo 

Flats and Persimmon Gap (Sperry, 1938); upper Lost Mine Trail, BBNP, 26 Jun 1978, 

Freeman, s.n. (UTEP); Chisos Mts., Moore & Steyermark 3170] 

Chrysothamnus pulchellus (Gray) Greene ssp. baileyi (Woot. & 

Standl.) Hall & Clements 



Chrysothamnus spathulatus L. C. Anderson 

[Chrysothamnus viscidiflorus (Hook.) Nutt. var. ludens Shinners] 


[Records: Chisos Mts, Laguna trail to Cattail Canyon (Powell, 1998)] 

Cirsium turneri Warnock 

[Warnock, 1960] 

[Records: Chisos Mts., 11 Aug 1915, Young s.n. (TEX-LL, MO); upper 

Pine Canyon, Chisos Mts., 5200 ft., 01 Aug 1950, Warnock 9175 (SRSC)] 

Cirsium undulatum (Nutt.) Spreng. Wavyleaf Thistle 

[Sperry, 1938; Wells, 1965] 

[Records: Dead Horse Mts. (Wells, 1965); Blue Creek and Oak Creek Canyons 

(Sperry, 1938)] 

Conyza canadensis L. Horseweed 


[Records: lower Cattail Canton, Green Gulch, Laguna, S slope of Mt. 

Emory, Boquillas, lower Willow Creek Canyon and Government Spring 

(Sperry, 1938)] 

Cosmos parviflorus (Jacq.) Pers. 


[Records: moist canyons of Chisos Mts. (Sperry, 1938)] 

Eclipta prostrata (L.) L. 


[Records: Rio Grande near San Vicente Canyon (Sperry, 1938)] 

Erigeron divergens Torr. & Gray Spreading Fleabane 


[Records: flats N of Chisos Mts. and Green Gulch (Sperry, 1938)] 

Erigeron flagellaris Gray Trailing Fleabane 

[Erigeron nudiflorus Buckl.] 


[Records: Green Gulch and Basin (Sperry, 1938)] 

Erigeron modestus Gray 

[Cronquist, 1947] 

[Records: Blue Creek Trail, 10 Sep 1933, Cory 7352 (GH); Chisos Mts., 1 Aug 1931,

Mueller 8191 (GH)] 

Eupatorium greggii Gray 

[Conoclinium greggii (Gray) Small] 


[Records: Basin and Green Gulch (Sperry, 1938)] 

Eupatorium solidaginifolium Gray 

[Koanophyllon solidaginifolia (Gray) King & Robins.] 


[Brewster Co.: Juniper Canyon (Sperry, 1938); Basin (Powell, 1998)] 

Flaveria trinervia (Spreng.) C. Mohr 


[Brewster Co.: lower Oak Canyon and Boquillas (Sperry, 1938)] 

Flourensia cernua DC. Tarbush; Hojase; Blackbrush 

[Sperry, 1938; Wells, 1965] 

[Brewster Co.: Dead Horse Mts. (Wells, 1965); dry flats N of Chisos Mts. (Sperry, 1938)] 

Flyriella parryi (Gray) King & H. E. Robins. 

[Eupatorium parryi Gray] 

[Sperry, 1938; Baker & Turner, 1986] 

[Records: end of Pine Canyon Trail, 12 Nov. 1976, Correll 35402 (TEX-LL; MO); 

Cattail Falls, 09 Nov 1964, Correll & Correll 30592 (TEX-LL); Juniper Canyon, 16 JUN 

1964, Correll et al. 29721 (TEX-LL); Pine Canyon, 17 Jun 1963, Correll & Wasshausen 

27865 (CAS; TEX-LL; MO); Chisos Mts., Aug 1935, Marsh 149 (F); Cattail Springs, 07 

Jul 1931, Moore & Steyermark 3408 (CAS; GH; TEX-LL; MO; US); Chisos Mts., o5 Aug 

1931, Mueller 8179 (GH; TEX-LL; US); Pine Canyon near waterfall, 27 SEP 1977, Powell 

et al. 3229 (TEX-LL; US); lower Pulliam Canyon, 04 Jul 1936, Sperry 193 (TEX-LL; US); 

Cattail Falls, 19 Apr 1955, Wallmo 7 (TEX-LL); waterfalls in Pine Canyon, 01 Jul 1937, 

Warnock 760 (GH; TEX-LL; US0; mid Green Gulch, 23 Jul 1950, Warnock 9146 (TEX-LL); 

upper Green Gulch, 28 Jun 1941, Warnock 20975 (GH; TEX-LL, MO); Green Gulch, 22 

Aug 1947, Warnock & Tharp 4005 (US); 1 mi. below Basin Campground on trail to Window, 

BBNP, 26 May 1982, Van Devender s.n. (UTEP)] 

Gaillardia pinnatifida Torr. Cut-leaved Blanket Flower 

[Sperry, 1938; Biddulph, 1944] 

[Records: Persimmon Gap, flats N of Chisos Mts. and Comanche Spring (Sperry, 1938); 

foothills of Chisos Mts., 12 Sep 1915, Young (MO); Comanche Springs, Chisos Mts, area, 

17 Jul 1937, Warnock 729 (US); Dog Flats, 31 Mar 1940, Sperry 1750 (GH)] 

Galinsoga parviflora Cav. 

[Canne, 1977] 

[Records: Chisos Mts., above Boot Spring, Warnock 9740 (TEX-LL)] 

Gnaphalium canescens DC. Wright Cudweed 

[Gnaphalium wrightii Gray] 


[Brewster Co.: slopes of Mt. Emory (Sperry, 1938)] 

Gnaphalium stramineum Kunth Cotton-batting Cudweed 

[Gnaphalium chilense Spreng.] 


[Brewster Co.: moist canyons of the Chisos Mts. (Sperry, 1938)] 

Gnaphalium viscosum Kunth 

[Gnaphalium macounii Greene] 


[Brewster Co.: N slope of Mt. Emory and trail from Boot Spring to South Rim 

(Sperry, 1938)] 

Note: Material reported by Sperry (1938) to be Gnaphalium microcephalum Nutt. must 

have been something as the taxon is not reported as occurring in Texas by Correll and 

Johnston (1970). 

Grindelia arizonica Gray var. arizonica 

[Nesom, 1990, mapped]

Grindelia havardii Steyermark 

[Steyermark, 1934; Sperry, 1938] 

[Brewster Co.: Panther Creek Canyon (Sperry, 1938); open valley, Nail Place, Chisos 

Mts., Aug 1916, Young (TEX-LL)] 

Gutierrezia microcephala (DC.) Gray Threadleaf Snakeweed 

[Gutierrezia lucida (Greene) Greene] 

[Sperry, 1938; Lane, 1985, mapped] 

[Brewster Co.: Painted Gap, Green Gulch, Basin and upper Pine Canyon (Sperry, 1938)] 

Gutierrezia sarothrae (Pursh) Britt. & Rusby Broom Snakeweed 

[Wells, 1965; Lane, 1985, mapped] 

[Brewster Co.: Dead Horse Mts. (Wells, 1965)] 

Gutierrezia sphaerocephala Gray Roundleaf Snakeweed 

[Gutierrezia eriocarpa Gray] 

[Sperry, 1938; Solbrig, 1960] 

[Brewster Co.: flats N of Chisos Mts. and Dog Creek Wash (Sperry, 1938); Chisos 

Mts., 15-18 Aug 1921, Ferris & Duncan as G. glutinosa (NY)] 

Gymnosperma glutinosum (Spreng.) Less. Tatalencho 

[Selloa glutinosa Spreng.] 

[Sperry, 1938; Wells, 1965] 

[Brewster Co.: Dead Horse Mts. (Wells, 1965); Terrapin Tank (Sperry, 1938); Basin 

near campground,Aug 1983, Stafford s.n. (UTEP)] 

Haploesthes greggii Gray var. texana (Coult.) I. M. Johnst. 

[Sperry, 1938; Turner, 1975; Powell, 1998] 

[Brewster Co.: near Lajitas and Terlingua (Sperry, 1938); Grapevine Hills, Alamo 

Springs and 15 mi. SE Costolon (Powell, 1998); Chisos Mts., 1889, Nealley 203 (Type: US)] 

Helenium elegans DC. var. amphibolum (Gray) Bierner 

[Sperry, 1938; Bierner, 1972, mapped] 

[Brewster Co.: Rio Grande near San Vicente Canyon (Sperry, 1938)] 

Note: Sperry (1938) also reports var. amphilobum (Gray) Bierner from S of the Chisos 

Mts. 

Helenium microcephalum DC. var. ooclinium (Gray) Bierner 

[Sperry, 1938; Bierner, 1972, mapped] 

[Brewster Co.: Boquillas (Sperry, 1938)] 

Note: Sperry (1938) reports var. ooclinium (Gray) Bierner from upper Green Gulch, 

Lajitas and Terrapin Tank. 

Helianthus annuus L. Common Sunflower 

[Sperry, 1938; Heiser et al., 1969, mapped] 


Helianthus ciliaris DC. 

[Heiser et al., 1969, mapped] 

Helianthus niveus (Benth.) Brandeg. ssp. canescens (Gray) Heiser 

[Helianthus canus (Britt.) Woot. & Standl.] 

[Sperry, 1938; Heiser et al., 1969, mapped ] 

[Brewster Co.: near Santa Elena Canyon (Sperry, 1938)] 

Heliomeris longifolia (Robins. & Greenm.) Cockerell Longleaf Goldeneye 

[Viguiera longifolia (Robins. & Greenm.) Blake] 



Heliopsis parvifolia Gray 

[Steyermark & Moore, 1933; Sperry, 1938; Turner, 1988, mapped] 

[Brewster Co.: Basin, Squaw Creek Canyon and Pulliam Canyon (Sperry, 1938); in 

Basin at saddle above Green Gulch, BBNP, 15 Jul 1981, Freeman s.n. (UTEP); Chisos 


Heterosperma pinnatum Cav. 


[Brewster Co.: wooded areas of Chisos Mts. (Sperry, 1938)]

Heterotheca fulcrata (E. L. Greene) Shinners Rocky Goldaster 

var. fulcrata 

[Semple, 1996] 

[Brewster Co.: Chisos Mts., Bick Rock Trail, Cory 44873 (TEX-LL) “indument 

approaching var. senilis”] 

var. arizonica Semple 


[Brewster Co.: Chisos Mts., Mt. Emory, Warnock 9706 (SRSC) “approaching var. senilis”] 

var. senilis (Woot. & Standl.) Semple 

[Steyermark & Moore, 1933; Semple, 1996] 

[Brewster Co.: Chisos Mts., Mueller s.n. (NY), Mueller 8199 (NY), Warnock s.n. (ASU); 

Boot Spring, Mueller s.n. (TEX-LL); Cattail Falls, Powell & Powell 4205 (SRSC, WAT); 

Emory Peak, Correll 29749 (TEX-LL), Moore & Steyermark 3212 (CAS, DS, MIN, RM, NY), 

Mueller s.n.(GH); Green Gulch, Lundell & Lundell 13266 (TEX-LL); crest of Juniper Canyon, 

Ferris & Duncan 2790 (CAS, DS, NY); Nail Place, Young s.n. (TEX-LL); North Wall, Cory 

44872 (TEX-LL, NY); Pulliam Canyon, Sperry 417 (SRSC), Warnock 6955 (SRSC); 

Pulliam Bluff, Warnock 767 (SRSC); Window Trail, Cory 44772 (TEX-LL); et al. approaching 

Other vars. In Semple (1996, p. 80)] 

Heterotheca subaxillaris (Lam.) Britt. & Rusby 

[Heterotheca psammophila Wagenkn.] 

[Wagenknecht, 1960; Harms, 1965, mapped] 

[Records: Santa Elena Canyon, Moore & Steyermark 3467 (GH; PH; UC)] 

Heterotheca viscida (Gray) Harms 


[Records: Mexico: Chihauhua: Fern Canyon side of Santa Elena Canyon, Wendt & Lott 96 

(TEX-LL)] 

Note: This taxon is expected to be part of the flora of BBNP. 

Hieracium wrightii (Gray) Robins. & Greenm. 


[Brewster Co.: upper Green Gulch (Sperry, 1938)] 

Hymenoclea monogyra Torr. & Gray ex Gray Burrobrush 


[Brewster Co.: to lower Chisos Mts. and Rio Grande (Powell, 1998)] 

Isocoma pleuriflora (Torr. & Gray) Greene Jimmyweed 

[Haplopappus heterophyllus (Gray) Blake; Isocoma wrightii (Gray) Greene] 

[Sperry, 1938; Nesom, 1991, mapped] 

[Brewster Co.: Boquillas, Hot Spring, Persimmon Gap and Pinnacle Mts. 

(Sperry, 1938)] 

Iva ambrosiifolia (Gray) Gray 


[Brewster Co.: low areas N of Chisos Mts., Basin and Robbers’ Roost area 

(Sperry, 1938)] 

Jefea brevifolia (Gray) Strother Shorthorn Jefea 

[Zexmenia brevifolia Gray] 

[Sperry, 1938; Wells, 1965; Strother, 1991; Powell, 1998] 

[Brewster Co.: Dead Horse Mts. (Wells, 1965); Chisos and Dead Horse Mts. 

(Powell, 1998); Kit to Ghost Mtn., lower Juniper Canyon and Cattail Falls 

(Sperry, 1938); 1 mi. SE of Headquarters on road to Boquillas, 04 Oct 1964, Flyr 

218A (OBI, TEX-LL); lower Juniper Spring, 23 Jun 1937, Warnock 724 (ARIZ, F, 

OBI, TEX-LL, UC)] 

Lactuca ludoviciana (Nutt.) Riddell 



Laennecia coulteri (Gray) Nesom Coulter Laennecia 

[Conyza coulteri Gray] 

[Sperry, 1938]

[Brewster Co.: Green Gulch and Basin (Sperry, 1938)] 

Laennecia sophiefolia (Kunth) Nesom 

[Conyza sophiifolia Kunth] 


[Records: S slope of Mt. Emory and Laguna (Sperry, 1938)] 

Liatris punctata Hook. Dotted Gayflower 

[Sperry, 1938; Gaiser, 1946] 

[Records: Laguna and Basin (Sperry, 1938); Pine Canyon, 5 Sep 1937, Marsh 330 (GH); 

Blue Creek, 19 Jun 1931, Mueller 8176 (GH); Chisos Mts., Young (GH)] 

Machaeranthera pinnatifida (Hook.) Shinners var. chihuahuana Turner Lacy Tansyaster 

& Hartman 

[Sperry, 1938; Turner & Hartman, 1976, mapped; Powell & Powell, 1978] 

[Records: Chisos Mts. (Sperry, 1938); BBNP, n=4, Turner s.n.(TEX-LL)] 

Machaeranthera tanacetifolia (H.B.K.) Nees Tansyleaf Machaeranthera 


[Records: Tornillo Flats (Sperry, 1938)] 

Melampodium leucanthum Torr. & Gray Plains Blackfoot 

[Sperry, 1938; Wells, 1965; Stuessy, 1971, 1972] 

[Records: Dead Horse Mts. (Wells, 1965); Basin, Green Gulch and N of 

Chisos Mts. (Sperry, 1938); 6 mi. N Chisos Basin Lodge, Tomb 137 (TEX-LL)] 

Nicolletia edwardsii Gray 

[Sperry, 1938; Strother, 1978, mapped in BBNP] 

[Records: Hot Springs (Sperry, 1938)] 

Palafoxia riograndensis Cory 

[Turner & Morris, 1976] 

[Records: near mouth of Santa Elena Canyon, Shinners 8792 (BRIT), Warnock 1160 (GH), 

Sperry 1160 (US), and McVaugh 12708 (MICH)] 

Palafoxia rosea (Bush) Cory var. macrolepis (Rydb.) Turner & Morris 

[Othake macrolepis Rydb.] 


[Records: Rio Grande near Santa Elena Canyon (Sperry, 1938)] 

Parthenium argentatum Gray Guayule 

[Wells, 1965; Powell, 1998] 


Parthenium confertum Gray var. lyratum (Gray) Rollins Lyerleaf Parthenium 


[Brewster Co.: flats around Chisos Mts. and in lower Juniper Canyon (Sperry, 

1938)] 

Parthenium incanum H.B.K. Mariola 

[Sperry, 1938; Wells, 1965] 

[Brewster Co.: Dead Horse Mts. (Wells, 1965); about the Chisos Mts. and in 

Juniper Canyon (Sperry, 1938)] 

Pectis angustifolia Torr. var. angustifolia 

[Sperry, 1938; Keil, 1977, mapped] 

[Records: area of Johnson’s Ranch (Sperry, 1938); 1.3 mi. N of pavement on 

Old Ore Rd. to Ernst Tinaja, 21 Aug 1982, Van Devender s.n. (UTEP); BBNP, Keil & 

McGill 7795a (OS); 15.1 mi. N of Castolon, Keil & McGill 7798a (OS)] 

Note: Sperry (1938) reports var. tenella from flats around the Chisos Mts. and Keil 

(1977) maps it from the Black Gap area and to the west of BBNP. 

Pectis filipes Harv. & Gray var. subnuda Fern. 

[Sperry, 1938; Keil, 1977] 

[Records: BBNP, Correll & Correll 35379 (ENCB, MICH, SD, TEX-LL, US); 

15.1 mi. N of Costolon, K & M 7797 (OS); 12.5 mi. N of Castalon, Keil & McGill 7811 

(OS); hills W of Burnham’s Ranch and Lone Mtn. (Sperry, 1938)] 

Pectis papposa Harvey & Gray in A. Gray var. grandis Keil 

[Sperry, 1938; Keil, 1974; Keil, 1977]

[Records: Tornillo Flats, 28 Jun 1941, Warnock 21681 (F, NY, PH, BRIT, TEX-LL); 

BBNP, Keil & McGill 7818 (OS); Smoky Creek (Sperry, 1938)] 

Pectis prostrata Cav. 


[Brewster Co.: Oak Creek Canyon (Sperry, 1938)] 

Pericome caudata Gray Tailleaf Pericome 

[Sperry, 1938; Powell, 1973; Powell, 1998] 

[Brewster Co.: slopes of Casa Grande (Sperry, 1938); upper Green Gulch, Chisos 

Mts., Powell 1554 (SRSC) (Powell, 1973, 1998)] 

Perityle aglossa Gray Rayless Perityle 


[Brewster Co.: near Big Bend tunnel, 4 mi. W of Hot Springs and San Vivente 

Canyon (Powell, 1998)] 

Perityle dissecta (Torr.) Gray Slimlobe Rockdaisy 


[Brewster Co.: Santa Elene Canyon (Powell, 1998)] 

Perityle parryi Gray 

[Steyermark & Moore, 1933; Sperry, 1938; Powell, 1998] 

[Brewster Co.: Pulliam Canyon and Oak Creek Canyon (Sperry, 1938, #301 & 753); 

Chisos Mts. and vicinity including Santa Elena Canyon (Powell, 1998); Santa Elena 

Canyon, Moore & Steyermark 3466 (MO)] 

Perityle rupestris (Gray) Shinners var. albiflora Powell 

[Laphamia rupestris Gray] 


[Records: top of Casa Grande (Sperry, 1938, #742)] 

Perityle vaseyi Coult. Margined Perityle 


[Records: hills N of Chisos Mts., Hot Springs and Fisk Canyon (Sperry, 1938); 

near Hot Springs to near Costolon (Powell, 1998); Tunnel W of Rio Grande Village, 

BBNP, 28 May 1982, Toolin s.n. (UTEP)] 

Pinaropappus roseus Less. White Rock Lettuce 


[Records: Basin (Sperry, 1938)] 

Pluchea sericea (Nutt.) Cav. Arroweed 

[Tessaria sericea (Nutt.) Shinners] 


[Records: Tornillo Flats and Chisos Pens (Sperry, 1938); Chisos Mts. area, 

Tornillo Creek and Chisos Pens (Powell, 1998)] 

Porophyllum scoparium Gray Poreleaf 

[Sperry, 1938; Johnson, 1969; Powell, 1998] 

[Records: Oak Creek Canyon, Muskhog Spring, Chisos Pens and W 

slope of Dead Horse Mts. (Sperry, 1938); Chisos Mts., Blue Creek, June 1931, Mueller 8240 

(F, GH, MICH, MO, NY, TEX-LL, UC, US)] 

Psathyrotes scaposa Gray 

[Sperry, 1938; Strother & Pilz, 1975, mapped] 

[Records: hills and slopes N of Hot Springs (Sperry, 1938)] 

Psilostrophe gnaphalioides DC. 

[Sperry, 1938; Heiser, 1944; Brown, 1977; Brown, 1978, mapped] 

[Brewster Co.: Persimmon Gap area and lower Green Gulch (Sperry, 1938); 1.2 mi. N of 

Persimmon Gap Ranger Station, Keil 7830 (ASU); Chisos Mts., Young (TEX-LL); Chisos 

Mts., Sperry 743 (US)] 

Psilostrophe tagetina (Nutt.) Greene Wooly Paperflower 

[Sperry, 1938; Heiser, 1944; Brown, 1977; Brown, 1978, mapped] 

[Brewster Co.: Basin, Green Gulch, Mesa de Anguila and Panther Spring (Sperry, 1938); 

1.5 mi. N of Castolon, Keil 7802A and 7802 (ASU); Chisos Mts., Mueler 8231 (GH; MO; NY; 

TEX-LL); Willow Creek and Green Gulch Canyons, Sperry 250 (US); lower Green Gulch,

Warnock 1232 (GH); near Castellon, Palmer 34216 (NY); Boquillas, Hanson 608 (MO; US), 650 

(US); Santa Elena Canyon, Innes & Warnock 501 (GH); east of Chisos Mts., Sperry 1709 (GH)] 

Ratibida columnifera (Nutt.) Woot. & Standl. Mexican Hat 


[Brewster Co.: Laguna (Sperry, 1938)] 

Schkuhria multiflora Hook. & Arn. 

[Bahia neomexicana (Gray) Gray] 

[Heiser, 1945] 

[Brewster Co.: Chisos Mts., Mueller 8232 (F, GH, MO)] 

Schkuhria pinnata (Lam.) Kuntze ex Thell. var. wislizeni (Gray) Turner 

[Schkuhria anthemoidea (DC.) Coult. var. wrightii Gray) Heiser] 

[Sperry, 1938; Turner, 1995, taxonomy] 

[Brewster Co.: moist canyons in the Chisos Mts. (Sperry, 1938)] 

Senecio flaccidus Less. var. flaccidus Threadleaf Groundsel 

[Senecio longilobus Benth.] 


[Brewster Co.: Green Gulch (Sperry, 1938)] 

Senecio millelobatus Rydb. 


[Records: Pulliam Canyon, Green Gulch and Boot Spring area (Sperry, 1938); Pine Canyon, 

Muller s.n. (CAS; KSC; MO; NY; PH; RSA)] 

Senecio parryi Gray 


[Records: upper Green Gulch (Sperry, 1938)] 

Solidago juliae Nesom 

[Nesom, 1989] 

[Brewster Co.: old Grapevine Hills Ranch, BBNP, 12 Nov 1947, Correll & Correll 35408 

(TEX-LL); Cattail Falls, BBNP, 09 Nov 1964, Correll & Correll 30585 (TEX-LL)] 

Solidago radula Nutt. 


[Brewster Co.: Green Gulch, upper slopes of Casa Grande and Boot Spring (Sperry, 

1938)] 

Solidago velutina DC. 

[Solidago trinervata Greene] 


[Brewster Co.: Boot Spring area (Sperry, 1938)] 

Sonchus oleraceus L. Annual Sow Thistle 


[Brewster Co.: along Rio Grande at Boquillas (Sperry, 1938)] 

Stephanomeria pauciflora (Torr.) A. Nels. Desert Straw; Skeleton Plant 

[Ptiloria pauciflora (Torr.) Raf.] 


[Brewster Co.: Basin, Chisos Pens and Boquillas (Sperry, 1938)] 

Stevia serrata Cav. 


[Brewster Co.: Saddle, Boot Springa sna Juniper Canyon (Sperry, 1938)] 

Synphotrichum subulatum (Michx.) Nesom 

[Aster exilis Ell.; Aster subulatus Michx.] 


[Brewster Co.: Gano Spring and along Rio Grande near Boquillas (Sperry, 1938)] 

Tagetes micrantha Cav. 


[Brewster Co.: moist canyons in the Chisos Mts. (Sperry, 1938)] 

Tetraneuris scaposa (DC.) Greene var. linearis (Nutt.) Parker Clustered Bitterweed 


[Brewster Co.: Basin and Green Gulch (Sperry, 1938); above Tunnel near Rio Grande

Village, BBNP, 21 Aug 1982, Van Devender s.n. (UTEP)] 

Thelesperma ambiguum Gray 



Thelesperma longipes Gray Longstalk Greenthread 


[Brewster Co.: McKinney Spring area (Sperry, 1938); Dead Horse Mts. (Powell, 1998)] 

Thelesperma megapotamicum (Spreng.) Kuntze Colorado Greenthread 

[Thelesperma gracile (Torr.) Gray] 


[Brewster Co.: arroyos on flats N of Chisos Mts., Muskhog Spring and Hot Springs 

[Sperry, 1938)] 

Thymophylla aurea (Gray) Greene ex Britt. var. polychaeta (Gray) Strother 

[Sperry, 1938; Keil & Stuessy, 1975] 

[Records:Tornillo Flats (Sperry, 1938); 8.7 mi. N of Headquarters, Keil & McGill (OS)] 

Thymophylla pentachaeta (DC.) Small var. hartwegii (Gray) Strother Parralena 

[Sperry, 1938; Wells, 1965; Strother, 1969] 

[Brewster Co.: Dead Horse Mts. (Wells, 1965); along Rio Grande, Solis’ Ranch 

to Mariscal Canyon and Tornillo Flats (Sperry, 1938); Chisos Mts., 15 Aug 1915, Young 

(MO, TEX-LL)] 

Trixis californica Kell. var. californica American Trixis 

[Sperry, 1938; Anderson, 1972; Keil & Stuessy, 1975; Powell, 1998] 

[Records: Panther Canyon, lower Juniper Canyon and Hot Springs area 

(Sperry, 1938); Chisos Mts. area and Dead Horse Mts. (Powell, 1998); 14 mi. E of 

Castolon, s edge of Chisos Mts., 11 May 1937, Cutler 1002 (NY); BBNP, roadside in 

Green Gulch, 04 Oct 1964, Flyr 215 (MSC); Goat Mtn., 6500 ft., 22 Aug 1935, Hinckley 

379 (NY); mouth of Santa Elena Canyon, 103 DEG 37’W, 640 m, 10 Apr 1952, McVaugh 

12678 (MICH); Chisos Mts., near Blue Creek, 1400 m, 06 Jul 1931, Moore & Steyermark 

3404 (CAS, MICH, NY); 1.5 mi. N of Costollon, Keil & McGill 7814 (OS)] 

Verbesina encelioides (Cav.) Gray Cowpen Daisy 


[Brewster Co.: Dog Creek Wash, Terrapin Tank and Boquillas (Sperry, 1938)] 

Verbesina oreophila Woot. & Standl. 


[Brewster Co.: Pilliam Canyon (Sperry, 1938)] 

Vernonia marginata (Torr.) Raf. Plains Ironweed 

[Vernonia tenuifolia Small] 


[Brewster Co.: Fisk Canyon and Pinnacle Mts. (Sperry, 1938)] 

Viguiera cordifolia Gray Heartleaf Goldeneye 



Viguiera dentata (Cav.) Spreng. Sunflower Goldeneye 


[Brewster Co.: lower Pulliam Canyon (Sperry, 1938)] 

Viguiera stenoloba Blake var. chihuahuensis Butterwick Skeleton Goldeneye 

[Sperry, 1938; Wells, 1965] 

[Brewster Co.: Dead Horse Mts. (Wells, 1965); Basin, lower Juniper Canyon and 

Green Gulch (Sperry, 1938)] 

Xylorhiza wrightii (Gray) Greene 

[Machaeranthera wrightii (Gray) Cronq. & Keck] 

[Watson, 1978] 

[Records: Brewster Co., Watson 401, 403, 626 (TEX-LL)] 

Xylothamia triantha (Blake) Nesom 

[Ericameria triantha (Blake) Shinners] 

[Sperry, 1938; Nesom, 1990; Powell, 1998]

[Brewster Co.: Study Butte area (Sperry, 1938); between Santa Elene and Costolon, 

1 mi. beyond Tornillo Flats toward the Chisos Mts. and near Hot Springs (Powell, 1998); 

along road from Study Butte to Terlingua, 31 Aug 1937, Warnock 1126 (US)] 

Zinnia acerosa (DC.) Gray Spinyleaf Zinnia 

[Zinnia pumila Gray] 

[Sperry, 1938; Wells, 1965; Powell, 1998] 

[Brewster Co.: Dead Horse Mts. (Wells, 1965); desert areas N of Chisos Mts. 

(Sperry, 1938)] 

BERBERIDACEAE (Barberry Family) 

Mahonia haematocarpa (Woot.) Fedde Red Berberis 

[Berberis haematocarpa Woot.] 


[Brewster Co.: Chisos Mts. (Powell, 1998); Laguna, Pulliam Canyon and Pulliam 

Bluff (Sperry, 1938); Chisos Mts., Moore & Steyermark 3311] 

Mahonia trifoliata (Moric.) Fedde Agarito 

[Berberis trifoliata Moric. var. glauca (I. M. Johnst.) M. C. Johnst. ex Laf.] 

[Sperry, 1938; Johnston, 1950; Ahrendt, 1961; Wells, 1965; Powell, 1998] 

[Brewster Co.: arroyos and washes E of Lone Mountain (Sperry, 1938); Dead 

Horse Mts. (Wells, 1965); Chisos Mts., Moore & Steyermark 3121,3291] 

BETULACEAE (Birch Family) 

Ostrya virginiana (Mill.) K. Koch var. chisosensis (Correll) Henrickson Big Bend Hophornbean 

[Ostrya chisosensis Correll] 


[Brewster Co.: Emory Peak, Crown Mt. and above Boot Spring in Chisos Mts. (Powell, 

1998); Emory Peak at base of north facing ledge, 17 Jun 1964, Correll 29733 (Type: TEX-LL; 

Isotypes: US; GH; TEX-LL); near Nail Place, 21 Aug 1915, Young s.n. (TEX-LL); north of 

Crown Mtn., 1 Jul 1937, Warnock 21633 (TEX-LL); Wmory-Boot Trail, 7500 ft. 8 Jun 1952, 

Warnock 10478; north side of Emory Peak, 15 Jul 1955, Warnock 12726] 

BIGNONIACEAE (Catalpa Family) 

Chilopsis linearis (Cav.) Sweet ssp. linearis Desertwillow 

[Sperry, 1938; Fosberg, 1936; Henrickson, 1985, mapped] 

[Chilopsis linearis (Cav.) Sweet var. glutinosa (Engelm.) Fosberg] 

[Brewster Co.: along Rio Grande, Basin, Glenn Spring and Gano Spring (Sperry, 

1938); Chisos Mts., Moore & Steyermark] 

Tecoma stans (L.) Juss. ex Kunth Trumpetflower; Esperanza 

[Stenolobium incisum Rose & Standl.] 


[Brewster Co.: Glenn Spring area, Green Gulch, Oak Creek Canyon and Costolon 

(Sperry, 1938); Cattail Falls, BBNP, Aug 1983, Stafford s.n. (UTEP)] 

BORAGINACEAE (Borage Family) 

Cryptantha albida (H.B.K.) I. M. Johnst. 


[Brewster Co.: Dog Creek Wash and Tornillo Flats (Sperry, 1938)] 

Cryptantha angustifolia (Torr.) Greene 

[Johnston, 1967] 

[Brewster Co.: 11 mi. S of Persimmon Gap, 02 Apr 1959, Correll 20642; 4 mi. N of

Tornillo Creek, hwy. 51, 02 Apr 1959, Correll 20670; mouth of Dog Canyon, 20 Mar 

1941, Warnock 552; hills at mouth of Santa Elena Canyon, 16 Mar 1947, Warnock 

47040] 

Cryptantha crassisepala (Torr. & Gray) Greene var. elachantha I. M. Johnston 

[Johnston, 1959, 1967] 

[Brewster Co.: 4 mi. N Tornillo Creek, hwy. 51, 02 Apr 1959, Correll 20671; mouth 

of Santa Elena Canyon, 16 Mar 1947, Warnock 47040 (TEX-LL); mouth of Terlingua Creek, 

14 Apr 1948, Whitehouse 19632; below Santa Elena Canyon, 14 Apr 1948, Whitehouse 

19600, 19648] 

Cryptantha mexicana (Brandeg.) I. M. Johnston Mexican Cryptantha 


[Brewster Co.: 11 mi. S of Persimmon Gap, 02 Apr 1959, Correll 20638, 20640, 20641; 

4 mi. N Tornillo Creek, hwy. 51, 02 Apr 1959, Correll 20672; Boquillas, 17 Apr 1919, 

Hanson 603; Persimmon Gap, 24 Apr 1931, McKelvey 1970; 3 mi. S Persimmon Gap, 

05 Apr 1947, McVaugh 7840; Santa Elena Canyon, 19 Mar 1941, Rose-Innes & Warnock 

21580; Tornillo Flats, 13 Feb 1937, Sperry 527, 27 Mar 1937, Sperry 550; between Dog 

Flats and Persimmon Gap, 02 Apr 1938, Warnock C-272] 

Cryptantha palmeri (Gray) Payson 

[Cryptantha coryi I. M. Johnston] 


[Brewster Co.: 4 mi. N Tornillo Creek, hwy. 51, 02 Apr 1959, Correll 20651; 25 mi. N of 

Boquillas, 17 Apr 1919, Hanson 593; near Persimmon Gap, 25 Apr 1931, McKelvey1979; 

Dog Flats, 02 Apr 1938, Warnock C-295; hills near Dog Flats, 10 Apr 1956, Warnock 

13917; between Castalon and Smoky Creek, 12 Jul 1955, Warnock 13918; 10 mi. N Hot 

Springs, BBNP, 20 Mar 1941, Warnock & Rose-Innes 524] 

Hackelia besseyi (Rydb.) J. L. Gentry 

[Hackelia grisea (Woot. & Standl.) I. M. Johnston] 

[Sperry, 1938; Johnston, 1967; Gentry, 1974; Gentry & Carr, 1976] 

[Brewster Co.: upper Green Gulch, Chisos Mts., 07 Sep 1937, Sperry 39, 03 Aug 

1941, Warnock 20036 (GH, TEX-LL)] 

Heliotropium confertifolium (Torr.) Torr. ex Gray Leafy Heliotrope 

[Steyermark & Moore, 1933; Sperry, 1938; Wells, 1965; Johnston, 1967] 

[Brewster Co.: 8 mi. NE Government Springs, 27 Jun 1931, Moore & Steyermark 

3264; Tornillo Flats, 08 Jun 1936, Sperry 123, 03 May 1937, Warnock 1221; Onion 

Springs, N side Paint Gap Hills, 05 Aug 1955, Warnock 129671 [sic]; W slope of 

Dead Horse Mts., 08 Jun 1937, Warnock 907; along Dead Horse Mts., 03 May 1937, 

Warnock s.n.; Sue Peak, Dead Horse Mts., 19 Jul 1952, Warnock 10848; Mesa near 

Chisos Mts., 30 Aug 1915, Young s.n.; rd. from Panther Jct. to Rio Grande Village 

1.7 rd. mi. N Boquillas canyon turnoff, 29 DEG 12’N, 102 DEG 59’W, 2000 ft., 

15 Mar 1983, Yatskievych (UTEP)] 

Heliotropium convolvulaceum (Nutt.) Gray 

[Steyermark & Moore, 1933; Sperry, 1938; Johnston, 1967] 

[Brewster Co.: W end Boquillas Canyon, dunes, Moore & Steyermark 3431; Boquillas 

Canyon, 05 Aug 1937, Warnock 686] 

Heliotropium curassavicum L. var. currasavicum Salt Heliotrope 


[Brewster Co.: W end of Boquillas Canyon, 1931, Moore & Steyermark 3436; 

Boquillas, Jun 1931, Tharp 8802; Boquillas Canyon, 04 Aug 1937, Warnock 943; 

Along Rio Grande at Mariscal Canyon, 22 Jun 1937, Warnock 942; banks of the 

Rio Grande, Big Bend, 26 Aug 1915, Young s.n.] 

Heliotropium glabrisculum (Torr.) Gray 


[Brewster Co.: Dog Flats, 30 Jul 1938, Warnock 550, Sperry 1440; flats on road to Rosillas 

Mts., Big Bend, 30 Aug 1937, Warnock 1089; near Maravillas, 05 Jun 1941, Warnock 20119] 

Heliotropium greggii Torr. Fragrant Heliotrope 

[Johnston, 1967]

[Brewster Co.: 3 mi. S Persimmon Gap, 05 Apr 1947, McVaugh 7827; Dog Flats, 30 Jul 

1938, Sperry 1441; between Persimmon Gap and Dog Flats, 02 Apr 1938, Warnock 291; 

near Maravillas, 05 Jun 1941, Warnock 20118] 

Heliotropium molle (Gray) I. M. Johnston 

[Tournefortia monclovana Wats.] 


[Brewster Co.: 4 mi. SE Persimmon Gap, 03 Apr 1959, Correll 20698; Dog Flats, 17 Oct 

1937, Sperry 1154; Rio Grande near Santa Elena Canyon, 07 Jun 1937, Warnock 680; 

et al.] 

Heliotropium procumbens Miller 


[Brewster Co.: Boquillas Canyon, 30 Jul 1937, Warnock 1089; Rio Grande near Mariscal 

Canyon, 22 Jun 1937, Warnock 942; Head of San Vicente Canyon, BBNP, Warnock & 

Parks 8763] 

Heliotropium torreyi I. M. Johnston Slimleaf Heliotrope 


[Brewster Co.: Mesa de Anguilla, 14 Jul 1937, Warnock 21121] 

Lappula occidentalis (S. Wats.) Greene 


[Brewster Co.: 11 mi. S of Persimmon Gap, 02 Apr 1959, Correll 20635, 20641; 4 mi. N 

of Tornillo Creek, 02 Apr 1959, Correll 20673; Tornillo Flats, 20 Feb 1937, Sperry 509; 

Tornillo Creek at Hot Springs, 26 Mar 1939, Sperry 1651; Maravillas Crossing, 02 Apr 

1938, Warnock T-282; hills at mouth of Santa Elene Canyon, 16 Mar 1947, Warnock 

47033; Dog Canyon, 16 Mar 1947, Warnock 47057] 

Lithospermum incisum Lehm. 

[Sperry, 1938; Johnston, 1967} 

[Brewster Co.: Chisos Mts., 02 Aug 1931, Muller 8278; Dog Flat, 10 Apr 1956, Warnock 

13884] 

Lithospermum multiflorum Torr. ex Gray 


[Brewster Co.: trail to Laguna on S side of Basin, 21 Jul 1957, Correll & Johnston 

18290; Boot Springs, 21 Jul 1957, Correll & Johnston 18297, 15-18 Jul 1921, Ferris 

& Duncan 2792 Chisos Mts., 06 Jul 1931, Muller 8137; et al. in Johnston, 1967] 

Lithospermum viride Greene 


[Brewster Co.: lower Pine Canyon, Chisos Mts., 12 Jul 1937, Warnock 185; Nail Place, 

Chisos Mts., 17 Aug 1915, Young s.n.; Pulliam Canyon and S slope Pulliam Bluff (Sperry, 

1938)] 

Omphalodes aliena Gray 


[Brewster Co.: Santa Elena Canyon, ledges, 19 Mar 1941, Warnock 21579; Dog Canyon, 

08 Apr 1958, Warnock & Johnston 15991] 

Tiquilia canescens (DC.) A. Richards. var. canescens Gray Coldenia 

[Sperry, 1938; Wells, 1965; Johnston, 1967; Richardson, 1977] 

[Brewster Co.: N side of Chisos Mts., 27 Jun 1931, Moore & Steyermark 3284; 

near Glenn Springs, 03 May 1937; Sperry 565; Dead Horse Mts., 08 Jun 1937, 

Warnock 1230; Paint Gap Hills, 05 Aug 1955, Warnock 12949 (UTEP)] 

Tiquilia gossypina (Woot. & Standl.) A. Richards. 

[Johnston, 1961, 1967] 

[Brewster Co.: Boquillas, 16 Apr 1919, Hanson 578; N of Chisos Mts., 21 Jun 1936, 

Sperry 158; Dead Horse Mts., 8-9 Jun 1937, Warnock 909; Tornillo Flats near Hulen 

Ranch, 09 Jun 1937, Warnock 1134; head of Boquillas Canyon, 10 Aug 1955, Warnock 

12899] 

Tiquilia greggii (Torr. & Gray) A. Richards. Plume Coldenia 

[Ptilocalyx greggii Torr.] 

[Steyermark & Moore, 1933; Sperry, 1938; Johnston, 1967; Richardson, 1977]

[Records: 14 mi. N of Boquillas, 22 May 1901, Bailey 338; 25 mi. N of Boquillas, 

17 Apr 1919, Hanson 594; 10 mi. SE Persimmon Gap, 25 Aug 1949, Hinckley & Hinckley 

274; between Persimmon Gap and Dog Flats, 30 Jul 1938, Sperry 1404; Persimmon Gap, 

16 Jun 1931, Tharp 8586; Blue Creek, 11 May 1955, Wallmo 39; near McKinney Springs, 

Dead Horse, Mts., 19 Jul 1952, Warnock 10800; et al.; hills about Chisos Mts. and Hot 

Springs (Sperry, 1938); Boquillas Canyon, Moore & Steyermark 3452] 

Tiquilia hispidissima (Torr. & Gray) A. Richards. Rough Coldenia 


[Records: near mouth of Terlingua Creek, 22 Jul 1957, Correll & Johnston 18359; 

Boquillas, 17 Apr 1919, Hanson 658; Boquillas Canyon, 12 Jul 1931, Moore & Steyermark 

3447; between Hot Springs and Boquillas, 29 Jun 1941, Warnock 21692; near mouth of 

Boquillas Canyon, 22 Jul 1952, Webster 4487] 

Tiquilia mexicana (S. Wats.) A. Richards. Mexican Coldenia 

[Sperry, 1938; Johnston, 1961, 1967; Richardson, 1977] 

[Records: Tornillo Flats, 08 Jun 1936, and Persimmon Flats, 06 Sep 1950, Warnock 

9638; N of Mariscal Mt., near Lindsey Mine, 25 May 1955, Warnock 12284] 

BRASSICACEAE (Mustard Family) 

Cardamine macrocarpa Brandegee var. texana Rollins 

[Rollins, 1940] 

[Brewster Co.: Chisos Mts., Camp Mountain, Sep 1933, Cory 7141 (GH-Type); Chisos Mts., 

Apr 1936, Cory 18670 (GH), July 1931, Mueller 8009 (GH), Oct 1937, Cory 26904 (GH), 

May 1938, Palmer 34097 (GH, NY), Jun 1931, Moore & Steyermark 3166 (GH, NY), Mar 

1937, Sperry 612 (US), Jun 1937, Warnock 920 (US)] 

Descurainia pinnata (Walt.) Britt. var. pinnata Tansy Mustard 

[Sperry, 1938; Detling, 1939] 

[Brewster Co.: Chisos Mts. (Sperry, 1938); Hot Springs, Sperry 532 (US); Tornillo Flats, 

Sperry 537 (US)] 

Descurainia incana (Benth. ex Fisch. & C. A. Mey.) Dorn. 

var. viscosa (Rydb.) Kartesz & Gandhi 

[Descurainia richardsonii O. E. Schulz var. viscosa (Rydb.) Detling] 


[Brewster Co.: upper Green Gulch (Sperry, 1938); Chisos Mts. (Correll & Johnston, 

1970)] 

Draba cuneifolia Nutt. ex Torr. & Gray var. cuneifolia Whitelow Grass 

[Sperry, 1938; Hartman, Bacon & Bohnstedt, 1975, mapped] 

[Records: Tornillo Flats, Green Gulch and Tornillo Creek bed at Hot 

Springs (Sperry, 1938)] 

Halimolobos diffusa (Gray) Schulz var. diffusa Spreading Fissurewort 

[Sperry, 1938; Rollins, 1943] 

[Brewster Co.: Chisos Mts. (Sperry, 1938)Chisos Mts., Mueller 8008 (GH, US)] 

Lepidium austrinum Small 

[Lepidium lasiocarpum Nutt. var. orbiculare (Thellung) C. L. Hitchc.] 


[Brewster Co.: from Persimmon Gap across Tornillo Flats (Sperry, 1938)] 

Lepidium montanum Nutt. var. angustifolium C. L. Hitchc. 


[Brewster Co.: near Santa Elena Canyon (Sperry, 1938)] 

Lesquerella argyraea (Gray) S. Wats. 


[Brewster Co.: Tornillo Flats (Sperry, 1938)] 

Lesquerella fendleri (Gray) S. Wats. Fendler Bladderpod 


[Brewster Co.: Neville Spring (Sperry, 1938)]

Lesquerella purpurea (Gray) S. Wats. Rose Bladderpod 

[Steyermark & Moore, 1933; Sperry, 1938; Lewis, Stripling & Ross, 1962] 

[Brewster Co.: Green Gulch, Divide and Oak Creek Canyon near Window (Sperry, 

1938); Chisos Mts., Moore & Steyermark 3197; BBNP, Ross 50 (BRIT; TEX-LL)] 

Nerisyrenia camporum (Gray) Greene Mesa Greggia 

[Sperry, 1938; Rollins, 1939; Bacon, 1978] 

[Records: near Rio Grande, Tornillo Flats and base of Lone Mtn. (Sperry, 

1938); BBNP, Brenckle & Brenckle 51072 (BRIT); Juniper Canyon, Marsh 269 (GH; TEX-LL); 

0.3 mi. S of Persimmon Gap, McVaugh 7834 (BRIT; TEX-LL); Dog Canyon Flats, Warnock 

47063 (BRIT; TEX-LL); 8-10 mi. S of Dagger Flat, Miller & Miller 1276 (BRIT); Oak Creek, 

Rollins & Correll 61810 (GH; TEX-LL); mouth of Terlingua Creek, Correll & Johnston 18352 

(GH; TEX-LL); Castolon Ranger Station, Bacon & Hartman 1437 (TEX-LL); mouth of Santa 

Elena Canyon, Shinners 8796 (BRIT); 1 mi. SW of rd. from Santa Elena Canyon and cutoff to 

Terlingua, Webster 4395 (BRIT); arroyo W of Johnson Ranch, Chisos Mts., Marsh 49 (TEX-LL); 

BBNP, Sep 1938, Rollins & Chambers 2767; near Persimmon Gap, Apr 1931, McKelvey 1972] 

Pennellia micrantha (Gray) Nieuwl. 

[Heterothrix micrantha (Gray) Rydb.] 


[Brewster Co.: upper Basin near Saddle (Sperry, 1938); Chisos Mts., Mueller 8004 (F, TEX-LL)] 

Rorippa nasturtium-aquaticum (L.) Hayek 

[Greene, 1962, mapped] 

Rorippa ramosa Rollins 

[Stuckey, 1972, mapped] 

Rorippa teres Michx. 

[Rorippa walteri (Ell.) C. Mohr] 

[Rollins, 1961; Stuckey, 1972, mapped] 

[Records: Santa Elena Canyon, Goodman & Waterfall 4653 (GH)] 

Schoenocrambe linearifolia (Gray) Rollins Mountain Mustard 


[Brewster Co.: Green Gulch, Oak Creek Canyon, lower Cattail Falls and 

near top of Casa Grande (Sperry, 1938); Basin at Saddle above Green Gulch, 

BBNP, 15 Jul 1981, Freeman s.n. (UTEP); Boot Spring, Chisos Mts., 21 Jul 1957, 

Correll & Johnston 18300 (GH)] 

Selenia dissecta Torr. & Gray Texas Selenia 



Sisymbrium irio L. London Rocket 


[Brewster Co.: Tornillo Creek bed at Hot Springs (Sperry, 1938)] 

Streptanthus carinatus Wright ex Gray ssp. carinatus Lyerleaf Twistflower 


[Brewster Co.: Avary Canyon (Sperry, 1938)] 

Streptanthus cutleri Cory 

[Brewster Co.: 0.7 mi. W of tunnel near Rio Grande Village, BBNP, 18 Mar 1985, 

Van Devender s.n. (UTEP)] 

Note: The type locality is 4 mi. NE of Black Gap in Maravillas Canyon (Cory, 1943; 

Type: GH collected by Hugh Cutler). 

Synthlipsis greggii Gray 

[Rollins, 1959] 

[Records: 44 mi. SE of Marathon at Maravillas Creek, Cory 31618 (GH); between Persimmon 

Gap and Dog Flat, Warnock C293 (GH)] 

Thelypodium texanum (Cory) Rollins Texas Thelypody 

[Sisymbrium texanum Cory] 

[Cory, 1937; Sperry, 1938; Al-Shehbaz, 1973] 

[Brewster Co.: near Hot Springs (Sperry, 1938); Tornillo Creek near Rio Grande Hot Springs, 

Goodman & Waterfall 4609 (COLO, GH, TEX-LL, UC); Cottonwood Creek, BBNP, Rollins

& Correll 6184A (GH); 18 mi. N of Terlingua on floodplain of Terlingua Creek, 13 Apr 1936, 

Cory 18564 (GH-type)] 

Thelypodium wrightii Gray ssp. wrightii Wright Thelypody 

[Stanleyella wrightii (Gray) Rydb.] 

[Sperry, 1938; Al-Shehbaz, 1973] 

[Brewster Co.: Wade Canyon (Sperry, 1938); Chisos Mts., Mueller 8003 (F, NY, TEX-LL)] 

BUDDLEJACEAE 

Buddleja marrubiifolia Benth. Woolly Butterflybush 

[Steyermark & Moore, 1933; Sperry, 1938; Wells, 1965; Powell, 1998] 

[Brewster Co.: Dead Horse Mts. (Wells, 1965); Hot Springs near Santa Elena 

Canyon (Sperry, 1938); in small ravine N of Boquillas Canyon, Moore & Steyermark 

3449 (MO)] 

Buddleja scordioides Kunth Escobilla Butterflybush 


Emorya suaveolens Torr. Emorybush 

[Warnock, 1964; Norman, 1964; Powell, 1998] 

[Brewster Co.: Expected:Black Gap Refuge, lower Maravillas Creek near Indian 

Cave, 17 Nov. 1962, Rogers 256 (SRSC)] 

CACTACEAE (Cactus Family) 

Ariocarpus fissuratus (Engelm.) K. Schum. Living Rock Cactus 

[Sperry, 1938; Anderson, 1963; Wells, 1965; Benson, 1969; Evans, 1998] 

[Brewster Co.: S of Headquarters, BBNP, Anderson 941; Tornillo Creek, Anderson 

933, 935, 20 Jun 1952, Blakley B1425; Dead Horse Mts. (Wells, 1965); hills N and 

W of Chisos Mts. (Sperry, 1938); 19 mi. E of Panther Junction, 2050 ft., Anderson 933 

(RSA-POM); ] 

Coryphantha albicolumnaria (Hester) A. D. Zimmerman 

[Escobaria albicolumnaria Hester; Coryphantha sneedii (Britt. & Rose) Berger 

var. albicolumnaria (Hester) A. Zimmerman nomen in edit.] 

[Benson, 1969; Worthington, 1993; Evans, 1998] 

[Brewster Co.: Mariposa Mine area, Schwenin 4138 (UNM), Pierce 4250 (UNM), 

Weniger 146, 147 (UNM); Mariscal Mtn., Smith 2211 (SRSC); Boquillas Canyon, 

Zimmerman 1541 (BBNP Herb.); dry wash 1 mi. W of Boquillas Canyon, Wauer s.n. 

(RSA)] 

Coryphantha cornifera (DC.) Lem. var. echinis (Engelm.) L. Benson Sea Urchin Cactus 

[Sperry, 1938; Benson, 1969; Evans, 1998] 

[Brewster Co.: Dead Horse Mts., Warnock 11061] 

Coryphantha dasyacantha (Engelm.) Orcutt. 

var. dasyacantha Mountain Cob Cactus 

[Escobaria dasyacantha (Engelm.) Britt. & Rose var. dasyacantha] 

[Benson, 1969; Evans, 1998] 

[Brewster Co.: Panther Pass, Chisos Mts., 16 Jul 1955, Benson & Benson 15511] 

var. duncanii (Hester) N. P. Taylor Duncan Cory Cactus 

[Coryphantha duncanii (Hester) L. Benson] 

[Benson, 1969; Weedin, Powell & Kolle, 1989; Evans, 1998] 

[Brewster Co.: BBNP, 1969, Wauer s.n.; BBNP, 2n=22, Weedin 1270 (SRSC)] 

Coryphantha macromeris (Engelm.) Lem. var. macromeris Long Mamma 


[Brewster Co.: Paint Gap Hills, BBNP, Warnock 12832; Tornillo Creek desert, BBNP, 

21 Jun 1952, Riley & Marshall B1505; Rio Grande near Hot Springs (Sperry, 1938)] 

Coryphantha ramillosa Cutak Big Bend Cory Cactus 

[Wells, 1965; Evans, 1998]


Coryphantha strobiliformis (Poselger) Orcutt 

[Escobaria tuberculosa (Engelm.) Britt. & Rose; Coryphantha dasyacantha (Engelm.) 

orcutt var. varicolor (Tiegel) L. Benson] 

[Sperry, 1938; Wells, 1965; Benson, 1969] 

[Brewster Co.: E of Hot Springs, 20 Apr 1948, Whitehouse 1907; Chisos Mts., 06 Jul 

1932, Mueller s.n.; W of Oak Creek Ranch house, BBNP, 02 Jul 1955, Wallmo 300; 

Dead Horse Mts. (Wells, 1965)] 

Coryphantha vivipara (Nutt.) Britt. & Rose var. radiosa (Engelm.) Backeberg Spiny Star 

[Escobaria vivipara (Nutt.) Buxbaum var. radiosa (Engelm.) D. R. Hunt] 


[Brewster Co.: Mt. Emory, Chisos Mts., 19 Jun 1952, Blakley s.n.; Basin (Sperry, 1938)] 

Echinocactus horizonthalonius Lem. var. horizonthalonius Turk’s Head Cactus 

[Sperry, 1938; Wells, 1965; Evans, 1998] 

[Brewster Co.: Dead Horse Mts. (Wells, 1965); flats around Chisos Mts. (Sperry, 1938)] 

Echinocactus texensis Hopffer Horse Crippler 

[Evans, 1998] 

Echinocereus chisoensis W. T. Marsh Chisos Mountain Hedgehog 

[Echinocereus reichenbachii (Terscheck ex Walp.) Haage f. var. chisoensis 

(W. T. Marsh) L. Benson] 

[Sperry, 1938; Benson 1969; Evans, 1998] 

[Brewster Co.: N of Boquillas Canyon, 06 May 1963, Weniger 126;N of Hot Springs, 20 Jun 

1952, Blakley B1423; lower Tornillo Creek, 11 Apr 1965, Benson 16505; San Vicente-Glenn 

Spring Road, 13 Apr 1941, Sperry 1907; Chisos Mts., Jun 1950, Hester s.n.] 

Echinocereus chloranthus (Engelm.) Haag f. 

var. chloracanthus 

[Benson, 1969; Evans, 1998] 

[Brewster Co.: San Vicente-Glenn Springs, 13 Apr 1941, Sperry 1907; Panther Pass, 

11 Apr 1965, Benson 16513; Chisos Mts., Jun 1950, Hester s.n.] 

var. cylindricus (Engelm.) N.P. Taylor 

[Echinocereus viridiflorus (Engelm.) Engelm. var. cylindricus Engelm.] 

[Benson, 1969; Weniger, 1969] 

[Brewster Co.: Chisos Mts., 1924, Ferris s.n.] 

var. russanthus (Weniger) Lamb ex Rowley 

[Weniger, 1969; Weedin, Powell & Kolle, 1989] 

[Brewster Co.: Green Gulch, BBNP, n=11, Kolle 46 (SRSC); 24 km S from S entrance of 

Old Ore Road, BBNP, n=11, Kolle 54 (SRSC); northern part of Chisos Mts. (Weniger, 1969)] 

Echinocereus coccineus Engelm. Claret-cup 

var. coccineus 

[Benson, 1969; Evans, 1998] 

[Brewster Co.: Panther Pass, 12 Apr 1965, Benson 16512; Lost Mine to Green 

Gulch, 12 Apr 1941, Sperry A03; Casa Grande, Moore 308, 311] 

var. arizonicus (Rose ex Orcutt) Ferguson 

[E. triglochidiatus Engelm. var. neomexicanus auct non (Standl.) W. T. Marsh] 

[Benson, 1969; Evans, 1998] 

[Brewster Co.: Lost Mine Mt., Chisos Mts., 12 Apr 1941, Sperry 1903; Glenn 

Spring, 12 Apr 1941, Sperry 1905] 

Echinocereus dasyacanthus Engelm. Texas Rainbow 

[Echinocereus pectinatus (Scheidw.) Engelm. var. neomexicanus 

(Coulter) L. Benson; E. steereae Clover] 

[Sperry, 1938; Clover, 1938; Wells, 1965; Benson, 1969; Evans, 1998] 

[Brewster Co.: 21 mi. s of entrance to BBNP, Fields, McBryde & King 2578; 

Chisos Mts., Jun 1950, Hester s.n., 13 Apr 1941, Sperry 1904; Persimmon Gap, 

02 May 1937, Sperry 2063; Dead Horse Mts. (Wells, 1965); Pulliam Bluff (Sperry, 

1938); Willow Springs Site, BBNP, 29 DEG 18.56’N, 103 DEG 02.119’W, 990 m, 

05 Apr 1995, Hodgson 8950 (UTEP; DES); Chisos Mts., Mar 1937, Steere (type of

E. steereae Clover)] 

Echinocereus enneacanthus Engelm. var. enneacanthus 

[Echinocereus dubius (Engelm.) Engelm. ex Rumpl.] 

[Sperry, 1938; Moore, 1967; Benson, 1969; Weedin, Powell & Kolle, 1989; Evans, 1998] 

[Records: Boquillas Canyon Road, Moore 671 (MICH); Santa Elena Canyon Road, 

Moore 290, 294, 305, 307, 683, 684, 689 (MICH); Boquillas Canyon, Moore 326; 5 mi. S 

Of Panther Junction, Moore 680, 681 (MICH); Glenn Springs, 12 Apr 1946, Sperry 1905; 

Costolon, BBNP, n=11, Weedin 437 (SRSC)] 

Echinocereus pectinatus (Scheidw.) Engelm. var. pectinatus 

[Benson, 1968] 

[Brewster Co.: road to Boquillas Canyon, BBNP, Benson 16904 (observed)] 

Echinocereus pseudopectinatus (N. P. Taylor) N. P. Taylor 

[Echinocerus pectinatus (Scheidw.) Engelm. var. minor auct non (Engelm.) 

L. Benson] 

[Benson, 1968, 1969] 

[Brewster Co.: lower Tornillo Creek, 11 Apr 1965, Benson 16504 (RSA-POM)] 

Echinocereus stramineus (Engelm.) F. Seitz. Strawberry Cactus 

[Sperry, 1938; Wells, 1965; Moore, 1967; Benson, 1969; Evans, 1998] 

[Records: Rt. 385, 21 mi. S entrance to BBNP, King 2577; Hot Springs, 20 Apr 

1948, Whitehouse 19704; Boquillas Canyon Road, Moore 668, 669, 670 (MICH); Santa Elena 

Canyon, Moore 301 (MICH); E edge of Chisos Mts., Cutler 761; Hot Springs, Jones 26310 

(US); 2 mi. from Boquillas Canyon, Moore 672 (MICH); Dead Horse Mts. (Wells, 1965)] 

Epithelantha bokei L. Benson Boke Button Cactus 

[Epithelantha micromeris (Engelm.) A. Weber ex Britt. & Rose var. bokei (Benson) 

Glass & Foster] 

[Benson, 1969; Glass & Foster, 1978; Evans, 1998] 

[Records: near Boquillas Canyon, 12 Jul 1955, Boke s.n.(RSA-POM, type)] 

Epithelantha micromeris (Engelm.) A. Weber ex Britt. & Rose Common Button Cactus 

[Epithelantha micromeris (Engelm.) Weber ex Britt. & Rose var. micromeris] 

[Sperry, 1938; Benson, 1969; Glass & Foster, 1978; Evans, 1998] 

[Brewster Co.: Chisos Mts. between Boot Spring and South Rim, Warnock 12705; 

Juniper Canyon, Chisos Mts., Ferris & Duncan 3591; near top of Crown Mtn. (Sperry, 

1938)] 

Ferocactus hamatacanthus (Muhlenpfordt) Britt. & Rose Texas Barrel Cactus 

var. hamatacanthus 


[Brewster Co.: N of Santa Elena, 16 Jul 1955, Benson & Benson 15507; Chisos 

Mts., 22-24 Nov 1922, Pilsbry s.n., Warnock 12393, 12394; Pinnacle Mts. and 

Boquillas (Sperry, 1938)] 

Lophophora williamsii (Lemaire) Coulter Peyote 

[Sperry, 1938} 

[Brewster Co.: Chilocital Mtn. (Sperry, 1938)] 

Mammillaria heyderi Muehlenpfordt Heyder Pincushion Cactus 

var. heyderi 

[Mammillaria gummifera Engelm. var. applanata (Engelm.) L. Benson] 


[Brewster Co.: Chisos Mts., 22-24 Nov 1922, Pilsbry s.n.] 

var. meiacantha (Engelm.) L. Benson 

[Mammillaria gummifera Engelm. var. meiacantha (Engelm.) L. Benson] 


[Brewster Co.: Panthe Pass, Chisos Mts., 16 Jul 1955, Benson & Benson 15510; 

junction to Panther Pass, 16 Jul 1955, Benson & Benson15501; Boot Spring Canyon, 

Warnock 12394; Dead Horse Mts., Warnock 11065] 

Mammillaria lasiacantha Engelm. Lacespine Cactus 

[Sperry, 1938; Evans, 1998] 

Mammillaria pottsii Scheer ex Salm-Dyck Potts Mammillaria

[Sperry, 1938; Evans, 1998] 

Neolloydia conoidea (DC.) Britt. & Rose Texas Cone Cactus 


[Brewster Co.: Daggar Flats, 22 Jun 1952, Blakley 31449; Dead Horse Mts., Warnock 

11055, 11058] 

Opuntia aggeria Ralston & Hilsenbeck Clumped Dog Cholla 

[Opuntia grahamii X O. schottii] 

[Ralston & Hilsenbeck, 1989; Powell, 1998] 

[Brewster Co.: on Tornillo Flats, BBNP, 2800 ft., 30 Jul 1948, M. Anthony 856 

(MICH:Type); 19 km E of Costolon on River Road, BBNP, Ralston 135 (SRSC); 

43.4 km E of Costolon on River Road, BBNP, Ralston 135 (SRSC); 33 km E of 

Costolon on River Road, BBNP, Ralston 152 (SRSC); 8.4 km N of Santa Elena 

Canyon on Maverick Rd., BBNP, Ralston 120 (SRSC); 22 km SE Panther Junction, 

BBNP, Ralston 118 (SRSC); Boquillas Canyon crossing area, BBNP, Ralston 136 

(SRSC); 15.3 mi. W Mariscal Mtn.., BBNP, Powell 5216 (SRSC);Big Bend Natl. 

Park “the predominant dog cholla” (Powell, 1998)] 

Opuntia atrispina Griffiths 

[Benson, 1969; Powell, 1998] 

[Brewster Co.: Chisos Mts., Anthony 764] 

Note: Powell (1998) says it may occur no closer to BBNP that Terrell and Val Verde 

Counties. 

Opuntia aureispina (Brack & Heil) Pinkava & Parfitt Golden-spined Pricklypear 

[Evans, 1998; Powell, 1998] 

[Brewster Co.: near Mariscal Mtn. to Boquillas Canyon (Powell, 1998)] 

Opuntia chisosensis (Anthony) Ferguson Texas Pricklypear 

[Opuntia lindheimeri Engelm. var. chisosensis Anthony] 

[Evans, 1998; Powell, 1998] 

[Brewster Co.: wooded slopes in the Chisos Mts. (Powell, 1998)] 

Opuntia densispina Ralston & Hilsenbeck Dense Dog Cholla 

[Ralston & Hilsenbeck, 1992; Powell, 1998] 

[Brewster Co.: BBNP, 5.3 mi. NE Solis’ Ranch on Old River Road, 15 May 1989, 

Ralston 200 (Type: SRSC; Isotypes: TEX-LL, ASC, SRSC] 

Opuntia engelmannii Engelm. Engelmann’s Pricklypear 

var. engelmannii 

[Opuntia phaeacantha Engelm. var. discata (Griffiths) Benson & Walkington] 

[Benson, 1969; Evans, 1998] 

[Brewster Co.: Chisos Mts., Anthony 54, 825, 1937, Cutler 960; Hot Springs, 

Anthony 53, 55] 

var. lindheimeri (Engelm.) Parfitt & Pinkava 

[Benson, 1969; Weedin, Powell & Kollw, 1989; Powell, 1998] 

[Brewster Co.: Solis Ranch, BBNP, 15 Apr 1947, Anthony 33b, 27 Jun 1954, 

Anthony 6; Boquillas area of BBNP (Powell, 1998); Rio Grande Village, BBNP, 

2n=66, Weedin 1171 (SRSC)] 

var. linguiformis (Griffiths) Parfitt & Pinkava Cow’s Tongue Pricklypear 

[Evans, 1998] 

Opuntia grahamii Engelm. Graham Club Cholla 

[Sperry, 1938; Benson, 1969; Weedin & Powell, 1978; Ralston & Hilsenbeck, 1989; 

Evans, 1998; Powell, 1998] 

[Brewster Co.: Solis Ranch, BBNP, Anthony 27; Costolon, Anthony 11; Old Ore 

Rd. near La Noris, BBNP, Weedin & Weedin 237 (SRSC)] 

Opuntia imbricata (Haw.) DC. Tree Cholla; Cane Cholla 

var. imbricata 

[Benson, 1969; Evans, 1998] 

[Brewster Co.: Boot Springs Canyon, Chisos Mts., Warnock 12386] 

var. argentea Anthony Silverspine Cholla 

[Benson, 1969; Evans, 1998; Powell, 1998]

[Brewster Co.: near Solis Ranch house, BBNP, Anthony 276, 29 Apr 1948, Anthony 

280 (Type), 27 Jun 1954, Anthony 12; near Mariscal Mtn. (Powell, 1998)] 

Opuntia kleiniae DC. Candle Cholla; Klein Cholla 

[Benson, 1969; Weedin, Powell & Kolle, 1989; Evans, 1998; Powell, 1998] 

[Brewster Co.: Solis Ranch house near the Rio Grande, BBNP, 28 Apr 1948, Anthony 

275; Costolon, BBNP, 19 Apr 1948, Whitehouse 19687; near Costolon, BBNP, n=24, 

Weedin 1256 (SRSC)] 

Opuntia leptocaulis DC. Pencil Cholla; Tesajo 

[Sperry, 1938; Wells, 1965; Benson, 1969; Evans, 1998] 

[Brewster Co.: Paint Gap Hills, BBNP, Warnock 12918; Dead Horse Mts. (Wells, 

1965)] 

Opuntia macrocentra Engelm. Purple Pricklypear 

[Sperry, 1938; Wells, 1965; Evans, 1998] 

[Brewster Co.: Sue Peak (Wells, 1965); desert surrounding Chisos Mts. (Sperry, 

1938)] 

Opuntia macrorhiza Engelm. var. macrorhiza Plains Pricklypear 

[Benson, 1969; Evans, 1998] 

[Brewster Co.: Hot Springs, 12 Apr 1941, Sperry 1910; Chisos Mts., Nov 1963, 

Weniger 414; road mile 11 on R Maxwell Scenic Drive, BBNP, 05 Apr 1995, 

29 DEG 11’N, 103 DEG 22.5’W, 1066 m, Hodgson 9023 (UTEP, DES)] 

Opuntia phaeacantha Engelm. Brownspine Pricklypear 

[Opuntia phaeacantha Engelm. var. major Engelm.] 

[Benson 1969; Weedin, Powell & Kolle, 1989] 

[Brewster Co.: 25 mi. S of Government Springs, 13 Apr 1948, Whitehouse 19582; 

Santa Elena Canyon, Anthony 1209; Willow Springs Site, BBNP, 29 DEG 18.560’ 

N, 103 DEG 02.119’W, 990 m, 05 Apr 1995, Hodgson 8952, 8968 (UTEP, DES); 

W of Johnson ranch house, BBNP, 2n=66, Weedin 430 (SRSC)] 

Opuntia rufida Engelm. Blind Pricklypear 

[Sperry, 1938; Wells, 1965; Benson, 1969; Weedin, Powell & Kolle, 1989; Evans, 1998] 

[Brewster Co.: Boquillas Canyon Lookout, 11 Apr 1965, Benson 16506; Boquillas, 

1901, Bailey s.n., 11 Apr 1948, Anthony 273; 17 mi. SE Panther Junction, road to 

Boquillas Canyon, Anderson 936, Travers 2272, 2273; E side of Chisos Mts., 20 Jun 

1952, Blakley B-1427; Hot Springs, BBNP, 18 Apr 1947, Anthony 39a, 39b, 39c, 39d, 

39e (39c is type of var. tortiflora); Chilicotal Mt., BBNP, 03 Apr 1947, Anthony 16, 17; 

Dagger Flat, Anthony 898-899; Dead Horse Mts. (Wells, 1965); River Road, Black Dike 

At Smoky Creek, BBNP, n=11, Weedin 434 (SRSC)] 

Opuntia santa-rita (Griffiths & Hare) Rose 

[Opuntia violacea Engelm. var. santa-rita (Griffiths & Hare) Benson] 


[Brewster Co.: Glenn Springs, 13 Apr 1941, Sperry 1911; Santa Elena Canyon, 16 Jul 

1955, Benson & Benson 15504; Costolon, 1937, Cutler 763; 25 mi. S of Government 

Springs, 13 Apr 1948, Whitehouse 19582; Costolon to Santa Elena Canyon, 29 Jun 

1954, Anthony 11] 

Opuntia schottii Engelm. Scott Club Cholla 

[Benson, 1969; Weedin & Kolle, 1989; Ralston & Hilsenbeck, 1989; Weedin, Powell 

& Kolle, 1989; Evans, 1998] 

[Brewster Co.: Tornillo Creek near Chisos Mts., 22 May 1928, Palmer 43961; 

jct. Old Ore Rd. and Ernst Tinaja Rd., BBNP, Kolle & Weedin 53 (SRSC); Boquillas 

Canyon, 2n=22, Weedin 1152 (SRSC); jct. Old Ore Road and Ernst Tinaja Road, 2n=22, 

Kolle 53 (SRSC)] 

Opuntia spinosibacca Anthony Spinyfruited Prickly Pear 


[Brewster Co.: between Mariscal Mtn. And Boquillas Canyon (Powell, 1998)] 

Peniocereus greggii (Engelm.) Britt. & Rose Nightblooming Cereus 

[Cereus greggii Engelm.] 

[Sperry, 1938, as Acanthocereus pentagonus (L.) Britt. & Rose (?); Benson, 1969]

[Brewster Co.: Ward Mt., Chisos Mts., Warnock 13075; W slope Dead Horse 

Mts. (Sperry, 1938)] 

Sclerocactus intertextus (Engelm.) N. P. Taylor Chihuahuan Pineapple Cactus 

[Echinomastus intertextus Engelm.] 

[Sperry, 1938; Evans, 1998] 

Sclerocactus mariposensis (Hester) N. P. Taylor Mariposa Cactus 

[Neolloydia mariposensis (Hester) L. Benson; Echinomastus mariposensis Hester] 


[Brewster Co.: near Mariposa, 06 May 1963, Weniger 148; Santa Elena Canyon Road, 

21 Jun 1952, Marshall 1428b, 21 Jun 1952, Blakley 1428a] 

Sclerocactus uncinatus Galeotti) N. P. Taylor var. wrightii (Engelm.) N. P. Taylor 

[Ancistrocactus uncinatus (Galeotti) L. Benson var. wrightii (Engelm.) L. Benson] 

[Sperry, 1938, as Ancistrocactus sheeri (Salm-Dyck) Britt. & Rose; Wells, 1965; 

Benson, 1969; Evans, 1998] 

[Brewster Co.: 15 mi. S entrance to BBNP, King 2581; Dead Horse Mts. (Wells, 1965); 

flats E of Burro Mesa (Sperry, 1938)] 

Note: Evans (1998) is using he name Glandulocactus uncinatus var. wrightii. 

Sclerocactus warnockii (L. Benson) N. P. Taylor Warnock Cactus 

[Neolloydia warnockii L. Benson; Echinomastus warnockii (Benson) Glass & Foster] 


[Brewster Co.: SW of Persimmon Gap W of BBNP, 12 Apr 1965, Benson 16514 (RSA-POM, 

type); opposite Boquillas, 11 Apr 1965, Benson 16508; Dog Canyon below Persimmon Gap, 16 

Mar 1947, Warnock & Buechner 47-012; between Hot Springs and Hannold’s, 23 Feb 1941, 

Sperry 1818; 15 mi. NW of Boquillas, 06 May 1963, Weniger s.n.; road to Boquillas, 13 Jul 1955 

Boke s.n.] 

Thelocactus bicolor (Galeotti ex Pfeiffer) Britt. & Rose Glory of Texas 

[Sperry, 1938; Weedin, Powell & Kolle, 1989; Evans, 1998] 

[Brewster Co.: Babb’s Hut in the Pinnacle Mts. (Sperry, 1938); W of Old McKinney 

Ranch House, BBNP, 2n=22, Kolle 56 (SRSC)] 

CAMPANULACEAE (Bellflower Family) 

Lobelia cardinalis L. Cardinal Flower 

[Lobelia splendens Humb. & Bonpl. ex Willd.] 


[Brewster Co.: lower Oak and Cattail Canyons (Sperry, 1938); Cattail Falls, BBNP, 

Aug 1983, Stafford s.n. (UTEP)] 

Note: Sperry (1938) reports Lobelia cliffortiana L. from Fresno Springs and Fisk 

Canyon. Correll & Johnston (1970) do not report this species from Texas. 

CAPPARIDACEAE (Caper Family) [=CAPPARACEAE] 

Cleomella longipes Hook. 

[Sperry, 1938; Iltis, 1956] 

[Records: along Rio Grande near Costolon (Sperry, 1938); Chisos Mts. area in 

ravine of Rio Grande, Sperry 136 (US)] 

Koeberlinia spinosa Zucc. Allthorn; Crown-of-thorn; Junco 

[Sperry, 1938; Muller, 1941, mapped; Wells, 1965] 

[Records: Dead Horse Mts. (Wells, 1965); Gano Spring and flats adjacent 

to the Chisos Mts. (Sperry, 1938)] 

Polanisia uniglandulosa (Cav.) DC. Mexican Clammyweed 


[Records: upper Green Gulch, Oak Creek Canyon and Fresno Spring (Sperry, 

1938)]

CAPRIFOLIACEAE (Honeysuckle Family) 

Lonicera albiflora Torr. & Gray White Honeysuckle 


[Brewster Co.: Dead Horse Mts. (Wells, 1965); Chisos Mts. (Powell, 1998); 

Squaw Creek and Basin (Sperry, 1938)] 

Sambucus cerulea Raf. Blue Elderberry 


[Brewster Co.: upper Blue Creek Canyon (Sperry, 1938); on talus at base of cliff, 

Emory Peak, 17 Jun 1964, Correll 29746 (TEX-LL)] 

Symphoricarpos palmeri Jones 

[Jones, 1940] 

[Records: Chisos Mts., Palmer 34176 (MO; NY), Havard (US)] 

Symphoricarpos rotundifolius Gray Roundleaf Snowberry 


[Records: Boot Canyon (Sperry, 1938); upper Lost Mine Trail, BBNP, 

26 Jun 1978, Freeman s.n. (UTEP)] 

CARYOPHYLLACEAE (Pink Family) 

Arenaria benthamii Frenzl. ex Torr. & Gray 



Cerastium axillare Correll 

[Sperry, 1938, as Cerastium viscosum L. ?; Turner, 1995, mapped; Correll, 1996] 

[Records: near Chinese Wall (Sperry, 1938); above Lost Mine Peak Trail, Correll 20688-B 

(TEX-LL); Chisos Mts., Mueller 7996 (TEX-LL); Boot Springs, Parks & Cory 7134 (TAES); 

Boot Creek area, Wolff 4552 (TAES)] 

Cerastium brachypodum (Engelm. ex Gray) B. L. Robins. 


[Brewster Co.: Chisos Mts. (Sperry, 1938); Chisos Mts., Moore & Steyermark 3168] 

Paronychia jamesii Torr. & Gray James Nailwort 

[Sperry, 1938; Core, 1941; Correll, 1966; Turner, 1983, mapped] 

[Brewster Co.: Divide, Chinese Wall and Panther Spring (Sperry, 1938); North Sunny Glenn, 

17 Jul 1938, Sperry T683 (TAES); Chisos Mts., Mueller 7998b (MO, NY)] 

Paronychia monticola Cory 

[Paronychia nudata Correll] 

[Turner, 1983, mapped] 

Silene antirrhina L. Sleepy Catchfly 



Silene laciniata Cav. Mexican Silene 


[Brewster Co.: Pulliam Canyon, upper Blue Creek and E slope of Casa 

Grande (Sperry, 1938); Chisos Mts., Moore & Steyermark] 

Stellaria prostrata Baldw. Llovisna 



CELASTRACEAE (Staff-tree Family) 

Celastrus scandens L. Climbing Bittersweet 

[Lundell, 1969] 

[Brewster Co.: in canyons near Emory Peak, Chisos Mts., 25 May 1928, Palmer 34170] 

Mortonia sempervirens Gray ssp. scabrella (Gray) Prigge Rough Mortonia

[Wells, 1965] 


Schaefferia cuneifolia Gray Desert Yaupon; Capul 

[Sperry, 1938; Lundell, 1969; Powell, 1998] 

[Brewster Co.: Oak Canyon, Chisos Mts., 23 May 1928, Palmer 34114; near Glenn 

Spring, 28 Oct 1939, Sperry s.n.; Juniper Canyon, Chisos Mts., 25 Jun 1937, Warnock 

663; near Dominguez Spring, BBNP, 27 May 1955, Warnock & Wallmo 12305; Fresno 

Spring Creek and upper Juniper Canyon (Sperry, 1938)] 

CHENOPODIACEAE (Goosefoot Family) 

Atriplex acanthocarpa (Torr.) S. Wats. Armed Saltbush 

[Reed, 1969; Henrickson, 1988, mapped; Powell, 1998] 

[Brewster Co.: 3 mi. E of Study Butte, 26 Jun 1931, Moore & Steyermark 3552; 

just W of Maverick Mtn. Ranger Station, 11 Sep 1961, Correl & Johnston 24479; 

1.5 mi. N of Costolon, 28 Sep 1938, Park & Cory 30506; Avary Canyon, BBNP, 

23 Jul 1950, Warnock 9131; Santa Elena Canyon area, 12 May 1961, Traverse 2219; 

(more records in Reed, 1969); 5 mi. W Chisos Mts. (Powell, 1998)] 

Atriplex canescens (Pursh) Nutt. var. angustifolia (Torr.) s. Wats. Fourwing Saltbush 

[Sperry, 1938; Reed, 1969] 

[Brewster Co.: Nails Ranch near Chisos Mts., 11 Sep 1933, Wolff 4719; Chisos 

Mts., 18 Aug 1955, Warnock 12999; Santa Elena Canyon and vic., 12 May 1961, 

Traverse 2186, 2211; Grapevine Spring Rd., BBNP, 13 May 1961, Traverse 2234, 

2235; Grapevine Hills, Painted Gap, Blue Creek, Persimmon Gap, Gano Springs 

and Smoky Creek (Sperry, 1938)] 

Atriplex obovata Moq. Silver Saltbush 

[Reed, 1969; Powell, 1998] 

[Brewster Co.: Santa Elena Canyon, BBNP, 12 May 1961, Traverse 2219; Avary 

Canyon, BBNP, 23 Jul 1950, Warnock 9132] 

Chenopodium albescens Small Pale Goosefoot 

[Reed, 1969] 

[Brewster Co.: Boot Spring to Basin, Chisos Mts., 15 Jul 1955, Warnock 12740] 

Chenopodium album L. 


[Brewster Co.: lower Willow Creek and creek bed near Lone Mtn. (Sperry, 1938)] 

Chenopodium berlandieri Moq. var. berlandieri Pitseed Goosefoot 

[Sperry, 1938; Reed, 1969] 

[Brewster Co.: Chisos Mts. State Park and vic., 03 May 1937, Sperry 560; upper Green 

Gulch, Chisos Mts., 14 Sep 1941, Sperry 1962; Willow Creek, Basin, Chisos Mts., 15 Aug 

1957, Warnock 1118; Terrapin Tank and Willow Creek Canyon (Sperry, 1938)] 

Chenopodium fremontii S. Wats.Small 

[Reed, 1969] 

[Brewster Co.: Oak Canyon, Chisos Mts., 06 Sep 1933, Wolff 4449; Blue Creek Canyon, 

12 Sep 1932, Mueller s.n.; Chisos Mts., 06 Aug 1937, Marsh 192; Lost Mine Trail, Basin, 

Chisos Mts., 22 Jul 1957, Correll & Johnston 18328; Window Trail, Chisos Mts., 07 Jul 1944, 

Cory 44709; Chisos Mts. State Park, Jul 1935, Marsh 91] 

Chenopodium incanum (S. Wats.) Heller var. elatum Crawford 

[Reed, 1969; Crawford, 1977] 

[Records: Chisos Mts., 11 Sep 1915, Young s.n.; Oak Canyon, Chisos Mts., 19 Jun 1931, 

Tharp s.n.; Chisos Mts., near Love Mtn., 30 Jun 1957, Warnock 1208 (GH; TEX-LL)] 

Chenopodium incisum Poir. 

[Sperry, 1938; Reed, 1969] 

[Brewster Co.: Nail Place, Chisos Mts., 21 Aug 1915, Young s.n.; Chisos Mts., o4 Aug 

1931, Mueller 7980; Green Gulch, Chisos Mts., 31 Aug 1938, Warnock C644; Boot Canyon, 

Chisos Mts., 15 Jul 1955, Warnock 12721; Boot Creek area, 08 Sep 1933, Wolf 4591; Boot

Springs, 09 Sep 1933, Parks & Cory 7138; upper Green Gulch, Basin and Laguna (Sperry, 

1938)] 

Chenopodium murale L. Nettleleaf Goosefoot 

[Reed, 1969] 

[Brewster Co.: Chisos Mts., 04 Sep 1937, Marsh 322; Chisos Mts. State Park, 03 May 1939, 

Warnock 560; Glenn Springs, 25 May 1933, Parks & Cory 6530; 5 mi. S of Persimmon Gap, 

05 Mar 1947, McVaugh 7821] 

Chenopodium neomexicanum Standl. 

[Reed, 1969] 

[Brewster Co.: Chisos Mts., Willow Creek Basin, 15 Aug 1937, Warnock 1118] 

Corispermum hyssopifolium L. Shining Tickseed 

[Reed, 1969] 

[Brewster Co.: Boquillas Canyon on Rio Grande, 11 Jul 1931, Moore & Steyermark 

3430 and other collectors] 

Cycloloma atriplicifolium (Spreng.) Coulter 

[Reed, 1969] 

[Brewster Co.: St. Helena Canyon, Chisos Mts., 08 Aug 1938, Warnock T504C] 

Monolepis nuttalliana (J. A. Schultes) Greene 


[Brewster Co.: Tornillo Creek bed and Hot Springs (Sperry, 1938)] 

Salsola australis R. Br. Tumbleweed 

[Salsola kali L.; S. iberica Senn. & Pau; S. pestifer A. Nels.] 

[Reed, 1969] 

[Brewster Co.: Chisos Mts., 03 Sep 1937, Marsh 341] 

Suaeda suffrutescens S. Wats. var. suffrutescens Desert Seepweed 


[Brewster Co.: Santa Elena Canyon (SRSC)] 

var. detonsa Johnston 

[Hopkins & Blackwell, 1977] 

[Brewster Co.: Tornillo Creek, Hot Springs, 26 Mar 1939, Sperry 1637] 

CISTACEAE 

Helianthemum glomeratum (Lag.) Lag. ex Dunal 

[Sperry, 1938; Daoud & Wilbur, 1965; Wilbur, 1969] 

[Records: Chisos Mts., 28 Jul 1932, Mueller s.n.; top of Mt. Emory, Chisos 

Mts., 26 Aug 1937, Warnock 20976 (TEX-LL); Mt. Emory, Chisos Mts., Barkley 14T749; 

Flattop Mountain, Warnock 1116 (US); Flattop (Sperry, 1938)] 

Lechea mensalis Hodgdon 

[Hodgdon, 1938; Sperry, 1938; Wilbur, 1969; Correll & Johnston, 1970] 

[Brewster Co.: Chisos Mts., 28 Jul 1931, Mueller 8102 (GH) Type] 

Note: Some consider this to be no more than a subspecies or variety of Lechea tenuifolia. 

CONVOLVULACEAE (Morning Glory Family) 

Bonamia ovalifolia (Torr.) Hallier 

[Myint & Ward, 1968; Austin & Staplles, 1985; Austin, 1992] 

[Brewster Co.: Boquillas Canyon, edge of slope of deep sand, 22 Jul 1952, Webster 

4482 (TEX-LL; BRIT, W)] 

Bonamia repens (I. M. Johnston) Austin & Staples 

[Pterogenia repens I. M. Johnston] 

[Austin & Staples, 1985] 

[Brewster Co.: [expected in BBNP] Reed Plateau, W of Terlingua, Turner 1156 

(TEX-LL); Reagan Canyon, McVaugh 7816 (GH, TEX-LL)] 

Convolvulus equitans Benth. Hoary Bindweed


[Brewster Co.: Green Gulch and Panther Spring Creek Canyon (Sperry, 1938)] 

Dichondra argentea Humb. & Bonpl. ex Willd. Silver Ponyfoot 

[Sperry, 1938; Johnston, 1963, mapped] 

[Brewster Co.: Bois d’arc Spring (Sperry, 1938)] 

Dichondra brachypoda Woot. & Standl. New Mexico Ponyfoot 

[Sperry, 1938; Johnston, 1963, mapped] 

[Brewster Co.: Oak Creek Canyon and Green Gulch (Sperry, 1938)] 

Evolvulus alsinoides (L.) L. var. angustifolius Torr. Slender Evolvulus 

[Evolvulus alsinoides (L.) L. var. acapulcensis (Willd.) van Ooststr.] 

[Sperry, 1938; Austin, 1990] 

[Brewster Co.: Panther Spring and lower Green Gulch (Sperry, 1938); Grand Canyon 

of the Rio Grande, Aug, Parry s.n. (Tupes: GH, US, NY; could not be located by Austin]] 

Evolvulus pilosus Nutt. Hairy Evolvulus 

[Evolvulus nuttallianus J. A. Schultes] 



Ipomoea capillacea (Kunth) G. Don 

[Ipomoea muricata Cav. non (L.) Jacq.] 



Ipomoea costellata Torr. Crestrib Morning Glory 



Ipomoea cristulata Hallier f. Scarlet Morning Glory 

[Ipomoea hederifolia L. of some authors] 


[Brewster Co.: Basin and lower Blue Creek Canyon (Sperry, 1938)] 

Ipomoea lindheimeri Gray Blue Morning Glory 


[Brewster Co.: Basin and Mt. Emory (Sperry, 1938); along Boot Spring trail on 

E face of Emory Peak, BBNP, 29 DEG 15’N, 103 DEG 17’30”W, 6150 ft., 25 Aug 

1985, Yatskievych (UTEP)] 

CRASSULACEAE (Stonecrop Family) 

Echeveria strictiflora Gray 


[Records: lower Juniper Spring area (Sperry, 1938); Chisos Mts., Moore & 

Steyermark 3336] 

Sedum havardii Rose 

[Sperry, 1938; Correll & Johnston, 1970; Uhl, 1972] 

[Records: Pulliam Canyon (Sperry, 1938); N of Boot Spring, 6900 ft., Uhl 875 (BH); 

1.25 mi. S of Boot Spring, 7000 ft., Uhl 876 (BH); E side of Emory Peak, Blakely (BH); 

E side of Panther Pass and N side of Casa Grande, 6200 ft., Uhl 1674 (BH); Green Gulch, 

Clausen 363 (BH)] 

Sedum parvum Hemley ssp. nanifolium (Frod.) Clausen 

[?Sedum robertsianum Alexander] 



Sedum wrightii Gray Wright Stonecrop 

[Sperry, 1938; Clausen & Uhl, 1943; Uhl, 1972] 

[Records: Boot Spring area (Sperry, 1938); Green Gulch, 5600 ft., Uhl 877, 1675 (BH); 

upper Boot Canyon, 6900 ft., Uhl 873 (BH); Chinese Wall, Clausen 5298 (BH)] 

Note: Steyermark and Moore (1933) report Sedum moranense H.B.K. from the

Chisos Mts. (coll. no. 3169), but the record is questioned or not accepted (Correll 

& Johnston, 1970; Clausen, 1975). 

CROSSOSOMATACEAE (Crossosoma Family) 

Glossopetalon spinescens Gray var. spinescens Spiny Greasbush 

[Forsellesia spinescens (Gray) Greene] 

[Wells, 1965; Lundell, 1969; Powell, 1998] 

[Brewster Co.: Dead Horse Mts. and Panther Canyon (Powell, 1998)] 

CUCURBITACEAE (Gourd Family) 

Ibervillea tenuisecta (Gray) Small Slimlobe Globeberry 


[Brewster Co.: Gano Spring area and Green Gulch (Sperry, 1938)] 

Sicyos sp. 

[Sperry, 1938, as Sicyos angulatus L.] 


CUSCUTACEAE (Dodder Family) 

Cuscuta indecora Choisy Pretty Dodder 



EBENACEAE (Ebony or Persimmon Family) 

Diospyros texana Scheele Texas Persimmon 


[Brewster Co.: common in the S part, Big Bend Park “ (Powell, 1998); Chisos 

Mts. as far N as Persimmon Gap (Sperry, 1938); along Rio Grande, BBNP, 29 DEG 

22’30”N, 102 DEG 51’W, 1800 ft., 09 Nov 1988, Hodgson 5346 (UTEP, DES); Lost 

Mine Trail, BBNP, 26 Jun 1978, Freeman, s.n. (UTEP)] 

ERICACEAE (Heath Family) 

Arbutus xalapensis Kunth Texas Madeone; Naked Indian 


[Brewster Co.: Pinnacle’s trail towards Boot Springs (Jones & Jones, 1993); most canyons 

of Chisos Mts., Gano Spring, Pinnacle Mts. and near Santa Elene Canyon (Sperry, 1938); 

Basin at Saddle above Green Gulch, BBNP, 15 Jul 1981, Freeman s.n. (UTEP)] 

EUPHORBIACEAE (Spurge Family) 

Acalypha lindheimeri Muell.-Arg. 

[Sperry, 1938Johnston & Warnock, 1962, mapped] 

[Brewster Co.: Willow Creek Canyon, Oak Creek Canyon and Green Gulch (Sperry, 

1938)] 

Acalypha monostachya Cav. 

[Acalypha hederacea Torr.] 

[Sperry, 1938; Johnston & Warnock, 1962, mapped] 

[Brewster Co.: Muskhog Spring and near Boquillas Canyon (Sperry, 1998)] 

Acalypha neomexicana Muell.-Arg. 


[Brewster Co.: Chisos Mts. (Sperry, 1938)]

Andrachne arida (Warnock & M. C. Johnst.) G. L. Webst. 

[Savia arida Warnock & Johnston] 

[Warnock & Johnston, 1960; Johnston & Warnock, 1963, mapped; Powell, 1998] 

[Brewster Co.: Dead Horse Mts., middle western slope at 3400-4000 ft., 29 DEG 25’N, 

103 Deg 02’W, 21 Sep 1958, Warnock & Johnston 16840 (Type: SRSC), also Warnock 

10767 and 16681 (SRSC); Dead Horse Mts. near Sue Peak and Roy’s Peak (Powell, 1998)] 

Argythamnia neomexicana Muell.-Arg. 

[Ditaxis neomexicana (Muell.-Arg.) Heller] 

[Steyermark & Moore, 1933; Sperry, 1938; Johnston & Warnock, 1962, mapped] 

[Brewster Co.: Avary Canyon, Green Gulch, Smoky Creek and Boquillas (Sperry, 

1938); Santa Elena Canyon, Moore & Steyermark 3464 (as Ditaxis serrata)] 

Bernardia obovata I. M. Johnst. Desert Myrtlecroton 

[Steyermark & Moore, 1933; Johnston, 1940; Wells, 1965; Powell, 1998] 

[Brewster Co.: Dead Horse Mts. (Wells, 1965); Chisos Mts., Moore & Steyermark 3289 

as B. myricaefolia; I. M. Johnston made this the type of B. obovata; (GH); Green Gulch, 

Palmer 34137 (A)] 

Chamaesyce albomarginata (Torr. & Gray) Small White-margin Spurge 

[Sperry, 1938; Wheeler, 1941] 

[Brewster Co.: Pummel Peak and Boquillas (Sperry, 1938); near Lajitas, Warnock 

665 (US)] 

Chamaesyce arizonica (Engelm.) Arthur Arizona Spurge 


[Brewster Co.: Avery Canyon and near Lone Mtn. (Sperry, 1938); Persimmon Gap, Cory 

6906 (GH); 14 mi. E of Castolon, Cutler 664 (GH); Lone Mtn., Warnock 657 (US)] 

Chamaesyce capitellata (Engelm.) Millsp. 

[Wheeler, 1941] 

[Brewster Co.: Chisos Mts., Blue Creek, Palmer 34199 (MO; NY); near Castolon, Cory 

1907 (GH)] 

Chamaesyce chaetocalyx (Boiss.) Woot. & Standl. 

[Euphorbia fendleri Torr. & Gray var. chaetocalyx Boiss. and var. triligulata 

L. C. Wheeler] 


[Brewster Co.: Basin, Boquillas Canyon and W slope of Goat Mtn. (Sperry, 1938); Chisos 

Mts., Mueller 8077 (MO)] 

Chamaesyce cinerascens (Engelm.) Small 

[Steyermark & Moore, 1933; Sperry, 1938; Wheeler, 1941] 

[Brewster Co.: lower Green Gulch and Chisos Pens (Sperry, 1938); Lechuguilla Flats 

out of Green Gulch, Chisos Mts., Sperry 449 (US); Chisos Mts., Mueller 8080 (MO; NY; 

US); Chisos Mts., Moore & Steyermark 3266] 

Chamaesyce glyptosperma (Engelm.) Small Ridge-seed Spurge 



Chamaesyce golondrina (L. C. Wheeler) Shinners 

[Wheeler, 1940, 1941] 

[Records: sandy beach at entrance to Boquillas Canyon, 05 Aug 1937, Warnock 

998 (Type: US)] 

Chamaesyce hyssopifolia (L.) Small Hyssopleaf Spurge 


[Brewster Co.: Basin, Oak Creek Canyon and lower Green Gulch (Sperry, 1938); Oak 

Canyon, Chisos Mts., Cory 7005 (GH); Lower Green Gulch, Chisos Mts., Sperry 450 

(US)] 

Chamaesyce micromera (Boiss.) Woot. & Standl. 


[Brewster Co.: Persimmon Gap area, Sperry 1464 (US)] 

Chamaesyce serpyllifolia Pers. Winged Spurge 

[Sperry, 1938]


Chamaesyce setiloba (Engelm. ex Torr.) Millsp. ex Parish 


[Brewster Co.: between Nugent Mtn. and Lone Mtn., Warnock T526c (US); Persimmon 

Gap, Warnock T553 (US)] 

Chamaesyce theriaca (L. C. Wheeler) Shinners 


[Brewster Co.: Chisos Mts., Blue Creek, 01 Aug 1931, Mueller 8080 (GH, Type)] 

Chamaesyce villifera (Scheele) Small Hairy Spurge 


[Brewster Co.: Chisos Mts. (Sperry, 1938); Blue Creek, Chisos Mts., Cory 7006 (GH); 

Blue Creek Trail, Chisos Mts., Cory 7329 (GH); Chisos Mts., Mueller 8081 (GH; NY); 

Basin, Chisos Mts., Warnock C646 (US)] 

Croton dioicus Cav. 

[Croton neomexicanus Cav.] 


[Brewster Co.: Boquillas, Smoky Creek and Lechuguilla Flats (Sperry, 1938)] 

Croton fruticulosus Torr. 

[Sperry, 1938; Johnston & Warnock, 1962, mapped; Wells, 1965; Powell, 1998] 

[Brewster Co.: Muskhog Spring, Oak Creek Canyon and Panther Spring (Sperry, 1938); 

Dead Horse Mts. (Powell, 1998); Sue Peak (Wells, 1965)] 

Croton lindheimerianus Scheele var. tharpii M. C. Johnst. 

[Johnston & Warnock, 1962, mapped] 

Croton incanus Kunth Torry Croton 


[Brewster Co.: Boquillas Canyon (Powell, 1998)] 

Croton pottsii (Klotz.) Muell.-Arg. 

[Croton corymbulosus Engelm.] 

[var. pottsii] 


[Brewster Co.: Green Gulch, Pinnacle Mts. and Mesa de Anguilla (Sperry, 1938)] 

var. thermophilus (M. C. Johnst.) M. C. Johnst. 

[Johnston & Warnock, 1962, mapped; Powell, 1998] 

[Brewster Co.: Boquillas Canyon, Dead Horse Mts., Tornillo Creek and Daggar Flats 

(Powell, 1998)] 

Ditaxis humilis (Engelm. & Gray) Pax 

[Argythamnia humilis (Engelm. & Gray) Muell.-Arg. var. humilis] 

[Johnston & Warnock, 1962, mapped; Turner, 2001, mapped] 

Note: the pubescent form is mapped for BBNP (Turner, 2001). 

Euphorbia antisyphilitica Zucc. Candelilla 


[Brewster Co.: near Glenn Spring and in Fresno Canyon (Sperry, 1998); Boquillas, 


Euphorbia brachycera Engelm. 


[Brewster Co.: Pine Canyon (Sperry, 1938); Chisos Mts., Moore & Steyermerk 3334] 

Euphorbia dentata Michx. Toothed Spurge 



Euphorbia eriantha Benth. 


[Brewster Co.: between Roy’s Peak and Banta Shut-in,, along creek near Kit Mtn., 

along Rio Grande S of Chisos Mts. (Sperry, 1938); Boquillas Canyon, Moore & Steyermark 

3440 (MO)] 

Euphorbia exstipulata Engelm. var. lata Warnock & Johnston 

[Warnock & Johnston, 1969]

[Brewster Co.: Ward Spring, Chisos Mts., 18 Aug 1955, Warnock 13104 (SRSC) 

(Type); 4 mi. W Hot Springs, BBNP, 31 Oct 1948, Turner & Warnock 8314 (SRSC); 

N end Burro Meso near Chisos Pens, BBNP, Warnock 12840 (SRSC); Onion Spring, 

N side Paint Gap Hills, BBNP, Warnock 13080 (SRSC)] 

Euphorbia lurida Engelm. 



Jatropha dioica Cav. var. graminea McVaugh Leatherstem; Sangre de Drago 

[Jatropha spathulata Muell.-Arg.] 

[Steyermark & Moore, 1933; Sperry, 1938; Johnston & Warnock, 1963, mapped; 

Wells, 1965; Miller & Webster, 1966] 

[Brewster Co.: Rio Grande to Persimmon Gap (Sperry, 1938); Dead Horse Mts. 

(Wells, 1965); Boquillas, Moore & Steyermark 3437; BBNP, Miller & Miller 1268] 

Phyllanthus polygonoides Spreng. Weed Leaf Flower 

[Sperry, 1938; Johnston & Warnock, 1963, mapped; Webster, 1970, mapped] 

[Brewster Co.: Basin, Willow Creek Canyon and near Neville Spring (Sperry, 1938)] 

Stillingia treculiana (Muell.-Arg.) I. M. Johnst. Queen’s Delight 



Tragia ramosa Torr. Branching Noseburn 

[Tragia stylaris Muell.-Arg.] 

[Sperry, 1938; Miller & Webster, 1967, mapped] 

[Brewster Co.: Basin, Oak Creek Canyon and the Window (Sperry, 1938)] 

FABACEAE (Legume Family) 

Acacia angustissima (P. Mill.) Kuntze var. chisosiana Isely Fern Acacia 

[Acacia cuspidata Schlecht.] 

[Steyermark & Moore, 1933; Isely, 1969; Turner, 1996, mapped] 

[Brewster Co.: “Top of Divide (of Wilson)-Chisos Mts., July 27, 1937” Warnock 20719 

(NY) (Type); Chisos Mts., Moore & Steyermark 3338] 

Acacia berlandieri Benth. Berlandier Acacia; Guajillo 


[Brewster Co.: Burro Falls W of Chisos Mts. (Powell, 1998); S of Chisos Mts. 

(Sperry, 1998)] 

Acacia constricta Benth. Mescat Acacia; Whitethorn Acacia 


Acacia farnesiana (L.) Willd. Huisache; Sweet Acacia 


[Brewster Co.: “naar the Rio Grande, Boquillas and east to the corner of Big Bend 

National Park” (Powell, 1998)] 

Acacia greggii Gray var. greggii Catclaw Acacia 


[Brewster Co.: Gano Spring, Hot Spring and near Chisos Mts. (Sperry, 1938)] 

Acacia neovernicosa Isely Viscid Acacia 


Acacia rigidula Benth. Blackbrush Acacia 

[Acacia armentacea DC] 

[Sperry, 1938; Lee, Seigler & Ebinger, 1989; Powell, 1998] 

[Brewster Co.: Mariscal Mt. (Powell, 1998); near Solis’ ranch and Mariscal 

Canyon (Sperry, 1938); near Solis, BBNP, 13 Sep 1961, Correll & Johnston 

24565 (MO)] 

Acacia roemeriana Scheele Roemer Acacia 


[Brewster Co.: Oak Creek and Pulliam Bluff (Sperry, 1938)] 

Acacia schottii Torr. Schott Acacia


[Brewster Co.: Boquillas area (Powell, 1998)] 

Astragalus emoryanus (Rydb.) Cory var. terlinguensis (Cory) Barneby 

[Astragalus terlinguensis Cory] 

[Cory, 1937] 

[Brewster Co.: 18 mi. N of Terlingua on floodplain of Terlingua Creek, 13 Apr 1936, 

Cory 18584 (GH-type)] 

Astragalus molissimus Torr. 

[Sperry, 1938; Jones & Jones, 1993] 

[Brewster Co.: Pinnacle’s Trail toward Boot Springs (Jones & Jones, 1993); Green 

Gulch (Sperry, 1938)] 

Brongniartia minutifolia S. Wats. 

[Sperry, 1938; Correll & Johnston, 1970; Powell, 1998] 

[Brewster Co.: Solis’ Ranch to Lindsey’s Mine (Sperry, 1938); Chisos 

Mts. (Correll & Johnston, 1970); more records in Powell (1998)] 

Caesalpinia gilliesii (Hook.) Wallich ex D. Dietr. Bird-of-paradise 

[Isely, 1975, mapped in Lajitas area] 

Note: The species is widely cultivated in the Trans-Pecos. 

Caesalpinia jamesii (Torr. & Gray) Fisher James Rushpea 

[Hoffmannseggia jamesii Torr. & Gray] 


[Brewster Co.: near Glenn Spring (Sperry, 1938)] 

Caesalpinia parryi (Fisher) Eifert Parry Caesalpinia 

[Isely, 1975; Powell, 1998] 

[Brewster Co.: Heath, Big Brushy and Horse Canyons in Dead Horse Mts., Glenn 

Springs and Boquillas Canyon (Powell, 1998); Cuesta Carlota area 1 mi. N jct. rd. to 

Rio Grande Village along Old Ore Rd., BBNP, 2400-2800 ft., 27 May 1982, Van 

Devender s.n. (UTEP)] 

Calliandra conferta Benth. Flamemesquite; Calliandra 


[Brewster Co.: lower Chisos Mts., Ward Spring, Burro Spring (Powell, 1998)] 

Cercidium texanum Gray Texas Paloverde 

[Isely, 1975, mapped; Powell, 1998] 

[Brewster Co.: Santa Elena Canyon below Costolon (Powell, 1998)] 

Cercis canadensis L. var. mexicana (Rose) M. Hopk. Mexican Redbud 

[Sperry, 1938, as Cercis reniformis; Powell, 1998] 

[Brewster Co.: Pine Canyon (Sperry, 1938); Chisos and Dead Horse Mts. 

(Powell, 1998)] 

Cologania angustifolia Kunth 

[Cologania longifolia Gray] 

[Steyermark & Moore, 1933; Sperry, 1938; Turner, 1992, mapped] 

[Brewster Co.: Chisos Mts. (Sperry, 1938); Chisos Mts., Moore & Steyermark 3163] 

Dalea aurea Nutt. ex Pursh Golden Dalea 



Dalea formosa Torr. Feather Dalea 

[Wells, 1965; Powell, 1998; Spellenberg, 1981] 

[Records: Dead Horse Mts. (Wells, 1965); 10.5 mi. E Study Butte, 25 Mar 1975, Spellenberg 

3970 (NMC); upper slopes of Chisos Mts., 4 Apr 1942, Nelson & Nelson 5101 (TEX-LL)] 

Dalea greggii Gray Gregg Dalea 


[Brewster Co.: Dead Horse Mts. (Powell, 1998)] 

Dalea frutescens Gray Black Dalea 

[Sperry, 1938; Barneby, 1977] 

[Brewster Co.: Oak Creek Canyon, Green Gulch and flats arounf Chisos Mts. 

(Sperry, 1938); Basin at Saddle above Green Gulch, 15 Jul 1981, Freeman s.n.

(UTEP)] 

Dalea jamesii (Torr.) Torr. & Gray 



Dalea lachnostachys Gray 


[Brewster Co.: Basin, Dead Horse Mountains, Juniper Springs and Fresno 

Canyon (Sperry, 1938)] 

Dalea lanata Spreng. var. terminalis (M. E. Jones) Barneby 

[Barneby, 1977] 

Expected: “Rio Grande to Big Bend” (Barneby, 1977). 

Dalea leporina (Ait.) Bullock Foxtail Dalea 

[Dalea alopecuroides Willd.; Petalostemon oreophilum Cory] 

[Sperry, 1938; Cory, 1939] 

[Brewster Co.: Boot Canyon (Sperry, 1938); Boot Spring Canyon, 6800 ft., 29 Sep 

1955, Wallmo 49 (UTEP); Boot Creek near summit less than a mile from South Rim, 

26 Sep 1938, Cory (Type of Petalostemon oreophilum, GH)] 

Dalea neomexicana (Gray) Cory var. neomexicana 

[Steyermark & Moore, 1933, as D. mollis var. neomexicana; Wiggins, 1940, mapped; Mahler, 

1976, mapped] 

[Brewster Co.: flats between Lone Mtn. and Nugent Mtn., BBNP, 4000 ft., 03 Sep 

1947, Warnock 7177 (UTEP); Rio Grande river milage 828.5, 29 DEG 03’N, 103 DEG 

06’W, 1900 ft., 05 Nov 1988, Hodgson 5218 (UTEP, DES); Chisos Mts., Moore & Steyermark 

3270] 

Dalea pogonathera Gray Bearded Dalea 



Dalea wrightii Gray Wright Dalea 


[Brewster Co.: Hot Springs (Sperry, 1998)] 

Desmanthus glandulosus (Turner) Luckow Velvet Bundleflower 

[Desmanthus velutinus Scheele var. glandulosus Turner] 

[Sperry, 1938; Luckow, 1993] 

[Brewster Co.: desert areas surrounding Chisos Mts. (Sperry, 1938); Oak Canyon 

below the Window, BBNP, Correll & Correll 30605 (TEX-LL); upper Juniper Canyon, 

Chisos Mts., Ferris & Duncan 5772 (DS); Basin, Fletcher 152 (SRSC); mine trail 

Below Casa Grande, Luckow 2733 (TEX-LL); Chisos Mts., Mueller s.n. (MICH, BRIT) 

and Mueller 8024 (MICH, F, NY, TEX-LL, US); Chisos Mts., Basin, Sperry s.n. (TAES); 

BBNP, W slope facing the Window, Von Reis 89 (MICH); Chisos Mts., Oak Creek, 

Warnock 711 (TEX-LL); Chisos Mts., BBNP, Warnock 13108 (SRSC, TAES)] 

Desmanthus virgatus (L.) Willdenow 

[Luckow, 1993] 

[Brewster Co.: [expected in BBNP] Study Butte, Correll 33874 (TEX-LL)] 

Desmodium grahamii Gray 


[Brewster Co.: Blue Creek and Green Gulch (Sperry, 1998)] 

Desmodium psilophyllum Schlecht. 

[Desmodium wrightii Gray] 


[Brewster Co.: upper Green Gulch (Sperry, 19380] 

Eysenhardtia texana Scheele Texas Kidneywood 

[Sperry, 1938; Lang & Isely, 1982, mapped; Powell, 1998] 

[Brewster Co.: Oak Creek Canyon and Juniper Canyon (Sperry, 1938); 

Green Gulch (Powell, 1998); Basin at Saddle above Green Gulch, BBNP, 

15 Jul 1981, Freeman s.n. (UTEP); Lost Mine Trail, BBNP, 6600 ft., 26 Jun 

1978, Freeman s.n. (UTEP)]

Genistidium dumosum I. M. Johnston 

[Lavin & Sousa, 1995] 

[Brewster Co.: [expected in BBNP] see Lavin and Sousa (1995) for records from the 

Terlingua, Lajitas and Reed Plateau area] 

Gleditsia triacanthos L. 

[Isely, 1975, mapped] 

Hoffmannseggia glauca (Ort.) Eifert Indian Rushpea 

[Hoffmannseggia densiflora Benth.] 

[Sperry, 1938; Isely, 1975, mapped; Simpson, 1999] 

[Brewster Co.: Tornillo Flats (Sperry, 1938); 3 mi. S of Persimmon Flats (29 Deg 03’N, 

103 Deg 09’W), 05 Apr 1947, McVaugh 7830 (DS; F; GH; SR)] 

Leucaenia retusa Gray Littleleaf Leadtree 

[Sperry, 1938; Wells, 1965; Powell, 1998; Hughes, 1998] 

[Brewster Co.: Green Gulch and west of Dead Horse Mts. (Sperry, 1938); Dagger 

Flats and near Sue Peak (Powell, 1998); Dead Horse Mts. (Wells, 1965); Chisos Mts., 

Green Gulch, BBNP, 29 DEG 18’N, 103 DEG 18’W, Correll 30625 (NY, TEX-LL); 

Arroyo 24 km S of Persimmon Gap, 29 DEG 21’N, 103 DEG 07’W, Sperry 1375 

(GH, US)] 

Lotus plebeius (Brandegee) Barneby Commonplace Lotus 

[Lotus oroboides (H.B.K.) Ottley; L. puberulus (Benth.) Greene] 

[Sperry, 1938; Ottley, 1944; Isely, 1981, mapped] 

[Records: Green Gulch (Sperry, 1938); Chisos Mts., foothills, Aug 1883, Havard (US); 

Blue Creek Trail, Cory 7318 (CA-GR; CLA)] 

Note: Steyermark & Moore (1933) report the closely related Lotus wrightii (Gray) 

Greene from the Chisos Mts. 

Lupinus havardii S. Wats. Chisos Bluebonnet 


[Brewster Co.: Tornillo Creek, Avery Canyon and Dog Creek wash (Sperry, 1938); 

mouth of Panther Canyon, 04 Apr 1976, Everitt s.n. (UTEP)] 

Medicago sativa L. Alfalfa 



Mimosa aculeaticarpa Ortega var. biuncifera (Benth.) Barneby Catclaw Mimosa 


[Brewster Co.: Basin, Pulliam Bluff and Blue Creek (Sperry, 1938)] 

Mimosa borealis Gray Fragrant Mimosa 


[Brewster Co.: Green Gulch and Gano Spring (Sperry, 1938)] 

Mimosa emoryana Benth. Emory Mimosa 


[Brewster Co.: Chisos Mts. and vicinity (Powell, 1998); Green Culch and Nail’s 

Ranch (Sperry, 1938)] 

Mimosa texana (Gray) Small var. texana Texas Mimosa 


Note: This species has been confused with M. aculeaticarpa with which it 

is sympatric. 

Mimosa turneri Barneby Turner Mimosa 

[Barneby, 1986; Powell, 1998] 

[Brewster Co.: between Costolon and Smoky Creek (Powell, 1998); Chisos Mts., 

Correll 25425 (TEX-LL)] 

Peteria scoparia Gray Rush Peteria 


Phaseolus angustissimus Gray var. angustissimus Slimleaf Bean 


[Brewster Co.: Pinnacle Mountains (Sperry, 1938)] 

Phaseolus filiformis Benth.

[Phaseolus wrightii Gray] 


[Brewster Co.: Green Gulch, Blue Creek Canyon and Pinnacle Mountains 

(Sperry, 1938)] 

Pomaria melanosticta S. Schauer 

[Hoffmannseggia melanosticta (S. Schauer) Gray] 

[Brewster Co.: BBNP, 0.7 mi. W of tunnel near Rio Grande Village, 746 m, 18 Mar 1985, 

Van Devender et al. 85-65 (ARIZ); 11 mi. from hwy. down old Boquillas Rd., lower slopes 

of Dead Horse Mts., 1200 m, 21 Sep 1958, Warnock & Johnston 16835 (SRSC); Boquilla 

Canyon, 666 m, 1 Apr 1961, Warnock 18410 (SRSC); Dead Horse Mts. at head of Heath 

Canyon, 1266 m, 19 Jul 1952, Warnock 10830 (SRSC); Green Basin, Chisos Mts., 23 Sep 

1938, Parks 30173 (AA)] 

Prosopis glandulosa Torr. var. torreyana (L. Benson) M. C. Johnston Honey Mesquite 

[Sperry, 1938; Johnsron, 1962, mapped; Powell, 1998] 

[Records: Oak Creek Canyon, Johnson’s Ranch and McKinney Spring 

(Sperry, 1938)] 

Prosopis pubescens Benth. Screwbean Mesquite 


[Brewster Co.: Tornillo Creek and Glenn Springs (Powell, 1998); near Johnson’s 

Ranch and Glenn Spring (Sperry, 1938)] 

Rhynchosia senna Gillies ex Hook. var. texana (Torr. & Gray) Texas Senna 

M. C. Johnst. 

[Sperry, 1938; Grear, 1978] 

[Brewster Co.: Green Gulch and Blue Creek Canyon (Sperry, 1938); Green Gulch, 08 Jul 1931, 

Moore & Steyermark 3420 (MICH, MO, NY)] 

Senna bauhinioides (Gray) Irwin & Barneby Shrubby Senna 

[Cassia bauhinioides Gray] 

[Sperry, 1938; Isely, 1975, mapped] 

[Brewster Co.: Croton Spring and Smoky Creek (Sperry, 1938)] 

Senna durangensis (Rose) Irwin & Barneby var. isleyi (Irwin & 

Barneby) Irwin & Barneby 

[Cassia durangensis Rose] 


[Brewster Co.: near Santa Elena Canyon, Hot Springs area and Croton 

Springs (Sperry, 1938)] 

Senna lindheimeriana (Scheele) Irwin & Barneby Lindheimer Senna 

[Cassia lindheimeri Scheele] 


[Brewster Co.: lower Green Gulch and Basin (Sperry, 1938); on Basin Rd. 0.5 mi. 

from Lost Mine Trail, Aug 1983, Stafford s.n. (UTEP)] 

Sophora secundiflora (Ort.) Lag. ex DC. Mescalbean; Texas Mountain Laurel 

[Sperry, 1938; Isely, 1981, mapped; Powell, 1998] 

[Brewster Co.: Oak Creek Canyon (Sperry, 1938); bank of Panther Canyon, 05 Apr 1976, 

Everitt s.n. (UTEP)] 

Vicia ludoviciana Nutt. ssp. ludoviciana Slim Vetch; Louisiana Vetch 

[Vicia exigua Nutt. in Torr. & Gray] 



FAGACEAE (Beech Family) 

Quercus carmenensis C. H. Mull. Delcarmen Oak 


[Brewster Co.: “Chisos Mts., Casa Grande, ca. 7300 ft., just below the 

SE bluffs” Sept. 1982, A. M. Powell (SRSC)] 

Quercus emoryi Torr. Emory Oak


[Brewster Co.: Chisos Mts. (Sperry, 1938); Lost Mine Trail, BBNP, 26 Jun 1978, 

Freeman s.n. (UTEP)] 

Quercus gambelii Nutt. Gambel Oak 


[Brewster Co.: Casa Grande and Lost Mine Peak (Powell, 1998)] 

Quercus graciliformis C. H. Mull. Chisos Oak 


[Brewster Co.: endemic to Chisos Mts.] 

Quercus gravesii Sudw. Graves Oak 


[Brewster Co.: Chisos Mts. (Sperry, 1938); upper end of Lost Mine Trail, 

BBNP, 26 Jun 1978, Freeman s.n. (UTEP)] 

Quercus grisea Liebm. Gray Oak 


[Brewster Co.: Chisos Mts. (Sperry, 1938); Lost Mine Trail, BBNP, 26 Jun 1978, 

Freeman s.n. (UTEP)] 

Quercus intricata Trel. Coahuila Scrub Oak 

[Sperry, 1938; Wells, 1965; Muller, 1979, mapped; Powell, 1998] 

[Brewster Co.: Sue Peaks (Wells, 1965); Cattail Canyon, Laguna Meadow, 

Boot Springs to South Rim, and top of Lost Pine Trail (Powell, 1998)] 

Qercus muehlenbergii Engelm. Chinkapin Oak 


[Brewster Co.: Chisos Mts., Maple or Pulliam Canyon (Powell, 1998)] 

Quercus pungens Liebm. 

var. pungens Sandpaper Oak; Scrub Oak 


[Brewster Co.: Dead Horse Mts. in Heath and Big Brushy Canyons (Powell, 1998)] 

var. vaseyana (Buckl.) C. H. Mull. Vasey Shin Oak 

[Wells, 1965; Powell, 1998] 

[Brewster Co.: Chisos Mts., Dead Horse Mts. (Powell, 1998)] 

Quercus rugosa Nee Netleaf Oak 


[Brewster Co.: Boot Canyon and the Pinnacles Trail (Powell, 1998)] 

Quercus tardifolia C. H. Mull. Lateleaf Oak 


[Brewster Co.: “between Boot Spring and the South Rim along upper Boot Canyon 

(Powell, 1998); Boot Spring, 7000 ft., 13 Sep 1997, Powell 6183 (UTEP)] 

FOUQUIERIACEAE (Ocotillo Family) 

Fouquieria splendens Engelm. Ocotillo 

[Sperry, 1938; Wells, 1965] 

[Brewster Co.: Dead Horse Mts. (Wells, 1965); Chisos Mts. and surrounding 

hills (Sperry, 1938)] 

GARRYACEAE (Silktassel Bush Family) 

Garrya ovata Benth. ssp. lindheimeri (Torr.) Dahling Eggleaf Silktassel 

[Sperry, 1938; Wells, 1965; Jones & Jones, 1993; Powell, 1998] 

[Brewster Co.: Chisos Mts. (records in Powell, 1998); Dead Horse Mts. (Wells, 1965); 

Pulliam Canyon, Boot Spring and S slope Pulliam Bluff (Sperry, 1938)] 

GENTIANACEAE (Gentian Family)

Centaurium calycosum (Buckl.) Fern. Rosita 

[Centaurium arizonicum (Gray) Heller] 

[Sperry, 1938; Turner, 1993, treated as C. arizonicum and mapped in BBNP] 

[Brewster Co.: Gano Spring (Sperry, 1938)] 

Eustoma exaltatum (L.) Salisb. ex G. Don Prarie Gentian 


[Brewster Co.: Boquillas and near Santa Elena Canyon (Sperry, 1938)] 

GERANIACEAE (Geranium Family) 

Erodium texanum Gray Texas Filaree 



Geranium caespitosum James Tufted Geranium; Purple Geranium 



Geranium lentum Woot. & Standl. 

[Jones & Jones, 1943] 

[Brewster Co.: Chisos Mts., Mueller 8050 (F)] 

HYDROPHYLLACEAE (Waterleaf Family) 

Nama havardii Gray Havard Nama 

[Steyermark & Moore, 1933; Hitchcock, 1933; Sperry, 1938; Bacon, 1974] 

[Records: Tornillo Creek, Rio Grande near Hot Springs and Tally 

Mountain (Sperry, 1938); 0.5 mi. E Tornillo Creek on rd. to Rio Grande Village, 

BBNP, 21 Aug 1982, Van Devender s.n. (UTEP); 3 mi. E of Study Butte , Moore & 

Steyermark 3248 (MO; UC); Hot Springs, Bacon 957 (TEX-LL); Hot Springs, 1928, 

Cory (GH); Hot Springs, Jones 25737 (RSA-POM); bank of Tornillo Creek, Havard 

95 ½ (Type: GH)] 

Nama hispidum Gray Rough Nama 

[Hitchcock, 1933; Sperry, 1938; Bacon, 1974] 

[Records: Tornillo Flats, Basin, Painted Gap, Terrapin Tank and deserts 

N of Chisos Mts. (Sperry, 1938); 48.8 mi. S of Marathon on hwy. 385, Bacon 

956 (TEX-LL); mouth of Santa Elena Canyon, Moore & Steyermark 3461 (MO)] 

Nama torynophyllum Greenm. 

[Cory, 1939] 

[Brewster Co.: 2 mi. E of Castolon, 4 Mar 1937, Cutler; 1 mi. N of Castolon, 28 Sep 1938, 

Cory; gravelly bed of Marvillas Creek, 5 Apr 1939, Cory] 

Nama undulatum Kunth 

[Hitchcock, 1933] 

[Records: mouth of Terlingua River, Havard 95 (GH)] 

Phacelia congesta Hook. Spike Phacelia 


[Brewster Co.: Pulliam Canyon and slopes of Casa Grande (Sperry, 1938)] 

Phacelia integrifolia Torr. var. integrifolia 

[Voss, 1937] 

[Brewster Co.: Chisos Mts., Moore & Steyermark 3232 (GH, UC, NY); Chisos Mts., 

Palmer 34077 (NY)] 

Phacelia popei Torr. & Gray Pope Phacelia 


[Brewster Co.: Dog Creek Wash and Tornillo Flats (Sperry, 1938)] 

Phacelia robusta (Macbr.) I. M. Johnst. Stout Phacelia 

[Sperry, 1938; Cave & Connstance, 1950] 

[Brewster Co.: Willow Creek Canyon, Oak Creek Canyon and slopes

of Casa Grande (Sperry, 1938); BBNP, Albers 46213] 

Phacelia rupestris Greene Rock Phacelia 

[Phacelia congesta Hook. var. rupestris (Greene) J. F. Macbr.] 

[Voss, 1937] 

[Brewster Co.: Chisos Mts., Moore & Steyermark 3325 (GH, NY, UC)] 

JUGLANDACEAE (Walnut Family) 

Juglans major (Torr.) Heller Arizona Walnut 

[Juglans rupestris Engelm. ex Torr. var. major Torr.] 


[Brewster Co.: Nail Ranch, BBNP (Powell, 1998)] 

Juglans microcarpa Berl. var. stewartii (I. M. Johnston) Little Walnut 

W. Manning 

[Correll & Johnston, 1970; Powell, 1998] 

[Brewster Co.: Chisos Mts. (Correll & Johnston, 1970)] 

KRAMERIACEAE (Ratany Family) 

Krameria erecta Willd. ex Schultes Range Krameria 

[Krameria glandulosa Rose & Painter] 

[Sperry, 1938; Simpson, 1989] 

[Brewster Co.: 3 mi. S of Persimmon Gap, 05 Apr 1947, McVaugh 7833 

(DS, G, GH, MICH, P, TEX-LL); along Blue Creek, 26 May 1928, Palmer 

34198 (A, MO); Persimmon Gap, 24 Apr 1931, McKelvey 1982 (A); lower 


Krameria grayi Rose & Painter White Krameria 

[Sperry, 1938; Simpson, 1989] 

[Brewster Co.: Blue Creek Tract, Chisos Mts., 03 Oct 1933,Cory 7323 (GH); 

6 mi. SW Glenn Spring, 27 May 1938, Cutler 1869 (CAS, GH, MO); 9 mi. N 

Santa Elena Rd., just SW Chisos Basin Rd., 13 May 1961, Traverse 2231 (A, 

TEX-LL); north of Chisos Mts., Juniper Canyon and Smoky Creek (Sperry, 1938)] 

LAMIACEAE (Mint Family) 

Agastache micrantha (Gray) Woot. & Standl. var. micrantha White Gianthyssop 

[Sperry, 1938; Lint & Epling, 1945; Sanders, 1987] 

[Records: Boot Spring area (Sperry, 1938); Chisos Mts., Correll 13709 

(TEX-LL; NY); Boot Spring, Warnock 1070, Cory 30421, Mueller 8145] 

Agastache pallidiflora (Heller) Rydb. ssp. neomexicana (Briq.) Lint 

& Epling var. havardii (Gray) Sanders 

[Sperry, 1938; Lint & Epling, 1945; Sanders, 1987] 

[Records: canyon on E slope of Casa Grande and Pulliam Canyon (Sperry, 1938); 

Chisos Mts., Correll & Johnsron 24514 (TEX-LL); Chisos Mts., cliffs and ravines , Aug 1883, 

Havard 29 (Type, GH); Pulliam Canyon, Cory 26514, Sperry 298; E slope of Casa Grande, 

6800 ft., Warnock 989] 

Hedeoma drummondii Benth. Drummond Pennyroyal 

[Sperry, 1938; Irving, 1980, mapped in BBNP] 

[Records: Muskhog Spring, Lone Mtn. and Green Gulch (Sperry, 1938)] 

Hedeoma mollis Torr. Hairy Hedeoma 

[Poliomintha mollis Gray] 

[Steyermark & Moore, 1933; Sperry, 1938; Irving, 1980, mapped in BBNP; Powell, 1998] 

[Brewster Co.: Chisos Mts. Powell, 1998); Pulliam Canyon and W slope of 

Casa Grande (Sperry, 1938); Chisos Mts., Moore & Steyermark 3409] 

Hedeoma nana (Torr.) Briq. Ssp. nana Low False Pennyroyal

[Hedeoma thymoides Gray] 

[Steyermark & Moore, 1933; Sperry, 1938; Irving, 1976; Irving, 1980, mapped in BBNP] 

[Brewster Co.: Oak Creek (Sperry, 1938); Pine Canyon, BBNP, Correll & Waushausen 

27882 (TEX-LL); Boquillas, Moore & Steyermark 3445] 

Hedeoma plicata Torr. 

[Steyermark & Moore, 1933; Irving, 1980, mapped] 


Marrubium vulgare L. Common Horehound 


[Brewster Co.: Terrapin Tank (Sperry, 1938)] 

Mentha canadensis L. 

[Mentha penardi (Briq.) Rydb.] 



Mentha spicata L. Spearmint 



Salazaria mexicana Torr. Bladder-sage 


[Brewster Co.: between Rio Grande and Chisos Mts., Mesa de Anguilla 

and Smoky Creek (Sperry, 1938); near Smoky Creek, between Smoky Creek 

and Costolon, near Alamo Spring, between Costolon and Santa Elene Canyon, 

and top of Mesa de Anguilla (Powell, 1998)] 

Salvia arizonica Gray 


[Brewster Co.: Willow Creek Canyon and Basin (Sperry, 1938)] 

Salvia greggii Gray Autumn Sage 


[Brewster Co.: Dead Horse Mts. (Powell, 1998)] 

Salvia lycioides Gray Canyon Sage 


[Brewster Co.: Pulliam Canyon, Laguna and Basin (Sperry, 1938); Chisos Mts. 

(Powell, 1998)] 

Salvia reflexa Hornem. Rocky Mountain Sage 

[Salvia lanceaefolia Poir.] 



Salvia regla Cav. Mountain Sage; Royal Sage 

[Steyermark & Moore, 1933; perry, 1938; Powell, 1998] 

[Brewster Co.: Laguna, Pulliam Canyon and Green Gulch (Sperry, 1938); Green 

Gulch, Pulliam, Pine, Boot and Wade Canyons, Lost Mine Peak, Casa Grande, Mt. 

Emory and elsewhere (Powell, 1998); Chisos Mts., Moore & Steyermark 3179] 

Salvia roemeriana Scheele Cedar Sage 


[Brewster Co.: Green Gulch and S slope Pulliam Bluff (Sperry, 1938)] 

Scutellaria drummondii Benth. 



Scutellaria potosina Brandeg. var. tessellata (Epling) Turner 

[Sperry, 1938, as S. resinosa Torr.; Turner, 1994] 

[Brewster Co.: Green Gulch, slopes of Pulliam Bluff and Panther Spring Creek 

(Sperry, 1938); upper Blue Creek, Chisos Mts., 1520 m, 25 Jun 1931, Moore & 

Steyermark 3231 (Type of ssp. parviflora: UC; Isotypes: CAS-DS, GH, MICH, MO)] 

Stachys bigelovii Gray Rock Betony 

[Sperry, 1938; Correll & Johnston, 1970; Turner, 1994, mapped] 

[Brewster Co.: Pulliam Canyon, slopes of Pulliam Bluff and Casa Grande

(Sperry, 1938)] 

Stachys coccinea Jacq. Texas Betony 



Teucrium cubense Jacq. ssp. laevigatum (Vahl) Shinners 

[Sperry, 1938; McClintock & Epling, 1946] 

[Records: Oak Creek Canyon (Sperry, 1938); Brewster Co., Mueller 8147] 

LINACEAE (Flax Family) 

Linum aristatum Engelm. 


[Brewster Co.: Chisos Mts. (Sperry, 1938); Basin at Saddle above Green Gulch, BBNP, 

15 Jul 1981, Freeman s.n. (UTEP)] 

Linum australe Heller var. australe 

[Sperry, 1938; Rogers, 1964, 1968] 

[Records: Muskhog Spring area (Sperry, 1938)] 

Linum berlandieri Hook. var. filifolium (Shinners) Rogers 

[Linum rigidum Pursh var. filifolium Shinners] 

[Rogers, 1968, 1979, mapped in BBNP] 

?Linum hunsonioides Planch. 

[Rogers, 1963] 

[Records: Brewster Co., Mueller 8056 (TEX-LL)] 

Linum imbricatum (Raf.) Shinners 

[Linum multicaule Hook.] 


[Records: open slopes of Green Gulch (Sperry, 1938)] 

Linum puberulum (Engelm.) Heller 

[Rogers, 1968, mapped in Brewster Co.] 

Linum lewisii Pursh Blue Flax 


[Records: Basin and near top of Lost Mine Peak (Sperry, 1938)] 

Linum rupestre (Gray) Engelm. Rock Flax 

[Sperry, 1938; Rogers, 1968] 

[Records: Basin, near top of Baldy Mtn. and Panther Spring (Sperry, 1938)] 

Linum schiedeanum Schlecht. & Cham. Schied Flax 

[Linum greggii Engelm.] 

[Steyermark & Moore, 1933; Sperry, 1938; Osborne, W. P. & W. H. Lewis, 1962; 

Rogers, 1964, 1968] 

[Records: Chisos Mts., BBNP, Osborne 73-76 (BRIT)]; below falls in Wade 

Canyon and slopes of Casa Grande (Sperry, 1938); near Boot Spring, Chisos Mts., 

Moore & Steyermark 3225 (MO)] 

Linum sulcatum Riddell Grooved Yellow Flax 


[Records: near base of Lone Mtn. (Sperry, 1938)] 

Linum vernale (Woot.) Small Spring Flax 

[Sperry, 1938; Rogers, 1968] 

[Records: Tornillo Flats to Persimmon Gap (Sperry, 1938)] 

LOASACEAE (Stickleaf Family) 

Cevallia sinuata Lag. Stinging Cevallia; Shirley’s Nettle 

[Steuyermark & Moore, 1933; Sperry, 1938; Powell, Powell & Tomb, 1977; Powell, 1998] 

[Brewster Co.: Rio Grande region (Sperry, 1938); 10 mi. E Panther Jct., Averett 141 

(SRSC); 6.1 mi. SE Panther Jct., High 128 (SRSC); 1 mi. E jct. Costolon and Mariscal

Mine roads, High 130 (SRSC); near San Vicente, High 131 (SRSC); Chisos Mts., Moore 

& Steyermark 3253] 

Eucnide bartonioides Zucc. Yellow Rocknettle 

[Steyermark & Moore, 1933; Sperry, 1938; Waterfall, 1959; Hufford, 1987, 1988] 

[Brewster Co.: banks of Tornillo Creek, Rio Grande at Hot Springs and Casa 

Grande (Sperry, 1938); Panther Canyon, 05 Apr 1976, Everitt s.n. (UTEP); BBNP, 

26 Jun 1962, Thompson & Ernst 3283 (LA); near Hot Springs, BBNP, 13 Jul 1931, 

Moore & Steyermark 3791 (UC); banks of Tornillo Creek near Hot Springs, Sperry 

519 (US)] 

Mentzelia asperula Woot. & Standl. 

[Thompson & Powell, 1981, mapped] 

?Mentzelia lindheimeri Urban & Gilg. 


Note: This collection by Sperry may not be properly identified. 

Mentzelia mexicana Thompson & Zavortink 

[Thompson & Zavortink, 1968; Thompson & Powell, 1981] 

[Brewster Co.: mouth of Terlingua Creek, Whitehouse 19621 (BRIT); Hot Springs, 

Sperry 1746 (GH; US); W of Glen Springs, Rose-Innes & Moon 1198 (GH); mouth 

of Santa Elena Canyon, Rollins & Chambers 2769 (GH); Hot Springs, Sperry 1679 (US); 

N of Castolon, Thompson 3548 (LA); Boquillas Canyon, Thompson 3547 (LA)] 

Mentzelia multiflora (Nutt.) Gray 

[Thompson & Powell, 1981, mapped] 

Note: Darlington (1934) reports Mentzelia pumila (Nutt.) T. & G. var. procera from 

mesa N of Chisos Mts. (Moore & Steyermark 3275 at MO). This specimen needs to 

be checked to see if it is annotated to something else (likely multiflora). 

Mentzelia oligosperma Sims. Chickenthief; Stickleaf 

[Sperry, 1938; Thompson & Powell, 1981] 

[Brewster Co.: lower Oak Creek (Sperry, 1938); 1.3 mi. N of pavement on Old 

Ore Rd. to Ernst Tinaja, 21 Aug 1982, Van Devender s.n. (UTEP); Pine Canyon, Powell 

et al., 3233 (SRSC)] 

Mentzelia pachyrhiza I. M. Johnston 

[Thompson & Powell, 1981] 

[Brewster Co.: BBNP near W entrance, Thompson 3727 (LA)] 

LYTHRACEAE (Loosestrife Family) 

Lythrum californicum Torr. & Gray California Lythrum 

[Lythrum linearifolium (Gray) Small] 


[Brewster Co.: near Chisos Mts. (Sperry, 1938)] 

Nesaea longipes Gray Stalkflower Nesaea 

[Heimia longipes (Gray) Cory] 

[Soerry, 1938; Powell, 1998] 

[Brewster Co.: near Glenn and Fresno Springs (Powell, 1998); Lechuguilla Pens 

and Fresno Springs (Sperry, 1938)] 

MALPIGHIACEAE (Malpighia Family) 

Janusia gracilis Gray Slender Janusia; Propellerbush 


[Brewster Co.: “S half of county to Rio Grande (Powell, 1998); lower Cattail 

Canyon, Glenn Spring, Persimmon Gap, Fresno Spring and Boquillas (Sperry, 1938); 

Chisos Mts., Moore & Steyermark 3358]

MALVACEAE (Mallow Family) 

Abutilon incanum (Link) Sweet 



Abutilon malacum S. Wats. 


[Brewster Co.: Blue Creek Canyon, flats N of Kit Mtn., lower Juniper Spring and 

Boquillas (Sperry, 1938)] 

Abutilon parvulum Gray Littleleaf Abutilon 


[Brewster Co.: S of Chisos Mts. (Sperry, 1938)] 

Allowissadula holosericea (Scheele) Bates 

[Abutilon marshii Standl.] 


[Brewster Co.: Chisos Mts. (Sperry, 1938; Correll & Johnston, 1970)] 

Anoda crenatiflora Ortega 

[Fryxell, 1987] 

[Records: North Sunny Glen, 22 Sep 1940, Sperry T840 (TAES)] 

Batesimalva violacea (Rose) Fryxell 

[Gaya violacea Rose] 

[Correll, 1996; Powell, 1998] 

[Records: W side of Chisos Mts. (Powell, 1998); Oak Canyon below the Window, Correll & 

Correll 30610 (TEX-LL)] 

Herissantia crispa (L.) Briz. 


[Brewster Co.: S of Chisos Mts. (Sperry, 1938)] 

Hibiscus coulteri Harv. ex Gray Desert Rosemallow 

[Sperry, 1938; Fryxell, 1980] 

[Brewster Co.: lower Green Gulch and N of Chisos Mts. (Sperry, 1938); Green Gulch, 

Marsh 272 (UC); near/in Chisos Mts., Sperry 115 (TAES, US), Parks & Cory 7016 (TAES), 

Muller 8091 (MO, US), Ferris & Duncan 2823 (DS), Warnock & Wallmo 12302 (TAES); 

Dead Horse Mts. near McKinney Spring, Warnock 10820 (BRIT)] 

Hibiscus denudatus Benth. Paleface Rosemallow 

[Sperry, 1938; Fryxell, 1980] 

[Brewster Co.: Glenn Spring and N and W side of Chisos Mts. (Sperry, 1938); near Castalon, 

Fryxell 1246 (TAES, US); near Chisos Mts., Sperry 339 (TAES, US); Juniper Canyon, 16 mi. 

NE of Castolon, Cutler 21834 (TAES), between San Vicente and Glenn Springs, Sperry 1561 

(TAES); 14.5 mi. SE of Terlingua, Parks & Cory 30271 (TAES); Ward Spring, W side Chisos 

Mts., Warnock 13111 (TEX-LL, TAES); between Burro and Wasp Springs, W side of Chisos 

Mts., Warnock & Wallmo 13233 (TAES); 14 mi. W BBNP HQ, Hess 817 (OKLA; BRIT); 

Heath Canyon, Warnock & Hinckley 235 (BRIT); Santa Elena Canyon, Rose-Innes & Warnock 

21599 (BRIT)] 

Rhynchosida physocalyx (Gray) Fryxell Spear-leaf Sida 



Sida abutifolia P. Mill. Spreading Sida 

[Sida procumbens Sw.] 


[Brewster Co.: low hills N of Chisos Mts., Pinnacle Mts., and Green Gulch (Sperry, 

1938)] 

Sida lindheimeri Engelm. & Gray 



Sphaeralcea angustifolia 9Cav.) G. Don Narrowleaf Globemallow 

var. cuspidata (Gray) Kearney

[Sperry, 1938; La Duke, 1981] 

[Brewster Co.: Boquillas, Terrapin Tank, Oak Creek Canyon, Green Gulch and 

along Rio Grande S of Chisos Mts. (Sperry, 1938); Basin Campground, BBNP, 13 Jul 1983, 

La Duke 919 (GFND, not to variety)] 

Sida neomexicana Gray 



MENISPERMACEAE (Moonseed Family) 

Cocculus carolinus (L.) DC. 


[Brewster Co.: lower Panther Spring Creek and Panther Canyon (Sperry, 1938); Chisos 


MORACEAE (Mulberry Family) 

Maclura pomifera (Raf.) Schneid. Osage Orange 


[Brewster Co.: Bois d’arc Spring, Sperry1125] 

Morus microphylla Buckl. Mountain Mulberry 


[Brewster Co.: Panther Canyon, Wade Canyon, Oak Creek Canyon and upper 

Juniper Canyon (Sperry, 1938)] 

NYCTAGINACEAE (Four O’Clock Family) 

Abronia angustifolia Greene 

[Sperry, 1938; Reed, 1969; Galloway, 1975] 

[Records: Glenn Springs, 20 Apr 1938, Sperry 1325; Glenn Springs along Rio Grande, 

30 Apr 1938, Warnock T384 (US; ARIZ; TEX-LL); Dagger Flats (Sperry, 1938)] 

Acleisanthes longiflora Gray Angel Trumpets 

[Sperry, 1938; Reed, 1969] 

[Brewster Co.: Mesa de Anguila, Chisos Mts., 14 Jul 1937, Warnock 859; 

Mariscal Mt. Near Lindsay Mine, 25 May 1959, Warnock & Wallmo 12279; 

Mesa de Anguila (Sperry, 1938)] 

Allionia choisyi Standl. Smooth Umbrellawort 

[Wedeliella glabra (Choisy) Cockerell] 

[Sperry, 1938; Reed, 1969] 

[Brewster Co.: mouth of Terlingua Creek, BBNP, 22 Jul 1957, Correll & 

Johnston 18344; Chisos Mts., 25 Jun 1937, Marsh 15a, 31 Aug 1936, Sperry 

349; Rio Grande S of Chisos Mts. (Sperry, 1938)] 

Allionia incarnata L. var. incarnata. Trailing Four O’Clock 

[Wedeliella incarnata (L.) Cockerell] 

[Sperry, 1938; Reed, 1969; Turner, 1994, mapped] 

[Brewster Co.: Paint Gap Hills, BBNP, 12 Aug 1955, Warnock 12883; Santa Elena 

Canyon, 22 May 1933, Parks & Cory 6379; Oak Canyon, Chisos Mts., 06 Sep 1933, 

Wolff 4622; et al. in Reed (1969)] 

Ammocodon chenopodioides (Gray) Standl. Goosefoot Moonpod 

[Reed, 1969; Fowler & Turner, 1977, mapped] 

[Brewster Co.: Green Gulch, Chisos Mts., 10 Sep 1925, Berkman 3411; Chisos 

Mts., 05 Sep 1937, Marsh 340; between Costolon and Johnson Ranch, 20 Apr 

1940, Sperry 1706; flats N of Chisos Mts., near Santa Elena Canyon, Boquillas 

and near Hot Springs (Sperry, 1938)] 

Anulocaulis eriosolenus (Gray) Standl. Big Bend Ringstem

[Standley, 1909; Steyermark & Moore, 1933; Sperry, 1938; Reed, 1969; Spellenberg, 

1993] 

[Brewster Co.: Hot Springs area, (numerous collections); et al. in Reed (1969); 

2 mi. NW of Boquillas, 20 Jul 1936, Correll & Johnston 24563 (ARIZ, F, GH, UC); 

Hot Springs, 13 Jul 1951, Moore & Steyermark 3456 (CAS, DUD, GH, NY, UC); 

Hot Springs, Muskhog Spring and Boquillas (Sperry, 1938); bluffs of the Rio Grande, 

1883, Havard] 

Anulocaulis leiosolenus (Torr,) Standl. 

var. leiosolenus 

[Reed, 1969; Spellenberg, 1993] 

[Brewster Co.: Tornillo Creek, 1883, Havard s.n.] 

Note: Spellenberg (1993) found no records of this variety in BBNP. 

var. lasianthus I. M. Johnston 

[Reed, 1969; Spellenberg, 1993] 

[Brewster Co.: Hot Springs, 1937, Warnock 701A; Tornillo Creek, 1883, Havard (this 

coll. not det. to var.)] 

Boerhavia anisophylla Torr. 

[Standley, 1909; Sperry, 1938; Reed, 1969] 

[Brewster Co.: Ward Spring, Chisos Mts., 18 Aug 1955, Warnock 13085; Lindsay 

Mine, Chisos Mts., 02 Jul 1949, Warnock & Parks 8751; Alamo Creek, BBNP, 

05 Aug 1955, Warnock 12856; Boquillas Canyon, 31 Oct 1948, Warnock, Turner 

& Allen 8335; Solis N of Mariscal Canyon, 13 Sep 1961, Correll & Wasshausen 

27894; Chisos Mts., 05 Sep 1937, Marsh 336; Muskhog Spring, Chisos Mts., 11 Jun 

1937, Warnock 1108; Dead Horse Mts., 02 Aug 1937, Warnock 21031; Rays Peak, 

Chisos Mts., 02 Aug 1937, Warnock & Sperry 698; Muskhog Spring and Tornillo 

Creek to Mail Box (Sperry, 1938); Tornillo Creek, Havard 63] 

Boerhavia coccinea P. Mill. Scarlet Spiderling 

[Boerhavia caribaea Jacq.] 

[Sperry, 1938; Reed, 1969] 

[Brewster Co.: Chisos Mts., 26 Aug 1915, Young s.n.; Oak Canyon, Chisos Mts., 

06 Sep 1933, Wolff 4456; Chisos Mts., 03 Sep 1947, Warnock 7187; Pinnacle Mts., 

Chisos Mts., 22 Jul 1937, Warnock 700; Juniper Springs, Chisos Mts., 25 Jun 1937, 

Warnock 20239, 05 Aug 1937, Warnock 20237; et al. in Reed (1969); Chisos Pens 

(Sperry, 1938)] 

Boerhavia erecta L. Erect Spiderling 

[Reed, 1969] 

[Brewster Co.: Ward Spring, Chisos Mts., 18 Aug 1955, Warnock 12998; Costolon, 

28 Sep 1938, Parks & Cory 30511; Costolon to Smoky Creek, 21 May 1955, Warnock 

& Wallmo 13215; Santa Elena Canyon, 21 Oct 1937, Parks & Cory 26445; Chisos 

Mts., Aug 1935, Marsh 80-118, 26 Jun 1937, Marsh 30] 

Boerhavia gracillima Heimerl Slimstalk Spiderling 

[Sperry, 1938; Reed, 1969] 

[Brewster Co.: Big Bend Natl. Pk., 12 Aug 1966, Hess 820; Todd Hill to Burro Mesa, 

01 Aug 1955, Warnock 12820; Pinnacle Mts. in the Chisos Mts., 22 Jul 1937, Warnock 

1080; Chisos Mts., 20 Aug 1931, Mueller 7992, 21 Jun 1937, Sperry 1937; Fisk Canyon 

and Fresno Springs (Sperry, 1938); Hot Springs, BBNP, 29 DEG 11’N, 102 DEG 59’ 

30”W, 1800 ft., 05 Nov 1988, Hodgson 5250 (UTEP, DES); 1.9 mi. S Study Butte on 

hwy. 118, 22 Aug 1982, Van Devender s.n. (UTEP)] 

Boerhavia intermedia M. E. Jones Spreading Spiderling 

[Sperry, 1938; eed, 1969] 

[Brewster Co.: Oak Canyon, 06 Sep 1933, Wolff 4413; 03 Aug 1955, Warnock 

12810; Hot Springs area, 08 Aug 1946, Shinners 8820; 08 Jun 1937, Warnock 1235, 

03 Sept 1940, Sperry 1782, 08 Jun 1937, Sperry 697, 24 Jun 1941, Warnock 21171; 

Chisos Mts., 31 Jun 1938, Sperry 1454, 1455, 01 Aug 1937, Marsh 211; et al. in 

Reed (1969); Tornillo Creek, Smoky Creek, Fresno Springs and Boquillas (Sperry, 

1938)]

Boerhavia linearifolia Gray 

[Reed, 1969] 

[Brewster Co.: Hot Springs, BBNP, 29 Jun 1941, Warnock 21670] 

Boerhavia purpurascens Gray Purple Spiderling 

[Reed, 1969] 

[Brewster Co.: Mt. Emory, 09 Sep 1933, Parks & Cory 7130; upper Green Gulch, 

14 Sep 1941, Sperry 1960] 

Boerhavia scandens L. Climbing Wortclub 

[Commicarpus scandens (L.) Standl.] 

[Reed, 1969] 

[Brewster Co.: Burro Mesa, BBNP, 23 Sep 1966, Correll 33845; Ward Spring, 

Chisos Mts., 18 Aug 1955, Warnock 13091] 

Boerhavia spicata Choisy Spicate Spiderling 

[Boerhavia torreyana (Wats.) Standl.] 


[Brewster Co.: Hot Springs, BBNP, 13 Sep 1931, Correll & Johnston 24558, 

10 Aug 1955, Warnock 13057; Santa Elena Canyon, 06 Aug 1946, Shinners 8798; 

Boquillas Canyon, 31 Oct 1948, Warnock & Turner 8315; Boquillas (Sperry, 1938); 

Tornillo Creek, 1883, Havard 63] 

Boerhavia wrightii Gray Wright Spiderling 


[Brewster Co.: Costolon Peak, BBNP, 23 Sep 1966, Correll 33836; Tornillo Creek 

at Banta Shut-in, 02 Aug 1937, Warnock 1235, 11 Aug 1941, Warnock 20346; near 

Roy’s Peak (Sperry, 1938); near great canyon of the Rio Grande, 1883, Havard 62 

(Type of B. bracteosa)] 

Cyphomeris gypsophiloides (Mart. & Gal.) Standl. Red Cyphomeris 

[Sperry, 1938; Reed, 1969] 

[Brewster Co.: Mt. Emory, Chisos Mts., 10 Sep 1915, Young s.n.; lower Blue 

Creek Canyon, 07 Sep 1933, Wolff 4511; et al. in Reed (1969); Saddle and Willow 

Creek Canyon (Sperry, 1938)] 

Mirabilis glabrifolia (Ortega) I. M. Johnst. 


Mirabilis grayana (Standl.) Standl. Gray’s Four O’Clock 

[Reed, 1969] 

[Brewster Co.: Blue Creek, BBNP, 23 Sep 1966, Correll 33830, 09 Oct 1936, 

Tharp s.n.; Muskhog Spring, Chisos Mts., 11 Jun 1937, Warnock & Sperry 692] 

Mirabilis linearis (Pursh) Heimerl Linearleaf Four O’Clock 


[Brewster Co.: BBNP, 29 Aug 1964, Hanson 530; Nail Place, Chisos Mts., 

21 Aug 1915, Young s.n.; Chisos Mts. (Sperry, 1938)] 

Mirabilis longiflora L. 

[Mirabilis longiflora L. var. wrightiana (Gray) Kearney & Peebles] 

[Le Duc, 1995] 

[Brewster Co.: Expected: distributed to the Big Bend area of Texas (Le Duc, 1995)] 

Mirabilis nyctaginea (Michx.) McMillan 


Mirabilis oblongifolia (Gary) Heimerl 

[Mirabilis albida (Walt.) Heimerl. in part; Mirabilis comata (Small) Standl.; 

Mirabilis grayana (Standl.) Standl.; Allionia melanotricha Standl.; Allionia 

Pratensis Standl. ] 


[Brewster Co.: Lost Mine Peak, 12 Sep 1961, Correll & Johnston 24506; 

Mt. Emory, Chisos Mts., 11 Jul 1955, Warnock 12377; upper Green Gulch, Chisos 

Mts., 31 Aug 1938, Warnock 20234; et al. in Reed (1969); Chisos Mts., Moore & 

Steyermark 3342, 3180] 

Mirabilis oxybaphoides (Gray) Gray Spreading Four O’Clock

[Sperry, 1938; Reed, 1969; Spellenberg & Rodriguez Tijerina, 2001] 

[Brewster Co.: Lost Mine Peak, BBNP, 12 Sep 1961, Correll & Johnston 24516 (GH, 

TEX-LL); Mt. Emory Trail, 27 Aug 1964, Hanson 526B; Mt. Emory slopes, 09 Sep 1933, 

Wolff 4650; Chisos Mts., 23 Aug 1931, Mueller 7994 (GH); upper slopes of Casa Grande 

(Sperry, 1938)] 

Mirabilis pseudaggregata Heimerl 

[Reed, 1969] 

[Brewster Co.: Boot Springs, Chisos Mts., 10 Sep 1950, Warnock 9756] 

Nyctaginea capitata Choisy Scarlet Muskflower 

[Sperry, 1938; Reed, 1969] 

[Brewster Co.: Maravillas Creek, Dog Canyon, 53 mi. SE of Marathon, 18 Jun 1961, 

Warnock & Wallmo s.n.; Chisos Mts., 21 Jun 1936, Sperry 155] 

Selinocarpus angustifolius Torr Narrowleaf Moonpod 

[Steyermark & Moore, 1933; Sperry, 1938; Reed, 1969; Fowler & Turner, 1977, mapped] 

[Brewster Co.: Talley Mt., BBNP, 10 May 1955, Wallmo 43; Santa Elena, 17 Jul 

1952, Webster 4372; et al. in Reed (1969); Persimmon Gap (Sperry, 1938); Santa 

Elena Canyon, Moore & Steyermark 3307] 

Selinocarpus parvifolius (Torr.) Standl. Littleleaf Moonpod 

[Standley, 1909; Sperry, 1938; Reed, 1969; Fowler & Turner, 1977, mapped] 

[Brewster Co.: Santa Elena Canyon, 22 Jul 1957, Correll & Johnston 18338; 

Tornillo Creek above Hot Springs, 22 Apr 1996, Correll & Rollins 32655; Bone 

Spring and Tornillo Creek, Aug 1883, Havard s.n.; et al. in Reed (1969); Muskhog 

Spring (Sperry, 1938); 0.5 mi. E Tornillo Creek on rd. to Rio Grande Village, BBNP, 

21 Aug 1982, Van Devender s.n. (UTEP)] 

OLEACEAE (Olive Family) 

Forestiera angustifolia Torr. Narrowleaf Forestiera ; Desert Olive 

[Sperry, 1938; Wells, 1965] 

[Brewster Co.: Dead Horse Mts. (Wells, 1965); Persimmon Gap and Boquillas 

(Sperry, 1938)] 

Forestiera pubescens Nutt. Elbowbush; Cruzilla 

[Forestiera neomexicana Gray] 


[Brewster Co.: Boot Spring, South Rim and Blue Creek (Sperry, 1938)] 

Fraxinus berlandieriana DC. Mexican Ash 

[Brewster Co.: 3 mi. above Canyon de Tale, BBNP, 02 Apr 1976, Everitt s.n. 

(UTEP)] 

Fraxinus cuspidata Torr. Fragrant Ash; Flowering Ash 


[Brewster Co.: Oak Creek Canyon, Casa Grande, Fresno Canyon and Juniper 

Canyon (Sperry, 1938); Dead Horse and Chisos Mts. (Powell, 1998)] 

Fraxinus greggii Gray Gregg Ash; Littleleaf Ash; Escobilla 

[Sperry, 1938; Little, 1952; Wells, 1965; Powell, 1998] 

[Brewster Co.: Dead Horse Mts. (Wells, 1965); Pinnacle Mts., Basin, Boot Canyon 

and Blue Creek (Sperry, 1938); Lost Mine Trail, BBNP, 6500 ft., 26 Jun 1978, Freeman 

s.n. (UTEP); Chisos Mts., Warnock 1078 (US)] 

Fraxinus velutina Torr. Velvet Ash; Arizona Ash 


[Brewster Co.: Rio Grande Village, BBNP (Powell, 1998)] 

Menodora decemfida (Gill.) Gray var. longifolia Steyermark 



Menodora longiflora Gray Showy Menodora 

[Sperry, 1938; Turner, 1991, mapped]

[Brewster Co.: Oak Creek Canyon near CCC camp (Sperry, 1938)] 

Menodora scabra Gray Rough Menodora 

[Steyermark & Moore, 1933; Sperry, 1938; Lewis, Stripking & Ross, 1962; Turner, 1991, 

mapped] 

[Records: flats below Pine Canyon, hills near Stillwell’s Ranch, on 

limestone hill in Basin, flats near Boquillas, top of Baldy Mtn., Comanche 

Springs area and between Chisos Mts. and Rio Grande (Sperry, 1938); Chisos 

Mts., Moore & Steyermark 3256; BBNP, Ross 52 (BRIT; TEX-LL)] 

ONAGRACEAE (Evening Primrose Family) 

Calylophus hartwegii (Benth.) Raven 

ssp. hartwegii 

[Sperry, 1938; Towner, 1977] 

[Brewster Co.: Comanche Spring area (Sperry, 1938); near Solis, just N of Mariscal 

Canyon, Johnston & Correll 24568 (TEX-LL)] 

Calylophus tubicula (Gray) Raven 


[Brewster Co.: near Persimmon Gap (Sperry, 1938)] 

Gaura boquillensis Raven & Gregory 

[Raven Y Gregory, 1972] 

[Brewster Co.: Bowuillas Canyon, 1919, Harrison (WISC), 1937, Warnock (TAES), 

Cory 2139 (RSA-POM); BBNP, Webster 4481 (DUKE, GH, OKLA, BRIT); mouth of 

Santa Elene Canyon, A. & R. A. Nelson 5052 (GH), Innes et al. 503 (GH); banks of Rio 

Grande in Grand Canyon, Palmer 34224 (MIN, NY); Pinnacle Mts., Chisos Mts. area, 

Warnock 1055 (US); mouth of Boquillas Canyon, 20 Apr 1960, Gregory 232 (Type: 

RSA-POM, Isotype: NY)] 

Gaura coccinea Nutt. ex Pursh Scarlet Gaura 

[Sperry, 1938; Raven & Gregory, mapped] 

[Brewster Co.: Terrapin Tank and lower Green Gulch (Sperry, 1938)] 

Gaura macrocarpa Rothrock 

[Sperry, 1938; Munz, 1938; Raven & Gregory, 1972] 

[Brewster Co.: Boquillas Canyon, Rosillas Mts. and Pinnacle Mts. (Sperry, 1938); Bois 

D’Arc Spring, Warnock 127 (PENN); Boquillas, Cory 2139 (RSA-POM)] 

Gaura mollis James Lizardtail 

[Gaura parviflora Hook.] 

[Raven & Gregory, 1972, mapped] 

Ludwigia peploides (Kunth) Raven ssp. glabrescens (Kuntze) Raven Yellow Water weed 

[Jussiaea repens L.] 


[Brewster Co.: Boquillas and San Francisco Canyon (Sperry, 1938)] 

Oenothera brachycarpa Gray Shortpod Evening Primrose 



Oenothera kunthiana (Spach) Nunz 

[Munz, 1932; Sperry, 1938] 

[Records: Tornillo Creek near Hot Springs and Dog Flats (Sperry, 1938); Glen Spring, 

Cory 2116 (RSA-POM)] 

Oenothera rosea L’Her. ex Ait. 


[Brewster Co.: lower Green Gulch and Terrapin Tank (Sperry, 1938)] 

Oenothera triloba Nutt. 


[Brewster Co.: Persimmon Gap to Tornillo Flats and Tornillo Creek bed near 

Hot Springs (Sperry, 1938)]

OROBANCHACEAE (Broomrape Family) 

Conopholis alpina Liebm. var. mexicana (Gray ex S. Wats.) Haynes Mexican Squaw-root 

[McVaugh, 1952, mapped; Thieret, 1969; Jones & Jones, 1993] 

[Records: Casa Grande Mt., BBNP, 26 May 1968, Haynes 2694; Pinnacle’s trail 

towards Boot Springs (Jones & Jones, 1993)] 

Orobanche ludoviciana Nutt. Louisiana Broomrape 


[Brewster Co.: “Boquillas, San Vicente,” Sperry (1938)] 

OXALIDACEAE (Wood Sorrel Family) 

Oxalis corniculata L. var. wrightii (Gray) Turner 

[Oxalis wrightii Gray] 


Oxalis drummondii Gray Drummond Wood Sorrel 

[Oxalis amplifolia auct. non (Trel.) Kunth] 

[Sperry, 1938; Denton, 1973] 

[Brewster Co.: Pulliam Canyon and near upper Cattail Falls (Sperry, 1938); Chisos Mts., 

Pulliam Canyon, 26 Aug 1936, Sperry 290 (US); Chisos Mts., rim rock, 29 Jun 1932, 

Muller (GH, MICH); 2 mi. SW of Boot Spring, 21 Jun 1931, Moore & Steyermark 3182 

(GH; MICH)] 

Oxalis stricta L. 



Oxalis violacea L. Violet Wood Sorrel 

[Sperry, 1938; Denton, 1973] 

[Brewster Co.: Chisos Mts. (Sperry, 1938); Chisos Mts., 29 Hun 1932, Muller; 

Chisos Mts., Lost Mine Peak, 15-18 Jul 1921, Ferris & Duncan 2869 (NY); Chisos Mts., 

17 Jun 1931, Muller 8052 (US)] 

PAPAVERACEAE (Poppy Family) 

Argemone chisosensis G. Ownbey 

[Sperry, 1938, as Argemone platyceras Link & Otto; Ownbey, 1958] 

[Brewster Co.: Rio Grande, Basin and Pulliam Bluff (Sperry, 1938); Chisos Mts., 

Juniper Canyon, 15-18 Jul 1921, Ferris & Duncan 3003 (NY Type, Isotypes CAS, DS, MO); 

Chisos Mts., Oak Canyon, Cory 6994 (GH); 25 mi. N of Boquillas, Hanson 588 (NY, US); 

Chisos Mts., Marsh 107 (F); The Basin, Chisos Mts., Sperry 383 (US); east of Nugent Mt., 

Sperry 1369 (US); Glenn Springs, Warnock 146 (TEX-LL)] 

Argemone sanguinea Greene 

[Argemone platyceras Link & Otto var. rosea Coulter] 


[Brewster Co.: near Johnson’s Ranch and near Nugent Mtn. (Sperry, 1938)] 

PEDALIACEAE [Sesame Family] 

Proboscidea louisianica (P. Mill.) Thellung ssp. louisianica 

[Bretting, 1983, mapped] 

Proboscidea parviflora (Woot.) Woot. & Standl. New Mexico Devil’s Claw 



PHILADELPHACEAE (Mock Orange Family)

Fendlera rigida I. M. Johnst. Narrowleaf Fendlerbush 

[Correll, 1965, as Fendlera linearis Rehd.; Powell, 1998] 

[Brewster Co.: Chisos Mts., on Crown Mtn. (Powell, 1998); Chisos Mts., near top of 

Nugent Mtn. 16 Jun 1964, Correll et al. 29729 (TEX-LL)] 

Philadelphus microphyllus Gray Littleleaf Mock Orange 

[Sperry, 1938; Hu, 1954-1956; Powell, 1998] 

[Brewster Co.: Chisos Mts. (Powell, 1998); N slope Mt. Emory (Sperry, 1938); Ferris & 

Duncan 2856 (MO; NY); Marsh 101 (F); Moore & Steyermark 3160 (A, GH, MO); Mueller 

8013 (A; F; MO); Warnock 866 (GH; US)] 

Philadelphus occidentalis A. Nels. 

[Hu, 1954-1956; Powell, 1998] 

[Brewster Co.: Chisos Mts. on NE slope Casa Grande (Powell, 1998); Chisos Mts., Cory 

7077 (GH)] 

PHYTOLACCACEAE (Pokeweed Family) 

Rivina humilis L. Pigeon Berry 



PLANTAGINACEAE (Plantain Family) 

Plantago ovata Forsk. 

[Plantago insularis Eastw. var. fastigiata (Morris) Jepson] 

[Correll, 1966] 

[Records:beds near Terlingus, BBNP, 18 Apr 1961, Correll & Rollins 23631 (TEX-LL); 

mouth of Tornillo Creek at Hot Springs, 22 Apr 1966, Correll & Rollins 32642 (TEX-LL)] 

Plantago patagonica Jacq. Bristle Bract Plantain; Woolly 

[Plantago parishii R. & S.] Plantain 


[Brewster Co.: Tornillo Creek near Hot Springs and Tornillo Flats (Sperry, 1938)] 

POLEMONIACEAE (Phlox Family) 

Gilia incisa Benth. 

[Steyermark & Moore, 1933; Sperry, 1938; Wherry, 1967; Turner, 1994, mapped] 

[Records: Green Gulch, Oak Canyon and Cattail Falls area (Sperry, 1938); 


Gilia insignis (Brand) Cory & Parks 

[Shinners, 1963; Wherry, 1967; Turner, 1994, mapped] 

[Records: 3 mi. S Persimmon Gap, 05 Apr 1947, McVaugh 7831 (BRIT); near Dog 

Flat, 02 Apr 1938, Warnock 21485; Tornillo Creek, Degenhardt 115; Persimmon Flat, 

Warnock 21566] 

Gilia laxiflora (Coult.) V. Grant 


[Records: S slope of Pulliam Bluff (Sperry, 1938)] 

Gilia rigidula Benth. ssp. acerosa (Gray) Wherry 

[Sperry, 1938; Wherry, 1967; Turner, 1994, mapped] 

[Records: Muskhog Spring area (Sperry, 1938)] 

Gilia stewartii I. M. Johnston 

[Shinners, 1963; Wherry, 1967; Turner, 1994, mapped] 

[Records: between Lone Mtn. and Nugent Mtn., BBNP, 03 Sep 1947, Warnock 7161 

(BRIT); 6 mi. N Hot Spring, BBNP, Rise-Innes & Warnock 546; McKinney Spring, Warnock 

10761; Santa Elena Canyon, Cory 44037; Muskhog Spring, Warnock 677] 

Ipomopsis aggregata (Pursh) V. Grant

[Sperry, 1938; Wherry, 1967] 

[Records: Boot Springs, Parks & Cory 7195; Burnt House Canyon, Warnock 156; 

Casa Grande to Mt. Emory, Sperry 458; Chisos Mts., Marsh 216, Mueller 8133; just past 

Lost mine trail on rd. to Basin, BBNP, summer 1983, collector unknown (UTEP); near 

Basin Campground, BBNP, Aug 1983, Stafford s.n. (UTEP)] 

Note: Pehhaps more than one ssp. is represented in the Chisos Mts.; clearly represented is 

ssp. formosissima (Greene) Wherry. 

Ipomopsis havardii (Gray) V. Grant 

[Sperry, 1938, as Ipomopsis havardii; Kearney & Peebles, 1943; Wherry, 1967; Powell, 1998] 

[Records: N end Mariscal Mtn. (SRSC); Avery Canyon, BBNP, 23 Jul 1950, Warnock 

9141 (BRIT); Dog Canyon Flats and Tornillo Flats (Sperry, 1938); vicinity of Chisos 

Mts., Sperry 623] 

Ipomopsis longiflora (Torr.) V. Grant ssp. longiflora 


[Records: Santa Elena Canyon, Moore & Steyermark 3305] 

Loeselia greggii S. Wats. 

[Loeselia scariosa (Mart. & Gal.) Walp.] 

[Steyermark & Moore, 1933; Sperry, 1938; Shinners, 1963; Van Schaack & Johnston, 

1964; Wherry, 1967; Turner, 1994, mapped] 

[Records: Wade Canyon, Chisos Mts., 30 Aug 1935, Marsh 12 (F; TEX-LL); Green 

Gulch, Sperry 361; Lost Mine Peak, Correll & Johnston 24553; Blue Creek Canyon, 02 Jul 

1931, Moore & Steyermark 3344 (MO); Panther Pass, 1958, Van Schaack, Warnock] 

Loeselia mexicana (Lam.) Brand 

[Wherry, 1967] 

[Records: Big Bend region, Apr 1930, Clark s.n., 01 Apr. 1930, Quillin s.n.] 

Phlox mesoluca Greene 

[Sperry, 1938; Wherry, 1967] 


Phlox nana Nutt. 


Phlox triovulata Thurb. ex Torr. 


POLYGALACEAE (Milkwort Family) 

Polygala alba Nutt. 

[Lewis & Davis, 1962] 

[Records: BBNP, Lewis & Oliver 5455 (BRIT)] 

Polygala barbeyana Chod. 

[Polygala longa Blake] 


[Records: near Green Gulch and W of Dead Horse Mts. (Sperry, 1938)] 

Polygala lindheimeri Gray 

[Polygala tweedyi Britt. ex Wheelock] 

[Steyermark & Moore, 1933; Sperry, 1938; Wendt, 1979, mapped] 

[Records: above Glenn Spring, Fresno Springs area, Chisos Pens, Burro 

Mesa and flats N of Chisos proper (Sperry, 1938); 0.8 mi. NW of Tunnel near 

Rio Grande Village, 20 Aug 1982, Van Devender s.n. (UTEP); Ernst Tinaja, 

BBNP, 21 Aug 1982, Van Devender s.n. (UTEP); Chisos Mts., Moore & Steyer- 

mark 3276] 

Note: Wendt (1979) mapped vars. linderheimeri and parvifolia Wheelock as occurring 

in BBNP. 

Polygala macradenia Gray Glandleaf Milkwort 


[Records: near Government Spring, Tornillo Flats and Roy’s Tank (Sperry, 1998); 

Chisos Mts., Moore & Steyermark 3274]

Polygala maravillasensis Correll 


[Records: in crevaces of ledges on summit of mountains west of Maravillas Creek, 

about 2 miles from mouth of Maravillas canyon, 24 Jun 1964, Correll & Hanson 29877 

(Type: TEX-LL; Isotypes: US; GH; TEX-LL)] 

Polygala nudata Brandegee 

[Wendt, 1979] 

[Records: Blue Creek, Cory 6999 (GH)] 

Polygala obscura Benth. 

[Polygala puberula Gray] 


[Records: Chisos Mts. (Sperry, 1938); Chisos Mts., Moore & Steyermark 3177, 3361] 

Polygala scoparioides Chod. 

[Steyermark & Moore, 1933; Sperry, 1938; Lewis & Davis, 1962] 

[Records: Basin, Lechuguilla Flats and near Lone Mtn. (Sperry, 1938); Chisos Mts., 

Moore & Steyermark 3251, 3273; Chisos Mts., Lewis 5545 (BRIT); Panther Junction, 

Lewis 5546 (BRIT)] 

POLYGONACEAE (Knotweed Family; Smartweed Family) 

Eriogonum abertianum Torr. Abert Wildbuckwheat 

[Eriogonum pinetorum Greene] 


[Brewster Co.: lower Oak Creek, lower Green Gulch and arroyos near Lone 

Mtn. (Sperry, 1938)] 

Eriogonum hemipterum (Torr. & Gray) Stokes 

[Eriogonum hieracifolium Benth. f. atropurpureum Standl.] 

[Sperry, 1938; Reveal, 1968; Correll & Johnston, 1970; Hess & Reveal, 1976] 

[Brewster Co.: Chisos Mts., Laguna Meadow trail to south, 3, Sep 1967, Reveal & Davidse 906 

(ARIZ, BRY, CAS, COLO, CS, GH, ISC, KSC, TEX-LL, MO, NY, RM, RSA-POM, BRIT, UC, 

US, UTC); Oak Creek Canyon, Willow Creek Canyon and Basin along trail to Laguna (Sperry, 

1938); Oak Creek near Window, 23 Aug 1936, Hinckley 835 (GH, TEX-LL, NY); Laguna, 28 Jul 

1932, Muller 46555 (MO, BRIT, TAES, TEX-LL, US); Oak and Willow Creek Canyons, 26 Sep 

1936, Sperry 216 (NEB, TAES, US)] 

Eriogonum jamesii Benth. var. undulatum (Benth.) S. Stokes ex M. E. Jones 

[Eriogonum undulatum Benth.] 

[Sperry, 1938; Reveal, 1968] 

[Brewster Co.: upper Green Gulch, Chisos Mts., Reveal & Davidse 906 (UTC); lower 

Pulliam Canyon and near top of Mt. Emory (Sperry, 1938)] 

Eriogonum polycladon Benth. 


[Brewster Co.: Painted Gap (Sperry, 1938)] 

Eriogonum suffruticosum S. Wats. 

[Reveal, 1968; Powell, 1998] 

[Brewster Co.: 5 mi. NW Nine Points Mesa, Muller 5197a, 5197b (TEX-LL; BRIT); 

“endemic … S Brewster and Presidio counties … common mostly in southern Brewster 

County” (records at SRSC)] 

Eriogonum tenellum Torr. var. platyphyllum (Torr. ex Benth.) Torr. 


[Brewster Co.: Pulliam Canyon, Green Gulch, slopes of Casa Grande, Banta Shut-in 

and Robbers’ Roost (Sperry, 1938)] 

Eriogonum wrightii Torr. ex Benth. Wright Wildbuckwheat 


[Brewster Co.: Green Gulch (Sperry, 1938); upper end Lost Mine Trail, BBNP, 

6500 ft., 26 Jun 1978, Freeman s.n. (UTEP)] 

Polygonum hydropiperoides Michx.

[Polygonum persicaroides H.B.K.] 



Polygonum lapathifolium L. Curltop Smartweed 


[Brewster Co.: Hot Springs (Sperry, 1938)] 

Polygonum pensylvanicum L. Pensylvania Smartweed 


[Brewster Co.: Boquillas Canyon (Sperry, 1938)] 

Polygonum persicaria L. var. persicaria Lady’s Thumb 

[Persicaria vulgaris Webb & Moq.] 



Rumex crispus L. 



Rumex salicifolius Weinm. var. mexicanus (Meisn.) C. L. Hitchc. 

[Rumex mexicanus Meisn.] 

[Brewster Co.: Rio Grande near San Vicente Canyon (Sperry, 1938)] 

PORTULACACEAE (Purslane Family) 

Portulaca oleracea L. Common Purslane 

[Portulaca retusa Engelm.] 


[Brewster Co.: lower Juniper Spring and flats north of Chisos Mts. (Sperry, 1938)] 

Portulaca pilosa L. Shaggy Portulaca 

[Sperry, 1938; Matthews, Ketron & Zane, 1992] 

[Brewster Co.: sandy flats N of Chisos Mts. (Sperry, 1938); BBNP, 29 Sep 1990, 

Powell & Powell 5650 (UNCC)] 

Portulaca suffrutescens Engelm. Shrubby Portulaca 


[Brewster Co.: sandy flats N of the Chisos Mts. (Sperry, 1938)] 

Talinum aurantiacum Engelm. Orange Flame-flower 


[Brewster Co.: below lower Juniper Spring (Sperry, 1938)] 

Talinum paniculatum (Jacq.) Gaertn. 


[Brewster Co.: upper Cattail Canyon (Sperry, 1938)] 

Talinum parviflorum Nutt. 



Talinum pulchellum Woot. & Standl. 



PRIMULACEAE (Primrose Family) 

Samolus ebracteatus H.B.K. var. cuneatus (Small) Henrickson Brookweed 

[Sperry, 1938; Henrickson, 1983, mapped] 

[Brewster Co.: Boquillas and Babb’s Hut in the Pinnacle Mts. (Sperry, 1938)] 

Samolus valerandi L. ssp. parviflorus (Raf.) Hulten 

[Samolus floribundus Kunth] 


[Brewster Co.: Gano Spring and lower Cattail Falls (Sperry, 1938)]

RANUNCULACEAE (Crowfoot Family) 

Anemone tuberosa Rydb. var. texana Enquist & Crozier 

[Enquist & Crozier, 1995] 

[Brewster Co.: E slope of Pulliam above Moss Well, BBNP, 21 Mar 1967, Warnock 

21142 (SRSC); Dog Canyon, BBNP, 08 Apr 1958, Warnock & Johnston 15986 (SRSC); 

Chisos Mts., 15 Mar 1949, Warnock 442 (TEX-LL)] 

Aquilegia chrysantha Gray Co;umbine 

[Steyermark & Moore, 1933; Sperry, 1938, var. longissima] 

[Brewster Co.: Pine and Pulliam Canyons (Sperry, 1938); Chisos Mts., Moore & 


Clematis drummondii Torr. & Gray Texas Virginsbower 

[Sperry, 1938; McVaugh, 1952, mapped] 

[Records: Near the Rio Grande, Oak Creek Canyon and base of Lone Mtn. 

(Sperry, 1938)] 

Clematis pitcheri Torr. & Gray var. pitcheri Leatherflower 



Delphinium wootonii Rydb. Plains Larkspur 

[Sperry, 1938, as Delphinium carolinianum Walt.] 

[Brewster Co.: Tornillo Creek N of Chisos Mts. (Sperry, 1938)] 

Note: Correll & Johnston (1970) do not report D. carolinianum from the Trans-Pecos 

hence the assignment to the above taxon. 

RHAMNACEAE (Buckthorn Family) 

Adolphia infesta (Kunth) Meisn. Texas Adolphia 

[Sperry, 1938; Johnston & Johnston, 1969] 

[Brewster Co.: Basin of Chisos Mts., 18 Jun 1937, Warnock 573; Oak Creek Canyon, 

(Sperry, 1938)] 

Ceanothus greggii Gray Desert Ceanothus 

[Sperry, 1938; Wells, 1965; Johnston & Johnston, 1969; Jones & Jones, 1993; Powell, 1998] 

[Brewster Co.: Chisos Mts., 22 Jun 1931, Muller 8016, and others; Pinnacle’s 

trail towards Boot Springs (Jones & Jones, 1993); Dead Horse Mts. (Wells, 1965)] 

Colubrina texensis (Torr. & Gray) Gray var. texensis Texas Colubrina 

[Records: Panther Canyon, BBNP, 05 Apr 1976, Everitt 76068 (UTEP)] 

Condalia ericoides (Gray) M. C. Johnston Javalinabush 

[Sperry, 1938; Johnston, 1962; Johnston & Johnston, 1969] 

[Records: Buro Mesa, BBNP, 10 Apr 1956, Warnock 13867 (SRSC); Chisos, 18 Mar 1941, 

Warnock 470 (TEX-LL); Pulliam Bluff (Sperry, 1938)] 

Condalia viridis I. M. Johnston Green Condalia 

[Johnston, 1962; Johnston & Johnston, 1969; Powell, 1998] 

[Records: between Burro Mesa and Chisos Mts., 06 Jul 1931, Moore & Steyermark 3402 

(A; GH; MICH); Gano Spring, 16 Jul 1937, Marsh Marsh 92a (F); hills above Hot 

Spring, 29 Jun 1941, Warnock 20947 (TEX-LL); Tornillo Creek near Hot Spring, 20 Apr 1948, 

Whitehouse 19722 (MICH; SRSC; US); Chisos Mts., 12 Jun 1942, Whitaker 439 (US)] 

Condalia warnockii M. C. Johnston Warnock Condalia 

var. warnockii 

[Johnston, 1962; Johnston & Johnston, 1969] 

[Records: E base Chisos Mts., 19 Jun 1901, Bailey 397 (US); arroyo E of Burro Mesa, 

06 Jul 1931, Moore & Steyermark 3399 (CAS; DS); Chisos Mts., 02 Aug 1931, Muller 8099 

(A; GH; TEX-LL; US); Nail Place, Chisos Mts., 15 Jul 1931, Muller s.n. (GH; TEX-LL); 

Heath Canyon, Dead Horse Mts., 19 Jul 1952, Warnock 10814 (TEX-LL; SRSC); Sam 

Nail’s, E side of Burro Mesa, 12 Aug 1955, Warnock 12907 (SRSC; TAES); near Dugout,

4 Jul 1938, Sperry 1458 (US); Gano Spring, 16 Jul 1937, Marsh 92 (F)] 

Karwinskia humboldtina (Schult.) Zucc. Coyotillo 

[Sperry, 1938; Johnston & Johnston, 1969; Powell, 1998] 

[Brewster Co.: Boquillas Canyon (SRSC); Rio Grande at milage 796.8, 29 DEG 

15’30”n, 102 DEG 55’30”W, 1700 ft., 06 Nov 1988, Hodgson 5316 (UTEP)] 

Rhamnus betulifolia Greene Birchleaf Buckthorn 

[Sperry, 1938; McVaugh, 1952, mapped; Johnston & Johnston, 1969; Powell, 1998] 

[Records: “Chisos Mts., above Boot Spring; Pulliam Bluff; upper Pine Canyon” 

Powell (1998) (SRSC); Mt. Emory, 07 Aug 1947, Correll 13683] 

Rhamnus serrata Humb. & Bonpl. ex J. A. Schultes var. serrata 

[Sperry, 1938; Johnston & Johnston, 1969; Johnston, 1975; Johnston & Johnston, 1978; 

Powell, 1998] 

[Brewster Co.: upper Cattail Canyon and above Boot Spring (SRSC); Chisos Mts., Aug 

1893, Havard 129; Chisos Mts., 1200-2000 m, 09 Sep 1937 Marsh 354 (TEX-LL); upper 

Green Gulch, 22 Aug 1947, Warnock & Tharp 6846; Chisos Mts., 09 Aug 1915, Young 

s.n. (TEX-LL); BBNP, 2100 m, 13 Jul 1972, Chiang, Wendt & Johnston 8420f (TEX-LL); 

below Emory Peak, BBNP, 17 Jun 1964, Correll 29751 (TEX-LL) and Correll & Hanson 

29821 (TEX-LL); Chisos Mts. State Park, 30 Aug 1935, Marsh s.n. (TEX-LL); above Boot 

Springs, 2120 m, 21 Aug 1931, Moore & Steyermark 3161 (CA, US, UC); upper Green 

Gulch, 22 Aug 1947, Tharp 5100 (TEX-LL); Green Gulch, 22 Aug 1947, Tharp, Warnock 

& Hinckley (TEX-LL); Mt. Emory, 2100-2400 m, Warnock 1023, 6846 (TEX-LL); between 

Boot Spring and South Rim along upper Boot Canyon, 2100 m, 15 Jul 1955, Warnock 12703 

(TEX-LL)] 

Ziziphus obtusifolia (Hook. ex Torr. & Gray) Gray var. obtusifolia Lotebush 

[Sperry, 1938; Johnston, 1963, mapped; Wells, 1965; Johnston & Johnston, 1969] 

[Brewster Co.: Boquillas, 28 May 1901, Bailey 358; Chisos Mts., Aug 1935, Marsh 

208; et al. in Johnston & Johnston (1969); Dead Horse Mts. (Wells, 1965)] 

RESEDACEAE (Mignonette Family) 

Oligomeris linifolia (Vahl) J. F. Macbr. Linearleaved Cambess 

[Dipetalia subulata (Webb. & Berth.) Ktze] 


[Brewster Co.: Persimmon Gap and across Tornillo Flats (Sperry, 1938)] 

ROSACEAE (Rose Family) 

Cercocarpus montanus Raf. Mountain Mahogany 

[Sperry, 1938, as Cercocarpus eximius; Powell, 1998] 

Cercocarpus breviflorus Gray var. breviflorus Mountain Mahogany 


[Brewster Co.: Lost Mine Trail, BBNP, 6500 ft., 26 Jun 1978, Freeman s.n.(UTEP)] 

Note: Denham (1996) has published a partial new interpretation of Cercocarpus. 

Fallugia paradoxa (D. Don) Endl. ex Torr. Apacheplume 

[Sperry, 1938; Wells, 1965] 

[Brewster Co.: Green Gulch, Basin and Blue Creek (Sperry, 1938); Dead Horse 

Mts. (Wells, 1965)] 

Holodiscus discolor (Pursh.) Maxim. Bush Rockspires 

[Sperry, 1938, as Seriotheca dumosa; Powell, 1998] 

[Brewster Co.: Emory Peak (Powell, 1998)] 

Malacomeles denticulata (Kunth) Engelm. Toothed Serviceberry 

[Amelanchier denticulata (Kunth) Koch.] 


[Brewster Co.: Chisos Mts, and Dead Horse Mts. (Powell, 1998)] 

Prunus havardii (W. Wight) S. C. Mason Havard Plum


[Brewster Co.: Green Gulch, Window Trail, and Pine Canyon (Powell, 1998); 

Oak Creek Canyon (Sperry, 1938); Dead Horse Mts. (Wells, 1965); Blue Creek 

Canyon, Chisos Mts. Moore & Steyermark 3230 (MO)] 

Prunus serotina Ehrh. var. virens (Woot. & Standl.) McVaugh Southwestern Chokecherry 

[Sperry, 1938; McVaugh, 1951, 1952] 

[Records: Green Gulch and Pine Canyon (Sperry, 1938); Chisos Mts., Mueller (TEX-LL; 

F; MICH; NY); Chisos Mts., Moore & Steyermark 3333 (MICH; NY)] 

Purshia ericifolia (Torr. ex A. Gray) Henrickson Heath Cliffrose 

[Cowania ericifolia Torr. ex A. Gray] 

[Steyermark & Moore, 1933; Wells, 1965; Powell, 1998] 

[Brewster Co.: Brushy Canyon and above Roy’s Peak in Dead Horse Mts., near 

Mckinney Springs, 0.75 mi. NE of Dagger Flats visitor area (Powell, 1998); Boquillas, 


Rubus persistens Rydb. Persistent Blackberry 


[Brewster Co.: near Cattail Falls in the Chisos Mts. (Powell, 1998)] 

Rubus trivialis Michx. 

[McVaugh, 1952, mapped; Powell, 1998] 

Note: Powell (1998) mentions this taxon as being on the BBNP list created by B. Amos 

(1991). 

Vaquelinia corymbosa Humb. & Bonpl. ssp. angustifolia (Rydb.) Hess Slimleaf Vaquelinia 

& Henrickson 

[Steyermark & Moore, 1933; Sperry, 1938; Wells, 1965; Hess & Henrickson, 1987; Powell, 

1998] 

[Brewster Co.: Basin and Oak Canyon in Chisos Mts., Dead Horse Mts. (Powell, 1998); 

Basin 1 mi. below campground on trail to Window, BBNP, 26 May 1982, Van Devinder 

s.n. (UTEP); Basin at Saddle above Green Gulch, BBNP, 15 Jul 1981, Freeman s.n. (UTEP); 

ridge on E side of Juniper Canyon, Chisos Mts., 15-18 Jul 1921, Ferris & Duncan2927 

(CAS, DS, NY); Blue Creek Canyon, Chisos Mts., 23 Jun 1931, Moore & Steyermark 3203 

(AA, CAS, DS, MICH, NY, UC); Oak Creek Canyon near camp and the Window, Chisos 

Mts., 05 Jul 1937, Sperry 168 (US); Dead Horse Mts. near Sue Peak, 19 Jul 1952, Warnock 

10778 (TEX-LL)] 

RUBIACEAE (Madder Family) 

Bouvardia ternifolia (Cav.) Schlecht. Sacrlet Bouvardia 

[Sperry, 1938; Lewis, 1962; Powell, 1998] 

[Records: Basin and Green Gulch (Sperry, 1938); Lost Mine Trail, BBNP, Aug 

1983, Stafford s.n. (UTEP); upper Lost Mine Trail, 26 Jun 1978, Freeman s.n. (UTEP); 

BBNP, Ross 51 (BRIT)] 

Cephalanthus occidentalis L. Common Buttonbush 

[Sperry, 1938; McVaugh, 1952, mapped; Powell, 1998] 

[Records: Gano Springs, Hot Springs and Rio Grande near Mariscal 

(Sperry, 1938)] 

Galium aparine L. 

[Powell, 1959, mapped] 

Galium mexicanum Kunth ssp. flexicum Dempster 

[Steyermark & Moore, 1933; Dempster, 1958, 1976, 1978] 

[Records: Chisos Mts., 18 Aug 1931, Mueller 8281 (Type: TEX-LL; Isotypes: 

GH, MICH, NY); Chisos Mts., above Boot Spring, Moore & Steyermark 3159 

(this coll. reported as var. asperulum by Dempster in 1958)] 

Galium microphyllum Gray 

[Relbunium microphyllum (Gray) Hemsl.] 

[Powell, 1959, mapped] 

Galium pilosum Ait.

[Steyermark & Moore, 1933; Jones & Jones, 1993] 

[Records: Pinacle’s trail towards Boot Springs (Jones & Jones, 1993); Chisos 


Galium proliferum Gray Great Basin Bedstraw 

[Sperry, 1938; Powell, 1959, mapped] 

[Records: Avary Canyon (Sperry, 1938)] 

Galium uncinulatum DC. 

[Galium uncinulatum DC. var. obstipum (Schlecht.) S. Wats.] 

[Powell, 1959, mapped; Dempster, 1978] 

Galium wrightii Gray Wright Bedstraw 

[Galium rothrockii Gray] 

[Steyermark & Moore, 1933; Sperry, 1938; Powell, 1959, mapped] 

[Records: upper slopes of Casa Grande (Sperry, 1938); Chisos Mts., Moore & 


Hedyotis intricata Fosb. Tangle Bedstraw 

[Houstonia fasciculata Gray] 


[Brewster Co.: Dead Horse Mts. (Wells, 1965); Pulliam Bluff, Basin, South Rim 

and lower Juniper Spring (Sperry, 1938); upper slopes Green Gulch, 27 Aug 1944, 

Lundell 13256 (UTEP); Chisos Mts., Moore & Steyermark 3323] 

Houstonia acerosa (Gray) Gray ex Benth. & Hook. f. 

[Sperry, 1938; Lewis, 1962, 1968; Terrell, 1996, 2001] 

[Brewster Co.: Basin, Dead Horse Mts., and areas N of Chisos Mts. (Sperry, 1938); 

Chisos Mts., Moore & Steyermark 3261 (CAS, DS, MICH, NY); Limestone hills in Basin, 

Sperry 455 (US); Chisos Mts., Mueller 8170 (GH, var. polypremoides); BBNP, Lewis 5541, 

Lewis & Oliver 5464, Ross 53 (all BRIT as var. polypremoides)] 

Note: Terrell (2001) shows vars. acerosa and polypremoides (Gray) Terrell occurring in 

Brewster Co. 

Stenaria mullerae (Fosberg) Terrell var. pooleana (B. L. Turner) Terrell 

[Hedyotis pooleana Turner; Hedyotis mullerae Fosberg] 

[Turner, 1995; Terrell, 1996, 2001; Turner, 1997] 

[Brewster Co.: Dead Horse Mts., Brushy Canyon Preserve, 0.7 mi. SW of adobe house 

on Brush Canyon divide, 29 DEG 27’N, 102 DEG 58’W, 4900 ft., 01 May 1987, Poole 

2942 (Type: TEX-LL), 25 May 1985, Poole 2527 (SRSC, TEX-LL)] 

Note: Turner (1997) maintains that H. pooleana is a distinct species. The species has been 

observed on the same ridge at 4840 ft. in BBNP which is about one mile from the type 

locality. 

Stenaria nigricans (Lam.) Terrell var. nigricans Bluets 

[Houstonia angustifolia Michx.; Hedyotis nigricans (Lam.) Terrell var. nigricans] 

[Sperry, 1938; Terrell, 2001, systematics] 

[Brewster Co.: Basin, Green Gulch, Muskhog Spring and Glenn Spring (Sperry, 

1938); 0.8 mi. NW of Tunnel near Rio Grande Village, 20 Aug 1982, Van Devender 

s.n. (UTEP); BBNP, Lewis 5543 (BRIT)] 

RUTACEAE (Citrus Family) 

Ptelea trifoliata L. ssp. angustifolia Skunkbush; Common Hoptree 

[Steyermark & Moore, 1933; Sperry, 1938; Bailey, 1962; Bailey & Bailey, 1965; 

Powell, 1998] 

[Records: Pulliam Canyon, base of Casa Grande, Laguna and Cattail Canyon (Sperry, 1938); 

Chisos Mts., above Boot Spring, Moore & Steyermark 3184 (CAS; DS; MICH; MO; UC; 

US); Chisos Mts., Tharp (TEX-LL; UC), Mueller 8286 (MICH; TEX-LL; US), Fletcher 

11992 (TAES); Correll 13725 (BRIT); upper Boot, 1933, Parks & Cory (TAES); Mt. Emory, 

10 Sep 1915, Young (TEX-LL); Casa Grande, Webster 4407 (BRIT); near base of Cap, Warnock 

(TEX-LL); US); Lost Mine Peak, Ferris & Duncan 2857 (CAS; DS; MO); Laguna, Warnock 

867 (US)]

Note: Powell (1998) treats these as above following Correll & Johnston (1970). Bailey (1962) 

and Bailey and Bailey (1965) treats them as ssp. pallida (Greene) V. Bailey var. confinis (Greene) 

V. Bailey. 

Thamnosma texana (Gray) Torr. Texas Desertrue; Rueda del Monte 

[Thamnosma aldrichii Tharp] 

[Sperry, 1938; Tharp, 1945; Powell, 1998] 

[Brewster Co.: W of Dead Horse Mts., near Muskhog Spring and above Hot Springs 

(Sperry, 1938); Chhisos Mts., Mueller s.n. and Tharp s.n. (TEX-LL); above Hot Springs, 

Warnock 949 (TEX-LL)] 

SALICACEAE (Willow and Cottonwood Family) 

Populus angustifolia James Narrowleaf Cottonwood 


[Brewster Co.: ‘Santa Elena Crossing below Costolon, Boquillas Crossing, Rio Grande 

Village, Chilicotal Spring, and McKinney Spprings” (Powell, 1998)] 

Populus fremontii S. Wats. ssp. mesetae Eckenw. Arizona Cottonwood; Chopo 

[Sperry, 1938, as Populus palmeri Sarg.; Powell, 1998] 

[Brewster Co.: Rio Grande in BBNP (Powell, 1998)] 

Populus tremuloides Michx. Quaking Aspen 

[Sperry, 1938; Powell, 1998} 

[Brewster Co.: Emory Peak of Chisos Mts. (Sperry, 1938; Powell, 1998)] 

Salix exigua Nutt. ssp. exigua var. exigua Coyote Willow 


[Brewster Co.: near Boquillas (Powell, 1998)] 

Note: The Sperry (1938) record of S. longifolia from Solis’ ranch may be this 

species. 

Salix exilifolia Dorn Yellowleaf Willow 

[Salix taxifolia H.B.K.] 

[Sperry, 1938; Augus & McJannet, 1992, mapped; Dorn, 1998] 

[Brewster Co.: Rio Grande at Boquillas (Sperry, 1938)] 

Salix gooddingii Ball Goodding Willow 

[Sperry, 1938; Powell, 1998; Dorn, 1998, mapped] 

[Brewster Co.: along Rio Grande, Boquillas and Fisk Canyon (Sperry, 1938)] 

Salix lasiolepis Benth. Arroyo Willow 

[Johnston, 1981;Powell, 1998] 

[Brewster Co.: Chisos Mts. (Johnston, 1981; Powell, 1998)] 

Salix thurberi Rowlee 

[Salix longifolia Muhl. non Lam. vars.] 

[Schneider, 1919; Dorn, 1998, mapped] 

[Brewster Co.: Rio Grande S of Chisos Mts., Aug 1883, Havard] 

Note: Dorn (1998) cites a Chisos Mts. collection by Mueller 7965 (TEX-LL, NA) as 

intermediate between Salix exigua exigua and S. thurberi. 

SAPINDACEAE (Soapberry Family) 

Sapindus saponaria L. var. drummondii (Hook. & Arn.) L. Benson Western Soapberry 


[Brewster Co.: Nail’s Ranch (Sperry, 1938)] 

Ungnadia speciosa Endl. Mexican Buckeye 


[Brewster Co.: Green Gulch, Boot Spring and Pine Canyon (Sperry, 1938)] 

SAPOTACEAE (Sapodilla Family)

Sideroxylon lanuginosum Michx. ssp. rigidum (Gray) T. D. Pennington Woollybucket Sideroxylon 

[Bumelia lanuginosa (Michx.) Pers. var. rigidum Gray] 


[Brewster Co.: Dog Canyon (Powell, 1998)] 

SAXIFRAGACEAE (Saxigrage Family) 

Heuchera rubescens Torr. 


[Brewster Co.: slopes of Casa Grande and Mt. Emory (Sperry, 1938); Chisos Mts., 

Moore & Steyermark] 

SCROPHULARIACEAE (Figwort Family) 

Castilleja integra Gray var. integra 

[Castilleja elongata Pennell] 

[Sperry, 1938; Pennell, 1940; Nesom, 1992] 

Brewster Co.: limestone slopes in Basin (Sperry, 1938); Panther Jct., Scott 2 (SRSC); 

Chisos Mts., Boot Canyon, Steiger 1212 (NY); northern Chisos Mts., 6000 ft., Aug 1934, 

Steiger 1689 (NY, Type of C. elongata Penn.); Chisos Mts., Sperry 263 (US; SRSC)] 

Castilleja lanata Gray Wooly Paintbrush 

[Sperry, 1938; Nesom, 1992, mapped] 

[Brewster Co.: Tornillo Creek near Hulen’s Ranch (Sperry, 1938)] 

Castilleja nervata Eastw. 


[Brewster Co.: Pinnacles trail towards Boot Springs (Jones & Jones, 1993)] 

Castileja rigida Eastw. 

[Castilleja latebracteata Pennell] 

[Sperry, 1938; Pennell, 1940; Nesom, 1992, mapped] 

[Brewster Co.: Juniper Canyon, flats N and W of Chisos Mts., Muskhog Spring 

and Mesa de Anguila (Sperry, 1938); Chisos Mts., Palmer 34144 (PH; NY), Sperry 

459 (US)] 

Castilleja sessiliflora Pursh Downy Paintbrush 

[Nesom, 1992, mapped] 

Leucophyllum candidum I. M. Johnst. Boquillas Silverleaf 

[Leucophyllum violaceum Penn.] 

[Pennell, 1940; Wells, 1965; Henrickson & Flyr, 1985; Powell, 1998] 

[Brewster Co.: “generally between the Chisos Mts. and Boquillas Canyon, E to Black 

Gap refuge … “(Powell, 1998); Dead Horse Mts. (Wells, 1965); above Tunnel near 

Rio Grande Village, BBNP, 17 Aug 1983, Van Devender s.n. (UTEP); Boquillas, Hanson 590 

(MO; US), Hanson 712 (US; NY); Glenn Springs to base of Chisos Mts., Ferris & Duncan 2758a 

(US); Hot Springs area, Sperry 1522, 1579 (PH), Warnock 1124 (PH)] 

Leucophyllum frutescens (Berl.) I. M. Johnst. Ceniza, Purple Sage 

[Sperry, 1938; Pennell, 1940; Henrickson & Flyr, 1985; Powell, 1998] 

[Brewster Co.: “vicinity of Chisos Mts., especially toward Boquillas, also NE toward 

Marathon” (Powell, 1998); Rock Spring, Chisos Mts., Young (TEX-LL); Grapevine Hills, 

Steiger 731 (NY); near Tornillo Creek, Palmer 34062 (PH)] 

Leucophyllum minus Gray Big Bend Silverleaf 

[Sperry, 1938; Pennell, 1940; Henrickson & Flyr, 1985; Powell, 1998] 

[Brewster Co.: “throughout in limestone” (Powell, 1998); adjacent to Chisos Mts., 

Boquillas and Fresno Springs (Sperry, 1938); E of Chisos Mts., Sperry 1580 (PH); Grapevine 

Hills N of Chisos Mts., Steiger 732 (NY); Persimmon Gap, Sperry 1518 (PH)] 

Maurandella antirrhiniflora (Humb. & Bonpl. ex Willd.) Rothm. Blue Snapdragon Vine 

[Maurandya antirrhiniflora Humb. & Bonpl. ex Willd.]

[Sperry, 1938; Pennell, 1940; Elisens, 1985] 

[Brewster Co.: Rio Grande near Costolon, lower Oak Creek Canyon and Rosillos 

Mts. (Sperry, 1938); Chisos Mts., Marsh 36 (F, GH, TEX-LL, UC), Muller 8161 (TEX-LL; 

MO; PH)] 

Mecardonia procumbens (P. Mill.) Small 

[Mecardonia vandellioides (H.B.K.) Penn.; Pagesia peduncularis (Benth.) Penn.] 



Mimulus glabratus Kunth var. glabratus Monkeyflower 

[Sperry, 1938Pennell, 1940] 

[Brewster Co.: Tornillo Creek bed near Hot Springs, Sperry 512 (US; SRSC)] 

Nuttallanthus texanus (Scheele) D. A. Sutton Texas Toad Flax 

[Linaria texana Scheele] 



Penstemon baccharifolius Hook. Baccharisleaf Penstemon 

[Steyermark & Moore, 1933; Pennell, 1940; Powell, 1998] 

[Brewster Co.: “Doubtful Canyon; Dead Horse Mts., Big Brushy Canyon, upper Heath 

Canyon; Maravillas Creek” (Powell, 1998); Boquillas Canyon, Moore & Steyermark 

3450 (MO; NY)] 

Penstemon barbatus (Cav.) Roth. ssp. torreyi (Benth.) Keck 

[Steyermark & Moore, 1933; Sperry, 1938; Pennell, 1940] 

[Brewster Co.: Green Gulch, Willow Creek Canyon and near top of Casa Grande 

(Sperry, 1938); Chisos Mts., Moore & Steyermark; Chisos Mts., Muller 8162 (PH), 

Steiger 1243 (NY)] 

Penstemon dasyophyllus Gray 

[Steyermark & Moore, 1933; Crosswhite, 1966] 

[Brewster Co.: Oak Canyon, Moore & Steyermark 3360 (GH)] 

Penstemon fendleri Torr. & Gray Fendler Penstemon 



Penstemon havardii Gray 

[Sperry, 1938; Pennell, 1940] 

[Brewster Co.: Willow Creek Canyon, Oak Creek Canyon and arroyo near Lone 

Mtn. (Sperry, 1938); Basin near campground, Aug 1983, Stafford s.n. (UTEP); Chisos Mts., 

Havard 94 (GH; US); Palmer 34149 (MO; PH); 4 mi. W of Government Springs, Cory (PH)] 

Penstemon jamesii Benth. 

[Pennell, 1940] 

[Records: Santa Elena Canyon, Steiger 1251 (NY)] 

Penstemon ramosus Crosswhite 

[Penstemon lanceolatus auct. non Benth.] 

[Crosswhite, 1966] 

[Brewster Co.: head of Heath Canyon, Dead Horse Mts., BBNP, Warnock 10725 

(TEX-LL; BRIT)] 

Seymeria scabra Gray 

[Sperry, 1938; Pennell, 1940] 

[Brewster Co.: slopes of Mt. Emory and Flattop (Sperry, 1938); Chisos Mts., Havard (US), 

Steiger 1217 (NY)] 

SIMAROUBACEAE (Quassia Family) 

Holacantha stewartii C. H. Muell. Crucifixion Thorn 

[Correll, 1959; Powell, 1998] 

[Brewster Co.: Paint Gap Hills, Dagger Flat, 1 mi. S Government Springs and 

between Glenn Springs and Lindsey Mine (Powell, 1998); Paint Gap Hills, BBNP, 

3500 ft., 12 Aug 1995, Warnock 12903 (TEX-LL)]

SOLANACEAE (Potato Family) 

Calibrachoa parviflora (Juss.) D’Arcy Wild Petunia 

[Petunia parviflora Juss.] 


[Brewster Co.: creek bed in Pinnacle Mts. and Glenn Spring (Sperry, 1938)] 

?Capsicum annuum L. var. glabrisculum (Dun.) Heiser & Pickersgill Bird Pepper; Chilipiquin 

[Sperrt, 1938] 

Note: The collection Sperry referenced (#TM52) from Smoky Creek needs to be 

Located and reviewed. 

Chamaesaracha conoides (Moric. ex Dunal) Britt. 

[Sperry, 1938; Averett, 1973] 

[Brewster Co.: Hot Springs, near Comanche Spring and Tornillo Flats (Sperry, 

1938)] 

Chamaesaracha coronopus (Dunal) Gray Small Groundcherry 

[Sperry, 1938; Averett, 1967,1973, mapped] 

[Brewster Co.: Basin, Tornillo Flats and Persimmon Gap (Sperry, 1938); 

Persimmon Gap, BBNP, Sperry 566; Basin, BBNP, Averett 192] 

Chamaesaracha crenata Rydb. 

[Averett, 1967, 1973] 

[Brewster Co.: Glenn Springs, BBNP, 25 May 1955, Warnock & Wallmo 

12300 (SRSC, TEX-LL); 8 mi. NE of Costolon, BBNP, Averett & Sikes 238; Hot 

Springs area, BBNP, Sperry 1727; Dog Flat, BBNP, Warnock 13909; Burro Mesa, 

BBNP. Warnock 18284; Roy Peak in Dead Horse Mts., BBNP, Warnock & Johnston 

15909; head of Boquillas Canyon, BBNP, Warnock & Turner 8327; Santa Elena 

Canyon, BBNP, Webster 4363] 

Chamaesaracha pallida Averett Pale Falsenightshade 

[Averett, 1967,1973, mapped] 

[Brewster Co.: McKinney Spring, BBNP, Warnock 10853] 

Chamaesaracha sordida (Dunal) Gray Hairy Falsenightshade 

[Averett, 1967, 1973] 

[Brewster Co.: foothills of Chisos Mts., 19 May 1931, Tharp s.n. (GH, MICH, 

TEX-LL, UC); Onion Spring, BBNP, Warnock 12946; Smoky Creek, BBNP, 

Warnock & Wallmo 13222] 

Chamaesaracha villosa Rydb. 

[Averett, 1967] 

[Brewster Co.: Glenn Spring, BBNP, Warnock & Wallmo 12300] 

Datura quercifolia H.B.K. Oakleaf Thornapple 



Datura wrightii Regel Sacred Datura 

[Datura meteloides auct. non Dunal] 


[Brewster Co.: near Glenn Spring and Pinnacle Mts. (Sperry, 1938)] 

Lycium berlandieri Dun. Berlandier Wolfberry 


[Brewster Co.: Johnson’s Ranch to Costolon, Dead Horse Mts., Boquillas, 

Oak Creek Canyon and Mail Box Tank area (Sperry, 1938)] 

Note: Most recors are probably var. parviflorum (Gray) Terracc. 

Lycium pallidum Miers. Pale Wolfberry 

[Hitchcock, 1932; Sperry, 1938] 

[Brewster Co.: Chisos Pens (Sperry, 1998); Boquillas, 28 May 1901, Bailey 357 (US)] 

Lycium puberulum Gray var. berberioides (Correll) Chiang Silvery Wolfberry 

[Hitchcock, 1932; Wells, 1965; Correll, 1965; Powell, 1998] 

[Brewster Co.: Dead Horse Mts. (Wells, 1965); lower slopes of Chisos Mts.,

Sublett Ranch, 3-4 mi. S of Persimmon Gap, Study Butte to Panther Jct., et al. 

(Powell, 1998); foothills of Chisos and plains, Aug 1883, Havard 91 (US); 3 mi. S 

of Persimmon Gap, 15 Jun 1964, Correll et al. 29712 (Type of berberioides: TEX-LL; 

Isotypes: US, GH, TEX-LL); 3 mi. S of Persimmon Gap, 5 Apr 1947, McVaugh 7835 

(NA; BRIT); Chisos Mts., Juniper Canyon, 16 Jun 1964, Correll et al. 29714 (TEX-LL)] 

Lycium torreyi Gray Torrey Wolfberry 


[Brewster Co.: along Rio Grande (Powell, 1998)] 

Nicotiana glauca Grah. Tree Tobacco; Tronadora 


[Brewster Co.: along Rio Grande from Terlingua to Boquillas (Sperry, 1938)] 

Nicotiana trigonophylla Dunal Desert Tobacco; Tobaquillo 



Physalis hederifolia Gray Heartleaf Groundcherry 


[Brewster Co.: S Chisos Mts., Green Gulch and Muskhog Spring (Sperry, 1938); Santa 


Note: Several varieties are probably represented. 

Physalis philadelphica Lam. var. immaculata Waterfall 

[Sperry, 1938, as P. ixocarpa] 


Physalis solanaceus (Schlecht.) Axelius Netted Globeberry 

[Margaranthus solanaceus Schlecht.] 



Note: I have not been able to attribute the Steyermark and Moore (1933) collection 

reported as Physalis subintegra Fernald from the Chisos Mts. to any of the known 

taxa. 

Quincula lobata (Torr.) Raf. Purple Groundcherry 


[Brewster Co.: Dog Flats (Sperry, 1938)] 

Solanum davisense M. D. Whalen 

[Sperry, 1938, as S. nigrum; Whalen, 1976; Lemke, 1991] 

[Brewster Co.: slopes of Casa Grande and Boquillas (Sperry, 1938); Chisos Mts. 

(Lemke, 1991); Chisos Mts., between Boot Spring and South Rim, igneous soil, 7000 ft., 

15 Jul 1955, Warnock 12717 (SRSC)] 

Solanum elaeagnifolium Cav. Purple Nightshade 


[Brewster Co.: Rio Grande near Boquillas (Sperry, 1938)] 

Solanum rostratum Dun. Buffalo Bur 


[Brewster Co.: Green Gulch and Pine Canyon (Sperry, 1998)] 

Solanum sarrachoides Sendtner 

[Sperry, 1938, as S. villosum; Lemke, 1991] 

[Brewster Co.: Chisos Mts., Sperry 475 (SRSC) (Lemke, 1991)] 

Note: Sperry (1938) reports other collections (Nos. 683 and 1049) from the Chisos 

Mts. that may not have been seen by Lemke. 

Solanum triquetrum Cav. Texas Nightshade; Hierba Mora 


[Brewster Co.: in most mountains S of the Glass Mts. (Powell, 1998)] 

Note: The report of Solanum sisymbriifolium Lam. From Green Gulch (Sperry, 1938, 

no. 390) is unlikely since Lemke (1991) found only one sheet of this adventive species 

for all of Texas.

STERCULIACEAE (Cacao Family) 

Ayenia filiformis S. Wats. 


[Brewster Co.: Onion Spring, Paint Gap Hills and Head of Boquillas Canyon (Powell, 

1998)] 

Ayenia microphylla Gray Dense Ayenia 


[Brewster Co.: “many localities in Big Bend Park” (Powell, 1998); Hot Springs 

(Sperry, 1938)] 

Ayenia pilosa Cristobal Dwarf Ayenia 


[Brewster Co.: Chisos Mts. and upper Big Brushy Canyon in Dead Horse Mts. 

(Powell, 1998)] 

Note: Sperry (1938) reports Ayenia pusilla L. from near the Window and near top of 

Baldy Mtn. (nos. 685 and 901). 

TAMARICACEAE (Tamarisk Family) 

Tamarix aphylla (L.) Karsten. Athel 


[Brewster Co.: Boquillas Canyon (Powell, 1998)] 

Tamarix chinensis Lour. Salt Cedar; Tamarisk 


Tamarix gallica L. Salt Cedar; Tamarisk 


[Brewster Co.: Rio Grande Village (Powell, 1998)] 

Tamarix ramosissima Ledeb. Salt Cedar; Tamarisk 


Note: This group is especially problematic. D. H. Wilken (1993) indicates that T. gallica 

and T. ramosissima may be conspecific. 

ULMACEAE (Elm Family) 

Celtis laevigata Willd. var. reticulata (Torr.) L. Benson Netleaf Hackberry 


[Brewster Co.: Oak Creek Canyon] 

Note: Sperry (1938) reports var. texana from Oak Creek Canyon, but Powell (1998) 

does not treat this taxon as part of the Trans-Pecos flora. 

Celtis pallida Torr. Spiny Hackberry; Granjeno 


[Brewster Co.: “Neville Spring, lower Juniper Canyon, creek bed north of Panther 

Mountain, Gano Spring (Sperry, 1938)] 

URTICACEAE (Nettle Family) 

Parietaria pensylvanica Muhl. ex Willd. Pellitory 



Urtica gracilenta Greene 

[Woodland, Bassett & Crompton, 1976, mapped in or close to BBNP] 

VERBENACEAE (Vervain Family)

Aloysia gratissima (Gillies & Hook.) Troncoso Whitebrush; Common Beebrush 

[Lippia ligustrina (Lag.) Britt.; Aloysia lycioides Cham.] 



Aloysia wrightii Heller ex Abrams Oreganillo 

[Lippia wrightii Gray] 


[Brewster Co.: Neville Spring area and lower Blue Creek (Sperry, 1938)] 

Bouchea longiflora (Mold.) Nesom Spoon-leaf 

[Bouchea spathulata Torr. var. longiflora Mold.] 

[Steyermark & Moore, 1933; Wells, 1965; Nesom, 1991, mapped and taxonomy; Powell, 

1998] 

[Brewster Co.: Dead Horse Mts. (Wells, 1965); head of Heath Canyon in Dead 

Horse Mts. , Old Boquillas Road opposite Alto Relex and E of Tornillo Creek 

(Powell, 1998); rocky ridge above Boquillas Canyon, Moore & Steyermark 3446 (MO?)] 

Glandularia bipinnatifida (Nutt.) Nutt. var. bipinnatifida 

[Verbena ciliata Benth.] 

[Sperry, 1938; Lewis & Oliver, 1961; Umber, 1979, mapped; Turner, 1998] 

[Brewster Co.: Tornillo Flats, Dog Creek Wash and Santa Elena Canyon (Sperry, 1938); 

BBNP, Lewis & Oliver 5454 (ASTC as G. wrightii)] 

Note: Turner (1998) calls this plant Glandularia bipinnatifida (Nutt.) Nutt. var. ciliata 

(Benth.) B. L. Turner and treats G. wrightii as a synonym of it. I do not know which 

taxonomic interpretation will prevail. 

Glandularia verecunda Umber 

[Umber, 1979, mapped; Turner, 1998] 

Note: Turner (1998) treats this species as a synonym Glandularia quadrangulata (Heller) Umber. 

It is not clear from Turner’s map if he saw material from within BBNP. 

Glandularia wrightii (Gray) Umber 

[Sperry, 1938; Lewis & Oliver, 1961; Umber, 1979] 

[Brewster Co.: BBNP, Lewis & Oliver 5454 (MO) (Lewis & Oliver, 1961)] 

Lantana achyranthifolia Desf. Veinyleaf Lantana; Desert Lantana 

[Lantana macropoda Torr.] 


[Brewster Co.: Chisos Mts. and vicinity (Powell, 1998); Oak Creek Canyon, 

Persimmon Gap and Glenn Spring (Sperry, 1938); Oak Canyon, 17 Jul 1932, Muller 

32016 (UTEP); 1.3 mi. N pavement on Old Ore Rd. to Ernst Tinaja, BBNP, 21 Aug 1982, 

Van Devender s.n. (UTEP)] 

Lantana urticoides Hayek. Common Lantana; Texas Lantana 


Lippia graveolens Kunth Scented Lippia; Redbrush Lippia 


[Brewster Co.: Boquillas area (Sperry, 1938); 4 mi. W of Hot Springs and Dead 

Horse Mts. (Powell, 1998); Old Ore Rd. 1 mi. N Tx rd. to Rio Grande Village at Cuesta 

Carlota area, 2400-2800 ft., 27 May 1982, Van Devender s.n. (UTEP)] 

Phyla nodiflora (L.) Greene Sawtooth Frogfruit 

[Lippia incisa (Small) Tidest.] 


[Brewster Co.: Boquillas (Sperry, 1938); mouth of Panther Canyon, BBNP, 04 Apr 1976, 

Everitt s.n. (UTEP)] 

Tetraclea coulteri Gray Stinkweed 


[Brewster Co.: Smoky Creek (Sperry, 1938)] 

Verbena canescens Kunth 

[Verbena canescens Kunth var. roemeriana (Scheele) Perry] 

[Lewis & Oliver, 1961] 

[Brewster Co.: BBNP, Lewis & Oliver 5456 (ASTC)]

Verbena halei Small 


[Brewster Co.: Rio Grande near Mariscal Canyon (Sperry, 1938)] 

Verbena neomexicana (Gray) Small 

[Sperry, 1938, as var. hirtella Perry and unknown vr.] 

[Brewster Co.: Basin and flats N of Chisos Mts. (Sperry, 1938); 1.3 mi. N of pavement 

on Old Ore Rd. to Ernst Tinaja, 21 Aug 1982, Van Devender s.n. (UTEP); BBNP, Lewis 

& Oliver 5456 (ASTC as var. hirtella)] 

Verbena plicata Greene 


[Brewster Co.: Santa Elena Canyon, Moore & Steyermark 3299] 

VISCACEAE (Mistletoe Family) 

Phoradendron hawksworthii (Wiens) Wiens Rough Mistletoe 

[Phoradendron bolleanum (Seem.) Eichler ssp. hawksworthii Wiens] 

[Sperry, 1938; Lewis, Stripling & Ross, 1962; Powell, 1998] 

[Records: “Basin, Green Gulch, Laguna” (Sperry, 1938); Basin Rd. 1.2 mi. 

above St. Hwy. 118, BBNP, 4200 ft., 16 Mar 1985, Van Devender (UTEP); 

BBNP, Lewis & Oliver 5452 (BRIT)] 

Phoradendron juniperinum Engelm. Juniper Mistletoe 


[Brewster Co.: Chisos Mts. on Juniperus and Cupressus (Powell, 1998)] 

Phoradendron tomentosum (DC.) Engelm. ex A. Gray Christmans Mistletoe 


Note: Collections by Sperry (1938) said to be Phoradendron cockerellii on 

Acacia and Prosopis at Hot Springs and Gano Springs are likely this taxon. 

Phoradendron villosum (Nutt.) Nutt. ssp. coryae (Trel.) Wiens Oak Mistletoe 

[Phoradendron coryae Trel.; P. havardianum Trel.] 


[Brewster Co.: Chisos Mts. and Dead Horse Mts. (Powell, 1998); Basin, Panther Creek 

Canyon and Pulliam Bluff on oak and hackberry (Sperry, 1938); Chisos Mts., Moore & 


VITACEAE (Grape Family) 

Cissus incisa Des Moul. Ivy Treebine 

[Sperry, 1938, as Cissus macrocarpa Rose] 

[Brewster Co.: Basin, Fresno Spring Crees and tank at Nail’s ranch (Sperry, 1938)] 

Parthenocissus vitacea (Kneer.) Hitchc. Thicket Creeper 


[Brewster Co.: “lower canyons of the Rio Grande” (Powell, 1998)] 

Vitis arizonica Engelm. Canyon Grape 


Note: This is probably the only grape in BBNP (Powell, 1998). 

ZYGOPHYLLACEAE (Caltrop Family) 

Guaiacum angustifolia Engelm. Guayacan; Soapbush 


[Brewster Co.: Gano Spring, McKinney Spring and Santa Elena Canyon (Sperry, 

1938)] 

Kallstroemia grandiflora Torr. ex Gray Orange Kaltrop 


[Brewster Co.: Smoky Creek (Sperry, 1938)]

Kallstroemia hirsutissima Vail ex Small 


[Brewster Co.: near Santa Elena Canyon, Lone Mtn., and above Hot Springs 

(Sperry, 1938)] 

Larrea tridentata (Sesse & Moc. ex DC.) Cov. Creosotebush; Gobernadora 


[Brewster Co.: Dead Horse Mts. (Wells, 1965); Basin (Sperry, 1938)] 

Tribulus terrestris L. Goat Head; Puncture Weed 


[Brewster Co.: lower Green Gulch and Boquillas (Sperry, 1938)] 

AGAVACEAE (Agave Family) 

LILIOPSIDA (Monocots) 

Agave glomeruliflora (Engelm.) Berger Chisos Agave 

[Agave chisosensis C. H. Mull.] 

[Muller, 1939; Powell, 1998] 

[Brewster Co.: Chisos Mts., Basin area (Powell, 1998); Chisos Mts., Basin, 13 Jul 1937, 

Sperry, Warnock & McDougall 840 (US, type of A. chisosensis)] 

Note: This species may be a hybrid (Powell, 1998). 

Agave havardiana Trel. Havard Agave 

[Mulford, 1896, as A. applanata Koch ex Jacobi;Trelease, 1911; Sperry, 1938, as Agave scabra 

Salm-Dyck; Powell, 1998] 

[Brewster Co.: Chisos Mts. (Powell, 1998); Basin (Sperry, 1938); Chisos Mts., Aug 1883, 

Havard (MO?)] 

Agave lechuguilla Torr. Lechuguilla 


[Brewster Co.: Dead Horse Mts. and Sue Peaks (Wells, 1965)] 

Dasylirion heteracanthum I. M. Johnston Sotol 

[Powell, 1998} 

[Brewster Co.: southern Brewster Co. (Powell, 1998); just N of Willow Spring, 

BBNP, 29 DEG 18.560’N, 103 DEG 01.119’W, 990 m, 05 Apr 1995, Hodgson 

8984 (UTEP, DES)] 

Dasylirion leiophyllum Engelm. ex Trel. Smooth Sotol; Desert Candle 

[Sperry, 1938; Wells, 1965] 

[Brewster Co.: Dead Horse Mts. and on Sue Peak (Wells, 1965); Basin and 


Nolina erumpens (Torr.) S. Wats. Foothill Nolina; Beargrass 

[Trelease, 1911; Sperry, 1938] 

[Brewster Co.: Chisos Mts., 1901, Bailey 391; slopes surrounding Chisos Mts. 

(Sperry, 1938)] 

Nolina texana S. Wats. var. texana Texas Sacahuista 

Yucca baccata Torr. Datil Yucca; Banana Yucca 


[Brewster Co.: flats and hills N of Chisos Mts. (Sperry, 1938)] 

Yucca elata (Engelm.) Engelm. Soaptree Yucca; Palmella 


[Brewster Co.: W od Dead Horse Mts. (Sperry, 1938)] 

Yucca faxoniana (Trel.) Sarg. Faxon Yucca 

[Yucca carnerosana (Trel.) McKelvey] 

[Wells, 1965; Robins, 1983; Powell, 1998] 

[Brewster Co.: Dagger Flat and northern Dead Horse Mts. (Powell, 1998); Dead 

Horse range and Sue Peak (Wells, 1965); Dagger Flat, 8.7 mi. from jct. hwy. to 

Marathon, BBNP, 29 DEG 31.45’N, 103 DEG 02.99’W, 1050 m, 07 Apr 1995,

Hodgson 9024 (UTEP, DES); south of Persimmon Gap on road from Marathon to 

Boquillas (TEX-LL; A)] 

Yucca thompsoniana Trel. Beaked Yucca 

[Yucca rostrata Engelm. ex Trel.] 

[Sperry, 1938; Wells, 1965] 

[Brewster Co.: Dead Horse Mts. (Wells, 1965); Persimmon Gap (Sperry, 1938); Dagger 

Flat, 8.7 mi. from jct. with hwy. to Marathon, BBNP, 29 DEG 31.45’N, 103 DEG 02.99’ 

W, 1050 m, 07 Apr 1995, Hodgson 9025 (UTEP, DES)] 

Yucca torreyi Shafer Torrey Yucca 

[Wells, 1965; Robins, 1983] 

[Brewster Co.: common on alluvium in Dead Horse Mts. (Wells, 1965); 5.8 rd. 

mi. W Panther Jct.-Boquillas Rd. on Old Oar Rd., BBNP, 750 m, 05 Apr 1995, 

Hodgson 8990 (UTEP, DES); 2.9 mi. N of Lajitas Gate, BBNP, 29 DEG 18.318’N, 

103 DEG 45.316’W, 810 m, Hodgson 9057 (UTEP, DES); north entrance to BBNP 

(TEX-LL); lower Green Gulch (TEX-LL)] 

Note: Robins (1983) considers this species conspecific with Yucca treculiana. 

BROMELIACEAE (Pineapple Family) 

Hechtia texensis S. Wats. Texas Falseagave 

[Hechtia scariosa L. B. Sm.] 

[Sperry, 1938; Wells, 1965; Burt-Utley & Utley, 1987; Powell, 1998] 

[Brewster Co.: near Frog Tank and McKinney Springs in Dead Horse Mts., 

above Boquillas and above Glenn Springs (Powell, 1998); near Glenn Springs 

(Sperry, 1938); rocky hills near Hot Springs, BBNP, Correll & Rollins 23619 

(TEX-LL); near McKinney Springs, BBNP, Warnock 14047, 21610 (TEX-LL)] 

Tillandsia recurvata (L.) L. 

[Steyermark & Moore, 1933; Sperry, 1938; White et al., 1988, mapped] 

[Brewster Co.: S slope of Pulliam Bluff and upper Juniper Spring (Sperry, 1938); 

Chisos Mts., on vertical cliff face below Emory Peak, Moore & Steyermark 3198 

(MO?)] 

COMMELINACEAE (Spiderwort Family) 

Commelina dianthifolia Delile var. dianthifolia Birdbill Dayflower 

[Sperry, 1938; Brashier, 1966] 

[Brewster Co.: Chisos Mts. (Sperry, 1938); Chisos Mts., 1931, Mueller 7956 (UC, 

MICH, MO)] 

Commelina erecta L. var. angustifolia (Michx.) Fern. Erect Dayflower 

[Commelina crispa Woot.] 


[Records: S slope of Mt. Emory and flats N of Chisos Mts. (Sperry, 1938)] 

Tradescantia brevifolia (Torr.) Rose 

[Tradescantia leiandra Torr.; Setcreasea leiandra (Torr.) Pilger var. brevifolia Torr.; 

Setcreasea ovata (Coult.) Faruqi] 

[Steyermark & Moore, 1933; Sperry, 1938; Faruqi, Celarier & Mehra, 1962] 

[Records: Laguna and E slope of Casa Grande (Sperry, 1938); Chisos Mts., 

Moore & Steyermark 3228 (GH; MO; F), Hinckley (GH; F), Havard 78 (GH; US), 

Warnock T574C (GH; US; F), Warnock 643 (US), Sperry 1476 (GH), Mueller 7854 

(F, US, MO, MICH), Young 122 (MO), McDougall 33 (GH)] 

CYPERACEAE (Sedge Family) 

Bulbostylis capillaris (L.) Kunth ex C. B. Clarke 

[Sperry, 1938; Kral, 1971, mapped in Brewster Co.]

[Brewster Co.: top of Flattop (Sperry, 1938); Chisos Mts., Mueller s.n. (UTEP)] 

Bulbostylis juncoides (Vahl) Alain 

[Kral, 1971, mapped in Brewster Co.] 

Carex geophila Mackenzie 

[Jpnes & Jones, 1993] 

[Brewster Co.: Pinnacle’s trail towards Boot Springs (Jones & Jones, 1993)] 

Carex lativena S. D. Jones and G. D. Jones Broad-veined Carex 


[Brewster Co.: BBNP, 0.8 km southward on the Pinnacle’s trail towards Boot Springs 

from the trail head near the Post Office, ca. 1735 m, 19 Apr 1991, S. & G. Jones 6523 

(MICH-Holotype, BRIT, TAES, TEX-LL, US, VDB); Pine Canyon, BBNP, 1735 m, 

21 Apr 1991, S. & G. Jones 6547 (SRSC)] 

Carex planostachys Kunze 

[Steyermark & Moore, 1933; Correll & Johnston, 1970] 


Cyperus elegans L. 


[Brewster Co.: W of Elephant Tusk Mtn. and Pinnacle Mts. (Sperry, 1938)] 

Cyperus fendlerianus Boeckl. 

[Brewster Co.: Chisos Mts., 28 Jul 1932, Mueller 8793 (UTEP)] 

Cyperus laevigatus L. 


[Brewster Co.: Glenn Spring and Tornillo Creek at Hot Spring (Sperry, 1938)] 

Cyperus odoratus L. Fragrant Flatsedge 


[Brewster Co.: Willow Creek Canyon, lower Oak Creek and Boquillas (Sperry, 1938)] 

Cyperus retroflexus Buckl. 

[Cyperus uniflorus Torr. & Hook. non Thunb.] 



Cyperus seslerioides Kunth 


[Brewster Co.: ridge above Green Gulch (Sperry, 1938); Chisos Mts., 28 Jul 1932, 

Mueller s.n. (UTEP)] 

Cyperys sphaerolepis Boeckl. 

[Cyperus rusbyi Britt.] 



Cyperus squarrosus L. Bearded Flatsedge 

[Cyperus aristatus Rottb.] 


[Brewster Co.: Boquillas Canyon (Sperry, 1938)] 

Eleocharis geniculata (L.) Roemer & Schult. 

[Eleocharis caribaea (Rottb.) Blake] 


[Brewster Co.: Pinnacle Mts. (Sperry, 1938); Boquillas, Moore & Steyermark 3453] 

Fimbristylis annua (All.) Roemer & J. A. Schultes 

[Kral, 1971, mapped in BBNP] 

Fuirena squarrosa Michx. Umbrella-grass 

[Fuirena hispida Ell.] 


[Brewster Co.: Pinnacle Mts. (Sperry, 1938)] 

JUNCACEAE (Rush Family) 

Juncus interior Wieg. var. arizonicus (Wieg.) F. J. Herm.

[Juncus arizonicus Wieg.] 


[Brewster Co.: upper Pulliam Canyon (Sperry, 1938)] 

Juncus nodosus L. 


[Brewster Co.: lower Oak Creek, Gano Spring, Glenn Spring and Pinnacle Mts. 

(Sperry, 1938); Chisos Mts., Moore & Steyermark 3390] 

Juncus saximontanus A. Nels. 

[Juncus brunnescens Rydb.] 


[Brewster Co.: Cattail Falls (Sperry, 1938)] 

LILIACEAE (Lily Family) 

Allium cernuum Roth. Nodding Onion 

[Sperry, 1938; Ownbey, 1950] 

[Brewster Co.: Boot Spring (Sperry, 1938); Mt. Emory, 10 Sep 1933, Cory 7279 (GH); 

Boot Creek, 26 Sep 1938, Cory 30369 (GH); upper Boot Canyon, 6 Aug 1931, Mueller 

7963 (GH, MO, NY, BRIT, US, TEX-LL); Boot Spring, 25 Aug 1937, Warnock 1094 

(GH, US, TEX-LL)] 

Allinm kunthii Don Kunth Onion 

[Sperry, 1938; Ownbey, 1950] 

[Brewster Co.: S slope of Mt. Emory and Bois D’Arc Spring (Sperry, 1938); Emory 

Peak, 14 Aug 1931, Mueller 7962 (GH, MO, NY, US, TEX-LL); Chisos Mts., 2 Aug 1932, 

Mueller 32012 (MO, NY, TEX-LL); near Muskhog Spring, 30 Jul 1938, Sperry 1466 (GH, 

US)] 

Cooperia drummondii Herbert 


[Brewster Co.: Laguna and N slope of Pulliam Bluff (Sperry, 1938)] 

Nothoscordum bivalve (L.) Britt. Crow Poison 

[Sperry, 1938; McVaugh, 1952] 

[Records: Bois D’Arc Spring (Sperry, 1938)] 

ORCHIDACEAE (Orchid Family) 

Epipactis gigantea Dougl. ex Hook. Giant Helleborine 


[Brewster Co.: Chisos Mts. (Sperry, 1938); Chisos Mts., head of Cat Tail Canyon, 

Moore & Steyermark 3406 (MO?)] 

Hexalectris revoluta Correll 


[Brewster Co.: rare in lower Willo Creek basin of Chisos Mts., 16 Jul 1937, Warnock 

21551] 

Hexalectris warnockii Ames & Corell 

[Steyermark & Moore, 1933, as Hexalectris spicata (Walt.) Barnh.; Correll, 1947, 1961] 

[Brewster Co.: Chisos Mts., Love Creek Basin, 20 Jul 1932, Mueller; Chisos Mts., near 

Lost Mine Peak, 6500 ft., 18 Jun 1937, Warnock 20774; Chisos Mts., upper Blue Creek 

Canyon, 25 Jun 1937, Warnock 21261; Chisos Mts., trail to Baldy Peak, Warnock 20776; 

Chisos Mts., Basin at Window, 04 Aug 1946, Lundell & Lundell 14609; Chisos Mts., 

Near Window of Basin, 04 Aug 1946, Correll 13630; Chisos Mts., Moore & Steyermark 

3391] 

Malaxis wendtii Salazar 

[Correll, 1968; Todsen, 1995] 

[Brewster Co.: Chisos Mts., Warnock 23382 (SRSC); Chisos Mts., Boot Canyon, 12 Aug 

1967, Whitson 302 (SRSC); Boot Canyon, Aug 1964, Whitson (BAYLU)]

Spiranthes cinabarina (Llave & Lex.) Hemsl. Scarlet Ladies’ Tresses 

[Sperry, 1938; Correll, 1961; Correll & Johnston, 1970] 

[Brewster Co.: trail above upper Cattail Falls (Sperry, 1938)] 

Spiranthes confusa (Garay) Kartesz & Gandhi 

[Spiranthes durangensis Ames & C. Schweinf.; Spiranthes saltensis Ames] 

[Ames, 1932; Steyermark & Moore, 1933; Correll, 1961, 1978] 

[Records: Chisos Mts., N slope Emory Peak, grassy soil pockets in rim rock, 22 Jun 1931, 

Moore & Steyermark 3214 (A?)] 

POACEAE (Grass Family) 

Achnatherum eminens (Cav.) Barkworth Southwestern Needlegrass 

[Stipa eminens Cav.] 



Achnatherum lobata (Swallen) Barkworth Needlegrass 

[Stipa lobata Swallen] 



Agrostis gigantea Roth Redtop 

[Agrostis alba L. of authors] 


[Brewster Co.: along creek from Boot Spring toward the Boot (Sperry, 1938)] 

Agrostis scabra Willd. var. scabra Rough Bentgrass 

[Agrostis hyemalis (Walt.) B.S.P.] 


[Brewster Co.: wooded canyons of Chisos Mts. (Sperry, 1938)] 

Andropogon gerardii Vitman Big Bluestem 

[Andropogon furcatus Muhl. ex Willd.] 



Aristida adscensionis L. Sixweeks Threeawn 


[Brewster Co.: Green Gulch, Pinnacle Mts. and Fresno Spring Creek (Sperry, 1938)] 

Aristida arizonica Vasey Arizona Threeawn 


[Brewster Co.: South Rim (Sperry, 1938)] 

Aristida divaricata Willd. Poverty Threeawn 


[Brewster Co.: S slopes of Chisos Mts. (Sperry, 1938)] 

Aristida orcuttiana Vasey Single Threeawn 


[Brewster Co.: Basin and lower Juniper Spring (Sperry, 1938)] 

Aristida purpurea Nutt. Purple Threeawn 

var. nealleyi (Vasey) Allred Nealley Threeawn 

[Aristida glauca (Nees) Walp.] 


[Brewster Co.: lower willow creek and N of Chisos Mts. (Sperry, 1938)] 

var. purpurea 

[Aristida purpurea micrantha; A. roemeriana Scheele] 

[Hitchcock, 1924] 

[Brewster Co.: Boquillas, Bailey 341] 

var. wrightii (Nash) Allred Wright’s Threeawn 


[Brewster Co.: Basin along trail to Laguna (Sperry, 1938)] 

Aristida ternipes Cav. Hook Threeawn



Note: The varieties remain to be sorted out. 

Blepharoneuron tricholepis (Torr.) Nash Pine Dropseed 

[Sperry, 1938; Peterson & Annable, 1990] 

[Brewster Co.: upper slopes of Basin and Pine Canyon (Sperry, 1938); Lost 

Mine Peak, BBNP, 12 Sep 1961, Correll & Johnston 24533 (NY)] 

Bothriochloa barbinodis (Lag.) Herter Cane Bluestem 


[Records: Boquillas Canyon (Sperry, 1938)] 

Bothriochloa laguroides (DC.) Herter ssp. torreyana (Steud.) Allred & Silver Bluestem 

Gould 

[Sperry, 1938, as Bothriochloa saccharoides (Swartz) Rydb.] 


Bothriochloa springfieldii (Gould) Parodi 

[Andropogon springfieldii Gould] 

[Records: Chisos Mts., Lundell 13286 (UC)] 

Bouteloua aristidoides (H.B.K.) Griseb. Needle Grama 


[Records: lower Green Gulch and Pinnacle Mts. (Sperry, 1938)] 

Bouteloua barbata Lag. Sixweeks Grama 


[Records: Oak Creek Canyon, E slope of Rosillos Mts. and Tornillo 

Flats (Sperry, 1938)] 

Bouteloua curtipendula (Michx.) Torr. 

[Sperry, 1938; Wells, 1965] 

[Records: Dead Horse Mts. (Wells, 1965); lower Juniper Canyon and Basin 

(Sperry, 1938)] 

var. curtipendula 

[Kapadia & Gould, 1964, mapped] 

var. caespitosa Gould & Kapadia 

[Kapadia & Gould, 1964, mapped] 

Bouteloua eriopoda Torr. Black Grama 



Bouteloua gracilis (H.B.K.) Lag. ex Steud. Blue Grams 

[Griffiths, 1912; Sperry, 1938] 

[Brewster Co.: Laguna and Mariscal Canyon (Sperry, 1938); Chisos Mts., Bailey 393] 

Bouteloua hirsuta Lag. Hairy Grama 


[Brewster Co.: Basin and Pine Canyon (Sperry, 1938)] 

Bouteloua ramosa Scribn. ex Vasey 

[Sperry, 1938, as Bouteloua breviseta; Wells, 1965; Reeder & Reeder, 1980] 

[Brewster Co.: Dead Horse Mts. (Wells, 1965); Basin, 08 Sep 1936, Sperry 395 (SRSC); 

Ward Spring on W side of Chisos Mts., 18 Aug 1955, Warnock 13110 (SRSC); Chisos 

Mts., 16 Sep 1936, Silveus 3405 (ARIZ, TEX-LL), 01 Aug 1931, Mueller 7880 (TEX-LL, 

US); Smoky Creek, 21 May 1955, Warnock & Wallmo 13219 (TEX-LL, SRSC); 10 mi. 

S of Persimmon Gap entrance, 03 Aug 1972, Reeder & Reeder 5827 (RM)] 

Bouteloua uniflora Vasey 


[Brewster Co.: Basin and Oak Creek Canyon (Sperry, 1938)] 

Brachiaria arizonica (Scribn. & Merr.) S. T. Blake Arizona Signalgrass 

[Panicum arizonicum Scribn. & Merr.] 


[Brewster Co.: Fresno Springs (Sperry, 1938)] 

Bromus anomalus Rupr. ex Fourn.

[Sperry, 1938; Wagon, 1952, mapped] 

[Records: upper slopes of Basin and Laguna (Sperry, 1938)] 

Bromus lanatipes (Shear) Rydb. Woollysheath Brome 

[Wagon, 1952, mapped; Powell, 1994] 

[Records: trail to Laguna, Lost Mine Peak (Powell, 1994)] 

Bromus marginatus Nees ex Steud. 


[Records: wooded canyons of the Chisos Mts. (Sperry, 1938)] 

Cathestechum erectum J. Presl. 


[Brewster Co.: Near Johnson’s Ranch and Avary Canyon (Sperry, 1938); Santa 


Cenchrus incertus M. A. Curtis Sandbur; Grassbur 

[De Lisle, 1964] 

[Brewster Co.: mouth of Santa Elena Canyon, De Lisle 837 (TEX-LL)] 

Cenchrus myosuroides Kunth 


[Brewster Co.: Fresno Creek (Sperry, 1938)] 

Cynodon dactylon (L.) Pers. Bermudagrass 


[Brewster Co.: Fresno Springs and along Rio Grande (Sperry, 1938)] 

Dasyochloa pulchella (Kunth.) Steud. Fluffgrass 

[Sperry, 1938; Wells, 1965] 

[Brewster Co.: Dead Horse Mts. (Wells, 1965); Oak Creek Canyon, Pinnacle Mts. 

and N of Chisos Mts. (Sperry, 1938)] 

Dichanthelium acuminatum (Sw.) Gould var. acuminatum Wooly Rosettegrass 

[Panicum lindheimeri Nash] 


[Brewster Co.: near Cattail Falls (Sperry, 1938)] 

Digitaria cognata (Schult.) Pilger ssp. pubiflora Wipff & Hatch Fall Witchgrass 



Digitaria sanguinalis (L.) Scop. Hairy Crabgrass 


[Brewster Co.: lower Green Gulch and Boquillas (Sperry, 1938)] 

Echinochloa colonum (L.) Link Junglerice 


[Brewster Co.: Pine Canyon and Fresno Creek (Sperry, 1938)] 

Elymus arizonicus (Scribn. & J. G. Sm.) Gould Arizona Wheatgrass 

[Agropyron arizonicum Scribn. & Smith] 


[Brewster Co.: S slope of Mt. Emory and between Boot Spring and South 

Rim (Sperry, 1938)] 

Elymus elymoides (Raf.) Swezey Longleaf Squirreltail 


[Brewster Co.: Pinnacle’s trail towards Boot Springs (Jones & Jones, 1993); Green 

Gulch, Cattail Canyon et al. (Powell, 1994); Laguna (Sperry, 1938)] 

Note: The correct name for this taxon may be Elymus longifolius (Smith) Gould 

(Powell, 1994). 

Elymus smithii (Rydb.) Gould Western Wheatgrass 

[Agropyron smithii Rydb.] 



Elymus spicatus (Pursh) Gould [the inermus phase] Bluebunch Wheatgrass 

[Agropyron inerme (Scribn. & Sm.) Rydb.] 

[Sperry, 1938]


Enneapogon desvauxii Beauv. Feather Pappusgrass 

[Pappophorum wrightii S. Wats.] 


[Brewster Co.: Green Gulch and Fresno Spring (Sperry, 1938)] 

Eragrostis cilianensis (All.) Lut. ex Janchen Stinkgrass 


[Brewster Co.: Basin and along Fresno Spring Creek (Sperry, 1938); Oak Canyon, 

17 Jul 1932, Mueller s.n. (UTEP)] 

Eragrostis intermedia A. S. Hitchc. Plains Lovegrass 



Eragrostis pectinacea (Michx.) Nees ex Steud. var. pectinacea Carolina Lovegrass 

[Eragrostis diffusa Buckl.] 


[Brewster Co.: Laguna, upper Cattail Falls, lower Juniper Spring, NW slope 

of Pulliam Bluff and Boquillas Canyon (Sperry, 1938)] 

Erioneuron avenaceum (H.B.K.) Tateoka Large-flowered Tridens 

[Tridens grandiflorus (Vasey) Woot. & Standl.] 


[Brewster Co.: Chisos Mts. (Sperry, 1938); Chisos Mts., Moore & Steyermark 3314, 3367] 

Hemarthria altissima (Poir.) Stapf & C. E Hubbard 

[Manisuris altissima (Poir.) A. S. Hitchc.] 


[Brewster Co.: Rio Grande near Solis’ Ranch (Sperry, 1938)] 

Hesperostipa neomexicana (Thurb.) Barkworth New Mexico Feathergrass 

[Stipa neomexicana (Thurb.) Scribn.] 



Heteropogon contortus (L.) Beauv. ex Roem. & Schult. Tanglehead 


[Brewster Co.: Juniper Canyon, Willow Creek Canyon, Santa Elena Canyon and 

Divide (Sperry, 1938)] 

Koeleria macrantha (Ledeb.) Schult. Junegrass 

[Koeleria cristata (L.) Pers. of authors] 


[Brewster Co.: Basin, Pine Canyon and Laguna (Sperry, 1938); Chisos Mts., Moore 

& Steyermark] 

Leptochloa dubia (H.B.K.) Nees Green Sprangletop 


[Brewster Co.: Green Gulch, slopes of Casa Grande, Oak Creek. Mariscal 

Canyon and upper Cattail Falls (Sperry, 1938)] 

Leptochloa fascicularis (Lam.) Gray Bearded Sprangletop 

[Manisuris fasciculata (Lam.) Hitchc.] 

[Steyermark & Moore, 1933; Powell, 1994] 

[Brewster Co.: Santa Elena Canyon, Moore & Steyermark 3434; Upper Cattail 

Falls and Boot Springs (Powell, 1994)] 

Leptochloa uninervia (Presl.) Hitchc. & Chase Mexican Sprangletop 


[Brewster Co.: bank of Rio Grande near Ogle Spring (Sperry, 1938)] 

?Lycurus phleoides H.B.K. Wokftain 


[Brewster Co.: top of Divide and Green Gulch (1938)] 

Note: At the time Sperry made this collection the Lycurus setosus was unknown. 

Lycurus setosus (Nutt.) C. Reeder 

[Reeder, 1985]

[Brewster Co.: Chisos Mts., Warnock 814 (ARIZ), Shinners 8771 (ARIZ)] 

Muhlenbergia crispiseta A. S. Hitchc. 

[Peterson & Annable, 1991] 

[Brewster Co.: BBNP, Chisos Mts., Cattail Canyon, Warnock 21268 (SRSC)] 

Muhlenbergia depauperata Scribn. 

[Peterson & Annable, 1991] 

[Brewster Co.: BBNP, Lost Mine Peak, Correll & Johnston 24492 (MO, NY); 

Green Gulch, BBNP, Warnock 20027 (MO, NY, RSA, US)] 

Muhlenbergia emersleyi Vasey Bullgrass; Bull Muhly 


[Brewster Co.: Basin, upper Cattail Canyon, near top of Casa Grande and Green Gulch 

(Sperry, 1938)] 

Muhlenbergia glauca (Nees) B. D. Jackson Desert Muhly 

[Muhlenbergia lemmonii Scribn.] 

[Lamson-Scribner, 1911; Sperry, 1938] 

[Brewster Co.: Basin and Laguna (Sperry, 1938); Chico sic Mts., 1889, Nealley 379 (US)] 

Muhlenbergia pauciflora Buckl. New Mexico Muhly 



Muhlenbergia polycaulis Scribn. Cliff Muhly 


[Brewster Co.: Boot Spring (Sperry, 1938); Chisos Mts., Moore & Steyermark 3215] 

Muhlenbergia porteri Scribn. ex Beal Bush Muhly 


[Brewster Co.: near Lechuguilla Pens and Kit Mtn. (Sperry, 1938)] 

Muhlenbergia repens (Presl.) Hitchc. Creeping Muhly; Mat Muhly 



Muhlenbergia rigida (H.B.K.) Kunth Purple Muhly 


[Brewster Co.: Laguna and Basin (Sperry, 1938)] 

Muhlenbergia tenuifolia (Kunth) Trin. Mesa Muhly 

[Muhlenbergia monticola Buckl.] 


[Brewster Co.: Willow Creek Canyon and near top of Casa Grande (Sperry, 1938)] 

Nassella tenuissima (Trin.) Barkworth Finestem Needlegrass 

[Stipa tenuissima Trin.] 

[Hitchcock, 1925; Sperry, 1938; Jones & Jones, 1993; Powell, 1994] 

[Brewster Co.: Pinnacle’s trail towards Boot Springs (Jones & Jones, 1993); Chisos 

Mts. et al. (Powell, 1994); Laguna and lower Green Gulch (Sperry, 1938); Chisos Mts., 

Havard 12] 

Panicum bulbosum H.B.K. Bulb Panicum 


[Brewster Co.: Willow Creek, Basin and Oak Creek Canyon (Sperry, 1938); Chisos 

Mts., Moore & Steyermark] 

Panicum hallii Vasey Filly Panicum 


[Brewster Co.: near Burnham’s Ranch and Kit Mountain (Sperry, 1938)] 

Panicum hirticaule Presl. Mexican Witchgrass 



Panicum obtusum H.B.K. Vine Mesquite 



Pappophorum vaginatum Buckl. Whiplash Pappusgrass 

[Powell, 1994]

[Brewster Co.: Grapevine Hills, Oak Canyon and Basin Campground (Powell, 1994); 

Oak Canyon, 17 Jul 1932, Mueller s.n. (UTEP)] 

Paspalum distichum L. Knotgrass 

[Chase, 1929] 

[Brewster Co.: Boquillas, Swallen 1117] 

Paspalum pubiflorum Rupr. ex Fourn. 


[Brewster Co.: Fisk Canyon and Pinnacle Mts. (Sperry, 1938)] 

Pappophorum bicolor Fourn. 


[Brewster Co.: Muskhog Spring (Sperry, 1938)] 

Pappophorum vaginatum Buckl. 

[Pappophorum mucronulatum auct. non Nees] 


[Brewster Co.: Chisos Pens (Sperry, 1938)] 

Phragmites australis (Cav.) Trin. ex Steud. Common Reed 

[Phragmites communis Trin.] 


[Brewster Co.: Rio Grande, Solis’ Ranch and Boquillas (Sperry, 1938)] 

Pipitochaetium fimbriatum (Kunth in H.B.K.) A. S. Hitchc. Pinyon Ricegrass 

[Oryzopsis fimbriata (Kunth in H.B.K.) Hemsl.] 


[Brewster Co.: Willow Creek Canyon and near Saddle (Sperry, 1938); Chisos 

Mts., Moore & Steyermark] 

Pleuraphis mutica Buckl. Tobosa; Tobosagrass 

[Hilaria mutica (Buckl.) Benth.] 


[Records: Tornillo Flats, Dig Flats and Avary Canyon (Sperry, 1938)] 

Poa bigelovii Vasey & Scribn. Bigelow Bluegrass 

[Jones & Jones, 1993; Powell, 1994] 

[Records: Pinnacle’s trail towards Boot Springs (Jones & Jones, 1993); E slope 

Pulliam Peak, Dog Canyon, Dog Canyon flats, et al. (Powell, 1994)] 

Poa fendleriana (Steud.) Vasey Mutton Bluegrass 


[Records: Pinnacle’s trail toward Boot Springs (Jones & Jones, 1993)] 

Poa strictiramea Hitchc. Chisos Bluegrass 

[Poa involuta Hitchc.] 

[Hitchcock, 1928; Powell, 1994] 

[Records: Chisos Mts., first ridge SW of Juniper Canyon, 2200 m, 15-18 Jul 1921, Ferris 

& Duncan 2811 (Type of P. involuta: US); Chisos Mts., Swallen 1110 (US); trail to Boot 

Spring in Boot Canyon, along trail to South Rim, Emory Peak, Laguna Meadow, upper 

Pine Spring Canyon and upper Cattail Canyon (Powell, 1994)] 

Polypogon viridis (Gouan) Breistr. Water Polypogon 

[Agrostis verticillata Vill.] 


[Brewster Co.: lower Oak Creek Canyon and Fresno Springs (Sperry, 1938)] 

Schizachyrium scoparium (Michx.) Nash 


[Brewster Co.: Casa Grande and Baldy Mtn. (Sperry, 1938)] 

Scleropogon brevifolius Phil. Burrograss 

[Sperry, 1938; Wells, 1965] 

[Brewster Co.: Dead Horse Mts. (Wells, 1965); Dog Flats, Tornillo Flats and 

flats along E base of Rosillos Mts. (Sperry, 1938)] 

Setaria grisebachii Fourn. Grisebach’s Bristlegrass 


[Brewster Co.: Chisos Mts. (Sperry, 1938)]

Setaria leucopila (Scribn. & Merr.) K. Schum. Plains Bristlegrass 

[Setaria macrostachya H.B.K. of most authors] 

[Hitchcock, 1920; Sperry, 1938] 

[Brewster Co.: Fresno Creek, Willow Creek and Mariscal Canyon (Sperry, 1938); Boquillas, 

Bailey 392] 

Setaria verticillata (L.) Beauv. Hooked Bristlegrass 


[Brewster Co.: lower Oak Creek Canyon (Sperry, 1938)] 

Setaria viridis (L.) Beauv. Green Bristlegrass 


[Brewster Co.: Pinnacle Mts. (Sperry, 1938)] 

Sorghum bicolor (L.) Moench 

[Sorghum vulgare Pers. var. sudanense (Piper) Hitchc.] 


[Brewster Co.: Rio Grande near Mariscal Canyon (Sperry, 1938)] 

Sphenopholis obtusata (Michx.) Scribn. Prarie Wedgescale 



Sporobolus cryptandrus (Torr.) Gray Sand Dropseed 



Sporobolus pyramidatus (Lam.) A. S. Hitchc. Whorled Dropseed 


[Brewster Co.: Boquillas Canyon and Chisos Pens (Sperry, 1938)] 

Tragus berteronianus Schult. Spike Burgrass 


[Brewster Co.: lower Green Gulch and Oak Creek Canyon (Sperry, 1938)] 

Tridens albescens (Vasey) Woot. & Standl. White Tridens 


[Brewster Co.: Dog Flats (Sperry, 1938)] 

Tridens muticus (Torr.) Nash Slim Tridens 


[Brewster Co.: 1 mi. S end of Old Ore Rd. at Candelilla Camp, BBNP, 15 Mar 1983, 

Yatskievych (UTEP)] 

Tripsacum dactyloids (L.) L. 

[Sperry, 1938; Newell & DeWet, 1974, mapped] 

[Brewster Co.: Oak Creek Canyon near Window (Sperry, 1938)] 

POTAMOGETONACEAE (Pondweed Family) 

Potamogeton diversifolius Raf. 


[Brewster Co.: “waters of two unwooded canyons of Chisos Mountains (Sperry, 1938); 

creek near Boot Spring, Moore & Steyermark 3186 (MO?); head of Cat Tail Canyon, 

Moore & Steyermark 3414 (MO?)] 

Potamogeton foliosus Raf. var. foliosus 

[Haynes, 1974, mapped in or close to park] 

Potamogeton nodosus Poir. 

[Ogden, 1943] 

[Records: Haley Ranch, Cory 9198] 

Potamogeton pectinatus L. 

[Ogden, 1967] 

Note: Ogden (1967) reviewed four sheets from Brewster Co. but did not five the specific 

localities; the taxon is expected in the park.

TYPHACEAE (Cattail Family) 

Typha domingensis Pers. 

[Steyermark & Moore, 1933; Hotchkiss & Dozier, 1949, mapped] 

[Records: Boquillas, Moore & Steyermark 3455] 

Typha latifolia L. Broad-leaved Cattail 


[Records: Boquillas (Sperry, 1938)] 

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