A type catalogue of Campylopodioideae and ... - Tropical Bryology

Tropical Bryology 16: 17-102, 1999 

A type catalogue of Campylopodioideae and Paraleucobryoideae 

(Musci, Dicranaceae), Part II, Campylopus 

Jan-Peter Frahm 

Botanisches Institut der Universität, Meckenheimer Allee 170, 53115 Bonn, Germany 

Abstract. All species of Campylopus Brid. are listed, including citations of basionyms and homotypic 

synonyms, completed by citation of the type specimen and its location, as well as the current use or 

other uses of the name. 

As pointed out in the first part of this catalogue 

(Frahm 1994), type registers are important tools 

especially for herbarium curators, because it is 

often difficult even for an experienced curator to 

check all herbarium specimens for potential 

types, and it requires usually good library 

facilities to confirm the status of type specimens 

with the protologue. So far only a few herbarium 

based catalogues existed, and no genus based 

catalogues. Therefore the taxonomic data of 

Atractylocarpus, Brothera, Bryohumbertia, 

Campylopodiella, Campylopodium, Microcamplyopus, 

Paraleucobryum, Pilopogon and 

Sphaerothecium were listed in the first part of 

this catalogues. The genus Campylopus 

remained, because it needed special efforts to 

compile the data for more than 1000 species. 

The register includes information on: 

Species name, author, publication and year 

B: the basionym 

HS: eventually homotypic synonyms, 

17 

T: information on the type (country, locality, 

collector, number) as well as information on the 

location of the type using abbreviations of the 

Index herbariorum (Homgren et al. 1990) and 

the status of the type (holo-, lecto- iso-, syn-, 

paratypes) 

M: Material, in the case of nomina nuda. 

OU: other use of the name, the name of a species 

to which this species was attributed before, with 

reference. 

CU: if the species has been synonymized, the 

name of the species (current use) with which it 

has been synonymized, including reference. 

Nomina nuda are included but species described 

in Campylopus but belonging to other genera 

(such as Grimmia or Racomitrium) are not 

completely included. 

Accepted names are in boldface, synonyms are 

in italics, nomina nuda and doubtful species are 

in normal print.

18 

The information is not 100% complete, there are 

some cases that protologues (published in 

journals which were not available) could not be 

studied, and there are also species, of which 

material could not be studied. Such unrevised 

species are marked with an asterix (*). 

Frahm, J.-P. 1994. A type catalogue of 

Campyopodioideae and Paraleucobryoideae 

(Musci, Dicranaceae), Part I. Fragmenta 

Floristica et Geobotanica 39(1): 85-101, 1994. 

Holmgren, P.K., Homgren, N.H., Barnett, L.C. 

(eds.) 1990. Index herbariorum Part I: The 

herbaria of the world. 8. ed. Regnum Vegetabile 

120, 693 pp., New York. 

A 

abbreviatus (Dix.) Manuel, Fed. Mus. Journ. 

26:161 (1981) . 

B: Thysanomitrion abbreviatum Dix., Bull. 

Torr. Bot. Club 51: 227 (1924). 

T: Malaysia, Pahang, summit of Gunong 

Jahan, Ridley 1012 (isotype PC). 

CU: C. exasperatus (Nees & Blum) Brid. cf. 

Frahm, Nova Hedwigia 39:590 (1984). 

aberrans Broth. in Skottsberg, Nat. Hist. Juan 

Fernandez 2(3):414 (1923). 

T: Juan Fernandez Isl., Skottsberg 11 

(lectotypus, S; isotypus H-BR). 

CU: C. purpureocaulis Dus. ssp. aberrans 

(Broth.) J.-P.Frahm, Bryol. Beitr. 7:63 (1987). 

abyssinicus Schimp. in Par., Ind. Bryol. 262 

(1894). 

abyssinicus De Not., Nuov. Giorn. Bot. Ital. 4: 

21 (1872) (*) 

acervatus Mitt., J. Linn. Soc. Bot. 12:87 (1869). 

T: „America australis“, Lobb 122 (holotype 

NY, isotypes C, H-BR). 

CU: C. occultus Mitt. cf. Frahm, Bryol. Beitr. 

7:9 (1987). 

acrocaulon (C.Müll.) Kindb., Enum. Bryin. Exot. 

106 (1891). 

B: Dicranum acrocaulon C.Müll., Flora 

71:110 (1888). 

T: Kilimandscharo 300 - 4000 m, Meyer s.n. 

(lectotypus PC). 

CU: C. pilifer Brid. cf. Frahm, Bryoph. 

Bibl.31:143 (1985). 

acuminatus Mitt., J. Linn. Soc. Bot. 12:90 

(1869). 

T: Fuegia, Hermite Island, Hooker s.no. 

(holotype NY). 

acuminatus Mitt var. kirkii J.-P. Frahm, Bryol. 

Beitr. 7:9 (1987). 

B: Campylopus kirkii Mitt. in Beck., Trans. 

New Zeal. Inst. 76: 280 (1894). 

OU: C. acuminatus var. pilosus J.-P. Frahm, 

Lindbergia 7: 30 (1981). 

acutirameus Dix. & Thér. ex Bartr., Philipp. 

Journ. Sc. 68:37 (1939). 

T: Philippines: on palm roofs, Mayaygay, 

Baker 3659 (isotypes NY, S) 

CU: C. savannarum (C. Müll.) Mitt. cf. 

Frahm, Cryptogamie Bryol. Lich. 6:207 

(1985). 

adustus De Not., Atti Univ. Genova 1:469 (1869) 

CU: C. atrovirens De Not. ssp. adustus (De 

Not.) Kindb. 

aemulans (Hampe) Jaeg., Ber. S. Gall. Naturw. 

Ges. 1870-71:444. 1872. 

B: Thysanomitrium aemulans Hampe, Vid. 

Medd. Naturh. Foer. Koebenh. ser.3,2:273. 

1870. 

T: Brazil, Rio de Janeiro, Glaziou 3307 

(isotypes, H-BR, NY, PC, S). 

aethiops Welw. & Duby Mem. Soc. Phys. Hist. 

Nat. Geneve 21:221 (1870). 

T: (Angola) „Ad rupes humedas in summis 

jugis Pedra Songue 3800 ft.“, Welwitsch 64 

(holotype G, isotype S) . 

CU: C. aureonitens (C. Müll.) Jaeg. cf. 

Frahm, Bryoph. Bibl. 31:24 (1985). 

afroconcolor .Müll. ex Broth. Bot. Jahrb. 24:235 

(1897). 

T: Type: Kamerun, „in montis Rumpi ad Jumbo 

pagum c. 1100m in truncis putridis, Dusén 

653 (holotype B, isotypes BM, S). 


Frahm, Lindbergia 8:48 (1982). 

akagiensis Broth. & Yas., Oefv. Finsk. Vet. Soc. 

Foerh. 62A(9):3 (1921) . 

T: Japan, Mt. Akagi, Tannoda 300 (holotype

H-BR). 

CU: C. fragilis (Brid.) B.S.G. cf. Takaki, 

Journ. Hatt. Bot. Lab. 30:240 (1967). 

alatus (Broth. ex Ihs.) Iwats. & Nog., Journ. Jap. 

Bot. 48(7):215 (1973). 

B: Thysanomitrion alatum Broth. ex Ihs., Cat. 

Moss. Japan 33 (1929). 

CU: C. umbellatus (Arn.) Par. cf. Frahm, 

Nova Hedwigia 39:601 (1984). 

albescens (C. Müll.) Jaeg.Ber. S. Gall. Naturw. 

Ges. 1870-71:429 (1872). 

B: Dicranum albescens C. Müll., Bot. Zeit. 

11: 36 (1853). 

T: India: Nilghiri Mtns., Schmid s.no. 

(holotype destroyed in B, lectotypus BM). 

CU: C. zollingerianus (C. Müll.) Bosch. & 

Lac. cf. Frahm, Bryol. Beitr. 7:10 (1987) = 

C. fragilis (Brid.) B.S.G. ssp. goughii (Mitt.) 

J.-P. Frahm, Trop. Bryol. 61, 1991. 

albidovirens Herz.Biblioth. Bot. 87:19 (1916). 

T: Bolivia, Quellwiese an der Waldgrenze 

ber Tablas 3400 m, Herzog 2782 (holotype 

JE, isotypes B, H-BR, FH, NY, S.) 

albofoliatus J.-P. Frahm, Bryol. Beitr. 7:10 

(1987) nom. nov. pro C. laetevirens (Broth.) 

J.-P.Frahm hom. illeg. 

albovaginatus Borth. in Bruehl, Rec. Bot. Surv. 

India 13: 122 (1931), nom. nud. 

alienus Dix. in Christ., Res. Norw. Sci. Exp. Tristan 

da Cunha 48:19 (1960) 

T: Tristan d’Acunha. Christophersen 1188 

(lectotype BM, isotype FH) 

CU: C. arctocarpus (Hornsch.) Mitt. ssp. 

madecassus (Besch.) J.-P.Frahm cf. Frahm, 

Bryoph. Bibl. 31:16 (1985). 

alleizettii Thér. & P. Varde, Ann. Crypt. Exot. 

1:279 (1928). 

T: China: Tonkin environs de Hanoi, 

Alleizette 117 (holotype PC) 

CU: C. flexuosus (Hedw.) Brid. cf. Frahm, 

Bryol. Beitr. 7: 97 (1987). 

alopecurus (C. Müll.) Kindberg, Enum. Bryin. 

Exot. 88. 1889. 

B: Dicranum alopecurum C. Müll., Linnaea 

43: 401. 1882. 

T: Type. Argentina. „Rio Secco inter Oran et 

S. Andres“, Lorentz s. no. (holotype destroyed 

at B; lectotypus H-BR, isotypes JE, S). 

OU: C. annotinus Mitt. cf. Frahm, Nova 

Hedwigia 30:837 (1978). 

19 

CU: C. heterostachys (Hampe) Jaeg. 

alopecurus (C. Müll.) Kindberg var. bolivianus 

Thér. Rev. Bryol. Lich. 11:48 (1939). 

CU: C. bolivianus Thér. ex J.-P. Frahm 

alpestris Schimp. in Lor., Bryol. Notizb. 13 

(1865) nom. nud. 

CU: Dicranodontium aristatum Schimp. cf. 

Milde, Bot. Zeit. 28:417 (1870). 

alpicola (C. Müll.) Jaeg.Ber. S. Gall. Naturw. 

Ges. 1870-71:431 (1872). 

B: Dicranum alpicola C. Müll. Bot. Zeit. 16: 

161 1858 

T: Aethiopia, in summis alpibus: Schimper 

12 (lectotypus PC). 

CU: C. fragilis cf. Frahm, Cryptogamie Bryol. 

Lich. 6:208 (1985). 

alpigena Broth., Sitzungsber. Ak. Wiss. Wien, 

math.nat. Kl. 133:562 (1924) . 

T: China, Yünnan, v. Handel-Mazetti 47 

(holotype H-BR, isotype S). 

CU: C. schimperi Milde cf. Frahm, 

Cryptogamie Bryol. Lich. 3(1):63 (1982). 

alpinus Schimp., Musci Eur. Nov. Bryol. Eur. 

Suppl. Campyl. 1-2:5 (1864) 

CU: Dicranodontium denudatum (Brid.) Hag. 

ssp. alpinum (Schimp.) Giac. 

altissimus (C. Müll.) Jaeg., Ber. S. Gall. Naturw. 

Ges. 1877-78:381 (1880). 

B: Dicranum altissimum C. Müll., Flora 58: 

530 (1875) 

T: Type: Colombia, Antiquoia, Wallis s.no. 

(holotype destroyed at B, lectotypus NY, 

isolectotype S). 

CU: C. jamesonii (Hook.) Jaeg. cf. Robinson, 

The Bryologist 70:14 (1967). 

alto-filifolius (C. Müll.) Par. Ind. Bryol. 

ed.2,1:297 (1904) 

B: Dicranum alto-filifolium C. Müll., 

Hedwigia 39: 253. (1900) 

T: Brazil, Serra Geral, Ule 835 (holotype 

destroyed at B; lectotype H-BR). 

CU: C. filifolius (Hornsch.) Mitt. cf. Frahm, 

Bryoph. Bibl. 5:16 (1975) = Bryohumbertia 

filifolia (Hornsch.) J.-P.Frahm cf. Frahm, 


alto-virescens (C. Müll.) Par. Ind. Bryol. Suppl. 

88 (1900) 

B: Dicranum alto-virescens C. Müll., 

Hedwigia 38: 152 (1881) 

T: Natal, Jammerlappen Dittrich s.n. .

20 

(holotype destroyed at B, lectotypus H-BR). 

CU: C. inerangae (C. Müll.) Par. cf. Sim, 

Trans. R. Soc. S. Afr. 15:173 (1926) = C. 

robillardei Besch. cf. Frahm, Bryoph. Bibl. 

31:164 (1985). 

alvarezianus Card., Trans. R. Soc. Edinburgh 

48:73 (1911). 

T: Gough Island: Rudmose Brown s.no. 

(holotype PC). 

CU: C. eximius Reichdt. cf. Frahm, Bryoph. 

Bibl. 31:67 (1985). 

amabilis (C. Müll.) Par., Ind. Bryol. 239 (1888). 

B: Dicranum amabile C. Müll., Linnaea 43: 

399 (1882). 

T: Argentina, Cuesta de Buyuyu, Lorentz 

s.no. (holotype destroyed). 

CU: C. pyriformis (Schultz) Brid. cf. Frahm, 

Lindbergia 8(1):49 (1982). 

ambiguus Thér., Rec. Publ. Soc. Havraise Etud. 

Div. 1922(2):114 (1922). 

T: Madagascar, Massif d’Adringitra, Perrier 

de la Bathie s.no. (holotype PC). 

amblyacron (Ren.) Ren. & Card., Prodr. Fl. 

Bryol. Madag. Suppl. 20 (1909) 

CU: C. cambouei var. amblyacron Ren. 

amboroensis Thér., Rev. Bryol. Lich. 11: 54. 

1939. 

T: Bolivia, Cerro Amboro, Herzog 273 

(holotype JE; isotypes S, PC). 

ampliretis (C. Müll.) Par., Ind. Bryol. Suppl. 88 

(1900). 

B: Dicranum amplirete C. Müll., Hedwigia 

38: 81 (1899). 

T: South Africa, Montagu Pass, Rehmann 

60 (holotype destroyed at B, lectotype PC). 

CU: = C. flaccidus Ren. & Card. cf. Frahm, 

Nova Hedwigia 39: 591 (1984) non C. 

lepidophyllus (C. Müll.) Jaeg. cf. Sim, Trans. 

R. Soc. S. Afr. 15:171 (1926). 

anderssonii (C. Müll.) Jaeg., Ber. St. Gall. 

Naturw. Ges. 1870-71: 436. (1872). 

B: Dicranum anderssonii C. Müll., Bot. 

Zeitschr. 14: 169. (1872). 

T: Ecuador, Galapagos Isl., Charles Isl., 

Andersson s. n. (holotype destroyed at B; 

lectotype NY; isolectotypes FH, H-SOL, 

S). 

andicola Thér., Rev. Bryol. Lich. 11: 43. 1939. 

T: Bolivia, Chojnacota, Herzog 2976a 

(isotype JE). 

CU: C. edithae Herz. cf. Frahm. Nova 

Hedwigia 39:157 (1984). 

andreanus Card. & P. Varde, Rev. Bryol. 49:37 

(1922). (*) 

T: India: Kodaikanal, Andre 70 (holotype 

PC). 

angolensis Mach. & Dix. Bull. Torrey Bot. Club 

43:65 (1916). 

T: Angola, Gambos, on the ground at 1250m 

alt. April 1915, Carvalho s.n. (holotype BM). 

CU: C. savannarum (C. Müll.) Mitt. cf. Frahm 

Bryoph. Bibl. 31:171 (1985). 

angustelimbatus Thér. & P. Varde, Bull. Soc. Bot. 

France 71:1055 (1924). 

T: Usambara, mission de la Gare, Seoul s.n. 


CU: = C. nivalis (Brid.) Brid. cf. Frahm. 

Bryoph. Bibl. 7: 11, 1987 

OU: C. leucochlorus (C. Müll.) Par. cf. Bizot 

& Kilbertus, Rev. Bryol. Lich. 45:85 (1979). 

angustialatus Thér. Smiths. Misc. Coll. 85(4):2. 

1931. 

T: Mexico. Río Frío, Brother Amable 1724 

(holotype PC; isotypes FH, H-BR). 

CU: C. albidovirens Herz. cf. Frahm, 

Lindbergia 7:29 (1981). 

angusticosta J.-P.Frahm Bryoph. Bibl. 5:17 

(1975) nom. nov. pro C. angustinervis Herz. 

hom. illeg. 

CU: C. viridatus (C. Müll.) Broth. cf. Frahm, 

Revue Bryol. Lich. 45:166 (1979). 

angustifolius Warnst. Hedwigia 57:81 (1915) 

T: Brazil, Sao Paulo Usteri s.n. (holotype B) 

CU: C. pilifer Brid. fo. angustifolius 

(Warnst.) J.-P.Frahm Rev. Bryol. Lich. 

45:158 (1979). 

angustilimbatus Bartr. Trans. Brit. Bryol. Soc. 

1(4):466 (1951) hom. illeg. 

T: Australia, Karri Forest, Pemberton, Smith 

81 (holotype FH). 

CU: C. australis Catcheside & J.-P. Frahm. 

angustinervis Dix.S. Afr. J. Sc. 18:303 (1922). 

T: Type: „Central Africa“, Bishop 

Hannington s.no. (isotype NY). 

CU: Campylopus perpusillus Mitt. cf. Sim, 

Trans. R. Soc. S. Afr. 15:180 (1926) non 

Microcampylopus perpusillus 

(Mitt.) Broth. 

angustinervis Herz., Mem. Soc. Fl. Fauna 

Fennica 25: 54. 1954, hom. illeg. non Dixon

1922. 

T: Brazil, Estacao Biologica Alto da Serra, 

Hoehne 528 (holotype JE; isotypes B, S) 

OU: C. angusticosta J.-P.Frahm nom. nov. 

CU: C. viridatus (C. Müll.) Broth. cf. Frahm, 


angustiretis (Aust.) Sull. & Jam., Man. Moss. N. 

Am. 80. 1884. 

B: Dicranum angustirete Aust., Bot. Gaz. 

4:150. 1879. 

T: Florida, Jacksonville, Austin s. no. 


annamensis Par. & Broth., Rev. Bryol. 34:42 

(1907). 

T: Annam, Langbian, ad ripa torr. Komly 

1450m, Eberhardt s.no. (isotype S). 

CU: C. surinamensis C. Müll. cf. Frahm, 

Cryptogamie Bryol. Lich. 3(1):61 (1982) =?= 

C. laxitextus Lac.cf. Frahm, Bryol. Beitr. 7: 

12 (1987). 

annotinus Mitt., J. Linn. Soc. Bot. 12: 80. 1869. 

T: Colombia, „Andes Bogotenses, in sylvis 

prope Pacho, Weir 214 (lectotype NY; 

isolectotype H-BR). 

CU: C. heterostachys (Hampe) Jaeg. cf. 

Frahm, J. Hatt. Bot. Lab. 44:485 (1978). 

aongstroemii C.Müll. in Aongstr., Oefv. K. Vet. 

Ak. Foerh. 33(4):54 (1876). 

T: St. Mauritius, Andersson s.n. (isotype PC, 

lectotype S, isolectotype G). 

CU: C. aureo-nitens (C. Müll.) Jaeg. cf. 


aoraiensis H. Whitt.,Mosses Society Islands 89 

(1976). 

T: Tahiti, Mt. Aorai 1500m, Whittier 2762A 

(holotype indicated to be in NY but not 

present). 

CU: ?C. eberhardtii Par. cf. Frahm, Bryol. 

Beitr. 7:12 (1987) =?= C. japonicus Broth. 

apollinairei Thér., Arch. Bot. 2(10): 187. 1928. 

T: Colombia, Bogotá, Apollinaire s.n. 

(holotype PC; isotype NY). 

CU: C. cavifolius Mitt. cf. Frahm, Nova 

Hedwigia 30:844 (1978). 

appressifolius Mitt. in Hook.f., Handb. New Zeal. 

Fl. 414 (1867). 

CU: C. clavatus (R.Brown) Wils. cf. Sainsb., 

Trans. New Zeal. Inst. 59(3):506 (1928). 

arachnoideus Herz.,Rep. Sp. Nov. Regn. Veg. 

21:29 (1925). 

21 

T: Brazil, Serra dos Orgaos, Morro Assu 

2400m, v. Lützelburg 6688 (holotype JE, 

isotypes B,S). 

CU: C. gemmatus (C. Müll.) Par. cf. Frahm, 


araucarieti (C. Müll.) Par., Ind. Bryol. Suppl. 88. 

1900. 

B: Dicranum araucarieti C. Müll., Hedwigia 

39: 254. 1900. 

T: Brazil. Serra Geral, „in terra araucarieti“, 

Ule 661 (lectotype, H-BR, isolectotypes FH, 

JE, KIEL, NY, S). 

OU: C. filifolius (Hornsch.) Mitt. var. humilis 

(Mont.) J.-P.Frahm cf. Frahm, Nova 

Hedwigia 29:247 (1978). 

CU: Bryohumbertia filifolia (Hornsch.) J.- 

P.Frahm var. humilis (Mont.) J.-P. Frahm. 

arbogastii Ren. & Card. in Ren., Rev. de Bot. 

9:289 (1891) 

T: Madagascar, „in terra arenosa“, Arbogast 

s. no. (holotype PC, isotype G). 

CU: C. julaceus Jaeg. ssp. arbogastii (Ren. 

& Card.) J.-P.Frahm, Bryoph. Bibl. 31:109 

(1985). 

arboricola Card. & Dix., New Zealand Inst. Bull. 

3(3): 90 1923 

T: New Zealand, Te Aroha, growing dense 

mats on tree trunk, Klaud & Chase 276 (BM). 

CU: C. purpureocaulis cf. Frahm 

Cryptogamie Bryol. Lich. 6:205 (1985) 

archboldii (Bartr.) Bartr. Brittonia 9:34 (1957) 

T: Indonesia West Irian, Jayawijaya. Mt. 

Wilhelmina, Lake Habbema,3225 m. Brass 

9450 (FH) 

CU: C. exasperatus cf. Frahm et al., Acta 

Bot. Fenn. 131:78 (1985). 

arctocarpus (Hornsch.) Mitt., J. Linn. Soc., Bot. 

12: 87. 1869. 

B: Dicranum arctocarpum Hornsch., Fl. bras. 

1(2): 12. 1840. 

T: Uruguay. Montevideo, Sellow s.n. 

(holotype destroyed at B; lectotype BM). 

arctocarpus (Hornsch.) Mitt. ssp. madecassus 

(Besch.) J.P. Frahm J. Linn. Soc. Bot. 12: 87 

(1869). 

B: Campylopus madecassus Besch. 

arctocarpus (Hornsch.) Mitt. var. caldensis 

(Aongstr.) J.-P. Frahm Fl. Neotr. 54: 55 

(1991). 

B: Campylopus caldensis Aongstroem,

22 

Öfvers F”rh. Kongl. Svenska Vetensk.-Akad. 

33(4): 5 (1876) 

T: Brazil without locality Widgren s.n. 

(holotype S ). 

arctocarpus (Hornsch.) Mitt. fo. strictifolius 

(Broth.)J.-P.Frahm Revue Bryol. Lich. 

45:138 (1979). 

B: C. strictifolius Broth. 

arcuatus (Brid.) Jaeg., Ber. S. Gall. Naturw. Ges. 

1870-71: 420 (1872). 

B: Dicranum arcuatum Brid. Brid. Spec. 

Musc. 1: 212 

T: „In insula Bourbonis monte excelso Piton 

des Neiges dicto...“, Bory s. no. (holotype 

in B-Bridel lacking, lectotypus nov. 

BM, isolectotype L). 

arcuatus R.Brown ter., Trans. New Zealand Inst. 

29:474 (1897) hom. illeg. 

CU: C. clavatus (R.Brown) Wils. cf. Dixon, 

New Zealand Inst. Bull. 3:88 (1923) 

arduennae Lib., Pl. Crypt. Arduenn. 106 (1831). 

CU: C. flexuosus (Hedw.) Brid. cf. Pire & 

March., Bull. Soc. R. Bot. Belg. 11:7 (1872). 

arenaceum (Broth.) J.-P. Frahm, J. Bryol. 8(2): 

258. (1974). 

B: Thysanomitrion arenaceum Broth., 

Denkschr. Akad. Wiss. Wien math.-nat. Kl. 

83: 264. 1926. 

T: Brazil, Sao Paulo, Faxina, Schiffner 1100 

(lectotype H-BR). 


Frahm, Bryol. Beitr. 7:13 (1987). 

arenarius R.Brown ter., Trans. New Zeal. Inst. 

29:475 (1897). 

CU: C. clavatus (R.Brown) Wils. p.pte. cf. 

Dixon, New Zeal. Inst. Bull. 3(3):77 (1923). 

arenicola (C. Müll.) Mitt. J. Linn. Soc. Bot. 12:77 

(1869). 

B: Dicranum arenicola C. Müll., Bot. Zeit. 

13: 762 (1855). 

T: Brazil, Itajahi, Pabst s.n. (holotype 

destroyed at B; lectotype BY; isolectotypes 

GOET, H-SOL, JE, S). 

CU: C. trachyblepharon (C. Müll.) Mitt. cf. 

Frahm, Rev. Bryol. Lich. 45:164 (1979). 

areodictyon (C.Müll.) Mitt., J. Linn. Soc. Bot. 

12: 77 (1869). 

B: Dicranum areodictyon C. Müll., Syn. 

Musc. 394. 1848. 

T: Type. Venezuela, Prov. Mérida, Sierra 

Nevada 9000 ft., Funck & Schlim 1082 

(holotype destroyed in B; lectotype 

NY; isolectotype H-BR). 

areodictyon Schimp. in Mandon, Pl. Boliv. n. 

1612 nom. nud. 

OU: C. leucognodes (C. Müll.) Par. cf. C. 

Müll., Nuov. Giorn. Bot. Ital. n.ser.4:164 

(1897). 

CU: C. argyrocaulon (C. Müll.) Broth. 

argutidens Broth. & Par. Rev. Bryol. 34:93 

(1907). 

T: Guinea, Diaguisse, Pobeguin s.n. (holotype 

PC). 


Frahm, Cryptogamie Bryol. Lich. 3(1):60 

(1982). 

argutidens Broth. & Par. var. angustifolius Thér. 

& P.Varde, Rev. Bryol. n.ser. 4: 66 (1932). 

T: Gabun, entre Moumba et Mbounga, Le 

Testu 3139 (holotype PC). 



(1982). 

argyrocaulon (C.Müll.) Broth., Nat. Pfl. 1(3): 

332. 1901. 

B: Dicranum argyrocaulon C.Müll. Linnaea 

38: 588 (1874). 

T: Colombia Prov. Antioquia, Páramo de 

Sonson 10-20 000 ft., Wallis s.n. (holotype 

destroyed in B; lectotype H-BR). 

aristatus Kindb. in Weim., Foert. Skand. Vaexter 

Moss. 2:60 (1937) 

CU = Dicranodontium asperulum (Mitt.) 

Broth. cf. Weim. l.c. 

arizonae Besch. in Ren. & Card., Rev. Bryol. 

19:77 (1892) nom. nud. 

arsenei Thér., Smiths. Misc. Coll. 78:5 (1926) 

T: Mexico, Cerro Azul, Arsene 4981 

(holotype PC) 

OU: C. destructilis (C. Müll.) Jaeg. cf. Theriot 

l.c. 85:2 (1931) = C. chrismarii (C. Müll.) 

Mitt. cf. Williams, N. Am. Fl. 15:136 (1913) 

CU: C. nivalis (Brid.) Brid. 

aspericuspes Thér. & P. Varde Rev. Bryol. Lich. 

6.136 (1934). 

T: „Africa trop. occ., Gabo, prope Assoc 

Ngoum, Le Testu , Musci selecti et critici 9 

(isotypes S,U,L,NY). 

CU = C. savannarum (C. Müll.) Mitt. cf. 

Frahm, Cryptogamie Bryol. Lich. 3:60

(1982). 

asperifolius Mitt., J. Linn. Soc. Bot. 12: 79. 1869. 

T: Ecuador. Abitagua 7000 ft., Spruce 54 

(lectotype PC; the original material in the 

Mittenherbarium , NY, is lacking). 

OU: Dicranodontium asperifolium (Mitt) 

Broth., Nat. Pflanzenfam. ed. 2, 10: 190 

(1924). 

asperulus (Mitt.) Kindb., Bih. K. Svensk Vet. Ak. 

Handl. 7:88 (1883) 

CU: Dicranodontium asperulum (Mitt.) 

Broth. 

assimilis Thér. & P. Varde Rev. Bryol. n. ser.l:89 

(1928). 

T: Gabun, propre Nzamalen, Le Testu 309 

(isotypes L,U). 



(1982). 

atlanticus B.H. Allen, The Bryologist 9: 447 

(1990). 

T: Panama, Bocas del Toro, Allen 5713 

(holotype MO, isotype PMA). 

atratus Bartr., Contr. U.S.Nat. Herb. 26:61 (1928) 

hom. illeg. 

T: Costa Rica, Cerro de las Vueltas 3000m, 

Standley 43686 (holotype,FH; isotypes H-BR, S). 

CU: C. paramoensis Bowers nomen novum, 

Bryologist 77(2): 152 (1974). 

atratus Broth., Trans. Linn. Soc. Bot. ser. 2,6: 

89. 1900. 

T: Venezuela, Mt. Roraima, McConnell & 

Quelch 527 (holotype H-BR; isotypes K, NY, 

S) 

CU: C. richardii Brid. cf. Frahm, Nova 

Hedwigia 39:598 (1984). 

atratus Schimp. in C. Müll., Gen. Musc. Fr. 265 

(1900) nom. nud. 

atrofuscescens Bartr. Contr. U.S.Nat. Herb. 

37:155 (1965) 

T: Papua New Guinea. Simbu, east slopes of 

Mt. Wilhelm 13100 ft, J.D. Collins 29941 

(holotype FH). 

CU: C. macgregorii Broth. & Geh. cf. Frahm 

et al., Acta Bot. Fenn. 131:82 (1985). 

atro-luteus (C. Müll.) Par. Ind. Bryol. Suppl. 89 

(1900) 

B. Dicranum atroluteum C. Müll., Hedwigia 

38: 80 (1899). 

T: Cape Town, Rehmann 63 (lectotypus nov., 

23 

PRE, isolectotypes BOL, G, H-BR, H-SOL, 

L, NH, NY, PC). 

CU: C. bicolor (C. Müll.) Wils. in Hook. f. 

ssp. atroluteus (C. Müll.) J.-P.Frahm Bryoph. 

Bibl. 31:37 (1985). 

atrosordidus C.Müll. ex Broth. Bot. Jahrb. 

24:233 (1897). 

T: Kamerun, supra Bueam pagum inter 2100 

et 3000m. Dusen 347 (lectotype. NY, isotypes 

PC,S,BM,G). 

CU: C. pilifer Brid. cf. De Sloover, Bull. Jard. 

Bot. Nat. Belg. 51:228 (1981). 

atrovirens De Not., Syll. Musc. Ital. 221 (1838). 

T: Italy in valle Intrasca, de Notaris s.n. 

(lectotype RO). 

atrovirens De Not. var. cucullatifolius J.-P. 

Frahm, Bryologist 83: 574 (1980). 

T: USA, Alaska, Washington Bay, Kuju Island, 

Eyerdam 5269 (holotype DUKE). 

attenuatus (Stirt.) Dix. J. Bot. 61:14 (1923) 

CU: C. brevipilus B.S.G. fid. Dix. l.c. 

attenuatus Schimp. in Jaeg., Ber. S. Gall. Naturw. 

Ges. 1870-71:444 (1872) nom. nud. 

aubertii (Bel.) Thér. in Broth., Nat. Pfl. 

ed.2,10:187 (1924) 

CU: C. cf. hildebrandtii (C. Müll.) Jaeg. cf. 


aureonitens (C.Müll.) Jaeg. Ber. S. Gall. Naturw. 

Ges. 1870-71:435 (1872). 

B: Dicranum aureonitens C. Müll. Syn. Musc. 

1:406 (1948). 

T: „Insula Bourbonis“ Bory St. Vincent s.n. 

(holotype destroyed at B, lectotypus nov. S, 

isolectotype G). 

aureonitens ssp. recurvifolius (Dus.) J.-P.Frahm, 

Bryol. Beitr. 7:14 (1987). 

B: Campylopus recurvifolius Dus. 

aureo-viridis (C. Müll.) Schimp. ex Par., Ind. 

Bryol. Suppl.89 (1900) 

B: Dicranum aureo-viride C. Müll., Hedwigia 

38: 85 (1899) 

T: Südafrika, Gnadenthal, Breutel s.n. 

(lecotytype PC) 


Bryoph. Bibl. 31:160 (1985). 

OU: C. atroluteus (C. Müll.) Par. cf. Sim, 

Trans. R. Soc. S. Afr. 15:178 (1926). 

aurescens Stirt., Ann. Scott. Nat. Hist. 8:104 

(1899). 

CU: C. brevipilus B.S.G. var. auriculatus Dix.

24 

cf. Dixon, J. Bot. 61:15 (1923). 

aureus Bosch & Lac., Bryol. Jav. 1:80 (1858). 

T: Indonesia, Java, Mandalawagie 9200' 

Junghuhn s.n. (lectotype L). 

CU: C. schmidii (C. Müll.) Jaeg. cf. Frahm, 

Bryol. Beitr. 7:15 (1987) 

auribrunneus (C.Müll.) Broth., Nat. Pfl. 

1(3):332. 1901. 

B: Dicranum auribrunneum C.Müll., 

Hedwigia 39:260. 1900. 

T: Brazil, Goyaz, Serra de Balisa, Ule 1533 

(holotype destroyed at B; lectotype PC). 

CU: C. aemulans (Hampe) Jaeg. cf. Frahm, 


auriculatus Wils. in MacKinlay, Proc.Nat. Hist. 

Soc. Glasgow 1:91 (1869). 

OU: C. schwarzii Schimp. fid. MacKinlay l.c. 

CU: C. gracilis. 

aurificus (C. Müll.) Kindb., Enum. Bryin. Exot. 

88 (1889). 

B: Dicranum aurificum C. Müll., Linnaea 43: 

399 (1882). 

T: Argentina uruguensis, in collibus 

australibus, Lorentz s.n. (lectotypus C). 

CU: C. introflexus (Hhedw.) Brid. cf. Frahm, 


australiensis Duby, Mem. Soc. Phys. Hist. Nat. 

Geneve 20:259 (1870) 

T: Australia, without locality (isotype S). 

CU: C. introflexus (Hedw.) Brid. cf. Frahm, 


australis Catcheside & J.-P. Frahm, J.Bryol. 13: 

360 (1985). nom. nov. pro C. angustilimbatus 

Bartr. 

austro-alpinus (C. Müll.) Kindb. Enum. Bryin. 

Exot. 88 (1889). 

B: Dicranum austro-alpinum C. Müll., 

Linnaea 43: 378 (1882). 

austrostramineus Thér., Bull. Soc. Linn. Seine 

Marit. 11: 103 (1924). 

T: Kerguelen, Baie au Nord du Cap Dauphin, 

Peau 16 (holotype PC). 

austrosubulatus Broth. & Geh., Öfv. Finska Vet. 

Soc. Förhandl. 37: 154 (1895). 

T: Papua N.G., Milne Bay, Mt. Musgrave, W. 

Macgregor 345 (holotype H-BR). 

azoricus Mitt. in Godm., Nat. Hist. Azores 292 

(1870). 

T: Azores, St. Miguel, Godman s.n. (holotype 

NY). 

OU: C. flexuosus (Hedw.) Brid. var. azoricus 

(Mitt.) Thér., C. pyriformis (Schultz) Brid. 

var. azoricus (Mitt.) A.J.Smith. 

CU: : C. pyriformis (Schultz) Brid. cf. Corley 

& Frahm, J. Bryol. 12:190 (1982). 

B 

balanseanus Besch., Ann. Sci. Nat. Bot. ser. 5, 

18: 199 (1873), C. balansae Besch. ex Kindb. 

Enum. Bryin. Exot. 49 (1888). 

T: „Nouvelle-Caledonie, Noumea, ad 

truncos“ Balansa 749 (holotype PC, isotype 

S) introflexus 

OU: C. aureus Bosch & Lac. cf. Fleisch., 

Musci Fl. Buitenzorg 1:110 (1904). 



barbuloides Broth. in Bruehl, Rec. Bot. Surv. 

India 13(1):123 (1931) nom. nud. 

M: NW Himalaya, Mussoorie, Dhanaulti 

7000 ft., on ground, Singh 3698 (S). 

CU: C. subulatus Schimp. cf. Frahm, 


bartlettii Bartr., J. Wash. Ac. Sc. 22:477 (1932) 

T: Honduras, Bartlett 12973 (holotype FH; 

isotypes DUKE, F, MICH, NY, S). 

OU: C. savannarum (C. Müll.) Mitt. ssp. 

bartlettii (Bartr.) Florsch. Mosses of 

Suriname 79 (1964). 

CU: C. savannarum mod. bartlettii. 

bartramiaceus (C. Müll.) Thér. in Broth. Nat. Pfl. 

ed. 2, 10: 185 (1924) hom. illeg. 

B: Dicranum b. C. Müll. Hedwigia 38: 86 

1899 

T: Cape Town, Rehmann 37 (lectotype BM). 

OU: Dicranum leucobasis C. Müll. Hedwigia 

38:78 (1899) var. bartramiaceus C. Müll.), 

C. subbartramiaceus Rehm. in sched., 

CU: C. introflexus (Hedw.) Brid. cf. Sim, 


bartramiaceus (C. Müll.) Thér. var. laevis P. 

Varde & Thér. Bull. Soc. Bot. France 87: 353 

( ) 

T: Congo bel. Kivu, volcan Mukavura, 

Humbert 7528 (holotype PC) 


bartramiaceus (C. Müll.) Par., Ind. Bryol. 240 

(1900). 

OU: C. bartramiaceus (C. Müll.) Dix. & 

Gepp. Bull. Misc. Inf. Kew 1923:197 (1923)

sensu Index Muscorum, non C. atroluteus (C. 

Müll.) Par. cf. Sim, Trans. R. Soc. S. Afr. 

15:171 (1926), Frahm, Revue Bryol. Lich. 

42:613 (1976), non C. stramineus cf. Bizot 

& Kilbertus, Revue Bryol. Lich. 45:94 (1979), 

excl. var. laevis P. Varde & Thér. Bull. Soc. 

Bot. France 87:353 (1940). 

CU: C. nivalis (Brid.) Brid. cf. Frahm, 

Bryoph. Bibl.31:35 (1985). 

bartramioides Broth. in Whitel., Proc. Linn. 

N.S.Wales 7:277 (1882) nom. nud. 

CU: Dicranoloma bartramioides (Broth.) Par. 

basalticolus (C. Müll.) Par. Ind. Bryol. 89 (1900) 

B: Dicranum basalticolum C. Müll., 

Hedwigia 38: 82 (1899). 

T: Cape, Somerset-East, Boschberg, 

MacOwen s.n. (lectotype BM). 

OU: Campylopus chlorophyllosus (C. Müll.) 

Jaeg. cf. Sim, Trans. R. Soc. S. Afr. 15:177 

(1926), C. subchlorophyllosus cf. Magill, Fl. 

S. Afr. Bryoph. 1(1):146 (1981). 

CU: Sphaerothecium subchlorophyllosum (C. 

Müll.) J.-P. Frahm. 

batjanensis Broth. in Warb., Monsunia 1:42 

(1899). 

CU: Dicranodontium uncinatum (Harv.) Jaeg. 

cf. Fleisch., Musci Fl. Buitenzorg 1:89 (1904). 

beauverdianus Thér., Bull. Soc. Bot. Geneve 

26:78 (1936). 

belangeri Thér. Bull. Mus. Hist. Nat. Paris 34:115 

(1928). (*) 

T: „Ins. Ascensionis, ex dono cl. Belanger“. 

bellii R.Brown ter., Trans. New Zeal. Inst. 29:469 

(1897). 

OU: C. capillatus Hook.f. & Wils. cf. Dix., 

New Zealand Inst. Bull. 3(3):86 (1923), C. 

pallidus Hook.f. & Wils. cf. R. Soc. N. 

Zealand Bull 5:116 (1955). 

CU: C. pyriformis (Schultz) Brid. 

benedictii Herz. Beih. Bot. Centralbl. 36:50 

(1910). 

T: Type. Bolivia, Cochabamba, Abra de San 

Benito, Herzog s. n. (holotype JE; isotype S). 

OU: C. ptychotheca Herz. cf. Frahm, Nova 

Hedwigia 39:161 (1984). 

CU: C. reflexisetus (C. Müll.) Broth. cf. 

Frahm, Bryol. Beitr. 7: 17 (1987). 

bequaertii Thér. & Nav. Bull. Soc. R. Bot. Belg. 

60:17 (1927). 

T: Zaire, Ruwenzori, Butagu, Bequaert 4974 

25 


CU: = C. savannarum (C. Müll.) Mitt. cf. 

Frahm, Lindbergia 8(1):48 (1982). 

bermudianus Williams in Britton, J. New York 

Bot. Garden 13: 193. 1913. 

T: Bermuda, Paget Marsh, Brown 651 

(holotype NY; isotypes CANM, DUKE) 

CU: C. trachyblepharon cf. Frahm, Herzogia 

5:518 (1981). 

berteroanus Duby, Flora 58:282 (1875). 

HS: C. berteroi Duby ex Kindb., Enum. 

Bryin. Exot. 49 (1888). 

T: Juan Fernandez Is., Betero s.n. (holotype 

G, isotype S). 

CU: C. introflexus (Hedw.) Brid. cf. Thér., 

Rec. Publ. Soc. Havraise Etud. Div. 88:30 

(1921). 

bertramii Hampe in Limpr., Laubm. Deutschl. 

1:375 (1886) nom. nud. 

CU: Paraleucobryum longifolium (Hedw.) 

Loeske var. subalpinum (Milde) Cas.Gil. cf. 

Limpr. l.c. 

bessonii Ren. & Card. in Ren., Rev. de Bot. 9:397 

(1891). 

T: Madagaskar, Fianarantsoa - Betsileo, 

Besson s.n. (holotype PC). 

OU: C. polytrichoides De Not. var. bessonii 

(Ren. & Card.) Ren. & Card. 

CU: C. pilifer Brid. ?var. lamellatus (Mont.) 

Gradstein & Sipman. 

bewsii Sim Trans. R. Soc. 5. Afr. 15:176 (1926). 

T: Natal, Knoll, Hilton Road, Sim 9838 

(lectotype L). 

CU: C. pilifer Brid. cf. Magill, Fl. S. Afr. 

Bryoph. 1(1):152 (1981). 

beyrichianus Duby, Mem. Soc. Phys. Hist. Nat. 

Geneve 20:359 (1870). 

HS: C. beyrichii Duby ex Kindb. Enum. 

Bryin. Exot. 49 (1888). 

T: „E-Brasilia“, hb Duby (holotype G). 


Bryol. Beitr. 7: 17 (1987). 

bicolor (C.Müll.) Wils. in Hook. f., Fl. Nov. Zel. 

2:69 (1854). 

B: Dicranum bicolor Hornsch. ex C. Müll., 

Syn.Musc. 1: 392. (1848). 

T: Nova Hollandia, Sieber 9. 

bicolor (C. Müll.) Wils. ssp atroluteus (C. Müll.) 

J.-P.Frahm, Bryophytorum Bibliotheca, 31: 

37 (1985).

26 

B: Dicranum atroluteum C. Müll., Hedwigia 

38: 80 (1899). 

OU: Campylopus atro-luteus (C. Müll.) Par. 

Ind. Bryol. Suppl. 89 (1900) . 

T: Cape Town, Rehmann 63 (lectotypus nov., 

PRE, isolectotypes BOL, G, H-BR, H-SOL, 

L, NH, NY, PC). 

CU: C. bicolor s.str. 

bicolor (C.Müll.) Wils. var. ericeticola (C. Müll.) 

Dix., New Zealand Inst. Bull. 3: 87 (1923). 

B: C. ericeticola C. Müll., Abh. Naturw. Ver. 

Bremen 16(3):496 (1900) 

birgeri Card., Bull. Herb. Boiss. ser.2,5:1000 

(1905). 

T: Islas Malwinas, Birger 207 (isotype S). 

CU: C. acuminatus Mitt. cf. Frahm, Bryol. 

Beitr. 7: 17 (1987). 

blindiaceus C. Müll. Gen. Musc. Fr. 274 (1900) 

nom. nud. 

blindioides Broth. in Skottsberg., Nat. Hist. Juan 


T: Juan Fernandez, Masafuera, Skottsberg 60 

(H-BR). 


Bryol. Beitr. 7: 17 (1987). 

blumei (Dozy & Molk.) Bosch & Lac., Bryol. 

Jav. 1:81 (1858). 

B: Trichostomum blumei Dozy & Molk., Ann. 

Sc. Nat. Bot. ser. 3,2: 316 (1844). 



OU: C. richardii Brid. cf. Dixon, J. Bot. 

60:289 (1922). Thysanomitrion blumei Card. 

Annuair. Cons. Jard. Bot. Geneve 15-16: 161 

(1912) 

bogotensis Thér., Rev. Bryol. Lich. 11:60 (1939). 

CU: Sphaerothecium phascoideum (Hampe) 

Williams cf. Robinson, The Bryologist 70:11 

(1967). 

boivinianus Besch., Ann. Sci. Nat. Bot. ser 6, 9: 

319 (1880). 

T: Ile Maurice, Boivin 89 (B,S). 

HS: C. boivinii Besch. ex Kindb. Enum. 

Bryin. Exot. 88 (1889). 



bolivianus Theriot ex J.-P. Frahm, Lindbergia 7: 

28 (1981). 

T: Bolivia, Sillunticava, Yungas de la Paz, 

Troll 122 a (holotype PC, isotype JE). 

CU: C. oblongus Thér. cf. Frahm, Bryol. 

Beitr. 7: 18 (1987). 

bolivarensis Bartr., Fieldiana Bot. 28: 4 (1951). 

T: Venezuela, Bolivar: Mt. Roraima, 

Steyermark 58895 (lectotype FH; isotype F). 

CU: C. cuspidatus (Hornsch.) Mitt. cf. Frahm, 

J. Hatt. Bot. Lab. 44:485 (1978). 

boryanus Besch. Ann. Sc. Nat. Bot. ser 5, 9: 

319 1880 Reunion Plaine des Chicots Bory 

s.n. PC nivalis 

boryanus Besch., Ann. Sc. Nat. Bot. ser. 6,9:319 

(1880). 

HS: C. boryi Besch. ex Kindb. Enum. Bryin. 

Exot. 88 (1889). 

T: Bourbon, Plaine des Chicots, Bory 


CU: C. chrismarii (C. Müll.) Mitt. cf. Frahm, 

Lindbergia 8(1):46 (1982) = C. nivalis (Brid.) 

Brid. 

boswellii (C. Müll.) Par. Ind. Bryol. Suppl. 89 

(1900). 

B: Dicranum boswellii C. Müll., Flora 82: 442 

(1896). 

T: Hawaii, without locality, Wheeler s.n. 

1879, Boswell s.n. (syntypes, could not be 

located). 

CU: C. fragilis (Brid.) B.S.G. ssp. 

zollingerianus (C. Müll.) J.-P. Frahm cf. 

Frahm, Trop. Bryol. 4: 61, (1991). 

brachymastyx C. Müll. ex Besch. Ann. Sc. Nat. 

Bot. ser. 6,9:324 (1880). 

T: Ile Maurice, Robillard s.n. (holotype PC, 

isotype S). 

OU: C. virescens Besch. cf. Card. in Grand., 

Hist. Madag. 39:112 (1915). 



brachymitrius (Geh. & Hampe) Kindberg, Enum. 

Bryin. Exot. 88. (1889). 

B: Dicranum brachymitrium Geheeb & 

Hampe, Flora 64: 343. 1881. 

T: Brazil, Apiahy, Puiggari 331 (lectotype, 

H-BR). 

CU: C. occultus Mitt. cf. Frahm, Revue Bryol. 

Lich. 45:155 (1979). 

brachyphyllulus (C. Müll.) Broth., Nat. pfl. 

1(3):332 (1901). 

B: Dicranum brachyphyllulum C. Müll., 

Hedwigia 39: 260 (1900). 

T: Brasilien, Minas Geraes, Caraca, ad rupes,

Ule 1360 (lectotype PC). 



(1981). 

brachyphyllus C. Müll. in Par. Ind. Bryol. 261 

(1894) nom. nud., hom. illeg. 

M: Cameroun, Mont Rumpi 1800m, Dusén 

s.n. (H-BR, PC-Thér., S). 

OU: C. subbrachyphyllus Par. nom. nud. 

CU: C. nanophyllus C. Müll. ex Broth. cf. 


brachyphyllus C.Muell. in Par. Ind. Bryol. 261 

(1894) nom. nud., hom. illeg. 

brachyphyllus Wils. ex Mitt. J. Linn. Soc. Bot. 

12:81 (1869). 

T: Andes Quitenses, Jameson 28 (holotype 

NY, isotypes B, BM). 

OU: C. rosulatus (Hampe) Mitt. cf. Robinson, 



Bryol. Beitr. 7: 18 (1987). 

brachythysanos (C. Müll.) Par., Ind. Bryol. Suppl. 

89 (1900). 

B: Dicranum brachythysanum C. Müll., 

Hedwigia 36: 97 (1897). 

T: Uruguay, Montevideo, Arechavaleta s. no. 

(holotype destroyed at B; lectotype NY). 

CU: C. occultus Mitt. cf. Frahm, Lindbergia 

7:31 (1981). 

brasiliensis Broth., Hedwigia 38 Beibl. 1:58 

(1899) nom. nud. 

M: Ule, Bryotheca bras. 109. 

OU: Dicranum brasiliense C. Müll., Bull. 

Herb. Boiss. 6: 39 (1898). 

CU: Atractylocarpus brasiliensis (C. Müll.) 

Williams cf. Frahm, Cryptogamie Bryol. 

Lich. 2(4):444 (1981). 

brassii Bartr. Farlowia 4:236 (1952). 

T: Australia, Queensland, Cape York, Brass. 

s.n. (holotype FH). 

CU: C. robillardei Besch. var. perauriculatus 

(Broth.) J.-P. Frahm Bryol. Beitr. 7: 89 

(1987). 

brevifolius Roell in Limpr., Laubm. Deutschl. 

1:393 (1897) nom. nud. 

CU: C. paradoxus Wils cf. Limpr. l.c. = C. 

flexuosus (Hedw.) Brid. cf. Hagen, K.Norsk. 

Vid. Selsk. Skrift. 1914(1):75 (1915). 

brevifolius Schimp., Musci Eur. Nov. Bryol. 

Eur.oSuppl. fasc.1-2,4:2 (1864). 

27 

T: Italien, Südtirol, Meran, Lorentz. 

CU: C. subulatus Schimp. cf. Braithw., Brit. 

Moss Fl. 1:130 (1882). 

brevipilus B.S.G. Bryol. Eur. 1:167 (1847). 

T: „Prope Fontainebleau praecedenti (C. 

longipilus) intermix...Alex. Braum (isotype 

B). 

brevipilus var. marginatulus (Geh.) Thér. in Pers 

Perss. Bot. Not. 1939: 570 (1939). 

B: Campylopus marginatulus Geh. 

CU: Ditrichum punctulatum. 

brevirameus Dix., Ann. Bryol. 2:6 (1929). 

T: Seychelles, Mahé, Delarié s.n. (S,PC). 

CU: C. pilifer Brid. x C. introflexus (Hedw.) 

Brid. cf. Frahm, Bryoph. Bibl. 31:149 (1985). 

brevis Rehm. in Geh., Rev. Bryol. 5:69 (1878) 

nom. nud. 

T: Cape, Stinkwater, Rehmann 44 (L, PC). 

OU: C. atroluteus (C. Müll.) Par. cf. C. Müll., 

Hedwigia 38:81 (1899). 

CU: C. bicolor (C. Müll.) Wils. in Hook. ssp. 

atroluteus (C. Müll.) J.-P.Frahm. 

breweri Bartr., Bol. Soc. Venez. Ci. Nat. 25: 47. 

(1963). 

T: Venezuela, Merida, Sierra de la Culata, 

Brewer s.n. (holotype FH; isotype VEN). 

CU: C. areodictyon (C. Müll.) Mitt. cf. Frahm, 

Bryol. Beitr. 7: 19 (1987). 

brittonae Williams, N.Am.Fl.15:141 (1913). 

T: Jamaica, Summet of St. John’s Peak, 

Britton 1909 (holotype NY, isotypes DUKE, 

FH, H, S). 

CU: C. shawii Wils. cf. Frahm, J. Bryol. 

13:329 (1985). 

brotherianus Par., Ind. Bryol. Suppl. 89 (1900) 

nom. nud. 

T: homotypic with C. procerus. 

OU: C. praealtus (C. Müll.)Par. cf. Broth., 

Denkschr. Akad. Wiss. Wien Math. Nat. Kl. 

83:260 (1926). 

CU: C. subcuspidatus (Hampe) Jaeg. 

brownii Par., Ind. Bryol. Suppl. 90 (1900). 


New Zeal. Inst. Bull. 3(3):88 (1923). 

brownii Par., Ind. Bryol. Suppl. 90 (1900). 

brunneo-bolax (C. Müll.) Broth., Nat. Pfl. 

1(3):333 (1901) nom. nud. 

OU: C. stricticaulis (C. Müll.) Par. cf. Frahm, 


CU: C. aemulans (Hampe) Jaeg. cf. Frahm,

28 


brunneus (C. Müll.) Par. Ind. Bryol. Suppl. 90 

(1900).(*) 

bryhnii Broth., Nat. Pfl. ed.2,10:188 (1924). 

T: Natal, Zululand, Ekombe, Titlestad s.n. 

(holotype H-BR). 

OU: C. inchangae cf. Magill, Fl. S. Afr. 

Bryoph. 1(1):146 (1981). 



bryotropii J.-P Frahm Nov Hedwigia 39: 152, 

(1984). 

T: Peru, Dep Ancash, Cordillera Blanca 

Laguna, Llanganuco 3850m, Frahm 825118 

(holotype B; isotypes C, EGR, F, FLAS, G, 

GRO, GZU, H, HBG, KR, KRA, M, MEXU, 

NAM, NFLD, NY, PC, S, U, UPS, USM). 

buchtienii Thér., Rev. Bryol. Lich. 11:53 (1939). 

T: Bolivien, Casana im Tipuanital, 1400m, 

Buchtien 265 (isotypes B, JE). 

CU: C. arctocarpus (Hornsch.) Mitt. cf. 


C 

cacti (C. Müll.) Kindb. Enum. Bryin. Exot. 88 

(1889). 

B: Dicranum cacti C. Müll., Linnaea XLIII: 

403 (1882). 

T: Uruguay, Concepion, Lorentz s. no. 

(holotype destroyed at B; lectotype BM; 

isolectotypes JE, PM, S). 


Lich. 42:902 (1976). 

cacuminis (C. Müll.) Par., Ind. Bryol. Suppl. 90 

(1900). 

B: Dicranum cacuminis C. Müll., Hedwigia 

37: 227 (1898). 

T: Jamaica, Blue Mtns. Peak, Hansen s.n. 

(holotype destroyed at B). 

CU: C. cygneus (Hedw.) Brid. cf. Thériot, 

Mem. Soc. Cubana Hist. Nat. 13:215 (1939). 

caducipilus Besch. in Par. Ind. Bryol. Suppl. 90 

(1900) nom. nud. 

CU: C. bicolor (C. Müll.) Wils. cf. Fleisch., 


caespiticius Brid., Mant. Musc. 77 (1819). 

CU: Grimmia caespiticia (Brid.) Brid. 

caespitosus (Mitt.) Jaeg. Ber. S. Gall. Naturw. 

Ges. 1870-71:434 (1872). 

CU: Dicranodontium caespitosum (Mitt.) Par. 

cagnii Negri, Annali Bot. 7. 162 (1908). 

T: „in monte Rewenzori in valle Lacuum ca. 

4000 m ad radices..“ 

CU: C. leucochlorus (C. Müll.) Par. cf. 


cailleae Ren. & Card., Bull. Soc. R. Bot. Belg. 

32: 102 (1894). 

T: Madagaskar, Nassi Comba, Caille s.n. 

(PC). 

CU: C. perpusillus Mitt. cf. Frahm, Bryoph. 

Bibl. 31:175 (1985). 

caldensis Angstr., Oefv. K. Vat. Ak. Foerh. 

33(4):5 (1876). 

T: Brazil, without locality, Widgren s.no. 

(holotype S). 


caldensis (Angstr.) J.-P.Frahm 

calochlorus Broth. & Thér. Bull. Mus. Hist. Nat. 

Paris 30:239 (1924). 

T:“British East Africa“, Foret de Lamorie, 

LePetit s.n. (PC). 

CU: C. pilifer Brid. cf. Frahm, Bryoph. Bibl. 

31:143 (1985). 

calodictyon Broth., Leafl. Philipp. Bot. 2:651 

(1909). 

T: „Philippines, Negros, Apr. 1908 nr. 9745“ 


CU: C. laxitextus Lac. cf. Frahm, Bryol. Beir. 

7: 87 (1987). 

calvatus Dix. in Luis., Broteria Cienc. Nat. 8: 46 

(1939). 

T: Tristan da Cunha, Milne s.n. (isotype NY). 

OU: C. atroluteus cf. Frahm, Herzogia 5: 521 

(1981). 

CU: C. pyriformis (K.F. Schultz) Brid. cf. 

Frahm, Bryoph. Bibl. 31: 160 (1985). 

calvus Ren. & Card., Bull. Soc. R. Bot. Belg. 

33(2):111 (1895). 

T: Madagascar, Betsileo, Ambohimatsara, sur 

terre humide, un peu tourbeuse, Berthieu s.n. 


CU: C. aureus Bosch & Lac. cf. Frahm, 

Bryoph. Bibl. 31:30 (1985) = C. schmidii C. 

Müll. 

calymperidictyon (Geh. & Hampe) Kindb., 

Enum. Bryin. Exot. 88 (1889). 

B: Dicranum calymperidictyon Geheeb & 

Hampe, Flora 64: 342 (1881). 

T: Brazil, Sao Paulo, Apiahy, Puiggari 250 

(isotypes FH, S).


Lich. 45:155 (1979). 

cambodgensis Thér., Rev. Bryol. n.ser. 2:17 

(1929). 

T: Cambodge, N de Kampot, rocher 800 m, 

Porlane s.n. (1928) (isotype BM). 



cambouei Ren. & Card. in Ren., Rev. de Bot. 

392 (1891). 

T: Madagaskar, près du lac d’Ambohipo, 

Camboue s.n. (holotype PC). 

campiadelphus (C. Müll.) Par. Ind. Bryol. 

ed.2,1:301 (1904). 

B: Dicranum campiadelphum C. Müll., 

Hedwigia 39: 252 (1900). 

CU: C. canaliculatus cf. Frahm, Revue Bryol. 

Lich. 45:140 (1979) = C. pyriformis (Schultz) 

Brid. 

campoanus Thér., Rev. Chil. Hist. Nat. 22:81 

(1918). 

T: Chile, Victoria, Campo s.n. (isotype H- 

BR). 

CU: C. modestus Card. cf. Frahm, Bryol. 

Beitr. 7: 20 (1987). 

canadensis Kindb., Rev. Bryol. 32:35 (1905). 


Loeske cf. Grout, Moss Fl. N. Am. 2:95 

(1937). 

canaliculatus (Geh. & Hampe) Kindb. Enum. 

Bryin. Exot. 88 (1898). 

B: Dicranum canaliculatum Geh. & Hampe, 

Vid. Medd. Naturh. Foer. Kjoebenh. ser. 4,1: 

92 (1879). 

T: Brasiia, prope Apiahy, Puiggari s.n. 

isotype PC). 

OU: C. pallidus cf. Frahm, Rev. Bryol. Lich. 

45:140 (1979). 


canariensis Schimp. in Geh. & Herz., Biblioth. 

Bot. 73:28 (1910) nom. nud. 

canescens (C. Müll.) Jaeg. Ber. S. Gall. Naturw. 

Ges. 1870-71:435 (1872). 

B: Dicranum canescens C. Müll. Bot. Zeit. 

17: 220 (1859). 

T: Falkland Inseln, Lechler 90 (isotype JE). 



canescens (C. Müll.) Jaeg. fo. perincanus (C. 

Müll.) J.-P. Frahm, Rev. Bryol. Lich. 42: 903 

29 

(1976). 

canescens Wils., Kew J. Bot. 9:298 (1857) nom. 

nud. 

capensis Broth., Nat. Pfl. ed.2,10:188 (1924). 

OU: C. chlorotrichus (C. Müll.) Par. cf. Sim, 


stenopelma (C. Müll.) Par. = C. controversus 

(Hampe) Jaeg. 

CU: C. thwaitesii (Mitt.) Jaeg. 

capensis Schimp. in Mitt., J. Linn. Soc. Bot. 12:64 

(1869) nom. nud. 

HS: capensis Schimp. in Par. Ind. Bryol. 242 

1894 nom. nud. 

M: „S. Afr., auf Sandboden Br(eutel) 

legit“(NY). 

OU: C. lepidophyllus (C. Müll.) Jaeg. cf. Par., 

Ind. Bryol. 242 (1894). 

CU: C. introflexus (Hedw.) Brid. 

capillaceus (Brid.) Jaeg. Ber. S. Gall. Naturw. 

Ges. 1870-71:429 (1872) hom. illeg. 

capillaceus Hook. f. & Wils. London J. Bot. 

3:543 (1844). (*) 

capillatus Hook.f. & Wils., Fl. Tasman. 2:172 

(1859). 

T: Tasmania, Weymouth s.n. (isotype S). 

OU: C. pallidus Hook.f. & Wils. cf. Sainsb., 

R. Soc. New Zeal. Bull. 5:116 (1955). 


capitiflorus Mont. Syll. 42 (1856). 

B: Dicranum capitiflorum C. Müll. Syn. 

Musc. 2: 598 (1851). 

T: Reunion, Ile Bourbon, Richard 682 


CU: C. arcuatus (Brid.) Jaeg. cf. Frahm, 

Bryoph. Bibl. 31:19 (1985). 

capitiflorus Mont. var. pachycomus Besch. Ann. 

Sci. Nat. Bot. ser. 6, 9: 322 (1880). 


Bryoph. Bibl. 31:19 (1985). 

capitulatus Bartr., Bull. Brit. Mus. (Nat. Hist.) 

Bot. 2: 52. (1955). 

T: Ecuador. Chaupi-Sagcha, Pululagua, on a 

stump, Bell 591 (holotype, FH; isotype BM). 

caracasanus (C. Müll.) Par. Ind. Bryol. 242 

(1894). 

T: Venezuela, Fendler 36 (holotype destroyed 

at B; lectotype NY; isotype S). 

OU: Thysanomitrion caracasanum C. Müll., 

Linnaea 42: 471 (1879). 

CU: C. richardii Brid. cf. Frahm, Revue

30 

Bryol. Lich. 42:603 (1976). 

carassensis (Broth.) Par., Ind. Bryol. 242. 1894. 

B: Thysanomitrion carassensis Broth., Act. 

Soc. Sc. Fenn. 19(5): 9 (1891). 

T: Brazil, Minas Gerais, Caraca, Wainio s. n. 

(holotype, H-BR; isotypes FH, JE, S). 

CU: C. cuspidatus (Hornsch.) Mitt. cf. Frahm, 


carbonicolus Thér., Rev. Chil. Hist. Nat. 22:81 

(1918). 

T: Chile, Victoria, Campo s.n. (isotype H- 

BR). 


Beitr. 7: 21 (1987). 

cardotii Thér., Rec. Publ. Soc. Havraise Etud. 

Div. 1922:116 (1922). 

T: Madagascar massif d’ Andringitra 1600- 

2500 m, Bathie 20 (holotype PC). 

OU: C. controversus (Hampe) Jaeg. cf. 

Frahm, Hikobia 11 (1994). 


carinatus C. Müll. Gen. Musc. Fr. 265 (1900) 

nom. nud. 

M: Argentinien, Barranca am Arroyo 

Cortapic, Lorentz (1881) (JE). 

CU: C. introflexus (Hedw.) Brid. var. 

cordobaensis Thér. cf. Frahm, Nova 

Hedwigia 30:843 (1978). 

carolinae Grout, Moss fl. N. Amer. 1: 249. 

(1939). 

T: U.S.A., North Carolina, Brunswick Co., 

near Southport, Anderson & Evans 6180 

(holotype DUKE). 

carreiroanus Card., Bull. Herb. Boiss. ser.2,5:202 

(1905). 

T: Azores, San Miguel, Carreiro 689 


CU: C. shawii Wils. cf. Frahm, Journ. Bryol. 

13:329 (1985). 

cataractarum (Fleisch.) Fleisch. Musci Fl. 

Buitenzorg 1:185 (1904). 

T: Java, am Wasserfall bei Tjiburnum 1800 

m, Fleischer 111 (isotype NY). 

OU: C. recurvus (Mitt.) Jaeg. ssp. 

cataractarum Fleischer, Musci Fl. Buitenzorg 

1:110 (1904), C. hemitrichius fide Eddy, 

Malesian Mosses vol. 1, 1988. 

CU: C. recurvus (Mitt.) Jaeg. cf. Frahm, 

Bryol. Beitr. 7: 22 (1987). 

catarractilis (C. Müll.) Par., Ind. Bryol. Suppl. 

90 (1900). 

B: Dicranum catarractile C. Müll., Hedwigia 

38: 80 (1899). 

T: South Africa, ad catharractam supra 

Rondebosch, Rehmann 64 (G, PC). 

catharinensis Par., Ind. Bryol. ed.2,1:302 (1904) 

nom. illeg. 

OU: C. perfalcatus Broth. 


caldensis (Aongstr.) J.-P.Frahm . 

catumbensis Broth., Bih. K. Svensk. Vet. Ak. 

Foerh. 21 afd. 3(3):8 (1895). 

T: Type. Brazil, Rio de Janeiro, Catumbez, 

Mosén 190 (holotype H-BR; isotype S). 

CU: C. surinamensis C. Müll. cf. Frahm, Revue 

Bryol. Lich. 45:163 (1979). 

caudatus (C. Müll.) Mont. in Dozy & Molk., 

Bryol. Jav. 1:78 (1858). 

B: Dicranum caudatum C. Müll. Syn. Musc. 

Frond. 1: 401 (1848). 

T: India, Nilgherri, Perrottet s.n. (lectotype 

H-BR). 

CU: C. comosus Bosch & Lac. cf. Frahm, 

Journ. Bryol. 13:166 (1984). 

caudatus C. Müll. nom. nud. 

CU: Chorisodontium sp. cf. Frahm, Nova 

Hedwigia (1994). 

cavernosus J.-P.Frahm, Nova Hedwigia 29:245 

(1978). 

T: Type: Brazil, Rio de Janeiro, Itatiaia Nat. 

Park, Agulhas Negras 2760 m, Vital 4921 

(holotype SP; isotypes B, U). 

CU: C. densicoma cf. Frahm, Nova Hedwigia 

39:155 (1984). 

cavifolius Mitt., J. Linn. Soc. Bot. 12: 87 (1869). 

T: Ecuador, „Andes Quitenses“, Chimborazo, 

Jameson s.n. (lectotype NY). 

celebesicus Card., Rev. Bryol. 28:113 (1911). 

T: Celebes, Buan Kräng, 5000 - 9000ped., 

Fruhstorfer 1895 (isotype FH). 

OU: C. zollingerianus (C. Müll.) Bosch & 

Lac. cf. Frahm, Bryol. Beitr. 7: 22 (1987). 


zollingerianus (C. Müll.) J.-P. Frahm, Trop. 

Bryol. 4: 61 (1991). 

cerradensis Vital, Cryptogamie Bryol. Lich. 5:20 

(1984). 

T: Brazil. Sao Paulo: Municipio de Itirapina, 

Vital 1714 (holotype, SP).

CU: C. carolinae Grout cf. Frahm , Nova 

Hedwigia 43 (1986). 

ceylonensis Broth. in Fleisch., Musci Fl. 

Buitenzorg 1:101 (1904) nom. nud. 

OU: C. zollingerianus (C. Müll.) Bosch. & 

Lac. cf. Fleischer, l.c. 

CU: C. fragilis (Brid.) B.S.G. ssp. goughii 

(Mitt.) J.-P. Frahm, Trop. Bryol. 4: 61 (1991). 

chalarobasens C. Müll. in Par., Ind. Bryol. 242 

(1894). (*) 

chalarobasis C. Müll. ex Thér., Rev. Bryol. Lich 

7:171 (1935). 

T: Chile, Corral, Quitaluto, Gunckel 1729 (S). 

CU: C. incrassatus C. Müll. cf. Frahm. Bryol. 

Beitr. 7: 22 (1987). 

chateauvertii Thér., Rev. Bryol. ser.4:76 (1931). 

T: Malawi, Mont Mlongo 2600 m, 

Chateauvert s.n. (holotype PC). 

CU: C. ampliretis cf. Frahm, Cryptogamie 

Bryol. Lich. 3(1):61 (1982) = C. flaccidus 

Ren. & Card. 

chevalieri Broth. & Thér., Bull. Mus. Hist. Nat. 

Paris 18: 476 (1912). 

T: Cote d’Ivoire, Aug. Chevalier s.n. (PC, 

S). 

chilensis De Not., Mem. R. Acc. Sc. Torino 

ser.2,18:453 (1859). 

T: Chile, Valparaiso, Puccio s.n. (isotype H- 

SOL). 

chionophilus (C. Müll.) Mitt. J. Linn. Soc. Bot. 

12:81 (1869). 

CU: Dicranodontium denudatum (Brid.) 

Britt. cf. Robinson, Bryologist 70:11 (1967). 

chlorophyllosus (C. Müll.) Jaeg. Ber. S. Gall. 

Naturw. Ges. 1870-71:428 (1872). 

B: Dicranum chlorophyllosum C. Müll. Syn. 

Musc. 1: 395 (1848). 

T: South Africa,“ Promont boni spei“, 

Guienzius s.n. ,1842 (could not be located). 

CU: C. bicolor (C. Müll.) Wils. ssp. atroluteus 

(C. Müll.) J.-P.Frahm cf. Frahm, Bryoph. 

Bibl. 31:37 (1985). 

chlorotrichus (C. Müll.) Par., Ind. Bryol. 

ed.2,1:303 (1904). 

T: Dicranum chlorotrichum C. Müll. 

Hedwigia 38: 87 (1899). 

T: Cape, Montagu Pass, Rehmann 53 

(isotypes PC, L). 

CU: C. stenopelma (C. Müll.) Par. cf. Magill, 

Fl. S. Afr. Bryoph. 1(1):141 (1981), C. 

31 

controversus (Hampe) Jaeg. cf. Frahm, 

Bryoph. Bibl. 31:180 (1985). 


chrismarii (C. Müll.) Mitt. J. Linn. Soc. Bot. 

12:88 (1869). 

B: Dicranum chrismarii C. Müll. Bot. Zeit 

13: 761 (1855). 

T: Mexico. Michoacan, Chrismar s. n. 

(holotype destroyed at B; lectotype NY; 

isolectotypes H-BR, S). 


Bryoph. Bibl. 31:124 (1985). 

chrismarii var. tolucensis (Bartr.) J.-P. Frahm, 

Bryologist 80: 120 (1977). 

B: C. tolucensis Bartr. 

CU: C. nivalis cf. Frahm, Fl. Neotr. 54 (1991). 

chrismarii var. suboblongus (Herz.) J.-P. Frahm, 

J. Hatt. Bot. Lab. 48:209 (1980) = C. 

nivalis (Brid.) Brid. cf. Frahm, Bryol. 

Beitr. 7: 23 (1987). 

chryseolus C. Müll. ex Besch., Ann. Sc. Nat. Bot. 

ser. 6,9:325 (1880). 

T: Comores, Boivin 18 (holotype PC, isotype 

G). 

HS: C. chrysocolus C. Müll. ex Besch., Rev. 

Bryol. 4:15 (1877). 

OU: C. aureus Bosch & Lac. cf. Frahm, 

Bryoph. Bibl. 31:30 (1985). 

CU: C. schmidii C. Müll. 

chrysodictyon (Hampe) Mitt., J. Linn. Soc. Bot. 

12:78 (1869). 

B: Dicranum chrysodictyon Hampe, Ann. Sc. 

Nat. Bot. ser. 5,5: 338 (1866). 

T: Type. Colombia, Bogotá, Tequendama, 

Lindig s. no. (holotype not seen; isotypes H- 

BR, H-SOL, GOET, NY, S) 

CU: C. pauper cf. Frahm, Bryol. Beitr. 7: 23 

(1987). 

cinchonae Par., Ind. Bryol. Suppl. 90 (1900). 

T: Jamaica, Wilson 813 (not seen). 

HS: Thysanomitrion jamaicense C. Müll., 

Bull. Herb. Boiss. 5:552 (1897). 

OU: C. introflexus (Hedw.) Brid. cf. Williams, 

N.Am. Fl. 15:144 (1913). 

CU: C. pilifer Brid. 

cirrhatus Brid., Bryol. Univ. 1:479 (1826). 

CU: Cynodontium gracilescens (Web. & 

Mohr) Schimp. cf. Limpr., Laubm. Deutschl. 

1:284 (1886). 

circinatus J.-P. Frahm, Cryptogamie Bryol.

32 

Lichènol. 19(1): 27, 1998. 

T: Australia, Victoria, Aireys Inlet, Distillery 

Creek, Fern Gully, on peaty ground in tree 

fern grove. Angahook Regeneration Study 

Group Quadrat M14, 24.10.93 (Holotypus 

MELU, isotypus hb. Frahm) 

citrescens Stirt., Trans. Bot. Soc. Edinburgh 

26:245 (1914). 

CU: C. fragilis (Brid.) B.S.G. cf. Dixon, J. 

Bot. 61:15 (1923). 

civa Lor., Moosstud. 159 (1864). 

T: „India orientalis“ (isotype S). 

CU: C. pilifer Brid. cf. Frahm, Cryptogamie 

Bryol. Lich. 3(1):60 (1982). 

clavatus (R. Brown) Wils. in Hook., Fl. Nov. 

Zeal. 2: 69 (1854). 

B: Dicranum clavatum R. Brown in 

Schwaegr., Spec. Musc. Suppl. 3 (2): 255a 

(1829). 

T: van Diemen Land, Brown & Sieber s.n. 

clavatus Rehm. in Sim, Trans. R. Soc. S. Afr. 

15:171 (1926) nom. nud. 

CU: introflexus (Hedw.) Brid. cf. Sim l.c. 

cleefii J.-P. Frahm, Cryptogamie Bryol. Lich. 7: 

441 (1986). 

T: Colombia. Boyaca: Paramo La Rusia, 

subparamo cerca del puente del Rio Surba, 

Cleef 6269 (holotype U). 

clemensiae Bartr. Rev. Bryol. Lichenol. 30: 187 

(1961). 

T: Papua N.G. Western Highlands, Wabag 

area, 10.000 ft. R.G. Robinson 3025a 


clementsii Dix., Ann. Bryol. 12:49 (1939). 

T: Sumatra, Gunung Goh Lemboek 3000 m, 

van Steenis 10185 (BM). 

CU: Atractylocarpus comosus Dix. cf. Frahm, 

Bryol. Beitr. 7: 24 (1987). 

cockyanii R. Brown ter., Trans. New Zeal. Inst. 

29:467 (1897). (*) 

collinus Par., Ind. Bryol. Suppl. 90 (1900). 

T: Brasilien, Sta. Catharina, Laguna, auf Hügeln 

im Campo, Ule 948 (isotype H-BR). 



comatulus Besch., Compt. Rend. Ac. Sc. Paris 

81:724 (1875). 

T: Ile de Amsterdam, sterile entre les touffes 

de lycopodium, G. d’Isle s.n. (holotype PC). 

CU: C. megalotus Besch. cf. Frahm, Bryoph. 

Bibl. 31: 118 (1985). 

comatus Ren. & Card. in Ren., Rev. de Bot. 9:290 

(1891). 

T: Madagascar, St. Marie, Arbogast (1890) 

s.n. (holotype PC, isotype G). 

CU: C. trachyblepharon ssp. comatus (Ren. 

& Card.) J.-P. Frahm, Bibl. Bot. 31: 187 

(1985). 

commersonii Besch. in Ren., Prodr. Fl. Bryol. 

Madag. 88 (1898). 

T: Reunion, Cratere Commerson, G. de I’Isle 

s.n. (holotype PC). 

OU: C. chrismarii (C. Müll.) Mitt. cf. Frahm, 

Lindbergia 8(1):46 (1982). 


comosus (Schwaegr.) Bosch & Lac., Bryol. Jav. 

1:75 (1858). 

B: Dicranum comosum Schwaegr. Spec. 

Musc. Suppl. 2(2): 114, tab. 185 (1827). 

T: Indonesia, Java, Reinwardt s.n. (holotype 

L). 

comosus var. compactus Fleisch. Nova Guinea 

8(2): 738 (1912). 

T: Indonesia,West Irian, Jayawijaya, am Gipfel 

d. Hellwig-Geb. 2500-3000 m, Römer 

1309 (holotype FH). CU: C. comosus 

(Schwaegr.) Bosch & Sande Lac. 

compactus Par. & Broth. Rev. Bryol. 31:117 

(1904) hom. illeg. 

T: Guinea, in valle fl. Bating, Pobeguin 

(1906) s.n. (holotype PC). 



(1982). 

compactus Schimp. in Hunt, Mem. Lit. Phil. Soc. 

Manchester ser. 3,3:234 (1868) nom. illeg. 

CU: C. schimperi Milde. 

compactus Wils., Kew J. Bot. 9:297 (1857) nom. 

nud. 

OU: C. goughii (Mitt.) Jaeg. cf. Mitt., J. Linn. 

Soc. Bot. Suppl. 1:17 (1859). 


(Mitt.) J.-P. Frahm, Trop. Bryol. 4: 61, 1991. 

concolor (Hook.) Brid., Bryol. Univ. 1: 476 

(1826). 

B: Dicranum concolor Hook., Musci Exot. 

139. (1816). 

T: Colombia, Alto de Aranda, Humboldt & 

Bonpland s. n. (isotypes H-BR, KIEL). 

connivens Broth. in Bruehl, Rec. Bot. Surv. India

13(1):123 (1931) nom. nud. 

consociatus Thér. Rec. Publ. Soc. Havraise Etud. 

Div. 2: 118 1922 

B: Thysanomitrion consociatum Broth Nat. 

Pfl. ed. 2, 10. 189 1924 

T: Madagascar. Massif d’ Adringita 1600- 

2500 m, Perrier de la Bathie s.n. (Holotypus 

H-BR) 

CU: C. flaccidus Ren. & Card. cf. Frahm, 

Nova Hedwigia 39 592 (1984). 

contortus (Wahlenb.) Brid., Mant. Musc. 74 

(1819) 

CU: Grimmia incurva Schwaegr. cf. C. Müll., 

Syn. Musc. 1:789 (1849). 

controversus (Hampe) Jaeg., Ber. St. Gall. 

Naturw. Ges. 1877-78: 385. 1880. 

B: Dicranum controversum Hampe, Vid. 

Medd. Naturh. For. Kjoebenh. ser. 3,4: 42. 

1872. 

T: Brazil, Glaziou 5204 (holotype BM, 

isotypes L, NY). 

CU: C. thwaitesii (Mitt.) Jaeg. cf. Frahm, 

Crypt. Bryol. Lichènol. 19: 32, 1998. 

convolutus Aongstr. in Kindb., Enum. Musc. 

Exot. 9 (1888) nom. nud. 

copelandii Broth., Leafl. Philip. Bot. 6:1973 

(1913). 

T: Philippines, Luzon, Mt. Banahao, 

Copeland 830 (holotype H-BR). 

OU: C. hemitrichius (C. Müll.) Jaeg. cf. Bartr., 

Philipp. J. Sc. 68:41 (1939). 

CU: C. schmidii C. Müll. ssp. hemitrichius 

(C. Müll.) J.-P. Frahm. 

coreensis Card., Bull. Herb. Boiss. ser. 2,7:715 

(1907). 

T: Corée, Fusan, Faurie 265 (holotype PC, 

isotypes NY, S). 


Nova Hedwigia 39: 600 (1984). 

corralensis Broth. in Par., Ind. Bryol. ed.2,1:304 

(1904) nom. nud. 

CU: C. clavatus (R.Brown) Wils. fc. Sainsb., 

R.Soc. New Zeal. Bull. 5:110 (1955). 

costaricensis Bartr., Contr. U.S. Nat. Herb. 26: 

62 (1928). 

T: Costa Rica, Finca Montechristo, Standley 

& Valerio 48579 (holotype FH; isotype US). 

OU: C. oerstedianus cf. Frahm, Lindbergia 

7:28 (1981). 

CU: C. surinamensis fide Allen, Trop. Bryol. 

33 

1: 91 (1989). 

crassinervis Wils., Kew J. Bot. 9:297 (1857) nom. 

nud. 

CU: Paraleucobryum enerve (Thed.) Loeske 

cf. Podpera, Consp. 155 (1954). 

crassissimus Besch., Bull. Soc. Bot. France 

32:LVIII (1885). 

T: Chile, Puerto Bueno, Savatier s.n. 

(holotype PC, isotype NY). 


Beitr. 7: 25 (1987). 

cratericola Par., Ind. Bryol. Suppl. 90 (1900) 

nom. nud. 

crateris Besch., Ann. Sc. Nat. Bot. ser. 6, 9: 327 

(1880). 

T: Reunion. cratere Commerson, Lepervanche 


OU: Thysanomitrion crateris (Besch.) Broth. 

in Card. in Grand Hist. Madag. 39: 130 

(1915). 

crenulatus Stirt., Trans. Bot. Soc. Edinburgh 

26:244 (1914). 

CU: C. flexuosus (Hedw.) Brid. cf. Dixon, J. 

Bot. 61:15 (1923). 

cribrosus (Brid.) Brid., Mant. Musc. 76 (1819). 

CU: Grimmia pulvinata (Hedw.) Sm. var. 

africana (Hedw.) Hook.f. & Wils. cf. Par., 

Ind. Bryol. ed.2,2:222 (1904). 

crishna Lor., Moosstud. 159 (1864). 

T: „India orientalis“ (S). 

CU: C. pilifer Brid. cf. Frahm, Cryptogamie 

Bryol. Lich. 3(1):60 (1982). 

crispatulus Thér., Rec. Publ. Soc. Havraise Etud. 

Div. 1925:129 (1926). (*) 

crispatus (C. Müll.) Broth. in Par., Ind. Bryol. 

Suppl. 91 (1900). 

B: Dicranum crispatum C. Müll., Bull. Herb. 

Boiss. 6: 36 (1898). 

T: Brazil, Ule 1799 (holotype destroyed at 

B; lectotype, H-BR). 


caldensis (Aongstr.) J.-P.Frahm cf. Frahm, 

Revue Bryol. Lich, 45(2):138 (1979). 

crispicoma (C. Müll.) Jaeg. Ber. S. Gall. Naturw. 

Ges. 1877-78:383 (1880). 

B: Dicranum crispicoma C. Müll., Flora 34: 

529 (1875). 

T: Venezuela, prope Caracas, Galipan 4500 - 

5000 ft., Funck & Schlim 366 (holotype 

destroyed at B; lectotype NY).

34 



crispifolius Bartr., Contr. U.S. Nat. Herb. 37:45 

(1965). 

T: Papua New Guinea. Simbu, east slopes, 

Mt. Wilhelm,3560 m. Brass 29894 (holotype 

FH). 

OU: C. boswellii (Hampe) Par. cf. Frahm, 

Bryologist 94: 62 (1991). 


zollingerianus (C. Müll.) J.-P. Frahm cf. 

Frahm, Trop. Bryol. 4: 61 (1991). 

crispo-filifolius Par., Ind. Bryol. Suppl. 90 (1900) 

nom. nud. 

M: Sumatra occid., in regiona lacus Toba, 

Modigliani 241 (FH, H-BR, JE, S). 

O: C. walkeri Mitt. cf. Frahm, J. Bryol. 

13:189 (1984) = Bryohumbertia walkeri 

(Mitt.) J.-P.Frahm. 

CU: Bryohumbertia subcomosa (Dix.) J.-P. 

Frahm cf. Frahm, Trop. Bryol. 1:9 (1989). 

cruegeri (C. Müll.) Par., Ind. Bryol. Suppl. 91 

(1900). (*) 

cryptopodioides Broth., Bih. K. Svensk Vet. 

Akad. Foerh. 26 Afd. III(7): 9 (1900). 

T: Brazil, Mato Grosso, Lindman 475 

(holotype H-BR; isotype S). 

cuatrecasii Robinson, Bryologist 70: 16 (1967). 

T: Colombia, Cuatrecasas 19090a (holotype 

US; isotype COLO). 

CU: C. trivialis Thér. cf. Frahm, Fl. Neotr. 

54: 187 (1991). 

cubensis Sull., Proc. Am. Arts sc. 5: 278 (1861). 

T: Cuba, Wright 39 (holotype NY). 

cubensis Sull. var. exaltatus (C. Müll.) J.-P. 

Frahm, Nova Hedwigia 41: 274 (1985). 

CU: C. lamellinervis var. exaltatus. 

cucullatifolius Herz., Bibl. Bot. 87:22 (1916). 

T: Bolivia, Tablas 3400 m, Herzog 2826/a 

(holotype JE). 

CU: C. cuspidatus (Hornsch.) Mitt. var. 

dicnemioides (C. Müll.) J.-P. Frahm cf. 


cummingii Duby, Mem. Soc. Phys. Hist. Nat. 

Geneve 20:361 (1869). 

T: Chile, Valdivia (holotype G, isotype S). 


Cryptogamie Bryol. Lich. 3:62 (1982). 

curvatifolius Card., Bull. Herb. Boiss. 

ser.2,5:1000 (1905). 

T: Iles Falkland, Port Standley, Birger 208 


CU: ? C. acuminatus Mitt. cf. Frahm, Bryol. 

Beitr. 7: 26 (1987). 

curvifolius Brid., Mant. Musc. 78 (1819). 

CU: Grimmia ovalis (Hedw.) Lindb. cf. C. 

Müll., Syn. 1:810 (1849). 

cuspidatus (Hornsch.) Mitt., J. Linn. Soc. Bot. 

12: 90 (1869). 

B: Dicranum cuspidatum Hornsch., Fl. Bras. 

13. 1840. 

T: Brazil, Sincora et Rio das Contas, Martius 

s. n. (holotype destroyed at B; lectotype BM; 

isolectotypes JE, NY). 

cuspidatus (Hornschuch), Mitten var. 

dicnemioides (C. Müll.) Frahm, Rev. Bryol. 

Lichenol. 45(2): 143 (1979). B: Dicranum 

dicnemioides C.Müll., Bull. Herb. Boiss. 6: 

36 (1898). 

T: Brazil, Serra do Itatiaia, Agulhas Negras, 

Ule 1788 (H-BR). 

cygneus (Hedwig) Brid., Mant. Musc. 72 (1819). 

B: Dicranum cygneum Hedwig, Spec. Musc. 

148. 1801. 

T: „India occ.“, Swartz s. n. (holotype G; 

isotypes H-SOL, NY, S). 

cylindrotheca R.Brown ter., Trans. New Zeal. 

Inst. 29:473 (1892). 


New Zeal. Inst. Bull. 3(3):88 (1923). 

D 

damazii Broth. Denkschr. Ak. Wiss. Wien Math. 

Nat. Kl. 83:260 (1926). 

T: Brazil, Minas Gerais, Pico da Serra de 

Itabira, Damaziou 2147 (holotype H-BR). 

OU: C. praealtus var. damazii (Broth.) J.- 

P.Frahm cf. Frahm, Revue Bryol. Lich. 

45:159 (1979). 

CU: C. subcuspidatus var. damazii cf. Frahm, 

Bryol. Beitr. 7:27 (1987). 

davalianus Watts. in Dix., J. Bot. 60:289 (1922) 

nom. nud. 

OU: C. ericeticola C. Müll. cf. Dixon, l.c. 

CU: C. bicolor (C. Müll.) Wils. var. 

ericeticola (C. Müll.) Dix. 

decaryii Thér., Rec. Publ. Soc. Havraese Etud. 

Div. 1923(2): 78 (1923). 

T: Madagascar, Moramanga, sur la terre humide, 

Decary s.n. (holotype PC).

deciduus Ren. & Card., Bull. Soc. Bot. R. Bot. 

Belg. 33(2): 111 (1895). 

T: Madagascar, Ambohimatsara, Betsilio 

Berthieu 1892 (holotype PC, isotype S). 

CU: C. aureus Bosch. & Lac. cf. Frahm, 

Bryoph. Bibl. 31:30 (1985). 

decipiens Lac., Ned. Kruidk. Arch. 5:294 (1870). 

CU: C. brevipilus B.S.G. cf. Milde, Bryologia 

silesiaca 78 (1869). 

declivium Par., Ind. Bryol. ed.2,1:305 (1904). 

OU: C. nano-filifolius (C. Müll.) Par., Ind. 

Bryol. Suppl. 94 (1900), C. filifolius 

(Hornsch.) Mitt. cf. Frahm, Bryoph. Bibl. 

5:42 (1975). 


P.Frahm cf. Frahm, Bryol. Beitr. 7:27 (1987). 

delagoae (C. Müll.) Par. Ind. Bryol. Suppl. 91 

(1900). 

B: Dicranum delagoae C. Müll. Hedwigia 38: 

86 (1899). 

T: Transvaal, between Delagoa Bay and 

Lydenburg, Wilms (1884) s.n. (holotype B). 



OU: Thysanomitrion delagoae (C. Müll.) 

Broth. Nat. pfl. 10: 188 (1924); C. stenopelma 

(C. Müll.) Par. cf. Sim, Trans. R. Soc. S. Afr. 

15:114 (1926). 

delicatulus Williams, N.Am. Fl. 15:137 (1913). 

T: Cuba,. Herradura,. Britton 6523 (holotype 

NY; isotype PC). 

CU: C. angustiretis (Aust.) Lesq. & Jam. cf. 


demangei Thér. & P.Varde, Rev. Bryol. 49:28 

(1922). 

T: Tonkin, Yen Lap prè s de Quang Yen, 

Demange 529 (holotype PC). 

CU: C. serratus Lac. cf. Frahm, Bryol. Beitr. 

7: 27 (1987). 

OU: C. aureus cf. Eddy, Malesian Mosses vol. 

1 (1988). 

densicoma (C. Müll.) Par., Ind. Bryol. Suppl. 91. 

(1900). 

B: Dicranum densicoma C. Müll., Nuovo 

Giorn. Bot. Ital. n. ser. 4: 33 (1897). 

T: Bolivia, Prov. Cochabamba, Germain 1120 

(holotype destroyed at B; lectotype NY; 

isolectotype H-BR). 

densicoma (C. Müll.) Par. var. yungarum (Herzog) 

J.-P. Frahm ,Nova Hedwigia 39: 156 

35 

(1984). 

T: Bolivia, Incacorral 2200 m, Herzog 1908 

(holtype JE isotype B). 

CU: C. densicoma 

densifolius Aongstr., Oefv. K. Svensk Vet. Ak. 

Foerh. 29(4):18 (1872). 

T: Hawaii, Honolulu, N.J. Anderson s.n. 

(isotypes H-BR, L). 

CU: C. hawaiicus (C. Müll.) Jaeg. var. 

densifolius (Aongstr.) J.-P. Frahm cf. Frahm, 

The Bryologist 94: 62 (1991). 

densifolius Aongstr. var. falcatus Broth., Bishop 

Mus. Bull. 40:6 (1927). 

CU: C. pupureo-flavescens (C. Müll.) Par. cf. 

Bartr., Bishop Mus. Oc. pap. 16:323 (1942). 

densinervis Broth. in Williams, Bull. New York 

Bot. Garden 8:335 (1914) nom. nud. 

densiretis Broth., Bull. Soc. Bot. Ital. 1904:15 

(1904) nom. nud. 

OU: C. densifolius Aongstr. cf. Broth., Bishop 

Mus. Bull. 40:36 (1927). 




densus Schimp. in Jaeg., Ber. S. Gall. Naturw. 

Ges. 1877-78: 386 (1880). 

T: Abyssinia, Schimper s.n. (PC) 

CU: C. fragilis 

densus B.S.G. Bryol. Eur. 1:166 (1847). 

CU: C. fragilis (Brid.) B.S.G. fo. densus cf. 

Moenkemeyer, Laubm. Eur. 222 (1927). 

dentato-acicularis C. Müll. Gen. Musc. Fr. 270 

(1900) nom. nud. 


Lich. 42:901 (1976). 

M: Brazil, Jaguarone, Ule 708 (H-BR). 

denticuspis Broth. in Mildbr., Wiss. Erg. Deutsch. 

Zentr. Afr. Exp. 2:140 (1910). 

T: Zaire Karisimbi, Gipfelkegel 4000 m, 

Mildbraed 2091 (holotype H-BR). 

OU: C. stramineus (Mitt.) Jaeg. cf. Bizot & 

Kilbertus, Revue Bryol. Lich. 45:94 (1979). 


denticuspis Broth., Proc. Linn. Soc. N.S. Wales 

41: 575 (1916). 

T: Australia, N.S.W., Heath, Wardell, 

Richmond, Watts 5260 (holotype H-BR). 

OU: C. kirkii Mitt. cf. Frahm, J. Bryol. 13: 

365 (1985). 

CU: C. acuminatus Mitt var. kirkii (Mitt.)

36 

J.-P. Frahm, Bryol. Beitr. 7: 95 (1987). 

denticuspis Broth. var. lutescens Broth. Proc. 

Linn. Soc. N.S. Wales 41: 575 (1916). 

T: Australia, N.S.W., Heath, Wardell, 

Richmond, Watts 1739, 4758 (isotypes H- 

BR). 

CU: C. acuminatus Mitt var. kirkii (Mitt.) J.-P. 


denudatus (Brid.) Kindb., Bih. K. Svensk Vet. 

Ak. Handl. 7(9):87 (1883). 


Britt. 

depilis C. Müll. Gen. Musc. Fr. 276 (1900) nom. 

nud. 

destructilis (C. Müll.) Jaeg. Ber. S. Gall. Naturw. 

Ges. 1870-71:430 (1872). 

B: Dicranum destructile C. Müll., Bot. Zeit. 

17: 220. 1859. 

T: Mexico, Schiede s. no. (holotype destroyed 

at B, isotypes could not be located). 

OU: C. chrismarii (C. Müll.) Mit. cf. 

Williams, N.Am. Fl. 15:136 (1913). 


detonsus (Hampe) Kindb., Enum. Bryin. Exot. 

88 (1894). 

B: Dicranum detonsum Hampe, Flora 64: 341 

(1881). 

T: Brazil. Rio de Janeiro, Glaziou 11746 

(isotypes H-BR, JE, S). 

CU: C. trachyblepharon (C. Müll.) Mitt. cf. 

Frahm, Revue Bryol. Lich. 45:164 (1979). 

dichrostis (C. Müll.) Par. in Broth., Nat. Pfl. 1(3): 

333. (1901). 

B: Dicranum dichrostis C. Müll., Hedwigia 

39: 265. 1900. 

T: Brazil, Minas Gerais, Ouro Preto, Ule 1359 

(holotype destroyed at B; lectotype H-BR). 

dickieanus Stirt., Ann. Scott. Nat. Hist. 15 

(58):108 (1906). (*) 

dicksoniae C. Müll., Gen. Musc. Fr. 270 (1900) 

nom. nud. 

M: Brazil, S. Catarina Ule 660,1125 (H-BR). 

OU: C. cacti (C. Müll.) Par. Ind. Bryol. Lich. 

42:902 (1976). 

CU: C. occultus Mitt. 

dicnemioides (C. Müll.) Par., Ind. Bryol. Suppl. 

91 (1900). 

B: Dicranum dicnemioides C. Müll., Bull. 

Herb. Boiss. 6: 36 (1898). 

T: Brazil. Serra do Itatiaya, Agulhas Negras, 

Ule 788 (holotype destroyed at B; lectotype 

H-BR). 

CU: C. cuspidatus var. dicnemioides (C. 

Müll.) J.-P.Frahm, Herzogia 5:108 (1979). 

dicranelloides Ren. & Card., Bull. Soc. R. Bot. 

Belg. 29(1):171 (1890). 

CU: C. perpusillus Mitt. 

OU: Microcampylopus perpusillus (Mitt.) 

Broth. cf. Sim, Trans. R. Soc. S. Afr.15:181 

(1926). 

dicranodontioides Broth. in Skottsb., Nat. Hist. 

Juan Fernandez 2:242 (1924). 

OU: C. hygrophilus Broth. cf. Frahm, Nova 

Hedwigia 34:392 (1981). 

CU: C. vesticaulis Mitt. cf. Frahm, Bryol. 

Beitr. 7: 28 (1987). 

dicranoides Thér. & Nav., Bull. Soc. R. Bot. 

Belg. 60: 15 (1927). 

T: Ruwenzori, Butagu 3000 m, sur tronc 

tombe, Bequaert 4017 (holotype PC, isotype 

BR). 

didictyon (Mitt.) Jaeg., Ber. S. Gall. Naturw. Ges. 

1870-71:434 (1872). 

B: Dicranum didictyon Mitt., J. Linn. Soc. 

Bot. Suppl. 1: 21 (1859). 

CU: Dicranodontium didictyon (Mitt.) Jaeg. 

didrichsenii (C. Müll.) Jaeg., Ber. S. Gall. 

Naturw. Ges. 1870-71:424 (1872). 

B: Dicranum didrichsenii C. Müll., Bot. Zeit. 

20: 329 (1862). 

T: Hawaii, without locality, Didrichsen s.n. 

in expeditione transatl. danica not be located). 

CU: C. richardii Brid. cf. Dixon, J. Bot. 

60:287 (1922). 

OU: C. umbellatus (Arn.) Par. cf. Bartr., 

Bishop Mus. 101:44 (1933). 

didymodon (Griff.) Jaeg. Ber. S. Gall. Naturw. 

Ges. 1870-71:434 (1872). 

B: Dicranum didymodon Griff., Calcutta J. 

Nat. Hist. 2: 499 (1842). 

CU: Dicranodontium didymodon (Griff.) Par. 

dietrichiae (C. Müll.) Jaeg. Ber. S. Gall. Naturw. 

Ges. 1877-78:383 (1880). (*) 

B: Dicranum dietrichiae C. Müll., J. Mus. 

Godeffroy 3: 62 (1874). 

discriminatus (Hampe) Jaeg. Ber. S. Gall. 

Naturw. Ges. 1877-78:497 (1880). 

B: Dicranum discriminatum Hampe, Vid. 

Medd. Naturh. Foer. Kjoebenh. ser. 3, 9-10: 

254 (1878).


(isolectotype H-BR). 



(1981). 

dissitus C. Müll. ex Broth., Bot. Jahrb. 24:234 

(1897). 

T: Kamerun, Buea, Dusen s.n. (holotype B, 




distractus (C. Müll.) C. Müll., Abh. Naturw. Ver. 

Bremen 16(3):495 (1900). 

B: Dicranum distractum C. Müll., Hedwigia 

36: 350 (1897). 

T: cannot be located. 

OU: C. capillaceus Hook.f. & Wils. cf. Dixon, 

New Zealand Inst. Bull. 3(3):85 (1923) 

CU: C. lonchochaete C. Müll. 

ditrichoides Broth., Act. Soc. Sc. Fenn. 19(5):7 

(1891). 

T: Type. Brazil. Minas Gerais, Wainio s.n. 

(holotype H-BR; isotype S). 

CU: C. angustiretis (Sull.) Lesq. & Jam. cf. 


ditrichoides var.robustior Broth., Bih. K. Vet. Ak. 

Handl. 21 afd. III (3):6 (1895). 

T: Brazil, Glaziou 11761 (holotype, H-BR). 

CU: C. angustiretis (Sull.) Lesq. & Jam. cf. 


divaricatus (Mitt.) Jaeg. Ber. S. Gall. Naturw. 

Ges. 1877-78:381 (1880). 

B: Dicranum divaricatum Mitt. J. Linn. Soc. 

Bot. 7:149 (1863). 

T: Princess Island, Niger River, Barter 1923 

(isotype PC , holotype NY ). 



(1982). 

diversifolius (Leroy & Thér.) Takaki, J. Hatt. Bot. 

Lab. 37:516 (1973). 

T: Ruanda-Urundi, Escarpements à 

Usambura 1000m, Cornelis 21 (PC). 

B: Campylopodiella diversifolia Leroy & 

Thér., Bull. Jard. Bot. Bruxelles 18:200 

(1947). 

CU: C. obrutus Thér. & P. Varde cf. Frahm, 

Bryol. Beitr. 7: 29 (1987). 

diversinervis Broth., Philipp. J. Sc. C. 5:140 

(1910). 

37 

T: Philippines, Luzon, Mt. Pulog, Curzon 

16423 (holotype H-BR). 

CU: C. hemitrichius (C. Müll.) Jaeg. cf. 


divisus (Geh. & Hampe) Kindb., Enum. Bryin. 

Exot. 88 (1889). 

B: Dicranum divisum Geh. & Hampe, Flora 

64: 345 (1881). 

T: Brazil, Apiahy, Puiggari 323 (isotypes H- 

BR, S). 


Lich. 45:155 (1979). 

dixonii Luis., Broteria ser. Bot. 24:18 (1930). 

T: Madeira, Rio Arieiro, Barreto s.n. (isotype 

BM). 

CU: eximius Reichdt. cf. Frahm, Bryoph. 

Bibl. 31:67 (1985). 

OU: C. introflexus (Hedw.) Brid. cf. Sim, 

Trans. R. Soc. S. Afr. 15:174 (1926); C. 

bartramiaceus (C. Müll.) Thér. cf. Persson, 

Bot. Not. 1939:570 (1939), C. atroluteus (C. 

Müll.) Par. pp. 

doliolidium C. Müll. in Par., Ind. Bryol. Suppl. 

91 (1900) nom. nud. 

M: Madagascar orient., Tenerive distr. 

Taniatave, Perrot 1568 (S). 

OU: C. inerangae (C. Müll.) Par. cf. Sim, 


inchangae (C. Müll.) Par. cf. Magill, Fl. S. 

Afr. Bryoph. 1(1):146 (1981); C. rigens Ren. 

& Card. cf. Renault in Levier, Revue Bryol. 

28:89 (1901).. 

CU: C. robillardei Besch. cf. renauld in 

levier, Rev. Bryol. 28: 89 (1901). 

dolosus Besch., Ann. Sc. Nat. Bot. ser. 6,9:320 

(1880). 

T: Reunion, dísle 211/G (holotype PC). 

CU: Bryohumbertia spec. cf. Frahm, Bryoph. 

Bibl. 31:196 (1985). 

donnellii (Aust.) Lesq. & Jam., Man. Moss. N. 

Am. 79 (1884). 

B: Dicranum donnellii Austin, Bot. Gaz. 4: 

150 (1879). 

T: Florida, Smith, Musci Appal. 470 



Bryologist 83:581 (1980). 

OU: C. zygodonticarpus (C. Müll.) Par. cf. 

Allen, Trop. Bryol. 1: 92 (1989); C. flexuosus 

(Hedw.) Brid. cf. Bartram, Fieldiana Bot.

38 

25:43 (1949). 

dozyanus (C. Müll.) Jaeg., Ber. S. Gall. Naturw. 

Ges. 1870-71:418 (1872). 

HS: C. dozyi C. Müll. ex Kindb., Enum. 

Bryin. Exot 50 (1888). 

B: Dicranum dozyanum C. Müll. Syn. Musc. 

1: 385 (1848). 

T: Java, Korthals s.n. (PC). 



OU: C. richardii Brid. cf. Dixon, J. Bot. 

60:287 (1922). 

drouhardii Card. & Copp. in Grand Hist. Madag. 

39:114 (1915). 

T: Madagascar, Diego Suarez, Drouhard s.n. 


CU: C. cf. aureo-nitens (C. Müll.) Jaeg. cf. 

Frahm. Bryoph. Bibl. 31:24 (1985). 

durelii Broth. ex Gangulee, Nova Hedwigia 8: 

146, 1964, nom. invalid. 

dusenii C. Müll. in Par., Ind. Bryol. 246 (1894) 

nom. nud. 

M: Kamerun: ad Victoriam emporium in 

saxis, Dusen 304 (BM,PC,S). 



(1982). 

E 

eberhardtii Par., Rev. Bryol. 35:43 (1908). 

T: China, Tonkin, Eberhardt (1907) s.n. 


CU: C. japonicus Broth. cf. Frahm, The 

Bryologist 96: 142 (1993). 

echernieri Besch., Ann. Sc. Nat. Bot. ser. 6,9:323 

(1880). 

T: La Reunion, Echernier s.n. (isotype NY). 

CU: C. jamesonii (Hook.) Jaeg. cf. Frahm, 

Bryoph. Bibl. 31:102 (1985). 

echinatus Rehm. ex Sim, Trans. R. Soc. S. Africa 

15:175 (1926) nom. illeg. incl. vars. 

brevipilus Rehm. in Dix. & Gepp, leucobasis 

(Par.) Par., turgescens Rehm. in Par., 

umbrosus Rehm. in Par., vallis gratiae Rehm. 

in Par. 

T: Cape, Rehmann 67,68 (isotype NY). 

HS: C. echinatus Sim Bryo. S. Afr. 175 

(1926). 

CU: C. introflexus (Hedw.) Brid. cf. Dixon, 


eckendorfii Thér. & P. Varde Rev. Bryol. Lich. 

11:168 (1939). 

T: Zentralafrikanische Republik, Bubangui, 

Eckendorff 1780 (holotype PC). 


31:143 (1985). 

edithae Broth., Bot. Jahrb. 49: 175. (1912). 

T: Bolivia, Cordillera Real, Knoche 43 

(holotype H-BR, isotype B). 

OU: C. subjugorum Broth. var. edithae 

(Broth.) Thér. 

edwardsii Sim, Trans. R. Soc. S. Afr. 15:168 

(1926). 

T: Transvaal, Johannesburg 6000 ft., Edwards 

9836 (PC). 


Bryoph. Bibl. 31:71 (1985). 

ekmannii Thér., Mem. Soc.Cubana Hist. Nat. 

13:216 (1939). 

T: Cuba. Sierra Maestra, La Gran Piedra, 

Ekman 1703 (lectotype H-BR, NY, S). 

CU: C. setaceus Card. cf. Frahm, 


OU: C. cygneus cf. Frahm, Nova Hedwigia 

41:277 (1985). 

elliottii Bartr., Bull. Brit. Mus. Hist. Bot. 2:38 

(1955). 

T: Dominica, summit of Mt. Trois Pitons, 

Elliott 478a (holotype FH, isotype BM). 

CU: C. underwoodii Williams cf. Frahm, 


OU: C. cygneus cf. Frahm, Nova Hedwigia 

41:277 (1985). 

ellipticothecium R. Brown ter., Trans. New Zeal. 

Inst. 29:473 (1897). 


Trans. New Zeal. Inst. 59(5):506 (1928). 

ellipticus (Turn.) Brid., Mant. Musc. 76 (1819) 

CU: Racomitrium ellipticum (Turn) B.S.G. 

elongatus Brid. Bryol. Univ. 4:815 (1827). 

T: „Grateloup, Dax“ ex hb. Candolle 

(Holotypus B, Isotypus G). 

CU: Pilopogon guadeloupensis cf. Frahm, 

Crypt. Bryol. Lichènol. 19: 30, 1998. 

erectus (C. Müll.) Mitt., J. Linn. Soc. Bot. 12: 

89. 1869. 

B: Dicranum erectum C. Müll., Syn. 1: 408. 

1849. 

T: Colombia, Galipan, Moritz 145. 

CU: C. pilifer Brid.

ericeticola C. Müll., Abh. Naturw. Ver. Bremen 

16(3):496 (1900) 

T: New Zealand, Chatham Islands, Heide 27 

(H-BR). 

CU: C. bicolor (C. Müll.) Wils. var. 

ericeticola (C. Müll.) Dix. 

ericetorum (Mitt.) Jaeg., Ber. S. Gall. Naturw. 

Ges. 1877-78:386 (1880). 

B: Dicranum ericetorum Mitt., J. Linn. Soc. 

Bot. Suppl. 1: 20 (1859). 

OU: C. introflexus (Hedw.) Brid. cf. Mitten, 

J. Linn. Soc. Bot. 12:84 (1869). 

CU: C. pilifer Brid. cf. Frahm, Journ. Bryol. 

13:179 (1984). 

ericetorum Hampe in C. Müll., Gen. Musc. Fr. 

271 (1900) nom. nud. 

HS: C. nano-exasperatus C. Müll. nom. nud. 

ericoides (Griff.) Jaeg., Ber. S. Gall. Naturw. 

Ges. 1870-71:424 (1872). 

B: Dicranum ericoides Griff., Calcutta J. Nat. 

Hist. 2: 499 (1842). 

T: „in monte Khasia, in sylvis Myrung, 

Griffith 65 (lectotype BM). 

erythrocaulon Broth. Bol. Soc. Broteria 8:174 

(1890). 

T: Insula S. Thome, Quintas s.nr. (holotype 

B, isotypes H-BR,S,PC). 



erythrodontius Jaeg., Ber. S. Gall. Naturw. Ges. 

1877-78:496 (1880). 

T: Brazil, Rio de Janeiro, Glaziou 7371 (H- 

BR, NY). 


Bryol. Beitr. 7: 32 (1987). 

erythrognaphalus (C. Müll.) Jaeg., Ber. S. Gall. 

Naturw. Ges. 1870-71:439 (1872). 

B: Dicranum erythrognaphalon C. Müll., Bot. 

Zeit. 11: 37 (1853). 

T: India, Montes Neilgherienses, „leg. Berth. 

Schmid“ (PC). 

OU: C. involutus cf. Frahm, Bryol. Beitr. 7: 

32 (1987). 

CU: C. ericoides (Griff.) Jaeg. 

erythropoma Duby, Mem. Soc. Phys. Hist. Nat. 

Geneve 20:360 (1869). 

T: Australia, Mt. Mandon, Müller (1867) s.n. 

(holotype G, isotype S). 


Cryptogamie... 

39 

eunanus (C. Müll.) Par., Ind. Bryol. Suppl. 91 

(1900). 

B. Dicranum eunanum C. Müll., Hedwigia 

36: 348 (1897). 

T: Nouvelle Caledonie, près de Ile de 

Noumea, Balansa s.n. (PC). 

CU: Campylopodium medium (Duby) Giese 

& J.-P.Frahm cf. Frah, Bryol. Beitr. 7: 88 

(1987). 

euphorocladus Bosch. & Lac., Bryol. Jav. 1:79 

(1858). 

T: Java, Zollinger 411 Z (BM, MY). 

OU: Campylopodium euphorocladum (C. 

Müll.) Besch. 


& Frahm. 

eurydictyon Dix. ex Bartr., Philipp. J. Sc. 68:42 

(1939). 

T: Philippines, Luzon, Mt. Maquiling, 

Kerklots P136 (holotype BM). 

CU: C. laxitextus Lac. cf. Frahm, Bryol. Beitr. 

7: 32 (1987). 

exalaris (Hampe) Par., Ind. Bryol. 246 (1894). 

B: Dicranum exalare Hampe, Vid. Medd. 

Naturh. For. Kjoebenh. ser. 3: 257 (1878). 

T: „Brasilia australis“, Glaziou 7473 (H-BR). 



exaltatus (C. Müll.) Par., Ind. Bryol. 246 (1894). 

B: Dicranum exaltatum C. Müll., Linnaea 42: 

472 (1879). 

T: Venezuela, without locality, Fendler s. n. 

(could not be located). 

HS: C. exaltatus (C. Müll.) Kindb., Enum. 

Bryin. Exot. 88 (1889). 

CU: C. lamellinervis (C. Müll.) Mitt. var. 

exaltatus (C. Müll.) J.-P.Frahm, Bryoph. Bibl. 

5:78 (1975). 

OU: C. cubensis Sull. var. exaltatus (C. Müll.) 

J.-P.Frahm, Nova Hedwigia 41:275 (1985). 

exasperatus (Nees & Blume) Brid., Bryol. Univ. 

1:473 (1826). 

B: Trichostomum exasperatum Nees & Blume, 

Nova Acta Leopold. Carol. 11: 134 

(1823). 

OU: Thysanomitrion exasperatum (Nees & 

Blume) Reinw.& Hornsch., Nova Acta Acad. 

Leop. Carol. 14(2): 704 1829 

T: Indonesia, Java, Blume s.n. (PC).

40 

exasperatus (Nees & Blume) Brid. var. minor C. 

Müll. in Fleisch., Musci Fl. Buitenzorg 

1:115 (1904) nom. nud. 

CU: C. umbellatus (Arn.) Par. 

OU: C. richardii cf. Dixon, Journ. Bot. 

60:289 (1922). 

excurrens Dix., Proc. R. Soc. Queensland 53:27 

(1941). 

T: Australia, Brown Bay, North Queensland, 

Flecker 703 (isotype B). 

CU: C. japonicus Broth. cf. Frahm, The 

Bryologist 96: 142 (1993). 

OU: C. umbellatus (Arn.) Par. cf. Frahm, 

Cryptogamie Bryol. Lich. 3(1):65 (1982); C. 

eberhardtii cf. Frahm, Bryol. Beitr. 7: 32 

(1987); C. richardii cf. Dixon, J. Bot. 60: 289 

(1922).. 

exfrimbriatus C. Müll., Flora 83:331 (1897).(*) 

eximius Reichdt., Verh. Zool. Bot. Ges. Wien 

18:193 (1868). 

T: St. Paul, Jelinek 106 (G,S). 

CU: C. incrassatus C. Müll. cf. Frahm, Bryol. 

Beitr. 7: 32 (1987). 

extinctus J.-P. Frahm, The Bryologist 99: 218, 

1996. 

T: Brazil, „auf Baumfarnstämmen cult. im 

Bot. Garten Göttingen. Sta. Catharina, 

Grossmann s.n. 1904 (GOET) 

exustus Mitt., J. Linn. Soc. Bot. 12:83 (1869). 

T: Andes Peruvianae, in monte Pingulla, 

Spruce 57 (holotype NY). 

OU: Campylopodium pusillum cf. Frahm, 


CU: Microcampylopus curvisetus (Hampe) 

Giese & J.-P.Frahm. 

F 

falcatulus Bartr., Contr. U.S. Nat. Herb. 26: 64 

(1928). 

T: Costa, Rica Cerro de las Lajas, Standley 

51507 (holotype FH, isotypes H-BR, S, NY). 

OU: C. arctocarpus (Hornsch.) Mitt. cf. 


CU: C. arctocarpus ssp. caldensis (Aongstr.) 

J.-P.Frahm cf. Frahm, Nova Hedwigia 31:431 

(1979). 

falcatus Ferg. in Par., Ind. Bryol. ed. 2,1:307 

(1904) nom. nud. 

CU: C. atrovirens De Not. fo. cf. 

Moenkemeyer, Laubm. Eur. 225 (1927). 

falcifolius Mitt., J. Linn. Soc. Bot. 14:481 (1875). 

T: St. Paul, Milne 23 (holotype NY). 

OU: C. eximius Reichdt. cf. Frahm, Bryoph. 

Bibl. 31:67 (1985). 

CU: C. incrassatus 

fallaciosus (Thér.) Podp., Consp. 128 (1954). 

CU: C. pyriformis var. fallaciosus (Thér.) 

Corley, J. Bryol. 9:201 (1976). 

fendleri (C. Müll.) Kindb., Enum. Bryin. Exot. 

88 (1889). 

B: Dicranum fendleri C. Müll., Linnaea 42: 

472 (1879). 

T: Venezuela, Fendler 41 (H-SOL, NY). 

OU: C. rosulatus (Hampe) Mitt. cf. Frahm, 

Bryol. Beitr. 7: 33 (1987). 

CU: C. pauper (Hampe) Mitt. 

fergussonii Stirt., Trans. Bot. Soc. Edinburgh 

26:243 (1914). 

CU: C. atrovirens De Not. cf. Dixon, J. Bot. 

61:15 (1923). 

fernandezianus Bartr., Ark. Bot. ser. 2,4:31 

(1957). 

CU: Dicranoloma menziesii (Tayl.) Par. cf. 

Robinson, Smiths. Contr. Bot. 27:22 (1975). 

fernandezianus Card. in Skottsb., K. Svensk. Vet. 

Ak. Handl. 51(9):65 (1914), nom. nud. 

ferrieri Broth. in Par., Ind. Suppl. 92 (1900) nom. 

nud. 

M: Japan, Nagasaki, Ferrié 206 (H-BR). 



ferromecoae Par., Ind. Bryol. ed.2,1:308 (1904). 

CU: Dicranoloma kunertii (C. Müll.) Ren. 

fibrobasius Dus., Ark. Bot. 4(13):1 (1905). 

OU: C. torquatus Mitt. in Hook. cf. Frahm, 

Bryoph. Bibl. 5:48 (1975). 


filescens Ren. & Card., Bull. Soc. R. Bot. Belg. 

29(1):169 (1890). 


Bibl. 31:196 (1985). 

filicaudatus (C. Müll.) Broth., Nat. Pfl. 1(3):332 

(1901). 

B: Dicranum filicaudatum C.Müll., Hedwigia 

39:260. (1900). 

T: Brazil, Minas Geraes, Serra de Ouro Preto, 

Ule 1367 (holotype destroyed at B; lectotype, 

H-BR). 

CU: C. aemulans cf. Frahm, Revue Bryol.

Lich. 45:135 (1979). 

filicaulis (Hampe) Mitt., J. Linn. Soc. Bot. 12:74 

(1869). 

OU: C. nano-filifolius cf. Robinson, The 

Bryologist 70:17 (1967). 


P.Frahm. 

filicuspis Broth. in Herz., Bibl. Bot. 87:22 (1916). 

T: Bolivia, Comarapa 2600 m, Herzog 4192 

(holotype JE; isotypes B, FH, H-BR, S). 


39:155 (1984). 

filifolioides C. Müll., Gen. Musc. Fr. 269 (1900), 

nom. nud. 

M: Sumatra, Singalang, Beccari s.n. (B, BM, 

JE). 


Frahm, Trop. Bryol. 1:9 (1989). 

filifolius (Hornsch.) Mitt., J. Linn. Soc. Bot. 12:76 

(1896). 

B: Dicranum filifolium Hornsch., Fl. bras. 1: 

12. (1840). 

T: Brazil, „prope Novo-Friburgum“, Beyrich 

s.n. (lectotype ). 


P.Frahm, Cryptogamie Bryol. Lich. 3:366 

(1982). 

filifolius (Hornsch.) Mitt. var. humilis (Mont.) J.- 

P.Frahm, Nova Hedwigia 29:249 (1978). 

CU: Bryohumbertia filifolia var. humilis 

(Mont.) J.-P.Frahm, Cryptogamie Bryol. 

Lich. 3:366 (1982). 

filifolius (Hornsch.) Mitt. var. longifolius Bartr., 

Contr. U.S.Nat. Herb. 26:64 (1928). 

OU: C. filifolius (Hornsch.) Mitt. var. 

longifolius (Bartr.) Bartr. 

OU: C. filifolius cf. Frahm, Nova 

Hedwigia 41:275 (1985). 


P.Frahm 

fimbriatus Mitt., J. Linn. Soc. Bot. 12:88 (1869). 

T: Type. Ecuador, Quito, Spruce 51 (lectotype 

NY). 

CU: C. fragilis (Brid.) B.S.G. cf. Robinson, 


flaccidus (C. Müll.) Broth., Nat. Pfl. 1(3):334 


B: Dicranum flaccidum C. Müll., Hedwigia 

39: 251. 1900, nom. invalid. 

HS: C. ouropretensis Par. 

41 

T: Brazil. Minas Gerais, Serra de Ouro Preto, 

Ule 1362 (holotype H-BR). 

OU: C. ?controversus (Hampe) Jaeg. cf. 

Frahm, Rev. Bryol. Lich. 45:168 (1979). 

CU: C. thwaitesii 

flaccidus Ren. & Card., Bull. Soc. Bot. Belg. 32 

(2): 11 1893 

T: Madagascar, Ambohimatsara pres 

Ambositra (Betsileo) Berthieu 329 (holotype 

PC) 

flaccidus Ren. & Card. var. cucullatifolius J.-P. 

Frahm in Frahm & O´Shea, J. Bryol. 19: 128, 

1996. 

T: Burundui, Mont Manga Mogongo, 

2350m, De Sloover 19.187 (holotype NAM). 

flagellaceus (C. Müll.) Mitt., J. Linn. Soc. Bot. 

12:77 (1869) 

B: Dicranum flagellaceum C. Müll., Syn. 

Musc. 2: 597 (1851). 

OU: Atractylocarpus flagellaceus (C. Müll.) 

Williams 

CU: A. stenocarpus cf. Padberg & Frahm, 


flagelliferus (C. Müll.) Jaeg., Ber. S. Gall. 

Naturw. Ges. 1870-71:423 (1872). 

B: Dicranum flagelliferum C. Müll., Bot. Zeit. 

11: 35 (1853). 

T: India, Nilghiri Mtns., Schmid s.n. (NY). 

flagelliferus (C. Müll.) Jaeg. var. brachyphyllulus 

Card. nom. dub. 



flageyi Ren. & Card., Bull. Soc. R. Bot. Belg. 

32:12 (1893) 

T: Madagascar, forêt dÁnamalaroatra, 

Camboué 251 (holotype PC). 

OU: Campylopus filifolius (Hornsch.) Mitt. 

cf. Frahm, Lindbergia 8:51 (1982). 


P.Frahm. 

flavescens Rehm. in Par. Ind. Bryol. 247 (1894) 

nom. nud. 

CU: C. aureonitens 

flavescens Rehm. in Kindb., Enum. Bryin. Exot. 

88 (1889) nom. nud. 

OU: C. leptotrichaceus (C. Müll.) Par. cf Sim, 



Bryoph. Bibl. 31:160 (1985). 

flavicoma C. Müll. ex Broth., Bot. Jahrb. 24:235

42 

(1897). 

T: Cameroun, supra Bueam pagum, Dusén 

299 (H-BR). 

CU: Bryohumbertia flavicoma (C. Müll.) J.- 


(1982). 

flavissimus (C. Müll.) Besch., Bull. Soc. Bot. 

France 32:LVI (1885). 

T: „Magellan, Churucca, Savatier“ (PC). 

CU: Chorisodontium fulvastrum Besch. cf. 


flavo-nigritus Dus., Ark. Bot. 4(13):8 (1905). 

T: Chile, Patagonia, in insulis Guaitecas, 

Dusén 610 (FH, NY). 

CU: C. chilensis De Not. cf. Frahm, Bryol. 

Beitr. 7: 34 (1987). 

flavo-viridis Dus., Ark. Bot. 4(13):2 (1905). 

T: Chile, Patagonia, ad Puerto Bueno in terra 

truncisque dejectis, Dusen 103 (H-BR). 

OU: C. eximius Reichdt. cf. Frahm, Bryoph. 

Bibl. 31:67 (1985). 

CU: C. incrassatus C. Müll. 

flavotextus Thér. in Broth., Nat. Pfl. ed.2,10:187 

(1924) nom. nud. 

CU: C. aureonitens 

flexifolius Bosch & Lac., Bryol. Jav. 1:76 (1858). 

T: Java, Reinhardt s.n. (holotype L). 

OU: C. caudatus (C. Müll.) Mont. var. 

flexifolius (Bosch & Lac.) Fleisch., Musci Fl. 


CU: C. comosus (Reinw. & Hornsch.) Bosch 

& Lac. 

flexipes (Mitt.) Fleisch., Musci Fl. Buitenzorg 

4:1685 (1923). 


Müll.) Besch. cf. Fleischer, Musci Fl. 



& J.-P.Frahm. 

flexipilis Dix., Bishop Mus. Occ. Pap. 19(11):222 

(1948). 

T: Fiji, Viti Levu, Greene 612 (BM). 


7: 35 (1987). 

flexuosus (Hedw.) Brid., Mant. musc. 4: 71 

(1819). 

B: Dicranum flexuosum Hedw., Spec. musc. 

frond. 145 (1801). 

T: „145 Dicranum flexuosum Dill. t. 47 f 33.“ 

Timm s. n. (lectotype G). 

flexuosus var. incacorralis (Herzog) J.P. Frahm 

Bryol. Beitr. 7: 35. (1987). 

B: C. incacorralis Herz. 

flindersii Catcheside & J.-P.Frahm, J. Bryol. 

13:363 (1985). 

T: S. Australia, Flinders range, Mt. 

remarkable, Melrose, Catcheside 53.169 

(holotype AD, isotype BM). 

foxworthii Broth., Philipp. J. Sc. 5:139 (1910). 

T: Philippines, Mt. Pinatubo, Luzon, 

Foxworth 2551, 2552, 2349, 2544, 5135 

(syntypes NY). 

OU: C. hemitrichius (C. Müll.) Jaeg. cf. Bartram, 

Philipp. J. Sc. 68:41 (1939). 


(C. Müll.) J.-P. Frahm. 

fragiliformis J.-P.Frahm, Rev. Bryol. Lich. 

45:147 (1979). 

T: Type. Brazil, Rio de Janeiro, Parque 

Nacional Itatiaia, Agulhas Negras, Frahm 

1637 (holotype hb. Frahm, isotypes ALTA, 

B, MO, SP, U). 


fragiliformis (J.-P. Frahm) J.-P. Frahm, Bryol. 

Beitr. 7: 35, 1987. 

fragilis (Brid.) B.S.G. Bryol. Eur. 1: 114 (1847). 

B: Dicranum fragile Brid. J. f. Bot. 1800 (2): 

296 (1801). 

T: without locality, hb. Dickson no. 40 

(lectotype BM). 

fragilis (Brid.) B.S.G. ssp fragiliformis (J.-P. 

Frahm) J.-P. Frahm, Flor. Neotropica 54: 101 

(1991). 

fragilis (Brid.) B.S.G. ssp. goughii (Mitt.) J.-P. 


fragilis (Brid.) B.S.G. ssp. zollingerianus (C. 

Müll.) J.-P. Frahm, Trop. Bryol. 4: 61 

(1991). 

friabilis (Hampe) Jaeg., Ber. S. Gall. Naturw. 

Ges. 1870-71: 432. (1872). 

B: Dicranum friabile Hampe in C. Müll., Bot. 

Zeit. 17: 220. (1859). 

T: Costa Rica, Las Nubes, Wendland s. no. 

(holotype not seen; isotype GOET). 

OU: C. chrismarii (C. Müll.) Mitt. cf. 

Williams, N.Am. Fl. 15:136 (1913). 


frigidus Lesq., Misc. Publ. U.S. Geol. Surv. 4:155 

(1874). 

CU: Paraleucobryum sauteri B.S.G. &

Campylopus schimperi Milde cf. Grout, Moss 

Fl. N. Am. 1:96, 134 (1937). 

fuegianus Dus., Ark. Bot. :15 (1905). 

T: Tierra del Fuego, Dusén 271 (H,S). 


Beitr. 7: 35 (1987). 

fulvoviridis Stirt., Ann. Scott. Nat. Hist. 

11(42):105 (1902). 

CU: C. brevipilus B.S.G. cf. Dixon, J. Bot. 

61:15 (1923). 

fulvus (Hook.) Kindb., Bih. K. Svensk Vet. Ak. 

Handl. 7(9):88 (1883). 

CU: Dicranum fulvum Hook. 

fulvus Herz., Bibl. Bot. 87:21 (1916), hom. illeg. 

T: Bolivia, Rio Paracti, 1800 m, Herzog 4996 

(holotype JE, isotypes B, FH, H-BR, S). 

OU: C. longisubulatus Thér. cf. Frahm, Bryol. 

Beitr. 7: 35 (1987). 

CU: C. anderssonii (C. Müll.) Jaeg. 

fumarioli C. Müll., Abh. Naturw. Ver. Bremen 

16(3):496 (1900). 

T: Hawaii, Kilauea, Schauinsland 150 

(BREM). 

CU: C. laxitextus Lac. cf. Frahm, The 

Bryologist 94: 64, 1991. 

funalis (Schwaegr.) Brid., Mant. Musc. 75 

(1819). 

CU: Grimmia funalis (Schwaegr.) B.S.G. 

fuscatus Besch., J. Bot. 5: 146. 1891. Type. 

fuscatus Besch., J. de Bot. 5:146 (1891). 

T: Uruguay, Montevideo, Courbon s. no. 


CU: C. cf. occultus Mitt. cf. Frahm, Bryol. 

Beitr. 7: 36 (1987). 

f usco-croceus (Hampe) Jaeg., Ber. S. Gall. 

Naturw. Ges. 1877-78:382 (1880). 

B: Dicranum fuscocroceum Hampe, Vid. 

Medd. Naturh. For.Kjoebenh. ser. 3,4: 45 

(1872). 

fusco-luteus Card. in Skottsb., K. Svensk. Vet. 

Ak. Handl. 51(9):68 (1914) nom. nud. 

fusco-luteus Stirt., Ann. Scott. Nat. Hist. 6:117 

(1897). 

CU: C. subulatus Schimp. fo. cf. Podpera, 

Consp. 127 (1954). 

fusco-viridis (Card.) Dix. & Thér. ex Hong & 

Ando, Biol. Inst. Cath. Med. Seoul Korea 

3:373 (1959). 

T: Corée, Quen San, Faurié 18 (isotype BM). 

OU: C. japonicus Broth. var. fusco-viridis 

43 

Card. 

CU: C. atrovirens De Not. cf. Frahm, J. 

Hattori Bot. Lab. 71: 135 (1992). 

fusco-viridis Hampe in C. Müll., Gen. Musc. Fr. 

272 (1900) nom. nud. 

fuscolutescens Ren. & Card., Bull. Soc. R. Bot. 

Belg. 35(1): 302 (1897). 

OU: Thysanomitrion fuscolutescens (Ren. & 

Card.)Broth Nat. pfl. ed 2, 10: 188 (1924). 

T: Maurice, Robillard (holotype PC). 


Frahm, Hikobia 11 (1994). 


G 

galapagensis J.-P.Frahm & Sipman, J. Bryol. 

10:61 (1978). 

T: Ecuador, Galapagos Islands, Santa Cruz, 

base of Mt. Crocker, Gradstein & Weber 17 

(holotype U; isotypes CDRS, COLO, QCA) 

CU: C. pilifer Brid. var. galapagensis (J.-P. 

Frahm & Sipman) J.-P. Frahm cf. Frahm, 

Bryol. Beitr. 7: 88, 1987. 

gallienii Par., Rev. Bryol. 27:90 (1900). 

T: Madagascar, Fort Dauphin, Offiziers s.n., 


CU: C. robillardei Besch. cf. Frahm, Bryoph. 

Bibl. 31:164 (1985). 

gardneri (C. Müll.) Mitt., J. Linn. Soc. Bot. 12: 

83 (1869). 

B: Dicranum gardneri C. Müll. in C. Müll. 

& Hampe, Bot. Zeit. 15: 379 (1857). 

T: Brazil, Pernambuco, Sierra de Araripe, 

Gardner 20 (lectotype NY; isolectotype, S). 

gastro-alaris (C. Müll.) Par., Ind. Bryol. ed. 2,1: 

310. (1904). 

B: Dicranum gastro-alare C. Mülller, 

Hedwigia 39: 256. (1900). 

T: Brazil, Minas Gerais, Serra de Caraca, Ule 

1361 (holotype destroyed at B; lectotype H- 

BR). 

gaudichaudii Besch., J. de Bot. 5:148 (1891) 

T: Chile, Ile S. Catherine (Holotypus PC). 

Location probably wrong. 


Frahm, Crypt. Bryol. Lichènol. 39: 32, 1998. 

gedehensis Baumg. & Froehl., Naturh. Mus.

44 

Wien 59:70 (1953) (*) 

T: Java, im Gedeh Krater 2700m, Schiffner 

10.186 (holotype W?, isotype S). 

CU: C. macgregorii Broth. & Geh. cf. Frahm, 

Trop. Bryol. 15 (in press). 

gemmatus (C. Müll.) Par., Ind. Bryol. 82. (1900). 

B: Dicranum gemmatum C. Müll., Bull. Herb. 

Boiss. 6: 34 (1898). 

T: Brazil. Serra do Itatiaia, Ule 1789 


geniculatus (Schwaegr.) Fuernr., Flora 12:595 

(1829). 

CU: Campylostelium saxicola (Web. & 

Mohr) B.S.G. cf. Paris, Ind. Bryol. 265 

(1894). 

geniculatus Aongstr., Oefv. K. Svensk. Vet. Ak. 

Foerh. 29(4):80 (1872) hom. illeg. 

T: Hawaii, Honululu, Anderson s.n. (S, B). 

CU: C. umbellatus (Arn.) Par. var. 

pterotoneuron (C. Müll.) J.-P.Frahm, Nova 

Hedwigia 39:602 (1984), OU: C. richardii 

Brid. cf. Dixon, J. Bot. 60:287 (1922). 

genuflexus C. Müll., Gen. Musc. Fr. 266 (1900) 

nom. nud. 

geraense Par., Ind. Bryol. ed.2,1:310 (1904) (*). 

germainii Demar. & Leroy, Bull. Jard. Bot. 

Bruxelles 19(1):85 (1948). 

T: Congo Belge, Distr. Lac Edouard et Kivu, 

Nyamurugira, Tshambene, Germain 3759 

(isotype PC). 

CU: Pilopogon africanus Broth. in Mildbr. 

cf. Frahm, Bryoph. Bibl. 31:196 (1985). 

geronticus C. Müll. Gen. Musc. Fr. 267 (1900) 

nom. nud. 

gertrudis Herz., Bibl. Bot. 87:21 (1916). 

T: Bolivia, Sillar 2000 m, Herzog 2731 

(holotype JE; isotypes B, FH, H-BR, S). 


39:155 (1984). 

gibboso-alaris (Broth. & Par.) Wijk & Marg., 

Taxon 8: 72 (1959). 

B: Thysanomitrion gibboso-alaris 

Broth.&Par. Nat. Pfl. ed. 2, 10: 189 (1924). 

T: „Nova Caledonia, plateau de Dogny, soil 

a. rock, 750-1050 m“ Le Rat 1402 (holotype 

PC). 



giganteus Sull., Proc. Am. Ac. Arts Sc. 5:278 

(1861). 

T: Cuba, Wright s. n. (not seen). 

OU: C. tortuosus (Hampe) Par. cf. Williams, 

N. Am. Fl. 15:147 (1913). 

CU: C. lamellinervis (C. Müll.) Mitt. 

glauco-pallidus C. Müll., Gen. Musc. fr. 270 

(1900) nom. nud. 

M: S. Catarina, Ule 703, 836 (H-BR). 

CU: occultus Mitt. cf. Frahm, Revue Bryol. 

Lich. 42:901 (1976). 

glauco-viridis (C. Müll.) Par., Ind. Bryol. Suppl. 

92 (1900). 

OU: C. capillatus Hook.f. & Wils. cf. Dixon, 

New Zeal. Inst. Bul.. 3(3):86 (1923); C. 

pallidus Hook.f. & Wils. cf. Sainsbury, R. 

Soc. New Zeal. Bull. 5:116 (1955). 


glaziovii (Hampe) Jaeg., Ber. S. Gall. Naturw. 

Ges. 1877-78:385 (1880). 

B: Dicranum glaziovii Hampe, Vid. Medd. 

Naturh. Foer. Kjoebenh. ser. 3, 6: 138. (1875). 

T: Type. Brazil, Rio de Janeiro, Glaziou 6368 

(isotype NY). 



goughii (Mitt.) Jaeg., Ber. S. Gall. Naturw. Ges. 

1870-71:424 (1872). 

B: Dicranum goughii Mitt., J. Linn. Bot. 

Suppl. 1: 17 (1859). 

T: Sikkim, Hooker 79 (holotype NY, isotype 

H-SOL). 

OU: C. zollingerianus Bosch & Lac. cf. 



(Mitt.) Frahm cf. Frahm, Trop. Bryol. 4: 61, 

1991. 

gracilentus Card., Bot. Centralbl. 19(2):94 

(1905). 

T: Formosa, Taitun, Faurié 35 (lectotype PC). 

CU: C. laxitextus Lac. cf. Frahm, J. Hattori 

Bot. Lab. 71: 147, 1992. 

gracilicaulis Mitt., J. Linn. Soc. Bot. 12:83 

(1869). 

T: Brazil, Rio Negro, Spruce 60 (holotype 

NY, isotypes BM, FH, NY, PC, S). 

CU: C. surinamensis C. Müll. cf. Frahm, The 

Bryologist 83:581 (1980). 

OU: C. flexuosus (Hedw.) Brid. cf. Bartram, 

Fieldiana Bot. 25:43 (1949). 

gracilicaulis Mitt. ssp. angustiretis (Aust.) Kind., 

Eur. N. Bryin. 2:204 (1897).

CU: C. angustiretis (Aus.) Lesq. & Jam. 

OU: C. surinamensis var. angustiretis (Aust.) 

J.-P.Frahm, The Bryologist 83:582 (1980). 

gracilicaulis Mitt. var. donnellii (Aust.) Grout, 

N.oAm. Fl. 2:92 (1937). 

CU: C. surinamensis C. Müll. cf. Frahm, The 

Bryologist 83:582 (1880). 

OU: C. flexuosus cf. Bartram, Fieldiana Bot. 

25:43 (1949). 

gracilis (Mitt.) Jaeg., Ber. S. Gall. Naturw. Ges. 

1870-71:427 (1872). 

B: Dicranum gracile Mitt. J. Linn. Bot. Suppl. 

1: 17 (1859). 

T: „In Himalayae orient. temp. Sikkim, 

Hooker 69, 72 (syntypes NY). 

OU: C. subulatus Schimp. cf. Frahm, J. 

Hattori Bot. Lab. 71: 154 (1992). 

grimmioides (C. Müll.) Kindb., Enum. Bryin. 

Exot. 88 (1869). 

B: Dicranum grimmioides C. Müll. Linnaea 

43: 397 (1882). 

T: Argentina, Las Punas in tierra de Cordoba, 

Lorentz s.n. Febr. 1871. 

CU: C. introflexus (Hedw.) Brid. var. 

cordobaensis Thér. cf. Frahm. Bryol. Beitr. 

7: 38 (1987). 

griseolus (C. Müll.) Par., Ind. Bryol. Suppl. 92 

(1900). 

B: Dicranum griseolum C.Müll. Hedwigia 

38:80 (1889). 

T: Transvaal, Lydenburg, Wilms s.n. 

(holotype B, isotype G). 

OU: C. polytrichoides De Not. cf. Frahm, 



griseus (Hornsch.) Jaeg., Ber. S. Gall. Naturw. 

Ges. 1870-71: 443 (1872). 

B: Thysanomitrion griseum Hornsch., Fl. 

Bras. 1:16 (1840). 

T: Uruguay, Montevideo, Sellow s.n. 

(holotype destroyed at B; lectotype KIEL). 

griseus Williams ssp ingeniensis (Williams) 

J.P.Frahm, Bull. New York Bot. Gard. 3(9): 

109 (1993). 

T: Bolivia, Ingenio, Williams 1772 (holotype 

NY, isotypes B, FH, H-BR). 

griseus Williams var. lapidicola (C. Müll.) J.- 

P.Frahm, Revue Bryol. Lich. 42:606 

(1976). 

CU: C. griseus s.str. cf. Frahm, Revue 

45 

Bryol. Lich. 45:150 (1979). 

guadeloupensis (Brid.) Mitt., J. Linn. Soc. Bot. 

12:77 (1869). 

B: Dicranum guadeloupense Brid., Spec. 

Musc. 1: 213 (1806). 

T: Guadeloupe,Balby s.n. (holotype B). 

OU: Campylopus guadeloupensis (Brid.) 

Mitt., J. Linn. Soc. Bot. 12: 77 (1869). 

Pilopogon gracilis (Hook.) Brid. cf. Frahm, 

J. Bryol. 11:213 (1980). 

CU: Pilopogon guadeloupensis Brid. 

guaitecae Dus., Ark. Bot. 4(13):13 (1905). 

T: Tierra del Fuego, Desolación, Dusén (L). 


Beitr. 7: 89 (1987). 

guatemalensis Bartr., Bryologist 49:110 (1946). 

T: Guatemala, Dep. Totonicapan, Standley 

86159 (holotype FH; isotypes F, FH, MICH). 

CU: C. albidovirens Herz. cf. Frahm, 


guineensis P. Varde & Thér., Bull. Mus. Hist. 

Nat. Paris ser.2,13:207 (1941) hom. illeg. 

T: Environs de Mali, Mont Luru nr. 2007 

(PC). 

CU: C. obrutus cf. Frahm, Bryol.Beitr. 7: 89 

(1987). 

guineensis Thér., Bull. Mus. Hist. Nat. Peris ser. 

2, 3: 772 (1931). 

T: Guinee fr., Deloba, Chevalier s.n. 


CU: C. obrutus P. Varde & Thér. cf. Frahm, 

Nova Hedwigia 39: 596 (1984) 

gulliveri R. Brown ter., Trans. New Zeal. Inst. 

29:472 (1897). 


Müll.) Besch. cf. Dixon, New Zeal. Inst. Bull. 

3(3):77 (1923) = CU: Campylopodium 

medium (Duby) Giese & J.-P.Frahm. 

H 

haitensis Thér., Rev. Bryol. Lich. 14:8 (1944). 

T: Haiti, Mt. Formon, Ekman 7561 (isotypes 

F, NY, S). 

OU: Campylopus filifolius var. humilis cf. 


CU: Bryohumbertia filifolia var. humilis 

hakoniae C. Müll., Gen. Musc. fr. 266 (1900) 

nom. nud. 

hallii Lesq., Misc. Publ. U.S.Geol. Geogr. Survey 

4:155 (1874).

46 

CU: Paraleucobryum enerve (Thed.) Loeske 

cf. Grout, Moss Fl. N.Am. 2:96 (1937). 

hamatus Broth. in Roiv., Ann. Bot. Soc. Fenn. 

Vanamo 9(2):29 (1937). 

T: Chile, Vulcan Uaima, Ruthsatz (1966) s.n. 


CU: ? Chorisodontium sp. 

handelii Broth., Symb. Sin. 4:18 (1929). 

T: China, Yünnan, Handel-Mazetti 7676 

(isotype S). 



handelii Broth. var. setschwanicus Broth., Symb. 

Sin. 4: 18 (1929) 

T: China, Yünnan, Handel-Mazetti 1515 




harpophyllus Herz., Bibl. Bot. 87:22 (1916). 

T: Bolivia, Cerros de Malaga, Herzog 4796 

(holotype JE; isotypes B, S). 

CU: C. edithae Broth. cf. Frahm, Bryol. Beitr. 

7: 39 (1987). 

harrisii (C. Müll.) Par. Ind. Bryol. Suppl. 92 

(1900). 

B: Dicranum harrisii C. Müll., Bull. Herb. 

Boiss. 5: 553 (1897). 

T: Jamaica, Newhaven Pass, Harris 11008 

(holotype destroyed; lectotype NY; 


CU: C. cubensis Sull. cf. Frahm, Nova 

Hedwigia 41:275 (1985). 

harrisii (C. Müll.) Par. var. longifolius (Bartr.) 

Bartr., J. Wash. Ac. Sc. 19:12 (1929). 

OU: C. filifolius (Hornsch.) Mit. var. 

longifolius (Bartr.) Bartr., C. filifolius cf. 

Frahm, Nova Hedwigia 41: 275 (1985). 

CU: Bryohumbertia filifolia (Hornsch.) 

Frahm 

hartmanii Schimp. ex Hartm., Bot. Not. 1855:49 

(1855). 

CU: Tortella fragilis (Hook. & Wils.) Limpr. 

cf. Lindb., Oefv. K. Vet. Akad. Foerh. 20:387 

(1863). 

hawaiico-flexuosus (C. Müll.) C. Müll., Abh. 

Naturw. Ver. Bremen 16(3):494 (1900). 

B: Dicranum hawaiico-flexuosum C. Müll., 

Flora 82: 443 (1896). 

T: Hawaii,without locality, Hillebrand s.n. 

(holotype destroyed at B, isotypes not 

known). 

OU: C. purpureoflavescens (C. Müll.) Par. cf. 

Bartr., Bishop Mus. oc. Pap. 16:323 (1942); 

C. densifolius Aongstr. var. hawaiicoflexuosus 

(C. Müll.) Mill. 

CU: C: hawaiicus (C. Müll.) Jaeg. var. 

hawaiico-flexuosus (C. Müll.) J.-P. Frahm cf 

Frahm, The Bryologist 94: 62 (1991). 

hawaiicus (C. Müll.) Jaeg., Ber. S. Gall. Naturw. 

Ges. 1870-71:436 (1872). 

B: Dicranum hawaiicum C. Müll., Bot. Zeit. 

20: 328 (1862). 

T: Hawaii, Didrichsen in exped. transatl. 

Danica 1861 (PC). 

HS: C. hawaianus (C.Müll.) Par., Ind. Bryol. 

249 (1894). 

OU: C. umbellatus (Arn.) Par. cf. Bartr., 

Bishop Mus. Bull. 101:44 (1933); C. richardii 

Brid. cf. Dixon, J. Bot. 60:287 (1922). 

hawaiicus (C. Müll.) Jaeg. var. hawaiicoflexuosus 

(C. Müll.) J.-P. Frahm cf Frahm, 


heinrici Ren. & Card. ex Kindb., Eur. N. Am. B 

helenicus (C. Müll.) Jaeg., Ber. S. Gall. Naturw. 

Ges. 1870-71:431 (1872). 

B: Dicranum helenicum C. Müll., Syn. Musc. 

2: 599 (1851). 

T: „St. Helena, Kunze donavit“ (could not be 

located). 


Bryoph. Bibl. 31:19 (1985). 

hellerianus (Hampe) Jaeg., Ber. S. Gall. Naturw. 

Ges. 1870-71:417 (1872). 

HS: C. helleri Hampe ex Kindb., Enum. 

Bryin. Exot. 50 (1888). 

B: Dicranum hellerianum Hampe, Verh. K.K. 

Zool. Bot. Ges. Wien 19: 507 (1870). 

T: Mexico, Huatusco, Heller s.n. (FH, NY). 

CU: C. heterostachys (Hampe) Jaeg., cf. 




hemitrichius (C. Müll.) Jaeg., Ber. S. Gall. 

Naturw. Ges. 1877-78:384 (1880). 

B: Dicranum hemitrichium C. Müll., Linnaea 

38: 553 (1874). 

T: Philippines, reg. montana, Wallis 

(lectotype FH). 


(C. Müll.) J.-P. Frahm cf. Frahm, The

Bryologist 94: 64 (1991). 

henrici Ren. & Card., Bot. Gaz. 13:197 (1888). 

CU: Dicranella heteromalla (Hedw.) Schimp. 

cf. Williams, N.Am.Fl. 15:89 (1919). 

hensii Ren. & Card., Bull. Soc. R. Bot. Belg. 

35:301 (1897). 

T: Congo, Bougalu, Hens 36 (isotype G) 

heribaudii Ren. & Card., Bull. Soc. R. Bot. Belg. 

209(1):168 (1890). 

T: Madagaskar circa Tananarive, Camboue 

s.n. (holotype PC, isotypes S). 


Bibl. 31:164 (1985). 

herzogii Broth., Hedwigia 50:122 (1910). 

T: Ceylon, Adams Peak, Herzog (holotype 

JE). 

OU: C. aureus Bosch. & Lac. cf. Frahm, J. 

Bryol. 13:165 (1984). 

CU: C. schmidii C. Müll.. 

heteroneuron Thér., Bull. Soc. Bot. Geneve 26:79 

(1936). 

T: Transvaal, Sanatorium, Junod s.n. 




heterophyllus Mitt., J. Linn. Soc. Bot. 12:77 

(1869). 

T: Type. Colombia. „Andes Bogotenses, in 

montibus inter Bucumaranga et Pamplona“, 

Weir 347 (holotype NY, isotypes FH, H-BR, 

S). 



heterostachys (Hampe) Jaeg., Ber. S. Gall. 

Naturw. Ges. 1870-71: 421 (1872). 

B: Dicranum heterostachys Hampe, Flora 48: 

581 (1865). 

T: Peru, Prov. Carabaya, Sandia, Hasskarl s. 

no. (holotype BM; isotype S). 

hildebrandtianus Fleisch., Musci Fl. Buitenzorg 

1:112 (1904). 

HS: C. hildebrandii Broth. in Fleisch. l.c. 

nom. nud. 

OU:? C. subulifolius cf. Frahm, J. Bryol. 

13:182 (1984). 

CU: C. comosus cf. Eddy, Malesian Mosses 

vol. 1 (1988). 

hildebrandtii (C. Müll.) Jaeg., Ber. S. Gall. 

Naturw. Ges. 1877-78:382 (1880). 

B: Dicranum hildebrandtii C. Müll. Linnaea 

47 

40: 236 (1876). 

T: Comoren Insel Johanna, Hildebrandt 

(1839) (holotype B, isotype G). 

hispidus Ren. & Card., Bull. Soc. R. Bot. Belg. 

29(1):171 (1890). 

T: Madagascar, Imerina, Ambohimalaza, 

Camboué s.n. (holotype PC). 

CU: C. pilifer Brid. cf. Frahm. Bryol. Beitr. 

7: 40 (1987). 

hoehnelii (C.Müll.) Broth. Bot. Jahrb. 20:177 

(1894). 

B: Dicranum hoehnelii C.Müll., Flora 73: 473 

(1890). 

T: Kilimandscharo, Höhnel (1889) s.n. 

(isotype PC, lectotypus nov. BM). 


31:143 (1985). 

hoehnelii (C.Müll.) Broth. var. subelamellatus 

Broth. in Mildbr., Wiss. Erg. Deutsch. Zentr. 

Afr. Exp. 2:141 (1910). 

T: T: Ninagongo, Lavaspalten des Kraters, 

ca. 3300 m, Mildbraed 2004 (H-BR). 

CU: C. johannis-meyeri (C. Müll.) Par. cf. 


hoffmannii (C. Müll.) Ren. & Card., Bull. Soc. 

R. Bot. Belg. 31(1): 147 (1893). 

B: Dicranum hoffmannii C. Müll., Linnaea 

38: 592 (1872). 

T: Costa Rica. Forets du Barba, Bondez 9909 


CU: C. cuspidatus cf. Frahm, J. Hattori Bot. 

Lab. 44:485 (1978) non Allen, Trop. Bryol. 

1: 84 (1989). 

holomitrium (C. Müll.) Jaeg., Ber. S. Gall. 

Naturw. Ges. 1870-71:421 (1872). 

B: Dicranum holomitrium C. Müll., Syn. 

Musc. 1: 389 (1848). 

T: „Ad Insel Bay, Nova Zeelandia“, Hooker 

s.n. (BM). 

OU: C. capillaceus Hook.f. & Wils. cf. Jaeg. 

l.c. 


& J.-P.Frahm cf. Frahm, Bryol. Beitr. 7: 41 

(1987). 

holomitrius auct. = C. lonchochaete C. Müll. 

homalobolax (C. Müll.) Par., Ind. Bryol. Suppl. 

92 (1900). (*) 

homomallus Leroy & Thér., Bull. Jard. Bot. 

Bruxelles 18:162 (1947). 

T: Congo bel., Distr. du Lac Albert, Djugu,

48 

Leroy 44B (holotype BR). 


Bryoph. Bibl. 31:76 (1985). 

hondurensis Bartr., Field Mus. Nat. Hist. Bot. 

4:351 (1929). 

T: Honduras, Standley 56157a (lectotype F). 

CU: C. flexuosus (Hedw.) Brid. cf. Bartram, 


horridus Welw. & Duby Mem. Soc. Phys. Hist. 

Nat. Geneve 21:222 (1870). 

T: Angola, Pungo Andongo 2300 ped., 

Soyana 76 (lectotypus BM), 

CU: savannarum (C. Müll.) Mitt. cf. Frahm, 

Bryoph. Bibl. 31:171 (1985). 

huallagensis Broth., Hedwigia 45:261 (1906). 

T: Peru. Rio Huallaga, Ule 2358 (holotype 

H-BR; isotypes JE, FH). 

huallagensis var. weberbaueri Brotherus Bot. 

Jahrb. 56, Beibl. 123: 5 (1920). 

T: Peru, Loreto: E of Moyobamba, Weberbauer 


humifugus (C. Müll.) Kindb., Enum. Bryin. Exot. 

88 (1889). 

B: Dicranum humifugum C. Müll., Linnaea 

XLIII: 400 (1882). 

T: Argentina, Gran Chaco, Rio Secco, Lorentz 

s. n. (holotype destroyed at B; lectotype H- 

BR; isolectotype FH). 

CU: C. occultus Mitt. cf. Frahm, Bryol. Beitr. 

7: 41 (1987). 

humilis Mont., Ann. Sc. Nat. Bot. ser. 3,4:110 

(1845). 

T: Type. Brazil, Blanchet 103 (holotype PC; 

isotype NY). 

OU: C. filifolius var. humilis (Mont.) J.- 

P.Frahm, Nova Hedwigia 29:249 (1978). 


P.Frahm var. humilis (Mont.) J.-P. Frahm. 

humoricola (C. Müll.) Par., Ind. Bryol. ed.2,1:313 

(1904). 

B: Dicranum humoricola C. Müll., Hedwigia 

39: 256 (1900). 

T: Brazil, Rio de Janeiro, Corcovado, Ule 

1650 (holotype destroyed at B; lectotypus 

nov., H-BR). 

OU: C. controversus cf. Frahm, Revue Bryol. 

Lich. 43:141 (1979). 


huntii Stirt., Ann. Scott. Nat. Hist. 8(30):106 

(1899). 

OU: C. schwarzii Schimp. var. huntii (Stirt.) 

Dix. = C. schwarzii s.str. ? 

CU: C. gracilis (Mitt.) Jaeg. 

hygrometricus Rehm. in Geh., Rev. Bryol. 5:69 

(1878) nom. nud. 

OU: C. purpurascens Lor. cf. Sim, Trans. R. 

Soc. S. Afr. 15:176 (1926); C. pilifer Brid. 

cf. Magill, Fl. S. Afr. Bryoph. 1(1):152 

(1981). 

CU: C. aureonitens (C. Müll.) Jaeg. cf. Frahm, 

Bryoph. Bibl. 31:21 (1985). 

hygrophilus Broth. in Skottsb., Nat. Hist. Juan 



Beitr. 7: 41 (1987). 

I 

inandae (C.Müll.) Par. Ind. Bryol. Suppl. 92 

(1900). 

B: Dicranum inandae C.Müll. Hedwigia 

38:95 (1899). 

T: Natal, Inanda, Rehmann 43 (lectotypus 

nov. NY, isotypes L, G, PC ). 




Soc. S. Afr. 15:176 (1926). 

incacorralis Herzog, Beih. Bot. Centralbl. 26: 52 

(1910). 

T: Bolivia, Incacorral, Herzog 285 (holotype 

JE, isotype B). 

CU: C. flexuosus (Hedw.) Brid. var. 

incacorralis (Herz.) J.-P.Frahm, Bryol. Beitr. 

7: 42 (1987) comb. invalid. 

incertus Thér., Rev. Bryol. Lichenol. 11: 63 

(1939). 

T: Colombia, Paramo El Boqueron near 

Bogota, Troll 2161 (lectotype PC, isotypes 

FH, JE). 

inchangae (C. Müll.) Par., Ind. Bryol. Suppl. 92 

(1900). 

B: Dicranum imerangae C. Müll. Hedwigia 

38: 83 (1899). 

HS: C. imerangae (C. Müll.) Par., Ind. Bryol. 

Suppl. 92 (1900), Campylopus incrangae (C. 

Müll.) Dix. & Gepp, Bull. Misc. Inf. Kew 

1923:197 (1923), Campylopus inchangae (C. 

Müll.) Thér. in Dix., Trans. R. Soc. S. Afr. 

8:185 (1920). 

T: Natal, Inchanga,Rehmann 42 (BM).


Bibl. 31:164 (1985). 

incrassatus C. Müll. Linnaea 18:686 (1845). 

T: Chile, Talcahuana, Poeepig 22 (H-BR, JE, 

NY). 

incurvatus J.-P.Frahm & Hoe, Bryologist 81:449 

(1978). 

T: Hawaii: E Mauna Kea, S. Hilo Distr., 

Kolekole Stream 585m, Balakishnan & 

Jacobi 30328 (holotype hb. Frahm, isotypes 

BISH, MICH, MO, NICH, U, hb. Hoe). 

ingeniensis Williams, Bull. New York Bot. Garden 

3(9):109 (1903). 

T: Bolivia, Ingenio, Williams 1772 (holotype, 

NY; isotypes B, FH, H-BR).CU: C. griseus 

(Hornsch.) Jaeg. ssp. ingeniensis (Williams) 

J.-P. Frahm. 

insignis Herz., Bibl. Bot. 87:23 (1916). 

T: Bolivia, Abra San Mateo. Herzog 3721 


OU: C. multicapsularis (C. Müll.) Schimp. 

cf. Theriot, Revue Bryol. 11:42 (1939). 

CU: C. nivalis (Brid.) Brid. var. 

multicapsularis (C. Müll.) J.-P. Frahm. 

insistitius Hook.f. & Wils., Fl. Tasman. 2:172 

(1859). 

HS: C. insidiosus Hook.f. & Wils. ex Mitt., 

Proc. R. Soc. Victoria 19:55 (1892) 


Trans. New Zeal. Inst. 59:507 (1928). 

insularis Bartr., Proc. California Ac. Sc. 

ser.4,21:79 (1933). 

T: Ecuador, Galapagos Isl.: Duncan Isl., 

Stewart 3323 (lectotype FH). 

CU: C. anderssonii Bartr. cf. Frahm, Fl. 

Neotr. 54: 47 (1991). 

intermedius Wils. in Hunt., Mem. Lit. Phil. Soc. 

Manchester ser.3,3:234 (1868) nom. nud. 

OU: Campylopus alpinus cf. Hunt l.c. 

CU: Dicranodontium denudatum (Brod.) 

Britt. ssp. alpinum (Schimp.) Giac. 

interruptulus C. Müll. ex Besch., Ann. Sc. Nat. 

Bot. ser. 6,9:324 (1880). 


Bibl. 31:197 (1985). 

intorquatus Watts. & Whitel., Proc. Linn. Soc. 

N.S.Wales Suppl. 27:45 (1902) nom. nud. 

intorquatus Watts. & Whitel. var. flavifolius 

Watts. & Whitel. l.c., nom. nud. 

introflexus (Hedw.) Brid., Mant. Musc. 72 

49 

(1819). 

B: Dicranum introflexum Hedw. Spec. Musc. 

147 (1801). 

T: „Nova Hollandia“ (G ). 

introflexus (Hedw.) Brid. var. cordobaensis Thér. 

in Bauer, Musci Eur. Am. Exsicc. ser. 45 

n. 2204 (1934). 

CU: C. pilifer x C. introflexus. 

involutus (C. Müll.) Jaeg., Ber. S. Gall. Naturw. 

Ges. 1870-71:418 (1872) . 

B: Dicranum involutum C. Müll., Bot. Zeit. 

11: 34 (1853). 

T: India, Montes Neilgherienses, Schmid s.n. 

(H-BR). 

CU: C. ericoides (Griff.) Jaeg. cf. Frahm, J. 

Hattori Bot. Lab. 71: 140, 1992. 

irregularis Lor., Verh. Zool. Bot. Ges. Wien 

17:683 (1867) nom. nud. 

CU: Ditrichum heteromallum (Hedw.) Britt. 

var. zonatum (Brid.) Podp. cf. Lor. l.c. 

irrigatus Thér., Bull. Ac. Geogr. Bot. 19:18 

(1909). 

T: China, Konyt Chen, Cavalerie 1673 

(holotype PC(. 

CU: C. japonicus Broth. cf. Frahm, Bryol. 

Beitr. 7: 43 (1987). 

isesanctus Sak., Bot. Mag. Tokyo 65:90 (1952). 

CU: Dicranella spec. cf. Takaki, J. Hatt. Bot. 

Lab. 30:247 (1967). 

itacolumitis (C. Müll.) Broth., Nat. Pfl. 1(3):334 

(1901). (*) 

J 

jamaicensis Mitt., J. Linn. Soc. Bot. 12: 82. 1869. 

T: Jamaica, Wilson 813 (holotype NY). 

CU: Campylopus cygneus (Hedw.) Brid. cf. 

Williams, N. Am. Fl. 15(2):137 (1913). 

jamesonii (Hook.) Jaeg. Ber. S. Gall. Naturw. 

Ges. 1870-71: 422 1872 

B: Dicranum jamesonii Hook., Pl. rar. 2: 179 

1837 

T: Ecuador „in montre Surruchcho“ Jameson 

s.n. 

japonicus Broth., Hedwigia 38:207 (1899). 

T: Japan, without detailed locality, Ankarcrona 

s. n. (holotype, H-BR). 

CU: Campylopus sinensis (C. Müll.) J.-P. 

Frahm cf. Frahm, Ann. Bot. Fenn. 34: 202, 

1997.

50 

jashii Broth. in Nav., Bull. Soc. R. Bot. Belg. 

60(1):21 (1927) nom. nud. 

CU: Dicranum jashii Thér. cf. Nav. l.c. 

johannis-meyeri (C. Müll.) Par., Ind. Bryol. 251 

1894 

B: Dicranum j.-m. C. Müll. Flora 71: 410 1888 

T: Kilimandscharo, ad finem silvestris 3000- 

4000 m Dr. Meyer s.n. (Lectotype PC) 

joinvilleanus (Hampe) Jaeg., Ber. S. Gall. 

Naturw. Ges. 1877-78:382 (1880) 

B: Dicranum joinvilleanum Hampe, Vid. Medd. 

Naturh. For. Kjoebenh. ser. 3,4: 45. 1872. 

OU: C. joinvillei Hampe ex Kindb., Enum. 

Bryin. Exot. 50 (1888) nom. illeg. 


Lich. 45:156 (1979). 

jucrangae Par., Ind. Bryol. 251 (1894) err. pro 

C. inerangae 

OU: C. inchangae (C. Müll.) Par. 

CU: C. robillardei cf. Frahm, Bryoph. Bibl. 

31:164 (1985). 

jugorum Herzog, Beih. Bot. Centralbl. 26(2): 51. 

1910. 

T: Bolivia, Cochabamba, Abra San Benito, 

Herzog 3356 (holotype, JE; isotype, B). 

julaceus Jaeg., Ber. S. Gall. Naturw. Ges. 1877- 

78: 384 1880 

B: Dicranum julaceum Hampe, Vidd. Medd. 

Naturh. For. Kjoebenhavn ser. 3,4:45. 1872 

hom. illeg. 

T: Brazil, Rio de Janeiro, Serra dos Orgaos, 

Glaziou 4554 (holotype BM? [not seen]; 

isotypes H-BR, PC). 

julaceus Jaeg. ssp. arbogastii (Ren. & Card.) J.- 

P. Frahm, Bryophytorum Bibliotheca 31: 109 

1985 

B: Campylopus arbogastii Ren. & Card. 

T: Sainte Marie de Madagascar, in terra 

arenosa, Arbogast s.n. (holotype PC, isotype 

G) 

julaceus Rehm. in Par., Ind. Bryol. 252 (1894) 

nom. nud. 

M: Orange Free State, Kadziberg, Rehmann 58 

(B, L). 

OU: C. pseudojulaceus Sim hom. illeg. cf. Sim, 

Trans. R. Soc. S. Afr. 15:169 (1926), C. simii 

Schelpe 

nom. nov. 

CU: C. julaceus Jaeg. ssp. arbogastii 

julicaulis Broth., Denkschr. Ak. Wiss. Wien 

math.-nat. Kl. 83:261. 1926. 

T: Brazil. Paraná, Roca Nova, Dusén 245 

(holotype H-BR; isotypes, FH, JE, NY). 

K 

kaalaasii Hag. in Kaal., K. Norsk Vid. Selsk. 

Skrifter 1910 (7): 55 (1911) 

CU: Dicranodontium denudatum (Brid.) Britt. 

var. kaalaasii (Hag.) C.Jens. 

kingii Robins., Bryologist 70(1): 18 (1967) 

T: Colombia, King & Guevara C-1132 

(holotype US, isotype COLO) 

OU: C. chrysodictyon (Hampe) Mitt. cf. Frahm, 

J. Hatt. Bot. Lab. 44:485 (1978) 

CU: C. pauper (Hampe) Mitt. cf. Frahm, Fl. 

Neotr. 54: 148 (1991). 

kirkii Mitt. in Beck., Trans. New Zeal. Inst. 

26:280 (1894) 

CU: C. acuminatus Mitt var. kirkii (Mitt.) J.-P. 


kirkii Mitt. in Beck. var. pilosus Frahm, 

Lindbergia 7: 30 (1981). 

T: New Zealand, North Island, Kaiiwi & 

Taharoa Lake, Barthlett s.n. (holotype CHR). 


Beitr. 7: 95 (1987). 

kirkii var. pilosus J.-P.Frahm, Lindbergia 7:30 

(1981). 


Beitr. 7: 95 (1987). 

kiusiuensis Broth. in Ihs., Nihon - Senrui Bunriu 

137, 1932. (*) 

kivuensis P.Varde & Thér., Bull. Soc. Bot. France 

87:352 (1940) 

T: Congo bel. Kivu, volcan Ninagongo 2900- 

2800 m Humbert 7651B (holotype PC). 

CU: ? C. johannis-meyeri (C. Müll.) Kindb. cf. 


kouroussensis Ren. & Par., Rev. Bryol. 30:66 

(1903) 

T: Guinea, Kouroussa, Pobeguin s.n. (holotype 

PC, isotype NY). 


Bibl. 31:139 (1985). 

krauseanus (Hampe & Lor.) Jaeg., Ber. S. Gall. 

Naturw. Ges. 1870-71:427 (1872). 

B: Dicranum krauseanum Hampe & Lorentz, 

Bot. Zeit. 27: 434. 1869. 

T: Ecuador, Krause s. n. (holotype not seen;

isotype GOET). 

OU: C. krausei Hampe & Lor. ex Kindb., 

Enum. Bryin. Exot. 88 (1889); C. chrismarii 

(C. Müll.) Mitt. cf. Williams, N. Am. Fl. 15:136 

(1913). 


kunertii Par., Ind. Bryol. Suppl. 93 (1900) nom. 

illeg. 

OU: C. pseudojulaceus C. Müll. 

CU: C. julaceus Jaeg. ssp. arbogastii (Ren. & 

Card.) J.-P. Frahm. 

kunkelii Bartr., Ark. Bot. ser.2,4:31 (1957). 

T: Juan Fernandez, Masafuera, Skottsberg 18 

(isoptype S). 

CU: C. pyriformis (Schultz) Brid. cf. Corley & 

Frahm, J. Bryol. 12:190 (1982). 

kurzii Hampe in C. Müll., Gen. Musc. Fr. 271 

(1900) nom. nud. 

L 

labeensis Broth. & Par., Rev. Bryol. 38: 25 

(1911). 

T: Guinea, Fouta Djallon, in jugo Labe, 

Pobeguin s.n. (isotype BR). 

CU: C. cambouei Ren. & Card. cf. Frahm, 

Bryoph. Bibl. 31:45 (1985). 

lacouturei Thér. in Broth., Nat. Pfl. ed.2,10:187 

(1924) nom. nud. 

laete-virens (Broth.) J.-P.Frahm, Revue Bryol. 

Lich. 41:328 (1975) hom. illeg. 

B: Thysanomitrion laete-virens Broth., Acta 

Hort. Gothenbzurg 1: 189 (1924). 

T: Chile Prov. Concepión, prope Talcahuano, 

Skottsberg 62 (lectotype H-BR). 

CU: C. laxoventralis Herz. ex J.-P. Frahm, 

Bryol. Beitr. 7: 45 (1987). 

laete-virens (C. Müll.) Jaeg., Ber. S. Gall. 

Naturw. Ges. 1870-71:424 (1872). 

B: Dicranum laete-virens C. Müll., Bot. Zeit. 

20: 12 (1862). 

CU: C. fragilis (Brid.) B.S.G. cf. Corbiere, 

Mem. Soc. Bot. France 7:8 (1907). 

laetus (Mitt.) Jaeg., Ber. S. Gall. Naturw. Ges. 

1870-71:417 (1872). 

B: Dicranum laetum Mitt., J. Linn. Soc. Bot. 

Suppl. 1: 19 (1859). 

T: East Bengal, Khasia Mtns., Hooker & 

Taylor 84 (holotype NY). 


Frahm, J. Bryol. 13:179 (1984). 

51 

laevigatus (C. Müll.) Schimp. in Besch., Mem. 

Soc. Sc. Nat. Cherbourg 16:167 (1872) hom. 

illeg. 

B: Dicranum laevigatum C. Müll., Syn. Musc. 

II: 601 (1851). 

T: Mexico, Liebmann s. n. (holotype 

destroyed at B; lectotype H). 

CU: C. richardii Brid. cf. Dixon, J. Bot. 

60:287 (1922). 

OU: C. umbellatus (Arn.) Par. cf. Williams, 

N. Am. Fl. 15:14 (1913). 

laevigatus Brid., Mant. Musc. 76 (1819). 

CU: Grimmia laevigata (Brid.) Brid. 

laevis Tayl., London J. Bot. 5:47 (1846). 

T: Ecuador, Mt. Pichincha, Jameson (FH). 

OU: Pilopogonella laevis (Tayl.) Bartr. 

CU: Pilopogon laevis (Tayl.) Thér. 

lamellatus Mont., Ann. Sc. Nat. Bot. ser. 2,9:52 

(1888). 

T: Bolivia (holotype PC; isotype BM). 

CU: C. pilifer Brid. var. lamellatus (Mont.) 

Gradstein & Sipman. 


J. Linn. Soc. Bot. 12:84 (1869). 

lamellicosta (C. Müll.) Schimp. in Besch., Mem. 

Soc. Sc. Nat. Cherbourg 16:167 (1872). 

B: Dicranum lamellicosta C. Müll., Syn. 

Musc. 2: 601 (1851). 


N. Am. Fl. 15:144 (1913). 


lamellinervis (C. Müll.) Mitt., J. Linn. Soc. Bot. 

12:82. (1869). 

B: Dicranum lamellinerve C. Müll., Syn. 

Musc. 1:390 (1848). 

T: Jamaica, Wilson s. n. (holotype destroyed 

at B; isotypes could not be located). 

lamellinervis (C. Müll.) Mitt. var. exaltatus 

(C.Müll.) J.P. Frahm, Bryoph. Bibl. 5: 78 

(1975). 

B: Dicranum exaltatum C. Müll., Linnaea 42: 

472 (1879). 

OU: Campylopus exaltatus 

lamprodictyon (Hampe) Mitt., J. Linn. Soc. Bot. 

12:78 (1869). 

B: Dicranum lamprodictyon Hampe, Ann. Sc. 

Nat. Bot. ser. 5,5: 339 (1866). 

T: Colombia, Bogotá La Penna, Lindig s. n. 

(holotype, BM; isotype S). 

CU: C. pauper (Hampe) Mitt. var.

52 

lamprodictyon (Hampe) J.-P. Frahm, Bryol. 

Beitr. 7: 96 (1987). 

laniger (C. Müll.) Besch., Bull. Soc. Bot. France 

32:LVI (1885). 

CU: Chorisodontium lanigerum (C. Müll.) 

Broth. 

lapidicola (C. Müll.) Par., Ind. Bryol. ed.2,1:315 

(1904). 

B: Dicranum lapidicola C. Müll., Hedwigia 

39: 358 (1900). 

T: Brazil. S. Catarina, Orleans, Ule 701 


OU: C. griseus var. lapidicola (C. Müll.) J.- 

P.Frahm, Rev. Bryol. Lich. 42:606 (1976). 

CU: C. griseus s.str. 

latifolius Card., Rev. Bryol. 36:16 (1909). 

T: Congo belg., Hens s.n. (holotype PC). 

CU: C. perpusillus Mitt. cf. Frahm, Bryph. 

Bibl. 31:139 (1985). 

latifolius Rehm. in Kindb., Enum. Bryin. Exot. 9 

(1888) nom. nud. 

M: Transvaal, in uliginosis montis Lechlaba, 

Rehmann 448 (PC). 


Bryoph. Bibl. 31:71 (1985). 

OU: C. trichodes Lor. (= C. pilifer Brid.) cf. 

Sim, Trans. R. Soc. S. Sfr. 15:168 (1926). 

latinervis (Mitt.) Jaeg., Ber. S. Gall. Naturw. Ges. 

1870-71:426 (1872). 

B: Dicranum latinerve Mitt., J. Linn. Soc. Bot. 

Suppl. 1: 17 (1859). 

T: Sikkim, Hooker 80. 

OU: C. gracilis cf. Frahm, J. Bryol. 13:176 

(1984). 

latinervis Herz., Bibl. Bot. 87:19 (1916) hom. 

illeg. 

CU: C. renneri Herz. 

latipilis C. Müll. In Kindb., Enum. Bryin. Exot. 

106 (1891) nom. nud. 

M: Chile, Valdivia, Han s.n. (H-BR). 



latolaminatus C. Müll. in Broth., Nat. Pfl. 

1(3):334 (1901) nom. nud. 



laureri Lor. in C. Müll., Gen. Musc. Fr. 266 

(1900) nom. nud. 

lavardei Thér. in P. Varde, Bull. Soc. Bot. France 

71:1055 (1924). 

T: Haute Forêt du Kilima-Njaro, no. 916 



Bryol. Beitr. 7: 46 (1987). 

laxiretis Bartr., Ark. Bot. ser.2,4:30 (1957) hom. 

illeg. 

OU: C. masafuerae J.-P.Frahm nom. nov., C. 

blindioides Broth. in Skottsb. cf. Robinson, 

Smiths. Contr. Bot. 27:26 (1975). 


Bryol. Beitr. 7: 46 (1987). 

laxiretis Herz., Beih. Bot. Centralbl. 26(2):54 ( 

). 

T: Bolivia, bei Incacorral Prov. Cochabamba, 

Herzog 15 (isotype L). 

OU: Atractylocarpus spec. cf. Frahm, J. Hatt. 

Bot. Lab. 44:484 (1978). 

CU: Atractylocarpus longisetus cf. Padberg 

& Frahm, Cryptogamie Bryol. Lich. 6:326 

(1985). 

laxitextus Sande Lac., Natuurk. Ver. Kon. 

Wetensch. Akad. Amsterdam 13: 10 (1872). 

T: Java, Lacoste s.n. hb. Dozy & Molkenboer 

(holotype L, isotypes B, BM, W). 

laxobasis Ren. & Card., Bull. Soc. R. Bot. Belg. 

35(1):299 (1897). 

T: Madagaskar, Imerina, Ambatomanga, 

Camboue s.n. (holotype PC). 


Bryoph. Bibl. 31:45 (1985). 

laxoventralis Herz. ex J.-P.Frahm, Nova 

Hedwigia 31:428 (1979). 

T: Nordchile: Fray George, Tigre Zapallar, 

Schwabe 232 (holotype JE). 

leanus (Sull.) Sull. & Lesq., Musci Bor. Am. 18 

(1859). 

CU: Brothera leana (Sull.) C. Müll. 

lechleri Schimp. in Lor., Bot. Zeit. 24:186 (1866) 

nom. nud. 


Bryoph. Bibl. 5:80 (1975). 

lecomtei Besch. in P. Varde, Rev. Bryol. Lich. 

7:231 (1935) nom. nud. 

leioneuron Thér. & P. Varde, Rev. Bryol. 491: 

40 (1922). 

T: India, Maduré, André 475 (holotype PC). 

CU: C. involutus (C. Müll.) Jaeg. cf. Frahm, 


lenormandii Thér., Bull. Soc. Bot. Geneve 26:76 

(1936).

T: Australia, Mt. Macedon, F. Müller ex hb. 

Lenormand nr. 19 (isotype H-BR). 

OU: C. pallidus cf. Frahm, Cryptogamie 

Bryol. Lich. 3(1):64 (1982). 


lepidophyllus (C. Müll.) Jaeg., Ber. S. Gall. 

Naturw. Ges. 1870-71:435 (1872). 

B: Dicranum lepidophyllum C. Müll., Syn. 

Musc. 407 (1848). 

T: Cape, Palmiet River, Ecklon s.n. (BM). 


Revue Bryol. Lich. 41(3):321 (1975). 

leptocarpus Wils. in Hook.f., Fl. Nov. Zel. 2:68 

(1854) nom. nud. 

OU: C. leptodus Mont. cf. Wilson l.c. 



leptocephalus (C. Müll.) Jaeg., Ber. S. Gall. 

Naturw. Ges. 1870-71:435 (1872). 

B: Dicranum leptocephalum C. Müll., Bot. 

Zeit. 9: 551 (1851). 

T: New Zealand, between Waitenabe and 

Kaipora, Mossman 726 (NY). 

CU: C. introflexus (Hedw.) Brid. cf. Mitten, 

J. Linn. Soc. Bot. 12:84 (1869). 

leptodictyon Broth., Denkschr. Ak. Wiss. Wien 

math. nat. Kl. 83:259 (1926). 

T: Brazil, Glaziou 11744 (lectotype, H-BR). 

CU: C. angustiretis Sull. & Lesq. cf. Frahm, 


leptodrepanium (Broth.) Broth., Nat. Pfl. 

1(3):332 (1901). 

B: Dicanodontium leptodrepannium C. Müll. 

ex Broth. Bot. Jahrb. 24: 233 (1897). 

T: Kamerun, Buea 2425 m, Dusen 348 

(holotype H-BR, isotype G). 

OU: C. stramineus cf. Bizot & Kilbertus, 

Revue Bryol. Lich. 45:90 (1979); C. 

chrismarii cf. Frahm, Lindbergia 8:46 (1982). 


leptodus Mont., Ann. Sc. Nat. Bot. ser. 3,4:111 

(1845). 

T: Chili, Valdivia (holotype PC). 



leptoneuron Broth. ex Ihs., Cat. Moss. Japan 32 

(1929). 

T: Japan, Nagasaki, Hirotsu 1486 (PC). 



53 

leptotrichaceus (C. Müll.) Par., Ind. Bryol. Suppl. 

93 (1900). 

B: Dicranum leptotrichaceum C. Müll., 

Hedwigia 38: 34 (1899). 

T: Südafrika, in sylvis Knysnae, Rehmann 33 

(PC). 


Bryoph. Bibl. 31:161 (1985). 

OU: C. atroluteus cf. Magill, Bryoph. S. Afr. 

1(1):148 (1981). 

letestui Thér. & P. Varde, Bull. Soc. Bot. France 

74:146 (1927). 

T: Gabun, Moucoussa, Le Testu 5363 




leucobasis Herz., Bibl. Bot. 87:23 (1916) nom. 

invalid. 

OU: C. insignis Herz. cf. Herz. l.c. 


leucobasis Par. Ind. Bryol. 93 (1900) hom. illeg. 

T: „Prom. bon. spei, in monte Boschberg 

prope Somerset-East“ Mac Owan s.n. 

Bryotheca europaea 1359 (isotype L). 

OU: C. introflexus (Hedw.) Brid. cf. Dixon, 



31:143 (1985). 

leucochlorus (C. Müll.) Par., Ind. Bryol. 253 

(1894). 

B: Dicranum leucochlorum C. Müll. Flora 71: 

411 (1888). 

T: Kilimand- scharo, ad finem sylvestrum 

superiorem inter 3000-4000 m. Meyer s.n. 

(PC) 

CU: C. hildebrandtii (C. Müll.) Jaeg. cf. 

Frahm, Trop. Bryol. 8: 157, 1993. 

leucogaster (C. Müll.) Mitt., J. Linn. Soc. Bot. 

12:75 (1869). 

B: Dicranum leucogaster C. Müll., Syn. 

Musc. 1: 387 (1848). 

T: Mexico, Jalapa, Deppe & Schiede s.n. 

(NY). 

OU: Campylopodium pusillum cf. Frahm, 




leucognodes (C. Müll.) Par. Ind. Bryol. Suppl. 

93 (1900). 

B: Dicranum leucognodes C.Müll., Nuovo

54 

Giorn. Bot. Ital. n. ser. 4: 32. (1897). 

T: Bolivia, Prov. Cochabamba, Germain 1105 

(lectotype H-BR; isolectotype NY). 

CU: C. argyrocaulon 

leucophaeus Stirt., Ann. Scott. Nat. Hist. 12:110 

(1903). 

CU: C. flexuosus (Hedw.) Brid. var. 

paradoxus (Wils.) Husn. cf. Hag., K. Norsk. 

Vid. Selsk. Skrift. 1914(1):75 (1915). 

leucopteryx C. Müll., Gen. Musc. Fr. 276 (1900) 

nom. nud. 

leucotrichus Sull. & Lesq., Musci Bor. Am. 17 

(1856). 

T: Alabama, Racoon montes, Musci amer. 

bor. 73 (NY). 


J. Linn. Soc. Bot. 12:84 (1869). 


levieri Broth. in Par., Ind. Bryol. Suppl. 92 (1900) 

nom. nud. 

CU: C. hensii Ren. & Card. cf. Frahm, 

Bryoph. Bibl. 31:87 (1985). 

liebmannii (C. Müll.) Schimp. in Besch., Mem. 

Soc. Sc. Nat. Cherbourg 16:167 (1872). 

B: Dicranum liebmannii C. Müll., Syn. Musc. 

2: 601 (1851). 

T: Mexico, Hahn s. n. (holotype destroyed at 

B; lectotype H-BR). 

HS: Pilopogon liebmannii (C. Müll.) Broth., 

Nat. Pfl. 1(3):336 (1901). 


N. Am. Fl. 15:144 (1913) = C. pilifer Brid. 

liliputanus (C. Müll.) Broth., Rev. Bryol. 47:3 

(1920). 

B: Pilopogon liliputanus C. Müll. 

T: Bolivia, prope Choquecamata, Germain 

1107 (NY). 

OU: Pilopogon gracilis (Hook.) Brid. cf. 


CU: Pilopogon guadeloupensis. 

limbatulus Broth., Bull. Soc. Bot. Ital. 1904:15 

(1904) nom. nud. 

lindavii Broth. in Engl., Pflanzenw. Ostafrika C 

68 (1895) nom. nud. 

T: Runssoro, Erica Wald 3000-3600 m, Stuhlmann 

3290 (isotype B ). 

CU: C. dicranoides Thér. & Nav. cf. Frahm, 

Bryoph. Bibl. 31:64 (1985). 

lonchochaete C. Müll., Abh. Naturw. Ver. Bremen 

16:495 (1900). 

T: New Zealand, Waiotapu, n.164 (lectotype 

H-BR, 172, 224, syntypes H-BR). 

OU: C. holomitrius auct. non C. capillaceus 

Hook.f. & Wils. cf. Dixon, New Zealand Inst. 

Bull. 3(3):85 (1923). 

lonchocladus C. Müll. in Geh. Rev. Bryol . 5:59 

(1878). 

T: „Insula Mauritius, Robillard leg. 1876“ (B, 

holotype). 



longescens (C. Müll.) Par., Ind. Bryol. Suppl. 93 

(1900). 

B: Dicranum longescens C. Müll. Hedwigia 

38: 82 (1889). 

T: Südafrika, Knysna, Rehmann 41 (holotype 

B, isotype L). 

OU:=?= C. echinatus Sim cf. Sim, Trans. R. 

Soc. S. Afr. 15:175 (1926); C. introflexus 

(Hedw.) Brid. cf. Dixon & Gepp, Bull. Misc. 

Inf. Kew 1923:198 (1923); C. inchangae (C. 

Müll.) Par. cf. Magill, Fl. S. Afr. Bryoph. 

1(1):146 (1981). 


Bibl. 31: 164 (1985). 

longicellularis J.-P. Frahm, Cryptogamie Bryol. 

Lich. 7(4):443 (1986). 

T: Colombia, Meta, Páramo de Sumapaz, 

Cleef 777a (holotype U) 

longifolius (Hedw.) Kindb., Bih. K. Svensk. Vet. 

Ak. Handl. 9(7):87 (1883) hom. illeg. 

B: Dicranum longifolium Ehrh. ex Hedew., 

Spec. Musc. 130 (1801). 


Loeske. 

longifolius Schimp. ex Besch., Ann. Sc. Nat. Bot. 

ser. 6,9:323 (1880). 

T: Reunion, Plaine des Chicots, Bory (BR). 

CU: ? C. arcuatus (Brid.) Jaeg. cf. Frahm, 

Bryoph. Bibl. 31:20 (1985). 

longigemmatus Gao in Gao, Zhang & Cao, Acta 

Bot. Yunn. 3:392 (1981). 

T: China, Xizang, Distr. Cona Xian, ad 

lignum cariosum 2900 m, Chen Shu-kun 

5152a (holotype ISPH, Isotypus HKAS). 

CU: Dicranodontium spec. cf. Frahm, Nova 

Hedwigia 41:273 (1985). 

longipilus Brid., Bryol. Univ. 1:477 (1826) nom. 

illeg. 

CU: C. introflexus (Hedw.) Brid. & C.

atrovirens De Not. cf. Grout, Moss Fl. N. Am. 

1:93 (1937). 

longipilus var. elongatus (C. Müll.) Schimp. in 

Jaeg., Ber. S. Gall. Naturw. Ges. 1870- 

71:437 (1872). 

OU: C. introflexus (Hedw.) Brid. fo. el. 

Podp., Consp. 131 (1954). 


longipilus var. falcatus (Braithw.) Par., Ind. 

Bryol. 254 (1894). 

CU: C. atrovirens De Not. fo. falcatus 

Podp., Consp. 130 (1954). 

longipilus var. muticus Milde, Bot. Zeit. 28:396 

(1870). 

CU: C. atrovirens De Not. fo. muticus 

Moenk, Laubm. Eur. 225 (1927). 

longirostris (Web. & Mohr) Spruce, Musci Pyren. 

202 (1847). 


Britt. cf. Williams, N. Am. Fl. 15:151 (1913). 

longisubulatus Thér., Rev. Bryol. Lich. 11: 49 

(1939). 

T: Bolivia, Tablas 1800 m, Herzog 4566 

(holotype PC; isotypes B, JE) 

CU: C. anderssonii (C. Müll.) Jaeg. cf. Frahm, 

Bryol. Beitr. 7: 49, 1987. 

lorentzii (Fleisch.) W. Schultze-Motel, 

Willdenowia 3: 407 (1963). 

B: Pilopogon lorentzii Fleisch., Nov. Guinea 

12(2): 13 81914). 

T: Holländisch-Guinea, In den Gipfelwäldern 

des Hellwiggebirges.. v. Römer 1261 

(holotype FH, isotype S). 

OU: Thysanomitrion lorentzii (Fleisch.) 

Fleisch., Hedwigia 50: 280 (1911). 

CU: C. exasperatus var. lorentzii (Fleisch.) 

J.-P. Frahm et al., Acta bot. Fenn. 131: 80, 

1985. 

lorentzii C. Müll. ex Par., Ind. Bryol. 254 (1894) 

err. pro Conostomum lorentzii C. Müll. 

ludovicae Broth. & Par., Oefv. Finsk. Vet. Soc. 

Foerh. 51A(17):3 (1909). 

OU: C. polyanthus Besch. cf. Thér., Rev. 

Bryol. 48:58 (1921). 

CU: C. laxitextus Lac. 

luetzelburgii Herz. ex J.-P.Frahm, Cryptogamie 

Bryol. Lich. 2(4):442 (1981). 

T: Brazil, Serra do Itatiaia, Agulhas Negras, 

Dusén s. no. (holotype, B). 


55 

fragiliformis (J.-P. Frahm) J.-P. Frahm cf. 

Frahm, Fl. Neotrop. 54: 101 (1991). 

luluensis Thér. & Nav., Rev. Bryol. Lich. 12:68 

(1942). 

T: Congo, Auverlaet oct. 1923 (BR). 

CU: C. savannarum (C. Müll.) Mitt. cf. ???? 

luridus Schimp. ex Besch., Mem. Soc. Sc. Nat. 

Cherbourg 16:168 (1872). 

T: Mexico, Orizaba, F. Müller s. n. (not seen). 


N. Am. Fl. 15(2):144 (1913). 


luscinialis Dix in Christ., Res. Norw. Sci. Exp. 

Trista da Cunha 48:19 (1960). 

T: Tristan da Cunha, Nightingale, 

Christophersen 2127,pp. (isotype FH). 


Bryoph. Bibl. 31:160 (1985). 

lutescens (C. Müll.) Jaeg., Ber. S. Gall. Naturw. 

Ges. 1870-71:440 (1872). 

B: Dicranum lutescens C. Müll., Syn. Musc. 

2: 602 (1851). 

T: Mexico, Orizaba, Liebmann s.n. 


J. Linn. Soc. Bot. 12:84 (1869). 


luteus (C. Müll.) Par., Ind. Bryol. 254 (1894). 

B: Thysanomitrion luteum C. Müll., Linnaea 

42: 470 (1879). 

T: Venezuela, Tovar, Fendler 39 (holotype 

destroyed at B; lectotype H-BR; isotype H- 

SOL). 

M 

macgregorii Broth. & Geh., Oefv. Finska Bet. 

Akad. Förh. 37: 155 (1895). 

OU: Thysanomitrion macgregorii Broth., Nat. 

Pflanzenfam. Ed 2, 10: 189 (1924). 

T: Papua New Guinea, Owen Stanley Range, 

13000 ft., Macgregor 335 (holotype H-BR, 


maclaudii Par. & Broth., Rev. Bryol. 30:66 

(1903). 

T: Guinea, Fouta Djallon, rochers au bord du 

Bafing, Maclaud s.n. 1901 (B, PC). 

HS: macleaudii Par. ex Broth., Nat. Pfl. 

1(3):1186 (1909). 



macleanus Rehm. in Thér., Rev. Bryol. n.ser. 2:75

56 

(1931) nom. invalid. 

T: Transvaal,in silvis monte Kwatlamba supra 

Pilgimsrest ad rupes, MacLea 447 (PC). 

HS: macleai Rehm. in Kindb., Enum. Bryin. 

Exot. 88 (1889) nom. nud. 




Afr. Bryoph. 1(1):146 (1981). 


Bibl. 31:164 (1985). 

macrogaster (C. Müll.) Broth., Nat. Pfl. 1(3):334 

(1901). 

B: Dicranum macrogaster C. Müll., Hedwigia 

39: 252 (1900). 

T: Brazil. S. Catarina, Serra Geral, Ule 1123 

(holotype destroyed at B; lectotype H- 

BR).OU: C. controversus (Hampe) Jaeg. cf. 


CU: C. thwaitesii. 

macrophyllus (C. Müll.) Mitt., J. Linn. Soc. Bot. 

12:80 (1869). 

T: Dicranum macrophyllum C. Müll., Syn. 

Musc. I: 402 (1848). 

T: Peru, Cuchero, Poeppig 213 (holotype 

destroyed at B; lectotype NY; isolectotype 

KIEL). 

OU: C. filifolius (Hornsch.) Mitt. cf. Frahm, 



P.Frahm. 

macrotis C. Müll. ex Broth., Bot. Jahrb. 24:233 

(1897). 

T: Kamerun, Wervoka 700 m alt., Dusen 699 

(holotype B). 



madagassus Dix., J. Bot. 80:43 (1942). 

T: Madagascar, circa 1814 Thompson 22 

(holotype BM). 



Bryoph. Bibl. 31:16 (1985). 

madecassus Besch., Ann. Sc. Nat. Bot. ser. 

6,9:321 (1880). 

T: Madagascar, Bernier s.n. (holotype PC, 

isotype B). 


madecassus (Besch.) J.-P.Frahm, Bryoph. 

Bibl. 31:16 (1985). 

madeirensis Geh. in Kindb., Enum. Bryin. Exot. 

9 (1888) nom. nud. 

CU: Ditrichum punctulatum Mitt. cf. Seppelt 

in Frahm, Bryoph. Bibl. 31:197 (1985). 

CU: C. brevipilus B.S.G. var. marginatulus 

(Geh.) Thér. cf. Luisier, Broteria Cienc. Nat. 

24:28 (1930). 

magniretis (C. Müll.) Par., Ind. Bryol. Suppl. 94 

(1900). 

B: Dicranum magnirete C. Müll., Bull. Herb. 

Boiss. 5: 186 (1897). 

T: Guatemala, Alta Vera Paz, in trunco, v. 

Türckheim s.n. 

OU: Campylopodium pusillum (Schimp.) 

Williams cf. Williams, N. Am. Fl. 15:91 

(1913). 

OU: Microcampylopus curvisetus (Hampe) 


malagensis Herz., Bibl. Bot. 87:23 (1916). 

T: Bolivien, Cerro de Malaga, Herzog 4367 

(holotype JE, isotype B). 

CU: Campylopodiella malagensis (Herz.) J.- 

P.Frahm, The Bryologist 87:250 (1984). 

mandoni Schimp. in Par., Ind. Bryol. 254 (1894) 

nom. nud. 

M: Mandon, Pl. Boliv. 1610 (GOET) 

CU: = C. nivalis (Brid.) Brid. var. 

multicapsularis (C. Müll.) J.-P. Frahm cf. 

Frahm, Nova Hedwigia (1994). 

mangenotii Biz. & Kilb., Rev. Bryol. Lich. 

4o:113 (1974) nom. invalid. 

T: Reunion, Piton des Neiges, Cadet 162 (hb. 

Bizot and Kilbertus). 

OU: C. parisii Ren. cf. Frahm, Bryoph. Bibl. 

31:195 (1985). 

CU: C. smaragdinus (Brid.) Jaeg. 

manginii Thér., Rec. Publ. Soc. Havraise Etud. 

Div. 89(2):113 (1922). 

T: Madagascar,massif d’Adringitra, Perrier 

de la Bathie s.n. (isotypes G, NY). 

OU: C. stramineus Mitt. cf. Bizot & Kilbertus, 

Rev. Bryol. Lich. 45:79 (1979). 


Bryoph. Bibl. 31:124 (1985). 

marginatulus Geh., Bibl. Bot. 73:12 (1910). 

T: Madeira, Fritze (1880) s.n. (isotype PC). 

CU: Ditrichum puntulatum Mitt. cf. Seppelt 

in Frahm, Bryoph. Bibl. 31:197 (1985). 

OU: C. brevipilus B.S.G. var. marginatulus 

(Geh.) Thér.

marillacii Rehm. in Par., Ind. Bryol. 254 (1894) 

nom. nud. 

T: Montagu Pass, Rehmann 65 (B,L). 


Bryoph. Bibl.31:71 (1985). 

marmellensis Broth., Hedwigia 45:262 (1906). 

T: Brazil, Amazonas, Rio Madeira, Ule 246 

(lectotype H-BR; isolectotypes FH, JE, NY). 

CU: C. surinamensis C. Müll. cf. Frahm, Revue 

Bryol. Lich. 45:163 (1979). 

martinae (martensi) Watts. & Whitel., Proc. Linn. 

Soc. N.S.Wales Suppl. 27:45 (1902) nom. 

nud. 

masafuerae Frahm, Bibl. Bot. 5:83 (1975) nom. 

nov. pro C. laxiretis Bartr. hom. illeg. 

OU: C. blindioides Broth. in Skottsb. cf. 

Robinson, Smiths. Contr. Bot. 27:26 (1975). 


Bryol. Beitr. 7: 51 (1987). 

matarensis Besch., Ann. Sc. Nat. Bot. ser. 6,9:323 

(1880). 

T: Reunion, Cilaos, Valentin s.n. (holotype 

PC). 

CU: C. arctocarpus ssp. madecassus (Besch.) 

J.-P.Frahm cf. Frahm, Bryoph. Bibl. 31:16 

(1985). 

mathieui Thér. Bull. Mens. Soc. Linn. Seine 

Marit. 12:239 (1925). 

T: Kamerun, Mathieu s.n.(isotype BR). 



megalotus Besch. cf. Frahm, Bryoph. Bibl. 

31:118 (1985). 

T: Ile St. Paul, sterile, dans les touffes de 

Sphagnum, G. de Isle 16 (holotype PC). 

melaphanus Stirt., Ann. Scott. Nat. Hist. 12:110 

(1903). 

CU: C. flexuosus (Hedw.) Brid. cf. Dixon, J. 

Bot. 61:15 (1923). 

merapicola C. Müll. Gen. Musc. Fr. 266 (1900) 

nom. nud. 

OU: C. aureus Bosch & Lac. cf. Fleischer, 



metzlerelloides (P. Varde & Thér.) Biz. in Biz. 

& Pocs, Acta Bot. Acad. Sci. Hung. 25:246 

(1979). 

B: Bryohumbertia metzlerelloides P. Varde 

& Thér., Bull. Soc. Bot. France 86: 422 

(1939). 

57 

OU: C. flavicoma C. Müll. ex Broth. cf. De 

Sloover, Bull. Jard. Bot. Nat. Belg. 51:227 

(1981). 

CU: Bryohumbertia flavicoma (C. Müll.) J.- 


(1982) 

mexicanus Thér., Smiths. Misc. Coll. 78:7 (1926). 

T: Mexico, Campanario, Arséne 7576 




micans Wulfsb., Forh. Vid. Selsk. Christiana 

1876:349 (1876). 

CU: C. flexuosus (Hedw.) Brid. var. micans 

(Wulfsb.) Hag. 

micholitzii C. Müll. ex Fleisch., Musci Fl. 


T: Sumatra, Tindjoelaut 3000p., Mieholitz 61 

(B, BM, FH). 

CU: C. laxitextus Lac. cf. Eddy, Malesian 

Mosses vol. 1 (1988). 

microcephalus (C. Müll.) Par., Ind. Bryol. Suppl. 

94 (1900). 

B: Dicranum microcephalum C. Müll., Flora 

82: 443 (1896). 

T: Hawaii, without locality, Hillebrand s.n. 

(not seen). 

CU: C. praemorsus cf. Frahm, Nova 

Hedwigia 39:597 (1984). 

OU: C. exasperatus cf. Bartr., Bishop Mus. 

Bull. 101:45 (1933). 

microjulaceus (C. Müll.) Par., Ind. Bryol. Suppl. 

94 (1900). 

B: Dicranum micro-julaceum C. Müll., Bull. 

Herb. Boiss. VI: 37 (1898). 

T: Brazil, Serra do Itatiaia, Ule 1796 

(holotype destroyed at B; isotype H-BR). 

CU: C. julaceus Jaeg. cf. Frahm, Bryol. Beitr. 

7: 52 (1987). 

microrhodon C. Müll. in Par., Ind. Bryol. Suppl. 

94 (1900) nom. nud. 

M: Sikkim-Himalaya, prope Kurseong 

1000m, Bretaudeau 16931 (PC). 



OU: C. introflexus (Hedw.) Brid. cf. P. Varde, 

Arch. Bot. 1:178 (1927). 

microstomus Mitt. in Spruce, Cat. Musc. Amaz. 

And. 2 (1867) nom. nud. 

OU: C. porphyreodictyon (C. Müll.) Mitt. cf.

58 

Mitten, J. Linn. Soc. Bot. 12:75 (1869). 

OU: C. filifolius (Hornsch.) Mitt. = 

Bryohumbertia filifolia (Hornsch.) J.-P.Frahm 

microtheca Herz., Beih. Bot. Centralbl. 26:55 

(1910). 

T: Bolivia, Cerro Amboro, Herzog 271 

(holotype JE; isotype B). 



mildei Limpr., Laubm. Deutschl. 1:396 (1887). 

T: Italy, ad una ripa lungo la strada dalla 

Madonna diSantino a Bieno, in val Intrasca, 

Lago Maggiore, De Notaris(1866) (lectotype 

BP). 

CU: C. oerstedianus cf. Frahm, J. Bryol. 

11:216 (1980). 

milleri Ren. & Card., Bull. Soc. R. Bot. Belg. 

41:17 (1905). 

minarum Par., Ind. Bryol. ed. 2,1: 318 (1904), 

nom. nud. 

M: the same as in the type of Dicranum 

laxobasis C. Müll., Hedwigia 39: 252. 1900, 

Brazil, Minas Gerais, Serra de Caraca, Ule 

1353 (holotype destroyed at B; lectotype H- 

BR) 



minor Besch., Compt. Rend. Ac. Sc. Paris 81:723 

(1875). 

T: Ile Amsterdam, G. del’ Isle (1874-75) s.n. 



Bryoph. Bibl. 31:160 (1985). 

minutus Lindb. in Aongstr., Oefv. K. vet. Ak. 

Foerh. 33:5 (1876) nom. nud. 

mittenii (C. Müll.) Jaeg., Ber. S. Gall. Naturw. 

Ges. 1870-71:433 (1872). 

B: Dicranum mittenii C, Müll., Syn. Musc. 

2: 600 (1851). 

T: Ecuador, prope Quito, Jameson s.n. 

(lectotype BM, isolectotypes FH, NY). 

CU: Chorisodontium mittenii (C. Müll.) 

Broth. 

modestus Card., Bull. Herb. Boiss. ser.2,5:1000 

(1905). 

T: Falkland Isl., Port Standley, Skottsberg 218 

(isotype BM). 

molkenboeri Lac. in Milde, Bryol. Siles. 79 

(1869) nom. nud. 

CU: C. brevipilus B.S.G. var. molkenboeri 

Milde 

mollis C. Müll. in Par., Ind. Bryol. 255 (1894) 

nom. nud. 

M: Natal, Inanda, Rehmann 36 (PC, B). 

OU: C. stenopelma (C. Müll.) Rehm. ex Par. 



montagnei De Not., Mem. R. Acc. Sc. Torino 

ser.2,18:452 (1895). 

T: Chili, Valparaiso, Puccio s.n. (isotype H- 

SOL). 

CU: C. clavatus (R.Brown) Wils. cf. Frahm, 


montanus Welw. & Duby, Mem. Soc. Phys. Hist. 

Nat. Geneve 21:221 (1870). 

T: Angola, Distr. Huilla, Welwitsch 3 

(lectotypus nov. G, isotypes BM,PC). 

CU: savannarum (C. Müll.) Mitt. cf. Frahm, 

Bryoph. Bibl. 31:171 (1985). 

morenoi (C. Müll.) Par., Ind. Bryol. Suppl. 94 

(1900). 

B: Dicranum morenoi C. Müll., Hedwigia 36: 

97 (1897). 

T: Patagonia 50° 53´, Moreno (1880) s.n. 

(holotype B) 

CU: Dicranum spec. cf. Frahm, Bryol. Beitr. 

7: 95 (1987). 

morobensis Bartr., Bryologist 48:111 (1945). 

T: Papua New Guinea, Morobe Prov., Matap, 

Clemens 11290 (holotype FH). 

OU: Bryohumbertia walkeri (Mitt.) J.- 

P.Frahm cf. Frahm et al., Acta Bot. Fennica 

131:68 (1985). 



mosenii Broth., Bih. K. Svensk. Vet. Ak. Handl. 

21 Afd. 3(3):6 (1895). 

T: Brazil, Sao Paulo, Alto da Serra, Mosén 



Lich. 45:141 (1979). 


mouensis Broth. & Par., Oefvers. Foerh. Finska 

Vetensk.-Soc. 53A: 6 (1910). 

T: Nouvelle-Caledonie, Mou 1100 m, Le Rat 

1463 (H-BR). 

OU: C. pallidus cf. Frahm, Lindbergia 8:49 

(1982). 


mozambicus P.Varde, Rev. Bryol. Lich. 26:3

(1957). 

T: Mozambique near Gogogo Peak, 

Gorongosa Mtns. Schelpe 5554 (holotype 

PC). 


Bryoph. Bibl. 31:45 (1985). 

mucorinus C. Müll., Gen. Musc. Fr. 265 (1900) 

nom. nud. 

muelleri (Hampe) Jaeg., Ber. S. Gall. Naturw. 

Ges. 1870-71:442 (1872) hom. illeg. 

B: Thysanomitrion muelleri Hampe, Ann. Sc. 

Nat. Bot. ser. 5,3: 363 (1865). 

T: Colombia, Triana & Planchon s. no. (not 

seen). 

CU: C. richardii Brid. cf. Mitten, J. Linn. Soc. 

Bot. 12:90 (1869). 

muelleri Jur. in Rabenh., Bryoth. Eur. fasc.12 nr. 

578 (1862). 

CU: C. pyriformis (Schultz) Brid. mod. 

muelleri 

OU: C. fragilis (Brid.) B.S.G. var. muelleri 

(Jur.) Wijk & Marg., C. fragilis (Brid.) B.S.G. 

fo. muelleri (Jur.) Smirnova. 

muelleri Lor., Verh. Zool. Bot. Ges. Wien 17:678 


T: Type. Brazil, Puiggari 176 (not seen). 

CU: C. filifolius (Hornsch.) Mitt. cf. Hampe, 

Vid. Medd. Naturh. For. Kjoebenh. ser.4,1:91 

(1879). 

multicapsularis (C. Müll.) Schimp. in Par., Ind. 

Bryol. Suppl. 94 (1900). 

B: Dicranum multicapsulare C. Müll., Prodr. 

Fl. Boliv. 34 (1897). 

T: Bolivia, Larecaja, Mandon 1611 (holotype 

B; isotypes GOET, NY). 

CU: C. nivalis (Brid.) Brid. var. 

multicapsularis (C. Müll.) J.-P. Frahm. 

multisulcatus Duby in Hampe, Vid. Medd. 

Naturh. For. Kjoebenh. ser.3,2:272 (1870) 

nom. invalid. 

CU: C. lamellinervis (C. Müll.) Mitt. cf. 

Hampe l.c. 

muricatus Dix., K. Norsk. Vid. Selsk. Forh. 4:180 

(1932). 

CU: Chorisodontium aciphyllum cf. Dixon, 

Det Konigl. Norske Vidensk. Selsk. Foerh. 

7:1 (1934). 

N 

nadeaudianus Besch., Ann. Sc. Nat. Bot. ser. 

59 

7,20: 16 (1894). 

T: Tahiti, Mamano 200m, Nadeaud 98 



7: 54 (1987). 

nagasakinus Broth. in Par., Ind. Bryol. 94 (1900) 

nom. nud. 

M: Japan, Nagasaki, Faurié207 (PC). 



nakamurae Sak., Bot. Mag. Tokyo 55: 212 (1941) 

(*). 

nanocaudatus C. Müll., Gen. Musc. Fr. 274 

(1900) nom. nud. 

M: „Südgeorgien, Dr. Wilken 1883“ (GOET) 

CU: = Chorisodontium sp. cf. Frahm, Nova 

Hedwigia 59: 150, 1994. 

nano-exasperatus C. Müll., Gen. Musc. Fr. 271 

(1900) nom. nud. 

nano-filifolius (C. Müll.) Par., Ind. Bryol. Suppl. 

94 (1900). 

B: Dicranum nano-filifolium C. Müll., Nuov. 

Giorn. Bot. Ital. N.S. 4: 35 (1897). 

T: Brazil, Sta. Catarina, Ule 439 (lectotype, 

H-BR). 


Bryoph. Bibl. 5: 86 (1975). 

CU: Bryohumbertia filifolia (Hornsch.) J.-P. 

Frahm 

nanophyllus C. Müll. ex Broth., Bot. Jahrb. 24: 

235 (1897). 

T: „Ad victoriam emporium in saxis die 26 

m. julii a. 1891“ Dusen 305 (holotype B, 


nano-tenax (C. Müll.) Par., Ind. Bryol. Suppl. 94 

(1900). 

B: Dicranum nano-tenax C. Müll. Hedwigia 

38: 82 (1899I. 

T: Natal, Umpumulo, Borgen 80 (PC, L). 




Trans. R. Soc. S. Afr. 15: 178 (1926); C. 


Afr. Bryoph. 1(1): 146 (1981). 

nanus C. Müll., Bot. Zeitschr. 5: 804 (1847). 

CU: Dicranella subsubulata (C. Müll.) Jaeg. 

cf. Magill, Fl. S. Afr. Bryoph. 1(1): 121 

(1981). 

OU: Microcampylopus nanus (C. Müll.)

60 

Broth. 

natalensis Rehm. in Par., Ind. Bryol. 255 (1894) 

nom. nud. 

M: Natal, Van Reenen Pass, Rehmann 51 

(L,G). 




Trans. R. Soc. S. Afr. 15: 178 (1926). 

naumannii (C. Müll.) Kindb., Enum. Bryin. Exot. 

89 (1889) 

B: Dicranum naumannii C.Müll. Bot. Jahrb. 

5:84 (1883). 

T: Insula Ascension, Green Mount 2000 ft., 

Dr. Naumann (1874) s.n. (holotype B). 

CU: C. aureonitens (C. Müll.) Jaeg. cf. Frahm, 

Bryoph. Bibl. 31: 24 (1985). 

nematophyllus Ren. & Card., Bull. Soc. R. Bot. 

Belg. 31: 148 (1893) nom. invalid. 

T: Costa Rica, Forêt de Barba, 6.2.1890 


OU: C. hoffmannii (C. Müll.) Ren. & Card. 

cf. Renauld & Cardot l.c. 

CU: C. cuspidatus cf. Frahm, J. Hattori Bot. 

Lab. 44: 485 (1978). 

nepalensis (Brid.) Jaeg., Ber. S. Gall. Naturw. 

Ges. 1870-71:416 (1872) (*) 

nietneri (C. Müll.) Jaeg., Ber. S. Gall. Naturw. 

Ges. 1870-71: 432 (1872). 

T: „Ceylon, Nietner“ (lectotype GOET) 

OU: ?C. involutus cf. Frahm, J. Bryol. 13: 

176 (1984). 


(Mitt.) J.-P. Frahm 

nigerrimus (C. Müll.) Par., Ind. Bryol. Suppl. 94 

(1900). 

B: Thysanomitrion nigerrimum C. Müll., Bull. 

Herb. Boiss. 6: 39 (1898). 

T: Brazil, Rio de Janeiro, Tijuca, Ule 1081 


CU: C. richardii Brid. cf. Frahm, Nova 

Hedwigia 28: 133 (1976). 

nigrescens (Mitt.) Jaeg., Ber. S. Gall. Naturw. 

Ges. 1870-71: 417 (1872) hom. illeg. 

B: Dicranum nigrescens Mitt., J. Linn. 

Soc.Bot. Suppl. 1: 19 (1859). 

T: Sikkim, Hooker 73, 78, 82 (isosyntypes 

PC). 

HS: Thysanomitrium nigrescens (Mitt.) P. 

Varde, Rev. Bryol. 49: 40 (1922). 

CU: C. umbellatus (Arn.) Par., cf. Frahm, 


OU: C. richardii Brid. cf. Dixon, J. Bot. 60: 

290 (1922). 

nigrescens Duby, Mem. Soc. Phys. Hist. Nat. 

Geneve 19: 292 (1867). 

CU: C. richardii Brid. cf. Mitten, J. Linn. Soc. 

Bot. 12: 90 (1869). 

nigro-flavus (C. Müll.) Par., Ind. Bryol. Suppl. 

94 (1900). 

T: Australia occ., Mt. Kindsay, Webb s.n. 


CU: C. kirkii Mitt. cf. Frahm, Cryptogamie 

Bryol. Lichénol. 3: 64 (1982). 

nilghiriensis (Mitt.) Jaeg., Ber. S. Gall. Naturw. 

Ges. 1870-71: 426 (1872). 

B:Dicranum nilghiriense Mitt., J. Linn. Soc. 

Bot. Suppl. 1: 16 (1859). 

T: India, In mont. Nilghiri, Perrottet, Gardner 

(isotype H-BR). 

CU: C. goughii p. pte. cf. Frahm, J. Bryol. 

13: 172 (1984) = C. fragilis (Brid.) B.S.G. 

ssp. goughii (Mitt.) J.-P. Frahm, Trop. Bryol. 

4: 61, 1991. 

nitidus Dozy & Molk., Musci Fr. Ined. Arch. 

Indici 5: 139 (1847). 

OU: Dicranodontium nitidum (Dozy & 

Molk.) Fleisch., hom. illeg. 

nitidus Thér. in Felipp., Rev. Bryol. n.ser. 2: 212 

hom illeg. 

T: Uruguay, Maldonado, Felippone 1214 

(isotypes B, H-BR). 

OU: C. uruguensis J.-P. Frahm, Bryoph. Bibl. 

5: 127 (1975). 


Bryol. Beitr. 7: 55 (1987). 

nitidus Wils. in Mitt., Kew J. Bot. 3: 52 (1851) 

nom. nud. 

CU: C. areodictyon (C. Müll.) Mitt. cf. Mitten 

l.c. 

nivalis (Brid.) Brid., Bryol. Univ. 1: 477 (1926). 

B: Weisia nivalis Brid., Spec. Musc. 1:123 

(1806). 

T: Ile Bourbon, Bory St. Vincent s.n. (holotype 

B). 

OU: Thysanomitrion nivale (Brid) Arnot, 

Mem. Soc. linn. Paris 5: 263 (1827). 

nivalis var. multicapsularis (C. Müll.) Flora 

Neotropica 54: 139 (1991). 

B: Dicranum multicapsulare C. Müll., Nuov.

Giorn. Bot. Ital. n. ser. 4: 34 (1897). 

nodiflorus (C. Müll.) Jaeg., Ber. S. Gall. Naturw. 

Ges. 1870-71: 440 (1872). 

B. Dicranum nodiflorum C. Müll., Bot. Zeit. 

11: 38 (1853). 

T: India orientalis, Perrottet (GOET). 

OU: C. aureus Bosch & Lac. cf. Frahm, J. 

Bryol. 13: 165 (1984). 


novae-guineae Dix. & Thér., Farlowia 1: 28 

(1943). 

T: Indonesia, West Irian, Paniai, Dajakweide, 

ca. 4000 m, Mt. Carstenz. Wissel 128 

(BZ,NY). 

CU: C. austro-subulatus Broth. & Geh. cf. 

Frahm et al., Acta Bot. Fenn. 131: 72 (1985). 

novae-valesiae Broth., Oefv. Finsk. Vet. Soc. 

Foerh. 40: 163 (1898). 

T: Australia, Bulli Pass, N.S.W., Watts s.n. 


OU: C. pallidus cf. Frahm, Cryptogamie 

Bryol. Lichénol. 3: 64 (1982). 


novae-zealandiae Bartr. & Dix., Bot. Not. 1937: 

72 (1937). 

T: Insula austr. Novae Zelandiae, Bluff, 

Berggren 697 (BM). 


Bryol. Beitr. 7: 55 (1987). 

nudicaulis Besch., Bull. Soc. Bot. France 45: 55 

(1898). 

T: Tahiti, Nadeaud 208 (isotype BM). 


Bryol. Beitr. 7: 55 (1987). 

nudus (Hampe) Jaeg., Ber. S. Gall. Naturw. Ges. 

1870-71: 421 (1872). 

CU: Dicranoloma menziesii (Tayl.) Par. cf. 

Dixon, New Zealand Inst. Bull. 3: 364 (1929). 

O 

oblongus Thér., Rev. Bryol. Lich. 11:62 (1939). 

T: Colombia, above Bogotá 3000 - 3200 m, 

Troll 2082 (holotype, PC) 

obrutus Thér. & P. Varde Rev. Bryol. n. ser. 1: 

90 (1930). 

T: Gabun, Ngwasse, Mobounga, Le Testu 

2940 (holotype PC). 

obscurus Aongstr., Oefv. K. Svensk. Vet. Ak. 

Foerh. 30(5):119 (1873). 


61 


obtectus Stirt., Trans. Bot. Soc. Edinburgh 26:426 

(1915). 

CU: C. flexuosus (Hedw.) Brid. „var. 

paradoxus (Wils.) Husn.“ cf. Dixon, J. Bot. 

61:16 (1923). 

occultoviridis Dus., Rep. Princeton Univ. Exp. 

Patag. 8:121 (1903) nom. nud. 

occultus Mitt., J. Linn. Soc. Bot. 12: 86 (1869). 

T: Brazil, Paraná, Weir 66 (holotype,NY; 

isoytypes F, H-BR). 

ochrodictyon Aongstr., Oefv. K. Svensk. Vet. Ak. 

Foerh. 30(5):146 (1873). 

T: St. Helena, Andersson s.n. (B, G). 


Bryoph. Bibl. 31:97 (1985). 

oedicaulis C. Müll. in Par., Ind. Bryol. Suppl. 95 

(1900) nom. nud. 

M: N.W. Himalaya, Dehra-Dun, Gollan 1760 

(PC). 



oerstedianus (C. Müll.) Mitt., J. Linn. Soc. Bot. 

12:81 (1869). 

B: Dicranum oerstedianum C. Müll., Bot. 

Zeit. 9(14): 259 (1851). 

T: Costa Rica, „in regione alpina de Irasu 

11000'“, Oersted s. n. (holotype destroyed at 

B; lectotype NY; isolectotypes FH, H-BR). 

ohingaitii R. Brown ter., Trans. New Zealand 

Inst. 29:470 (1897). 

OU: C. pallidus Hook.f. & Wils. cf. Dixon, 

New Zealand Inst. Bull. 3:86 (1923). 


olivaceo-nigricans (C. Müll.) Par., Ind. Bryol. 

Suppl. 95 (1900). 

B: Dicranum olivaceo-nigricans C. Müll., 

Hedwigia 38: 81 (1899). 

T: Transvaal, Wilms s.n. (PC, G). 

OU: C. lepidophyllus (C. Müll.) Jaeg. cf. Sim, 



Bryoph. Bibl. 31:71 (1985). 

orizabae C. Müll., Gen. Musc. Fr. 266 (1900) 

nom. nud. 

orthocomus (C. Müll.) Besch., Bull. Soc. Bot. 

France 32:LVII (1885). 

CU: Chorisodontium aciphyllum (Hook.f. & 

Wils.) Broth. cf. Broth, Nat. Pfl. 1(3):1183 

(1909).

62 

orthopelma (C. Müll.) Par., Ind. Bryol. ed.2,1:320 

(1904). 

B: Dicranum orthopelma C. Müll., Hedwigia 

39: 257 (1900). 

T: Brazil, S. Catarina, Serra Geral, Ule 1128 

(holotype destroyed at B; lectotypus H-BR). 


Lich. 45:141 (1979). 


orthopodius (C. Müll.) Broth., Nat. Pfl. 1(3): 334 

(1901). (*) 

B: Dicranum orthopodium C. Müll., 

Hedwigia 39: 249 (1900). 

orzeszkoanus Ren. & Card., Bull. Soc. R. Bot. 

Belg. 41:20 (1905). 

T: Gr. Comore, Kortala 2000 m, Mace s.n. 

(holotype PC, isotype NY). 

CU: C. crateris cf. Frahm, Bryoph. Bibl. 

31:60 (1985). 

otaramaii R. Brown ter., Trans. New Zealand 

Inst. 29:474 (1897). 

OU: C. appressifolius Mitt. cf. Dixon, New 

Zealand Bull. 3:89 (1923). 


Trans. New Zealand Inst. 59:506 (1928). 

ouro-pretensis Par. in Broth., Nat. Pfl. 1(3):334 

(1901) nom. nud. 


Frahm, Fl. Neotr. 54: 75 (1991). 


ovalis (Hedw.) Wahlenb., Fl. Suec. 2:748 (1926). 

CU: Grimmia ovalis (Hedw.) Lindb. 

ovatus Brid., Mant. Musc. 76 (1819) nom. illeg. 

CU: Grimmia ovalis (Hedw.) Lindb. 

P 

pachycomus Besch. in Par., Ind. Bryol. 256 

(1894) nom. nud.. 


N. Am. Fl. 15:144 (1913). 


pachyneuros (Mol.) Walth. & Mol., Laubm. 

Oberfrank. 106 (1868). 

B: Campylopus flexuosus var. pachyneuros 

Mol. 

OU: Dicranum pachyneuron (;ol.) Kindb., 

Eur. N. Am. Bryin. 190 (1897). 


Britt. ssp. alpinum (Schimp.) Giac. cf. Milde, 

Bot. Zeit. 28:416 (1870). 

pallescens Besch., Ann. Sc. Nat. Bot. ser. 6,9:320 

(1880). 

T: La Reunion, de l’Isle 200 (isotype G). 

CU: C. hildebrandtii cf. Frahm, Bryoph. Bibl. 

31:91 (1985). 

pallidisetus Thér., Rec. Publ. Soc. Havraise Etud. 

Div. 1925:130 (1926). 

T: Madagaskar, Mt. Tsaratanara 1200 - 2000 

m, Perrier de la Bathie s.n. (PC). 


Bibl. 31:197 (1985). 

pallido-nitens Thér., Rec. Soc. Havraise Etud. 

Div. 1932:134 (1932). 

T: Madagaskar, Sikora s.n. (holotype PC). 


Bibl. 31:164 (1985). 

pallidus Hook.f. & Wils., Fl. Nov. Zel. 2:68 

(1854). 

T: Campbell Island, F.D. Hooker, antarctic 

voyage of Erebus and Teror (holotype BM). 

CU: C. pyriformis (Schultz) Brid. cf. Corley 

& Frahm, J. Bryol. 12:190 (1982). 

paludicola Broth. in Mildbr., Wiss. Erg. Deutsch. 

Zentr. Afr. Exp. 2:140 (1910) 

T: Ruanda, Rugege Wald, Rukarara Moor zw. 

Carex-Bülten 1800 m, Mildbraed 985a 

(holotype B-BR). CU: C. pyriformis 

(Schultz) Brid. cf. Frahm, Nova Hedwigia 59: 

150 (1994). 

paludosus J.-P.Frahm & J.K.Bartlett, Journ. 

Bryol. 12:376 (1983). 

T: New Zealand, South Island, bog near 

Jacksons 42°S, Barthlett 18 (holotype WELL, 

isotypes BM, NICH, NY). 

CU: C. vesticaulis Mitt. cf. Frahm. Bryol. 

Beitr. 7: 57 (1987). 

papuensis Bartr., Brittonia 9:33 (1957). 

T: Papua New Guinea, Milne Bay District, 

N-slopes of Mt. Dayman, Maneau Range, 

Brass 22966 (holotype FH). 

CU: C. exasperatus (Nees & Blum) Brid. var. 

lorentzii (Fleisch.) J.-P.Frahm cf. Frahm et 

al., Acta Bot. Fennica 131:80 (1985). 

paradoxus Wils. in Hardy, Berwickshire Nat. 

Hist. Cl. 1868:48 (1868). 

CU: C. flexuosus (Hedw.) Brid. cf. Hagen, 

Norsk. Vid. Selsk. Skrift. 1914:75 (1915). 

paraguayensis Broth., Nat. Pfl. 1(3):332 (1901) 

nom. nud. 

parallelus Thér. in Broth., Nat. Pfl. ed.2,10:188

(1909). 

T: „Ile Bourbon, hb. Despreaux“ 

CU: C. arcuatus cf. Frahm, Bryoph. Bibl. 

31:20 81985). 

paramoensis F.Bowers, Bryologist 77:152 

(1974) nom. nov. pro C. atratus Bartr. hom. 

illeg. 

CU: C. subcuspidatus (Hampe) Jaeg. f. 

Frahm, Fl. Neotr. 54: 176 (1991). 

parisii Ren., Prodr. Fl. Bryol. Madag. Suppl. 25 

(1909). 

T: Madagaskar prov. d’Ambositra Salvan s.n. 


CU: C. smaragdinus (Brid.) Jaeg. cf. Frahm, 

Bryoph. Bibl. 31:177 (1985). 

parvicaespitosus (C. Müll.) Par., Ind. Bryol. 

ed.2,1:321 (1904). 

B: Dicranum parvicaespitosum C. Müll., 

Hedwigia 39: 259 (1900). 



parvulus Schimp. in Besch., Mem. Sc. Nat. Cherbourg 

16:165 (1872) nom. nud. 

pascuanus J.-P.Frahm, Bryoph. Bibl. 5:91 (1975) 

nom. nov. pro C. saxicola Broth. in Skottsb. 

hom. illeg. 


Hedwigia 34:392 (1981). 


Beitr. 7: 58 (1987). 

patagonicus Broth. ex Dus., Ark. Bot. 4(13):14 

(1905). 

CU: C. chilensis cf. Frahm, Nova Hedwigia 

29:244 (1978). 

patens (Hedw.) Brid., Mant. Musc. 73 (1819). 

CU: Racomitrium patens (Hedw.) Hüb. 

pauper (Hampe) Mitt., J. Linn. Soc. Bot. 12: 74 

(1869). 

B: Dicranum pauper Hampe, Linnaea 32: 137 

(1863). 

T: Colombia. Manzanos, Lindig 2013 

(holotype not seen; isotypes H-BR, H-SOL, 

GOET, JE, NY). 

pauper (Hampe) J.-P. Frahm var. lamprodictyon 

(Hampe) Frahm, Bryol. Beitr. 7: 58 (1987). 

B: Dicranum lamprodictyon Hampe, Prodr. 

Fl. Nov. Granat. 339 (1866). 

OU: Campylopus lamprodictyon (Hampe) 


pelichucensis Williams, Bull. New York Bot. 

63 

Gard. 3:110 (1903). 

T: Bolivia, Pelichuco River, Williams 2835 




pelidnus (Stirt.) Stirt., Ann. Scott. Nat. Hist. 

9(38):113 (18901). 

CU: C. brevipilus B.S.G. cf. Dixon, J. Bot. 

61:16 (1923). 

penicillatus (Hornsch.) Jaeg., Ber. S. Gall. 

Naturw. Ges. 1870-71:442 (1872) hom. illeg.. 

B: Dicranum penicillatum Hornsch., Fl. Bras. 

1(2): 13 (1840). 

T: Brazil, Serra dos Orgaos, Gardner 25 

(holotype destroyed at B; lectotype B; 

isolectotypes GOET, JE). 

CU: C. lamellinervis (C. Müll.) Mitt. cf. 

Hampe, Vid. Medd. Naturh. For. Kjoebenh. 

ser.3,2:272 (1870). 

penicillatus var. exaltatus (C. Müll.) Thér., Mem. 

Soc. Cubana Hist. Nat. 13:217 (1939). 

CU: C. lamellinervis (C. Müll.) Mitt. var. 

exaltatus (C. Müll.) J.-P. Frahm cf. Frahm, 

Fl. Neotr. 54: 129 (1991). 

OU: C. cubensis Sull. var. exaltatus (C. 

Müll.)J.-P.Frahm, Nova Hedwigia 41:275 

(1985). 

penicillatus Brid., Mant. Musc. 73 (1819). 

CU: C. fragilis (Brid.) B.S.G. cf. C.Mueller, 

Syn. Musc. 1:393 (1848). 

perangustifolius C. Müll. Flora 82:479 (1869). 

T: „Sandwich, am Mastigobryo Remyano“ 

Remy 712 (BM, FH). 

OU: C. hawaiico-flexuosus (C. Müll.) Par. cf. 

Bartram, Bishop Mus. Bull. 101:40 (1933); 

C. purpureo-flavescens (C. Müll.) Par. 

CU: hawaiicus (C. Müll.) Jaeg. var. hawaiicoflexuosus 

(C. Müll.) J.-P. Frahm cf. Frahm, 


perauriculatus Broth., Oefv. Finsk. Vet. Ak. 

Foerh. 42:93 (1896). 

T: Australia, N.S.W., Richmond R., Watts 


CU: C. robillardei var. perauriculatus 

(Broth.) J.-P. Frahm, Bryol. Beitr. 7: 89 

(1987). 

percristatus Thér., Bull. Soc. Bot. Geneve 26:80 

(1936). 

T: Iles Philippines, Cumingham 2217 

(holotype PC).

64 



percurvatus C. Müll. Flora 83:331 (1897). (*) 

perexilis (C. Müll.) Par., Ind. Bryol. Suppl. 95 

(1900). 

B: Dicranum perexile C. Müll., Nuov. Giorn. 

Bot. Ital. n. ser. 4: 34 (1897). 

T: Bolivia, Cochabamba, Germain 1006 

(holotype destroyed at B; lectotype H-BR; 

isotypes JE, NY). 

perfalcatus (C. Müll.) Broth., Nat. Pfl. 1(3): 334 

(1901). 

B: Dicranum perfalcatum C. Müll., Hedwigia 

39: 250 (1900). 

T: Brazil, Sta. Catarina, Ule 1124 (holotype 

destroyed at B; lectotype, H-BR). 

OU: C. crispatus Broth. cf. Frahm, Bryoph. 

Bibl. 5:93 (1973). 


caldensis (Aongstr.) J.-P.Frahm. 

pergracilis Stirt., Ann. Scott. Nat. Hist. 14:105 

(1905). 

CU: C. pyriformis cf. Dixon, J. Bot. 61:16 

(1923). 

perhorridus Dus., Ark. Bot. 4(13):16 (1905). 

T: Patagonia occ., Rio Aysen, Dusén s.n. (B): 

CU: Chorisodontium cf. fulvastrum (Besch.) 

Broth.. 

perichaetialis P. Varde & Thér., Bull. Soc. Bot. 

France 87: 352 (1940). 

T: Zaire, massif du Kahuzi 3000 m, Humbert 


perincanus (C. Müll.) Kindb., Enum. Bryin. Exot. 

51 (1889). 

B: Dicranum perincanum C. Müll., Flora 68: 

410 (1885). 

OU: C. canescens fo. perincanus (C. Müll.) 

J.-P.Frahm, Revue Bryol. Lich. 42:903 

(1976). 



perinvolutus Thér. in Froehl., Arch Hydrobiol. 

Suppl. 21:307 (1955). 

T: Sumatra, Fort de Koek supra Padäng, 

Ruttner (1929) s.n. (holotype PC). 

CU: C. macgregorii cf. Frahm, Nova 

hedwigia 39: 595 (1984). 

permollis Thér., Rev. Chil. Hist. Nat. 30: 341 

(1926). 

T: Chile, Santiago, Jaffuel s.n. (FH, W). 


Beitr. 7: 59 (1987). 

perplexans Stirt., Trans. Bot. Soc. Edinburgh 26: 

247 (1914). 

CU: C. atrovirens De Not. cf. J. Bot. 61: 16 

(1923). 

perporosus Broth., Denkschr. Ak. Wiss. Wien 

Math. Nat. Kl. 83: 263 (1926). 

CU: C. arctocarpus ssp. caldensis cf. Frahm, 

Revue Bryol. Lichénol. 45: 131 (1979). 

perpusillus Mitt., J. Linn. Soc. Bot. 22: 301 

(1886). 

T: Zentr.Afr., Ugogo, Bishop Hannington s.n. 


perreduncus (C. Müll.) Par., Ind. Bryol. Suppl. 

95 (1900). 

CU: Chorisodontium speciosum cf. Frahm. 

J. Hattori Bot. Lab. 44:484 (1978). 

perrieri Thér., Rec. Publ. Soc. Havraise 

Etud. Div. 89:117 (1922). 

T: Madagascar, Perrier de la Bathie 


CU: C. flaccidus Ren. & Card. cf. 

Frahm, Nova Hedwigia 39:592 

(1984). 

perrottetii (Mont.) Fleisch., Musci Fl. 


CU: Dicranella perrottetii (Mont.) 

Mitt. 

persimplex C. Müll., Abh. Naturw. Ver. 

Bremen 16(3):496 (1900). 

OU: C. insititius Hook.f. & Wils. cf. 

Dixon, New Zealand Inst. Bull. 

3(3):88 (1923). 

CU: C. clavatus (R.Brown) Wils. cf. 

Sainsb., Trans. New Zealand Inst. 

59:507 (1928). 

pertristis (C. Müll.) Par., Ind. Bryol. 

Suppl. 95 (1900). 

B: Dicranum pertriste C. Müll., Flora 

82: 444 (1896). 

T: Hawaii, Oahu, ad rupes, Clakre s.n. 

(could not be located). 

CU: C. umbellatus (Arn.) Par. cf. Bartram, 

Bishop Mus. Bull. 101:44 

(1933). 

OU: C. richardii Brid. cf. Dixon, J. 

Bot. 60:287 (1922). 

peruvianus Williams, Bull. Torr. Bot. Cl. 

42:393 (1915).

T: Peru, near Molendo, Rose & Rose 

18997 (holotype NY). 

CU: Pilopogon peruvianus (Williams) 

J.-P.Frahm, Lindbergia 9:104 (1983). 

OU: Pilopogon laevis (Tayl.) Thér. cf. 

Hegewald & Hegewald, Journ. Hattori 

Bot. Lab. 43:73-80 (1977). 

pes funariae (C. Müll.) Par., Ind. Bryol. 

257 (1894). 

T: Philippines, without locality, Wallis 

s.n. (BM, H-BR). 

CU: C. laxitextus Lac. cf. Frahm, 

Bryol. Beitr. 7: 60 (1987). 

pilifer Brid. Mant. Musc. 72 (1819). 

T: Italien, „in insula Ischia ad rupes, Auguste 

1806“ (holotype B). 

pilifer ssp. galapagensis (J.P. Frahm & Sipman) 

Bryol. Beitr. 7: 60 (1987). 

B: galapagensis J.-P.Frahm & Sipman, J. 

Bryol. 10:61 (1978). 

T: Ecuador, Galapagos Islands: Santa Cruz, 

base of Mt. Crocker, Gradstein & Weber 17 

(U CDRS, COLO, QCA). 

pilifer var. lamellatus (Montagne) Grad.& Sip., 

Bryologist 81: 119 (1978). 

B: Campylopus lamellatus Mont., Ann. Sc. 

Nat. Bot. ser. 2,9: 52 (1838). 

OU: Dicranum lamellatum (Mont.) C. Müll., 

Syn. Musc. 1: 411 (1848). 

T: Bolivia (holotype PC; isotype BM). 

pilopogon C. Müll., Gen. Musc. Fr. 270 

(1900) nom. nud. 

pilosissimus Schimp. ex Besch., Mem. 

Soc. Sc. Nat. Cherbourg 16:168 

(1872). 

T: Mexico, Mirador, Sartorius s. n. 

(not seen). 

OU: C. introflexus (Hedw.) Brid. cf. 

Williams, N. Am. Fl. 15(2):144 

(1913). 


pinangensis Thér. in Henry, Rev. Bryol. 

n.ser. 1:42 (1928). 

T: Pinang. Montagne, Deschamp s.n. 


CU: C. serratus Lac. cf. Frahm, Bryol. 

Beitr. 7: 60 (1987). 

pinfaensis Thér., Bull. Ac. Int. Geogr. 

Bot. 19:18 (1909). 

T: China „Prov. Kouy, Tcheou, Pin- 

65 

fa, leg. Cavalerie no. 1941“ (Isotype 

S). 

CU: C. fragilis ssp. fragilis cf. Frahm, 

Trop. Bryol. 15 (in press). 

pintasi Broth. in C. Müll., Gen. Musc. Fr. 

275 (1900) nom. nud. 

pittieri Williams, Bull. Torrey Bot. Club 34: 56 

(1908). 

T: Colombia, Rio Lopez, Pittier 1088 


pittieri var. congestus (Thér.) Florsch.-de Waard 

& Florsch., Bryologist 82:223 (1979). 

pittieri var. latilimbatus (Thér.) Florsch.-de 

Waard & Florsch., Bryologist 82:223 

(1979). 

platyneuron (Hampe) Jaeg., Ber. S. Gall. Naturw. 

Ges. 1877-78: 382 (1880). 

B: Dicranum platyneuron Hampe, Vid. 

Medd. Naturh. Foer. Kjoebenh. ser. 3,6: 139 

(1875). 

OU: Thysanomitrion platyneuron (Hampe) C. 

Müll., Gen. Musc. Fr. 260 (1900). 


(isotype, NY). 


Lich. 45:156 (1979). 

pleurocarpus (C. Müll.) Par., Ind. Bryol. Suppl. 

95 (1900). 

B: Dicranum pleurocarpum C. Müll., Bull. 

Herb. Boiss. VI: 35 (1898). 




Frahm, Fl. Neotr. 54: 75 (1991). 


plumosus Sull. ex Par., Ind. Bryol. ed.2,1:305 

(1904) nom. nud. 

OU: C. densifolius Aongstr. cf. Paris l.c. 




poasensis Ren. & Card., Bull. Soc. R. Bot. Belg. 

31(1): 148 (1893). 

T: Costa Rica, Volcán de Poás, Pittier s. n. 

(not seen). 

OU: C. chrismarii (C. Müll.) Mitt. cf. 

Williams, N. Am. Fl. 15:136 (1913). 


pobeguinii Par. & Broth. Rev. Bryol. 31:84 

(1904).

66 

T: „Guinea gall., Pobequin“ s.n. (BM,NY,PC, 

L, B) . 



poicilophyllus Nav. Bull. Soc. R. Bot. Belg. 60; 

19 (1928). 

T: Zaire, entre Angia et Tango, Bequaert 6194 

(isotype L). 

OU: C. introflexus (Hedw.) Brid. cf. Naveau 

l.c. 


polyanthos Besch. Ann. Sci. Nat. Bot. ser. 5, 18: 

199 (1873). 

T: „in sylvis, prope sinum dictum du Pony, 

ad terram humidam“ Balansa 748 (lectotype 

PC). 


polytrichoides De Not., Syll. Musc. 222 (1838). 

CU: C. pilifer Brid. cf. Gradstein & Sipman, 


porophorus Stirt., Ann. Scott. Nat. Hist. 10:113 

81901). 

CU: C. setifolius Wils. cf. Dixon, J. Bot. 61:16 

(1923). 

porphyreocaulis (C. Müll.) Kindb., Enum. Bryin. 

Exot. 89 (1889). 

B: Dicranum porphyreocaule C. Müll., 

Linnaea 42: 472 (1879). 

T: Venezuela, Fendler 34 (holotype destroyed 

at B; lectotype BM; isolectotypes MICH, NY, 

PC, S). 

CU: C. heterostachys cf. Frahm, J. Hattori 

Bot. Lab. 44:485 (1978). 

porphyreodictyon (C. Müll.) Mitt., J. Linn. Soc. 

Bot. 12:75 (1869) 

T: Brasilia, Caldas, Widgren (GOET). 

B: Dicranum porphyreodictyon C. Müll., Syn. 

Musc. I: 395 (1848). 

OU: Campylopus filifolius (Hornsch.) Mitt. 



P.Frahm. 

powellii (C. Müll.) Par., Ind. Bryol. Suppl. 95 

(1900). 

B: Thysanomitrion powellii C. Müll., Bot. 

Jahrb. 23: 230 (1836). 

T: Samoa, Powell (H-BR). 



OU: C. richardii cf. Dixon, J. Bot. 60:290 

(1922). 

praealtus (C. Müll.) Par., Ind. Bryol. Suppl. 96 

(1900). 

B: Dicranum praealtum C. Müll., Hedwigia 

37: 227 (1898). 

T: Puerto Rico, Sierra de Luquillo, Sintensis 

s. n. (holotype destroyed at B; lectotype NY; 

isolectotypes H-BR; MO). 

CU: C. subcuspidatus (Hampe) Jaeg. cf. 


(1981). 

praealtus var. damazii (Broth.) J.-P.Frahm, 

Bryol. Beitr. 7: 90, 1987. 

B: C. damazii Broth. 

praemorsus (C. Müll.) Jaeg., Ber. S. Gall. 

Naturw. Ges. 1870-71:437 (1872). 

B: Dicranum praemorsum C. Müll. Bot. Zeit. 

20: 337 (1862). 

OU: Thysanomitrion praemorsum (C. Müll.) 

Broth., Bishop Mus. Bull. 40: 7 (1927). 

T: Ins. Sandvicensis (Hawaii), Ditrichsen in 

exped. transatl. danica (C). 

praetermissus J.-P. Frahm, Bryoph. Bibl. 31: 152 

(1985). 

T: Gabun, Mocambo, Le Testu 4753 


prasino-rufus Stirt., Ann. Scott. Nat. Hist. 15:108 

(1906). 

CU: C. atrovirens De Not. var. muticus Milde 

cf. Dixon, J. Bot. 61:16 (1923). 

preussii Broth. in Par., Ind. Bryol. Suppl. 96 

(1900) nom. nud. 

M: Kamerun, Buea, Preuss s.n. (isotype B). 

CU: C. johannis-meyeri (C. Müll.) Par. cf. 


procerus (C. Müll.) Par., Ind. Bryol. 258 (1894). 

B: Dicranum procerum C. Müll., Flora 73: 

472 (1890). 

CU: C. jamesonii cf. Frahm, J. Bryol. 11:217 

(1980). 

procerus Broth. in Par., Ind. Bryol. Suppl. 89 

(1900). 

T: Brazil, Rio de Janeiro, Pico de Papageio, 

Ule Bryothec. bras. 110 (holotype, H-BR) 

HS: C. procerus Broth., Hedwigia 38 (Beibl.): 

58 (1899) nom. nud. 

OU: C. praealtus (C. Müll.) Par. cf. Broth., 

Denkschr. Ak. Wiss. Wien, Math. Nat. Kl. 

83:260 (1926). 

CU: C. subcuspidatus (C. Müll.) Par.

proliferus (C. Müll.) Mitt., J. Linn. Soc. Bot. 

12:85 (1869). 

B: Dicranum proliferum C. Müll., Syn. Musc. 

2: 602 (1851). 

T: Venezuela, Tovar, Wagner s. n. (not seen). 


N. Am. Fl. 15:144 (1913). 

OU: C. pilifer Brid. 

propinquus (Hampe) Mitt., J. Linn. Soc. Bot. 

12:75 (1869). 

B: Dicranum propinquum Hampe, Ann. Sc. 

Nat. Bot. ser. 5,3: 367 (1865). 

T: Colombia. Manzanos 2700 m, Lindig 2119 

(holotype not seen; isotypes, H-BR, H-SOL, 

GOET, JE, NY) 

CU: C. pauper cf. Frahm. Bryol. Beitr. 7: 61 

(1987). 

pseudobicolor C. Müll. ex Ren. & Card., Bull. 

Soc. R. Bot. Belg. 34: 59 (1896). 

T: Madagaskar, Boswell s.n. (holotype PC). 

OU: C. bicolor (C. Müll.) Wils. cf. Dixon in 

Sim, Trans. R. Soc. S. Afr. 15:180 (1926). 

pseudobrachymitrius (C. Müll.) Par., Ind. Bryol. 

ed.2,1:324 (1904). 

B: Dicranum pseudobrachymitrium C. Müll., 

Hedwigia 39: 249 (1900). 

T: Brazil, Minas Gerais, Serra do Itatiaia, Ule 

1803 (lectotype H-BR). 


Lich. 45:156 (1979). 

pseudoclavatus Par., Ind. Bryol. 243 (1894) nom. 

nud. 

CU: C. introflexus (Hedw.) Brid. cf. Sim, 


pseudodichrostis Broth., Denkschr. Ak. Wiss. 

Wien, Math. Nat. Kl. 83:263 (1926). 

T: Brazil, Sao Paulo, Schiffner 402 (lectotype, 

H-BR). 

CU: C. viridatus cf. Frahm, Revue Bryol. 

Lich. 45:166 (1979). 

pseudodicranum Herz., Beih. Bot. Centralbl. 

26(2):53 (1910). 

T: Bolivia, Cerro Amboro, Herzog s. n. 

(holotype, JE; isotypes B, H-BR). 

CU: C. lamellinervis cf. Frahm, Bryoph. Bibl. 

5:100 (1975). 

pseudofilifolius (C. Müll.) Par., Ind. Bryol. 258 

(1894). 

B: Dicranum pseudofilifolium C. Müll., 

Linnaea 42: 471 (1879). 

67 

T: Venezuela, Fendler 40a (holotype 

destroyed at B; lectotype H-BR; isolectotype 

JE). 

CU: C. filifolius (Hornsch.) Mitt. cf. Frahm, 

Bryoph. Bibl. 5:100 (1975). 

pseudogracilis Card. & Dix., Rec. Bot. Surv. 

India 6:76 (1914). 

T: Ceylon, Unasgiriya (H-BR). 

CU: C. goughii cf. Frahm, J. Bryol. 13:172 

(1984); C. fragilis (Brid.) B.S.G. ssp. goughii 


pseudogriseus C. Müll., Gen. Musc. Fr. 265 

(1900) nom. nud. 

pseudojulaceus Sim, Trans. R. Soc. S. Afr. 

15:169 (1926) hom. illeg. 

OU: C. simii Schelpe in Magill & Schelpe, 

Mem. Bot. Surv. S. Afr. 43:5 (1979). 

CU: C. julaceus Jaeg. ssp. arbogastii (Ren. 

& Card.) J.-P.Frahm, Bryoph. Bibl. 31:109 

(1985). 

pseudojulaceus C. Müll., Bull. Herb. Boiss. 6:38 

(1898). 

T: Orange Free State, Kadziberg, Rehmann 

58 (holotype B, isotypes PRE, L). 



pseudolongipilus C. Müll., Gen. Musc. Fr. 266 

(1900) nom. nud. 

pseudomuelleri Card., Bull. Herb. Boiss. 

ser.2,7:715 (1907) 

T: Japon, Futatabisan, à terre 500m, n. 2247 


CU: C. japonicus Broth. cf. Takaki, J. Hatt. 

Bot. Lab. 30:240 (1967). 

pseudo-nanus (C. Müll.) Jaeg., Ber. S. Gall. 

Naturw. Ges. 1870-71:416 (1872). 

B: Dicranum pseudo-nanum C. Müll., Bot. 

Zeit. 17: 190 (1859). 

T: Java, Blume s.n. 


Müll.) Besch. cf. Fleischer, Musci Fl. 



& J.-P.Frahm cf. Giese & Frahm, Lindbergia 

11: 126 (1985). 

pseudonigrescens Thér., Rec. Publ. Soc. Havraise 

Etud. Div. 1922:119 (1922). 

T: Madagascar, Massif d’ Andringika, 2500 

m, Perrier de la Bathie s.n. (isotype H-BR). 

CU: C. flaccidus Ren. & Card. cf. Frahm,

68 


pseudovirescens Ren. & Card., Bull. Soc. R. Bot. 

Belg. 41:19 (1905). 

T: Madagascar, Maroantsetra, sur un arbre 

pourré, Mathieu s.n. (holotype PC). 

CU:? C. arcuatus (Brid.) Jaeg. cf. Frahm. 

Bryol. Beitr. 7: 63 (1987). 

pterotoneuron (C. Müll.) Jaeg., Ber. S. Gall. 

Naturw, Ges. 1870-71:417 (1872). 

B: Dicranum pterotoneuron C. Müll., 

Linnaea 36: 34 (1869). 

T: Insula Ceylon prope Punduloya, Nietner 

s.n. (H-BR). 

OU: Thysanomitrion pterotoneuron (C. Müll.) 

C. Müll., Gen. Musc. Fr. 260 (1900). 

CU: C. umbellatus (Arn.) Par. var. 

pterotoneuron (C. Müll.) J.-P.Frahm, Nova 

Hedwigia 39:602 (1994). 

ptychotheca Herz., Beih. Bot. Centralbl. 26:49 

(1910). 

T: Bolivia, Cochabamba, Incacorral, Herzog 

284 (holotype JE; isotype B). 

CU: C. reflexisetus (C. Müll.) Broth. cf. 


pudicus (C. Müll.) Jaeg., Ber. S. Gall. Naturw. 

Ges. 1870-71:435 (1872). 

B: Dicranum pudicum Hornsch. ex C. Müll., 

Syn. Musc. 1: 407 (1848). 

T: Nova Hollandia, Sieber 24 (JE). 

CU: C. introflexus (Hedw.) Brid. cf. Sainsb., 

Pap. Proc. R. Soc. Tasmania 89:70 (1955). 

OU: Campylopus introflexus (C. Müll.) Brid. 

ssp. pudicus (C. Müll.) Dix. 

puiggarii (Geh. & Hampe) Par., Ind. Bryol. 258 

(1894). 

B: Thysanomitrion puiggarii Geheeb & 

Hampe, Flora 64: 346 (1881). 

T: Brazil, Sao Paulo, Puiggari s. n. (isotype 

JE). 

CU: C. richardii Brid. cf. Frahm, Revue 

Bryol. Lich. 41:323 (1975). 

pulvinaris Broth., Nat. Pfl. 1(3):334 (1901) nom. 

nud. 

pulvinatus (C. Müll.) Rehm. in Par., Ind. Bryol. 

96 (1900) hom. illeg. 

B: Dicranum pulvinatum Rehm.ex Müll., 

Hedwigia 38: 80, (1899). 

T: South Africa, Stinkwater, Rehmann 62 

(L,NY). 

OU: C. pallidus Hook.f. & Wils. cf. Magill, 

Fl. S. Afr. Bryoph. 1(1):147 (1981). 

CU: C. catarractilis (C. Müll.) Par. cf. Frahm, 

Bryoph. Bibl. 31:51 (1985). 

pulvinatus (Hedw.) Brid., Mant. Musc. 75 (1819). 

CU: Grimmia pulvinata (Hedw.) Sm. 

purpurascens Lor. Moosstud. 159 (1864). 

T: Cape, Table Mountain, Ecklon s.n. (isotype 

NY). 


31:143 (1985). 

purpurascens Stirt., Ann. Scott. Nat. Hist. 10:109 


CU: C. schwarzii Schimp. cf. Dixon, J. Bot. 

61:17 (1923). 

purpureo-alaris Broth., Bull. Soc. Bot. Ital. 

1904:15 (1904) nom. nud. 

CU: C. wheeleri (Hampe ex C. Müll.) Par. cf. 

Frahm, The Bryologist 94: 66, 1991. 

purpureo-aureus (C. Müll.) Par., Ind. Bryol. 

Suppl. 96 (1900) 

B: Dicranum purpureo-aureum C. Müll., 

Hedwigia 38: 82 (1899). 

T: Transvaal, Duivels Krackler bei 

Lydenburg, Wilms (1884) s.n. (holotype B, 

isotype G). 

CU: C. aureo-nitens cf. Frahm, Bryoph. Bibl. 

31:24 (1985). 


Soc. S. Afr. 15:176 (1926), C. pilifer Brid. 

cf. Magill, Fl. S. Afr. Bryoph. 1(1):152 

(1981). 

purpureoflavescens (C. Müll.) Par., Ind. Bryol. 

Suppl. 96 (1900). 

B: Dicranum purpureo-flavescens C. Müll., 

Flora 82: 442 (1896). 

T: Hawaii, without locality, Wheeler s.n. 

(lectotype B). 

OU: C. densifolius Aongstr. var. purpureoflavescens 

(Hampe ex C. Müller) H. Miller, 

J. Hattori Bot. Lab. 30: 271 (1967). 

CU: C. hawaiicus (C. Müll.) Jaeg cf. Frahm, 

Revue Bryol. Lichénol. 44: 50, 1978. 

purpureocaulis Dus., Ark. Bot. 4(13): 11 (1905). 

T: Chile aus. in litore lacus Llanquihue, Dusen 

157 (H-BR). 

purpureocaulis Dus. ssp. aberrans (Broth.) J.- 

P.Frahm. Bryol. Beitr. 7: 63 (1987) comb. 

inval. 

B: Campylopus aberrans Broth. 

pusillus Schimp. ex C. Müll., Hedwigia 38:78


OU: Microcampylopus nanus (C. Müll.) Par. 

cf. Dixon, Trans. R. Soc. S. Afr. 8:187 (1920). 

CU: Dicranella subsubulata cf. Magill, F. S. 

Afr. Bryoph. 1(1):121 (1981). 

pusillus Schimp. in Besch., Mem. Sc. Nat. Cherbourg 

16:165 (1872). 


Giese & J.-P.Frahm cf. Giese & Frahm, 

Lindbergia..... 


Williams. 

pycnangius C. Müll. in Kindb., Enum. Bryin. 

Exot. 106 (1891) nom. nud. 

CU: C. clavatus (R. Brown) Wils. in Hook. 

cf. Frahm, Bryol. Beitr. 7: 87 (1987). 

pygmaeo-filifolius C. Müll., Gen. Musc. Fr. 269 

(1900) nom. nud. 

M: Madagaskar, Andrangokaha, Sikora 188 

(B). 

CU: Bryohumbertia cf. flavicoma cf. Frahm, 

Bryoph. Bibl. 31:198 (1985). 

pyriformis (Schultz) Brid., Bryol. Univ. 1: 469 

(1826). 

B: Dicranum pyriforme Schultz, Prodr. Fl. 

Starg. Suppl. 73 (1819). 

T: „ad pagum Ballin in dem Holz hinter dem 

Hofgarten“ Schultz s.n. (B). 

OU: C. fragilis (Brid.) B.S.G. var. pyriformis 

(Schultz) Agst. cf. Index Muscorum. 

pyriformis var. azoricus (Mitt.) Corley, Journ. 

Bryol. 9:201 (1976). 

B: Campylopus azoricus Mitt. 

pyriformis var. fallaciosus (Thér.) Corley, Journ. 

Bryol. 9:201 (1976). 

pyriformis var. fragilis Corb., Mem. Soc. Nat. 

Cherbourg 26:223 (1889). 

CU: C. pyriformis var. fragilis Husn. cf. 


pyriformis var. fragilis Husn., Muscol. Gall. 43 

(1882). 

CU: C. pyriformis mod. fragilis cf. Frahm, 


pyriformis var. hamatus J.-P.Frahm & 

J.K.Bartlett, Journ. Bryol. 12:378 (1983). 

T: New Zealand, South Island, Denniston 

Plateau near Westfort, in bog, Bartlett 26 

(holotype WELL, isotype bb. Frahm). 

pyriformis var. muelleri (Jur.) Braithw., J. Bot. 

8:33 (1870) = C. pyriformis stat. muelleri 

69 

cf. Frahm, Nova Hedwigia 34:405 (1981). 

Q 

queszelii Desplanques & Hebrard, Riviera 

scientif. 8:61 (1972). 

OU: Ditrichaceae cf. Frahm, Bryoph. Bibl. 

31:197 (1985). 

CU: Verrucidens spec. vel Dicranoweisia 

spec. cf. Vitt (in litt.) 

quintasii Broth., Bol. Soc. Broter. 8:175 (1890). 

T: „Afr. occ. trop., Ins. S. Thome, leg. Fr. 

Quintas“ (isotypes PC, S). 



R 

rabenii Lor., Moosstud. 158 (1864). 

T: Brasilia, com. Raben (B). 

CU: C. filifolius var. humilis cf. Frahm, 

Cryptogamie Bryol. Lich. 2:444 (1981) = 


var. humilis (Mont.) J.-P.Frahm. 

rallieri Card., Bull. Mus. Hist. Nat. Paris 22:337 

(1916). 

T: Kerguelen Expedition, Rallier de Baty 

(1913-14) 30,31 (syntypes PC). 

CU: C. eximius Reichdt. cf. Frahm, Bryoph. 

Bibl. 31:67 (1985). 

ramuliger Broth., Denkschr. Ak. Wiss. Wien 

Math. Nat. Kl. 83:262 (1926). 

T: Type. Brazil, Sao Paulo, Apiahy, Schiffner 

110 (holotype not present in H-BR; isotypes 

BM, PC). 

CU: C. cryptopodioides Broth. cf. Frahm, Fl. 

Neotr. 54: 77 (1991). 

rarus R.Brown ter., Trans. New Zealand Inst. 

29:470 (1897). 


Zealand Inst. Bull. 3(3):89 (1923). 


Trans. New Zealand Inst. 59(3):506 (1928). 

raspannae C. Müll. in Bruehl., Rec. Bot. Surv. 

India 13(1):123 (1931) nom. nud. 

rauei (Aust.) Par., Ind. Bryol. 259 (1894). 

CU: Dicranum muehlenbeckii B.S.G. cf. Par. 

l.c. 

reconditus (Thwait. & Mitt.) Jaeg., Ber. S. Gall.

70 

Naturw. Ges. 1877-78:384 (1880). 

B: Sphaerothecium reconditum Thwait. & 


T: Ceylon, Central Province, Thwaites 11 

(FH). 

CU: Sphaerothecium reconditum Thwait. & 

Mitt. 

rectipes C. Müll., Gen. Musc. Fr. 270 (1900) nom. 

nud. 

OU: C. pleurocarpus cf. Frahm, Revue Bryol. 

Lich. 42:609 (1976) = C. controversus 

(Hampe) Jaeg. 


rectipilis C. Müll. in Kindb., Enum. Bryin. Exot. 

106 (1891) nom. nud. 

OU: C. xanthophyllus Mont. cf. Theriot, Rev. 

Chil. Hist. Nat. 22:82 (1918). 


J. Linn. Soc. Bot. 12:84 (1869). 

rectisetus (Hampe) Jaeg., Ber. S. Gall. Naturw. 

Ges. 1877-78:385 (1880). 

B: Dicranum rectisetum Hampe, Vidd. Med. 

Naturh. Foer. Kjoebenh. ser. 3, 6: 137 (1875). 





(1981). 

recurvifolius Dus., Ark. Bot. 4:9 (1905). 

CU: C. aureo-nitens ssp. recurvifolius (Dus.) 

J.-P.Frahm cf. Frahm, Bryol. Beitr. 7: 95 

(1987).. 

recurvipilus (C. Müll.) Par., Ind. Bryol. 

ed.2,1:325 (1904). 

B: Dicranum recurvipilum C. Müll., 

Hedwigia 39: 259 (1900). 

T: Brazil, S. Catarina, Novo Friburgo, Peters 

1871 (not seen). 

CU: C. griseus cf. Frahm, Revue Bryol. Lich. 

42:606 (1976). 

recurvus (Mitt.) Jaeg., Ber. S. Gall. Naturw. Ges. 

1870-71:418 (1872). 

B: Dicranum recurvum Mitt., J. Linn. Soc. 

Bot. Suppl. 1: 20 (1859). 

T: India, Nilghiri Mtns., Gardner 16 


recurvus ssp. cataractarum Fleisch., Musci Fl. 


OU: C. cataractarum (Fleisch.) Fleisch., C. 

hemitrichius (C. Müll.) Jaeg. cf. Eddy, 

Malesian Mosses vol. 1 (1988). 

CU: C. recurvus (Mitt.) Jaeg. cf. Frahm, 

Bryol. Beitr. 7: 22 (1987), 

reduncus (Reinw. & Hornsch.) Bosch & Lac., 

Bryol. Jav. 1:75 (1858) . 

B: Dicranum reduncum Reinw. & Hornsch., 

Nov. Act. Leop. Car. 14: 703 (1829). 

T: Java, Gedeh, Junghuhn (H-BR). 

CU: C. comosus cf. Frahm, J. Bryol. 13:166 

(1894). 

reflexisetus (C. Müll.) Broth., Nat. Pfl. 1(3): 332 

(1901). 

B: Dicranum reflexisetum C. Müll., Linnaea 

38: 586 (1874). 

T: Colombia, Antioquia, Páramo de Sonsón, 

Wallis s. n. (holotype destroyed at B; 

lectotype H-BR) 

reflexus Broth. in Herz., Biblioth. Bot. 87:23 

(1916). 

T: Bolivia, Incacorral, Herzog 4988 (holotype 

JE, isotypes B, FH, H-BR). 

CU: C. incacorralis Herz. cf. Frahm, J. 

Hattori Bot. Lab. 44:485 (1978) = C. 

flexuosus (Hedw.) Brid. var. incacorralis 

(Herz.) J.-P. Frahm. 

refractus J.-P. Frahm, Nova Hedwigia 59: 147, 

1994. 

T: New Caledonia, Mt. Panie, low, mostly 

mossy forest 1300-1640m, H.M. Moore jr. 

10051, 30.9.72 (holotype MO, isotype hb. 

Frahm). 

renneri Herz., Bibl. Bot. 88: 5 (1920). 

T: Bolivia, Moränental Torreni, Herzog 3738 


CU: C. pittieri Williams cf. Frahm, Fl. Neotr. 

54: 159 (1991). 

renneri Herz. var. latelimbata Thér., Rev. Bryol. 

Lichenol. Ind. 1-22: 18 (1954). 

T: Bolivia, Herzog 4318b, pp. (holotype JE). 

CU: C. pittieri cf.......... 

restingae C. Müll. ex Broth., Denkschr. Ak. Wiss. 

Wien Math. Nat. Kl. 83:260 (1926). 

T: Brazil, Rio de Janeiro, Mauá, Ule 212 

(holotype H-BR; isotypes JE, KIEL, NY). 

CU: C. acervatus Mitt. cf. Frahm, Revue 

Bryol. Lich. 45:135 (1979). 

CU: occultus cf. ??? 

reticulatus Par. & Broth., Rev. Bryol. 29:63 

(1902). 

T: Guinée francaise, Iles de Los, ile Kassa,

Pobeguin 1 (PC). 

CU: Bryaceae cf. Frahm, Bryoph. Bibl. 

31:197 (1985). 

retinervis (C. Müll.) Par., Ind. Bryol. Suppl. 96 

(1900). 

B: Dicranum retinerve C. Müll., Bull. Herb. 

Boiss. 5: 552. 1897. 

T: Jamaica, Cinchona, Harris 11025 

(holotype destroyed at B; isotype, NY). 

CU: C. fragilis (Brid.) B.S.G. cf. Frahm, 


revolvens Herz. in Thér., Rev. Bryol. Lich. 11:59 

(1939). 

T: Colommbia, Florencia, Woronow 317/318 

(JE, B). 

CU: no Campylopus cf. Frahm, Bryol. Beitr. 

7: 66 (1987). 

richardii Brid., Mant. Musc. 73. (1819). 

T: Guadelupe, Parker s. n. (holotype, B). 

OU: Thysanomitrion richardii (Brid.) 

Schwaegr., Spec. Musc. Suppl. 1(2): 61 

(1823). 

ridleyi (Dix.) Manuel, Fed. Mus. Journ. 26:161 

(1981). 

B: Thysanomitrion ridleyi Dix., Bull. Torr. 

Bot. Cl. 51: 227 (1924). 

T: Malay Peninsula, Pahang, Gunong Jahan, 

Ridley s.n. (PC). 

CU: C. lorentzii cf. Frahm, Nova Hedwigia 

39:595 (1984) = C. exasperatus var. lorentzii 

cf. Frahm et al., Acta Bot. Fenn. 131:80 

(1985). 

rigens Ren. & Card., Bull. Soc. R. Bot. Belg. 

35:300 (1897). 

T: Madagascar, Perrot s.n. (holotype PC). 



Bryoph. Bibl. 31:16 81985). 



rigescens Kindb., Enum. Bryin. Exot. 89 (1889) 

nom. nud. 

rigidiusculus (Hampe) Jaeg., Ber. S. Gall. 

Naturw. Ges. 1870-71:432 (1872). 

B: Dicranum rigidiusculum Hampe, Vid. 

Medd. Naturh. Foer. Kjoebenh. ser. 3, 2: 272 

(1870). 

T: Brazil, Sierra de Piedade, Warming s.n. 



71 


rigidus (Hornsch.) Jaeg., Ber. S. Gall. Naturw. 

Ges. 1870-71:443 (1872). 

B: Thysanomitrion rigidum Hornsch., Fl. 

Bras. 1(2): 15 (1840). 

T: Brazil, Ouro Preto, Martius s. n. (not seen). 

CU: C. richardii Brid. cf. Frahm, Revue 

Bryol. Lich. 45:160 (1979). 

ripicola Besch., Ann. Sc. Nat. Bot. ser. 6,9:326 

(1880). 

T: Bourbon, Laperranche 23 (holotype PC). 

CU: C. aureus Bosch & Lac. cf. Frahm, 

Bryoph. Bibl. 31:30 (1985) = C. schmidii (C. 

Müll.) Jaeg. 

robbinsii Bartr., Rev. Bryol. Lich. 3o:187 (1962). 

T: Papua N.G., Western Highlands, Wabag 

area, near Tabanaka Wage Valley.... Robbins 

3304 (holotype FH). 

CU: C. aureus Bosch & Lac. cf. Frahm et al., 

Acta Bot. Fenn. 131:70 (1985) = C. schmidii 

(C. Müll.) Jaeg. 

roberti Par., Ind. Bryol. Suppl. 96 (1900). 





robillardei Besch., Ann. Sc. Nat. Bot. ser. 6, 9: 

323 (1880). 

T: Maurice, Robillard s.n. (holotype PC). 

robillardei var. perauriculatus (Broth.) J.-P. 


rodwayi Broth. in Rodw., Pap. Proc. R. Soc. 

Tasmania 1913:260 (1914). 


roellii Ren. & Card., Bull. Soc. R. Bot. Belg. 38:9 

(1900). 

T: Costa Rica, Juan Vinas 3400 ped., Sarg 

s.n. (not seen). 

OU: C. flexuosus (Hedw.) Brid. cf. Hagen, 

K. Norsk. Vid. Selsk. Skrift. 1914(1):75 

(1915). 

CU: C. tallulensis Sull. & Lesq. cf. Allen, 

Trop. Bryol. 1: 78 (1987). 

roinei Card. & P. Varde, Rev. Bryol. 49:37 

(1922). 

T: India, Kodaikanal, Foreau & Roiné, Musci 

Madurenses 51 (holotype PC, isotype BR). 

CU: C. involutus (C. Müll.) Jaeg. cf. Frahm, 

Bryol. Beitr. 7: 67 (1987) = C. ericoides 

(Griff.) Jaeg.

72 

roraimae Broth., Trans. Linn. Soc. Bot. ser. 

2,6:89 (1901). 

T: Venezuela. McConnell & Quelch 527 


CU: C. arctocarpus (Hornsch.) Mitt. cf. Florschütz, 

Mosses of Suriname 73 (1964). 

rosulatus (Hampe) Mitt., J. Linn. Soc. Bot. 12:79 

(1869). 

B: Dicranum rosulatum Hampe, Linnaea 32: 

139 (1863). 

T: Colombia, Bogotá la Penna et Pacho, 

Lindig 2012 (holotype not seen; isotypes H- 

BR, GOET, NY). 

CU: C. pauper (Hampe) Mitt. cf. Frahm, Fl. 

Neotr. 54: 148 (1991). 

rubiginosus Stirt., Ann. Scott. Nat. Hist. 15:109 

(1906). 

CU: C. flexuosus (Hedw.) Brid. cf. Hagen, 

K. Norsk Vid. Selsk. Skrift. 1914(1):75 

(1915). 

rubricaulis Broth. & Par., Oefv. Finsk. Vet. Soc. 

Foerh. 51A:4 (1909). 

T: Nouvelle Caledonie, Baie du Sud, Le Rat 

s.n. (isotypes B, FH, L). 

CU: C. purpureo-flavescens (C. Müll.) Par. 

cf. Frahm, Bryol. Beitr. 7: 94 (1987) = C. 

hawaiicus (C. Müll.) Jaeg. cf. Frahm, The 

Bryologist 94: 62 (1991). 

rubricaulis Lindb. in C. Müll., Gen. Musc. Fr. 

169 (1900) nom. nud. 

M: Brazil, Caldas, Lindberg s.n.(B). 

CU: C. filifolius (Hornsch.) Mitt. var. humilis 

(Mont.) J.-P. Frahm cf. Frahm, Cryptogamie 

Bryol. Lich. 2(4):445 (1981) = 

Bryohumbertia filifolia (Hornsch.) J.-P. 

Frahm var. humilis (Mont.) J.-P.Frahm. 

rubricaulis Lindberg in C. Müll., Gen. Musc. Fr. 

169. 1900, nom. nud. 

rufescens Broth. in Par., Ind. Bryol. Suppl. 96 

(1900). 

B: Dicranum rufescens C. Müll., Bull. Herb. 

Boiss. VI:34 (1898). 

T: Brazil, Minas Gerais, Serra do Ouro Preto, 

Ule 1348 (lectotype, H-BR). 


Lich. 45:156 (1979). 

rugosus Besch., Ann. Sc. Nat. Bot. ser. 5,18:198 

(1873). 

T: „in monte Cougui, Balansa 2560 cum C. 

Balansaeano socius“ (holotype PC). 


ruttneri Thér. in Froehl., Arch. Hydrobiol. Suppl. 

21:307 (1955). 

T: Sumatra, Fort de Koek supra Padäng, 

Ruttner s.n. (PC). 

CU: C. macgregorii cf. Frahm, Nova 

Hedwigia 39:595 (1984). 

ruwenzoriensis Demar. & P. Varde. Bull. Jard. 

Bot. Bruxelles 26:265 (1956). 

T: Zaire, entre Kalonge et Mahanga, Demaret 

516p (holotype BR). 


Bryoph. Bibl.31:81 (1985). 

ryparochlorus C. Müll., Gen. Musc. Fr. 276 

(1900) nom. nud. 


Bryoph. Bibl. 31:196 (1985). 

S 

saddleanus (C. Müll.) Besch., Bull. Soc. Bot. 

France 32: LVIII (1885). 

T: Saddle-Inseln, Wollaston, Hariot 163 (H). 

CU: C. vesticaulis Mitt. cf. Dixon in Christ., 

Res. Norw. Sc. Exp. Tristan da Cunha 48:18 

(1960). 

saint-pierrei Thér., Smiths. Misc. Coll. 85(4):3,2 

(1931). 

T: Mexico, Hidalgo, El Chico Saint Pierre 

1589 (PC). 

CU: C. japonicus Broth. 

salesseanus Par. & Broth., Rev. Bryol. 29:64 

(1902). 

T: Guinea, Fouta Djallon, Normand s.n. 

(isotype B, holotype PC). 


Frahm, Lindbergia 8(1):48 (1982). 

samoanus Broth. in Rech., Denkschr. Ak. Wiss. 

Wien Math. Nat. Kl. 84:387 (1908). 

T: Samoa, Berg Launtre, Rechinger 355 

(lectotypus H-BR). 


7: 68 (1987).. 

sancti-caroli Besch., J. de Bot. 5:147 (1891). 

T: Chili, San Carlos, Gay s.n. (BM). 


Bryol. Beitr. 7: 68 (1987). 

sandwicensis Par., Ind. Bryol. 97 (1900). 

T: Insulae Hawaiicae, sine loco speciali, 

Hillebrand s.n. (could not be located). 

CU: C. umbellatus (Arn.) Par. cf. Bartr.,

Bishop Mus. Bull. 101:44 (1933). 

OU: C. richardii Brid. cf. Dixon. J. Bot. 

60:287 (1922). 

santae-helenae Schimp. in Par., Ind. Bryol. 

ed.2,1:326 (1904) nom. nud. 

OU: C. helenicus (C. Müll.) Jaeg. cf. Paris 

l.c. 

CU: C. arcuatus (Brid.) Jaeg. 

sargii Roell ex Ren. & Card., Bull. Soc. R. Bot. 

Belg. 38(1):8 (1900) . 

T: Costa Rica, Sarg s. n. (isotype H-BR). 

OU: C. flexuosus (Hedw.) Brid. cf. Bartr., 


CU: C. zygodonticarpus (C. Müll.) Par. cf. 

Allen, Trop. Bryol. 1: 92 (1987). 

sauteri (B.S.G.) Par., Ind. Bryol. ed.2,1:236 

(1904). 

CU: Paraleucobryum sauteri B.S.G. 

savannarum (C. Müll.) Mitt., J. Linn. Soc. Bot. 

12:85 (1869). 

B: Dicranum savannarum C. Müll., Syn. 

Musc. 2: 596 (1851). 

T: Type. Suriname, Kegel s. n. (holotype 

destroyed at B; lectotype BM; isolectotypes 

H-BR, JE, L). 

savannarum ssp. bartletii (Bartr.) Florsch., 

Mosses Suriname 79 (1964). 

B: Campylopus bartletii Bartr. 

CU: C. savannarum mod. bartlettii. 

saxatilis Williams, N. Am. Fl. 15:137 (1913). 

T: Jamaica, Cinchona, Harris 11143 

(holotype NY, isotypes DUKE, F). 

CU: C. cygneus cf. Frahm, Nova Hedwigia 

41:277 (1985). 

saxicola (Web. & Mohr) Brid., Mant. Musc. 72 

(1819). 

CU: Campylostelium saxicola (Web. & 

Mohr) B.S.G. 

saxicola Broth. in Skottsb., Nat. Hist. Juan 

Fernandez 2(3):243 (1924) hom. illeg. 

OU: C. pascuanus Frahm nom. nov. 

CU: C. hygrophilus Broth. cf. Frahm, Nova 

Hedwigia 34:392 (1981) = C. vesticaulis Mitt. 

cf. Frahm, Bryol. Beitr. 7: 68 (1987). 

scaberrimus Broth. in Dus., Rep. Princeton Univ. 

Exp. Patag. 8:72 (1903) nom. nud. 

CU: Chorisodontium aciphyllum (Hook.f. & 

Wils.) Broth. cf. Card. in Broth., Nat. Pfl. 

1(3):1183 (1909). 

scabrellus Mitt., J. Linn. Soc. Bot. 12:86 (1869). 

73 

T: Brazil, without locality, Lobb s. n. 



Lichen. 45:156 (1979). 

scabridorsus Dix., Hong Kong Natural. Suppl. 

2:5 (1933). 

T: Hong Kong, repusle Bay, Ah. Nin H22A 

(isotypes BR, L). 


7: 91 (1987). 

scabridii Bealey in Dix., New Zeal. Inst. Bull. 

3(3):88 (1923) nom. nud. 

CU: C. clavatus (R.Brown) Wils. cf. Dixon 

l.c. 

scabricuspis Mitt. in Mell., S. Helena 358 (1875). 

T: St. Helena, Melliss 9 (holotype NY). 

CU: C. flexuosus cf. Frahm, Bryol. Beitr. 7: 

90 (1987). 

scabridorsus Dix., Hong Kong Natural. Suppl. 

2:5 (1933). 


7: 68 (1987). 

scabripilus Warnst., Hedwigia 57:80 (1915) . 



OU: C. richardii cf. Takaki, J. Hatt. Bot. Lab. 

30:236 (1967). 

scabrophyllus (C. Müll.) Par., Ind. Bryol. Suppl. 

97 (1900). 

B: Dicranum scabrophyllum C. Müll., 

Hedwigia 37: 98 (1898). 

T: Uruguay, Montevideo, Arechavaleta 232 

(holotype destroyed at B, lectotypus nov., 

NY). 

CU: C. griseus (Hornsch.) Mitt. cf. Frahm, 


schenkei C. Müll. in Par., Ind. Bryol. ed.2,1:326 

(1904) nom. nud. 

HS: C.schemkei Par., Ind. Bryol. 260, (1894). 

OU: C. bicolor ssp. atroluteus cf. Frahm, 

Bryol. Beitr. 7: 90 (1987). 

CU: C. bicolor (C. Müll.) Wils. 

schiffneri (Broth.) J.-P.Frahm, J. Bryol. 8:260 

(1974). 

B: Thysanomitrion schiffneri Broth., 

Denkschr. Akad. Wiss. Wien math.-nat. Kl. 

83: 265 (1926). 

T: Brazil, Sao Paulo, Campo Grande, 

Schiffner 560 (holotype H-BR). 

CU: C. savannarum (C. Müll.) Mitt. cf.

74 


schimperi Milde, Bot. Zeit. 22 Beil.:13 (1864). 

T: Italien: Am Fuß der Zielalpe bei Partschins 

bei Meran 1600ám Wasser einer Wasserleitung. 

Milde (20.10.1863) Bryoth. europ. 658 

(FB, FR, GOET, JE, U). 

schmidii (C. Müll.) Jaeg., Ber. S. Gall. Naturw. 

Ges. 1870-71:439 (1872) (39). 

B: Dicranum schmidii C. Müll., Bot. Zeit. 11: 

37 (1853). 

T: „Ost-Ind., Neilgheries, Schmid s.n. (BM, 

L). 

schroederi Broth., Hedwigia 52:307 (1912). 

T: Kilimandscharo, Schroeder (9/1910) s.n. 


CU: Campylopus filifolius cf. Frahm, 

Lindbergia 8:51 (1982) = Bryohumbertia 

spec. cf. Frahm, Bryoph. Bibl. 31:197 (1985). 

schunkei Rehm. in Dix. & Gepp, Bull. Misc. Inf. 

Kew 1923:197 (1923) nom. nud. 

OU: C. schenkei C. Müll. 

CU: C. bicolor ssp. atroluteus cf. Frahm, 

Bryoph. Bibl. 31:191 (1985). 

schwaegrichenii Duby, Mem. Soc. Phys. Hist. 

Nat. Geneve 20:360 (1870). 

T: Brazil, Porto de Estrelle, Beyrich s. no. 

(holotype, G). 

CU: C. trachyblepharon cf. Frahm, Revue 

Bryol. Lich. 45:164 (1979). 

schwarzii Schimp., Musci Eur. Nov. Bryol. Eur. 

Suppl. fasc.1-2, Campylop. 1 (1864). 

T: Österreich, Velbertauern, Dr. Schwarz 

(16.8.1858) s.n. (JE). 

CU: C. gracilis (Mitt.) Jaeg. cf. Frahm, Nova 

Hedwigia 59: 152 (1994). 

sciuroides Welw. & Duby, Mem. Phys. Hist. Nat. 

Geneve 21:220 (1870). 

T: Angola, Distr. Pungo Adongo, Welwitsch 

10 (G, lectotype). 



sclerodictyum Card., Annuair. Cons. Jard. Bot. 

Geneve 15-16:160 (1912). 

T: Java, M. Gedeh 2700-2800 m, Hochreuter 

278 p. pte (isotypes FH, L). 

CU: C. exasperatus cf. Frahm, Bryol. Beitr. 

7: 70 (1987). 

scopelliformis (C. Müll.) Broth., Nat. Pfl. 

1(3):334 (1901). 

B: Dicranum scopelliforme C. Müll., 

Hedwigia 39: 248 (1900). 

T: Sta. Catarina, Serra do Oratorio, Ule 706 

(holotype destroyed at B; lectotype, H-BR). 


caldensis (Aongstr.) J.-P.Frahm cf. Frahm, 


scoposum Hampe ex C. Müll., Gen. Musc. Fr. 

265 (1900). (*) 

scottianus (Turn.) Brid., Mant. Musc. 72(1819), 

C. scoltianus Brid. l.c. 

CU: Dicranum scottianum Turn. 

scrabricuspis Mitt. in Mell., S. Helena 358 

(1875). 

CU: C. flexuosus (Hedw.)Brid. cf. Frahm, 

Bryol. Beitr. 7: 68 (1987). 

searellii R.Brown ter., Trans. New Zealand Inst. 

29:468 (1897).(*) 

sedgwickii Dix., J. Bot. 48:303 (1910). 

T: India, Western Ghats, Mahableshevar, on 

earth, Sedgwick s.n. (isotype H-BR). 

CU: C. recurvus (Mitt.) Jaeg. cf. Frahm, Trop. 

Bryol. 4: 60 (1991). 

sehnemii Bartr., J. Wash. Ac. Soc. 42(6): 179. 

1952. 

T: Brazil, Rio Grande do Sul, Campestre 

Montenegro, Sehnem 2310 (holotype FH). 

CU: C. julicaulis Broth. cf. Frahm, Nova 

Hedwigia 59: 152 (1994). 

sellowianus (Hampe) Jaeg., Ber. S. Gall. Naturw. 

Ges. 1877-78:385 (1880). 

HS: C. sellowii C. Müll. ex Kindb., Enum, 

Bryin. Exot. 51 (1888). 

B: Dicranum sellowianum Hampe, Vidensk. 

Medd. Naturh. For. Kjoebenh. ser. 3,4: 43 

(1872), nom. illeg. pro Thysanomitrion 

scabrisetum Hampe. 

T: Brazil. Sellow s. n. (not seen). 

CU: C. arctocarpus (Hornsch.) Mitt. cf. Mitten, 

J. Linn. Soc. Bot. 12:8 (1869). 

semidorsum Rehm. ex Par., Ind. Bryol. 260 

(1894) nom. nud. 

CU: C. catarractilis (C. Müll.) Par. cf. Sim, 


OU: C. atroluteus (C. Müll.) Par. cf. Magill, 

Fl. S. Afr. Bryoph. 1(1):148 (1981). 

senex (C. Müll.) Par., Ind. Bryol. Suppl. 97 

(1900). 

B: Dicranum senex C. Müll. Hedwigia 36: 

351 (1897). 

T: Australia, Victoria, Luehmann s.n.; Hume

River, Campbell s.n. (syntypes, not seen). 


J. Bryol. 13:359 (1985). 

senilis Lor. in C. Müll., Gen. Musc. Fr. 265 

(1900) nom. nud. 

sericeoides Dix., J. Linn. Soc. Bot. 50:71 (1935). 

T: N-Borneo (Mt. Kinabalu), Pakka (Cave) 

10200´, Holthum 25/89 (holotype BM). 

sericeus Fleisch., Musci Fl. Buitenzorg 1:99 

(1904). 

T: Java, Pangerango, Fleischer (1898) s.n. 

(holotype FH, isotype L). 



sericeus Negri, Annali Bot. 7:162 (1908) hom. 

illeg. 

T: M. Ruwenzori prope Bujongolo ca. 3800 

m ad cortices. 

CU: C. stramineus (Mitt.) Jaeg. cf. Dixon, 

Smiths. Misc. Coll. 69(2):10 (1918) = C. 

nivalis (Brid.) Brid. 

serratus Lac., Natuurk. K. Ak. Wet. Amsterdam 

13:11 (1872). 

T: Banca VII/58 leg. Kurz ex hb. Sande 

Lacoste (isotype H-BR). 

serricuspes Thér. & P. Varde Rev. Bryol. Lich. 

6:135 ( 1934). 

T:“Africa trop. occ., Gabon“ Le Testu, Musci 

selecti et critici 12 (isotypes NY,L,U,B). 



(1985). 

serridorsus (C. Müll.) Par., Ind. Bryol. Suppl. 97 

(1900). 

B: Dicranum serridorsum C. Müll., Hedwigia 

38: 84 (1899). 

T: Cape, Table Mountain, Rehmann 45. 

CU: C. inchangae cf. Magill, Bryoph. S. Afr. 

1(1):146 (1981) = C. robillardei Besch. 

OU: C. catarractilis (C. Müll.) Par. cf. Sim, 


serrifolius (Broth. & Par.) Frahm, 

B: Pilopogon serrifolius B roth. & Par., Oefv. 

Finsk. Vet. Akad. Soc. Foerh. 53A(11): 7 

(1911). 

T: Neukaledonien, Le rat (7/1909) s.n. (H- 

BR, S). 

OU: Thysanomitrion serrifolium (Broth. & 

Par.) Broth., Nat. Pfl. ed. 2,10: 189 (1924). 

serripilus C. Müll., Gen. Musc. Fr. 265 (1900) 

75 

nom. nud. 

setaceo-rigidus (Hampe) Jaeg., Ber. S. Gall. 

Naturw. Ges. 1877-78:496 (1880). 

B: Dicranum setaceo-rigidum Hampe, Vid. 

Medd. Naturh. Foer. Kjoebenhavn ser. 3,9- 

10:257 (1878). T: Brazil, „ad Rio Preto“, 

Glaziou 9077 (isotypes NY, PC). 

CU: C. aemulans cf. Frahm, Revue Bryol. 

Lich. 45:135 (1979). 

setaceus Card., Ann. Missouri Bot. Garden 8:54 

(1897). 

T: Azores, Fayal, C.S. Brown (1894) s.n. 


CU: C. cygneus cf. Frahm, Nova Hedwigia 

41:277 (1985). 

setifolius Wils., Bryol. Brit. 89 (1855). 

T: In rocky places on mountains. On Carrig 

Mountain near Dunkerron, Ireland, Dr. 

Taylor (holotype BM). 

seuberti C. Müll., Gen. Musc. fr. 271 (1900) nom. 

nud. 

sharpii J.P. Frahm, Horton & Vitt Bryologist 

82: 623 (1979). 

T: Mexico, Oaxaca: Near S. Pedro Yolox, 

17°36' N, 96°24' W, Vitt 17715 (ALTA 

TENN). 

shawii Wilson in Hunt, Mem. Lit. Phil. Soc. 

Manchester ser. 3,3: 234 (1868). 

T: Outer Hebrides, Shaw s. no. (holotype 

BM). 

siamensis Dix. & Thér., J. Siam Soc. Nat. Hist. 

Suppl. 9(1):8 (1932). 

T: Siam, Udawu Loi, Kao Krading 1200 m, 

Kerr 85 (BM). 



siamensis var. subsiamensis Dix. & Thér. l.c. 

T: Rangala, Ceylon, Alston 1502 (BM). 



silvaticus Rehm. in Par., Ind. Bryol. ed.2,1:327 

(1904) nom. nud. 

M: Südafrika, Rehmann 50 (PC). 

CU: C. praetermissus J.-P.Frahm cf. Frahm, 

Bryoph. Bibl. 31:152 (1985). 

simii Schelpe, Mem. bot. Surv. S. Afr. 43:5 

(1979) nom. nov. pro C. pseudojulaceus Sim 

hom. illeg. 

T: as in C. pseudojulaceus. 

CU: C. julaceus ssp. arbogastii cf. Frahm,

76 

Bryoph. Bibl. 31:109 (1985). 

sinensis (C. Müll.) J.-P. Frahm, Ann. Bot. Fenn. 

34: 202, 1997. 

B: Dicranum sinense C. Müll., Nuov. Giorn. 

Bot. Ital. n. ser. 4: 249, 1897 

OU: Dicranodontium sinense (C. Müll.) Par., 

Ind. Bryol. Suppl. 119, 1900 

T: China, „China interior, provincia Schensi 

sept., in monte Ki-san“, Goraldi 1490 

(Lectotypus H-BR, cf. Frahm 1997) 

singaporensis Fleisch. ex Par., Rev. Bryol. 33:25 

(1906). 

T: Singapore, im botan. Garten auf Wiesen 

zwischen Gras wachsend, Fleischer Musci Fr. 

Arch. Ind. 403 (NY, PC). 

CU: C. serratus Lac. cf. Dixon, Gard. Bull. 

Straits. Settl. 4:5 (1926). 

skottsbergii Broth., Bishop Mus. Bull. 40:6 

(1927). 

T: Hawaii, summit of Puu Kukui, Skottsberg 

1740 (lectotype H-BR). 

CU: C. schmidii ssp. hemitrichius (C. Müll.) 

J.-P. Frahm cf. Frahm, The Bryologist 94: 64 

(1991). 

smaragdinus (Brid.) Jaeg., Ber. S. Gall. 

Naturwiss. Ges. 1870-71: 432 (1872). 

B: Didymodon smaragdinus Brid. 

T: Ascension, Lesson s.n. (isotypes BM, PC). 

OU: Dicranum smaragdinum (Brid.). C. 

Müll., Syn. Musc. 1: 401 (1848). 

sommieri Bott. in Giac., Atti Ist. Univ. Lab. Critt. 

Pavia ser.5,13:63 (1955) nom. nud. 

OU: C. introflexus (Hedw.) Brid. var. 

vaporarius (De Not.) Moenk. cf. Giacomini 

l.c. 


sordidus (Mitt.) Jaeg., Ber. S. Gall. Naturw. ges. 

1870-71:434 (1872). 

CU: Blindia sordida (Mitt.) C. Müll. 

sparksii R. Brown ter., Trans. New Zealand Inst. 

29:467 (1897). 

CU: C. pallidus Hook.f. & Wils. cf. Dixon, 

New Zealand Inst. Bull. 3(3):86 (1923) = C. 

pyriformis (Schultz) Brid. 

spegazzinii (C. Müll.) Par., Ind. Bryol. 261 

(1894). 

CU: Chorisodontium spegazzinii (C. Müll.) 

Roiv. 

sphagnicola Herz., Rep. Spec. Nov. Regn. Veg. 

21:28 (1925). 

T: Brazil, Serra dos Orgaos, Morro Assu, v. 

Lützelburg 6760b (holotype, JE). 


fragiliformis (J.-P. Frahm) J.-P. Frahm . 

spiralis Dus., Ark. Bot. 4(13):4 (1905). 

spirifolius Herz., Beih. Bot. Centralbl. 26(2):55 

(1909). 

CU: Ditrichum bogotense (Hampe) Broth. 

spirothecius C. Müll. in Polak., Verh. Bot. Ver. 

Brandenburg 19:68 (1878) nom. nud. 

sprucei Mitt., J. Linn. Soc. Bot. 12:81 (1869). 

T: Brazil, Rio Negro, Spruce 59 (holotype 

NY; isotypes F, FH, H-BR, H-SOL). 


Frahm, J. Bryol. 11:213 (1980). 

spurio-concolor (C. Müll.) Par., Ind. Bryol. 

Suppl. 97 (1900). 

B: Dicranum spurio-concolor C. Müll., Nuov. 

Giorn. Bot. Ital., n. ser. 4:34 (1897). 

T: Bolivia, Yungas, Rusby 3115 (holotype 



39:155 (1984). 

squalidus Brid., Mant. Musc. 77 (1819). 

CU: Racomitrium lanuginosum (Hedw.) Brid. 

var. squalidum (Brid.) Brid. 

standaertii Broth. & Par., Nat. Pfl. 1(3):1185 

(1909) nom. nud. 

CU: Pilopogon laevis (Tayl.) Thér. cf. Frahm, 

J. Hattori Bot. Lab. 44:484 (1978). 

standleyi Bartr., Contr. U.S. Nat. Herb. 26:59 

(1928). 

T: Costa Rica, Cerro de las Vueltas, Standley 

& Valerio 43939 (holotype FH; isotypes JE, 

NY). 

CU: C. jamesonii (Hook.) Jaeg. cf. Bartr., 


OU: C. standleyi cf. Allen, Trop. Bryol. 1: 

90 (1989). 

stenocarpus (Hampe) Jaeg., Ber. S. Gall. Naturw. 

Ges. 1871-72:434 (1872). 

B: Thysanomitrium stenocarpum Hampe 

T: Brasilia, Joinville (GOET). 

CU: Dicranum stenocarpum (Hampe) C. 

Müll. Syn. Musc. 2: 600 (1851). 

stenopelma (C. Müll.) Rehm. ex Par., Ind. Bryol. 

Suppl. 97 (1900). 

B: Dicranum stenopelma C. Müll. Hedwigia 

38: 83 (1899). 

T: Cape, Knysna, near Esternek, Rehmann 52

(G, PC). 

CU: C. controversus (Hampe) Jaeg. cf. 

Frahm, Hikobia 11 (1994) = C. thwaitesii. 

stewartii R.Brown ter., Trans. New Zeal. Inst. 

29:472 (1897). 

CU: C. bicolor (C. Müll.) Wils. cf. Dixon, 

New Zealand Inst. Bull. 3(3):86 (1923). 

steyermarkii Bartr., Bot. Soc. Venez. Cienc. Nat. 

106:35 (1963). 

T: Venezuela. Cabeceras de Rio Chicanán, 

Steyermark 89600 (holotype FH; isotype 

VEN). 

CU: C. richardii Brid. cf. Frahm, Bryol. Beitr. 

7: 97 (1987). 

stramineolus C. Müll., Gen. Musc. Fr. 270 (1900) 

nom. nud. 

CU: C. pleurocarpus cf. Frahm, Revue Bryol. 

Lich. 42:609 (1976) = C. controversus 

(Hampe) Jaeg. = C. thwaitesii. 

stramineus (Mitt.) Jaeg., Ber. S. Gall. Naturw. 

Ges. 1877-78:382 (1880). 

T: Dicranum stramineum Mitt. J. Linn. Soc. 

Bot. 7: 148 (1864). 

T: Mount Cameroon 9000 ft., Mann s.n. 

(FH,NY). 


Lindbergia 8:46 (1982) = C. nivalis (Brid.) 

Brid. 

straminifolius Bartr., Contr. U.S. Nat. Herb. 

26(3):63 (1928). 

T: Costa Rica, Standley 51212 (holotype FH). 

OU: C. flexuosus (Hedw.) Brid. cf. Bartr., 


CU: C. densicoma (C. Müll.) Par. cf. Allen, 

Trop. Bryol. 1: 82 (1989). 

stricticaulis (C. Müll.) Par., Ind. Bryol. Suppl. 

97 (1900). 

B: Dicranum stricticaule C.Müll., Bull. Herb. 

Boiss. 6:38. 1898. 


(holotype destroyed at B; lectotype, H-BR, 



Lich. 45:135 (1979). 

strictifolius Broth., Act. Soc. Sc. Fenn. 19(5):9 

(1891). 

T: Brazil Minas Gerais: Caraca, Waino s.n. 


CU: C. arctocarpus (Hornsch.) Mitt. fo. 

strictifolius (Broth.) J.-P.Frahm, Revue Bryol. 

77 

Lich. 45:138 (1979). 

strictisetus (C. Müll.) Broth., Nat. Pfl. 1(3):334 

(1901). 

B: Dicranum strictisetum C. Müll., Hedwigia 

39: 253 (1900). 

T: Brazil, Minas Gerais, Caraca, Ule 1357 


CU: Campylopus filifolius (Hornsch.) Mitt. 

var. humilis (Mont.) J.-P.Frahm cf. Frahm, 

Revue Bryol. Lich. 45:146 (1979) = 


var. humilis (Mont.) J.-P.Frahm. 

strictissimus C. Müll., Gen. Musc. Fr. 273 (1900) 

nom. nud. 

CU: C. appressifolius Mitt. cf. Dixon, New 

Zealand Inst. Bull. 3(3):89 (1923) = C. 

clavatus (R. Brown) Wils. cf. Sainsb., Trans. 

New Zealand Inst. 59:506 (1928). 

strictulus (C. Müll.) Robinson, The Bryologist 

70:20 (1967) 

B: Dicranum strictulum C.Müll., Linnaea 38: 

587 (1874). 

T: Mexico, Rio de Orizaba, F. Müll. s. n. (not 

seen). 

OU: Atractylocarpus strictulus (C. Müll.) J.- 

P.Frahm, Journ. Hattori Bot. Lab. 44:484 

(1978). 

CU: A. longisetus (Hook.) Bartr. cf. Padberg 

& Frahm, Cryptogamie Bryol. Lich. 6:326 

(1985). 

strictus C. Müll., Linnaea 19:207 (1846). 

T: Colombia, Moritz s.n. (BM). 

CU: C. pilifer Brid. cf. Frahm, Bryol. Beitr. 

7: 73 (1987). 

strictus Schimp. ex Besch., Mem. Soc. Sc. Nat. 

Cherbourg 16:167 (1872) hom. illeg. 

T: as in C. strictus C. Müll. 


N. Am. Fl. 15(2):144 (1913). 


strictus Tayl. in Hook.f., Fl. Nov. Zel. 2:68 (1854) 

nom. nud. 

OU: C. xanthophyllus Mont. cf. Hooker f. l.c. 


J. Linn. Soc. Bot. 12:84 (1869). 

stygius Schimp. in C. Müll., Gen. Musc. Fr. 270 

(1900) nom. nud. 

subambiguus Thér., Rec. Publ. Soc. Havraise 

Etud. Div. 1925:127 (1926). 

T: Madagascar, Mt. Tsarantanana 1200 m,

78 

Perrier de la Bathie s.n. (holotype PC). 



subannotinus Thér. & P. Varde, Rev. Bryol. Lich. 

11:48 (1939). 

T: Bolivia, Comarapa, Laguna Verde, Herzog 

4301 (holotype PC; isotype JE). 

CU: C. multicapsularis cf. Frahm, Journ. 

Hattori Bot. Lab. 48:209 (1980) = C. nivalis 

(Brid.) Brid. var. multicapsularis (C. Müll.) 

J.-P. Frahm. 

subappressifolius Broth. & Geh. in Rodw., Pap. 

Proc. R. Soc. Tasmania 1912:97 (1913). 

T: Tasmania, Jones Track 700 ft., Moore 59 

(lectotype H-BR). 

CU: C. flavonigritus cf. Catcheside & Frahm, 

J. Bryol. 13:365 (1985). 

subarctocarpus (Hampe) Jaeg., Ber. S. Gall. 

Naturw. Ges. 1870-71:496 (1880). 

B: Dicranum subarctocarpum Hampe, 

Vidensk. Medd. Naturh. For. Kjoebenh. ser. 

3, 9-10: 255 (1878). T: Brazil. Rio de Janeiro, 

Rio Preto, Glaziou 9078 (lectotype, BM; 

isolectotypes H-BR, NY). 

CU: C. filifolius (Hornsch.) Mitt. var. humilis 

(Mont.) J.-P.Frahm cf. Frahm, Revue Bryol. 

Lich. 45:146 (1979) = Bryohumbertia filifolia 

var. humilis. 

subarenicola C. Müll. ex Broth., Bih. K. Svensk. 

Vet. Ak. Foerh. 21 Afd. 3(3):7 (1895). 

T: Brazil, Santos, Mosén 94 (holotype 



Bryol. Lich. 45:164 (1979). 

subareodictyon Broth. in Skottsb., Nat. Hist. Juan 

Fernandez 2(3):415 (1924) . 

CU: C. blindioides Broth. in Skottsb. cf. 

Robinson, Smiths. Contr. Bot. 27:26 (1975) 

= C. pyriformis (Schultz) Brid. cf. Frahm, 

Bryol.Beitr. 7: 88 (1987). 

subbartramiaceus C. Müll. in Geh., Rev. Bryol. 

5:69 (1878). 

M: Cape Town, Rehmann 55 (lectotypus L, 

isolectotypes NY,PC). 

OU: C. leucobasis Par. var. bartramiaceus (C. 

Müll.) Par., C. introflexus (Hedw.) Brid. cf. 

Sim, Trans. R. Soc. S. Afr. 15:171 (1926). 


31:143 (1985). 

subbicolor Geh. & Broth. in C. Müll., Gen. Musc. 

Fr. 275 (1900) nom. nud. 

CU: C. bicolor cf. herb. mat. in NSW. 

subbrachymitrius C. Müll., Hedwigia 38 

Beibl.:57 (1899) nom. nud. 

M: Brazil, Ule 289, 433, 636 (H-BR). 

CU: C. brachymitrius = C. occultus. 

subbrachyphyllus Par., Ind. Bryol. 261 (1894) 

nom. nud. 

M: Cameroun, Mt. Rumpi, 1800 m, Dusén 

s.n. (BR). 


Frahm, Brol. Beitr. 7: 90 (1987).. 

subcapillatus Watts. & Whitel., Proc. Linn. Soc. 

N.S. Wales Suppl. 27:45 (1902) nom. nud. 

subcaudatus Herz., Hedwigia 66:343 (1926). 

T: Sumatra, Sibaganding, Tobasee, Heusser 

s.n. (holotypeJE, isotype H-BR). 

OU: ? Campylopus walkeri cf. Frahm, J. 

Bryol. 13:185 (1984); ? Bryohumbertia 

walkeri (Mitt.) J.-P. Frahm. 



subchlorophyllosus C. Müll. ex Rabenh., Bryoth. 

Eur. 28 nr. 1362 (1884). 

T: Cape, Somerset-East, Boschberg, 

MacOwan s.n. (BM,L). 

CU: Sphaerothecium subchlorophyllosum (C. 

Müll. ex Rabenh.) J.-P. Frahm. 

subcinereus Stirt., Ann. Scott. Nat. Hist. 8:105 

(1899). 

CU: C. flexuosus (Hedw.) Brid. cf. Dixon, 

Stud. Handb. Brit. Moss. ed 3:95 (1924). 

subclavatus Kindb., Enum. Bryin. Exot. 89 

(1889) nom. nud. 

CU: C. introflexus cf. Sim, Trans. R. Soc. S. 

Afr. 15:171 (1926). 

subcomatus (Ren. & Card.) Thér. Bull. Soc. R. 

Bot. Belg. 35 (1): 302 (1897). 

T: Madagascar, Andevorante, Perrot s.n. 


CU: C. trachyblepharon (C. Müll.) Mitt. ssp. 

comatus (Ren. & Card.) J.-P.Frahm, Bryoph. 

Bibl. 31:187 (1985). 

subcomosus Dix., J. Linn. Soc. Bot. 45:498 

(1922). 

T: New Guinea, Port Moresby Distr., Mt. 

Darigolo, Clark 1 (holotype BM). 


P.Frahm cf. Frahm, Bryol. Beitr. 7: 74, 1987, 

Campylopus comosus cf. Eddy, Malesian

Mosses vol. 1 (1988). 



subconcolor (Hampe) Mitt., J. Linn. Soc. Bot. 

12: 86 (1869). 

B: Dicranum subconcolor Hampe, Linnaea 

32: 138 (1863). 

T: Colombia, Andes Bogotenses, La Penna, 

„in sylvis ad latera Barrancas 2900 m“, Lindig 

213b (isotype FH). 

CU: C. leucognodes (C. Müll.) Par. cf. Florschütz-de 

Waard & Worrell-Schets, Proc. 

Kon. Nederl. Akad. Wetensch. ser C vol. 83 

(1):42 (1980) = C. areodictyon (C. Müll.) 

Mitt. cf. Frahm, Bryol. Beitr. 7: 75 (1987). 

subcubitus Williams, Bull. New York Bot. Garden 

3(9):111 (1903). 

T: Bolivia, lower Sorata River, Williams 1749 

(holotype NY; isotypes FH JE, H-BR). 



subcuspidatus (Hampe) Jaeg., Ber. S. Gall. 

Naturw. Ges. 1870-71: 441 (1872). 

B: Dicranum subcuspidatum Hampe, 

Vidensk. Meddel. Dansk Naturhist. 

Foren. Kjobenhavn 1870: 273. 

T: Brazil, Rio de Janeiro, Tijuca, Glaziou 

7096 (holotype BM; isotypes NY,PC). 

subcuspidatus (Hampe) Jaeg. var. damazii 

(Broth.) J.P.Frahm, Bryol. Beitr. 7: 75 (1987). 

B: C. damazii Broth. 

subericoides Williams, Bull. New York Bot. 

Garden 8:336 (1914). 

T: Philippines, Mt. Santo Tomas, leg. ? 1857 

(NY). 



suberythrocaulon Broth. in Mildbr., Deutsch. 

Zentral Afr. Exp. 2:141 (1910). 

T: Bukoba, Totensinsel im Viktoria-See, 

Mildbraed 38 (B). 

CU: C. hensii Ren. & Card. cf. Frahm, 

Bryoph. Bibl. 31:87 (1985). 

subexasperatus (C. Müll.) Jaeg., Ber. S. Gall. 

Naturw. Ges. 1877-78:384 (1880). 

B: Dicranum subexasperatum C. Müll., 

Linnaea 37: 169 (1872). 

CU: C. umbellatus (Arn.) Par. cf. Bartr., Philip. 

J. Sc. 68:44 (1939). 

OU: Thysanomitrion subexasperatum (C. 

79 

Müll.) Broth., Nat. Pfl. ed. 2,10: 189 (1924); 

C. richardii Brid. cf. Dixon, J. Bot. 60:287 

(1922). 

subfalcatus (Hornsch.) Jaeg., Ber. S. Gall. 

Naturw. Ges. 1870-71:442 (1872). (*) 

subfimbriatus Thér., Rev. Bryol. Lich. 11:44 

(1939). 

T: Bolivia, Illampu, Troll 69 (lectotype, PC). 


J. Hattori Bot. Lab. 48:209 (1980) = C. nivalis 

(Brid.) Brid. 

subflexuosus Par., Ind. Bryol. 262 (1894) nom. 

nud., nom. nov. pro C. abyssinicus Schimp. 

nom. nud. 


Bryoph. Bibl. 31:187 (1985). 

subfragilis Ren. & Card., Bull. Soc. R. Bot. Belg. 

34(2):59 (1896). 


Bryol. Beitr. 7: 95 (1987). 

subfulvus Thér., Rev. Bryol. Lich. 11:45 (1939). 

T: Bolivia, Yungas of La Paz, Troll 38a 

(holotype, PC). 

CU: C. luteus cf. Frahm, Bryol. Beitr. 7: 75 

(1987). 

subgiganteus Broth., Bull. Soc. Bot. Ital. 1904:15 

(1904) nom. nud. 

subgracilis Ren. & Card. ex Gang., Bull. Soc. 

Bot. Bengal. 14:26 (1960). 

T: „Sikkim, Gollan 3222“. 

OU: C. zollingerianus Mitt. cf. Frahm, Bryol. 

Beitr. 7: 75 (1991). 



subgriseus (Hampe) Jaeg., Ber. S. Gall. Naturw. 

Ges. 1877-78:384 (1880). 

B: Dicranum subgriseum Hampe, Vid. Medd. 

Naturh. For. Kjoebenh. ser. 3,4: 47 (1872). 

T: Brazil, Petropolis, Glaziou 4543 (isotypes 

H-BR,NY, PC). 

CU: C. griseus cf. Frahm, Revue Bryol. Lich. 

42:606 (1976). 

subincacorralis Thér., Rev. Bryol. Lich. 11:44 

(1939). 

T: Bolivia, Tablas 3400 m, Herzog 2799 

(isotype B). 

CU: C. longisubulatus cf. Frahm, Nova 

Hedwigia 39:159 (1984) = C. anderssonii (C. 

Müll.) Jaeg. 

subincrassatus (Hampe) Jaeg., Ber. S. Gall.

80 

Naturw. Ges. 1877-78:384 (1880). 

B: Dicranum subincrassatum Hampe, Vid. 

Medd. Naturh. Foer. Kjoebenh. ser. 3,4:47 

(1872). 

T: „Brasilia australis“, Glaziou 4507 (isotype 

H-BR). 


Lich. 45:135 (1979). 

subintroflexus Kindb. Rev. Bryol. 25:91 (1898). 

T: Azoren, S. Miguel, Machado s.n. (S). 

CU:C. polytrichoides De Not. cf. Frahm, Revue 

Bryol. Lich. 41:331 (1975) = C. pilifer 

Brid. 

subjugorum Broth., Bot. Jahrb. 49:174 (1912). 

T: Bolivia, Cordillera Real, Knoche 60a 

(holotype H-BR) 

subleptodrepanium Broth. & Thér., Bull. Mus. 

Hist. Nat. Paris 18:476 (1912). 

T: Cote dÍvoire, Chevalier s.n. (BM). 

CU: C. nivalis cf. descr. in Bizot & Kilbertus, 

Revue Bryol. Lich. 45:81-85 (1979). 

subleucogaster (C. Müll.) Jaeg., Ber. S. Gall. 

Naturw. Ges. 1877-78:381 (1880). 

B: Dicranum subleucogaster C. Müll., Bull. 

Torrey Bot. Club 5: 49 (1879). 

T: Alabama, Mobile, Mohr 1868 (holotype 

destroyed at B; lectotype NY). 

CU: C. surinamensis cf. Frahm, Bryologist 

83:581 (1980). 

subluteus (Mitt.) Jaeg., Ber. S. Gall. Naturw. Ges. 

1870-71:417 (1872). 

B: Dicranum subluteum Mitt., Kew J. Bot. 8: 

353 (1856). 

T: Birma, Moulmein, Rev. Parish (BM). 

subnanus (C. Müll.) Jaeg., Ber. S. Gall. Naturw. 

Ges. 1870-71:415 (1872). 

B: Dicranum subnanum C. Müll., Bot. Zeit. 

17: 190 (1859). 

OU: Microcampylopus subnanus (C. Müll.) 

Fleisch. 

CU: Microcampylopus khasianus (Griff.) 

Giese & J.-P.Frahm cf. Giese & J.-P.Frahm, 

Lindbergia 11: 119 (1985). 

subnitens Kaal., Nyt Mag. Naturvid. 50: 100 

(1912). 

suboblongus Herz., Beih. Bot. Centralbl. 61B:585 

(1942). 

T: Ecuador, without locality, Espinoza 29 


CU: C. chrismarii var. suboblongus (Herz.) 

J.-P.Frahm, Journ. Hattori Bot. Lab. 48:209 

(1980) = C. nivalis (Brid.) Brid. 

subobrutus Thér. & P. Varde, Rev. Bryol. n. ser. 

4:66 (1931). 

T: Gabun, Issele, Le Testu 3882 (holotype 

PC). 

CU: C. obrutus cf. Frahm, Nova Hedwigia 

39:595 (1984). 

subpenicillatus (C. Müll.) Broth., Nat. Pfl. 

1(3):333 (1901). 

B: Dicranum subpenicillatum C. Müll., 

Hedwigia 39: 257 (1900). 

T: Brazil, Sa. Catarina, Blumenau, Ule s. n. 

(holotype destroyed at B; isoytype, H-BR). 

CU: C. lamellinervis cf. Frahm, Rev. Bryol. 

Lich. 45:154 (1979). 

subperichaetialis Biz. & Kilb. in Biz., Rev. Bryol. 

Lich. 40:113 (1974) nom. invalid. 

T: Tanzania, Bondwa Peak 1950-2100 m, 

Pocs & Gibbon 6052 (B, PC). 

OU: C. subperichaetialis Biz. & Kilb. ex 

Pocs, Folia Hist. Nat. Mus. Matr. 4:28 (1977). 

CU: C. incacorralis Herz. cf. Frahm, 

Lindbergia 8(1):48 (1982) = C. flexuosus 

(Hedw.) Brid. ssp. incacorralis (Herz.) J.-P. 

Frahm. 

subpolyanthus Par., Ind. Bryol. 262 (1894) nom. 

nud. 

subproliferus C. Müll. in Ren. & Card., Bull. Soc. 

R. Bot. Belg. 31(1):149 (1893). 

OU: C. introflexus (Hedw.) Brid. cf. Crum & 

Steere, Sc. Surv. Portorico 7:435 (1957). 


subpulvinatus P. Varde & Thér., Bull. Soc. Bot. 

France 87:353 (1940). 

T: massif du Ruwenzori, Ituri 4000-4400 m, 

Humbert 7756 (PC). 

CU: C. smaragdinus cf. Frahm, Bryoph. Bibl. 

31:177 (1985). 

subpusillus Par., Ind. Bryol. Suppl. 98 (1900), 

OU: Microcampylopus nanus (C. Müll.) Par. 

cf. Dixon, Trans. R. Soc. S. Afr. 8:187 (1920). 

CU: Dicranella subsubulata cf. Magill. Fl. 

S. Afr. Bryoph. 1(1):121 (1981). 

subreconditus (Geh. & Hampe) Kindb., Enum. 

Bryin. Exot. 89 (1889). 

B: Dicranum subreconditum Geh. & Hampe, 

Flora 64: 343 (1881). 

CU: C. erythrodontius cf. Frahm, Revue 

Bryol. Lich. 45:145 (1979) = C. pyriformis

(Schultz) Brid. 

subsecundus Lindb. in Aongstr., Oefv. K. Vet. 

Ak. Foerh. 33:5 (1876) nom. nud. 

substramineus Broth. in Mildbr., Deutsch. Zentral 

Afr. Exp. 2:139 (1910). 

T: Africa occ, Karisimbi 4000 m, Mildbraed 

2088 (B). 

CU: C. stramineus (Mitt.) Jaeg. cf. P. Varde, 

Bull. Mus. Hist. Nat. Paris ser.2,14:363 

(1943) = C. nivalis (Brid.) Brid. 

substrictus Hampe in Fleisch., Musci Fl. 

Buitenzorg 1:87 (1904) nom. nud. 

CU: Dicranodontium nitidum (Dozy & 

Molk.) Fleisch. cf. Fleischer l.c. 

substygius C. Müll. in Kindb., Enum. M. Exot. 9 

(1888) nom. nud. 

subtenuifolius Hampe in C. Müll., Gen. Musc. 

Fr. 274 (1900) nom. nud. 

subtorquatus C. Müll., Rev. Bryol. 3:3 (1876) 

nom. nud. 

CU: C. woolsii (C. Müll.) Par. var. cylindrica 

(C. Müll.) Par. cf. Par., Ind. Bryol. ed.2,1:334 

(1904) = C. pyriformis. 

subtrachyblepharus C. Müll., Gen. Musc. Fr. 270 

(1900). (*) 

subtricolor Lor., Moosstud. 159 (1864). 

T: India Orientalis no. 6 (M). 

CU: C. aureus cf. Frahm, Journ. Bryol. 

13:165 (1984) = C. schmidii. 

subturfaceus Card., Rev. Bryol. 37:119 (1910). 

T: Mexico, Chiapas, San Cristobal, Münch s. 

n. (holotype PC). 

CU: C. fragilis cf. Frahm, Bryologist 80:120 

(1977). 

subulatus Schimp. in Rabenh., Bryoth. Eur. 

9:n.451 (1861). 

T: Italien: um Gratsch undAlgund bei Meran 

(Tirol), Milde s.n. (lectotype S). 

subulatus ssp. schimperi (Milde) Dix., Stud. 

Handb. Brit. Moss. 91 (1896). 

B: Campylopus schimperi Milde 

CU: C. schimperi Milde cf. Frahm & Vitt, 

Brittonia 30:365-373 (1978). 

subulifolius Thwait. & Mitt., J. Linn. Soc. Bot. 

13:297 (1873) 


Frahm, Hikobia 11 (1994) = C. thwaitesii cf. 

Frahm, Crypt. Bryol. Lichènol. 19: 32, 1998. 

subulifolius Schimp. in C. Müll., Gen. Musc. Fr. 

269 (1900) nom. nud. 

81 

subvirescens Ren. & Card., Bull. Soc. R. Bot. 

Belg. 33(2):109 (1895). 

T: Madagascar, Andevorante, Camboué s.n. 

(PC). 



Bryoph. Bibl. 31:16 (1985). 

sudabyanus Rehm. in Dix. & Ggepp., Misc. Inf. 

Kew 1923:197 (1923) nom. nud. 

CU: C. robillardei Ren. & Card. cf. Frahm, 

Bryoph. Bibl. 31:199 (1985). 

sudeticus (Funck) Fuernr., Flora 12:595 (1829). 

CU: Racomitrium heterostichum (Hedw.) 

Brid. ssp. sudeticum (Funck) Dix. 

sulfureo-flavus (C. Müll.) Par., Ind. Bryol. Suppl. 

98 (1900). 

CU: C. apressifolius Mitt. cf. Dixon, New 

Zealand Inst. Bull. 3(3):89 (1923) = C. 

clavatus (R.Brown) Wils. cf. Sainsb., Trans. 

New Zealand Inst. 59(3):506 (1928). 

sulfureo-nigritus Dus., Ark. Bot. 4(13):6 (1905). 


Beitr. 7: 78 (1987). 

sulfureus Besch., Bull. Soc. Bot. France 45:55 

(1898). 

T: Tahiti, Nadeuad 207 (isotype BM). 


7: 78 (1987). 

surinamensis C. Müll., Linnaea 21: 186 (1848). 

T: Suriname, near Paramaibo, Kegel 516 

(holotype destroyed at B; lectotype H-BR; 

isolectotypes GOET, PC) 

surinamensis var. angustiretis (Aust.) J.-P.Frahm, 

Bryologist 83:582 (1980). 

CU: C. angustiretis Aust. cf. Frahm, Nova 

Hedwigia 41:276 (1985). 

symonsii Sim Trans. R. Soc. S. Afr. 15:167 

(1926). 

T: S. Africa, Natal, Giant s Castle, top, 8000 

ft., Symons (1912) s.n., hb. Sim 9843 

(lectotypus L). 

CU: C. aureonitens cf. Frahm, Bryoph. Bibl. 

31: 24 (1985). 

OU: C. bicolor var. ericeticola cf. Frahm, 


symplectus Stirt., Scott. Natural. 11:234 (1886). 

CU: C. schwarzii Schimp. cf. Dixon, J. Bot. 

61:17 (1923).

82 

T 

tablasensis Thér., Rev. Bryol. Lich. 11: 152 

(1939). 

T: Bolivia, Rio Tablas, Herzog 4534a 

(holotype PC; isotype JE). 


J. Bryol. 11:215 (1980). 

tabularis Thér. in Broth., Nat. Pfl. ed. 2,10:1888 

(1924) nom. nud. 

taiwanensis Sak., Bot. Mag. Tokyo 55:206 

(1941). (*) 

tallulensis Sullivant & Lesquereux, Musci bor.amer. 

17 (1865). 

T: U.S.A. Georgia: Tallulah Falls, Small s.n. 

(NY). 


Fieldiana Bot. 45: 43 (1949). 

tallulensis var. subleucogaster (C. Müll.) Grout, 

Moss Fl. N. Am. 1:92 (1937). 



OU: C. flexuosus cf. Bartram l.c. 

tasmanicus Par., Ind. Bryol. Suppl. 98 (1900) 

CU: C. introflexus (Hedw.) Brid. cf. Dixon, 


taylorii Dix., J. Bot. 76:224 (1938). 

T: Kenia, in burnt thicket 10.500 ft., 

Kinangop, Aberdare Mtns. Taylor 1630 

(holotype BM). 


Bryoph. Bibl. 31:124 (1985). 

tenax (C. Müll.) Par., Ind. Bryol. Suppl. 98 

(1900). 

B: Dicranum tenax C. Müll. Hedwigia 38: 83 

(1899). 

T: Cape, ad Blanco, Rehmann 54 (PC,G). 

CU: C. stenopelma (C. Müll.) Par. cf. Sim, 


thwaitesii. 

tenellus Rehm. in Dix. & Gepp, Bull. Misc. Inf. 

Kew 1923:196 (1923) nom. nud. 


31:199 (1985). 

tener Broth., Rev. Bryol. 47:2 (1920). 

T: Ecuador prov. Azuay, Rio Gualaquiza, 

Allioni s. n. (lectotypus H-BR). 

CU: C. trivialis cf. Frahm, Nova Hedwigia 

39:163 (1984). 

tenuinervis Fleisch., Musci Fl. Buitenzorg 1:120 

(1920). 

T: Indonesia Java, Gedeh 2400 m, Fleischer 

107 (isotype PC). 

CU: C. involutus cf. Frahm, Nova Hedwigia 

39:593 (1984) = C. ericoides (Griff.) Jaeg. 

tenuis Wils., Kew J. Bot. 9:296 (1857) nom. nud. 

CU: Campylopodium khasianum (Griff.) Par. 

cf. Mitt., J. Linn. Soc. Bot. Suppl. 1:8 (1859). 

tenuissimus Sull., Proc. Am. Ac. Arts Sc. 5:279 

(1861). 

T: Cuba, Wright s.n. (isotype GOET). 

OU: C. porphyreodictyon (C. Müll.) Mitt. cf. 

Mitten, J. Linn. Soc. Bot. 12:75 (1869). 

CU: Campylopus filifolius (Hornsch.) Mitt. 

cf. Frahm, Bryoph. Bibl. 5:98 (1975) = 


tequendamensis Thér., Rev. Bryol. Lich. 11:58 

(1939). 

T: Colombia, Tequendama, Troll 2036 


CU: C. tallulensis cf. Frahm, Nova Hedwigia 

5:528 (1981). 

terebrifolius (C. Müll.) Jaeg., Ber. S. Gall. 

Naturw. Ges. 1877-78:385 (1880). (*) 

thwaitesii (Mitt.) Jaeg., Ber. S. Gall. Naturw. Ges. 

1870-71:417 (1872). 

B: Dicranum thwaitesii Mitt., J. Linn. Soc. 

Bot. Suppl. 1: 19 (1859). 

T: Ceylon, Thwaites s.n. (holotype NY). 

thysanomitrioides Thér., Rev. Bryol. Lich. 11:51 

(1939). 

T: Bolivia, Cerros de Malaga, Herzog 4400 


CU: C. harpophyllus cf. Frahm, J. Bryol. 

11:214 (1980) = C. edithae Broth. 

tijucae Broth. in Par., Ind. Bryol. Suppl. 98 (1900) 

nom. nud. 

M: Brazil, Rio de Janeiro, „ad rupes prope 

Tijucae“, Ule Bryotheca bras. 213 (GOET). 

CU: C. widgrenii (C. Müll.) Mitt. cf. Frahm, 


OU: C. erythrodontius cf. Frahm, Rev. Bryol. 

Lich. 45:145 (1979). 

tisserantii P. Varde, Rev. Bryol. Lich. 18: 108 

1949 

T: Oubangui, station de Boukoko, Tisserant 

B 28,124 (PC) 

CU: C. nanophyllus C. Müll. ex Broth., 

Bryoph. Bibl. 31:122 (1985). 

tolucensis Bartr., Rev. Bryol. Lich. 15:21 (1946).

T: Mexico, Nevado de Toluca, Bartram 4114 


OU: C. chrismarii (C. Müll.) Mitt. var. 

tolucensis (Bartr.) J.-P.Frahm, Bryologist 

80:120 (1977). 

CU: C. nivalis (Brid.) Brid. s.str. cf. Frahm, 

Fl. Neotrop. 54: 137 (1991). 

torfaceus B.S.G., Bryol. Eur. 1:164 (1847). 


torquatus Mitt. in Hook.f., Fl. Tasman. 2:173 

(1859). 

T: New Zealand, E-Coast and Auckland, 

Sinclair (1850) s.n. (lectotype BM). 

CU: C. pallidus Hook.f. & Wils = C. 


torrentis Thér. & P. Varde, Rev. Bryol. Lich. 5: 

202 (1933). 

T: Gabun, cascade du Mougoungoulou, Le 

Testu s.n. (holotype PC). 

tortilipilus J.-P.Frahm, Cryptogamie Bryol. Lich. 

2:446 (1981). 

T: Brazil, Piaui, Mc. de Corrente, Vital 8232 

(holotype SP). 


Frahm, Fl. Neotr. 54: 167 (1991). 

tortilisetus Ule in C. Müll., Bull. Herb. Boiss. 6:34 

(1898) nom. nud. 

OU: C. rufescens Broth. cf. Par. l.c. 

T: C. occultus Mitt. cf. Frahm, Revue Bryol. 

Lich. 45:156 (1979). 

tortorae Herz., Beih. Bot. Centralbl. 26(2):52 

(1910). 

CU: Pilopogon spec. cf. Frahm, Journ. Hattori 

Bot. Lab. 44:485 (1978). 

tortuosus (Hampe) Par., Ind. Bryol. Suppl. 98 

(1900). 

B: Dicranum tortuosum Hampe, Linnaea 25: 

361 (1853). 

T: Puerto Rico (not seen). 

CU: C. lamellinervis cf. Frahm, Bryoph. Bibl. 

5:124 (1975). 

trachyblepharon (C. Müll.) Mitt., J. Linn. Soc. 

Bot. 12: 80 (1869). 

B: Dicranum trachyblepharon C. Müll., Syn. 

Musc. 1: 389 (1848). 

T: Brazil, Beyrich s.n. (holotype destroyed at 

B; lectotype NY). 

trachyblepharon ssp. comatus (Ren. & Card.) 

Frahm, Bryophytorum Bibliotheca 31: 187 

(1985). 

83 

B: Campylopus comatus Ren. & Card. 

T: St. Marie de Madagascar, Rev. Arbogast 

s.n. (isotype G, holotype PC). 

trachynotus (C. Müll.) Par., Ind. Bryol. Suppl. 

98 (1900) . 

B: Dicranum trachynotum C. Müll., Bull. 

Herb. Boiss. 6: 33 (1898). 

HS: trachynotus C. Müll., Gen. Musc. Fr. 276 

(1900) nom. nud. 

CU: C. beyrichianus cf. Frahm, Revue Bryol. 

Lich. 45:140 (1979) = C. pyriformis (Schultz) 

Brid. cf. Frahm, Fl. Neotrop. 54: 145 (1991). 

trachythecius (C. Müll.) Jaeg., Ber. S. Gall. 

Naturw. Ges. 1870-71:423 (1872). 

B: Dicranum trachythecium (C. Müll., 

Linnaea 36: 33 (1869). 

T: Ceylon, Ounduloay, Nietner s.n. (H-BR; 

FH). 

CU: C. comosus cf. Frahm, J. Bryol. 13:166 

(1984). 

trailii R. Brown ter., Trans. New Zealand Inst. 

29:468 (1897). 

CU: C. clavatus (R,Brown) Wils. cf. Dixon, 


transvaaliense Herz. et Dix. in Sim, Trans. R. 

Soc. Afr. 15:166 (1926). 


Bryoph. Bibl. 31:71 (1985). 

transvaaliensis Rehm. in Par. Ind. Bryol. 254 

(1894). 

T: Transvaal, in monte Kwatlamba, MacLea 

s.n. (L). 

HS: C. transvaaliensis Rehm. in Kindb., 

Enum. Bryin. Exot.89 (1889) nom. nud. 


31:145 (1985). 

OU: C. stenopelma (C. Müll.) Par. cf. Sim, 


trichodes Lor., Moosstud. 159 (1864). 

T: Cape, Table Mountain, Ecklon s.n. (H-BR). 

CU: C. pilifer Brid. cf. Frahm, Revue Bryol. 

Lich. 42:607 (1976). 

trichophorus Hampe ex Herz., Biblioth. Bot. 

87:20 (1916). 

T: Ecuador Prov. Loja, Krause s.n. (not seen). 

CU: C. asperifolius cf. Frahm, Lindbergia 

7:29 (1981). 

trichophylloides Thér. in Herzog, Rep. Spec. 

Novi Regni Veget. 95 (1938). 

T: Peru, Cordillera Blanca, Kinzl s. n.

84 

(holotype JE; isotype, B). 

tricolor (C. Müll.) Jaeg., Ber. S. Gall. Naturw. 

Ges. 1870-71:437 (1872). 

B: Dicranum tricolor C. Müll., Bot. Zeit. 11: 

36 (1853). 

CU: C. pilifer Brid. cf. Frahm, J. Bryol. 

13:179 (1984). 

trivialis C. Müll. ex Britton, Bull. Torrey Bot. 

Club 23: 476 (1896). 

T: Bolivia, Mapiri, Rusby s. n. (holotype, NY). 

trollii Thér., Rev. Bryol. Lich. 11:46 (1939). 

T: Bolivia, Yungas von La Paz, Troll 111 

(holotype PC; isotype, B). 

CU: C. pauper (Hampe) Mitt. var. 

lamprodictyon (Hampe) J.-P. Frahm cf. 

Frahm, Fl. Neotr. 54: 150 (1987). 

truncatus C. Müll., Linnaea 18:685 (1845). 


J. Linn. Soc. 12:84 (1869). 

tsaratananae Thér., Rec. Publ. Soc. havraise 

Etud. Div. 1925:128 (1926). 

T: Madagascar, Mt. Tsaratanana alt. 1200- 

2400 m, Perrier de la Bathie s.n. (holotype 

PC, isotype G). CU: C. flexuosus (Hedw.) 

Brid. cf. Frahm, Bryoph. Bibl. 31:76 (1985). 

tubulosus Bartr., Bishop Mis. Occ. Pap. 16:324 

(1942). 

T: Hawaii, Kauai, summit of Waialeale, 

Skottsberg et al. 4033 (holotype FH). 

OU: C. skottsbergii cf. Frahm, Revue Bryol. 

Lich. 44:51 (1978). 

CU: C. schmidii (C. Müll.) Jaeg. ssp. 

hemitrichius (C. Müll.) J.-P. Frahm, The 

Bryologist 94: 64, 1991. 

tuerckheimii (C. Müll.) Par., Ind. Bryol. Suppl. 

98 (1900). 

B: Dicranella tuerckheimii C. Müll., Bull. 

Herb. Boissier 5: 186 (1897). 

T: Guatemala, Alta Vera Paz: Coban, 

Tuerckheim 6652 (NY BM). 


Williams cf. Williams, N. Am. Fl. 15:94 

(1913), Microcampylopus curvisetus 

(Hampe) Giese & J.-P.Frahm cf. Giese & 

Frahm, Lindbergia ..............., 

CU: Campylopus zygodonticarpus (C. Müll.) 

Par. cf. Allen, Trop. Bryol. 1: 92 (1989). 

tullgrenii Ren. & Card., Bull. Herb. Boiss. ser. 

2,2:435 (1902). 

T: Teneriffe, Guarranato, Tullgren 101 (PC). 


31:145 (1985). 

tunariensis Herz., Bibl. Bot. 87:19 (1916). 

T: Bolivia, Tunarisee, Herzog 3430 (lectotype 

JE; isolectotypes B, H-BR). 

CU: C. fragilis (Brid.) B.S.G. cf. Frahm, Nova 

Hedwigia 39:158 (1984). 

turficola Broth. in Skottsb., Nat. Hist. Juan 



Hedwigia 34:392 (1981). 


Beitr. 7: 81 (1987). 

turgidus Rehm. in Dix. & Gepp, Bull. Misc. Inf. 

Kew 1923:197 (1923) nom. nud. 

M: Orange Free State, Kadziberg, Rehmann 

57 (PC). 


31:199 (1985). 

U 

uii Broth., Oefv. Finsk. Vet. Soc. Foerh. 62A: 3 

(1921). 

T: Japan, Kii, Ui 1103 (holtype H-BR). 

CU: C. japonicus Broth. cf. Takaki, J. 

Hatt. Bot. Lab. 30:240 (1967). 

uleanus (C. Müll.) Broth., Nat. Pfl. 1(3): 333 

(1901). 

B: Dicranum uleanum C. Müll., Hedwigia 

39: 258 (1900). 

T: Brazil, S. Catarina, Flaggenberg, Ule 

148 (holotype destroyed at B; lectotypus nov. H- 

BR). 

umbellatus (Arn.) Par., Ind. Bryol.: 264 (1894). 

B: Thysanomitrium umbellatum Arnott, 

Nouv. Dispos. Meth. Especes Mousses 34 

(1825). 

T: USA, „Ins. Sandwich“ (Hawaii) 

Gaudichaud s.n. (holotype PC). 

OU: C. richardii cf. Dixon, J. Bot. 60:287 

(1922). 

umbellatus (Arn.) Par., Ind. Bryol.: 264 (1894) 

var. pterotoneuron (C. Müll.) J.-P. Frahm, 

Nova Hedwigia 39: 602, 1984. 

B: Campylopus pterotoneuron (C. Müll.) 

Jaeg., Ber. S. Gall. Naturw, Ges. 1870- 

71:417 (1872), Dicranum pterotoneuron 

C. Müll, Linnaea 36: 34, 1869. 

uncinatus (Harv.) Dozy & Molk., Bryol. Jav. 1:78 

(1858).

CU: Dicranodontium uncinatum (Harv.) 

Jaeg. 

underwoodii Williams N. Amer. Fl. 15(2): 142 

(1913). 

T: Jamaica, Summit of Blue Mt. Peak, 

Underwood 2539 (holotype NY, isotypes 

F, H-BR). 

OU: C. brittonae (non setaceus cf. Frahm, 

Cryptogamie 3(1):62, 1982) cf. Frahm, 


CU: C. shawii cf. Frahm, J. Bryol. 13:329 

(1985). 

uruguensis Frahm, Bryoph. Bibl. 5:127 (1975) 

nom. nov. pro C. nitidus Thér. in Felipp. 

hom. illeg. 

T: Uruguay, Maldonado, Felippone 1214 

(B, H-BR). 

CU: C. pyriformis (Schultz) Brid. cf. 


usambaricus P. Varde, Ark. Bot. ser.2,3:135 

(1955). 

T: Tanganyika, Usambara Mt. near Amani 

950 m, Hedberg 2037h (holotype PC). 

CU: C. obrutus cf. Frahm, Bryol. Beitr. 

7:82 (1987). 

V 

valentinii Besch. Ann. Sci. Nat. Bot. ser. 6,9:325 

(1880). 

T: Bourbon, Valentin s.nr. (holotype PC). 



valerioi Allen, Trop. Bryol. 1:68 (1989) nom. 

nov. pro. Dicranum costaricense Bartr., J. 

Wasj Acad. Sci 19:13 (1929) hom. illeg. pro 

Campylopus costaricensis Bartr., Contr. U.S. 

Nat. Herb. 26: 62 (1928). 

T: Costa Rica, la Palma 1500m, Valerio 148 

(holotype FH, isotype US). 

validinervis (Dix.) B.C. Tan, Willdenowia 18: 

510, 1989. 

vallis-gratiae Hampe in Breutel, Musci Capenses, 

nom. invalid. 

OU: C. leucobasis (C. Müll.) Par., var. 

longescens (C. Müll.) Par. cf. C. Müll., 

Hedwigia 38: 19, 1899; C. echinatus Sim cf. 

Sim, Trans. R. Soc. S. Afr. 15: 175, 1926. 

CU: C. introflexus (Hedw.) Brid. 

vaporarius Bolle, Verh. Bot. Ver. Brandenburg 

85 

7:20 (1865) nom. nud. 

OU: C. introflexus (Hedw.) Brid. var. 

vaporarius (De Not.) Mönk. 


variegatus Dix. & Wagn. in Sim, Trans. R. Soc. 

S. Afr. 15:174 (1926) nom. invalid. 

T: Transvaal, Kaapsche Hoek, Wager 209 

(isotypes PC, L). 

OU: C. inerangae (C. Müll.) Par. cf. Sim l.c., 

C. inchangae (C. Müll.) Par. cf. Magill, Fl. 

S. Afr. Bryoph. 1(1):146 (1981). 

CU: C. robillardei Besch. cf. Frahm. Bryol. 

Beitr. 7: 82, 1987. 

ventanus C. Müll., Gen. Musc. Fr. 265 (1900) 

nom. nud. 

M: Argentinien, Felsen in einem Thale der 

Sierra Ventana, Lorentz (1881) (JE). 

ventri-alaris (C. Müll.) Broth., Nat. Pfl. 1(3):334 

(1901). 

B: Dicranum ventri-alare C. Müll., Hedwigia 

39: 251 (1900). 

T: Brazil, Minas Gerais, Ule 1347 (holotype 

destroyed at B; lectotype, H-BR). 



ventrialaris Herz. in Broth., Nat. Pfl. ed.2,10:188 

(1924) nom. nud. 

verdoliniii Par. & Ren. („verdillonii“), Rev. 

Bryol. 29:3 (1902). 

T: Madagascar, vallée de l’Ofika près 

Anaribe, Verdillon 12 (holotype PC). 


Bryoph. Bibl. 31:76 (1985). 

vernieri Duby, Flora 58: 283 (1875). 

T: Tahiti, Vernier s.n. (PC). 

OU: C. obscurus Aongstr. cf. Besch., Ann. 

Sc. Nat. ser. 7,20:17 (1895). 



verrucosus Besch., Ann. Sc. Nat. Bot. ser. 

5,18:200 (1873). 

T: Nouvelle- Caledonie, Mt. Mie 1000 m, 

Balansa 914 (isotypes BM, L). 


Bryol.Beitr. 7: 91 (1987). 

verticillatus (Hampe) Kindb., Enum. Bryin. Exot. 

89 (1889). 

B: Dicranum verticillatum Hampe, Vid. 

Medd. Naturh. For. Kjoebenh. ser. 3, 4: 44 

(1872).

86 




P.Frahm. 

vesticaulis Mitt. in Mell. S. Helena 359 (1875). 

T: Tristan d’Acunha, Milne s.n. (holotype 

NY, isotype B). 

villicaulis (Hampe) Jaeg., Ber. S. Gall. Naturw. 

Ges. 1877-78:381 (1880). 

B: Dicranum villicaule Hampe, Vidensk. 

Medd. Naturh. For. Kjoebenh. er. 3,4: 44. 

1872. 


(isotype, H-BR). 


Bryol. Lich. 45:165 (1979). 

virescens Besch., Ann. Sc. Nat. Bot. ser. 6,9:325 

(1880). 

T: La Reunion, Valentin s.n. (holotype PC). 


Bryoph. Bibl. 31:76 (1985). 

virginicus (Aust.) Lesq. & Jam., Man. Moss. N. 

Am. 80 (1884). 

CU: Dicranodontium asperulum (Mitt.) 

Broth. cf. Williams, N. Am. Fl. 15:152 (1913). 

viridatulus C.Müll. ex Broth. Bot. Jahrb. 24:234 

(1897). 

T: „Guinea gall. Kouroussa prope pagum 

Diendien“, Pobeguin s.n. (isotype PC, 

lectotypus nov. L). 



viridatus (C. Müll.) Broth., Nat. Pfl. 1(3): 333 

(1901). 

B: Dicranum viridatum C. Müll., Hedwigia 

39: 257 (1900). 

T: Brazil, Sao Francisco, Rio Passucao, Ule 

31 (holotype destroyed at B; lectotypus nov. 

H-BR). 


Frahm, Hikobia 11 (1994) = C. thwaitesii. 

viridicatus (C. Müll.) Par., Ind. Bryol. Suppl. 99 

(1900). 

B: Dicranum viridicatum C. Müll., Hedwigia 

36: 352, 1897. 

T: Australia, Waterloo Marsches, Whitelegge 

3369; Northwood, Whitelegge 21; Sidney, 

Whitelegge s.n. (syntypes ). 

CU: C. pyriformis (Schultz) Brid. cf. 

Catcheside in litt. 

viridis Sull. & Lesq., Musci Bor. Am. 18 (1856). 

CU: Dicranum viride. 

viridulus Card., Bull. Herb. Boiss. ser. 2,7:715 

(1907). 

T: Japan, „Arita, rochers, n. 2439“ (holotype 

PC). 

CU: =?= C. umbellatus (Arn.) Par. cf. Frahm, 

Revue Bryol. Lich. 42:907 (1976), Takaki, J. 

Hatt. Bot. Lab. 30:236 (1967) sub C. 

richardii. 

vitzliputzli Lor., Moosstud. 158 (1864). 


N. Am. Fl. 15:144 (1913). 


voeltzkowii Broth. in Voeltzkow, Reise Ostafr. 

3:52 (1908) 

T: Madagascar, St. Marie, Voeltzkow s.n. 


OU: C. cailleae Ren. & Card. cf. Broth., Nat. 

Pfl. ed.2,1o:187 (1924). 


Bibl. 31:139 (1985). 

W 

wabagensis Bartr., Rev. Bryol. Lich. 30: 187 

(1962). 

T: Papua N.G., Western Highlands, along 

track to Marimuni...10.000 ft. Robbins 3025 


CU: C. comosus (Schwaegr.) Bosch & Lac. 

cf. Frahm et al., Acta Not. Fennica 131:76 

(1985). 

walkeri (Mitt.) Jaeg., Ber. S. Gall. Naturw. Ges. 

1870-71:429 (1872). 

B: Dicranum walkeri Mitt., J. Linn. Soc. Bot. 

Suppl. 1: 18, 1859. 

T: Ceylon, Walker, Thwaites 21 (isosyntypes 

H-BR). 


P.Frahm et al., Acta Bot. Fennica 131:68 

(1985). 



walkeri R.Brown ter., Trans. New Zealand Inst. 

29:469 (1897) hom. illeg. 

OU: C. appressifolius Mitt. cf. Dixon. New 




wallisii Sauerb. in Jaeg., Ber. S. Gall. Naturw.

Ges. 1877-78:382 (1880). 

OU: Dicranum hemitrichius (C. Müll.) Jaeg. 

cf. Bartr., Philipp. J. Sc. 68:41 (1939) =?= C. 

hemitrichius (C. Müll.) Jaeg. = C. schmidii 

(C. Müll.) Jaeg. ssp. hemitrichius (C. Müll.) 

J.-P. Frahm, The Bryologist 94: 64, 1991. 

wattsii Broth., Proc. Linn. Soc. N.S.Wales 43:547 

(1918). 

T: Australia, Queensland, Watts 366 



Bryol. Beitr. 7: 84 (1987). 

wawraeanus Reichdt., Sitzungsber. Ak. Wiss. 

Wien 75:565 (1877). 

OU: C. densifolius Aongstr. cf. Bartram, 

Bishop Mus. Bull. 101:38 (1933). 




weberbaueri Broth., Bot. Jahrb. 56 Beibl. 123:5 

(1920). 

T: Type. Peru, Loreto, E of Moyobamba, 

Weberbauer 4718 (holotype, H-BR). 

CU: C. huallagensis Broth. var. weberbaueri 

(Broth.) J.-P. Frahm, Fl. Neotr. 54: 114 

(1991). 

weddelii Besch., J. de Bot. 5:147 (1891). 

T: Peru, Carabaya: Weddel s s.n. (isotype 

BM). 

CU: C. asperifolius Mitt. cf. Frahm,. Bryol. 

Beitr. 7: 84, 1987. 

weisiopsis (C. Müll.) Par., Ind. Bryol. Suppl. 99 

(1900). 

B: Dicranum w. C. Müll., Hedwigia 38: 78 

(1899). 

T: Cape Town, Rehmann 61 (lectotype PC 

). 

OU: C. lepidophyllus (C. Müll.) Jaeg. cf. 

Dixon in Sim, Trans. R. Soc. S. Afr. 15:170 

(1926), C. introflexus (Hedw.) Brid. cf. 

Magill, Fl. S. Afr. Bryoph. 1(1):150 (1981). 


31:143 (1985). 

wheeleri (C. Müll.) Hampe ex Par., Ind. Bryol. 

Suppl. 99 (1900). 

B: Dicranum wheeleri Hampe ex C. Müll., 

Flora 82: 441 (1896) 

T: Hawaii, without locality, Wheeler s.n., 

Boswell s.n. (syntypes, not available). 

OU: Dicranoloma wheeleri (C. Müll.) Par. 

87 

Ind. Bryol. ed. 2,31 (1904). 

whiteleggei C. Müll. in Geh., Rev. Bryol. 24:67 

(1897) nom. nud. 

CU: Dicranoloma argutum (Hampe) Par. cf. 

Dixon, New Zealand Inst. Bull. 3(1):14 

(1913). 

wichurae Broth. in Fleisch., Musci Fl. Buitenzorg 

1:110 (1904) nom. nud. 

CU: C. recurvus (Mitt.) Jaeg. cf. Fleischer l.c. 

widgrenii (C. Müll.) Mitt., J. Linn. Soc. Bot. 12: 

88 (1869). 

B: Dicranum widgrenii C. Müll., Bot. Zeit. 

14: 418 (1856). 

T: Brazil, Minas Gerais, Caldas, Widgren s. 

no. (holotype destroyed at B; lectotype NY; 

isotypes H-BR, H-SOL). 

wilhelmensis Bartr. Contr. U.S. Nat. Herb. 37 

(1965). 

T: Papua New Guinea,. Simbu, east slopes of 

Mt. Wilhelm, 2740 m. Brass 30237 (holotype 

FH). 

CU: C. comosus cf. Frahm et al., Acta Bot. 

Fenn. 131:76 (1985). 

woolsii (C. Müll.) Par., Ind. Bryol. Suppl. 99 

(1900). 

B: Dicranum woolsii C. Müll., Hedwigia 36: 

348 (1897). 

T: Australia, Queensland, Brisbane, Bailey 5 

(holotype B). 

HS: Campylopus woolsianus Broth., Nat. Pfl. 

1(3): 332 (1901). 


Bryol. Beitr. 7: 94(1987).. 

wurdackii Robins. The Bryologist 70:319 (1967). 

T: Peru, Amazonas: Rio Ventilla, Wurdack 

1524 (holotype US). 

CU: C. weberbaueri cf. Frahm, J. Hattori Bot. 

Lab. 44:485 (1978) = C. huallagensis Broth. 

var. weberbaueri (Broth.) J.-P. Frahm, Fl. 

Neotr. 54: 114 (1991). 

X 

xanthophyllus Mont., Ann. Sc. Nat. Bot. ser. 

3,4:111 (1845) 

T: Chili, Vooyage Gay, hb. Montagne 



J. Linn. Soc. Bot. 12:84 (1869). 

CU: C. clavatus cf. Thér., Rev. Bryol. Lich. 

7:172 (1935), Frahm, Nova Hedwigia 34:394

88 

(1981). 

Y 

yakushimensis Sak., Bot. Mag. Tokyo 47:332 

(1933) 

CU: C. atrovirens De Not. cf. Frahm, J. 

Hattori Bot. Laboratory 71: 146, 1992. 

yungarum Herzog, Beih. Bot. Centralbl. 26(2): 

51. 1910. 

T: Bolivia, Incacorral 2200 m, Herzog 1908 

(holotype JE, isotype B). 

CU: C. densicoma var. yungarum (Herz.) J.- 

P. Frahm, Nova Hedwigia 39:156 (1984). 

yunqueanus (C. Müll.) Par., Ind. Bryol. Suppl. 

99. 1900. 

B: Thysanomitrion yunqueanum C. Müll., 

Hedwigia 37: 225. 1898. 

T: Puerto Rico, Sierra Luquillo, Sintenius F 

29 (holotype destroyed at B; lectotype H-BR) 

CU: C. richardii Brid. cf. Williams, N. Am. 

Fl. 15:149 (1913). 

Z 

zollingerianus (C. Müll.) Bosch & Lac., Bryol. 

Jav. 1:77 (1858). 

B. Dicranum zollinerianum C. Müll., Syn. 

Musc. 2: 599 (1851). 

HS: C. zollingeri C. Müll. ex Kindb., Enum. 

Bryin. Exot. 51 (1888) 

T: Insula Zumbawa, ad rupes altitud,. 4000´, 

Zollinger 1184 (holotype L). 


zollingerianus (C. Müll.) J.-P. Frahm, Trop. 

Bryol. 4: 61, 1991. 

zonatus Mol., Ber. Naturh. Ver. Passau 10:53 

(1875) 

CU: C. flexuosus (Hedw.) Brid. 

zygodonticarpus (C. Müll.) Par., Ind. Bryol. 265 

(1894). 

B: Dicranum zygodonticarpum C. Müll., 

Linnaea 42: 471 (1879). 

T: Venezuela, Tovar, Fendler 35 (holotype 

destroyed at B; lectotype NY). 

OU: C. tallulensis Sull. & Lesq. var. 

subleucogaster (C. Müll.) Grout, Moss Fl. N. 

Am. 1:92 (1937), C. flexuosus (Hedw.) Brid. 

cf. Bartram, Fieldiana Bot. 25:43 (1949) 

APPENDIX: LIST OF LEGITIMATE 

SPECIES WITH SYNONYMS 

Unrevised taxa, of which the type material is not 

available, in brackets. 

(abyssinicus) De Not. 

acuminatus Mitt. 

birgeri Card. 

crassissimus Besch. 

curvatifolius Card. 

fuegianus Dus. 

kirkii Mitt. in Beck var. pilosus J.-P. Frahm 

sulfureo-nigritus Dus. 

var. kirkii (Mitt.) J.-P. Frahm 

denticuspis Broth. 

nigroflavus (C. Müll.) Par. 

aemulans (Hampe) Jaeg. 

auribrunneus (C. Müll.) Broth. 

brunneo-bolax (C. Müll.) Broth. 

filicaudatus (C. Müll.) Broth. 

setaceo-rigidus (Hampe) Jaeg. 

stricticaulis (C. Müll.) Par. 

subincrassatus (Hampe) Jaeg. 

albidovirens Herz. 

angusti-alatus Thér. 

guatemalensis Bartr. 

(ambiguus) Thér. 

amboroensis Thér. 

anderssonii (C. Müll.) Jaeg. 

fulvus Herz. hom. illeg. 

insularis Bartr. 

longisubulatus Thér. 

subincacorralis Thér. 

(andreanus) Card. & P. Varde 

angustiretis (Aust.) Lesq. & Jam. 

delicatulus Williams 

ditrichoides Broth. 

leptodictyon Broth. 

arctocarpus (Hornsch.) Mitt. 

buchtienii Thér. 

discriminatus (Hampe) Jaeg. 

crispicoma (C. Müll.) Jaeg. 

glaziovii (Hampe) Jaeg. 

microtheca Herz. 

pelichucensis Williams 

rectisetus (Hampe) Jaeg. 

rigidiusculus (Hampe) Jaeg. 

roraimae Broth.

sellowianus (Hampe) Jaeg. 

strictifolius Broth. 

ventri-alaris (C. Müll.) Broth. 

ssp. caldensis (Aongstr.) J.-P. Frahm 

catharinensis Par. 

crispatus (C. Müll.) Broth. 

falcatulus Bartr. 

perfalcatus Broth. 

perporosus Broth. 

scopelliformis (C. Müll.) Broth. 

subcubitus Williams 

tablasensis Thér. 

ssp. madecassus (Besch.) J.-P. Frahm 

madagassus Dix. 

matarensis Besch. 

rigens Ren. & Card. 

subvirescens Ren. & Card. 

areodictyon (C. Müll.) Mitt. 

breweri Bartr. 

nitidus Wils. in Mitt. 

subconcolor (Hampe) Mitt. 

arcuatus (Brid.) Jaeg. 

capitiflorus Mont. 

helenicus (C. Müll.) Jaeg. 

longifolius Schimp. ex Besch. 

parallelus Thér. in Broth. 

pseudovirescens Ren. & Card. 

santae-helenae Schimp. in Par. 

argyrocaulon (C. Müll.) Broth. 

leucognodes (C. Müll.) Par. 

areodictyon Schimp. in Mandon 

asperifolius Mitt. 

trichophorus Hampe ex Herz. 

weddelii Besch. 

atlanticus B.H. Allen 

atro-virens De Not. 

adustus De Not. 

falcatus Ferg. in Par. 

fergussonii Stirt. 

fusco-viridis (Card.) Dix. & Thér. 

perplexans Stirt. 

prasino-rufus Stirt. 

yakushimensis Sak. 

aureonitens (C. Müll.) Jaeg. 

aethiops Welw. & Duby 

aongstroemii C. Müll. in Aongstr. 

drouhardii Card. & Coop. 

hygrometricus Rehm. in Geh. 

lonchocladus C. Müll. in Geh. 

naumannii (C. Müll.) Kindb. 

purpureo-aureus (C. Müll.) Par. 

symonsii Sim 

valentinii Besch. 

ssp. recurvifolius (Dus.) J.-P. Frahm 

australis Catcheside & J.-P. Frahm 

angustilimbatus Bartr. hom. illeg. 

(austro-alpinus) (C. Müll.) Kindb. 

(austro-stramineus) Thér. 

austro-subulatus Broth. & Geh. 

novae-guineae Dix. & Thér. 

sericeus Fleisch. 

subericoides Williams 

bartramiaceus (C. Müll.) Par. 

bartramiaceus (C. Müll.) Dix. & Gepp 

(beauverdianus) Thér. 

(belangeri) Thér. 

bicolor (C. Müll.) Wils. in Hook.f. 

ssp. bicolor 

bartramiaceus (C. Müll.) Par. 

basalticolus (C. Müll.) Par. 

caducipilus Besch. in Par. 

stewartii R. Brown ter. 

subbicolor Geh. & Broth. 

var. ericeticola (C. Müll.) Dix. 

davallianus Watts. in Dix. 

ssp. atroluteus (C. Müll.) J.-P. Frahm 

brevis Rehm. in Geh. 

chlorophyllosus (C. Müll.) Jaeg. 

schenkei C. Müll. in Par. 

schunkei Rehm. in Dix. & Gepp 

brevipilus B.S.G. 

attenuatus (Stirt.) Dix. 

aurescens Stirt. 

decipiens Lac. 

fulvoviridis Stirt. 

molkenboeri Lac. 

(brunneus) (C. Müll.) Par. 

bryotropii J.-P. Frahm 

cambouei Ren. & Card. 

amblyacron (Ren.) Ren. & Card. 

labeensis Broth. & Par. 

lavardei Thér. in P. Varde 

laxobasis Ren. & Card. 

mozambicus P. Varde 

capitulatus Bartr. 

carolinae Grout 

89

90 

cerradensis Vital 

catarractilis (C. Müll.) Par. 

cataractitis Par. 

pulvinatus (C. Müll.) Rehm. in Par. 

semidorsum Rehm. ex Par. 

cavifolius Mitt. 

apollinairei Thér. 

(chalarobasens) C. Müll. in Par. 

chevalieri Broth. & Par. 

chilensis De Not. 

flavo-nigritus Dus. 

subappressifolius Broth. & Geh. in Rodw. 

circinatus J.-P. Frahm 

clavatus (R. Brown) Wils. in Hook.f. 

appressifolius Mitt. in Hook.f. 

arcuatus R. Brown ter. 

arenarius R. Brown ter. 

brownii Par. 

corralensis Broth. in Par. 

cylindrothecium R. Brown ter. 

elliptothecium R. Brown ter. 

insistitius Hook.f. & Wils. 

insidiosus Hook.f. & Wils. 

leptocarpus Wils. in Hook.f. 

leptodus Mont. 

montagnei De Not. 

otaramaii R. Brown ter. 

persimplex C. Müll. 

pycnangius C. Müll. in Kindb. 

rarus R. Brown ter. 

robertii Par. 

rodwayi Broth. 

scabridii Bealey in Dix. 

strictissimus C. Müll. 

subclavatus Kindb. 

sulfureo-flavus (C. Müll.) Par. 

trailii R. Brown ter. 

walkeri R. Brown ter. 

xanthophyllus Mont. 

clemensiae Bartr. 

cleefii J.-P. Frahm 

(cockyanii) R. Brown ter. 

comosus (Reinw. & Hornsch.) Bosch & Lac. 

caudatus (C. Müll.) Mont. in Dozy & Molk. 

flexifolius Bosch. & Lac. 

reduncus (Reinw. & Hornsch.) Bosch. & Lac. 

trachythecius (C. Müll.) Jaeg. 

wabagensis Bartr. 

walkeri (Mitt.) Jaeg. 

wilhelmensis Bartr. 

concolor (Hook.) Brid. 

crateris Besch. 

orzeszkoanus Ren. & Card. 

(crispatulus) Thér. 

cryptopodioides Broth. 

ramuliger Broth. 

(cruegeri) (C. Müll.) Par. 

cubensis Sull. 

harrisii (C. Müll.) Par. 

cuspidatus (Hornsch.) Mitt. 

bolivarensis Bartr. 

carassensis (Broth.) Par. 

hoffmannii (C. Müll.) Ren. & Card. 

nematophyllus Ren. & Card. 

var. dicnemioides (C. Müll.) J.-P. Frahm 

cucullatifolius Herz. 

cygneus (Hedw.) Brid. 

cacuminis (C. Müll.) Par. 

ekmannii Thér. 

elliottii Bartr. 

jamaicensis Mitt. 

saxatilis Williams 

setaceus Card. 

decaryi Thér. 

densicoma (C. Müll.) Par. 

cavernosus J.-P. Frahm 

filicuspis Broth. in Herz. 

gertrudis Herz. 

spurio-concolor (C. Müll.) Par. 

var. yungarum (Herz.) J.-P. Frahm 

dichrostis (C. Müll.) Par. 

(dieckianus) Stirt. 

dicranoides Thér. & Nav. 

lindavii Broth. in Engl. 

(dietrichiae) (C. Müll.) Jaeg. 

edithae Broth. 

andicola Thér. 

harpophyllus Herz. 

thysanomitrioides Thér. 

ericoides (Griff.) Jaeg. 

erythrognaphalus (C.Müll.) Jaeg. 

involutus (C.Müll.) Jaeg. 

roinei Card. & P. Varde 

tenuinervis Fleisch. 

exasperatus (Nees & Blum) Brid.

abbreviatus (Dix.) Manuel 

archboldii (Bartr.) Bartr. 

sclerodictyum Card. 

var. lorentzii (Fleisch.) J.-P. Frahm 

papuensis Bartr. 

ridleyi (Dix.) Manuel 

(exfimbriatus) C. Müll. 

extinctus J.-P. Frahm 

flaccidus Ren. & Card. 

ampliretis (C. Müll.) Par. 

chateauvertii Thér. 

consociatus Thér. 

edwardsii Sim 

heteroneuron Thér. 

latifolius Rehm. in Kindb. 

marillacii Rehm. in Par. 

olivaceo-nigricans (C. Müll.) Par. 

perrieri Thér. 

pseudonigrescens Thér. 

transvaaliense Herz. & Dix. 

var. cucullatifolius J.-P. Frahm 

flagelliferus (C. Müll.) Jaeg. 

flexuosus (Hedw.) Brid. 

alleizettii Thér. & P. Varde 

arduennae Lib. 

boivinianus Besch. 

brachymastyx C. Müll. ex Besch. 

brachyphyllus Wils. ex Mitt. 

brevifolius Roell in Limpr. 

crenulatus Stirt. 

heterophyllus Mitt. 

homomallus Leroy & Thér. 

hondurensis Bartr. 

leucophaeus Stirt. 

melaphanus Stirt. 

mexicanus Thér. 

micans Wulfsb. 

obtectus Stirt. 

paradoxus Wils. in Harsy 

quintasii Broth. 

roellii Ren. & Card. 

rubiginosus Stirt. 

ryparochlorus C. Müll. 

sargii Roell ex Ren. & Card. 

scabricuspis Mitt. in Mell. 

straminifolius Bartr. 

subcinereus Stirt. 

tsaratananae Thér. 

verdolinii Par. & Ren. 

virescens Besch. 

wattsii Broth. 

zonatus Mol. 

var. incacorralis (Herz.) J.-P. Frahm 

reflexus Broth. in Herz. 

subperichaetialis Biz. & Kilb. 

flindersii Catcheside & J.-P. Frahm 

fragilis (Brid.) B.S.G. 

ssp. fragilis 

abyssinicus Schimp. in Par. 

akagiensis Broth. & Yas. 

alpicola (C. Müll.) Jaeg. 

citrescens Stirt. 

densus B.S.G. 

fimbriatus Mitt. 

laete-virens (C. Müll.) Jaeg. 

penicillatus Brid. 

pinfaensis Thér. 

retinervis (C. Müll.) Par. 

ruwenzoriensis Demar. & P. Varde 

subflexuosus Par. 

subturfaceus Card. 

tunariensis Herz. 

ssp. fragiliformis (J.-P. Frahm) J.-P. Frahm 

luetzelburgii Herz. ex J.-P. Frahm 

? sphagnicola Herz. 

ssp. goughii (Mitt.) J.-P. Frahm 

goughii (Mitt.) Jaeg. 

albescens (C. Müll.) Jaeg. 

ceylonensis Broth. in Fleisch. 

compactus Wils. 

latinervis (Mitt.) Jaeg. 

nietneri (C. Müll.) Jaeg. 

nilghiriensis (Mitt.) Jaeg. 

pseudogracilis Card. & Dix. 

subgracilis Ren. & Card. ex Gang. 

ssp. zollingerianus (C. Müll.) J.-P. Frahm 

celebesicus Card. 

zollingerianus (C. Müll.) Bosch. & Lac. 

boswellii (C. Müll.) Par. 

crispifolius Bartr. 

celebesicus Card. 

fusco-lutescens Ren. & Card. 

gardneri (C. Müll.) Mitt. 

gastro-alaris (C. Müll.) Par. 

gemmatus (C. Müll.) Par. 

arachnoideus Herz. 

91

92 

(geraense) Par. 

gracilis (Mitt.) Jaeg. 

schwarzii Schimp. 

auriculatus Wils. in MacKinlay 

huntii Stirt. 

purpurascens Stirt. 

symplectus Stirt. 

griseus (Hornsch.) Jaeg. 

lapidicola (C. Müll.) Par. 

recurvipilus (C. Müll.) Par. 

scabrophyllus (C. Müll.) Par. 

subgriseus (Hampe) Jaeg. 

ssp. ingeniensis (Williams) J.-P. Frahm 

hawaiicus (C.Müll.) Jaeg. 

var. hawaiicus 

purpureo-flavescens (C. Müll.) Par. 

rubricaulis Broth. & Par. 

var. hawaiico-flexuosus (C. Müll.) J.-P. Frahm 

perangustifolius C. Müll. 

var. densifolius (Aongstr.) J.-P. Frahm 

densiretis Broth. 

plumosus Sull. ex Par. 

wawreanus Reichdt. 

hensii Ren. & Card. 

levieri Broth. in Par. 

suberythrocaulon Broth. in Mildbr. 

heterostachys (Hampe) Jaeg. 

alopecurus (C. Müll.) Kindb. 

annotinus Mitt. 

hellerianus (Hampe) Jaeg. 

porphyreocaulis (C. Müll.) Kindb. 

hildebrandtii (C. Müll.) Kindb. 

aubertii (Bel.) Thér. in Broth. 

cagnii Negri 

leucochlorus (C. Müll.) Par. 

nano-tenax (C. Müll.) Par. 

natalensis Rehm. in Par. 

pallescens Besch. 

(homalobolax) (C. Müll.) Par. 

huallagensis Broth. 

var. weberbaueri (Broth.) J.-P. Frahm 

wurdackii Robins. 

incertus Thér. 

incrassatus C. Müll. 

alvarezianus Card. 

chalarobasis C. Müll. ex Thér. 

eximius Reichdt. 

dixonii Luis. 

falcifolius Mitt. 

flavoviridis Dus. 

flavotextus Thér. in Broth. 

rallieri Card. 

incurvatus J.-P. Frahm & Hoe 

introflexus (Hedw.) Brid. 

aurificus (C. Müll.) Kindb. 

australiensis Duby 

balanseanus Besch. 

bartramiaceus (C. Müll.) Thér. in Broth. 

berteroanus Duby 

canescens (C. Müll.) Jaeg. 

capensis Schimp. in Mitt. 

clavatus Rehm. in Sim 

collinus Par. 

cummingii Duby 

echinatus Sim 

erythropoma Duby 

exalaris (Hampe) Par. 

latipilis C. Müll. in Kindb. 

lechleri Schimp. in Lor. 

lepidophyllus (C. Müll.) Jaeg. 

leptocephalus (C. Müll.) Jaeg. 

liebmannii (C. Müll.) Schimp. in Besch. 

longescens (C. Müll.) Par. 

ochrodictyon Aongstr. 

parvicaespitosus (C. Müll.) Par. 

perincanus (C. Müll.) Kindb. 

pseudoclavatus Par. 

pseudojulaceus C. Müll. 

pudicus (C. Müll.) Jaeg. 

rectipilis C. Müll. in Kindb. 

senex (C. Müll.) Par. 

strictus Tayl. 

tasmanicus Par. 

truncatus C. Müll. 

verrucosus Besch. 

var. cordobaensis Thér. 

carinatus C. Müll. 

grimmioides (C. Müll.) Kindb. 

(itacolumitis) (C. Müll.) Broth. 

jamesonii (Hook.) Jaeg. 

altissimus (Hook.) Jaeg.

echernieri Besch. 

procerus (C. Müll.) Par. 

standleyi Bartr. 

johannis-meyeri (C. Müll.) Kindb. 

kivuensis P. Varde & Thér. 

preussii Broth. in Par. 

hoehnelii (C. Müll.) Kindb. var. 

subelamellatus Broth. in Mildbr. 

jugorum Herz. 

julaceus Jaeg. 

microjulaceus (C. Müll.) Par. 

ssp. arbogastii (Ren. & Card.) J.-P. Frahm 

julaceus Rehm. in Par. 

kunertii Par. 

pseudo-julaceus Sim 

simii Schelpe 

julicaulis Broth. 

sehnemii Bartr. 

lamellinervis (C. Müll.) Mitt. 

giganteus Sull. 

multisulcatus Duby in Hampe 

penicillatus (Hornsch.) Jaeg. 

pseudodicranum Herz. 

subpenicillatus (C. Müll.) Broth. 

tortuosus (Hampe) Par. 

var. exaltatus (C. Müll.) J.-P. Frahm 

laxitextus Lac. 

? annamensis Par. & Broth. 

calodictyon Broth. 

? celebesicus Card. 

eurydictyon Dix. ex Bartr. 

flexipilis Dix. 

fumarioli C. Müll. 

gracilentus Card. 

ludoviciae Broth. & Par. 

micholitzii C. Müll. ex Fleisch. 

nadeaudianus Besch. 

pes-funariae (C. Müll.) Par. 

polyanthus Besch. 

rugosus Besch. 

samoanus Broth. in Rech. 

sulfureus Besch. 

laxoventralis Herz. ex J.-P. Frahm 

laetevirens (Broth.) J.-P. Frahm hom. illeg. 

lonchochaete C. Müll. 

distractus C. Müll. 

holomitrius auct. 

longicellularis J.-P. Frahm 

luteus (C. Müll.) Par. 

subfulvus Thér. 

macgregorii Broth. & Geh. 

atrofuscescens Bartr. 

gedehensis Baumg. & Froehl. 

perinvolutus Thér. in Froehl. 

ruttneri Thér. in Froehl. 

megalotus Besch. 

comatulus Besch. 

milleri Ren. & Card. 

modestus Card. 

campoanus Thér. 

carbonicola Thér. 

permollis Thér. 

(nakamurae) Sak. 

nanophyllus C. Müll. ex Broth. 

brachyphyllus C. Müll. in Par. 

bryhnii Broth. 

letestui Thér. & P. Varde 

maclaudii Par. & Broth. 

subambiguus Thér. 

subbrachyphyllus Par. 

tisserantii P. Varde 

(nepalensis) (Brid.) C. Jaeg. 

nivalis (Brid.) Brid. 

anguste-limbatus Thér. & P. Varde 

arsenei Thér. 

boryanus Besch. 

chrismarii (C. Müll.) Mitt. 

commersonii Besch. in Ren. 

denticuspis Broth. in Mildbr. 

destructilis (C. Müll.) Jaeg. 

friabilis (Hampe) Jaeg. 

krauseanus (Hampe & Lor.) Jaeg. 

leptodrepanium (Broth.) Broth. 

mandonii Schimp. in Par. 

manginii Thér. 

poasensis Ren. & Card. 

sericeus Negri 

stramineus (Mitt.) Jaeg. 

subfimbriatus Thér. 

93

94 

subleptodrepanium Broth. & Thér. 

suboblongus Herz. 

substramineus Broth. in Mildbr. 

taylorii Dix. 

tolucensis Bartr. 

var. multicapsularis (C. Müll.) J.-P. Frahm 

insignis Herz. 

subannotinus Thér. & P. Varde 

leucobasis Herz. 

oblongus Thér. 

bolivianus Thér. ex J.-P. Frahm 

obrutus Thér. & P. Varde 

diversifolius (Leroy & Thér.) Takaki 

guineensis Thér. 

subobrutus Thér. & P. Varde 

usambaricus P. Varde 

occultus Mitt. 

acervatus Mitt. 

brachymitrius (Geh. & Hampe) Kindb. 

brachythysanos (C. Müll.) Par. 

cacti (C. Müll.) Par. 

calymperidictyon (Geh. & Hampe) Kindb. 

dentato-acicularis C. Müll. nom. nud. 

dicksoniae C. Müll. nom. nud. 

divisus (Geh. & Hampe) Kindb. 

fuscatus Besch. 

glauco-pallidus C. Müll. 

humifugus (C. Müll.) Kindb. 

joinvilleanus (Hampe) Jaeg. 

platyneuron (Hampe) Jaeg. 

pseudo-brachymitrius (C. Müll.) Par. 

restingae C. Müll. ex Broth. 

rufescens Broth. in Par. 

scabrellus Mitt. 

subbrachymitrius C. Müll. 

tortillisetus Ule in C. Müll. 

oerstedianus (C. Müll.) Mitt. 

mildei Limpr. 

pauper (Hampe) Mitt. 

chrysodictyon (Hampe) Mitt. 

fendleri (C. Müll.) Par. 

kingii Robins. 

propinquus (Hampe) Mitt. 

rosulatus (Hampe) Mitt. 

var. lamprodictyon (Hampe) J.-P. Frahm 

trollii Thér. 

perexilis (C. Müll.) Par. 

perichaetialis P. Varde & Thér. 

perpusillus Mitt. 

angustinervis Dix. 

cailleae Ren. & Card. 

dicranelloides Ren. & Card. 

kouroussensis Ren. & Par. 

latifolius Card. 

voeltzkowii Broth. in Voeltzkow 

pilifer Brid. 

acrocaulon (C. Müll.) Kindb. 

angustifolius Warnst. 

atro-sordidus C. Müll. ex Broth. 

bessonii Ren. & Card. 

bewsii Sim 

calochlorus Broth. & Thér. 

cinchonae Par. 

civa Lor. 

crishna Lor. 

eckendorfii Thér. & P. Varde 

erectus (C. Müll.) Mitt. 

ericetorum (Mitt.) Jaeg. 

griseolus (C. Müll.) Par. 

hoehnelii (C. Müll.) Kindb. 

hispidus Ren. & Card. 

lamellicosta (C. Müll.) Schimp. in Beschliebmannii 

(C. Müll.) Schimp. in Besch. 

leucobasis Par. 

leucotrichus Sull. & Lesq. 

longipilus Brid. p.pte. 

luridus Schimp. ex Besch. 

lutescens (C. Müll.) Jaeg. 

pachycomus Besch. in Par. 

pilosissimus Schimp. ex Besch. 

poicilophyllus Nav. 

polytrichoides De Not. 

proliferus (C. Müll.) Mitt. 

purpurascens Lor. 

sommieri Bott. in Giac. 

strictus Schimp. ex Besch. 

subbartramiaceus C. Müll. in Geh. 

subintroflexus Kindb. 

subproliferus C. Müll. in Ren. & Card. 

tenellus Rehm. in Dix. & Gepp 

transvaaliensis Rehm. in Kindb. 

tricolor (C. Müll.) Jaeg.

trichodes Lor. 

tullgrenii Ren. & Card. 

turgidus Rehm. in Dix. & Gepp 

vaporarius Bolle 

vitzliputzli Lor. 

weisiopsis (C. Müll.) Par. 

var. lamellatus (Mont.) Gradstein & Sipman 

ssp. galapagensis (J.-P. Frahm & Sipman) J.- 

P. Frahm 

pittieri Williams 

latinervis Herz. 

renneri Herz. 

praemorsus (C. Müll.) Jaeg. 

microcephalus (C. Müll.) Par. 

praetermissus J.-P. Frahm 

silvaticus Rehm. in Par. 

pseudobicolor C. Müll. ex Ren. & Card. 

purpureocaulis Dus. 

aberrans Broth. 

arboricola Card. & Dix. 


amabilis (C. Müll.) Par. 

aureo-viridis (C. Müll.) Schimp. ex Par. 

azoricus Mitt. 

bellii R. Brown ter. 

beyrichianus Duby 

blindioides Broth. in Skottsb. 

calvatus Dix. in Luis. 

campiadelphus (C. Müll.) Par. 

canaliculatus (Geh. & Hampe) Kindb. 

capillatus Hook.f. & Wils. 

distractus (C. Müll.) C. Müll. 

erythrodontius Jaeg. 

glauco-viridis (C. Müll.) Par. 

fallaciosus (Thér.) Podp. 

fibrobasius Dus. 

flavescens Rehm. in Kindb. 

kunkelii Bartr. 

laxiretis Bartr. 

lenormandii Thér. 

leptotrichaceus (C. Müll.) Par. 

lonchochaete C. Müll. 

luscinialis Dix. in Christ. 

masafuerae J.-P. Frahm 

minor Besch. 

mouensis Broth. & Par. 

muelleri Jur. in Rabenh. 

nitidus Thér. in Felipp. 

novae-valesiae Broth. 

novae-zealandiae Bartr. & Dix. 

ohingaitii R. Brown ter. 

pallidus Hook.f. & Wils. 

paludicola Broth. in Mildbr. 

pergracilis Stirt. 

sancti-caroli Besch. 

sparksii R. Brown ter. 

subareodictyon Broth. in Skottsb. 

subreconditus (Geh. & Hampe) Kindb. 

subtorquatus C. Müll. 

tijucae Broth. in Par. 

torfaceus B.S.G. 

torquatus Mitt. in Hook.f. 

trachynotus (C. Müll.) Par. 

uruguensis J.-P. Frahm 

viridicatus (C. Müll.) Par. 

woolsii (C. Müll.) Par. 

var. hamatus J.-P. Frahm & Bartlett 

recurvus (Mitt.) Jaeg. 

cataractarum (Fleisch.) Fleisch. 

wichurae Broth. in Fleisch. 

sedgwickii Dix. 

reflexisetus (C. Müll.) Broth. 

benedictii Herz. 

ptychotheca Herz. 

refractus J.-P. Frahm 

richardii Brid. 

atratus Broth. 

caracasanus (C. Müll.) Par. 

laevigatus (C. Müll.) Schimp. in Besch. 

muelleri (Hampe) Jaeg. 

nigerrimus (C. Müll.) Par. 

nigrescens Duby 

puiggarii (Geh. & Hampe) Par. 

rigidus (Hornsch.) Jaeg. 

steyermarkii Bartr. 

yunqueanus (C. Müll.) Par. 

robillardei Besch. 

alto-virescens (C. Müll.) Par. 

doliolidium C. Müll. in Par. 

heribaudii Ren. & Card. 

gallienii Par. 

inchangae (C. Müll.) Par. 

inerangae (C. Müll.) Par. 

jucrangae Par. 

longescens (C. Müll.) Par. 

macleanus Rehm. in Thér. 

pallido-nitens Thér. 

95

96 

serridorsus (C. Müll.) Par. 

sudabyanus Rehm. in Dix. & Gepp 

variegatus Dix. & Wagn. in Sim 

var. perauriculatus (Broth.) J.-P. Frahm 

brassii Bartr. 

savannarum (C. Müll.) Mitt. 

afroconcolor C. Müll. ex Broth. 

acutirameus Dix. & Thér. ex Bartr. 

angolensis Mach. & Dix. 

arenaceum (Broth.) J.-P. Frahm 

argutidens Broth. & Par. 

aspericuspis Thér. & P. Varde 

assimilis Thér. & P. Varde 

bartlettii Bartr. 

bequaertii Thér. & Nav. 

brachyphyllulus (C. Müll.) Broth. 

cambodgensis Thér. 

compactus Par. & Broth. 

delagoae (C. Müll.) Par. 

dissitus C. Müll. ex Broth. 

divaricatus (Mitt.) Jaeg. 

dusenii C. Müll. in Par. 

erythrocaulon Broth. 

gaudichaudii Besch. 

horridus Welw. & Duby 

inandae (C. Müll.) Par. 

laetus (Mitt.) Jaeg. 

latolaminatus C. Müll. in Broth. 

luluensis Thér. & Nav. 

macrotis C. Müll. ex Broth. 

mathieui Thér. 

montanus Welw. & Duby 

pobeguinii Par. & Broth. 

salesseanus Par. & Broth. 

serricuspis Thér. & P. Varde 

schiffneri (Broth.) J.-P. Frahm 

sciuroides Welw. & Duby 

siamensis Dix. & Thér. 

sprucei Mitt. 

? tortilipilus J.-P. Frahm 

viridatulus C. Müll. ex Broth. 

(scoposum) Hampe ex C. Müll. 

(searellii) R. Brown ter. 

sericeoides Dix. 

serratus Lac. 

demangei Thér. & P. Varde 

scabridorsus Dix. 

singaporensis Fleisch. & Par. 

pinangensis Thér. in Henry 

schimperi Milde 

alpigena Broth. 

compactus Schimp. in Hunt. 

handelii Broth. 

schmidii (C. Müll.) Jaeg. 

aureus Bosch & Lac. 

calvus Ren. & Card. 

chryseolus C. Müll. ex Besch. 

deciduus Ren. & Card. 

herzogii Broth. 

merapicola C. Müll. 

nodiflorus (C. Müll.) Jaeg. 

ripicola Besch. 

robbinsii Bartr. 

subtricolor Lor. 

ssp. hemitrichius (C. Müll.) J.-P. Frahm 

copelandii Broth. 

diversinervis Broth. 

foxworthii Broth. 

skottsbergii Broth. 

tubulosus Bartr. 

wallisii Sauerb. in Jaeg. 

setifolius Wils. 

porophorus Stirt. 

sharpii J.-P. Frahm, Horton & Vitt 

shawii Wils. in Hunt 

brittonae Williams 

carreiroanus Card. 

underwoodii Williams 

sinensis (C. Müll.) J.-P. Frahm 

? aoraiensis H. Whittier 

eberhardtii Par. 

excurrens Dix. 

fusco-viridis (Card.) Dix. & Thér. 

irrigatus Thér. 

japonicus Broth. 

pseudomuelleri Card. 

saint-pierrei Thér. 

uii Broth. 

smaragdinus (Brid.) Jaeg. 

mangenotii Biz. & Kilb. 

mollis C. Müll. in Par. 

parisii Ren. 

subpulvinatus P. Varde & Thér. 

spiralis Dus. 

subcuspidatus (Hampe) Jaeg. 

atratus Bartr. 

brotherianus Par.

paramoensis Bowers 

praealtus (C. Müll.) Par. 

procerus Broth. 

var. damazii (Broth.) J.-P. Frahm 

(subfalcatus) (Hornsch.) Jaeg. 

subfragilis Ren. & Card. 

subjugorum Broth. 

subluteus (Mitt.) Jaeg. 

subnitens Kaal. 

(subtrachyblepharus) C. Müll. 

subulatus Schimp. in Rabenh. 

barbuloides Broth. in Bruehl 

brevifolius Schimp. 

gracilis (Mitt.) Jaeg. 

fusco-luteus Stirt. 

latinervis (Mitt.) Jaeg. 

surinamensis C. Müll. 

catumbensis Broth. 

costaricensis Bartr. 

donnellii (Aust.) Lesq. & Jam. 

gracilicaulis Mitt. 

marmellensis Broth. 

minarum Par. 

subleucogaster (C. Müll.) Jaeg. 

(taiwanensis) Sak. 

tallulensis Sull. & Lesq. 

tequendamensis Thér. 

terebrifolius (C. Müll.) Jaeg. 

thwaitesii (Mitt.) Jaeg. 

angusticosta J.-P. Frahm 

angustinervis Herz. 

capensis Broth. 

chlorotrichus (C. Müll.) Par. 

cardotii Thér. 

controversus (Hampe) Jaeg. 

flaccidus (C. Müll.) Broth. 

fusco-croceus (Hampe) Jaeg. 

hildebrandtianus Fleisch. 

humoricola (C. Müll.) Par. 

macrogaster (C. Müll.) Broth. 

mosenii Broth. 

orthopelma (C. Müll.) Par. 

ouropretensis Par. in Broth. 

pleurocarpus (C. Müll.) Par. 

pseudodichrostis Broth. 

stenopelma (C. Müll.) Rehm. ex Par. 

stramineolus C. Müll. 

subulifolius Thwait. & Mitt. 

tenax (C. Müll.) Par. 

viridatus (C. Müll.) Broth. 

torrentis Thér. & P. Varde 

trachyblepharon (C. Müll.) Mitt. 

arenicola (C. Müll.) Mitt. 

bermudianus Williams in Britt. 

detonsus (Hampe) Kindb. 

schwaegrichenii Duby 

subarenicola C. Müll. ex Broth. 

villicaulis (Hampe) Jaeg. 

ssp. comatus (Ren. & Card.) J.-P. Frahm 

subcomatus Ren. & Card. 

trichophylloides Thér. in Herz. 

trivialis C. Müll. ex Britt. 

tener Broth. 

cuatrecasii Robins. 

97 

uleanus (C. Müll.) Broth. 

umbellatus (Arn.) Par. 

alatus (Broth. ex Ihs.) Iwats. & Nog. 

blumei (Dozy & Molk.) Bosch & Lac. 

coreensis Card. 

depallieri Card. & P. Varde 

didrichsenii (C. Müll.) Jaeg. 

dozyanus (C. Müll.) Jaeg. 

ferrieri Broth. in Par. 

foreauanum Card. & P. Varde 

gibboso-alaris (Broth. & Par.) Wijk & Marg. 

hawaiicus (C. Müll.) Jaeg. 

leptoneuron Broth. ex Ihs. 

microrhodon C. Müll. in Par. 

nagasakinus Broth. in Par. 

nigrescens (Mitt.) Jaeg. 

nudicaulis Besch. 

obscurus Aongstr. 

oedicaulis C. Müll. in Par. 

percristatus Thér. 

pertristis (C. Müll.) Par. 

powellii (C. Müll.) Par. 

? sandwicensis Par. 

scabripilus Warnst. 

subexasperatus (C. Müll.) Jaeg. 

vernieri Duby 

viridulus Card. 

var. pterotoneuron (C. Müll.) J.-P. Frahm 

geniculatus Aongstr.

98 

valerioi B.H. Allen 

validinervis (Dix.) B.C. Tan 

vesticaulis Mitt. in Mell. 

dicranodontioides Broth. in Skottsb. 

guaitecae Dus. 

hygrophilus Broth. in Skottsb. 

paludosus J.-P. Frahm & Bartlett 

saddleanus (C. Müll.) Besch. 

saxicola Broth. in Skottsb. 

turficola Broth. in Skottsb. 

widgrenii (C. Müll.) Mitt. 

tijucae Broth. in Par. 

wheeleri (C. Müll.) Hampe ex Par. 

zygodonticarpus (C. Müll.) Par. 

LITERATURE 

Allen, B. 1990. Campylopus atlanticus (Musci: 

Dicranaceae), a New Species from Panama, with 

Comments on the Taxonomy of Campylopus. 

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Allen, B. 1989. A preliminary treatment of the genus 

Campylopus in Central America. Tropical 

Bryology 1: 63-94. 

Arts, T. 1987. The occurrence of tubers in Campylopus 

pyriformis. Lindbergia 12: 125-128. 

Arts, T. 1989. The occurrence of rhizoidal tubers in 

the genus Campylopus. Lindbergia 15:60-64. 

Arts, T. 1993. Rhizoidal tubers in the genus 

Campylopus and their systematic relevance. 

Lindbergia 18: 31-40. 

Arts, T. & Frahm, J.-P. 1990. Campylopus pyriformis 

new to North America. The Bryologist 93: 290- 

294. 

Bartlett, J.K. and J.-P. Frahm. 1983. Notes on 

Campylopus and Chorisodontium from New 

Zealand. J. Bryol. 12: 337-341. 

Bizot, M., G. Kilbertus 1979. Les Campylopus 

africains subg. Lucidus Biz. & Kilb. Rev. Bryol. 

Lich. 45: 61-95. 

Bizot, M., R.B. Pierrot & T. Pócs 1974. Trois genres 

nouveaux des muscinées. Rev. bryol. lichénol. 

N.S. 40: 25-31. 

Catcheside, D.G. and J.-P. Frahm. 1985. Additions 

to the Campylopus flora of Australia. J. Bryol. 

13: 359-367. 

Corley, M.V.F. 1976. The taxonomy of Campylopus 

pyriformis (Schultz) Brid. and related species. 

Journal of Bryology 9: 193-212. 

Corley, M.F.V. & J.-P. Frahm. 1982. Taxonomy and 

world distribution of Campylopus pyriformis 

(Schultz) Brid. Journal of Bryology 12(2): 187- 

190. 

Crundwell, A.C. 1979. Rhizoids and Moss 

Taxonomy. In: G.C.S. Clarke and J.G. Duckett, 

Bryophyte Systematics pp. 347-363. London. 

Egunyomi, A. 1981. How reliable are the specific 

characters in the genus Campylopus? Misc. 

Bryol. Lich. 9: 41-44. 

Florschütz, P.A. & J. Florschütz-de Waard 1974. 

Studies on Colombian Cryptogams I. Variation 

of characters in Southamerican species of 

Campylopus. J. Hattori Bot. Lab. 38: 111-114. 

Florschütz - de Waard & M. Worrell - Schets 1980. 

Studies on Colombian Cryptogams VII. Culture 

studies on the taxonomic relevance of costal 

anatomy in the Campylopus leucognodes - 

subconcolor complex and in Campylopus 

pittieri. Proc. Koninkl. Nederl. Akad. Wetensch. 

Series C, vol. 83(1): 37-45. 

Frahm, J.-P. 1974. Zur Unterscheidung und Verbreitung 

von Campylopus introflexus (Hedw.) Brid. 

und C. polytrichoides De Not. Revue Bryol. 

Lichénol. 40: 33-44. 

Frahm, J.-P. 1975. Conspectus der mittel- und südamerikanischen 

Campylopus-Arten 

(Dicranaceae). Bryoph. Bibl. 5, 144 pp. Vaduz. 

Frahm, J.-P. 1975a Revision der Gattung 

Thysanomitrium in Südamerika. J. Bryol. 8: 255- 

264. 

Frahm, J.-P. 1975b Taxonomische Notizen zur Gattung 

Campylopus. Rev. Bryol. Lichénol. 41: 

321-332. 

Frahm, J.-P. 1976. Taxonomische Notizen zur Gattung 

Campylopus II. Rev. Bryol. Lichénol. 42: 

603-616. 

Frahm, J.-P. 1976a. Taxonomische Notizen zur Gattung 

Campylopus III. Rev. Bryol. Lichénol. 42:

891-908. 

Frahm, J.-P. 1976b. Taxonomische Notizen zur Gattung 

Campylopus IV. Nova Hedwigia 28: 131- 

136. 

Frahm, J.-P. 1977. Zur Campylopus-Flora von 

Mexico. Bryologist 80: 119-124. 

Frahm, J.-P. 1977a. Ventrale stereidale Bildungen 

im Rippenquerschnitt von Campylopus flexuosus 

(Hedw.) Brid. Herzogia 4: 249-264. 

Frahm, J.-P. 1978. Übersicht der Campylopus-Arten 

der Anden. J. Hattori Bot. Lab. 44: 483-524. 


Campylopus V. Nova Hedwigia 29: 243- 

250. 


Campylopus VI. Nova Hedwigia 30: 835- 

849. 

Frahm, J.-P. 1979. Die Campylopus-Arten Brasiliens. 

Rev. Bryol. Lichénol. 45: 127-178. 

Frahm, J.-P. 1979a. New Campylopus records from 

Mexico, including C. sharpii sp. nov. Bryologist 

82: 623-625. 

Frahm, J.-P. 1979b. Ergänzungen zur Campylopusflora 

von Mexico. Herzogia 5: 111-117. 

Frahm, J.-P. 1979c. Taxonomische Notizen zur Gattung 

zur Gattung Campylopus VII. Nova 

Hedwigia 31: 427-433. 

Frahm, J.-P. 1979c. Campylopodes Brasiliae 

Exsiccatae. Herzogia 5: 103-110. 

Frahm, J.-P. 1980. An Analysis of the Campylopus 

cavifolius Complex. J. Hattori Bot. Lab. 48: 195- 

210. 


Campylopus IX. J. Bryol. 11: 213-218. 

Frahm, J.-P. 1980b. Synopsis of the genus 

Campylopus in North America north of Mexico. 

Bryologist 83: 570-588. 

Frahm, J.-P. 1981. Ergänzungen zur Campylopus- 

Flora von Brasilien. Cryptogamie Bryol. 

Lichénol. 2: 441-448. 

Frahm, J.-P. 1981a. New Campylopus records from 

South and Central America. Bryologist 84: 242- 

243 


Campylopus X. Lindbergia 7: 27-31. 


Campylopus VIII. Herzogia 5: 517-529. 

Frahm, J.-P. 1981d. Taxonomische Notizen zur Gattung 

Campylopus XI. Nova Hedwigia 34: 391- 

395. 

99 

Frahm, J.-P. 1982. A reinterpretation of 

Bryohumbertia P. de la Varde et Thér. 

Cryptogamie Bryol. Lichénol. 3: 365-369. 

Frahm, J.-P. 1982a. Grossdisjunktionen von Arealen 

südamerikanischer und afrikanischer 

Campylopus-Arten. Lindbergia 8:36-45. 

Frahm, J.-P. 1982b. New Campylopus Records from 

South and Central America II. Bryologist 85: 

322-323. 


Campylopus XII. Cryptogamie Bryol. 

Lichénol. 3: 59-66. 

Frahm, J.-P. 1982d. Problems of infrageneric 

classification in Campylopus. Nova Hedwigia 

Beih. 71: 219-224. 

Frahm, J.-P. 1983. A monograph of Pilopogon Brid. 


Frahm, J.-P. 1983a. A new infrageneric classification 

of the genus Campylopus Brid. J. Hattori Bot. 

Lab. 54: 207-228. 

Frahm, J.-P. 1984. Campylopus Brid. 1. Subgenus 

Thysanomitrion (Schwaegr.) Kindb. emend. J.- 

P. Frahm. Nova Hedwigia 39: 585-621. 

Frahm, J.-P. 1984a. A survey of the genus 

Campylopus in Sri Lanka. J. Bryol. 13: 163-192. 

Frahm, J.-P. 1984b. Dicranaceen aus Peru. Nova 

Hedwigia 39: 145-167. 

Frahm, J.-P. 1984c. A Review of Campylopodiella. 

Bryologist 87: 249-250. 

Frahm, J.-P. 1985. Afrikanische Campylopus-Arten. 

Bryophyt. Bibl. 31. 216 pp. 


Campylopus XIII. Cryptogamie Bryol. 

Lichénol. 6: 205-208. 


Campylopus XIV. Nova Hedwigia 41: 273- 

277. 

Frahm, J.-P. 1986. Ecosystematics and Geotaxonomy 

of the Campylopodioideae. Bryol. Times 36: 1- 

4. 

Frahm, J.-P. 1986a. A review of Sphaerothecium 

Hampe. Bryologist 89: 152-154. 

Frahm, J.-P. 1986b. Campylopus Brid. 2. Subg. 

Campylopidulum Vital. Nova Hedwigia 43: 221- 

227. 

Frahm, J.-P. 1986c. New Campylopus Records for 

Central and South America III, with the 

description of Campylopus cleefii J.-P. Frahm 

spec. nov. and C. longicellularis J.-P. Frahm 

spec. nov. Cryptogamie Bryol. Lichénol. 7: 439-

100 

446. 

Frahm, J.-P. 1987. A revised list of the Campylopus 

species of the world. Bryologische Beiträge 7: 

1-117. 

Frahm, J.-P. 1987. A survey of the Campylopus 

species of Australia. J. Bryol. 14: 701-728. 

Frahm, J.-P. 1987a. The ecological significance of 

the costal anatomy in the genus Campylopus. 

Abstracta Bot. 9 Suppl. 2: 159-169. 

Frahm, J.-P. 1988. The subantarctic and southern 

hemispheric species of Campylopus 

(Dicranaceae), with contributions to the origin 

and speciation of the genus. J. Hattori Bot. Lab. 

64: 367-387. 

Frahm, J.-P. 1989. Bryohumbertia subcomosa (Dix.) 

J.-P. Frahm, a new name for Bryohumbertia 

walkeri (Mitt.) J.-P. Frahm. Trop. Bryol. 1: 9- 

10. 

Frahm, J.-P. 1990. Campylopus, a modern and 

successful genus !? Tropical Bryology 2: 91-101. 

Frahm, J.-P. 1990. The origin and distribution of 

Neotropical species of Campylopus. Tropical 


Frahm, J.-P. 1990. A short survey of the Campylopus 

flora of New Caledonia. Cryptogamie, Bryol. 

Lichénol. 11: 369-375. 

Frahm, J.-P. 1990. Campylopus laxitextus Lac. new 

to Australia. J. Bryology 16: 305-306. 

Frahm, J.-P. 1990. Phytogeography of African 

species of Campylopus. J. Bryology 16: 289- 

304. 

Frahm, J.-P. 1991. Campylopodioideae, 

Paraleucobryoideae. Flora Neotropica 

Monograph 54.. 238 SS. 

Frahm, J.-P. 1991. A survey of the Campylopus flora 

of Hawaii. Bryologist 94: 60-66. 


Campylopus XV. Tropical Bryology 4: 59- 

61. 

Frahm, J.-P. 1992. The SE-Asian species of 

Campylopus. Origin, evolution and distribution. 

Bryobrothera 1: 221-230. 

Frahm, J.-P. 1992. A revision of the East-Asian 

species of Campylopus. J. Hattori Bot. Lab. 71: 

133-164. 

Frahm, J.-P. 1993a. Campylopus japonicus New to 

North America North of Mexico. Bryologist 96: 

142-144. 

Frahm, J.-P. 1993b. Taxonomic Results of the 

BRYOTROP Expedition to Zaire and Rwanda. 

17. Andreaeaceae, Bruchiaceae, Dicranaceae, 

Rhizogoniaceae, Bartramiaceae, 

Rhacocarpaceae, Hedwigiaceae, Cryphaeaceae, 

Leucodontaceae. Ztrop. Bryol. 8: 153-169. 

Frahm, J.-P. 1994a. The affinities between the 

Campylopus floras of Sri Lanka and Madagascar 

- or: which species survived on Noah´s arc? 

Hikobia 11: 371-376. 

Frahm, J.-P. 1994b. A new synopsis of the 

Campylopus species of Australia. Jouirnal of 


Frahm, J.-P. 1994c. New records of Campylopus 

(Musci, Dicranaceae) from Central and South 

America IV. Fragm. Flor. Geobot. 39: 77-84. 

Frahm, J.-P. 1994d. A contribution to the 

differentiation of Campylopus subulatus and C. 

tallulensis in North America. Evansia 11(3): 95- 

99. 

Frahm, J.-P. 1994e. Taxonomische Notizen zur Gattung 

Campylopus XVI. Nova Hedwigia 59: 147- 

155. 

Frahm, J.-P. 1996. Campylopus extinctus n. sp. 

(Musci, Dicranaceae), an apparently extinct 

species from Brazil. The Bryologist 99: 218-220. 

Frahm, J.-P. 1997. A taxonomic revision of 

Dicranodontium (Musci). Acta Botanica Fennici 

34: 179-204. 


Campylopus XVII. Cryptogamie, Bryol. 

Lichènol. 39: 27-34. 

Frahm, J.-P. & Arts, Th. 1990. Campylopus 

pyriformis (K.F.Schultz) Brid. New to North 

America. Bryologist 93: 290-294. 

Frahm, J.-P., M. Giese, M. Padberg, T. Koponen 

and D.H. Norris. 1985. Bryophyte flora of Huon 

Peninsula, Papua New Guinea IX. 

Atractylocarpus, Bryohumbertia, 

Campylopodium and Campylopus (Dicranaceae, 

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Frahm, J.-P. & P. Geissler 1985. Lectotypification 

von Dicranum flexuosum Hedw. (Campylopus 

flexuosus (Hedw.) Brid.) Cryptogamie Bryol. 

Lich. 6:287-290. 

Frahm, J.-P. and S.R. Gradstein. 1987. The genera 

Bryohumbertia and Campylopus (Musci) in the 

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320. 

Frahm, J.-P. & E. Hegewald, 1979. Eine Moossammlung 

aus Peru II. Campylopus. Nova 

Hedwigia 31: 435-447.

Frahm, J.-P. & Hoe, W.J. 1978. Campylopus 

incurvatus: A new species of the Dicranaceae 

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Frahm, J.-P. & P. Isoviita 1988.Type revision of 

European mosses 2. Atractylocarpus, 

Campylopus and Paraleucobryum. Lindbergia 

14: 89-96. 

Frahm, J.-P. and H. Mohamed. 1987. A survey of 

Campylopus and Bryohumbertia (Dicranaceae) 

in Malaysia. Mem. New York Bot. Garden 45: 

470-491. 

Frahm, J.-P., Norris, D.H. & T. Koponen 1988. 

Bryophyte Flora of the Huon Peninsula, Papua 

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Frahm, J.-P. & H.J.M. Sipman 1978. Eine neue 

Campylopus-Art von den Galapagos Inseln. J. 

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Frahm, J.-P. & Vitt, D.H. 1978. A taxonomic study 

of Campylopus schimperi and C. subulatus 

(Bryopsida: Dicranaceae) in North America. 

Brittonia 30: 365-372. 

Frey, W. & J.-P. Frahm 1987. The twist mechanism 

in the cygneous setae of the genus Campylopus. 

Morphology, structure and function. Nova 

Hedwigia 44: 291-304. 

Geiger, H. & Markham, K.R. 1992. 

Campylosaurone, an auronoflavonone 

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Campylopodium (C. Müll.) Besch. Lindbergia 

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101 

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165. 

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two Cladonia species and Campylopus 

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102

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