GRAPHIS ScnIPTA - Universitetet i Oslo

GRAPHIS ScnIPTA 

Volym 4, hiifte I, 1992 

Nordisk Lichenologrsk Forening

Nordisk Lichenologisk Fiirening (NLF) 

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Stockholm juni L99z 

ISSN 0901-7593

Lichens from the coast of Eniseiskii Gulf, W. Taimyr, Siberia 

MICHAIL P. ZHURBENKO 

Zhurbenko, M. P. L992: Lichens from the coast of Eniseiskii gulf, 'W. 

Taimyr, Siberia. Graphis Scripta 4: I-4. Stockholm. ISSN 0901 -7593. 

A list of 141 tara of lichens from the coast of Eniseiskii gulf, 'W. Taimyr, 

Siberia and their ecotopic distribution is presented. Eight species, including a 

few amphiberingian, are reported as new to Taimyr. The "active" (thriving) 

species of the local lichen flora are selected. 

Michail P. Zhurbenko, Laboratory of Lichenologt and Bryolog, Komarov 

Botanical Institute, Prof. Popov str. 2, St. Petersbrrg, 197376, Russia. 

Although the lichen flora of the Taimyr 

peninsula is better studied than most of the 

other regional floras of the Russian Arctic 

(Piin I979a, L979b, 1984, Piin & Martin 1978, 

Piin & Trass l97l), there still remains much 

to do for lichenologists. 

In July 1990, the author investigated the 

lichen flora at the coast of Eniseiskii gulf close 

to the mouth of Ragozinka river, 72" 48' N, 

80o 53' E, W. Taimyr, Siberia. This paper 

covers only part of the collected specimens, 

mainly macrolichens. 

Materials and methods 

As a rule, the material was collected on 

sample plots situated within a definite type of 

habitat. Usually such plots were measuring 25 

x 25 m. The total number of plots was 31. 

Some specimens were also collected outside 

the plots. The lichen flora was investigated 

within the radius of about five km from the 

base camp. Specimens were collected and 

determined by the author, unless othenvise 

stated. They are deposited at the Cryptogamic 

Herbarium of the Komarov Botanical Institute 

St. Petersburg (LE). 

For the chemical investigation, only spot 

tests were used. 

Climate and vegetation 

The climate is continental with a large temperature 

amplitude, a cold winter and a small 

annual precipitation (300-400 mm/year). The 

mean temperature in January is about minus 

27 oC, in July about plus 7 oC (Atlas of the 

Arctic 1985). A continuous snow cover melts 

in the middle of June and forms again at the 

end of September. According to the zonal 

classification (Yurtsev et al. l97S) the vegetation 

is referred to the southern arctic tundra. 

Favourable mesic habitats on placors 

(interfluves) are occupied by herb-dwarf 

shrub-moss(-lichen) tundras dominated by 

e.g. Carex ensifulia, Dryas punctata and 

Hylocomium splendens. Scrub communities 

occur only in the depressions of the relief. 

Type of habitats 

The following set of habitats was used for the 

ecotopic analysis of the local lichen flora. 

Accepted abbreviations are used hereafter in 

the annotated list of taxa. Number of plots in 

brackets: 

WS - willow shrubs (Salix reptans, S. lanata) 

along streams in valleys and in other depressions 

of the relief. (0)

2 Michail P. Zhurbenko GRAPHIS SCRIPTA 4 (199?) 

T - Polygonal spotted tundra (herb/dwarf St - Stone fields, of no calciferous content. (3) 

shrub/moss (lichen)) on placors (interfluves). Sw - stone fields in wet situations, e.g. along 

(4) water courses. (1) 

Md - herb-grass meadows on S-facing SS - stone fields covered by snow beds or 

slopes. (2) bordering them. (2) 

Dr - Dryas communities on well-drained, SR - stone fields together with rocky out- 

open tops ofhills. (2) crops. (3) 

W - sedge-cotton-grass tundra in wet stands, 

e.g. temporarily flushed sites along streams. 

(1) 

Sn - snow beds where snow persists until the The taxa and their habitsts 

middle of July or later. (3) 

Et edge of the coastal terrace, in rather 

moist environments. (1) 

Mr - grass-hypnum and polygonal mires. (2) 

Dw - drift-wood and old wood of the traps 

for polar fox in tundra. (5) 

EOc 

812 

44 

11 

34 

22 

11 

43+ 

44 

44 

78 

7 1t 

44 

47 

44 

14 

31 

10 15 

713 

69 

45 

t2 24 

44 

22 

14 

56 

R - outcrops of bed-rocks (basalt, diabases). 

(2) 

E total number of the developed Upes of 

habitats. 

Oc - occurrences in the 31 sample plots. 

+ = 1 finding outside sample plots (ecotype). 

C. odontella: SR 1 1 

C. pinasln: Dw 1 5 

C. sepincola: Dw 1 4 

Cladina arbuscula:T, ET, Mr, Dw, 

SI,SR,R 7 10 

C. rangifeina: T, Mr 2 4 

C.stygia:T 1 1 

Cladonia acuminata: Md 1 1 

C. amaurocraea: W, Mr, Dw, St, SS, 

SR,R 7 

C. bacilliformis: T, Mr 2 

C. bellidiflora: T, ET, Mr, Dw, St, SR, R 7 

11 

2 

8 

C. cariosa: Dw, R 

2 2 

C. canteola: Dw 

1 1 

C. cenotea: Mr, Dw 

2 2 

C. chlorophaea s. lat.: T, Md, 

Dr, St, SS, SR, R 

78 

C. coccrftro: T, Dr, Sn, ET, Mr, 

Dw, St, SS, SR, R 

C. cornuta: Mr, Dv 

C. qanipes: T, Mr, Dw 

C. deformrs: Dw 

C. ecmocyna: T, Md, Dr, Sn, ET, 

Mr, Dw, St, SS, SR, R 

C. fimbriata: T, Dw, R 

C. furcata: T, Mr 

C. gracilrs: T, R 

C. macrophylla: ET, Dw, St, SR, R 

C. phyllophora: St, SR 

10 16 

22 

36 

11 

11 20 

35 

23 

21 

55 

22

GRAPHTS SCRTPTA 4 (1992) 

C. pleurota: Mt, Dw, St, SR, R 

C. pocillum: T, Md, Dr, Sn, ET, 

Dw, SS, SR, R 

C. pyxidata: T, Md, Dr, Sn, ET, 


C. squamosa: T, Mr, Dw, St, SR, R 

C. sticta: T, Dr, W, Sn, ET, 


C. cf. subcervicomls: SS 

C. subfurcata: T, Sn, SS 

C. sulphurina: Dw 

C. uncialrs: T, ET, Mr, Dw, St, SR, R 

Coelocaulon aculeatum: Dr 

Coriscum virid,e: Mr 

Dactylina arctica: T, Md, Dr, Sn, 

Mr, Dw, St, SR, R 

D. beingica: Md 

D. ramulosa: T, Md, Dr, Dw, SR, R 

Evernia mesomorpha: Dw 

Foraminella ambrya: Dw 

F. hyperopta: Dw 

Hypogmnia austerodes: Dw 

H. physodes: Dw, St, SR 

H. subobscura: T, Dr, Dw, St, SR, R 

Lobaia linita: T, Md, Dr, W, Sn, ET, 

Mr, Dw, St, SR, R 

Melanelia granulosa: St, SR, R 

M. infumata: Dw, St, SR, R 

M. pannifurmis: St, SR, R 

M. septentionalis: Dw 

M. sorediata: St, SS, R 

M. stygia: St, SS, SR, R 

M. substygta: St, SR 

Nephroma arcticumi T, Mr, St 

N. expallidum: T, Md, W, Sn, Mr, 

Dw, St, SS, SR, R r 

Ochrolechia frigida: T, Md, Dr, Sn, 

ET, Mr, Dw, St, SS, SR, R 

Ochrolechia upsaliensrs: T, Md, Dr, 

Dw, St 

Pannaia pezizoides: T, Md, Dr, W, 

ET, Dw, SS, R 

P. praetermissa: Md, Sn, St, R 

Parmelia fraudans: St, SR, R 

P. omphalodes: T, Dr, Dw, St, SR, R 

P. saxatilis: Dw, St, SS, SR, R 

P. sulcata: Dw, St, SR, R 

P arme lie II a triptophylla f . onegen.rrr 

56 

913 

11 t9 

66 

11 20 

11 

33 

12 

79 

11 

11 

9 

1 

6 

1 

I 

1 

1 

6 

6 

L4 

1 

8 

1 

4 

1 

1 

11 

11 

11 L7 

36 

47 

35 

12 

33 

46 

22 

32 

10 L7 

11 L9 

5 

9 

4 

3 

5 

5 

4 

1 

13 

4 

4 

L2 

11 

9 

1 

Lichens from Eniseiskii 3 

Peltigera aphthosa:T,Md, Dr, Sn, 

Mr, Dw, St 

P. didactyla: T, Md, Dr, Sn, Dw, 

St, SS, SR, R 

P. lepidophora: T, Dr, Sn, St, R 

P. leucophlebia: T, Md, Dr, W, Sn, 

Mr, Dw, St 

P. malacea: T, Md, Dr, W, Sn, Dw, 

St, SR, R 

P. rufescens: T, Md, Dr, W, Sn, 

Mr, Dw, St, R 

P. scabrosa: T, Md, Mr, Dw, St, SR 

P. venosa: T, Dw, R 

Pertusaia bryontha: T, Md 

P. dactylina: T, Mr, Dw, St, SR 

P. geminipara: T, ET, SS, SR, R 

P. oculata: Dw, SS, SR 

P. panyrga: T, Dr, St, SR 

P. subdactylina: T, Dr, Dw, St,R 

Phaeocalicium praecedens: WS 

Phaeophyscia cf. endococcina: SS 

P. kairamoi: R 

P. cf. nigricanr: R 

P. sciastra: St, SR, R 

Physcia aipolia v. alnophila: Ws, Dw 

P. caesia: WS, St, SR, R 

P. dubia: WS, Dw, St, SS, SR, R 

Physconia detersa: St, R 

P. muscigena: WS, Dr, St, SR, R 

Pseudephebe minuscula: SR 

P. pubescens: SR 

Psoroma hypnontm: T, Md, Dr, W, Sn, 

ET, Mr, Dw, St, SS, SR, R 

Mizoplaca chrysoleuca: R 

Soloina crocea: Dw, St, SR, R 

Sphaeorophorus fragilis: St, SR, R 

S. globo,rr{.r: T, Dr, Mr, Dw, St, SR, R 

Stereocaulon alpinum: T, Md, Dr, Sn, 


S. botryosum: Dw, St, SS, SR, R 

S. capitellatum: R 

S. glareosum: W, Sn, Dw, SS 

S. groenlandicurn: St, SS, SR, R 

S. ivulorum: T, Dr, W, Sn, ET, Dw, 

SS, SR 

S. subcoralloides: SS, SR, R 

Sticta arctica: T, Dr, Sn, Dw, R 

710 

915 

46 

89 

913 

916 

68 

33 

22 

56 

s5 

34 

48 

57 

11 

11 

12 

11 

33 

22 

47 

68 

23 

59 

11 

11 

t2 18 

11 

44 

33 

712 

10 22 

56 

11 

44 

44 

813 

33 

59 

I

4 Michail P. Zhurbenko 

Thamnolia subultformis: T, Md, Dr, 

Mr, St, SR, R 7 15 

T. vermicularis: T, Md, Dr, Mr, St, SR, R7 13 

Umbilicaia arctica: SR 1 1 

U. qlindrica: SS, SR, R 3 6 

U. decussata: SR, R 2 4 

U. deusra: SS, SR 2 3 

U. hyperborea: SR, R 2 4 

Discussion 

The following species are new to Taimyr: 

Dactylina beringica, Melanelia substygia, 

Parmeliella tiptophylla f. onegensis, Phaeocalicium 

praecedens, Physcia aipolia v. 

alnophila, Physconia detersa, Stereocaulon 

capitellatum and Umbilicaria tyngei. Three 

species, which were previously known from 

single findings were confirmed: Asahinea 

scholanderi (known from Tareia), Corbcum 

viride and Phaeophyscia kairamor (both 

species were known from Kresty Taimyrskie). 

The occurrence of Asahinea scholanderi, and 

Dactylina beringica at western Taimyr is of 

interest taking into account the amphiberingian 

pattern of distribution of these 

species. The most "active" (thriving) lichen 

species sensu B. A. Yurtsev (Yurtsev et al. 

1978) are those which thrive in the highest 

number of habitats in the landscape and play 

the most important role in the plant cover. 

These are Cetraia islandica, Stereocaulon 

alpinum, Cladonia ecmoqna, C. sticta, 

Ochrolechia figida, Cladonia plaidata, 

Psoroma hypnotuffi, Lobaia linita, Nephroma 

etcpalliduffi , Peltigera rufescens, Cladonia cocctfera, 

Cetraria cucullata, Peltigera didactyla, 

Thamnolia subuliformis and Dactylina arctica. 

Acknowledgements 

Thanks are due to the following persons help 

with determinations: R. Moberg (Physciaceae), 

E. S. Hansen (Coriscum viride), G. B. 

Feige (Umbilicaria spp.), P. M. Jorgensen 

(Parmeliella triptophylla) and A. N- Titov 

GRAPHIS SCRIPTA 4 (Leez) 

U. Iyngei: SR 

U. proboscidea: Dw, SR, R 

U. torrefacta: St, SR 

Usnea sp.: Dw 

Xanthoia candelaia: Dw, SR, R 

X. elegans: St, SW, SS, SR, R 

X. sorediata: St 

(Phaeocalicium praecedens). I am indebted to 

N. Matveeva and L. Zanoha for the opportunity 

to take part in their expedition to 

Taimyr in 1990. The English was revised by U. 

Sochting. 

References 

11 

34 

z1 

11 

36 

56 

11 

Atlas of the Arctic. 1985: Treshnikov, A. F. 

(ed.).Moscow. 

Piin, T. L979a: Terricolous lichens of the cape 

Cheliyskin. In: Arctic tundra and polar 

deserts of Taimyr: 6'l'-73. I-eningrad. 

Piin, T. 1979b: Terricolous lichens in the 

vicinity of the bay Mariia Pronchisceva. 

In: Arctic tundra and polar deserts of 

Taimyr: l4-I43. lrningrad. 

Piin, T. 1984: Flora and distribution of terricolous 

lichens in the southern tundra of 

Taimyr. In: Flora and groupings of lower 

plants in natural and antropogenous 

extreme environment conditions: L34-172. 

Tallinn. 

Piin, T. & Martin, Y. 1978: Flora of terricolous 

lichens. In: Ary-Mas: 101- I23. 

kningrad. 

Piin, T. & Trass, H. L97l: Terricolous lichens 

in the vicinity of Tareia (W. Taimyr). In: 

Biogeocenoses of Taimyr tundra and their 

productivity: 15 1 - 159. Irningrad. 

Yurtsev, B. A., Tolmatchev, A. I. & Rebristaja, 

O. V. 1978: The floristic delimitation 

and subdivision of the Arctic. In: Yurtsry 

B. A. (ed.), The Arctic floristic region: 9- 

105. Irningrad.

Anteckningar om nfrgra arter av Peltigera canina-gruppen i 

Sverige 

GUNNAR CARLIN 

Carlin, G. 1992: Anteckningar om n8gra arter av Peltigera canin4-gruppen i 

Sverige. [Notes on the Swedish species of the Peltigera canina group.J 

Graphis Scipta 4: 5-17. Stockholm. ISSN 0901-7593. 

During recent years, several new species of Peltigera have attracted attention 

in Sweden. As a consequence, several of the specific characters commonly 

used in examination keys and short descriptions are now obsolete. The present 

paper describes the large tomentose Peltigera-species P. canina, P. 

kristinssonii, P. membranacee, P. ponojenrrJ, P. praetextata, P. retifuveata and 

P. rufescens. The authors ambition was to describe characters useful for the 

identification of the species, rather than attempting to cover the morphology 

completely. Short notes on the ecology and distribution within Sweden are 

included. 

Gunnar Carlin. The Herbaium, University of Uppsala, Box 541, 5-751 21 

Uppsala, Sweden. 

Det har varit relativt Htt att med korta uttryck 

karaktiirisera arterna i Peltigera caninagruppen: 

P. praetactata har isidielika bildningar, 

P. rufescens har smala, krusiga lober 

med uppvikt kant, i princip iir allt Ovrigt P. 

canina speciellt om det finns tufsiga rhiziner. 

Det har f6ljaktligen varit Htt att gora en enkel 

bestdmningstabell. Men sedan Vitikainen 

(1981, 1985) uppmiirksammat arterna P. 

I

6 Gunnar Carlin GRAPHIS SCRIPTA 4 (19e2) 

rr''il 

!,///r,uW 

\l \\ /, \\'\. -t/ / 

\/ 

1---"t1 

Pdli3ewa cctrztl,tcl 

41"" Scln"Zter nL lq3z V b D orsjD 

ta(/ I (t, 

r/l ///, t,t/ t 

rr// ///, ,ttt. 

'4 ' //t rt 

:/e 

->,.fr 

RrJluffi*;\ff^A\,^., E V \qqs 

Figur 1. Schematiska bilder av P. canina-b61ar, efter hArt pressade exemplar. loberna 5r korta, 

breda, rundade och 6verlappande. Den ovre bilen visar typiska nedbdjda lobiindar och en antydd 

radiiir tillviixt, medan den undre bllen viixt mer oregelbundet och ger ett mer "krispigt' intryctq 

vilket fdrstiirks av att bllkanterna skadats blde i naturen och i herbariel 

/2/' 

"r,/(r* 

z


belu. H6r anses att b6len helt och hfillet best6r 

av lober (vilka genom forgreningar och 

inskiirningar delas upp i nya "lober" av ltigre 

ordning - hiirav dubbeltydigheten i begreppet 

lob), snarare 6n att lober skulle vara utskott 

fr6n en central bAl. 

Eftersom btlen, eller "hogsta ordningens" 

lober, ofta utbreder sig divergerat och forgrenat 

mot periferin uppster problemet hur 

man definierar lobernas bredd. Den praktiska 

lOsningen verkar bli det bredaste, av 

insk6rningar ej avbrutna, avstlndet mellan tv6 

lobkanter, som man kan se. Problemet kan 

tyckas konstruerat, men iir inte sA trivialt n6r 

man tvingas ttinka efter. Vilken ordning av 

insktirningar skall man stanna vid? (Jiimf6r 

med de kdnda fraktalgeometriska (definitions- 

) problemet: hur lAng 6r en kuststrticka! 

Kusten mellan t.ex. Hudiksvall och S6derhamn 

tir 5 mil om man anvdnder en 1 mil l8ng 

miitsticka, men siikert 50 eller kanske 500 mil 

om man anviinder en L cm lAng miitsticka, osv. 

ad inftnitum.) 

Andra anviinda begrepp som rydlig, knottig, 

smdgropig, matt, gltinsande 6r relativa och 

diirfor iinnu hopplosare att definiera och 

kriiver kontrasterande jiimforelsematerial f6r 

att kunna f6rst6s rdtt. Detta 5r ju egentligen 

inget konstigt. De flesta karakttirer tir i 

grunden relativa, och f6r alla sliikten gdller 

mer eller mindre att det iir omojligt att 

besttimma den f6rsta ensamma kollekten efter 

floran. Trots att det tar emot f5r man som 

vanligt sviilja att ingen bestiimningsnyckel kan 

erstitta erfarenhet och arbete. Jag har tilHtit 

mig att ganska ogenerat anvii-nda jdmfdrande 

uttryck utan att klart ange vad jag jiimfor med. 

T. ex. n6r det siigs att loberna hos P. 

membranacea [r ganska korta och breda 

(canina-lika), jiimfor jag forst6s med P. 

rufescens, P. praeteJcta och P. ponojensis. 

Miirk att beskrivningen av rhiziner alltid giiller 

friliggande sidana, aldrig rhiziner som tir 

fastvrxna vid underlaget. 

Peltigera canina 

B6len kan bli 6ver 20 cm stor. 

Peltigera canina-gmppen i Sverige 7 

l.oberna tillvtixer vanligen divergerande, 

de forgrenar sig upprepat och breder ut sig + 

6ver varandra ("taktegellagda"), se figur L. 

Loberna iir typiskt mycket breda 2-3(-4) cm, 

runda eller n6got tviirt avhuggna i 6nden, och 

6r riitt korta (har korta insktirningar). 

Lobkanterna 5r konvexa, "tslika", nedb6jda 

och + krusiga i ytteriinden. l,oberna kan i 

vissa fall vara lfrngstrSckta och ha uppetvikta, 

krusiga radiiira "parallella" sidor, men detta iir 

ovanligt och aldrig lika tydligt som hos vissa 

former av P. praetextata (figur 2). (Bilden av 

"P. canina" i Moberg-Holrnisen: Lavar, 

forestiiller ddrfor troligen P. praetextata). Det 

forekommer ocks6 b6lar med mindre, krusigare 

lober, som kan vara sv6ra att skilja fr6n 

P. rufescens (bed6m rhizinerna!). 

B6lytan tir matt sm6knottrig, gre till 

brun, hos tunna bfrlar {- bucklig efter [dermonstret 

p6 undersidan. Ytans matthbt iir 

niir man liirt kiinna denna karaktiir - vanligen 

ett mycket gott skiljetecken gentemot P. praetextata, 

som har mer fettgHnsande yta (dock 

lSngt ifrtn glansen hos t. ex. P. potydactyla). 

B8len iir vanligen tjockare och sl6tare (dvs. 

mindre bucklig) An hos P. membranacea, men 

kan vara lika tunn som hos denna, och undersidan 

kan ha ett gr8aktigt fdrgstick fr6n 

alglagret. Undersidan tir alltid Uus. 

Tomentum tdcker vanligen st6rre delen av 

b6len, men kan saknas i centrala delar och 

faller ibland av sjokvis. Det 6r tjockast mot 

lobiindarna, alltid kraftigt utvecklat, alltid 

kraftigare och mer omfattande dn hos P. 

praetextata. 

Adrorna iir ljusa, vita svagt ockragula, 

men blir ibland morkare i centrala delar, se 

figur 5. De bildar ett niitverk, som 6r tydligt 

och blir finare och finare fOrgrenat ut mot 

bilens kant. Adrorna tir distinkiupphojda och 

vll avgrdnsade fr6n bilens undre W, (men 

plattare 6n hos P. membranacea), tiimligen 

smala, de grovsta blir ca 0,5 mm breda. Ytan 

iir sldt (dvs hyferna 5r l6ngsg6ende och -i' 

sammanklibbade) eller ibland sammetsluddiga, 

liknande den hos P. membranacea. (Luddiga 

6dror tycks vara h6gre och skarpare avsatta 6n 

sliita Adror.) Mellanrummen mellan 6drorna 6r

8 Gunnar Carlin 

11,, ' ', , 

//- 

a 

'tz/t)/)>'.2 

-?/: ,2r- 

4/i 

;",{ 

\ t1 

rri 

?', Uii 

l'/l 

Pel{ige,ra pta.\ur{a}c\ L.'^do.[^l t{.lV. lqS3 

ttlt.Fu.-bo 

pe\{isen^ vu{esce"' U;fU"X'Y;l;1, 

Figur 2. P. praetextata. Raditirt utvecklad b6ltyp med avl6nga, 

overlappande lober med uppvikt, krusig kant. 

Figur 3. Smallobig b6l av P. rufescens med uppvikta, krusiga lober. 

GRAPHIS SCRIPTA 4 (19e2) 

hoptrlingda, ej markant

GRAPHTS SCRTPTA 4 (Leez) 

ljusa, alltid minst dubbelt si brett som de 

omkringliggande 6drorna, och ! rombformiga, 

ca 2-3 ggr si ltnga som breda, ibland iinnu 

mer avl6nga. Adror p6 fertila lobutskott lr, 

som hos alla Peltigera-arter, betydligt kraftigare. 

Rhizinerna, figur 6, dr m6ngformiga, sitter 

betydligt tdtare 6n hos P. praetextata och p. 

membranacea, ibland med kortare avstind tin 

0,5 ffiffi, ibland flera stycken i en tat rad, 

ibland som en kam av smA korta buskiga 

rhiziner mellan tv6 liingre. 

Rhizinerna 5r ungefiir jiimntjocka med Adran 

de sitter pe och tir vanligen kortare iin 5 mm. 

Friliggande rhiziner iir alltid vita. De varierar 

frin i lOst sammanh6llna uppsplittrade 

knippen av + parallella hyfer, till yvigt f6rgrenade, 

buskiga rhiziner med rikligt med 

mm-l6nga vinkelrdtt eller snett "fram6triktade" 

(mot marken riktade) sm6grenar. Det 

forekommer iiven membranacea-lika enkla, 

l6nga rhiziner, som tir ttickta av en sammetsliknande 

matta av vinkelratt utstaende korta 

hyfer. 

Rhizinetiickningen iir mycket olika pi 

olika delar av samma b8l, fr6n helt glest sittande 

till en kompakt matta av buskiga 

rhiziner, som inte liimnar b6lens undersida 

synlig. Olika Wper av rhiziner iir inte huflrx 

sammanblandade utan det finns gradienter 

fr6n den korta buskiga typen till den l6nga 

smala. 

Apothecierna iir morkbruna. 7 ffiffi, 

ungefiir cirkelrunda eller n6got bredare iin 

h6ga, pe I upprtittstiende lober. Vanliga. 

Ekologi och utbredning. P. canina vtixer 

bflde bland mossa pA sten och triidstammar 

och p6 marken i skuggiga skogar, men 6ven p6 

torrare stiillen som viigsklrningar (se Lindahl 

1953). Sk-TL. Utbredningens tyngdpunkt 

ligger norrut av herbariematerialet att d6ma. 

Det finns t ex f6 kollekter fr6n Bohusltin, och 

arten tycks vara ovanligare tin P. praetextata i 

Svealand och Gotaland. 

Sammanfattning. Peltigera canina iir en 

ttimligen enhetlig art, utmiirkt av breda lober, 

f6rekomst (6tm. htir och dar) av buskiga 

rhiziner och viil utvecklat tomentum. 

Peltigera canina-gntppen i Sveige 9 

Typiskt f6r P. canina 6r den oj6mna, 

fliickvisa forekomsten av tiitt sammanpackade 

korta, buskiga, "lurviga" rhiziner. Stdllen med 

glesa l6nga rhiziner och tydligt upphOjda 6dror 

blir mycket lika P. membranacea. Observera 

att rhizinernas utseende varierar mer inom 

individen zin mellan individerna. 

Peltigera kristinssonii 

B6len 6r ca 10 cffi, alltsA mindre 6n hos p. 

canina. 

Loberna tir t-L,s-Z cm breda med 

uppAtbojda (eller inrullade) kanter, liknar 

mycket P. rufescens. Lobkanterna biir ofta smi 

isidielika lober. Bfilytan iir striiv i kanten och 

bdr scabrosa-lika vflrtor och ett sparsamt, fint 

tomentum av uppriitta, korta hir. Mot centrum 

blir ytan blank, helt olik ytan hos p. 

rufescens och P. canina, och inte riktigt lika 

gltinsande som hos P. potydactyla. Bilen ir 

grfibrun - brun, ofta karaktiiristiskt gulaktig. 

Adrorna 6r m6rka, tamhgen plana och 

luddiga av uppriittst6ende hyfer. De bildar ett 

fint och tiitt niitverk av polydacUla .W. Mellanrummen 

iir ljusa och rundade. Adrorna 6r 

jtimforelsevis flatare och niitverket iir finare 6n 

hos P. canina och P. rufescens. 

Rhizinerna 6r m6rka - svarta, upp till 3- 

4 mm l6nga, tufsiga, buskiga eller enkla, de 

sitter viil 6tskilda. 

Ekologi och utbredning. P. kristinssonii 

vtixer i huvudsak ovan trtidgr6nsen. Jmt-TL. 

Utom Sverige tir arten k6nd fr6n Gr6nland, 

Island, Nordamerika, Sovjet, Norge, Turkiet, 

Alperna och Pyren6erna. 

P. kistinssonii kiinns bdst igen pl den i 

centrala delar blanka b6lytan, och de fina 

hSren i de yttre delarna. 

Denna beskrivning 5r extraherad ur Vitikainen 

(1985) och Kristinsson (1968, som p. 

occidentalis). Jag har sjtilv inte sett n6got 

svenskt material, endast ett fStal isliindska 

kollekter. Dessa stdmmer viil med ovanstiende 

beskrivning vad gtiller loberingens likhet med 

P. rufescens, b8lytans glans och det fina 

tomentumet. Diiremot iir inte alls undersidans 

karaktiirer lika klart skilda frAn p. rufescens, 

som antyds i beskrivningarna. Hos det


NW 

Ntrfi 

.$' tr'1,i.,*,- 

'{ i, //'7 

', 

il 

V v+^ 

Peffiget^". wrernb s am6L €A 

tt,rqJ* s to,\ t{. x' tct Sct 

o'[o t,.lcr'^'^'A c1 

Bat"". 

Vlt': 

/ 7t 

ffi \i 

ft 

,ry 

aa 

i3, 

w4 

'*, i4i 

.' i.:7:> 

\ v,t -\ 

ffiJ il,Tl 

I r, I ,,i,' l,r/! 

Figur 4. P. membranacea. Tunn, bredlobig caninc-lik b6l. 

i i /'t" 

,,rr' 

TA 

GRAPHIS SCzuPTA 4 (1992)

GRAPHTS SCRTPTA 4 (19e2) 

material jag sett tir iderntitet finmaskigt och 

tydligt i lobkanten, men det iir ljust, ej m6rkt 

som hos P. potydactyla. Innanfor kantzonerna 

iir 8drorna bredare, sammanflytande och delvis 

morka. Rhizinerna likn ar P. ntfescens. Av det 

lilla material jag undersOkt ?ir det svf,rt att fe 

en ktinsla for det karaktiiristiska hos p. 

I


Stna A r r, 

Pe t I i qe,,rc\ vn eM bravr r,tc e c^ 

Vatt{uist Vg. E' \qrq 

Ye\*lc(era ca!.r/lc\ 

v 

Lindc.hl Up\. \.U. l9tl9 

GRAPHIS SCRIPTA 4 (re92) 

fa 

) Pr:'r 

Fisur 5. Aderniit (utan rhiziner) hos Peltigera membranacea och P. canina. Adrorna ar vita. 

Ai"tna*, ai gf.tit. hos P. membranoiea och forhAllandet mellan ldrornas bredd och 

mellanrummenJ bredd Ar mindre. P. canina kan ha Adror och rhiziner som iir praktiskt taget 

identiska med P. membranacea, men Aderniitet er altid tatare' 

Observera att storleksskillnaden pA teckningarna inte betyder annat dn att en hogre ordning av 

lober iir avbildad hos P. membranacea.Tacif1r tvl tredjedelar f6r en direkt j6mf6relse!

GRAPHIS SCRTPTA 4 (ree?) 

6r tunnare och mindre utbrett, finns endast i 

lobernas yttersta delar och kan vara mycket 

otydligt. 

Adringen faller helt inom variationen , for 

P. rufescens. 

Rhiziner, figur 8. Typiska ponojenrurhiziner 

6r vita, viil Stslilda men ibland 

ttittsittande (flera per mm-), upp till 4 mm 

linga, tunna - 0,1-0,2 mm breda, och bestir 

av tiitt sammanklibbade liingsgiende hyfer. 

Dessutom fdrekommer, liksom hos P. 

rufescens, kamlika rader av tatt sittande korta 

rhiziner, buskiga-borstlika-uppsplittrade rhiziner, 

och till och med enstaka bruna rhiziner. 

Apothecier som hos P. canina och P. 

rufescens. Saknas ofta. Enligt Vitikainen 

(muntligt) tenderar apothecierna hos P. ponojensis 

att vara bredare, st6rre och ljusare tin 

hos P. rufescens. 

En sliit bAlyta med svagt tomentum korrelerar 

viil med f6rekomst av m6nga linga, 

tunna, vita rhiziner, vilket definierar P. ponojensis. 

Formodligen skiljer man P. rufescens 

och P. ponojensrs liittast genom de ovre belytornas 

klart olika karakt?irer. (Jtimfor ocksA 

P. kristinssonii, som 6r mycket svArare att 

skilja frln P. ponojensis). 

Ekologi (?) och utbredning som hos P. 

rufescenJ. De ekologiska preferenserna kommer 

sdkert att kunna formuleras med st6rre 

skhrpa, sedan nu arten uppmiirksammats av 

Vitikainen (1981). Sk-TL. 

Peltigera praetextata 

B6len kan bli ca 30 cm stor. 

Loberingen kan vara av tve riitt skilda 

slag, som kan f6rekomma bredvid varandra i 

naturen. Dels finns en typ vars lober iir 

lingstriickta, vdxer ut radi6rt fr6n b6lens mitt 

och vidgar sig gradvis mot b8lperiferin, se figur 

2. De iir i sin inre del ca 1 cm breda och nSgot 

bredare mot bfrlperiferin. Inskdrningarna mellan 

loberna Ar * radiiira och tydliga. Loberna 

iir ej tydligt Overlappande, ofta pressar intillliggande 

lober varandras sidor uppit sA att 

loberna blir riinnformiga. Kanterna iir n6got 

uppbojda och ofta mycket krusiga, men nedb6jda 

i den avrundade lobtinden. Den andra 

Peltigera canina-gntppen i Sverige 13 

typen av lobering liknar typisk P. canina, dvs 

loberna 6r breda och overlappande, har mer 

cirkuliir omkrets (-inte lAngsmala) och har f 

nedbojda kanter runt om. 

Bfilytan iir helt sliit till liiderartat sm[bucklig, 

jiimnt gr8tiirgad till kraftigt brun - det 

senare 6r typiskt fbr arten, ibland med 

obetydliga ljusare fliickar. B6len 5r vanligen sl 

tjock att undersidans fldermonster ej antyds 

genom bucklor pl ovansidan, men tunnare 

och buckliga bAlar f6rekommer. 

Tomentum varierar, iir alltid viilutvecklat 

men tiimligen tunt, finns alltid p6 lobernas 

yttersta 3-5 ffiffi, saknas vanligen centralt. 

Den tomentumfria ytan i centrala delar av 

bilen utgor vanligen en betydligt st6rre andel 

av b6len och iir mer gltinsande tin hos P. 

canina. 

Fyllidier 5r utmiirkande for arten och kan 

tiicka kvadratcentimeter-stora omrAden, men 

5r ofta begriinsade till vissa sprickor och 

kanter. Kantfyllidier b6rjar som smi diskreta 

korn, och utvecklas till t6tt sammantr6ngda, 

upprepat loberade ! korallika {511, med ca 0,L 

mm breda ! platta lober. Aven lobytans ryllidier 

(de som ej 6r uppenbart associerade till 

skador i bSlen) tycks b6rja som diskreta fia[ 

vilka utvecklas till platta korallika tiita 

"buskar". Ibland finns bara sm6 korn och ej 

ordentligt utvecklade inskurna, loberade qail. 

Ibland saknas fyllidier helt. Observera att 

m6nga (alla?) Peltigera-arter kan ha smt fialliga 

utskott pA lobkanten (men inte liings 

sprickor och pl b8lytan). F6rekomst av fyllidier 

iir siledes ej ett stikert kSnnetecken f6r 

P. praetextata. Det iir framf6r allt undersidan, 

speciellt rhizinerna som bdr de arttypiska 

kiinnetecknen som hos de flesta Peltigeraarterna 

men Sven den n6got gliinsande 

barken iir en god karaktiir. 

Adringen liknar i allt viisentli gt P. caninas. 

Kanske har P. praetextata ett tiitare f6rgrenat 

ndtverk iin P. canina. 

Rhizinerna, figur 11, iir ttimligen l6nga, 

3-5 ffiffi, kortare mot lobdnden, vita eller 

basalt svartbruna. De sitter 8tskilda (diskreta), 

med ca 1 mm mellanruffi, och iir ungefiir 

jiimntjocka med idrorna de kommer ifrin. De


I 

,,"- 

Figur 7. P. membranacea har mycket l6nga, 

smala, vita rhiziner tiickta med en sammetslik 

filt av vinkelrtitt utstflende hyfer. 


Figur 6. Rhizinerna hos P. canina dr vita och 

mycket m6ngformiga men typiskt korta, 

buskiga och tufsiga. Aven liingre rhiziner iir 

f6rgrenade eller borstlika. Ibland forekommer 

rhiziner av membranacea-typ. 

lmm 

I, 

Figur 8. Peltigera ponojensu rhiziner 6r vita 

och smala. De bestfir av mycket tiitt 

sammanfogade hyfer. Andra typer forekommer 

ocks6.


tycks bestA av en bunt tatt sammanklibbade 

hyfer, men iir ibland tomentosa i den basala 

delen eller har snett ut6triktade korta borst. 

Ibland iir knippet uppsplitsat i 2-3 nya buntar. 

Apothecier tir vanliga. De sitter pn 

uppriittst8ende lober, -6-7 mm breda och 

hOga. 

Ekologi och utbredning. Viixer pl tradstammar 

och mossiga stenar i lundar och lovskogar, 

mer sbllan i barrskogar. Foredrar 

oceaniskt klimat (Degelius 1935). 

Sk-LuL. Mest samlad i Svealand och 

GOtaland. 

Kollekten H. Smith, Jtl, Are, Enaforsholm 

19.VII.1965, UPS, 6r ett intressant mellanting 

mellan P. praetextata och P. canina. Den 

saknar fyllidier, har breda overlappande lober 

med uppiwikta kanter, tunt tomentum, praetacta-gllnsande 

yta och canina-borstiga 

rhiziner (Figur t1). 

Peltigera retifoveata 

B6len kan bli 20 cm stor. 

Loberna 61 2-4 cm breda, canina-lika, 

med nedvikta -f b6ljande, krusiga kanter. 

Inbernas sidor 6r ej nedvikta. B6lytan 6r 916, 

matt och sm6gropig, i st6rre luppf6rstoring 

sm6fl5ckig - krakelerad, (delvis med sm6 ljusa 

fltickar, "pseudocyphell.t"), och enligt Vitikainen 

(1985) vanligen gronare 5n hos P. 

canina. 

Tomentum 5r fint och tunt, tticker en stor 

del av b6len, men avtar fltickvis mot b6lcentrum. 

Adrornu iir ljusa, nigot ockragula, riitt 

otydligt avgr6nsade fr6n den vita bAlundersidan. 

Adrornas och mellanrummens storlek blir 

dtirf6r oklar, men bidas bredd iir av storleksordningen 

1 mrn. Mellanrummen tir + 

runda-ovala, upp till dubbelt s6 l6nga som 

breda, de ser ut som gropar (till skillnad mot 

P. canina, P. membranacea och P. praetextata, 

vars 6dror ser ut att ligga ovanpA en sliit yta, 

och vars mellanrum iir romboida-kantiga). 

Undersidans hyfer, b6de Adror och mellanrum, 

bildar en lucker filtmatta. 

Rhizinerna, figur 10, tir morka bruna 

och sitter viil Atskilda (diskreta) med i stor- 

Peltigera canina-gruppen i Sveige 15 

leksordningen 3 mm mellanruffi, men tiitare 

mot lobiinden. De 6r ofta 0,5 mm breda, ej 

siillan cirka 1 mm breda och 3-5 mm lfinga. 

De best6r av glest sammanfogade liingsglende 

hyfer och kring dessa en lucker filt av korta 

utstAende trfldar. Undersidans utseende lr 

mycket karaktiiristiskt med mdrka fitskilda 

rhiziner pA en ljus botten av otydliga 6dror. 

Apothecier ej sedda i denna studie. 

Beskrivningen bygger i stort sett p[ 

dubletter av typkollekten (Finland, Kuusamo), 

varf6r den inte behOver vara helt representativ 

for arten. Beskrivningen sttlmmer inte i alla 

delar med Vitikainens (1985), vilken bygger pi 

ett mycket st6rre material. 

Ekologi och utbredning. P. retifuveata 

viixer enligt Vitikainen (1985) pi jord och 

bland mossa i boreala skogar p& nigot sura 

jordarter. Endast ett fynd 6r giort i Sverige 

(Uppsala, 186L), men fler borde kunna g6ras i 

norra Sverige. Arten finns annars i Finland, 

Sibirien, Canada och USA (Washington och 

Alaska). 

P. retifuveata inneh8ller lavsubstanser 

(tenuiorin, metylgyroforat, zeorin och dolichorrizin) 

i motsats till de Ovriga behandlade 

arterna (Vitikainen 1985). 

Peltigera rufescens 

B6len iir vanligen ca L0 cm eller mindre. 

l,oberna utg6r i princip radiiirt frin b6lens 

centrum, men detta iir aldrig speciellt tydligt 

ens hos v6lutvecklade, fritt vdxande b8lar 

(figur 3). Loberna tir (upprepat) f6rgrenade, 

giirna med sA brett divergerande (sekundiira) 

lober att angrdnsande lober till stor del overlappar 

varandra. l.oberna 6r ca 0,5-1 cm 

breda, vanligen 3-5 cm l6nga. Typiskt f6r P. 

rufescens Ar den mycket krusiga, "for stora" 

kanten, vilken alltid tir upp8tbojd. Ej sAllan 

med ungetiir 0,1-0,3 mm stora "regenerationslober" 

i kanten. l-oberna ser i kanten 

tjockare ut hos P. ntfescens iin hos P. canina. 

Enligt Lindahl (1953) iir detta en synvilla. 

Tomentum tir kraftigt och tatt, men 

tunnar ut mot centrum diir det kan saknas. 

Ibland finns en kraftig pruina. Bfilytan iir 

matt, ofta fint knottrig (grovre iin hos P.

L6 Gunnar Carlin 

Lmm 

Figur 9, Storre rhiziner hos P. rufescens 5r 

morka och best6r av + l6st sammanfogade 

hyfer. Rhizinerna sitter ofta tiitt tillsammans i 

rader. 

I'mm 

GRAPHTS SCRTPTA 4 (reez) 

Figur 10. Rhizinerna hos P. retifuveata 

m6rka och tdckta av en lucker filt 

utstiende tr6dar. 

Smith, Jtl, Are 1965 

(se text) 

Figur 11. P. praetevata har vita rhiziner, som i huvudsak best6r av smala, 

viil sammanh6llna buntar av hyfer. 

iir 

av


praetextata, men forst6s sldtare iin hos P. 

scabrosa). Fiirgen iir ljusgr6, brungrA eller 

kraftigt brun. 

Adrorna 5r ljusa, vita eller nAgot gula 

eller mdrka, speciellt mot b6lens centrum. De 

iir upphojda men vanligen rtitt platta - stillan 

lika hoga som breda inte alltid speciellt 

skarpt avgriinsade och inte alltid tydliga iinda 

ut i b6lens periferi. Ytan tir tiimligen sliit. 

Mellanrummen mellan idrorna iir ljusa, ofta 

mycket l6ngstrdckta och ungefiir lika breda till 

dubbelt s6 breda som tdrorna, dvs ca 0,5- L 

mm. I centrala delar blir mellanrummen 

stOrre, bredare och rundare. Skillnaden i 

Adring mellan P. canina och P. ntfescens dr 

inte alls sA tydlig som det ibland gors giillande. 

T. ex. rhizinernas frirg och form iir biittre skiljekaraktdrer 

5n 8dringen. 

Rhizinerna, figur 9, 6r m6rka (i alla fall 

de liingre) eller har m6rk ktirna. Ofta finns i 

yttre delen av bilen mycket typiska l6nga 

kamlika rader av uppstickande vita

Boganmeldelse 

Introduktion til svampe 

Dissing, H., Hansen, L., Olson L. & Sochting, 

U. 1992: Introduldion til svampe. 3. udgave. 

VIII + 1S9 pp. Nucleus. ISBN 87 87661 67 5. 

120 DKK. 

Bogen giver en elementrer indfOring i 

svampenes systematik og biologi. Der er 

generelle indledende afsnit om bl. a. livscykler, 

navngivning og ernethg, samt et afsluttende 

kapitel om fflogenetiske betragtninger. Bogen 

er ellers systematisk opbygget over eksempler 

pt arter fra hver orden og med indskudte 

biologiske afsnit. 

Bogen indeholder kun fi og smi afsnit om 

licheniserede svampe, og disse er for korte til 

Exkursion 

En lavexkursion arrangeras pi Visings6 i 

Smiland den 26 27 september 1992. Hiir 

finns bl. a. Caloplaca biatoina, C. lucifuga, C. 

obscurella, Candelariella retlexa, Cyphelium 

notarisii, Opegrapha sorediifera, Physconia 

at tjene som en fyldestggrende introduktion til 

laverne. L&serne kan blive forvirret af at 

bladlaver siges at have rhiziner, men senere 

omtales Hypogmnia physodes som en bladlav. 

Calicium regnes til Plectascales, og der gives 

ingen eksempler pi licheniserede pyrenomyceter. 

En omtale af. Vemtcaria maura og 

strandzonering havde vreret oplagt. 

Eftersom laver er svampe kan man f& et 

storre udbytte af lavstudierne, n6r man har et 

generelt kendskab til svampene, og bogen er 

velegnet til at bibringe lreseren et s6dant 

kendskab. 

Vagn Alstrup 

grisea och Ramalina obtusata. Intresserade 

kan kontakta Goran Thor, Botaniska institutionen, 

Stockholms Universitet, 5-106 91 

Stockholm, Sverige.

The 9th meeting of the Nordic Lichen Sociefy, Sweden L99I 

cOnaN THoR 

Thor, G. L992: The 9th meeting of the Nordic Lichen Society, Sweden 1991. 

Graphis scripta 4: 19-29. Stockholm. ISSN 090L-7593. 

A fiye-day meeting of 55 lichenologists from the Nordic countries was held 

in Asele l-appmark, Srurniis in August 199L. The meeting included 

excursions, a one day symposium and the general meeting of the Nordic 

Lichen Society. A list of lichens and lichenicolous fungi recorded is presented 

here. A total of 558 species were observed of which 3 (Lecidea tepraioides, 

Leucocarpia biatorelir . Zwackhiomyces berengeianus) are new io Sweden 

and 9L are new to Asele I-appmark. 

Goran Thor, Botaniska Institutionen, Stockholms (Iniversitet, 5-106 gI 

Stockholm, Sweden. 

From August 5th to 9th, IggI the Nordic 

Lichen Society held its 9th meeting in the area 

surrounding Saxnds (270 km NW of UmeA 

and 70 km NW of Vilhelmina) in Asele 

I-appmark, northern Sweden. 55 lichenologists 

from the Nordic countries attended the 

meeting, which included four day-long 

excursions (5th, 6th, Sth and 9th), a one-day 

symposium (7th) and on the evening of the 

7th, a general meeting where the firsi charter 

of the association was adopted and the first 

council was elected. The symposium had the 

theme: "Nordic lichens, current taxonomical 

and ecological research" and 11 speakers 

contributed (Teuvo Ahti, Per Anders Esseen, 

Frc Froberg, Eric Steen Hansen, per Magnus 

Jorgensen, H6rdur Kristinsson, Jan-Eric 

Mattsson, Henrik Skult, Ulrik SOchting, Einar 

Timdal and Orvo Vitikainen). During the 

evenings it was possible to work with and to 

determine the collected material. 

visited localities are presented after the species 

list with short notes on the habitats. Each 

locality is identified by a number. Geologically 

the area is dominated by acid bedrock and 

moraine with the exception of local ity 4 with 

ultrabasic peridotite, locality 5 with dolomite 

and locality 8 with calcareous rocks. The 

visited habitats included one alpine heath 

(locality 1), one Pinus sylvestris forest (locality 

3), one Picea abies forest (locality 6), nvo talus 

slopes with deciduous trees (locality 4, 9) and 

four mixed coniferous forests with stands of 

deciduous trees (Betula, Populus temula, 

Salix) (locality 2, 5, 7, 8).Most of the forests 

at lower altitudes in the area have been 

influenced by forestry and there are only small 

remnants of natural forest left (e. g. tocatity 6). 

The altitude of the visited localities varies 

between 390 and 960 m. 

Earlier investigations 

Visited habitats 

The visited area is one of the least explored 

rhe area su'ounding saxniis is covered with iliul|l":yffi.Ijll#tilf"rH"li:1"::t:fl 

forests.in the valley bottoms and has alpine lichen flora auailuble. Lists of species (almost 

vegetation on the surrounding mountains. The e:

20 Goran Thor 

been published from locality 1 (Degelius 

1932), locality 2 (Ahlner 1938, Degelius 

1954), locality 3 (Ahlner 1938, Degelius 

1932), and in Santesson (1984) Endocarpon 

psorodeum is reported as new to Sweden from 

locality 5. All of the previously published 

lichen species were not found on the 

excursions. The most interesting species 

reported earlier is perhaps Collema curtispontm 

from locality 2 (Degelius 1954). We 

did not rediscover this species. Some interesting 

vascular plants have been reported 

from locality 4 (Rune 1953) and locality 5 

(Stenberg & Westerberg 1983). 

The species list 

Seven localities were visited during the 

meeting (see visited localities). Material 

collected from two localities on the 

reconnaissance excursions in 1988 and 1990 

have been included in the list. The participants 

of the excursions were in 1988 Per Anders 

Esseen, Jan-Eric Mattsson, Roland Moberg 

and Goran Thor, in 1990 Per Anders Esseen, 

Per-Olof Martinsson and Goran Thor. A 

total of 558 ta,xa were collected or observed of 

which 2 are new to Sweden and 92 are new to 

Asele Lappmark. The species list includes 

both collected species (always marked with the 

acronym of the collector) and species only 

observed in the field (lacking acronym). Some 

field determinations which should be 

confirmed with collected material - have been 

excluded from the species list. For some 

species short notes on the chemistry or the 

habitat are given. In addition to lichens, the 

family Mycocaliciaceae and some lichenicolous 

fungi have been included. I have tried to use 

up-to-date names. The names used by 

Santesson (1984) are given as synonyms when 

they differ from the names in the list. 

Contributions to the species list were 

made by several persons during the 

excursions. The persons listed below have 

later contributed with information about their 

collections. The herbarium in which the 

collections are housed is specified after the 

acronyms of the names: Teuvo Ahti (TA) (H), 


Per Anders Esseen (PAE) (UME), TonY 

Foucard (TF) (private herbariuil, two 

collections in UPS), Hans-Erik Gustavsson 

(HEG) (private herbarium), Veli Haikonen 

(VH) (H), Ingemar Herber (IH) (private 

herbarium), H6kon Holien (HH) (private 

herbarium, will be deposited in TRH), Sofie 

Isbrand (SI) (private herbarium), Per Magnus 

Jprgensen (PMJ) (BG), Erik I-arsen (EL) 

(private herbarit*), Anders Nordin (AN) 

(private herbariu*), Rolf Santesson (RS) 

(UPS), Rikard Sundin (RSU) (S), Ulrik 

Sochting (US) (C), G6ran Thor (GT) (S), 

Orvo Vitikainen (OU (H), Heino V6nskii 

(H\|) (collections will be deposited in "I{), 

Mats Wedin (MUl) (UPS), Ake Agren (AA) 

(private herbarium). 

* = New to Asele L^appmark. 

* * = New to Sweden. 

Abrothallus parmeliarum (A. cetrariae): 5 

(MW, conf. RS). -On Platismatia glauca. 

Acarospora badiofusca: 1 (TF) 

A. glaucocarpa: 5 (GT),7 (TF) 

A. sinopica: I 

A. veronensis:4 (TF) 

*Agonimia tristicula:5 (GT), 9 (GT) 

Alectoia nigicans: 1 (RSU) 

A. ochroleuca: 1 (RSU) 

A. sarmentosa: 2, 5 (TA), 6,7 

Allantoparmelia (Parmelia) alpicola: t 

Amygdalaia (Huilia) panaeola: 12 (TF) 

*Anisomeridium nyssaegenum: 2 (GT), 5 

(GT). -In shaded and wet habitats on base 

of Alnus incana (2) and on base of- Picea 

abies (5). 

*Anzina carneonivea: I (TA), 2 (GT)' 3 

(RSU), 5,6 (GT) 

Arctomia delicatula: 1 (HH), 2 (GTr det. 

PMJ), 5 (GT). -Not reported from Al Uy 

Santesson 1984 but by Foucard L990. 

Arthonia dispersai 5 (EL) 

A. mediella: 2 (TF), 5 (TF)'7 (TF) 

*A. molendoi:4 (RS). -On Xanthoria elegans. 

A. radiata:2 (GT), 5 (GI) 

*A. vinosa: 2 (GT). -On base of Picea abies 

in dens e Picea abies forest.

GRAPHTS SCRIPTA 4 (1992\ 

Arthopyrenia lapponina: 2 (TF), 5 (HEG, TF), 

6 (TF), 7 (TF), 9 (cT, det. TF) 

A. puncttformis: 7 (TF) 

*A. salicr: 5 (GT,det. TF) 

Anhrorhaphb citinella: I,2,7 (TF) 

Aspicilia aquatica: ? (TF) 

*4. mashiginensis:2 (HH, TA, GT) 

A. melanaspis;2 (HH), 3 

A. montana: 1 (TF) 

A. myinii: I (HH, RS) 

A. supertegens:2 (HH, GT, TA, det. RS) 

Bachmanniomyces uncialicola-. 2 (MW, conf. 

RS). -On Cladonia amaurocraea. 

Bacidia baglieuoana: 5 (GT) 

B. (Catillaia) globulosa: 2 (TF, GT), 4 (TF), 

5 (HEG,TF) 

B. igniarii: 6 (TF) 

B. subincomptat 2 (qD, 5 (GT, HH), 7 

B. trachona: 5 (TF, AA; 

Bacidina inundata: 7 GF) 

Baeomyces carneus: I 

B. placophyllus: 1 (TA), 2 (SI), 3 (SI) 

B. roseus: 1, 6 (Ou 

B. rufus coll.: I (TA),2,3, 4, 5, 6,7 

Bellemerea (Aspicilia) alpina: 1 (TF, AA; 

B. (Aspicilia) cinereorufescens: 2,3, 6,7 

B. (Aspicilia) diamarta: 1 (EL) 

B. (Aspicilia) subsorediza: L,2 (HH), 6 

Belonia russula: 2 (GT), 3 (Gf), 5 (TF), 9 

(GT). -Not reported from At- Uy Santesson 

1984 but by Foucard 1990. 

Biatora carneoalbida (Bacidia sphaeroides): L, 

2,5 (HH, IH, GT) 

B. (Lecidea) cuprea: 5 (TA) 

*8. (Lecidea) ffiorescensi 2 (GT, HH), 5 

(GT, TA), 6, 7 (TF).-Argopsin (TLC, 

collections by GT). 

*8. (Lecidea) epixanthoidiza: 2 (GT), 3 (GT), 

5 (TF), 6 (GT). -Thallus C+ red 

(gyrophoric acid). 

B. (Lecidea) helvola: 1 (HH, TF), Z (TF), 5 

oF),6 (OV, TA),7 (TF) 

B. (Bacidia) pallens: 2 (HH), 4, 5 (TF), 6 

(RSU, cT) 

B. (Lecidea) vernalis: 2 (IH, RSU), 5 (TA, 

GT) 

Bryonora curyescens; 1 (GT) 

NLF meeting 1991 2L 

*Bryonora pruinosai 1 (AN). -For a morphological 

description, see Holtan-Hartwig 

1991, 

Bryoria capillaris: 2, 5,7 

B. furcellata: 1 (TF) 

B. fuscescens:2r 51 617 

B. simplicior: 3, 4, 5, 6,7 

Buellia discifurmis: 2,3,6 (TF), 9 (GT) 

B. (Diplotomma) epipolia: 5 

B. (Diplotomma) nivalis: 2,3 (GT) 

B. punctata: 2 (TF), 5 (TF) 

*Calicium adaequatum: 2 (HH, MW, conf. L. 

Tibell). -Both on Alnus incana and 

Populus tremula. 

*C. glaucellum: 5,6,'7 

C. salicinum: 2, 5,'l 

C. trabinellum:2,6 

C. viridez 2,5,6 

*Caloplaca ammiospila: l, 4 (RS), 5 (US) 

C. approximata: 3 (US) 

C. arenaia: a (GT) 

*C. borealis: 1 (AN, OV, US), 2 (HH, TA), 5 

(GT, HH, OV, US),6 

C. ceina: 1 (HH, TA, US), 2, 4,5 (US), 6, 9 

(GT) 

C. chloina: 5 (US) 

*C. (Leproplaca) chrysodeta: 5 (RSU, GT) 

C. citina: 4 (TF), 5 (US) 

*C. tlavorubescens:2 (GT, US), 5 (TA). 

C. flavovirescens: 5 (US) 

C. herbi"della: 5 (EL) 

C. holocarpa: 1 (TF), 4 (TF, US), 5 

C. jungermanniae: 4, 5 (TA), 8 (GT, conf. 

US) 

*C. magni-ftlii:1 (US) 

C. nivalis: 1 (US) 

C. obliterans: L,2 (GT, US), 3 (OV), 4 (US), 

s (cr) 

C. sacicola:5 (US), 9 (GT) 

C. sinapisperma: 5 (TA, US) 

*C. sorocafpa: 1 (HH, US),5 (US) 

C. tiroliensis: 2 (GT, conf. US), 3 (GT), 9 

(GT, det. US) 

C. vitellinula: 2, 3 (GT). -Not reported from 

Al by Santesson '1984 but by Foucard 

199A. 

Candelaiella aurella: 4 

C. coralliza: L

22 Goran Thor 

C. kuusamoiinsrs: 3 (RSU) 

C. placodizans:4 (TF) 

C. vitellina: | (TF), 2 (TF), 3 (TF) 

C. xanthostigma:3 

Carbonea vitellinaia: 4 (TA). -On Candelaiella 

aurella. 

Catapyrenium lachneum: 5 (GT) 

*Catillaia nigroclavata: 2 (SI, conf. V. 

Alstrup) 

Catinaria atropufpurea: 5 (HH) 

C atolechia w ahlenbergii: L 

Cetraia chlorophylla: 2 (RSU, GT), 3 (OV), 5 

C. commixta:1, 2 (SI), 3 (GT) 

C. cucullata: t 

C. delbei:3 (RSU) 

C. eicetorum: 1,3 (OV), 4 

C. hepatizon: 1, 3 

C. islandica ssp. crupiformis: 4 

C. islandica ssp. ts/a ndica: L,2, 4 

C. junipeina: 1 (RSU, GT, OV, TA), 3 

(PAE), 4 (H\) 

C. nivahs: L, 4 

C. odontella (Coelocaulon odontellum): 2,3 

C. pinasti: I,2,3, 4, 5, 6 

C. sepincola: 1 (SI), 5,6 

Chaenotheca brachypolo. (Conioqbe sulphurea):2 

(GT), 5, 6 (AA) 

C. brunneola: 2 (TF) 

C. chlorella (C. carthusiae)z 2 

C. chrysocephala: 2 (GT), 5, 6,7 GF) 

C. (Conioqbe) fu(uracea:2, 5 (MW, conf. L. 

Tibell, GT),6 

C. (Conioqbe) gracillima: 2 (MW, conf. L. 

Tibell), 5 (MW, conf. L. Tibell) 

C. stemonea: 5 (MW, conf. L. Tibell, GT), 6 

C. subrosci.da: 2 (MW, conf. L. Tibell), 5 

(HH),6 (RSU, GT) 

C. tichialis:2 (GT), 5 (RSU), 6 (GT), 9 (GT) 

C. ryloxena:2 (HH) 

Chaenothecopsis epithallina: 5 (MW, conf. L. 

Tibell) 

C. nana: 2 (MW, conf. L. Tibell), 5 (RSU)' 6 

C. viridialba: 2 (MW, conf. L. Tibell), 5 (MW, 

conf. L. Tibell), 6 (MW,conf. L. Tibell) 

*Chrysothrix chloina: 2 (GT) 

Cladonia acuminata: 2 (HH, TA), 3, 4 (TA), 

s (rA) 

C. amaurocraea: lr 21 3, 4, 5 

GRAPHTS SCRIPTA 4 (19e2) 

C. arbuscula: t,2 (TA), 3,4,5 (TA), 6 

C. bacillifurmis;2,3,5 

C. bellidiflora: t,2,3, 4 

C. borealis: 1, 2 GA), 3, 5,6 

C. botrytes: 5,6 

C. cariosa: 2 (TA), 4 (TA), 7 (TF) 

C. carneola: 1,2, 5, 6,7 

C. cenotea: 2,3, 5, 6 

C. cervicornis ssp. cet?icornis: 1 (TA) 

C. ceruicornis ssp. verticillata: 2,6 

C. chlorophaea: 1 (s. str.) OA), 2 (coll.) (TA), 

5 

C. cocctfera s. str.: 1 (TA), 2 GA),4 

C. coniocraea: L, 5 

C. cornuta: 2,3, 5, 6 

C. crispata: 1 (TA), 2,5,6 

C. cyanipes: L,5 

c. decorticata:2 (TA), 3, 4 (HH) 

C. deformts: 1 (TA), 2 (TA), 3,4,5 

C. digitata: L,5 

C. ecmocyna: 1 (TA),7 (Ou 

C. fimbriata:2 (TA), 5 (TA) 

C. furcata: t,3, 4, 5 

C. gracilrs ssp. elongata: l,-4,5 

C. gracilrs ssp. gracilis:4 (TA) 

C. gracilrs ssp. turbinata: 2, 5,6 

*C. grayi: 1 (TA),2 (TA) 

*C. luteoalba: 1 (TA), 3 

C. macrophyllaz 1,2, 6 

C. macrophyllodes: 1 (TA) 

C. marima: L (TA) 

*C. metacorallifera: L,2,3,9 (GT) 

C. mitis: 1 (TA), 2 (TA), 3, 4,5,6 

C. ochrochlora:2 (TA), 3, 4, 5 (TA), 6 (TA) 

C. phyllophora: t,2,3, 5, 6 

C. pleurota: 1,2 (TA), 3,4,5 (TA), 6 

C. pocillum: 4,5, 8 (GT) 

C. pyxidata: 1 (TA), 2,3, 4, 5, 6,7 

C. rangiferina: 1,2,3, 4, 5, 6 

C. squamosa: t,2,3, 5 (TA) 

C. stellaru: L (TA), 2, 5 

*C. strep,si/rs: 1 (TA) 

C. sticta v. stricta: 1,2 

C. stygia: I,2 (TA), 4, 5 (TA) 

C. subulata:7 

C. sulphuina: 2,3, 5, 6,7 

C. symphycarpa: 2 (GT), 4,5 (TA) 

C. tutgida:2,3,41 5 (GT)


C, uncialls: L, 2r 3, 4 

Clauzadea (.ecidea) monticola: 5 (TF) 

*Cliostomum co,rugatum (Catillaria graniformis): 

5 (HH). -On Sorbus aucuparia 

close to rock-face. 

Coelocaulon aculeatum: L 

C. muricatum: 1 (AA) 

Collema cristatum: 5 (GT) 

C. flaccidum: 5, 9 (GT, conf. G. Degelius) 

C. furfuraceum:2 (PAE, RSU) 

C. glebulentum: 2 (AN), 5, 9 (GT, conf. G. 

Degelius) 

C. nigrescensi 5 (PAE, HH) 

C. occultatum: 2 (GT),5 (GT), 7 

C. tuniforme:2 

*Conotrema populorum: 2 

Coriscium viride: 5 (GT) 

*Cornutispora lichenicola: 6 (RS). -On Cetraia 

sepincola. 

*Cybebe (Coniocybe) gracilenta: 2, 5 (HH) 

*Cyphelium inquinans: 5 (HH, MW, conf. L. 

Tibell), 6 (MW, conf. L. Tibell) 

C. karelicum: 6 (GT, HH, RSU, MW, conf. L. 

Tibell) 

Cystocoleus ebeneus: 2 (GT), 5 

Dactylospora athallina: 5 (EL). -On 

Baeomyces rufus. 

*D. deminuta: 2 (GT, oo Lopadium pezboideum),s 

(RS, on Biatora vernalis; GT, on 

Gyalecta peziza). 

Dermatocarpon arnoldianum: 4 (IH, det. RS) 

D. bachmannii: 4 (IH, RS, GT, all det. RS).- 

First published from Scandinavia and this 

locality in Moberg (1986). 

*D. luri.dum (D. weberi): 5 (along the stream) 

(VH) 

D. miniatum:5 

*Dimerella pineti: 5 

*Diploschistes glpsaceus: 1,5 (RSU) 

*D. muscorum: 1 (TF),2,3 (SI) 

D. scntposusi 1 (TF), 3, 5 (TF) , 6,7 (TF) 

Endocafpon psorodeum:4 (HH, GT) 

Enterographa (Opegrapha) zonata: Z (GT) 

Ephebe hispidula:2 

Epilichen scabrosus: 1 (HEG). -On Baeomyces 

placophyllus, 

Euopsis (furenopsis) pulvinata: 3, 4 

NLF meeting 1991 23 

+ Everniicola fletispora: 5 (RS, RSU, TA, det. 

RS). -On Nephroma arcticum. 

Farnoldia (Melanolecia) jurana: 5 (SI, conf. V. 

Alstrup, EL) 

*F. (Melanolecia) micropsrs: 5 tAAt 

Foraminella (Parmeliopsis) ambigua: 1, 2, 3, 

5,6,7 

F. (Parmeliopsis) hyperopta: 1,2,3, 4, 5, 6,7 

*Graphium aphtosae: 5 (RS). -On Peltigera 

aphthosa. 

*Gyalecta 

erythrozona: 3 (GT) 

*G. 

fiesii: 5 

G. geoica: t,5 (RSU, GT) 

G. jenensrs: 5 (RSU, GT) 

G. peziza: 5 (GT) 

*G. ulmi: 5 (GT). -In South and Central 

Sweden this species is found on old trunks 

of broad-leaved deciduous trees. On the 

very few localities in northern Sweden 

(earlier known from Jmt) it is found on 

shaded, wet rocks. On the mountain 

Kalvberget it was rather commonly found 

on shaded and wet dolomite rocks. This is 

the hitherto northernmost locality for this 

species in Sweden. Totally as many as five 

Gyalecta species was found on Kalvberget. 

*Gyalidea 

fiuei: 5 (along the stream) OF) 

(uPS) 

*Hobsonia santessonii: 2 (HEG, det. RS). 

On Peltigera scabrosa. 

Hymenelia (Aspicilia) lacustris: 2,3 (SI) 

Hypocenomyce leucococca: 2 (CtT), 5 (TA), 6 

(RSU) 

H. scalaris: 2r 31 7 

*H. sorophora: 2 (GT), 3, 5 (GT), 6 (GT). - 

Alectorialic acid (TLC). This species is 

reported as "Sterile sp." in Karstrdm & 

Thor 199t. 

Hypogmnia austerodes: 2,6 OA) 

H. bitteri:2 (GT), 5, 6 (MW, RSU) 

H. intestinifurmis: 1 (OV, RSU), 2, 3 (GT, 

oV),6 

H. oroarctica: I (OV). -On rock outcrop. 

H. physodes: 1,2, 3 ESU, GT), 4 (HEG, c. 

ap.), 5, 6,7,9 (GT, c. ap.) 

H, tubulosa:2,3 (OV), 5 (GT), 7 

H. vittata:2 

Icmadophila eicetontm: 1,2,3,5 (TA), 6,7

24 Goran Thor 

Imshaugia (Parmeliopsis) aleuitesz 3 (OD 

Ionaspis arctica: 3 (RS) 

I. chrysophana (1. suaveolens): 3 (PMJ) 

I. qanocarpa: 5 

I. heteromorpha: 5 (GT) 

I. odora: 3 (AN) 

*1. rhodopis:5 0H) 

Iapewia subauiferaz 2 (GT), 5 (GT, TA), 6 

(RSU),7 

I. (-ecidea) tomo1nsb: 5,6 (RSU) 

Koerbeiella wimmeiana (Aspicilia leucophyma): 

2 (RS, TF) 

Lecania rabenhorstii: 5 (TF) 

L. suavrs: 4 (AN,TF),5 (RS) 

*Lecanora albella (L. pallida)z 5 (SI, conf. V. 

Alstrup, TF) 

L. albescens:4 (TF) 

L. anopta: 5 (TF) 

L. argopholis:2,3 (HV, TF, VH), 4 (HV) 

L. boligera: | (TF), 5 (TA), 6 (SI, TA), 7 (TF) 

L. cadubriae:3 (HH), 6 (TF) 

L. cateileaz 2,5 (HH, OV, TA), 6 (OV) 

L. cenisia: t GF), 3,4 (RS), 7 (TF) 

L. chlarotera: 7 (TF) 

L. chloroleprosa:3 (HH, HV, RSU), 6 

L. chlorophaeodes; 2 (HV) 

L. circumborealis (L. coilocarpa): 2, 3 (IH, 

oV),5 (OV),6 

L. dispersa:2 (GT), 3 (GT), 5 (TA) 

L. epibryon; 5 (GT) 

*L. expallens:5 (HH, TA). -The collection by 

HH on Sorbus aucuparia. 

L. frustulosa s. str.: 2 (Hu 

L. fuscescensi 1, 2 (SI) , 5, 6 

L. hypoptoidesz 5 (TF) 

L. inticata: 1 (HV), 2 (HV) ,3, 4 (HV), 6 

L. muralis: 4 (HV, GT), 5 (TF) 

L. phaeostigma: 2 (TF) 

L. piniperda: 6 (TF) 

L. polytropa: I (HV), 2 (HV, TF)' 3 (TF), 5 

(GT),6, 7 (TF) 

L. pulicaris:2 (TF), 4, (TF), 5 (TF), 7 (TF) 

L. reagens:2 (GT, HV) 

L. septentionalls: 5 (Ou 

L. subintricata: 2 (TF), 5 (HV),6 (HV) 

L. rymmictaz t,2 (GT), 5 (GT, HV), 6,7 (TF) 

Lecanora sp. (L. chlorophaeodes agg.): 2 

(HV) 


Lecidea albohyalina: 2 (HH), 5 (TF). -On 

locality 2 collected on Sorbus aucupaia. 

L. auiculata: 1 (TF) 

L. caesioatra: 1,2 (TA), 3 (AA) 

L. cinnabarina: 2 (GT), 5 (HH, MW, RSU, 

GT),6 (GT) 

L. conferenda: a (EL) 

L. confluens:1 (SL conf. V. AlstruP), 7 (TF) 

L. cuprea: I (TF) 

L. erythrophaea: 2 (GT) 

L. fuscoatra: 1 (EL, TF), 5 (TF) 

L. lapicida (including L. lactea): I (EL' TF) 

**L. lepraioides:5. (fA, det. T. TOnsberg) 

L. leucophaea: 1 (AA) 

L. limosa: (EL) 

L. lithophila: 1 (HH), 2 (EL) 5 (SI, conf. V. 

Alstrup). -On locality 1 it was collected 

on siliceous rocks. 

*L. margaritella:6 

L. paupercula: 1 (EL) 

L. pullata: 1 (RSU, TA), 2 (GT, TA), 3, 5 

(GT,TA),6(TA),7(TF) 

L. pycnocarpa: 1 (TF) 

L. turgidula: 5,6 

Lecidella euphoregr"l (TF), 4 (TF) 

L. inamoena: 1 (AA) 

L. stigmatea: 4 

Lecidoma demissum (Lecidea demissa): 1 

(RSU),3 (TA) 

Lempholemma potyanthes (L. myriococcum): 

5 (TA, det. PMJ) 

*Lepraia incana coll: 2, 5 (TA) 

*L. lobiftcans: 5 (GT). Thallus with a 

distinct medulla. Two collections, one on 

base of. Picea abies, one on mossy, vertical 

shaded rock. Both have atranorin, zeorin, 

divaricatic and stictic acid complex (3 

spots) (TLC). 

L. neglectai 1, 2 (TF), 3 (TA),4,6 (TA) 

Leproloma membranacea (Lepraria membranacea): 

9 (GT). -Pannaric acid and roccellic 

acid (TLC). 

L. vouauxii (Lepraia arctica):2 (GT), 5 (GT). 

-Methylpannaric acid, pannaric acid 

(trace), unknown (trace) (TLC). 

Leptogium lichenoi.d,es:2 (GT), 5 (GT) 

L. saturninum: 2 (RSU),5 (OV), 9 (GT) 

L. sinuatum: 9 (GT)


*L. teretiusculum:9 (GT). -On Ulmus glabra. 

L. tenuissimum: 5 (HH) 

*Leptogium sp.: 5 (GT, det. PMJ) (an undescribed 

species, tentatively called L. 

imbricatum). 

**Leucocarpia biatorella:5 (RS).-For a 

morphological description, see Poelt & 

Vdzda 198L. It was found on mosses on a 

wet, vertical dolomite rock accompanied 

with e.g. Gyalecta jenensis. Also collected 

in Finland (Muhr 1987). 

Lobaia pulmonaia: 2 (GT), 5 

L. scrobiculata: 2 (RSU, GT, OV), 5,6 

Lopadium coralloideum: 1 (AN), 4 (HH), 5 

(HH, GT) 

L. pezizoideum: 1, 5 (HH, TA) 

Massalongia carnosa: 2 (GT), 5 

Megaspora (Pachyospora) veffucosa: 3 (RSU, 

TA),s (TF) 

Melanelia (Parmelia) exasperata: 5 (TA) 

M. (Parmelia) exasperatula: 2 (GT). -On 

Picea abies. 

M. (Parmelia) infumata: 2 (RSU), 3 (GT), 4 

M. (Parmelia) olivacea: 1,2,3,4,5, 6 

M. (Parmelia) pannifurmis:2 (RSU) 

M. (Parmelta) septentionalis: 5 (TA) 

M. (Parmelia) sorediosa: 3, 4 

M. (Parmelia) stygin: 1,,2 

M. (Parmelia) subauifera: 5 (SI, conf. V. 

Alstrup) 

*Micarea globulosella: 5 (HH). -At bases of 

large Picea abies. Thallus contains gyrophoric 

acid (TLC). With apothecia and 

stalked pycnidia. Only reported once earlier 

from Sweden (Angermanland, LB}Z) 

but probably overlooked. 

*M. granulans Wf. subleprosula): 5 (HH). 

On mosses on rocks. No substances 

detected by TLC. 

M. hgnaia:2 

M. melaena:2 (TF) 

*M. prasina: 5 (GT) 

Microcalicium disseminatumt Z, 5, 6 (MW, 

conf. L. Tibell, RSU, GT) 

Miriquidic a (Leci^de a) atrofulv a : Z 

M. (Aspicilia) complanata: 3 (TA, det. RS) 

M. (Lecidea) leucophaeai 1 (TF) 

M. (Lecidea) lulensis: 1 (TF) 


M. (Lecidea) nigroleprosa: 1 (TF), 3 (RSU) 

Muellerella lichenicola: 4 (RS). -On Xanthoria 

borealrs and X. elegans. 

Mycobilimbia (Lecidea) berengeriana: 1 (GT), 

s (Gr) 

M. (Lecidea) hypnontm: 5 (TF) 

M. tetramera (Bacidia fusca, Mycobilimbia 

obscurata): 5 (GT, HH) 

M. (Bacidia) sabuletorum: 5 (GT) 

Mycoblastus affinis:2 

M. sanguinarius: 2,3 (TF), 4 (TF), 5, 6 (GT), 

7 (rF) 

Mycocalicium subtile : 5 

Nanostictis peltigerae:2 (MW, conf. RS) 

Nephroma arcticum: l, 2, 5 (RSU), 6, 7 

N. bellum: | (TA), 2 (GT), 4,5,7 (Ou 

N. expallidum: 4,5 (PAE) 

N. parile: 1 (OV), 2 (OV, RSU), 3 (GT), 4 

(RSU), 5,6 

N. resupinatum: 2 (RSU), 4 (RSU, GT, OV), 

5,6 

*Nesolechia 

neglecta: 4 (RS). -On Leprarin 

neglecta. 

Ochrolechia androgna coll.: 1 (TA, TF), 2 

(RSU, GT, MW, TA), 3 (RSU, GT), 5 

(RSU, GT), 6,7 (TF) 

O. figida: I (TA), 2 

O. gimmiae:1 (TF) 

O. inaequatula: 1 (HH), 2 (TA) 

O. microstictoides: 2 (GT), 7 (TF). 

Lichesterinic and variolaric acids (locality 

2,TLC). 

O. pallescens:5 (TA), 6 (TF), 7 (TF), 9 (GT) 

Omphalina hudsoniana:3 (RSU), 5 

Opegrapha (Lecanactis) abscondita: 2 (GT),9 

(GT) 

*O. glrocarpa: | (GT), 2 (GT, conf. L. Ka[sten), 

9 (GT) 

Ophiopaffna ventosa (Haematomma ventosum): 

1(OV),2,3 

Pannaria conoplea: 2 (GT) 

P. leucophaea: 2 (OV) ,3, 5 

*P. meditenanea: 4 (AN) 

P. pezizoides: 1,2,3,4,5 (MW, OV), 9 (GT) 

P. praetermissa: 2,3, 5 

Parmelia fraudans: L,2 (GT),3 (PAE), 4 

P. omphalodes: 1,2, 4

26 Gdran Thor 

P, saxatilis: 1,2 (GT), 3, 4 (TA), 5, 6,9 (GT, 

c. ap.) 

P. sulcata: L,2, 5 (GT) 

Parmeliella tiptophylla: 2, 5 (GT, Ou 

Peltigera aphthosa: 1,, 2 (OV), 5 (OV), 6 

(OV), 7 (OV). -On locality 5 also bluegreen 

morphotype. 

P. canina:2 (OV), 4, 5 (OV), 6 (OV), 7 (Ou 

P. collina:4,5 (OV, TA) 

*P. degenii:2,5 (OV), 6 (OV), 7 (OV) 

P. didactyla (P. spuria)' 1, 2 (RSU), 3 (OV), 4, 

5,6,7 (OV, TF) 

P. elisabethae: 2 (OV), 3 (OV),5 (TA) 

*P. kristinssonii: 1 (OV). -On soil on steep 

slope. 

P. lepidophora: 5 (PAE) 

P. leucophlebia: | (OV), 2 (OV), 4 (OV), 5 

(oV),6 (OV) 

P. malacea:1 (OV),2 (OV, RSU), 4 (OV) 

*P. membranacea: 2 (OV). -Among mosses 

on rocky outcrop. 

*P. neckei:2 (OV), 7 (OV) 

P. neopotydactyla: 2 (OV, RSU), 5 (OV), 6 

(o\.),7 (O\r) 

P. polydactylon: 1 (OV),2,5,7 (Ou 

*P. ponojensis:3 (OV), 5 (OV), 7 (OV) 

P. praetextata: 5 (OV) 

P. rufescensi 1 (OV), 2 (OV), 4, 5 (OV), 6,7 

(ov) 

P. scabrosa: 2 (OV), 6 (OV) 

P. venosa:2,5 

Pertusaia albescensi 2 (GT), 5 (TF) 

*P. amara: 5 (TA, GT), 6 

P. carneopallida: t (OV), 2 QA, GT), 5 

*P. coccodes v. phymatodes; 2 (GT). -On old 

Picea abies in a marsh. 

P. dactylina: 1 (RSU, TA), 5,6 (SI) 

P. (Ochrolechia) geminipara: 1, 2 (TA), 3, 5 

(GT),6 

P. lactea: "1. (TF), 2 (TF), 5 (TF) 

P. oculataz | (TA), 5 

*P. ophthalmiza:2,5 

P. panyrga: 1 (TA) 

*P. sommerfeltii: 2 (HH), 3 

*P. stenhammaii:2 (TF), 5 (GT) 

*Phaeocalicium populneum: 2 (MW, conf. L. 

Tibell) 

Phaeophyscia ciliata: 2 (RSU) 

GRAPHIS SCRIPTA 4 (reez) 

P. endococcina: 4 (GT), 5 

*P. kairamoi:5 (HH, TA) 

P. sciastra:2 (OV), 3, 4 (OV), 5 (GT) 

Phaeonhiza nimbosa: 5 (GT), 8 (GT) 

Phaeopyxis puncta: 2 (TA, conf. RS). -On 

Cladonia borealis. 

*Phlyctis argena: 5 (GT). -On Sorbus intermedia. 

Phylliscum demangeonii: 3 (TA) 

Physcia aipolia:2 

P. caesia:2 (GT), 4 (GT, OV), 5 

P. dubia: t,4 (TA), 5 (TA) 

Physconia detersa: 4 (HH). -On mosses on 

ultrabasic (peridotite) rocks. 

P. muscigena: 2 (RSU), 3, 4 (RSU),5 (GT) 

P. perisidiosa: 4 

Pilophorus cereolus: 5 

*Plaqnthiella icmalea (included in Lecidea 

uliginosa in Santesson 1984): 1, 3 (TA), 5 

(GT),6 

P. (Lecidea) oligotropha: | (TA), 3, 5 (TA), 6 

P. (Lecidea) uliginosa:3 (TF), 7 (TF) 

Plaqnthium asperellum: 5 (GT) 

*P, 

flabellosum: 2 (HH), 3 OA), 4 (TF), 7 

(TF) 

P. nigrumz 3,5, 8 (GT) 

P. pannariellum:3 

Platismatia glaucaz 2,3, 4, 5, 6 

P. norvegica: 5 (MW) 

Plectocarpon cladoniae R. Sant. sp. nov. 

(ined.): 2 (GT, RS). -On Cladonia pyxidata. 

Polyblastia gelatinosa: 5 (TA) 

P. hyperborea: 5 

P. sendtneri:5 (GT) 

Potychidium muscicola: 2, 5 (RSU) 

*Poroqphus coccodes: 3 (TA) 

Porpidia (Huilia) cinereoatra: 2 (SI, conf. V. 

Alstrup) 

P. (Huilia) crustulata: 2 (TF), 3 OF), 4 (SD, 

6,7 (TF) 

P. (Huilia) tlavocaerulescens:2 (RS, TF). 

P. (Huilia) macrocarpa: 2 (TF), 3 (TF, VH)' 5 

(TF),6 (TA), 7 (TF) 

P. pseudomelinodes (Huilia melinodes auct. 

p.p.): 2 (TA),3 

P. (Lecidea) speirea: 2 (TF), 4 (EL) 

P. (Huilia) tuberculosaz 2,3,7 (TF)

GRAPHTS SCRTPTA 4 (ree?) 

Protoblastenia rupestrb: 5 (GT) 

*P. terricola:5 (GT) 

Protoparmelia (-ecanora) badia: 1 (OV) 

Protothelenella (Microglaena) coffosa: 2 (RS) 

*P. (Microglaena) leucothelia: 6 (RS) 

P. (Microglaena) sphinctinoidella: 2 (AN) 

P. (Microglaena) sphinctrinoides: 2 (HEG) 

Pseudephebe minuscula: 1 (RSU) 

P. pubescens: L 

Psora decipienr: 3 (VH), 4 

Psoroma hypnorum: t,2 (RSU, GT), 5 

Psorula (Lecidea) rufonigra: 4. -Found by E. 

Timdal. 

frcnothelia papillaria: 1 (HEG) 

Ramalina pollinaria: 2,3 (IH), 4, 5 (TA) 

Mizocafpon alpicola: I 

R. badioatrum: 1, 2 (TF), 3 (TF), 4 (TF), 6 

(sr), 7 (TF) 

*R cinereonigrum: L. -Det. E. Timdal. 

R cinereovirens: 2 (TF), 5 (SI, conf. V. 

Alstrup) 

R. concenticum: 4 (EL), 5 (TF) 

R. copelandii: L 

R distinctum: 5 (TF),6 (TF), 7 (TF) 

R eupetraeum: 3 (TF) 

R expallescens: 1 (TF) 

R. geminatum:2 (GT), 4 (GT), 9 (GT) 

R geographicum: 1., 3 

R. grande: 2 (GT) 

R hochstetteri:5 (SI, conf. V. Alstrup), 7 (TF) 

R. inarense: I 

R. lavatum: 2 (EL, TF), 3 (SI, conf. V. 

Alstrup) 

R. obscuratum: 7 (TF) 

R. oederi:2 (IH) 

R polycatpum: 2 (TF), 3 (EL) 

R roridulum: 1., 5 

R. umbilicatum: 5 

*Rinodina cinereovirens: 1 (HH) , Z (GT), 3, 5 

(GT, HH, TA), 6 (RS). -One of the collections 

by GT from locality 5 were epiphytic 

on Lobaia scrobiculata. 

R efflorescens: 2 (GT, HH), 5 (TA, det. T. 

Tonsberg), 6 (GT). -pannarin and unknown 

(TLC, locality 6). Not reported 

from AL by Santesson Lgg4 but by Foucard 

1990. 

R efigua: 5 (TF) 


R. pyina: 5 (TF) 

R septentrionalis: 2 (TF), 4 (TF), 5 (TF). 

*Sagediopsis ftssurisedens: 1 (RS). -On Aspicilia 

myiniL 

Sagiolechia protuberanEi 5 (GT) 

S. rhexoblephara: 1 (GT) 

Sarcogtne regularu: 5 

Schaereria fuscocinerea (5. tenebrosa): L (EL, 

TF) 

*Sclerophora . (Coniocybe) coniophaea: z 

(cT),6 (AA), e (cT) 

Scoliciosporum umbinum: 7 (TF) 

Slqttea gregaia: 2 (SI, det. V. Alstrup) 

Solorina crocea: 1, 5 (TF), 6,7 

S, saccata: 5 (RSU), S (GT) 

Sphaerophorus fragilis:1 (RSU), 2 

S. globo,srls: 1 (RSU) 

Spilonema revertens: 3 

Sporastatia testudinea: L 

Staurothele clopima: 2 (GT), 4 (GT) 

S. fissa: 7 (TF) 

S. fuscocuprea:2, 4 

*5. 

succedens:5 (RS) 

Stenoqtbe pullatula: 2, 3 

Stereocaulon alpinum: I 

*5. dactylophyllum: 4 (TA) 

S. glareosum: a (TA) 

S. grande: 1r 2,3, 4 

S. nanodes:2 

S. paschale: 3 (SI), 4 

*5. pileatum: L,2 (GT), 6 (GT) 

S. ivulorum: 1 (TA),6 (TA) 

S. sacatile: I,2,3 

S. subcoralloides: 4 

S. tomentosum: lr 2r 3, 4, 61 7 

S. vesuvianum:2 

*Stigmidium 

peltideae: 5 (RS). -On peltigera 

aphthosa and Sctlorina saccata. 

*5. pumilum: a (RS). -On Physcia caesia. 

Tephromela (Lecidea) aglaea: 1 (TF) 

T. (Lecanora) atra: 1.,2, 3, 4, s 

Thamnolia vermicularis : I 

*Thelenella (Microglaena) pertusaiella: 5. 

On Salu sp. 

Theli^dium decipiens : 5 

T. pyrenophorum: 5 (TF) 

Thelocarpon epibolum:5 (TA), 6 (TA). -On 

locality 5 was var. epibolum collected on

28 Goran Thor 

Icmadophila eicetorum and var. longisporum 

on Peltigera leucophlebia. On 

locality 6 was var. epibolum collected on 

Soloina crocea. 

*7. impressulum: 5 (GT). -On base of old 

Picea abies at small stream. 

*Thelopsis 

flaveola: 5 CfF, collection in UPS, 

HEG). -Found on base of old Picea abies 

in a shaded and wet habitat. A surprising 

finding indicating that this species is not 

confined to broad-leaved forests in South 

Sweden. 

*7. melathelia: 2 (GT), 5 (GT) 

Thermutis velutina: 9 (GT) 

Thrombium epigaeum: 2 (TA) 

Toninia cumulata: 1 (TA, conf. E. Timdal, 

AN),3. 

T. rosulata: 5 (GT, det. E. Timdal). 

Trapelia coarctata:2,5 (GT), 7 (TF) 

T. granulosa: 3 (GT) 

T. involuta:2,7 (TF) 

T. placodioi^des: 7 (TF) 

Trapeliopsis flexuosa (Lecidea aeruginosa): 7 

(TF) 

T. (Lecidea) granulosa: 1 (TA), 2, 3 (GT), 4, 

5, 6,7 

*Tremella cetrariicola Diederich & Coppins 

ined.: 2 (RSU, det. Diederich).-On Cetraia 

chlorophylla. 

Tremolecia atrata: L (SI, TF), 2 

Umbilicaria crustulosa: I (PAE) 

U. cylindrica; I 

U. deusta: L,3, 6 

U. hirsuta:2 

U. hyperborea: t,2,3, 6 

U. proboscidea: I 

U. vellea: 1,2 

Usnea filipendula:2 

Varicellaia rhodocarpa: 1,2,5 (TA), 6 (TA) 

Vemtcaria anziana: 7 (TF) 

*V. 

foveolata: 5 (EL) 

V. nigrescensi 5 (TF) 

*Wentiomyces peltigeicola: 5 (RS). -On 

Peltigera aphthosa. 

Xanthoparmelia (Parmelia) centifuga: L, 2, 3, 

6 

X. (Parmelia) incuruaz 1,2 


Xanthoia borealis: 4 (AN, GT, HH, TA, all 

det. RS), 5 (TA). -For a morphological 

description, see Poelt & Petutschnig L992. 

X. candelaria: 1, 5 (TA) 

X. elegans: | (TF), 2 (RSU, GT), 3, 4,5 

X. sorediata: 3 (OV), 4 (GT), 5 

Xylographa parallela (X. abietina): 2 (GT)' 3 

(RSU), 4,5,6 (TA), 7 CfF) 

X. vitiligo: 1 (GT), 2 (GT), 3 (RSU), 5 (GT' 

TA),6 (RSU, GT,TA),7 (TF) 

**Zwackhiomyces berengerianus coll.: 8 (GT' 

det. M. Grube).-On a sterile, crustose 

lichen (cf. Mycobilimbia sp.) on mosses on 

a shaded wet rock. 

Visited localities 

All localities: Sweden, Asele I-appmark. 

The positions of most localities are given 

in long/lat, Swedish local coordinate system 

(non-numerical and numerical) and UTM. 

1. Vilhelmina parish, SE slope of the 

mountain Satsfiallet 6 km SW the village 

Sarnbs, alt. 880 - 960 ffi, 5 August L991, &" 

56' N 15" 15' E, 23FOe3d,7204 L474, UTM 

33W WN 118015. Alpine heath on acid soil. 

2. Vilhelmina parish, 20 km ESE Sa:rniis, 

N and S shore of the rapid Dimforsen (near 

the bridge) in the stream Kultsjotn, alt. 390 m, 

6 August 199L, 64" 56' N 1.5' 44' E,23FO|3- 

4c, 7205 1498, UTM 33W WN 35ffi2I. 

Siliceous rocks, including alum shale, in mixed 

coniferous/deciduous forest along the river. 

3. Vilhelmina parish, 6 km ESE Saxniis, 

Kultsj6luspen, W shore of the stream Kultsj6in, 

alt. 520 m, 6 August 1991 , 64" 57' N 15o 

28' E, 23F1gLd, 7206 L484, UTM 33W WN 

2L9041,. Mesic to dry Pinus rylvestrb forest 

with siliceous rock outcrops by the river. 

4. Vilhelmina parish, 10 km E Klimp{eil' 

S slope of the hill R6berget at the [-ake Kultsjon, 

100 m N the road, alt. 600 6?0 il, 6 

August !99L,65o 04' N 15" 01 '8, 23F3c3c, 

72L9 !464, UTM 33W WN 0L?160. Ultrabasic 

peridotite cliff with talus-slope. 

5. Risbiick parish, NE sloPe of the 

dolomite mountain Kalvberget L0.5 km WNW 

the village Risbiick, alt. 560 - 620 ffi, 8 August

GRAPHTS SCRIPTA 4 (r99?) 

I99L,64" 44' N 15' 20' E,22F59c-6Oc, 7I8L 

1477, UTM 33W WM 161, 779. Steep rockface 

with a talus-slope in mesic Picea abies 

forest with scattered deciduous trees. A few 

collections were also made along the small 

stream between the N slope of Kalvberget 

(starting 500 m NNW the small lake) and lake 

Mevattnet (these collections marked "along 

the stream"). 

6. Vilhelmina parish, 18 km N Stalon, E 

slope of the mountain Morrosjoliden near the 

road Eriksberg Vtistansjo, alt. 625 m, 9 

August 199t,65" 06' N tr 5" 53' E, 23G4aAc-d, 

7223 1504, UTM 33W WN 208206. 

Oldgrowth, open Picea abies forest. 

7. Vilhelmina parish, Sarnhs, in the 

surrounding of Sa,xnas FjdllgArd (Stiftsgfirden) 

100 m E the church, 5 - 9 August \991, 64" 

58' N 150 20', E, UTM 33W WN 155055. 

Roadsides and open Betula and Picea forests. 

8. Vilhelmina parish, NW of the village 

Storvallen, 1.5 km E the hotel in Klimpfiiill, 

along the stream Doda biicken 300 - 600 m N 

the road, 29 August L988, alt. 580 630 m, 

65" 04' N 1.4" 51' E, 23F 3a, UTM 33W VN 

925L62. Calcareous rocks along the stream. 

9. Vilhelmina parish, B6ngntis, 

Bjdrnskalet, SSE exposed talus-slope, 30 

August 1988 and 1 August 1990, alt. 560 

590 m, 64o 56' N 15o 37' E, 23F 0i. Open 

talus-slope with siliceous rocks and small 

water fall surrounded by trees. 

Acknowledgments 

I am most grateful to all participants in the 

meeting who have supplied additions to the 

species list. In particular I want to thank my 

colleagues in the organizing committee, Per 

Anders Esseen, Jan-Eric Mattsson and 

Rikard Sundin. Information on lichenicolous 

fungi new to Asele Iappmark was supplied by 

Rolf Santesson who also kindly made various 

comments on the manuscript. 

References 


Ahlner, S. 1938: Weitere Beitrtige zur 

Strauch- und l-aubflechtenflora von 

Asele lappmark. Arkiv for Botanik 2gA, 

no 9. 

Degelius, G. 1932: Zur Flechtenflora des 

siidlichsten lapplands (,A,sele lappmark). 

1. Strauch- und laubflechten. Arkiv for 

Botanik 25A, no 1.. 

DegeliuS, G. 1954: The lichen genus Collema 

in Europe. Morphology, ta:ronoffiy, ecology. 

Symbolae Botanicae Upsaliensis 13: 

?. 

Foucard, T. L990: Svensk skorplavsflora. 

Interpublishing. 

Holtan-Hartwig, J. L99L: A revision of the 

lichens Bryonora castanea and B. curvescens. 

Mycotaxon 40: 295-305. 

Karstr6ffi, M. & Thor, G. L991,: Field meeting 

in Vuollerim, Lule I-appmark by Swedish 

lichenologists. Graphis Scipta: 3: 84-93. 

Moberg, R. L986: Lichenes selecti exsiccati 

Upsaliensis, Fascicle L. Thunberyia 2: 1- 

10. 

Muhr, L.-E. L987: Floristic notes from Sweden 

and Finland. Graphis Scripta 1: &- 

65. 

Poelt, J. & Petutschnig, W. 1"992: Beitrdge zur 

Kenntnis der Flechtenflora des Himalaya 

IV. Die Gattungen Xanthoria und Teloschistes 

zugleich Versuch einer Revision 

der Xanthoria candelaria-Gruppe. Nova 

Hedvigia 54: L-36. 

Poelt, J. & Ydzda, A. 1981 : Bestimmungsschlilssel 

europiiischer Flechten. Ergiinzungsheft 

2. Yaduz. 

Rune, O. L953: Plant life on serpentines and 

related rocks in the north of Swed en Acta 

Phytogeographica Suecica 3 I : L-139. 

Santesson, R. L984: The lichens of Sweden 

and Norway. Stockholm & Uppsala 

Swedish Museum of Natural History. 

Stenberg,L. & Westerberg, S. 1983: Murruta, 

Asplenium ruta-muraria funnen i Asele 

lappmark. Natur t Non 2: 72 - 74.

Notes on the genus Caloplaca in the Siberian arctic 

ULRIK SOCHTING, MICHATLZHURBENKO and ERIC STEEN HANSEN 

Sochting, U., Zhurbenko, M. & Hansen E. S. 1992: Notes on the genus 

Caloplaca in the Siberian arctic. Graphis Scipta 4: 30-32. Stockholm. ISSN 

0901 -7593. 

Data are presented on the occurrence of 13 species of Caloplaca collected by 

the second author during expeditions to Bennett Island, New Siberian Islands 

in the summer of 1989 and to the Taimyr Peninsula in the summer of 1990. 

Ulnk SOchting, Institut for Sporeplanter, Univ. of Copenhagen, @. Farimagsgade 

2D, DK-1353 Copenhagen K Denmark. 

Michail Zhurbenko, Komarov Botanical Institute, Laboratory of Lichenologt 

and Bryologt, Academy of Sciences of Russia, Prof. Popov str. 2, St 

Petersburg 197376, Russia. 

Eric Steen Hansen, Botanisk Museum, Univ. of Copenhagen, Gothersgade 

130, DK-1123 Copenhagen K Denmark. 

The genus Caloplaca is well represented in 

polar regions of the world. Recently some 

papers have increased the knowledge of this 

genus in the Arctic and Subarctic (Hansen et 

al. L987, SOchting 1989) and in the Antarctic 

(SOchtin g & @vstedal 1992, Olech & SOchting 

in prep.). However, there are still large areas, 

where the Caloplaca flora is still unsufficiently 

known. One of these is the Siberian Arctic, 

from where some information, however, was 

provided by Lynge (1928). About 20 species 

have been reported from that region 

(Zhurbenko 1991). 

The present paper deals with Caloplaca 

species collected by the second author in 

Siberia. The specimens are preserved in the 

Cryptogamic Herbarium of the Komarov 

Botanical Inst. (LE). Additional data on 

climate and and occurrence of other lichen 

genera were published by Zhurbenko & 

Hansen (1992). 

Localities 

1. W. Taimyr: Mouth of Ragozinka River, 

coast of Eniseiskii Gulf. 72" 48' N, 80o 53' 

E. 

2 W. Taimyr: Mouth of Uboinaia River, 

coast of Cara Sea. 73o 39' N, 82o 22' E. 

3 'W. Taimyr: Near river Right Uboinaia, 

small mountain up to 2I7 m. 73" 25' N, 

82" 51'E. 

4. New Siberian Islands: Bennett Island. 76" 

37' - 76o 45'N, 148o 28' - L49" 27',E. 

Species list 

Caloplaca alcarum: l. C. alcarum was described 

by Poelt (1954) from Novaia 

Zemlya from where it was reported by 

Lynge (1928) under the name C. sacicola 

var. obliterata. It was also recorded from 

Greenland (Hansen et al. L987) and 

Svalbard (Hertel & Ullrich 1976, Hertel 

L977, SOchting 1989). It grows on strongly 

eutrophicated maritime coasts in the Arctic.


C. ammiospila: t-4. This species, which was 

formerly often recorded under the name 

C. cinnamomea (Th. Fr.), Oliv., is widespread 

in arctic and boreai regions of the 

Northern Hemisphere (Hansen et al. 

L987, SOchting 1989, Santesson 1984, 

Egan L987). It was recently recorded from 

Antarctica (SOchting & @vstedal l99Z). 

Some of the material collected in Siberia 

on lignum has affinity to C. spitsbergensis 

H. Magn. The tanonomic status of this 

species is however doubtful, and the 

specimens may be only modifications of C. 

ammiospila (SOchting 1989). C. spitsbrrgensis 

was reported from Svalbard 

(Magnusson 1944) and Greenland 

(Hansen et al. 1987). 

C. caesiorufella: l-4. Apparently widespread 

in arctic, subarctic and boreal regions. It is 

recorded from arctic Canada (Ahti et al. 

1973), Greenland (Hansen et al. 1987, as 

C. phaeocarpella), Svalbard (SOchting 

1989), Scandinavia (Santesson l9S4) and 

Siberia (Nylander 1885). 

C. ceina: l, 2,4. Seems to be widespread in 

the Northern Hemisphere, from where it 

is recorded from a variety of substrates. It 

is not known from Antarctica. 

C. fraudans:1. This is an arctic species 

previously recorded from North America, 

Greenland, Svalbard, Scandinavia and 

Novaya Zemlya (Magnusson 1944). 

C. jungermanniae: l. An arctic-alpine species, 

which is widely distributed over the 

northern hemisphere and is recorded from 

North America (Egan 1987), Greenland 

(Hansen et al. L987), Svalbard (Lynge 

1924), Scandinavia (Santesson L9S4) and 

Siberia (Zhurbenko t99t). 

C. lithophila coll. : 4. Specimens belonging to 

the C. holocatpa group and growing on 

rocks are presently little understood. 

Accordingly any comments on distribution 

has little meaning. 

C. nivalis:1. This arctic-alpine species, is 

recorded from North America (Egan 

t987), Greenland (Hansen et al. l9B7), 

Svalbard (Elvebakk I9g4), Scandinavia 

Caloplaca in Siberia 31 

(Santesson 1984) and Siberia (Zhurbenko 

1ee1). 

C. phaeocarpella: L, 3. C. phaeocarpella has so 

far not been recorded from North America, 

and is not with certainty identified 

from Great Britain and Scandinavia. It is, 

however, recorded from Greenland 

(Hansen et al. 1987) and Svalbard 

(SOchting 1989) and the holotype is from 

the Kola Peninsula in Russia. It is previously 

recorded from Siberia (Zhurbenko 

lggL). 

C. sibiricai 1, 4. Described by Magnusson 

(L952) from Torne lappmark and arctic 

Siberia. The holotype is from Minin Island 

and additional specimens are from Taymir 

Island, Cap Cheljuskin and Dudinka. 

Hansen et al. (L987) were unable to trace 

any material from Greenland, but it was 

recently published from Svalbard 

(SOchting 1989). The specimens from 

Bennett Island grew on both moss and 

lignum. 

C, tetraspora: l-4. Appears to be a common 

and widespread arctic-alpine species 

which has been recorded from both the 

Northern Hemisphere (Egan 1997, 

Hansen et al. 1987, Santesson 1984, Lynge 

L924, Zhurbenko 1991). It is also known 

from Antarctica (Jakobsen & Kappen 

1988, SOchtin g & Olech, unpublished). 

C. tiroliensls: l, 2,4. This species is together 

with C. tetraspora common in arcticalpine 

areas of the Northern Hemisphere 

(for references se under C. tetraspora). It 

is also a common species in Antarctica 

(Sochting & ovstedat t9g2). 

C. torno€nsis: 1, 2. C. torno€nsb is recorded 

from Northwest Territories, Canada 

(Thomson & Scotter L984), from Greenland 

(Hansen et al. L9B7), Svalbard (poelt 

1969) and Scandinavia (Santesson 1984). 

It is new to Siberia.

32 Ulnk Suchting et al. 

References 

Ahti, T., Scotter, G. 'W. & Viinsk6, H. L973: 

Lichens of the Reindeer Preseru€, Northwest 

Territories, Canada. Bryologist 76: 

48-76. 

Egan, R. S. L987 z A fifth checklist of the 

lichen-forming, lichenicolous and allied 

fungi of the continental United States and 

Canada . Bryologist 90: 77 -173. 

Elvebakk, A. 1984: Contributions to the 

lichen flora and ecologry of Svalbard, Arctic 

Nonvay. Bryologist 87: 308-313. 

Hansen, E. S., Poelt, J. & Sgchting, U.. L987: 

Die Flechtengattung Caloplaca in GrOnland. 

Meddr. Grqnland Biosci. 25: L-52. 

Hertel, H. L977: Bemerkenswerte Flechtenfunde 

aus dem Gebiet des Kongsfiordes 

und des Is{ordes (Spitzbergen). Hercogia 

4:367-40L. 

Hertel, H. & Ullrich, H. L976: Flechten von 

Amsterdamoya (Svalbard). Mitt. Bot. 

Miinchen 12: 4Ll -512. 

Jacobseo, P. & Kappen, L. 1988: Lichens from 

the Admiralty Bay region, King George 

Island (South Shetland Islands, Antarctica). 

Nova Hedwigia 46: 503-510. 

Lynge, B. 1924: Lichens from Spitsbergen I. 

Result. Statsunderst. Spitsberyenelcsped. I 

(5). 

Lynge, B. I9Z8: Lichens from Novaya kmlya 

(excl. of Acarospora and Lecanora). Rep. 

Scr Results Norwegian Exped. Novaya 

Zemlya 1921, 43: l-299. 

GRAPHTS SCRIPTA 4 (1992) 

Magnusson, A. H. L944: Studies in the ferruginea-group 

of the genus Caloplaca. K 

Vet. Vitterh.-Samh. Handl., 6. Ser. B 3 no. 

1. 

Magnusson, A. H. L952: Lichens from Torne 

lappmark. Ark. Bot. Ser. 2 Bd. 2: 45-249. 

Nylander, W. 1885: Lichenes novic Freto 

Behringti. Flora 68: 439-446. 

Poelt, J. L954: Die gelappten Arten der 

Flechtengattung Caloplaca in Europa. 

Mitt. Bot. Staatssamml. Milnchen 2: 11- 

31. 

Poelt, J. L969: Bestimmungsschliissel europiiischer 

Flechten. J. Kramer, khre. 

Santesson, R. 1984: The lichens of Sweden 

and Norway. Stockholm & UPPsala. 


Sochting, U. 1989: Lignicolous species of the 

lichen genus Caloplaca from Svalbard. 

Opera Bot. 100: 241-257. 

Sgchting, IJ. & Ovstedal, D. O. L992: 

Contribution to the Caloplaca flora of the 

western Antarctic region. Nord. f. Bot. 12: 

L2L-134. 

Thomson, J. 'W. & Scotter, G. 'W. L984: 

Lichens of Bylot and northern Baffin 

Islands, Northwest Territories, Canada- 

Bryologist 87:228-232. 

Zhurbenko, M. P. L99I. The preliminary noncritical 

checklist of the lichens of Siberian 

arctic (incl. Novaia Zemlia). Manuscipt- 

Zhurbenko, M. P. & Hansen, E. S. t992: 

New, rare or othenvise interesting lichen 

species from the Siberian Arctic. Mycotcaon 

45:275-284.

Protothelenella rylina new to Finland 

PIETER. P. G. VAN DEN BOOM 

van den Boom, P. P. G. 1992: Protothelenella rylina new to Finla nd. Graphis 

Scipta 4: 33-34. Stockholm. ISSN 0901 -7593. 

Protothelenella xyltry! Mayrh. & Poelt, so far known only from the Alps, is 

reported as new to Finland, from the Pallastunturi reserve near Muonio. 

Pieter van den Boom, Azi€laan 12, s69ILC son, The Netherlands. 

In the summer of 1988 I made a trip mainly 

through central and northern Fennoscandia 

with the aim to study lichens in this part of 

Europe. A few days were spent in northern 

Finland, the area of Muonio, within the 

Pallastunturi reserve. ca. 170 collections were 

made. 

The most important collection was 

Protothelenella xylina. The specimen was 

collected 15 km E of Muonio near the 

entrance of the Pallastunturi reserve from 

weathered lignum of a high stump in a damp 

Betula-Picea-Pinus forest with scattered 

boulders, 67o 57' N, 24" 00, E, alt. 300 m, 24 

July 1988. P. rylina was growing in a 

moderately shaded place. 

The morphological characters of the 

species are: thallus endoxylic, yellowish greygreen; 

perithecia black, up to 0.3 mm in diam., 

numerous; ascospores submuriform to 

muriform, colourless, Z0-ZS x L}-_LZ ltm 

(Figure 1). 

Associated with Protothelenella rylina 

were: Buellia grbeovirens, Calicium 

denigratum, Cladonia botrytes, C. cenotea, 

Hypocenomyce scalaris, Lecanora cf. hypopta, 

Mycoblastus sanguinaius, Ochrolechia 

androgtna, Parmeliopsis ambigua, p. 

hyperopta, a Porpid.ia sp. which refers to p. 

crustulata, Stereocaulon alpinum (atranorin, 

lobaric acid and unknown trace RF6 detected 

by TLC) and Trapeliopsis granulosa. 

The lichen genus Protothelenella is 

separated from Microglaena by having amyloid 

asci, a distinct amyloid apical apparatus, 

amyloid jelly and by the structure of the 

ascocarps (Mayrhofer & Poelt 1935). These 

authors provide the first description of p. 

rylina and mention five other species of 

Protothelenella. Howevef a few years later 

three additional species have been published 

in Mayrhofer (1987). 

Distribution 

Protothelenella rylina is known from two 

localities in the Alps of Switzerland and 

Austria. Collected only from lignum of pinus 

cembra (Mayrhofer & Poelt 1985). 

Although the occurence of relative many 

species of Protothelenella have been 

mentioned from northern Europe in 

Mayrhofer (1957) there were no previous 

records of P. rylina. 

A duplicate of the collection of p. rylina is 

deposited in the herbarium of the Botanical 

Museum at Helsinki. 

Acknowledgements 

Thanks are due to Dr. H. Mayrhofer for 

confirming the identification of p. rylina and 

to Dr. H. Sipman for the identification of 

Stereocaulon alpinum.

34 P. P. G. van den Boom 

Refercnces 

# L0;rm 

Figure 1. Protothelenella rylina, ascospores. 

Mayrhofer, H. & Poelt, P. 1985: Die 

Flechtengattung Microglaena sensu 

Zahlbruckner in Europa. Hercogia 7: 13- 

79. 


Mayrhofer, H. 1987: Ergdnzende Studien zur 

Ta:

Caloplaca soropelta (8. S. Hansen, Poelt & Sgchting) Sgchting 

comb. nov. 

ULRIK SOcHTING 

Sochting, U. L992: Caloplacq soropelta (E. S. Hansen, Poelt & SOchting) 

Sochting comb. nov. Graphis Scipta 4: 35-36. Stockholm. ISSN 0901-7 Sg{' 

Caloplaca citina var. soropelta was described by Hansen, Poelt & SOchting 

(1987) from Greenland. Subsequent finds from Svalbard has proved th; 

ta:ron to be well delimited from C. ci.trina and it is accordingly heie raised to 

species level. New material, which is partly fertile, allows 1o make a more 

comprehensive description, and thus to establish a better understanding of 

the species. 

UMk Sochting, _In1tltut for Sporeplanter, University of Copenhagen, O. 

Faimagsgade 2D, DK-1353 Copenhagen K. 

Caloplaca soropelta €. S. Hansen, poelt 

& Sgchting) SOchting comb. nov. 

Basionym: Caloplaca citrina var. soropelta E. 

S. Hansen, Poelt & Sochting. l9g7; Meddr. 

Gr^nland Biosc. 25: 26. 

Morphology 

Thallus up to 2 cm in diameter, consisting of 

centrally crowded, laterally dispersed, irregular, 

but hardly incised squamules, which are up 

to about I mm, bright orange yellow to bright 

golden yellow and often slightly pruinose. fhe 

squamules are peltate in the centre, but 

smaller, more convex and more appressed to 

the substratum towards the periphery of the 

thallus. Older squamules are concave, with 

often greyish cortex, and most often golden 

yellow soralia developing from the under surfgce 

of the periphery. soralia never develop at 

the surface of the squamules, which remain 

covered by a strongly concave cortex. Soredia 

20-30pcm diam. 

Apothecia lecanorine to zeorine, very rare 

and unripe in the material. They are sessile 

centrally on the squamules, at the material 

studied about 0.6 mm diam. Disc initially concave, 

orange. The margin is about 0.1 mm 

thick, concolorous with disc. The outer surface 

of the margin is sorediate. 

Anatomy 

Cortex thin, little differentiated. Algal layer 

about 300 pm thick, diffuse. A fine epipsamma 

of anthraquinones is very abundant at the 

soralia, less so on the cortex. 

Thalline margin with many algae producing 

soredia laterally. Hymenium immature, but 

paraphyses seem to be little branched and 

hardly inflated apically. 

Ecologr 

Caloplaca soropelta is a species of highly 

eutrophicated, calcareous rock. It prefers 

cracked surfaces. on rocks with soil accumulated 

in the cracks it may form peltate 

squamules with rhizinae penetrating the soil. 

c. soropelta is often associated with xanthoria 

elegans, X. sorediata, X. candelaria, Caloplaca 

sacicola and C. decipiens.

36 Ulrik SPchting 

Discussion 

There are a number of sorediate species of 

Caloplaca in Svalbard including C. citrina and 

C. epiphyta. Several of these species colonize 

both rocks, soil, detritus and lignum. Their 

ta:ronomy is not always clear, and some of 

them are probably undescribed. C. soropelta, 

however, appears to be a very well delimited 

species. Its closest relative may prove to be C. 

tominii, which grows on soil and has been 

recorded from more or less continental regions 

in several continents. C. tominii differs in 

having less regularly rounded squamules and 

larger soredia, YV.30-40 pm. 

Specimens studied 

GRAPHIS SCRIPTA 4 (Lee?) 

Svalbard, Sabine [and, Tempel{orden: Below 

Fjordnibba, UTM: WH 450L, Alt.: 10 m. Bird 

perching rock of limestone, 20 m from the sea. 

L2. vii. I9frc, US 5623 (C); Fjordnibba below 

bird cliffs, NW-exposed, UTM WG 4499, Alt 

180 m. Rock of chert. 13. viii. L986, US 5642 

(c). 

Reference 

Hansen, E. S., Poelt, J. & Sochting, U. L987: 

Die Flechtengattung Caloplaca in GrOnland 

Meddr. Grqnland Biosc.25: I-52.

Boganmeldelse 

Toninia 

Timdal, E. L99L: A monograph of the genus 

Toninia (lrcideaceae, Ascomycetes). Opera 

Botanica 110: L37 pp. Pris 4L0 DKK. 

Revisionen omfatter ca. 6500 indsamlinger fra 

alle kontinenter med undtagelse af Antarktis, 

isrer fra Europa og Nordamerika, og er 

baseret pA anatomiske og morfologiske 

karakterer. TLC er udfort pi ca. 2000 kollektioner, 

de fleste arter indeholder ikke lavstoffer. 

15 arter har et eller flere stoffer, der ikke 

er bestemt. Flere arter omfatter mere end en 

kemotype. 

Slregtsafgrrensningen er baseret pA apothecietype, 

asci, paraffser og pyknider. Apothecierne 

er lecideine med amyloid hymenialgel, 

asci kglleformede med ikkeamyloid va,g 

omgivet af en amyloid gelatinOs kappe. Der er 

en tydelig tolus med okularrum omgivet af en 

svagt amyloid inderzone og en strerkt amyloid 

yderzone. ParaSserne er lige, kun lidt 

grenede, tyndvrggede, kun lidt sammenklrebede 

og 2-2,5 pm tykke forneden, 

endecellen er opsvulmet til 4-6 pm og omgivet 

af en diffust afgrrenset gelatinos pigmentkappe, 

hos nogle arter er pigmentet dog 

indlejret i den overste del af endecellens vr;g. 

Ascosporerne er 6ncellede eller er delt af 

tvrervregge. Pykniderne er flaskeformede, 

konidierne trAdformede, krumme og ca. 12-20 

x I pm. Thalluskaraktererne spiller ingen rolle 

for slregtsafgrrensningen, b6de arter med 

skorpeformet thallus som ulicheniserede, 

lavboende svampe medtages i slregten. 

48 arter anerkendes pfi verdensplan, L9 

arter findes i Skandinavien. 13 nye arter og 5 

underarter beskrives. 18 nye kombinationer 

foreslfis, heraf 2 i andre shgter. Sbgterne 

Kiliasia og Lobiona og Leci^dea sect. Toniniopsis 

inddrages i Toninia. De vigtigste 

rendringer er (gamle navne i parentes): 

T. auiculata Timdal (Lobiona albomarginata) 

T. athallina (Hepp) Timdal (Kiliasia athallina) 

T. episema (Nyl.) Timdal (Kiliasia episema) 

T. philippea (Mont.) Timdal (Kiliasia phihpp€a, 

K riparia) 

T. lutosa (Ach.) Timdal (Z vemtcosa\ 

T. dffiacta (Massal.) hhlbr. (7. candida ssp. 

diffracta) 

T. massata (Tuck.) Herre (7. kellei, T. 

glaucomela) 

T. albilabra (Dufour) H. Olivier (7i albomarginata) 

T. physaroides (Opiz) hhlbr. g. lurida, T. 

alluviicola) 

T. verrucaroides (Nyl.) Timdal Q. lcolaV T. 

aromatica ssp. ceruina) 

T. taurica (Szat.) Oxner (7. clemens) 

T. sedifolia (Scop.) Timdal (7. caentleonigricans) 

T. squalida (Ach.) Massal._ (inkl. T. veffuculosa, 

T. multiseptata, T. havaasii ssp. 

havaasii) 

T. coelestina (Anzi) VEzda (7. havaasii ssp. 

aggregata) 

T. cinereovirens (Schaerer) Massal. (7. sbarbaronis) 

T. aromatica (Sm.) Massal. (incl. T. fusispora) 

Micarea granulans (Vainio) Timdal (M. subleprosula) 

I et indledende kapitel diskuteres anvendelsen 

af illustrationer som typemateriale og Timdal 

g@r opmrerksom at enhver h8ndlavet tegning 

er et udtryk for kunstnerens indtryk af det 

studerede materiale, og ikke et materiale som 

forfatteren baserer sit indtryk p6. Derfor 

accepteres sSdanne tegninger ikke som typemateriale. 

Efter beskrivelse af slregten og 

diskussion af afgrrensningen fOlger en nogle til 

arterne. Det kunne her have veret nyttigt 

ogsA med en nogle til de 39 nrertst6ende eller 

overfladisk lignende slregter, som diskuteres. 

De enkelte arter er grundigt beskrevet med 

henvisninger til publicering, synonymer, typemateriale, 

taxonomiske bemrerkninger, habi-

38 Boganmeldelse 

tatsangivelser, udbredelseskort og fremragende 

sorthvide fotos. Studerede exsiccater 

samt antallet af kollektioner fra forskellige 

lande og regioner er opremset. 107 ekskluderede 

eller ikke undersogte arter omtales 

kort. 

Revisionen er baseret pi sunde taksonomiske 

principper, den er verdensomsprendende 

og bygger pi et stort materiale, 

GRAPHTS SCRTPTA 4 (r99z) 

den er konsekvent gennemfart, klart skrevet 

og prent trykt. Einar skal have en stor tak for 

dette arbejde. Vi er mange som har delt hans 

irritation over ikke at kunne bestemme almindelige 

Toninia-arter. Den tid er nu forbi, i 

hvert fald i vores del af verden. 

Vagn Alstrup

Deichmann Branth - A portrait based mainly on his letters 

VAGN AI-STRUP 

Alstrup, V. L992: Deichmann Branth A portrait based mainly on his 

letters. Graphis Scripta 4: 39-43. Stockholm. ISSN 0901 -7593. 

A biography is given of the Danish amateur lichenologist Jacob Severin 

Deichmann Branth (1831-l9ll) based mainly on his correspondence for 

more than 60 years with his friend Otto Moller. 

Vagn Alstrup, Institut for gkologisk botanih Q. Farimagsgade 2D, DK-1353 

Kqbenhavn K Danmark. 

Branth was born 1831 in NykObing Mors, 

Denmark. After his mother's death two years 

later he was brought up by his uncle, the rural 

dean R. S. Deichmann. In L849 he graduated 

from secondary school and began to study 

theology at Copenhagen University, where he 

met Otto MOller. The correspondence 

between the two students began in 1853 and 

continued until Christmas 1914, shortly before 

Mgller died. 242 letters were exchanged 

between them. 

The letters are all preserved and have 

been published in full length (NOrr 1936). In 

the letters the two friends exchange opinions 

about all aspects of their personal life, 

religious matters, culture, politjcs and science. 

Being personal letters they are often written 

in a straightfonvard, powerful style, and they 

contain expressions and personal opinions 

about colleagues and other public persons, 

which they would hardly have published. As a 

historical source they are therefore valuable, 

although it sometimes appears, that they kept 

a debate open by provoking exaggerations or 

ambiguities to tease each other. They often 

used words which cannot be found in ordinary 

dictionaries and my translation into English 

does probably not give the writers full justice. 

The following portrait is based on the 

letters as well as on Norr's introduction, 

Christensen (L924-L926) and on my personal 

contact with Branth's youngest daughter Ruth 

Deichmann Branth, born L903. Several 

photographs of Branth at different ages are 

included in NOrr (1986). Christensen also 

brought a portrait. 

Deichmann Branth's professional caneer. 

Branth worked as a private tutor 1358-L86L, 

then he became headmaster at Tonder 

Realskole. After the German occupation of 

Slewig L864 he was dismissed and was 

unoccupied until he became assistent vicar in 

SkrelskOr, Zealand, L866. I87l to 1880 he was 

vicar at Elling in north Jutland and 1880- 1905 

vicar at Sneptrup, east Jutland. After his 

retirement he moved to Kolding where he 

died. 

Family life. 

Deichmann Branth married Sophie 

Dumreicher 1862. Branth was sorry, that they 

did not get any children, but 1872 they had a 

foster-girl, who brought some life to the 

house. Sophie was very quiet and religious, 

and she turned more and more inwards. 

Branth wrote to Moller, that she had almost 

left this world long before she died shortly 

after their silver jubilee in L887.

40 Vagn Alstrup 

In 1889 Branth married Emmy Boldsen, at 

that time 25 years old. MOller was shocked to 

hear that. Mgller understood that his friend 

was very lonesome without any family, and 

that he wanted a wife to take care of the 

house. But he should have married an older 

woman, and the marriage could hardly be 

based on love, as according to Moller people 

above 30 years did not fall in love. But 

Deichmann Branth declared he had never 

been so much in love and that the love was 

mutual. The happy couple produced 7 

children, the last one was born in L907. 

Branth was very fond af the children, but the 

large family also gave some economic 

problems. 

Branth had spent much money on books 

and travelling, but on his old days he was 

afraid to die before the children grew up, so 

that the family would have to live on a widow's 

pension, which was much lower than his own. 

Branth complained that his wife would give 

out a coin after turning it only once in the 

hand, although she ought to turn it trvice! The 

daughter Ruth remembered, that her father 

was always worried about money, when they 

lived in Kolding. She was interested in 

photography but was not allowed to keep the 

tapwater running, and she had to rinse the 

prints when her father was out. 

Branth travelled much. He participated in 

a large student meeting in Uppsala in 1856. 

After finishing his studies he went on a 

journey to ltaly in 1857. He participated in 

scientific meetings in Copenhagen and 

Stockholm and in several church meetings and 

conventions. In 1891 he participated in a 

botanical excursion to Bornholm to teach the 

students about lichens. On his older days he 

went to stay at a health resort every year, to 

prevent his rheumatism to develop. 

Life and economy of a vicar. 

As payment for the religious duties the church 

gave the vicar a farm to live oo, and it was 

then up to the vicar, how much he could earn 

as a farmer. Often they had to pay pension to 

the former vicar or his widow. Branth payed 

GRAPHIS SCRTPTA 4 (reez) 

1000 kr per year to his predecessor, which he 

thought was far too much, as the old fellow 

was completely insipid, avaricious and pettish 

and absolutely had no pleasure of or need for 

the money. Branth was very happy when the 

predecessor died and the widow should have 

only 500 kr per year. Branths successor on the 

contrary paid too little, as the amount was 

based on the last 5 years relatively low taxes. 

Branth called his successor a beast, an 

agreeable party-man, ventriloquist, and 

christmas-he-goat for the parish council. The 

wife was called self-important, boasting and 

obtrusive. 

Many vicars let their land on lease, but in 

long periods Branth took care of the farming 

himself, as it was difficult othenvise to earn 

enough. He had many problems with the 

farm-hands and often complained about 

falling prices, other troubles and the time 

spent on farming, which could othenvise have 

been spent on church duties or science. He 

was also proud that he could raise the farm 

production, and the farmers of the parish 

looked up to him for his farming. Branth 

supplemented his income by writing books and 

articles about religious matters. 

Branth was an eager preacher and 

seriously interested in the religious life of his 

congregation and tried to improve their 

chastity, but he often complained about how 

little effect all his efforts made. At one time he 

wrote that only two members of the 

congregation found favour in his eyes. He also 

took eager part in the public religious debate, 

e.g. he advocated that sundays should be free 

from duties, giving ordinary people time to 

deal with something else. His work with the 

congregation was impeded by the activities of 

several sects, lay preachers, and strong 

political turbulences. 

Branth did not belong to any of the 

normal theological schools. He had his o\tn 

strong opinions and he did not always promote 

his views with diplomary. His personal 

thinking combined with a sharp and perhaps 

patronizing style was not always welcome for a 

religious debate, and in the end, only the

GRAPHIS SCRTPTA 4 (re92) 

Swedish journal "Facklan" (The torch) would 

print his straightforward extreme letters. 

Branth was not intolerant to people of 

another religious opinion but he detested 

halfteartedness and ignorance. When he 

retired he became friends with the catholic 

priest at Kolding. Neither Mgller nor Branth 

was satisfied with the priesthood. Branth 

preferred the company of scientists as the 

clergymen were characterized by unctuous 

powerlessness and unmanliness, afraid of 

voting, and dared not to oppose the rural 

deans or bishops. Branth called his bishop a 

nonentity and a frog blowing himself up to the 

size of an ox. 

The bishops went on episcopal visitations 

and reported to the ministry of culture. About 

Branth they wrote e.g. "numerous peculiarities 

and oddities", "his exterior endowments are 

very small, but he preaches christianly, 

carefully and comprehensible", "not without 

oddities and sickly critisism", "his character is 

open and determined, his conduct honest". 

Branth as a scientist 

As a student Branth also followed lectures 

of botany. At that time he called his scientific 

interest allotrial, that means irrelevant 

occupation, which could disturbe his 

theological studies. Later he claimed to be a 

scientist which had become a vicar. Time gave 

him right. His contribution to lichenology is of 

lasting value while his participation in the 

theological debate long since has been 

forgotten. His scientific approach to 

theological matters sometimes brought him 

into fruitless disputes. 

While being headmaster he wrote a 

textbook on physics, which was used for 40 

years. During his time in north Jutland he 

became interested in geology. Branth studied 

the layering of sand and clay in the pits of the 

area, and together with a professor of geology 

from Oslo he studied ttre origin bf tfie 

icetransported moraine-blocks of north 

Jutland. In this period he also published about 

the native pine forest of the island La;sg, 

which had then disappeared. Branth found 

Deichmann Branth 4l 

pine-cones in a turf pit in north Jutland, and 

concluded, that the pine-forest would have 

had a greater distribution in ealier times. After 

his retirement, Branth studied the flora of the 

ruined castle of Koldinghus. 

Branth first published about lichens in 

1860. In the beginning he had his 

identifications checked by Nylander and Th. 

Fries. In 1866 he was found worthy to receive 

Copenhagen University's golden medal for a 

paper on lichenology, as was also Emil 

Rostrup, Bt that time teacher at a training 

college. But since they both had got 

permanent jobs at that time, they could not 

receive the medal according to the rules. The 

two decided to combined their papers into the 

first Danish lichen flora: Lichenes Daniae 

eller Danmarks laver, published L869. 

Branth's concept of species was much broader 

than Rostrup's, but Rostrup accepted Branth's 

view, and Rostrup's contributed only with 

illustrations to the flora. Only six species af 

Cladonia and one of Ramalina was accepted, 

all of them having numerous varieties and 

forms. The work had a modern ta

42 Vagn Alstrup 

time. Normally he did not get a salary for his 

botanical work. Branth wrote that it should 

probably be seen as an honour to be allowed 

to work for the Botanical Museum, and he 

concluded that the more he worked the 

greater his debt of honour would be. In 1898 

he asked (squeezed with his own word) the 

Botanical Museum for a salary, since for Lz 

years he had identified all lichens collected in 

Denmark, Iceland, Greenland, and elsewhere 

by Danish collectors, which had taken 

inexcusable long time. He got 500 kr. once for 

all. Branth mentioned that it was about half of 

the amount a worker would have earned on a 

factory during the same time and found the 

small amount to be an example of "the 

parliament's dirty stinginess". 

Finally Branth was honoured in 1908 with 

the cross of order of a knighthood (Ridder af 

Dannebrog) for his botanical work after 

recommendation of Prof. Eug. Warming. Most 

vicars were honoured this wolr when they 

retired, and that would give their eventual 

widow a small pension, but Branth was not on 

good terms with his last bishop, who did not 

nominate him for this honour. 

Branth's view of Kopernikus and Darwin 

The apparent schism between theology and 

science was one of the most hot questions of 

the period. As a student Mgller became 

interested in astronomy, but he got scruples, 

as he found the theories of Kopernikus to 

interfere with the theological picture of the 

universe. Branth, on the contrary, was of the 

opinion, that Kopernikus was not in contrast 

to the Revelation. The case was different 

when it came to Danpin. Branth never 

accepted "the ape theory", which did not 

influence his taronomic view. Branth found, 

that Kopernikus' ideas were empirical founded 

on observations, while Danvin's were pure 

speculation. Mgller had some sympathy with 

Daruvin's ideas, but, apparently in a strong 

religious moment, he renunciated them. "It is 

a lewd stink coming from the abyss, in which 

the Darwinists, the Literature Society and the 

Socialists waltz around each to their own 

GRAPHTS SCRTPTA 4 (r9e2) 

melody; all this ugly fuss, resembling science 

as the ape resembles man, is no doubt only the 

toes and claws of the animal from Jordan" 

Mgller wrote. He admitted that he had 

tendencies towards free-thinking, and 

therefore he would not listen to such a 

eloquent person as John Stuart Mill. Branth 

never admitted to have such doubts. 

J. S. Deichmann Branth's lichenological 

publications: 

1860: l-avarterne. Tidsslvift for populere 

Fremstillinger af N aturvidenskabente 2 (2) : 

339-3U. 

1867: I-avernes udbredelse i den nordlige del 

af Jylland. Bot. Tidsslcr. 2: 68-88. 

- & Rostrup, E. 1869: Lichenes Daniae eller 

Danmarks laver. Bot. Tidsskr. 3: 127-284, 

t.lII-IV. 

L872: Collemaceae. Lichinaceae. Epiconiaceae 

. In: lange, J., Oversigt over sieldne 

arter. Bot. Tidssl


1908: Lichens. In: Duc d'Orl6ans, CroisiCre 

ocdanographique ... de la Belgtca 1905. 

Botanique 13. 

1909: Soredium, I-epra, Isidium. Bot. Tidsskr. 

29: L6-1,69 

References 

Deichmann Branth 43 

Christensen, C. 1924-1926: Den Danske 

BotaniJcs Historie I + II. Kgbenhavn. 884 

+ 690 pp. 

NOrr, E. 1986: Mellem Oao og facob I + II. 

Selskabet for Kirkehistorie, Gads Forlag. 

Kpbenhavn. 573 pp.

44 GRAPHTS SCRIPTA 4 (r9e2) 

Ove Almborn

Ett liv iignat frt lavar - Ove Almborn in memoriam 

Ove Almborn foddes den 30 juli L91,4 i Ronneby 

dir han ocksi viixte upp. Efter studentexamen 

i Karlskrona piborjade han 1933 sina 

akademiska studier vid Lunds universitet. De 

ledde fram till en doktorsavhandling i botanik, 

1948. Den behandlade en grupp sydsvenska 

lavars utbredning och ekologi. Det viildokumenterade 

materialet utg6r nu niira 50 6r 

senare grunden f6r ett nytt och st6rre forskningsprojekt 

over hotade lavar i sodra Sverige. 

Ove Almborn anstiilldes redan hosten 

L936 som amanuens i systematisk botanik och 

med undantag for ett par perioder da han 

tjiinstgjorde som gymnasieliirare blev tran 6nda 

fram till sin dod institutionen trogen, f6rst som 

docent och frin L966 som L:e museiintendent. 

Inte ens under sin tid som lektor vid 

d6varande Malm6 h6gre allmdnna liroverk, 

L956-L966, sliippte han greppet om lavarna. 

Han iignade de ail sin lediga tid inklusive 

helger och lov 6t att bygga upp ett omfattande 

lavherbarium. Pa 3O-talet de han kom till 

institutionen utgjordes detta av ett litet antal 

ganska d6ligt skotta samlingar. Idag 5r det en 

av de storsta och biist ordnade lavsamlingarna 

i viirlden. 

Ove foljde ocksi den tradition som redan 

under L9?}-talet funnits vid institutionen att 

forska kring sbdra Afrikas rika flora. Det var 

hans ambition att en ging ffl se projektet om 

en lavflora over sodra Afrika i hamn, men 

politiskt morker och oro som forhindrade stdd 

fr6n forskningsrid och andra forskare gjorde 

att det fick drivas pi sparl8ga. Sedan han tilltriitt 

tjiinsten som 1:e museiintendent fick han 

ocksi mindre tid till egen forskning. Museisamlingarna 

skotte han med stor pliktkiinsla 

och vid sin pensionering 1980 efterliimnade 

han ett mycket viilorganiserat herbarium. 

Ove Almborns vetenskapliga produktion 

omfattar ca 100 publikationer frin det fdrsta 

arbetet over l-avarna pi Lunds domkyrka 

publicerat i Skdnes Natur (1935) till det sista 

som han sjiilv aldrig fick se i tryck, Some 

overlooked or misidentified species of Teloschistes 

from South America and a key to the 

South American species, i Nordic lournal of 

Botany 12: 351-354. Den mest aktiva perioden 

var under 40- och 5O-talen di hans wA 

stdrsta arbeten kom ut, avhandlingen Distribution 

and Ecology of Some South 

Scandinavian Lichens, Botaniska Notiser, 

Supplement l:2 (1948) samt lawegetation och 

lavflora pA Hallands V6der6, Ktngliga Svenska 

Vetenskapsakademiens Avhandlingar i 

Naturslqddstirenden 11 (L955). I anslutning till 

dessa bida arbeten publicerades 6ven artiklar 

over sterila barklavar samt 6ver sltiktet Pertusaia. 

1956 kom den forsta av tillsammans 6 

fasciklar ut av exiccatet Lichenes Aficani och 

under samma 6r publicerades The South 

African Species of Xanthoria i Botaniska 

Notiser. Under 60-talet publicerades flera 

artiklar som behandlade sddra Afrikas lavflora 

bl.a. Studies in the lichen family Teloschistaceae 

fr6n 1963 samt Revision of some lichen 

genera in Southern Africa I frin 196F., bida 

publicerade i Botaniska Notiser. Fascikel 2 och 

3 av av Lichenes Africani kom ut 1960 och 

1965. 

Ove Almborn var inte bara en engagerad 

lavforskare och museichef. Han hade ocksA ett 

genuint intresse for botanisk litteratur, sirskilt 

Sldre sidan. Han 6gde sj6lv och var mycket 

stolt 6ver en niistan fullstiindig samling av 

Linneana. Hans omfattande bibliotek av 

lavlitteratur dr ett av de tre forniimsta i 

viirlden. Han stiillde vtilvilligt detta bibliotek 

till forfogande for nya generationer av lavforskare 

vid institutionen och var internationellt 

mycket anlitad i bibliografiska sammanhang. 

Ove Almborns litteraturintresse 

Aterspeglas dven i det stora antal vetenskapliga 

recensioner, jag har riiknat till 63, som publicerades 

under irens lopp, den fdrsta over 

Motykas Usnea-monografi i Botaniska

46 Ingvar KArnefelt 

Notiser (1940) till det sista 6ver monografin av 

Pyrenulaceae av Aptroot i Taxon (L992). 

Ove skaffade sig tidigt ett stort kontaktnit 

frdmst genom en omfattande korrespondens 

och han blev med tiden en internationellt 

mycket viilkSnd botanist. Han deltog flitigt i 

botaniska kongresser. I dessa sammanhang 

levde han verkligen upp och pA kongressmiddagar 

brukade han ofta forhoja stiimningen 

med goda historier ur sin stora repertoar om 

svunna tiders botanister. Ove Almborns stora 

personintresse resulterade dven i ett antal 

biografier bide utgivna som minnesteckningar 

samt i Svenskt Biografukt Lexikon dir han var 

en medarbetare sedan ett antal 6r tillbaka. 

Under 70- och 80-talen sammarbetade Ove 

dven flitigt med Frarz Stafleu i hans stora 

projekt Taxonomic Literature. 

Ove Almborns sista 6r blev inte siirskilt 

lyckliga. Han kom ofta in till mig och anf6r- 

trodde mig saker som var sv6rt att tala om. 

Han led bide ffsiskt och mentalt, dels av svflr 

viirk i ben och rygg, men friimst av Sngest och 

oro for ensamhet och morker. Periodvis f6ll 

han in i djupa depressioner och han var 

mycket sv6r att fe kontakt med under dessa 

tider. Vi talade annars nlstan alltid med 

varandra varje dag. Det hade blivit en vana 

sedan mlnga 5r, siirskilt sedan 1980 de han 

efter sin pensionering frf,n museitjiinsten flyttade 

upp till mig vid lavherbariet. Han pl6gades 

sSrskilt av flera oavslutade projekt, uDgdomsambitioner 

som de flesta av oss biir som 

ett ok genom livet. Tiden, moderna metoder, 

andra stora internationella projekt, hade 

hunnit i fatt honom, som i fallen med 

Pertusaria och The Lichen Flora of Southern 

Afica. Det hjiilpte f6ga att jag talade om f6r 

honom att minga andra personer hade rikat 

ut for samma problem, eller for att anviinda 

Oves egna ord "satt i samma bit". Trots dessa 

perioder av sv6rmod under de senare Sren vet 

jag emellertid att Ove siirskilt gladde sig 6t de 

yngre forskare vid institutionen som s6rjer for 

Sterviixten inom den del av botaniken som 

blev hans livsgtirning och som han sjiilv introducerat 

i Lund. 

GRAPHTS SCRTPTA 4 (L992) 

Pt morgonen den 6 mars 1992 fick jag det 

telefonmeddelande som jag se liinge mentalt 

hade varit forbredd p6. Vid begravningsakten 

den 23 mars var hans f6rste elev Hans Runemark 

officiant och sjiilv som hans andre elev, 

h6ll jag det tal som jag vet Ove hade Onskat 

nigon niira vln skulle hfllla. 

Du undrade ibland hur det skulle bli. Vem 

skulle grdta vid din bdr? For ndgra mdnader 

sedan tog jag farvtil av min far. Det var en 

smiirtsam upplevelse. Han var omgiven av sina 

barn in i det sbta. Du dog helt ensam. Vad iir 

det egentligen som gor ett mdnniskoliv, en viiv 

av kontakter och relationer som man sjiilv 

skapar. Du hade en rik sddan viiv, och du blev 

vida kiind i internationella sammanhang men 

iiven pd hemmaplan. Din itiaelika efyerltimnade 

korrespondens vittnar speciellt om detta. 

De iildsta arken har for ltingesedan borjat 

blelaru och mdnga av dessa viinner som 

formade din viiv finns inte heller liingre ibland 

o.r.r. For mig blev det niira 22 dr som vi var 

tillsammans, 22 dr med ldnga diskussioner om 

lavar, nomenklatur, typifteringat, artiklar eller 

personer. Iag fick ocl

GRAPHTS SCRTPTA 4 (re92) 

aldrig ha htnnat overge ditt tilskade Lund fhr 

ndgonting. Nu mdste jag ta farviil och iag 

tackar dig igen for alla de ldnga dren och ljusa 

dagarna och kanske jag trdffar dig igen ndgon 

gdng diir solen stdr hogt pd himmelen i 

Namaqualand, som vi sa niir compositerna 

Ove Almbom in memoriam 47 

blommar och diir buskama iir fulla med Teloschistaceer. 

Ingvar Kiimefelt, Institutionen fhr Systematisk 

Botanih Lunds (Jnivercitet, O. VoW. 18, 223 

61 Lund

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Manuslcripr. Manuskript siinds till redaktoren 

(Vagn Alstrup) och skrivs pi engelska eller ett 

skandinaviskt sprik med en kort engelsk 

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diskett, helst skriven i MS Word (IBM eller 

Macintosh) alternativt som ASCII-fiI. Bifoga 

alltid en utskrift. Engelska manuskript skall 

vara spr6kgranskade. 

Citerad litterarur. FOlj dessa exempel: 

Krog, H. 1991: Lichenological observations in 

low montane rainforests of eastern 

Tanzania. In: Galloway, D. J. (ed.), 

Tropical Lichens: Their systematics, 

conseryation and ecologt. The Systematics 

Association Special Volume 43: 85-94. 

Santesson, R. 1984: The lichens of Sweden 

and Norway. Stockholm & Uppsala. 


Hansen, E. S. I99L: The lichen flora near a 

lead-zinc mine at Maarmorilik in West 

Greenland. Lichenologist 23 : 381 -391. 

Siirtryck Forfattaren/forfattarna erhiller gratis 

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The manuscripts should be type-written and 

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If possible, send also the manuscript on a 

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GRAPHIS SCnIPTA 

Volym 4, hiifte 7,1992 

Innehfrll 

1 Lichens from the coast of Eniseiskii Gulf, W. Taimyr, Siberia 

M. P. Zhurbenko 

5 Anteckningar om nigra arter av Peltigera canina-gruppen i Sverige 

[Notes on the Swedish species of the Peltigera canina group] 

G. Carlin 

18 Boganmeldelse (Introduktion til svampe) 

18 Exkursion till Visings6 

L9 The 9th meeting of the Nordic Lichen Society, Sweden 1991 

G. Thor 

30 Notes on the genus Caloplaca in the Siberian arctic 

U. SOthting, M. Zhurbenlco & E. S. Hansen 

33 Protothelenella rylina new to Finland 

P. P. G. van den Boom 

35 Caloplaca soropelta (E. S. Hansen, Poelt & S6chting) Sochting comb. nov. 

U. Sqchting 

37 Boganmeldelse (Toninia) 

39 Deichmann Branth - A portrait based mainly on his letters 

V.Alstrup 

44 Ett liv 6gnat it lavar - Ove Almborn in memoriam 

I. Kiimefelt

GRAPHIS ScnIPTA - Universitetet i Oslo

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