Family 208. BEGONIACEAE 1. Begonia Linnaeus CAMIGUIN ...

Family 208. BEGONIACEAELiterature:Smith, LB, DC Wasshausen, J Golding & CE Karegeannes. 1986. Begoniaceae. Part. I:Illustrated Key; Part II. Annotated species list. Smithsonian Contributions to Botany no.60: 1-584.Golding, J & DC Wasshausen. 2002. Begoniaceae, Edition 2. Part. I: Annotated species list; PartII.Illustrated key, Abridgement and supplement. Contributions from the United StatesNational Herbarium 43: 1-289.1. Begonia LinnaeusBegonia acuminatissima Merr., PJS 6 c (1911) Bot. 395; EPFP 3 (1923) 119.CAMIGUIN, MINDANAO: Agusan del Norte (Mt Urdaneta), BALUT. Lowland primaryforests and stream embankments up to 400m.Begonia aequata A Gray, Bot. Wilkes US Explor. Exped. (1854) 658; --Merr., EPFP 3 (1923) 119.LUZON: Laguna (Mt Makiling). In primary forests, 400-1000m, climbing on tree trunks.Locally common.Begonia affinis Merr., PJS 7 c (1912) Bot. 308; EPFP 3 (1923) 119.MINDANAO: Zamboanga. Damp forests, c. 900m.Begonia agusanensis Merr., PJS 6 c (1911) Bot. 377; EPFP 3 (1923) 119. --Type: BS 7312 Merrill(US*, iso). Mindanao: Agusan del Norte prov., Agusan River near Waloe, on banks along smallstreams in forests, altitude c. 50m, 2 Oct 1910.MINDANAO: Agusan del Norte, Agusan del Sur (Talacogon), Bukidnon. Along smallstreams in lowland primary forests.Begonia alba Merr., PJS 10 c (1915) Bot. 45; EPFP 3 (1923) 120. --Type: BS 19963 McGregor(US*, iso). Luzon: Ifugao prov., Feb. 1913.LUZON: Ifugao. Probably in forests above 1000m.Begonia alvarezii Merr., PJS 6 c (1911) Bot. 405; EPFP 3 (1923) 120. –Type: FB 24446 Alvarez (notseen). Luzon: Nueva Ecija prov., Santor River, Feb 1911.LUZON: Apayao, Nueva Ecija, Bulacan (Angat), Quezon (Mt Pular). Low and mediumaltitude forest stream embankments.Begonia androturba Coyle, Edinburgh J. Bot. 66(2) (2009) 206-209, Fig. 2, map. –Type: PPI 38072Pipoly, Romero et al (BRIT*, holo). Palawan: Palawan prov., Mt Gantung, district of Tindogan,NNW slopes at headwaters of Tagwaya River, 8 59.0’N, 117 49.0’E, steep slopes frequentlysubjected to large landslides, on ultramafic soil, altitude 800m, 19 Aug 1996.PALAWAN: Mt Gantung. Known only from the type locality.Begonia angilogensis Merr., PJS 26 (1925) 477; EPFP 4 (1925) 250. --Type: BS 40777 Ramos(B*, K*, US*, iso). Luzon: Rizal prov., Mt Angilog, forested slopes, altitude 1200m, Apr 1922.LUZON: Rizal (Mt Angilog). Known only from the type collection.Begonia anisoptera Merr., PJS 6 c (1911) Bot. 398; EPFP 3 (1923) 120. --Type: BS 5482 Merrillravines, altitude less than 100m, 10 Oct 1906.MINDANAO: Zamboanga. Semi-shaded cliffs and boulders in low altitude forests.(NY*, iso; US*, isosynty

Begonia apayaoensis Merr., PJS 13 c (1918) Bot. 39; EPFP 3 (1923) 120. --Type: BS 28403 EFenix (US*, iso). Luzon: Apayao prov., Mt. Sulu, May 1917.LUZON: Apayao. Rocky shaded slopes, c. 1000m.Begonia biliranensis Merr., PJS 10 c (1915) Bot. 46; EPFP 3 (1923) 120. --Type: BS 18822McGregor (US*, iso). Biliran, Jun 1914.BILIRAN, SAMAR. Epilithic in low and medium altitude forests.Begonia binuangensis Merr., PJS 13 c (1918) Bot. 40; EPFP 3 (1923) 120. --Type: BS 28813Ramos & Edaño (US*, iso). Luzon: Quezon prov., Mt. Binuang, May 1917.LUZON: Quezon, Sorsogon (Mt Bulusan). Climbing on tree trunks in primary forests atmedium altitudes.Begonia bolsteri Merr., PJS 6 c (1911) Bot. 387; EPFP 3 (1923) 120. --Type: Bolster 310 (US*, iso).Mindanao: Surigao del Norte prov., Apr 1906.LUZON: Laguna, Sorsogon, BILIRAN, LEYTE, MINDANAO: Surigao. Along streams inprimary lowland forests.Begonia brevipes Merr., PJS 6 c (1911) Bot. 378; EPFP 3 (1923) 378. --Type: BS 7431 Ramos (US*,iso). Luzon: Cagayan prov., Pamplona, in forests, Mar 1909.LUZON: Cagayan, MINDANAO: Lanao. Lowland forests.Begonia calcicola Merr., PJS 6 c (1911) Bot. 400; EPFP 3 (1923) 120. --Type: BS 7062 Merrill (US*,iso). Luzon: Rizal prov., Jan 1910.LUZON: Rizal, Laguna, Batangas. Cliffs at low altitudes.Begonia casiguranensis Quisumb. & Merr., PJS 37 (1928) 172. --Type: BS 45277 Ramos &Edaño (NY*, US*, iso). Luzon: Aurora prov., Casiguran, Cabulig River, May-Jun 1925.LUZON: Aurora (Casiguran, Cabulig River). Along streams in open places at lowaltitudes.Begonia castilloi Merr., PJS 13 c (1918) Bot. 38; EPFP 3 (1923) 120. --Type: BS 22723 Castillo (US*,iso). Luzon: Cagayan prov., Apr-May 1915.LUZON: Cagayan. Streamside cliffs, c. 100m.Begonia caudata Merr., PJS 13 c (1918) Bot. 41; EPFP 3 (1923) 120. --Type: BS 28414 E Fenix(NY*, US*, iso). Luzon: Apayao prov., Mt. Sulu, May 1917.LUZON: Apayao. Rocky forested slopes, climbing on trees.Begonia chloroneura P Wilkie & Sands, New Plantsman 6 (1999) 132, fig. –Type: P Wilkie 29015et al. (PNH, holo; iso: K, E*, 2 sheets). Luzon: Isabela prov., San Mariano municip., Barangay San Jose,vertical rock face in crack, 6 Mar 1997.LUZON: Isabela (San Mariano).Begonia ciliifera Merr., PJS 6 c (1911) Bot. 376; EPFP 3 (1923) 121. --Type: FB 9318 Whitford &Hutchinson (US*, iso). Mindanao: Zamboanga del Sur prov., Port Banga, in canyons, altitude 20m,Jan 1908.MINDANAO: Zamboanga del Sur. Damp shaded ravines up to 400m altitudes.Begonia collisiae Merr., PJS 14 (1919) 424; EPFP 3 (1923) 121. --Type: BS 31469 Ramos & Edaño(US*, iso). Panay: Aklan prov., Libacao, Apr-May 1918.PANAY: Aklan. Forested stream embankment, c. 100m.

Begonia colorata Warb. in Perkins, Frag. Fl. Philip. (1904) 51; --Merr., EPFP 3 (1923) 121. –Type:Warburg 14633MINDANAO: Davao. Or rocky bluffs and banks at medium altitudes; apparently close toB nigritarum Steud.Begonia contracta Warb. in Perkins, Frag. Fl. Philip. (1904) 54; --Merr., EPFP 3 (1923) 121. –Type:Warburg 13085 (B, holo). Luzon: Quezon prov., Sampaloc,LUZON: Ilocos Norte, Cagayan, Isabela, Bataan, Bulacan, Rizal, Quezon, Sorsogon,MINDORO, PANAY, NEGROS, LEYTE, SAMAR, MINDANAO: Zamboanga. Low andmedium altitude primary forests.Begonia copelandii Merr., PJS 6 c (1911) Bot. 401; EPFP 3 (1923) 121. --Type: Copeland 1255LUZON: Cagayan, PANAY, MINDANAO: Davao. On boulders along streams up to850m.Begonia coronensis Merr., PJS 26 (1925) 480; EPFP 4 (1925) 250. --Type: BS 41164 Ramos (US*,iso*). Coron Island, 17 Sept 1922.CORON. Lowland forests on limestone, on rocks and ledges along small streams.Begonia crispipila Elmer, LPB 2 (1910) 737; --Merr., EPFP 3 (1923) 121. --Type: Elmer 8687 (NY*, US*, iso). Luzon: BengLUZON: Apayao, Benguet. In damp shaded ravines, altitude up to 1500m.Begonia cumingiana (Klotz.) DC, Prodr. 15 1 (1864) 320; --Fern.-Villar, Novis. App. (1880) 98; --Vidal, Phan. Cuming. Philip. (1885) 116; Rev. Pl. Vasc. Filip. (1886) 143; --Merr., PJS 6 c (1911)Bot. 388; EPFP 3 (1923) 121; --Diploclinium cumingianum (Klotz.) Miq., Fl. Ind. Bat. 1, 1 (1857)691; --Petermannia cumingiana Klotz., Monatab. Berl. Akad. (1854) 124; Abh. Akad. Berl. (1855)75, t. 6, fig. C; --Walp., Ann. 4 (1857) 906. –Type: Cuming 856.LUZON: Albay, Sorsogon, CATANDUANES. Primary forests, usually along streams.Begonia cumingii A Gray, Bot. Wilkes US Explor. Exped. (1854) 658; --Merr., PJS 3 c (1908) Bot.84, t. 3; idem 6 c (1911) Bot. 384; --EPFP 3 (1923) 121. –Type:Begonia philippinensis A DC, Prodr. 15, 1 (1864) 320; --Fern.-Villar, Novis. App. (1880) 98; --Vidal, Phan.Cuming. Philip. (1885) 116; Rev. Pl. Vasc. Filiip. (1886) 143. –Type:LUZON: Laguna, Quezon. Primary forests, altitude 700-1400m.Begonia dolichotricha Merr., PJS 17 (1920) 292; EPFP 3 (1923) 122. --Type: BS 33926 Ramos &Edaño (US*, iso). Luzon: Camarines Sur prov., Mt. Bagacay, forested slopes, altitude c. 650m,18 Dec 1918.LUZON: Aurora (Mt Alzapan, BS 45673, Ramos & Edaño, B*), Camarines Sur (MtBagacay).Begonia edanoi Merr., PJS 13 c (1918) Bot. 314; EPFP 3 (1923) 122. --Type: BS 29374 Ramos &Edaño (NY*, US*, iso). Luzon: Rizal prov., Mt. Susong-Dalaga, Aug 1917.LUZON: Rizal (Mt Susong Dalaga; Mt Irid). Primary forests at medium altitudes, climbingon trees.Begonia elatostematoides Merr., PJS 7 c (1912) Bot. 309; EPFP 3 (1923) 122. --Type: BS 8232Merrill (US*, iso). Mindanao: Zamboanga prov., Nov-Dec 1911.MINDANAO: Zamboanga. Damp shaded ravines and forested slopes, 800-1000m.Begonia elmeri Merr., PJS 13 c (1918) Bot. 39; EPFP 3 (1923) 122; --Begonia peltata Elmer, LPB7 (1915) 2556, non A DC. –Type: Elmer 14183.

MINDANAO: Agusan del Norte (Mt Hilong-hilong). Cliffs in shaded ravines, altitude c.1050m.Begonia esculenta Merr., PJS 6 c (1911) Bot. 389; EPFP 3 (1923) 122. --Type: BS 9449 CBAug 1909.LUZON: Quezon (Mt Binuang). Mossy forests, c. 800m.Robinson (US*, iso). LuBegonia everettii Merr., PJS 6 c (1911) Bot. 390; EPFP 3 (1923) 122. –Type: FB 5587 Everett (notseen). Negros: Negros Occidental prov., Sicaba, Daluapan River, 12 Nov 1906.LUZON: Quezon, PANAY, NEGROS, SAMAR. Low altitude primary forests.Begonia fasciculiflora Merr., PJS 6 c (1911) Bot. 376; EPFP 3 (1923) 122.—Type: FB 9316Whitford & Hutchinson (not seen). Mindanao: Zamboanga del Sur prov., Port Banga, in canyons,altitude 5-30m, 13 Jan 1906.MINDANAO: Zamboanga del Sur. Forested ravines at low altitudes.Begonia fenicis Merr., PJS 3 c (1908) Bot. 421; EPFP 3 (1923) 122. --Type: BS 3619 E Fenix (US*,isosyntype). Batan: Sto Domingo de Basco, 30 May 1907.S Ryukyus, Taiwan (Lanyu) and the Philippines.BATAN, BABUYAN ISLS, BALABAC (Merrill 5375, B*), SAMAR (Merrill 5237bis, B*). Oncliffs and boulders at low altitudes.Begonia georgei Coyle, Edinburgh J. Bot. 66(2) (2009) 209-210, Fig. 2, map.Begonia gitingensis Elmer, LPB 2 (1910) 738; --Merr., PJS 6 c (1911) Bot. 401; EPFP 3 (1923) 122.--Type: Elmer 12368 (US*, NY*, iso). Sibuyan: Mt. Guiting-guiting, Apr 1910.SIBUYAN (Mt Giting-giting). Shaded ravines along small streams, c. 525m.Begonia gracilipes Merr., PJS 6 c (1911) Bot. 405; EPFP 3 (1923) 123. --Type: BS 7395 Ramos(US*, iso). Luzon: Cagayan prov., Claveria, on trees in forest, 13 Mar 1909.LUZON: Cagayan, CATANDUANES. Climbing on trees in damp primary forests at lowaltitudes.Begonia halconensis Merr., PJS 6 c (1911) Bot. 385; EPFP 3 (1923) 123. --Type: Merrill 5515 (NY*,US*, iso). Mindoro: Mindoro Oriental prov., Mt. Halcon, on steep, damp forested slopes,altitude 1800m, Nov 1906.MINDORO: Mindoro Oriental (Mt Halcon). Primary forests, altitude 1100-1800m.Begonia hernandioides Merr., PJS 6 c (1911) Bot. 392; EPFP 3 (1923) 123. --Type: BS 7393Ramos (US*, iso). Luzon: Cagayan prov., Claveria, on rocks along streams, Mar 1909.LUZON: Ilocos Norte (Bangui, BS 43571 McGregor, K*), Cagayan, Tayabas. On cliffsand bowlers in shaded ravines at low or medium altitudes.Begonia incisa DC, Ann. Sci. Nat. Bot. iv, 11 (1859) 129; Prodr. 15, 1 (1864) 321; --Fern.-Villar, Novis.App. (1880) 99; --Merr., PJS 6 c (1911) Bot. 382; EPFP 3 (1923) 123. –Type:LUZON: Zambales, Quezon, Camarines, Sorsogon, PANAY, NEGROS. On banks andcliffs in primary forests at low and medium altitudes.Begonia isabelensis Quisumb. & Merr., PJS 37 (1928) 173. --Type: BS 47207 Ramos & Edaño(NY*, US*, iso). Luzon: Isabela prov., San Mariano, on rocks along forested streams, 14 Mar 1926.LUZON: Isabela (San Mariano).

Begonia jagori Warb. in Perkins’ Frag. Fl. Philip. (1904) 54; --Merr., PJS 6 c (1911) Bot. 382; EPFP 3(1923) 123. –Syntypes: Jagor 889 (B*) and 890 (B*). Luzon: no further locality data but accordingto Merrill (1923) ‘probably Tayabas’ [=Quezon prov.]. Coll. 1861,LUZON: prob. Tayabas, MINDORO. In primary forests at low altitudes.Begonia klemmei Merr., PJS 6 c (1911) Bot. 402; EPFP 3 (1923) 123. --Type: FB 5677 KlemmeNov 1906.LUZON: Mountain Province, Benguet. On cliffs and boulders, altitude 1400-2000m.(US*, iso). Luzon: MouBegonia lacera Merr., PJS 10 c (1915) Bot. 49; EPFP 3 (1923) 123.—Type: BS 16111 ReilloBASILAN. Low altitude primary forests.Begonia lagunensis Elmer, LPB 2 (1910) 735; --Merr., PJS 6 c (1911) Bot. 406; EPFP 3 (1923) 123.--Type: Elmer 9327 (US*, NY*, syntypes). Luzon: Quezon prov., Lucban, May 1907.LUZON: Rizal, Laguna, Quezon, Camarines, MINDORO, NEGROS, SAMAR. Mediumaltitude primary forests, climbing on trees.Begonia lancifolia Merr., PJS 10 c (1915) Bot. 48; EPFP 3 (1923) 123. --Type: BS 16162 Reillo.BASILAN. Low altitude primary forests.Begonia lancilimba Merr., PJS 14 (1919) 424; EPFP 3 (1923) 123. --Type: BS 32232 McGregor(US*, iso). Panay: Antique prov., May-Aug 1918.PANAY: Antique, Mossy forests, altitude 900-1000m.Begonia latistipula Merr., PJS 10 c (1915) Bot. 51; EPFP 3 (1923) 123. --Type: BS 15367 Ramos(US*, iso). Leyte: Dagami, forest near the summit of Mt Buraui, Aug 1912.BILIRAN, LEYTE, SAMAR. Medium altitude primary forests.Begonia leptantha C Robinson, PJS 6 c (1911) Bot. 211; idem. 281; EPFP 3 (1923) 124. –Type: BS6857 CB Robinson (B*, NY*, US*, iso). Polillo, Aug 1909.LUZON: Quezon (Mt Binuang; Mahabang Sugsugan River along Siniloan-Infanta Road),POLILLO. Lowland forests along streams.Begonia leucosticta Warb. in Perkins’ Frag. Fl. Philip. (1904) 55; --Merr., EPFP 3 (1923) 124. –Type:Warburg 12004 (B*, holo). Luzon: Isabela prov., Digamai [=Dicamay in present-day Jonesmunicipality].LUZON: Benguet, Isabela, BUCAS GRANDE. In thickets on cliffs, from near sea-level to1400m altitude,Begonia littleri Merr., PJS 6 c (1911) Bot. 379; EPFP 3 (1923) 124. –Type: DeVore & Hoover 92 (US*,iso). Basilan: Basilan prov., Isabela, along banks of streams at an altitude of c. 90m, Apr 1903.Note: Although the type number is cited in the protologue as ‘DeVore & Hoover 94’, LB Smith (inannot. on the US sheet cited above) believes this to be a typographical error.BASILAN, MINDANAO: Zamboanga. Lowland forests along streams.Begonia loheri Merr., PJS 6 c (1911) Bot. 382; EPFP 3 (1923) 124. –Syntypes: Loher 6090 (B*, US*)and 6098 (K*). Luzon: Rizal prov., Angilog, 15 Mar 1906LUZON: Rizal (Angilog). Apparently known only from a few collections at the typelocality.Begonia longibractea Merr., PJS 17 (1920) 293; EPFP 3 (1923) 124 (as ‘longibracteata’). --Type:BS 34870 Ramos & Pascasio (US*, iso). Siargao. Jun 1919.SIARGAO. Forested slopes at low altitudes.

Begonia longinoda Merr., PJS 6 c (1911) Bot. 397; EPFP 3 (1923) 124. –Type: BS 13372 Ramos.Luzon: Quezon prov. Tagkawayan, c. 200m, Mar 1911.LUZON: Quezon (Tagkawayan). Lowland primary forests, c. 200m.Begonia longiscapa Warb. in Perkins’ Frag. Fl. Philip. (1904) 52; --Merr., EPFP 3 (1923) 124. –Type:Begonia longistipula Merr., PJS 6 c (1911) Bot. 379; EPFP 3 (1923) 124. –Type: Bolster 248 (notseen). Mindanao: Surigao del Norte prov., near Surigao, altitude c. 125m, Feb and Apr 1906.SAMAR, MINDANAO: Surigao del Norte. Low altitude primary forests.Begonia longivillosa DC, Ann. Sci. Nat. Bot. iv, 11 (1859) 130; --Merr., EPFP 3 (1923) 124. –Type:Begonia luzonensis Warb. in Perkins’ Frag. Fl. Philip. (1904) 52; --Merr., EPFP 3 (1923) 124.Begonia bakeri Elmer, non DC, –Type:Begonia macgregorii Merr., PJS 7 c (1912) Bot. 310; EPFP 3 (1923) 124. --Type: BS 11334 RCMcGregor (US*, iso). Luzon: Nueva Vizcaya prov., Dupax, Mar-Apr 1912.LUZON: Nueva Vizcaya. Along small streams in primary forests at medium altitudes.Begonia malindangensis Merr., PJS 6 c (1911) Bot. 391; EPFP 3 (1923) 125. --Type: FB 4563Mearns & Hutchinson (NY*, US*, iso). Mindanao: Misamis Occidental prov., Mt. Malindang, inforest, altitude c. 1800m, May 1906.MINDANAO: Misamis Occidental (Mt Malindang), Bukidnon, Davao. Forests, altitude c.1800m.Begonia manillensis DC, Ann. Sci. Nat. Bot. iv, 11 (1859) 129; --Merr., EPFP 3 (1923) 125. –Type:Begonia mearnsii Merr., PJS 6 c (1911) Bot. 383; EPFP 3 (1923) 125. –Type:Begonia megacarpa Merr., PJS 9 c (1914) Bot. 378; EPFP 3 (1923) 125. –Type:LUZON: Nueva Ecija, Camarines, LEYTE, MINDANAO: Agusan. Climbing on trees inprimary forests at low and medium altitudes.Begonia magalantha Merr., PJS 10 c (1915) Bot. 47; EPFP 3 (1923) 125. –Type:Begonia merrittii Merr., PJS 5 c (1910) Bot. 365; EPFP 3 (1923) 125. --Type: Elmer 6254 (US*, iso).Luzon: Benguet prov., Mt Sto Tomas, May 1904.LUZON: Mountain Province, Benguet (Pauai; Mt Pulag; Mt Lusod; Mt Sto Tomas; MtUgo). Mossy forests above the pine regions on the higher mountains, altitude 1500-2400m. Common.Begonia mindanaensis Warb. in Perkins’ Frag. Fl. Philip. (1904) 55; --Merr., PJS 6 c (1911) Bot.376; EPFP 3 (1923) 125. –Type: Warburg 14633 (B, holo). Mindanao: Davao del Sur prov., MtDagatpan (near Mt Apo), Jun 1888.LEYTE, MINDANAO: Zamboanga, Lanao, Bukidnon, Davao del Sur, Agusan del Norte,Surigao del Norte, SIARGAO. Low and medium altitude primary forests, apparentlycommon.

Begonia mindorensis Merr., PJS 6 c (1911) Bot. 396; EPFP 3 (1923) 125. –Type:Begonia negrosensis Elmer, LPB 2 (1910) 736; --Merr., PJS 6 c (1911) Bot. 385; EPFP 3 (1923)125. --Type: Elmer 9903 (NY*, MO*, US*, iso). Negros: Negros Oriental prov., Cuernos Mtns., Jun1908 (Apr 1908 according to protologue).LUZON: Pampanga (Mt Pinatubo, Elmer 21492, K*), PANAY: Aklan (Libacao),NEGROS: Negros Occidental (Mt Kanlaon), Negros Oriental (Cuernos Mtns). Forests upto 1200m.Begonia neopurpurea LB Sm. & DC Wasshausen, Phytologia 52 (1983) 445.Begonia purpurea Elmer, LPB 10 (1939) 3707, non Sw., 1788. --Type: Elmer 16565 (NY*, iso). Luzon:Sorsogon prov., Mt. Bulusan, Jul 1916.Begonia nigritarum Steudel, Nomencl. (1821) 104; --Merr., EPFP 3 (1923) 126.Begonia oblongata Merr., PJS 7 c (1912) Bot. 310; EPFP 3 (1923) 126. --Type: Merrill 8166 (US*,iso). Mindanao: Zamboanga District, Nov-Dec 1911.Begonia obtusifolia Merr., PJS 14 (1919) 425; EPFP 3 (1923) 126. --Type: BS 30803 Ramos &Edaño (US*, iso). Panay: Capiz prov., Mt. Macosolon.PANAY: Capiz. On boulders in damp forests, c. 150m.Begonia oligantha Merr., PJS 10 c (1915) Bot. 50; EPFP 3 (1923) 126. Type: BS 15733 E Fenix(US*, iso). Mindanao: Bukidnon prov.Begonia oxysperma DC, Ann. Sci. Nat. Bot. iv, 11 (1859) 122; --Merr., EPFP 3 (1923) 126. –Type:LUZON: Mountain Province, Ifugao (Mt Polis), Benguet (Baguio; Mt Sto. Tomas), Rizal,Quezon (Mt Banahaw & San Cristobal), Laguna (Mt Makiling). Primary forests, on treesor cliffs, 400-1900m.Begonia palawanensis Merr., PJS 6 c (1911) Bot. 380; EPFP 3 (1923) 126. --Type: BS 7232 Merrill(US*, L*, Iso). Palawan (west coast): Puerto Princesa, Napsahan, altitude c. 4m, 19 Sept 1910.PALAWAN. Along small streams in low altitude primary forests.Begonia panayensis Merr., PJS 14 (1919) 428; EPFP 3 (1923) 127. --Type: BS 32309 RC McGregor(US*, iso). Panay: Antique prov., Culasi, May-Aug 1918.PANAY: Antique. Along shaded streams, c. 250m.Begonia parva Merr., PJS 6 c (1911) Bot. 402; EPFP 3 (1923) 127. --Type: BS 5551 Ramos (US*,iso). Luzon: Benguet prov., Rio Trinidad (=Balili River), Dec 1908.LUZON: Benguet (La Trinidad, along Balili River). On stream boulders, c. 1400m.Begonia parvilimba Merr., PJS 26 (1925) 481; EPFP 4 (1925) 250. --Type: BS 36936 Ramos &Edaño (US*, iso). Mindanao: Zamboanga del Sur prov., Malangas, Oct-Nov 1919.Begonia perryae LB Sm. & DC Wasshausen, Phytologia 52 (1983) 445; --Begonia robinsoniiMerr., PJS 6 c (1911) Bot. 375, non Ridley, 1909. –Type: BS 6340 Robinson (not seen). Luzon:Camarines Sur prov., Maagnas, 28 Aug 1908.

LUZON: Camarines Sur. Low altitude forests.Begonia pinamalayensis Merr., PJS 26 (1925) 482; EPFP 4 (1925) 250. --Type: BS 40856 Ramos(US), Mindoro: Mindoro Oriental prov., Pinamalayan.MINDORO. Along forest streams.Begonia platyphylla Merr., PJS 10 c (1915) Bot. 46; EPFP 3 (1923) 127. --Type: BS 20074 RCMcGregor (US*, iso). LBegonia polilloensis Tebbitt, Edinburgh J. Bot. 61(2-3) (2004, publ. 12 Aug 2005) 99-101, Fig. 2. –Type: PNH 42651 (L, holo).POLILLO, NEGROSBegonia pseudolateralis Warb. in Perkins’ Frag. Fl. Philip. (1904) 51; --Merr., PJS 2 c (1907) Bot.285; idem. 6 c (1911) Bot. 374; EPFP 3 (1923) 127. –Type:Philippines, Sulawesi.LUZON: Cagayan to Sorsogon, POLILLO, MINDORO, PANAY, LEYTE, CAMIGUIN,JOLO, BASILAN, MINDANAO. Low and medium altitude primary forests, ascending to1200m.Begonia quercifolia DC, Ann. Sci. Nat. Bot. iv, 11 (1859) 129; --Merr., EPFP 3 (1923) 127. –Type:Cuming 1696 (K*, 2 sheets, holo and iso not specified). Samar, coll. 1841.Begonia leytensis Elmer, LPB 2 (1910) 730; --Merr., PJS 6 c (1911) Bot. 384. –Type: Elmer 7255 (K*, MO*,NY*, US*, iso). Leyte: Leyte prov., Palo, Jan 1906.LEYTE (Palo), SAMAR (Catubig River). Lowland forests along streams.Begonia ramosii Merr., PJS 6 c (1911) Bot. 388; EPFP 3 (1923) 127. –Type: BS 10942 Ramos (notseen). Luzon: Laguna prov., San Antonio, in forest, Aug 1910. Additional materials seen BS 10941Ramos (K*).LUZON: Laguna (San Antonio; Paete). Low altitude damp primary forest.Begonia rizalensis Merr., PJS 6 c (1911) Bot. 388; EPFP 3 (1923) 127. --Type: BS 12109 RamosLUZON: Rizal (Montalban; San Isidro). In primary forests, probably at médium altitudes.(US*, iso). Luzon: RizaBegonia rubrifolia Merr., PJS 14 (1919) 426; EPFP 3 (1923) 127. –Type: BS 32430 RC McGregor(B*, iso). Panay: Antique prov., Culasi, alt. c. 1000m, 7 Jun 1918.PANAY: Antique (Culasi). Forests, c. 1000m.Begonia rufipila Merr., PJS 6 c (1911) Bot. 393; EPFP 3 (1923) 127. --Type: FB 5665 Klemme(US*, iso). Luzon: Ilocos Sur prov., Dolores (NB: this is now part of the province of Abra), 31Oct 1906.LUZON: Ilocos Range along W Abra. Damp limestone boulders in shaded places, c.200m.Begonia samarensis Merr., PJS 30 (1926) 413. --Type: BS 43757 McGregor (NY*, US*, iso) Samar:Western Samar prov., Loquilocon, hillside forests, c. 250m. Jun 1924.SAMAR (Loquilocon). Low altitude forests on limestone.Begonia sarmentosa LB Sm. & DC Wasshausen, Phytologia 52 (1983) 443; --Begonia elegansElmer, LPB 7 (1915) 2554; --Merr., EPFP 3 (1923) 122, non Humboldt, Bonpland & Kunth (1825).--Type: Elmer 13672 (iso in MO*, NY (2 sheets)*, US*). Mindanao: Agusan del Norte prov.,Cabadbaran, Mt Urdaneta, Cawilanan, alt. 4250 ft [=1295m], Sept 1912.MINDANAO: Agusan del Norte (Mt Urdaneta). Forested ravines, alt. 1250m. Probablynot distinct from the very close B. aequata A. Gray of Luzon.

Begonia serpens Merr., PJS 14 (1919) 427; EPFP 3 (1923) 128. --Type: BS 32588 RC McGregor(US, iso). Panay: Antique prov., May-Aug 1918.Begonia sorsogonensis Elmer ex Merr., EPFP 3 (1923) 121. –Type:LUZON: SorsogonBegonia suborbiculata Merr., PJS 6 c (1911) Bot. 398; EPFP 3 (1923) 128. --Type: Merrill 7229(US*, iso). Palawan, Sept 1910.Begonia subprostrata Merr., PJS 26 (1925) 478; EPFP 4 (1925) 250. --Type: BS 36977 Ramos& Edaño (US*, iso). Mindanao: Zamboanga del Sur prov., Malangas, Oct-Nov 1919.Begonia subtruncata Merr., PJS 6 c (1911) Bot. 390; EPFP 3 (1923) 128. --Type: Loher 6076 (notseen). Luzon: La Union prov., Castilla, Mar 1906.LUZON: La Union. In forests.Begonia tayabensis Merr., PJS 13 c (1918) Bot. 38; EPFP 3 (1923) 128. --Type: BS 29054 Ramos &Edaño (NY*, US*, iso). Luzon: Tayabas prov., May-Jun 1917.Begonia trichochila Warb. in Perkins’ Frag. Fl. Philip. (1904) 53; --Merr., EPFP 3 (1923) 128. –Type:Begonia urdanetensis Elmer, LPB 7 (1915) 2559; --Merr., EPFP 3 (1923) 128. --Type: Elmer 13819(NY*, MO*, US*, iso). Mindanao: Agusan del Norte prov., Mt. Urdaneta (=Mt Hilong-hilong), Sept1912.MINDANAO: Agusan del Norte. Primary forests, c. 1500m.Begonia vanoverberghii Merr., PJS 6 c (1911) Bot. 403; EPFP 3 (1923) 128. –Type:Begonia wadei Merr. & Quisumb., Addisonia 17 (1932) 57, pl. 573. --Type: BS 78301 WH Brown(NY*, iso). Coron Island, Dec 1929.CORON. Exposed coastal coralline limestone, which at times may be subject to saltspray. Known only from the type locality.Begonia weberi Merr., PJS 6 c (1911) Bot. 381; EPFP 3 (1923) 128. –Type: Weber 1210 (not seen).Mindanao: Agusan del Norte prov., Mt Hilong-hilong, rocky slopes in ravines, c. 450m, Mar 1911.MINDANAO: Agusan del Norte (Mt Hilong-hilong). Rocky slopes in forested ravines, c.450m.Begonia wenzelii Merr., PJS 10 c (1915) Bot. 277; EPFP 3 (1923) 128. –Type:Begonia wilkiei C Coyle,--Type: C Coyle sn (E, holo). PalawanBegonia woodii Merr., PJS 26 (1925) 478; EPFP 4 (1925) 250. --Type: Merrill 11589 (US). Palawan:Malampaya Bay (Sound).PALAWAN. Primary forests on cliffs.Begonia zamboangensis Merr., PJS 26 (1925) 481; EPFP 4 (1925) 250. --Type: BS 37371 Ramos& Edaño (US, iso). Mindanao: Zamboanga del Sur prov., Malangas, Nov 1919.MINDANAO: Zamboanga del Sur (Malangas). Forested small stream banks at low

altitudes.Dubious taxon.Begonia porteana van Geert, Cat. (1881) 90, descriptione inchoata; --LB Sm. et al., Smiths. Contr.Bot. no. 60 (1986) 218.

Family 208. BEGONIACEAE 1. Begonia Linnaeus CAMIGUIN ...

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