The genus Pyrenopeziza Fuck. - ASCOfrance

TAXONOMICAL NOTES ON MOLLISIACEOUS FUNGI-VI 

The genus Pyrenopeziza Fuck. 

]. GREMMEN 

Forest Research Station, "De Dorschkamp", TVageningen 

A historical review of the genus Pyrenopeziza Fuck. is given. It appears that 

Pyrenopeziza takes precedence over Mollisia (Fr.) Karst. The diagnosis of the 

genus is emended. A subdivision based on differences of the receptacle is 

attempted, and eight new sections and two subsections are proposed. 

INTRODUCTION.-In the course of the study of the mollisiaceous fungi it soon 

appeared that asci and ascospores are of little use for the identification of the 

species. Measuring of the spores is helpful only in a limited number of longspored 

species. The receptacle, however, shows characteristic features which 

may be of taxonomic value, but the variation of its elements should be interpreted 

very carefully, since young and old receptacles may be totally different 

within the same species. In Pyrenopeziza rubi (Fr.) Rehm for instance, when 

growing on its natural substratum of raspberry stems, young excipular cells 

may be seen to range from 4-8 f1. in diameter, whereas fully developed excipular 

cells measure 8-15 f1. (Gremmen, 1954). In Pyrenopeziza artemisiae (Lasch) 

Rehm the variability of the excipular cells may even be studied in herbarium 

material (Gremmen, 1955). It should also be born in mind that the excipular 

cells of apothecia grown in culture may differ from those of apothecia found in 

nature. Apothecia of Pyrenopeziza plantaginis Fuck. grown in vitro have 

excipular cells larger than those of apothecia developed on their natural 

medium. Similarly, ascospores produced in vitro are longer (Gremmen, 1952). 

However, in spite of certain limitations, the receptacle will be used largely as a 

basis for the following division of the genus Pyrenopeziza. 

For the study of the present genus to be successful, it is first of all necessary to 

investigate such species as are known to form abundant apothecia in nature, 

since that alleviates the comparative study of the various developmental 

stages of the exciple. Species of which few apothecia can be found, should, for 

the time being, be avoided. Such species which Nannfeldt (1932) called "verirrt" 

in general defy any attempt at identification. 

The following abbreviations denote institutions and private collections where material 

mentioned in this paper is located. BM: British Museum, Natural History, London; 

C: Botanical Museum, Copenhagen; G: Conservatoire et Jardin Botaniques, Geneve; 

Grd: herbarium W. D. Graddon, Congleton; Grm: author's herbarium; H: Botanical 

Museum, Helsinki; K: The Herbarium, Royal Botanic Gardens, Kew; L: Rijksherbarium, 

Leiden; PC: Museum National d'Histoire Naturelle, Laboratoire de 

Cryptogamie, Paris; PR: Botanical Department of the National Museum, Prague; 

S: Naturhistoriska Riksmuseum, Stockholm; UPS: Universitetets Institution for Systematisk 

Botanik, Uppsala, and ZT: Institut fur Spezielle Botanik der Eidgenossischen 

Technischen Hochschule, Zurich. 

HISTORICAL REVIEw.-Pyrenopeziza was first used by Fuckel (1869: 293), 

whereas the name Mollisia originated from Elias Fries (1822: 116), who by it

FUNGUS [JAARG. 28 No. 1-4 

indicated a tribus of his genus Peziza. This tribus was raised to generic rank 

by P. A. Karsten (1871: IS), and two sections were distinguished, viz. "Sect. 1. 

Apothecia subiculo nullo vel minus distincto insidentia" and "Sect. n. 

Apothecia subiculo distincto subtomentoso insidentia Tapesia Fr. p.pr." 

Phillips (1887) regarded Mollisia sensu Fr. as a good genus, dividing it in the 

subgenera Niptera Fuck., Pyrenopeziza Fuck., Dilutella Phill., Mollisiella Phill., 

Hysteropeziza Rabh., Pseudopeziza Fuck., and Peristomialis (without author's 

name). In part this scheme is based on the apothecial structure, e.g. Niptera 

being characterized by "texture rather firm; cups plane" and Pyrenopeziza by 

"texture soft; cups globose (mostly black) ," but on the whole the genus is very 

heterogeneous. 

Rehm (r896) recognized in his Mollisieae the group Eumollisieae with 

"Apothecien von Anfang an sitzend" and the Pyrenopezizeae characterized by 

"Apothecien zuerst in die oberen Zellschichten eingesenkt, durch dieselben 

hervorbrechend und dann sitzend." The Eumollisieae are split up into (a) genera 

without a subiculum, comprising Mollisia Fr., Niptera Fr., Belonidium Mont. & 

Dur., and Belonopsis Sacc., and (b) genera with a subiculum such as Tapesia 

Pers. and Trichobelonium Sacc. The Pyrenopezizeae comprise the Pseudopezizeae 

with the genera Pseudopeziza Fuck., and Fabraea Sacc., and the Eupyrenopezizeae 

with Pyrenopeziza Fuck., Pirottaea Sacc. & Speg., Beloniella Sacc., 

and Velutaria Fuck. 

Boudier (r907) mentioned in his family Mollisiacees the following genera: 

Pyrenopeziza Fuck., Ephelina Sacc., Pirottaea Sacc., Coronellaria Karst., 

Mollisia Fr., Tapesia Pers., Niptera Fr., Mollisiella Boud., and Spilopodia Boud. 

He characterized Mollisia as follows: "L'aspect general aplati, plus Hale, des 

receptacles qui sont toujours sessiles, ... " and Pyrenopeziza: "Les receptacles 

sont generalement noirs exterieurement avec la marge plus blanche." According 

to Boudier the paraphyses in M ollisia are filled with an oily protoplasm, lacking 

in Pyrenopeziza. 

In r9I2 Rehm revised the Pyrenopezizeae and the genus Pyrenopeziza was 

again discussed in detail. 

Nannfeldt (1932) recognized two subfamilies in his Mollisioideae, viz. the 

Mollisieae and the Pyrenopezizeae. The first category dealt with the species 

having the "Apothecien ganz oberfHichlich oder nur mit zapfen-formiger Basis 

im Substrat eingesenkt," those of the second group having the "Apothecien im 

Substrat eingesenkt, hervorbrechend." In the former are found the genera 

Haglundia Nannf., Belonium Sacc., Coronellaria Karst., Tapesia Fuck., Trichobelonium 

Rehm, Motlisia (Fr.) Karst., and Belonopsis Rehm; in the latter 

Pirottaea Fuck. and Hysteropezizella v. Hohn. 

The present author already demonstrated that it is hardly possible to delimit 

the genus Pyrenopeziza Fuck. from Mollisia (Fr.) Karst. merely on the basis of 

its erumpent behaviour during development. Nannfeldt occasionally arrived 

at the same conclusion (r932: 123,156), but probably maintained the separation 

for practical purposes. Gremmen (1954) united both genera, assuming Mollisia 

Fr. to be the correct name. It appears, however, that Pyrenopeziza Fuck. was 

published two years prior to Mollisia (Fr.) Karst., so the former takes precedence. 

Some corrections and additions to the diagnosis previously published 

appear necessary.

Nov. 1958J J. GREMMEN: On mollisiaceous fungi-VI 

PYRENOPEZIZA Fuck. em. Gremmen 

Pyrenopeziza Fuck., Symb. myc. 293. 1869. 

Peziza ser. III Phialea Trib. XI Mollisia Fr., Syst. myc. 2: 116. 1822. - Mollisia (Fr.) Karst., 

Myc. fenn. 1: IS. 1871. - Mollisia Fr. sensu Gremmen in Fungus 24:2. 1954. 

Apothecia scattered or gregarious, small, ceraceous, sessile, shortly stipitate or broadly 

based, erumpent or superficial from the beginning, cupuliform to disciform. Receptacle 

even, verrucose, fimbriate or hairy. Ectal excipulum consisting of textura globulosa, 

i.e. of roundish, oblong or polyhedral cells which may be variously coloured or colourless; 

marginally with or without elongate or club-shaped, coloured or colourless cells. 

Hymenium variously coloured. Hypothecium colourless, undefined. Asci minute, clavate 

or cylindrical. Ascospores 4 or 8 per ascus, I-celled, or 2- or 4-septate, bacilliform, 

cylindrical, ovoid, comma-shaped or acicular. Paraphyses filiform or lanceolate, 

colourless. Type species: Peziza chailletii Pers. (vide Nannfeldt, 1932: 137). 

Already Nannfeldt distinguished certain groups in the genus Pyrenopeziza, 

one of which is " ... u.a. charakterisiert durch verhaltnismassig grosse, mehr 

oder weniger niedergedriickt ellipsoidische Apothecien, ..." A number of species 

were considered to belong to this group. Another group" ... bildet eine Anzahl 

von Arten, die samtlich auf vorjahrigen Stengeln wachsen und deren Apothecien 

mit breiter Basis ihren Holzzylindern aufsitzen." 

As appears from the following it is possible to distinguish eight sections which 

are mainly based on differences of the colour and structure of the receptacle. 

KEY TO THE SECTIONS 

I a. Paraphyses filiform 

2a. Receptacle even, glabrous 

3a. Receptacle dark brown or red-brown 

4a. Receptacle sessile on a hypostroma 

4b. Receptacle without a hypostroma 

3b. Receptacle differently coloured 

sa. Receptacle olivaceous or greenish 

brown 

5b. Receptacle melleous or light brown 

2b. Receptacle verrucose, fimbriate or hairy 

6a. Receptacle verrucose 

7a. Ascospores bacilliform 

7b. Ascospores ovoid 

6b. Receptacle with appendages 

8a. Receptacle fimb riate 

8b. Receptacle hairy 

Ib. Paraphyses lanceolate . . . . . . . 

Sectio 1. PYRENOPEZIZA 

· Sectio 2. Hypostromatina 

· Sectio 3. Astromatina 

· Sectio 4. Elaiodiscina 

· Sectio 5. Melipeziza 

· Sectio I. Pyrenopeziza, subsectio a. 

· Sectio I. Pyrenopeziza, subsectio b. 

· Sectio 6. Fimbriaria 

· Sectio 7. Trichantina 

· Sectio 8. Hemipirottaea 

Apothecia erumpentia, sessilia vel basi stipite fusca. Receptaculum verrucosum. In caulibus 

anni praeteriti emortuis. Sectionis species typica: P. chailletii (Pers.) Fuck. 

Apothecia erumpent, with or without dark brown base submerged in host. Receptacle 

verrucose, owing to presence of groups of protruding cells. Inhabiting last year's sterns. 

Type species: P. chailletii (Pers.) Fuck. 

The section is divided in two subsections. 

39

40 FUNGUS [JAARG. 28 No. r-4 

Sabsectio a. Bacillares Gremmen, subsect. novo 

Ascosporae bacilliformes, unicellulares vel pluricellulares. 

Ascospores bacillifonn, 1- or more-celled. 

1. PYRENOPEZIZA CHAILLETII (Pers.) Fuck., Symb. myc. 294. 1869. 

Peziza chailletii Pers., Myc. eur. 1: 288. 1822 (basionym). 

Mat.erial: Pyrenopeziza chailletii Fuck. (G). 

Host: Chaerophyllum aureum. 

2. PYRENOPEZIZA POTENTILLAE (Rostr.) Nannf. in Svensk bot. Tidskr. 22: 136. 

1928. 

Trochilapotentillae Rostr. in Medd. om Gronl. 3: 54°.1888 (basionym). 

Material: Trochila potentillae Rostr., type (C). 

Host: Potentilla nivea. 

3. PYRENOPEZIZA THALICTRI (Peck) Sacc., Syll. Disc. 360. 1889. 

Peziza thalictri Peck in Ann. Rep. N.Y. State Mus. 29: 55. 1878 (basionym). - Pyrenopeziza 

osiliensis Vgr. in Bot. Not. 40. 1900. - Beloniella osiliensis (Vgr.) Rehm in Ber. bayr. bot. Ges. 13: 

18I. 1912. 

Material: no. 517, Pyrenopeziza thalictri (Grd). 

Host: Thalictrum spec. 

4. PYRENOPEZIZA PYRENOCARPOIDES Rehm in Ber. bayr. bot. Ges. 13: 175· 

1912. 

Mollisia pyrenocarpoides (Rehm) Gremmen in Fungus 25: 9. 1955. 

Material: Pyrenopeziza pyrenocarpoides Rehm (S). 

Host: Cirsium spec. 

5. Pyrenopeziza buniadis (Nannf.) ex Gremmen, comb. novo 

Mollisia buniadis Nannf. spec. novo inedit. (basionym). 

Material: Flora suec. 1913, Mollisia bumiadis Nannf., Uppsala, Slottsbacken, 

11.6. 1925, leg. ]. A. Nannfeldt, type (UPS). 

Host: Bunias orientalis. 

Apothecia 400-500 ft diam., basi 80 ft lata. Receptaculum verrucosum, melleum, e textura 

globulosa. Cellulae excipuli 7-10ft diam. Margo cellulis minutis formata. Asci 38-50 X 5·5 ft· 

Ascosporae 9.5-10.5 X 2 ft' unicellulares, incoloratae. Paraphyses filiformes, incoloratae, 

guttulatae. 

Apothecia 400-500 ft across, with 80 ft broad base. Receptacle consisting of textura 

globulosa, honey-brown, verrucose owing to presence ofprotruding cells. Excipular cells 

7-10 ft in diameter, marginally with short cell-excrescences. Asci 38-50 X 5·5 ft. Ascospores 

9.5-10.5 X 2 ft, colourless, 1-celled. Paraphyses colourless, filiform, filled with 

numerous, minute guttulae. 

6. PYRENOPEZIZA LYCHNIDIS (Sacc.) Rehm in Krypt. Fl. 3: 1265. 1896. 

Pyrenopeziza sphaerioides (Pers.) Fuck. f. lychnidis Sacc., Syll. Disc. 365. 1889. 

Material: Rabenh.-Pazschke, Fgi. eur. & extraeur. 4173, Pyrenopeziza 

lychnidis Sacc. (L). 

Host: Lychnis spec. 

7· PYRENOPEZIZA LAVATERAE E. Muller in Sydowia 9: 239· 1955· 

Material: Fgi. Pakistan 4387, Pyrenopeziza lavaterae E. Mull., type (ZT). 

Host: Lavatera spec.

Nov. I958J J. GREMMEN: On mollisiaceous fungi-VI 

Subsectio b. Ovoideae Gremmen, subsect. novo 

Ascosporae ovoideae vel formam commatis aemulantes, unicellulares. 

Ascospores ovoid or comma-shaped, I-celled. 

8. PYRENOPEZIZA NIGRELLA Fuck, Symb. myc. Nachtr. 3: 30. I87!. 

Material: Pyrenopeziza nigrella Fuck (G). 

Host: Succisa pratensis. 

9. PYRENOPEZIZA LYTHRI Fautr. in Rev. myc. 14: 3. 1892. 

Material: Rabenh.-Pazschke, Fgi. eur. & extraeur. 4470, PYl'enopeziza 

lycopodis f. lythri Rehm (L). 

Host: Lythrum salicaria. 

10. PYRENOPEZIZA UNI (Rostr.) Petr. & Syd. in Ann. myc. 22: 356. 1924. 

Laestadia lini Rostr. in Bot. Tidsskr. 28: 217.1907 (basionym). 

Material: Pyrenopeziza lini (Rostr.) Petr. & Syd. (ZT). 

Host: Linum viscosum. 

I!. PYRENOPEZIZA BUBAKII Klika in Ann. myc. 24: 136. 1926. 

Material: Krypt. exs.

42 FUNGUS [JAARG. 28 No. 1-4 

14. Pyrenopeziza revincta (Karst.) Gremmen, comb. novo 

Peziza revincta Karst., Mon. Pez. 157. 1869 (basionym). - Mollisia revincta (Karst.) Rehm 

in Krypt. Fl. 3: 1264. 1896. 

Material: Peziza revincta Karst., type (H). 

Host: Rubus, Vlmaria, Epilobium, and various Compositae. 

IS. Pyrenopeziza benesuada (Tu!.) Gremmen, comb. novo 

Peziza benesuada Tul. in Bot. Zeit. 11: 55. 1853 (basionym). - Mollisia benesuada (Tul.) 

Phill., Man. Brit. Disc. 174. 1887. 

. Material: Flora suec. 9452, Mollisia cf. benesuada (UPS). 

Host: Alnus spec. 

16. Pyrenopeziza adenostylidis (Rehm) Gremmen, cumb. novo 

Mollisia adenostylidis Rehm in Krypt. Fl. 3: 526, 1896 (basionym). 

Material: Rehm, Ascom. 49, Mollisia adenostylidis Rehm (S). 

Host: Adenostyles spec. 

17. Pyrenopeziza arundinacea (DC.) Gremmen, comb. novo 

Xyloma arundinacea DC., Fl. France 6: 162. 1815 (basionym). - Mollisia arundinacea (DC.) 

Phill., Man. Brit. Disc. 177. 1887. 

Material: Rabenh. 423, Peziza cinerea Batsch (L). 

Host: Phragmites communis. 

Sectio 4. Elaiodiscina Gremmen, sect. novo 

Receptaculum olivaceum vel virido-fuscum. In caulibus anni praeteriti emortuis. Sectionis 

species typica: P. carduorum Rehm. 

Receptacle olivaceous or brownish green. Inhabiting last year's stems. Type species: 

P. carduorum Rehm. 

18. PYRENOPEZIZA CARDUORUM Rehm in Winter, Diag. u. Notiz. zu Rehm's 

Ascom. 562. 1872. 

Niptera carduorum (Rehm) Winter in Krypt. Fl. 3: 555. 1896. - Ephelina carduorum (Rehm) 

Rehm in Ber. bayr. bot. Ges. 13: 183. 1912. - Mollisia carduorum (Rehm) Gremmen in Fungus 

25: 5· 1955· 

Material: Rehm, Ascom. 68, Pyrenopeziza carduorum Rehm, type (S). 

Host: Cirsium spec. 

19. Pyrenopeziza pastinacae (Nann£.) Gremmen, comb. novo 

Mollisia pastinacae Nannf. in Nova Acta Reg. Soc. Sci. Ups. ser. 4. 8 (2): 127. 1932 (basionym). 

Material: Flora suec. 1906, Mollisia pastinacae Nannf., isotype (UPS). 

Host: Pastinaca, Anthriscus, Bidens, Cirsium, Heracleum, Angelica. 

20. PYRENOPEZIZA URTICICOLA (Phill.) Boud., Disc. d'Eur. 135. 1907. 

Mollisia urticicola Phill., Man. Brit. Disc. 177. 1887 (basionym). 

Material: no. 1109, Mollisia urticicola Phill., teste ]. A. Nannfeldt (Grd). 

Host: Vrtica dioica. 

21. Pyrenopeziza depressuloides (Gremmen) Gremmen, comb. novo 

Mollisia depressuloides Gremmen in Fungus 25: 9. 1955 (basionym).

44 FUNGUS [JAARG. 28 No. 1-4 

29. PYRENOPEZIZA RUBI (Fr.) Rehm in Krypt. Fl. 3: 611. 1896. 

Excipula rubi.Fr., Syst. myc. 2: 190. "1-822 (basionym). - Mollisia rubi (Fr.) Karst., Rev· 

Mon. 136. 1885. 

Material: Rabenh., Herb. myc. 2, 3°5, Peziza phacidioides Fr. (L). 

Host: Rubus spec. 

30. PYRENOPEZIZA MILLEGRANA Boud., Disc. d'Eur. 133. 1907. 

Mollisia millegrana (Boud.) Nannf. in Nova Acta Reg. Soc. Sci. Ups. ser. 4. 8 (2): 127. 1932. 

Material: Flora suec. 4315, Mollisia millegrana (Boud.) Nannf. (UPS). 

Host: Ulmaria. 

31. PYRENOPEZIZA ARCTII (PhiU.) Nannf. in Nova Acta Reg. Soc. Sci. Ups. ser. 

4. 8 (2): 142. 1932. 

Peziza arctii Phill. in Proc. Bristol Nat. Soc. N.S. 4: 58. 1883 (basionym). - Mollisia arctii 

(Philt) Phill., Man. Brit. Disc. 183. 1887. - Belonium arctii (Phill.) Sacc., Syll. Disc. 495. 1889. 

- Belonidium arctii (Phill.) Massee, Brit. Fung. Fl. 4: 225. 1895. 

Material: Peziza arctii Phill., type (BM). 

Host: Arctium spec. 

32. PYRENOPEZIZA EBULI (Fr.) Sacc., Syll. Disc. 360. 1889. 

Peziza atrata var. ebuli Fr., Syst. myc. 2: 148. 1822 (basionym). - Mollisia ebuli (Fr.) Karst., 

Myc. fenn. 1: 202. 1871. 

Material: Rehm, Ascom. 656-b, Mollisia ebuli (Fr.) Karst. (S). 

Host: Sambucus ebulus. 

33. PYRENOPEZIZA PLANTAGINIS Fuck., Symb. myc. 294. 1870. 

Mollisia plantaginis (Fuck.) Phill., Man. Brit. Disc. 183. 1887. - Peziza atrata var.foliicola 

Desm. in Ann. Sci. Nat. ser. 2, Bot. 368-369.1843. 

Material: Desm. Crypt. Fr. Ser. I, 1313, Peziza atrata var. joliicola Desm. 

(K). 

Host: Plantago lanceolata. 

34. Pyrenopeziza pulveracea (Fuck.) Gremmen, comb. novo 

Peziza pulveracea Fuck., Symb. myc. 297. 1870 (basionym). - Mollisia pulveracea (Fuck.) 

Rehm in Krypt. Ft 3: 532. 1896. 

Material: Fgi. rhen. 2191, Peziza pulveracea Fuck., type (G). 


35. Pyrenopeziza polygoni (Lasch) Gremmen, comb. novo 

Peziza polygoni Lasch in Klotzsch, Herb. vivo myc., in Rabenh., Herb. myc. 1842 (basionym). 

- Mollisiapolygoni (Lasch) Rehm in Krypt. Fl. 3: 527.1896. 

Material: Rehm, Ascom. 70, Niptera polygcni Rehm (L). 

Host: Polygonum amphibium. 

36. PYRENOPEZIZA PETIOLARIS (A. & S. ex Fr.) Nannf. in Nova Acta Reg. Soc. 

Sci. Ups. ser. 4. 8 (2): 158. 1932. 

Hysterium petiolare A. & S., Consp. Fung. Nisk. 59. 1805; ex Fr., Syst. myc 2: 593. 1823 

(basionym). - Trochila petiolaris (A. & S. ex Fr.) Rehm in Krypt. Fl. 3: 132. 1896. - Mollisia 

petiolaris (A. & S. ex Fr.) Gremmen in Fungus 24: 9. 1954. 

Material: no. 9700, Mollisia petiolaris (A. & S. ex Fr.) Gremmen (L). 

Host: Acer spec.

FUNGUS [JAARG. 28 No. 1-4 

Receptacle marginally as well as laterally provided with brown, septate hairs. 

Inhabiting last year's stems. Type species: P. escharodes (B. & Br.) Rehm. 

This group forms a transition to the genus T apesia Fuck. 

44. PYRENOPEZIZA ESCHARODES (B. & Br.) Rehm in Krypt. Fl. 3: 612. 1896. 

Peziza escharodes B. & Br. in Ann. Mag. nat. Hist. ser. 4. 7: 433. 1871 (basionym). 

Lachnella escharodes (B. & Br.) Phill., Man. Brit. Disc. 262. 1887. - Mollisia escharodes (B. & Br.) 

Gremmen in Fungus 24: 5. 1954. 

Material: Peziza escharodes B. & Br., type (BM). 

Host: Rubus spec. 

Sectio 8. Hemipirottaea Gremmen, sect. novo 

Apothecia paraphysibus lanceolatis instructa. In caulibus anni praeteriti emortuis. Sectionis 

species typica: P. lanceolata (Gremmen) Gremmen. 

Apothecia with characteristic lanceolate paraphyses. Inhabiting last year's stems. 

Type species: P. lanceolata (Gremmen) Gremmen. 

This section is intermediate between the section Astromatina and the genus 

Pirottaea Sacc. 

45. Pyrenopeziza lanceolata (Gremmen) Gremmen, comb. novo 

Mollisia lanceolata Gremmen in Fungus 26: 35. 1956 (basionym). 

Material: no. 690, Mollisia lanceolata Gremmen, type (Grm). 


ACKNOWLEDGEMENT.-Dr R. A. Maas .Geesteranus (Leiden) critically 

revised the manuscript for which I am very thankful. 

REFERENCES 

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FRIES, E. 1822-1823. Systema mycologicum 2. Lundae. 

FUCKEL, L. (1869) 1870. Symbolae mycologicae. Wiesbaden. 

GREMMEN, j. 1952. Het genus Pyrenopeziza. Levenswijze en cultuur van Pyrenopeziza plantaginis 

Fuck. in Fungus 22: 22-24. 

--- 1954. Taxonomical notes on Mollisiaceous fungi-I. A study on some Dutch species 

growing on Rubus stems. In Fungus 24: 1-8. 

--- 1955. Taxonomical notes on Mollisiaceous fungi-H. Some caulicolous Mollisia species 

inhabiting various hosts, mainly Compositae. In Fungus 25: 1-12. 

KARSTEN, P. A. 1871. Mycologia fennica, Pars I. Discomycetes. In Bidr. Kann. Fin!. Nat. Folk 

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NANNFELDT,j. A. 1932. Studien iiber die Morphologie und Systematik der nicht-lichenisierten 

inoperculaten Discomyceten. In Nova Acta Reg. Soc. Sci. Ups. ser. 4. 8 (2). 

PHILLlP3, W. 1887. A manual of the British Discomycetes. London. 

REHM, H. 1896. Die Pilze Deutschlands, Oesterreichs und der Schweiz. In Rabenhorst's 

Kryptogamenflora, ed. 2, 1 (3). Leipzig. 

--- 1912. Zur Kenntnis der Discomyceten Deutschlands, Deutsch-Oesterreichs und der 

Schweiz. In Ber. bayr. bot. Ges. 13: 102-206.

The genus Pyrenopeziza Fuck. - ASCOfrance

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