Floristic and chorological news from north Albania - Botanica Serbica

33 (2): 177-183 (2009) 

Floristic and chorological news from north Albania 

Marash Rakaj 

Department of Biology, Faculty of Natural Sciences, University of Shkodra “Luigj Gurakuqi”, Sheshi “2 Prilli” 

Shkoder, Albania 

Received 12 September 2009 Revision accepted 07 December 2009 

Original Scientifi c Paper 

ABSTRACT: Th e North Albania region includes Albanian part of Mt Bjeshkët e Nemuna (Prokletije) massif, 

Mt Pashtrik on the east, Mt Tarabosh (southernmost part Mt Rumija) on the west, as well as the 

right side of the river Drin/Drim valley with surrounding mountain areas. It represents an area 

with the noticeably richer fl ora than any other region in Albania. Six new vascular plant species for 

Albania were reported: Adenostyles alpina (L.) Bluff & Fingerh, Calamagrostis varia (Schrad.) Host, 

Grafi a golaka (Hacq.) Rech., Melampyrum nemorosum L. Crocus weldenii Hoppe & Fürnrohr and 

Viburnum maculatum Pant., of which last two species are Balkan endemics. 

New fl oristic records for the Albanian fl ora are also presented for fi ve taxa: Amphoricarpus autariatus 

Blečić. & May. subsp. bertisceus Blečić & E. Mayer, Iris reichenbachii Heuff ., Rumex balcanicus Rech. 

fi l., Senecio squalidus L. subsp. rupestris (Walds. & Kit.) Greuter and Potentilla heptaphylla L. subsp. 

australis (Krašan ex Nyman) Gams var. malyana (Borbás ex Malý) G. Beck. 

Key words: New fl oristic and chorological additions, North Albania, Bjeshkët e Nemuna (Prokletije), 

Mt Pashtrik. 

INTRODUCTION 

Flora of Albania shows a considerable fl oristic richness 

with a great number of interesting local and regional 

endemics. Th e 4-volumes of Flora e Shqiperise (Paparisto 

et al. 1988) comprises 3250 vascular plant species or 3758 

taxa, including 175 cultivated species, of which 450 are 

Balkan endemic taxa, 46 are local endemic taxa, while 190 

are sub-endemic taxa (V. Stevanovic pers. comm.). Th ey 

constitute 29.5% of the species of the European fl ora, and 

47% of the species of the Balkan Peninsula fl ora. 

Th e North Albania region is located in the southeastern 

part of the Dinaric mountain range including 

mountain blocks of Albanian part of Mt Bjeshkët e 

Nemuna (Prokletije), Mt Pashtrik, on the east, Mt Tarabosh 

(southernmost part Mt Rumija), on the west, as well as the 

right side of the river Drin/Drim valley with surrounding 

mountains where the highest peaks are between 1200 and 

1500 m (Fig. 1). Th e southern border of the North Albania 

region is more than 150 km long valley of the river Drin/ 

Drim. It represents an area with the noticeably richer fl ora 

✳ correspondence: marashrakaj@yahoo.com 

UDK 581.95(496.5) 

than any other region in Albania. Th is area is characterized, 

predominately, by calcareous and serpentine rocks. 

Geological and climatic characteristics. Bjeshkët e 

Nemuna (Prokletije) or so-called Albanian Alps is a 

transboundary high mountain chain which includes c. 

40 separated mountains over 2.000 m high, including 17 

peaks over 2.500 m. Th e highest peaks are Jezerca and 

Gjeravica at 2,694 m and 2,656 m, respectively (Gruda 

1981). Bjeshkët e Nemuna (Prokletije) is mainly composed 

of limestone, although the siliceous rocks also abound. 

Th e Albanian Alps range was formed by the folding 

actions of the African Plate which lies under the European 

one. Pleistocene glaciations are strongly expressed and 

caused actual mountain orography with the numerous 

traces of former glaciers and formation of fl uvial-glacial 

erosion and deposits (moraines, glacial lakes, former 

glacial cirques and valleys). 

Valleys are deep and extended almost to the mountain 

summits, while peaks are very edgy and high. Sources 

of the rivers Valbona, Shala, Cemi/Cijevna, Bistrica of 

© 2009 Institute of Botany and Botanical Garden Jevremovac, Belgrade

178 vol. 33 (2) 

Fig. 1. Th e North Albania region 

Peja/Pečka Bistrica) and some other, are situated in high 

mountain parts of Bjeshket e Nemuna (Prokletije). Th ese 

rivers formed deep canyons and gorges of which some are 

c. 1000 m deep. A plenty of glacial lakes are also present 

in these mountains. For instance, only in the area of the 

mountain peak Jezerce (Buni of Jezerca), six small glacial 

lakes occurred. 

Th e geological substrate of the North Albania is 

complex: limestone and dolomites, diabase-chert 

formation, argiloschists, ultrabasic or serpentine rocks, 

sandstone and conglomerate are to be found (Vranaj 

1990). 

Th e North Albania mountain region belongs to 

Dinaric Mountain transitional eco-region (Hoda 1992). 

It is characterized mainly by Alpine relief, alpine karst 

hydrography, as well as by very rich fl ora with numerous 

endemics and Alpine plant species. A transitional climate 

between perhumid sub-Mediterranean and humid 

mountain climate of the Alpine type is dominant (Gruda 

1981). 

Phytogeographical characteristics. Flora of the Albanian 

Alps is one of the richest in the Balkan. Approximately 

1.611 wild vascular plant species and intraspecifi c taxa 

are described and recorded, which constitutes over 50% of 

Albanian fl ora. Th ese plants belong to 690 genera and 150 

families. Particularly signifi cant are the numerous Tertiary 

and glacial relics, while 250 species are Balkan endemics, 

comprising 136 regional and 17 local endemics distributed 

only in Mt Bjeshkët e Nemuna (Prokletije), such as: Crepis 

bertiscea Jáv., Ligusticum albanicum Jáv., Lunaria telekiana 

Jáv., Valeriana bertiscea Pančić, Pedicularis ernesti-mayeri 

Stevanović, Heliosperma oliverae, Wulfenia baldaccii Degen, 

etc. (Ruci & Vangjeli 2001; Stevanović 2002). 

Th e natural potential vegetation is represented by 

the following communities Fagetum montanum, Abieti- 

Fagetum, Piceetum excelsae montanum, Pinetum mugi, 

Pinetum peucis, Pinetum heldreichii, Oxytropidon dinaricae. 

According to Ruci & Vangjeli (2001) 161 plant communities 

are described in this region of Albania. 

Th e dominant habitat types are forests of the Turkey 

oak, Macedonian oak, Sessile oak, beech, Bosnian pine, 

Macedonian pine, as well as alpine and sub-alpine calcareous 

grasslands. Above the forest belt, there are various shrubby 

communities, mainly dominated by Siberian juniper 

(Juniperus sibirica) and blueberries (Vaccinium spp.). 

Riparian or fl ooded vegetation, as Salicetum eleagni, 

Salicetum fragilis and Alnetum incanae is distributed along 

the river valleys (Ruci & Vangjeli 2001). 

Th e North Albanian region, situated at the crossroads of 

several plant migration routes, is one of the most important 

refugium in the Balkans and, probably, in whole S. Europe. 

It is the most important region both for preservation of 

plants during Ice Age and for plant speciation. Mountains 

of the North Albania could be regarded as a corridor for 

the migrations of plants from the Alps towards the south- 

and eastward regions. 

According to Ruci & Vangjeli (2001) the following 

fl oristic spectrum was recorded in the investigated area: 

European mountains (Central and Alpine) fl oristic 

element 22%, Eurasian 24.5% (Wulfenia, Forsythia, 

Gymnospermium, Ramonda, Aesculus, Dioscorea, Aster 

etc.), Boreal 9.3%, Mediterranean 12.5%, Balkan endemics 

28% and 3.7% of other ones. 

Th e presence taxonomically isolated endemo-relict and 

relict species (Pinus heldreichii Christ, Pinus peuce Griseb., 

Narthecium scardicum Košanin, Taxus baccata L., Forsythia 

europaea Deg. & Bald., Silene asterias Griseb., Edraianthus 

graminifolius (L.) DC., Acer heldreichii Orph., Juglans regia 

L., Asarum europaeum L., Campanula lingulata Waldst. 

et Kit., Valeriana pancicii Hal. et Bald., Rumex balcanicus 

Rech., Ramonda serbica Pančić, Ostrya carpinifolia 

Scop., Jasione orbiculata Griseb. ex Velen.), as well as the 

existence of fl oristic affi nities with ancient Mediterranean 

and sub-Mediterranean fl ora indicate that speciation of 

the Albanian fl ora have began since the Tertiary period. 

Th e occurrence of common sub-endemic taxa in 

the Balkans and Apennine peninsula point to ancient 

amphiadriatic fl oristic link. Th e typical sub-endemic 

species having disjunct ranges (or Apennine-Scardo- 

Pindhic, Illyrian-Scardo-Pindhic-Apennine, Illyrian- 

Apennine, Subscardo-Pindhic-South Apennine, etc), are: 

Pinus heldreichii Crist., Pinus nigra Arnold, Quercus trojana 

Webb, Drypis spinosa L. subsp. linnaeana Murb., Geum 

molle Vis. et Pančić, Potentilla apennina Ten., Anemone 

apennina L., Hypericum barbatum Jacq., Pinguicula 

hirtifl ora Ten., Asyneuma pichleri (Vis.) D. Lakušić &

F. Conti, Campanula lingulata Waldst. et Kit., Jasione 

orbiculata Griseb., Cardamine glauca Spreng., Doronicum 

columnae Ten., etc. 

Pleistocene glaciations strongly eff ected on the current 

composition of the North Albanian mountain fl ora, as 

well as on the whole Balkan fl ora. Th e numerous species 

remained here aft er glaciers melted, such as: Potentilla 

crantzii (Crantz)Beck, Arabis alpina L., Arctostaphylos 

alpinus (L.) Spreng., Aster alpinus L., Saxifraga aizoides L., 

Saxifraga oppositifolia L., Salix retusa L., Salix reticulata L., 

Dryas octopetala L., Androsace villosa L., Geum montanum 

L., Polygonum viviparum L., Trollius europaeus L., Parnassia 

palustris L., Veronica alpina L., Pedicularis verticillata L. 

Nigritella nigra (L.) Reichenb., Juncus trifi dus L., Poa alpina 

L., Luzula spicata DC., Selaginella selaginoides (L.) Schrank. 

& C. F. P. Mart etc. Stevanović et al. (2009) recorded c. 50 

Arctic-Alpine plants in Mt Prokletije. Th e majority of the 

above mentioned plants are glacial relicts. 

Th e North Albanian fl ora is characterised by a high 

number of isolated endemic species. Approximately 110 

Balkan endemics have northern, eastern or southern 

limits of distribution in N Albania that is of special 

phytogeographical interest (Ruci & Vangjeli 2001). Th e 

northernmost limit of distribution is characteristic for 

Alchemilla albanica Rothm., Minuartia baldaccii subsp. 

skutariensis Hayek, Ranunculus degenii Kümmerle & 

Jav., Ranunculus hayekii Dörfl er, Viola kosaninii (Degen) 

Hayek etc), Viola dukadjinica W. Becker et Bornm., while 

the eastern- or southernmost limits have Cerastium 

dinaricum G. Beck et Szyszyl., Edraianthus serpyllifolius 

(Vis.) A. DC., Edraianthus tenuifolius (Waldst. & Kit) A. 

DC., Gentiana dinarica G. Beck, Petasites doerfl eri Hayek, 

Minuartia velenovskyi (Rohlena) Hayek, Teucrium arduini 

L., Verbascum nicolai Rohlena etc. 

Th e number of plant endemic species in whole Mt 

Prokletije massif is particularly high. Approximately 17 

local endemic species are known, as: Alchemilla bertiscea 

Martinčič, Crepis bertiscea Jáv., Dianthus behriorum 

Bornm, Hieracium wettsteinianum Hayek & Zahn subsp. 

wettsteinianum, H. guentheri-beckii subsp. phaedroleucum 

Hayek & Zahn., H. chloropannosum Zahn, H. geminum 

Hayek & Zahn, H. fritschianum Hayek & Zahn, Ligusticum 

albanicum Jav., Lunaria telekiana Jav., Pedicularis ernestimayeri 

Stevanović, Niketić & D. Lakušić, Sesleria wettsteinii 

Dörfl er & Hayek, Heliosperma macranthum Pančić, 

Valeriana bertiscea Pančić, Wulfenia baldaccii Degen, 

Viola vilaensis Hayek, etc. Th ree centres of endemism can 

be distinguished according to number and distribution of 

the regional and local endemic plant taxa from the North 

Albania region: 

1. 

Mt Bjeshkët e Nemuna (Prokletije) – a big isolated 

mountain massif mainly made of calcareous (limestone 

and dolomite) substratum, represented by numerous 

2. 

3. 

M. Rakaj: Floristic and chorological news from north Albania 

regional and local paleo- and neo-endemic species, 

being an important centre of plant speciation and 

refugium; 

Tropoja Valley, Bytyçi and White Drin Valley – a great 

serpentine area, represented also by numerous regional 

paleo- and neo-endemic taxa and few local endemics. 

It is important refugium (south-north corridor) of 

serpentine elements, as well as the speciation centre. 

Has region, including Mt Pashtrik – mainly made of 

limestone, with few stenoendemic species. 

However, the occurrence of the endemic species, such 

as Asperula scutellaris Vis., Crepis baldaccii Halácsy subsp. 

albanica Jáv., Edraianthus serpyllifolius (Vis.) A. DC., 

Phyteuma pseudorbiculare Pant., Pimpinella serbica (Vis.) 

Bentham & Hooker fi l. ex Drude, Potentilla montenegrina 

Pant., Valeriana bertiscea Pančić, etc. confi rm fl oristic 

connection between N Albania and surrounding parts of 

the Balkans (Dinaric Alps and Scardo-Pindhian mountain 

system). 

Th erefore, this region is of a particular phytogeographical 

interest as a barrier for a numerous Mediterranean 

plant species, from one side, and an important 

distribution corridor for orophytes from N Dinaric Alps 

towards south (Albanids, Pindhos) and east (Mt Shar 

Planina), as well as vice-versa, from the other side. 

History of investigation. Th e fi rst fl oristic data for North 

Albanian fl ora have been obtained by Grisebach (1843, 

1844) in the middle of the XIX century. Th e serious work on 

fl ora and vegetation begun in the fi rst part of the XX century, 

when noumerous botanists, such as Degen & Dörfler 

(1897), Baldacci (1904), Janchen (1916), Hayek (1917, 

1924), Jávorka (1921), Jávorka et al. (1926), Markgraf 

(1931), Bornmüller (1933), Rechinger (1935), Lemperg 

(1937), Košanin (1939), Rohlena (1942), etc., started the 

botanical explorations of this region. Th ey recorded and 

described numerous new plant species, including local 

endemics, as Wulfenia baldaccii Degen, Petasites doerfl eri 

Hayek, Centaurea kosaninii Hayek, Crepis bertiscea Jáv., 

Ligusticum albanicum Jáv., Polygala doerfl eri Hayek, 

Sanguisorba albanica Andrasovszky and Jáv., etc. 

In the last three decades a number of Albanian 

botanists have also investigated this area and published 

their own studies, such as Demiri (1983), Ruci (1986), 

Paparisto et al. (1988), Hoda (1992), Qosja et al. (1992, 

1996), Vangjeli et al. (2000), Ruci & Vangjeli (2001), 

Rakaj & Kashta (2003, 2007), Rakaj (2006), Kashta 

(2007), Barina & Pifko (2008a, 2008b) etc. In this region 

several new taxa for Albanian fl ora were recorded during 

the last decade: Dianthus behriorum Bornm, Hyacinthella 

dalmatica (Baker) Chouard., Lathraea squamaria L., 

Parietaria lusitanica L., Prunus tenella Batsch, Ranunculus 

montanus Willd. 

179

180 vol. 33 (2) 

It is worth to mention that the fl ora and vegetation 

of the northern parts of Mt Prokletije and Mt Pashtrik, 

outside Albania, were also investigated in more details by 

some ex-Yugoslav botanists: Lakušić (1971), Wraber 

(1986, 1989), Martinčič (1990), Lakušić & Stevanović 

(1995), Stevanović (1999), Stevanović & Lakušić 

(2000), Stevanović et al. (2002, 2003), Lakušić et 

al. (2004), Niketić (2005), Niketić & Stevanović 

(2006), etc. Th ey found seven new local endemic species: 

Alchemilla bertiscea Martinčič, Cynoglossum krasniqii 

T. Wraber, Draba bertiscea D. Lakušić & Stevanović, 

Draba kuemmerlei Stevanović & D. Lakušić, Hieracium 

bertisceum Niketić, Heliosperma oliverae Niketić & 

Stevanović, Pedicularis ernesti-mayeri Stevanović, Niketić 

& D. Lakušić. Most of them were found in close proximity 

of the Albanian territory, so it is probable to assume that 

some of these taxa will be recorded in Albania too. 

MATERIALS AND METHODS 

Th e results of the present paper are based mainly on 

personal investigations and collecting plants material 

during the period of 2000-2009, and partly on the plant 

material stored in herbarium of Faculty of Natural Science, 

University of Shkodra, Albania. In addition, the literature 

data were used for the presentation of the distribution of 

the investigated taxa. Th e new plant species for Albania are 

mapped on UTM grid 10 x 10 sq. km. 

RESULTS AND DISCUSSION 

New taxa for the Albanian fl ora. Albanian fl ora, particularly, 

in mountain is not suffi ciently known. In this 

paper six new vascular plant taxa for Albanian fl ora were 

reported, including one new genus (Grafi a) (Fig. 2-4).: 

1. Adenostyles alpina (L.) Bluff & Fingerh: limestone, in 

high-mountain rocky grounds and screes on Mt Shtegu 

i Dhenve (UTM CM99), 1850m, coll. T. Wraber, M. 

Rakaj, 15.07.2002; Mt Maja Jezerce (UTM DM09), 

2340m, coll. B. Surina, M. Rakaj 16.08.2007; Mt 

Parun (UTM CM88), 1740m, coll. B. Surina, M. Rakaj 

18.08.2007 (Fig. 2). 

Th is species was known from Alps, N Apennines 

and NW Dinarides (Slovenia and Croatia). Th e new 

localities in Albanian Alps represent a very isolated 

disjunction on the eastern and southern part of the 

species range. 

2. Calamagrostis varia (Schrad.) Host: limestone, in 

high-mountain rocky grounds on Mt Shtegu i Dhenve 

(UTM CM99), 1700-1850m, coll. B. Surina, M. Rakaj 

15.08.2007; Mt Parun, (UTM CM88), 1640m, coll. B. 

Surina, M. Rakaj 18.08.2007 (Fig. 3). 

Eurasian species widespread in mountain areas of C and 

Fig. 2. – Distribution of Adenostyles alpina (L.) Bluff & Fingerh 

in Albania. 

Fig. 3. – Distribution of Calamagrostis varia (Schrad.) Host in 

Albania. 

S Europe. It has been recorded in all Balkan countries, 

except in Albania. Collected samples belong to the type 

subspecies (subsp. varia). 

3. Crocus weldenii Hoppe & Fürnrohr: sunny hill slopes, 

grassy and stony places; Tarabosh, Goraj, Vrith (UTM 

CM65), 100 - 200m, coll. M. Rakaj, 22.02.2009; 

14.03.2009 (Fig. 4). 

Balkan endemic with general distribution in Slovenia, 

Croatia, Herzegovina, Montenegro, Serbia and 

Macedonia.s North-western border of the species range 

reaches the vicinity of Trieste in NE Italy (Pulević 

1977, Randjelović et al. 1990). It is interesting to note 

that Randjelović et al. (1990) also cited Albania for 

general distribution but without any locality. Species 

belongs to C. bifl orus Mill. aggregate. 

4. Grafi a golaka (Hacq.) Rech.: limestone, on rocky 

ground next to Mt Shtegu i Dhenve (UTM CM99), 

1700m, coll. B. Surina, M. Rakaj, 15.07.2007 (Fig. 4). 

Apennine-Balkan (Dinaric) mountain species. Th e new 

discovered population in Albania is situated on the 

easternmost part of the species area, and also provides 

the south-easternmost border of the Balkan part of 

the area. Th is species has been also found in adjacent 

parts of Mt Prokletije in Montenegro (M. Niketić, pers. 

comm.).

Fig 4. – Distribution of Crocus weldenii Hoppe & Fürnrohr 

(black circle) and Grafi a golaka (Hacq.) Rech., Melampyrum 

nemorosum L. and Viburnum maculatum Pant. (triangle) in 

Albania. 

5. Melampyrum nemorosum L.: on calcareous rocks of 

west slopes under Shpella upper Okol (Th eth) (UTM 

CM99), 1450m, coll.: T. Wraber, M. Rakaj 16.07.2002; 

M. Rakaj, B. Surina 16.08. 2007 (Fig. 4). 

Widespread species in Europe, except on the West. It 

is also known from adjacent parts of Mt Prokletije in 

Montenegro (Rohlena 1942). 

6. Viburnum maculatum Pant.: limestone, on rocky 

grounds on Mt Maja Shtegut (Th eth) (UTM CM99), 

1500m; coll. S. Barina, M. Rakaj, 16.08.2007 (Fig. 4). 

Regional endemic outer chain of SE Dinaric Alps under 

strong Mediterranean climatic infl uence along Adriatic 

coast from Mt Orjen to Mt Rumija. 

Th e new discovered locality in Albania provides the 

southeasternmost border of the species area. 

New fl oristic records for the Albanian fl ora from North 

Albania 

1. Amphoricarpus autariatus Blečić. & May. ssp. bertisceus: 

in stony places in the beech and pine forest on Th ore, 

1650m; bellow Qafa Pejes, 1600m; coll: B. Surina, M. 

Rakaj, 15, 08. 07. It is distributed southward to NW 

Greece. Regional endemic. 

2. Iris reichenbachii Heuff . (=Iris bosniaca Beck): 

limestone, in high-mountain pastures and rocky 

grounds on Mt Pashtrik (UTM DM67), 1750m, coll. 

T. Wraber, M. Rakaj, L. Kashta 07.06. 2001. Literature 

data: Mt Maja Gjallices, 1840m (Košanin 1939: 75). 

Balkan – S Carpatian fl oristic element. 

3. Rumex balcanicus Rech. fi l.: limestone, high mountains 

and rocky grounds of Mt Maja Malisores (UTM DM09), 

2300m; coll: M. Rakaj, 15.08. 08. Tertiary relict and 

Balkan endemic distributed in SE Dinaric Alps and N 

Scardo-Pindhic mountains. 

4. Potentilla heptaphylla L. subsp. australis (Krašan ex 

Nyman) Gams var. malyana (Borbás ex Malý) G. Beck: 


serpentine, in stony places in the bush on Cahan, 

Mici-Has (Has) (530-800m) and (Kolsh 470m) (UTM 

DM57), coll. T. Wraber, M. Rakaj, L. Kashta 23.07. 

2002. Literature data: Skanderbeg (Stane të Frenkthit, 

serpentine) (Markgraf 1931, sub. P. australis). 

P. heptaphylla subsp. australis is a south European 

orophyte, with an ecotype (var. malyana) characteristic 

for serpentinite rocky ground in the W and C Balkan. 

5. Senecio squalidus L. subsp. rupestris (Walds. & Kit.) 

Greuter: on open grassy and calcareous rocks on Fusha 

e Runices 1750m (UTM CN90), coll. T. Wraber, M. 

Rakaj, 18.07.2000; Kroj i Kuq (Mt Gjallice) 2100m 

(UTM DM55); coll.: T. Wraber, M. Rakaj, 18.07.2000; 

05.06.2001. Literature data: Mt Maja Pashtrik (Košanin 

1939:75-105). Th is C and SE European orophyte is 

widespread through the Balkan mountains, particularly 

in the Dinaric Alps. 

CONCLUSIONS 

Flora of the North Albania is one of the richest in the 

region, even though it is not suffi ciently known. Six new 

vascular plant taxa for Albanian fl ora were reported from 

North Albania: Adenostyles alpina (L.) Bluff & Fingerh, 

Calamagrostis varia (Schrad.) Host., Grafi a golaka (Hacq.) 

Rech., Melampyrum nemorosum L., Crocus weldenii Hoppe 

& Fürnrohr and Viburnum maculatum Pant., of which last 

two are Balkan endemics. Th e genus Grafi a is a new for the 

fl ora of Albania. 

Th e new fl oristic records for fi ve taxa of the Albanian 

fl ora are also presented: Amphoricarpus autariatus Blečić. 

& May. subsp. bertisceus, Iris reichenbachii Heuff ., Rumex 

balcanicus Rech. fi l., Senecio squalidus L. subsp. rupestris 

(Walds. & Kit.) Greuter and Potentilla heptaphylla L. subsp. 

australis (Krašan ex Nyman) Gams var. malyana (Borbás 

ex Malý) G. Beck. 

Th e occurrence of numerous Tertiary and glacial 

relicts, as well as the high number of regional and local 

endemics establish North Albanian region, particularly 

Mt Bjeshkët e Nemuna (Prokletije), as a one of the most 

important centre of diversity and evolution of vascular 

fl ora in SE Europe. 

Acknowledgements – I would like to express the highest 

gratitude to Prof. Tone Wraber, University of Ljubljana 

(Slovenia) and dr Boštjan Surina, National History 

Museum, Rijeka (Croatia), for their sincere support 

during our fi eld trip in North Albanian mountains. I 

am also gratefully acknowledged to dr Marjan Niketić, 

Natural History Museum, Belgrade (Serbia) and Prof. 

Vladimir Stevanović, Department of Plant Ecology and 

Phytogeography, Faculty of Biology, University of Belgrade 

(Serbia) for valuable comments and suggestions. 

181

182 vol. 33 (2) 

REFERENCES 

Baldacci A. 1904. Resultati botanici e fi togeografi ci 

della due misione scientifi che italiane del 1902 e 1903 

al Montenegro e nell’Albania fi nitima. (Rend.d.sess.d.R. 

Acc.d.Sc.di Bologna. 

Barina Z & Pifko D. 2008a. Additions and amendments 

to the fl ora of Albania. Wildenowia 38: 455-464. 

Barina Z & Pifko D. (2008b): Data on the fl ora of Albania. 

Acta Botanica Hungarica 50: 231-236. 

Bornmüller J. 1933. Zur Flora von Montenegro, Albanien 

und Mazedonien. Mag. Bot. Lapok. 32: 109-142. 

Degen A & Doerfler J. 1897. Beitrang zur Flora 

Albaniens und Mazedoniens. Wien. 

Demiri M. 1983. Flora eskursioniste e Shqiperise. Shtepia 

Botuese e Shqiperise, pp. 986 Tirana 

Grisebach A. 1843. Spicilegium Florae rumelicae et 

bithynicae. Vol. I. pp. 407. Braunsvigae. 

Grisebach A. 1844. Spicilegium Florae rumelicae et 

bithynicae. Vol. II. pp. 539. Braunsvigae. 

Gruda GJ. 1981. Alpet e Shqiperise (PhD Th esis), pp. 178. 

University of Tirana (in Albanian) 

Hayek A. 1917. Beitrag zur Kenntnis der Flora des 

albanisch-montenegrinischen Grenzgebietes. Denkschrift 

der Akademie der Wissenschaft e, Mathematisch- 

Naturwissenschaft liche Klass, Wien 94: 127-210. 

Hayek A. 1924. Zweiter Beitrag zur Kenntnis der Flora von 

Albanien. Denkschrift der Akademie der Wissenschaft e, 

Mathematisch-Naturwissenschaft liche Klass, Wien 

99:101-224. 

Hoda P. 1992. A geobotanical survey of the black pine 

(Pinus nigra Arn.) forests in Albania. Annali di Botanica 

51: 77-92. 

Janchen E. 1916. Notizen zur Herbstfl ora der 

Nordwestlichen Albanien. Oesterr. Bot. Zeitschr. 66: 

386-397. 

Jávorka S. 1921. Uj adatok Albania fl orajahoz 

(Novitates fl orae Albanicae). Bot. kozlem. 19: 17-29. (in 

Hungarian) 

Jávorka S, Degen A, Gayer G, Soó R, Trautmann R 

& Zahn KH. 1926. Additamenta ad fl oram Albaniae 

Reisebericht, VII. Anthophyta. In: Teleki P & Csiki E. 

(eds.) A Magyar Tudomanyos Akademia Kiadasa, pp. 

74-89, 219-346. Budapest. 

Košanin N. 1939. Über die Vegetation von Nordalbanien. 

Spomenik LXXXIX (Prvi Razed) 20: 75-105. 

Lakušić R. 1971. Noch eine Art der Gattung Wulfenia 

Jacq. auf dem Prokletije Gebirges. Glasn. Republ. Zav. 

Zažt. Prir.- Prir. Muz. Titograd 4: 15-33. 

Lakušić R. 1973a. Rumex balcanicus Rech. fi l. – novi 

tercijerni relikt u fl ori Crne Gore. Glas. Rep. Zavoda 

Zašt. Prir. Titograd 5: 29–32. (in Serbo-Croat) 

Lakušić D, Stevanović V, Bulić Z, Jovanović S, 

Tomović G & Vukojević S. 2004. Florological and 

chorological contributions to the vascular fl ora of 

Montenegro. Glas. Rep. Zav. Zašt. Prirode Muz. 

Podgorica, 27-28: 33-42. 

Lemperg F. 1937. Verzeichnis der 1936 in Nordalbanien 

und Westmazedonien gesammelten Pfl anzen. 

Hatzendorf. (Manuscript aufb ewart im Inst. für Bot. 

Uni. Graz, Österreich).. 

Markgraf F. 1931. Pfl anzen aus Albanien 1928. Denkschrift 

der Akademie der Wissenschaft e, Mathematisch- 

Naturwissenschaft liche Klass, Wien 102: 317-360. 

Martinčič A. 1990. Prispevek k poznavanju rodu 

Alchemilla L. v Jugoslaviji. Biološki Vestnik 38: 23-38. (in 

Slovenian) 

Niketić M. 2005. Hieracium bertisceum (Compositae), 

a new species from the State Union of Serbia and 

Montenegro. Willdenowia 35: 265-269. 

Niketić M & Stevanović V. 2006. A new species 

of Heliosperma (Caryophyllaceae) from Serbia and 

Montenegro. Botanical Journal of the Linnean Society 

154: 55–63. 

Paparisto K, Demiri M, Mitrushi I & Qosja XH. 1988. 

Flora e Shqiperise, Vol. 1. Akademia e Shkencave e RPS 

te Shqiperise, Qendra e Kerkimeve Biologjike, pp. 457. 

Tirana. (in Albanian) 

Pulević V. 1977. Prilog taksonomiji i horologiji nekih 

vrsta Crocus L. iz Jugoslavije (C. weldenii, C. alexandrii 

i C. adamii sens lat.). Glasn. Republ. Zavoda za zaštitu 

Prir. Prirodnjačke zbirke Titograda 10: 81–99. (in Serbo- 

Croat) 

Rakaj M. 2006. “Njoft imi i parë mbi gjetjen e tre llojeve 

bimore në Shqipëri: Parietaria lusitanica L. subsp. 

lusitanica, Oenothera parodiana Munz subsp. parodiana 

dhe Lathrea squamaria L.” Buletin Shkencor i USH “Luigj 

Gurakuqi”, Seria e Shkencave të Natyrës, Shkodër 56: 88– 

95. (in Albanian). 

Rakaj M & Kashta L. 2007. “Njoft imi i parë mbi gjetjen 

e katër specieve të bimëve me lule (Angiospermae) 

në Shqipëri: Hyacinthela dalmatica (Baker) Chouard, 

Prunus tenella Batsch. Ranunculus montanus Wild. dhe 

Dianthus behriorum Bornm.” Buletin Shkencor i USH 

“Luigj Gurakuqi”, Seria e Shkencave të Natyrës, Shkodër 

57: 111-119. (in Albanian). 

Randjelović N, Hill DA & Randjelović V. 1990. Th e 

genus Crocus L. in Serbia. Th e Serbian Academy of 

Sciences and Arts, pp.52. Belgrade. 

Rechinger KH. 1935. Ergebnisse einer botanischen Reise 

in den Bertiscus (Nordalbanische Alpen). Feddes Report. 

38: 137- 152, 319-389. 

Rohlena J. 1942. Conspectus Florae Montenegrinae. 

Preslia 20/21: 1-506.

Qosja XH, Paparisto K, Demiri M, Vangjeli J & Balza 

E. 1992. Flora e Shqiperise, Vol. 2. Akademia e Shkencave 

e Republikes se Shqiperise, Qendra e Kerkimeve 

Biologjike, pp. 446. Tirana. (in Albanian) 

Qosja XH, Paparisto K, Vangjeli J & Ruci B. 1996. Flora 

e Shqiperise, Vol 3. – Akademia e Shkencave e Republikes 

se Shqiperise, Instituti i Kerkimeve Biologjike, pp 331. 

Tirana. (in Albanian) 

Ruci B. 1986: Konsiderata mbi bimësisnë dhe fl orën e 

rrethit të Shkodrës (PhD Th esis, munscript), pp 666. 

Tirana. (in Albanian). 

Ruci B & Vangjeli J. 2001. Te dhena mbi fl oren dhe 

vegjetacionin e Alpeve te Shqiperise. Studime Biologjike 

(Instituti i Kërkimeve Biologjike. Ak.Shk. Tirana) 7: 90- 

96. (in Albanian). 

Stevanović V (ed.). 1999. Crvena knijga fl ore Srbije 1. 

Iščlezli i krajnje ugroženi taksoni. (Th e Red Data Book 

of Flora of Serbia) pp. 566. Beograd. 

Stevanović V & Lakušić D. 2000. Draba kuemmerlei 

(sect. Aizoides), nom. nov. (Brassicaceae) – taxonomic, 

chorological and ecological characteristics. Botanika 

Chronika 13: 87-93. 

Stevanović V, Niketić M & Lakušić D. 2002. Pedicularis 

ernesti-mayeri (P. subset. Comosae, Scrophulariaceae), a 

REZIME 


Novi fl oristički i horološki podaci za Severnu 

Albaniju 

Marash Rakaj 

Severna Albanija obuhvata i deo Prokletijskog masiva, Paštrik na istoku i planinu Taraboš kao krajnji južni 

deo Runije na zapadu, te prostor desno od reke Drim sa dolinom i okolnim planinama. To je prostor koji 

ima primetno bogatiju fl oru od drugih albanskih regiona. Po prvi put se navodi šest novih vrsta za Albaniju: 

Adenostyles alpina (L.) Bluff & Fingerh, Calamagrostis varia (Schrad.) Host, Grafi a golaka (Hacq.) Rech., 

Melampyrum nemorosum L. Crocus weldenii Hoppe & Fürnrohr i Viburnum maculatum Pant. Od kojih su 

poslednje dve Balkanski endemiti. 

Novi podaci za Albansku fl oru dati su za pet taksona: Amphoricarpus autariatus Blečić. & May. subsp. bertisceus 

Blečić & E. Mayer, Iris reichenbachii Heuff ., Rumex balcanicus Rech. Fil., Senecio squalidus L. subsp. rupestris 

(Walds. & Kit.) Greuter i Potentilla heptaphylla L. subsp. australis (Krašan ex Nyman) Gams var. malyana (Borbás 

ex Malý) G. Beck. 

Ključne reči: Novi fl oristički i horološki podaci, Severna Albanija, Prokletije, Paštrik. 

new species from Mt Prokletije (Yugoslavia). Razprave 4 

Razreda SAZU (Ljubljana) 42: 209-226. 

Stevanović V, Tan K & Iatrou G. 2003. Distribution 

of the endemic Balkan fl ora on serpenitine. Obligate 

serpentines endemics. Plant Syst. Evol. 242: 149-170. 

Stevanović V, Vukojičić S, Šinžar-Sekulić J, 

Lazarević M, Tomović G & Tan K. 2009. Distribution 

and diversity of Arctic-Alpine species in the Balkans. 

Plant Syst. Evol. 283: 219-235. 

Vangjeli J, Ruci B, Mullaj A, Paparisto K & Qosja XH. 

2000. Flora e Shqiperise, Vol. 4. Akademia e Shkencave 

e Republikes se Shqiperise, Instituti i Kerkimeve 

Biologjike, pp 502. Tirane. (in Albanian). 

Vranaj A. 1990. Gjeologjia e Shqiperise, pp.420. Tirana. 

(in Albanian) 

Wraber T. 1989. Botanični izlet v Gusinjske Prokletije. 

Proteus 51: 291-297. (in Slovenian) 

Wraber T. 1986. Ein neue Cynoglosum (C. krasniqii) aus 

Jugoslawien. Candollea 41: 145-150. 

183

Floristic and chorological news from north Albania - Botanica Serbica

Create successful ePaper yourself

Delete template?

Save as template?