1 Check-list of Cladosporium names Frank M. DUGAN, Konstanze ...

Check-list of Cladosporium names 

Frank M. DUGAN, Konstanze SCHUBERT & Uwe BRAUN 

Abstract: DUGAN, F.M., SCHUBERT, K. & BRAUN; U. (2004): Check-list of Cladosporium 

names. Schlechtendalia 11: 1–119. 

Names of species and subspecific taxa referred to the hyphomycetous genus Cladosporium 

are listed. Citations for original descriptions, types, synonyms, teleomorphs (if known), 

references of important redescriptions in literature, illustrations as well as notes are given. 

This list contains data of 772 taxa, i.e., valid, invalid and illegitime species, varieties and 

formae as well as herbarium names. 

Zusammenfassung: DUGAN, F.M., SCHUBERT, K. & BRAUN; U. (2004): Checkliste der 

Cladosporium-Namen. Schlechtendalia 11: 1–119. 

Namen von Arten und subspezifischen Taxa der Hyphomycetengattung Cladosporium 

werden aufgelistet. Bibliographische Angaben zur Erstbeschreibung, Typusangaben, 

Synonyme, die Teleomorphe (falls bekannt), wichtige Literaturhinweise und Abbildungen 

sowie Anmerkungen werden angegeben. Die vorliegende Liste enthält Namen von 772 Taxa, 

d. h. gültige, ungültige und illegitime Arten, Varietäten, Formen und auch Herbarnamen. 

Introduction: 

Cladosporium Link (LINK 1816) is one of the largest genera of hyphomycetes, comprising 

more than 772 names, but also one of the most heterogeneous ones, which is not very 

surprising since all early circumscriptions and delimitations from similar genera were rather 

vague and imprecise (FRIES 1832, 1849; SACCARDO 1886; LINDAU 1907, etc.). All kinds of 

superficially similar cladosporioid fungi, i.e., amero- to phragmosporous dematiaceous 

hyphomycetes with conidia formed in acropetal chains, were assigned to Cladosporium s. lat., 

ranging from saprobes to plant pathogens as well as human-pathogenic taxa. DE VRIES (1952) 

and ELLIS (1971, 1976) maintained broad concepts of Cladosporium. ARX (1983), MORGAN- 

JONES & JACOBSEN (1988), MCKEMY & MORGAN-JONES (1990), MORGAN-JONES & MCKEMY 

(1990) and DAVID (1997) discussed the heterogeneity of Cladosporium and contributed 

towards a more natural circumscription of this genus. DAVID (1997) carried out detailed 

examinations of the conidiogenous loci and conidial hila of Cladosporium (incl. Heterosporium 

Klotzsch ex Cooke) species by means of SEM microscopy and showed that true 

species of the latter genus are confined to anamorphs of mycosphaerella-like ascomycetes 

with a unique scar type. The protuberant conidiogenous loci (scars) and conidial hila are 

characterised by having a central convex dome surrounded by a raised rim (coronate). 

Molecular examinations of cladosporioid fungi, carried out during the course of the past 

decade, proved the heterogeneity of Cladosporium s. lat. and confirmed the approach of 

DAVID (1997) to confine this genus to species with coronate conidiogenous loci, which are, as 

far as known, anamorphs of mycosphaerella-like ascomycetes. BRAUN et al. (2003) provided 

results of comprehensive ITS sequence analyses of cladosporioid fungi, summarised 

outcomes of molecular examinations published by other authors and discussed phylograms of 

cladosporium-like fungi in detail. Human-pathogenic cladophialophora-like hyphomycetes 

[Herpotrichiellaceae], Sorocybe resinae (Fr.) Fr. (Amorphotheca resinae Parbery) [Amorphothecaceae], 

Alternaria malorum (Rühle) U. Braun, Crous & Dugan (Cladosporium malorum 

Rühle) [Pleosporaceae] and cladosporioid Venturia Sacc. anamorphs (Fusicladium Bonord.) 

[Venturiaceae] formed separate monophyletic clades and could be excluded from 

Cladosporium s. str. Within a big clade formed by members of the Mycosphaerellaceae, 

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species with anamorphs belonging to the genus Cladosporium s. str. were shown to represent 

a sister clade to Mycosphaerella Johanson s. str. (with cercosporoid anamorphs), for which 

the teleomorph genus Davidiella Crous & U. Braun was proposed. The results of these 

examinations led to a reassessment and new phylogenetic circumscription of Cladosporium s. 

str. Now we are confronted with the big task to prove the generic affinity of hundreds of 

names assigned to Cladosporium and to contribute to comprehensive, monographic examinations 

of the latter genus. The present annotated list is meant to be a first step towards a 

monograph of Cladosporium. 

List of names 

This list encompasses names from SACCARDO (1882–1972), KIRK (1985), ‘Botanischer 

Jahresbericht’ (Vol. 1, for 1873, to 10, for 1882) and ‘Just´s Botanischer Jahresbericht’ (Vol. 

11, for 1883, to 63, for 1935), OUDEMANS (1919–1924), ‘Petrak´s Lists’ (1920–1939), ‘A 

Supplement to Petrak´s List’ (1920–1939), ‘Index of Fungi’ (1940–2003), ‘Index Fungorum’ 

(http://www.indexfungorum.org/Names/Names.asp), plus names from other sources. All taxa 

which have been put into Cladosporium are alphabetically arranged, independent of their 

nomenclatural status as valid, invalid, illegitime or herbarium names (if a name was found in 

herbarium or other records, but no place of publication is recorded, the specific epithet is also 

placed in alphabethical order within the list, along with the name of the author if known and 

the source of the name (herbarium label, catalog, etc.). The name of the genus 

(Cladosporium) is left out and is only cited for the synonyms. 

The list provides citations for original descriptions, types, synonyms, important redescriptions 

and illustrations by subsequent authors as well as commentaries. The names of authors and 

their abbreviations follow KIRK (2003), and those of periodicals are from LAWRENCE et al. 

(1968) and BRIDSON & SMITH (1991). Original data for the types (primarily in Latin, French, 

German or Italian) are translated into English. Other highly pertinent data or comments 

excerpted from SACCARDO (l.c.), ‘Index of Fungi’ and other sources are usually presented 

unaltered in original languages. Type specimens and ex-type cultures are noted as such when 

possible. Abbreviations of herbaria follow HOLMGREN et al. (1990) and abbreviations of 

culture collections are from JONG et al. (1996). Names of accepted, already checked, true 

members of Cladosporium s. str. and the current, accepted names for synonyms and excluded 

taxa are printed in bold. Host range and distribution data are only given for species of the 

latter category (if more specimens and records than the type collection are known). 

Inconsistencies and mistakes in the quotation of names and references have often been 

encounted in the original publications inspected. Therefore, we have tried to find and examine 

the original papers whenever possible (checked original publications are marked by ‘!’). 

PFISTER (1985) and STAFLEU & COWAN (1976–1988) list herbaria containing specified sets of 

exsiccatae. KOHLMEYER (1962) was consulted with reference to specimens from Rabenhorst. 

STEVENSON (1971) was an important source of data dealing with the publication of numerous 

exsiccatae. 

Abbreviations 

General: 

cf. = 

comb. inval. = combination invalidum (invalid combination). 

Ill. = illustrations. 

Lit. = literature, references (important publications dealing with the species concerned, mostly 

with redescriptions). 

nom. anamorph. = nomen anamorphum (). 

nom. illeg. = nomen illegitimum (illegitime name). 

nom. inval. = nomen invalidum (invalid name). 

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nom. nov. = nomen novum (new name). 

nom. nud. = nomen nudum (name without any description or diagnosis). 

nom. prov. = nomen provisorium (name proposed provisionally) 

nom. superfl. = nomen superfluum (superfluous name) 

s. lat. = sensu lato (in a wide sense). 

s. str. = sensu stricto (in a narrower sense). 

T = type (typus). 

! = original publication seen and checked. 

Exsiccata 

Barthol., F. columb. = Bartholomew, Fungi columbiani. 

Barthol., N. Am. F. = Bartholomew, North American Fungi. 

Braun, F. sel. exs. = U. Braun, Fungi selecti exsiccati. 

Briosi & Cav., F. paras. = Briosi & Cavara, I funghi parassiti delle piante coltivate od utili, 

essicati, delineati e descritti. 

Cooke, F. brit. exs. = Cooke, Fungi britannici exsiccati. 

Desm., Pl. crypt. N. France = Desmazières, Plantes cryptogames Nord du France. ? 

Ellis, N. Am. F. = Ellis, North American Fungi. 

Ellis & Everh., F. columb. = Ellis & Everhardt, Fungi columbiani. 

Erb. Critt. Ital. = Erbario Crittogamico Italiano. 

Erikss., F. paras. scand. = Eriksson, Fungi parasitici scandinavici exsiccati. 

Fuckel, F. rhen. = Fuckel, Fungi rhenani. 

Herb. Mycol. Rom. = Herbarium Mycologicum Romanicum. 

Jaap, F. sel. exs. = Jaap, Fungi selecti exsiccati. 

Kab. & Bub., F. imp. exs. = Kabát & Bubák, Fungi imperfecti exsiccati. 

Klotzsch, Herb. viv. myc. = Herbarium vivum mycologicum. 

Lib., Pl. crypt. ard. = Libert, Plantae cryptogamicae guas in Arduenna collegit. 

Petr., Mycoth. gen. = Petrak, Mycotheca generalis. 

Rabenh., F. eur. = Rabenhorst, Fungi europaei exsiccati. 

Rabenh., Herb. mycol. = Rabenhorst, Herbarium mycologicum. 

Rav., F. amer. exs. = Ravenel, Fungi americani exsiccati. 

Roum., F. gall. exs. = Roumeguère, Fungi gallici exsiccati. 

Roum., F. sel. gall. exs. = Roumeguère, Fungi selecti gallici exsiccati. 

Sacc., Mycoth. ital. = Saccardo, Mycotheca italica. 

Sacc., Mycoth. ven. = Saccardo, Mycotheca veneta. 

Syd., F. exot. exs. = H. Sydow, Fungi exotici exsiccati. 

Syd., Mycoth. germ. = H. & P. Sydow, Mycotheca germanica. 

Syd., Mycoth. march. = P. Sydow, Mycotheca marchica. 

Thüm., F. austr. = de Thümen, Fungi austriaci. 

Thüm., Herb. myc. oec. = de Thümen, Herbarium mycologicum oeconomicum. 

Thüm., Mycoth. univ. = de Thümen, Mycotheca universalis. 

abietinum (Pers.) Link, Ges. Naturf. Freunde Berlin Mag. Neuesten Entdeck. Gesammten 

Naturk. 7: 37 (1816)!. 

T: on bark of Pinus abies. 

≡ Dematium abietinum Pers., Neues Mag. Bot. 1: 121 (1794)!. 

≡ Sporotrichum abietinum (Pers.) Link, Ges. Naturf. Freunde Berlin Mag. Neuesten Entdeck. 

Gesammten Naturk. 3: 13 (1809)!. 

Lit.: DE VRIES (1952: 89). 

Notes: According to HUGHES (1958) this species has to be reduced to synonymy with Trentepohlia 

abietina (Flot.) Hansg. 

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abietinum Zukal, Verh. K. K. Zool.-Bot. Ges. Wien 37: 44–45 (1887)!, nom. illeg., 

homonym, non C. abietinum (Pers.) Link, 1816. 

T: on moist tuber skins of Solanum tuberosum (Solanaceae), Austria, 1885, associated with 

Chaetomium crispatum. 

≡ Spondylocladium abietinum Sacc., Syll. fung. 10: 662 (1892)!, as ‘(Zukal) Sacc.’. 

Lit.: LINDAU (1910: 142, as synonym of ‘Spondylocladium atrovirens Harz’). 

Ill.: ZUKAL (1887: Tab. 1, Fig. 8). 

Notes: Zukal’s type, housed at GZU could not be located there (personal communication with 

Christian Scheuer). 

acaciae Panwar → acaciicola. 

acaciae Reichert, Bot. Jahrb. Syst. 56: 720 (1921)!. 

T: on dry fruits of Acacia farnesiana (Mimosaceae), Egypt, ‘in insula Rhoda ad Kahiram’, 

Feb. 1822/25, Ehrenberg (B). 

Lit.: SACCARDO (1972: 1336). 

Ill.: REICHERT (1921: Tab. 4, Fig. 2). 

acaciicola M.B. Ellis, More Dematiaceous Hyphomycetes: 342 (1976)!. 

T: from rhizosphere of Acacia nilotica subsp. indica (Mimosaceae), India, Rajastan, Jodhpur, 

sand dunes of Masuria (IMI 104172). 

≡ C. acaciae Panwar, Curr. Sci. 39(18): 422 (1970)!, non C. acaciae Reichert, 1921. 

Ill.: ELLIS (1976: 343, Fig. 260 A). 

Notes: Excluded, black yeast-like. 

acerinum Noelli, Nuovo Giorn. Bot. Ital., N.S. 24(3): 195 (1917)!. 

T: on branches of Acer negundo (Aceraceae), Italy, Piemont, Bruere (Rivoli), 29 Mar. 1915. 

Lit.: SACCARDO (1931: 787). 

acutum Ellis & Dearn., Proc. Canad. Inst., N.S., 3, 1: 91 (1897)!. 

T: on fallen leaves of Fraxinus sp. (Oleaceae), Canada, Ontario, London, Victoria park, 

Oct./Nov. 1896 (DAOM). 

= Cladosporium herbarum (Pers.: Fr.) Link, Ges. Naturf. Freunde Berlin Mag. Neuesten 

Entdeck. Gesammten Naturk. 7: 37 (1816)!. 

Lit.: SACCARDO (1899: 1080). 

adianticola R.F. Castañeda, Fungi Cubenses II: 3 (1987)!, as ‘adianticolum’. 

T: on living leaves of Adiantum sp. (Adiantaceae), Cuba, prov. Matanzas, San Miguel de los 

Baños, 23 Jan. 1987, R.F. Castañeda (INIFAT C87/40: holotype). 

Lit.: CROUS et al. (2005). 

Ill.: CASTAÑEDA (1987: Figs 3, 5). 

Notes: Excluded, generic affinity unclear. 

aecidiicola Thüm., Mycoth. univ., Cent. IV, No. 373 (1876)!. 

T: on aecia of rust on living leaves of Euphorbia cyparissias (Euphorbiaceae), Germany, 

Bavaria, Bayreuth, 1874, Thümen, Thüm., Mycoth. univ. 373 (e.g., B 70-6144, B 70-6146; 

BPI 426074; HAL; HBG; M-57483: syntypes). 

= Cladosporium vincae Moesz, Bot. Közlem. 23: 123 (1926)!, nom. illeg., homonym, non C. 

vincae Fairm., 1911. [W 10216], syn. nov. 

= Cladosporium aecidii Pass., in herb. (B 70-6132). 

Lit.: SACCARDO (1886: 368, as ‘aecidiicolum’, 1913: 1371), LINDAU (1907: 806–807, 1910: 

796), FERRARIS (1912: 350), GONZÁLES-FRAGOSO (1927: 211), ELLIS (1976: 330), ELLIS & 

ELLIS (1985: 571, 1988), BRAUN & ROGERSON (1995: 142), HEUCHERT et al. (2005: 14–17). 

Ill.: ELLIS (1976: 330, Fig. 248), HEUCHERT et al. (2005: 15, Fig. 1). 

Host(s)/substrate(s) & distribution: on aecia of rusts including those of Puccinia phragmitis 

and Uromyces limonii on different host plants; Europe (Austria, Germany, Hungary, Italy, 

Montenegro, Romania, Switzerland), North America (USA). 

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Notes: BRAUN & ROGERSON (1995) mention the first record of the species from North 

America on aecia of Puccinia grindeliae on leaves of Chrysothamnus viscidiflorus var. 

lanceolatus as new host. 

aequatoriense Petr., Sydowia 2: 380 (1948)!. 

T: on living leaves of Mikania sp. (Asteraceae), Ecuador, prov. Pichincha, Guarumos near 

Nono, 17 Oct. 1937 (M-57478: syntype). 

≡ Parastenella aequatoriensis (Petr.) K. Schub. & U. Braun, Mycol. Progr. 4(2): 103 (2005)!. 

Ill.: SCHUBERT & BRAUN (2005a: 104, Fig. 3). 

aeruginosum F. Patt., Bull. Torrey Bot. Club 27: 284 (1900)!. 

T: on living leaves of Olea fragrans (Oleaceae) affected by Gloeosporium oleae, USA, 

Department of Agriculture, greenhouse, Jan. 1900, F.W. Patterson (type in herb. of ‘Division 

of Vegetable Physiology and Pathology, U.S. Department of Agriculture’). Topotype: BPI 

426094 (from 6 Feb. 1900). 

Lit.: SACCARDO (1902: 1058). 

agoseridis U. Braun & Rogerson, Sydowia 47(2): 142 (1995)!. 

T: on living leaves of Agoseris glauca (Asteraceae), USA, Utah, Washington Co., northwest 

side of Pine Valley Mountain, northeast of Diamond Valley, vicinity of Mud Spring, 7 Jun. 

1994, C.T. Rogerson (NY: holotype; HAL 1556: isotype). 

Ill.: BRAUN & ROGERSON (1995: 143, Fig. 1). 

Host(s)/substrate(s) & distribution: on Agoseris glauca; North America (USA). 

albicans Hallier, Parasitologische Untersuchungen bezüglich auf die pflanzlichen Organismen 

bei Masern, Hungertyphus, Darmtyphus, Blattern, Kuhpocken, Schafpocken, Cholera Nostra 

etc.: 27 (1868)!. 

T: from man, causing parasitic stomatitis. 

≡ Cladosporium albicans Hallier, Flora, Neue Reihe, 26(19): 293 (1868)!, nom. prov. 

Ill.: HALLIER (1868b: Pl. II, Fig. 2). 

Notes: Considered to be a state (morphe) of Oidium albicans C.P. Robin [≡ Candida albicans 

(C.P. Robin) Berkhout] by HALLIER (1868b: 27); human pathogenic, undoubtedly not 

belonging to Cladosporium s. str. HALLIER (1866: 86) discussed this fungus under 

‘Stempylium polymophum ? (Oidium albicans auct.)’. 

albiziae S.N. Khan & B.M. Misra, Indian Forester 125(7): 746 (1999)!. 

T: on leaves of Albizia lebbek (Mimosaceae), India, Uttar Pradesh, Kalsi (Dehra Dun) (IMI 

282484). 

Ill.: KHAN & MISRA (1999: 745, Fig. 2). 

Notes: The species described herein is allied to C. psoraleae M.B. Ellis (1976) but differs in 

its virulence, lesion formation and conidial characters (KHAN & MISRA 1999). 

album Dowson, J. Roy. Hort. Soc. 49(2): 211 (1924). 

T: on living leaves of Lathyrus odoratus (Fabaceae), Great Britain. 

≡ Hyalodendron album (Dowson) Diddens, Zentralbl. Bakteriol., 2. Abt., 90: 316 (1934)!. 

≡ Ramularia alba (Dowson) Nannf., in Lundell & Nannfeldt, Fungi exs. suec., Fasc. XXXIX– 

XL, No. 17 (1950)!. 

= Ramularia galegae f. lathyri Ferraris, Malphigia 20: 153 (1906)!. 

≡ Ramularia lathyri Ferraris, Flora Ital. Crypt., Pars I, Fungi: 812 (1913)!. 

≡ Ramularia lathyri (Ferraris) Bubák, Ann. Mycol. 14: 350 (1916)!. 

= Ramularia lathyri Hollós, Bot. Közlem. 1910, 2: 112 (1910)!. 

= Ramularia deusta f. odorati W.C. Snyder & W.H. Davis, Mycologia 42: 417 (1950)!. 

= Ramularia deusta [(Fuckel) Karak.] var. alba U. Braun, Nova Hedwigia 56: 429 (1993)!. 

Lit.: BRAUN (1998: 157). 

algarum Cooke & Massee, in Cooke, Grevillea 16(79): 80 (1888)!. 

T: on rotting fronds of Laminaria flexicaulis (= Laminaria digitata) (Laminariaceae), Great 

Britain, Sussex, Hastings, Bexhill [as ‘Baxhill’], Dec. 1887, E.M. Holmes (K: lectotype, 

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selected by Kohlmeyer; isolectotypes: NY 72455, 72456). 

≡ Heterosporium algarum (Cooke & Massee) Cooke & Massee, in Cooke, Grevillea 18(88): 

74 (1890)!. 

= Cladosporium herbarum [(Pers.: Fr.) Link] var. macrocarpum (Preuss) M.H.M. Ho & 

Dugan, in Ho, Castañeda, Dugan & Jong, Mycotaxon 72: 131 (1999)!. 

Lit.: KOHLMEYER & KOHLMEYER (1979: 482), DAVID (1997: 71–73). 

algeriense (Montpell. & Catanei) Vuill., Champ. paras.: 78 (1931)!. 

T: isolated from man. 

≡ Hormodendrum algeriense Montpell. & Catanei, Ann. Dermatol. Syphiligr., Sér. 6, 8: 626– 

635 (1927). 

Lit.: DODGE (1935: 845), DE HOOG et al. (2000: 1014). 

Notes: DE HOOG et al. (2000) describe it as a doubtful species close to or identical with 

Fonsecaea pedrosoi (Brumpt) Negroni [current name Phialophora pedrosoi (Brumpt) 

Redaelli & Cif.]. 

allii (Ellis & G. Martin) P.M. Kirk & J.G. Crompton, Pl. Pathol. 33: 320 (1984)!. 

T: on leaves of Allium vineale (Alliaceae), USA, New Jersey, Newfield, Jun. 1882, Ellis 

collection No. 3608a (NY: neotype, selected by KIRK & CROMPTON, 1984; IMI 270432, 

slide). 

≡ Heterosporium allii Ellis & G. Martin, J. Mycol. 1: 100 (1885)!. 

= Heterosporium ornithogali var. allii-porri Sacc. & Briard, in Briard, Rev. Mycol. 

(Toulouse) 8: 25 (1886)!. [T: PAD]. 

≡ Heterosporium allii var. allii-porri (Sacc. & Briard) Sacc., Syll. fung. 4: 480 (1886)!. 

≡ Heterosporium allii-porri (Sacc. & Briard) Nicolas & Aggéry, Rev. Pathol. Vég. Entomol. 

Agric. France 14: 197 (1927). 

≡ Cladosporium allii-porri (Sacc. & Briard) Boerema, Verslagen Meded. Plantenziektenk. 

Dienst Wageningen 152: 15, 1977 (1978)!. 

≡ Heterosporium allii-porri (Sacc. & Briard) Arx, Genera Fungi Sporul. Pure Cult., ed. 3: 305 

(1981)!, comb. inval. 

= Heterosporium allii var. allii-sativi Bontea & Dumitraș, Rev. Roumaine Biol., Sér. Bot. 

12(6): 389 (1967)!. [T: ?BUCM; K; IMI]. 

Teleomorph: ? Didymellina intermedia Cif., Not. Malatt. Piante 2: 6–7 (1949) (?, not 

confirmed). 

Lit.: KIRK (1986a), DAVID (1997: 29–33), HO et al. (1999: 116–118). 

Ill.: KIRK & CROMPTON (1984: 321, Fig. 3), DAVID (1997: 31, Fig. 4; 32, Fig. 5 A–C). 

Host(s)/substrate(s) & distribution: on leaves of Allium including A. ampeloprasum, A. ascalonicum, 

A. canadense, A. fistulosum, A. oleraceum, A. porrum, A. sativum, A. schoenoprasum 

(including A. sibiricum), A. scorodoprasum, A. vallidum, A. vineale; Europe (Belgium, Czech 

Republic, Denmark, France, Germany, Great Britain, Greece, Latvia, Norway, Romania, 

Russia, Ukraine), Asia [Armenia, China, Georgia, Japan, Kazakhstan, Russia (Amur Oblast)], 

North America (Canada, USA). 

Notes: In OUDEMANS (1919) Kniphofia uvaria (Asphodeliaceae) is mentioned as host of 

Heterosporium allii, which is very doubtful. 

allii-cepae (Ranoj.) M.B. Ellis, More Dematiaceous Hyphomycetes: 337 (1976)!. 

T: on dying stems of Allium cepa (Alliaceae), Serbia, Sumadija, Rebschule von Veliko 

Oraschje, Jun. 1905, N. Ranojević (BPI: holotype; IMI 270434, slide ex herb. BPI). 

≡ Heterosporium allii-cepae Ranoj., Ann. Mycol. 8: 399 (1910)!. 

= ?Heterosporium allii var. cepivorum Nicolas & Aggéry, Rev. Pathol. Vég. Entomol. Agric. 

France 14: 197 (1927). 

Teleomorph: Davidiella allii-cepae (M.M. Jord., Maude & Burchill) Crous & U. Braun, in 

Braun, Crous, Dugan, Groenewald & de Hoog, Mycol. Progr. 2(1): 10 (2003)!. 

Lit.: ELLIS & ELLIS (1985: 305), KIRK (1986b), DAVID (1997: 33–34). 

Ill.: KIRK & CROMPTON (1984: 322, Fig. 4; 323, Fig. 5), SHIN (1995: 92, Fig. 1 a–d), DAVID 

(1997: 32, Fig. 5 D–G; 35, Fig. 6). 

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Host(s)/substrate(s) & distribution: usually Allium cepa, also A. fistulosum; Asia (India, Korea, 

Japan), Europe (Great Britain, Greece, Ireland, Norway, Serbia), North America (USA?). 

alliicola H.D. Shin & U. Braun, Korean J. Mycol. 23(2): 141 (1995)!. 

T: on Allium victorialis var. platyphyllum (Alliaceae), Korea, Suwon, 3 Sept. 1993, H.D. Shin 

(SMK 12597: holotype; HAL 1533: isotype; SMK 12761: paratype). 

= Cladosporium victorialis (Thüm.) U. Braun & H.D. Shin, in Braun & Melnik, Proc. 

Komarov Bot. Inst. (St. Petersburg) 20: 101 (1997)!. 

Ill.: SHIN & BRAUN (1995: 140, Fig. 1; 141, Fig. 2 a–b). 

allii-porri (Sacc. & Briard) Boerema, Verslagen Meded. Plantenziektenk. Dienst Wageningen 

152: 15, 1977 (1978)!. 

T: on dead leaves of Allium porrum (Alliaceae), France, Troyes, Briard (PAD: holotype). 

≡ Heterosporium ornithogali var. allii-porri Sacc. & Briard, in Briard, Rev. Mycol. 

(Toulouse) 8: 25 (1886)!. 

≡ Heterosporium allii var. allii-porri (Sacc. & Briard) Sacc., Syll. fung. 4: 480 (1886)!. 

≡ Heterosporium allii-porri (Sacc. & Briard) Nicolas & Aggéry, Rev. Pathol. Vég. Entomol. 

Agric. France 14: 197 (1927). 

≡ Heterosporium allii-porri (Sacc. & Briard) Arx, Genera Fungi Sporul. Pure Cult., ed. 3: 305 

(1981), comb. inval. 

= Cladosporium allii (Ellis & G. Martin) P.M. Kirk & J.G. Crompton, Pl. Pathol. 33: 320 

(1984)!. 

Lit.: DAVID (1997: 29–30), HO et al. (1999: 116–118). 

Ill.: HO et al. (1999: 117, Fig. 1). 

alliorum Hanzawa, Mycol. Centralbl. 5: 11 (1914)!. 

T: on leaves of Allium cepa (Alliaceae), Japan, Sapporo. 

Lit.: SACCARDO (1931: 793). 

Ill.: HANZAWA (1914: 6, Fig. 2). 

Notes: HANZAWA (1914) described the species as closely related to Cladosporium herbarum 

but distinct by having somewhat swollen conidiophores and thick conidia. 

alneum Pass. ex K. Schub., sp. nov. 

T: on living leaves of Alnus glutinosa (Betulaceae), Italy, Emilia Romagna, Parma, 1879, G. 

Passerini (B 70-6156: holotype). 

≡ Cladosporium alneum Pass., in herb. 

Host(s)/substrate(s) & distribution: on Alnus glutinosa; Europe (Italy). 

alnicola Bubák & Vleugel, in Vleugel, Svensk Bot. Tidskr. 11(3–4): 322 (1917)!, nom. illeg., 

homonym, non C. alnicola Corda, 1837. 

T: on Alnus incana var. borealis (Betulaceae), Sweden, prov. Vesterbotten, Umeå, Sept. 1911, 

J. Vleugel (BPI 426104: holotype). 

= Cladosporium herbarum (Pers.: Fr.) Link s. lat. 

Lit.: SACCARDO (1931: 789). 

alnicola Corda, Icon. fung. 1: 14 (1837)!, as ‘alnicolum’. 

T: on rotten wood of Alnus sp. (Betulaceae), Czech Republic (PRM). 

≡ Didymotrichum alnicola (Corda) Bonord., Handb. Mykol.: 89 (1851)!. 



Lit.: SACCARDO (1886: 354, as ‘alnicolum’), LINDAU (1907: 819), OUDEMANS (1920), 

HUGHES (1958: 750). 

Ill.: CORDA (1837: Tab. 4, Fig. 211). 

alopecuri (Ellis & Everh.) U. Braun, Schlechtendalia 5: 32 (2000)!. 

T: on Alopecurus geniculatus (Poaceae), USA, Montana, Columbia Falls, 20 May 1887, B.T. 

Galloway (NY: holotype). 

≡ Fusicladium alopecuri Ellis & Everh., J. Mycol. 4: 53 (1888)!. 

7

Ill.: BRAUN (2000: 33, Fig. 2). 

Host(s)/substrate(s) & distribution: on Alopecurus geniculatus; North America (USA). 

Notes: In FARR et al. (1989) Malus sylvestris (USA, MT) is given as a further host of the 

species, which is very doubtful. 

alpiniae T. Zhang & Z.Y. Zhang, Plant Diseases and Their Control: 108 (1998)!. 

T: on living leaves of Alpinia galanga (Zingiberaceae), China, Hubei, Wuchang, 22 Sept. 

1980, J.Y. Li & T.Y. Zhang, No. 4074 (MHYAU 03947: holotype). 

Ill.: ZHANG & ZHANG (1998b: 109, Fig. 1), ZHANG et al. (2003: 38, Fig. 12). 

Notes: ZHANG & ZHANG (1998b) cited Alpinia zerumbet as an additional host species and 

compared C. alpiniae with C. cladosporioides, the only other species occurring on a host 

belonging to the Zingiberaceae (on Zingiber officinale), stating that the latter species possesses 

longer, limoniform or ellipsoid conidia. Since type material was not available the status of this 

taxon remains unclear. 

alternicoloratum R.F. Castañeda & W.B. Kendr., Univ. Waterloo Biol. Ser. 35: 20 (1991)!. 

T: on leaves and stems of Cyperus alternifolium (Cyperaceae), Cuba, Pinar del Río, Cuchillas 

de San Simón, 24 Mar. 1990, R.F. Castañeda (INIFAT C90/129: holotype). 

Ill.: CASTAÑEDA & KENDRICK (1991: 21, Fig. 10). 


amaranticola Opiz, Lotos 5: 41 (1855)!, nom. nud. 

T: on Amaranthus retroflexus (Amaranthaceae), Czech Republic, Prag, park Stromovka, 5 Jan. 

1853, Opiz (PRM 657400). 

= Cladosporium herbarum (Pers.: Fr.) Link s. lat. 

Lit.: LINDAU (1907: 832). 

ambrosiae House, in herb. 

On dead stems of Ambrosia trifida (Asteraceae),USA, New York, Albany Co. Selkirk, 10 Apr. 

1925, H.D. House (NYS). 

americanum H.C. Greene, Amer. Midl. Naturalist 41(3): 723 (1949)!. 

T: on living leaves of Prunus americana (Rosaceae), USA, Wisconsin, Dane Co., Madison, 

University of Wisconsin Arboretum, 18 Sept. 1944, H.C. Greene (BPI 426105; WIS: 

syntypes). 

= Fusicladium carpophilum (Thüm.) Oudem., Verh. Kon. Ned. Akad. Wetensch., Afd. 

Natuurk. 1900: 388 (1900)!. 

Lit.: BRAUN (2001: 53), SCHUBERT (2005: 56–57). 

Ill.: SCHUBERT (2005: 57, Fig. 1). 

amoenum R.F. Castañeda, in Untereiner et al., BCCM MUCL Agro-industrial fungi-yeasts 

(1998), nom. nud. 

T: on fallen leaves of Eucalyptus sp. (Myrtaceae), Cuba, Santiago de Cuba, La Gran Piedra, 2 

Nov. 1994, R.F. Castañeda (HO et al. 1999: 117, Figs 2–3: iconotype; ATCC 200947: epitype; 

INIFAT C94/155, CBS 254.95, MUCL 39143, IMI 367525: isoepitypes). 

≡ Anungitopsis amoena R.F. Castañeda & F.M. Dugan, in Ho, Castañeda & Dugan, Mycotaxon 

72: 118 (1999)!. 

amorphae Thüm., Rev. Mycol. (Toulouse) 1: 59 (1879)!. 

T: on dead stems of Amorpha herbacea (Fabaceae), USA, South Carolina, Aiken, H.W. 

Ravenel, Thüm., Mycoth. univ. 1572 (e.g., BPI 426106; BR-MYC 81368,82; HAL; M; NY). 

Lit.: SACCARDO (1886: 362). 

ampelinum Pass., Erb. Critt. Ital., Ser. 2, No. 595 (1872)!. 

T: on leaves of Vitis sp. (Vitaceae), Italy, Erb. Critt. Ital. 595 (E; IMI 112146). 

= Pseudocercospora vitis (Lév.) Speg., Anales Mus. Nac. Buenos Aires 20: 438 (1910)!. 

Lit.: SACCARDO (1886: 458), LINDAU (1910: 116), OUDEMANS (1921), SIVANESAN (1984: 

210), CROUS & BRAUN (2003: 427). 

Notes: see C. vitis (Lév.) Sacc. 

8

amphitrichum Sacc., Syll. fung. 4: 354 (1886)!. 

T: on rotten wood of Pinus sp. (Pinaceae), Czech Republic, near Reichenberg (PRM). 

≡ Amphitrichum olivaceum Corda, Icon. fung. 1: 16 (1837)!, non C. olivaceum (Corda) 

Bonord., 1851. 

Lit.: LINDAU (1907: 811). 


anomalum Berk. & M.A. Curtis, in Berkeley, J. Linn. Soc., Bot. 10: 362 (1869)!. 

T: on the underside of leaves of a Malvaceae, Cuba, C. Wright, Fungi cubensis Wrightiani, 

No. 639 (K). 

≡ Pseudocercospora anomala (Berk. & M.A. Curtis) de Hoog, Persoonia 15(1): 68 (1992)!. 

Lit.: SACCARDO (1886: 363). 

Notes: Authentic material is also deposited at PC (C. Wright collection, 1870, North Pacific 

Expedition 1853–56). 

annonae Nann., Atti Reale Accad. Fisiocrit. Siena, Ser. 10, 4(1–2): 91 (1929)!. 

T: on living leaves of Annona sp. (Annonaceae), Italy, Siena, botanical garden, Aug. 1928 

(SIENA). 

Lit.: SACCARDO (1972: 1336). 

Host(s)/substrate(s) & distribution: on Annona sp.; Europe (Italy). 

antillanum R.F. Castañeda, Fungi Cubensis II: 3 (1987)!. 

T: on fallen leaves of Clusia rosea (Clusiaceae), Cuba, prov. Guantánamo, Imias, 3 May 1986, 

Mayra Camino (INIFAT C86/128: holotype). 

Ill.: CASTAÑEDA (1987: Fig. 6). 


aphidis Thüm., Herb. myc. oec., Fasc. X, No. 484 (1877)!, with description on the label. 

T: on dead carcass of Aphis symphyti (Homoptera, Aphididae) on leaves of Symphytum officinale, 

Austria, Klosterneuburg, Aug. 1875, Thümen, Thüm., Herb. myc. oec. 484 (e.g., M). 

= Cladosporium aphidis Thüm., Oesterr. Bot. Z. 27: 12 (1877)!, homonym. 

aphidis Thüm., Oesterr. Bot. Z. 27: 12 (1877)!, homonym. 

T: on dead carcass of Aphis symphyti (Homoptera, Aphididae), Austria, Klosterneuburg, Aug. 

1876, Thümen, Thüm., Mycoth. univ. 672 (e.g., HAL; M; MICH). 

= Cladosporium aphidis Thüm., Herb. myc. oec., Fasc. X, No. 484 (1877). 

Lit.: SACCARDO (1886: 369), LINDAU (1907: 830), FERRARIS (1912: 351), LIND (1913: 522). 

aphidis [Thüm.] var. muscae Briard & Har., Rev. Mycol. (Toulouse) 12: 132 (1890)!. 

T: on dead carcass of fly (Musca), France, Méry-sur-Seine, P. Hariot. 

Lit.: SACCARDO (1892: 605). 

apicale Berk. & Broome, J. Linn. Soc., Bot. 14: 99, 1873 (1875)!. 

T: on leaves of Cycas circinalis (Cycadaceae), Ceylon (Sri Lanka), Peradeniya, Jan. 1868, 

G.H.K. Thwaites (K 121544: isotype). 

Lit.: SACCARDO (1886: 367), ELLIS (1976: 332). 

Ill.: ELLIS (1976: 334, Fig. 252 A). 

Host(s)/substrate(s) & distribution: on Cycas circinalis; Asia (Sri Lanka). 

Notes: DE & CHATTOPADHYAY (1994) describe and illustrate this species from West Bengal, 

India, on Swietenia mahogoni (Meliaceae) [IMI 209595]. 

apiculatum Berk., in herb. 

On Helianthus sp. (Asteraceae), USA, Carolina (K). 

Lit.: SACCARDO (1895: 621). 

aquilinum, in herb. 

On Pteridium sp. (Dennstaedtiaceae); on Pteridium and Salib., Northampton (PH). 

araguatum (Syd.) Arx, Genera Fungi Sporul. Pure Cult., ed. 2: 224 (1974)!. 

T: on living leaves of Pithecellobium lanceolatum (Mimosaceae), Venezuela, Aragua, La Vic- 

9

toria, between La Victoria and Suata, Jan. 1928, H. Sydow (BPI 443420, 443421, 443422; IMI 

15728, IMI 34905). 

≡ Stenella araguata Syd., Ann. Mycol. 28: 205 (1930)!. 

= Cladosporium castellanii Borelli & Marcano, Castellania 1(5): 154 (1973). [T: IMI]. 

Lit.: MCGINNIS & PADHYE (1978). 

araliae Sawada, Rep. Gov. Res. Inst. Formosa 85: 91 (1943)!, nom. inval. 

T: on Aralia decaisneana (Araliaceae), Taiwan, 26 Nov. 1928, K. Sawada (BPI 426122; 

PPMH: syntypes). 

≡ Stenella araliae Sawada ex K. Schub. & U. Braun, sp. nov. 

arcticum Berl. & Voglino, Syll. fung. 4a: 170 (1886)!. 

T: on Epilobium latifolium (Onagraceae), Danmark, Greenland, Kaiser-Franz-Joseph-Fjord. 

Lit.: SACCARDO (1891: 882). 

Notes: The species is described by BERLESE & VOGLINO (1886) as anamorphic state of 

Pleospora arctica Fuckel. 

argillaceum Minoura, J. Ferment. Technol. 44: 140 (1966)!. 

T: isolated from a decayed myxomycete, Japan, Yaku Island, 21 Oct. 1961, K. Tubaki (Dept. 

Fermentation Technology, Faculty Engineering, Osaka, Japan). ATCC 38103 (= CBS 241.67) 

(Ex-type). 

Lit.: HEUCHERT et al. (2005: 53). 

Ill.: MINOURA (1966: 142, Fig. 6 E). 


aristolochiae H. Zhang & Z.Y. Zhang, Mycosystema 17(4): 304 (1998)!. 

T: on living leaves of Aristolochia kwangsiensis (Aristolochiaceae), China, Hubei, Wuchang, 

22 Sept. 1980, J.Y. Li & T.Y. Zhang (MHYAU 03956: holotype). 

Ill.: ZHANG & ZHANG (1998a: 304, Fig. 1). 

aromaticum Ellis & Everh., Proc. Acad. Nat. Sci. Philadelphia 47(3): 439 (1895)!. 

T: on living leaves of Rhus aromatica (Anacardiaceae), USA, California, Pasadena, Aug. 

1894, A.J. McClatchie (BPI 426124; NY: syntypes). 

≡ Fusicladium aromaticum (Ellis & Everh.) K. Schub. & U. Braun, Fungal Diversity (2005)!, 

in press. 

= Cladosporium nervale Ellis & Dearn., in Barthol., F. columb., Cent. XXI, No. 2010 (1905)!. 

Lit.: SACCARDO (1899: 1079), CASH (1952: 69). 

Ill.: SCHUBERT & BRAUN (2005b, Fig. 2). 

artemisiae H.C. Greene, Amer. Midl. Naturalist 48(3): 757 (1952)!. 

T: on living leaves of Artemisia caudata (= A. campestris spp. caudata) (Asteraceae), USA, 

Wisconsin, Dane Co., Madison, University of Wisconsin Arboretum, sandy slope, 7 Jul. 1951, 

H.C. Greene (BPI 426134, WIS: syntypes). 

≡ Fusicladium artemisiae (H.C. Greene) K. Schub. & U. Braun, in Schubert, Mycotaxon 92: 

57 (2005). 


arthoniae M.S. Christ. & D. Hawksw., in Hawksworth, Bull. Brit. Mus. (Nat. Hist.), Bot. 

6(3): 210 (1979)!. 

T: on apothecia of Arthonia impolita (Arthoniaceae) on Quercus (Fagaceae), Sweden, Skåne, 

Genarp, Häckeberga, 24 Apr. 1946, M. Skytte Christiansen (herb. Christiansen 570: holotype). 


Ill.: HAWKSWORTH (1979: 211, Fig. 10). 

Notes: Excluded, will be treated and reassessed separately. 

arthrinioides Thüm. & Beltr., in Thümen, Nuovo Giorn. Bot. Ital. 8: 252 (1876)!. 

T: on living and wilted leaves of Bougainvillea spectabilis (Nyctaginaceae), Italy, Sicily, 

Palermo, botanical garden, Mar. 1875, V. Beltrani-Pisani (M-57465), and Thüm., Mycoth. 

univ. 873 (BPI 426137; HAL; HBG; M-57466; MICH: syntypes). 

10

Lit.: SACCARDO (1886: 359), FERRARIS (1912). 

Host(s)/substrate(s) & distribution: on Bougainvillea spectabilis; Europe (Italy). 

arthropodii K. Schub. & C.F. Hill, sp. nov. 

T: on Arthropodium cirratum (Anthericaceae, Liliaceae s. lat.), New Zealand, Auckland, Glen 

Funes, University of Auckland, Tamaki Campus, 1 Jul. 2004, C.F. Hill, No. 1054, mixed 

infection with Alternaria sp. (HAL 1828 F: holotype). Paratypes: on Arthropodium cirratum, 

New Zealand, East Tamaki, Auckland University Campus, 4 Sept. 2003, E.H.C. McKenzie 

(PDD 78376) and Little Huia, 3 Dec. 1963, J.M. Dingley (PDD 23039). 

Host(s)/substrate(s) & distribution: on Arthropodium cirratum; New Zealand. 

artocarpi Gonz. Frag. & Cif., Bol. Soc. Esp. Hist. Nat. 25: 366 (1925)! and Publ. Estac. 

Agron. Haina, Ser. B, Bot. 1: 14 (1925). 

T: on faded leaves of Artocarpus incisa (Moraceae), Dominican Republic, near Haina, 19 Apr. 

1925, R. Ciferri (MA 06393: holotype). 

artocarpi Kulhara & Singh. (?). 

On Artocarpus heterophyllus (Moraceae), India, M.P., Jabalpur. 

arundinaceum P. Karst. – OUDEMANS (1924). An error; Clasterosporium was intended. 

arundinaceum Mont., Ann. Sci. Nat. Bot., Sér. 3, 12: 299 (1849)!. 

T: on sheaths and leaves of Arundo mauritanica (Poaceae), France, Marseille, Castagne. 

Lit.: SACCARDO (1886: 364). 

arundinicola Berl., Riv. Patol. Veg. 4: 19 (1895)!, as ‘arundinicolum’. 

T: on rotting culms of Arundo donax (Poaceae), Italy, Avellino. 

Lit.: SACCARDO (1913: 1371), FERRARIS (1914: 883), GONZÁLES-FRAGOSO (1927: 210). 

Ill.: BERLESE (1895: Tab. 3, Fig. 17). 

arundinis (Corda) Sacc., Syll. fung. 4: 364 (1886)!. 

T: on rotten stems of Arundo sp. (Poaceae), Czech Republic, Prag, Lieben, Corda (PRM 

155582). 

≡ Myxocladium arundinis Corda, Icon. fung. 1: 12 (1837). 



Lit.: LINDAU (1907: 814), HUGHES (1958: 750), ZHANG, WEI & ZHANG (1998: 196). 

Ill.: CORDA (Tab. 3, Fig. 172), ZHANG, WEI & ZHANG (1998: 196, Fig. 2). 

Notes: ZHANG, WEI & ZHANG (1998) treated C. arundinis as a distinct species and not as a 

synonym of the widespread Cladosporium herbarum. The fungus described and illustrated 

from China by ZHANG et al. (1998, 2003) as ‘Cladosporium arundinis’ is quite distinct and the 

name is misapplied. 

asperococcum Oudem., in Roum., F. gall. exs., Cent. XXXXVI, No. 4592 (1888)!. 

T: on dead branches of Sambucus nigra (Caprifoliaceae), the Netherlands, La Haye, May 

1888, C. Destrée, Roum., F. gall. exs. 4592 (L; FH). 

Lit.: OUDEMANS (1923; 1924, as ‘asperococcos’). 

Notes: ‘Contrib. Fl. Myc. Pays-Bas.’ (given on the label as place of publication). 

astericola Davis, Trans. Wisconsin Acad. Sci. 20: 428 (1922)!. 

T: on upper leaves and upper portions of stems of Aster umbellatus (Asteraceae), USA, 

Wisconsin, Mellen, 4 Aug. 1919, J.J. Davis (BPI 426143, WIS: syntypes). 

≡ Fusicladium astericola (Davis) K. Schub. & U. Braun, in Schubert, Mycotaxon 92: 59 

(2005)!. 


asterinae Deighton, Mycol. Pap. 118: 30 (1969)!. 

T: on colonies of Asterina contigua (Asterinaceae) on leaves of Dialium dinklagei (Caesalpi- 

niaceae), Sierra Leone, Kenema (Nongowa), 6 Dec. 1937, F.C. Deighton (IMI 11851b: 

holotype). 

11

≡ Parapericoniella asterinae (Deighton) U. Braun, Heuchert & K. Schub., in Heuchert, Braun 

& Schubert, Schlechtendalia 13: 59 (2005)!. 

Lit.: ELLIS (1976: 331), KHAN & SHAMSI (1982: 111–112). 

Ill.: DEIGHTON (1969: 31, Fig. 17), ELLIS (1976: 332, Fig. 250 A), KHAN & SHAMSI (1982: 

112, Fig. 1), HEUCHERT et al. (2005: 60, Fig. 23). 

asteroma Fuckel, Jahrb. Nassauischen Vereins Naturk. 23–24: 355 ‘1869’ (1870)!. 

T: on living leaves of Populus tremula (Salicaceae), Germany, between Homburg and Wehrheim, 

Fuckel, F. rhen. 2208 (e.g., HAL). 

≡ Napicladium asteroma (Fuckel) Allesch., Ber. Bayer. Bot. Ges. 5: 25 (1897)!. 

≡ Napicladium asteroma (Fuckel) Sacc., Malphigia 17: 421 (1902)!. 

= Oidium radiosum Lib., Pl. crypt. ard., Fasc. 3, 285 (1834). [T: BR; Lib., Pl. crypt. ard. 285]. 

≡ Fusicladium radiosum (Lib.) Lind var. radiosum, Ann. Mycol. 3: 430 (1905)!. 

≡ Fusicladium radiosum (Lib.) Lindau, in Rabenhorst, Krypt.-Fl., ed. 2, 1(8): 777 (1907)!. 

≡ Stigmina radiosa (Lib.) Goid., Ann. Bot. (Rome) 21: 11 (1936). 

≡ Pollaccia radiosa (Lib.) E. Bald. & Cif., in E. Baldacci, Atti Ist. Bot. “Giovanni Briosi” 10: 

61 (1937)!. 

≡ Venturia radiosa (Lib.) Ferd. & C. A. Jørg., Skovtraeernes Sygdomme 1: 125 (1938) (nom. 

anamorph.). 

= Cladosporium ramulosum Roberge ex Desm., Ann. Sci. Nat. Bot., Sér. 2, 18: 361 (1852)!, 

non Reissek, 1851. [T: PC]. 

≡ Fusicladium ramulosum Rostr., Tidsskr. Skovbr. 6: 294 (1883), nom. nov., as ‘(Roberge, in 

Desm.) Rostr.’. 

≡ Pollaccia ramulosa (Rostr.) Ondřej, Eur. J. Forest Pathol. 2: 143 (1972)!, nom. nov., as 

‘(Desm.) Ondřej’. 

= ? Cladosporium asteroma [Fuckel] var. macrosporum Sacc., Michelia 2(6): 126 (1882)!. 

= Fusicladium tremulae A.B. Frank, Hedwigia 22: 127 (1883)!. [T: B]. 

≡ Napicladium tremulae (A. B. Frank) Sacc., Syll. fung. 4: 482 (1886)!. 

= ? Cladosporium asteroma [Fuckel] var. microsporum Sacc., Syll. fung. 4: 357 (1886)!. 

≡ Fusicladium radiosum [(Lib.) Lindau] var. microsporum (Sacc.) Lindau, in Rabenhorst, 

Krypt.-Fl., ed. 2, 1(8): 777 (1907)!. 

= Fusariella populi Garb., Bull. Soc. Mycol. France 33: 89 (1917). 

Teleomorph: Venturia tremulae Aderh., Hedwigia 36: 81 (1897)! var. tremulae. 

Lit.: SACCARDO (1886: 357, 1913: 1376), LINDAU (1907: 777), LIND (1913: 520), BALDACCI 

& CIFERRI (1937: 61), SIVANESAN (1984: 618), SCHUBERT et al. (2003: 85). 

asteroma [Fuckel] var. macrosporum Sacc., Michelia 2(6): 126 (1880)!. 

T: on leaves of Populus alba (Salicaceae), France. 

= ? Fusicladium radiosum (Lib.) Lind, Ann. Mycol. 3: 429 (1905)!. 

Lit.: SCHUBERT et al. (2003: 85). 

asteroma [Fuckel] var. microsporum Sacc., Syll. fung. 4: 357 (1886)!. 

T: on leaves of Populus tremula (Salicaceae), Italy, Conegliano. 

= ? Fusicladium radiosum (Lib.) Lind, Ann. Mycol. 3: 429 (1905)!. 

Lit.: LINDAU (1907: 777), BALDACCI & CIFERRI (1937: 61), SIVANESAN (1984: 618), 

SCHUBERT et al. (2003: 85). 

asteromatoides Sacc., in Saccardo & Berlese, Atti Reale Ist. Veneto Sci. Lett. Arti, Ser. 6, 

3(4): 722 (1885)!. 

T: on legumes of Erythrina sp. (?) (Caesalpiniaceae), Tahiti, G. Brunaud, May 1884, G. 

Brunaud, Roum., F. gall. exs. 3292 (e.g., B). 

≡ Cladosporium asteromatoides Sacc. & Roum., in Roum., F. gall. exs., Cent. 33, No. 3292 

(1885)!, nom. illeg., homonym [host given as ‘Cassia sp.’ and authors as ‘Sacc. & Roum.’]. 

Lit.: SACCARDO (1886: 353). 

astroideum Ces., Flora 36: 204 (1853)!. 

T: on stems of Alisma sp. (Alismataceae), on stems and leaves of Typha sp. (Typhaceae), Italy, 

12

1852, Cesati, Klotzsch, Herb. viv. myc., No. 1787, mixed infection with Cladosporium 

lanciforme Ces. (e.g., B; HAL; M). 

Lit.: SACCARDO (1886: 366), LINDAU (1907: 813), FERRARIS (1912: 337), ZHANG et al. 

(1999: 37−38). 

Ill.: ZHANG et al. (1999: 37, Fig. 2). 

Notes: ZHANG et al. (1999) described this species as causing leaf spots on Sagittaria 

sagittifolia (Alismataceae) from China, Shaanxi. 

aterrimum Ellis & Everh., Proc. Acad. Nat. Sci. Philadelphia 1894: 378 (1895)!. 

T: on rotten wood, USA, Kansas, Rockport, Nov. 1893, E. Bartholomew, No. 1256 (NY 

313201; BPI 426152). 

= Helminthosporium binum Corda, Icon. fung. 6: 9 (1854), as ‘Helmisporium’. [T: PRM]. 

≡ Scolicotrichum binum (Corda) Sacc., Syll. fung. 4: 349 (1886)!. 

≡ Spadicoides bina (Corda) S. Hughes, Canad. J. Bot. 36: 806 (1958)!, as ‘binum’. 

= Virgaria uniseptata Berk. & M.A. Curtis, in Berkeley, Grevillea 3(28): 145 (1875)!. [T: K; 

NYS]. 

≡ Cladotrichum uniseptatum (Berk. & M.A. Curtis) Sacc., Syll. fung. 4: 373 (1886)!. 

≡ Scolicotrichum uniseptatum (Berk. & M.A. Curtis) Cooke, Grevillea 17(82): 41 (1888)!. 

≡ Diplococcium uniseptatum (Berk. & M.A. Curtis) S. Hughes, Canad. J. Bot. 31: 634 (1953)!. 

= Cladotrichum simplex Sacc., Ann. Mycol. 4: 278 (1906). [T: PAD]. 

= Cladotrichum tapesiae Sacc., Ann. Mycol. 6: 565 (1908). 

Lit.: SACCARDO (1895: 620), CASH (1952: 68), HUGHES (1953: 634), ELLIS (1963: 8–9). 

atriellum Cooke, Grevillea 6(40): 139 (1878)!. 

T: on decayed fruits of Yucca aloifolia (Agavaceae), USA, South Carolina, Aiken, Rav., F. 

amer. exs. 296 (K; NY; PH 01020433: syntypes). 

Lit.: SACCARDO (1886: 366). 

atriplicis Massee & Rodway, in herb. (?). 

On Atriplex cinerea (Chenopodiaceae), Australia, Tasmania. 

atroseptum Pidopl. & Deniak, in Pidoplichko, Gribnaja Flora Grubych Kormov: 268 (1953)!, 

nom. inval. 

T: isolated from damp straw, Ukraine. 

Ill.: PIDOPLICHKO (1953: 268, Fig. 69). 

atrum Link, Ges. Naturf. Freunde Berlin Mag. Neuesten Entdeck. Naturk. 7: 38 (1816)!: Fr., 

Syst. mycol. 3(2): 371 (1832)!. 

T: on dry stems of a herbaceous plant, Germany, Berlin, Link (B: examined by Hughes). 

= Dematium herbarum δ lignorum Alb. & Schwein., Consp. fung. lusat.: 368 (1805)!. 

Lit.: SACCARDO (1886: 362), LINDAU (1907: 831), HUGHES (1958: Mycelia sterilia). 

aureum Link, Ges. Naturf. Freunde Berlin Mag. Neuesten Entdeck. Gesammten Naturk. 7: 38 

(1816)!. 

T: on rocks, sent by Nees von Esenbeck. 

= ? Trentepohlia aurea (L.) Mart., Fl. crypt. erlang.: 351 (1817)! 

Notes: In NEES (1817): “Cladosporium aureum Link ... bildet mit Dematium petraeum und 

strigosum Pers. eine eigene, den Lichenen beizuordnende Gattung, die ich Amphiconium 

nenne.”. The genus Amphiconium Nees is synonymous with Trentepohlia Mart. 

auriculae (Cooke) J.C. David, Mycol. Pap. 172: 98 (1997)!. 

T: on leaves of Primula auricula (Primulaceae), Great Britain, Royal Horticultural Society 

(K). 

≡ Heterosporium auriculae Cooke, J. Roy. Hort. Soc. 27: 380 (1902). 

Ill.: DAVID (1997: 84, Fig. 19 A–D). 

Host(s)/substrate(s) & distribution: on Primula auricula; Europe (Great Britain). 

autumnale Kübler, Arch. Sci. Phys. Nat., Sér. 3, 2: 699 (1879)!. 

T: on leaves of Vitis sp. (Vitaceae), Switzerland, 1876, M. Kübler. 

13

avellaneum G.A. de Vries, Contr. Knowl. Genus Cladosporium: 56 (1952)! (f. avellaneum). 

T: isolated from ‘Nivea’ ointment, Netherlands, Utrecht, 19 May 1947 (ATCC 11273 = CBS 

186.54 = IMI 49620). 

≡ Cladosporium resinae [(Lindau) G.A. de Vries] f. avellaneum (G.A. de Vries) G.A. de 

Vries, Antonie van Leeuwenhoek J. Microbiol. Serol. 21: 167 (1955)!. 

= Sorocybe resinae (Fr.) Fr., Summa veg. Scand. 2: 468 (1849)!. 

Lit.: HO et al. (1999: 150), PARTRIDGE & MORGAN-JONES (2002: 344−348). 

Ill.: PARTRIDGE & MORGAN-JONES (2002: 347, Fig. 4). 

Notes: DOMSCH et al. (1980) have C. avellaneum G.A. de Vries as a synonym of C. resinae 

(Lindau) G.A. de Vries (≡ Hormoconis resinae (Lindau) Arx & G.A. de Vries). However, 

FARR et al. (1989) list Hormoconis resinae and Cladosporium avellaneum separately and do 

not list either as a synonym of the other. 

avellaneum f. albidum G.A. de Vries, Contr. Knowl. Genus Cladosporium: 56 (1952)!. 

T: isolated from ‘Nivea’ ointment, Netherlands, Utrecht, 19 May 1947 (CBS 185.54). 

≡ Cladosporium resinae [(Lindau) G.A. de Vries] f. albidum (G.A. de Vries) G.A. de Vries, 

Antonie van Leeuwenhoek J. Microbiol. Serol. 21: 167 (1955)!. 


Notes: monosporous isolate from the parent culture (DE VRIES 1952). 

avellaneum f. sterile G.A. de Vries, Contr. Knowl. Genus Cladosporium: 56 (1952)!. 

T: isolated from ‘Nivea’ ointment, Netherlands, Utrecht, 19 May 1947. 

≡ Cladosporium resinae [(Lindau) G.A. de Vries] f. sterile (G.A. de Vries) G.A. de Vries, 

Antonie van Leeuwenhoek J. Microbiol. Serol. 21: 167 (1955)!. 


Notes: monosporous isolate from the parent culture (DE VRIES 1952). 

avellaneum f. viride G.A. de Vries, Contr. Knowl. Genus Cladosporium: 56 (1952)!. 

T: isolated from ‘Nivea’ ointment, Netherlands, Utrecht, 19 May 1947 (ATCC 11274 ex-type 

= CBS 187.54 = IMI 49621). 


Lit.: DE VRIES (1955: 167). 

Notes: monosporous isolate from the parent culture (DE VRIES 1952). Hormodendrum resinae 

Lindau and Cladosporium avellaneum f. viride G.A. de Vries are identical (DE VRIES 1955). 

baccae Verwoerd & Dippen., S. African J. Sci. 27: 327 (1930)!. 

T: on fruits of Vitis vinifera (Vitaceae), South Africa, Stellenbosch, B.J. Dippenaar, No. 392 in 

herbarium of Len Verwoerd at Stellenbosch. 

Notes: Type material could not be traced in South Africa and is probably not preserved. 

bacilligerum Mont. & Fr., in Montagne, Ann. Sci. Nat. Bot., Sér. 2, 6: 31 (1836)!. 

T: on Alnus glutinosa (Betulaceae), France, Lyons, Rochecardon near ‘Lugdunum’. 

≡ Passalora bacilligera (Mont. & Fr.) Mont. & Fr., in Montagne, Syll. gen. sp. crypt.: 305 

(1856). 

≡ Scolecotrichum bacilligerum (Mont. & Fr.) J. Schröt., in Cohn, Krypt.-Fl. Schlesien, Bd. 

3(2), Heft 4: 498 (1897)!. 

Lit.: COOKE (1871: 584), DEIGHTON (1967: 5–8), CROUS & BRAUN (2003: 440). 

balladynae Deighton, Mycol. Pap. 118: 32 (1969)!. 

T: on Balladyna magnifica (Parodiopsidaceae) on leaves of Canthium vulgare (Rubiaceae), 

Uganda, near Masaka, May 1962, C.L.A. Leakey (IMI 98798i: holotype). 

Lit.: ELLIS (1976: 331), HEUCHERT et al.(2005: 61–62). 

Ill.: DEIGHTON (1969: 33, Fig. 18), ELLIS (1976: 332, Fig. 250 B), HEUCHERT et al. (2005: 62, 

Fig. 24). 

Notes: This species is clearly to be excluded from Cladosporium s. str. and seems to be 

congeneric with C. asterinae, but type material is too meagre for a final conclusion. 

14

anaticum Săvul., Bul. Sti. Acad. Republ. Populare Române 3(2): 227 (1951)!. 

T: on dry leaves of Dianthus kitaibelii (= Dianthus petraeus Waldst. & Kit. subsp. petraeus) 

(Caryophyllaceae), Romania, Severin, Moldova Nouă, 11 Jul. 1948. 

bantianum (Sacc.) Borelli, Riv. Anat. Patol. Oncol. 17: 618 (1960)!. 

T: isolated from cerebral granulomata, Italy, Florenz, 1912 (PAD: photomicrographs prepared 

by Saccardo). 

≡ Torula bantiana Sacc., Ann. Mycol. 10: 320 (1912)!. 

≡ Xylohypha bantiana (Sacc.) McGinnis, A.A. Padhye, Borelli & Ajello, J. Clin. Microbiol. 

23: 1150 (1986)!. 

≡ Cladophialophora bantiana (Sacc.) de Hoog, Kwon-Chung & McGinnis, in de Hoog, 

Guého, Masclaux, Gerrits van den Ende, Kwon-Chung & McGinnis, J. Med. Veterin. 

Mycol. 33: 343 (1995)!. 

= Cladosporium trichoides C.W. Emmons, in Binford, Thompson & Gorham, Amer. J. Clin. 

Pathol. 22: 541 (1952)!. 

= Cladosporium trichoides [C.W. Emmons] var. chlamydosporum Kwon-Chung, Mycologia 

75(2): 320 (1983)!. 

Lit.: MATSUSHIMA (1975), MCGINNIS & BORELLI (1981), KWON-CHUNG & BENNETT (1992: 

639), HO et al. (1999: 146), DE HOOG et al. (2000: 564), SCHELL (2003: 577). 

Notes: Xylohypha emmonsii A.A. Padhye, McGinnis & Ajello, formerly reduced to a synonym 

of Cladophialophora bantiana, was re-established as a separate species despite high nDNA 

homology values, as Cladophialophora emmonsii (A.A. Padhye, McGinnis & Ajello) de Hoog 

& A.A. Padhye, in Gerrits van den Ende & de Hoog, Stud. Mycol. 43: 160 (1999). 

baptisiae H.C. Greene, Amer. Midl. Naturalist 39(2): 456 (1948)!. 

T: on living leaves of Baptisia leucophaea (Fabaceae), USA, Wisconsin, Dane Co., Madison, 

University of Wisconsin Arboretum, 14 Jul. 1947, H.C. Greene (BPI 426163, WIS: syntypes). 

≡ Fusicladium baptisiae (H.C. Greene) K. Schub. & U. Braun, in Schubert, Mycotaxon 92: 62 

(2005)!. 


beijerinckii Oudem., on CABI page, Kirk et al. (n. d.). 

Notes: An error, Coryneum beyerinckii Oudem., Hedwigia 22: 115 (1883) [= Stigmina carpophila 

(Lév.) M.B. Ellis, Mycol. Pap. 72: 56 (1959)!] is intended. 

bellynckii Westend., Bull. Acad. Roy. Sci. Belgique 21(8): 240 (1854)!. 

T: on faded leaves of Cynanchum vincetoxicum (= Vincetoxicum hirundinaria Medik.) 

(Asclepiadaceae), Belgium, Bois de Dave, near Namur, Prof. Bellynck (BR). 

≡ Cercospora bellynckii (Westend.) Niessl, Hedwigia 15: 1 (1876)!. 

≡ Cercospora bellynckii (Westend.) Sacc., Nuovo Giorn. Bot. Ital. 8: 818 (1876)!. 

≡ Cercosporidium bellynckii (Westend.) X.J. Liu & Y.L. Guo, Acta Mycol. Sin. 1(2): 93 

(1982)!. 

≡ Mycovellosiella bellynckii (Westend.) Constant., Cryptog. Mycol. 3(1): 67 (1982)!. 

≡ Passalora bellynckii (Westend.) U. Braun, Mycotaxon 55: 228 (1995)!. 

= Cercospora vincetoxici Sacc. Syll. fung 15: 85 (1901)!. 

Lit.: SACCARDO (1886: 450), LINDAU (1910: 129), CHUPP (1954: 69), CROUS & BRAUN 

(2003: 78). 

berkheyae Syd., Ann. Mycol. 12: 267 (1914)!. 

T: on leaves of Berkheya sp. (Asteraceae), South Africa, Natal, Cramond, 2 Dec. 1913, No. 

6852 (S). 

≡ Fulvia berkheyae (Syd.) M.B. Ellis, More Dematiaceous Hyphomycetes: 315 (1976)!. 

≡ Mycovellosiella berkheyae (Syd.) U. Braun & Crous, Mycol. Res. 99(1): 32 (1995)!. 

≡ Passalora berkheyae (Syd.) U. Braun & Crous, in Crous & Braun, Mycosphaerella and its 

anamorphs: 1. Names published in Cercospora and Passalora, CBS Biodiversity Ser. 1: 

441 (2003)!. 

Lit.: SACCARDO (1931: 789), CROUS & BRAUN (1996: 250). 

15

etulignum Ellis & Galloway, in herb. 

On Betula nigra (Betulaceae), USA, Washington District of Columbia, 2 Oct. 1887, B.T. 

Galloway (BPI 426165). 

≡ Fusicladium betulignum Ellis & Galloway ex K. Schub. & U. Braun, sp. nov. 

bignoniae Schwein., Trans. Amer. Philos. Soc., N.S., 4(2): 277 (1832)!. 

T: on legumes of Bignonia radicans (Bignoniaceae), USA, Carolina, Pennsylvania, Bethlehem, 

No. 2600 (PH 01020422, 01020423: syntypes). 

Lit.: SACCARDO (1886: 353), GONZÁLES-FRAGOSO (1927: 206), SCHUBERT & BRAUN (2004: 

297–298). 

Notes: nomen dubium. 

bisporum Matsush., Icones Microfungorum a Matsushima Lectorum: 33 (1975)!. 

T: on a dead leaf of Musa ×paradisiaca (Musaceae), Japan, Iriomote Island, Okinawa, Mar. 

1974 (Matsushima herb. 4861). 

≡ Beejadwaya bispora (Matsush.) Subram., Kavaka 5: 97 (1977)!. 

Ill.: MATSUSHIMA (1975: Pl. 188, Figs 1–2). 

Notes: SUBRAMANIAN (1977) transferred the species into a new genus because of the 

characteristically shaped, 1-celled conidia consistently formed in chains of two. 

boenninghauseniae Togashi & Katsuki, in Katsuki, Kyushu Agric. Res. 8: 84 (1951)!. 

T: on leaves of Boenninghausenia albiflora (Rutaceae), Japan, Kyushu, Pref. Fukuoka, 

Mizunashi, Ito-mura, 15 Oct. 1950, S. Katsuki. 

Ill.: KATSUKI (1951: 84, Fig. 1). 

borassi Hasija, Indian Phytopathol. 19(1966): 373 (1967)!, as ‘borassii’. 

T: on leaves of Borassus flabellifer (Arecaceae), India, Madhya Pradesh, Jabalpur, Howbagh, 

Coll. Garden, 7 Sept. 1964, S.K. Hasija (IMI 109416c). 

Ill.: HASIJA (1967: 375, Fig. 2). 

Host(s)/substrate(s) & distribution: on Borassus flabellifer; Asia (India). 

bosciae (Sacc.) K. Schub., comb. et stat. nov. 

T: on leaves of Boscia senegalensis (Capparidaceae), Eritrea, Barca, Agordat, 23 Feb. 1909, 

A. Fiori (PAD). 

≡ Cladosporium compactum [Berk. & M.A. Curtis] f. bosciae Sacc., Ann. Mycol. 8: 340 

(1910)!, as ‘f. Bosciae’. 

≡ Cladosporium compactum [Berk. & M.A. Curtis] var. bosciae (Sacc.) Sacc., Syll. Fung. 22: 

1367 (1913).! 

Host(s)/substrate(s) & distribution: on Boscia senegalensis; Africa (Eritrea). 

brachormium Berk. & Broome, Ann. Mag. Nat. Hist., Ser. 2, VII: 99 (1851)!. 

T: on leaves of Fumaria officinalis (Fumariaceae), King’s Cliffe. 

Lit.: COOKE (1871: 584), SACCARDO (1886: 363). 

brachyelytri H.C. Greene, Trans. Wisconsin Acad. Sci. 53: 214 (1964)!. 

T: on living leaves of Brachyelytrum erectum (Poaceae), USA, Wisconsin, Sawyer Co., 

Flambeau State Forest near Oxbow, 22 Jul. 1964, H.C. Greene (BPI 426166, WIS: syntypes). 

= Passalora fusimaculans [(G.F. Atk.) U. Braun & Crous] var. barretoana U. Braun & 

Crous, in Crous & Braun, Mycosphaerella and its anamorphs: 1. Names published in 

Cercospora and Passalora. CBS Biodiversity Ser. 1: 453 (2003)!. 

Lit.: SCHUBERT (2005: 63–65). 


brachytrichum Corda, Icon. fung. 1: 14 (1837)!. 

T: on the inner side of the bark of Fagus sp. (Fagaceae), Czech Republic, near Reichenberg, 

Corda (PRM 155419, 155420: syntypes). 

≡ Didymotrichum brachytrichum (Corda) Bonord., Handb. Mykol.: 89 (1851)!. 

Lit.: SACCARDO (1886: 354), LINDAU (1907: 821). 

Ill.: CORDA (Tab. 4, Fig. 209). 

16

Notes: Excluded, no Cladosporium s. str. This species is cladosporium-like, but the conidiogenous 

loci and conidia hila are subdenticulate and not cladosporioid (coronate). The pale 

conidia remind one of those of ‘Cladosporium elatum’. 

brassicae (Ellis & Barthol.) M.B. Ellis, More Dematiaceous Hyphomycetes: 340 (1976)!. 

T: on old leaves of Brassica oleracea (Brassicaceae), USA, Kansas, Rooks Co., 22 Apr. 1897, 

E. Bartholomew 2399 (NY: holotype; Barthol., N. Am. F. 3589; K, IMI 99694: isotypes). 

≡ Cladotrichum brassicae Ellis & Barthol., Trans. Kansas Acad. Sci. 16: 167 (1899). 

≡ Heterosporium brassicae (Ellis & Barthol.) Arx, Genera Fungi Sporul. Pure Cult., ed. 3: 305 

(1981). 

Lit.: DAVID (1997: 82). 

Ill.: ELLIS (1976: 341, Fig. 259 A), DAVID (1997: 83, Fig. 18; 84, Fig. 19 E–G). 

Host(s)/substrate(s) & distribution: on Brassica oleracea; North America (USA). 

brassicicola Sawada, Special Publ. Coll. Agric. Natl. Taiwan Univ. 8: 195 (1959)!, nom. 

inval. 

T: on leaves of Brassica juncea (Brassicaceae), Taiwan, Pref. Taichung, Taichung, 17 Feb. 

1913, K. Sawada. 

Ill.: SAWADA (1959: Pl. 3, Figs 4–5). 

Notes: published without Latin diagnosis. 

brevicatenulatum Rebr. & Sizova, Novosti Sist. Nizsh. Rast. 15: 137 (1978)!, as ‘brevicatenulatum’. 

T: isolated from ancient cloth, Russia, Yaroslavskaya, Rostov, 1973, T.P. Sizova. 

Ill.: REBRICOVA & SIZOVA (1978: 138, Figs a–b). 

Notes: REBRICOVA & SIZOVA (1978) mention type material as ‘Cultura typica No. 10P in 

Laboratorio Centrali’. 

brevicompactum Pidopl. & Deniak, Mikrobiol. Zhurn. 5(2): 186, 194 (1938)!, as ‘brevicompactum’, 

nom. inval. 

T: isolated from soil. 

Lit.: PIDOPLICHKO (1953: 271). 

Ill.: PIDOPLICHKO & DENIAK (1938: 186, Fig. 3). 

Notes: Neither in Mikrobiol. Zhurn. (1938) nor in PIDOPLICHKO (1953) a Latin diagnosis is 

given. 

brevicompactum var. tabacinum Pidopl. & Deniak, in Pidoplichko, Gribnaja Flora Grubych 

Kormov: 272 (1953)!, nom. inval. 

T: isolated from fermentated tobacco leaves, Ukraine. 


brevipes Ellis & Barthol. → subsessile. 

brevipes House, Bull. New York State Mus. Nat. Hist. 219/220: 62 (1919–1920)!. 

Notes: Petrak appears to have errored. Although page 62 of vol 219/220 reads 

“CLADOSPORIUM BREVIPES, N. sp.” this portion of vol 219/220 is actually a reprint of 

Peck’s report of 1886 (1887), reproduced in vol 219/220 for reasons stated on page 36. Homer 

D. House authored in 1921 “Notes on Fungi, IV…from New York State Museum Bulletins 

219, 220” but House’s notes make no reference to C. brevipes or any other Cladosporium. It 

seems that Petrak’s reference to C. brevipes House is an error. C. letiferum House was likely 

created via the same error. Both actually pertain to Peck’s fungi of the same name. 

brevipes Peck, Rep. (Annual) New York State Mus. Nat. Hist. 40: 64 (1887)!. 

T: on living leaves of Quercus alba (Fagaceae), USA, New York, Menands, July, C.H. Peck 

(NYS 523: holotype). 

Lit.: SACCARDO (1892: 604). 

Notes: Excluded, taxonomic status unclear. 

breviramosum Morgan-Jones, in Morgan-Jones & Jacobsen, Mycotaxon 32(1): 228 (1988)!. 

T: on discoloured wallpaper, USA, Georgia, St. Simon’s Island, King and Prince Hotel, Dec. 

17

1987, B.J. Jacobsen (AUA: holotype). [ATCC 64696 (Ex-type culture) = ATCC 76215]. 

Lit.: HO et al. (1999: 119). 

Ill.: MORGAN-JONES & JACOBSEN (1988: 229, Fig. 2; 231, Pl. 1), HO et al. (1999: 117, Figs 4– 

5). 

Notes: Presumably not a Cladosporium, species clusters together with Amorphotheca resinae 

not within the large Davidiella-subclade (BRAUN et al. 2003). 

britannicum M.B. Ellis, More Dematiaceous Hyphomycetes: 328 (1976)!. 

T: on dead wood of Quercus sp. (Fagaceae), Great Britain, Wales, Pwee-y-Faeda Estate, 13 

May 1973, M.B. Ellis (IMI 175936). 

Lit.: ELLIS & ELLIS (1985: 51). 

Ill.: ELLIS (1976: 327, Fig. 245 C). 

bruhnei Linder, Bull. Natl. Mus. Canada 97: 259 (1947)!. 

T: on Hordeum vulgare (Poaceae), Germany, Halle and Berlin, K. Bruhne. 

≡ Hormodendrum hordei Bruhne, in W. Zopf, Beitr. Physiol. Morph. nied. Org. 4: 1 (1894), 

non C. hordei Pass., 1887. 

≡ Cladosporium herbarum [(Pers.: Fr.) Link] δ cerealium [Sacc.] f. hordei (Bruhne) Ferraris, 

Flora Ital. Crypt., Pars I, Fungi, Fasc.13: 882 (1914)!. 

≡ Cladosporium hordei (Bruhne) Pidopl., Gribnaja Flora Grubych Kormov: 268 (1953)!, nom. 

illeg., homonym, non C. hordei Pass., 1887. 

Lit.: SACCARDO (1899: 1076). 

Ill.: LINDER (1947: 289, Pl. 14, Fig. C). 

Notes: LINDER (1947) examined No. 1481a-5, presumably in the National Museum and stated 

the species closely resembling C. herbarum. 

brunneoatrum McAlpine, Fungus Dis. Citrus Trees Austral.: 15, 78 (1899)!. 

T: on fruits of Citrus aurantium (Rutaceae), Australia, New South Wales, Sydney, Jul. 1898 

(VPRI). 

Lit.: SACCARDO (1902: 1057). 

Ill.: MCALPINE (1899: Pl. 1; Figs 17–18). 

Notes: Excluded, no Cladosporium s. str., but status still unclear; type material very sparse. 

brunneolum Sacc., Syll. fung. 4: 358 (1886)!. 

T: on dead leaves of Hedera helix (Araliaceae), USA, California, Dec. 1880, Henkney, No. 

1954 (K 121546: holotype). Isotype: BPI 426168. 

≡ Cladosporium brunneum Cooke & Harkn., Grevillea 12: 96 (1884)!, nom. illeg., homonym, 

non C. brunneum Corda, 1837. 



brunneum Cooke & Harkn. → brunneolum. 

brunneum Corda, Icon. fung. 1: 15 (1837)!, as ‘bruneum’. 

T: on rotten leaves of Populus sp. (Salicaceae), Czech Republic, near Prag (not preserved). 

Neotype: on rotten leaves of Populus sp., Czech Republic, Prague, 1847, E.D. Hofmann (PRM 

657440). 

= Cladosporium herbarum var. macrocarpum (Preuss) M.H.M. Ho & Dugan, in Ho, 

Castañeda, Dugan & Jong, Mycotaxon 72: 131 (1999). 



buteicola Cooke, Grevillea 5(33): 15 (1876)!, as ‘buteacolum’. 

T: on legumes of Butea frondosa (Fabaceae), India, 1876, Colonel Hobsen (K 121548: 

holotype). 


Ill.: COOKE (1876, Pl. 74, Fig. 10), ELLIS (1976: 343, Fig. 260 B). 

butyri O. Jensen, Centralbl. Bakteriol., 2. Abth., 8: 311–312 (1902)!. 

T: on butter. 

18

= Monilia nigra Burri & W. Staub, Landw. Jahrb. Schweiz 23: 479 (1909)!. 

Notes: A separate, yeast-like form and other characters do not enable this species to be placed 

in Cladosporium (DE VRIES 1952: 90). This species was cited by DE VRIES (1952) as 

published in ‘Landw. Jahrb. Schweiz 15: 329 (1901)!’, but that reference did not contain a 

description of this species. The whole paper of BURRI & STAUB (1909: 479–513) dealt with 

Monilia nigra, but on page 479 the new name appeared for the first time. 

caducum Davis, Trans. Wisconsin Acad. Sci. 21: 298 (1924)!. 

T: on leaves of Betula nigra (Betulaceae), USA, Wisconsin, along the Wisconsin river, 21 Jul. 

1922, J.J. Davis (BPI 426179; WIS: syntypes). 

≡ Fusicladium caducum (Davis) K. Schub. & U. Braun, in Schubert, Mycotaxon 92: 65 

(2005). 


caesalpiniae Sawada, Rep. Gov. Res. Inst. Formosa 85: 91 (1943)!, nom. inval. 

T: on Caesalpinia nuga (Caesalpiniaceae), Taiwan, Kaohsiung Prov., 24 Mar. 1930, K. 

Sawada (BPI 426182, PPMH: syntypes). 

≡ Fusicladium caesalpiniae [Sawada] ex K. Schub. & U. Braun, Fungal Diversity (2005), in 

press. 


caespiticium Rabenh., F. eur., Cent. VI, No. 579 (1863)! and Bot. Zeitung (Berlin) 21: 230 

(1863)!, nom. nud. 

T: on leaves of Rhamnus alaternus (Rhamnaceae), ‘ad Gandam’, winter 1861, E. Coemans, 

Rabenh., F. eur. 579 (M-57455: lectotype). Isolectotypes: e.g., HAL; HBG; M-57454. 




caespitosum Ell. & Everh., in herb. 

On Arabis holboelii (Brassicaceae), USA, Utah, M.E. Jones, Utah Fungi 5743. 

Notes: Specimen with descriptive notes in Myc. Coll. (CASH 1952: 68). 

calamigenum Berk. & Broome, J. Linn. Soc., Bot. 14: 99, 1873 (1875)!, as ‘calamigena’. 

T: on the fruit of Calamus sp., India, Ceylon, south of the island, Jul. 1868, G.H.K. Thwaites 

(K 121549: isotype). 

Lit.: SACCARDO (1886: 367). 

Notes: Excluded, probably galls. 

calcareum Beeli, Bull. Soc. Roy. Bot. Belgique 56: 68 (1924)!. 

T: on lime-coated wall and wood in a cellar, Belgium, Brussels. 

Lit.: SACCARDO (1972: 1337). 

Ill.: BEELI (1923: Tab. 4, Fig. 14). 

callae Peck & Clinton, in herb. 

On leaves of Calla sp., USA, New York, Buffalo, G.W. Clinton (BPI 426184: syntype 

material of Cercospora callae Peck & Clinton). 

= Cercospora callae Peck & Clinton, Rep. (Annual) New York State Mus. Nat. Hist. 29: 52 

(1876). 

Lit.: CHUPP (1954: 57). 

calotropidis F. Stevens, Trans. Illinois State Acad. Sci. 10: 207 (1917)!. 

T: on leaves of Calotropis procera (Asclepiadaceae), Puerto Rico, Jul. 1915 (ILL 15842; IMI 

19791; K; MICH; PC: syntypes). 

= Cercospora calotropidis Ellis & Everh., Rep. (Annual) Missouri Bot. Gard. 9: 120 (1898)!. 

[T: BPI 433953, 433956; NY; IMI 7752 (slide)]. 

≡ Phaeoramularia calotropidis (Ellis & Everh.) Kamal, A.S. Moses & R. Chaudhary, Mycol. 

Res. 94: 716 (1990)!. 

≡ Passalora calotropidis (Ellis & Everh.) U. Braun, Schlechtendalia 5: 60 (2000)!. 

19

= Cercospora microsora Pat., in R.P. Duss, Champignons de la Guadeloupe, 3 e Sér.: 91 

(1902)!, nom. illeg., homonym, non C. microsora Sacc., 1880. 

≡ Cercospora patouillardii Sacc., Syll. fung. 18: 608 (1906)!. 

= Cercospora calotropidis Lingelsh., Bot. Jahrb. Syst. 39: 605 (1907)!, nom. illeg., homonym, 

non C. calotropidis Ellis & Everh., 1898. 

≡ Cercospora lingelsheimii Săvul. & Rayss, Ann. Cryptog. Exot. 8: 49 (1935)!. 

= Cercospora inconspicua Pat. & Har., Bull. Soc. Mycol. France 24: 16 (1909)!. [T: FH 

7807]. 

= Napicladium calotropidis H. Morstatt, Ann. Mycol. 10: 451 (1912)!. 

= Cercospora calotropidis Speg., Anales Mus. Nac. Hist. Nat. Buenos Aires 26: 132 (1914), 

nom. illeg., homonym, non C. calotropidis Ellis & Everh., 1898. 

≡ Cercosporina calotropidis Sacc., in Trotter, Syll. fung. 25: 897 (1931)!, as ‘(Speg.) Sacc.’. 

= Cercospora domingensis Gonz. Frag. & Cif., Bol. Soc. Esp. Hist. Nat. 26: 339 (1926). [T: 

BPI 435826; MA]. 

Lit.: SACCARDO (1931: 789), SUBRAMANIAN (1971: 293), CROUS & BRAUN (2003: 96). 

‘cancerogenes von Niessen, Canceromyces Auct. 1894’ (no reference to an original description 

has been found). 

T: isolated from a cancer of uterus. 

Lit.: VUILLEMIN (1931: 77), NANNIZZI (1934). 

Notes: “Wartmann credette di identificarlo con il C. herbarum o con una specie vicina, cioè 

con un saprofita banale” (NANNIZZI 1934). 

capsici Kovatsch., Z. Pflanzenkrankh. Pflanzenschutz 48(7): 335 (1938)!, nom. nov., as ‘(É.J. 

Marchal & Steyaert) Kovatsch.’. 

T: on Capsicum frutescens (Solanaceae), Congo belge, Prov. di l’Equateur. 

≡ Cercospora capsici É.J. Marchal & Steyaert, Bull. Soc. Roy. Bot. Belgique 61: 167 (1929), 

nom. illeg., homonym, non C. capsici Heald & F.A. Wolf, 1911. 

= Cercospora capsici Unamuno, Bol. Soc. Esp. Hist. Nat. 32: 161 (1932)!, nom. illeg., homonym, 

non C. capsici Heald & F.A. Wolf, 1911. 

≡ Cercospora unamunoi Castell., Rivista Agric. Subtrop. Trop. 42: 20 (1948). 

≡ Phaeoramularia unamunoi (Castell.) Munt.-Cvetk., Lilloa 30: 183 (1960)!, nom. inval. 

= Cercospora capsicicola Vassiljevsky, in Vassiljevsky & Karakulin, Parazitnye nesovershennye 

griby, Ch. I, Gifomicety: 344 (1937)!. 

≡ Phaeoramularia capsicicola (Vassiljevsky) Deighton, in Ellis, More Dematiaceous Hyphomycetes: 

323 (1976)!. 

≡ Phaeoramularia capsicicola (Vassiljevsky) Deighton, Trans. Brit. Mycol. Soc. 67: 140 

(1976)!, comb. superfl. 

≡ Passalora capsicicola (Vassiljevsky) U. Braun & F. Freire, Cryptog. Mycol. 23: 299 

(2002)!. 

Lit.: CHUPP (1954: 553), SACCARDO (1972: 1336), CROUS & BRAUN (2003: 103). 

cardariae Opiz, Lotos 5: 41 (1855)!. 

T: on silicles of Cardaria draba (Brassicaceae), Czech Republic, Prag, park Vimrovské sady, 

23 Nov. 1853, Opiz (PRM 657446, 657447: syntypes). 



Lit.: LINDAU (1907: 832). 

caricicola Corda, Icon. fung. 1: 14 (1837)!, as ‘caricicolum’. 

T: on leaves and culms of Carex (Cyperaceae), Czech Republic, near Reichenberg (PRM). 

≡ Didymotrichum caricicola (Corda) Bonord., Handb. Mykol.: 89 (1851)!. 



Lit.: SACCARDO (1886: 365), LINDAU (1907: 816), LIND (1913: 523), HUGHES (1958: 750). 


20

caricinum C.F. Zhang & P.K. Chi, Guangdong Guoshu Zhenjun Binghai Zhi.: 54 (2000)!. 

T: on living leaves and fruits of Carica papaya (Caricaceae), China, Guangzhou, 1993, C.F. 

Zhang, No. 01371. 

Lit.: ZHANG et al. (2003: 185–186). 

Notes: The name C. caricinum is not validly published since there is no indication where the 

type material has been deposited. 

carpesii Sawada, Bull. Gov. Forest Exp. Sta. 105: 93 (1958)!. 

T: on leaves of Carpesium abrotanoides var. thunbergianum (Asteraceae), Japan, Tohoku 

District, 12 Sept. 1947, K. Sawada. 

Notes: Type material of this species could not be traced, neither in PPMH nor BPI, and is 

probably not preserved. 

carpophilum Thüm., Oesterr. Bot. Z. 27: 12 (1877)!. 

T: on fruits of Prunus persica (Rosaceae), Austria, Klosterneuburg, Aug. 1877, Thümen 

(PAD: neotype). 

≡ Fusicladium carpophilum (Thüm.) Oudem., Verh. Kon. Ned. Akad. Wetensch., Afd. 

Natuurk., Tweede Sect. 1900: 388 (1900). 

≡ Megacladosporium carpophilum (Thüm.) Vienn.-Bourg., Les champignons parasite des 

plantes cultivées 1: 489 (1949). 

≡ Fusicladosporium carpophilum (Thüm.) Partridge & Morgan-Jones, Mycotaxon 85: 362 

(2003)!. 

= Fusicladium pruni Ducomet, Thèse Fac. Sci. Paris: 137 (1907). 

= Fusicladium amygdali Ducomet, Ann. École, Natl. Agric. Rennes 4: 11 (1911). 

= Cladosporium americanum H.C. Greene, Amer. Midl. Naturalist 41(3): 723 (1949)!. 

Teleomorph: Venturia carpophila E.E. Fisher, Trans. Brit. Mycol. Soc. 44: 339 (1961)!. 

Lit.: SACCARDO (1886: 353), LIND (1913), ELLIS (1971: 317), SIVANESAN (1974: 75; 1984: 

609), SCHUBERT et al. (2003: 26, 28–30), SCHUBERT (2005: 56–57). 

Notes: “Herb. myc. oec. 599, Wien, 1877 (LE: syntype)” was cited as type by SCHUBERT et al. 

(2003), but this collection is not the type. 

carrionii Trejos, Revista Biol. Trop. 2: 106 (1954). 

T: isolated from a case of chromoblastomycosis, Australia, Venezuela. 

≡ Cladophialophora carrionii (Trejos) de Hoog, Kwon-Chung & McGinnis, in de Hoog, 


Mycol. 33: 345 (1995)!. 

= Cladophialophora ajelloi Borelli, Pan Amer. Health Organ. Sci. Publ. 396: 335 (1980). 

Lit.: KWON-CHUNG & BENNETT (1992: 350), HO et al. (1999: 147), DE HOOG et al. (2000: 

570), SCHELL (2003: 578). 

caryigenum (Ellis & Langl.) Gottwald, Mycologia 74(3): 388 (1982)!. 

T: on leaves of Carya illinoensis (Juglandaceae), USA, Louisiana, St. Martin, 3 Sept. 1888, 

A.B. Langlois, Fl. Ludov. 1499 (NY: lectotype, selected by SCHUBERT et al., 2003). 

Isolectotypes: on leaves of Carya illinoensis (C. olivaeformis), USA, Louisiana, St. Martinsville, 

Sept. 1888, A.B. Langlois (BPI 426315, 426333; M). 

≡ Fusicladium caryigenum Ellis & Langl., J. Mycol. 4: 124 (1888)!. 

= Fusicladium effusum G. Winter, J. Mycol. 1: 101 (1885)!. 

Lit.: HO et al. (1999), SCHUBERT & BRAUN (2002a). 

caryigenum var. carpineum (Ellis & Everh.) Gottwald, Mycologia 74(3): 389 (1982)!, as 

‘carpinum’, comb. inval. 

T: on Carpinus americana (Corylaceae), Canada, London, Oct. 1889, J. Dearness (NY: 

lectotype, selected by SCHUBERT et al., 2003; DAOM, M: isolectotypes). 

≡ Fusicladium effusum var. carpineum Ellis & Everh., Proc. Acad. Nat. Sci. Philadelphia 

1891: 91 (1891)!. 

≡ Fusicladium carpineum (Ellis & Everh.) U. Braun & K. Schub., IMI Descriptions of Fungi 

and Bacteria 152, No. 1512 (2002)!. 

21

casei (Johan-Olsen) Sacc. & Traverso, in Saccardo, Syll. fung. 19: 296 (1910)!. 

T: in cheese, Norway. 

≡ Dematium casei Johan-Olsen, Centralbl. Bakteriol., 2. Abth., 3: 280 (1897)!. 

Lit.: SACCARDO (1913: 1371). 

Ill.: JOHAN-OLSEN (1897: Tab. 4–5, Figs 7–13). 

cassiae-surathensis J.M. Yen, Bull. Trimestriel Soc. Mycol. France 97(3): 130–131 (1981)!. 

T: on living leaves of Cassia surathensis (Caesalpiniaceae), Singapore, Bukit Timah, 29 Mar. 

1970, G. Lim, No. 45 (LAM: Yen Herb., No. 10541). 

Lit.: BRAUN & FREIRE (2004: 222). 

Ill.: YEN (1981: 130, Fig. 1). 

Host(s)/substrate(s) & distribution: on Cassia surattensis and Chamaecrista sp.; Asia (Singapore), 

South America (Brazil). 

castellanii Borelli & Marcano, Castellania 1(5): 154 (1973). 

T: isolated from a patient with ‘tinea nigra’, Venezuela (IMI 183818). 

= Stenella araguata Syd., Ann. Mycol. 28: 205 (1930)!. 

Lit.: MCGINNIS & PADHYE (1978: 415). 

catamarcense Speg., Anales Soc. Ci. Argent. 10: 63 (1880)!, as ‘catamarcensis’. 

T: on wilting leaves of Pachylaena atriplicifolia (Asteraceae), Argentina, ‘in arenosis alpinis 

de Catamarca’ (LPS 13.129: holotype). 

Lit.: SACCARDO (1886: 363), FARR (1973). 

Notes: Excluded, but taxonomic status remains unclear. 

cattleyae Verpl., Meded. Landbouwhoogeschool Opzoekingsstat. Staat Gent 3: 103 (1935). 

T: on dead leaves of Cattleya mossia (Orchidaceae), Belgium, Antwerpen, botanical garden, 

12 Mar. 1935 (GENT: holotype). 

= Dendryphiella vinosa (Berk. & M.A. Curtis) Reisinger, Bull. Soc. Mycol. France 84(1): 27 

(1968). 

Lit.: SCHUBERT & BRAUN (2004: 313–314). 

cellare (Pers.) Schanderl, Zentralbl. Bakteriol., 2. Abt., 94: 117 (1936)!. 

T: from a wine vault. 

≡ Racodium cellare Pers., Neues Mag. Bot. 1: 123 (1794)!. 

≡ Zasmidium cellare (Pers.) Fr., Summa veg. Scand. 2: 407 (1849)!. 

≡ Rhinocladiella cellaris (Pers.) M.B. Ellis, Dematiaceous Hyphomycetes: 248 (1971)!. 

= Byssus septica Roth, in Tent. Fl. Germ. 1: 516 (1788). 

= Byssus mollissima Ehrb., P. crypt., No. 217 (1790). 

= Byssus cryptarum DC., F. franç. 2: 67 (1805). 

= Antennaria cellaris Fr., Syst. mycol. 3: 229 (1832). 

= Rhinocladiella ellisii Hawksw., in Hawksworth & Riedl, Taxon 26(2–3): 208 (1977)!. 

Lit.: GONZÁLES-FRAGOSO (1927: 249), DE VRIES (1952), BARRON (1968: 266). 

cerasi (Rabenh.) Aderh., Centralbl. Bakteriol., 2. Abth., 7: 656 (1901)!. 

T: on fruits of Prunus cerasus (Rosaceae), Germany, Borussia (BRAUN 1853: Tab. 1, B, 1–2: 

iconotype). 

≡ Acrosporium cerasi Rabenh., in Braun, Verh. Vereins Beförd. Gartenbaues Königl. Preuss. 

Staaten 1: 176 (1853)!. 

≡ Fusicladium cerasi (Rabenh.) Erikss., Meddeland. Kongl. Lantbruksakad. Exp.-fält 1: 73 

(1885). 

≡ Fusicladium cerasi (Rabenh.) Sacc., Syll. fung. 4: 346 (1886)!, comb. superfl. 

≡ Fusicladiopsis cerasi (Rabenh.) Karak. & Vassiljevsky, in Vassiljevsky & Karakulin, Parazitnye 

nesovershennye griby, Ch. I. Gifomicety: 210 (1937)!. 

≡ Megacladosporium cerasi (Rabenh.) Vienn.-Bourg., Les champignons parasites des plantes 

cultivées 1: 537 (1949). 

≡ Karakulinia cerasi (Rabenh.) N.P. Golovina, Novosti Sist. Nizsh. Rast. 1: 213 (1964). 

Teleomorph: Venturia cerasi Aderh., Landw. Jahrb. 29: 541 (1900)!. 

Lit.: SIVANESAN & HOLLIDAY (1981), SIVANESAN (1984), SCHUBERT et al. (2003: 33–35). 

22

cercestidis Deighton, Mycol. Res. 94(4): 570 (1990)!. 

T: on living leaves of Cercestis congensis (Araceae), Sierra Leone, Njala (Kori), 25 Apr. 

1934, F.C. Deighton (IMI 7735: holotype). 

≡ Stenella cercestidis (Deighton) U. Braun, Schlechtendalia 5: 54 (2000)!, as ‘cercestis’, nom. 

illeg, non S. cercestidis (J.M. Yen & Gilles) Deighton (1979). 

≡ Stenella deightoniana (Deighton) U. Braun, in Braun & Crous, Mycotaxon (2005), in press. 

cerophilum (Tubaki) Matsush., Icones Microfungorum a Matsushima Lectorum: 34 (1975)!. 

T: on the blackened (originally white), powdery wax layer under the leaf sheathes of Sasa sp. 

(Poaceae), Japan, May 1955 (preserved in Nagao Institute). 

≡ Acrotheca cerophila Tubaki, J. Hattori Bot. Lab. 20: 143 (1958)!. 

≡ Ramichloridium cerophilum (Tubaki) de Hoog, Stud. Mycol. 15: 74 (1977)!. 

chaetomium Cooke, Grevillea 17(83): 66 (1889)!. 

T: on leaves of Euphorbia sp. (Euphorbiaceae), USA, New Jersey, Newfield, J.B. Ellis, No. 

2289 (K). 

≡ Cercosporidium chaetomium (Cooke) Deighton, Mycol. Pap. 112: 27 (1967)!. 

≡ Passalora chaetomium (Cooke) Arx, Proc. Kon. Ned. Akad. Wetensch. C, 86(1): 44 

(1983)!. 

≡ Passalora chaetomium (Cooke) Poonam Srivast., J. Liv. World 1(2): 114 (1994)!, comb. 

inval. 

= Scolecotrichum ? euphorbiae Tracy & Earle, Bull. Torrey Bot. Club 23: 209 (1896). [T: 

NY]. 

≡ Piricularia euphorbiae (Tracy & Earle), G.F. Atk., Cornell Univ. Sci. Bull. 3(1): 40 (1897)!. 

Lit.: SACCARDO (1892: 602), ELLIS (1971: 281), CROUS & BRAUN (2003: 445). 

chamaeropis (Unamuno) K. Schub., comb. nov. 

T: on leaves of Chamaerops humilis (Arecaceae), Spain, near Oriedo, May 1921, P. Unamuno 

(MA 06416: holotype). 

≡ Cladosporium fasciculare f. chamaeropis Unamuno, Trab. Secc. Cienc. Nat. Congr. Assoc. 

Progr. Cienc. Oporto 1921: 60 (1922). 

Host(s)/substrate(s) & distribution: on Chamaerops humilis; Europe (Spain). 

cheonis (Chupp & Linder) U. Braun, Biblioth. Lichenol. 86: 85 (2003)!. 

T: on leaves of Ilex sp. (Aquifoliaceae), China, Kiangsi Prov., Huang Yen Ssu, Hsing Tzu 

Hsien, 13 Sept. 1932, S.Y. Cheo, No. 922 (CUP 39400: holotype). 

≡ Cercospora cheonis Chupp & Linder, Mycologia 29: 27 (1937)!. 

Lit.: CHUPP (1954: 52), CROUS & BRAUN (2003: 119). 

Ill.: BRAUN (2003: 94, Fig. 8). 

Host(s)/substrate(s) & distribution: on Ilex sp.; Asia (China). 

chlamydeum Cif. & Redaelli, Mycopathol. Mycol. Appl. 8: 187 (1957)!. 

T: from skin of Canis. 

Notes: “Material probably lost; judging from the description this was Moniliella suaveolens” 

(DE HOOG et al. 2000). 

chlamydospora Matsush., Icones Microfungorum a Matsushima Lectorum: 34 (1975)!, as 

‘chlamydosporis’. 

T: from garden soil, Japan, Osaka, Ibaraki City, May 1967 (Herb. Osaka 1047). 

≡ Devriesia chlamydospora (Matsush.) Seifert & N.L. Nickerson, in Seifert et al., Canad. J. 

Bot. 82: 922 (2004)!. 

Ill.: MATSUSHIMA (1975: Pl. 55, Fig. 3), SEIFERT et al. (2004: ). 

chlorocephalum (Fresen.) E.W. Mason & M.B. Ellis, Mycol. Pap. 56: 123 (1953)!. 

T: on dead stems of Paeonia sp. (Paeoniaceae) (not preserved). 

≡ Periconia chlorocephala Fresen., Beitr. Mykol. 1: 21 (1850)!. 

≡ Haplographium chlorocephalum (Fresen.) Grove, Sci. Gossip 21: 198 (1885). 

≡ Graphiopsis chlorocephala (Fresen.) Trail, Scott. Naturalist (Perth) 10: 75 (1889). 

23

= ? Cladosporium paeoniae Pass., in Thümen, Herb. myc. oec., Fasc. IX, No. 416 (1876)! and 

Just’s Bot. Jahresber. 4: 235 (1876)!. 

= Periconia ellipsospora Penz. & Sacc., Atti Reale Ist. Veneto Sci. Lett. Arti, Ser. 6, 2: 596 

(1883–1884). 

= ? Cladosporium paeoniae [Pass.] var. paeoniae-anomalae Sacc., Syll. fung. 4: 362 (1886)!. 

= Haplographium chlorocephalum [(Fresen.) Grove] var. ovalisporum Ferraris, Flora Ital. 

Crypt., Pars I, Fungi, Fasc. 13: 875 (1914). 

Lit.: SACCARDO (1886: 362), LINDAU (1907: 822), FERRARIS (1912: 348), LIND (1913: 524), 

DE VRIES (1952: 94), ELLIS (1971: 309), SUBRAMANIAN (1971: 296–297), ELLIS & ELLIS 

(1985: 395), MCKEMY & MORGAN-JONES (1991a), HO et al. (1999: 120). 

Ill.: FRESENIUS (1850: Taf. 4, Figs 10–15), MASON & ELLIS (1953: 124–125, Figs 42–43), 

ELLIS (1971: 310, Fig. 214 B), MCKEMY & MORGAN-JONES (1991a: 137, Fig. 1; 139, Pl. 1; 

141, Fig. 2; 143, Pl. 2), HO et al. (1999: 122, Fig. 7). 

Notes: Excluded, possibly congeneric with the new genus Parapericoniella U. Braun, 

Heuchert & K. Schub. 

chodatii (Nechitsche) Sacc. & D. Sacc., Syll. fung. 18: 577 (1906)!. 

T: on fermentated rice (Oryza, Poaceae). 

≡ Dematium chodatii Nechitsche, Inst. Bot. Univ. Genève, Ser. 6, 5: 22 (1904)!, as ‘chodati’. 

≡ Candida chodati (Nechitsche) Berkhout, Die schimmelgeschlachten Monilia, Oidium, 

Oospora en Torula: 54(1923). 

Lit.: BILGRAMI et al. (1991). 

Ill.: NECHITSCHE (1904: 23–25, Figs 4–6). 

Notes: SUBRAMANIAN (1971) cited the species as possible synonym of Aureobasidium oleae 

(Castagne) Subram., now regarded as A. pullulans var. pullulans (de Bary) G. Arnaud. 

chrysanthemi Pidopl. & Deniak, in Pidoplichko, Gribnaja Flora Grubych Kormov: 272 

(1953)!, nom. inval. 

T: on fallen petals of chrysanthemum, Ukraine. 


chrysophylli Thaung, Trans. Brit. Mycol. Soc. 63(3): 620 (1974)!. 

T: on living leaves of Chrysophyllum cainito (Sapotaceae), Burma, Sintoung, east of Thazi, 24 

May 1973, Mya Thaung (IMI 17741: holotype). 

Ill.: THAUNG (1974: 620, Fig. 1). 

Host(s)/substrate(s) & distribution: on Chrysophyllum cainito; Asia (Burma). 

cinnamomeum (Racib.) Höhn., in Kabát & Bubák, F. imp. exs., Fasc. XIII, No. 643 (1910)!. 

T: on Cinnamomum sp. (Lauraceae), Indonesia, Java, Buitenzorg, Tjenkumeh, 1908, F. v. 

Höhnel, Kab. & Bub., F. imp. exs. 643 (PC). 

≡ Scolecotrichum cinnamomeum Racib., Paras. Alg. Pilz. Javas: 40 (1900)!. 

≡ Stenella cinnamomea (Racib.) U. Braun, Schlechtendalia 8: 37 (2002)!. 

= Stenella cinnamomi Hosag. & U. Braun, Indian Phytopathol. 48: 261 (1995). 

circaeae Y. Qin & Z.Y. Zhang, Mycosystema 18(2): 135 (1999)!. 

T: on living leaves of Circaea mollis (Onagraceae), China, Jiangxi, Lushan, 5 Oct. 1980, J.Y. 

Li & T.Y. Zhang, No. 41440 (MHYAU 03953: holotype). 

Ill.: QIN & ZHANG (1999: 135, Fig. 1), ZHANG et al. (2003: 69, Fig. 38; Pl. 13, Fig. 4). 

circinalis Grüss, Wochenschr. Brauerei 48(7): 67 (1931)!. 

T: on the surface of wort. 

Notes: From GRÜSS (1931): “Die Sporen entstehen am Ende der Hyphen in Kettenform oder 

durch Verschiebung in kleine Häufchen.” This collapse of chains into heads does not sound 

like Cladosporium. 

citri Briosi & Farneti, Atti Ist. Bot. Univ. Pavia, Ser. 2, 10: 19 (1907)!, nom. illeg., homonym, 

non C. citri Massee, 1899. 

T: on fruits of Citrus limon (Rutaceae), Italy, Sicily. 

≡ Cladosporium farnetianum Sacc., Syll. fung. 22: 1366 (1913)!. 

24

≡ Kurosawaia citri Hara, List of Japanese Fungi, ed. 4: 172 (1954), nom. nov., as ‘(Briosi & 

Farneti) Hara’. 

Lit.: FERRARIS (1912: 347). 

Notes: It is not a Cladosporium, but maybe Sphaceloma fawcetii Jenkins. 

citri Massee, Text book Pl. Diseas. 1899: 310 (1899)!. 

T: on leaves and fruits of Citrus limon (Rutaceae), USA (K ?). 

Lit.: SACCARDO (1913: 1367), JENKINS (1925). 

Notes: Massee gave no specimen or herbarium designations, nor a description, but on p. 311 

refers to “USDA Bull. No. 8” (SWINGLE & WEBER, 1896), in which the disease of Citrus is 

attributed to a Cladosporium, for which a short description is rendered without specifying 

specimens or herbaria. JENKINS (1925) discussed the taxonomy and nomenclature of the citrus 

scab fungus in detail and showed that C. citri was based on a true Cladosporium s. str., which 

was found by SWINGLE & WEBER (1896) on old lesions. This fungus was often confused with 

the causal agent of the citrus scab disease (e.g., FAWCETT 1907, 1916), for which JENKINS 

(1925) introduced the name Sphaceloma fawcettii Jenkins (= Cladosporium citri sensu 

Fawcett, non Massee). Nevertheless, FAWCETT (1936: 535) still wrote on the identity of C. 

citri Massee that the fungus is now regarded as a Sphaceloma. 

citri Penz. – an error. See FAWCETT (1910 and 1936: 536). 

cladosporioides (Fresen.) G.A. de Vries, Contr. Knowl. Genus Cladosporium: 57 (1952)!. 

T: on overwintered leaves of Hydrangea sp. (Hydrangeaceae), Germany (not preserved). 

≡ Penicillium cladosporioides Fresen., Beitr. Mykol. 1: 22 (1850)!. 

≡ Hormodendrum cladosporioides (Fresen.) Sacc., Michelia 2(6): 148 (1880). 

≡ Cladosporium herbarum [(Pers.: Fr.) Link] f. hormodendroides Ferraris, Flora Ital. Crypt., 

Pars I, Fungi, Fasc. 8: 332 (1912)!. 

= ? Ramularia meliloti Ellis & Everh., Erythea 2: 26 (1894). 

= Monilia humicola Oudem., Arch. Néerl. Sci. Exact. Nat. 7: 286 (1902). 

Lit.: YAMAMOTO (1959: 3), ELLIS (1971: 319), SUBRAMANIAN (1971: 285), DOMSCH et al. 

(1980: 202), ELLIS & ELLIS (1985: 290, 468), WANG & ZABEL (1990), BRAUN (1998: 301), 

HO et al. (1999: 121), SAMSON et al. (2000: 108), DE HOOG et al. (2000: 583), SAMSON et al. 

(2001: 340), SCHUBERT & BRAUN (2004: 304). 

Ill.: FRESENIUS (1850: Taf. 3, Figs 23–28), DE VRIES (1952: 58–59, Figs 10–11), YAMAMOTO 

(1959: 4, Figs 9–12), ELLIS (1971: 318, Fig. 219 C), DOMSCH et al. (1980: 203, Fig. 82), HO et 

al. (1999: 122, Figs 8–9), DE HOOG et al. (2000: 583–584, Figs), SAMSON et al. (2000: 108, 

Fig. 48; 109, Pl. 46), SCHELL (2003: 582, Fig. 16). 

Host(s)/substrate(s) & distribution: parasitic causing leaf spots and as a secondary invader on 

many different plants, and saprobic, isolated from air, soil, textiles, etc.; cosmopolitan. 

Notes: Type material of Fresenius is housed in the Senckenberg-Herbarium Frankfurt, but type 

material of P. cladosporioides has not yet been traced. DE VRIES (1952) cited Cladosporium 

hypophyllum Fuckel as a synonym of C. cladosporioides but this species proved to be quite 

distinct from C. cladosporioides. 

cladosporioides [(Fresen.) G.A. de Vries] f. sp. pisicola (W.C. Snyder) G.A. de Vries → 

pisicola. 

cladrastidis Naumov, Bull. Soc. Mycol. France 30: 80 (1914)!. 

T: on leaves of Cladrastis amurensis (= Maackia amurensis) (Fabaceae), Russia, Far east, 

Juzhno-Ussurijskij Kray, 24 Jul. 1912, No. 10, N. Naumov (PC: holotype). 

= Cercospora cladrastidis Jacz., Hedwigia 39: 123 (1900). [T: HBG; LE 40382]. 

≡ Pseudocercospora cladrastidis (Jacz.) J.K. Bai & M.Y. Cheng, Acta Mycol. Sin. 11: 121 

(1992)!. 


clappieri – listed in Unesco (1955). 

clavatum Schwabe, Fl. anhalt. 2: 349 (1839)!. 

T: on dry wood of Quercus sp., Germany. 

25

Lit.: SACCARDO (1913: 1369). 

Ill.: SCHWABE (1839: Tab. 6, Fig. 13). 

clemensiae P.W. Graff, in Merrill, Philipp. J. Sci. 9: 40 (1914)!. 

T: on Eragrostis tenella (Poaceae), Guam, Agaña, 27 Nov. 1910, M.S. Clemens. 

coelosporum Spreng., Syst. veg. 4(1): 553 (1827)!. 

T: on stems of Gramineae, Germany (?). 

= Dematium articulatum Pers., Neues Mag. Bot. 1: 121 (1794)!. 

= Helminthosporium carispermum Link, ?, as ‘Helmisporium’. 

Notes: Dematium articulatum Pers. has also been cited as synonym of C. fasciculare (Pers.) 

Fr. 

colocasiae Sawada, Trans. Nat. Hist. Soc. Taiwan 25: 125 (1916). 

T: on Colocasia antiquorum (= C. esculenta) (Araceae), Taiwan, 2 Jun. 1910, K. Sawada 

(PPMH). 

= Cladosporium colocasiicola Sawada, Special Publ. Coll. Agric. Natl. Taiwan Univ. 8: 195 

(1959)!, nom. inval., syn. nov. 

Lit.: BUGNICOURT (1958), ELLIS (1971: 312), MATSUSHIMA (1975: 34), DAVID (1988a), HO 

et al. (1999: 123). 

Ill.: BUGNICOURT (1958: 235, Fig. 1), ELLIS (1971: 313, Fig. 216 B), MATSUSHIMA (1975: Pl. 

77), DAVID (1988a: Fig.), DE & CHATTOPADHYAY (1994: 228, Fig. 4), HO et al. (1999: 124, 

Figs 10–11). 

Host(s)/substrate(s) & distribution: on leaves of Colocasia esculenta; Africa, Asia, Europe, 

North America, Australia, New Zealand. 

colocasiicola Sawada, Special Publ. Coll. Agric. Natl. Taiwan Univ. 8: 195 (1959)!, nom. 

inval. 

T: on leaves of Colocasia esculenta (Araceae), Taiwan, Pref. Taipei, Taipei, 15 Oct. 1919, 

E.K. and Pref. Chiayi, Shuishan, 2 Nov. 1909, K. Sawada. 

= Cladosporium colocasiae Sawada, Trans. Nat. Hist. Soc. Taiwan 25: 125 (1916)!. 


Notes: without a Latin diagnosis, not validly published; two different collections cited without 

designating a type specimen. 

comesii Carbone, Atti Ist. Bot. Univ. Pavia, Ser. 2, 14: 322 (1914)!. 

T: isolated from sausage [‘in botulis (Salame crudo)’], Italy, Pavia. 

Lit.: SACCARDO (1931: 799). 

compactiusculum Sacc. & P. Syd., Syll. fung. 14: 1082 (1899)!. 

T: on twigs of Sterculia foetida (Sterculiaceae) and Frusinalia (Combretaceae), Ceylon. 

≡ Cladosporium subcompactum Roum. & P. Karst., in Karsten, Roumeguère & Hariot, Rev. 

Mycol. (Toulouse) 12: 80 (1890)!, nom. illeg., homonym, non C. subcompactum Sacc., 

1886. 

≡ Cladosporium zeylanicum Sacc. & Trotter, Syll. fung. 22: 1371 (1913)!, nom. superfl. 

Notes: The original material from PC is deposited under ‘Cladosporium subcompactum B. et 

C.’ (on Sterculia foetida, Ceylon) 

compactum Berk. & M.A. Curtis, in Berkeley, Grevillea 3(27): 106 (1875)!. 

T: on leaves of Arundinaria sp. (Poaceae), North America, No. 3767 (IMI 69771; K; STR). 

≡ Cercosporidium compactum (Berk. & M.A. Curtis) Deighton, Mycol. Pap. 112: 59 (1967)!. 

≡ Passalora compacta (Berk. & M.A. Curtis) U. Braun & Crous, in Crous & Braun, Mycosphaerella 

and its anamorphs: 1. Names published in Cercospora and Passalora, CBS 

Biodiversity Ser. 1: 133 (2003)!. 

= Cercospora scolecotrichoides G.F. Atk., Cornell Univ. Sci. Bull. 3(1): 46 (1897)!. [T: CUP; 

IMI 95405]. 


compactum [Berk. & M.A. Curtis] f. bosciae Sacc., Ann. Mycol. 8: 340 (1910)!. 

T: on leaves of Boscia senegalensis (Capparidaceae), Eritrea, Barca, Agordat, 23 Feb. 1909, 

26

A. Fiori (PAD). 

≡ Cladosporium compactum [Berk. & M.A. Curtis] var. bosciae (Sacc.) Sacc., Syll. Fung. 22: 

1367 (1913)!. 

≡ Cladosporium bosciae (Sacc.) K. Schub., comb. et stat. nov. 

compactum Sacc. → subcompactum Sacc. 

compactum [Sacc.] *punctatum Sacc. → punctatum Sacc. 

condylonema Pass., in Briosi & Cavara, F. paras., No. 79 (1889)!. 

T: on leaves of Prunus domestica (Rosaceae), Italy, Parma, Jun. 1889, Briosi & Cav., F. paras. 

79 (BPI 426388; HAL; MICH: syntypes). 



Lit.: SACCARDO (1892: 604), ADERHOLD (1901: 814), LINDAU (1907: 824), FERRARIS (1912: 

341), BRAUN (2001: 53). 

Ill.: ADERHOLD (1901: Tab. 18, Fig. 10). 

Notes: The species belongs to C. herbarum s. lat. and is morphologically intermediate between 

var. herbarum and var. macrocarpum (BRAUN 2001). 

confusum Matsush., Matsushima Mycol. Mem. 3: 4 (1983)!. 

T: on bark of Acer saccharum (Aceraceae), Canada, Ontario. 

Ill.: MATSUSHIMA (1983: 28–29, Figs 145–147). 

Notes: MATSUSHIMA (1983) depicts phialosporous form. 

congestum Berk. & Broome, J. Linn. Soc., Bot. 14: 99, 1873 (1875)!. 

T: on leaves of Litsea (Lauraceae), India, Ceylon (K 115280; PC). 

= Spiropes scopiformis (Berk.) M.B. Ellis, Mycol. Pap. 114: 30 (1968)!. 

Lit.: SACCARDO (1886: 359). 

Notes: see C. scopiforme Berk. The material from PC reads: in Litzoa, Ceylon, Dothidea, 

Berkeley. 

coralloides W. Yamam., Sci. Rep. Hyogo Univ. Agric., Ser. Agric. 4(1): 5 (1959)!, nom. 

inval. 

T: isolated from Ficus carica (Moraceae) and Oryza sativa (Poaceae), Japan. 

Lit.: HO et al. (1999: 125). 

Ill.: YAMAMOTO (1959: 6, Figs 17–20), HO et al. (1999: 124, Figs 12–13). 

Notes: This species was not validly published, because the author did not designate a type. 

The ‘lectotype’ chosen in HO et al. (1999) is also incorrect since it is not an element from the 

protologue of the original description. Hence, a formal validation of this name is necessary, 

which will be proposed in a separate paper based on a re-examination of this fungus. 

Excluded, but generic affinity not yet clear. 

corchori Z.Y. Zhang & T. Zhang, Plant Diseases and Their Control: 103 (1998)!. 

T: on Corchorus capsularis (Tiliaceae), China, Shaanxi, Yizhan, 20 Aug. 1973, J.Y. Li & T.Y. 

Zhang, No. 0412 (MHYAU 03955: holotype). 

Ill.: ZHANG, LIU, WEI & HE (1998: 104, Fig. 1), ZHANG et al. (2003: 79, Fig. 43). 

coreopsidis H.C. Greene, Trans. Wisconsin Acad. Sci. 45: 190 (1956)!. 

T: on living leaves of Coreopsis palmata (Asteraceae), USA, Wisconsin, Dane Co., Madison, 

University of Wisconsin Arboretum, 27 Jun. 1955 (BPI 426392; WIS: syntypes). 

≡ Fusicladium coreopsidis (H.C. Greene) K. Schub. & U. Braun, in Schubert, Mycotaxon 92: 

67 (2005)!. 


cornigenum Bubák, in Handel-Mazzetti, Ann. K.K. Naturhist. Hofmus. 23: 106 (1909)!. 

T: on living leaves of Cornus australis (= C. sanguinea) (Cornaceae), Turkey, Stephanos, near 

Trapezunt, 7 Jul. 1907, Handel-Mazzetti, No. 214 (BPI 426393: holotype). 

Lit.: SACCARDO (1913: 1367). 

Notes: Excluded, conidiogenous cells probably monophialidic. 

27

corrugatum McAlpine, Fungus Dis. Citrus Trees Austral.: 88 (1899)!. 

T: on both surfaces of green orange leaves (Citrus aurantium, Rutaceae), Australia, Armadale 

near Melbourne, Jan. 1899 (VPRI 5924: holotype). 

Lit.: SACCARDO (1913: 1367). 

Ill.: MCALPINE (1899: Fig. 57). 

Host(s)/substrate(s) & distribution: on Citrus aurantium; Australia. 

corynitrichum Ellis & Everh., in Millspaugh, West Virginia Geol. Surv., Ser. A, 5: 36 (1913), 

nom. nud. 

T: on dead fallen leaves of Magnolia fraseri (Magnoliaceae), USA, West Virginia, Fayette 

Co., 29 Nov. 1895, L.W. Nuttall (NY; WIS). 

Lit.: CASH (1952: 68). 

coryphae (Syd. & P. Syd.) J.C. David, Mycol. Pap. 172: 99 (1997)!. 

T: on leaves of Corypha elata (Arecaceae), Philippines, Mindoro, San José, Jan. 1912, P.W. 

Graff, Syd., F. exot. exs. 48 (S: lectotype, selected by DAVID, 1997; M, K, IMI 10041: 

isolectotypes). 

≡ Heterosporium coryphae Syd. & P. Syd., Philipp. J. Sci. 8: 196 (1913). 

Ill.: DAVID (1997: 89, Fig. 22 I–K; 101, Fig. 27). 

Host(s)/substrate(s) & distribution: on Corypha elata; Asia (Philippines). 

cubense R.F. Castañeda, Fungi Cubensis II: 4 (1987)!. 

T: on fallen leaves of Ficus sp. (Moraceae), Cuba, prov. Guantánamo, Maisí, 24 Apr. 1986, 

Mayra Camino (INIFAT C86/134: holotype). 



cubisporum Berk. & M.A. Curtis, in Berkeley, Grevillea 3(27): 107 (1875)!. 

T: on Ribes (Grossulariaceae), USA, Maine, Rev. J. Blake, No. 6318 (K). 

≡ Coremiella cubispora (Berk. & M.A. Curtis) M.B. Ellis, Dematiaceous Hyphomycetes: 33 

(1971)!. 

≡ Briosia cubispora (Berk. & M.A. Curtis) Arx, Antonie van Leeuwenhoek J. Microbiol. 

Serol. 38(3): 293 (1972)!. 

Lit.: SACCARDO (1886: 355). 

cucumerinum Ellis & Arthur, Bull. Agric. Exp. Sta., Indiana 19: 9–10 (1889). 

T: on fruits of Cucumis sativus (Cucurbitaceae), USA, New York, Geneva, J.C. Arthur (NY). 

= Scolecotrichum melophthorum Prill. & Delacr., Bull. Soc. Mycol. France 7(1): 219 (1891). 

≡ Macrosporium melophthorum (Prill. & Delacr.) Rostr., Gartn.-Tidende 24: 18 (1893). 

= Cladosporium cucumeris A.B. Frank, Z. Pflanzenkrankh. 3: 31 (1893)!. 

= Cladosporium scabies Cooke, Gard. Chron., Ser. 3, 34: 100 (1903)!. 

= Cladosporium cucumerinum var. europaeum Bubák, in herb. (BPI), syn. nov. 

Lit.: SACCARDO (1892: 601), LINDAU (1907: 830, 1910: 797), FERRARIS (1912: 349), 

GONZÁLES-FRAGOSO (1927: 206), CASH (1952: 68), DE VRIES (1952: 62), ELLIS (1971: 318), 

ELLIS & HOLLIDAY (1972), ELLIS & ELLIS (1985: 339), VON ARX (1987: 193), MCKEMY & 

MORGAN-JONES (1992), HO et al. (1999: 125). 

Ill.: DE VRIES (1952: 63, Fig. 12), ELLIS (1971: 318, Fig. 219 B), ELLIS & HOLLIDAY (1972: 

Fig.), VON ARX (1987: 194, Fig. 83b), MCKEMY & MORGAN-JONES (1992: 165, Fig. 1; 167, 

Pl. 1), HO et al. (1999: 124, Fig. 14). 

Host(s)/substrate(s) & distribution: on leaves, stems and fruits of Cucurbitaceae, especially 

Cucumis sativus, C. melo and Cucurbita pepo, other host genera Citrullus, Lagenaria, Luffa, 

Momordica, Sechium; cosmopolitan. 

Notes: In DE VRIES (1952) and MCKEMY & MORGAN-JONES (1992) ‘Macrosporium 

cucumerinum Ellis & Everh., Hedwigia 7: 49 (1896)’ is cited as a synonym of Cladosporium 

cucumerinum, but in Hedwigia, vol. 7 (published in 1868, not in 1896) there is no reference to 

this name. On CABI page (Index fungorum) the original citation of Macrosporium cucumeri- 

28

num is given as ‘Proc. Acad. Nat. Sci. Philadelphia 1895: 440 (1895)’ and ‘Alternaria 

cucumerina (Ellis & Everh.) A. Elliott, Amer. J. Bot. 4: 472 (1917)’ is given as current name. 

HASIJA (1967) described this species from India on Solanum tuberosum. In Korea it was 

isolated from Solanum melongena (KWON et al. 1999), indicating that C. cucumerinum is not 

confined to hosts of the family Cucurbitaceae. However, additional detailed inoculation 

experiments and molecular examinations are necessary to prove the host range of this species. 

cucumerinum var. europaeum Bubák, in herb. 

On Cucumis sativus (Cucurbitaceae), Austria, Carinthia, Poertschack at the Wörther See, Aug. 

1902, E. Cerny (BPI 426422). 

= Cladosporium cucumerinum Ellis & Arthur, Bull. Agric. Exp. Sta., Indiana 19: 9–10 

(1889). 

cucumeris A.B. Frank, Z. Pflanzenkrankh. 3: 31 (1893)!. 

T: on fruits of Cucumis sativus (Cucurbitaceae), Germany, Erkner near Berlin (B 70-6228: 

holotype). 


(1889). 

Lit.: LINDAU (1907: 830), LIND (1913: 524). 

cumulus Preuss, Linnaea 25: 726 (1851)!, as ‘cumulum’. 

T: on fallen branches, Germany, Hoyerswerda (B). 


cycadis Marcolongo, Riv. Patol. Veg., Ser. 2, 7(1): 8 (1914)!. 

T: on leaves of Cycas revoluta (Cycadaceae), Italy, Napoli (type at ‘R. Istituto Botanico di 

Napoli’?). 

Lit.: SACCARDO (1931: 790), ZHANG, PENG, LIU & ZHANG (1998: 6). 

Ill.: ZHANG, PENG, LIU & ZHANG (1998: 5, Fig. 2). 

Notes: “Cladosporium cycadis C. Massal., Riv. Patol. Veg. 7: 6 (1914)”, on Cycas revoluta, 

Italy (CIFERRI, Quaderno 19: 324 (1961), an error, see C. cycadis Marcolongo). 

cyclaminis Massey & Tilford, Phytopathology 22(1): 19 (1932)!. 

T: on Cyclamen sp. (Primulaceae), USA, ‘received Aug. 1932 from L.M. Massey’ (BPI 

426434: lectotype, part of type culture). 

= Ramularia cyclaminicola Trel., Trans. Illinois State Acad. Sci. 9: 145 (1916). [T: ILL 

14246: holotype]. 

Lit.: BAKER et al. (1950), BRAUN (1998: 226), ZHANG, ZHANG, LIU & HE (1999: 38). 

Notes: This species, previously only known from North America, has recently been reported 

from Asia (China) by ZHANG, ZHANG, LIU & HE (1999). 

cyrtomii Z.Y. Zhang, H.H. Peng & H. Zhang, in Zhang, Peng, Liu & Zhang, Mycosystema 

17(1): 4 (1998)!. 

T: on living leaves of Cyrtomium caryotideum (Dryopteridaceae), China, Prov. Yunnan, Gejiu, 

9 Dec. 1994, Wang Ying-xiang & Li Mao-lan (MHYAU 04048). 

Ill.: ZHANG, PENG, LIU & ZHANG (1998: 4, Fig. 1), ZHANG et al. (2003: 84, Fig. 48; Pl. 11, 

Fig. 4). 

cyttariicola Speg., Physis (Buenos Aires) 7(23): 20 (1923)!, as ‘cyttariicolum’. 

T: on Cyttaria harioti (Cyttariaceae, Ascomycetes), Argentina, Tierra del Fuego, Puerto 

Garibaldi (LPS 13.078). 

Lit.: FARR (1973: 251, as ‘cyttariicolum’), HEUCHERT et al. (2005: 63). 

Notes: excluded. 

daphniphylli Sawada, Rep. Gov. Res. Inst. Formosa 85: 91 (1943)!, nom. inval. 

T: on Daphniphyllum glaucescens (Daphniphyllaceae), Taiwan, 8 Feb. 1931, K. Sawada 

(PPMH: holotype). 

Notes: diagnosis only in Japanese, not validly published. Excluded, but generic affinity not yet 

clear. 

29

decolorans McAlpine, in herb. 

On Cynosurus cristatus (Poaceae), Australia (VPRI). 

delectum Cooke & Ellis, Grevillea 6(37): 6 (1877)!. 

T: on dead leaves of Magnolia glauca (Magnoliaceae), USA, New Jersey, Newfield (K; NY). 

Lit.: SACCARDO (1886: 358), LINDAU (1907: 822), FERRARIS (1912: 341), CASH (1952: 68). 

Ill.: COOKE & ELLIS (1877: Pl. 96, Fig. 36). 

delectum [Cooke & Ellis] f. ailanthi-glandulosae Thüm., Mycoth. univ., Cent. XVII, No. 

1666 (1880)!, nom. nud. 

T: on leaves of Ailanthus glandulosa (= A. altissima) (Simaroubaceae), USA, South Carolina, 

Aiken, H.W. Ravenel, Thüm., Mycoth. univ. 1666 (e.g., HAL). 

delicatulum Cooke, Grevillea 5(33): 17 (1876)!. 

T: on dead leaves, India, Colonel Hobsen, No. 23 (K 121551: isotype). 

Lit.: SACCARDO (1886: 361). 

dematiosum Ellis & Langl., in herb. 

On rotten stems of oak (Quercus sp., Fagaceae), USA, Louisiana, St. Martin, 23 Mar. 1888, 

A.B. Langlois, Fl. Ludoviciana 1264 (NY). 

Notes: Specimen in Myc. Coll. (CASH 1952: 68). 

dendriticum Desm. – GOLA (1930). 

dendriticum Wallr., Fl. crypt. Germ. 2: 169 (1833)!. 

T: on leaves of Pyrus malus (= Malus sylvestris) (Rosaceae), Germany, Thuringia (B, STR: 

syntypes). 

≡ Fusicladium dendriticum (Wallr.) Fuckel, Jahrb. Nassauischen Vereins Naturk. 23–24: 357 

‘1869’ (1870)!. 

≡ Passalora dendritica (Wallr.) Sacc., Mycoth. ven., Cent. XII, No. 1246 (1876)!, [Michelia 

1(2): 265 (1878)]. 

= Fusicladium pomi (Fr.) Lind, Dan. fung.: 521 (1913)!. 

Teleomorph: Venturia inaequalis (Cooke) G. Winter, Hedwigia 36: 81 (1897)!. 

Lit.: COOKE (1871: 583), SACCARDO (1886: 345), LINDAU (1907: 779), SIVANESAN (1984: 

616, as Spilocaea pomi), RITSCHEL (2001), CROUS & BRAUN (2003: 485), SCHUBERT et al. 

(2003: 76). 

Notes: Further synonyms are given in SCHUBERT et al. (2003). 

dendriticum [Wallr.] var. ß orbiculatum Berk., Gard. Chron. 1848: 716 (1848)! (?). 

Lit.: COOKE (1871: 583). 

Notes: ‘Gard. Chron. 1848: 716 (1848)’ has been checked, but the name ‘var. orbiculatum’ is 

not present. 

dendriticum [Wallr.] var. heteromeles Harkn. (1881), in herb. 

On Heteromeles arbutifolia (Rosaceae), USA, California, Jun. 1881 (BPI 426448). 


Lit.: RITSCHEL (2001), SCHUBERT et al. (2003: 76). 

dendryphioides Ellis, in herb. 

On Phytolacca sp. (Phytolaccaceae), USA, New Jersey. 

Notes: Specimen in Myc. Coll. (CASH 1952). 

densum Sacc., Bull. Orto Bot. Regia Univ. Napoli 6: 71 (1921)!. 

T: on dead stems of Ricinus communis (Euphorbiaceae), Italy, Salerno, Scafati, A. Trotter. 

Lit.: SACCARDO (1931: 791). 

depressum Berk. & Broome, Ann. Mag. Nat. Hist., Ser. 2, 7: 99 (1851)!. 

T: on Angelica sylvestris (Apiaceae), Great Britain (K). 

≡ Passalora depressa (Berk. & Broome) Sacc., Nuovo Giorn. Bot. Ital. 8: 187 (1876)!. 

≡ Fusicladium depressum (Berk. & Broome) Roum., F. gall. exs., No. 86 (1879)!. 

30

≡ Scolecotrichum depressum (Berk. & Broome) J. Schröt., in Cohn, Krypt.-Fl. Schlesien, Bd. 

3(2), Heft 4: 497 (1897)!. 

≡ Cercospora depressa (Berk. & Broome) Vassiljevsky, in Vassiljevsky & Karakulin, Parazitnye 

nesovershennye griby, Ch. I, Gifomicety: 385 (1937)!. 

≡ Megacladosporium depressum (Berk. & Broome) Vienn.-Bourg., Les champignons parasites 

des plantes cultivées 2: 1488 (1949), comb. inval. 

≡ Cercosporidium depressum (Berk. & Broome) Deighton, Mycol. Pap. 112: 37 (1967)!. 

≡ Passalora depressa (Berk. & Broome) Poonam Srivast., J. Liv. World 1(2): 114 (1994)!, 

comb. inval. 

= Passalora polythrincioides Fuckel, Jahrb. Nassauischen Vereins Naturk. 23–24: 353 ‘1869’ 

(1870). [T: Fuckel, F. rhen. 103, e.g., HAL]. 

= Fusicladium peucedani Syd. & P. Syd., Ann. Mycol. 5: 340 (1907), nom. illeg., homonym, 

non F. peucedani Ellis & Holw., 1895. [T: B; S]. 

= ? Cercospora depressa f. angelicae Dzhanuz., Trudy Vsesoyuzn. Inst. Zashch. Rast. 19: 9 

‘1963’ (1964). 

Teleomorph: Mycosphaerella angelicae Woron., Věstn. Tiflissk. Bot. Sada 28: 17 (1913). 

Lit.: COOKE (1871: 584), LINDAU (1907: 786), OUDEMANS (1923), CROUS & BRAUN (2003: 

157). 

desmotrichum Desm., Ann. Sci. Nat. Bot., Sér. 3, 16: 297 (1851)!, as ‘desmitrichum’. 

T: on dry leaves of Fraxinus ornus (Oleaceae), France (PC). 

Lit.: SACCARDO (1886: 360), FERRARIS (1912: 346), OUDEMANS (1923). 

devriesii A.A. Padhye & Ajello, in Gonzalez, Alfonso, Seckinger, Padhye & Ajello, 

Sabouraudia 22(5): 430 (1984)!. 

T: from breast of Homo sapiens, Cayman Islands. [ATCC 56280 ex-type (= CBS 147.84)]. 

≡ Cladophialophora devriesii (A.A. Padhye & Ajello) de Hoog, Kwon-Chung & McGinnis, in 

de Hoog, Guého, Masclaux, Gerrits van den Ende, Kwon-Chung & McGinnis, J. Med. 

Veterin. Mycol. 33: 344 (1995)!. 

Lit.: KWON-CHUNG & BENNETT (1992: 645), HO et al. (1999: 147), DE HOOG et al. (2000: 

573), SCHELL (2003: 578). 

dianellicola Y. Cui & Z.Y. Zhang, in He & Zhang, Mycosystema 20(4): 470 (2001)!. 

T: on living leaves of Dianella ensifolia (Phormiaceae), China, Zhejiang Prov., Hangzhou, 2 

Nov. 1980, J.Y. Li & T.Y. Zhang (MHYAU 03922: holotype). 

≡ Cladosporium dianellicola Z.Y. Zhang & Y. Cui, in Zhang et al., Flora Fungorum Sinicorum, 

Vol. 14: 88 (2003)!, nom. superfl. 

Ill.: HE & ZHANG (2001: 470, Fig. 2), ZHANG et al. (2003: 88, Fig. 52). 

diaphanum Thüm., Mycoth. univ., Cent. XIX, No. 1868 (1881)!. 

T: on dead leaves of Photinia serrulata (Rosaceae), France, Lyon, Jun. 1880, J. Therry, 

Thüm., Mycoth. univ. 1868 (HAL: lectotype, selected by BRAUN 2001; Thüm., Mycoth. univ. 

1868: isolectotypes). Topotypes: Roum., F. sel. gal. exs. 5591. 

Lit.: SACCARDO (1892: 603), ELLIS (1976: 342), BRAUN (2001: 56). 

Ill.: ELLIS (1976: 341, Fig. 259 B), BRAUN (2001: 55, Fig. 2). 

Host(s)/substrate(s) & distribution: on Photinia glabra; Europe (France). 

dieffenbachiae Verpl. & Van den Broecke, Ann. Soc. Sci. Bruxelles, Sér. B, 56: 105 (1936)!. 

T: on dead leaves of Dieffenbachia magnifica (Araceae), Belgium, Gent, botanical garden, 

Sept. 1935, associated with Septoria dieffenbachiae Verpl. & Van den Broecke (GENT: 

holotype). 

digitalicola Z.Y. Zhang, T. Zhang & W.Q. Pu, in Zhang, Wei & Zhang, Mycosystema 17(3): 

195 (1998)!. 

T: on living flowers of Digitalis purpurea (Scrophulariaceae), China, Prov. Yunnan, 

Kunming, 25 Jun. 1990, Li Hua (MHYAU 03934). 

Ill.: ZHANG, WEI & ZHANG (1998: 196, Fig. 1). 

31

dracaenatum Thüm., Mycoth. univ., Cent. XIX, No. 1869 (1881)!. 

T: on living leaves of Dracaena cooperi (Dracaenaceae), USA, South Carolina, Aiken, 1876, 

H.W. Ravenel, Thüm., Mycoth. univ. 1869 (B 70-6248; BPI 426454; BR-MYC 8172,86; 

HAL; M-57504: syntypes). 

Lit.: SACCARDO (1892: 605). 

Host(s)/substrate(s) & distribution: on Cordyline and Dracaena spp.; North America (USA). 

Notes: FARR et al. (n.d.) reduced C. dracaenatum to synonymy with C. elatum (Harz) Nannf. 

However, the latter species is quite distinct and do not even belong in Cladosporium s. str. It is 

morphologically indistinguishable from the human-pathogenic species of Cladophialophora, 

but differs in being sensitive to cycloheximide (DE HOOG et al. 2000). The taxonomic position 

of C. elatum remains to be determined (HO et al. 1999). 

dufourii Brond., Arch. Fl. 1: 60 (1854)!. 

T: on decaying fruits of cucurbits (Cucurbitaceae), France, autumn and winter (TL). 

Lit.: SACCARDO (1886: 370). 

Notes: Types of Louis de Brondeau are at TL [according to STAFLEU & MENNEGA (1995)]. 

echinulatum (Berk.) Cooke – ATCC 56129 (JONG et al. 1996). 

Note: There is no such validly published name as C. echinulatum (Berk.) Cooke. Was actually 

deposited (Form 1F in ATCC records) as C. echinulatum (Berkeley) de Vries. 

echinulatum (Berk.) G.A. de Vries, Contr. Knowl. Genus Cladosporium: 49 (1952)!. 

T: on Dianthus caryophyllus (Caryophyllaceae), Great Britain, Feb. 1870, ex herb. M.J. 

Berkeley (K: lectotype, selected by DAVID, 1997). 

≡ Helminthosporium echinulatum Berk., Gard. Chron. 1870: 382 (1870). 

≡ Heterosporium echinulatum (Berk.) Berk. & Broome, in Cooke, Grevillea 5(35): 123 

(1877)!. 

= Heterosporium circinale Klotzsch, Herb. viv. myc., No. 188 (1832)!, nom. inval. 

= Helminthosporium exasperatum Berk. & Broome, Ann. Mag. Nat. Hist., Ser. 4, 11: 345 

(1873). 

≡ Heterosporium exasperatum (Berk. & Broome) Cooke, Grevillea 16(80): 109 (1888)!. 

= Heterosporium dianthi Sacc. & Roum., Rev. Mycol. (Toulouse) 3(11): 57 (1881). 

= Heterosporium echinulatum var. dianthi Losa, Collect. Bot. (Barcelona) 3: 149 (1952). 

= ? Heterosporium dianthi Sawada, Bull. Gov. Forest Exp. Sta. 105: 98 (1958), nom. illeg., 

homonym, non H. dianthi Sacc. & Roum., 1881. 

Teleomorph: Davidiella dianthi (C.C. Burt) Crous & U. Braun, in Braun, Crous, Dugan, 

Groenewald & de Hoog, Mycol. Progr. 2(1): 10 (2003)!. 

Lit.: ELLIS (1971: 311), SUBRAMANIAN (1971: 291–293), SIVANESAN (1984: 222), DAVID 

(1988b, 1997: 34–40), HO et al. (1999: 127). 

Ill.: ELLIS (1971: 311, Fig. 215 A), SUBRAMANIAN (1971: 292, Fig. 226), SIVANESAN (1984: 

223, Fig. 119), DAVID (1988b: Fig.; 1997: 37–38, Figs 7, 8 A–C), HO et al. (1999: 126, Figs 

15–16). 

Host(s)/substrate(s) & distribution: on leaves and sometimes also inflorescences of various 

species of Dianthus, but also other members of the Caryophyllaceae, including e.g. Cerastium, 

Lychnis, Saponaria, Silene and Viscaria; widespread. 

edgeworthiae H. Zhang & Z.Y. Zhang, Mycosystema 17(4): 305 (1998)!. 

T: on living leaves of Edgeworthia chrysantha (Thymelaeaceae), China, Jiangxi, Lushan, 15 

Oct. 1980, J.Y. Li & T.Y. Zhang (MHYAU 03957: holotype). 

Ill.: ZHANG & ZHANG (1998a: 305, Fig. 2), ZHANG et al. (2003: 92, Fig. 55). 

effusum Berk. & M.A. Curtis, in Berkeley, Grevillea 3(27): 106 (1875)!. 

T: on Polygonum punctatum (Polygonaceae), USA, South Carolina, Society Hill, No. 3775 (K: 

lectotype; IMI 104922: isolectotype). 

≡ Cercospora effusa (Berk. & M.A. Curtis) Ellis, J. Mycol. 1: 53 (1885). 

≡ Didymaria effusa (Berk. & M.A. Curtis) Solheim, Illinois Biol. Monogr. 12: 65 (1930). 

≡ Passalora effusa (Berk. & M.A. Curtis) U. Braun, Mycotaxon 55: 231 (1995)!. 

32

= Cercospora polygonorum Cooke, Hedwigia 17: 39 (1878). [T: K]. 

≡ Pseudocercospora polygonorum (Cooke) Y.L. Guo & X.J. Liu, Mycosystema 4: 110 (1991). 

= Helminthosporium hydropiperis Thüm., Rev. Mycol. (Toulouse) 1: 60 (1879). [T: Thüm., 

Mycoth. univ. 1087, e.g., HAL]. 

≡ Cercospora hydropiperis (Thüm.) Speg., Bol. Acad. Nac. Ci. 9: 191 (1880). 

Lit.: SACCARDO (1886: 362, 447), CHUPP (1954: 451, as Cercospora polygonorum), 

DEIGHTON (1986: 637), CROUS & BRAUN (2003: 170). 

Notes: In the original diagnosis three collections on different hosts were mentioned, 

DEIGHTON (1986) designated the collection on Polygonum as lectotype. The specimens on 

Lobelia species refer to a similar, but distinct Passalora. 

effusum G. Winter (GOTTWALD 1982). 

Notes: This entry belongs to material examined from the National Fungus Collection. See C. 

caryigenum and the species below. 

effusum (G. Winter) Demaree, J. Agric. Res. 37: 186 (1928)!, nom. illeg., homonym, non C. 

effusum Berk. & M.A. Curtis, 1875. 

T: on Carya tomentosa (= Carya alba) (Juglandaceae), USA, Illinois, Cobden Zels., 1 Oct. 

1882, F.S. Earle (B: holotype). 

≡ Fusicladium effusum G. Winter, J. Mycol. 1: 101 (1885). 

≡ Fusicladosporium effusum (G. Winter) Partridge & Morgan-Jones, Mycotaxon 85: 364 

(2003)!. 

= Fusicladium caryigenum Ellis & Langl., J. Mycol. 4: 124 (1888)!. [T: NY; BPI; M]. 

≡ Cladosporium caryigenum (Ellis & Langl.) Gottwald, Mycologia 74(3): 388 (1982)!. 

Lit.: SCHUBERT & BRAUN (2002a), SCHUBERT et al. (2003: 41–43). 

elaeagnus Gapon., Uzbekistan Biol. Zurn. (1964). 

T: on Elaeagnus (Elaeagnaceae), Uzbekistan, Bucharskoj Oblast. 

Lit.: SAGDULLAEVA et al. (1990: 47). 

Ill.: SAGDULLAEVA et al. (1990: 48). 

elatum (Harz) Nannf., in Melin & Nannfeldt, Svenska Skogsvardsfoereren Tidskr. 32(3–4): 

397 (1934). 

T: on an old stump, Germany. 

≡ Hormodendrum elatum Harz, Bull. Soc. Imp. Naturalistes Moscou 44(1): 140 (1871). 

≡ Cadophora elatum (Harz) Nannf., in Melin & Nannfeldt, Svenska Skogsvardsfoereren 

Tidskr. 32(3–4): 422 (1934). 

Lit.: DE VRIES (1952: 67), ELLIS (1976: 326), MATSUSHIMA (1983: 4), HO et al. (1999: 127), 

DE HOOG et al. (2000: 585). 

Ill.: DE VRIES (1952: 67, Fig. 14), MINOURA (1966: 141, Fig. 4A), ELLIS (1976: 327, Fig. 245 

B), MATSUSHIMA (1983: 26–27, Figs 143–144), HO et al. (1999: 126, Fig. 17), DE HOOG et al. 

(2000: 585–586, Figs). 

Notes: C. elatum is quite distinct from Cladosporium s. str. It is morphologically indistinguishable 

from the human-pathogenic species of Cladophialophora, but differs in being 

sensitive to cycloheximide (DE HOOG et al. 2000). The taxonomic position of C. elatum 

remains to be determined (HO et al. 1999). 

elegans Matsush., Icones Microfungorum a Matsushima Lectorum: 35 (1975)!, nom. illeg., 

homonym, non C. elegans Penzig, 1882. 

T: isolated from garden soil, Japan, Yaku Island, Kagoshima, Jul. 1971 (Matsush. herb. 4109). 

Ill.: MATSUSHIMA (1975: Pl. 43). 

Notes: Excluded, but generic affinity not yet clear. ZHANG et al. (2003) reduced C. elegans 

Matsush. to synonymy with C. elegans Penz., but the latter species proved to be a synonym of 

C. herbarum and is therefore quite distinct. 

elegans Penz., in Saccardo, Michelia 2(8): 471 (1882)!. 

T: on living leaves of Citrus sp. (Rutaceae), Italy, Padova, Mar. 1882, O. Penzig (HBG: 

neotype; PAD: isoneotype). 

33



Lit.: SACCARDO (1886: 358), LINDAU (1907: 825), FERRARIS (1912: 342), GONZÁLES- 

FRAGOSO (1927: 201). 

Notes: “Subaffine Cladosporio noduloso Corda” (see SACCARDO 1886). OUDEMANS (1920– 

1923) listed Urtica dioica, Hedera helix and Lonicera tatarica as hosts of this species, 

although it was described as parasitic on Citrus. 

elegans [Penz.] var. singaporense Sacc., Bull. Orto. Bot. Regia Univ. Napoli 6: 60 (1921)!. 

T: on leaves of Citrus acida (Rutaceae), Singapore, Botanical Garden, Aug. 1917, Baker, No. 

4985 (PAD). 

= Helminthosporium guareicola F. Stevens, Bot. Gaz. 65: 241 (1918). 

≡ Spiropes guareicola (F. Stevens) Cif., Sydowia 9: 303 (1955). 

≡ Pleurophragmium guareicola (F. Stevens) S. Hughes, Canad. J. Bot. 36: 797 (1958). 

Lit.: SACCARDO (1931: 795), SCHUBERT & BRAUN (2005a: 107). 

elegantulum Pidopl. & Deniak, Mikrobiol. Zhurn. 5(2): 182, 193 (1938)!, nom. inval. 

T: from a rotting fruit of Malus sp. (Rosaceae) and from meat, Ukraine. 



elsinoes H.C. Greene, Trans. Wisconsin Acad. Sci. 47: 127 (1958)!. 

T: on fructifications of Elsinoe wisconsinensis (Elsinoaceae) on Desmodium illinoense 

(Fabaceae), USA, Wisconsin, Lafayette Co., Ipswich near Platteville, 16 Aug. 1951, H.C. 

Greene (BPI 426465A, BPI 426465B; WIS: isotypes). 

= Dendryphiella infuscans (Thüm.) M.B. Ellis, Dematiaceous Hyphomycetes: 500 (1971). 

Lit.: HEUCHERT et al. (2005: 63–64). 

Ill.: HEUCHERT et al. (2005: 64, Fig. 25). 

entoxylinum Corda, Icon. fung. 1: 14 (1837)!. 

T: on wood of Pinus sp. (Pinaceae), Czech Republic, Reichenberg, Corda (PRM 155726). 

Lit.: SACCARDO (1886: 353), LINDAU (1907: 811, 1910: 796), OUDEMANS (1919, 1923). 


Notes: Excluded, no Cladosporium s. str. This species has to be excluded from Cladosporium 

s. str. The conidiogenous cells are unilocal, determinate or occasionally percurrent, the 

conidiogenous loci and conidial hila are unthickened and non-coronate. This species is rather 

taeniolella/xylohypha-like. Several collection under ‘Cladosporium entoxylinum’ proved to be 

Cladosporium herbarum s. lat. (e.g., on wood of Abies excelsa, Bohemia, Krypt. exs. 2837, 

PRM 481966; on wood of Carpinus betulus, Austria, Wiener Wald, May 1936, K. Keissler, 

PRM 657474). The fungus described and illustrated from China by ZHANG et al. (2003) as 

‘Cladosporium entoxylinum’ is quite distinct and the name is misapplied. OUDEMANS (1923) 

listed C. entoxylinum as pathogen of Sambucus racemosa (Caprifoliaceae). 

epacridis McAlpine, Victoria Naturalist 17(10): 186 (1901)!. 

T: on living leaves of Epacris impressa (Epacridaceae), Australia, Victoria, Caulfield, Aug. 

1900, C. French (VPRI). 

Lit.: SACCARDO (1902: 1058). 

Notes: Excluded, but taxonomic status not yet clear. 

epibryum Cooke & Massee, in Cooke, Grevillea 17(84): 76 (1889)!. 

T: on capsules of various mosses, USA, E.G. Britton (K). 

Lit.: SACCARDO (1892: 605). 

epichloës Lobik, Bolezni Rast. 17(3–4): 189 (1928)!. 

T: on stroma of Epichloë typhina (Clavicipitaceae) on Bromus inermis (Poaceae), Russia, 

‘Leyss., Bezirk v. Piatigorsk, im Walde am Abhange d. Berges Maschuk, bei d. Lermontowschen 

Haltestelle’, 26 Sept. 1923 (LE 40522: holotype). 


Ill.: LOBIK (1928: Tab. 8, Fig. 86), HEUCHERT et al. (2005: 18, Fig. 2). 

Host(s)/substrate(s) & distribution: on Epichloë typhina; Europe (Austria, Germany, Russia). 

34

epimyces Cooke, Grevillea 12(61): 31 (1883)!. 

T: on pileus of Agaricus melleus (Agaricaceae), USA, South Carolina, Aiken, Rav., F. amer. 

exs. 596 (NY: lectotype, selected by HEUCHERT et al., 2005). Isolectotypes: Rav., F. amer. 

exs. 596 (e.g., BPI 426471; PH 01020432). 


Ill.: HEUCHERT et al. (2005: 20–21, Fig. 3−4). 

Host(s)/substrate(s) & distribution: on gills of several Agaricales and Russulales, including 

Armillaria mellea, Hypholoma sublatentum, Lactarius sp. and Leucopaxillus gentianeus; 

Europe (Poland), North America (USA). 

epiphyllum Peck – DOV [A. Commons 1379 (SUMSTINE 1949: 18)]. 

epiphyllum (Pers.: Fr.) Nees, Syst. Pilze 1: 67 (1817)! and in Mart., Fl. crypt. erlang.: 351 

(1817)!: Fr., Syst. mycol. 3(2): 370 (1832)!. 

T: on dry leaves (L 910.225-646). 

≡ Dematium epiphyllum Pers., Syn. meth. fung. 2: 695 (1801)!. 

≡ Cladosporium epiphyllum (Pers.) Link, in Willd., Sp. pl. 6(1): 42 (1824)!. 

≡ Chloridium epiphyllum (Pers.) Chevall., Fl. gén. env. Paris 1: 35 (1826)!. 

≡ Cladosporium epiphyllum (Pers.) Fries, Syst. mycol. 3(2): 370 (1832)!. 



Lit.: CORDA (1837: 14), COOKE (1871: 583), SACCARDO (1886: 360), LINDAU (1907: 804; 

1910: 795), FERRARIS (1912: 334; 1914: 882), GONZÁLES-FRAGOSO (1927: 199), LIND (1913: 

523), DE VRIES (1952: 71), HUGHES (1958: 750). 

epiphyllum [(Pers.: Fr.) Nees] β chionanthi (Pers.) Link, in Willd., Sp. pl. 6(1): 42 (1824)!. 

T: on leaves of Chionanthus (Oleaceae). 

≡ Dematium epiphyllum [Pers.] β chionanthi Pers., Mycol. eur. 1: 16 (1822)!. 

epiphyllum [(Pers.: Fr.) Nees] var. acerinum Sacc., Nuovo Giorn. Bot. Ital., N.S., 27: 86 

(1920)!. 

T: on leaves, partly becoming dry, of Acer platanoides (Aceraceae), USA, Idaho, Coeur d’ 

Alene, Sept. 1918, N.W. Scherer, No. 10027 (PAD). 


Castañeda, Dugan & Jong, Mycotaxon 72: 131 (1999)!. 

Lit.: SACCARDO (1931: 787). 

episclerotiale Bubák, Ann. Mycol. 12: 213 (1914)!. 

T: on sclerotia of Sclerotinia cinerea (Sclerotiniaceae) on fruits of Prunus domestica (Rosaceae), 

Italy, Tyrol, Arco, Mar. 1913, E. Diettrich-Kalkhoff (BPI 426531: holotype). 

Lit.: SACCARDO (1931: 797), HEUCHERT et al. (2005: 22–24). 

Ill.: HEUCHERT et al. (2005: 23, Fig. 5; Pl. 2, Fig. 9). 

Host(s)/substrate(s) & distribution: on Boletales, including Monilinia laxa and Suillus bovinus; 

Europe (Germany, Italy). 

episphaerium Schwein., Trans. Amer. Philos. Soc., N.S., 4(2): 278 (1832)!, as ‘episphaeria’. 

T: on Sphaeria species, USA, Pennsylvania, Bethlehem, No. 2607 (PH 01020420, 01020421: 

syntypes). 

= ? Dematium episphaerium Alb. & Schwein., Consp. fung. lusat.: 369 (1805)!. 

Lit.: SACCARDO (1886: 369), OUDEMANS (1919, 1920: on Hypoxylon rubiginosum; 1924). 

epixilinum Corda – GOLA (1930: 20). 

Notes: see C. herbarum Link var. epixylinum Corda. 

equiseti Pass., in herb. 

On Equisetum ramosum (Equisetaceae), Italy, Emilia Romagna, Parma, 1873, G. Passerini (B 

70-6324). 



35

erianthi Thüm., Rev. Mycol. (Toulouse) 1: 59 (1879)!. 

T: on dry culms and leaves of Erianthus saccharoides (Poaceae), USA, South Carolina, Aiken, 

H.W. Ravenel, Thüm., Mycoth. univ. 1766 (e.g., BPI 426532; BR-MYC 81373,87; HAL; M; 

NY). 

Lit.: SACCARDO (1886: 364). 

eriobotryae Pass. & Beltrani, Transunti, Ser. 3, 7(1): 38 (1882)!. 

T: on dry leaves of Eriobotrya japonica (Rosaceae), Italy, Sicily, Monostalla, Sept. 1878. 

Lit.: SACCARDO (1886: 356), LINDAU (1907: 824), FERRARIS (1912: 344). 

eriolobi Thaung, Trans. Brit. Mycol. Soc. 63(3): 620 (1974)!. 

T: on living leaves of Eriolobus indica (Rosaceae), Myanmar (Burma), Maymyo, Botanical 

Garden, 28 Dec. 1972, Mya Thaung (IMI 175732: holotype). 

≡ Stenella eriolobi (Thaung) K. Schub. & U. Braun, comb. nov. 

Ill.: THAUNG (1974: 621, Fig. 2). 

eschscholtziae (Harkn.) Dingley, nom. ined. 

T: on leaves of Eschscholtzia californica (Papaveraceae), USA, California, San Francisco, Jan. 

[1884], Harkness, No. 3116 (holotype destroyed). 

≡ Heterosporium eschscholtziae Harkn., Bull. Calif. Acad. Sci. 1: 38 (1884). 

≡ Acroconidiella eschscholtziae (Harkn.) M.B. Ellis, More Dematiaceous Hyphomycetes: 407 

(1976)!. 

Lit.: DAVID (1997: 111). 

eucalypti Tassi, Bull. Lab. Orto Bot. Reale Univ. Siena 3(1): 20 (1900). 

T: on frozen leaves of Eucalyptus globulus (Myrtaceae), Italy, Siena, botanical garden. 


FRAGOSO (1927: 202). 

Ill.: GONZÁLES-FRAGOSO (1927: 202, Fig. 45). 

Notes: Type material is not deposited at herb. SIENA. 

eucalypticola M.B. Ellis, on CABI page, in herb.?, Kirk et al. (n. d.). 

euphorbiae Politis, Pragmat. Akad. Athen 4: 39 (1935)!. 

T: on Euphorbia sibthorpii (Euphorbiaceae), Greece, Attica, near Athina. 

Lit.: SACCARDO (1972: 1337). 

exoasci Ellis & Barthol., in Shear, F. columb., Cent. XV, No. 1493 (1901)!, nom. nud. 

T: on Exoascus communis on Prunus americana (Rosaceae), USA, Colorado, Walsenberg, Jul. 

1900, C.L. Shear, Shear, F. columb. 1493 (e.g., NY). 

= Cladosporium phyllophilum McAlpine, Agric. Gaz. New South Wales 7: 153 (1896)!. 

exoasci Lindau, in Rabenhorst, Krypt.-Fl., ed. 2, 1(8): 808 (1907)!. 

T: on Taphrina pruni (= Exoascus rostrupianus) (Taphrinaceae) on Prunus spinosa (Rosaceae), 

Germany, Hessen-Nassau, Rhön, near Gersfeld, ca. 500 m, 31 Jul. 1906, O. Jaap, F. sel. 

exs. 248 (B: lectotype; Jaap, F. sel. exs. 248: isolectotypes). 

= Cladosporium exoasci Ellis & Barthol., in Shear, F. columb., Cent. XV, No. 1493 (1901)!, 

nom. nud. 

= Cladosporium phyllophilum McAlpine, Agric. Gaz. New South Wales 7: 153 (1896)!. 

Lit.: LINDAU (1910: 796), SACCARDO (1913: 1370), BRAUN (2001: 54–56), HEUCHERT et al. 

(2005: 36–40). 

Ill.: BRAUN (2001: 55, Fig. 1), HEUCHERT et al. (2005: 38, Fig. 14). 

exobasidii Jaap, Verh. Bot. Vereins Prov. Brandenburg 49: 29 (1907)!. 

T: on Exobasidium vaccinii (Exobasidiaceae) galls on Vaccinium uliginosum (Ericaceae), 

Germany, Rhön, Gersfeld, Rotes Moor, 30 Jul. 1906, O. Jaap (B 70-6339: lectotype, selected 

by HEUCHERT et al., 2005). Isolectotypes: Jaap, F. sel. exs. 200 (e.g., HAL, M-57603). 

Paratype: on galls of Exobasidium vaccinii var. vaccinii on Vaccinium uliginosum, Germany, 

Brandenburg, Kr. Prignitz, Putlitzer Heide bei Putlitz, 1 Sept. 1900, O. Jaap (B 70-6340). 

36

Lit.: LINDAU (1907: 808, 1910: 796), LIND (1913: 522), SACCARDO (1913: 1371), BRAUN 

(2001: 57), HEUCHERT et al. (2005: 24–26). 

Ill.: BRAUN (2001: 55, Fig. 3), HEUCHERT et al. (2005: 25, Fig. 6). 

Host(s)/substrate(s) & distribution: on Exobasidium vaccinii; Europe (Denmark, Germany, 

Sweden, Ukraine). 

Notes: DE VRIES (1952) and HO et al. (1999) cite C. exobasidii as a synonym of Cladosporium 

cladosporioides, but the former species is well-distinguished by having frequently nodulose, 

hardly geniculate-sinuous conidiogenous cells with numerous aggregated scars (BRAUN 2001). 

In OUDEMANS (1919) this species is treated as a synonym of Cladosporium fuligineum 

Bonord. (= C. herbarum). 

exobasidii [Jaap] var. verruculosum Heuchert, U. Braun & K. Schub., Schlechtendalia 13: 27 

(2005)!. 

T: on Exobasidium vaccinii var. vaccinii, Czech Republic, near Pontresina, 3 Sept. 1899, P. 

Magnus (HBG: holotype). 


Host(s)/substrate(s) & distribution: on Exobasidium vaccinii and E. warmingii; Europe (Czech 

Republic, Switzerland). 

extoma Sacc. – GOLA (1930: 20) listed the type at PAD. 

Notes: Maybe this is an error in GOLA (1930) and C. extorre was intended. 

extorre Sacc., Nuovo Giorn. Bot. Ital., N.S., 27: 86 (1920)!. 

T: on the bark of young twigs of Whitney Crab Apple (= Pyrus coronaria) (Rosaceae), USA, 

Wyoming, Tonington, 12 Jun. 1917, V. Simmons, No. 10335 (PAD). 

Lit.: SACCARDO (1931: 795). 

Notes: Species is to be excluded. 

fagi Oudem., Ned. Kruidk. Arch., Ser. 3, 2(3): 768 (1902)! and Beih. Bot. Centralbl. 11: 538 

(1902)!. 

T: on fallen, dry leaves of Fagus silvatica (Fagaceae), Netherlands, Bussum, 11 Oct. 1901, 

C.J. Koning (L). 

= ? Cladosporium herbarum (Pers.: Fr.) Link, Ges. Naturf. Freunde Berlin Mag. Neuesten 


Lit.: SACCARDO (1906: 577), LINDAU (1907: 821), OUDEMANS (1920), DE VRIES (1952: 71). 

farnetianum Sacc., Syll. fung. 22: 1366 (1913)!. 

T: on fruits of Citrus limon (Rutaceae), Italy, Sicily. 

≡ Cladosporium citri Briosi & Farneti, Atti Ist. Bot. Univ. Pavia, Ser. 2, 10: 19 (1907)!, nom. 

illeg., homonym, non C. citri Massee, 1899. 

≡ Kurosawaia citri Hara, List of Japanese Fungi, ed. 4: 172 (1954), nom. nov., as ‘(Briosi & 

Farneti) Hara’. 

Lit.: FERRARIS (1914: 884). 

Notes: see comments on C. citri Massee above. 

fasciculare Fr., Syst. mycol. 3(2): 370 (1832)!. 

T: on herbal stems. 

≡ Dematium articulatum Pers., Neues Mag. Bot. 1: 121 (1794)!. 

= Helminthosporium vesicarium Wallr., Fl. crypt. Germ. 2: 166 (1833)!. 

Lit.: COOKE (1871: 583), SACCARDO (1886: 367), LINDAU (1907: 817), FERRARIS (1912: 

339), OUDEMANS (1919–1924), GONZÁLES-FRAGOSO (1927: 207). 

Ill.: PERSOON (1794: Tab. 4, Fig. 2). 

fasciculare f. chamaeropis Unamuno, Trab. Secc. Cienc. Nat. Congr. Assoc. Progr. Cienc. 

Oporto 1921: 60 (1922). 

T: on leaves of Chamaerops humilis (Arecaceae), Spain, near Oriedo, May 1921, P. Unamuno 

(MA 06416: holotype). 

≡ Cladosporium chamaeropis (Unamuno) K. Schub., comb. nov. 

37

fasciculatum Corda, Icon. fung. 1: 15 (1837)!. 

T: on Scirpus (Cyperaceae), Czech Republic, ‘prope Okoř Bohemiae’ (PRM). 



Lit.: SACCARDO & BERLESE (1884: 100), SACCARDO (1886: 366), LINDAU (1907: 816), 

FERRARIS (1912: 338; 1914: 883), GONZÁLES-FRAGOSO (1927: 210). 


fasciculatum [Corda] f. amerotrichum Traverso, Malpighia 19: 149 (1905)!. 

T: on faded leaves of Gladiolus sp. (Iridaceae), Italy, Prov. Di Como, Villa Stroppa in Tradate, 

Sept. 1902. 

Lit.: SACCARDO (1906: 577), LINDAU (1907: 817), OUDEMANS (1919). 

Notes: TRAVERSO (1905) described it as differing from typical C. fasciculatum in having nonseptate 

conidiophores. 

fasciculatum var. densum, in herb. 

On Euonymus japonicus (Celastraceae), USA, South Carolina (BPI 426554). 

fermentans Goto, Yamak. & Yokots., J. Agric. Chem. Soc. Japan 49(7): 380 (1975)!. 

T: from olive fruit waste, Japan, Kofu, 12 Jun. 1968, S. Goto (RIFY 0587). 

= Pichia burtonii Boidin, Pignal, Lehodey, Vey & Abadie, Bull. Soc. Mycol. Fr. 80: 437 

‘1964’ (1965). 

≡ Endomycopsis burtonii (Boidin, Pignal, Lehodey, Vey & Abadie) Kreger, The Yeasts, Ed. 2: 

174 (1970), nom. inval. 

≡ Hyphopichia burtonii (Boidin, Pignal, Lehodey, Vey & Abadie) Arx & Van der Walt, 

Antonie van Leeuwenhoek J. Microbiol. Serol. 42(3): 310 (1976)!. 

ferox (Kabát & Bubák ex Lindau) J.C. David, Mycol. Pap. 172: 40 (1997)!. 

T: on leaves of Ranunculus arvensis (Ranunculaceae), Czech Republic, Tábor, in the botanical 

garden, 12 Jul. 1906, F. Bubák, Kab. & Bub., F. imp. exs. 444 (BPI: lectotype). 

≡ Heterosporium ferox Kabát & Bubák ex Lindau, in Rabenhorst, Krypt.-Fl., ed. 2, 1(9): 83 

(1910)!. 

≡ Heterosporium ferox Kabát & Bubák, F. imp. exs., Fasc. 9, No. 444 (1907)!, nom. inval. 

Ill.: DAVID (1997: 41–42, Figs 9–10). 

Host(s)/substrate(s) & distribution: on Ranunculus arvensis; Europe (Czech Republic). 

ferrugineum Allesch., in Hennings, Hedwigia 34: 116 (1895)!. 

T: on leaves of Sweetia bijuga (Fabaceae), Brazil, Minas Geraës, Paranahyba, Jul. 1892, E. 

Ule, No. 1905 (M-57571: lectotype; HBG, PC: isolectotypes). 

≡ Passalora sweetiae K. Schub. & U. Braun, Mycol. Progr. 4(2): 105 (2005)!. 

Lit.: SACCARDO (1895: 619). 


ferrugineum R.F. Castañeda, Fungi Cubensis II: 4 (1987)!, nom. illeg., homonym, non C. 

ferrugineum Allescher, 1895. 

T: on living leaves of Nectandra coriacea (Lauraceae), Cuba, prov. Matanzas, Calimete, 24 

Jan. 1987, R.F. Castañeda [INIFAT C87/45: holotype; CBS 784.87 (ex-type)]. 



ferrugineum Sacc. – GOLA (1930: 20) listed type at PAD. 

Notes: Maybe an error in GOLA (1930) and C. fumagineum Sacc. was intended. 

festucae Sawada, Bull. Gov. Forest Exp. Sta. 105: 95 (1958)!. 

T: on leaves of Festuca japonica (Poaceae), Japan, Tohoku District, 29 Sept. 1947. 

Notes: Type material could not be traced, neither in herb. PPMH not BPI, and is probably not 

preserved. 

fici F. Patt., Bull. Torrey Bot. Club 27: 285 (1900)!. 

T: on living leaves of Ficus parcelli (Moraceae), USA, Department of Agriculture, green- 

38

house, Jan. 1900, F.W. Patterson (herb. of the ‘Division of Vegetable Physiology and 

Pathology, U.S. Department of Agriculture’). 

Lit.: SACCARDO (1902: 1059). 

Notes: Type material of this species could not be traced and was not available for a reexamination. 

flueggeae Thüm., ad. int., in Rabenhorst, F. eur., Cent. XVI, No. 1571 (1872)!, nom. nud. 

T: on leaves of Flueggea japonica (Euphorbiaceae), Greece, Athina, 20 Sept. 1869, de 

Heldreich, Rabenh., F. eur. 1571 (e.g., HAL; HBG: syntypes). 

Notes: Excluded, mixed infection with Alternaria sp., Colletotrichum sp. and an unknown 

phialidic hyphomycete. 

foliorum Ellis & Everh. ex K. Schub., sp. nov. 

T: on living leaves of Angelica breweri (Apiaceae), USA, California, Amador Co., Pine 

Grove, Aug. 1896, G. Hansen (BPI 426580: holotype). Isotype: BPI 426581. 

≡ Cladosporium foliorum Ellis & Everh., in herb. 

Host(s)/substrate(s) & distribution: on Angelica breweri; North America (USA). 

forsythiae Z.Y. Zhang & T. Zhang, in Zhang, Zhang, Liu, & He, J. Anhui Agric. Univ. 26: 36 

(1999)!. 

T: on living leaves of Forsythia suspensa (Oleaceae), China, Henan, Zhengzhou, 20 Aug. 

1992, Qing Yun (MHYAU 07030: holotype). 

Ill.: ZHANG, ZHANG, LIU & HE (1999: 37, Fig. 1), ZHANG et al. (2003: 101, Fig. 63). 

Notes: Second collection is mentioned by ZHANG et al. (2003) on Forsythia sp. from China, 

Liaoning. 

foveolicola Speg., Anales Mus. Nac. Buenos Aires 20: 437 (1910)!. 

T: on living leaves of Eupatorium bupleurifolium (Asteraceae), Argentina, near San Javier, 

Misiones, Aug. 1909 (LPS 13.136: holotype). 

= Passalora foveolicola (Speg.) U. Braun & K. Schub., comb. nov. 


fraxinicola K. Schub. & W. Mułenko, sp. nov. 

T: on Fraxinus excelsior (Oleaceae), Germany, Sachsen-Anhalt, Halle (Saale), Neuwerk/ 

Jägerplatz, kindergarten, 23 Jun. 2004, K. Schubert (HAL 1829: holotype). Paratypes: on 

Fraxinus excelsior, Germany, Sachsen-Anhalt, Halle (Saale), Neuwerk/Jägerplatz, kindergarten, 

2 Aug. 2004, K. Schubert (HAL 1830 F) and U. Braun, Fungi selecti exsiccati 47; 

Poland, Lublin, street margin, 10 Aug. 2004, W. Mułenko (HAL 1831 F) and U. Braun, Fungi 

selecti exsiccati 48. 

Host(s)/substrate(s) & distribution: on Fraxinus excelsior; Europe (Germany, Poland). 

fuligineum Bonord., Abh. Mykol. 1: 92 (1864)!. 

T: on decaying gills of Boletus subtomentosus (Boletaceae), Germany, Westfalen, Siegen, 

Hainichen, 16 Jul. 1922, A. Ludwig (B: neotype, selected by BRAUN, 2001). 

= Cladosporium herbarum (Pers.: Fr.) Link (var. herbarum), Ges. Naturf. Freunde Berlin 

Mag. Neuesten Entdeck. Gesammten Naturk. 7: 37 (1816)!. 

Lit.: SACCARDO (1886: 368; 1913: 1371), LINDAU (1907: 808; 1910: 796), JAAP (1908: 217), 

LIND (1913: 523), GONZÁLES-FRAGOSO (1927: 212), BRAUN (2001: 56). 

fuligineum f. racemosa, in herb. 

Substrate Undetermined, Czech Republic, Bohemia, Kokocko, near Plzen, 10 Oct. 1910, F. 

Bubák (BPI 426582). 

fuligo Bonord., in herb. 

On Tilia europaea (Tiliaceae), Funk (B). 

fulvum (Arx) K. Bhalla & A.K. Sarbhoy → fulvum Cooke. 

fulvum Cooke, Grevillea 12(61): 32 (1883)!. 

T: on leaves of tomato (Lycopersicon esculentum) (Solanaceae), USA, South Carolina, Rav., 

39

F. amer. exs. 599 (K). 

≡ Fulvia fulva (Cooke) Cif., Atti Ist. Bot. Lab. Crittog. Univ. Pavia 10(1): 245 (1954)!. 

≡ Mycovellosiella fulva (Cooke) Arx, Proc. Kon. Ned. Akad. Wetensch. C, 86(1): 48 (1983)!. 

≡ ‘Cladosporium fulvum (Arx) K. Bhalla & A.K. Sarbhoy’, Indian Phytopathol. 53(3): 262 

(2000)!, nom. superfl. 

≡ Passalora fulva (Cooke) U. Braun & Crous, in Crous & Braun, Mycosphaerella and its anamorphs: 

1. Names published in Cercospora and Passalora, CBS Biodiversity Ser. 1: 453 

(2003)!. 

Lit.: SACCARDO (1886: 363), LINDAU (1907: 829; 1910: 797), FERRARIS (1912: 349; 1914: 

885), DE VRIES (1952: 70), ELLIS (1971: 306–307), HOLIDAY & MULDER (1976), VON ARX 

(1987: 195), HO et al. (1999: 128). 

fulvum [Cooke] var. violaceum Voglino, Ann. Reale Accad. Agric. Torino 55: 381, 1912 

(1913)!. 

T: on leaves of Solanum lycopersicum (= Lycopersicon esculentum) (Solanaceae), Italy, 

Liguria, Albenga. 

Lit.: SACCARDO (1931: 794). 

Notes: Very probably synonymous with Passalora fulva (Cooke) U. Braun & Crous. 

fumagineum Sacc., Nuovo Giorn. Bot. Ital., N.S., 27: 86 (1920)!. 

T: on living leaves of Quercus sp. (Fagaceae), USA, Wyoming, Cook Co., Devils Tower, 28 

Jul. 1918, J.R. Weir, No. 10025 (BPI 426763; PAD). 

Lit.: SACCARDO (1931: 791). 

Host(s)/substrate(s) & distribution: on Quercus sp.; North America (USA). 

Notes: SACCARDO (1931) cited “Spokane, Washington” as locality, not agreeing with the data 

given in two collections from BPI and PAD, which are marked as ‘type’. 

fumago Link, in Willd., Sp. pl. 6(1): 40–41 (1824)!: Fr., Syst. mycol. 3(2): 372 (1832)!. 

T: on leaves of various plant species, Europe (B ?). 

≡ Torula fumago (Link) Chevall., Fl. gén. env. Paris 1: 34 (1826)!. 

≡ Caldariomyces fumago (Link) Woron., Ann. Mycol. 25: 261 (1927). 

≡ Leptoxyphium fumago (Link) R.C. Srivast., Arch. Protistenk. 125(1–4): 333 (1982), as 

‘(Woron.) R.C. Srivast.’, nom. inval. 

= ? Fumago vagans Pers., Mycol. eur. 1: 9 (1822)!. 

≡ Cladosporium vagans (Pers.) Desz., Pl. crypt. N. France, Ed. 1, Fasc. I, No. 6 (1825), as 

‘Pers.’. 

= Syncollesia foliorum C. Agardh, Syst. alg.: 32 (1824). 

Lit.: FRIES (1832: 372), SACCARDO (1886: 547), LINDAU (1910: 267), LIND (1913: 166), 

OUDEMANS (1919–1924). 

Notes: LINK (in WILLDENOW, 1824) mentioned Fumago Pers. (“Fumago Pers. ex hoc 

Cladosporio saepe oritur”), but did not refer to Fumago vagans, so that Link´s species was not 

based on the latter name. 

fumago [Link] f. artemisiae abrotani Thüm., Herb. myc. oec., Fasc. XIV, No. 657 (1879)!, 

nom. nud. 

T: on Artemisia abrotanum (Asteraceae), Austria, near Kalksburg, Sept. 1879, von Thümen, 

Thüm., Herb. myc. oec. 657 (B). 

fumago [Link] f. carpini betuli Thüm., Herb. myc. oec., Fasc. VII, No. 339 (1875)!, nom. nud. 

T: on living leaves of Carpinus betulus (Corylaceae), Czech Republic, Bohemia, near Teplitz- 

Schönau, Aug. 1873, von Thümen, Thüm., Herb. myc. oec. 339 (B). 

fumago [Link] f. corticicola Rabenh., Herb. mycol., Ed. nova, Ser. prima, Cent. IV, No. 330 

(1856)! and Flora 15(9): 134 (1857)!, nom. nud. 

T: on Salix twigs, Germany, Giessen, Hoffmann, Rabenh., Herb. mycol. 330 (e.g., HAL). 

fumago [Link] f. coryli Thüm., Herb. myc. oec., Fasc. III, No. 131 (1873)!, nom. nud. 

T: on leaves and twigs of Corylus avellana (Coryllaceae), Czech Republic, Bohemia, near 

Teplitz-Schönau, Jul. 1873, von Thümen, Thüm., Herb. myc. oec. 131 (B). 

40

fumago [Link] f. fragariae-vescae Thüm., F. austr., Cent. XI, No. 1085 (1874)! and Herb. 

myc. oec., Fasc. IV, No. 169 (1874)!, nom. nud. 

T: on living leaves of Fragaria vesca (Rosaceae), Czech Republic, Teplitz-Schönau, summer 

1873, de Thümen, Thüm., F. austr. 1085, Herb. myc. oec. 169 (B; HAL). 

fumago [Link] f. fraxini Thüm., Herb. myc. oec., Fasc. VIII, No. 375 (1875)!, nom. nud. 

T: on living leaves of Fraxinus excelsior (Oleaceae), Czech Republic, Bohemia, near Teplitz- 

Schönau, Aug. 1873, von Thümen, Thüm., Herb. myc. oec. 375 (B 70-6422). 

Notes: Excluded, several hyphomycetes, incl. a trimmatostroma-like hyphomycete and C. 

herbarum. 

fumago [Link] f. grossulariae Thüm., Herb. myc. oec., Fasc. III, No. 130 (1873)!, nom. nud. 

T: on leaves and twigs of Ribes grossularia (Grossulariaceae), Czech Republic, Bohemia, near 

Teplitz-Schönau, Jul. 1873, de Thümen, Thüm., Herb. myc. oec. 130 (B). 

fumago [Link] f. humuli-lupuli Thüm., Herb. myc. oec., Fasc. XIII, No. 606 (1878)!, nom. 

nud. 

T: on living leaves of Humulus lupulus (Cannabaceae), Austria, near Klosterneuburg, Aug. 

1878, von Thümen, Thüm., Herb. myc. oec. 606 (B). 

fumago [Link] f. poae-pratensis Thüm., Herb. myc. oec., Fasc. IV, No. 160 (1874)!, nom. 

nud. 

T: on Poa pratensis (Poaceae), Czech Republic, Bohemia, near Teplitz-Schönau, summer 

1873, de Thümen, Thüm., Herb. myc. oec. 160 (B). 

fumago [Link] f. quercus, in herb. 

On leaves of Quercus pedunculata (Fagaceae), Germany, Bavaria, Bayreuth, Aug. 1877, A. 

Walther (BPI 426739). 

fumago [Link] f. rosae-acutifoliae, in herb. 

On Rosa sp. (Rosaceae), Germany, Bavaria, Bayreuth, Sept. 1877, A. Walther (BPI 426751). 

fumago [Link] f. rosae-albae Thüm., Herb. myc. oec., Fasc. IX, No. 418 (1876)!, nom. nud. 

T: on leaves and twigs of Rosa alba (Rosaceae), Germany, Bavaria, near Bayreuth, Sept. 

1874, von Thümen, Thüm., Herb. myc. oec. 418 (B). 

fumago [Link] f. syringae-vulgaris Thüm., Herb. myc. oec., Fasc. VIII, No. 393 (1875)!, nom. 

nud. 

T: on living leaves of Syringa vulgaris (Oleaceae), Czech Republic, Bohemia, near Teplitz- 

Schönau, Aug. 1873, von Thümen, Thüm., Herb. myc. oec. 393 (B; M-57668). 

Notes: Excluded, trimmatostroma-like hyphomycete. 

fumago [Link] f. ulmi Rabenh., Herb. mycol., Ed. nova, Ser. prima, No. 329 (1856)! and Flora 

15(9): 134 (1857)!, nom. nud. 

T: on Ulmus campestris (Ulmaceae), Germany, Dresden, 1856, Rabenh., Herb. mycol. 329 

(e.g., HAL). 

fumago [Link] f. ulmi-effusae Thüm., Herb. myc. oec., Fasc. V, No. 237 (1874)!, nom. nud. 

T: on leaves and young twigs of Ulmus sp. (Ulmaceae), Czech Republic, Bohemia, near 

Bilina, Jul. 1873, de Thümen, Thüm., Herb. myc. oec. 237 (B 70-6428). 

fumago [Link] f. vitis Thüm., Herb. myc. oec., Fasc. II, No. 76 (1872)!, nom. nud. 

T: on Vitis vinifera (Vitaceae), Germany, Lübeck, near Travemünde, Oct. 1872, Behrens, 

Thüm., Herb. myc. oec. 76 (B). 

fumago [Link] var. betulae L.A. Kirchn., Lotos 6: 183 (1856)!, nom. nud. 

T: on living leaves of Betula alba (Betulaceae), Czech Republic, Haasmüllnerberg near 

Kaplitz. 

fumago [Link] ‘var. corticola Hoffm.’, Herb. mycol., Ed. nova, Ser. prima, No. 330 (1857)!, 

nom. nud. [see OUDEMANS 1920: 189]. 

41

Notes: OUDEMANS (l.c.) spelled the name differently than C. fumago [Link] f. corticicola 

Rabenh., changed ‘f.’ to ‘var.’ and attributed this name to ‘Hoffm.’ although the exsiccatus 

has the same number. Correct quotation → C. fumago f. corticicola. 

fumago [Link] var. elongatum Mont., ‘Fl. J. Fern., n. 53’ (SACCARDO 1882: 81), reference not 

found. 

T: on leaves, especially of ferns, Chile, Juan Fernandez. 

≡ Antennaria robinsonii Berk. & Mont., in Berkeley, London J. Bot. 2: 641 (1843). 

Lit.: SACCARDO (1882: 81). 

fumago [Link] var. epiphyllum Rabenh. – Exsiccatus (68). I specified in KOHLMEYER (1962: 

39). 

fumago [Link] var. maculaeforme Thüm., Mycoth. univ., Cent. VII, No. 673 (1877)! and 

Flora 61(7): 108 (1878)!. 

T: on living leaves of Syringa vulgaris (Oleaceae), Germany, Bavaria, Bayreuth, autumn 

1875, de Thümen, Thüm., Mycoth. univ. 673 (B; HAL; MICH). 

Notes: Excluded, generic affinity not yet clear. 

fumago [Link] var. padi L.A. Kirchn., Lotos 6: 184 (1856)!, nom. nud. 

T: on living leaves of Padus vulgaris (= ?Prunus padus) (Rosaceae), Czech Republic, Kaplitz. 

fumago [Link] var. rubi L.A. Kirchn., Lotos 6: 184 (1856)!, nom. nud. 

T: on leaves and stems of Rubus idaeus (Rosaceae), Czech Republic, Dreisesselberg, Jac. 

Jungbauer. 

fumago Mont., in Gay, Fl. chil. 8(1): 32 (1852), nom. illeg., homonym, non C. fumago Link, 

1824. 

T: on living leaves of Eugenia (Myrtaceae), Chile. 

= ? Napicladium fumago Speg., Revista Fac. Agron. Vet. La Plata, Ser. 2, 6: 190 (1910). 

≡ Hyphosoma fumago (Speg.) M.B. Ellis, in herb. 

Lit.: SACCARDO (1913: 1396). 

fumosa Preuss, in herb. 

Without detailed data, Nr. 386 (Mappe 274), herb. C.G.T. Preuss (B). 

fungorum (Pers.) Roum., F. sel. gall. exs., Cent. XXXIII, No. 3293 (1885)!, as ‘Pers.’. 

T: on an old carpophore (L 910.225-732). 

≡ Dematium herbarum [Pers.] γ fungorum Pers., Syn. meth. fung. 2: 699 (1801)!. 

≡ Dematium vulgare [Pers.] γ fungorum (Pers.) Pers., Mycol. eur. 1: 14 (1822)!. 

≡ Cladosporium herbarum [(Pers.: Fr.) Link] β fungorum (Pers.) Chevall., Fl. gén. env. Paris 

1: 36 (1826)!. 



Lit.: OUDEMANS (1919: 49), PRASIL & DE HOOG (1988: 52), HEUCHERT et al. (2005: 53). 


‘fungorum’ Sacc., Syll. fung. 2: 406 (1883)!. 

T: on Polyporus sp., USA, Pennsylvania, Bethlehem. 

Notes: Cited by SACCARDO (l.c.) in the text under Sphaeria cladosporiosa Schwein., Trans. 

Amer. Philos. Soc., N.S., 4(2): 211 (1832)!. It is unclear if Saccardo intended to introduce a 

new name or if he referred to Dematium herbarum [Pers.] γ fungorum Pers. 

funiculosum W. Yamam., Sci. Rep. Hyogo Univ. Agric., Ser. Agric. 4(1): 5 (1959)!. 

T: on leaves of Phaseolus chrysanthos (Fabaceae), Japan, Prov. Tamba, Tannan-cho, 

Komakura, 29 Oct. 1958, W. Yamamoto. 

Ill.: YAMAMOTO (1959: 6, Figs 21–24). 

Notes: This name is valid since the author only cited a single collection, which can be 

considered the holotype. 

42

furfuraceum McAlpine, Fungus Dis. Citrus Trees Austral.: 78 (1899)!. 

T: on fruits of Citrus medica (Rutaceae), Australia, Victoria, Doncaster and elsewhere. 

Lit.: SACCARDO (1902: 1058). 

Ill.: MCALPINE (1899: Figs 19–20). 

Notes: Type material is not preserved in the McAlpine herbarium (VPRI). 

fuscatum Link, in Willd., Sp. pl. 6(1): 41 (1824)!, non C. fuscum Link, 1824. 

T: on herbal stems [on Passiflora edulis (Passifloraceae) and Avena sativa (Poaceae)] (L 

910.225-720). 

≡ Dematium fuscum Pers., Mycol. eur. 1: 16 (1822)!. 



Lit.: CROUS et al. (2000). 

Notes: PRASIL & DE HOOG (1988) re-examined type material of Dematium fuscum Pers. but 

did not find any Cladosporium species. 

fuscum Link, in Willd., Sp. pl. 6(1): 40 (1824)!: Fr., Syst. mycol. 3(2): 372 (1832)!. 

T: on stems of Rosa sp. (Rosaceae), Germany, Leipzig (Kunze) and Berlin (Link) (B). 

Lit.: FRIES (1832: 372), SACCARDO (1886: 352), LINDAU (1907: 805), FERRARIS (1912: 335), 

OUDEMANS (1921), GONZÁLES-FRAGOSO (1927: 201). 

fusicladiiformis Gonz. Frag., Trab. Mus. Nac. Ci. Nat., Ser. Bot. 10: 188 (1916). 

T: on living leaves of Eriobotrya japonica (Rosaceae), Spain, near Dos Hermanas, Sevilla, 22 

Mar. 1915, Gonzáles Fragoso (MA 06413: holotype). 

Lit.: GONZÁLES-FRAGOSO (1927: 205), SACCARDO (1931: 795). 

Host(s)/substrate(s) & distribution: on Eriobotrya japonica; Europe (Spain). 

fusicladium Sacc., in Bresadola & Saccardo, Malpighia 11: 321 (1897)!. 

T: on leaves of Salix incana (Salicaceae), Italy, Riva-Valsesia, 28 Apr. 1891, Carestia, No. 

770 (PAD: holotype). 




fusiforme (S.M. Reddy & Bilgrami) de Hoog & Morgan-Jones (?). 

≡ Hyalodendron fusiforme S.M. Reddy & Bilgrami, Curr. Sci. 40(24): 668 (1971). 

≡ Retroconis fusiformis (S.M. Reddy & Bilgrami) de Hoog & Bat. Vegte, Stud. Mycol. 31: 99 

(1989). 

fusisporum Berk. & M.A. Curtis, in herb. 

On stems of Chionanchus sp. (?), USA, Alabama (NYS). 

galii Mułenko, K. Schub. & M. Kozlowska, Mycotaxon 90(2): 272 (2004). 

T: on Galium odoratum (Rubiaceae), Poland, Nizina Północnopodlaska, Białowieża Forest, 

Białowieża National Park, Forest Compartment 342, Permanent plot No 40 BSG UW, single 

collection in oak-linden-hornbeam forest (Tilio-Carpinetum), 26 Sept. 1992, leg. W. Mułenko 

(LBLM-8459: holotype). Isotype: HAL 1811 (F). 

Ill.: MUŁENKO et al. (2004: 273, Fig. 1). 

Host(s)/substrate(s) & distribution: on Galium odoratum; Europe (Poland). 

gallicola B. Sutton, Mycol. Pap. 132: 37 (1973)!. 

T: on globose galls of Endocronartium harknessii (Cronartiaceae) on twigs of Pinus 

banksiana (Pinaceae), Canada, Saskatchewan, 27 mi S. Meadow Lake, 25 May 1967, C. 

Rentz, WINF (M) 6898e (IMI 145204: holotype). 

Lit.: ELLIS (1976: 329), HEUCHERT et al. (2005: 28–31). 

Ill.: SUTTON (1973: 38–39, Figs 17–18), ELLIS (1976: 329, Fig. 247), HEUCHERT et al. (2005: 

29–30, Figs 8−9; Pl. 1, Fig. 5; Pl. 2, Fig. 8). 

Host(s)/substrate(s) & distribution: hyperparasitic on galls of Endocronartium harknessii and 

Cronartium comandrae on Pinus, including P. banksiana and P. contorta var. latifolia, also on 

43

Pucciniastrum goeppertianum on Abies grandis; North America (Canada: very common, 

USA). 

geniculatum Morgan-Jones, in Morgan-Jones & Jacobsen, Mycotaxon 32(1): 226 (1988)!. 

T: on discoloured wall plaster, USA, Florida, Jacksonville, Airport Holiday Inn, Dec. 1987, 

B.J. Jacobsen (AUA: holotype). 

Ill.: MORGAN-JONES & JACOBSEN (1988: 227, Fig. 1). 

Notes: Listed as C. geniculatum Morgan-Jones & B.J. Jacobsen by KIRK et al. (n.d.). 

gentianae Lobik, Bolezni Rast. 17(3–4): 189 (1928)!. 

T: on leaves of Gentiana cruciata (Gentianaceae), Russia (LE 40527: holotype). 

Ill.: LOBIK (1928: Tab. 8, Fig. 87). 

Host(s)/substrate(s) & distribution: on Gentiana cruciata; Asia (Kazakhstan, Russia). 

georginae Fuss, Archiv Verein siebenb. Landesk., N.F., 14(2): 431 (1878)!, nom. nud. 

On overwintered stems of Georgina variabilis (= Dahlia pinnata) (Asteraceae), Romania, 

Giresau, Fuss. 

Notes: a formae of C. herbarum. 

gerwasiae Heuchert, U. Braun & K. Schub., Schlechtendalia 13: 31 (2005). 

T: on Gerwasia (Uredinales) sp. on leaves of Rubus urticifolius (Rosaceae), South America, 

Guatemala, Chimaltenango, Tecpan, Xecoxol, ‘plantation ex Rubus urticifolius’, G.A. Alvarez 

[N 140°51.8; W 90°59] (KR 5684: holotype). 

Ill.: HEUCHERT et al. (2005: 32, Fig. 10; Pl. 1, Figs 1–2). 

Host(s)/substrate(s) & distribution: on Gerwasia sp. on Rubus urticifolius; South America 

(Guatemala). 

gleditschiae Cooke, Grevillea 17(83): 66 (1889)!. 

T: on legumes of Gleditsia (Caesalpiniaceae), USA, South Carolina, Aiken, Rav., F. amer. 

exs. 297 (BPI 426765; K 121559; PH 01020431: syntypes). 

Lit.: SACCARDO (1892: 603, 1895: 621). 

glochidionis C.D. Sharma, Gadp., Firdousi, A.N. Rai & K.M. Vyas, Indian Phytopathol. 

51(2): 152 (1998)!. 

T: on living leaves of Glochidion sp. (Euphorbiaceae), India, Madhya Pradesh, Shahdol circle, 

Amarkantak (south forest division), Kapil dhara, Jan. 1993, C.D. Sharma (S.U. Herb. No. 

C.S.1: holotype; IMI 356765: isotype). 

Ill.: SHARMA et al. (1998: 153, Fig. 1). 

gloeosporioides G.F. Atk., Cornell Univ. Sci. Bull. 3(1): 39 (1897)!. 

T: on leaves of Ascyrum stans (= Hypericum stans), USA, Alabama, Lee Co., Auburn, 29 

Aug. 1891, G.F. Atkinson (CUP-A2064: syntype); on leaves and stems of Hypericum mutilum 

(Hypericaceae), USA, Alabama, Lee Co., Auburn, 2 Sept. 1891, Duggar. (CUP-A2170: 

lectotype). 

≡ Dischloridium gloeosporioides (G.F. Atk.) U. Braun & K. Schub., in Schubert & Braun, 

Fungal Diversity (2005), in press. 

Lit.: SACCARDO (1899: 1080). 


gonorrhoicum Hallier, Flora, Neue Reihe, 26(19): 293 (1868)!. 

T: isolated from man (gonorrhea). 

≡ Cladosporium coniothecii-gonorrhoici Hallier, Flora, Neue Reihe, 26(19): 293 (1868)! 

(alternative name). 

Ill.: HALLIER (1868: Tab. 3, Fig. 4, 5, 8, 9). 

Notes: Undoubtedly not belonging to Cladosporium s. str. Described as state (morphe) of 

Coniothecium gonorrhoicum Hallier (p. 294). “Forma penicillare. Hab. Associato a casidi 

gonorrhea”(NANNIZZI 1934). “Species incertae” (SACCARDO 1911). 

gossypii Jacz., Bolezni khlpchatnica: 181–182 (1931)!. 

T: on fibres of Gossypium sp. (Malvaceae), Russia (LEP). 

Ill.: JACZEWSKI (1931: 181, Fig. 16). 

44

gossypiicola Pidopl. & Deniak, in Pidoplichko, Gribnaja Flora Grubych Kormov: 273 

(1953)!, as ‘gossypicola’, nom. inval. 

T: on seeds of cotton (Gossypium sp., Malvaceae), also found on damp straw, Ukraine. 

Lit.: HO et al. (1999: 128). 

Ill.: PIDOPLICHKO (1953: 274, Fig. 75), HO et al. (1999: 130, Fig. 20). 

Notes: ZHANG et al. (2003) cited collections on Bombax malabaricum (China, Shaanxi), 

Gossypium hirsutum (China, Henan), and Gossypium sp. (China, Hubei). 

gossypiicola var. minor Pidopl. & Deniak, in Pidoplichko, Gribnaja Flora Grubych Kormov: 

273 (1953)!, nom. inval. 

T: on seeds of cotton (Gossypium sp., Malvaceae), Ukraine, Krim. 

gougerotii (Matr.) G. Carrión & Marg. Silva, Arch. Inst. Pasteur Algérie 72: 532 (1955)!. 

T: isolated from cystic lesion of human patient, USA, Memphis (CBS 526.76: neotype, 

selected by BORELLI, Acta Ci. Venez. 6(2): 81, 1955). 

≡ Sporotrichum gougerotii Matr., Compt. Rend. Hebd. Séances Acad. Sci. 150: 545 (1910)!, 

as ‘gougeroti’, nom. ambig. et dub. 

≡ Rhinocladium gougerotii (Matr.) Verdun, Précis de Parasitologie Humaine, éd. 2: 677 

(1913). 

≡ Dematium gougerotii (Matr.) Grigoraki, Bull. Soc. Mycol. France 40: 274 (1924), as 

‘gougeroti’. 

≡ Torula gougerotii (Matr.) Brumpt, Précis Parasitol., ed. 5: 1791 (1936). 

≡ Oospora gougerotii (Matr.) D. Janke, Archiv Dermatol. Syph. 187: 693 (1949), as 

‘Gougeroti’. 

≡ Phialophora gougerotii (Matr.) Borelli, Acta Ci. Venez. 6(2): 81 (1955). 

Lit.: DE HOOG & HERMANIDES-NIJHOF (1977: 114), MCGINNIS & AJELLO (1982). 

Notes: Of S. gougeroti: “This name has been applied to different fungi and has therefore been 

abandoned.” (DE HOOG et al. 2000: 1022). MCGINNIS & AJELLO (1982) cite a lack of type 

material and illustrations, and inadequate descriptions as reasons for considering S. gougeroti 

a dubious name. CARRIÓN & SILVA (1955) designate No. 1792 (Kennedy Hospital, Memphis), 

No. 7031 (NIH � CBS) and No. 7028 (NIH) as specimens of Cladosporium gougerotii. They 

illustrate in photomicrographs Nos. 1792 and 7028 as possessing phialides. 

gracile Corda, Icon. fung. 1: 15 (1837)!. 

T: on rotten leaves of Quercus sp. (Fagaceae), Czech Republic, near Prag, Corda (PRM). 

≡ Didymotrichum gracile (Corda) Bonord., Handb. Mykol.: 89 (1851)!. 



Lit.: SACCARDO (1886: 361), LINDAU (1907: 820), FERRARIS (1912: 343; 1914: 884), 

GONZÁLES-FRAGOSO (1927: 203). 


gramineum Link (FRIES 1832) under “Species dubias”. 

graminum Cooke, DOV [A. Commons 30,2060 (SUMSTINE 1949: 19)]. 

graminum Corda, Icon. fung. 1: 14 (1837)!, nom. illeg., homonym, non C. graminum (Pers.) 

Link, 1824. 

T: on rotten leaves of Gramineae, Czech Republic, near Prag. 




Notes: In CORDA (1837): ‘C. graminum (Link. p. 42 ?)’. 

graminum [Corda] f. bambusae Roum., F. sel. gall. exs., Cent. XLII, No. 4191 (1887)!, nom. 

nud. 

T: on leaves of Bambusa sp., France, Dept. Haute-Garonne, Vallee du Lys, Bagnères-de- 

Luchon, park of casino, Dec. 1886, Ch. Fourcade, Roum., F. sel. gall. exs. 4191 (B). 

45

graminum [Corda] f. inflorescentiae Sacc., Syll. fung. 25: 791 (1931)!. 

T: on flowers of Baldingera arundinacea (= Phalaris arundinacea) (Poaceae), France, Hariot. 

graminum [Corda] var. ‘moliniae’ Sacc., Ann. Mycol. 3: 169 (1905)!. 

Lit.: SACCARDO (1906: 577), FERRARIS (1912: 338). 

Notes: This reference is actually to var. moliniae-caeruleae. See next entry. 

graminum [Corda] var. moliniae-caeruleae Sacc., Ann. Mycol. 3: 169 (1905)!. 

T: on leaves of Molinia caerulea (Poaceae), Italy, Treviso, Montello, Aug. 1904, P.A. 

Saccardo, mixed infection with Sphaerella montellica Sacc. (BPI 426831A). 

Lit.: LINDAU (1907: 815). 

graminum (Pers.) Link, in Willd., Sp. pl. 6(1): 42 (1824)!. 

T: on dry leaves and culms of Gramineae (L 910.225-723). 

≡ Dematium graminum Pers., Mycol. eur. 1: 16 (1822)!. 

≡ Chloridium graminum (Pers.) Chevall., Fl. gén. env. Paris 1: 35 (1826)!. 

= Cladosporium graminum Corda, Icon. fung. 1: 14 (1837)!, nom. illeg., homonym, non C. 

graminum (Pers.) Link, 1824. 



Lit.: SACCARDO (1886: 365, 1972: 1352), LINDAU (1907: 815), FERRARIS (1912: 337), LIND 

(1913: 523), GONZÁLES-FRAGOSO (1927: 208), DE VRIES (1952: 71, as ‘C. graminum Pers. ex 

Corda’), DAVID (1997: 10). 

Ill.: GONZÁLES-FRAGOSO (1927: 209, Fig. 46). 

graminum [(Pers.) Link] f. poae-pratensis Thüm., Mycoth. univ., Cent. V, No. 490 (1876)!, 

nom. nud. 

T: on dry culms of Poa pratensis (Poaceae), Germany, Bavaria, Bayreuth, summer 1874, de 

Thümen, Thüm., Mycoth. univ. 490 (e.g., HAL; HBG). 

graminum var. scirpi, in herb. 

On dead plants of Scirpus sp., USA, California, Apr. 1880 (BPI 426819). 

graminum var. sorghi Rav., in herb. 

On Sorghum saccharatum (Poaceae), USA, South Carolina, H.W. Ravenel (BPI 426822). 

grech-delicatae Sacc., Ann. Mycol. 11: 564 (1913)! and Nuovo Giorn. Bot. Ital., N.S. 21(1): 

125 (1914)!. 

T: on still living stems of Ranunculus aquatilis (Ranunculaceae), Malta, Uied Bufula, Apr. 

1913, Doct. Borg (PAD). 

Lit.: SACCARDO (1931: 794). 

Host(s)/substrate(s) & distribution: on Ranunculus aquatilis; Europe (Malta). 

grewiae Bacc., Ann. Bot. (Rome) 4: 277 (1906)!, as ‘graeviae’. 

T: on dead leaves of Grewia (Tiliaceae), Eritrea, Mai Hiryi, Pappi. 

Lit.: SACCARDO (1913: 1368). 

griseo-olivaceum Pidopl. & Deniak, Mikrobiol. Zhurn. 5(2): 183, 193–194 (1938)!, nom. 

inval. 

T: on grains of Zea mays (Poaceae), Ukraine. 



Notes: Neither in Mikrobiol. Zhurnal (1938) nor in PIDOPLICHKO (1953) is a Latin diagnosis 

given. 

griseum (Berk. & Broome) S. Hughes, Canad. J. Bot. 31: 587 (1953)!. 

T: on dead stems of Urtica sp., Great Britain, Northamptonshire, Kings Cliff, Mar. 1850 (K 

121543: isotype). 

≡ Dendryphion griseum Berk. & Broome, Ann. Mag. Nat. Hist., Ser. 2, 7: 177 (1851), as 

‘Dendryphium’, non Polyscytalum griseum Sacc., 1886. 

46

≡ Polyscytalum berkeleyi M.B. Ellis, More Dematiaceous Hyphomycetes: 158 (1976)!, nom. 

nov. 

Notes: HUGHES (1953) apparently based his new combination and illustration on IMI 4333. 

grumosum (Pers.) Link, in Willd., Sp. pl. 6(1): 42 (1824)!. 

T: not specified in the original publication. 

≡ Dematium grumosum Pers., Mycol. eur. 1: 16 (1822)!. 

Lit.: FRIES (1832: 373), CORDA (1837: 14), SACCARDO (1886: 363), LINDAU (1907: 822). 

Notes: In OUDEMANS (1919) Pteris aquilina (= Pteridium aquilinum) is mentioned as host 

species. See C. aquilinum. 

guanicense F. Stevens, Trans. Illinois Acad. Sci. 10: 207 (1917), as ‘guanicensis’. 

T: on leaves of Argemone mexicana (Papaveraceae), Puerto Rico, Guanica, 2 Mar. 1913, F.L. 

Stevens, Porto Rican Fungi 347 (a) (BPI 426841, 426844, 426487; ILL 15873; MICH; PC). 

≡ Polythrincium guanicense (F. Stevens) Cif., Ann. Mycol. 36: 233 (1938)!. 

≡ Cercosporidium guanicense (F. Stevens) Deighton, Mycol. Pap. 112: 34 (1967)!. 

≡ Passalora guanicensis (F. Stevens) U. Braun & R.F. Castañeda, in Castañeda Ruiz & 

Braun, Cryptog. Bot. 1(1): 46 (1989)!. 

≡ Passalora guanicensis (F. Stevens) Poonam Srivast., J. Liv. World 1(2): 116 (1994), as 

‘guanicense’, comb. inval. 

= Cercospora whetzelii Chupp, J. Dept. Agric. Porto Rico 15: 16 (1931). [T: CUP]. 

≡ Piricularia whetzelii (Chupp) Bat. & R. Garnier, Publ. Univ. Recife Inst. Micol. 278: 18 

(1960). 


gynoxidicola Petr., Sydowia 2: 381 (1948)!, as ‘gynoxidicolum’. 

T: on living leaves of Gynoxys sp. (Asteraceae), Ecuador, Pichincha mountains near Quito, 30 

Nov. 1937 (M-57615; IMI 88949: slide). 

≡ Stenella gynoxidicola (Petr.) J.L. Mulder, Trans. Brit. Mycol. Soc. 79(3): 478 (1982)!. 

≡ Passalora gynoxidicola (Petr.) K. Schub. & U. Braun, Fungal Diversity (2005), in press. 

Lit.: ELLIS (1976: 342). 


haplophylli (Vasyag. & Tartenova) J.C. David, Mycol. Pap. 172: 85 (1997)!. 

T: on leaves of Haplophyllum latifolium (Rutaceae), Kazakhstan, Chimkentskaya Oblast, in 

the southern Kyzyl-Kum desert, 10 Jun. 1957, Tartenova (holotype: location not known); on 

leaves of Haplophyllum latifolium, Kazakhstan, Chimkentskaya Oblast, Talasskii Alatau, 

Zapovednik Aksu-Dzhabagly, by the river Aksu, 23 Jul. 1968, M.P. Vasjagina (AA: paratype). 

≡ Heterosporium haplophylli Vasyag. & Tartenova, in Shvartsman et al., Fl. sporov. rast. 

Kazakhstana 8(2): 164 (1975). 

Ill.: DAVID (1997: 87, Fig. 20). 

Host(s)/substrate(s) & distribution: on species of Haplophyllum, including H. acutifolium and 

H. latifolium; Central Asia (Kazakhstan, Turkmenistan). 

harknessii (Peck) S. Hughes, Canad. J. Bot. 31: 586 (1953)!. 

T: on decaying wood, USA, New York, Helderberg Mountains, C.H. Peck (NYS 1443: 

holotype; DAOM 31921: isotype, slide). 

≡ Monilia harknessii Peck, Rep. (Annual) New York State Mus. Nat. Hist. 34: 49 (1883). 

Ill.: HUGHES (1953: 586, Fig. 7). 

Notes: HUGHES (1953) apparently based his new combination and illustration on DAOM 

28997. 

hederae, in herb. (without author). 

On leaves of Hedera sp. (HBG). 



Notes: on the web site of HBG as ‘rederse’ 

47

heleophilum J.C. David, Mycol. Pap. 172: 80 (1997)!. 

T: on Typha latifolia (Typhaceae), USA, Washington, Seattle, 26 Aug. 1912, Bartholomew, 

Barthol., F. columb. 4407, sub C. herbarum (K: holotype). 

Ill.: DAVID (1997: 62, Fig. 17 J–L). 

Host(s)/substrate(s) & distribution: on Typha latifolia; Europe (Germany, Great Britain), 

North America (USA). 

Notes: Presumably more common than indicated, because it was confused with C. herbarum 

or similar species occurring on Typha spp. 

helicosporum R.F. Castañeda & W.B. Kendr., in Castañeda Ruiz, Kendrick & Gené, Mycotaxon 

63: 183 (1997)!. 

T: on fallen leaves of Laciasis divaricata from a rainforest, Cuba, La Habana, La Chorrera, 14 

Apr. 1992, R.F. Castañeda & B. Kendrick (INIFAT C92/207-1: holotype; MUCL 39868: 

isotype). 

Ill.: CASTAÑEDA et al. (1997: 184–185, Figs 1–2). 

heliotropii Erikss., Bot. Centralbl. 47: 299 (1891)! and F. paras. scand., Fasc. 8, No. 396 

(1891)!, with description on the label. 

T: on leaves of Heliotropium peruvianum (Boraginaceae), Sweden, Stockholm, Rosendal, 

1882, J. Eriksson, F. paras. scand. 396 (e.g., BPI 426853; HAL; HBG). 


Host(s)/substrate(s) & distribution: on species of Heliotropium, including H. peruvianum; 

Europe (Sweden), North America (USA). 

helminthosporioides (Corda) Fr., Summa veg. Scand. 2: 499 (1849)!. 

T: on dead leaves of conifers, Czech Republic. 

≡ Azosma helminthosporioides Corda, in Sturm, Deutsch. Fl. 3(12): 35 (1831)!. 

≡ Macrosporium helminthosporioides (Corda) Sacc. & Traverso, Syll. fung. 20: 8 (1911)!. 

Ill.: CORDA (1831: Tab. 18). 

Notes: Type material could not be located in herb. PRM. 

hemileiae Steyaert, Bull. Soc. Roy. Bot. Belgique 63(1): 47 (1930)!. 

T: on uredosoris of Hemileia vastatrix (Uredinales) on Coffea robusta (Rubiaceae), Zaire, 

Prov. Orientale, Biaro, near Kisangani (Stanleyville), Oct. 1929, R.L. Steyaert (BPI 426854: 

holotype). 

≡ Digitopodium hemileiae (Steyaert) U. Braun, Heuchert & K. Schub., in Heuchert et al., 

Schlechtendalia 13: 65 (2005). 

Lit.: SACCARDO (1972: 1337), SUTTON (1973: 40). 

Ill.: STEYAERT (1930: Tab. 4–5), HEUCHERT et al. (2005: 65, Fig. 26; Pl. 1, Figs 4,7). 

herbarum (Pers.: Fr.) Link, Ges. Naturf. Freunde Berlin Mag. Neuesten Entdeck. Gesammten 

Naturk. 7: 37 (1816)!: Fr., Syst. mycol. 3(2): 370 (1832)!. 

T: (L 910.225-733: lectotype). 

≡ Dematium herbarum Pers., Ann. Bot. (Usteri) 11: 32 (1794): Fr., Syst. mycol. 3(2): 370 

(1832)!. 

≡ Acladium herbarum (Pers.) Link, Ges. Naturf. Freunde Berlin Mag. Neuesten Entdeck. 

Gesammten Naturk. 3: 12 (1809)!. 

≡ Byssus herbarum (Pers.) DC., Fl. franç., Ed. 3, 5: 11 (1815)!, as ‚Bissus’. 

≡ Dematium vulgare Pers., Mycol. eur. 1: 13 (1822)!, nom superfl. 

≡ Dematium vulgare [Pers.] α herbarum (Pers.) Pers., Mycol. eur. 1: 13 (1822)!. 

= ? Byssus caespitosa Roth, Catal. Bot. 1: 215 (1797). 

= ? Dematium herbarum [Pers.] β brassicae Pers., Syn. meth. fung. 2: 699 (1801)!. 

= ? Dematium cinnabarinum Pers., Syn. meth. fung. 2: 697 (1801)!. 

= Dematium epiphyllum Pers., Syn. meth. fung. 2: 695 (1801)!. 

≡ Cladosporium epiphyllum (Pers.) Nees, Syst. Pilze 1: 67 (1817)!. 

≡ Cladosporium epiphyllum (Pers.) Link, in Willd., Sp. pl. 6(1): 42 (1824)!. 

≡ Chloridium epiphyllum (Pers.) Chevall., Fl. gén. env. Paris 1: 35 (1826)!. 

48

≡ Cladosporium epiphyllum (Pers.) Fries, Syst. mycol. 3(2): 370 (1832)!. 

= Dematium herbarum [Pers.] γ fungorum Pers., Syn. meth. fung. 2: 699 (1801)!. [T: L 

910.225-732]. 

≡ Dematium vulgare [Pers.] γ fungorum (Pers.) Pers., Mycol. eur. 1: 14 (1822)!. 

≡ Cladosporium herbarum [(Pers.: Fr) Link] β fungorum (Pers.) Chevall., Fl. gén. env. Paris 

1: 36 (1826)!. 

≡ Cladosporium fungorum (Pers.) Roum., F. sel. gall. exs., Cent. 33, No. 3293 (1885)!, as 

‘Pers.’. 

= ? Dematium conicum Schum., Emun. Saell. 2: 445 (1803). 

= Dematium graminum Pers., Mycol. eur. 1: 16 (1822)!. 

≡ Cladosporium graminum (Pers.) Link, in Willd., Sp. pl. 6(1): 42 (1824)!. 

≡ Chloridium graminum (Pers.) Chevall., Fl. gén. env. Paris 1: 36 (1826)!. 

= Dematium fuscum Pers., Mycol. eur. 1: 16 (1822)!. 

≡ Cladosporium fuscatum Link, in Willd., Sp. pl. 6(1): 41 (1824)!, non C. fuscum Link, 1824. 

= Cladosporium typharum Desm., Pl. crypt. N. France, Ed. 1, Ser. 2, Fasc. VII, No. 304 

(1828). 

= Cladosporium sparsum Schwein., Trans. Amer. Philos. Soc., N.S., 4(2): 277 (1832)!. [T: PH 

1020413, 1020414] 

= Helminthosporium herbarum Schwein., Trans. Amer. Philos. Soc., N.S. 4(2): 279 (1832)!. 

= Acladium heterosporum Wallr., Flora crypt. Germ. 2: 287 (1833)!. 

= Myxocladium arundinis Corda, Icon. fung. 1: 12 (1837)!. [T: PRM]. 

≡ Cladosporium arundinis (Corda) Sacc., Syll. fung. 4: 364 (1886)!. 

= Helminthosporium flexuosum Corda, Icon. fung. 1: 13 (1837)!. [T: PRM]. 

≡ Brachysporium flexuosum (Corda) Sacc., Syll. fung. 4: 429 (1886)!. 

= Cladosporium alnicola Corda, Icon. fung. 1: 14 (1837)!. [T: PRM]. 

≡ Didymotrichum alnicola (Corda) Bonord., Handb. Mykol.: 89 (1851)!. 

= Cladosporium caricicola Corda, Icon. fung. 1: 14 (1837)!. [T: PRM]. 

≡ Didymotrichum caricicola (Corda) Bonord., Handb. Mykol.: 89 (1851)!. 

= Cladosporium graminum Corda, Icon. fung. 1: 14 (1837)!, nom. illeg., homonym, non C. 

graminum (Pers.) Link, 1824. 

= Cladosporium fasciculatum Corda, Icon. fung. 1: 15 (1837)!. [T: PRM]. 

= Cladosporium lignicola Corda, Icon. fung. 1: 14 (1837)!. [T: PRM]. 

= Cladosporium nodulosum Corda, Icon. fung. 1: 15 (1837)!. [T: PRM]. 

≡ Didymotrichum nodulosum (Corda) Bonord., Handb. Mykol.: 89 (1851)!. 

= Cladosporium tomentosum Corda, Icon. fung. 1: 15 (1837)!. [T: PRM]. 

= ? Cladosporium herbarum var. typharum Westend., Cryptogames: 26 (1854), nom. inval. 

= Cladosporium amaranticola Opiz, Lotos 5: 41 (1855)!, nom. nud. [T: PRM]. 

= Cladosporium caespiticium Rabenh., F. eur., Cent. VI, No. 579 (1863) and Bot. Zeitung 

(Berlin) 21: 230 (1863) (nom. nud.), syn. nov. [T: HAL; HBG; M]. 

= Cladosporium fuligineum Bonord., Abh. Mykol. 1: 92 (1864)!. 

= Cladosporium microporum Rabenh., Unio itin. crypt., No. 42 (1866) (nom. nud.), syn. nov. 

[T: HBG; M]. 

≡ Cladosporium microporum Rabenh., in Cooke, Grevillea 17(83): 66 (1889), syn. nov. 

= Helminthosporium vesiculosum Thüm., Mycoth. univ., Cent. VIII, No. 784 (1877)!. [T: 

HAL; PAD]. 

≡ Brachysporium vesiculosum (Thüm.) Sacc., Syll. fung. 4: 429 (1886)!. 

= Cladosporium elegans Penz., in Saccardo, Michelia 2(8): 471 (1882)!. [T: HBG; PAD]. 

= Cladosporium punctulatum Sacc. & Ellis, Michelia 2(8): 578 (1882), syn. nov. [T: BPI; NY; 

PAD]. 

= Cladosporium brunneum Cooke & Harkn., Grevillea 12: 96 (1884)!, nom. illeg., homonym, 

non C. brunneum Corda, 1837, syn. nov. [T: BPI; K]. 

≡ Cladosporium brunneolum Sacc., Syll. fung. 4: 358 (1886)!, syn. nov. 

= Helminthosporium phyllophilum P. Karst., Hedwigia 23: 41 (1884)!. [T: H]. 

= Heterosporium abroniae Harkn., Bull. Calif. Acad. Sci. 1: 38 (1884). [T: BPI]. 

= Cladosporium kniphofiae Cooke, Grevillea 14(70): 40 (1885), syn. nov. [T: K]. 

49

= Helminthosporium nanum [Nees] f. petiolicola Roum., F. gall. exs., No. 3391 (1885). [T: G; 

PC]. 

= Heterosporium epimyces Cooke & Massee, in Cooke, Grevillea 16(79): 80 (1888)!. [T: K 

121555]. 

= Heterosporium laburni Oudem., Ned. Kruidk. Arch., Ser. 2, 5(2): 174 (1888)!. [T: L]. 

= Cladosporium condylonema Pass., in Briosi & Cavara, F. paras., No. 79 (1889)!. 

= ? Heterosporium fungicola Ellis & Everh., J. Mycol. 5: 70 (1889)!, as ‘fungicolum’. 

= Heterosporium goiranicum C. Massal., Nuovo Giorn. Bot. Ital. 21: 170 (1889)! and Mem. 

Accad. Agric. Verona, Ser. III, Fasc. 2, 65: 117 (1889). [T: VER]. 

= Cladosporium velutinum Ellis & Tracy, J. Mycol. 6: 76 (1890)!, syn. nov. [T: NY]. 

= Cladosporium phoenicis Roum., F. sel. gall. exs., Cent. 58, No. 5798 (1891) and Rev. 

Mycol. (Toulouse) 13: 133 (1891), as ‘phaenicis’, syn. nov. [T: B; FH]. 

= Helminthosporium acuum P. Karst., Hedwigia 31: 295 (1892)!. [T: H]. 

= Helminthosporium compactum P. Karst., Hedwigia 31: 295 (1892)!. [T: H]. 

= Heterosporium galii Fautrey & Roum., Rev. Mycol. (Toulouse) 14: 106 (1892). [T: PC]. 

= Heterosporium caulicola Ellis & Everh., Proc. Acad. Nat. Sci. Philadelphia 46: 381 (1894)!, 

as ‘caulicolum’. [T: NY]. 

= Heterosporium cladosporioides Ellis & Everh., Proc. Acad. Nat. Sci. Philadelphia 46: 382 

(1894)!. [T: NY]. 

= Heterosporium didymosporum Clem., Bot. Surv. Nebraska 3, 1893(2): 11 (1894). [T: NEB]. 

= Cladosporium menispermi Allesch., Hedwigia 34: 220 (1895), syn. nov. [T: M]. 

= Cladosporium acutum Ellis & Dearn., Proc. Canad. Inst., N.S., 3, 1: 91 (1897)!. [T: 

DAOM]. 

= Cladosporium fusicladium Sacc., in Bresadola & Saccardo, Malpighia 11: 321 (1897), syn. 

nov. [T: PAD]. 

= Heterosporium avenae Oudem., Hedwigia 37: 318 (1898)!. [T: L]. 

= Heterosporium syringae Oudem., Hedwigia 37: 183 (1898)!. [T: L]. 

= ? Cladosporium fagi Oudem., Ned. Kruidk. Arch., Ser. 3, 2(3): 768 (1902)! and Beih. Bot. 

Centralbl. 11: 538 (1902)!. 

= Cladosporium laricis Sacc., Ann. Mycol. 3: 515 (1905), syn. nov. [T: PAD]. 

= Heterosporium ephedrae Potebnia, Ann. Mycol. 5: 21 (1907)! and Trudy Obshch. Isp. Prir. 

Imp. Khar’kovsk. Univ. (Trudy Oshch. Estest. imp. Khar’kov Univ.) 1907: 43 (1907). [T: 

PAD]. 

= Heterosporium fraxini Ferd. & Winge, Bot. Tidsskr. 28(2): 256 (1907)!. [T: C]. 

= Heterosporium opuntiae Lindau, in Rabenhorst, Krypt.-Fl., ed. 2, 1(9): 84 (1910)!. [T: B]. 

= Heterosporium berberidis Ranoj., Ann. Mycol. 8: 399 (1910)!. [T: BPI]. 

= Heterosporium cytisi Ranoj., Ann. Mycol. 8: 398 (1910)!. 

= Cladosporium vincae Fairm., Ann. Mycol. 9: 148 (1911)!, syn. nov. [T: CUP-F2873(24- 

68)]. 

= Heterosporium asperatum Massee ex Sacc., Syll. fung. 22: 1388 (1913)!. [T: K]. 

= Heterosporium spiraeae Syd. & P. Syd., Ann. Mycol. 11: 406 (1913)!. [T: S]. 

= Heterosporium yuccae Bubák, Ann. Mycol. 12: 214 (1914)!. [T: BPI]. 

= Heterosporium sorghi Ranoj., Ann. Mycol. 12: 418 (1914)!. [T: BPI]. 

= Heterosporium stromatigenum Bubák & Vleugel, in Bubák, Ann. Mycol. 14: 351 (1916)!. 

[T: BPI]. 

= Cladosporium alnicola Bubák & Vleugel, in Vleugel, Svensk Bot. Tidskr. 11(3–4): 322 

(1917), nom. illeg., homonym, non C. alnicola Corda, 1837, syn. nov. [T: BPI]. 

= Heterosporium atopomerum Kirschst., Ann. Mycol. 37: 122 (1939)!. [T: B]. 

= Cladosporium moldavicum Fosteris, Bull. Sect. Sci. Acad. Roumaine 26(7): 494 (1944)? 

and in Herb. Mycol. Rom., Fasc. 27, No. 1341 (1944) (nom. inval.), syn. nov. [T: BPI; 

MA]. 

= Heterosporium equiseti H.C. Greene, Amer. Midl. Naturalist 44(3): 642 (1950)!. [T: WIS]. 

= Cladosporium hederae, in herb. (HBG), syn. nov. 

= Heterosporium caricis Grove, in herb. 

= Heterosporium pseudoplatani Grove, in herb. 

50

Teleomorph: Davidiella tassiana (De Not.) Crous & U. Braun, in Braun, Crous, Dugan, 


Lit.: COOKE (1871: 582), SACCARDO (1886: 350, 1972: 327, 1304), LINDAU (1907: 800, 1910: 

795), FERRARIS (1912: 331), GONZÁLES-FRAGOSO (1927: 194), DE VRIES (1952: 71), 

HUGHES (1958: 750), ELLIS (1971: 313), DOMSCH et al. (1980: 204), SIVANESAN (1984: 225), 

ELLIS & ELLIS (1985: 290, 468, 1988: 168), PRASIL & DE HOOG (1988), WANG & ZABEL 

(1990: 202), MCKEMY & MORGAN-JONES (1991c), DUGAN & ROBERTS (1994), DAVID 

(1997: 59), HO et al. (1999: 129), DE HOOG et al. (2000: 587), SAMSON et al. (2000: 110), 

SAMSON et al. (2001). 

Ill.: FERRARIS (1912: 327, Fig. 101), DE VRIES (1952: 73, Fig. 15), YAMAMOTO (1959: 2, Figs 

1–4), ELLIS (1971: 314, Fig. 217 A), DOMSCH et al. (1980: 206, Fig. 83), VON ARX (1987: 57, 

Fig. 27), PRASIL & DE HOOG (1988: 51, Fig. 3), MCKEMY & MORGAN-JONES (1991c: 311, Pl. 

1; 313, Fig. 1), DUGAN & ROBERTS (1994: 516, Figs 4–7), DAVID (1997: 62, Fig. 17 F, G, I), 

HO et al. (1999: 130, Figs 21–22), DE HOOG et al. (2000: 587–588, Figs), SAMSON et al. (2000: 

110, Fig. 49; 111, Pl. 47). 

Host(s)/substrate(s) & distribution: on different hosts and substrates, commonly on dead 

herbaceous plants; cosmopolitan. 

Notes: See PRASIL & DE HOOG (1988) on CBS 177.71, L no. 910.225-733 (lectotype) and for 

comments on the incorrect typification of one of Link’s secondary collections at herb. B de 

Vries (1952). In SACCARDO (1886), LINDAU (1907) and FERRARIS (1912) ‘Dematium 

pullulans de Bary & Löwenthal’ is cited as synonym of Cladosporium herbarum, but the 

current name is Aureobasidium pullulans (de Bary) G. Arnaud var. pullulans. 

herbarum [(Pers.: Fr.) Link] α aphidis Fuckel, Jahrb. Nassauischen Vereins Naturk. 23–24: 

356 ‘1869’ (1870)!, nom. nud. 

Lit.: OUDEMANS (1923). 

Notes: see C. herbarum var. aphidis. 

herbarum [(Pers.: Fr.) Link] α foliorum (Pers.) Chevall., Fl. gén. env. Paris 1: 36 (1826)!. 

T: on leaves of Iris, Brassica etc. 

≡ Dematium vulgare [Pers.] β foliorum Pers., Mycol. eur. 1: 14 (1822)!. 

herbarum [(Pers.: Fr.) Link] β fasciculare Corda, Icon. fung. 3: 9 (1839)!. 

Lit.: FERRARIS (1914: 882),OUDEMANS (1923). 

Notes: see C. herbarum var. fasciculare. 

herbarum [(Pers.: Fr.) Link] β fungorum (Pers.) Chevall. → fungorum (Pers.) Roum. 

herbarum [(Pers.: Fr.) Link] β nigricans (Roth: Fr.) Fr., Syst. mycol. 3(2): 371 (1832)!. 

Notes: see C. herbarum var. nigricans. 

herbarum [(Pers.: Fr.) Link] δ cerealium [Sacc.] f. hordei (Bruhne) Ferraris → bruhnei. 

herbarum [(Pers.: Fr.) Link] b solutum (Link) Rabenh. → solutum. 

herbarum [(Pers.: Fr.) Link] f. agaves-echeveriae Savelli, Ann. Reale Accad. Agric. Torino 

56: 113 (1914)! (dated 1913). 

T: on living leaves of Agave americana (Agavaceae) and Echeveria sp. (Crassulaceae), Italy, 

Torino, Mar. 1913. 

Lit.: FERRARIS (1914: 882), PRASIL & DE HOOG (1988: 53). 

Notes: PRASIL & DE HOOG (1988) found no authentic material in FI, PDA, PAV, RO, ROPV 

and TO. They think it unlikely that this was a Cladosporium. 

herbarum [(Pers.: Fr.) Link] f. amaranthi, in herb. 

On Amaranthus retroflexus (Amaranthaceae), Czech Republic, Moravia, Hranice (Maehr.weisskirchen), 

Jesernice, 3 Sept. 1914, F. Petrak (BPI 426907). 

herbarum [(Pers.: Fr.) Link] f. asparagi Thüm., Herb. myc. oec., Fasc. VII, No. 320 (1875)!, 

nom. nud. 

T: on Asparagus sp. (Asparagaceae), USA, Thüm., Herb. myc. oec. 320. 

Lit.: STEVENSON (1971). 

51

herbarum [(Pers.: Fr.) Link] f. camellia-japonicae Bubák, in Rabenhorst-Pazschke, F. eur., 

Cent. 43, No. 4289 (1901)!, nom. nud. 

T: on leaves of Camellia japonica (Theaceae), Czech Republic, Bohemia, Tabór, Mar. 1901, 

F. Bubák, Rabenh., F. eur. 4289 (BPI 426941; HBG: syntypes). 

= Cladosporium cladosporioides (Fresen.) G.A. de Vries, Contr. Knowl. Genus Cladosporium: 

57 (1952). 

herbarum [(Pers.: Fr.) Link] f. carpophilum Bacc., Ann. Bot. (Rome) 4: 277 (1906)!, as 

‘carpophylum’. 

T: on inflorescences of Panicum maximum (Poaceae), Eritrea. 

Lit.: SACCARDO (1913: 1366), PRASIL & DE HOOG (1988: 53). 

Notes: No material in PAD. 

herbarum [(Pers.: Fr.) Link] f. cerealium, in herb. 

On Koeleria cristata (Poaceae), USA, New Mexico, 25 Aug. 1916, P.C. Standley (BPI 

427189). 

Notes: See C. herbarum var. cerealium? 

herbarum [(Pers.: Fr.) Link] f. dianthi, in herb. 

On leaves of Dianthus caryophyllus (Caryophyllaceae), Italy, Treviso, Selva, Aug. 1903 (BPI 

427194, 426981). 

herbarum [(Pers.: Fr.) Link] f. epixylon Sacc., Mycoth. ven., Fasc. III, No. 286 (1876)!, nom. 

nud. 

T: on decorticated branches of Robinia pseudacacia (Fabaceae), Italy, Selva (Treviso), Sept. 

1874, Sacc., Mycoth. ven. 286 (e.g., BPI 427093; HAL). 

Notes: In OUDEMANS (1919: 570) wrongly cited as ‘C. herbarum [Lk. (fa)] var. epixylon 

Sacc., in Thüm., F. austr., No. 891’. However, the only name cited in ‘Thüm., F. austr. 891’ is 

Cladosporium herbarum var. lignicola Thüm. 

herbarum [(Pers.: Fr.) Link] f. fimicola Marchal, Bull. Soc. Roy. Bot. Belgique 24: 67 

(1885)!. 

T: on dung of shrew and mice, Belgium, Brussels, summer 1883 and winter 1883–1884 (in 

fimo soricino et murino). 

Lit.: PRASIL & DE HOOG (1988: 53). 

Notes: no authentic material at BR. 

herbarum [(Pers.: Fr.) Link] f. flosculorum Thüm., F. austr., Cent. XII, No. 1173 (1874)!. 

T: on Anthemis austriaca (Asteraceae), Czech Republic, Teplitz, summer 1873, von Thümen, 

Thüm., F. austr. 1173 (e.g., HAL). 


Notes: see C. herbarum var. flosculorum Thüm. 

herbarum [(Pers.: Fr.) Link] f. fructicola, in herb. 

On fruits of Lonicera etrusca (Caprifoliaceae), Italy, Padua, Aug. 1902 (BPI 427197). 

On fruits of Datura stramonium (Solanaceae), Austria, Lower Austria, Krems, 1871, A. Boller 

(BPI 427198). 

herbarum [(Pers.: Fr.) Link] f. hormodendroides Ferraris → cladosporioides. 

herbarum [(Pers.: Fr.) Link] f. mesembrianthemi – Exsiccatus 767. II specified in KOHL- 

MEYER (1962: 39). 

herbarum [(Pers.: Fr.) Link] f. parasiticum Sacc., Ann. Mycol. 13(2): 133 (1915)!, as ‘parasitica’. 

T: on Exoascus tosquinetii on leaves of Alnus glutinosa (Betulaceae), M. Weisskirchen (PAD: 

holotype). 

Lit: SACCARDO (1931: 797), PRASIL & DE HOOG (1988: 54), HEUCHERT et al. (2005: 53–55). 


52

Notes: PRASIL & DE HOOG (1988) examined authentic material at PAD, which contains a 

mixture of C. herbarum and C. cladosporioides. 

herbarum [(Pers.: Fr.) Link] f. psoraleae Gonz. Frag., Intr. al. est. de la flor. de micr. de Cat.: 

152 (1917). 

T: on Psoralea sp. (Fabaceae), Spain. 

Lit.: GONZÁLES-FRAGOSO (1927: 198). 

herbarum [(Pers.: Fr.) Link] f. repens Fresen., Beitr. Mykol. 1: 24 (1850)!. 

T: (FRESENIUS 1850: Tab. 3, Fig. 29: iconotype). 

Notes: FRESENIUS (1850) described a ‘forma’ repens without indicating any morphological 

differences. A type collection was not mentioned. LINDAU (1907: 801) cited it as a synonym 

of C. herbarum (Pers.: Fr.) Link. 

herbarum [(Pers.: Fr.) Link] f. rubi Gonz. Frag., Mem. Real Acad. Ci. Barcelona, Ser. 3, 

15(17): 458 (32) (1920)!. 

T: on leaves of Rubus rusticanus (= R. ulmifolius) and Rubus sp. (Rosaceae), Spain, 

Barcelona, Bonanova and Vallvidrera, Sept. 1915 and Jul. 1918, Fr. Sennen (MA 06328, 

06329: syntypes). 

= Pseudocercospora rubi (Sacc.) Deighton, Mycol. Pap. 140: 152 (1976)!. 

Lit.: SACCARDO (1931: 795), PRASIL & DE HOOG (1988: 53, as ‘var.’). 

herbarum [(Pers.: Fr.) Link] f. saxicola Sacc., Michelia 2(8): 578 (1882)!. 

T: on wet stones, USA, New Jersey, Newfield, Ellis no. 3551 (PAD). 

Lit.: PRASIL & DE HOOG (1988: 54)!. 

Notes: Original specimen at PAD contains no fungus (PRASIL & DE HOOG 1988). 

herbarum [(Pers.: Fr.) Link] f. sechii-edulis Thüm., Herb. myc. oec., Fasc. X, No. 466 (1877), 

nom. nud. 

T: on Sechium edule (Cucurbitaceae), USA, Thüm., Herb. myc. oec. 466. 

Lit.: STEVENSON (1971). 

herbarum [(Pers.: Fr.) Link] f. stellariae Unamuno, Bol. Soc. Esp. Hist. Nat. 34: 146 (1934)!. 

T: on dead leaves of Stellaria uliginosa (Caryophyllaceae), Spain, León, Cerca de Baña, 

Arroyo, Sierra Baja, 1200 m, 7 Aug. 1933, W. Rothmaler, Iter Hispanicum 583 (MA 06330). 


herbarum [(Pers.: Fr.) Link] var. agaves-echeveriae Savelli, Ann. Reale Accad. Agric. Torino 

56: 113 (1914)! (dated 1913), as ‘f. agaves-echeveriae’. 

Lit.: SACCARDO (1931: 796). 

Notes: see C. herbarum f. agaves-echeveriae. 

herbarum [(Pers.: Fr.) Link] var. aphidicola C. Massal., in Sacc., Madonna Verona 1918: 21 

(1918). 

T: parasitic on aphids, Italy (PAD). 


Notes: No aphids nor Cladosporium conidia could be found in authentic specimen at PAD 

(PRASIL & DE HOOG 1988). 

herbarum [(Pers.: Fr.) Link] var. aphidis Fuckel, Jahrb. Nassauischen Vereins Naturk. 23–24: 

356 ‘1869’ (1870)!, as ‘α aphidis’, nom. nud. 

T: on dead carcass of Aphidina (aphids) on Cornus sanguinea, Germany, near Oestrich. 

Lit.: SACCARDO (1886: 369), LINDAU (1907: 830), PRASIL & DE HOOG (1988: 52). 

Notes: Type material is not preserved in herb. G (PRASIL & DE HOOG 1988). 

herbarum [(Pers.: Fr.) Link] var. brassicae, in herb. 

On leaves of Brassica napus (Brassicaceae), Czech Republic, Bohemia N., Decin (Tetschen), 

spring 1873, de Thümen (BPI 427187). 

53

herbarum [(Pers.: Fr.) Link] var. carpophilum, in herb. 

On Cheiranthus cheiri (= Erysimum cheiri) (Brassicaceae), Italy, Treviso, Selva, Jul. 1899, de 

Thümen (BPI 427188A, 427188B). 

herbarum [(Pers.: Fr.) Link] var. cellulosae Sartory, R. Sartory, J. Mey. & Baumli, Papier 

38(1): 43 (1935). ? (not in this volume, probably not in this journal). 

T: on rotten paper, France. 


Notes: location of material unknown (no material in PC or STR); possible identity with C. 

sphaerospermum. 

herbarum [(Pers.: Fr.) Link] var. cerealium Sacc., in Ferraris, Ann. Mycol. 7: 285 (1909)!. 

T: on leaves, culms etc. of cereals (Triticum, Hordeum, Secale etc.) (Poaceae), Europe (PAD 

?). 

Lit.: LINDAU (1910: 795), FERRARIS (1912: 333), SACCARDO (1913: 1366), PRASIL & DE 

HOOG (1988: 52). 

Notes: The species was described as a substrate form on cereals without any real morphological 

difference (PRASIL & DE HOOG 1988). 

herbarum [(Pers.: Fr.) Link] var. citricola H.S. Fawc. & O.F. Burger, Phytopathology 1: 165 

(1911)!. 

T: on Citrus sp. (Rutaceae), USA, Florida (FLAS). 


Notes: This variety, sufficiently different from C. herbarum, is to be considered as a separate 

species (PRASIL & DE HOOG 1988). 

herbarum [(Pers.: Fr.) Link] var. densum Roum., F. sel. gall. exs., Cent. X, No. 950 (1880). 

T: on leaves of Rhus cotinus (= Cotinus coggygria) (Anacardiaceae), France, Roum., F. sel. 

gall. exs. 950. 


herbarum [(Pers.: Fr.) Link] var. epixylinum Corda 

Lit.: SACCARDO & BERLESE (1884: 100). (No place of publication is provided). 

herbarum [(Pers.: Fr.) Link] var. fasciculare Corda, Icon. fung. 3: 9 (1839)!, as ‘β fasciculare’. 

T: on stems of umbellifers, Lilliaceae and Equisetum sp., Czech Republic. 

Lit.: SACCARDO (1886: 351), LINDAU (1907: 803), FERRARIS (1912: 333), PRASIL & DE 

HOOG (1988: 52). 


Notes: Type material is not preserved in PRM, but from the figures it seems to be 

indistinguishable from C. herbarum (PRASIL & DE HOOG 1988). RABENHORST (1844) listed it 

as synonym of C. fasciculare (Pers.) Fr. 

herbarum [(Pers.: Fr.) Link] var. fimicola Marchal, Bull. Soc. Roy. Bot. Belgique 24(1): 67 

(1885)!, as ‘forma fimicola’. 

Lit.: SACCARDO (1886: 351, as ‘fimicolum’), LINDAU (1907: 803). 

Notes: see C. herbarum f. fimicola. 

herbarum [(Pers.: Fr.) Link] var. flosculorum, in herb. 

On florets of Anthemis austriaca (Asteraceae), Czech Republic, Bohemia N., Teplice, summer 

1873, von Thümen (BPI 427196). 

herbarum [(Pers.: Fr.) Link] var. hypharum (sic) Westend. & Van Haes., LINDAU (1910 – in 

index). 

An error. Refers to C. herbarum var. typharum Westend. & Van Haes., in LINDAU (1907: 

801). 

herbarum [(Pers.: Fr.) Link] var. indutum Thüm., Mycoth. univ., Cent. XVI, No. 1571 

(1880)!. 

54

T: on dry stems of Zea mays (Poaceae), USA, South Carolina, Aiken, 1877, H.W. Ravenel, 

Thüm., Mycoth. univ. 1571 (e.g., HAL). 

herbarum [(Pers.: Fr.) Link] var. lablab Sacc., Philipp. J. Sci. 18: 604 (1921). 

T: on legumes of Dolichos lablab (Fabaceae), China, Foochow (PAD). 


Notes: Authentic material at PAD contains C. herbarum, C. cladosporioides, Alternaria 

alternata and Penicillium-like conidia (PRASIL & DE HOOG 1988). 

herbarum [(Pers.: Fr.) Link] var. lignicola Thüm., F. austr., Fasc. 9, No. 891 (1874)!, nom. 

nud. 

T: on wood of Pinus sp. (Pinaceae), Czech Republic, Bohemia, Libouchec (Königswald), 

summer 1873, von Thümen, Thüm., F. austr. 891 (e.g., BPI 427199; HAL). 

herbarum [(Pers.: Fr.) Link] var. macrocarpum (Preuss) M.H.M. Ho & Dugan, in Ho, 

Castañeda, Dugan & Jong, Mycotaxon 72: 131 (1999)!, as ‘Cladosporium herbarum (Pers.) 

Link var. macrocarpum M.H.M. Ho & F.M. Dugan, comb. nov.’. 

T: on dead leaves of Eryngium pandanifolium (Apiaceae), Italy, Saccardo (Herb. mycol.) 419 

(PAD: neotype), selected by DE VRIES, 1952, as ‘lectotype’. 

≡ Cladosporium macrocarpum Preuss, in Sturm, Deutschl. Fl. 3(6): 27 (1848)!. 

Lit.: MCKEMY & MORGAN-JONES (1991c), DAVID (1997: 71). 

Ill.: HO et al. (1999: 130, Fig. 23). 

Notes: see C. macrocarpum Preuss. 

herbarum [(Pers.: Fr.) Link] var. macrosporum Lagière, Ann. École Natl. Agric. Grignon, 

Sér. 3, 5: 159 (1945–1946)!. 

T: on leaves of Phleum pratense (Poaceae), France, Seine-et-Oise, Grignon, 1944, Lagière. 

= ? Cladosporium phlei (C.T. Greg.) G.A. de Vries, Contr. Knowl. Genus Cladosporium: 49 

(1952)!. 


Notes: Original material seems to be lost. PRASIL & DE HOOG (1988) consider it to be 

probably a Heterosporium species. 

herbarum [(Pers.: Fr.) Link] var. nigricans (Roth: Fr.) Fr., Syst. mycol. 3(2): 371 (1832)!, as 

‘β nigricans’. 

T: on wood and other hard substrates. 

≡ Byssus nigricans Roth, Catal. Bot. 1: 216 (1797). 

= ? Dematium hippocastani Pers., Ann. Bot. (Usteri) 11: 32 (1794)!; Syn. meth. fung. 2: 698 

(1801)!. 

Lit.: SACCARDO (1886: 351, 1931: 798), LINDAU (1907: 803), ARENS (1945), PRASIL & DE 

HOOG (1988: 53). 

Notes: Material could not be traced neither in herb. B nor PRM (PRASIL & DE HOOG 1988). 

herbarum [(Pers.: Fr.) Link] var. phaseoli, in herb. 

On leaves of Phaseolus vulgaris (Fabaceae), Czech Republic, Bohemia N., near Decin 

(Tetschen), autumn 1873, von Thümen (BPI 427201). 

herbarum [(Pers.: Fr.) Link] var. phlei Lagière, Ann. École Natl. Agric. Grignon, Sér. 3, 5: 

159 (1945–1946)!. 

T: on leaves of Phleum pratense (Poaceae), France, Seine-et-Oise, Grignon, summer 1944, R. 

Lagière. 


Notes: PRASIL & DE HOOG (1988): Original material is probably lost, possible identity with C. 

macrocarpum.’ 

herbarum [(Pers.: Fr.) Link] var. rhois, in herb. 

On dead branches of Rhus typhina (Anacardiaceae), Austria, Lower Austria, Krems, winter 

1870, A. Boller (BPI 427203). 

55

herbarum [(Pers.: Fr.) Link] var. rubi Gonz. Frag., Mem. Real Acad. Ci. Barcelona, Ser. 3, 

15(17): 458 (32) (1920)!. 

Notes: see C. herbarum f. rubi Gonz. Frag. 

herbarum [(Pers.: Fr.) Link] var. saxicola Sacc., Michelia 2(8): 578 (1882)!, as ‘forma’ 

saxicola. 

Lit.: SACCARDO (1886: 351). 

Notes: see C. herbarum f. saxicola. 

herbarum [(Pers.: Fr.) Link] var. solutum (Link) Sacc. → solutum. 

herbarum [(Pers.: Fr.) Link] var. torulosum Berk. & Broome, The Fungi of Ceylon, no. 886 

(1870); J. Linn. Soc., Bot. 14: 99 ‘1873’ (1875)!. 

T: on leaves of Anamirta (Menispermaceae), India, Ceylon (K). 

Lit.: SACCARDO (1886: 351), PETCH (1927), PRASIL & DE HOOG (1988: 53). 

Notes: PRASIL & DE HOOG (1988) considered this variety to be identical with C. herbarum. 

herbarum [(Pers.: Fr.) Link] var. typharum Westend., Cryptogames: 26 (1854), nom. inval. 


Notes: see entry below. 

herbarum [(Pers.: Fr.) Link] var. typharum Westend. & Van Haes., Cat. crypt.: 8 (1838), no. 

173. 

Notes: cited in LINDAU (1907: 801) as synonym of C. herbarum. 

herbarum [(Pers.: Fr.) Link] var. vincetoxici Allesch., in P. Sydow, Hedwigia 36(6), Beibl.: 

163 (1897)!. 

T: on dead stems of Vincetoxicum purpurascens (Asclepiadaceae), Germany, near Berlin (M). 

Lit.: SACCARDO (1899: 1080), LINDAU (1907: 803), PRASIL & DE HOOG (1988: 53). 

Notes: indistinguishable from C. herbarum (PRASIL & DE HOOG 1988). 

herbarum [(Pers.: Fr.) Link] var. vitricola Sacc., in Ferraris, Ann. Mycol. 7: 286 (1909)!. 

T: on dirty, wet glass, Italy (PAD?). 

Lit.: LINDAU (1910: 795), FERRARIS (1912: 334), SACCARDO (1913: 1366), PRASIL & DE 

HOOG (1988: 53). 

Notes: “Byssocladium fenestrale auct. p.p. non Mart. nec Link” (SACCARDO 1913: 1366). 

heteronemum (Desm.) Oudem., Arch. Néerl. Sci. Exact. Nat. 11: 363 (1876)!. 

T: on dead leaves of Sagittaria sagittifolia (Alismataceae), France, Desm., Pl. crypt. N. 

France, Sér. 2, No. 7. 

≡ Macrosporium heteronemum Desm., Ann. Sci. Nat. Bot., Sér. 3, 20: 216 (1853)!. 

= ? Ascospora solidaginis Fr., Summa veg. Scand. 2: 425 (1849)!. 

Notes: Synonomy and exsiccati in OUDEMANS (1923). 

heterophragmatis S.A. Khan & Kamal, Mycopathol. Mycol. Appl. 18(4): 246 (1962)!. 

T: on leaves of Heterophragma adenophyllum (Bignoniaceae), Pakistan, Tandoja, Campus 

A.R.I., 15 Nov. 1961, Shakil Ahmad Khan (IMI 90787: holotype). 

Lit.: SCHUBERT & BRAUN (2004: 298–300). 

Ill.: KHAN & KAMAL (1962: 247, Fig.), SCHUBERT & BRAUN (2004: 299, Fig. 1). 

Host(s)/substrate(s) & distribution: on Heterophragma adenophyllum; Asia (Pakistan). 

heterosporium, in herb. 

On an unidentified host plant, USA, New York, Buffalo, G.W. Clinton (BPI 427204). 

Notes: Excluded, conidiophores with tretic conidiogenous loci. Taxonomic affinity remains 

unclear. 

heuglinianum Thüm., Rev. Mycol. (Toulouse) 1: 11 (1879)!. 

T: on leaves of Buddleja polstachya (Buddlejaceae), near Nakfa on Red Sea (Nakfa ad Mare 

Rubrum). 

Lit.: SACCARDO (1892: 603). 

Notes: Type material could not be located. 

56

hibisci Reichert, Bot. Jahrb. Syst. 56: 721 (1921)!. 

T: on dry stems of Hibiscus esculentus (Malvaceae), Egypt, near Siut, Oct. 1822/25, 

Ehrenberg (B). 

Lit.: SACCARDO (1972: 1338). 


Notes: ZHANG et al. (2003) cited collections on Hibiscus syriacus (China, Shaanxi) and H. 

tiliaceus (China, Heilongjiang). 

hordei (Bruhne) Pidopl. → bruhnei. 

hordei Pass., in Brunaud, Fragments Mycologiques 1884–1885: 32 (1887). 

T: on faded leaves of Hordeum distichum (= distichon) (Poaceae), France, Saintes. 

Lit.: SACCARDO (1892: 605). 

Notes: Type at B missing (Burghard Hein, personal communication). 

hoveae Syd. & P. Syd., Ann. Mycol. 15: 148 (1917)!. 

T: on leaves of Hovea longifolia var. pannosa (Fabaceae), Australia, Brisbane River, 1863– 

1865, A. Dietrich, comm. J. Bornmüller (S: holotype). 

≡ Pseudocercospora hoveae (Syd. & P. Syd.) K. Schub. & U. Braun, Mycol. Progr. 4(2): 106 

(2005)!. 

Lit.: SACCARDO (1931: 792). 


humile Davis, Trans. Wisconsin Acad. Sci. 19: 702 (1919)!. 

T: on leaves of Acer rubrum (Aceraceae), USA, Wisconsin, Luck, 25 Aug. 1916, J.J. Davis 

(WIS: lectotype; BPI 427214: isolectotype). 

≡ Fusicladium humile (Davis) K. Schub. & U. Braun, IMI Descriptions of Fungi and Bacteria 

152, No. 1520 (2002)!. 

≡ Fusicladosporium humile (Davis) Partridge & Morgan-Jones, Mycotaxon 85: 366 (2003)!. 

Teleomorph: Venturia acerina Plakidas ex M.E. Barr, Canad. J. Bot. 46: 814 (1968)!. 

Lit.: SACCARDO (1931: 788), ELLIS (1976: 340), SIVANESAN (1984: 607, as Cladosporium 

state of Venturia acerina), SCHUBERT et al. (2003: 57). 

hydrangeae Z.Y. Zhang & T.F. Li, in Zhang, Li, Zhang & Wang, J. Anhui Agric. Univ. 26: 

40 (1999)!. 

T: on living leaves of Hydrangea macrophylla (Hydrangeaceae), China, Liaoning, 16 Sept. 

1992, Wang & Li (MHYAU 07029: holotype). 

Ill.: ZHANG, LI, ZHANG & WANG (1999: 41, Fig. 1), ZHANG et al. (2003: 110, Fig. 70). 

Notes: The article is published in a supplement to volume 26. Further collections on 

Hydrangea macrophylla are reported from China, Liaoning and Yunnan (ZHANG et al. 2003). 

hypophloeum Berk. & M.A.Curtis, in Berkeley, J. Linn. Soc., Bot. 10: 362 (1869)!. 

T: on leaves of a Sapindaceae, Cuba, February. 

Lit.: SACCARDO (1886: 359). 

hypophyllum Fuckel, Jahrb. Nassauischen Vereins Naturk. 23–24: 356 ‘1869’ (1870)!. 

T: on the lower surface of living leaves of Ulmus campestris (= U. minor) (Ulmaceae), 

Germany, ‘auf Grünau bei Hattenheim am Rheinufer’, Fuckel, F. rhen. 1629 (e.g., BPI 

427228, 427229, 427234; HAL; M-57614: syntypes). 

= Cladosporium microstictum Sacc. & D. Sacc., Ann. Mycol. 3: 169 (1905)!, syn. nov. 


Host(s)/substrate(s) & distribution: on Ulmus minor; Europe (Germany, Italy). 

Notes: This species was discussed by DE VRIES (1952) to be a synonym of Cladosporium 

cladosporioides. A re-examination of type material showed that both species are quite distinct. 

hyridis – BPI 427627 TYPE 

[probably misspelling of ‘xyridis’, i.e., Cladosporium xyridis Tracy & Earl; Xyris fimbriata 

(Xyridaceae), USA, Mississippi, 29 Sept. 1895, F.S. Earle]. 

57

idesiae Bres., in Sydow, Hedwigia 35(1), Beih.: 62 (1896)!. 

T: on leaves of Idesia sp. (Flacourtiaceae), Germany, Berlin, Späth’sche Baumschule, Sydow, 

Mycoth. march. 4498 (B 70-6556). 

= Cladosporium herbarum [(Pers.: Fr.) Link] var. macrocarpum (Preuss) M.H.M. Ho & 

Dugan in Ho, Castañeda, Dugan & Jong, Mycotaxon 72: 131 (1999)!. 


inaequiseptatum Matsush., Icones Microfungorum a Matsushima Lectorum: 35 (1975)!. 

T: on a dead leaf of Quercus phillyraeoides (Fagaceae), Japan, Miyajima, Hiroshima, May 

1972 (Matsush. Herb. 4428). 

≡ Parapleurotheciopsis inaequiseptata (Matsush.) P.M. Kirk, Trans. Brit. Mycol. Soc. 78(1): 

65 (1982)!. 

inconspicuum Thüm., Contributiones ad florum mycologicam lusitanicam, Ser. 2, No. 193 

(1879) and Hedwigia 19: 133 (1880)!. 

T: on living leaves of Styrax officinalis (Styracaceae), Portugal, Coimbra, botanical garden, 

Oct. 1878, G.A. Moller (B 70-6557). 

Lit.: SACCARDO (1886: 359), OUDEMANS (1923). 

Host(s)/substrate(s) & distribution: on Styrax officinalis; Europe (Portugal). 

indicum J.N. Rai, J.P. Tewari & Mukerji, Mycopathol. Mycol. Appl. 38: 22 (1969)!. 

T: from mangrove mud, India, West Bengal, via Lucknow University, 26 Mar. 1962, J.N. Rai 

(IMI 92675, IMI 92676). 

Ill.: RAI et al. (1969: 25, Figs 12–16). 

indigoferae Sawada, Special Publ. Coll. Agric. Natl. Taiwan Univ. 8: 196 (1959)!, nom. inval. 

T: on stems of Indigofera tinctoria (Fabaceae), Taiwan, Pref. Taipei, Taipei, 18 Aug. 1909, K. 

Sawada (PPMH; BPI 427230). 


infuscans Thüm., Rev. Mycol. (Toulouse) 1: 59 (1879)!. 

T: on living stems of Desmodium strictum (Fabaceae), USA, South Carolina, Aiken, H.W. 

Ravenel, Thüm., Mycoth. univ. 1573 (e.g., BPI 427232, 427231; HAL; M; NY: syntypes). 

≡ Dendryphiella infuscans (Thüm.) M.B. Ellis, Dematiaceous Hyphomycetes: 500 (1971)!. 

Lit.: SACCARDO (1886: 361). 

inopinum (Petr.) U. Braun, Mycotaxon 55: 224 (1995)!. 

T: on leaves of Gynoxys hallii (Asteraceae), Ecuador, Pichincha, near Quito, 20 Sept. 1937, H. 

Sydow, Reliquiae Petrakianae 1350 (GZU: syntype). 

≡ Cercospora inopina Petr., Sydowia 4: 570 (1950)!. 

Lit.: CROUS & BRAUN (2003: 227). 

Ill.: BRAUN (1995: 225, Fig. 2). 

Host(s)/substrate(s) & distribution: on Gynoxys hallii; South America (Ecuador). 

insectorum Gonz. Frag., Broteria, Sér. Bot. 22: 69 (1926)!, as ‘insectarum’. 

T: on Saissetia hemisphaerica (Homoptera, Coccidae) on leaves of Asplenium lineatum 

(Aspleniaceae) and Cordyline congesta (Dracaenaceae), Spain, Madrid, botanical garden, Mar. 

1925, Menor. 

Lit.: SACCARDO (1972: 1338). 

Notes: Type collections could not be located in herb. MA. 

iridicola Schwein., Trans. Amer. Philos. Soc., N.S., 4(2): 277 (1832)!. 

T: on leaves of Iris virginica (Iridaceae), USA, Pennsylvania, Bethlehem, No. 2604. 

Lit.: SACCARDO (1886: 367, as ‘iridicolum’), ZHANG, LI, ZHANG & WANG (1999: 41−42). 

Ill.: ZHANG, LI, ZHANG & WANG (1999: 41, Fig. 2). 

Notes: Type material could not be located in herb. PH, it is probably not preserved. A record 

of this species from China is reported by ZHANG, LI, ZHANG & WANG et al. (1999). 

iridis (Fautrey & Roum.) G.A. de Vries, Contr. Knowl. Genus Cladosporium: 49 (1952)!. 

T: on leaves of Iris germanica (Iridaceae), France, Cote d’Or, Jardin de Noidan, Jul. 1880, F. 

58

Fautrey, Roum., F. sel. gall. exs. 5689 (PC: lectotype, selected by DAVID, 1997; K: isolectotype). 

≡ Scolecotrichum iridis Fautrey & Roum., Rev. Mycol. (Toulouse) 13: 82 (1891). 

≡ Heterosporium iridis (Fautrey & Roum.) J.E. Jacques, Contr. Inst. Bot. Univ. Montréal 39: 

18 (1941). 

= Heterosporium gracile Sacc., Syll. fung. 4: 480 (1886)!, as ‘(Wallr. ?) Sacc.’. [T: PAD: 

lectotype]. 

= Heterosporium montenegrinum Bubák, Sber. K. böhm. Ges. Wiss. Math.-Nat. Kl. 1903(12): 

21 (1903). [T: BPI: lectotype]. 

= Scolecotrichum cladosporioideum Maire, Ann. Mycol. 4: 329 (1906)!, as ‘sp. prov. nov.’. 

[T: MPU: holotype; PC, K: isotypes]. 

= ? Heterosporium pruneti Nicolas & Aggéry, Rev. Pathol. Vég. Entomol. Agric. France 15: 

66 (1928)!. 

= Heterosporium iridis-pumilae Săvul. & Sandu, Hedwigia 75: 222 (1935)!. [T: K, IMI 

10047: isotypes]. 

Teleomorph: Davidiella macrospora (Kleb.) Crous & U. Braun, in Braun, Crous, Dugan, 


Lit.: ELLIS (1971: 312), ELLIS & WALLER (1974), SIVANESAN (1984: 222, as Cladosporium 

state of Mycosphaerella macrospora), MCKEMY & MORGAN-JONES (1990), DAVID (1997: 

43), SHIN et al. (1999). 

Ill.: ELLIS (1971: 311, Fig. 215 B), SIVANESAN (1984: 224, Fig. 120), MCKEMY & MORGAN- 

JONES (1990: 427, Fig. 1; 429, Pl. 1; 431, Fig. 2; 433, Fig. 3; 437, Pl. 2), DAVID (1997: 45, 

Fig. 11), SHIN et al. (1999: Fig. 1). 

Host(s)/substrate(s) & distribution: on Iris spp., including I. florentina, I. foetidissima, I. 

germanica, I. gueldenstaedtiana, I. pallida, I. plicata (= I. swertii Hort.), I. pseudacorus and 

other species, also on Belacamanda chinensis (= Gemmingia chinensis); Europe, Asia, Africa, 

North America, Central & South America, Australia. 

jacarandae Viégas, Bragantia 7(2): 33 (1947)!. 

T: on living leaves of Jacaranda sp. (Bignoniaceae), Brazil, Minas Gerais, Agua Limpa, Exp. 

de Agua Limpa, 21 May 1945, E.P. Heringer (IACM: holotype). 

≡ Fusicladium jacarandae (Viégas) K. Schub., U. Braun & F. Freire, in Schubert & Braun, 

Sydowia 56(2): 302 (2004). 

Ill.: VIÉGAS (1947: 46, Fig. 11), SCHUBERT & BRAUN (2004: 303, Fig. 3). 

jacarandicola K. Schub., U. Braun & C.F. Hill, in Schubert & Braun, Sydowia 56(2): 300– 

301(2004). 

T: on living leaves of Jacaranda mimosifolia (Bignoniaceae), New Zealand, Mt. Albert, 

Ruarangi Road, 6 Aug. 2002, C.F. Hill 693 [HAL 1812 (F): holotype]. 

Ill.: SCHUBERT & BRAUN (2004: 300, Fig. 2). 

Host(s)/substrate(s) & distribution: on Jacaranda mimosifolia; New Zealand. 

jasmini Schwein. → maculans Schwein. 

javanicum Wakker, Meded. Proefstat. Oost-Java, N.S., 28: 1–9 (1896)!. 

T: on cortex of (in corticibus) Saccharum officinarum (Poaceae), Java. 

Lit.: SACCARDO (1899: 1082). 

Ill.: WAKKER (1896: Tab. 1). 

juglandinum Cooke, Grevillea 16(79): 80 (1888)!. 

T: on fading leaves of walnut (Juglans regia, Juglandaceae), Great Britain, Highgate (K). 


juglandis Pass., in herb. 

On leaves of Juglans regia (Juglandaceae), Italy (B). 

Notes: on the label: “a Cladosporium epiphyllo Nees”. 

59

juncicolum Rabenh. / P. Magnus, in herb. 

On Juncus acutus? (Cyperaceae), Egypt, P. Ascherson, iter aegyptiacum quartum, ex herb. P. 

Magnus (HBG). 

Notes: on the label “viell. Brachysporium juncicolum Rabenh.; Helminthosporium juncicolum 

Rabenh.“. SACCARDO (1886: 430): “Brachysporium juncicolum (Rabenh.) Sacc. Helminthosporium 

juncicolum Rabenh., Bot. Zeit. 1851, p. 626 – An Cladosporium ?“. 

kapildharens C.D. Sharma, Gadp., Firdousi, A.N. Rai & K.M. Vyas, Indian Phytopathol. 

51(2): 160 (1998)!. 

T: on living leaves of Hospitis ignoti, India, Madhya Pradesh, Shahdol circle, Amarkantak 

(south forest division), Kapil dhara, Jan. 1993, C.D. Sharma (S.U. Herb. No. C.S.2: holotype; 

IMI 356766: isotype). 


kniphofiae Cooke, Grevillea 14(70): 40 (1885)!. 

T: on dead leaves of Kniphofia aloides (Asphodelaceae), Great Britain, Surrey, Kew, Royal 

Botanic Gardens, Aug. 1885, M.C. Cooke (K 121560: holotype). 

= Cladosporium herbarum s. lat. 

Lit.: SACCARDO (1886: 367). 

lacroixii Desm., Pl. crypt. N. France, Ed. 3, Fasc. 15–16, No. 755 (1860). 

T: on Narcissus jonquilla (Amaryllidaceae), France, Vienne, L.-S.V. de Lacroix, Desm., Pl. 

crypt. N. France 755 (PC: lectotype, selected by DAVID, 1997; BR, K, IMI 118411: isolectotypes). 

Lit.: SACCARDO (1886: 370), ELLIS (1976: 333), DAVID (1997: 50). 

Ill.: ELLIS (1976: 335, Fig. 253), DAVID (1997: 51, Fig. 13; 53, Fig. 14 G–I). 

Host(s)/substrate(s) & distribution: on Narcissus jonquilla; Europe (France). 

Notes: OUDEMANS (1919) listed Allium sphaerocephalum and Ornithogalum umbellatum as 

hosts, but these records seem to be based on misidentifications referring to Cladosporium allii 

and Cladosporium ornithogali (Klotzsch ex Cooke) G.A. de Vries. 

lactucae Sawada, Rep. Gov. Res. Inst. Formosa 85: 92 (1943)!, nom. inval. 

T: on Lactuca indica (Asteraceae), Taiwan, Taipeh, 9 Mar. 1924, K. Sawada (BPI 427238; 

PPMH). 

= Cladosporium lactucicola Y. Cui & Z.Y. Zhang, in He & Zhang, Mycosystema 21(1): 22 

(2002). 

≡ Cladosporium lactucicola Z.Y. Zhang & Y. Cui, in Zhang et al., Flora Fungorum 

Sinicorum, Vol. 14: 114 (2003)!, nom. superfl. 

≡ Passalora lactucicola (Y. Cui & Z.Y. Zhang) K. Schub. & U. Braun, Mycol. Progr. 4(2): 

105 (2005)!. 


lactucicola Y. Cui & Z.Y. Zhang, in He & Zhang, Mycosystema 21(1): 22 (2002)!. 

T: on living leaves of Lactuca indica (Asteraceae), China, Sichuan Prov., Chengdu, 15 Aug. 

1985, Y. Qing (MHYAU 03881: holotype). 

≡ Cladosporium lactucicola Z.Y. Zhang & Y. Cui, in Zhang et al., Flora Fungorum Sinicorum, 

Vol. 14: 114 (2003)!, nom. superfl. 

≡ Passalora lactucicola (Y. Cui & Z.Y. Zhang) K. Schub. & U. Braun, Mycol. Progr. 4(2): 

105 (2005). 

= Cladosporium lactucae Sawada, Rep. Gov. Res. Inst. Formosa 85: 92 (1943)!, nom. inval. 

Ill.: HE & ZHANG (2002: 22, Fig. 2), ZHANG et al. (2003: 114, Fig. 73), SCHUBERT & BRAUN 

(2005a: 105, Fig. 4). 

ladinum E. Müll., Sydowia 4: 294 (1950)!. 

T: on dead stems of Laserpitium halleri (Apiaceae), Switzerland, Graubünden, Scuol, 17 Jul. 

1948, E. Müller (ZT). 

Teleomorph:? Leptosphaeria ladina E. Müll., Sydowia 4: 293 (1950)!. 

≡ Nodulosphaeria ladina (E. Müll.) L. Holm, Symb. Bot. Upsal. 14(3): 83 (1957)!. 

60

Notes: ‘It must be considered a culture contaminent rather than an anamorph’ (CRANE & 

SHEARER 1991). 

lanciforme Ces., Flora 36: 204 (1853)!. 

T: on Typha sp. (Typhaceae), Italy, mixed infection with Cladosporium astroideum Ces. 

≡ Cladosporium typharum [Desm.] f. lanciforme (Ces.) Ferraris, Flora Ital. Crypt., Pars I, 

Fungi, Fasc. 8: 337 (1912)!. 

Lit.: SACCARDO (1886: 366), LINDAU [1907: 813, indexed as ‘laniforme Ces.’ in LINDAU 

(1910)]. 

langeronii (Fonseca, Leão & Nogueira) Vuill., Champ. paras.: 78 (1931)!. 

T: isolated from an ‘ulcero-nodular mycosis’. 

≡ Hormodendron langeronii Fonseca, Leão & Nogueira, Sci. Med. 5: 563 (1927). 

≡ Cladosporium langeronii (Fonseca, Leão & Nogueira) Cif., Manuale di Micologia Medica, 

ed. 2: 488 (1960)!, as ‘langeroni’, comb. superfl. 

= Cladosporium sphaerospermum Penz., in Sacc., Michelia 2(8): 473 (1882)!. 

Lit.: DODGE (1935: 847–848), DE HOOG et al. (2000: 592). 

lantanae K. Bhalla & A.K. Sarbhoy, Indian Phytopathol. 53(3): 263 (2000)!. 

T: on living leaves of Lantana camara (Verbenaceae), Cuba, Bayamo, Mar. 1967, R. Urtiaga 

(IMI 126781b). 

≡ Mycovellosiella lantanae [Chupp] var. cubensis Deighton, Mycol. Pap. 137: 36 (1974)!. 

≡ Passalora lantanae var. cubensis (Deighton) U. Braun & Crous, in Crous & Braun, Mycosphaerella 


Biodiversity Ser. 1: 243 (2003)!. 

laricis Sacc., Ann. Mycol. 3: 515 (1905)!. 

T: on still living leaves of Larix europaea (= Larix decidua) (Pinaceae), Italy, Apenninen, 

Giogo di Scarparia (Mugello), Sept. 1905, Prof. V. Perona (PAD: holotype). 



Lit.: LINDAU (1907: 812), FERRARIS (1912: 336; 1914: 883), SACCARDO (1913: 1369, 1931: 

790). 

Notes: See Saccardo’s comments on relation to Meria (1931: 1370). 

laricis [Sacc.] var. pini-pineae Sacc. & Trotter, I Funghi dell’Avellinese, Avellino: 154 

(1920). 

T: on sheaths of living leaves or leaves becoming dry (In vaginis adhuc vivis vel arescentibus 

foliorum) of Pinus pinea (Pinaceae), Italy, Campania, near Avellino, mixed infection with 

Pestalozzia hartigi. 

Lit.: SACCARDO (1931: 790). 

lathyri Z.Y. Zhang & Y.L. Liu, J. Yunnan Agric. Univ. 15(3): 219–221 (2000)!. 

T: on leaves and stems of Lathyrus quinquenervius (Fabaceae), China, Liaoning, Shenyang, 20 

Oct. 1992, Y.X. Wang & H. Li (MHYAU 07835: holotype). 

Ill.: ZHANG & LIU (2000: 219, Fig. 1), ZHANG et al. (2003: 115, Fig. 74). 

lauri Raybaud, Congr. Pathol. Végét. (Cent. Pasteur), Strausbourg: 48 (1923)!. 

T: on leaves of Laurus nobilis (Lauraceae), on Aonidia lauri (Hemiptera, Diaspididae) and 

Lecanium hesperidum, France. 

laxicapitulatum Matsush., Icones Microfungorum a Matsushima Lectorum: 35 (1975)!. 

T: on a rotting leaf of Pasania edulis (Fagaceae), Japan, Hyogo, Kobe City, Apr. 1969 

(Matsush. herb. 2570). 

Lit.: HO et al. (1999: 131). 

Ill.: MATSUSHIMA (1975: Pl. 42, Figs 1–2), HO et al. (1999: 133, Figs 24–25). 

laxum Kalchbr. & Cooke, Grevillea 9(49): 24 (1880)!. 

T: on fading leaves of Printzia pyrifolia (Asteraceae), South Africa (IMI 115272; K). 

61

≡ Mycovellosiella laxa (Kalchbr. & Cooke) Deighton, Mycol. Pap. 137: 65 (1974)!. 

≡ Passalora laxa (Kalchbr. & Cooke) U. Braun & Crous, in Crous & Braun, Mycosphaerella 

and its anamorphs: 1. Names published in Cercospora and Passalora, CBS Biodiversity 

Ser. 1: 458 (2003)!. 

Lit.: SACCARDO (1886: 358). 

leproides (L. Léger & Nogueira) Nann., Trattato di Micopatologia Umana 4: 409 (1934)!, as 

‘(Léger & Nogueira) Thom 1930’. 

T: isolated from two patients with mild lesions resembling leprosy. 

≡ Scopulariopsis leproides L. Léger & Nogueira, Bull. Soc. Pathol. Exot. 15: 656 (1922). 

≡ Hormodendrum leproides (L. Léger & Nogueira) C.W. Dodge, Med. Mycol.: 848 (1935)!. 

Notes: “Certainly not a Scopulariopsis” (Mycol. Pap. 86: 87, 1963). 

leprosum Morgan-Jones, Mycotaxon 6(1): 1 (1977)!. 

T: on painted surface of veneer, USA, Alabama, Lee Co., Auburn, Apr. 1976, G. Morgan- 

Jones (BPI: holotype; AUA). 

Ill.: MORGAN-JONES (1977: 2, Fig. 1). 

lethiferum House, Bull. New York State Mus. Nat. Hist. 219/220: 62 (1919–1920)!. 

Notes: Petrak (Petrak's Lists 1: 192) seems to have errored. C. lethiferum Peck is the published 

name. See comments under Cladosporium brevipes House. 

lethiferum Peck, Rep. (Annual) New York State Mus. Nat. Hist. 40: 64 (1887)!. 

T: on living leaves of Populus tremuloides (Salicaceae), USA, New Hampshire, Keene, Jun. 

1887, Peck (NYS: holotype; BPI 427241: isotype). 

≡ Pollaccia lethifera (Peck) M. Morelet, Bull. Soc. Sci. Nat. Archéol. Toulon & Var 34(219): 

12 (1978)!. 

≡ Pollaccia radiosa [(Lib.) E. Bald. & Cif.] var. lethifera (Peck) M. Morelet, Cryptog. Mycol. 

6: 113 (1985)!. 

≡ Fusicladium radiosum [(Lib.) Lind] var. lethiferum (Peck) Ritschel & U. Braun, in 

Schubert, Ritschel & Braun, Schlechtendalia 9: 87 (2003)!. 

= Clasterosporium populi Ellis & Everh., J. Mycol. 7: 134 (1892), nom. illeg., homonym, non 

C. populi Sacc., 1886. [T: NY]. 

≡ Stigmina populi Pound & Clem., Bull. Geol. Nat. Hist. Surv. 9: 662 (1896), as ‘(Ellis & 

Everh.) Pound & Clem.’. 

≡ Stigmina populi Peck, Bull. New York State Mus. Nat. Hist. 157: 34 (1912), as ‘(Ellis & 

Everh.) Peck’. 

= Dicoccum populinum Ellis & Everh., Proc. Acad. Nat. Sci. Philadelphia 1893: 462 (1894). 

[T: NY]. 

= Fusicladium lageniforme Solheim & Hadfield, in Hadfield, Univ. Wyoming Pupl. (1946): 

18–19 (1946), nom. inval. 

= Pollaccia americana Ondřej, Eur. J. Forest Pathol. 2: 144 (1972). [T: DAOM, as 

Fusicladium radiosum (Lib.) Lind]. 

Teleomorph: Venturia tremulae [Aderh.] var. grandidentatae M. Morelet, Cryptog. Mycol. 6: 

113 (1985). 

Lit.: SACCARDO (1892: 604). 

levieri (Magnus) Hara, Agric. & Hort. 12: 2706 (1937)!. 

T: on Diospyros lotus (Ebenaceae), Caucasus, Georgia, Batum, ‘in silvis litoris Euscini’, 16 

Jun. 1890 (HBG: holotype). 

≡ Fusicladium levieri Magnus, in Sommier & Lévier, Trudy Imp. S.-Peterburgsk. Bot. Sada 

16: 543 (1900)!. 

≡ Ragnhildiana levieri (Magnus) Vassiljevsky, in Vassiljevsky & Karakulin, Parazitnye nesovershennye 

griby, Ch. 1., Gifomicety: 373 (1937)!. 

≡ Phaeoramularia levieri (Magnus) U. Braun, in Braun & Melnik, Proc. Komarov Bot. Inst. 

(St. Petersburg) 20: 69 (1997)!. 

= Fusicladium kaki Hori & Yoshino, Bot. Mag. (Tokyo) 19: 220 (1905). 

62

= Fusicladium diospyros Chona, Munjal & J.N. Kapoor, Indian Phytopathol. 9: 129 (1956). 

[T: HCIO?]. 

= Fusicladium diospyros Hori & Yoshino, in herb. (B). 

Lit.: SCHUBERT (2001), CROUS & BRAUN (2003: 482), SCHUBERT et al. (2003: 61–62), 

SCHOLLER et al. (2004: 132–134). 

lichenicola Linds., Quart. J. Microscop. Sci., N.S., 11: 42 (1871)!, as ‘lichenicolum’, nom. 

inval. 

T: on thallus of Peltigera aphthosa (Peltigeraceae), Great Britain, Scotland, S. Aberdeenshire, 

Falls of the Garrawalt, Aug. 1856, W.L. Lindsay. 

Lit.: HAWKSWORTH (1979: 269), HEUCHERT et al. (2005: 55). 

Notes: “...refers only to sterile mycelium. Lindsay was hesitant in introducing the name as he 

stated that the fungus ‘... if it is entitled to specific distinction, may be fitly denominated C. 

lichenicolum’.” (HAWKSWORTH 1979). 

lichenum Keissl., Centralbl. Bakteriol., 2. Abth., 37: 389 (1913)!. 

T: on apothecia of Haematomma cismonicum (Haematommataceae), Austria, Steiermark, valle 

See-Aü at Leopoldsteiner See near Eisenerz, alt. 700 m, Jul. 1912, K. von Keissler 

(W1912/117: holotype). 

≡ Pseudocercospora lichenum (Keissl.) D. Hawksw., Bull. Brit. Mus. (Nat. Hist.), Bot. 6(3): 

246 (1979)!. 

Lit.: SACCARDO (1931: 796), HEUCHERT et al. (2005: 69). 

Ill.: HAWKSWORTH (1979: 247, Fig. 31). 

Notes: Excluded. 

lignatile Schwein., Trans. Amer. Philos. Soc., N.S., 4(2): 277 (1832)!. 

T: on rotten wood, USA, Pennsylvania, Bethlehem, No. 2601 (PH 01020417, as ‘lignatilis’). 

≡ Virgaria lignatilis (Schwein.) S. Hughes, Canad. J. Bot. 31: 603 (1953)!. 

= Botrytis nigra Link, Ges. Naturf. Freunde Berlin Mag. Neuesten Entdeck. Gesammten 

Naturk. 3: 14 (1809). 

≡ Virgaria nigra (Link) Nees, Syst. Pilze 1: 54 (1817)!. 

≡ Sporotrichum nigrum (Link) Fr., Syst. mycol. 3(2): 416 (1832). 

≡ Trichosporum nigrum (Link) Fr., Summa veg. scand. 2: 492 (1849). 

= Sporotrichum fuliginosum Pers., Mycol. eur. 1: 77 (1822). [T: L]. 

= Botrytis atrofumosa Cooke & Ellis, Grevillea 6(39): 90 (1878)!. [T: K; NY]. 

≡ Virgaria atrofumosa (Cooke & Ellis) Sacc., Syll. fung. 4: 281 (1886)!. 

= Trichosporum splenicum Sacc. & Berl., Atti Reale Ist. Veneto Sci. Lett. Arti, Ser. 6, 3(4): 

741 (1885). [T: PAD]. 

Lit.: SACCARDO (1886: 356), HUGHES (1958: 751, 823). 

lignicola Corda, Icon. fung. 1: 14 (1837)!, as ‘lignicolum’. 

T: on rotten wood, Czech Republic, near Prag (PRM 155424). 


Lit.: COOKE (1871: 584), SACCARDO (1886: 356), LINDAU (1907: 809), FERRARIS (1912: 

340), HO et al. (1999: 132). 

Ill.: CORDA (1837: Tab. 3, Fig. 206), HO et al. (1999: 133, Figs 26–27). 

Notes: HUGHES (1958) reduced this species to a synonym of Cladosporium herbarum, which 

could be confirmed, based on a re-examination of the type material. ‘C. lignicola’ from North 

America, described and illustrated by HO et al. (1999), is quite distinct and the name is 

undoubtedly misapplied. 

lineolatum Sacc., Ann. Mycol. 12: 313 (1914)!. 

T: on faded or dead leaves of Capparis micracantha (Capparidaceae), Philippines, Prov. Rizal, 

Alabang, Mar. 1912, P.W. Graff, No. 16748 (PAD: lectotype; isolectotype: BPI 427242). 

Lit.: SACCARDO (1931: 789). 

Host(s)/substrate(s) & distribution: on Capparis micracantha; Asia (Philippines). 

63

linicola Pidopl. & Deniak, in Pidoplichko, Gribnaja Flora Grubych Kormov: 267 (1953)!, 

nom. inval. 

T: on seeds of flax (Linaceae), also found on damp hay, Ukraine, Kievsk. Oblast. 


liriodendri K. Schub. & U. Braun, sp. nov. 

T: on Liriodendron tulipifera, USA, New York, Tomkins co., Six Mile Ravine, 11 Sept. 1949, 

C. T. Rogerson, as ‘Cercospora or Cladosporium’ (NY: holotype). 

Host(s)/substrate(s) & distribution: on Liriodendron tulipifera; North America (USA). 

longipes Sorokīn, On nekot. bolez. vinograda i drug. rast. Kavkazk. Kraja, Tiflis: 26 (1892), 

also in Z. Pflanzenkrankh. 3: 154 (1893)!. 

T: on leaves of Vitis vinifera (Vitaceae), Caucasus, ‘im kubanischen Bezirk’. 

Lit.: SACCARDO (1895: 619). 

Ill.: SOROKĪN (1893: Tab. 4, Figs 49–50). 

lonicerae Sawada, Rep. Gov. Res. Inst. Formosa 86: 163 (1943), nom. inval. 

T: on Lonicera japonica var. sempervillosa (Caprifoliaceae), Taiwan, Taipeh, 20 Dec. 1914, 

K. Sawada (BPI 427243). 

= Stenella lonicericola (Y.H. He & Z.Y. Zhang) K. Schub., H.D. Shin & U. Braun, in 

Schubert & Braun, Fungal Diversity (2005), in press. 

lonicericola Y.H. He & Z.Y. Zhang, Mycosystema 20(4): 469 (2001)! and in Zhang et al., 

Flora Fungorum Sinicorum, Vol. 14: 116 (2003)!. 

T: on living leaves of Lonicera japonica (Caprifoliaceae), China, Yunnan Prov., Kunming, 3 

Aug. 1990, H. Li (MHYAU 03533: holotype). 

≡ Stenella lonicericola (Y.H. He & Z.Y. Zhang) K. Schub., H.D. Shin & U. Braun, in 

Schubert & Braun, Fungal Diversity (2005). 

Ill.: HE & ZHANG (2001: 469, Fig. 1), ZHANG et al. (2003: 117, Fig. 75), SCHUBERT & BRAUN 

(2005b, Figs 7–8). 

Notes: Abelia biflora and Leycesteria formosa are reported as additional hosts. Cladosporium 

lonicerae Sawada is cited as synonym, which could be confirmed based on a re-examination. 

lophodermii Georgescu & Tutunaru, Rev. Biol. (Bucharest) 3(1): 61 (1958)!. 

T: on apothecia of Lophodermium pinastrum (Rhytismataceae) on leaves of Pinus silvestris 

(Pinaceae), Romania, Poiana Stalin at Postăvaru, Jun. 1956. 


Ill.: GEORGESCU & TUTUNARU (1958: 60, Fig. 14). 

lupiniphilum U. Braun, A monograph of Cercosporella, Ramularia and allied genera (Phytopathogenic 

Hyphomycetes) 2: 410 (1998)!. 

T: on Lupinus luteus (Fabaceae), Byelorussia, Minsk area, 1959, Čenaškskaja (?) (LEP: 

holotype, as ‘Ramularia lupini’). 

Ill.: BRAUN (1998: 411, Fig. 664). 

Host(s)/substrate(s) & distribution: on Lupinus luteus; Europe (Byelorussia). 

lychnidis Z.Y. Zhang & Y.L. Liu, in Zhang, Liu, Wei & He, Plant Diseases and Their 

Control: 104 (1998)!. 

T: on Lychnis coronata (Caryophyllaceae), China, Hubei, Wuchang, 21 Sept 1980, J.Y. Li & 

T.Y. Zhang, No. 13632 (MHYAU 03958: holotype). 

Ill.: ZHANG, LIU, WEI & HE (1998: 105, Fig. 2); ZHANG et al. (2003: 118, Fig. 76). 

lycoperdinum Cooke, Grevillea 12(61): 32 (1883)!. 

T: on Lycoperdon, USA, South Carolina, Aiken, Rav., F. amer. exs. 595 (K 121561: lectotype, 

selected by HEUCHERT et al., 2005). Isolectotypes: Rav., F. amer. exs. 595 (e.g., BPI 427244; 

NY; PH 01020430). 


Ill.: HEUCHERT et al. (2005: 34–35, Figs 11−12). 

64

Host(s)/substrate(s) & distribution: on gills of several Agaricales, Boletales, Polyporales and 

Russulales, including Gomphidius viscidus, Lactarius volemus, Lycoperdon and Polyporus 

spp.; Europe (Germany), North America (USA), South America (Uruguay). 

lycopersici Plowr., Gard. Chron. 16: 621 (1881)!. 

T: on fruits of Solanum lycopersicum (= Lycopersicon esculentum) (Solanaceae), Great 

Britain. 


Ill.: PLOWRIGHT (1881: 621, Fig. 121). 

lysimachiae H.C. Greene, Trans. Wisconsin Acad. Sci. 38: 232 (1946)!, nom. illeg., homonym, 

non C. lysimachiae Guba, 1939. 

T: on living leaves and stems of Lysimachia terrestris (Primulaceae), USA, Wisconsin, Dane 

Co., Madison, Univ. Wisconsin Arboretum, Marsh, 24 Aug. 1943, H.C. Greene (BPI 427245, 

WIS: syntypes). 

≡ Fusicladium lysimachiae (Guba) K. Schub. & U. Braun, Mycol. Progr. 4(2): 102 (2005)!. 

lysimachiae Guba, Rhodora 41: 513 (1939)!. 

T: on living leaves, rarely on the stems of Lysimachia vulgaris (Primulaceae), USA, Massachusetts, 

Nantucket Co., Nantucket, in waste places near the waterfront east of Main Street, 15 

Aug. 1936, E.F. Guba, No. 115 (ILL 21101: holotype). 

≡ Fusicladium lysimachiae (Guba) K. Schub. & U. Braun, Mycol. Progr. 4(2): 102 (2005)!. 

= Cladosporium lysimachiae H. C. Greene, Trans. Wisconsin Acad. Sci. 38: 232 (1946)!, nom. 

illeg. 


lythri Westend., Bull. Acad. Roy. Sci. Belgique 21(8): 240 (1854)!. 

T: on leaves of Lythrum salicaria (Lythraceae), Belgium, near Courtrai, Westendorp, No. 

1091 (BR). 

≡ Cercospora lythri (Westend.) Niessl, Hedwigia 15: 1 (1876)!. 

≡ Stenella lythri (Westend.) J.L. Mulder, Trans. Brit. Mycol. Soc. 65: 517 (1975)!. 

= Cercospora sanguinea Fuckel, Hedwigia 5: 30 (1866)!. [T: Fuckel, F. rhen. 1630, e.g., 

HAL]. 

Lit.: SACCARDO (1886: 452), LINDAU (1910: 122, 803), CHUPP (1954: 362), CROUS & BRAUN 

(2003: 259). 

machili Sawada, Special Publ. Coll. Agric. Natl. Taiwan Univ. 8: 196 (1959)!, nom. inval. 

T: on leaves of Machilus thunbergii (= Persea) (Lauraceae), Taiwan, Pref. Taipei, Taipei, 15 

Nov. 1914, Y.F (PPMH: holotype). 

Notes: Excluded, taxonomic status remains unclear. 

macrocarpum Pers. (sic) – listed by SACCARDO & BERLESE (1884: 100). 

macrocarpum Preuss, in Sturm, Deutsch. Fl. 3(26): 27 (1848)!. 

T: on dead leaves of Eryngium pandanifolium (Apiaceae), Italy, Saccardo (Herb. mycol.) 419 

[PAD: neotype, selected by DE VRIES (1952) as ‘lectotype’]. Isoneotypes: Sacc., Mycoth. ital. 

1396, e.g., HBG. 

≡ Cladosporium herbarum [(Pers.: Fr.) Link] var. macrocarpum (Preuss) M.H.M. Ho & 

Dugan, in Ho, Castañeda, Dugan & Jong, Mycotaxon 72: 131 (1999)!. 

= Cladosporium brunneum Corda, Icon. fung. 1: 15 (1837)!, syn. nov. [T: PRM 657440]. 

= Cladosporium gracile Corda, Icon. fung. 1: 15 (1837)!, syn. nov. [T: PRM]. 

≡ Didymotrichum gracile (Corda) Bonord., Handb. Mykol.: 89 (1851)!. 

= Cladosporium cardariae Opiz, Lotos 5: 41 (1855)!, syn. nov. [T: PRM 657446, 657447]. 

= Heterosporium minutulum Cooke & Massee, in Cooke, Grevillea 16(77): 11 (1887)!. [T: K]. 

= Cladosporium algarum Cooke & Massee, in Cooke, Grevillea 16(79): 80 (1888)!. [T: K]. 

≡ Heterosporium algarum (Cooke & Massee) Cooke & Massee, in Cooke, Grevillea 18(88): 

74 (1890)!. 

= Heterosporium typharum Cooke & Massee, in Cooke, Grevillea 16(79): 80 (1888)!. [T: 

NY]. 

65

= Heterosporium phragmitis var. typharum Cooke, Grevillea 16(80): 109 (1888)!, nom. inval. 

[T: K]. 

= Heterosporium maculatum Klotzsch ex Cooke, Grevillea 17(83): 65 (1888)!. [T: K]. 

= Heterosporium hybridum Ellis & Everh., J. Mycol. 5: 70 (1889)!. [T: NY]. 

= Heterosporium cleomis Ellis & Everh., in Kelsey, J. Mycol. 5: 82 (1889)!, nom.inval. 

= Fusicladium destruens Peck, Rep. (Annual) New York State Mus. Nat. Hist. 43: 30 (1890). 

[T: NYS]. 

= Heterosporium tuberculans Ellis & Everh., Erythea 1: 203 (1893). [T: NY, K]. 

= Heterosporium sphaeriiforme Ellis & Everh., Proc. Acad. Nat. Sci. Philadelphia 46: 381 

(1894)!, as ‘sphaeriaeforme’. [T: NY]. 

= Heterosporium proteus Starbäck, Bot. Centralbl. 64: 382 (1895)!. [T: K]. 

= Cladosporium idesiae Bres., in Sydow, Hedwigia 35(1), Beih.: 62 (1896)!. [T: B 70-655]. 

= Heterosporium oxybaphi F. Patt., Bull. Torrey Bot. Club 27: 285 (1900)!. [T: BPI]. 

= Heterosporium calandriniae Massee, Bull. Misc. Inform. 175–177: 168 (1901). [T: K]. 

= Heterosporium chamaeropis Oudem., Beih. Bot. Centralbl. 11: 539 (1902)!. [T: L]. 

= Heterosporium hordei Bubák, Sber. K. böhm. Ges. Wiss., Math.-Nat. Kl. 1903(12): 20 

(1903). [T: BPI]. 

= Cladosporium magnoliae Lindau, Verh. Bot. Vereins Prov. Brandenburg 47: 74 (1905)!, 

syn. nov. [T: B]. 

= Heterosporium amsoniae Kabát & Bubák, in Bubák & Kabát, Hedwigia 47: 362 (1908)!. [T: 

BPI]. 

= Heterosporium tortuoso-inflatum Bubák, Bot. Közlem. 15(3–4): 82 (1915)!. [T: BPI]. 

= Cladosporium epiphyllum [(Pers.: Fr.) Nees] var. acerinum Sacc., Nuovo Giorn. Bot. Ital., 

N.S., 27: 86 (1920)!. [T: PAD]. 

= Heterosporium laricinum Dearn., Mycologia 21: 328 (1929)!. [T: DAOM]. 

= ? Heterosporium maydis Lobik, Trudy Severo-Kavkazsk. Inst. Zashch. Rast. 1(2): 42 

(1933). [T: LEP]. 

= Heterosporium thapsiae Petr., in Rechinger, Denkschr. Akad. Wiss. Wien 105(2): 26 

(1943)!. [T: W]. 

= ? Cladosporium herbarum var. phlei Lagière, Ann. École Natl. Agric. Grignon, Sér. 3, 5: 

159 (1946)!. 

= Heterosporium petuniae R. Sprague, Mycologia 57: 658 (1965)!. [T: WSP]. 

= Cladosporium equiseti Pass., in herb. (B 70-6324), syn. nov. 

= Heterosporium cerastis Jacz., nom. herb. [VIR ex LEP]. 

= Heterosporium trichostematis Jacz., nom. herb. [LE 41142]. 


FRAGOSO (1927: 200), DE VRIES (1952: 76), ELLIS (1971: 315), DOMSCH et al. (1980: 208), 

ELLIS & ELLIS (1985: 290, 468), MATSUSHIMA (1985: 5), MCKEMY & MORGAN-JONES 

(1991c), DUGAN & ROBERTS (1994), DAVID (1997: 71), SAMSON et al. (2000: 112). 

Ill.: PREUSS (1848: Tab. 14), GONZÁLES-FRAGOSO (1927: 200, Fig. 44), DE VRIES (1952: 77– 

78, Figs 16–17), ELLIS (1971: 314, Fig. 217 C), DOMSCH et al. (1980: 209, Fig. 84), 

MATSUSHIMA (1985: 58, Fig. 311), DUGAN & ROBERTS (1994: 516, Figs 1–3), DAVID (1997: 

62, Fig. 17 H–I; 89, Fig. 22 G–H), SAMSON et al. (2000: 112, Fig. 50; 113, Pl. 48). 

maculans (Catt.) Sacc. → oryzae Sacc. & P. Syd. 

maculans Schwein., Trans. Amer. Philos. Soc., N.S., 4(2): 277 (1832)!. 

T: under the epidermis of Jasminum fructicans (Oleaceae), USA, Pennsylvania, Bethlehem, 

No. 2599 (PH 01020418). 

≡ Cladosporium jasmini Schwein., in herb. 

Lit.: SACCARDO (1886: 355). 

maculatum Cooke, Fungi britannici exsiccati, No. 162 (?). 

On an unidentified host, Great Britain, Channel Islands, Jersey, Cooke, F. brit. exs. 162 (B 70- 

6613). 


Notes: Excluded. Material immature and poor, cercosporoid. 

66

maculicola Ellis & Barthol., in herb. 

On leaves of Populus monilifera, USA, Kansas, Rockport, Sept. 1894, E. Bartholomew, ex 

herb. Ellis (BPI 427256, type material of Cladosporium subsessile). 

= Fusicladium subsessile (Ellis & Barthol.) K. Schub. & U. Braun, IMI Descriptions of Fungi 


Notes: Specimen in Myc. Coll. (CASH 1952: 69). 

maculicola (Romell & Sacc.) M. Morelet, Bull. Soc. Sci. Nat. Archéol. Toulon & Var 201: 4 

(1972)!, as ‘maculicolum’. 

T: on living leaves of Populus tremula (Salicaceae), Sweden, Nacka Vikdalen, 24 Jun. 1890, 

L. Romell (S: lectotype; WINF(M) 11082, IMI 17008, PAD: isolectotypes). 

≡ Torula maculicola Romell & Sacc., in Saccardo, Grevillea 21(99): 69 (1893)!, non 

Fusicladium maculicola (Ellis & Kellerm.) Ondřej, 1973. 

≡ Phaeoramularia maculicola (Romell & Sacc.) B. Sutton, Canad. J. Bot. 48: 471 (1970)!. 

≡ Fusicladium romellianum Ondřej, Česká Mykol. 27(4): 237 (1973)!. 

= Pollacia borealis A. Funk, Canad. J. Bot. 67: 776 (1989)!. [T: DAVFP 23609]. 

Lit.: ELLIS (1976: 322), SCHUBERT & BRAUN (2002b, c), SCHUBERT et al. (2003: 90–92). 

madagascarense Sartory, Champignons Parasites de l’Homme et des Animaux 11: 729 

(1923)!, as ‘madagascarensis’. 

T: infection of human, Madagascar. 

≡ Cladosporium sp. (‘madagascariensis’) Guég., Compt. Rend. Hebd. Séances Acad. Sci. 

152(7): 412–413 (1911)!. 

≡ Cladosporium madagascariense Verdun, Précis de Parasitologie Humaine, éd. 2: ? (1912) 

[publication not found]. 

≡ Hormodendrum madagascarense (Verdun) C.W. Dodge, Med. Mycol.: 845 (1935)!. 

Lit.: DE HOOG et al. (2000: 1025). 

Notes: The name C. madagascariensis was not coined by Guéguen (C. sp.). A description for 

this species was given in SARTORY (1923: 729), who cited ‘Verdun, 1912’ without any details. 

But this publication could not be traced, and it is unclear if Verdun introduced the name C. 

madagascarense and if he published a first description. Doubtful species, no material known 

(DE HOOG et al. 2000). 

magnoliae Lindau, Verh. Bot. Vereins Prov. Brandenburg 47: 74 (1905)!. 

T: on leaves of Magnolia soulangeana (Magnoliaceae), Germany, Brandenburg, Tamsel, Dec. 

1904, Vogel (B 70-6614: holotype). 




magnusianum (Jaap) M.B. Ellis, More Dematiaceous Hyphomycetes: 337 (1976)!. 

T: on leaves of Narthecium ossifragum (Melanthiaceae), Denmark, Tǿnder, Rǿmǿ, peatbog by 

Twismark, Jul.-Aug. 1901, Jaap. 

≡ Heterosporium magnusianum Jaap, Schriften Naturwiss. Vereins Schleswig-Holstein 12: 

346 (1902)!. 

= Napicladium ossifragi Rostr., Bot. Fǽröes 1: 316 (1901). 

≡ Heterosporium ossifragi (Rostr.) Lind, Dan. fung.: 531 (1913)!. 

Lit.: ELLIS & ELLIS (1985: 390), DAVID (1995a; 1997: 85), HO et al. (1999). 

Ill.: DAVID (1995a: 1, Fig.; 1997: 88–89, Figs 21, 22 A–B), HO et al. (1999: 133, Figs 28–29) 

Host(s)/substrate(s) & distribution: on Narthecium ossifragum; Europe (Austria, Denmark, 

Germany, Great Britain, Ireland, Norway). 

Notes: Location of the types is unknown (DAVID 1997). 

malopis Fuss, Archiv Verein siebenb. Landesk., N.F., 14(2): 431 (1878)!, nom. nud. 

On overwintered stems of Malope malacoides (Malvaceae), Romania, Giresau, Fuss. 


malorum Rühle, Phytopathology 21: 1146 (1931)!. 

T: on living fruits of Pyrus malus (= Malus sylvestris) (Rosaceae), USA, Washington. 

67

≡ Cladosporium malorum Heald & Rühle, Wash. State Agric. Exp. Sta. Bull. 245: 48 (1930)!, 

nom. nud. 

≡ Alternaria malorum (Rühle) U. Braun, Crous & Dugan, in Braun, Crous, Dugan, 


= Phaeoramularia kellermaniana Marasas & I.H. Bredell, Bothalia 11: 217 (1974)! [IMI 

165252; PREM 44703]. 

≡ Cladophialophora kellermaniana (Marasas & I.H. Bredell) U. Braun & Feiler, Micro-biol. 

Res. 150: 83 (1995)!. 

≡ Pseudocladosporium kellermaniana (Marasas & I.H. Bredell) U. Braun, A monograph of 

Cercospora, Ramularia and allied genera 2: 393 (1998)!. 

= Cladosporium porophorum Matsush., Icones Fungorum a Matsushima Lectorum: 36 

(1975)!. 

Lit.: HO et al. (1999: 134). 

malvacearum C.D. Sharma, Gadp., Firdousi, A.N. Rai & K.M. Vyas, Indian Phytopathol. 

51(2): 156 (1998)!. 

T: on living leaves of Kydia calycina (Malvaceae), India, Madhya Pradesh, Shahdol circle, 

Amarkantak (south forest division), Jan. 1993, C.D. Sharma (S.U. Herb. No. C.S.5: holotype; 

IMI 254691: isotype?). 


Notes: Data given on the label of the collection in herb. IMI (254691) [on Grewia sp. 

(Tiliaceae), India, Univ. Gorakhpur, A.N. Rai] deviate from the data cited in the original 

description. 

manoutchehrii Esfand., Sydowia 5: 368 (1951)!. 

T: on living leaves of Quercus atropatena (Fagaceae), Iran, Polé Zangouleh, 27 Jul. 1948, 

Manoutchehri. 

mansonii (Castell.) Pinoy, Ann. Dermatol. Syphiligr., Sér. 5, 3: 341 (1912)! ? (not on this 

page). 

T: isolated from a human patient, India, Ceylon (CBS 158.58 = IMI 134457 = ATCC 18657: 

neotype, selected by DE HOOG, 1977). 

≡ Microsporum mansonii Castell., Brit. Med. J. 2: 1271 (1905). 

≡ Foxia mansonii (Castell.) Castell., J. Trop. Med. Hyg. 11: 261 (1908). 

≡ Malassezia mansonii (Castell.) Verdun, Précis de Parasitologie Humaine, éd. 2: 698 (1912). 

≡ Torula mansonii (Castell.) Vuill., Compt. Rend. Hebd. Séances Acad. Sci. 89: 406 (1929). 

≡ Sporotrichum mansonii (Castell.) Toro, Sci. Surv. Porto Rico & Virgin Islands 8: 222 

(1932), as ‘mansoni’. 

≡ Dematium mansonii (Castell.) C.W. Dodge, Med. Mycol.: 678 (1935)!. 

≡ Pullularia mansonii (Castell.) Borelli, Riv. Anat. Patol. Oncol. 17: 617 (1960). 

≡ Aureobasidium mansonii (Castell.) W.B. Cooke, Mycopathol. Mycol. Appl. 17: 34 (1962)!. 

≡ Rhinocladiella mansonii (Castell.) Schol-Schwarz, Antonie van Leeuwenhoek J. Microbiol. 

Serol. 34: 122 (1968). 

≡ Exophiala mansonii (Castell.) de Hoog, Stud. Mycol. 15: 114 (1977)!. 

≡ Wangiella mansonii (Castell.) Bièvre & Mariat, Bull. Soc. Franç. Mycol. Méd. 8(2): 127 

(1979). 

Lit.: NANNIZZI (1934: 405), KWON-CHUNG & BENNETT (1992: 191–192), DE HOOG et al. 

(2000: 1026). 

Notes: DE HOOG & HERMANIDES-NIJHOF (1977) and ‘Index fungorum’ give ‘Castell. & 

Chalm.’ as authors of the combination Cladosporium mansonii (Castell.), Manual of Tropical 

Medicine, ed. 2: 1100 (1913). KWON-CHUNG & BENNETT (1992) summarise nomenclatural 

controversies surrounding the name C. mansonii. According to these authors, it was Castellani 

who proposed the combination C. mansonii and attributed it to Pinoy. This name has been 

abandoned because of confusion. The neotype strain, CBS 158.58, is also the type strain of 

Exophiala castellani (DE HOOG et al. 2000). 

68

maracuja Viégas, Bragantia 6: 367 (1947)!. 

T: on Passiflora (Passifloraceae), Brazil, Prov. St. Pauli, Pindorama, Est. Exp. de Pindorama, 

19 Jul. 1935, A.S. Costa (IACM: holotype). 

Ill.: VIÉGAS (1947: 368, Fig. 5). 

Host(s)/substrate(s) & distribution: on Passiflora sp.; South America (Brazil). 

marinum A.K. Pal & Purkay., J. Mycopathol. Res. 30(2): 175 (1992)!. 

T: on living leaves of Avicennia marina (Avicenniaceae), India, West Bengal, Sundarban, 

Bakkhali, 10 Jun. 1991 (on the label) / 14 May 1991, A.K. Pal (IMI 351331: holotype). 

Ill.: PAL & PURKAYASTHA (1992: 174, Figs 3–5). 

Notes: Excluded, status unclear. 

martianoffianum Thüm., Byull. Moskovsk. Obshch. Isp. Prir., Otd. Biol. 55(1): 74 (1880). 

T: on living leaves of Populus laurifolia (Salicaceae), Russia, Sibiria, Minussinsk, near river 

Jenissei, Aug. 1879, N. Martianoff (M: lectotype; Thüm., Mycoth. univ. 2067: isolectotypes). 

≡ Fusicladium martianoffianum (Thüm.) K. Schub. & U. Braun, IMI Descriptions of Fungi 


= Fusicladium asiaticum Ondřej, Česká Mycol. 27(4): 237 (1973)!. [T: LE 161361]. 

Lit.: SACCARDO (1886: 357), LINDAU (1907: 818; 1910: 796), SCHUBERT et al. (2003: 64–65). 

melanophlaei Thüm., Flora 60: 412 (1877)!. 

T: on living leaves of Myrsine melanophlaeos (Myrsinaceae), South Africa, Promont, near 

Grahamstown, summer 1876, P. Mac. Owan, No. 1255. 

Lit.: SACCARDO (1886: 358). 

Notes: Type material could not be located. 

menispermi Allesch., Hedwigia 34: 220 (1895)!. 

T: on fading leaves of Menispermum canadense (Menispermaceae), Germany, Munich, 

botanical garden, Sept. 1894, Allescher (M-57686: holotype). 



metaniger (Castell.) Ferraris, Atti Ist. Bot. “Giovanni Briosi” 3: 183 (1932)!. 

T: isolated from a case of ‘trichomycosis nigra’ on human skin, Castellani. 

≡ Cryptococcus metaniger Castell., Archiv Dermatol. Syph. 16(4): 402 (1927)!. 

= Hortaea werneckii (Horta) Nishim. & Miyaji, Jap. J. Med. Mycol. 26(2): 145 (1984). 

Lit.: NANNIZZI (1934: 406), DODGE (1935: 675), COOKE (1962: 27), KWON-CHUNG & 

BENNETT (1992: 195). 

Notes: In Unesco (1955: 38) as ‘metanigrum’. See C. werneckii. 

metaplexis Z.Y. Zhang & X.Y. Wang, in Zhang, Wang, Liu & Li, Mycosystema 19(2): 165 

(2000)!. 

T: on living leaves of Metaplexis japonica (Asclepiadaceae), China, Heilongjiang, Harbin, 

Xiangfang, 2 Sept. 1992, Y.X. Wang & H. Li (MHYAU 07830: holotype). 

Ill.: ZHANG, WANG, LIU & LI (2000: 166, Fig. 1), ZHANG et al. (2003: 123, Fig. 80). 

micropilum (sic) Syd. – see C. microspilum. 

microporum Rabenh., in Cooke, Grevillea 17(83): 66 (1889)!. 

T: on leaves of Nerium oleander (Apocynaceae), Italy, Sardinia, Gonnos-Fanadiga, Dr. 

Marcucci, Unio itin. crypt. 42 (e.g., HBG; M-57685: syntypes). 

≡ Cladosporium microporum Rabenh., Unio itin. crypt., No. 42 (1866)!, nom. nud. 



Notes: Unio Itin. crypt. (1866) is a selection of species made by Marcucci and determinated by 

Rabenhorst. New species were not validly published, because descriptions were not given. 

COOKE (1889): ‘In our specimens only a minute species of Coniothyrium can be found.’ See 

C. microsporum Rabenh. 

microporum Roth – GOLA (1930: 21). 

Notes: See also C. microsporum Rabenh. 

69

microspermum Berk. & M.A. Curtis, in Berkeley, Grevillea 3(27): 107 (1875)!. 

T: on leaves of Quercus obtusiloba (Fagaceae), USA, South Carolina, No. 1686. 

Lit.: SACCARDO (1886: 360). 

Ill.: ZHANG, WEI & ZHANG (1998: 197, Fig. 3). 

Notes: ZHANG, WEI & ZHANG (1998) describe this species on Lithocarpus viridis from China, 

Yunnan. 

microspilum Syd. & P. Syd., Ann. Mycol. 18: 102 (1920)!. 

T: on leaves of Cissampelos pareira (Menispermaceae), Phillipines, Luzon, Prov. Laguna, Mt. 

Maquiling, 26 Apr. 1919, T. Collado, No. 6271. 

Lit.: SACCARDO (1931: 793). 

Notes: Type collection is not deposited at herb. S. 

microsporum Rabenh., sensu Saccardo. 

Lit.: SACCARDO (1892: 603, 1895: 617), FERRARIS (1912: 344), OUDEMANS (1923). 

Notes: SACCARDO (1892) changed the epithet ‘microporum’ in ‘microsporum’, because he 

thought that this name was misspelled by Rabenhorst. However, it is not quite clear what 

Rabenhorst meant, minute spores or pores. 

microsporum Trab., in Roum., F. sel. gall. exs., Cent. XV, No. 1426 (1881). 

T: on leaves of Nerium oleander (Apocynaceae), Algeria, Trabut. 

≡ Bispora trabutiana Sacc., Bull. Soc. Roy. Bot. Belgique 31(2): 237 (1892). 


microstictum Sacc. & D. Sacc., Ann. Mycol. 3: 169 (1905)!. 

T: on living leaves of Ulmus campestris (Ulmaceae), Italy, Treviso, Vittorio, Oct. 1899, Sacc., 

Mycoth. ital. 589 (e.g., BPI 427261; HBG). 

= Cladosporium hyophyllum Fuckel, Jahrb. Nassauischen Vereins Naturk. 23–24: 356 ‘1869’ 

(1870)!. 

Lit.: SACCARDO (1905: 165; 1906: 576), LINDAU (1907: 820), FERRARIS (1912: 345). 

mikaniae F. Stevens, Trans. Illinois Acad. Sci. 10: 208 (1917)!. 

T: on leaves of Mikania sp. (Asteraceae), Puerto Rico, Las Marias, 22 Mar. 1913, F.L. Stevens 

(ILL 314; IMI 119607). 

≡ Mycovellosiella mikaniae (F. Stevens) Deighton, Mycol. Pap. 137: 45 (1974)!. 

≡ Passalora mikaniae (F. Stevens) U. Braun & F. Freire, Cryptog. Mycol. 23: 300 (2002)!. 


milii Syd., Ann. Mycol. 12: 538 (1914)!. 

T: on leaves of Milium effusum (Poaceae), France, ‘Lothringen’, ‘Wald am Oetinger Tälchen’ 

near Forbach, 22 Jun. 1913, A. Ludwig, Syd., Mycoth. germ. 1295; on leaves of Millium 

effusum, ‘Ostpreussen, Warnicken, Samland’, 11 Jul. 1914, H. Sydow, Syd., Mycoth. germ. 

1296 (BPI 427263, HBG, M: syntypes). 

≡ Passalora milii (Syd.) G.A. de Vries, Contr. Knowl. Genus Cladosporium: 94 (1952)!. 


mimulicola U. Braun, Nova Hedwigia 58(1–2): 196 (1994)!. 

T: on Mimulus sp. (Scrophulariaceae), USA, California, 18 Jul. 1895, J.J. Davis (NY: holotype). 

Ill.: BRAUN (1994: 193, Pl. 1, Fig. 7). 

Host(s)/substrate(s) & distribution: on Mimulus sp.; North America (USA). 

minor R.F. Castañeda, Fungi Cubensis III: 22 (1988)!, as ‘minus’, nom illeg., homonym, non 

C. minor Spreng. 1827. 

T: on an unidentified leaf, Cuba, Prov. Pinar del Río, Viñales, 25 Sept. 1987, R.F. Castañeda 

(INIFAT C87/292-2: holotype). 

≡ Alysidium minus R.F. Castañeda & W.B. Kendr., Univ. Waterloo Biol. Ser. 35: 6 (1991)!, 

nom. nov., as ‘(R.F. Castañeda) R.F. Castañeda & W.B. Kendr.’. 

70

≡ Castanedaea minor A. Baker & Partridge, in Partridge, Baker & Morgan-Jones, Mycotaxon 

78: 178 (2001)!, nom. nov., as ‘(R.F. Castañeda) A. Baker & Partridge’. 

minor Spreng., Syst. veg. 4(1): 553 (1827)!, as ‘minus’. 

T: on fallen twigs and on stems of herbaceous plants. 

= ? Helminthosporium nanum Nees, Syst. Pilze 1: 67 (1817)!. 

Notes: cited in LINDAU (1910: 40). 

minourae Iwatsu, Mycotaxon 20(2): 523 (1984)!. 

T: from rotting wood, Japan, Chiba, Shiroi, 11 Feb. 1979, T. Iwatsu (IFM 4700: holotype in 

Depart. Pathog. Fungi, Research Inst. Chemobiodynamics, Chiba, Japan). [Culture ex-type: 

ATCC 52853 (= CBS 556.83 = IMI 298056)]. 

≡ Cladophialophora minourae (Iwatsu) Haase & de Hoog, in Haase, Sonntag, Melzer-Krick 

& de Hoog, Stud. Mycol. 43: 94 (1999)!. 

Lit.: HO et al. (1999: 134). 

minus R.F. Castañeda → minor R.F. Castañeda. 

minus Spreng. → minor Spreng. 

minusculum Sacc., Ann. Mycol. 11: 20 (1913)!. 

T: on excrements of insects on living leaves of Salix alba (Salicaceae), Malta, Ghain el Gbira, 

Oct. 1911, Caruana Gatto (PAD: holotype; IMI 70294: isotype). 

Lit.: FERRARIS (1914: 886), SACCARDO (1931: 798), BRAUN (2000: 34). 

Ill.: BRAUN (2000: 37, Fig. 5). 

Host(s)/substrate(s) & distribution: on Salix alba; Europe (Malta). 

Notes: C. minusculum is morphologically close to Cladosporium cladosporioides, but differs 

in having strongly geniculate-sinuous conidiophores and consistently verruculose conidia 

(BRAUN 2000). 

miyakei Sacc. & Trotter, Syll. fung. 22: 1370 (1913)!. 

T: on leaves of Oryza sativa (Poaceae), Japan. 

≡ Cladosporium oryzae I. Miyake, J. Coll. Agric. Imp. Univ. Tokyo 2: 262 (1910), nom. illeg., 

homonym, non C. oryzae Sacc. & P. Syd., 1899. 

Lit.: PADWICK (1950: 170). 

Ill.: MIYAKE (1910: Tab. 14, Figs 68–70). 

Notes: ‘A. Cl. maculani mycelio superficiali distinctum.’ (SACCARDO 1913). 

modestum Syd., Ann. Mycol. 37: 252 (1939)!. 

T: on living leaves of Anthostema senegalensis (Euphorbiaceae), Sierra Leone, Kenema, 5 

Dec. 1938, F.C. Deighton (IMI 7520). 

≡ Denticularia modesta (Syd.) Deighton, Trans. Brit. Mycol. Soc. 59(3): 422 (1972)!. 

Lit.: ELLIS (1976: 183). 

moldavicum Fosteris, Bull. Sect. Sci. Acad. Roumaine 26(7): 494 (1944)? and in Herb. 

Mycol. Rom., Fasc. 27, No. 1341 (1944)!, nom. inval. 

T: on Festuca ovina (Poaceae), Romania, Moldova, Neamț District, Broșteni, Neagră Valley, 

15 Aug. 1943, S. Fosteris, Herb. Mycol. Rom. 1341 (BPI 427266, MA-Fungi 8381: syntypes). 



Notes: Page 494 does not exist, name not published in this volume, and the descriptions in 

Herb. Mycol. Rom. 1341 are not in Latin. 

molle Cooke, Grevillea 6(40): 139 (1878)!. 

T: on under surface of dead leaves of Asclepias (Asclepiadaceae), USA, South Carolina, 

Aiken [IMI (slide); K; M]. 

≡ Cercospora molle (Cooke) Deighton, in herb. (IMI). 

= Cercospora venturioides Peck, Rep. (Annual) New York State Mus. Nat. Hist. 34: 47 

(1881). [T: NYS]. 

71

≡ Mycovellosiella venturioides (Peck) U. Braun, Proc. Komarov Bot. Inst. (St. Petersburg) 20: 

99 (1997)!. 

≡ Passalora venturioides (Peck) U. Braun & Crous, in Crous & Braun, Mycosphaerella and 

its anamorphs: 1. Names published in Cercospora and Passalora, CBS Biodiversity Ser. 1: 

419 (2003)!. 

= Cercospora illionensis Barthol., F. columb., No. 2611 (1908). [T: Barthol., F. columb. 2611, 

e.g., HBG; NY]. 


monardae H.C. Greene, Amer. Midl. Naturalist 50(2): 508 (1953)!. 

T: on living leaves of Monarda punctata (Lamiaceae), USA, Wisconsin, Dane Co. Madison, 

Univ. Wisconsin Arboretum, Prairie, 10 Aug. 1952, H.C. Greene (BPI 427271, WIS: 

syntypes). 

≡ Fusicladium monardae (H.C. Greene) K. Schub. & U. Braun, in Schubert, Mycotaxon 92: 

71 (2005). 


mori (Yendo) H. Zhang & Z.Y. Zhang, Proceedings of Phytopathological Symposium 

Organized by Phytopathology Laboratory of Yunnan Province 2: 306 (1998). 

T: on Morus sp. (Moraceae), Japan. 

≡ Hormodendrum mori Yendo, Dai-Nihon Sanshi Kaiho 335: 6 (1919)!. 


Notes: recorded from China on Morus alba. 

multigeniculatum W. Yamam., Sci. Rep. Hyogo Univ. Agric., Ser. Agric. 4(1): 3 (1959)!, 

nom. inval. 

T: on Phyllostachys reticulata, Japan. 

Lit.: HO et al. (1999: 136). 

Ill.: YAMAMOTO (1959: 4, Figs 13–16), HO et al. (1999: 135, Figs 33–34). 

Notes: Yamamoto did not designate a type. HO et al. (1999) select figures 33/34 as ‘lectotype’ 

(iconotype) of this species, with lyophilized culture of ATCC 38012, batch 12-13-78 as 

epitype. This ‘lectotypification’ is, however, incorrect since the figures cited are not elements 

from the protologue. Hence, the name C. multigeniculatum was not validated by Ho et al. 

(1999). A formal validation is necessary and will be published elsewhere in connection with a 

re-examination of this fungus. 

murorum Petr., Ann. Naturhist. Mus. Wien 52: 288 (1941)!. 

T: on a kitchen wall, Austria, Wien, Apr. 1941, H. Lohwag (M). 

Lit.: RIEDL (1968). 

musae E.W. Mason, in E.B. Martyn, Mycol. Pap. 13: 2 (1945)!. 

T: on leaves of Musa sp. (Musaceae), Jamaica, 7 Sept. 1942, E.B. Martyn (IMI 7521, slide, ex 

type collection). 

≡ Periconiella sapientumicola Siboe, African J. Mycol. Biotechnol. 1994: 4 (1994), non 

Periconiella musae M.B. Ellis, 1967. 

≡ Metulocladosporiella musae (E.W. Mason) Crous, Schroers, Groenewald, U. Braun & K. 

Schub., Mycol. Res. (2005), in press. 

Lit.: ELLIS (1971: 317), DAVID (1988c), HO et al. (1999: 136). 

Ill.: MARTYN (1945: 3, Fig. 1), ELLIS (1971: 318, Fig. 219 A), DAVID (1988c: 1, Fig.), HO et 

al. (1999: 138, Figs 35–37), CROUS et al. (2005, Figs 3–14). 

myriosporum Ellis & Dearn., Proc. Canad. Inst., N.S., 3, 1: 90 (1897)!. 

T: on legumes of Pisum (Fabaceae), Canada, British Columbia (DAOM). 

Lit.: SACCARDO (1899: 1080), CASH (1952: 69). 

myrmecophilum (Fresen.) Bayl. Ell., Trans. Brit. Mycol. Soc. 5(1): 138 (1914)!. 

T: in a nest of Lasius fuliginosus (Insecta), Germany. 

≡ Septosporium myrmecophilum Fresen., Beitr. Mykol. 3: 49 (1863)!. 

≡ Macrosporium myrmecophilum (Fresen.) Sacc., Syll. fung. 4: 538 (1886)!. 

72

≡ Cladotrichum myrmecophilum (Fresen.) Lagerh., Entomol. Tidskr. 1900: 17 (1900). 

Lit.: SACCARDO (1931: 798). 

Ill.: FRESENIUS (1863: Tab. 6, Figs 29–32), BAYLISS ELLIOTT (1914: Tab. 2, Figs 1–4). 

Notes: Material examined by Jessie S. Bayliss Elliott was supplied by H. Donisthorpe from 

freshly excavated ant (L. fuliginosus and L. umbratus) nests. Further designation or deposit of 

material not specified by Elliott. FRESENIUS (1863) described his new species from the surface 

of a nest of Formica fuliginosa (= Lasius fuliginosus) consisting of wood fibres of Picea. 

myrtacearum K. Schub., U. Braun & R.G. Shivas, in Braun, Cunnington, Priest, Shivas, & 

Schubert, Australas. Pl. Pathol. (2005), in press. 

T: on Corymbia polycarpa (Myrtaceae), Australia, Northern Territory, Millingimbi, Townsite, 

17 Aug. 1999, A.A. Mitchel (BRIP 26527: holotype; DNAP 26527: isotype). 

Ill.: BRAUN et al. (2005, Fig. 5). 

Host(s)/substrate(s) & distribution: on Corymbia polycarpa; Australia. 

myrticola Bubák, in Bubák & Kabát, Ann. Mycol. 13: 113 (1915)!, as ‘myrticolum’. 

T: on leaves of Myrtus communis (Myrtaceae), Italy, Tyrol, Gries near Bozen, 30 May 1914, 

Dr. W. Pfaff (BPI 427273: holotype). 

≡ Fusicladium myrticola (Bubák) K. Schub. & U. Braun, Fungal Diversity (2005), in press. 

Lit.: SACCARDO (1931: 793). 


Notes: In the type material, the month of the collecting date is given as June. 

myrticola R.F. Castañeda & W.B. Kendr., Univ. Waterloo Biol. Ser. 35: 20 (1991)!, nom. 

illeg., homonym, non C. myrticola Bubák, 1915. 

T: on dead leaves of Syzygium jambos (Myrtaceae), Cuba, Pinar del Río, Cuchillas de San 

Simón, 24 Mar. 1990, R.F. Castañeda (INIFAT C90/108: holotype). 



neocheiropteridis Y.L. Liu & Z.Y. Zhang, in Liu, He & Zhang, Mycosystema 19(2): 169 

(2000)!. 

T: on living leaves of Neocheiropteris palmatopedata (Polypodiaceae), China, Yunnan, 

Kunming, Jindian, 4 Apr. 1997, K. Li & H. Xiong (MHYAU 07827: holotype). 

Ill.: LIU et al. (2000: 170, Fig. 1), ZHANG et al. (2003: 129, Fig. 85). 

neottopteridis Y.L. Liu & Y.H. He, in Liu, He & Zhang, Mycosystema 19(2): 169 (2000)!. 

T: on leaves of Neottopteris nidus, China, Yunnan (MHYAU 07828: holotype). 

Ill.: LIU et al. (2000: 170, Fig. 2) , ZHANG et al. (2003: 130, Fig. 86). 

nerii Gonz. Frag., Mem. Real Acad. Ci. Barcelona, Ser. 3, 15(17): 459 (33) (1920)!. 

T: on faded and dried leaves of Nerium oleander (Apocynaceae), Spain, Baleares, Ibiza, near 

S. Juan, 24 Mar. 1918, Dr. Font Quer. 


Notes: Type material could not be located in herb. MA. ZHANG et al. (2003) reported the 

species on Nerium indicum from Sichuan, China. 

neriicola S.A. Khan & M.A. Kamal, Mycopathol. Mycol. Appl. 52(1): 33 (1974)!. 

T: on leaves of Nerium indicum (Apocynaceae), Pakistan, Tando Mohd Khan, Faugi Sugarcane 

Farm, 11 Oct. 1966, S.A. Khan (IMI 123901: holotype). 

Ill.: KHAN & KAMAL (1974: 34, Fig. 4). 

Host(s)/substrate(s) & distribution: on Nerium indicum; Asia (Pakistan). 

nervale Ellis & Dearn., in Bartholomew, F. columb., Cent. 21, No. 2010 (1905)!. 

T: on living leaves of Rhus typhina (Anacardiaceae), Canada, London, Jul./ Aug. 1904, J. 

Dearness, Barthol., F. columb. 2010 (BPI 427277–427278; ILL; NY: syntypes). 

= Cladosporium aromaticum Ellis & Everh., Proc. Acad. Nat. Sci. Philadelphia 1895: 439 

(1895)!. 

73

≡ Fusicladium aromaticum (Ellis & Everh.) K. Schub. & U. Braun, Fungal Diversity (2005), 

in press. 

Lit.: SACCARDO (1913: 1367; 1931: 788), CASH (1952: 69), ZHANG et al. (1999: 42). 

Notes: ZHANG et al. (1999) reported this species from China as causal agent of leaf spots on 

Rohdea japonica (Convallariaceae), which is very doubtful. 

nervisequum Mont., Ann. Sci. Nat. Bot., Sér. 4, 8: 298 (1857)!. 

T: on leaves of Eriobotrya japonica (Rosaceae), France, L. Castagne, No. 2789. 


nicotianae Oudem., Ned. Kruidk. Arch., Ser. 3, 2(3): 769 (1902)!. 

T: on decaying leaves of Nicotiana tabacum (Solanaceae), Netherlands, Amerongen, Jul. 

1901, C.J. Koning (L). 

= Cladosporium tabaci Oudem., Beih. Bot. Centralbl. 11: 538 (1902)!. 


Notes: The type is represented by only two drawings, the description is short and the drawings 

are so poor that this species was regarded as doubtful [a nomen dubium] (DE VRIES 1952). 

nigrelloides U. Braun & Mouch., in Braun, Mouchacca & McKenzie, New Zealand J. Bot. 

37(2): 302 (1999)!. 

T: on leaves of Manihot utilissima (Euphorbiaceae), French Polynesia, Isles Gambier, 

Mangareva, Rikitaea, 2 Apr. 1966, Huguenin (PS 66.684, PC: holotype, as ‘Cercospora 

henningsii’). 

Ill.: BRAUN et al. (1999: 300, Fig. 2). 

Host(s)/substrate(s) & distribution: on Manihot utilissima; Oceania (French Polynesia). 

nigrellum Ellis & Everh., Proc. Acad. Nat. Sci. Philadelphia 1893: 463 (1894)!. 

T: on inner bark of railroad ties, USA, West Virginia, Fayette Co., Nuttallburg, 20 Oct. 1893, 

L.W. Nutall, Flora of Fayette County, No. 172 (BPI; NY); Ellis & Everh., N. Amer. F. 3086 

(e.g., PH) and Ellis & Everh., F. columb. 382 (e.g., PH). 

Lit.: SACCARDO (1895: 620), ELLIS (1976: 329), MORGAN-JONES (1977: 5), HO et al. (1999: 

137). 

Ill.: ELLIS (1976: 328, Fig. 246), MORGAN-JONES (1977: 4, Fig. 2), HO et al. (1999: 138, Fig. 

38). 

Notes: OUDEMANS (1920) listed Salix as substrate. 

nitrariae Dumitraș & Bontea, in Bontea & Dumitraș, Rev. Roumaine Biol., Sér. Bot. 12(6): 

387 (1967)!. 

T: on living leaves, fruits and twigs of Nitraria schoberi (Zygophyllaceae), Romania, Ploești, 

between Pîclele Mari and Pîclele Mici, Jul. 1962, Tiberiu Oprescu (BUCM). 

Ill.: BONTEA & DUMITRAȘ (1967: 388, Fig. 2). 

nodulosum Corda, Icon. fung. 1: 15 (1837)!. 

T: on rotting wood of Carpinus sp. (Corylaceae), Czech Republic, Bohemia (PRM). 

≡ Didymotrichum nodulosum (Corda) Bonord., Handb. Mykol.: 89 (1851)!. 



Lit.: COOKE (1871: 585), SACCARDO (1886: 351, 358), LINDAU (1907: 806), FERRARIS (1912: 

335; 1914: 883), OUDEMANS (1919), HUGHES (1958: 751). 


oblongum Bres., Stud. Trent. 7(2): 74 (1926)!. 

T: on stems of Zinnia elegans (Asteraceae), Italy, Trento, Feb. 1923, G. Bresadola (BPI 

427289). 

obtectum Rabenh. ex Cooke, Grevillea 17(83): 66 (1889)!. 

T: on fadding leaves of Artemisia maritima (Asteraceae), Italy, Sardinia, Alghero, Dr. 

Marcucci, Unio itin. crypt. 36 (e.g., HBG) and Rabenh., F. eur. 2783 (e.g., HBG: syntypes). 

≡ Cladosporium obtectum Rabenh., Unio itin. crypt., No. 36 (1866)!, nom. nud. 

74

Lit.: SACCARDO (1892: 602, 1895: 619), FERRARIS (1912: 348), ELLIS (1976: 342). 

Ill.: ELLIS (1976: 341, Fig. 259 C). 

Host(s)/substrate(s) & distribution: on Artemisia maritima; Europe (Italy). 

Notes: Unio itin. crypt. (1866) is a selection of species made by Marcucci and determinated by 

Rabenhorst. New species were not validly published, because descriptions were not given. 

obtectum Roth – listed by GOLA (1930: 21). 

occultum Ces., Atti Accad. Sci. Fis. 8(4): 25 (1879)!. 

T: on hidden glumes of Sporobolus sp. (Poaceae), Borneo, Pulo-Pinang, May 1865, C.O. 

Beccari. 

≡ Dactylosporium occultum (Ces.) Cif. & Vegni, in Vegni, Riv. Patol. Veg., Sér. 3, 3: 207 

(1963)!. 

Lit.: SACCARDO (1886: 364). 

Notes: Two specimens in Cesati collection at RO (Palmer Marchi, personal communication). 

oleacinum – Sacc., Syll. fung. 14: 900 (1899). 

(= ? Chaetophoma oleacina Cooke). [Under description of Ch. oleacina: “Peritheciis (sic) 

gregariis, subglobosis, inter hyphas fuligineas…cladosporioideas, conidia fuligenea, uniseptata, 

8,4–10,5 x 3½–6 gerentes (Cladosporium oleacinum) oriundis, …; sporulis ovoideis, 2,3– 

2,6 x 1–1,3 …” etc. 

oleae Ellis & Everh., in herb. 

On Olea europaea, USA, California, Santa Barbara, 1 Jun. 1895, J.J. Davis (WIS). 

oligocarpum Corda, Icon. fung. 1: 14 (1837)!. 

T: on rotting wood, Czech Republic, near Reichenberg (PRM 155428). 

≡ Didymotrichum oligocarpum (Corda) Bonord., Handb. Mykol.: 89 (1851)!. 



Notes: Excluded, no Cladosporium s. str., but status still unclear. 

oligocarpum [Corda] var. malvacearum Berk., J. Linn. Soc., Bot. 14: 353 (1875)!. 

T: on leaves of some malvaceous plant, Cape-Verdes, St. Vincent, Aug. 1873. 

Lit.: SACCARDO (1886: 352)!. 

Notes: This seems to be an error in SACCARDO (1886), because BERKELEY (1875) did not 

describe a new variety rather he considered it to be C. oligocarpum Corda though it is very 

difficult to speak positively without authentic specimens. 

olivaceum (Corda) Bonord., Handb. Mykol.: 72 (1851)!. 

T: on rotten wood of Picea (Pinaceae), Czech Republic. 

≡ Mydonosporium olivaceum Corda, in Sturm, Deutschl. Fl. 3(13): 95 (1833)!. 


Ill.: CORDA (1833: Tab. 48). 

Notes: Type material could not be located in herb. PRM. 

oncobae K. Schub. & C.F. Hill, sp. nov. 

T: on leaves of Oncoba spinosa (Flacourtiaceae), New Zealand, Auckland, Princes Street, 

Auckland University Campus, 19 Sept. 2004, C. F. Hill, No. 1076 (HAL 1832 F: holotype). 

Host(s)/substrate(s) & distribution: on Oncoba spinosa; New Zealand. 

opacum Schulzer & Sacc. – cited in OUDEMANS (1919, 1924). 

Notes: ‘Cladosporium opacum Schulzer & Sacc.’ appears to be an error in OUDEMANS (1919, 

1924). Cladotrichum opacum Schulzer & Sacc., Hedwigia 23: 127 (1884)! was intended. 

ophiopogonis T. Zhang & Z.Y. Zhang, Plant Diseases and Their Control: 110 (1998)!. 

T: on living leaves of Ophiopogon mairei (Liliaceae s. lat.), China, Zhejiang, Hangshou, 3 

Nov. 1980, J.Y. Li & T.Y. Zhang, No. 44132 (MHYAU 03951). 

Ill.: ZHANG & ZHANG (1998b: 110, Fig. 2), ZHANG et al. (2003: 136, Fig. 91). 

Notes: Chlorophytum elatum was recorded from Hubei, China as an additional host. 

75

oplismeni Syd., Philipp. J. Sci., (Ser. C., Bot.) 8: 507 (1913). 

T: on spikes of Oplismenus undulatifolius (Poaceae), Philippines, Luzon, Ifugao, Mt. Polis, 

McGregor. 

Lit.: SACCARDO (1931: 792). 

Notes: Type material is not in herb. S. 

orbiculans Schwein.?, in herb. 

On leaves of Ixora sp. (Rubiaceae), Surinam (PH 01020416). 

Notes: Excluded, but status unclear. 

orbiculatum Desm., Ann. Sci. Nat. Bot., Sér. 3, 11: 275 (1849)!. 

T: on living leaves of Sorbus domestica (Rosaceae), herb. Desmazières (PC: lectotype). 

≡ Fusicladium orbiculatum (Desm.) Thüm., F. austr., Cent. VIII, No. 774 (1873)!. 

≡ Passalora dendritica var. orbiculata (Desm.) Berk., in Saccardo, Mycoth. ven., Cent. XII, 

No. 1246 (1876)! and Michelia 1(2): 265 (1878)!. 

≡ Fusicladium dendriticum var. orbiculatum (Desm.) Sacc., Syll. fung. 4: 345 (1886)!. 


Teleomorph: Venturia inaequalis (Cooke) G. Winter, Hedwigia 36: 81 (1897)!. 

Lit.: LINDAU (1907: 782), OUDEMANS (1921), RITSCHEL (2001), CROUS & BRAUN (2003: 

485), SCHUBERT et al. (2003: 76). 

Notes: For further synonyms please see SCHUBERT et al. (2003). 

orchidearum Cooke & Massee, in Cooke, Grevillea 16(79): 80 (1888)!. 

T: on leaves of orchids (Oncidium crispum) (Orchidaceae), Great Britain, Surrey, Kew, Kew 

Gardens, H. Low’s nursery, Borough of Richmond (NY 72454: holotype). 

Lit.: SACCARDO (1892: 605), ELLIS (1976: 339), SCHUBERT & BRAUN (2004: 305–306). 

Ill.: ELLIS (1976: 339, Fig. 257 B), SCHUBERT & BRAUN (2004: 306, Fig. 4). 

Host(s)/substrate(s) & distribution: on Oncidium crispum; Europe (Great Britain). 

orchidiphilum K. Schub. & U. Braun, Sydowia 56(2): 306–308 (2004). 

T: on leaves of Cypripedium sp. (Orchidaceae), Australia, Canterbury, 19 Aug. 1914 (VPRI 

2488: holotype). 


Host(s)/substrate(s) & distribution: on Cypripedium sp.; Australia. 

orchidis E.A. Ellis & M.B. Ellis, in M.B. Ellis, Mycol. Pap. 131: 17 (1972)!. 

T: on living leaves of Orchis praetermissa (= Dactylorhiza majalis subsp. praetermissa) 

(Orchidaceae), Great Britain, Norfolk, Horsey Warren, 17 Jul. 1955, E.A. Ellis (IMI 60545: 

holotype). 

≡ Fusicladium orchidis (E.A. Ellis & M.B. Ellis) K. Schub. & U. Braun, Sydowia 56(2): 314 

(2004). 

Lit.: ELLIS (1976: 338), ELLIS & ELLIS (1985). 

Ill.: ELLIS (1972: 18, Fig. 17), ELLIS (1976: 339, Fig. 257 A), SCHUBERT & BRAUN (2004: 

315, Fig. 8). 

oreodaphnes Allesch. ex K. Schub., sp. nov. 

T: on faded leaves of Oreodaphne foetens (= Ocotea foetens) (Lauraceae), Germany, Berlin, 

botanical garden, 1894, P. Hennings, No. 40 (M-57756: holotype). 

≡ Cladosporium oreodaphnes Allesch., in herb. 

Host(s)/substrate(s) & distribution: on Ocotea foetens; Europe (Germany). 

ornithogali (Klotzsch ex Cooke) G.A. de Vries, Contr. Knowl. Genus Cladosporium: 49 

(1952)!. 

T: on leaves of Ornithogalum umbellatum (Hyacinthaceae), Germany, spring, Klotzsch, Herb. 

viv. myc. 69 (M: lectotype, selected by DAVID, 1997; K: isolectotype). 

≡ Heterosporium ornithogali Klotzsch ex Cooke, Grevillea 5(35): 123 (1877)!. 

≡ Heterosporium ornithogali Klotzsch, Herb. viv. myc., Cent. I, No. 69 (1832)!, nom. inval. 

76

= Heterosporium ornithogali var. minus Bäumler, in Beck & Zahlbruckner, Ann. K.K. 

Naturhist. Hofmus. 12(2): 81 (1897)!, as ‘f. minus’. [T: K]. 

Teleomorph: Davidiella ornithogali (J.E. Jacques) Crous & U. Braun, in Braun, Crous, 

Dugan, Groenewald & de Hoog, Mycol. Progr. 2(1): 10 (2003)!. 

Lit.: ELLIS (1976: 338), DAVID (1995b, 1997: 52). 

Ill.: ELLIS (1976: 337, Fig. 256 B), DAVID (1995b: 1, Fig.; 1997: 53–54, Figs 14 A–F, 15). 

Host(s)/substrate(s) & distribution: on leaves of Ornithogalum and Gagea spp.; Europe (Czech 

Republic, France, Germany, Great Britain, Hungary, Italy, Netherlands, Poland, Romania, 

Serbia, Slovakia, Russia), Africa (Kenya, South Africa), North America (USA: IL, PA, WA). 

oryzae I. Miyake → miyakei. 

oryzae Sacc. & P. Syd., Syll. fung. 14: 1082 (1899)!. 

T: on rotten culms of Oryza sativa (Poaceae), Italy. 

≡ Helminthosporium maculans Catt., Arch. Triennale Lab. Bot. Crittog. 2/3: 122 (1879)!. 

≡ Cladosporium maculans (Catt.) Sacc., Syll. fung. 4: 365 (1886)!, nom. illeg., homonym, non 

C. maculans Schwein., 1832. 

Lit.: LINDAU (1907: 814), FERRARIS (1912: 339), PADWICK (1950: 170). 

Ill.: CATTANEO (1879: Tab. 14, Figs 7–9). 

Notes: OUDEMANS (1923) cited Jasminum fruticans as host species, but this record presumably 

refers to C. maculans Schwein. 

osterici Ces., in herb. 

On Ostericum verticillare (= Peucedanum verticillare) (Apiaceae), Italy (B). 

= Passalora depressa (Berk. & Broome) Sacc., Nuovo Giorn. Bot. Ital. 8: 187 (1876). 

oudemansii Kupka, Oesterr. Bot. Z. 67: 157 (1918)!, nom. nov. 

T: on leaves of Phragmites communis (= P. australis) (Poaceae), Netherlands. 

≡ Cladosporium phragmitis J. Opiz ex Oudem., Ned. Kruidk. Arch., Ser. II, 6: 57 (1892)!, non 

C. phragmitis J. Opiz, 1852. 

= Deightoniella arundinacea (Corda) S. Hughes, Mycol. Pap. 48: 29 (1952)!. 

Lit.: LINDAU (1907: 814), DE VRIES (1952: 96), DAVID (1997: 137). 

Notes: see C. phragmitis. 

ovorum Pidopl. & Deniak, in Pidoplichko, Gribnaja Flora Grubych Kormov: 272 (1953)!, 

nom. inval. 

T: on a hen’s egg, Ukraine. 


oxycocci Shear, Bull. Torrey Bot. Club 34(6): 306 (1907)!. 

T: on living leaves of Vaccinium macrocarpon (Ericaceae), Canada, Nova Scotia, Arichat, 21 

Jun. 1902, C.L. Shear (BPI 427299; 1492 C.L.S.). 

≡ Stenella oxycocci (Shear) U. Braun & K. Schub., comb. nov. 

Lit.: SACCARDO (1913: 1368). 

oxysporum Berk. & M.A. Curtis, in Berkeley, J. Linn. Soc., Bot. 10: 362 (1869)!. 

T: on dead leaves of Passiflora sp. (Passifloraceae), Cuba, C. Wright, Fungi cubensis 

Wrightiani, No. 489 (K 121562: holotype). 

= Cladosporium subtile Rabenh., F. eur., Ed. nov., Ser. 2, Cent. 24, No. 2364 (1876)!, nom. 

nud. 

= Cladosporium sorghi S.R. Chowdhury, Sydowia 23(6): 50 ‘1969’ (1970), syn. nov. [T: 

IMI]. 

Lit.: SACCARDO (1886: 363), ELLIS (1971: 312), MCKEMY & MORGAN-JONES (1991b), 

DAVID (1997: 81), BAGYANARAYANA & BRAUN (1999: 13), HO et al. (1999: 137), DE HOOG 

et al. (2000: 589), SCHUBERT & BRAUN (2004: 308–309). 

Ill.: ELLIS (1971: 313, Fig. 216 A), MCKEMY & MORGAN-JONES (1991b: 399, Pl. 1; 401, Fig. 

1; 403, fig. 2), DAVID (1997: 62, Fig. 17 A–E), HO et al. (1999: 138, Fig. 39), DE HOOG et al. 

(2000: 589–590, Figs). 

77

Host(s)/substrate(s) & distribution: on dead parts of leaves and stems of herbaceous and 

woody plants and other organic matter; common and widespread in the tropics and subtropics. 

paeoniae Pass., in Thümen, Herb. myc. oec., Fasc. IX, No. 416 (1876)! and Just’s Bot. 

Jahresber. 4: 235 (1876)!. 

T: on living leaves of Paeonia edulis (= P. lactiflora) (Paeoniaceae), Italy, Thüm., Herb. myc. 

oec. 416 (e.g., M-57753) and Thüm., Mycoth. univ. 670 (e.g., HAL; M-57752). 

= ? Cladosporium chlorocephalum (Fresen.) E.W. Mason & M.B. Ellis, Mycol. Pap. 56: 123 

(1953)!. 

Lit.: SACCARDO (1886: 362), LINDAU (1907: 822), FERRARIS (1912: 348), LIND (1913: 524), 

DE VRIES (1952: 94), MCKEMY & MORGAN-JONES (1991a). 

Notes: Excluded, but generic affinity not yet clear. 

paeoniae [Pass.] var. paeoniae-anomalae Sacc., Syll. fung. 4: 362 (1886)!. 

T: on faded leaves of Paeonia anomala (Paeoniaceae), Russia, Sibiria. 

≡ Cladosporium paeoniae (“paeoniae-anomalae”) Sacc., Michelia 2(6): 148 (1880)!. 

= ? Cladosporium chlorocephalum (Fresen.) E.W. Mason & M.B. Ellis, Mycol. Pap. 56: 123 

(1953)!. 

Lit.: LINDAU (1907: 822), MCKEMY & MORGAN-JONES (1991a). 

‘paeoniae-anomalae’ Sacc., Michelia 2(6): 148 (1880)!. 

Notes: Michelia (1880) refers to C. paeoniae Pass. reported on Paeonia anomala. 

pallidum Berk. & M.A. Curtis, Proc. Amer. Acad. Arts 4: 127, 1858 (1860)!. 

T: on leaves of an unidentified host, Nicaragua, Greytown, 1856, U.S. Pac. Ex. 354 (K). 

≡ Cercospora pallida (Berk. & M.A. Curtis) Cooke, Grevillea 17(81): 21 (1888)!, nom. illeg., 

homonym, non C. pallida Ellis & Everh., 1887. 

Lit.: SACCARDO (1886: 361; 1892: 638), CHUPP (1954: 609), CROUS & BRAUN (2003: 304). 

pallidum (Oudem.) H. Zhang & Z.Y. Zhang, Proceedings of Phytopathological Symposium 

Organized by Phytopathology Laboratory of Yunnan Province 2: 306 (1998), nom. illeg., 

homonym, non C. pallidum Berk. & M.A. Curtis, 1860. 

T: from soil, the Netherlands, Bussum. 

≡ Hormodendrum pallidum Oudem., Arch. Néerl. Sci. Exact. Nat. 7: 293 (1902). 

Lit.: SACCARDO (1906: 581), ZHANG et al. (2003: 233). 


Notes: ZHANG et al. (2003) recorded this species from China on Antirrhinum majus and 

Enkianthus quinqueflorus. In any case, the description and illustration of ‘C. pallidum’ from 

China do not agree with the original description of Hormodendrum pallidum by Oudemans, 

who described branched (dendroid) conidiophores and much wider conidia, 12−20 x 5−8 µm. 

palmetto Ger. – an error in VANEV & TASEVA (1990). 

Notes: In Index fungorum (CABI page) only Helminthosporium palmetto W.R. Gerard, 

Grevillea 17: 68 (1889), is listed. This species was transferred to the genus Pleurophragmium 

by HUGHES (1958) and later to Spiropes by ELLIS (1968). 

pannosum Cooke, Grevillea 12(61): 24 (1883)!. 

T: on Musa sp. (Musaceae), USA, South Carolina, H.W. Ravenel, No. 3056 (K 121564: 

holotype). 



Host(s)/substrate(s) & distribution: on Musa sp.; North America (USA). 

Notes: Under description (l.c.) of Chaetophoma musae Cooke, Grevillea 12: 24 (1883): 

“Effusa atro-fusca, pannosa; hyphis intertextis, cladosporioideis, (Cladosporium pannosum 

Cooke); peritheciis (sic) globoso-depressis…fuscis…”. 

papyricola Berk. & Broome, Trans. Linn. Soc., Ser. 2, Bot., 2: 68 (1883)!, as ‘papyricolor’. 

T: on damp paper, Australia, Queensland, Brisbane, F.M. Bailey, no. 128 (K 121565: isotype). 

Lit.: SACCARDO (1886: 369, as ‘papyricolum’; 1897: 879). 

78

parasiticum Sorokīn, Mikol. ocherki: 30 (1871). 

T: on abdomen of Melolontha fullonis (Coleoptera), Russia, Saratow. 

Lit.: SACCARDO (1906: 681, 1913: 1370). 

Notes: In SACCARDO (1906) under Strumella parasitica C. Wize (l.c.), [Pilze des Cleon. 

punctiventr. Crac. 1905: 725, fig. 11 (Bull. Acad. Crac. )] “An Cladosporium parasiticum 

Sorok? (ubi?)”. Also cited in GUÉGUEN, Champ. paras. homme: 256 (1904). 

paulliniae Deighton, Mycol. Pap. 144: 54 (1979)!. 

T: on leaves of Paullinia pinnata (Sapindaceae), Ghana, Essipun, 9 May 1949, S.J. Hughes 

(IMI 37238a: holotype). 

≡ Stenella paulliniae (Deighton) K. Schub. & U. Braun, Mycol. Progr. 4(2): 107 (2005)!. 

Ill.: DEIGHTON (1979: 54, Fig. 28), SCHUBERT & BRAUN (2005a: 107, Fig. 7). 

pelliculosum Berk. & M.A. Curtis, in herb. 

On leaves of Lobelia puberulosa (Campanulaceae), USA, South Carolina, No. 1742 (K 

121567). 

≡ Passalora lobeliae-cardinalis (Schwein.) U. Braun & Crous, in Crous & Braun, Mycosphaerella 


Biodiversity Ser. 1: 254 (2003). 

Lit.: COOKE (1889), SACCARDO (1892: 602, 1895: 621), SCHUBERT & BRAUN (2005b). 

Notes: COOKE (1889): “Scarcely appears to differ from Cladosporium effusum Berk. & M.A. 

Curtis, and does not seem to have been described”. 

penicilloides Preuss, in Sturm, Deutschl. Fl. 3(26): 31 (1848)!. 

T: on Tubercularia granulata and T. vulgaris, Germany, C.G.T. Preuss, Nr. 396 (B: holotype). 

Lit.: SACCARDO (1886: 369), LINDAU (1907: 807), FERRARIS (1912: 351), OUDEMANS (1920: 

Betula verrucosa as host, 1921: Prunus domestica as host), NANNIZZI (1934: 407), HEUCHERT 

et al. (2005: 56). 

Ill.: PREUSS (1848: Tab. 16). 

persicum, in herb. 

On Prunus persica (Rosaceae), Japan, Iwate, Morioka, 27 Oct. 1927, K. Togashi (BPI 

427388). 

= Stenella persicae T. Yokoy. & Nasu, Mycoscience 41: 92 (2000)!. 

perfragile R.F. Castañeda, unpublished name. CABI-page, Kirk et al (n. d.). 

pericarpium Cooke, Grevillea 12(61): 31 (1883)!. 

T: on husks of walnut (Juglans nigra, Juglandaceae), USA, South Carolina, Aiken, Rav., F. 

amer. exs. 597 (e.g., BPI 427378; K; NY; PH 01020429: syntypes). 

Lit.: SACCARDO (1886: 353). 

peridermiicola, in herb. 

On Gymnosporangium nelsonii, Peridermium columnare (BPI). 

= Cladosporium galliicola Sutton, Mycol. Pap. 132: 37 (1973). 

perpusillum Sacc., Atti Reale Ist. Veneto Sci. Lett. Arti, Ser. 6, 2(3): 449 (1883–1884)!. 

T: on culms of Ammophila sp. (Poaceae), France, Vendée, Malbranche, no. 78. 

Lit.: SACCARDO (1886: 364). 

personatum Berk. & M.A. Curtis, in Berkeley, Grevillea 3(27): 106 (1875)!. 

T: on leaves of Arachis hypogaea (Fabaceae), USA, Santee River, Ravenel, No. 1612 (IMI 

104553; K). 

≡ Cercospora personata (Berk. & M.A. Curtis) Ellis & Everh., J. Mycol. 1: 63 (1885)!. 

≡ Cercosporiopsis personata (Berk. & M.A. Curtis) Miura, Flora of Manchuria and East 

Mongolia, III. Cryptog. Fungi: 529 (1928), as ‘personatum’. 

≡ Passalora personata (Berk. & M.A. Curtis) S.A. Khan & M. Kamal, Pakistan J. Sci. Res. 

13: 188 (1961). 

≡ Cercosporidium personatum (Berk. & M.A. Curtis) Deighton, Mycol. Pap. 112: 71 (1967)!. 

79

≡ Phaeoisariopsis personata (Berk. & M.A. Curtis) Arx, Proc. Kon. Ned. Akad. Wetensch. C, 

86(1): 43 (1983). 

≡ Passalora personata (Berk. & M.A. Curtis) Poonam Srivast., J. Liv. World 1(2): 117 

(1994)!, comb. inval. et nom. illeg. 

= Septogloeum arachidis Racib., Z. Pflanzenkrankh. 8: 66 (1898). 

= Cercospora arachidis Henn., Hedwigia 41: 18 (1902). [T: N]. 

Teleomorph: Mycosphaerella berkeleyi W.A. Jenkins, J. Agric. Res. 56: 330 (1938). 

Lit.: SACCARDO (1886: 439), SIVANESAN (1984: 219), CROUS & BRAUN (2003: 317). 

personatum [Berk. & M.A. Curtis] f. arachidis-hypogaeae Thüm., Herb. myc. oec., Fasc. 

XIII, No. 608 (1878)!, nom. nud. 

T: on living leaves of Arachis hypogaea (Fabaceae), USA, South Carolina, Thüm., Herb. myc. 

oec. 608. 

Notes: Original material could not be examined, but this forma is very probably synonymous 

with Passalora personata. 

personatum [Berk. & M.A. Curtis] var. cassiae Thüm., Mycoth. univ., Cent. XX, No. 1964 

(1881)!. 

T: on living leaves of Cassia occidentalis (Caesalpiniaceae), USA, South Carolina, Aiken, 

1876, H.W. Ravenel, Thüm., Mycoth. univ. 1964 (e.g., BPI; G; HBG; HAL; S; W: syntypes). 

= Cercospora personata var. cassiae-occidentalis Berk. & M.A. Curtis, Grevillea 3(27): 106 

(1875). [T: IMI 104555; K]. 

= Cercospora occidentalis Cooke, Hedwigia 17: 39 (1878). [T: BPI 438975; K; IMI 92632a; 

Ellis, N. Am. F. 642; Rav., F. amer. exs. 65]. 

≡ Ramularia cassiicola Heald & F.A. Wolf, U.S.D.A. Bur. Pl. Industr. Bull. 226: 101 (1912), 

nom. nov., non Ramularia occidentalis Ellis & Kellerm., 1883. 

≡ Phaeoramularia occidentalis (Cooke) Deighton, in Ellis, More Dematiaceous Hyphomycetes: 

322 (1976)!. 

≡ Passalora occidentalis (Cooke) U. Braun, Schlechtendalia 5: 70 (2000)!. 

= Cercospora sphaeroidea Speg., Anales Soc. Ci. Argent. 9: 279 (1880). [T: IMI 206774; LPS 

911]. 

≡ Phaeoisariopsis sphaeroidea (Speg.) L.G. Br. & Morgan-Jones, Mycotaxon, 4: 303 (1976). 

= Cercospora cassiicola Roum., F. sel. gall. exs., No. 4486 (1886), as ‘cassiaecola’, nom. 

nud. [T: LEP]. 

= Cercospora paulensis Henn., Hedwigia 48: 18 (1909). [T: B]. 

= Cercospora iponemensis Henn., Hedwigia 48: 18 (1909). [T: B]. 

≡ Cercosporina iponemensis (Henn.) Sacc., Syll. fung. 25: 906 (1931). 

= Cercospora occidentalis Ellis & Kellerm., U.S.D.A. Bur. Pl. Industr. Bull. 226: 101 (1912), 

nom. illeg., homonym, non C. occidentalis Cooke, 1878. 

= Cercospora somalensis Curzi, Boll. Staz. Patol. Veg. Roma, N.S., 12: 158 (1932). 

Lit.: ELLIS (1976: 322), CROUS et al. (2000), CROUS & BRAUN (2003: 294). 

peruamazonicum Matsush., Matsushima Mycol. Mem. 7: 47 (1993)!. 

T: on rotten leaves, Peru. 

pestis Thüm., Herb. myc. oec., Fasc. IX, No. 419 (1876)! and Oesterr. Bot. Z. 27: 12 (1877)!. 

T: on living leaves of Vitis vinifera (Vitaceae), Austria, Klosterneuburg, Krems, 1876, von 

Thümen, Thüm., Herb. myc. oec. 419 (e.g., M). 

= Passalora dissiliens (Duby) U. Braun & Crous, in Crous & Braun, Mycosphaerella and its 


164 (2003)!. 


Notes: see also C. roesleri. 

phaseoli Fuss, Archiv Verein siebenb. Landesk., N.F., 14(2): 431 (1878)!, nom. nud. 

On overwintered fruits of Phaseolus vulgaris (Fabaceae), Romania, Giresau, Fuss. 


80

phlei (C.T. Greg.) G.A. de Vries, Contr. Knowl. Genus Cladosporium: 49 (1952)!. 

T: on Phleum pratense (Poaceae), USA, New York, Ithaca, greenhouse, 6 Apr. 1918, Gregory 

10498 (CUP: holotype). 

≡ Heterosporium phlei C.T. Greg., Phytopathology 9: 580 (1919)!. 

= ? Cladosporium herbarum var. macrosporum Lagière, Ann. École Natl. Agric. Grignon, Sér. 

3, 5: 159 (1946)!. 

Lit.: MATSUSHIMA (1975: 36), ELLIS (1976: 334), ELLIS & ELLIS (1985: 508), DAVID (1988d, 

1997: 90), HO et al. (1999: 139). 

Ill.: GREGORY (1919: 579, Fig. 2), MATSUSHIMA (1975: Pl. 53, Figs 1–2), ELLIS (1976: 335, 

Fig. 254), DAVID (1988d: 1, Fig.; 1997: 91, Fig. 23), HO et al. (1999: 141, Figs 40–41). 

Host(s)/substrate(s) & distribution: on Phleum pratense; Asia (Armenia, Japan), Europe 

(Byelorussia, Germany, Denmark, Eire, Estonia, Great Britain, Norway, Russia, Sweden), 

North America (Canada: Que., USA: most states), New Zealand. 

phlei-pratensis Sawada, Bull. Gov. Forest Exp. Sta. 105: 96 (1958)!, as ‘phlei-pratense’, nom. 

inval. 

T: on leaves of Phleum pratense (Poaceae), Japan, Tohoku District, 27 Jul. 1940, Iizuka 

(10516) and 18 Jul. 1941, Ikeda (10440). 

Notes: In the original diagnosis two collections were mentioned but the author did not 

designate a type. 

phoenicis ‘Fautrey’, Rev. Mycol. (Toulouse) 13: 133 (1891)!. 

Lit.: SACCARDO (1892: 604). 

Notes: This species was not described by Fautrey, rather it refers to Roumeguère, see next 

entry. 

phoenicis Roum., F. sel. gall. exs., Cent. 58, No. 5798 (1891)! and Rev. Mycol. (Toulouse) 

13: 133 (1891)!, as ‘phaenicis’. 

T: on leaves of Phoenix tenuis (Arecaceae), France, Toulouse, 1891, G. Machado, Roum., F. 

sel. gall. exs. 5798 (e.g., B; FH). 



phragmitis J. Opiz, in Opiz, Seznam: 117 (1852), nom. nud., non C. phragmitis J. Opiz ex 

Oudem., 1892. 

T: on leaves of Phragmites communis (= P. australis) (Poaceae), Czech Republic (PRM). 

= Heterosporium phragmitis Sacc., in Saccardo & Roumeguère, Rev. Mycol. (Toulouse) 6: 37 

(1884), as ‘(Opiz?) Sacc.’. 


phragmitis J. Opiz ex Oudem., Ned. Kruidk. Arch., Ser. 2, 6(1): 57 (1892)!. 

T: on leaves of Phragmites communis (= P. australis) (Poaceae), Netherlands, Bien de 

Campagne Zorgvlied, near Haye, Jul. 1889, C.E. Destrée. 

≡ Cladosporium oudemansii Kupka, Oesterr. Bot. Z. 67: 157 (1918)!. 

= Helminthosporium arundinaceum Corda, Icon. fung. 3: 10 (1839), as ‘Helmisporium’. 

≡ Napicladium arundinaceum (Corda) Sacc., Syll. fung. 4: 482 (1886)!. 

≡ Deightoniella arundinacea (Corda) S. Hughes, Mycol. Pap. 48: 29 (1952)!. 

Lit.: DE VRIES (1952: 96), DAVID (1997: 137). 

Notes: DAVID (1997): ‘OUDEMANS (1892) provided a description of what he took to be Opiz’s 

species. This description, also given by LINDAU (1907), is of an entirely different fungus, as 

noted by KUPKA (1918) and DE VRIES (1952).’. 

phyllachorae M.B. Ellis, More Dematiaceous Hyphomycetes: 332 (1976)!. 

T: on Catacauma apoensis (= Phyllachora) (Phyllachoraceae) on leaves of Ficus nervosa 

(Moraceae), Philippines, Samar, Mar.–Apr. 1914, M. Ramos (Bur. Sc. 17616). 

≡ Monotospora parasitica Syd. & P. Syd., Ann. Mycol. 15: 263 (1917)!, non Cladosporium 

parasiticum Sorokīn, 1891. 


81

Ill.: ELLIS (1976: 333, Fig. 251). 

Notes: A further collection for Monotospora parasitica is mentioned in the original diagnosis 

on Phyllachora pseudis on leaves of Ficus nota, Phillipines, Luzon, Prov. Laguna, San 

Antonio, Oct. 1915, M. Ramos (Bur. Sc. 23781). 

phyllogenum K. Schub., sp. nov. 

T: on living leaves of Ulmus laevis, Germany, Sachsen-Anhalt, Halle (Saale), botanical 

garden, 9 Jul. 2004, K. Schubert (HAL 1845 F). 

Host(s)/substrate(s) & distribution: on Ulmus laevis; Europe (Germany). 

phyllophilum McAlpine, Agric. Gaz. New South Wales 7: 153 (1896)!. 

T: on leaves and twigs of Prunus persica (= Persica vulgaris) (Rosaceae) infected with and 

deformed by Taphrina deformans (= Exoascus deformans), Australia, Victoria, Armadale, 16 

Feb. 1896, D. McAlpine (VPRI 2490: lectotype). 

= Cladosporium exoasci Ellis & Barthol., in Shear, F. columb., Cent. XV, No. 1493 (1901)!, 

nom. nud. 

= Cladosporium exoasci Lindau, in Rabenhorst, Krypt.-Fl., ed. 2, 1(8): 808 (1907)!. 

Lit.: MCALPINE (1902: 100), SACCARDO (1906: 575), BRAUN (2001: 53), HEUCHERT et al. 

(2005: 36–40). 

Ill.: MCALPINE (1902: Figs 87, 88), HEUCHERT et al. (2005: 37–38, Figs 13–14; Pl. 2, Fig. 

12). 

Host(s)/substrate(s) & distribution: on species of Taphrina, including T. cerasi, T. communis, 

T. deformans and T. pruni on Prunus s. lat. species; Asia (Armenia, Georgia, Kazakhstan, 

Uzbekistan), Australia, Europe (Czech Republic, France, Germany, Romania, Switzerland), 


pilicola Richon, Cat. Champ. Marne: 452 (1889). 

T: on dry stems of Galium mollugo (Rubiaceae), France, Marne, Soulanges. 

Lit.: SACCARDO (1892: 602, as ‘pilicolum’). 

pipericola R.A. Singh & Shankar, Mycopathol. Mycol. Appl. 43(1): 110 (1971)!, as ‘pipericolum’. 

T: on living leaves of Piper betle (Piperaceae), India, Uttar Pradesh, Varanasi, 15 Jan. 1965, 

G. Shankar (MSP no. 342; IMI 116933). 

Lit.: DAVID (1988e). 

Ill.: SINGH & SHANKAR (1971: 111, Pl. 1, Figs 3–4), DAVID (1988e: 1, Fig.). 

Host(s)/substrate(s) & distribution: on Piper betle; Asia (India). 

piricularioides Dearn. & House, Circ. New York State Mus. 24: 57 (1940), nom. inval. 

T: on leaves of Panicum boreale (Poaceae), USA, New York, Essex Co., Newcomb, 17 Aug. 

1924, H.D. House (NYS 2365: holotype; DAOM 5741, NY: isotypes). 

= Passalora fusimaculans [(G.F. Atk.) U. Braun & Crous] var. barretoana U. Braun & 

Crous, in Crous & Braun, Mycosphaerella and its anamorphs: 1. Names published in 

Cercospora and Passalora. CBS Biodiversity Ser. 1: 453 (2003). 

Lit.: SCHUBERT & BRAUN (2005a: 104–105). 

Notes: without latin diagnosis. In NYS label: C. piricularioidis. 

piriforme Reichert – SACCARDO (1972: 1338). 

Notes: see C. pyriforme. 

pirorum Berk. – LINDAU (1907: 779). 

Notes: see C. pyrorum Berk. 

pisi Cugini & Macch., Boll. Reale Staz. Agric. Modena, N.S., 10(1890): 104 (1891). 

T: on legumes of Pisum sativum (Fabaceae), Italy, Vaciglio near Modena. 

= ? Cladosporium herbarum (Pers.: Fr.) Link, Ges. Naturf. Freunde Berlin Mag. Neuesten 


Lit.: SACCARDO (1892: 601), LINDAU (1907: 825), FERRARIS (1912: 349), OUDEMANS (1921: 

on Vicia faba), SNYDER (1934: 890), MORGAN-JONES & MCKEMY (1992: 11). 

Ill.: CUGINI & MACCHIATI (1891: Tab. 5). 

Notes: The status of this taxon is uncertain (FARR et al. 1989). 

82

pisicola W.C. Snyder, Phytopathology 24: 899 (1934)!, as ‘pisicolum’. 

T: on Pisum sativum (Fabaceae), USA, California. [UBC 41245, W. Snyder collector, Dec. 

1940, infected pea pod: topotype ?]. 

≡ Cladosporium cladosporioides [(Fresen.) G.A. de Vries] f. sp. pisicola (W.C. Snyder) G.A. 

de Vries, Contr. Knowl. Genus Cladosporium: 61 (1952)!. 

Lit.: FARR et al. (1989: 629). 

Ill.: SNYDER (1934: 893, Fig. 2). 

pithecolobii Kulhara & Singh. 

On Pithecolobium dulce (Mimosaceae), India, M. Pradesh, Jabalpur. 

platycodonis Z.Y. Zhang & H. Zhang, in Zhang, Zhang & Li, Mycosystema 19(3): 308 

(2000)!. 

T: on living leaves and petals of Platycodon grandiflorus (Campanulaceae), China, 

Heilongjiang, Monte Maoer, 5 Sept. 1992, H. Li & Y.X. Wang (MHYAU 07826: holotype). 

Ill.: ZHANG, ZHANG & LI (2000: 309, Fig. 1), ZHANG et al. (2003: 145, Fig. 97). 

Notes: ZHANG et al. (2003) reported the species also on Adenophora stricta from Jilin, China. 

polygonati M.B. Ellis, More Dematiaceous Hyphomycetes: 338 (1976)!. 

T: on leaves of Polygonatum sp. (Convallariaceae), Ireland, Wicklow Co., Eriskerry, Bray, 

Powerscourt, Oct. 1965, C.H. Dickinson (IMI 116694: holotype). 

≡ Heterosporium polygonati (M.B. Ellis) Arx, Genera Fungi Sporul. Pure Cult., ed. 3: 305 

(1981)!. 

Lit.: DAVID (1997: 57). 

Ill.: ELLIS (1976: 337, Fig. 256 C), DAVID (1997: 38, Fig. 8 D–G; 56, Fig. 16). 

Host(s)/substrate(s) & distribution: on Polygonatum sp.; Europe (Ireland). 

polygonaticola Z.Y. Zhang & W.Q. Pu, in Zhang, Liu, Wei & He, Plant Diseases and Their 

Control: 105 (1998)!. 

T: on living leaves of Polygonatum cirrhifolium (Liliaceae s. lat.), China, Jiangxi, Lushan, 6 

Oct. 1980, J.Y. Li & T.Y. Zhang, No. 41551 (MHYAU 03949: holotype). 

Ill.: ZHANG, LIU, WEI & HE (1998: 106, Fig. 3), ZHANG et al. (2003: 146, Fig. 98). 

Notes: ZHANG et al. (2003) cited three hosts, viz., Polygonatum cirrhifolium, P. cyrtonema 

(China, Hubei) and P. sibiricum (China, Yunnan). 

polymorphosporum R.F. Castañeda & W.B. Kendr., Univ. Waterloo Biol. Ser. 35: 24 (1991)!. 

T: on stem of an unidentified grass, Cuba, Pinar del Río, Sandino, 24 Mar. 1990, R.F. 

Castañeda (INIFAT C90/139: holotype). 


polymorphum Peyl, Lotos 15: 18 (1865)! and Hedwigia 5: 60 (1866)!. 

T: on fruits of Pyrus sp. (Rosaceae), Czech Republic, Kačina, near Neuhof, 1864. 

= Helminthosporium pyrorum Lib. (p.p.), Pl. crypt. ard., Fasc. 2, No. 188 (1832). [T: DAOM]. 

≡ Fusicladium pyrorum (Lib.) Fuckel, Jahrb. Nassauischen Vereins Naturk. 23–24: 357 

‘1869’ (1870)!, as ‘Fusicladium pyrinum’. 

≡ Passalora pyrina (Lib.) Sacc., Michelia 1: 537 (1879). 

≡ Megacladosporium pyrorum (Lib.) Vienn.-Bourg., Les Champignons parasites des plantes 

cultivèes 1: 489 (1949), as ‘Megacladosporium pirinum’. 

= Arthrinium pyrinum Wallr., Fl. crypt. Germ. 2: 163 (1833). [T: STR; IMI 68300]. 

= Fusidium pyrinum Corda, Icon. Fung. 1: 3 (1837). [T: PRM]. 

= Fusicladium virescens Bonord., Handb. Mykol.: 80 (1851)!. [T: Bonorden, 1851: Fig. 94 

(iconotype)]. 

= Fusicladium fuscescens Rabenh., Bot. Zeitung (Berlin) 15: 430 (1857)!. [T: Rabenh., Herb. 

mycol. 588, e.g., HAL; HBG]. 

= Passalora pomi G.H. Otth, Mitteil. Naturf. Ges. Bern 1868: 66 (1868). [T: BERN]. 

= Fusicladium pyrorum [(Lib.) Fuckel] var. cladophilum Ellis & Everh., N. Am. F., No. 2791 

(1892)!. [T: Ellis & Everh., N. Am. F. 2791, e.g., BPI; M; NY]. 

83

= Cercospora porrigo Speg., Anales Mus. Nac. Buenos Aires, Ser. 2, 6: 341 (1899). [T: LPS 

934]. 

= Fusicladium pyrorum [(Lib.) Fuckel] f. carpophila Sacc., Mycoth. ital., No. 992 (1901)!. [T: 

Sacc., Mycoth. ital. 992, e.g., B]. 

= Acrotheca dearnessiana Sacc., Ann. Mycol. 10: 314 (1912). [T: Barthol., F. columb. 5001, 

e.g., IMI 7073]. 

≡ Fusicladium dearnessianum (Sacc.) M.B. Ellis, in herb. 

Teleomorph: Venturia pyrina Aderh., Landw. Jahrb. 25: 875 (1896)!, as ‘pirina’. 

Lit.: LINDAU (1907: 781), OUDEMANS (1921), SCHUBERT et al. (2003: 82). 

polysporum Link, in Willd., Sp. pl. 6(1): 40 (1824)!. 

T: on rotten wood, Germany, Berlin, Link (B). 

= Trichoderma globosum Schwein., Syn. fung. Carol. sup.: 77 (1822): Fr., Syst. Mycol. 3(1): 

215 (1829). [T: PH; BPI; K; UPS]. 

≡ Oidium inquinans Schwein., Trans. Amer. Philos. Soc., N.S., 4(2): 286 (1832), nom. nov. 

≡ Torula inquinans (Schwein.) Sacc., Syll. fung. 4: 251 (1886)!. 

≡ Streptothrix globosa (Schwein.: Fr.) S. Hughes, Canad. J. Bot. 31: 606 (1953)!. 

≡ Conoplea globosa (Schwein.: Fr.) S. Hughes, Canad. J. Bot. 36: 755 (1958)!. 

= Steptothrix atra Berk. & M.A. Curtis, in Berkeley, Grevillea 3(27): 107 (1875)!. 

= Strumella coryneoidea Sacc. & G. Winter, in Rabenhorst, F. eur., Cent. XXX, No. 2984 

(1883) and Hedwigia 22(11): 175 (1883)!. [T: CUP]. 

= Trichosporium densum P. Karst., Hedwigia 23(4): 59 (1884)!. 

= Streptothrix pereffusa Sumst., Mycologia 6: 34 (1914). 

Lit.: SACCARDO (1886: 354), LINDAU (1907: 831), DE VRIES (1952: 96). 

polytrichorum Reichardt, Verh. K.K. Zool.-Bot. Ges. Wien 27: 844, 1877 (1878)!. 

T: on Polytrichum formosum (Polytrichaceae), Austria, Tobelbad, near Graz, Sept. 1875, H.W. 

Reichardt. 

Notes: REICHARDT (1878): “an status conidiophorus Lizoniae emperigoniae Ces. ?”. 

populicola K. Schub. & U. Braun, sp. nov. 

T: on Populus tremula (Salicaceae), Germany, Schleswig-Holstein, Missunde, Schlei ferry, 

river bank, 30 Aug. 2004, U. Braun (HAL 1833 F: holotype). 

Host(s)/substrate(s) & distribution: on Populus tremula; Europe (Germany). 

porophorum Matsush., Icones Microfungorum a Matsushima Lectorum: 36 (1975)!. 

T: from seeds of Raphanus sativus (Brassicaceae), Japan, Feb. 1969 (Matsush. herb. 2578). 

= Alternaria malorum (Rühle) U. Braun, Crous & Dugan, in Braun, Crous, Dugan, Groenewald 

& de Hoog, Mycol. Progr. 2(1): 5 (2003)!. 

Lit.: HO et al. (1999: 134). 

Notes: see C. malorum Rühle. 

potebniae Pidopl. & Deniak, Mikrobiol. Zhurn. 5(2): 189, 194 (1938)!, as ‘potebnjae’, nom. 

inval. 

T: from rotting fruit of Malus sp. (Rosaceae), Ukraine. 



praecox (Niessl) U. Braun, Schlechtendalia 5: 34 (2000)!. 

T: on leaves of Tragopogon orientalis (Asteraceae), Czech Republik, ‘pr. Bistenz ad Brunnam 

Moraviae’, May, G. de Niessl, Rabenh., F. eur. 1166 (B, HAL, HBG: syntypes). 

≡ Fusicladium praecox Niessl, in Rabenhorst, F. eur., Ed. Nov., Ser. II, No. 1166 (1868)! and 

Hedwigia 7: 124 (1868)!. 

Ill.: BRAUN (2000: 33, Fig. 3). 

Host(s)/substrate(s) & distribution: on Tragopogon orientalis; Europe (Czech Republic). 

profusum Desm. ex Sacc., Syll. fung. 10: 602–603 (1892)!. 

T: France, Desm., Pl. crypt. N. France 755 (e.g., PC). 

84

≡ Cladosporium profusum Desm., Pl. crypt. N. France, No. 755 (1836), nom. nud. 

≡ Cladosporium profusum Desm., in Rabenhorst, F. eur., Cent. 6, No. 578 (1863), nom. nud. 

Lit.: SACCARDO (1886: 370, 1892: 602), LINDAU (1907: 832), GONZÁLES-FRAGOSO (1927: 

211). 

Notes: LINDAU (1907): “...Ebensowenig war es mir möglich festzustellen, wo Rabenhorst die 

Diagnose veröffentlicht hat und ob dies überhaupt geschehen ist. Woher Saccardo die 

Diagnose hat, ist mir nicht bekannt. Cooke hat in seinem zitierten Aufsatz nur die Abbildung, 

nimmt aber im Text auf die Art keinen Bezug. Am besten läßt man sie vielleicht ganz fort”. 

GONZÁLES-FRAGOSO (1927: 211) provided a description and considered this species to be 

very close to C. herbarum. 

profusum [Desm. ex Sacc.] var. robustior Roum. & Pat., Rev. Mycol. (Toulouse) 5: Tab. 35, 

Fig. 6 (1883)!. 

prunicola McAlpine, Fungus Dis. Stone-fruit Trees Austral.: 100 (1902)!, as ‘prunicolum’. 

T: on partly decayed leaves of Prunus armeniaca (= Armeniaca vulgaris) (Rosaceae), 

Australia, Victoria, Armadale near Melbourne, Dec. 1899. 

Lit.: SACCARDO (1906: 575), BRAUN (2001: 53). 

Ill.: MCALPINE (1902: Figs 89, 90). 

Notes: Type material is not preserved in VPRI, but description is close to Cladosporium 

herbarum var. macrocarpum (J. Cunnington, in litt.) (BRAUN 2001). 

psammicola (Sacc.) Morgan-Jones & W.B. Kendr., Canad. J. Bot. 50(9): 1817 (1972)!. 

T: on dead leaves of Psamma arenaria (= Ammophila arenaria) (Poaceae), North Africa, 

Libya, Ras Carrac in Magna Syrte, 18 May (PAD: holotype). 

≡ Exosporium psammicola Sacc., in Saccardo & Trotter, Ann. Mycol. 11: 420 1913)!. 

Ill.: MORGAN-JONES & KENDRICK (1972: 1818, Fig. 1). 

psidiicola J.M. Yen, Bull. Trimestriel Soc. Mycol. France 95(3): 188 ‘1979’ (1980)!, as 

‘psidicolum’. 

T: on living leaves of Psidium guajava (Myrtaceae), Hong Kong, Tai-Yuan-Yu-Tsun, Hsin- 

Chiai, Kowloon, 13 Nov. 1971, Jo-min Yen, No. 71334 (LAM). 

Ill.: YEN (1980: 187, Fig. 2). 

psoraleae M.B. Ellis, Mycol. Pap. 131: 16 (1972)!. 

T: on living leaves of Psoralea corylifolia (Fabaceae), Myanmar (Burma), Mandalay, 25 Nov. 

1971, M. Thaung (IMI 163005: holotype). 

Lit.: ELLIS (1976: 344). 

Ill.: ELLIS (1972: 17, Fig. 16). 

Host(s)/substrate(s) & distribution: on Psoralea corylifolia; Asia (Myanmar). 

Notes: BILGRAMI et al. (1991) cited a record on living leaves of Gardenia turgida from India 

(U.P.), which seems to be very doubtful. 

puccinioides Cooke, Grevillea 5(33): 15 (1876)!. 

T: on under side of living leaves of an unidentified host plant, India, 1876, Colonel Hobsen, 

No. 57 (K 121568: holotype). 

≡ Prathigada puccinoides (Cooke) M.B. Ellis, in herb. 

≡ Pseudoasperisporium puccinioides (Cooke) K. Schub. & U. Braun, Fungal Diversity 

(2005), in press. 

Lit.: SACCARDO (1886: 361), SUBRAMANIAN (1971: 291). 

Ill.: COOKE (1876: Pl. 74, Fig. 11), SCHUBERT & BRAUN (2005b, Fig. 6). 

Notes: COOKE (1876): ‘This is certainly intermediate between Cladosporium and Helminthosporium.’. 

pulcherrimum Ellis & Everh., N. Am. F., Ser. 2, Cent. XXIX, No. 2877 (1893), nom. nud. 

T: on Carpinus sp. (Corylaceae), USA, Ontario, Ellis & Everh., N. Am. F. 2877. 

Lit.: CASH (1952: 69). 

85

pullulans (de Bary) Sacc., Syll. fung. 22: 1250 (1913)!. 

T: on fruits of Vitis vinifera (Vitaceae), France (ex-neotype: CBS 584.75). 

≡ Dematium pullans de Bary, Vergl. Morph. Biol. Pilze: 182 (1884). 

≡ Oidium pullulans (de Bary) Lindner, Wochenschr. Brauerei 15: 209–213 (1898). 

≡ Oospora pullulans (de Bary) Sacc., Syll. fung. 18: 499 (1906)!. 

≡ Aureobasidium pullulans (de Bary) G. Arnaud var. pullulans, Ann. Écol. Nat. Agric. Montpellier, 

N.S., 16: 39 (1918). 

≡ Pullularia pullulans (de Bary) Berkhout, Die schimmelgeschlachten Monilia, Oidium, 

Oospora en Torula: 55 (1923). 

≡ Hormonema pullulans (de Bary) Lagerb. & Melin ex/ in Robak, Nyt Mag. Naturvidensk 71: 

256 (1932). 

Notes: Cited by SACCARDO (l.c.) under Oidium erysiphoides [Fr.] f. cordiae Sacc., Ann. 

Mycol. 8: 339 (1910): “Hab. In foliis Cordiae suboppositae … Socium adest Cladosporium 

(Demat.) pullulans.”. In SACCARDO (1886), LINDAU (1907) and FERRARIS (1912) ‘Dematium 

pullulans de Bary & Löwenthal’ is cited as synonym of Cladosporium herbarum. For further 

synonyms and comments on Aureobasidium pullulans see SUBRAMANIAN (1971), DE HOOG & 

YURLOVA (1994) and YURLOVA et al. (1999). 

punctatum Dearn. & House → sarraceniae. 

punctatum (Sacc.) Sacc., Syll. fung. 4: 355 (1886)!. 

T: on Euonymus japonicus (Celastraceae), France, Rouen, Malbranche (PAD). 

≡ Cladosporium compactum [Sacc.] *punctatum Sacc., Michelia 2(7): 363 (1881)!. 

Lit.: SACCARDO (1886: 359). 

Notes: “Affine Cl. subcompacto Sacc.” (SACCARDO 1886). 

punctiforme Fuckel, F. rhen., Fasc. II, No. 116 (1863)!. 

T: on living leaves of Sanicula europaea (Apiaceae), Germany, ‘auf der Geis im Hattenheimer 

Wald’, Fuckel, F. rhen. 116 (e.g., HAL). 

≡ Cercospora punctiformis (Fuckel) G.A. de Vries, Contr. Knowl. Genus Cladosporium: 97 

(1952)!, nom. illeg., homonym, non C. punctiformis Sacc. & Roum., 1881. 

= Cercospora saniculae-europaeae E. Müll. & Arx, Phytopathol. Z. 24: 356 (1955)!. 

≡ Pseudocercospora saniculae-europaeae (E. Müll. & Arx) U. Braun & Crous, in Crous & 

Braun, Mycosphaerella and its anamorphs: 1. Names published in Cercospora and 

Passalora, CBS Biodiversity Ser. 1: 365 (2003)!. 

Lit.: FUCKEL (1870: 355), SACCARDO (1886: 362), LINDAU (1907: 825), OUDEMANS (1923). 

punctulatum Sacc. – listed by SACCARDO & BERLESE (1884: 100). 

Notes: See C. punctulatum Sacc. & Ellis. 

punctulatum Sacc. & Ellis, Michelia 2(8): 578 (1882)!. 

T: on leaves of Euonymus japonicus (Celastraceae), USA, New Jersey, Newfield, 17 Apr. 

1881, J.B. Ellis, no. 3585 (NY: lectotype). Isolectotypes: BPI 427402; PAD. 




FRAGOSO (1927: 207). 

Notes: “Cladosporio punctato Sacc. subaffine.” (SACCARDO 1886). 

punctulatum var. xylogenum Fairm., Proc. Rochester Acad. Sci. 6: 131 (1922)!. 

T: on the outside of a cigar box exposed to damp weather, USA, New York, Lyndonville, 14 

Dec. 1920, C.E. Fairman (associated with Epicoccum agyrioides Corda). 

putrefaciens – Z. Pflanzenkrankh. 4: 333 (1894). 

Notes: without author on Beta sp. – maybe an error and Clasterosporium putrefaciens (Fuckel) 

Sacc. was intended. 

pygmaeum Ellis & Everh. (in Exs.: Flora Sequoia Gigantea Region, No. 1235, nom. nud.). 

T: on Vitis californica (Vitaceae), USA, California, Amador Co., Pine Grove, Jul. 1893, G.E. 

86

Hansen (B; BPI 427408–427409; NY: syntypes). 

= Asperisporium minutulum (Sacc.) Deighton, in Ellis, More dematiaceous hyphomycetes: 

242 (1976). 

Lit.: SCHUBERT & BRAUN (2005b). 

pyllophilum – see phyllophilum. 

pyriforme Reichert, Bot. Jahrb. Syst. 56: 721 (1921)!, as ‘pyriformum’. 

T: on cladodes of Opuntia ficus-indica (Cactaceae), Egypt, near Bulak, 1822/25, Ehrenberg 

(B). 

Lit.: SACCARDO (1972: 1338, as ‘piriforme I. Reichert). 


‘pyrorum Berk.’, Gard. Chron. 1848: 398 (1848)!. 

Lit.: LINDAU (1907: 779), OUDEMANS (1921). 

Notes: LINDAU cited Gard. Chron. p. 398 (1848) for C. pirorum Berk. and listed it as synonym 

of Fusicladium dendriticum (Wallr.) Fuckel [= Fusicladium pomi (Fr.) Lind, Dan. fung.: 521 

(1913)]. The page concerned has been examined, but the name “Cladosporium pyrorum 

Berk.” was not found. 

qinghaiense T. Zhang & Z.Y. Zhang, in Zhang, Zhang & Liu, Proceedings of Phytopathological 

Symposium Organized by Phytopathology Laboratory of Yunnan Province 2: 285 

(1998)!, as ‘qinghaiensis’. 

T: on Pisum sativum (Fabaceae), China, Qinghai, Huzhu, 3 Aug, 1989, H. Li & T.F. Li, No. 

087 (MHYAU 03925: holotype). 

Ill.: ZHANG, ZHANG & LIU (1998: 286, Fig. 1), ZHANG et al. (2003: 148, Fig. 100). 

quitense Syd., Ann. Mycol. 37: 420 (1939)!. 

T: on leaves of Berberis schwerinii (Berberidaceae), Ecuador, Pichincha mountains near 

Quito, 11 Sept. 1937, H. Sydow (B 70-6694: lectotype). Isolectotypes: B 70-6693 and Syd., F. 

exot. exs. 1232, e.g., B 70-6695; BPI 427427; M-57728. 

≡ Stenella quitensis (Syd.) K. Schub. & U. Braun, Mycol. Progr. 4(2): 108 (2005)!. 


radians Sacc. & D. Sacc., in Saccardo & P. Sydow, Syll. fung. 16: 1059 (1902)!. 

T: on leaves of Abies pinsapo (Pinaceae), Italy, Padua, Apr. 1900, Sacc., Mycoth. ital. 787 (B 

70-6696; BPI 427428; HBG: syntypes). 

≡ Cladosporium radians Sacc. & D. Sacc., Mycoth. ital., Cent. VIII., No. 787 (1901)!, nom. 

nud. 

≡ Septonema radians (Sacc. & D. Sacc.) U. Braun & K. Schub., comb. nov. 

Lit.: LINDAU (1907: 812), FERRARIS (1912: 336). 

ramulosum Rab. – OUDEMANS (1924). 

Notes: An error. C. ramulosum Roberge ex Desm. was intended. 

ramulosum Reissek, Sitzungsber. Kaiserl. Akad. Wiss., Math.-Naturwiss. Cl. 7(2): 336 

(1851)!. 

T: on pollen of Pinus sylvestris (Pinaceae), Austria. 

Lit.: SACCARDO (1886: 370, 1913: 1370), LINDAU (1907: 833). 

Notes: REISSEK (1851): ‘(C. entoxylinum Corda var. ?)’. “An diversum a Cladosporio 

ramuloso Desm.?” (SACCARDO 1886). “Species omnino dubia, cfr. Lindau l.c.” (SACCARDO 

1913). See also C. ramulosum Roberge below. 

ramulosum Roberge ex Desm., Ann. Sci. Nat. Bot., Sér. 3, 18: 361 (1852)! nom. illeg., homonym, 

non C. ramulosum Reissek, 1851. 

T: on Populus alba (Salicaceae), France, Paris, Parc du Libisy, May 1851, Roberge [PC 1518: 

holotype; herb. Desmazières 2135 (PC): isotype]. 

≡ Fusicladium ramulosum Rostr., Tidsskr. Skovbr. 6: 294 (1883), nom. nov., as ‘(Roberge, in 

Desm.) Rostr.’. 

87

≡ Pollaccia ramulosa (Rostr.) Ondřej, Eur. J. Forest Pathol. 2: 143 (1972)!, nom. nov., as 

‘(Desm.) Ondřej’. 

= Fusicladium radiosum (Lib.) Lind, Ann. Mycol. 3: 429 (1905) var. radiosum. 

Lit.: SACCARDO (1886: 357), LINDAU (1907: 777), OUDEMANS (1920), BALDACCI & CIFERRI 

(1937: 61), RITSCHEL (2001), SCHUBERT et al. (2003: 85). 

Notes: see C. asteroma. 

raphanicola Opiz, Seznam: 117 (1852), nom. nud. 

T: on Raphanus ?, Czech Republic. 

Lit.: SACCARDO (1886: 370, as ‘raphanicolum’), LINDAU (1907: 832), KUPKA (1918: 156). 

Notes: Type is not preserved at PRM. KUPKA (1918): “Von dem variablen C. herbarum kaum 

verschieden...“. 

rectangulare K. Schub. & U. Braun, Sydowia 56(2): 309 (2004). 

T: on still living and fading leaves of Epidendrum prismatocarpum (Orchidaceae), from ex 

England, intercepted at Hoboken, New Jersey, USA, 29 Sept. 1941, D. P. Limber (BPI 

427292: holotype, as ‘C. orchidearum’). 


Host(s)/substrate(s) & distribution: on Epidendrum prismatocarpum; North America (USA). 

rectum Preuss, in Sturm, Deutschl. Fl. 3(26): 29 (1848)!. 

T: on the innerside of bark of Pinus (Pinaceae), Germany, near Hoyerswerda (B). 

= Helminthosporium fasciculare Corda, Icon. fung. 1: 14 (1837)!, as ‘Helmisporium’. [T: 

PRM]. 

≡ Septonema fasciculare (Corda) S. Hughes, Canad. J. Bot. 36: 803 (1958)!. 

= Dendryphion pini Höhn., Sitzungsber. Kaiserl. Akad. Wiss., Math.-Naturwiss. Cl., Abt. 1, 

116: 153 (1907). [T: FH]. 

Lit.: SACCARDO (1886: 354, 374; 1906: 577), LINDAU (1907: 810). 

rederse, in herb. HBG (without author and description). 

An error on the web-site of the herbarium HBG and on the label of the collection in HBG// 

misinterpretation of the hand writing, see hederae. 

resinae (Lindau) G.A. de Vries (f. resinae), Antonie van Leeuwenhoek J. Microbiol. Serol. 

21: 167 (1955)!. 

≡ Hormodendrum resinae Lindau, in Rabenhorst, Krypt.-Fl., ed. 2, 1(8): 699 (1907)!, as 

‘Hormodendron’. 

≡ Hormoconis resinae (Lindau) Arx & G.A. de Vries, in Arx, Verh. Kon. Ned. Akad. 

Wetensch., Afd. Natuurk., Tweede Sect., 61(4): 62 (1973)!. 

= Racodium resinae Fr., Observ. mycol. 1: 216 (1815). 

≡ Sporocybe resinae (Fr.) Fr., Syst. mycol. 3(2): 341 (1832). 

≡ Sorocybe resinae (Fr.) Fr., Summa veg. Scand. 2: 468 (1849)!. 

≡ Dendryphion resinae (Fr.) Corda, Icon. fung. 6: 11 (1854). 

≡ Stysanopsis resinae (Fr.) Ferraris, Flora Ital. Crypt., Pars I, Fungi, Fasc. 6: 187 (1910). 

= Pycnostysanus resinae Lindau, Verh. Bot. Vereins Prov. Brandenburg 45: 160 (1904). 

≡ Stysanus resinae (Lindau) Sacc., Syll. fung. 18: 651 (1906). 

= Cladosporium avellaneum [G.A. de Vries] f. viride G.A. de Vries, Contr. Knowl. Genus 

Cladosporium: 56 (1952)!. 

Teleomorph: Amorphotheca resinae Parbery, Austral. J. Bot. 17: 340 (1969)!. 

Lit.: ELLIS (1971), DOMSCH et al. (1980), DAVID & KELLEY (1995), HO et al. (1999: 149), 

PARTRIDGE et al. (2001: 179), PARTRIDGE & MORGAN-JONES (2002: 344–348). 

resinae [(Lindau) G.A. de Vries] f. albidum (G.A. de Vries) G.A. de Vries, Antonie van 

Leeuwenhoek J. Microbiol. Serol. 21: 167 (1955)!. 

≡ Cladosporium avellaneum [G.A. de Vries] f. albidum G.A. de Vries, Contr. Knowl. Genus 



88

esinae [(Lindau) G.A. de Vries] f. avellaneum (G.A. de Vries) G.A. de Vries, Antonie van 


≡ Cladosporium avellaneum [G.A. de Vries] f. avellaneum G.A. de Vries, Contr. Knowl. 

Genus Cladosporium: 56 (1952)!. 


resinae [(Lindau) G.A. de Vries] f. sterile (G.A. de Vries) G.A. de Vries, Antonie van 


≡ Cladosporium avellaneum [G.A. de Vries] f. sterile G.A. de Vries, Contr. Knowl. Genus 



rhododendri K. Schub. & U. Braun, sp. nov. 

T: on Rhododendron sp. (Ericaceae), Germany, Sachsen-Anhalt, Halle (Saale), Pyrastraße, 11 

Jun. 2004, D. Seidel (HAL 1834 F: holotype). 

Host(s)/substrate(s) & distribution: on Rhododendron sp.; Europe (Germany). 

rhododendri, in herb. 

On leaves of Rhododendron sp. (Ericaceae), Switzerland, Bern, Berner Oberland, vom 

Faulhorn, 7/58 (B 70-6700). 

Notes: Excluded, generic affinity not yet clear. 

rhodomyrti Sawada, Rep. Gov. Res. Inst. Formosa 87: 74 (1944)!, nom. inval. 

T: on Rhodomyrtus tomentosa (Myrtaceae), Taiwan (PPMH). 

Notes: description only in Japanese, not validly published. Excluded, generic affinity remains 

unclear. 

rhois Arcang., in Thümen, Mycoth. univ., Cent. XIV, No. 1371 (1879)!. 

T: on living leaves of Rhus coriaria (Anacardiaceae), Italy, Etruria, Tuscany, Settignano, near 

Florence, Nov. 1878, Arcangeli, Thüm., Mycoth. univ. 1371 and Erb. Critt. Ital. 849 (e.g., BPI 

427440; E; HAL; K: syntypes). 

= Cercospora marmorata Tranzschel, in Tranzschel & Serebrianikow, Mycotheca Rossica, 

Fasc. 5, No. 250 (1911)!. [T: e.g., K; LE; W]. 

≡ Cercosporina marmorata (Tranzschel) Sacc., Syll. fung. 25: 895 (1931)!. 

≡ Phaeoramularia marmorata (Tranzschel) Deighton, Mycol. Pap. 144: 34 (1979)!. 

≡ Passalora marmorata (Tranzschel) U. Braun & Crous, in Crous & Braun, Mycosphaerella 


Ser. 1: 267 (2003)!. 

= Cercospora rhois-coriariae Kuhnh.-Lord., Ann. Éphiphyt., Ser. 2, 13: 54 (1947). 


rietmanni Sart. & Syd., Rev. Pat. Malad. Pays Chauds 15(1): 9–44 (1935). 

T: isolated from man (mycosis of the epidermis). 

= Hortaea werneckii (Horta) Nishim. & Miyaji, Jap. J. Med. Mycol. 26(2): 145 (1984). 

Lit.: CIFERRI (1960: 501), DE HOOG et al. (2000: 721). 

Notes: see C. werneckii. 

rigidiphorum R.F. Castañeda, in herb. 

On dead leaves of Smilax sp. (Smilacaceae) (CBS 314.95; MUCL 39142). 

rivinae Speg., Anales Mus. Nac. Buenos Aires 20: 437 (1910)!. 

T: on living leaves of Rivina laevis (= R. humilis) (Phytolaccaceae), Argentina, near Metán, 

Salta, Jun. 1905, C. Spegazzini, Mycetes Argent. Nr. 1108 (LPS 13.137: holotype). 

Lit.: SACCARDO (1913: 1369), FARR (1973: 251). 

Host(s)/substrate(s) & distribution: on Rivina humilis; South America (Argentina). 

robiniae (Kabát & Bubák) J.C. David, Mycol. Pap. 172: 92 (1997)!. 

T: on fallen leaves of Robinia pseudacacia (Fabaceae), Czech Republic, Turnau, 10/20 Nov. 

1903, J.E. Kabát (BPI: holotype; Kab. & Bub., F. imp. exs., No. 596: isotype, e.g., BPI; K). 

89

≡ Heterosporium robinae Kabát & Bubák, in Bubák & Kabát, Hedwigia 43: 421 (1904)!. 

Ill.: DAVID (1997: 84, Fig. 19 H–K; 93, Fig. 24). 

Host(s)/substrate(s) & distribution: on Robinia pseudacacia; Europe (Czech Republic), North 

America (USA: NY). 

roesleri Catt., Bol. Commiss. Agrar. Voghera 13: 263 (1876). 

T: on Vitis vinifera (Vitaceae), France, Dep. de l’Eure, Eburense, A. Malbranche. 

≡ Cercospora roesleri (Catt.) Sacc., Michelia 2(6): 128 (1880)!. 

≡ Cercospora roesleri ‘f. typica (Catt.)’ Elenkin, Bolezni Rast. 4: 67 (1909), nom. inval. 

≡ Ragnhildiana roesleri (Catt.) Vassiljevsky, in Vassiljevsky & Karakulin, Parazitnye nesovershennye 

griby, Ch. I, Gifomicety: 375 (1937)!. 

= Torula dissiliens Duby, Mem. Soc. Phys. Genève 7: 128 (1835). 

≡ Septocylindrium dissiliens (Duby) Sacc., Mycoth. ven., No. 583 (1876). 

≡ Phaeoramularia dissiliens (Duby) Deighton, in Ellis, More Dematiaceous Hyphomycetes: 

324 (1976)!. 

≡ Passalora dissiliens (Duby) U. Braun & Crous, in Crous & Braun, Mycospaerella and its 

anamorphs: 1. names published in Cercospora and Passalora, CBS Biodiversity Ser. 1: 164 

(2003)!. 

= ? Septocylindrium virens Sacc., Nuovo Giorn. Bot. Ital. 8: 186 (1876). 

= Septosporium fuckelii Thüm., Oesterr. Bot. Z. 27: 137 (1877). 

≡ Cercospora fuckelii (Thüm.) Jacz., Parasitic fungal diseases of grape vine, ed. 2: 81 (1906). 

≡ Cercospora roesleri f. fuckelii (Thüm.) Elenkin, Bolezni Rast. 4: 68 (1909). 

≡ Isariopsis fuckelii (Thüm.) du Plessis, Farming South Africa 17: 62 (1942). 

= Cladosporium pestis Thüm., Oesterr. Bot. Z. 27: 12 (1877)!. 

= ? Cercospora coryneoides Săvul. & Rayss, Rev. Pathol. Vég. Entomol. Agric. France 22: 

223 (1935). 

= Cercospora leoni Săvul. & Rayss, Rev. Pathol. Vég. Entomol. Agric. France 22: 222 (1935). 

[T: HUJ]. 

= Cercospora judaica Rayss, Palestine J. Bot, Jerusalem Ser. III, 50: 22 (1943). [T: HUJ]. 

Lit.: SACCARDO (1886: 458, as ‘rosleri’), LINDAU (1910: 117, as ‘Rösleri’), CHUPP (1954: 

604). 

‘roumegueri Speg.’, Rev. Mycol. (Toulouse) 1: 149 (1879)!. 

Notes: An error on the web-site of the herbarium HBG (as ‘rouegnesis’) and on the label of 

the collection in HBG, Cladotrichum roumegueri Speg., in Roumeguère, Rev. Mycol. 

(Toulouse) 1: 148–149 (1879) is intended. 

rubi Ellis & Everh. (Exs.: Flora Sequoia Gigantea Region, No. 1487, nom. nud.) 

T: on Rubus parviflorus (Rosaceae), USA, California, Jul. 1893, G. Hansen (B; BPI). 

= Cladosporium cladosporioides (Fresen.) G.A. de Vries, Contr. Knowl. Genus Cladosporium: 

57 (1952)!. 

rutae (T.M. Achundov) U. Braun, A Monograph of Cercosporella, Ramularia and Allied 

Genera (Phytopathogenic Hyphomycetes), vol. 2: 306 (1998)!. 

T: on Ruta graveolens (Rutaceae), Azerbaijan, Apscheron, botanical garden, 25 Mar. 1961, 

Achundov (BAK: holotype; LE 42008: isotype). 

≡ Ramularia rutae T.M. Achundov, Novosti Sist. Nizsh. Rast. 24: 96 (1987)!. 

Ill.: BRAUN (1998: 307, Fig. 571). 

Host(s)/substrate(s) & distribution: on Ruta graveolens; Asia (Azerbaijan). 

salicis Moesz & Smarods, in Moesz, Magyar Bot. Lapok 31: 42 (1932)!. 

T: on branches of Salix cinerea (Salicaceae), Latvia, near Adaži, 10 Jun. 1930, J. Smarods 

(M). Topotypes: Petr., Mycoth. gen. 1808 (e.g., BPI 427452); Behr, Plantae rarae et novae 

134-1939 (e.g., HAL), material collected at the type locality in 1937. 

Lit.: SACCARDO (1972: 1339). 

Ill.: MOESZ (1932: 43, Fig. 6). 

90

salicis-sitchensis Dearn. & Barthol., in Dearness, Mycologia 16: 174 (1924)!. 

T: on living leaves of Salix sitchensis (Salicaceae), USA, Washington, Langley, Sept. 1922, 

Grant, No. 5011 (DAOM: lectotype). 

= Ramulaspera salicina var. tirolense Bubák & Kabát, Oesterr. Bot. Z. 55: 243 (1905). 

≡ Ramularia salicina var. tirolense (Bubák & Kabát) Deighton, Trans. Brit. Mycol. Soc. 

90(2): 330 (1988). 

≡ Phacellium salicinum var. tirolense (Bubák & Kabát) U. Braun, Nova Hedwigia 56: 438 

(1993)!. 

Lit.: SACCARDO (1972: 1339), BRAUN (1998: 337). 

sambuci Brunaud, Champ. Charente-Infer. 1892: 38 (1892). 

T: on leaves of Sambucus nigra (Caprifoliaceae), France, Saintes. 

Lit.: SACCARDO (1895: 620). 

Notes: ZHANG et al. (2003) reported a collection on Sambucus williamsii from China 

determinated as Cladosporium sambuci, but the conidia of the Chinese collection were 

described to be much narrower, 2.6–5.1 µm. 

sambuci Pass., in herb. 

On living leaves of Sambucus nigra (Caprifoliaceae), Italy (B 70-6710: paratype material of 

Pseudocercospora sambucigena U. Braun & Crous). 

= Pseudocercospora sambucigena U. Braun & Crous, Mycotaxon (2005), in press. 

Notes: The herbarium name Cladosporium sambuci proved to be a true and new species of the 

genus Pseudocercospora. The well-growing collection was sparsely intermixed with a Cladosporium 

sp. 

‘sarcopodioides Sacc.’ – OUDEMANS (1924). 

Notes: An error. Clasterosporium sarcopodioides was intended. 

sarraceniae Dearn. & House, Circ. New York State Mus. 24: 58 (1940), nom. inval. 

T: on dead petioles and blades of Sarracenia purpurea (Sarraceniaceae), USA, New York, 

Albany Co., Voorheesville, 30 Aug. 1924, H.D. House (NYS 2716: holotype; DAOM: 

isotype). 

≡ Cladosporium punctatum Dearn. & House, in herb., non C. punctatum (Sacc.) Sacc., 1882. 

Notes: published without Latin diagnosis. 

savastani Carbone, Atti Ist. Bot. Univ. Pavia, Ser. 2, 14: 322 (1914)!. 

T: isolated from sausage [‘in botulis (Salame crudo)’], Italy, Pavia. 

Lit.: SACCARDO (1931: 799). 

scabies Cooke, Gard. Chron., Ser. 3, 34: 100 (1903)!. 

T: on fruits of cucumber (Cucurbitaceae), Great Britain. 


(1889). 

Lit.: MCKEMY & MORGAN-JONES (1992), CROUS et al. (2000). 

scillae Deighton, in Laundon, New Zealand J. Bot. 8(1): 55 (1970)!. 

T: on living leaves of Scilla peruviana (Hyacinthaceae), New Zealand, Levin, 21 Dec. 1965, 

G.F. Laundon, LEV 477 (IMI 116997: holotype). 

≡ Fusicladium scillae (Deighton) U. Braun & K. Schub., IMI Descriptions of Fungi and 

Bacteria 152, No. 1518 (2002)!. 

Lit.: SCHUBERT et al. (2003: 94–96). 

sclerotiophilum Sawada, Rep. Gov. Res. Inst. Formosa 51: 112 (1931)!, nom. inval. 

T: on twigs of Citrus grandis var. butan (Rutaceae), Taiwan, 25 Nov. 1928, K. Sawada 

(PPMH). 

Notes: description only in Japanese. Excluded, but taxonomic status not yet clear. 

scopiforme Berk., Hooker’s J. Bot. Kew Gard. Misc. 6: 208 (1854), as ‘scopæforme’. 

T: on leaves of Myristica sp. (Myristicaceae), India, Khasia (Churra), Hooker (K 115206; 

UPS). 

91

≡ Helminthosporium scopiforme (Berk.) Subram., J. Indian Bot. Soc. 35: 450 (1956), as 

‘scopæforme’. 

≡ Pleurophragmium scopiforme (Berk.) S. Hughes, Canad. J. Bot. 36: 798 (1958)!, as ‘scopæforme’. 

≡ Spiropes scopiformis (Berk.) M.B. Ellis, Mycol. Pap. 114: 30 (1968)!. 

= Cladosporium congestum Berk. & Broome, J. Linn. Soc., Bot. 14: 99, 1873 (1875)!. [T: K]. 

= Helminthosporium iteodaphnes Thüm., Rev. Mycol. (Toulouse) 2: 38 (1880). [T: W 89009]. 

≡ Cercospora iteodaphnes (Thüm.) Sacc., Syll. fung. 4: 464 (1886)!. 

Lit.: SACCARDO (1886: 358). 

scribnerianum Cavara, in Briosi & Cavara, F. paras., Fasc. 7/8, No. 187 (1892)! and Hedwigia 

31: 143 (1892)!. 

T: on leaves of Betula populifolia (Betulaceae), Italy, Pavia, 1890, F.L. Scribner, Briosi & 

Cav., F. paras. 187 (HAL: syntype). 

≡ Fusicladium scribnerianum (Cavara) M.B. Ellis, More Dematiaceous Hyphomycetes: 238 

(1976)!. 

Lit.: SACCARDO (1895: 620), LINDAU (1907: 819), FERRARIS (1912: 340), SCHUBERT et al. 

(2003: 96–97). 

secedens Fr., Summa veg. Scand. 2: 499 (1849)!. 

T: on rotten Sterea (= Stereum), Scandinavia. 

Lit.: SACCARDO (1886: 368). 

sericeum, in herb. 

On a decorticated limb of Magnolia fraseri (Magnoliaceae), USA, West Virginia, 18 Sept. 

1895, L.W. Nutall (BPI 427456, BPI 427457). 

sidae Cif. & Gonz. Frag., Bol. Real Soc. Esp. Hist. Nat. 25: 455 (1925)! and Publ. Estac. 

Agron. Haina, Ser. B, Bot., 2: 12 (1926). 

T: on dry stems of Sida sp. (Malvaceae), Dominican Republic, Haina, 28 Jun. 1925, Dr. R. 

Ciferri (MA 06453; BPI 427458: isotype). 

Lit.: SACCARDO (1972: 1339). 

simplex Schwein., Trans. Amer. Philos. Soc., N.S., 4(2): 277 (1832)!. 

T: on fallen leaves of Fraxinus sp. (Oleaceae), USA, Pennsylvania, Bethlehem, No. 2606 (PH 

1020415). 

Lit.: SACCARDO (1886: 360). 

Notes: The taxonomic status of this species is quite unclear since the type material is too 

meagre for a final conclusion. It was not possible to find sufficient fructification for a reevaluation 

of this species. The original description (SCHWEINITZ 1832) is too brief and noninformative 

for any conclusions about the status of this fungus. 

smilacicola K. Schub., sp. nov. 

T: on living leaves of Smilax grandiflora (Smilacaceae), Germany, Munic, botanical garden, 

cold house, Mar. 1895, Allescher (M-57718: holotype). 

Host(s)/substrate(s) & distribution: on Smilax grandiflora; Europe (Germany). 

smilacis (Schwein.) Fr., Syst. mycol. 3(2): 369 (1832)!. 

T: in the epidermis of Smilax rotundifolia (Smilacaceae), USA, Salem, Bethlehem (PH 

01020424, 01020425: syntypes). 

≡ Dematium smilacis Schwein., Syn. fung. Carol. sup.: 102 (1822)!. 

Lit.: SACCARDO (1886: 367). 

Notes: No Cladosporium found, type material in bad condition, excluded. ZHANG et al. (2003) 

cited a collection on Smilax china from Shaanxi, China. 

solanicola Viégas, Bragantia 6: 368 (1946)!, as ‘solanicolum’. 

T: on Solanum lycocarpum (Solanaceae), Brazil, Prov. St. Pauli, Campinas, Bosque de 

Jequìtibás, 27 Jun. 1913, A.P. Viégas (IACM: holotype). 

≡ Mycovellosiella solanicola (Viégas) Munt.-Cvetk., Lilloa 30: 178 (1960)!. 

92

= Cercospora brachycarpa Syd., Ann. Mycol. 28: 207 (1930)!. [T: IMI 8500a]. 

≡ Mycovellosiella brachycarpa (Syd.) Deighton, Mycol. Pap. 137: 8 (1974)!. 

≡ Passalora brachycarpa (Syd.) U. Braun & Crous, in Crous & Braun, Mycosphaerella and 

its anamorphs: 1. Names published in Cercospora and Passalora, CBS Biodiversity Ser. 1: 

87 (2003)!. 

= Cercospora jaguarensis Chupp & A.S. Mull., Bol. Soc. Venez. Ci. Nat. 8: 48 (1942), nom. 

inval. [T: CUP; IMI 105210a]. 

Notes: ZHANG et al. (1998) mentioned a first record of this species from China on Solanum 

melongena. 

soldanellae Jaap, Ann. Mycol. 5: 270 (1907)!. 

T: on dead leaves of Soldanella alpina (Primulaceae), Switzerland, Simplonhospiz, 2010 m 

alt. 

= Cladosporium stysanoides Bubák, Bot. Közlem. 15(3–4): 81 (1915)!, syn. nov. 

Lit.: LINDAU (1910: 796), FERRARIS (1912: 348), SACCARDO (1913: 1368). 

Host(s)/substrate(s) & distribution: on Soldanella alpina; Europe (Italy, Montenegro, Switzerland). 

Notes: Type at B missing (Burghard Hein, personal communication). 

solutum Link, in Willd., Sp. pl., 6(1): 39 (1824)!. 

T: on stems of Hibiscus esculentus (≡ Abelmoschus esculentus) (Malvaceae), Egypt, near Sjut, 

Oct., Ehrenberg. 

≡ Cladosporium herbarum b solutum (Link) Rabenh., Krypt.-Fl. 1: 113 (1844)! : Fr. (1832). 

≡ Cladosporium herbarum var. solutum (Link) Sacc., Syll. fung. 4: 351 (1886)!. 

Lit.: PRASIL & DE HOOG (1988: 53, as ‘C. herbarum var. solutum’). 

Notes: Authentic material is not preserved in herb. B, so the identity of the species cannot be 

established (PRASIL & DE HOOG 1988). In OUDEMANS (1923), Ricinus communis (Euphorbiaceae) 

is given as host. 

sorghi S.R. Chowdhury, Sydowia 23(6): 50 ‘1969’ (1970)!. 

T: on living inflorescences of Sorghum vulgare (Poaceae), India, Madhya Pradesh, Raipur, 

Coll. of Science, Dec. 1966, S.R. Chowdhury (IMI 125190b: holotype). 

= Cladosporium oxysporum Berk. & M.A. Curtis, in Berkeley, J. Linn. Soc. Bot. 10: 362 

(1869)!. 

Ill.: CHOWDHURY (1970: 51, Fig. 2). 

sparsum Schwein., Trans. Amer. Philos. Soc., N.S., 4(2): 277 (1832)!. 

T: on leaves of Allium cepae and Allium sp. (Alliaceae), USA, Pennsylvania, Bethlehem, No. 

2602 (PH 01020413, 01020414: syntypes). 



Lit.: SACCARDO (1886: 367). 

sphaeroideum Cooke, Grevillea 8(46): 60 (1879)!. 

T: on leaves of Poa foliosa (Poaceae), New Zealand, Canterbury Alps, No. 398 (K 121569: 

holotype). 

= Passalora graminis (Fuckel) Höhn., Zentralbl. Bakteriol. Parasitenk., Abt. 2, 60: 6 (1923). 

Lit.: SACCARDO (1886: 365), LIND (1913), SCHUBERT & BRAUN (2005b). 

Notes: COOKE (1879) stated that the habit of this species resembles a Sphaeria or Venturia. 

sphaerospermum Penz., Michelia 2(8): 473 (1882)!. 

T: on faded leaves and branches of Citrus sp. (Rutaceae), Italy, Padova, Feb. 1882, O. Penzig. 

= Torula lichenopsis Höhn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 83: 36 

(1927)!. [T: FH-Höhnel 1275: holotype]. 

= Hormodendron langeronii Fonseca, Leão & Nogueira, Sci. Med. 5: 563 (1927). 

≡ Cladosporium langeronii (Fonseca, Leão & Nogueira) Vuill., Champ. paras.: 78 (1931)!. 

≡ Cladosporium langeronii (Fonseca, Leão & Nogueira) Cif., Manuale di Micologia Medica, 

ed. 2: 488 (1960)!, as ‘langeroni’, comb. superfl. 

93

Lit.: SACCARDO (1886: 355), LINDAU (1907: 826), FERRARIS (1912: 345), DE VRIES (1952: 

81), YAMAMOTO (1959: 3), ELLIS (1971: 315), HAWKSWORTH (1979: 287), DOMSCH et al. 

(1980: 209), ELLIS & ELLIS (1985: 290, 1988), WANG & ZABEL (1990: 200), HO et al. (1999: 

139), DE HOOG et al. (2000: 591), SAMSON et al. (2000: 114), SAMSON et al. (2001: 340). 

Ill.: DE VRIES (1952: 82, Fig. 18), YAMAMOTO (1959: 2, Figs 5–8), MINOURA (1966: 141, Fig. 

5C), ELLIS (1971: 316, Fig. 218 A), DOMSCH et al. (1980: 209, Fig. 85), HO et al. (1999: 141, 

Figs 42–43), DE HOOG et al. (2000: 591–592, Figs), SAMSON et al. (2000: 114, Fig. 51; 115, 

Pl. 49), SCHELL (2003: 583, Fig. 18). 

Host(s)/substrate(s) & distribution: as a secondary invader on many different plants as well as 

in air, soil, foodstuffs, paint, textiles and occasionally isolated from man and animals; 

cosmopolitan. 

sphaerosporum (sic) – BARRON (1968: 130), Fig. 55. 

Notes: Neither C. sphaerospermum nor C. ‘sphaerosporum’ are amongst the Cladosporium 

species indexed by BARRON (1968), but Figure 55, of which a detail provides the illustration 

for the cover and an enlargement the frontispiece for BARRON (1968), is a commendable 

illustration of typical C. sphaerospermum. 

sphondilii Fuss, Archiv Verein siebenb. Landesk., N.F., 14(2): 431 (1878)!, nom. nud. 

On dry stems of Heracleum sphondylium (Apiaceae), Romania, Grosscheuern, Giresau, Fuss. 


spinae-christi M.B. Ellis, in herb., not published, cited on CABI page. 

spongiosum Berk. & M.A. Curtis, in Berkeley, J. Linn. Soc., Bot. 10(46): 362 (1869)!. 

T: on inflorescences of Cenchrus (Poaceae), Cuba, C. Wright, No. 287 (K 121570: holotype). 

≡ Helminthosporium spongiosum (Berk. & M.A. Curtis) Cif., Atti Ist. Bot. Lab. Crittog. Univ. 

Pavia, Ser. 5, 19: 109 (1962)!. 


Ill.: ELLIS (1971: 316, Fig. 218 B). 

Host(s)/substrate(s) & distribution: on Cenchrus sp.; Caribbean (Cuba). 

Notes: BERKELEY (1869) compared the habit of his newly described species with 

Helminthosporium ravenelii M.A. Curtis. CIFERRI (1962) did not examine type material of 

Cladosporium spongiosum Berk. & M.A. Curtis, but rather based his new combination on 

material collected by himself on Cenchrus echinatus in the Dominican Republic (in comparing 

it with the description given by Berkeley and Curtis). 

stanhopeae Allesch., Hedwigia 34: 221 (1895)!. 

T: on faded leaves of Stanhopea sp. (Orchidaceae), Germany, Munich, botanical garden, Sept. 

1894, Allescher (M: holotype). 

Lit.: SACCARDO (1895: xlviii; 1899: 1081), LINDAU (1907: 817), SCHUBERT & BRAUN (2004: 

311). 


Host(s)/substrate(s) & distribution: on Stanhopea sp.; Europe (Germany). 

staurophorum (W.B. Kendr.) M.B. Ellis, More Dematiaceous Hyphomycetes: 333 (1976)!. 

T: on fallen needles of Pinus sylvestris (Pinaceae), Great Britain, Cheshire, Delamere Forest, 

1957, isol. B. Kendrick (IMI 71590: lectotype). 

≡ Hormodendrum staurophorum W.B. Kendr., Canad. J. Bot. 39: 835 (1961)!. 

≡ Devriesia staurophora (W.B. Kendr.) Seifert & N.L. Nickerson, in Seifert et al., Canad. J. 

Bot. 82: 919 (2004). 

Lit.: ELLIS & ELLIS (1985: 175), HO et al. (1999: 140). 

Ill.: KENDRICK (1961: 833–834, Figs 1–2; Pl. 1, Figs 3–5), ELLIS (1976: 334, Fig. 252 B), HO 

et al. (1999: 141, Figs 44–45), SEIFERT et al. (2004: 920, Figs 2–12). 

stenosporum Berk. & M.A. Curtis, in Berkeley, Grevillea 3(27): 107 (1875)!. 

T: on Stylosanthes (Fabaceae), USA, South Carolina, No. 2067 and on leaves of Malus sp. 

(Rosaceae), USA, No. 2529 (not in herb. K). 

Lit.: SACCARDO (1886: 352). 

94

Notes: BUBÁK (1916) examined material of Acrotheca dearnessiana Sacc. sent to him by J. 

Dearness and thought it to be Cladosporium stenosporum. Acrotheca dearnessiana is a 

synonym of Fusicladium pyrorum (Lib.) Fuckel (SCHUBERT 2001, CROUS & BRAUN 2003: 

488). ZHANG et al. (2003) reported and illustrated a collection determinated as C. stenosporum 

from Zhejiang, China on Pyrus calleryana. 

stercorarium Corda, Icon. fung. 1: 14 (1837)!. 

T: on bird dung, Czech Republic. 



Notes: Type material is not preserved in herb. PRM. 

stercoris Speg., Anales Mus. Nac. Buenos Aires, Ser. 2, 6: 338 (1899)!. 

T: on old rabbit dung, Argentina, Parque de La Plata, May 1888. 

Lit.: SACCARDO (1902: 1059), FARR (1973: 251). 

stevensii (G. Arnaud) Toro (STEVENSON 1975) 

An error, Clasterosporium was intended. 

stipae H.C. Greene, Trans. Wisconsin Acad. Sci. 41: 127 (1952)!. 

T: on living leaves of Stipa spartea (Poaceae), USA, Wisconsin, Dane Co., Madison, Univ. 

Wisconsin Arboretum, Oak opening, 6 Sept. 1951, H.C. Greene (BPI 427474, WIS: syntypes). 

= Stenella stipae (H.C. Greene) K. Schub. & U. Braun, in Schubert, Mycotaxon 92: 71 (2005). 


straminicola Pidopl. & Deniak, in Pidoplichko, Gribnaja Flora Grubych Kormov: 269 

(1953)!, nom. inval. 

T: on straw and on hay, Ukraine. 


‘strictum Sacc.’ − GOLA (1930), Cladotrichum strictum Sacc. is intended. 

strobilanthis H.J. Lu, Y.L. Liu & Z.Y. Zhang, Mycosystema 22: 49 (2003)!. 

T: on living leaves of Strobilanthes cusia (Acanthaceae), China, Shaanxi Prov., Xian, 29 Aug. 

1989, T.F. Li & H. Li (MHYAU 07908: holotype). 

Notes: The publication of this name is connected with a severe misprint. The text starts with 

the Latin description, but the name of the new species and ‘sp. nov.’ is missing. However, in 

the Chinese summary the name ‘C. strobilanthis’as new species appears, so that it can 

nevertheless be considered a valid name. 

‘stromatum Pers.’ – listed by SACCARDO & BERLESE (1884: 100). 

stromatum Preuss, in Sturm, Deutschl. Fl. 3(26): 25 (1848)!. 

T: on wood of Pinus sp. (Pinaceae), Germany, Hoyerwerda. 

Lit.: SACCARDO (1886: 352, 355), LINDAU (1907: 811), FERRARIS (1912: 339), OUDEMANS 

(1919: on Eutypa lejoplaca, 1920: on Juglans regia, 1921: on Acer campestris). 

Ill.: PREUSS (1848: Tab. 13). 

strumelloideum Milko & Dunaev, Novosti Sist. Nizsh. Rast. 23: 134 (1986)!. 

T: on leaves of Carex sp. (Cyperaceae) from stagnent water, Russia, Yaroslavskaya Oblast, 

Rybinskoe, Sutka (BKMF-2534). 

stysanoides Bubák, Bot. Közlem. 15(3–4): 81 (1915)!. 

T: on leaves of Soldanella alpina (Primulaceae), Montenegro, Durmitor, Lokvice, ca. 2200 m, 

15 Aug. 1904, F. Bubák (BPI 427476: holotype). 

= Cladosporium soldanellae Jaap, Ann. Mycol. 5: 270 (1907)!. 

Lit.: SACCARDO (1931: 794). 

suaveolens (Lindner) Delitsch, Ergebnisse der theoretischen und angewandten Mikrobiologie, 

Ed. Lembke, Bd. 1, Systematik der Schimmelpilze: 135 (1943). 

T: in a distillery. 

95

≡ Sachsia suaveolens Lindner, Mikroskopische Betriebskontrolle in den Gährungsgewerben: 

153 (1895). 

≡ Oospora suaveolens (Lindner) Lindau, in Rabenhorst, Krypt.-Fl., ed. 2, 1(8): 35 (1907)!. 

≡ Candida suaveolens (Lindner) Langeron & Guerra, ? 

≡ Geotrichum suaveolens (Lindner) Cif., in Caretta, Atti Ist. Bot. Lab. Crittog. Univ. Pavia, 

Ser. 5, 19: 6 (1962)! (cited as ‘Cif., in Diddens & Lodder, 1942’ ?). 

≡ Moniliella suaveolens (Lindner) Arx, Antonie van Leeuwenhoek J. Microbiol. Serol. 38(3): 

294 (1972)!. 

Notes: “The species does not belong in the genus Cladosporium” (DE VRIES 1952: 97, with 

comments on type and other specimens). 

subcompactum Roum. & P. Karst. → compactiusculum. 

subcompactum Sacc., Syll. fung. 4: 361 (1886)!. 

T: on dead stems of Delphinium ajacis (Ranunculaceae), France, Rouen, Letendre. 

≡ Cladosporium compactum Sacc., Michelia 2(6): 127 (1880)!, nom. illeg., non C. compactum 

Berk. & M.A. Curtis, 1875. 

Lit.: LINDAU (1907: 825), FERRARIS (1912: 344; 1914: 884), OUDEMANS (1921), GONZÁLES- 

FRAGOSO (1927: 203). 

subfusoideum McAlpine, Fungus Dis. Citrus Trees Austral.: 79 (1899)!. 

T: on fruits of Citrus medica (Rutaceae), Australia, Victoria, Wandin Yallok, Sept. 1898 and 

New South Wales, Parramatta, Feb. 1899, Cairnes. 

Lit.: SACCARDO (1902: 1058). 

Ill.: MCALPINE (1899: Pl. 5, Figs 21–22). 

Notes: Types are lost, no specimen in VPRI. ‘Type details agree with Diplodia citricola 

McAlpine (in part)’ (personal communication with J. Cunnington / I. Pascoe). 

subnodosum Cooke, Grevillea 17(83): 67 (1889)!. 

T: on leaves of Spinacia oleracea (Chenopodiaceae), USA, South Carolina, Aiken, Rav., F. 

amer. exs. 294 (NY: lectotype). Isolectotypes: Rav., F. amer. exs. 294, e.g., BPI 427478; NY; 

PH. 

= Cladosporium variabile (Cooke) G.A. de Vries, Contr. Knowl. Genus Cladosporium: 85 

(1952). 

Lit.: SACCARDO (1892: 601, 1895: 621). 

subsclerotioideum Bubák & Dearn., in Bubák, Hedwigia 58: 33 (1916)!. 

T: on living leaves of Turritis glabra (= Arabis glabra) (Brassicaceae), Canada, Ontario, 

London, Jun.–Jul. 1910, J. Dearness, mixed infection with Peronospora parasitica and Albugo 

candida (BPI 427479). 

Lit.: SACCARDO (1931: 790). 

Host(s)/substrate(s) & distribution: on Arabis glabra; North America (Canada). 

subsessile Ellis & Barthol., Erythea 4: 83 (1896). 

T: on living leaves of Populus monilifera (= Populus deltoides subsp. monilifera) (Salicaceae), 

USA, Kansas, 18 Sept. 1894, Bartholomew (NY: lectotype, selected by SCHUBERT et al. 

2003); isolectotypes: on leaves of Populus deltoides subsp. monilifera, USA, Kansas, Rockport, 

Sept. 1894, E. Bartholomew, Ellis & Everh., N. Am. F. 3288 (e.g., M; NY). 

≡ Cladosporium brevipes Ellis & Barthol., Erythea 4: 27 (1896), nom. illeg., non C. brevipes 

Peck, 1887. 

≡ Fusicladium subsessile (Ellis & Barthol.) K. Schub. & U. Braun, IMI Descriptions of Fungi 


≡ Cladosporium maculicola Ellis & Barthol., in herb. (BPI 427256). 

Lit.: SACCARDO (1899: 1081), FERRARIS (1912: 345), SCHUBERT et al. (2003: 99–100). 

subtile Rabenh., F. eur., Ed. Nov., Ser. II, Cent. XXIV, No. 2364 (1876)!, nom. nud. 

T: on legumes of Leucaena glauca (Mimosaceae), India, Calcutta, Rabenh., F. eur. 2364 (e.g., 

HAL). 

96

= Cladosporium oxysporum Berk. & M.A. Curtis, in Berkeley, J. Linn. Soc., Bot. 10: 362 

(1869)!. 

Lit.: SACCARDO (1895: 621), BAGYANARAYANA & BRAUN (1999: 13). 

superficiale Petch, Ann. Roy. Bot. Gard. (Peradeniya) 9: 327 (1925). 

T: on leaves of Cinnamomum ovalifolium (Lauraceae), India, Ceylon, Hakgala, 27 Feb. 1922, 

No. 6570 (K 121571: holotype). 

Lit.: SACCARDO (1972: 1339). 

Notes: Excluded, mycelial sooty mould. 

sycophilum Farneti, Atti Ist. Bot. Univ. Pavia, Ser. 2, 8: 517 (1904)!, as ‘sicophilum’. 

T: on living fruits of Ficus carica (Moraceae), Italy, Pavia. 


Notes: In OUDEMANS (1920) Morus alba is given as a further host. 

symphoricarpi Dearn., in herb. 

On Symphoricarpos acutus (Caprifoliaceae), Canada, British Columbia, Salmo, 11 Jul. 1935, 

G.G. Hedgcock (BPI 427503). 

= Cercospora symphoricarpi Ellis & Everh., J. Mycol. 5: 70 (1889). 

≡ Phaeoramularia symphoricarpi (Ellis& Everh.) Deighton, on Ellis, More dematiaceous 

hyphomycetes: 317 (1976). 

≡ Passalora symphoricarpi (Ellis & Everh.) U. Braun & Crous, in Crous & Braun, Mycosphaerella 

and its anamorphs. 1. Names published in Cercospora and Passalora. CBS 

Biodiversity Ser. 1: 393 (2003). 

Notes: As secondary invader Cladosporium herbarum s. lat and a cladosporioides-like Cladosporium 

species are occasionally intermixed. 

syphiliticum Hallier, Flora, Neu Reihe, 26(19): 294 (1868)!. 

T: isolated from man associated with syphilis. 

≡ Cladosporium coniothecii-syphilitici Hallier, Flora, Neu Reihe, 26(19): 294 (1868)! (alternative 

name). 

Lit.: SACCARDO (1913: 1371), NANNIZZI (1934: 409). 

Ill.: HALLIER (1868a: Tab. 3, Fig. 13). 

Notes: Introduced as state (morphe) of Coniothecium syphiliticum Hallier and Penicillium 

syphiliticum Hallier (p. 295). A doubtful, human pathogenic fungus associated with syphilis, 

undoubtedly not belonging to Cladosporium s. str. “Est species omnini obsura et vix 

Cladosporium.” (SACCARDO 1913). “Species incertae” (SACCARDO 1911: 282). Of E. Hallier, 

“Herbarium and types: unknown” (STAFLEU & COWAN 1979). 

syringae Montem., Riv. Patol. Veg., Ser. 2, 1915: 226 (1915). 

T: on leaves of Syringa vulgaris (Oleaceae), Italy, Montubeccaria, Pavia. 

Lit.: SACCARDO (1931: 793). 

syringicola K. Schub. & U. Braun, sp. nov. 

T: on Syringa ×chinensis (Oleaceae), Germany, Sachsen-Anhalt, Halle (Saale), botanical 

garden, 2 Aug. 2004, K. Schubert, mixed infection with Erysiphe syringae-japonicae (U. 

Braun) U. Braun & S. Takam. (HAL 1835 F: holotype). Isotypes: U. Braun, Fungi selecti 

exsiccati 36. 

Host(s)/substrate(s) & distribution: on Syringa ×chinensis; Europe (Germany). 

tabaci Oudem., Beih. Bot. Centralbl. 11: 538 (1902)!. 

T: on decaying leaves of Nicotiana tabaca (Solanaceae), Netherlands, Bussum, Aug. 1901, 

C.J. Koning (L). 

= Cladosporium nicotianae Oudem., Ned. Kruidk. Arch., Ser. 3, 2(3): 769 (1902)!. 


taphrinae Bubák, Bot. Közlem. 15(3–4): 81 (1915)!. 

T: on Taphrina coerulescens on living leaves of Quercus cerris (Fagaceae), Montenegro, 

Šavnik, 30 Sept. 1911, L. Vlach (BPI 427506: holotype). 

97

Lit.: SACCARDO (1931: 797), HEUCHERT et al. (2005: 40). 

Ill.: HEUCHERT et al. (2005: 41, Fig. 15; Pl. 2, Figs 10–11). 

Host(s)/substrate(s) & distribution: on asci of Taphrina coerulescens; Europe (Montenegro), 


tectonae Sawada, Rep. Gov. Res. Inst. Formosa 85: 92 (1943)!, nom. inval. 

T: on Tectona grandis (Verbenaceae), Taiwan, Taipeh, 6 May 1930, K. Sawada (BPI 427507; 

PPMH: syntypes). 

= Cladosporium tectonicola Y.H. He & Z.Y. Zhang, Mycosystema 21(1): 21 (2002) and in 

Zhang et al., Flora Fungorum Sinicorum, Vol. 14: 164 (2003). 

Notes: description only in Japanese, not validly published. Excluded, but generic affinity not 

yet clear. 

tectonicola Y.H. He & Z.Y. Zhang, Mycosystema 21(1): 21 (2002)! and in Zhang et al., Flora 

Fungorum Sinicorum, Vol. 14: 164 (2003)!. 

T: on living leaves of Tectona grandis (Verbenaceae), China, Guangdong, Ledong, 30 Aug. 

1978, D.R. Duan (HMAS 38603: holotype). 

Ill.: HE & ZHANG (2002: 21, Fig. 1), ZHANG et al. (2003: 165, Fig. 114). 

Notes: ZHANG et al. (2003) cited Cladosporium tectonae Sawada as synonym. Excluded, 

taxonomic status unknown. 

tenerrimum Link, in Willd., Sp. pl. 6(1): 41 (1824)!. 

T: on rotting bark, Germany, Berlin, Link (B). 


tenerum (Link) E.W. Mason, in herb.?, Kirk et al. (n. d.) CABI page. 

tenuis – GOLA (1930: 21). 

tenuissimum Cooke, Grevillea 6(40): 140 (1878)!. 

T: on sheats of Zea mays (Poaceae), USA, South Carolina, Aiken, H.W. Ravenel, Rav., F. 

amer. exs. 160 (NY: lectotype). Isolectotypes: Rav., F. amer. exs. 160 (e.g., K, PH 01020427). 

Lit.: SACCARDO (1886: 365), OUDEMANS (1919), ELLIS (1976: 326), HO et al. (1999: 140), 

HEUCHERT et al. (2005: 50–52). 

Ill.: ELLIS (1976: 327, Fig. 245 A), HO et al. (1999: 143, Figs 46–47), HEUCHERT et al. (2005: 

51, Fig. 20). 

Host(s)/substrate(s) & distribution: on different host plants, also isolated from air, bread and 

soil; cosmopolitan but especially common in the tropics. 

tetrapanacis D.X. Wu & Z.Y. Zhang, Mycosystema 22: 48 (2003)!. 

T: on living leaves of Tetrapanax papyriferus (Araliaceae), China, Shaanxi Prov., Xian, 29 

Aug. 1989, T.F. Li (MHYAU 07906: holotype). 

Ill.: WU & ZHANG (2003: 49, Fig. 1). 

teucrii Y.L. Liu & Z.Y. Zhang, Plant Diseases and Their Control: 101 (1998)!. 

T: on living leaves of Teucrium viscidum (Lamiaceae), China, Hubei, Wuchang, 24 Sept. 

1980, J.Y. Li & T.Y. Zhang, No. 41007 (MHYAU 03954: holotype). 

Ill.: LIU & ZHANG (1998: 102, Fig. 1), ZHANG et al. (2003: 168, Fig. 116). 

theobromicola Av.-Saccá, Bol. Agric. (São Paulo) 21: 59 (1920)!, as ‘theobromicolum’. 

T: on Theobroma cacao (Sterculiaceae), Brazil. 

Ill.: AVENA-SACCÁ (l.c.: Figs. 3–5). 

tomentosum Corda, Icon. fung. 1: 15 (1837)!. 

T: on wood and rotten bark, Czech Republic, near Reichenberg (PRM). 



Lit.: SACCARDO (1886: 356), LINDAU (1907: 809), HUGHES (1958: 751). 


98

tortuosum Fr., Summa veg. Scand. 2: 499 (1849)!. 

T: on wood of Quercus (Fagaceae), Scandinavia. 

Lit.: SACCARDO (1886: 354). 

transchelii Pidopl. & Deniak, Mikrobiol. Zhurn. 5(2): 188, 194 (1938)!, nom. inval. 

T: from stalks of Zea mays (Poaceae), Ukraine. 



transchelii var. semenicola (Pidopl. & Deniak) Pidopl. & Bilai, in Pidoplichko, Gribnaja 

Flora Grubych Kormov: 274 (1953)!, nom. inval. 

T: on grains of oats, Ukraine. 

≡ Cladosporium viridiolivaceum var. semenicola Pidopl. & Deniak ? 

Notes: It is unknown where the authors published this variety of C. viridiolivaceum Pidopl. & 

Deniak. 

transchelii var. viridi-olivacearum (Pidopl. & Deniak) Pidopl. & Bilai, in Pidoplichko, 

Gribnaja Flora Grubych Kormov: 274 (1953)!, nom. inval. 

T: isolated from rotting apples (Malus sp., Rosaceae), Ukraine. 

≡ Cladosporium viridiolivaceum Pidopl. & Deniak, Mikrobiol. Zhurn. 5(2): 187, 194 (1938)!, 

as ‘viridi-olivaceum’, nom. inval. 


trichellum Sacc. – GOLA (1930: 21). 

trichoides C.W. Emmons, in Binford, Thompson & Gorham, Amer. J. Clin. Pathol. 22: 541 

(1952)!. 

T: isolated from man, USA. 

= Cladophialophora bantiana (Sacc.) de Hoog, Kwon-Chung & McGinnis, in de Hoog, 


Mycol. 33: 343 (1995)!. 

Lit.: MCGINNIS & BORELLI (1981), HO et al. (1999: 146). 

trichoides [C.W. Emmons] var. chlamydosporum Kwon-Chung, Mycologia 75(2): 320 

(1983)!. 

T: from brain abscess in man, USA, Maryland. 

= Cladophialophora bantiana (Sacc.) de Hoog, Kwon-Chung & McGinnis, in de Hoog, 


Mycol. 33: 343 (1995)!. 

trichophilum H.C. Greene, Amer. Midl. Naturalist 48(3): 756 (1952)!, nom. illeg., homonym, 

non C. trichophilum Petr. & Cif., 1932. 

T: on living leaves of Lonicera hirsuta (Caprifoliaceae), USA, Wisconsin, Rusk Co., Hawkins, 

26 Aug. 1918, J.J. Davis (BPI 427512, WIS: syntypes). 

= Mycovellosiella nopomingensis B. Sutton, Mycol. Pap. 132: 77 (1973). 

≡ Passalora nopomingensis (B. Sutton) U. Braun & Crous, in Crous & Braun, Mycosphaerella 

and its anamorphs: 1. Names published in Cercospora and Passalora. CBS Biodiversity 

Ser. 1: 462 (2003). 

Lit.: SHVARTSMAN et al. (1975: 96), SCHUBERT (2005: 73–75). 


trichophilum Petr. & Cif., Ann. Mycol. 30: 337 (1932)!. 

T: on living leaves of Lantana trifolia (Verbenaceae), Dominican Republic, Valle del Cibao, 

Prov. Santiago, Las Lagunas, at Pozo Hediondo, 7 Dec. 1930, R. Ciferri & E.L. Ekman (BPI 

427513A, 43696A; IMI 127138a; M; W). 

≡ Mycovellosiella trichophila (Petr. & Cif.) Deighton, in herb. ? 

= Cercospora lantanae Chupp, in Toro, J. Dept. Agric. Porto Rico 15: 10 (1931). [T: CUP-PR 

1200; IMI 132050]. 

≡ Mycovellosiella lantanae (Chupp) Deighton, Mycol. Pap. 137: 33 (1974)!. 

99

≡ Passalora lantanae (Chupp) U. Braun & Crous, in Crous & Braun, Mycosphaerella and its 


242 (2003)!. 

= Chaetotrichum lantanae Petr., Sydowia 5: 38 (1951), nom. nov., non Chaetotrichum trichophilum 

(Stev.) Petr., 1951. 

= Mycovellosiella lantanae var. verbenacearum K. Bhalla, S.K. Singh & A.K. Srivast., 

Australas. Syst. Bot. 12: 369 (1999). [T: IMI 373101]. 

trillii Ellis & Everh., Proc. Acad. Nat. Sci. Philadelphia 47: 430 (1895)!. 

T: on leaves of Trillium petiolatum (Trilliaceae), USA, Washington, Pullman, Jun. 1894, C.V. 

Piper, no. 341, mixed infection with Phyllosticta trillii Ellis & Everh. (NY: holotype). 

= Heterosporium trillii Ellis & Everh., Proc. Acad. Nat. Sci. Philadelphia 46: 382 (1894)!. 

≡ Cladosporium trillii (Ellis & Everh.) J.C. David, Mycol. Pap. 172: 94 (1997)!, nom. illeg., 

homonym, non C. trillii Ellis & Everh., 1895. 

≡ Cladosporium trilliicola J.C. David, Schlechtendalia 11: 88 (2004). 

Lit.: SACCARDO (1899: 1081). 

Host(s)/substrate(s) & distribution: on species of Trillium, including T. ovatum and T. petiolatum; 


trilliicola J.C. David, Schlechtendalia 11: 88 (2004). 

T: on leaves (partly dead) of Trillium ovatum (Trilliaceae), USA, Idaho, Latah Co., 14 Jul. 

1893, C.V. Piper (NY: holotype). 

≡ Heterosporium trillii Ellis & Everh., Proc. Acad. Nat. Sci. Philadelphia 46: 382 (1894)!. 

≡ Cladosporium trillii (Ellis & Everh.) J.C. David, Mycol. Pap. 172: 94 (1997)!, nom. illeg., 

homonym, non C. trillii Ellis & Everh., 1895. 

= Cladosporium trillii Ellis & Everh., Proc. Acad. Nat. Sci. Philadelphia 47: 430 (1895)!. 

Ill.: DAVID (1997: 89, Fig. 22 C–F; 95, Fig. 25). 

triostei Peck, in Trelease, Trans. Wisconsin Acad. Sci. 6: 119 (1885)! and J. Mycol. 1: 13 

(1885)!. 

T: on leaves of Triosteum perfoliatum (Caprifoliaceae), USA, Wisconsin, La Crosse, L.H. 

Pammel (NYS 3219: holotype). 

≡ Fusicladium triostei (Peck) K. Schub. & U. Braun, Mycol. Progr. 4(2): 102 (2005)!. 

Lit.: SACCARDO (1886: 359), DE VRIES (1952: 99). 


tropicale Sartory, R. Sartory, J. Mey. & R. Weiss, Bull. Acad. Roy. Méd. 113(24): 890 

(1935), as ‘tropicalis’, nom. inval. 

T: ‘dermatomycosis tropicalis’, disease caused in man, Central Africa. 

Notes: Latin diagnosis lacking, description rudimentary. “Doubtful, probably an Exophiala” 

(DE HOOG et al. 2000: 1033) or an Aureobasidium (CIFERRI 1960: 501). 

tuberculatum Fr., Summa veg. Scand. 2: 499 (1849)!. 

T: on ostiola of Cytospora leucosperma, Scandinavia. 

Lit.: SACCARDO (1886: 368). 

tuberum Cooke, Grevillea 12(61): 31 (1883)!. 

T: on tubers of Batata edulis (= Ipomoea batatas) (Convolvulaceae), USA, South Carolina, 

Aiken, Rav., F. amer. exs. 600 (BPI 427529; K; NY; PH 01020426: syntypes). 

Lit.: SACCARDO (1886: 362). 

typhae Schwein., Trans. Amer. Philos. Soc., N.S., 4(2): 277 (1832)!. 

T: on dead leaves of Typha sp. (Typhaceae), USA, Pennsylvania, Bethlehem, No. 2603 (PH 

01020410). 

Lit.: SACCARDO (1886: 366). 

typharum Desm., Pl. crypt. N. France, Ed. 1, Ser. 2, Fasc. VII, No. 304 (1828). 

T: on Typha sp. (Typhaceae), France, Desm., Pl. crypt. N. France 304 (K: isotype). 

100



Lit.: SACCARDO (1886: 366, 1899: 1081), LINDAU (1907: 813, 1910: 796), FERRARIS (1912: 

337), LIND (1913: 523), OUDEMANS (1919), GONZÁLES-FRAGOSO (1927: 211), DAVID (1997: 

59, 70). 

Notes: DAVID (1997): ‘The name C. typharum was introduced by Desmazieres in 1828, giving 

Dematium vulgare [sic] ?var. γ typharum Pers. as a synonym. …Authentic Persoon material 

has a mixed population of conidia comprising both C. herbarum and C. macrocarpum.’. 

‘typharum [Desm.] f. fuscum P. Karst.’, Hedwigia 35: 48 (1896)!. 

Notes: Cited by OUDEMANS (1919); see C. typarum var. fuscum. The actual reference in 

Hedwigia is to Brachysporium typharum [(Desm.) Karst.] var. fuscum n. var. 

typharum [Desm.] f. lanciforme (Ces.) Ferraris → lanciforme. 

typharum [Desm.] f. minor Brunaud, Bull. Soc. Bot. France 36: 340 (1889)!. 

T: on leaves of Typha latifolia (Typhaceae), France, Rochefort. 


‘typharum [Desm.] var. fuscum P. Karst.’, Hedwigia 35: 48 (1896)!. 

Lit.: SACCARDO (1899: 1081)!. 

Notes: See comment under Cladosporium typharum [Desm.] f. fuscum P. Karst. 

uleanum Henn., Hedwigia 34: 116 (1895)!. 

T: on living leaves of a Myrtaceae, Brazil, Uberaba in Minas Geraës, Jun. 1892, E. Ule, no. 

1927 (B; HBG: syntypes). 

Lit.: SACCARDO (1895: 620), LINDAU (1907: 828), LIND (1913: 524), OUDEMANS (1923). 

Notes: Excluded, taxonomic status remains unclear. 

ulmariae Grognot, in Roum., F. sel. gall. exs., Cent. XXXVII, No. 3697 (1886)!. 

T: on Spiraea ulmaria (= Filipendula ulmaria subsp. ulmaria) (Rosaceae), France, Roum., F. 

sel. gall. exs. 3697 (FH). 

Lit.: SACCARDO (1895: 621). 

umbrinum Fr., Syst. mycol. 3(2): 372 (1832)!. 

T: on Agaricus olearius, France, Montagne. 

= ? Botrytis pulvinata Link, in Willd., Sp. pl. 6(1): 61 (1824). 

Lit.: SACCARDO (1886: 369), LINDAU (1907: 807), OUDEMANS (1919), HEUCHERT et al. 

(2005: 57–58). 

unedonis Gonz. Frag., Mem. Real Acad. Ci. Barcelona, Ser. 3, 15(17): 459 (33) (1920)!. 

T: on living leaves, becoming dry, of Arbutus unedo (Ericaceae), Spain, near Barcelona, Las 

Planas, 28 Mar. 1918, Prof. A. Caballero (MA 06466). 



uniseptosporum Matsush., Icones Microfungorum a Matsushima Lectorum: 36 (1975)!. 

T: on rotten wood, Japan, Chiba, Kiyozumi Exp. Forest, Univ. Tokyo, Oct. 1967 (Matsush. 

herb. 2147). 

Lit.: HO et al. (1999: 142). 

Ill.: MATSUSHIMA (1975: Pl. 59, Fig. 3), HO et al. (1999: 143, Fig. 48). 

uredinicola Speg., Anales Mus. Nac. Hist. Nat. Buenos Aires 23: 122–123 (1912)!. 

T: on living acervuli of Puccinia cestri (Pucciniaceae) on Cestrum pubescens (Solanaceae), 

Argentina, Salta, near Calilegua, Nov. 1911, C. Spegazzini (LPS 13073: holotype; slides IMI 

87162a; AUA). 

Lit.: SACCARDO (1931: 798), SUTTON (1973: 40), ELLIS (1976: 330), ELLIS & ELLIS (1985: 

571, 1988), MORGAN-JONES & MCKEMY (1990), HO et al. (1999: 142), HEUCHERT et al. 

(2005: 41–46). 

Ill.: SUTTON (1973: 41, Fig. 19A), ELLIS (1976: 331, Fig. 249), MORGAN-JONES & MCKEMY 

(1990: 189, Pl. 1; 191, Fig. 1; 193, Fig. 2; 195, Fig. 3; 197, Pl. 2; 199, Pl. 3), HO et al. (1999: 

101

143, Fig. 49), HEUCHERT et al. (2005: 42–44, Figs 16–18). 

Host(s)/substrate(s) & distribution: hyperparasitic on telia and uredia of rusts, especially 

Cronartium and Puccinia, also on downy mildews and powdery mildew fungi; Asia (Hong 

Kong, India, Iran), Australia, Caribbean, Europe (Germany, Great Britain), New Zealand, 

North America (Canada, USA), South America (Argentina, Brazil). 

urediniphilum Speg., Anales Mus. Nac. Hist. Nat. Buenos Aires 31: 438–439 (1923)!. 

T: on living acervuli of Uredo cyclotrauma (Uredinales) on leaves of Pithecolobium 

cauliflorum, Paraguay, Asunción, Puerto Sajonia, Oct. 1919, C. Spegazzini. 

Lit.: DEIGHTON (1969), SACCARDO (1972: 1340), FARR (1973: 252), SUTTON (1973: 40), 

HEUCHERT et al. (2005: 58). 

Ill.: SUTTON (1973: 41, Fig. 19 B). 

uredinis Deighton – listed on CABI-page, refers to Cladosporiella uredinis Deighton, Mycol. 

Pap. 118: 36 (1969). 

ushuwaiense Speg., Bol. Acad. Nac. Ci. 27(4): 399 (1924)!, as ‘ushuwaiensis’. 

T: on dead leaves of Berberis ilicifolia (Berberidaceae), Argentina, Tierra del Fuego, 

Ushuwaia, 18 Jan. 1924, C. Spegazzini (LPS 13.144). 


Host(s)/substrate(s) & distribution: on Berberis ilicifolia; South America (Argentina). 

uvarum McAlpine, Add. fungi vine Australia: 47 (1898)!. 

T: on berries dried up and shrunken of Vitis vinifera (Vitaceae), Australia (VPRI). 

Lit.: SACCARDO (1899: 1079). 

Ill.: MCALPINE (1898: Figs 72–74). 

Notes: ZHANG et al. (2003) reported the species on Vitis vinifera from Shaanxi, China. 

vagans (Pers.) Desm., Pl. crypt. N. France, Ed. 1, Fasc. I, No. 6 (1825), as ‘Pers.’. 

T: on leaves of Acer, Tilia, etc. 

≡ Fumago vagans Pers., Mycol. eur. 1: 9 (1822)!. 


Notes: see C. fumago Link. 

vangueriae (Thirum. & Mishra) Arx, Genera Fungi Sporul. Pure Cult., Ed. 2: 222 (1974)!. 

T: on leaves of Vangueria spinosa (= Meyna laxiflora) (Rubiaceae), India, Bihar, Darbhanga 

(BPI 442756; IMI 51482). 

≡ Biharia vangueriae Thirum. & Mishra, Sydowia 7(1–4): 79 (1963)!. 

≡ Stenella vangueriae (Thirum. & Mishra) Deighton, Mycol. Pap. 144: 53 (1979)!. 

variabile (Cooke) G.A. de Vries, Contr. Knowl. Genus Cladosporium: 85 (1952)!. 

T: on Spinacia oleracea (Chenopodiaceae), Great Britain, Wales, Montgomeryshire, Welshpool, 

Forden Vicarage, J.E. Vize, Cooke, F. brit. exs. 360 (K: holotype). 

≡ Helminthosporium variabile Cooke, F. brit. exs., Ed. 2, No. 360 (1870), nom. inval. 

≡ Heterosporium variabile Cooke, Grevillea 5(35): 123 (1877)!. 

= Cladosporium subnodosum Cooke, Grevillea 17(83): 67 (1889)!, syn. nov. [T: BPI; NY; 

PH]. 

Lit.: ELLIS (1971: 315), ELLIS & ELLIS (1985: 429), DAVID (1995c, 1997: 94), HO et al. 

(1999: 144). 

Ill.: MINOURA (1966: 141, Fig. 5D), ELLIS (1971: 314, Fig. 217 B), DAVID (1995c: 1, Fig.; 

1997: 97, Fig. 26), FUENTES-DAVILA & GABRIELSON (1996: 54–55, Figs 1–2), HO et al. 

(1999: 145, Fig. 50). 

Host(s)/substrate(s) & distribution: on Spinacia oleracea; Asia (China, India, Iraq, Pakistan), 

Europe (Austria, Belgium, Denmark, France, Germany, Great Britain, Hungary, Italy, Montenegro, 

Netherlands, Norway, Romania, Spain, Turkey; Cyprus), North America (USA: CA, 

MA, VA, WA). 

velutinum Ellis & Tracy, J. Mycol. 6: 76 (1890)!. 

T: on leaves of Phalaris canariensis (Poaceae), USA, Mississippi, Starkville, 25 Mar. 1890, 

S.M. Tracy, No. 1323 (NY: holotype). Topotypes: 1 Apr. 1890, S.M. Tracy (BPI 427595, 

102

427597; NY) and 2 May 1890, S.M. Tracy (BPI 427596; NY). 


Entdeck. Gesammten Naturk. 7: 36 (1816). 

Lit.: SACCARDO (1892: 605). 

venturioides Sacc., Nuovo Giorn. Bot. Ital., N.S., 22: 71 (1915)!. 

T: on faded or dead branches of Amaranthus caudatus (Amaranthaceae), Malta, Zebbih, Jan. 

1914, no. 643 (PAD, according to GOLA 1930). 

Lit.: SACCARDO (1931: 796). 

Notes: ZHANG et al. (2003) reported the species on Amaranthus caudatus (China, Zhejiang) 

and A. tricolor (China, Hubei). 

venturioides [Sacc.] var. citricola Sacc., Nuovo Giorn. Bot. Ital., N.S., 22: 72 (1915)!. 

T: on leaves (‘in foliis morientibus’) of Citrus aurantium (Rutaceae), Malta, Imtahleb, C. 

Balzan, 1014, no. 501. 

Lit.: SACCARDO (1931: 796). 

versicolor P.A. Dang., Botaniste 22: 455 (1931)!. 

T: in a glass of water containing a piece of potato, France. 

Ill.: DANGEARD (1931: 489, Pl. 17; 491, Pl. 18). 

Notes: DANGEARD (1931) described and illustrated this species as a pycnidial fungus. 

versicolor T.E.T. Bond, Ceylon J. Sci., Sect. A, Bot. 12: 183 (1947), nom. illeg., homonym, 

non C. versicolor P.A. Dang., 1931. 

T: on Ageratum conyzoides (Asteraceae), India, Ceylon, St. Coombs, Dec. 1943 (IMI 676). 

= Cercospora perfoliati Ellis & Everh., J. Mycol. 5: 71 (1889), as ‘perfoliata’. [T: NY]. 

≡ Mycovellosiella perfoliati (Ellis & Everh.) Munt.-Cvetk., Lilloa 30: 201 (1960). 

≡ Passalora perfoliati (Ellis & Everh.) U. Braun & Crous, in Crous & Braun, Mycosphaerella 


Ser. 1: 314 (2003)!. 

= Cercospora agerati F. Stevens, Bernice P. Bishop Mus. Bull. 19: 154 (1925). [T: ILL 

16297]. 

≡ Ragnhildiana agerati (F. Stevens) F. Stevens & Solheim, Mycologia 23: 402 (1931). 

= Ramularia agerati Sawada, Special Publ. Coll. Agric. Natl. Taiwan Univ. 8: 190 (1959), 

nom. inval. [T: NTU-PPE, herb. Sawada; IMI 123997a (slide)]. 

= Cercosporella coorgica Muthappa, Mycopathol. Mycol. Appl. 34: 194 (1968). [T: IMI 

937100]. 

Lit.: DEIGHTON (1974: 69). 

victorialis (Thüm.) U. Braun & H.D. Shin, in Braun & Melnik, Proc. Komarov Bot. Inst. (St. 

Petersburg) 20: 101 (1997)!. 

T: on Allium victorialis (Alliaceae), Russia, West Sibiria, Mt. Kerlygan, Martjanov (LE 

40451: lectotype; LE 40452: isolectotype). 

≡ Cercospora victorialis Thüm., Hedwigia 21: 172 (1882)!. 

= Cladosporium alliicola H.D. Shin & U. Braun, Korean J. Mycol. 23(2): 141 (1995)!. 

Lit.: CROUS & BRAUN (2003: 422). 

Ill.: BRAUN & MELNIK (1997: Fig. 71). 

Host(s)/substrate(s) & distribution: ? Allium cepa, A. ochotense, ? A. sativum, A. victorialis; 

Asia (Korea, Russia), ? South America (Brazil). 

vignae M.W. Gardner, Phytopathology 15(8): 457 (1925)!. 

T: on Vigna sinensis (= V. unguiculata) (Fabaceae), USA, Indiana, LaFayette, Sept. 1924, 

M.W. Gardner (BPI 427608: lectotype). Isolectotype: BPI 427604. Topotype: BPI 427602. 

Lit.: DE VRIES (1952: 99), MORGAN-JONES & MCKEMY (1992), HO et al. (1999: 144). 

Ill.: MORGAN-JONES & MCKEMY (1992: 13, Fig. 1; 15, Pl. 1; 17, Fig. 2), HO et al. (1999: 145, 

Fig. 51). 

Host(s)/substrate(s) & distribution: on Vigna unguiculata, Lespedeza bicolor and Lespedeza 

thunbergii; Africa (South Africa, Zimbabwe), Asia (China), Australia (New South Wales, 

Queensland), North America (USA), South America (Brazil). 

103

vincae Fairm., Ann. Mycol. 9: 148 (1911)!. 

T: on dead leaves of Vinca minor (Apocynaceae), USA, New York, Lyndonville, 6 May 1910, 

C.E. Fairman [CUP-F2873(24-68)]. 


Entdeck. Gesammten Naturk. 7: 36 (1816). 

Lit.: SACCARDO (1931: 788). 

Ill.: FAIRMAN (1911: 150, Figs 3–5). 

vincae Moesz, Bot. Közlem. 23: 123 (1926)!, nom. illeg., homonym, non C. vincae Fairm., 

1911. 

T: on living leaves of Vinca herbacea (Apocynaceae) infected with Puccinia vincae (DC) 

Plowr., Hungary, near Budapest, 25 Apr. 1926, Dr. W. Moesz (W 10216: lectotype). 

= Cladosporium aecidiicola Thüm., Mycoth. univ., Cent. IV, No. 373 (1876)!. 

Lit.: SACCARDO (1972: 1340). 

Ill.: MOESZ (1926: Fig. 7). 

Notes: This species was described and illustrated as phytopathogenic, causing leaf spots, with 

conidiophores emerging through stomata. 

‘virescens Pers.’, Mycol. eur. 1: 14 (1822)!. 

Notes: Cladosporium virescens is cited in LINDAU (1907: 206) as synonym of Sporotrichum 

virescens (Pers.) Link (Bas.: Dematium virescens Pers.) which seems to be an error. PERSOON 

(1822) refers to Dematium virescens Pers.; the name Cladosporium virescens Pers. was never 

published. 

virgultorum Schwein., Trans. Amer. Philos. Soc., N.S., 4(2): 277 (1832)!. 

T: on branches, USA, Pennsylvania, Bethlehem, No. 2605 (PH 01020411, 01020412: syntypes). 

Lit.: SACCARDO (1886: 356). 

viride (Fresen.) Z.Y. Zhang & T. Zhang, Proceedings of Phytopathological Symposium 

Organized by Phytopathology Laboratory of Yunnan Province 2: 306 (1998). 

T: on rotten fruits of Vitis (Vitaceae). 

≡ Penicillium viride Fresen., Beitr. Mykol. 1: 21 (1850)!. 

≡ Hormodendrum viride (Fresen.) Sacc., Syll. fung. 4: 311 (1886). 


Notes: recorded from China (Jiangxi) on Vitis balanseana. 

viridiolivaceum Pidopl. & Deniak → transchelii var. viridi-olivacearum. 

viridiolivaceum [Pidopl. & Deniak] var. semenicola Pidopl. & Deniak → transchelii var. 

semenicola. 

viticola Ces., in Rabenhorst, Flora 37: 206 (1854)! and in Klotzsch, Herb. viv. myc., Cent. 

XIX, No. 1877 (1854)!, as ‘viticolum’. 

T: on Vitis sp. (Vitaceae), Italy, Klotzsch, Herb. viv. myc. 1877 (e.g., BPI 797134; HAL: 

syntypes). 

≡ Cercospora viticola (Ces.) Sacc., Syll. fung. 4: 485 (1886)!. 

= Pseudocercospora vitis (Lév.) Speg., Anales Mus. Nac. Buenos Aires 20: 438 (1910)!. 

Lit.: LINDAU (1910: 116, as ‘viticolum’), CHUPP (1954: 605), SIVANESAN (1984: 210). 

vitis (Lév.) Sacc., Mycoth. ven., Cent. III, No. 284 (1875)!. 

T: on leaves of Vitis vinifera (Vitaceae), Italy, Treviso, Selva, Sept. 1874, Sacc., Mycoth. ven. 

284 (HAL). 

≡ Septonema vitis Lév., Ann. Sci. Nat. Bot., Sér. 3, 9: 261 (1848)!. 

≡ Cercospora vitis (Lév.) Sacc., Nuovo Giorn. Bot. Ital. 8: 188 (1876)!. 

≡ Helminthosporium vitis (Lév.) Pirotta, Rev. Mycol. (Toulouse) 11: 185 (1889)!. 

≡ Pseudocercospora vitis (Lév.) Speg., Anales Mus. Nac. Buenos Aires 20: 438 (1910)!. 

≡ Phaeoisariopsis vitis (Lév.) Sawada, Rep. Dept. Agric. Gov. Res. Inst. Formosa 2: 164 

(1922). 

104

≡ Cercosporiopsis vitis (Lév.) Miura, Flora of Manchuria and East Mongolia, III. Cryptog. 

Fungi: 527 (1928). 

= Cladosporium viticola Ces., in Rabenhorst, Flora 38: 206 (1854)! and in Klotzsch, Herb. 

viv. myc., Cent. XIX, No. 1877 (1854)!, as ‘viticolum’. 

≡ Cercospora viticola (Ces.) Sacc., Syll. fung. 4: 485 (1886)!. 

= Cladosporium ampelinum Pass., Erb. Critt. Ital., Ser. 2, No. 595 (1872)!. 

= Graphium clavisporium Berk. & Cooke, Grevillea 3(27): 100 (1874)!. [T: K]. 

≡ Isariopsis clavispora (Berk. & Cooke) Sacc., Syll. fung. 4: 631 (1886)!. 

= Cercospora vitis [(Lév.) Sacc.] var. rupestris Cif., Ann. Mycol. 20: 45 (1922). 

= Cercospora vitis f. parthenocissi Docea, Lucr. Ști. Inst. Agron. ‘N. Bălescu’, Ser. A., 11: 

406 (1968). 

Teleomorph: Mycosphaerella personata B.B. Higgins, Amer. J. Bot. 16: 287 (1929). 

Lit.: LINDAU (1910: 116, as ‘vitis Sacc.’), CHUPP (1954: 605), DEIGHTON (1976: 131), 

SIVANESAN (1984: 210), CROUS & BRAUN (2003: 427). 

vitis-frutigeni, in herb. 

On Vitis sp. (Vitaceae), USA, North Carolina, 17 Sept. 1890, G.E. Boggs (BPI 427626). 

vulgaris – PACMA lists one collection in CM herbarium. 

wedrila – on CABI page without author and citation, in herb.? (Kirk et al. n.d.). 

werneckii Horta, Revista Med. Cirugía Brasil 29: 274 (1921), as ‘Wernecki’. 

T: on man, Brazil. [ATCC 36317: ex-type]. 

≡ Dematium werneckii (Horta) C.W. Dodge, Med. Mycol.: 676 (1935)!. 

≡ Pullularia werneckii (Horta) G.A. de Vries, Contr. Knowl. Genus Cladosporium: 101 

(1952)!. 

≡ Exophiala werneckii (Horta) Arx, Genera Fungi Sporul. Pure Cult.: 180 (1970)!. 

≡ Hortaea werneckii (Horta) Nishim. & Miyaji, Jap. J. Med. Mycol. 26(2): 145 (1984). 

≡ Phaeoannellomyces werneckii (Horta) McGinnis & Schell, in McGinnis, Schell & Carson, 

Sabouraudia 23(3): 184 (1985). 

= Cryptococcus metaniger Castell., Archiv Dermatol. Syph. 16(4): 402 (1927)!. 

≡ Cladosporium metaniger (Castell.) Ferraris, Atti Ist. Bot. “Giovanni Briosi” 3: 183 (1932)!. 

= Cladosporium rietmanni Sart. & Syd., Rev. Pat. Malad. Pays Chauds 15(1): 9–44 (1935). 

Lit.: NANNIZZI (1934: 408), DE VRIES (1952: 100), COOKE (1962: 34), KWON-CHUNG & 

BENNETT (1992: 195), SCHELL (2003: 606). 

wikstroemiae (Sawada) H. Zhang & Z.Y. Zhang, Proceedings of Phytopathological 

Symposium Organized by Phytopathology Laboratory of Yunnan Province 2: 306 (1998), 

comb. inval. 

T: on Wikstroemia indica (Thymeleaceae), Taiwan, Taipei, 19 Feb. 1913, K. Sawada (TNS 

F218930: lectotype; PPMH: isolectotype). 

≡ Heterosporium wikstroemiae Sawada, Rep. Gov. Res. Inst. Formosa 87: 77 (1944), nom. 

inval. et illeg., homonym, non H. wikstroemiae Petch, 1922. 

= Heterosporium wikstroemiae Petch, Ann. Roy. Bot. Gard. (Peradeniya) 7: 319 (1922). [T: 

K; PAD]. 

≡ Stenella wikstroemiae (Petch) J. Walker, in Walker & White, Mycol. Res. 95: 1010 (1991)!. 

Lit.: DAVID (1997: 126). 

xyridis Tracy & Earle, Bull. Torrey Bot. Club 23(5): 206 (1896)!, as ‘Gladisporium (sic) 

xyridis’. 

T: on petals of Xyris fimbriata (Xyridaceae), USA, Mississippi, Ocean Springs, 29 Sept. 1895, 

F.S. Earle (NY: lectotype). Isolectotypes: BPI 427627–427628; NY. 

Lit.: SACCARDO (1899: 1081). 

Host(s)/substrate(s) & distribution: on species of Xyris, including X. fimbriata and X. jupicai; 

Caribbean (Cuba), North America (USA). 

105

zeae Lobik, Trudy Severo-Kavkazsk. Inst. Zashch. Rast. 1(2): 41 (1933), nom. illeg., homonym, 

non C. zeae Peck, 1894. 

T: on Zea mays (Poaceae), former USSR, Caucasus (LE). 

zeae Peck, Rep. (Annual) New York State Mus. Nat. Hist. 46: 114 (1894). 

T: on unripened grains of Zea mays (Poaceae), USA, Menands, Sept., C.H. Peck (NYS 3441: 

holotype). 

Lit.: SACCARDO (1895: 620). 

zeylanicum Sacc. & Trotter → compactiusculum. 

zizyphi P. Karst. & Roum., Rev. Mycol. (Toulouse) 12(46): 78 (1890)!, non Pseudocercospora 

zizyphi (Petch) Crous & Braun, 1996. 

T: on faded leaves of Zizyphus (Rhamnaceae), ‘ad Sontag’, Dec. 1887 (PC and Roum., F. sel. 

exs. 5500). 

= Cercospora jujubae S. Chowdhury, Indian J. Agric. Sci. 16: 525 (1946). [T: IMI 113803]. 

≡ Pseudocercospora jujubae (S. Chowdhury) N. Khan & Shamsi, Bangladesh J. Bot. 12: 117 

(1983)!. 

Lit.: SACCARDO (1892: 604), FERRARIS (1912: 342), OUDEMANS (1923), CROUS & BRAUN 

(2003: 233). 

Unnamed Cladosporium states of named teleomorphs: 

Cladosporium state of Apiosporina collinsii (Schwein.) Höhn. 

≡ Fusicladium state of Apiosporina collinsii (Schwein.) Höhn. 

Lit.: SIVANESAN (1984: 598), SCHUBERT (2001), SCHUBERT et al. (2003: 105–106). 

Acknowledgements: The authors gratefully acknowledge the cooperation of the following 

institutions in locating type and authentic material: B, BPI, CUP, DAOM, FH, FLAS, GENT, 

GZU, HBG, ILL, IMI, K, LE, LEP, M, MA, NY, NYS, PAD, PH, PPMH, PRM, RO, S, 

SIENA, VPRI, WIS. We are very grateful to the directors and curators. And we are much 

obliged to P.W. Crous (Utrecht, the Netherlands), J.H. Cunnington (Burnley, Australia), J. 

David (Egham, UK), Guo (China), S. de Hoog (Utrecht, the Netherlands), V.A. Melnik (St. 

Petersburg, Russia), J. Mouchacca (Paris, France), D. Seidel (Halle, Germany) and Z.Y. 

Zhang (Kunming, China) for the support of our search for rare publications. 

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Addresses of authors: 

Dr. F.M. Dugan, USDA-ARS, 59 Johnon Hall, Washington State University, Pullman WA 

99164-6402 USA. e-mail: fdugan@mail.wsu.edu. 

K. Schubert and Prof. U. Braun, Martin-Luther-University, Institute for Geobotany and 

Botanical Garden, Herbarium, Neuwerk 21, D-06099 Halle, Germany. 

e-mail: schubert@botanik.uni-halle.de and braun@botanik.uni-halle.de. 

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