Rust Phylogenetics - Plant Management Network
Rust Phylogenetics - Plant Management Network
Rust Phylogenetics - Plant Management Network
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<strong>Rust</strong> <strong>Phylogenetics</strong><br />
Recent advances in systematics, taxonomy,<br />
and evolution of rust fungi<br />
M. Catherine Aime<br />
Louisiana State University AgCenter<br />
<strong>Plant</strong> Pathology & Crop Physiology
Systematics<br />
of Pucciniomycotina
Pucciniomycotina<br />
• <strong>Rust</strong>s and their allies<br />
• ~8,000 species, i.e., 1/3 rd of described Basidiomycota<br />
• Mostly parasitic: phytopathogens (e.g., Pucciniales), entomoparasites entomoparasites<br />
(e.g.,<br />
Septobasidium), Septobasidium),<br />
mycoparasites (e.g., Tuberculina)<br />
Tuberculina<br />
• Most higher-level higher level groups not formally recognized or named until 2006<br />
Figures from: Aime et al. 2006. (2007). Mycologia 98:896-905.
Pucciniales<br />
Aime et al. 2006. An overview of the higher-level classification of<br />
Pucciniomycotina based on combined analyses of nuclear large<br />
and small subunit rDNA sequences. Mycologia 98(6):896-905<br />
Pucciniomycetes<br />
Pucciniomycotina<br />
Pucciniomycotina<br />
Combined nuc-lsu nuc lsu & nuc-ssu nuc ssu<br />
8 classes, 18 orders<br />
Pucciniomycotina
Pucciniales<br />
Aime et al. 2006. An overview of the higher-level classification of<br />
Pucciniomycotina based on combined analyses of nuclear large<br />
and small subunit rDNA sequences. Mycologia 98(6):896-905<br />
Pucciniomycetes<br />
Pucciniomycotina<br />
Pucciniomycetes:<br />
© Matthias Lutz:<br />
Bio Pro website<br />
Pucciniomycotina
Systematics<br />
of Pucciniales
Pucciniales Clem. & Shear (1931) =<br />
rust fungi (= Uredinales)<br />
All obligate pathogens (largest natural group of<br />
phytopathogens)<br />
Most Most complex life cycles in Eumycota (in Eukaryotes?)<br />
Very Very plastic (morphologically; genomically—haploid<br />
genomically haploid<br />
estimates in Pucciniaceae alone from 64-418 64 418 Mbp)<br />
Life Life cycle data incomplete or missing for most<br />
AFTOL classification: Hibbett DS et al. 2007. A higher-level phylogenetic<br />
classification of the Fungi. Mycological Research 111:509-547.
Pucciniales Taxonomy<br />
~ 7000 described species<br />
~ 150 genera<br />
~ 13 families<br />
CONTENTIOUS.<br />
CONTENTIOUS.<br />
1. “primitive primitive” hosts harbor “primitive primitive” rusts<br />
2. Telia<br />
3. Spermogonia morphology<br />
3.<br />
Cummins & Hiratsuka. 2003. Illustrated Genera of the <strong>Rust</strong> Fungi
<strong>Rust</strong> Molecular Systematics<br />
Sjamsuridzal et al. 1999. Mycoscience 40:21-27. 40:21 27.<br />
7 taxa Pucciniales; 18S rDNA seqs.<br />
Fern rusts are not ancestral<br />
Maier et al. 2003. Can. J. Bot. 81:12-23. 81:12 23.<br />
54 taxa Pucciniales (all temperate); nLSU rDNA seqs.<br />
Recovered: Melampsoraceae; Pucciniastraceae s.l.; s.l. ;<br />
Phragmidiaceae; Gymnosporangium; Gymnosporangium;<br />
Pucciniaceae;<br />
Tranzschelia-Ochropsora<br />
Tranzschelia Ochropsora<br />
Melampsora as basal<br />
Wingfield et al. 2004. Australasian <strong>Plant</strong> Pathology 33:327-335. 33:327 335.<br />
64 taxa Pucciniales; 18S rDNA seqs.<br />
Recovered: 2 families<br />
Racospermyces as basal
Pucciniales<br />
rDNA LSU<br />
1229 bp.; 857 incl. char.; 263 PI char<br />
470 seq; 350 taxa (+4 outgroup)<br />
13 of 13 families (100%)<br />
58 of 117 genera (49.5%)<br />
~500 of 7000 species (7%)<br />
(classifications fide<br />
Cummins & Hiratsuka 2003)<br />
1 change<br />
Helicobasidium purpureum AY885168<br />
Eocronartium muscicola AY512844<br />
Puccinia mariae-wilsoniae U-005 Claytonia (Portulaceae I-III) MD<br />
Puccinia mariae-wilsoniae MCA2497 Claytonia (Portulaceae) MD<br />
Puccinia lagenophorae U-384 (#569) Asteraceae (Osteocarpum) South Africa<br />
Puccinia lagenophora U-128 Senecio (I-III Asteraceae) Tunisia<br />
Puccinia lagenophorae U-107 Senecio (I, III Asteraceae) Argentina<br />
Puccinia violae U-023 Viola (Violaceae II-III) Norway<br />
Puccinia violae U-269 Viola (Violaceae II-III) NY<br />
Uromyces sp. MCA 2870 Juncaceae (Juncus? III) VA<br />
Puccinia? U397 Asteraceae MD<br />
Puccinia? U-390 Asteraceae MD<br />
Puccinia cf. violae U-4 Violaceae (Viola cucullata )-I, autoecious) MD<br />
Uromyces vignae? U-015 Macroptilium (Fabaceae II) Argentina<br />
Uromyces vignae JRH-472 Fabaceae (Vigna II-III) Puerto Rico<br />
Uromyces vignae JRH-468 Fabaceae (Vigna II-III) Puerto Rico<br />
Uromyces vignae U-125 Vigna (Fabaceae II) C.R.<br />
Uromyces vignae U-126 Vigna (Fabaceae II) C.R.<br />
Uromyces appendiculatus U-122 Phaseolus (II Fabaceae) C.R.<br />
Uromyces appendiculatus U-123 Phaseolus (II Fabaceae) Mexico<br />
Uromyces appendiculatus U-00123 Phaseolus (Fabaceae) Mexico<br />
Uromyces neotropicalis sp.nov MCA2533 Cucurbitaceae II,III Ecuador<br />
Dietelia mesoamericana sp.nov. U-320 (W1909) Asteraceae (M. micrantha microcyclic I) Mexico<br />
Dietelia portoricensis u-322 (no #) Asteraceae (Mikania micrantha I-microcyclic) Costa Rica<br />
Puccinia granularis U-443 Geraniaceae (Pelargonium O,I,II,III) South Africa<br />
Puccinia granularis U-444 (#581) Geraniaceae (Pelargonium II,III) South Africa<br />
Puccinia granularis U-444 (#581) Geraniaceae (Pelargonium II,III) South Africa<br />
Puccinia granularis U-443 (#580) Geraniaceae (Pelargonium O,I,II,III) South Africa<br />
Puccinia spegazzinii JRH-226 Asteraceae (Mikania micrantha III) Honduras<br />
Puccinia spegazzinii U-319 (W2102) Asteraceae (Mikania micrantha microcyclic III) Peru<br />
Puccinia spegazzinii U-315 (W2020) Asteraceae (Mikania micrantha IIImicrocyclic) Argentina<br />
Dietelia mesoamericana sp. nov. U-321 (mislabelled as U143) Mikania Mexico<br />
Puccinia hydrocotyles U-079 Hydrocotyle (II-III Apiaceae) AL<br />
Puccinia? MCA-2864 Asteraceae (Helianthus) VA<br />
Puccinia pimpinellae? MCA2576 Apiaceae( Osmorhiza I) ID<br />
Puccinia pimpinellae U-43 Osmorhiza (Apiaceae III) UT<br />
Puccinia pimpinellae MCA2604 Osmorhiza (Apiaceae III) TN<br />
Puccinia berkheyicola U-314 Berkheya (Asteraceae) South Africa<br />
Puccinia notobasidis U-229 Cirsium (Asteraceae) Turkey<br />
Puccinia variabilis var. lapsanae U-37 Asteraceae (Lapsana II,III) UT<br />
?? U-412 Asteraceae (Youngia) Guam<br />
?? JRH199 Wedelia (Asteraceae) Hondorus (inc seq)<br />
Puccinia menthae MCA 2601 Monarda (Lamiaceae II) TN<br />
Puccinia menthae U-343 Origanum (Lamiaceae II) Argentina<br />
Puccinia menthae MCA 2601 Monarda (Lamiaceae II) TN<br />
Puccinia marylandica MCA2602 Sanicula (Apiaceae II-III) TN<br />
Puccinia marylandica MCA2606 Sanicula (Apiaceae II-III?) TN<br />
Puccinia commelinae U-049 Commelina (Commelinaceae II) Guyana<br />
Puccinia synedrellae JRH152 Synedrella (III Asteraceae) Costa Rica<br />
Puccinia cnici-oleracei GUY-6 Asteraceae (Emilia sonchifolia III) Guyana<br />
Puccinia synedrellae JRH-152 Asteraceae (Synedrella nodiflora III) Costa Rica<br />
Puccinia cnici-oleracei JRH-204 Asteraceae (Emilia fosbergii III) Honduras<br />
Puccinia cnici-oleracei JRH-490 Asteraceae (Emilia fosbergii III) Puerto Rico<br />
Puccinia MCA-2864 Asteraceae (Helianthus) VA<br />
Uromyces ficariae AF426204 Ranunculus (Ranunculaceae)<br />
Puccinia sp. MCA2575 Asteraceae Idaho<br />
Puccinia sp. MCA-2575 Asteraceae (II) ID<br />
Puccinia chondrillina U-202 Asteraceae (Chondrilla) Turkey [BPI extraction]<br />
Puccinia balsamorrhizae U-332Asteraceae (Balsamorhiza III?) B.C.<br />
Puccinia balsamorrhizae U-354 Asteraceae (Balsamorhiza) ID<br />
Puccinia balsamorrhizae U-355 Asteraceae (Balsamorhiza) ID<br />
Miyagia pseudosphaeria U-63 (BPI) Asteraceae (Sonchus oleraceus)<br />
Puccinia calcitrapae v. centaureae U-201 Asteraceae (Centaurea depressa) Turkey [BPI extraction]<br />
Puccinia calcitrapae U-218 Carduus (Asteraceae) Turkey<br />
Puccinia calcitrapae U-218 Asteraceae (Carduus nutans) Turkey<br />
Puccinia carthami complex MCA-2881 Asteraceae (Cirsium?) Costa Rica<br />
Puccinia acroptili U-224(confirmed no.) Asteraceae (Acroptilon) Turkey<br />
Puccinia acroptili U-578 Asteraceae (Acroptilon repens III) NM<br />
Puccinia acroptili U-135 (Dana Berner 02-048) Asteraceae (Acroptilon repens II-III?) Turkey<br />
Puccinia acroptili U-224 Asteraceae (Acroptilon repens) Turkey<br />
Puccinia acroptili U-577 Asteraceae (Centaurea repens) NM<br />
Puccinia U-203 Asteraceae (Cardueae II-III) Turkey [host & rust confirmed from BPI]<br />
Puccinia punctiiformis U-225 Cirsium (Asteraceae) Turkey<br />
Puccinia punctiiformis U-225 Asteraceae (Cirsium arvense) Turkey<br />
Puccinia calcitrapae U-164 Asteraceae (Carthamus tinctorius II-III) Argentina<br />
Puccinia centaurea U-278 (FDWSRU 04-010) Asteraceae (Centaurea hybrid) OR<br />
Puccinia calcitrapae v. centaureae U-140 (Dana Berner 03-051) Asteraceae (Centaurea macedonica) Greece<br />
Puccinia calcitrapae v. centaurea U-209 Asteraceae (Centaurea calcitrapa) Turkey<br />
Puccinia carthami U-323 Asteraceae (Carthamus tinctorius III) Oman<br />
Puccinia atropuncta MCA2609 Veratrum (Liliaceae III) TN<br />
?? U492 Antirrhinum (Scrophulariaceae) Germany<br />
?? MCA2529 Geranium (Geraniaceae II) Ecuador<br />
Endophyllum stachytarphetae GUY-55 Stachytarpheta (III Verbenaceae)<br />
?? MCA2558 Phragmites? (Poaceae) Ecuador<br />
Endophyllum cassiae U-525 Fabaceae (Cassia III?) Nigeria<br />
?? JRH-466 Cyperaceae Puerto Rico<br />
?? U-265 Cyperus (Cyperaceae II?) Guam<br />
Puccinia cnici-oleracei JRH-200 Asteraceae (Melampodium divaricatum III) Honduras<br />
Puccinia cnici-oleracei JRH-207 Asteraceae (Malampodium divaricatum III) Honduras<br />
Puccinia ursiniae U-463 (#590) Asteraceae (Ursinia) South Africa<br />
Puccinia tanaceti U-040 Artemisia (Asteraceae II-III) UT<br />
Puccinia rubefaciens? U-330 Galium (Rubiaceae III?) B.C. Canada<br />
Puccinia medellinensis JRH-154 Lamiaceae (Hyptis II) Costa Rica<br />
Puccinia melampodii JRH138 Emilia (Asteraceae III) Costa Rica<br />
Uromyces rumicis U-227 Rumex (Polygonaceae) Turkey<br />
Uromyces lespedezae-procumbentis U-007 Trifolium (Fabaceae III) MD<br />
Puccinia convolvuli? MCA2778 Calystegia (Convolvulaceae II) MD<br />
Puccinia convolvuli U-068 Calystegia (II-III Convolvulaceae) MD<br />
Puccinia scleriicola GUY-57 Scleria (I Cyperaceae)<br />
Puccinia ibrae JRH-228 Convolvulaceae (Ipomoea I-III) Honduras<br />
Puccinia thaliae JRH141 Canna (Cannaceae II) Costa Rica<br />
Puccinia thaliae GUY-3 Canna (Cannaceae II) Guyana<br />
Puccinia kalchbrenneriana U-312 (#560) Asteraceae (Helichrysum III?) South Africa<br />
Puccinia kalchbrenneriana U-313 (#555) Asteraceae (Helichrysum III?) South Africa<br />
Puccinia kalchbrenneri? U-258 (#539) Asteraceae (Helichrysum III) South Africa<br />
Uromyces colombianus JRH-145 Asteraceae (Melanthera II-III) Costa Rica<br />
Uromyces psoraleae U-029 Psoralidium (Fabaceae I-III) UT<br />
Uromyces sp. JRH-482 Cucurbitaceae (Cayaponia) Puerto Rico<br />
Puccinia gigantea AF426198 Epilobium (Onagraceae)<br />
Puccinia enceliae JRH-166 Asteraceae (Tithonia II-III) Costa Rica<br />
Puccinia thunbergiae U-260 (543) Acanthaceae (Thunbergia O,I,III) South Africa<br />
Puccinia hydrocotyles U-105 Hydrocotyle (Apiaceae) Argentina<br />
Uromyces galegae U-242 Galega (Fabaceae) Turkey<br />
Uredo cajani U-119 Cajanus (Fabaceae II) Honduras<br />
Uredo cajani MCA-2895 Fabaceae (Cajanus) Costa Rica<br />
Uredo cajani JRH-489 Fabaceae (Cajanus II) Puerto Rico<br />
Uromyces dolicholi JRH-469 Fabaceae (Rhynchosia II) Puerto Rico<br />
Uromyces scutellatus U-204(presumed) Euphorbiaceae (Euphorbia) Turkey<br />
Uromyces euphorbiae GUY-7 Euphorbia (II Euphorbiaceae)<br />
Uromyces pisi AF426201 Euphorbia<br />
Uromyces pisi-sativi? MCA2377 Euphorbia (Euphorbiaceae) Italy<br />
Uromyces? U-471 Euphorbiaceae (Euphorbia) Germany<br />
Endophyllum euphorbiae-sylvaticae AF426200 Euphorbia<br />
Puccinia abrupta JRH-146 Viguiera (Asteraceae II-III) Costa Rica<br />
Uromyces viciae-fabae AF426199 Vicia (Fabaceae)<br />
Puccinia sp.nov.? U-210 Tragopagon (Asteraceae I-III) Turkey<br />
Puccinia stoboeae U-259 (#550) Asteraceae (Berkheya O,I,II) South Africa<br />
Uromyces hedysari-paniculati U-124 Desmodium (Fabaceae II-III ) Honduras<br />
Uromyces hedysari-paniculati U-052 Desmodium (Fabaceae II) Guyana<br />
Aecidium sp. U-211 Salsola (Chenopodiaceae I) Turkey<br />
Uromyces ciceris-arietini U-183 Cicer (Fabaceae II-III ) SD<br />
Uromyces junci AF426203 Pulicaria (Asteraceae)<br />
Puccinia? MCA2593 (Asteraceae O?) ID<br />
Puccinia diplachnis MCA2375 Cirsium (I Asteraceae) Switzerland {T=Poaceae}<br />
Puccinia verruca U-130 Centaurea (Asteraceae III) Tunisia<br />
Puccinia collinsiae U-044 Collinsiae (Scrophulariaceae) UT<br />
Uromyces trifolii-repentis U-207 Trifolium (Fabaceae) Turkey<br />
Puccinia urticae-acutiformis AF426202 Carex (Cyperaceae)<br />
Puccinia hemerocallidis U498 Hemerocallis (Lilioaceae) TX<br />
Puccinia hemerocallidis U-072 Hemerocallis (II Liliaceae) AL<br />
Puccinia hemerocallidis U-073 Hemerocallis (II Liliaceae) AL<br />
Uromyces euryopsidicola U-381 (#575) Asteraceae (Euryops III) South Africa<br />
Uromyces euryopsidicola U-216 (#295) Asteraceae (Euryops) South Africa<br />
Uromyces euryopsidicola U-261 (#295) Asteraceae (Euryops) South Africa<br />
Uromyces euryopsidicola U-216 (#295) Asteraceae (Euryops) South Africa<br />
Puccinia sparganioides U-368 Poaceae (Spartina II,III) NY<br />
Puccinia buxi U-418 Buxus (Buxaceae) Georgia<br />
Puccinia tillandsiae U419 Tillandsia (Bromeliaceae) Guatemala<br />
Puccinia arechaveletae MCA2430 Serjania (Sapindaceae) Guyana<br />
Puccinia arechavelaete GUY-66 Serjania (III Sapindaceae)<br />
Puccinia arechavaletae JRH175 Unvillea (II-III Sapindaceae) Costa Rica<br />
Puccinia physalidis U-189 Physalis (Solanaceae III) ND<br />
Uromyces ixiae U452 Romulea (Iridaceae) S. Africa<br />
Uromyces transversalis U-386 Iridaceae (Gladiolus) Mexico<br />
Puccinia hordei MCA2391 (Poaceae) CA<br />
Puccinia? U-288 Poaceae (Pennisetum II) Guam<br />
Uromyces setaria-italicae JRH220 Melinis (II Poaceae) Hondorus<br />
Uromyces setaria-italicae JRH205 Panicum (II Poaceae) Hondorus<br />
Uromyces setaria-italicae JRH202 Panicum (II Poaceae) Honduras<br />
Puccinia gouaniae JRH159 Gouania (II Rhamnaceae) Costa Rica<br />
Puccinia arundinariae? MCA2494 Arundinaria (Poaceae) VA<br />
Puccinia purpurea JRH198 Sorghum (Poaceae II-III) Honduras<br />
Puccinia purpurea JRH213 Sorghum (Poaceae II) Hondurus<br />
Puccinia purpurea JRH218 Sorghum (Poaceae II) Hondurus<br />
Puccinia duthiae GUY8 Bothriochloa (Poaceae II) Guyana<br />
Puccinia duthiae U-264 Dichanthium (Poaceae II) Guam<br />
Puccinia exhauriens U-538 (#600) Oleaceae (Jasminum) South Africa<br />
Puccinia exhauriens U-539 (#605) Oleaceae (Jasminum) South Africa<br />
Puccinia moraeae U462 Moraea (Iridaceae) S. Africa<br />
Uromyces massoniae? U-446(#586) Liliaceae (Massonia II-III) South Africa<br />
Uromyces salsolae U-134 Salsola (Chenopodiaceae) Syria<br />
Puccinia sp. U-006 Allium? (Liliaceae III) MD<br />
Uromyces aloes U-00372 (#567) Aloaceae (Aloe) South Africa<br />
Puccinia obliquo-septata MCA2431 Olyra (Poaceae) Guyana<br />
Puccinia obliquo-septata GUY-10 Olyra (II Poaceae)<br />
Puccinia peperomiae JRH192 Piper (II Piperaceae) Costa Rica<br />
Puccinia fallax MCA2305 Palicourea (Rubiaceae II-III) Guyana<br />
Puccinia fallax GUY-58 Palicourea (II Rubiaceae)<br />
Sphenospora kevorkianii U-10 Orchidaceae (Stanhopea II) Peru<br />
Puccinia coronata U-583 Poaceae (Avena) ND<br />
Puccinia coronata U400 Festuca (Poaceae II, III) VA<br />
Puccinia coronata U-244 Rhamnus (Rhamnaceae O-O) ND<br />
Puccinia coronata U-245 Scolochloa (Poaceae III?) ND<br />
Puccinia coronata AF426207 Rhamnus (Rhamnaceae)<br />
Puccinia pampeanum U-109 Salpichroa (III Solanales) Argentina<br />
Endophyllum pampeanum U-108 Salpichroa (III Solanales) Argentina<br />
Uromyces gageae AF426208 Gagea (Liliaceae)<br />
Aecidium wiborgiae U-380 (#576) Fabaceae (Wiborgia) South Africa<br />
Aecidium wiborgiae U-379 (#573) Fabaceae (Wiborgia) South Africa<br />
Uromyces sp. MCA2383 Lilium (III Liliaceae) Switzerland<br />
Puccinia mogiphanis U-111 Alternanthera (Amaranthaceae) Nigeria<br />
Puccinia stonemanniae U-310 Thesium (Santalaceae III?) South Africa<br />
Puccinia sorghi U-362 Poaceae (Zea mays II, III) VA<br />
Uromyces polygoni U-214 Polygonum (Polygonaceae) Turkey<br />
Uromyces polygoni U-214 Polygonum (Polygonaceae) Turkey<br />
Puccinia mogiphanis JRH 180 Amarantaceae (Alternanthera II) Costa Rica<br />
Puccinia cynodontis JRH 219 Cynodon (Poaceae II-III) Hondorus<br />
Aecidium kalanchoe U-18 Kalanchoe<br />
Puccinia horiana U-391 Asteraceae (Chrysanthemum III) DE<br />
Puccinia horiana U-392 Asteraceae (Chrysanthemum III) MD<br />
Puccinia horiana U-070 Chrysanthemum (Asteraceae) NJ<br />
Aecidium sp. U-465 (#579) Fabaceae (Lebeckia) South Africa<br />
Puccinia arenariae U142 Dianthus (Caryophyllaceae) Ecuador<br />
Puccinia graminis U-478 Berberis (Berberidaceae) Germany<br />
Puccinia.poarum FROM BRANDON on Tussilago (iii on Poaceae)<br />
Puccinia MCA-2882 Asteraceae? Costa Rica<br />
Puccinia MCA-2886 Asteraceae? Costa Rica<br />
Puccinia graminis (as Aecidium sp.) PDD78317 Berberis I N.Z.<br />
Puccinia graminis U-469 Agropyron (Poaceae) Germany<br />
Puccinia graminis U-470 Agropyron? (Poaceae) Germany<br />
Puccinia graminis U-470 Poaceae (Agropyron?) Germany<br />
Uredo baruensis GUY-71 Sapotaceae (Chrysophyllum II) Guyana<br />
[IDd as Coleosporium on Merremia (Convolvulaceae)] JRH-484 P.R. (seq. confirmed 2X)<br />
Sphaerophragmium acacia JRH-486 Fabaceae (Albizia II-III) Puerto Rico<br />
Sphaerophragmium acaciae JRH-470 Fabaceae (Albizia II) Puerto Rico<br />
Pileolaria toxicodendri AY 745698 (AFTOL ID 988)<br />
Pileolaria toxicodendri u-607 Anacardiaceae (Toxicodendron) MN<br />
Puccinia tetragonia U-442 Aizoaceae (Tetragonia) South Africa<br />
?? U-476 Allium (Liliaceae) Germany<br />
Uromyces albucae U-447 (#591) Liliaceae (Albuca) South Africa<br />
Puccinia dispersa U-234 Anchusa (Boraginaceae) Turkey<br />
Puccinia? MCA-2885 ?? Costa Rica<br />
Chrysocyclus cestri U-156 Cestrum (Solanaceae III) Argentina<br />
Puccinia smilacis U-12 Smilacaceae (Smilax II-III) MD<br />
Puccinia smilacis U-393 Smilacaceae (Smilax) MD<br />
Maravalia sapotae JRH-172<br />
Stereostratum corticiodes U-027 Bambusa (Poaceae II-III) China<br />
Puccinia asarina MCA2374 Asarum (III Aristolochiaceae) Italy<br />
Puccinia pampeana JRH135 Capsicum (Solanaceae II-III aecioid) Costa Rica<br />
Endophyllum macowanii U-255 Rhamnus (Rhamnaceae III?) South Africa<br />
Puccinia thlaspeos U-054 Boechera (III Brassicaceae) WA<br />
Uromyces bulbinis BRIP-28988 Liliaceae (Bulbine) Australia<br />
Uromyces heteromorphae U-309 Peucedanum (Apiaceae) South Africa<br />
Uromyces trollipii U-445 Zygophyllaceae (Zygophyllum I, III) South Africa<br />
Endophyllum osteospermi from Alan Wood South Africa<br />
Endophyllum elytropappi U254 Elytropappus (Asteraceae III?) South Africa<br />
Puccinia magnusiana PUR-19 Ranunculaceae (Clematis O,I) Italy<br />
Puccinia festucae MCA2380 Lonicera (I Caprifoliaceae) Italy<br />
Puccinia hordei U-129 Taeniatherum (Poaceae III) Turkey<br />
Uromyces zygadeni MCA2577 Zigadenus (Liliaceae I-III) Idaho<br />
Puccinia phragmitis U-495 Poaceae (Phragmites) Austria<br />
Puccinia phragmitis U-423 Poaceae (Phragmites III) MN<br />
Puccinia recondita MCA2376 Aquilegia (Ranunculaceae) Italy<br />
?? U473 Narcissus (Liliaceae) Germany<br />
?? U473 Narcissus (Liliaceae) Germany<br />
Uredo dioscoreae-filiformis U458 Dioscorea (Discoreaceae) S. Africa<br />
Uromyces kentaniensis U-385 (s.n.) Iridaceae (Chasmanthe O,I,II,III) South Africa<br />
Puccinia recondita U-363 Poaceae (Agrostis II,III) VA<br />
Puccinia extensicola var. hydnoidea U-186 Dirca (Thymelaeaceae I) MN<br />
Cumminsiella mirabilissima AF426206 Mahonia (Berberidaceae)<br />
Cumminsiella U-480 Berberidaceae (Mahonia) Germany<br />
Puccinia recondita U-364 Poaceae (Agropyron II,III) VA<br />
Puccinia myrsiphylli U-377 (#571) Liliaceae (Asparagus) South Africa<br />
Puccinia hordei U-238 Lolium (Poaceae II-III) Turkey<br />
Uromyces geranii MCA2381 Geranium (O-I Geraniaceae) Italy<br />
Puccinia malvacearum? U474 Alcea (Malvaceae) Germany<br />
Puccinia sorghi? U-326 Oxalidaceae (Oxalis I) MD<br />
Puccinia malvacearum AF426205 Alcea (Malvaceae)<br />
Puccinia malvacearum JRH-225 Malvaceae (Malvastrum III) Honduras<br />
Puccinia isiacae U-340 Phragmites (Poaceae) Turkey<br />
Puccinia sp. U-139 Cardaria (Brassicaceae I) Turkey<br />
Puccinia magnusiana? U-223 Poaceae (Phragmites) Turkey<br />
Puccinia magnusiana? U-223 Poaceae (Phragmites) Turkey<br />
Uromycladium tepperianum U-466 Fabaceae (Acacia O,III) South Africa<br />
Uromycladium fusisporum BRIP-27608 Fabaceae (Acacia II-III) Australia<br />
Phakopsora arthuriana U-8 Euphorbiaceae (Jatropha II) P.R.<br />
Phakopsora JRH-488 Euphorbiaceae P.R.<br />
Phakopsora arthuriana GUY-2 Euphorbiaceae (Jatropha II) Guyana<br />
Phakopsora meibomiae JRH-456 Fabaceae (Lablab purpureus II) P.R.<br />
Phakopsora meibomiae JRH-471 Fabaceae (Lablab purpureus II) P.R.<br />
Phakopsora meibomiae JRH-475 Fabaceae (Lablab purpureus II) P.R.<br />
Phakopsora meibomiae U-102 Fabaceae (Phaseolus II) Honduras<br />
Phakopsora meibomiae JRH-284 Fabaceae (Desmodium II) Panama<br />
Phakopsora meibomiae JRH-314 Fabaceae (Phaseolus multiflorus II) Panama<br />
Phakopsora tecta JRH-457 Commelinaceae (Commelina II) P.R.<br />
Phakopsora tecta JRH-137 Commelinaceae (Commelina II) Costa Rica<br />
Phakopsora tecta JRH-476 Commelinaceae (Commelina II) Puerto Rico<br />
Phakopsora tecta U-536 (#610) Commelinaceae (Aneilema) South Africa<br />
Phakopsora pachyrhizi U-573 Fabaceae (Glycine max) LA<br />
Phakopsora pachyrhizi U-574 Fabaceae (Glycine max) LA<br />
Phakopsora pachyrhizi U-644 Fabaceae (Glycine max) Zimbabwe<br />
Phakopsora gossypii BRIP-39200 Malvaceae (Gossypium II) Australia<br />
Uredopeltis sp nov? U-454 (#582) Tiliaceae (Grewia) South Africa<br />
Phakopsora nishidana U-14 (JRH 1996-051) Moraceae (Ficus II) Argentina<br />
Batistopsora crucis-filii GUY-36 Annonaceae (Annona II-III) Guyana<br />
Phakopsora nishidana U-9 Moraceae (Ficus) P.R.<br />
[IDd as]Ravenelia platensis U-91 Fabaceae (Erythrina O) Argentina<br />
Prospodium sp nov? JRH-183 Bignoniaceae (Cydista II-II<br />
?? JRH160 Clytostoma (Bignoniaceae) Costa Rica<br />
Prospodium? JRH-160 Bignoniaceae (Clytostoma) Costa Rica<br />
Prospodium? JRH-178 Bignoniaceae Costa Rica<br />
?? JRH178 ?? (Bignoniaceae) Costa Rica<br />
?? JRH 181 Bignoniaceae Costa Rica<br />
Prospodium? JRH-181 Bignoniaceae Costa Rica<br />
Prospodium lippiae U-152 Verbenaceae (Aloysia II-III) Argentina<br />
Prospodium appendiculatum JRH-251 Bignoniaceae (Tecoma) Mexica<br />
Raveneliaceae U-410 Fabaceae (Derris II) Guam<br />
Raveneliaceae U-416 Fabaceae (Derris II) Guam<br />
Ravenellia havanensis U-19 Fabaceae (Enterolobium II-III) Argentina<br />
Ravenelia havanensis U-114 Fabaceae (Enterlobium II-III) Argentina<br />
Ravenelia echinata var. ectypa U-86 Fabaceae (Calliandra) Argentina<br />
Ravenelia sp. U-00118 Acacia (Fabaceae) C.R.<br />
Uredo sp. nov? (Ravenelia) U-121 Fabaceae (Acacia II) Costa Rica<br />
Uredo sp. nov? (Ravenelia) U-121 Acacia (Fabaceae II) C.R.<br />
Ravenelia acaciae-pennatulae U-115 Fabaceae (Acacia II) Mexico<br />
Uredo sp.nov? U-120 Diphysa (Fabaceae II) C.R.<br />
Uredo sp.nov? U-120 Fabaceae (Diphysa II) Costa Rica<br />
Ravenelia macrocarpa U-88 Fabaceae (Senna II) Argentina<br />
Puccinia spegazzinii U-317 (W1705) Asteraceae (M. micrantha III-microcyclic) Trinidad<br />
Ravenelia macowaniana U-376t Fabaceae (Acacia III) South Africa<br />
Ravenelai macowaniana U-449 (#588) Fabaceae (Acacia III) South Africa<br />
Ravenelia inornata U-376a (#570) Fabaceae (Acacia I) South Africa<br />
[from Phakopsora coca U-65 Erythroxylum (Erythroxylaceae II)] Argentina<br />
Gymnosporaingium MCA2568B Rosaceae Amelanchier I) VA<br />
Gymnoconia peckiana U-349 Rosaceae (Rubus allegheniensis O,I) MN<br />
Gymnoconia peckiana U-349 Rosaceae (Rubus allegheniensis O,I) MN<br />
Gymnosporangium japonicum U-24 Cupressaceae (Juniperus III) CA<br />
Gymnosporangium sp. MCA 2776 Rosaceae (Malus) VA<br />
Gymnosporangium juniperi-virginianae AY629316 AFTOL-712<br />
Gymnosporangium juniperi-virginianae AF522167<br />
Gymnosporangium exiguum? U-25 Rosaceae (Heteromales I) CA<br />
Gymnosporangium clavariiforme AF426211 Cupressaceae (Juniperus communis)<br />
Gymnosporangium sabinae AY512845<br />
Gymnosporangium sabinae AF426209 Rosaceae (Pyrus communis)<br />
Gymnosporangium cornutum AF426210 Rosaceae (Sorbus aucuparia)<br />
enodphytic rust? U-665 Poaceae (Bouteloua eriopoda, asymptomatic) NM<br />
Puccinia podophylli U-2 Berberidaceae (Podophyllum I-III) MD<br />
Puccinia podophylli U-13 Berberidaceae (Podophyllum O-III) MD<br />
Tranzschelia discolor JRH-139 Rosaceae (Prunus domestica II) Costa Rica<br />
Tranzschelia discolor U-507 (KR-0010962) Ranunculaceae (Anemone coronaria O,I) Greece<br />
Tranzschelia discolor U-509 (KR-0010965) Rosaceae (Prunus persica) Iran<br />
Tranzschelia discolor U-510 (KR-0010966) Rosaceae (Prunus domestica) Iran<br />
Tranzschelia discolor U-517 (KR-0010973) Rosaceae (Prunus persica) Iran<br />
Tranzschelia discolor AF426223 Rosaceae (Prunus domestica)<br />
Tranzschelia hyrcanica U-512 (KR-0010968) Rosaceae (Prunus cerasifera) Iran<br />
Tranzschelia hyrcanica U-513 (KR-0010969) Rosaceae (Prunus cerasifera) Iran<br />
Tranzschelia hyrcanica U-514 (KR-0010970) Rosaceae (Prunus cerasifera) Iran<br />
Tranzschelia hyrcanica U-515 (KR-0010971) Rosaceae (Prunus cerasifera) Iran<br />
Tranzschelia hyrcanica U-511 (KR-0010967) Rosaceae (Prunus spinosa) Iran<br />
Tranzschelia pruni-spinosae U-516 (KR-0010972) Rosaceae (Prunus cerasifera) Iran<br />
Tranzschelia pruni-spinosae AF426224 Ranunculaceae (Anemone ranunculoides)<br />
Tranzschelia fusca AF426225 Ranunculaceae (Anemone nemorosa)<br />
Phakopsora? JRH-463 Poaceae Puerto Rico<br />
Melampsora epitea complex U-562 Salicaceae (Salix sp.) WA<br />
melampsora epitea complex (SAL-53) Salix bebbiana ID<br />
Melampsora epitea? U-172 S. sp. Norway<br />
Melampsora laricis-epitea f.sp. laricis-daphnoides AY444789 (&AY444777ITS2) S. ac<br />
Melampsora epitea complex U-559 Salicaceae (Salix sp.) WA<br />
Melampsora (SAL-2) S. melanopsis ID<br />
Melampsora coleosporioides AY652951 (&AY652948ITS2) S. babylonica, Japan<br />
Melampsora amygdalinae AY444782 (AY444776ITS2) S. triandra N.Ireland<br />
Melampsora epitea U-284 [seq. inc.] S. humboldtiana Venezuela<br />
Melampsora ribesii-purpureae AY444791 (&AY444770ITS2) S. purpurea England<br />
Melampsora hypericorum U-394 Clusiaceae (Hypericum) WA<br />
Melampsora euphorbiae-gerardianae? U-485 Euphorbiaceae (Euphorbia) Germany<br />
Melampsora euphorbiae U-138 [LSU only] Euphorbiaceae (Euphorbia II) Syria<br />
Melampsora epitea complex U-557 [multiple PPs] Salix sp. WA<br />
Melampsora sp. (SAL-83) S. lemmonii ID<br />
Melampsora laricis AB116809 Populus davidiana, Tibet, China<br />
Melampsora occidentalis U-395 Populus balsamifera II WA<br />
Melampsora (SAL-49) [LSU only] Pinaceae (Abies grandis I) ID<br />
Pucciniastrum agrimoniae AF426234 Rosaceae (Agrimonia eupatoria)<br />
Cronartium quercuum U-62 Fagaceae (Quercus macrocarpa) IN<br />
Cronartium quercuum U-432 Fagaceae (Quercus muehlenbergii II) IN<br />
Cronartium quercuum U-434 Fagaceae (Quercus alba II) IN<br />
Cronartium quercuum U-435 Fagaceae (Quercus bicolor II) IN<br />
Cronartium quercuum u-436 Fagaceae (Quercus macrocarpa II) IN<br />
Cronartium ribicola U-396/U-268 Ribes VA/Pinus WY<br />
Endocronartium harknessii AF522175<br />
Cronartium flaccidum AF426239 Asclepiadaceae (Vincetoxicum hirundinaria)<br />
Endocronartium harknessii AY700193<br />
Pucciniastrum agrimoniae? U-304 Rosaceae (Leucoside) South Africa [inc. seq.]<br />
Chrysomyxa arctostaphyli AF522163<br />
Chrysomyxa arctostaphyli AY700192<br />
Chrysomyxa ledi AF426246 Pinaceae (Picea abies)<br />
Chrysomyxa rhododendri AF426245 Ericaceae (Rhododendron ferrugineum)<br />
Caeoma tsukubaensis AB19124 Ericaceae (Rhododendron kaempferi) Japan<br />
Thekopsora areolata AF426235 Rosaceae (Prunus padus)<br />
Hyalopsora U-361 undet. fern (Woodsiaceae?) UT<br />
Hyalopsora polypodii AF426229 Dryopteridaceae (Cystopteris fragilis)<br />
Hyalopsora polypodii AY512852<br />
Pucciniastrum? MCA2884 ?? Costa Rica<br />
Melampsoridium betulinum AF426228 Fagaceae (Betula pendula)<br />
Naohidemyces vaccinii MCA 2780 Ericaceae (Vaccinium) WA<br />
Naohidemyces vaccinii AF426238 Ericaceae (Vaccinium uliginosum)<br />
Uredinopsis sp. AF522181<br />
Uredinopsis filicina AF426237 Thelypteridaceae (Phegopteris connectilis)<br />
Milesina (as Milesia) scolopendri AF426236 Aspleniaceae (Asplenium scolopendrium)<br />
Pucciniastrum epilobii AF426226 Onagraceae (Epilobium angustifolium)<br />
Pucciniastrum epilobii AF522178<br />
Pucciniastrum circaeae AY745697<br />
Pucciniastrum epilobii AF522179<br />
Pucciniastrum circaeae AF426227 Onagraceae (Circaea lutetiana)<br />
Melampsorella caryophyllacearum AF426232 Pinaceae (Abies alba)<br />
Pucciniastrum goeppertianum AF522180<br />
Coleosporium ipomoeae JRH-195 Convolvulaceae (Ipomoea nil II) Costa Rica<br />
Coleosporium ipomoeae JRH-485 Convolvulaceae (Merremia quinquefolia II-III) Puerto Rico<br />
Coleosporium ipomoeae JRH-465 Convolvulaceae (Ipomoea triloba II) P.R.<br />
Coleosporium ipomoeae JRH-216 Convolvulaceae (Ipomoea cholulensis II-III) Honduras<br />
Coleosporium ipomoeae JRH-168 Convolvulaceae (Ipomoea trifida II-III) Costa Rica<br />
Coleosporium ipomoeae JRH-230 Convolvulaceae (Ipomoea anisomeres II) Honduras<br />
Coleosporium ipomoeae JRH-143 Convolvulaceae (Ipomoea neeiII) Costa Rica<br />
Coleosporium plumierae U-112 Apocynaceae (Plumeria rubra II-III) Nigeria<br />
Coleosporium plumeriae U-50 Apocynaceae (Plumeria rubra II) Guyana<br />
Coleosporium plumeriae U-266 Apocynaceae (Plumeria II) GUAM<br />
Coleosporium (asterum or delicatulum?) U-369 Asteraceae (Solidago) NY<br />
Coleosporium asterum U-67 Asteraceae (Symphyotrichum (=Aster) racemosum II-III) MD<br />
Coleosporium asterum AF522165<br />
Coleosporium asterum AF426241 Asteraceae (Aster ciliolatus)<br />
Coleosporium (asterum or delicatulum?) MCA-2600 Asteraceae (Solidago II) TN<br />
Coleosporium U-430 Asteraceae (Solidago?) NJ<br />
Coleosporium U-389 Asteraceae (Solidago?) MD<br />
Coleosporium U-388 Asteraceae (Solidago?) PA<br />
Coleosporium (asterum or delicatulum?) U-428 Asteraceae (Solidago) MD<br />
Coleosporium (asterum or delicatulum?) MCA-2777 S(Asteraceae (Solidago canadensis II-III) MD<br />
Coleosporium asterum AF522164<br />
Coleosporium sp. U-398 Asteraceae MD<br />
Coleosporium asterum U-366 Asteraceae (Aster novae-angliae II-III?) MD<br />
Coleosporium (asterum or delicatulum?) U-347 Asteraceae (Solidago) PA<br />
Coleosporium vernoniae U-371 Asteraceae (Vernonia) IL<br />
Coleosporium tussilaginis U-74 Asteraceae (Senecio triangularis) OR<br />
Coleosporium tussilaginis MCA-2389 Asteraceae (Sonchus?) CA<br />
Coleosporium tussilaginis MCA-2392 Asteraceae (Sonchus or Lactuca) CA<br />
Coleosporium vernoniae GUY-25 Asteraceae (Elephantopus mollis II)<br />
Coleosporium senecionis AY512840<br />
Coleosporium tussilaginis AF426242 Asteraceae (Tussilago farfara)<br />
Coleosporium cacaliae AF426243 Asteraceae (Adenostyles glabra)<br />
Coleosporium campanulae AF426244 Campanulaceae (Campanula scheuchzeri)<br />
Thekopsora symphyti AF426230 Boraginaceae (Symphytum officinale)<br />
Thekopsora guttata AF426231 Rubiaceae (Galium odoratum)<br />
Gymnosporangium libocedri AF522168<br />
Phakopsora apoda PDD-72076 Poaceae (Pennisetum II) New Zealand<br />
Phakopsora apoda PDD-68451 Poaceae (Pennisetum II) New Zealand<br />
Dasturella divina BRIP-39289 Poaceae (Babusa II) Australia<br />
Dasturella divina MCA-2887 Poaceae (bamboo) Costa Rica<br />
Phakopsora? MCA2617 Poaceae (Panicum) NC<br />
Phakopsora phakopsoroides JRH-184 Poaceae (Olyra II) Costa Rica<br />
Phakopsora euvitis BRIP-27996 Vitaceae (Vitis II) Australia<br />
Phakopsora vernoniae U-256 [#368] Asteraceae (Vernonia) South Africa<br />
Uredo alpestris AF426212 Violaceae (Viola alpestris)<br />
Kuehneola loeseneriana? MCA-2879 Rosaceae (Rubus trichomallus?) Costa Rica<br />
Gerwasia (as Mainsia) pittieriana U-78 Rosaceae (Rubus) Guatemala<br />
Kuehneola uredinis U-429 Rosaceae (Rubus) MD<br />
Kuehneola uredinis U-303 Rosaceae (Rubus pinnatus) South Africa<br />
Kuehneola uredinis U-348 Rosaceae (Rubus) PA<br />
Kuehneola uredinis AY745696 AFTOL-987<br />
Kuehneola uredinis AF426218 Rosaceae (Rubus fruticosus)<br />
Kuehneola uredinis MCA-2393 Rosaceae (Rubus III) MD<br />
Kuehneola uredinis MCA-2867 Rosaceae (Rubus) MD<br />
Kuehneola uredinis MCA-2863 Rosaceae (Rubus) VA<br />
Frommeella mexicana var. indicae U-3 Rosaceae (Duchesnea II) MD<br />
FrommeÎlla mexicana MCA-2496 Rosaceae (Duchesnea) MD<br />
Frommeella mexicana MCA-2495 Rosaceae (Duchesnea II?) VA<br />
Phragmidium mucronatum U-106 Rosaceae (Rosa II) Argentina<br />
Phragmidium mucronatum? U-353 Rosaceae (Rosa) OR<br />
Phragmidium sanguisorbae AF426216 Rosaceae (Sanguisorba minor)<br />
Phragmidium violaceum AF426214 Rosaceae (Rubus fruticosus)<br />
Phragmidium violaceaum MCA-2782 Rosaceae (Rubus fruticosus? II-III) France<br />
Phragmidium violaceum U-645 Rosaceae (Rubus armeniacus II) OR<br />
Phragmidium rubi-idaei AF426215 Rosaceae (Rubus idaeus)<br />
Phragmidium fragariae AF426217 Rosaceae (Potentilla sterilis)<br />
Phragmidium montivagum AF426213 Rosaceae (Rosa cf. woodsii)<br />
Triphragmium ulmariae AF426219 Rosaceae (Filipendula ulmaria)<br />
Ochropsora ariae AF426221 Ranunculaceae (Anemone nemorosa)<br />
Ochropsora ariae AF426222 Rosaceae (Aruncus dioicus)<br />
Trachyspora intrusa AF426220 Rosaceae (Alchemilla vulgaris)<br />
Trachyspora intrusa MCA2384 Alchemilla (Rosaceae) Switzerland<br />
Caeoma torreyae AF522183<br />
Platygloea vestita AY512872<br />
Rhodotorula fushanensis AB176591<br />
16 GROUPS<br />
Dataset: Aime. 2006. Toward<br />
resolving family-level relationships in<br />
rust fungi (Uredinales). Mycoscience<br />
47(3):112-122. and unpublished.
Pucciniales<br />
rDNA LSU<br />
1229 bp.; 857 incl. char.; 263 PI char<br />
470 seq; 350 taxa (+4 outgroup)<br />
13 of 13 families (100%)<br />
58 of 117 genera (49.5%)<br />
~500 of 7000 species (7%)<br />
(classifications fide<br />
Cummins & Hiratsuka 2003)<br />
1 change<br />
Helicobasidium purpureum AY885168<br />
Eocronartium muscicola AY512844<br />
Puccinia mariae-wilsoniae U-005 Claytonia (Portulaceae I-III) MD<br />
Puccinia mariae-wilsoniae MCA2497 Claytonia (Portulaceae) MD<br />
Puccinia lagenophorae U-384 (#569) Asteraceae (Osteocarpum) South Africa<br />
Puccinia lagenophora U-128 Senecio (I-III Asteraceae) Tunisia<br />
Puccinia lagenophorae U-107 Senecio (I, III Asteraceae) Argentina<br />
Puccinia violae U-023 Viola (Violaceae II-III) Norway<br />
Puccinia violae U-269 Viola (Violaceae II-III) NY<br />
Uromyces sp. MCA 2870 Juncaceae (Juncus? III) VA<br />
Puccinia? U397 Asteraceae MD<br />
Puccinia? U-390 Asteraceae MD<br />
Puccinia cf. violae U-4 Violaceae (Viola cucullata )-I, autoecious) MD<br />
Uromyces vignae? U-015 Macroptilium (Fabaceae II) Argentina<br />
Uromyces vignae JRH-472 Fabaceae (Vigna II-III) Puerto Rico<br />
Uromyces vignae JRH-468 Fabaceae (Vigna II-III) Puerto Rico<br />
Uromyces vignae U-125 Vigna (Fabaceae II) C.R.<br />
Uromyces vignae U-126 Vigna (Fabaceae II) C.R.<br />
Uromyces appendiculatus U-122 Phaseolus (II Fabaceae) C.R.<br />
Uromyces appendiculatus U-123 Phaseolus (II Fabaceae) Mexico<br />
Uromyces appendiculatus U-00123 Phaseolus (Fabaceae) Mexico<br />
Uromyces neotropicalis sp.nov MCA2533 Cucurbitaceae II,III Ecuador<br />
Dietelia mesoamericana sp.nov. U-320 (W1909) Asteraceae (M. micrantha microcyclic I) Mexico<br />
Dietelia portoricensis u-322 (no #) Asteraceae (Mikania micrantha I-microcyclic) Costa Rica<br />
Puccinia granularis U-443 Geraniaceae (Pelargonium O,I,II,III) South Africa<br />
Puccinia granularis U-444 (#581) Geraniaceae (Pelargonium II,III) South Africa<br />
Puccinia granularis U-444 (#581) Geraniaceae (Pelargonium II,III) South Africa<br />
Puccinia granularis U-443 (#580) Geraniaceae (Pelargonium O,I,II,III) South Africa<br />
Puccinia spegazzinii JRH-226 Asteraceae (Mikania micrantha III) Honduras<br />
Puccinia spegazzinii U-319 (W2102) Asteraceae (Mikania micrantha microcyclic III) Peru<br />
Puccinia spegazzinii U-315 (W2020) Asteraceae (Mikania micrantha IIImicrocyclic) Argentina<br />
Dietelia mesoamericana sp. nov. U-321 (mislabelled as U143) Mikania Mexico<br />
Puccinia hydrocotyles U-079 Hydrocotyle (II-III Apiaceae) AL<br />
Puccinia? MCA-2864 Asteraceae (Helianthus) VA<br />
Puccinia pimpinellae? MCA2576 Apiaceae( Osmorhiza I) ID<br />
Puccinia pimpinellae U-43 Osmorhiza (Apiaceae III) UT<br />
Puccinia pimpinellae MCA2604 Osmorhiza (Apiaceae III) TN<br />
Puccinia berkheyicola U-314 Berkheya (Asteraceae) South Africa<br />
Puccinia notobasidis U-229 Cirsium (Asteraceae) Turkey<br />
Puccinia variabilis var. lapsanae U-37 Asteraceae (Lapsana II,III) UT<br />
?? U-412 Asteraceae (Youngia) Guam<br />
?? JRH199 Wedelia (Asteraceae) Hondorus (inc seq)<br />
Puccinia menthae MCA 2601 Monarda (Lamiaceae II) TN<br />
Puccinia menthae U-343 Origanum (Lamiaceae II) Argentina<br />
Puccinia menthae MCA 2601 Monarda (Lamiaceae II) TN<br />
Puccinia marylandica MCA2602 Sanicula (Apiaceae II-III) TN<br />
Puccinia marylandica MCA2606 Sanicula (Apiaceae II-III?) TN<br />
Puccinia commelinae U-049 Commelina (Commelinaceae II) Guyana<br />
Puccinia synedrellae JRH152 Synedrella (III Asteraceae) Costa Rica<br />
Puccinia cnici-oleracei GUY-6 Asteraceae (Emilia sonchifolia III) Guyana<br />
Puccinia synedrellae JRH-152 Asteraceae (Synedrella nodiflora III) Costa Rica<br />
Puccinia cnici-oleracei JRH-204 Asteraceae (Emilia fosbergii III) Honduras<br />
Puccinia cnici-oleracei JRH-490 Asteraceae (Emilia fosbergii III) Puerto Rico<br />
Puccinia MCA-2864 Asteraceae (Helianthus) VA<br />
Uromyces ficariae AF426204 Ranunculus (Ranunculaceae)<br />
Puccinia sp. MCA2575 Asteraceae Idaho<br />
Puccinia sp. MCA-2575 Asteraceae (II) ID<br />
Puccinia chondrillina U-202 Asteraceae (Chondrilla) Turkey [BPI extraction]<br />
Puccinia balsamorrhizae U-332Asteraceae (Balsamorhiza III?) B.C.<br />
Puccinia balsamorrhizae U-354 Asteraceae (Balsamorhiza) ID<br />
Puccinia balsamorrhizae U-355 Asteraceae (Balsamorhiza) ID<br />
Miyagia pseudosphaeria U-63 (BPI) Asteraceae (Sonchus oleraceus)<br />
Puccinia calcitrapae v. centaureae U-201 Asteraceae (Centaurea depressa) Turkey [BPI extraction]<br />
Puccinia calcitrapae U-218 Carduus (Asteraceae) Turkey<br />
Puccinia calcitrapae U-218 Asteraceae (Carduus nutans) Turkey<br />
Puccinia carthami complex MCA-2881 Asteraceae (Cirsium?) Costa Rica<br />
Puccinia acroptili U-224(confirmed no.) Asteraceae (Acroptilon) Turkey<br />
Puccinia acroptili U-578 Asteraceae (Acroptilon repens III) NM<br />
Puccinia acroptili U-135 (Dana Berner 02-048) Asteraceae (Acroptilon repens II-III?) Turkey<br />
Puccinia acroptili U-224 Asteraceae (Acroptilon repens) Turkey<br />
Puccinia acroptili U-577 Asteraceae (Centaurea repens) NM<br />
Puccinia U-203 Asteraceae (Cardueae II-III) Turkey [host & rust confirmed from BPI]<br />
Puccinia punctiiformis U-225 Cirsium (Asteraceae) Turkey<br />
Puccinia punctiiformis U-225 Asteraceae (Cirsium arvense) Turkey<br />
Puccinia calcitrapae U-164 Asteraceae (Carthamus tinctorius II-III) Argentina<br />
Puccinia centaurea U-278 (FDWSRU 04-010) Asteraceae (Centaurea hybrid) OR<br />
Puccinia calcitrapae v. centaureae U-140 (Dana Berner 03-051) Asteraceae (Centaurea macedonica) Greece<br />
Puccinia calcitrapae v. centaurea U-209 Asteraceae (Centaurea calcitrapa) Turkey<br />
Puccinia carthami U-323 Asteraceae (Carthamus tinctorius III) Oman<br />
Puccinia atropuncta MCA2609 Veratrum (Liliaceae III) TN<br />
?? U492 Antirrhinum (Scrophulariaceae) Germany<br />
?? MCA2529 Geranium (Geraniaceae II) Ecuador<br />
Endophyllum stachytarphetae GUY-55 Stachytarpheta (III Verbenaceae)<br />
?? MCA2558 Phragmites? (Poaceae) Ecuador<br />
Endophyllum cassiae U-525 Fabaceae (Cassia III?) Nigeria<br />
?? JRH-466 Cyperaceae Puerto Rico<br />
?? U-265 Cyperus (Cyperaceae II?) Guam<br />
Puccinia cnici-oleracei JRH-200 Asteraceae (Melampodium divaricatum III) Honduras<br />
Puccinia cnici-oleracei JRH-207 Asteraceae (Malampodium divaricatum III) Honduras<br />
Puccinia ursiniae U-463 (#590) Asteraceae (Ursinia) South Africa<br />
Puccinia tanaceti U-040 Artemisia (Asteraceae II-III) UT<br />
Puccinia rubefaciens? U-330 Galium (Rubiaceae III?) B.C. Canada<br />
Puccinia medellinensis JRH-154 Lamiaceae (Hyptis II) Costa Rica<br />
Puccinia melampodii JRH138 Emilia (Asteraceae III) Costa Rica<br />
Uromyces rumicis U-227 Rumex (Polygonaceae) Turkey<br />
Uromyces lespedezae-procumbentis U-007 Trifolium (Fabaceae III) MD<br />
Puccinia convolvuli? MCA2778 Calystegia (Convolvulaceae II) MD<br />
Puccinia convolvuli U-068 Calystegia (II-III Convolvulaceae) MD<br />
Puccinia scleriicola GUY-57 Scleria (I Cyperaceae)<br />
Puccinia ibrae JRH-228 Convolvulaceae (Ipomoea I-III) Honduras<br />
Puccinia thaliae JRH141 Canna (Cannaceae II) Costa Rica<br />
Puccinia thaliae GUY-3 Canna (Cannaceae II) Guyana<br />
Puccinia kalchbrenneriana U-312 (#560) Asteraceae (Helichrysum III?) South Africa<br />
Puccinia kalchbrenneriana U-313 (#555) Asteraceae (Helichrysum III?) South Africa<br />
Puccinia kalchbrenneri? U-258 (#539) Asteraceae (Helichrysum III) South Africa<br />
Uromyces colombianus JRH-145 Asteraceae (Melanthera II-III) Costa Rica<br />
Uromyces psoraleae U-029 Psoralidium (Fabaceae I-III) UT<br />
Uromyces sp. JRH-482 Cucurbitaceae (Cayaponia) Puerto Rico<br />
Puccinia gigantea AF426198 Epilobium (Onagraceae)<br />
Puccinia enceliae JRH-166 Asteraceae (Tithonia II-III) Costa Rica<br />
Puccinia thunbergiae U-260 (543) Acanthaceae (Thunbergia O,I,III) South Africa<br />
Puccinia hydrocotyles U-105 Hydrocotyle (Apiaceae) Argentina<br />
Uromyces galegae U-242 Galega (Fabaceae) Turkey<br />
Uredo cajani U-119 Cajanus (Fabaceae II) Honduras<br />
Uredo cajani MCA-2895 Fabaceae (Cajanus) Costa Rica<br />
Uredo cajani JRH-489 Fabaceae (Cajanus II) Puerto Rico<br />
Uromyces dolicholi JRH-469 Fabaceae (Rhynchosia II) Puerto Rico<br />
Uromyces scutellatus U-204(presumed) Euphorbiaceae (Euphorbia) Turkey<br />
Uromyces euphorbiae GUY-7 Euphorbia (II Euphorbiaceae)<br />
Uromyces pisi AF426201 Euphorbia<br />
Uromyces pisi-sativi? MCA2377 Euphorbia (Euphorbiaceae) Italy<br />
Uromyces? U-471 Euphorbiaceae (Euphorbia) Germany<br />
Endophyllum euphorbiae-sylvaticae AF426200 Euphorbia<br />
Puccinia abrupta JRH-146 Viguiera (Asteraceae II-III) Costa Rica<br />
Uromyces viciae-fabae AF426199 Vicia (Fabaceae)<br />
Puccinia sp.nov.? U-210 Tragopagon (Asteraceae I-III) Turkey<br />
Puccinia stoboeae U-259 (#550) Asteraceae (Berkheya O,I,II) South Africa<br />
Uromyces hedysari-paniculati U-124 Desmodium (Fabaceae II-III ) Honduras<br />
Uromyces hedysari-paniculati U-052 Desmodium (Fabaceae II) Guyana<br />
Aecidium sp. U-211 Salsola (Chenopodiaceae I) Turkey<br />
Uromyces ciceris-arietini U-183 Cicer (Fabaceae II-III ) SD<br />
Uromyces junci AF426203 Pulicaria (Asteraceae)<br />
Puccinia? MCA2593 (Asteraceae O?) ID<br />
Puccinia diplachnis MCA2375 Cirsium (I Asteraceae) Switzerland {T=Poaceae}<br />
Puccinia verruca U-130 Centaurea (Asteraceae III) Tunisia<br />
Puccinia collinsiae U-044 Collinsiae (Scrophulariaceae) UT<br />
Uromyces trifolii-repentis U-207 Trifolium (Fabaceae) Turkey<br />
Puccinia urticae-acutiformis AF426202 Carex (Cyperaceae)<br />
Puccinia hemerocallidis U498 Hemerocallis (Lilioaceae) TX<br />
Puccinia hemerocallidis U-072 Hemerocallis (II Liliaceae) AL<br />
Puccinia hemerocallidis U-073 Hemerocallis (II Liliaceae) AL<br />
Uromyces euryopsidicola U-381 (#575) Asteraceae (Euryops III) South Africa<br />
Uromyces euryopsidicola U-216 (#295) Asteraceae (Euryops) South Africa<br />
Uromyces euryopsidicola U-261 (#295) Asteraceae (Euryops) South Africa<br />
Uromyces euryopsidicola U-216 (#295) Asteraceae (Euryops) South Africa<br />
Puccinia sparganioides U-368 Poaceae (Spartina II,III) NY<br />
Puccinia buxi U-418 Buxus (Buxaceae) Georgia<br />
Puccinia tillandsiae U419 Tillandsia (Bromeliaceae) Guatemala<br />
Puccinia arechaveletae MCA2430 Serjania (Sapindaceae) Guyana<br />
Puccinia arechavelaete GUY-66 Serjania (III Sapindaceae)<br />
Puccinia arechavaletae JRH175 Unvillea (II-III Sapindaceae) Costa Rica<br />
Puccinia physalidis U-189 Physalis (Solanaceae III) ND<br />
Uromyces ixiae U452 Romulea (Iridaceae) S. Africa<br />
Uromyces transversalis U-386 Iridaceae (Gladiolus) Mexico<br />
Puccinia hordei MCA2391 (Poaceae) CA<br />
Puccinia? U-288 Poaceae (Pennisetum II) Guam<br />
Uromyces setaria-italicae JRH220 Melinis (II Poaceae) Hondorus<br />
Uromyces setaria-italicae JRH205 Panicum (II Poaceae) Hondorus<br />
Uromyces setaria-italicae JRH202 Panicum (II Poaceae) Honduras<br />
Puccinia gouaniae JRH159 Gouania (II Rhamnaceae) Costa Rica<br />
Puccinia arundinariae? MCA2494 Arundinaria (Poaceae) VA<br />
Puccinia purpurea JRH198 Sorghum (Poaceae II-III) Honduras<br />
Puccinia purpurea JRH213 Sorghum (Poaceae II) Hondurus<br />
Puccinia purpurea JRH218 Sorghum (Poaceae II) Hondurus<br />
Puccinia duthiae GUY8 Bothriochloa (Poaceae II) Guyana<br />
Puccinia duthiae U-264 Dichanthium (Poaceae II) Guam<br />
Puccinia exhauriens U-538 (#600) Oleaceae (Jasminum) South Africa<br />
Puccinia exhauriens U-539 (#605) Oleaceae (Jasminum) South Africa<br />
Puccinia moraeae U462 Moraea (Iridaceae) S. Africa<br />
Uromyces massoniae? U-446(#586) Liliaceae (Massonia II-III) South Africa<br />
Uromyces salsolae U-134 Salsola (Chenopodiaceae) Syria<br />
Puccinia sp. U-006 Allium? (Liliaceae III) MD<br />
Uromyces aloes U-00372 (#567) Aloaceae (Aloe) South Africa<br />
Puccinia obliquo-septata MCA2431 Olyra (Poaceae) Guyana<br />
Puccinia obliquo-septata GUY-10 Olyra (II Poaceae)<br />
Puccinia peperomiae JRH192 Piper (II Piperaceae) Costa Rica<br />
Puccinia fallax MCA2305 Palicourea (Rubiaceae II-III) Guyana<br />
Puccinia fallax GUY-58 Palicourea (II Rubiaceae)<br />
Sphenospora kevorkianii U-10 Orchidaceae (Stanhopea II) Peru<br />
Puccinia coronata U-583 Poaceae (Avena) ND<br />
Puccinia coronata U400 Festuca (Poaceae II, III) VA<br />
Puccinia coronata U-244 Rhamnus (Rhamnaceae O-O) ND<br />
Puccinia coronata U-245 Scolochloa (Poaceae III?) ND<br />
Puccinia coronata AF426207 Rhamnus (Rhamnaceae)<br />
Puccinia pampeanum U-109 Salpichroa (III Solanales) Argentina<br />
Endophyllum pampeanum U-108 Salpichroa (III Solanales) Argentina<br />
Uromyces gageae AF426208 Gagea (Liliaceae)<br />
Aecidium wiborgiae U-380 (#576) Fabaceae (Wiborgia) South Africa<br />
Aecidium wiborgiae U-379 (#573) Fabaceae (Wiborgia) South Africa<br />
Uromyces sp. MCA2383 Lilium (III Liliaceae) Switzerland<br />
Puccinia mogiphanis U-111 Alternanthera (Amaranthaceae) Nigeria<br />
Puccinia stonemanniae U-310 Thesium (Santalaceae III?) South Africa<br />
Puccinia sorghi U-362 Poaceae (Zea mays II, III) VA<br />
Uromyces polygoni U-214 Polygonum (Polygonaceae) Turkey<br />
Uromyces polygoni U-214 Polygonum (Polygonaceae) Turkey<br />
Puccinia mogiphanis JRH 180 Amarantaceae (Alternanthera II) Costa Rica<br />
Puccinia cynodontis JRH 219 Cynodon (Poaceae II-III) Hondorus<br />
Aecidium kalanchoe U-18 Kalanchoe<br />
Puccinia horiana U-391 Asteraceae (Chrysanthemum III) DE<br />
Puccinia horiana U-392 Asteraceae (Chrysanthemum III) MD<br />
Puccinia horiana U-070 Chrysanthemum (Asteraceae) NJ<br />
Aecidium sp. U-465 (#579) Fabaceae (Lebeckia) South Africa<br />
Puccinia arenariae U142 Dianthus (Caryophyllaceae) Ecuador<br />
Puccinia graminis U-478 Berberis (Berberidaceae) Germany<br />
Puccinia.poarum FROM BRANDON on Tussilago (iii on Poaceae)<br />
Puccinia MCA-2882 Asteraceae? Costa Rica<br />
Puccinia MCA-2886 Asteraceae? Costa Rica<br />
Puccinia graminis (as Aecidium sp.) PDD78317 Berberis I N.Z.<br />
Puccinia graminis U-469 Agropyron (Poaceae) Germany<br />
Puccinia graminis U-470 Agropyron? (Poaceae) Germany<br />
Puccinia graminis U-470 Poaceae (Agropyron?) Germany<br />
Uredo baruensis GUY-71 Sapotaceae (Chrysophyllum II) Guyana<br />
[IDd as Coleosporium on Merremia (Convolvulaceae)] JRH-484 P.R. (seq. confirmed 2X)<br />
Sphaerophragmium acacia JRH-486 Fabaceae (Albizia II-III) Puerto Rico<br />
Sphaerophragmium acaciae JRH-470 Fabaceae (Albizia II) Puerto Rico<br />
Pileolaria toxicodendri AY 745698 (AFTOL ID 988)<br />
Pileolaria toxicodendri u-607 Anacardiaceae (Toxicodendron) MN<br />
Puccinia tetragonia U-442 Aizoaceae (Tetragonia) South Africa<br />
?? U-476 Allium (Liliaceae) Germany<br />
Uromyces albucae U-447 (#591) Liliaceae (Albuca) South Africa<br />
Puccinia dispersa U-234 Anchusa (Boraginaceae) Turkey<br />
Puccinia? MCA-2885 ?? Costa Rica<br />
Chrysocyclus cestri U-156 Cestrum (Solanaceae III) Argentina<br />
Puccinia smilacis U-12 Smilacaceae (Smilax II-III) MD<br />
Puccinia smilacis U-393 Smilacaceae (Smilax) MD<br />
Maravalia sapotae JRH-172<br />
Stereostratum corticiodes U-027 Bambusa (Poaceae II-III) China<br />
Puccinia asarina MCA2374 Asarum (III Aristolochiaceae) Italy<br />
Puccinia pampeana JRH135 Capsicum (Solanaceae II-III aecioid) Costa Rica<br />
Endophyllum macowanii U-255 Rhamnus (Rhamnaceae III?) South Africa<br />
Puccinia thlaspeos U-054 Boechera (III Brassicaceae) WA<br />
Uromyces bulbinis BRIP-28988 Liliaceae (Bulbine) Australia<br />
Uromyces heteromorphae U-309 Peucedanum (Apiaceae) South Africa<br />
Uromyces trollipii U-445 Zygophyllaceae (Zygophyllum I, III) South Africa<br />
Endophyllum osteospermi from Alan Wood South Africa<br />
Endophyllum elytropappi U254 Elytropappus (Asteraceae III?) South Africa<br />
Puccinia magnusiana PUR-19 Ranunculaceae (Clematis O,I) Italy<br />
Puccinia festucae MCA2380 Lonicera (I Caprifoliaceae) Italy<br />
Puccinia hordei U-129 Taeniatherum (Poaceae III) Turkey<br />
Uromyces zygadeni MCA2577 Zigadenus (Liliaceae I-III) Idaho<br />
Puccinia phragmitis U-495 Poaceae (Phragmites) Austria<br />
Puccinia phragmitis U-423 Poaceae (Phragmites III) MN<br />
Puccinia recondita MCA2376 Aquilegia (Ranunculaceae) Italy<br />
?? U473 Narcissus (Liliaceae) Germany<br />
?? U473 Narcissus (Liliaceae) Germany<br />
Uredo dioscoreae-filiformis U458 Dioscorea (Discoreaceae) S. Africa<br />
Uromyces kentaniensis U-385 (s.n.) Iridaceae (Chasmanthe O,I,II,III) South Africa<br />
Puccinia recondita U-363 Poaceae (Agrostis II,III) VA<br />
Puccinia extensicola var. hydnoidea U-186 Dirca (Thymelaeaceae I) MN<br />
Cumminsiella mirabilissima AF426206 Mahonia (Berberidaceae)<br />
Cumminsiella U-480 Berberidaceae (Mahonia) Germany<br />
Puccinia recondita U-364 Poaceae (Agropyron II,III) VA<br />
Puccinia myrsiphylli U-377 (#571) Liliaceae (Asparagus) South Africa<br />
Puccinia hordei U-238 Lolium (Poaceae II-III) Turkey<br />
Uromyces geranii MCA2381 Geranium (O-I Geraniaceae) Italy<br />
Puccinia malvacearum? U474 Alcea (Malvaceae) Germany<br />
Puccinia sorghi? U-326 Oxalidaceae (Oxalis I) MD<br />
Puccinia malvacearum AF426205 Alcea (Malvaceae)<br />
Puccinia malvacearum JRH-225 Malvaceae (Malvastrum III) Honduras<br />
Puccinia isiacae U-340 Phragmites (Poaceae) Turkey<br />
Puccinia sp. U-139 Cardaria (Brassicaceae I) Turkey<br />
Puccinia magnusiana? U-223 Poaceae (Phragmites) Turkey<br />
Puccinia magnusiana? U-223 Poaceae (Phragmites) Turkey<br />
Uromycladium tepperianum U-466 Fabaceae (Acacia O,III) South Africa<br />
Uromycladium fusisporum BRIP-27608 Fabaceae (Acacia II-III) Australia<br />
Phakopsora arthuriana U-8 Euphorbiaceae (Jatropha II) P.R.<br />
Phakopsora JRH-488 Euphorbiaceae P.R.<br />
Phakopsora arthuriana GUY-2 Euphorbiaceae (Jatropha II) Guyana<br />
Phakopsora meibomiae JRH-456 Fabaceae (Lablab purpureus II) P.R.<br />
Phakopsora meibomiae JRH-471 Fabaceae (Lablab purpureus II) P.R.<br />
Phakopsora meibomiae JRH-475 Fabaceae (Lablab purpureus II) P.R.<br />
Phakopsora meibomiae U-102 Fabaceae (Phaseolus II) Honduras<br />
Phakopsora meibomiae JRH-284 Fabaceae (Desmodium II) Panama<br />
Phakopsora meibomiae JRH-314 Fabaceae (Phaseolus multiflorus II) Panama<br />
Phakopsora tecta JRH-457 Commelinaceae (Commelina II) P.R.<br />
Phakopsora tecta JRH-137 Commelinaceae (Commelina II) Costa Rica<br />
Phakopsora tecta JRH-476 Commelinaceae (Commelina II) Puerto Rico<br />
Phakopsora tecta U-536 (#610) Commelinaceae (Aneilema) South Africa<br />
Phakopsora pachyrhizi U-573 Fabaceae (Glycine max) LA<br />
Phakopsora pachyrhizi U-574 Fabaceae (Glycine max) LA<br />
Phakopsora pachyrhizi U-644 Fabaceae (Glycine max) Zimbabwe<br />
Phakopsora gossypii BRIP-39200 Malvaceae (Gossypium II) Australia<br />
Uredopeltis sp nov? U-454 (#582) Tiliaceae (Grewia) South Africa<br />
Phakopsora nishidana U-14 (JRH 1996-051) Moraceae (Ficus II) Argentina<br />
Batistopsora crucis-filii GUY-36 Annonaceae (Annona II-III) Guyana<br />
Phakopsora nishidana U-9 Moraceae (Ficus) P.R.<br />
[IDd as]Ravenelia platensis U-91 Fabaceae (Erythrina O) Argentina<br />
Prospodium sp nov? JRH-183 Bignoniaceae (Cydista II-II<br />
?? JRH160 Clytostoma (Bignoniaceae) Costa Rica<br />
Prospodium? JRH-160 Bignoniaceae (Clytostoma) Costa Rica<br />
Prospodium? JRH-178 Bignoniaceae Costa Rica<br />
?? JRH178 ?? (Bignoniaceae) Costa Rica<br />
?? JRH 181 Bignoniaceae Costa Rica<br />
Prospodium? JRH-181 Bignoniaceae Costa Rica<br />
Prospodium lippiae U-152 Verbenaceae (Aloysia II-III) Argentina<br />
Prospodium appendiculatum JRH-251 Bignoniaceae (Tecoma) Mexica<br />
Raveneliaceae U-410 Fabaceae (Derris II) Guam<br />
Raveneliaceae U-416 Fabaceae (Derris II) Guam<br />
Ravenellia havanensis U-19 Fabaceae (Enterolobium II-III) Argentina<br />
Ravenelia havanensis U-114 Fabaceae (Enterlobium II-III) Argentina<br />
Ravenelia echinata var. ectypa U-86 Fabaceae (Calliandra) Argentina<br />
Ravenelia sp. U-00118 Acacia (Fabaceae) C.R.<br />
Uredo sp. nov? (Ravenelia) U-121 Fabaceae (Acacia II) Costa Rica<br />
Uredo sp. nov? (Ravenelia) U-121 Acacia (Fabaceae II) C.R.<br />
Ravenelia acaciae-pennatulae U-115 Fabaceae (Acacia II) Mexico<br />
Uredo sp.nov? U-120 Diphysa (Fabaceae II) C.R.<br />
Uredo sp.nov? U-120 Fabaceae (Diphysa II) Costa Rica<br />
Ravenelia macrocarpa U-88 Fabaceae (Senna II) Argentina<br />
Puccinia spegazzinii U-317 (W1705) Asteraceae (M. micrantha III-microcyclic) Trinidad<br />
Ravenelia macowaniana U-376t Fabaceae (Acacia III) South Africa<br />
Ravenelai macowaniana U-449 (#588) Fabaceae (Acacia III) South Africa<br />
Ravenelia inornata U-376a (#570) Fabaceae (Acacia I) South Africa<br />
[from Phakopsora coca U-65 Erythroxylum (Erythroxylaceae II)] Argentina<br />
Gymnosporaingium MCA2568B Rosaceae Amelanchier I) VA<br />
Gymnoconia peckiana U-349 Rosaceae (Rubus allegheniensis O,I) MN<br />
Gymnoconia peckiana U-349 Rosaceae (Rubus allegheniensis O,I) MN<br />
Gymnosporangium japonicum U-24 Cupressaceae (Juniperus III) CA<br />
Gymnosporangium sp. MCA 2776 Rosaceae (Malus) VA<br />
Gymnosporangium juniperi-virginianae AY629316 AFTOL-712<br />
Gymnosporangium juniperi-virginianae AF522167<br />
Gymnosporangium exiguum? U-25 Rosaceae (Heteromales I) CA<br />
Gymnosporangium clavariiforme AF426211 Cupressaceae (Juniperus communis)<br />
Gymnosporangium sabinae AY512845<br />
Gymnosporangium sabinae AF426209 Rosaceae (Pyrus communis)<br />
Gymnosporangium cornutum AF426210 Rosaceae (Sorbus aucuparia)<br />
enodphytic rust? U-665 Poaceae (Bouteloua eriopoda, asymptomatic) NM<br />
Puccinia podophylli U-2 Berberidaceae (Podophyllum I-III) MD<br />
Puccinia podophylli U-13 Berberidaceae (Podophyllum O-III) MD<br />
Tranzschelia discolor JRH-139 Rosaceae (Prunus domestica II) Costa Rica<br />
Tranzschelia discolor U-507 (KR-0010962) Ranunculaceae (Anemone coronaria O,I) Greece<br />
Tranzschelia discolor U-509 (KR-0010965) Rosaceae (Prunus persica) Iran<br />
Tranzschelia discolor U-510 (KR-0010966) Rosaceae (Prunus domestica) Iran<br />
Tranzschelia discolor U-517 (KR-0010973) Rosaceae (Prunus persica) Iran<br />
Tranzschelia discolor AF426223 Rosaceae (Prunus domestica)<br />
Tranzschelia hyrcanica U-512 (KR-0010968) Rosaceae (Prunus cerasifera) Iran<br />
Tranzschelia hyrcanica U-513 (KR-0010969) Rosaceae (Prunus cerasifera) Iran<br />
Tranzschelia hyrcanica U-514 (KR-0010970) Rosaceae (Prunus cerasifera) Iran<br />
Tranzschelia hyrcanica U-515 (KR-0010971) Rosaceae (Prunus cerasifera) Iran<br />
Tranzschelia hyrcanica U-511 (KR-0010967) Rosaceae (Prunus spinosa) Iran<br />
Tranzschelia pruni-spinosae U-516 (KR-0010972) Rosaceae (Prunus cerasifera) Iran<br />
Tranzschelia pruni-spinosae AF426224 Ranunculaceae (Anemone ranunculoides)<br />
Tranzschelia fusca AF426225 Ranunculaceae (Anemone nemorosa)<br />
Phakopsora? JRH-463 Poaceae Puerto Rico<br />
Melampsora epitea complex U-562 Salicaceae (Salix sp.) WA<br />
melampsora epitea complex (SAL-53) Salix bebbiana ID<br />
Melampsora epitea? U-172 S. sp. Norway<br />
Melampsora laricis-epitea f.sp. laricis-daphnoides AY444789 (&AY444777ITS2) S. ac<br />
Melampsora epitea complex U-559 Salicaceae (Salix sp.) WA<br />
Melampsora (SAL-2) S. melanopsis ID<br />
Melampsora coleosporioides AY652951 (&AY652948ITS2) S. babylonica, Japan<br />
Melampsora amygdalinae AY444782 (AY444776ITS2) S. triandra N.Ireland<br />
Melampsora epitea U-284 [seq. inc.] S. humboldtiana Venezuela<br />
Melampsora ribesii-purpureae AY444791 (&AY444770ITS2) S. purpurea England<br />
Melampsora hypericorum U-394 Clusiaceae (Hypericum) WA<br />
Melampsora euphorbiae-gerardianae? U-485 Euphorbiaceae (Euphorbia) Germany<br />
Melampsora euphorbiae U-138 [LSU only] Euphorbiaceae (Euphorbia II) Syria<br />
Melampsora epitea complex U-557 [multiple PPs] Salix sp. WA<br />
Melampsora sp. (SAL-83) S. lemmonii ID<br />
Melampsora laricis AB116809 Populus davidiana, Tibet, China<br />
Melampsora occidentalis U-395 Populus balsamifera II WA<br />
Melampsora (SAL-49) [LSU only] Pinaceae (Abies grandis I) ID<br />
Pucciniastrum agrimoniae AF426234 Rosaceae (Agrimonia eupatoria)<br />
Cronartium quercuum U-62 Fagaceae (Quercus macrocarpa) IN<br />
Cronartium quercuum U-432 Fagaceae (Quercus muehlenbergii II) IN<br />
Cronartium quercuum U-434 Fagaceae (Quercus alba II) IN<br />
Cronartium quercuum U-435 Fagaceae (Quercus bicolor II) IN<br />
Cronartium quercuum u-436 Fagaceae (Quercus macrocarpa II) IN<br />
Cronartium ribicola U-396/U-268 Ribes VA/Pinus WY<br />
Endocronartium harknessii AF522175<br />
Cronartium flaccidum AF426239 Asclepiadaceae (Vincetoxicum hirundinaria)<br />
Endocronartium harknessii AY700193<br />
Pucciniastrum agrimoniae? U-304 Rosaceae (Leucoside) South Africa [inc. seq.]<br />
Chrysomyxa arctostaphyli AF522163<br />
Chrysomyxa arctostaphyli AY700192<br />
Chrysomyxa ledi AF426246 Pinaceae (Picea abies)<br />
Chrysomyxa rhododendri AF426245 Ericaceae (Rhododendron ferrugineum)<br />
Caeoma tsukubaensis AB19124 Ericaceae (Rhododendron kaempferi) Japan<br />
Thekopsora areolata AF426235 Rosaceae (Prunus padus)<br />
Hyalopsora U-361 undet. fern (Woodsiaceae?) UT<br />
Hyalopsora polypodii AF426229 Dryopteridaceae (Cystopteris fragilis)<br />
Hyalopsora polypodii AY512852<br />
Pucciniastrum? MCA2884 ?? Costa Rica<br />
Melampsoridium betulinum AF426228 Fagaceae (Betula pendula)<br />
Naohidemyces vaccinii MCA 2780 Ericaceae (Vaccinium) WA<br />
Naohidemyces vaccinii AF426238 Ericaceae (Vaccinium uliginosum)<br />
Uredinopsis sp. AF522181<br />
Uredinopsis filicina AF426237 Thelypteridaceae (Phegopteris connectilis)<br />
Milesina (as Milesia) scolopendri AF426236 Aspleniaceae (Asplenium scolopendrium)<br />
Pucciniastrum epilobii AF426226 Onagraceae (Epilobium angustifolium)<br />
Pucciniastrum epilobii AF522178<br />
Pucciniastrum circaeae AY745697<br />
Pucciniastrum epilobii AF522179<br />
Pucciniastrum circaeae AF426227 Onagraceae (Circaea lutetiana)<br />
Melampsorella caryophyllacearum AF426232 Pinaceae (Abies alba)<br />
Pucciniastrum goeppertianum AF522180<br />
Coleosporium ipomoeae JRH-195 Convolvulaceae (Ipomoea nil II) Costa Rica<br />
Coleosporium ipomoeae JRH-485 Convolvulaceae (Merremia quinquefolia II-III) Puerto Rico<br />
Coleosporium ipomoeae JRH-465 Convolvulaceae (Ipomoea triloba II) P.R.<br />
Coleosporium ipomoeae JRH-216 Convolvulaceae (Ipomoea cholulensis II-III) Honduras<br />
Coleosporium ipomoeae JRH-168 Convolvulaceae (Ipomoea trifida II-III) Costa Rica<br />
Coleosporium ipomoeae JRH-230 Convolvulaceae (Ipomoea anisomeres II) Honduras<br />
Coleosporium ipomoeae JRH-143 Convolvulaceae (Ipomoea neeiII) Costa Rica<br />
Coleosporium plumierae U-112 Apocynaceae (Plumeria rubra II-III) Nigeria<br />
Coleosporium plumeriae U-50 Apocynaceae (Plumeria rubra II) Guyana<br />
Coleosporium plumeriae U-266 Apocynaceae (Plumeria II) GUAM<br />
Coleosporium (asterum or delicatulum?) U-369 Asteraceae (Solidago) NY<br />
Coleosporium asterum U-67 Asteraceae (Symphyotrichum (=Aster) racemosum II-III) MD<br />
Coleosporium asterum AF522165<br />
Coleosporium asterum AF426241 Asteraceae (Aster ciliolatus)<br />
Coleosporium (asterum or delicatulum?) MCA-2600 Asteraceae (Solidago II) TN<br />
Coleosporium U-430 Asteraceae (Solidago?) NJ<br />
Coleosporium U-389 Asteraceae (Solidago?) MD<br />
Coleosporium U-388 Asteraceae (Solidago?) PA<br />
Coleosporium (asterum or delicatulum?) U-428 Asteraceae (Solidago) MD<br />
Coleosporium (asterum or delicatulum?) MCA-2777 S(Asteraceae (Solidago canadensis II-III) MD<br />
Coleosporium asterum AF522164<br />
Coleosporium sp. U-398 Asteraceae MD<br />
Coleosporium asterum U-366 Asteraceae (Aster novae-angliae II-III?) MD<br />
Coleosporium (asterum or delicatulum?) U-347 Asteraceae (Solidago) PA<br />
Coleosporium vernoniae U-371 Asteraceae (Vernonia) IL<br />
Coleosporium tussilaginis U-74 Asteraceae (Senecio triangularis) OR<br />
Coleosporium tussilaginis MCA-2389 Asteraceae (Sonchus?) CA<br />
Coleosporium tussilaginis MCA-2392 Asteraceae (Sonchus or Lactuca) CA<br />
Coleosporium vernoniae GUY-25 Asteraceae (Elephantopus mollis II)<br />
Coleosporium senecionis AY512840<br />
Coleosporium tussilaginis AF426242 Asteraceae (Tussilago farfara)<br />
Coleosporium cacaliae AF426243 Asteraceae (Adenostyles glabra)<br />
Coleosporium campanulae AF426244 Campanulaceae (Campanula scheuchzeri)<br />
Thekopsora symphyti AF426230 Boraginaceae (Symphytum officinale)<br />
Thekopsora guttata AF426231 Rubiaceae (Galium odoratum)<br />
Gymnosporangium libocedri AF522168<br />
Phakopsora apoda PDD-72076 Poaceae (Pennisetum II) New Zealand<br />
Phakopsora apoda PDD-68451 Poaceae (Pennisetum II) New Zealand<br />
Dasturella divina BRIP-39289 Poaceae (Babusa II) Australia<br />
Dasturella divina MCA-2887 Poaceae (bamboo) Costa Rica<br />
Phakopsora? MCA2617 Poaceae (Panicum) NC<br />
Phakopsora phakopsoroides JRH-184 Poaceae (Olyra II) Costa Rica<br />
Phakopsora euvitis BRIP-27996 Vitaceae (Vitis II) Australia<br />
Phakopsora vernoniae U-256 [#368] Asteraceae (Vernonia) South Africa<br />
Uredo alpestris AF426212 Violaceae (Viola alpestris)<br />
Kuehneola loeseneriana? MCA-2879 Rosaceae (Rubus trichomallus?) Costa Rica<br />
Gerwasia (as Mainsia) pittieriana U-78 Rosaceae (Rubus) Guatemala<br />
Kuehneola uredinis U-429 Rosaceae (Rubus) MD<br />
Kuehneola uredinis U-303 Rosaceae (Rubus pinnatus) South Africa<br />
Kuehneola uredinis U-348 Rosaceae (Rubus) PA<br />
Kuehneola uredinis AY745696 AFTOL-987<br />
Kuehneola uredinis AF426218 Rosaceae (Rubus fruticosus)<br />
Kuehneola uredinis MCA-2393 Rosaceae (Rubus III) MD<br />
Kuehneola uredinis MCA-2867 Rosaceae (Rubus) MD<br />
Kuehneola uredinis MCA-2863 Rosaceae (Rubus) VA<br />
Frommeella mexicana var. indicae U-3 Rosaceae (Duchesnea II) MD<br />
FrommeÎlla mexicana MCA-2496 Rosaceae (Duchesnea) MD<br />
Frommeella mexicana MCA-2495 Rosaceae (Duchesnea II?) VA<br />
Phragmidium mucronatum U-106 Rosaceae (Rosa II) Argentina<br />
Phragmidium mucronatum? U-353 Rosaceae (Rosa) OR<br />
Phragmidium sanguisorbae AF426216 Rosaceae (Sanguisorba minor)<br />
Phragmidium violaceum AF426214 Rosaceae (Rubus fruticosus)<br />
Phragmidium violaceaum MCA-2782 Rosaceae (Rubus fruticosus? II-III) France<br />
Phragmidium violaceum U-645 Rosaceae (Rubus armeniacus II) OR<br />
Phragmidium rubi-idaei AF426215 Rosaceae (Rubus idaeus)<br />
Phragmidium fragariae AF426217 Rosaceae (Potentilla sterilis)<br />
Phragmidium montivagum AF426213 Rosaceae (Rosa cf. woodsii)<br />
Triphragmium ulmariae AF426219 Rosaceae (Filipendula ulmaria)<br />
Ochropsora ariae AF426221 Ranunculaceae (Anemone nemorosa)<br />
Ochropsora ariae AF426222 Rosaceae (Aruncus dioicus)<br />
Trachyspora intrusa AF426220 Rosaceae (Alchemilla vulgaris)<br />
Trachyspora intrusa MCA2384 Alchemilla (Rosaceae) Switzerland<br />
Caeoma torreyae AF522183<br />
Platygloea vestita AY512872<br />
Rhodotorula fushanensis AB176591<br />
16 GROUPS<br />
Identification<br />
Predictive Value<br />
Host Inference<br />
Dataset: Aime. 2006. Toward<br />
resolving family-level relationships in<br />
rust fungi (Uredinales). Mycoscience<br />
47(3):112-122. And unpublished.
Pucciniales<br />
rDNA SSU & LSU<br />
Data: Aime, M.C. 2006. Toward<br />
resolving family-level relationships<br />
in rust fungi (Uredinales).<br />
Mycoscience 47(3):112-122.<br />
Uropyxidaceae ?<br />
Chaconiaceae<br />
Pucciniosiraceae<br />
0.01 substitutions/site<br />
Pucciniaceae<br />
Phakopsoraceae<br />
Raveneliaceae<br />
Pileolariaceae<br />
Phakopsoraceae pp<br />
Gymnosporangium<br />
Phragmidiaceae<br />
Coleosporiaceae<br />
Cronartiaceae<br />
Melampsoraceae<br />
Mikronegeriaceae<br />
Caeoma torreyae<br />
Outgroups
Pucciniales<br />
rDNA SSU & LSU<br />
Data: Aime, M.C. 2006. Toward<br />
resolving family-level relationships<br />
in rust fungi (Uredinales).<br />
Mycoscience 47(3):112-122.<br />
Roman numeral = Group<br />
Arabic numeral = Type<br />
0.01 substitutions/site<br />
Pucciniaceae<br />
Phakopsoraceae<br />
Raveneliaceae<br />
Pileolariaceae<br />
Phakopsoraceae pp<br />
Gymnosporangium<br />
Phragmidiaceae<br />
Coleosporiaceae<br />
Cronartiaceae<br />
Melampsoraceae<br />
Mikronegeriaceae<br />
Caeoma torreyae<br />
Outgroups<br />
Spermagonia<br />
V (4)<br />
VI (7)<br />
VI (5,7)<br />
VI (5,7)<br />
VI (7)<br />
V (4)<br />
IV (6,10,11)<br />
VI (7)<br />
I (2)<br />
I (1,2,3)<br />
II (9)<br />
I (3)<br />
? [V (4); VI (5,7)]<br />
V (4)
Definitions for the purposes<br />
of this talk<br />
A host jump occurs when:<br />
a rust colonizes a host in a new family<br />
A host shift occurs when:<br />
A rust colonizes a new host within the<br />
same family, whether or not the two<br />
hosts are each others closest relatives.
Telial host/<br />
stage<br />
http://www.biology.ed.ac.uk/research/groups/jdeacon/FungalBiology/rust.htm<br />
Aecial host/<br />
stage
Pucciniales<br />
rDNA SSU & LSU<br />
Data: Aime, M.C. 2006. Toward<br />
resolving family-level relationships<br />
in rust fungi (Uredinales).<br />
Mycoscience 47(3):112-122.<br />
0.01 substitutions/site<br />
Pucciniaceae<br />
Phakopsoraceae<br />
Raveneliaceae<br />
Pileolariaceae<br />
Phakopsoraceae pp<br />
Gymnosporangium<br />
Phragmidiaceae<br />
Coleosporiaceae<br />
Cronartiaceae<br />
Melampsoraceae<br />
Mikronegeriaceae<br />
Caeoma torreyae<br />
Outgroups
Pucciniales<br />
rDNA SSU & LSU<br />
Data: Aime, M.C. 2006. Toward<br />
resolving family-level relationships<br />
in rust fungi (Uredinales).<br />
Mycoscience 47(3):112-122.<br />
Host JUMP versus<br />
Host SHIFT<br />
Jumps rare in aecial stage<br />
0.01 substitutions/site<br />
Pucciniaceae<br />
Phakopsoraceae<br />
Raveneliaceae<br />
Pileolariaceae<br />
Phakopsoraceae pp<br />
Gymnosporangium<br />
Phragmidiaceae<br />
Coleosporiaceae<br />
Cronartiaceae<br />
Melampsoraceae<br />
Mikronegeriaceae<br />
Caeoma torreyae<br />
Outgroups<br />
Aecial Host<br />
Angiosperm rusts<br />
(I-Angiosperms)<br />
Pinaceae rusts<br />
(I-Pinaceae)<br />
Basal lineages<br />
(I-non-Pinaceae conifers)
Pucciniales<br />
rDNA SSU & LSU<br />
Data: Aime, M.C. 2006. Toward<br />
resolving family-level relationships<br />
in rust fungi (Uredinales).<br />
Mycoscience 47(3):112-122.<br />
0.01 substitutions/site<br />
Pucciniaceae<br />
Phakopsoraceae<br />
Raveneliaceae<br />
Pileolariaceae<br />
Phakopsoraceae pp<br />
Gymnosporangium<br />
Phragmidiaceae<br />
Coleosporiaceae<br />
Cronartiaceae<br />
Melampsoraceae<br />
Mikronegeriaceae<br />
Caeoma torreyae<br />
Outgroups<br />
Ancestral rusts?<br />
Sjamsuridzal et al. 1999.<br />
Mycoscience 40:21-27.<br />
SSU 7 rusts, incl. Hyalospora<br />
and Uredinopsis
Pucciniales<br />
rDNA SSU & LSU<br />
Data: Aime, M.C. 2006. Toward<br />
resolving family-level relationships<br />
in rust fungi (Uredinales).<br />
Mycoscience 47(3):112-122.<br />
0.01 substitutions/site<br />
Pucciniaceae<br />
Phakopsoraceae<br />
Raveneliaceae<br />
Pileolariaceae<br />
Phakopsoraceae pp<br />
Gymnosporangium<br />
Phragmidiaceae<br />
Coleosporiaceae<br />
Cronartiaceae<br />
Melampsoraceae<br />
Mikronegeriaceae<br />
Caeoma torreyae<br />
Outgroups<br />
Ancestral rusts?<br />
Sjamsuridzal et al. 1999.<br />
Mycoscience 40:21-27.<br />
SSU 7 rusts, incl. Hyalospora<br />
and Uredinopsis<br />
Uredinopsis<br />
Photos: plantpath.cornell.edu
Ono & Hennen 1983 Mycol. Soc. Japan 24:369-402<br />
“But rust taxa on extant primitive host taxa<br />
are not necessarily more primitive than those<br />
on advanced hosts. <strong>Rust</strong>s, in both specific and<br />
generic levels, may jump to unrelated hosts in<br />
the course of evolution (Jackson 1931b; Leppik<br />
1953, 1955, 1959, 1967, 1972). Directions of<br />
this jump are oriented mostly by ecophysiological<br />
factors, and not always from primitive<br />
to advanced host taxa”.
Pucciniales<br />
rDNA SSU & LSU<br />
Data: Aime, M.C. 2006. Toward<br />
resolving family-level relationships<br />
in rust fungi (Uredinales).<br />
Mycoscience 47(3):112-122.<br />
Ancient conifers:<br />
Taxaceae (Torreya)<br />
Araucariaceae (Agathis,<br />
Araucaria—Southern<br />
hemisphere)<br />
Jurassic & Cretaceous<br />
(worldwide ~200-145<br />
mya)<br />
Heteroecious?<br />
0.01 substitutions/site<br />
Pucciniaceae<br />
Phakopsoraceae<br />
Raveneliaceae<br />
Pileolariaceae<br />
Phakopsoraceae pp<br />
Gymnosporangium<br />
Phragmidiaceae<br />
Coleosporiaceae<br />
Cronartiaceae<br />
Melampsoraceae<br />
Mikronegeriaceae<br />
Caeoma torreyae<br />
Outgroups<br />
Ancestral rusts?<br />
Blastospora<br />
Mikronegeria<br />
/Caeoma<br />
Photo: araucaria.org.uk
Pucciniales<br />
rDNA SSU & LSU<br />
Data: Aime, M.C. 2006. Toward<br />
resolving family-level relationships<br />
in rust fungi (Uredinales).<br />
Mycoscience 47(3):112-122.<br />
Co-evolution (? or just<br />
host shifting) with host<br />
in the telial stage<br />
0.01 substitutions/site<br />
Pucciniaceae<br />
Phakopsoraceae<br />
Raveneliaceae<br />
Pileolariaceae<br />
Phakopsoraceae pp<br />
Gymnosporangium<br />
Phragmidiaceae<br />
Coleosporiaceae<br />
Cronartiaceae<br />
Melampsoraceae<br />
Mikronegeriaceae<br />
Caeoma torreyae<br />
Outgroups<br />
Autoecious rusts?<br />
Multiple genera on Rosidae
Pucciniales<br />
rDNA SSU & LSU<br />
Data: Aime, M.C. 2006. Toward<br />
resolving family-level relationships<br />
in rust fungi (Uredinales).<br />
Mycoscience 47(3):112-122.<br />
Pucciniaceae<br />
excl.<br />
Gymnosporangium<br />
0.01 substitutions/site<br />
Pucciniaceae<br />
Phakopsoraceae<br />
Raveneliaceae<br />
Pileolariaceae<br />
Phakopsoraceae pp<br />
Gymnosporangium<br />
Phragmidiaceae<br />
Coleosporiaceae<br />
Cronartiaceae<br />
Melampsoraceae<br />
Mikronegeriaceae<br />
Caeoma torreyae<br />
Outgroups<br />
Pucciniaceae<br />
Puccinia<br />
Uromyces<br />
Cumminsiella<br />
etc<br />
Aecidium<br />
Endophyllum<br />
Dietelia<br />
Pucciniosira
Pucciniales<br />
rDNA SSU & LSU<br />
Data: Aime, M.C. 2006. Toward<br />
resolving family-level relationships<br />
in rust fungi (Uredinales).<br />
Mycoscience 47(3):112-122.<br />
Gymnosporangium<br />
0.01 substitutions/site<br />
Pucciniaceae<br />
Phakopsoraceae<br />
Raveneliaceae<br />
Pileolariaceae<br />
Phakopsoraceae pp<br />
Gymnosporangium<br />
Phragmidiaceae<br />
Coleosporiaceae<br />
Cronartiaceae<br />
Melampsoraceae<br />
Mikronegeriaceae<br />
Caeoma torreyae<br />
Outgroups<br />
Rosaceae <strong>Rust</strong>s<br />
Phragmidiaceae<br />
Incl. Triphragmium &<br />
Trachyspora<br />
Phragmidium<br />
Gymnoconium<br />
Kuehneola<br />
Etc.<br />
Rosidae vs.<br />
Maloideae
Pucciniales<br />
rDNA SSU & LSU<br />
Data: Aime, M.C. 2006. Toward<br />
resolving family-level relationships<br />
in rust fungi (Uredinales).<br />
Mycoscience 47(3):112-122.<br />
0.01 substitutions/site<br />
Pucciniaceae<br />
Phakopsoraceae<br />
Raveneliaceae<br />
Pileolariaceae<br />
Phakopsoraceae pp<br />
Gymnosporangium<br />
Phragmidiaceae<br />
Coleosporiaceae<br />
Cronartiaceae<br />
Melampsoraceae<br />
Mikronegeriaceae<br />
Caeoma torreyae<br />
Outgroups<br />
Pinaceae <strong>Rust</strong>s<br />
Coleosporium<br />
Chrysomyxa<br />
Cronartium (Peridermium<br />
& Endocronartium)<br />
Fern rusts (Hyalospora,<br />
Uredinopsis, Milesinia)<br />
Melampsoridium<br />
Melampsora
Pucciniales<br />
rDNA SSU & LSU<br />
Data: Aime, M.C. 2006. Toward<br />
resolving family-level relationships<br />
in rust fungi (Uredinales).<br />
Mycoscience 47(3):112-122.<br />
excl.<br />
Non-Mimosoidea<br />
(e.g. Sphenospora,<br />
Triphragmium)<br />
incl.<br />
Chaconiaceae p.p.<br />
0.01 substitutions/site<br />
Pucciniaceae<br />
Phakopsoraceae<br />
Raveneliaceae<br />
Pileolariaceae<br />
Phakopsoraceae pp<br />
Gymnosporangium<br />
Phragmidiaceae<br />
Coleosporiaceae<br />
Cronartiaceae<br />
Melampsoraceae<br />
Mikronegeriaceae<br />
Caeoma torreyae<br />
Outgroups<br />
Raveneliaceae<br />
Ravenelia<br />
Lipocystis<br />
Ypsilopsora<br />
Chaconia p.p.<br />
Etc.<br />
Uredo<br />
Images: Systematic Mycology and Microbiology, USDA
Pucciniales<br />
rDNA SSU & LSU<br />
Data: Aime, M.C. 2006. Toward<br />
resolving family-level relationships<br />
in rust fungi (Uredinales).<br />
Mycoscience 47(3):112-122.<br />
0.01 substitutions/site<br />
Pucciniaceae<br />
Phakopsoraceae<br />
Raveneliaceae<br />
Pileolariaceae<br />
Phakopsoraceae pp<br />
Gymnosporangium<br />
Phragmidiaceae<br />
Coleosporiaceae<br />
Cronartiaceae<br />
Melampsoraceae<br />
Mikronegeriaceae<br />
Caeoma torreyae<br />
Outgroups<br />
Phakopsoraceae<br />
Fabaceae,<br />
Euphorbiaceae,<br />
Moraceae, etc.<br />
Poaceae, Vitaceae
Systematics<br />
of Phakopsoraceae<br />
and Phakopsora
Phakopsoraceae Systematics<br />
~ 150 species<br />
~ 13 genera<br />
CONTENTIOUS. CONTENTIOUS. Based on morphology<br />
Spermogonia Group VI. Aecia Aecidium, Caeoma, or Uredo type<br />
(with or without peridium); aeciospores catenulate or borne singly. singly.<br />
Uredinia Malupa type, mostly with paraphyses;Telia erumpent or<br />
embedded, 2 to several spores deep; teliospores unicellular, sessile, sessile,<br />
catenulate or irregularly arranged. Heteroecious.<br />
Cummins & Hiratsuka. 2003. Illustrated Genera of the <strong>Rust</strong> Fungi
~ 90 species<br />
Phakopsora Systematics<br />
In the past, split into at least 5 genera (Physopella ( Physopella, ,<br />
Angiopsora, Bubakia, Stakmania) Stakmania)<br />
now considered synonyms<br />
> > 30 host families in >150 genera (monocots & dicots)<br />
Primarily Primarily tropical or subtropical<br />
CONTENTIOUS. CONTENTIOUS. Based on morphology<br />
Spermogonia Group VI (type 7). Aecia Calidion or Milesia type<br />
(subepidermal and paraphysate). Uredinia with or without paraphyses;Telia<br />
paraphyses;Telia<br />
subepidermal, non-erupent.<br />
non erupent.<br />
Cummins & Hiratsuka. 2003. Illustrated Genera of the <strong>Rust</strong> Fungi
Pucciniales<br />
rDNA SSU & LSU<br />
Data: Aime, M.C. 2006. Toward<br />
resolving family-level relationships<br />
in rust fungi (Uredinales).<br />
Mycoscience 47(3):112-122.<br />
0.01 substitutions/site<br />
Pucciniaceae<br />
Phakopsoraceae<br />
Raveneliaceae<br />
Pileolariaceae<br />
Phakopsoraceae pp<br />
Gymnosporangium<br />
Phragmidiaceae<br />
Coleosporiaceae<br />
Cronartiaceae<br />
Melampsoraceae<br />
Mikronegeriaceae<br />
Caeoma torreyae<br />
Outgroups<br />
Phakopsoraceae<br />
Fabaceae,<br />
Euphorbiaceae,<br />
Moraceae, etc.<br />
Poaceae, Vitaceae
Phakopsora sensu stricto<br />
“complete complete life cycles of most species are unknown” unknown – Ono et<br />
al. 1992. Mycol. Res. 96:825-850. 96:825 850.<br />
“production production of telia may not be necessary for most species to<br />
persist” persist – Ono et al. 1992. Mycol. Res. 96:825-850. 96:825 850.<br />
“Spermogonia Spermogonia and aecia are not known [for Phakopsora sensu<br />
stricto] and it is possible that some or all of them are<br />
heteroecious” heteroecious – Cummins & Hiratsuka 2003.
Phakopsoraceae<br />
rDNA LSU<br />
Phakopsoraceae s.s.<br />
Source: Aime unpubl.<br />
Phakopsora arthuriana Euphorbiaceae (Jatropha II) P.R.<br />
Phakopsora arthuriana Euphorbiaceae (Jatropha II) Guyana<br />
Phakopsora arthuriana Euphorbiaceae (Jatropha II) Guyana<br />
Phakopsora arthuriana Euphorbiaceae P.R.<br />
Phakopsora meibomiae Fabaceae (Lablab purpureus II) P.R.<br />
Phakopsora meibomiae Fabaceae (Phaseolus II) Honduras<br />
Phakopsora meibomiae Fabaceae (Phaseolus multiflorus II) Panama<br />
Phakopsora meibomiae Fabaceae (Desmodium II) Panama<br />
Phakopsora meibomiae Fabaceae (Lablab purpureus II) P.R.<br />
Phakopsora meibomiae Fabaceae (Lablab purpureus II) P.R.<br />
Phakopsora tecta Commelinaceae (Commelina II) P.R.<br />
Phakopsora tecta Commelinaceae (Commelina II) Costa Rica<br />
Phakopsora tecta Commelinaceae (Commelina II) Costa Rica<br />
Phakopsora tecta Commelinaceae (Commelina II) Puerto Rico<br />
Phakopsora tecta Commelinaceae (Aneilema) South Africa<br />
Phakopsora pachyrhizi Fabaceae (Glycine max) LA<br />
Phakopsora pachyrhizi Fabaceae (Glycine max) LA<br />
Phakopsora pachyrhizi Fabaceae (Glycine max) Zimbabwe<br />
Phakopsora gossypii Malvaceae (Gossypium II) Australia<br />
Uredopeltis sp nov? Tiliaceae (Grewia) South Africa<br />
Uredopeltis sp nov? Tiliaceae (Grewia) South Africa<br />
Batistopsora crucis-filii Annonaceae (Annona II-III) Guyana<br />
Phakopsora nishidana Moraceae (Ficus) P.R.<br />
Cerotelium fici Moraceae (Ficus II) Australia<br />
Phakopsora nishidana Moraceae (Ficus II) Argentina<br />
Prospodium cydistae Bignoniaceae (Cydista II-III) Costa Rica<br />
Prospodium Bignoniaceae (Clytostoma) Costa Rica<br />
Prospodium pithecoctenii Bignoniaceae (Pithecoctenium II-III) Costa Rica<br />
Prospodium pithecoctenii Bignoniaceae (Pithecoctenium II-III) Costa Rica<br />
Puccinia smilacis Smilacaceae (Smilax II-III) MD<br />
Puccinia smilacis Smilacaceae (Smilax) MD<br />
Prospodium lippiae Verbenaceae (Aloysia II-III) Argentina<br />
Prospodium appendiculatum Bignoniaceae (Tecoma) Mexica<br />
P. arthuriana / Euphorbiaceae<br />
P. meibomiae / Fabaceae<br />
P. tecta / Commelinaceae<br />
P. pachyrhizi / Fabaceae<br />
P. gossypii / Malvaceae<br />
Uredopeltis / Tiliaceae<br />
Batistopsora / Annonaceae<br />
P. nishidana & Cerotelium fici / Moraceae
Phakopsora pachyrhizi<br />
Autoecious lineages (e.g., Phragmidiaceae) appear restricted in in<br />
host<br />
family or subfamily<br />
Telial Telial stage known from nature on: Derris, Desmodium, Erythrina,<br />
Glycine, Ormocarpu, Pachyrhizus, Pueraria, Amphicarpa species in<br />
Philippines, China, Burma, Thailand, Japan, Taiwan and Nepal<br />
(as of Ono 1992).<br />
1 st description 1903, Japan; 1 st<br />
description of telia 1931, Taiwan)<br />
assuming assuming point of origin – aecial host (if<br />
it exists) likely in East Asia<br />
Map: Phuket.net
Acknowledgments<br />
Assembling the Fungal Tree of Life<br />
National Science Foundation<br />
DEB 0228657 and DEB 0732968<br />
Deep Hypha<br />
NSF Research Coordination <strong>Network</strong>: A Phylogeny for the<br />
Kingdom Fungi<br />
DEB 0090301<br />
Cindy Park & USDA-ARS-SBML; Ramandeep Kaur (LSU)<br />
David Hibbett & Brandon Matheny (Clark University)<br />
Deep Hypha<br />
Vol. 98(6).2006.