Cecidomyiidae - Ephydridae; pp. 98 - Nearctica

98 

Zeuxidiplosis Kieffer 1904 

Zeuxidiplosis giardi Kieffer 1896 (Diplosis) 

Diplosis giardiana Kieffer 1897 Syn. 

CERATOPOGONIDAE 

Allohelea Kieffer 1917 

Allohelea bottimeri Wirth 1991 (Allohelea) 

Allohelea distortifemur Wirth 1991 (Allohelea) 

Allohelea johannseni Wirth 1953 (Monohelea) 

Allohelea nebulosus Coquillett 1901 (Ceratopogon) 

Allohelea pedicellata Wirth 1991 (Allohelea) 

Allohelea weemsi Wirth 1991 (Allohelea) 

Alluaudomyia Kieffer 1913 

Neoceratopogon Malloch 1915 Syn. 

Prionognathus Carter, Ingram, and Macfie 1921 Homo. 

Thysanognathus Ingram and MacFie 1922 Syn. 

Isoecacta Garrett 1925 Syn. 

Alluaudomyia bellus Coquillett 1902 (Ceratopogon) 

Isoecacta poeyi Garrett 1925 Syn. 

Alluaudomyia footei Wirth 1952 (Alluaudomyia) 

Alluaudomyia megaparamera Williams 1957 (Alluaudomyia) 

Alluaudomyia needhami Thomsen 1935 (Alluaudomyia) 

Alluaudomyia paraspina Wirth 1952 (Alluaudomyia) 

Alluaudomyia parva Wirth 1952 (Alluaudomyia) 

Alluaudomyia downesi Wirth 1952 Syn. 

Alluaudomyia sitictipennis Wirth 1952 (Alluaudomyia) 

Alluaudomyia variegata Glick and Mullen 1982 (Alluadomyia) 

Alluaudomyia wirthi Williams 1957 (Alluaudomyia) 

Amerohelea Grogan and Wirth 1981 

Amerohelea frontispina Dow and Turner 1976 (Bezzia) 

Atrichopogon Kieffer 1906 

Gymnohelea Kieffer 1921 Syn. 

Dolichohelea Edwards 1929 Syn. 

Lophomyidium Cordero 1929 Syn. 

Psilokempia Enderlein 1936 Syn. 

Meloehelea Wirth 1956 Syn. 

Psammopogon Remm 1979 Syn. 

Rostropogon Remm 1979 Syn. 

Atrichopogon archboldi Wirth 1995 (Atrichopogon) 

Atrichopogon arcticus Coquillett 1900 (Ceratopogon) 

Atrichopogon borkenti Wirth 1994 (Atrichopogon) 

Atrichopogon corpulentus Ewen 1958 (Atrichopogon) 

Atrichopogon crinitus Ewen 1958 (Atrichopogon) 

Atrichopogon deyrupi Wirth 1994 (Atrichopogon) 

Atrichopogon downesi Wirth 1980 (Atrichopogon) 

Atrichopogon epicautae Wirth 1956 (Atrichopogon) 

Atrichopogon falcatus Boesel 1973 (Atrichopogon) 

Atrichopogon farri Wirth 1956 (Atrichopogon) 

Atrichopogon flavus Ewen 1958 (Atrichopogon) 

Atrichopogon fuscinervis Malloch 1915 (Ceratopogon) 

Atrichopogon fusculus Coquillett 1901 (Ceratopogn) 

Atrichopogon polydactylus Nielsen 1951 Syn. 

Atrichopogon geminus Boesel 1973 (Atrichopogon) 

Atrichopogon gilvus Coquillett 1905 (Ceratopogon) 

NOMINA INSECTA NEARCTICA 

DIPTERA: CECIDOMYIIDAE, CERATOPOGONIDAE 

Atrichopogon humicolus Ewen 1958 (Atrichopogon) 

Atrichopogon inconspicuus Ewen 1958 (Atrichopogon) 

Atrichopogon jamnbacki Wirth 1994 (Atrichopogon) 

Atrichopogon levis Coquillett 1901 (Ceratopogon) 

Ceratopogon exilis Coquillett 1902 Syn. 

Atrichopogon lucorum Meigen 1818 (Ceratopogon) 

Atrichopogon maculosus Ewen 1958 (Atrichopogon) 

Atrichopogon minutus Meigen 1830 (Ceratopogon) 

Atrichopogon occidentalis Wirth 1952 (Atrichopogon) 

Atrichopogon oedemerarum Storå 1939 (Atrichopogon) 

Atrichopogon peregrinus Johannsen 1908 (Ceratopogon) 

Atrichopogon titanus Boesel 1973 (Atrichopogon) 

Atrichopogon transiens Walker 1848 (Ceratopogon) 

Atrichopogon transversus Wirth 1952 (Atrichopogon) 

Atrichopogon warmkei Wirth 1956 (Atrichopogon) 

Atrichopogon websteri Coquillett 1901 (Ceratopogon) 

Atrichopogon wirthi Chan and Linley 1988 (Atrichopogon) 

Bezzia Kieffer 1899 

Pseudobezzia Malloch 1915 Syn. 

Allobezia Kieffer 1917 Syn. 

Lasiobezzia Kieffer 1925 Syn. 

Homobezzia Macfie 1932 Syn. 

Pygobezzia Remm 1974 Syn. 

Sivabezzia Remm 1974 Syn. 

Pygobezzia Remm 1974 Syn. 

Aspinabezzia Dow and Turner 1976 Syn. 

Aspinabezzia Dow and Turner 1976 Syn. 

Bezzia aklavikensis Wirth and Grogan 1983 (Bezzia) 

Bezzia albidorsata Malloch 1915 (Bezzia) 

Bezzia andersonorum Wirth and Grogan 1983 (Bezzia) 

Bezzia apicata Malloch 1914 (Bezzia) 

Bezzia biannulata Wirth 1952 (Bezzia) 

Bezzia bicolor Meigen 1804 (Ceratopogon) 

Probezzia typhae Kieffer 1919 Syn. 

Probezzia brehmiana Kieffer 1924 Syn. 

Probezzia algarum Zilahi-Sebess 1930 Syn. 

Probezzia rufithorax Goetghebuer 1932 Syn. 

Probezzia copiosa Thomsen 1935 Syn. 

Bezzia bilineata Wirth 1952 (Bezzia) 

Bezzia bivittatus Coquillett 1905 (Ceratopogon) 

Bezzia chelistyla Wirth and Grogan 1983 (Bezzia) 

Bezzia cockerelli Malloch 1915 (Bezzia) 

Bezzia obscura Malloch 1915 Syn. 

Bezzia coloradensis Wirth 1952 (Bezzia) 

Bezzia dentata Malloch 1914 (Bezzia) 

Bezzia dorsasetula Dow and Turner 1976 (Bezzia) 

Bezzia downesi Dow and Turner 1976 (Bezzia) 

Bezzia expolitus Coquillett 1901 (Ceratopogon) 

Ceratopogon johnsoni Coquillett 1901 Syn. 

Bezzia fairchildi Wirth 1983 (Bezzia) 

Bezzia fascispinosa Clastrier 1962 (Bezzia) 

Bezzia longiradia Dow and Turner 1976 Syn. 

Bezzia flavitarsis Malloch 1914 (Bezzia) 

Bezzia flavitibia Dow and Turner 1976 (Bezzia) 

Bezzia gibber Coquillett 1905 (Ceratopogon) 

Bezzia gibberella Wirth and Grogan 1983 (Bezzia) 

Bezzia glaber Coquillett 1902 (Ceratopogon) 

Bezzia imbifida Dow and Turner 1976 (Bezzia) 

Bezzia japonica Tokunaga 1939 (Bezzia) 

Bezzia suffusa Dow and Turner 1976 Syn. 

Bezzia laciniastyla Dow and Turner 1976 (Bezzia) 

Bezzia luteiventris Wirth and Grogan 1983 (Bezzia) 

Bezzia magnisetula Dow and Turner 1976 (Bezzia) 

Bezzia mallochi Wirth 1951 (Bezzia)

Bezzia medius Coquillett 1904 (Ceratopogon) 

Bezzia mohave Wirth and Grogan 1983 (Bezzia) 

Bezzia nigripes Wirth and Grogan 1983 (Bezzia) 

Bezzia nobilis Winnertz 1852 (Ceratopogon) 

Ceratopogon setulosa Loew 1861 Syn. 

Ceratopogon barberi Coquillett 1901 Syn. 

Bezzia armaticrus Kieffer 1919 Syn. 

Bezzia cinerella Kieffer 1919 Syn. 

Bezzia csikiana Kieffer 1919 Syn. 

Bezzia leucosticta Kieffer 1919 Syn. 

Bezzia chrysocoma Kieffer 1922 Syn. 

Bezzia atlantica Wirth and Williams 1956 Syn. 

Bezzia nobiliformis Clastrier 1962 Syn. 

Bezzia obelisca Dow and Turner 1976 (Bezzia) 

Bezzia perplexa Dow and Turner 1976 (Bezzia) 

Bezzia pruinosus Coquillett 1905 (Ceratopogon) 

Bezzia modocensis Wirth 1952 Syn. 

Bezzia pseudobscura Wirth 1951 (Bezzia) 

Bezzia pulvereus Coquillett 1901 (Ceratopogon) 

Bezzia punctipennis Williston 1896 (Ceratopogon) 

Bezzia saileri Wirth 1983 (Bezzia) 

Bezzia sandersoni Wirth and Grogan 1983 (Bezzia) 

Bezzia setosinotum Wirth and Grogan 1983 (Bezzia) 

Bezzia sordida Wirth 1952 (Bezzia) 

Bezzia spathula Wirth and Grogan 1983 (Bezzia) 

Bezzia spicata Dow and Turner 1976 (Bezzia) 

Bezzia texensis Wirth and Grogan 1983 (Bezzia) 

Bezzia twinni Wirth 1983 (Bezzia) 

Bezzia uncistyla Dow and Turner 1976 (Bezzia) 

Bezzia unispina Dow and Turner 1976 (Bezzia) 

Bezzia varicolor Coquillett 1902 (Ceratopogon) 

Brachypogon Kieffer 1899 

Isohelea Kieffer 1917 Syn. 

Nilohelea Kieffer 1921 Syn. 

Trishelea Kieffer 1924 Syn. 

Sarissohelea Debenham 1991 Syn. 

Brachypogon canadensis Downes 1976 (Brachypogon) 

Brachypogon pruinosa Wirth 1952 (Helea) 

Brachypogon serrata Lewis 1956 (Helea) 

Brachypogon stigmalis Coquillett 1902 (Ceratopogon) 

Ceratoculicoides Wirth and Ratanaworabhan 1971 

Ceratoculicoides blantoni Wirth and Ratanaworabhan 1971 

(Ceratoculicoides) 

Ceratoculicoides longipennis Wirth 1952 (Helea) 

Ceratoculicoides virginiana Wirth 1951 (Helea) 

Ceratopogon Meigen 1803 

Helea Meigen 1800 Unav. 

Psilohelea Kieffer 1915 Syn. 

Anakempia Kieffer 1924 Syn. 

Diplohelea Kieffer 1925 Syn. 


Ceratopogon abstrusus Borkent and Grogan 1995 (Ceratopogon) 

Ceratopogon algidus Borkent and Grogan 1995 (Ceratopogon) 

Ceratopogon annettae Borkent and Grogan 1995 (Ceratopogon) 

Ceratopogon arcanus Borkent and Grogan 1995 (Ceratopogon) 

Ceratopogon boomerangus Borkent and Grogan 1995 

(Ceratopogon) 

Ceratopogon cinaedicus Borkent and Grogran 1995 (Ceratopogon) 

Ceratopogon culicoidithorax Hoffman 1926 (Ceratopogon) 

Ceratopogon curtus Borkent and Grogan 1995 (Ceratopogon) 

Ceratopogon gigaforceps Remm 1973 (Ceratopogon) 

Ceratopogon inverecundus Borkent and Grogan 1995 

(Ceratopogon) 

Ceratopogon mallochi Cole 1921 (Hartomyia) 

Ceratopogon mammulus Borkent and Grogan 1995 (Ceratopogon) 

Ceratopogon multisetosus Borkent and Grogan 1995 (Ceratopogon) 

Ceratopogon pallicoleatus Borkent and Cole 1995 (Ceratopogon) 

Ceratopogon pandus Borkent and Grogan 1995 (Ceratopogon) 

Ceratopogon pontis Borkent and Grogan 1995 (Ceratopogon) 

Ceratopogon pubiantennalis Borkent and Grogan 1995 

(Ceratopogon) 

Ceratopogon rectus Borkent and Grogran 1995 (Ceratopogon) 

Ceratopogon relictus Borkent and Grogan 1995 (Ceratopogon) 

Ceratopogon scutellatus Say 1829 (Ceratopogon) 

Ceratopogon seculus Borkent and Grogan 1995 (Ceratopogon) 

Ceratopogon willisi Borkent and Grogan 1995 (Ceratopogon) 

Clinohelea Kieffer 1917 

Clinohelea bimaculatus Loew 1861 (Ceratopogon) 

Clinohelea curriei Coquillett 1905 (Ceratopogon) 

Palpomyia nebulosa Malloch 1915 Syn. 

Clinohelea dimidiatus Adams 1903 (Ceratopogon) 

Clinohelea longitheca Grogan and Wirth 1975 (Clinohelea) 

Clinohelea nubiferus Coquillett 1905 (Ceratopogon) 

Clinohelea pseudonubifera Grogan and Wirth 1975 (Clinohelea) 

Clinohelea usingeri Wirth 1952 (Clinohelea) 

Culicoides Latreille 1809 

Oecacta Poey 1851 Syn. 

Psychophaena Philippi 1865 Syn. 

Haematomyidium Goeldi 1905 Syn. 

Cotocripus Brethes 1912 Syn. 

Diplosella Kieffer 1922 Syn. 

Prosapelma Kieffer 1925 Syn. 

Haemophoructus Macfie 1925 Syn. 

Hoffmania Fox 1948 Syn. 

Beltranmyia Vargas 1953 Syn. 

Monoculicoides Khalaf 1954 Syn. 

Selfia Khalaf 1954 Syn. 

Avaritia Fox 1955 Syn. 

Macfiella Fox 1955 Syn. 

Trithecoides Wirth and Hubert 1959 Syn. 

Drymodesmyia Vargas 1960 Syn. 

Diphaomyia Vargas 1960 Syn. 

Anilomyia Vargas 1960 Syn. 

Glaphiromyia Vargas 1960 Syn. 

Mataemyia Vargas 1960 Syn. 

Meijerehelea Wirth and Hubert 1961 Syn. 

Pontoculicoides Remm and Zhogolev 1968 Syn. 

Callotia Vargas and Kremer 1972 Syn. 

Wirthomyia Vargas 1973 Syn. 

Sensiculicoides Shevchenko 1974 Syn. 

Silvatoculicoides Glukhova 1975 Syn. 

Remmia Glukhova 1977 Syn. 

Silvaticulicoides Glukhova 1977 Syn. 

Nullicella Lee 1982 Syn. 

Jilinocoides Chu 1983 Syn. 

Sinocoides Chu 1983 Syn. 

Silvicola Mirzaeva and Isaev 1990 Syn. 

Culicoides alachua Jamnback and Wirth 1963 (Culicoides) 

Culicoides alaskensis Wirth 1951 (Culicoides) 

Culicoides alexanderi Wirth and Hubert 1962 (Culicoides) 

Culicoides arboricola Root and Hoffman 1937 (Culicoides) 

Culicoides arizonensis Wirth and Hubert 1960 (Culicoides) 

Culicoides arubae Fox and Hoffman 1944 (Culicoides) 

Culicoides atchleyi Wirth and Blanton 1969 (Culicoides) 

Culicoides barbosai wirth and Blanton 1956 (Culicoides) 

Culicoides baueri Hoffman 1925 (Culicoides) 

Culicoides beckae Wirth and Blanton 1967 (Culicoides) 

DIPTERA: CERATOPOGONIDAE 

99

100 

Culicoides bergi Cochrane 1973 (Culicoides) 

Culicoides bermudensis Williams 1956 (Culicoides) 

Culicoides bickleyi Wirth and Hubert 1962 (Culicoides) 

Culicoides biguttatus Coquillett 1901 (Ceratopogon) 

Culicoides blantoni Vargas and Wirth 1955 (Culicoides) 

Culicoides bottimeri Wirth 1955 (Culicoides) 

Culicoides boydi Wirth and Mullens 1992 (Culicoides) 

Culicoides brookmani Wirth 1952 (Culicoides) 

Culicoides butleri Wirth and Hubert 1960 (Culicoides) 

Culicoides byersi Atchley 1967 (Culicoides) 

Culicoides cacticola Wirth and Hubert 1960 (Culicoides) 

Culicoides calexicanus Wirth and Rowley 1971 (Culicoides) 

Culicoides californiensis Wirth and Blanton 1967 (Culicoides) 

Culicoides canadensis Wirth and Blanton 1969 (Culicoides) 

Culicoides cavaticus Wirth and Jones 1956 (Culicoides) 

Culicoides chiopterus Meigen 1830 (Ceratopogon) 

Culicoides cochisensis Wirth and Blanton 1967 (Culicoides) 

Culicoides cockerellii Coquillett 1901 (Ceratopogon) 

Culicoides copiosus Root and Hoffman 1937 (Culicoides) 

Culicoides crepuscularis Malloch 1915 (Culicoides) 

Culicoides daedalus Macfie 1948 (Culicoides) 

Culicoides davisi Wirth and Rowley 1971 (Culicoides) 

Culicoides debilipalpis Lutz 1913 (Culicoides) 

Culicoides khalafi Beck 1957 Syn. 

Culicoides defoliarti Atchley and Wirth 1979 (Culicoides) 

Culicoides denningi Foote and Pratt 1954 (Culicoides) 

Culicoides denticulatus Wirth and Hubert 1962 (Culicoides) 

Culicoides dickei Jones 1956 (Culicoides) 

Culicoides doeringae Atchley 1967 (Culicoides) 

Culicoides downesi Wirth and Hubert 1962 (Culicoides) 

Culicoides eadsi Wirth and Blanton 1971 (Culicoides) 

Culicoides edeni Wirth and Blanton 1974 (Culicoides) 

Culicoides erikae Atchley and Wirth 1979 (Culicoides) 

Culicoides floridensis Beck 1951 (Culicoides) 

Culicoides flukei Jones 1956 (Culicoides) 

Culicoides footei Wirth and Jones 1956 (Culicoides) 

Culicoides franclemonti Cochrane 1974 (Culicoides) 

Culicoides freeborni Wirth and Blanton 1969 (Culicoides) 

Culicoides frohnei Wirth and Blanton 1969 (Culicoides) 

Culicoides furens Poey 1851 (Oecata) 

Ceratopogon maculithorax Williston 1896 Syn. 

Culicoides dovei Hall 1932 Syn. 

Culicoides furensoides Williams 1955 (Culicoides) 

Culicoides gigas Root and Hoffman 1937 (Culicoides) 

Culicoides gregsoni Wirth and Blanton 1969 (Culicoides) 

Culicoides guttipennis Coquillett 1901 (Ceratopogon) 

Culicoides haematopotus Malloch 1915 (Culicoides) 

Culicoides hawsi Wirth and Rowley 1971 (Culicoides) 

Culicoides hieroglyphicus Malloch 1915 (Culicoides) 

Culicoides hinmani Khalaf 1952 (Culicoides) 

Culicoides hirtulus Coquillett 1900 (Ceratopogon) 

Culicoides hoguei Wirth and Assis de Moraes 1979 (Culicoides) 

Culicoides hollensis Melander and Brues 1903 (Ceratopogon) 

Culicoides canithorax Hoffman 1925 Syn. 

Culicoides husseyi Wirth and Blanton 1971 (Culicoides) 

Culicoides insignis Lutz 1913 (Culicoides) 

Culicoides inamollae Fox and Hoffman 1944 Syn. 

Culicoides inyoensis Wirth and Blanton 1969 (Culicoides) 

Culicoides jacksoni Atchley 1970 (Culicoides) 

Culicoides jamaicensis Edwards 1922 (Culicoides) 

Culicoides jamesi Fox 1946 (Culicoides) 

Culicoides jamnbacki Wirth and Hubert 1962 (Culicoides) 

Culicoides jonesi Wirth and Hubert 1960 (Culicoides) 

Culicoides juddi Cochrane 1974(Culicoides) 

Culicoides knowltoni Beck 1956 (Culicoides) 

Culicoides kubunensis Tokunaga 1937 (Culicoides) 

Culicoides cubitalis Edwards 1939 Syn. 

Culicoides lahontan Wirth and Blanton 1969 (Culicoides) 



Culicoides leechi Wirth 1977 (Culicoides) 

Culicoides loisae Jamnback 1965 (Culicoides) 

Culicoides lophortygis Atchley and Wirth 1975 (Culicoides) 

Culicoides loughnani Edwards 1922 (Culicoides) 

Culicoides luglani Jones and Wirth 1958 (Culicoides) 

Culicoides luteovenus Root and Hoffman 1937 (Culicoides) 

Culicoides melleus Coquillett 1901 (Ceratopogon) 

Culicoides minutissimus Zetterstedt 1855 (Ceratopogon) 

Culicoides mississippiensis Hoffman 1926 (Culicoides) 

Culicoides mohave Wirth 1952 (Culicoides) 

Culicoides monoensis Wirth 1952 (Culicoides) 

Culicoides mortivallis Wirth and Blanton 1971 (Culicoides) 

Culicoides mulrennani Beck 1957 (Culicoides) 

Culicoides multidentatus Atchley and Wirth 1975 (Culicoides) 

Culicoides multipunctatus Malloch 1915 (Culicoides) 

Culicoides nanellus Wirth and Blanton 1969 (Culicoides) 

Culicoides nanus Root and Hoffman 1937 (Culicoides) 

Culicoides neofagineus Wirth and Blanton 1969 (Culicoides) 

Culicoides neomontanus Wirth 1976 (Culicoides) 

Culicoides montanus Wirth and Blanton 1969 Homo. 

Culicoides neopulicaris Wirth 1955 (Culicoides) 

Culicoides niger Root and Hoffman 1937 (Culicoides) 

Culicoides novamexicanus Atchley 1967 (Culicoides) 

Culicoides obsoletus Meigen 1818 (Ceratopogon) 

Culicoides occidentalis Wirth and Jones 1957 (Culicoides) 

Culicoides albertensis Wirth and Jones 1957 Syn. 

Culicoides australis Wirth and Jones 1957 Syn. 

Culicoides sonorensis Wirth and Jones 1957 Syn. 

Culicoides oklahomensis Khalaf 1952 (Culicoides) 

Culicoides oregonensis Wirth and Rowley 1971 (Culicoides) 

Culicoides ousairani Khalaf 1952 (Culicoides) 

Culicoides owyheenis Jones and Wirth 1978 (Culicoides) 

Culicoides palmerae James 1943 (Culicoides) 

Culicoides paraensis Goeldi 1905 (Haematomyidium) 

Culicoides parapiliferus Wirth and Blanton 1974 (Culicoides) 

Culicoides pechumani Cochrane 1974 (Culicoides) 

Culicoides pecosensis Wirth 1955 (Culicoides) 

Culicoides pifanoi Ortiz 1951 (Culicoides) 

Culicoides piliferus Root and Hoffman 1937 (Culicoides) 

Culicoides posoensis Wirth and Blanton 1969 (Culicoides) 

Culicoides pseudopiliferus Wirth and Hubert 1962 (Culicoides) 

Culicoides pusillus Lutz 1913 (Culicoides) 

Culicoides reevesi Wirth 1952 (Culicoides) 

Culicoides riggsi Khalaf 1957 (Culicoides) 

Culicoides ryckmani Wirth and Hubert 1960 (Culicoides) 

Culicoides salihi Khalaf 1952 (Culicoides) 

Culicoides saltonensis Wirth 1952 (Culicoides) 

Culicoides sanguisuga Coquillett 1901 (Ceratopogon) 

Simulium nocivum Harris 1862 Unav. 

Culicoides saundersi Wirth and Blanton 1969 (Culicoides) 

Culicoides scanloni Wirth and Hubert 1962 (Culicoides) 

Culicoides sierrensis Wirth and Blanton 1969 (Culicoides) 

Culicoides sitiens Wirth and Hubert 1960 (Culicoides) 

Culicoides snowi Wirth and Jones 1956 (Culicoides) 

Culicoides sommermanae Wirth and Blanton 1969 (Culicoides) 

Culicoides sordidellus Zetterstedt 1838 (Ceratopogon) 

Culicoides sphagnumensis Williams 1955 (Culicoides) 

Culicoides spinosus Root and Hoffman 1937 (Culicoides) 

Culicoides stellifer Coquillett 1901 (Ceratopogon) 

Culicoides stilobezzioides Foote and Pratt 1954 (Culicoides) 

Culicoides stonei James 1943 (Culicoides) 

Culicoides weesei Khalaf 1952 Syn. 

Culicoides sublettei Atchley 1967 (Culicoides) 

Culicoides tenuistylus Wirth 1952 (Culicoides) 

Culicoides testudinalis Wirth and Hubert 1962 (Culicoides) 

Culicoides tissoti Wirth and Blanton 1966 (Culicoides) 

Culicoides torreyae Wirth and Blanton 1971 (Culicoides) 

Culicoides travisi Vargas 1949 (Culicoides)

Culicoides simulans Root and Hoffman 1937 Homo. 

Culicoides horneae Foote and Pratt 1954 Syn. 

Culicoides tristriatulus Hoffman 1925 (Culicoides) 

Culicoides unicolor Coquillett 1905 (Ceratopogon) 

Culicoides usingeri Wirth 1952 (Culicoides) 

Culicoides utahensis Fox 1946 (Culicoides) 

Culicoides utowana Jamnback 1965 (Culicoides) 

Culicoides variipennis Coquillett 1901 (Ceratopogon) 

Culicoides venustus Hoffman 1925 (Culicoides) 

Culicoides villosipennis Root and Hoffman 1937 (Culicoides) 

Culicoides werneri Wirth and Blanton 1971 (Culicoides) 

Culicoides wirthi Foote and Pratt 1954 (Culicoides) 

Culicoides wisconsinensis Jones 1956 (Culicoides) 

Culicoides yukonensis Hoffman 1925 (Culicoides) 

Dasyhelea Kieffer 1911 

Prokempia Kieffer 1913 Syn. 

Pseudoculicoides Malloch 1915 Syn. 

Tetrahelea Kieffer 1925 Syn. 

Cryptoscena Enderlein 1936 Syn. 

Dicryptoscena Enderlein 1936 Syn. 

Sebessia Remm 1979 Syn. 

Dasyhelea ancorus Coquillett 1902 (Ceratopogon) 

Dasyhelea atlantis Wirth and Williams 1957 (Dasyhelea) 

[Dasyhelea atlantis Waugh and Wirth 1976 (Dasyhelea) Homo.] 

Dasyhelea atrata Wirth 1952 (Dasyhelea) 

Dasyhelea bahamensis Johnson 1908 (Ceratopogon) 

Dasyhelea bajensis Wirth 1978 (Dasyhelea) 

Dasyhelea bermudae Wirth and Williams 1957 (Dasyhelea) 

Dasyhelea brevicornis Waugh and Wirth 1976 (Dasyhelea) 

Dasyhelea brevicosta Waugh and Wirth 1976 (Dasyhelea) 

Dasyhelea brookmani Wirth 1952 (Dasyhelea) 

Dasyhelea cactorum Wirth and Hubert 1960 (Dasyhelea) 

Dasyhelea chani Wirth and Linley 1990 (Dasyhelea) 

Dasyhelea cinctus Coquillett 1901 (Ceratopogon) 

Dasyhelea crassiseta Borkent and Forster 1986 (Dasyhelea) 

Dasyhelea festiva Wirth 1952 (Dasyhelea) 

Dasyhelea griseus Coquillett 1901 (Ceratopogon) 

Dasyhelea subcaerulea Thomsen 1935 Syn. 

Dasyhelea johannseni Malloch 1915 (Pseudoculicoides) 

Dasyhelea leptobranchia Waugh and Wirth 1976 (Dasyhelea) 

Dasyhelea luteogrisea Wirth and Williams 1957 (Dasyhelea) 

Dasyhelea major Malloch 1915 (Pseudoculicoides) 

Dasyhelea messersmithi Waugh and Wirth 1976 (Dasyhelea) 

Dasyhelea mutabilis Coquillett 1901 (Ceratopogon) 

Dasyhelea navaiae Waugh and Wirth 1976 (Dasyhelea) 

Dasyhelea neobifurcata Wirth 1976 (Dasyhelea) 

Dasyhelea bifurcata Wirth 1952 Homo. 

Dasyhelea oppressa Thomsen 1935 (Dasyhelea) 

Dasyhelea pallens Wirth 1952 (Dasyhelea) 

Dasyhelea pentalineata Wirth and Hubert 1960 (Dasyhelea) 

Dasyhelea pollinosa Wirth 1952 (Dasyhelea) 

Dasyhelea pritchardi Wirth 1952 (Dasyhelea) 

Dasyhelea pseudocincta Waugh and Wirth 1976 (Dasyhelea) 

Dasyhelea pseudoincisurata Waugh and Wirth 1976 (Dasyhelea) 

Dasyhelea ryckmani Wirth and Hubert 1960 (Dasyhelea) 

Dasyhelea sanctaemariae Wirth 1952 (Dasyhelea) 

Dasyhelea scissurae Macfie 1937 (Dasyhelea) 

Dasyhelea scutellatus Meigen 1830 (Ceratopogon) 

Dasyhelea sonorensis Wirth 1978 (Dasyhelea) 

Dasyhelea spiniforma Waugh and Wirth 1976 (Dasyhelea) 

Dasyhelea stemlerae Waugh and Wirth 1976 (Dasyhelea) 

Dasyhelea sublettei Wirth 1987 (Dasyhelea) 

Dasyhelea tenebrosus Coquillett 1905 (Ceratopogon) 

Dasyhelea thomsenae Wirth 1952 (Dasyhelea) 

Dasyhelea traverae Thomson 1935 (Dasyhelea) 

Dasyhelea tristyla Wirth 1952 (Dasyhelea) 


Downeshelea Wirth and Grogan 1988 

Downeshelea stonei Wirth 1953 (Monohelea) 

Echinohelea Macfie 1940 

Echinoideshelea Wirth 1994 Syn. 

Echinohelea lanei Wirth 1951 (Echinohelea) 

Forcipomyia Meigen 1818 

Labidomyia Stephens 1829 Syn. 

Euprojoannisia Brethes 1914 Syn. 

Euforcipomyia Malloch 1915 Syn. 

Microhelea Kieffer 1917 Syn. 

Lepidohelea Kieffer 1917 Syn. 

Eupelma Kieffer 1919 Syn. 

Trichohelea Goetghebuer 1920 Syn. 

Lasiohelea Kieffer 1921 Syn. 

Thyridomyia Saunders 1925 Syn. 

Pterobosca Macfie 1932 Syn. 

Synthyridomyia Saunders 1956 Syn. 

Proforcipomyia Saunders 1956 Syn. 

Metaforcipomyia Saunders 1956 Syn. 

Caloforcipomyia Saunders 1956 Syn. 

Warmkea Saunders 1956 Syn. 

Neoforcipomyia Takunaga 1959 Syn. 

Dacnoforcipomyia Chan and Saunders 1965 Syn. 

Blantonia Wirth and Dow 1971 Syn. 

Schizoforcipomyia Chan and LeRoux 1971 Syn. 

Phytohelea Remm 1971 Syn. 

Rhynchoforcipomyia Wirth and Dow 1972 Syn. 

Saliohelea Wirth and Ratanaworabhan 1978 Syn. 

Rhinohelea Meillon and Wirth 1979 Syn. 

Pedilohelea Meillon and Wirth 1980 Syn. 

Panhelea Remm 1981 Syn. 

Collessohelea Debenham 1987 Syn. 

Herakleohelea Debenham 1987 Syn. 

Kattangomyia Debenham 1987 Syn. 

Schineromyia Debenham 1987 Syn. 

Typhonomyia Debenham 1987 Syn. 

Bassoforcipomyia Debenham 1987 Syn. 

Dycea Debenham 1987 Syn. 

Forcipohelea Clastrier and Delecolle 1991 Syn. 

Forcipomyia acinacis Wirth and Spinelli 1993 (Forcipomyia) 

Forcipomyia aurea Malloch 1915 (Forcipomyia) 

Forcipomyia basifemoralis Wirth and Spinelli 1992 (Forcipomyia) 

Forcipomyia baueri Wirth 1956 (Forcipomyia) 

Forcipomyia beckae Wirth 1976 (Forcipomyia) 

Forcipomyia bipunctata Linnaeus 1767 (Tipula) 

Forcipomyia bipunctatopropinqua Chan and LeRoux 1971 

(Forcipomyia) 

Forcipomyia blantoni Soria and Bystrak 1975 (Forcipomyia) 

Forcipomyia brevipennis Macquart 1826 (Ceratopogon) 

Ceratopogon specularis Coquillett 1901 Syn. 

Forcipomyia bystraki Grogan and Wirth 1975 (Forcipomyia) 

Forcipomyia calcaratus Coquillett 1905 (Ceratopogon) 

Forcipomyia canadensis Bystrak and Wirth 1978 (Forcipomyia) 

Forcipomyia christiansoni Wirth and Hubert 1960 (Forcipomyia) 

Forcipomyia ciliatus Winnertz 1852 (Ceratopogon) 

Forcipomyia cilipes Coquillett 1900 (Ceratopogon) 

Forcipomyia cinctipes Coquillett 1905 (Ceratopogon) 

Forcipomyia cochisei Wirth and Spinelli 1993 (Forcipomyia) 

Forcipomyia colemani Wirth 1952 (Forcipomyia) 

Forcipomyia concolor Malloch 1915 (Forcipomyia) 

Forcipomyia crinita Saunders 1964 (Forcipomyia) 

Forcipomyia desertensis Wirth and Hubert 1960 (Forcipomyia) 


101

102 



Forcipomyia dowi Bystrak and Wirth 1978 (Forcipomyia) 

Forcipomyia dubiamima Wirth and Spinelli 1993 (Forcipomyia) 

Forcipomyia eadsi Wirth and Spinelli 1993 (Forcipomyia) 

Forcipomyia edmistoni Wirth and Spinelli 1993 (Forcipomyia) 

Forcipomyia elegantula Malloch 1915 (Forcipomyia) 

Forcipomyia eques Johannsen 1908 (Ceratopogon) 

Forcipomyia chrysopae Mayer 1934 Syn. 

Lasiohelea chrysopivora Tokunaga 1939 Syn. 

Forcipomyia fairfaxensis Wirth 1951 (Forcipomyia) 

Forcipomyia fimbriatus Coquillett 1901 (Ceratopogon) 

Forcipomyia floridensis Dow and Wirth 1972 (Forcipomyia) 

Forcipomyia frutetorum Winnertz 1852 (Ceratopogon) 

Forcipomyia aethiopiae Ingram and Macfie 1924 Syn. 

Lasiohelea japonica Tokunaga 1937 Syn. 

Forcipomyia aspinosa Saunders 1956 Syn. 

Forcipomyia fuliginosus Meigen 1818 (Ceratopogon) 

Forcipomyia erucicida Knab 1914 Syn. 

Ceratopogon coquilletti Kieffer 1917 Syn. 

Forcipomyia brookmani Wirth 1952 Syn. 

Forcipomyia intonsa Chan and LeRoux 1971 Syn. 

Forcipomyia fuscicalcarata Bystrak and Wirth 1978 (Forcipomyia) 

Forcipomyia fusicornis Coquillett 1905 (Ceratopogon) 

Ceratopogon fur Johnson 1913 Syn. 

Pterobosca foridana Johannsen 1950 Syn. 

Forcipomyia genualis Loew 1866 (Ceratopogon) 

Forcipomyia raleighi Macfie 1938 Syn. 

Forcipomyia glauca Macfie 1934 (Forcipomyia) 

Forcipomyia monilis Goetghebuer 1934 Syn. 

Forcipomyia diversipes Goetghebuer 1936 Syn. 

Forcipomyia splendida Wirth 1951 Syn. 

Forcipomyia goniognatha Wirth and Messersmith 1971 

(Forcipomyia) 

Forcipomyia gravesi Wirth and Spinelli 1992 (Forcipomyia) 

Forcipomyia hirtipennis Malloch 1915 (Euforcipomyia) 

Forcipomyia turfacea Kieffer 1924 Syn. 

Forcipomyia hurdi Wirth 1952 (Forcipomyia) 

Forcipomyia hygrophila Kieffer 1924 (Forcipomyia) 

Forcipomyia occidentalis Wirth 1952 Syn. 

Forcipomyia indecora Kieffer 1914 (Forcipomyia) 

Forcipomyia ingrami Carter 1919 Syn. 

Forcipomyia johannseni Thomsen 1935 (Forcipomyia) 

Forcipomyia leptognatha Wirth and Messersmith 1971 

(Forcipomyia) 

Forcipomyia longitarsis Malloch 1915 (Euforcipomyia) 

Forcipomyia luteigenua Wirth and Spinelli 1992 (Forcipomyia) 

Forcipomyia macswaini Wirth 1952 (Forcipomyia) 

Forcipomyia mcateei Wirth 1956 (Forcipomyia) 

Forcipomyia saundersi Chan and LeRoux 1965 Syn. 

Forcipomyia mexicana Wirth 1956 (Forcipomyia) 

Forcipomyia monilicornis Coquillett 1905 (Ceratopogon) 

Ceratopogon hirtus Lundstrom 1910 Syn. 

Thyridomyia palustris Saunders 1925 Syn. 

Forcipomyia curticornis Goetghebuer 1933 Syn. 

Forcipomyia mortuifolii Saunders 1959 (Forcipomyia) 

Forcipomyia murinus Winnertz 1852 (Ceratopogon) 

Forcipomyia sulfurea Kieffer 1923 Syn. 

Forcipomyia hirtipalpis Kieffer 1924 Syn. 

Forcipomyia sate Kieffer 1925 Syn. 

Forcipomyia longitarsis Tokunaga 1940 Syn. 

Forcipomyia moascari Macfie 1943 Syn. 

Forcipomyia attonsa Goetghebuer 1950 Syn. 

Forcipomyia sibmurina Remm 1980 Syn. 

Forcipomyia navaiae Bystrak and Wirth 1978 (Forcipomyia) 

Forcipomyia nodosa Saunders 1959 (Forcipomyia) 

Forcipomyia obscurus Walker 1848 (Ceratopogon) 

Forcipomyia pallidus Winnertz 1852 (Ceratopogon) 

Forcipomyia papilionivora Edwards 1923 (Forcipomyia) 

Forcipomyia parvus Walker 1848 (Ceratopogon) 

Forcipomyia pechumani Bystrak and Wirth 1978 (Forcipomyia) 

Forcipomyia pergandei Coquillett 1901 (Ceratopogon) 

Forcipomyia piceus Winnertz 1852 (Ceratopogon) 

Forcipomyia simulata Walley 1932 Syn. 

Forcipomyia pictoni Macfie 1938 (Forcipomyia) 

Forcipomyia pilosus Coquillett 1902 (Ceratopogon) 

Forcipomyia pinicola Bystrak and Messersmith 1980 (Forcipomyia) 

Forcipomyia pluvialis Malloch 1923 (Forcipomyia) 

Forcipomyia pricei Wirth and Spinelli 1993 (Forcipomyia) 

Forcipomyia quadiingrami Macfie 1939 (Forcipomyia) 

Forcipomyia quatei Wirth 1952 (Forcipomyia) 

Forcipomyia rugosa Chan and LeRoux 1970 (Forcipomyia) 

Forcipomyia seminole Wirth 1976 (Forcipomyia) 

Forcipomyia solonensis Wirth 1951 (Forcipomyia) 

Forcipomyia sonora Wirth 1952 (Forcipomyia) 

Forcipomyia squamipes Coquillett 1902 (Ceratopogon) 

Ceratopogon brumalis Long 1902 Syn. 

Forcipomyia stenammatis Long 1902 (Ceratopogon) 

Forcipomyia swezeyana Tokunaga and Murachi 1959 

(Forcipomyia) 

Forcipomyia tenuichela Dow and Wirth 1972 (Forcipomyia) 

Forcipomyia tenuiforceps Macfie 1939 (Forcipomyia) 

Forcipomyia texanus Long 1902 (Ceratopogon) 

Forcipomyia titillans Winnertz 1852 (Ceratopogon) 

Forcipomyia townesi Wirth 1952 (Forcipomyia) 

Forcipomyia unica Bystrak and Wirth 1978 (Forcipomyia) 

Forcipomyia usingeri Wirth and Spinelli 1993 (Forcipomyia) 

Forcipomyia varipennis Wirth and Williams 1957 (Forcipomyia) 

Forcipomyia veroensis Wirth and Messersmith 1971 (Forcipomyia) 

Forcipomyia weemsi Wirth and Spinelli 1992 (Forcipomyia) 

Forcipomyia wheeleri Long 1902 (Ceratopogon) 

Forcipomyia wirthi Saunders 1956 (Forcipomyia) 

Heteromyia Say 1825 

Pachyleptus Walker 1856 Syn. 

Heteromyia fasciata Say 1825 (Heteromyia) 

Ceratopogon festiva Loew 1861 Syn. 

Heteromyia prattii Coquillett 1902 (Heteromyia) 

Jenkinshelea Macfie 1934 

Jenkinsia Kieffer 1913 Homo. 

Jenkinshelea albarius Coquillett 1895 (Ceratopogon) 

Johannsenomyia aequalis Malloch 1915 Syn. 

Jenkinshelea blantoni Grogan and Wirth 1977 (Jenkinshelea) 

Jenkinshelea magnipennis Johannsen 1908 (Johannseniella) 

Jenkinshelea stonei Grogan and Wirth 1977 (Jenkinshelea) 

Johannsenomyia Malloch 1915 

Dicrohelea Kieffer 1917 Syn. 

Johannsenomyia annulicornis Malloch 1918 (Johannsenomyia) 

Johannsenomyia argentatus Loew 1861 (Ceratopogon) 

Johannsenomyia halteralis Malloch 1915 Syn. 

Leptoconops Skuse 1889 

Tersesthes Townsend 1893 Syn. 

Mycterotypus Noe 1905 Syn. 

Holoconops Kieffer 1918 Syn. 

Schizoconops Kieffer 1918 Syn. 

Styloconops Kieffer 1921 Syn. 

Microconops Kieffer 1921 Syn. 

Brachyconops Wirth 1973 Syn. 

Megaconops Wirth 1973 Syn. 

Proleptoconops Clastrier 1974 Syn. 

Leptoconops andersoni Clastrier and Wirth 1978 (Leptoconops)

Leptoconops arnaudi Clastrier and Wirth 1978 (Leptoconops) 

Leptoconops asilomar Clastrier and Wirth 1978 (Leptoconops) 

Leptoconops atchleyi Clastrier and Wirth 1978 (Leptoconops) 

Leptoconops becquaerti Kieffer 1925 (Holoconops) 

Leptoconops bequaerti Wirth 1965 Emend. 

Leptoconops belkini Wirth 1973 (Leptoconops) 

Leptoconops californiensis Wirth 1973 (Leptoconops) 

Leptoconops catawbae Boesel 1948 (Holoconops) 

Leptoconops floridensis Wirth 1951 (Leptoconops) 

Leptoconops foulki Clastrier and Wirth 1978 (Leptoconops) 

Leptoconops freeborni Wirth 1952 (Leptoconops) 

Leptoconops kerteszi Kieffer 1908 (Leptoconops) 

Leptoconops americana Carter 1921 Syn. 

Leptoconops knowltoni Clastrier and Wirth 1978 (Leptoconops) 

Leptoconops linleyi Wirth 1973 (Leptoconops) 

Leptoconops melanderi Wirth 1973 (Leptoconops) 

Leptoconops mohavensis Wirth 1973 (Leptoconops) 

Leptoconops reesi Clastrier and Wirth 1978 (Leptoconops) 

Leptoconops sublettei Clastrier and Wirth 1978 (Leptoconops) 

Leptoconops torrens Townsend 1893 (Tersesthes) 

Leptoconops carteri Hoffman 1926 Syn. 

Leptoconops werneri Wirth 1973 (Leptoconops) 

Leptoconops whitseli Clastrier and Wirth 1978 (Leptoconops) 

Macropeza Meigen 1818 

Macroptilum Becker 1903 Syn. 

Haasiella Kieffer 1913 Syn. 


Macropeza blantoni Wirth and Ratanaworabhan 1972 (Macropeza) 

Mallochohelea Wirth 1962 

Mallochohelea albibasis Malloch 1915 (Johannsenomyia) 

Mallochohelea albihalter Wirth 1962 (Mallochohelea) 

Mallochohelea atripes Wirth 1962 (Mallochohelea) 

Mallochohelea caudellii Coquillett 1905 (Ceratopogon) 

Mallochohelea flavidula Malloch 1914 (Johannseniella) 

Mallochohelea pullata Wirth 1952 (Johannsenomyia) 

Mallochohelea smithi Lewis 1956 (Johannsenomyia) 

Mallochohelea spinipes wirth 1962 (Mallochohelea) 

Mallochohelea sybleae Wirth 1952 (Johannsenomyia) 

Mallochohelea texensis Wirth 1962 (Mallochohelea) 

Mallochohelea variegata Wirth 1962 (Mallochohelea) 

Monohelea Kieffer 1917 

Monohelea andersoni Wirth and Grogan 1981 (Monohelea) 

Monohelea bifurcata Wirth and Williams 1964 (Monohelea) 

Monohelea floridensis Wirth and Williams 1964 (Monohelea) 

Monohelea hirsuta Wirth and Grogan 1981 (Monohelea) 

Monohelea jamnbacki Wirth and Williams 1964 (Monohelea) 

Monohelea knighti Wirth and Williams 1964 (Monohelea) 

Monohelea lanei Wirth 1953 (Monohelea) 

Monohelea macfiei Wirth 1953 (Monohelea) 

Monohelea maculipennis Coquillett 1905 (Ceratopogon) 

Monohelea magnitheca Wirth and Grogan 1981 (Monohelea) 

Monohelea obscura Wirth and Williams 1964 (Monohelea) 

Monohelea ornata Wirth 1953 (Monohelea) 

Monohelea texana Wirth 1953 (Monohelea) 

Monohelea wirthi Khalaf 1969 (Monohelea) 

Neurobezzia Wirth and Ratanaworabhan 1972 

Neurobezzia granulosa Wirth 1952 (Bezzia) 

Neurohelea Kieffer 1925 

Neurohelea macroneura Malloch 1915 (Johannsenomyia) 

Neurohelea nigra Wirth 1952 (Neurohelea) 

Nilobezzia Kieffer 1921 

Parrotia Kieffer 1924 Syn. 

Sphaerobezzia Zilahi-Sebess 1940 Syn. 

Nilobezzia brevicornis Wirth 1952 (Sphaeromias) 

Nilobezzia mallochi Wirth 1962 (Nilobezzia) 

Nilobezzia minor Wirth 1952 (Sphaeromias) 

Nilobezzia schwarzii Coquillett 1901 (Ceratopogon) 

Ceratopogon setipes Coquillett 1905 Syn. 

Pachyhelea Wirth 1959 

Pachyhelea pachymerus Williston 1900 (Ceratopogon) 

Ceratopogon magnus Coquillett 1905 Syn. 

Palpomyia Meigen 1818 

Apogon Rondani 1856 Homo. 

Alasion Rondani 1857 Syn. 

Miopalpomyia Pierce 1966 Syn. 

Neopalpomyia Pierce 1966 Syn. 

Parapalpomyia Pierce 1966 Syn. 

Gluhhovia Remm 1976 Syn. 

Palpomyia aldrichi Malloch 1915 (Heteromyia) 

Palpomyia altispina Grogan and Wirth 1975 (Palpomyia) 

Palpomyia armatipes Wirth 1952 (Palpomyia) 

Palpomyia aspina Grogan and Wirth 1979 (Palpomyia) 

Palpomyia basalis Walker 1848 (Ceratopogon) 

Ceratopogon trivialis Loew 1861 Syn. 

Heteromyia tenuicornis Malloch 1915 Syn. 

Palpomyia belkini Grogan and Wirth 1979 (Palpomyia) 

Palpomyia bottimeri Grogan and Wirth 1979 (Palpomyia) 

Palpomyia canadensis Grogan and Wirth 1979 (Palpomyia) 

Palpomyia cressoni Malloch 1915 (Heteromyia) 

Palpomyia downesi Grogan and Wirth 1979 (Palpomyia) 

Palpomyia flaviceps Johannsen 1908 (Johannseniella) 

Heteromyia hirta Malloch 1915 Syn. 

Heteromyia opacithorax Malloch 1915 Syn. 

Palpomyia hastata Grogan and Wirth 1975 (Palpomyia) 

Palpomyia jamnbacki Grogan and Wirth 1979 (Palpomyia) 

Palpomyia jonesi Grogan and Wirth 1975 (Palpomyia) 

Palpomyia kernensis Wirth 1952 (Palpomyia) 

Palpomyia lineatus Meigen 1818 (Ceratopogon) 

Ceratopogon slossonae Coquillett 1905 Syn. 

Palpomyia illinoisensis Malloch 1914 Syn. 

Palpomyia pruinescens Thomsen 1935 Syn. 

Palpomyia linsleyi Wirth 1952 (Palpomia) 

Palpomyia nigripes Meigen 1830 (Ceratopogon) 

Palpomyia novitibalis Grogan and Wirth 1975 (Palpomyia) 

Palpomyia occidentalis Grogan and Wirth 1979 (Palpomyia) 

Palpomyia plebeiella Grogan and Wirth 1975 (Palpomyia) 

Palpomyia plebeius Loew 1861 (Ceratopogon) 

Palpomyia praeustus Loew 1869 (Ceratopogon) 

Palpomyia pseudorufa Grogan and Wirth 1975 (Palpomyia) 

Palpomyia rubiginosa Grogan and Wirth 1975 (Palpomyia) 

Palpomyia rufus Loew 1861 (Ceratopogon) 

Palpomyia scalpellifera Grogan and Wirth 1975 (Palpomyia) 

Palpomyia stonei Wirth 1951 (Palpomyia) 

Palpomyia subasper Coquillett 1901 (Ceratopogon) 

Palpomyia essigi Wirth 1952 Syn. 

Palpomyia tibialis Meigen 1818 (Ceratopogon) 

Palpomyia trifasciata Wirth 1952 (Palpomyia) 

Palpomyia walteri Grogan and Wirth 1979 (Palpomyia) 

Palpomyia weemsi Grogan and Wirth 1979 (Palpomyia) 

Parabezzia Malloch 1915 


103

104 

Parabezzia alexanderi Wirth 1965 (Parabezzia) 

Parabezzia atchleyi Grogan and Wirth 1977 (Parabezzia) 

Parabezzia bystraki Grogan and Wirth 1977 (Parabezzia) 

Parabezzia downesi Wirth 1965 (Parabezzia) 

Parabezzia eupetiolata Grogan and Wirth 1977 (Parabezzia) 

Parabezzia horvathi Grogan and Wirth 1977 (Parabezzia) 

Parabezzia huberti Grogan and Wirth 1977 (Parabezzia) 

Parabezzia inermis Coquillett 1902 (Ceratopogon) 

Parabezzia neunguis Grogan and Wirth 1977 (Parabezzia) 

Parabezzia petiolata Malloch 1915 (Parabezzia) 

Stilobezzia uncinata Johannsen 1943 Syn. 

Parabezzia texensis Grogan and Wirth 1977 (Parabezzia) 

Parabezzia unguis Wirth 1965 (Parabezzia) 

Parabezzia williamsi Wirth 1965 (Parabezzia) 

Paradasyhelea Macfie 1940 



Paradasyhelea olympiae Wirth and Blanton 1969 (Paradasyhelea) 

Paryphoconus Enderlein 1912 

Paryphoconus sonorensis Wirth and Ratanaworabhan 1972 

(Paryphoconus) 

Phaenobezzia Haeselbarth 1965 

Phaenobezzia fulvithorax Malloch 1915 (Probezzia) 

Phaenobezzia opacus Loew 1861 (Ceratopogon) 

Ceratopogon pachymerus Williston 1900 Syn. 

Probezzia incerta Malloch 1915 Syn. 

Phaenobezzia sabroskyi Wirth and Grogan 1982 (Phaenobezzia) 

Probezzia Kieffer 1906 

Dicrobezzia Kieffer 1919 Syn. 

Probezzia albitibia Wirth 1971 (Probezzia) 

Probezzia albiventris Loew 1861 (Ceratopogon) 

Probezzia atriventris Wirth 1951 (Probezzia) 

Probezzia bottimeri Wirth 1971 (Probezzia) 

Probezzia concinnus Meigen 1818 (Ceratopogon) 

Probezzia fairchildi Wirth 1994 (Probezzia) 

Probezzia flavoniger Coquillett 1905 (Ceratopogon) 

Probezzia fuscipennis Wirth 1971 (Probezzia) 

Probezzia glicki Wirth 1994 (Probezzia) 

Probezzia infuscata Malloch 1915 (Probezzia) 

Probezzia jamnbacki Wirth 1971 (Probezzia) 

Probezzia ludoviciana Wirth 1951 (Probezzia) 

Probezzia meadi Wirth 1994 (Probezzia) 

Probezzia nigra Wirth 1971 (Probezzia) 

Probezzia pallida Malloch 1914 (Probezzia) 

Probezzia rosewalli Wirth 1951 (Probezzia) 

Probezzia sabroskyi Wirth 1951 (Probezzia) 

Probezzia seminigra Panzer 1798 (Tipula) 

Ceratopogon venustus Meigen 1818 Syn. 

Probezzia smithii Coquillett 1901 (Ceratopogon) 

Ceratopogon mundus Coquillett 1900 Unav. 

Probezzia unica Johannsen 1934 (Lasiobezzia) 

Probezzia weemsi Wirth 1994 (Probezzia) 

Probezzia williamsi Wirth 1971 (Probezzia) 

Probezzia xanthogaster Kieffer 1917 (Bezzia) 

Ceratopogon elegans Coquillett 1901 Homo. 

Rhynchohelea Wirth and Blanton 1970 

Rhynchohelea monilicornis Wirth and Blanton 1970 

(Rhynchohelea) 

Schizohelea Kieffer 1917 

Kiefferomyia Mayer 1937 Syn. 

Schizohelea leucopeza Meigen 1804 (Ceratopgon) 

Ceratopogon albitarsis Wiedemann 1817 Syn. 

Ceratopogon copiosa Winnertz 1852 Syn. 

Ceratopogon politus Coquillett 1901 Syn. 

Bezzia stecki Kieffer 1915 Syn. 

Kiefferomyia gorana Mayer 1937 Syn. 

Serromyia Meigen 1818 

Prionomyia Stephens 1829 Syn. 

Ceratolophus Kieffer 1899 Homo. 

Johannseniella Williston 1907 Syn. 

Ceratolophana Strand 1928 Syn. 

Serromyia barberi Wirth 1952 (Serromyia) 

Serromyia borealis Borkent 1990 (Serromyia) 

Serromyia crassifemorata Malloch 1914 (Serromyia) 

Serromyia ledicola Kieffer 1925 (Serromyia) 

Serromyia macronyx Goetghebuer 1933 Syn. 

Serromyia europaea Clastrier 1963 Syn. 

Serromyia nudicolis Borkent 1990 (Serromyia) 

Serromyia sierrensis Borkent 1990 (Serromyia) 

Serromyia vockerothi Borkent 1990 (Serromyia) 

Sphaeromias Curtis 1829 

Xylocrypta Kieffer 1899 Syn. 

Sphaeromias bifidus Wirth and Grogan 1979 (Sphaeromias) 

Sphaeromias longipennis Loew 1861 (Ceratopogon) 

Stenoxenus Coquillett 1899 

Stenoxenus johnsoni Coquillett 1899 (Stenoxenus) 

Stilobezzia Kieffer 1911 

Hartomyia Malloch 1915 Syn. 

Acanthohelea Kieffer 1917 Syn. 

Eukraiohelea Ingram and Macfie 1921 Syn. 

Neostilobezzia Goetghebuer 1934 Syn. 

Debenhamia Wirth and Grogan 1988 Syn. 

Stilobezzia amnigena Macfie 1935 (Eukraiohelea) 

Stilobezzia antennalis Coquillett 1901 (Ceratopogon) 

Stilobezzia beckae Wirth 1953 (Stilobezzia) 

Stilobezzia bicolor Lane 1947 (Stilobezzia) 

Stilobezzia bulla Thomson 1935 (Stilobezzia) 

Stilobezzia coquilletti Kieffer 1917 (Stilobezzia) 

Ceratopogon pictus Coquillett 1905 Homo. 

Stilobezzia diversus Coquillett 1901 (Ceratopogon) 

Stilobezzia elegantula Johannsen 1907 (Bezzia) 

Stilobezzia maculitibia Lane and Forattini 1956 Syn. 

Stilobezzia fuscula Wirth 1952 (Stilobezzia) 

Stilobezzia glauca Macfie 1939 (Stilobezzia)

Stilobezzia kiefferi Lane 1947 (Stilobezzia) 

Stilobezzia punctipes Wirth 1953 Syn. 

Stilobezzia lutea Malloch 1918 (Hartomyia) 

Stilobezzia mallochi Hoffman 1924 Syn. 

Stilobezzia navaiae Wirth and Grogan 1981 (Stilobezzia) 

Stilobezzia pallidiventris Malloch 1915 (Hartomyia) 

Stilobezzia pruinosa Wirth 1952 (Stilobezzia) 

Stilobezzia rabelloi Lane 1947 (Stilobezzia) 

Stilobezzia stonei Wirth 1953 (Stilobezzia) 

Stilobezzia sybleae Wirth 1953 (Stilobezzia) 

Stilobezzia thomsenae Wirth 1953 (Stilobezzia) 

Stilobezzia viridis Coquillett 1901 (Ceratopogon) 

Washingtonhelea Wirth and Grogan 1988 

Washingtonhelea frommeri Wirth and Grogan 1988 

(Washingtonhelea) 

Zeuxidiplosis Kieffer 1904 

Zeuxidiplosis giardi Kieffer 1896 (Diplosis) 

Diplosis giardiana Kieffer 1897 Syn. 

CHAMAEMYIIDAE 

Chamaemyia Meigen 1803 

Ochtiphila Fallen 1823 Syn. 

Chamaemyia aridella Fallen 1823 (Ochtiphila) 

Chamaemyia flavipalpis Haliday 1838 (Ochtiphila) 

Ochtiphila maritima Zetterstedt 1846 Syn. 

Chamaemyia fumicosta Malloch 1940 (Chamaemyia) 

Chamaemyia geniculata Zetterstedt 1838 (Ochtiphila) 

Chamaemyia herbarum Robineau-Desvoidy 1830 (Estelia) 

Chamaemyia juncorum Fallen 1823 (Ochtiphila) 

Ochtiphila polystigma Meigen 1830 Syn. 

Cremifania Czerny 1904 

Cremifania nearctica McAlpine 1963 (Cremifania) 

Cremifania nigrocellulata Czerny 1904 (Cremifania) 

Leucopis Meigen 1830 

Neoleucopis Malloch 1921 Syn. 

Leucopomyia Malloch 1921 Syn. 

Leucopina Malloch 1927 Syn. 

Xenoleucopis Malloch 1933 Syn. 


Leucopis americana Malloch 1921 (Leucopis) 

Leucopis ancilla McAlpine 1971 (Leucopis) 

Leucopis argenticollis Zetterstedt 1848 (Leucopis) 

Leucopis orbitalis Malloch 1921 Syn. 

Leucopis atratula Ratzeburg 1844 (Musca) 

Leucopis atrifacies Aldrich 1925 (Leucopis) 

Leucopis bellula Williston 1889 (Leucopis) 

Leucopis bivittata Malloch 1940 (Leucopis) 

Leucopis conciliata McAlpine and Tanasijtschuk 1972 (Leucopis) 

Leucopis flavicornis Aldrich 1914 (Leucopis) 

Leucopis hennigrata McAlpine 1978 (Leucopis) 

Leucopis maculata Thompson 1910 (Leucopis) 

Leucopis major Malloch 1921 (Leucopis) 

Leucopis militia McAlpine 1971 (Leucopis) 

Leucopis minor Malloch 1921 (Leucopis) 

Leucopis nigraluna McAlpine 1971 (Leucopis) 

Leucopis obscura Haliday 1833 (Leucopis) 

Leucopis hyalipennis Zetterstedt 1845 Syn. 

Leucopis ocellaris Malloch 1940 (Leucopis) 

Leucopis parallela Malloch 1921 (Leucopis) 

Leucopis pemphigae Malloch 1921 (Leucopis) 

Leucopis pinicola Malloch 1921 (Leucopis) 

Leucopis piniperda Malloch 1921 (Leucopis) 

Leucopis pulvinariae Malloch 1921 (Leucopis) 

Leucopis simplex Loew 1869 (Leucopis) 

Leucopis phylloxerae Riley 1883 Syn. 

Leucopis tapiae Blanchard 1964 (Leucopis) 

Leucopis verticalis Malloch 1940 (Leucopis) 

Lipoleucopis Meijere 1928 

Lipoleucopis praecox Meijere 1928 (Lipoleucopis) 

Paraleucopis Malloch 1913 

Paraleucopis boydensis Steyskal 1971 (Paraleucopis) 

Paraleucopis corvina Malloch 1913 (Paraleucopis) 

Plunomia Curran 1934 

Plunomia elegans Curran 1934 (Plunomia) 

Plunomia flavicornis Malloch 1940 Syn. 

Plunomia obtusa Malloch 1940 (Plunomia) 

Plunomia tibialis Malloch 1940 (Plunomia) 

Plunomia transversa Malloch 1940 (Plunomia) 

Pseudodinia Coquillett 1902 

Pseudodinia angelica Barber 1985 (Pseudodinia) 

Pseudodinia angustata Barber 1985 (Pseudodinia) 

Pseudodinia antennalis Malloch 1940 (Pseudodinia) 

Pseudodinia hamata Barber 1985 (Pseudodinia) 

Pseudodinia latiphallis Barber 1985 (Pseudodinia) 

Pseudodinia melanitida Barber 1985 (Pseudodinia) 

Pseudodinia nitida Melander 1913 Homo. 

Pseudodinia nigritarsis Barber 1985 (Pseudodinia) 

Pseudodinia nitens Melander and Spuler 1917 (Piophila) 

Piophila nitida Wulp 1867 Homo. 

Pseudodinia occidentalis Barber 1985 (Pseudodinia) 

Pseudodinia polita Malloch 1915 (Pseudodinia) 

Pseudodinia pruinosa Melander 1913 (Pseudodinia) 

Pseudodinia slussi Barber 1985 (Pseudodinia) 

Pseudodinia varipes Coquillett 1902 (Pseudodinia) 

Pseudoleucopis Malloch 1925 

Pseudoleucopis benefica Malloch 1930 (Pseudoleucopis) 

Toropamecia Cogan 1978 

Toropamecia punctata Coquillett 1902 (Acrometopia) 

Tigonometopus reticulata Johnson 1913 Syn. 

DIPTERA: CERATOPOGONIDAE, CHAMAEMYIIDAE 

105

106 

CHAOBORIDAE 

Chaoborus Lichtenstein 1800 

Corethra Meigen 1803 Syn. 

Culicites Heyden 1862 Syn. 

Sayomyia Coquillett 1903 Syn. 

Corethrinda Giles 1904 Syn. 

Schadonophasma Dyar and Shannon 1924 Syn. 

Neochaoborus Edwards 1930 Syn. 

Edwardsops Lane 1942 Syn. 

Peusomyia Saether 1970 Syn. 

Chaoborus albatus Johnson 1921 (Chaoborus) 

Chaoborus americana Johannsen 1903 (Corethra) 

Sayomyia hudsoni Felt 1904 Syn. 

Chaoborus sonjai Saether 1970 Unav. 

Chaoborus typica Saether 1970 Unav. 

Chaoborus astictopus Dyar and Shannon 1924 (Chaoborus) 

Chaoborus lacustris Freeborn 1926 Syn. 

Chaoborus cooki Saether 1970 (Chaoborus) 

Chaoborus crystallina De Geer 1776 (Tipula) 

Tipula pilicornis Fabricius 1787 Syn. 

Tipula hafniensis Gmelin 1790 Syn. 

Tipula plumicornis Fabricius 1794 Syn. 

Chaoborus antisepticus Lichtenstein 1800 Syn. 

Corethra lateralis Meigen 1804 Syn. 

Corethra fusca Staeger 1839 Syn. 

Corethra pilipes Gimmerthal 1845 Syn. 

Proboscistoma pellucens Saccardo 1864 Syn. 

Chaoborus borealis Cook 1956 Syn. 

Chaoborus festivus Dyar and Shannon 1924 (Chaoborus) 

Chaoborus annulatus Cook 1956 Syn. 

Chaoborus flavicans Meigen 1830 (Corethra) 

Corethra albipes Johannsen 1903 Syn. 

Sayomyia rotundifolia Felt 1904 Syn. 

Chaoborus eluthera Dyar and Shannon 1924 Syn. 

Chaoborus alpinus Peus 1938 Syn. 

Chaoborus maculipes Stone 1965 (Chaoborus) 

Chaoborus maculipes Stone 1965 (Chaoborus) 

Chaoborus punctipennis Say 1823 (Corethra) 

Corethra appendiculata Herrick 1884 Syn. 

Chaoborus trivittata Loew 1862 (Corethra) 

Sayomyia knabi Dyar 1905 Syn. 

Chaoborus brunskilli Saether 1970 Syn. 

Eucorethra Underwood 1903 

Pelorempis Johannsen 1903 Syn. 

Eucorethra underwoodi Underwood 1903 (Eucorethra) 

Pelorempis americana Johannsen 1903 Syn. 

Mochlonyx Loew 1844 

Mochlonyx cinctipes Coquillett 1903 (Corethra) 

Corethra obscura Dyar and Shannon 1924 Syn. 

Mochlonyx fuliginosa Felt 1905 (Corethra) 

Mochlonyx martinii Edwards 1930 Syn. 

Mochlonyx simplex Sasa and Takahasi 1948 Syn. 

Mochlonyx velutina Ruthe 1831 (Corethra) 

Tipula culiciformis De Geer 1776 Homo. 

Corethra rufa Zetterstedt 1838 Syn. 

Mochlonyx effoetus Walker 1856 Syn. 

Corethra karnerensis Felt 1904 Syn. 

Corethra lintneri Felt 1904 Syn. 


DIPTERA: CHAOBORIDAE, CHIRONOMIDAE 

Mochlonyx vernalis Mondchadsky 1953 Syn. 

CHIRONOMIDAE 

Abiskomyia Edwards 1937 

Abiskomyia virgo Edwards 1937 (Abiskomyia) 

Ablabesmyia Johannsen 1905 

Karelia Roback 1971 Syn. 

Asayia Roback 1985 Syn. 

Ablabesmyia alaskensis Roback 1971 (Ablabesmyia) 

Ablabesmyia annulatus Say 1823 (Tanypus) 

Ablabesmyia aspera Roback 1959 (Pentaneura) 

Pentaneura gera Roback 1959 Syn. 

Ablabesmyia aurea Johannsen 1907 (Ablabesmyia) 

Ablabesmyia cinctipes Johannsen 1946 (Pentaneura) 

Ablabesmyia hauberi Beck and Beck 1966 (Ablabesmyia) 

Ablabesmyia idei Walley 1925 (Tanypus) 

Ablabesmyia illinoensis Malloch 1915 (Tanypus) 

Ablabesmyia janta Roback 1959 (Pentaneura) 

Ablabesmyia johannseni Roback 1959 (Pentaneura) 

Ablabesmyia mallochi Walley 1925 (Tanypus) 

Pentaneura auriensis Roback 1957 Syn. 

Pentaneura aequifasciata Dendy and Sublette 1959 Syn. 

Ablabesmyia ornata Beck and Beck 1966 Syn. 

Ablabesmyia tarella Roback 1971 Syn. 

Ablabesmyia monilis Linnaeus 1758 (Tipula) 

Tanypus basalis Walley 1925 Syn. 

Ablabesmyia americana Fittkau 1962 Syn. 

Ablabesmyia nigra Roback 1971 Unav. 

Ablabesmyia parajanta Roback 1971 (Ablabesmyia) 

Ablabesmyia peleensis Walley 1926 (Tanypus) 

Ablabesmyia philosphagnos Beck and Beck 1966 (Ablabesmyia) 

Ablabesmyia pulchripennis Lundbeck 1898 (Tanypus) 

Tanypus prudens Walley 1925 Syn. 

Ablabesmyia rasha Roback 1971 (Ablabesmyia) 

Ablabesmyia rhamphe Sublette 1964 (Ablabesmyia) 

Ablabesmyia simpsoni Roback 1985 (Ablabesmyia) 

Acamptocladius Brundin 1956 

Phycoidella Saether 1971 Syn. 

Acamptocladius dentolatens Saether 1971 (Phycoidella) 

Acricotopus Kieffer 1921 

Acricotopus lucens Zetterstedt 1850 (Chironomus) 

Chironomus baueri Goetghebuer 1955 Syn. 

Acricotopus nitidellus Malloch 1915 (Orthocladius) 

Spaniotoma senex Johannsen 1937 Syn. 

Alotanypus Roback 1971 

Alotanypus aris Roback 1971 (Alotanypus) 

Alotanypus venustus Coquillett 1902 (Tanypus) 

Antillocladius Saether 1981 

Antillocladius arcuatus Saether 1982 (Antillocladius)

Antillocladius pluspilalus Saether 1982 (Antillocladius) 

Apedilum Townes 1945 

Apedilum elachistus Townes 1945 (Apedilum) 

Apedilum subcinctum Townes 1945 (Apedilum) 

Paralauterborniella alamedensis Sublette 1960 Syn. 

Apometriocnemus Saether 1985 

Apometriocnemus beringensis Cranston and Oliver 1988 

(Apometriocnemus) 

Apometriocnemus fontinalis Saether 1985 (Apometriocnemus) 

Apsectrotanypus Fittkau 1962 

Apsectrotanypus algens Coquillett 1902 (Tanypus) 

Apsectrotanypus florens Johannsen 1908 (Tanypus) 

Apsectrotanypus johnsoni Coquillett 1901 (Tanypus) 

Arctodiamesa Makarchenko 1982 

Arctodiamesa appendiculata Lundström 1915 (Diamesa) 

Arctopelopia Fittkau 1962 

Arctopelopia cana Roback 1971 (Arctopelopia) 

Arctopelopia melanosoma Goetghebuer 1933 (Ablabesmyia) 

Asheum Sublette and Sublette 1983 

Pedionomus Sublette 1964 Homo. 

Asheum beckae Sublette 1964 (Pedionomus) 

Axarus Roback 1980 

Anceus Roback 1963 Homo. 

Axarus dorneri Malloch 1915 (Chironomus) 

Axarus festivus Say 1823 (Chironomus) 

Chironomus lineatus Say 1823 Syn. 

Chironomus lineola Wiedemann 1828 Emend. 

Chironomus lasiomerus Walker 1848 Syn. 

Axarus rogersi Beck and Beck 1958 (Xenochironomus) 

Axarus scopula Townes 1945 (Xenochironomus) 

Axarus taenionotus Say 1829 (Chironomus) 

Baeoctenus Saether 1971 

Baeoctenus bicolor Saether 1976 (Baeoctenus) 

Beardius Reiss and Sublette 1985 

Beardius truncatus Reiss and Sublette 1985 (Beardius) 

Beckidia Saether 1979 

Beckiella Saether 1977 Homo. 

Beckidia tethys Townes 1945 (Harnischia) 

Bethbilbeckia Fittkau and Murray 1988 


Bethbilbeckia floridensis Fittkau and Murray 1988 (Bethbilbeckia) 

Boreochlus Edwards 1938 

Boreochlus gracilistylus Brundin 1966 (Boreochlus) 

Boreochlus persimilis Johannsen 1926 (Trichotanypus) 

Boreochlus longisetus Brundin 1966 Syn. 

Boreochlus sinuaticornis Brundin 1966 (Boreochlus) 

Boreoheptagyia Brundin 1966 

Neopodonomus Chandhuri and Ghosh 1981 Syn. 

Boreoheptagyia lurida Garrett 1925 (Diamesa) 

Brillia Kieffer 1913 

Brillia flavifrons Johannsen 1905 (Metriocnemus) 

Brillia laculata Oliver and Roussel 1983 (Brillia) 

Brillia longifurca Kieffer 1921 (Brillia) 

Brillia parva Johannsen 1934 (Brillia) 

Brillia retifinis Saether 1969 (Brillia) 

Brillia sera Roback 1957 (Brillia) 

Brundiniella Roback 1978 

Brundinia Roback 1978 Homo. 

Brundiniella eumorpha Sublette 1964 (Anatopynia) 

Bryophaenocladius Thienemann 1934 

Clinocladius Sublette 1970 Syn. 

Platycladius Sublette 1970 Syn. 

Bryophaenocladius aestivus Brundin 1947 (Eudactylocladius) 

Bryophaenocladius astis Roback 1957 (Hydrobaenus) 

Hydrobaenus traenis Roback 1957 Syn. 

Bryophaenocladius digitatus Saether 1973 (Bryophaenocladius) 

Bryophaenocladius flavoscutellatus Malloch 1915 (Orthocladius) 

Bryophaenocladius fumosinus Curran 1930 (Camptocladius) 

Bryophaenocladius furcatus Kieffer 1916 (Dactylocladius) 

Bryophaenocladius impectinus Saether 1976 (Bryophaenocladius) 

Bryophaenocladius laticaudus Saether 1973 (Bryophaenocladius) 

Bryophaenocladius pleuralis Malloch 1915 (Orthocladius) 

Bryophaenocladius psilacrus Saether 1982 (Bryophaenocladius) 

Bryophaenocladius subparallelus Malloch 1915 (Orthocladius) 

Camptocladius Wulp 1874 

Camptocladius stercorarius De Geer 1776 (Tipula) 

Tipula byssina Schrank 1803 Syn. 

Cantopelopia Roback 1971 

Cantopelopia gesta Roback 1971 (Cantopelopia) 

Cardiocladius Kieffer 1912 

Cardiocladius albiplumus Saether 1969 (Cardiocladius) 

Cardiocladius fulva Johannsen 1908 (Thalassomya) 

Thalassomya fusca Johannsen 1903 Unav. 

Cardiocladius obscura Johannsen 1903 (Thalassomya) 

Cardiocladius platypus Coquillett 1902 (Orthocladius) 

Chaetocladius Kieffer 1911 

Chaetocladius adsimilis Goetghebuer 1933 (Orthocladius) 

Chaetocladius grandilobus Brundin 1956 (Chaetocladius) 

Chaetocladius holmgreni Jacobson 1898 (Chironomus) 

Chironomus festivus Holmgren 1869 Homo. 

Chaetocladius ligni Cranston and Oliver 1988 (Chaetocladius) 

Chaetocladius oliveri Saether 1969 (Chaetocladius) 

Chaetocladius perennis Meigen 1830 (Chironomus) 

Chaetocladius stamfordi Johannsen 1947 (Hydrobaenus) 

Chasmatonotus Loew 1864 

DIPTERA: CHIRONOMIDAE 

107

108 



Chasmatonotus atripes Rempel 1937 (Chasmatonotus) 

Chasmatonotus bicolor Rempel 1937 (Chasmatonotus) 

Chasmatonotus bimaculatus Osten Sacken 1877 (Chasmatonotus) 

Chasmatonotus fascipennis Coquillett 1905 (Chasmatonotus) 

Chasmatonotus hyalinus Coquillett 1905 (Chasmatonotus) 

Chasmatonotus maculipennis Rempel 1937 (Chasmatonotus) 

Chasmatonotus unimaculatus Loew 1864 (Chasmatonotus) 

Chasmatonotus univittatus Coquillett 1900 (Chasmatonotus) 

Chernovskiia Saether 1977 

Chernovskiia amphitrite Townes 1945 (Harnischia) 

Chernovskiia orbicus Townes 1945 (Harnischia) 

Chironomus Meigen 1803 

Tendipes Meigen 1800 Unav. 

Camptochironomus Kieffer 1918 Syn. 

Holtedahlia Kieffer 1922 Syn. 

Chaetolabis Townes 1945 Syn. 

Lobochironomus Ryser, Wülker, Scholl 1985 Syn. 

Chironomus anonymus Williston 1896 (Chironomus) 

Chironomus anthracinus Zetterstedt 1860 (Chironomus) 

Chironomus meridionalis Johannsen 1908 Syn. 

Chironomus rempelii Thienemann 1941 Syn. 

Chironomus athalassicus Cannings 1975 (Chironomus) 

Chironomus atrella Townes 1945 (Tendipes) 

Chironomus atritibia Malloch 1934 (Chironomus) 

Chironomus atroviridis Townes 1945 (Tendipes) 

Chironomus attenuatus Walker 1848 (Chironomus) 

Chironomus bifimbriatus Kieffer 1922 (Chironomus) 

Chironomus bimaculus Walker 1848 (Chironomus) 

Chironomus biseta Townes 1945 (Tendipes) 

Chironomus borealis Curtis 1835 (Chironomus) 

Chironomus brevipalpis Kieffer 1926 (Holtedahlia) 

Chironomus crassicaudatus Malloch 1915 (Chironomus) 

Chironomus cucini Webb 1969 (Chironomus) 

Chironomus decorus Johannsen 1905 (Chironomus) 

Chironomus similis Johannsen 1905 Syn. 

Chironomus cayugae Johannsen 1913 Syn. 

Chironomus distinguendus Kieffer 1917 Syn. 

Chironomus decumbens Malloch 1934 (Chironomus) 

Chironomus excavatus Kieffer 1917 (Chironomus) 

Chironomus excisus Kieffer 1917 (Chironomus) 

Chironomus frommeri Atchley and Martin 1971 (Chironomus) 

Chironomus grandivalva Shilova 1957 (Tendipes) 

Chironomus halophilus Packard 1873 (Chironomus) 

Chironomus harti Malloch 1915 (Chironomus) 

Chironomus hyperboreus Staeger 1845 (Chironomus) 

Chironomus lasiopus Walker 1848 (Chironomus) 

Chironomus longipes Staeger 1839 (Chironomus) 

Chironomus major Wülker and Butler 1983 (Chironomus) 

Chironomus maturus Johannsen 1908 (Chironomus) 

Chironomus melanderi Kieffer 1917 (Chironomus) 

Chironomus mendax Storå 1936 (Chironomus) 

Chironomus milleri Townes 1945 (Tendipes) 

Chironomus nigritibia Walker 1848 (Chironomus) 

Chironomus ochreatus Townes 1945 (Tendipes) 

Chironomus pallidivittatus Malloch 1915 (Chironomus) 

Chironomus pilicornis Fabricius 1794 (Tipula) 

Chironomus polaris Kirby 1824 Syn. 

Chironomus plumosa Linnaeus 1758 (Tipula) 

Tipula annularis De Geer 1776 Syn. 

Chironomus cristatus Fabricius 1805 Syn. 

Chironomus ferrugineovittatus Zetterstedt 1850 Syn. 

Chironomus imperator Walley 1926 Syn. 

Chironomus prasinus Meigen 1804 (Chironomus) 

Chironomus prior Butler 1982 (Chironomus) 

Chironomus pungens Townes 1945 (Tendipes) 

Chironomus redeuns Walker 1856 (Chironomus) 

Chironomus riparius Meigen 1804 (Chironomus) 

Chironomus albistria Walker 1848 Syn. 

Chironomus serus Malloch 1915 Syn. 

Chironomus militaris Johannsen 1937 Syn. 

Chironomus sanctipauli Sublette 1966 (Chironomus) 

Chironomus conformis Malloch 1923 Homo. 

Chironomus staegeri Lundbeck 1898 (Chironomus) 

Chironomus fasciventris Malloch 1915 Syn. 

Chironomus stigmaterus Say 1823 (Chironomus) 

Chironomus glaucurus Wiedemann 1828 Syn. 

Chironomus tardus Butler 1982 (Chironomus) 

Chironomus tentans Fabricius 1805 (Chironomus) 

Chironomus trichomerus Walker 1848 (Chironomus) 

Chironomus tuberculatus Townes 1945 (Tendipes) 

Chironomus tuxis Curran 1930 (Chironomus) 

Chironomus utahensis Malloch 1915 (Chironomus) 

Chironomus vancouveri Michailova and Fischer 1986 (Chironomus) 

Chironomus vockerothi Rasmussen 1984 (Chironomus) 

Chironomus whitseli Sublette and Sublette 1974 (Chironomus) 

Cladopelma Kieffer 1921 

Cryptocladopelma Lenz 1941 Unav. 

Cladopelma amachaerus Townes 1945 (Harnischia) 

Cladopelma boydi Beck 1962 (Chironomus) 

Cladopelma collator Townes 1945 (Harnischia) 

Cladopelma edwardsi Kruseman 1933 (Tendipes) 

Cladopelma galeator Townes 1945 (Harnischia) 

Cladopelma spectabilis Townes 1945 (Harnischia) 

Cladopelma viridula Linnaeus 1767 (Tipula) 

Cladotanytarsus Kieffer 1922 

Lenziella Kieffer 1922 Syn. 

Cladotanytarsus aeiparthenus Bilyj 1989 (Cladotanytarsus) 

Cladotanytarsus atridorsum Kieffer 1924 (Cladotanytarsus) 

Cladotanytarsus cruscula Saether 1971 (Lenziella) 

Cladotanytarsus daviesi Bilyj 1989 (Cladotanytarsus) 

Cladotanytarsus elaensis Bilyj 1989 (Cladotanytarsus) 

Cladotanytarsus fusiformis Bilyj 1989 (Cladotanytarsus) 

Cladotanytarsus mancus Walker 1856 (Chironomus) 

Cladotanytarsus muricatus Bilyj 1989 (Cladotanytarsus) 

Cladotanytarsus pinnaticornis Bilyj 1989 (Cladotanytarsus) 

Cladotanytarsus tribelus Bilyj 1989 (Cladotanytarsus) 

Cladotanytarsus viridiventris Malloch 1915 (Tanytarsus) 

Clinotanypus Kieffer 1913 

Aponteus Roback 1971 Syn. 

Clinotanypus aureus Roback 1971 (Clinotanypus) 

Clinotanypus pinguis Loew 1861 (Tanypus) 

Tanypus flavicinctus Loew 1861 Syn. 

Tanypus thoracicus Loew 1866 Syn. 

Procladius caliginosus Johannsen 1905 Syn. 

Clinotanypus planus Roback 1971 (Clinotanypus) 

Clinotanypus sabensis Roback 1971 (Clinotanypus) 

Clinotanypus wirthi Roback 1971 (Clinotanypus) 

Clunio Haliday 1855 

Clunio californiensis Hashimoto 1974 (Clunio) 

Clunio marshalli Stone and Wirth 1947 (Clunio) 

Coelotanypus Kieffer 1913 

Coelotanypus atus Roback 1971 (Coelotanypus)

Coelotanypus concinnus Coquillett 1895 (Tanypus) 

Coelotanypus flavus Kieffer 1923 Syn. 

Coelotanypus naelis Roback 1963 (Coelotanypus) 

Coelotanypus scapularis Loew 1866 (Tanypus) 

Coelotanypus tricolor Loew 1861 (Tanypus) 

Compteromesa Saether 1981 

Compteromesa oconeensis Saether 1981 (Compteromesa) 

Compterosmittia Saether 1981 

Compterosmittia clavigera Saether 1982 (Compterosmittia) 

Compterosmittia nerius Curran 1930 (Camptocladius) 

Conchapelopia Fittkau 1957 

Mesopelopia Roback 1971 Syn. 

Conchapelopia aleta Roback 1971 (Conchapelopia) 

Conchapelopia bruna Roback 1971 (Conchapelopia) 

Conchapelopia currani Walley 1925 (Tanypus) 

Conchapelopia fasciata Beck and Beck 1966 (Conchapelopia) 

Conchapelopia dusena Roback 1971 Syn. 

Conchapelopia mera Roback 1971 (Conchapelopia) 

Conchapelopia pallens Coquillett 1902 (Tanypus) 

Pentaneura goniodes Sublette 1964 Syn. 

Conchapelopia trifida Roback 1971 Unav. 

Conchapelopia paramelanops Roback 1971 (Conchapelopia) 

Conchapelopia rurika Roback 1957 (Pentaneura) 

Conchapelopia telema Roback 1971 (Conchapelopia) 

Conchapelopia varna Roback 1981 (Conchapelopia) 

Constempellina Brundin 1947 

Constempellina brevicosta Edwards 1937 (Tanytarsus) 

Corynocera Zetterstedt 1838 

Corynocera ambigua Zetterstedt 1838 (Corynocera) 

Corynoneura Winnertz 1846 

Corynoneura arctica Kieffer 1923 (Corynoneura) 

Corynoneura celeripes Winnertz 1852 (Corynoneura) 

Corynoneura atra Winnertz 1852 Syn. 

Corynoneura diara Roback 1957 (Corynoneura) 

Corynoneura fittkaui Schlee 1968 (Corynoneura) 

Corynoneura lacustris Edwards 1924 (Corynoneura) 

Corynoneura lobata Edwards 1924 (Corynoneura) 

Corynoneura minuscula Brundin 1949 (Corynoneura) 

Corynoneura oxfordana Boesel and Winner 1980 (Corynoneura) 

Corynoneura scutellata Winnertz 1846 (Corynoneura) 

Corynoneura taris Roback 1957 (Corynoneura) 

Cricotopus Wulp 1874 

Isocladius Kieffer 1909 Syn. 

Nostococladius Ashe and Murray 1980 Syn. 

Pseudocricotopus Nishida 1987 Syn. 

Cricotopus abana Curran 1929 (Cricotopus) 

Cricotopus absurdus Johannsen 1905 (Orthocladius) 

Cricotopus annulator Goetghebuer 1927 (Cricotopus) 

Cricotopus irwini Sublette and Sublette 1971 Syn. 

Cricotopus baptistensis Boesel 1983 (Cricotopus) 

Cricotopus belkini Dendy and Sublette 1959 (Cricotopus) 

Cricotopus beringensis Oliver and Dillon 1988 (Cricotopus) 

Cricotopus bicinctus Meigen 1818 (Chironomus) 

Cricotopus bifurcatus Cranston and Oliver 1988 (Cricotopus) 


Cricotopus brunnicans Walley 1928 (Cricotopus) 

Cricotopus coronatus Hirvenoja 1973 (Cricotopus) 

Cricotopus cumulatus Hirvenoja 1973 (Cricotopus) 

Cricotopus curtis Hirvenoja 1973 (Cricotopus) 

Cricotopus cylindraceus Kieffer 1908 (Trichocladius) 

Cricotopus diversus Boesel 1983 (Cricotopus) 

Cricotopus edurus Sublette and Sublette 1971 (Cricotopus) 

Cricotopus elegans Johannsen 1943 (Cricotopus) 

Cricotopus ephippium Zetterstedt 1838 (Chironomus) 

Ceratopogon humeralis Zetterstedt 1838 Syn. 

Cricotopus festivellus Kieffer 1906 (Chironomus) 

Chironomus festivus Meigen 1830 Homo. 

Cricotopus flavibasis Malloch 1915 (Cricotopus) 

Cricotopus flavipes Johannsen 1942 (Cricotopus) 

Cricotopus flavocinctus Kieffer 1924 (Trichocladius) 

Cricotopus fugax Johannsen 1905 (Orthocladius) 

Cricotopus furtivus Sublette and Sublette 1971 (Cricotopus) 

Cricotopus fuscatus Wirth 1957 (Cricotopus) 

Cricotopus fuscus Kieffer 1909 (Trichocladius) 

Cricotopus gelidus Kieffer 1922 (Trichocladius) 

Cricotopus geminatus Say 1823 (Chironomus) 

Cricotopus globistylus Roback 1957 (Cricotopus) 

Cricotopus infuscatus Malloch 1915 (Orthocladius) 

Cricotopus aratus Roback 1957 Syn. 

Cricotopus ceris Roback 1957 Syn. 

Cricotopus intersectus Staeger 1839 (Chironomus) 

Cricotopus junus Roback 1957 (Cricotopus) 

Cricotopus laetus Hirvenoja 1973 (Cricotopus) 

Cricotopus laricomalis Edwards 1932 (Cricotopus) 

Cricotopus lestralis Edwards 1924 (Trichocladius) 

Cricotopus luciae LeSage 1980 (Cricotopus) 

Cricotopus mackenziensis Oliver 1977 (Cricotopus) 

Cricotopus macraei Saether 1971 (Cricotopus) 

Cricotopus flannagani Saether 1971 Syn. 

Cricotopus myriophylli Oliver 1984 (Cricotopus) 

Cricotopus nostocicola Wirth 1957 (Cricotopus) 

Cricotopus obscurifuscus Sublette and Sublette 1971 (Cricotopus) 

Cricotopus oceanicus Packard 1869 (Chironomus) 

Cricotopus olivetus Boesel 1983 (Cricotopus) 

Cricotopus ornatus Meigen 1818 (Chironomus) 

Cricotopus parafuscatus Sublette and Sublette 1971 (Cricotopus) 

Cricotopus patens Hirvenoja 1973 (Cricotopus) 

Cricotopus pedatus Sublette 1967 (Cricotopus) 

Cricotopus pilitarsis Zetterstedt 1850 (Chironomus) 

Cricotopus pilosellus Brundin 1956 (Cricotopus) 

Cricotopus politus Coquillett 1902 (Orthocladius) 

Cricotopus pulchripes Verrall 1912 (Cricotopus) 

Cricotopus slossonae Malloch 1915 (Cricotopus) 

Cricotopus subfuscus Sublette and Sublette 1971 (Cricotopus) 

Cricotopus sylvestris Fabricius 1794 (Tipula) 

Cricotopus tibialis Meigen 1804 (Chironomus) 

Chironomus basalis Staeger 1845 Syn. 

Chironomus pavidus Holmgren 1869 Syn. 

Cricotopus polaris Kieffer 1926 Syn. 

Trichocladius bacilliger Kieffer 1926 Syn. 

Cricotopus tremula Linnaeus 1758 (Tipula) 

Cricotopus triannulatus Macquart 1826 (Chironomus) 

Cricotopus exilis Johannsen 1905 Syn. 

Cricotopus tricinctus Meigen 1818 (Chironomus) 

Cricotopus lebetis Sublette 1964 Syn. 

Cricotopus trifascia Edwards 1929 (Cricotopus) 

Cricotopus ithacanensis Sublette 1967 Syn. 

Cricotopus trifasciatus Meigen 1813 (Chironomus) 

Cricotopus remus Sublette 1964 Syn. 

Cricotopus trilobus Oliver and Dillon 1988 (Cricotopus) 

Cricotopus tristis Hirvenoja 1973 (Cricotopus) 

Cricotopus varipes Coquillett 1902 (Cricotopus) 

Cricotopus vierriensis Goetghebuer 1935 (Cricotopus) 


109

110 

Cryptochironomus Kieffer 1918 



Cryptochironomus argus Roback 1957 (Cryptochironomus) 

Cryptochironomus blarina Townes 1945 (Cryptochironomus) 

Cryptochironomus conus Mason 1986 (Cryptochironomus) 

Cryptochironomus curryi Mason 1986 (Cryptochironomus) 

Cryptochironomus digitatus Malloch 1915 (Chironomus) 

Cryptochironomus eminentia Mason 1986 (Cryptochironomus) 

Cryptochironomus fulvus Johannsen 1905 (Chironomus) 

Chironomus abbreviatus Malloch 1915 Homo. 

Chironomus parvilamellatus Malloch 1915 Syn. 

Chironomus mallochi Kieffer 1919 Syn. 

Cryptochironomus parafulvus Beck and Beck 1964 (Chironomus) 

Cryptochironomus ponderosus Sublette 1964 (Chironomus) 

Cryptochironomus psittacinus Meigen 1830 (Chironomus) 

Chironomus stylifera Johannsen 1908 Syn. 

Chironomus obtusilobus Malloch 1923 Syn. 

Chironomus farinalis Walley 1926 Syn. 

Cryptochironomus ramus Mason 1986 (Cryptochironomus) 

Cryptochironomus scimitarus Townes 1945 (Cryptochironomus) 

Cryptochironomus sorex Townes 1945 (Cryptochironomus) 

Cryptotendipes Lenz 1941 

Cryptotendipes ariel Sublette 1960 (Tendipes) 

Cryptotendipes causuaria Townes 1945 (Harnischia) 

Cryptotendipes darbyi Sublette 1960 (Tendipes) 

Cryptotendipes emorsa Townes 1945 (Harnischia) 

Cryptotendipes pilicuspis Saether 1977 (Cryptotendipes) 

Cryptotendipes pseudotener Goetghebuer 1922 (Cryptochironomus) 

Cyphomella Saether 1977 

Cyphomella argentea Townes 1945 (Harnischia) 

Cyphomella cornea Saether 1977 (Cyphomella) 

Cyphomella gibbera Saether 1977 (Cyphomella) 

Cyphomella griseus Malloch 1915 (Chironomus) 

Demeijerea Kruseman 1933 

Demeijerea abruptus Townes 1945 (Glyptotendipes) 

Demeijerea atrimanus Coquillett 1902 (Chironomus) 

Demeijerea brachialis Coquillett 1901 (Chironomus) 

Demeijerea obreptus Townes 1945 (Glyptotendipes) 

Demicryptochironomus Lenz 1941 

Schadinia Lipina 1939 Nomob. 

Irmakia Reiss 1988 Syn. 

Demicryptochironomus cuneata Townes 1945 (Harnischia) 

Demicryptochironomus fastigata Townes 1945 (Harnischia) 

Denopelopia Roback and Rutter 1988 

Denopelopia atria Roback and Rutter 1988 (Denopelopia) 

Derotanypus Roback 1971 

Merotanypus Roback 1971 Syn. 

Derotanypus aclines Sublette 1964 (Anatopynia) 

Derotanypus alaskensis Malloch 1919 (Tanypus) 

Anatopynia centralis Malloch 1934 Syn. 

Diamesa Meigen 1835 

Eutanypus Coquillett 1899 Syn. 

Adiamesa Kieffer 1906 Syn. 

Psilodiasmea Kieffer 1918 Syn. 

Diamesa aberrata Lundbeck 1898 (Diamesa) 

Diamesa alpina Tokunaga 1936 (Diamesa) 

Diamesa amplexivirilia Hansen 1976 (Diamesa) 

Diamesa ancysta Roback 1959 (Diamesa) 

Diamesa arcticus Boheman 1865 (Chironomus) 

Diamesa bertrami Edwards 1935 (Diamesa) 

Diamesa bohemani Goetghebuer 1932 (Diamesa) 

Diamesa cheimatophila Hansen 1976 (Diamesa) 

Diamesa chiobates Hansen 1976 (Diamesa) 

Diamesa chorea Lundbeck 1898 (Diamesa) 

Diamesa clavata Edwards 1933 (Diamesa) 

Diamesa colenae Hansen 1976 (Diamesa) 

Diamesa davisi Edwards 1933 (Diamesa) 

Diamesa garretti Sublette and Sublette 1965 (Diamesa) 

Diamesa borealis Garrett 1925 Homo. 

Diamesa geminata Kieffer 1926 (Diamesa) 

Diamesa biappendiculata Goetghebuer 1928 Syn. 

Diamesa furcata Edwards 1933 Syn. 

Diamesa gregsoni Edwards 1933 (Diamesa) 

Diamesa haydaki Hansen 1976 (Diamesa) 

Diamesa heteropus Coquillett 1905 (Tanypus) 

Adiamesa confusa Garrett 1925 Syn. 

Diamesa banana Garrett 1925 Syn. 

Diamesa onteona Roback 1957 Syn. 

Diamesa incallidus Walker 1856 (Chironomus) 

Diamesa fonticola Saether 1969 Syn. 

Diamesa insignipes Kieffer 1908 (Diamesa) 

Diamesa japonica Tokunaga 1936 (Diamesa) 

Diamesa leoniella Hansen 1976 Syn. 

Diamesa leona Roback 1957 (Diamesa) 

Diamesa caena Roback 1957 Syn. 

Diamesa pieta Roback 1957 Syn. 

Diamesa lindrothi Goetghebuer 1931 (Diamesa) 

Diamesa lupus Willassen 1985 (Diamesa) 

Diamesa mendotae Muttkowski 1915 (Diamesa) 

Diamesa nivicavernicola Hansen 1976 (Diamesa) 

Diamesa nivoriundus Fitch 1847 (Chironomus) 

Eutanypus borealis Coquillett 1899 Syn. 

Diamesa polaris Kieffer 1926 (Diamesa) 

Diamesa simplex Kieffer 1926 (Diamesa) 

Diamesa sommermani Hansen 1976 (Diamesa) 

Diamesa sonorae Willassen 1985 (Diamesa) 

Diamesa spinacies Saether 1969 (Diamesa) 

Diamesa subletti Makarchenko 1986 (Diamesa) 

Diamesa vockerothi Hansen 1976 (Diamesa) 

Dicrotendipes Kieffer 1913 

Limnochironomus Kieffer 1920 Syn. 

Dicrotendipes adnilus Epler 1987 (Dicrotendipes) 

Dicrotendipes aethiops Townes 1945 (Tendipes) 

Dicrotendipes botaurus Townes 1945 (Tendipes) 

Dicrotendipes californicus Johannsen 1905 (Chironomus) 

Dicrotendipes crypticus Epler 1987 (Dicrotendipes) 

Dicrotendipes fumidus Johannsen 1905 (Chironomus) 

Chironomus incognitus Malloch 1915 Syn. 

Dicrotendipes leucoscelis Townes 1945 (Tendipes) 

Dicrotendipes lobiger Kieffer 1921 (Limnochironomus) 

Dicrotendipes lobus Beck 1962 (Chironomus) 

Dicrotendipes lucifer Johannsen 1907 (Chironomus) 

Chironomus indistinctus Malloch 1915 Syn. 

Dicrotendipes modestus Say 1823 (Chironomus) 

Dicrotendipes neomodestus Malloch 1915 (Chironomus) 

Dicrotendipes nervosus Staeger 1839 (Chironomus) 

Chironomus brevitibialis Zetterstedt 1850 Syn. 

Dicrotendipes simpsoni Epler 1987 (Dicrotendipes) 

Dicrotendipes thanatogratus Epler 1987 (Dicrotendipes)

Dicrotendipes tritomus Kieffer 1916 (Chironomus) 

Chironomus incurvus Sublette 1964 Syn. 

Diplocladius Kieffer 1908 

Diplocladius cultriger Kieffer 1908 (Diplocladius) 

Diplocladius bilobatus Brundin 1956 Syn. 

Diplosmittia Saether 1981 

Diplosmittia carinata Saether 1985 (Diplosmittia) 

Djalmabatista Fittkau 1968 

Calotanypus Roback 1971 Syn. 

Djalmabatista pulcher Johannsen 1908 (Protenthes) 

Tanypus apicalis Walley 1926 Syn. 

Protenthes fasciger Curran 1930 Syn. 

Procladius maculatus Roback 1971 Syn. 

Doithrix Saether and Sublette 1983 

Doithrix barberi Saether and Sublette 1983 (Doithrix) 

Doithrix dillonae Cranston and Oliver 1988 (Doithrix) 

Doithrix ensifer Saether and Sublette 1983 (Doithrix) 

Doithrix hamiltoni Saether and Sublette 1983 (Doithrix) 

Doithrix parcivillosa Saether and Sublette 1983 (Doithrix) 

Doithrix villosa Saether and Sublette 1983 (Doithrix) 

Doncricotopus Saether 1981 

Doncricotopus bicaudatus Saether 1981 (Doncricotopus) 

Einfeldia Kieffer 1924 

Einfeldia austini Beck and Beck 1970 (Einfeldia) 

Einfeldia brunneipennis Johannsen 1905 (Chironomus) 

Einfeldia chelonia Townes 1956 (Tendipes) 

Einfeldia natchitocheae Sublette 1964 (Chironomus) 

Einfeldia paganus Meigen 1838 (Chironomus) 

Einfeldia synchrona Oliver 1971 Syn. 

Endochironomus Kieffer 1918 


Endochironomus nigricans Johannsen 1905 (Chironomus) 

Chironomus johnsoni Kieffer 1906 Unav. 

Endochironomus oldenbergi Goetghebuer 1932 (Endochironomus) 

Endochironomus subtendens Townes 1945 (Tanytarsus) 

Endotribelos Grodhaus 1987 

Endotribelos hesperius Sublette 1960 (Tendipes) 

Epoicocladius Zavrel 1924 

Epoicocladius flavens Malloch 1915 (Camptocladius) 

Camptocladius ephemerae Kieffer 1924 Syn. 

Eretmoptera Kellogg 1900 

Eretmoptera browni Kellogg 1900 (Eretmoptera) 

Eukiefferiella Thienemann 1926 

Adactylocladius Saether 1968 Syn. 

Eukiefferiella brevinervis Malloch 1915 (Orthocladius) 

Eukiefferiella claripennis Lundbeck 1898 (Chironomus) 

Spaniotoma hospita Edwards 1929 Syn. 

Eukiefferiella cyanea Thienemann 1936 (Eukiefferiella) 

Eukiefferiella devonica Edwards 1929 (Spaniotoma) 

Eukiefferiella unicalcar Saether 1969 (Adactylocladius) 

Euryhapsis Oliver 1981 

Euryhapsis annuliventris Malloch 1915 (Metriocnemus) 

Euryhapsis cilium Olivier 1981 (Euryhapsis) 

Euryhapsis illoba Oliver 1981 (Euryhapsis) 

Fittkauimyia Karunakaran 1969 

Parapelopia Roback 1971 Syn. 

Fittkauimyia serta Roback 1971 (Parapelopia) 

Georthocladius Strenzke 1941 

Atelopodella Saether 1982 Syn. 

Georthocladius curticornus Saether 1982 (Georthocladius) 

Georthocladius fimbriosus Saether and Sublette 1983 

(Georthocladius) 

Georthocladius platystylus Saether and Sublette 1983 


Georthocladius triquetrus Saether and Sublette 1983 


Georthocladius wirthi Saether and Sublette 1983 (Georthocladius) 

Gillotia Kieffer 1921 

Gillotia alboviridis Malloch 1915 (Chironomus) 

Glyptotendipes Kieffer 1913 

Glyptotendipes amplus Townes 1945 (Glyptotendipes) 

Glyptotendipes barbipes Staeger 1839 (Chironomus) 

Glyptotendipes dreisbachi Townes 1945 (Glyptotendipes) 

Glyptotendipes lobiferus Say 1823 (Chironomus) 

Chironomus caliginosus Johannsen 1905 Homo. 

Chironomus ithacanensis Johannsen 1908 Syn. 

Glyptotendipes americanus Kieffer 1917 Syn. 

Glyptotendipes meridionalis Dendy and Sublette 1959 

(Glyptotendipes) 

Glyptotendipes paripes Edwards 1929 (Chironomus) 

Glyptotendipes seminole Townes 1945 (Glyptotendipes) 

Glyptotendipes senilis Johannsen 1937 (Chironomus) 

Glyptotendipes testaceus Townes 1945 (Glyptotendipes) 

Glyptotendipes unacus Townes 1945 (Glyptotendipes) 

Goeldichironomus Fittkau 1965 

Stolimyia Fittkau 1968 Syn. 

Goeldichironomus amazonica Fittkau 1968 (Stolimyia) 

Goeldichironomus carus Townes 1945 (Tendipes) 

Goeldichironomus devineyae Beck 1961 (Chironomus) 

Goeldichironomus holoprasinus Goeldi 1905 (Chironomus) 

Chironomus fulvipilis Rempel 1939 Syn. 

Goeldichironomus pictus Reiss 1974 (Goeldichironomus) 

Guttipelopia Fittkau 1962 

Guttipelopia guttipennis Wulp 1874 (Tanypus) 

Tanypus multipunctatus Curran 1930 Syn. 

Guttipelopia currani Beck and Beck 1966 Syn. 

Guttipelopia rosenbergi Bilyj 1984 (Guttipelopia) 

Gymnometriocnemus Goetghebuer 1932 

Raphidocladius Saether 1983 Syn. 


111

112 

Gymnometriocnemus brumalis Edwards 1929 (Metriocnemus) 

Gymnometriocnemus acigus Saether 1983 Syn. 

Gymnometriocnemus subnudus Edwards 1929 (Metriocnemus) 

Gymnometriocnemus marionensis Saether 1969 Syn. 

Halocladius Hirvenoja 1973 

Halocladius variabilis Staeger 1839 (Chironomus) 

Halocladius vitripennis Meigen 1818 (Chironomus) 

Harnischia Kieffer 1921 

Harnischia curtilamellatus Malloch 1915 (Chironomus) 

Harnischia incidata Townes 1945 (Harnischia) 

Hayesomyia Murray and Fittkau 1985 

Hayesomyia senatus Walley 1925 (Tanypus) 

Chironomus unicolor Walker 1848 Syn. ? 

Pentaneura chrysos Sublette 1964 Syn. 

Heleniella Gowin 1943 

Heleniella curtistila Saether 1969 (Heleniella) 

Heleniella hirta Saether 1969 (Heleniella) 

Heleniella parva Saether 1985 (Heleniella) 

Helopelopia Roback 1971 

Helopelopia cornuticaudatus Walley 1925 (Tanypus) 

Conchapelopia gigas Beck and Beck 1966 Syn. 

Helopelopia pilicaudatus Walley 1925 (Tanypus) 

Heterotanytarsus Spärck 1923 

Heterotanytarsus nudalus Saether 1975 (Heterotanytaruss) 

Heterotanytarsus perennis Saether 1975 (Heterotanytarsus) 

Heterotrissocladius Spärck 1923 



Heterotrissocladius boltoni Saether 1993 (Heterotrissocladius) 

Heterotrissocladius changi Saether 1975 (Heterotrissocladius) 

Heterotrissocladius cooki Saether 1975 (Heterotrissocladius) 

Heterotrissocladius hirtapex Saether 1975 (Heterotrissocladius) 

Heterotrissocladius latilaminus Saether 1975 (Heterotrissocladius) 

Heterotrissocladius maari Brundin 1949 (Heterotrissocladius) 

Heterotrissocladius maeaeri Oliver et al. 1990 Emend. 

Heterotrissocladius marcidus Walker 1856 (Chironomus) 

Heterotrissocladius oliveri Saether 1975 (Heterotrissocladius) 

Heterotrissocladius subpilosus Kieffer 1911 (Dactylocladius) 

Hudsonimyia Roback 1979 

Hudsonimyia karelena Roback 1979 (Hudsonimyia) 

Hudsonimyia parrishi Caldwell and Soponis 1982 (Hudsonimyia) 

Hydrobaenus Fries 1830 

Hydrobaenus calvescens Saether 1976 (Hydrobaenus) 

Hydrobaenus conformis Holmgren 1869 (Chironomus) 

Orthocladius natvigi Goetghebuer 1933 Syn. 

Orthocladius natvigi Goetghebuer 1933 Syn. 

Hydrobaenus labradorensis Saether 1976 Syn. 

Hydrobaenus domus Sublette 1966 (Orthocladius) 

Orthocladius nivoriundus Johannsen 1934 Homo. 

Orthocladius johannseni Sublette 1967 Syn. 

Trissocladius hamiltoni Saether 1969 Syn. 

Hydrobaenus fusistylus Goetghebuer 1933 (Orthocladius) 

Hydrobaenus octomerus Saether 1976 Unav. 

Hydrobaenus hudsoni Saether 1976 (Hydrobaenus) 

Hydrobaenus laticaudus Saether 1976 (Hydrobaenus) 

Hydrobaenus martini Saether 1976 (Hydrobaenus) 

Hydrobaenus pilipes Malloch 1915 (Orthocladius) 

Hydrobaenus pilipodex Saether 1976 (Hydrobaenus) 

Hydrobaenus scapulapilosus Saether 1976 (Hydrobaenus) 

Hydrobaenus spinnatis Saether 1976 (Hydrobaenus) 

Hydrobaenus travisi Saether 1989 (Hydrobaenus) 

Hydrobaenus tumidistylus Saether 1976 (Hydrobaenus) 

Hydrobaenus virgo Saether 1976 (Hydrobaenus) 

Hyporhygma Reiss 1982 

Hyporhygma quadripunctatus Malloch 1915 (Chironomus) 

Kiefferulus Goetghebuer 1922 

Wirthiella Sublette 1960 Syn. 

Kiefferulus dux Johannsen 1905 (Chironomus) 

Kiefferulus modocensis Sublette 1960 (Tendipes) 

Kloosia Kruseman 1933 

Oschia Saether 1983 Syn. 

Kloosia dorsenna Saether 1983 (Oschia) 

Krenopelopia Fittkau 1962 

Krenopelopia hudsoni Roback 1983 (Krenopelopia) 

Krenopelopia narda Roback 1971 (Krenopelopia) 

Krenosmittia Thienemann and Krüger 1939 

Krenosmittia boreoalpina Goetghebuer 1944 (Smittia) 

Krenosmittia camptophelps Edwards 1929 (Spaniotoma) 

Krenosmittia halvorseni Cranston and Saether 1986 (Rheosmittia) 

Labrundinia Fittkau 1962 

Labrundinia becki Roback 1971 (Labrundinia) 

Labrundinia johannseni Beck and Beck 1966 (Labrundinia) 

Labrundinia maculata Roback 1971 (Labrundinia) 

Labrundinia neopilosella Beck and Beck 1966 (Labrundinia) 

Labrundinia pilosellus Loew 1866 (Tanypus) 

Labrundinia floridana Beck and Beck 1966 Syn. 

Labrundinia virescens Beck and Beck 1966 (Labrundinia) 

Lappodiamesa Serra-Tosio 1968 

Lappodiamesa boltoni Saether and Willassen 1988 (Lappodiamesa) 

Larsia Fittkau 1962 

Larsia berneri Beck and Beck 1966 (Larsia) 

Larsia canadensis Bilyj 1984 (Larsia) 

Larsia decoloratus Malloch 1915 (Tanypus) 

Larsia lurida Beck and Beck 1966 Syn. 

Larsia indistincta Beck and Beck 1966 Syn. 

Larsia lyra Sublette 1964 (Pentaneura) 

Larsia marginellus Malloch 1915 (Tanypus) 

Larsia planensis Johannsen 1946 (Pentaneura) 

Larsia sequoiaensis Sublette 1964 (Pentaneura) 

Lasiodiamesa Kieffer 1924 

Lasiodiamesa arietinus Coquillett 1908 (Tanypus) 

Lasiodiamesa brusti Saether 1969 (Lasiodiamesa)

Lasiodiamesa rawsoni Brundin 1966 (Lasiodiamesa) 

Lasiodiamesa sphagnicola Kieffer 1925 (Isoplastus) 

Lasiodiamesa tenebrosus Coquillett 1905 (Tanypus) 

Linacerus piloala Garrett 1925 Syn. 

Lasiodiamesa nearctica Brundin 1966 Syn. 

Lauterborniella Bause 1913 

Lauterborniella agrayloides Kieffer 1911 (Tanytarsus) 

Limnophyes Eaton 1875 

Limnophyes anderseni Saether 1990 (Limnophyes) 

Limnophyes asquamatus Andersen 1937 (Limnophyes) 

Limnophyes septentrionalis Goetghebuer 1940 Syn. 

Limnophyes smolandicus Brundin 1947 Syn. 

Limnophyes vernalis Brundin 1947 Syn. 

Limnophyes franzi Goetghebuer 1949 Syn. 

Limnophyes hamiltoni Saether 1969 Syn. 

Limnophyes atomarius Zetterstedt 1850 (Chironomus) 

Limnophyes brachytomus Kieffer 1922 (Camptocladius) 

Limnophyes borealis Goetghebuer 1933 Syn. 

Limnophyes squamatus Andersen 1937 Syn. 

Limnophyes spatulosus Saether 1975 Syn. 

Limnophyes carolinensis Saether 1975 (Limnophyes) 

Limnophyes coloradensis Saether 1975 (Limnophyes) 

Limnophyes doughmani Saether 1990 (Limnophyes) 

Limnophyes eltoni Edwards 1922 (Camptocladius) 

Limnophyes fumosus Johannsen 1905 (Camptocladius) 

Limnophyes cristatissimus Saether 1975 Syn. 

Limnophyes hastulatus Saether 1975 (Limnophyes) 

Limnophyes margaretae Saether 1975 (Limnophyes) 

Limnophyes minimus Meigen 1818 (Chironomus) 

Limnophyes pusillus Eaton 1875 Syn. 

Limnophyes groenlandiensis Andersen 1937 Syn. 

Limnophyes groenlandicus Andersen 1937 Missp. 

Limnophyes interruptus Goetghebuer 1938 Syn. 

Limnophyes bipunctatus Goetghebuer 1941 Syn. 

Limnophyes praecox Goetghebuer 1944 Syn. 

Limnophyes immucronatus Saether 1969 Syn. 

Limnophyes hudsoni Saether 1975 Syn. 

Limnophyes natalensis Kieffer 1914 (Camptocladius) 

Camptocladius palmensis Santos-Abreu 1918 Syn. 

Limnophyes brevis Goetghebuer 1934 Syn. 

Limnophyes bequaerti Goetghebuer 1939 Syn. 

Limnophyes spinosa Freeman 1953 Syn. 

Limnophyes nudiradius Saether 1975 Syn. 

Limnophyes ninae Saether 1975 (Limnophyes) 

Limnophyes pilicistulus Saether 1975 (Limnophyes) 

Limnophyes prolongatus Kieffer 1921 (Camptocladius) 

Limnophyes pumilio Holmgren 1869 (Chironomus) 

Camptocladius globifer Lundström 1915 Syn. 

Camptocladius crescens Kieffer 1933 Syn. 

Limnophyes folliculatus Saether 1975 Syn. 

Limnophyes recisus Saether 1975 (Limnophyes) 

Lipurometriocnemus Saether 1981 

Lipurometriocnemus vixlobatus Saether 1982 

(Lipurometriocnemus) 

Lopescladius Oliveira 1967 

Cordiella Coffman and Roback 1984 Syn. 

Lopescladius hyporheicus Coffman and Roback 1984 

(Lopescladius) 

Lopescladius inermis Saether 1983 (Lopescladius) 

Lopescladius verruculosus Saether 1983 (Lopescladius) 


Macropelopia Thienemann 1916 

Macropelopia decedens Walker 1848 (Tanypus) 

Tanypus hirtipennis Loew 1866 Syn. 

Anatopynia dena Roback 1955 Syn. 

Anatopynia brunnea Roback 1965 Homo. 

Meropelopia Roback 1971 

Meropelopia americana Fittkau 1957 (Conchapelopia) 

Meropelopia flavifrons Johannsen 1905 (Ablabesmyia) 

Pentaneura alba Roback 1957 Homo. 

Pentaneura gesta Roback 1965 Syn. 

Arctopelopia fittkaui Beck and Beck 1966 Syn. 

Mesocricotopus Brundin 1956 

Mesocricotopus thienemanni Goetghebuer 1940 (Trichocladius) 

Mesosmittia Brundin 1956 

Mesosmittia acutistylus Saether 1985 (Mesosmittia) 

Mesosmittia lobiga Saether 1985 (Mesosmittia) 

Mesosmittia mina Saether 1985 (Mesosmittia) 

Mesosmittia patrihortae Saether 1985 (Mesosmittia) 

Mesosmittia prolixa Saether 1985 (Mesosmittia) 

Mesosmittia tora Saether 1985 (Mesosmittia) 

Metriocnemus Wulp 1874 

Metriocnemus abdominoflavatus Picado 1913 (Metriocnemus) 

Metriocnemus aequalis Johannsen 1934 (Metriocnemus) 

Metriocnemus albolineatus Meigen 1818 (Chironomus) 

Chironomus atratulus Zetterstedt 1850 Syn. 

Metriocnemus brusti Saether 1898 (Metriocnemus) 

Metriocnemus edwardsi Jones 1916 (Metriocnemus) 

Metriocnemus fuscipes Meigen 1818 (Chironomus) 

Metriocnemus knabi Coquillett 1904 (Metriocnemus) 

Metriocnemus longipennis Holmgren 1883 (Smittia) 

Metriocnemus obscuripes Holmgren 1869 (Chironomus) 

Chironomus eurynotus Holmgren 1883 Syn. 

Chironomus ripicola Holmgren 1883 Syn. 

Chironomus transgressus Holmgren 1883 Syn. 

Orthocladius hygropetricus Kieffer 1911 Syn. 

Metriocnemus cataractum Kieffer 1919 Syn. 

Metriocnemus longitarsus Goetghebuer 1921 Syn. 

Metriocnemus flavipilus Kieffer 1925 Syn. 

Metriocnemus grossus Kieffer 1925 Syn. 

Metriocnemus violaceus Kieffer 1935 Syn. 

Metriocnemus perfuscus Malloch 1934 (Metriocnemus) 

Metriocnemus picipes Meigen 1818 (Chironomus) 

Metriocnemus polaris Kieffer 1926 (Metriocnemus) 

Metriocnemus similis Kieffer 1922 (Metriocnemus) 

Metriocnemus tristellus Edwards 1929 (Metriocnemus) 

Metriocnemus ursinus Holmgren 1869 (Chironomus) 

Metriocnemus yaquina Cranston and Judd 1987 (Metriocnemus) 

Microchironomus Kieffer 1918 

Leptochironomus Pagast 1931 Syn. 

Microchironomus nigrovittatus Malloch 1915 (Chironomus) 

Micropsectra Kieffer 1909 

Lauterbornia Kieffer 1911 Syn. 

Lundstroemia Kieffer 1921 Syn. 

Oeklandia Kieffer 1922 Syn. 


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114 

Micropsectra apposita Walker 1856 (Chironomus) 

Micropsectra attenuata Reiss 1969 (Micropsectra) 

Micropsectra borealis Kieffer 1922 (Oeklandia) 

Micropsectra brundini Säwedal 1979 (Micropsectra) 

Micropsectra connexus Kieffer 1906 (Chironomus) 

Chironomus confinis Walker 1848 Homo. 

Chironomus brunneus Walker 1848 Syn. (?) 

Micropsectra deflectus Johannsen 1905 (Tanytarsus) 

Micropsectra dives Johannsen 1905 (Tanytarsus) 

Micropsectra geminata Oliver and Dillon 1994 (Micropsectra) 

Micropsectra groenlandica Andersen 1937 (Micropsectra) 

Micropsectra insignilobus Kieffer 1924 (Micropsectra) 

Micropsectra junci Meigen 1818 (Chironomus) 

Chironomus brunnipes Zetterstedt 1850 Syn. 

Chironomus gmundensis Egger 1863 Syn. 

Micropsectra lindrothi Goetghebuer 1931 (Micropsectra) 

Micropsectra logani Johannsen 1928 (Tanytarsus) 

Micropsectra nigripilus Johannsen 1905 (Tanytarsus) 

Micropsectra politus Malloch 1915 (Tanytarsus) 

Micropsectra recurvata Goetghebuer 1928 (Micropsectra) 

Micropsectra sedna Oliver 1976 (Lauterbornia) 

Micropsectra tori Säwedal 1981 (Micropsectra) 

Micropsectra xantha Roback 1955 (Calopsectra) 

Microtendipes Kieffer 1915 

Microtendipes anticus Walker 1848 (Chironomus) 

Microtendipes caducus Townes 1945 (Microtendipes) 

Chironomus pallidus Johannsen 1905 Homo. 

Microtendipes caelum Townes 1945 (Microtendipes) 

Microtendipes pedella De Geer 1776 (Tipula) 

Chironomus aberrans Johannsen 1905 Syn. 

Chironomus fascipes Coquillett 1908 Syn. 

Chironomus deflexa Walley 1926 Syn. 

Microtendipes stygius Townes 1945 Syn. 

Monodiamesa Kieffer 1922 

Monodiamesa bathyphila Kieffer 1918 (Prodiamesa) 

Monodiamesa depectinata Saether 1973 (Monodiamesa) 

Monodiamesa prolilobata Saether 1973 (Monodiamesa) 

Monodiamesa tuberculata Saether 1973 (Monodiamesa) 

Monopelopia Fittkau 1962 

Monopelopia boliekae Beck and Beck 1966 (Monopelopia) 

Monopelopia tenuicalcar Kieffer 1918 (Tanypus) 

Monopelopia tillandsia Beck and Beck 1966 (Monopelopia) 

Nanocladius Kieffer 1913 

Microcricotopus Thienemann and Harnisch 1932 Syn. 

Plecopteracoluthus Steffan 1965 Syn. 



Nanocladius alternantherae Dendy and Sublette 1959 (Nanocladius) 

Nanocladius anderseni Saether 1977 (Nanocladius) 

Nanocladius branchicolus Saether 1977 (Nanocladius) 

Nanocladius crassicornus Saether 1977 (Nanocladius) 

Nanocladius distinctus Malloch 1915 (Orthocladius) 

Orthocladius basalis Malloch 1915 Syn. 

Nanocladius downesi Steffan 1965 (Plecopteracoluthus) 

Nanocladius incomptus Saether 1977 (Nanocladius) 

Nanocladius mallochi Sublette 1970 (Nanocladius) 

Orthocladius bicolor Malloch 1915 Homo. 

Nanocladius minimus Saether 1977 (Nanocladius) 

Nanocladius rectinervis Kieffer 1911 (Cricotopus) 

Nanocladius sordens Johannsen 1905 (Orthocladius) 

Nanocladius spiniplenus Saether 1977 (Nanocladius) 

Natarsia Fittkau 1962 

Natarsia baltimoreus Macquart 1855 (Tanypus) 

Ablabesmyia fastuosa Johannsen 1905 Syn. 

Natarsia miripes Coquillett 1905 (Tanypus) 

Nilotanypus Kieffer 1923 

Nilotanypus americanus Beck and Beck 1966 (Nilotanypus) 

Nilotanypus fimbriatus Walker 1848 (Chironomus) 

Nilotanypus kansensis Roback 1986 (Nilotanypus) 

Nilothauma Kieffer 1921 

Nilothauma babiyi Rempel 1937 (Chironomus) 

Nilothauma bicornis Townes 1945 (Kribioxenus) 

Nilothauma mirabilis Townes 1945 (Kribioxenus) 

Nimbocera Reiss 1972 

Nimbocera pinderi Steiner and Hulbert 1982 (Nimbocera) 

Odontomesa Pagast 1947 

Odontomesa ferringtoni Saether 1985 (Odontomesa) 

Odontomesa fulva Kieffer 1919 (Prodiamesa) 

Odontomesa nearctica Saether 1985 Syn. 

Odontomesa lutosopra Garrett 1925 (Prodiamesa) 

Oliveridia Saether 1980 

Oliveria Saether 1976 Homo. 

Oliveridia hugginsi Ferrington and Saether 1987 (Oliveridia) 

Oliveridia tricornis Oliver 1976 (Trissocladius) 

Omisus Townes 1945 

Omisus pica Townes 1945 (Omisus) 

Oreadomyia Kevan and Cutten-Ali-Khan 1975 

Oreadomyia albertae Kevan and Cutten-Ali-Khan 1975 

(Oreadomyia) 

Orthocladius Wulp 1874 

Eudactylocladius Thienemann 1935 Syn. 

Euorthocladius Thienemann 1935 Syn. 

Rheorthocladius Thienemann 1935 Syn. 

Lapporthocladius Thienemann 1937 Syn. 

Pogonocladius Brundin 1956 Syn. 

Symposiocladius Cranston 1982 Syn. 

Orthocladius abiskoensis Edwards 1937 (Orthocladius) 

Orthocladius albidohalteralis Malloch 1915 (Orthocladius) 

Orthocladius annectens Saether 1969 (Orthocladius) 

Orthocladius anteilis Roback 1957 (Hydrobaenus) 

Orthocladius appersoni Soponis 1977 (Orthocladius) 

Orthocladius barbicornis Linnaeus 1767 (Tipula) 

Orthocladius bifasciatus Malloch 1918 (Orthocladius) 

Orthocladius carlatus Roback 1957 (Hydrobaenus) 

Orthocladius charensis Soponis 1977 (Orthocladius) 

Orthocladius clarkei Soponis 1977 (Orthocladius) 

Orthocladius clepsydrus Coquillett 1902 (Orthocladius) 

Orthocladius coffmani Soponis 1990 (Orthocladius) 

Orthocladius consobrinus Holmgren 1869 (Chironomus) 

Orthocladius cooki Soponis 1977 (Orthocladius) 

Orthocladius decoratus Holmgren 1869 (Chironomus) 

Orthocladius dentifer Brundin 1947 (Orthocladius)

Orthocladius difficilis Lundbeck 1898 (Chironomus) 

Orthocladius dorenus Roback 1957 (Hydrobaenus) 

Orthocladius currani Sublette 1967 Syn. 

Orthocladius dubitatus Johannsen 1942 (Orthocladius) 

Orthocladius ferringtoni Soponis 1983 (Orthocladius) 

Orthocladius frigidus Zetterstedt 1838 (Chironomus) 

Orthocladius gelidus Kieffer 1922 (Orthocladius) 

Orthocladius glacialis Kieffer 1926 (Camptocladius) 

Orthocladius groenlandensis Goetghebuer 1933 (Orthocladius) 

Orthocladius hazenensis Soponis 1977 (Orthocladius) 

Orthocladius hellenthali Soponis 1977 (Orthocladius) 

Orthocladius knabeni Goetghebuer 1933 (Orthocladius) 

Orthocladius knuthi Soponis 1977 (Orthocladius) 

Orthocladius lapponicus Goetghebuer 1940 (Orthocladius) 

Orthocladius lignicola Kieffer 1915 (Orthocladius) 

Orthocladius tryoni Soponis 1977 Syn. 

Orthocladius mallochi Kieffer 1919 (Orthocladius) 

Orthocladius lacteipennis Malloch 1915 Homo. 

Orthocladius manitobensis Saether 1969 (Orthocladius) 

Orthocladius minutus Zetterstedt 1850 (Chironomus) 

Orthocladius mixtus Holmgren 1869 (Chironomus) 

Orthocladius nanseni Kieffer 1926 (Orthocladius) 

Orthocladius nigritus Malloch 1915 (Orthocladius) 

Orthocladius oblidens Walker 1856 (Chironomus) 

Orthocladius lenzi Kieffer 1924 Syn. 

Orthocladius obumbratus Johannsen 1905 (Orthocladius) 

Orthocladius rhyacobius Kieffer 1911 Syn. 

Orthocladius rhyacophilus Kieffer 1911 Syn. 

Orthocladius dispar Goetghebuer 1942 Syn. 

Orthocladius excavatus Brundin 1947 Syn. 

Hydrobaenus paradorenus Roback 1957 Syn. 

Orthocladius oliveri Soponis 1977 (Orthocladius) 

Orthocladius rivicola Kieffer 1911 (Orthocladius) 

Orthocladius rivulorum Kieffer 1909 (Orthocladius) 

Orthocladius robacki Soponis 1977 (Orthocladius) 

Orthocladius roussellae Soponis 1990 (Orthocladius) 

Orthocladius rubicundus Meigen 1818 (Chironomus) 

Orthocladius saxicola Kieffer 1911 Syn. 

Orthocladius breviseta Saether 1969 Homo. 

Orthocladius curtiseta Saether 1973 Syn. 

Orthocladius saxosus Tokunaga 1939 (Spaniotoma) 

Orthocladius smolandicus Brundin 1947 (Orthocladius) 

Orthocladius spitzbergensis Kieffer 1919 (Dactylocladius) 

Orthocladius subletti Soponis 1977 (Orthocladius) 

Orthocladius thienemanni Kieffer 1906 (Orthocladius) 

Orthocladius trigonolabis Edwards 1924 (Orthocladius) 

Orthocladius wiensi Saether 1969 (Orthocladius) 

Pagastia Oliver 1959 

Hesperodiamesa Sublette 1967 Syn. 

Pagastia orthogonia Oliver 1959 (Pagastia) 

Pagastia partica Roback 1957 (Syndiamesa) 

Syndiamesa artisia Roback 1957 Syn. 

Pagastia sequax Garrett 1925 (Prodiamesa) 

Pagastiella Brundin 1949 

Pagastiella orophilus Edwards 1929 (Chironomus) 

Pagastiella ostansa Webb 1969 (Paralauterborniella) 

Paralauterborniella ostensa Boerger et al. 1982 Missp. 

Paraboreochlus Thienemann 1939 

Paraboreochlus stahli Coffman 1988 (Paraboreochlus) 

Parachaetocladius Wülker 1959 

Habrobaenus Saether 1977 Syn. 


Parachaetocladius abnobaeus Wülker 1959 (Chaetocladius) 

Habrobaenus hudsoni Saether 1977 Syn. 

Parachaetocladius abnobeus Saether and Sublette 1983 Missp. 

Parachaetocladius hirtipectus Saether 1969 (Parachaetocladius) 

Parachaetocladius imberbus Saether and Sublette 1983 

(Parachaetocladius) 

Parachironomus Lenz 1921 

Parachironomus abortivus Malloch 1915 (Chironomus) 

Parachironomus alatus Beck 1962 (Chironomus) 

Parachironomus carinatus Townes 1945 (Harnischia) 

Parachironomus chaetoala Sublette 1960 (Tendipes) 

Parachironomus directus Dendy and Sublette 1959 (Tendipes) 

Parachironomus elodeae Townes 1945 (Harnischia) 

Parachironomus forceps Townes 1945 (Harnischia) 

Parachironomus frequens Johannsen 1905 (Chironomus) 

Parachironomus hirtalatus Beck and Beck 1964 (Chironomus) 

Parachironomus monochromus Wulp 1874 (Chironomus) 

Chironomus unicolor Wulp 1858 Homo. 

Parachironomus pectinatellae Dendy and Sublette 1959 (Tendipes) 

Parachironomus potamogeti Townes 1945 (Harnischia) 

Parachironomus schneideri Beck and Beck 1969 (Parachironomus) 

Parachironomus sublettei Beck 1961 (Chironomus) 

Parachironomus tenuicaudatus Malloch 1915 (Chironomus) 

Parachironomus varus Goetghebuer 1921 (Chironomus) 

Parachironomus vitiosus Goetghebuer 1921 (Chironomus) 

Paracladius Hirvenoja 1973 

Paratrichocladius Thienemann 1942 Homo. 

Paracladius alpicola Zetterstedt 1850 (Chironomus) 

Paracladius conversus Walker 1856 (Chironomus) 

Paracladius quadrinodosus Hirvenoja 1973 (Paracladius) 

Paracladopelma Harnisch 1923 

Paracladopelma alphaeus Sublette 1960 (Tendipes) 

Paracladopelma camptolabis Kieffer 1913 (Tendipes) 

Paracladopelma doris Townes 1945 (Harnischia) 

Paracladopelma galaptera Townes 1945 (Harnischia) 

Chironomus claripennis Malloch 1915 Homo. 

Paracladopelma loganae Beck and Beck 1969 (Paracladopelma) 

Paracladopelma nais Townes 1945 (Harnischia) 

Paracladopelma nereis Townes 1945 (Harnischia) 

Paracladopelma nixe Townes 1945 (Harnischia) 

Paracladopelma rolli Chernovskij 1949 (Cryptochironomus) 

Paracladopelma undine Townes 1945 (Harnischia) 

Paracladopelma winnelli Jackson 1977 (Paracladopelma) 

Paracricotopus Thienemann and Harnisch 1932 

Paracricotopus glaber Saether 1980 (Paracricotopus) 

Paracricotopus millrockensis Caldwell 1985 (Paracricotopus) 

Paracricotopus mozleyi Steiner 1983 (Paracricotopus) 

Parakiefferiella Thienemann 1936 

Maricladius Sublette 1970 Syn. 

Parakiefferiella bathophilus Kieffer 1912 (Dactylocladius) 

Parakiefferiella coronata Edwards 1929 (Spaniotoma) 

Parakiefferiella gracillimus Kieffer 1924 (Camptocladius) 

Parakiefferiella nigra Brundin 1949 (Parakiefferiella) 

Parakiefferiella subaterrimus Malloch 1915 (Camptocladius) 

Parakiefferiella torulata Saether 1969 Syn. 

Paralauterborniella Lenz 1941 


115

116 

Paralauterborniella nigrohalteralis Malloch 1915 (Chironomus) 

Paralimnophyes Brundin 1956 

Paralimnophyes arcticus Brundin 1956 (Paralimnophyes) 

Paramerina Fittkau 1962 

Paramerina anomala Beck and Beck 1966 (Paramerina) 

Paramerina fragilis Walley 1926 (Tanypus) 

Paramerina hanseni Roback 1971 (Paramerina) 

Paramerina smithae Sublette 1964 (Pentaneura) 

Paramerina testa Roback 1971 (Paramerina) 

Parametriocnemus Goetghebuer 1932 

Parametriocnemus eoclivus Saether 1969 (Parametriocnemus) 

Parametriocnemus graminicola Lundbeck 1898 (Chironomus) 

Parametriocnemus hamatus Johannsen 1934 (Metriocnemus) 

Parametriocnemus lundbeckii Johannsen 1905 (Metriocnemus) 

Chironomus nanus Lundbeck 1898 Homo. 

Metriocnemus innocuus Curran 1930 Syn. 

Parametriocnemus stylatus Kieffer 1924 (Metriocnemus) 

Parametriocnemus vespertinus Saether 1969 (Parametriconemus) 

Paraphaenocladius Thienemann 1926 

Paraphaenocladius brevinervis Holmgren 1869 (Chironomus) 

Metriocnemus despectus Kieffer 1926 Syn. 

Paraphaenocladius debilipennis Lundbeck 1898 (Chironomus) 

Paraphaenocladius exagitans Johannsen 1905 (Metriocnemus) 

Metriocnemus brachyneura Malloch 1915 Syn. 

Paraphaenocladius impensus Walker 1856 (Chironomus) 

Paraphaenocladius nasthecus Saether 1969 (Paraphaenocladius) 

Parasmittia Strenzke 1950 

Parasmittia carinata Strenzke 1950 (Parasmittia) 

Paratanytarsus Thienemann and Bause 1913 

Stylotanytarsus Kieffer 1921 Syn. 

Paratanytarsus dissimilis Johannsen 1905 (Tanytarsus) 

Tanytarsus confusus Malloch 1915 Syn. 

Paratanytarsus dubius Malloch 1915 (Tanytarsus) 

Paratanytarsus grimmii Schneider 1885 (Chironomus) 

Tanytarsus boiemicus Bause 1913 Syn. 

Paratanytarsus inopertus Walker 1856 (Chironomus) 

Paratanytarsus intricatus Goetghebuer 1921 (Tanytarsus) 

Paratanytarsus laccophilus Edwards 1929 (Tanytarsus) 

Paratanytarsus natvigi Goetghebuer 1933 (Micropsectra) 

Paratanytarsus pencillatus Goetghebuer 1928 (Tanytarsus) 

Paratanytarsus setosimanus Goetghebuer 1933 (Ditanytarsus) 

Paratanytarsus similatus Malloch 1915 (Tanytarsus) 

Paratanytarsus tenuis Meigen 1830 (Chironomus) 

Paratanytarsus tolucensis Reiss 1972 (Paratanytarsus) 

Paratendipes Kieffer 1911 

Paratendipes albimanus Meigen 1818 (Chironomus) 

Chironomus annularis Meigen 1804 Homo. 

Paratendipes basidens Townes 1945 (Paratendipes) 

Paratendipes duplicatus Johannsen 1937 (Chironomus) 

Chironomus basalis Malloch 1915 Homo. 

Paratendipes fuscitibia Sublette 1960 (Paratendipes) 

Paratendipes nitidulus Coquillett 1901 (Chironomus) 

Chironomus nitidellus Malloch 1915 Missp. 



Paratendipes nudisquama Edwards 1929 (Chironomus) 

Paratendipes subaequalis Malloch 1915 (Chironomus) 

Paratendipes thermophilus Townes 1945 (Paratendipes) 

Paratrichocladius Santos Abreu 1918 

Syncricotopus Brundin 1956 Syn. 

Paratrichocladius nitidus Malloch 1915 (Orthocladius) 

Trichocladius helis Roback 1957 Syn. 

Syncricotopus fontinalis Saether 1969 Syn. 

Paratrichocladius rufiventris Meigen 1830 (Chironomus) 

Paratrichocladius skirwithensis Edwards 1929 (Spaniotoma) 

Paratrissocladius Zavrel 1937 

Paratrissocladius lawrencensis Cranston and Oliver 1988 

(Paratrissocladius) 

Parochlus Enderlein 1912 

Paratanypus Garrett 1925 Syn. 

Parochlus kiefferi Garrett 1925 (Paratanypus) 

Tanypus tibialis Staeger 1845 Homo. 

Podonomus peregrinus Edwards 1929 Syn. 

Pentaneura Philippi 1865 

Pentaneura futilis Wulp 1867 (Tanypus) 

Pentaneura inconspicuus Malloch 1915 (Tanypus) 

Pentaneura comosa Sublette 1964 Syn. 

Pentaneura inculta Beck and Beck 1966 Syn. 

Pentaneura indecisus Williston 1896 (Tanypus) 

Pentaneura inyoensis Sublette 1964 (Pentaneura) 

Phaenopsectra Kieffer 1921 

Phaenopsectra dyari Townes 1945 (Tanytarsus) 

Phaenopsectra flavipes Meigen 1818 (Chironomus) 

Tanytarsus flavicauda Malloch 1915 Syn. 

Phaenopsectra incomptus Zetterstedt 1838 (Chironomus) 

Phaenopsectra mortensoni Grodhaus 1987 (Phaenopsectra) 

Phaenopsectra obediens Johannsen 1905 (Tanytarsus) 

Phaenopsectra pilicellata Grodhaus 1976 (Phaenopsectra) 

Phaenopsectra profusus Townes 1945 (Tanytarsus) 

Phaenopsectra punctipes Wiedemann 1817 (Chironomus) 

Phaenopsectra vittatus Townes 1945 (Tanytarsus) 

Platysmittia Saether 1982 

Platysmittia bilyji Saether 1985 (Platysmittia) 

Platysmittia fimbriata Saether 1982 (Platysmittia) 

Plhudsonia Saether 1982 

Plhudsonia partita Saether 1982 (Plhudsonia) 

Polypedilum Kieffer 1913 

Pentapedilum Kieffer 1913 Syn. 

Tripodura Townes 1945 Syn. 

Polypedilum acifer Townes 1945 (Polypedilum) 

Polypedilum albicornis Meigen 1838 (Chironomus) 

Polypedilum nigritum Townes 1945 Syn. 

Polypedilum albinodus Townes 1945 (Polypedilum) 

Polypedilum angustum Townes 1945 (Polypedilum) 

Polypedilum apicatum Townes 1945 (Polypedilum) 

Polypedilum artifer Curran 1930 (Chironomus) 

Polypedilum aviceps Townes 1945 (Polypedilum)

Polypedilum braseniae Leathers 1922 (Chironomus) 

Polypedilum californicum Sublette 1960 (Polypedilum) 

Polypedilum calopterus Mitchell 1908 (Chironomus) 

Polypedilum walleyi Townes 1945 Syn. 

Polypedilum cinctum Townes 1945 (Polypedilum) 

Polypedilum convictus Walker 1856 (Chironomus) 

Chironomus flavus Johannsen 1905 Syn. 

Chironomus flaviventris Johannsen 1907 Syn. 

Polypedilum obtusum Townes 1945 Syn. 

Polypedilum fallax Johannsen 1905 (Chironomus) 

Polypedilum floridense Townes 1945 (Polypedilum) 

Polypedilum fuscipennis Meigen 1818 (Chironomus) 

Polypedilum gomphus Townes 1945 (Polypedilum) 

Polypedilum halteralis Coquillett 1901 (Chironomus) 

Chironomus griseopunctatus Malloch 1915 Syn. 

Polypedilum digitifer Townes 1945 Syn. 

Polypedilum simulans Townes 1945 Syn. 

Polypedilum illinoensis Malloch 1915 (Chironomus) 

Chironomus decoloratus Malloch 1915 Syn. 

Polypedilum isocerus Townes 1945 (Polypedilum) 

Polypedilum labeculosus Mitchell 1908 (Chironomus) 

Polypedilum laetus Meigen 1818 (Chironomus) 

Polypedilum nubeculosus Meigen 1804 (Chironomus) 

Polypedilum ontario Walley 1926 (Chironomus) 

Chironomus hirtipes Mitchell 1908 Homo. 

Polypedilum ophioides Townes 1945 (Polypedilum) 

Polypedilum parascalaenum Beck 1962 (Polypedilum) 

Polypedilum pardus Townes 1945 (Polypedilum) 

Polypedilum parvum Townes 1945 (Polypedilum) 

Polypedilum pedatum Townes 1945 (Polypedilum) 

Polypedilum excelsius Townes 1945 Syn. 

Polypedilum pedestris Meigen 1830 (Chironomus) 

Polypedilum pterospilus Townes 1945 (Polypedilum) 

Polypedilum scalaena Schrank 1803 (Tipula) 

Chironomus needhamii Johannsen 1908 Syn. 

Polypedilum sordens Wulp 1874 (Tanytarsus) 

Tanytarsus fulvescens Johannsen 1905 Syn. 

Calopsectra americana Kieffer 1917 Syn. 

Polypedilum subcultellatum Sublette 1960 (Polypedilum) 

Polypedilum sulaceps Townes 1945 (Polypedilum) 

Polypedilum trigonus Townes 1945 (Polypedilum) 

Polypedilum tritus Walker 1856 (Chironomus) 

Polypedilum albulum Townes 1945 Syn. 

Polypedilum vibex Townes 1945 (Polypedilum) 

Potthastia Kieffer 1922 

Potthastia gaedii Meigen 1838 (Diamesa) 

Potthastia longimanus Kieffer 1922 (Potthastia) 

Potthastia montium Edwards 1929 (Diamesa) 

Potthastia iberica Serra-Tosio 1971 Syn. 

Procladius Skuse 1889 

Psilotanypus Kieffer 1906 Syn. 

Holotanypus Roback 1982 Syn. 

Procladius abetus Roback 1971 (Procladius) 

Procladius abrupta Garrett 1925 (Tanypus) 

Procladius arcuata Garrett 1925 (Tanypus) 

Procladius barbatulus Sublette 1964 (Procladius) 

Procladius bellus Loew 1866 (Tanypus) 

Tanypus pusillus Loew 1866 Syn. 

Procladius adumbratus Johannsen 1905 Syn. 

Procladius flavidus Kieffer 1923 Syn. 

Procladius malifero Garrett 1925 Syn. 

Procladius bifida Garrett 1925 (Tanypus) 

Procladius clavus Roback 1971 (Procladius) 

Procladius culiciformis Linnaeus 1767 (Tipula) 


Tanypus crassinervis Zetterstedt 1838 Syn. 

Protenthes claripennis Malloch 1915 Syn. 

Procladius curtus Roback 1971 (Procladius) 

Procladius deltaensis Roback 1987 (Procladius) 

Procladius denticulatus Sublette 1964 (Procladius) 

Procladius dentus Roback 1971 (Procladius) 

Procladius desis Roback 1971 (Procladius) 

Procladius freemani Sublette 1964 (Procladius) 

Procladius asetus Roback 1971 Unav. 

Procladius brunneus Roback 1971 Unav. 

Procladius cretis Roback 1971 Unav. 

Procladius elongatus Roback 1971 Unav. 

Procladius gretis Roback 1971 (Procladius) 

Procladius jeris Roback 1971 (Procladius) 

Procladius johnsoni Roback 1980 (Procladius) 

Procladius lundstromi Goetghebuer 1936 (Procladius) 

Procladius macrotrichus Roback 1971 (Procladius) 

Procladius nietus Roback 1971 (Procladius) 

Procladius paragretis Roback 1971 (Procladius) 

Procladius prolongatus Roback 1971 (Procladius) 

Procladius riparius Malloch 1915 (Protenthes) 

Procladius ruris Roback 1971 (Procladius) 

Procladius grandis Roback 1971 Unav. 

Procladius sublettei Roback 1971 (Procladius) 

Procladius grandis Roback 1971 Unav. 

Procladius minuta Roback 1971 Unav. 

Procladius trifolia Garrett 1925 (Tanypus) 

Procladius vestitipennnis Kieffer 1917 (Trichotanypus) 

Procladius vesus Roback 1971 (Procladius) 

Procladius wilhmi Roback 1966 (Procladius) 

Prodiamesa Kieffer 1906 

Prodiamesa cubita Garrett 1925 (Prodiamesa) 

Prodiamesa olivaceus Meigen 1818 (Chironomus) 

Chironomus turpis Zetterstedt 1838 Syn. 

Tanypus occidentalis Coquillett 1902 Syn. 

Protanypus Kieffer 1906 

Protanypus caudatus Edwards 1924 (Protanypus) 

Protanypus hamiltoni Saether 1975 (Protanypus) 

Protanypus morio Zetterstedt 1838 (Tanypus) 

Protanypus pseudomorio Makarchenko 1982 (Protanypus) 

Protanypus ramosus Saether 1975 (Protanypus) 

Protanypus saetheri Wiederholm 1975 (Protanypus) 

Psectrocladius Kieffer 1906 

Allopsectrocladius Wülker 1956 Syn. 

Monopsectrocladius Wülker 1956 Syn. 

Psectrocladius barbatimanus Kieffer 1926 (Psectrocladius) 

Psectrocladius barbimanus Edwards 1929 (Spaniotoma) 

Psectrocladius armatus Andersen 1937 Syn. 

Psectrocladius brevicosta Kieffer 1917 (Psectrocladius) 

Psectrocladius calcarata Edwards 1929 (Spaniotoma) 

Psectrocladius elatus Roback 1957 (Psectrocladius) 

Psectrocladius fennicus Storå 1939 (Psectrocladius) 

Psectrocladius flavus Johannsen 1905 (Orthocladius) 

Psectrocladius limbatellus Holmgren 1869 (Chironomus) 

Psectrocladius edwardsi Brundin 1949 Syn. 

Psectrocladius nigrus Roback 1957 (Psectrocladius) 

Psectrocladius obvius Walker 1856 (Chironomus) 

Psectrocladius octomaculatus Wülker 1956 (Psectrocladius) 

Psectrocladius pilosus Roback 1957 (Psectrocladius) 

Psectrocladius polaris Kieffer 1926 (Psectrocladius) 

Psectrocladius psilopterus Kieffer 1906 (Orthocladius) 

Psectrocladius semicirculatus Saether 1969 (Psectrocladius) 


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118 

Psectrocladius simulans Johannsen 1937 (Spaniotoma) 

Psectrocladius sordidellus Zetterstedt 1838 (Chironomus) 

Psectrocladius spinifer Johannsen 1928 (Orthocladius) 

Psectrocladius vernalis Malloch 1915 (Orthocladius) 

Psectrocladius zetterstedti Brundin 1949 (Psectrocladius) 

Psectrotanypus Kieffer 1909 

Psectrotanypus discolor Coquillett 1902 (Tanypus) 

Psectrotanypus dyari Coquillett 1902 (Tanypus) 

Tanypus guttularis Coquillett 1902 Syn. 

Tanypus garretti Walley 1925 Syn. 

Psectrotanypus pictipennis Zetterstedt 1838 (Tanypus) 

Pseudochironomus Malloch 1915 



Pseudochironomus aix Townes 1945 (Pseudochironomus) 

Pseudochironomus anas Townes 1945 (Pseudochironomus) 

Pseudochironomus atricaudus Saether 1977 (Pseudochironomus) 

Pseudochironomus aureus Johannsen 1908 (Psectrocladius) 

Pseudochironomus badius Saether 1977 (Pseudochironomus) 

Pseudochironomus banksi Townes 1945 (Pseudochironomus) 

Pseudochironomus chen Townes 1945 (Pseudochironomus) 

Pseudochironomus crassus Townes 1945 (Pseudochironomus) 

Pseudochironomus fulviventris Johannsen 1905 (Chironomus) 

Pseudochironomus julia Curran 1930 (Orthocladius) 

Pseudochironomus middlekauffi Townes 1945 (Pseudochironomus) 

Pseudochironomus netta Townes 1945 (Pseudochironomus) 

Pseudochironomus pseudoviridis Malloch 1915 (Chironomus) 

Pseudochironomus rex Hauber 1947 (Pseudochironomus) 

Pseudochironomus richardsoni Malloch 1915 (Pseudochironomus) 

Pseudodiamesa Goetghebuer 1939 

Pachydiamesa Oliver 1959 Syn. 

Pseudodiamesa arctica Malloch 1919 (Diamesa) 

Pseudodiamesa branickii Nowicki 1873 (Diamesa) 

Pseudodiamesa pertinax Garrett 1925 (Prodiamesa) 

Pseudokiefferiella Zavrel 1941 

Diplomesa Pagast 1947 Syn. 

Pseudokiefferiella parva Edwards 1932 (Diamesa) 

Syndiamesa polaris Kieffer 1926 Homo. 

Pseudorthocladius Goetghebuer 1932 

Lordella Saether and Sublette 1983 Syn. 

Pseudorthocladius amplicaudus Saether and Sublette 1983 

(Pseudorthocladius) 

Pseudorthocladius clavatosus Saether and Sublette 1983 


Pseudorthocladius comans Saether and Sublette 1983 


Pseudorthocladius curticornus Saether and Sublette 1983 


Pseudorthocladius curtistylus Goetghebuer 1921 


Pseudorthocladius destitutus Saether and Sublette 1983 


Pseudorthocladius dumicaudus Saether 1969 (Pseudorthocladius) 

Pseudorthocladius lunatus Saether and Sublette 1983 


Pseudorthocladius macrostomus Soponis 1980 (Pseudorthocladius) 

Pseudorthocladius macrovirgatus Saether and Sublette 1983 


Pseudorthocladius cranstoni Saether and Sublette 1983 Syn. 

Pseudorthocladius morsei Saether and Sublette 1983 


Pseudorthocladius paravirgatus Saether and Sublette 1983 


Pseudorthocladius pilosipennis Brundin 1956 (Pseudorthocladius) 

Pseudorthocladius rectilobus Saether and Sublette 1983 


Pseudorthocladius tricanthus Saether and Sublette 1983 


Pseudorthocladius uniserratus Saether and Sublette 1983 


Pseudorthocladius virgatus Saether and Sublette 1983 


Pseudorthocladius wingoi Saether and Sublette 1983 


Pseudosmittia Goetghebuer 1932 

Prosmittia Brundin 1956 Syn. 

Pseudosmittia conjuncta Edwards 1929 (Spaniotoma) 

Pseudosmittia digitata Saether 1981 (Pseudosmittia) 

Pseudosmittia forcipatus Goetghebuer 1921 (Camptochironomus) 

Smittia triappendiculata Goetghebuer 1931 Syn. 

Pseudosmittia gracilis Goetghebuer 1913 (Comptocladius) 

Pseudosmittia nanseni Kieffer 1926 (Camptocladius) 

Pseudosmittia oxoniana Edwards 1922 (Camptocladius) 

Pseudosmittia setavena Saether 1969 (Pseudosmittia) 

Psilometriocnemus Saether 1969 

Psilometriocnemus triannulatus Saether 1969 (Psilometriocnemus) 

Psilometriocnemus cristatus Saether 1982 Syn. 

Radotanypus Fittkau and Murray 1985 

Radotanypus submarginella Sublette 1964 (Anatopynia) 

Rheocricotopus Thienemann and Harnisch 1932 

Psilocricotopus Saether 1985 Syn. 

Rheocricotopus amplicristatus Saether 1985 (Rheocricotopus) 

Rheocricotopus chapmani Edwards 1935 (Spaniotoma) 

Rheocricotopus conflusirus Saether 1985 (Rheocricotopus) 

Rheocricotopus effusoides Saether 1985 (Rheocricotopus) 

Rheocricotopus effusus Walker 1856 (Chironomus) 

Orthocladius striatus Malloch 1915 Syn. 

Rheocricotopus eminellobus Saether 1969 (Rheocricotopus) 

Rheocricotopus glabricollis Meigen 1830 (Chironomus) 

Trichocladius gouini Goetghebuer 1936 Syn. 

Trichocladius extatus Roback 1957 Syn. 

Rheocricotopus pauciseta Saether 1969 (Rheocricotopus) 

Rheocricotopus robacki Beck and Beck 1964 (Trichocladius) 

Rheocricotopus kenorensis Saether 1969 Syn. 

Rheocricotopus tuberculatus Caldwell 1984 (Rheocricotopus) 

Rheocricotopus unidentatus Saether and Schnell 1988 

(Rheocricotopus) 

Rheomyia Roback 1986 

Rheomyia wartinbei Roback 1984 (Zavrelimyia) 

Rheopelopia Fittkau 1962 

Rheopelopia acra Roback 1971 (Thienemannimyia) 

Rheopelopia paramaculipennis Roback 1971 (Thienemannimyia) 

Rheopelopia perda Roback 1971 (Thienemannimyia) 

Rheotanytarsus Bause 1913


Rheotanytarsus akrina Roback 1960 (Calopsectra) 

Rheotanytarsus distinctissima Brundin 1947 (Stempellinella) 

Rheotanytarsus exiguus Johannsen 1905 (Tanytarsus) 

Rheotanytarsus magnini Cloutier and Harper 1986 (Rheotanytarsus) 

Rheotanytarsus pellucidus Walker 1848 (Chironomus) 

Robackia Saether 1977 

Robackia claviger Townes 1945 (Harnischia) 

Robackia demeijerei Kruseman 1933 (Tendipes) 

Robackia pilicauda Saether 1977 (Robackia) 

Saetheria Jackson 1977 

Saetheria hirta Saether 1983 (Saetheria) 

Saetheria tylus Townes 1945 (Harnischia) 

Saetheriella Halvorsen 1982 

Saetheriella amplicristata Halvorsen 1982 (Saetheriella) 

Sergentia Kieffer 1922 

Sergentia albescens Townes 1945 (Tanytarsus) 

Sergentia coracinus Zetterstedt 1850 (Chironomus) 

Skutzia Reiss 1985 

Skutzia inopinata Reiss 1985 (Skutzia) 

Smittia Holmgren 1869 

Phaenocladius Kieffer 1921 Syn. 

Smittia arctica Malloch 1923 (Smittia) 

Smittia aterrima Meigen 1818 (Chironomus) 

Smittia crassicollis Walker 1848 (Chironomus) 

Smittia edwardsi Goetghebuer 1932 (Smittia) 

Smittia extremus Holmgren 1869 (Chironomus) 

Smittia flavibasis Malloch 1915 (Camptocladius) 

Smittia fulvipluma Kieffer 1926 (Camptocladius) 

Smittia lasiophthalmus Malloch 1915 (Camptocladius) 

Smittia lasiops Malloch 1915 (Camptocladius) 

Smittia longitibia Goetghebuer 1933 (Smittia) 

Smittia parvus Lundbeck 1898 (Chironomus) 

Smittia polaris Kieffer 1926 (Camptocladius) 

Smittia polymorpha Andersen 1937 (Smittia) 

Smittia roena Roback 1957 (Smittia) 

Smittia subnudipennis Goetghebuer 1933 (Smittia) 

Smittia velutinus Lundbeck 1898 (Chironomus) 

Stackelbergina Shilova and Zelentsov 1978 

Stackelbergina praeclara Shilova and Zelentsov 1978 

(Stackelbergina) 

Stelechomyia Reiss 1982 

Stelechomyia perpulcher Mitchell 1908 (Chironomus) 

Stempellina Bause 1913 

Stempellina johannsenii Thienemann and Bause 1913 (Tanytarsus) 

Stempellina ranota Webb 1969 (Stempellina) 

Stempellina rodesta Webb 1969 (Stempellina) 

Stempellinella Brundin 1947 

Stempellinella brevis Edwards 1929 (Tanytarsus) 

Stempellinella leptocelloides Webb 1969 (Stempellina) 

Stempellinella minor Edwards 1929 (Tanytarsus) 

Stenochironomus Kieffer 1919 

Petalopholeus Borkent 1984 Syn. 

Stenochironomus aestivalis Townes 1945 (Stenochironomus) 

Stenochironomus albipalpus Borkent 1984 (Stenochironomus) 

Stenochironomus annettae Borkent 1984 (Stenochironomus) 

Stenochironomus bisetosus Borkent 1984 (Stenochironomus) 

Stenochironomus browni Townes 1945 (Stenochironomus) 

Stenochironomus cinctus Townes 1945 (Stenochironomus) 

Stenochironomus colei Malloch 1919 (Chironomus) 

Stenochironomus fuscipatellus Borkent 1984 (Stenochironomus) 

Stenochironomus hilaris Walker 1848 (Chironomus) 

Chironomus taeniapennis Coquillett 1901 Syn. 

Chironomus nephopterus Mitchell 1908 Syn. 

Chironomus exquisitus Mitchell 1908 Syn. 

Chironomus zonopterus Mitchell 1908 Syn. 

Stenochironomus macateei Malloch 1915 (Chironomus) 

Stenochironomus maculatus Borkent 1984 (Stenochironomus) 

Stenochironomus niger Borkent 1984 (Stenochironomus) 

Stenochironomus poecilopterus Mitchell 1908 (Chironomus) 

Stenochironomus pulchripennis Coquillett 1902 (Chironomus) 

Stenochironomus totifuscus Sublette 1960 (Stenochironomus) 

Stenochironomus unictus Townes 1945 (Stenochironomus) 

Stenochironomus woodi Borkent 1984 (Stenochironomus) 

Stictochironomus Kieffer 1919 

Stictochironomus albicrus Townes 1945 (Tanytarsus) 

Stictochironomus annulicrus Townes 1945 (Tanytarsus) 

Stictochironomus devinctus Say 1829 (Chironomus) 

Chironomus compes Coquillett 1908 Syn. 

Chironomus ornatipes Kieffer 1917 Syn. 

Stictochironomus flavicingula Walker 1848 (Chironomus) 

Stictochironomus lutosus Townes 1945 (Tanytarsus) 

Stictochironomus marmoreus Townes 1945 (Tanytarsus) 

Stictochironomus naevus Mitchell 1908 (Chironomus) 

Stictochironomus palliatus Coquillett 1902 (Chironomus) 

Stictochironomus pictulus Meigen 1830 (Chironomus) 

Chironomus sticticus Meigen 1818 Homo. 

Stictochironomus quagga Townes 1945 (Tanytarsus) 

Stictochironomus rosenschoeldi Zetterstedt 1838 (Chironomus) 

Stictochironomus stictica Fabricius 1781 (Tipula) 

Tipula histrio Fabricius 1794 Syn. 

Stictochironomus unguiculatus Malloch 1934 (Chironomus) 

Stictochironomus varius Townes 1945 (Tanytarsus) 

Stictochironomus virgatus Townes 1945 (Tanytarsus) 

Stilocladius Rossaro 1979 

Stilocladius clinopecten Saether 1982 (Stilocladius) 

Sublettea Roback 1975 

Sublettea coffamni Roback 1975 (Tanytarsus) 

Sublettiella Saether 1983 

Sublettiella calvata Saether 1983 (Sublettiella) 

Symbiocladius Kieffer 1925 

Symbiocladius chattahoocheensis Caldwell 1984 (Symbiocladius) 

Symbiocladius equitans Claassen 1922 (Trissocladius) 


119

120 

Sympotthastia Pagast 1947 

Sympotthastia diastena Sublette 1964 (Pseudodiamesa) 

Sympotthastia fulva Johannsen 1921 (Diamesa) 

Sympotthastia zavreli Pagast 1947 (Sympotthastia) 

Synendotendipes Grodhaus 1987 

Synendotendipes luski Grodhaus 1987 (Synendotendipes) 

Synorthocladius Thienemann 1935 

Synorthocladius semivirens Kieffer 1909 (Dactylocladius) 

Tanypus Meigen 1803 

Pelopia Meigen 1800 Unav. 

Protenthes Johannsen 1907 Syn. 

Apelopia Roback 1971 Syn. 

Tanypus carinatus Sublette 1964 (Tanypus) 

Tanypus clavatus Beck 1962 (Tanypus) 

Tanypus concavus Roback 1971 (Tanypus) 

Tanypus grodhausi Sublette 1964 (Tanypus) 

Tanypus imperialis Sublette 1964 (Tanypus) 

Tanypus neopunctipennis Sublette 1964 (Tanypus) 

Tanypus nubifer Coquillett 1905 (Tanypus) 

Tanypus parastellatus Sublette 1964 (Tanypus) 

Tanypus pubitarsis Zetterstedt 1850 (Tanypus) 

Tanypus punctipennis Meigen 1818 (Tanypus) 

Protenthes americanus Kieffer 1923 Syn. 

Tanypus stellatus Coquillett 1902 (Tanypus) 

Tanypus telus Roback 1971 (Tanypus) 

Tanypus tibialis Say 1823 (Tanypus) 

Tanypus vilipennis Kieffer 1918 (Protenthes) 

Tanytarsus Wulp 1874 

Calopsectra Kieffer 1909 Syn. 

Tanytarsus allicis Sublette 1964 (Tanytarsus) 

Tanytarsus anderseni Reiss and Fittkau 1971 (Tanytarsus) 

Tanytarsus brundini Lindeberg 1963 (Tanytarsus) 

Tanytarsus buckleyi Sublette 1964 (Tanytarsus) 

Tanytarsus curticornis Kieffer 1911 (Tanytarsus) 

Tanytarsus debilis Meigen 1830 (Chironomus) 

Tanytarsus dendyi Sublette 1964 (Tanytarsus) 

Tanytarsus dispar Lindeberg 1967 (Tanytarsus) 

Tanytarsus fatigans Johannsen 1905 (Tanytarsus) 

Tanytarsus fimbriatus Reiss and Fittkau 1971 (Tanytarsus) 

Tanytarsus glabrescens Edwards 1929 (Tanytarsus) 

Tanytarsus gracilentus Holmgren 1883 (Tanytarsus) 

Chironomus deviatus Malloch 1923 Syn. 

Tanytarsus guerla Roback 1957 (Calopsectra) 

Tanytarsus inaequalis Goetghebuer 1921 (Tanytarsus) 

Tanytarsus islandica Malloch 1934 (Tanytarsus) 

Tanytarsus lapponicus Lindeberg 1970 (Tanytarsus) 

Tanytarsus limneticus Sublette 1964 (Tanytarsus) 

Tanytarsus mendax Kieffer 1925 (Tanytarsus) 

Tanytarsus holochlorus Edwards 1929 Syn. 

Tanytarsus muticus Kieffer 1905 (Tanytarsus) 

Tanytarsus nemorosus Edwards 1929 (Tanytarsus) 

Tanytarsus neoflavellus Malloch 1915 (Tanytarsus) 

Tanytarsus niger Andersen 1937 (Tanytarsus) 

Tanytarsus norvegica Kieffer 1924 (Fournieria) 

Tanytarsus pallidicornis Walker 1856 (Chironomus) 

Tanytarsus quadratus Sublette 1964 (Tanytarsus) 

Tanytarsus quadridentatus Brundin 1947 (Tanytarsus) 

Tanytarsus recens Sublette 1964 (Tanytarsus) 

Tanytarsus recurvatus Brundin 1947 (Tanytarsus) 



Tanytarsus signatus Wulp 1858 (Chironomus) 

Tanytarsus tibialis Webb 1969 (Tanytarsus) 

Tanytarsus trilobus Webb 1969 (Tanytarsus) 

Tanytarsus usmaensis Pagast 1931 (Tanytarsus) 

Tanytarsus varela Roback 1957 (Calopsectra) 

Tanytarsus xanthus Sublette 1964 (Tanytarsus) 

Tavastia Tuiskunen 1985 

Tavastia cristacauda Saether 1992 (Tavastia) 

Telmatogeton Schiner 1866 

Paraclunio Kieffer 1911 Syn. 

Telmatogeton alaskensis Coquillett 1900 (Telmatogeton) 

Telmatogeton japonicus Tokunaga 1933 (Telmatogeton) 

Telmatogeton macswaini Wirth 1949 (Telmatogeton) 

Telmatogeton spinosus Hashimoto 1973 (Paraclunio) 

Telmatogeton trilobatus Kieffer 1911 (Paraclunio) 

Telopelopia Roback 1971 

Telopelopia okoboji Walley 1928 (Tanypus) 

Tethymyia Wirth 1949 

Tethymyia aptena Wirth 1949 (Tethymyia) 

Thalassomya Schiner 1856 

Thalassomya bureni Wirth 1949 (Thalassomya) 

Thalassosmittia Strenzke and Remmert 1957 

Saunderia Sublette 1967 Syn. 

Thalassosmittia clavicornis Saunders 1928 (Camptocladius) 

Thalassosmittia marinus Saunders 1928 (Camptocladius) 

Thalassosmittia pacificus Saunders 1928 (Camptocladius)

Thienemannia Kieffer 1909 

Thienemannia pilinucha Saether 1985 (Thienemannia) 

Thienemanniella Kieffer 1911 

Thienemanniella cubitus Garrett 1925 (Corynoneura) 

Thienemanniella elana Roback 1957 (Corynoneura) 

Thienemanniella majuscula Edwards 1924 (Corynoneura) 

Thienemanniella mallochi Sublette 1970 (Thienemanniella) 

Thienemanniella similis Malloch 1915 (Corynoneura) 

Thienemanniella xena Roback 1957 (Corynoneura) 

Thienemannimyia Fittkau 1957 

Thienemannimyia barberi Coquillett 1902 (Tanypus) 

Thienemannimyia fusciceps Edwards 1929 (Pentaneura) 

Thienemannimyia norena Roback 1957 (Pentaneura) 

Thienemannimyia quadrata Roback 1971 Unav. 

Tokunagaia Saether 1973 

Tokunagaia excellens Brundin 1956 (Eukiefferiella) 

Tokunagaia obriani Hayes and Murray 1988 (Tokunagaia) 

Tokunagaia rowensis Saether 1969 (Adactylocladius) 

Tribelos Townes 1945 

Tribelos ater Townes 1945 (Tanytarsus) 

Tribelos fuscicornis Malloch 1915 (Chironomus) 

Tribelos jucundus Walker 1848 (Chironomus) 

Chironomus dimorphus Malloch 1915 Syn. 

Chironomus fusciventris Malloch 1915 Syn. 

Tanytarsus protextus Townes 1945 Syn. 

Tribelos subatrum Grodhaus 1987 (Tribelos) 

Tribelos subletteorum Grodhaus 1987 (Tribelos) 

Trichochilus Saether 1985 

Trichochilus lacteipennnis Johannsen 1908 (Trichocladius) 

Trichocladius of authors 


Trichocladius of authors ater Goetghebuer 1955 (Trichocladius) 

Trichocladius of authors pubitarsis Zetterstedt 1838 (Chironomus) 

Trichocladius of authors septris Roback 1957 (Trichocladius) 

Trichotanypus Kieffer 1906 

Trichotanypus alaskensis Brundin 1966 (Trichotanypus) 

Trichotanypus foliaceus Wirth and Sublette 1970 (Trichotanypus) 

Trichotanypus hanseni Wirth and Sublette 1970 (Trichotanypus) 

Trichotanypus mariae Wirth and Sublette 1970 (Trichotanypus) 

Trichotanypus posticalis Lundbeck 1898 (Tanypus) 

Trissopelopia Kieffer 1923 

Trissopelopia ogemawi Roback 1971 (Trissopelopia) 

Tvetenia Kieffer 1922 

Tvetenia calvescens Edwards 1929 (Spaniotoma) 

Tvetenia duodenaria Kieffer 1922 (Tvetenia) 

Eukiefferiella saanensis Wülker 1959 Syn. 

Tvetenia paucunca Saether 1969 (Eukiefferiella) 

Tvetenia vitracies Saether 1969 (Eukiefferiella) 

Unniella Saether 1982 

Unniella multivirga Saether 1982 (Unniella) 

Vivacricotopus Schnell and Saether 1988 

Vivacricotopus piloculus Cranston and Oliver 1988 (Hydrobaenus) 

Xenochironomus Kieffer 1921 

Xenochironomus xenolabis Kieffer 1916 (Chironomus) 

Xenopelopia Fittkau 1962 

Xenopelopia tincta Roback 1971 (Xenopelopia) 

Xestochironomus Sublette and Wirth 1972 

Insulanus Sublette 1967 Homo. 

Xestochironomus brunneus Borkent 1984 (Xestochironomus) 

Xestochironomus subletti Borkent 1984 (Xestochironomus) 

Xylotopus Oliver 1982 

Xylotopus par Coquillett 1901 (Orthocladius) 

Eurycnemus scitulus Coquillett 1901 Syn. 

Metriocnemus par Johannsen 1905 Homo. 

Zalutschia Lipina 1939 

Zalutschia briani Soponis 1979 (Zalutschia) 

Zalutschia furcarca Saether 1976 (Zalutschia) 

Zalutschia lingulata Saether 1976 (Zalutschia) 

Zalutschia pauca Saether 1976 Syn. 

Zalutschia obseptus Webb 1969 (Orthocladius) 

Zalutschia pusa Saether 1976 (Zalutschia) 

Zalutschia sivertseni Aagard 1979 (Eukiefferiella) 

Zalutschia tatrica Pagast 1935 (Spaniotoma) 

Zalutschia trigonacies Saether 1976 (Zalutschia) 

Zalutschia vockerothi Saether 1976 (Zalutschia) 

Zalutschia xethis Roback 1957 (Hydrobaenus) 

Zalutschia zalutschicola Lipina 1939 (Zalutschia) 

Zavreliella Kieffer 1920 

Zavreliella varipennis Coquillett 1902 (Chironomus) 

Zavrelimyia Fittkau 1962 

Zavrelimyia bifasciatus Coquillett 1901 (Tanypus) 

Zavrelimyia sinuosus Coquillett 1905 (Tanypus) 

Zavrelimyia carneosa Fittkau 1962 Syn. 

Pentaneura fluminalis Sublette 1964 Syn. 

Zavrelimyia thryptica Sublette 1964 (Pentaneura) 

CHLOROPIDAE 

Apallates Sabrosky 1980 

Apallates convexus Loew 1866 (Hippelates) 

Apallates coxendix Fitch 1856 (Oscinis) 

DIPTERA: CHIRONOMIDAE, CHLOROPIDAE 

121

122 

Oscinella pullicornis Sabrosky 1936 Syn. 

Apallates dissidens Tucker 1908 (Oscinis) 

Hippelates texanus Malloch 1913 Syn. 

Apallates hermsi Sabrosky 1941 (Hippelates) 

Apallates microcentrus Coquillett 1904 (Hippelates) 

Apallates montanus Sabrosky 1941 (Hippelates) 

Apallates neocoxendix Sabrosky 1940 (Oscinella) 

Apallates ochripes Sabrosky 1940 (Oscinella) 

Apallates particeps Becker 1912 (Oscinella) 

Hippelates subvittatus Malloch 1913 Syn. 

Apallates tener Coquillett 1900 (Hippelates) 

Aphanotrigonum Duda 1932 

Aphanotrigonella Narchuk 1964 Syn. 

Aphanotrigonum eulophus Loew 1872 (Hippelates) 

Aphanotrigonum scabra Aldrich 1918 (Oscinis) 

Biorbitella Sabrosky 1980 

Biorbitella frontoorbitalis Sabrosky 1940 (Oscinella) 

Biorbitella hesperia Sabrosky 1940 (Oscinella) 

Biorbitella pectoralis Coquillett 1898 (Oscinis) 

Biorbitella virgata Coquillett 1898 (Oscinis) 

Cadrema Walker 1859 

Prohippelates Malloch 1913 Syn. 

Cadrema pallidus Loew 1866 (Hippelates) 

Caviceps Malloch 1924 

Caviceps darlingtoniae Jones 1916 (Botanobia) 

Cetema Hendel 1907 

Centor Loew 1866 Homo. 

Centema Collin 1911 Emend. 

Centorella Strand 1928 Syn. 

Archecetema Narchuk 1976 Syn. 

Cetema procera Loew 1872 (Chlorops) 

Cetema subvittata Loew 1863 (Oscinis) 

Cetema hypocera Becker 1912 Syn. 

Chaetochlorops Malloch 1914 

Chaetochlorops inquilina Coquillett 1898 (Siphonella) 

Chlorops Meigen 1803 

Oscinis Latreille 1804 Syn. 

Cotilea Lioy 1864 Syn. 

Anthracophaga Loew 1866 Syn. 

Chloropsina Becker 1911 Syn. 

Lasiochlorops Duda 1934 Syn. 

Globiops Andersson 1977 Syn. 

Anthracophagella Andersson 1977 Syn. 

Sclerophallus Beschovski 1978 Syn. 

Asianochlorops Kanmiya 1984 Syn. 

Chlorops adamsi Sabrosky 1935 (Chlorops) 

Chlorops annulata Adams 1904 Homo. 

Chlorops atra Macquart 1851 (Chlorops) 

Chlorops brunnipennis Becker 1912 (Chlorops) 

Chlorops certima Adams 1904 (Chlorops) 

Chlorops cinerapennis Adams 1903 (Chlorops) 

Chlorops albifascies Adams 1903 Syn. 

Chlorops constricta Becker 1912 (Chlorops) 

Chlorops crocota Loew 1863 (Chlorops) 

DIPTERA: CHLOROPIDAE 


Chlorops declinata Becker 1912 (Anthracophaga) 

Chlorops pulvera Harbeck 1905 Unav. 

Chlorops egregia Becker 1912 (Chlorops) 

Chlorops genarum Becker 1912 (Chlorops) 

Chlorops laevis Becker 1912 (Chlorops) 

Chlorops languida Becker 1912 (Chlorops) 

Chlorops lasciva Adams 1904 (Chlorops) 

Chlorops liturata Adams 1903 (Chlorops) 

Chlorops melanocera Loew 1863 (Chlorops) 

Chlorops oblita Becker 1912 (Chlorops) 

Chlorops obscuricornis Loew 1863 (Chlorops) 

Chlorops obscuripennis Cole and Lovett 1921 Missp. 

Chlorops palpalis Adams 1903 (Chlorops) 

Chlorops perflava Walker 1849 (Chlorops) 

Chlorops producta Loew 1863 (Chlorops) 

Chlorops proximus Say 1830 (Chlorops) 

Chlorops pubescens Loew 1863 (Chlorops) 

Chlorops rectinervis Becker 1912 (Chlorops) 

Chlorops rubicunda Adams 1903 (Chlorops) 

Chlorops rubrivittata Adams 1904 (Chlorops) 

Chlorops rufescens Coquillett 1910 (Chlorops) 

Chlorops ruginosa Becker 1912 (Chlorops) 

Chlorops sabulona Becker 1912 (Chlorops) 

Chlorops sahlbergii Loew 1863 (Chlorops) 

Chlorops seminigra Becker 1912 (Chlorops) 

Chlorops sordidella Becker 1912 (Chlorops) 

Chlorops stigmata Becker 1912 (Chlorops) 

Chlorops stigmaticalis Becker 1912 (Chlorops) 

Chlorops quadrimaculata Becker 1912 Homo. 

Chlorops subnigra Coquillett 1910 (Chlorops) 

Chlorops sulphurea Loew 1863 (Chlorops) 

Chlorops tarsalis Becker 1912 (Chlorops) 

Chlorops testacea Macquart 1851 (Chlorops) 

Conioscinella Duda 1929 

Conioscinella flavescens Tucker 1908 (Oscinis) 

Conioscinella frontalis Tucker 1908 (Oscinis) 

Conioscinella grisescens Sabrosky 1940 (Oscinella) 

Conioscinella hinkleyi Malloch 1915 (Botanobia) 

Conioscinella nuda Adams 1905 (Oscinis) 

Conioscinella triorbiculata Sabrosky 1940 (Oscinella) 

Dasyopa Malloch 1918 

Trachysiphonella Enderlein 1936 Syn. 

Dasyopa latifrons Loew 1872 (Siphonella) 

Dasyopa pleuralis Malloch 1918 Syn. 

Dasyopa triangulata Becker 1912 (Siphonella) 

Dicraeus Loew 1873 

Parastia Pandelle 1898 Syn. 

Oxyapium Becker 1912 Syn. 

Dicraeinus Enderlein 1936 Syn. 

Paroedesiella Enderlein 1936 Syn. 

Eudicraeus Nartshuk 1967 Syn. 

Dicraeus aberrans Sabrosky 1950 (Dicraeus) 

Dicraeus elongatus Sabrosky 1950 (Dicraeus) 

Dicraeus incongruus Aldrich 1918 (Dicraeus) 

Dicraeus ingrata Loew 1866 (Eutropha) 

Dicraeus tibialis Macquart 1835 (Oscinis) 

Dicraeus wilburi Sabrosky 1950 (Dicraeus) 

Diplotoxa Loew 1863 

Diplotoxa alternata Loew 1872 (Chlorops) 

Diplotoxa inclinata Becker 1912 (Diplotoxa)

Chlorops confluens Loew 1872 Homo. 

Diplotoxa messoria Fallen 1820 (Oscinis) 

Chlorops bilineata Adams 1903 Syn. 

Diplotoxa nigripes Coquillett 1910 (Chlorops) 

Diplotoxa major Becker 1912 Syn. 

Diplotoxa recurva Adams 1903 (Chlorops) 

Diplotoxa unicolor Becker 1912 (Diplotoxa) 

Diplotoxa versicolor Loew 1863 (Chlorops) 

Dysartia Sabrosky 1991 

Dysartia latigena Sabrosky 1991 (Dysartia) 

Ectecephala Macquart 1851 

Leptotrigonum Becker 1912 

Ectecephala albistylum Macquart 1851 (Ectecephala) 

Ectecephala laticornis Coquillett 1910 (Ectecephala) 

Ectecephala sulcata Sabrosky 1941 (Ectecephala) 

Ectecephala sulcifrons Coquillett 1910 (Ectecephala) 

Ectecephala unicolor Loew 1863 (Chlorops) 

Ectecephala similis Becker 1912 Syn. 

Elachiptera Macquart 1835 

Crassiseta Roser 1840 Syn. 

Pachychaeta Loew 1845 Syn. 

Macrochetum Rondani 1856 Syn. 

Pachychoeta Bezzi 1895 Homo. 

Macrochaetum Corti 1909 Syn. 

Myrmecomorpha Corti 1909 Syn. 

Doliomyia Johannsen 1924 Syn. 

Neoelachiptera Seguy 1938 Syn. 

Elachiptera angusta Sabrosky 1948 (Elachiptera) 

Elachiptera angustifrons Sabrosky 1948 (Elachiptera) 

Elachiptera angustistylum Sabrosky 1948 (Elachiptera) 

Elachiptera californica Sabrosky 1948 (Elachiptera) 

Elachiptera costata Loew 1863 (Crassiseta) 

Elachiptera decipiens Loew 1863 (Oscinis) 

Elachiptera erythropleura Sabrosky 1948 (Elachiptera) 

Elachiptera flaviceps Sabrosky 1948 (Elachiptera) 

Elachiptera formosa Loew 1863 (Crassiseta) 

Elachiptera knowltoni Sabrosky 1948 (Elachiptera) 

Elachiptera longiventris Johannsen 1924 (Melanochaeta) 

Elachiptera nigriceps Loew 1863 (Crassiseta) 

Elachiptera pechumani Sabrosky 1948 (Elachiptera) 

Elachiptera penita Adams 1908 (Crassiseta) 

Elachiptera punctulata Becker 1912 (Elachiptera) 

Elachiptera nigroscutellata Becker 1912 Homo. 

Elachiptera salinaria Sabrosky and Valley 1987 (Elachiptera) 

Elachiptera tau Sabrosky 1948 (Elachiptera) 

Elachiptera vittata Sabrosky 1948 (Elachiptera) 

Elachiptera bilineata Adams 1904 Homo. 

Elachiptera willistoni Sabrosky 1948 (Elachiptera) 

Elliponeura Loew 1869 

Elliponeura debilis Loew 1869 (Elliponeura) 

Elliponeura diplotoxoides Becker 1912 (Elliponeura) 

Epichlorops Becker 1910 

Epichlorops exilis Coquillett 1898 (Eurina) 

Epichlorops puncticollis Zetterstedt 1848 (Oscinis) 

Epichlorops scabra Coquillett 1898 (Chlorops) 

Eribolus Becker 1910 


Eribolus californicus Sabrosky 1950 (Eribolus) 

Eribolus longulus Loew 1863 (Crassiseta) 

Melanochaeta intermedia Becker 1912 Syn. 

Eribolus nana Zetterstedt 1848 (Oscinis) 

Eribolus sudeticus Becker 1910 Syn. 

Oscinella planicollis Becker 1912 Syn. 

Eribolus nearcticus Sabrosky 1948 (Eribolus) 

Eugaurax Malloch 1913 

Eugaurax floridensis Malloch 1913 (Eugaurax) 

Eugaurax vittatus Sabrosky 1950 Syn. 

Fiebrigella Duda 1921 

Teleocoma Aldrich 1924 Syn. 

Goniopsis Duda 1929 Syn. 

Goniopsita Duda 1930 Syn. 

Laecometopa Duda 1930 Syn. 

Mimoscinis Malloch 1934 Syn. 

Lasiambia Sabrosky 1941 Syn. 

Fiebrigella catalpae Malloch 1913 (Botanobia) 

Fiebrigella magnipalpis Becker 1912 (Oscinella) 

Fiebrigella oophaga Sabrosky 1967 (Goniopsita) 

Gaurax Loew 1863 

Botanobia Lioy 1864 Syn. 

Neogaurax Malloch 1914 Syn. 

Pseudochlorops Duda 1930 

Pseudoscinella Duda 1931 

Gaurax apicalis Malloch 1915 (Gaurax) 

Gaurax atrilinea Sabrosky 1951 (Gaurax) 

Gaurax atripalpus Sabrosky 1951 (Gaurax) 

Gaurax basitarsalis Sabrosky 1951 (Gaurax) 

Gaurax dorsalis Loew 1863 (Oscinis) 

Gaurax dubia Macquart 1835 (Oscinis) 

Oscinis ephippium Zetterstedt 1848 Syn. 

Gaurax festivus Loew 1863 (Gaurax) 

Gaurax fumipennis Malloch 1915 (Neogaurax) 

Gaurax maculicornis Sabrosky 1951 (Gaurax) 

Gaurax maculipes Sabrosky 1951 (Gaurax) 

Gaurax melanotum Sabrosky 1951 (Gaurax) 

Gaurax mesopedalis Sabrosky 1951 (Gaurax) 

Gaurax montanus Coquillett 1898 (Gaurax) 

Gaurax obscuripennis Johnson 1913 (Gaurax) 

Gaurax ocellaris Sabrosky 1951 (Gaurax) 

Gaurax pallidipes Malloch 1915 (Gaurax) 

Gaurax patelainae Peterson 1967 (Gaurax) 

Gaurax pilosula Becker 1912 (Oscinella) 

Oscinella pilosella Becker 1912 Missp. 

Gaurax pseudostigma Johnson 1913 (Gaurax) 

Gaurax flavidulus Malloch 1915 Syn. 

Gaurax interruptus Malloch 1915 Homo. 

Gaurax semivittatus Sabrosky 1951 (Gaurax) 

Gaurax shannoni Sabrosky 1951 (Gaurax) 

Gaurax splendidus Malloch 1915 (Gaurax) 

Gaurax tripus Sabrosky 1951 (Gaurax) 

Gaurax varihalterata Malloch 1913 (Botanobia) 

Gaurax vittipes Sabrosky 1951 (Gaurax) 

Hapleginella Duda 1933 

Hapleginella conicola Greene 1918 (Madiza) 

Hippelates Loew 1863 

Siphomyia Williston 1896 Syn. 

Palaeoconioscinella Duda 1930 


123

124 

Hippelates dorsalis Loew 1869 (Hippelates) 

Hippelates impressus Becker 1912 (Hippelates) 

Hippelates nobilis Loew 1863 (Hippelates) 

Hippelates plebejus Loew 1863 (Hippelates) 

Hippelates proboscideus Williston 1896 (Hippelates) 

Hippelates saundersi Kumm 1936 (Hippelates) 

Homaluroides Sabrosky 1980 

Homaluroides abdominalis Coquillett 1895 (Chlorops) 

Homaluroides distichliae Malloch 1918 (Anthracophaga) 

Homaluroides graminea Coquillett 1898 (Chlorops) 

Homaluroides ingratus Williston 1893 (Chlorops) 

Anthracophaga interrupta Becker 1912 Syn. 

Chlorops fossae Becker 1912 Syn. 

Homaluroides mellea Loew 1872 (Chlorops) 

Homaluroides pilosulus Becker 1912 (Chlorops) 

Chlorops horrida Becker 1912 Homo. 

Homaluroides quinquepunctata Loew 1863 (Chlorops) 

Homaluroides surda Curran 1930 (Chlorops) 

Incertella Sabrosky 1980 

Incertella bispina Malloch 1918 (Botanobia) 

Incertella dorsata Loew 1872 (Oscinis) 

Oscinis dorsalis Loew 1869 Homo. 

Botanobia spiniger Malloch 1918 Syn. 

Incertella incerta Becker 1912 (Oscinella) 

Oscinella opacifrons Aldrich 1936 Unav. 

Incertella infesta Becker 1912 (Oscinella) 

Incertella insularis Malloch 1914 (Botanobia) 

Incertella minor Adams 1905 (Oscinis) 

Incertella ovalis Adams 1905 (Oscinis) 

Botanobia marginalis Malloch 1914 Syn. 

Botanobia proxima Malloch 1914 Syn. 

Lasiopleura Becker 1910 

Parahippelates Becker 1911 Syn. 

Pseudohippelates Malloch 1913 Syn. 

Liomochaeta Duda 1934 Syn. 

Oscinelloides Malloch 1940 Syn. 

Lasiopleura barberi Sabrosky 1951 (Lasiopleura) 

Lasiopleura capax Coquillett 1898 (Hippelates) 

Lasiopleura grisea Malloch 1934 (Lasiopleura) 

Lasiopleura hirta Loew 1863 (Oscinis) 

Lasiopleura hirtoides Sabrosky 1951 (Lasiopleura) 

Lasiopleura itascae Sabrosky 1951 (Lasiopleura) 

Lasiopleura shewelli Sabrosky 1951 (Lasiopleura) 

Lasiopleura willistoni Sabrosky 1951 (Lasiopleura) 

Lasiosina Becker 1910 

Euchlorops Malloch 1913 Syn. 

Lasiosina approximatonervis Zetterstedt 1848 (Oscinis) 

Chlorops microcera Loew 1872 Syn. 

Chlorops parva Adams 1903 Syn. 

Leptocera transversalis Malloch 1919 Syn. 

Lasiosina canadensis Aldrich 1918 (Lasiosina) 

Lasiosina similis Malloch 1913 (Euchlorops) 

Lasiosina vittata Malloch 1913 (Euchlorops) 

Liohippelates Duda 1929 

Stenprosopon Duda 1930 Syn. 

Statriomicroneurum Duda 1933 

Liohippelates apicatus Malloch 1913 (Hippelates) 



Liohippelates bicolor Coquillett 1898 (Hippelates) 

Liohippelates bishoppi Sabrosky 1941 (Hippelates) 

Liohippelates collusor Townsend 1895 (Oscinis) 

Liohippelates flaviceps Loew 1863 (Oscinis) 

Liohippelates flavipes Loew 1866 (Hippelates) 

Liohippelates pallipes Loew 1863 (Oscinis) 

Hippelates partitus Becker 1912 Syn. 

Hippelates nitidifrons Malloch 1913 Syn. 

Liohippelates peruanus Becker 1912 (Hippelates) 

Liohippelates pusio Loew 1872 (Hippelates) 

Hippelates splendens Adams 1904 Syn. 

Hippelates lituratus Becker 1912 Syn. 

Liohippelates robertsoni Sabrosky 1941 (Hippelates) 

Lipara Meigen 1830 

Gymnopoda Macquart 1835 

Lipara lucens Meigen 1830 (Lipara) 

Malloewia Sabrosky 1980 

Malloewia abdominalis Becker 1912 (Siphonella) 

Oscinella bifurca Becker 1912 Syn. 

Malloewia aequa Becker 1912 (Siphonella) 

Malloewia diabolus Becker 1912 (Siphonella) 

Malloewia excipiens Becker 1912 (Siphonella) 

Malloewia extrema Becker 1912 (Siphonella) 

Malloewia neglecta Becker 1912 (Siphonella) 

Malloewia nigripalpis Malloch 1913 (Madiza) 

Malloewia setulosa Malloch 1918 (Madiza) 

Melanochaeta Bezzi 1906 

Pachychaeta Bezzi 1895 Homo. 

Pachychaetina Hendel 1907 Syn. 

Lasiochaeta Corti 1909 Syn. 

Melanochaeta eunota Loew 1872 (Crassiseta) 

Melanochaeta kaw Sabrosky 1948 (Melanochaeta) 

Melanochaeta melampus Becker 1912 (Melanochaeta) 

Meromyza Meigen 1830 

Meromyia Zetterstedt 1860 Emend. 

Aschabadicola Frey 1921 Syn. 

Nippomera Fedoseeva and Nartshuk 1983 Syn. 

Meromyza affinis Fedoseeva 1971 (Meromyza) 

Meromyza americana Fitch 1856 (Meromyza) 

Meromyza flavipalpis Malloch 1914 Syn. 

Meromyza arizonica Fedoseeva 1971 (Meromyza) 

Meromyza bipunctata Fedoseeva 1971 (Meromyza) 

Meromyza canadensis Fedoseeva 1971 (Meromyza) 

Meromyza columbi Fedoseeva 1971 (Meromyza) 

Meromyza communis Fedoseeva 1971 (Meromyza) 

Meromyza frontosa Fedoseeva 1971 (Meromyza) 

Meromyza laurelae Fedoseeva 1971 (Meromyza) 

Meromyza modesta Fedoseeva 1978 (Meromyza) 

Meromyza mutabilis Fedoseeva 1971 (Meromyza) 

Meromyza pilosa Fedoseeva 1971 (Meromyza) 

Meromyza pratorum Meigen 1830 (Meromyza) 

Meromyza lineola Curran 1923 Syn. 

Meromyza rara Fedoseeva 1971 (Meromyza) 

Meromyza sabroskyi Fedoseeva 1971 (Meromyza) 

Meromyza saltatrix Linnaeus 1761 (Musca) 

Meromyza nigriventris Macquart 1835 Syn. 

Meromyza punctifer Becker 1912 Syn. 

Meromyza marginata Becker 1912 Syn. 

Meromyza truncata Fedoseeva 1971 (Meromyza) 

Meromyza vladimirovae Fedoseeva 1978 (Meromyza)

Monochaetoscinella Duda 1930 

Monochaetoscinella anonyma Williston 1896 (Oscinis) 

Monochaetoscinella nigricornis Loew 1863 (Crassiseta) 

Neodiplotoxa Malloch 1914 

Neodiplotoxa nigricans Loew 1872 (Chlorops) 

Neodiplotoxa pulchripes Loew 1872 (Chlorops) 

Neoscinella Sabrosky 1980 

Neoscinella fuscipalpis Becker 1912 (Oscinella) 

Neoscinella gigas Sabrosky 1940 (Oscinella) 

Neoscinella lugubria Sabrosky 1940 (Oscinella) 

Olcella Enderlein 1911 

Olcella cinerea Loew 1863 (Siphonella) 

Madiza trivittata Sabrosky 1935 Syn. 

Olcella difficilis Becker 1912 (Oscinella) 

Oscinella apparens Becker 1912 Syn. 

Olcella finalis Becker 1912 (Siphonella) 

Olcella parva Adams 1904 (Siphonella) 

Olcella projecta Malloch 1913 (Madiza) 

Olcella provocans Becker 1912 (Siphonella) 

Olcella punctifrons Becker 1912 (Siphonella) 

Olcella pygmaea Becker 1912 (Siphonella) 

Olcella quadrivittata Sabrosky 1935 (Madiza) 

Olcella submarginalis Sabrosky 1935 (Madiza) 

Olcella trigramma Loew 1863 (Oscinis) 

Siphonella quinquelineata Adams 1904 Syn. 

Opetiophora Loew 1872 

Opetiophora straminea Loew 1872 (Opetiophora) 

Oscinella Becker 1909 

Cyclocerculla Beschovski 1978 Syn. 

Microcercis Beschovski 1978 Syn. 

Oscinella frit Linnaeus 1758 (Musca) 

Oscinis nigra Tucker 1908 Syn. 

Oscinella grandissima Sabrosky 1940 (Oscinella) 

Oscinella nitidissima Meigen 1838 (Chlorops) 

Oscinella painteri Sabrosky 1940 (Oscinella) 

Oscinisoma Lioy 1864 

Oscinosoma Enderlein 1911 Emend. 

Oscinisoma aliena Becker 1912 (Elachiptera) 

Oscinoides Malloch 1916 

Oscinoides arpidia Malloch 1916 (Oscinoides) 

Oscinoides ater Malloch 1918 Syn. 

Oscinoides elegans Malloch 1918 Syn. 

Oscinoides humeralis Malloch 1918 Syn. 

Gaurax dorri Johnson 1925 Syn. 

Parectecephala Becker 1910 

Parectecephala aristalis Coquillett 1898 (Chlorops) 

Parectecephala dissimilis Malloch 1914 (Parectecephala) 

Parectecephala eucera Loew 1863 (Chlorops) 

Parectecephala maculiceps Becker 1912 (Parectecephala) 

Parectecephala maculosa Loew 1872 (Chlorops) 


Parectecephala sanguinolenta Loew 1863 (Chlorops) 

Pseudogaurax Malloch 1915 

Mimogaurax Hall 1937 Syn. 

Pseudogaurax anchora Loew 1866 (Gaurax) 

Elachiptera dispar Williston 1896 Syn. 

Pseudogaurax floridensis Sabrosky 1950 (Pseudogaurax) 

Pseudogaurax signata Loew 1876 (Gaurax) 

Gaurax araneae Coquillett 1896 Syn. 

Gaurax mallochi Duda 1930 Syn. 

Rhodesiella Adams 1905 

Macrostyla Lioy 1864 Homo. 

Meroscinis Meijere 1906 Syn. 

Prionoscelus Becker 1911 Syn. 

Lonchonotes Lamb 1918 Syn. 

Euthyridium Frey 1923 Syn. 

Tachinoceras Meijere 1924 Syn. 

Aspistyla Duda 1933 Syn. 

Rhodesiella brimleyi Sabrosky 1947 (Rhodesiella) 

Rhopalopterum Duda 1929 

Rhopalopterum atriceps Loew 1863 (Oscinis) 

Rhopalopterum beameri Sabrosky 1940 (Oscinella) 

Rhopalopterum carbonaria Loew 1869 (Oscinis) 

Botanobia salinaria Curran 1927 Syn. 

Rhopalopterum criddlei Aldrich 1918 (Oscinis) 

Rhopalopterum limitata Becker 1912 (Oscinella) 

Rhopalopterum luteiceps Sabrosky 1940 (Oscinella) 

Rhopalopterum nudiuscula Loew 1863 (Oscinis) 

Rhopalopterum soror Macquart 1851 (Chlorops) 

Oscinis tibialis Fitch 1856 Syn. 

Oscinis variabilis Loew 1863 Syn. 

Botanobia ainsliei Curran 1927 Syn. 

Rhopalopterum umbrosa Loew 1863 (Oscinis) 

Sacatonia Sabrosky 1967 

Sacatonia graminivora Sabrosky 1967 (Sacatonia) 

Siphonella Macquart 1835 

Syphonella Perris 1839 Emend. 

Eurinella Meunier 1893 Syn. 

Siphonella oscinina Fallen 1820 (Madiza) 

Speccafrons Sabrosky 1980 

Speccafrons mallochi Sabrosky 1938 (Oscinella) 

Botanobia halterata Malloch 1913 Homo. 

Stenoscinis Malloch 1918 

Stenoscinis adachiae Sabrosky 1961 (Stenoscinis) 

Stenoscinis longipes Loew 1863 (Oscinis) 


125

126 

Thaumatomyia Zenker 1833 

Chloropisca Loew 1866 Syn. 

Pseudochlorops Malloch 1914 Syn. 

Thaumatomyia annulata Walker 1849 (Chlorops) 

Chlorops variceps Loew 1863 Syn. 

Chloropisca prolifica Osten Sacken 1888 Syn. 

Chloropisca marianapolitana Ouellet 1934 Syn. 

Thaumatomyia apache Sabrosky 1943 (Thaumatomyia) 

Thaumatomyia appropinqua Adams 1903 (Chlorops) 

Thaumatomyia bistriata Walker 1849 (Chlorops) 

Chloropisca clypeata Malloch 1914 Syn. 

Thaumatomyia glabra Meigen 1830 (Chlorops) 

Chlorops assimilis Macquart 1851 Syn. 

Siphonella obesa Fitch 1856 Syn. 

Chlorops trivialis Loew 1863 Syn. 

Chlorops hortensis Fitch 1872 Syn. 

Chlorops halteralis Adams 1903 Syn. 

Thaumatomyia grata Loew 1863 (Chlorops) 

Thaumatomyia obtusa Malloch 1914 (Chloropisca) 

Thaumatomyia parviceps Malloch 1915 (Chloropisca) 

Thaumatomyia pulla Adams 1904 (Chlorops) 

Chloropisca monticola Becker 1912 Syn. 

Chloropisca punctum Becker 1912 Syn. 

Chlorops integra Becker 1912 Syn. 

Thaumatomyia pullipes Coquillett 1898 (Chlorops) 

Thaumatomyia rubida Coquillett 1898 (Chlorops) 

Thaumatomyia rubrivittata Sabrosky 1943 (Thaumatomyia) 

Thaumatomyia trifasciata Zetterstedt 1848 (Oscinis) 

Tricimba Lioy 1864 

Tricymba Marschall 1873 Emend. 

Notanaulax Becker 1903 Syn. 

Euhippelates Malloch 1925 Syn. 

Hammaspis Duda 1930 Syn. 

Eutricimba Malloch 1931 Syn. 

Echimba Duda 1935 Syn. 

Syphonerina Seguy 1938 Syn. 

Nartshukiella Beschovski 1981 Syn. 

Schumanniella Beschovski 1981 Syn. 

Tricimba brunnicollis Becker 1912 (Notonaulax) 

Tricimba cincta Meigen 1830 (Chlorops) 

Tricimba lineella Fallen 1820 (Oscinis) 

Tricimba melancholica Becker 1912 (Oscinella) 

Tricimba occidentalis Sabrosky 1938 (Tricimba) 

Tricimba spinigera Malloch 1913 (Tricimba) 

Tricimba trisulcata Adams 1905 (Oscinis) 

Trigonomma Enderlein 1911 

Beckerella Enderlein 1911 Syn. 

Trigonomma fossulata Loew 1863 (Chlorops) 

Chloropisca atra Curran 1926 Syn. 

CHYROMYIDAE 

Aphaniosoma Becker 1903 

Aphaniosoma aldrichi Wheeler 1961 (Aphaniosoma) 

Aphaniosoma frontatum Wheeler 1961 (Aphaniosoma) 

Aphaniosoma quadrivittatum Malloch 1915 (Aphaniosoma) 


DIPTERA: CHLOROPIDAE, CHYROMYIDAE, CLUSIIDAE 

Chyromya Robineau-Desvoidy 1830 

Lisella Robineau-Desvoidy 1830 Syn. 

Scyphella Robineau-Desvoidy 1830 Syn. 

Thyrimyza Zetterstedt 1847 Syn. 

Chyromya femorella Fallen 1820 (Sapromyza) 

Chyromya flava Linnaeus 1758 (Musca) 

Musca ochrapesus Rathvon 1855 Syn. 

Chyromya oppidana Scopoli 1763 (Musca) 

Gymnochiromyia Hendel 1933 

Gymnochiromyia concolor Malloch 1914 (Chyromya) 

Gymnochiromyia minima Becker 1904 (Peletophila) 

Gymnochiromyia nigrimana Malloch 1914 (Chyromya) 

CLUSIIDAE 

Clusia Haliday 1838 

Macrochira Zetterstedt 1838 Syn. 

Peratochetus Rondni 1856 Unav. 

Stomphastica Loew 1864 Syn. 

Peratochetus Rondani 1874 Syn. 

Clusia czernyi Johnson 1913 (Clusia) 

Clusia lateralis Walker 1849 (Helomyza) 

Tetanura pallida Harris 1835 Unav. 

Heteroneura spectabilis Loew 1860 Syn. 

Clusia occidentalis Malloch 1918 (Clusia) 

Clusiodes Coquillett 1904 

Heteroneura Fallen 1823 Homo. 

Columbiella Malloch 1922 Syn. 

Clusiaria Malloch 1922 Syn. 

Clusiodes albimanus Meigen 1830 (Heteroneura) 

Clusiodes americanus Malloch 1922 (Clusiodes) 

Clusiodes apicalis Zetterstedt 1848 (Heteroneura) 

Clusiodes apiculatus Malloch 1922 (Clusiodes) 

Clusiodes ater Melander and Argo 1924 (Clusiodes) 

Clusiodes geomyzinus Fallen 1823 (Heteroneura) 

Clusiodes johnsoni Malloch 1922 (Clusiodes) 

Clusiodes nigripalpis Malloch 1922 Syn. 

Clusiodes melanostoma Loew 1864 (Heteroneura) 

Heteroneura flavifascies Coquillett 1897 Unav. 

Clusiodes duplicata Malloch 1922 Syn. 

Clusiodes niger Melander and Argo 1924 (Clusiodes) 

Clusiodes nitidus Melander and Argo 1924 (Clusiodes) 

Clusiodes scutellatus Melander and Argo 1924 Syn. 

Clusiodes orbitalis Malloch 1922 (Clusiodes) 

Clusiodes pictipes Zetterstedt 1855 (Heteroneura) 

Clusiodes ruficollis Meigen 1830 (Heteroneura) 

Clusiodes terminalis Melander and Argo 1924 (Clusiodes) 

Heteromeringia Czerny 1903 

Heteromeringia annulipes Johnson 1913 (Heteromeringia) 

Heteromeringia flavipes Williston 1896 (Heteroneura) 

Heteromeringia nitida Johnson 1913 (Heteromeringia) 

Heteromeringia nigripes Melander and Argo 1924 Syn.

Sobarocephala Czerny 1903 

Chaetoclusia Coquillett 1904 Syn. 

Sobarocephala affinis Johnson 1913 (Chaetoclusia) 

Sobarocephala atricornis Sabrosky and Steyskal 1974 

(Sobarocephala) 

Sobarocephala atrifacies Sabrosky and Steyskal 1974 


Sobarocephala cruciger Sabrosky and Steyskal 1974 


Sobarocephala dreisbachi Sabrosky and Steyskal 1974 


Sobarocephala flava Melander and Argo 1924 (Sobarocephala) 

Sobarocephala populi Steyskal 1951 Syn. 

Sobarocephala flaviseta Johnson 1913 (Heteromeringia) 

Heteromeringia convergens Malloch 1922 Syn. 

Sobarocephala interrupta Sabrosky and Steyskal 1974 


Sobarocephala lachnosternum Melander and Argo 1924 


Sobarocephala latifacies Sabrosky and Steyskal 1974 


Sobarocephala latifrons Loew 1860 (Heteroneura) 

Sobarocephala muesebecki Sabrosky and Steyskal 1974 


Sobarocephala quadrimaculata Soos 1963 (Sobarocephala) 

Sobarocephala nitida Soos 1963 Syn. 

Sobarocephala setipes Melander and Argo 1924 (Sobarocephala) 

Sobarocephala testacea Soos 1964 (Sobarocephala) 

Sobarocephala texensis Sabrosky and Steyskal 1974 


COELOPIDAE 

Coelopa Meigen 1830 

Fucomyia Haliday 1838 Syn. 

Neocoelopa Malloch 1933 Syn. 

Coelopa frigida Fabricius 1805 (Musca) 

Coelopa gravis Haliday 1833 Syn. 

Coelopa parvula Haliday 1833 Syn. 

Coelopa nitidula Zetterstedt 1847 Syn. 

Coelopa eximia Stenhammar 1854 Syn. 

Coelopa nebularum Aldrich 1929 (Coelopa) 

Coelopa stejnegeri Aldrich 1929 (Coelopa) 

Coelopa vanduzeei Cresson 1914 (Coelopa) 

Coelopina Malloch 1933 

Coelopina anomala Cole 1923 (Coelopa) 

CONOPIDAE 

Conops Linnaeus 1758 

Conopaejus Rondani 1845 Syn. 

Conopilla Rondani 1856 Syn. 

Sphixosoma Rondani 1857 Syn. 

Bombidia Lioy 1864 Syn. 


Asiconops Chen 1939 Syn. 

Conops bermudensis Parsons 1940 (Conops) 

Conops flaviceps Macquart 1843 (Conops) 

Dalmannia Robineau-Desvoidy 1830 

Stachynia Macquart 1835 Syn. 

Dalmannia blaisdelli Cresson 1919 (Dalmannia) 

Dalmannia heterotricha Bohart 1938 (Dalmannia) 

Dalmannia nigriceps Loew 1866 (Dalmannia) 

Dalmannia pacifica Banks 1916 (Dalmannia) 

Dalmannia hirsuta Van Duzee 1927 Syn. 

Dalmannia picta Williston 1883 (Dalmannia) 

Dalmannia vitiosa Coquillett 1892 (Dalmannia) 

Myopa Fabricius 1775 

Pictinia Robineau-Desvoidy 1853 Syn. 

Myopella Robineau-Desvoidy 1853 Syn. 

Phorosia Robineau-Desvoidy 1853 Syn. 

Fairmairia Robineau-Desvoidy 1853 Syn. 

Lonchopalpus Robineau-Desvoidy 1853 Syn. 

Purpurella Robineau-Desvoidy 1853 Syn. 

Myopina Robineau-Desvoidy 1853 Syn. 

Haustellia Robineau-Desvoidy 1853 Syn. 

Gonirhyncus Rondani 1856 Syn. 

Glossigona Rondani 1856 Syn. 

Gonirhynchus Rondani 1857 Syn. 

Arpagita Lioy 1864 Syn. 

Myopa bohartorum Camras 1953 (Myopa) 

Myopa castanea Bigot 1887 (Gonirhyncus) 

Myopa clausa Loew 1866 (Myopa) 

Myopa curticornis Kröber 1916 (Myopa) 

Myopa flavopilosa Kröber 1916 (Myopa) 

Myopa varians Banks 1916 Syn. 

Myopa longipilis Banks 1916 (Myopa) 

Myopa melanderi Banks 1916 (Myopa) 

Myopa perplexa Camras 1953 (Myopa) 

Myopa plebeia Williston 1885 (Myopa) 

Myopa rubida Bigot 1887 (Glossigona) 

Myopa aperta Röder 1889 Syn. 

Myopa seminuda Banks 1916 Syn. 

Myopa vesiculosa Say 1823 (Myopa) 

Myopa apicalis Walker 1849 Syn. 

Myopa conjuncta Thomson 1869 Syn. 

Glossigona maculifrons Bigot 1887 Syn. 

Myopa utahensis Stains and Knowlton 1939 Syn. 

Myopa vicaria Walker 1849 (Myopa) 

Myopa pilosa Williston 1885 Syn. 

Myopa virginica Banks 1916 (Myopa) 

Myopa willistoni Banks 1916 (Myopa) 

Myopa pictipennis Williston 1885 Homo. 

Physocephala Schiner 1861 

Physocephala burgessi Williston 1882 (Conops) 

Physocephala brevirostris Van Duzee 1927 Syn. 

Physocephala floridana Camras 1957 (Physocephala) 

Physocephala furcillata Williston 1882 (Conops) 

Physocephala lucida Van Duzee 1931 Syn. 

Physocephala marginata Say 1823 (Conops) 

Physocephala dakotensis Van Duzee 1934 Syn. 

Physocephala stylifer Van Duzee 1934 Syn. 

Physocephala sagittaria Say 1823 (Conops) 

Conops aethiops Walker 1849 Syn. 

Conops dimidiata Walker 1852 Syn. 

DIPTERA: CLUSIIDAE, COELOPIDAE, CONOPIDAE 

127

128 

Conops genualis Loew 1853 Syn. 

Conops catanoptera Loew 1853 Syn. 

Physocephala ruficornis Van Duzee 1934 Syn. 

Physocephala texana Williston 1882 (Conops) 

Conops affinis Williston 1882 Syn. 

Conops ochreiceps Bigot 1887 Syn. 

Physocephala humeralis Van Duzee 1927 Syn. 

Physocephala simulans Van Duzee 1927 Syn. 

Physocephala aurifacies Van Duzee 1927 Syn. 

Physocephala buccalis Van Duzee 1927 Syn. 

Physocephala rubida Van Duzee 1934 Syn. 

Physocephala tibialis Say 1829 (Conops) 

Conops nigricornis Wiedemann 1830 Syn. 

Conops fulvipennis Macquart 1843 Syn. 

Conops lugubris Macquart 1843 Syn. 

Physoconops Szilady 1926 

Pachyconops Camras 1955 Syn. 

Gyroconops Camras 1955 Syn. 

Physoconops analis Fabricius 1805 (Conops) 

Conops angustifrons Williston 1892 Syn. 

Physoconops brachyrhynchus Macquart 1843 (Conops) 

Conops xanthopareus Williston 1882 Syn. 

Physoconops xanthostomus Banks 1916 Missp. 

Conops fenestratus Kröber 1915 Syn. 

Physoconops bulbirostris Loew 1853 (Conops) 

Physoconops discalis Williston 1892 (Conops) 

Conops semifuscus Banks 1916 Syn. 

Physoconops excisus Wiedemann 1830 (Conops) 

Conops sugens Wiedemann 1830 Syn. 

Physoconops floridanus Camras 1955 (Physoconops) 

Physoconops fronto Williston 1885 (Conops) 

Conops pulchellus Kröber 1915 Syn. 

Conops argentifacies Van Duzee 1927 Syn. 

Conops fraterculus Van Duzee 1927 Syn. 

Conops rubicundulus Van Duzee 1927 Syn. 

Physoconops gracilis Williston 1885 (Conops) 

Physoconops nigrimanus Bigot 1887 (Conops) 

Conops striatifrons Kröber 1915 Syn. 

Conops limuva Brimley 1927 Syn. 

Physoconops obscuripennis Williston 1882 (Conops) 

Conops foxi Van Duzee 1927 Syn. 

Physoconops sylvosus Williston 1882 (Conops) 

Conops arizonicus Banks 1916 Syn. 

Physoconops townsendi Camras 1955 (Physoconops) 

Conops auratus Townsend 1901 Homo. 

Robertsonomyia Malloch 1919 

Robertsonomyia palpalis Robertson 1901 (Zodion) 

Zodion scapularis Adams 1903 Syn. 

Robertsonomyia parva Adams 1903 (Zodion) 

Zodion lovetti Van Duzee 1934 Syn. 

Stylogaster Macquart 1835 

Stylomyia Westwood 1852 Syn. 

Stylogaster biannulata Say 1823 (Myopa) 

Stylomyia confusa Westwood 1852 Syn. 

Stylogaster neglecta Williston 1883 (Stylogaster) 

Thecophora Rondani 1845 

Occemya Robineau-Desvoidy 1853 Syn. 

Thecophora abbreviata Loew 1866 (Oncomyia) 

Oncomyia melanopoda Williston 1885 Syn. 

Thecophora longicornis Say 1823 (Myopa) 


DIPTERA: CONOPIDAE, CORETHRELLIDAE 

Oncomyia infuscipes Van Duzee 1927 Syn. 

Thecophora luteipes Camras 1945 (Occemya) 

Thecophora modesta Williston 1883 (Oncomyia) 

Zodion basalis Van Duzee 1927 Syn. 

Thecophora nigra Van Duzee 1927 (Oncomyia) 

Thecophora nigripes Camras 1945 (Occemya) 

Thecophora occidensis Walker 1849 (Zodion) 

Oncomyia loraria Loew 1866 Syn. 

Oncomyia baroni Williston 1883 Syn. 

Oncomyia brevirostris Van Duzee 1927 Syn. 

Oncomyia aequalis Van Duzee 1927 Syn. 

Oncomyia terminalis Van Duzee 1927 Syn. 

Zodion bimacula Curran 1933 Syn. 

Oncomyia frontalis Van Duzee 1934 Syn. 

Thecophora propinqua Adams 1903 (Oncomyia) 

Zodion angusticornis Van Duzee 1927 Syn. 

Oncomyia angusticornis Van Duzee 1927 Homo. 

Oncomyia longipalpis Van Duzee 1934 Syn. 

Zodion Latreille 1796 

Zodion abitus Adams 1903 (Zodion) 

Zodion bicolor Adams 1903 Syn. 

Zodion americanum Wiedemann 1830 (Zodion) 

Zodion nanellum Loew 1866 Syn. 

Zodion pygmaeum Williston 1885 Syn. 

Zodion albifacies Van Duzee 1927 Syn. 

Zodion anale Kröber 1915 (Zodion) 

Zodion auricaudatum Williston 1892 (Zodion) 

Zodion albonotatum Townsend 1897 Syn. 

Zodion californicum Camras 1954 (Zodion) 

Zodion cinereiventre Van Duzee 1927 (Zodion) 

Zodion cyanescens Camras 1943 (Zodion) 

Zodion fulvifrons Say 1823 (Zodion) 

Zodion abdominalis Say 1823 Syn. 

Myopa rubrifrons Robineau-Desvoidy 1830 Syn. 

Myopa bistria Walker 1849 Syn. 

Zodion lativentre Graenicher 1910 Syn. 

Zodion obscurum Banks 1916 Syn. 

Zodion sayi Banks 1916 Syn. 

Zodion reclusum Banks 1916 Syn. 

Zodion bilineata Van Duzee 1927 Syn. 

Zodion intermedium Banks 1916 (Zodion) 

Zodion occidentale Banks 1916 Syn. 

Zodion nigrifrons Kröber 1915 (Zodion) 

Zodion hirtipes Van Duzee 1927 Syn. 

Zodion obliquefasciata Macquart 1846 (Myopa) 

Zodion leucostoma Williston 1885 Syn. 

Myopa tectura Adams 1903 Syn. 

Zodion perlongum Coquillett 1902 (Zodion) 

Zodion pictulum Williston 1885 (Zodion) 

Zodion triste Bigot 1887 (Zodion) 

CORETHRELLIDAE 

Corethrella Coquillett 1902 

Ramica Annandale 1911 Syn. 

Lutzomiops Lane 1942 Syn. 

Notocorethrella Belkin 1968 Syn. 

Corethrella appendiculata Grabham 1906 (Corethrella)

Corethrella arborealis Shannon and Del Ponte 1928 Syn. 

Corethrella confusa Lane 1939 Syn. 

Corethrella brakeleyi Coquillett 1902 (Corethra) 

Corethrella kerrvillensis Stone 1965 (Lutzomiops) 

Corethrella laneana Vargas 1946 (Corethrella) 

Corethrella wirthi Stone 1968 (Corethrella) 

CRYPTOCHETIDAE 

Cryptochetum Rondani 1875 

Lestophonus Williston 1888 Syn. 

Cryptochetum iceryae Williston 1888 (Lestophonus) 

CULICIDAE 

Aedes Meigen 1818 

Ochlerotatus Lynch Arribalzaga 1891 Syn. 

Taeniorhynchus Lynch Arribalzaga 1891 Unav. 

Stegomyia Theobald 1901 Syn. 

Mucidus Theobald 1901 Syn. 

Finlaya Theobald 1903 Syn. 

Aedimorphus Theobald 1903 Syn. 

Catageiomyia Theobald 1903 Syn. 

Howardina Theobald 1903 Syn. 

Macleaya Theobald 1903 Syn. 

Gilesia Theobald 1903 Syn. 

Acartomyia Theobald 1903 Syn. 

Skusea Theobald 1903 Syn. 

Verrallina Theobald 1903 Syn. 

Culicada Felt 1904 Syn. 

Ecculex Felt 1904 Syn. 

Danielsia Theobald 1904 Syn. 

Hulecoetomyia Theobald 1904 Syn. 

Culicelsa Felt 1904 Syn. 

Protoculex Felt 1904 Syn. 

Gualteria Lutz 1904 Homo. 

Scutomyia Theobald 1904 Syn. 

Pseudoculex Dyar 1905 Syn. 

Reedomyia Ludlow 1905 Syn. 

Lepidotomyia Theobald 1905 Syn. 

Pecomyia Theobald 1905 Syn. 

Polyleptiomyia Theobald 1905 Syn. 

Pseudograbhamia Theobald 1905 Syn. 

Popea Ludlow 1905 Syn. 

Phagomyia Theobald 1905 Syn. 

Leptosomatomyia Theobald 1905 Syn. 

Chrysoconops Goeldi 1905 Syn. 

Gymnometopa Coquillett 1906 Syn. 

Lepidoplatys Coquillett 1906 Syn. 

Quasistegomyia Theobald 1906 Syn. 

Protomacleaya Theobald 1907 Syn. 

Duttonia Newstead 1907 Syn. 

Pardomyia Theobald 1907 Syn. 

Neomacleaya Theobald 1907 Syn. 

Neomelaniconion Newstead 1907 Syn. 

Banksinella Theobald 1907 Syn. 

Pseudohowardina Theobald 1907 Syn. 

Pseudoskusea Theobald 1907 Syn. 


Mimeteculex Theobald 1908 Syn. 

Geitonomyia Leicester 1908 Syn. 

Ekrinomyia Leicester 1908 Syn. 

Aioretomyia Leicester 1908 Syn. 

Pseudostegomyia Ludlow 1908 Syn. 

Bathosomyia Theobald 1909 Syn. 

Myxosquamus Theobald 1909 Syn. 

Neopecomyia Theobald 1909 Syn. 

Stenoscutus Theobald 1909 Syn. 

Chaetocruiomyia Theobald 1910 Syn. 

Pseudocarrollia Theobald 1910 Syn. 

Molpemyia Theobald 1910 Syn. 

Kingia Theobald 1910 Homo. 

Leslieomyia Christophers 1911 Syn. 

Diceromyia Theobald 1911 Syn. 

Andersonia Strickland 1911 Syn. 

Calomyia Taylor 1914 Homo. 

Caenocephalus Taylor 1914 Homo. 

Christophersiomyia Barraud 1923 Syn. 

Aniella Enderlein 1923 Syn. 

Catatassomyia Dyar and Shannon 1925 Syn. 

Cancraedes Edwards 1929 Syn. 

Dendroskusea Edwards 1929 Syn. 

Geoskusea Edwards 1929 Syn. 

Rhinoskusea Edwards 1929 Syn. 

Hyparctius Martini 1930 Syn. 

Feltianus Martini 1930 Syn. 

Hyparctius Martini 1930 Syn. 

Dunnius Edwards 1930 Homo. 

Indusius Edwards 1934 Syn. 

Paraedes Edwards 1934 Syn. 

Soperia Komp 1936 Syn. 

Kompia Aitken 1941 Syn. 

Levua Stone and Bohart 1944 Syn. 

Downsiomyia Vargas 1950 Syn. 

Kummyia Vargas 1950 Syn. 

Ayurakitia Thurman 1954 Syn. 

Pseudarmigeres Stone and Knight 1956 Syn. 

Alanstonea Mattingly 1960 Syn. 

Edwardseades Belkin 1962 Syn. 

Halaedes Belkin 1962 Syn. 

Lorrainea Belkin 1962 Syn. 

Nothoskusea Dumbleton 1962 Syn. 

Huaedes Huang 1968 Syn. 

Abraedes Zavortink 1970 Syn. 

Aztecaedes Zavortink 1972 Syn. 

Bothaella Reinert 1973 Syn. 

Rusticoidus Shevchenko and Prudkina 1973 Syn. 

Isoaedes Reinert 1979 Syn. 

Belkinius Reinert 1982 Syn. 

Albuginosus Reinert 1987 Syn. 

Kenknightia Reinert 1990 Syn. 

Sinoaedes Gong and Lu 1991 Syn. 

Aedes aboriginis Dyar 1917 (Aedes) 

Aedes abserratus Felt and Young 1904 (Culex) 

Aedes centrotus Howard, Dyar, and Knab 1917 Syn. 

Aedes dysanor Dyar 1921 Syn. 

Aedes aegypti Linnaeus 1762 (Culex) 

Culex argenteus Poiret 1787 Syn. 

Culex fasciatus Fabricius 1805 Syn. 

Culex calopus Meigen 1818 Syn. 

Culex frater Robineau-Desvoidy 1827 Syn. 

Culex mosquito Robineau-Desvoidy 1827 Syn. 

Culex taeniatus Wiedemann 1828 Syn. 

Culex sugens Wiedemann 1828 Syn. 

DIPTERA: CORETHRELLIDAE, CRYPTOCHETIDAE, CULICIDAE 

129

130 

Culex kounoupi Brulle 1833 Syn. 

Culex toxorhynchus Macquart 1838 Syn. 

Culex annulitarsis Macquart 1846 Syn. 

Culex excitans Walker 1848 Syn. 

Culex viridifrons Walker 1848 Syn. 

Culex inexorabilis Walker 1848 Syn. 

Culex formosus Walker 1848 Syn. 

Culex exagitans Walker 1856 Syn. 

Culex insatiabilis Bigot 1859 Syn. 

Culex bancrofti Skuse 1889 Syn. 

Culex elegans Ficalbi 1890 Syn. 

Culex rossii Giles 1899 Syn. 

Stegomyia luciensis Theobald 1901 Syn. 

Stegomyia nigeria Theobald 1901 Syn. 

Stegomyia queenslandensis Theobald 1901 Syn. 

Stegomyia lamberti Ventrillon 1904 Syn. 

Steomyia canariensis Pittaluga 1905 Syn. 

Stegomyia persistans Banks 1906 Syn. 

Culex albopalposus Becker 1908 Syn. 

Culex angustealatus Becker 1908 Syn. 

Duttonia alboannulis Ludlow 1911 Syn. 

Mimeteomyia pulcherrima Taylor 1919 Syn. 

Stegomyia atritarsis Edwards 1920 Syn. 

Aedes albopictus Skuse 1895 (Culex) 

Stegomyia samarensis Ludlow 1903 Syn. 

Stegomyia nigritia Ludlow 1910 Syn. 

Stegomyia quasinigritia Ludlow 1911 Syn. 

Aedes aloponotum Dyar 1917 (Aedes) 

Aedes atlanticus Dyar and Knab 1906 (Aedes) 

Aedes atropalpus Coquillett 1902 (Culex) 

Aedes aurifer Coquillett 1903 (Culex) 

Aedes bicristatus Thurman and Winkler 1950 (Aedes) 

Aedes bimaculatus Coquillett 1902 (Culex) 

Aedes rozeboomi Vargas 1941 Syn. 

Aedes brelandi Zavortink 1972 (Aedes) 

Aedes burgeri Zavortink 1972 (Aedes) 

Aedes campestris Dyar and Knab 1907 (Aedes) 

Aedes callithotrys Dyar 1920 Syn. 

Aedes canadensis Theobald 1901 (Culex) 

Culex nivitarsis Coquillett 1904 Syn. 

Aedes mathesoni Middlekauff 1944 Syn. 

Aedes cantator Coquillett 1903 (Culex) 

Aedes cataphylla Dyar 1916 (Aedes) 

Aedes prodotes Dyar 1917 Syn. 

Aedes rostochiensis Martini 1920 Syn. 

Aedes pacificensis Hearle 1927 Syn. 

Aedes churchillensis Ellis and Brust 1973 (Aedes) 

Aedes cinereus Meigen 1818 (Aedes) 

Aedes rufus Gimmerthal 1845 Syn. 

Culex nigritulus Zetterstedt 1850 Syn. 

Aedes fuscus Osten Sacken 1877 Syn. 

Aedes leucopygos Eysell 1903 Syn. 

Culex pallidohirta Grossbeck 1905 Syn. 

Aedes hemiteleus Dyar 1924 Syn. 

Aedes clivis Lanzaro and Eldridge 1992 (Aedes) 

Aedes communis De Geer 1776 (Culex) 

Culex fasciatus Meigen 1804 Homo. 

Culex nemorosus Meigen 1818 Syn. 

Culex sylvaticus Meigen 1818 Syn. 

Aedes obscurus Meigen 1830 Syn. 

Culex lazarensis Felt and Young 1904 Syn. 

Culex borealis Ludlow 1911 Syn. 

Culicada diplolineata Schneider 1913 Syn. 

Aedes tahoensis Dyar 1916 Syn. 

Aedes altiusculus Dyar 1917 Syn. 

Ochlerotatus palmeni Edwards 1921 Syn. 

Aedes prolixus Dyar 1922 Syn. 

Aedes decticus Howard, Dyar, and Knab 1917 (Aedes) 

DIPTERA: CULICIDAE 


Aedes pseudodiantaeus Smith 1952 Syn. 

Aedes deserticola Zavortink 1969 (Aedes) 

Aedes diantaeus Howard, Dyar, and Knab 1913 (Aedes) 

Aedes serus Martini 1920 Syn. 

Aedes dorsalis Meigen 1830 (Culex) 

Culex curriei Coquillett 1901 Syn. 

Culex onondagensis Felt 1904 Syn. 

Culex lativittatus Coquillett 1906 Syn. 

Aedes quaylei Dyar and Knab 1906 Syn. 

Grabhamia broquetii Theobald 1913 Syn. 

Aedes grahami Ludlow 1920 Syn. 

Aedes dupreei Coquillett 1904 (Culex) 

Aedes epactius Dyar and Knab 1908 (Aedes) 

Aedes perichares Dyar 1921 Syn. 

Aedes nielseni O'Meara and Craig 1970 Syn. 

Aedes euedes Howard, Dyar, and Knab 1913 (Aedes) 

Aedes barri Rueger 1958 Syn. 

Aedes excrucians Walker 1856 (Culex) 

Culex abfitchii Felt 1904 Syn. 

Culex siphonalis Grossbeck 1904 Syn. 

Aedes sansoni Dyar and Knab 1909 Syn. 

Culicada sourcoufi Theobald 1912 Syn. 

Aedes dytes Martini 1922 Syn. 

Aedes fitchii Felt and Young 1904 (Culex) 

Aedes palustris Dyar 1916 Syn. 

Aedes pricei Dyar 1917 Syn. 

Aedes mimesis Dyar 1917 Syn. 

Aedes flavescens Müller 1764 (Culex) 

Culex fletcheri Coquillett 1902 Syn. 

Aedes fulvus Wiedemann 1828 (Culex) 

Culex flavicosta Walker 1856 Syn. 

Culex pallens Ross 1943 Syn. 

Aedes grossbecki Dyar and Knab 1906 (Aedes) 

Culex sylvicola Grossbeck 1906 Syn. 

Aedes hendersoni Cockerell 1918 (Aedes) 

Aedes hexodontus Dyar 1916 (Culex) 

Aedes masamae Dyar 1920 Syn. 

Aedes cyclocerculus Dyar 1920 Syn. 

Aedes leuconotips Dyar 1920 Syn. 

Aedes labradorensis Dyar and Shannon 1925 Syn. 

Aedes hokkaidensis Tanaka et al. 1979 Syn. 

Aedes impiger Walker 1848 (Culex) 

Aedes nearcticus Dyar 1919 Syn. 

Aedes parvulus Edwards 1921 Syn. 

Aedes daisetsuzanus Tanaka et al. 1979 Syn. 

Aedes implicatus Vockeroth 1954 (Aedes) 

Aedes increpitus Dyar 1916 (Aedes) 

Grabhamia vittata Theobald 1903 Homo. 

Aedes mutatus Dyar 1919 Syn. 

Aedes hewitti Hearle 1923 Syn. 

Aedes infirmatus Dyar and Knab 1906 (Aedes) 

Aedes intrudens Dyar 1919 (Aedes) 

Aedes kompi Vargas and Downs 1950 (Aedes) 

Aedes melanimon Dyar 1924 (Aedes) 

Grabhamia mediolineata Ludlow 1907 Homo. 

Aedes klotsi Matheson 1933 Syn. 

Aedes mercurator Dyar 1920 (Aedes) 

Aedes albertae Dyar 1920 Syn. 

Aedes mitchellae Dyar 1905 (Culex) 

Aedes monticola Belkin and McDonald 1957 (Aedes) 

Aedes muelleri Dyar 1920 (Aedes) 

Aedes iridipennis Dyar 1922 Syn. 

Aedes nevadensis Chapman and Barr 1964 (Aedes) 

Aedes nigripes Zetterstedt 1838 (Culex) 

Aedes innuitus Dyar and Knab 1918 Syn. 

Aedes nigromaculis Ludlow 1906 (Grabhamia) 

Grabhamia grisea Ludlow 1907 Syn. 

Aedes niphadopsis Dyar and Knab 1918 (Aedes)

Aedes pionips Dyar 1919 (Aedes) 

Aedes provocans Walker 1848 (Culex) 

Culex trichurus Dyar 1904 Syn. 

Culex cinereoborealis Felt and Young 1904 Syn. 

Aedes pagetonotum Dyar and Knab 1909 Syn. 

Aedes poliochros Dyar 1919 Syn. 

Aedes pullatus Coquillett 1904 (Culex) 

Aedes acrophilus Dyar 1917 Syn. 

Culex jugorum Villeneuve 1919 Syn. 

Aedes metalepticus Dyar 1920 Syn. 

Aedes gallii Martini 1920 Syn. 

Aedes pearyi Dyar and Shannon 1925 Syn. 

Aedes seguyi Apfelbeck 1929 Syn. 

Aedes punctodes Dyar 1922 (Aedes) 

Aedes punctor Kirby 1837 (Culex) 

Culex implacabilis Walker 1848 Syn. 

Culex fusculus Zetterstedt 1850 Syn. 

Culicelsa auroides Felt 1905 Syn. 

Culex wahlgreni Theobald 1910 Syn. 

Culicada alineata Schneider 1913 Syn. 

Culicauda haplolineata Schneider 1913 Syn. 

Aedes meigenanus Dyar 1921 Syn. 

Aedes purpureipes Aitken 1941 (Aedes) 

Aedes rempeli Vockeroth 1954 (Aedes) 

Aedes riparius Dyar and Knab 1907 (Aedes) 

Aedes semicantans Martini 1920 Syn. 

Aedes ater Gutzevich 1955 Syn. 

Aedes scapularis Rondani 1848 (Culex) 

Ochlerotatus confirmatus Lynch Arribalzaga 1891 Syn. 

Aedes hemisurus Dyar and Knab 1906 Syn. 

Aedes camposanus Dyar 1918 Syn. 

Aedes rhyacophilus Lima 1933 Syn. 

Aedes schizopinax Dyar 1929 (Aedes) 

Aedes sierrensis Ludlow 1905 (Taeniorhynchus) 

Aedes sollicitans Walker 1856 (Culex) 

Culex colon Harris 1835 Unav. 

Aedes spencerii Theobald 1901 (Culex) 

Grabhamia idahoensis Theobald 1903 Syn. 

Aedes squamiger Coquillett 1902 (Culex) 

Grabhamia deniedmannii Ludlow 1904 Syn. 

Aedes sticticus Meigen 1838 (Culex) 

Culex concinnus Stephens 1846 Syn. 

Culex hirsuteron Theobald 1901 Syn. 

Culex sylvae Theobald 1901 Syn. 

Culex pretans Grossbeck 1904 Syn. 

Culex aestivalis Dyar 1904 Syn. 

Aedes aldrichi Dyar and Knab 1908 Syn. 

Aedes gonimus Dyar and Knab 1918 Syn. 

Aedes vinnipegensis Dyar 1919 Syn. 

Culex dorsovittatus Villeneuve 1919 Syn. 

Ochlerotatus lesnei Seguy 1920 Syn. 

Aedes paradiantaeus Apfelbeck 1929 Syn. 

Aedes stimulans Walker 1848 (Culex) 

Culicada subcantans Felt 1905 Syn. 

Aedes classicus Dyar 1920 Syn. 

Aedes mississippii Dyar 1920 Syn. 

Aedes taeniorhynchus Wiedemann 1821 (Culex) 

Culex damnosus Say 1823 Syn. 

Taeniorhynchus niger Giles 1904 Syn. 

Culex portoricensis Ludlow 1905 Syn. 

Aedes epinolus Dyar and Knab 1914 Syn. 

Aedes thelcter Dyar 1918 (Aedes) 

Aedes keyensis Buren 1947 Syn. 

Aedes thibaulti Dyar and Knab 1910 (Aedes) 

Aedes togoi Theobald 1907 (Culiselsa) 

Aedes tormentor Dyar and Knab 1906 (Aedes) 

Aedes tortilis Theobald 1903 (Culex) 

Culex bracteatus Coquillett 1906 Syn. 


Aedes habanicus Dyar and Knab 1906 Syn. 

Aedes balteatus Dyar and Knab 1907 Syn. 

Aedes plutocraticus Dyar and Knab 1907 Syn. 

Aedes virginensis Dyar 1922 Syn. 

Aedes triseriatus Say 1823 (Culex) 

Finlaya nigra Ludlow 1905 Homo. 

Aedes trivittatus Coquillett 1902 (Culex) 

Culex inconspicuus Grossbeck 1904 Syn. 

Aedes varipalpus Coquillett 1902 (Culex) 

Aedes ventrovittis Dyar 1916 (Aedes) 

Aedes fisheri Dyar 1917 Syn. 

Aedes cacothius Dyar 1923 Syn. 

Aedes vexans Meigen 1830 (Culex) 

Culex parvus Macquart 1834 Syn. 

Culex articulatus Rondani 1872 Syn. 

Culex malariae Grassi 1898 Syn. 

Culex sylvestris Theobald 1901 Homo. 

Culex nocturnus Theobald 1903 Syn. 

Culex montcalmi Blanchard 1905 Syn. 

Culex arabiensis Patton 1905 Syn. 

Culicada minuta Theobald 1907 Syn. 

Culicada eruthroscopa Theobald 1910 Syn. 

Culex sudanensis Theobald 1911 Syn. 

Culex niger Theobald 1913 Homo. 

Aedes euochrus Howard, Dyar, and Knab 1917 Syn. 

Culicada nipponii Theobald 2907 Syn. 

Aedes washinoi Lanzaro and Eldridge 1992 (Aedes) 

Aedes zoosophus Dyar and Knab 1918 (Aedes) 

Aedes alleni Turner 1924 Syn. 

Anopheles Meigen 1818 

Cyclolepidopteron Blanchard 1901 Syn. 

Laverania Theobald 1902 Homo. 

Nyssorhynchus Blanchard 1902 Syn. 

Rossia Theobald 1902 Homo. 

Myzorhynchus Blanchard 1902 Syn. 

Grassia Theobald 1902 Homo. 

Howardia Theobald 1902 Homo. 

Cellia Theobald 1902 Syn. 

Mezomyia Blanchard 1902 Syn. 

Pyretophorus Blanchard 1902 Syn. 

Stethomyia Theobald 1902 Syn. 

Christya Theobald 1903 Syn. 

Arribalzagia Theobald 1903 Syn. 

Aldrichia Theobald 1903 Homo. 

Lophoscelomyia Theobald 1904 Syn. 

Lophomyia Giles 1904 Homo. 

Arribalzagaia Blanchard 1905 Emend. 

Kerteszia Theobald 1905 Syn. 

Coelodiazesis Dyar and Knab 1906 Syn. 

Nototricha Coquillett 1906 Syn. 

Feltinella Theobald 1907 Syn. 

Neocellia Theobald 1907 Syn. 

Pseudomyzomyia Theobald 1907 Syn. 

Myzorhynchella Theobald 1907 Syn. 

Manguinhosia Cruz 1907 Syn. 

Calvertia Ludlow 1908 Homo. 

Calvertina Ludlow 1909 Syn. 

Cycloleppteron Theobald 1910 Syn. 

Neostethopheles James 1910 Syn. 

Patagiamyia James 1910 Syn. 

Neomyzomyia Theobald 1910 Syn. 

Aldrichinella Theobald 1910 Syn. 

Nyssomyzomyia James 1910 Syn. 

Christophersia James 1910 Syn. 

Dactylomyia Newstead and Carter 1910 Homo. 

Cyclophorus Eysell 1912 Homo. 

Proterorhynchus Brethes 1912 Syn. 


131

132 



Memnemyia Strickland 1915 Syn. 

Dendropaedium Dyar and Knab 1918 Syn. 

Calviana Strand 1928 Syn. 

Paramyzomyia Christophers and Barraud 1931 Syn. 

Palaeomyzomyia Swellengrebel and Rodenwaldt 1932 Syn. 

Eomyzomyia Evans 1934 Syn. 

Lophopodomyia Antunes 1937 Syn. 

Shannoniella Fonseca and Da Silva Ramos 1939 Syn. 

Shannonesia Fonseca and Da Silva Ramos 1940 Syn. 

Arthuromyia Galvao 1941 Syn. 

Ayrozamyia Coutinho and Farias 1942 Syn. 

Russellia Vargas 1943 Syn. 

Missirolimyia Vargas 1950 Syn. 

Maculipennia Buonomini and Mariani 1953 Syn. 

Anopheles albimanus Wiedemann 1820 (Anopheles) 

Anopheles cubensis Agramonte 1900 Syn. 

Anopheles albipes Theobald 1901 Syn. 

Cellia tarsimaculata Goedli 1905 Syn. 

Anopheles bisignatus Hoffmann 1938 Syn. 

Anopheles trisignatus Hoffmann 1938 Syn. 

Anopheles atropos Dyar and Knab 1906 (Anopheles) 

Anopheles ferruginosus Wiedemann 1828 Nomob. 

Anopheles barberi Coquillett 1903 (Anopheles) 

Anopheles niger Herrick 1905 Syn. 

Anopheles bradleyi King 1939 (Anopheles) 

Anopheles crucians Wiedemann 1828 (Anopheles) 

Anopheles earlei Vargas 1943 (Anopheles) 

Anopheles franciscanus McCracken 1904 (Anopheles) 

Anopheles boydi Vargas 1939 Syn. 

Anopheles willardi Vargas 1941 Syn. 

Anopheles freeborni Aitken 1939 (Anopheles) 

Anopheles georgianus King 1939 (Anopheles) 

Anopheles hermsi Barr and Guptavanij 1988 (Anopheles) 

Anopheles judithae Zavortink 1969 (Anopheles) 

Anopheles occidentalis Dyar and Knab 1906 (Anopheles) 

Anopheles perplexens Ludlow 1907 (Anopheles) 

Anopheles pseudopunctipennis Theobald 1901 (Anopheles) 

Anopheles peruvianus Tamayo 1907 Syn. 

Proterorhynchus argentinus Brethes 1912 Syn. 

Anopheles tucumanus Lahille 1912 Syn. 

Anopheles punctipennis Say 1823 (Culex) 

Culex hyemalis Fitch 1847 Syn. 

Anopheles stonei Vargas 1941 Syn. 

Anopheles quadrimaculatus Say 1824 (Anopheles) 

Anopheles annulimanus Wulp 1867 Syn. 

Anopheles walkeri Theobald 1901 (Anopheles) 

Culex Linnaeus 1758 

Heteronycha Lynch Arribalzaga 1891 Syn. 

Melanoconion Theobald 1903 Syn. 

Lasioconops Theobald 1903 Syn. 

Heptaphlebomyia Theobald 1903 Syn. 

Lutzia Theobald 1903 Syn. 

Aedinus Lutz 1904 Syn. 

Neoculex Dyar 1905 Syn. 

Carrollia Lutz 1905 Syn. 

Pseudoheptaphlebomyia Ventrillon 1905 Syn. 

Trichopronomyia Theobald 1905 Syn. 

Trichorhynchus Theobald 1905 Homo. 

Lophoceraomyia Theobald 1905 Syn. 

Melanoconium Blanchard 1905 Emend. 

Gnophodeomyia Theobald 1905 Syn. 

Mochlostyrax Dyar and Knab 1906 Syn. 

Jamesia Christophers 1906 Syn. 

Micraedes Coquillett 1906 Syn. 

Tinolestes Coquillett 1906 Syn. 

Aporoculex Theobald 1907 Syn. 

Leucomyia Theobald 1907 Homo. 

Oculeomyia Theobald 1907 Syn. 

Culiciomyia Theobald 1907 Syn. 

Neomelanoconion Theobald 1907 Syn. 

Philodendromyia Theobald 1907 Syn. 

Maillotia Theobald 1907 Syn. 

Microculex Theobald 1907 Syn. 

Acalleomyia Leicester 1908 Syn. 

Pectinopalpus Theobald 1909 Syn. 

Eumelanomyia Theobald 1909 Syn. 

Protomelanoconion Theobald 1909 Syn. 

Cyathomyia Meijere 1910 Syn. 

Asebeomyia Aiken 1911 Syn. 

Theobaldiomyia Brunetti 1912 Syn. 

Trichorhynchomyia Burnetti 1912 Syn. 

Phalangomyia Dyar and Knab 1914 Syn. 

Laiomyia Izquierdo 1916 Syn. 

Transculicia Dyar 1918 Syn. 

Choeroporpa Dyar 1918 Syn. 

Transculicia Dyar 1918 Syn. 

Cacoculex Dyar 1918 Syn. 

Helcoporpa Dyar 1918 Syn. 

Barraudius Edwards 1921 Syn. 

Carrollella Lutz 1921 Syn. 

Anoedioporpa Dyar 1923 Syn. 

Lasiosiphon Kirkpatrick 1925 Syn. 

Dinoporpa Dyar 1928 Syn. 

Mochthogenes Edwards 1930 Syn. 

Upsiloporpa Komp and Curry 1932 Syn. 

Acallyntrum Stone and Penn 1948 Syn. 

Thaiomyia Bram 1966 Syn. 

Allimanta Casal and Garcia 1968 Syn. 

Belkinomyia Adames and Galindo 1973 Syn. 

Afroculex Danilov 1989 Syn. 

Kitzmilleria Danilov 1989 Syn. 

Phenacomyia Harbach and Peyton 1992 Syn. 

Culex abominator Dyar and Knab 1909 (Culex) 

Culex anips Dyar 1916 (Culex) 

Culex apicalis Adams 1903 (Culex) 

Culex arizonensis Bohart 1949 (Culex) 

Culex atratus Theobald 1901 (Culex) 

Culex falsificator Dyar and Knab 1909 Syn. 

Culex advieri Senevet 1938 Syn. 

Culex bahamensis Dyar and Knab 1906 (Culex) 

Culex eleuthera Dyar 1918 Syn. 

Culex petersoni Dyar 1920 Syn. 

Culex boharti Brookman and Reeves 1950 (Culex) 

Culex reevesi Bohart 1949 Homo. 

Culex chidesteri Dyar 1912 (Culex) 

Culex finlayi Perez Vigueras 1956 Syn. 

Culex deanei Correa and Ramalho 1959 Syn. 

Culex coronator Dyar and Knab 1906 (Culex) 

Culex mosseri Vargas and Martinez 1954 Syn. 

Culex declarator Dyar and Knab 1906 (Culex) 

Culex inquisitor Dyar and Knab 1906 Syn. 

Culex proclamator Dyar and Knab 1906 Syn. 

Culex jubilator Dyar and Knab 1907 Syn. 

Culex revelator Dyar and Knab 1907 Syn. 

Culex dictator Dyar and Knab 1909 Syn. 

Culex vindicator Dyar and Knab 1909 Syn. 

Culex forattinii Correa and Ramalho 1959 Syn. 

Culex erraticus Dyar and Knab 1906 (Mochlostyrax) 

Culex egberti Dyar and Knab 1907 Syn. 

Culex leprincei Dyar and Knab 1907 Syn. 

Culex invocator Pazos 1908 Syn. 

Culex trachycampa Dyar and Knab 1909 Syn. 

Culex peribleptus Dyar and Knab 1918 Syn.

Culex pose Dyar and Knab 1918 Syn. 

Culex degustator Dyar 1921 Syn. 

Culex homoepas Dyar and Ludlow 1921 Syn. 

Culex borinqueni Root 1922 Syn. 

Culex tovari Evans 1924 Syn. 

Culex erythrothorax Dyar 1907 (Culex) 

Culex federalis Dyar 1923 Syn. 

Culex badgeri Dyar 1924 Syn. 

Culex interrogator Dyar and Knab 1906 (Culex) 

Culex reflector Dyar and Knab 1909 Syn. 

Culex iolambdis Dyar 1918 (Culex) 

Culex latisquama Coquillett 1906 (Tinolestes) 

Culex molestus Forskal 1775 (Culex) 

Culex domesticus Germar 1817 Syn. 

Culex haematophagus Ficalbi 1893 Syn. 

Culex quasimodestus Theobald 1905 Syn. 

Culex autogenicus Roubaud 1935 Syn. 

Culex berbericus Roubaud 1935 Syn. 

Culex sternopallidus Roubaud 1945 Syn. 

Culex sternopunctatus Roubaud 1945 Syn. 

Culex mulrennani Basham 1948 (Culex) 

Culex nigripalpus Theobald 1901 (Culex) 

Culex palus Theobald 1903 Syn. 

Culex similis Theobald 1903 Syn. 

Culex biocellatus Theobald 1903 Syn. 

Culex microsquamosus Theobald 1905 Syn. 

Culex mortificator Dyar and Knab 1906 syn. 

Culex carmodyae Dyar and Knab 1906 Syn. 

Culex factor Dyar and Knab 1906 Syn. 

Culex regulator Dyar and Knab 1906 Syn. 

Trichopronomyia microannulata Theobald 1907 Syn. 

Culex proximus Dyar and Knab 1909 Syn. 

Culex caraibeus Howard, Dyar, Knab 1913 Syn. 

Culex prasinopleurus Martini 1914 Syn. 

Culex azuayus Levi-Castillo 1954 Syn. 

Culex opisthopus Komp 1926 (Culex) 

Melanoconion annulipes Theobald 1907 Syn. (?) 

Culex mychonde Komp 1928 Syn. 

Culex cedecei Stone and Hair 1968 Syn. 

Culex peccator Dyar and Knab 1909 (Culex) 

Culex incriminator Dyar and Knab 1909 Syn. 

Culex peus Speiser 1904 (Culex) 

Culex affinis Adams 1903 Homo. 

Culex thriambus Dyar 1921 Syn. 

Culex pilosus Dyar and Knab 1906 (Mochlostyrax) 

Mochlostyrax floridanus Dyar and Knab 1906 Syn. 

Mochlostyrax cubensis Dyar and Knab 1906 Homo. 

Mochlostyrax jamaicensis Grabham 1906 Homo. 

Culex agitator Dyar and Knab 1907 Syn. 

Culex hesitator Dyar and Knab 1907 Syn. 

Culex deceptor Dyar and Knab 1909 Syn. 

Culex reductor Dyar and Knab 1909 Syn. 

Culex ignobilis Dyar and Knab 1909 Syn. 

Culex mastigia Howard, Dyar, Knab 1913 Syn. 

Culex curopinensis Bonne-Wepster and Bonne 1920 Syn. 

Culex colombiensis Dyar 1924 Syn. 

Culex radiatus Senevet and Abonnenc 1939 Syn. 

Culex pipiens Linnaeus 1758 (Culex) 

Culex bifurcatus Linnaeus 1758 Syn. 

Culex fasciatus Mueller 1764 Syn. 

Culex trifurcatus Fabricius 1794 Syn. 

Culex luteus Meigen 1804 Syn. 

Culex rufus Meigen 1818 Syn. 

Culex bicolor Meigen 1818 Syn. 

Culex marginalis Stephens 1825 Syn. 

Culex meridionalis Leach 1825 Syn. 

Culex consobrinus Robineau-Desvoidy 1827 Syn. 

Culex calcitrans Robineau-Desvoidy 1827 Syn. 


Culex thoracicus Robineau-Desvoidy 1827 Syn. 

Culex pallipes Waltl 1835 Syn. 

Culex unistriatus Curtis 1837 Syn. 

Culex pallipes Macquart 1838 Syn. 

Culex rufinus Bigot 1888 Syn. 

Culex agilis Bigot 1889 Syn. 

Culex phytophagus Ficalbi 1890 Syn. 

Aedes pallens Coquillett 1898 Syn. 

Culex melanorhinus Giles 1900 Syn. 

Culex varioannulatus Theobald 1903 Syn. 

Culex azoriensis Theobald 1903 Syn. 

Culex longefurcatus Becker 1903 Syn. 

Culex osakensis Theobald 1907 Syn. 

Culex dipseticus Dyar and Knab 1909 Syn. 

Culex comitatus Dyar and Knab 1909 Syn. 

Culex doliorum Edwards 1912 Syn. 

Culex disjunctus Roubaud 1957 Syn. 

Culex calloti Rioux and Pech 1959 Syn. 

Culex torridus Iglisch 1977 Syn. 

Culex erectus Iglisch 1977 Syn. 

Culex quinquefasciatus Say 1823 (Culex) 

Culex pungens Wiedemann 1828 Syn. 

Culex fatigans Wiedemann 1828 Syn. 

Culex aestuans Wiedemann 1828 Syn. 

Culex acer Walker 1848 Syn. 

Culex cingulatus Doleschall 1856 Syn. 

Culex cubensis Bigot 1857 Syn. 

Culex anxifer Bigot 1859 Syn. 

Culex serotinus Philippi 1865 Syn. 

Culex autumnalis Weyenbergh 1882 Syn. 

Culex penafieli Sanchez 1885 Syn. 

Culex macleayi Skuse 1889 Syn. 

Culex skusii giles 1900 Syn. 

Culex doleschallii Giles 1900 Syn. 

Culex albolineatus Giles 1901 Syn. 

Culex quasipipiens Theobald 1901 Syn. 

Culex fouchowensis Theobald 1901 Syn. 

Culex reesi Theobald 1901 Syn. 

Culex sericeus Theobald 1901 Syn. 

Culex luteoannulatus Theobald 1901 Syn. 

Culex trilineatus Theobald 1901 Syn. 

Culex pallidocephala Theobald 1904 Syn. 

Culex cartroni Ventrillon 1905 Syn. 

Culex barbarus Dyar and Knab 1906 Syn. 

Culex didieri Neveu-Lemaire 1906 Syn. 

Culex pygmaeus Neveu-Lemaire 1906 Syn. 

Culex zeltneri Neveu-Lemaire 1906 Syn. 

Culex quasilinealis Theobald 1907 Syn. 

Culex stoehri Theobald 1907 Syn. 

Culex christophersii Theobald 1907 Syn. 

Culex raymondii Tamayo 1907 Syn. 

Culex aikenii Dyar and Knab 1908 Syn. 

Culex minor Theobald 1908 Syn. 

Culex revocator Dyar and Knab 1909 Syn. 

Culex lachrimans Dyar and Knab 1909 Syn. 

Culex goughii Theobald 1911 Syn. 

Culicelsa fuscus Taylor 1914 Syn. 

Culex aseyehae Dyar and Knab 1915 Syn. 

Culex townsvillensis Taylor 1919 Syn. 

Culex hensemaeon Dyar 1920 Syn. 

Culex nigrirostris Enderlein 1920 Syn. 

Culex reevesi Wirth 1948 (Culex) 

Culex restuans Theobald 1901 (Culex) 

Culex brehmei Knab 1916 Syn. 

Culex salinarius Coquillett 1904 (Culex) 

Culex stigmatosoma Dyar 1907 (Culex) 

Culex eumimetes Dyar and Knab 1908 Syn. 

Culex tarsalis Coquillett 1896 (Culex) 


133

134 

Culex willistoni Giles 1900 Syn. 

Culex kelloggii Theobald 1903 Syn. 

Culex territans Walker 1856 (Culex) 

Culex saxatilis Grossbeck 1905 Syn. 

Culex frickii Ludlow 1906 Syn. 

Culex pyrenaicus Brolemann 1919 Syn. 

Culex nematoides Dyar and Shannon 1925 Syn. 

Culiseta Felt 1904 

Theobaldia Neveu-Lemaire 1902 Homo. 

Culicella Felt 1904 Syn. 

Theobaldinella Blanchard 1905 Syn. 

Pseudotheobaldia Theobald 1907 Syn. 

Climacura Howard, Dyar, and Knab 1915 Syn. 

Allotheobaldia Broelemann 1919 Syn. 

Theomyia Edwards 1930 Syn. 

Austrotheobaldia Dobrotworsky 1954 Syn. 

Neotheobaldia Dobrotworsky 1958 Syn. 

Culiseta alaskaensis Ludlow 1906 (Theobaldia) 

Theobaldia arctica Edwards 1920 Syn. 

Culiseta siberiensis Ludlow 1920 Syn. 

Theobaldia indica Edwards 1920 Syn. 

Theobaldia wassilievi Shingarev 1927 Syn. 

Culiseta impatiens Walker 1848 (Culex) 

Culex pinguis Walker 1866 Syn. 

Culex absobrinus Felt 1904 Syn. 

Culiseta incidens Thomson 1869 (Culex) 

Culiseta inornata Williston 1893 (Culex) 

Culex magnipennis Felt 1904 Syn. 

Culiseta melanurus Coquillett 1902 (Culex) 

Culiseta minnesotae Barr 1957 (Culiseta) 

Culiseta morsitans Theobald 1901 (Culex) 

Culex dyari Coquillett 1902 Syn. 

Culex brittoni Felt 1905 Syn. 

Culicella parodites Dyar 1928 Syn. 

Culiseta particeps Adams 1903 (Culex) 

Culiseta maccrackenae Dyar and Knab 1906 Syn. 

Culiseta dugesi Dyar and Knab 1906 Syn. 

Culiseta silvestris Shingarev 1928 (Culiseta) 

Theobaldia ochroptera Peus 1935 Syn. 

Culiseta amurensis Maslov 1964 Syn. 

Deinocerites Theobald 1901 

Brachiosoma Theobald 1901 Syn. 

Brachyiomyia Theobald 1901 Syn. 

Deinokerides Giles 1902 Missp. 

Dinocerites Blanchard 1905 Emend. 

Dinomimetes Knab 1907 Syn. 

Dinanamesus Dyar and Knab 1909 Syn. 

Deinocerites cancer Theobald 1901 (Deinocerites) 

Deinocerites tetraspathus Dyar and Knab 1909 Syn. 

Deinocerites mathesoni Belkin and Hogue 1959 (Deinocerites) 

Deinocerites pseudes Dyar and Knab 1909 (Deinocerites) 

Haemagogus Williston 1896 

Stegoconops Lutz 1905 Syn. 

Cacomyia Coquillett 1906 Syn. 

Conopostegus Dyar 1925 Syn. 

Longipalpifer Levi-Castillo 1951 Syn. 

Haemagogus equinus Theobald 1903 (Haemagogus) 

Aedes affirmatus Dyar and Knab 1906 Syn. 

Aedes philosophicus Dyar and Knab 1906 Syn. 

Mansonia Blanchard 1901 

Panoplites Theobald 1900 Homo. 



Coquillettidia Dyar 1905 Syn. 

Mansonioides Theobald 1907 Syn. 

Mansonia dyari Belkin 1970 (Mansonia) 

Mansonia perturbans Walker 1856 (Culex) 

Culex testaceus Wulp 1867 Syn. 

Culex ochropus Dyar and Knab 1907 Syn. 

Mansonia titillans Walker 1848 (Culex) 

Orthopodomyia Theobald 1904 

Bancroftia Lutz 1904 Syn. 

Pneumaculex Dyar 1905 Syn. 

Newsteadina Theobald 1909 Syn. 

Thomasina Newstead and Carter 1911 Syn. 

Orthopodomyia alba Baker 1936 (Orthopodomyia) 

Orthopodomyia kummi Edwards 1939 (Orthopodomyia) 

Orthopodomyia signiferus Coquillett 1896 (Culex) 

Orthopodomyia californica Bohart 1950 Syn. 

Psorophora Robineau-Desvoidy 1827 

Janthinosoma Lynch Arribalzaga 1891 Syn. 

Conchyliastes Theobald 1901 Syn. 

Ianthinosoma Lynch Arribalzaga 1902 Emend. 

Grabhamia Theobald 1903 Syn. 

Feltidia Dyar 1905 Syn. 

Lepidosia Coquillett 1906 Syn. 

Ceratocystia Dyar and Knab 1906 Syn. 

Lanesia Levi-Castillo 1953 Syn. 

Psorophora ciliata Fabricius 1794 (Culex) 

Culex cyanopennis Humboldt 1819 Syn. 

Culex molestus Wiedemann 1820 Syn. 

Culex rubidus Robineau-Desvoidy 1827 Syn. 

Culex boscii Robineau-Desvoidy 1827 Syn. 

Culex tibialis Robineau-Desvoidy 1827 Syn. 

Culex conterrens Walker 1856 Syn. 

Psorophora lynchi Brethes 1916 Syn. 

Psorophora ctites Dyar 1918 Syn. 

Psorophora columbiae Dyar and Knab 1906 (Janthinosoma) 

Janthinosoma floridense Dyar and Knab 1906 Syn. 

Janthinosoma texanum Dyar and Knab 1906 Syn. 

Janthinosoma toltecum Dyar and Knab 1906 Syn. 

Psorophora cyanescens Coquillett 1902 (Culex) 

Psorophora purpurascens Edwards 1922 Syn. 

Psorophora tovari Evans 1922 Syn. 

Psorophora discolor Coquillett 1903 (Culex) 

Psorophora ferox Humboldt 1819 (Culex) 

Culex posticatus Wiedemann 1821 Syn. 

Culex musicus Say 1829 Homo. 

Janthinosoma sayi Dyar and Knab 1906 Syn. 

Janthinosoma echinata Grabham 1906 Syn. 

Janthinosoma terminalis Coquillett 1906 Syn. 

Janthinosoma vanhalli Dyar and Knab 1906 Syn. 

Janthinosoma sayi Theobald 1907 Syn. 

Janthinosoma coquillettii Theobald 1907 Syn. 

Janthinosoma jamaicensis Theobald 1907 Syn. 

Aedes pazosi Pazos 1908 Syn.


Janthinosoma centralis Brethes 1910 Syn. 

Psorophora horridus Dyar and Knab 1908 (Aedes) 

Psorophora howardii Coquillett 1901 (Psorophora) 

Psorophora virescens Dyar and Knab 1906 Syn. 

Psorophora simplex Martini 1935 Syn. 

Psorophora johnstonii Grabham 1905 (Janthinosoma) 

Janthinosoma coffini Dyar and Knab 1906 Syn. 

Janthinosoma schwarzi Dyar and Knab 1906 Syn. 

Psorophora longipalpus Randolph and O'Neill 1944 (Psorophora) 

Psorophora longipalpis Roth 1945 Syn. 

Psorophora mathesoni Belkin 1975 (Psorophora) 

Psorophora mexicanus Bellardi 1859 (Culex) 

Psorophora pygmaea Theobald 1903 (Grabhamia) 

Culex nanus Coquillett 1903 Syn. 

Taeniorhynchus antiguae Giles 1904 Syn. 

Psorophora signipennis Coquillett 1904 (Taeniorhynchus) 

Psorophora varipes Coquillett 1904 (Conchyliastes) 

Toxorhynchites Theobald 1901 

Megarhinus Robineau-Desvoidy 1827 Homo. 

Lynchiella Lahille 1904 Syn. 

Ankylorhynchus Lutz 1904 Syn. 

Worcesteria Banks 1906 Syn. 

Teromyia Leicester 1908 Syn. 

Afrorhynchus Ribeiro 1991 Syn. 

Toxorhynchites rutilus Coquillett 1896 (Megarhinus) 

Megarhinus septentrionalis Dyar and Knab 1906 Syn. 

Megarhinus herrickii Theobald 1906 Syn. 

Uranotaenia Lynch Arribalzaga 1891 

Anisocheleomyia Theobald 1905 Syn. 

Pseudouranotaenia Theobald 1905 Syn. 

Pseudoficalbia Theobald 1912 Syn. 

Uranotaenia anhydor Dyar 1907 (Uranotaenia) 

Uranotaenia syntheta Dyar and Shannon 1924 Syn. 

Uranotaenia lowii Theobald 1901 (Uranotaenia) 

Uranotaenia continentalis Dyar and Knab 1906 Syn. 

Uranotaenia minuta Theobald 1907 Syn. 

Uranotaenia monilis Shannon and Del Ponte 1928 Syn. 

Uranotaenia sapphirina Osten Sacken 1868 (Aedes) 

Uranotaenia coquilletti Dyar and Knab 1906 Syn. 

Wyeomyia Theobald 1901 

Dendromyia Theobald 1903 Syn. 

Prosopolepis Lutz 1905 Syn. 

Menolepis Lutz 1905 Syn. 

Calladimyia Dyar 1919 Syn. 

Cleobonnea Dyar 1919 Syn. 

Decamyia Dyar 1919 Syn. 

Dinomyia Dyar 1919 Syn. 

Heliconiamyia Dyar 1919 Syn. 

Hystatamyia Dyar 1919 Syn. 

Triamyia Dyar 1919 Syn. 

Diphalangarpe Dyar 1919 Syn. 

Dodecamyia Dyar 1919 Syn. 

Miamyia Dyar 1919 Syn. 

Pentemyia Dyar 1919 Syn. 

Shropshirea Dyar 1922 Syn. 

Phyllozomyia Dyar 1924 Syn. 

Eunicemyia Dyar and Shannon 1924 Syn. 

Janicemyia Dyar and Shannon 1924 Syn. 

Phyllozomyia Dyar 1924 Syn. 

Technicomyia Dyar 1925 Syn. 

Melanolepis Dyar 1928 Syn. 

Nunezia Dyar 1928 Syn. 

Antunesmyia Lane and Cerqueira 1942 Syn. 

Cruzmyia Lane and Cerqueira 1942 Syn. 

Davismyia Lane and Cerqueira 1942 Syn. 

Zinzala Zavortink 1986 Syn. 

Caenomyiella Harbach and Peyton 1990 Syn. 

Exallomyia Harbach and Peyton 1991 Syn. 

Wyeomyia haynei Dodge 1947 (Wyeomyia) 

Wyeomyia mitchellii Theobald 1905 (Dendromyia) 

Dendromyia jamaicensis Theobald 1905 Syn. 

Wyeomyia ochrura Dyar and Knab 1906 Syn. 

Wyeomyia violescens Dyar and Knab 1906 Syn. 

Wyeomyia glaucocephala Dyar and Knab 1906 Syn. 

Wyeomyia abia Dyar and Knab 1908 Syn. 

Wyeomyia antoinetta Dyar and Knab 1909 Syn. 

Wyeomyia smithii Coquillett 1901 (Aedes) 

Wyeomyia vanduzeei Dyar and Knab 1906 (Wyeomyia) 

Wyeomyia argyrura Dyar and Knab 1908 Syn. 

Wyeomyia conchita Dyar and Knab 1909 Syn. 

CURTONOTIDAE 

Curtonotum Macquart 1843 

Diplocentra Loew 1862 Syn. 

Curtonotum helva Loew 1862 (Diplocentra) 

CYPSELOSOMATIDAE 

Latheticomyia Wheeler 1956 

Latheticomyia lineata Wheeler 1956 (Latheticomyia) 

Latheticomyia tricolor Wheeler 1956 (Latheticomyia) 

DEUTEROPHLEBIIDAE 

Deuterophlebia Edwards 1922 

Deuterophlebia coloradensis Pennak 1945 (Deuterophlebia) 

Deuterophlebia inyoensis Kennedy 1960 (Deuterophlebia) 

Deuterophlebia nielsoni Kennedy 1958 (Deuterophlebia) 

Deuterophlebia personata Courtney 1990 (Deuterophlebia) 

Deuterophlebia shasta Wirth 1951 (Deuterophlebia) 

Deuterophlebia vernalis Courtney 1990 (Deuterophlebia) 

DIASTATIDAE 

Campichoeta Macquart 1835 

DIPTERA: CULICIDAE, CURTONOTIDAE, CYPSELOSOMATIDAE, DEUTEROPHLEBIIDAE, 

DIASTATIDAE 

135

136 

Thryptocheta Rondani 1856 Syn. 

Campichoeta griseola Zetterstedt 1855 (Geomyza) 

Thryptocheta micans Hendel 1911 Syn. 

Campichoeta latigena McAlpine 1962 (Campichoeta) 

Diastata Meigen 1830 

Trichoptera Lioy 1864 Homo. 

Calopterella Coquillett 1910 Syn. 

Diastata eluta Loew 1863 (Diastata) 

Diastata modesta Melander 1913 (Diastata) 

Diastata repleta Walker 1849 (Notiphila) 

Drosophila decemguttata Walker 1852 Syn. 

Diastata pulchra Loew 1861 Syn. 

Diastata vagans Loew 1864 (Diastata) 

DIOPSIDAE 

Sphyracephala Say 1828 

Hexechopsis Rondani 1875 Syn. 

Zygocephala Rondani 1875 Syn. 

Pseudodiopsis Hendel 1917 Syn. 

Microdiopsis Curran 1934 Syn. 

Sphyracephala brevicornis Say 1817 (Diopsis) 

Sphyracephala subbifasciata Fitch 1855 (Sphyracephala) 

DIXIDAE 

Dixa Meigen 1818 

Eriopterites Meunier 1915 Syn. 

Dixa adleri Peters 1987 (Dixa) 

Dixa arge Dyar and Shannon 1924 (Dixa) 

Dixa blax Dyar and Shannon 1924 (Dixa) 

Dixa brevis Garrett 1924 (Dixa) 

Dixa distincta Garrett 1925 (Dixa) 

Dixa fluvica Peters 1966 (Dixa) 

Dixa fraterna Garrett 1924 (Dixa) 

Dixa fusca Loew 1863 (Dixa) 

Dixa mystica Dyar and Shannon 1924 Syn. 

Dixa hegemonica Dyar and Shannon 1924 (Dixa) 

Dixa inextricata Dyar and Shannon 1924 (Dixa) 

Dixa johannseni Garrett 1924 (Dixa) 

Dixa lobatus Garrett 1924 (Dixa) 

Dixa melanderi Peters 1966 (Dixa) 

Dixa modesta Johannsen 1903 (Dixa) 

Dixa neohegemonica Peters 1966 (Dixa) 

Dixa notata Loew 1863 (Dixa) 

Dixa pseudindiana Peters 1981 (Dixa) 

Dixa pullogruma Peters 1966 (Dixa) 

Dixa recens Walker 1848 (Dixa) 

Dixa rhathyme Dyar and Shannon 1924 (Dixa) 

Dixa montanus Garrett 1924 Syn. 

Dixa parvus Garrett 1924 Syn. 

Dixa garretti Cooper and Rapp 1944 Syn. 

Dixa rudis Garrett 1924 (Dixa) 


DIPTERA: DIASTATIDAE, DIOPSIDAE, DIXIDAE, DOLICHOPODIDAE 

Dixa similis Johannsen 1923 (Dixa) 

Dixa terna Loew 1863 (Dixa) 

Dixa venosa Loew 1872 (Dixa) 

Dixa xavia Dyar and Shannon 1924 (Dixa) 

Dixella Dyar and Shannon 1924 

Dixapuella Dyar and Shannon 1924 Syn. 

Paradixa Tonnoir 1924 Syn. 

Dixella alexanderi Peters 1970 (Dixella) 

Dixella aliciae Johannsen 1924 (Dixa) 

Dixella aurora Peters 1966 (Dixella) 

Dixella californica Johannsen 1923 (Dixa) 

Dixa thones Dyar and Shannon 1924 Syn. 

Dixella clavata Loew 1869 (Dixa) 

Dixella cornuta Johannsen 1923 (Dixa) 

Dixa somnolenta Dyar and Shannon 1924 Syn. 

Dixa spiralis Garrett 1924 Syn. 

Dixa plexipus Garrett 1925 Syn. 

Dixa simplex Garrett 1925 Syn. 

Dixella cumbrica Peters 1966 (Dixella) 

Dixella deltoura Peters 1966 (Dixella) 

Dixella dorsalis Garrett 1924 (Dixa) 

Dixella dyari Garrett 1924 (Dixa) 

Dixella indiana Dyar 1925 (Dixa) 

Dixella marginata Loew 1863 (Dixa) 

Dixella naevia Peus 1934 (Dixa) 

Dixella neoaliciae Garrett 1924 (Dixa) 

Dixella nova Walker 1848 (Dixa) 

Dixa centralis Loew 1863 Syn. 

Dixa nocheles Dyar and Shannon 1924 Syn. 

Dixa occidentalis Garrett 1924 Syn. 

Dixella serrata Garrett 1924 (Dixa) 

Dixella techana Peters 1966 (Dixella) 

Dixella universitatis Cockerell 1926 (Dixa) 

Dixella vespertina Peters 1966 (Dixella) 

Meringodixa Nowell 1951 

Meringodixa chalonensis Nowell 1951 (Meringodixa) 

DOLICHOPODIDAE 

Achalcus Loew 1857 

Achalcus oregonensis Harmston and Miller 1966 (Systenus) 

Achalcus utahensis Harmston and Miller 1966 (Systenus) 

Aphrosylus Haliday 1851 

Paraphrosylus Becker 1922 Syn. 

Aphrosylus californicus Harmston 1952 (Aphrosylus) 

Aphrosylus direptor Wheeler 1897 (Aphrosylus) 

Aphrosylus grassator Wheeler 1897 (Aphrosylus) 

Aphrosylus nigripennis Van Duzee 1924 (Aphrosylus) 

Aphrosylus praedator Wheeler 1897 (Aphrosylus) 

Aphrosylus wirthi Harmston 1951 (Aphrosylus) 

Argyra Macquart 1834 

Porphyrops Meigen 1824 Syn. (?) 

Leucostola Loew 1857 Syn.

Argyra abdominalis Say 1829 (Dolichopus) 

Argyra albicans Loew 1861 (Argyra) 

Porphyrops quadriplagiatus Harris 1835 Unav. 

Argyra albicoxa Van Duzee 1925 (Argyra) 

Argyra albiventris Loew 1864 (Argyra) 

Argyra aldrichi Johnson 1904 (Argyra) 

Argyra angustata Van Duzee 1925 (Argyra) 

Argyra argentiventris Van Duzee 1925 (Argyra) 

Argyra barbipes Van Duzee 1925 (Argyra) 

Argyra basalis Van Duzee 1932 (Argyra) 

Argyra bimaculata Van Duzee 1925 (Argyra) 

Argyra brevipes Van Duzee 1925 (Argyra) 

Argyra calceata Loew 1861 (Argyra) 

Argyra calcitrans Loew 1861 (Argyra) 

Argyra californica Van Duzee 1925 (Argyra) 

Argyra ciliata Van Duzee 1924 (Argyra) 

Argyra cingulata Loew 1861 (Leucostola) 

Argyra condomina Harmston and Knowlton 1946 (Argyra) 

Argyra currani Van Duzee 1925 (Argyra) 

Argyra cylindrica Loew 1864 (Argyra) 

Argyra dakotensis Harmston and Knowlton 1939 (Argyra) 

Argyra fasciventris Van Duzee 1930 (Argyra) 

Argyra femoralis Van Duzee 1925 (Argyra) 

Argyra flavicornis Van Duzee 1925 (Argyra) 

Argyra flavicoxa Van Duzee 1925 (Argyra) 

Argyra flavipes Van Duzee 1925 (Argyra) 

Argyra grayi Robinson 1964 (Argyra) 

Argyra hirta Van Duzee 1925 (Argyra) 

Argyra idahona Harmston and Knowlton 1946 (Argyra) 

Argyra inaequalis Van Duzee 1925 (Argyra) 

Argyra involuta Van Duzee 1925 (Argyra) 

Argyra johnsoni Van Duzee 1925 (Argyra) 

Argyra minuta Loew 1861 (Argyra) 

Argyra nigricoxa Van Duzee 1925 (Argyra) 

Argyra nigripes Loew 1864 (Argyra) 

Argyra nigriventris Van Duzee 1925 (Argyra) 

Argyra obscura Van Duzee 1925 (Argyra) 

Argyra pilosicornis Walker 1849 (Porphyrops) 

Argyra robusta Johnson 1906 (Argyra) 

Argyra scutellaris Van Duzee 1925 (Argyra) 

Argyra sericata Van Duzee 1925 (Argyra) 

Argyra setipes Van Duzee 1925 (Argyra) 

Argyra similis Harmston and Knowlton 1940 (Argyra) 

Argyra spina Van Duzee 1925 (Argyra) 

Argyra splendens Van Duzee 1933 (Argyra) 

Argyra splendida Van Duzee 1925 Homo. 

Argyra thoracica Van Duzee 1925 (Argyra) 

Argyra utahna Harmston 1951 (Argyra) 

Argyra velutina Van Duzee 1925 (Argyra) 

Asyndetus Loew 1869 

Anchineura Thomson 1869 Syn. 

Meringopherusa Becker 1902 Syn. 

Asyndetus ammophilus Loew 1869 (Asyndetus) 

Asyndetus appendiculatus Loew 1869 (Asyndetus) 

Asyndetus caudatus Van Duzee 1916 (Asyndetus) 

Asyndetus cornutus Van Duzee 1916 (Asyndetus) 

Asyndetus crassipodus Harmston 1968 (Asyndetus) 

Asyndetus flavipalpus Van Duzee 1932 (Asyndetus) 

Asyndetus harbeckii Van Duzee 1914 (Asyndetus) 

Asyndetus hardyi Robinson 1964 (Asyndetus) 

Asyndetus interruptus Loew 1861 (Diaphorus) 

Asyndetus johnsoni Van Duzee 1916 (Asyndetus) 

Asyndetus latus Van Duzee 1916 (Asyndetus) 

Asyndetus nevadensis Harmston 1968 (Asyndetus) 

Asyndetus nigripes Van Duzee 1916 (Asyndetus) 

Asyndetus occidentalis Van Duzee 1919 (Asyndetus) 


Asyndetus oregonensis Harmston 1966 (Asyndetus) 

Asyndetus parvicornis Van Duzee 1932 (Asyndetus) 

Asyndetus scopifer Harmston 1952 (Asyndetus) 

Asyndetus severini Harmston and Knowlton 1939 (Asyndetus) 

Asyndetus spinitarsis Harmston 1951 (Asyndetus) 

Asyndetus syntormoides Wheeler 1899 (Asyndetus) 

Asyndetus texanus Van Duzee 1916 (Asyndetus) 

Asyndetus utahensis Harmston and Knowlton 1942 (Asyndetus) 

Calyxochaetus Bigot 1888 

Nothosympycnus Wheeler 1899 Syn. 

Calyxochaetus cilifemoratus Van Duzee 1924 (Nothosympycnus) 

Sympycnus fuscitibialis Harmston and Knowlton 1940 Syn. 

Calyxochaetus clavicornis Van Duzee 1930 (Sympycnus) 

Calyxochaetus distortus Van Duzee 1930 (Sympycnus) 

Calyxochaetus fortunatus Wheeler 1899 (Nothosympycnus) 

Calyxochaetus frontalis Loew 1861 (Sympycnus) 

Calyxochaetus hastatus Van Duzee 1930 (Sympycnus) 

Calyxochaetus inornatus Van Duzee 1917 (Nothosympcnus) 

Calyxochaetus insolitus Van Duzee 1932 (Sympycnus) 

Calyxochaetus luteipes Van Duzee 1923 (Nothosympycnus) 

Nothosympycnus tarsalis Curran 1924 Syn. 

Calyxochaetus monticola Van Duzee 1932 (Sympycnus) 

Calyxochaetus nodatus Loew 1862 (Sympycnus) 

Nothosympycnus abbreviatus Van Duzee 1917 Syn. 

Calyxochaetus oreas Wheeler 1899 (Nothosympycnus) 

Sympycnus spatulatus Harmston and Knowlton 1940 Syn. 

Calyxochaetus sobrinus Wheeler 1899 (Nothosympycnus) 

Calyxochaetus vegetus Wheeler 1899 (Nothosympycnus) 

Campsicnemus Haliday 1851 

Camptosceles Haliday 1832 Unav. 

Ectomus Mik 1878 Syn. 

Emperoptera Grimshaw 1902 Syn. 

Campsicnemus alaskensis Harmston and Miller 1966 

(Campsicnemus) 

Campsicnemus alexanderi Harmston and Miller 1966 


Campsicnemus americanus Van Duzee 1924 (Campsicnemus) 

Campsicnemus arcuatus Van Duzee 1917 (Campsicnemus) 

Campsicnemus bryanti Malloch 1932 (Campsicnemus) 

Campsicnemus cinctipus Harmston 1968 (Campsicnemus) 

Campsicnemus claudicans Loew 1864 (Campsicnemus) 

Campsicnemus coloradensis Harmston and Miller 1966 


Campsicnemus curvispina Van Duzee 1930 (Campsicnemus) 

Campsicnemus calcaratus Van Duzee 1924 Homo. 

Campsicnemus degener Wheeler 1899 (Campsicnemus) 

Campsicnemus hirtipes Loew 1861 (Campsicnemus) 

Campsicnemus melanus Harmston and Knowlton 1942 


Campsicnemus milleri Harmston 1966 (Campsicnemus) 

Campsicnemus montanus Harmston and Knowlton 1942 


Campsicnemus nigripes Van Duzee 1917 (Campsicnemus) 

Campsicnemus oedipus Wheeler 1899 (Campsicnemus) 

Campsicnemus oregonensis Harmston and Miller 1966 


Campsicnemus philoctetes Wheeler 1899 (Campsicnemus) 

Campsicnemus thersites Wheeler 1899 (Campsicnemus) 

Campsicnemus utahensis Harmston and Knowlton 1942 


Campsicnemus vanduzeei Curran 1933 (Campsicnemus) 

Campsicnemus wheeleri Van Duzee 1923 (Campsicnemus) 

Thinophilus variabilis Van Duzee 1932 Syn. 

DIPTERA: DOLICHOPODIDAE 

137

138 

Chrysotimus Loew 1857 

Chrysotimus arizonicus Robinson 1967 (Chrysotimus) 

Chrysotimus delicatus Loew 1861 (Chrysotimus) 

Chrysotimus flavicornis Van Duzee 1916 (Chrysotimus) 

Chrysotimus luteopalpus Curran 1923 (Chrysotimus) 

Chrysotimus luteus Curran 1930 (Chrysotimus) 

Chrysotimus occidentalis Harmston 1951 (Chrysotimus) 

Chrysotimus pusio Loew 1861 (Chrysotimus) 

Chrysotus Meigen 1824 

Achradocera Becker 1922 Syn. 



Chrysotus affinis Loew 1861 (Chrysotus) 

Chrysotus agalmus Harmston and Rapp 1968 (Chrysotus) 

Chrysotus albohirtus Van Duzee 1924 (Chrysotus) 

Chrysotus aldrichi Van Duzee 1924 (Chrysotus) 

Chrysotus annulatus Van Duzee 1924 (Chrysotus) 

Chrysotus anomalus Malloch 1914 (Chrysotus) 

Chrysotus arcuatus Van Duzee 1924 (Chrysotus) 

Chrysotus argentatus Van Duzee 1924 (Chrysotus) 

Chrysotus arkansensis Van Duzee 1930 (Chrysotus) 

Chrysotus atratus Van Duzee 1930 (Chrysotus) 

Chrysotus auratus Loew 1861 (Chrysotus) 

Chrysotus badius Van Duzee 1932 (Chrysotus) 

Chrysotus barbatus Loew 1861 (Synarthrus) 

Chrysotus validus Loew 1861 Syn. 

Xiphandrium americanum Wheeler 1896 Syn. 

Chrysotus barbipes Van Duzee 1932 (Chrysotus) 

Chrysotus bellulus Van Duzee 1932 (Chrysotus) 

Chrysotus bellus Van Duzee 1924 (Chrysotus) 

Chrysotus bracteatus Van Duzee 1924 (Chrysotus) 

Chrysotus caerulus Van Duzee 1924 (Chrysotus) 

Chrysotus californicus Van Duzee 1924 (Chrysotus) 

Chrysotus canadensis Van Duzee 1924 (Chrysotus) 

Chrysotus caroliniensis Robinson 1964 (Chrysotus) 

Chrysotus caudatus Van Duzee 1924 (Chrysotus) 

Chrysotus chlanoflavus Harmston and Knowlton 1940 (Chrysotus) 

Chrysotus choricus Wheeler 1890 (Chrysotus) 

Chrysotus ciliatus Malloch 1914 Syn. 

Chrysotus clypeatus Robinson 1967 (Chrysotus) 

Chrysotus coloradensis Van Duzee 1924 (Chrysotus) 

Chrysotus convergens Van Duzee 1924 (Chrysotus) 

Chrysotus cornutus Loew 1862 (Chrysotus) 

Chrysotus costalis Loew 1861 (Chrysotus) 

Chrysotus cressoni Van Duzee 1924 (Chrysotus) 

Chrysotus crosbyi Van Duzee 1924 (Chrysotus) 

Chrysotus currani Van Duzee 1924 (Chrysotus) 

Chrysotus dakotensis Harmston 1952 (Chrysotus) 

Chrysotus discolor Loew 1861 (Chrysotus) 

Chrysotus cobaltinus Van Duzee 1924 Syn. 

Chrysotus disjunctus Van Duzee 1924 (Chrysotus) 

Chrysotus distinctus Van Duzee 1924 (Chrysotus) 

Chrysotus dividuus Van Duzee 1924 (Chrysotus) 

Chrysotus dorsalis Van Duzee 1924 (Chrysotus) 

Chrysotus emarginatus Van Duzee 1928 (Chrysotus) 

Chrysotus excisus Aldrich 1896 (Chrysotus) 

Chrysotus exilis Van Duzee 1924 (Chrysotus) 

Chrysotus flavicauda Van Duzee 1928 (Chrysotus) 

Chrysotus flavisetus Malloch 1914 (Chrysotus) 

Chrysotus frontalis Van Duzee 1924 (Chrysotus) 

Chrysotus fulvohirtus Van Duzee 1915 (Chrysotus) 

Chrysotus furcatus Robinson 1964 (Chrysotus) 

Chrysotus gilvipes Van Duzee 1924 (Chrysotus) 

Chrysotus halteralis Van Duzee 1924 (Chrysotus) 

Chrysotus hastatus Van Duzee 1924 (Chrysotus) 

Chrysotus hirtipes Van Duzee 1924 (Chrysotus) 

Chrysotus dubius Van Duzee 1924 Syn. 

Chrysotus humilis Parent 1928 (Chrysotus) 

Chrysotus idahoensis Van Duzee 1924 (Chrysotus) 

Chrysotus incertus Walker 1849 (Chrysotus) 

Chrysotus intrudus Harmston 1951 (Chrysotus) 

Chrysotus johnsoni Van Duzee 1924 (Chrysotus) 

Chrysotus junctus Van Duzee 1924 (Chrysotus) 

Chrysotus kansensis Harmston 1952 (Chrysotus) 

Chrysotus lamellifer Robinson 1964 (Chrysotus) 

Chrysotus litoralis Robinson 1964 (Chrysotus) 

Chrysotus longihirtus Van Duzee 1932 (Chrysotus) 

Chrysotus longimanus Loew 1861 (Chrysotus) 

Chrysotus magnicornis Van Duzee 1924 (Chrysotus) 

Chrysotus major Van Duzee 1924 (Chrysotus) 

Chrysotus neopicticornis Robinson 1967 (Chrysotus) 

Chrysotus nigriciliatus Van Duzee 1933 (Chrysotus) 

Chrysotus nigrifrons Parent 1929 (Chrysotus) 

Chrysotus nigripalpis Van Duzee 1924 (Chrysotus) 

Chrysotus nubilus Say 1829 (Chrysotus) 

Chrysotus nudipes Van Duzee 1924 (Chrysotus) 

Chrysotus nudus Harmston and Knowlton 1963 (Chrysotus) 

Chrysotus obliquus Loew 1861 (Chrysotus) 

Chrysotus pallipes Loew 1861 (Chrysotus) 

Chrysotus palpiger Wheeler 1890 (Diaphorus) 

Chrysotus spinifer Malloch 1914 Syn. 

Chrysotus parvicornis Van Duzee 1924 (Chrysotus) 

Chrysotus exiguus Van Duzee 1924 Syn. 

Chrysotus parvulus Van Duzee 1924 (Chrysotus) 

Chrysotus pectoralis Van Duzee 1924 (Chrysotus) 

Chrysotus philtrum Melander 1903 (Chrysotus) 

Chrysotus picticornis Loew 1862 (Chrysotus) 

Chrysotus pomeroyi Parent 1934 (Chrysotus) 

Chrysotus rauterbergi Wheeler 1890 (Diaphorus) 

Chrysotus sagittarius Van Duzee 1924 (Chrysotus) 

Chrysotus silvicolus Harmston 1951 (Chrysotus) 

Chrysotus simulans Van Duzee 1924 (Chrysotus) 

Chrysotus subcostatus Loew 1864 (Chrysotus) 

Chrysotus pratincola Wheeler 1890 Syn. 

Chrysotus subjectus Van Duzee 1924 (Chrysotus) 

Chrysotus tarsalis Van Duzee 1924 (Chrysotus) 

Chrysotus teapanus Aldrich 1901 (Chrysotus) 

Chrysotus tennesseensis Robinson 1964 (Chrysotus) 

Chrysotus terminalis Van Duzee 1924 (Chrysotus) 

Chrysotus tibialis Van Duzee 1924 (Chrysotus) 

Chrysotus triangularis Van Duzee 1924 (Chrysotus) 

Chrysotus varipes Van Duzee 1924 (Chrysotus) 

Chrysotus viridifemora Macquart 1850 (Chrysotus) 

Chrysotus vividus Loew 1864 (Chrysotus) 

Chrysotus vulgaris Van Duzee 1924 (Chrysotus) 

Chrysotus wiconsinensis Wheeler 1890 (Chrysotus) 

Chrysotus xanthocal Harmston and Knowlton 1940 (Chrysotus) 

Condylostylus Bigot 1859 

Dasypilopus Bigot 1859 Syn. 

Aedisilopus Bigot 1859 Syn. 

Oedipsilopus Bigot 1859 Syn. 

Tylochaetus Bigot 1888 Syn. 

Laxina Curran 1934 Syn. 

Condylostylus albicoxa Walker 1849 (Psilopus) 

Condylostylus banksii Van Duzee 1915 (Sciapus) 

Condylostylus brimleyi Robinson 1964 (Condylostylus) 

Condylostylus calcaratus Loew 1861 (Psilopus) 

Condylostylus caudatus Wiedemann 1830 (Psilopus) 

Psilopus virgo Wiedemann 1830 Syn. 

Psilopus caudatulus Loew 1861 Syn. 

Condylostylus chalybeus Van Duzee 1914 (Sciapus) 

Condylostylus chrysoprasi Walker 1849 (Psilopus) 

Psilopus ciliipes Aldrich 1901 Syn.

Condylostylus clavatus Van Duzee 1929 (Psilopus) 

Condylostylus coloradensis Van Duzee 1932 (Condylostylus) 

Condylostylus comatus Loew 1861 (Psilopus) 

Condylostylus connectans Curran 1942 (Laxina) 

Condylostylus crinitus Aldrich 1904 (Psilopodinus) 

Condylostylus albiapicatus Parent 1929 Syn. 

Condylostylus currani Parent 1929 (Condylostylus) 

Condylostylus debilis Becker 1922 (Condylostylus) 

Condylostylus dimidiatus Loew 1862 (Psilopus) 

Condylostylus erectus Becker 1922 (Condylostylus) 

Condylostylus femoratus Say 1823 (Dolichopus) 

Condylostylus flavipes Aldrich 1904 (Psilopodinus) 

Condylostylus furcatus Van Duzee 1915 (Sciapus) 

Condylostylus fusitarsis Van Duzee 1933 (Condylostylus) 

Condylostylus graenicheri Van Duzee 1927 (Psilopus) 

Condylostylus inermis Loew 1861 (Psilopus) 

Condylostylus inornatus Aldrich 1901 (Psilopus) 

Condylostylus jucundus Loew 1861 (Psilopus) 

Condylostylus leonardi Van Duzee 1915 (Sciapus) 

Condylostylus longitalus Van Duzee 1923 (Psilopus) 

Condylostylus melampus Loew 1862 (Psilopus) 

Condylostylus mundus Wiedemann 1830 (Psilopus) 

Psilopus ciliatus Loew 1861 Syn. 

Condylostylus nigrofemoratus Walker 1849 (Psilopus) 

Psilopus scutellatus Harris 1835 Unav. 

Psilopus scobinator Loew 1861 Syn. 

Psilopus cockerelli Van Duzee 1927 Syn. 

Condylostylus occidentalis Bigot 1888 (Psilopodinus) 

Condylostylus ogilvii Malloch 1932 (Sciapus) 

Condylostylus parvicauda Van Duzee 1927 (Psilopus) 

Condylostylus patibulatus Say 1823 (Dolichopus) 

Psilopus amatus Walker 1849 Syn. 

Psilopodinus carolinensis Bigot 1888 Syn. 

Condylostylus pilicornis Aldrich 1904 (Psilopodinus) 

Condylostylus portoricensis Macquart 1834 (Psilopus) 

Condylostylus pruinosus Coquillett 1904 (Sciapus) 

Condylostylus quintusflavus Gunther 1980 (Condylostylus) 

Condylostylus radians Macquart 1834 (Psilopus) 

Condylostylus semicomatus Van Duzee 1929 (Psilopus) 

Condylostylus similis Aldrich 1901 (Psilopus) 

Condylostylus sipho Say 1823 (Dolichopus) 

Psilopus gemmifer Walker 1849 Syn. 

Psilopus scaber Loew 1861 Syn. 

Condylostylus tonsus Aldrich 1901 (Psilopus) 

Condylostylus viridicoxa Aldrich 1904 (Psilopodinus) 

Condylostylus viridis Parent 1929 (Condylostylus) 

Diaphorus Meigen 1824 

Diaphora Macquart 1834 Syn. 

Lyroneurus Loew 1857 Syn. 

Diaphorus aldrichi Van Duzee 1915 (Diaphorus) 

Diaphorus albifacies Parent 1929 Syn. 

Diaphorus alienus Van Duzee 1915 (Diaphorus) 

Diaphorus argentifacies Van Duzee 1932 (Diaphorus) 

Diaphorus australis Van Duzee 1915 (Diaphorus) 

Diaphorus femoratus Van Duzee 1915 Homo. 

Diaphorus bakeri Robinson 1967 (Diaphorus) 

Diaphorus basalis Van Duzee 1915 (Diaphorus) 

Diaphorus brevinervis Van Duzee 1924 (Diaphorus) 

Diaphorus californicus Van Duzee 1917 (Diaphorus) 

Diaphorus canus Robinson 1964 (Diaphorus) 

Diaphorus consanguineus Harmston and Knowlton 1963 

(Diaphorus) 

Diaphorus funeralis Parent 1929 (Diaphorus) 

Diaphorus fuscus Van Duzee 1921 (Diaphorus) 

Diaphorus adustus Van Duzee 1915 Homo. 

Diaphorus gibbosus Van Duzee 1915 (Diaphorus) 


Diaphorus hirsutus Van Duzee 1921 (Diaphorus) 

Diaphorus inornatus Van Duzee 1917 (Diaphorus) 

Diaphorus intermedius Robinson 1964 (Diaphorus) 

Diaphorus iowensis Robinson 1964 (Diaphorus) 

Diaphorus junctus Van Duzee 1917 (Diaphorus) 

Diaphorus laffooni Robinson 1964 (Diaphorus) 

Diaphorus lamellatus Loew 1864 (Diaphorus) 

Diaphorus latifacies Van Duzee 1932 (Diaphorus) 

Diaphorus leucostoma Loew 1861 (Diaphorus) 

Diaphorus vittatas Van Duzee 1915 Syn. 

Diaphorus infuscatus Van Duzee 1915 Syn. 

Diaphorus longilamellus Harmston 1971 (Diaphorus) 

Diaphorus mundus Loew 1861 (Diaphorus) 

Diaphorus nigricans Meigen 1824 (Diaphorus) 

Diaphorus opacus Loew 1861 Syn. 

Diaphorus nudus Van Duzee 1917 (Diaphorus) 

Diaphorus occidentalis Van Duzee 1915 (Diaphorus) 

Diaphorus parmatus Van Duzee 1915 (Diaphorus) 

Diaphorus pseudopacus Robinson 1964 (Diaphorus) 

Diaphorus crinipes Robinson 1964 Syn. 

Diaphorus quadratus Van Duzee 1915 (Diaphorus) 

Diaphorus remulus Van Duzee 1915 (Diaphorus) 

Diaphorus repandus Van Duzee 1915 (Diaphorus) 

Diaphorus similis Van Duzee 1915 (Diaphorus) 

Diaphorus slossonae Van Duzee 1932 (Diaphorus) 

Diaphorus snowii Van Duzee 1917 (Diaphorus) 

Diaphorus sodalis Loew 1861 (Diaphorus) 

Diaphorus sparsus Van Duzee 1917 (Diaphorus) 

Diaphorus spectabilis Loew 1861 (Diaphorus) 

Diaphorus spinitalus Van Duzee 1923 (Diaphorus) 

Diaphorus texanus Van Duzee 1932 (Diaphorus) 

Diaphorus triangulatus Van Duzee 1915 (Diaphorus) 

Diaphorus trivittatus Van Duzee 1915 (Diaphorus) 

Diaphorus usitatus Van Duzee 1915 (Diaphorus) 

Diaphorus vanduzeei Robinson 1964 (Diaphorus) 

Diaphorus communis Van Duzee 1932 Homo. 

Diaphorus variabilis Van Duzee 1915 (Diaphorus) 

Diaphorus versicolor Van Duzee 1932 (Diaphorus) 

Diaphorus vulsus Van Duzee 1917 (Diaphorus) 

Diostracus Loew 1861 

Asphyrotarsus Oldenberg 1916 Syn. 

Diostracus mchughi Harmston 1966 (Diostracus) 

Diostracus olga Aldrich 1911 (Diostracus) 

Diostracus prasinus Loew 1861 (Diostracus) 

Dolichopus Latreille 1796 

Ragheneura Rondani 1856 Syn. 

Hygroceleuthus Loew 1857 Syn. 

Spathichira Bigot 1888 Syn. 

Macrodolichopus Stackelberg 1933 Syn. 

Dolichopus abaftanus Harmston 1966 (Dolichopus) 

Dolichopus abbreviatus Van Duzee 1921 (Dolichopus) 

Dolichopus aboriginis Harmston and Knowlton 1943 (Dolichopus) 

Dolichopus abrasus Van Duzee 1921 (Dolichopus) 

Dolichopus abruptus Aldrich 1922 (Dolichopus) 

Dolichopus absonus Van Duzee 1921 (Dolichopus) 

Dolichopus accidentalis Harmston and Knowlton 1941 

(Dolichopus) 

Dolichopus acricola Van Duzee 1921 (Dolichopus) 

Dolichopus acuminatus Loew 1861 (Dolichopus) 

Dolichopus acutus Van Duzee 1921 (Dolichopus) 

Dolichopus adaequatus Van Duzee 1921 (Dolichopus) 

Dolichopus adjacens Walker 1849 (Dolichopus) 

Dolichopus adultus Van Duzee 1921 (Dolichopus) 

Dolichopus aequalis Van Duzee 1921 (Dolichopus) 


139

140 



Dolichopus aeratus Van Duzee 1921 (Dolichopus) 

Dolichopus aethiops Van Duzee 1921 (Dolichopus) 

Dolichopus affinis Walker 1849 (Dolichopus) 

Dolichopus splendidulus Loew 1864 Syn. 

Dolichopus afflictus Osten Sacken 1877 (Hygroceleuthus) 

Dolichopus affluens Van Duzee 1921 (Dolichopus) 

Dolichopus agronomus Melander and Brues 1900 (Dolichopus) 

Dolichopus ainsliei Van Duzee 1921 (Dolichopus) 

Dolichopus alacer Van Duzee 1921 (Dolichopus) 

Dolichopus alberecrus Gunther 1979 (Dolichopus) 

Dolichopus albertensis Curran 1922 (Dolichopus) 

Dolichopus albiciliatus Loew 1862 (Dolichopus) 

Dolichopus atrovirens Johnson 1925 Unav. 

Dolichopus albicoxa Aldrich 1893 (Dolichopus) 

Dolichopus albivestitarsis Robinson 1964 (Dolichopus) 

Dolichopus aldrichii Wheeler 1899 (Hygroceleuthus) 

Dolichopus amnicola Melander and Brues 1900 (Hygroceleuthus) 

Dolichopus robertsoni Curran 1923 Syn. 

Dolichopus amphericus Melander and Brues 1900 (Dolichopus) 

Dolichopus amplipennis Van Duzee 1921 (Dolichopus) 

Dolichopus andersoni Curran 1924 (Dolichopus) 

Dolichopus angustatus Aldrich 1893 (Dolichopus) 

Dolichopus angusticornis Van Duzee 1921 (Dolichopus) 

Dolichopus annaclareii Gunther 1979 (Dolichopus) 

Dolichopus apheles Melander and Brues 1900 (Dolichopus) 

Dolichopus appendiculatus Van Duzee 1921 (Dolichopus) 

Dolichopus argentipes Van Duzee 1921 (Dolichopus) 

Dolichopus arizonicus Harmston 1951 (Dolichopus) 

Dolichopus aurifacies Aldrich 1893 (Dolichopus) 

Dolichopus aurifex Van Duzee 1921 (Dolichopus) 

Dolichopus bakeri Cole 1912 (Dolichopus) 

Dolichopus barbaricus Van Duzee 1921 (Dolichopus) 

Dolichopus barbicauda Van Duzee 1921 (Dolichopus) 

Dolichopus barbipes Van Duzee 1921 (Dolichopus) 

Dolichopus barycnemus Coquillett 1900 (Dolichopus) 

Dolichopus beameri Harmston and Knowlton 1941 (Dolichopus) 

Dolichopus beatus Van Duzee 1921 (Dolichopus) 

Dolichopus bifractus Loew 1861 (Dolichopus) 

Dolichopus bisetosus Van Duzee 1921 (Dolichopus) 

Dolichopus blandus Van Duzee 1921 (Dolichopus) 

Dolichopus bolsteri Van Duzee 1921 (Dolichopus) 

Dolichopus brevicauda Van Duzee 1921 (Dolichopus) 

Dolichopus breviciliatus Van Duzee 1930 (Dolichopus) 

Dolichopus brevimanus Loew 1861 (Dolichopus) 

Dolichopus brevipennis Meigen 1824 (Dolichopus) 

Dolichopus brevipilosus Van Duzee 1933 (Dolichopus) 

Dolichopus breviciliatus Van Duzee 1933 Homo. 

Dolichopus bruesi Van Duzee 1921 (Dolichopus) 

Hygroceleuthus propinquus Melander and Brues 1900 Homo. 

Dolichopus bruneifacies Van Duzee 1933 (Dolichopus) 

Dolichopus brunneus Aldrich 1893 (Dolichopus) 

Dolichopus bryanti Van Duzee 1921 (Dolichopus) 

Dolichopus burnesi Van Duzee 1921 (Dolichopus) 

Dolichopus calainus Melander and Brues 1900 (Dolichopus) 

Dolichopus calcaratus Aldrich 1893 (Dolichopus) 

Dolichopus californicus Van Duzee 1921 (Dolichopus) 

Dolichopus calvimontis James 1939 (Dolichopus) 

Dolichopus canadensis Van Duzee 1921 (Dolichopus) 

Dolichopus canaliculatus Thomson 1869 (Dolichopus) 

Dolichopus carolinensis Van Duzee 1921 (Dolichopus) 

Dolichopus cavatus Van Duzee 1921 (Dolichopus) 

Dolichopus celeripes Van Duzee 1921 (Dolichopus) 

Dolichopus chrysostomus Loew 1861 (Dolichopus) 

Dolichopus coercens Walker 1849 (Dolichopus) 

Dolichopus conterminus Walker 1849 Syn. 

Dolichopus batillifer Loew 1861 Syn. 

Dolichopus coloradensis Aldrich 1893 (Dolichopus) 

Dolichopus agilis Aldrich 1893 Homo. 

Dolichopus comatus Loew 1861 (Dolichopus) 

Dolichopus cornutus Johnson 1910 Missp. 

Dolichopus compactus Van Duzee 1921 (Dolichopus) 

Dolichopus completus Van Duzee 1921 (Dolichopus) 

Dolichopus comptus Van Duzee 1921 (Dolichopus) 

Dolichopus confinis Walker 1849 (Dolichopus) 

Dolichopus consanguineus Wheeler 1899 (Hygroceleuthus) 

Dolichopus conspectus Van Duzee 1921 (Dolichopus) 

Dolichopus conspicuus Van Duzee 1921 Missp. 

Dolichopus contiguus Walker 1849 (Dolichopus) 

Dolichopus splendidus Loew 1861 Syn. 

Dolichopus contingens Walker 1852 (Dolichopus) 

Dolichopus convergens Aldrich 1893 (Dolichopus) 

Dolichopus coquilletti Aldrich 1893 (Dolichopus) 

Dolichopus corax Osten Sacken 1877 (Dolichopus) 

Dolichopus correus Steyskal 1959 (Dolichopus) 

Dolichopus crassicornis Aldrich 1922 (Dolichopus) 

Dolichopus crassitibia Robinson 1967 (Dolichopus) 

Dolichopus crenatus Osten Sacken 1877 (Hygroceleuthus) 

Dolichopus cuprinus Wiedemann 1830 (Dolichopus) 

Dolichopus cupreus Say 1823 Homo. 

Dolichopus czekanowskii Stackelberg 1928 (Dolichopus) 

Dolichopus dakotensis Aldrich 1893 (Dolichopus) 

Dolichopus dasyops Malloch 1919 (Dolichopus) 

Dolichopus dasypodus Coquillett 1910 (Dolichopus) 

Dolichopus decorus Van Duzee 1921 (Dolichopus) 

Dolichopus defectus Van Duzee 1921 (Dolichopus) 

Dolichopus delicatus Aldrich 1922 (Dolichopus) 

Dolichopus demissus Van Duzee 1921 (Dolichopus) 

Dolichopus detersus Loew 1866 (Dolichopus) 

Dolichopus digitus Van Duzee 1921 (Dolichopus) 

Dolichopus discessus Walker 1849 (Dolichopus) 

Dolichopus discolor Van Duzee 1921 (Dolichopus) 

Dolichopus distractus Walker 1849 (Dolichopus) 

Dolichopus diversipennis Curran 1922 (Dolichopus) 

Dolichopus divigatus Harmston 1952 (Dolichopus) 

Dolichopus dolosus Parent 1934 (Dolichopus) 

Dolichopus domesticus Van Duzee 1921 (Dolichopus) 

Dolichopus dorsalis Van Duzee 1921 (Dolichopus) 

Dolichopus dorycerus Loew 1864 (Dolichopus) 

Dolichopus duplicatus Aldrich 1893 (Dolichopus) 

Dolichopus elegans Aldrich 1922 (Dolichopus) 

Dolichopus enigma Melander and Brues 1900 (Dolichopus) 

Dolichopus eudactylus Loew 1861 (Dolichopus) 

Dolichopus evolvens Parent 1929 (Dolichopus) 

Dolichopus exclusus Walker 1849 (Dolichopus) 

Dolichopus facirecedens Harmston and Knowlton 1939 

(Dolichopus) 

Dolichopus factivittatus Harmston 1966 (Dolichopus) 

Dolichopus fallax Van Duzee 1933 (Dolichopus) 

Dolichopus finitus Walker 1849 (Dolichopus) 

Dolichopus atricornis Harris 1835 Unav. 

Dolichopus scoparius Loew 1864 Syn. 

Dolichopus flagellitenens Wheeler 1890 (Dolichopus) 

Dolichopus flaviciliatus Van Duzee 1921 (Dolichopus) 

Dolichopus flavicoxa Van Duzee 1921 (Dolichopus) 

Dolichopus flavifacies Van Duzee 1933 (Dolichopus) 

Dolichopus flavilacertus Van Duzee 1921 (Dolichopus) 

Dolichopus footei Harmston 1966 (Dolichopus) 

Dolichopus formosus Van Duzee 1921 (Dolichopus) 

Dolichopus fortis Aldrich 1922 (Dolichopus) 

Dolichopus frontalis Van Duzee 1928 (Dolichopus) 

Dolichopus fucatus Van Duzee 1921 (Dolichopus) 

Dolichopus fulgerus Harmston 1966 (Dolichopus) 

Dolichopus fulvipes Loew 1862 (Dolichopus) 

Dolichopus fumosus Van Duzee 1921 (Dolichopus) 

Dolichopus funditor Loew 1861 (Dolichopus) 

Dolichopus distinctus Van Duzee 1921 Syn.


Dolichopus gaigei Steyskal 1973 (Dolichopus) 

Dolichopus genualis Van Duzee 1921 (Dolichopus) 

Dolichopus gladius Van Duzee 1921 (Dolichopus) 

Dolichopus grandis Aldrich 1893 (Dolichopus) 

Dolichopus gratiolus Steyskal 1973 (Dolichopus) 

Dolichopus gratus Loew 1861 (Dolichopus) 

Dolichopus mercieri Parent 1929 Syn. 

Dolichopus groenlandicus Zetterstedt 1843 (Dolichopus) 

Dolichopus harbecki Van Duzee 1921 (Dolichopus) 

Dolichopus hardyi Harmston 1951 (Dolichopus) 

Dolichopus hastatus Loew 1864 (Dolichopus) 

Dolichopus helenae James 1939 (Dolichopus) 

Dolichopus hirsutitarsis Harmston 1952 (Dolichopus) 

Dolichopus humilis Van Duzee 1921 (Dolichopus) 

Dolichopus idahoensis Aldrich 1894 (Hygroceleuthus) 

Dolichopus idoneus Van Duzee 1921 (Dolichopus) 

Dolichopus imperfectus Van Duzee 1921 (Dolichopus) 

Dolichopus incisuralis Loew 1861 (Dolichopus) 

Dolichopus platyprosopus Loew 1866 Syn. 

Dolichopus incongruus Wheeler 1890 (Dolichopus) 

Dolichopus indianus Harmston and Knowlton 1946 (Dolichopus) 

Dolichopus indigenus Van Duzee 1921 (Dolichopus) 

Dolichopus inflatus Aldrich 1922 (Dolichopus) 

Dolichopus integripes Parent 1929 (Dolichopus) 

Dolichopus intentus Melander and Brues 1900 (Dolichopus) 

Dolichopus interjectus Van Duzee 1923 (Dolichopus) 

Dolichopus iowaensis Harmston and Knowlton 1939 (Dolichopus) 

Dolichopus jaquesi Harmston and Knowlton 1939 (Dolichopus) 

Dolichopus johnsoni Aldrich 1893 (Dolichopus) 

Dolichopus jugalis Tucker 1911 (Dolichopus) 

Dolichopus procerus Van Duzee 1921 Syn. 

Dolichopus kansensis Aldrich 1893 (Dolichopus) 

Dolichopus kleini Curran 1934 (Dolichopus) 

Dolichopus kyphotus Harmston 1966 (Dolichopus) 

Dolichopus laciniatus Coquillett 1910 (Dolichopus) 

Dolichopus lamellicornis Thomson 1869 (Dolichopus) 

Dolichopus lamellipes Walker 1849 (Dolichopus) 

Dolichopus boreus Van Duzee 1921 Syn. 

Dolichopus laticornis Loew 1861 (Dolichopus) 

Dolichopus latipes Loew 1861 (Hygroceleuthus) 

Hygroceleuthus cognatus Melander and Brues 1900 Syn. 

Dolichopus latronis Van Duzee 1921 (Dolichopus) 

Dolichopus leucacra James 1939 (Dolichopus) 

Dolichopus litoralis Van Duzee 1921 (Dolichopus) 

Dolichopus lobatus Loew 1861 (Dolichopus) 

Dolichopus longicornis Stannius 1831 (Dolichopus) 

Dolichopus longimanus Loew 1861 (Dolichopus) 

Dolichopus longipennis Loew 1861 (Dolichopus) 

Dolichopus longus Aldrich 1922 (Dolichopus) 

Dolichopus lundbecki Curran 1923 (Dolichopus) 

Dolichopus luteipennis Loew 1861 (Dolichopus) 

Dolichopus greenei Van Duzee 1921 Syn. 

Dolichopus maculitarsis Van Duzee 1925 (Dolichopus) 

Dolichopus magnantenna James 1939 (Dolichopus) 

Dolichopus manicula Van Duzee 1921 (Dolichopus) 

Dolichopus mannerheimi Zetterstedt 1838 (Dolichopus) 

Dolichopus marginatus Aldrich 1893 (Dolichopus) 

Dolichopus melanderi Van Duzee 1921 (Dolichopus) 

Dolichopus melanocerus Loew 1864 (Dolichopus) 

Dolichopus micropygus Wahlberg 1851 (Dolichopus) 

Dolichopus monarchus Harmston 1968 (Dolichopus) 

Dolichopus monticola Van Duzee 1921 (Dolichopus) 

Dolichopus multisetosus Van Duzee 1921 (Dolichopus) 

Dolichopus myosotus Osten Sacken 1887 (Dolichopus) 

Dolichopus neomexicanus Harmston 1951 (Dolichopus) 

Dolichopus nigricauda Van Duzee 1921 (Dolichopus) 

Dolichopus nigricornis Meigen 1824 (Dolichopus) 

Dolichopus discifer Stannius 1831 Syn. 

Dolichopus tanypus Loew 1861 Syn. 

Dolichopus nigricoxa Van Duzee 1926 (Dolichopus) 

Dolichopus nigrilineatus Van Duzee 1924 (Dolichopus) 

Dolichopus nigrimanus Van Duzee 1921 (Dolichopus) 

Dolichopus nigroapicalis Van Duzee 1930 (Dolichopus) 

Dolichopus nigropleurus Harmston 1966 (Dolichopus) 

Dolichopus nodipennis Van Duzee 1921 (Dolichopus) 

Dolichopus nomadus Harmston and Knowlton 1942 (Dolichopus) 

Dolichopus nubifer Van Duzee 1921 (Dolichopus) 

Dolichopus nudus Loew 1864 (Dolichopus) 

Dolichopus obcordatus Aldrich 1893 (Dolichopus) 

Dolichopus obsoletus Van Duzee 1921 (Dolichopus) 

Dolichopus occidentalis Aldrich 1893 (Dolichopus) 

Dolichopus reticulus Van Duzee 1926 Syn. 

Dolichopus omnivagus Van Duzee 1921 (Dolichopus) 

Dolichopus opportunus Van Duzee 1921 (Dolichopus) 

Dolichopus oregonensis Van Duzee 1927 (Dolichopus) 

Dolichopus ornatipennis Van Duzee 1921 (Dolichopus) 

Dolichopus ovatus Loew 1861 (Dolichopus) 

Dolichopus pachycnemus Loew 1861 (Dolichopus) 

Dolichopus packardi Van Duzee 1921 (Dolichopus) 

Dolichopus palaestricus Loew 1864 (Dolichopus) 

Dolichopus paluster Melander and Brues 1900 (Dolichopus) 

Dolichopus pantomimus Melander and Brues 1900 (Dolichopus) 

Dolichopus partitus Melander and Brues 1900 (Dolichopus) 

Dolichopus parvicornis Van Duzee 1921 (Dolichopus) 

Dolichopus parvimanus Van Duzee 1933 (Dolichopus) 

Dolichopus penicillatus Van Duzee 1921 (Dolichopus) 

Hygroceleuthus ciliatus Aldrich 1893 Homo. 

Dolichopus pensus Aldrich 1922 (Dolichopus) 

Dolichopus pernix Melander and Brues 1900 (Dolichopus) 

Dolichopus perplexus Van Duzee 1923 (Dolichopus) 

Dolichopus misellus Melander 1900 Homo. 

Dolichopus melandri Becker 1922 Syn. (?) 

Dolichopus melanderi Foote et al. 1965 Emend. 

Dolichopus phyllocerus Vockeroth 1962 (Dolichopus) 

Dolichopus pilatus Van Duzee 1921 (Dolichopus) 

Dolichopus pingreensis James 1939 (Dolichopus) 

Dolichopus plumipes Scopoli 1763 (Musca) 

Dolichopus ciliatus Walker 1849 Syn. 

Dolichopus sequax Walker 1849 Syn. 

Dolichopus plumitarsis Fallen 1823 (Dolichopus) 

Dolichopus plumosus Aldrich 1893 (Dolichopus) 

Dolichopus pollex Osten Sacken 1877 (Dolichopus) 

Dolichopus porphyrops Van Duzee 1921 (Dolichopus) 

Dolichopus praeustus Loew 1862 (Dolichopus) 

Dolichopus puberiseta Parent 1934 (Dolichopus) 

Dolichopus pugil Loew 1866 (Dolichopus) 

Dolichopus henshawi Wheeler 1890 Syn. 

Dolichopus pulcher Walker 1852 (Dolichopus) 

Dolichopus pulchrimana Bigot 1888 (Spathichira) 

Dolichopus willistonii Aldrich 1893 Syn. 

Dolichopus quadrilamellatus Loew 1864 (Dolichopus) 

Dolichopus ramifer Loew 1861 (Dolichopus) 

Dolichopus recticosta Aldrich 1922 (Dolichopus) 

Dolichopus reflectus Aldrich 1893 (Dolichopus) 

Dolichopus refulgens Harmston and Knowlton 1939 (Dolichopus) 

Dolichopus remipes Wahlberg 1839 (Dolichopus) 

Dolichopus remotus Walker 1849 (Dolichopus) 

Dolichopus cuniculus Van Duzee 1921 Syn. 

Dolichopus remus Van Duzee 1921 (Dolichopus) 

Dolichopus renidescens Melander and Brues 1900 (Dolichopus) 

Dolichopus retinens Van Duzee 1921 (Dolichopus) 

Dolichopus ruficornis Loew 1861 (Dolichopus) 

Dolichopus rupestris Haliday 1833 (Dolichopus) 

Dolichopus festinans Zetterstedt 1838 Syn. 

Dolichopus sarotes Loew 1866 (Dolichopus) 

Dolichopus scapularis Loew 1861 (Dolichopus) 


141

142 



Dolichopus scopifer James 1939 (Dolichopus) 

Dolichopus sedulus Van Duzee 1921 (Dolichopus) 

Dolichopus separatus Walker 1849 (Dolichopus) 

Dolichopus serratus Van Duzee 1921 (Dolichopus) 

Dolichopus setifer Loew 1861 (Dolichopus) 

Dolichopus michiganus Harmston and Knowlton 1945 Syn. 

Dolichopus setosus Loew 1862 (Dolichopus) 

Dolichopus sexarticulatus Loew 1864 (Dolichopus) 

Dolichopus shastaensis Harmston 1966 (Dolichopus) 

Dolichopus shelfordi Curran 1934 (Dolichopus) 

Dolichopus sicarius Van Duzee 1921 (Dolichopus) 

Dolichopus silvicola Harmston 1951 (Dolichopus) 

Dolichopus simplicipes Aldrich 1922 (Dolichopus) 

Dolichopus simulans Van Duzee 1926 (Dolichopus) 

Dolichopus sincerus Melander 1900 (Dolichopus) 

Dolichopus subdirectus Van Duzee 1921 Syn. 

Dolichopus sinualaris Harmston 1966 (Dolichopus) 

Dolichopus slossonae Van Duzee 1921 (Dolichopus) 

Dolichopus smithae Harmston 1966 (Dolichopus) 

Dolichopus soccatus Walker 1849 (Dolichopus) 

Dolichopus socius Loew 1862 (Dolichopus) 

Dolichopus solidus Van Duzee 1921 (Dolichopus) 

Dolichopus sordidatus Van Duzee 1921 (Dolichopus) 

Dolichopus speciosus Van Duzee 1921 (Dolichopus) 

Dolichopus sphaeristes Brues 1901 (Dolichopus) 

Dolichopus sporadicum Harmston and Knowlton 1942 (Dolichopus) 

Dolichopus squamiciliatus Harmston 1966 (Dolichopus) 

Dolichopus squamosus Van Duzee 1921 (Dolichopus) 

Dolichopus stenhammari Zetterstedt 1843 (Dolichopus) 

Dolichopus annulipes Zetterstedt 1838 Homo. 

Dolichopus steyskali Robinson 1964 (Dolichopus) 

Dolichopus stricklandi Harmston and Knowlton 1939 (Dolichopus) 

Dolichopus subciliatus Loew 1864 (Dolichopus) 

Dolichopus subcostatus Van Duzee 1930 (Dolichopus) 

Dolichopus subspina Van Duzee 1928 (Dolichopus) 

Dolichopus sufflavus Van Duzee 1921 (Dolichopus) 

Dolichopus superbus Van Duzee 1921 (Dolichopus) 

Dolichopus talus Van Duzee 1921 (Dolichopus) 

Dolichopus tarsipictis Harmston and Knowlton 1963 (Dolichopus) 

Dolichopus tener Loew 1861 (Dolichopus) 

Dolichopus tenuimanus Van Duzee 1932 (Dolichopus) 

Dolichopus tenuipes Aldrich 1894 (Dolichopus) 

Dolichopus terminalis Loew 1866 (Dolichopus) 

Dolichopus germanus Wheeler 1890 Homo. 

Dolichopus terminatus Walker 1849 (Dolichopus) 

Dolichopus tetricus Loew 1864 (Dolichopus) 

Dolichopus tonsus Loew 1861 (Dolichopus) 

Dolichopus townsendi Aldrich 1922 (Dolichopus) 

Dolichopus trisetosus Van Duzee 1921 (Dolichopus) 

Dolichopus uliginosus Van Duzee 1923 (Dolichopus) 

Dolichopus umbrosus Van Duzee 1921 (Dolichopus) 

Dolichopus ungulata Linnaeus 1758 (Musca) 

Nemotelus aeneus De Geer 1776 Syn. 

Dolichopus utahensis Harmston and Knowlton 1943 (Dolichopus) 

Dolichopus uxorcula Van Duzee 1921 (Dolichopus) 

Dolichopus vanduzeei Curran 1922 (Dolichopus) 

Dolichopus variabilis Loew 1861 (Dolichopus) 

Dolichopus gracilis Aldrich 1893 Syn. 

Dolichopus varipes Coquillett 1900 (Dolichopus) 

Dolichopus vegetus Harmston 1952 (Dolichopus) 

Dolichopus vernaae Harmston and Knowlton 1940 (Dolichopus) 

Dolichopus versutus Van Duzee 1921 (Dolichopus) 

Dolichopus vigilans Aldrich 1893 (Dolichopus) 

Dolichopus virga Coquillett 1910 (Dolichopus) 

Dolichopus virginiensis Van Duzee 1921 (Dolichopus) 

Dolichopus lobipennis Van Duzee 1930 Syn. 

Dolichopus viridis Van Duzee 1921 (Dolichopus) 

Dolichopus vittatus Loew 1861 (Dolichopus) 

Dolichopus walkeri Van Duzee 1921 (Dolichopus) 

Dolichopus wheeleri Melander and Brues 1900 (Hygroceleuthus) 

Dolichopus xanthocnemus Loew 1864 (Dolichopus) 

Dolichopus zygomus Harmston 1966 (Dolichopus) 

Enlinia Aldrich 1933 

Collinellula Aldrich 1932 Homo. 

Enlinia arizonica Robinson 1973 (Enlinia) 

Enlinia californica Robinson and Arnaud 1970 (Enlinia) 

Enlinia ciliata Robinson 1964 (Enlinia) 

Enlinia magistri Aldrich 1932 (Collinellula) 

Enlinia saxicola Robinson 1964 (Enlinia) 

Enlinia taeniocaudata Robinson and Arnaud 1970 (Enlinia) 

Enlinia texana Robinson 1973 (Enlinia) 

Harmstonia Robinson 1964 

Harmstonia intricata Robinson 1964 (Harmstonia) 

Harmstonia pectinicauda Robinson 1964 (Harmstonia) 

Hercostomus Loew 1857 

Acanthipodus Rondani 1856 Syn. (?) 

Gymnopternus Loew 1857 Syn. 

Dasyarthrus Mik 1878 Syn. 

Poecilobothrus Mik 1878 Syn. 

Hercostomus albiceps Loew 1861 (Gymnopternus) 

Hercostomus albipodus Harmston and Knowlton 1941 

(Hercostomus) 

Hercostomus aldrichi Robinson 1964 (Gymnopternus) 

Hercostomus anarmostus Melander 1900 (Hercostomus) 

Hercostomus annulatus Van Duzee 1926 (Gymnopternus) 

Hercostomus annulipes Robinson 1964 (Gymnopternus) 

Hercostomus anomalocerus Robinson 1964 (Gymnopternus) 

Hercostomus aurifer Thomson 1869 (Dolichopus) 

Dolichopus metatarsalis Thomson 1869 Syn. 

Hercostomus procerus Wheeler 1899 Syn. 

Hercostomus barbatulus Loew 1861 (Gymnopternus) 

Hercostomus brevipes Van Duzee 1933 (Hercostomus) 

Hercostomus brunneifacies Robinson 1960 (Hercostomus) 

Hercostomus cacheae Harmston and Knowlton 1941 (Hercostomus) 

Hercostomus californicus Van Duzee 1920 (Gymnopternus) 

Hercostomus campsicnemoides Robinson 1964 (Gymnopternus) 

Hercostomus chaetilamellus Harmston and Knowlton 1941 

(Hercostomus) 

Hercostomus chetifer Walker 1849 (Hercostomus) 

Neurigona ornatus Van Duzee 1921 Syn. 

Paraclius ornatipes Curran 1925 Missp. 

Hercostomus dreisbachi Harmston and Knowlton 1945 Syn. 

Hercostomus coloradensis Harmston 1952 (Hercostomus) 

Hercostomus consanguineus Harmston 1952 (Hercostomus) 

Hercostomus constrictus Robinson 1964 (Gymnopternus) 

Hercostomus costalis Van Duzee 1923 (Hercostomus) 

Hercostomus coxalis Loew 1864 (Gymnopternus) 

Hercostomus crassicauda Loew 1861 (Gymnopternus) 

Hercostomus cryptus Harmston and Knowlton 1941 (Hercostomus) 

Hercostomus cumberlandensis Robinson 1964 (Gymnopternus) 

Hercostomus cuneicornis Robinson 1964 (Gymnopternus) 

Hercostomus currani Van Duzee 1930 (Hercostomus) 

Hercostomus debilis Loew 1861 (Gymnopternus) 

Hercostomus despicatus Loew 1861 (Gymnopternus) 

Hercostomus difficilis Loew 1861 (Gymnopternus) 

Hercostomus dorsalis Van Duzee 1914 (Neurigona) 

Hercostomus exilis Loew 1861 (Gymnopternus) 

Hercostomus fimbriatus Loew 1861 (Gymnopternus) 

Hercostomus flaviciliatus Van Duzee 1914 (Gymnopternus) 

Hercostomus flavitarsis Van Duzee 1925 (Hercostomus)


Hercostomus flaviventris Robinson 1964 (Gymnopternus) 

Hercostomus flavus Loew 1861 (Gymnopternus) 

Hercostomus floridensis Robinson 1964 (Gymnopternus) 

Hercostomus flutatus Harmston and Knowlton 1945 (Hercostomus) 

Hercostomus frequens Loew 1861 (Gymnopternus) 

Hercostomus humilis Loew 1864 (Gymnopternus) 

Gymnopternus humeralis Criddle 1921 Missp. 

Hercostomus hybridus Robinson 1964 (Gymnopternus) 

Hercostomus impudicus Wheeler 1899 (Hercostomus) 

Hercostomus indianus Harmston 1952 (Hercostomus) 

Gymnopternus tibialis Van Duzee 1928 Homo. 

Hercostomus laevigatus Loew 1861 (Gymnopternus) 

Hercostomus laffooni Robinson 1964 (Gymnopternus) 

Hercostomus lividifrons Van Duzee 1926 (Gymnopternus) 

Hercostomus longilamellus Harmston and Knowlton 1940 

(Hercostomus) 

Hercostomus lunifer Loew 1861 (Gymnopternus) 

Gymnopternus exiguus Loew 1864 Syn. 

Hercostomus maculiventris Van Duzee 1925 (Hercostomus) 

Hercostomus meniscoides Robinson 1964 (Gymnopternus) 

Hercostomus meniscus Loew 1864 (Gymnopternus) 

Hercostomus minutus Loew 1861 (Gymnopternus) 

Hercostomus mirificus Melander 1900 (Gymnopternus) 

Hercostomus neocryptus Harmston and Knowlton 1941 

(Hercostomus) 

Hercostomus nigribarbus Loew 1861 (Gymnopternus) 

Gymnopternus phyllophorus Loew 1866 Syn. 

Hercostomus nigricomus Robinson 1964 (Gymnopternus) 

Hercostomus nigricoxa Van Duzee 1924 (Gymnopternus) 

Gymnopternus nigricera Leonard 1928 Missp. 

Hercostomus nigrifacies Van Duzee 1933 (Hercostomus) 

Hercostomus obscurus Say 1823 (Dolichopus) 

Hercostomus obtusicauda Van Duzee 1924 (Gymnopternus) 

Hercostomus occidentalis Cole 1912 (Hercostomus) 

Paraclius flavicornis Van Duzee 1918 Syn. 

Hercostomus ohioensis Robinson 1964 (Gymnopternus) 

Hercostomus opacus Loew 1861 (Gymnopternus) 

Hercostomus orbicularis Harmston 1952 (Hercostomus) 

Hercostomus ovaticornis Van Duzee 1933 (Hercostomus) 

Hercostomus pallidiciliatus Van Duzee 1930 (Hercostomus) 

Hercostomus parvicornis Loew 1861 (Gymnopternus) 

Hercostomus politus Loew 1861 (Gymnopternus) 

Hercostomus propriofacies Robinson 1964 (Gymnopternus) 

Hercostomus pseudodebilis Robinson 1964 (Gymnopternus) 

Hercostomus purpuratus Van Duzee 1925 (Hercostomus) 

Hercostomus purus Harmston and Knowlton 1963 (Hercostomus) 

Hercostomus pusillus Loew 1864 (Gymnopternus) 

Hercostomus robustus Van Duzee 1925 (Hercostomus) 

Hercostomus schlingeri Harmston and Knowlton 1963 

(Hercostomus) 

Hercostomus scotias Loew 1861 (Gymnopternus) 

Hercostomus browni Van Duzee 1933 Syn. 

Hercostomus setosa Van Duzee 1913 (Neurigona) 

Hercostomus torridus Harmston and Knowlton 1941 Syn. 

Hercostomus sharpi Robinson 1964 (Gymnopternus) 

Hercostomus singularis Van Duzee 1924 (Gymnopternus) 

Hercostomus spectabilis Loew 1861 (Gymnopternus) 

Gymnopternus chalcochrus Loew 1864 Syn. 

Hercostomus stanfordi Harmston and Knowlton 1940 

(Hercostomus) 

Hercostomus subdilatatus Loew 1861 (Gymnopternus) 

Hercostomus subulatus Loew 1861 (Gymnopternus) 

Hercostomus synolcus Steyskal 1966 (Hercostomus) 

Gymnopternus convergens Van Duzee 1920 Homo. 

Hercostomus tennesseensis Robinson 1964 (Gymnopternus) 

Hercostomus tenuicauda Van Duzee 1928 (Gymnopternus) 

Hercostomus tibialis Van Duzee 1913 (Neurigona) 

Hercostomus tristis Loew 1864 (Gymnopternus) 

Hercostomus truncatus Harmston and Knowlton 1940 

(Hercostomus) 

Hercostomus unicolor Loew 1864 (Hercostomus) 

Gymnopternus poenitens Wheeler 1890 Syn. 

Hercostomus ornaticauda Van Duzee 1933 Syn. 

Hercostomus utahensis Harmston and Knowlton 1940 

(Hercostomus) 

Hercostomus vanduzeei Curran 1930 (Gymnopternus) 

Hercostomus ventralis Loew 1861 (Gymnopternus) 

Hercostomus vernaculus Van Duzee 1924 (Gymnopternus) 

Hercostomus vetitus Melander 1900 (Hercostomus) 

Hercostomus violaceus Van Duzee 1921 (Proarchus) 

Hercostomus vockerothi Robinson 1964 (Gymnopternus) 

Hercostomus wasatchensis Harmston and Knowlton 1943 

(Hercostomus) 

Hercostomus weemsi Robinson 1964 (Gymnopternus) 

Hydatostega Philippi 1865 

Millardia Curran 1934 Homo. 

Hydatostega cerutias Loew 1872 (Hydrophorus) 

Hydatostega plumbeus Aldrich 1911 (Hydrophorus) 

Hydrophorus albomaculatus Van Duzee 1926 Syn. 

Hydatostega viridiflos Walker 1852 (Medeterus) 

Hydrophorus gratiosus Aldrich 1911 Syn. 

Hydrophorus intentus Aldrich 1911 Syn. 

Hydrophorus argentatus Van Duzee 1918 Syn. 

Hydrophorus fulvidorsum Van Duzee 1925 Syn. 

Hydrophorus browni Curran 1930 Syn. 

Hydrophorus Fallen 1823 

Aphrozeta Perris 1847 Syn. 

Parhydrophorus Wheeler 1896 Syn. 

Hydrophorus aestuum Loew 1869 (Hydrophorus) 

Hydrophorus agalma Wheeler 1899 (Hydrophorus) 

Hydrophorus alboflorens Walker 1849 (Medeterus) 

Hydrophorus pirata Loew 1861 Syn. 

Hydrophorus algens Wheeler 1899 (Hydrophorus) 

Hydrophorus hirtipes Van Duzee 1933 Syn. 

Hydrophorus alpinus Wahlberg 1844 (Hydrophorus) 

Hydrophorus epakter Hurley 1985 Syn. 

Hydrophorus altivagus Aldrich 1911 (Hydrophorus) 

Hydrophorus wahlgreni Frey 1915 Syn. 

Hydrophorus lividipes Van Duzee 1926 Syn. 

Hydrophorus amplectens Aldrich 1911 (Hydrophorus) 

Hydrophorus ampullaceus Van Duzee 1924 (Hydrophorus) 

Hydrophorus angustifacies Hurley 1985 (Hydrophorus) 

Hydrophorus aquatilis Aldrich 1922 (Hydrophorus) 

Hydrophorus criddlei Van Duzee 1925 Syn. 

Hydrophorus argentifacies Van Duzee 1926 Syn. 

Hydrophorus arnaudi Hurley 1985 (Hydrophorus) 

Hydrophorus californicus Hurley 1985 Syn. 

Hydrophorus bovatus Hurley 1985 (Hydrophorus) 

Hydrophorus brevicauda Van Duzee 1923 (Hydrophorus) 

Hydrophorus breviseta Thomson 1869 (Medeterus) 

Hydrophorus canescens Wheeler 1896 (Parhydrophorus) 

Hydrophorus canities Van Duzee 1923 (Hydrophorus) 

Hydrophorus chrysologus Walker 1849 (Medeterus) 

Hydrophorus alter Parent 1934 Syn. 

Hydrophorus claripennis Van Duzee 1924 (Hydrophorus) 

Hydrophorus nigrinervis Van Duzee 1926 Syn. 

Hydrophorus dioktes Hurley 1985 (Hydrophorus) 

Hydrophorus dreisbachi Harmston and Knowlton 1963 

(Hydrohorus) 

Hydrophorus eldoradensis Wheeler 1899 (Hydrophorus) 

Hydrophorus extrarius Aldrich 1911 (Hydrophorus) 

Hydrophorus purus Curran 1924 Syn. 


143

144 

Hydrophorus ferruginus Hurley 1985 (Hydrophorus) 

Hydrophorus flavihirtus Van Duzee 1923 (Hydrophorus) 

Hydrophorus flavipennis Van Duzee 1926 (Hydrophorus) 

Hydrophorus fumipennis Van Duzee 1921 (Hydrophorus) 

Hydrophorus glaber Walker 1849 (Medeterus) 

Hydrophorus propinquus Van Duzee 1923 Syn. 

Hydrophorus harmstoni Hurley 1985 (Hydrophorus) 

Hydrophorus hesperius Hurley 1985 (Hydrophorus) 

Hydrophorus hirpicifer Hurley 1985 (Hydrophorus) 

Hydrophorus innotatus Loew 1864 (Hydrophorus) 

Hydrophorus vandykei Van Duzee 1926 Syn. 

Hydrophorus maculipennis Van Duzee 1926 (Hydrophorus) 

Hydrophorus magdalenae Wheeler 1899 (Hydrophorus) 

Hydrophorus manicatus Collin 1935 (Hydrophorus) 

Hydrophorus minimus Van Duzee 1924 (Hydrophorus) 

Hydrophorus myllomerus Hurley 1985 (Hydrophorus) 

Hydrophorus parvus Loew 1862 (Hydrophorus) 

Hydrophorus pectinipes Van Duzee 1923 (Hydrophorus) 

Hydrophorus pensus Aldrich 1911 (Hydrophorus) 

Hydrophorus phaeopteryx Hurley 1985 (Hydrophorus) 

Hydrophorus phalarus Hurley 1985 (Hydrophorus) 

Hydrophorus philombrius Wheeler 1890 (Hydrophorus) 

Hydrophorus phoca Aldrich 1911 (Hydrophorus) 

Hydrophorus oregonensis Van Duzee 1933 Syn. 

Hydrophorus spinosus Van Duzee 1933 Syn. 

Hydrophorus pilitarsis Malloch 1919 (Hydrophorus) 

Hydrophorus plautus Hurley 1985 (Hydrophorus) 

Hydrophorus rhionopous Hurley 1985 (Hydrophorus) 

Hydrophorus signiferus Coquillett 1899 (Hydrophorus) 

Hydrophorus magnicornis Frey 1915 Syn. 

Hydrophorus nigribarbus Van Duzee 1923 Syn. 

Hydrophorus signifer Foote et al. 1965 Emend. 

Hydrophorus sodalis Wheeler 1899 (Hydrophorus) 

Hydrophorus trichaspis Hurley 1985 (Hydrophorus) 

Hydrophorus viridifacies Van Duzee 1923 (Hydrophorus) 

Hypocharassus Mik 1878 

Drepanomyia Wheeler 1898 Syn. 

Hypocharassus gladiator Mik 1878 (Hypocharassus) 

Drepanomyia johnsonii Wheeler 1898 Syn. 

Hypocharassus pruinosa Wheeler 1898 (Drepanomyia) 

Keirosoma Van Duzee 1929 

Keirosoma slossonae Van Duzee 1933 (Keirosoma) 

Lamprochromus Mik 1878 

Lamprochromus canadensis Van Duzee 1917 (Sympycnus) 

Telmaturgus brevicornis Robinson 1960 Syn. 

Lamprochromus occidentalis Robinson 1967 (Lamprochromus) 

Lamprochromus satrapa Wheeler 1890 (Diaphorus) 

Liancalus Loew 1857 

Anoplomerus Rondani 1856 Homo. 

Anoplopus Rondani 1857 Homo. 

Liancalus genualis Loew 1861 (Liancalus) 

Liancalus hydrophilus Aldrich 1893 (Liancalus) 

Liancalus limbatus Osten Sacken 1917 (Liancalus) 

Liancalus querulus Osten Sacken 1877 (Liancalus) 

Liancalus similis Aldrich 1893 (Liancalus) 

Medetera Fischer von Waldheim 1819 

Medeterium Berthold 1827 Syn. 

Taechobates Haliday 1832 Syn. 

Orthobates Wahlberg 1844 Syn. 



Anorthus Loew 1850 Syn. 

Oligochaetus Mik 1878 Syn. 

Asioligochaetus Negrobov 1966 Syn. 

Lorea Negrobov 1966 Syn. 

Medetera aberrans Wheeler 1899 (Medetera) 

Medeterus lobatus Van Duzee 1914 Syn. 

Medeterus flavicosta Van Duzee 1932 Syn. 

Medetera aeneiventris Van Duzee 1933 (Medetera) 

Medeterus aeneus Van Duzee 1919 Homo. 

Medeterus obesus Van Duzee 1919 Homo. 

Medeterus univittatus Van Duzee 1933 Syn. 

Medetera obsoletas Megrobov and Thuneberg 1970 Syn. 

Medetera aequalis Van Duzee 1919 (Medetera) 

Medetera aldrichii Wheeler 1899 (Medetera) 

Medetera oregonensis Van Duzee 1919 Syn. 

Medetera alpina Harmston and Knowlton 1941 (Medetera) 

Medetera apicalis Zetterstedt 1843 (Medetera) 

Medeterus aurivittata Wheeler 1899 Syn. 

Medeterus caerulescens Malloch 1919 Syn. 

Medeterus distinctus Van Duzee 1919 Syn. 

Medeterus frontalis Van Duzee 1919 Syn. 

Medeterus bicolor Van Duzee 1923 Homo. 

Medeterus parvus Van Duzee 1923 Syn. 

Medeterus ciliatus Van Duzee 1928 Syn. 

Medeterus simplicipes Curran 1928 Syn. 

Medeterus venatus Curran 1928 Syn. 

Medetera orbiculata Van Duzee 1932 Syn. 

Medetera albiciliata Van Duzee 1933 Syn. 

Medetera arctica Van Duzee 1933 Syn. 

Medetera arnaudi Harmston 1951 (Medetera) 

Medetera bistriata Parent 1929 (Medetera) 

Medetera californiensis Wheeler 1899 (Medetera) 

Medetera longimana Van Duzee 1933 Syn. 

Medetera canadensis Bickel 1985 (Medetera) 

Medetera crassivenis Curran 1928 (Medetera) 

Medetera idahoensis Harmston and Knowlton 1943 Syn. 

Medetera cyanogaster Wheeler 1899 (Medetera) 

Medetera diadema Linnaeus 1767 (Musca) 

Medetera princeps Wheeler 1899 Syn. 

Medetera dorycondylus Bickel 1985 (Medetera) 

Medetera excipiens Becker 1922 (Medetera) 

Medetera falcata Van Duzee 1919 (Medetera) 

Medeterus longinquus Van Duzee 1919 Syn. 

Medetera flinflon Bickel 1985 (Medetera) 

Medetera furcata Curran 1928 (Medetera) 

Medetera gaspensis Bickel 1985 (Medetera) 

Medetera halteralis Van Duzee 1919 (Medetera) 

Medetera tarasovae Negrobov 1974 Syn. 

Medetera isobellae Bickel 1985 (Medetera) 

Medetera longinervis Van Duzee 1928 (Medetera) 

Medetera marylandica Robinson 1967 (Medetera) 

Medetera maura Wheeler 1899 (Medetera) 

Medetera modesta Van Duzee 1914 (Medetera) 

Medetera neomelancholia Bickel 1985 (Medetera) 

Medetera nigripes Loew 1861 (Medetera) 

Medetera nova Van Duzee 1919 (Medetera) 

Medetera petulca Wheeler 1899 (Medetera) 

Medetera nitidiventris Van Duzee 1919 Syn. 

Medetera physothrix Bickel 1985 (Medetera) 

Medetera pinicola Kowarz 1877 (Medetera) 

Medetera platythrix Bickel 1985 (Medetera) 

Medetera postminima Steyskal 1967 (Medetera) 

Medetera minima Van Duzee 1925 Homo. 

Medetera potomac Bickel 1985 (Medetera) 

Medetera pseudosibirica Bickel 1985 (Medetera) 

Medetera saguaroicola Bickel 1985 (Medetera) 

Medetera signaticornis Loew 1857 (Medetera)

Medeterus viridifacies Van Duzee 1923 Syn. 

Medeterus trisetosus Van Duzee 1924 Syn. 

Medeterus vanduzeei Curran 1928 Syn. 

Medetera similis Van Duzee 1919 (Medetera) 

Medetera cuneiformis Van Duzee 1919 Syn. 

Medetera subsignaticornis Bickel 1985 (Medetera) 

Medetera truncorum Meigen 1824 (Medetera) 

Medetera tuktoyaktuk Bickel 1985 (Medetera) 

Medetera utahensis Bickel 1985 (Medetera) 

Medetera veles Loew 1861 (Medetera) 

Medeterus appendiculatus Wheeler 1899 Homo. 

Medetera bilineata Frey 1915 Syn. 

Medetera albosetosa Van Duzee 1928 Syn. 

Medetera intermedia Van Duzee 1928 Syn. 

Medetera wheeleri Foote, Coulson, and Robinson 1965 Syn. 

Medetera vidua Wheeler 1899 (Medetera) 

Medeterus emarginatus Van Duzee 1914 Syn. 

Medetera vittata Van Duzee 1919 (Medetera) 

Medetera vockerothi Bickel 1985 (Medetera) 

Medetera xerophila Wheeler 1899 (Medetera) 

Medetera obscuripennis Van Duzee 1919 Syn. 

Melanderia Aldrich 1922 

Wirthia Arnaud 1958 Syn. 

Melanderia californica Harmston 1972 (Melanderia) 

Melanderia crepuscula Arnaud 1958 (Melanderia) 

Melanderia curvipes Van Duzee 1918 (Hydrophorus) 

Melanderia mandibulata Aldrich 1922 (Melanderia) 

Mesorhaga Schiner 1868 

Aptorthus Aldrich 1893 Syn. 

Mesorhaga albiciliatus Aldrich 1893 (Aptorthus) 

Mesorhaga borealis Aldrich 1893 (Aptorthus) 

Mesorhaga caerulea Van Duzee 1930 (Mesorhaga) 

Mesorhaga caudata Van Duzee 1915 (Mesorhaga) 

Mesorhaga clavicauda Van Duzee 1925 (Mesorhaga) 

Mesorhaga flavipes Van Duzee 1932 (Mesorhaga) 

Mesorhaga jucunda Becker 1922 (Mesorhaga) 

Mesorhaga nigripes Aldrich 1893 (Aptorthus) 

Mesorhaga pallidicornis Van Duzee 1925 (Mesorhaga) 

Mesorhaga townsendi Aldrich 1893 (Aptorthus) 

Mesorhaga tricorniflavrai Gunther 1982 (Mesorhaga) 

Mesorhaga varipes Van Duzee 1917 (Mesorhaga) 

Micromorphus Mik 1878 

Micromorphus albipes Zetterstedt 1843 (Hydrophorus) 

Micromorphus bifrons Robinson 1964 (Micromorphus) 

Micromorphus fulvosetosus Parent 1929 (Micromorphus) 

Micromorphus knowltoni Robinson 1967 (Micromorphus) 

Micromorphus leucostoma Robinson 1967 (Micromorphus) 

Micromorphus longilamellatus Robinson 1964 (Micromorphus) 

Micromorphus minima Van Duzee 1925 (Neurigona) 

Nanomyina Robinson 1964 

Nanomyina litorea Robinson 1964 (Nanomyina) 

Nematoproctus Loew 1857 

Nematoproctus cylindricus Van Duzee 1924 (Diaphorus) 

Nematoproctus flavicoxa Van Duzee 1930 (Nematoproctus) 

Nematoproctus jucundus Van Duzee 1927 (Nematoproctus) 

Nematoproctus junctus Van Duzee 1930 Missp. 

Nematoproctus metallicus Van Duzee 1930 (Nematoproctus) 

Nematoproctus terminalis Van Duzee 1914 (Leucostola) 


Nematoproctus varicoxa Van Duzee 1930 (Nematoproctus) 

Nematoproctus venustus Melander 1900 (Nematoproctus) 

Neoparentia Robinson 1967 

Neoparentia caudatus Van Duzee 1917 (Sympycnus) 

Neurigona Rondani 1856 

Saucropus Loew 1857 Syn. 

Dactylomyia Aldrich 1894 Syn. 

Neurigona aestiva Van Duzee 1913 (Neurigona) 

Neurigona albospinosa Van Duzee 1913 (Neurigona) 

Neurigona aldrichii Van Duzee 1913 (Neurigona) 

Neurigona arcuata Van Duzee 1913 (Neurigona) 

Neurigona australis Van Duzee 1913 (Neurigona) 

Neurigona bivittata Van Duzee 1913 (Neurigona) 

Neurigona californica Harmston 1972 (Neurigona) 

Neurigona carbonifer Loew 1869 (Saucropus) 

Neurigona ciliata Van Duzee 1913 (Neurigona) 

Neurigona cilimanus Van Duzee 1924 (Neurigona) 

Neurigona deformis Van Duzee 1913 (Neurigona) 

Neurigona dimidiatus Loew 1861 (Saucropus) 

Neurigona disjuncta Van Duzee 1913 (Neurigona) 

Neurigona flava Van Duzee 1913 (Neurigona) 

Neurigona uinta Harmston and Knowlton 1942 Syn. 

Neurigona floridula Wheeler 1899 (Neurigona) 

Neurigona infuscata Van Duzee 1913 Syn. 

Neurigona fuscalaris Harmston 1968 (Neurigona) 

Neurigona georgianus Harmston and Knowlton 1963 (Neurigona) 

Neurigona kesseli Hendrickson 1961 (Neurigona) 

Neurigona lateralis Say 1829 (Medeterus) 

Saucropus superbiens Loew 1861 Syn. 

Dactylomyia gracilipes Aldrich 1894 Syn. 

Neurigona lienosa Wheeler 1899 (Neurigona) 

Neurigona maculata Van Duzee 1913 (Neurigona) 

Neurigona nigrimanus Van Duzee 1930 (Neurigona) 

Neurigona nigritibalis Robinson 1964 (Neurigona) 

Neurigona nitida Van Duzee 1913 (Neurigona) 

Neurigona ornata Van Duzee 1930 (Neurigona) 

Neurigona pectoralis Van Duzee 1913 (Neurigona) 

Neurigona perbrevis Van Duzee 1913 (Neurigona) 

Neurigona perplexa Van Duzee 1913 (Neurigona) 

Neurigona planipes Van Duzee 1924 (Neurigona) 

Neurigona rubellus Loew 1861 (Saucropus) 

Neurigona scutitarsis Robinson 1970 (Neurigona) 

Neurigona smithi Robinson 1970 (Neurigona) 

Neurigona spiculifera Robinson 1970 (Neurigona) 

Neurigona tarsalis Van Duzee 1913 (Neurigona) 

Neurigona tenuis Loew 1864 (Saucropus) 

Neurigona terminalis Van Duzee 1924 (Neurigona) 

Neurigona torrida Harmston 1951 (Neurigona) 

Neurigona transversa Van Duzee 1913 (Neurigona) 

Neurigona tridens Van Duzee 1913 (Neurigona) 

Neurigona valgusa Harmston and Rapp 1968 (Neurigona) 

Neurigona viridis Van Duzee 1913 (Neurigona) 

Neurigona zionensis Harmston and Knowlton 1942 (Neurigona) 

Neurigonella Robinson 1964 

Neurigonella nigricornis Van Duzee 1914 (Neurigona) 

Neurigonella sombrea Harmston and Knowlton 1945 (Neurigona) 

Orthochile of authors 

Orthochile of authors deremptus Walker 1849 (Orthochile) 

Paraclius Loew 1864 


145

146 



Paracleius Bigot 1859 Unav. 

Leptocorypha Aldrich 1896 Syn. 

The generic name Paraclius is technically an emendation of 

Paracleius. This useage, in line with current authors, preserves the 

name Paraclius even though Paracleius Bigot Loew 1864 is an 

available name and perhaps not congeneric with the species currently 

standing under the name Paraclius. This case needs to be resolved by 

workers in the family. 

Paraclius aberrans Robinson 1964 (Paraclius) 

Paraclius alternans Loew 1864 (Pelastoneurus) 

Paracleius vicinus Aldrich 1904 Syn. 

Paraclius claviculatus Loew 1866 (Paracleius) 

Paracleius fraternus Van Duzee 1933 Syn. 

Paraclius consors Walker 1852 (Dolichopus) 

Paraclius filifer Aldrich 1896 (Paracleius) 

Paraclius flagellatus Harmston 1952 (Hercostomus) 

Paraclius floridensis Robinson 1964 (Paraclius) 

Paraclius heteroneurus Macquart 1850 (Dolichopus) 

Paraclius hybridus Melander 1900 (Paracleius) 

Paraclius magnicornis Van Duzee 1927 (Paracleius) 

Paraclius minutus Van Duzee 1921 (Paracleius) 

Paraclius nigrocaudatus Van Duzee 1918 (Paracleius) 

Paraclius ovatus Van Duzee 1914 (Paracleius) 

Paraclius propinquus Wheeler 1899 (Paracleius) 

Paraclius pumilio Loew 1872 (Paracleius) 

Paraclius quadrinotatus Aldrich 1902 (Paracleius) 

Paraclius utahensis Harmston and Knowlton 1946 (Paracleius) 

Parasyntormon Wheeler 1899 

Neosyntormon Curran 1934 Syn. 

Parasyntormon appendiculatum Harmston and Knowlton 1943 

(Parasyntormon) 

Parasyntormon asellus Wheeler 1899 (Parasyntormon) 

Parasyntormon classicum Harmston and Knowlton 1943 


Parasyntormon emarginatum Wheeler 1899 (Parasyntormon) 

Parasyntormon emarginicorne Curran 1923 (Parasyntormon) 

Parasyntormon flavicoxa Van Duzee 1922 (Parasyntormon) 

Parasyntormon fraterculus Van Duzee 1922 (Parasyntormon) 

Parasyntormon hendersoni Harmston and Knowlton 1939 


Parasyntormon hinnulus Wheeler 1899 (Parasyntormon) 

Parasyntormon lagotis Wheeler 1899 (Parasyntormon) 

Parasyntormon lepus Van Duzee 1918 (Parasyntormon) 

Parasyntormon longicorne Van Duzee 1933 (Parasyntormon) 

Parasyntormon montivagum Wheeler 1899 (Parasyntormon) 

Parasyntormon mulinum Van Duzee 1922 (Parasyntormon) 

Parasyntormon nigripes Harmston and Knowlton 1943 


Parasyntormon occidentalis Aldrich 1894 (Sympycnus) 

Parasyntormon petiolatum Van Duzee 1933 (Parasyntormon) 

Parasyntormon rotundicorne Van Duzee 1926 (Parasyntormon) 

Parasyntormon utahnum Van Duzee 1933 (Parasyntormon) 

Parasyntormon virens Harmston and Knowlton 1943 


Pelastoneurus Loew 1861 

Metapelastoneurus Aldrich 1894 Syn. 

Pelastoneurus abbreviatus Loew 1864 (Pelastoneurus) 

Pelastoneurus aldrichi Van Duzee 1923 (Pelastoneurus) 

Pelastoneurus angulatus Robinson 1964 (Pelastoneurus) 

Pelastoneurus asciaeformis Becker 1922 (Pelastoneurus) 

Pelastoneurus aurifacies Van Duzee 1923 (Pelastoneurus) 

Pelastoneurus minutus Van Duzee 1933 Syn. 

Pelastoneurus barri Harmston 1972 (Pelastoneurus) 

Pelastoneurus bifrons Walker 1852 (Dolichopus) 

Pelastoneurus brevis Robinson 1964 (Pelastoneurus) 

Pelastoneurus cognatus Loew 1861 (Pelastoneurus) 

Pelastoneurus comatus Robinson 1992 (Pelastoneurus) 

Pelastoneurus cyaneus Wheeler 1899 (Pelastoneurus) 

Pelastoneurus dissimilipes Wheeler 1899 (Pelastoneurus) 

Pelastoneurus nigrescens Wheeler 1899 Syn. 

Pelastoneurus floridanus Wheeler 1899 (Pelastoneurus) 

Pelastoneurus furcifer Loew 1872 (Pelastoneurus) 

Pelastoneurus hebes Walker 1852 (Dolichopus) 

Pelastoneurus ineptus Walker 1852 (Dolichopus) 

Pelastoneurus irrasus Walker 1849 (Dolichopus) 

Pelastoneurus kansensis Aldrich 1894 (Metapelastoneurus) 

Pelastoneurus laetus Loew 1861 (Pelastoneurus) 

Pelastoneurus falcatus Aldrich 1904 Syn. 

Pelastoneurus ramosus Van Duzee 1923 Syn. 

Pelastoneurus lamellatus Loew 1864 (Pelastoneurus) 

Pelastoneurus cristatus Van Duzee 1924 Syn. 

Pelastoneurus latifacies Van Duzee 1930 (Pelastoneurus) 

Pelastoneurus longicauda Loew 1861 (Pelastoneurus) 

Pelastoneurus quadricincta Van Duzee 1928 Syn. 

Pelastoneurus lugubris Loew 1861 (Pelastoneurus) 

Pelastoneurus maculipes Walker 1852 (Dolichopus) 

Pelastoneurus neglectus Wheeler 1899 (Pelastoneurus) 

Pelastoneurus nigricornis Van Duzee 1923 (Pelastoneurus) 

Pelastoneurus penicillatus Parent 1929 Syn. 

Pelastoneurus occidentalis Wheeler 1899 (Pelastoneurus) 

Pelastoneurus parvus Aldrich 1904 (Pelastoneurus) 

Pelastoneurus potomacus Robinson 1992 (Pelastoneurus) 

Pelastoneurus proximus Aldrich 1904 (Pelastoneurus) 

Pelastoneurus arboreus Van Duzee 1923 Syn. 

Pelastoneurus scutatus Aldrich 1904 (Pelastoneurus) 

Pelastoneurus semiplumatus Becker 1922 (Pelastoneurus) 

Pelastoneurus seticauda Van Duzee 1930 (Pelastoneurus) 

Pelastoneurus stentorius Harmston 1971 (Pelastoneurus) 

Pelastoneurus tibialis Van Duzee 1923 (Pelastoneurus) 

Pelastoneurus umbripictus Becker 1922 (Pelastoneurus) 

Pelastoneurus vagans Loew 1861 (Pelastoneurus) 

Pelastoneurus longilamellatus Parent 1929 Syn. 

Pelastoneurus varius Walker 1852 (Dolichopus) 

Pelastoneurus pictipennis Wheeler 1899 Syn. 

Pelastoneurus wheeleri Melander 1900 (Pelastoneurus) 

Peloropeodes Wheeler 1890 

Anomalopyga Oldenberg 1916 Syn. 

Kophosoma Van Duzee 1926 Syn. 

Pachypyga Parent 1928 Syn. 

Peloropeodes acuticornis Van Duzee 1926 (Kophosoma) 

Peloropeodes apicales Harmston and Knowlton 1946 

(Peloropeodes) 

Peloropeodes bicolor Van Duzee 1926 (Kophosoma) 

Peloropeodes brevis Van Duzee 1926 (Kophosoma) 

Campsicnemus crassitibia Van Duzee 1930 Syn. 

Peloropeodes cornuta Van Duzee 1926 (Kophosoma) 

Peloropeodes discolor Robinson 1964 (Peloropeodes) 

Peloropeodes frater Robinson 1960 Homo. 

Peloropeodes fuscipes Van Duzee 1926 (Kophosoma) 

Peloropeodes magnicornis Harmston and Rapp 1968 (Peloropeodes) 

Peloropeodes pygidus Harmston and Rapp 1968 (Peloropeodes) 

Peloropeodes salax Wheeler 1890 (Peloropeodes) 

Peloropeodes flavipes Van Duzee 1914 Syn. 

Plagioneurus Loew 1857 

Plagioneurus univittatus Loew 1857 (Plagioneurus) 

Psilopiella Van Duzee 1914

Psilopiella nebraskaense Harmston and Rapp 1968 (Psilopiella) 

Psilopiella rutila Van Duzee 1914 (Psilopiella) 

Rhaphium Meigen 1803 

Anglearia Carlier 1835 Syn. 

Xiphandrium Loew 1857 Syn. 

Rhaphium aequale Van Duzee 1927 (Rhaphium) 

Rhaphium albibarba Van Duzee 1924 (Porphyrops) 

Rhaphium aldrichi Van Duzee 1922 (Xiphandrium) 

Rhaphium arboreum Curran 1924 (Rhaphium) 

Rhaphium armatum Curran 1924 (Rhaphium) 

Rhaphium atkinsoni Curran 1926 (Rhaphium) 

Rhaphium banksi Van Duzee 1926 (Rhaphium) 

Rhaphium barbipes Van Duzee 1923 (Porphyrops) 

Rhaphium borealis Van Duzee 1923 (Porphyrops) 

Rhaphium brevicoris Van Duzee 1923 (Porphyrops) 

Rhaphium brevilamellatum Van Duzee 1926 (Rhaphium) 

Rhaphium browni Curran 1931 (Rhaphium) 

Rhaphium calcaratum Van Duzee 1928 (Rhaphium) 

Rhaphium campestris Curran 1924 (Rhaphium) 

Rhaphium canadense Curran 1924 (Rhaphium) 

Rhaphium caudatum Van Duzee 1926 (Rhaphium) 

Rhaphium ciliatum Curran 1929 (Rhaphium) 

Rhaphium coloradense Curran 1926 (Rhaphium) 

Xiphandrium longicorne Van Duzee 1922 Homo. 

Rhaphium colutis Harmston and James 1942 (Rhaphium) 

Rhaphium crassipes Meigen 1824 (Porphyrops) 

Rhaphium discolor Zetterstedt 1838 (Rhaphium) 

Rhaphium consobrinum Zetterstedt 1843 Syn. 

Rhaphium dubium Van Duzee 1922 (Xiphandrium) 

Rhaphium effilatus Wheeler 1899 (Porphyrops) 

Rhaphium elegantulus Meigen 1824 (Porphyrops) 

Rhaphium elongatum Van Duzee 1933 (Rhaphium) 

Rhaphium exile Curran 1926 (Rhaphium) 

Rhaphium fascipes Meigen 1824 (Porphyrops) 

Rhaphium femineum Van Duzee 1922 (Xiphandrium) 

Rhaphium femoratum Van Duzee 1922 (Xiphandrium) 

Rhaphium flavicoxa Van Duzee 1922 (Xiphandrium) 

Rhaphium foliatum Curran 1926 (Rhaphium) 

Rhaphium fuscicosta Van Duzee 1928 Syn. 

Rhaphium furcifer Curran 1931 (Rhaphium) 

Rhaphium gibsoni Curran 1926 (Rhaphium) 

Rhaphium gracilis Curran 1924 (Rhaphium) 

Rhaphium grandis Curran 1923 (Porphyrops) 

Rhaphium hirtimanus Van Duzee 1933 (Rhaphium) 

Rhaphium impetuum Curran 1926 (Rhaphium) 

Rhaphium insolitum Curran 1926 (Rhaphium) 

Rhaphium johnsoni Van Duzee 1923 (Porphyrops) 

Rhaphium latifacies Van Duzee 1930 (Rhaphium) 

Rhaphium longibara Van Duzee 1930 (Rhaphium) 

Rhaphium longipalpis Curran 1926 (Rhaphium) 

Rhaphium longipes Loew 1864 (Porphyrops) 

Rhaphium lugubre Loew 1861 (Rhaphium) 

Rhaphium melampus Loew 1861 (Porphyrops) 

Systenus americanus Van Duzee 1914 Syn. 

Rhaphium montanus Van Duzee 1920 (Porphyrops) 

Rhaphium nigra Van Duzee 1923 (Porphyrops) 

Rhaphium nigricoxa Loew 1861 (Porphyrops) 

Rhaphium nigrociliatum Curran 1926 (Rhaphium) 

Rhaphium nigrovittatum Curran 1926 (Rhaphium) 

Rhaphium nudus Van Duzee 1924 (Porphyrops) 

Rhaphium obtusum Van Duzee 1928 (Rhaphium) 

Rhaphium occipitale Curran 1927 (Rhaphium) 

Rhaphium orientale Curran 1926 (Rhaphium) 

Rhaphium ornatus Van Duzee 1923 (Porphyrops) 

Rhaphium petchi Curran 1924 Syn. 


Rhaphium palpale Curran 1926 (Rhaphium) 

Rhaphium pollex Van Duzee 1922 (Xiphandrium) 

Rhaphium punctitarsis Curran 1924 (Rhaphium) 

Rhaphium robustum Curran 1926 (Rhaphium) 

Rhaphium rossi Harmston and Knowlton 1940 (Rhaphium) 

Rhaphium rotundiceps Loew 1861 (Porphyrops) 

Porphyrops fumipennis Loew 1861 Syn. 

Rhaphium septentrionale Curran 1931 (Rhaphium) 

Rhaphium shannoni Curran 1926 (Rhaphium) 

Rhaphium signiferus Osten Sacken 1878 (Porphyrops) 

Rhaphium simplicipes Curran 1926 (Rhaphium) 

Rhaphium slossonae Johnson 1906 (Leucostola) 

Rhaphium spinitarsis Curran 1924 (Rhaphium) 

Rhaphium steyskali Robinson 1967 (Rhaphium) 

Rhaphium subarmatum Curran 1924 (Rhaphium) 

Rhaphium subfurcatum Van Duzee 1932 (Rhaphium) 

Rhaphium temerarium Becker 1922 (Xiphandrium) 

Rhaphium terminalis Van Duzee 1924 (Porphyrops) 

Rhaphium triangulatum Van Duzee 1922 (Xiphandrium) 

Rhaphium tricaudata Van Duzee 1923 (Porphyrops) 

Rhaphium tripartita Frey 1913 (Porphyrops) 

Rhaphium unistylatum Vockeroth 1952 (Rhaphium) 

Rhaphium vanduzeei Curran 1926 (Rhaphium) 

Rhaphium vockerothi Robinson 1964 (Rhaphium) 

Rhaphium wheeleri Van Duzee 1932 (Rhaphium) 

Rhaphium xipheres Wheeler 1899 (Porphyrops) 

Scellus Loew 1857 

Scellus abditus Hurley 1995 (Scellus) 

Scellus amplus Curran 1923 (Scellus) 

Scellus asaroticus Hurley 1995 (Scellus) 

Scellus avidus Loew 1864 (Scellus) 

Scellus coloradensis Harmston and James 1942 (Scellus) 

Scellus crinipes Van Duzee 1925 (Scellus) 

Scellus dyscritus Hurley 1995 (Scellus) 

Scellus exustus Walker 1852 (Medeterus) 

Scellus filifer Loew 1864 (Scellus) 

Scellus knowltoni Harmston 1939 (Scellus) 

Scellus monstrosus Osten Sacken 1877 (Scellus) 

Scellus spinimanus Zetterstedt 1843 (Hydrophorus) 

Scellus varipennis Van Duzee 1925 (Scellus) 

Scellus vigil Osten Sacken 1877 (Scellus) 

Scellus virago Aldrich 1907 (Scellus) 

Sciapus Zeller 1842 

Psilopus Meigen 1824 Homo. 

Stenarus Gistel 1848 Syn. 

Psilopodius Rondani 1861 Syn. 

Psilopodinus Bigot 1888 Syn. 

Sciapus bicolor Loew 1861 (Psilopus) 

Sciapus bradleii Van Duzee 1915 (Sciapus) 

Sciapus californicus Steyskal 1966 (Sciapus) 

Sciapus costalis Aldrich 1904 (Agonosoma) 

Sciapus delicatus Walker 1849 (Psilopus) 

Psilopodinus pallescens Bigot 1888 Syn. 

Sciapus noveboracensis Van Duzee 1915 Syn. 

Sciapus divergens Van Duzee 1933 (Sciapus) 

Sciapus dorsalis Loew 1866 (Psilopus) 

Condylostylus viridivittatus Robinson 1960 Syn. 

Sciapus filipes Loew 1861 (Psilopus) 

Sciapus floridanus Harmston 1971 (Sciapus) 

Sciapus fuscinervis Van Duzee 1926 (Psilopus) 

Sciapus infumatus Aldrich 1901 (Gnamptopsilopus) 

Sciapus pallens Wiedemann 1830 (Psilopus) 

Sciapus pollinosus Van Duzee 1915 (Sciapus) 

Sciapus pressipes Parent 1929 (Sciapus) 


147

148 

Sciapus psittacinus Loew 1861 (Psilopus) 

Sciapus rotundiceps Aldrich 1904 (Agonosoma) 

Sciapus scintillans Loew 1861 (Psilopus) 

Sciapus fulgidus Parent 1929 Syn. 

Sciapus dubiosus Van Duzee 1932 Syn. 

Sciapus tener Loew 1862 (Psilopus) 

Sciapus trisetosus Van Duzee 1932 (Sciapus) 

Sciapus unicoiensis Robinson 1964 (Sciapus) 

Sciapus unifasciatus Say 1823 (Dolichopus) 

Psilopus sayi Wiedemann 1830 Syn. 

Psilopus ungulivena Walker 1857 Syn. 

Sciapus amabilis Parent 1929 Syn. 

Chrysosoma plumosa Van Duzee 1932 Syn. 

Sciapus variegatus Loew 1861 (Psilopus) 

Sympycnidelphus Robinson 1964 



Sympycnidelphus californicus Harmston 1968 (Sympycnidelphus) 

Sympycnidelphus sharpi Robinson 1964 (Sympycnidelphus) 

Sympycnidelphus texanus Harmston 1968 (Sympycnidelphus) 

Sympycnus Loew 1857 

Gymnoceromyia Bigot 1890 Syn. 

Subsympycnus Becker 1922 Syn. 

Pycsymnus Frey 1925 Syn. 

Hopliognusus Vaillant 1953 Syn. 

Sympycnus aldrichi Van Duzee 1930 (Sympycnus) 

Sympycnus arizonicus Harmston 1968 (Sympycnus) 

Sympycnus aurifacies Van Duzee 1923 (Sympycnus) 

Sympycnus basistylatus Parent 1929 (Sympycnus) 

Sympycnus binodatus Harmston and Knowlton 1940 (Sympycnus) 

Sympycnus brevicauda Van Duzee 1932 (Sympycnus) 

Sympycnus brevipes Van Duzee 1933 (Sympycnus) 

Sympycnus breviventris Van Duzee 1930 (Sympycnus) 

Sympycnus calcaratus Van Duzee 1930 (Sympycnus) 

Sympycnus clavatus Van Duzee 1913 (Sympycnus) 

Sympycnus cuprinus Wheeler 1899 (Sympycnus) 

Sympycnus fasciventris Van Duzee 1917 (Sympycnus) 

Sympycnus globulicauda Van Duzee 1930 (Sympycnus) 

Sympycnus hardyi Harmston and Knowlton 1940 (Sympycnus) 

Sympycnus inaequalis Van Duzee 1930 (Sympycnus) 

Sympycnus isoaristus Harmston and Knowlton 1940 (Sympycnus) 

Sympycnus laevigatus Van Duzee 1930 (Sympycnus) 

Sympycnus latitarsis Van Duzee 1930 (Sympycnus) 

Peloropeodes jamesi Harmston and Knowlton 1939 Syn. 

Sympycnus lineatus Loew 1861 (Sympycnus) 

Sympycnus longinervis Van Duzee 1932 (Sympycnus) 

Sympycnus marcidus Wheeler 1899 (Sympycnus) 

Sympycnus minuticornis Van Duzee 1932 (Sympycnus) 

Sympycnus montanus Van Duzee 1930 (Sympycnus) 

Sympycnus pectoralis Van Duzee 1933 (Sympycnus) 

Sympycnus pennarista Harmston and Knowlton 1940 (Sympycnus) 

Sympycnus pictipes Harmston and Knowlton 1940 (Sympycnus) 

Sympycnus pugil Wheeler 1899 (Sympycnus) 

Sympycnus pulvillus Van Duzee 1930 (Sympycnus) 

Sympycnus setosus Van Duzee 1930 (Sympycnus) 

Sympycnus tertianus Loew 1864 (Sympycnus) 

Sympycnus tripilus Van Duzee 1930 (Sympycnus) 

Sympycnus utahensis Harmston and Knowlton 1939 (Sympycnus) 

Syntormon Loew 1857 

Synarthrus Loew 1857 Syn. 

Eutarsus Loew 1857 Syn. 

Drymonoeca Becker 1907 Syn. 

Syntormon affinis Wheeler 1899 (Synarthrus) 

Syntormon bisinuatum Van Duzee 1925 (Syntormon) 

Syntormon californicum Harmston 1951 (Syntormon) 

Syntormon cinereiventris Loew 1861 (Synarthrus) 

Syntormon clavatum Van Duzee 1925 (Syntormon) 

Syntormon dissimilipes Van Duzee 1925 (Syntormon) 

Syntormon femoratum Van Duzee 1925 (Syntormon) 

Syntormon kennedyi Harmston and Knowlton 1942 (Syntormon) 

Syntormon myklebusti Harmston and Miller 1966 (Syntormon) 

Syntormon nubilum Van Duzee 1933 (Syntormon) 

Syntormon oregonensis Harmston and Knowlton 1942 (Syntormon) 

Syntormon ornatipes Van Duzee 1925 (Syntormon) 

Syntormon palmaris Loew 1864 (Synarthrus) 

Syntormon simplicitarse Van Duzee 1925 (Syntormon) 

Syntormon strataegus Wheeler 1899 (Synarthrus) 

Syntormon tricoloripes Curran 1923 (Syntormon) 

Syntormon uintaensis Harmston and Knowlton 1940 (Syntormon) 

Syntormon utahensis Harmston and Knowlton 1942 (Syntormon) 

Syntormon vanduzeei Curran 1931 (Syntormon) 

Syntormon variegatum Harmston 1952 (Syntormon) 

Systenus Loew 1857 

Systenus albimanus Wirth 1952 (Systenus) 

Systenus apicalis Wirth 1952 (Systenus) 

Systenus californicus Harmston 1968 (Systenus) 

Systenus eucercus Steyskal 1970 (Systenus) 

Systenus minuta Van Duzee 1913 (Neurigona) 

Systenus shannoni Wirth 1952 (Systenus) 

Tachytrechus Haliday 1851 

Ammobates Stannius 1831 Homo. 

Gongrophora Philippi 1875 Syn. 

Polymedon Osten Sacken 1877 Syn. 

Macellocerus Mik 1878 Syn. 

Psilischium Becker 1922 Syn. 

Tachytrechus albonotatus Loew 1864 (Paraclius) 

Tachytrechus angulatus Van Duzee 1914 (Paraclius) 

Tachytrechus angustipennis Loew 1862 (Tachytrechus) 

Tachytrechus argyropus Becker 1922 (Tachytrechus) 

Tachytrechus auratus Aldrich 1896 (Macellocerus) 

Tachytrechus binodatus Loew 1866 (Tachytrechus) 

Tachytrechus boharti Harmston 1968 (Tachytrechus)

Tachytrechus californicus Harmston and Knowlton 1943 

(Polymedon) 

Tachytrechus californicus Harmston and Knowlton 1963 

(Tachytrechus) 

Tachytrechus castus Wheeler 1899 (Polymedon) 

Tachytrechus dilaticosta Van Duzee 1927 (Polymedon) 

Tachytrechus duplicatus Harmston 1972 (Tachytrechus) 

Tachytrechus flabellifer Osten Sacken 1877 (Polymedon) 

Tachytrechus flavitibialis Van Duzee 1930 (Polymedon) 

Tachytrechus floridensis Aldrich 1896 (Tachytrechus) 

Tachytrechus granditarsis Greene 1922 (Tachytrechus) 

Tachytrechus greeni Foote, Coulson, and Robinson 1965 


Tachytrechus bipunctatus Greene 1922 Homo. 

Tachytrechus indianus Harmston and Knowlton 1946 (Paraclius) 

Tachytrechus laticrus Van Duzee 1918 (Tachytrechus) 

Tachytrechus mchughi Harmston 1972 (Tachytrechus) 

Tachytrechus milleri Harmston 1966 (Tachytrechus) 

Tachytrechus moechus Loew 1861 (Tachytrechus) 

Tachytrechus mysticus Becker 1922 (Paraclius) 

Tachytrechus nigrifemoratus Van Duzee 1927 (Polymedon) 

Tachytrechus nimius Aldrich 1901 (Polymedon) 

Tachytrechus nitidus Van Duzee 1927 (Polymedon) 

Tachytrechus olympiae Aldrich 1896 (Macellocerus) 

Tachytrechus protervus Melander 1900 (Tachytrechus) 

Tachytrechus junctus Coquillett 1910 Syn. 

Tachytrechus rotundipennis Greene 1922 (Tachytrechus) 

Tachytrechus sanus Osten Sacken 1877 (Tachytrechus) 

Tachytrechus simulatus Greene 1922 (Tachytrechus) 

Tachytrechus spinitarsis Van Duzee 1924 (Tetrechus) 

Tachytrechus tahoensis Harmston and Knowlton 1940 


Tachytrechus tenuiseta Greene 1922 (Tachytrechus) 

Tachytrechus utahensis Harmston and Knowlton 1940 


Tachytrechus volitans Melander 1900 (Tachytrechus) 

Tachytrechus vorax Loew 1861 (Tachytrechus) 

Telmaturgus Mik 1874 

Telmaturgus parvus Van Duzee 1924 (Chrysotus) 

Telmaturgus perparvus Johannsen 1928 Missp. 

Teuchophorus Loew 1857 

Teuchophorus clavigerellus Wheeler 1899 (Teuchophorus) 

Teuchophorus condylus Harmston and Knowlton 1946 

(Teuchophorus) 

Teuchophorus diminucosta Harmston and Knowlton 1942 


Teuchophorus utahensis Harmston and Knowlton 1942 


Thinophilus Wahlberg 1844 

Schoenophilus Mik 1878 Syn. 

Thinophilus armiger Van Duzee 1926 (Thinophilus) 

Thinophilus bimaculatus Johnson 1921 (Thinophilus) 

Thinophilus brevipes Van Duzee 1932 (Thinophilus) 

Thinophilus canities Van Duzee 1926 (Thinophilus) 

Thinophilus cantities Van Duzee 1926 Missp. 

Thinophilus delicatus Van Duzee 1926 (Thinophilus) 

Thinophilus depressus Van Duzee 1926 (Thinophilus) 

Thinophilus flavicaudatus Robinson 1964 (Thinophilus) 

Thinophilus frontalis Van Duzee 1914 (Thinophilus) 

Thinophilus insulanus Van Duzee 1926 (Thinophilus) 

Thinophilus latimanus Van Duzee 1926 (Thinophilus) 

Thinophilus magnipalpus Van Duzee 1926 (Thinophilus) 


Thinophilus neglectus Wheeler 1899 (Thinophilus) 

Thinophilus ochrifacies Van Duzee 1924 (Thinophilus) 

Thinophilus pectinifer Wheeler 1896 (Thinophilus) 

Thinophilus prasinus Johnson 1921 (Thinophilus) 

Thinophilus pruinosus Van Duzee 1930 (Thinophilus) 

Thinophilus rufibarbis Van Duzee 1926 (Thinophilus) 

Thinophilus russelli Curran 1927 (Thinophilus) 

Thinophilus scopiventris Harmston and Knowlton 1940 

(Thinophilus) 

Thinophilus semipallidus Robinson 1964 (Thinophilus) 

Thinophilus spinipes Van Duzee 1926 (Thinophilus) 

Thinophilus thallasinus Van Duzee 1926 (Thinophilus) 

Thinophilus vespertinus Robinson 1964 (Thinophilus) 

Thinophilus vinculatus Harmston and Rapp 1968 (Thinophilus) 

Thinophilus viridifacies Van Duzee 1924 (Thinophilus) 

Chrysotus quadratus Van Duzee 1924 Syn. 

Thinophilus nigripilosus Van Duzee 1926 Syn. 

Thrypticus Gerstaecker 1864 

Aphantotimus Wheeler 1890 Syn. 

Xanthotrichia Aldrich 1896 Syn. 

Submedeterus Becker 1917 Syn. 

DIPTERA: DOLICHOPODIDAE, DROSOPHILIDAE 

149 

Thrypticus abdominalis Say 1829 (Chrysotus) 

Thrypticus aurinotatus Van Duzee 1915 (Thrypticus) 

Thrypticus comosus Van Duzee 1915 (Thrypticus) 

Thrypticus deficiens Robinson 1980 (Thrypticus) 

Thrypticus dissectus Robinson 1964 (Thrypticus) 

Thrypticus fosteri Robinson 1980 (Thrypticus) 

Thrypticus fraterculus Wheeler 1890 (Aphantotimus) 

Thrypticus grogani Robinson 1980 (Thrypticus) 

Thrypticus longicauda Van Duzee 1921 (Thrypticus) 

Thrypticus minutus Parent 1929 (Thrypticus) 

Thrypticus muhlenbergiae Johannsen and Crosby 1913 (Thrypticus) 

Thrypticus nigripes Van Duzee 1921 (Thrypticus) 

Thrypticus setosus Robinson 1964 (Thrypticus) 

Thrypticus squamiciliatus Harmston and Rapp 1968 (Thrypticus) 

Thrypticus tectus Van Duzee 1915 (Thrypticus) 

Thrypticus vietus Van Duzee 1915 (Thrypticus) 

Thrypticus willistoni Wheeler 1890 (Aphantotimus) 

Xanthochlorus Loew 1857 

Xanthochlorus helvinus Loew 1861 (Xanthochlorus) 

DROSOPHILIDAE 

Amiota Loew 1862 

Phortica Schiner 1862 Syn. 

Erima Kertesz 1899 Syn. 

Sinophthalmus Coquillett 1904 Syn. 

Paraphortica Duda 1934 Syn. 

Apsiphortica Okada 1971 Syn. 

Amiota buccata Wheeler 1952 (Amiota) 

Amiota huachucae Wheeler 1960 (Amiota) 

Amiota humeralis Loew 1862 (Amiota) 

Amiota leucostoma Loew 1862 (Amiota) 

Amiota minor Malloch 1921 (Phortica) 

Amiota nigrescens Wheeler 1952 (Amiota) 

Amiota picta Coquillett 1904 (Sinophthalmus)

150 

Amiota setigera Malloch 1924 (Amiota) 

Amiota steganoptera Malloch 1926 (Amiota) 

Cacoxenus Loew 1858 

Gitonides Knab 1914 Syn. 

Eudrosophila Malloch 1924 Syn. 

Paracacoxenus Hardy 1960 Syn. 

Cacoxenus guttatus Hardy and Wheeler 1960 (Paracacoxenus) 

Chymomyza Czerny 1903 

Amphoroneura Meijere 1911 Syn. 

Zygodrosophila Hendel 1917 Syn. 

Chymomyza aldrichii Sturtevant 1916 (Chymomyza) 

Chymomyza tetonensis Wheeler 1949 Syn. 

Chymomyza amoena Loew 1862 (Drosophila) 

[none] subfasciata Harris 1835 Unav. 

Chymomyza caudatula Oldenberg 1914 (Chymomyza) 

Chymomyza coxata Wheeler 1952 (Chymomyza) 

Chymomyza olympia Wheeler 1960 (Chymomyza) 

Chymomyza procnemis Williston 1896 (Drosophila) 

Chymomyza procnemoides Wheeler 1952 (Chymomyza) 

Chymomyza wirthi Wheeler 1954 (Chymomyza) 

Cladochaeta Coquillett 1900 

Clastopteromyia Malloch 1924 Syn. 

Cladochaeta floridana Malloch 1924 (Clastopteromyia) 

Cladochaeta inversa Walker 1861 (Drosophila) 

Cladochaeta nebulosa Coquillett 1900 (Cladochaeta) 

Cladochaeta sturtevanti Wheeler and Takada 1971 (Cladochaeta) 

Dettopsomyia Lamb 1914 

Pictostyloptera Duda 1924 Syn. 

Dettopsomyia nigrovittata Malloch 1924 (Drosophila) 

Diathoneura Duda 1924 

Calatila Strand 1927 Syn. 

Macroptera Duda 1927 Homo. 

Diathoneura opaca Williston 1896 (Drosophila) 

Drosophila Fallen 1823 

Hypenomyia Grimshaw 1901 Syn. 

Idiomyia Grimshaw 1901 Syn. 

Chaetodrosophilella Duda 1923 Syn. 

Spinulophila Duda 1923 Syn. 

Hirtodrosophila Duda 1923 Syn. 

Scaptodrosophila Duda 1923 Syn. 

Paradrosophila Duda 1923 Syn. 

Spuriostyloptera Duda 1923 Syn. 

Acrodrosophila Duda 1924 Syn. 

Chaetodrosophila Duda 1924 Syn. 

Pugiodrosophila Duda 1924 Syn. 

Tanygastrella Duda 1924 Syn. 

Spinodrosophila Duda 1924 Syn. 

Acanthophila Duda 1925 Homo. 

Dasydrosophila Duda 1925 Syn. 

Xiphidiochaeta Duda 1925 Syn. 

Acanthoterna Duda 1925 Syn. 

Macropalpus Duda 1926 Homo. 

Adrosophila Seguy 1938 Syn. 

Sophophora Sturtevant 1939 Syn. 

Dichaetophora Duda 1940 Syn. 

Dichaetophila Duda 1940 Missp. 

Dorsilopha Sturtevant 1942 Syn. 

DIPTERA: DROSOPHILIDAE 


Phloridosa Sturtevant 1942 Syn. 

Pholadoris Sturtevant 1942 Syn. 

Dudaica Strand 1943 Syn. 

Siphlodora Patterson and Mainland 1944 Syn. 

Sordophila Wheeler 1949 Syn. 

Chusqueophila Brncic 1957 Syn. 

Lordiophsa Basden 1961 Syn. 

Trichotobregma Hardy 1965 Syn. 

Antopocerus Hardy 1965 Syn. 

Psilodorha Okada 1968 Syn. 

Engiscaptomyza Kaneshiro 1969 Syn. 

Drosophila acutilabella Stalker 1953 (Drosophila) 

Drosophila affinis Sturtevant 1916 (Drosophila) 

Drosophila iroquois Sturtevant and Dobzhansky 1936 Syn. 

Drosophila alabamensis Sturtevant 1918 (Drosophila) 

Drosophila albipes Walker 1852 (Drosophila) 

Drosophila aldrichi Patterson and Crow 1940 (Drosophila) 

Drosophila algonquin Sturtevant and Dobzhansky 1936 

(Drosophila) 

Drosophila americana Spencer 1938 (Drosophila) 

Drosophila texana Patterson 1940 Syn. 

Drosophila ananassae Doleschall 1858 (Drosophila) 

Drosophila imparata Walker 1859 Syn. 

Drosophila similis Lamb 1914 Homo. 

Drosophila caribea Sturtevant 1916 Syn. 

Drosophila errans Malloch 1933 Syn. 

Drosophila arizonae Ruiz and Wasserman 1990 (Drosophila) 

Drosophila athabasca Sturtevant and Dobzhansky 1936 

(Drosophila) 

Drosophila mahican Sturtevant and Dobzhansky 1936 Syn. 

Drosophila azteca Sturtevant and Dobzhansky 1936 (Drosophila) 

Drosophila basdeni Wheeler 1957 (Drosophila) 

Drosophila bifurca Patterson and Wheeler 1942 (Drosophila) 

Drosophila borealis Patterson 1952 (Drosophila) 

Drosophila brevis Walker 1852 (Drosophila) 

Drosophila brooksi Pipkin 1961 (Drosophila) 

Drosophila brooksae Wheeler 1981 Emend. 

Drosophila buskii Coquillett 1901 (Drosophila) 

Drosophila rubrostriata Becker 1908 Syn. 

Drosophila plurilineata Villeneuve 1911 Syn. 

Drosophila busckii Wheeler 1965 Emend. 

Drosophila californica Sturtevant 1923 (Drosophila) 

Drosophila fuliginea Patterson and Wheeler 1942 Syn. 

Drosophila carbonaria Patterson and Wheeler 1942 (Drosophila) 

Drosophila cardini Sturtevant 1916 (Drosophila) 

Drosophila carsoni Wheeler 1957 (Drosophila) 

Drosophila chagrinensis Stalker and Spencer 1939 (Drosophila) 

Drosophila cinerea Patterson and Wheeler 1942 (Drosophila) 

Drosophila colorata Walker 1849 (Drosophila) 

Drosophila sulcata Sturtevant 1916 Syn. 

Drosophila deflecta Malloch 1924 (Drosophila) 

Drosophila duncani Sturtevant 1918 (Drosophila) 

Drosophila euronotus Patterson and Ward 1952 (Drosophila) 

Drosophila falleni Wheeler 1960 (Drosophila) 

Drosophila flavomontana Patterson 1952 (Drosophila) 

Drosophila flavopinicola Wheeler 1954 (Drosophila) 

Drosophila floricola Sturtevant 1942 (Drosophila) 

Drosophila fronto Walker 1852 (Drosophila) 

Drosophila fulvalineata Patterson and Wheeler 1942 (Drosophila) 

Drosophila funebris Fabricius 1787 (Musca) 

Musca erythrophthalmua Panzer 1794 Syn. 

Drosophila clarkii Hutton 1901 Syn. 

Leucophenga atkinsoni Miller 1921 Syn. 

Drosophila dudai Malloch 1934 Syn. 

Drosophila grisea Patterson and Wheeler 1942 (Drosophila) 

Drosophila guttifera Walker 1849 (Drosophila) 

Drosophila multipunctata Loew 1866 Syn.


Drosophila hamatofila Patterson and Wheeler 1942 (Drosophila) 

Drosophila hydei Sturtevant 1921 (Drosophila) 

Drosophila yucatanensis Spencer 1940 Syn. 

Drosophila setosa Dobzhansky and Pavan 1943 Homo. 

Drosophila immigrans Sturtevant 1921 (Drosophila) 

Drosophila brouni Hutton 1901 Unav. 

Drosophila cilifemur Villeneuve 1923 Syn. 

Drosophila flexipilosa Pipkin 1964 Syn. 

Drosophila innubila Spencer 1943 (Drosophila) 

Drosophila lacicola Patterson 1944 (Drosophila) 

Drosophila latifasciaeformis Duda 1940 (Drosophila) 

Drosophila gracilipes Duda 1940 Syn. 

Drosophila mirim Dobzhansky and Pavan 1943 Syn. 

Drosophila baeomyia Wheeler 1949 Syn. 

Drosophila lebanonensis Wheeler 1949 (Drosophila) 

Drosophila casteeli Pipkin 1961 Syn. 

Drosophila linearis Walker 1852 (Drosophila) 

Drosophila longala Patterson and Wheeler 1942 (Drosophila) 

Drosophila longicornis Patterson and Wheeler 1942 (Drosophila) 

Drosophila lowei Heed, Crumpacker, Ehrman 1968 (Drosophila) 

Drosophila lutzii Sturtevant 1916 (Drosophila) 

Drosophila macroptera Patterson and Wheeler 1942 (Drosophila) 

Drosophila macrospina Stalker and Spencer 1939 (Drosophila) 

Drosophila ohioensis Spencer 1940 Syn. 

Drosophila limpiensis Mainland 1941 Syn. 

Drosophila magnaquinaria Wheeler 1954 (Drosophila) 

Drosophila mainlandi Patterson 1943 (Drosophila) 

Drosophila melanica Sturtevant 1916 (Drosophila) 

Drosophila melanissima Sturtevant 1916 (Drosophila) 

Drosophila melanogaster Meigen 1830 (Drosophila) 

Drosophila fasciata Meigen 1830 Syn. 

Drosophila nigriventris Macquart 1843 Syn. 

Drosophila approximata Zetterstedt 1847 Syn. 

Drosophila immatura Walker 1849 Syn. 

Drosophila ampelophila Loew 1862 Syn. 

Drosophila eumulata Ray-Chaudhuri and Mukherjee 1941 Syn. 

Drosophila melanura Miller 1944 (Drosophila) 

Drosophila mercatorum Patterson and Wheeler 1942 (Drosophila) 

Drosophila carinata Grimshaw 1901 Unav. 

Drosophila pararepleta Dobzhansky and Pavan 1943 Syn. 

Drosophila meridiana Patterson and Wheeler 1942 (Drosophila) 

Drosophila rioensis Patterson 1943 Syn. 

Drosophila mettleri Heed 1977 (Drosophila) 

Drosophila micromelanica Patterson 1941 (Drosophila) 

Drosophila minuta Walker 1852 (Drosophila) 

Drosophila miranda Dobzhansky 1935 (Drosophila) 

Drosophila mojavensis Patterson and Crow 1940 (Drosophila) 

Drosophila arizonensis Patterson and Wheeler 1942 Syn. 

Drosophila baja Mettler 1963 Syn. 

Drosophila montana Stone, Griffen, and Patterson 1941 

(Drosophila) 

Drosophila montana Patterson and Wheeler 1942 Homo. 

Drosophila ovivororum Lakovaara and Hackman 1973 Syn. 

Drosophila mulleri Sturtevant 1921 (Drosophila) 

Drosophila munda Spencer 1942 (Drosophila) 

Drosophila narragansett Sturtevant and Dobzhansky 1936 

(Drosophila) 

Drosophila nebulosa Sturtevant 1916 (Drosophila) 

Drosophila limbata Williston 1896 Homo. 

Drosophila nigrohalterata Duda 1925 (Drosophila) 

Drosophila nigrohydei Patterson and Wheeler 1942 (Drosophila) 

Drosophila hydeoides Patterson and Wheeler 1942 Syn. 

Drosophila nigromelanica Patterson and Wheeler 1942 (Drosophila) 

Drosophila nigrospiracula Patterson and Wheeler 1942 (Drosophila) 

Drosophila novamexicana Patterson 1941 (Drosophila) 

Drosophila novitskii Sulerud and Miller 1966 (Drosophila) 

Drosophila occidentalis Spencer 1942 (Drosophila) 

Drosophila orbospiracula Patterson and Wheeler 1942 (Drosophila) 


151 

Drosophila ordinaria Coquillett 1904 (Drosophila) 

Drosophila melanderi Sturtevant 1916 Syn. 

Drosophila magnafumosa Stalker and Spencer 1939 Syn. 

Drosophila pachea Patterson and Wheeler 1942 (Drosophila) 

Drosophila palustris Spencer 1942 (Drosophila) 

Drosophila lativittata Malloch 1924 Homo. 

Drosophila mallochi Frota-Pessoa 1946 Syn. 

Drosophila paramelanica Patterson 1942 (Drosophila) 

Drosophila peninsularis Patterson and Wheeler 1942 (Drosophila) 

Drosophila persimilis Dobzhansky and Epling 1944 (Drosophila) 

Drosophila pictiventris Duda 1925 (Drosophila) 

Drosophila pinicola Sturtevant 1942 (Drosophila) 

Drosophila polychaeta Patterson and Wheeler 1942 (Drosophila) 

Drosophila populi Wheeler and Throckmorton 1961 (Drosophila) 

Drosophila pseudoobscura Frolova 1929 (Drosophila) 

Drosophila bogotana Ayala and Dobzhansky 1974 Syn. 

Drosophila putrida Sturtevant 1916 (Drosophila) 

Drosophila pseudomelanica Sturtevant 1916 Syn. 

Drosophila quinaria Loew 1866 (Drosophila) 

Drosophila recens Wheeler 1960 (Drosophila) 

Drosophila rellima Wheeler 1960 (Drosophila) 

Drosophila repleta Wollaston 1858 (Drosophila) 

Drosophila punctulata Loew 1862 Syn. 

Drosophila adspersa Mik 1886 Syn. 

Drosophila nigropunctata Wulp 1892 Syn. 

Drosophila maculiventris Wulp 1897 Syn. 

Drosophila prorepleta Duda 1925 Syn. 

Drosophila pygmaea Duda 1927 Homo. 

Drosophila melanopalpa Patterson and Wheeler 1942 Syn. 

Drosophila austrorepleta Dobzhansky and Pavan 1943 Syn. 

Drosophila betari Dobzhansky and Pavan 1943 Syn. 

Drosophila brunneipalpa Dobzhansky and Pavan 1943 Syn. 

Drosophila pumiliaris Wheeler 1981 Syn. 

Drosophila ritae Patterson and Wheeler 1942 (Drosophila) 

Drosophila tira Wasserman 1962 Syn. 

Drosophila robusta Sturtevant 1916 (Drosophila) 

Drosophila rubrifrons Patterson and Wheeler 1942 (Drosophila) 

Drosophila seminole Sturtevant and Dobzhansky 1936 (Drosophila) 

Drosophila sigmoides Loew 1872 (Drosophila) 

Drosophila simulans Sturtevant 1919 (Drosophila) 

Drosophila stalkeri Wheeler 1954 (Drosophila) 

Drosophila subfunebris Stalker and Spencer 1939 (Drosophila) 

Drosophila suboccidentalis Spencer 1942 (Drosophila) 

Drosophila subpalustris Spencer 1942 (Drosophila) 

Drosophila suffusca Spencer 1943 (Drosophila) 

Drosophila tenebrosa Spencer 1943 (Drosophila) 

Drosophila tenuipes Walker 1849 (Diastata) 

Drosophila testacea Roser 1840 (Drosophila) 

Drosophila nigrithorax Strobl 1894 Syn. 

Drosophila setosa Villeneuve 1921 Syn. 

Drosophila thoracis Williston 1896 (Drosophila) 

Drosophila transversa Fallen 1823 (Drosophila) 

Drosophila laeta Zetterstedt 1847 Syn. 

Drosophila subquinaria Spencer 1942 Syn. 

Drosophila tripunctata Loew 1862 (Drosophila) 

Drosophila modesta Sturtevant 1916 Syn. 

Drosophila trispina Wheeler 1949 (Drosophila) 

Drosophila victoria Sturtevant 1942 (Drosophila) 

Drosophila virilis Sturtevant 1916 (Drosophila) 

Drosophila wheeleri Patterson and Alexander 1952 (Drosophila) 

Drosophila willistoni Sturtevant 1916 (Drosophila) 

Drosophila pallida Williston 1896 Homo. 

Drosophila paulista Dobzhansky and Pavan 1943 Syn. 

Drosophila quechua Ayala 1973 Syn. 

Gitona Meigen 1830 

Gitona americana Patterson 1943 (Gitona)

152 

Gitona bivisualis Patterson 1943 (Gitona) 

Gitona sonoita Wheeler 1949 (Gitona) 

Leucophenga Mik 1886 

Oxyleucophenga Hendel 1913 Syn. 

Drosomyiella Hendel 1914 Syn. 

Paraleucophenga Oldenberg 1914 Homo. 

Neoleucophenga Oldenberg 1915 Syn. 

Ptyelusimyia Seguy 1932 Syn. 

Drosophilopsis Seguy 1951 Syn. 

Leucophenga guttata Wheeler 1952 (Leucophenga) 

Leucophenga maculosa Coquillett 1895 (Drosophila) 

Leucophenga fenestrata Duda 1927 Syn. 

Leucophenga schnuseana Duda 1927 Syn. 

Leucophenga montana Wheeler 1952 (Leucophenga) 

Leucophenga neovaria Wheeler 1960 (Leucophenga) 

Leucophenga pulcherrima Patterson and Mainland 1944 

(Leucophenga) 

Leucophenga trisphenata Wheeler 1952 (Leucophenga) 

Leucophenga varia Walker 1849 (Drosophila) 

Drosophila quadrimaculata Walker 1852 Syn. 

Opomyza signicosta Walker 1861 Syn. 

Microdrosophila Malloch 1921 

Oxystyloptera Duda 1924 Syn. 

Incisurifrons Duda 1924 Syn. 

Hopkinsomyia Malloch 1934 Syn. 

Microdrosophila quadrata Sturtevant 1916 (Drosophila) 

Mycodrosophila Oldenberg 1914 

Mycodrosophila claytonae Wheeler and Takada 1963 

(Mycodrosophila) 

Mycodrosophila dimidiata Loew 1862 (Drosophila) 

Mycodrosophila stalkeri Wheeler and Takada 1963 

(Mycodrosophila) 

Paramycodrosophila Duda 1924 

Upolymyia Malloch 1934 Syn. 



Paramycodrosophila anomala Wheeler 1954 (Paramycodrosophila) 

Paramycodrosophila centralis Wheeler 1954 (Paramycodrosophila) 

Pseudiastata Coquillett 1908 

Hyalistata Wheeler 1960 Syn. 

Pseudiastata nebulosa Coquillett 1908 (Pseudiastata) 

Pseudiastata pictiventris Wheeler 1960 (Pseudiastata) 

Rhinoleucophenga Hendel 1917 

Pseudophortica Sturtevant 1918 Syn. 

Rhinoleucophenga obesa Loew 1872 (Drosophila) 

Phortica hirtifrons Johnson 1913 Syn. 

Scaptomyza Hardy 1849 

Parascaptomyza Duda 1924 Syn. 

Scaptomyzella Hendel 1928 Syn. 

Scaptomyzetta Hendel 1928 Missp. 

Bunostoma Malloch 1932 Syn. 

Rosenwaldia Malloch 1934 Syn. 

Euscaptomyza Seguy 1938 Syn. 

Tantalia Malloch 1938 Syn. 

Macroscaptomyza Frey 1954 Syn. 

Ctenoscaptomyza Frey 1954 Syn. 

Tristanomyia Frey 1954 Syn. 

Trogloscaptomyza Frey 1954 Syn. 

Hemiscaptomyza Hackman 1959 Syn. 

Mesoscaptomyza Hackman 1959 Syn. 

Metascaptomyza Hackman 1959 Syn. 

Alloscaptomyza Hackman 1962 Syn. 

Dentiscaptomyza Takada 1965 Syn. 

Exalloscaptomyza Hardy 1965 Syn. 

Boninoscaptomyza Okada 1973 Syn. 

Lauxanomyza Tsacas and Cogan 1976 Syn. 

Elmomyza Hackman 1982 Syn. 

Scaptomyza adusta Loew 1862 (Drosophila) 

Scaptomyza apicata Thomson 1869 (Drosophila) 

Scaptomyza bipunctipennis Wheeler 1952 (Scaptomyza) 

Scaptomyza flaviventris Hackman 1959 (Scaptomyza) 

Scaptomyza graminum Fallen 1823 (Drosophila) 

Drosophila incana Meigen 1830 Syn. 

Drosophila rufipes Meigen 1830 Syn. 

Hydrellia amoena Meigen 1838 Syn. 

Drosophila flavipennis Zetterstedt 1838 Syn. 

Drosophila sordida Zetterstedt 1838 Syn. 

Scaptomyza tetrasticha Becker 1908 Syn. 

Drosophila semiatricornis Duda 1935 Syn. 

Scaptomyza borealis Wheeler 1952 Syn. 

Scaptomyza norica Hackman 1955 Syn. 

Scaptomyza hirsuta Wheeler 1949 (Scaptomyza) 

Scaptomyza hsui Hackman 1955 (Scaptomyza) 

Scaptomyza maculifera Becker 1919 (Scaptomyza) 

Scaptomyza parapicata Hackman 1959 Syn. 

Scaptomyza malada Wheeler and Takada 1966 (Scaptomyza) 

Scaptomyza montana Wheeler 1949 (Scaptomyza) 

Scaptomyza nigrocella Wheeler 1949 Syn. 

Scaptomyza nigrita Wheeler 1952 (Scaptomyza) 

Scaptomyza pallida Zetterstedt 1847 (Drosophila) 

Drosophila disticha Duda 1921 Syn. 

Scaptomyza paradusta Wheeler 1952 (Scaptomyza) 

Scaptomyza paravittata Wheeler 1952 (Scaptomyza) 

Scaptomyza teinoptera Hackman 1955 (Scaptomyza) 

Scaptomyza terminalis Loew 1863 (Drosophila) 

Scaptomyza trochanterata Collin 1953 (Scaptomyza) 

Scaptomyza unipunctum Zetterstedt 1847 (Geomyza) 

Scaptomyza bocharensis Hackman 1959 Syn. 

Scaptomyza vittata Coquillett 1895 (Drosophila) 

Scaptomyza wheeleri Hackman 1959 (Scaptomyza) 

Stegana Meigen 1830 

Orthostegana Hendel 1913 Syn. 

Ceratostylus Enderlein 1922 Syn. 

Oxyphortica Duda 1923 Syn. 

Chaetocnema Duda 1926 Homo. 

Steganina Wheeler 1960 Syn. 

Parastegana Okada 1971 Syn. 

Pseudostegana Okada 1978 Syn. 

Stegana antigua Wheeler 1960 (Stegana) 

Stegana coleoptrata Scopoli 1763 (Musca) 

Stegana annulata Haliday 1833 Syn. 

Stegana dentifera Lastovka and Maca 1982 (Stegana) 

Stegana vittata Coquillett 1901 (Phortica) 

Stegana wheeleri Lastovka and Maca 1982 (Stegana)

DRYOMYZIDAE 

Dryomyza Fallen 1820 

Dryope Robineau-Desvoidy 1830 Syn. 

Neuroctena Rondani 1868 Syn. 

Stenodryomyza Hendel 1923 Syn. 

Dryomyza anilis Fallen 1820 (Dryomyza) 

Dryomyza pallida Day 1881 Syn. 

Dryomyza bergi Steyskal 1957 (Dryomyza) 

Dryomyza ferruginea Melander 1920 (Dryomyza) 

Dryomyza flaveola Fabricius 1794 (Musca) 

Dryomyza dayi Cresson 1920 Syn. 

Dryomyza melanderi Steyskal 1957 (Dryomyza) 

Dryomyza maculipennis Melander 1920 Homo. 

Dryomyza setosa Bigot 1886 (Odontomera) 

Neuroctena fumida Coquillett 1901 Syn. 

Dryomyza simplex Loew 1862 (Dryomyza) 

Neuroctena ruthae Brimley 1925 Syn. 

Helcomyza Curtis 1825 

Actora Meigen 1826 Syn. 

Macromelanderia Curran 1934 Syn. 

Helcomyza mirabilis Melander 1920 (Helcomyza) 

Heterocheila Rondani 1857 

Heterostoma Rondani 1856 Homo. 

Oedoparea Loew 1862 Syn. 

Pseudosciomyza Malloch 1923 Syn. 

Heterocheila hannai Cole 1921 (Dryomyza) 

Heterocheila nudiseta Curran 1933 Syn. 

Oedoparena Curran 1934 

Oedoparena glauca Coquillett 1900 (Oedoparea) 

Oedoparena nigrifrons Mathis and Steyskal 1980 (Oedoparena) 

EMPIDIDAE 

Allanthalia Melander 1928 

Allanthalia pallida Zetterstedt 1838 (Anthalia) 

Allodromia Smith 1962 

Allodromia testacea Melander 1928 (Micrempis) 

Anomalempis Melander 1928 

Anomalempis archon Melander 1945 (Anomalempis) 

Anomalempis tacomae Melander 1928 (Anomalempis) 

Anthalia Zetterstedt 1838 

Anthalia bulbosa Melander 1902 (Euthyneura) 

Anthalia femorata Melander 1928 (Anthalia) 

Anthalia flava Coquillett 1903 (Anthalia) 

Anthalia gilvihirtus Coquillett 1903 (Microphorus) 


Anthalia inornata Melander 1928 (Anthalia) 

Anthalia interrupta Melander 1928 (Anthalia) 

Anthalia lacteipennis Melander 1928 (Anthalia) 

Anthalia mandalota Melander 1928 (Anthalia) 

Anthalia schoenherri Zetterstedt 1838 (Anthalia) 

Anthalia scutellaris Melander 1928 (Anthalia) 

Anthalia stigmalis Coquillett 1903 (Anthalia) 

Anthalia petiolata Melander 1928 Syn. 

Anthepiscopus Becker 1891 

Anthepiscopus flavicoxa Melander 1928 (Anthepiscopus) 

Anthepiscopus flavipilosus Coquillett 1900 (Microphorus) 

Anthepiscopus hirsutus Melander 1928 (Anthepiscopus) 

Anthepiscopus longipalpis Melander 1928 (Anthepiscopus) 

Anthepiscopus nuptus Melander 1928 (Anthepiscopus) 

Anthepiscopus polygynus Melander 1928 (Anthepiscopus) 

Anthepiscopus stentor Melander 1902 (Euthyneura) 

Apalocnemis Philippi 1865 

Apalocnemis hirsuta Melander 1946 (Apalocnemis) 

Apalocnemis oreas Melander 1946 (Apalocnemis) 

Bicellaria Macquart 1823 

Cyrtoma Meigen 1824 Syn. 

Enicopteryx Stephens 1829 Unav. 

Bicellaria angustifurca Melander 1928 (Bicellaria) 

Bicellaria brevifurca Melander 1928 (Bicellaria) 

Bicellaria furcifer Melander 1928 (Bicellaria) 

Bicellaria halteralis Loew 1862 (Cyrtoma) 

Bicellaria longipes Loew 1862 (Cyrtoma) 

Bicellaria lugubris Melander 1928 (Bicellaria) 

Bicellaria pectinata Melander 1928 (Bicellaria) 

Bicellaria pilipes Loew 1862 (Cyrtoma) 

Bicellaria spuria Fallen 1816 (Empis) 

Bicellaria uvens Melander 1928 (Bicellaria) 

Boreodromia Coquillett 1903 

Boreodromia bicolor Loew 1863 (Synamphotera) 

Brachystoma Meigen 1822 

Blepharoprocta Loew 1862 Syn. 

Brachystoma nigrimanus Loew 1862 (Brachystoma) 

Brachystoma occidentalis Melander 1902 (Brachystoma) 

Brachystoma robertsonii Coquillett 1895 (Brachystoma) 

Brachystoma serrulata Loew 1861 (Brachystoma) 

Brachystoma binummus Loew 1862 Syn. 

Brochella Melander 1928 

Brochella monticola Melander 1928 (Brochella) 

Ceratempis Melander 1928 

Ceratempis longicornis Melander 1928 (Ceratempis) 

Charadrodromia Melander 1928 

Charadrodromia arnaudi Melander 1960 (Charadrodromia) 

Charadrodromia microphona Melander 1928 (Charadrodromia) 

Charadrodromia syletor Melander 1928 (Charadrodromia) 

DIPTERA: DRYOMYZIDAE, EMPIDIDAE 

153

154 

Chelifera Macquart 1823 

Mantipeza Rondani 1856 Syn. 

Polydromia Bigot 1857 Syn. 

Thanategia Melander 1928 Syn. 

Chelifera banksi Melander 1947 (Chelifera) 

Chelifera bidenta MacDonald 1994 (Chelifera) 

Chelifera caliga Lavallee 1975 (Chelifera) 

Chelifera chvalai Wagner 1984 (Chelifera) 

Chelifera circinata MacDonald 1994 (Chelifera) 

Chelifera cirrata Melander 1947 (Chelifera) 

Chelifera defecta Loew 1862 (Hemerodromia) 

Chelifera knutsoni Lavallee 1975 Syn. 

Chelifera ensifera Melander 1947 (Chelifera) 

Chelifera lovetti Melander 1947 (Chelifera) 

Chelifera mana Lavallee 1975 (Chelifera) 

Chelifera multidenta MacDonald 1994 (Chelifera) 

Chelifera neangusta MacDonald 1994 (Chelifera) 

Chelifera notata Loew 1862 (Hemerodromia) 

Chelifera obsoleta Loew 1862 (Hemerodromia) 

Chelifera palloris Coquillett 1895 (Mantipeza) 

Chelifera rastrifera Melander 1947 (Chelifera) 

Chelifera recurvata Melander 1947 (Thanategia) 

Chelifera stuprator Melander 1947 (Thanategia) 

Chelifera subensifera MacDonald 1994 (Chelifera) 

Chelifera subnotata MacDonald 1994 (Chelifera) 

Chelifera valida Loew 1862 (Hemerodromia) 

Chelifera scrotifera Melander 1947 Syn. 

Chelifera varix Melander 1947 (Chelifera) 

Chelipoda Macquart 1823 

Phyllodromia Zetterstedt 1837 Syn. 

Chiromantis Rondani 1856 Homo. 

Chyromantis Rondani 1856 Missp. 

Lepidomya Bigot 1857 Homo. 

Thamnodromia Mik 1886 Syn. 

Litanomyia Melander 1902 Syn. 

Lepidomyia Kertesz 1909 Emend. 

Chelipoda albiseta Zetterstedt 1838 (Hemerodromia) 

Chelipoda americana Melander 1947 (Phyllodromia) 

Chelipoda contracta Melander 1947 (Chelipoda) 

Chelipoda elongata Melander 1902 (Litanomyia) 

Chelipoda limitaria MacDonald 1993 (Chelipoda) 

Chelipoda praestans Melander 1947 (Chelipoda) 

Chelipoda sicaria Melander 1947 (Chelipoda) 

Chelipoda truncata MacDonald 1993 (Chelipoda) 

Chelipoda vocatoria Fallen 1815 (Tachydromia) 

Chersodromia Walker 1851 

Coloboneura Melander 1902 Syn. 

Halsanalotes Becker 1902 Syn. 

Thinodromia Melander 1906 Syn. 

Chersodromia cana Melander 1945 (Chersodromia) 

Chersodromia houghii Melander 1902 (Stilpon) 

Chersodromia inchoata Melander 1906 (Thinodromia) 

Chersodromia insignita Melander 1945 (Chersodromia) 

Chersodromia inusitata Melander 1902 (Coloboneura) 

Chersodromia magacetes Melander 1945 (Chersodromia) 

Chersodromia nana Coquillett 1903 (Coloboneura) 

Chersodromia parallela Melander 1928 (Thinodromia) 

Clinocera Meigen 1803 

Atalanta Meigen 1800 Unav. 

Hydrodromia Macquart 1835 Syn. 

Paramesia Macquart 1835 Syn. 

Wiedemannia Bigot 1856 Homo. 

DIPTERA: EMPIDIDAE 


Bergenstammia Mik 1881 Syn. 

Phaeobalia Mik 1881 Syn. 

Kowarzia Mik 1881 Syn. 

Archiclinocera Frey 1945 Syn. 

Hypoclinocera Frey 1945 Syn. 

Clinocera appendiculata Zetterstedt 1838 (Wiedemannia) 

Brachystoma aucta Zetterstedt 1849 Syn. 

Clinocera binotata Loew 1876 (Clinocera) 

Clinocera brevitibia Melander 1928 (Clinocera) 

Clinocera conjuncta Loew 1860 (Clinocera) 

Clinocera fuscipennis Loew 1876 (Clinocera) 

Clinocera genualis Coquillett 1910 (Clinocera) 

Clinocera lecta Melander 1902 (Clinocera) 

Clinocera lineata Loew 1862 (Clinocera) 

Clinocera longifurca Melander 1928 (Clinocera) 

Clinocera maculata Loew 1860 (Clinocera) 

Clinocera olivacea Melander 1928 (Clinocera) 

Clinocera prasinata Melander 1928 (Clinocera) 

Clinocera stagnalis Haliday 1833 (Heleodromia) 

Clinocera taos Melander 1902 (Clinocera) 

Clinocera trunca Melander 1928 (Clinocera) 

Clinocera undulata Melander 1928 (Clinocera) 

Dolichocephala Macquart 1823 

Ardoptera Macquart 1827 Syn. 

Leptosceles Haliday 1833 Syn. 

Dolichocephala argus Melander 1928 (Dolichocephala) 

Dolichocephala irrorata Fallen 1815 (Tachydromia) 

Drapetis Meigen 1822 

Elaphropeza Macquart 1827 Syn. 

Caecula Gistl 1848 Syn. 

Arbicola Gistl 1848 Syn. 

Crossopalpus Bigot 1857 Syn. 

Eudrapetis Melander 1918 Syn. 

Drapetis aliternigra Melander 1918 (Drapetis) 

Drapetis armata Melander 1918 (Drapetis) 

Drapetis assimilis Fallen 1815 (Tachydromia) 

Drapetis nigra Meigen 1830 Syn. 

Drapetis bispina Melander 1918 (Drapetis) 

Drapetis cerina Rogers 1989 (Drapetis) 

Drapetis deceptor Curran 1929 (Drapetis) 

Drapetis discalis Melander 1918 (Drapetis) 

Drapetis divergens Loew 1872 (Drapetis) 

Drapetis diversa Melander 1918 (Drapetis) 

Drapetis diversipes Melander 1918 (Drapetis) 

Drapetis dividua Melander 1902 (Drapetis) 

Stilpon nigripes Melander 1902 Syn. 

Drapetis facialis Melander 1918 (Drapetis) 

Drapetis gilvipes Loew 1872 (Drapetis) 

Drapetis incultus Coquillett 1895 (Platypalpus) 

Drapetis infumata Melander 1918 (Drapetis) 

Drapetis lata Coquillett 1903 (Tachydromia) 

Drapetis latipennis Melander 1902 (Drapetis) 

Drapetis mariae Steyskal 1953 (Drapetis) 

Drapetis medetera Melander 1902 (Drapetis) 

Drapetis micropyga Melander 1918 (Drapetis) 

Drapetis naica Melander 1918 (Drapetis) 

Drapetis parvicornis Melander 1918 (Drapetis) 

Drapetis pilosa Melander 1918 (Drapetis) 

Drapetis plumipes Melander 1918 (Drapetis) 

Drapetis populi Steyskal 1953 (Drapetis) 

Drapetis scissa Melander 1918 (Drapetis) 

Drapetis septentrionalis Melander 1902 (Drapetis) 

Drapetis setulosa Melander 1918 (Drapetis)

Drapetis spectabilis Melander 1902 (Drapetis) 

Drapetis trichura Melander 1918 (Drapetis) 

Drapetis unipila Loew 1872 (Drapetis) 

Drapetis nitida Melander 1902 Syn. 

Drapetis vittata Melander 1918 (Drapetis) 

Drapetis xanthopoda Williston 1896 (Drapetis) 

Empis Linnaeus 1758 

Platyptera Meigen 1803 Syn. 

Pachymeria Stephens 1829 Syn. 

Platypterygia Stephens 1829 Syn. 

Pachymerina Macquart 1834 Syn. 

Eriogaster Macquart 1838 Homo. 

Zapolia Gistl 1848 Syn. 

Enoplempis Bigot 1880 Syn. 

Empimorpha Coquillett 1895 Syn. 

Pterempis Bezzi 1909 Syn. 

Xanthempis Bezzi 1909 Syn. 

Anacrostichus Bezzi 1909 Syn. 

Polyblepharis Bezzi 1909 Syn. 

Argyrandrus Bezzi 1909 Syn. 

Coptophlebia Bezzi 1909 Syn. 

Lissempis Bezzi 1909 Syn. 

Kritempis Collin 1926 Syn. 

Leptempis Collin 1926 Syn. 

Rhadinempis Collin 1926 Syn. 

Acallomyia Melander 1928 Syn. 

Pyrrempis Melander 1928 Syn. 

Rhagionempis Engel 1943 Syn. 

Euempis Frey 1953 Syn. 

Planempis Frey 1953 Syn. 

Disneyempis Smith 1976 Syn. 

Empis abcirus Walker 1849 (Empis) 

Empis aeripes Melander 1902 (Empis) 

Empis aerobatica Melander 1902 (Empis) 

Empis agasthus Walker 1849 (Empis) 

Empis aldrichii Melander 1902 (Empis) 

Empis amytis Walker 1849 (Empis) 

Empis anthophila Melander 1946 (Empis) 

Empis armipes Loew 1861 (Empis) 

Empis arthrica Melander 1902 (Empis) 

Empis asema Melander 1902 (Empis) 

Empis barbatoides Melander 1965 (Empis) 

Empis barbata Loew 1862 Homo. 

Empis bigoti Melander 1902 (Empis) 

Enoplempis cinerea Bigot 1882 Homo. 

Empis brachysoma Coquillett 1900 (Empis) 

Empis brevis Loew 1862 (Pachymeria) 

Empis browni Curran 1931 (Empis) 

Empis brunnea Coquillett 1903 (Empis) 

Empis bullifera Kessel and Kessel 1951 (Empis) 

Empis cacuminifer Melander 1902 (Empis) 

Empis caeligena Melander 1902 (Empis) 

Empis canaster Melander 1902 (Empis) 

Empis captus Coquillett 1895 (Empis) 

Empis clausa Coquillett 1895 (Empis) 

Empis colonica Walker 1849 (Empis) 

Empis comantis Coquillett 1895 (Empimorpha) 

Empis cometes Steyskal 1965 (Empis) 

Empis comantis Coquillett 1895 Homo. 

Empis compta Coquillett 1895 (Empis) 

Empis ctenocnema Melander 1945 (Empis) 

Empis dactylica Melander 1946 (Empis) 

Empis delumbis Melander 1965 (Empis) 

Empis clauda Coquillett 1900 Homo. 

Empis desiderata Melander 1965 (Empis) 

Empis avida Coquillett 1895 Homo. 


Empis deterra Walley 1928 (Empis) 

Empis distans Loew 1869 (Empis) 

Empis dolabraria Melander 1902 (Empis) 

Empis disconvenita Melander 1902 Syn. 

Empis enodis Melander 1902 (Empis) 

Empis eudamides Walker 1849 (Empis) 

Empis exilis Coquillett 1903 (Empis) 

Empis falcata Melander 1902 (Empis) 

Empis frontalis Coquillett 1903 (Empis) 

Empis fumida Coquillett 1900 (Empis) 

Empis geneatis Melander 1902 (Empimorpha) 

Empis gladiator Melander 1902 (Empis) 

Empis gulosa Coquillett 1895 (Empis) 

Empis hirticrus Melander 1928 (Empis) 

Empis hirtipes Coquillett 1903 Homo. 

Empis tenebrosa Coquillett 1903 Homo. 

Empis humilis Coquillett 1895 (Empis) 

Empis infumata Coquillett 1900 (Empis) 

Empis johnsoni Melander 1902 (Empis) 

Empis labiata Loew 1861 (Empis) 

Empis laevigata Loew 1864 (Empis) 

Empis laniventris Eschscholtz 1823 (Empis) 

Empis latrappensis Ouellet 1942 (Empis) 

Empis leptogastra Loew 1863 (Empis) 

Empis levicula Coquillett 1895 (Empis) 

Empis longeoblita Steyskal 1965 (Empis) 

Empis longipes Loew 1864 Homo. 

Empis loripedis Coquillett 1895 (Empis) 

Empis manca Coquillett 1895 (Empis) 

Empis melanderi Arnaud and Birchim 1966 (Empis) 

Empis mira Bigot 1880 (Enoplempis) 

Empis mixopolia Melander 1902 (Empis) 

Empis montiradicis James 1942 (Empis) 

Empis nodipoplitea Steyskal 1965 (Empis) 

Empis nodipes Melander 1902 Homo. 

Empis nuda Loew 1862 (Empis) 

Empis obesa Loew 1861 (Empis) 

Empis ravida Coquillett 1895 Syn. 

Empis ollius Walker 1849 (Empis) 

Empis otiosa Coquillett 1895 (Empis) 

Empis pallida Loew 1861 (Empis) 

Empis pellucida Coquillett 1900 (Empis) 

Empis peregrina Melander 1902 (Iteaphila) 

Iteaphila perigrina Melander 1902 Missp. 

Empis plectrum Melander 1946 (Empis) 

Empis podagra Melander 1902 (Empis) 

Empis poeciloptera Loew 1861 (Empis) 

Empis poplitea Loew 1863 (Empis) 

Empis serperastrorum Melander 1902 Syn. 

Empis pudica Loew 1861 (Pachymeria) 

Empis reciproca Walker 1857 (Empis) 

Empis rufescens Loew 1864 (Empis) 

Empis scatophagina Melander 1902 (Empis) 

Empis scoparia Coquillett 1903 (Empis) 

Empis snoddyi Steyskal 1969 (Empis) 

Empis sordida Loew 1861 (Empis) 

Empis spectabilis Loew 1862 (Empis) 

Empis stenoptera Loew 1864 (Empis) 

Empis subinfumata Malloch 1923 (Empis) 

Empis tenebrosa Coquillett 1895 (Empis) 

Empis teres Melander 1902 (Empis) 

Empis tersa Coquillett 1895 (Empis) 

Empis tridentata Coquillett 1901 (Empis) 

Empis vaginifer Melander 1902 (Empis) 

Empis valentis Coquillett 1895 (Empis) 

Empis varipes Loew 1861 (Empis) 

Empis virgata Coquillett 1895 (Empis) 


155

156 

Euhybus Coquillett 1895 

Euhybus baropeodes Melander 1928 (Euhybus) 

Euhybus coquilletti Melander 1928 (Euhybus) 

Euhybus cuspidatus Melander 1928 (Euhybus) 

Euhybus duplex Walker 1849 (Hybos) 

Euhybus genitivus Melander 1928 (Euhybus) 

Euhybus metatarsalis Melander 1928 (Euhybus) 

Euhybus nigripes Melander 1928 (Euhybus) 

Euhybus purpureus Walker 1849 (Hybos) 

Euhybus sordipes Melander 1928 (Euhybus) 

Euhybus strumaticus Melander 1928 (Euhybus) 

Euhybus subjectus Walker 1849 (Hybos) 

Euhybus triplex Walker 1849 (Hybos) 

Euhybus simplex Melander 1928 Syn. 

Euthyneura Macquart 1836 

Euthyneura aperta Melander 1902 (Euthyneura) 

Euthyneura argyria Melander 1928 (Euthyneura) 

Euthyneura bucinator Melander 1902 (Euthyneura) 

Euthyneura crocatus Coquillett 1900 (Microphorus) 

Euthyneura matura Melander 1928 (Euthyneura) 

Euthyneura spinipes Melander 1928 (Euthyneura) 

Gloma Meigen 1822 

Gloma fuscipes Melander 1945 (Gloma) 

Gloma luctuosa Melander 1928 (Gloma) 

Gloma pectinipes Melander 1945 (Gloma) 

Heleodromia Haliday 1833 

Microcera Zetterstedt 1838 Syn. 

Sciodromia Westwood 1840 Syn. 

Illiesiella Wagner 1985 Syn. 

Heleodromia pullata Melander 1902 (Sciodromia) 

Hemerodromia Meigen 1822 

Microdromia Bigot 1857 Syn. 

Hemerodromia brevifrons Melander 1947 (Hemerodromia) 

Hemerodromia brunnea Melander 1928 (Hemerodromia) 

Hemerodromia captus Coquillett 1895 (Hemerodromia) 

Hemerodromia chillcotti Harper 1974 (Hemerodromia) 

Hemerodromia coleophora Melander 1928 (Hemerodromia) 

Hemerodromia empiformis Say 1823 (Ochthera) 

Hemerodromia exhibitor Melander 1947 (Hemerodromia) 

Hemerodromia fibrina Landry and Harper 1985 (Hemerodromia) 

Hemerodromia haruspex Melander 1947 (Hemerodromia) 

Hemerodromia jugulator Melander 1928 (Hemerodromia) 

Hemerodromia melanosoma Melander 1947 (Hemerodromia) 

Hemerodromia rogatoris Coquillett 1895 (Hemerodromia) 

Hemerodromia stellaris Melander 1947 (Hemerodromia) 

Hemerodromia sufflexa Melander 1947 (Hemerodromia) 

Hemerodromia superstitiosa Say 1824 (Hemerodromia) 

Hemerodromia vates Melander 1947 (Hemerodromia) 

Hemerodromia vittata Loew 1862 (Hemerodromia) 

Hesperempis Melander 1906 

Hesperempis mabelae Melander 1902 (Ragas) 

Hesperempis sanduca Melander 1928 (Hesperempis) 

Hilara Meigen 1822 

Pseudoragas Frey 1952 Syn. 

Platyhilara Frey 1952 Syn. 

Ochtherohilara Frey 1952 Syn. 



Meroneurula Frey 1952 Syn. 

Pseudorhamphomyia Frey 1953 Syn. 

Hilara argyrata Curran 1930 (Hilara) 

Hilara atra Loew 1862 (Hilara) 

Hilara aurata Coquillett 1900 (Hilara) 

Hilara auripila Curran 1926 (Hilara) 

Hilara baculifer Melander 1902 (Hilara) 

Hilara basalis Loew 1862 (Hilara) 

Hilara bella Melander 1902 (Hilara) 

Hilara brevipila Loew 1862 (Hilara) 

Hilara cana Coquillett 1895 (Hilara) 

Hilara carbonaria Melander 1902 (Hilara) 

Hilara cavernicola Melander 1945 (Hilara) 

Hilara congregaria Melander 1902 (Hilara) 

Hilara crickmayi Curran 1926 (Hilara) 

Hilara dixi Steyskal 1969 (Hilara) 

Hilara femorata Loew 1862 (Hilara) 

Hilara garretti Curran 1926 (Hilara) 

Hilara gracilis Loew 1862 (Hilara) 

Hilara granditarsis Curran 1926 (Hilara) 

Hilara johnsoni Coquillett 1895 (Hilara) 

Hilara juno Curran 1930 (Hilara) 

Hilara leucoptera Loew 1862 (Hilara) 

Hilara lutea Loew 1863 (Hilara) 

Hilara macroptera Loew 1863 (Hilara) 

Hilara migrata Walker 1849 (Hilara) 

Hilara mutabilis Loew 1862 (Hilara) 

Hilara nearctica Shewell 1955 (Hilara) 

Hilara nigriventris Loew 1862 (Hilara) 

Hilara nugax Melander 1902 (Hilara) 

Hilara plebeia Walker 1857 (Hilara) 

Hilara quadrivittata Meigen 1822 (Hilara) 

Hilara rufopuncta Curran 1926 (Hilara) 

Hilara seriata Loew 1864 (Hilara) 

Hilara testacea Loew 1864 (Hilara) 

Hilara transfuga Walker 1849 (Hilara) 

Hilara tristis Loew 1864 (Hilara) 

Hilara trivittata Loew 1862 (Hilara) 

Hilara umbrosa Loew 1862 (Hilara) 

Empis brachystoma Coquillett 1900 Unav. 

Hilara unicolor Loew 1862 (Hilara) 

Hilara varipennis Curran 1930 (Empis) 

Hilara velutina Loew 1862 (Hilara) 

Hilara wheeleri Melander 1901 (Hilara) 

Hoplocyrtoma Melander 1928 

Hoplocyrtoma femorata Loew 1864 (Cyrtoma) 

Hoplocyrtoma procera Loew 1864 (Cyrtoma) 

Hormopeza Zetterstedt 1838 

Hormopeza brevicornis Loew 1864 (Hormopeza) 

Hormopeza bullata Melander 1902 (Hormopeza) 

Hormopeza copulifera Melander 1928 (Hormopeza) 

Hormopeza fumicola Steyskal 1969 (Hormopeza) 

Hormopeza nigricans Loew 1864 (Hormopeza) 

Hormopeza senator Melander 1928 (Hormopeza) 

Hormopeza virgator Melander 1928 (Hormopeza) 

Hybos Meigen 1803 

Noeza Meigen 1800 Unav. 

Hybos reversus Walker 1849 (Hybos) 

Hybos slossonae Coquillett 1895 Syn.

Iteaphila Zetterstedt 1838 

Stelochaeta Becker 1887 Syn. 

Iteaphila cana Melander 1946 (Iteaphila) 

Iteaphila conjuncta Coquillett 1900 (Empis) 

Iteaphila cormus Walker 1849 (Empis) 

Iteaphila fuliginosa Melander 1946 (Iteaphila) 

Iteaphila luctuosa Kirby 1837 (Empis) 

Empis geniculata Kirby 1837 Syn. 

Iteaphila macquarti Zetterstedt 1838 (Iteaphila) 

Iteaphila curva Curran 1925 Syn. 

Iteaphila napaea Melander 1946 (Iteaphila) 

Iteaphila nitidula Zetterstedt 1838 (Iteaphila) 

Iteaphila americana Melander 1946 Syn. 

Iteaphila orchestris Melander 1902 (Iteaphila) 

Iteaphila testacea Melander 1946 (Iteaphila) 

Iteaphila triangula Coquillett 1900 (Empis) 

Iteaphila vetula Melander 1946 (Iteaphila) 

Leptopeza Macquart 1827 

Leptopeza antennalis Melander 1928 (Leptopeza) 

Leptopeza borealis Zetterstedt 1842 (Leptopeza) 

Leptopeza compta Coquillett 1895 (Leptopeza) 

Leptopeza disparilis Melander 1902 (Leptopeza) 

Leptopeza ruficollis Meigen 1820 (Ocydromia) 

Ocydromia flavipes Meigen 1820 Syn. 

Megagrapha Melander 1928 

Megagrapha exquisita Malloch 1923 (Coloboneura) 

Megagrapha platytarsis Chillcott 1983 (Megagrapha) 

Megagrapha pubescens Loew 1862 (Drapetis) 

Meghyperus Loew 1850 

Meghyperus nitidus Melander 1902 (Meghyperus) 

Meghyperus occidens Coquillett 1895 (Meghyperus) 

Metachela Coquillett 1903 

Metachela albipes Walker 1849 (Hemerodromia) 

Metachela collusor Melander 1902 (Hemerodromia) 

Metachela convexa MacDonald 1989 (Metachela) 

Micrempis Melander 1928 

Micrempis arnasoni Chillcott 1983 (Micrempis) 

Micrempis bomboxynon Chillcott 1983 (Micrempis) 

Micrempis anatolica Chillcott 1983 Syn. 

Micrempis flava Chillcott 1983 (Micrempis) 

Micrempis melina Chillcott 1983 (Micrempis) 

Micrempis millepalmae Chillcott 1983 (Micrempis) 

Micrempis minuta Melander 1902 (Stilpon) 

Micrempis nana Melander 1928 (Micrempis) 

Micrempis obliqua Melander 1928 (Micrempis) 

Micrempis pseudopalmae Teskey 1983 (Micrempis) 

Micrempis richardsi Chillcott 1983 (Micrempis) 

Microphorella Becker 1909 

Microphorella acroptera Melander 1928 (Microphorella) 

Microphorella chiragra Melander 1928 (Microphorella) 

Microphorella longitarsis Melander 1928 (Microphorella) 

Microphorella ornatipes Melander 1928 (Microphorella) 

Microphorella tubifera Melander 1928 (Microphorella) 

Microphorus Macquart 1827 


Schistosoma Becker 1902 Syn. 

Microphorus armipes Melander 1928 (Microphorus) 

Microphorus atratus Coquillett 1900 (Microphorus) 

Microphorus bilineata Melander 1902 (Holoclera) 

Microphorus cirripes Melander 1940 (Microphorus) 

Microphorus discalis Melander 1940 (Microphorus) 

Microphorus drapetoides Walker 1849 (Microphorus) 

Microphorus evisceratus Melander 1940 (Microphorus) 

Microphorus isommatus Melander 1928 (Microphorus) 

Microphorus obscurus Coquillett 1903 (Microphorus) 

Microphorus ravidus Coquillett 1895 (Microphorus) 

Microphorus ravus Melander 1940 (Microphorus) 

Microphorus robustus Melander 1928 (Microphorus) 

Microphorus strigilifer Melander 1940 (Microphorus) 

Microphorus sycophantor Melander 1902 (Holoclera) 

Microphorus tacomae Melander 1940 (Microphorus) 

Microphorus yakimensis Melander 1928 (Microphorus) 

Neoplasta Coquillett 1895 

Neoplasta bifida MacDonald and Turner 1993 (Neoplasta) 

Neoplasta chrysopleura MacDonald and Turner 1993 (Neoplasta) 

Neoplasta concava MacDonald and Turner 1993 (Neoplasta) 

Neoplasta deyrupi MacDonald and Turner 1993 (Neoplasta) 

Neoplasta hansoni MacDonald and Turner 1993 (Neoplasta) 

Neoplasta hebes Melander 1947 (Neoplasta) 

Neoplasta megorchis Melander 1947 (Neoplasta) 

Neoplasta octoterga MacDonald and Turner 1993 (Neoplasta) 

Neoplasta parahebes MacDonald and Turner 1993 (Neoplasta) 

Neoplasta paramegorchis MacDonald and Turner 1993 (Neoplasta) 

Neoplasta scapularis Loew 1862 (Hemerodromia) 

Clinocera maculipes Bigot 1887 Syn. 

Neoplasta alleghani Melander 1947 Syn. 

Neoplasta radialis Melander 1947 Syn. 

Neoplasta scapuliformis MacDonald and Turner 1993 (Neoplasta) 

Niphogenia Melander 1928 

Niphogenia eucera Melander 1928 (Niphogenia) 

Niphogenia turneri Wilder 1981 (Niphogenia) 

Ocydromia Meigen 1820 

Eucinesia Gistl 1848 Syn. 

Ocydromia glabricula Fallen 1816 (Empis) 

Ocydromia peregrinata Walker 1849 Syn. 

Oedalea Meigen 1820 

Oedalea astylata Melander 1928 (Oedalea) 

Oedalea lanceolata Melander 1928 (Oedalea) 

Oedalea testacea Melander 1928 Syn. 

Oedalea ohioensis Melander 1902 (Oedalea) 

Oedalea pruinosa Coquillett 1903 (Oedalea) 

Oreogeton Schiner 1860 

Oreogeton capnopterus Melander 1928 (Oreogeton) 

Oreogeton cymballista Melander 1928 (Oreogeton) 

Oreogeton heterogamus Melander 1928 (Oreogeton) 

Oreogeton mitrephorus Melander 1928 (Oreogeton) 

Oreogeton obscura Loew 1864 (Gloma) 

Oreogeton rufa Loew 1864 (Gloma) 

Oreogeton scopifera Coquillett 1900 (Gloma) 

Oreogeton xanthus Melander 1945 (Oreogeton) 

Oreothalia Melander 1902 


157

158 

Oreothalia chillcotti Wilder 1981 (Oreothalia) 

Oreothalia pelops Melander 1902 (Oreothalia) 

Oreothalia rupestris Vaillant 1960 (Oreothalia) 

Oreothalia sierrensis Wilder 1981 (Oreothalia) 

Oreothalia spinitarsis Wilder 1981 (Oreothalia) 

Parathalassius Mik 1891 

Parathalassius aldrichi Melander 1906 (Parathalassius) 

Parathalassius candidatus Melander 1906 (Parathalassius) 

Parathalassius melanderi Cole 1912 (Parathalassius) 

Philetus Melander 1837 

Philetus memorandus Melander 1928 (Philetus) 

Philetus schizophorus Melander 1928 (Philetus) 

Platypalpus Macquart 1827 

Phoroxypha Rondani 1856 Syn. 

Cleptodromia Corti 1907 Syn. 

Platypalpus achlytarsis Chillcott 1962 (Platypalpus) 

Platypalpus aequalis Loew 1864 (Platypalpus) 

Platypalpus alexippus Walker 1849 (Platypalpus) 

Platypalpus alumnus Melander 1928 (Platypalpus) 

Platypalpus anatolicus Chillcott 1962 (Platypalpus) 

Platypalpus apicalis Loew 1864 (Platypalpus) 

Platypalpus arcticus Melander 1928 (Platypalpus) 

Platypalpus armillatus Melander 1928 (Platypalpus) 

Platypalpus ballistrarius Melander 1928 (Platypalpus) 

Platypalpus ballucatus Melander 1928 (Platypalpus) 

Platypalpus bicornis Melander 1928 (Platypalpus) 

Platypalpus callithrix Melander 1928 (Platypalpus) 

Platypalpus canus Melander 1902 (Platypalpus) 

Platypalpus carectorum Chillcott 1962 (Platypalpus) 

Platypalpus cellarius Melander 1928 (Platypalpus) 

Platypalpus churchillensis Chillcott 1962 (Platypalpus) 

Platypalpus continguus Melander 1928 (Platypalpus) 

Platypalpus coquilletti Melander 1924 (Platypalpus) 

Platypalpus crassifemoris Fitch 1856 (Oscinis) 

Platypalpus debilis Loew 1863 Syn. 

Platypalpus melanocerus Melander 1928 Syn. 

Platypalpus mollis Melander 1928 Syn. 

Platypalpus crepidarius Melander 1928 (Platypalpus) 

Platypalpus cuneipennis Melander 1924 (Platypalpus) 

Platypalpus decolor Melander 1928 (Platypalpus) 

Platypalpus direptor Melander 1928 (Platypalpus) 

Platypalpus discifer Loew 1863 (Platypalpus) 

Platypalpus dissimilipes Melander 1928 (Platypalpus) 

Platypalpus diversipes Coquillett 1900 (Platypalpus) 

Platypalpus enervatus Melander 1928 (Platypalpus) 

Platypalpus fasciventris Melander 1928 (Platypalpus) 

Platypalpus flammifer Melander 1924 (Platypalpus) 

Platypalpus flavirostris Loew 1864 (Platypalpus) 

Platypalpus dilutior Melander 1928 Syn. 

Platypalpus microcerus Melander 1928 Syn. 

Platypalpus vittiger Melander 1928 Syn. 

Platypalpus gesticulor Melander 1928 (Platypalpus) 

Platypalpus glacialis Melander 1928 (Platypalpus) 

Platypalpus gravidus Melander 1902 (Platypalpus) 

Platypalpus harpestylis Chillcott 1962 (Platypalpus) 

Platypalpus harpiger Melander 1924 (Platypalpus) 

Platypalpus hastatus Melander 1902 (Platypalpus) 

Platypalpus hians Melander 1902 (Platypalpus) 

Platypalpus fuscohalteratus Melander 1924 Syn. 

Platypalpus holosericus Melander 1924 (Platypalpus) 

Platypalpus hyaenoides Melander 1928 (Platypalpus) 



Platypalpus impexus Melander 1902 (Platypalpus) 

Platypalpus incurvus Melander 1902 (Platypalpus) 

Platypalpus inferialis Melander 1928 (Platypalpus) 

Platypalpus inops Melander 1902 (Platypalpus) 

Platypalpus aequicornis Melander 1928 Syn. 

Platypalpus juvenis Melander 1928 (Platypalpus) 

Platypalpus lacertosus Melander 1928 (Platypalpus) 

Platypalpus laetabilis Melander 1928 (Platypalpus) 

Platypalpus laetus Loew 1864 (Platypalpus) 

Platypalpus lateralis Loew 1864 (Platypalpus) 

Platypalpus collateralis Melander 1928 Syn. 

Platypalpus luctator Melander 1928 (Platypalpus) 

Platypalpus lupatus Melander 1902 (Platypalpus) 

Platypalpus lyristes Melander 1928 (Platypalpus) 

Platypalpus manni Chillcott 1962 (Platypalpus) 

Platypalpus masoni Chillcott 1962 (Platypalpus) 

Platypalpus melanogaster Melander 1928 (Platypalpus) 

Platypalpus melleus Melander 1928 (Platypalpus) 

Platypalpus mesogrammus Loew 1863 (Platypalpus) 

Platypalpus mimus Melander 1928 (Platypalpus) 

Platypalpus monticola Melander 1902 (Platypalpus) 

Platypalpus murphyi Chillcott 1962 (Platypalpus) 

Platypalpus nitidipleura Melander 1928 (Platypalpus) 

Platypalpus ochricollis Melander 1928 (Platypalpus) 

Platypalpus oculeus Melander 1928 (Platypalpus) 

Platypalpus pachycnemus Loew 1864 (Platypalpus) 

Platypalpus paluster Chillcott 1962 (Platypalpus) 

Platypalpus pectinator Melander 1924 (Platypalpus) 

Platypalpus peneprorsus Chillcott 1962 (Platypalpus) 

Platypalpus pilatus Melander 1928 (Platypalpus) 

Platypalpus pluto Melander 1902 (Platypalpus) 

Platypalpus politellus Melander 1928 (Platypalpus) 

Platypalpus politus Melander 1928 Homo. 

Platypalpus nitens Melander 1928 Syn., 

Platypalpus porrectus Melander 1924 (Platypalpus) 

Platypalpus suffasciatus Melander 1928 Syn. 

Platypalpus postpositus Melander 1928 (Platypalpus) 

Platypalpus prorsus Melander 1928 (Platypalpus) 

Platypalpus pubescens Melander 1928 (Platypalpus) 

Platypalpus pudens Melander 1928 (Platypalpus) 

Platypalpus puerinus Melander 1928 (Platypalpus) 

Platypalpus pulverulentus Melander 1928 (Platypalpus) 

Platypalpus recurvus Melander 1928 (Platypalpus) 

Platypalpus richardsi Chillcott 1962 (Platypalpus) 

Platypalpus rubefactus Melander 1928 (Platypalpus) 

Platypalpus rufiventris Melander 1902 (Platypalpus) 

Platypalpus satyriacus Melander 1928 (Platypalpus) 

Platypalpus sericatus Melander 1928 (Platypalpus) 

Platypalpus simplicipes Melander 1928 (Platypalpus) 

Platypalpus soccatus Melander 1928 (Platypalpus) 

Platypalpus spinosus Melander 1928 (Platypalpus) 

Platypalpus splendens Melander 1928 (Platypalpus) 

Elaphropeza montana Melander 1902 Homo. 

Platypalpus sutor Melander 1924 (Platypalpus) 

Platypalpus tachistiformis Melander 1928 (Platypalpus) 

Platypalpus talaris Melander 1928 (Platypalpus) 

Platypalpus tenax Melander 1928 (Platypalpus) 

Platypalpus tenellus Melander 1902 (Platypalpus) 

Platypalpus tenuis Melander 1928 (Platypalpus) 

Platypalpus tersus Coquillett 1895 (Platypalpus) 

Platypalpus trivialis Loew 1864 (Platypalpus) 

Platypalpus truncatus Chillcott 1962 (Platypalpus) 

Platypalpus valgus Melander 1928 (Platypalpus) 

Platypalpus velox Melander 1928 (Platypalpus) 

Platypalpus venaticus Melander 1928 (Platypalpus) 

Platypalpus verpus Melander 1928 (Platypalpus) 

Platypalpus versipes Melander 1928 (Platypalpus) 

Platypalpus versutus Melander 1924 (Platypalpus)

Platypalpus vicarius Walker 1857 (Platypalpus) 

Platypalpus vierecki Melander 1902 (Platypalpus) 

Platypalpus vittatus Melander 1928 (Platypalpus) 

Platypalpus perimerus Melander 1928 Syn. 

Platypalpus vulnificus Melander 1928 (Platypalpus) 

Platypalpus xanthochiton Melander 1928 (Platypalpus) 

Platypalpus xanthopodus Melander 1928 (Platypalpus) 

Platypalpus gilvipes Coquillett 1900 Homo. 

Proclinopyga Melander 1928 

Proclinopyga amplectens Melander 1928 (Proclinopyga) 

Proclinopyga exporrecta Melander 1928 (Proclinopyga) 

Proclinopyga fistulator Melander 1928 (Proclinopyga) 

Proclinopyga monogramma Melander 1928 (Proclinopyga) 

Proclinopyga flavicoxa Melander 1928 Syn. 

Proclinopyga solivaga Melander 1928 (Proclinopyga) 

Ragas Walker 1837 

Rhagas Agassiz 1846 Emend. 

Ragas primigenia Melander 1945 (Ragas) 

Rhamphomyia Meigen 1822 

Dionnaea Meigen 1800 Unav. 

Holoclera Schiner 1860 Syn. 

Megacyttarus Bigot 1880 Syn. 

Neocota Coquillett 1895 Syn. 

Pararhamphomyia Frey 1922 Syn. 

Dasyrhamphomyia Frey 1922 Syn. 

Choreodromia Frey 1922 Syn. 

Lundstroemiella Frey 1922 Syn. 

Aclonempis Collin 1926 Syn. 

Amydroneura Collin 1926 Syn. 

Ctenempis Frey 1935 Syn. 

Eorhamphomyia Frey 1950 Syn. 

Alpinoyia Frey 1950 Syn. 

Collinaria Frey 1950 Syn. 

Orientomyia Saigusa 1963 Syn. 

Calorhamphomyia Saigusa 1963 Syn. 

Rhamphomyia adversa Coquillett 1900 (Rhamphomyia) 

Rhamphomyia agasicles Walker 1849 (Rhamphomyia) 

Rhamphomyia albata Coquillett 1902 (Rhamphomyia) 

Rhamphomyia albopilosa Coquillett 1900 (Rhamphomyia) 

Rhamphomyia ambocnema Chillcott 1959 (Rhamphomyia) 

Rhamphomyia americana Wiedemann 1830 (Rhamphomyia) 

Rhamphomyia amplicella Coquillett 1895 (Rhamphomyia) 

Rhamphomyia amplipedis Coquillett 1895 (Rhamphomyia) 

Rhamphomyia ethellia Curran 1933 Syn. 

Rhamphomyia anaxo Walker 1849 (Rhamphomyia) 

Rhamphomyia angustipennis Loew 1861 (Rhamphomyia) 

Rhamphomyia anthracodes Coquillett 1900 (Rhamphomyia) 

Rhamphomyia aperta Loew 1862 (Rhamphomyia) 

Rhamphomyia arctotibia Chillcott 1959 (Rhamphomyia) 

Rhamphomyia arcuata Coquillett 1895 (Rhamphomyia) 

Rhamphomyia atrata Coquillett 1900 (Rhamphomyia) 

Rhamphomyia avida Coquillett 1895 (Rhamphomyia) 

Rhamphomyia barypoda Coquillett 1900 (Rhamphomyia) 

Rhamphomyia basalis Loew 1864 (Rhamphomyia) 

Rhamphomyia bifilata Coquillett 1895 (Rhamphomyia) 

Rhamphomyia bigelowi Walley 1928 (Rhamphomyia) 

Rhamphomyia birdi Curran 1929 (Rhamphomyia) 

Rhamphomyia borealis Fabricius 1780 (Empis) 

Rhamphomyia brevis Loew 1861 (Rhamphomyia) 

Rhamphomyia corvina Loew 1861 Syn. 

Rhamphomyia cervina Loew 1861 Missp. 

Rhamphomyia californica Coquillett 1895 (Rhamphomyia) 


Rhamphomyia calvimontis Cockerell 1916 (Rhamphomyia) 

Rhamphomyia candicans Loew 1864 (Rhamphomyia) 

Rhamphomyia ciliata Coquillett 1895 (Rhamphomyia) 

Rhamphomyia cilipes Say 1823 (Empis) 

Rhamphomyia cinefacta Coquillett 1900 (Rhamphomyia) 

Rhamphomyia cineracea Coquillett 1900 (Rhamphomyia) 

Rhamphomyia clauda Coquillett 1901 (Rhamphomyia) 

Rhamphomyia clavator Coquillett 1901 (Rhamphomyia) 

Rhamphomyia macrura Coquillett 1900 Homo. 

Rhamphomyia clavigera Loew 1861 (Rhamphomyia) 

Rhamphomyia colorata Coquillett 1895 (Rhamphomyia) 

Rhamphomyia compta Coquillett 1895 (Rhamphomyia) 

Rhamphomyia conjuncta Loew 1861 (Rhamphomyia) 

Rhamphomyia conservativa Malloch 1919 (Rhamphomyia) 

Rhamphomyia cophas Walker 1849 (Rhamphomyia) 

Rhamphomyia currani Steyskal 1964 (Rhamphomyia) 

Neocota rufipes Curran 1933 Homo. 

Rhamphomyia curvipes Coquillett 1904 (Rhamphomyia) 

Rhamphomyia dana Walker 1849 (Rhamphomyia) 

Rhamphomyia daria Walker 1849 (Rhamphomyia) 

Rhamphomyia dasypoda Steyskal 1964 (Rhamphomyia) 

Rhamphomyia debilis Loew 1861 (Rhamphomyia) 

Rhamphomyia dimidiata Loew 1861 (Rhamphomyia) 

Rhamphomyia disconcerta Curran 1930 (Rhamphomyia) 

Rhamphomyia disparilis Coquillett 1900 (Rhamphomyia) 

Rhamphomyia diversa Coquillett 1901 (Rhamphomyia) 

Rhamphomyia duplicis Coquillett 1895 (Rhamphomyia) 

Rhamphomyia ecetra Walker 1849 (Rhamphomyia) 

Rhamphomyia effera Coquillett 1895 (Rhamphomyia) 

Rhamphomyia erinacioides Malloch 1919 (Rhamphomyia) 

Rhamphomyia exigua Loew 1862 (Rhamphomyia) 

Rhamphomyia expulsa Walker 1857 (Rhamphomyia) 

Rhamphomyia falcipedia Chillcott 1959 (Rhamphomyia) 

Rhamphomyia ficana Walker 1849 (Rhamphomyia) 

Rhamphomyia filicauda Henriksen and Lundbeck 1917 

(Rhamphomyia) 

Rhamphomyia fimbriata Coquillett 1895 (Rhamphomyia) 

Rhamphomyia flavirostris Walker 1849 (Rhamphomyia) 

Rhamphomyia flexuosa Coquillett 1895 (Rhamphomyia) 

Rhamphomyia formosula Melander 1965 (Rhamphomyia) 

Rhamphomyia pulchra Loew 1861 Homo. 

Rhamphomyia frontalis Loew 1862 (Rhamphomyia) 

Rhamphomyia fumosa Loew 1861 (Rhamphomyia) 

Rhamphomyia gilvipes Loew 1861 (Rhamphomyia) 

Rhamphomyia gilvipilosa Coquillett 1895 (Rhamphomyia) 

Rhamphomyia glabra Loew 1861 (Rhamphomyia) 

Rhamphomyia glauca Coquillett 1900 (Rhamphomyia) 

Rhamphomyia gracilis Loew 1861 (Rhamphomyia) 

Rhamphomyia bipunctata Curran 1930 Syn. 

Rhamphomyia herschelli Malloch 1919 (Rhamphomyia) 

Rhamphomyia hirticula Collin 1937 (Rhamphomyia) 

Rhamphomyia hirtipes Loew 1864 (Rhamphomyia) 

Rhamphomyia hirtula Zetterstedt 1842 (Rhamphomyia) 

Rhamphomyia hoelsi Frey 1950 (Rhamphomyia) 

Rhamphomyia hoeli Frey 1955 Emend. 

Rhamphomyia hovgaardii Holmgren 1880 (Rhamphomyia) 

Rhamphomyia impedita Loew 1862 (Rhamphomyia) 

Rhamphomyia incompleta Loew 1863 (Rhamphomyia) 

Rhamphomyia insecta Coquillett 1895 (Rhamphomyia) 

Rhamphomyia intonsa Steyskal 1964 (Rhamphomyia) 

Rhamphomyia irregularis Loew 1864 (Rhamphomyia) 

Rhamphomyia jubata Chillcott 1959 (Rhamphomyia) 

Rhamphomyia laevigata Loew 1861 (Rhamphomyia) 

Rhamphomyia leucoptera Loew 1861 (Rhamphomyia) 

Rhamphomyia limata Coquillett 1900 (Rhamphomyia) 

Rhamphomyia limbata Loew 1861 (Rhamphomyia) 

Megacyttarus argentea Bigot 1880 Syn. 

Rhamphomyia liturata Loew 1861 (Rhamphomyia) 


159

160 

Rhamphomyia longicauda Loew 1861 (Rhamphomyia) 

Rhamphomyia longicornis Loew 1861 (Rhamphomyia) 

Rhamphomyia longipennis Loew 1861 (Rhamphomyia) 

Rhamphomyia loripedis Coquillett 1895 (Rhamphomyia) 

Rhamphomyia luctifera Loew 1861 (Rhamphomyia) 

Rhamphomyia luctuosa Loew 1872 (Rhamphomyia) 

Rhamphomyia lugens Loew 1862 Homo. 

Rhamphomyia luteiventer Curran 1929 (Rhamphomyia) 

Rhamphomyia luteiventris Loew 1864 (Rhamphomyia) 

Rhamphomyia macilenta Loew 1864 (Rhamphomyia) 

Rhamphomyia mallos Walker 1849 (Rhamphomyia) 

Rhamphomyia manca Coquillett 1895 (Rhamphomyia) 

Rhamphomyia minytus Walker 1849 (Rhamphomyia) 

Rhamphomyia minutus Johnson 1910 Missp. 

Rhamphomyia mutabilis Loew 1862 (Rhamphomyia) 

Rhamphomyia nana Loew 1861 (Rhamphomyia) 

Rhamphomyia nasoni Coquillett 1895 (Rhamphomyia) 

Rhamphomyia nigricans Loew 1864 (Rhamphomyia) 

Rhamphomyia nigrita Zetterstedt 1838 (Rhamphomyia) 

Rhamphomyia nitidivittata Macquart 1846 (Rhamphomyia) 

Rhamphomyia novecarolina Beutenmüller 1913 (Rhamphomyia) 

Rhamphomyia opacithorax Malloch 1923 (Rhamphomyia) 

Rhamphomyia otiosa Coquillett 1895 (Rhamphomyia) 

Rhamphomyia pachymera Bigot 1887 (Rhamphomyia) 

Rhamphomyia parva Coquillett 1895 (Rhamphomyia) 

Rhamphomyia pectinata Loew 1861 (Rhamphomyia) 

Rhamphomyia pectoris Coquillett 1895 (Rhamphomyia) 

Rhamphomyia phemius Walker 1849 (Rhamphomyia) 

Rhamphomyia piligeronis Coquillett 1895 (Rhamphomyia) 

Rhamphomyia polita Loew 1862 (Rhamphomyia) 

Rhamphomyia prava Chillcott 1959 (Rhamphomyia) 

Rhamphomyia priapulus Loew 1861 (Rhamphomyia) 

Rhamphomyia pulla Loew 1861 (Rhamphomyia) 

Rhamphomyia pusio Loew 1861 (Rhamphomyia) 

Rhamphomyia quinquelineata Say 1823 (Empis) 

Rhamphomyia rava Loew 1862 (Rhamphomyia) 

Rhamphomyia morrisoni Bigot 1887 Syn. 

Rhamphomyia ravida Coquillett 1895 (Rhamphomyia) 

Rhamphomyia rufirostra Say 1829 (Rhamphomyia) 

Rhamphomyia rustica Loew 1864 (Rhamphomyia) 

Rhamphomyia scaurissima Wheeler 1896 (Rhamphomyia) 

Rhamphomyia scolopacea Say 1823 (Empis) 

Rhamphomyia scutellaris Coquillett 1895 (Rhamphomyia) 

Rhamphomyia geniculata Bigot 1887 Homo. 

Rhamphomyia sellata Loew 1861 (Rhamphomyia) 

Rhamphomyia setosa Coquillett 1895 (Rhamphomyia) 

Rhamphomyia similata Malloch 1919 (Rhamphomyia) 

Rhamphomyia soccata Loew 1861 (Rhamphomyia) 

Rhamphomyia sociabilis Williston 1893 (Empis) 

Rhamphomyia abdita Coquillett 1895 Syn. 

Rhamphomyia soleata Melander 1965 (Rhamphomyia) 

Rhamphomyia argentea Curran 1930 Homo. 

Rhamphomyia sordida Loew 1861 (Rhamphomyia) 

Rhamphomyia crassinervis Loew 1861 Syn. 

Rhamphomyia stylata Coquillett 1895 (Rhamphomyia) 

Rhamphomyia sudigeronis Coquillett 1895 (Rhamphomyia) 

Rhamphomyia nigrita Bigot 1887 Homo. 

Rhamphomyia tersa Coquillett 1895 (Rhamphomyia) 

Rhamphomyia testacea Loew 1862 (Rhamphomyia) 

Rhamphomyia tristis Walker 1857 (Rhamphomyia) 

Rhamphomyia umbilicata Loew 1861 (Rhamphomyia) 

Rhamphomyia ungulata Loew 1861 Syn. 

Rhamphomyia umbrosa Loew 1864 (Rhamphomyia) 

Rhamphomyia unimaculata Loew 1862 (Rhamphomyia) 

Rhamphomyia ursinella Melander 1928 (Rhamphomyia) 

Rhamphomyia ursina Malloch 1919 Homo. 

Rhamphomyia valga Coquillett 1895 (Rhamphomyia) 

Rhamphomyia vara Loew 1861 (Rhamphomyia) 



Rhamphomyia versicolor Chillcott 1959 (Rhamphomyia) 

Rhamphomyia villipes Coquillett 1900 (Rhamphomyia) 

Rhamphomyia virgata Coquillett 1895 (Rhamphomyia) 

Rhamphomyia vittata Loew 1862 (Rhamphomyia) 

Rhamphomyia weedii Coquillett 1895 (Neocota) 

Roederiodes Coquillett 1901 

Roederioides Curran 1934 Missp. 

Roederiodes distincta Chillcott 1961 (Roederiodes) 

Roederiodes juncta Coquillett 1901 (Roederiodes) 

Roederiodes recurvata Chillcott 1961 (Roederiodes) 

Roederiodes vockerothi Chillcott 1961 (Roederiodes) 

Roederiodes wirthi Chillcott 1961 (Roederiodes) 

Roederiodes retroversa Chillcott 1961 Syn. 

Sabroskyella Wilder 1984 

Sabroskyella rancheria Wilder 1984 (Sabroskyella) 

Stilpon Loew 1859 

Agatachys Meigen 1830 Unav. 

Tetraneurella Dahl 1909 Syn. 

Pseudostilpon Seguy 1950 Syn. 

Stilpon curvipes Melander 1928 (Stilpon) 

Stilpon pauciseta Melander 1928 (Stilpon) 

Stilpon pectiniger Melander 1902 (Stilpon) 

Stilpon pleuritica Melander 1928 (Stilpon) 

Stilpon spinipes Melander 1928 (Stilpon) 

Stilpon varipes Loew 1862 (Stilpon) 

Symballophthalmus Becker 1889 

Macroptera Becker 1889 Homo. 

Symballophthalmus masoni Chillcott 1958 (Symballophthalmus) 

Syndyas Loew 1857 

Syndyas austropolita Teskey and Chillcott 1977 (Syndyas) 

Syndyas dorsalis Loew 1861 (Syndyas) 

Syndyas lustricola Teskey and Chillcott 1977 (Syndyas) 

Syndyas merbleuensis Teskey and Chillcott 1977 (Syndyas) 

Syndyas pleuripolita Teskey and Chillcott 1977 (Syndyas) 

Syndyas polita Loew 1861 (Syndyas) 

Syneches Walker 1852 

Pterospilus Rondani 1856 Syn. 

Syneches albonotatus Loew 1862 (Syneches) 

Syneches ater Melander 1928 (Syneches) 

Syneches debilis Coquillett 1895 (Syneches) 

Syneches frosti Wilder 1974 (Syneches) 

Syneches hyalinus Coquillett 1895 (Syneches)

Syneches longipennis Melander 1902 (Syneches) 

Syneches pusillus Loew 1861 (Syneches) 

Syneches rufus Loew 1861 (Syneches) 

Syneches simplex Walker 1852 (Syneches) 

Syneches punctipennis Wulp 1867 Syn. 

Syneches thoracicus Say 1823 (Hybos) 

Gloma phthia Walker 1849 Syn. 

Syneches testaceus Melander 1928 Syn. 

Syneches vittatus Wilder 1974 (Syneches) 

Tachydromia Meigen 1803 

Sicus Latreille 1796 Homo. 

Coryneta Meigen 1800 Unav. 

Sicodus Rafinesque 1815 Syn. 

Danistes Gistl 1848 Syn. 

Phoneutisca Loew 1863 Syn. 

Tachista Loew 1864 Syn. 

Tachysta Loew 1864 Missp. 

Tachydromia bimaculata Loew 1863 (Phoneutisca) 

Tachydromia chelana Melander 1928 (Tachydromia) 

Tachydromia diversipes Melander 1910 (Tachydromia) 

Tachydromia enecator Melander 1902 (Tachydromia) 

Tachydromia harti Malloch 1919 (Tachydromia) 

Tachydromia hirtipes Melander 1928 (Tachydromia) 

Tachydromia maculipennis Walker 1849 (Tachydromia) 

Tachydromia pusilla Loew 1864 Syn. 

Tachydromia monaca Melander 1928 (Tachydromia) 

Tachydromia phengites Melander 1928 (Tachydromia) 

Tachydromia pseliophora Melander 1928 (Tachydromia) 

Tachydromia schwarzii Coquillett 1895 (Tachydromia) 

Tachydromia tacoma Melander 1928 (Tachydromia) 

Tachydromia varipennis Coquillett 1903 (Tachydromia) 

Tachyempis Melander 1928 

Tachyempis agens Melander 1910 (Tachydromia) 

Tachyempis calva Melander 1910 (Tachydromia) 

Tachyempis cinerea Melander 1928 (Tachyempis) 

Tachyempis longipennis Melander 1958 (Tachyempis) 

Tachyempis nervosa Melander 1928 (Tachyempis) 

Tachyempis universalis Melander 1910 (Tachydromia) 

Tachypeza Meigen 1830 

Cormodromia Zetterstedt 1838 Syn. 

Tachypeza annularis Melander 1928 (Tachypeza) 

Tachypeza binotata Melander 1928 (Tachypeza) 

Tachypeza brachialis Melander 1902 (Tachydromia) 

Tachypeza clavipes Loew 1864 (Tachypeza) 

Tachypeza corticalis Melander 1902 (Tachydromia) 

Tachypeza discifera Melander 1928 (Tachypeza) 

Tachypeza distans Melander 1928 (Tachypeza) 

Tachypeza dolorosa Melander 1928 (Tachypeza) 

Tachypeza excisa Melander 1928 (Tachypeza) 

Tachypeza fenestratus Say 1823 (Sicus) 

Tachydromia similis Walker 1849 Syn. 

Tachypeza rapax Loew 1864 Syn. 

Tachypeza humeralis Melander 1928 (Tachypeza) 

Tachypeza inusta Melander 1902 (Tachydromia) 

Tachypeza portaecola Walker 1849 (Tachydromia) 

Tachypeza postica Walker 1857 (Tachydromia) 

Tachypeza pruinosa Coquillett 1903 (Tachypeza) 

Tachypeza rostrata Loew 1864 (Tachypeza) 

Tachypeza vittipennis Walker 1857 (Tachydromia) 

Tachypeza winthemi Zetterstedt 1838 (Tachypeza) 


Toreus Melander 1906 

Toreus neomexicana Melander 1902 (Empis) 

Trichina Meigen 1830 

Pipistrellus Gistl 1848 Syn. 

Oedaleopsis Tuomikoski 1959 Syn. 

Trichina atripes Melander 1902 (Euthyneura) 

Trichina basalis Melander 1928 (Trichina) 

Trichina clavipes Meigen 1830 (Trichina) 

Trichina flavipes Meigen 1830 (Trichina) 

Trichina nitida Melander 1928 (Trichina) 

Trichina nura Melander 1902 (Euthyneura) 

Trichina pullata Melander 1928 (Trichina) 

Trichoclinocera Collin 1941 

Seguyella Vaillant 1960 Syn. 

Acanthoclinocera Saigusa 1965 Syn. 

Trichoclinocera agilis Sinclair 1994 (Trichoclinocera) 

Trichoclinocera cascadensis Sinclair 1994 (Trichoclinocera) 

Trichoclinocera comata Melander 1928 (Wiedemannia) 

Trichoclinocera ctenistes Melander 1928 (Wiedemannia) 

Trichoclinocera cummingi Sinclair 1994 (Trichoclinocera) 

Trichoclinocera dasycoxa Sinclair 1994 (Trichoclinocera) 

Trichoclinocera dolicheretma Melander 1902 (Clinocera) 

Clinocera brunnipennis Melander 1928 Syn. 

Trichoclinocera falcata Sinclair 1994 (Trichoclinocera) 

Trichoclinocera fumosa Vaillant 1960 (Wiedemannia) 

Trichoclinocera hamifera Melander 1928 (Wiedemannia) 

Wiedemannia saltans Vaillant 1960 Syn. 

Trichoclinocera lapponica Ringdahl 1933 (Atalanta) 

Trichoclinocera longipes Walker 1849 (Heleodromia) 

Trichoclinocera minor Melander 1928 (Wiedemannia) 

Trichoclinocera ozarkensis Sinclair 1994 (Trichoclinocera) 

Trichoclinocera pectinifemur Sinclair 1994 (Trichoclinocera) 

Trichoclinocera rupestris Sinclair 1994 (Trichoclinocera) 

Wiedemannia Zetterstedt 1838 

Agyrtomyia Gistl 1848 Syn. 

Macellaria Gistl 1848 Syn. 

Philolutra Mik 1881 Syn. 

Chamaedipsia Mik 1881 Syn. 

Roederia Mik 1881 Syn. 

Eucelidia Mik 1881 Syn. 

Roederella Engel 1918 Syn. 

Camelopis Engel 1918 Syn. 

Pseudowiedemannia Engel 1918 Syn. 

Clinocerella Engel 1918 Syn. 

Wiedemannia gubernans Melander 1928 (Wiedemannia) 

Wiedemannia lepida Melander 1902 (Clinocera) 

Wiedemannia simplex Loew 1862 (Clinocera) 

Zanclotus Wilder 1984 

Zanclotus dioktes Wilder 1984 (Zanclotus) 


161

162 

EPHYDRIDAE 

Allotrichoma Becker 1896 

Epiphasis Becker 1907 Syn. 

Pseudohecamede Hendel 1936 Syn. 

Allotrichoma abdominalis Williston 1896 (Hecamede) 

Allotrichoma longirostre Hendel 1930 Syn. 

Allotrichoma adustum Mathis 1991 (Allotrichoma) 

Allotrichoma atrilabris Cresson 1926 (Allotrichoma) 

Allotrichoma facialis Hendel 1936 (Pseudohecamede) 

Allotrichoma lacteum Cresson 1926 (Allotrichoma) 

Allotrichoma lasiocercum Cresson 1926 (Allotrichoma) 

Allotrichoma lateralis Loew 1860 (Hecamede) 

Allotrichoma oceanum Becker 1926 Syn. 

Allotrichoma impudicum Duda 1942 Syn. 

Allotrichoma valkanovi Beschovski 1966 Syn. 

Allotrichoma simplex Loew 1861 (Discocerina) 

Allotrichoma slossonae Cresson 1942 (Allotrichoma) 

Allotrichoma yosemite Cresson 1926 (Allotrichoma) 

Asmeringa Becker 1903 

Asmeringa lindsleyi Sturtevant and Wheeler 1954 (Asmeringa) 

Athyroglossa Loew 1860 

Ochtheroidea Williston 1896 Syn. 

Parathyroglossa Hendel 1931 Syn. 

Standitrichoma Duda 1942 Syn. 

DIPTERA: EPHYDRIDAE 


Athyroglossa cressoni Wirth 1980 (Athyroglossa) 

Athyroglossa dinorata Mathis and Zatwarnicki 1990 (Athyroglossa) 

Athyroglossa glabra Meigen 1830 (Notiphila) 

Clasiopa brevipectinata Becker 1897 Syn. 

Athyroglossa glaphyropus Loew 1878 (Athyroglossa) 

Athyroglossa granulosa Cresson 1922 (Ochtheroidea) 

Athyroglossa laevis Cresson 1918 (Ochtheroidea) 

Athyroglossa melanderi Cresson 1922 (Ochtheroidea) 

Athyroglossa transversa Sturtevant and Wheeler 1954 

(Athyroglossa) 

Atissa Haliday 1839 

Atissa Haliday 1837 Unav. 

Parephydra Coquillett 1902 Syn. 

Pelignellus Sturtevant and Wheeler 1954 Syn. 

Atissa atlantica Cresson 1926 (Atissa) 

Atissa humilis Coquillett 1902 (Parephydra) 

Atissa litoralis Cole 1912 (Allotrichoma) 

Atissa luteipes Cresson 1944 (Atissa) 

Atissa subnudus Sturtevant and Wheeler 1954 (Pelignellus) 

Axysta Haliday 1839 

Microlytogaster Clausen 1983 Syn. 

Axysta americana Clausen 1983 (Axysta) 

Axysta bradleyi Cresson 1930 (Axysta) 

Axysta extera Cresson 1942 (Lytogaster) 

Brachydeutera Loew 1862 

Brachydeutera argentata Walker 1852 (Notiphila) 

Brachydeutera dimidiata Loew 1862 Syn. 

Brachydeutera longipes Hendel 1913 (Brachydeutera) 

Brachydeutera neotropica Wirth 1964 (Brachydeutera) 

Brachydeutera sturtevanti Wirth 1964 (Brachydeutera) 

Callinapaea Sturtevant and Wheeler 1954 

Callinapaea aldrichi Sturtevant and Wheeler 1954 (Napaea) 

Callinapaea laurentiana Wirth 1965 (Parydra) 

Calocoenia Mathis 1975 

Leptocoenia Mathis 1975 Syn. 

Calocoenia paurosoma Sturtevant and Wheeler 1954 (Coenia) 

Coenia borealis Dahl 1975 Syn. 

Calocoenia platypelta Cresson 1935 (Paracoenia) 

Ceropsilopa Cresson 1917 

Pelex Cresson 1925 Syn. 

Batula Cresson 1940 Syn. 

Ceropsilopa adjuncta Cresson 1925 (Ceropsilopa) 

Ceropsilopa coquilletti Cresson 1922 (Ceropsilopa) 

Ceropsilopa costalis Wirth 1956 (Ceropsilopa) 

Ceropsilopa dispar Cresson 1922 (Ceropsilopa) 

Ceropsilopa mellipes Coquillett 1900 (Psilopa) 

Ceropsilopa nasuta Cresson 1917 (Ceropsilopa) 

Ceropsilopa staffordi Cresson 1925 (Ceropsilopa) 

Cirrula Cresson 1915 

Pogonephydra Hendel 1917 Syn. 

Hydropyrus Cresson 1934 Syn. 

Cirrula austrina Coquillett 1900 (Ephydra) 

Coenia virida Hine 1904 Syn. 

Cirrula currani Wirth 1971 (Ephydra) 

Ephydra salina Curran 1931 Homo. 

Cirrula gigantea Cresson 1915 (Cirrula) 

Pogonephydra chalybea Hendel 1917 Syn. 

Cirrula hians Say 1830 (Ephydra) 

Ephydra crassimana Loew 1866 Syn. 

Ephydra californica Packard 1871 Syn. 

Ephydra tarsata Williston 1893 Syn. 

Clanoneurum Becker 1903 

Cyclocephala Strobl 1902 Homo. 

Cyclocephalomyia Hendel 1907 Syn. 

Discomyzoides Becker 1926 Syn. 

Heringium Enderlein 1934 Syn. 

Clanoneurum americanum Cresson 1940 (Clanoneurum) 

Clasiopella Hendel 1914 

Clasiopella uncinata Hendel 1914 (Clasiopella) 

Psilopa giloipes Becker 1924 Syn. 

Coenia Robineau-Desvoidy 1830 

Coenia alpina Mathis 1975 (Coenia) 

Coenia curvicauda Meigen 1830 (Ephydra) 

Cressonomyia Arnaud 1958 

Plagiops Cresson 1918 Homo. 

Plagiopsis Cresson 1934 Homo. 

Cressonomyia aciculata Loew 1862 (Psilopa) 

Ephygrobia metallica Schiner 1868 Syn. 

Cressonomyia aeneonigra Loew 1878 (Psilopa) 

Psilopa fulvipennis Hine 1904 Syn. 

Cressonomyia hinei Cresson 1922 (Plagiops)

Cressonomyia skinneri Cresson 1922 (Psilopa) 

Diclasiopa Hendel 1917 

Diclasiopa frontalis Cresson 1933 (Diclasiopa) 

Diclasiopa lacteipennis Loew 1862 (Discocerina) 

Hecamede xanthocera Loew 1869 Syn. 

Hecamede xanthocera Loew 1869 Syn. 

Hecamede aurella Strobl 1893 Syn. 

Dimecoenia Cresson 1916 

Dimecoenia fuscifemur Steyskal 1970 (Dimecoenia) 

Dimecoenia spinosa Loew 1864 (Caenia [sic]) 

Diphuia Cresson 1944 

Diphuia nasalis Wirth 1956 (Diphuia) 

Diphuia nitida Sturtevant and Wheeler 1954 (Diphuia) 

Discocerina Macquart 1835 

Dioscocerina Macquart 1835 Missp. 

Clasiopa Stenhammar 1844 Syn. 

Lamproclasiopa Hendel 1933 Syn. 

Basila Cresson 1942 Syn. 

Discocerina brunneonitens Cresson 1940 (Discocerina) 

Discocerina flavipes Cresson 1941 (Discocerina) 

Discocerina nadineae Cresson 1925 (Ditrichophora) 

Discocerina obscura Williston 1896 (Discocerina) 

Discocerina obscurella Fallen 1813 (Notiphila) 

Drosophila cinerella Fallen 1823 Syn. 

Diastata luctuosa Meigen 1830 Syn. 

Notiphila nigrina Meigen 1830 Syn. 

Notiphila pusilla Meigen 1830 Syn. 

Notiphila tristis Meigen 1830 Syn. 

Psilopa pallidula Stenhammar 1844 Syn. 

Discocerina parva Loew 1862 Syn. 

Discocerina nigriventris Cresson 1916 Syn. 

Hippelates porteri Brethes 1925 Syn. 

Discocerina trochanterata Cresson 1940 (Discocerina) 

Discocerina turgidula Cresson 1940 (Discocerina) 

Discomyza Meigen 1830 

Discomyza maculipennis Wiedemann 1824 (Notiphila) 

Discomyza obscurata Walker 1860 Syn. 

Discomyza balioptera Loew 1862 Syn. 

Discomyza pelagica Frauenfeld 1867 Syn. 

Discomyza amabilis Kertesz 1901 Syn. 

Discomyza u-signata Cresson 1926 (Discomyza) 

Ditrichophora Cresson 1924 

Standiscocera Duda 1942 Syn. 

Ditrichophora atrata Cresson 1940 (Ditrichophora) 

Ditrichophora exigua Cresson 1924 (Ditrichophora) 

Ditrichophora lenis Cresson 1940 (Ditrichophora) 

Ditrichophora lugubris Cresson 1940 (Ditrichophora) 

Ditrichophora occidentalis Cresson 1942 (Ditrichophora) 

Ditrichophora simiaceps Cresson 1940 (Ditrichophora) 

Ditrichophora valens Cresson 1942 (Ditrichophora) 

Ephydra Fallen 1810 

Halephydra Wirth 1971 Syn. 

Ephydra auripes Aldrich 1912 (Ephydra) 

Ephydra brevis Walker 1858 (Ephydra) 


Ephydra bruesi Cresson 1934 (Ephydra) 

Ephydra goedeni Wirth 1971 (Ephydra) 

Ephydra gracilis Packard 1871 (Ephydra) 

Ephydra cinerea Jones 1906 Syn. 

Ephydra millbrae Jones 1906 (Ephydra) 

Ephydra nana Walker 1858 (Ephydra) 

Ephydra niveiceps Cresson 1916 (Ephydra) 

Ephydra obscuripes Loew 1866 (Ephydra) 

Ephydra oscitans Walker 1858 (Ephydra) 

Ephydra packardi Wirth 1971 (Ephydra) 

Ephydra halophila Packard 1869 Homo. 

Ephydra pectinulata Cresson 1916 (Ephydra) 

Ephydra riparia Fallen 1813 (Ephydra) 

Ephydra albula Meigen 1830 Syn. 

Ephydra salina Heyden 1843 Syn. 

Coenia halophila Heyden 1844 Syn. 

Ephydra salinae Zetterstedt 1846 Syn. 

Ephydra lata Walker 1858 Syn. 

Ephydra strenzkei Giordani Soika 1960 Syn. 

Ephydra subopaca Loew 1864 (Ephydra) 

Ephydra thermophila Cresson 1934 (Ephydra) 

Ephydra usingeri Wirth 1976 (Ephydra) 

Eutaenionotum Oldenberg 1923 

Eutaenionotum guttipennis Stenhammar 1844 (Notiphila) 

Ephydra octonotata Walker 1849 Syn. 

Notiphila producta Walker 1849 Syn. 

Napaea yukonensis Cresson 1949 Syn. 

Gastrops Williston 1897 

Ventrops Grossbeck 1912 Unav. 

Gastrops nebulosus Coquillett 1900 (Gastrops) 

Gastrops niger Williston 1897 (Gastrops) 

Glenanthe Haliday 1839 

Glenanthe fascipennis Sturtevant and Wheeler 1954 (Glenanthe) 

Glenanthe interior Chillcott 1964 (Glenanthe) 

Glenanthe litorea Cresson 1925 (Glenanthe) 

Glenanthe ruetzleri Mathis 1995 (Glenanthe) 

Glenanthe salina Mathis 1995 (Glenanthe) 

Guttipsilopa Wirth 1956 

Neopsilopa Mathis and Wirth 1977 Syn. 

Guttipsilopa umbrosa Loew 1862 (Psilopa) 

Rhysophora unbrosa Cresson 1946 Missp. 

Guttipsilopa wirthi Mathis and Freidberg 1980 (Neopsilopa) 

Gymnoclasiopa Cresson 1930 

Gymnoclasiopa argyrostoma Cresson 1916 (Discocerina) 

Discocerina aliena Cole and Lovett 1921 Unav. 

Discocerina aliena Cresson 1922 Syn. 

Gymnoclasiopa cana Cresson 1940 (Ditrichophora) 

Gymnoclasiopa canifrons Cresson 1926 (Ditrichophora) 

Gymnoclasiopa montana Cresson 1942 (Ditrichophora) 

Gymnoclasiopa parilis Cresson 1924 (Ditrichophora) 

Gymnoclasiopa pulchella Meigen 1830 (Notiphila) 

Clasiopa fulgida Becker 1896 Syn. 

Gymnoclasiopa subnubila Cresson 1940 (Ditrichophora) 

Gymnoclasiopa tacoma Cresson 1924 (Ditrichophora) 

Haloscatella Mathis 1979 

Haloscatella arichaeta Mathis 1979 (Lamproscatella) 


163

164 

Haloscatella cephalotes Cresson 1935 (Lamproscatella) 

Haloscatella muria Mathis 1979 (Lamproscatella) 

Haloscatella nivosa Cresson 1935 (Lamproscatella) 

Haloscatella salinaria Sturtevant and Wheeler 1954 

(Lamproscatella) 

Hecamede Haliday 1837 

Hemicyclops Meijere 1913 Syn. 

Soikia Canzoneri and Meneghini 1969 Syn. 

Hecamede albicans Meigen 1830 (Notiphila) 

Psilopa globifera Boheman 1853 Syn. 

Hecamede grisescens Becker 1903 Syn. 

Hecamede granifera Thomson 1868 (Notiphila) 

Hecamede lacteipennis Lamb 1912 Syn. 

Hecamede persimilis Hendel 1913 Syn. 

Hecamede nivea Meijere 1916 Syn. 

Hecamede femoralis Malloch 1930 Syn. 

Hecamedoides Hendel 1917 

Hecamedoides unispinosa Collin 1943 (Discocerina) 

Helaeomyia Cresson 1941 

Helaeomyia petrolei Coquillett 1899 (Psilopa) 

Hyadina Haliday 1839 

Hydrina Robineau-Desvoidy 1830 Homo. 

Hyadina Haliday 1837 Unav. 

Ephydrosoma Lioy 1864 Syn. 

Neohydrellia Malloch 1933 Syn. 

Hyadina albovenosa Coquillett 1900 (Hyadina) 

Hyadina binotata Cresson 1926 (Hydrina) 

Hyadina macquarti Cresson 1930 Syn. 

Hyadina corona Cresson 1926 (Hydrina) 

Hyadina furva Cresson 1926 (Lytogaster) 

Hyadina flavipes Sturtevant and Wheeler 1954 Syn. 

Hyadina pruinosa Cresson 1926 (Hydrina) 

Hyadina subnitida Sturtevant and Wheeler 1954 (Hyadina) 

Hyadina vockerothi Clausen 1984 (Hyadina) 

Hydrellia Robineau-Desvoidy 1830 

Hydropota Rondani 1861 Syn. 

Hydrellia advenae Cresson 1934 (Hydrellia) 

Hydrellia agitator Deonier 1971 (Hydrellia) 

Hydrellia ainsworthi Deonier 1971 (Hydrellia) 

Hydrellia alabamae Deonier 1994 (Hydrellia) 

Hydrellia americana Cresson 1931 (Hydrellia) 

Hydrellia amnicola Deonier 1971 (Hydrellia) 

Hydrellia amplecta Deonier 1995 (Hydrellia) 

Hydrellia apalachee Deonier 1994 (Hydrellia) 

Hydrellia ascita Cresson 1942 (Hydrellia) 

Hydrellia atroglauca Coquillett 1910 (Hydrellia) 

Hydrellia bergi Cresson 1941 (Hydrellia) 

Hydrellia bicarina Deonier 1995 (Hydrellia) 

Hydrellia bilobifera Cresson 1936 (Hydrellia) 

Hydrellia biloxiae Deonier 1971 (Hydrellia) 

Hydrellia borealis Cresson 1944 (Hydrellia) 

Hydrellia bryani Deonier 1995 (Hydrellia) 

Hydrellia caliginosa Cresson 1936 (Hydrellia) 

Hydrellia cavator Deonier 1971 (Hydrellia) 

Hydrellia cessator Deonier 1971 (Hydrellia) 

Hydrellia columbata Deonier 1971 (Hydrellia) 

Hydrellia conformis Loew 1869 (Hydrellia) 

Hydrellia crassipes Cresson 1931 (Hydrellia) 



Hydrellia decens Cresson 1931 (Hydrellia) 

Hydrellia deceptor Deonier 1971 (Hydrellia) 

Hydrellia definita Cresson 1944 (Hydrellia) 

Hydrellia discursa Deonier 1971 (Hydrellia) 

Hydrellia fascitibia Roser 1840 (Notiphila) 

Notiphila grisea Stenhammar 1844 Syn. 

Hydrellia cruralis Coquillett 1910 Syn. 

Hydrellia cylindrica Cresson 1932 Syn. 

Hydrellia ruficornis Hendel 1933 Homo. 

Hydrellia halophila Canzoneri and Meneghini 1979 Syn. 

Hydrellia flavicoxalis Cresson 1944 (Hydrellia) 

Hydrellia floridana Deonier 1971 (Hydrellia) 

Hydrellia formosa Loew 1861 (Hydrellia) 

Hydrellia gladiator Deonier 1971 (Hydrellia) 

Hydrellia griseola Fallen 1813 (Notiphila) 

Tephritis hordei Oliver 1813 Syn. 

Tephritis pallida Oliver 1813 Syn. 

Notiphila chrysostoma Meigen 1830 Syn. 

Ephydra obscura Meigen 1830 Homo. 

Hydrellia communis Robineau-Desvoidy 1830 Syn. 

Hydrellia hypoleuca Loew 1862 Syn. 

Hydrellia scapularis Loew 1862 Syn. 

Hydrellia obscuriceps Loew 1862 Syn. 

Psilopa incerta Becker 1924 Syn. 

Hydrellia chinensis Qu and Li 1983 Syn. 

Hydrellia sinica Fall and Xia 1983 Syn. 

Hydrellia harti Cresson 1936 (Hydrellia) 

Hydrellia idolator Deonier 1971 (Hydrellia) 

Hydrellia insulata Deonier 1971 (Hydrellia) 

Hydrellia ischiaca Loew 1862 (Hydrellia) 

Hydrellia fallax Duda 1942 Syn. 

Hydrellia appendiculata Collin 1966 Syn. 

Hydrellia tomiokai Miyagi 1977 Syn. 

Hydrellia itascae Deonier 1971 (Hydrellia) 

Hydrellia johnsoni Cresson 1941 (Hydrellia) 

Hydrellia lata Cresson 1944 (Hydrellia) 

Hydrellia limnobii Deonier 1994 (Hydrellia) 

Hydrellia manitobae Deonier 1971 (Hydrellia) 

Hydrellia melanderi Deonier 1971 (Hydrellia) 

Hydrellia morrisoni Cresson 1924 (Hydrellia) 

Hydrellia najadis Deonier 1994 (Hydrellia) 

Hydrellia nobilis Loew 1862 (Psilopa) 

Hydrellia notata Deonier 1971 (Hydrellia) 

Hydrellia notiphiloides Cresson 1924 (Hydrellia) 

Hydrellia ocalae Deonier 1995 (Hydrellia) 

Hydrellia penicilli Cresson 1944 (Hydrellia) 

Hydrellia personata Deonier 1971 (Hydrellia) 

Hydrellia platygastra Cresson 1931 (Hydrellia) 

Hydrellia pontederiae Deonier 1994 (Hydrellia) 

Hydrellia proclinata Cresson 1915 (Hydrellia) 

Hydrellia proctori Cresson 1934 (Hydrellia) 

Hydrellia procteri Cresson 1936 Emend. 

Hydrellia prudens Curran 1930 (Hydrellia) 

Hydrellia pulla Cresson 1931 (Hydrellia) 

Hydrellia rixator Deonier 1971 (Hydrellia) 

Hydrellia saltator Deonier 1971 (Hydrellia) 

Hydrellia serena Cresson 1931 (Hydrellia) 

Hydrellia spinicornis Cresson 1918 (Hydrellia) 

Hydrellia subnitens Cresson 1931 (Hydrellia) 

Hydrellia surata Deonier 1971 (Hydrellia) 

Hydrellia suspecta Cresson 1936 (Hydrellia) 

Hydrellia tenebricosa Collin 1939 (Hydrellia) 

Hydrellia luctuosa Cresson 1942 Syn. 

Hydrellia nigra Miyagi 1977 Syn. 

Hydrellia tibialis Cresson 1917 (Hydrellia) 

Hydrellia diadema Frey 1930 Syn. 

Hydrellia tibiospica Deonier 1995 (Hydrellia) 

Hydrellia trichaeta Cresson 1944 (Hydrellia)

Hydrellia valida Loew 1862 (Hydrellia) 

Hydrellia wilburi Cresson 1944 (Hydrellia) 

Diclasiopa frontalis Curran 1933 Homo. 

Hydrochasma Hendel 1936 

Hydrochasma buccata Cresson 1930 (Hecamedoides) 

Hydrochasma facialis Williston 1896 (Discocerina) 

Hydrochasma zernyi Hendel 1936 Syn. 

Hydrochasma capax Cresson 1938 Syn. 

Hydrochasma incisa Coquillett 1902 (Discocerina) 

Discocerina poecilogastra Hendel 1930 Syn. 

Hydrochasma leucoprocta Loew 1861 (Discocerina) 

Ilythea Haliday 1839 

Ilythea Haliday 1837 Unav. 

Epipela Stenhammar 1844 Syn. 

Dichaetopareia Oldenberg 1923 Syn. 

Ilythea caniceps Cresson 1918 (Ilythea) 

Ilythea flaviceps Cresson 1916 (Ilythea) 

Ilythea spilota Curtis 1832 (Ephydra) 

Ephydra notata Stenhammar 1844 Syn. 

Lamproscatella Hendel 1917 

Lamproscatella aklavik Mathis 1979 (Lamproscatella) 

Lamproscatella bimaculata Hendel 1933 (Lamproscatella) 

Lamproscatella brunnipennis Malloch 1923 (Scatella) 

Lamproscatella norvegica Andersson 1975 Syn. 

Lamproscatella occidentalis Mathis 1979 (Lamproscatella) 

Lamproscatella sibilans Haliday 1833 (Ephydra) 

Scatella pilosigenis Becker 1896 Syn. 

Scatella faeroeensis Becker 1915 Syn. 

Lemnaphila Cresson 1933 

Lemnaphila scotlandae Cresson 1933 (Lemnaphila) 

Leptopsilopa Cresson 1922 

Leptopsilopa atrimana Loew 1878 (Psilopa) 

Leptopsilopa similis Coquillett 1900 (Psilopa) 

Leptopsilopa varipes Coquillett 1900 (Psilopa) 

Halaeomyia californica Cresson 1941 Syn. 

Limnellia Malloch 1925 

Eustigoptera Cresson 1930 Syn. 

Stictoscatella Collin 1930 Syn. 

Stranditella Duda 1942 Syn. 

Limnellia anderssoni Mathis 1978 (Limnellia) 

Limnellia anna Cresson 1935 (Limnellia) 

Limnellia balioptera Mathis 1978 (Limnellia) 

Limnellia huachuca Mathis 1978 (Limnellia) 

Limnellia lactea Mathis 1978 (Limnellia) 

Limnellia lecocercus Mathis 1978 (Limnellia) 

Limnellia rainier Mathis and Zack 1980 (Limnellia) 

Limnellia sejuncta Loew 1863 (Scatella) 

Limnellia stenhammari Zetterstedt 1846 (Ephydra) 

Ephydra meleagris Zetterstedt 1846 Unav. 

Ephydra oscitans Walker 1849 Syn. 

Limnellia sticta Mathis 1978 (Limnellia) 

Limnellia turneri Mathis 1978 (Limnellia) 

Lipochaeta Coquillett 1896 

Lipochaeta slossonae Coquillett 1896 (Lipochaeta) 


Lipochaeta texensis Townsend 1898 Syn. 

Lytogaster Becker 1896 

Lytogaster abdominalis Stenhammar 1844 (Notiphila) 

Pelina ventruosa Loew 1873 Syn. 

Lytogaster cervenkae Clausen 1982 (Lytogaster) 

Lytogaster excavata Sturtevant and Wheeler 1954 (Hyadina) 

Lytogaster gravida Loew 1863 (Hyadina) 

Lytogaster willistoni Cresson 1916 Syn. 

Lytogaster mexicana Clausen 1982 (Lytogaster) 

Lytogaster neglecta Sturtevant and Wheeler 1954 (Hyadina) 

Lytogaster obscura Clausen 1982 (Lytogaster) 

Lytogaster perplexa Clausen 1982 (Lytogaster) 

Mimapsilopa Cresson 1941 

Mimapsilopa cressoni Lizarralde de Grosso 1982 (Mimapsilopa) 

Mosillus Latreille 1804 

Gymnomyza Fallen 1810 Syn. 

Gymnopa Fallen 1820 Syn. 

Glabrinus Rondani 1856 Syn. 

Mosillus bidentata Cresson 1926 (Gymnopa) 

Mosillus nigroaenea Walker 1852 (Gymnopa) 

Mosillus stegmaieri Wirth 1969 (Mosillus) 

Mosillus tarsalis Walker 1852 (Gymnopa) 

Mosillus tibialis Cresson 1916 (Mosillus) 

Nostima Coquillett 1900 

Philygriola Hendel 1917 Syn. 

Nostima approximata Sturtevant and Wheeler 1954 (Nostima) 

Nostima gilvipes Coquillett 1900 (Hydrellia) 

Nostima immaculata Cresson 1918 Syn. 

Nostima giovannolii Wirth 1956 (Nostima) 

Nostima niveivenosa Cresson 1930 (Nostima) 

Nostima picta Fallen 1813 (Notiphila) 

Notiphila pullula Fallen 1823 Syn. 

Philygria nigripes Strobl 1880 Syn. 

Nostima quinquenotata Cresson 1930 (Nostima) 

Nostima schildi Cresson 1941 (Nostima) 

Nostima scutellaris Cresson 1933 (Nostima) 

Nostima occidentalis Sturtevant and Wheeler 1954 Syn. 

Nostima slossonae Coquillett 1900 (Nostima) 

Philygria calverti Cresson 1914 Syn. 

Notiphila Fallen 1810 

Dichaeta Meigen 1830 Syn. 

Keratocera Robineau-Desvoidy 1830 Syn. 

Notophila Agassiz 1846 Emend. 

Pegophila Rondani 1856 Syn. 

Notiphilacantha Hendel 1914 Syn. 

Agrolimna Cresson 1917 Syn. 

Notiphila adusta Mathis 1979 (Notiphila) 

Notiphila aenigma Cresson 1917 (Notiphila) 

Notiphila atrata Mathis 1979 (Notiphila) 

Notiphila atripes Cresson 1917 (Notiphila) 

Notiphila atriventris Cresson 1915 (Dichaeta) 

Notiphila avia Loew 1878 (Notiphila) 

Notiphila bella Loew 1862 (Notiphila) 

Notiphila biseriata Cresson 1917 (Notiphila) 

Notiphila bispinosa Cresson 1917 (Notiphila) 

Notiphila brevis Walker 1858 (Notiphila) 

Notiphila carinata Loew 1862 (Notiphila) 

Notiphila caudata Fallen 1813 (Notiphila) 


165

166 

Dichaeta tibialis Brulle 1832 Syn. 

Dichaeta brevicauda Loew 1860 Syn. 

Notiphila cognata Cresson 1917 (Notiphila) 

Notiphila cressoni Mathis 1979 (Notiphila) 

Notiphila bicolor Cresson 1917 Homo. 

Notiphila decorata Williston 1896 (Notiphila) 

Dichaeta furcata Coquillett 1902 Syn. 

Notiphila decoris Williston 1893 (Notiphila) 

Notiphila atrisetis Cresson 1917 Syn. 

Notiphila deonieri Mathis 1979 (Notiphila) 

Notiphila deserta Mathis 1979 (Notiphila) 

Notiphila eleomyia Mathis 1979 (Notiphila) 

Notiphila elophila Mathis 1979 (Notiphila) 

Notiphila erythrocera Loew 1878 (Notiphila) 

Notiphila bellula Williston 1896 Syn. 

Notiphila varia Jones 1906 Syn. 

Notiphila punctifera Cresson 1917 Syn. 

Notiphila pura Cresson 1917 Syn. 

Notiphila rubricornis Becker 1919 Syn. 

Notiphila floridensis Cresson 1917 (Notiphila) 

Notiphila footei Mathis 1979 (Notiphila) 

Notiphila hamifera Wheeler 1961 (Notiphila) 

Notiphila kentensis Huryn 1987 (Notiphila) 

Notiphila latelimbata Curran 1930 (Notiphila) 

Notiphila latigena Mathis 1979 (Notiphila) 

Notiphila loewi Cresson 1917 (Notiphila) 

Notiphila macrochaeta Loew 1878 (Notiphila) 

Notiphila brachychaeta Cresson 1946 Syn. 

Notiphila mathisi Huryn 1984 (Notiphila) 

Notiphila minima Cresson 1917 (Notiphila) 

Notiphila nanosoma Mathis 1979 (Notiphila) 

Notiphila nudipes Cresson 1917 (Notiphila) 

Notiphila olivacea Cresson 1917 (Notiphila) 

Notiphila oriens Mathis 1979 (Notiphila) 

Notiphila pallicornis Mathis 1979 (Notiphila) 

Notiphila pallidipalpis Cresson 1940 (Notiphila) 

Notiphila paludia Mathis 1979 (Notiphila) 

Notiphila pauroura Mathis 1979 (Notiphila) 

Notiphila phaeopsis Mathis 1979 (Notiphila) 

Notiphila poliosoma Mathis 1979 (Notiphila) 

Notiphila pulchra Mathis 1979 (Notiphila) 

Notiphila pulchrifrons Loew 1872 (Notiphila) 

Notiphila signata Cresson 1917 Syn. 

Notiphila quadrisetosa Thomson 1869 (Notiphila) 

Notiphila occidentalis Cresson 1917 Syn. 

Notiphila robusta Mathis 1979 (Notiphila) 

Notiphila scalaris Loew 1862 (Notiphila) 

Notiphila scoliochaeta Mathis 1979 (Notiphila) 

Notiphila shewelli Mathis 1979 (Notiphila) 

Notiphila sicca Cresson 1940 (Notiphila) 

Notiphila solita Walker 1852 (Notiphila) 

Notiphila vittata Loew 1862 Syn. 

Notiphila taenia Mathis 1979 (Notiphila) 

Notiphila theonae Huryn 1984 (Notiphila) 

Notiphila transversa Walker 1852 (Notiphila) 

Notiphila uliginosa Haliday 1839 (Notiphila) 

Notiphila tarsata Stenhammar 1844 Syn. 

Notiphila unicolor Loew 1862 (Notiphila) 

Ochthera Latreille 1802 

Macrochira Meigen 1803 Syn. 

Haplochila Enderlein 1922 Syn. 

Stenochthera Hendel 1930 Syn. 

Ochthera acta Clausen 1977 (Ochthera) 

Ochthera anatolikos Clausen 1977 (Ochthera) 

Ochthera baia Cresson 1931 (Ochthera) 

Ochthera borealis Clausen 1977 (Ochthera) 



Ochthera collina Clausen 1977 (Ochthera) 

Ochthera cuprilineata Wheeler 1896 (Ochthera) 

Ochthera cuprilineata Williston 1896 Homo. 

Ochthera humilis Williston 1897 Syn. 

Ochthera melanderi Cresson 1944 Syn. 

Ochthera exsculpta Loew 1862 (Ochthera) 

Ochthera lauta Wheeler 1896 (Ochthera) 

Ochthera loreta Cresson 1931 (Ochthera) 

Ochthera mantis De Geer 1776 (Musca) 

Ochthera occidentalis Clausen 1977 (Ochthera) 

Ochthera pilosa Cresson 1926 (Ochthera) 

Ochthera tuberculata Loew 1862 (Ochthera) 

Ochthera rapax Loew 1862 Syn. 

Oedenops Becker 1903 

Oedenops nuda Coquillett 1902 (Paralimna) 

Paracoenia Cresson 1935 

Thiomyia Wirth 1954 Syn. 

Paracoenia ampla Mathis 1975 (Paracoenia) 

Paracoenia bisetosa Coquillett 1902 (Coenia) 

Paracoenia calida Mathis 1976 (Paracoenia) 

Paracoenia fumosalis Cresson 1935 (Paracoenia) 

Paracoenia quatei Wirth 1954 (Thiomyia) 

Paracoenia turbida Curran 1927 (Coenia) 

Paracoenia wirthi Mathis 1975 (Paracoenia) 

Paralimna Loew 1862 

Phaiosterna Cresson 1916 Syn. 

Poecilothorax Becker 1922 Syn. 

Afrolimna Cogan 1968 Syn. 

Paralimna decipiens Loew 1878 (Paralimna) 

Paralimna multipunctata Williston 1896 (Paralimna) 

Paralimna ciliata Cresson 1916 Syn. 

Paralimna obscura Williston 1896 (Paralimna) 

Paralimna punctipennis Wiedemann 1830 (Notiphila) 

Paralimna appendiculata Loew 1862 Syn. 

Paralimna texana Cresson 1915 (Paralimna) 

Paratissa Coquillett 1900 

Acanthonotiphila Lamb 1912 Syn. 

Paratissa semilutea Loew 1869 (Cacoxenus) 

Drosophila pollinosa Williston 1896 Syn. 

Parydra Stenhammar 1844 

Napaea Robineau-Desvoidy 1830 Homo. 

Falosoma Lioy 1864 Syn. 

Hygrophila Lioy 1864 Syn. 

Chaetoapnaea Hendel 1930 Syn. 

Paranapaea Hendel 1930 Syn. 

Parydra abbreviata Loew 1861 (Parydra) 

Napaea hulli Cresson 1934 Syn. 

Parydra acuta Clausen 1971 (Parydra) 

Parydra alpina Cresson 1924 (Napaea) 

Parydra appendiculata Loew 1878 (Parydra) 

Parydra aquila Fallen 1813 (Ephydra) 

Parydra bituberculata Loew 1862 Syn. 

Parydra nitida Cresson 1915 Syn. 

Parydra tibialis Cresson 1916 Syn. 

Parydra papulata Cresson 1949 Syn. 

Parydra arctica Clausen 1971 (Parydra) 

Parydra aurata Jones 1906 (Parydra) 

Parydra borealis Cresson 1949 (Napaea)

Parydra breviceps Loew 1862 (Parydra) 

Parydra limpidipennis Loew 1878 Syn. 

Parydra vicina Cresson 1940 Syn. 

Parydra copis Clausen 1971 (Parydra) 

Parydra fossarum Haliday 1833 (Ephydra) 

Napaea minor Robineau-Desvoidy 1830 Syn. (?) 

Ephydra affinis Stenhammar 1844 Syn. 

Parydra parasocia Clausen 1971 Syn. 

Parydra halteralis Cresson 1930 (Napaea) 

Parydra hamata Clausen 1971 (Parydra) 

Parydra humilis Williston 1897 (Parydra) 

Parydra imitans Loew 1878 (Parydra) 

Parydra trituberculata Sturtevant and Wheeler 1954 Syn. 

Parydra incommoda Cresson 1930 (Parydra) 

Parydra joaquinensis Clausen 1971 (Parydra) 

Parydra lingulata Clausen 1971 (Parydra) 

Parydra parva Cresson 1949 (Parydra) 

Napaea undulata Cresson 1934 Homo. 

Parydra paullula Loew 1862 (Parydra) 

Parydra pedalis Clausen 1971 (Parydra) 

Parydra penabbreviata Clausen 1971 (Parydra) 

Parydra penisica Clausen 1971 (Parydra) 

Parydra pinguis Walker 1852 (Ephydra) 

Parydra quadriloba Clausen 1971 (Parydra) 

Parydra quadrituberculata Loew 1862 (Parydra) 

Parydra socia Cresson 1934 (Napaea) 

Parydra spinosa Clausen 1971 (Parydra) 

Parydra succurva Clausen 1971 (Parydra) 

Parydra transversa Cresson 1940 (Parydra) 

Parydra unituberculata Loew 1878 (Parydra) 

Parydra vanduzeei Cresson 1933 (Napaea) 

Parydra varia Loew 1863 (Parydra) 

Parydra vulgaris Cresson 1949 (Napaea) 

Pelina Haliday 1839 

Pelina Haliday 1837 Unav. 

Telmatobia Stenhammar 1844 Syn. 

Calocephala Zetterstedt 1846 Unav. 

Pelina biloba Clausen 1973 (Pelina) 

Pelina bispinosa Clausen 1973 (Pelina) 

Pelina canadensis Cresson 1934 (Pelina) 

Pelina compar Cresson 1934 (Pelina) 

Pelina rudis Cresson 1940 Syn. 

Pelina latiforma Clausen 1973 (Pelina) 

Pelina neaenescens Clausen 1973 (Pelina) 

Pelina prospinosa Clausen 1973 (Pelina) 

Pelina subulata Clausen 1973 (Pelina) 

Pelina truncatula Loew 1878 (Pelina) 

Pelina uncata Clausen 1973 (Pelina) 

Philotelma Becker 1896 

Pseudoscatella Becker 1902 Syn. 

Philotelma alaskensis Cresson 1935 (Philotelma) 

Philygria Stenhammar 1844 

Cressoniella Saether 1970 Homo. 

Philygria debilis Loew 1861 (Philygria) 

Philygria fuscicornis Loew 1862 Syn. 

Cressoniella montana Saether 1970 Syn. 

Philygria dimidiata Sturtevant and Wheeler 1954 (Hydrina) 

Philygria nigrescens Cresson 1930 (Hydrina) 

Philygria punctatonervosa Fallen 1813 (Notiphila) 

Philygria opposita Loew 1861 Syn. 

Philygria vittipennis Zetterstedt 1837 (Notiphila) 


Placopsidella Kertesz 1901 

Enchastes Lamb 1912 Syn. 

Oscinomima Enderlein 1912 Syn. 

Placopsidella grandis Cresson 1925 (Gymnopa) 

Placopsidella opaca Miyagi 1977 Syn. 

Placopsidella rossii Canzoneri 1986 Syn. 

Platygymnopa Wirth 1971 

Platygymnopa helicis Wirth 1971 (Platygymnopa) 

Polytrichophora Cresson 1924 

Polytrichophora agens Cresson 1924 (Polytrichophora) 

Polytrichophora conciliata Cresson 1924 (Polytrichophora) 

Polytrichophora orbitalis Loew 1861 (Discocerina) 

Polytrichophora setigera Cresson 1916 (Discocerina) 

Pseudohyadina Clausen 1983 

Pseudohyadina longicornis Sturtevant and Wheeler 1954 (Hyadina) 

Psilopa Fallen 1823 

Hygrella Haliday 1839 Syn. 

Ephygrobia Schiner 1862 Syn. 

Diasemocera Bezzi 1895 Syn. 

Domina Hutton 1901 Syn. 

Discocerinella Mercier 1927 Syn. 

Psilopa compta Meigen 1830 (Notiphila) 

Psilopa gaudens Cresson 1946 Unav. 

Psilopa dupla Cresson 1940 (Psilopa) 

Psilopa flavida Coquillett 1900 (Psilopa) 

Psilopa girschneri Röder 1889 (Psilopa) 

Psilopa nigropuncta Williston 1896 Syn. 

Psilopa olga Cresson 1922 Syn. 

Psilopa dimidiata Cresson 1922 Syn. 

Psilopa leucostoma Meigen 1830 (Notiphila) 

Psilopa pulchripes Loew 1878 (Psilopa) 

Psilopa unica Cresson 1926 Syn. 

Ptilomyia Coquillett 1900 

Atissiella Cresson 1918 Syn. 

Alocatissa Hendel 1930 Syn. 

Ptilomyia alkalinella Cresson 1942 (Atissiella) 

Ptilomyia enigma Coquillett 1900 (Ptilomyia) 

Atissiella falsisetae Cresson 1942 Syn. 

Ptilomyia lobiochaeta Sturtevant and Wheeler 1954 (Ptilomyia) 

Ptilomyia mabelae Cresson 1926 (Atissiella) 

Ptilomyia occidentalis Sturtevant and Wheeler 1954 (Ptilomyia) 

Ptilomyia parva Williston 1896 (Hydrellia) 

Ptilomyia pleuriseta Cresson 1942 (Atissiella) 

Rhinonapaea Wirth 1965 

Rhinonapaea metallica Cole 1921 (Parydra) 

Rhysophora Cresson 1924 

Rhysophora robusta Cresson 1924 (Rhysophora) 

Discocerina magna Coquillett 1910 Unav. 

Scatella Robineau-Desvoidy 1830 

Teichomyza Macquart 1835 Syn. 

Trixostomus Rondani 1856 Syn. 

Synhoplos Lamb 1917 Syn. 


167

168 

Neoscatella Malloch 1933 Syn. 

Apulvillus Malloch 1934 Syn. 

Chaetoscatella Malloch 1934 Syn. 

Parascatella Cresson 1935 Syn. 

Strandella Duda 1942 Syn. 

Scatella arizonensis Cresson 1935 (Scatella) 

Scatella crassicosta Becker 1896 (Scatella) 

Scatella lindbergi Dahl 1959 Syn. 

Scatella favillacea Loew 1862 (Scatella) 

Scatella nitidifrons Cresson 1918 Syn. 

Scatella laxa Cresson 1933 (Scatella) 

Scatella marinensis Cresson 1935 (Parascatella) 

Scatella melanderi Cresson 1935 (Parascatella) 

Scatella obscura Williston 1896 (Scatella) 

Scatella laetifica Cresson 1934 Syn. 

Scatella obscuriceps Cresson 1915 (Scatella) 

Scatella obsoleta Loew 1861 (Scatella) 

Scatella callosicosta Bezzi 1895 Syn. 

Scatella strandi Duda 1942 Syn. 

Scatella paludum Meigen 1830 (Ephydra) 

Ephydra leucostoma Meigen 1830 Syn. 

Ephydra sorbillans Haliday 1839 Syn. 

Ephydra argyrostoma Stenhammar 1844 Syn. 

Scatella pentastigma Thomson 1869 (Ephydra) 

Scatella picea Walker 1849 (Ephydra) 

Scatella lugens Loew 1862 Syn. 

Scatella quadrinotata Cresson 1933 (Scatella) 

Scatella setosa Coquillett 1900 (Scatella) 

Scatella intermedia Cresson 1915 Syn. 

Scatella stagnalis Fallen 1813 (Ephydra) 

Ephydra lacustris Meigen 1830 Syn. 

Scatella buccata Robineau-Desvoidy 1830 Syn. 

Scatella tateyamana Matsumura 1915 Syn. 

Scatella tenuicosta Collin 1930 (Scatella) 

Scatella thermarum Collin 1930 Syn. 

Scatella triseta Coquillett 1902 (Scatella) 

Scatella troi Cresson 1933 (Scatella) 

Scatophila Becker 1896 

Centromeromyia Frey 1954 Syn. 



Scatophila adamsi Cresson 1935 (Scatophila) 

Scatophila arenaria Cresson 1935 (Scatophila) 

Scatophila bipiliaris Sturtevant and Wheeler 1954 (Scatophila) 

Scatophila bisignata Cresson 1935 (Scatophila) 

Scatophila carinata Sturtevant and Wheeler 1954 (Scatophila) 

Scatophila conifera Sturtevant and Wheeler 1954 (Scatophila) 

Scatophila cribrata Stenhammar 1844 (Ephydra) 

Ephydra striata Walker 1849 Syn. 

Scatophila despecta Haliday 1839 (Ephydra) 

Ephydra fenestrata Stenhammar 1844 Syn. 

Scatophila hamifera Becker 1896 Syn. 

Scatophila disjuncta Cresson 1935 (Scatophila) 

Scatophila exilis Cresson 1935 (Scatophila) 

Scatophila hesperia Sturtevant and Wheeler 1954 (Scatophila) 

Scatophila hirsuta Sturtevant and Wheeler 1954 (Scatophila) 

Scatophila hirtirostris Sturtevant and Wheeler 1954 (Scatophila) 

Scatophila iowana Wheeler 1961 (Scatophila) 

Scatophila mesogramma Loew 1869 (Scatella) 

Scatophila infantina Becker 1926 Syn. 

Scatophila ordinaria Sturtevant and Wheeler 1954 (Scatophila) 

Scatophila picta Sturtevant and Wheeler 1954 (Scatophila) 

Scatophila pulchra Sturtevant and Wheeler 1954 (Scatophila) 

Scatophila rubribrunnea Sturtevant and Wheeler 1954 (Scatophila) 

Scatophila tuberculosa Cresson 1935 (Scatophila) 

Scatophila unicornis Czerny 1900 (Scatophila) 

Scatophila kuntzei Kramer 1917 Syn. 

Scatophila affinis Bolwig 1940 Unav. 

Scatophila poeciloptera Papp 1975 Syn. 

Scatophila variabilis Cresson 1917 (Scatophila) 

Scatophila variofacialis Sturtevant and Wheeler 1954 (Scatophila) 

Scatophila viridella Sturtevant and Wheeler 1954 (Scatophila) 

Schema Becker 1907 

Pelignus Cresson 1926 Syn. 

Atissina Enderlein 1936 Unav. 

Atissina Cresson 1936 Unav. 

Pseudoedenops Seguy 1951 Syn. 

Schema salinus Cresson 1942 (Pelignus) 

Setacera Cresson 1930 

Setacera aldrichi Cresson 1935 (Setacera) 

Setacera atrovirens Loew 1862 (Ephydra) 

Setacera durani Cresson 1935 (Setacera) 

Setacera jamesi Mathis 1982 (Setacera) 

Setacera micans Haliday 1833 (Ephydra) 

Setacera needhami Johannsen 1935 (Setacera) 

Setacera pacifica Cresson 1925 (Ephydra) 

Setacera pilicornis Coquillett 1902 (Ephydra) 

Setacera knabi Cresson 1935 Syn. 

Thinoscatella Mathis 1979 

Thinoscatella quadrisetosa Becker 1896 (Scatella) 

Trimerina Macquart 1835 

Trimerina madizans Fallen 1813 (Notiphila) 

Notiphila rufipes Meigen 1830 Syn. 

Notiphila nigella Meigen 1830 Syn. 

Hydrellia lepida Meigen 1838 Syn. 

Trimerinoides Cresson 1925 

Trimerinoides adfinis Cresson 1922 (Trimerina) 

Typopsilopa Cresson 1916 

Psilopina Becker 1926 Syn. 

Typopsilopa antennalis Wirth 1968 (Typopsilopa) 

Typopsilopa arnaudi Wirth 1968 (Typopsilopa) 

Typopsilopa atra Loew 1862 (Psilopa) 

Psilopa scoriacea Loew 1862 Syn. 

Typopsilopa nigra Williston 1896 (Psilopa) 

Typopsilopa flavitarsis Cresson 1916 Syn.

Cecidomyiidae - Ephydridae; pp. 98 - Nearctica

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