Section 2. Checklist of Plants and Fungi by smeathmanella

Annotated Catalogue of Biodiversity of the Doward and Symonds Yat West Ian Draycott

August’23

Section 1. The Plant and Fungal Kingdoms Overview. According to data presented in the National Biodiversity Network (NBN) Atlas in 2017 at least 1,258 species of plants and 1,556 species of fungi and lichens have been recorded within 2km of the centre of Great Doward. This total reflects the range of terrestrial macro and micro habitats arising from the combination of both Carboniferous limestone and Devonian sandstone underlaying the site. Most of the habitats are woodland or dry open areas of grassland and rock exposures but the small marshy areas around Dropping Wells and the seasonally flooded zone along the riverbank also adds to the plant diversity. Looking at the NBN plant data in more detail gives 784 species of flowering plants (Angiosperms and Gymnosperms) and 26 ferns (Pteridophytes) and spore-forming allies, 344 species are bryophytes (mosses and liverworts) and a further 5 are species of green algae. The data on fungi includes lichens which are a symbiotic association between and a fungal thallus and an algae or blue-green bacteria. It should be stressed that this dataset will not be a precise figure of what is actually present on the Doward today as a good number of these records are quite old or do not have adequate information about the location. Some records may not be on the Doward side of the river however, other species will be present but have yet to be found and recorded.

To simplify the production of checklists this publication uses the following main divisions:

Vascular Plants

Non-vascular Plants

Fungi

Angiosperms – Flowering Plants

Page 2.

Gymnosperms – Conifers and allies

Page 5.

Pteridophytes – Ferns and allies

Page 6.

Bryophytes: Mosses

Page 10.

Liverworts

Page 12.

Chlorophyta – Algae

Page 15.

Larger Fungi

Page 16.

Lichen

Page 21.

Supplementary Note: Endemic Plants of the Doward

Page 22.

These plant checklists cover species considered to be ‘native’ or ‘naturalised in UK. They excluded a range of species deliberately planted in gardens, horticulture and forestry estates. They also do not include details of some aggregates of species where morphological differences are minor and hard to discern in the field. As a consequence, these lists and their species totals do not fully match all species listed in the larger NBN or BSBI databases where expert interpretation has been required.

1.1 Checklist of Vascular Plants The list below is generated from a range of ‘informal’ observations by various recorders on the Doward rather than a set of formal records. Although each species listed has been reported on the Doward not all have been verified so should be treated as being for guidance purposes only. Angiosperms - Flowering Plants Ranunclulaceae Creeping Buttercup Meadow Buttercup Lesser Celandine Goldilocks Buttercup Wood anemone Columbine Travellers Joy Stinking Hellebore Papaveraceae Opium Poppy Greater Celandine Fumiariaceae Yellow Corydalis Common Fumitory Ulmaceae Wych Elm Viscacae Mistletoe Urticaceae Common nettle Pellitory-of-the-Wall Fagaceae Sessile Oak Sweet Chestnut Beech Betulaceae Silver/Downy Birch Alder Grey Alder Hazel Chenopodiaceae Fat-hen Caryophyllaceae Red Campion White campion Bladder Campion Common Chickweed Greater Stichwort Common Mouse-ear Thyme-leaved Sandwort Annual Pearlwort Knotted/procumbent pearlwort Polygonacea Knotgrass Redshank Water Pepper Common Sorrel Broad-leaved dock Curled Dock Japanese knotweed Black Bindweed

Ranunculus repens Ranunculus acris Ranunclus ficaria Ranunclulus auricomus Anemone nemorosa Aquilega vulgaris Clematis vitalba Heleborus viridis Papavera somniferum Chelidonium majus Pseudofumaria lutea Fumaria officinalis Ulmus glabra Viscum album Urtica dioica Parietaria judaica Quercus petraea Castanea sativa Fagus sylvatica Betula pendula/pubescens Alnus glutinosa Alnus incana Corylus avellana Chenopodium album Silene dioca Silene latifolia Silene vulgaris Stellaria media Stellaria holostea Cerastium fontanum ag Arenaria serpyllifolia Sagina apetala Sagina nodosa/procumens Polygonum aviculare Percicaria maculosa Percicaria hydropiper Rumex acetosa Rumex obtusifolius Rumex crispus Fallopia japonica Fallopia convolvulus

Clusiaceae Hairy St Johns-wort Tutsan Perforate St John's-wort Slender St John's-wort Pale St John's-wort Square-stalked St John's-wort Malvaceae Musk Mallow Common Mallow Tiliaceae Large-leaved Lime Small-leaved Lime Cistaceae Common Rock-rose Salicaceae White Poplar Sallow/Grey Willow Creeping Willow Violaceae Common Dog-violet Early Dog-violet Hairy Violet Sweet Violet Brassicaceae Hairy Bittercress Wavy Bittercress Narrow-leaved Bittercress Lady's smock Common Whitlowgrass Hutchinsia Thale Cress Treacle Mustard Swine-cress Hairy Rock-cress Honesty Shepard's Purse Hedge Mustard Garlic Mustard Charlock Rape Creeping Yellow-cress Resedaceae Wild mignonette Weld Dipsacaceae Wild Teasel Small Teasel Field Scabious Small Scabious Devil’s-bit Scabious Astercea Daisy Ox-eye Daisy Blue Fleabane Scentless Mayweed Common Fleabane Common Ragwort Hoary Ragwort Groundsel Ploughman's-spikenard Yarrow Dandelion

Hypericum hirsutum Hypericum androsaemum Hypericum perforatum Hypericum pulchrum Hypericum montanum Hypericum tetrapterum Malva moschata Malva sylvestris Tilia platyphyllos Tilia cordata Helianthemum apenninum Populus alba Salix cinerea Salix repens Viola riviana Viola riechenbachiana Viola hirta Viola odorata Cardamine hirsuta Cardamine flexuosa Cardamine impatiens Cardamine pratensis Erophila verna Hornungia petraea Arabidopsis thaliana Erysimum cheiranthoides Coronopus squamatus Arabis hirsuta Lunaria annua Capsella bursa-pastoris Sisimbryum officinale Alliaria petiolata Sinapsis arvensis Brassica napus Rorippa sylvestris Reseda lutea Reseda luteola Dipsacus fullonum Dipsacus pilosus Knautia arvensis Scabiosa columbaria Succisa pratensis Bellis perrenis Leucanthemum vulgare Erigeron acer Tripleurospermum inodorum Pulicaria dysenterica Senecio jacobaea Senecio eruciformis Senecio vulgaris Inula conyzae Achillea millefolium Taraxacum officinale

Canadian Goldenrod Goldenrod Pineappleweed Hemp Agrimony Mugwort Greater Burdock Nipplewort Mouse-ear Hawkweed Goat's-beard Hawkweed Oxtongue Creeping Thistle Spear Thistle Marsh Thistle Common Knapweed Cat's-ear Rough Hawksbit Autumn Hawkbit Smooth Hawks-beard ag Hawkweed ag. Prickly Sow-thistle Smooth Sow-thistle Wall Lettuce Great Lettuce Coltsfoot Plantaginaceae Ribwort Plantain Greater Plantain Crassulacea Wall Pennywort or Navelwort English/Biting Stonecrop Reflexed Stonecrop Gentianaceae Yellowwort Common Centuary Saxifragaceae Opposite-leaved Golden Saxifrage Rosaceae Bramble Field rose Dog rose Agrimony Fragrant Agrimony Salad Burnet Meadowsweet Dropwort Wood Avens Silverweed Creeping Cinquefoil Trailing Tormentil Barren Strawberry Wild Strawberry Parsley-piert Lady's Mantle Wild Cherry Blackthorn Crab Apple Wild Service Tree Whitebeam group Rowan Cotoneaster Fabeacea Tufted Vetch Bush Vetch

Solidago canadensis Solidago virguarea Matricaria discoides Eupatorium cannabinum Artemesia vulgaris Arctium lappa Lapsana communis Philosella officinarum Tragopogon pratensis Picris hieracoides Cirsium arvense Cirsium vulgare Cirsium palustre Centaurea nigra Hypochaeris radicata Leontodon hispidus Leontodon autumnalis Crepis spp Hieraceum spp Sonchus asper Sonchus oleraceus Mycelis muralis Lactuca virosa Tussilago farfara Plantago lanceolata Plantago major Umbilicus rupestris Sedum anglicum/acre Sedum rupestre Blackstonia perfoliata Centaurium erythraea Chryosplenium alternifolium Rubus fruticosus agg. Rosa arvensis Rosa canina agg. Agrimonia eupatoria Agrimonia procera Sanguisorba officinalis Filipendula ulmaria Filipendula vulgaris Geum urbanum Potentilla anserina Potentilla reptens Potentilla anglica Potentilla sterilis Fragaria vesca Aphanes arvensis Alchemilla sp agg. Prunus avium Prunus spinosa Malus sylvestris Sorbus torminalis Sorbus aria ag see text Sorbus aucuparia Cotoneaster agg. Vicia cracca Vicia sepium

Hairy tare Common Vetch Wood Bitter-vetch Kidney Vetch Bitter Vetch(ling) Meadow Vetchling Spotted Medick Black Medick Red Clover Zigzag Clover White Clover Lesser Trefoil Hop Trefoil Common Birds-foot-trefoil Greater Bird's-foot-trefoil Common Restharrow Horseshoe Vetch Broom Western Gorse Geraniaceae Bloody Cranesbill Herb-Robert French Crane's-bill Cut-leaved Cranesbill Linaceae Fairy Flax Euphoriaceae Dog's Mercury Wood Spurge Petty Spurge Polygalaceae Common Milkwort Thymelaeaceae Spurge Laurel Aceracea Field Maple Sycamore Oxalidaceae Wood Sorrel Cucurbitacea White Bryony Onagraceae Enchanters Nightshade Large-flowered Evening primrose Rosebay Willowherb Great Willowherb Broad-leaved Willowherb Hoary Willowherb Cornaceae Dogwood Viscaceae Mistletoe Celastraceae Spindle Aquifoliaceae Holly Apiaceae Upright Hedge Parsley Cow Parsley Wild Angelica Burnet-saxifrage Wild Carrot Hogweed

Vicia tetrasperma Vicia sativa Vicia orobus Anthyllis vulneraria Lathyrus linifolius Lathyrus pratensis Medicago arabica Medicogo lupulina Trifolium pratense Trifolium medium Trifolium repens Trifolium dubium Trifolium campestre Lotus corniculatus Lotus pedunculatus Ononis repens Hippocrepis camosa Cytisus scoparius Ulex gallii Geranium sanguineum Geranium robertianum Geranium endressi Geranium dissectum Linium catharticum Mercurialis annua Euphorbia amygdaloides Euphorbia peplus Polygala vulgaris Daphne laurelo Acer campestre Acer pseudoplatanus Oxalis acetosella Bryonia dioica Circaea lutetiana Oenothera erythrosepala Chamerion angustifolium Epilobium hirsutum Epilobium montanum Epilobium parviflorum Cornus sanguinea Viscum album Euonymus europeaus Ilex aquifolium Torilis japonica Anthriscus sylvestris Angelica sylvestris Pimpinrella saxifraga Daucus carota Heracleum sphondylium

Fool's Parsley Sanicle Pignut Ericaceae Ling Heather Bilberry Primulaceae Primrose Cowslip Yellow Pimpernel Scarlet Pimpernel Apocyanaceae Greater Periwinkle Rubiaceae Field Madder Wild Madder Lady's Bedstraw Hedge bedstraw Cleavers Woodruff Crosswort Convolvulaceae Hedge bindweed Field Bindweed Boraginaceae Common Gromwell Common Comfrey Green Alkanet Field Forgetmenot Lamiaceae White Deadnettle Red Deadnettle Common Hemp-nettle Bugle Yellow Archangel Betony Wild Clary Hedge Woundwort Marsh Woundwort Ground ivy Selfheal Wild Basil Common Calamint Wild Marjoram Thyme Wood Sage Gypsywort Solanaceae Bittersweet Deadly Nightshade Scrophulariaceae Wall Speedwell Heath Speedwell Germander Speedwell Common Field speedwell Thyme-leaved Speedwell Lousewort Great Mullein Common Figwort Ivy-leaved Toadflax Common Cow-wheat Red Bartsia Foxglove

Aethusa cynapium Sanicula europaea Conopodium majus Calluna vulgaris Vaccinium myrtillus Primula vulgaris Primula veris ag Lysimachia nemorum Anagallis arvensis Vinca major Sherardia arvensis Rubia peregrina Galium verum Galium mollugo Galium aparine Galium odoratum Cruciata laevipes Calystegia sepium Convolvulus arvensis Lithospermum offinale Symthytum offininalis Pentaglottis sempervirens Myosotes arvensis Lamium album Lamium purpureum Galeopsis tetrahit Ajuga reptans Lamiastrum galeobdolon Stachys officinalis Salvia verbenacea Stachys sylvatica Stachys palustris Glechoma hederacea Prunella vularis Clinopodium vulgare Clinopodium ascendens Origanum vulgare Thymus polytrichus Teucrium scorodonia Lycopus europaeus Solanum dulcamara Atropa belladonna Veronica arvensis Veronica officinalis Veronica chamaedrys Veronia persica Veronica serpyllifolia Pedicularis sylvatica Verbascum thapsus Scropularia nodosa Cymalaria muralis Melampyrum pratense Odontites vernus Digitalis purpurea

Yellow-rattle Eyebright Caprofoliaceae Honeysuckle Red Valerian Campanulaceae Hairbell Nettle-leaved Bellflower Monotropidae Yellow Birds-nest Araceae Lords-and-Ladies Amaryllidaceae Wild Daffodil Snowdrop Summer Snowflake Iridacea Yellow Flag Stinking Iris Montbretia Dioscoreaceae Black Bryony Liliacea Three-cornered Garlic Ramsons Bluebell Meadow Saffron Solomon's Seal Orchidacea Common Spotted Orchid Early Purple Orchid Bird's-nest Orchid Greater Butterfly Orchid Common Twayblade Bee Orchid Pyramidal Orchid Fly Orchid Broad-leaved Helleborine White Helleborine Poaceae Sweet Vernal grass Crested Dogs tail Meadow Foxtail Smooth Meadow-grass Rough Meadow-grass Annual Meadow-grass Flattened Meadow-grass Wood Meadow-grass Yorkshire Fog Creeping Soft-grass Cocksfoot Fern-grass Tufted Hair-grass Wavy Hair-grass False Brome Sheep's Fescue Red Fescue Rat's-tail Fescue Wood Melick Mountain Melick False Oat Grass Quaking Grass Perennial Rye grass

Rhinanthus minor Euphrasia ag Lonicera pericyclamenum Centranthus ruber Campanula rotundifolisa Campanula trachelium Monotropa hypopitys Arum maculatum Narcissus sp Galanthus nivalis Leucojum aestivum Iris pseudacorus Iris foetidissima Crocosmia xcrocosmiiflora Tamus communis Allium triquestrum Allium ursinum Hyacinthoides non-scripta Colchicum autumnale Polygonatum multiflorum Dactylorhiza fuchsii Orchis mascula Neottia nidus-avis Platanthera chlorantha Neottia ovata Ophrys apifera Anacamptis pyramidalis Ophrys insectifera Epipactis helleborine Cephalanthera damasonium Anthroxanthum odoratum Cynosurus cristatus Alopecurus pratensis Poa pratensis Poa trivialis Poa annua Poa compressa Poa nemoralis Holcus lanatus Holcus mollis Dactylis glomerata Catapodium rigidum Deschampsia caespitosa Deschampsia flexuosa Brachypodium sylvaticum sl. Festuca ovina Festuca rubra Vulpia myuros Melica uniflora Melica nutans Arrhenatherum elatius Briza media Lolium perene

Soft Brome Upright Brome Blue Moor Grass Wood Barley Timothy Heath Grass Common Reed Cyperaceae Glaucous Sedge Pendulous Sedge Soft-leaved Sedge Fingered Sedge Dwarf Sedge Spring Sedge Grey Sedge Remote Sedge Pale Sedge Wood Sedge Yellow Sedge Juncaceae Field Wood-rush Hairy Wood-rush Jointed Rush Slender Rush Hard Rush Compact rush Toad Rush

Bromus mollis Bromus erectus Sesleria caerulens Hordelymus europaeus Phleum pratense Danthonia decumbens Phragmites australis Carex flacca Carex pendula Carex montana Carex digitata Carex humilis Carex caryophyllacea Carex divulsa Carex remota Carex pallescens Carex sylvatica Carex viridula Luzula campestris Luzula pilosa Juncus articulatus Juncus tenuis Juncus inflexus Juncus conglomeratus Junus bufonius

Gynmosperms - Native or naturalised Coniferous trees and shrubs Taxaceae Yew

Taxus baccata

Pinaceae Larches Pine Spruces

Larix dedcidua agg. Pinus sylvestris agg. Picea abies agg.

Cupressaceae Leyland or Lawson Cypress

Cypress agg.

Checklist of Pteridophyta - Ferns and allies Dryopteridacae Male Fern Scaly Male Fern Broad Buckler Fern Hard Shield Fern Soft Shield Fern Thelypteridaceae Lemon-Scented Fern Woodsiacae Lady Fern Dennstaedtiacae Bracken Blechnacae Hard Fern Polypodiacae Common Polypody Ophioglossacea Adder's-tongue Aspleniacae Hart's Tongue Fern Wall Rue Maidenhair Spleenwort Black Spleenwort Rustyback

Dryopteris filix-mas Dryopteris affinis Dryopteris dilatata Polystichum aculeatum Polystichum setiferum Oreopteris limbosperma Athyrium filix-femina Pteridium aquilinum Blechnum spicant Polypodium vulgare Ophioglossum vulgatum Phyllitis scolopendrium Asplenium ruta-muraria Asplenium trichomanes Asplenium adiantum-nigrum Ceterach officinarum

Checklist of Equisetaceae – Horsetails Marsh Horsetail Field Horsetail

Equisetum palustre Equisetum arvense

1.2 Checklist of Non-vascular Plants Bryophytes – Mosses and Liverworts Taxonomy and nomenclature from Blockeel et al. A new checklist of the bryophytes of Britian and Ireland, 2020. Journal of Bryology- BBS. Local status - c= commonly found. f = frequently found. u = uncommon or rare with few sites or occurrences National status - *** Rare or uncommon - a Doward speciality. # = Difficult to identity in field.

Bryophyta - Mosses - Species recorded since 1970 on the Doward. - Listed by order in BBS sequence. Sphagnales Blunt-leaved Bogmoss Lustrous Bog-moss

Sphagnum palustre Sphagnum subnitens

1.06 1.20

u u

Tetraphidales Pellucid Four-tooth Moss

Tetraphis pellucida

4.01

Polytrichales Common Smoothcap Dwarf Haircap Wood Hair-moss Juniper Haircap

Atrichum undulatum Pogonatum aloides Polytrichum formosum Polytrichum juniperinum

6.03 9.02 10.02 10.06

c u f u

Encalyptales Spiral Extinguisher-moss Common Extinguisher-moss

Encalypta streptocarpa Encalypta vulgaris

14.01 14.04

Funariales Bonfire Moss

Funaria hygrometrica

15.01

Dicranales Bendy Ditrichum Slender Ditrichum Brittle Swan-neck Moss Dwarf Swan-neck-moss Rusty Swan-neck Moss Heath Star-moss Large/Smaller White-moss Variable Forklet-moss Silky Forklet-moss Green Pocket-moss Petty Pocket-moss Narrow-leaved Pocket-moss Short-leaved Pocket-moss Lesser Pocket-moss Flatfoot Pocket-moss Slender Pocket-moss Common/great Pocket-moss Rock Pocket-moss Maidenhair Pocket-moss Broom Fork-moss Greater Fork-moss Common Pincushion Taper-leaved Earth-moss Redshank

Flexitrichum flexicaule Flexitrichum (Ditrichum) gracile Campylopus fragilis Campylopus pyriformis Campylophus flexuosus Campylophus introflexus Leucobryum glaucum Dicranella varia Dicranella heteromalla Fissidens viridulus Fissidens pusillus Fissidens gracilifolius Fissidens incurvus Fissidens bryoides Fissidens crassipes Fissidens exilis Fissidens taxifolius Fissidens dubius Fissidens adianthoides Dicranum scoparium Dicranum majus Dicranoweisia cirrata Pleuridium acuminatum sl. Ceratodon purpureus

21.01 21.20 26.04 26.05 26.06 26.11 27.01 31.05 31.10 32.01 32.03 32.04 32.05 32.06 32.09 32.11 32.15 32.16 32.17 34.04 34.06 39.01 46.01 50.01

u f u

Pottiales Whorled Tufa Moss Small-mouthed Beardless-moss Green-tufted Stubble-moss Variable Crisp-moss Curly Crisp-moss

Eucladium verticillatum Weissia brachycarpa Weissia controversa Trichostomum brachydontium Trichostomum crispulum

53.01 54.06 54.01 55.01 55.02

f u u u f

u f

u u c

u u

f u u

c f u

f u u u

note 1.

Neat Crisp-moss Frizzled Crisp-moss Slender Stubble-moss Luisier's Tufa-moss Blunt-leaved Tufa Moss Verdigris Tufa-moss Hornschuch's Beard-moss Revolute Beard Moss Red Beard-moss Lesser Bird's-claw Beard-moss Bird's-claw Beard-moss Wavy Beard-moss False Beard-moss Dusky Beard-moss Olive Beard-moss Nicholson's Beard-moss Soft-tufted Beard-moss Cylindrical Beard-moss Rigid Beard-moss Aloe Moss Bordered Screw-moss Wall Screw-moss Common Pottia Cuspidate Earth-moss Stanford Screw-moss Great Hairy Screw-moss Intermediate Screw-moss Small Hairy Screw-moss Marble Screw-moss Water Screw-moss Smaller Lattice-moss

Tortella nitida Tortella tortuosa Gyroweisa tenuis Gymnostomum viridulum Gymnostomum calcareum Gymnostumum aeruginosum Pseudocrossidium hornschuchianum Pseudocrossidium revolutum Bryoerythrophyllum recurvirostrum Streblotrichum (Barbula) convoluta Barbula unguiculata Didymodon sinuosus Didymodon fallax Didymodon luridus Didymodon tophaceus Didymodon nicholsonii Didymodon vinealis Didymodon insulanus Didymodon rigidulus Aloina aloides Tortula marginata Tortula muralis Tortula truncata Tortula (Phascum) acaulon Hennediella stanfordensis Syntrichia (Tortula) ruralis Syntrichia montana Syntrichia laeviphila Syntrichia papillosa Syntrichia latifolia Cinclidotus fontinaloides

57.02 57.05 61.01 62.01 62.02 62.03 67.01 67.02 68.01 70.01 71.01 72.01 72.04 72.06 72.07 72.08 72.09 72.10 72.13 76.03 77.07 77.10 77.19 77.21 79.01 82.01 82.04 82.07 82.08 82.09 83.01

u f u u

Gimmiales Thickpoint Grimmia Elegant Grimmia Grey-cushioned Grimmia Hair-pointed Grimmia

Schistidium crassipilum Schistidium elegantulum Grimmia pulvinata Grimmia trichophylla

92.17 92.18 93.16 93.20

Bryales Silver Moss Tall-clustered Thread-moss Bicoloured Bryum Wall Thread-moss Capillary Thread-moss Crimson-tuber thread-moss Marsh Bryum Pink-fruited Thread-moss Pale-glaucous Thread-moss Forest Star Bordered Thyme-moss Starry Thyme-moss Dotted Thyme-moss Woodsy Thyme-moss Many-fruited Thyme-moss Hart's-tongue Thyme moss Long-beaked Thyme-moss

Bryum argenteum Bryum pallens Bryum dichotomum Bryum radiculosum Bryum capillare Bryum rubens Bryum pseudotriquetrum Pohlia melanodon Pohlia wahlenbergii Mnium hornum Mnium marginatum Mnium stellare Rhizomnium punctatum Plagiomnium cuspidatum Plagiomnium affine Plagiomnium undulatum Plagiomnium rostrata

110.05 110.06 110.09 110.12 110.42 110.52 110.48 113.18 113.19 115.01 115.05 115.06 117.01 118.01 118.02 118.06 118.07

c u f u

Orthotrichales Green Yoke Moss Lesser Yoke-moss Stirton's Yoke-moss Wood Bristle-moss Lyell's Bristle moss

Zygodon viridissimus Zygodon conoideus Zygodon stirtonii Lewinskya (Orthotrichum) affinis Pulvigera (Orthothrichum) lyellii

120.01 120.04 120.02 123.03 125.01

u u

u u u c c f

f f u u

u c f

u c f u u u c u u u u

u ***

c u

c f u u u

c u u

u f f

c f

u c u

River Bristle-moss Elegant Bristle-moss Straw Bristle-moss Anomalous Bristle-moss White-tipped Bristle-moss Bruch’s Pincisshion Cape Thread-moss Drumsticks

Orthotrichum rivulare Orthotrichum pulchellum Orthotricum stramineum Orthotrichum anomalum Orthotrichum diaphanum Ulota bruchii Orthodontium lineare Aulacomnium androgynum

122.05 122.03 122.16 122.01 122.07 126.04 128.01 131.03

Hypnales Starry Feather-moss Dull-starry Feather-moss Greater Water-moss Dentated Silk-moss Juicy Silk-moss Woodsy Silk-moss Elegant Silk-moss Fern-leaved Hook-moss Curled Hook-moss Creeping Feather-moss Tiny Feather-moss Chalk Feather-moss Sickle-leaved Hook-moss Many-fruited Leskea Tamarisk Moss Neat Feather Moss Lesser-striate Feather-moss Common Striated Feather-moss Wall Feather-moss Clustered Feather-moss Tender Feather-moss Hair-pointed Feather-moss Beech Feather-moss Dwarf Feather Moss Swartz's Feather-moss Twist-tip Feather-moss Common Feather-moss Matted Feather-moss White Feather-moss Streaky Feather-moss Rough-stalked Feather-moss Tufted Feather-moss Velvet Feather-moss Silky Wall Feather-moss Yellow Feather-moss Cypress-leaved Plait-moss Mamillate Plait-moss Heath Plait-moss Depressed Feather-moss Flat-brochade Moss Pointed Spear-moss Lindberg’s Plait-moss Springy Turf Moss Little Shaggy Moss Big Shaggy Moss Short-beaked Wood-moss Lateral Cryphaea Flat Neckera Crisped Neckera Blunt Feather-moss Fox-tail Feather-moss Mouse-tail moss Larger Mouse-tail Moss

Campylium stellatum Campylium protensum Fontinalis antipyretica Plagiothecium denticulatum Plagiothecium succulentum Plagiothecium nemorale Pseudotaxiphyllum elegans Cratoneuron filicinum Palustriella commutata Amblystegium serpens Serpoleskea confervoides Campylophyllopsis calcarea Sanionia uncinata Leskea polycarpa Thuidium tamariscinum Pseudoscleropodium purum Plasteurhychium striatulum Eurhynchium striatum Rhynchostegium murale Rhychostegium confertum Rhychostegiella tenella Cirriphyllum piliferum Cirriphyllum crassinervium Microeurychium pumilum Oxyrrhynchium hians Oxyrhychium schleicheri Kindbergia (Eurynchium) praelonga Sciuro-hypnum populeum Brachythecium albicans Brachythecium glareosum Brachythecium rutabulum Scleropodium cespitans Brachytheciastrum velutinum Homalothecium sericeum Homalothecium lutescens Hypnum supressiforme Hypnum andoi Hypnum jutlandicum Taxiphyllum wissgrillii Platygyrium repens Calliergonella cuspidata Calliergonella lindbergii Rhytidiadelphus squarrosus Rhytidiadelphus loreus Hylocomiadelphus triquetus Loeskeobryum brevirostre Cryhaea heteromala Neckera complanata Neckera crispa Homalia trichomanoides Thamnobryum alopecurum Isothecium myosuroides Isothecium alopecuroides

134.01

u u f

134.02 137.01 141.03 141.08 141.09 143.01 150.01 151.01 153.01 161.01 162.01 170.01 174.01 178.01 180.01 182.02 183.01 184.03 184.04 185.01 186.01 186.02 187.01 188.01 188.02 189.01 190.04 191.01 191.03 191.06 192.01 194.01 195.01 195.02 196.01 196.02 196.04 197.01 198.01 201.01 201.02 210.01 210.03 211.01 212.01 218.01 223.04

223.03 224.01 225.01 229.01 229.03

f f f f c u u

u f

u f u

u c u *** u *** u u c

c u *** c

u f f f u

f u u ***

c u

u f c

BBS

u c f f u

u c u

c u c u c c f u f c c

# note 2.

Comb Moss Rambling Tail-moss

Ctenidium molluscum Anomodon viticulosus

230.01 233.02

c c

162 spp Notes to above list: 1. All or Leucobryum glaucum records could in fact be L. juniperoideum as field determination is not easy. 2. This now includes Hypnum resupinatum, H. lacunosum due to recent taxonomic changes

Additional Moss records – Old records prior to 1970 or not reliably seen since. Listed in alphabetic order. AL – Rev. Augustin Ley ca. 1880 Aloina ambigua Anomodon longifolius Amphidium mougeotii Bartramia pomiformis Sciuro-hypnum plumosum Bryum cespiticium Bryum canariensis Bryum donianum Bryum kunzei Bryum torquescens Bryum turbinatum Campylophyllum chrysophyllum Climacium dendroides Dialytrichia mucronate Dichodontium pellucidum Dicranum fuscescens Dicranum spadiceum Dicranum scottianum Entosthodon fascicularis Entostodon obtusus Ephemerum stoloniferum Grimmia montana Grimmia orbicularis Hedwigia ciliata Heterocladium heteropterum Hygrohypnum luridum Hylocomium splendens Leptodictyum riparium Leucodon scuiroides Microbryum davallianum Orthotrichum tenellum Orthotrichum sprucei Physcomitrium pyriforme Philonotis fontana Platydictya jungermanniodes Rhychostegium ripariodes Pleurochaete squarrosa Pohlia proligera Pohlia nutans Polytrichum piliferum Pseudephemerum nitidum Nogopterium gracile Ptychomitrium polyphyllum Racomitrium fasciculare Racomitrium heterostichum Rhodobryum roseum Rhychostegiella teneriffae Rhytidium rugosum Schistidium apocarpum Scorpiurium circinatum Seligeria pusilla

BBS no. 76.04 216.01 28.02 106.02 190.05 110.49 110.04 110.41 110.50 110.45 110.32 152.04 147.01 66.01 29.01 34.04 34.05 34.12 16.04 16.06 65.05 93.05 93.17 95.01 228.01 159.01 214.01 155.01 221.01 78.02 122.05 122.06 17.01 107.05 139.01 184.01 64.01 113.12 113.04 10.05 47.01 22.01 88.01 94.04 94.09 111.01 185.04 216.01 92.05 181.01 87.01

AL 19thC, Symonds Yat. Status Unclear if ever on Doward side of Wye.

BBS AL AL AL AL

AL AL

Seligeria donniana Seligeria acutifolia Trichodon cylindricus Ulota crispa 55spp

87.06 28.02 48.01 126.07

BBS status unclear cf. U bruchii

Marchantiophyta - Liverworts Species recorded since 1970 - Listed by order in BBS sequence

Lunulariales Crescent-cup Liverwort Common Liverwort

Lunularia cruciata Marchantia polymorpha

4.01 5.01

f f

Marchantiales Great Scented Liverwort Hemisphaeric Liverwort Lesser Crystalwort

Conocephalum conicum Reboulia hemisphearica Riccia subbifurca sl.

9.01 6.01 10.12

u u u

Pelliales Endive Pellia Overleaf Pellia

Pellia endiviifolia Pellia epiphylla

13.03 13.01

f f

Fossombroniales Common Frillwort

Fossombronia pusilla sl

14.05

Metzgeriales Bluish Veilwort Whiskered Veilwort Forked Veilwort Downy Veilwort Greasewort Jagged Germanderwort

Metzgeria violacea Metzeria consanguinea Metzgeria furcata Metzgeria pubescens Aneura pinguis Riccardia chamedryfolia sl.

19.01 19.02 19.03 19.06 20.01 21.02

f f c u *** u u

Porellales Wall Scalewort Bitter Scalewort Even Scalewort Dilated Scalewort Minute Pouncewort Micheli’s Least Pouncewort Mackay's Pouncewort Fairy Beads

Porella platyphylla Porella arboris-vitae Radula complanata Frullania dilatata Myriocoleopsis minutissima Lejeunea cavifolia Marchesinia mackaii Microlejeunea ulicina

22.01 22.03 23.01 24.05 27.01 31.01 32.01 33.01

f f f c u u u *** f

Jungermanniales Creeping Fingerwort Bifid Crestwort Variable-leaved Crestwort Greater Featherwort Lesser Featherwort Common Pouchwort Mueller's Pouchwort Notched Pouchwort Scarce Notchwort Top Notchwort Dark-green Flapwort Two-horned Pincerwort Rustwort White Earwort Grove Earwort Heath Earwort

Lepidozia reptans Lophocolea bidentata Lophocolea heterophylla Plagiochila asplenoides Plagiochila porelloides Calypogeia fissa Calypogeia muelleriana Calypogeia arguta Mesoptychia badensis Mesoptychia turbinata Jungermannia atrovirens Cephalozia bicuspidata Cephalozia curvifolia Diplophyllum albicans Scapania nemorea Scapania irrigua

40.01 44.01 44.03 48.03 48.02 54.01 54.02 54.08 57.07 57.08 60.01 71.01 71.02 92.01 94.11 94.12

f c f c c c u c u u u f u c f u

R. multifida

Liverworts continued – Old Records prior to 1970 or not reliably seen since. Listed in alphabetic order. Cololejeunea rosettiana Frullania tamarisci Lophosia excisa Marsupella emmarginata Metzgeria conjugata Orthocaulis attenuatus Plagiochila Britannica Riccia glauca Riccia sorocarpa Scapania compacta

26.03 24.01 87.05 67.02 19.04 76.03 48.04 10.09 10.07 94.01

Anthocerotophyta - Hornworts Anthoceros punctatus 1.01

Algae This group is largely outside the scope of this publication as most species require advance microscopic techniques for determination. However, some species are important for various ecological reasons and can be noticeable when growing on various land and aquatic substrates. Trentapohlia aurea, a filamentous green alga, is a common sight common on trees in The Doward and noticeable due its bright orange colouration caused by the production of carotinoid chemicals. This and the genus Trebouxia are from the class of Chlorophyta are the partner alga or photobiont of many Lichen species such as Xanthoria parietina. Four other species of algae from Phylum Chlorophyta are also listed by NBN from 1977. Gongrosira incrustans, Paulschulzia, Cladophora glomerata, Ulva flexuosa. Phylum Charophyta (aquatic Stoneworts) reported by NBN just outside Doward at Dixton, Monmouth with two species Chara vulgaris and C. globularis listed (1891 and 1944 respectively and again, but not to species level, in 1979).

1.3. Checklist of Fungi and Lichens The list below is generated from a range of ‘informal’ observations by various recorders on the Doward rather than a set of formal records. It should therefore be treated as for guidance purposes only. The family classification and sequence are taken from that used in Sterry, P & Hughes B. (2009). Collins Complete Guide to British Mushrooms & Toadstools. It has NOT been checked with the latest version from the British Mycological Society. Most families of larger fungi are in the Phylum Basidomycota although some are in Ascomycota. Lichens are treated separately. *** denotes a nationally uncommon or notable species. Boletacea Bay Bolete type Cep Devil's Bolete Lurid Bolete Scarletina Bolete type Bitter Beech Bolete Rooting Bolete Orange Oak Bolete Brown Birch Bolete Larch Bolete Hollow Bolete n/a Bovine Bolete n/a Matt Bolete Russulaceae Blackening Brittlegill Ochre Brittlegill Charcoal Burner Golden Brittlegill Birch Brittlegill Crowded Brittlegill Fragile Brittlegill Milk White Brittlegill Scarlet Brittlegill Bare-toothed Brittlegill Beech Milkcap Mild Milkcap Fiery Milkcap Oakbug Milkcap Tawny Milkcap Woolly Milkcap Yellowdrop Milkcap Pale Milkcap Birch Milkcap Agaricaceae George’s Mushroom Yellow Stainer Wood Mushroom n/a Freckled Dapperling Stinking Dapperling Shield Dapperling Parasol Slender Parasol n/a Hygrophoraceae Snowy Waxcap Spangle Waxcap

Boletus badius Boletus edulis Boletus satanas *** Boletus luridus Boletus luridiformis/erythropus ag Boletus calopus Boletus radicans Leccinum aurantiacum Leccinum scabrum Suillus grevillea Suillus cavipes *** Suillus tridentinus Suillus bovinus Xerocomellus cisalpinus Xerocomellus pruinatus Russula nigricans Russula ochroleuca or ag Russula cyanoxantha Russula risigallina Russula betularum Russula densifolia Russula fragilis Russula delica Russula pseudointegra Russula vesca Lactarius blennius Lactarius subdulcis Lactarius pyrogalus Lactarius quietus lactarius fulvissimus Lacterius torminosus Lactarius chrysorrheus Lactarius pallidus Lactarius tabidus Calocybe gambosa Agaricus xanthodermus Agaricus silvicola Lepiota hystrix Lepiota aspera Lepiota cristata Lepiota clypeolaria Macrolepiota procera Macrolepiota mastoides Cystolepiota hetieri Hygrocybe virginea Hygrocybe insipida

Scarlet waxcap Butter Waxcap Blackening Waxcap Honey Waxcap Parrot Waxcap Meadow Waxcap Cedarwood Waxcap Hygrophoropsidaceae Ivory Woodwax Cantharellaceae False Chanterelle Trumpet Chanterelle Chanterelle Coprinaceae Glistening Inkcap Shaggy Inkcap Pleated Inkcap Fairy Inkcap Magpie Inkcap Common Inkcap Psathyrellaceae Stump Brittlestem Conical Brittlestem Pale Brittlestem Brown Mottlegill Petticoat Brittlestem Weeping Widow Tricholomataceae Common Bonnet Drab Bonnet Burgundydrop Bonnet Saffrondrop Bonnet Lilac Bonnet Ivory Bonnet Milking Bonnet Blackedge Bonnet Pink Bonnet n/a Snapping Bonnet n/a Russet Toughshank Whitelaced Shank Sulphur Knight Blue spot Knight Soapy Knight n/a Clustered Domecap Club Foot Velvet Shank The Deceiver Amythyst Deceiver Wood Blewit Tawny Funnel Porcelain Fungus Trooping Funnel Common Funnel Ivory/Frosty Funnel Honey Fungus Plums and Custard Common Cavalier Twig Parachute Collared Parachute Orange Mosscap

Hygrocybe coccinea Hygrocybe ceracea Hygrocybe conica Hygrocybe reidii Hygrocybe psittacina Hygrocybe pratensis Hygrocybe russocoriacea Hygrophorus eburneus Hygrophoropsis aurantiaca Cantharellus tubaeformis Cantharellus cibarius Coprinus micacea Coprinus comatus Parasolus (Coprinus) plicatilis Coprinellus disseminatus Coprinus picacea Coprinus atramentaria Psathyrella piluliformis Psathyrella conophilus Psathyrella candoleana Panaeolina foenisecii Psathyrella artemisia Lacrymaria lacrymabunda Mycena galericulata Mycene aetites sl. Mycena haematopus Mycena crocata Mycena pura Mycena flavoalba Mycena galopus Mycena pelianthina Mycena rosella Mycena stipata Mycena vitilis Hydropus subalpinus Collybia (Gymnopus) dryophila Megacollybia platyphylla Tricholoma sulphureum Tricholoma columbretta Tricholoma saponaceum Tricholoma ustale Lyophyllum decastes Ampulloclitocybe clavipes Flamulina velutipes Laccaria laccata Laccaria amethystina Lepista nuda Lepista flaccida Oudemansiella mucida Clitocybe geotrupa Clitocybe gibba Clitocybe dealabata/phyllophila Armillaria mellea Tricholomopsis rutilans Melanoleuca polioleuca Marasmiellus ramealis Marasmiellus rotula Rickenella fibula

White Domecap Clustered Domecap Strophariaceae Blue Roundhead Verdigris Roundhead Sulfur Tuft Shaggy Scalycap Golden Scalycap Petticoat Mottlegill Cortinariaceae Bulbous/Star Fibrecap type White/Lilac Fibrecap Scaly Fibrecap n/a n/a Common Rustgill n/a Variable Webcap Dappled Webcap Stocking Webcap n/a n/a Poisonpie Sweet Poisonpie Funeral Bell Moss Bell Entolomataceae Wood Pinkgill Pimple Pinkgill n/a n/a The Miller Bolbitaceae Yellow Fieldcap Velvet Toughshank Pluteaceae Deer Shield n/a Paxillaceae Brown Rollrim Auriscalpiaceae Aniseed Cockleshell Amanitaceae Snakeskin Grisette Rooting Shank Fly Agaric Grey-spotted Amanita Death cap False Deathcap Tawny Grisette Psathyrellaceae n/a n/a Lentinaceae Oyster Mushroom Pale Oyster Crepidotaceae Variable Oysterling n/a n/a Scurfy Twiglet Felted Twiglet

Lyophyllum connatum Lyophyllum decastes Stropharia caerulea Stropharia aeriginosa Hypholoma fasciculare Pholiota squarrosa Pholiota aurivella Panaeolus papilionaceus Inocybe napipes/asterospora Inocybe geophylla Inocybe hystrix Inocybe cincinnata var. major Inocybe patouillardii Gymnopilus penetrans Conocybe tenera Cortinarius anomalus Cortinarius bolaris Cortinarius torvus Cortinarius olivaceofuscus Cortinarius elegantissimus Hebeloma crustuliniforme Heboloma sacchariolens Galerina marginata (was G. autumnalis) Galerina hypnorum Entoloma rhodopolium Entoloma hebes Entoloma incanum Entoloma conferendum Clitopilus prunulus Bolbitius titubans Kuehneromyces mutabilis Pluteus cervinus Pluteus salicinus Paxillus involutus Lentinellus cochleatus Amanita ceciliae Xerula radicata Amanita muscaria Amanita excelsa Amanita phalloides Amanita citrina Amanita fulva Psathyrella candolleana Psathyrella leucotephra Pleurotus ostreatus Pleurotus pulmonarius Crepidotus variabilis Crepidotus lundellii Crepidotus cesatii Tubaria furfuracea Tuberia conspersa

Xylariacae Dead Man's Fingers Dead Moll's Finger Candlesnuff Beech Woodwart Hazel Woodwart Cramp Balls Heliotiales insertae sedis Green Elfcup Helotiacea Beech Jellydisc Purple Jellydisc Common Grey Disco Lemon Disco Bulgariaceae Black Bulgar Auriculariaceae Jelly (Jew's) Ear Witches Butter Leotiacea Jellybaby Clavariaceae Yellow Club Meadow Coral Crested Coral White Spindles Slender Club Coral sp Upright Coral Clavariadelphaceae Giant Club Phallaceae Stinkhorn Dacrymycetaceae Yellow Stagshorn Small Stagshorn Yellow Brain Orange Brain Helvellacea Elfin Saddle Common White Saddle Orange Peel Fungus Geoglossaceae Earthtongue Earthtongue Sarcoscyphaceae Scarlet Elfcups Geastraceae Collared Eathstar Nidulariacea Fluted Bird's Nest Lycoperdacea Common Puffball Stump Puffball Spiny Puffball Meadow/grassland Puffball Sclerodermataceae Common Earthball Polyporacaea Dryad's Saddle Blackfoot Polypore Tuberous Polypore Winter Polypore

Xylaria polymorpha Xylaria longipes Xylaria hypoxxylon Hypoxylon fragiforme Hypoxylon fuscum Daldinia concentrica Chlorociboria aeruginascens Neobulgaria pura Ascocoryne sarcoides Mollisia cinerea Bisporella citrina Bulgaria inquinans Auricularia auricula-judae Exidia glandulosa Leotia lubrica Clavulinopsis helvola/ luteoalba Clavulinopsis corniculata Clavulina coralloides Clavaria fragilis Macrotyphula juncea Ramaria spp poss. curta or decurrens Ramaria stricta Clavariadelphus pistillaris Phallus impudicus Calocera viscosa Calocera cornea Tremella mesenterica Tremella aurantia Helvella lacunosa Helvella crispa Aleuria aurantia Geoglossum cookeanum ag Geoglossum viride or olivacum Sarcoscypha austraca Geastrum triplex Cyathus olla Lycoperdon perlatum Lycoperdon pyriforme Lycoperdon echinatum Lycoperdon pratense/lividum Scleroderma citrinum Polyporus squamosus Polyporus leptocephalus/varius Polyporus tuberaster Polyporus brumalis

Ganodermateae Artists Bracket n/a Turkeytail Lacquered Bracket Purplepore Bracket Hazel Bracket Fistulinaceae Beefsteak Fungus Hymenochaetaceae Oak Bracket Hydnaceae Wood hedgehog Terracota Hedgehog Coriolaceae Birch Polypore Fomitopsidaceae Powderpuff Bracket Resupinates Wrinkled Crust Cobalt Crust Bleeding Broadleaf Crust Bleeding Oak Crust n/a Silverleaf Fungus Morchellacea Bleach Cup Semifree Morel

Ganoderma applanatum Ganoderma australe Trametes versicola Ganoderma lucidum Trichaptum abietinum Skeletocutis nivea Fistulina hepatica Inonotus dryadeus Hydnum repandum Hydnum rufescens Piptoporus betulinus Postia ptychogaster Phlebia radiata Terana caerulea Stereum rugosum Stereum gausapatum Stereum hirsutum Chondrostereum purpureum Disciota venosa Morchella semilibera

Rust on Pendulous Sedge

Puccinia caricina var ribesii-pendulae

Tar spot

Rhytisma acerinum

A checklist of Lichens A substantial number of species are listed on NBN and in other sources but due to recent taxonomic changes a detailed review would be required to provide an updated list. The following list is a set of common species known on the Doward using old nomenclature and are all unverified records so should be treated as such. Arthonia radiata Graphis scripta ag. Verrucaria nigrescens ag. Caloplaca citrina ag. Caloplaca flavescens Lecidella elaeochroma Lecanora expallens ag. Lecanora chlarotera Lecanora albella sl. Lecanora sp prob dispersa sl. Xanthoria parietina Physcia tennella-adscendens Physcia aipolia Parmelia sulcata Punctelia subrudecta Hypnogymia physodes Evernia prunastri Ramalina fastigiata Ramalina farinacea Usnea subfloridana Usnea florida Peltigera membranacea ag. Cladonia coniocraea ag. Hypotrachyna revoluta Collema auriforme Protoblastenia rupestris

Supplementary Notes 1.

Endemics plants of the Doward

The two taxa groups where the local speciation is deemed to have occurred are: 1. Whitebeams: In a contribution to Herefordshire Wildlife Trust’s March 2015 Flycatcher magazine Dave Green asserted that the Doward now has 14 species of Sorbus of which 5 are endemic to the Lower Wye Valley with two being found only on the Doward. He also described the Doward is the second most important site in the British Isles for its Sorbus diversity after the Avon Gorge. Studies by Tim Rich of Cardiff University aided by Dave Green and others describe an ongoing evolutionary process for this genus in this area of the Wye Valley. Hybridisation and then back-crossing has taken place as revealed by genetic study techniques such as flow cytometry and with virtually all Whitebeam trees on the Doward having now been examined and mapped. It seems that subtly different habitats in closed canopy woodland and woodland edges on rocky ground have led to a range of new species being able to survive. The full explanation of whitebeams on the Doward is given in scientific publications. In summary, there are three well known and well distributed species of Sorbus in UK, the Common Whitebeam Sorbus aria, the Rowan Sorbus aucuparia and the Wild Service Tree Sorbus torminalis and all occur on the Doward in reasonable numbers. Two species are considered endemic to the Doward: Sorbus herefordensis - Herefordshire Whitebeam, a woodland canopy species found in Miner’s Rest reserve and other central areas of Great Doward. Sorbus greenii - Green’s Whitebeam, a woodland edge species found particularly around old quarries in the west and north of Great Doward. Nine further species occurring on the Doward are considered near-endemic as they also occur in nearby areas of the Lower Wye Valley: Sorbus evansii Evans Whitebeam Cliff edge or woodland fringe Sorbus eminentiformis Doward Whitebeam Open high woodland Sorbus rupicola Rock Whitebeam Cliff edge Sorbus saxicola Symonds Yat Whitebeam. Cliff edge Sorbus porrigentiformis Grey-leaved Whitebeam Rock faces Sorbus anglica English Whitebeam Rock faces Sorbus eminens Round-leaved Whitebeam Cliff face and woodland fringe Sorbus thuringiacia unnamed Cliff face (= S. aria x S. aucuparia) Sorbus x decipiens Wye Whitebeam (old name) Open high woodland (= S. aria x S.torminalis previously called S. x tomentella) Also not recorded on Doward but found nearby on Coldhams Rocks: Sorbus parviloba Ship Rock Whitebeam.

One of the rare cliff-edge Sorbus species at Seven Sisters.

2. Hawkweeds – Hieracium species: To most people Hawkweed is just another yellow Dandelion-like plant but to people who like to dig deeper into the taxonomy and morphology of the family Aceracea (or Compositae to the old school) there is a range of interesting micro-species. One of them, Hieracium subplanifolium, is endemic to the Doward but known only from a small patch. It is unclear how many other similar Hawkweed species occur here, but this this aggregate can be numerous especially around rocky areas such as in Lords Wood Quarry NR and on certain other thin-soiled track verges.

A Hawkweed Hieracium sp in Lord’s Wood Quarry