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Euphorbia subg. Chamaesyce

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Euphorbia in Southern Africa

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Abstract

Bisexual (sometimes unisexual), annual herbs, succulent shrubs with slender cylindrical stem and branches (rarely with spines developing from tips of branches), rarely geophytes or small trees. Leaves alternate or opposite, ovate to lanceolate and sometimes reduced to minute caducous rudiments, rarely arising on raised tubercles (these irregularly arranged along branches); small stipules often present and glandular or filiform. Synflorescences in axils of leaves towards apices of stem and branches or terminal on peduncle, peduncles usually simple, often with alternating or opposite bracts often differently shaped from leaves and sometimes larger just beneath cyathium, further peduncles with terminal cyathia rarely developing from axils of uppermost bracts. Cyathia bisexual or unisexual (then males larger than females); glands 4–5, usually elliptic and flat with entire, crenulate or few-toothed outer margins, sometimes with petaloid appendages; male florets with glabrous or pubescent pedicels. Capsule obtusely 3-lobed, smooth, glabrous to pubescent, 2–10 mm diam., sessile to exserted, dehiscing explosively. Seeds 3 per capsule, 4-angled to ellipsoidal or oblong, smooth to tuberculate, with or without caruncle.

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References

  • Aiton, W. (1789). Hortus Kewensis. Ed. 1, 2. : G. Nicol.

    Google Scholar 

  • Alpinus, P. (1735). Historiae naturalis Aegypti pars secunda, sive, de Plantis Aegypti Liber. Cum Observationibus & Notis Joannis Veslingii. Leiden: G. Potvliet.

    Google Scholar 

  • Anthony, H. J. (1949). A succulent enthusiast in Nyasaland. Journal of the New York Botanical Garden, 50, 97–103.

    Google Scholar 

  • APG IV. (2016). An update of the Angiosperm Phylogeny Group classification for the orders and families of flowering plants: APG IV. Botanical Journal of the Linnean Society, 181, 1–20.

    Article  Google Scholar 

  • Arakaki, M., Christin, P.-A., Nyffeler, R., Lendel, A., Eggli, U., Ogburn, R. M., Spriggs, E., Moore, M. J., & Edwards, E. J. (2011). Contemporaneous and recent radiations of the world’s major succulent plant lineages. Proceedings of the National Academy of Sciences, 108, 8379–8384.

    Article  CAS  Google Scholar 

  • Archer, R. H. (1998). Euphorbia leistneri (Euphorbiaceae), a new species from the Kaokoveld (Namibia). South African Journal of Botany, 64, 258–260.

    Article  Google Scholar 

  • Archer, R. H., & Carter, S. (2001). Euphorbia venteri. Flowering Plants of Africa, 57, 86–90.

    Google Scholar 

  • Axelrod, D. J. (1983). Biogeography of oaks in the Arcto-Tertiary Province. Annals of the Missouri Botanical Garden, 70, 629–657.

    Article  Google Scholar 

  • Axelrod, D. J., & Raven, P. H. (1978). Late Cretaceous and Tertiary vegetation history of Africa. In M. J. A. Werger (Ed.), Biogeography & ecology of Southern Africa (pp. 77–130). W. Junk.

    Chapter  Google Scholar 

  • Baker, J.G. 1869. Euphorbia tetragona, Haw. In W.W. Saunders (Ed.) Refugium Botanicum 1: t. 39.

    Google Scholar 

  • Bally, P. R. O. (1961). THE GENUS MONADENIUM. With descriptions of 18 new species 32 plates in natural size and 88 figures. Benteli Publishers.

    Google Scholar 

  • Bally, P. R. O. (1963). Miscellaneous notes on the flora of Tropical East Africa including description of new taxa. 8–15. Candollea, 18, 335–357.

    Google Scholar 

  • Bauer, A. M., Günther, R., & Klippel, M. (1995). The herpetological contributions of Wilhelm Peters (1815–1883). Society for the Study of Amphibians and Reptiles.

    Google Scholar 

  • Berger, A. (1899). Zwei verwechselte Euphorbien. Monatsschr Kakteenk, 9, 88–92.

    Google Scholar 

  • Berger, A. (1902). Eine neue Euphorbia. Monatsschr Kakteenk, 12, 39–40.

    Google Scholar 

  • Berger, A. (1906). Sukkulente Euphorbien. E. Ulmer.

    Google Scholar 

  • Berger, A. (1910). Einige neue afrikanische Sukkulenten. Botanische Jahrbücher für Systematik, 45, 223–233.

    Google Scholar 

  • Bergius, P. J. (1767). Descriptiones Plantarum ex Capite Bonae Spei. Laurentius Salvius.

    Google Scholar 

  • Boissier, P. E. (1860). Centuria Euphorbiarum. Paris: B. Hermann, Leipzig & J.-B. Baillière.

    Google Scholar 

  • Boissier, E. (1862). Euphorbieae. In A. P. De Candolle (Ed.), Prodromus 15 (2): 3–188. Paris: V. Masson & Sons.

    Google Scholar 

  • Brown, R. (1814). General remarks, geographical and systematical, on the botany of Terra Australis. In M. Flinders (Ed.), A voyage to Terra Australis: Undertaken for the purpose of completing the discovery of that vast country (Appendix III) (Vol. 2, pp. 533–613). G. W. Bulmer.

    Google Scholar 

  • Brown, N. E. (1897). Euphorbia grandicornis, Goebel. Hooker’s Icones Plantarum, 26, t. 2531, 2532.

    Google Scholar 

  • Brown, N. E. (1911–12). Euphorbia. In W. T. Thiselton-Dyer (Ed.), Flora Tropical Africa 5 (2): 470–603. London: L. Reeve & Co.

    Google Scholar 

  • Brown, N. E. (1915). Euphorbia. In W. T. Thiselton-Dyer (Ed.), Flora Capensis 5 (2): 222–375. London: L. Reeve & Co.

    Google Scholar 

  • Bruyns, P. V. (1992a). Notes on Euphorbia monteiroi. Aloe, 29, 36–38.

    Google Scholar 

  • Bruyns, P. V. (1992b). Notes on African plants. Euphorbiaceae. Notes on Euphorbia species from the Northwestern Cape. Bothalia, 22, 37–51.

    Article  Google Scholar 

  • Bruyns, P. V. (2000). Euphorbia. In P. Goldblatt & J. Manning (Eds.), Cape Plants. Strelitzia 9: 455–458. Pretoria: South African National Biodiversity Institute.

    Google Scholar 

  • Bruyns, P. V. (2012). Nomenclature and typification of southern African species of Euphorbia. Bothalia, 42, 217–245.

    Article  Google Scholar 

  • Bruyns, P. V. (2013). Euphorbiaceae. In D. A. Snijman (Ed.), Plants of the Greater Cape Floristic Region. 2: The Extra Cape Flora. Strelitzia 30: 371–379. Pretoria: South African National Biodiversity Institute.

    Google Scholar 

  • Bruyns, P. V. (2018). New taxa in Euphorbia (Euphorbiaceae) in southern Africa. Haseltonia, 25, 30–56.

    Google Scholar 

  • Bruyns, P. V., & Berry, P. E. (2019). The nomenclature and application of the names Euphorbia candelabrum Welw. and Euphorbia ingens E. Mey ex Boiss. in tropical Africa. Taxon, 68, 828–838.

    Article  Google Scholar 

  • Bruyns, P. V., Mapaya, R. J., & Hedderson, T. (2006). A new subgeneric classification for Euphorbia (Euphorbiaceae) in southern Africa based on ITS and psbA-trnH sequence data. Taxon, 55, 397–420.

    Article  Google Scholar 

  • Bruyns, P. V., Al-Farsi, A., & Hedderson, T. (2010). Phylogenetic relationships of Caralluma R. Br. (Apocynaceae). Taxon, 59, 1031–1043.

    Article  Google Scholar 

  • Bruyns, P. V., Klak, C., & Hanáček, P. (2011). Age and diversity in Old World succulent species of Euphorbia (Euphorbiaceae). Taxon, 60, 1717–1733.

    Article  Google Scholar 

  • Bruyns, P. V., Oliveira-Neto, M., Melo-de-Pinna, F., & Klak, C. (2014). Phylogenetic relationships in the Didiereaceae with special reference to subfamily Portulacarioideae. Taxon, 63, 1053–1064.

    Article  Google Scholar 

  • Burman, J. (1737). Thesaurus Zeylanicus. Janssonius-Waesberg. & S. Schouten.

    Google Scholar 

  • Burman, J. (1738–9). Rariorum Africanarum Plantarum. Amsterdam: Boussière.

    Google Scholar 

  • Burrows, J. E., Burrows, S., Schmidt, E., Lötter, M., & Wilson, E. O. (2018). Trees and shrubs Mozambique. Print Matters Heritage.

    Google Scholar 

  • Carter, S. (1988). Euphorbia in Euphorbiaceae (Part 2). In R. M. Polhill (Ed.), Flora of tropical East Africa (pp. 409–533). Kew: Royal Botanic Gardens.

    Google Scholar 

  • Carter, S. (1993). Euphorbia, In: M. Thulin (Ed.), Flora of Somalia 1: 306–337. Kew: Royal Botanic Gardens.

    Google Scholar 

  • Carter, S. (1994). A preliminary classification of Euphorbia subgenus Euphorbia. Annals of the Missouri Botanical Garden, 81, 368–379.

    Article  Google Scholar 

  • Carter, S. (1999). New Euphorbia taxa in the Flora Zambesiaca area. Kew Bulletin, 54, 959–965.

    Article  Google Scholar 

  • Carter, S. (2002). Euphorbia. In U. Eggli (Ed.), Illustrated handbook of succulent plants: Dicotyledons (pp. 102–203). Springer.

    Google Scholar 

  • Carter, S. (2014). Typification of Euphorbia xylophylloides and Euphorbia polygona. Euphorbia World, 10, 18–20.

    Google Scholar 

  • Carter, S., & Leach, L. C. (2001). Tribe Euphorbieae. In G. V. Pope (Ed.), Flora Zambesiaca 9 (5): 339–465. Kew: Royal Botanic Gardens.

    Google Scholar 

  • Chapman, J. D. (1962). The vegetation of Mlanje Mountains, Nyasaland. The Government Press.

    Google Scholar 

  • Coe, M. J., & Skinner, J. D. (1993). Connections, disjunctions and endemism in the eastern and southern African mammal faunas. Transactions of the Royal Society of South Africa, 48, 233–255.

    Article  Google Scholar 

  • Coetzee, J. A. (1980). Tertiary environmental changes along the south-western African coast. Palaeontologia Africana, 23, 197–203.

    Google Scholar 

  • Commelijn, C. (1697). Horti Medici Amstelodamensis 1. : A. van Someren.

    Google Scholar 

  • Commelijn, C. (1703). Praeludia botanica. Lugduni Batavorum (Leiden). F. Haringh.

    Google Scholar 

  • Compton, R. H. (1929). The vegetation of the Karoo. Journal of the Botanical Society of South Africa, 15, 13–21.

    Google Scholar 

  • Compton, R. H. (1931). The Flora of the Whitehill district. Transactions of the Royal Society of South Africa, 19, 269–329.

    Article  Google Scholar 

  • Compton, R. H. (1976). The Flora of Swaziland (Journal of South African Botany, Supplementary Volume 11). Trustees of National Botanc Gardens.

    Google Scholar 

  • Corner, E. J. H. (1976). The Seeds of Dicotyledons. Cambridge University Press.

    Google Scholar 

  • Court, D. (1988). Euphorbia polycephala at Cranemere. Euphorbia Journal, 5, 39–42.

    Google Scholar 

  • Cracraft, J. (1992). The species of the birds-of-paradise (Paradisaeidae): Applying the phylogenetic species concept to a complex pattern of diversification. Cladistics, 8, 1–43.

    Article  PubMed  Google Scholar 

  • Croizat, L. (1932). A list of annotated observations on the remarks of Dr. K. von Poellnitz concerning A. Berger’s classification of succulent Euphorbiae. Cactus and Succulent Journal (US), 4, 291–293.

    Google Scholar 

  • Croizat, L. (1933). A list of annotated observations on the remarks of Dr. K. von Poellnitz concerning A. Berger’s classification of succulent Euphorbiae. Part III. Cactus and Succulent Journal (US), 4, 330–334.

    Google Scholar 

  • Croizat, L. (1934). De Euphorbio Antiquorum Atque Officinarum. A study of succulent Euphorbiae long known in cultivation. .

    Google Scholar 

  • Croizat, L. (1937). On the classification of Euphorbia. II. How should the cyathium be interpreted? Bulletin of the Torrey Botanical Club, 64, 523–536.

    Article  Google Scholar 

  • Croizat, L. (1945). “Euphorbia Esula” in North America. American Midland Naturalist, 33, 231–243.

    Article  Google Scholar 

  • Croizat, L. (1965). An introduction to the subgeneric classification of Euphorbia L., with stress on the South African and Malagasy species. I. Webbia, 20, 473–706.

    Article  Google Scholar 

  • Croizat, L. (1967). Two new semi-succulent Euphorbias from Venezuela: Euphorbia lutzenbergeriana and E. lagunillarum. Cactus and Succulent Journal (US), 39, 142–144.

    Google Scholar 

  • Croizat, L. (1972). An introduction to the subgeneric classification of Euphorbia L., with stress on the South African and Malagasy species III. Webbia, 27, 1–221.

    Article  Google Scholar 

  • Cronquist, A. (1988). The evolution and classification of flowering plants (2nd ed.). New York Botanical Garden.

    Google Scholar 

  • De Candolle, A. P. (1805). Plantarum Historia Succulentarum, Fasc. 27. Paris: Garnery.

    Google Scholar 

  • D’Isnard, A.-T. D. (1720). Etablissement d’un Genre de Plante appellé Euphorbe. Histoire de l’Academie Royale des Sciences. Avec les Mémoires de Mathematique & de Physique. Paris, 1720, 384–399.

    Google Scholar 

  • Desfontaines, R. L. (1829). Tableau de l’École de Botanique du Muséum d’Histoire Naturelle (3rd ed.). J.S. Chaudé.

    Google Scholar 

  • Dillenius, J. J. (1732). Hortus elthamensis 2. London.

    Google Scholar 

  • Dinter, M. K. (1921). Botanische Reisen in Deutsch-Südwest-Afrika. Feddes Repert. Spec. Nov. Regni Veg., Beih. Band 3.

    Google Scholar 

  • Dinter, M. K. (1923). Succulentenforschung in Südwestafrika. I. Teil. Feddes Repert. Spec. Nov. Regni Veg., Beih. Band 23.

    Google Scholar 

  • Dinter, M. K. (1928). Sukkulentenforschung in Südwestafrika. II. Teil. Feddes Repert. Spec. Nov. Regni Veg., Beih. Band 53.

    Google Scholar 

  • Dorsey, B. L., Haevermans, T., Aubriot, X., Morawetz, J. J., Riina, R., Steinmann, V. W., & Berry, P. E. (2013). Phylogenetics, morphological evolution, and classification of Euphorbia subgenus Euphorbia. Taxon, 62, 291–315.

    Article  Google Scholar 

  • Drège, J. F. (1843). Zwei planzengeographische Documente. Besondere Beigabe zur Flora 1843, Band II.

    Google Scholar 

  • Dressler, R. L. (1957). The genus Pedilanthus (Euphorbiaceae). Contributions from the Gray Herbarium, 182, 1–188.

    Google Scholar 

  • Dudley, C. O. (1997). The candelabra tree (Euphorbia ingens): A source of water for black rhinoceros in Liwonde National Park, Malawi. Koedoe, 40, 57–62.

    Article  Google Scholar 

  • Dyer, R. A. (1931). Notes on Euphorbia species of the Eastern Cape Province with descriptions of three new species. Records of the Albany Museum, 4, 64–110.

    Google Scholar 

  • Dyer, R. A. (1934a). Miscellaneous new species. Bulletin of Miscellaneous Information, 1934, 265–270.

    Google Scholar 

  • Dyer, R. A. (1934b). Euphorbia duseimata. Flowering Plants of South Africa, 14, t. 530.

    Google Scholar 

  • Dyer, R. A. (1935a). The seed germination of certain species of Euphorbia. South African Journal of Science, 32, 313–319.

    Google Scholar 

  • Dyer, R. A. (1935b). Note on the meloformia group of Euphorbia. Records of the Albany Museum, 4, 254–255.

    Google Scholar 

  • Dyer, R. A. (1937a). Euphorbia complexa. Flowering Plants of South Africa, 17, t. 643.

    Google Scholar 

  • Dyer, R. A. (1937b). Euphorbia tubiglans. Flowering Plants of South Africa, 17, t. 646.

    Google Scholar 

  • Dyer, R. A. (1938a). Euphorbia fasciculata Thunb. and E. schoenlandii Pax. South African Journal of Science, 35, 298–299.

    Google Scholar 

  • Dyer, R. A. (1938b). Euphorbia zoutpansbergensis. Flowering Plants of South Africa, 18, t. 715.

    Google Scholar 

  • Dyer, R. A. (1940a). Euphorbia schoenlandii. Flowering Plants of South Africa, 20, t. 772.

    Google Scholar 

  • Dyer, R. A. (1940b). Euphorbia obesa. Flowering Plants of South Africa, 20, t. 788.

    Google Scholar 

  • Dyer, R. A. (1949). Col. Robert Jacob Gordon’s contribution to South African Botany. South African Biological Society Pamphlet, 14, 1–20.

    Google Scholar 

  • Dyer, R. A. (1951). Newly described plants. Euphorbiaceae. Bothalia, 6, 221–227.

    Google Scholar 

  • Dyer, R. A. (1953). A new succulent Euphorbia from the Western Cape Province. Journal of South African Botany, 19, 135–136.

    Google Scholar 

  • Dyer, R. A. (1957). The classification of species of Euphorbia with stipular spines. Bulletin du Jardin Botanique de l’état, Bruxelles, 27, 487–493.

    Google Scholar 

  • Dyer, R. A. (1974a). Euphorbia triangularis. Flowering Plants of Africa, 43, t. 1687.

    Google Scholar 

  • Dyer, R. A. (1974b). Notes on African plants. Euphorbiaceae. A new species of Euphorbia. Bothalia, 11, 278–280.

    Article  Google Scholar 

  • Dyer, R. A. (1979). Looking back. Cactus and Succulent Journal (US), 51, 116–117, 119.

    Google Scholar 

  • Flores, G. (1970). Suggested origin of the Mozambique channel. Transactions of the Geological Society of South Africa, 73, 1–16.

    Google Scholar 

  • Fourie, S. P. (1987). An introduction to the succulent Euphorbias of the Transvaal, Part 2: Shrubs. Euphorbia Journal, 4, 48–68.

    Google Scholar 

  • Fourie, S. P. (1988). An introduction to the succulent Euphorbias of the Transvaal, Part 3: Dwarf shrubs I. Euphorbia Journal, 5, 83–93.

    Google Scholar 

  • Fourie, S. P. (1989). An introduction to the succulent Euphorbias of the Transvaal, Part 3: Dwarf Shrubs II. Euphorbia Journal, 6, 113–125.

    Google Scholar 

  • Frick, G. A. (1931). Euphorbia polycephala. Cactus and Succulent Journal (US), 3, 97–98.

    Google Scholar 

  • Frick, G. A. (1936). Euphorbia Dinteri Berger. Euphorbia Review, 2, 2–3.

    Google Scholar 

  • Frick, G. A. (1937). Euphorbia halleri Dtr. Cactus and Succulent Journal (US), 9, 39.

    Google Scholar 

  • Frick, G. A. (1938). Habitat of Euphorbia pseudocactus. Cactus and Succulent Journal (US), 9, 143–144.

    Google Scholar 

  • Genç, Z., & Rauh, W. (1984). Vergleichend-anatomische Untersuchungen an den Honigdrüsen einiger Euphorbia-Arten. In W. Rauh (Ed.), Anatomisch-Biochemische Untersuchungen an Euphorbien Teil 1. Trop. subtrop. Pflanzenw. 45: 9–54.

    Google Scholar 

  • Gilbert, M. G. (1987). Two new geophytic species of Euphorbia with comments on the subgeneric grouping of its African members. Kew Bulletin, 42, 231–244.

    Article  Google Scholar 

  • Gilbert, M. G. (1995). Euphorbiaceae. In S. Edwards, M. Tadesse, & I. Hedberg (Eds.), Flora of Ethiopia and Eritrea 2 (2): 265–380. Addis Ababa: National Herbarium.

    Google Scholar 

  • Glen, H. F., & Germishuizen, G. (2010). Botanical Exploration of southern Africa (Strelitzia 26) (2nd ed.). South African National Biodiversity Institute.

    Google Scholar 

  • Goebel, K. (1889). Pflanzenbiologische Schilderungen 1. : N.G. Elwert’sche Verlagsbuchhandlung.

    Google Scholar 

  • Grace, O. M., Buerki, S., Symonds, M. R. E., Forrest, F., Van Wyk, A. E., Smith, G. F., Klopper, R. R., Bjora, C. S., Neale, S., Demissew, S., Simmonds, M. S. J., & Rønstead, N. (2016). Evolutionary history and leaf succulence as explanations for medicinal use in aloes and the global popularity of Aloe vera. Evolutionary Biology, 2016, 16–29.

    Google Scholar 

  • Grove, A. T. (1969). Landform and climatic change in the Kalahari and Ngamiland. Geographical Journal, 135, 191–212.

    Article  Google Scholar 

  • Gschwendt, M., & Hecker, E. (1973). Über die Wirkstoffe der Euphorbiaceen I. Z. Krebsforsch., 80, 335–350.

    Article  CAS  Google Scholar 

  • Gschwendt, M., & Hecker, E. (1974). Über die Wirkstoffe der Euphorbiaceen II. Z. Krebsforsch., 81, 193–210.

    Article  CAS  Google Scholar 

  • Gunn, M., & Codd, L. E. (1981). Botanical exploration of Southern Africa. Cape Town: A.A. Balkema.

    Google Scholar 

  • Hall, H. (1949). Euphorbia fasciculata rediscovered. Cactus and Succulent Journal (US), 21, 140–141.

    Google Scholar 

  • Hammer, S. A. (1993). The genus Conophytum. Succulent Plant Publications.

    Google Scholar 

  • Hans, A. S. (1973). Chromosomal conspectus of the Euphorbiaceae. Taxon, 22, 591–636.

    Article  Google Scholar 

  • Hargreaves, B. J. (1976). Killing and curing. Cactus and Succulent Journal (US), 48, 190–196.

    Google Scholar 

  • Hargreaves, B. J. (1984). Is “Chikhawo” Euphorbia decidua? Euphorbia Journal, 2, 91–94.

    Google Scholar 

  • Hargreaves, B. J. (1992a). The succulent Euphorbias of Botswana. Euphorbia Journal, 8, 45–50.

    Google Scholar 

  • Hargreaves, B. J. (1992b). Volcanos & spurges. Euphorbia Journal, 8, 103–110.

    Google Scholar 

  • Hargreaves, B. J. (1992c). The other spurges of Lesotho (at least allegedly). Euphorbia Journal, 8, 126–132.

    Google Scholar 

  • Hargreaves, B. J. (1994). The Euphorbia schinzii complex. Euphorbia Journal, 9, 147–153.

    Google Scholar 

  • Hässler, A. (1931). Verwantschaftliche Gliederung der afrikanischen Euphorbien aus der Sektionen Trichadenia Pax und Rhizanthium Boiss. Botaniska Notiser, 1931, 317–338.

    Google Scholar 

  • Haworth, A. H. (1812). Synopsis Plantarum Succulentarum. R. Taylor.

    Google Scholar 

  • Haworth, A. H. (1827). LVI. Description of new succulent plants. Philosophical Magazine Annals of Chemistry, 1, 271–277.

    Article  Google Scholar 

  • Hecker, E. (1981). Cocarcinogenesis and tumor promoters of the diterpene ester type as possible carcinogenic risk factors. Journal of Cancer Research and Clinical Oncology, 99, 103–124.

    Article  CAS  PubMed  Google Scholar 

  • Hedberg, O. (1957). Afro-alpine vascular plants: a taxonomic revision. Symbolea Botanicae Upsalienses, 15, 1–411.

    Google Scholar 

  • Hemsley, W. B. (1904). Euphorbia viperina. Curtis’s Botanical Magazine, 130, t. 7971.

    Google Scholar 

  • Hendey, Q. B. (1982). Langebaanweg. A record of past life. South African Museum.

    Google Scholar 

  • Herre, H. (1936). Euphorbia Stapelioides Boiss. Euphorbia Review, 2, 18–19.

    Google Scholar 

  • Herre, H. (1950). Euphorbia superans Nel spec. nov. Desert Plants Life, 22, 15.

    Google Scholar 

  • Herre, H. (1973). Notes on South African succulents. Cactus and Succulent Journal (US), 45, 238–239.

    Google Scholar 

  • Holmes, S. (1993). Euphorbia. In M. Thulin (Ed.), Flora of Somalia (Vol. 1, pp. 306–337). Royal Botanic Gardens.

    Google Scholar 

  • Hooker, J. D. (1865). Euphorbia monteiri. Curtis’s Botanical Magazine, 91, t. 5534.

    Google Scholar 

  • Hooker, J. D. (1903). Euphorbia obesa. Curtis’s Botanical Magazine, 129, t. 7888.

    Google Scholar 

  • Hoppe, J. R. (1985). Die Morphogenese der Cyathiendrüsen und ihrer Anhänge, ihre blattypologische Deutung und Bedeutung. Botanische Jahrbücher fur Systematik, 105, 497–581.

    Google Scholar 

  • Hoppe, J. R., & Uhlarz, H. (1982). Morphogenese und typologische Interpretation des Cyathiums von Euphorbia-Arten. Beiträge zur Biologie der Pflanzen, 56, 63–98.

    Google Scholar 

  • Horn, J. W., van Ee, B. W., Morawetz, J. J., Riina, R., Steinmann, V. W., Berry, P. E., & Wurdack, K. J. (2012). Phylogenetics and the evolution of major structural characters in the giant genus Euphorbia L. (Euphorbiaceae). Molecular Phylogenetics and Evolution, 63, 305–326.

    Article  PubMed  Google Scholar 

  • Hutchinson, J. (1946). A botanist in Southern Africa. P.R. Gawthorn.

    Google Scholar 

  • Ihlenfeld, H.-D., & Gerbaulet, M. (1990). Untersuchungen zum Merkmalsbestand und zur Taxonomie der Gattungen Apatesia N.E. Br., Carpanthea N.E. Br., Conicosia N.E. Br., Herrea Schwantes und Hymenogyne Haw. (Mesembryanthemaceae Fenzl). Botanische Jahrbücher fur Systematik, 111, 457–498.

    Google Scholar 

  • Jackson, S. P. (1961). Climatological Atlas of Africa. CCTA/CSA.

    Google Scholar 

  • Jacobsen, N. H. G. (1988). Euphorbias of the Skeleton Coast National Park: Namibia and adjacent areas. Euphorbia Journal, 5, 59–67.

    Google Scholar 

  • Jacot Guillarmod, A. (1971). Flora of Lesotho (Basutoland). J. Cramer.

    Google Scholar 

  • Jones, K., & Smith, J. B. (1969). The chromosome identity of Monadenium Pax and Synadenium Pax (Euphorbiaceae). Kew Bulletin, 23, 491–498.

    Article  Google Scholar 

  • Jordaan, P. G. (1973). Le Vaillant’s botanical paintings. In J. C. Quinton, A. M. Lewin Robinson, & P. W. M. Sellicks (Eds.), François le Vaillant: Traveller in South Africa, and his collection of 165 Water-Colour Paintings, 1781–1784, 2: 43–81. : Library of Parliament.

    Google Scholar 

  • Kimberley, M. J. (1982). Dr R.F. Rand. Excelsa, 10, 85–87.

    Google Scholar 

  • Klak, C., & Linder, H. P. (1998). Systematics of Psilocaulon N.E. Br. & Caulipsolon Klak, gen. nov. (Mesembryanthemoideae, Aizoaceae). Botanische Jahrbücher fur Systematik, 120, 301–375.

    Google Scholar 

  • Klotzsch, J. F. (1860a). Linne’s natürliche Pflanzenklasse Tricoccae des Berliner Herbarium’s im Allgemeinen und die natürliche Ordnung Euphorbiaceae insbesondere. Monatsberichte der Königlichen Preussische Akademie des Wissenschaften zu Berlin, 1859, 236–254.

    Google Scholar 

  • Klotzsch, J. F. (1860b). Linne’s natürliche Pflanzenklasse Tricoccae des Berliner Herbarium’s im Allgemeinen und die natürliche Ordnung Euphorbiaceae insbesondere. Physikalische Abhandlungen der Königlichen Akademie der Wissenschaften zu Berlin, 1859, 1–108.

    Google Scholar 

  • Koutnik, D. L. (1984a). A Brief Taxonomy of the Euphorbia clava-loricata complex (Treisia). Euphorbia Journal, 2, 38–50.

    Google Scholar 

  • Koutnik, D. L. (1984b). Chamaesyce (Euphorbiaceae) – A newly recognized genus in southern Africa. South African Journal of Botany, 3, 262–264.

    Article  Google Scholar 

  • Koutnik, D. L. (1987). A taxonomic revision of the Hawaiian species of the genus Chamaesyce (Euphorbiaceae). Allertonia, 4, 331–388.

    Google Scholar 

  • Krauss, C. F. F. (1844). Pflanzen des Cap- und Natal-landes, gesammelt und zusammengestellt von Dr. Ferdinand Krauss (Fortsetzung). Flora, 27, 277–307.

    Google Scholar 

  • Krenzelok, E. P., Jacobsen, T. D., & Aronis, J. M. (1996). Poinsettia exposures have good outcomes…Just as we thought. American Journal of Emergency Medicine, 14, 671–674.

    Article  CAS  PubMed  Google Scholar 

  • Kunkel, G. (1978). Flora de Gran Canaria 3. Las plantas suculentas. Seix y Barral Hnos.

    Google Scholar 

  • Lamarck, J.-B. (1786). Encyclopédie Méthodique. Botanique 2. Paris/Liege: Panckoucke/Plomteux.

    Google Scholar 

  • Lawant, P., & Van Veldhuisen, R. (2014a). Euphorbia leachii Lawant & van Veldhuisen sp. nov., a new species from South Africa. Euphorbia World, 10, 5–8.

    Google Scholar 

  • Lawant, P., & Van Veldhuisen, R. (2014b). Some dubious interpretations of Euphorbia patula Mill. Euphorbia World, 10, 12–15.

    Google Scholar 

  • Leach, L. C. (1963). Euphorbia johnsonii. Its rediscovery and an amplified description. Kirkia, 3, 34–36.

    Google Scholar 

  • Leach, L. C. (1964). Euphorbia species from the Flora Zambesiaca area III. Journal of South African Botany, 30, 209–217.

    Google Scholar 

  • Leach, L. C. (1966). Euphorbia species from the Flora Zambesiaca area: V. Journal of South African Botany, 33, 173–182.

    Google Scholar 

  • Leach, L. C. (1967). Euphorbia species from the Flora Zambesiaca area: VII. Journal of South African Botany, 33, 247–262.

    Google Scholar 

  • Leach, L. C. (1968a). Euphorbia species from the Flora Zambesiaca area: VI. Kirkia, 6, 133–143.

    Google Scholar 

  • Leach, L. C. (1968b). Euphorbiae succulentae angolenses–I. Boletim da Sociedade Broteriana, Series 2, 42, 161–179.

    Google Scholar 

  • Leach, L. C. (1969). Euphorbiae succulentae angolenses: II. Boletim da Sociedade Broteriana, Series 2, 43, 163–182.

    Google Scholar 

  • Leach, L. C. (1970a). Euphorbia species from the Flora Zambesiaca Area: IX. Journal of South African Botany, 36, 13–52.

    Google Scholar 

  • Leach, L. C. (1970b). Euphorbiae Succulentae Angolensis: III. Bol. Soc. Brot., 44, 185–206.

    Google Scholar 

  • Leach, L. C. (1971). Euphorbia virosa Willd. with a new synonym and two new taxa from Angola. Boletim da Sociedade Broteriana, Series 2, 45, 349–362.

    Google Scholar 

  • Leach, L. C. (1973a). Euphorbia tirucalli L, its typification, synonymy and relationships, with notes on ‘Almeidina’ and ‘Cassoneira’. Kirkia, 9, 69–86.

    Google Scholar 

  • Leach, L. C. (1973b). Euphorbia species from the Flora Zambesiaca Area: 10. Journal of South African Botany, 39, 3–22.

    Google Scholar 

  • Leach, L. C. (1974). Euphorbia succulentae Angolenses: IV. Garcia de Orta, Sér. Bot., 2, 31–54.

    Google Scholar 

  • Leach, L. C. (1975a). Notes on Euphorbia mauritanica, E. gossypina and some related species with an amplified description of E. berotica. Bothalia, 11, 505–510.

    Article  Google Scholar 

  • Leach, L. C. (1975b). Euphorbia succulentae Angolenses: V. Garcia de Orta, Sér. Bot., 2, 111–116.

    Google Scholar 

  • Leach, L. C. (1976a). Euphorbia (Tetracanthae) in Angola and northern Kaokoland. Dinteria, 12, 1–35.

    Google Scholar 

  • Leach, L. C. (1976b). A review of the Euphorbia brevis-imitata-decidua complex with descriptions of five new species. Bulletin du Jardin Botanique National de Belgique, 46, 241–263.

    Article  Google Scholar 

  • Leach, L. C. (1976c). Distributional and morphological studies of the tribe Euphorbieae (Euphorbiaceae) and their relevance to its classification and possible evolution. Excelsa, 6, 3–18.

    Google Scholar 

  • Leach, L. C. (1977). Euphorbiae succulentae Angolenses: VI. Garcia da Orta, Sér. Bot., 3, 99–102.

    Google Scholar 

  • Leach, L. C. (1980). A taxonomic review of Euphorbia gariepina Boiss., its distribution, variability and synonymy. Excelsa Taxon Series, 2, 74–81.

    Google Scholar 

  • Leach, L. C. (1981). A new species of Euphorbia from the Karoo. Journal of South African Botany, 47, 103–107.

    Google Scholar 

  • Leach, L. C. (1982). A new species of Euphorbia from the Namib, South West Africa. Dinteria, 16, 27–31.

    Google Scholar 

  • Leach, L. C. (1983). A new Euphorbia from South West Africa. Journal of South African Botany, 49, 189–192.

    Google Scholar 

  • Leach, L. C. (1984a). A new Euphorbia from South Africa. Journal of South African Botany, 50, 341–345.

    Google Scholar 

  • Leach, L. C. (1984b). A new Euphorbia from the Richtersveld. Journal of South African Botany, 50, 563–568.

    Google Scholar 

  • Leach, L. C. (1985). A new species of Euphorbia (Euphorbiaceae) from the Cape Province. South African Journal of Botany, 51, 281–283.

    Article  Google Scholar 

  • Leach, L. C. (1986a). Euphorbia filiflora. Flowering Plants of Africa, 49, t. 1927.

    Google Scholar 

  • Leach, L. C. (1986b). A new Euphorbia (Euphorbiaceae) from the western Cape Province. South African Journal of Botany, 52, 10–12.

    Article  Google Scholar 

  • Leach, L. C. (1986c). A new Euphorbia from the western Knersvlakte, Cape Province. South African Journal of Botany, 52, 369–371.

    Article  Google Scholar 

  • Leach, L. C. (1988a). A new species of Euphorbia (Euphorbiaceae) from the south-western Cape. South African Journal of Botany, 54, 501–503.

    Article  Google Scholar 

  • Leach, L. C. (1988b). The Euphorbia juttae-gentilis complex, with a new species and a new subspecies. South African Journal of Botany, 54, 534–538.

    Article  Google Scholar 

  • Leach, L. C. (1991). Euphorbia griseola Pax: Its subspecies & relationships. The Euphorbia Journal, 7, 131–137.

    Google Scholar 

  • Leach, L. C. (1992). Euphorbia knuthii Pax: Its distribution and other matters. Euphorbia Journal, 8, 72–73.

    Google Scholar 

  • Leach, L. C., & Williamson, G. (1990). The identities of two confused species of Euphorbia (Euphorbiaceae) with descriptions of two closely related new species from Namaqualand. South African Journal of Botany, 56, 71–78.

    Article  Google Scholar 

  • Lemaire, C. (1855). Observations diagnostico-nomenclaturales sur les Euphorbes charnues du Cap. Ill. Hort. 2: misc. 65–69.

    Google Scholar 

  • Lemaire, C. (1858). Nouvelles Euphorbes. Ill. Hort. 5: misc. 63–64.

    Google Scholar 

  • Le Vaillant, F. (1790). Voyage de M. le Vaillant dans l’intérier de l/Afrique par Le Cap de Bonne Espérence, dans les années 1783, 84 & 85. Leroy.

    Google Scholar 

  • Le Vaillant, F. (1795). Second voyage dans l’intérieur de l’Afrique: par le Cap de Bonne-Espérance, dans les années 1783, 84, et 85. H.J. Jansen.

    Google Scholar 

  • Le Vaillant, F. (1797). Reise in das Innere con Afrika, vom Vorgebürge der guten Hoffnung aus, in den Jahren 1780 bis 85 (Vols. 3, 4). P.H. Guilhauman.

    Google Scholar 

  • Liede, S., & Meve, U. (1993). Towards an understanding of the Sarcostemma viminale (Asclepiadaceae) complex. Botanical Journal of the Linnean Society, 112, 1–15.

    Article  Google Scholar 

  • Linneaus, C. (1753). Species Plantarum. Laurentius Salvius.

    Google Scholar 

  • Löffler, E., & Uhlarz, H. (1969). Observations on some Euphorbias from Tropical West-Africa. Cactus and Succulent Journal (US), 41, 210–220.

    Google Scholar 

  • Loffler, L., & Loffler, P. (2005). Swaziland tree Atlas, including selected shrubs and climbers. SABONET.

    Google Scholar 

  • Lotsy, J. P., & Goddijn, W. A. (1928). Hybrisization in the native flora. Genetica, 10, 1–129.

    Article  Google Scholar 

  • Loutit, B. D., Louw, G. N., & Seely, M. K. (1987). First approximation of food preferences and the chemical composition of the diet of the desert-dwelling black rhinoceros, Diceros bicornis L. Madoqua, 15, 35–54.

    Google Scholar 

  • Luckow, M. (1995). Species concepts: assumptions, methods and applications. Systematic Botany, 20, 589–605.

    Article  Google Scholar 

  • Lugard, E. (1938). Monadenium lugardae N.E.Br. Cactus and Succulent Journal (US), 9, 178–179.

    Google Scholar 

  • Macnae, M. M., & Davidson, L. E. (1969). The volume “Icones Plantarum et Animalium” in the Africana Museum, Johannesburg, and its relationship to the Codex Witsenii quoted by Jan Burman in his “Decades Rariorum Africanarum Plantarum”. Journal of South African Botany, 35, 65–81.

    Google Scholar 

  • Manning, J., & Goldblatt, P. (Eds.). (2012). Plants of the Greater Cape Floristic Region. 1: The Core Cape Flora. (Ed.). Strelitzia 29. : South African National Biodiversity Institute.

    Google Scholar 

  • Marloth, R. (1908). Das Kapland, insonderheit das Reich der Kapflora, das Waldgebiet und die Karroo, pflanzengeographisch dargestellt. G. Fischer.

    Google Scholar 

  • Marloth, R. (1910). Some new South African succulents. Part III. Transactions of the Royal Society of South Africa, 2, 33–39.

    Article  Google Scholar 

  • Marloth, R. (1925). The flora of South Africa. 2 (2). Cambridge: Cambridge University Press.

    Google Scholar 

  • Marloth, R. (1928). The meloformia group of Euphorbias. South African Gardening & Country Life, 28, 45–46.

    Google Scholar 

  • Marloth, R. (1930). A revision of the group Virosae of the genus Euphorbia as far as represented in South Africa. South African Journal of Science, 27, 331–340.

    Google Scholar 

  • Marloth, R. (1931). Euphorbias III. South African Gardening & Country Life, 21(127), 133.

    Google Scholar 

  • Marx, J. G. (1987). Euphorbia albipollinifera Leach. A recently described species from the South-Eastern Cape Province. Aloe, 24, 18–19.

    Google Scholar 

  • Marx, J. G. (1988). Euphorbia meloformis Aiton and Euphorbia valida N.E. Brown: Some observations in habitat. Euphorbia Journal, 5, 94–103.

    Google Scholar 

  • Marx, J. G. (1992). The succulent Euphorbias of the southeastern Cape Province, Part 1: Dwarf species and smaller shrubs. Euphorbia Journal, 8, 74–102.

    Google Scholar 

  • Marx, J. G. (1993). The subglobose euphorbias and relatives. Aloe, 30, 103–107.

    Google Scholar 

  • Marx, J. G. (1994). Euphorbia brevirama N.E. Brown: The quest continues. Euphorbia Journal, 9, 221–222.

    Google Scholar 

  • Marx, J. G. (1999a). The South African Melon-shaped Euphorbias: The full picture as known to date. Euphorbiaceae Study Group Bulletin, 12, 13–34.

    Google Scholar 

  • Marx, J. G. (1999b). Euphorbia suppressa J.G. Marx and Euphorbia gamkensis J.G. Marx, two hitherto-unnamed species from the Western Cape Province, South Africa. Cactus and Succulent Journal (US), 71, 33–40.

    Google Scholar 

  • Marx, J. G. (2015). Accumulation of errors regarding Euphorbia crassipes Marloth. Euphorbia World, 11, 20–26.

    Google Scholar 

  • Mauseth, J. D. (2004a). The structure of photosynthetic succulent stems in plants other than cacti. International Journal of Plant Sciences, 165, 1–9.

    Article  Google Scholar 

  • Mauseth, J. D. (2004b). Cacti and other succulents: Stem anatomy of ‘other succulents’ has little in common with that of cact. Bradleya, 22, 131–140.

    Article  Google Scholar 

  • Mayr, E. (1964). Systematics and the Origin of Species from the Viewpoint of a Zoologist. Dover Publications Inc..

    Google Scholar 

  • Meerburg, N. (1789). Plantae rariores vivis coloribus depictae. J. Meerburg.

    Book  Google Scholar 

  • Meyer, P. G. (1967). Euphorbiaceae. In H. Merxmüller (Ed.), Prodromus einer Flora von Südwestafrika (p. 67). Lehre: J. Cramer.

    Google Scholar 

  • Miller, P. (1768). The gardeners dictionary (8th ed., p. London).

    Google Scholar 

  • Millspaugh, C. H. (1909). Praenunciae bahamensis II. Publ. Field Columb. Mus. Bot Ser, 2, 298–322.

    Google Scholar 

  • Monteiro, J. J. (1875). Angola and the River Congo. 2 Vols. : MacMillan & Co.

    Google Scholar 

  • Nel, G. C. (1935). Jahrb. Deutsch. Kakteen-Ges, 1, 29–32, 42–43.

    Google Scholar 

  • Nordenstam, B. (1969). Phytogeography of the genus Euryops (Compositae). Opera Botanica, 23, 5–77.

    Google Scholar 

  • Nordenstam, B. (1974). The flora of the Brandberg. Dinteria, 11, 1–67.

    Google Scholar 

  • Oudemans, C. A. J. A. (Ed.). (1865). Neerland’s Plantentuin (Vol. 1). J.B. Wolters.

    Google Scholar 

  • Palmer, E. (1966). The plains of Camdeboo. Collins.

    Google Scholar 

  • Panhuysen, L. (2011). Een Nederlander in de Wildernis. De Ontdekkingsreizen van Robert Jakob Gordon (1743–1795) in Zuid-Afrika. Rijksmuseum, Amsterdam (Derde, Herziene Druk).

    Google Scholar 

  • Park, K.-R. (1996). Phylogeny of New World subtribe Euphorbiinae (Euphorbiaceae). Korean Journal of Plant Taxonomy, 26, 235–256.

    Article  Google Scholar 

  • Park, K.-R., & Elisens, W. J. (2000). A phylogenetic study of tribe Euphorbieae (Euphorbiaceae). International Journal of Plant Sciences, 161, 425–434.

    Article  CAS  PubMed  Google Scholar 

  • Park, K.-R., & Jansen, R. K. (2007). A phylogeny of Euphorbieae subtribe Euphorbiinae (Euphorbiaceae) based on molecular data. Journal of Plant Biology, 50, 644–649.

    Article  CAS  Google Scholar 

  • Partridge, T. C., & Maud, R. R. (2000). Macro-scale geomorphic evolution of southern Africa. In T. C. Partridge & R. R. Maud (Eds.), The Cenozoic of southern Africa (pp. 3–18). Oxford University Press.

    Google Scholar 

  • Paterson, W. (1789). A narrative of four journeys into the country of the Hottentots, and Caffraria: In the years one thousand seven hundred and seventy-seven, eight, and nine. J. Johnson.

    Google Scholar 

  • Paterson, W. (1790). Reisen in das Land der Hottentotten und der Kaffern während der Jahre 1777, 1778 und 1779. C.F. Voss, Berlin [translated from the English by J.R. Forster].

    Google Scholar 

  • Pax, F. (1894). Euphorbiaceae africanae. II. Bot. Jahrb. Syst., 19, 76–127.

    Google Scholar 

  • Pax, F. (1900). Euphorbiaceae. In Zahlbruckner, A. (Ed.), Plantae Pentherianae. Ann. K. Naturhist. Hofmus. 15: 48–51.

    Google Scholar 

  • Pax, F. (1904). Monographische Übersicht über die afrikanische Arten aus der Sektion Diacanthium der Gattung Euphorbia. Botanische Jahrbücher fur Systematik, 34, 61–85.

    Google Scholar 

  • Pax, F. (1905). XXIII. Euphorbia schoenlandii Pax… Feddes Repert. Spec. Nov Regni Veg. 1: 59.

    Google Scholar 

  • Pax, F. (1908). Euphorbiaceae. In Schinz, H. (Ed.), Beiträge zur Kenntnis der Afrikanischen-Flora 21. Bull. Herb. Boiss., Sér. 2, 8: 634–637.

    Google Scholar 

  • Pax, F. (1909). Euphorbiaceae africanae. IX. Botanische Jahrbücher fur Systematik, 43, 75–90.

    Google Scholar 

  • Pax, F. (1921). Euphorbiaceae. In A. Engler & O. Drude (Eds.): 1–168. Die Vegetation der Erde, 9, Die Pflanzenwelt Afrikas, insbesondere seiner tropischen Gebiete 3 (2). Leipzig: Engelmann.

    Google Scholar 

  • Pearson, H. H. W. (1914). Observations on the internal temperatures of Euphorbia virosa and Aloe dichotoma. Annals of the Bolus Herbarium, 1, 41–66.

    Google Scholar 

  • Peirson, J. A., Bruyns, P. V., Riina, R., Morawetz, J. J., & Berry, P. E. (2013). A molecular phylogeny and classification of the largely succulent and mainly African Euphorbia subg. Athymalus (Euphorbiaceae). Taxon, 62, 1178–1199.

    Article  Google Scholar 

  • Petiver, J. (1709–11). Catalogus Classicus & Topicus Omnium Rerum Figuratum in V. Decadibus, Seu primo [-secundo] volumine Gazophylacii Naturae & Artis. London: C. Bateman.

    Google Scholar 

  • Phillips, E. P. (1924). Euphorbia cooperi. Flowering Plants of South Africa, 4, t. 157.

    Google Scholar 

  • Phillips, E. P. (1925). Euphorbia tridentata. Flowering Plants of South Africa, 5, t. 197.

    Google Scholar 

  • Phillips, E. P. (1926). Euphorbia monteiri. Flowering Plants of South Africa, 6, t. 218.

    Google Scholar 

  • Phillips, E. P. (1928a). Euphorbia bubalina. Flowering Plants of South Africa, 8, t. 285.

    Google Scholar 

  • Phillips, E. P. (1928b). Euphorbia trichadenia. Flowering Plants of South Africa, 8, t. 288.

    Google Scholar 

  • Phillips, E. P. (1929a). Euphorbia stellaespina. Flowering Plants of South Africa, 9, t. 344.

    Google Scholar 

  • Phillips, E. P. (1929b). Euphorbia kunthii. Flowering Plants of South Africa, 9, t. 348.

    Google Scholar 

  • Phillips, E. P. (1929c). Euphorbia fusca. Flowering Plants of South Africa, 9, t. 350.

    Google Scholar 

  • Phillips, E. P. (1930). Euphorbia heptagona. Flowering Plants of South Africa, 10, t. 390.

    Google Scholar 

  • Phillips, E. P. (1934). Euphorbia valida. Flowering Plants of South Africa, 14, t. 526, t. 527.

    Google Scholar 

  • Pickford, M., & Senut, B. (2000). Geology and palaeobiology of the central and southern Namib Desert, southwestern Africa, Vol. 1. Geology and history of study. Memoir Geological Survey of Namibia, 18, 1–155.

    Google Scholar 

  • Plukenet, L. (1692). Phytographia seu plantae quamplurimae novae… 3. .

    Google Scholar 

  • Poiret, J. L. M. (1812). Encyclopédie Méthodique, Botanique (Lamarck), Supplément, 2 (2). : H. Agasse.

    Google Scholar 

  • Porembski, S., & Barthlott, W. (2000). Granitic and gneissic outcrops (inselbergs) as centers of diversity for dessication-tolerant vascular plants. Plant Ecology, 151, 19–28.

    Article  Google Scholar 

  • Prain, D. (1909). Euphorbia ledienii. Curtis’s Botanical Magazine, 135, t. 8275.

    Google Scholar 

  • Prenner, G., & Rudall, P. J. (2007). Comparative Ontogeny of the cyathium in Euphorbia (Euphorbiaceae) and its allies: Exploring the organ-flower-inflorescence boundary. American Journal of Botany, 94, 1612–1629.

    Article  PubMed  PubMed Central  Google Scholar 

  • Prenner, G., Hopper, S. D., & Rudall, P. J. (2008). Pseudanthium development in Calycopeplus paucifolius, with particular reference to the evolution of the cyathium in Euphorbieae (Euphorbiaceae-Malpighiales). Australian Systematic Botany, 21, 153–161.

    Article  PubMed  PubMed Central  Google Scholar 

  • Rabinowitz, P. D., Coffin, M. F., & Falvey, D. (1983). The separation of Madagascar and Africa. Science, 220, 67–69.

    Article  CAS  PubMed  Google Scholar 

  • Raper, P. E., & Boucher, M. (Eds.). (1988). Robert Jacob Gordon: Cape Travels 1777–1786. Brenthurst Press.

    Google Scholar 

  • Rauh, W. (1967). Die grossartige Welt der Sukkulenten. Paul Parey.

    Google Scholar 

  • Rauh, W., Loffler, E., & Uhlarz, H. (1969). Observations on some Euphorbias from tropical West Africa. Cactus and Succulent Journal (US), 41, 210–220.

    Google Scholar 

  • Raven, P. H. (1975). The bases of angiosperm phylogeny: Cytology. Annals of the Missouri Botanical Garden, 62, 724–764.

    Article  Google Scholar 

  • Raven, P. H., & Axelrod, D. J. (1974). Angiosperm biogeography and past continental movements. Annals of the Missouri Botanical Garden, 61, 539–657.

    Article  Google Scholar 

  • Renner, S. (2004). Plant dispersal across the tropical Atlantic by wind and sea currents. International Journal of Plant Sciences, 165(Suppl), 523–533.

    Google Scholar 

  • Reynolds, G. W. (1966). The Aloes of tropical Africa and Madagascar. The Aloes Book Fund.

    Google Scholar 

  • Riina, R., Peirson, J. A., Geltman, D. V., Molero, J., Frajman, B., Pahlevani, A., Barres, L., Morawetz, J. J., Salmaki, Y., Zarre, S., Kryukov, A., Bruyns, P. V., & Berry, P. E. (2013). A worldwide molecular phylogeny and classification of the leafy spurges, Euphorbia subgenus Esula (Euphorbiaceae). Taxon, 62, 316–342.

    Article  Google Scholar 

  • Röper, J. A. C. (1824). Enumeratio Euphorbiarum quae in Germannia et Pannonia gignuntur. C.E. Rosenbusch.

    Google Scholar 

  • Rolfe, R. A. (1889). Matabele Land and the Victoria Falls: A naturalist’s wanderings in the interior of South Africa. From the letters & journals of the late Frank Oates, F.R.G.S. (C. G. Oates, Ed.). 2nd ed. Appendix V. Botany: 390–413. London: Kegan Paul, Trench & Co.

    Google Scholar 

  • Rutherford, M. C., & Westfall, R. H. (1986). Biomes of southern Africa – An objective categorisation. Memoirs of the Botanical Survey of South Africa, 54, 1–98.

    Google Scholar 

  • Schnabel, D. H. (2013). A morphology based taxonomic revision of the Euphorbia polygona species complex. Euphorbia World, 9, 5–25.

    Google Scholar 

  • Schnabel, D. H. (2018). A monograph of Euphorbia polygona from the perspective of an amateur botanist. D.H. Schnabel.

    Google Scholar 

  • Schnepf, E., & Deichgräber, G. (1984). Electron microscopical studies of nectaries of some Euphorbia species. In W. Rauh (Ed.), Anatomisch-Biochemische Untersuchungen an Euphorbien Teil 1. Trop. subtrop. Pflanzenw. 45: 55–93.

    Google Scholar 

  • Schweickerdt, H. G. (1935). In H. G. Schweickerdt & I. C. Verdoorn (Eds.), Notes on the Flora of southern Africa VI. Bulletin of Miscellaneous Information 1935: 204–209.

    Google Scholar 

  • Seely, M. K., & Jacobson, K. M. (1994). Desertification and Namibia: A perspective. Journal of African Zoology, 108, 21–36.

    Google Scholar 

  • Shah, J. J., & Jani, P. M. (1964). Shoot apex of Euphorbia neriifolia L. In Proceedings of the National Academy of Sciences, India 30, B, no. 2: 81–91.

    Google Scholar 

  • Sidwell, K. (1999). Monographing Brillantaisia P. Beauv. (Acanthaceae): How to delimit species and higher taxa. In J. Timberlake & S. Kativu (Eds.), African Plants: Biodiversity, Taxonomy and Uses. Proceedings of the 1997 AETFAT Congress (pp. 367–382). Royal Botanic Gardens.

    Google Scholar 

  • Siesser, W. G. (1980). Late Miocene origin of the Benguela upswelling system off northern Namibia. Science, 208, 283–285.

    Article  CAS  PubMed  Google Scholar 

  • Sloane, B. L. (1952). Alain White 1880–1951. Cactus and Succulent Journal (US), 24, 70–71.

    Google Scholar 

  • Snijman, D. A. (Ed.). (2013). Plants of the Greater Cape Floristic Region. 2: The Extra Cape Flora. Strelitzia 30: 371–379. : South African National Biodiversity Institute.

    Google Scholar 

  • Srivastava, S., Narain, P., & Srivastava, G. S. (1987). A new basic chromosome number in Euphorbia based on male meiosis of E. clandestina Jacquin Hort. Cytologica, 52, 627–630.

    Article  Google Scholar 

  • Stebbins, G. L. (1950). Variation and Evolution in Plants. Columbia University Press.

    Book  Google Scholar 

  • Steinmann, V. W. (2003). The submersion of Pedilanthus into Euphorbia (Euphorbiaceae). Acta Botánica Mexicana, 65, 45–50.

    Article  Google Scholar 

  • Steinmann, V. W., & Porter, J. M. (2002). Phylogenetic relationships in Euphorbieae (Euphorbiaceae) based on ITS and ndhF sequence data. Annals of the Missouri Botanical Garden, 89, 453–490.

    Article  Google Scholar 

  • Steinmann, V. W., Van Ee, B., Berry, P. E., & Gutiérrez, J. (2007). The systematic position of Cubanthus and other shrubby endemic species of Euphorbia (Euphorbiaceae) in Cuba. Anales del Jardín Botánico de Madrid, 64, 123–133.

    Article  Google Scholar 

  • Struck, M. (1992). Pollination ecology in the arid winter rainfall region of southern Africa: A case study. Mitteilungen aus dem Institut für allgemeine Botanik in Hamburg, 24, 61–90.

    Google Scholar 

  • Swanepoel, W. (2009a). Euphorbia ohiva (Euphorbiaceae), a new species from Namibia and Angola. South African Journal of Botany, 75, 249–255.

    Article  Google Scholar 

  • Swanepoel, W. (2009b). Euphorbia otjingandu (Euphorbiaceae), a new species from the Kaokoveld, Namibia. South African Journal of Botany, 75, 497–504.

    Article  Google Scholar 

  • Swanepoel, W. (2013). Euphorbia otjipembana subsp. okakoraensis (Euphorbiaceae), a new subspecies from the Kaokoveld, Namibia, with an amplified description of Euphorbia otjipembana. Phytotaxa, 117, 51–57.

    Article  Google Scholar 

  • Thellung, A. (1916). Euphorbiaceae. In H. Schinz (Ed.), Beiträge zur Kenntnis der afrikanischen Flora. (XXVII). Vierteljahrsschr. Naturforsch. Gesellsch. Zürich 61: 431– 461.

    Google Scholar 

  • Thunberg, C. P. (1800). Prodomus Plantarum Capensium 2. : J.F. Edman.

    Google Scholar 

  • Thunberg, C. P. (1823). In J. A. Schultes (Ed.), Flora Capensis, sistens plantas promontorii bonae spei Africes 2. Stuttgart: J.G. Cotta.

    Google Scholar 

  • Tian, X., Wang, Q., & Zhou, Y. (2018). Euphorbia section Hainanensis (Euphorbiaceae), a new section endemic to the Hainan Island of China from biogeographical, karyological, and phenotypical evidence. Frontiers in Plant Science, 9, 1–9.

    Article  Google Scholar 

  • Troll, W. (1935–7). Vergleichende Morphologie der höheren Pflanzen 1 (1). Berlin: Gebrüder Borntraeger.

    Google Scholar 

  • Uhlarz, H. (1974). Entwicklungsgeschichtliche Untersuchungen zur Morphologie der basalen Blatteffigurationen sukkulenter Euphorbien aus den Subsektionen Diacanthium Boiss. und Goniostema Baill. Tropische und Subtropische Pflanzenwelt, 9, 1–69.

    Google Scholar 

  • Uhlarz, H. (1975a). Über die strittige Homologie sogenannter Stipulardrüsen bei einiger Euphorbia-Arten der Sektion Euphorbium. Plant Systematics and Evolution, 124, 229–250.

    Article  Google Scholar 

  • Uhlarz, H. (1975b). Über die Dorsalstacheln der Podarien von Monadenium guentheri Pax. Beiträge zur Biologie der Pflanzen, 51, 335–352.

    Google Scholar 

  • Uhlarz, H. (1978). Über die Stipularorgane der Euphorbiaceae, unter besonderer Berücksichtigung ihrer Rudimentation. Tropische und Subtropische Pflanzenwelt, 23, 1–65.

    Google Scholar 

  • Venter, K. (1987). A new field hybrid from the Transvaal?: A suspected hybrid between Euphorbia excelsa & Euphorbia enormis. Euphorbia Journal, 4, 69–70.

    Google Scholar 

  • Vesling, J. (1638). Ioannis Veslingii Mindani…De Plantis Aegyptis observationes et notae as Proseperum Alpinum. Leiden: P. Frambottum.

    Google Scholar 

  • Veslingius, J. (1638). Ioannis Veslingii Mindani…De Plantis Aegyptis observationes et notae as Proseperum Alpinum. P. Frambottum.

    Google Scholar 

  • Visser, J. (1981). South African parasitic flowering plants. Juta.

    Google Scholar 

  • Vlok, J., & Schutte-Vlok, A. L. (2010). Plants of the Klein Karoo. Hatfield: Umdaus Press.

    Google Scholar 

  • Vogel, S. (1954). Blütenbiologische Typen als Elemente der Sippengliederung. Dargestellt anhand der Flora Südafrikas. Botanische Studien, 1, 1–338.

    Google Scholar 

  • Von Willert, D. J., Eller, B. M., Werger, M. J. A., Brinckmann, E., & Ihlenfeldt, H.-D. (1992). Life strategies of succulents in deserts with special reference to the Namib Desert. Cambridge University Press.

    Google Scholar 

  • Vosa, C. G., & Bassi, P. (1991). Cromosome (sic) studies in the Southern African flora. 95–102. The basic caryotype of eight species of succulent Euphorbia L. Caryologia, 44, 27–33.

    Article  Google Scholar 

  • Ward, J. D., & Corbett, I. (1990). Towards an age for the Namib. In M. K. Seely (Ed.), Namib ecology: 25 years of Namib research (Transvaal Museum Monograph) (Vol. 7, pp. 17–26). Transvaal Museum.

    Google Scholar 

  • Ward, J. D., Seely, M. K., & Lancaster, N. (1983). On the antiquity of the Namib. South African Journal of Science, 79, 175–183.

    Google Scholar 

  • Watt, J. M., & Breyer-Brandwijk, M. G. (1962). The medicinal and poisonous plants of Southern and Eastern Africa (2nd ed.). E. & S. Livingstone.

    Google Scholar 

  • Webster, G. L. (1967). The genera of Euphorbiaceae in the southeastern United States. Journal of the Arnold Arboretum, 48, 363–430.

    Article  Google Scholar 

  • Webster, G. L. (1975). Conspectus of a new classification of the Euphorbiaceae. Taxon, 24, 593–601.

    Article  Google Scholar 

  • Webster, G. L. (1994). Synopsis of the genera and suprageneric taxa of Euphorbiaceae. Annals of the Missouri Botanical Garden, 81, 33–144.

    Article  Google Scholar 

  • Weiss, J. E. (1893). Empfehlenswerte Kakteen. Neubert’s Deutsch. Garten-Magazin, 46, 286–292.

    Google Scholar 

  • Wheeler, L. C. (1939). A miscellany of New World Euphorbiaceae 2. Contributions from the Gray Herbarium, 127, 48–78.

    Article  Google Scholar 

  • Wheeler, L. C. (1941). Euphorbia subg. Chamaesyce in Canada and the Unites States exclusive of southern Florida. Rhodora, 43, 97–154.

    Google Scholar 

  • Wheeler, L. C. (1943). The genera of the living Euphorbieae. American Midland Naturalist, 30, 456–503.

    Article  Google Scholar 

  • White, A. C., Dyer, R. A., & Sloane, B. L. (1941). The Succulent Euphorbieae (Southern Africa). Pasadena: Abbey Garden Press.

    Google Scholar 

  • Wijnands, D. O. (1983). The botany of the Commelins. A.A.Balkema.

    Google Scholar 

  • Wijnands, D. O., Wilson, M. L., & Toussant van Hove, T. (Eds.). (1996). Jan Commelin’s monograph on Cape Flora. The Editors.

    Google Scholar 

  • Wiley, E. O. (1981). Phylogenetics. The theory and practice of phylogenetic systematics. Wiley.

    Google Scholar 

  • Williamson, G. (1995). Euphorbia versicolores, a new species from the northwestern Cape, South Africa. Cactus and Succulent Journal (US), 67, 284–287.

    Google Scholar 

  • Williamson, G. (1999). A new species of Monadenium (Euphorbiaceae) from south-east Zimbabwe. Excelsa, 19, 56–58.

    Google Scholar 

  • Williamson, G. (2003). A new variety of Euphorbia filiflora Marloth from the Northern Cape Province of South Africa. Bradleya, 21, 49–52.

    Article  Google Scholar 

  • Williamson, G. (2007). Notes on Euphorbia multiramosa Nel (Euphorbiaceae) and related species. Euphorbia World, 3, 8–18.

    Google Scholar 

  • Wilman, M. (1946). Preliminary check list of the flowering plants and ferns of Griqualand West (Southern Africa). Deighton, Bell & Co.

    Google Scholar 

  • Wilson, M. L., Toussant van Hove-Exalto, T., & van Rijssen, W. J. J. (Eds.). (2002). Codex Witsenii. Iziko Museums of Cape Town/Davidii Media.

    Google Scholar 

  • Wiss, H.-J. (1957). Brandberg Expedition 1955. Ein Bericht ueber die gesammelten und beobachteten Pflanzen. Journal of the South West Africa Scientific Society, 12, 45–68.

    Google Scholar 

  • Worsdell, W. C. (1914). On some points in the stem-anatomy of Euphorbia virosa and Aloe dichotoma. Annals of the Bolus Herbarium, 1, 67–71.

    Google Scholar 

  • Yang, Y., Riina, R., Morawetz, J. J., Haevermans, T., Aubriot, X., & Berry, P. E. (2012). Molecular phylogenetics and classification of Euphorbia subgenus Chamaesyce (Euphorbiaceae). Taxon, 61, 764–789.

    Article  Google Scholar 

  • Zimmermann, N. F. A., Ritz, C. M., & Hellwig, F. H. (2010). Further support for the phylogenetic relationships within Euphorbia L. (Euphorbiaceae) from nrITS and trnL-trnF IGS sequence data. Plant Systematics and Evolution, 286, 39–58.

    Article  Google Scholar 

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    Peter V. Bruyns

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Bruyns, P.V. (2022). Euphorbia subg. Chamaesyce. In: Euphorbia in Southern Africa. Springer, Cham. https://doi.org/10.1007/978-3-030-49399-8_1

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