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WIKTROP - Weed Identification and Knowledge in the Tropical and Mediterranean areas
WIKTROP - Weed Identification and Knowledge in the Tropical and Mediterranean areas
SpeciesMapsDocumentsIDAO

Eclipta prostrata (L.) L.

Accepted
Eclipta prostrata (L.) L.
Eclipta prostrata (L.) L.
Eclipta prostrata (L.) L.
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Eclipta prostrata (L.) L.
Eclipta prostrata (L.) L.
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Eclipta prostrata (L.) L.
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🗒 Synonyms
synonymAcmella lanceolata Link ex Spreng.
synonymAmellus carolinianus Walter
synonymAnthemis abyssinica J.Gay ex A.Rich.
synonymAnthemis abyssinica var. abyssinica
synonymAnthemis bornmuelleri var. brachyota (Eig) Feinbrun & Fertig
synonymAnthemis bornmuelleri var. galilaea (Eig) Feinbrun & Fertig
synonymAnthemis cotula Blanco
synonymAnthemis cotula Blanco [Illegitimate]
synonymAnthemis cotula subsp. lithuanica (DC.) Tzvelev
synonymAnthemis cotula var. hierosolymitana Eig
synonymAnthemis cotula-foetida Crantz
synonymAnthemis cotuloides Raf. ex DC.
synonymAnthemis galilaea var. brachyota Eig
synonymAnthemis galilaea var. hierosolymitana (Eig) Yavin
synonymAnthemis sulphurea Wall. ex Nyman
synonymAnthemis viridis Blanco
synonymArtemisia viridis Blanco
synonymArtemisia viridis Blanco [Illegitimate]
synonymBellis racemosa Steud.
synonymBellis ramosa Jacq.
synonymBuphthalmum diffusum Vahl ex DC.
synonymChamaemelum foetidum Baumg.
synonymChamaemelum foetidum Garsault
synonymCotula alba (L.) L.
synonymCotula alva (L.) L.
synonymCotula oederi Murray
synonymCotula prostrata (L.) L.
synonymEclipta adpressa Moench
synonymEclipta adpressa Moench [Illegitimate]
synonymEclipta alba (L.) Hassk.
synonymEclipta alba f. erecta (L.) Hassk.
synonymEclipta alba f. longifolia Hassk.
synonymEclipta alba f. prostrata (L.) Hassk.
synonymEclipta alba f. prostrata Huber
synonymEclipta alba f. zippeliana (Blume) Hassk.
synonymEclipta alba var. alba
synonymEclipta alba var. erecta (L.) Hassl.
synonymEclipta alba var. erecta (L.) Miq.
synonymEclipta alba var. longifolia Bettfr.
synonymEclipta alba var. parviflora (Wall. ex DC.) Miq.
synonymEclipta alba var. prostrata (L.) Hassl.
synonymEclipta alba var. prostrata (L.) Miq.
synonymEclipta alba var. zippeliana (Blume) Miq.
synonymEclipta angustifolia C.Presl
synonymEclipta arabica Steud.
synonymEclipta arabica Steud. [Illegitimate]
synonymEclipta brachypoda Michx.
synonymEclipta ciliata Raf.
synonymEclipta dentata B.Heyne ex Wall.
synonymEclipta dentata Wall. [Invalid]
synonymEclipta dichotoma Raf.
synonymEclipta dubia Raf.
synonymEclipta erecta L.
synonymEclipta erecta L. [Illegitimate]
synonymEclipta erecta var. brachypoda (Michx.) Torr. & A.Gray
synonymEclipta erecta var. diffusa DC.
synonymEclipta erecta var. erecta
synonymEclipta erecta var. latifolia Willd. ex Walp.
synonymEclipta erecta var. prostrata (L.) Baker
synonymEclipta erecta var. zippeliana (Blume) J.Kost.
synonymEclipta flexuosa Raf.
synonymEclipta heterophylla Bartl.
synonymEclipta hirsuta Bartl.
synonymEclipta linearis Otto ex Sweet
synonymEclipta longifolia Schrad.
synonymEclipta longifolia Schrad. ex DC.
synonymEclipta marginata Boiss.
synonymEclipta marginata Steud.
synonymEclipta marginata Steud. [Illegitimate]
synonymEclipta nutans Raf.
synonymEclipta nutans var. nutans
synonymEclipta oederi (Murr.) Weigel
synonymEclipta oederi (Murray) Weigel
synonymEclipta palustris DC.
synonymEclipta parviflora Wall. ex DC.
synonymEclipta patula Schrad.
synonymEclipta patula Schrad. ex DC.
synonymEclipta philippinensis Gand.
synonymEclipta procumbens Michx.
synonymEclipta procumbens var. brachypoda (Michx.) A.Gray
synonymEclipta procumbens var. patula (Schrad.) DC.
synonymEclipta prostrata f. aureoreticulata Y.T.Chang
synonymEclipta prostrata f. prostrata
synonymEclipta prostrata var. prostrata
synonymEclipta prostrata var. undulata (Willd.) DC.
synonymEclipta prostrata var. zippeliana (Blume) J.Kost.
synonymEclipta pumila Raf.
synonymEclipta punctata Jacq.
synonymEclipta punctata L.
synonymEclipta punctata L. [Illegitimate]
synonymEclipta simplex Raf.
synonymEclipta spicata Spreng.
synonymEclipta strumosa Salisb.
synonymEclipta sulcata Raf.
synonymEclipta thermalis Bunge
synonymEclipta tinctoria Raf.
synonymEclipta undulata Willd.
synonymEclipta zippeliana bl.
synonymEclipta zippeliana Bl.
synonymEclipta zippeliana Blume
synonymEcliptica alba (L.) Kuntze
synonymEcliptica alba var. alba
synonymEcliptica alba var. erecta (Hassk.) Kuntze
synonymEcliptica alba var. erecta Kuntze
synonymEcliptica alba var. parviflora (Wall. ex DC.) Kuntze
synonymEcliptica alba var. prostrata (L.) Kuntze
synonymEcliptica alba var. zippeliana (Blume) Kuntze
synonymEleutheranthera prostrata (L.) Sch.Bip.
synonymEleutheranthera prostrata var. prostrata
synonymEupatoriophalacron album (L.) Hitchc.
synonymGalinsoga oblonga DC.
synonymGalinsoga oblongifolia (Hook.) DC.
synonymGrangea lanceolata Poir.
synonymGrangea lanceolata Poir. [Illegitimate]
synonymMicrelium tolak Forssk.
synonymPaleista brachypoda (Michx.) Raf.
synonymPaleista brachypoda (Michx.) Raf. [Illegitimate]
synonymPolygyne inconspicua Phil.
synonymSpilanthes pseudo-acmella (L.) Murray
synonymVerbesina alba L.
synonymVerbesina alba subsp. alba
synonymVerbesina alba subsp. neapolitana
synonymVerbesina conyzoides Trew
synonymVerbesina prostrata L.
synonymVerbesina pseudoacmella L.
synonymVerbesina spicata Lour.
synonymWedelia psammophila Poepp.
synonymWilborgia oblongifolia Hook.
🗒 Common Names
Bengali
  • Keshuti
Chinese
  • 鳢肠, 鳢肠属, 卤地菊, Li chang, Lǐ cháng shǔ, Lǔ dì jú
Creoles and pidgins; French-based
  • Langue poule (Guyane)
English
  • False daisy, White eclipta, American false daisy
  • Eclipta (USA)
  • Tattoo plant
Français / French
  • Herbe à l'eau
  • Eclipte blanche
  • Eclipte droite
Hindi
  • Bhringraj
  • Bhangra
Indonesian
  • Aring aring, Keremek
Italian
  • Eclipta, Gramigna indiana
Malagasy
  • Ahimarana
  • Kelibehazo
Other
  • M'gawa (Shimaore, Mayotte)
  • Mgawa (Kibushi, Mayotte)
Portuguese
  • Verbesina
  • Erva portão, Erva de botao, Agriao do brejo, Pimenta d'agua, Suricuina, Tangaraca, Ervanço, Cravo bravo (Brazil)
Spanish; Castilian
  • Botoncillo, Botón blanco, Clavel de pozo, Hierba de la muela, Mata de los quesitos, Pestosa
  • Cabeza de pollo, Yerba de tago (Argentina)
  • Florecita (Bolivia)
  • Eclipta blana (Cuba)
  • Hierba prieta (Honduras)
  • Congo tala, Hoja de tinta, Naparo cimarrón (Mexico)
Urdu
  • Daryai buti
📚 Overview
Overview
Brief
Code

ECLAL

Growth form

broadleaf

Biological cycle

annual

Habitat

marshland

Wiktrop
AttributionsWiktrop
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Lovena Nowbut
StatusUNDER_CREATION
LicensesCC_BY
References
    Diagnostic Keys
    Description
    Global description
     
    Eclipta prostrata is an herbaceous plant, usually annual, branched, erect or prostrate, which measures up to 80 cm high. It is entirely covered with short stiff and white hairs. The leaves are opposite, simple, sessile or very shortly-stalked, with elongated oval to lanceolate lamina, rough to the touch. The white flowers are grouped into sub-globular long pedicellate capitulum, in solitary or by 2 or 3, terminal or axillary. The flowers are of 2 types: tubular flowers in the center, more numerous, ligulate flowers on the periphery. The fruit is a compressed warty achene, brown or black in color.
     
    Cotyledons
     
    Cotyledons are stalked (1.5 to 2.5 mm long), glabrous. The lamina is largely oval (4 to 4.8 mm long and 3.2 to 3.6 mm wide), with obtuse base, rounded or truncated apex, with entire margin.
     
    First leaves
     
    The first leaves are short-stalked, simple and opposite. The petiole measures 1 to 1.5 mm long. It is densely covered with applied hairs. The lamina is oval (4.5 to 6.8 mm long and 3.2 to 3.8 mm wide), with obtuse base and rounded tip. It is covered on both sides with short stiff and white hairs. Venation is pinnate. The margin is slightly serrated near the base and ciliated.
     
    General habit
     
    Annual or briefly perennial herbaceous plant, branched, erect or prostrate, then erect, measuring 10 to 80 cm high. The part of the stem in contact with the ground take roots at the nodes. The plant is completely covered with short white stiff and applied hair. The whole plant turns black as it dries.
     
    Underground system
     
    The plant has a taproot system.

    Stem
     
    The stem is cylindrical, slightly fleshy, more or less woody at the base. It is densely covered with short stiff white hairs, applied and directed upwards. Its color varies from green to violet, is in general rather reddish.
     
    Leaf
     
    The leaves are opposite and simple. The upper leaves are sessile, the lower ones have a short petiole. The leaf blade is oval-lanceolate, with abruptly narrowed base and pointed apex. The margin is entire or marked with fine spaced tines. Both sides of the leaf blade are quite densely covered with short and applied hairs making them rough to the touch. The leaf measure 2 to 15.5 cm long and 0.5 to 4 cm wide. It has three nerves at the base.
     
    Inflorescence
     
    The flowers are grouped in white extensively pedicellate capitulum, solitary or grouped by 2 or 3, at the ends of branches or leaf axils. The pedicels measure 1 to 7 cm long. They are covered with stiff applied hairs facing up. The flower heads are sub-globular in the shape of largely open bell, about 1 cm in diameter. The bracts of the receptacle are few (5-6) and nested. They are ovate to orbicular, pointed, covered with dense applied hairs. External bracts are 4 to 6 mm long, the internals are usually shorter.
     
    Flower
     
    The flowers are white, of 2 types. The flowers in the center are tubular, bisexual, small and numerous. These of the margin are ligulate, female or neutral. They are less numerous and form a fringe around the capitulum.
     
    Fruit
     
    The fruit is a cuneiform compressed achene, 2 to 3 mm long, with more or less thickened margin, very warty. It is brown or black. It lacks egret but have short white hairs at the top. This plant that flowers throughout the year has a very important seed production (a plant can produce up to 17,000 seeds).

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      No Data
      📚 Natural History
      Life Cycle

      Life cycle

      Annual
      Annual

      The seeds of Eclipta prostrata have no dormancy and can germinate early in the growing season.

      Brazil: Eclipta prostrata has a cycle that can be very rapid, flowering 4 to 6 weeks after germination and fruiting after 100 to 120 days.
      China: Eclipta prostrata flowers and fruits from June to September.
      Mayotte: Eclipta prostrata flowers from August to February and bears fruit from September to March.
      Morocco: Eclipta prostrata flowers from June to October.
      Nicaragua: Eclipta prostrata flowers and fruits all year round.

       

      Wiktrop
      AttributionsWiktrop
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      StatusUNDER_CREATION
      LicensesCC_BY
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        Cyclicity

        Eclipta prostata is mainly an annual plant (sometimes briefly sustainable), reproducing by seed. The seeds produced in large quantities are dispersed by wind, water and animals.

        Wiktrop
        AttributionsWiktrop
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        StatusUNDER_CREATION
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        References
          Morphology

          Growth form

          Erected
          Erected
          Prostrated
          Prostrated

          Type of prefoliation

          Leaf ratio medium
          Leaf ratio medium
          Narrow leaf
          Narrow leaf

          Equality of opposite leaves

          Opposite leaves equal
          Opposite leaves equal

          Latex

          Without latex
          Without latex

          Root type

          Taproot
          Taproot

          Stipule type

          No stipule
          No stipule

          Leaf attachment type

          sessile
          sessile
          with petiole
          with petiole

          Achene type

          Achene without pappus
          Achene without pappus

          Lamina base

          attenuate
          attenuate

          Lamina margin

          denticulate
          denticulate
          entire
          entire

          Lamina apex

          attenuate
          attenuate

          Simple leaf type

          Lamina elliptic
          Lamina elliptic

          Inflorescence type

          Capitule with tubular and ligulate flowers
          Capitule with tubular and ligulate flowers

          Stem pilosity

          Less hairy
          Less hairy

          Stem hair type

          Bifidus
          Bifidus
          Pubescent
          Pubescent

          Life form

          Broadleaf plant
          Broadleaf plant
          Ecology

          Eclipta prostata adapts well to changing environmental conditions. Often on little draining land, along the rivers and ditches, marshes, in full sun. Weed common in flooded rice fields. From sea level up to 2000 m altitude.

          Brazil: Eclipta prostrata grows in regions with a warm climate and damp soil. It is particularly fond of permanently flooded areas with low water levels. It tolerates salty soils where it can become dominant.
          Central Africa: Eclita prostrata is found in more or less humid ruderal and post-cultivation environments, wet ditches, riverbanks, lakes and swamps; up to 2000 m altitude.
          French Guiana: Eclipata prostrata likes very humid environments. It is very frequent in the rice fields of Mana, especially along the edges of the plots and along the canals. It is rather rare in vegetables and fruit growing plots, except in very humid vegetable plots.
          Madagascar: E. prostrata grows on slightly flooded hydromorphic soils, moist alluvial soils, rich enough, in the sub-humid and humid region. It is present from sea level to an altitude of 1200 m on the edges of canals in the Highlands, the Middle West, West, North-western and eastern side of Madagascar. It is a weed of crops on humid soil on sunny and or lightly shaded ground. It is particularly common in pluvial rice cultivation on more or less humid terrace, or in lowland without water control; vegetables crops, sugar cane, taro, in more or less extensive cropping systems.
          Mauritius: ruderal and weed of crops, at all altitudes, but especially common along the coast.
          Mayotte: E. prostrata is a naturalized species in a wide range of secondarized and generally humid environments. It is present from the coast, to crops, ditches and urban areas.
          Morocco: Eclipta prostrata is an exotic, naturalized and very rare species. It grows in marshes, ditches, wet wastelands and riparian forests up to 500 m altitude.
          South Africa: Eclipta prostrata grows along rivers, water furrows and around ponds. Wet sites such as the edges of marshes, the banks of rivers or lakes, the edges of rice paddies, black clay soils with cotton.
          West Indies: Eclipta prostrata is a ruderal and shrubby species found in flat, damp environments, especially clay, gutters and ditches.

           

          Wiktrop
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            No Data
            📚 Habitat and Distribution
            General Habitat

            Habitat

            Terrestrial
            Terrestrial
            Marshland
            Marshland
            Description

            Origin

            Eclipta prostrata is native to tropical America.

            Worldwide distribution

            Eclipta prostata is currently widespread in the tropics and sub-tropics worldwide and warm temperate regions. Common in the rice fields at altitude. Southeast Asia, Indonesia, China, India, Pakistan, Sri Lanka, Southern Africa, North, Central and Western, South America, USA, Southern Europe, Australia, the Pacific Islands.

            Wiktrop
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            StatusUNDER_CREATION
            LicensesCC_BY_SA
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              No Data
              📚 Occurrence
              No Data
              📚 Demography and Conservation
              Risk Statement

              Global harmfulness

              Eclipta prostata is a common weed in flooded rice fields, it can become problematic, particularly in region of low altitude with high rainfall.
               
              Local harmfulness

              Benin: E. prostrata is rare and scarce in rice fields.
              Brazil: Eclipta prostrata is an important weed in the North and North-East regions, for example in onion crops.
              Burkina Faso: frequent and scarce.
              Ivory Coast: rare and scarce.
              Ghana: frequent and generally abundant.
              Mali: frequent and generally abundant.
              Madagascar: It is a weed quite frequent with a relatively low abundance. It does not present any particular difficulties in the fight except in directly sown crops (nurseries, direct seeding of rice, vegetables).
              Mauritius: A weed of crops with a very low frequency and abundance and therefore quite minimal harmfulness.
              Nigeria: rare and scarce.
              Senegal: frequent and generally abundant.
              Uganda: rare and scarce.
              Tanzania: frequent and scarce.

               

              Wiktrop
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              StatusUNDER_CREATION
              LicensesCC_BY
              References
                Threats

                Madagascar: Eclipta prostata is an secundary host of nematodes as Meloidogyne incognita.

                 

                Wiktrop
                AttributionsWiktrop
                Contributors
                StatusUNDER_CREATION
                LicensesCC_BY
                References
                  No Data
                  📚 Uses and Management
                  Uses
                  Medicinal: Eclipta prostata is a source-based compounds "coumestan" used in pharmaceutical preparations prescribed in cases of liver cirrhosis and liver disease. It is used in traditional medicine to treat diabetes, asthma and liver diseases and scalp.
                  Food: It is also used as wild edible plant.
                  Other: The leaves are used as black coloring or hair toning.
                  Wiktrop
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                  StatusUNDER_CREATION
                  LicensesCC_BY_SA
                  References
                    Management
                    Global control
                     
                    Mechanical control: Eclipta prostrata is easily controlled manual by weeding and by cultivation.
                    Chemical Controls: amitrole is very effective. 2,4-D (1 to 1.5 kg / ha dissolved in 200 liters of water) and MCPA (0.5 to 1 kg / ha dissolved in 200 liters of water) are less effective and more applications are necessary.
                     
                    For weeding Advice broadleaf annual weeds of irrigated rice and lowland in Africa, visit: http://portal.wiktrop.org/document/show/20
                     
                    Local control
                     
                    Madagascar: The manual control of Eclipta prostrata is time consuming when it is abundant.
                    Chemically it is easily controlled to 2,4-D amine or glyphosate on young and old plants.
                    E. prostrata is well controlled by mulching and gradually disappears through direct seeding mulch-provided you have sufficient coverage.
                    Mauritius: The fight against this weed is not listed in Mauritius, but its control in sugar cane can be obtained by the recommended standard herbicides.
                     
                     

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                      References
                      1. Grard, P., T. Le Bourgeois, J. Rodenburg, P. Marnotte, A. Carrara, R. Irakiza, D. Makokha, G. kyalo, K. Aloys, K. Iswaria, N. Nguyen and G. Tzelepoglou (2012). AFROweeds V.1.0: African weeds of rice. Cédérom. Montpellier, France & Cotonou, Bénin, Cirad-AfricaRice eds.
                      2. Barthelat, F. 2019. La Flore illustrée de Mayotte. Meze, Paris, France, Collection Inventaires et Biodiversité, Biotope – Muséum national d’Histoire naturelle. 487 p.
                      3. Bosser J., Fergusson I.K. & Soopramanien C., Mult. an. Flore des Mascareignes. La Réunion, Maurice, Rodrigues, MSIRI, IRD, Kew.
                      4. Flora Maroccana (Dobignard A.) https://www.floramaroccana.fr/eclipta-prostrata.html
                      1. Marita I.G., Keith Moody, Colin M. Piggin. 1999. Upland Rice Weeds of Southeast Asia, IRRI.
                      1. Husson, O., H. Charpentier, F.-X. Chabaud, K. Naudin, Rakotondramanana et L. Séguy (2010). Flore des jachères et adventices des cultures. Annexe 1 : les principales plantes de jachères et adventices des cultures à Madagascar. In : Manuel pratique du semis direct à Madagascar. Annexe 1 - Antananarivo : GSDM/CIRAD, 2010 : 64 p.
                      2. Berton, A. (2020). Flore spontanée des cultures maraichères et fruitières de Guyane. Guide de reconnaissance des 140 adventices les plus communes des parcelles cultivées. Cayenne, Guyane, FREDON Guyane: 186.https://portal.wiktrop.org/document/show/173
                      3. Marnotte, P. and A. Carrara. (2007). "Plantes des rizières de Guyane." from http://plantes-rizieres-guyane.cirad.fr/.
                      1. Soerjani M., Kostermans A. J. G. H., Tjitrosoepomo G. 1987. Weeds of rice in Indonesia. Balai Pustaka. Jakarta.
                      1. Moody K., Munroe C.E., Lubigan R.T., Paller E.C.Jr. 1984. Major Weeds of the Philippines.
                      1. MOODY K. 1989. Weeds reported in Rice in South and Southeast Asia. IRRI Manila Phillippines 442p.
                      2. ANTOINE R., BOSSER J., FERGUSSON I.K., 1993. FLORE DES MASCAREIGNES : La Réunion, Maurice, Rodrigues. 80 Légumineuses. MSIRI, ORSTOM, KEW. p 181
                      1. Johnson, D.E., (1997). Weeds of rice in West Africa. WARDA, Bouaké. 52p;
                      2. Grard, P., Homsombath, K., Kessler, P., Khuon, E., Le Bourgeois, T., Prospéri, J., Risdale, C. 2006. Oswald V.1.0: A multimedia identification system of the major weeds of rice paddy fields of Cambodia and Lao P.D.R. In Cirad [ed.]. Cirad, Montpellier, France. Cdrom. ISBN 978-2-87614-653-2;
                      3. Supaluk P., Orapin W., Thummaruk S., Somsak R., Virapong P., (2010). Bioactivity Evaluation of Eclipta prostrata Linn: A Potential Vasorelaxant. European Journal of Scientific Research Vol.44 No.2, pp.167-176
                      4. Plants of the World Online https://powo.science.kew.org/taxon/urn:lsid:ipni.org:names:1186403-2
                      5. The World Flora Online https://www.worldfloraonline.org/taxon/wfo-0000108811
                      6. Flora of China http://www.efloras.org/florataxon.aspx?flora_id=2&taxon_id=200023875
                      7. CABI https://www.cabidigitallibrary.org/doi/10.1079/cabicompendium.20395
                      8. Kissmann, K.G. & Groth, D. 1992. Plantas Infestantes e Nocivas. Sao Paulo.
                      Information Listing > References
                      1. Grard, P., T. Le Bourgeois, J. Rodenburg, P. Marnotte, A. Carrara, R. Irakiza, D. Makokha, G. kyalo, K. Aloys, K. Iswaria, N. Nguyen and G. Tzelepoglou (2012). AFROweeds V.1.0: African weeds of rice. Cédérom. Montpellier, France & Cotonou, Bénin, Cirad-AfricaRice eds.
                      2. Barthelat, F. 2019. La Flore illustrée de Mayotte. Meze, Paris, France, Collection Inventaires et Biodiversité, Biotope – Muséum national d’Histoire naturelle. 487 p.
                      3. Bosser J., Fergusson I.K. & Soopramanien C., Mult. an. Flore des Mascareignes. La Réunion, Maurice, Rodrigues, MSIRI, IRD, Kew.
                      4. Flora Maroccana (Dobignard A.) https://www.floramaroccana.fr/eclipta-prostrata.html
                      5. Marita I.G., Keith Moody, Colin M. Piggin. 1999. Upland Rice Weeds of Southeast Asia, IRRI.
                      6. Husson, O., H. Charpentier, F.-X. Chabaud, K. Naudin, Rakotondramanana et L. Séguy (2010). Flore des jachères et adventices des cultures. Annexe 1 : les principales plantes de jachères et adventices des cultures à Madagascar. In : Manuel pratique du semis direct à Madagascar. Annexe 1 - Antananarivo : GSDM/CIRAD, 2010 : 64 p.
                      7. Berton, A. (2020). Flore spontanée des cultures maraichères et fruitières de Guyane. Guide de reconnaissance des 140 adventices les plus communes des parcelles cultivées. Cayenne, Guyane, FREDON Guyane: 186.https://portal.wiktrop.org/document/show/173
                      8. Marnotte, P. and A. Carrara. (2007). "Plantes des rizières de Guyane." from http://plantes-rizieres-guyane.cirad.fr/.
                      9. Soerjani M., Kostermans A. J. G. H., Tjitrosoepomo G. 1987. Weeds of rice in Indonesia. Balai Pustaka. Jakarta.
                      10. Moody K., Munroe C.E., Lubigan R.T., Paller E.C.Jr. 1984. Major Weeds of the Philippines.
                      11. MOODY K. 1989. Weeds reported in Rice in South and Southeast Asia. IRRI Manila Phillippines 442p.
                      12. ANTOINE R., BOSSER J., FERGUSSON I.K., 1993. FLORE DES MASCAREIGNES : La Réunion, Maurice, Rodrigues. 80 Légumineuses. MSIRI, ORSTOM, KEW. p 181
                      13. Johnson, D.E., (1997). Weeds of rice in West Africa. WARDA, Bouaké. 52p;
                      14. Grard, P., Homsombath, K., Kessler, P., Khuon, E., Le Bourgeois, T., Prospéri, J., Risdale, C. 2006. Oswald V.1.0: A multimedia identification system of the major weeds of rice paddy fields of Cambodia and Lao P.D.R. In Cirad [ed.]. Cirad, Montpellier, France. Cdrom. ISBN 978-2-87614-653-2;
                      15. Supaluk P., Orapin W., Thummaruk S., Somsak R., Virapong P., (2010). Bioactivity Evaluation of Eclipta prostrata Linn: A Potential Vasorelaxant. European Journal of Scientific Research Vol.44 No.2, pp.167-176
                      16. Plants of the World Online https://powo.science.kew.org/taxon/urn:lsid:ipni.org:names:1186403-2
                      17. The World Flora Online https://www.worldfloraonline.org/taxon/wfo-0000108811
                      18. Flora of China http://www.efloras.org/florataxon.aspx?flora_id=2&taxon_id=200023875
                      19. CABI https://www.cabidigitallibrary.org/doi/10.1079/cabicompendium.20395
                      20. Kissmann, K.G. & Groth, D. 1992. Plantas Infestantes e Nocivas. Sao Paulo.

                      Etude floristique et phytoécologique des adventices des complexes sucriers de Ferké 1 et 2, de Borotou-Koro et de Zuenoula, en Côte d'Ivoire

                      Thomas Le Bourgeois
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