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WIKTROP - Weed Identification and Knowledge in the Tropical and Mediterranean areas
WIKTROP - Weed Identification and Knowledge in the Tropical and Mediterranean areas
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Imperata cylindrica (L.) Raeusch.

Accepted
Imperata cylindrica (L.) P.Beauv.
Imperata cylindrica (L.) P.Beauv.
Imperata cylindrica (L.) P.Beauv.
Imperata cylindrica (L.) P.Beauv.
Imperata cylindrica (L.) P.Beauv.
Imperata cylindrica (L.) P.Beauv.
Imperata cylindrica (L.) P.Beauv.
Imperata cylindrica (L.) P.Beauv.
Imperata cylindrica (L.) P.Beauv.
Imperata cylindrica (L.) P.Beauv.
Imperata cylindrica (L.) P.Beauv.
Imperata cylindrica (L.) P.Beauv.
Imperata cylindrica (L.) P.Beauv.
Imperata cylindrica (L.) P.Beauv.
Imperata cylindrica (L.) P.Beauv.
Imperata cylindrica (L.) P.Beauv.
Imperata cylindrica (L.) P.Beauv.
Imperata cylindrica (L.) P.Beauv.
Imperata cylindrica (L.) P.Beauv.
Imperata cylindrica (L.) P.Beauv.
Imperata cylindrica (L.) P.Beauv.
Imperata cylindrica (L.) P.Beauv.
Imperata cylindrica (L.) P.Beauv.
Imperata cylindrica (L.) P.Beauv.
Imperata cylindrica (L.) P.Beauv.
Imperata cylindrica (L.) P.Beauv.
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Imperata cylindrica (L.) P.Beauv.
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🗒 Synonyms
synonymArundo epigeios Forssk. ex Steud. [Invalid]
synonymArundo epigeios Forssk. ex Steud., pro syn.
synonymCalamagrostis lagurus Koeler [Illegitimate]
synonymCalamagrostis lagurus Koeler, nom. superfl.
synonymImperata allang Jungh.
synonymImperata angolensis Fritsch
synonymImperata arundinacea Cirillo
synonymImperata arundinacea var. africana Andersson
synonymImperata arundinacea var. europaea Andersson
synonymImperata arundinacea var. glabrescens Büse
synonymImperata arundinacea var. indica Andersson
synonymImperata arundinacea var. koenigii (Retz.) Benth.
synonymImperata arundinacea var. latifolia Hook.f.
synonymImperata arundinacea var. pedicellata (Steud.) Debeaux
synonymImperata arundinacea var. thunbergii (Retz.) Stapf, nom. superfl.
synonymImperata cylindrica f. pallida Honda
synonymImperata cylindrica subsp. koenigii (Retz.) Tzvelev
synonymImperata cylindrica var. africana (Andersson) C.E.Hubb.
synonymImperata cylindrica var. cylindrica
synonymImperata cylindrica var. europaea (Andersson) Asch. & Graebn.
synonymImperata cylindrica var. koenigii (Retz.) Pilg.
synonymImperata cylindrica var. latifolia (Hook.f.) C.E.Hubb.
synonymImperata cylindrica var. major (Nees) C.E.Hubb.
synonymImperata cylindrica var. parviflora Batt. & Trab.
synonymImperata cylindrica var. pedicellata (Steud.) Debeaux
synonymImperata cylindrica var. thunbergii (Retz.) T.Durand & Schinz
synonymImperata cylindrica var. thunbergii (Retz.) T.Durand & Schinz, nom. superfl.
synonymImperata dinteri Pilg.
synonymImperata filifolia Nees ex Steud.
synonymImperata koenigii (Retz.) P.Beauv.
synonymImperata koenigii var. major Nees
synonymImperata laguroides (Pourr.) J.Roux
synonymImperata latifolia (Hook.f.) L.Liou
synonymImperata latifolia (Hook.f.) L.Liu
synonymImperata pedicellata Steud.
synonymImperata praecoquis Honda
synonymImperata robustior A.Chev.
synonymImperata sieberi Opiz
synonymImperata sisca P.Beauv. ex Steud. [Invalid]
synonymImperata sisca P.Beauv. ex Steud., pro syn.
synonymImperata thunbergii (Retz.) Nees
synonymLagurus cylindricus L.
synonymSaccharum cylindricum (L.) Lam.
synonymSaccharum cylindricum var. europaeum Pers.
synonymSaccharum diandrum J.Koenig ex Retz. [Invalid]
synonymSaccharum diandrum J.Koenig ex Retz., nom. nud.
synonymSaccharum europaeum Pers. [Illegitimate]
synonymSaccharum europaeum Pers., nom. superfl.
synonymSaccharum indum Pers. [Invalid]
synonymSaccharum indum Pers., pro syn.
synonymSaccharum koenigii Retz.
synonymSaccharum laguroides Pourr.
synonymSaccharum sisca Cav.
synonymSaccharum spicatum Burm. ex Kunth, pro syn.
synonymSaccharum spicatum J.Presl [Illegitimate]
synonymSaccharum spicatum J.Presl, nom. superfl.
synonymSaccharum thunbergii Retz.
🗒 Common Names
Comorian
  • Bidambé, Mbidambe
English
  • Blady grass, Cogon grass
  • Bedding grass, Japanese blood grass, Silver spike, Spear grass, Sword grass, Thatch grass
French
  • Paille de diss (Nouvelle-Calédonie)
  • Impérata cylindrique, Impérate, Paillotte
Malagasy
  • Manevika, Tenina
Other
  • Unyasi (Shimaore, Mayotte)
  • Maneviki (Kibushi, Mayotte)
Spanish; Castilian
  • Carrizo, Cisca, Marciega
📚 Overview
Overview
Brief
Code

IMPCY

Growth  form

grass

Biological cycle

vivacious

Habitat

terrestrial

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    Diagnostic Keys
    Description
    Global description
     
    Imperata cylindrica is a vivacious grass with rhizomes. It grows in clumps with ribbon-like leaves, vertically erect, pointed at the top. The culm is rigid, cylindrical bearing a fusiform inflorescence which is fluffy, and silvery white in colour.
     
    First leaves
     
    Seedlings from germination are rare due to the low fertility of the seeds. The young plant comes from a rhizome that straightens vertically out of the ground. The underground leaves are reduced to nested scales. Above the ground surface, the leaf blade develops into a vertical ribbon, rigid, very acute at the apex. The prefoliation is rolled.
     
    General habit
     
    Erect vivacious grass. The plant has many erect culms, connected by creeping rhizomes.
     
    Underground system
     
    The underground system is made up of underground stems consisting of creeping rhizomes, branched, from which tufts of erect shoots  regularly emerge.
     
    Culm
     
    The culm is rigid, slender, cylindrical, reaching up to 0.5 m to 1 m high. At the base of the clumps, the remnants of fibrous sheaths of the previous year persist.
     
    Leaf
     
    The leaves are mostly basal. They are vertically erect. The sheath is fibrous and short. It has a  striped leaf blade, which attenuates in a sharp point, pungent at the top. The base of the blade is narrowed into a pseudo-petiole. The leaves are up to 1 m long and 5 to 18 mm wide. The leaf blade is smooth on both sides. The margin is more or less scabrous. The central rib, clearer, appears slightly convex on the underside. Ligule is membranous and inconspicuous (1 mm long). It is lined with a few stiff hairs at the base of the margins of the leaf blade.
     
    Inflorescence
     
    The inflorescence is a cylindrical fusiform panicle, loose and silky. It is silvery-white in colour, dotted with orange stamens and brown stigmas. It measures 5 to 20 cm long and 0.5 to 2.5 cm wide.
     
    Spikelet
     
    The spikelet, 2.5 to 5 mm long, is surrounded by long white bristles of 1 cm. The lower and upper glumes are lanceolate, ciliated on the top.
     
    Grain
     
    The grain is ellipsoid, pointed at the top, 1 mm long.

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      No Data
      📚 Natural History
      Life Cycle

      Life cycle

      Vivacious
      Vivacious

      Mayotte: Imperata cylindrica flowers and fruits all year round.
      New Caledonia:
      Imperata cylindrica enters into vegetation as soon as the soil is wet. Flowering and fruiting take place from September to December, at the beginning of the dry season.

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        Reproduction
        Imperata cylindrica is a vivacious species. It multiplies mainly vegetatively by extension of vigorous rhizomes system. It can also reproduce by seeds spread by wind over long distances. Some populations produce flowers regularly, while others don’t. Fire, mowing, grazing, induce renewed vegetation and stimulate flowering. If the aerial parts burn easily, rhizomes withstand high temperatures. The rhizomes can penetrate the soil to a depth of 120 cm, but the majority are in the top 15 centimeters of heavy ground and in the first 40 centimeters in light soil. Fragments of rhizomes of 1 to 5 cm long are adapted to bud. Rhizomes fragments placed on the surface of the ground withstand desiccation very badly. They cannot bud after two days at a humidity lower than 47%. The optimum temperature for development is 30 ° C.

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          Morphology

          Growth form

          Tuft plant with narrow leaves
          Tuft plant with narrow leaves

          Leaf type

          Grass or grass-like
          Grass or grass-like

          Latex

          Without latex
          Without latex

          Root type

          Rhizome
          Rhizome

          Ligule type

          Ligule membranous large
          Ligule membranous large

          Stipule type

          No stipule
          No stipule

          Leaf attachment type

          with graminate sheathing
          with graminate sheathing

          Fruit type

          Grain of grasses
          Grain of grasses

          Lamina base

          sheathing grass-like smaller
          sheathing grass-like smaller

          Lamina margin

          scabrous
          scabrous
          entire
          entire

          Lamina apex

          attenuate
          attenuate

          Simple leaf type

          Lamina linear
          Lamina linear

          Life form

          Grass
          Grass
          Geophytic plant
          Geophytic plant
          Look Alikes

          Imperata cylindrica is very similar to I. brasiliensis Trin. (present from southern USA to northern Argentina and in the Caribbean) and I. contracta (Kunth) Hitchc. (present from southern Mexico to northern Argentina, in the Caribbean and introduced in the West Indies as an ornamental plant). However, I. cylindrica is the only species of this genus to have 2 stamens per flower (the other species have only one), moreover, it is essentially present in Africa and Asia.

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            Ecology

            Imperata cylindrica is a plant of tropical zone to hot temperate. It develops in extremely varied environments. It can be found in arid, sandy areas as well as in swampy areas or along waterways, in cultivated areas (formidable weed of many annual or perennial crops), in peri-urban areas, from sea level to about 2700 m of altitude, under rainy regimes varying from 500 to 5000 mm per year.
            Primary natural forests and secondary forests with a lot of shade are not favorable to the development of this plant but can be colonized in case of disturbance; it is a very frequent weed of the clearing-burn areas. Indeed the activity of the cauline buds of the rhizomes is greatly favored by the passage of the fire or by the ploughings, in particular with the plough with discs, which by splitting the rhizomes, multiplies and disseminates the plant.

            Benin: Imperata cylindrica is found throughout the country from the coastal zone around Cotonou to the northern savannahs (Tanguiéta, Segbana).
            Northern Cameroon
            : Imperata cylindrica is a species of humid climate, very abundant in transition forest in Sudan and Guinea savanna zones. It grows preferentially in climates where annual rainfall exceeds 1300 mm.It occasionally grows in drier climates, in humid soils along the river. It tolerates short periods of flooding. This species is very sun-loving, and it grows mainly as weed of crops or as ruderal on roadsides and in open areas. It has no significant preference for the soil type.
            Comoros: Sun loving plant, frequent in wetlands, especially at low altitudes up to 400 m. It is present especially in zones regularly set on fire. It is observed in the three islands.
            Madagascar: Weed with powerful rhizomes, fairly widespread in the highlands and the eastern side of the island. It infests fallows and crops on terraces with deep soil down slopes and on alluvial plains. The presence of this weed is often considered as a degradation of soil fertility.
            Mauritius: absent.
            Mayotte: Imperata cylindrica is an exotic species. Weed of crops in burned areas, because fire favors the germination of seeds and limits the competition. It is present in crops, pastures and also in padza (deforested and strongly gullied areas with red lateritic soil).
            New Caledonia: Species of humid climate (greater than 1500 mm rainfall), it prefers deep and draining soils. It is found preferentially on acidic soils in New Caledonia where it could be native. It spreads in areas often burned, because fire stimulates the regrowth of the plant.
            Reunion: absent.
            Seychelles: absent.

             

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              📚 Habitat and Distribution
              Description

              Geographical distibution

              Madagascar
              Madagascar
              Comoros
              Comoros

              Origin

              Imperata cylindrica is native of the tropical regions of the old world (tropical and southern Africa until Madagascar); ubiquitous species for the countries of this zone.

              Worldwide distribution

              Imperata cylindrica is a very widespread species in Africa, Australia, South Asia, India, North America (Alabama, Florida, Georgia, Louisiana, Mississippi, Oregon, South Carolina, Texas) and Central America (Mexico). It is present, but not a problem, in Southern Europe and around the Mediterranean.


              Northern Cameroon: Imperata cylindrica is present south of Garoua and occasionally further north in riverside plots.

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                📚 Occurrence
                No Data
                📚 Demography and Conservation
                Risk Statement

                Global harmfulness
                 
                Imperata cylindrica is included in the IUCN list of the 100 most invasive species in the world, as well as the 18 most damaging weeds in the world.
                I. cylindrica is capable of invading many different types of environments and preventing the natural regeneration of native species. One of the major reasons for the invasiveness of this species is its rhizome network, concentrated in the first 20 centimeters of soil, particularly dense and efficient and immune to fire and some control methods. The colonization of a new environment by this plant induces deep changes in the functioning of the concerned ecosystem. Finally, it is a recognized and feared weed for many crops, even in its native area. I. cylindrica is a rhizomatous species whose expansion in the plots is greatly favored by the actions of splitting at the time of the cultural operations (ploughing, weeding, ridging). Ploughing with a disc plow during rainy periods is the most favorable case of multiplication.
                Brook (1989), cites up to 64.5 million ha of overgrown grassland in Indonesia, 5 million ha in Papua New Guinea and 300,000 ha in Fiji; more than 40% of rubber plantations in Java and 1.5 to 2 million ha of plantations in Malaysia are affected by the plant. Millions of hectares of grassland in West and Central Africa are abandoned due to the presence of this weed. The area of plantations and agricultural land colonized by this species worldwide is estimated at more than 200 million hectares.
                 
                Local harmfulness
                 
                Northern Cameroon: I. cylindrica is considered a major local weed.
                Comoros: This weed grows in cassava crops and in all cultures with tubers plant, for which it represents a major problem because the pointed rhizomes of I. cylindrica that perforate tubers in the cultures. Moreover, it makes the soil compact and difficult for weeding. It is also an important weed of grasses in meadows.
                Kenya: Rare but abondant when present
                Madagascar: Invasive weed and harmful for rainfed crops slightly intensified (cassava, upland rice Tavy, sweet potato). It often occurs as a dense stand of variable size and poses a serious problem to the development of already infested land (weak equipment and facilities available to farmers).
                Mauritius: absent.
                New Caledonia: It remains moderately invasive on pasture, and is frequently found in waste lands and savannas in small stands.
                Reunion: absent.
                Seychelles: absent.
                Tanzania: Frequent but not abundant.
                Uganda: Frequent and usually abundant.

                 

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                  📚 Uses and Management
                  Uses
                  Medicinal: Known in Malawi as a treatment for digestive complaints and in India it is an important drug of 'Trinpanchmool' used for treatment of various ailments urinary calculi, retention of urine, cardiac disorder, gout, common cough and cold, menorrhegia, inflammation, burning sensation, fever, anemia, and as aphrodisiac, rejuvenator etc.
                  Fodder: Due to its poor foraging value, I. cylindrica is consumed only at the early stage and it is recommended to eradicate it because of its potential spread.
                  Other: Imperata cylindrica is used to cover the traditional Kanak huts in New-Caledonia.
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                    Management

                    Global control
                     
                    Imperata cylindrica is a rhizomatous species whose expansion in the plots is greatly favored by splitting parts of it at the time of cultural operations (plowing, weeding, hilling). Imperata cylindrica is a difficult species to control. Tillage with a plow at the beginning of the dry season, brings many rhizomes at the surface, causing them to dry out. This species is insensitive to pre-emergence herbicides and contact, however it is sensitive to systemic post-emergence herbicides.
                    For perennial grass weed control tips for irrigated rice and lowland in Africa, visit: http://portal.wiktrop.org/document/show/33

                    Chemical control: A chemical control can be considered during the vegetative phase with fairly high dose of glyphosate (2880 g / ha) absorbed by the leaves and penetrating the rhizomes, killing of the plant within 4 to 6 weeks. The haloxyfop (864 g / ha) also proves to be effective, however, regrowth is possible. In the case of discharges, it is advisable to deal with localized applications. In all cases, ensure the rapid implementation of new forage species able to cover the space quickly.

                     
                    Local control


                    Comoros: The management of this species is practiced with weeding stakes made of iron or wood, to extract the rhizomes.
                    Madagascar: Tillage aids to the multiplication of I. cylindrica by splitting the rhizomes. It is also favored by fires and / or overgrazing, re-developing very rapidly from its rhizomes, which allows it to dominate other plants. It is insensitive to most selective herbicides of crops and supports short floods. However, it can be quickly controlled during seedlings: glyphosate treatment (1800 to 2160 g / ha) and maintain the mulch.(which seems to have a limiting effect on its recovery)  that have been formed allow the growth of a culture that one can associate with a vegetal cover (Mucuna, stylosanthes, brachiaria, etc.), which thus allow to keep this vegetal pest in shade and destroy it.
                    New Caledonia: Although it is difficult to control, I. cylindrica can be treated mechanically by plowing during dry weather, so that the rhizomes dry in the sun, repeating the operation if necessary. Never burn as it increases its proliferation.

                     

                     

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                      📚 Information Listing
                      References
                      1. Ivens G. W., Moody K. & Egunjobi J. K., 1978. West African Weeds. Oxford University Press, Ibadan, Nigeria, 255 p.
                      1. Le Bourgeois Th., 1993. Les mauvaises herbes dans la rotation cotonnière au Nord-Cameroun (Afrique) - Amplitude d'habitat et degré d'infestation - Cycle de développement. Thèse USTL Montpellier II, Montpellier, France, 241 p.
                      1. Le Bourgeois, T., A. Carrara, M. Dodet, W. Dogley, A. Gaungoo, P. Grard, Y. Ibrahim, E. Jeuffrault, G. Lebreton, P. Poilecot, J. Prosperi, J. A. Randriamampianina, A. P. Andrianaivo and F. Théveny (2008). Advent-OI : Principales adventices des îles du sud-ouest de l'Océan Indien. Cédérom. Montpellier, France, Cirad ed.
                      1. Merlier H. & Montégut J., 1982. Adventices tropicales. ORSTOM-GERDAT-ENSH éd., Montpellier, France, 490p.
                      1. Zon van der A.P.M., 1992. Graminées du Cameroun, Vol. II, Flore. Wageningen Agric. Univ. Papers 92 - 1, Wageningen, 557 p.
                      1. Berhaut J., 1967. Flore du Sénégal. 2ème éd. Clairafrique éd., Dakar, Sénégal, 485 p.
                      1. Blanfort, V., Desmoulins, F., Prosperi, J., Le Bourgeois, T., Guiglion, R., Grard, P. 2010. AdvenPaC V.1.0 : Adventices et plantes à conflit d'intérêt des Pâturages de Nouvelle-Calédonie. IAC, Cirad, Montpellier, France. Cédérom. ISBN 978-2-87614-670-9.
                      1. Grard, P., T. Le Bourgeois, J. Rodenburg, P. Marnotte, A. Carrara, R. Irakiza, D. Makokha, G. kyalo, K. Aloys, K. Iswaria, N. Nguyen and G. Tzelepoglou (2012). AFROweeds V.1.0: African weeds of rice. Cédérom. Montpellier, France & Cotonou, Bénin, Cirad-AfricaRice eds.Le Bourgeois, T., Merlier, H. 1995. Adventrop - Les adventices d'Afrique soudano-sahélienne. Cirad, Montpellier, France.
                      1. Holm L. G., Plucknett D. L., Pancho J. V. & Herberger J. P., 1977. The World's Worst Weeds : Distribution and Biologie. East-West Center, University Press of Hawaii, Honolulu, 609 p.
                      1. Husson, O., H. Charpentier, F.-X. Chabaud, K. Naudin, Rakotondramanana et L. Séguy (2010). Flore des jachères et adventices des cultures. Annexe 1 : les principales plantes de jachères et adventices des cultures à Madagascar. In : Manuel pratique du semis direct à Madagascar. Annexe 1 - Antananarivo : GSDM/CIRAD, 2010 : 64 p.
                      1. Akobundu I.O. & Agyakwa C.W., 1989. Guide des adventices d'Afrique de l'Ouest. Institut international d'agriculture tropicale. Ibadan, Nigeria, 521p.
                      1. Le Bourgeois, T., Camou, R. & Ehret, P. 2006. Analyse de risque phytosanitaire - Appui à la rédaction de la réglementation spécifique aux départements d'outre-mer. Cas des plantes envahissantes. Montpellier, France, Poseidom, Cirad, Dgal-Sdqpv. CDrom
                      2. Barthelat, F. 2019. La Flore illustrée de Mayotte. Meze, Paris, France, Collection Inventaires et Biodiversité, Biotope – Muséum national d’Histoire naturelle.
                      3. Brook, R. M. ; 1989. Review of literature on Imperata cylindrica (L.) Raeuschel with particular reference to South East Asia. Tropical Pest Management, 35 (1) : 12-25.
                      4. Koger,C. H., Bryson, C. T. ; 2004. Effect of cogongrass (Imperata cylindrica) extracts on germination and seedling growth of selected grass and broadleaf species. Weed Technology. 18 (2) : 236-242.
                      5. MacDonald, G. ; 2004. Cogongrass (Imperata cylindrica) - biology, ecology, and management. Critical Reviews in Plant Sciences. 23 (5) : 367-380.
                      6. Radanachaless T., Maxwell J.F. 1994. Weeds of soybean fields in Thailand. Multiple Cropping Center ed., Chiang Mai Univ., Chiang Mai, Thailand, 408.
                      1. S. Jayalakshmi1, Arjun Patra1, V.K. Lal, A.K. Ghosh (2010). Pharmacognostical standardization of roots of Imperata cylindrica Linn (poaceae). J. Pharm. Sci. & Res. Vol.2 (8), 472-476;
                      2. Chikoye, D., Manyong, V.M., Ekeleme, F., (2000). Characteristics of speargrass (Imperata cylindrica) dominated fields in West Africa: crops, soil properties, farmer perceptions and management strategies. Crop Prot. 19, 481-487; Chikoye, D., Ekeleme, F., Ambe, J.T., 1999. Survey of distribution and farmer perceptions of speargrass (Imperata cylindrica) in cassava based cropping systems in West Africa. International Journal of Pest Management 45, 305-312;
                      3. Johnson, D.E., Riches, C.R., Diallo, R., Jones, M.J., (1997). Striga on rice in West Africa; crop host range and the potential of host resistance. Crop Prot. 16, 153-157;
                      4. K. A. Nyarko and S. K. Detta (1991). A hand book for weed control in Rice. International Rice Research Institute. Los Banos, Phillipine. 102p;
                      5. R. J. HILLOCKS (1998). The potential benefits of weeds with reference to small holder agriculture in Africa. Natural Resources Institute, University of Greenwich. Integrated Pest Management Reviews 3, 155-167;
                      6. Townson, J., 1991. Imperata cylindrica and its control. Weed Abstracts 40, 457-468; Troupin G. (1989). Flore du Rwanda, Spermatophyte (Volume IV). Musée Royal de l'Afrique centrale, Tervuren, Belgique. 299-300 p
                      7. I O AKOBUNDU & F E EKELEME (2000). Effect of method of Imperata cylindrica management on maize grain yield in the derived savannah of south-western Nigeria. International Institute of Tropical Agriculture, Ibadan Nigeria. Blackwell Science Weed Research 40, 335-341;
                      1. https://www.cabi.org/isc/datasheet/28580
                      2. https://www.feedipedia.org/node/425
                      1. https://powo.science.kew.org/taxon/urn:lsid:ipni.org:names:30138371-2
                      2. http://www.issg.org/database/species/ecology.asp?si=16&fr=1&sts=sss&lang=FR
                      Information Listing > References
                      1. Ivens G. W., Moody K. & Egunjobi J. K., 1978. West African Weeds. Oxford University Press, Ibadan, Nigeria, 255 p.
                      2. Le Bourgeois Th., 1993. Les mauvaises herbes dans la rotation cotonnière au Nord-Cameroun (Afrique) - Amplitude d'habitat et degré d'infestation - Cycle de développement. Thèse USTL Montpellier II, Montpellier, France, 241 p.
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                      13. Barthelat, F. 2019. La Flore illustrée de Mayotte. Meze, Paris, France, Collection Inventaires et Biodiversité, Biotope – Muséum national d’Histoire naturelle.
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                      25. https://www.cabi.org/isc/datasheet/28580
                      26. https://www.feedipedia.org/node/425
                      27. https://powo.science.kew.org/taxon/urn:lsid:ipni.org:names:30138371-2
                      28. http://www.issg.org/database/species/ecology.asp?si=16&fr=1&sts=sss&lang=FR

                      Etude floristique et phytoécologique des adventices des complexes sucriers de Ferké 1 et 2, de Borotou-Koro et de Zuenoula, en Côte d'Ivoire

                      Thomas Le Bourgeois
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                        WIKTROP - Weed Identification and Knowledge in the Tropical and Mediterranean areasWIKTROP - Weed Identification and Knowledge in the Tropical and Mediterranean areas
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