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Vascular Plants of California
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Woodsia scopulina


Higher Taxonomy
Family: WoodsiaceaeView Description 
Common Name: CLIFF FERN FAMILY
Habit: Perennial herb in rock crevices, occasionally in soil; rhizome generally short-creeping. Leaf: short-spaced, 5--40 cm, +- alike; stipe firm or fleshy (easily crushed), base darker or not, with 2 vascular strands; blade 1-pinnate to 2-pinnate-pinnately lobed, herbaceous, generally with hairs, hair-like scales, or gland-tipped hairs on axes, veins free, terminating before leaf margin; rachis, costa generally grooved adaxially. Sporangia: sori round; indusia of many segmented hair- or scale-like fragments or lobes encircling sorus from below [sphere-like].
Genera In Family: 1 genus, +- 35 species: worldwide, generally temperate. All species often treated in Woodsia (Shao et al. 2015 PLoS ONE 10; Pteridophyte Phylogeny Group 2016 J Syst Evol in press); four to six segregate genera could be recognized (Schmakov 2015 Turczaninowia 18:11--16). Note: A relatively isolated family within the broad eupolypods II clade (suborder Aspleniineae; Pteridophyte Phylogeny Group 2016 J Syst Evol in press), only distantly related to Cystopteridaceae or Athyriaceae, both often included here (Smith et al. 2016 Taxon 55:705--731).
eFlora Treatment Author: Carl J. Rothfels & Alan R. Smith
Scientific Editor: Bruce G. Baldwin & Thomas J. Rosatti.
Genus: WoodsiaView DescriptionDichotomous Key


Common Name: CLIFF FERN
Habit: Rhizome generally ascending to suberect, short, old stipe bases many. Leaf: often glandular or hairy; stipe base ×-section with 2 vascular strands; blade 1--2-pinnate, segments +- toothed to pinnately lobed, veins free, ending just short of margin. Sporangia: sori round, generally not at margins; indusium cup-like, often of many segmented hair- or scale-like fragments or lobes encircling sorus from below [sphere-like], often of crusty, +- white beads, often obscure in age.
Etymology: (J. Woods, Britain, b. 1776)
eFlora Treatment Author: John C. Game, Alan R. Smith and Thomas Lemieux
Reference: Windham 1993 FNANM 2:270--280
Unabridged Reference: Brown 1964 Beih Nova Hedwigia 16:1--154
Woodsia scopulina D.C. Eaton
NATIVE
Leaf: < 32 cm, 1.5--2 cm wide, tip +- acute, unforked; hairs on abaxial leaf axes +- 0.5--1 mm, +- flat, segmented, nonglandular, and +- 0.1 mm, cylindric, non-segmented, glandular; pinnae < 12--27 mm, 5--12 mm wide, pinnately lobed to 1-pinnate, margin toothed to shallowly lobed. Sporangia: indusium of narrow scale-like lobes. Chromosomes: 2n=76,152.
Ecology: Crevices, rock bases; Elevation: 1300--3500 m. Bioregional Distribution: KR, CaRH, SNH, SnBr, MP, W&I; Distribution Outside California: to Alaska, eastern Canada, western United States. Note: If recognized taxonomically, plants differing in chromosome number, spore size [Windham 1993], perhaps other ways assignable to W. scopulina subsp. laurentiana Windham (2n = 152), W. scopulina subsp. scopulina (2n = 76); study needed.
Jepson eFlora Author: John C. Game, Alan R. Smith and Thomas Lemieux
Reference: Windham 1993 FNANM 2:270--280
Index of California Plant Names (ICPN; linked via the Jepson Online Interchange)

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Botanical illustration including Woodsia scopulina

botanical illustration including Woodsia scopulina

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Citation for this treatment: John C. Game, Alan R. Smith and Thomas Lemieux 2012, Woodsia scopulina, in Jepson Flora Project (eds.) Jepson eFlora, https://ucjeps.berkeley.edu/eflora/eflora_display.php?tid=48616, accessed on April 27, 2024.

Citation for the whole project: Jepson Flora Project (eds.) 2024, Jepson eFlora, https://ucjeps.berkeley.edu/eflora/, accessed on April 27, 2024.

Woodsia scopulina
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©2006 Steve Matson
Woodsia scopulina
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©2006 Steve Matson
Woodsia scopulina
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©2017 Neal Kramer
Woodsia scopulina
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©2017 Neal Kramer
Woodsia scopulina
click for enlargement
©2017 Neal Kramer

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Geographic subdivisions for Woodsia scopulina:
KR, CaRH, SNH, SnBr, MP, W&I
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map of distribution 1
(Note: any qualifiers in the taxon distribution description, such as 'northern', 'southern', 'adjacent' etc., are not reflected in the map above, and in some cases indication of a taxon in a subdivision is based on a single collection or author-verified occurence).





 

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CCH collections by month

Duplicates counted once; synonyms included.
Species do not include records of infraspecific taxa, if there are more than 1 infraspecific taxon in CA.
Blue line denotes eFlora flowering time (fruiting time in some monocot genera).